2004

Asiatic Herpetological ResearchVol. 10, pp. 191-201

Annotated Checklist of Amphibians and Reptiles of Pakistan
MUHAMMAD SHARIF KHAN
Apt # A17, 151-S. Bishop Ave., Secane, PA 19018, USA
E-mail: typhlops99@hotmail.com
Abstract. - From recent herpetological collections several new amphibian and reptilian taxa have been added to theherpetofauna of Pakistan. Thus raising the number of species from Minton's 144 and Mertens' 178 to 225.
Key words. - Checklist, Pakistan, herpetofauna.

Introduction
Following is the checklist of amphibian and reptile
species that, so far, have been recorded and described in
the major works on the herpetology of the subcontinent
from the areas now included in Pakistan (Günther, 1864;
Murray, 1884, 1886, 1892; Boulenger, 1890, 1896;
Smith, 1931, 1935, 1943). Recently described andrecorded species from Pakistan are also included in it(Anderson and Leviton, 1966, 1967, 1969; Andersonand Minton, 1963; Cherlin, 1981, 1983; Szczerbak,
1991; Golubev and Szczerbak, 1981; Ingoldby, 1922;
Mertens 1969a,b, 1970, 1971, 1972, 1974; Baig, 1988,
1989, 1998, 1999; Baig and Böhme, 1996; Baig and
Gvozdik, 1998; Dubois and Khan, 1979; Minton, 1966;
Minton and Anderson, J. 1965; Minton et al., 1970;
Khan, A. Q. and Khan, M.S. 1996; Khan, 1972, 1974,
1980, 1984a,b, 1988, 1989, 1991, 1992, 1993a,b, 1994,
1997a,b,c, 1998, 1999a,b, 2000, 2001a,b, 2003a,b; Khanand Baig, 1992; Khan and Khan, R.Z., 1997; Khan andTasnim, 1989, 1990; Khan and Rösler, 1999; RastegarPouyani, 1999.

Frogs and Toads
Family: Bufonidae
Bufo Laurenti, 1768
Bufo himalayanus Günther, 1864
Bufo latastii Boulenger, 1882
Bufo melanostictus hazarensis Khan, 2001
Bufo olivaceus Blanford, 1874
Bufo pseudoraddei pseudoraddei Mertens, 1971
Bufo pseudoraddei baturae Stock, Schmid,
Steinlein, and Grosse, 1999
Bufo siacheninsis Khan, 1997
© 2004 by Asiatic Herpetological Research

Bufo stomaticus Lütkin, 1862
Bufo surdus Boulenger, 1891
Bufo viridis zugmayeri Eiselt and Schmidtler, 1973

Family: Microhylidae
Microhyla Tschudi, 1828
Microhyla ornata (Duméril and Bibron, 1841)
Uperodon Dúmeril and Bibron, 1841
Uperodon systoma (Schneider, 1799)

Family: Ranidae
Euphlyctis Fitzinger, 1843
Euphlyctis cyanophlyctis cyanophlyctis (Schneider,
1799)
Euphlyctis cyanophlyctis microspinulata Khan,
1997
Euphlyctis cyanophlyctis seistanica (Nikolsky, 1900)
Hoplobatrachus Peters, 1863
Hoplobatrachus tigerinus (Daudin, 1802)
Fejervarya Bolkay, 1915
Fejervarya limnocharis (Boie, 1834)
Fejervarya syhadrensis (Annandale, 1919)
Nanorana Günther, 1836
Nanorana pleskei (Günther, 1896)

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Paa Dubois, 1975

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Kachuga tecta (Gray, 1831)

Paa barmoachensis (Khan and Tasnim, 1989)
Paa hazarensis Dubois and Khan, 1979

Family: Testudinidae

Paa sternosignata (Murray, 1885)

Agrionemys Khozatsky and Mlynarsky, 1966

Paa vicina (Stoliczka, 1872)

Agrionemys horsfieldii (Gray, 1844)

Sphaerotheca Duméril and Bibron, 1841.

Geochelone Fitzinger, 1835

Sphaerotheca breviceps (Schneider, 1799)

Geochelone elegans (Schopff, 1792)

Turtles and Tortoises

Family: Trionychidae
Aspideretes Hay, 1835

Family: Cheloniidae

Aspideretes gangeticus (Cuvier, 1825)

Caretta Rafiuesque, 1814

Aspideretes hurum (Gray, 1831)

Caretta caretta (Linnaeus, 1758)
Chitra Gray, 1844
Chelonia Brongniart, 1800

Chitra indica (Gray, 1831)

Chelonia mydas (Linnaeus, 1758)
Lissemys Smith, 1931
Eretmochelys Fitzinger, 1843

Lissemys punctata andersoni Webb, 1980

Eretmochelys imbricata (Linnaeus, 1766)

Crocodiles and Gavials
Lepidochelys Fitzinger, 1843
Lepidochelys olivacea (Eschscholtz, 1824)

Family: Crocodylidae
Crocodylus Laurenti, 1768

Family: Dermochelyidae

Crocodylus palustris Lesson, 1831

Dermochelys Blainville, 1816
Dermochelys coriacea (Vandelli, 1761)

Family: Gavialidae
Gavialis Oppel, 1811

Family: Emydidae

Gavialis gangeticus (Gmelin, 1789)

Geoclemys (Gray, 1821)

Lizards

Geoclemys hamiltonii (Gray, 1821)
Hardella Gray, 1870
Hardella thurjii Gray, 1870
Kachuga Gray, 1856
Kachuga smithii (Gray, 1863)

Family: Agamidae
Brachysaura Blyth, 1856
Brachysaura minor (Hardwicke and Gray, 1827)
Calotes Cuvier, 1817
Calotes versicolor versicolor (Daudin, 1802)

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Calotes versicolor farooqi Auffenberg and Rehman,
1995

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Trapelus agilis pakistanensis Rastegar-Pouyani, 1999
Trapelus megalonyx Günther, 1864
Trapelus rubrigularis Blanford, 1876

Japalura Gray, 1853

Trapelus ruderatus baluchianus (Smith, 1935)

Japalura kumaonensis (Annandale, 1907)

Family: Chameleonidae
Laudakia Gray, 1845

Chamaeleo Laurenti, 1768

Laudakia agrorensis (Stoliczka, 1872)

Chamaeleo zeylanicus Laurenti, 1768

Laudakia badakhshana (Anderson and Leviton, 1969)
Laudakia caucasia (Eichwald, 1831)
Laudakia fusca (Blanford, 1876)
Laudakia himalayana (Steindachner, 1869)
Laudakia lirata (Blanford, 1874)
Laudakia melanura nasiri Baig, 1999
Laudakia melanura melanura Blyth, 1854

Family: Eublepharidae
Eublepharis Gray, 1827
Eublepharis macularius (Blyth, 1854)

Family: Gekkonidae
Agamura Blanford, 1874
Agamura persica (Duméril, 1856)

Laudakia microlepis (Blanford, 1874)
Laudakia nupta (de Filippi, 1843)
Laudakia nuristanica (Anderson and Leviton, 1969)
Laudakia pakistanica (Baig, 1989)
Laudakia pakistanica auffenbergi Baig and Böhme,Altigekko M.S. Khan, 2003
Altigekko baturensis (Khan and Baig, 1992)
Altigekko boehmei (Szczerbak, 1991)
Altigekko stoliczkai (Steindachner, 1869)

1996
Laudakia pakistanica khani Baig and Böhme, 1996
Laudakia tuberculata (Hardwicke and Gray, 1827)
Phrynocephalus Kaup, 1825
Phrynocephalus clarkorum (Anderson and Leviton,
1967)
Phrynocephalus euptilopus Alcock and Finn, 1896

Bunopus Blanford, 1874
Bunopus tuberculatus Blanford, 1874
Crossobamon Boettger, 1888
Crossobamon lumsdeni (Boulenger, 1887)
Crossobamon maynardi (Smith, 1933)
Crossobamon orientalis (Blanford, 1876)

Phrynocephalus luteoguttatus Boulenger, 1887
Phrynocephalus maculatus Anderson, 1872
Phrynocephalus ornatus Boulenger, 1887
Phrynocephalus scutellatus Olivier, 1807

Cyrtopodion Fitzinger, 1843
Cyrtopodion agamuroides (Nikolsky, 1900)
Cyrtopodion kachhense kachhense (Stoliczka, 1872)
Cyrtopodion kachhense ingoldbyi Khan, 1997

Trapelus Cuvier, 1816
Trapelus agilis Olivier, 1804
Trapelus agilis agilis (Olivier, 1804)

Cyrtopodion kohsulaimanai (Khan, 1991d)
Cyrtopodion montiumsalsorum (Annandale, 1913)
Cyrtopodion potoharensis Khan, 2001

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Cyrtopodion scabrum (Heyden, 1827)

Teratoscincus Strauch, 1863

Cyrtopodion watsoni (Murray, 1892)

Teratoscincus microlepis Nikolsky, 1899

2004

Teratoscincus scincus (Schlegel, 1858)
Hemidactylus Oken, 1817

Teratoscincus scincus keyserlingi Strauch, 1863

Hemidactylus brookii Gray, 1845
Hemidactylus flaviviridis Rüppell, 1835

Tropiocolotes Peters, 1880

Hemidactylus frenatus Schlegel, 1836

Tropiocolotes depressus Minton and J. A. Anderson,

Hemidactylus leschenaultii Duméril and Bibron, 1836

1965

Hemidactylus persicus J. Anderson, 1872

Tropiocolotes persicus persicus (Nikolsky, 1903)

Hemidactylus triedrus (Daudin, 1802)

Tropiocolotes persicus euphorbiacola Minton, S.

Hemidactylus turcicus (Linnaeus, 1758)

Anderson, and J. A. Anderson, 1970

Indogekko M.S. Khan, 2003

Family: Lacertidae

Indogekko fortmunroi (Khan, 1993)

Acanthodactylus Wiegmann, 1834

Indogekko indusoani (Khan, 1980)

Acanthodactylus blanfordii Boulenger, 1918

Indogekko rhodocaudus (Baig, 1998)

Acanthodactylus cantoris Günther, 1864

Indogekko rohtasfortai (Khan and Tasnim, 1990)

Acanthodactylus micropholis Blanford, 1874

Mediodactylus Szczerbak and Golubev, 1977

Eremias Wiegmann, 1834

Indogekko walli (Ingoldby, 1922)

Eremias acutirostris (Boulenger, 1887)
Eremias aporosceles (Alcock and Finn, 1896)

Ptyodactylus Goldfuss, 1820

Eremias fasciata Blanford, 1874

Ptyodactylus homolepis Blanford, 1876

Eremias persica Blanford, 1874
Eremias scripta (Strauch, 1867)

Rhinogecko de Witte, 1973
Rhinogecko femoralis (Smith, 1933)

Mesalina Gray, 1838

Rhinogecko misonnei de Witte, 1973

Mesalina brevirostris Blanford, 1874
Mesalina watsonana (Stoliczka, 1872)

Siwaligekko M. S. Khan, 2003
Siwaligekko battalensis (Khan, 1993)

Ophisops Ménétriés, 1832

Siwaligekko dattanensis (Khan, 1980)

Ophisops elegans Ménétriés, 1832

Siwaligekko mintoni (Golubev and Szczerbak, 1981)

Ophisops jerdonii Blyth, 1853

Teratolepis Günther, 1870
Teratolepis fasciata (Blyth, 1853)

Family: Scincidae
Ablepharus Fitzinger, 1823
Ablepharus grayanus (Stoliczka, 1872)
Ablepharus pannonicus (Fitzinger, 1823)

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Chalcides Laurenti, 1768

Varanus bengalensis (Daudin, 1802)

Chalcides ocellatus (Forskål, 1775)

Varanus flavescens (Hardwicke and Gray, 1827)
Varanus griseus (Daudin, 1803)

Eurylepis Blyth, 1854

Varanus griseus caspius (Eichwald, 1831)

Eurylepis taeniolatus taeniolatus (Blyth, 1854)

Varanus griseus koniecznyi Mertens, 1954

Ophidia: Snakes

Lygosoma Hardwick and Gray, 1827
Lygosoma punctata (Linnaeus, 1766)
Mabuya Fitzinger, 1826
Mabuya dissimilis (Hallowell, 1860)
Mabuya macularia (Blyth, 1853)

Family: Leptotyphlopidae
Leptotyphlops Fitzinger, 1843
Leptotyphlops blanfordii (Boulenger, 1890)
Leptotyphlops macrorhynchus (Jan, 1862)

Family: Typhlopidae
Novoeumeces Griffith, Ngo, and Murphy, 2000
Novoeumeces blythianus (J. Anderson, 1871)

Ramphotyphlops Fitzinger, 1843
Ramphotyphlops braminus (Daudin, 1803)

Novoeumeces indothalensis (M.S. Khan and M.R.Z.
Khan, 1997)
Novoeumeces schneiderii zarudnyi (Nikolsky, 1900)

Typhlops Oppel, 1811
Typhlops ahsanai M.S. Khan, 1999
Typhlops diardii Schlegel, 1839

Ophiomorus Dúmeril and Bibron, 1839
Ophiomorus blanfordi Boulenger, 1887
Ophiomorus brevipes (Blanford, 1874)
Ophiomorus raithmai S. Anderson and Leviton, 1966
Ophiomorus tridactylus (Blyth, 1853)

Typhlops diardii platyventris M.S. Khan, 1998
Typhlops ductuliformes M.S. Khan, 1999
Typhlops madgemintonai madgemintonai M.S. Khan,
1999
Typhlops madgemintonai shermanai M.S. Khan, 1999

Scincella Mittleman, 1950

Family: Boidae

Scincella himalayana (Günther, 1864)

Eryx Daudin, 1803

Scincella ladacensis (Günther, 1864)

Eryx conicus (Schneider, 1801)

Family: Uromastycidae

Eryx johnii (Russell, 1801)
Eryx tataricus speciosus Zarevsky, 1915

Uromastyx Merrem, 1820
Uromastyx asmussi (Strauch, 1863)
Uromastyx hardwickii Gray, 1827

Family: Varanidae
Varanus Merrem, 1820

Python Daudin, 1803
Python molurus (Linnaeus, 1758)

Family: Colubridae
Amphiesma Dúmeril, Bibron and Dúmeril, 1854

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Amphiesma platyceps (Blyth, 1854)

Oligodon Boie, 1827

Amphiesma sieboldii (Günther, 1860)

Oligodon arnensis (Shaw, 1802)

Amphiesma stolatum (Linnaeus, 1758)

Oligodon taeniolatus (Jerdon, 1853)

Argyrogena Werner, 1924

Platyceps Blyth, 1860

Argyrogena fasciolata (Shaw, 1802)

Platyceps rhodorachis kashmirensis (M. S. Khan and
A. Q.Khan, 2000)

Boiga Fitzinger, 1826

Platyceps rhodorachis ladacensis (J. Anderson, 1871)

Boiga melanocephala (Annandale, 1904)

Platyceps rhodorachis rhodorachis (Jan, 1865)

Boiga trigonata (Schneider, 1802)

Platyceps ventromaculatus bengalensis (M. S. Khan and
A. Q.Khan, 2000)

Coluber Linnaeus, 1758
Coluber karelini karelini Brandt, 1838
Coluber karelini mintonorum Mertens, 1969

Platyceps ventromaculatus indusai (M. S. Khan and A.
Q. Khan, 2000)
Platyceps ventromaculatus ventromaculatus (Gray andHardwicke, 1834)

Enhydris Sonnini and Latreille, 1802
Enhydris pakistanica Mertens, 1959

Psammophis Fitzinger, 1826
Psammophis condanarus (Merrem, 1820)

Hemorrhois Boie, 1826

Psammophis leithii leithii Günther, 1869

Hemorrhois ravergieri (Ménétriés, 1832)

Psammophis lineolatus lineolatus (Brandt, 1838)
Psammophis schokari schokari (Forskål, 1775)

Lycodon Boie, 1826
Lycodon aulicus aulicus (Linnaeus, 1758)

Pseudocyclophis Boettger, 1888

Lycodon striatus Shaw, 1802

Pseudocyclophis persicus (J.Anderson, 1872)

Lycodon striatus bicolor (Nikolsky, 1903)
Lycodon striatus striatus (Shaw, 1802)

Ptyas Fitzinger, 1843

Lycodon travancoricus (Beddome, 1870)

Ptyas mucosus (Linnaeus, 1758)

Lytorhynchus Peters, 1862

Sibynophis Fitzinger, 1843

Lytorhynchus maynardi Alcock and Finn, 1896

Sibynophis sagittarius (Cantor, 1839)

Lytorhynchus paradoxus (Günther, 1875)
Lytorhynchus ridgewayi Boulenger, 1887

Spalerosophis Jan, 1865
Spalerosophis arenarius (Boulenger, 1890)

Natrix Laurenti, 1768

Spalerosophis diadema diadema (Schlegel, 1837)

Natrix tessellata (Laurenti, 1768)

Spalerosophis diadema atriceps (Fisher, 1885)
Spalerosophis schirazianus (Jan, 1865)

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Telescopus Wagner, 1830

Microcephalophis Lesson, 1834

Telescopus rhinopoma (Blanford, 1874)

Microcephalophis cantoris (Günther, 1864)
Microcephalophis gracilis (Shaw, 1802)

Xenochrophis Günther, 1864
Xenochrophis cerasogaster cerasogaster (Cantor, 1839)

Pelamis Daudin, 1803

Xenochrophis piscator piscator (Schneider, 1799)

Pelamis platurus (Linnaeus, 1766)

Xenochrophis sanctijohannis (Boulenger, 1890)
Praescutata Wall, 1921

Family: Elapidae

Praescutata viperina (Ph. Schmidt, 1852)

Bungarus Daudin, 1803
Bungarus caeruleus caeruleus (Schneider, 1801)

Family: Viperidae

Bungarus sindanus razai M. S. Khan, 1985

Daboia Gray, 1842

Bungarus sindanus sindanus Boulenger, 1847

Daboia russelii russelii (Shaw and Nodder, 1797)

Naja Laurenti, 1768

Echis Merrem, 1820

Naja naja (Linnaeus, 1758)

Echis carinatus (Schneider, 1820)

Naja oxiana (Eichwald, 1831)

Echis carinatus astolae Mertens, 1969
Echis carinatus multisquamatus Cherlin, 1981

Family: Hydrophiidae

Echis carinatus sochureki Stemmler, 1964

Astrotia Fisher, 1856
Astrotia stokesii (Gray, 1846)

Eristicophis Alcock and Finn, 1896
Eristicophis macmahonii Alcock and Finn, 1897

Enhydrina Gray, 1849
Enhydrina schistosa (Daudin, 1803)

Macrovipera Reuss, 1927
Macrovipera lebetina obtusa (Dwigubsky, 1832

Hydrophis Latreille, 1802
Hydrophis caerulescens (Shaw, 1802)

Pseudocerastes Boulenger, 1896

Hydrophis cyanocinctus Daudin, 1803

Pseudocerastes bicornis Wall, 1913

Hydrophis fasciatus (Schneider, 1799)

Pseudocerastes persicus (Duméril, Bibron, andHydrophis lapemoides (Gray, 1849)

Duméril, 1854)

Hydrophis mamillaris (Daudin, 1803)
Hydrophis ornatus (Gray, 1842)

Family: Crotalidae

Hydrophis spiralis (Shaw, 1802)

Gloydius Hoge and Romano-Hoge, 1981
Gloydius himalayanus (Günther, 1864)

Lapemis Gray, 1835
Lapemis curtus (Shaw, 1802)

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Literature Cited
Anderson, S. C. and A. E. Leviton. 1966. A review of thegenus Ophiomorus (Sauria: Scincidae), withdescriptions of three new forms. Proceedings
California Academy of Sciences. 4th Ser. 33:499534.
Anderson, S. C. and A. E. Leviton, 1967. A new speciesof Eremias (Reptilia: Lacertidae) from Afghanistan.
Proceedings California Academy of Sciences(64):1-4.
Anderson, S. C. and A. E. Leviton.1969. Amphibians
and reptiles collected by the Street Expedition
to Afghanistan, 1965. Proceedings California
Academy of Sciences, (4th Ser.) 37:25-56.
Anderson, S. C., and S. A. Minton. 1963. Two noteworthy herpetological records from the Thar Parker
Desert, West Pakistan. Herpetologica, 19:152.
Baig, K. J. 1988. New record of Agama nuristanica(Sauria:Agamidae) from Pakistan. Biologia(Lahore) 34:199-200.
Baig, K.J. 1989. A new species of Agama (Sauria,
Agamidae) from northern Pakistan. Bulletin
Kitakyushu Museum Natural History 9:117-122.

2004

Boulenger, G. A. (1897): A new krait from Sind
(Bungarus sindanus). Journal Bombay Natural
History Society 9:73-74.
Cherlin, V. A. 1981. A new saw-scaled viper, Echis multisquamatus, sp. nov. from southwestern andMiddle Asia. Trudy, Zoology Institut Akademy
Nauk USSR, Leningrd 101:92-95.
Cherlin, V. A. 1983. New facts on the taxonomy ofsnakes of the genus Echis. Vest. Zool. 1983:42-46.
(English tranalation: Smithsonian Herpetological
Information Ser. No. 61, 1984).
Dubois, A. and Khan, M. S. 1979. A new species offrog (genus Rana, subgenus Paa) from northern
Pakistan (Amphibia, Anura). Journal Herpetology
13:403-410
Fitzinger, L. J. 1823. In: Lichtenstein, M. H. C.,
Verzeichniss der Doubletten des zoologischen
Museums...UniversitatzuBerlinnebst
Beschreibung vieler biserunbekannter Arten vonSaugethieren, Vogeln, Amphibien und Fischen.
Berlin.
Fitzinger, L. J. 1843. Systema reptilium. Fasciculus
Primus. Vienna.

Baig, K. J. 1998. A new species of Tenuidactylus(Sauria: geckonidae) from Balochistan, Pakistan.
Hamadryad 23(2):127-132.

Fühn, I. E. 1969. Revision and Redefinition of the genus
Ablepharus Lichtenstein, 1823 (Reptilia,
Scincidae). Revue Rouma. De Biologia, ser. Zool.,
14:23-41.

Baig, K. J. 1999. Description and ecology of a new sub
species of black rock agama, Laudakia melanura(Sauria: Agamidae) from Balochistan, Pakistan.
Russian Journal of Herpetology 6(2):81-86.

Golubev, M, and N. Szczerbak. 1981. A new species ofGenus Gymnodactylus Spix, 1823 (Reptilia, Sauria,
Gekkonidae) - from Pakistan Vestnik Zoology, 1981
(3):40-45.

Baig, K.J. and W. Bohme. 1996. Description of two newsubspecies of Laudakia pakistanica (Sauria:
Agamidae). Russian Journal Herpetology 3:1-10.

Griffith, H., A. Ngo, and R.W. Murphy. (2000): A cladistic evaluation of the cosmopolitan genus Eumeces
Wiegmann (Reptilia, Squamata, Scincidae).
Russian Journal Herpetology, Moscow; 7(1):1-16.

Baig, K.J. and L. Gvozdik. 1998. Uperodon systoma
Schneider: record of a new microhylid frog from
Pakistan. Pakistan Journal Zoology 30:155-156.

Günther, A. 1864. The reptiles of British India. Oxford& IBH Publishing Co. Bombay.

Boulenger, G. A. 1890. Fauna of British India, including
Ceylon and Burma. Reptilia and Batrachia.
London.

Ingoldby, C.M. 1922. A new stone gecko from
Himalayas. Journal Bombay Natural History
Society 28:1051.

Boulenger, G. A. 1896. On the lizards of the genus
Eremias, section Boulengerina. Proceedings of the
Zoological Society of London, 1896: 920-930.

Jan, . 1865. In: De-Filippi, Viagg. in Persia 356.
Khan, A.Q. and M.S. Khan, 1996. Snakes of State of2004

Asiatic Herpetological Research

Azad Jammu and Kashmir. Proc. Pakistan
Zoological Congress 16:173-182.
Khan, M. S. 1980. A new species of gecko from northern Pakistan. Pakistan Journal Zoology 12(1): 1116.
Khan, M. S. 1984a. Validity of the natricine taxon Natrix
sancti-johannis Boulenger. Journal Herpetology
18:198-200.
Khan, M. S. 1984b.
Rediscovery and validity of
Bungarus sindanus Boulenger. The Snake 16:4348.
Khan, M. S. 1985b. Taxonomic notes on Bungarus
caeruleus Schneider and Bungarus sindanus
Boulenger. The Snake 17:71-78.
Khan, M. S. 1988. A new Cyrtodactylid gecko from
northwestern
Punjab,Pakistan.
Journal
Herpetology 22:241-243.
Khan, M. S. 1989. Rediscovery and redescription of thehighland ground gecko, Tenuidactylus montiumsalsorum Annandale, l913. Herpetologica 45:46-54.
Khan, M. S. 1991 d. A new Tenuidactylus gecko fromthe Sulaiman Range, Punjab, Pakistan. Journal
Herpetology 25:199-204.
Khan, M. S. 1992. Validity of the mountain gecko
Gymnodactylus walli Ingoldby, 1922. Herptology
Journal 2:106-109.
Khan, M. S. 1993a. A new angular-toed gecko from
Pakistan, with remarks on the taxonomy and a key
to the species belonging to genus Cyrtodactylus(Reptilia: Sauria: Geckkonidae). Pakistan Journal of
Zoology 25:67-73.
Khan, M. S. 1993b. A new sandstone gecko from Fort
Munro, Dera Ghazi Khan district, Punjab, Pakistan.
Pakistan Journal of Zoology 25:217-221.
Khan, M. S. 1994. Validity and redescription ofTenuidactylus yarkandensis (J. Anderson, 1872).
Pakistan Journal of Zoology, 26(2):139-143.
Khan, M. S. 1997a. A new toad from the foot of Siachin
Glacier, Baltistan, northeastern Pakistan. Pakistan
Journal Zoology 29(1):43-48.

Vol. 10, p. 199

Khan, M. S. 1997b. A new subspecies of common skittering frog Euphlyctis cyanophlyctis (Schneider,1799) from Balochistan, Pakistan. Pakistan Journal
Zoology 29(2):107-112.
Khan, M. S. 1997c. Validity, generic redesignation, andtaxonomy of western rock gecko Gymnodactylus
ingoldbyi Proctor, 1923. Russian Journal
Herpetology 4(2):83-88.
Khan, M. S. 1998. Notes on Typhlops diardi Schlegel,
1839, with description of a new subspecies (squamata, Serpentes, Scolecophidia). Pakistan Journal
Zoology 30(3):213-221.
Khan, M. S. 1999a. New species of blind snakes ofgenus Typhlops from Azad Kashmir and Punjab,
Pakistan (Serpentes: Typhlopidae). Russian Journalof Herpetology, 6(3):231-240.
Khan, M. S. 1999b. Typhlops ductuliformes, a new
species of blind snakes from Pakistan and a note on
T. porrectus Stoliczka, 1871, (Squamata, Serpentes,
Scolecophidia). Pakistan Journal of Zoology,
31(4):385-390.
Khan, M. S. 2000. Redescription and generic redesignation of Gymnodactylus stoliczkai Steindachner,
1869. Pakistan Journal Zoology 32(2): 157-163.
Khan, M. S. 2001a. Taxonomic notes on angular-toedgekkota of Pakistan, with description of a new
species of genus Cyrtopodion. Pakistan Journal
Zoology33(1):13-24.
Khan, M. S. 2001b. Notes on Cranial-Ridged Toads ofPakistan and Description of a new subspecies(Amphibia: Bufonidae). Pakistan Journal of
Zoology 33(4):293-298.
Khan, M. S. 2003a. Notes on circum Indus geckos ofgenus Cyrtopodion (Squamata: Gekkonidae).
Gekkota 4:
Khan, M. S. 2003b. Questions of generic designation ofangular-toed geckos of Pakistan with descriptionsof three new genera (Reptilia: Gekkonidae). Journalof Natural History and Wildlife (Karachi), Vol. 2(2):9-29.
Khan, M. S., and K. J. Baig. 1992. A new tenuidactylid
gecko from northeastern Gilgit Agency, North
Pakistan. Pakistan Journal of Zoology, 24:273-277.

Vol. 10, p. 200

Asiatic Herpetological Research

Khan, M. S. and M. R. Z. Khan. 1997. A new skink fromthe Thal Desert of Pakistan. Asiatic HerpetologicalResearch 7:61-67.
Khan, M. S., and R. Tasnim. 1989. A new frog of thegenus Rana, subgenus Paa, from southwestern
Azad Kashmir. Journal Herpetology 23:419-423.
Khan, M. S., and R. Tasnim. 1990. A new gecko of thegenus Tenuidactylus from northeastern Punjab,
Pakistan, and southwestern Azad Kashmir.
Herpetologica 46:142-148.

2004

Pakistans. 2. Nachtrag. Senckenberg Biology52(12):7-15.
Mertens, R. 1972. Nachtrage zum Krokodil-Katalog dersenckenbergischen Sammlungen. 3. Nachtrag.
Senck. biol. 53:21-35.
Mertens, R. 1974. Die Amphibien und Reptilien WestPakistans. Senckenb. biol. 55(1-3):35-38.
Minton, S. A. 1966. A contribution to the herpetology ofWest Pakistan. Bulletin of the American Museum of
Natural History 134(2):31-184.

Khan, M. S., and Khan, A. Q. 2000. Three newsubspecies of snakes of genus Coluber from
Pakistan. Pakistan Journal of Zoology 32(1):4952.

Minton, S. A. 1966. A contribution to the herpetology ofWest Pakistan. Bulletin American Museum of
Natural History 134(2):31-184.

Khozatsky, L.I., and M. Mlynarski. 1966. Agrionemys nouveau genre de tortues terrestres [Testudinidae].
Bull. Acad. Pol. Sci. Ser. Sci. Biol., [2]:123-125.

Minton, S. A., and Anderson, S. 1965. A new dwarf
gecko(Tropiocolotes)
fromBaluchistan.
Herpetological 21:59-61.

Leviton, A., and Anderson, S. C. 1972. Description of a
new species of Tropiocolotes (Reptiliua: geckonidae) with a revised key to the genus. Occasional
Papers of the California Academy of Science 96:17.

Minton, S. A., S. Anderson, and J. A. Anderson. 1970.
Remarks on some geckos from Southwest Asia,with description of three new forms and a key to thegenus Tropiocoloties. Proceedings of the California
Academy Sciences, (4th Ser.) 37:333-362.

Malnate, E. V. 1966. Amphiesma platyceps (Blyth) andAmphieasma sieboldii (Günther): sibling species(Reptilia:Serpentes). Journal of Bombay Natural
History Society 63:1-17.

Murray, J. A. 1884. The vertebrate zoology of Sind.
London & Bombay.

Malnate, E. V., and S. A. Minton. 1965. A redescriptionof the natricine snake Xenochrophis cerasogaster,with comments on its taxonomic status.
Proceedings of Academy National Science,
Philadelphia 117:19-43.
Mertens, R. 1954. Uber die Rassen des Wustenwarans(Varanus griseus). Senckenberg Biology 35:353357.
Mertens, R. 1969a. Die Amphibien und Reptilien WestPakistans. Stuttgart Beitrag Naturkundi 197:1-96.
Mertens, R. 1969b. Eine neue Rasse derDachschildkrote, Kachuga tecta. Seckenberg
Biology 50:23-30.
Mertens, R. 1970. Die Amphibien und Reptilien WestPakistans. Stuttgart Beitrag Naturkundi 216:1-5.
Mertens, R. 1971. Die Amphibien und Reptilien West-

Murray, J. A. 1886. The zoology of Sind. London &Bombay.
Murray, J. A. 1892. The zoology of Baloochistan andsouthern Afghanistan. London & Bombay.
Rastegar-Pouyani, N. 1999. Two new subspecies ofTrapelus agilis complex (Sauria:Agamidae) fromlowland southwestern Iran and southeastern
Pakistan. Asiatic Herpetological Research 8:90101.
Schmidtler, J. J. and J. F. Schmidtler. 1969. Uber Bufo
surdus; mit einem Schlussel und Anmerkungen zu
den ubrigen Kroten Irans und West-Pakistans.
Salamandra 5:113-123.
Schätti, B., and U. Utiger. 2001. Hemerophis, a new
genus for Zamenis socotrae Günther, and a contribution to the phylogeny of Old World racers, whip
snakes, and related genera (Reptilia: Squamata:Colubrinae). Revue Suisse de Zoologie 108(4):919948.

2004

Asiatic Herpetological Research

Smith, M. A. 1931. The fauna of British India, including
Ceylon and Burma. Reptilia and amphibia. Vol. I:
Loricata, Testudines. Taylor and Francis Ltd.
London.
Smith, M. A. 1935. The Fauna of British India, including Ceylon and Burma. Reptilia and amphibia. Vol.
II: Sauria. Taylor and Francis Ltd. London.
Smith, M. A. 1943. The fauna of British India, Ceylon,
andBurma. Reptilia and amphibia. Vol.
III:Serpentes. Taylor and Francis, London.
Steindachner, F. 1869. Reptilia. In: Reise der osterreichischen
Fregatte
Novara...Zoologischer
Theil.Vienna, 1:1-98.
Stock, M., M. Schmid, C. Steinlein, and W. R. Grosse.
1999. Mosaicism in somatic triploid specimens of
the Bufo viridis complex in the Karakoram withexamination of calls, morphology and taxonomic
conclusions. Italian Journal of Zoology 66:215-32.
Szczerbak, N. N. 1991. Eine neue Gecko-Art aus
Pakistan: Alsophylax (Altiphylax) boehmi sp. nov.
Salamandra 27(1):53-57.
Tuck, R. G. 1971. Rediscovery and redescription of the
Khuzistan dwarf gecko Microgecko helenae
Nikolsky (Sauria:Geckkonidae). Proceedings of the
Biological Society of Washington 83:477-482.
Webb, R. G. 1980. The identity of Testudo punctata
Lacépède, 1788 (Testudines, Trionychidae).
Bulletin du Muséum National d’Histoire Naturelle.
Paris. 4(2):547-557.

Vol. 10, p. 201