Sequoiadendron giganteum

Sequoiadendron giganteum, also known as the giant sequoia, giant redwood or Sierra redwood is a coniferous tree, classified in the family Cupressaceae in the subfamily Sequoioideae. Giant sequoia specimens are the most massive trees on Earth. They occur naturally only in groves on the western slopes of the Sierra Nevada mountain range of California.
The giant sequoia is listed as an endangered species by the IUCN, with fewer than 80,000 trees remaining. Since its last assessment as an endangered species in 2011, it was estimated that another 13–19% of the population (or 9,761–13,637 mature trees) were destroyed during the Castle Fire of 2020 and the KNP Complex & Windy Fire in 2021, events attributed to fire suppression and drought. Prescribed burns to reduce available fuel load may be crucial for saving the species.
The Giant sequoia grow to an average height of 50–85 m (164–279 ft) with trunk diameters ranging from 6–8 m (20–26 ft). Record trees have been measured at 94.8 m (311 ft) tall. The specimen known to have the greatest diameter at breast height is the General Grant tree at 8.8 m (28.9 ft). Giant sequoias are among the oldest living organisms on Earth. The oldest known giant sequoia is 3,200–3,266 years old.
Wood from mature giant sequoias is fibrous and brittle; trees would often shatter after they were felled. The wood was unsuitable for construction and instead used for fence posts or match sticks. The giant sequoia is a very popular ornamental tree in many parts of the world. The species is named in honor of the Cherokee Chief Sequoyah.
Etymology
The etymology of the genus name has been presumedinitially in The Yosemite Book by Josiah Whitney in 1868to be in honor of Sequoyah (1767–1843), who was the inventor of the Cherokee syllabary. An etymological study published in 2012 concluded that Austrian Stephen L. Endlicher is actually responsible for the name. A linguist and botanist, Endlicher corresponded with experts in the Cherokee language including Sequoyah, whom he admired. He also realized that coincidentally the genus could be described in Latin as sequi (meaning to follow) because the number of seeds per cone in the newly classified genus aligned in mathematical sequence with the other four genera in the suborder. Endlicher thus coined the name "Sequoia" as both a description of the tree's genus and an honor to the indigenous man he admired.
Description
Giant sequoia specimens are the most massive individual trees in the world. They grow to an average height of with trunk diameters ranging from . Record trees have been measured at tall. Trunk diameters of have been claimed via research figures taken out of context. The specimen known to have the greatest diameter at breast height is the General Grant tree at . Between 2014 and 2016, it is claimed that specimens of coast redwood were found to have greater trunk diameters than all known giant sequoias - though this has not been independently verified or affirmed in any academic literature. The trunks of coast redwoods taper at lower heights than those of giant sequoias which have more columnar trunks that maintain larger diameters to greater heights.
The oldest known giant sequoia is 3,200–3,266 years old based on dendrochronology. Giant sequoias are among the oldest living organisms on Earth, and are the verified third longest-lived tree species after the Great Basin bristlecone pine and alerce. Giant sequoia bark is fibrous, furrowed, and may be thick at the base of the columnar trunk. The sap contains tannic acid, which provides significant protection from fire damage. The leaves are evergreen, awl-shaped, long, and arranged spirally on the shoots.
The giant sequoia regenerates by seed. The seed cones are long and mature in 18–20 months, though they typically remain green and closed for as long as 20 years. Each cone has 30–50 spirally arranged scales, with several seeds on each scale, giving an average of 230 seeds per cone. Seeds are dark brown, long, and broad, with a wide, yellow-brown wing along each side. Some seeds shed when the cone scales shrink during hot weather in late summer, but most are liberated by insect damage or when the cone dries from the heat of fire. Young trees start to bear cones after 12 years.
Trees may produce sprouts from their stumps subsequent to injury, until about 20 years old; however, shoots do not form on the stumps of mature trees as they do on coast redwoods. Giant sequoias of all ages may sprout from their boles when branches are lost to fire or breakage.
A large tree may have as many as 11,000 cones. Cone production is greatest in the upper portion of the canopy. A mature giant sequoia disperses an estimated 300–400 thousand seeds annually. The winged seeds may fly as far as from the parent tree.
Lower branches die readily from being shaded, but trees younger than 100 years retain most of their dead branches. Trunks of mature trees in groves are generally free of branches to a height of , but solitary trees retain lower branches.
Distribution
The natural distribution of giant sequoias is restricted to a limited area of the western Sierra Nevada, California. As a paleoendemic species, they occur in scattered groves, with a total of 81 groves (see list of sequoia groves for a full inventory), comprising a total area of only . Nowhere does it grow in pure stands, although in a few small areas, stands do approach a pure condition. The northern two-thirds of its range, from the American River in Placer County southward to the Kings River, has only eight disjunct groves. The remaining southern groves are concentrated between the Kings River and the Deer Creek Grove in southern Tulare County. Groves range in size from with 20,000 mature trees, to small groves with only six living trees. Many are protected in Sequoia and Kings Canyon National Parks and Giant Sequoia National Monument.
The giant sequoia is usually found in a humid climate characterized by dry summers and snowy winters. Most giant sequoia groves are on granitic-based residual and alluvial soils. The elevation of the giant sequoia groves generally ranges from in the north, to to the south. Giant sequoias generally occur on the south-facing sides of northern mountains, and on the northern faces of more southerly slopes.
High levels of reproduction are not necessary to maintain the present population levels. Few groves, however, have sufficient young trees to maintain the present density of mature giant sequoias for the future. The majority of giant sequoia groves are currently undergoing a gradual decline in density since European settlement.
Pre-historic range
While the present day distribution of this species is limited to a small area of California, it was once much more widely distributed in prehistoric times, and was a reasonably common species in North American and Eurasian coniferous forests until its range was greatly reduced by the last ice age. Older fossil specimens reliably identified as giant sequoia have been found in Cretaceous era sediments from a number of sites in North America and Europe, and even as far afield as New Zealand and Australia.
Artificial groves
In 1974, a group of giant sequoias was planted by the United States Forest Service in the San Jacinto Mountains of Southern California in the immediate aftermath of a wildfire that left the landscape barren. The giant sequoias were rediscovered in 2008 by botanist Rudolf Schmid and his daughter Mena Schmidt while hiking on Black Mountain Trail through Hall Canyon. Black Mountain Grove is home to over 150 giant sequoias, some of which stand over tall. This grove is not to be confused with the Black Mountain Grove in the southern Sierra. Nearby Lake Fulmor Grove is home to seven giant sequoias, the largest of which is tall. The two groves are located approximately southeast of the southernmost naturally occurring giant sequoia grove, Deer Creek Grove.
It was later discovered that the United States Forest Service had planted giant sequoias across Southern California. However, the giant sequoias of Black Mountain Grove and nearby Lake Fulmor Grove are the only ones known to be reproducing and propagating free of human intervention. The conditions of the San Jacinto Mountains mimic those of the Sierra Nevada, allowing the trees to naturally propagate throughout the canyon.
Ecology
Giant sequoias are in many ways adapted to forest fires. Their bark is unusually fire resistant, and their cones will normally open immediately after a fire. Giant sequoias are a pioneer species, and are having difficulty reproducing in their original habitat (and very rarely reproduce in cultivation) due to the seeds only being able to grow successfully in full sun and in mineral-rich soils, free from competing vegetation. Although the seeds can germinate in moist needle humus in the spring, these seedlings will die as the duff dries in the summer. They therefore require periodic wildfire to clear competing vegetation and soil humus before successful regeneration can occur. Without fire, shade-loving species will crowd out young sequoia seedlings, and sequoia seeds will not germinate. When fully grown, these trees typically require large amounts of water and are therefore often concentrated near streams. Squirrels, chipmunks, finches and sparrows consume the freshly sprouted seedlings, preventing their growth.
Fires also bring hot air high into the canopy via convection, which in turn dries and opens the cones. The subsequent release of large quantities of seeds coincides with the optimal postfire seedbed conditions. Loose ground ash may also act as a cover to protect the fallen seeds from ultraviolet radiation damage.
Due to fire suppression efforts and livestock grazing during the early and mid-20th century, low-intensity fires no longer occurred naturally in many groves, and still do not occur in some groves today. The suppression of fires leads to ground fuel build-up and the dense growth of fire-sensitive white fir, which increases the risk of more intense fires that can use the firs as ladders to threaten mature giant sequoia crowns. Natural fires may also be important in keeping carpenter ants in check.
In 1970, the National Park Service began controlled burns of its groves to correct these problems. Current policies also allow natural fires to burn. One of these untamed burns severely damaged the second-largest tree in the world, the Washington tree, in September 2003, 45 days after the fire started. This damage made it unable to withstand the snowstorm of January 2005, leading to the collapse of over half the trunk.
In addition to fire, two animal agents also assist giant sequoia seed release. The more significant of the two is a longhorn beetle (Phymatodes nitidus) that lays eggs on the cones, into which the larvae then bore holes. Reduction of the vascular water supply to the cone scales allows the cones to dry and open for the seeds to fall. Cones damaged by the beetles during the summer will slowly open over the next several months. Some research indicates many cones, particularly higher in the crowns, may need to be partially dried by beetle damage before fire can fully open them. The other agent is the Douglas squirrel (Tamiasciurus douglasi) that gnaws on the fleshy green scales of younger cones. The squirrels are active year-round, and some seeds are dislodged and dropped as the cone is eaten.
Genome
The genome of the giant sequoia was published in 2020. The size of the giant sequoia genome is 8.125 Gb (8.125 billion base pairs) which were assembled into eleven chromosome-scale scaffolds, the largest to date of any organism.
This is the first genome sequenced in the Cupressaceae family, and it provides insights into disease resistance and survival for this robust species on a genetic basis. The genome was found to contain over 900 complete or partial predicted NLR genes used by plants to prevent the spread of infection by microbial pathogens.
The genome sequence was extracted from a single fertilized seed harvested from a 1,360-year-old tree specimen in Sequoia/Kings Canyon National Park identified as SEGI 21. It was sequenced over a three-year period by researchers at University of California, Davis, Johns Hopkins University, University of Connecticut, and Northern Arizona University and was supported by grants from Save the Redwoods League and the National Institute of Food and Agriculture as part of a species conservation, restoration and management effort.
Discovery and naming
The giant sequoia was well known to Native American tribes living in its area. Native American names for the species include wawona, toos-pung-ish and hea-mi-withic, the latter two in the language of the Tule River Tribe.
The first reference to the giant sequoia by Europeans is in 1833, in the diary of the explorer J. K. Leonard; the reference does not mention any locality, but his route would have taken him through the Calaveras Grove. Leonard's observation was not publicized. The next European to see the species was John M. Wooster, who carved his initials in the bark of the 'Hercules' tree in the Calaveras Grove in 1850; again, this observation received no publicity. Much more publicity was given to the "discovery" by Augustus T. Dowd of the Calaveras Grove in 1852, and this is commonly cited as the species' discovery by non-natives. The tree found by Dowd, christened the 'Discovery Tree', was felled in 1853.
The first scientific naming of the species was by John Lindley in December 1853, who named it Wellingtonia gigantea, without realizing this was an invalid name under the botanical code as the name Wellingtonia had already been used earlier for another unrelated plant (Wellingtonia arnottiana in the family Sabiaceae). The name "Wellingtonia" has persisted in England as a common name. The following year, Joseph Decaisne transferred it to the same genus as the coast redwood, naming it Sequoia gigantea, but again this name was invalid, having been applied earlier (in 1847, by Endlicher) to the coast redwood. The name Washingtonia californica was also applied to it by Winslow in 1854, though this too is invalid, belonging to the palm genus Washingtonia.
In 1907, it was placed by Carl Ernst Otto Kuntze in the otherwise fossil genus Steinhauera, but doubt as to whether the giant sequoia is related to the fossil originally so named makes this name invalid.
The nomenclatural oversights were finally corrected in 1939 by John Theodore Buchholz, who also pointed out the giant sequoia is distinct from the coast redwood at the genus level and coined the name Sequoiadendron giganteum for it.
The etymology of the genus name has been presumedinitially in The Yosemite Book by Josiah Whitney in 1868to be in honor of Sequoyah (1767–1843), who was the inventor of the Cherokee syllabary. An etymological study published in 2012, however, concluded that the name was more likely to have originated from the Latin sequi (meaning to follow) since the number of seeds per cone in the newly classified genus fell in mathematical sequence with the other four genera in the suborder.
John Muir wrote of the species in about 1870:"Do behold the King in his glory, King Sequoia! Behold! Behold! seems all I can say. Some time ago I left all for Sequoia and have been and am at his feet, fasting and praying for light, for is he not the greatest light in the woods, in the world? Where are such columns of sunshine, tangible, accessible, terrestrialized?'
Uses
Wood from mature giant sequoias is highly resistant to decay, but due to being fibrous and brittle, it is generally unsuitable for construction. From the 1880s through the 1920s, logging took place in many groves in spite of marginal commercial returns. The Hume-Bennett Lumber Company was the last to harvest giant sequoia, going out of business in 1924. Due to their weight and brittleness, trees would often shatter when they hit the ground, wasting much of the wood. Loggers attempted to cushion the impact by digging trenches and filling them with branches. Still, as little as 50% of the timber is estimated to have made it from groves to the mill. The wood was used mainly for shingles and fence posts, or even for matchsticks.
Pictures of the once majestic trees broken and abandoned in formerly pristine groves, and the thought of the giants put to such modest use, spurred the public outcry that caused most of the groves to be preserved as protected land. The public can visit an example of 1880s clear-cutting at Big Stump Grove near General Grant Grove. As late as the 1980s, some immature trees were logged in Sequoia National Forest, publicity of which helped lead to the creation of Giant Sequoia National Monument.
The wood from immature trees is less brittle, with recent tests on young plantation-grown trees showing it similar to coast redwood wood in quality. This is resulting in some interest in cultivating giant sequoia as a very high-yielding timber crop tree, both in California and also in parts of western Europe, where it may grow more efficiently than coast redwoods. In the northwest United States, some entrepreneurs have also begun growing giant sequoias for Christmas trees. Besides these attempts at tree farming, the principal economic uses for giant sequoia today are tourism and horticulture.
Threats
Formerly exploited by logging, sequoias are now threatened by both a combination of fire and fire suppression, along with drought. Fire suppression caused by European land management practices promotes the excessive growth of smaller trees such as fir, which can dry out in droughts, and serve as a fuel load for devastating crown fires fueled by strong winds, which can kill large amounts of sequoias. One such fire, the Castle Fire of 2020, is thought to have wiped out almost 10–14% of the species' whole population, or 7500–10,600 mature trees, including potentially the King Arthur Tree, the 10th tallest known individual. 3 to 5% more, or 2,261–3,637 sequoias, are thought to have died during the KNP Complex and Windy Fire in 2021, with a combined total of 13–19%. The most effective remedy for these impacts is heavy prescribed burning, which will need to be multiplied by 30 times current burning practices in order to ensure heathy sequoia groves.
Cultivation
Giant sequoia is a very popular ornamental tree in many areas. It is successfully grown in most of western and southern Europe, the Pacific Northwest of North America north to southwest British Columbia, the southern United States, southeast Australia, New Zealand and central-southern Chile. It is also grown, though less successfully, in parts of eastern North America.
Trees can withstand temperatures of −31 °C (−25 °F) or colder for short periods of time, provided the ground around the roots is insulated with either heavy snow or mulch. Outside its natural range, the foliage can suffer from damaging windburn.
A wide range of horticultural varieties have been selected, especially in Europe, including blue, compact blue, powder blue, hazel smith, pendulumor weepingvarieties, and grafted cultivars.
France
The tallest giant sequoia ever measured outside of the United States is a specimen planted near Ribeauvillé in France in 1856 and measured in 2014 at a height between and at age 158 years.
United Kingdom
The giant sequoia was first brought into cultivation in Britain in 1853 by the horticulturist Patrick Matthew of Perthshire from seeds sent by his botanist son John in California. A much larger shipment of seed collected from the Calaveras Grove by William Lobb, acting for the Veitch Nursery near Exeter, arrived in England in December 1853; seed from this batch was widely distributed throughout Europe.
Growth in Britain is very fast, with the tallest tree, at Benmore in southwest Scotland, reaching in 2014 at age 150 years, and several others from tall; the stoutest is around in girth and in diameter, in Perthshire. The Royal Botanic Gardens at Kew, and in their second campus at Wakehurst, contain multiple large specimens of the species. Biddulph Grange Garden in Staffordshire holds a fine collection of both Sequoiadendron giganteum and Sequoia sempervirens (coast redwood). The General Sherman of California has a volume of ; by way of comparison, the largest giant sequoias in Great Britain have volumes no greater than , one example being the specimen in the New Forest.
Sequoiadendron giganteum has gained the Royal Horticultural Society's Award of Garden Merit.
An avenue of 218 giant sequoias was planted in 1865 near the town of Camberley, Surrey, England. The trees have since been surrounded by modern real estate development.
Italy
Numerous giant sequoia were planted in Italy from 1860 through 1905. Several regions contain specimens that range from in height. The largest tree is in Roccavione, in the Piedmont, with a basal circumference of . One notable tree survived a tall flood wave in 1963 that was caused by a landslide at Vajont Dam. There are numerous giant sequoia in parks and reserves.
Growth rates in some areas of Europe are remarkable. One young tree in Italy reached tall and trunk diameter in 17 years (Mitchell, 1972).
Northern and Central Europe
Growth further northeast in Europe is limited by winter cold. In Denmark, where extreme winters can reach , the largest tree was tall and diameter in 1976 and is bigger today. One in Poland has purportedly survived temperatures down to with heavy snow cover.
Two members of the German Dendrology Society, E. J. Martin and Illa Martin, introduced the giant sequoia into German forestry at the Sequoiafarm Kaldenkirchen in 1952.
Twenty-nine giant sequoias, measuring around in height, grow in Belgrade's municipality of Lazarevac in Serbia.
The oldest Sequoiadendron in the Czech Republic, at , grows in Ratměřice u Votic castle garden.
United States and Canada
Giant sequoias are grown successfully in the Pacific Northwest and southern US, and less successfully in eastern North America. Giant sequoia cultivation is very successful in the Pacific Northwest from western Oregon north to southwest British Columbia, with fast growth rates. In Washington and Oregon, it is common to find giant sequoias that have been successfully planted in both urban and rural areas.
In Seattle, a sequoia stands as a prominent landmark at the entrance to Seattle's downtown retail core. Other large specimens exceeding are located on the University of Washington and Seattle University campuses, in the Evergreen Washelli Memorial Park cemetery, and in the Leschi, Madrona, and Magnolia neighborhoods.
In the northeastern US there has been some limited success in growing the species, but growth is much slower there, and it is prone to Cercospora and Kabatina fungal diseases due to the hot, humid summer climate there. A tree at Blithewold Gardens, in Bristol, Rhode Island, is reported to be tall, reportedly the tallest in the New England states. The tree at the Tyler Arboretum in Delaware County, Pennsylvania, at may be the tallest in the northeast. Specimens also grow in the Arnold Arboretum in Boston, Massachusetts (planted 1972, 18 m tall in 1998), at Longwood Gardens near Wilmington, Delaware, in the New Jersey State Botanical Garden at Skylands in Ringwood State Park, Ringwood, New Jersey, and in the Finger Lakes region of New York. Private plantings of giant sequoias around the Middle Atlantic States are not uncommon, and other publicly accessible specimens can be visited at the U.S. National Arboretum in Washington, D.C. A few trees have been established in Colorado as well. Additionally, numerous sequoias have been planted with success in the state of Michigan.
A cold-tolerant cultivar 'Hazel Smith' selected in about 1960 is proving more successful in the northeastern US. This clone was the sole survivor of several hundred seedlings grown at a nursery in New Jersey. The U.S. National Arboretum has a specimen grown from a cutting in 1970 that can be seen in the Gotelli Conifer Collection.
Australia
The Ballarat Botanical Gardens contain a significant collection, many of them about 150 years old. Jubilee Park and the Hepburn Mineral Springs Reserve in Daylesford, Cook Park in Orange, New South Wales and Carisbrook's Deep Creek park in Victoria both have specimens. Jamieson Township in the Victorian high country has two specimens which were planted in the early 1860s.
In Tasmania, specimens can be seen in private and public gardens, as sequoias were popular in the mid-Victorian era. The Westbury Village Green has mature specimens with more in Deloraine. The Tasmanian Arboretum contains both Sequoiadendron giganteum and Sequoia sempervirens specimens.
The Pialligo Redwood Forest consists of 3,000 surviving redwood specimens, of 122,000 planted, 500 meters east of the Canberra Airport. The forest was laid out by the city's designer Walter Burley Griffin, though the city's arborist, Thomas Charles Weston, advised against it. The National Arboretum Canberra began a grove of Sequoiadendron giganteum in 2008. They also grow in the abandoned arboretum at Mount Banda Banda in New South Wales.
New Zealand
Several impressive specimens of Sequoiadendron giganteum can be found in the South Island of New Zealand. Notable examples include a set of trees in a public park of Picton, as well as robust specimens in the public and botanical parks of Christchurch and Queenstown. There are also several in private gardens in Wanaka. Other locations in Christchurch and nearby include a number of mature trees at the Riccarton Park Racecourse, three large trees on the roadside bordering private properties on Clyde Road, near Wai-Iti Terrace—these are at least 150 years old. The suburb of ‘Redwood’ is named after a 160 years old Giant Redwood tree in the grounds of a local hotel. At St James Church, Harewood, is a protected very large specimen believed to be about 160 years old. A grove of about sixteen Redwood trees of varying ages is in Sheldon Park in the Belfast, New Zealand suburb. Some of these trees are in poor condition because of indifferent care. There is also a very large tree at Rangiora High School, which was planted for Queen Victoria's Golden Jubilee and is thus over 130 years old.
Record trees
Largest by trunk volume
As of 2009, the top ten largest giant sequoias sorted by volume of their trunks are:
The General Sherman tree is estimated to weigh about 2100 tonnes.
The Washington Tree was previously arguably the second largest tree with a volume of (although the upper half of its trunk was hollow, making the calculated volume debatable), but after losing the hollow upper half of its trunk in January 2005 following a fire, it is no longer of great size.
The largest giant sequoia ever recorded was the Father of the Forest from Calaveras Grove, an exceedingly massive tree which fell many centuries ago in the North Grove. Reportedly, the tree was once over 435ft tall, and 110ft in circumference, with a minimum height of 365 feet. Now, what is left of the tree is a popular tourist attraction.
Tallest
Redwood – Redwood Mountain Grove –
tallest outside the United States: specimen near Ribeauvillé, France, measured in 2014 at a height between and at age 158 years.
Oldest
Muir Snag – Converse Basin Grove – more than 3500 years
Greatest girth
Waterfall Tree – Alder Creek Grove – – tree with enormous basal buttress on very steep ground.
Greatest base diameter
Waterfall Tree – Alder Creek Grove – – tree with enormous basal buttress on very steep ground.
Tunnel Tree – Atwell Mill Grove – – tree with a huge flared base that has burned all the way through.
Greatest mean diameter at breast height
General Grant – General Grant Grove –
Largest limb
Arm Tree – Atwell Mill, East Fork Grove – in diameter
Thickest bark
or more
See also
Mother of the Forest
The House (trees)
Largest organisms
Kings Canyon National Park
Yosemite National Park
Calaveras Big Trees State Park
Old growth forest
List of superlative trees
Notes
References
Sources
Listed as Vulnerable (VU A1cd v2.3)
Further reading
External links
ARKive: Sequoiadendron giganteum, arkive.org
Arboretum de Villardebelle – photos of cones & shoots with phenology notes, pinetum.org
Giant Sequoia Fire History in Mariposa Grove, Yosemite National Park, nps.gov
Save-the-Redwoods League, savethewoods.org
"Sequoias of Yosemite National Park" 1949 by James W. McFarland
Redwoodworld.co.uk giant redwoods in the U.K.
Scenes of Wonder and Curiosity in California (1862) – The Mammoth Trees Of Calaveras, yosemite.ca.us
Short radio episode Woody Gospel Letter in which John Muir extols "King Sequoia" from The Life and Letters of John Muir, 1924. California Legacy Project.
Conifers Around the World: Sequoiadendron giganteum – Giant Sequoia.
Giants of Sequoia and Kings Canyon National Park, On YouTube by Sequoia Parks Conservancy
giganteum
Endemic flora of California
Flora of the Sierra Nevada (United States)
Trees of the Southwestern United States
Trees of the United States
Symbols of California
Garden plants of North America
Ornamental trees
Plants described in 1939
Taxa named by John Lindley