Vol. 4 (1): 77'88

ORIENTAL INSECTS

I\Iarch 1970

INDIANCRANEFLIESNEWANDLITTLE-KNOWN(DIPTERA: TIPULIDAE)_III
By CharlesP. Alexander
39OldTot;ttRoad,Amherst,Massachusetts,U'S'A'
India are described,these
Assrnlcr. Five new speciesof Tipulidae from South(Xipholimnobia) madrasensis,
being Tipula (Indotipula) tetradolos, Xipholimnobia
and S' pentachaeta' In
Gonimyfi (dioceral megastigma, Styringomyia monochaeta,of several previously described
addition figuresof the male hypopygiumand venation
to the genusToxorhina
belongingspeciesoi Iodiun cranefliesurc prouid.d, virtually all
Loew, none of which had beenillustrated previously'

3:part underthis generaltitle appearedin oriental Ins''
Thepreceding
five new speciesof craneflies'
1-13,March 1969. At this time I am describing
by Dr. Fernand
colrected
all from variouspartsof south India wheretheywere
expressmy sincerethanks
Schmidand Mr. p. SusaiNathan,to whomI again
Tipulidae of the Indianfor their appreciatedefforts to nrake knorvn thefauna.
Ialsoamprovidingfiguresthatshowthemalehypopygiumandvenatio
Thereremainsomeof severalother speciisihat neverhavebeenillustrated'in this greatfamily of flies that never have been
hundredsof still otherspeciesof papersI hopeto beadequateryfigured und-in subsequeitpartsof this series
ableto treatmanyof thesein similarmanner'
the yearslgll and 1924'describedsome
ThelateEnricoBrunetti,between
andBurma'in this family of fliesfrom the formerBritishIndia
240 new species
writer (Rec' Indiqn Mus''thesehavingbeenlistedin an earlier paper by the
speciesappearto havebeen
59 : lg-34, March 1963). A mu;oiity of these
there remain a considerable
describedsatisfactorilyand are reclgnizabrebutof Brunetti'sfailure
becausenumber of obscureformsthat arein doubt,chiefly
appears
andfigurethe malegenitaliain thesecases' It
describe
to adequately
may proveto be a serious
speciesthat our inaUiiity to recogiizesuchdoubtful
h a n d i c a p t o l a t e r w o r t o n t h i , f a u n a . I t i s v e r y d e s i r a b l e t htreateda t i n t andhespecies
carefully
removed'beshouldhypopygium
in questionthe maleslide' such slides
a microscope
dissected,and then *ount.o p.rtun,ntly onof the doubtfulforms but
would not only helpto providecertaindetirminations
than arepinnedmaterials'
arelessliableto damageor destruction
PublishedAPril 1970

78

ORIENTAL INSBCTS

Vol. 4, No. I

Sunrluny Trpurwml.

Tipula (Indotipula)tetradolos,sp. nov. (Fig.

3)

closely related to beringana,differing chiefly inhypopygiar structure,includingthe tergiteand innerdististyle,th; Iatterwith four spinoidsetaeon thedorsal crest.
Male"rFrontalprolongationof headlight yellow,
sidesrestrictedly
darkened,
nasuslong,tippedwith conspicuous
black ietae,proximaltwo segmentsof palpi
brownish-yellow,
remainderbrown, outer end of terminal segmentyellowed.
Antennaewith proximal three segments
obscureyellow,remaindervery weakly
bicolored,Iight brown,basalenlargements
darker brown, outer segmentsmore
uniformly darkened ; flagellar segmentsmuch
as in belingana,the outerenlargementsubtriangularin outline, verticils ofouter segmentsvery long.
Headwith front whitenedwith a restrictedbrown markbetweenthe antennalbases; vertex more brownish-yellowwith a very
narrow darkenedmedianline.
Pronotumlight yellow,sides narrowly darkened.
Mesonotalpraescutumorange' disk rvith four narrowlight brown stripesthat are vaguelyborderedby
darkerbrown,sidesof scleritebroadlylight yellow; scutumwith centerof eachlobe obscureorange,marginedinternurryuv iight bro*n; scutellumlight bro'n,parascutella
brownish-orange; mediotergiteyellowed on central part, with avaguepalebrownstripeon eithersideof midiine,posterior
third -o.. uniformly
light brown, pleurotergiteyellowed,katapleurotergite
uro*,. ;;;;;#,
light yellow,rvith brownareason propleuru,un.pisternum,
u.n,.ut ,il;;;'ffi;
and ventral meron. Haltereswith stem brownish_yellow,
knob small,darker
brown' Legs with fore coxae light brown, remaining
coxae light yellow;
trochanters
yellow; femoraand tibiae light brown,tarsi darker
brown; terminal
tarsal segmentat base with a conspicuous
epicondylar
brushof setae; outer
tooth of claw long. Wingsbrownish-yellow,
pre-arcularand costalfieldsclearer
yellow; stigma light brown ; veinslight brown. venation: cell 2nd Aslightly
narrowerthan in belingana.
Abdominaltergitesbrown,posteriorand lateralborders
narrowly yellowed,
sternites
yellow; hypopygiumwith the large eighth sternite
brown, conspicuously patternedwith obscureyellow ur lo certain otheralliedspecies,
tergitechieflyyellowed' Male hypopygium(Fig. 3) generallyas
in telingana,differing
especiallyin the tergiteand innerdististyle. Niottt tlrgite, r,with the blackened
outer end restrictedin area, the lateral projectioor-*id.ly
separatedby a
U-shapedemargination,lateral spinesfew. Innerdististyle, ,1,*ithfour setoid
spineson dorsalcrest,theseunusuallystout posteriorlobe;
of style very large.
Eighth sternite,8s, rarge, chiefly darkened,with a vague
yelrowedpatternan
outer half includingthe posteriorborder,

1970

ALEXANDER: ON TIPULIDAE

79

about3.5mm'
Length.'j, about15mm, wing15 mm, antenna
Holotype6f,INorl : Cherangode,Nilgiri Hills, South India, 3500 ft. xi.
1950,Coll. SusaiNathan.
Atthoughthe presentfly is closelyrelated to Tipula (Indotipula)belingana
venation;figs'47,48, male
Alexander(PhilippineJ. Sci.,94 : 274-276,frg.21,
describedfrom SouthCoorg,South India, I consider the
hypopygium; 19,65),
as
two flies to be distinct, especiallyshown by the hypopygialcharacters,
described.
SunrmltlY LnloultNAE
Intgs LEcsnttNt
sp. nov. (Figs.l, 2)
madrasensis,(Xipholimnobia)
2. Xipholimnobia
General coloration of mesonotum brownish-yellow,pleurotergiteand
pleura clear yellow, the latter without a darkenedstripe; wings weakly
iinged with brown, broadestoppositecell lst A ; male hypopygiumwith"p[r, of interbasesvery obtuse; outer dististyle with base expanded,
the slenderlateral ann with two or three small spinesbeforethe terminal
point.
thewith scapetestaceous,Male.. Rostrumbrown ; palpiblack. Antennae
the segmentslong-oval' Headenlargedpedicelblack,flagellumbrownish-black,light brownabove,narrowlylight grayon orbits'
pronotum brownish-yellow,pretergiteswhitened. Mesonotalpraescutum
light brownish-yellow,pale laterally; scutal lobes slightly darker brown,darker posteriorly,parascutella; scutellumtestaceous,,*aiun regiontestaceous
yellow; mediotergitebrownish-yellow,pleurotergiteand pleura clear yellow,
th. lutt., without a darkenedstripe. Haltereswith stem yellow, knob lightyellow,remainderlight brown. Wings
brown. Legswith coxaeand trochanters(Fig. 1) **t ty tinged with brown, costal regionmoreyellowed; veinslight
Ur*o. Wingsof malebroadestoppositecell /sl z{ shortly beyond terminationof vein 2nd A. Longitudinalveinsbeyondcord with trichia, includingalso theoutertwo-thirds of Rs, lacking on both anal veins. Venation: Sc, ending
about opposite five-sixthRs, ,Sczslightly removed; m-cu at neartwo-thirds
Mr+n'(Fig' 2)
Abdomenlight brown,hypopygiumyellowed. Male hypopygium
tip very obtuse. Outerdististyle,d,with the interbase,f, nearlypaiatGt-sided,
lateralarm slender,its outerendgentlycurved, terminatingwith baseexpanded,
beforeapex;
in a subacutepoint, with two or threesmallspineson outermarginapexwith ainner style nearly as long,slender,narrowedoutwardly,at endnear
qmalltuberclethat bearsa long bristlg'

80

ORIENTAL INSECTS

Vol. 4, No. I

Length.'J, about 2.5 rlrl, wing 3 mm.
HolotypeJ, INota: VaraiyattuTittu, Madras, 1500ft., 21.xii.1958,Coll.
Schmid.
The regionalXipholimnobia(Xipholimnobia)(Alexander)has theumbripennfs
wings narrowerand much darker,the thoracicpleurastripedlongitudinallywith
darker,and with the hypopygial details, especiallythe outer dististyle, quitedistinct.
Tnrse EnloprsnNr
3. Gonomyia(Idiocera)megastigma,
sp. nov.
Generalcolorationof thorax gray, praescutalstripesdarker brownish-gray,pleura with a conspicuousventralyellowstripe; legsyellowed; wingsfaintly
yellow,stigmalarge,dark brown ; macrotrichiaof veinssparse,on veins/?o toMs,lackingon Rs and the anals.
Female.'Rostrum black ; palpi dull black. Antennaewith scape darkbrown above, light yellow beneath,pedicel yellow, flagellum dark brown;
proximalflagellarsegments
oval, the outer ones progressivelylonger, subequal
to their verticils Headlight gray, front and anteriorvertexyellow.
Pronotumgray,narrowlyyellowedlaterallyand on pretergites,sidesbrown.
Mesonotal praescutumlight gray with broad intermediatedarkerbrownishgray stripes, humeral and lateral borders light yellow,pseudosutural
foveaeshiningblack,conspicuous; scutumgray,posteriorcallositiesobscureyellow;
scutellumbrownish-gray,posteriorborderobscurebrownish-yellow; postnotum
dark gray. Pleuradark graywith a conspicuous
ventrallight yellow longitudinal stripe, extendingfrom behind the fore coxaeto baseof abdomen,behindmore expandedto includeparts of the metapleura,ventralpteropleuriteand resdorsopleural,trictedlyon pleurotergite;
regionyellow. Halteresrvith stemyellow,
knob palebrown. Legswith forecoxaeand trochantersdark brown, remaining
yellow; femoralight yellow,
coxaeyellow,basesnarrowlydarkened,trochanters
tips narrowly very pale brown, remainderof legs yellowed. Wings faintly
yellowed, pre-arcularand costal fields clearer yellow ; stigma dark brown,
unusuallylarge,subrectangularin outline,reachingcosta in front, behind vein
Xs i vague darkeningsat arculus, origin of Rs and the cord, indicatedchiefly
by darkeningof the veins,remainingveinslight brown, costa brownish-yellow.
Macrotrichiaon veins Rn, Ro and all branchesof M,lackingon Rsand theanals. Venation: ,Sc1ending opposite origin of Rs, S., far retracted, Sc,
alone about four-fifths Rs i Rz+s+rshort, about one-thirdRs, cell R3relatively
small ; cell 2ndM2 shallow,lessthan its petiole, the veins slightly divergent,wideningthe cell at wing margin.
Abdomen dark brown, extreme posterior borders of the sternites
yellowed.
Length.' ?, about6,8-7glr, wing 5.5-5.7mm.

1970

ALEXANDER: ON TIPULIDAE

8l

Madras,1500feet,l.xii'1958,Coll' Schmid'
Holotype9, Inota,: Sathuparai,coll. Schmid.
ParatypeQ,Nagodi,Mysore,2500feet,28.i.1959,
The most similar regionalspeciesis Gonomyia(Idiocera) recensAlexander'of thelikewisefrom SouthIndia, whicirin the femalesexdiffersin the venation
wing veins,outer radial and medialfieldsand in the abundanttrichiationof theon bothincludingall longitudinalveinsbeyond origin of Rs and extensively
anal veins.
sp. nov. (Fig' a)
4. Styringomyiamonochaeta,
Allied to kempiana,differingespeciallyin hypopygialstructure'pedicelMale.' Rostrum and palpi black. Antennae with scape andslightlybrownish-black,basal flagellar segmentyellow, succeedingsegments
variegateddarker,outer onesbrownish-black. Head dark brown, restrictedly
with obscureyellowon anteriororbits.
with
Pronotumchieflylight yellow,darkeron.sides. Mesonotalpraescutum
brownish'black,scutum similarly dark
threedark gray strip.r, 1n. interspaces
obscuregray; posierior scleritesof notum darkened,scutellumvariegatedwith
yellowed vitta. Pleura brownishletiow, mediotergitewith a vague central
slightly
black. Halteresinfuscated. Lels with coxaedark brown, trochanterspaler ; femora brownish-blackwith a narrow obscureyellownearlycomplete
a subtermiring beforeapex; tibia similarlydark brown,the extremebaseandproximalhalf ofnal ring yellow; fore tarsi brownish-black,mid tarsi withproximal threebasitarsiyellowcd,posteriortarsi moreuniformly pale, tips of
Wings
segmentsnarrowly darkened,outer segmentsuniformly infuscated'in casesthe basal fourth clearer
to palebrown,unpatterned,
brownish-yellow
yellow ; veinslight brown,moreyellowedin the basalfield'
as in kempiana,
Abdomendark brown. Male hypopygium(Fig, 4) generally
in the structureof the complex dististyle,the major outerdifferingespeciallythe narrow innerarm with asetiferousarm broad and entire, as in kempiana,
in kempiana
singlevery largeblack spinoidsetaon margin at near midlength,with five or more' Other
with two much smaller such setae,in pentachaeta
structuresof the hypopygiumabout as shown'
Length.'$, about7.8-8illrl, wing 5-5.5mm'
feet'
Holotypeil, Ixptl : Naduvatam, Nilgiri Hills, South India' 6000
with type ; I d'
20.v.1950, coll' Susai Nathan' Paratopotypes7 dd
ix.1950
Formerly I had placed this fly with styringomyia kempiana Alexander,that infrom the North Kanara District, South India, but it now appears
similarbut distinct species,including theSouthIndia thereare othergeneralty
sp.nov.' al! nolt readilytold amongthenppresent fly and also ,S.pentachaeto,selvesby the hypopygialstructure.

.l

82

ORIENTAL INSECTS

Vol.4 , No. I

5. Styringomyia
pentachaeta,
sp. nov. (Fig. 5)
Allied to kempiana,
differingespeciallyin hypopygialstrucr.ure.
Male.' Foreand middlefemorachieflydark brownwith a narrow
incomplete
obscureyellowring morethan its own lengthbeforeapex tibia
with base and;a subterminalring narrowlyyellow; fore tarsi blacl, mid tarsi
with proximal
two segmentsobscureyellow, tips broadly brownish-black,ourer segments
uniformly black ; posterior femora obscuiebrownish-yellow
with an obscureyellow subterminalring, the darkenedtip slightly brtader; tibia almost
uniformly medium brown, the pale rings diffuse to scarcely
evident; tarsi
obscure yellow, tips of the individual segmentsslightly
darker. Wings
brorvnish-yellow,
unpatterned,
pre-arcularand costalfieldslt.ur., yellow veins;
Iight brown. Venation: Cell 2nd M, short-sessile
vein
2nct
A curved very;
gentlyto margin,the cell narrow.
Abdomenbrown. Mare hypopygium(Fig. 5) generailyas in kempiana,
the basistyle,b, trispinous,one spinemuchimaller. oististyle,r/, distinctive,
onearm deeplyemarginate
to form two narrow blades,each with long black
setae; inner arm shorter,with similar blacksctae,its marginwith fivemore
powerfulblackbristles,the outermostshorterand slightly stJuter,lower pair
Iongest,certain of the remainingsetaeon disk virtuallyas longandsimilarly
conspicuous
outerrod of stylewith numerousblack setaeat base; elongate
includingcomb
aof aboutl0 stouterspinoidsetae.
Length.'j, about6 mm, wingabout4.2 mm.
lolqtype -d, INola : cinchona, Anamalai Hills, South India, 1067m,
v . 1 9 5 9C, o l l .S u s aN
i athan.
The deeplyemarginatearm of the dististylediffers from the condition
found in ,styrrirgomyia
kempionaAlexanderand s. monochaeta,
sp. nov.
6. Styringomyia
tarsatraAlexander(Fig. 6)
st;ringomyiatarsatraAlcxander,1966,J. Newyork Ent. soc.,7l: lg3-1g4.

Typesfrom Parewavirand Amlekhgang,
Nepal, 1957,coll. coher.
Male hypopygium(Fig. 6).
7. Toxorhina(Ceratccheilus)
bistylaAlexander(Figs7, ll)
Toxorhina (Ceratocheilus) bistS,/cAlexander, 1967, J,I 84-l 85

NewYork Ent.

Soc., 7 5 :Typ, J, Bilo La, Kameng,North.East Frontier Agency,Assam, 6000 ft,
l0.vi.1961,Coll.Schmid.
Venation(Fig.7), malehypopygium(Fig. I l).

1970: oN TIPULIDAE
ALExANDER't

83

8. Toxorhina(Ceratocheilus)
brevifrons(Brunetti)
Conithoraxbrevifi'onsBrunetti, 1918,Rcc. Indian Mus.,15 : 300.
Toxorhina (Ceratocheilus)brevifrons: Alexander, 1936, Philippine ,I. Sci., 6l:(re-descriptionof type e ).
Toxorhina (Ceratocheilus)brevifrons: Alexander, 1952, Rec. Indian Mrs., 50:
fig. I5 e (venation).

202
357.

9. Toxorhina (Ceratocheilus)
fulvicolorAlexander (Figs. 8, 12)
Toxorhina (Ceratocheilus)fulvicolor Alexander, 1967, J. New York Ent. Soc., 75:
185.

Type d, Khaorum, Manipur, Assam, 3750 ft., 28.viii.1960,Coll.
Schmid.
(Fig. l2).
Venation(Fig.8), malehypopygium
fuscolimbata
10. Toxorhina(Ceratocheilus)
Alexander(Figs.9, l3)Toxorhina(Ceratochcilus)
fuscolimbataAlexander, 1967, f . New York Ent. Soc., 75:
r85-186.

Type d, Hkyam Boum, Manipur, Assam, 7500 ft., 20.vi.1960,Coll.
Schmid.
(Fig.l3).
(Fig.9), malehypopygium
Venation(Figs.10, l4)
Alexander
11. Toxorhina(Ceratocheilus)
luteibasis
Toxorhina(Ceratocheilus\luteibassAlexander,1962,Ent. News,73: 12+-125.

Coll.Schmid.
Typej, Chateng,
Sikkim,8700ft., 22.v.1959,(Fig. l4).
Venation(Fig. l0), malehypopygium
mesorrhyncha
Alexander(Fig. l9)
12. Toxorhina(Ceratocheilus)
Toxorhina(Ceratocheilus)mesorhynchaAlexander, 1936, Philippine J. ^Scf.,6l : 200201,pl. l, fig. 24 (venation),pl. 2,fig.34 (male hypopygium).

Type d, Cherrapunji,Khasi Hills, Assam, 4000 ft, viii.l935, Coll.
Sircar. Further material from Lathong, Sikkim, 6560 ft.,26.vii.1959,Coll.
Schmid.
The following descriptionand figure are more accuratethan those provided
earlier. Male hypopygium (Fig. 19) with mesal face of proximal end ofbasistyle,6, producedinto a conspicuous
tuberclethat is tufted with about 12
strong setae in a compactgroup. Interbase,i, unusuallyslender,especiallyat
base. Dististyle, d, single, beak long-produced,narrowed gradually to theobtnse tip, on its face u,ith a long low flange; outer margin of proximal third
with a slender tubercle. Arms of aedeagus,e, rglatively long, slender,tips
gentlydivergent

84

oRTBNTAL
rNsgcrstuo,. 4, No. I

13. Toxorhina(ceratocheilus)
monostylaAlexander(Figs. 15, 20)
Toxorhina(Ceratocheilus)
monostylaAlexander,1962,Ent. News,13: 123-124.

TypeS, Dikchu,Sikkim,2300ft., 9.v.1959,Coll. Schmid.
Venation(Fig. l5), malehypopygium(Fig. 20).
14. Toxorhina(Ceratocheilus)
simplicistylaAlexander(Fig. 22)
Toxorhina(Ceratocheilus)
simplicistylaAlexander,1967,f. New York Ent. Soc., 75:
186187.

Typed, Nakhu, Kameng,North-East Frontier Agency, Assam, 4800 ft.
3.vii.1961,Coll. Schmid.
Male hypopygium(Fig. 22).
t5. Toxorhina(Ceratocheilus)
tuberiferaAlexander(Figs.16,21)
Toxorhina(Ceratocheilus)
tuberifera Alexander, 1966,J. Nr', York Ent. Soc.,74 :184.

TypeJ, Lathing,sikkim, 6560ft. vii.26.lg5g,coll. Schmid.
Venation(Fig. 16),malehypopygium(Fig.2 t).
16. Toxorhina(Toxorhina)brevirama
Alexander
Toxorhirta (Toxorhitna) brevirann Alexander, 1953, Artn. Ivlag. Nat. Hist., (12) 6 z

t9t-192.

To.torhina(Toxorhina\
breviranu:Alexander,1953,Rec.IndianMus.,50: 357,fig.15 f(malehypopygium).

Type d, Naduvatam,Nilgiri Hills, South India, ix.1950, coll. Susai
Nathan.
17. Toxorhina(Toxorhina)scitaAlexander(Figs.17, 23)
Toxorhina(Toxorhina)scitaAlexander,1962,Ent. News,73z 126-127.

Type$, Cherangode,Nilgiri Hills, South India, 13.ix.1950,Coll. Susai
Nathan.
Venation(Fig. l7), malehypopygium(Fig. 23).
18. Toxorbina(Toxorhina)sparsiseta
Alexander(Figs. 18,24)
Toxorhina(Toxorhina)sparsisetaAlexander,1962,Ent. News,73 : 127-129.

Type6f, Cinchona,Anamalai Hills, South India, 1067 m, v.1959,Coll.
SusaiNathan.
lgnatioq (Fig. l8), malehypopygium(Fig. 24).

ALEXANDER: ON TIPULIDAE

le70

85

.t

3

is/
rl
i:
\1.

: i
: I
t:

5\

sp. nov' Figs'
Figs. l-6. venation of xiphoti,,;nobia (xipholinmobia) madrasensis,
sp' nov'; 3,2-6. Male hypopvgiu* of 2,Xipholimnobia(Xipholimnobia)madrasensis,
SP' nov' ; 5,
Tipula (Indotipula) letradolos, sp. nov. ; 4, Styringomyia monochaeta,: 6t(Symbols
Styringomyia pentachaeta, sp. nov. ; 6, Styringomyia tarsapa liexander't' tergite)'basistyle; d, dististyle ; i, interbase; p, phallosome; s, sternite;

86

ORIENTAL INSECTS

Vol. 4, No.

t

r3

t4

Figs. 7'14. Venation of 7, Toxorhina (Ceratocheilus) bistyla Alexancler; g,Toxorhina (Ceratocheilus) fulvicolor Alexander ; 9, Toxorhina (Ceratocheilusj
fuscolimbala Alexander ; 10, Toxorhina (Ceratocheilus)luteibasrs Alexander. Figs.
ll-14. Male hypopygium of ll, Toxorhina (Ceratocheitus)bistyla Alexander; iZ,
Toxorhina (Ceratocheilus) fulvicolor Alexander i 73, Toxorhina (Ceratocheilus)
fuscolimbalaAlexander ; 14, Toxorhina(Ceratocheitus)luteibasl'sAlexander. lSymbols:c, aedeagusi b, basistyle; d, dististyle; i, interbase).

r970

87

ALEXANDER: ON TIPULIDAB

.?

(-

r9

2l
23

24
monostvlaAlexander; 16,
15, Toxorhina (ce^ratocheilus)-i,,iuifrro.'ei.*ioori
Fies. 15-24. Venationofscita
i r7.,.toxo.rhiia_(roxorhina\
roriifiio'" <tirotiiiliiiii'i;;';;ii^t"'liiiiini,1i7.
Male
Figs'-19-24'
Alexander.
sparsisita
Alexander. 18,; 20, Toxorhina
oijs,-iiiiliino -etllinde-r';-ijeratochiilrli ^r'orhvnc.haAlexandefi')ffi;;;iilnberifera(.Ceratocheilys-)
Tgx{rhi.na
Zi,(Ceratocheitus) ^;;;;i;roxorhina
23,
itmpticistyta--iixorh.tna,Alexander;hl;ffi,i;;;'-iz,'foiiiiiio'(d;'aigieitustsparsisetn(T'oxorhina\.Alexander.
(Toxorhina)scita fidii'nari-- .z+,.
p, phallosome;i,interbase;dististyle;d,fo,-iia"?eil;T:'uasistyie
is;;6il;'l, tergite).