605
APRTL,
194,61ALEXAND ER_TANYDERIDAENOTESON rHEto*o"llBfolr:r51

rHE AUSTRALASTAN( Diptera )PART IBY CHARLES P. ALEXANDER

Maasanhusetts
Stote College, Amherst
The uncommon and primitive flies that constitute the family
Tanyderidae are better representedin the Australasian region
than elsewhere.By far the largest generic group within the family is RadinoderusHandlirsch, including about ten speciesin theAustralian and Papuansubregions.In large collectionsof Tipulidae received from various friends and correspondentsa few
scattered specimensof these flies were included, representing
various speciesthat are discussedin this report. Where not specified to the contrary the types of the new speciesare preservedin my collection of Tipulidae and allied families. For a consideration of the charactersand scopeof the Tanyderidae,the latest
comprehensive
papers may be consulted.l
RaorNopERUS
soloMoNrs (AmxaNDER)
L924. Tangderu.s (Radinoderus)
scit. Menst., t2:743.

solomonis Alexander, Insec. In-The type, a female,was from Guadalcanal(Guadalcanar), collectedin January,I92L, by J. A. Kusche; preservedin the BishopMuseum,Honolulu. One further female,Solomon Islands (exactisland unspecified),November,1944 (JeanLaffoon).
Radinoderus holwayi Alexander, new species
Sizelarge (wing, female,over 18 mm.); thoracicdorsumchiefly
dark brown, the posterior scleritesof the notum and the pleuravariegated by gray; antennae,including the flagellum, brownish black; legs yellow, the femoral tips and tibial ba.sesbroadly
brownish black; wings subfaleate, whitish subhyaline, handsomely patterned with brown, the dark areas in the basal
third and along the costal border with pale centers and narrow
rAlexander, C. P.
7927. Tanyderidae, Genera Insectorum, Fasc. 189:1-13, pl.
1928. The
Tanyderidae of Australia (Diptera).
Proc. Linn. Soc. New South'Wales,
53:367-374,4 fiss.
1932. The Dipterous family Tanyderidae in Japan (Insecta).
Aunot. Zool.
Japonenses, 132278-281,2 figs. (detailed bibliography).

52

pAN-pACrFrCENTOMOLOGTST

IvolxxII, No. 2

dark borders; oval area in outer end of cell R unusually wellmarked and delimited by curvature of the enclosing veins; vein
Rr angulated and spurred at outer end; eell Mn strongly widened
outwardly, at margin a little less than three times as wide as cellM'; abdomen dark brown, the more proximal tergites and sternites
variegated by grayish or yellowish spots.
Female. Length about 29 mm.; wing 18.2 mm.
Frontal prolongation of head brownish black, the surface
sparsely pruinose, the dorsal surface protuberant; mouthparts
black; palpi brownish black, the intermediate segments a trifle
paler. Antennae brownish black; flagellum broken at the sixth
segment; segments cylindrical, with a dense ereet pubescenceandsparse elongate unilaterally distributed verticils. Head above on
front, anterior vertex, cephalic portion of posterior vertex and theorbits gray, the restricted posterior vertex and occiput darkblown; anterior vertex on its eephalic half very narrow, not muchwider than a single row of ommatidia, on the posterior portion
beeoming about four times as rvide.
Cervical sclerites very elongate, dark brown above, somewhat
paler on sides. Pronotum dark brown above, paler and morepruinose on sides. Mesonotal prescutum and scutum with thedisk occupied by three confluent brown stripes, the humeral andlateral borders yellowish gray; scutellum grayish pruinose, with acentral brown line; mediotergrte brown on central portion, broadly
gray pruinose on the eephalic lateral angles. Pleura and pleurotergite dark brown varigated with light Br'&y, including a majorarea on posterior sternopleurite and cephalic and dorsal pteropleurite. Halteres with stem light yellow, kncb brownish black.
Legs with the coxae and trochanters dark brown, sparsely pruinose; femora light yellow, the tips rather narrowly brownish
black; remainder of legs light yellow, the tibial bases brownish
biack, the amount a little .greater than the femoral tips. Win.qs
whitish subhyaline, handsomely patterned with brown, the markirgr arranged about as in terrae-reginae yet with all details distinct; dark areas of basal third of wing and along the costal
border much paler on their central portions, narrowly borderedby dark brown, the paler centers more or less freckled with paleyellow spots; the subbasal clear band in terrae-reginae is here virtually eliminated in the radical and medial fields; ground costal
interspace opposite the cord very extensive, wider than either subtending dark area I veins yellow in the ground fields, chiefly darkened in the patterned areas, this includin E C, Sc and outer radialveins. Wing outline strongly subfaleate, the margin being strongly
emarginate opposite the termination of veins R' to Mr, inclusive.
Yenation: Outer two-fifths of Rs and the corresponding portion ofvein M behind it strongly convex to delimit an unusually distinct
oval area in the outer fourth of cell ^R; Sc long, Sc, terminatingnearly opposite the fork of R,*r; distal section of vein Rr moderately sinuous, angulated and spurred at outer end; cell /st Mz aAPRTL,L9461

ALEXANDER_TANYDERIDAE53

little shorter than in terrae-reginae, the distal section of vein Mg
more than one-half the length of the basal seetion; rn angvlated
at near midlength; cell Mn strongly widened outwardly as interrae-reginae, at margin a little less than three times that ofcell M'.
Abdomen elongate; basal tergite dark brown, with a major grayarea at the eephalic portion and another less evident one near theposterior border; succeeding tergites dark brown, with a large
silvery gray area on the side beyond midlength, this smaller on
the outer segments, becoming obsolete at near the sixth tergite;
secondto fourth tergites, inclusive, with a more yellowish medianbrightening just before the posterior border; proximal sternites
dark brown, the lateral margins patterned with vag'ue, more yelIowed areas; outer segments uniformly dark brown.
Habitat SoroMoN Isuxos.
Holotype, 9, Guao.q,LCANAL,
M"y 6, L}LB (n. T. Holway).
Named for the collector, Dr. Richard T. Holway, who served
as ensign and naval malariologist in the Pacific area. By my key
to the species of Radinoderus ( Insec. Inscit. Menst., 12 :141;
L924), this fly runs to terrae-reginae ( Alexander) , of southern
Queensland, with rvhich species it has been compared throughout the above description. The most conspicuous characters of
this species include the darkened antennal flagellum and thewing shape, patterrt and venation. The oval area set off in cell R
is more distinct here than in other speciesknown to me.
Radinoderus pictipes Alexander, new species
Legs variegated with brown and yellow, the tibiae medium
brown at either end, enclosinga broad yellow band; antennae 23segmented,flagellum yellcw, the incisures a little more darkened;wings relatively broad, whitish subh1,'aline,
conspicuouslybandedwith brown, the areas only inconspicuouslymargined with still
darker brown; outer dark band sending a long arm to the wingtip in the outer radial eells; major pale areas in outer radial field
and beyond cord with smooth margins, the latter marking lying
almost transversely to the wing; veins enelosing:outer end of eell
R arcuated to delimit an oval area near the outer end of cell R;
vein R' beyond Sc' unusually sinuous, curved at, tip; cell -ZstM,long, approximately twice vein M, beyond it; cell Mn widened outwardly, at margin fully four times m-cLL.
Fentale. Length about 15 mm.; wing 15 mm.
Frontal prolongation of head brown, very sparsely pruinoseI
mouthparts darker brown, the first segment of palpus paler. Antennae with scapebrown, pedicel more yellowed, flagellar segmentsa trifle darker at the incisures; 23-segmented,including: the reduced terminal unit; flagellar segments subcylindrical, with long54

pAN_pACrFrC
ENTOMOLOGTST[vor,. xxu, No. 2

outspreading yellow setae that much exceed the diameter of thesegment opposite their point of insertion. Head light gray; anterior vertex reduced to a capillary strip that is only about as wide
as a single row of ommatidia; eyes relatively large, ommatidia
very fine.
Cervical region dark brownl pronotum somewhat paler brown.
Mesonotal praescutum and scutum with the restricted ground color
light graVr most evident as lateral borders just behind the pseudosutural fovea; posterior interspaces a little paler than three darker
brown stripes, the center of the median stripe more brownish
gray; interspaces with conspicuous erect setae; scutellum lightyellow, gray pruinose, the apex with long yellow setae; mediotergite light brown, paler on sides and on caudal half of pleurotergite. Pleura conspicuously variegated pale yellow and darkbrown, the latter including a conspicuous atea on sternopleurite,anepisternum and again on meron, leaving an extensive pale markon the pteropleurite and posterior border of sternopleurite; dorsopleural membrane surrounding the spiracle yellow, darker behind.
Halteres with stem pale yellow, knob dark brown. Leg:s with thefore coxae and trochanters light yellow; middle and hind coxae
infuscated; trochanters obscure yellow; femora yellow, with anearly terminal brown ring that occupies about the outer sixth toeighth of segment; tibiae brown at both ends, enclosing a broad
yellow ring, this narrower than the darkened outer portion, provided with pale setael extreme tip of tibia paling to yellow; tarsi
yellow. Wings relatively broad, whitish subhyaline, conspicuously
banded with brown, the latter appearing as two broad crossbands,
with additional darkenings at base and again at apex; a verysmall ocelliform mark at origin of Rs; the two major crossbands
interconnected in cells C and Sc only; outer band sending an arm
obliquely outward to the wing tip, crossing the distal ends of theouter radial cells, extending from vein .E' to l?', with pale marg:inal droplets in ends of all cells; in outer radial field an unusually conspicuous oval pale area extending from costa to vein Rn,
pale band beyond cord almost transverse to the wing, both paleareas with almost smooth margins; dark areas very weakly bordered by brown, more conspicuously so in the vicinity of the cord;veins brown, yellow in the patterned areas excepting C, Sc and R,
origin of Rr, and most of the elements comprising the eord. Venation: Outer twe.fifths of Rs strongly arcuate, the eorTesponding
portion of M slightly bent, the two enclosing a conspicuous suboval outline at the distal fourth of cell .E; vein R' beyond Sc, unusually sinuous, at tip curved strongly cephalad, vein E, similarlyupcurved; na-su just before midlength of the nearly square Ms*aicell /st Mz ver1ylong, approximately twice vein M' beyond it; cell
Mn widened outwardly, at margin fully four times r/L-c1'(,.
Abdominal tergites dark brown, segments two to six each witha conspicuous silvery white sublateral spot near base, this smallest
on the sixth segment; sternites silvery white on basal lateral Dor-APRTL,19461

ALEXAND ER-TANYDERIDA E55

tions, the remainder pale brown; outer abdominal segments,including ovipositor, dark brown.
Habita. NETHERLANDs
Nrw GurxuL.
Holotype, 9, Horr,aNDrA,M"y, lgg4 (w. stiiber); to be returned to the Zoological Museum, Buiten zorg, Java.
By -y key to the speciesof Ra.dinoderus(Insec. Inscit. Menst.,
l2:l4I; 1924), the presentfly runs to couplet 3, including orrub
tissimus ( Doleschal) and terrae-reginan Alexander, differingfrom both, and from the speciessubsequentlydescribed,by thepattern of the legs and wings, and in the venational details, asdescribed. The oval area at the outer end of cell R is unusuallyconspicuousas comparedwith the related species,with the exception oL holwayi new species.

A NEW RECORDFOR AULICUSTERRESTRISLINSLEY
While collecting on the floor of San Joaquin Valley about ten
miles north of McKittrick, California, two specimensof Aulicus
terrestris Linsley were taken on the flowers of a Phacelia sp.
The Phacelia were growing in a damp wash some eight feetbelow the general surfaceof the valley floor. The specimenswere
taken at 4:45 p.M., while overcastand cool. The fact that theadults were feeding upon flowers is interesti.g, as Linsleyl has
recorded the adults as predaceousupon the larvae of certain
moths.
As is pointed out by Linsleyl, Aulicus terrestris is found inassociation with the lubber grasshopper, Esselenia aanduzeei
Hebard. This grasshopperhas been found in Monterey Countr,
on Mount Hamilton, on Mount Diablo, in the Livermore Valley,
and in the foothills of Kern County. Aulicus terrestris Linsley
has been recordedfrom all the above areasbut one, that of Kern
County. The new record eompletesthe locality relationship. Al.
though Esselenia.aanduzeei Hebard was not taken in the same
spot with the clerids, the specieswas found in abundancea few
miles to the south.-K. E. FnrcK.
l Linsley, E. G., 19t6. Studies in thc Genur Aalieu
Publ., 6(9) :249-162.

Spinolr.

Univ. Calll.