# 호저아목

| **호저아목** |
| --- |
|  카피바라 |
| **생물 분류**ℹ️ |
| 계: | 동물계 |
| 문: | 척삭동물문 |
| 강: | 포유강 |
| 목: | 쥐목 |
| 아목: | **호저아목** (Hystricomorpha) Brandt, 1855 |
| **과** |
| * 본문 참조

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호저아목의 대부분의 종에서 나타나는, 확장된 안와하관을 보여주는 카피바라의 두개골. 이런 형태를 천축서형이라고 부른다.
**호저아목** 또는 **바늘두더지아목**(Hystricomorpha)은 역사적으로 다양한 정의가 있다. 넓은 의미에서는 천축서형의 협골궁형 교근을 지닌 뛰는쥐과를 제외한 대부분의 설치류를 가리킨다. 이 아목은 산미치광이하목과 군디과, 비늘꼬리청서과 그리고 뜀토끼과를 포함하고 있다. 분자생물학과 형태학적 연구 결과는 호저아목 내에 비늘꼬리청서과와 뜀토끼과를 포함시키는 것에 의문을 제기하고 있다. 2005년 칼레톤(Carleton)과 무세르(Musser)는 이 2개 과를 별도의 아목인 비늘꼬리청서아목으로 분류하여 취급하고 있다.

## 하위 분류

* 군디상과 (Ctenodactyloidea)
	+ 군디과 (Ctenodactylidae)
	+ † Tammquammyidae
	+ 라오스바위쥐과 (Diatomyidae)
	+ † Yuomyidae
	+ † Chapattimyidae
* Hystricognathiformes
	+ † Tsaganomyidae
	+ 호저하목 또는 산미치광이하목 (Hystricognathi)
		- † "Baluchimyinae"
		- 호저과 (Hystricidae)
		- 피오미스소목 (Phiomorpha)
			* † Myophiomyidae
			* † Diamantomyidae
			* † 피오미스과 (Phiomyidae)
			* † Kenyamyidae
			* 아프리카바위쥐과 (Petromuridae)
			* 사탕수수쥐과 (Thryonomyidae)
			* 두더지쥐과 (Bathyergidae)
			* † Bathyergoididae
		- 천축서소목 (Caviomorpha)
			* 아메리카호저상과 (Erethizontoidea)
				+ 아메리카호저과 (Erethizontidae)
			* 천축서상과 (Cavioidea)
				+ 아구티과 (Dasyproctidae)
				+ 파카과 (Cuniculidae)
				+ † Eocardiidae
				+ 파카라나과 (Dinomyidae)
				+ † Cephalomyidae
				+ 천축서과 (Caviidae)
			* 데구상과 (Octodontoidea)
				+ 데구과 (Octodontidae)
				+ 투코투코과 (Ctenomyidae)
				+ 아메리카가시쥐과 (Echimyidae)
				+ 후티아과 (Capromyidae)
				+ † Heptaxodontidae
			* 친칠라상과 (Chinchilloidea)
				+ 친칠라과 (Chinchillidae)
				+ † Neoepiblemidae
				+ 친칠라쥐과(Abrocomidae)

## 계통 분류

다음은 쥐목의 계통 분류이다.

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| --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- |
| 쥐목 |

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| 다람쥐아목 |

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겨울잠쥐과 |
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| 다람쥐상과 |

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산비버과 |
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다람쥐과 |
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호저아목 |
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| --- | --- | --- | --- | --- | --- | --- | --- |
| 비늘꼬리청서아목 |

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뜀토끼과 |
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비늘꼬리청서과 |
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쥐아목 |
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| 비버아목 |

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비버과 |
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| 주머니생쥐상과 |

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흙파는쥐과 |
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주머니생쥐과 |
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다음은 호저아목의 계통 분류이다.

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| --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- |
| 호저아목 |

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라오스바위쥐과 |
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군디과 |
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| --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- |
| 호저하목 |

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호저과 |
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| 피오미스소목 |

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사탕수수쥐과 |
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아프리카바위쥐과 |
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뻐드렁니쥐과 |
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| 천축서소목 |

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아메리카호저과 |
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파카과 |
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천축서과 |
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아구티과 |
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파카라나과 |
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친칠라과 |
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| 데구상과 |

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친칠라쥐과 |
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후티아과 |
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가시쥐과 |
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투코투코과 |
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데구과 |
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* Bergsten, J. 2005. A review of long-branch attraction. Cladistics, 21:163-193.
* Cao, Y., Adachi, J., Yano, T. and Hasegawa, M. 1994. Phylogenetic place of guinea pigs: No support of the rodent-polyphyly hypothesis from maximum-likelihood analyses of multiple protein sequences. Molecular Biology and Evolution, 11: 593-604.
* Carleton, M. D. and G. G. Musser. 2005. Order Rodentia. Pp745–752 in Mammal Species of the World A Taxonomic and Geographic Reference (D. E. Wilson and D. M. Reeder eds.). Baltimore, Johns Hopkins University Press.
* D'Erchia, A., Gissi, C., Pesole, G., Saccone, C. and Arnason, U. 1996. The guinea-pig is not a rodent. Nature, 381 (6583): 597-600.
* Flynn, J. J., Wyss, A. R., Croft, D. A., and Charrier, R. 2003. The Tinguiririca Fauna, Chile: biochronology, paleoecology, biogeography, and a new earliest Oligocene South American Land Mammal ‘Age’. Palaeogeography, Palaeoclimatology, Palaeoecology, 195:229-259.
* Graur, D., Hide, W. and Li, W. 1991. Is the guinea-pig a rodent? Nature, 351: 649-652.
* Huchon, D. E. J. P. Douzery. 2001. From the Old World to the New World: A molecular chronicle of the phylogeny and biogeography of hystricognath rodents. Molecular Phylogenetics and Evolution, 20:238-251.
* Kuma, K. and Miyata, T. 1994. Mammalian phylogeny inferred from multiple protein data. Japanese Journal of Genetics, 69 (5): 555-66.
* Landry, S. O. J. 1999. A proposal for a new classification and nomenclature for the glires. Mitt. Mus. Nat. Kd. Berl. Zool. Reihe, 75:283-316.
* Lin, Y-H, et al. 2002. Four new mitochondrial genomes and the increased stability of evolutionary trees of mammals from improved taxon sampling. Molecular Biology and Evolution, 19: 2060-2070.
* Marivaux, L., M. Vianey-Liaud, and J.-J. Jaeger. 2004. High-level phylogeny of early Tertiary rodents: dental evidence. Zoological Journal of the Linnean Society, 142:105-134.
* Marivaux, L. J. L. Welcomme, M. Vianey-Liaud, and J.J. Jaeger. 2002. The role of Asia in the origin and diversification of hystricognathous rodents. Zoologica Scripta, 31:225-239.
* McKenna, Malcolm C., and Bell, Susan K. 1997. *Classification of Mammals Above the Species Level.* Columbia University Press, New York, 631 pp. ISBN 0-231-11013-8
* Reyes, A., Pesole, G. and Saccone, C. 2000. Long-branch attraction phenomenon and the impact of among-site rate variation on rodent phylogeny. Gene, 259 (1-2): 177-87.
* Reyes, A., Gissi, C., Catzeflis, F., Nevo, E. Pesole, G. and Saccone, C. 2004. Congruent mammalian trees from mitochondrial and nuclear genes using Bayesian methods. Molecular Biology and Evolution, 21 (2): 397-403.
* Robinson-Rechavi, M., Ponger, L. and Mouchiroud, D. 2000. Nuclear gene LCAT supports rodent monophyly. Molecular Biology and Evolution, 17: 1410-1412.
* Sullivan, J. and Swofford, D.L. 1997. Are guinea pigs rodents? the importance of adequate models in molecular phylogenetics. Journal of Mammalian Evolution, 4: 77-86.