# Chordate

| Chordates Temporal range: Ediacaran–Present, (Possible Ediacaran record, 555 Ma) |
| --- |
|  |
| Scientific classification  |
| Kingdom: | Animalia |
| Subkingdom: | Eumetazoa |
| *Clade*: | ParaHoxozoa |
| *Clade*: | Bilateria |
| *Clade*: | Nephrozoa |
| Superphylum: | Deuterostomia |
| Phylum: | Chordata Haeckel, 1874 |
| Subgroups |
| * †Vetulocystidae?
* †Vetulicolia? (Paraphyletic?)
* †*Yunnanozoon*?
* †*Cathaymyrus*
* †*Pikaia*
* Cephalochordata
* Olfactores
	+ Tunicata
	+ Total group Vertebrata/Craniata
		- †*Emmonsaspis*
		- †*Metaspriggina*
		- †*Nuucichthys*
		- †Myllokunmingiidae
		- †Conodonta?
		- Vertebrata

And see text |

**Chordates**, phylum **Chordata**, are a phylum of animals which have a notochord. The group includes vertebrates, with some closely related invertebrates.

## Classification

### Taxonomy

* **Phylum Chordata**
	+ **Tunicata** – (tunicates, formerly Urochordata; 3,000 species)
	+ †**Vetulicolia** (8 Cambrian species of fossil)
	+ **Cephalochordata** – (lancelets, 30 species)
	+ **Vertebrata** (vertebrates – animals with backbones; 57,739 species)
		- **Agnatha** (jawless vertebrates; 100+ species)
			* **Myxini** or Hyperotreti (hagfish; 65 species)
			* **Conodonta** (Conodonts)
			* **Hyperoartia** (Lampreys)
			* †**Cephalaspidomorphi** (Paleozoic jawless fish)
			* †**Pteraspidomorphi** (Paleozoic jawless fish)
		- **Gnathostomata** (jawed vertebrates)
			* †**Placodermi** (Paleozoic armoured forms)
			* **Chondrichthyes** (cartilaginous fish; 300+ species)
			* †**Acanthodii** (Paleozoic "spiny sharks")
			* **Osteichthyes** (bony fishes; 30,000+ species)
				+ **Actinopterygii** (ray-finned fish; about 30,000 species)
				+ **Sarcopterygii** (lobe-finned fish)
			* **Tetrapoda** (four-legged vertebrates; 18,000+ species)

### Phylogeny

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| **Chordata** |

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|  Tunicata  |

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|  |  Appendicularia (formerly Larvacea) |
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|  |  Thaliacea  |
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|  |  Ascidiacea  |
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|  |  Cephalochordata |
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|  Craniata  |

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| --- | --- |
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Myxini |
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|  Vertebrata  |

|  |  |
| --- | --- |
|  |  Conodonta† |
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|  |  Cephalaspidomorphi† |
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|  |  Hyperoartia |
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|  |  Pteraspidomorphi† |
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|  Gnathostomata  |

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|  |  Placodermi† |
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|  |  Chondrichthyes |
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|  Teleostomi  |

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|  |  Acanthodii† |
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|  Osteichthyes  |

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|  |  Actinopterygii |
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|  Sarcopterygii  |

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| --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- |
| void |

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| --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- |
|  Tetrapoda  |

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|  |  Amphibia |
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|  Amniota  |

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|  Synapsida  |

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| void |

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|  |  Mammalia |
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|  Sauropsida  |

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| void |

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|  |  Aves |
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**Note:** Lines show likely evolutionary relationships. Extinct groups are marked with a "†". Extinct animals are ones that have completely died out.

A X-ray tetra is one of the few chordates with a visible backbone
### Origin of chordates

According to a long-standing theory, the origin of chordates may be found in transformed larvae of sea-squirts (tunicates). Adult tunicates are sessile, but their larvae are motile, and have some features found in early vertebrates. The process of paedomorphosis, where juvenile features are retained in the adult, is the proposed mechanism. Genome analysis shows that tunicates are the closest living relatives of vertebrates.

## Other websites

* Chordate node at Tree Of Life Archived 2007-02-24 at the Wayback Machine
* Chordate node at NCBI Taxonomy