Bembidion

Bembidion is the largest genus of beetles in the family Carabidae by number of species. All species are small (less than 7.5 mm) and move very fast. Most of them live close to water. The genus has a biantitropical distribution, meaning they are found in both the Northern and Southern Hemispheres, but not in the tropics. In warmer regions it is substituted by closely related Tachys and other genera.
Taxonomy
There have been many attempts to divide it into smaller genera, most notably by René Jeannel in 1941 and by G.G. Perrault in 1981, but none of them have been generally accepted.
This genus is divided into numerus subgenera, some of which are elevated to full genus rank by various authors; as noted above, however, no universally accepted way of splitting the genus exists yet. Bembidion subgenera include:
Actedium Motschulsky, 1864
Ananotaphus Netolitzky, 1931
Andrewesa Netolitzky, 1931
Antiperyphanes Jeannel, 1962
Antiperyphus Jeannel, 1962
Aptenidium Habu & Ueno, 1955
Apteromimus Wollaston, 1877
Armatocillenus Dupuis, 1912
Asioperyphus Vysoky, 1986
Aureoplataphus Netolitzky, 1943
Australoemphanes Toledano, 2005
Bembidion Latreille, 1802
Bembidionetolitzkya E.Strand, 1929
Bembidromus Toledano, 2000
Blepharoplataphus Netolitzky, 1920
Bracteon Bedel, 1879
Chilioperyphus Jeannel, 1962
Chinocillenus Netolitzky, 1942
Chlorodium Motschulsky, 1864
Cillenus Samouelle, 1819
Cyclolopha Casey, 1918
Desarmatocillenus Netolitzky, 1939
Diplocampa Bedel, 1896
Ecuadion Moret & Toledano, 2002
Emphanes Motschulsky, 1850
Endosomatium Wollaston, 1877
Euperyphus Jeannel, 1941
Eupetedromus Netolitzky, 1911
Furcacampa Netolitzky, 1931
Gnatholymnaeum Sharp, 1903
Gondwanabembidion Toledano, 2005
Hirmoplataphus Lindroth, 1963
Hoquedela Müller-Motzfeld, 1988
Hydriomicrus Casey, 1918
Hydrium LeConte, 1847
Hypsipezum Alluaud, 1917
Jammuphanes J.Schmidt & Marggi, 2014
Jedlickion Toledano, 2008
Josefia Toledano, 2000
Leuchydrium Casey, 1918
Limnaeoperyphus Nakane, 1963
Lindrochthus Maddison, 2012
Liocosmius Casey, 1918
Lymnaeum Stephens, 1828
Melomalus Casey, 1918
Metallina Motschulsky, 1850
Microserrullula Netolitzky, 1921
Microsinocys Toledano, 1998
Necpericompsus Netolitzky, 1935
Neja Motschulsky, 1864
Neobembidion Bousquet, 2006
Neoemphanes Habu, 1978
Nepha Motschulsky, 1864
Nesocidium Sharp, 1903
Nipponobembidion Habu & Baba, 1968
Notaphemphanes Netolitzky, 1920
Notaphocampa Netolitzky, 1914
Notaphus Dejean, 1821
Nothocys Jeannel, 1962
Notholopha Jeannel, 1962
Nothonepha Jeannel, 1962
Notoperyphus Bonniard de Saludo, 1970
Ochthedromus LeConte, 1847
Ocydromus Clairville, 1806
Ocyturanes Müller-Motzfeld, 1986
Odontium LeConte, 1847
Omoperyphus Netolitzky, 1931
Omotaphus Netolitzky, 1914
Pacmophena Jeannel, 1962
Pamirium Netolitzky, 1920
Paraprincidium Netolitzky, 1914
Pekinium Csiki, 1901
Peryphanes Jeannel, 1941
Peryphidium Tschitscherine, 1895
Peryphiolus Jeannel, 1941
Peryphodes Casey, 1918
Peryphophila Netolitzky, 1939
Peryphus Dejean, 1821
Philochthemphanes Netolitzky, 1943
Philochthus Stephens, 1828
Phyla Motschulsky, 1844
Plataphus Motschulsky, 1864
Plocamoperyphus Jeannel, 1962
Politophanes Müller-Motzfeld, 1998
Princidium Motschulsky, 1864
Pseudometallina Netolitzky, 1920
Pseudoperyphus Hatch, 1950
Pseudophilochthus Wollaston, 1877
Pseudosinocys Toledano, 2005
Pseudotrepanes Jeannel, 1962
Sakagutia Ueno, 1954
Semicampa Netolitzky, 1910
Sloanephila Netolitzky, 1931
Taiwanobembidion Habu, 1973
Talanes Motschulsky, 1864
Terminophanes Müller-Motzfeld, 1998
Testediolum Ganglbauer, 1891
Testedium Motschulsky, 1864
Thaumatoperyphus Netolitzky, 1935
Trechonepha Casey, 1918
Trepanedoris Netolitzky, 1918
Trepanes Motschulsky, 1864
Trichoplataphus Netolitzky, 1914
Zeactedium Netolitzky, 1931
Zecillenus Lindroth, 1980
Zemetallina Lindroth, 1976
Zeperyphodes Lindroth, 1976
Zeperyphus Lindroth, 1976
Zeplataphus Lindroth, 1976
† Archaeophilochthus Ortuño & Arillo, 2010
† Eodontium J.Schmidt & Michalik, 2017
References and notes
External links
Tree of Life A list of species