Colydiinae

Colydiinae is a subfamily of beetles, commonly known as cylindrical bark beetles. They have been treated historically as a family Colydiidae, but have been moved into the Zopheridae, where they constitute the bulk of the diversity of the newly expanded family, with about 140 genera worldwide. They are diverse for example in the Australian region, from where about 35 genera are known; in Europe, though, only 20 genera are found and many of these only with few species.
Little is known about the biology of these beetles. Most feed on fungi, others are carnivores and eat small arthropods such as bark beetles.
Systematics and taxonomy
Up to 9 tribes are accepted by various authors; others, however, synonymize some of these. Formerly, many additional tribes were recognized, but the Synchitini, for example, are today generally held to include a number of these tribes, and are even sometimes merged into the Colydiini.
The tribes are:
Acropini
Adimerini
Colydiini Billberg, 1820
Gempylodini
Nematidiini
Orthocerini Blanchard, 1845 (= Sarrotriini Billberg, 1820 (nom. rej.))
Rhagoderini
Rhopalocerini
Synchitini Erichson, 1845 (sometimes in Colydiini)
Delimitation of the Colydiinae against the other lineages of Zopheridae is usually unproblematic. The only significant case of dispute may be the Pycnomerini, which is a small lineage of Zopheridae incertae sedis and was formerly considered an independent family like the "Colydiidae". That treatment is almost certainly wrong, but whether these beetles should be placed in Zopheridae as an additional subfamily Pycnomerinae, or treated as tribe Pycnomerini – and if the latter, whether they are better included in the Colydiinae or the Zopherinae – is still disputed.
Selected genera
Genera of cylindrical bark beetles include:
Ablabus Broun, 1880
Acolobicus
Acolophoides Slipinski & Lawrence, 1997
Acolophus Sharp, 1885
Acostonotus Slipinski & Lawrence, 1997
Acropis Burmeister, 1840
Afrorthocerus
Allobitoma Broun, 1921
Alluauditoma
Anosyana
Antilissus Sharp, 1879
Ascomma
Asprotera
Asynchita
Atyscus
Aulonium Erichson, 1845
Bhutania
Bitoma Herbst, 1793
Bolcocius Dajoz, 1977
Bulasconotus Ślipiński & Lawrence, 1997
Bupala
Cacotarphius
Caprodes Pascoe, 1863
Catolaemus
Cebia Pascoe, 1863
Cerchanotus Erichson, 1845
Chorasus Sharp, 1882 (including Vitiacus)
Cicablabus Slipinski & Lawrence, 1997
Cicones
Ciconissus Broun, 1893 (including Caanthus)
Colobicones Grouvelle, 1918
Colobicus Latreille, 1807
Colydium Fabricius, 1792
Colydodes
Corticus Germar, 1824
Coxelus Dejean, 1821
Dechomus
Denophloeus
Diodesma Latreille, 1829
Diplagia Reitter, 1882
Diplotoma
Ditomoidea
Emilka
Endeitoma Sharp, 1894
Endocoxelus
Endophloeus Dejean, 1834
Enhypnon Carter, 1919
Epistranodes Slipinski & Lawrence, 1997
Epistranus Sharp, 1878
Erylus
Eucicones
Eudesma
Eulachus
Faecula Slipinski & Lawrence, 1997
Fenerivia
Gempylodes Pascoe 1863
Glenentela Broun, 1893
Glyphocryptus Sharp, [1885]
Helioctamenus Schaufuss, 1882
Heterargus Sharp, 1886 (including Gathocles, Protarphius)
Holopleuridia
Hyberis
Hybonotus Ślipiński & Lawrence, 1997
Hystricones
Isotarphius
Kanantsia
Labrotrichus
Langelandia Aubé, 1843
Lascobitoma Slipinski & Lawrence, 1997
Lasconotus Erichson, 1845
Lascotonus
Lascotrichus
Lastrema Reitter, 1882
Linophloeus
Lobogestoria
Lobomesa Slipinski & Lawrence, 1997
Lyreus Aubé, 1861
Madacones
Madadesia
Madenphloeus
Mamakius
Megataphrus
Microprius Fairmaire, 1868
Microsicus
Mnionychus
Monoedus Horn, 1882 (including Adimerus)
Munaria Reitter, 1882
Namunaria Reitter, 1882 (including Sympanotus)
Nematidium Erichson, 1845
Neotrichus Sharp, 1885
Niphopelta Reitter, 1882
Norix Broun, 1893
Nosodomodes Reitter, 1922
Notocoxelus Ślipiński & Lawrence, 1997
Orthocerus Latreille, 1796 (including Sarrotrium)
Paha
Paratarphius
Paryphus
Pharax
Phloeodalis
Phloeonemus
Phormesa
Phorminx Carter & Zeck, 1937
Phreatus
Priolomopsis
Priolomus
Pristoderus Hope, 1840 (including Dryptops, Enarsus, Recyntus, Sparactus)
Prosteca Wollaston, 1860
Pseudendestes Lawrence, 1980
Pseudocorticus
Pseudotarphius
Rechodes
Rhagodera Mannerheim 1843
Rhopalocerus Redtenbacher, 1842
Rytinotus Broun, 1880
Sallachius
Sassaka
Sechellotoma
Sprecodes
Synagathis Carter & Zeck, 1937
Syncalus Sharp, 1876 (including Acosmetus)
Synchita Hellwig, 1792
Tarphiablabus Ślipiński & Lawrence, 1997
Tarphiomimus Wollaston, 1873
Tarphiosoma
Tarphius Erichson, 1845
Tentablabus Slipinski & Lawrence, 1997
Todima Grouvelle, 1893
Todimopsis Ślipiński & Lawrence, 1997
Trachypholis Erichson, 1845
Trigonophloeus
Xylolaemus Redtenbacher, 1858
Zanclea
The genera Pycnomerodes, Pycnomerus and Rhizonium are sometimes included in the Colydiinae too. Other authors consider the latter incertae sedis among the Tenebrionoidea; for the former two, see above. The species Xylolaemus sakhnovi was described in 2014 from a fossil preserved in Baltic amber, which dates to the Middle Eocene. This was the first species of Xylolaemus described from the fossil record. The extinct genus Paleoendeitoma, belonging to the extant tribe Synchitini is known from the Cenomanian aged Burmese amber of Myanmar.
See also
List of beetle species recorded in Britain – superfamily Tenebrionoidea
References
External links
Atlas of cylindrical bark beetles (Colydiidae) of Russia - Zoological Institute of Russian Academy of Sciences
Zopheridae
Articles containing video clips