ASIAN MYRMECOLOGY Volume 8, 17 – 48, 2016

ISSN 1985-1944 © Weeyawat Jaitrong, Benoit Guénard, Evan P. Economo,
Nopparat Buddhakala and Seiki Yamane

DOI: 10.20362/am.008019

A checklist of known ant species of Laos
(Hymenoptera: Formicidae)
Weeyawat Jaitrong1, Benoit Guénard2, Evan P. Economo3,
Nopparat Buddhakala4 and Seiki Yamane5*
Thailand Natural History Museum, National Science Museum,Technopolis, Khlong 5, Khlong Luang, Pathum Thani, 12120 Thailand
E-mail: polyrhachis@yahoo.com
2
School of Biological Sciences, The University of Hong Kong,
Pok Fu Lam Road, Hong Kong SAR, China
3
Okinawa Institute of Science and Technology Graduate University,
1919-1 Tancha, Onna, Okinawa 904-0495, Japan
4
Biology Divisions, Faculty of Science and Technology, Rajamangala University of Technology Tanyaburi, Pathum Thani 12120 Thailand
E-mail: nopparat_b@rmutt.ac.th5
Kagoshima University Museum, Korimoto 1-21-30, Kagoshima-shi,
890-0065 Japan
1*

Corresponding author’s email: mayiopa0@gmail.com

ABSTRACT. Laos is one of the most undersampled areas for ant biodiversity.
We begin to address this knowledge gap by presenting the first checklist of
Laotian ants. The list is based on a literature review and on specimens collected from several localities in Laos. In total, 123 species with three additional
subspecies in 47 genera belonging to nine subfamilies are listed, including 62
species recorded for the first time in the country. Comparisons with neighboring
countries suggest that this list is still very incomplete. The provincial distribution of ants within Laos also show that most species recorded are from Vientiane Province, the central part of Laos while the majority of other provinces
have received very little, if any, ant sampling. Many ant specimens collectedfrom Laos are awaiting identification and future collections should provide
many new species records.
Keywords: Formicidae, ants, checklist, new records, Laos
INTRODUCTION
Laos People’s Democratic Republic (Laos) is alandlocked country located in Southeast Asia
which borders China, Vietnam, Myanmar, Cambodia and Thailand. Although ant faunas are now
known to some extent in several areas in Southeast Asia (e.g. China: Guénard & Dunn 2012;
Vietnam: Eguchi et al. 2011; Thailand: Jaitrong& Nabhitabhata 2005; Cambodia: Hosoishi et al.
2013; Borneo: Pfeiffer et al. 2011; Yunnan: Liu et

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al. 2015), the ants of Laos are largely unknown.
A recent global analysis of gaps in our knowledge of ant biodiversity identified Laos as oneof the least known countries in the world (Guénard et al. 2012), thus placing it as a top priority
for ant biodiversity research. Santschi (1920b)
first reported 28 ant species from several localities in this country. Over the following century,
only a few species of ants have been describedand added in the country (e.g. Ward 2001; Dill
2002; Roncin & Deharveng 2003; Bolton 2007;

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Weeyawat Jaitrong, Benoit Guénard, Evan P. Economo, Nopparat Buddhakala & Seiki Yamane

Jaitrong et al. 2011; Yamane & Jaitrong 2011;
Jaitrong & Yamane 2013; Laciny et al. 2015; Hosoishi & Ogata 2016); and currently, only 61 species have been listed.
We sought to increase knowledge of theant fauna in Laos by compiling data on ant geographic distributions from a review of literature,and through new field surveys during 1997-2015.
With this material we present the first preliminary
checklist of the known ant species from the country in order to encourage future research on ants
and biodiversity conservation in the region.
MATERIALS AND METHODS
The checklist was assembled through literature/
data compilation, in addition to new field surveys.
The data compilation included records from the
GABI database, which includes data from over
8670 publications and other existing databases(e.g. Antweb 2015), with species distribution
available for visualization through the website
antmaps.org (Janicki et al. 2016). Separately, information given by other websites available (e.g.
Antwiki) were also added here.
In addition, an extensive collection was
acquired from several localities in Laos which are
now deposited in the Natural History Museum of
National Science Museum, Thailand (THNHM)
and Seiki Yamane’s Collection at Kitakyushu
Museum of Natural History and Human History, Japan (SKYC). Most specimens were collected by Weeyawat Jaitrong and Seiki Yamanefrom two sites within Vientiane Province during
May and June 2010. The first site was in Sivilay
Village (18ᵒ15/N, 102ᵒ27/E, ca. 200 m alt.), Naxaythong District, located approximately 20 km
northwest of Vientiane City. The second site wasin Phang Dang Village (18ᵒ14/N, 103ᵒ01/E, ca.
500 m alt.), Pak Ngum District, approximately
30 km northeast of Vientiane City. Specimens
collected in various parts of the country by Mr.
Haruo Fukuda and Dr. Yukino Ochiai were also
examined. The specimens were identified by W.
Jaitrong and Sk. Yamane using identification
keys created for other Southeast Asian ants (e.g.
Ward 2001; Bolton 2007; Eguchi 2008; Hosoishi & Ogata 2009a; Jaitrong & Yamane 2011;
Jaitrong et al. 2011; Schmidt & Shattuck 2014;

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Hosoishi & Ogata 2016). Furthermore, Dr. Stefan
P. Cover of the Museum of Comparative Zoology, Cambridge, Massachusetts, U.S.A. (MCZC)
kindly offered us records of specimens of somespecies collected by David Pick in Laos that wereidentified and managed by Dr. Cover.
Supposed biological species (not identified as known species) are not included in thelist. Species validity, spelling and authority havebeen checked in Bolton’s Synopsis of the Formicidae and Catalogue of Ants of the World (Bolton2015). The list is arranged alphabetically by subfamily, genus and species. Material examinedand distributions are given for the listed species.
The species previously recorded from Laos aremarked with “*”.
RESULTS AND DISCUSSION
In the present paper, 123 named species and additional three subspecies are listed for 47 genera
belonging to nine subfamilies from Laos (Table
1). Among them, 62 species are recorded for the
first time in the country. The most speciose subfamily is the Myrmicinae with around 37 percent
of all genera and 30 percent of all species, followed by Formicinae, Ponerinae, Dolichoderinae and Dorylinae respectively (see Table 1).
Worldwide, currently 16 subfamilies, 327 generaand 15,050 species are known in Formicidae,
excluding fossil taxa (Bolton, 2015). Thus, todate the record in Laos covers only 0.8 percent
of all known ant species in the world, though thismay in part be a result of the incompleteness ofinvestigation. Our material contains many otherspecies identified only at the genus level; andthus the number of species is likely to increase
once these are identified. The genera collected
for which no species could be identified are:Aphaenogaster, Calyptomyrmex, Hypoponera,
Nylanderia, Paraparatrechina, Philidris, Plagiolepis, Prenolepis, Pseudolasius, Rhopalomastix, Strumigenys and Vollenhovia. As a result, 58
genera have been recorded from Laos for now.
Many other genera widespread in the region(e.g. Acanthomyrmex, Buniapone, Chronoxenus, Cryptopone, Dilobocondyla, Discothyrea,
Emeryopone, Gauromyrmex, Gesomyrmex,
Kartidris, Lepisiota, Leptanilla, Lordomyrma,

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A Checklist of known ant species of Laos (Hymenoptera: Formicidae)
Table 1. Numbers and percentages of named species and genera recorded for each subfamily from Laos.
Subfamilies

No. of genera%

No. of species%

Amblyoponinae

1

2.2

1

0.8

Dolichoderinae

4

8.7

13

10.6

Dorylinae

3

6.5

10

8.1

Ectatomminae

1

2.2

2

1.6

Formicinae

7

15.2

33

26.8

Myrmicinae

17

37.0

37

30.0

Ponerinae

11

23.9

20

16.3

Proceratiinae

1

2.2

1

0.8

Pseudomyrmecinae

1

2.2

6

4.9

Total

46

100

123

100

Fig. 1. Known species diversity in the eighteen Laotian provinces: 1) Phongsali, 2) Luang Namtha, 3) Bokeo, 4)
Oudomxay, 5) Luang Prabang, 6) Houaphan, 7) Sayabouly, 8) Vientiane capital & province, 9) Xaisomboun, 10)
Xieng Khouang, 11) Bolikgamsai, 12) Khammouane, 13) Savannakhet, 14) Salavan, 15) Sekong, 16) Attapeu,
17) Champasak.

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Mesoponera, Myopias, Myrmecina, Myrmicaria,
Myrmoteras, Mystrium, Parvaponera, Ponera,
Proatta, Proceratium, Simopone, Stigmatomma,
Syllophopsis, Vombisidris) are expected in Laos.
The predicted final figure for Laos is between 64
and 78 genera, based on an earlier study by Guénard and collaborators. (2012).
The most speciose and commonly encountered genera are Polyrhachis (19 species),
Camponotus (8), Aenictus (7), Crematogaster(7), Tetramorium (6), Dolichoderus (5), Technomyrmex (5), Carebara (5), Pheidole (5) andTetraponera (5). Together, the ten most diverse
genera include 59 percent of the known species
from Laos. Twenty-seven genera (59%) are so
far represented in Laos by a single species. There
are some genera with large numbers of Lao species (according to the specimens deposited in
THNHM and SKYC), but most of these remain
unknown, as they have not been studied in detailand identified yet, and therefore are not represented in this paper. For instance, Camponotus,
Pheidole, Tetramorium and Monomorium all
have several unidentified species.
Santschi (1920b) recorded Odontoponera transversa (F. Smith, 1857) from Lad-Homand Luang Prabang, Laos. According to Yamane(2009), this species is confined to Sundaland, i.e.,
Borneo, Java, Sumatra, Singapore and Malay
Peninsula (including southern Thailand). Odontoponera denticulata has a wider distribution,
ranging from the Philippines, through Sundaland
to continental Southeast Asia such as Thailand,
Myanmar, Vietnam, Laos, S. China, Bangladesh,
India and Pakistan (Eguchi et al. 2005; Hannan
2007; Jaitrong & Nabhitabhata 2005; Yamane etal. 2003). Thus Lao specimens (Lat-Hom and Luang Prabang) cited as O. transversa in Santschi(1920b) should be identified as O. denticulata.
Over 80 percent of the known ant species from Laos are common in Southeast Asia.
There are a few species, e.g. Carebara petulens(Santschi 1920), Dolichoderus brevis Santschi,
1920, Polyrhachis emmae Santschi, 1920 and
Leptogenys khammouanensis Roncin et Deharveng, 2003, that thus far can be considered as
endemic to Laos. Some species which to datehave been recorded only from regions adjacentto Laos might occur in the country, but they areomitted from the list because their presence has

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not been confirmed. For example, the common
species, e.g. Monomorium chinense Santschi,
1925, Ectomomyrmex astuta (F. Smith, 1858) (=Pachycondyla astuta), and Pheidole pieli Santschi, 1925, are recorded from China, Vietnam andThailand (Eguchi et al. 2005; Jaitrong & Nabhitabhata 2005; Guénard & Dunn 2012), but notfrom Laos. However, it is likely that this distribution gap exists due to incompleteness of samplingin the country.
In the ant collections of THNHM and
SKYC, we recognized about 70 unidentified species from Laos, some of which seem to be new
to science; a new species of Calyptomyrmex will
be described soon (Jaitrong & Yamane, in prep.).
Specimens of some genera, e.g., Nylanderia, Philidris, Paraparatrechina, Hypoponera, Vollenhovia and Strumigenys, have not yet been identified
to species.
The under-sampling in the country
is particularly striking at the provincial level.
Among the 18 provinces of Laos, seven regions
have no ant records and six more have fewer than
10 records. The “best” sampled province, Vientiane, has only 91 recorded species, while the second best-sampled province, Luang Prabang, hasonly 34 (Fig. 1). These results show that the entire
country is highly undersampled with virtually nosampling efforts for the southern and the northwestern regions in particular. Yet, in light of theant species richness found in neighboring regions(e.g. Vietnam: Eguchi et al. 2011, 2014; Yunnan
province in China: Liu et al. 2015), many more
species than mentioned in this paper (123+70)
should be recognized in Laos in the future.
We carefully studied all the ant specimens collected by us from Laos. However, we
present here only a first checklist of known species while intensive surveys are being planned.
Future work performed in Laos should greatly
expand the knowledge on ant diversity in thiscountry. New intensive surveys of Laos areplanned during 2016-2018 by Weeyawat Jaitrong
and Seiki Yamane, which should help address
this gap in knowledge. Without a doubt, many
species and distribution records as well as biological information on ants from the country will
increase knowledge beyond the initial checklist
presented here.

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21

Figs 2 – 19. 2-3, Dolichoderus laotius; 4-5, Dolichoderus thoracicus; 6-7, Iridomyrmex anceps; 8-9, Tapinoma
indicum; 10-11, Tapinoma melanocephalum; 12-13, Technomyrmex albipes; 14-15, Technomyrmex difficilis; 1617, Technomyrmex kraepelini; 18-19, Technomyrmex modiglianii.

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Weeyawat Jaitrong, Benoit Guénard, Evan P. Economo, Nopparat Buddhakala & Seiki Yamane

Checklist of species
AMBLYOPONINAE [1 genus, 1 species]
Myopopone castanea (F. Smith, 1860b)*
Material examined. None.
Distribution in Laos. No further records (Santschi 1920b: 3; referred to as M. mulleri Bingh.)
DOLICHODERINAE [4 genera, 13 species]
Dolichoderus affinis Emery, 1889*
Material examined. None.
Distribution in Laos. Muong Pek (Santschi
1920b: 169, as Dolichoderus affinis var. nigricans, synonym of D. affinis).
Dolichoderus brevis Santschi, 1920b*
Material examined. None.
Distribution in Laos. Vientiane (Santschi
1920b: 167) and Luang Prabang (Santschi
1920b: 167; Dill & Maschwitz 2002: 62).
Dolichoderus laotius Santschi, 1920b* (Figs 2-3)
Material examined. Vientiane: Pak NgumDist., 300 m alt., Phang Dang Village,
12.VI.2010, Sk. Yamane leg. (SKYC).
Distribution in Laos. Luang Prabang(Santschi 1920b: 170; Dill & Maschwitz
2002: 46), Muong Nya (Santschi 1920b:
170; Dill & Maschwitz 2002: 46), Hah Sampong (Dill & Maschwitz 2002: 46), Vientiane (new record).
Dolichoderus taprobanae F. Smith, 1858*
Material examined. None.
Distribution in Laos. Muong Pek (Santschi,
1920b: 169, as Dolichoderus taprobanae
var. obscuripes, synonym of D. taprobanae).
Dolichoderus thoracicus (F. Smith, 1860a)*(Figs 4-5)
Material examined. Vientiane: Pak NgumDist., Phang Dang Village, 12.VI.2010, W.
Jaitrong leg., WJT10-LAO-102 (THNHM);
same loc., 12.VI.2010, Sk. Yamane leg.
(SKYC); Naxaythong Dist., Sivilay Village, 11.VI.2010, W. Jaitrong leg., WJT10LAO-99 (THNHM); same loc., 9.VI.2010,
W. Jaitrong leg., WJT10-LAO-40 (SKYC,THNHM); same loc., 9.VI.2010, Sk. Yamane leg., LA10-SKY-004 (THNHM,
SKYC); Tangon, 22.III.1998, H. Fukuda(SKYC). Vang Vieng: 10.III.1997, H. Fu-

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kuda leg. (SKYC, THNHM). Phong Sali:
Muang Khoua, 22-24.VIII.2005, Y. Ochiai leg. (SKYC); Khoa dist., 1,600 m alt.,
23.XII.2015, Y. Ochiai, LA15/16-YO-01(SKYC). Bolikhamxay: Lak Sao (600-700m alt.), 2.IV.1988, H. Fukuda leg. (SKYC).
Luang Prabang: Kuangsi Waterfall Park,
16.XII.2015, W. Jaitrong leg. (THNHM).
Distribution in Laos. Muong Pek (Santschi
1920b: 169, as Dolichoderus bituberculatus
var. emarginata synonym of D. thoracicus);
Phong Sali, Bolikhamxay, Vientiane, Vang
Vieng, Luang Prabang (new record) and
Xieng Khouang (Yoshitomi et al. 2014).
Iridomyrmex anceps (Roger, 1863) (Figs 6-7)
Material examined. Vientiane: Pak NgumDist., Phang Dang Village, 14.VI.2010, W.
Jaitrong leg. (SKYC, THNHM). Oudomxay:
Namo Dist., Mai, III.2006, Y. Ochiai leg.
Distribution in Laos. Oudomxay and Vientiane (new record).
Tapinoma indicum Forel, 1895 (Figs 8-9)
Material examined. Vientiane: Naxaythong Dist., Sivilay Village, 9.VI.2010, W.
Jaitrong leg., WJT10-LAO-48 (THNHM,
SKYC); same loc., 9.VI.2010, Sk. Yamaneleg., LA10-SKY-018 (THNHM, SKYC);
Somnuek Guesthouse, 3.VII.2012, D. Pickleg., #347 (MCZC). Phong Sali: Huei Pee(1000 m alt.), 19.VIII.2004, Y. Ochiai leg.
(SKYC); Phongsali Hotel, 15.VIII.2012, D.
Pick leg., #348 (MCZC). Luang Prabang:
Santi Resort, 16.XII.2015, W. Jaitrong leg.,
LAO15-WJT-005 (THNHM).
Distribution in Laos. Luang Prabang,
Phong Sali and Vientiane (new record).
Tapinoma melanocephalum (Fabricius, 1793)(Figs 10-11)
Material examined. Vientiane: Vientiane, 24.VIII.2003, Y. Ochiai leg. (SKYC);
Pak Ngum Dist., Phang Dang Village,
14.VI.2010, Sk. Yamane leg. (THNHM,
SKYC); Naxaythong Dist., Sivilay Village, 11.VI.2010, W. Jaitrong leg., WJT10LAO-126 (THNHM, SKYC).
Distribution in Laos. Vientiane (new record).
Technomyrmex albipes (F. Smith, 1861)* (Figs12-13)
Material examined. Vientiane: Pak NgumDist., Phang Dang Village, 12.VI.2010, Sk.

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Figs 20 – 37. 20-21, Technomyrmex yamanei; 22-23, Aenictus binghami; 24-25, Aenictus doydeei; 26-27, Aenictus fuchuanensis; 28-29, Aenictus hodgsoni; 30-31, Aenictus nishimurai; 32-33, Aenictus paradentatus; 34-35,
Aenictus parahuonicus; 36-37, Cerapachys sulcinodis.

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Weeyawat Jaitrong, Benoit Guénard, Evan P. Economo, Nopparat Buddhakala & Seiki Yamane

Yamane leg. (THNHM, SKYC); Naxaythong Dist., Sivilay Village, 11.VI.2010, W.
Jaitrong leg., WJT10-LAO-95 (THNHM,
SKYC); same loc., 11.VI.2010, Sk. Yamane leg., ex dead twig, LA10-SKY-083(THNHM, SKYC). Luang Prabang: Santi Resort, 16.XII.2015, W. Jaitrong leg.,
LAO15-WJT-006 (THNHM).
Distribution in Laos. Vientiane and Luang
Prabang (new record), Xieng Khouang (Yoshitomi et al. 2014).
Technomyrmex difficilis Forel, 1892a (Figs 14-15)
Material examined. Vientiane: Pak
Ngum Dist., Phang Dang Village, 170 malt., 14.VI.2010, Sk. Yamane leg., LA10SKY-121 (THNHM, SKYC); HadsaiphongDist., Thana Village, 6.V.2010, W. Jaitrongleg., WJT10-LAO-206 (THNHM, SKYC).
Luang Prabang: Santi Resort, 16.XII.2015,
W.
Jaitrongleg.,
LAO15-WJT-003(THNHM).
Distribution in Laos. Vientiane and Luang
Prabang (new record).
Technomyrmex kraepelini Forel, 1905 (Figs
16-17)
Material examined. Vientiane: Pak NgumDist., Phang Dang Village, 14.VI.2010, Sk.
Yamane leg. (THNHM, SKYC); same loc.,
14.VI.2010, W. Jaitrong leg. (SKYC); Naxaythong Dist., Sivilay Village, 10.VI.2010,
W. Jaitrong leg. (THNHM).
Distribution in Laos. Vientiane (new record).
Technomyrmex modiglianii Emery, 1900* (Figs
18-19)
Material examined. Bolikhamxay: Lak Sao,
6.III.1997, H. Fukuda leg. (THNHM, SKYC).
Distribution in Laos. Bolikhamxay (Bolton
2007: 92).
Technomyrmex yamanei Bolton, 2007 (Figs
20-21)
Material examined. Vientiane: Naxaythong Dist., Sivilay Village, 10.VI.2010, W.
Jaitrong leg., WJT10-LAO-69 (THNHM,
SKYC); same loc., WJT10-LAO-71(THNHM, SKYC); same loc., 7.V.2010, W.
Jaitrong leg., WJT10-LAO-236 (THNHM);
same loc., 10.VI.2010, Sk. Yamane leg.,
LA10-SKY-033 (THNHM, SKYC); samedata., LA10-SKY-034 (SKYC); same data,
ex bamboo stem, nest in soil, LA10-SKY-039

2 - 19 - AM8 - A Checklist of known ant species of Laos.indd 24(SKYC); same data, nest in soil, LA10-SKY
043 (SKYC); same data, night walk, LA10SKY 063 (SKYC); same data, LA10-SKY
069 (SKYC). Phong Sali: Muang Khoua,
22-24.VIII.2005, Y. Ochiai leg. (SKYC).
Distribution in Laos. Phong Sali and Vientiane (new record).
DORYLINAE [3 genera, 10 species]
Aenictus binghami Forel, 1900a* (Figs 22-23)
Material examined. Vientiane: Pak NgumDist., Phang Dang Village, 12.VI.2010, W.
Jaitrong leg., WJT10-LAO-15 (THNHM,
SKYC); same loc. and date, Sk. Yamane,
LA10-SKY-104 (THNHM, SKYC); sameloc., 13.VI.2010, W. Jaitrong leg., WJT10LAO-18 (THNHM).
Distribution in Laos. Vientiane (Jaitrong etal. 2011: 318; Jaitrong & Yamane 2011: 25).
Aenictus doydeei Jaitrong et Yamane, 2011*(Figs 24-25)
Material examined. Vientiane: Naxaythong Dist., Sivilay Village, 10.VI.2010, W.
Jaitrong leg., WJT10-LAO-13 (type series,
THNHM, SKYC); same loc. and date, Sk.
Yamane, night walk, LA10-SKY-056 (typeseries, THNHM, SKYC).
Distribution in Laos. Vientiane (Jaitrong etal. 2011: 319; Jaitrong & Yamane 2012: 53).
Aenictus fuchuanensis Zhou, 2001* (Figs 26-27)
Material examined. Vientiane: Naxaythong Dist., Sivilay Village, 10.VI.2010, Sk.
Yamane leg., LA10-SKY-058 (THNHM,
SKYC).
Distribution in Laos. Vientiane (Jaitrong &Yamane 2012: 187).
Aenictus hodgsoni Forel, 1901* (Figs 28-29)
Material examined. Vientiane: Naxaythong Dist., Sivilay Village, 10.VI.2010, W.
Jaitrong leg., WJT10-LAO-11 (THNHM,
SKYC); same loc. and date, Sk. Yamane leg.,
LA10-SKY-007 (THNHM, SKYC). Luang
Prabang: Santi Resort, 16.XII.2015, W.
Jaitrong leg., LAO15-WJT-001 (THNHM).
Distribution in Laos. Vientiane (Jaitrong etal. 2011: 321; Jaitrong & Yamane 2011: 35)
and Luang Prabang (new record).
Aenictus nishimurai Terayama et Kubota,
1993* (Figs 30-31)

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Figs 38 – 55. 38-39, Dorylus orientalis; 40-41, Gnamptogenys bicolor; 42-43, Gnamptogenys binghamii; 44-45,
Acropyga butteli; 46-47, Anoplolepis gracilipes; 48-49, Camponotus carin; 50-51, Camponotus nicobarensis;
52-53, Camponotus rufoglaucus; 54-55, Oecophylla smaragdina.

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Weeyawat Jaitrong, Benoit Guénard, Evan P. Economo, Nopparat Buddhakala & Seiki Yamane

Material examined. Vientiane: Pak NgumDist., Phang Dang Village, 14.VI.2010, W.
Jaitrong leg., WJT10-LAO-19 (THNHM,
SKYC); same loc. and date, Sk. Yamane leg.,
LA10-SKY-129 (THNHM, SKYC).
Distribution in Laos. Vientiane (Jaitrong etal. 2011: 321; Jaitrong & Yamane 2012: 61).
Aenictus paradentatus Jaitrong et Yamane,
2012* (Figs 32-33)
Material examined. Vientiane: Pak NgumDist., Phang Dang Village, 12.VI.2010, W.
Jaitrong leg., WJT10-LAO-16 (THNHM);
same loc., 14.VI.2010, WJT10-LAO-20(THNHM, SKYC); same loc. and date, Sk.
Yamane leg., LA10-SKY-126 (THNHM,
SKYC); Naxaythong Dist., Sivilay Village, 10.VI.2010, W. Jaitrong leg., WJT10LAO-12 (THNHM, SKYC); same loc. anddate, Sk. Yamane leg., (THNHM, SKYC).
Distribution in Laos. Vientiane (Jaitrong etal. 2012: 136).
Aenictus parahuonicus Jaitrong et Yamane,
2011* (Figs 34-35)
Material examined. Vientiane: Naxaythong Dist., Sivilay Village, 9.VI.2010, W.
Jaitrong leg., WJT10-LAO-10 (THNHM);
same loc. and date, Sk. Yamane, night walk,LA10-SKY-028 (THNHM, SKYC).
Distribution in Laos. Vientiane (Jaitrong &Yamane 2011: 19).
Cerapachys sulcinodis Emery, 1889 (Figs 36-37)
Material examined. Vientiane: Pak Gnum
Dist., Phag Dang Village, W. Jaitrong leg.,
WJT10-LAO-107 (THNHM, SKYC) ; same
loc. and date, Sk. Yamane, LA10-SKY-095(SKYC).
Distribution in Laos. Vientiane (new record).
Dorylus laevigatus F. Smith, 1857*
Material examined. None.
Distribution in Laos. Luang Prabang(Santschi 1920b: 160).
Dorylus orientalis Westwood, 1835 (Figs 38-39)
Material examined. Vientiane: Naxaythong Dist., Sivilay Village, 10.VI.2010, W.
Jaitrong leg., WJT10-LAO-54 (THNHM,
SKYC); same loc. and date, Sk. Yamane leg.,
LA10-SKY-035 (SKYC).
Distribution in Laos. Vientiane (new record).

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ECTATOMMINAE [1 genus, 2 species]
Gnamptogenys bicolor (Emery, 1889) (Figs 40-41)
Material examined. Vientiane: Pak Gnum
Dist., Phang Dang Village, 14.VI.2010, Sk.
Yamane (SKYC).
Distribution in Laos. Vientiane (new record).
Gnamptogenys binghamii (Forel, 1900b) (Figs42-43)
Material examined. Vientiane: Pak NgumDist., Phang Dang Village, 13.VI.2010, W.
Jaitrong leg., WJT10-LAO-131 (THNHM,
SKYC); same loc. and date, Sk. Yamane, rotting wood, LA10-SKY-117 (SKYC); Naxaythong Dist., Sivilay Village, 7.V.2010, W.
Jaitrong leg., WJT10-LAO-250 (THNHM,
SKYC). Phong Sali, Muang Khoua, 22-24.
VIII.2005, Y. Ochiai (SKYC).
Distribution in Laos. Phong Sali and Vientiane (new record).
FORMICINAE [8 genera, 32 species, 1 subspecies]
Acropyga butteli Forel, 1912 (Figs 44-45)
Material examined. Vientiane: Pak NgumDist., Phang Dang Village, 13.VI.2010, W.
Jaitrong leg., WJT10-LAO-143 (THNHM,
SKYC); same loc., 14.VI.2010, Sk. Yamaneleg., LA10-SKY-134 (SKYC).
Distribution in Laos. Vientiane (new record).
Anoplolepis gracilipes (F. Smith, 1857) (Figs
46-47)
Material examined. Vientiane: Pak NgumDist., Phang Dang Village, 12.VI.2010, W.
Jaitrong leg. (THNHM); Naxaythong Dist.,
Sivilay Village, 9.VI.2010, Sk. Yamane leg.
(SKYC); vicinity of Vientiane, 5.V.1997, H.
Fukuda leg. (SKYC). Phong Sali: Phongsali Hotel, 15.VIII.2012, D. Pick leg., #348
(MCZC). Luang Prabang: Santi Resort,
16.XII.2015, W. Jaitrong leg. (THNHM);
Luang Prabang City, 290 m alt., 03.I.2016,
Y. Ochiai leg. (SKYC).
Distribution in Laos. Vientiane, Phong Sali
and Luang Prabang (new record).
Camponotus camelinus (F. Smith, 1857)
Material examined. Oudomxay: NamoDist., Mai Naatan, 950 m alt., 21-26.
VIII.2004, Y. Ochiai leg. (THNHM, SKYC).

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27

Figs 56 – 73. 56-57, Paratrechina longicornis; 58-59, Polyrhachis abdominalis; 60-61, Polyrhachis armata;62-63, Polyrhachis cryptoceroides; 64-65, Polyrhachis dives; 66-67, Polyrhachis flavicornis; 68-69, Polyrhachis
illaudata; 70-71, Polyrhachis laevissima; 72-73, Polyrhachis phalerata.

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Weeyawat Jaitrong, Benoit Guénard, Evan P. Economo, Nopparat Buddhakala & Seiki Yamane

Phong Sali: Muang Khoua, 22-24.VIII.2005,
Y. Ochiai leg. (SKYC).
Distribution in Laos. Oudoxmai and Phong
Sali (new record).
Camponotus carin Emery, 1889 (Figs 48-49)
Material examined. Vientiane: Pak NgumDist., Phang Dang Village, 12.VI.2010, W.
Jaitrong leg., WJT10-LAO-117 (THNHM);
same loc. and date, Sk. Yamane leg., LA10SKY-107 (SKYC); Naxaythong Dist., Sivilay Village, 9.VI.2010, Sk. Yamane leg.,nest in soil, LA10-SKY-006 (SKYC); sameloc., 10.VI.2010, W. Jaitrong leg., WJT10LAO-92 (THNHM, SKYC); same loc.,
11.VI.2010, Sk. Yamane, ex termite mound(LA10-SKY-073); same loc., 7.V.2010, W.
Jaitrong leg., WJT10-LAO-229 (THNHM,
SKYC); same loc., 9.VI.2010, W. Jaitrongleg., WJT10-LAO-36 (THNHM, SKYC);
same loc., 10.VI.2010, W. Jaitrong leg.,
WJT10-LAO-72 (THNHM, SKYC); Sadsaiphong Dist., Thana Village, 6.V.2010, W.
Jaitrong leg., WJT10-LAO-204 (THNHM).
Oudomxay; Namo Dist., Mai Naatan (950m alt.), 21-26.VIII.2004, Y. Ochiai (SKYC).
Distribution in Laos. Oudoxmai and Vientiane (new record).
Camponotus dolendus Forel, 1892b*
Material examined. None.
Distribution in Laos. Thatom (Santschi
1920b: 174).
Camponotus nicobarensis Mayr, 1865 (Figs50-51)
Material examined. Bolikhamxay: W. footof Annamite Range nr Vientiane, 8.V.1997,
H. Fukuda leg. (SKYC); Naxaythong Dist.,
Sivilay Village, 10.VI.2010, W. Jaitrong leg.,
WJT10-LAO-68 (THNHM, SKYC); same
loc. and date, Sk. Yamane leg. (SKYC).
Phong Sali: Muang Khoua, 22-24.VIII.2005,
Y. Ochiai (SKYC).
Distribution in Laos. Bolikhamxay, Phong
Sali and Vientiane (new record).
Camponotus parius Emery, 1889*
Material examined. None.
Distribution in Laos. Vientiane (Santschi
1924: 22, cited as Camponotus rufoglaucus paria).
Camponotus rufoglaucus (Jerdon, 1851)* (Figs52-53)

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Material examined. Vientiane: Pak
Ngum Dist., Phang Dang Village, 1213.VI.2010, Sk. Yamane leg. (THNHM,
SKYC); Naxaythong Dist., Sivilay Village,
10.VI.2010, Sk. Yamane leg. (SKYC); sameloc., 11.VI.2010, W. Jaitrong leg. (THNHM). Phong Sali: Khoua, 1000 m alt.,
18.VIII.2004, Y. Ochiai leg. (SKYC).
Distribution in Laos. Phonga Sali, Vientiane (new record) and Xieng Khouang (Yoshitomi et al. 2014).
Remarks. No taxonomic revision is available on the C. rufoglaucus group (Myrmosericus) in continental Asia. According toBingham (1903) C. dolendus is a highland
species, while C. rufoglaucus and C. parius
occur in lower elevations and found together.
We have applied the name rufoglaucus to the
populations of continental Asia, adopting awider concept of rufoglaucus. Careful comparison of all the castes and sexes is needed
to evaluate the status of these three forms.
Camponotus singularis (F. Smith, 1858)*
Material examined. Oudomxay: NamoDist., Mai Naatan, 950 m alt., 21-26.
VIII.2004, Y. Ochiai leg. (SKYC). Phong
Sali: Muang Khoua, 22-24.VIII.2005, Y.
Ochiai leg. (SKYC).
Distribution in Laos. Luang Prabang(Santschi 1920b: 174, as Camponotus camelinus var. singularis); Oudomxay and Phong
Sali (new record).
Camponotus variegatus fuscithorax Dalla
Torre, 1893*
Material examined. None.
Distribution in Laos. Luang Prabang [PakVet] (Santschi 1920b: 173), Pak-Vet (Santschi 1924: 22).
Camponotus variegatus proles Emery, 1925*
Material examined. None.
Distribution in Laos. Luang Prabang(Santschi 1920b: 173; Emery 1925: 96).
Colobopsis leonardi Emery, 1889
Material examined. Vientiane: Pak NgumDist., Phang Dang Village, 12.VI.2010, W.
Jaitrong leg. (THNHM); same loc. and date,
Sk. Yamane leg. (SKYC).
Distribution in Laos. Vientiane (new record).
Echinopla cherapunjiensis Bharti et Gul, 2012*Material examined. None.

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29

Figs 74 – 91. 74-75, Polyrhachis proxima; 76-77, Polyrhachis tibialis; 78-79, Cardiocondyla wroughtonii; 80-83,
Carebara affinis; 84-85, Carebara castanea; 86-89, Carebara diversa; 90-91, Carebara pygmaea.

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Weeyawat Jaitrong, Benoit Guénard, Evan P. Economo, Nopparat Buddhakala & Seiki Yamane

Distribution in Laos. Luang Nam Tha(Zettel & Laciny 2015: 113).
Oecophylla smaragdina (Fabricius, 1775)*(Figs 54-55)
Material examined. Vientiane: Pak NgumDist., Phang Dang Village, 12-13.VI.2010,
Sk. Yamane leg. (SKYC, THNHM); Naxaythong Dist., Sivilay Village, 7.V.2010,
W. Jaitrong leg. (SKYC, THNHM); Hadsaiphong Dist., Thana Village, 6.V.2010,
W. Jaitrong leg. (THNHM); same loc.,
9.VI.2010, Sk. Yamane leg. (SKYC);
Ang Nam Ngum, 24.VIII.2003, Y. Ochiaileg. (THNHM). Oudomxay: Namo, Mai
Naatan, 21-26.VIII.2004, Y. Ochiai leg.
(SKYC, THNHM). Lak Sao: 600-700 m
alt., 22.III.1998, H. Fukuda leg. (SKYC,THNHM). Phong Sali: Khoa, Huei Pee(1000 m alt.), Y. Ochiai leg. (SKYC); Muang
Khoua, 22-24.VIII.2005, Y. Ochiai leg.
(SKYC). Luang Prabang: Santi Resort,
17.XII.2015, W. Jaitrong leg. (THNHM);
Luang Prabang City, 290 m alt., 03.I.2016,
Y. Ochiai leg.
Distribution in Laos. Vientiane (Boulidam 2010;
Itterbeeck et al. 2014), Bolikhamxay, Oudomxay, Luang Prabang (https://www.flickr.
com/photos/onefatmonkey/6542531847),
Khammouan(https://www.flickr.com/
photos/micche/6993419826),
Savannakhet (Yhoung-Aree & Viwatpanich
2005), Champasak (https://www.flickr.
com/photos/101244975)
and
Bokeo
(https://www.flickr.com/photos/bartholomae/4060120523).
Paratrechina longicornis (Latreille, 1802) (Figs
56-57)
Material examined. Vientiane: Hadsaiphong, Naxaythong Dist., Thana subdist. 6.V.2010, W. Jaitrong leg. (SKYC);
Sivilay Village, 9.V.2010, W. Jaitrong leg.
(THNHM) same loc. and date, Sk. Yamane(SKYC); Somnuek Guesthouse, 3.VII.2012,
D. Pick leg., #347 (MCZC). Phong Sali:
Phongsali Hotel, 15.VIII.2012, D. Pickleg., #348 (MCZC). Luang Prabang:
Santi Resort, 16.XII.2015, W. Jaitrong leg.
(THNHM). Luang Prabang: Santi Resort,
16.XII.2015, W. Jaitrong leg. (THNHM).

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Distribution in Laos. Vientiane, Phong Sali
and Luang Prabang (new record).
Polyrhachis abdominalis F. Smith, 1858 (Figs
58-59)
Material examined. Vientiane: Pak NgumDist., Phang Dang Village, 13.VI.2010, W.
Jaitrong leg. (THNHM); same loc. and date,
Sk. Yamane leg. (SKYC). Luang Prabang:
Kuangsi Waterfall Park, 16.XII.2015, W.
Jaitrong leg. (THNHM).
Distribution in Laos. Vientiane and Luang
Prabang (new record).
Polyrhachis armata (Le Guillou, 1842) (Figs60-61)
Material examined. Vientiane: Pak NgumDist., Phang Dang Village, 12-13.VI.2010,
W. Jaitrong leg. (THNHM, SKYC); same
loc. 12-14.VI.2010, Sk. Yamane leg.
(SKYC); Naxaythong Dist., Sivilay Village,
7.V.2010, W. Jaitrong leg., WJT10-LAO-233(THNHM).
Distribution in Laos. Vientiane (new record).
Polyrhachis bellicosa F. Smith, 1859*
Material examined. None.
Distribution in Laos. Bolikhamxay [BanVan Heue 20km E of Phou-kowkuei] (Hung,1970).
Remarks. According to Kohout (2014) the
populations of continental Asia belong to
P. olybria Forel, 1912, not to P. bellicosa.
We have never seen P. bellicosa specimensfrom the Indo-Chinese Peninsula. Although
Kohout did not refer to the above-mentioned
record from Laos cited by Hung (1970), thisrecord was most probably based on misidentification.
Polyrhachis bicolor F. Smith, 1858*
Material examined. None.
Distribution in Laos. No further record(Ran & Zhou 2013).
Polyrhachis bihamata (Drury, 1773)*
Material examined. Oudomxay: NamoDist., Mai Naatan (950 m alt.), 22-23.
VIII.2004, Y. Ochiai leg. (SKYC).
Distribution in Laos. Luang Prabang,
Muang Sai (Santschi 1920b: 174); Muang
Pek (Hung 1970: 16; Kohout 2014: 10) andOudomxay (new record).
Polyrhachis cryptoceroides Emery, 1887a (Figs
62-63)

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31

Figs 92 – 109. 92-93, Cataulacus granulatus; 94-95, Crematogaster aurita; 96-97, Crematogaster ferrarii; 9899, Crematogaster modiglianii; 100-101, Crematogaster rogenhoferi; 102-103, Lophomyrmex lucidus; 104-105,
Mayriella transfuga; 106-107, Meranoplus bicolor; 108-109, Monomorium floricola.

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Weeyawat Jaitrong, Benoit Guénard, Evan P. Economo, Nopparat Buddhakala & Seiki Yamane

Material examined. Vientiane: Pak NgumDist., Phang Dang Village, 14.VI.2010, Sk.
Yamane leg., LA10-SKY-127 (THNHM,
SKYC).
Distribution in Laos. Vientiane (new record).
Polyrhachis dives F. Smith, 1857* (Figs 64-65)
Material examined. Bolikhamxay: Lak
Sao, 5.IV.1998, 600-700 m alt., H. Fukuda
leg. (SKYC, THNHM).
Distribution in Laos. This species was recorded from Laos without further information (Ran & Zhou 2013; Wang et al. 2012;
Li 2006). In the present paper we record thisspecies for the first time from Bolikhamxay.
Polyrhachis emmae Santschi, 1920b*
Material examined. None.
Distribution in Laos. Luang Prabang(Santschi 1920b: 175).
Polyrhachis flavicornis F. Smith, 1857 (Figs
66-67)
Material examined. Vientiane: Pak NgumDist., Phang Dang Village, 14.VI.2010, W.
Jaitrong leg., WJT10-LAO-145 (THNHM,
SKYC); Naxaythong Dist., Sivilay Village, 11.VI.2010, W. Jaitrong leg., WJT10LAO-80 (THNHM, SKYC); same loc. anddate, Sk. Yamane leg., carton nest on leaf,
LA10-SKY-071, -077, -086 (THNHM,
SKYC).
Distribution in Laos. Vientiane (new record).
Polyrhachis halidayi Emery, 1889*
Material examined. None.
Distribution in Laos. Vientiane (Yoshitomi et al., 2014), Luang Prabang, Tong-La(Santschi 1920b: 174) and Xieng Khousang(Yoshitomi et al. 2014).
Polyrhachis hemiopticoides Mukerjee, 1930*
Material examined. None.
Distribution in Laos. Bolikhamxay (Kohout 2013: 150; locality name is misspelled
as Borikhane).
Polyrhachis illaudata Walker, 1859* (Figs 68-69)
Material examined. Vientiane: Pak NgumDist., Phang Dang Village, 13.VI.2010, W.
Jaitrong leg. (THNHM).
Distribution in Laos. Vientiane (new record), Ban Khoumkhan and Ban Vangheua[Pakse?] (Mezger & Moreau 2015).

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Polyrhachis laevissima F. Smith, 1858* (Figs70-71)
Material examined. Phong Sali: Huei Pee,
1000 m alt., 18.VIII.2004, Y. Ochiai leg.
(THNHM, SKYC).
Distribution in Laos. Phong Sali (new record).
Polyrhachis phalerata Menozzi, 1926 (Figs
72-73)
Material examined. Vientiane: Pak NgumDist., Phang Dang Village, 14.VI.2010, W.
Jaitrong leg., WJT10-LAO-156 (THNHM,
SKYC); same loc. and date, Sk. Yamane,
carton nest, LA10-SKY-132 (SKYC).
Distribution in Laos. Vientiane (new record).
Remarks. Polyrhachis mucronata F. Smith,1859 recorded from Bolaven Plateau and
Ban Vangheus by Mezger and Moreau(2015) may be this species or P. moesta Emery, 1887. P. mucronata is a species ranging
from northern Australia through New Guinea
to eastern Indonesia (Kohout 2010).
Polyrhachis proxima Roger, 1863 (Figs 74-75)
Material examined. Vientiane: Pak NgumDist., Phang Dang Village, 12.VI.2010, Sk.
Yamane leg., attacked by Aenictus (THNHM, SKYC); Naxaythong Dist., Sivilay Village, 10.VI.2010, Sk. Yamane leg. (SKYC);
same loc., 11.VI.2010, W. Jaitrong leg.
(THNHM).
Distribution in Laos. Vientiane (new record).
Polyrhachis rufipes F. Smith, 1858
Material examined. Vientiane: Pak NgumDist., Phang Dang Village, 14.VI.2010, Sk.
Yamane leg. (SKYC).
Distribution in Laos. Vientiane (new record).
Polyrhachis tibialis F. Smith, 1858-complex(Figs 76-77)
Material examined. Vientiane: Pak NgumDist., Phang Dang Village, 13.VI.2010,
W. Jaitrong leg. (THNHM); same loc.,
14.VI.2010, Sk. Yamane (SKYC).
Distribution in Laos. Vientiane (new record).
Polyrhachis vitalisi Santschi, 1920a*
Material examined. None.
Distribution in Laos. Muong Pek (Santschi
1920a: 567; 1920b: 175).
MYRMICINAE [17 genera, 37 species, 2 subspecies]

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Figs 110 – 127. 110-111, Monomorium pharaonis; 112-113, Paratopula macta; 114-115, Pheidole dugasi; 116119, Pheidole hongkongensis; 120-123, Pheidole plagiaria; 124-127, Pheidole tumida.

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Weeyawat Jaitrong, Benoit Guénard, Evan P. Economo, Nopparat Buddhakala & Seiki Yamane

Cardiocondyla wroughtonii (Forel, 1890) (Figs
78-79)
Material examined. Vientiane: Naxaythong Dist., Sivilay Village, 11.VI.2010,
Sk. Yamane leg., LA10-SKY-070 (THNHM,
SKYC).
Distribution in Laos. Vientiane (new record).
Carebara affinis (Jerdon, 1851) (Figs 80-83)
Material examined. Vientiane: Pak NgumDist., Phang Dang Village, 13.VI.2010, Sk.
Yamane leg. (SKYC); same loc., 14.VI.2010,
W. Jaitrong leg. (THNHM, SKYC); Naxaythong Dist., Sivilay Village, 9.VI.2010, Sk.
Yamane leg., LA10-SKY-010 (SKYC); sameloc., 11.VI.2010, W. Jaitrong leg., WJT10LAO-91 (THNHM).
Distribution in Laos. Vientiane (new record).
Carebara castanea F. Smith, 1858 (Figs 84-85)
Material examined. Vientiane: Naxaythong Dist., Sivilay Village, 10.VI.2010, W.
Jaitrong leg., WJT-LAO-078 (THNHM);
same loc., 11.VI.2010, Sk. Yamane leg.
(SKYC).
Distribution in Laos. Vientiane (new record).
Carebara castanea augustata Santschi, 1920b*
Material examined. None.
Distribution in Laos. Thatom (Santschi
1920b: 162).
Carebara diversa (Jerdon, 1851) (Figs 86-89)
Material examined. Vientiane: Pak NgumDist., Phang Dang Village, 14.VI.2010, W.
Jaitrong leg. (SKYC, THNHM); same loc.
and date, Sk. Yamane leg. (SKYC); Naxaythong Dist., Sivilay Village, 9.VI.2010,
Sk. Yamane leg., LA10-SKY-020 (SKYC,THNHM); Hadsaiphong Dist., Dondu Village, 6.V.2010, W. Jaitrong leg., WJT10LAO-239 (SKYC, THNHM). Luang Prabang: Kuangsi Waterfall Park, 16.XII.2015,
W.
Jaitrongleg.,
LAO15-WJT-008(THNHM); Luang Prabang City, 290 m alt.,
02.I.2016, Y. Ochiai leg., LA15/16-YO-02(SKYC).
Distribution in Laos. Vientiane and Luang
Prabang (new record).
Carebara diversa laotina (Santschi, 1920b)*
Material examined. None.
Distribution in Laos. Pakvet, Luang Prabang, Thatom (Santschi 1920b: 162).

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Carebara petulens (Santschi, 1920b)*
Material examined. None.
Distribution in Laos. Muong Pek (Santschi 1920b: 164, as Pheidologeton petulens
nov. sp.).
Carebara pygmaea (Emery, 1887b)-complex(Figs 90-91)
Material examined. Vientiane: Naxaythong Dist., Sivilay Village, 10.VI.2010,
W. Jaitrong leg., WJT-LAO-059 (SKYC,THNHM); same loc. and date, Sk. Yamaneleg. (SKYC).
Distribution in Laos. Vientiane (new record).
Cataulacus granulatus (Latreille, 1802) (Figs
92-93)
Material examined. Vientiane: Pak NgumDist., Phang Dang Village, 14.VI.2010,
W. Jaitrong leg. (THNHM, SKYC); Naxaythong Dist., Sivilay Village, 10.VI.2010,
W. Jaitrong leg., WJT-LAO-079 (THNHM,
SKYC); same loc. and date, Sk. Yamaneleg., nest in bamboo stem, LA10-SKY-029(SKYC); same loc., 11.VI.2010, Sk. Yamane (SKYC); Houi Nyang near Tha Ngon,
26.VIII.2003, Y. Ochiai leg. (SKYC).
Distribution in Laos. Vientiane (new record).
Crematogaster aurita Karavaiev, 1935* (Figs
94-95)
Material examined. Vientiane: Pak NgumDist., Phang Dang Village, 12.VI.2010, Sk.
Yamane leg., LA10-SKY-99 (SKYC, THNHM); Saythany Dist., Dongdok Village,
6.V.2010, W. Jaitrong leg., WJT10-ALO-203(THNHM, SKYC); Naxaythong Dist., Sivilay Village, 9.VI.2010, Sk. Yamane leg.,
LA10-SKY-015 (SKYC); same loc.,10.
VI.2010, Sk. Yamane leg., LA10-SKY-032,-041 (SKYC, THNHM); Ang Nam Ngum,
24.VIII.2003, Y. Ochiai leg. (SKYC).
Distribution in Laos. Vientiane (Hosoishi &Ogata 2009a, b).
Crematogaster bouvardi Santschi, 1920*
Material examined. None.
Distribution in Laos. Xiang Mouang (Santschi 1920b: 161; described as var. salvazai of
C. walshi st. bouvardi).
Crematogaster ferrarii Emery, 1888 (Figs 96-97)
Material examined. Vientiane: Naxaythong Dist., Sivilay Village, 10.VI.2010,
10.VI.2010, Sk. Yamane leg., LA10-

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Figs 128 – 145. 128-131, Pheidole yeensis; 132-133, Pristomyrmex punctatus; 134-135, Proatta butteli; 136-139,
Solenopsis geminata; 140-141, Tetramorium flavipes; 142-143, Tetramorium insolens; 144-145, Tetramorium
lanuginosum.

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Weeyawat Jaitrong, Benoit Guénard, Evan P. Economo, Nopparat Buddhakala & Seiki Yamane

SKY-047, -048, -054 (SKYC); same loc. anddate, W. Jaitrong leg., WJT10-LAO-051,-076 (SKYC); Hadsaiphong Dist., Dondu
Village, 8.V.2010, W. Jaitrong leg., WJT10LAO-237 (SKYC); Ang Nam Ngun,
24.VIII.2003, Y. Ochiai leg. (SKYC). Phong
Sali: Muang Khoua, 22-24.VIII.2005, Y.
Ochiai leg. (SKYC).
Distribution in Laos. Phong Sali and Vientiane (new record).
Crematogaster modiglianii Emery, 1900* (Figs
98-99)
Material examined. Vientiane: Pak NgumDist., Phag Dang Village, 13.VI.2010, W.
Jaitrong leg., WJT10-LAO-136 (THNHM);
same loc., 12.VI.2010, Sk. Yamane leg.,
LA10-SKY-089 (THNHM, SKYC).
Distribution in Laos. Vang Vieng (Hosoishi
et al. 2010) and Vientiane (new record).
Crematogaster quadriruga Forel, 1911a*
Material examined. Vientiane: Pak NgumDist., Phag Dang Village, 13.VI.2010, Sk.
Yamane leg. (SKYC).
Distribution in Laos. Vientiane (Hosoishi &Ogata 2016).
Crematogaster rogenhoferi Mayr, 1879-complex* (Figs 100-101)
Material examined. Vientiane: Pak NgumDist., Phag Dang Village, 12.VI.2010, W. Jaitrong leg., WJT10-LAO-100 (SKYC, THNHM); same loc. and date, Sk. Yamane leg.,
LA10-SKY-103 (SKYC); same loc. and date,
Sk. Yamane leg., night walk (SKYC); Naxaythong Dist., Sivilay Village, 10.VI.2010, W.
Jaitrong leg (THNHM); same loc. and date,
Sk. Yamane leg, night walk, LA10-SKY-051(SKYC). Bolikhamxay: Lak Sao, 600-700 m
alt., 2.IV.1998, H. Fukuda leg. (SKYC). Luang Prabang: Santi Resort, 16.XII.2015, W.
Jaitrong leg., LAO15-WJT-002 (THNHM).
Distribution in Laos. Vientiane, Luang Prabang (new record) and Xieng Khouang (Yoshitomi et al. 2014).
Crematogaster treubi Emery, 1896
Material examined. Vientiane: Pak NgumDist., Phag Dang Village, 12.VI.2010, Sk.
Yamane leg. (SKYC); Naxaythong Dist.,
Sivilay Village, 9.VI.2010, Sk. Yamane leg.,
LA10-SKY-002, -009, -053 (SKYC); same
loc. 10.VI.2010, W. Jaitrong leg., WJT10-

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LAO-59 (SKYC); Hadsaiphong dist., Dondu Village, 8.V.2010, WJT10-LAO-238(SKYC).
Distribution in Laos. Vientiane (new record).
Liomyrmex gestroi (Emery, 1887b)*
Material examined. None.
Distribution in Laos. Muong Pek (Santschi,
1920b: 160, as Liomyrmex aurianus, synonym of L. gestroi).
Lophomyrmex lucidus Menozzi, 1930 (Figs102-103)
Material examined. Vientiane: Naxaythong Dist., Sivilay Village, 7.V.2010, W.
Jaitrong leg. (THNHM, SKYC); sameloc., 10.VI.2010, W. Jaitrong leg., WJT10LAO-121 (THNHM); same loc., 11.VI.2010,
Sk. Yamane, LA10-SKY-081 (SKYC).
Distribution in Laos. Vientiane (new record).
Mayriella transfuga Baroni Urbani, 1977 (Figs
104-105)
Pak Ngum Dist., Phag Dang Village,
13.VI.2010, Sk. Yamane leg. (SKYC).
Distribution in Laos. Vientiane (new record).
Meranoplus bicolor Guérin-Méneville, 1844)*(Figs 106-107)
Material examined. Vientiane: Naxaythong Dist., Sivilay Village, 9.VI.2010, W.
Jaitrong leg. (SKYC); same loc. and date,
Sk. Yamane leg. (SKYC); same loc.,10.
VI.2010, Sk. Yamane leg., LA10-SKY-038(THNHM, SKYC). Oudomxay: Namo Dist.,
Mai Naatan, 26.VIII.2004, Y. Ochiai leg.
(SKYC).
Distribution in Laos. Sedone, Pakse(Schödl 1999: 64); Oudomxay and Vientiane(new record).
Meranoplus laeviventris Emery, 1889*
Material examined. None.
Distribution in Laos. 25 km SE Muang
Sing (Schödl, 1998: 380).
Monomorium floricola (Jerdon, 1851) (Figs
108-109)
Material examined. Vientiane: Naxaythong Dist., Sivilay Village, 7.V.2010, W.
Jaitrong leg., WJT10-LAO-255 (THNHM,
SKYC); Hadsaiphong Dist., Thana Village, 7.V.2010, W. Jaitrong leg. (THNHM,
SKYC); Saytharry dist., Dongdock Village,
6.V.2010, W. Jaitrong leg. (SKYC).

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37

Figs 146 – 163. 146-149, Tetramorium polymorphum; 150-151, Tetramorium smithi; 152-153, Trichomyrmex destructor; 154-155, Anochetus graeffei; 156-157, Brachyponera chinensis; 158-159, Centromyrmex feae; 160-161,
Diacamma longitudinale; 162-163, Diacamma rugosum.

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Weeyawat Jaitrong, Benoit Guénard, Evan P. Economo, Nopparat Buddhakala & Seiki Yamane

Distribution in Laos. Vientiane (new record) and no locality (Rapusas et al. 2006).
Monomorium pharaonis (Linnaeus, 1758) (Figs
110-111)
Material examined. Vientiane: Naxaythong Dist., Sivilay Village, 9.VI.2010, W.
Jaitrong leg. (THNHM); Hadsaiphong Dist.,Thana Village, 6.V.2010, W. Jaitrong leg.,
WJT10-LAO-210 (THNHM, SKYC).
Distribution in Laos. Vientiane (new record).
Paratopula macta Bolton, 1988 (Figs 112-113)
Material examined. Vientiane: Pak NgumDist., Phang Dang Village, 14.VI.2010, W.
Jaitrong leg. (THNHM, SKYC).
Distribution in Laos. Vientiane (new record).
Pheidole dugasi Forel, 1911b (Figs 114-115)
Material examined. Vientiane: Naxaythong Dist., Sivilay Village, 9.VI.2010,
Sk. Yamane leg. (THNHM).
Distribution in Laos. Vientiane (new record).
Pheidole hongkongensis Wheeler, 1928 (Figs116-119)
Material examined. Vientiane: Naxaythong Dist., Sivilay Village, 10.VII.2010,
Sk. Yamane, LA10-SKY-046 (SKYC); sameloc., 11.VI.2010, W. Jaitrong leg., WJT10LAO-125 (THNHM, SKYC).
Distribution in Laos. Vientiane (new record).
Pheidole plagiaria F. Smith, 1860b* (Figs
120-123)
Material examined. Vientiane: Pak NgumDist., Phang Dang Village, 12.VI.2010, W.
Jaitrong leg., WJT10-LAO-116 (THNHM,
SKYC).
Distribution in Laos. Vientiane (new record) and Xieng Khouang (Yoshitomi et al.
2014).
Pheidole tumida Eguchi, 2008 (Figs 124-127)
Material examined. Vientiane: Pak NgumDist., Phag Dang Village, 14.VI.2010, W.
Jaitrong leg., WJT10-LAO-147 (THNHM);
same loc., 14.VI.2010, Sk. Yamane leg.,
LA10-SKY-120 (SKYC, THNHM); Naxaythong Dist., Sivilay Village, 11.VI.2010, W.
Jaitrong leg., WJT10-LAO-82 (THNHM);
same loc., 11.VI.2010, W. Jaitrong leg.,
WJT10-LAO-98 (THNHM).
Distribution in Laos. Vientiane (new record).
Pheidole yeensis Forel, 1902 (Figs 128-131)
Material examined. Vientiane: Naxaythong2 - 19 - AM8 - A Checklist of known ant species of Laos.indd 38

Dist., Sivilay Village, 7.V.2010, W. Jaitrongleg., WJT10-LAO-234 (THNHM). Luang
Prabang: Santi Resort, 17.XII.2015, W.
Jaitrong leg., LAO15-WJT-007 (THNHM).
Distribution in Laos. Vientiane and Luang
Prabang (new record).
Pristomyrmex punctatus (F. Smith, 1860b)*(Figs 132-133)
Material examined. Vientiane: Naxaythong Dist., Sivilay Village, 7.V.2010, W.
Jaitrong leg., WJT10-LAO-226 (SKYC,THNHM); same loc., 9.VI.2010, W. Jaitrongleg., WJT10-LAO-30 (SKYC, THNHM);
same loc., 11.VI.2010, Sk. Yamane leg.
(SKYC).
Distribution in Laos. Muong Pek (Santschi 1920b: 160, as Pristomyrmex japonicus
Forel, 1900 junior synonym of P. punctatus);
Vientiane (new record).
Proatta butteli Forel, 1912 (Figs 134-135)
Material examined. Vientiane: Pak NgumDist., Phang Dang Village, 12.VI.2010, W.
Jaitrong leg. (THNHM); Naxaythong Dist.,
Sivilay Village, 10.VI.2010, W. Jaitrongleg., WJT10-LAO-70 (THNHM); sameloc., 10.VI.2010, Sk. Yamane leg., LA10SKY-042 (THNHM, SKYC); same loc.,
11.VI.2010, Sk. Yamane leg. (SKYC).
Distribution in Laos. Vientiane (new record).
Recurvidris recurvispinosa (Forel, 1890)
Material examined. Pak Ngum Dist., Phang
Dang Village, 13.VI.2010, Sk. Yamane leg.
(SKYC).
Distribution in Laos. Vientiane (new record).
Solenopsis geminata (Fabricius, 1804)* (Figs
136-139)
Material examined. Vientiane: Pak NgumDist., Phang Dang Village, 12.VI.2010,
Sk. Yamane leg. (SKYC); same loc.,
14.VI.2010, W. Jaitrong leg. (THNHM); Naxaythong Dist., Sivilay Village, 9.VI.2010,
W. Jaitrong leg., WJT10-LAO-33 (THNHM,
SKYC); Hadsaiphong Dist., Thana Village,
6.V.2010, W. Jaitrong leg. (THNHM); Somnuek Guesthouse, 3.VII.2012, D. Pick leg.,#347 (MCZC).
Distribution in Laos. Luang Prabang (Santschi, 1920b: 161); Vientiane (new record).
Tetramorium flavipes Emery, 1893 (Figs 140-141)
Material examined. Vientiane: Pak Ngum

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39

Figs 164 – 181. 164-165, Ectomomyrmex leeuwenhoeki; 166-167, Harpegnathos venator; 168-169, Leptogenys
diminuta; 170-171, Leptogenys hysterica; 172-173, Leptogenys kraepelini; 174-175, Pseudoneoponera rufipes;176-177, Odontomachus rixosus; 178-179, Odontomachus simillimus; 180-181, Odontoponera denticulata.

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Weeyawat Jaitrong, Benoit Guénard, Evan P. Economo, Nopparat Buddhakala & Seiki Yamane

Dist., Phang Dang Village, 12.VI.2010, Sk.
Yamane leg. (SKYC); same loc. and date,
W. Jaitrong leg. (SKYC); Naxaythong Dist,
Sivilay Village, 9.VI.2010, W. Jaitrong leg.,
WJT10-LAO-043 (THNHM, SKYC); same
loc. and date, Sk. Yamane leg. , ex termite
mound, LA10-SKY-022, -24 (SKYC); sameloc., 10.VI.2010, W. Jaitrong leg., WJT10LAO-064 (THNHM); same loc. and date,
Sk. Yamane leg., nest under log, LA10SKY-055); same loc., 12.VI.2010, Sk. Yamane leg., rotting wood, LA10-SKY-093,-102 (SKYC).
Distribution in Laos. Vientiane (new record).
Tetramorium insolens (F. Smith, 1861) (Figs142-143)
Material examined. Vientiane: Nasaythong Dist., Sivilay Village, 9.VI.2010, W.
Jaitrong leg., WJT-LAO-31, -039 (THNHM,
SKYC); same loc. and date, Sk. Yamane leg.,ex termite mound, LA10-SKY-014, -026
(SKYC); same loc., 10.VI.2010, W. Jaitrongleg., WJT10-LAO-062 (THNHM).
Distribution in Laos. Vientiane (new record).
Tetramorium lanuginosum Mayr, 1870 (Figs144-145)
Material examined. Vientiane: Naxaythong Dist., Sivilay Village, 9.VI.2010, W.
Jaitrong leg., WJT10-LAO-045 (THNHM);
same loc. and date, Sk. Yamane leg. (SKYC).
Phong Sali: Phongsali Hotel, 15.VIII.2012,
D. Pick leg., #348 (MCZC).
Distribution in Laos. Vientiane and Phong
Sali (new record).
Tetramorium polymorphum Yamane et
Jaitrong, 2011* (Figs 146-149)
Material examined. Vientiane: Pak NgumDist., Phang Dang Village, 12.VI.2010, W.
Jaitrong leg., WJT10-LAO-114 (THNHM,
SKYC, type series); same colony, Sk. Yamane leg. LA10-SKY-100 (THNHM,
SKYC, type series).
Distribution in Laos. Vientiane (Yamane &Jaitrong 2011: 68).
Tetramorium smithi Mayr, 1879 (Figs 150-151)
Material examined. Hadsaiphong Dist.,Thana Village, 6.V.2010, W. Jaitrong leg.
(THNHM).
Distribution in Laos. Vientiane (new record).

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Trichomyrmex destructor (Jerdon, 1851) (Figs
152-153)
Material examined. Vientiane: NaxaythongDist., Sivilay Village, 10.VI.2010, W. Jaitrongleg., WJT10-LAO-120 (THNHM, SKYC);
same loc., 10.VI.2010, Sk. Yamane leg.,
LA10-SKY-059, -133 (THNHM, SKYC).
Phong Sali: Phongsali Hotel, 15.VIII.2012,
D. Pick leg., #348 (MCZC). Bolikhamxay:
Lak Sao, 600-700 m alt., 2.IV.1998, H. Fukuda leg. (THNHM, SKYC).
Distribution in Laos. Bolikhamxay, Vientiane and Phong Sali (new record).
PONERINAE [11 genera, 20 species]
Anochetus graeffei Mayr, 1870 (Figs 154-155)
Material examined. Vientiane: Naxaythong Dist., Sivilay Village, 10.VI.2010,
W. Jaitrong leg. (THNHM); same loc.,
11.VI.2010, Sk. Yamane leg., ex soil, LA10SKY-036 (SKYC); same loc., 5.V.2010,
W. Jaitrong leg, WJT10-LAO30 (SKYC);
Pak Ngum Dist., Phang Dang Village,
12.VI.2010, Sk. Yamane leg., LA10SKY-094 (THNHM, SKYC).
Distribution in Laos. Vientiane (new record).
Brachyponera chinensis (Emery, 1895)-complex (Figs 156-157)
Material examined. Vientiane: Pak NgumDist., Phang Dang Village, 13.VI.2010,
W.
Jaitrong
leg.,
WJT10-LAO-128(THNHM); Naxaythong Dist., Sivilay Village, 11.VI.2010, Sk. Yamane leg., LA10SKY-075 (THNHM, SKYC).
Distribution in Laos. Vientiane (new record).
Brachyponera luteipes Mayr, 1862*
Material examined. None.
Distribution in Laos. Tuong La (Santschi
1920b: 159, cited as Euponera (Brachyponera) luteipes).
Centromyrmex feae (Emery, 1889) (Figs 158-159)
Vientiane: Naxaythong Dist., Sivilay Village, 10.VI.2010, W. Jaitrong leg., WJT10LAO-63 (THNHM); same loc. and date, Sk.
Yamane leg. (SKYC); same loc., 7.V.2010,
WJT10-LAO-251 (THNHM, SKYC).
Distribution in Laos. Vientiane (new record).
Diacamma ceylonense oribiculata Santschi,
1932*

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41

Figs 182 – 197. 182-183, Platythyrea parallela; 184-185, Probolomyrmex longiscapus; 186-187, Tetraponera allaborans; 188-189, Tetraponera attenuata; 190-191, Tetraponera bingami; 192-193, Tetraponera nigra; 194-195,
Tetraponera nitida; 196-197, Tetraponera rufonigra.

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Weeyawat Jaitrong, Benoit Guénard, Evan P. Economo, Nopparat Buddhakala & Seiki Yamane

Material examined. None.
Distribution in Laos. Luang Prabang(Santschi 1932: 14).
Diacamma holzschuhi Laciny, Pal et Zettel,
2015*
Material examined. None.
Distribution in Laos. Hua Phan (Laciny et
al. 2015: 115).
Diacamma longitudinale Emery, 1889* (Figs
160-161)
Material examined. Vientiane: NaxaythongDist., Sivilay Village, 10.VI.2010, W. Jaitrong leg., WJT10-LAO-66 (THNHM).
Distribution in Laos. Luang Prabang (Santschi 1920b: 158); Vientiane (new record).
Diacamma rugosum (Le Guillou, 1842) (Figs
162-163)
Material examined. Vientiane: Pak NgumDist., Phang Dang Village, 14.VI.2010, W.
Jaitrong leg., WJT10-LAO-153 (THNHM);
same loc. and date, Sk. Yamane leg., nest under log, LA10-SKY-123 (SKYC).
Distribution in Laos. Vientiane (new record).
Ectomomyrmex leeuwenhoeki (Forel, 1886)*(Figs 164-165)
Material examined. Vientiane: Pak NgumDist., Phang Dang Village, 12.VII.2010, Sk.
Yamane leg., surface soil, LA10-SKY-088(SKYC); same loc., 14.VI.2010, W. Jaitrongleg., WJT10-LAO-155 (THNHM, SKYC).
Phong Sali, Khoua, Hoei Pee (1000 m alt.),
19.VIII.2004, Y. Ochiai leg. (SKYC).
Distribution in Laos. Phong Sali and Vientiane (Jaitrong et al. 2011: 319, cited as
Pachycondyla leeuwenhoeki, prey of Aenictus cf. dentatus.).
Harpegnathos venator F. Smith, 1858 (Figs
166-167)
Material examined. Phong Sali: Khoa,
Huei Pee, 1000 m alt., 19.VIII.2004, Y.
Ochiai leg. (THNHM).
Distribution in Laos. Phong Sali (newrecord).
Leptogenys diminuta (F. Smith, 1857)-complex(Figs 168-169)
Material examined. Vientiane: Pak NgumDist., Phang Dang Village, 14.VI.2010, W.
Jaitrong leg. (THNHM); same loc. and date,
Sk. Yamane leg. (SKYC).

2 - 19 - AM8 - A Checklist of known ant species of Laos.indd 42

Distribution in Laos. Vientiane (new record).
Leptogenys hysterica Forel, 1900b (Figs 170-171)
Material examined. Vientiane: Pak NgumDist., Phang Dang Village, 14.VI.2010, W.
Jaitrong leg., WJT10-LAO-149 (THNHM,
SKYC); same loc. and date, Sk. Yamane leg.
(SKYC); Naxaythong Dist., Sivilay Village,
7.V.2010, W. Jaitrong leg., WJT10-LAO-223(THNHM, SKYC).
Distribution in Laos. Vientiane (new record).
Leptogenys khammouanensis Roncin et Deharveng, 2003*
Material examined. None.
Distribution in Laos. Khammouan (Roncin& Deharveng 2003: 920).
Remarks. This is a unique species specialized in caves.
Leptogenys kraepelini Forel, 1905 (Figs 172-173)
Material examined. Vientiane: Pak NgumDist., Phang Dang Village, 13.VI.2010, rotting wood, Sk. Yamane leg., LA10-SKY-110(SKYC); same loc. 14.VI.2010, W. Jaitrong
leg. (THNHM).
Distribution in Laos. Vientiane (new record).
Odontomachus monticola Emery, 1892*
Material examined. Oudomxay: NamoDist., Mai Naatan, 950 m alt., 21-26.
VIII.2004, Y. Ochiai leg. (THNHM, SKYC).
Distribution in Laos. Muong Pek (Santschi
1920b: 159) and Oudomxay (new record).
Odontomachus rixosus F. Smith, 1857 (Figs
176-177)
Material examined. Vientiane: Pak NgumDist., Phang Dang Village, 12.VI.2010, Sk.
Yamane leg., LA10-SKY-098 (THNHM,
SKYC); Naxaythong Dist., Sivilay Village,
9.VI.2010, W. Jaitrong leg., WJT10-LAO-38(THNHM, SKYC).
Distribution in Laos. Vientiane (new record).
Odontomachus simillimus F. Smith, 1858 (Figs
178-179)
Material examined. Vientiane: Hadsaiphong Dist., Thana Village, 6.V.2010, W.
Jaitrong leg., WJT10-LAO-205 (THNHM,
SKYC).
Distribution in Laos. Vientiane (new record).
Odontoponera denticulata (F. Smith, 1858)*(Figs 180-181)
Material examined. Vientiane: Pak NgumDist., Phang Dang Village, 14.VI.2010,09-Dec-16 3:14:01 PM

A Checklist of known ant species of Laos (Hymenoptera: Formicidae)

W. Jaitrong leg. (THNHM); same loc. anddate, Sk. Yamane leg. (SKYC); NaxaythongDist., Sivilay Village, 9.VI.2010, Sk. Yamane leg. (SKYC); some loc., 10.VI.2010,
W. Jaitrong leg. (THNHM); Houi Nyang
near Tha Ngon, 26.VIII.2003, Y. Ochiaileg. (SKYC). Oudomxay: Namo Dist., Mai
Naatan, 26.VIII.2004, Y. Ochiai leg. (THNHM, SKYC). Phong Sali: Khoua, Huei Pee,
1000 m alt., 19.VIII.2004, Y. Ochiai leg.,(THNHM, SKYC); Muang Khoua, 22-24.
VIII.2005, Y. Ochiai leg. (THNHM, SKYC).
Luang Prabang: Kuangsi Waterfall Park,
16.XII.2015, W. Jaitrong leg. (THNHM).
Remarks. Lao specimens (Lat-Hom and Luang Prabang) cited as Odontoponera transversa (F. Smith, 1857) in Santschi (1920b)
should be reidentified as O. denticulata (sensu Yamane 2009).
Distribution in Laos. Vientiane, Oudomxay
Phong Sali and Luang Prabang.
Platythyrea parallela (F. Smith, 1859) (Figs
182-183)
Material examined. Vientiane: Pak NgumDist., Phang Dang Village, 14.VI.2010, W.
Jaitrong leg. (THNHM, SKYC); NaxaythongDist., Sivilay Village, 7.V.2010, W. Jaitrong
leg. (THNHM); same loc., 10.VI.2010, Sk.
Yamane leg. (SKYC).
Distribution in Laos. Vientiane (new record).
Pseudoneoponera rufipes (Jerdon, 1851)* (Figs
174-175)
Material examined. Vientiane: Pak NgumDist., Phang Dang Village, 14.VI.2010, W.
Jaitrong leg., WJT10-LAO-144 (THNHM,
SKYC); same loc. and date, Sk. Yamane leg.
(SKYC); Naxaythong Dist., Sivilay Village,
11.VI.2010, Sk. Yamane leg. (SKYC). Luang Prabang: Santi Resort, 17.XII.2015, W.
Jaitrong leg. (THNHM).
Distribution in Laos. Thatom (Santschi
1920b: 159, cited as Pachycondyla (Bothroponera) rufipes) and Luang Prabang (newrecord).
PROCERATIINAE [1 genus, 1 species]
Probolomyrmex longiscapus Xu et Zeng, 2000
(Figs 184-185)
Material examined. Vientiane: Naxay-

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43thong Dist., Sivilay Village, 11.VI.2010, W.
Jaitrong leg., WJT10-LAO-97 (THNHM);
same loc. and date, Sk. Yamane leg., nest in
soil, LA10-SKY-078 (SKYC).
Distribution in Laos. Vientiane (new record).
PSEUDOMYRMICINAE [1 genus, 6 species]
Tetraponera allaborans (Walker, 1859)* (Figs
186-187)
Material examined. Vientiane: Naxaythong Dist., Sivilay Village, 9.VI.2010, W.
Jaitrong leg., WJT10-LAO-32 (THNHM,
SKYC); same loc. and date, Sk. Yamane leg.
nest in dead twig, LA10-SKY-003 (SKYC);
same loc., 10.VI.2010, W. Jaitrong leg.
(THNHM); Hadsaiphong Dist., Thana Village, 6.V.2010, W. Jaitrong loc. (THNHM);
Houi Nyang near Tha Ngon, 26.VIII.2003,
Y. Ochiai (SKYC).
Distribution in Laos. Bolikhamxay (Ward2001) and Vientiane (new record).
Tetraponera attenuata F. Smith, 1877* (Figs
188-189)
Material examined. Vientiane: Pak NgumDist., Phang Dang Village, 12.VI.2010, Sk.
Yamane leg. (SKYC); same loc., 13.VI.2010,
W. Jaitrong leg. (THNHM).
Distribution in Laos. Bolikhamxay [PhouKhao Khouay N.P.] (Ward 2001) and Vientiane (new record).
Tetraponera binghami (Forel, 1902)* (Figs
190-191)
Material examined. Vientiane: Naxaythong Dist., Sivilay Village, 7.V.2010, W.
Jaitrong leg., WJT10-LAO-227 (THNHM,
SKYC); Hadsaiphong Dist., Dondu Village,
8.V.2010, W. Jaitrong leg., WJT10-LAO-240(THNHM, SKYC).
Distribution in Laos. Luang Prabang(Santschi 1920b: 160, as Sima binghami);
Vientiane (new record).
Tetraponera nigra (Jerdon, 1851) (Figs 192-193)
Material examined. Vientiane: Naxaythong Dist., Sivilay Village, 11.VI.2010, W.
Jaitrong leg., WJT10-LAO-93 (THNHM,
SKYC); same loc. and date, W. Jaitrongleg., WJT10-LAO-123 (THNHM, SKYC);
same loc. and date, Sk. Yamane leg., LA10SKY-085 (SKYC).

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Weeyawat Jaitrong, Benoit Guénard, Evan P. Economo, Nopparat Buddhakala & Seiki Yamane

Distribution in Laos. Vientiane (new record).
Tetraponera nitida (F. Smith, 1860) (Figs 194-195)
Material examined. Vientiane: Pak NgumDist., Phang Dang Village, 14.VI.2010, W.
Jaitrong leg. (THNHM).
Distribution in Laos. Vientiane (new record).
Tetraponera rufonigra (Jerdon, 1851)* (Figs
196-197)
Material examined. Vientiane: NaxaythongDist., Sivilay Village, 10.VI.2010, Sk. Yamane leg. (SKYC); same loc., 11.VI.2010, W.
Jaitrong leg., WJT10-LAO-94 (THNHM,
SKYC). Oudomxay, Namo, Mai Naatan (950m alt.), Y. Ochiai leg. (SKYC).
Distribution in Laos. Champasak (Ward2001: 649); Oudomxay and Vientiane (newrecord).
ACKNOWLEDGEMENTS
We thank Mr. Cholawit Thongcharoenchaikit andMiss Scarlett Rose McGrail (National Science
Museum, Thailand) for reading through an earlier
draft of the manuscript. We also thank Dr. Yukino
Ochiai (Ryukoku University, Japan) and Mr. Haruo Fukuda (Kagoshima-shi, Japan) for providing
us with valuable specimens. Dr. Stefan P. Cover(Museum of Comparative Zoology, Cambridge,
Massachusetts, USA) kindly offered us valuable
records of some ant species from Laos. EPE wassupported by subsidy funding to OIST. Sincere
thanks are also extended to one anonymous reviewer, Dr. James Wetterer (Wilkes Honors College, Florida Atlantic University, USA) and Dr.
Stefan P. Cover for their valuable comments.
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