Pholcus

Pholcus is a genus of spiders in the family Pholcidae, with 375 described species as of January 2023.
It includes the cellar spider P. phalangioides, often called the "daddy longlegs". This may cause confusion because the name "daddy longlegs" is also applied to two other unrelated arthropods: the harvestman and the crane fly.
Description
Pholcus, like Pholcidae in general, have extremely long and thin legs. The genus can be distinguished from other pholcid genera by its large size (body length >4 mm), eight eyes, evenly domed prosoma (lacking a median furrow or pit) and cylindrical opisthosoma (longer than it is high).
Habitat
In the wild, Pholcus live in environments such as caves, under rocks, forest shrubs and deep limestone cracks. Synanthropic species such as P. phalangioides live in and around buildings and other disturbed habitats.
Species
Pholcus abstrusus Yao & Li, 2012 — China
Pholcus acutulus Paik, 1978 — Korea
Pholcus aduncus Yao & Li, 2012 — China
Pholcus afghanus Senglet, 2008 — Afghanistan
Pholcus agilis Yao & Li, 2012 — China
Pholcus alagarkoil (Huber, 2011) — India
Pholcus alloctospilus Zhu & Gong, 1991 — China
Pholcus alpinus Yao & Li, 2012 — China
Pholcus alticeps Spassky, 1932 — Poland to Russia, Iran
Pholcus amani Huber, 2011 — Tanzania
Pholcus anachoreta Dimitrov & Ribera, 2006 — Canary Islands
Pholcus ancoralis L. Koch, 1865 — Ryukyu Islands to Hawaii, New Caledonia, Marquesas Islands, Rapa
Pholcus anlong Chen, Zhang & Zhu, 2011 — China
Pholcus arayat Huber, 2011 — Philippines
Pholcus arcuatilis Yao & Li, 2013 — Laos
Pholcus arkit Huber, 2011 — Central Asia
Pholcus armeniacus Senglet, 1974 — Iran
Pholcus arsacius Senglet, 2008 — Iran
Pholcus attuleh Huber, 2011 — Cameroon
Pholcus auricularis Zhang, Zhang & Liu, 2016 — China
Pholcus babao Tong & Li, 2010 — China
Pholcus baguio Huber, 2016 — Philippines
Pholcus bailongensis Yao & Li, 2012 — China
Pholcus bajia (Lu, Yao & He, 2022) — China
Pholcus baka Huber, 2011 — West, Central Africa
Pholcus bakweri Huber, 2011 — Bioko, Cameroon
Pholcus baldiosensis Wunderlich, 1992 — Canary Islands
Pholcus bamboutos Huber, 2011 — Cameroon
Pholcus bangfai Huber, 2011 — Laos
Pholcus bantouensis Yao & Li, 2012 — China
Pholcus bat Lan & Li, 2021 — China
Pholcus batepa Huber, 2011 — Sao Tome
Pholcus beijingensis Zhu & Song, 1999 — China
Pholcus berlandi Millot, 1941 — Senegal
Pholcus bessus Zhu & Gong, 1991 — China
Pholcus bicornutus Simon, 1892 — Philippines
Pholcus bidentatus Zhu et al., 2005 — China, Laos
Pholcus bifidus Yao, Pham & Li, 2015 — Vietnam
Pholcus bikilai Huber, 2011 — Ethiopia
Pholcus bimbache Dimitrov & Ribera, 2006 — Canary Islands
Pholcus bing Yao & Li, 2012 — China
Pholcus bolikhamsai Huber, 2011 — Laos
Pholcus bourgini Millot, 1941 — Guinea
Pholcus brevis Yao & Li, 2012 — China
Pholcus bulacanensis Yao & Li, 2017 — Philippines (Luzon)
Pholcus caecus Yao, Pham & Li, 2015 — Vietnam
Pholcus calcar Wunderlich, 1987 — Canary Islands
Pholcus calligaster Thorell, 1895 — Myanmar, Nepal
Pholcus camba Huber, 2011 — Sulawesi
Pholcus caspius Senglet, 2008 — Iran
Pholcus ceheng Chen, Zhang & Zhu, 2011 — China
Pholcus cenranaensis Yao & Li, 2016 — Indonesia (Sulawesi)
Pholcus ceylonicus O. Pickard-Cambridge, 1869 — Sri Lanka, possibly Malaysia
Pholcus chang Yao & Li, 2012 — China
Pholcus changchi Yao, Li & Lu, 2022 — China
Pholcus chappuisi Fage, 1936 — Kenya
Pholcus chattoni Millot, 1941 — Guinea, Ivory Coast
Pholcus cheaha Huber, 2011 — USA
Pholcus chengde Yao, Li & Lu, 2022 — China
Pholcus cheongogensis Kim & Ye, 2015 — Korea
Pholcus chevronus Yin, Xu & Bao, 2012 — China
Pholcus chiakensis Seo, 2014 — Korea
Pholcus chicheng Tong & Li, 2010 — China
Pholcus chilgapsanensis J. G. Lee & J. H. Lee, 2021 — Korea
Pholcus choctaw Huber, 2011 — USA
Pholcus chuncheonensis J. G. Lee, Choi & S. K. Kim, 2021 — Korea
Pholcus circularis Kraus, 1960 — Sao Tome
Pholcus clavatus Schenkel, 1936 — China
Pholcus clavimaculatus Zhu & Song, 1999 — China
Pholcus cophenius Senglet, 2008 — Afghanistan
Pholcus corcho Wunderlich, 1987 — Canary Islands
Pholcus corniger Dimitrov & Ribera, 2006 — Canary Islands
Pholcus crassipalpis Spassky, 1937 — Bulgaria, Ukraine, Russia (Europe, Caucasus), Turkey, Kazakhstan
Pholcus crassus Paik, 1978 — Korea
Pholcus creticus Senglet, 1971 — Crete
Pholcus crypticolenoides Kim, Lee & Lee, 2015 — Korea
Pholcus crypticolens Bösenberg & Strand, 1906 — Japan
Pholcus cuneatus Yao & Li, 2012 — China
Pholcus curvus Zhang, Zhang & Liu, 2016 — China
Pholcus dade Huber, 2011 — USA
Pholcus dali Zhang & Zhu, 2009 — China
Pholcus datan Tong & Li, 2010 — China
Pholcus datong (Yao, Li & Lu, 2022) — China
Pholcus debilis (Thorell, 1899) — Bioko, Cameroon
Pholcus decorus Yao & Li, 2012 — China
Pholcus dentatus Wunderlich, 1995 — Madeira
Pholcus deunggolensis Kim & Kim, 2016 — Korea
Pholcus dieban Yao & Li, 2012 — China
Pholcus difengensis Yao & Li, 2016 — China
Pholcus dixie Huber, 2011 — USA
Pholcus djelalabad Senglet, 2008 — Afghanistan, India
Pholcus dongxue Yao & Li, 2017 — Thailand
Pholcus doucki Huber, 2011 — Guinea
Pholcus duan Yao & Li, 2017 — Thailand
Pholcus dungara Huber, 2001 — Queensland
Pholcus edentatus Campos & Wunderlich, 1995 — Canary Islands
Pholcus elymaeus Senglet, 2008 — Iran
Pholcus exilis Tong & Li, 2010 — China
Pholcus extumidus Paik, 1978 — Korea, Japan
Pholcus fagei (Kratochvíl, 1940) — Kenya
Pholcus faveauxi (Lawrence, 1967) — Congo
Pholcus fengcheng Zhang & Zhu, 2009 — China
Pholcus fengning (Yao, Li & Lu, 2022) — China
Pholcus foliaceus Peng & Zhang, 2013 — China
Pholcus fragillimus Strand, 1907 — Sri Lanka, India to Japan
Pholcus fuerteventurensis Wunderlich, 1992 — Canary Islands, Morocco
Pholcus gaizhou Yao & Li, 2021 — China
Pholcus gajiensis Seo, 2014 — Korea
Pholcus ganziensis Yao & Li, 2012 — China
Pholcus gaoi Song & Ren, 1994 — China
Pholcus genuiformis Wunderlich, 1995 — Algeria
Pholcus gomerae Wunderlich, 1980 — Canary Islands
Pholcus gonggarensis Yao & Li, 2016 — China
Pholcus gosuensis Kim & Lee, 2004 — Korea
Pholcus gracillimus Thorell, 1890 — Malaysia, Singapore, Sumatra, Java
Pholcus guadarfia Dimitrov & Ribera, 2007 — Canary Islands
Pholcus guangling (Yao, Li & Lu, 2022) — China
Pholcus guani Song & Ren, 1994 — China
Pholcus guanshui Yao & Li, 2021 — China
Pholcus gui Zhu & Song, 1999 — China
Pholcus guineensis Millot, 1941 — Guinea, Sierra Leone
Pholcus hamaensis Yao & Li, 2016 — China
Pholcus hamatus Tong & Ji, 2010 — China
Pholcus hamuchal Yao & Li, 2020 — Pakistan
Pholcus harveyi Zhang & Zhu, 2009 — China
Pholcus helenae Wunderlich, 1987 — Canary Islands
Pholcus henanensis Zhu & Mao, 1983 — China
Pholcus hieroglyphicus Pavesi, 1883 — Eritrea
Pholcus higoensis Irie & Ono, 2008 — Japan
Pholcus hinsonensis Yao & Li, 2016 — Thailand
Pholcus hochiminhi Yao, Pham & Li, 2015 — Vietnam
Pholcus hoyo Huber, 2011 — Congo
Pholcus huailai Yao, Li & Lu, 2022 — China
Pholcus huapingensis Yao & Li, 2012 — China
Pholcus huberi Zhang & Zhu, 2009 — China
Pholcus hunyuan (Yao, Li & Lu, 2022) — China
Pholcus huoxiaerensis Yao & Li, 2016 — China
Pholcus hyrcanus Senglet, 1974 — Iran
Pholcus hytaspus Senglet, 2008 — Iran
Pholcus imbricatus Yao & Li, 2012 — China
Pholcus incheonensis J. G. Lee & J. H. Lee, 2021 — Korea
Pholcus intricatus Dimitrov & Ribera, 2003 — Canary Islands
Pholcus jaegeri Huber, 2011 — Laos
Pholcus jiaotu Yao & Li, 2012 — China
Pholcus jiguanshan Yao & Li, 2021 — China
Pholcus jindongensis Seo, 2018 — Korea
Pholcus jingnan Yao & Li, 2020 — China
Pholcus jingyangensis Yao & Li, 2016 — China
Pholcus jinniu Tong & Li, 2010 — China
Pholcus jinwum Huber, 2001 — Queensland
Pholcus jiulong Tong & Li, 2010 — China
Pholcus jiuwei Tong & Ji, 2010 — China China
Pholcus jixianensis Zhu & Yu, 1983 — China
Pholcus joreongensis Seo, 2004 — Korea
Pholcus jusahi Huber, 2011 — USA
Pholcus juwangensis Seo, 2014 — Korea
Pholcus kaebyaiensis Yao & Li, 2016 — Thailand
Pholcus kakum Huber, 2009 — Ghana, Ivory Coast, Guinea, Congo
Pholcus kalam Yao & Li, 2020 — Pakistan
Pholcus kamkaly Huber, 2011 — Kazakhstan
Pholcus kandahar Senglet, 2008 — Afghanistan
Pholcus kangding Zhang & Zhu, 2009 — China
Pholcus kapuri Tikader, 1977 — Andaman Islands
Pholcus karawari Huber, 2011 — New Guinea
Pholcus kawit Huber, 2016 — Philippines
Pholcus kihansi Huber, 2011 — Tanzania
Pholcus kimi Song & Zhu, 1994 — China, Laos
Pholcus kindia Huber, 2011 — Guinea
Pholcus kingi Huber, 2011 — USA
Pholcus knoeseli Wunderlich, 1992 — Canary Islands
Pholcus koah Huber, 2001 — Queensland
Pholcus koasati Huber, 2011 — USA
Pholcus kohi Huber, 2011 — Malaysia, Singapore, Sumatra
Pholcus krachensis Yao & Li, 2016 — Thailand
Pholcus kribi Huber, 2011 — Cameroon
Pholcus kuaile (Yao, Li & Lu, 2022) — China
Pholcus kui Yao & Li, 2012 — China
Pholcus kunming Zhang & Zhu, 2009 — China
Pholcus kwamgumi Huber, 2011 — Kenya, Tanzania
Pholcus kwanaksanensis Namkung & Kim, 1990 — Korea
Pholcus kwangkyosanensis Kim & Park, 2009 — Korea
Pholcus kyondo Huber, 2011 — Congo
Pholcus laksao Huber, 2011 — Laos
Pholcus lamperti Strand, 1907 — Tanzania, possibly Zanzibar
Pholcus langensis Yao & Li, 2016 — China
Pholcus lanieri Huber, 2011 — USA
Pholcus leruthi Lessert, 1935 — Congo, East Africa
Pholcus lexuancanhi Yao, Pham & Li, 2012 — Vietnam
Pholcus lijiangensis Yao & Li, 2012 — China
Pholcus lilangai Huber, 2011 — Tanzania
Pholcus lingguanensis Yao & Li, 2016 — China
Pholcus lingulatus Gao, Gao & Zhu, 2002 — China
Pholcus linzhou Zhang & Zhang, 2000 — China
Pholcus liui Yao & Li, 2012 — China
Pholcus liutu Yao & Li, 2012 — China
Pholcus longlin Yao & Li, 2020 — China
Pholcus longus Yao & Li, 2016 — China
Pholcus longxigu Yao & Li, 2021 — China
Pholcus lualaba Huber, 2011 — Congo
Pholcus luanping (Yao & Li, 2022) — China
Pholcus luding Tong & Li, 2010 — China
Pholcus luki Huber, 2011 — Congo
Pholcus luoquanbei Yao & Li, 2021 — China
Pholcus lupanga Huber, 2011 — Tanzania
Pholcus luya Peng & Zhang, 2013 — China
Pholcus madeirensis Wunderlich, 1987 — Madeira
Pholcus magnus Wunderlich, 1987 — Madeira
Pholcus malpaisensis Wunderlich, 1992 — Canary Islands
Pholcus manueli Gertsch, 1937 — Kazakhstan, Turkmenistan, Russia (Far East), China, Korea, Japan. Introduced to USA
Pholcus mao Yao & Li, 2012 — China
Pholcus maronita Brignoli, 1977 — Lebanon
Pholcus mascaensis Wunderlich, 1987 — Canary Islands
Pholcus maxian Lu, Yang & He, 2021 — China
Pholcus mazumbai Huber, 2011 — Tanzania
Pholcus mbuti Huber, 2011 — Congo
Pholcus mecheria Huber, 2011 — Algeria
Pholcus medicus Senglet, 1974 — Iran
Pholcus medog Zhang, Zhu & Song, 2006 — China, India
Pholcus mengla Song & Zhu, 1999 — China
Pholcus mentawir Huber, 2011 — Borneo
Pholcus metta Huber, 2019 — Sri Lanka
Pholcus mianshanensis Zhang & Zhu, 2009 — China
Pholcus mirabilis Yao & Li, 2012 — China
Pholcus mixiaoqii Xu, Zhang & Yao, 2019 — China
Pholcus moca Huber, 2011 — Bioko, Cameroon
Pholcus montanus Paik, 1978 — Korea
Pholcus multidentatus Wunderlich, 1987 — Canary Islands
Pholcus mulu Huber, 2016 — Philippines
Pholcus muralicola Maughan & Fitch, 1976 — USA
Pholcus musensis Yao & Li, 2016 — Thailand
Pholcus nagasakiensis Strand, 1918 — Japan
Pholcus namkhan Huber, 2011 — Laos
Pholcus negara Huber, 2011 — Bali
Pholcus nenjukovi Spassky, 1936 — Central Asia
Pholcus ningan Yao & Li, 2018 — China
Pholcus nkoetye Huber, 2011 — Cameroon
Pholcus nodong Huber, 2011 — Korea
Pholcus obscurus Yao & Li, 2012 — China
Pholcus oculosus Zhang & Zhang, 2000 — China
Pholcus okgye Huber, 2011 — Korea
Pholcus olangapo Huber, 2016 — Philippines
Pholcus opilionoides (Schrank, 1781) — Europe to Azerbaijan
Pholcus ornatus Bösenberg, 1895 — Canary Islands
Pholcus otomi Huber, 2011 — Japan
Pholcus ovatus Yao & Li, 2012 — China
Pholcus pagbilao Huber, 2011 — Philippines
Pholcus pajuensis J. G. Lee, Choi & S. K. Kim, 2021 — Korea
Pholcus palgongensis Seo, 2014 — Korea
Pholcus papilionis Peng & Zhang, 2011 — China
Pholcus papillatus Zhang, Zhang & Liu, 2016 — China
Pholcus paralinzhou Zhang & Zhu, 2009 — China
Pholcus parayichengicus Zhang & Zhu, 2009 — China
Pholcus parkyeonensis Kim & Yoo, 2009 — Korea
Pholcus parthicus Senglet, 2008 — Iran
Pholcus parvus Wunderlich, 1987 — Madeira
Pholcus pennatus Zhang, Zhu & Song, 2005 — China
Pholcus persicus Senglet, 1974 — Iran
Pholcus phalangioides (Fuesslin, 1775) — Cosmopolitan
Pholcus phnombak Lan, Jäger & Li, 2021 — Cambodia
Pholcus phoenixus Zhang & Zhu, 2009 — China
Pholcus phungiformes Oliger, 1983 — Russia
Pholcus piagolensis Seo, 2018 — Korea
Pholcus ping Yao & Li, 2017 — Vietnam
Pholcus pocheonensis J. G. Lee, Choi & S. K. Kim, 2021 — Korea
Pholcus pojeonensis Kim & Yoo, 2008 — Korea
Pholcus ponticus Thorell, 1875 — Romania, Bulgaria to China
Pholcus punu Huber, 2014 — Gabon
Pholcus puranappui Huber, 2019 — Sri Lanka
Pholcus pyeongchangensis Seo, 2018 — Korea
Pholcus qingchengensis Gao, Gao & Zhu, 2002 — China
Pholcus qingyunensis Yao & Li, 2016 — China
Pholcus quinghaiensis Song & Zhu, 1999 — China
Pholcus rawiriae Huber, 2014 — Gabon
Pholcus reevesi Huber, 2011 — USA
Pholcus roquensis Wunderlich, 1992 — Canary Islands
Pholcus ruteng Huber, 2011 — Flores
Pholcus saaristoi Zhang & Zhu, 2009 — China
Pholcus saidovi Yao & Li, 2017 — Tajikistan
Pholcus sakaew Yao & Li, 2018 — Thailand
Pholcus schawalleri Yao, Li & Jäger, 2014 — Philippines
Pholcus seokmodoensis J. G. Lee & J. H. Lee, 2021 — China
Pholcus seorakensis Seo, 2018 — Korea
Pholcus seoulensis J. G. Lee & J. H. Lee, 2021 — Korea
Pholcus shangrila Zhang & Zhu, 2009 — China
Pholcus shenshi Yao & Li, 2021 — Korea
Pholcus shuangtu Yao & Li, 2012 — China
Pholcus shuguanensis Yao & Li, 2017 — Tajikistan
Pholcus sidorenkoi Dunin, 1994 — Russia, Tajikistan
Pholcus silvai Wunderlich, 1995 — Madeira
Pholcus simbok Huber, 2011 — Korea
Pholcus socheunensis Paik, 1978 — Korea
Pholcus sogdianae Brignoli, 1978 — Central Asia
Pholcus sokkrisanensis Paik, 1978 — Korea
Pholcus songi Zhang & Zhu, 2009 — China
Pholcus songkhonensis Yao & Li, 2016 — Thailand
Pholcus songxian Zhang & Zhu, 2009 — China
Pholcus soukous Huber, 2011 — Congo
Pholcus spasskyi Brignoli, 1978 — Turkey
Pholcus spiliensis Wunderlich, 1995 — Crete
Pholcus spilis Zhu & Gong, 1991 — China
Pholcus steineri Huber, 2011 — Laos
Pholcus strandi Caporiacco, 1941 — Ethiopia
Pholcus sublaksao Yao & Li, 2013 — Laos
Pholcus sublingulatus Zhang & Zhu, 2009 — China
Pholcus suboculosus Peng & Zhang, 2011 — China
Pholcus subwuyiensis Zhang & Zhu, 2009 — China
Pholcus suizhongicus Zhu & Song, 1999 — China
Pholcus sumatraensis Wunderlich, 1995 — Sumatra
Pholcus suraksanensis J. G. Lee & J. H. Lee, 2021 — Korea
Pholcus sveni Wunderlich, 1987 — Canary Islands
Pholcus taarab Huber, 2011 — Tanzania, Malawi
Pholcus taibaiensis Wang & Zhu, 1992 — China
Pholcus taibeli Caporiacco, 1949 — Ethiopia
Pholcus taishan Song & Zhu, 1999 — China
Pholcus taita Huber, 2011 — Kenya
Pholcus tang (Yao & Li, 2012) — China
Pholcus tangyuensis Yao & Li, 2016 — China
Pholcus tenerifensis Wunderlich, 1987 — Canary Islands
Pholcus thakek Huber, 2011 — Laos
Pholcus tianmenshan Yao & Li, 2021 — China
Pholcus tianmuensis Yao & Li, 2016 — China
Pholcus tongi Yao & Li, 2012 — China
Pholcus tongyaoi Wang & Yao, 2020 — China
Pholcus triangulatus Zhang & Zhang, 2000 — China
Pholcus tuoyuan Yao & Li, 2012 — China
Pholcus turcicus Wunderlich, 1980 — Turkey
Pholcus tuyan Yao & Li, 2012 — China
Pholcus twa Huber, 2011 — East Africa
Pholcus uiseongensis Seo, 2018 — Korea
Pholcus umphang Yao & Li, 2018 — Thailand
Pholcus unaksanensis J. G. Lee, Choi & S. K. Kim, 2021 — Korea
Pholcus undatus Yao & Li, 2012 — China
Pholcus uva Huber, 2019 — Sri Lanka
Pholcus varirata Huber, 2011 — New Guinea
Pholcus vatovae Caporiacco, 1940 — East Africa
Pholcus velitchkovskyi Kulczynski, 1913 — Russia, Ukraine, Iran
Pholcus vietnamensis Yao & Li, 2017 — Vietnam
Pholcus viveki Sen et al., 2015 — China
Pholcus wahehe Huber, 2011 — Tanzania
Pholcus wangi Yao & Li, 2012 — China
Pholcus wangjiang Yao & Li, 2021 — China
Pholcus wangtian Tong & Ji, 2010 — China
Pholcus wangxidong Zhang & Zhu, 2009 — China
Pholcus woongil Huber, 2011 — Korea
Pholcus wuling Tong & Li, 2010 — China
Pholcus wuyiensis Zhu & Gong, 1991 — China
Pholcus xianrendong Liu & Tong, 2015 — China
Pholcus xiaotu Yao & Li, 2012 — China
Pholcus xinglong (Yao, Li & Lu, 2022) — China
Pholcus xingqi Yao & Li, 2021 — China
Pholcus xingren Chen, Zhang & Zhu, 2011 — China
Pholcus xingyi Chen, Zhang & Zhu, 2011 — China
Pholcus xinzhou (Yao, Li & Lu, 2022) — China
Pholcus yangi Zhang & Zhu, 2009 — China
Pholcus yanjinensis Yao & Li, 2016 — China
Pholcus yanqing (Yao, Li & Lu, 2022) — China
Pholcus yaoshan Yao & Li, 2021 — China
Pholcus yeoncheonensis m, Lee & Lee, 2015 — Korea
Pholcus yeongwol Huber, 2011 — Korea
Pholcus yi Yao & Li, 2012 — China
Pholcus yichengicus Zhu, Tu & Shi, 1986 — China
Pholcus yongshun Yao & Li, 2018 — China
Pholcus yoshikurai Irie, 1997 — Japan
Pholcus yuantu Yao & Li, 2012 — China
Pholcus yugong Zhang & Zhu, 2009 — China
Pholcus yuhuangshan Yao & Li, 2021 — China
Pholcus yunnanensis Yao & Li, 2012 — China
Pholcus yuxi Yao & Li, 2018 — China
Pholcus zham Zhang, Zhu & Song, 2006 — China, Nepal
Pholcus zhangae Zhang & Zhu, 2009 — China
Pholcus zhaoi Yao, Pham & Li, 2015 — Vietnam
Pholcus zhongdongensis Yao & Li, 2016 — China
Pholcus zhui Yao & Li, 2012 — China
Pholcus zhuolu Zhang & Zhu, 2009 — China
Pholcus zichyi Kulczynski, 1901 — Russia, China, Korea
Identification resources
Pholcus of Europe: https://araneae.nmbe.ch/specieskey/263/Pholcus
Pholcus of North America: https://bugguide.net/node/view/9609
References
Pholcidae
Araneomorphae genera
Cosmopolitan spiders