Title: Theodor Schwann
Content:  Theodor Schwann was born in Neuss on 7 December 1810 to Leonard Schwann and Elisabeth Rottels. Leonard Schwann was a goldsmith and later a printer. Theodor Schwann studied at the Dreikönigsgymnasium (also known as the Tricoronatum or Three Kings School), a Jesuit school in Cologne. Schwann was a devout Roman Catholic. In Cologne his religious instructor, a priest and novelist, emphasized the individuality of the human soul and the importance of free will. In 1829, Schwann enrolled at the University of Bonn in the premedical curriculum. He received a bachelor of philosophy in 1831. While at Bonn, Schwann met and worked with physiologist Johannes Peter Müller. Müller is considered to have founded scientific medicine in 
Title: Theodor Schwann
Content:  publishing his Handbuch der Physiologie des Menschen für Vorlesungen in 1837–1840. It was translated into English as Elements of Physiology in 1837–1843 and became the leading physiology textbook of the 1800s. In 1831, Schwann moved to the University of Würzburg for clinical training in medicine. In 1833, he went to the University of Berlin, where Müller was now Professor of Anatomy and Physiology. Schwann graduated with an M.D. degree in medicine from the University of Berlin in 1834. He did his thesis work in 1833–1834, with Müller as his advisor. Schwann's thesis involved a careful study of the necessity for oxygen during the embryonic development of the chicken. To carry it out, he designed and 
Title: Theodor Schwann
Content:  1838 and again in 1846, but was disappointed. Instead, in 1839, Schwann accepted the chair of anatomy at the Université Catholique de Louvain in Leuven, Belgium, another Catholic city. Schwann proved to be a dedicated and conscientious professor. With his new teaching duties, he had less time for new scientific work. He spent considerable time perfecting experimental techniques and instruments for use in experiments. He produced few papers. One exception was a paper in 1844 that reported on a series of experiments on dogs and established the importance of bile in digestion. In examining processes such as muscle contraction, fermentation, digestion, and putrefaction, Schwann sought to show that living phenomena were the result of 
Title: Theodor Schwann
Content:  From 1834 to 1839, Schwann worked as an assistant to Müller in at the Anatomisch-zootomische Museum at the University of Berlin. Schwann carried out a series of microscopic and physiological experiments focused on studying the structure and function of nerves, muscles and blood vessels. In addition to performing experiments in preparation for Müller's book on physiology, Schwann did research of his own. Many of his important contributions were made during the time that he worked with Müller in Berlin. Schwann used newly powerful microscopes to examine animal tissues. This enabled him to observe animal cells and note their different properties. His work complemented that of Matthias Jakob Schleiden in plants and was informed by 
Title: Theodor Schwann
Content:  the two were close friends. Described as quiet and serious, Schwann was particularly gifted in the construction and use of apparatus for his experiments. He was also able to identify important scientific questions and design experiments to systematically test them. His writing has been described as accessible, and his logic as a "clear progression". He identified the question that he wanted to answer and communicated the importance of his findings effectively to others. His co-worker Jakob Henle spoke of him as having an "inborn drive" to experiment. By 1838, Schwann needed a position with a more substantial salary. He hoped to return to Bonn, a Catholic city. He attempted to gain a professorship there 
Title: Theodor Schwann
Content:  Theodor Schwann ( 7 December 1810 – 11 January 1882) was a German physician and physiologist. His most significant contribution to biology is considered to be the extension of cell theory to animals. Other contributions include the discovery of Schwann cells in the peripheral nervous system, the discovery and study of pepsin, the discovery of the organic nature of yeast, and the invention of the term metabolism.
Title: Theodor Schwann
Content:  as a "unifying principle of organic development", related to the philosophical, religious, and political ideas of various proponents including Schwann. In 1848, Schwann's compatriot Antoine Frédéric Spring convinced him to transfer to the University of Liège, also in Belgium. At Liège, Schwann continued to follow the latest advances in anatomy and physiology but did not himself make major new discoveries. He became something of an inventor. One of his projects was a portable respirator, designed as a closed system to support human life in environments where the surroundings cannot be breathed. By 1858 he was serving as professor of physiology, general anatomy and embryology. In 1863, the American Philosophical Society elected him an international 
Title: Eduard Schwartz
Content:  Eduard Schwartz (22 August 1858 – 13 February 1940) was a German classical philologist. Born in Kiel, he studied under Hermann Sauppe in Göttingen, under Hermann Usener and Franz Bücheler in Bonn, under Theodor Mommsen in Berlin and under Ulrich von Wilamowitz-Moellendorff in Greifswald. In 1880 he obtained his doctorate from the University of Bonn. In 1884 he became a lecturer in Bonn, afterwards being appointed professor of classical philology at the University of Rostock (1887). This was followed by professorships at the Universities of Giessen (1893), Strasbourg (1897), Göttingen (1902) and Freiburg (1909). In 1914 he returned to Strasbourg, where he served as university rector in 1915/16. In 1919 he was a successor to Otto 
Title: Theodor Schwann
Content:  As of 1872, he ceased to teach general anatomy, and as of 1877, embryology. He retired fully in 1879. Schwann was deeply respected by his peers. In 1878, a festival was held to celebrate his years of teaching and his many contributions. He was presented with a unique gift: a book containing 263 autographed photographic portraits of scientists from various countries, each of them sent by the scientist to be part of the gift for Schwann. The volume was dedicated "To the creator of the cell theory, the contemporary biologists." Three years after retiring, Schwann died in Cologne, on 11 January 1882. He was buried in the family tomb in Cologne's Melaten Cemetery. 
Title: Theodor Schwann
Content:  an apparatus that enabled him to pump the gases oxygen and hydrogen out of the incubation chamber at specific times. This enabled him to establish the critical period in which the eggs needed oxygen. Schwann passed the state examination to practice medicine in the summer of 1834, but he chose to continue to work with Müller, doing research rather than practicing medicine. He could afford to do so, at least in the short term, because of a family inheritance. His salary as an assistant was only 120 taler. For the next five years, Schwann would pay the other three-quarters of his expenses out of his inheritance. As a long-term strategy, it was not sustainable.

Question: where did theodor schwann study?
Answer: Humboldt University of Berlin.