2004

Asiatic Herpetological ResearchVol. 10, pp. 129-150

Recent Records of Turtles and Tortoises from
Laos, Cambodia, and Vietnam
BRYAN L. STUART1,2,3,4,* AND STEVEN G. PLATT4,5
1Field Museum of Natural History, Department of Zoology, Division of Amphibians & Reptiles,

1400 S. Lake Shore Drive, Chicago, Illinois 60605 USA. E-mail: bstuart@fieldmuseum.org*Corresponding author and address
2University of Illinois at Chicago, Department of Biological Sciences (M/C 066),
845 W. Taylor, Chicago, Illinois 60607 USA
3Wildlife Conservation Society, P.O Box 6712, Vientiane, Lao PDR
4Wildlife Conservation Society, P.O. Box 1620, Phnom Penh, Cambodia
5Present address: Department of Math and Science, Oglala Lakota College,
P.O. Box 490, Kyle, South Dakota 57752-0490 USA
Abstract. - The chelonian fauna of Laos, Cambodia, and Vietnam remains poorly known and is currently threatened
by widespread and intensive exploitation for food and traditional Chinese medicine. The distributions of many species
are uncertain owing to a paucity of records. Because turtles are so extensively traded in the region, most records now
come from animals in trade. We emphasize that authors must be explicit about how their records were obtained toallow other workers the ability to critically evaluate the accuracy of the distribution record. We here present detailed
information on recent (1993-2002), vouchered records of 19 species of freshwater turtles, tortoises, and marine turtles collected in the field or obtained from hunters, abandoned hunting camps, villages, or markets in Laos, Cambodia,
and Vietnam.
Key words. - Testudines, turtles, tortoises, Laos, Cambodia, Vietnam, distribution.

Introduction
Although mainland Southeast Asia has long been
regarded as a hotspot of chelonian diversity (van Dijk etal., 2000), the turtle and tortoise fauna of Laos,
Cambodia, and Vietnam (formerly known as French
Indochina) remains poorly known. Biological investigation was limited prior to World War II, and since thendecades of civil unrest, political instability, and military
conflict have largely prevented fieldwork. Consequently
few museum records exist (summarized by Iverson,1992) and with the exceptions of Smith (1931) and
Bourret (1941), little information is available on theoccurrence and distribution of chelonians in former
French Indochina. A recent photographic identification
guide to the region (Stuart et al., 2001) and country
reviews of Laos (Stuart and Timmins, 2000), Cambodia(Touch et al., 2000), and Vietnam (Hendrie, 2000) summarized information on chelonian distributions, but provided few details concerning new records on whichthese accounts are based.
Exploitation of chelonians for food and medicinal
markets is widespread in Laos, Cambodia and Vietnam(Jenkins, 1995; Le Dien Duc and Broad, 1995; Lehr,
1997; Timmins and Khounboline, 1999; van Dijk et al.,
2000; Ziegler, 2002; Holloway 2003). Hunters in ruralvillages capture turtles and tortoises for local consump© 2004 by Asiatic Herpetological Researchtion or to sell to traders who periodically visit villages topurchase wildlife. Although turtles and tortoises are
locally consumed and domestically traded in Laos,
Cambodia, and Vietnam, most are exported to markets
in southern China (Stuart et al., 2000; van Dijk et al.,2000). Chelonians from Laos and Cambodia are usuallytransported to Vietnam, where they join with
Vietnamese turtles on northward routes to China (Stuart
et al., 2000). Because Laos and Cambodia are source
rather than destination or transfer countries, specimens
obtained from markets in Laos or Cambodia usuallyoriginated from that country. However, trade specimens
in Vietnam may have originated from Vietnam, Laos,
Cambodia, or beyond. The volume of this trade isbelieved to pose a serious threat to the continued viability of wild chelonian populations throughout Southeast
Asia (van Dijk et al., 2000).
Because turtles and tortoises are extensively andvisibly traded in Southeast Asia, most recent distribution
records are based on animals observed in trade rather
than collected from the wild. The geographic origin ofmany trade specimens can be difficult if not impossible
to determine, especially those obtained in urban markets. Uncritical acceptance of these records by workers
has led to inaccurate characterization of species distributions, with serious biological, conservation, legal, andregulatory implications. Additional confusion has result-Vol. 10, p. 130

Asiatic Herpetological Researched from the realization that some species of Asian turtles
described during the last two decades were based ontype specimens obtained from Hong Kong animal dealers who provided inaccurate or fabricated locality data,
leaving the geographic origin of many in doubt (Dalton,
2003; Parham et al., 2001). Distribution records that
explicitly state how the turtles were obtained are therefore clearly important, given the historical paucity of
information, uncertainties in recent literature, and theserious conservation threats faced by these taxa.
We here report recent distribution records of chelonians from Laos, Cambodia, and Vietnam that can beverified with voucher specimens or photographs. These
records were obtained by (1) us during herpetologicalsurveys conducted from February 1998 through May
2001, (2) other workers in the region between 1993 andJune 2002, and provided to us, or (3) other workers who
deposited specimens at the Field Museum of Natural
History, Chicago, USA since 1993. For each record wenote whether the specimen was collected in the field,
found in abandoned hunting camps, obtained fromhunters or residents in rural villages, or purchased frommarkets.
It should be emphasized that our collecting activities had little if any detrimental impact on populations ofwild chelonians. The number of collected specimens on
which we report is insignificant when compared to themillions of chelonians annually consumed by the
wildlife markets of southern China (Lau et al., 2000).
Furthermore, the majority of specimens we and others
collected in the field were shells of animals consumed
by rural villagers. Chelonian shells are commonplaceand easily obtained in villages; shells are retained byhunters as trophies, sold or kept for medicinal purposes,
and used as food containers for domestic animals andrice scoops. Our collecting activities certainly provided
no stimulus for the additional harvesting of wild chelonians. Finally, we believe that further scientific collecting is warranted in the region, as most species remain
under-represented in museum collections, and taxonomic study can affect conservation priorities (Parham andShi, 2001; Stuart and Thorbjarnarson, 2003).

Methods and Conventions
Measurements were taken to the nearest 0.1 cm with 80
cm sliding calipers. We use the following abbreviations:
CL = maximum straight carapace length, including
spines or projections (i.e. not necessarily the mid-line);
CW = maximum carapace width, including spines or
projections; PL = maximum plastron length, including
spines or projections; BD = maximum depth of complete
specimen (head and neck extended in trionychid specimens). Measurements were not reported if shell damage
precluded accurate measurement.

2004

Records are presented under each species account in
the following format where applicable: Field Museum of
Natural History (FMNH) or figure number, type ofrecord, measurements (defined above), locality including coordinates if available, approximate elevation andbrief habitat description if field-collected, circumstancesof origin if not field-collected by collector or photographer, date specimen was collected or photographed, andname of collector or photographer. In the case of records
obtained from hunters or villages, the name of collector
refers to the person who preserved the specimen or provided the record to us rather than to the name of the person who actually captured it from the wild. In the samecases, the date of collection refers to the date the collector (as previously defined) obtained the specimen or
took the photograph rather than to the date it wasremoved from the wild.
GPS coordinates are presented only if the originalcollector provided them, and in the same format as originally provided. Coordinates that we generated for the
purposes of mapping the records are not presented.
Marine turtle records are not mapped.
Species Accounts
Platysternidae
Platysternon megacephalum Gray, 1831
[Map 1]
Laos. - Fig. 1, photograph only, Huaphahn Province,
Vieng Tong District, Ban Sa Kok Village, 20°11' N
103°12' E, captured by resident of Ban Sa Kok, 29 April
1998, B. L. Stuart. FMNH 258749, complete specimen,
CL = 9.3, CW = 7.3, PL = 6.9, BD = 2.6, Bolikhamxay
Province, Khamkeut District, Nape border area, stream
in wet evergreen forest, 19 March 1997, D. Davenport.
Fig. 2 (and Fig. 4 in Stuart and Timmins, 2000), photograph only, Khammouan Province, Nakai District,Nakai-Nam Theun National Biodiversity Conservation
Area, Ban Xiangthong Village, 17° 54' 05'' N 105° 23'
50'' E, one of eleven individuals in the possession of aVietnamese trader leaving Ban Xiangthong, 17
November 1998, B. L. Stuart. Fig. 3, photograph only,
Xe Kong Province, Dakchung District, Ban Daklan
Village, 15° 21.61' N 107° 01.70' E, captured by residents of Ban Daklan, D. Showler, December 1997.
Vietnam. - FMNH 252164, complete specimen, CL =15.5, CW = 11.6, PL =12.7, BD = 5.2, Gia-Lai Province,
Ankhe District, Buon Loi Village, 20 km northwest of
Kannack town, Annamite Mountains, 14° 20' N 108° 36'
E, 700-750 m, found in burrow under overhanging
stream bank, 31 March 1995, I. Darevsky and N. L.
Orlov.

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Asiatic Herpetological Research

Remarks. - Ziegler (2002) reported the species in local
trade in Ha Tinh Province, Vietnam.
Geoemydidae
Batagur baska (Gray, 1831 “1830-35”)[Map 2]
Cambodia. - Platt et al. (2003) reviewed the status of B.
baska in Cambodia and reported on a breeding population in the Sre Ambel River System of Koh Kong
Province.
Cuora amboinensis (Daudin, 1802)[Map 3]
Cambodia. - Fig. 4, photograph only, Battambang
Province, Ek Phnom District, Koh Chivang Commune,
Prek Toal Village on Tonle Sap Lake, 13°14' 28'' N 103°
39' 32'' E, captured by residents of Prek Toal, 27 August
1999, B. L. Stuart, J. Smith, and K. Davey. FMNH
259411, broken carapace and plastron, Kandal Province,
Trayo Village, 11° 19' 02'' N 105°09' 47'' E, obtained
from hunter in Trayo, 05 July 2000, S. G. Platt. Fig. 5,
photograph only, two living turtles, CL = 19.3 and PL =17.5, CL = 20.6 and PL = 18.9, Kampong Thom
Province, Sary Village, 12° 48.48' N 104° 44.19' E, collected by Sary residents in Tonle Sap, 21 June 2000, S.
G. Platt, Heng Sovannara, and Long Kheng. Fig. 6, photograph only, CL = 20.4 cm, PL = 18.5, Koh Kong
Province, Sre Ambel Town, in house of wildlife trader,
11° 07.30' N; 103° 44.73' E, 27 August 2000, S. G. Platt,
B. L. Stuart, and Vuthy Monyrath. Fig. 7, photographonly, CL = 11.6, PL = 11.0, Koh Kong Province, KohKong Town near municipal airport, 11° 37.11' N; 103°
00.98' E, crossing road in open grassland bordered by
Melaleuca and Rhizophora swamp, 07 February 2001,
S. G. Platt, Heng Sovannara, and Long Kheng.
Laos. - FMNH 255262, complete specimen, CL = 16.7,
CW = 11.7, PL = 15.2, BD = 7.5, Champasak Province,
Mounlapamok District, Dong Khanthung Proposed
National Biodiversity Conservation Area, Ban Khiem
Village, 14°14' N 105° 20' E, captured by residents of
Ban Khiem for food, 24 July 1998, B. L. Stuart.
Vietnam. - Fig. 8, photograph only, Kien Giang
Province, An Minh District, photographed in a reptile
trade shop, 09° 45' 04'' N 104° 59' 35'' E, 31 October
2000, B. L. Stuart.
Cuora galbinifrons Bourret, 1939
[Map 4]
Laos. - FMNH 256544, complete specimen, CL = 16.6,
CW = 12.2, PL = 16.2, BD = 8.0, Khammouan Province,Vol. 10, p. 131

Nakai District, Nakai-Nam Theun National Biodiversity
Conservation Area, 17° 50' N 105° 35' E, 900 m, wetevergreen forest, found under brush in leaf litter on hillside 500 m from nearest stream, 13 December 1998, B.
L. Stuart. FMNH 255273, carapace only, CL = 18.1,
CW = 12.9, Khammouan Province, Yommalat District,
Khammouan Limestone (= Phou Hin Poun) National
Biodiversity Conservation Area, Ban That Mouang Khai
Village, 17° 32' N 105° 04' E, consumed by residents of
Ban That Mouang Khai, 01 April 1998, B. L. Stuart.
Vietnam. - FMNH 255694, complete specimen, CL =19.2, CW = 13.0, PL = 18.0, BD = 8.5, Nghe An
Province, Tuong Duong District, Pu Mat Nature
Reserve, 19° 03' N 104° 37' E, 600 m, wet evergreen hill
forest, in leaf litter along Khe Mat Stream, 14 September
1998, B. L. Stuart. FMNH 255695, complete specimen,
repatriated to Vietnam before measurements could betaken, collecting information same as FMNH 255694.
Remarks. - The subspecies C. galbinifrons galbinifrons
Bourret, 1939 is treated here as a full species following
the recommendation of Stuart and Parham (2004).
Ziegler (2002) reported the species in local trade in Ha
Tinh Province, Vietnam, and Fritz et al. (2002) reported
hybrids of C. galbinifrons and C. bourreti in local tradein Ha Tinh and Quang Binh Provinces, Vietnam.
Cuora mouhotii (Gray, 1862)[Map 5]
Laos. - FMNH 258880, carapace only, CL = 16.1, CW= 11.0, Khammouan Province, Boualapha District, Hin
Nam No National Biodiversity Conservation Area, Ban
Tasang Village, eaten by residents of Ban Tasang, 30
December 1995, R. J. Timmins. FMNH 258881, carapace only, CL = 17.3, CW = 13.0, collecting information
same as FMNH 258880. FMNH 258887, plastron only,
PL = 16.8, Bolikhamxay Province, Nam Kading
National Biodiversity Conservation Area, eaten by villagers living in Nam Kading, 01 May 1995, R. J.
Timmins.
Remarks. - The species mouhotii was previously placedin the monotypic genus Pyxidea Gray, 1863, but we allocate it to the genus Cuora following Honda et al. (2002)
and Stuart and Parham (2004). Ziegler (2002) reported
the species in local trade in Ha Tinh Province, Vietnam.
Cyclemys atripons Iverson and McCord, 1997
[Map 6]
Cambodia. - FMNH 259050, complete specimen, CL =21.4, CW = 16.2, PL = 20.7, BD = 8.4, Mondolkiri
Province, Pichrada District, Phnom Nam Lyr Wildlife

Vol. 10, p. 132

Asiatic Herpetological Research

Sanctuary, near 12° 32' 16'' N 107° 32' 00'' E, 600-700
m, evergreen gallery forest, found on sand bank at baseof large boulder 1.5 m from swift, shallow stream, 21
June 2000, B. L. Stuart. FMNH 259051, complete specimen, CL = 22.7, CW = 17.0, PL = 21.2, BD = 8.0, KohKong Province, Sre Ambel District, Sre Ambel Town,
11° 07' 20'' N 103° 44' 45'' E, obtained from turtle trader who reported specimen came from Sophat Village,
downstream from Sre Ambel Town, 27 August 2000, B.
L. Stuart and S. G. Platt. FMNH 259052, complete specimen, CL = 18.0, CW = 15.0, PL =16.7, BD = 6.2, collecting information same as FMNH 259051. FMNH
259412, carapace and incomplete plastron, CL = 19.3,
CW = 15.2, Koh Kong Province, Sre Ambel District,Boeung Tradok Pong Village, 11° 31' 10'' N 103° 46' 55''
E, obtained from hunter in Boeung Tradok Pong, 24
August 2000, B. L. Stuart and S. G. Platt. FMNH
259414, plastron only, PL = 16.5, collecting information
same as FMNH 259412. FMNH 259415, plastron only,
PL = 16.2, collecting information same as FMNH
259412. FMNH 259416, plastron only, PL = 14.9, KohKong Province, Sre Ambel District, Chaouethail Plous
Village on Sre Ambel River, 11° 18' 03' N, 103° 44'
56”E, obtained from hunter in Chaouethail Plous, 21
August 2000, B. L. Stuart and S. G. Platt. FMNH
259417, plastron only, PL = 19.2, collecting information
same as FMNH 259416. FMNH 259422, plastron only,
PL = 20.2, Koh Kong Province, Sre Ambel District,
Chay Reap Village, west bank of Sre Ambel River, 11°
29' 10'' N 103° 47' 00'' E, <10 m, obtained from hunterin Chay Reap, 23 August 2000, B. L. Stuart and S. G.
Platt. Fig. 9, photograph only, two living animals, CL =13.7 cm and PL = 13.0 cm, CL = 20.8 cm and PL = 19.6
cm, Koh Kong Province, Kaoh Pao River, 11° 44.46' N;
103° 04.80' E, surrounding hills covered in dense evergreen forest with some mangrove along shoreline,
obtained from fishermen, taken in crab traps set in river,
10 May 2001, S. G. Platt, Heng Sovannara, and Long
Kheng.
Remarks. - Fritz and Ziegler (1999) reviewed records ofCyclemys from the region. Species boundaries within
the genus Cyclemys remain uncertain (Fritz and Ziegler,
1999; Guicking et al., 2002). The specimens weassigned to C. atripons have plastra that are largely yellow with densely pigmented bridges; complete specimens exhibit nearly immaculate chins. These characteristics are typical of both C. atripons and C. pulchristriata Fritz, Gaulke & Lehr, 1997, two species that were
described almost concurrently in 1997. Cyclemys
atripons and C. pulchristriata have been considered thesame taxon (Iverson in Guicking et al., 2002). However,Fritz et al. (2001) concluded that C. atripons has more
ventral neck stripes (7-8 light and 7-9 dark stripes when

2004

counted from one mouth corner to the other) than C. pulchristriata (5-7 light and 5-7 dark stripes). FMNH
259050 has 8 dark and 7 light ventral neck stripes,
FMNH 259051 has 10 dark and 9 light ventral neck
stripes, but in FMNH 259052 the ventral side of the neckis nearly immaculate like the chin and completely lacks
striping. These few samples demonstrate that ventral
neck stripes are more variable than stated by Fritz et al.
(2001). In a phylogenetic analysis of a 982 bp fragmentof the mitochondrial cytochrome b gene, Guicking et al.
(2002) recovered two clades in the atripons-pulchristriata complex that differed by up to 4.5% sequence divergence. Samples referred to atripons and pulchristriata
appeared in both clades, but the authors assigned thesenames according to whether the sample originated from
Cambodia (atripons) or Vietnam (pulchristriata), rather
than based on their morphology. The findings of
Guicking et al. (2002) suggest that more than one
species of Cyclemys with mostly yellow plastra, densely
pigmented bridges, and immaculate chins could exist,
but it remains unclear whether the two clades correspond to what have been described as atripons and pulchristriata. We assign the name C. atripons rather thanC. pulchristriata to our samples because the type locality of C. atripons is geographically closer to most of oursamples than to that of C. pulchristriata. Clearly, further
studies into the morphological and genetic variation in
Cyclemys are warranted, particularly with samples ofcertain provenance.
Cyclemys tcheponensis (Bourret, 1939)[Map 7]
Laos. - Fig. 10 (and Fig. 7d in Stuart et al., 2001), photograph only, Bolikhamxay Province, Thaphabat
District, Phou Khao Khouay National Biodiversity
Conservation Area, near That Xay Waterfall, 18° 27' N
103° 10' E, 300 m, dry evergreen forest mixed with bamboo, sleeping on bottom of 4 x 4 m pool in forested
stream, 26 June 1998, B. L. Stuart. FMNH 258870,
complete specimen, CL = 9.7, CW = 8.0, PL = 8.7, BD= 3.6, Bolikhamxay Province, Khamkeut District, purchased in Lac Xao Market, 14 December 1996, D.
Davenport. FMNH 258871, complete specimen, CL =9.0, CW = 8.1, PL = 8.3, BD = 3.6, collecting information same as FMNH 258870. FMNH 258875, complete
specimen, CL = 20.6, CW = 15.2, PL = 20.3, BD = 8.5,
Khammouan Province, Nakai District, Houay Moey
Stream (tributary of Nam Pheo River), Ban Na Meo
Village, dry evergreen forest, 07 March 1997, D.
Davenport and J. Chamberlain. FMNH 255263, complete specimen, CL = 17.1, CW = 13.6, PL = 16.1, BD =6.5, Khammouan Province, Nakai District, Khammouan
Limestone (= Phou Hin Poun) National Biodiversity

2004

Asiatic Herpetological ResearchConservation Area, 17° 53' N 104° 52' E, 570 m, dryevergreen forest mixed with deciduous trees and pine,caught on streambank by hunter using dog, 26 March
1998, B. L. Stuart and T. Chan-ard.
Remarks. - Species boundaries within the genus
Cyclemys remain uncertain (Fritz and Ziegler, 1999;
Guicking et al., 2002). The specimens we assigned to C.
tcheponensis have dark radiating patterns of the plastra,pigmented chins, head and neck stripes, and dorsal spotting on the crown of the head, as illustrated by Fritz andZiegler (1999) and Fritz et al. (1997). Fritz and Ziegler(1999) reviewed records of Cyclemys from the region,
and Ziegler (2002) reported the species in local trade in
Ha Tinh Province, Vietnam.
Cyclemys sp.
[Map 8]
Cambodia. - FMNH 259418, carapace only, CL = 19.5,
CW = 14.7, Kampong Speu Province, Koh Kong
Samling Logging Concession, 11° 24' 15'' N 103° 49'
47'' E, 200 m, recovered from hunter’s camp, mixed
deciduous forest and grassland, 15 February 2000, J.
Walston. FMNH 259419, carapace only, CL = 22.1, CW= 16.1, collecting information same as FMNH 259418.
FMNH 259420, carapace only, CL = 19.1, CW = 15.9,
collecting information same as FMNH 259418. FMNH
259421, plastron only, PL = 21.2, collecting information
same as FMNH 259418. FMNH 259423, plastron only,
PL = 21.0, Koh Kong Province, Sre Ambel District,
Chay Reap Village, west bank of Prek Sre Ambel River,11° 29' 10'' N 103° 47' 00'' E, <10 m, obtained fromhunter in Chay Reap, 23 August 2000, B. L. Stuart and
S. G. Platt.
Laos. - FMNH 258893, carapace only, CL = 21.8, CW= 16.6, Champasak Province, Pakxong District, Ban
Latsasin Village, near Xe Nam Noy River, 800 m, eaten
by residents of Ban Latsasin Village, 02 April 1995, T.
D. Evans.
Remarks. - The condition of these shell fragments precludes identifying them to species. They are not necessarily a species different from atripons or tcheponensis.
Heosemys grandis (Gray, 1860)[Map 9]Cambodia. - FMNH 259409, carapace only, CL = 30.5,
CW = 22.6, Phnom Penh, Oreussay Market, purchasedin market, 17 May 1999, S. G. Platt. FMNH 259405,
plastron only, PL = 26.5, Koh Kong Province, Sre
Ambel District, Boeung Tradok Pong Village, 11° 31'
10' N 103° 46' 55'' E, obtained from hunter in Boeung

Vol. 10, p. 133

Tradok Pong, 24 August 2000, B. L. Stuart and S. G.
Platt. FMNH 259406, plastron only, PL = 20.8, KohKong Province, Sre Ambel District, Chaouethail Plous
Village on Sre Ambel River, 11° 18' 03'' N 103° 44' 56''
E, obtained from hunter in Chaouethail Plous, 21 August
2000, B. L. Stuart and S. G. Platt. Fig. 11, photographonly, CL = 31.8, PL = 28.6, Koh Kong Province, Thmor
Andart Village along Stoeng Metoek River, 11° 49.23'
N, 102° 53.62' E, captured by residents of Thmor
Andart, 10 May 2001, S. G. Platt, Heng Sovannara, andLong Kheng. FMNH 259407, plastron only, PL = 19.4,
collecting information same as FMNH 259406.
Laos. - FMNH 255271, carapace only, CL = 23.6, CW= 18.3, Khammouan Province, Thakhek District,
Khammouan Limestone (= Phou Hin Poun) National
Biodiversity Conservation Area, Ban Na Village, 17° 33'
N, 104° 52' E, eaten by residents of Ban Na, 02 April
1998, B. L. Stuart. FMNH 258885, plastron only, PL =27.0, Khammouan Province, Khammouan Limestone (=Phou Hin Poun) National Biodiversity Conservation
Area, Ban Namphick Village, eaten by residents of Ban
Namphick, 22 May 1994, R. J. Timmins. FMNH
258894, carapace only, CL = 36.3, CW = 24.7, collecting information same as FMNH 258885. FMNH
258889, plastron only, PL = 13.9, Laos, Khammouan
Province, Khammouan Limestone (= Phou Hin Poun)
National Biodiversity Conservation Area, Ban
Chocksavang Village, eaten by residents of Ban
Chocksavang, 22 May 1994, R. J. Timmins. FMNH
258882, carapace only, CL = 23.2, CW = 19.1,
Savannakhet Province, Thaphangthong District, Xe
Bang Nouan National Biodiversity Conservation Area,
Ban Houay Meun Village, eaten by residents of Ban
Houay, 20 June 1994, R. J. Timmins. FMNH 258883,
carapace (broken) and plastron only, PL = 36.6, collecting information same as FMNH 258882 except collected 19 June 1994. FMNH 258877, carapace only, CL =34.1, CW = 24.2, Salavan Province, Toumlan District,
Xe Bang Nouan National Biodiversity Conservation
Area, Ban Nalan Village, eaten by residents of Ban
Nalan, 15 June 1994, R. J. Timmins. FMNH 258878,
intact shell only, CL = 36.8, CW = 25.8, PL = 35.3, BD= 14.7, Salavan Province, Xe Bang Nouan National
Biodiversity Conservation Area, Ban Konglur Village,
eaten by residents of Ban Konglur, 10 June 1994, R. J.
Timmins. FMNH 258891, carapace only, CL = 17.3,
CW = 15.1, Salavan Province, Xe Bang Nouan National
Biodiversity Conservation Area, Ban Nasompeng
Village, eaten by residents of Ban Nasompeng, 09 June
1994, R. J. Timmins. FMNH 258890, carapace (broken)
only, CL = 19.4, Champasak Province, Pathoumphon
District, Xe Pian National Biodiversity Conservation
Area, Xe Pian River upstream from Ban Phonsaat
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Asiatic Herpetological Research

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Figures 1-15. See text for locality details and circumstances of the record. 1. Platysternon megacephalum Huaphahn
Province, Laos (photo B. L. Stuart); 2. Platysternon megacephalum Khammouan Province, Laos (photo B. L. Stuart);
3. Platysternon megacephalum Xe Kong Province, Laos (photo D. Showler); 4. Cuora amboinensis Battambang
Province, Cambodia (photo B. L. Stuart); 5. Cuora amboinensis Kampong Thom Province, Cambodia (photo S. G.
Platt); 6. Cuora amboinensis Koh Kong Province, Cambodia (photo S. G. Platt); 7. Cuora amboinensis Koh Kong
Province, Cambodia (photo S. G. Platt). 8. Cuora amboinensis Kien Giang Province, Vietnam (photo B. L. Stuart); 9.
Cyclemys atripons Koh Kong Province, Cambodia (photo S. G. Platt); 10. Cyclemys tcheponensis Bolikhamxay
Province, Laos (photo B. L. Stuart); 11. Heosemys grandis Koh Kong Province, Cambodia (photo S. G. Platt); 12.
Hieremys annandalii Battambang Province, Cambodia (photo B. L. Stuart); 13. Hieremys annandalii Siem Reap
Province, Cambodia (photo S. G. Platt); 14. Hieremys annandalii Kampong Thom Province, Cambodia (photo S. G.
Platt); 15. Malayemys subtrijuga Battambang Province, Cambodia (photo B. L. Stuart).

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Figures 16-27. See text for locality details and circumstances of the record. 16. Malayemys subtrijuga Kampong Thom
Province, Cambodia (photo S. G. Platt); 17. Malayemys subtrijuga Kandal Province, Cambodia (photo S. G. Platt); 18.
Malayemys subtrijuga Vientiane, Laos (photo W. G. Robichaud). 19. Manouria impressa Koh Kong Province,
Cambodia (photo S. G. Platt). 20. Manouria impressa Ratanakiri Province, Cambodia (photo Suon Phalla/TRAFFIC).
21. Manouria impressa Xe Kong Province, Laos (photo B. L. Stuart). 22. Chelonia mydas Kampong Speu Province,
Cambodia (photo B. L. Stuart). 23. Eretmochelys imbricata Sihanoukville Province, Cambodia (photo S. G. Platt). 24.
Dermochelys coriacea Gulf of Thailand, near Sihanoukville, Cambodia (photo Vanna Nhem). 25. Amyda cartilaginea
Koh Kong Province, Cambodia (photo S. G. Platt). 26. Amyda cartilaginea Khammouan Province, Laos (photo B. L.
Stuart). 27. Pelochelys cantorii Kratie Province, Cambodia (photo D. Gambade).

after being eaten by hunters, May 1995, T. D. Evans.
FMNH 255266, complete specimen, CL = 7.5, CW =6.4, PL = 6.2, BD = 2.5, Champasak Province,
Mounlapamok District, Dong Khanthung Proposed
National Biodiversity Conservation Area, 14° 07' N 105°
29' E, 60 m, grassland with dry dipterocarp and evergreen forest along Xe Lepou River, found in mud at bot-

tom of flooded marsh, water depth about 15 cm, 11 July
1998, B. L. Stuart. FMNH 255272, carapace only, CL =28.6, CW = 22.2, Champasak Province, Mounlapamok
District, Dong Khanthung Proposed National
Biodiversity Conservation Area, Ban Thahin Village, on
Xe Lepou River, 14° 08' N 105° 35' E, 60 m, eaten byresidents of Ban Thahin, 17 July 1998, B. L. Stuart.

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Maps 1-17 of records given in the text: shaded circles represent records collected in the field; half-shaded circles represent records obtained from hunters, abandoned hunting camps, or villages; unshaded circles represent records purchased from markets.

Hieremys annandalii (Boulenger, 1903)[Map 10]
Cambodia. - FMNH 259408, carapace only, CL = 32.3,
CW = 24.8, Phnom Penh, Oreussay Market, purchasedin market, 17 May 1999, S. G. Platt. FMNH 258876,
carapace and plastron, CL = 40.3, CW = 26.9, PL = 35.5,
Battambang Province, Ek Phnom District, Koh Chivang
Commune, Prek Toal Village on Tonle Sap Lake, 13° 14'
28'' N 103° 39' 32'' E, <10 m, eaten by residents of Prek
Toal who captured it from nearby flooded forest, 27
August 1999, B. L. Stuart, J. Smith, K. Davey. Fig. 12,
photograph only, same collecting information as FMNH
258876 except not yet eaten. FMNH 259398, carapace
only, CL = 37.4, CW = 26.7, Siem Reap Province, Siem
Reap District, Choeng Khneas Port on Tonle Sap Lake,found in fisherman’s house, 28 June 2000, B. L. Stuart

and S. G. Platt. Fig. 13, photograph only, two living animals, CL = 42.1 and PL = 34.7, CL = 33.4 and PL = 28.2,
Siem Reap Province, Siem Reap District, Choeng
Khneas Village, 13° 15.18' N; 103° 49.37' E, collectedby residents of Choeng Khneas from Tonle Sap, 4October 2000, S. G. Platt, Heng Sovannara, Long
Kheng, and Vuthy Monyrath. Fig. 14, photograph only,
CL = 37.2, PL = 28.4, Kampong Thom Province,
Kampong Thom Town, 12° 42.69' N; 104° 53.31' E,
photographed in market, 21 June 2000, S. G. Platt, Heng
Sovannara, and Long Kheng.
Laos. - FMNH 258879, carapace and plastron only, CL= 33.3, CW = 23.0, PL = 31.3, Attapu Province,
Sanamsai District, Xe Pian River Basin, Ban Chanto
Village, 100-150 m, eaten by residents of Ban Chanto,
21 April 1995, T. D. Evans. FMNH 259399, carapace
only, CL = 29.7, CW = 22.0, Attapu Province, Sanamxai

2004

Asiatic Herpetological ResearchDistrict, Ban Mai Village, 14° 42' 30'' E 106° 29' 50'' E,
obtained from hunter in Ban Mai, 18 September 2000,
B. L. Stuart and S. G. Platt.
Vietnam. - FMNH 259074, complete specimen, CL =12.4, CW = 10.8, PL = 11.5, BD = 5.6, Kien Giang
Province, An Minh District, Dong Hoa Town, 09° 45'
25'' N 105° 00' 03'' E, <10m, purchased from a reptile
trade shop, 10 November 2000, B. L. Stuart.
Malayemys subtrijuga (Schlegel and Müller, 1844)[Map 11]
Cambodia. - FMNH 259404, carapace only, CL = 19.2,
CW = 14.1, Phnom Penh, purchased in market, 07 July
2000, S. G. Platt. FMNH 259403, carapace only, CL =21.5, CW = 16.9, Siem Reap Province, Siem Reap
District, Siem Reap Town, purchased from turtle restaurant, 11 December 1999, S. G. Platt. Fig. 15, photographonly, Battambang Province, Ek Phnom District, Koh
Chivang Commune, Prek Toal Village on Tonle Sap
Lake, 13° 14' 28'' N 103° 39' 32'' E, captured by residents of Prek Toal, 27 August 1999, B. L. Stuart, J.
Smith, K. Davey. FMNH 259402, plastron only, PL =17.8, collecting information same as FMNH 259403.
FMNH 259401, carapace (broken) and plastron, CL =23.2, PL = 20.0, Kampong Thom Province, Kampong
Thom Town, obtained from restaurant, 22 June 2000, S.
G. Platt. Fig. 16, photograph only, CL = 21.3, PL = 17.5,
Kampong Thom Province, Sary Village, 12° 48.48' N
104° 44.19' E, collected by residents of Sary in Tonle
Sap, 21 June 2000, S. G. Platt, Heng Sovannara, andLong Kheng. Fig. 17, photograph only, CL = 19.8, PL =16.6, Kandal Province, Bassac Marshes, Prasat Village,
11° 17.72' N, 105° 08.61' E, captured by residents of
Prasat, 5 July 2000, S. G. Platt, Heng Sovannara, andLong Kheng. FMNH 259400, plastron only, PL = 12.6,
Koh Kong Province, Sre Ambel District, Chay Reap
Village, west bank of Sre Ambel River, 11° 29' 10'' N
103° 47' 00'' E, <10 m, obtained from hunter in Chay
Reap, 23° August 2000, B. L. Stuart and S. G. Platt.
Laos. - Fig. 18, photograph only, Vientiane
Municipality, Vientiane, crossing road near culvert, May
1999, W. G. Robichaud. FMNH 258868, complete specimen, CL = 8.4, CW = 6.9, PL = 7.3, BD = 3.6, Vientiane
Municipality, Vientiane, purchased in That Luang Fresh
Food Market, 27 February 2000, B. L. Stuart. FMNH
255269, carapace only, CL = 15.9, CW = 12.9,
Khammouan Province, Yommalat District, Khammouan
Limestone (= Phou Hin Poun) National Biodiversity
Conservation Area, Ban Vieng Village, 17° 20' N, 104°
57' E, eaten by residents of Ban Vieng, 30 March 1998,
B. L. Stuart. FMNH 258888, plastron only, PL = 12.8,
Khammouan Province, Khammouan Limestone (= Phou
Hin Poun) National Biodiversity Conservation Area,

Vol. 10, p. 145

Ban Chocksavang Village, eaten by residents of Ban
Chocksavang, 22 May 1994, R. J. Timmins. FMNH
259653 plastron (broken) only, collecting information
same as FMNH 258888. FMNH 259654, plastron only,
PL = 12.5, collecting information same as FMNH
258888. FMNH 255268, intact shell, CL = 10.9, CW =8.8, PL = 9.2, BD = 4.8, Champasak Province,
Mounlapamok District, Dong Khanthung Proposed
National Biodiversity Conservation Area, Ban Thahin
Village, on Xe Lepou River, 14° 08' N 105° 35' E, 60 m,
eaten by residents of Ban Thahin, 17 July 1998, B. L.
Stuart. FMNH 255267, complete specimen, CL = 10.9,
CW = 8.4, PL = 9.3, BD = 4.7, Champasak Province,
Mounlapamok District, Dong Khanthung Proposed
National Biodiversity Conservation Area, Ban Tap Seng
Village, 14° 15' N 105° 41' E, mixed deciduous forest,
captured by resident of Ban Tap Seng for food, 25 July
1998, B. L. Stuart.
Vietnam. - FMNH 259075, complete specimen, CL =5.3, CW = 4.3, PL = 4.4, BD = 2.7, Kien Giang
Province, Vinh Thuang District, U Minh Thuong Nature
Reserve, 09° 32' 40'' N 105° 05' 11'' E, <10m, flooded
grassland and agricultural fields, caught in fishing net
set in canal, 02 November 2000, B. L. Stuart. FMNH
259394, complete specimen, CL = 4.8, CW = 3.8, PL =3.9, BD = 2.4, Kien Giang Province, An Minh District,
U Minh Thuong Nature Reserve, 09° 37' 29'' N 105° 07'
59'' E, <10m, flooded grassland and agricultural fields,
taken in fishing net, 16 November 2000, B. L. Stuart.
Sacalia quadriocellata (Siebenrock, 1903)[Map 12]
Laos. - FMNH 255270, carapace only, CL = 13.8, CW= 10.6, Huaphahn Province, Vieng Tong District, Nam
Et National Biodiversity Conservation Area, Nam PeunRiver, 20° 17' N 103° 25' E, 985 m, found in hunter’scamp, 01 April 1998, P. Davidson. FMNH 256542,
complete specimen, CL = 14.2, CW = 10.3, PL =12.5,
BD = 4.5, Khammouan Province, Nakai District, NakaiNam Theun National Biodiversity Conservation Area,
Annamite Mountains, 17° 50' N 105° 35' E, 600 m, wetevergreen forest along Houay Dreng Stream, found 1 m
deep in pool with sandy substrate, 01 December 1999,
B. L. Stuart. FMNH 256543, complete specimen, CL =7.2, CW = 6.7, PL = 5.9, BD = 2.8, Khammouan
Province, Nakai District, Nakai-Nam Theun National
Biodiversity Conservation Area, Annamite Mountains,
17° 56' N 105° 31' E, 600 m, Houay Maka-Noi Stream
near Ban Maka Village, collected by resident of Ban
Maka, 15 November 1999, B. L. Stuart.
Remarks. - Ziegler (2002) field-collected and reported
the species in local trade in Ha Tinh Province, Vietnam.

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Siebenrockiella crassicollis (Gray, 1831)[Map 13]
Cambodia. - FMNH 259054, complete specimen, CL =14.3, CW = 10.5, PL = 11.7, BD = 5.8, Khampong Thom
Province, Khampong Thom Town, 12° 42' N 104° 53' E,
purchased from turtle trader in Khampong Thom Town,
21 June 2000, S. G. Platt. FMNH 259053, whole specimen, CL = 15.9, CW = 12.0, PL =13.7, BD = 6.8, KohKong Province, Sre Ambel District, Chay Reap Village,
west bank of Sre Ambel River, 11° 29' 10'' N 103° 47'
00'' E, <10 m, captured by residents of Chay Reap, 24
August 2000, B. L. Stuart and S. G. Platt. FMNH
259396, intact shell, CL = 16.4, CW = 12.1, PL = 13.7,
BD = 6.6, Koh Kong Province, Sre Ambel District, Chay
Reap Village, west bank of Sre Ambel River, 11° 29' 10''
N, 103° 47' 00'' E, <10 m, obtained from hunter in Chay
Reap, 24 August 2000, B. L. Stuart and S. G. Platt.
FMNH 259397, plastron only, PL = 16.1, Koh Kong
Province, Sre Ambel District, Kohriem Village on Sre
Ambel River, obtained from hunter in Kohriem, 27
August 2000, B. L. Stuart and S. G. Platt. FMNH
259055, complete specimen, CL = 16.2, CW = 11.5, PL= 12.8, BD = 5.9, Koh Kong Province, Sre Ambel
District, Prek Kroch River (tributary of Sre Ambel
River), 11° 06' 20'' N, 103° 39' 35'' E, <10 m, flooded
paddy at edge of mangrove and Melaleuca forest, captured by fisherman in bamboo fish trap set at that location, 27 August 2000, B. L. Stuart and S. G. Platt.
Remarks. - FMNH 259053 has a pale streak extending
from the ear to the lower jaw, but the heads of FMNH
259054-55 are entirely dark, with no pale markings.
Testudinidae
Indotestudo elongata (Blyth, 1853)[Map 14]
Cambodia: FMNH 262316, carapace only, CL = 14.7,
CW = 10.0, Mondolkiri Province, Keo S’Marr District,
Samling Logging Concession, 200 m, mixed deciduous
forest and grassland, found in hunter’s camp, 13 May
2000, J. Walston. FMNH 262315, plastron only, PL =19.8, Koh Kong Province, Sre Ambel District, Chay
Reap Village, west bank of Sre Ambel River, 11° 29' 10''
N, 103° 47' 00'' E, <10 m, obtained from hunter in Chay
Reap, 23 August 2000, B. L. Stuart and S. G. Platt.
TC001 (held in the Field Museum of Natural History,
Division of Amphibians & Reptiles), blood sample only,
collecting information same as FMNH 262315, obtained
from hunter in Chay Reap and released after taking
blood sample, 23 August 2000, B. L. Stuart and S. G.
Platt. FMNH 262312, plastron only, PL = 18.2, collect-2004

ing information same as FMNH 262315. FMNH
262307, carapace only, CL = 25.6, CW = 15.1, collecting information same as FMNH 262315. FMNH
262297, carapace only, CL = 24.7, CW = 14.6, KohKong Province, Sre Ambel District, Boeung Tradok
Pong Village, 11° 31' 10'' N, 103° 46' 55'' E, obtained
from hunter in Boeung Tradok Pong, 24 August 2000, B.
L. Stuart and S. G. Platt. FMNH 262313, plastron only,
PL = 19.4, collecting information same as FMNH
262297. FMNH 262314, plastron only, PL = 18.2, collecting information same as FMNH 262297. FMNH
262302, carapace and plastron, CL = 19.5, CW = 13.0,
PL = 16.5, Koh Kong Province, Sre Ambel District,
Kohriem Village on Sre Ambel River, obtained fromhunter in Kohriem, 27 August 2000, B. L. Stuart and S.
G. Platt. FMNH 262311, plastron only, PL = 15.4, KohKong Province, Sre Ambel District, Chaouethail Plous
Village on Sre Ambel River, 11° 18' 03'' N, 103° 44' 56''
E, obtained from hunter in Chaouethail Plous, 21 August
2000, B. L. Stuart and S. G. Platt. FMNH 262310, plastron only, PL = 14.8, collecting information same as
FMNH 262311.
Laos. - FMNH 262294, carapace only, CL = 22.8, CW= 14.0, Khammouan Province, Boualapha District, Hin
Nam No National Biodiversity Conservation Area, Ban
Tasang Village, eaten by residents of Ban Tasang, 30
December 1995, R. J. Timmins. FMNH 259056, complete specimen, CL = 17.0, CW = 10.9, PL = 14.8, BD =7.4, Khammouan Province, Nakai District, Khammouan
Limestone (= Phou Hin Poun) National Biodiversity
Conservation Area, Ban Na Bon Village, limestone karst
with dry evergreen/mixed deciduous forest, 17° 54' N
104° 51' E, captured by residents of Ban Na Bon for
food, 22 March 1998, B. L. Stuart and T. Chan-ard.
FMNH 262304, carapace only, CL = 17.9, CW = 12.4,
Khammouan Province, Thakhek District, Khammouan
Limestone (= Phou Hin Poun) National Biodiversity
Conservation Area, Ban Na Village, 17° 33' N, 104° 52'
E, eaten by residents of Ban Na, 02 April 1998, B. L.
Stuart. FMNH 262299, carapace only, CL = 24.2, CW =15.4, Khammouan Province, Khammouan Limestone (=Phou Hin Poun) National Biodiversity Conservation
Area, Ban Namphick Village, eaten by residents of Ban
Namphick, 22 May 1994, R. J. Timmins. FMNH
262295, carapace only, CL = 25.2, CW = 15.1, collecting information same as FMNH 262299. FMNH
262296, carapace only, CL = 26.9, CW = 16.7, Salavan
Province, Xe Bang Nouan National Biodiversity
Conservation Area, Ban Konglur Village, eaten by residents of Ban Konglur, 10 June 1994, R. J. Timmins.
FMNH 262301, carapace only, CL = 29.0, CW = 18.1,
Attapu Province, Xe Nam Noy River, Ban Mun Houa
Muang Village, 300 m, eaten by residents of Ban Mun
Houa Muang, April 1995, D. Showler. FMNH 262303,

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Remarks. - Ziegler (2002) reported the species in local
trade in Ha Tinh Province, Vietnam.
Manouria impressa (Günther, 1882)[Map 15]

Figure 28. See text for locality details and circumstancesof the record. Pelochelys cantorii Champasak Province,
Laos (photo I. G. Baird).

carapace only, CL = 23.3, CW = 14.4, Attapu Province,
Sanamsai District, Xe Kong River, near Ban Sompoy
Village and Cambodian border, 90 m, discarded carapace found in camp after being eaten by hunters, May
1995, T. D. Evans. FMNH 262298, carapace only, CL =22.3, CW = 14.3, Champasak Province, Pathoumphon
District, Xe Pian National Biodiversity Conservation
Area, Xe Pian River upstream from Ban Phonsaat
Village, 100 m, discarded carapace found in camp along
Xe Pian River after being eaten by hunters, May 1995,
T. D. Evans. FMNH 262308, carapace only, CL = 17.7,
CW = 11.4, Champasak Province, Mounlapamok
District, Dong Khanthung Proposed National
Biodiversity Conservation Area, found in hunter’s campnear Nong Sathevada Wetland, 14° 13' N 105° 36' E, 100m, 04 March 1998, C. Francis. FMNH 262306, carapace
only, CL = 18.9, CW = 12.0, Champasak Province,
Mounlapamok District, Dong Khanthung Proposed
National Biodiversity Conservation Area, Ban Baw
Village, 14° 14' N, 105° 27' E, eaten by residents of Ban
Baw, 18 July 1998, B. L. Stuart. FMNH 262305, carapace only, CL = 26.6, CW = 15.9, Champasak Province,
Mounlapamok District, Dong Khanthung Proposed
National Biodiversity Conservation Area, Ban Khiem
Village, 14° 14' N, 105° 20' E, eaten by residents of Ban
Khiem, 24 July 1998, B. L. Stuart. FMNH 262300,
carapace only, CL = 24.9, CW = 14.7, Champasak
Province, Mounlapamok District, Dong Khanthung
Proposed National Biodiversity Conservation Area, Ban
Kadian Village, 14° 26' N, 105° 42' E, eaten by residentsof Ban Kadian, 29 July 1998, B. L. Stuart.

Cambodia. - Fig. 19, photograph only of two intact
shells, CL = 26.6 and PL = 23.8, CL = 22.8 and PL =19.9, Koh Kong Province, Thmor Andart Village, along
Stoeng Metoek River, 11° 49.23' N, 102° 53.62' E, collected by residents of Thmor Andart, 10 May 2001, S. G.
Platt, Heng Sovannara, and Long Kheng. Fig. 20, photograph only, Ratanakiri Province, Oyadao District,found in house of Vietnamese wildlife trader on
Cambodian side of Oyadao border checkpoint, Suon
Phalla/TRAFFIC, 16 June 2002.
Laos. - FMNH 262321, carapace only, CL = 23.7, CW= 17.3, Phongsaly Province, Phongsaly District, Phou
Dendin National Biodiversity Conservation Area, Ban
Sopkang Village, 22° 05' 51'' N, 102° 15' 02'' E, eaten byresidents of Ban Sopkang, 17 October 1999, B. L. Stuart
and H. F. Heatwole. FMNH 262317, carapace only, CL= 27.7, CW = 19.9, collecting information same as
FMNH 262321. FMNH 262324, carapace only, CL =28.6, CW = 20.2, Huaphahn Province, Vieng Tong
District, Phou Louey National Biodiversity
Conservation Area, Ban Sa Kok Village, 20° 11'' N, 103°
12'' E, eaten by residents of Ban Sa Kok, 30 April 1998,
B. L. Stuart. FMNH 262319, carapace only, CL = 27.0,
CW = 19.9, collecting information same as FMNH
262324. FMNH 262323, carapace and plastron, CL =20.8, CW = 16.3, PL = 19.4, collecting information sameas FMNH 26234. FMNH 262318, carapace only, CL =25.0, CW = 19.2, Huaphahn Province, Vieng Tong
District, Phou Louey National Biodiversity
Conservation Area, Ban Phone Xong Village, eaten byresidents of Ban Phone Xong, 04 May 1998, D. Showler.
FMNH 262322, plastron only, PL = 21.1, Khammouan
Province, Hin Nam No National Biodiversity
Conservation Area, Ban Katok Village, eaten by residents of Ban Katok, 12 January 1996, R. J. Timmins.
Fig. 21, photograph only, Xe Kong Province, Kaleum
District, Ban Talouy-Ngai Village, 15° 59' 50'' N, 106°
57' 22'' E, captured by residents of Ban Talouy-Ngai, 27
June 1999, B. L. Stuart. FMNH 262320, carapace only,
CL = 27.6, CW = 20.6, Attapu Province, Xe Nam Noy
River, Ban Taot Village, 800 m, eaten by residents of
Ban Taot, April 1995, D. Showler.
Remarks. - Lehr and Holloway (2000) obtained a single
carapace of M. impressa from a hunter in Ratanakiri
Province, Cambodia, who claimed to have captured theturtle in mountains north of Siem Pang, Stung Treng
Province.

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Cheloniidae
Chelonia mydas (Linnaeus, 1758)
Cambodia. - Fig. 22, photograph only, Kampong Speu
Province, Phnom Srouch District, Srei Khlong Market,
07 June 2000, B. L. Stuart.
Eretmochelys imbricata (Linnaeus, 1766)
Cambodia. - Fig. 23, photograph only, Sihanoukville
Province, Mitta Pheap District, Koh Rong Island, Bagnu
Village, obtained from fisherman in Bagnu, 01 January
2000, F. Goes.

2004

complete specimen, CL = 18.7, CW = 16.7, PL = 15.2,
BD = 4.9, Khammouan Province, Nakai District,
Khammouan Limestone (= Phou Hin Poun) National
Biodiversity Conservation Area, 17° 54' N, 104° 51' E,
obtained from villager who caught it in the Nam Thon
River, 28 March 1998, B. L. Stuart.
Vietnam. - FMNH 252163, whole specimen, CL = 12.5,
CW = 10.7, PL = 10.4, BD = 3.8, Gia-Lai Province,
Ankhe District, Buon Loi Village, 20 km northwest of
Kannack Town, Annamite Mountains, 14° 20' N, 108°
36' E, 700-750 m, Daklest River in the village, 07 May
1995, I. Darevsky and N. L. Orlov.
Remarks. - Farkas and Ziegler (2002) reviewed recordsof A. cartilaginea from Vietnam, Laos, and Cambodia.

Dermochelyidae
Dermochelys coriacea (Vandelli, 1761)
Cambodia. - Fig. 24, photograph only. Stuart et al.
(2002) discussed in detail this record from offshore of
the southern point of Koh Sra Mauch Island in the Gulfof Thailand, near Sihanoukville.
Trionychidae
Amyda cartilaginea (Boddaert, 1770)[Map 16]
Cambodia. - Fig. 25, photograph only, CL = 7.4, KohKong Province, Koh Kong Knong Village, 11° 25.97' N;
103° 09.77' E, captured locally by villager in Stoeng
Kep River, 11 May 2001, S. G. Platt, Heng Sovannara,and Long Kheng.
Laos. - FMNH 258869, complete specimen, CL = 7.4,
CW = 6.8, PL = 2.1, BD = 5.7, Vientiane Municipality,
Vientiane, purchased in That Luang Fresh Food Market,
27 February 2000, B. L. Stuart. FMNH 258874, complete specimen, CL = 29.5, CW = 26.0, PL = 20.6, BD =9.5, Bolikhamxay Province, Northern Extension
Proposed National Biodiversity Conservation Area, Ban
Chom-Tong Village, near Nam Mouan River, captured
by hunters in Nam Mouan River, 18 March 1995, R. J.
Timmins. Fig. 26, photograph only, Khammouan
Province, Nakai District, Nakai-Nam Theun National
Biodiversity Conservation Area, Ban Kao-Oy Village,
17° 43' 39'' N 105° 20' 05'' E, captured by resident of
Ban Kao-Oy, 03 November 1998, B. L. Stuart. FMNH
255264, complete specimen, CL = 18.2, CW = 16.4, PL= 13.8, BD = 4.5, Khammouan Province, Thakhek
District, Khammouan Limestone (= Phou Hin Poun)
National Biodiversity Conservation Area, 17° 33' N,
104° 52' E, obtained from villager who captured it whilefishing, 01 April 1998, B. L. Stuart. FMNH 255265,

Pelochelys cantorii Gray, 1864
[Map 17]
Cambodia. - Fig. 27, photograph only, Kratie Province,
Kratie District, Kratie Town, purchased from fisherman
in a Kratie market and released in the Mekong Riverafter being photographed, September 2000, D.
Gambade.
Laos. - Fig. 28, photograph only, Champasak Province,
Khong District, Ban Hang Khone Village, captured byresidents of Ban Hang Khone in the Mekong River, I. G.
Baird.

Acknowledgments
We thank Ian Baird, James Chamberlain, Ilya Darevsky,
David Davenport, Peter Davidson, Thomas Evans,
Charles Francis, Heng Sovannara, Long Kheng, Denis
Gambade, Troy Hansel, Frederic Goes, Nikolai Orlov,
William Robichaud, Sergei Ryabov, Shi Haitao, David
Showler, Suon Phalla/TRAFFIC, and Robert Timminsfor providing us their specimens or photographs. We
thank An Dara, Bui Huu Manh, Tanya Chan-ard,
Benjamin Hayes, Harold Heatwole, Heng Sovannara,
Khamkoun Khounboline, Long Kheng, Nguyen Van
Cuong, Augustus McCrae, William Robichaud, PhilipRound, Suon Phalla, Bee Thaovanseng, and Vuthy
Monyrath for assistance with collecting specimens in thefield. The opportunity for BLS to work in Laos wasmade possible by the Wildlife Conservation Society /Division of Forest Resource Conservation Cooperative
Program, and in Vietnam by Fauna & FloraInternational, CARE International in Vietnam, and thestaff of Pu Mat and U Minh Thuong Nature Reserves.
The opportunity for BLS and SGP to work in Cambodia
was made possible by the Wildlife Conservation Society/Ministry of Agriculture, Forestry and Fisheries/

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Ministry of Environment Collaborative Program.
Financial support was provided by the National
Geographic Society (Grant no. 6247-98 with Harold
Heatwole), the Wildlife Conservation Society, and The
John D. and Catherine T. MacArthur Foundation. Harold
Voris, Alan Resetar, and Jamie Ladonski facilitatedexamining specimens at the Field Museum of Natural
History. Sean Bober constructed the maps. Two anonymous reviewers improved the manuscript.

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