Bull. Natl. Mus. Nat. Sci., Ser. B, 34(1), pp. 23–29, March 21, 2008

Type Collection in the Herbarium of National Museum of Nature andScience (TNS). Salicales, Fagales, Urticales, Proteales, Santalales,
Polygonales and Chenopodiales
Atsushi Ebihara and Kyoko Miyashita
Department of Botany, National Museum of Nature and Science, Amakubo 4–1–1,
Tsukuba 305–0005, Japan
E-mail: ebihara@kahaku.go.jp

Abstract Type specimens of seven angiosperm orders deposited in the herbarium of National
Museum of Nature and Science (TNS) were explored and reexamined. In total, 85 sheets are confirmed as type collections for 56 taxa including 24 holotypes. The 85 sheets consist of seven Salicaceae, five Betulaceae, seven Fagaceae, two Ulmaceae, three Moraceae, three Urticaceae, 37
Polygonaceae, 16 Caryophyllaceae, two Amaranthaceae and one each of Proteaceae, Balanophoraceae and Chenopodiaceae.
Key words : TNS-herbarium, type collection.

SALICACEAE F. 7
Salix L.
Salix kamikotica Kimura
in Sci. Rep. Tohoku Imp. Univ., Ser. 4, Biol. 6:
194 (1931).
Jap. Name. Kinuyanagi.
Japan, Honshu, Prov. Shinano, Kamikochi,
1930-5-16, A. Kimura 782 [TNS-758703,
785704Isosyntypes].
Salix kassanogluensis Kotschy ex Anderss.
in Prodr. (DC.) 16(2): 212 (1868).
Turkey, montibus Kassan Oghlu, 1859-5-13, T.
Kotschy 64 [TNS-VS-239616 Isotype].
Salix nakamurae Koidz.
in Bot. Mag. (Tokyo) 27: 96 (1913).
Jap. Name. Renge-iwayanagi.
Japan, Prov. Shinano, Dailengezan, 1908-8-12,
M. Nakamura s.n. [TNS-VS-380883Isotype].
Salix paludicola Koidz.
in Bot. Mag. (Tokyo) 33: 219 (1919).
Jap. Name. Miyama-yachiyanagi.
Japan, Yeso, Prov. Ishikari, Mt. Chiubetudake,
1917-7-31, H. Koidzumi 39 (Herb. H. Koidzumi 66685) [TNS-VS-416390 Isotype].
Salix sichotensis Kharkevich et I.B.Vyshin.

in Bot. Zhurn. (Moscow & Leningrad) 70:
1120 (1985).
Russia, Khabarovskiy territory, North SeikhoteAlin, Nanayskiy district, Anyouy river, mountain Tardokyyany, 1983-7-22, S. Kharkevich,
T. Buch & I. Vyshin “Plantae Vasculares Orientis Extremi Rossici (VLA) Flora Exsiccata”
No. 345 [TNS-VS-685590 Isotype].
Salix tagawana Koidz.
in Acta Phytotax. Geobot. 9: 75 (1940).
Taiwan, Prov. Karenkô, Tamazato-gun,
between Sasarabi and Miyasan, 1940-4-8, M.
Tagawa 3646 [TNS-VS-155150 Isosyntype].
Taiwan, Prov. Karenkô, Tamazato-gun,
between Miyasan and Daisuikutsu, 1940-4-9,
M. Tagawa 3647 [TNS-VS-155148 Isosyntype].
BETULACEAE F. 8
Alnus Mill.
Alnus hakkodensis Hayashi
in Bull. Gov. For. Exp. Stat. 57: 153 (1952).
Jap. Name. Sarukura-hannoki.
Japan, Hondo, Prov. Mutsu, Mt. Hakkoda,
Sarukura, 1951-8-7, Y. Hayashi s.n. [TNS-VS-24

Atsushi Ebihara and Kyoko Miyashita

255581Holotype].
Note. According to the original descriptions,
Yasaka Hayashi (1911–1991)’s holotypes aredeposited in the herbarium of Government
Forest Experiment Station, and thus previous
publications of the TNS type collection catalogues treated his types as isotypes. But thecollections in TNS match perfectly the photographs of the specimens cited in their original descriptions. Therefore, it is highly likelythat most of his holotypes had been transferred to TNS in the past.
Betula L.
Betula costata Trautv. var. cuneifolia Kitag.
in J. Jap. Bot. 44: 183 (1969).
Jap. Name. Nagaba-chosen-minebari.
China, Manshuria, Prov. Hei-lung-ching, Mt.
Hsiao-hsing-an-ling, ?-?-?, M. Takahashi s.n.
[TNS-VS-931994 Holotype].
Carpinus L.
Carpinus erosa Blume var. microcarpa Hayashi
in Bull. Gov. For. Exp. Stat. 57: 154 (1952).
Jap. Name. Hime-sawashiba.
Japan, Ezo, Prov. Hiyama, Mt. Gamushi, 19528-8, Y. Hayashi s.n. [TNS-VS-255878 Holotype].
Carpinus erosa Blume var. velutina Hayashi
in Bull. Gov. For. Exp. Stat. 77: 26 (1955).
Jap. Name. Birodo-sawashiba.
Japan, Hondo, Prov. Rikuchiu, Shimohei-gun,
Yamagata[-mura] in Kuji, 1952-8-15, Y.
Hayashi s.n. [TNS-VS-255582 Holotype].
Carpinus tschonoskii Maxim. f. pendula
Hayashiin Bull. Gov. For. Exp. Stat. 125: 72 (1960).
Jap. Name. Shidare-inushide.
Japan, Hondo, Prov. Musashi, Kokubunji,
1958-5-10, Y. Hayashi s.n. [TNS-VS255855Holotype].

FAGACEAE F. 9
Cyclobalanopsis Oerst.
Cyclobalanopsis acuta (Thunb.) Oerst. var.
megaphylla Hayashi
in J. Jap. Bot. 29: 149 (1954).
Jap. Name. O-akagashi.
Japan, Hondo, Prov. Musashi, Meguro[-ku],
Ohshima Shrine, 1953-6-20, Y. Hayashi s.n.
[TNS-VS-255872 Holotype].
Fagus L.
Fagus japonica Maxim. var. pleiosperma
Hayashiin Bull. Gov. For. Exp. Stat. 107: 27 (1958).
Jap. Name. Hako-inubuna.
Japan, Hondo, Prov. Sagami, Mt. Tanzawa,
1956-7-15, Y. Hayashi s.n. [TNS-VS-254528
Holotype].
Pasania Oerst.
Pasania glabra (Thunb.) Oerst. f. microphylla
Hayashiin J. Geobot. 11: 117 (1963).
Jap. Name. Koba-no-shiribuka-gashi.
Japan, Hondo, Prov. Suo, Iwakuni, Mt. Shiro,
1962-11-14, Y. Hayashi s.n. [TNS-VS255875Holotype].
Quercus L.
Quercus grosseserrata Blume var. macrocarpa
Nakai
in Bull. Natl. Sci. Mus., Tokyo 27: 29 (1949).
Jap. Name. Nagaba-mizunara.
Japan, Hondo, Prov. Uzen, tractus NisiTagawa, oppidi Nedzugaseki, Wasada, 194810-21, T. Nakai s.n. [TNS-VS-79743,
79750Holotypes in 2 sheets].
Quercus mongolica Fisch. ex Ledeb. var. grosseserrata (Blume) Rehder et E.H.Wilson f.
laciniata Hayashi
in Bull. Gov. For. Exp. Stat. 107: 27 (1958).
Jap. Name. Hagoromo-mizunara.
Japan, Hondo, Prov. Uzen, Nishimurayamagun, Nishikawa-cho, 1956-8-31, T. Mizuta s.n.

TNS type collections[TNS-VS-256395 Holotype].
Quercus takaoyamensis Makino
in J. Jap. Bot. 2: 13 (1920).
Jap. Name. O-tsukubanegashi.
Japan, Prov. Musashi, Mt. Takao, 1920-6-30,
K. Hisauchi s.n. [TNS-VS-35910 Isotype].
ULMACEAE F. 11
Celtis L.
Celtis hashimotoi Koidz.
in Acta Phytotax. Geobot. 1: 19 (1932).
Jap. Name. Kansai-enoki.
Japan, Honto, Prov. Ohmi, Higashi-asai-gun,Higashi-Kusano-mura, Kami-itanami, Yasakajinja, 1930-10-26, C. Hashimoto [TNS-VS51209Isotype].
Ulmus L.
Ulmus davidiana Planch. f. glabra Kitag.
in J. Jap. Bot. 47: 354 (1972).
Jap. Name. Kenashi-tonire.
China, Shan-tung, Chi-nan, Mt. Ch’ien-foshan, 1940-4-28, T. Shinagawa 6 [TNS-VS76875Holotype].

25

URTICACEAE F. 14
Boehmeria Jacq.
Boehmeria maximowiczii Nakai et Satakein J. Fac. Sci. Univ. Tokyo, Sect. 3, Bot. 4: 522(1936).
Jap. Name. O-meyabumao.
Japan, Prov. Nagato, prope Hagi, Kauchi,
1926-10-4, J. Nikai 2970 [TNS-VS-48919 
Isotype].
Laportea Gaudich.
Laportea saipanensis Kaneh.
in Bot. Mag. (Tokyo) 45: 277 (1931).
Mariana Islands, Saipan, 1930-7-?, R. Kanehira
973 [TNS-VS-9525 Isosyntype].
Pilea Lindl.
Pilea amamiana Ohwi
in Bull. Natl. Sci. Mus., Tokyo n.s. 1: 6 (1954).
Jap. Name. Yuwan-mizu.
Japan, Ryukyu, Ins. Amamioshima, Mt. Yuwandake, 1951-5-20, T. Amano 6465 [TNS-VS87299Holotype].
PROTEACEAE F. 15

MORACEAE F. 13
Ficus L.
Ficus erecta Thunb. f. rhodovenia Nakai
in Jap. J. Bot. 25: 131 (1950).
Jap. Name. Akajiku-inubiwa.
Japan, Prov. Ohsumi, Izashiki, 1948-4-16, T.
Nakai & N. Maruyama s.n. [TNS-VS-81822 
Holotype].
Morus L.
Morus amamiana Nakai
in J. Jap. Bot. 25: 131 (1950).
Jap. Name. Amami-guwa.
Japan, Prov. Ohsumi, Sata, Izashiki (cult.),transplanted from Amami-Ohshima, 1948-416, T. Nakai & N. Maruyama s.n. [TNS-VS81824, 81825Syntypes].

Helicia Lour.
Helicia henryi Diels
in Repert. Spec. Nov. Regni Veg. 13: 528 (1915).
China, Yunnan, Szemao, ?-?-?, A. Henry
11910A [TNS-VS-193248 Isotype].
BALANOPHORACEAE F. 23
Balanophora J.R. Forst. et G. Forst.
Balanophora kiusiana Ohwi
in Fl. Jap. Rev. ed. (Ohwi) [Japanese ed.]1437 (1965).
Jap. Name. Kyushu-tsuchitorimochi.
Japan, Kyushu, Prov. Bungo, Mt. Yuhudake,
1964-8-?, M. Endo s.n. [TNS-VS-216475 
Holotype].

26

Atsushi Ebihara and Kyoko Miyashita

CHENOPODIACEAE F. 36
Atriplex L.
Atriplex franktonii Tascher.
in Can. J. Bot. 50: 1586 (1972).
Canada, Nova Scotia, Pictou county, north sideof Pictou Harbour, 1968-8-29, P. M. Taschereau 272 [TNS-VS-442246 Isosyntype].
POLYGONACEAE F. 25
Bistorta (L.) Adans.
Bistorta albiflora Miyam. et H. Ohba
in Bull. Natl. Sci. Mus., Tokyo, Ser. B 28: 141(2002).
China, Sichuan, Daocheng, around Gongga
Shan, 1997-8-6, S. Wu, H. Ikeda, S. Akiyama,
F. Miyamoto & W. Chen 459 [TNS-VS717846Isotype].
Bistorta attenuatifolia Miyam. et H. Ohba
in Bull. Natl. Sci. Mus., Tokyo, Ser. B 30: 109(2004).
China, Xizang (Tibet), Baxoi Xian, Baxoi
Xian, Rawu–Baxoi, 2000-8-4, S. Akiyama,
M. Wakabayasi, F. Miyamoto, H. Ikeda, S.K.
Wu, X. Chen, L. Gao, J. Yue & Y. Zhen
105176 [TNS-VS-730258 Isotype]. China,
Xizang (Tibet), Zogang Xian, Langla
Shan–Nianla Shan, 2000-7-19, S. Akiyama,
M. Wakabayasi, F. Miyamoto, H. Ikeda, S.K.
Wu, X. Chen, L. Gao, J. Yue & Y. Zhen
105057 [TNS-VS-730259 Paratype].
Bistorta longispicata Yonek. et H. Ohashi
in J. Jap. Bot. 76: 346 (2001).
China, Tibet, Tsa Tang, between Xigaze andLhasa, W. of Qüxü, 1914-8-1, E. Kawaguchi
62 [TNS-VS-104058 Paratype]. China,
Tibet, vicinity of Lhasa, Elba Temple, 1914-823, E. Kawaguchi 138 [TNS-VS-104053 
Paratype]; E. Kawaguchi 146 [TNS-VS104049Paratype]; E. Kawaguchi 148 [TNSVS-104051 Paratype]; E. Kawaguchi 152[TNS-VS-104070 Paratype]; E. Kawaguchi
156 [TNS-VS-104054 Paratype]. China,
Tibet, N. of Lhasa, Mt. Penpo Kola, 1914-910, E. Kawaguchi 267 [TNS-VS-104076 
Paratype]. China, Tibet, 14–15 miles NE. of

Lhasa, Mt. Chomo Sese, 1914-9-6, E.
Kawaguchi 244 [TNS-VS-104071Paratype];
E. Kawaguchi 245 [TNS-VS-104074 
Paratype]; E. Kawaguchi s.n. [TNS-VS104059Paratype]. China, Tibet, 14–15
miles S. of Lhasa, De la, 1914-9-16, E.
Kawaguchi 321 [TNS-VS-104062 Paratype];
E. Kawaguchi s.n. [TNS-VS-104056 
Paratype]; E. Kawaguchi 340 [TNS-VS104057Paratype].
Bistorta pacifica (Petrov ex Kom.) Kom. f.
velutina Kitag.
in J. Jap. Bot. 47: 354 (1972).
China, Manshuria, Yen-chi, Yanji, 1939-7-20,
K. Yoshimizu s.n. [TNS-VS-77557 Holotype]. China, Manshuria, Prov. Chi-lin, circa
T’u-mén-ling, 1932-6-1, K. Yamatsuta 1840
[TNS-VS-75704Paratype]. Korea, Prov.
Kan-hoku, Yu-hei, 1930-6-19, J. Ohwi 1464
[TNS-VS-219602 Paratype].
Bistorta tubistipulis Miyam. et H. Ohba
in Bull. Natl. Sci. Mus., Tokyo, Ser. B 28: 143(2002).
China, Sichuan, Bowa Shan (near Daocheng),
1996-8-16, S. Wu, H. Ikeda, M. Wakabayashi,
F. Miyamoto, Y. Yang & T. Kikuchi 1499
[TNS-VS-717845 Isotype].
Persicaria (L.) Mill.
Persicaria musashinoensis Hiyama
in J. Jap. Bot. 32: 190 (1957).
Jap. Name. Inu-hanatade.
Japan, Hondo, Prov. Musashi, Tokyo [Pref.],
Nerima-ku, Egoda, 1956-10-4, K. Hiyama s.n.
[TNS-VS-133475 Holotype].
Persicaria ramosa Nakai
in Bot. Mag. (Tokyo) 44: 518 (1930).
Jap. Name. Hai-yanonegusa.
Japan, Prov. Musashi, Itsuka-ichi, Kariyose,
1929-8-24, K. Hisauchi s.n. [TNS-VS-35907
and 35908Isoparatypes].
Persicaria satsumensis Nakai
in J. Jap. Bot. 25: 129 (1950).
Jap. Name. Satsuma-tade.
Japan, Kjusju, Prov. Satsuma, Ibusuki, 1948-4-

TNS type collections

15, T. Nakai & N. Maruyama s.n. [TNS-VS81806Possible type].
Note. The collection date is different from theoriginal description (1948-4-14 in the description).
Persicaria sterilis Nakai f. leucantha Nakai
in J. Jap. Bot. 25: 130 (1950).
Jap. Name. Shirobana-hime-sakuratade.
Japan, Hondo, Prov. Bingo, Kurashiki[-shi],
1943-10-26, Y. Ikegami 3021 [TNS-VS79749Holotype].
Polygonum L.
Polygonum auriculatum Makino
in Bot. Mag. (Tokyo) 17: 117 (1903).
Jap. Name. Hosoba-no-unagitsukami.
Japan, Prov. Tosa, Asakura[-mura], 1892-1015, T. Makino s.n. [TNS-VS-5128 Isosyntype].
Polygonum breviochreatum Makino
in Bot. Mag. (Tokyo) 17: 117 (1903).
Jap. Name. Nagaba-no-yanonegusa.
Japan, Prov. Tosa, Hage-mura, 1892-10-?, T.
Makino s.n. [TNS-VS-5144Possible syntype].
Note. The type locality is “Idzuma, Tosa” inthe original description.
Polygonum insulare M.I. Maximova
in Novosti Sist. Vyssh. Rast. 15: 142 (1979).
Russia, Jacutia, Insulae Ursinae maris Sibirisi
Orientaris, insula Krestovskij, 1971-7-17, M.
Maximova [TNS-VS-442103 Isotype].
Polygonum paludicola Makino var. nikaii
Makinoin Bot. Mag. (Tokyo) 28: 114 (1914).
Jap. Name. Saikoku-nukabo.
Japan, Prov. Bitchù, Kibi-gôri, Makane-mura,
Itakura, 1902-10-26, J. Nikai 939 [TNS-VS48757Isolectotype].
Polygonum pteranthum Kitag.
in J. Geobot. 20: 32 (1972).
Jap. Name. Tsukushi-michiyanagi.
Japan, Kyusyu, Prov. Ohsumi, Isl. Tanegashima,
Annô, 1954-12-21, S. Ouchiyama 69 [TNSVS-130070 Holotype].

27

Polygonum ramuliflorum Kitag.
in J. Jap. Bot. 39: 357 (1964).
Jap. Name. Nukabo-michiyanagi.
Japan, Prov. Oomi, Gamô[-gun], Kamakakemura, 1951-8-24, C. Hashimoto 14895 [TNSVS-103026 Holotype].
Polygonum ryukyuense Kitag.
in J. Geobot. 20: 34 (1972).
Jap. Name. Okinawa-michiyanagi.
Japan, Ryukyu, Prov. Okinawa, Isl. Okinawa,
prope Nawa [=Naha-shi], Mt. Ohno-yama,
1967-9-2, E. Takamine 2804 [TNS-VS215336Holotype].
Polygonum yamatutae Kitag.
in J. Jap. Bot. 39: 358 (1964).
Jap. Name. Nagaba-michiyanagi.
China, Manshuria, Prov. Fêng-t’ien, circa Peiling, 1924-7-10, K. Yamatuta 1832 [TNS-VS75752 Holotype].
Reynoutria Houtt.
Reynoutria sachalinensis (F. Schmidt) Nakai
var. brachyphylla Honda
in Bot. Mag. (Tokyo) 50: 389 (1936).
Jap. Name. Haruna-itadori.
Japan, Honshū, Prov. Kôzuke, monte Haruna,
1935-6-12, T. Sakai s.n. [TNS-VS-51185 
Isotype].
Rumex L.
Rumex daiwoo Makino
in Bot. Mag. (Tokyo) 15: 124 (1901).
Jap. Name. Madaio.
Japan, Prov. Tosa, Sakawa-mura, Nanokawa,
1893-6-?, T. Makino s.n. [TNS-VS-5295 
Syntype]. Japan, Prov. Tosa, Nanokawa, 18885-17, K. Watanabe s.n. [TNS-VS-54971Possible syntype].
Note. The collection date of TNS-VS-54971 isdifferent from the original description (18895-18).
Rumex ursinus M.I. Maximova
in Novosti Sist. Vyssh. Rast. 15: 113 (1979).
Russia, Insulae Ursinae maris Sibirici Orientaris, insula Leontjevii, 1971-8-24, M. Maxi-

28

Atsushi Ebihara and Kyoko Miyashita

mova s.n. [TNS-VS-442102 Isotype].
CARYOPHYLLACEAE F. 34
Brachystemma D. Don
Brachystemma ovatifolium M. Mizus.
in Acta Phytotax. Geobot. 16: 42 (1955).
Nepal, Gumrung to Sarti, 1953-7-26, S. Nakaos.n. [TNS-VS-263814 Isotype].
Cerastium L.
Cerastium calcicola Ohwi
in Repert. Spec. Nov. Regni Veg. 36: 46 (1934).
Taiwan, Karenko, prope Kiraikei, Nokogoe,
1933-6-?, J. Ohwi 2955 [TNS-VS-222195b 
Isotype].
Dianthus L.
Dianthus japonicus Thunb. f. albiflorus Konta
et S. Matsumotoin Bull. Natl. Sci. Mus., Tokyo, Ser. B 31: 23(2005).
Jap. Name. Shirobana-hama-nadeshiko.
Japan, Honshu, Shizuoka [Pref.], Shimoda
City, Suzaki Peninsula, Suzaki, 2004-6-24, F.
Konta, S. Matsumoto and K. Hashiba 22847-a[TNS-VS-744461 Holotype];
22847-b[TNS-VS-744460 Isotype]; 22847-c [TNSVS-744459 Isotype].
Melandrium Rohl.
Melandrium nubigenum Ohwi
in Repert. Spec. Nov. Regni Veg. 36: 48 (1934).
Taiwan, Mt. Nankotaisan, 1933-7-20, J. Ohwi
4159 [TNS-VS-222205a, 222205b Isotypes].
Pseudostellaria Pax
Pseudostellaria musashiensis Hiyama
in J. Jap. Bot. 26: 287 (1951).
Jap. Name. Musashi-wachigaiso.
Japan, Hondo, Prov. Musashi, Kiyose[-mura],
1950-4-11, K. Hiyama s.n. [TNS-VS110129Syntype]; same locality, 1950-5-6, F.
Tuda [TNS-VS-90299 Syntype].

Silene L.
Silene andersonii Clokey
in Bull. S. Calif. Acad. Sci. 38: 2 (1939).
U.S.A., Nevada, Clark Co., Charleston Mts.,
Level Canyon, 1937-8-9, I. W. Clokey 7514
[TNS-VS-241109 Isotype].
Silene tokachiensis Kadota
in Mem. Natl. Sci. Mus., Tokyo 27: 64 (1994).
Jap. Name. Tokachi-biranji.
Japan, Tokachi-shicho, Hokkaido, Kato-gun,Kamisihoro-cho, the Ishikari Range, Mt.
Nipesotsuyama, 1985-8-4, Y. Kadota & K.
Midorikawa 12730 [TNS-VS-9018827 
Holotype; TNS-VS-9018829 Isotype].
Stellaria L.
Stellaria iinumae Ohwi
in Acta Phytotax. Geobot. 6: 145 (1937).
Jap. Name. Ao-hakobe.
Japan, Hondo, Prov. Yamashiro, Kibune, 19376-9, J. Ohwi 9204 [TNS-VS-222398a,
222398bIsotypes]; same locality, 1937-6-9, J.
Ohwi [TNS-VS-57217Isotype]. Japan, Prov.
Tosa, Tosayama-mura, Kashiyama-toge, 19376-14, T. Yoshinaga 9196 [TNS-VS-222397
Isoparatype].
AMARANTHACEAE F. 37
Achyranthes L.
Achyranthes fauriei H. Lév. et Vaniot f.
rotundifolia Ohwi
in Fl. Jap. Rev. ed. (Ohwi) [Japanese ed.] 562,
1437 (1965).
Jap. Name. Maruba-inokozuchi.
Japan, Honshu, Prov. Iwashiro, Hobara-machi,
1963-10-1, T. Oishi [TNS-VS-163350 Holotype; TNS-VS-235330 Isotype].
Following specimens have not yet found in theherbarium, though they are cited as holotype
and syntypes in the original description.
Bistorta suffulta (Maxim.) H. Gross f. pubescens Hiyama
in J. Jap. Bot. 17: 318 (1941).
Jap. Name. Ke-kurin-yukifude.

TNS type collections

Japan, Prov. Kai, Mt. Mitsu-toge, 1934-6-10,
K. Hiyama s.n. [TNS-VS-?Syntype]. Japan,
Prov. Musashi, Mt. Kuromori, 1935-7-15, K.
Hiyama s.n. [TNS-VS-?Syntype].
Tovara filiformis (Thunb.) Nakai f. albiflora
Hiyama29

in J. Jap. Bot. 17: 319 (1941).
Jap. Name. Gin-mizuhiki.
Japan, Prov. Prov. Musashi, Agano-mura,
1928-9-25, K. Hiyama s.n. [TNS-VS-? 
Holotype].