Sstl

DIPTERA COLLECTED ON SOUTEAIIPTOIV ISTANDBY GEORGE MIKSCE SIITTONTRICEOCERID.IEAIVD TIPI'IID.T

By Cuenr,nsP. Ar,ExaNDDnl{assachusettsState College,Amherst,lVlassachusetts

R,EPBINTEDFBOM TEE MEMOINS OF TEE CARNEGIE MUSEUM
voIJ. xII, PABT rr, gEgfloN 4
IssusdJune18,1984

.

II.

DIPTERA COLLECTED ON SOUTHANIPTON ISLAND BY
GEORGE MIKSCH SUTTON.
TRICHOCERID,O AND TIPULID,E.
Bv Cnenr,psP. Ar,pxeNopn.
Mess,c.cHUSETTSStenp Colr,ncn,

Ausnnst,MessecHUSETTS.

Through the kind interest of Dr. William J. Holland, I have been privilegedto study
the Crane-fliescollectedon Southampton Island by Mr. GeorgeMiksch Sutton and now
preservedin the collectionof the CarnegieMuseum.
The presentreport is divided into two divisions,the first of which considersthe Craneflies securedby Mr. Sutton on SouthamptonIsland, the seconddivision being a discussion,
with list and bibliography,of the Tipulidre of the CanadianArctic.
All types and uniquesresulting from the presentinteresting seriesof Tipulida are preservedin the CarnegieMuseum. I wish to expressmy indebtednessto Drs. Holland andAvinoff, and to Messrs.Kahl and Sutton, for the opportunity of studying this relatively
extensiveseriesof Arctic AmericanTipulidn.
DrvtslorvI.
THE TIPULID,A AND TRICHOCERID,O OF SOUTHAMPTO}{ ISLAI{D.
Family TRICHOCERID,E.
GenusTnrcHocpneMeigen.
1. Trichocerahiemalis(de Geer).
de Geer;M6m. pour serv.i l'Hist. desIns., VI:360, pl.21, figs. 1-5.
1776. Tiputahiemal:isSouthamptonIsland, 1930(Sutton);5 specimens.
The fly had beenpreviouslyrecordedfrom Greenland.
Family TIPULID,fl.
SuereMrr,vTr pur,rw,o.
GenusPnroNocnneLoew.
2. PrionoceraelectaAlexander.
1927. PrionoceraelectaAlexander;CanadianEnt., LIX: 188-189.
SouthamptonIsland, July 3, 1930(Sutton);19 .
Describedfrom Hopedale,Labrador (Lat. 55rh" l{.). The presentfemalediffersfrom theunique type, a male, chiefly in the short-petiolateto nearly sessilecell M, of the wings,but
certainly appearsto representthe same species. The unusually broad and conspicuous
obliterative streak before the cord, in conjunction with the infumed outer radial field andshort upcurvedvein R, distinguishthe presentfly from allied regionalspeciesof Prianocera.

ed June 18, 1934.

MEMOIRS OF THE CARNEGIE MUSEUM

Genus NppunotoMA Meigen.

3. Nephrotomalundbecki (Nielsen)'Nielsen;Meddeielserom Grlnland, XXIX: 390-391'
1909. Pachyrrlina Lund,becki
Arctic Exped. 1913-18,vol' III,
1919. I{ephrotomi'irri"oto at""uid?;^R6il-Canadianpart C: 10e-1lc.
SouthamptonIsland,box closedAugust 1, 1930(Suttun);19'
from the Canadian
The type of.tund,beckiwas from East Greenland, that of arct;icol'aextensivegap in the range
Northwest Territories. The presentstation bridgesthe hithertoof the fly.
GenusTrPur'e Linneus'
4. Tipula (Vestiplex) arctica Curtis'
trome by commander J'
1g31. Tipula arcticacurtis; Description.of the insectsbr-o.ught
p. lxxvii'
Clark Ross. Ross'sVoyagbto the^Arcticregioll, Ap2endi2,

Hist.,(10)VIII: 80'
1981. TiputnfiitiriU:"t";frti;; na*utat; An"n.M'ag.'Nat.

Island,July 3, 1930(Sutton);7,&, 9'Southampton(Alexander,1919' 1925;
widely distributed in the Arctic Regions'of the New World.
Recordsfor Arctic Europe
Curtis, 1831;Henriksenand Lundbeck,1917;Nielsen,1907)'*
requireconfirmation.
5. TiPula diflava Alexander'
1913-18,vol' III' part1919. Ti.pula d,iftaaaAlexander;Report canad,ianArctic Exped'
C: 12c.
SouthamptonIsland, July 3, 1930(Sutton);8, c/ ' 9 'Describedfrom the CanadianNorthwest and Yukon Territories'
6. Tiputa hollandi sp. nov', (Plate XXV, figs' 1-5)'
clearergray stripes,which
Generalcolorationyellowishgr&y, the preescutumwith three the entire length of mesoare borderedby brown; a capillar?-meai""bryryn nittu extending*lt* frt"iiv u.t*nish, vari-egatedby dark brown andnotum; antenne fr4ii'trrr*igtr""t'vatoit', with three continuous'longiwhitish subhyatineareas;ubdorrrirtli;.igtt*:;b!#;margin of tergite with a broadtudinat black stripes; male n'popiliit;: *ih tnp *"aal sterniteunarmed'
U-shaped.rot.rr; tultiltvi.^pr.ir'y-r-"tEawith sternite; eighth
Uitn: Length,about 12-13mm'-i*itg, 13-14'-i;i"""* in. iength.to remainderof head, light gray;
Frontal proro"gutio" or heud-al]o"T-.'q"ut
prui-

nasusstout;pur;;Tlu;k rili,Eh";;.

the basalsegment
uiu.[ throggho.ut,

6ururenlargementonly feebl-ydeveloned:terminal segment^;[;;;;;
nose;flagellar.d;;tr;;ilth%tnu" the segniells. U.ia btoi'nish g?v,' somewhatclearervery smafr; ,ru*?Jifi
medianbrown vitta'
grav on front and on the narro* p"rt.ri6? oruitr; u rrotr*Iy-indieatid prescutum. yellowish
til;. Nlesottotal
pronotum gray,with a .r,pi[ui]"Lro*"-r""di;;are borderedbv brown lines, these*hich.,ririir"*r;;.;t'ridr
grey,with thre;il;irrt,-*pittrty bto*tt vitia extendsthe entire
being narrowerin the paratypetnarr-irr'tnpti'pui'uro*nu..ur. pleura clearBr&y,the dorsolength of mesonol;; ;;;h 99"1"^ir"t; *itt i"*ovellow.
tti" r."obsdarker,tippedwirh obscure

preurar*.*u.uirJu"rii.- ii;fi;;"ffy,
the tips
cox&.r;d6;h;;t"rr rignt-grav;l"q"id fulvous-vellow,
fffi i;#";;d' slender;.
wings
small.
very
claws
2;
2tibieeu.ratulri^;1";[;";i[ur'rp"is tbroadlvbrackened;*Dates in parenthesis refer to Bibliography at end of paper'

ALEXANDER:DIPTERAcoLLEcTEDoNSoUTHAMPTONISI,AND5(Pt.xxv,fig.l)withtheground'-colorfaintlybrowlish,variegatedwithdarkerbrownand
origin of Rs, end ofi".r"ortrg'smarl"spotsa,tdarkei;*ili;gq
theareas,
subhyarine
whitishitrJr"t"rioi.ord : furrher dark areasino"tou.;;;;;"dia;;r,agsb.t
th;
sc and stigma,*n c; ;;i;;;ia-reirqth; rhewhitishsubM oneithe*td;;i ;;hdi#il;;"a1"cerrU;; as a narrow'incorioletecrossband beand as ahvalineareaslie beforeand beyoniihiltfii;;Tspofat nog.,tJJ-trrira. tn6 lengttrof cell M, sc more
ybnd the cord,as a large,conspicuous'^Uoriutar,;"u.?tt;ffii
commonarea,dfi

brown,
Jil"ri.^c" ;"4 t;t A; veins-dark
beyondcord veryt"u.rot.ichia of veinsyerowishbrown.
tilg; ;ffi and dense;oi Rn*uanda nearlvcomplete
ioarse,including,;;;i", oi-u6or.t'i:#;;-Jryd.".ii""
Rr+zcorrplete,with from sixto
cur;-.qui*i-itrr."aof.
-y"outioor
sectionsbrieson distal"t ."ir M,,;f;q"al-to m; m-cuat fork of Ms+q'
ditr"
macrotric#;;t;;uin"ir;eiehtyellow, trivittate with;"t$ie;- obsc-ure
Basal abdominartergite gruv,'"it".-..ri...ai"g.
blackmediallv;outersegments,
brownishbrack;.T.r-"it"J"usi,r1gy;ii;*5;il1y-b;-r'ftri.tt
tp,r[ nnrp hvpopvgiut"(pr. xxv, fis. 2)
includinghypopygirlm,moreunifor#ii'ryril-rrsternite-bv-paleniembia"e:basistylenot comseparated?r"'rr-trrl
entirerytergitethewitho" ttt" dorsal' and a little lessthe sutures i"li;fi'dpletely separated frorn- the sternite,

areais cut
roughrytriansurar
onthe ventrSrportionil il;il;;;r[;.r""gut.,
dark.grav,
extensivery
sl
tis.
xxv.
Nirrh';?isi6 iFi
thedJrJai;ilil.
fromthe sterniteadjoiningof ripuln;
species
Arcticth.^;;;"i"-..".*t
arrdpori.n""dtiu.[;;rl.
not heav*y screrotized
u-.rtup.J "oi.rt; c.audalthiid of tergitewith
margihyitf;;;^t;9rd.
the caudar
transverse,
fri"*;:"R-.*i"tt bf ttittth sternite (Pl' xxv';;"Air;;incr"jr."tsetule,delicate
small,
verv
dististvle (Pl' XXV' fig'
nigf;til;;.-"tt.,r'ur1i..a.--b"*t;;;;;;d;broadry
fie.-4)
ut uu.".'^'i"n.rJirtistvle(PI.xxv, fig' 5, id)b, od) rerarivery';iffi;;;i31a,te,i;G;d;J;t""iaeO

wittr numerouserect pale sete'broad, the outer margin wrinkled
H abitat: Canadian Arctic'
H o l n t y p e : d ,so u th a mp to n l sl a n d ' July3' 1930( Sutton) 'itriy}i:if;;g."reof Dr. williamJ. Holland'rronorthishand,somein naming.riputnin
Al""rrrder, rikewise from the canadian

The crosestany appearsto be \ri"n subporari, uniformly blackened,the tibial spurs
antennre
Arctic. Both speciesagreein having thesimilarappearance.The presentfly
somewhatofput"t",,,
with.u.*i,,g
andslender,
Iongandquite distinct, and with the detailsof the hypohasthe ye*ow uoJ bhrk ubdo*i-rrurpatternof thepa"ily fusedwith the sternite' and the notch
pygium t"ry aiii"rent, the Uu.i.tvf" U"ingi"tsitt morebroadlYU-shaPed'

6-10)'
7. Tipula suttonisp' nov' (Pl' XXV' figs'
with two intermediatebrown stripes;antennee
Generarcolorationgray, the prresc-utugr in.'ffi;Gma. smattand ielativelvill-

brackthroughout;wingss1uy,qf,,'ffih;r'ri;u,otigi" of latter; malehypoconnectingwittr iZ., .o*" distancJ;A;;;, tilE

defined; rn-cu
t.i"a"a, q"pu1ui"dby a narrowmedian'altl*]fy'Ufutk;;;tt"'itiit
pygiumwith till;6;-o-t tn" t.'.srt""r""tv-uir.r.,
bearsfbur slenderspines;
witt
surfaceventralonnotch,;;ilhittistvie buffv-veltow'verv broad'mm'
Mak: Length, unottt 9-12 mm'; wing'

10'5-14

parpi
stout;
nasus
duTk
srav;
andstout,
I!#:Hil,'.ii3l,i;,hl.lrti,#i;;rl[f;l]:,ilftthrouqhout'ii'oit' black$G *{h I;;v'btack,the nasii;;;;'';;t.":-t};;;";Jut1"'tvu"iutg"*i"t-oi tut-! su[equalsgqmentl;f;;;.';;uf
titit.-pruinose;flagellaruHead Br&Y'
the sesme't.
..$',.r't, the.verticiil;'h;;![u"llJtesments'eyes
in tenstht" th;;";;il#;itubercle;
without
wide.
vertex"nterior.upitturf^*".iiu"'"iitr;
indicatedwith a barery
ott tft" posteriorvertex' lacking on;;li;t...'.t^r.i*-tttui;t;,U,rrrdurrt
small; gene *iif,pair being
the medianare&.
distinct brown stripes'the lateral
Mesonotumgray, the prrscutum with two

6

MEMoIRS oF THE cARNEGIE MUSEUM

obsoleteor nearly so; postnotal mediotergite with a capillary darkened median line. Pleura
Br&y, the dorso-pleuralmembrane more buffy. Notum and pleura, together with the coxe,with abundant long seta, which are chiefly pale in color. Halteres dusky, the base of stem
restrictedly pale, the apex of knob more or lessyellowish. Legs with the eoxaeand trochanters
gray; femora reddish brown, passing to black at tips; tibie and tarsi black; legs relatively
short and stout; tibiai spurs | -2 -2; claws with a single small tooth near base. Wings grayish
subhyaline; stigma relatively small and ill-defined, brown; obliterative areas extensive, but
not conspicuous;veins brown. Costal fringe short and dense; macrotrichia of veins very
sparse,including a weak seriesalong the entire length of distal sectionof E,+r and for almost
the whole length of vein ?nd,A. Squama naked but with a seriesof more elongate sete on
margin immediateiy beyond it. Venation (Pl. XXV, fig. 6) 1 Rr+z entirely preserved;rn-cu
con:recting with Mn shortly beyond base, leaving a short basal section; Cu, very pale; cell
2nd A wide.
Abdomen Bray, the caudal margins of intermediate tergites narrowly buffy, the lateral
margins more broadly so, in some specimensthis pale color obsoleteor nearly so; outer dististyie of hypopygium buffy; tergal lobes sooty-black. Male hypopygium (Pl. XXV, fig.7)
with the eighth tergite not concealedbeneath the seventh. Ninth tergite entirely distinct
from the sternite; basistyle partly fused with sternite, the suture indicated both dorsally andventrally, the fusion being complete on the mid-third. Ninth sternite extensive,with a deep
U-shaped median notch, the remainder of the mid-ventral region narrowly membranous;
outer dorsal portion of sternite denselyset with short, erect setulre. Ninth tergite (Pl. XXV,
fig. 8) entirely blackened above, with two conspicuouslobes, which are separatedby a deep
median notch; lobes rounded, their dorsai surfacedensely coveredwith short seta; on ventral surface of tergite a narrow blackened sclerotized band, rvhich bears two slender spines
on either side, the more lateral pair being directed caudad, the sublateral spines smaller anddirected more dorsad. Eighth sternite unarmed. Outer dististyle (Pl. XXV, fig. 9, od) unusually broad and flattened, only a little narrower than long. fnner dististyle heavily
blackened,narrow, shapedas shown (Pl. XXV, fig. 9, id).
H abitat: Canadian Arctic.
Holotype: d, Southampton Island, July 1930 (Sutton).
Allotopotype:9, Julv 3, 1930.
ParatopotApes:
4 d &, Julv 3, 1930.
The holotype male is the specimen having the smallest measurements,as given above.
Tipula suttoni is named in honor of the collector, Dr. George Miksch Sutton. In its
general appearance, almost uniform gray coloration, unpatterned wings and rather eonspicuously hairy body, the speciessomewhat resembles?. besselsiOsten Sacken and allied
membersof this immediate group, but is very distinct in the details of the male hypopygium.
The female, which I refer to this speciesas allotype, has an ovipositor of peculiar type
(Pl. XXV, fig. 10). The cerci are very thin, non-sclerotized,theapex of each acutely pointed,
the margins smooth; hypovalva with a dorsal flangeJilie lobe at base,thesevalves obtuse attheir apices. This type of ovipositor remotely suggeststhe corrdition found in the subgenus
Vestipleabut is quite distinct.
Sunreurr,y LruoxrrN,t.
Tribe Eriopterini.
Genus Hpr,onre Saint Fargeau.
8. Helobia hybrida (Meigen).
1804. Limonia hybrida Meigen; Klassificat., 1: 57, pl. 3, fig. 17.
1900. Hel,obiahybrida Coquillett; Proc. Washington Acad. Sci., 2: 399.
SouthamptonIsland (Sutton); | &, 19 .
The present fly has a vast range throughout the Holarctic region.

ALEXANDER:DIPTERA COLLECTED ON SOUTHAMPTON ISLAND

Drvrsrox II.
THE TIPULID,SOF ARCTIC AMERICA.

The true Arctic Tipulide,of the World are mostly very similar in their generalappearance.
This statement is especially true concerning the dominant genus Tipula, where several
subgeneraare involved, yet all speciesappear at first sight to be much alike. Gray is the
preJominant color, while shortened legs are very characteristic. Some speciesof this genus
are strikingly pilose but others have a more normai vestiture. Subapterous speciesare frequent on tn" pribilof Islands (Alexander, 1923), but elsewherein the far north have notb"u1 taken in any numbers. Among the characteristic Arctic Tipulide in the New World
may be mentioned: Prionocera parrii, P. parrioides, Nephrotoma lundbecki, Tipuln (Vesti'
T. diflaua, T. hewitti, T. h,ollnndi, T. johanseni, T.
plei) arctica, T.bessels,i,T. besselsoides,
suttoni, Dactylotnbisrhicnopt;iloides and Rhabdomastir (Sacandaga)caudata.
A Lrst oF THETrpur,ro,o or Anctrc Auonrce.
Areas inctud,ed,:(l) Alnska,north of the Arctic Circle but also including the Pribilof
Islands. (2) Canad,ian NorthwestTerritories, north of the Arctic Circle, including ail of theDistrict of Franklin (Baffinland and Ellesmereland) but including also Southampton Islandand coastalLabrador, as far south as 55o N. Latitude. (3) Greenlnnd'
Areasomitteil: (l) Alnsko, south of the Arctic Circle, including the Alaska Peninsula andAleutian Islands (this omitted area includ.esnumerous Tipul;ida from Sitka, desoibed byBergroth; the Harriman Alaska collections, reported on by Coquillett; the Katmai collecby Alexander; and other smaller series. These speciesare not Arctic species,
tions, d.iscussed
are more especially Hudsonian and. Vancouveran types). (2) Canad;ian Nortlrusest
but'Territories,
south of th; Arctic Circle, including the Distriets of Mackenzie and Keewatin,with the exception of Southampton Island (this omitted area includes numerous high Hudsonian types laken by Bryant, Harper, Kennicott, and others, but with few or no strictly
Arctic i;p"Uany. (B) Southern Labrad,or. Most of the speciesof Tipulid,e taken by Perrett
at Hopedut. (t ut. s5%" North) are wid.elydistributed Hudsonian types, but at the sametime
includ.ea lesser ,rrrrob.. of apparently true Arctic species. Southern Labrador, Canadian. Labrador and Newfoundland have produced no strictly Arctic Tipulide.
SouthamptanIslnnd,. Despite the-fact that it lies south of the Circle, all speciesof.Ti'pulacerand Nephrito*o taken by Mr. Sutton represent characteristic Aretic species. ft seems
tain thai the above artificial and arbitrar-r-boundarieslisted above will require much modification as a result of future discoveries.
(The date in parenthesispertains to the Bibliography)'

TIPULIN,0.
Alexander. Labrador; SouthamptonIsland. (Alexander,1927)
Prionocerael,ecta
P. parrii (Kirbv). CanadianNorthwest. (Alexander,1919)
p.binioides (Alexander). CanadianNorthwest. (Alexander,1919)(Nielsen)llNephrotoma'x.'rundbeckiarct'icolaAlexander: ltl. Lundbeclcz(Niljr."l.=-Cr"uaiun Northwest;SouthamptonIsland; Greenland(Nielsen,
1907;Alexander,1919)
Alexander. Pribilof Islands. (Alexander,1923)
ffuitn atnscaensis
Alexander. Pribilof Islands. (Alexander,1923)
T. al,eutica
T. apertaAlexander. Labrador. (Alexander,1918a)

8

MEMOIRS OF THE CARNEGIE MUSEUM

Alexander. Northern Alaska. (Alexander,1918a)
T. bergrotlviana
OstenSacken. CanadianArctic. (OstenSacken,1876)
T. besselsi
Alexander. CanadianNorthwest. (Alexander,1919)
besselsoides
T.
Loew. Labrador. (Alexander,1928)
T. canadensis
T. coracinoAlexander. Northern Alaska. (Alexander,1918a)
T. diflnuaAlexander. CanadianArctic. (Alexandir, lg_19)
Alexander. Labrador. (Alexander,1928)
T. entomophthora
lT. gtomerataWaIkEr: T. arcticaCurtis]
T. grenfelliAlexander. Labradol. (Alexander.,.1928)
f .hewitti Alexander. CanadianNorthwest. (Alexander,1919)
sp. nov. SouthamptonIsland. (thi's report)
T. h,oltandz,[7. imperfectaAlexander: T. apertaAlexander]
T. joianseni .Llexander. CanadianNorthwest._ (Alexander,1919)
T.hirbyarzoAlexander. I.{orthernAlaska. (Alexander,1918b)lT. lnbradoricaAlexander: T. platymeraWalker]
T. macrolabisLoew. Labrador. (Alexander,1928)-lT. noduticomisZetterstedt: T. arcticaCurtisl
T. packardi Alexander. Labrador. (Alexander,^192.8)
T. perrettr.Alexander. Labrador. (Alexan{er, t.92p)
T. platymeraWalker. Labrador._ (Nexander,.1928)
T. priiiloferzsfsAlexander. Pribilof Islands. (Alexander,1923)
T. prod,uitellaAlexander. Labrador. (Alexander,1928)
Loew. Labrador. (Alexander,1928)
T. ieptentrional:is(Alexander,1928)
Laborador.
T. sertaLoew.
T. subarcticaAlexander. CanadianNorthwest. (Alexander,1919)
T. wbpotorisAlexander. CanadianNorthwest. (Alexander,1919)
Alexander. Labrador. (Alexander,1928)
T. subserto
T. suttorvisp. nov. SouthamptonIsland- (this report)
Loew: T. platArneraWalker][7.
' tesselalo
T. wlvitneyiAlexander. Pribilof Islands. (Alexa,nder,1923)
pribilouiaAlex?nder. Pribilof Islands-. (Alexander,1921b).
f . (Ues'oil,puln)
f . iVesttpler) aictico Curtis. CanadianArctic; Greenland;Arctic Europe (?). (Alexander,
1919;Curtis,1831;Nielsen,1907)
LIMOI'{IIN,O.
LruoNuNr.
(Steger). CanadianArctic.
Limonia (Diffanomyi,a)decor,oL. (D.) halterata(oiten sacken). Labrador. (osten sacken,1869)
PporcrrNr.
Alexa_n{er.CanadianNorthwest. - (Alexander,1919)
Tricyphonabreuifu,rcata
T. g[;cial;isAlexinder. Pribilof Tslands. (Alexander,1921b)
T.hannai Alexander. Pribilof Islands. (Alexander,1923)
Hr:xetourNr.
rhimoptitoid,es(Alexander). Canadian Northwest; Ellesmereland;Labrador.
Dactyl,oln1is(Albxander,1919,1924;Johnson,-19?9)
Coquillett. Bering Island. (Coquillett, 1899)(Phalid,orea)'fiiiiiili
fuluocostal:isEnroptpnrNr.
ClvioneawaugluiCurran. Labrador' (Curran, 1925)
tl iiii"ophiia ul,tima(OstenSacken). Northern Alaska; CanadianNorthwest.

ALEXANDER:DIPTERA COLLECTED ON SOUTHAMPTON rSleNn(Sacand,aga)
caud,ata(Lundbeek). Baffinland; Greenland. (Alexander,1914;
Rhabd,omastir
Lundbeck,1898)
H etobia hybrid,a(Meigen). CanadianN orth_west; Southampton Island.
(Lundbelk,
1898)-Ormosiadfflns (Lundbeek). Greenland-.
Alexander. CanadianNorthwest. (Alexander,1919)
nrtoftrio'(Ptiloionopa) angustipennis
which have beenrecordedfor Greenland,
Recordsof a few Europeanspeciesof Tipul;idce,
apparently in error, are omitted from the above list. Such recordsinclude Limoni'a (Dic,iiomyioj mod,esta(Wiedemann),Ormosiafascipenrvis(Zetterstedt), Nephrotomal:ineata(Scopoli),and others.
A BIBLIOGRAPHY OF ARCTIC AMERICAI{ TIPULID,O.
from the United Statesand Canada. Tipulida, Diptera'Ar,oxeNunn,C. p. 1914. New or little-knowncrane-flies
579-606.
1914:
Phila.,
Nat.
Sci.
Acad.
Proc.
from the United Statesand Canada. Tipulida, Diptera. Op. cit., t9I5:
lglb. New or little-known crane-flies
458-514.
lg18a. New Nearcticcrane-flies(Tiputide, Diptera) Part IV CanadianEnt., L: 60-71'
1918b. New Nearcticcrane-flies(Tipulide, Diptera) Part V Op. cit., L:242-246.
lg1g. The crane-fliescollectedbV the CanadianArctic expedition,1913-1918.Report of the CanadianArctic
Yol. III, Insects;part C, Diptera,pp' 3c-30c'
Expedition,1913-18,
1920. The Crane-fliesof New York. part l. Distribution and Taxonomy of the Adult Flies. Cornell Univ.
Agr. Expt. Sta., Mem. 25: 765-993.
tgzl;. The Crane-fliesof New York. Part II. Biologyand phylogeny. Op. cit., Mem. 38: 691-1133'
pribilof Islands,Alaska. Proc' CaliforniaAcad. Sci.,
192tb. Dipterousinsectsof the family Tipulide fromlhe(a) XI: 183-184.
159-169'
t1zi.' Diptera of tfru Pribilof Islands,Alaska(Tipulidr and Rhyphida)' \orth AmericanFauna,46:
Tidsskr',
Entomol'Norsk
1898-1902.
inthe'iFram,"
Expedition
ArcticNorwegian
1924. Reportof the Second
I: 296-297.
lg21. New NearcticCrane-flies(Tipulida, Diptera) Part XII. CanadianEnt', LIX: 188-189'of th" g.our Tipuia from Labracior.(Tipulidre,Diptera) Op'cit',LXI:95-101'
1g28. New or little-knownspecies"bontributions to the Natural Hislory of Arctic America." Bull. United
Kumlien's
Buncnss, E. 1g79. Inludwig
StatesNat. Mus., XY: l-179.
Coeurr,lrtt, D. W. 1899. Fur-sealsand Fur-sealIslands,IY: 342'
CunneN,C.'H. 1925. Three new Diptera from Labrador. CanadianEnt', LVII: 24-26'
Voyageto theCunrls, Jonw. l83l. Descriptionof the insectsbroughthomeby CommanderJ. Clark Ross' Ross's
Arctic Regions,Appentlia,p. lxxvii, pl. A, fig. 15'-& Fauna Grenlandica'pp. 1-452'
Fernrcrus, Otto. 1780'
LuNosocx, Wrlr,. r^SiZ. ConspectusFauna Grcenlandica' Pars secunda.II' LandHpxngsp*, Ker L.,
XXII: _481-8-?3-.
arthropoder(insectaet,Arachnida) Meddelelserom Grlnland,"Danmark" Expedition.
Op' cit', XLIII:35-68'
Jon,rNsnu,Fnrrs, & NrnlsnN, I. C. 19f0. The insectsof thepeople. Appendix,thetheandcountry
The
Labrador:others.
JoxNsoN,C. W. 1g0g. /n, Grenfell,W. T., andpp.427-441.
thebeetles,
insects,excluding
1929. Diptera of Labrador. Psyche,XXXVI: 129-146'
North-west
Krnnv, Wu. 1824. Supplementto ttt" Appendix of Capt. Parry's Voyage for the Discovery of app.
ccxiv-ccxix'
1819-1820,
in the Years
Passage
I. Vfd. Meddel naturhist. Foren.,Kjdb-enha-vn,-pp'-236-314'
Lurnrncr, Wrr,r,. 1898. Diptera grcenfandica.
om Grlnland, XIX: 95-120'
LuNnnrcr, Wrlr,., & DnrcnueNN,I. H. 1896. OstgrlnlandskeInsekter' Meddelelser
13-72'
V:
Arctica,
Fauna
Inseln.
derarktischen
Mpr.lpnson, J. C. H. 1910. Die Dipteren
Nrnr,snN,I. C. 1g07. The Insectsoi East-Greenland.Meddelelserom Grlnland, XXIX: 363-409'
Miscell' Colls'
OsrnN S,rcrsN, C. R. 1869. Monographroitn. Diptera of North America. Part IV. Smithson'
219:1-345.
"Polaris" in 1872'
1g26. Report on the Diptera brought home by Dr. Besselsfrom the Arctic voyageof theProc.BostonSoc.Nat. Hist., XIX: 41-43.
(2) 1: 346-369.
St.acnn, C. 1845. Beskrivelseaf Grlnlands Antliater' Kroyer's Naturhist. Tidsskrift.,

10

MEMOIRS OI. TIIE CARNXIGIE IIUSEUM

EXPLANATIONOF PLATE XXV.

Tiptfu lwllortddsP. Dov.
Fre. 1. Wing. Fig. 2. Male hypopygrum,lateral aspect'
Fro. 8. Male hypopygium, 9th tergite; dorsal aspect.
Fra. 4. Male hypopygium, 9th sternite and basistyle, ventral aspeot.
Fro. 5. Male hypopygium, inner and outer dististyles.
sternite; t, tergite).
tSy-U,ifiii, ia."gor; f, tasietyle; id, inner dististyle; od, outer dististyle; s,
Tiptrb su&or.dEP.uov.

'

Fro. 6. Wing-venation. Fig. 7. Male hypopySum,lateral aspect'
Fro. 8. Male hypopygium, 9th tergite, dorsil and ventral aspects.
Frc. 9. Male hypopygiun, inner and outer dististytes.
X'ra. 10. Ovipositor, lrteral aspect.
(Svfo[ols:'b, basistyie;id, inner diatistyle; od, outer dististyle; s, sternite; t, tergite).

MEMOIRS CARNEGIE MUSEUM, Vol. XII.

M
t l n

(,ta
8{*''*

Plate XXV.

I

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