みちのくべントス
第 8号

2024年 3月

みちのくベントス研究所

みちのくベントス
第８号

２０２４年３月
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Gould, A.A.1859. Descriptions of new species of shells brought home by the North
Pacific Exploring Expedition. Proceedings of the Boston Society of Natural History
7: 138–142.
Š Ȗ Ď ș 1960. A.A. Gould * Ǭ ȅ   ĳ ł ƅ * Ǻ ȷ Ʌ Venus 21(1): 10–31.
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National Museum.
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animals XVI, Reeve Brothers, London.
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Choe, B.L., Park, J.K. (1992) Nine unrecorded mesogastropodous species (Gastropoda: Mollusca) from
Korean waters –Superfamilies Turritellacea, Calyptraeacea, Cypraeacea, and Tonnacea–. Korean
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Becoming a limpet: An ‘intermittent limpetization’ process driven by host features in the
kleptoparasitic gastropod family Capulidae. Molecular Phylogenetics and Evolution 155: 107014.
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English title: Ibuki Miyazaki1, Toyomu Yasunaga2, Hirokazu Abe3, Masahiko Washio3, Tadashi Sugimoto4,
Ryo Nakayama5, Hiroaki Fukumori3 (2024) Records of Neoiphinoe unicarinata (Gastropoda: Capulidae)
from Asamushi, Aomori based on field surveys and from Mutsu Bay based on specimens in the shellcollection at the Asamushi Aquarium.
1

Faculty of Agriculture, Kyoto University; 2 Faculty of Science, Tohoku University; 3 Research Center for
Marine Biology, Graduate School of Life Sciences, Tohoku University; 4 Asamushi Aquarium, Aomori
Prefecture; 5 Fisheries Research Institute, Aomori Prefectural Industrial Technology Research Center 

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Clout MN, Kawamichi M, De Poorter M, Iwatsuki K (eds) Assessment and control of biologicalinvasion risks. Shoukadoh Book Sellers, Kyoto, Japan and IUCN, pp. 104–112.
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Peterson CH (1983) Interactions between two infaunal bivalves, Chione undatella (Sowerby) andProtothaca staminea (Conrad), and two potential enemies, Crepidula onyx Sowerby and Cancer
anthonyi (Rathbun). Journal of Experimental Marine Biology and Ecology 68: 145–158.
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English title: Hiroaki Fukumori1, Hirokazu Abe1, Masahiko Washio1, Ryo Nakayama2 (2024) Records ofan invasive alien species Crepidula onyx (Gastropoda: Calyptraeidae) from Shimokita Peninsula andMutsu Bay with its host gastropod species in Aomori Prefecture, Japan.
1

Research Center for Marine Biology, Graduate School of Life Sciences, Tohoku University; 2 Fisheries
Research Institute, Aomori Prefectural Industrial Technology Research Center 

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6. G i t = g = I Goniobranchus aureopurpureus (Colingwood, 1881)Ⱦ Ý 2Fȿß © ¯ ęɅ¾ ƃ ȗ

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7. I n n < r = g = I Goniobranchus tumuliferus ( C o l l i n g w o o d , 1 8 8 1 )Ⱦ Ý 2 G ȿŲ š Ʌ 1   ɀ 2022 ĝ 9 ś 9 œ ɀ 10%c p f o t Þ ÿ u 70%> N Z v p  ú ß © ¯ ęɅČ Ħ ƬȾ ȴ Ē ȿ À ' œ š ƃ ż Đ ɀĺ ǐ ¿ Ē  À ' ñ Ĝ ƀ ż ĐȾ Ǟ ĩ uĩ Ǯ 1993ȿƞ ® ƙ Ž Ʌ Č Ħ Ƭ Ⱦ Ƞ ş 1979Ɇ š Ʊ ƾ ȿ

8. ; @ = g = I Hypselodoris festiva (A. Adams, 1861)Ⱦ Ý 2Hȿ
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4  ĕ ɀ§ x ĒȾ Ǟ ĩuĩ Ǯ 1993Ɇć Ȟu· Ǯ 2020Ɇũ ĉ uČ ē 2021ȿƞ ® ƙ ŽɅȯ Ŭ ƬȾ Baba 1935ȿɀČ Ħ ƬȾ Ƞ ş 1979Ɇš Ʊ ƾ ȿɀŏ Ǝ ƬȾ Ǥ ť 1969ɆǤ ť u Ŧ 1975bɆ ȷ Ď Ɲ Ɨ Ʊ ƾ  1999ȿ

Ƚ 57 Ƚ

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< d = g = I y ƹ Phylidioidea Rafinesque, 1814
D r I N W I = g = I ƹ Dendrodorididae O'Donoghue, 1924
10. f O n = g = I Dendrodoris fumata (Rüppell & Leuckart, 1830)Ⱦ Ý 2Jȿ
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N T J f = g = I y ƹ Arminoidea Iredale & O'Donoghue, 1923
N T J f = g = I ƹ Arminidae Iredale & O'Donoghue, 1923
11. @ U h = g = I Dermatobranchus otome Baba, 1992 Ⱦ Ý 2Kȿ
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@ @ g Z = g = I y ƹ Aeolidioidea Gray, 1827
m R K J g Z = g = I ƹ Facelinidae Bergh, 1889
12. ; A > n g Z = g = I Sakuraeolis enosimensis (Baba, 1930)Ⱦ Ý 2Lȿß©¯ęɅ¾ƃȗ

4  ĕ ɀ § x Ē Ⱦ Ǟ ĩ u ĩ Ǯ 1993Ɇ · Ǯ 2009ȿƞ ® ƙ Ž Ʌ ȯ Ŭ Ƭ Ⱦ Baba 1957ȿɀ Č Ħ Ƭ Ⱦ Ƞ ş 1979Ɇ ȷ Ď Ɲ Ɨ Ʊ ƾ  1999Ɇ šƱ ƾ ȿɀ ŏ Ǝ Ƭ Ⱦ Ǥ ť 1969Ɇ ȷ Ď Ɲ Ɨ Ʊ ƾ  1999ȿȲ ů ƪ Cephalaspidea P. Fischer, 1883
` V = B < y ƹ Haminoeoidea Pilsbry, 1895
` V = B < ƹ Haminoeidae Pilsbry, 1895
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Y  W qr u& ? : B 0 Apl ysia ku ro daip   R  Y  W qr u Elysia atrovirid isp   R
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; h _ n I ƪ Aplysiida Pelsener, 1906
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14. D r a o ; h _ n I Aplysia japonica G. B. Sowerby II, 1869Ⱦ Ý 3Bȿß©¯ęɅ¾ƃȗ

4  ĕ ɀ § x Ē Ⱦ Ǟ ĩ u ĩ Ǯ 1993Ɇ · Ǯ 2009ȿƞ ® ƙ Ž Ʌ ȯ Ŭ Ƭ Ⱦ Baba 1957Ɇ ȷ Ď Ɲ Ɨ Ʊ ƾ  1999ȿɀ Č Ħ Ƭ Ⱦ Ƞ ş 1979Ɇ ȷĎ Ɲ Ɨ Ʊ ƾ  1999Ɇ š Ʊ ƾ ȿɀ ŏ Ǝ Ƭ Ⱦ Ǥ ť 1969Ɇ Ǥ ť u Ŧ 1975bɆ ȷ Ď Ɲ Ɨ Ʊƾ  1999ȿǜɅš ƻ ' ǩ ħ ì ƥ " ǜ
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Ƚ 59 ȽÚ Ǧ ƪ Sacoglosa Ihering, 1876
P V o g V o B < y ƹ Plakobranchoidea Gray, 1840
P V o g V o B < ƹ Plakobranchidae Gray, 1840
16. D r g V o B < Elysia atroviridis Baba, 1955 Ⱦ Ý 3DȿŲ š Ʌ 3   ɀ 2020 ĝ 7 ś 3 œ ɀ 85%E o M o t Ɔ Ƅ Ų š ß©¯ęɅ¾ƃȗ

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17.  @ U h g V o B < Elysia obtusa Baba, 1938 Ⱦ Ý 3Eȿ
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[ O A i = g = I ƹ Limapontidae Gray, 1847
18. ; o i = g = I Ercolania boodleae (Baba, 1938)Ⱦ Ý 3FȿŲ š Ʌ 1   ɀ 2020 ĝ 7 ś 3 œ ɀ 85%E o M o t Ɔ Ƅ Ų š ß©¯ęɅ¾ƃȗ

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S`U_Baba K (1935) Report of the biological survey of Mutsu Bay (27) Nudibranchia of Mutsu
Bay. Science Reports of the Tohoku Imperial University, Fourth series, Biology
10(2): 331–360, pls. 5–7.
Baba K (1957) A revised list of the species of Opisthobranchia from the northern partof Japan, with some additional descriptions. Journal of the Faculty of Science,
Hokkaido University, series 6, Zoology 13(1–4): 8–14.
Baba K (1993) Two new species of Carminodoris (Nudibranchia: Dorididae) from Japan.
Venus 52(3): 223–234.
ƶ Ɵ þ Ⱦ 2021ȿ Biology and Evolution of the Mollusca ! ņ ×  7  ȍ  º Ɨ ' ¯ȳ  ǆ " Õ Ï ' ą Į ɂ Molluscan Diversity 6(2): 89–180.
Ǟ ĩ  wuĩ Ǯ Ǫ òȾ 1993ȿœ š Ç + Ó ȑ á ã ƞ ȍ  º Ɨ ǐ ƪ ȡ ɂ> p Ȉ ȳ ® Ɩ Ċ ɀ§ ĉ ɂ 693 pp.
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1RFMBgIQYȘĈŪƓ Țʔ!ƸĘ țʔǷǬTh ȚșȘȘ12ŭ­*ŖªǬ ŗőŵŵĄġķİŮżÓ ķİŗÎƄŵǬǲThe northernmost records of four heterobranch sea slugs (Mollusca: Gastropoda) in Miyagi
Prefecture, northeastern Japan.
Mao Kobayashi1, Hiroki Sato2, Hirokazu Abe1

1

Department of Biological Sciences, Faculty of Science and Engineering, Ishinomaki Senshu University
2

Department of Marine Ecosystem Science, Atmosphere and Ocean Research Institute,
The University of Tokyo*

Corresponding author: habe@isenshu-u.ac.jp, abehiro13@gmail.com


During the survey of heterobranch sea slugs in Miyagi Prefecture, northeastern Japan, from November
2023 to February 2024, Aplysia argus Rüppell & Leuckart, 1830 were collected from Moune Bay,Kesennuma City and Ogatsu Bay, Ishinomaki City and A. gigantea Sowerby I, 1869, Platydoris ellioti(Alder & Hancock, 1864), and Pleurobranchus peronii Cuvier, 1804 were collected from Ogatsu Bay. On
the Pacific side of Japan, these four species were previously reported to be distributed south of the Boso
Peninsula, Chiba Prefecture. The records of A. argus, Platydoris ellioti, and Pleurobranchus peronii from
Miyagi Prefecture in the present study updated the northernmost records of these species, but the informal
collection records for A. argus and Pleurobranchus peronii indicate distribution of the two species as far
north as the Oga Peninsula in Akita Prefecture, on the Sea of Japan side of Japan. A. gigantea has been
recently shown to be a senior synonym of A. extraordinaria (Allan, 1932), and the record from Mutsu
Bay, Aomori Prefecture as A. extraordinaria is the northernmost record for this species. The record of A.
gigantea from Miyagi Prefecture in the present study is not the northernmost record of the species, but
extended the distribution record of this species on the Pacific Ocean side of Japan northward. The
specimens of A. argus, A. gigantea, and Pleurobranchus peronii collected in the present study were
deposited in the Natural History Museum and Institute, Chiba (CBM) under specimen numbers CBM-ZM
203011, CBM-ZM 203012, and CBM-ZM 203013, respectively.

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žĴłŀʓɂĕāȴɓȵɓ 1 +"ɃʔĕāŠ] CBM-ZM 203011ʔCBM-ZM 203012ʔCBM-ZM
203013 ȾȯȼPƲũžTőȍʒCBMʓɁYƵȯȶȜ

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ŋĒŧʒɘʄɼʈɦŧʓȘ Order Aplysiida Pelseneer, 1906ɘʄɼʈɦŵȘ Family Aplysiidae Lamarck, 1809
ɘʄɼʈɦ

Ș Genus Aplysia Linnaeus, 1767ɧʆɸʄɘʄɼʈɦ

Aplysia argus Rüppell & Leuckart, 1830ʒo 2 A–Dʓ

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Aplysia gigantea Sowerby I, 1869
ʒo 2 E–Gʓ

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Ș Genus Platydoris Bergh, 1877ɠʅɟɬɚʃɚɦ

Platydoris ellioti (Alder & Hancock, 1864)
ʒo 2HʔIʓ

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Northernmost records of Penaeus (Penaeus) monodon Fabricius, 1798 and Penaeus (Eopenaeus)
semisulcatus De Haan, 1844 from Miyagi and Iwate prefectures, northeastern Japan
Haruto Oda1, Koichi Hatakeyama1, Takuzo Abe2, Shota Suzuki2, Takahiro Akaike3, Hirokazu Abe1

1

Department of Biological Sciences, Faculty of Science and Engineering, Ishinomaki Senshu University
2

Minamisanriku Nature Center
3

Ecoris Inc.

*Corresponding author: habe@isenshu-u.ac.jp, abehiro13@gmail.com


Penaeus (Penaeus) monodon Fabricius, 1798 and Penaeus (Eopenaeus) semisulcatus De Haan, 1844
were firstly recorded from Miyagi Prefecture, northeastern Japan based on specimens collected in
Moune Bay (two females), Nanakita River mouth (two males and one female) and Natori River mouth(one female) from 2–5 November 2023 for the former, and in Mitobe River mouth (one female) on 13
October 2023 for the latter. In addition, P. semisulcatus was firstly recorded from Iwate Prefecture basedon a photograph of an individual caught by a fixed trap net in the coastal area off Ofunato Bay andlanded at the Ofunato Fish Market in 12 October 2023. The northernmost records for P. monodon and P.
semisulcatus were previously reported from Ibaraki and Chiba prefectures, respectively. Therefore, ourrecords updated the northernmost records of P. monodon and P. semisulcatus to Miyagi and Iwate
prefectures, respectively. All specimens of both species collected from Miyagi Prefecture wereimmatures because their sizes less than their biological minimum sizes. These results indicate that all
specimens were spawned in 2023. It would be important to continue monitoring the occurrence of both
species to determine whether they can overwinter and establish a population in Miyagi and Iwate
prefectures. The specimens of P. monodon and P. semisulcatus are deposited in the Natural History
Museum and Institute, Chiba (CBM) and the Minamisanriku Nature Center (MNC) under specimen
numbers CBM-ZC 17790–17795 and MNC-AR-02533, respectively.

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S\oȼ ɿʢʛɺʓƘ Penaeidae Rafinesque, 1815 ɡɺʓȮɠɢŵĦȑǝɟƜɇɉqɧɱʬŬɠɿʢʛɺʓ® Penaeus Fabricius, 1798 ɢnƘɡɜĊɫɉɡĻŵȑǝƜɴqɩHȮǀɜɁɰʪȜʧē

2016ʫȽĒ®ɴqɩĠȹƆɡHȮƃBĥǦɴǙɘɕ Pérez Farfante & Kensleyʪ1997ʫɢʬɓɱɧ
ɜɿʢʛɺʓ®ɠqɧɱɛɂɕ 6 əɡ® Farfantepenaeus Burukovsky, 1997ʬFenneropenaeus
Pérez Farfante, 1969ʬLitopenaeus Pérez Farfante, 1969ʬMarsupenaeus Tirmizi, 1971ʬMelicertus
Rafinesque, 1814ʬPenaeus sensu stricto ɴ®ɠÿġɎɒɕȽȜʧēʪ2016ʫɢʬPérez Farfante &Kensleyʪ1997ʫɇÿġɎɒɕǧɡ 6 ®ɠwoɴìyɏʬɓɡɃɗ Penaeus sensu stricto ɠ¦ɏɛɢɹʄɺʓ®ɝɂɃùɕɟwoɴɄɕȽɏɆɏɟɇɮʬPérez Farfante & Kensleyʪ1997ʫȟʬƎƞǄɠɭɘɛĒ®ɡHȮƃãɂɢſɟɯʬƳɏɕǞǣɇÑɮɱɛɂɟɂŭÝɇƴɂɛɂɕʪYang et al.
2023ʫȽɌɡɭɃɟŒɴeɊʬYang et al.ʪ2023ʫɢʬ7 əɡ DNA ȪɴŶɂɛÄǁɡ Penaeusʪ=ɿʢʛɺʓ®ʫɠqɧɱɰ 32 Ɯ

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îì$1+!3Ýǜ$+͠Ƕņǘĳ ʁ 1 9 3 3 ͡ $ +  . 1

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n'3%UOn.+lf]BG
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8ȁƿ9ǜ2Nǒɘ˹Ǿ8ʩǹ

ɉ͠1927͡«̺͈ȅɶL8ǜɿɯ5u~[r8˫̰

3 ¬ ɬ ˢ # O + ͥ ɢ Ď ̦ Ű Ȩ ū ͠ ͕ ȼ 1 9 5 7 ͡ ò  ɇ ͠ ͧ ò  ȿ ͡͠ ī Ǳ 1 9 5 7 ͡) $ 1 ̬ Ò Ű ȥ ą ͠  ő 1 9 8 3 ͡ 2  N ͠ ĥ 1 ͡ ͕ ȼ ͠ 1 9 5 7 ͡ 9  ɢ Ď ̦ Ű Ȩ ū

Ƚ 82 Ƚ

8Ȟɢ2ʍ6˝řRN˙8ȎǱRÍƺ$
̖²&NTdu̾ņɁL˫̰MƒƱɯ6[r3ț̏$1âɤ̦Ń6ȸʳɅ8N!33

Ãʰ6

3̯ȗ8NɎLu~[r3čŔ$+ǃ̓ƀǩǘIǃ̓ĪɎ8˰ʦ9ǜ
2MˬƿȎǱGĥɿ#O15

ŇƐ2N+F̨͠˫8čŔȁƿ9ʌ8˾ÞɎ3$1

1936ͦSakai 1939͡ čŔ8½͎Ƨ9͘3N5

˓ʻ8͕ȼ͉łȜƐǡƹŧ$1+̉ɂʉŐɼʐƹ9)8ƗĦʑʉŐøɔ͔

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čƀ7ǩ30ǘ8òɇ8˹Ǿ6İ0u~[rRđF+Ǳɲ̰9ęĎ$˫̐5ŝţ͠1916͡ǈĝ$+H. tridens3Cha. convexusR)O*Ou~[r3u~[rq\3&NęĎ´ʟ9

ɉ͠1927͡8Øɓ«̺AB·ɡ#O15

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54ͪ

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Ǳʌ8˫̐IˬƿȎǱ8ĥɿ95

)8čŔ8ɸÄRɽFN&@95  ő ͠ 1 9 8 3 : 1 8 6 ͡ 9  ý ˇ ǘ Ǳ Ŀ Ĭ ɤ ș ͐ ĥ ̱ ͠ I I ͡ 8 u ~ [ r 8 ˦ ˶ 8
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3ȎǱ9͂͠KMNH IvR 100024͡3Ƽû̈́͠KMNH IvR 100025͡Ċ1À´

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102¬8ǒɘRʭˣ$+ʩǹR˜16ɿ&Ǳʌ8ð̻Ùŷ6/1ĿÞ&N3
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Ǳʌ8Ùŷð̻9͈ȅ3#ON!3Ľ.+͠14¬9¬͡ 1930ƀªɱKMZr͐Ù͐Ő8

Ƚ 83 Ƚ

< Q   0  QTRS   I l

1^scU|}v{mBE\Yr'3?VHNleZgnUarn*qZ1mptU<M\`hq##JleZg

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No.

ǒɘ

Ùŷð̻8ƻʜ

1
2
3
4
5
6
7
8
9
10
11
12
13
14
15
16
17
18
19
20
21

 ɉ ͠ 1927: 1118 ͡͞ ɢ ͠ 1931: 134 ͡ ɉ ͠ 1931: 804 ͡ȱ ̬ ͠ 1933: 557 ͡Ƕ£øɔŐɼʐ®͠1933: 227͡Ȑ Ŧ ͠ 1933: 174 ͡ǘǱëɔɼʐŐ®͠1934: 558͡Š ̬ ɢ  Š ̬ ɢ ͠ 1936: 51 ͡
̨ ͠ 1936: 233 ͡Š ̬ ɢ  Ǵ ̊ ͠ 1937: 219 ͡
Sakai (1939: 691)
Sakai (1940: 51 )
Š ̬ ɢ ͠ 1943: 657 ͡ ɉ  ̨ ͠ 1947: 652 ̨͡ ͩ 1956: 59 ͠ ɲ ̰ ͪ͡Ò ȶ ͠ 1956: 90 ͡
̨ ͠ 1957: 85 ͡
̨ ͠ 1958: 154 ͡ɢ  ͠ 1962: 96 ͡
̨ ͠ 1965a: 720 ͡
Sakai (1965b: 205 )22 ő ͠ 1970: 293 ͡

23
24
25
26
27
28
29
30

͕ ȼ L ͠ 1971: 127 ͡ˏ ̈ L ͠ 1972: 223 ͡
Baba & Fukuda (1972: 90 )
Ò ɢ ͠ 1972: 511 ͡
ſ Ǳ ͠ 1973: 65 ͡
³ Ȫ ͠ 1974: 71 ͡ː ̬ ͠ 1974: 95 ͡
Ȗ ɢ ͠ 1975: 289 ͡

31Sakai [1976: 415 (in Japanese);
668 (in English)]

32
33
34
35
36
37
38
39
40
41
42
43
44
45
46
47
48

͕ ȼ  ʍ ˒ ͠ 1982: 127 ͡Ǵ ũ ͠ 1982: 87 ͡Ȗ ɢ ͠ 1982: 222 ͡ ő ͠ 1983: 186 ͡
Nakasone et al. (1983: 37)Ȗ ɢ ͠ 1983: 289 ͡ˏ ̈ L ͠ 1984: 215 ͡Ȗ ɢ ͠ 1985: 171 ͡
̨ L ͠ 1987: 246 ̮͡ ǯ ͠ 1987: 265 ͡Ȗ ɢ ͠ 1990: 287 ͡Ũ ą  ͕ Ĵ ͠ 1993: 488 ͡Ǫ Ũ ͠ 1994: 118 ͡Ȗ ɢ ͠ 1994a: 263͡Ȗ ɢ ͠ 1994b: 217 ͡Ǵ ũ  Š ɢ ý ͠ 1995: 49 ͡ę ɢ ͠ 1995: 410 ͡

͈ȅ
͈ȅ
͈ȅ
͈ȅ
͈ȅ
ǱűöǶ̦
͈ȅ
͈ȅ
Ƹű
͈ȅ
“Bôsyû (Tateyama Bay)”“Tokyo B ay”͈ȅƸű«ö
Ƹű«ö
Ƹű«ö
Ƹʭ«ö
Ƹʭ«ö
Ƹʭ«ö
Ƹʭ«ö
“Sagami Bay”
Ǳű͠CO6͈ȅɶƸű͔ŨɁ«ö͡ó˒ɶ«öƸű«ö
“Sagami Bay”Ƹʭ«ö
ɴȏɁƸű͔͠Ũ͡«öǱű̦
Ƹʭôŭ«öǶ£Ɂ«ö…ǘǱȶȨū29ʅ ɶȨūͦ“Sagami Bay…on the continental side, from
Fukui”
ó˒ɶ«öǶ£ɁLöƸʭôŭ«ö͈ȅɶ“Aomori”Ƹʭôŭ«öƸű«ö
Ƹʭôŭ«öɴȏɁǶ£Ɂ«öƸʭôŭ«öƸʭôŭ«ö͈ȅɶ͢Ƹʭôŭʅ ɶ«öƸʭôŭ«ö͈ȅɶ«öƸʭôŭ«ö͈ȅɶ«öȽ 84 Ƚ

Ùŷ8Ñƿ3$1Ƌɡ#O+Ǳ˜8¨ǒɘ͠ǜɽ5Ƌɡ6̻N͡

21

33
31

34

35

49
50
51
52
53
54

ę ɢ ͠ 1996: 77 ͡
ʅ ͠ 1998: 47 ͡
Ƿ  ¼ ͠ 1999: 166 ͡ǭ ¿ ͠ 2000a: 335͡ǭ ¿ ͠ 2000b: 372 ͡Š Ǹ ͠ 2000: 122 ͡55

Ŭ Ȟ ͠ 2000: 300 ͡

56
57
58
59
60
61
62
63
64
65
66

ņ ̀  Ȗ ɢ ͠ 2000: 28 ̨͡  ʛ Ř ͠ 2001: 191 ͡ɭ ɋ ͠ 2001: 89 ͡ˏ ̈ L ͠ 2002: 215 ͡ǭ ¿ ͠ 2002: 285 ͡ ͠ 2002: 67 ͌͡ ̈ ͠ 2002: 224 ̮͡ ǯ ͠ 2002: 336 ͡Ǖ ̬ ͠ 2004: 188 ͡Ȗ ɢ ͠ 2004: 198 ͡Ȑ ĵ ͠ 2004: 135 ͡

67

Sakai et al. (2006: 13)68ǭ ¿  ȅ  ͠ 2007: 363 ͡69

Ǘ ͠ 2007: 13 ͡

70
71
72
73
74
75
76
77
78
79
80
81

ņ ̀  Ȗ ɢ ͠ 2008: 32 ͡ņ ̀  Ȗ ɢ ͠ 2010: 12 ͡ǭ ¿ ͠ 2011: 387 ͡Ȑ ĵ ͠ 2011: 135 ͡ǭ ¿ ͠ 2012: 79 ̮͡ ǯ ͠ 2012: 200 ͡Ŀ ȅ ͠ 2013: 146 ͡ů ɢ   ť ͠ 2013: 179 ͡ɭ Ũ L ͠ 2013: 126 ͡ç ˗ ͠ 2014: 97 ͡
Ȅ ý ͠ 2014: 233 ͡
Ŀ ȅ ͠ 2014: 287 ͡82́ ɢ  ̶ ͠ 2014: 206 ͡

83
84
85

ȼ ̦ ͠ 2014a: 113͡ȼ ̦ ͠ 2014b: 72 ͡
ˏ ̈ L ͠ 2016: 215 ͡

86Ǫ Č ͠ 2016: 54 ͡

87
88
89
90
91
92
93
94

ȅ ̬  ȃ ý ͠ 2016: 313 ͡Ȗ ɢ ͠ 2016: 44 ͡þ ŭ L ͠ 2017: I_1173 ͡ɧ ɢ ͠ 2018: 126 ͡Č Ů ͠ 2018: 125 ͡Ǽ ý  ̮ ǯ ͠ 2018: 49 ̟͡ ˢ ͠ 2019: 530 ͡Ȗ ɢ ͠ 2019: 38 ͡95

́ ɢ  ̶ ͠ 2019: 271 ͡

96
97
98
99
100
101
102

Ǡ Ǻ ͠ 2020: 160 ͡ȼ ̦ ͠ 2020: 733 ͡Ŀ ɉ ͠ 2022: 39 ͡Č Ů ͠ 2022: 155 ͡ę ɢ ͠ 2022: 451 ̭͡ ̀ L ͠ 2023: 46 ͡
Ƿ ţ ͠ 2023: 264 ͡Řĭɶ«öŘĭɶK<ʅ ɶƸʭôŭ«ö
͈ȅɶ
Ƕ£Ɂ
͈ȅɶ«öǶ£Ɂ«ö͠ŁſȭÃ͢͡ʅ ɶ«ö͠ǘǱȶÃ͡Ƹʭôŭ«öɴȏɁ…ʅ ŘĭɶƸű«ö͈ȅɶǶ£Ɂ«ö͈ȅɶ«öǶ£ɁǶ£Ɂ ʅ ɶ«öƸʭôŭ«öƸʭôŭ«ö
“Sagami Bay…on the Japan Sea sidesouthwards from Fukui prefecture”
͈ȅ͈ȅʲ…ĩǘǱȶÃ͢ɣʅ ʲ«öƸʭôŭ«öƸʭôŭ«ö͈ȅɶƸʭôŭ«öǶðŘĭɶ͈ȅɶ«öǶ£Ɂʅ ɶ«öó˒ɶ«öŘĭɶŘĭɶ͈ȅɶ«ö͈ȅɶ«ö8ŁſȭÃ ʅ ɶ«ö8ǘǱȶÃŘĭɶ«öŘĭƸű«ö
Ƕ£Ɂ…ǘǱȶÃ29ʅ ɶȨū6Ùŷ͈ȅƸʭôŭ«ö ̶ Ƕ « ö ͦ ɴ ȏ Ɂ « ö 
Řĭɶ
Řĭɶ
ǶðĪǖŁſȭÃ8ŘĭɶȨūĮŘĭɶƸʭôŭ«ö Ř ĭ ɶ ͠ Ł ſ ȭ Ã ͡ ͈ ȅ ɶ͠ ǘ Ǳ ȶ Ã ͡Řĭɶ«ö͈ȅɶ«öŘĭɶʅ ɶŘĭɶŘĭɶ«öŘĭɶŁſȭū28Ùŷð̻Řĭ͠ŁſȭÃ͡͈ȅ͠ǘǱȶÃ͡

Ƚ 85 Ƚ

25

28

49

28

67

48

35

25

75

35

75

35

75

67

75

75

68

ĿŚ2N̨ƩȜ͍˥Rɚ$̨͠ó˒3#O1+ɕ6

1987͡ Ȝ8Øɓɔ29̆$1ð̻9

ƶƗ6Øɓ#O+Ǌ˨Ĺ˞ǘǱëɔẖ̂6

1

Ù ŷ   Ƹ ű « ö ͠  ɉ  ̨1947͡6ĻǦ#O+Ɏ̂ƞɯ2N

C+

1960ƀªƗôKMZr͐8Ù͐ŐɯɼʐR̴ň$ɚĩ6˅NC2ẖ̂͐

8ɰ¾RđEĽǑ8ǒɘRØɓ$1NȖɢȕÁȜ͠Komatsu et al. 2012͡GC+

1¬8¸ļ͠Ȗɢ 1994b͡R̽$1+˹Ǿ$+ǒɘ8ùF+ʾǣ69

̆$1Ǳʌ8Ùŷð̻Ró˒3Ùŷð̻ó˒3#O+G8ôǑ«R

˫8K5ƬǆŜ#ONǖ

8Ùŷð̻RŘĭ3&NǒɘɚOňF

69ƐǡC,Ȱ2.+

ʌ8ð̻Ró˒3&Nẖ̂͐8˫̗6Ş$1
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1990ƀªɱKMǱʌ
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ª « ̺ 9 ʡ 4 ä R ù F N K  6 5 . + ͠ 3 2 ¬  1 4 ¬ ͡ ! 8 Ǯ ̵ 6 Ø ɓ # O + ǒ ɘÙŷ8Ñƿ3$1Ƌɡ$1N¨ǒɘLǆŜ&N3!8Ļï69žɇ

8ʪɆúƭëɔẖ̂6N̮ǯ͠2012͡8˫̗ƕ͊$1NĆʿƧN
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ð̻R͈ȅ3&Nǒɘ9ɍ65.1

u~[r8Ùŷ

)82ő͠1970: 293͡ ð ̻ R  C O 6 ͈ ȅ ɶ  3 ˫ ̗ $ 1  + 8 9 ȫ ɲ # O N ͠ ˜ 1 ͡  ő ͠ 1 9 7 0 ͡
61Ǳ˫̗8ȁƿ9ǜL6#O15ǃ̓ƀǩǘL!O9

˫ 8 ̬ Ò Ű ɠ Ȏ Ǳ 6 İ 0  1  + ! 3  Q  N  ͕ ȼ ͠ 1957 ͡  K < ī Ǳ
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͕͠ȼ 1957ͦīǱ 1957ͦő 1983͡ ʡ70ƀRʨ+ɚĩG͈ȅɶÒ6Ǳʌɟƪ$1NĐ8ɽ˲9čƹɯ6Øɚ$¦Ɨ8˸͏2N36$:$:u~[r3ɶÒÀ´ʵ)O*OʪɆ8)O8NĪĮÀ´ʵ̾͠ņɁ ͡͠ ̮ ǯ 2 0 1 2 ͡  K <  ̫ ˡ Ÿ š ̬ ɟ ɟ ɔ ͠ Ȗ ɢ 2 0 2 0 ͡ 6 ̤ Ŕ # O 1  N
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ɟ ƪ  ɽ ˲ # O 5  . + ͠ Ǽ ý  ̮ ǯ 2 0 2 1 ͡  ǖ
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