Darkseid (pronounced "Darkside") is a fictional supervillain appearing in American comic books published by DC Comics.
 Created by writer-artist Jack Kirby, the character made a cameo appearance in "Superman's Pal Jimmy Olsen" #134 (December 1970) before making his full first appearance in "Forever People" #1 (Feb.
 1971).
Steppenwolf is a fictional supervillain appearing in American comic books published by DC Comics.
 The character was created by Jack Kirby and made his first appearance in "New Gods" #7 (February 1972).
 Steppenwolf (which is German for "wolf of the steppe", or coyote) is one of the New Gods, the uncle of the archvillain Darkseid, and a member of Darkseid's Elite.
Amazo is a fictional supervillain appearing in American comic books published by DC Comics.
 The character first appeared in "The Brave and the Bold" #30 (June 1960) and was created by Gardner Fox and Murphy Anderson.
 Debuting in the Silver Age of Comic Books, the character has appeared in comic books and other DC Comics-related products, including animated television series, trading cards and video games.
Harley Quinn (Harleen Frances Quinzel) is a fictional supervillain and occasionally antihero appearing in American comic books published by DC Comics.
 The character was created by Paul Dini and Bruce Timm, and first appeared in "" in September 1992.
 She later appeared in DC Comics' Batman comic books, with the character's first comic book appearance in "The Batman Adventures" #12 (Sept. 1993).
Heinrich Zemo is a fictional supervillain that appeared in various American comic books published by Marvel Comics.
 He was first seen in "The Avengers" #4 (March 1964) in the Silver Age of Comic Books as the first depicted Baron Zemo.
 The character remains dead but his legacy lives on in his son.
Ra's al Ghul (Arabic: رأس الغول‎ ‎ "Raʾs al-Ġūl"; "Ghoul's Head" or "Demon's Head") is a fictional supervillain appearing in American comic books published by DC Comics, commonly as an adversary of the superhero Batman.
 Created by editor Julius Schwartz, writer Dennis O'Neil and artist Neal Adams, the character first appeared in "Batman" #232's "Daughter of the Demon" (June 1971).
 The character is one of Batman's most enduring enemies and belongs to the collective of adversaries that make up Batman's rogues gallery, though given his high status as a supervillain, he has also come into conflict with Superman and other superheroes in the DC Universe.
Bloodshed (real name Wyndell Dichinson) is a fictional supervillain appearing in American comic books published by Marvel Comics.
 It is also the name of a supervillain in Marvel's Razorline imprint, as well as a character in comics from an acquired company, Malibu Comics.
Massacre is a fictional supervillain appearing in American comic books published by Marvel Comics.
 He primarily appears in Spider-Man related publications.
 The character is responsible for the murder of Ashley Kafka, a supporting character in various Spider-Man comic books and adapted media.
Isbisa is a fictional supervillain appearing in American comic books published by Marvel Comics.
 He first appeared in "All Winners Comics" vol.
 1, #19 (Fall 1946), published by Marvel predecessor Timely Comics during the 1940s period fans and historians call the Golden Age of Comic Books.
 He was created by writer Bill Finger and one or more of the five artists who contributed one or more chapters to his 44-page debut story.
Kalibak is a fictional deity and supervillain appearing in American comic books published by DC Comics.
 Kalibak is the eldest son of Darkseid and one of the main enemies of Superman and the Justice League of America.

Question: Fictional supervillain Steppenwolf is the uncle of Darkseid, who appears in comic books published by what company?
Answer: DC Comics