Journal ol NannopJankton
Research

A PubUcadon of the International Nannoplankton Assocladon

JOURNAL OF NANNOPLANKTON RESEARCH
Published by the International Nannoplankton Association
Volume18

Number2

November 1996

NEWS AND GOSSIP Compiled byJackie Bumett ........... ............................ ........................................... ......... ... ..........
NEWP()SffiONS ......................... ....... ............................................ ..... ............... .................... .......... ....... ..... .. ............
FOR1HCOMIN'G EVENTS............ ....................................................................................................................... ... .....
7thiNAConference.................................................................. .'.................... .. ........... .... .................................
Copenhagen Conference Proceedings Volume........................................................... ...... ....... .. ........... ..... ..
Recoveries '97 Conference.. ... ............................................................. ..... ...... ........... ... ... .. .. ............................

42
42
42
42
42
42
E-Inaillist. .......................................... .......................... ...................................................................................... 42
PASTEVENTS.................................................................................................................. ................ ........ ...... ............... 42
6th International Global Emiliania Modelling Initiative Workshop.... .............. ....... ....... ...... ...... ............. 42
6th INA Conference Workshop (continued)............. ................. .... .......... ..... .......... ... .................................. 44
Palaeopelagos Meeting................................ ............ ... ...................... .......... .. ..... .............................. ................ 45
AREMlNDER................................................................................... ... ....... ................................................. .. ............... 45
LETTERS......................................................................................................... ... ... ... ... ...... ............................. ................. 45
B()()KS & REVIEWS.......................................................................................................... ........ ... .. ... ...... ........ ............. 46
Microfossils and Oceanic Environments Tim Bra lower................ ... :.............. ..... ............ .... ............. ..... .. ... . 46

OCEAN DRilLING PROGRAM UPDATE John Firth....... ...... .................................................... .......... .......................... 47
THEICBN: TIIINGSYOUNEEDTOKNOW-14

ShirleyE. van Heck. ...... ......... .... ............. ....... ........... ................ ..... 48

TREATING STRONGLY-LI1HIFlED ROCKS WI1HULTRASONIC WAVES
Jorijntje Henderiks & Patrizia Ziveri........... .......... ... ................ .............. ............. .... :.......... .. .... .... ..... .... ..... .... .... .... .. ... .... .... .. 51
NEW NAMES FOR OLD: TAXONOMIC CLARIFICATION OF SOMEEARLY CRETACEOUS NANNOFOSSILMARKER-SPECIES David Rut/edge & Paul Bown........ .. .............. ........ 53
UPPERZANCLEAN SILICOFLAGELLATES FROMMlLOS ISLAND(CYCLADES, GREECE) Dimitris Frydas.. ...... ..... .. ...... ...... .. ..... ...... .. .. .. .. .... .. ..... .. .. .. .. ............... .. ........ .... .. .. ... .......... .. .. .. .. .. . 61
BIBLIOGRAPIN AND TAXA OF CALCAREOUS NANNOPLANKTON- 27
Wi//iam G. Siesser ... .. .. .. .. .. .. .. .. .. ... .. .. .. .. .. .. .. .. .. ... .. .. .. .. .. .. .. .. ... .. .. .. .. .. .. .. .. .. ...... ... .. .. .. .. .. .. .. ... .. .. .. .. .. .. .. .. ... .. .. .. .. .. .. .. .. ... .. .. .. .. .. .. .. .... . 69
INA~ERSHIP CfiANGESKevin Cooper.......................................................................... ... ....................... ... ........ 84==========================41==========================

Journal of Nannoplankton Research, 18, 2, 1996.

News and Gossip, p. 42 - 46.

NEWS AND GOSSIP
Compiled by Jackie Burnett on behalfofthe INA Committee

NEWPOSffiONS
Harald Andruleit, formerly ofKiel, was successful in his
bid for the nannopalaeontology position at the German
Geological Survey.
FORmCOMING EVENTS
7th International Nannoplankton Association
Conference, Puerto Rico 1998
It seems like we just get over one when the next starts toloom on the horizon- yes, I'm talking about the next INA
Conference! At Copenhagen it was decided that we would
finally let Amos Winter drag us over to Puerto Rico forstm, sea and science. So, make space in your diaries for
February 1998 (hopefully outside of the wet and windy
season). The meeting will be held 'most likely' at La
Parguera, Puerto Rico. Amos needs 125 people to attend
to make it economic. Airfares from Europe should be lessthan $1000 (Amos knows of someone who paid $600) but it
may be worthwhile to check out fares to Miami, since this isa major holiday destination and it may be possible to getcheap flights through tour operators, and then get a flightfrom there (Miami-Puerto Rico around $200). From San
Francisco, flights cost around $650. Very cheap! The hotel
should cost around $40-50, shared occupancy. So, startsaving and applying for conference funding!
on behalfofAmos WinterCopenhagen Conference Proceedings Volume
Dave Jutsonlthe DGGU has finally produced author contribution guidelines for the conference volume. Contributions will appear in the Geological Survey of Denmarkand Greenland Bulletins. Attendees of the conference
should have received a letter containing the guidelines
from Dave himself around October. If you presented a paper at the conference and wish to contribute to the volume
but did not receive these guidelines, then you must inform
either Dave or myself immediately. Dave looks forward toreceiving your manuscripts and apologises for the delay.
Jackie Burnett

Recoveries '97
Final meeting ofthe UNESCO IGCP Project 335'Biotic Recoveries from Mass Extinctions'12th-14th September, 1997, Prague, Czech Republic
Call for participation- call for papen
About the project: In the history of the Earth (including
the Recent), numerous events of ecosystem collapses occurred that were followed by recoveries and origination of
new ecosystems. This significant transformation could be
realised in numerous ways. The project aims to be a platform for the study of survival and recovery of the biosphere, and restructuring of global environments, follow-

ing mass extinctions. This international project is headed
by Douglas H. Erwin, Smithsonian Institution, Washington,
DC, and Erie G. Kauffinan, University of Colorado, Boulder. Over sixty countries are involved in the project. The
meeting should bring together palaeobiologists, palaeontologists, biologists, ecologists, systems theorists, and otherpersons that are interested in the topic. The conference
will be held under the auspices of the Geological Institute,
Academy of Sciences, and is being organised by Petr
Cejchan & Jindfich Hladil, Geological Institute, Academyof Sciences, Rozvojova 135, CZ 165 02, Praha6 Lysolaje,
Czech Republic.
Deadlines: Pre-registration by January 31st, 1997
(printedfonn, or WWW: http://www.glicas.crJconf/recovery/preform.htm); abstracts by May 31st, 1997.
For further information: Please direct all your correspondence related to the conference to the ConferenceManager: Petra Hovorkova, Recoveries '97, Eurocongress
Centre, Budejovic.ka 15, CZ 140 00 Praha 4. For conference
update and details on the programme please consult:
http://www.glicas.cz/coof/recovery/recovery.htm
ore-mail: recovery@gli.cas.cz
E-maillist
I would like to compile an up-to-date list of e-mail addresses
for publication in the next issue. Although I know a few, I

am sure there are many more which I am unaware of. So,
could you please e- mail me and tell me you would like to be
included on the list. Thank you.
j. burnett@ucl.ac. uk

PAST EVENTS
6th International Global Emiliania Modelling Initiative(GEM) Workshop
Chateau de Blagnac, Cabara, France
September 4th-8th, 1996,
Convenor: Patrizia Ziveri.
Hosts: Dorien and Jan van Hinte'Global Emiliania Modelling Initiative:
the geological aspects'
Seven years ago, in September 1989, an interdisciplinary
meeting was held at the Chateau de Blagnac to discuss theimpact on global climate of a single organism, thecoccolithophorid alga, Emiliania huxleyi. This event
marked the creation of the Global Emiliania Modelling
Initiative, with the objective of developing Emiliania as amodel organism for studying the general role of the marine
pelagic biota in the climate system and in global dynamics.
GEM integrates experimental, observational and modelling
research on the Emiliania phenomenon, and is designed
to produce a nested suite of models at the molecular,organismal, ecological and global level of organisation,
ranging in time-scale between milliseconds and megayears.

========================= 42 =========================

Journal of Nannoplankton Research, 18, 2, 1996.

News and Gossip, p. 42 - 46.

This initial meeting triggered extensive collaborations, including more than 50 scientists from a wide range of institutions in many countries, with funding from both the EU
and national research councils. The results of this work
have been discussed at four subsequent meetings in
Blagnac, and in international conferencesheld in Bergen(Norway), Plymouth and London (UK). In total, more than
100 papers have been published, many Of which were collected together in the following volumes:
The Haptophyte Algae. 1994. J.C. Green & B.S .C.
Leadbeater (Eds). The Systematics Association Special Volume 51, Oxford University Press.
The 1992 Norwegian Emiliania huxleyi experiment. 1994.
B.R. Heirndal (Ed.). Sarsia, 79(4).
EHUX (Emiliania huxleyi). 1996. J.C. Green & R. Harris(Eds). Journal ofMarine Systems, 9(1-2).

Now GEM, and particularly the geological branch of GEM,
has moved from E. huxleyi as a target organism to include
all coccolithophores.
This year's GEM meeting, the 6th International Global Emiliania Workshop, concentrated on coccolithophore dynamics at the geological scale. The major goals of
this informal meeting were to establish working relationships concerning the following topics: ( 1) to quantify and

DMS

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1
archive-map on a global scale the variety of the coccolithophorecommunities in the Recent ocean by season or month, anddetermine the coccolith-CaC03 export fluxes; (2) to quantify and map the coccolithophore-induced CaC03 transport and dissolution characteristics in the water-column
and top sediments, and define thanatocoenosis composition in different climate regimes; (3) to develop and test
models representing the regional distribution, sedimentation behaviour and archive preservation of coccolithophore-induced CaC03 . In addition to providing an understanding of the Recent system, these models sllould serve
as a key to interpret the deep-sea archive over geologicaltime. The integration of these geologically oriented models with the biological GEM models is also to be implemented.
Following the traditions of the previous GEM meetings, overview talks by Peter Westbroek (Leiden University, NL) andPatriziaZiveri (Free University Amsterdam,
NL) provided a general picture of the interactions between
coccolithophores and climate, from the molecular biological to global geological levels. A range of more specialised
topics were then reviewed: coccolithophore production,
fluxes, sedimentation (Michael Knappertsbusch, Natural
History Museum, Basel, SW); overview of work on
coccolith fluxes in the Pacific Ocean (Kozo Takahashi,
Hokkaido Tokai University, Sapporo, J); the practicalities
of collecting coccolithophorids in sediment traps (GeertJan Drummer, NIOZ, NL); Saharan dust input and its
impact on phytoplankton production (Cemal Saydam,
Middle East Technical University, Turkey); prymnesiophyte molecular palaeontology (Gerard J.M.
Versteegb, NIOZ, NL); modelling marine snow dynamics(Gavin Ruddy, PML, UK); coccolith carbonate mass estimation (Jeremy R. Young, Natural History Museum, London, UK); the contribution of E. huxleyi to mass fluxes(Luc Beaufort, LGQ-CNRS, FR). Great attention was given
to the poster session, allowing us to focus on the resultsfrom different researches. The workshop was attended by
25 invited participants, including the majority of the specialists on sediment trapping of coccolithophores, anditgave us a unique opportunity to share experiences, suggestions and data with biologists and modellers. The 4-day
workshop was very productive: open discussion, shortand long-term plans have been made.
The main aim of the two workshops on 'Sediment
trapping of coccolithophorids, collection and preparation
procedures in the field and in the laboratory' and'Coccolithophore counting and carbonate flux calculations'
was to improve quantitative techniques and protocols toallow more-accurate and consistent estimation of thecoccolith carbonate mass-fluxes per species. Numerous
aspects where discussed: from the dispersion of the particles with chemical agents, through size fractionation due
to sieving, and mechanical destruction of particles. Practical outcomes we hope to achieve include: (1) developmentof a document on standard preparation and counting methods, a first draft of which is in preparation by Pat:rizia Ziveri;(2) An intercalibration experiment-will be carried out by============================43 ============================

Journal of Nannoplankton Research, 18, 2, 1996.

News and Gossip, p. 42 - 46.

(~_H_o_l_o_ce_n_e_h_is_to_r_y_a_n_d_c_li_m_a_te__®==s:...____..J

coccoliths while they sink to the sea
floor and settle on the ocean floor.
Figure 1 (page 43) shows the approach of investigating the coccolith
record in the sediment archive as aproxy for POC production and DMS
emission over time-scales varying
from seasons to geological time, andin different climate regimes. During
the workshop, the practicalities ofthis approach were widely discussed.
Different levels must be considered,including export of coccolith carbonate production recorded below
the euphotic zone; the fate of thismaterial whilst it sinks down throughthe dark ocean and settles on theocean floor; transformation and dissolution at the sea floor.
During our final discussion in
Blagnac, nine different major topics
were defined for near-future collaboration and for discussion at the next
Blagnac meeting (scheduled for
September, 1997) - as outlined in
Figure 2.
Above all, the meeting had ahighly stimulating atmospherenotonly because of the great expertise of the participants, but also very
importantly due to the hospitality
and friendship ofDorien and Jan van
Hinte at their Chateau de Blagnac.

Fig. 2

Patrizia Ziveri, Geomarine
Research Centre, Free University,
Amsterdam, zivp@geo. vu.n/

Coccoliths as a proxy to:total biomass, .
calcium carbonate,
andDMS

Modelling geographical maps of coccolnh andbiomarker productivity

2

Coccouth and blomarker production
as influenced by:

Oust
Ocean conditions

Bogs

®
(~

(:§)

Informed counting and mapping

Modelling geographical maps of coccolithand biomarker transformation

Mapping and laboratory studies on
coccoltth and biomarker transfonmation

6th International Nannoplankton Association
around 20 specialists, in order to provide a practical test ofConference, Copenhagen, 2nd-7th September, 1995
the consistency of our results. Two samples have been
Copenhagen Conference Workshops (continued)
selected, a sediment trap sample provided by Geert-JanA
reporton the goings-on at the Living/Quaternary
Brummer (NIOZ) and a Holocene sediment sample provided
Nannoplankton
Workshop, held at the Copenhagen conby Jacques Giraudeau (CNRS, Bordeaux, FR). Each oftheseference,wasnotsolicited in time for the last issue. So, a
has been distributed among interested participants andlittle late but still interesting, here is Ric's summary:other relevant colleagues who are supposed to estimate
individual species abundances per gram of sediment. In· Living/Quaternary: The workshop was held in two sesformation on this intercalibration experiment will be posted
sions and dealt with ( 1) problematic genera in the Quateron the 'coccoliths' e-maillist. The comparison of thesenary record, and (2) Coccolithus pelagicus and microresults should at the least allow an estimation of the error scope work. However, only part of the discussion derived
between different preparation and counting methods; (3) from the first session will be mentioned here.
Within the organisation of the international program, aPontosphaera: The coccoliths of this genus arecompilation of all available sediment trap data will be carrarely found in sediments (or the water-column), but nevriedout.
ertheless still need to be identified correctly. There are two
An important issue for the international geologicalmain problems facing the nannoworker. Firstly, there is a
GEM network is that, in order to understand the impact of need to recognise the discoliths of Scyphosphaera
cocco1ithophores on the oceanic carbon cycle, it is neces- apsteinii, and secondly, one must allow for a reasonable
sary to understand, quantify, and predict the fate of amount of variation in the discolith morphology of each========================~44

News and Gossip, p. 42 - 46.

Journal of Nannoplankton Research, 18, 2, 1996.

species. For instance, a few extra pores should not necessarily constitute a new species. A quick glance at the DSDP/
ODP volumes shows that the problem of splitting has occurred within this genus - I have found at least 80 fossil'species' from the literature (and there are probably more).
This is an enormous number, considering that we have
only five recognised extant species. As the genus originated in the Palaeogene, some of these species will be partof the evolutionary series, however, many are clearly fromthe Neogene/Quatemary and are possibly synonymouswith other species.
Syracosphaera: The coccoliths of this genus are
difficult to identify with the light-microscope, and are usually visible as just a dull ring (an exception to this is S.
pulchra). At present, there are over 20 extant species of
Syracosphaera, and in the Syracosphaeraceae there areseveral genera with caneolith-type coccoliths. From posters displayed at recent INA meetings, it is obvious thatmany of the extant species have records back into the Late
Miocene. However, isolated caneoliths (and cyrtoliths) are
relatively difficult to identify, even with the SEM, if one
does not have knowledge of the coccospheres of extant
species. For this reason, many taxa have been effectively
redescribed. Once again, a recent review of Syracosphaeraspp., this time from Loeblich & Tappan and INA bibliographies, came up with about 70 species (mostly living) which
are not considered in recent check-lists of extant taxa. It isalso a fact that many workers refer to these isolated
coccoliths in the sediments as Syracosphaera sp. and so
the DSDP /ODP volumes are probably littered with specimens which could have been identified.
The workshop members have decided within thenext year to gather LM and SEM micrographs ofPontosphaera and Syracosphaera spp. from Quaternary
sediments, with the aim ofbuilding up an atlas of the fossil
species and comparing them with extant data. It is hopedthat in this way we can progress in our understanding of
how many species exist in the Quaternary and how far backcertain families can be traced. If anyone is interested injoining this project then please contact me.
RicJordan
sh08J@kdw.kj.yamagata-u.ac.jp

Palaeopelagos Meeting
Rome, June, 1996
At the beginning ofJune the Palaeopelagos (Group ofltalian Palaeontologists) meeting was held in Rome. As usual,a meeting of the Italian nanno-workers was also organized.
Our main purpose is to keep everybody updated on theongoing research activities of all colleagues and to favour
cooperation. As a general task we would like to collect, in
an atlas, all the available data on Italian nannofossil
stratigraphy. In order to do this, we are divided into working groups, subdivided for time periods. Each group meets
periodically and will hopefully produce reports on the performed activity.
Andrea Fiorentino

otello@epidote.dmp. unipd. it===========================45

A REMINDERThe JNR bibliographers should be providing a comprehensive listing of all nannoplankton and silicoflagellate
papers being published around the world (not everyone
has the facility to scan the WWW databases). They can
only do this with your help - please send your reprints to
the relevant bibliographer so that they can broadcast yourwork. You might also check to see if all of your past references have appeared in the JNRIINA Newsletter. If you
subscribe to the 'coccoliths' mailing list, you will alreadybe aware that Wuchang Wei has recently compiled a bibliography (nanno-related works since 1990) which you canfind on his web site (http://wwei.ucsd.edu).
Jackie Burnett:U:TIERS
An open letter to all calcareous nannofossil specialists
The following letter was sent to the Editor and is here reproduced in its full and unedited form. Any comments may
be directed either to Martin or Jeremy, or to the Editor if
you wish them to be published (I think the contents should
inspire some discussion in the Journal).

The Natural History Museum
Cromwell Road
LondonSW75BD
October 1996
Martin Jakubowski has recently taken up the position of
Head of Curation in the Department of Palaeontology at
The Natural History Museum in London, England after
spending almost 14 years working in the oil industry with
The Robertson Group, Shell Exploration and Production(U.K.) and Shell International. As some of you will alreadyknow during that time Martin's responsibilities centredaround the analysis and evaluation of calcareous
nannofossil assemblages obtained from exploration andproduction well samples. They are now somewhat different, being concerned with the management of the curators
at the Musewn and it's fossil collections as well as carryingout research. In addition to the overall responsibility for
curation in the Palaeontology Department, Martin specifically manages the micropalaeontological collections. It isconcerning these collections and in particular the calcareous nannofossil collection, that we are writing to you.
In the Palaeontological Department the nannofossil
activities have now been effectively doubled and together
we would like to extend this activity further by establishinga collection of type species and/or sample material. This is
in line with the Museum's mission statemen\; "to maintainand develop its collections and use them to promote thediscovery, understanding, responsible use and enjoymentof the natural world". Already some authors have agreed
to donate material to the museum. Remember that Article
37.5 of the I.C.B.N. states that "For the name of a new
species or infraspecific taxon to be published on or after 1
January 1990 whose type is a specimen or unpublishedJournal of Nannoplankton Research, 18, 2, 1996.

illustration, the hetbarium or institution in which the type
is conserved must be specified." Everyone needs to consider a repository and so we are asking you to place The
Natural History Museum high on your list. Our ultimate
goal is to develop the museum as a major centre (at least in
Europe) for calcareous nannofossil collections.
We are therefore making a request (plea, beg orwhatever you wish to call it) for fellow nannofossil specialists to send material to the museum in order to enhance andfurther develop the collection. We would gladly accept
original slides containing type species and/or type sample
material together with, ifpossible, a copy of the paper documenting the supporting data. In those case where only
sample material can be offered we will guarantee to prepare
slides and place these within the collections. All donations
will be suitably acknowledged and the information relevantto the donation recorded on the Palaeontology Department's digital database. In addition, we will provide regular updates on the state of the collection throughthe Journal ofNannoplankton Research and, in line with Museum
policy, on the Internet.
In making your donation you will ensure that yourwork and collection are preserved and made available for
future use by the next generation of nannofossil workers.
If you require any further information please do
not hesitate to contact us.
Yours Faithfully

Mr. Martin Jakubowski
Head of Curation & Calcareous Nannofossil Research
Dep~en~ of Palaeontology .. ~ ___ .
E-mail: maiJ@nhmac.uk
I." ' · ...] o.~!S\t.....
Tel: +44-(0)171 938 8902=--- ...:.:=:-..:.:,___

Dr. Jeremy Young
Calcareous Nannofossil Research
Department of Palaeontology
E-mail: jy@nhm.ac.uk
Tel: +44-(0)171938 8996

BOOKS & REVIEWS

Microfossils and Oceanic Environments
Edited by Alicia Moguilevsky & Robin Whatley (1996)
University ofWales, Aberystwyth-Press: 434pp.
ISBN 0 903 878 74 7

£25 Softback
Reviewed by Tim Bra/ower, University ofNorth
Carolina, USA
This volume serves as a Proceedings for the International
Conference on "ODP and the Marine Biosphere" held in
Aberystwyth on 19th-21st April, 1994. The volume contains 25 papers dealing with the application ofbenthic andplanktonic microfossils in palaeoceanography, newmicrofossil techniques and applications, ecology,
biostratigraphy and evolution, and includes studies of
ostracods, planktonic and benthic fomminifera, radiolaria,
diatoms and calcareous nannofossils.
Seven separate papers address aspects of nannoplankton research. Most of these papers concern applica-

News and Gossip, p. 42 - 46.

tions ofnannofossils to palaeoceanography. Ric Jordan
and others describe a combined study of coccolith andalkenone stratigmphy for the last 130,000 years of an ODP
site off NW Africa. The authors recognize Heinrich andother cooling events in the alkenone record and postulate
relationships between the relative abundance of major
nannofossil species, temperature, depth and productivity.
This is a superb paper for those interested in the application of nannofossils in high-resolution palaeoceanogmphy.
Karl-Heinz Bauman and Helge Meggers use nannofossil
assemblages to shed light on the palaeoceanogmphic evolution of the Labrador Sea during the Plio-Pleistocene.
These authors show conclusively that nannofossil assemblages respond dmmatically to water-temperature changes
during glacial and interglacial inteiVals and provide one of
the clearest demonstrations of the effect of temperature on
coccolith size. Hisa Okada and Mariko Matsuoka use theabundanceof nannofossils which inhabit the lower part ofthe photic zone to infer relative monsoonal wind-strength
in the tropical Indian Ocean during the Late Quaternary.
Using spectral analysis, they demonstrate that wind strength
fluctuates cyclically in tune with the eccentricity of the
Earth's orbit. An interesting twist to their research is the
surprising result that glacial inteiVals show less intensified
circulation than interglacials . Alex Chepstow-Lusty provides a useful summary of studies of Discoaster abundance,
which has been previously been used by himself and otherauthors as an indicator of warm and oligotrophic watermasses. He shows high-resolution Late Pliocene data setsfrom 10 sites that support other evidence for major highlatitude cooling during this time inteiVal with less dramatic
tempemture changes in lower latitude areas. Jim Pospichal
provides yet another comprehensive paper on the
nannofossil assemblage changes at the Cretaceous/Tertiary boundary. He shows conclusively that there was little
precursor environmental pressure on the nannoplankton
at a high-latitude site prior to the KIT event as has been
postulated for the planktonic fomminifem, but that the surface-water productivity decreased substantially across theboundary. The paper selVes as a excellent summary of Jim's
long-term work on nannofossil assemblage changes acrossthe KIT boundary in a host of different environments andocean basins.
Jeremy Young describes in detail the study of the
shape of Emiliania huxleyi using a light microscope-based
image analysis system. He elegantly describes how the technique works and demonstrates that the LM-based system
can provide high-precision measurements of a variety ofmorphocharacters. Jeremy convinced me that I have wasted
thousands of dollars on scanning electron microscopy!
Finally Dave W atkins and others provide a stateof-the-art compilation of high southern latitude Upper Cretaceous nannofossil biostratigmphy. They demonstrate that
it is possible to obtain relatively high stratigraphic resolution in this intetVal. The paper includes an updated list of
species considered to be high-latitude in affinity and ananalysis of how high southern latitude surface-water masses
evolved both spatially and temporally in the latter stagesof the Cretaceous.
In summary, the nannoplankton community did an
excellent job providing a showcase of innovative and provocative papers for the rest of the micropalaeontologicaland palaeoceanographic community.

========================~46==========================

J. Firth: ODP News, p. 47- 48.

Journal of Nannoplimkton Research, 18, 2, 1996.

OCEAN DRILLING PROGRAM UPDATEJohn Firth, ODP, Texas A &M University, College Station, TX 77840, USA

Leg 162 (July 9- Sept. 3, 1995) was the second leg oftheNorth Atlantic Arctic Gateways program (the first being
Leg 151), and studied the gateway development between
the Arctic Ocean, Norwegian-Greenland Sea, and Atlantic
Ocean from the Palaeogene to Quaternary, as well as thepalaeoceanographic and palaeoclimatic history of this region. Wuchang Wei was the nannofossil specialist for thisleg.
Leg 163 (Sept. 7- Oct. 28, 1995) drilled the SE Greenland
margin in order to study the seaward-dipping basalt reflectors that formed massive flows along this margin during theearly rifting of the Norwegian-Greenland Sea.
Leg 164 (Nov. 1- Dec. 19, 1995) drilled the Blake Ridge
and Carolina Rise in order to study gas-hydrate formation.
Hisatake Okada was the nannofossil specialist on this leg.
Leg 165 (Dec. 24, 1995- Feb. 18, 1996) was drilled in the
Caribbean Sea and the Cariaco Basin in order to study
Cretaceous, Palaeogene and Neogene ocean history, theK!f boundary event, and high-resolution Quaternary sedimentation in an anoxic basin. The K!f boundary was recovered in three holes, and a long history of Tertiary explosive volcanism in this region was documented by numerous volcanic ash layers. Tim Bralower and Koji Kameo
were the nannofossil specialists for this leg.
Leg 166 (Feb. 23- Aprilll, 1996) studied the sea-level andfluid-flow changes in the Bahamas carbonate platform, andthe history of changes in oceanic circulation and climate
from the mid-Cretaceous to Recent Toki Sato was thenannofossil specialist for this leg.
Leg 167 (April21- June 16, 1996) studied the variability instrength and heat/salt transport capacity of the California
Current due to climatic and tectonic changes in the PacificBasin, and its relationship to fluctuations in upwelling andprimary productivity, and CCD changes in the NE Pacific.
Eliana Fornaciari was the nannofossil specialist for this leg.
Leg 168 (June 21- Aug. 16, 1996) investigated the natureand consequences of hydrothermal circulation in oceanic
crust, specifically to obtain information on lateral gradients in fluid composition, formation pressures and temperatures, formation-scale permeability, and circulation vigour.
Xin Su was the nannofbssil specialist for this leg.
Leg 169(Aug. 22- Oct. 17, 1996) drilled the JuandeFucaGorda spreading system in order to ascertain the interrelationships between tectonic, igneous, and sedimentary processes in controlling fluid-flow, energy and mass flux, andformation ofhydrothermal deposits at sediment-dominatedrift environments.
Leg 170 (Oct. 22 -Dec. 17, 1996) will drill the Costa Rica

accretionary prism with the aim of studying the mass- andfluid-flow patterns through the prism in order to establish
the mechanical and chemical behaviour of accretion andunderplating, and tectonic erosion, and to determine how
deformation and dewatering are distributed throughout anaccretionary prism. Staffing is now confirmed for this leg.
Leg 171 (Dec. 30, 1996- Feb. 16, 1997) is a two-part leg.
The first part (Dec. 30- Jan. 11) will be dedicated to using a
Logging-while-Drilling tool to retrieve logs of the Barbados accretionary prism. The second part (Jan. 11 -Feb. 16)
will drill the Blake Nose to recover Palaeogene and Creta·ceous soft sediments with the APC/XCB, in order to study
the older ocean history of the North Atlantic. Stafijng iscurrently underway for this leg.
Leg 172(Feb. 21- April18, 1997)will core a depth-transect
down the Blake Bahama Outer Rise and the Bermuda Rise
which will allow study of high sedimentation-rate drifts.
This will elucidate the history of changes in bottom- andintermediate-waters on extremely high-resolution timescales for the Late Neogene and Quaternary. Staffl.ng is
now underway for this leg.
Leg 173 (April23- June 18, 1997) will drill the Iberianmargin to study the history of early ri:fting along this nonvolcanic margin, and the ocean-continent basement transition. Staffing is now underway for this leg.
Leg 174 (June 23 - Aug. 18, 1997) is another two-part leg.
The first part (June 23 - July 17) will core in shallow waters
on the New Jersey Shelf to complete a transect of holes
previously drilled on Leg 150. The second part (July 20Aug. 18) will place a CORK in DSDP Hole 395A for a longterm study on the nature of fluid-flow through the crust at
this site. An engineering experiment will also be done totest a hammer drill-in casing system on bare, hard-rock outcrops on the Mid-Atlantic Ridge. Staffing is now underway
for this leg.
Leg 175 (Aug. 23 - Oct. 18, 1997) will study thepalaeoceanographic history of the Benguela Current, andthe history of upwelling along the SW African margin.
Staffing is now underway for this leg.
Leg 176 (Oct. 23- Dec. 18, 1997) will re-enter Hole 735B todeepen the hole through the lower crusta! gabbros. Staffing is now underway for this leg.
Scientific Prospectuses for upcoming ODP legs, beginningwith Leg 171, and Preliminary Reports of past legs, beginning with Leg 159, are now available on the World Wide
Web at http://www-odp.tamtLedu
For more information on ODP and other JOIDES
activities, write to Joint Oceanographic Institutions, Inc.,
1755 Massachusetts Ave., NW, Suite 800, Washington,

47============================

Journal of Nannoplankton Research, 18, 2, 1996.

DC, 20036-2102, USA. Phone: 202-232 3900 ore-mail:joi@iris.edu to request copies of the JOIDES Journal.
To apply for participation as a shipboard scientist
on an ODP cruise, send a letter of request and a resume to
the Manager of Science Operations, Ocean Drilling Program, Texas A&M University Research Park, College Sta-J. Firth: ODP News, p. 47 - 48. S. van Heck: The ICBN, p. 48 - 50.

tion, TX 77845, USA You will receive an application form
to fill in and return to ODP.
To request samples from DSDP /ODP cores, send a
letter of request to Chris Mato, Assistant Curator, Ocean
Drilling Program, Texas A&M University Research Park,
College Station, TX 77845, USA You will receive an application form to fill in and return to ODP.

THE ICBN: THINGS YOU NEED TO KNOW - 14
Shirley E. van Heck, Sarawak Shell Berhad, Lutong, Sarawak, Malaysia

The reorganisation of the new ICBN has slightly confused
the sequence I was following, starting from Article 6 andfollowing up on all references. However, since we havebasically finished with the articles dealing with validity, we
can return to Article 6, which has not changed much. One
thing that always causes a tremendous amount of confusion is the difference between valid and legitimate, and thisdifference is addressed in Article 6. I shall repeat the ruleon validity:
6.2. Valid publication ofnames is publication in accordance with Art.32- 45 ...

(as usual, I omit items that are not very relevant tonannoplankton).
This rule is a bit misleading because references in
Art.32- 45lead to a great many other articles, which therefore also have an impact on validity. These have been discussed in numbers 7 to 12 of this series of contributions.
6.3. A legitimate name is one that is in accordance withthe rules.

That seems a simple statement, but the following
article confuses the issue:
6.4. An illegitimate name is one that is designated assuch in Art. lB. 3, 19.6, or 52 - 54 (see also Art. 21 Note 1
andArt.24 Note 2). A name which according to this Codewas illegitimate when published cannot become legitimate later unless it is conserved or sanctioned.

In the book, the reference is to Art.19. 5 instead of
19 .6, but this is clearly a printing error. The question arises:
how about a name that does not comply with a rule notmentioned in Art.6.2 or Art. 6.4? It is notlegitimate, according to Art.6.3, but not illegitimate either. Actually, all therules are referred to, one way or another, when following
the references in the cited articles, so it is usually not a real
problem. Another interesting feature is that an invalid namecan be validated, but an illegitimate name cannot be madelegitimate (except by conservation). Now, let's follow up
on the references.
Art.18.3 has been discussed in van Heck(l991), so I shall..merely quote it here:
18. 3. A name ofa family based on an illegitimate
generic name is illegitimate unless conserved...

Art.l9.6 is a new one, and basically says the same for aname of a subfamily:
19. 6. A name ofa subdivision ofa family based on
an illegitimate generic name that is not the base of aconservedfamily name is illegitimate.. .

Now to some more interesting rules. The article refers to
Art. 52 - 54, which are in Chapter V, but I shall discuss therest of Chapter V as well, otherwise the other rules in thatchapter might get left out altogether.
Chapter V: Rejection of names
ARTICLE 51
51.1. A legitimate name must not be rejected merely because it, or its epithet, is inappropriate or disagreeable,or because another is preferable or better known (butsee Art.56. I), or because it has lost its original meaning,or...

Time and again people are tempted to do just that.
Noellaerhabdaceae has been considered inappropriate,
since the genus Noellaerhabdus is not a typical representative of the family (it also is a junior synonym, but thatis another story). I know someone who used Helicosphaera
not because it has priority, but because it was shorter than
Helicopontosphaera (and how about Pseudotriquetrorhabdulus?). And of course, most people use Discoasterbecause it is better known/preferable/easier to use thanEudiscoaster.
ARTICLE 52
52.1. A name, unless conserved (Art.J4) or sanctioned(Art.J5), is illegitimate and is to be rejected if it wasnomenclatura/ly superfluous when published, i.e. if thetaxon to which it was applied, as circumscribed by its
author, definitely included the type (as qualified in
Art.52.2) of a name which ought to have been adopted,
or whose epithet ought to have been adopted. under therules (but see Art.52.3).

This is a lot of words to say that an objective juniorsynonym is illegitimate. It is further qualified in the next,
although it does not add much:
52.2. For the purpose ofArt. 52.], definite inclusion ofthe
type ofa name is effected by citation (a) of the ho/otype
under Art.9. I or the original type under Art.] 0 or all========================~48==========================

S. van Heck: The ICBN, p. 48- 50.

syntypes under Art. 9.4 or all elements eligible as types
under Art.10.2; or (b) of the previously designated typeunder Art.9.9 or 10.2; or (c) ofthe illustrations ofthese. It
is also effected (d) by citation of the name itself, unlessthe type is at the same time excluded either explicitly or
by implication.
Example: Schwartz (1894) proposes to introduce anofficial name for all coccoliths, and because he cannot distinguish different species 'since they are so minute' he proposes 'for all the forms hitherto described, recent and fossil, the one name Coccolithus oceanicus'. This makes it ajunior synonym of C. pelagicus and C. carteri and hence
illegitimate.
ARTICLE 53
53. 1. A name of a family, genus or species, unless conserved or sanctioned, is illegitimate if it is a later homonym, that is, ifit is spelled exactly like a name based on adifferent type that was previously and validly published
for a taxon ofthe same rank.
Note 1: Even if the earlier homonym is illegitimate, or is generally treated as a synonym on taxonomic
grounds, the later homonym must be rejected.
One of the earliest examples is the nameCoccosphaera Wallich 1877, which is a homonym of
Coccosphaera Perty 1852 (algae). Introducing a new combination might result in a homonym, which must be avoided
by introducing a nomen novum instead (see Art.58).
53.3. When two or more generic, specific, or infraspecificnames based on different types are so similar that they
are likely to be confused (because they are applied torelated taxa orfor any other reason) they are to be treatedas homonyms.
This rule is likely to cause problems, as it is very
obviously subjective. The examples given in the ICBN are
not all that illuminating either, because under the names
they quote as not likely to be confused are several that I
had to read twice to see the difference. This, however, may
depend on the degree in which the taxa are related. However, I hereby propose to apply it to the following species:
Discoaster bergenii Kniittel et al. 1989 as a homonym of
Discoaster berggrenii Bukry 1971. Both species are rather
similar, have similar ranges, and the names are similar
enough to cause confusion to the uninitiated. I readily
agree that D. bergenii is a useful species to have (stratigraphically), but I would suggest that the authors chose a
different name. I further propose that anybody who wants
to discuss homonymy under this rule do so in the JNR, so
a wide nannoplankton readership is made aware of it andcan react to it if people disagree.
53. 4. When it is doubtful whether names are sufficiently
alike to be confused, a request for a decision may besubmitted to the General Committee ( ..)which will refer
itfor examination to the committee or committees for theappropriate taxonomic group or groups. A recommendation may then be put forward to an International Botanical Congress, and, ifratified, will become a binding decision.

===========================49Journal of Nannoplankton Research, 18, 2, 1996.

This process, of course, takes several years, so it
may be more efficient to decide amongst ourselves.

53.5. The names oftwo subdivisions ofthe same genus, oroftwo infraspecific taxa within the same species, even if
they are ofdifforent rank, are treated as homonyms ifthey
have the same epithet and are not based on the same type.
Compare this one carefully with Art.53 .1, for thismight cause confusion. A species and a subspecies of adifferent species may have the same name (see the discussion in van Heck, 1980, p.33 as an example) but, for example, a subspecies and a variety within the same species may
not have the same name.
53.6. When two or more homonyms have equal priority,
the first of them that is adopted in an effectively published text (Art.29-31) by an author who simultaneously
rejects the other(s) is treated as having priority. Likewise, ifan author in an effectively published text substitutes other names for all but one of thesehomonyms, thehomonym for the taxon that is not renamed is treated ashaving priority.
I have no example for this happening in nannoplankton literature, but it is something to keep in mind forthe future.
ARTICLE 54
54. 1. Consideration ofhomonymy does not extend to thenames oftaxa not treated as plants, except as stated below:(a) Later homonyms of the names of taxa once
treated as plants are illegitimate, even though the taxa
have been reassigned to a different group oforganisms towhich this Code does not apply.
(b) A name originally published for a taxon otherthan a plant, even if validly published under Art.32- 45
ofthis Code, is illegitimate ifit becomes a homonym ofa
plant name when the taxon to which it applies is firsttreated as a plant (see also Art. 45.5).
Finding out about the first, under (a), may be difficult, but not more so than finding out if, for instance, ageneric name has already been used for another plant.
However, a botanical institute may have the required
records. More likely to apply though is (b), as many
coccoliths were originally published under the zoologicalcode. However, I cannot find an example where this rulewould apply. The articles referred to have already been
discussed in previous issues.
ARTICLE 55
55.1. A name of a species or subdivision of a genus,
autonyms excepted (Art.22.1), may be legitimate even if
its epithet was originally placed under an illegitimate
name.
The 'may be' in this rule makes it rather difficult tounderstand, and the examples quoted do not help. The restof the rules under Art. 55 are as obscure, or do not apply.
ARTICLE 56
56.1. Any name that would cause a disadvantageous
nomenc/atural change (Art. 14.1) may be proposed forrejection. A name thus rejected, or its basionym if it has

Journal of Nannoplankton Research, 18, 2, 1996.

one, is placed on a list ofnomina utique rejicienda (App./V).
Along with the listed names, all combinations based onthem are similarly rejected, and none is to be used.

I know of no nannoplankton names that have been
officially rejected, for a good reason: it is a long legal battle
to achieve rejection, and it is far easier to simply agree not
to use a name. However, a time may come when we need toclean up our lists of taxa, and in that case this is the procedure to follow. Art.14 .1 d~als with conserved names, whichequally does not apply. Only if you intend to offer names
for rejecting need you bother with Art.56.2.
ARTICLE 58
58.1. A name rejected or otherwise unavailable for use
under Art. 52- 54 or 56- 57 is replaced by the name thathas priority (Art.ll) in the rank concerned. Ifnone exists
in any rank a new name must be chosen: (a) the taxon
may be treated as new andanother name publishedfor it,or (b) if the illegitimate name is a later homonym, anavowed substitute (nomen novum) based on the same typeas the rejected name may be publishedfor it. Ifa name isavailable in another rank, one of the above alternatives
may be chosen, or (c) a new combination, based on thename in the other rank, may be published.

We have several examples in nannoplankton. The
first I found serves as well as any other: Crepidolithus
p/iensbachensis was introduced as a nomen novum to replace Crepidolithus oce/latus Crux 1984 non Crepidolithus
ocellatus (Bramlette & Sullivan 1961) Noel1965.

S. van Heck: The ICBN, p. 48 - 50.

58. 2. Similar action is to be taken iftransfor ofan epithet
ofa legitimate name would result in a combination thatcannot be validly published under Art.21.3, 22.4, 23.4 or
27, or in a later homonym.

The only two of interest to us here are the cases forArt.27 (a species name may not be the same as the genus
name to which it is assigned) and the homonym. An example for the first is Tribrachiatus orthostylus, for which thetype is Discoaster tribrachiatus. A well-known example
for the second case, where a nomen novum was introduced
to avoid homonymy, is Coccolithus he/is. This has been
discussed in detail in van Heck (1979, p.C-3).
This concludes the discussion of Chapter V. Also, thislengthy contribution covers the last of 'what you need toknow' about the ICBN. I would encourage anyone to readthrough the volume to discover what more there is, and to
see the rules in their proper sequence. If you have any
questions, or want to discuss (or argue against) any points
that I have made, please write to me or send an e-mail!
REFERENCES
Greuter, W. et al. 1994. International Code ofBotanical Nomenclature (Tokyo Code) . Koeltz Scientific Books: 389 pp.
Heck, S.E. van 1979. Taxonomieal notes. INA Newsletter, 1(2):C3-C4.
Heck, S.E. van 1980. Bibliography and taxa of calcareous
nannoplankton./NA Newsletter, 2(1): 5-34.
Heck, S.E. van 1991. The ICBN: things you need to know- 5.
INA Newsletter, 13(3): 101-102.

========================~50 ========================~

J. Henderiks, P. Ziveri: Treating strongly-lithified rocks ... , p. 51.

Journal of Nannoplanldon Research, 18, 2, 1996.

TREATING STRONGLY- LITHIFIED ROCKS WITH ULTRASONICWAVES
Jorijntje Henderiks & Patrizia Ziveri, Geomarine Center, Vrije Universiteit, De Boe/e/aan 1085,
1081 HV Amsterdam, The Netherlands

A detailed calcareous nannofossil biostratigraphy has been
established for the Cenomanian-Turonian of the Russian
Craton Platform and the Crimean Peninsula, using the Cretaceous nannofossil zonation of Sissingh (1977) and modifications by Perch-Nielsen (1985). Quantitative andsemiquantitative analyses of six sections created abiostratigraphical framework for palaeoenvironmental andpalaeoceanographical interpretations. Results will be discussed in a separate paper, but we would like to make ashort note on the methodology used during this study whichmight be useful to other nannofossil investigators who workwith strongly-indurated rocks.
Strongly-lithified limestones from the Crimean sections proved to be barren ofcoccoliths after normal smearslide preparation. Experiments were performed to obtainsufficient material from these limestones in a quick and easy
way, as opposed to the laborious crushing and centrifuging technique, as described by Taylor & Hamilton ( 1982),
or the crushing and decanting method described by
Monechi & Thierstein (1985). It was found that treatmentof rocks with convergent, high-frequency ultrasonic waves
does not deleterously affect the small nannofossil fraction,and that sufficient nannofossiliferous material can be recovered by this method, if subsequently concentrated bycentrifuging. So-called sonifiers are occasionally used incleaning nannofossil material, but the frequency andstrength of the ultrasonic waves is far less than used forthis study.
Rock samples were placed into plastic bottles whichwere then filled with distilled, buffered water. The bottles
were put under a Branson Sonifier 450. This machine, that
can also be described as a ultrasonic 'disruptor', was operated for four minutes with the maximum output control ( 10)======================~51

and a duty cycle of700/o. To check for the reliability of thismethod in keeping the small fraction intact, a comparison
was made: smear-slides of a well-preserved, nannofossilrich sample were prepared using both the 'normal' method(from a scratched rock-surface) and from an ultrasonicwaves preparation. No differences between the two preparations could be appreciated under the polarising lightmicroscope, and SEM images showed no differences inpreservation, thus giving support for the non-destructive
nature of this method. Under the SEM, samples treatedwith this method also showed intact coccospheres, giving
additional support for the non-destructive character of themethod with respect to the nannofossil fraction.
Acknowledgements
The enthusiastic support and suggestions made by Peter
Willekes during laboratory procedures are gratefully acknowledged.
REFERENCES
Monechi, S. & Thierstein, H.R. 1985. Late Cretaceous-Eocene
nannofossil and magnetostratigraphic correlations near
Gubbio, Italy. Marine Micropaleontology, 9: 419-440.
Perch-Nielsen, K. 1985. Mesozoic calcareous nannofossils. In :H .M. Bolli, J.B. Saunders & K. Perch-Nielsen (Eds).
Plankton Stratigraphy: 329-426. Cambridge University
Press.
Sissingh, W. 1977. Biostratigraphy of Cretaceous calcareous
nannoplankton. Geol. Mijnbouw, 56(1): 37-65.
Taylor, R.J. & Hamilton, G.B. 1982. Techniques. In: A.R. Lord
(Ed. ). A Stratigraphical Index ofCa/careous Nannofossils:
11-15. British Micropalaeontological Society Series. Ellis
Horwood Limited, Chichester.

Journal of Nannoplankton Research, 18, 2, 1996.

============================52

D. Rut/edge, P. Bown: New names for old...... , p. 53 - 59.

Journal of Nannoplankton Research, 18, 2, 1996.

NEW NAMES FOR OLD: TAXONOMIC CLARIFICATION OF
SOME EARLY CRETACEOUS NANNOFOSSIL MARKER- SPECIESDavid Rut/edge, Robertson Research International, L/andudno, Gwynedd LL30 JSA, UK &Paul Bown, Department of Geological Sciences, University College London, Gower Street, London WCJ E 6BT, UK

Abstract: Five new boreal Early Cretaceous calcareous nannofossil species are described and illustrated:
Clepsilithus maculosus, N annoconus inornatus, Nannoconus pseudoseptentrionalis, Rhagodiscus
gallagheri and Rucinolithus windleyae. All five species have previously been misnamed or included within
the concepts of other species. The more precise taxonomic concepts introduced here provide improved
constraints for important marker-species events. One new combination, Zeugrhabdotus scutula, isintroduced.

Introduction
A detailed study of calcareous nannofossils from the boreal
area has shown that a number ofbiostratigraphically important taxa require more-precise definition. This paper
describes five new species that previous workers havegrouped with widely used marker-species, resulting in extended ranges and reduced biostratigraphical potential.
Recognition of these new taxa enables clarification of anumber ofbiostratigraphical ambiguities.
These observations are based on the detailed
biostratigraphical study of nine sections of Ryazanian toAptian age: three outcrop sections (two in England and acomposite German section) and six cores (two from onshore England, one from the southern North Sea, one from
offshore mid-Norway, and two from the Barents Sea). Allof the outcrop material and some of the core material isaccurately dated by means of cephalopods. The comprehensive results of this study will be presented in Rutledge& Bown (in prep.).
Systematic Palaeontology
Unless otherwise stated, all type-material is deposited in
the Postgraduate Unit ofMicropalaeontology, University
College London.
Genus CLEPSILITHUS Crux, 1987
Type species: Clepsilithus polystreptus Crux, 1987
Clepsilithus macu/osus sp. nov.

Plate l,Figures4-7
1982
1987
1988
1989
1991
1991

Stradnerlithus comptus Black, 1971; Taylor, p.71, pl.4.6,
figs 13-14 (EM).
Stradnerlithus comptus Black, 1971; Jakubowsk.i, p.117,
pl.2, figs 23-24 (LM).
Thurmanolithion clatratum GrOn & Zweili, 1980;
Applegate & Bergen, p.336, pl.17, fig.2 (EM).
Stradnerlithus comptus Black, 1971; Crux, p.207, pl.8.13,
figs 26-28 (LM).
Stradnerlithus comptus Black, 1971; Mutterlose (notillustrated).
Stradnerlithus comptus Black, 1971; Bralower, p.430,
fig.6.41-42 (LM).

platform; in crossed-polars the bars appear as a cycle ofregularly-spaced bright spots.
Description: The slightly flaring outer rim-cycle consists
ofabout 25 dextrally-imbricate elements, and is only weakly
birefringent (grey). The arrangement of elements in the
inner rim-cycle is uncertain, but this cycle is highly
birefringent (white) in crossed-polars. Each of the 12-16
regularly-arranged, radial bars consists of two laterallyfused elements, and exhibits white birefringence; these bars
support a rather less-birefringent (grey-white), diamondshaped platform that lies along the long axis of the central
area. There is no distal process.
Dimensions: Length 4.0-5.1J.UO, width 3.0-3.6).llll.
Remarks: This Upper Hauterivian marker species has beenpreviously misnamed Stradnerlithus comptus (e.g.
Jakubowski, 1987; Crux, 1989). S. comptuswasoriginallydescribed from the Kimmeridgian by Black (1971, pl.31,
fig.1 0) and has a similar number of radially-arranged bars,
but these are more delicately constructed and support aslender longitudinal bar, not a diamond-shaped platform.
However, the most significant difference between the two
species is their rim structure: S. comptus has a narrow,monocyclic-appearing rim composed of non-imbricate elements. In the light-microscope, the two are easily distinguished, S. comptus being weakly birefringent and inconspicuous, while C. macu/osus has a highly birefringent inner rim-cycle and central-area bars.
C/epsilithus polystreptus differs from C. maculosus
in having a less-developed inner rim-cycle, and fewer central bars (that are differently constructed), but this small
form has not yet been observed in the light-microscope.
The only contemporary species with which C. maculosus
might be confused is the similarly proportioned
Cretarhabdus inaequalis, but the latter species has a lessregular central structure.
Derivation of name: Latin, macu/osus meaning spotted,
referring to the brightly-spotted appearance of the cycleof central-area bars in crossed-polars.

Diagnosis: Small (4.0-5.0j.Ull), elliptical murolith with a bicyclic rim, the outer cycle of which is dextrally imbricate,

Holotype: Pl.l, fig.4 (EM).

and a central area occupied by about 14 short, radiallyarranged bars which support a diamond-shaped central

Paratype: PI. I, fig.6 (LM).

========================~53==========================

Journal of Nannoplankton Research, 18, 2, 1996.

Type locality & level: BGS Borehole 81/43 (southern NorthSea), 32.80m (Upper Hauterivian).

D. Rut/edge, P. Bown: New names for old ...... , p. 53 -59.

Holotype: Pl.l, fig.8 (EM).
Paratype: Pl.l, fig.9 (EM); Pl.l, fig.l6 (LM).

Occurrence: Hauterivian of the North Sea area,
occurringconsistently within the uppermost Hauterivian(above the last occurrence of Tegulalithus septentrionalis). The last occurrence of this species (basal
variabilis Amm. Zone) provides an excellent approximationof the Hauterivian-Barremian boundary, as currently defined at Speeton.
Genus NANNOCONUS Kamptner, 1931
Type species: Nannoconus steinmannii Kamptner,
1931
Nannoconus inornatus sp. nov.

Plate 1,Figures 8-12, 14-18
1982
1987

Nannoconus abundans Stradner & Griln, 1973; Taylor
pl.4.6, fig.19 (EM).
Nannoconus abundans Stradner & Griin, 1973; Thoms~n,pl.6, figs 10-11 (EM).

Diagnosis: A short, biconcave, pillar-like nannoconid constructed from thin plates arranged in a very low-angled
spiral, with a narrow axial canal.
Description: The sides of this nannoconid are convex, andthe ends unflared, slightly concave and identical, thus
apical/basal ends cannot be distinguished. The height isgenerally less than the diameter, often considerably less.
The axial canal is very much narrower than the wall of thenannoconid. The outer surface of the wall is smooth, giving a circular profile in end view. The thin plates composing the wall are arranged in a very low-angled spiral.
Dimensions: Diameter 5.0-9.01!ffi. height2.5-6.0Jllll
Remarks: Several previous authors (e.g. Taylor, 1982;
Thomsen, 1987) have included Nannoconus inornatus
within their concept of Nannoconus abundans, and consequently recorded an anomalously early first occurrence
for this marker-species. N. inornatus is considered to be
the ancestor ofN. abundans, but it is important to differentiate between the two species clearly, in order to maintainthe biostratigraphical integrity ofN. abundans.
N. inornatus differs from N. abundans in lacking a?basal flange, from Nannoconus circularis, Nannoconus
globulus and Nannoconus inconspicuus in having a narrower central canal, and from Nannoconus ligius in its
non-petalloid profile.
Two varieties of this species were recorded: a larger,
highly birefringent form, and a smaller, shorter variety yielding only white birefringence in end view. N. inornatus
developed vertical ribs in the Early Barremian (latefissicostatum Amm. Zone), giving it a regularly scalloped
margin in end view. These ribbed forms probably gave rise
to Nannoconus pseudoseptentrionalis sp. nov. in the late
Early Barremian (elegans Amm. Zone).
Derivation of name: Latin, inornatus meaning unadorned,
referring to the simple, un:flanged form of this species.

Type locality & level: BGS Borehole 81/43 (southern NorthSea); 14. 70m (Lower Barremian).
Occurrence: Upper Hauterivian-Barremian of the North
Sea area; rare through the lower part of this range, but
becoming common/abundant in the Lower Barremian,justprior to the inception of N. abundans.
Nannoconus pseudoseptentrionalis sp. nov.

Plate 1, Figures20-22?1987 Nannoconus quadriangulus quadriangulus Deflandre &Deflandre-Rigaud, 1967; Thomsen, pl.6, figs 6, 7, 9 (EM).
1989 Tegulalithus septentrionalis (Stradner, 1963) Crux, 1986;
Crux, p1.8.9, figs 5, 6 (EM).

Diagnosis: A short, flangeless nannoconid with a very
narrow axial canal, and an irregular, ragged margin, as seen
in end/plan view.
Description: This speCies is so short that it has only been
seen in end/plan view. The thin, shallowly-spiralling plates
composing the test overlap irregularly to give a ragged
margin. This form lacks regularly-spaced vertical ribs, andthus its margin is not regularly scalloped. The axial canal ismuch narrower than the width of the wall. It is highly
birefringent in the light-microscope.
Dimensions: Diameter: 4.5-7.0Jllll
Remarks: N. pseudoseptentriona/is is differentiated fromother similar nannoconids by its irregular, ragged margin.
This renders it similar to Tegula/ithus septentrionalis incrossed-polars, and it has been identified as such in several
previous studies (e.g. Crux, 1989). These optically-similar
species are readily differentiated in phase-contrast (in which
T. septentrionalis displays regularly-spaced wall elements)
and in side view (T. septentriona/is is commonly encountered in highly-distinctive side view). In end view, N.
pseudoseptentriona/is lacks the weakly birefringent flanges
often (but not always) seen on T. septentriona/is (see Pl.l,
fig.23).
Derivation of name: Greek, pseudes meaning fulse, i.e. falseseptentrionalis.
Holotype: Pl.1, :fig.20 (LM).
Paratypes: Crux(l989), p.l99,pl.8.9,:fig.5 (EM); Crux(1989),
p.199,pl.8.9,:fig.6(EM)(OttoGott, Germany, Bed 109, 'Middle' Barrernian, Aulacoteuthis Belemnite Zone).
Type locality & level: Speeton, NE England, Speeton Clay
Formation,LowerCementBed49,LowerBarrernian,elegans
Amm.Zone.
Occurrence: 'Middle' Barremian of the North Sea area.
This species is common in the Lower Cement Beds of

54==========================

D. Rut/edge, P. Bown: New names for old...... , p. 53 - 59.

Speeton (elegans Amm. Zone) and at a corresponding level in Germany. It is associated with common/abundant
Zeugrhabdotus scutula nov. comb. at both localities.

Journal of Nannoplankton Research, 18, 2, 1996.

GenusRUCJNOLITHUSStover, 1966
Type species: Rucinolithus hayi Stover, 1966
Rucino/ithus wind/eyae sp. nov.

Genus RHAGODJSCUS Reinhardt, 1967
Type species: Discolithus asper Stradner, 1963,
designated by Reinhardt, 1967
Rhagodiscus gallagheri sp. nov.

Plate I, Figures l-3
1987

Rhagodiscus angustus (Stradner, 1963) Reinhardt, 1971 ;Thomsen, p.77, pl.l6, figs 6-8 (EM).

Diagnosis: Small (<5 .OIJ.Illlong), eliipticalRhagodiscus witha length/width ratio of <2.5, and straight or slightly convex
longer sides. The central area is filled with a proximallysituated granular plate and spanned by short, transverse
struts which support a relatively large, hollow spine-base.
Dimensions: Length 3.5-4.5j.Ull.
Remarks: Previously included within the species conceptof Rhagodiscus angustus by several authors (Thomsen,
1987; Mutterlose, 1991), Rhagodiscus gallagheri occurs
earlier than Rhagodiscus angustus and is distinguished byits smaller size and more-elliptical outline. Morphologies
transitional between these two species have been observed,
confirming the difficulty in using R. angustus as a zonal
marker. Note that Stradner's (1963) holotype drawing of
Rhagodiscus angustus is parallel-sided with dimensions
5.0J.lm x 2.0J.lm. The oldest forms with these dimensions
were recorded, in our study, from the Chale Clay, Isle ofWight, from the lower forbesi Amm. Zone (intra-Lower
Aptian), although R. gal/agheri is still the dominant form
at this level. This first occurrence for R. angustus is still
very much earlier than the basal Upper Aptian event recorded by previous authors, perhaps reflecting application of an even narrower species concept, i.e. very elongate forms with a length/width ratio of> 3.
Derivation of name: Named inhonourofDr. Liam Gallagher,

nannopalaeontologist.
Holotype: Pl.1, fig.1 (LM).
Paratype: Thomsen (1987), p.77, pl.l6, fig.6 (EM) (Well
E-1, Danish Sector of the North Sea, 8193', basal Aptian).
Reproduced here as PI. I, fig.3.
Type locality & level: Atherfield Point (Isle of Wight),
Atherfield Clay Formation (Sample AC 13), Lower Aptian,forbesi Amm. Zone.
Occurrence: Common throughout the Aptian of the North
Sea area; particularly abundant in nannofossiliferous sampies from the Atherfield Clay, Isle ofWight, southern England (fissicostatus-forbesi Amm. Zones).

===========================55Plate I, Figures 24-29? 1987 Lithastrinus sp. Bralower, p.298 (not illustrated).

Diagnosis: Nannolith consisting of two inclined cycles ofbroad, petal-like elements: a wider ?proximal cycle of aboutnine regularly-imbricate elements, and a ?distal cycle formed
from a lesser nUf!lber of irregularly-overlapping elements.
Description: Two inclined cycles ofbroad, petal-like elements join at the solid centre of the nannolith; there is nocentral-area diaphragm structure. The wider ?proximal cycle consists of about nine sinistrally-imbricate, broadlypetalloid elements. The ?distal cycle consists of a lesser
number of irregularly-arranged elements, at least one ofwhich projects distally, overlapping the other elements ofthis cycle. In proximal view in the light-microscope, thisform appears as a large (7. 0-11.0 J.Lm), highly- birefringent
rosette, with imbricating, petalloid elements; distal focusing reveals the irregular distal cycle. In side view, it exhibitsa typical polycyclolith construction (sensu Varol, 1992),
with two cycles of inclined elements. The fairly high andirregular form of this species ensure that it is often encountered in oblique view in the light-microscope, where it appears as an irregular, radiating birefringent body.
Dimensions: Diameter 7.8-11.0j.Lill.
Remarks: This species is only tentatively assigned to ·Rucinolithus, due to its bicyclic construction. It would
seem to be intermediate between previously described species of Rucinolithus (mostly older) and fully formed
polycyclolithaceans, in which the two cycles of thenannolith are equally developed.
Differentiation: R. windleyae differs fromLithastrinus (asemended by Varol, 1992) in lacking a small central diaphragm. It is similar and possibly related to Rucino/ithus
pinnatus Bergen, 1994, but this older species (middle
Tithonian to latest Berriasian) has only eight imbricate elements and lacks a well-developed, irregular ?distal cycle,
having instead, a small-element cycle surrounding a central perforation. R windleyae is differentiated from the lightmicroscopically similar Assipetra terebrodentarius (PI. I,
figs 30, 31) by its larger size, and petalloid, rather than globular, habit. It is impOrtant to make this distinction in order toretain the biostratigraphical integrity of the latter species,
which has a first occurrence in the uppermost Hauterivian.
Derivation ofname: Named in honour of Dr. Dawn E.
Windley, erstwhile nannopalaeontologist.
Holotype: Pl.1, fig.25 (EM).
Paratype: Pl.l, fig.27 (EM); Pl.1, fig.28 (LM).

Journal of Nannoplankton Research, 18, 2, 1996.

D. Rutledge, P. Bown: New names for old......, p. 53 - 59.

Type locality & level: BGS Borehole 81/43 (southern NorthSea), 4l.l6m (Upper Hauterivian).

Occurrence: Lower/Upper Hauterivian boundary to Lower
Barremian ofBGS Borehole 81/43 and the Speeton section;rare through most of this range, but occasionally common
in the Upper Hauterivian to basal Barremian. Bralower (1987)
recorded the first occurrence of 'Lithastrinus sp.' (which
included the later-appearingAssipetra terebrodentarius)
in the Upper Hauterivian of a number of tethyan sites.
Genus ZEUGRHABDOTUS Reinhardt, 1965
Type species: Zygolithus erectus Dejlandre in
Dejlandre & Fert, 1954, designated by Reinhardt,

1965
Zeugrhabdotus scutula (Bergen, 1994) comb. nov.
Basionym: Reinhardites scutula Bergen, 1994 (Journal of
Nannoplankton Research, 16(2): 59-69, p.64, pl.l, figs
24a-c (holotype ), 25a-b ).
1989
1994

Zeugrhabdotus sisyphus (Gartner, 1968) Crux, p.198,
pl.8.7, fig.l (EM); pl.8. 12, fig.30 (LM).
Reinhardites scutula Bergen, 1994, p.64, pl.l, figs 24a-c(holotype), 25a-b (EM and LM; same specimen).

Remarks: Previous authors have assigned this species to
Zeugrhabdotus sisyphus, but included several other species (e.g. Z. trivectis) in this category, giving a much longer,

composite range. We believe that this species is best classified within the genus Zeugrhabdotus although it may
well be the ancestral species to the Reinhardtites lineageof the Late Cretaceous, as suggested by Bergen ( 1994).
Acknowledgements
We would like to thank DENI for funding of this research
project, IKU and BGS for the provision of sample materialand stratigraphic information, and Tim Bralower, Jim Bergen,
Osman Varol and Peter Rawson for advice, discussion andhelp along the way.
RERERENCES
Black, M. 1971. Coccoliths of the Speeton Clay and Sutterby
Mar!. Proceedings of the Yorkshire Geological Society,38(3): 381-424.
Bergen, J.A. 1994 . Berriasian to Early Aptian calcareous
nannofossils from the Vocontian Trough (S.E. France)and deep sea drilling site 534: new nannofossil taxa and asummary oflow-latitude biostratigraphic events. JournalofNannoplankton Research, 16(2): 59-69.
Bralower, T.J. 1987. Valanginian toAptiancalcareousnannofossil
stratigraphy and correlation with the upper M-sequencemagnetic anomalies. Marine Micropaleontology, 11: 293310.
Crux, J.A. 1987. Six new species of calcareous nannofossils fromthe Lower Cretaceous strata of England and Germany.
INA Newsletter, 9: 30-35.

Crux, J .A. 1989. Biostratigraphy and palaeogeographical applications of Lower Cretaceous nannofossils from northwestern Europe: 143-211. In: J.A. Crux & S.E. van Heck
(Eds). Nannofossils and their Applications. British
Micropalaeontological Society Series, Ellis Horwood Ltd.,
Chichester.
Jakubowski, M. 1987. A proposed Lower Cretaceous calcareous nannofossil zonation scheme for the Moray Firtharea of the North Sea. Abhandlungen Geologischen
Bundesanstalt, 39: 99-119.
Mutterlose, J. 1991. Das Verteilungs- und Migrationsmuster deskalkigen Nannoplanktons in der borealen Unterkreide(Valangin-Apt) NW-Deutschlands. Palaeontographica,
B221: 27-152.
Rutledge, D. & Bown, P.R. (in prep.). Calcareous nannofossilsof the boreal Lower Cretaceous.
Stradner, H. 1963. New contributions to Mesozoic stratigraphy
by means of nannofossils. Proc. 6th World Petroleum
Congress Frankfurt, Sect.]: 167-182.
Taylor, R.J. 1982. Lower Cretaceous (Ryazanian to Albian)calcareous nannofossils: 40-80. In: A.R. Lord (Ed.). A
Stratigraphical Index of Calcareous Nannofossils.
British Micropalaeontological Society Series, Ellis
Horwood Ltd., Chichester.
Thomsen, E. 1987. Lower Cretaceous calcareous nannofossil
biostratigraphy in the Danish Central Trough. Danmarks
Geologiske Undersegelse, Series A, 20: 1-89.
Varol, 0. 1992. Taxonomic revision of the Polycyclolithaceae
and its contribution to Cretaceous biostratigraphy. Newsletters on Stratigraphy, 27(3): 93-127.

List of species cited in the text
Assipetra terebrodentarius (Applegate et al., 1987 in
Covington& Wise, 1987) Rutledge & Bergen in
Bergen, 1994
Clepsilithus maculosus sp. nov.
Clepsilithus polystreptus Crux, 1987
Cretarhabdus inaequalis Crux, 1987
Nannoconus abundans Stradner & Griin, 1973
Nannoconus circularis Deres and Acheriteguy, 1980
Nannoconus globulus Bronnimann, 1955
Nannoconus inconspicuus Deflandre in Deflandre &Deflandre-Rigaud, 1962
Nannoconus inomatus sp. nov.
Nannoconus pseudoseptentrionalis sp. nov.
Nannoconus ligius Applegate & Bergen, 1988
Rhagodiscus angustus (Stradner, 1963) Reinhardt, 1971
Rhagodiscus gallagheri sp. nov.
Rucinolithus windleyae sp. nov.
Stradnerlithus comptus Black, 1971
Tegulalithus septentrionalis (Stradner, 1963) Crux, 1986
Zeugrhabdotus scutula nov. comb. (= Zeugrhabdotus
sisyphus Burnett in Gale et al. in press)

56========================~

D. Rut/edge, P. Bown: New names for old...... , p. .53 - .59.

Journal of Nannoplankton Research, 18, 2, 1996.

57

Journal of Nannoplankton Research, 18, 2, 1996.

D. Rut/edge, P. Bown: New names for old......, p. 53 - 59.

PLATE 1
Light micrographs x2112 magnification; electron micrographs, magnification as stated. Negatives are stored in the Postgraduate Unit ofMicropalaeontology, University College London. Neg.- negative number; LM- light-micrograph; SEM- scanning electron micrograph; xp - crossed-polars; pc - phase-contrast.
Figs 1-3:
Figs 1, 2:
Fig.1:
Fig.2:
Fig.3:

Rhagodiscus gal/agheri sp. nov.
Atherfield Clay Formation (Isle of Wight), AC 13, Lower Aptian, forbesi Amm. Zone.
Holotype, LM-xp,- Neg.5051-9.
LM-xp, Neg.5051-8.
Paratype, from Thomsen, 1987, p.77, pl.16, fig.6; Well El, 8193', Lower Aptian.

Figs 4-7:

Clepsi/ithus maculosus sp. nov.
BGS Borehole 81/43 (southern North Sea), 32.80m, Upper Hauterivian.
Holotype, SEM, distal view, x9180, Neg.4061-11.
SEM, distal oblique view, x8500, SEM, Neg.4061-12, same specimen as Fig.4.
Paratype, LM-xp, Neg.4071-19.
LM-pc, Neg.4071-21 , same specimen as Fig.6.

Fig.4:
Fig.5:
Fig.6:
Fig.7:

Figs 8-12, 14-18: Nannoconus inornatus sp. nov.

Figs 8-12:
Fig.8:
Fig.9:
Fig.10:Fig.ll:Fig.12:
Fig.14:
Fig.15:
Fig.16:
Fig.17:
Fig.18:BGS Borehole 81/43 (southern North Sea), 14.70m, Lower Barremian.
Holotype, SEM, oblique view, x5000, Neg.4079-17.
Paratype, SEM, oblique view, x5250, Neg.4079-16.
SEM, oblique view, x4875, Neg.4079-24.
SEM, oblique view, x5000, Neg.4079-22.
SEM, end view ofFig.11, x5000, Neg.4079-21.
LM-xp, side and end views, 'Neg.4099-6, Speeton, Bed LBSC.ID, Lower Barremian, rarocinctum Amm. Zone.
LM-pc, Neg.4099-7, same' specimens as Fig.l4.
Paratype, LM-xp, Neg.4099-4, side view, Speeton, Bed LBSC.ID, Lower Barremian, rarocinctum Amm. Zone.
LM-xp, Neg.3545-28, end view, Speeton, Bed LB1F (10), Lower Barremian, fissicostatum Amm. Zone.
LM-pc, Neg.3723-11, end view, Speeton, Bed LB5C.ID, Lower Barremian, rarocinctum.Amm. Zone.

Figs 13, 19:

Nannoconus abundans Stradner & Giiln, 1973.
Speeton, Bed LB1F (13), Lower Barremian, fissicostatum Amm. Zone.
SEM, oblique view, x4000, Neg.4069-29.
LM-xp, side view, Neg.3545-4.

Fig.13:
Fig.19:Figs 20-22:
Fig.20:
Fig.21:
Fig.22:

Nannoconus pseudoseptentrionalis sp. nov.
Speeton, Lower Cement Bed 49, Lower Barremian, elegans Amm. Zone.
Holotype, LM-xp, end view, Neg.4094-20.
LM-pc, Neg.4094-21, same specimen as Fig.20.
LM-xp, end view, Neg.4065-32.

Fig.23:Tegulalithus septentrionalis (Stradner, 1963) Crux, 1986.
LM-xp, Neg.4023-27, side and end views, BGS Borehole 81/43 (southern North Sea), 41.16m, Upper Hauterivian.

Figs 24-29:
Fig.24:
Fig.25:Fig.26:
Fig.27:
Fig.l8;
Fig.29:Rucinolithus windleyae sp. nov.
BGS Borehole 81143 (southern North Sea), 41.16m, Upper Hauterivian.
SEM, oblique view, x3875, Neg.4063-28.
Holotype, SEM, same specimen as Fig.24, x4000, Neg.4063-29.
SEM, ?distal view, x4000, Neg.4063-31 .
Paratype, SEM, side view, x4500, Neg.4060-30.
Paratype, LM-xp, Neg.4023-32.
LM-xp, Neg.4023-24.

Figs 30,31:
Fig.30:
Fig.31:

Assipetra terebrodentarius (Applegate et al., 1987 in Covington & Wise, 1987) Rutledge & Bergen in Bergen, 1994
LM-xp, Neg.4071-17, BGS Borehole 81/43 (southern North Sea), 11.92m, Lower Barrremian.
LM-pc, Neg.4071-18, same specimen as Fig.30.

Figs 32,33:
Fig.32:
Fig.33:

Assipetra infracretacea (Thierstein, 1973) Roth, 1973.
LM-xp, Neg.4071-2, BGS Borehole 81/43 (southern North Sea), 62.38m, Lower Hauterivian.
LM-pc, Neg.4071-3, same specimen as Fig.32.

========================~58

D. Rutledge, P. Bown:ew names for old. ..... , p. 53 - 59.

Journal of Nannoplanklon Research, 18, 2, 1996.

PlATE 159======================~

Journal of Nannoplankton Research, 18, 2, 1996.

========================~60

D. Frydas: Upper Zanclean silicoflagellates ... , p. 61 - 67.

Journal of Nannoplankton Research, 18, 2, 1996.

UPPER ZANCLEAN SILICOFLAGELLATES FROMMILOS ISLAND (CYCLADES, GREECE)
Dimitris Frydas, Department ofGeology, University ofPatras, GR-2611 0 Patras!Rion, Greece

Abstract: A calcareous and siliceous assemblage of phytoplankton from diatomaceous, laminated martsof the Frago section, at the southern part of Milos Island, was studied. It contains principally calcareous
nannofossils and silicoflagellates. The calcareous nannofossil association belongs to subzone CN11b(Discoaster assymetricus) from the Upper Zanclean (sensu Okada & Bukry, 1980), whereas thesilicoflagellates correspond, more or less, to the upper part of the Dictyocha extensa Interval Zone (sensu
McCartney et al., 1995). Dictyocha arbutusensis (Bukry, 1982) cycladica n. subsp. and Dictyocha(?) sp.
were found and are described from the silicoflagellates of the Frago section. The percentage variation of
the silicoflagellate species reflects the deposition of the sequence in subtropical to tropical conditions.

Introduction
The island of Milos belongs to the southern part of theactive Cyclades volcanic arc (Figure lA), which extends
from Soussaki (adjacent to the Isthmus of Corinth) up to
Nissyros and Giali, through Aegina, Methana, Poros, Milosand Santorini islands (Fytikas et al., 1976). Numerous scientists have studied the geology of Milos since the lastcentury. Sauvage (1846) compiled a geological map of theisland. Philippson (1897) published the first synthetic workon the geology of the Cyclades. The significant work of
Sonder (1925) relates to the volcanic rocks. Liatsikas (1949)
studied the important minerals and rocks of the island.
Angelier et al. (1977), Fytikas (1977a), Jacobshagen (1986)
and Pe-Piper & Piper (1989) studied the geology ofMilos
with respect to the evolution of the Hellenides. A detailed
geological map of the island (at 1:25 000 scale) was compiled by Fytikas (1977b). Neogene carbonate deposits
transgressively overlie the Upper Cretaceous-Palaeogene
metamorphic basement. Volcanic deposits are the dominant lithology of the island. This volcanic activity has leadto the development of hydrothermal fields and fumerolesat Chora and Adamas (Fytikas, 1977a), related to the increased geothermal gradients. Frydas (1994a) studied the'?-'stratigraphy and palaeoecology of Lower Pleistocene
silicoflagellate and diatom assemblages near Adamas. Bellas& Frydas (1994), in a preliminary report, described the
Neogene deposits of the Tsouvala and Frago sections insouthern Milos. They assigned to the calcareous
nannofossil assemblage of the former section a Messinianage.
The subject of this paper is a stratigraphical studyofNeogene deposits from the Frago section, using calcareous nannofossils in conjuction with silicoflagellates.
Lithostratigraphy
The sequence outcrops along the southern coast ofMilos
Island.It comprises approximately 8m of sediment above
present sea-level and is located on the Frago Peninsula,
between the cape Mesa Akrotiraki and the Pounta Peninsula (Figure IB). Three coarsening-upwards sedimentation
cycles can be distinguished in the section (Figure 2). The
beds of the section dip S-SE at an angle of 5-15°. The basal

2r

A

~
AEGEAN p

Chlos

Halepa

S E A
38'

294.4

...

38'

Andros

272.1

...

~~

, l?

~
/)
,<;;:>CYCLADES

IJ

37'

Milos
0

50

100lan22'~.f(7 ().Naxos

37'c'"'-?

~· " ' ' tJ

'l>

t)

Santorini

..

0

SOOm

1:::=:--olv·Fig. 1A: Map showing the island of Milos among the Cycladesgroup in the southern part of the volcanic arc of the Aegean Sea.

Fig. 1B: Location of the Frago section on the island of Milos,
Greece.

========================~61========================~

Journal of Nannoplankton Research, 18, 2, 1996.

D. Frydas: Upper Zanclean silicoflagellates... ; p. 61 - 67.

FRAGO~~~~ Lahar~~~~laminated dlatomaceous marls~~ ~~ Diatomaceous burrowed marls
I: ":: : : : :1Sandstoner"\JV\.1\ Unconformity...:_·_:._· _sea level

Fig. 2: Lithostratigraphy of the Frago section.

part consists of a sandstone bank which probably represents the uppermost part of an incomplete sedimentation
cycle. This is overlain by light green to yellowish, mediumcemented, laminated marls which pass upwards throughyellowish, sandy marls with burrows and iron oxides, tomedium-grained, light green, slightly cemented sandstone.
White in colour, stratigraphically important diatomaceous
layers are also interbedded in the second complete sedimentation cycle. Moderately laminated, white to yellowish
diatomaceous marls overlay the complete cycle. The upwardly-increasing percentage of sand, heavily burrowed
traces, and an internally, almost-destroyed structure dueto the bioturbation, are indications of water shallowing
during the deposition of these diatomites. The sandstone,

11
3
14

9

4

1
5

9

8

2

5

7

6

7

')

7

10

12

7

8
6

9

17

4
6

3
4

.1

h

6

7

9
4
10

10

Calcareous nannofossils:1Ml.fRRAY & RI.ArKMAN£akidiSL'US
C. macintvredBU_KRY_&_RRAMl .F.TTF.)
Coccolithus velaf!icus (WALL! CH)_
Discoaster asvmmetricus GARTNER

6

6

hrouweri

l

6

5
8

()

4

7

2

~LN..l:i lK

D. variabilis_ MARTINI & BRAMLETTE
Heticosvhaera carterUWALLlCH~

5'iilinnin

3

~

8

12

10

12

14

12

4
13

'I

2
5

8

34

4

4

8

'I

6

11

14

I K AJ\l_)i/;,.· l!DARTNF.R)

Scvvho.'mhaeraso.

2

4
5

48

10

23

22

4
19

48

9 D. bmim.t~illzLnq_(L_EMMERMANNl
46 D. extensa extenso (lOCKER l

'i

4
6

lh

20

39

R

n

5

8
23
22

Silicoflagellatessnnrh· .,,,.,,ra·i< IRI JKRY) ~vcladic, n subsn.

')j, •flu I. •hn 1'1

In7

AN ~IN HliKD.
·TAN SINHOK.D surculus MARTlliJ~ BRAMLETTE
D. tamalis KAMPTNER

3

4

1

4

Nannofossils
are indicatedas_p_ercents (%)

Sample number
Age

1

h

Biostratigraphy
The studies ofDeflandre ( 1950), Stradner (1961 ), Stradner&Bachmann(l978), Bukry(l981, 1982),Bukry &Foster(1973),Dmnitrica(1973),Perch-Nielsen(l985a,b)andF:tydas(1990, 1991, 1993) were used for the identification of thevarious calcareous and siliceous phytoplankton species.
The most characteristic species are shown in Plate 1. Table
1 shows the stratigraphical distribution of the calcareousand siliceous nannofloras along the studied section.
Calcareous nannofossils, as well as somesilicoflagellate species (samples Ml to M6), were recovered from the laminated, marly diatomites of the lower pintof the Frago section. According to the common presenceof Discoaster asymmi!lricus, Discoaster tamalis andReticu/ofenestra pseudoumbilica, the rich calcareous
nannoplankton association (Table 1) is assigned to NN1415B (Lower Pliocene) sensu Driever (1988), which corresponds to CN11b ofOkada & Bukty (1980) and to the upperpartofNN15 (Martini, 1971).
The diatomites of the upper part of the Frago section (samples B1-B6) yielded silicoflagellates and diatomsof higher abundance compared to the underlying, lower
part, while the occurrence of calcareous nannoplankton
decreased towards the top. In this upper part, Reticulofenestra pseudoumbi/ica is absent, whereas Discoasterl Table I J

Frago-section, Milos Island
MB
4
5
1 2 3 43
6
5
Uppennost Zanclean
Discoaster asymmetricus Subzone

2

I

which probably covered them as part of athird sedimentation cycle, has been removed by erosional processes which
followed another regressional event.
Finally, at the top of the section, an approximatelyl.Sm thick, green-coloured volcanic deposit ('lahar' after
Fytikas, 1977b) unconformably overlies the older sequence.
Recent studies consider lahar to be the result of one ormore discrete processes, rather than a single depositional
product (Fisher & Smith. 1991).

rnt>oonm>n •· • n<•ninll<n

7

2

IK~Y

Xr M

,R

11\."

D. cf verlaevisve~FNGlJFI.JJ)/)i.
. · holiviensis boliviensis(FRENGVELLil
Ds. ~culum minutw.J.BACHMANN.l' " " ' ' " • JIYI IEHK.tNJj.tKLr
13. .25.. 28. 'Js
20 10 17 Mesocena circulus LEHKJ::ilitlER.Ul

100 lOO lOO lOO 100 100 100 100 100 100 lOO 1.00===========================62==========================

D. Frydas: Upper Zanclean silicoflagellates ... , p. 61 - 67.

tamalis is scarcely present. In our material, the silico-

flagellate association corresponds to the upper part of the
Dictyocha fibula Zone (Bukry, 1981) and, more or less, tothe upper part of the Dictyocha extensa Interval Zone of
the eastern equatorial Pacific (McCartney et al., 1995).
The above-mentioned Dichtyocha extenso Interval Zone is not congruent to the Mesocena circulus Zone
ofMartini & Miiller ( 1976, DSDP Leg 38) and Perch-Nielsen(1985b), which really represents the Paramesocena
apiculata!Paramesocena circulus apiculata Zone. Locker& Martini (1986, DSDP Leg 90) have introduced aParamesocena circulus Zone for Lower to Upper Pliocene
sediments of the southwestern Pacific Ocean. This zone
lies above the uppermost Miocene-Lower Pliocene
Neonaviculopsis neonautica neonautica Zone (Locker&Martini, 1986).
Common to abundant silicoflagellates are:Dictyocha extensa extensa, Distephanus speculum
speculum and Mesocena circulus, the latter constituting

more than 20% in the studied samples. In smaller amounts
are also to be found: Dictyocha (?) sp., Dictyocha
arbutusensis cyc/adica n. subsp., Dictyocha brevispina
brevispina, Dictyocha cf. D. perlaevis perlaevis andDichtyocha messanensis aspinosa, while Distephanus
boliviensis boliviensis and Distephanus speculum minutus
are rare. Most of the silicoflagellate species encountered in
the present study have been reviewed in detail (Frydas,
1990, 1991, 1993). Silicoflagellate taxonomy is fully given inthese works. Therefore, full synonymies are not presented
here. One new subspecies of the genus Dictyocha, as well
as a Dictyocha(?) species of late Early Pliocene samples
from the Frago section, were determined and are described
here.

Journal of Nannoplankton Research, 18, 2, 1996.

has an elongated to elliptical ring lacking minor-axis spines,
and a very short apical bar. Two short, radial spines arealigned atthe major axis of the elongated basal ring. The
basal-ring bars show a distinctly rounded to half-circularform in the part on the ring where the lateral bars instate.
The maximum from inner diameter ranges from 39 to 52)llll(holotype = 49.3~-tm). The length/width (LIW) ratio frominner diameter ranges from 1. 7 to 2. 5.
Discussion: Dictyocha arbutusensis cyc/adica n.subsp.
is distinguished fromDictyocha arbutusensis (Bukry, 1982)
by the distinctly more semi-circular, rounded basal bars in
the part of the ring where these are joined with the obliquelateral bars; by the obliquely-oriented apical bars; by thevery short, radial spines; and by its generally smaller sizeand less broad outline. Dictyocha arbutusensis (Bukry,
1982) has a maximum inner diameter from 45 to 651-Ull, and a
L/W ratio from 2.8 to 3.4.
Age: Dictyocha arbutusensis cycladica n.subsp. first appears in the Frago section at the stratigraphical level ofSubzone CNllb (Discoaster asymmetricus) in the Lower
Pliocene (Zanclean). Dictyocha arbutusensis (Bukry, 1982)
occurs in low frequencies in the Upper Pliocene in the eastem equatorial Pacific. Its stratigraphical range is not yet
exactly established.
Occurrence: Dictyocha arbutusensis cyc/adica n.subsp.
is a common species in Zanclean samples M5 and M6 (290340cm above sea-level) in the Frago section. Other common species in these samples are: Dictyocha extensa
extensa, Dictyocha messanensis aspinosa, Distephanus
speculum minutus, Distephanus speculum speculum and
Mesocena circulus.

Systematic descriptions
Order Silicojlagellata Borgert, 1891
Family Dictyochaceae Lemmermann, 1901
GenusDICTYOCHA Ehrenberg, 1840
Dictyocha arbutusensis (Bukry, 1982) cyc/adica

n. subsp.
Plate 1, Figures 1, 2
Derivation of name: 'Cyclades' =group of islands in the
Aegean Sea to which the island ofMilos belongs.

Dictyocha(?) sp.
Plate 1, Figures 3, 4

Holotype: Plate 1, Fig.3; sampleBl.
Paratype: Plate 1, Fig.4; sample B4, from type locality.
Type locality: Frago section, samples B 1 and B4 (470cm
and 570cm above sea-level, respectively).
Type horizon: Lower Pliocene (Zanclean), Sub:zone CN11b(Discoasterasymmetricus) (orupperNN15).

Holotype: Plate 1, Fig.l; sampleM5.
Paratype: Plate 1; Fig.2, sampleM6, from type locality.
Type locality: Frago section, samples M5, M6 (290-340cm
above sea-level), Milos Island, central Aegean Sea.
Type horizon: Lower Pliocene (Zanclean), Sub:zone CN1lb(Discoaster asymmetricus) (or upper NN15).

Diagnosis: Elongated basal ring with two short radial spines
and a very short apical bar.
Description: Dictyocha arbutusensis cycladica n.subsp.

Diagnosis: Subparallel basal ring with two short spines
and a convex apical bar.
Description: Dictyocha(?) sp. has a subparallel to slightly
oblong basal ring with only two short spines aligned at the
major axis of the elongated ring. A slightly convex apicalbar connects the subparallel sides of the ring along theminor axis. The maximum from inner diameter ranges from42 to 62~-tm (holotype = 51.3~-tm). The L/W (length/width)
ratio from the inner diameter ranges from 2.1 to 2.6.
Discussion: Dictyocha(?) sp. is distinguished from
Dictyocha neonautica (Bukry, 1981), from the Upper========================~63==========================

Journal of Nannoplankton Research, 18, 2, 1996.

Miocene of the Carnegie Ridge, Pacific Ocean, DSDP 15738-CC (343m) and the Panama Basin, by its essentiallysmaller size. Dictyocha neonautica (Bulay, 1981) hasamaximwn inner diameter of70-901lffi
Age: Dictyocha neonautica defines the homonymous
subzone (Bukry, 1981) which is equivalent to the lower
part of the silicoflagellate Dictyochafibula Zone (Upper
Miocene). On the island of Crete, Dictyocha cf. D.
neonautica (sensu Bukry, 1981)has been estabiished as
the marker for an acme-subzone in the Lower Piacentian in
the Marathiti; Aghios Vlassios (Frydas, 1990, 1996) and
Gournes (Frydas, 1994b) sections of the Heraklion Province, and also in the Stavromenos section (Frydas & Keupp,
1992) of the Rethymnon Province.
Occurrence: Dictyocha(?) sp. is a common species in Upper Zanclean samples B I and B4 in the Frago section. Its
stratigraphical range is not yet exactly known. Other common species in these samples are: Dictyocha extensa
extensa, Dictyocha brevispina brevispina, Dictyocha cf.
D. perlaevis perlaevis, Dictyocha messanensis aspinosa,
Distephanus boliviensis boliviensis, Distephanus
speculum minutus, Distephanus speculum speculum and
Mesocena circulus.
Acknowledgements
I wish to thank the German Academic Exchange Service(DAAD), Bonn, Germany, for supporting my two-monthresearch (summer, 1995) at the Institute of Palaeontologyof the Free University ofBerlin. Thanks are due to Prof. Dr.
H. Keupp (Free University ofBerlin) and two reviewers forconstructive comments and critical reading of the manuscript: Assoc. Prof. Dr. K. McCartney, University ofMaine
atPresque Isle, USA, and Dr. Jackie A. Burnett, University
College London. English language problems were kindly
corrected by Assoc. Prof. Dr. Mrs. K. Kotopouli (Instituteof Petrography, University ofPatras).
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de Microscopic theorique et appliquee, 2 (1950): 72 etsuiv.).
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=========================65

Journal of Nannoplankton Research, 18, 2, 1996.

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D. Frydas: Upper Zanclean silicoflagellates... , p. 61 - 67.

Journal of Nannoplankton Research, 18, 2, 1996.

PLATE 1

Figs 1-17: Silicoflagellates. Figs 1-6, 12, 14 and 15-17 magnified x850 approximately; Figs 7-11 and 13 magnified x950
approximately. Figs 18-24: Calcareous nannofossils. All specimens magnified 1400x approximately. Holotypes are stored
in the Dept. of Geology, University ofPatras.
Figs 1, 2:
Fig.1:
Fig.2:

Dictyocha arbutusensis (Bukry) cycladica n.subsp ..

Figs 3, 4:
Fig.J:
Fig.4:

Dictyocho(?) sp..

Figs 5, 6:
Fig.5:
Fig.6:

Dictyocho brevispino brevispino (Lemmermann).

Figs 7-9:
Figs 7, 9:
Fig.8:

Dictyocho extenso extenso (Locker) Locker & Martini.

Holotype, sample M5.
Paratype, sample M6.

Holotype, sample Bl.
Paratype, sampleB4.

SampleM6.
SampleB4.

SampleB4.
SampleM6.

Figs 10, 11: Dictyocho messonensisaspinosa (Bukry) Locker & Martini.
Fig.10:SampleM6.
Fig.ll:SampleB4.
Fig.12:Distephonus boliviensis boliviensis (Frenguelli). Sample B4.

Fig.13:Distephonus speculum mlnutus (Bachmann). Sample M6.

Fig.14:Distephonus speculum speculum (Ehrenberg). Sample B4.

Figs 15-17: Mesoceno circulus (Ehrenberg) [syn: Paradictyocho circulus (Ehrenberg) Dumitrica].
SampleB4.
Fig.15:
SampleM5.
Fig.16:
SampleM6.
Fig.17:
Figs 18-19: Discoaster asymmetricus Gartner.
Fig.18:SampleMl.
Fig.19:SampleM6.
Fig.20:Discoaster brouweri Tan Sin Hok. Sample M6.

Fig.21:

Discoaster surcu/us Martini & Bramlette. Sample M6.

Fig.22:

Discooster variabilis Martini & Bramlette. Sample M6.

Fig.23:

Discoaster tama/is Kamptner. Sample M6.

Fig.24:

Discoaster triradiatus Tan SinHok. SampleM6.

=========================66=========================

D. Frydas: Upper Zanclean silicoflagellates ... , p. 61 - 67.

Journal of Nannoplankton Research, 18, 2, 1996.

PLATE 1

. . . . . '"'1 ,,.

·-~-V.:·,'·.···... ;,·;..,,,~,(.::
·•

-;~~

..I. . _'-.... ':".

«;..

,

,

• w.

.,~.=~~-:-:;,~·_ . . -~~-fj. ·-~ -~---. '1 .
. '~.:..~. '(' t., . , ·~..
..,

..

! •

()

;

'

' t...:.::- .
• .....~~

1)·;-- . ---.. '--:--.. ·__\ ,: t... ........
,~ .... . ,. ""
·

I

{'

•

=========================67========================~

Journal of Nannoplankton Research, 18, 2, 1996.

============================68

W G. Siesser: Calcareous nannoplankton bibliography 27, p. 69 - 83.

Journal of Nannoplankton Research,l8, 2.

BIBLIOGRAPHY AND TAXA OF CALCAREOUS NANNOPLANKTON - 27
Compiled by Wil/iam G. Siesser

A5227

AGUADO, R., MOLINA, J.M., & O'DOGHERTY, L.
Biostratigraphy and lithostratigraphy of the Carbonero Formation (upper Barremian-lower
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A5228

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AKSU,A.E, YASAR, D., MUDIE, P.J.,& GILLESPIE, H.
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A5230

ALCOBER,J. & JORDAN, R. W.
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1996

Abst.
Syst.

A5231

AMORE,F.O.
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QUAT.;TERT.U.
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A5232

ANDRULEIT, R
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A5233

BARKER, G.L.A., GREEN,J.C., HAYES, P.K,& MEDLIN, L.K
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1994

Bioi.
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A5234

BEAUFORT, L. & OLAFSSON, G.
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A5235

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1994

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1994

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A5238

BLEIJSWIJK, J.D.L. van, KEMPERS, R.S., WAL, P. van der,WESTBROEK,J.K, EGGE,J.K, & LUKK, T.
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1994

Bioi.
Living==========================69==========================

Journal of Nannoplankton Research,l8, 2.

W. G. Siesser: Calcareous nannoplankton bibliography 27, p. 69 - 83 .

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DOWN, P.R. & ELLISON, C.
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1996

Abst.
Strat.
CRET.L.;
JURA.U.
AmericaS.

A5240

BOWN, P.R., YOUNG, J.R., EHRENDORFER, T.E., DAVIES, S.A.,DIDYMUS, J.M., & MANN, S.
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1996

Abst.
Bioi.

A5241

BRALOWER. T.J.,ZACHOS,J.C., THOMAS,E.,PARROW,M.,PAULL,C.K.,
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Late Paleocene to Eocene paleoceanography of the equatorial Pacific Ocean: stable isotopes
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1995

Strat.;
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A5242

BROWNLEE, C., NIMER, N., DONG, LF., & MERRETT, M.J.
Cellular regulation during calcification in Emiliania huxleyi.
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1994

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A5243

BURNETT,J.
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1996

Abst.
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Worldwide

AS244

BURNETT,J.A.
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Worldwide

A5245

BUSSON, G., NOitL, D., MONNIOT, F., & CORNEE, A.
Some calcareous nannofossils misinterpreted in literature as Teualithus, Triquerrorhabdulus,
Fusellinus sp. are actually spicules of Didemnum ssp. (Tunicata, Ascidian).
Evidence from Upper Jurassic sediments of Charentes (W. France).
+ 6th INA Conf., Copenhagen, 1995, abstracts. J. Nannoplankton Res., 17(2): 49-50.

1996

Abst.
Syst.
JURA.U.
Europe W.

A5246

BYBELL, L.M. & GIBSON, T.G.
Paleogene stratigraphy of the Putneys Mill, New Kent County, Virginia, corehole.
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A5247

BYBELL, L.M., SELF-TRAIL,J.M., & GIBSON, T.G.
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A5248

CACIIA.O, M. & MOITA, M.T.
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1996

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CAVALIER-SMITH, T.
Origin and relationships of Haptophyta.
+ In: Green, J.C. & Leadbeater, B.S.C. (Eds.), The Haptophyte Algae. Systematics Assn.
Spec. Vol. No. 51, Clarendon Press, Oxford, pp. 413-435.

1994

Evol.
Syst.

CHEPSTOW-LUSTY A. & CHAPMAN, M.

1996

Abst.
Strat;Ecol.
TERT.U.
Worldwide

1994

Bioi.
Living

A5250

An examination of the global relationship between the last two "species" of Discoasters

at the end ofthe Pliocene.
+ 6th INA Conf., Copenhagen, 1995, abstracts. J. Nannoplankton Res., 17(2): 51-52.
A5251

CONTE, M. H., THOMPSON, A., & EGLINTON, G. .
Primary production of lipid biomarker compounds by Emiliania hux/eyi. Results froman experimental mesocosm study in fjords of southwestern Norway.
+ Sarsia, 79: 319-331.

===========================70===========================

W. G. Siesser: Calcareous nannoplankton bibliography 27, p. 69 - 83 .

Journal of Nannoplankton Research,l8, 2.

A5252

CONTE, M. H., VOLKMAN, J.K, & EGLINTON, G.
Lipid biomarkers of the Haptophyta.
• In: Green, J.C. & Lead beater, B.S.C. (Eds.), The Haptophyte Algae. Systematics Assn.
Spec. Vol. No. 51, Clarendon Press, Oxford, pp. 351-377.

1994

Bioi.

A5253

CROS, L.
Coccolithophorid distribution across a transect between the Iberian Peninsula and the Balearic
Islands (northwestern Mediterranean Sea).
t 6th INA Conf., Copenhagen, 1995, abstracts. J. Nannoplankton Res., 17(2): 53.

1996

Abst.
Ecol.
LivingMediterr.

A5254

CURRY, W.B., SHACKLETON, N.J., RICHTER. C., & THE
SHIPBOARD SCIENTIFIC PARTY.
t Proc. ODP, !nit. Repts., v. 154 (Ceara Rise; Sites 925-929). 1111 pp.
[Nannofossils by J.E. Backman & I. Raffi]

1995

Strat.
QUAT .;TERT .L.-U.
Atlantic C.

A5255

D' AT RI, A., NOVARETTI,A., RUFFINI, R., COSCA, M.A. & HUNZIKER J.C.
Calcareous plankton biostratigraphy and 40Arf3 9Ar Dating of volcanic ash layers
from Monferrato (NW Italy).
t 6th INA Conf., Copenhagen, 1995, abstracts. J. Nannoplankton Res., 17(2): 53.

1996

Abst.
(Strat.)TERT .U.
Europe W

A5256

DE VRIND-DE JONG, E.W., VAN EMBURG, P.R., & DE VRIND, J.P.M.
Mechanisms of calcification: Emiliania huxleyi as a model system.
t In: Green, J.C. & Leadbeater, B.S.C. (Eds.), The Haptophyte Algae. Systematics Assn.
Spec. Vol. No. 51, Clarendon Press, Oxford, pp. 149-166.

1994

Bioi.

A5257

DIDYMUS, J.M., YOUNG, J.R., & MANN, S.
Construction and morphogenesis of the chiral ultrastructure of coccoliths from the marine
algaEmiliania huxleyi.
• Proc. Roy. Soc. B., 258: 237-245.

1994

Bioi.
Living

A5258

DMITRENKO, O.B.
Paleoecology of the Paleocene nannoplankton.
t 6th INA Conf. , Copenhagen, 1995, abstracts. J. Nannoplankton Res., 17(2): 55.

1996

Abst.
Ecol.
TERT.L.
Atlantic;
IndianA5259

Dl\IITRENKO, O.B.
Neogene-Quatemary nannoplankton ofDSDP Site 362, the South Atlantic.
t Stratigr. Geol. Correlation, 3(4): 406-409.

1995

Strat.
QUAT .;TERT.U.

A5260

EGGE,J.K & HEIMDAL, B.R.
Blooms of phytoplankton including Emiliania huxleyi (Haptophyta). Effects of nutrient
supply in different N :P ratios.
t Sarsia, 79: 333-348.

1994

Bioi.
Living

A5261

ESHET, Y.
Obtaining rich nannofossil assemblages from "barren" samples: processing organic-rich rocks
in nannofossil investigations.
t J. Nannoplankton Res., 18(1): 17-21.

1996

Tech.

A5262

ESHET, Y. & ALMOGI-LABIN,A.
Calcareous nannofossils as paleoproductivity indicators in Campanian-Maastrichtian
upwelling systems.
t 6th INA Conf., Copenhagen, 1995, abstracts. J. Nannoplankton Res., 17(2): 55-56.

1996

Abst.
Ecol.
CRET.U.
Asia SW

A5263

ESHET, Y., MINSTER. T., & MOSHKOVITZ, S.
Calcareous na~nofossil datum levels: a key for the understanding of the genesis ofUpper Cretaceous oil shale sections in Israel.
t 6th INA Conf., Copenhagen, 1995, abstracts. J. Nannoplankton Res., 17(2): 56.

1996

Abst.
Strat;Ecol.
CRET.U.
Asia SW

A5264

ESHET, Y. & MOSHKOVITZ, S.
New nannofossil biostratigraphy for Upper Cretaceous organic-rich carbonates in Israel.
t Micropaleontology, 41(4): 321 -341.

1995(Strat.);(Syst);Ecol
CRET .U .
Asia SW=========================71==========================

Journal of Nannoplankton Research,l8, 2.

W G. Siesser: Calcareous nannoplankton bibliography 27, p. 69 - 83 .

-----------------------------------A5265

ESHET, Y. & MOSHKOVITZ, S.
Calcareous nannofossil biostratigraphy in Santonian-Maastrichtian organic-rich carbonate
sections in Israel.
+6th INA Conf., Copenhagen, 1995, abstracts. J. Nannoplankton Res., 17(2): 56-57.

1996

Abst.
Strat.
CRET.U.
Asia SW

A5266

FAGERBAKKE, KM., HELDAL, M., NORLAND, S., HEIMDAL, B.R.,& BATVIK, H.
Emi/iania huxleyi. Chemical composition and size of coccoliths from enclosure experiments
and a Norwegian fjord.
+ Sarsia, 79: 349-355.

1994

Bioi.
Living

A5267

FINDLAY,C.
Calcareous nannoplankton from the Australian sector of the Southern Ocean.
+ 6th INA Conf., Copenhagen, 1995, abstracts. J. Nannoplankton Res., 17(2): 57.

1996

Abst.
Ecol.
LivingPacific S.

A5268

FIORENTINO,A.
Maastrichtian nannofossil biogeography in the southern Tethys.
+6th INA Conf., Copenhagen, 1995, abstracts. J. Nannoplankton Res., 17(2): 57.

1996

Abst.
Ecol.
CRET.U.
Europe W.

A5269

FLOOD, R.D., PIPER, D.J.W., KLAUS,A., & THE SHIPBOARD
SCIENTIFIC PARTY.
+ Proc. ODP, Init. Repts., v. 155 (Amazon Fan; Sites 930-946). 1233 pp.

1995

Strat.
QUAT.;TERT.U.

A5270

FLORES, J.A., SIERRO, F.J., & RAFFI, L
Evolution of the calcareous nannofossil assemblage as a response to the paleoceanographic
changes in the eastern equatorial Pacific Ocean from 4 to 2Ma (Leg 138, Sites 349 and 852).
+ In: Pisias, N. G., Mayer, L.A., Janecek, T.R., Palmer-Julson, A., & van Andel, T.H. (Eds.),
138: 163-176.

1995

Ecol.;Diag.
TERT.U.
Pacific C.

A5271

FLORES, J.A., SIERRO, F.J., ZAMARRENO,A.V., UTRILLA, R. & FRANCES, G.
An example of eutrophic-oligotrophic alternates in the lower Messinian: the Sorbas Basin(western mediterranean).
+ 6th INA Conf., Copenhagen, 1995, abstracts. J. Nannoplankton Res., 17(2): 58.

1996

Abst.
Ecol.
TERT .U.
Europe W.

A5272

FRYDAS. P.D.
Biostratigraphy using silicoflagellates and calcareous nannofossils ofNeogene-Pleistocene
deposits of Greece.
+6th INA Conf., Copenhagen, 1995, abstracts. J. Nannoplankton Res., 17(2): 58.

1996

Abst.
(Strat.)QUAT .;TERT.U.
Europe W.

A5273

GARCIASOTO, C., FERNANDEZ, E., PINGREE, R.D., & HARBOUR, D.S.
Evolution and structure of a shelf coccolithophore bloom in the western English Channel.
+ J. Plankton Res., 17(11): 2011-2036.

1995

Ecol.
LivingAtlantic N.

A5274

GARDIN, S. & PERILLI, N.
Early Cretaceous calcareous nannofossil biostratigraphy of the internal Ligurids (northern
Apennines, Italy) and the onset of coccolith-bearing sediments in the Ligurian-Piedmont
oceanic domain.
+ 6th INA Conf., Copenhagen, 1995, abstracts. J. Nannoplankton Res., 17(2): 59-60.

1996

Abst.
Strat.
CRET.L.
Europe W.

A527 5GAYOSO, A. M.
Bloom of Emiliania huxleyi (Prymnesiophyceae) in western South Atlantic Ocean.
+ J. Plankton Res., 17(8): 1717-1 722.

1995

Ecol.
LivingAtlantic S.

A5276

GIBSON, T.G. & BYBELL, L. M.
Paleogene stratigraphy of the Solomons Island, Maryland corehole.
+ U.S. Geol. Survey Open-file Rept. 94-708, pp. 1-39.

1994

Strat.
TERT.L.
America N.

A5277

GIBSON, T.G. & BYBELL, L.M.
Sedimentary patterns across the Paleocene-Eocene boundary in the Atlantic andGulf Coastal Plains of the United States.
+Bull. Soc. Beige Geol., 103(3-4): 237-265.

1994

Strat.
TERT .L.
America N.

A5278

GIRAUDEAU,J. & BAILEY, G.W.
Sediment trap studies of coccolith and planktonic foraminiferal fluxes in the Benguela Upwelling System.
+ 6th IN'A Conf. , Copenhagen, 1995, abstracts. J. Nannoplankton Res., 17(2): 61.

1996

Abst.
Ecol.
LivingAtlantic S.

========================~ 72 ========================~

W. G. Siesser: Calcareous nannoplankton bibliography 27, p. 69 - 83.

A5279

Journal of Nannoplankton Research,l8, 2.

GREEN, J.C. & HO RI, T.

1994

Bioi.

GREEN, J .C. & JORDAN, R. W.
Systematic history and taxonomy.
+ In: Green, J.C. & Lead beater, B.S.C. (Eds.), The Haptophyte Algae. Systematics Assn.
Spec. Vol. No. 51, Clarendon Press, Oxford, pp. 1-2 1.

1994

Overview
Syst.

HAY,W.W.

1996

Abst.

Flagella and flagellar roots.
• In: Green, J.C . & Lead beater, B.S.C. (Eds.), The Haptophyte Algae. Systematics Assn.
Spec. Vol. No. 51, Clarendon Press, Oxford, pp. 47-71.
A5280

A5281

Oceanic fronts in the Mesozoic.
• 6th INA Conf. , Copenhagen, 1995, abstracts. J. Nannoplankton Res., 17(2): 61-62.

Ecol.
CREf.;JURAWorldwide

A5282

HECK, S. E. van
The ICBN: things you need to know- 13.
+ J. Nannoplankton Res., 18(1): 12-13.

1996

Syst.

A5283

HECK, S.E. vanResults of the survey on mounting media.
·· J. Nannoplankton Res., 18(1): 15-16.

1996

Tech. Prep.

HEIMDAL, B.R., EGGE, J.K., VELDHUIS, MJ.W.,& WESTBROEK, P.

1994

Overview
Living

1995

Ecol.
QUAT .
Pacific N.

1996

Abst.
Ecol.
CRET .L.-U.
Europe W.

1994

Bioi.

1996

Abst.
Strat.
CRET.L-U.
Australasia

HRADECKA, L. & SVABENICKA, L.
Foraminifera and calcareous nannoplankton assemblages from the Cenomanian-Turonianboundary interval of the Knoviz section, Bohemian Cretaceous Basin.
+ Geologica Carpathica, 46(5): 267-276.

1995

Strat.;(Syst.);
Ecol.
CRET.L.-U.
Europe E.

HUBER,B.T.&WATKINS,D.K

1992

Ecol. •
CRET.U.
Atlantic S
Indian

1994

Bioi.

1996

Abst.
Overview

A5284

The 1992 Norwegian Emiliania huxleyi experiment. An overview.
• Sarsia, 79: 285-290.
A5285

HERBERT, T.D., YASUDA, M., & BURNETT, C.
Glacial-interglacial sea-surface temperature record inferred from alkenone unsaturation
indices, Site 893, Santa Barbara Basin.
+ In: Kennett, J.P., Baldauf, J.G., Lyle, M. (Eds). Proc. ODP, Sci. Results, 146(2): 257-264.

A5286

HOMEIER, G.
Mid-Cretaceous calcareous nannoplankton in an organic-rich facies of the Subbetic andPenibetic (Betic Cordillera, Southern Spain).
+ 6th INA Conf., Copenhagen, 1995, abstracts. J. Nannoplankton Res., 17(2): 63 .

A5287

HO RI, T. & GREEN, J.C.
Mitosis and cell division.
+ In: Green, J.C. & Lead beater, B.S.C. (Eds.), The Haptophyte Algae. Systematics Assn.
Spec. Vol. No. 51, Clarendon Press, Oxford, pp. 91-109.

A5288

HOWE, R.
Mid-Late Cretaceous nannofossil biostratigraphy of the Papuan Basin, Papua, New Guinea.
• 6th INA Conf., Copenhagen, 1995, abstracts. J. Nannoplankton Res., 17(2): 64.

A5289

A5290

Biogeography of Campanian-Maastrichtian calcareous plankton in the region of the Southern
Ocean: paleogeographic and paleoclimatic implications.
• The Antarctic ptlooenvironment a perspective on global change. Antaretic Res,Ser., 56: 31-60.
A5291

INOUYE,L&KAWACHI,M.
The haptonema.
+ In: Green, J.C. & Leadbeater, B.S.C . (Eds.), The Haptophyte Algae. Systematics Assn.
Spec. Vol. No. 51, Clarendon Press, Oxford, pp. 73-89.

A5292

JAKUBOWSKI, M.
Biostratigraphy in the oil industry.
+ 6th INA Conf., Copenhagen, 1995, abstracts. J. Nannoplankton Res., 17(2): 63.

=========================73==========================

Journal of Nannoplankton Research, IS, 2.

W. G. Siesser: Calcareous nannoplankton bibliography 27, p. 69 - 83 .

------------------------------------A5293

JANIN, M.-C., SAMSON, Y., SAINT-GERMES, M., BAUDIN, F., & BIGNOT, G.
Comparative variation of calcareous nannofossils (especially Ellipsagelosphaeraceae),
foraminifers and organic matter concentration in the Kimmeridgian sequence from La
Briere (Normandy, France).
+ 6th INA Conf., Copenhagen, 1995, abstracts. J. Nannoplankton Res., 17(2): 65 .

1996

Abst.
Ecol.
JURA.U.
Europe W.

A5294

JEFFREY,S.W.&WRIGHT,S.W.
Photosynthetic pigments in the Haptophyta.
+ In: Green, J.C. & Leadbeater, B.S.C. (Eds.), The Haptophyte Algae. Systematics Assn.
Spec. Vol. No. 51, Clarendon Press, Oxford, pp. 111-132.

1994

Bioi.

A5295

JEFFREY, S.W., BROWN, M.R., & VOLKMAN, J.K.
Haptophytes as feedstocks in mariculture.
+ In: Green, J.C. & Leadbeater, B .S.C. (Eds.), The Haptophyte Algae. Systematics Assn.
Spec. Vol. No. 51, Clarendon Press, Oxford, pp. 287-302.

1994

Ecol.

A5296

JONES, H.L.J., LEAD BEATER, B.S.C., & GREEN, J.C.
Mixotrophy in haptophytes.
+ In: Green, J.C. & Leadbeater, B.S.C. (Eds.), The Haptophyte Algae. Systematics Assn.
Spec. Vol. No. 51, Clarendon Press, Oxford, pp. 247-263.

1994

Bioi.

A5297

JORDAN, R. W. & GREEN, J.C.
A check-list of the extant Haptophyta of the world.
+ J. Mar. Bioi. Assn. U.K., 74: 149-174.

1994

Overview
Syst.

A5298

JORDAN, R. W., KLELJNE,A., HEIMDAL, B.R., & GREEN,J.C.
A glossary of the extant Haptophyta of the world.
+ J. Mar. Bioi. Assn. U.K., 75: 769-814.

1995

Overview
Living

A5299

JUTSON, D.
The case of the missing nannofossil or "Now you see it, now you don'f'+ 6th INA Conf., Copenhagen, 1995, abstracts. J. Nannoplankton Res., 17(2): 66.

1996

Abst.
Diag.

A5300

KAENEL, E. dePaleocene to early Eocene calcareous nannofossil biostratigraphy of the El Kef section (Tunisia).
+6th INA Conf., Copenhagen, 1995, abstracts. J. Nannoplankton Res., 17(2): 54.

1996

Abst.
Strat.
TERT.L.
Africa N .

A530 1KAENEL, E. de, BERGEN, J.A., & von SALIS PERCH-NIELSEN, K.
Jurassic calcareous nannofossil biostratigraphy of western Europe. Compilation of recent
studies and calibration of bioevents.
+ Bull. Soc. Geol. France, 167(1): 15-28.

1996(Strat.);Syst.
JURA.L.-U.
Europe W.

A5302

KAMEO, K., SATO, T. & TAKAYAMA, T.
Late Pliocene nannofossil bioevents and their biostratigraphic and paleoceanographic meanings.
+ 6th INA Conf., Copenhagen, 1995, abstracts. J. Nannoplankton Res., 17(2): 66-67.

1996

Abst.
Strat.
TERT.U .
Atlantic N.

A5303

KINKEL, H., DITTERT, N. & BAUMANN, K.H.
A 300 kyrs calcareous plankton record of the equatorial South Atlantic: reconstructions ofsurface water productivity and bottom water dissolution cycles.
+ 6th INA Conf., Copenhagen, 1995, abstracts. J. Nannoplankton Res., 17(2): 69.

1996

Abst.
Ecol;Diag.
QUAT .
Atlantic S.

A5304

KNAPPERTSBUSCH, M. & BRUMMER, G ..J.A.
A sediment trap investigation of sinking coccolithophorids in the North Atlantic.
+ Deep-Sea Res. I, 42(7): 1083-1109.

1995

Ecol.;
Sedim.
LivingAtlantic N

A5305

KRISTIANSEN, S., THINGSTAN, T.F., WAL, P. van der, FARBROT, T., &SK.JOLDAL, E. F.
An Emiliania huxleyi dominated subsurface bloom in Samnanger-fjorden, western Norway.
Importance of hydrography and nutrients.
+ Sarsia, 79: 357-368.

1994

Bioi.
LivingEurope W.

A530<;iLEADBEATER, B.S.C.
Cell coverings.
• In: Green, J.C. & Leadbeater, B.S.C. (Eds.), The Haptophyte Algae. Systematics Assn.
Spec. Vol. No. 51, Clarendon Press, Oxford, pp. 23-46.

1994

Bioi.

==========================74=========================

W. G. Siesser: Calcareous nannoplankton bibliography 27, p. 69 - 83 .

A5307

Journal of Nannoplankton Research,l8, 2.

LECOURT, M., MUGGLI, D.L., & HARRISON, P.J.

1996

Bioi.;
Ecol.
Living

1996

Abst.
Ecol.
QUAT.
Atlantic N.

1996

Abst.
Strat.
KffBDY;
CRET.L.-U.
Europe W.

1995

Strat.
TERT.L.-U
Africa N.

1996

Abst.
(Strat.)
QUAT.
Pacific N.

1994

Ecol.

1994

Ecol.

1996

Abst.
Strat.
QUAT.
Europe W.

1996

Abst.
Strat.
KffBDY;TERT.L.
Africa N.

1996

Abst.
Strat.
JURAL-M
Europe W.

1994

Bioi.
Syst.

MERTINIENE, R.
Calcareous nannofossil assemblages in the Cenomanian and Turonian deposits in Lithuania.
+ 6th INA Conf., Copenhagen, 1995, abstracts. J. Nannoplankton Res., 17(2): 74.

1996

Abst.
Strat.
CRET.U.
Europe E.

MESzAROS, N.

1996

Abst.
Strat.
TERT .U.
Europe E.

Comparison of growth and sinking rates of non-coccolith-and coccolith-forrning strains ofEmiliania huxleyi (Prymnesiophyceae) grown under different irradiances and nitrogen sources.
+ J. Phycol., 32: 17-21.
A5308

LOTOSKA YA,A. & ZIVERI, P.
Calcareous nannoplankton response to Termination II and Heinrich Event (140-110 ka BP)
in the NE Atlantic.
• 6th INA Conf., Copenhagen, 1995, abstracts. J. Nannoplankton Res., 17(2): 70-71.

A5309

L07AR, F.
Calcareous nannoplankton biostratigraphic analysis as a tool for the study of discontinuity
surfaces in the Cretaceous of the Trento Plateau (southern Alpine, Italy).
+ 6th INA Conf., Copenhagen, 1995, abstracts. J. Nannoplankton Res., 17(2): 71.

A531 0

MAA TE,A., MARTiN-PEREZ,J.A., MARTiN-ALGARRA, SERRANO, F.,
AGUADO, R, MARTiN-MARTiN, M., & EL HAJJAJI, K.
The lower Burdigalian ofBoujarrah (northern Rif, Morocco) and the palaeotectonic significanceof the Miocene transgressive series on the Betic-Rifian internal zones.
• C.R. Acad. Sci. Paris, 320, Ser.II: 15-22.
(In French with English abstract and English abridged version)

A5311

MAIORANO, P., MARINO, M., & MONECHI, S.
Pleistocene calcareous nannofossil biostratigraphy of Site 577 Leg 86, northwestern Pacific Ocean.
+6th INA Conf., Copenhagen, 1995, abstracts. J. Nannoplankton Res., 17(2):71-72.

A5312

MALIN, G., LISS, P.S., & TURNER, S.M.
Dimethyl sulfide: production and atmospheric consequences.
+ In: Green, J.C. & Lead beater, B.S.C. (Eds.), The Haptophyte Algae. Systematics Assn.
Spec. Vol. No. 51, Clarendon Press, Oxford, pp. 303-320.

A5313

MARCHANT, H.J. & THOMSEN, H.A.
Haptophytes in polar waters.
+ In: Green, J.C. & Leadbeater, B.S.C. (Eds.), The Haptophyte Algae. Systematics Assn.
Spec. Vol. No. 51, Clarendon Press, Oxford, pp. 209-228.

A5314

MARINO,M.
Calcareous nannofossil biostratigraphy of early-middle Pleistocene terrigenous sediments(southern Italy).
t 6th INA Conf., Copenhagen, 1995, abstracts. J. Nannoplankton Res., 17(2):72-73.

A5315

MARZOUK,A.M. &ABOU-EL-ENEIN, M.K.
Calcareous nannofossil biostratigraphy of the late Cretaceous-early Tertiary ofWadi Feiran and Gebel Qabeliat, south western Sinai, Egypt.
• 6th JNA ConL Copenhagen, 1995, abstracts. J. Nannoplankton Res., 17(2): 73.

A5316

MA.TTIOLI, E. & PERILLI, N.
Calcareous nannofossils biostratigraphy of the Fuentelsaz (Spain) and Fiurninata (Italy)
sections: Late Toarcian-Early Aalenian.
+6th INA Conf., Copenhagen, 1995, abstracts. J. Nannoplankton Res., 17(2): 73-74.

A5317

MEDLIN, L.K, BARKER, G.L.A., BAUMANN, M., HA YES, P.K., & LANGE, M.
Molecular biology and systematics.
+ In: Green, J.C. & Leadbeater, B.S.C. (Eds.), The Haptophyte Algae. Systematics Assn.
Spec. Vol. No. 51, Clarendon Press, Oxford, pp. 393-411.

A5318

A5319

Nannoplankton zones in the volcanic tufTs of the Transylvanian Basin (Miocene).
t 6th INA Conf., Copenhagen, 1995, abstracts. J. Nannoplankton Res., 17(2): 75.

=========================75==========================

Journal of Nannoplankton Research, 18, 2.

W G. Siesser: Calcareous nannoplankton bibliography 27, p. 69 - 83 .

------------------------------------A5320

MIKKELSEN, N.
High resolution biostratigraphy of the Amazon Fan+ 6th INA Conf., Copenhagen, 1995, abstracts. J. Nannoplankton Res., 17(2): 76.

1996

Abst.
Strat.
QUAT.
Atlantic C.

A5321

MILITANTE-MATIAS,P. &DeLEON,M.M.
A study of the foraminiferal and calcareous nannoplankton assemblages of core samples
taken from the lahar-affected portions of the south China Sea in Zambales Province.
+ 6th INA Conf., Copenhagen, 1995, abstracts. J. Nannoplankton Res., 17(2): 76.

1996

Abst.
Strat.
QUAT.
Pacific C.
Atlantic C.

A5322

MOESTRUP, 0 .
Economic aspects: ' blooms', nuisance species, and toxins.
+ In: Green, J.C. & Leadbeater, B.S.C. (Eds.), The Haptophyte Algae. Systematics Assn.
Spec. Vol. No 51, Clarendon Press, Oxford, pp. 265-285 .

1994

Ecol.

A5323

MOSTAJO, E., CONCHEYRO,A., & DZQUEZ, F.
Calcareous nannofossils (Hauterivian) from "Agua De La Mula" section, Neuquen
Province, Argentina.
+ 6th INA Conf., Copenhagen, 1995, abstracts. J. Nannoplankton Res., 17(2): 76.

1996

Abst.
Strat.
CRET.L.
AmericaS.

A5324

MUTTERLOSE,J.
Calcareous nannofossil paleoceanography of the Lower Cretaceous of northwest Europe.
+ 6th INA Conf., Copenhagen, 1995, abstracts. J. Nannoplankton Res., 17(2): 77.

1996

Abst.
Ecol.
CRET.L.
Europe W.

A5325

MYHERE,A.M., THIEDE,J., FIRTH,J.V.,& THE SHIPBOARD SCIENTIFIC PARTY.
+ Proc. ODP, Init. Repts., v. 151 (North Atlantic -Arctic Gateways I; Sites 907-913).
926 pp.
[Nannofossils by T. Sato]

1995

Strat.
QUAT.;TERT.L.-U.
Atlantic N.

A5326

NAGYMAROSY,A.
The nannoplankton of the Jaslo Limestone horizons,+ 6th INA Conf., Copenhagen, 1995, abstracts. J. Nannoplankton Res., 17(2): 77-78.

1996

Abst.
Strat.
TERT .L.
Europe W.

A5327

NAJI, F.
Calcareous nannoplankton biostratigraphy of the Salzgitter-Salder limestone quarry,
Lower Saxony, Germany.
+6th INA Conf., Copenhagen, 1995, abstracts. J. Nannoplankton Res., 17(2): 78-79.

1996

Abst.
Strat.;(Strat.)CRET.U.
Europe W.

A5328

NEJSTGAARD, J.C., WITTE, H.J., WAL, P. van der, & JACOBSEN,A.
Copepod grazing during a mesocosm study of an Emi/iania huxleyi (Prymnesiophyceae) bloom.
+ Sarsia, 79: 369-377.

1994

Bioi.
Living

A5329

NffiLSEN, M. V.
Photosynthetic characteristics of the coccolithophorid Emiliania hwcleyi (Prymnesiophyceae)
exposed to elevated concentrations of dissolved inorganic carbon.
+ J. Phycol., 31 : 715-719.

1995

Bioi.
Living

A5330

OLAFSSON, G. & BEAUFORT, L.
Data Report: Oligocene-Pieistocene calcareous nannofossils from Leg 145.

1995

Strat.
QUAT.;+ In: Rea, DK., Baoov, lA., Scholl, D.W., &Allan, JF. Eds.), Proc. ODP, (Sci. Results, 145: 599-632.

TERT.L.-U.

A5331

0ZKAN, S., & ALTINER, D.
Nannofossils of the Maastrichtian-Paleocene Germav Formation, southeast Anatolia, Turkey.
+ 6th INA Conf., Copenhagen, 1995, abstracts. J. Nannoplankton Res., 17(2): 46.

1996

Abst.
Strat.
Kff BDY
Asia SW

A5332

PAVSIC,J. & DOLENEC, T.
Floristic and isotopic changes at the Paleocene-Eocene boundary in Slovenia.
+ 6th INA Conf., Copenhagen, 1995, abstracts. J. Nannoplankton Res., 17(2): 79.

1996

Abst.
Ecol.
TERT.L.
Europe E.

A5333

PELEO-AIAMPAY,A.
Morphometric and biochronologic study of Coccolithus miope/agicus.
+ 6th INA Conf, Copenhagen, 1995, abstracts. J. Nannoplankton Res., 17(2): 80.

1996

Abst.
Strat.
TERT.U.
Worldwide==========================76===========================

W G. Siesser: Calcareous nannoplank.ton bibliography 27, p. 69 - 83 .

Journal of Nannoplankton Research, 18, 2.

------------------------------------A5334

POSPICHAL, J.J.
Calcareous nannofossils, tsunamites, and the Krr boundary of northeastern Mexico.
+6th INA Conf, Copenhagen, I995, abstracts. J. Nannoplankton Res., I7(2): 80-81.

1996

Abst.
Strat.
Kff BDY
America N.

A5335

POSPICHAL, J.J.
Calcareous nannofossils and clastic sediments at the Cretaceous-Ter1iruy boundary, northeastern Mexico.
+ Geology, 24(3): 255-258.

1996

Strat.
Kff BDY
America N .

A5336

PRAMPARO, M., VOLKHEIMER, W. & MOSTAJO, E.
Calcareous nannofossils and palynomorphs from Valanginian and Hauterivian (Lower
Cretaceous) Strata of Cerro De La Parva, central-western Neuquen, Argentina.
+ 6th INA Conf., Copenhagen, I 995, abstracts. J. Nannoplankton Res., I 7(2): 8 I.

1996

Abst.
Strat.
CRET.L.
AmericaS.

A5337

PURDIE, D.A. & FINCH, M.S.
Impact of a coccolithophorid bloom on dissolved carbon dioxide in sea water enclosuresin a Norwegian fjord.
+ Sarsia, 79: 379-387.

1994

Bioi.
LivingEurope W.

A 5338RAFFI, L & FLORES, J.-A.
Pleistocene through Miocene calcareous nannofossils from eastern equatorial
Pacific Ocean (Leg 138).
+ In: Pisias, N.G., Mayer, L.A., Janecek, T.R., Palmer-Julson, A., &van Andel, T.H. (Eds.). Proc. ODP, Sci. Results, 138: 233-286.

1995(Strat.);(Syst.)QUAT .;TERT.L.-U.
Pacific C.

A5339

RAFFI,L, RIO, D., d' ATRI,A., FORNACIARI, E., & ROCCHETTI, S.
Quantitative distribution patterns and biomagnetostratigraphy of middle and late Miocene
calcareous nannofossils from equatorial Indian and Pacific Oceans (Legs I 15, 130, and 138).
+ In: Pisias, N.G., Mayer, L.A., Janecek, T.R., Palmer-Julson, A., &van Andel, T.H. (Eds.). Proc. ODP, Sci. Results, 138: 479-502.

1995(Strat.)
TERT.U.
Pacific C;
Indian

A5340

RAJU, D.S.N., JAIPRAKASH, B. C., KUMAR, A., SAXENA, R.K., DAVE,A.,
CHATTERJEE, T.K.,& MISHRA,.C.M.
Age of Deccan volcanism across KTB in Krishna-Godavari Basin: new evidence.
• J. Geol. Soc. India, 45: 229-233.

1995(Strat.)Kff BDY
Asia S.

A534 IRAJU, D.S.N., RAVINDRAN, C.N., MISHRA, P.K, CHIDAMBARAM, L, & SAXENA, R.K.
Stratigraphy and paleoenvironments of the Godavari Clay in the Krishna-Godavari Basin, India.
+ Indian J. Petrol. Geol., 3(2): 33-43.

1994

Strat.
TERT.U.
Asia S.

A5342

REALE, V.
Paleoecology of calcareousnannofossil assemblages and its relationship to sea-level variationsin the Figols Groups (early Eocene, Spanish Pyrenees).
• 6th INA Conf, Copenhagen, I995, abstracts. J. Nannoplankton Res., I7(2): 8I-82.

1996

Abst.
(Strat).
TERT.L.
Europe W.

A5343

RHODES, L.L., PEAKE, B. M., MACKENZIE,A.L.,& MARWICK, S.
Coccolithophores Gephyrocapsa oceanica and Emi/iania huxleyi Prymnesiophyceae equals(Haptophyceae) in New Zealand' s coastal waters: characteristics of blooms and growth inlaboratory cultures.
+ New Zealand J. Mar. Freshwater Res. , 29(3): 345-358.

1995

Bioi.
Living

A5344

ROMANIV,A.
Calcareous nannoplankton from the boundary deposits of Maastrichtian and Danianof the Ukrainian Carpathians.
+ 6th INA Conf, Copenhagen, I995, abstracts. J. Nannoplankton Res., 17(2): 82.

1996

Abst.
Strat.
TERT.L.
CRET .U.
CIS

A5345

SANCHEZ.RIOS,A., RUIZ-RUIZ, lL,PADILLAK, P., & GRAJALES..NISHIMURA,J.M.
Nannofossils and planktic foraminifera across the Kff boundary in two sections ofnorthwestern Mexico.
• 6th INA Conf., Copenhagen, I995, abstracts. J. Nannoplankton Res. , 17(2): 83.

1996

Abst.
(Strat.)Kff BDY
America N.

A5346

SATO, T. & KAMEO, K.
Late Pliocene nannofossil events recognized in the uppermost sequence in theJapan Sea side, with reference to th eglaciation in the Arctic Sea.
• 6th INA Conf., Copenhagen, 1995, abstracts. J. Nannoplanlcton Res., 17(2): 83.

1996

Abst.
Strat.
QUAT .
Pacific N.

========================~77========================~

Journal of NannoplaiJ.ktOn Research,l8, 2.

W. G. Siesser: Calcareous nannoplankton bibliography 27, p. 69 - 83.

A5347

SAXENA,R.K.
High resolution nannoplankton biostratigraphy of late Hauterivian through Daniansubsurface sediments in Krishna-Godavari Basin, India.
+ Proc. PETROTECH II-95, Techn. Trends in Petrol. Ind., New Delhi, pp. 35-42.

1995

Strat.
TERT .L.;
CRET.L.-U.
Asia S.

A5348

SAXENA, R.K. & MISRA, C. M.
Time and duration of Deccan volcanism in Razole area, Krishna-Godavari Basin, India.
+Current Sci., 66(1): 74-76.

1994

Strat.;(Syst.)KIT BDY
Asia S.

A5349

SCHMINCKE, H.-U., WEAVER, P.P.E., FIRTH,J.V., & THE SHIPBOARD
SCIENTIFIC PARTY.
+ Proc. ODP, Jnit. Repts., v. 157 (Gran Canaria and Madeira Abyssal Plain; Sites 950..956). 843 pp.
[Nannofossils by R. Howe and J. Sblendorio-Levy]

1995

Strat.
QUAT.;TERT.L.-U.
Atlantic N .

A5350

SELF-TRAll..,J.
Late Cretaceous biostratigraphy and paleoceanography of the Atlantic Coastal Plains, USA.
+6th INA Conf., Copenhagen, 1995, abstracts. J. Nannoplankton Res., 17(2): 83-84.

1996

Abst.
Strat;Ecol.
CRET.U.
America N.

A5351

SELF-TRAIL,J.M.,BYBELL,LM.
Cretaceous and Paleogene calcareous nannofossil biostratigraphy of New Jersey.
+In: Baker, J.E.B. (Ed.), Contributions to the Paleontology of New Jersey.
Geol. Assn. of New Jersey, XII: 102-139.

1995

Strat.;(Syst.)TERT.L.;
America N.

A5352

SHACKLETON, N.J., BALDAUF, J.G., FLORES, J.-A., lWAI, M., MOORE; T.C. Jr.,RAFFI, l, & VINCENT, E.
Biostratigraphic summary for Leg 138.
+ In: Pisias, N.G., Mayer, L.A., Janecek, T.R., Palmer-Julson, A. & van Andel, T.H. (Eds.),
Proc. ODP, Sci. Results, 138: 517-536.

1995(Stmt.)
QUAT.;TERT.U.
Pacific C.

A5353

SHIPLEY, T.H., OGAWA, Y., BLUM, P., & THE SHIPBOARD SCIENTIFIC PARTY+ Proc. ODP Init. Repts., v. 156 (Northern Barbados Ridge; Sites 947-949). 301 pp.
[Nannofossils by Y. Xu.]

1995

Strat.
QUAT;TERT .L.-U.
Atlantic C.

A5354

SIESSER, W.G.
Paleogene and Neogene productivity in the Indian Ocean.
+ 6th INA Conf., Copenhagen, 1995, abstracts. J. Nannoplankton Res., 17(2): 84.

1996

Abst.
Ecol.
TERT.L.-U.
Indian

A5355

SIESSER, W.G.
Bibliography and taxa of calcareous nannoplankton - 26.
+ J. Nannoplankton Res., 18(1): 33-35.

1996

Bib!.

A5356

SMITH, C.C.
Regional lithostratigraphy and calcareous nannofossil biostratigraphy of the Areola
Limestone Member, Mooreville Chalk, of eastern Mississippi and Alabama.
+ Gulf Coast Assn. Geol. Soc. Trans., 45: 529-536.

1995

Strat.
CRET.U.
America N .

A5357

SPIEGLER,D.& MULLER, C.
Correlation of calcareous nannoplankton and planktonic foraminifer biostratigraphy off
the coast of southern Chile.
+ Iti: Lewis, S.D., Behrmann, J.H., Musgrave, R.J., & Cande, S.C. (Eds.), Proc. ODP,
Sci.-Results, 141 : 193-211.

1995(Stmt.)
QUAT. ;TERT.U.
Pacific S.

A5358

STEFANO, A. d i
Calcareous nannofossils biostratigraphy of the Plio-Pleistocene deposits of thePeloritani Mountains (NE Sicily).
+ 6th INA Conf., Copenhagen, 1995, abstracts. J. Nannoplankton Res., 17(2): 54-55.

1996

Abst.
Strat.
QUAT .
TERT .U.
Europe W.

A5359

SU,X.
Development of the Plio-Pleistocene coccolith assemblages in the NE Atlantic correlated
to climatic changes.
• 6th INA Conf., Copenhagen, 1995, abstracts. J. Nannoplankton Res., 17(2): 84-85.

1996

Abst.
Ecol.
QUAT.;TERT.U.
Atlantic N.

==========================78===========================

W. G. Siesser: Calcareous nannoplankton bibliography 27, p. 69 - 83.

Journal of Nannoplankton Research, I&, 2.

A5360

SUMMERHA YES, C.P., KROON, D., ROSELL-MELE,A., ORDAN, R. W.,
SCHRADER,H.-J.,JHEARN, R., VILLANUEVA,J.,GRIMALT,J.O.,& EGLINTON,G.
Variability in the Benguela Current Upwelling System over the past 70,000 years.
+ Prog. Oceanogr., 35: 207-251.

1995

Strat.
QUAT.
Atlantic S

A5361

SVABENICKA, L.
The stratigraphical correlation of the Campanian low- and high-latitude calcareous
nannofossils in southern Moravia (western Carpathians).
+ Geologica Carpathica, 46(5): 297-302.

1995

Strat.;(Syst.)
CRET.U.
Europe E.

A5362

SVABENICKA, L.
Rare occurrences of Petrarhabdus copulatus Deflandre on the northern hemisphere(Central Europe).
+ 6th INA Conf., Copenhagen, 1995, abstracts. J. Nannoplankton Res., 17(2): 85-86.

1996

Abst.
Ecol.
CRET.U.
Europe E.

A5363

TAKAHASHI,K
Coccolithophorid biocoenosis: production and fluxes to the deep sea.
+ In: Green, J.C . & Leadbeater, B.S.C. (Eds.), The Haptophyte Algae. Systematics Assn.
Spec. Vol. No. 51, Clarendon Press, Oxford, pp. 335-350.

1994

Ecol.
Sedim.

A5364

TAVERA,J.M.,AGUADO, R., COMPANY, M., & OLORIZ, F.
Integrated biostratigraphy of the Durangites and Jacobi Zones (J/K boundary) at the
Puerto Escano section in southern Spain (Province of Cordoba).
+ Geobios, 17: 469-476.

1994(Strat.);(Syst.)
CRET.L.;
Europe W.

A5365

THOMSEN, E.
The Danian (lower Paleocene) limestone deposits in Denmark: basin development
and sequence stratigraphy+ 6th INA Conf, Copenhagen, 1995, abstracts. J. Nannoplankton Res., 17(2): 86.

1996

Abst.
(Strat.)
TERT.L.
Europe W.

A5366

THOMSEN, H.A.
Polar coccolithophorids.
+ 6th INA Con f., Copenhagen, 1995, abstracts. J. Nannoplankton Res., 17(2): 87.

1996

Abst.
Syst.;Ecol.
LivingAntarctic

A5367

THOMSEN, H.A., BUCK, KR., & CHA VEZ, F.P.
Haptophytes as components of marine phytoplankton.
+ In: Green, J.C. & Leadbeater, B.S.C. (Eds.), The Haptophyte Algae. Systematics Assn.
Spec. Vol. No. 51, Clarendon Press, Oxford, pp. 187-208.

1994

Ecol.

A5368

TYRRELL,T.&TAYLOR,A.H.
Latitudinal and seasonal variations in carbon dioxide and oxygen in the northeast Atlantic
and the effects on Emiliania huxleyi and other phytoplankton.
+ Global Biogeochem. Cycles, 9(4): 585-604.

1995

Ecol.
LivingAtlantic N.

A5369

USHAKOVA, M.
Changes of calcareous nannoplankton evolutionary diversity during the Paleogene.
+ 6th INA Conf., Copenhagen, 1995, abstracts. J. Nannoplankton Res., 17(2): 88.

1996

Abst.
Strat;Ecol.
TERT.L.-U.
Worldwide

A5370

VELDHUIS, M.J.W., STOLL, M., BAKKER, D., DRUMMER, G.-J., KRAAK, M.,
KOP,A, WEERLEE, E. van, KOUTRIK,A. van,& HEIMDAL, B.R.
Calcifying phytoplankton in Bjornafjorden, Norway. The prebloom situation.
+ Sarsia, 79: 389-399.

1994

Bioi.
LivingEurope W.

A5371

VOLIN, S., ESHET, Y. & KISH, H.
Utility of calcareous nannofossils and organic remains: in the reconstruction of thermal
metamorphism in the Hatrurim Formation (Tertiary, southern Israel).
+ 6th INA Conf., Copenhagen, 1995, abstracts. J. Nannoplankton Res., 17(2): 88.

1996

Abst.
Diag.
CRET.U.
Asia SW.

A5372

VON SALIS, K
Calcareous nannofossils in the arts.
+ 6th INA Conf., Copenhagen, 1995, abstracts. J. Nannoplankton Res., 17(2): 88-90.

1996

Abst.
(Strat.);Ecol.
Worldwide

A5373

WAL, P. van der, KRAAY, G.W., & MEER,J. van derCommunity structure and nutritional state of phytoplankton growing in mesocosms withdifferent initial N:P ratios studied with high performance liquid chromatography.
• Sarsia, 79: 409-416.

1994

Bioi.
Living===========================79============================

Journal of Nannoplankton Research,l8, 2.

W. G. Siesser: Calcareous nannoplankton bibliography 27, p. 69 - 83.

AS374

WAL,P. vander,DLEIJSWIJK,J.D.L. van,& EGGE,J.K.
Primary productivity and calcification rate in blooms of the coccolithophorid Emiliania
huxleyi (Lohmann) Hay et Mohler developing in mesocosms.
+ Sarsia, 79: 401-408.

1994

Bioi.
Living

AS375

WATKINS, D.K., DRALOWER, T.J., COVINGTON, J.M., & FISHER, C. G.
Biostratigraphy and paleoecology of the Upper Cretaceous calcareous nannofossils in theWestern Interior Basin, North America.
+In: Caldwell, W.G.E. & Kauffman, E.G. (Eds.), Evolution of the Western Interior
Basin, Geol. Assn. Canada Spec. Paper 39, pp. 521-537.

1993

Strat.;(Syst.);
Ecol.
CRET.U.
America N.

AS376

WEI,W.
The initiation of North Atlantic Deep Water as dated by nannofossils.
+ 6th INA Conf., Copenhagen, 1995, abstracts. J. Nannoplankton Res., 17(2): 90-91.

1996

Abst.
Ecol.
TERT.U.
Atlantic N.

A5377

WEI, W. & PELEO-ALAMPAY,A.
Data Report: calcareous nannofossils of Hole 893A in the Santa Barbara Basin.
+ In: Kennett, J.P., Baldauf, J.G. & Lyle, M. (Eds.), Proc. ODP., Sci. Results,
146(2): 329-330.

1995

Strat.
QUAT.;TERT.U.
Pacific N.

A5378

WEI, W. & ZHONG, S.
Taxonomy and magnetobiochronology of Tribrachiatus and R.homboaster, two generaof calcareous nannofossils.
+ J. Paleontol., 70(1): 7-22.

1996

Syst.
TERT.L.

AS379

WESTDROEK, P., V AN HINTE,J.E., DRUMMER, G.-J., VELDHUIS, M.,
DROWNLEE,C., GREEN,J.C., HARRIS, R., & HEIMDAL D.R.
Emiliania huxleyi as a key to biosphere-geosphere interactions.
+ In: Green, J.C. & Leadbeater, B.S.C. (Eds.), The Haptophyte Algae. Systematics Assn.
Spec. Vol. No. 51, Clarendon Press, Oxford, pp. 321-334.

1994

Ecol.

AS380

WINTER, A. & JORDAN, R. W.
Living coccolithophorid assemblages off the Puerto Rican Shelf.
+ 6th INA Conf., Copenhagen, 1995, abstracts. J. Nannoplankton Res., 17(2): 91-92.

1996

Abst.
Ecol.
LivingAtlantic C.

A5381

WISE, JR., S.W., DARREN, L.T., MILMAN, D., PAYTON, W.C., POSPICHAL, J.J.,
MUZA, J., MAIORANO, P., CONCHEYRO,A., VILLA, G., PERCH-NIELSEN, K.,
DYDELL, L.M., SELF-TRAIL,J.M., MURPHY, W.L., RIEDEL, W.R.,& XU, Y.
Nanno notes: an interactive digital image catalog of Cenozoic calcareous nannofossils.
+ 6th INA Conf., Copenhagen, 1995, abstracts. J. Nannoplankton Res., 17(2): 90.

1996

Abst.
Tech.

AS382

YOUNG,C.R.
Non-conventional nannoplankton biostratigraphy applied to biosteering horizontal wells.
+ 6th INA Conf., Copenhagen, 1995, abstracts. J. Nannoplankton Res., 17(2): 92.

1996

Abst.
Strat.
Tech.
Atlantic N .

A5383

YOUNG,J.R.
Variation in Emiliania huxleyi coccolith morphology in samples from the Norwegian
EIDJX experiment, 1992.
+ Sarsia, 79: 417-425 .

1994

Bioi.
LivingEurope W.

AS384

YOUNG, J.R., DOWN, P.R., & DURNETT, J.A.
Palaeontological perspectives.
+ In: Green, J.C. & Leadbeater, B.S.C. (Eds.), The Haptophyte Algae. Systematics Assn.
Spec. Vol. No. 51, Clarendon Press, Oxford, pp. 379-392.

1994

Overview

A5385

YOUNG,J.R., DURNETT, J.A. & KUCERA, M.
Coccobiom - a light microscope based image capture and morphometries application for
Emi/iania huxleyi and other coccoliths.
+ 6th INA Conf., Copenhagen, 1995, abstracts. J. Nannoplankton Res., 17(2): 92-94.

1996

Abst.
Tech.

AS386

ZALAT,A.A.
Calcareous nannoplankton and diatoms from the Eocene-Pliocene sediments,

1995

Strat.
TERT.L.-U.

Africa N.

Fayoum depression, Egypt.
+ J. African Earth Sci., 20(3-4): 227-244.

80

W. G. Siesser: Calcareous nannoplankton bibliography 27, p. 69 - 83.

Journal of Nannoplankton Research,l8, 2.

A5387

ZELTNER,A.
Studies of coccolithophorids from the Arabian Sea and the north Indian Ocean.
• 6th INA Conf., Copenhagen, 1995, abstracts. J. Nannoplankton Res., 17(2):94.

1996

Abst.
Ecol.
LivingIndian

A5388

ZIVERI,P., THUNELL,R.C.,& RIO, D.
Seasonal changes in coccolithophore densities in the Southern California Bight during
1991-1992.
• Deep-Sea Res., 42(11112): 1881-1903.

1995(Syst.);
Ecol.
LivingPacific N.

CALCISPHERES
A5389

HILDEBRAND-HABEL, T. & WILLEMS, H.
Calcdinocysts from the middle Coniacian to upper Santonian Chalks at Laegerdorf(N Germany).
• 6th INA Conf., Copenhagen, 1995, abstracts. J. Nannoplankton Res., 17(2): 62.

1996

Abst.
Syst.;Strat.
Ecol.
CRET.U.
Europe W.

A5390

HOELL, C.
Correlation of calcareous and organic-walled dinoflagellate cyst associations inglacial/interglacial cycles, an example of the Equatorial Atlantic.
• 6th INA Conf., Copenhagen, 1995, abstracts. J. Nannoplankton Res., 17(2): 63 .

1996

Abst.
Ecol.
QUAT.
Atlantic C .

A5391

JANOFSKE, D., KEUPP, H., & WILLEMS, H.
Calcareous dinoflagellate cysts: ultrastructure types and systematic application.
• 6th INA Conf., Copenhagen, 1995, abstracts. J. Nannoplankton Res., 17(2): 64-65 .

1996

Abst.
Bioi.;Syst.

A5392

KERNTOPF, B.
Calcareous dinoflagellates of the equatorial Atlantic Ocean.
• 6th INA Conf., Copenhagen, 1995, abstracts. J. Nannoplankton Res., 17(2): 67.

1996

Abst.
Ecol.
LivingAtlantic C.

A5393

KEUPP,IL
Calcareous dinoflagellate cysts of the late Albian and their environmental deductions(borehole Kirchrode 1/91, Germany).
• 6th INA Conf., Copenhagen, 1995, abstracts. J. Nannoplankton Res., 17(2): 68.

1996

Abst.
Ecol.
CRET.L.
Europe W.

A5394

KIENEL, V.
Development of calcareous dinof•agellate cysts (Calciodinellaceae Deflandre, 1947)
at the Cretaceous-Tertiary boundary (NE Germany, Denmark).
• 6th INA Conf., Copenhagen, 1995, abstracts. J. Nannoplankton Res., 17(2): 68-69.

1996

Abst.
Strat;Syst.
Kff BDY
Europe W.

A5395

KIENEL, U., REHFELD, U., BELLAS, S. & KOHRING, R.
The Miocene Blue Clay Formation of the Maltese Islands: sequence-stratigraphic and paleoceanographic
implications based on calcareous nannofossil stratigraphy and calcareous dinoflagellate cysts.
• 6th INA Conf., Copenhagen, 1995, abstracts. J. Nannoplankton Res., 17(2): 69-70.

1996

Abst.
Strat;Ecol.
TERT.U.
Europe W.

A5396

MONTRESOR. M.
Scrippsiella ramonii sp. nov. (Peridiniales, Dinophyceae), a marine dinoflagellate,
producing a calcareous resting cyst.
• Phycologia, 34(1 ): 87-91.

1995

Syst.
LivingMediterr.

A5397

MONTRESOR, M., ZINGONE,A., & MARINO, D.
The calcareous resting cyst of Pentapharsodinium tyrrhenicum comb. nov. (Dinophyceae).
+ J. Phycol., 29: 223-230.

1993

Syst.
LivingMediterr.

A5398

MONTRESOR, M., MONTESARCHIO, E., MARINO, D., & ZINGONE,A.
Calcareous dinoflagellate cysts in marine sediments of the Gulf of Naples (Mediterranean Sea).
• Rev. Palaeobot. Palynol., 84: 45-56.

1994

Syst.;Ecol.
QUAT.
Mediterr.

A5399

WILLEMS, H.
Calcareous dinoflagellate cysts from the Upper Cretaceous white chalk facies of N. Germany(Coniacian to Maastrichtian).
+Senckenberg. Lethaea, 68(5/6): 433-477.
(In German with English abstract)

1988

Syst.
CRET.U.
Europe W.

===========================81============================

Journal of Nannoplankton Research, IS, 2.
A5400

W G. Siesser: Calcareous nannoplankton bibliography 27, p. 69 - 83.

WILLEMS, R

1994

Syst.
CRET.U.
Europe W.

1996

Abst.
Strat.
KIT BDY
Europe W.

1995

Strat.
KIT BDY
Worldwide

New calcareous dinoflagellates from the Upper Cretaceous white chalk of northern Germany.
+ Rev. Palaeobot. Palynol., 84: 57-72.
A540 1WILLEMS, R
Calcareous dinoflagellate cysts at the KIT boundary of the Geulhemmerberg(Limburg, The Netherlands).
+ 6th INA Conf., Copenhagen, 1995, abstracts. J. Nannoplankton Res., 17(2): 91.

A5402

WILLEMS, H. & KIENEL, U.
Development of calcareous dinocysts (Calciodinellaceae, Deflandre, 1947) at theCretaceous/Tertiary boundary.
+In: Terra Nostra, Global Change and Marine Geology, 85: 49-50.

OTHER TITLES
A5403

BACKMANN,J.
Book Review - Coccolithophores.
+ Palaeogeogr., Palaeoclimatol., Palaeoecol., 118: 291-295.

1995

Review

A5404

BOWN, P.R.
Book Review - Coccolithophores.
+ BMS Newsltr. Micropalaeontol., 52: 32-33.

1995

Review

A5405

BOWN, P. & BURNETT, J.
Subcommission on Cretaceous stratigraphy 2nd International Symposium on Cretaceousstage boundaries, Brussels 8th-16th September 1995.
+ J. Nannoplankton Res., 18(1): 10-11.

1996

Report

A5406

BOWN, P., BURNETT, J., & YOUNG, J.
The 6th International NannoplanktonAssociation Conference, Copenhagen, 2nd-7th September 1995.
+ J. Nannoplankton Res., 18(1): 5-9.

1996

Report

A5407

BURNETT, J. on behalf of the INA committee.
News and gossip.
+ J. Nannoplankton Res., 18(1): 4-5

1996

Report

A5408

HECK, S.E. vanBook Review - International Code of Botanical Nomenclature (Tokyo Code).
+ J. Nannoplankton Res., 18(1): 4-5.

1996

Review==========================82=========================

W G. Siesser: Calcareous nannoplankton bibliography 27, p. 69 - 83 .

Journal of Nannoplankton Research,l8, 2.

Bl84

TAXA
Parhabolithus cavus (PRINS 1969 ex ROOD, HAY & BARNARD 1973 KAENEL, BERGEN& von SALIS PERCH-NIELSEN 1996, p. 27: (ex Crepidolithus).
Rotelapillus helotatus (WIND & WISE in WISE & WIND 1970 KAENEL, BERGEN& von SALIS PERCH-NIELSEN 1996, p. 27 (ex Corollithion).

A5301
A5301

Calcispheres
Amphora WILLEMS 1994, pp. 66-67. Type species: Amphora corona/a WILLEMS 1994.
Amphora corona/a WILLEMS 1994, pp. 67-69, pi. 3, figs. 1~; pi. 4, figs 1-4.
Northern Germany; lower Campanian.
Bicarinellum tumulosum WILLEMS 1988, pp. 458-461, pi. 6, figs. 28-29.
Northern Germany; upper Campanian.
Obliquipithonella echinosa (KEUPP 1981) WILLEMS 1988, pp. 456-457; (ex Pithonel/a ).
Orthopithone/la aequi/amellata WILLEMS 1988, pp. 446-449, pi. 3, figs. 11-16.
Northern Germany; middle/upper Santonian.
Orthopithone/la wi/liambensonii (BOLLI 1978) WILLEMS 1988, pp. 449; (ex Bonetocardie/la).
Pentapharsodinium INDELICATO & LOEBLICH 1986 emend. MONTRESOR,ZINGONE & MARINO 1993, p. 229.
Pentapharsodinium tyrrhenicum (BALECH 1990) MONTRESOR, ZINGONE & MARINO 1993,
p. 229; (ex Peridinium).
Pithone/la pyramidalis WILLEMS 1994, pp. 63-66, pi. 2, figs. 1-7. Northern Germany; lower Campanian.
Scrippsie/la ramonii MONTRESOR 1995, p. 88, figs. 1-13. Gulf of Naples; living.

A5400
A5400

A5399
A5399
A5399
A5399
A5397
A5397
A5400
A5396

NEW CALCISPHERE TAXA
GeneraAmphoraSpeciesaequilamellata, Orthopithonella
coronata, Amphorapyramidalis, Pithonellaramonii, Scrippsiellatumulosum, Bicarinellum==============~=========83==========================

Journal of Nannoplank.ton Research, IS, 2.

INA membership changes, p. 84- 85.

-----------------------------------INA MEMBERSHIP CHANGES
compiled by Kevin Cooper

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=========================84=========================

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==========================85==========================