Homer Paine (September 20, 1923 – July 5, 2010) was an American football tackle.
 He played college football at the University of Tulsa for one season and at the University of Oklahoma for three seasons.
 Paine was named to the All-Missouri Valley Conference first team while at Tulsa, and he was twice named to all-conference first teams while at Oklahoma.
 After college, Paine played professional football for one season with the Chicago Hornets of the All-America Football Conference.
 He was selected in the 14th round of the 1946 NFL Draft.
The horse-collar tackle is an American football maneuver in which a defender tackles another player by grabbing the back collar or the back-inside of an opponent's shoulder pads and pulling the ball carrier directly downward in order to pull his feet from underneath him.
 The technique is most closely associated with Pro Bowl safety Roy Williams.
 This kind of tackle was banned from the NFL during the 2005 off-season.
 The rule forbidding it is often referred to in the press as "The Roy Williams Rule".
 The rule, with modifications, was adopted in college football in 2008 and high school football in 2009.
Roy Lee Williams (born August 14, 1980), is a former American college and professional football player who was a safety in the National Football League (NFL) for nine seasons.
 He played college football for the University of Oklahoma, and was recognized as a unanimous All-American.
 He was drafted by the Dallas Cowboys eighth overall in the 2002 NFL Draft, and played professionally for the Cowboys and Cincinnati Bengals.
 He earned five straight Pro Bowl selections from 2003 to 2007.
 Williams is currently a sideline reporter for Oklahoma football games.
Lewis Pate "Bud" McFadin (August 21, 1928 – February 13, 2006) was an American football player.
 He played college football at the University of Texas and was a unanimous selection at the guard position on the 1950 College Football All-America Team.
 He later played professional football in the National Football League (NFL) for the Los Angeles Rams (1952–1956) and in the American Football League (AFL) for the Denver Broncos (1960–1963) and Houston Oilers (1964–1965).
 A versatile player, he played tackle and linebacker on defense, as well as tackle and guard on offense.
 He was a Pro Bowl pick in 1955 and 1956, a "Sporting News" All-AFL defensive tackle in 1960, 1961 and 1962, and an American Football League West Division All-Star in 1963.
Theodore Anthony Rosequist (April 17, 1908 – November 29, 1988) was an American football player.
 Rosequist played college football at John Carroll University and Ohio State University.
 He was selected by the International News Service as a third-team tackle on the 1932 College Football All-America Team.
 He also played professional football as a tackle in the National Football League for the Chicago Bears (1934–1936) and Cleveland Rams (1937).
Rex Reed Boggan (March 27, 1930 - December 8, 1985) was an American football player.
 Boggan was born in Tupelo, Mississippi, and attended the University of Mississippi.
 He played college football at the tackle position for the Ole Miss Rebels football team.
 He was selected by the Associated Press as a first-team player on its 1954 College Football All-America Team.
 He was drafted by the New York Giants in the 20th round of the 1952 NFL Draft and played for the Giants during the 1955 season at the defensive tackle position.
Robert Glen "Hogan" Wharton (December 13, 1935 – October 8, 2008) was an American football player.
 He attended the University of Houston where he played college football at the tackle position for the Houston Cougars football team from 1956 to 1958.
 He was named lineman of the year in the Missouri Valley Conference in 1957, and the following year he was selected by the American Football Coaches Association as a first-team tackle on its 1958 College Football All-America Team.
 Wharton later played professional football in the newly formed American Football League, playing at the guard position for the Houston Oilers during the first four years of the club's existence from 1960 to 1963, including the 1960 Houston Oilers team that won the first AFL championship.
 He was cut by the Oilers in September 1964.
Alvin Lawrence "Moose" Wistert (June 26, 1916 – October 3, 2005) was an American football player.
 A native of Chicago, Illinois, he played college football at the tackle position for Boston University in 1946 and at the University of Michigan from 1947 to 1949.
 He began his collegiate football career at age 30 following 12 years of working in a factory and serving in the United States Marine Corps during World War II.
 He played at the defensive tackle position for the undefeated 1947 and 1948 Michigan Wolverines football teams, both of which finished the season ranked No. 1 in the final Associated Press poll.
 He also holds the distinction of being the oldest college football player ever selected as a College Football All-American, having been selected to the 1948 College Football All-America Team at age 32 and the 1949 Team at age 33.
Harold Dwayne Akin (born January 11, 1945 in McAlester, Oklahoma) was an American football player.
 An offensive tackle, he played college football at Oklahoma State University, and played professionally in the American Football League for the San Diego Chargers in 1967 and 1968.
 He later went on to found The Budget Floor Store in Oklahoma City, which now has 4 locations across Oklahoma.
Emerson William "Spike" Nelson (April 2, 1906 – October 20, 1998) was an American football player and coach.
 He served as the head football coach at Mississippi State College, now Mississippi State University, in 1938 and at Yale University in 1941, compiling a career college football record of 5–13.
 He played college football as a tackle at the University of Iowa and was selected as a first-team tackle by the "New York Sun" on its 1926 College Football All-America Team.
 He was also selected as a second-team All-American by the Associated Press and Central Press.

Question: The horse-collar tackle is closely associated with what safety who played college football for the University of Oklahoma?
Answer: Roy Lee Williams