ISSN 2187-0403

Vol. 5

Mikurensis
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a_AELǍƱǺɁ(ƿȮ:¥ĸ7#®&ɎÏĐa_AELǁ&Ȳ!ǳĘ(Ǆȁ:»ȎȖ(ÆǨ&§ɎĥÏĐa_AELǁ(Č(ȃő&ȕ7#:ŝĤɏĨǸĐƮa_AELǍƿȮ Ĩ Ǹ Đ Ʈ a _ A E L Ǎ Ʊ Ǻ & Ȳ  ! ) ɎWatanabeɌ 1961ɍ& 4 7 23 ǎɌ ¯ ɐ ǎ )Ï þ ƺ Ü ǎ ɍ( Ȇ Ȯř 1 Ê  1 ( # į 9 8 7 ɏ ( ĥ ɎWatanabeɌ 1962ɍɎƛ ư ~ƒ ȞɌ 1969ɍɎƒ Ȟ~ȯ Ȑ ĒɌ 1983ɍɎWatanabeɌ 2000ɍɎNomura & KomatsuɌ 2012ɍɎ
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& 4 7 ŏ Š Ʈ a _ A E L Ǎ Ǟ ƿ ȮɌ Ő Ǻ ǝɑƱ Ǻ ƿ ɍ& Ɨ  !
ɎǎÐ©Ʒ(ɐǎ)ǂɏ%ȆȮǩÐ

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)ÙÐ:Ȇɏ
Family Staphylinidae a _ A E L Ǎ
Subfamily Omaliinae MacLeay v W p a _ A E L  Ǎ
1ɏ Eusphalerum (Eusphalerum) kojimai Y. Watanabe I M m v W p a ] o F x a _ AELȆ Ȯ Ʈ ä ɑ Otome-tôge~KurosakitakaoɌ Watanabe, 2015ɍɏ
³ ĕ ä ɑ  ȑ Ȏ Đ Ɍ Ĩ Ǹ Đ ɍɏ2ɏ Eusphalerum (Eusphalerum) mikurajimaense Y. Watanabe n E w M m a ] o F x v
Wpa_AELȆ Ȯ Ʈ ä ɑ Obannoo, Otome-tôge~KurosakitakaoɌ Watanabe, 2015ɍɏ
³ ĕ ä ɑ  ȑ Ȏ Đ Ɍ Ĩ Ǹ Đ ɍɏ
3ɏ Geodromicus harmaniɌ Y. Watanabeɍ a  m | n O D { v W p a _ A E LȆ Ȯ Ʈ ä ɑ Sato: Mikurajima-OhashiɌ Y. Watanabe, 2000: Psephidonus harmaniɍɏ³ ĕ ä ɑ ¾ ƌ ȧ , Š ē ,  ȑ Ȏ Đ Ɍ Ĩ Ǹ Đ ɍɎ ß â Ɏ  ē ɒ Ƚ â ɏ
4ɏ Nipponophloeostiba verrucifera Y. Watanabe U e d w S v W p a _ A E LȆ Ȯ Ʈ ä ɑ Sato, KawadaɌ Watanabe, 1962ɍɒ ƛ ư ~ ƒ Ȟ Ɍ 1969ɍɏ
³ ĕ ä ɑ  ȑ Ȏ Đ Ɍ  ü Đ Ɏ Ĩ Ǹ Đ Ɏ ­  Đ ɍɎ Ċ  Đ Ɏ Ǿ Ǽ Đ ɏ

4Subfamily Pselaphinae Latreille < x X A o L  Ǎ
5ɏ Batriscenellus (Batriscenellus) fallax (Sharp) d p a w E l < x X A o LȆ Ȯ Ʈ ä ɑ Mt. Oyama, SatoɌ Nomura, Kamezawa & Arai, 2013ɍɏ
³ ĕ ä ɑ ¾ ƌ ȧ Ɏ Š ē Ɏ  ȑ Ȏ Đ Ɍ  ü Đ Ɏ Ĩ Ǹ Đ ɍɏ
6ɏ Trissemus alienus (Sharp) ] n ? | m < x X A o LȆ Ȯ Ʈ ä ɑ Mt. Oyama Ɍ ƛ ư ~ ƒ Ȟ Ɏ 1969ɒ Nomura, Kamezawa & Arai, 2013ɍɏ
³ ĕ ä ɑ Š ē Ɏ  ȑ Ȏ Đ Ɍ  ü Đ Ɏ Ĩ Ǹ Đ ɍɎ ß â Ɏ  ē Ɏ ę Ĺ ɒ ¾ Ŝ ɉ Ɏ z L<ɏ7ɏ Tychus dichotomus Nomura et Lee S = x E q x < x X A o LȆ Ȯ Ʈ ä ɑ Mt. OyamaɌ Nomura, Kamezawa & Arai, 2013ɍɏ
³ ĕ ä ɑ â ĥ Đ Ɏ ¾ ƌ ȧ Ɏ Š ē Ɏ  ȑ Ȏ Đ Ɍ Ĩ Ǹ Đ ɍɎ  ē Ɏ Ą Ɇ ɒ Ƚ â ɏ
8ɏ Ctenisodes discedens (Sharp) k Q E L d H < x X A o LȆ Ȯ Ʈ ä ɑ SatoɌ Nomura, Kamezawa & Arai, 2013ɍɏ
³ ĕ ä ɑ Š ē Ɏ Ǚ Đ Ɏ £ ƒ } Đ Ɏ  ȑ Ȏ Đ Ɍ Ĩ Ǹ Đ ɍɎ  ē Ɏ Ą Ɇ ɏ
9ɏ Diartiger toshioi Nomura et Komatsu n E w s m [ d H g [ < x X A o LȆ Ȯ Ʈ ä ɑ Borozawa, Mt. OyamaɌ Nomura & Komatsu, 2012ɑ Diartiger fossulatus
toshioiɍɒ SatoɌ Nomura, Kamezawa & Arai, 2013ɍɏ
³ ĕ ä ɑ  ȑ Ȏ Đ Ɍ Ĩ Ǹ Đ ɍɏ
10ɏ Diartiger fossulatus izuinsulicola Nomura I s m [ d H g [ < x X A o LȆ Ȯ Ʈ ä ɑ Sato~KawadaɌ Nomura, Kamezawa & Arai, 2013ɍɏ
³ ĕ ä ɑ  ȑ Ȏ Đ Ɍ ǋ Ɗ Đ Ɏ Ĩ Ǹ Đ ɍɏ
Subfamily Tachyporinae MacLeay L x k Q a _ A E L  Ǎ
11ɏ Coproporus scitulus (Weise) E z U e m y E e a _ A E LȆ Ȯ Ʈ ä ɑ Kawada Ɍ Watanabe, 1961; Erchomus scitulus Weise ɍɒ Kawada, near
Akazawa, near Miyogaike Ɍ ƛ ư ~ ƒ Ȟ , 1969ɑ Erchomus scitulus Weiseɍɏ³ ĕ ä ɑ  ȑ Ȏ Đ Ɍ  ü Đ Ɏ Ĩ Ǹ Đ Ɏ ­  Đ ɍɎ ß â Ɏ  ē ɒ  â Ɏ z L < Ɍ Űť ɍɏ12ɏ Sepedophilus armatus (Sharp) E z H d p C ` I a _ A E LȆ Ȯ Ʈ äɑSato, KawadaɌ Watanabe, 1961; Conosoma armatum SharpɍɒSato, KawadaɌ ƛ ư ~ ƒ Ȟ Ɏ 1969ɑ Conosoma armatum Sharpɍɏ
³ ĕ ä ɑ ¾ ƌ ȧ Ɏ Š ē Ɏ  ȑ Ȏ Đ Ɍ  ü Đ Ɏ Ĩ Ǹ Đ Ɏ ­  Đ ɍɎ ß â Ɏ  ē Ɏ ĊĐɎǎùĐɎƅƃǰȩĐɒâɏ
13ɏ Sepedophilus germanus (Sharp) o E H d p C ` I a _ A E LȆ Ȯ Ʈ ä ɑ near KawadaɌ ƒ Ȟ ~ ȯ Ȑ Ē , 1983ɍɏ

5

³ ĕ äɑ¾ ƌ ȧ ɎŠ ē Ɏ ȑ Ȏ ĐɌ ò Đ ɎĨ Ǹ Đ ɍɎƏ Ș Đ Ɏȵ Ď Ɏß â Ɏ ē ɎƬĐɎĊĐɒâɏ
14ɏ Sepedophilus kobensis (Cameron) A S q | d p C ` I a _ A E LȆ Ȯ Ʈ ä ɑ KawadaɌ Watanabe, 1961ɍɏ
³ ĕ ä ɑ Š ē Ɏ  ȑ Ȏ Đ Ɍ  ü Đ Ɏ Ĩ Ǹ Đ ɍɏ
15ɏ Sepedophilus pumilus (Sharp) a N q | d p C ` I a _ A E LȆ Ȯ Ʈ ä ɑ Kawada, near AkazawaɌ ƛ ư ~ ƒ Ȟ , 1969ɍɏ
³ ĕ ä ɑ ¾ ƌ ȧ Ɏ Š ē Ɏ  ȑ Ȏ Đ Ɍ Ĩ Ǹ Đ Ɏ ­  Đ ɍɎ ß â Ɏ  ē ɎƬĐɎǎùĐɎĊĐɎóǥòĐɏ16ɏ Sepedophilus tristiculus (Weise) W s d p C ` I a _ A E LȆ Ȯ Ʈ ä ɑ KawadaɌ Watanabe, 1961: Conosoma tristiculum Sharpɍɒ Kawada, near
Akazawa Ɍ ƛ ư ~ ƒ Ȟ Ɏ 1969ɍɏ
³ ĕ äɑŠ ē Ɏ ȑ Ȏ ĐɌ  ü Đ ɎĨ Ǹ Đ ɍɎß âɒz L < Ɍ L j x < , Ű ť ɍɏ
17ɏ Sepedophilus varicornis (Sharp) E z d p C ` I a _ A E LȆ Ȯ Ʈ ä ɑ KawadaɌ Watanabe, 1961: Conosoma vasicorne Sharpɍɒ Kawada, near
Akazawa Ɍ ƛ ư ~ ƒ Ȟ Ɏ 1969ɍɏ
³ ĕ ä ɑ ¾ ƌ ȧ Ɏ Š ē Ɏ  ȑ Ȏ Đ Ɍ  ü Đ Ɏ Ĩ Ǹ Đ ɍɎ ȵ Ď Ɏ ß â Ɏ  ē ɎƬĐɎĊĐɎóǥòĐɒȽâɎâɏ
18ɏ Tachinus (Tachinoderus) diminutus Sharp I E z m y E e a _ A E LȆ Ȯ Ʈ ä ɑ Kawada, near Miyogaike Ɍ ƛ ư ~ ƒ Ȟ , 1969ɍɏ
³ ĕ ä ɑ Š ē Ɏ  ȑ Ȏ Đ Ɍ Ĩ Ǹ Đ ɍɎ ß â Ɏ  ē ɏ
19ɏ Tachinus (Tachinus) mimulus Sharp C j x m y E e a _ A E LȆ Ȯ Ʈ ä ɑ KawadaɌ Watanabe, 1961ɍɒ Sato, Kawada, Mt. Oyama, near MiyogaikeɌ ƛ ư ~ ƒ Ȟ Ɏ 1961ɍɏ
³ ĕ ä ɑ ¾ ƌ ȧ Ɏ Š ē Ɏ  ȑ Ȏ Đ Ɍ Ĩ Ǹ Đ ɍɎ £ ƒ } Đ Ɏ ß â Ɏ  ē ɏ
Subfamily Aleocharinae Fleming d H g [ a _ A E L  Ǎ
20ɏ Myrmecocephalus japonicus Bernhauer > N U r P n R a _ A E LȆ Ȯ Ʈ ä ɑ Kawada Ɍ ƛ ư ~ ƒ Ȟ Ɏ 1969ɍɏ
³ ĕ ä ɑ ¾ ƌ ȧ Ɏ Š ē Ɏ  ȑ Ȏ Đ Ɍ Ĩ Ǹ Đ ɍɎ ß â Ɏ  ē ɏ
21ɏ Aleochara (Aleochara) parens Sharp I E z d H g [ a _ A E LȆ Ȯ Ʈ ä ɑ KawadaɌ Watanabe, 1961ɍɒ Sato, Kawada, near MiyogaikeɌ ƛ ư ~ ƒ
Ȟ , 1969ɍɏ
³ ĕ ä ɑ ¾ ƌ ȧ Ɏ Š ē Ɏ  ȑ Ȏ Đ Ɍ  ü Đ Ɏ Ĩ Ǹ Đ ɍɎ ¾ Ǉ Ɋ Đ Ɏ ß â Ɏ  ē ɎĊĐɒȽâɎ¾ŜɉɎâɏ

6

Subfamily Scaphidiinae Latreille Z @ C ` I o L  Ǎ
22ɏ Scaphisoma haemaorrhoidale Reitter W s G L Z @ C ` I o LȆ Ȯ Ʈ ä ɑ Kawada, near Akazawa Ɍ ƛ ư ~ ƒ Ȟ , 1969ɍɏ
³ ĕ äɑŠ ē Ɏ ȑ Ȏ ĐɌ  ü Đ ɎĨ Ǹ Đ ɍɎß â Ɏ ē ɎĄ Ɇɒ¾ Ŝ ɉ Ɏ â Ɏz L < Ɍ L j x < ɍɏ
23ɏ Scaphisoma galloisi Achard B z < G L Z @ C ` I o LȆ Ȯ Ʈ ä ɑ near Miyogaike Ɍ ƛ ư ~ ƒ Ȟ Ɏ 1969ɒ Scaphisoma harmandi Achardɍɏ³ ĕ ä ɑ Š ē Ɏ  ȑ Ȏ Đ Ɍ Ĩ Ǹ Đ ɍɎ ß â Ɏ Ą Ɇ ɏ
24ɏ Scaphisoma rufum Achard < A G L Z @ C ` I o LȆ Ȯ Ʈ ä ɑ Kawada Ɍ ƛ ư ~ ƒ Ȟ , 1969ɍɏ
³ ĕ ä ɑ ¾ ƌ ȧ Ɏ Š ē Ɏ  ȑ Ȏ Đ Ɍ Ĩ Ǹ Đ ɍɎ ß â Ɏ  ē Ɏ Ą Ɇ Ɏ ó ǥ ò Đ Ɏ ƅǣŠĐɎǅåĐɎǾǼĐɒȽâɎâɏ
25ɏ Scaphobaeocera japonica (Reitter) [ e = z k Y = G L Z @ C ` I o LȆ Ȯ Ʈ äɑKawada Ɍ ƛ ư~ƒ Ȟ Ɏ1969ɒToxidium japonicum Reitter [ e = z k
Q Z @ C ` I o L ɍɏ
³ ĕ ä ɑ Š ē ,  ȑ Ȏ Đ Ɍ Ĩ Ǹ Đ ɍɎ  ē ɏƇɑȆ(ûÐ&Ȳ!)©ǍǴÂî(Ņǉ&è
1981 Ě & Löbl & 4  ! Š ǎ & Ƞ ¡ ( Ň ǎ!ȆȮɏ¢ɎȆ ț~ƻ Ǽ  8 !  7 ( " Ɏ 8 5(ǎ#(ſȚŮȅ&4!ǎÐ:ǈþ7Ĭǿ

7#(ľł:Èɏ" )  ĭ 1969 Ě & Ȇ Ȯ  8  ǎ ą Ð : ĝ Ư   ɏ
Subfamily Piestinae Erichson d w S a _ A E L  Ǎ
26ɏ Siagonium gracile Sharp k Q d w S a _ A E LȆ Ȯ Ʈ ä ɑ near Kawada Ɍ ƒ Ȟ ~ ȯ Ȑ Ē Ɏ 1983ɍɏ
³ ĕ ä ɑ ¾ ƌ ȧ Ɏ Š ē Ɏ  ȑ Ȏ Đ Ɍ Ĩ Ǹ Đ Ɏ ­  Đ ɍɎ ß â Ɏ  ē Ɏ Ċ  Đ ɏ
27ɏ Siagonium haroldi Weise < A q | d w S a _ A E LȆ Ȯ Ʈ ä ɑ Kawada Ɍ ƛ ư ~ ƒ Ȟ Ɏ 1969ɑ Siagonium vittatum Fauvel d w S a _A E L ɍɏ³ ĕ ä ɑ Š ē Ɏ  ȑ Ȏ Đ Ɍ  ü Đ Ɏ Ĩ Ǹ Đ ɍɎ Ċ  Đ ɒ Ì ƕ ɏ
Subfamily Osoriinae Erichson W W a _ A E L  Ǎ
28ɏ Osorius angustulus Sharp f [ W W a _ A E LȆ Ȯ Ʈ ä ɑ near Miyogaike Ɍ ƛ ư ~ ƒ Ȟ , 1969ɍɏ
³ ĕ ä ɑ ¾ ƌ ȧ Ɏ Š ē Ɏ  ȑ Ȏ Đ Ɍ Ĩ Ǹ Đ ɍɎ ȵ Ď Ɏ ß â Ɏ  ē Ɏ ǘ ¸ ƅ ` Đ ɎǎùĐɎĄɆɎƅƃǰȩĐɒȽâɏ

7

29ɏ Osorius taurus Sharp W ` f [ W W a _ A E LȆ Ȯ Ʈ äɑKawadaɌ Watanabe, 1961ɍ
ɒKawada, near MiyogaikeɌ ƛ ư~ƒ Ȟ Ɏ1969ɍɏ
³ ĕ ä ɑ Š ē Ɏ  ȑ Ȏ Đ Ɍ  ü Đ Ɏ Ĩ Ǹ Đ ɍɎ ß â Ɏ  ē Ɏ Ą Ɇ Ɏ Ċ  Đ Ɏ É ƃǰȩĐɒȽâɏ30ɏ Nacaeus longulus (Sharp) ] B q x W W a _ A E LȆ Ȯ Ʈ äɑKawadaɌ Watanabe, 1961ɒLispinus impressicollis var. longulus Sharpɍɒ
Kawada, near Akazawa, near MiyogaikeɌ ƛ ư~ƒ Ȟ Ɏ1969ɑLispinus impressicollis
longulus Sharp U e k Q a _ A E L ɍɏ
³ ĕ äɑŠ ē Ɏ ȑ Ȏ ĐɌ  ü Đ ɎĨ Ǹ Đ Ɏ­  Đ ɍɎȵ Ď Ɏß â Ɏ ē ɎĄ Ɇ ɎƬĐɎĊĐɒȽâɎÌƕɏ
31ɏ Thoracopholus certatus Sharp s m [ k Q W W a _ A E LȆ Ȯ Ʈ äɑKawadaɌ Watanabe, 1961ɍ
ɒKawada, near MiyogaikeɌ ƛ ư~ƒ Ȟ Ɏ1969ɍɏ
³ ĕ äɑŠ ē Ɏ ȑ Ȏ ĐɌ Ĩ Ǹ Đ Ɏ­  Đ ɍɎą ǔ Å Ȏ ĐɌ ž Đ ɍɎß â Ɏ ē ɎȵĎɎĊĐɎ[AwĐɎóǥòĐɎǾǼĐɒÌƕɏ
Subfamily Oxytelinae Fleming P N M a _ A E L  Ǎ
32ɏ Anotylus japonicus (Cameron) d w S P N M a _ A E LȆ Ȯ Ʈ ä ɑ KawadaɌ ƛ ư ~ ƒ Ȟ Ɏ 1969ɒ Rimba japonica (Cameron)ɍɏ
³ ĕ ä ɑ Š ē Ɏ  ȑ Ȏ Đ Ɍ Ĩ Ǹ Đ ɍɎ ȵ Ď Ɏ ß â ɎƬĐɎǎùĐɏ33ɏ Anotylus laticornis (Sharp) d p E z P N M a _ A E LȆ Ȯ Ʈ äɑSato, KawadaɌ Watanabe, 1961ɑOxytelus (Anotylus) latiusculus Kraatz ?ɒƛ ư ~ ƒ Ȟ Ɏ 1969ɑ Oxytelus laticornis Sharpɍɏ
³ ĕ ä : Š ē Ɏ  ȑ Ȏ Đ Ɍ Ĩ Ǹ Đ Ɏ ­  Đ ɍɎ ß â Ɏ  ē Ɏ Ą Ɇ ɒ  â ɏ
34ɏ Anotylus lewisius (Sharp) y = N W s P N M a _ A E LȆ Ȯ Ʈ ä ɑ Sato, KawadaɌ Watanabe, 1961ɑ Oxytelus (Anotylus) lewisius Sharpɒ ƛư ~ ƒ Ȟ , 1969ɑ Oxytelus lewisius Sharpɍɏ
³ ĕ ä ɑ Š ē Ɏ  ȑ Ȏ Đ Ɍ  ü Đ Ɏ Ĩ Ǹ Đ Ɏ ­  Đ ɍɎ ß â Ɏ  ē Ɏ Ą Ɇ ɎƬĐɎǎùĐɎĊĐɎóǥòĐɎƅǣŠĐɎęĊĐɒÌƕɎâɏ
35ɏ Oxytelus piceus Linnaeus C b _ P N M a _ A E LȆ Ȯ Ʈ ä ɑ SatoɌ ƛ ư ~ ƒ Ȟ Ɏ 1969ɍɏ
³ ĕ ä ɑ ¾ ƌ ȧ Ɏ Š ē Ɏ  ȑ Ȏ Đ Ɍ  ü Đ Ɏ Ĩ Ǹ Đ Ɏ ­  Đ ɍɎ ą ǔ Å Ȏ Đ Ɍ ƣĐ ɍɎ ß â Ɏ  ē Ɏ [ A w   Đ ɒ Ƚ â Ɏ ¾ Ŝ ɉ Ɏ  â Ɏ ť Ɖ ¿ Ɏ v  z V c Ɏ<fxAɏSubfamily Steninae MacLeay p T A a _ A E L  Ǎ
36ɏ Stenus macies Sharp \ > l Q p T A a _ A E L

8Ȇ Ȯ Ʈ ä ɑ KawadaɌ Watanabe, 1961ɒ ƛ ư ~ ƒ Ȟ Ɏ 1969ɍɏ
³ ĕ ä ɑ ¾ ƌ ȧ Ɏ Š ē Ɏ  ȑ Ȏ Đ Ɍ  ü Đ Ɏ Ĩ Ǹ Đ Ɏ ­  Đ ɍɎ ȵ Ď Ɏ ß â Ɏ ēɎĄɆɎĊĐɒâɏ
Subfamily Paederinae Fleming < x B S a _ A E L  Ǎ
37ɏ Astenus brevipes (Sharp) d p L x F z a _ A E LȆ Ȯ Ʈ ä ɑ SatoɌ Watanabe, 1961ɒ ƛ ư ~ ƒ Ȟ Ɏ 1969ɍɏ
³ ĕ ä ɑ Š ē Ɏ  ȑ Ȏ Đ Ɍ  ü Đ Ɏ Ĩ Ǹ Đ Ɏ Ⱥ } Đ ɍɎ ȵ Ď Ɏ ß â Ɏ  ē Ɏ ó ǥòĐɎƅǣŠĐɒȽâɏ
38ɏ Homaeotarsus (Hesperobium) japonicus (Sharp) s m [ ] B ? a _ A E LȆ Ȯ Ʈ ä ɑ near MiyogaikeɌ ƛ ư ~ ƒ Ȟ Ɏ 1969ɑ Ochthephilum japonicum Sharpɍɏ
³ ĕ ä ɑ ¾ ƌ ȧ Ɏ Š ē Ɏ  ȑ Ȏ Đ Ɍ Ĩ Ǹ Đ ɍɎ  ē Ɏ ƅ ǣ Š Đ ɒ Ì ƕ Ɏ Ƚ â Ɏ ¾
Ŝɉɏ39ɏ Lithocharis nigriceps Kraatz E z O [ B x a _ A E LȆ Ȯ Ʈ äɑSatoɌ Watanabe, 1961ɑLithocharis ochracea Gravenhorstɒƛ ư~ƒ Ȟ Ɏ
1969ɍɏ
³ ĕ äɑŠ ē Ɏ ȑ Ȏ ĐɌ  ü Đ ɎĨ Ǹ Đ Ɏ­  Đ ɍɎ£ ƒ } Đ Ɏą ǔ Å Ȏ ĐɌ ƣĐ ɍɎ ȵ Ď Ɏ ß â Ɏ  ē Ɏ ǎ ù Đ Ɏ Ċ  Đ Ɏ ó ǥ ò Đ Ɏ Ǿ Ǽ Đ Ɏ Á ò ť Đ Ɏ ¾ òťĐɒÌƕɎȽâɎâɎ=|\ɎťƉ¿ɏ
40ɏ Scopaeus virilis Sharp U e d p E e l Q a _ A E LȆ Ȯ Ʈ ä ɑ SatoɌ Watanabe, 1961ɒ ƛ ư ~ ƒ Ȟ Ɏ 1969ɍɏ
³ ĕ äɑŠ ē Ɏ ȑ Ȏ ĐɌ Ĩ Ǹ Đ Ɏ­  Đ ɍɎȵ Ď Ɏß â Ɏß â ƅ ` Đ Ɏǎ ù Đ ɎĊĐɎƅǣŠĐɒâ, ťƉ¿ɏ41ɏ Rugilus (Eurystilicus) rufescens (Sharp) E e l Q a _ A E LȆ Ȯ Ʈ ä ɑ KawadaɌ Watanabe, 1961ɒ ƛ ư ~ ƒ Ȟ Ɏ 1969ɍɏ
³ ĕ ä ɑ ¾ ƌ ȧ Ɏ Š ē Ɏ  ȑ Ȏ Đ Ɍ  ü Đ Ɏ Ĩ Ǹ Đ Ɏ ­  Đ ɍɎ ȵ Ď Ɏ ß â Ɏ ēɎĄɆɎĊĐɒÌƕɎȽâɎ¾ŜɉɎâɎzL<ɎťƉ¿ɏ
42ɏ Rugilus (Rugilus) japonicus Y. Watanabe W m C E e l Q a _ A E LȆ Ȯ Ʈ ä ɑ Kawada, Kawaguchi Ɍ ƛ ư ~ ƒ Ȟ Ɏ 1969ɍɏ
³ ĕ ä ɑ  ȑ Ȏ Đ Ɍ  ü Đ Ɏ Ĩ Ǹ Đ Ɏ ­  Đ ɍɎ ǎ ù Đ Ɏ Ċ  Đ Ɏ ó ǥ ò Đ Ɏ ƅǣŠĐɏ43ɏ Sunesta setiger (Sharp) S U H E e l Q a _ A E LȆ Ȯ Ʈ äɑKawada, near Akazawa Ɍ ƛ ư~ƒ Ȟ Ɏ1969ɑStilicopsis setigera Sharpɍɏ
³ ĕ ä ɑ Š ē Ɏ  ȑ Ȏ Đ Ɍ  ü Đ Ɏ Ĩ Ǹ Đ Ɏ ­  Đ ɍɎ ą ǔ Å Ȏ Đ Ɍ ƣ Đ ɍɎ ßâɎēɎĄɆɎǎùĐɎ[AwŶğĐɎóǥòĐɒÌƕɎȽâɎS=ɏ

9

44ɏ Pinophilus lewisius Sharp y = N E e g [ a _ A E LȆ Ȯ Ʈ äɑKawadaɌ Watanabe, 1961ɒƛ ư~ƒ Ȟ Ɏ1969ɒPinophilus lewisi Sharpɍɏ
³ ĕ ä ɑ Š ē Ɏ  ȑ Ȏ Đ Ɍ  ü Đ Ɏ Ĩ Ǹ Đ Ɏ ­  Đ ɍɎ ß â Ɏ  ē Ɏ Ą Ɇ Ɏ ǎ ùĐɎĊĐɒÌƕɎȽâɏ
45ɏ Oedechirus lewisius Sharp E z b _ < x B S a _ A E LȆ Ȯ Ʈ ä ɑ KawadaɌ ƛ ư ~ ƒ Ȟ Ɏ 1969ɍɏ
³ ĕ äɑŠ ē Ɏ ȑ Ȏ ĐɌ  ü Đ ɎĨ Ǹ Đ Ɏ­  Đ ɍɎǙ Đ Ɏȵ Ď Ɏß â Ɏ ē ɎĄɆɎǎùĐɎĊĐɎƅǣŠĐɒȽâɎ¾Ŝɉɏ
Subfamily Staphylininae Latreille a _ A E L  Ǎ
46ɏ Heterothopus cognatus Sharp k Q U e W s o ] a _ A E LȆ Ȯ Ʈ ä ɑ Sato, KawadaɌ Watanabe, 1961ɒ ƛ ư ~ ƒ Ȟ Ɏ 1969ɍɏ
³ ĕ äɑ¾ ƌ ȧ ɎŠ ē Ɏ ȑ Ȏ ĐɌ  ü Đ ɎĨ Ǹ Đ Ɏ­  Đ ɍɎǙ Đ Ɏ£ ƒ } Đ ɎßâɎēɎ

ƬĐɎĊĐɒȽâɎâɏ

47ɏ Bisnius germanus (Sharp) U r = z ^ P I B L w a _ A E LȆ Ȯ Ʈ äɑSato, Kawada, near MiyogaikeɌ ƛ ư~ƒ Ȟ Ɏ1969ɑPhilonthus germanus
Sharpɍɏ
³ ĕ ä ɑ Š ē Ɏ  ȑ Ȏ Đ Ɍ  ü Đ Ɏ Ĩ Ǹ Đ Ɏ ­  Đ ɍɎ ȵ Ď Ɏ ß â Ɏ  ē ɎƬĐɎĊĐɎóǥòĐɏ48ɏ Cafius (Pseudoremus) algarum (Sharp) d p < b S > n j a _ A E LȆ Ȯ Ʈ ä ɑ SatoɌ Watanabe, 1961ɒ ƛ ư ~ ƒ Ȟ Ɏ 1969ɍɏ
³ ĕ ä ɑ Š ē Ɏ  ȑ Ȏ Đ Ɍ  ü Đ Ɏ Ĩ Ǹ Đ Ɏ ­  Đ ɍɎ £ ƒ } Đ Ɏ ȵ Ď Ɏ ß â ɎēɎóǥòĐɒȽâɎɅƔɎâɎzL<ɏ49ɏ Cafius (Pseudoremus) histrio (Sharp) k Q < b S > n j a _ A E LȆ Ȯ Ʈ ä ɑ SatoɌ Watanabe, 1961; ƛ ư ~ ƒ Ȟ Ɏ 1969ɍɏ
³ ĕ äɑ¾ ƌ ȧ ɎŠ ē Ɏ ȑ Ȏ ĐɌ  ü Đ ɎĨ Ǹ Đ Ɏ­  Đ ɍɎ£ ƒ } Đ Ɏß â ɎǎùĐɎĊĐɒȽâɎ¾ŜɉɎɅƔɎzL<ɏ50ɏ Philonthus (Philonthus) microcephalus Sharp < A d w S I B L w a _ A E LȆ Ȯ Ʈ ä ɑ KawadaɌ ƛ ư ~ ƒ Ȟ Ɏ 1969ɍɏ
³ ĕ ä ɑ Š ē Ɏ  ȑ Ȏ Đ Ɍ Ĩ Ǹ Đ ɍɎ  ē ɏ
51ɏ Philonthus (Philonthus) tardus Kraatz i x < A b I B L w a _ A E LȆ Ȯ Ʈ äɑKawadaɌ Watanabe, 1961ɍ
ɒSato, Kawada, near AkazawaɌ ƛ ư~ƒ Ȟ Ɏ
1969ɍɏ
³ ĕ ä ɑ Š ē Ɏ  ȑ Ȏ Đ Ɍ  ü Đ Ɏ Ĩ Ǹ Đ Ɏ ­  Đ ɍɎ £ ƒ } Đ Ɏ ȵ Ď Ɏ ß â ɎąȑĐɎēɎĄɆɎƬĐɎĊĐɎƅǣŠĐɎñǰȱĐɒÌƕɎâɎǾ

10

=|\ɎNxw|Aɏ52ɏ Quedius (Microsaurus) simulans Sharp ] n W s a _ A E LȆ Ȯ Ʈ ä ɑ Sato, KawadaɌ ƛ ư ~ ƒ Ȟ Ɏ 1969ɍɏ
³ ĕ ä ɑ ¾ ƌ ȧ Ɏ Ǌ ň Đ Ɏ · ć Đ Ɏ Š ē Ɏ  ȑ Ȏ Đ Ɍ  ü Đ Ɏ Ĩ Ǹ Đ ɍɎ £ ƒ }ĐɎąȑĐɎēɒȽâɎâɏ
53ɏ Creophilus maxillosus (Linnaeus) @ @ a _ A E LȆ Ȯ Ʈ ä ɑ SatoɌ ƒ Ȟ ~ ȯ Ȑ Ē Ɏ 1983ɍɏ
³ ĕ ä ɑ ¾ ƌ ȧ Ɏ Š ē Ɏ  ȑ Ȏ Đ Ɍ Ĩ Ǹ Đ Ɏ ­  Đ ɍɎ ß â Ɏ  ē Ɏ Ċ  Đ Ɏ óǥòĐɎƅǣŠĐɎęĊĐɎǅåĐɎǾǼĐɒÌƕɎȽâɎ¾ŜɉɎâɎz L < Ɍ L j x < ɍɎ = | \ Ɏ a { = Ɏ ź ē Ɏ ¾ < f x A Ɏ < p x A ɏ
54ɏ Megalinus suffusus (Sharp) C b _ ] B a _ A E LȆ Ȯ Ʈ ä ɑ KawadaɌ Watanabe, 1961ɍɒ Sato, KawadaɌ ƛ ư ~ ƒ Ȟ Ɏ 1969ɍɏ
³ ĕ ä ɑ ¾ ƌ ȧ Ɏ Š ē Ɏ  ȑ Ȏ Đ Ɍ ò Đ Ɏ  ü Đ Ɏ Ĩ Ǹ Đ ɍɎ ß â Ɏ ą ȑ Đ Ɏ ēɎĄɆɎƬĐɎĊĐɒÌƕɎâɏ

55ɏ Liotesba punctiventris (Sharp) < A b W s E e ] B a _ A E LȆ Ȯ Ʈ ä ɑ KawadaɌ ƛ ư ~ ƒ Ȟ Ɏ 1969ɑ Megalinus punctiventris Sharpɍɏ
³ ĕ ä ɑ Š ē Ɏ  ȑ Ȏ Đ Ɍ Ĩ Ǹ Đ ɍɎ ß â Ɏ  ē Ɏ Ą Ɇ ɏ
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Nomura, S. & T. Komatsu, 2012. Two new subspecies of Diartiger fossulatus Sharp
Ɍ Coleoptera, Staphylinidae, Pselaphinaeɍ from Japanɏ Esakia, Fukuoka, Ɍ 52ɍ: 9-15ɏ
Nomura, S., H. Kamezawa & S. Arai, 2013. A reviw of psalaphine beetles (Coleoptera,
Staphylinidae) from the Izu-shoto Isls., Tokyo, Japan. Elytra, Tokyo, New Series, 3:31-43ɏ
Sawada, H. & Y. Watanabe, 1960. Description of a new species of lucanid-beetles from
Mikura Island in the Izu Islands, Japan.

J. agric. Sci. Tokyo Nogyo Daigaku, 6: 102ɏƛ ư ƨ Ż ~ ƒ Ȟ ƈ ő Ɏ 1969. Ĩ Ǹ Đ ( Ő Ǻ ǁ ɏ ť  ȝ ů ò û ȝ û ȸ ê , 14: 1-48ɏ
ũ ư ƈ ·~ č ý ·Ɍ Ǡ ȸ ɍɎ2013. ŏ Š Ʈ a _ A E L Ǎ Ǟ ƿ ȮɌ Ő Ǻ ǝɑƱ Ǻ ƿ ɍɏ ē ò û Ǟ Î ǆ Ǒ Â Ƥ Ʉ ǆ Ǒ ê Ô ,Ɍ 11ɍ : 69-218ɏŬ Ȑ Ƨ  Ɏ1954ɏ ȑ  Đ &ǆǑ

 7 Lamellicornia ( ³ ĕ &   ! ɏǺ êɌ ŏ Š Ő Ǻê Ô ɍɎɌ 10ɍɑ 17 pp., 3 plsɏŬ Ȑ Ƨ  Ɏ1961ɏ  ȑ  Đ & 7 A n C x o L Ǎ Ʊ Ǻ ( ³ ĕ ɏ Ő Ǻ , 29: 215-225ɏ

Watanabe, Y., 1961. The staphylinid-fauna of the middle and southern Izu Islands. J. agric.
Tokyo Nogyo Daigaku, 6: 348-356.
Watanabe, Y., 1962. Descriptions of a new genus and species of Omaliinae from the Izu
Islands, Japan (Coleoptera, Staphylinidae).

J. agric. Sci. Tokyo Nogyo Daigaku, 8:77-80.
Watanabe, Y., 1990. A taxonomic study on the subfamily Omaliinae from JapanɌ Coleoptera, Staphylinidaeɍɏ Mem. Tokyo Univ. Agric., 31: 55-391.
Watanabe, Y., 2000. Occurrence of Psephidonus harmani Y. Watanabe (Coleoptera,
Staphylinidae) on the Island of Mikurajima of the Izu Islands, central Japan. Elytra,
Tokyo, 28: 54.

13ƒ Ȟ ƈ ő Ɏ 2014ɏ v W p a _ A E L  Ǎ Pp. 2-4ɏ č Š Õ ę , ŏ Š ( a _ A E L  Ǎǁ ȃ ő & /  ! Ɍ 1ɍɑ ŏ Š Ʈ a _ A E L Ǎ  Ǎ ¶ ű Ȃ ɑ ȃ ő Ĝ ~ ä æ ȱ ſ Ț ~ ³
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Watanabe, Y., 2015. Two new species of Eusphalerum (Coleoptera, Staphylinidae) from
Mikura-jima Island of the Izu Islands off central Honshu, Japan. Elytra, Tokyo, New
Series, 5: 17-22ɏƒ Ȟ ƈ ő~ȯ Ȑ Ē ȧ ő Ɏ1983ɏ  ȑ Ĩ Ǹ Đ

5 ŋ Ȇ Ȯ ( a _ A E LɌ a _ A E L Ǎ³ ĕ ȕ ŉ 11ɍɏ Ʊ Ǻ ^ t  N ɎɌ 62ɍ : 4ɏ± ǃ 1-3ɏ Ĩ Ǹ Đ Ʈ v W p a _ A E L  Ǎ ( 3 ǎ ɏ
1. Geodromicus harmani (Y. Watanabe) a  m | n O D { v W p a _ A E L
2. Eusphalerum (Eusphalerum) kojimai Y. Watanabe I M m a ] o F x v W p a _ A
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3. Eusphalerum (Eusphalerum) mikurajimaense Y. Watanabe n E w M m a ] o F x v
Wpa_AEL14

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u_g{ˆ7ƜƎĪ˒pp. 41-62˒ŭƖȤãȅʱ˒Ƨ£˓đʎŢˊŃƣ ț˓2007˓Ƨ£ǦǝŅʣ6JǯʯŒ§ȗȅǺɑʼ7ǿǼ˓Sessile

Organisms, 24:9-19.
ȟʣǔ˓2011˓ǝˈň6ɋèHK,ªʅʁņŨɦņƇɲ˓ĉʌm` 679:23-31˓ńŇƈ¡˓1999˓ʇ7rdQ`˗ȟ7ʇ,.7ɪÚ3ȅů˓294 pp.˒ƧǝĞĪÍǸ«˒Ƨ£˓ńŇƈ¡ơƥĥħơ¥ƑħłêĺɯŊʓƗɯƭ¡ʢłɯɟŖƨ ɗĠĞʎ»ÿļɢǎʨơĩȋʜɲǓˇ÷ȈÑĠö¢ ĭ. 2004a. Ƒƣ6JǝȆȅǺ7¤ǴȐȩÄ3ÎƇ˗Ƒƣyj`Ī«ɛǶȂĘµÅĦü«6GJP[jɿƬ7ȿƪH.Ƒƣyj`Ī«ɼ˒59˗22-44.
ńŇƈ¡ơ¥ƑħƑƣyj`Ī«ɛǶȂĘµÅĦü«˓2004b˓Ƒƣ6¤ǴȐ6ȩ
Ä$K,ˀĈƦǝȆÚǺ7ÎŐźĞ601˓ƑƣwYjĪ«ē˒42:132-144.
ńŇƈ¡˓2009˓ǝ7ĜƦȅǺ Q & A. ƑƣwYjĪ«Ƒƣyj`Ī«ˏɊː˒
ǝ7ĜƦȅǺ˗¤ʯ6G/1Ƅ$K,ĉȀ7ǝ˒pp.3-18˒ƧǝĞĪÍǸ«˒Ƨ£.
Ʊäƿ˓1994˓Ƽǚǥ6JǯʯŒ§ȗȅǺ7ȫˆȾŲ3ǿĨʧ˓§ȗȅǺțȮ˒11: 1-10˓Øųʸ¦˓2006˓XcPVu_g{˗ÝȹH1ÝƑƣ7ǯʯŒNőŎ%,Ǝ%
ȩÄȫ˓Ƒƣ§ȗȅǺĪ«ˏɊː˒u_g{ˆ7ƜƎĪ˒pp. 80-92˒ŭƖȤãȅʱ˒Ƨ£˓ØɧƓģʦǡÃ˓2001˓TtVmWQkvfY˗ªʅʁņÆņ7ǝH˓157 pp.˒ʲū\~m[^a˒Ƨ£˓
Koike, F. and K. Iwasaki. 2010. A simple range expansion model of multiple pathways: the case ofnonindigenous green crab Carcinus aestuarii in Japanese waters. Biological Invasions,
13:459-470.

22Ʈƣʢ˓2015˓BH"39ʼ˓řǲȶɟˏɊɤː˒BH,I˒pp. 105-125˒ŨɦņɵÁà«˒Ũɦņƥ˓ɯƥû˓1998˓ªʅļȲäʁņ7cVW

Q601˓;,.<˒78&79 òŎ: 24-28˓ɯƥû˓1999˓ªʅļȲäʁņǝč6

JʇˆÎŐ˓Ƨ£ʤǇȆɻˌĔɿƬțȮēø˒211:1-124.
ɯƥûŵĖɡǎ˓2000˓īņȆ\^{WQ601˓.I@,O˒31: 67-68˓ɯƥûƟƝȦĠ˓2014˓ªʅĞņǝǔǠŁǇƐɹ2Žʼ%,XxQ_Tt`ŀ7Ƣɸʒȫ˙ ;,.<˒119: 18-21.
ģʦǡÃƞˋý¤˓2004˓ªʅʁņªʅĞņJǙǝŬkVˆȕˏȉǄɅ˒Þəȓ˒Ȏľȓː˓ƑƣȅǺĉȁĪ««ē˒59:49-69˓
Okutani˒T. 1972. Molluscan fauna on the submarine banks Zenisu, Hyotanse and Takase, near the
Izu-Shichito Islands. Bulletin of Tokai Regional Fisheries Research Laboratory 72:63-144.
ģʄþðóɣɌʻ˓2004˓ªʅļȲäʁņùʕ7ȠG<ņƲHŽʼ$K,Q
WQˆ˓.I@,O˒35:29-36˓Ğ ʄ ʞ Ġ ˓ 2004. Ƒ ƣ 7 ǝ ǔ ȩ Ä ȅ Ǻ 3 * 7 ȩ Ä ʝ Ȫ 6 0  1 . Ƒ ƣ y  j ` Ī « ɼ ,59:45-57.
ǟɲŰĭ˓2005˓Æņ2Žʼ$K,ʿBNŒ<,Y^}dV˓.I@,O˒35: 110-113˓
Sousa, W. P. 1979. Disturbance in marine inter-tidal boulder fields: the non-equilibrium
maintenance of species-diversity. Ecology, 60:1225-1239.
ʨơÂɐ¸Ȉǔ¡˓1967˓ªʅĞņG<*7§ʖǝč7Vm601˓ȉǄˆ7țȮ˒
3:86-104.
ÈǝķĚĠ˓1995˓ʿņ7ɥəˆ˓ʋǨȨĪțȮŴŠē˓



 ˓

Wu, S. and T.Okutani. 1984.The deepsea Chitons (Mollusca: Polyplacophora) collected by the R/V
Soyo-Maru fom Japan, 1, Lepidopleuridae. Venus, 43:1-31.
Yamaguchi, T. 1973. On the Megabalanus in Japan. Publications from the Seto Marine Biological
Laboratory, 21:115-140.
Yamaguchi, T. 1977. Taxonomic studies on some fossil and recent Japanese Balanoidea (Part 1).
Transactions on Proceedings of Plaeontological Society of Japan, 107:135-160.
Yamaguchi, T. 1987. Change sin the barnacle fauna since the Miocene and the infraspecificstructure of Tetraclita in Japan (Cirripedia; Balanomorpha). Bulletin of Marine Science,
41:337-350.
łêĺ˓1989˓ĜćHƑƣ6ȩ­%,u_g{ˆ˒ǻ6ĉȁȐÎŐG<ȅů7ěÜ˓ȥĢŊɛǶɼʋƋ˒10:17-32.
łêĺ˓1995˓AOˏɥːəɅ CIRRIPEDIA˓ɯƥʢˏɊɤː˒äɠƵȽƑƣǝŅÚǺąʬ II˒pp. 116-133˒µɗȤ˒Ğʲ˓

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Arase, T. and Okano, T. (2015) Influence of fertilization on native plants and exotic pasture grasseson the fascined landslide slopes in Mikura-Jima Island, Japan. International Journal of
GEOMATE, 8(2): 1316-1322
Bailey, D.R. (1988) Weed control in tropical pastures. In: Tropical forege legumes (eds. Skerman,
P.J., Cameron, D.G. and Riveros, F.) pp. 136-147, FAO, Rome.
Bazzaz, F. A. (1996) Plants in changing environment. 320 pp. Cambridge University Press,
Cambridge.
Holl, K. D., Loik, M. E., Lin, E. H. and Samuels, I. A. (2000) Tropical montane forest restoration in
Costa Rica: overcoming barriers to dispersal and establishment. Restoration Ecology, 8(4):339-349.
ĩ ʤ˽͋1984͌)Cǭȳ1ɀȐ͍̊ª͍38(4)͓26-31.
¡ǗôʞƌʳȒɉǈĸ͆˝!;͋2003͌ȬġıšɎȋ̱,ōǁ%ĴǜȲʭ44

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ʥɐˉ͋1980͌ƎʲţʴȦȌţú4̗ȏĤħ1Ĥ˧͍Ĥ˧˕ǥɗɣīĎ͋5 Ø1 1Ğų͍͌Ĥ˧˕ǥƤ͍35 pp.
Kamijo, T., Isogai, T., Hoshino, Y. and Hakamada, H. (2001) Altitudinal zonation and structure ofwarm-temperature forests on Mikurajima Island, Izu Islands, Japan. Vegetation Science, 18(1):13-22.
ÌȻ»đɃ;)ŧ Ƒðʰė͋2002͌ȋ̱0CŖÍɠ1̇¢ǵɠ
1¸ÍŌɔ0ú9Ɖ̳͍ǂǗʈíūŅ©ˏ͍28(1)͓139-142.

ŭǵ1©ʊ͋2004͌ĤȽGňC͋#1 1͌Ǝʲţ0CʟȭěƏ1ōˮ͍ŭǵ1©͍48 pp.
Loope, L. L., Hamann, O. and Stone, C. P. (1988) Comparative conservation biology of oceanic
archipelagoes –Hawaii and the Galapagos. BioScience, 38(4): 272-282.
̒ʻć˂ʖƈ͋2004͌˂þB˕ǥ0@CņţÙġȋ̱ʈí1ǭɀưɞ͍ǂǗʈíūŅ©ˏ͍30(2)͓383-388.
ȈɃ ɓ͋1956̬͌ʭʐʯ1Ɉ͎̆ɪǸ̈ɞ1ÛǓǿ̧1ˇǟ ͍ǂǗɀƠŅ©ˏ͍6(2)͓
62-66.
ȈɃ ɓȑ̒ˡĸ͋1969͌ǂǗǭȳɀƠĞ̛ ʇ˖͍ɳĤǎ͍̿39 pp.
ĶĬ̆͋1971a͌Ǝʲţ1ǭɀ͍ɛĻũɒɧôȳ̿ɗɣīĎ͍1(4)͓1-53.
ĶĬ̆͋1971b͌rNdXyUiˀɺ1ǭȳ͍ɛĻũɒɧôȳ̿˕ǥɗɣīĎ͍2͓1-63.
Š̒ē̌ʮȥ˳ĸ̜˝ũÅǚɃʘƚ̔ǔ͇ʏŁ ǋįǚæ (2005) Ǝʲţ1Ą̼ʺŏŦİĤ0CǭɀěƏ ()͎Ķʡƣ͐ĆŦİĤ0C˕ǥɾǡ͎͍̎ǬǠɗɣ͍53͓11-12.
ŗ̒Ŷ̪͋1994͌ńţ1ɀȳ%'͍šȌǎż͍239 pp.
ƔɂʀˈmhsdTʊ̫ŀĒ©ʊ͋1978͌ƔɂʀˈmhsdT͍̾˦Ī͍827 pp.
ʿǪĽƀɖũ̆ʦ͋1986͌̅˭ȋ̱1ǭɀɱȾ0̠Cɗɣ͋͌Ôșűĵ̰ħ1͂ɍġȋ̱0C 10 ŵ̟1ǭɀ1Ĳí͍ş̡ĶŅ˶Ņ̎ɗɣīĎ͍51͓363-374.
͈ʳɜǵ͋1990͌ǽġÖȾĬ0Ƣɧ%OOn~Z}sZŷ͉ʐʯ1ȳɀɁ1ȴƐ͎0ʍǾĘŋ0)+͎͍ǂǗǠŅ©ˏ͍72(3)͓208-215.
°ʳ Žʿǚ Ǩ͂Ƕǹƃ͋1965͌ʭĤ0Cɥɺʄƫ0̠
1 ī ɥɺ1ĵɂȲʭĤ1ý̓-ȅʃƘ0ú9CɀƠɀȾŅɋɗɣ ɪƉ̳͍ǂǗʭĤŅ©ˏ͍11(1)͓14-19.

ȢɃ³ŊɦɃģĵÏ£ʻ̌ʿʜ ƟȎɃ Ǎ͋2008͌ȟˤɒ0CʢɤōƳ
ū0@CȲʭƳɠ/FD%ŞȧˬĤ͋ǸǠȮĺĤ͌1ǭɀ͍¼ÎɀƠŅɗɣ͍
13(1)͓29-36.
ĂǔʤĚƥʅʪ͋1963͌ƹ̓ɋ̟¨0̠īĎ͍34͓1-63.

45

CɀƠŅɋɗɣ͍̏ĶŅ˶Ņ̎ȠʒǠŇɃũƃè̒øɵȤɃȒʏ͋2002͌Ǝʲţġı1ȾíŅƘ͍ǂǗġıʗƽŅ©˘ȠˀǄ̫͍48͓220.
·Ƀƙ͋1971͌ǬǠ1ɀƠŅ͍ɳĤǎ͍̿331 pp.
ĉũèɜ͋2006͌ɀƠŅɋ/ƾ̱1Ḇūȋ͎DA1ʈíƨʶ͍ŞȓĪ͍208 pp.
(ˢ) ǂǗȃˠò©ʊ͋1996͌1996 ŵȱ ȃˠŵ̛͍Ķʲɑöàś͍265 pp.
Zhou, J., Naito, K. and Takahashi, Y. (2002) Secondary vegetation and succession of abandoned
grazed grasslands in upland Southwest Honshu, Japan. Vegetation Science, 19 (1): 11-23.

46

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