Bull. Natl. Mus. Nat. Sci., Ser. B, 40(4), pp. 145–151, November 22, 2014

Nomenclatural Changes in the Genus Calanthe (Orchidaceae)
Tomohisa Yukawa1,* and Phillip Cribb2
1

Department of Botany, National Museum of Nature and Science,
Amakubo 4–1–1, Tsukuba, Ibaraki 305–0005, Japan
2
Royal Botanic Gardens, Kew, Richmond, Surrey, UK* E-mail: yukawa@kahaku.go.jp(Received 12 August 2014; accepted 24 September 2014)

Abstract Results of recent molecular phylogenetic analyses revealed that the genus Calanthe(Orchidaceae) represents a polyphyletic assemblage. To resolve this issue, we resurrect twogroups, Preptanthe and Styloglossum, at generic rank and make the necessary new combinations.
Key words : Calanthe, new combinations, Orchidaceae, taxonomy.

Introduction
The genus Calanthe R.Br. has been recognised
as one of the larger orchid genera in which 207
species are currently accepted (Clayton andCribb, 2013). Circumscription of the genus isstable in recent taxonomic treatments andregional ﬂoras (e.g. Chen et al., 2009; Comber,
2001; Nakajima and Ohba, 2011; Wood et al.,2011). However, phylogenetic relationships
within this genus and afﬁnities with allied genera
such as Cephalantheropsis Guillaumin, Gastrorchis Thouars, and Phaius Lour. have remainedunresolved.
Yukawa and Ishida (2008) and Yukawa (2013)
conducted molecular phylogenetic analyses of 53
species of Calanthe that cover major infragenericgroups and representative species of allied genera. These studies found the following sevenclades: 1) Calanthe sections Calanthe, Ghiesbrechtia, Monophylla, and Vernae; 2) Calanthe
sections Styloglossum and Rhodochilus; 3) Cephalantheropsis; 4) Phaius tankervilleae group; 5)
Gastrorchis and Phaius pulchellus; 6) Phaius ﬂavus group; 7) Calanthe subgenus Preptanthe.
Since the results deﬁnitely showed that Calantheand Phaius do not represent monophyletic

groups, current generic classiﬁcation breaks theconsistency of Calanthe and Phaius. Subsequently, Xiang et al. (2014) and Zhai et al.
(2014) analysed relationships in Calanthe andallied groups by using different species and different genetic regions and results of these studies
were consistent with the above-mentioned relationships.
Two alternative scenarios can be proposed torestore the monophyly of Calanthe and Phaius:
one is to redeﬁne the four genera, Calanthe,
Cephalantheropsis, Gastrorchis, and Phaius, as asingle genus. The other is to recircumscribe
Calanthe and Phaius in a narrow sense. In theformer option, the oldest available generic namefor this group is Phaius (Loureiro, 1790). This
treatment necessitates a lot of changes for wellestablished names of Calanthe. It is thus
contended that a broad deﬁnition of Phaius
including Calanthe, Cephalantheropsis, andGastrorchis violates the aim of nomenclatural
stability. Consequently, we adopt the latter option
in which Calanthe and Phaius are narrowly
deﬁned.
In that case, the redeﬁned Calanthe corresponds to the clade comprising sections
Calanthe, Ghiesbrechtia, Monophylla, and Ver-146

Tomohisa Yukawa and Phillip Cribbnae (Clade 1) because the type species of thegenus, C. veratrifolia (now accepted as a synonym of C. triplicata) is included in this clade.
As a result, the two remaining clades of thegenus Calanthe, one comprising sections Styloglossum and Rhodochilus (Clade 2) and the othercorresponding to subgenus Preptanthe (Clade 7),need to be treated as independent genera. Sections Styloglossum and Rhodochilus should be
replaced by genus Styloglossum and subgenus
Preptanthe should be elevated to genus Preptanthe. In summary, the current concept ofCalanthe is necessarily to be subdivided into
three genera, Calanthe, Styloglossum, and Preptanthe from the standpoint of taxonomic consistency. Here we present new combinations inaccordance with this treatment.
Taxonomic Treatment
Preptanthe Rchb.f. in Fl. Serres Jard. Eur. 8:245 (1853). Type: Preptanthe vestita (Lindl.)
Rchb.f.
Calanthe subgenus Preptanthe (Rchb.f.) Schltr.
in Repert. Spec. Nov. Regni Veg. Beih. 1: 377(1912).
Preptanthe cardioglossa (Schltr.) T.Yukawa &P.J.Cribb, comb. nov.
Basionym: Calanthe cardioglossa Schltr. in Repert. Spec. Nov. Regni Veg. Beih. 2: 85 (1906).
Calanthe hosseusiana Kraenzl. in Repert. Spec.
Nov. Regni Veg. Beih. 7: 82 (1909), nom.
superﬂ.
Calanthe cardiochila Kraenzl. in Repert. Spec.
Nov. Regni Veg. Beih. 7: 83 (1909), orth. var.
Preptanthe elmeri (Ames) T.Yukawa &P.J.Cribb, comb. nov.
Basionym: Calanthe elmeri Ames, Orch. 2: 154(1908).
Preptanthe hirsuta (Seidenf.) T.Yukawa &P.J.Cribb, comb. nov.
Basionym: Calanthe hirsuta Seidenf. in Dansk

Bot. Arkiv 29: 39 (1975).
Preptanthe labrosa (Rchb.f.) T.Yukawa &P.J.Cribb, comb. nov.
Basionym: Limatodis labrosa Rchb.f. in Gard.
Chron. n.s. 11: 202 (1879).
Calanthidium labrosum (Rchb.f.) Pﬁtz., Nat.
Pﬂanzenfam. 2, 6: 153 (1889).
Calanthe labrosa (Rchb.f.) Hook.f., Fl. Brit.
India 5: 856 (1890).
Alismorkis labrosa (Rchb.f.) Kuntze, Revis. Gen.
Pl. 2: 650 (1891).
Preptanthe papuana (Schltr.) T.Yukawa &P.J.Cribb, comb. nov.
Basionym: Aulostylis papuana Schltr. in Repert.
Spec. Nov. Regni Veg. Beih. 1: 392 (1912).
Calanthe papuana (Schltr.) J.J.Sm. in Nova
Guinea 14: 359 (1929).
Preptanthe punctata (Kurzweil) T.Yukawa &P.J.Cribb, comb. nov.
Basionym: Calanthe punctata Kurzweil in Gard.
Bull. Singapore 65: 163 (2013).
Preptanthe rosea (Lindl.) T.Yukawa &P.J.Cribb, comb. nov.
Basionym: Limatodis rosea Lindl. in Paxt., Fl.
Gard. 3: 35 (1852).
Calanthe rosea (Lindl.) Benth. in J. Linn. Soc.
Bot. 18: 309 (1880).
Alismorkis rosea (Lindl.) Kuntze, Revis. Gen. Pl.
2: 650 (1891).
Preptanthe succedanea (Gagnep.) T.Yukawa &P.J.Cribb, comb. nov.
Basionym: Calanthe succedanea Gagnep. in
Bull. Mus. Natl. Hist. Nat. Paris 2, 3: 324(1931).
Styloglossum Breda, Gen. Sp. Orchid. Asclep. 2:
t. 7 (1829). Type: Styloglossum nervosum
Breda.
Calanthe section Styloglossum (Breda) J.J.Sm.,
Fl. Buitenzorg 6: 202 (1905).

Nomenclatural Changes in Calanthe

Calanthe section Rhodochilus Schltr., in Repert.
Spec. Nov. Regni Veg. Beih. 1: 376 (1912),
syn. nov.
Styloglossum alboluteum (Ridl.) T.Yukawa &P.J.Cribb, comb. nov.
Basionym: Calanthe albolutea Ridl. in J. Straits
Branch Roy. Asiat. Soc. 39: 80 (1903).
Styloglossum angustifolium (Blume) T.Yukawa& P.J.Cribb, comb. nov.
Basionym: Amblyglottis angustifolia Blume,
Bijdr.: 369 (1825).
Calanthe angustifolia (Blume) Lindl., Gen. Spec.
Orch. Pl.: 251 (1833).
Calanthe phajoides Rchb.f., Xenia Orch. 1: 207(1856).
Alismorkis angustifolia (Blume) Kuntze, Revis.
Gen. Pl. 2: 650 (1891).
Alismorkis phajoides (Rchb.f.) Kuntze, Revis.
Gen. Pl. 2: 650 (1891).
Calanthe angustifolia var. ﬂava Ridl., Fl. Malay
Penins. 4: 120 (1924).
Calanthe angustifolia var. sumatrana Carr in J.
Malayan Branch Roy. Asiat. Soc. 11: 79(1933).
Calanthe pumila Fukuy. in Bot. Mag. Tokyo 49:
438 (1935).
Calanthe striata var. pumila (Fukuy.) S.S.Ying,
Col. Ill. Indig. Orch. Taiwan 1: 430 (1977).
Styloglossum aquamarinum (Schuit. & de
Vogel) T.Yukawa & P.J.Cribb, comb. nov.
Basionym: Calanthe aquamarina Schuit. & de
Vogel in Malesian Orchid J. 4: 102 (2009).
Styloglossum aurantiacum (Ridl.) T.Yukawa &P.J.Cribb, comb. nov.
Basionym: Calanthe aurantiaca Ridl. in J.
Straits Branch Roy. Asiat. Soc. 39: 80 (1903).
Calanthe rajana J.J.Sm. in Mitt. Inst. Allg. Bot.
Hamburg 7: 39 (1927).
Calanthe atjehensis J.J.Sm. in Blumea 5: 308(1943).

147

Styloglossum aureiﬂorum (J.J.Sm.) T.Yukawa& P.J.Cribb, comb. nov.
Basionym: Calanthe aureiﬂora J.J.Sm. in Bull.
Jard. Bot. Buitenzorg 3, 5: 67 (1922).
Styloglossum balansae (Finet) T.Yukawa &P.J.Cribb, comb. nov.
Basionym: Calanthe balansae Finet in Bull. Soc.
Bot. France 46: 435 (1900).
Calanthe saccifera Kraenzl. in Vierteljahrsschr.
Naturf. Ges. Zürich 74: 79 (1929).
Styloglossum carrii (Seidenf. & J.J.Wood ex
Govaerts) T.Yukawa & P.J.Cribb, comb. nov.
Basionym: Calanthe carrii Seidenf. & J.J.Wood
ex Govaerts in World Checklist Seed Pl. 3, 1:
11 (1999).
Calanthe pusilla Carr in Gard. Bull. Straits Settlem. 5: 147 (1930), non Finet (1899).
Styloglossum caulescens (J.J.Sm.) T.Yukawa &P.J.Cribb, comb. nov.
Basionym: Calanthe caulescens J.J.Sm. in Bull.
Jard. Bot. Buitenzorg 2, 2: 2 (1911).
Calanthe apostasioides Schltr. in Repert. Spec.
Nov. Regni Veg. Beih. 1: 377 (1912).
Calanthe salmoniviridis P.Royen, Alp. Fl. New
Guinea 2: 134 (1979).
Styloglossum caulodes (J.J.Sm.) T.Yukawa &P.J.Cribb, comb. nov.
Basionym: Calanthe caulodes J.J.Sm. in Bot.
Jahrb. Syst. 65: 468 (1933).
Styloglossum chevalieri (Gagnep.) T.Yukawa &P.J.Cribb, comb. nov.
Basionym: Calanthe chevalieri Gagnep. in Bull.
Mus. Nat. Hist. Paris 2, 3: 322 (1931).
Styloglossum chrysanthum (Schltr.) T.Yukawa& P.J.Cribb, comb. nov.
Basionym: Calanthe chrysantha Schltr. in K.
Schum. & C.A.G.Lauterbach, Fl. Schutzgeb.
Deutsch Südsee, Nachtr.: 141 (1905).
Calanthe chrysantha var. orbicularis J.J.Sm. in148

Tomohisa Yukawa and Phillip Cribb

Bot. Jahrb. Syst. 66: 178 (1934).

16: 15 (2006).

Styloglossum chrysoleucum (Schltr.) T.Yukawa& P.J.Cribb, comb. nov.
Basionym: Calanthe chrysoleuca Schltr. in Repert. Spec. Nov. Regni Veg. Beih. 1: 386 (1912).

Styloglossum ecallosum (J.J.Sm.) T.Yukawa &P.J.Cribb, comb. nov.
Basionym: Calanthe ecallosa J.J.Sm. in Bull.
Jard. Bot. Buitenzorg 2, 9: 26 (1913).

Styloglossum clavatum (Lindl.) T.Yukawa &P.J.Cribb, comb. nov.
Basionym: Calanthe clavata Lindl., Gen. Sp.
Orch. Pl.: 251 (1833), non Kraenzl. (1898).
Alismorkis clavata (Lindl.) Kuntze, Rev. Gen. Pl.
2: 650 (1891).
Calanthe clavata var. malipoensis Z.H.Tsi in
Acta Phytotax. Sin. 19: 508 (1981).

Styloglossum englishii (Rolfe) T.Yukawa &P.J.Cribb, comb. nov.
Basionym: Calanthe englishii Rolfe in Bull.
Misc. Inf. Kew 1899: 111 (1899).

Styloglossum clavicalcar (J.J.Sm.) T.Yukawa &P.J.Cribb, comb. nov.
Basionym: Calanthe clavicalcar J.J.Sm. in Bull.
Jard. Bot. Buitenzorg 3, 5: 68 (1922).
Styloglossum cleistogamum (Holttum) T.
Yukawa & P.J.Cribb, comb. nov.
Basionym: Calanthe cleistogama Holttum in
Gard. Bull. Sing. 11: 277 (1947).
Styloglossum cruciatum (Schltr.) T.Yukawa &P.J.Cribb, comb. nov.
Basionym: Calanthe cruciata Schltr. in Repert.
Spec. Nov. Regni Veg. Beih. 1: 389 (1912).
Styloglossum densiﬂorum (Lindl.) T.Yukawa &P.J.Cribb, comb. nov.
Basionym: Calanthe densiﬂora Lindl., Gen. Sp.
Orch. Pl.: 250 (1833).
Alismorkis densiﬂora (Lindl.) Kuntze, Rev. Gen.
Pl. 2: 650 (1891).
Calanthe kazuoi Yamam. in Trans. Nat. Hist.
Soc. Formosa 20: 38 (1930).
Phaius epiphyticus Seidenf. in Nord. J. Bot. 5:
159 (1985).
Styloglossum duyanum (Aver.) T.Yukawa &P.J.Cribb, comb. nov.
Basionym: Calanthe duyana Aver. in Rheedea

Styloglossum epiphyticum (Carr) T.Yukawa &P.J.Cribb, comb. nov.
Basionym: Calanthe epiphytica Carr in J.
Malayan Branch Roy. Asiat. Soc.11: 80 (1933).
Styloglossum habbemense (Ormerod &P.J.Cribb) T.Yukawa & P.J.Cribb, comb. nov.
Basionym: Calanthe habbemensis Ormerod &P.J.Cribb in Malesian Orchid. J. 8: 51 (2011).
Styloglossum halconensis (Ames) T.Yukawa &P.J.Cribb, comb. nov.
Basionym: Calanthe halconensis Ames in Phil.
J. Sci. C 2: 324 (1907).
Styloglossum hololeucum (Rchb.f.) T.Yukawa& P.J.Cribb, comb. nov.
Basionym: Calanthe hololeuca Rchb.f. in B.
Seemann, Fl. Vit.: 298 (1868).
Calanthe vaupeliana Kraenzl. in Notizbl. Bot.
Gart. Berlin 5: 111 (1909).
Calanthe neocaledonica Rendle in J. Linn. Soc.
Bot. 45: 251 (1921).
Styloglossum hyacinthinum (Schltr.) T.Yukawa& P.J.Cribb, comb. nov.
Basionym: Calanthe hyacinthina Schltr. in Repert. Spec. Nov. Regni Veg. Beih. 21: 178(1925).
Styloglossum inﬂatum (Schltr.) T.Yukawa &P.J.Cribb, comb. nov.
Basionym: Calanthe inﬂata Schltr. in Repert.

Nomenclatural Changes in CalantheSpec. Nov. Regni Veg. Beih. 1: 390 (1912).
Styloglossum johorense (Holttum) T.Yukawa &P.J.Cribb, comb. nov.
Basionym: Calanthe johorensis Holttum in Gard.
Bull. Sing. 11: 277 (1947).
Styloglossum lancilabre (Ormerod & P.J.Cribb)
T.Yukawa & P.J.Cribb, comb. nov.
Basionym: Calanthe lancilabris Ormerod &P.J.Cribb in Malesian Orchid J. 8: 51 (2011).
Styloglossum leucosceptrum (Schltr.) T.Yukawa& P.J.Cribb, comb. nov.
Basionym: Calanthe leucosceptrum Schltr. in
Repert. Spec. Nov. Regni Veg. Beih. 1: 388(1912).
Calanthe labellicauda Gilli in Ann. Naturhist.
Mus. Wien B 84: 18 (1980, publ. 1983).
Styloglossum leucosceptrum (Schltr.) T.Yukawa& P.J.Cribb var. bisubuliferum (J.J.Sm.) T.
Yukawa & P.J.Cribb, comb. nov.
Basionym: Calanthe leucosceptrum Schltr. var.
bisubulifera J.J.Sm. in Nova Guinea 14: 358(1929).
Styloglossum longifolium (Schltr.) T.Yukawa &P.J.Cribb, comb. nov.
Basionym: Calanthe longifolia Schltr. in Repert.
Spec. Nov. Regni Veg. Beih. 1: 387 (1912).
Styloglossum lyroglossum (Rchb.f.) T.Yukawa& P.J.Cribb, comb. nov.
Basionym: Calanthe lyroglossa Rchb.f., Otia
Bot. Hamburg. 1: 53 (1878).
Calanthe foerstermannii Rchb.f. in Gard. Chron.
n.s. 19: 814 (1883).
Alismorkis foerstermannii (Rchb.f.) Kuntze, Rev.
Gen. Pl. 2: 650 (1891).
Alismorkis lyroglossa (Rchb.f.) Kuntze, Rev.
Gen. Pl. 2: 650 (1891).
Calanthe forsythiiﬂora Hayata, Icon. Pl. Formos.
4: 67 (1914).
Calanthe liukiuensis Schltr. in Repert. Spec. Nov.

149

Regni Veg. Beih. 4: 67 (1919).
Calanthe nephroidea Gagnep. in Bull. Mus. Natl.
Hist. Nat., 2, 22: 625 (1951).
Calanthe lyroglossa var. forsythiiﬂora (Hayata)
S.S.Ying, Col. Ill. Indig. Orch. Taiwan 1: 88(1977).
Styloglossum lyroglossum (Rchb.f.) T.Yukawa& P.J.Cribb var. longibracteatum (OʼByrne)
T.Yukawa & P.J.Cribb, comb. nov.
Basionym: Calanthe lyroglossa var. longibracteata OʼByrne in Malayan Orchid Rev. 31: 29(1997).
Styloglossum lyroglossum (Rchb.f.) T.Yukawa& P.J.Cribb var. actinomorphum (Fukuy.) T.
Yukawa & P.J.Cribb, comb. nov.
Basionym: Calanthe actinomorpha Fukuy. in
Bot. Mag. Tokyo 49 : 668 (1935).
Phaius actinomorphus (Fukuy.) T.P.Lin, Nat.
Orch. Taiwan: 224 (1975).
Calanthe lyroglossa var. actinomorpha (Fukuy.)
P.J.Cribb & D.A.Clayton, The Genus Calanthe:
85 (2013).
Styloglossum maquilingense (Ames) T.Yukawa& P.J.Cribb, comb. nov.
Basionym: Calanthe maquilingensis Ames, Orch.
5: 97 (1915).
Styloglossum microglossum (Ridl.) T.Yukawa& P.J.Cribb, comb. nov.
Basionym: Calanthe microglossa Ridl. in J.
Straits Branch Roy. Asiat. Soc. 39: 81 (1903).
Styloglossum musa-amanii (J.J.Wood &C.L.Chan) T.Yukawa & P.J.Cribb, comb. nov.
Basionym: Calanthe musa-amanii J.J.Wood &C.L.Chan in J.J.Wood et al., Orch. Mount
Kinabalu 2: 162 (2011).
Styloglossum neohibernicum (Schltr.) T.Yukawa& P.J.Cribb, comb. nov.
Basionym: Calanthe neohibernica Schltr. in
K.M.Schumann & C.A.G.Lauterbach, Fl.

150

Tomohisa Yukawa and Phillip Cribb

Schutzgeb. Deutsch Südsee, Nachtr.: 142 (1905).
Styloglossum nicolae (OʼByrne) T.Yukawa &P.J.Cribb, comb. nov.
Basionym: Calanthe nicolae OʼByrne in
Malayan Orchid Rev. 30: 69 (1996).
Styloglossum oreadum (Rendle) T.Yukawa &P.J.Cribb, comb. nov.
Basionym: Calanthe oreadum Rendle in J. Linn.
Soc. Bot. 45: 250 (1921).
Styloglossum parvilabre (Schltr.) T.Yukawa &P.J.Cribb, comb. nov.
Basionym: Calanthe parvilabris Schltr. in
K.M.Schumann & C.A.G.Lauterbach, Fl.
Schutzgeb. Deutsch Südsee, Nachtr.: 143 (1905).
Styloglossum pulchrum (Blume) T.Yukawa &P.J.Cribb, comb. nov.
Basionym: Amblyglottis pulchra Blume, Bijdr.:
371 (1825).
Styloglossum nervosum Breda, Gen. Sp. Orchid.
Asclep. 2: t. 7 (1829).
Calanthe pulchra (Blume) Lindl., Gen. Sp. Orch.
Pl.: 250 (1833).
Calanthe curculigoides Lindl., Gen. Sp. Orch.
Pl.: 251 (1833).
Alismorkis pulchra (Blume) Kuntze, Rev. Gen.
Pl. 2: 650 (1891).
Alismorkis curculigoides (Lindl.) Kuntze, Rev.
Gen. Pl. 2: 650 (1891).
Calanthe shelfordii Ridl. in J. Straits BranchRoy. Asiat. Soc. 33: 23 (1900).
Styloglossum rhodochilum (Schltr.) T.Yukawa& P.J.Cribb, comb. nov.
Basionym: Calanthe rhodochila Schltr. in
K.M.Schumann & C.A.G.Lauterbach, Fl.
Schutzgeb. Deutsch Südsee, Nachtr.: 143 (1905).
Calanthe breviscapa J.J.Sm. in Bull. Jard. Bot.
Buitenzorg 2, 2: 1 (1911).
Calanthe bracteosa Schltr. in Bot. Jahrb. Syst.
58: 128 (1923), non Rchb.f. (1882).

Styloglossum rhodochilum (Schltr.) T.Yukawa& P.J.Cribb var. reconditiﬂorum (J.J.Sm.) T.
Yukawa & P.J.Cribb, comb. nov.
Basionym: Calanthe reconditiﬂora J.J.Sm. in
Bull. Jard. Bot. Buitenzorg 3, 10: 108 (1928).
Calanthe manis J.J.Sm., Nova Guinea 14: 356(1929).
Calanthe rhodochila var. reconditiﬂora (J.J.Sm.)
S.Thomas in Blumea 44: 29 (1999).
Styloglossum rigidum (Carr) T.Yukawa &P.J.Cribb, comb. nov.
Basionym: Calanthe rigida Carr in Gard. Bull.
Straits Settlem. 5: 145 (1930).
Calanthe rigida Carr var. gracilis Carr in J.
Malayan Branch Roy. Asiat. Soc.11: 81 (1933).
Styloglossum ruttenii (J.J.Sm.) T.Yukawa &P.J.Cribb, comb. nov.
Basionym: Calanthe ruttenii J.J.Sm. in Bull.
Jard. Bot. Buitenzorg 3, 10: 109 (1928).
Styloglossum saccatum (J.J.Sm.) T.Yukawa &P.J.Cribb, comb. nov.
Basionym: Calanthe saccata J.J.Sm. in Bull.
Inst. Bot. Buitenzorg 7: 3 (1900).
Styloglossum salaccense (J.J.Sm.) T.Yukawa &P.J.Cribb, comb. nov.
Basionym: Calanthe salaccensis J.J.Sm. in Bull.
Dép. Agric. Indes Néerl. 43: 21 (1910).
Styloglossum speciosum (Blume) T.Yukawa &P.J.Cribb, comb. nov.
Basionym: Amblyglottis speciosa Blume, Bijdr.:
371 (1825).
Calanthe speciosa (Blume) Lindl., Gen. Sp.
Orch. Pl.: 250 (1833).
Calanthe scortechinii Hook.f., Fl. Brit. India 5:854 (1890).
Alismorkis speciosa (Blume) Kuntze, Rev. Gen.
Pl. 2 : 650 (1891).
Calanthe formosana Rolfe in Ann. Bot. London
9: 157 (1895).
Calanthe yushunii K.Mori & Yamam. in J. Soc.

Nomenclatural Changes in Calanthe

Trop. Agric. Taiwan 4: 188 (1932).
Calanthe patsinensis S.Y.Hu in Chung Chi J. 11:
13 (1972).
Calanthe disticha Tang & F.T.Wang in Acta Phytotax. Sin. 12: 43 (1974).
Calanthe pulchra var. formosana (Rolfe)
S.S.Ying, Col. Illust. Indig. Orch. Taiwan 1:
430 (1977).
Styloglossum stella (P.J. Cribb) T.Yukawa &P.J.Cribb, comb. nov.
Basionym: Calanthe stella P.J.Cribb in MalesianOrchid J. 4: 13 (2009).
Styloglossum stenophyllum (Schltr.) T.Yukawa& P.J.Cribb, comb. nov.
Basionym: Calanthe stenophylla Schltr. in Repert. Spec. Nov. Regni Veg. Beih. 1: 388 (1912).
Styloglossum truncicolum (Schltr.) T.Yukawa& P.J.Cribb, comb. nov.
Basionym: Calanthe truncicola Schltr. in Beibl.
Bot. Jahrb. Syst. 104: 26 (1911).
Styloglossum tunense (J.J.Sm.) T.Yukawa &P.J.Cribb, comb. nov.
Basionym: Calanthe tunensis J.J.Sm. in Bull.
Inst. Bot. Buitenzorg 7: 3 (1900).
Calanthe ﬂabelliformis R.S.Rogers in Trans. &Proc. Roy. Soc. S. Austr. 49: 259 (1925).
Styloglossum ventilabrum (Rchb.f.) T.Yukawa& P.J.Cribb, comb. nov.
Basionym: Calanthe ventilabrum Rchb.f. in B.
Seemann, Fl. Vit.: 298 (1868).
Calanthe langei F.Muell., S. Sci. Rec. n.s., 1885:
1 (1885).
Calanthe bigibba Schltr. in Repert. Spec. Nov.
Regni Veg. Beih. 9: 99 (1911).
Styloglossum versteegii (J.J.Sm.) T.Yukawa &P.J.Cribb, comb. nov.
Basionym: Calanthe versteegii J.J.Sm. in Bull.
Jard. Bot. Buitenzorg 2, 13: 55 (1914).
Guilianettia disticha Ridl. in Trans. Linn. Soc.

151

London, Bot. 9: 201 (1916).
Calanthe arundinoides J.J.Sm. in Nova Guinea
14: 356 (1929).
Calanthe aurantimacula P.Royen, Alp. Fl. New
Guinea 2: 137 (1979).
Acknowledgments
This study is partly supported by a Grant-inAid to Scientiﬁc Research from the Japan Society for Promotion of Science (No. 24370040) for
T.Yukawa.
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