f,/2
Reprinted from Bur,r,nrrN oF rHE Bnoorr,vN ENrouor,ocrcAr, Socrntv,
Vol. XLIf, No. 1, pp. 19-24. Febluary, 1947.

NEW SPECTESOF PTYCHOPTERIDAE (DIPTERA).
PART III.
Bv CuIRLES P. ATBxANDER,Amherst, Mass.

The precedingparts under this title were publishedin the BurLErrN oF rHE Bnooxryx ExrouolocrcAl Socrnry, 32:I4V143,
1937,and 38 : 3712, 1943. At this time I wish to describethreefurther new speciesfrom Western North America,as well as a further noveltyfrorn Burma. A few additionalrecordsof distribution
for certainrare and little-known Nearctiespeciesof Ptychopteridae
are given. The typesof the new species
herewithdescribedare preservedin my personalcollectionof Tipuloidea.
Ptychoptera uta sp.n.
Male.-Length, about9-9.5 mm.; wing 8-8.5 mrn.; antenna
about5 mm.
Generally similar to Ptychoptera lenis coloradensisAlexander (Bull. Brooklyn Ent. Soc.,32: L4I-I42, 1937),differing
especiallyin detailsof structure of the male hypopySum.
Ninth tergite with the lateral lobesdistad of the outer spine
very small, exceededby the spine; subtergal spinuloselobe
large and clavate; lowermost tergal lobe reduced. Dististylelonger and more slender,sinuous,blackened,the tips pale.
large and massive,blackened,the apex broadly
Gonapophyses
obtuse. Ninth sternitewith the setaeof the median spatula
thelong and slender; subtendinglobes nearly parallel-sided,tips truncatedor weaklyexpanded.
Habitat.-Utah.
Holotype,3, Willard, April 29, 1939(Knowlton & Harmston).
Paratoltotypes,
3J ; parotyfc, 3, M"y 1,1939 (Knowlton & Harmston).
I am greatly indebtedto GeorgeKnowlton and Fred Harrnstonand for manyotherTipuloideafrom Utah. Thefor thesespecimens
detailedrecordfor the stateis in press(Arner. Midl. Nat., 1947).
Ptychopterasculleni,Alexander.
Deseribedfrom Washingtonand Oregon. Additional records:
Oregon.'PeavineRidge,nearMcMinnville,Station3 (605 feet).
S e p t e r n b eMr 4 , 1 9 4 5 ( K . M . F e n d e r ); B a l d M o u n t a i n ,C o a s t
Range,Yamhill Co.,July 19,1942(K. M. Fender).
California: Orick, Hurnboldt Co., June 21,1935 (A. L. Melander).

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Bulletin of the Brooklyn EntomologicalSociety vot.XLrI

P tychoft era pendula Alexander.
Utah; KimballsFort, June 29, 1943(G. F.Knowlton).
Wyoning : YellowstoneNational Park-Roosevelt Station, July5,1923;Old Faithful,July 14, 1923;SpringCreek,Jrly 15,L923;
Turbid I-ake,July 20, 1923(all A. L. Melander).
Ptychopteratownesi Alexander.
Washingtort..
Everett,July 6,1924; Pluvius,July 16,1922;Puget,August 4, 1925; Swauk Creek, June 28, 1924; Toledo, June 27,1935(all A. L. Melander).
Oregou Hood River (Leroy Childs).
Ptychoptera monoensissp. n.
Allied to pendula;generalcolorationof body polishedblack,the pronotutn and nresonotalscutellumyellow; antenlraewith
scapeand pedicelyellow; all coxaeyellow; wings with a weakbrownish tinge, the prearcularfield yellow; rnale hypopygiumwith the ninth tergite deeply notched,each lobe bearing two
blackenedlobules,in addition to the apical point ; dististyleconspicuouslytrilobed, the outelone a very large flattened
yellow blade, the intermediatearm a darkenedsubcylindricalrod ; innermost arm nlore compressed,bearing four or five
powerful spinoussetae.
Il[ale.-Length about8.5 mm.; wing 8 mm.; antennaabotrt
4.1 mm.
Rostrum and mouthpartsyellow; palpi yellow, the terminal
segmentbrownish black. Antennae of moderatelength, approximatelyone-halfas long as wing; scapeand pedicelyellow,
flagellum black; flagellar segmentscylindrical, the verticils
shorterthan the segrnents. Head polishedblack.
Pronotum obscureyellow. Mesonotumpolishedblack, thecentral portion of the scutal region and the scutellumobscureyellow, the parascutellablackened; postnotumblack,the dorsal
portion of the suture betweenthe mediotergiteand pleurotergite more reddened; dorsal pleurotergitewith conspicuoussetae. Pleura black,sparselypruinose,more heavily so on thepteropleurite; dorsopleural region buffy yellow. Halteresyellow, the knobs weakly darkened. Legs with all coxae andtrochantersyellow; femorayellow, the tips rather narrowly but
conspicuouslyblackened,the amount subequalon all legs;
tibiae obscureyellow,the tips narrowly blackened; tarsi brownish blackto black,the narrow proximal portionsof the basitarsi

Feb.,7e47Bulletin of the Brooklyn EntomologicalSociety

2lvaguely obscureyellow. Wings with a weak brownish tinge,
the prearcular field yellorv; a very restricted brown pattern,
especiallyevident over the central cord ; very restricted darkenings at forks of veins Rr*, and Rn*,; veins brownish black,yellow in the prearcularfield. Macrotrichia of cells relatively
abundant,including all cells beyond the general level of forkof R**" and as restrictedseriesin cell R, basalportions of cells
Rs and R; and as evenmore restrictedgroups in cellsC, Rt andM ; no trichia in basesof cellsM 2 or M r. Venation: Rs relatively long ; r-nl connectingwith Rs at fork or in Rn*, shortlybeyond; cell Znd A broad.
Abdomen polishedblack,the posteriorbordersof the secorrcl
and third tergites narrowly pale; hypopygium chiefly black.
Male hypopygium of the general type of ltendttla but differing
in important regards, especially of the tergite and dististyle.
Ninth tergite with an unusuallydeepU-shapednotch,the laterallobesproducedinto small conicalpoints, on nlesaledgeof apex
further produced into two blackeneclhairy lobules,the outerone more slenderand elongate. Dististyle trilobed, the outer
lobe a very large flattened yellow blade, on inner margin nearbase bearing a small tubercle; interrrrediatearm a darkened
subcylindrical rod, the distal half and especiallythe apex withconspicuousdark-colored setae; innermost or lowest arm aflattened-compressed
blade that bears four or five strong spinous setae,in casesthe terminal one isolatedand slightly larger;in other instancesthe spinesarrallged more definitely in pairs.
H obitat.-California.
Holotype,3, Coleville,Slinkard'sCanyon,Mono County,May 28,
1939 (Mont C-azier& T. H. G. Aitken).
Ptychopteralnonoetrsisis most nearly allied to P. pendulaAlexanderand P . toantesrAlexander,beingsomewhatcloserto the former
yet very distinct in the structure of the rnale hypopygium, particularly the tergite and dististyle.

Ptychoptera persimilis sp. n.
Male.-Length, about 8 mur.; wing 7 mm.
Color eharactersalmost as jn P. annandalciBrunetti, 1918,
that is, the mesonotum uniformly black excepting the yellow
scutellumand adjoining portion of the mediotergite. Pleura
yellow, apparently darker on the mesepisterum. Femora yellow, the tips narrowly and inconspicuouslyinfuscated, theamount subequalon all legs; tibiae cleareryellow, the tips still22

Bulletin of the Broolilyn EntomologicalSociety Tot.XLU

more narrowly darkened; basitarsiobscureyellow, the tips andremainderof tarsi blackened. Wings with the outer darkened
crossbandbroken, nlost distinct at stigma and over eachof tl-reforks. Venation : Rs short and straight. Abdominal tergites
yellow, the caudal borders ringed with brownish black; basal
tergite uniformly blackened; a dark ring on proximal half oftergite two ; darkenedareas on outer tergites rnore extensive;
sternitesand hypopygium yellorv. Male hypopygium with thetergite profoundlybifid, as in annandaleiand allied species,but
the arrns much stouter, especiallyat bases; arms bent at near
midlength, the apical portion. stout, provided with abundantpale setae,before apex with a small tuberele. Dististyle ofentirely different conformation; basal half expanded into a
broad lobe, additional to the two basal lobulesin annandalci;
the main lobe withouter lobesnot forceps-like,as in annandalei,,the outer blackenedsetaelong and abundant; separatedfrom
the outer group and nearer the base of style a linear row ofabout five shorter and strongerspines; outer lobe of dististylepale, clavate,provided 'uvithlong slendersetae. Sternal lobes
long and pale,denslyhairy, the setaeof inner margin near baseof unusuallength,the more proximal onesprogressivelylonger.
In annandalei,the tergal arnrs are long and slender,provided
at apex with a densebrush of blackenedsetae. Dististylenrore
the two outer lobes being opposedto
or less forceps-shapecl,
one another at their free ends; no dilation on basalportion ofstyle; spinesof the axial portion all short and stout. Sternallobesrnuch more slenderand fingerlike.
Habitat.-Bu1m3.
Hototyp€,3, Shwenyaung,SouthernShan States,August 1930.
of the Oriental Ptychopteridae,
For the nrostrecentconsicleration
seeAlexander, Arkiv for Zoologi,38 A, No. 2: 1-10, ffi&P,1946.
Bittacomorphella fenderiana sp. n.
Generally sin-rilarto sachenii; antennae black throughout ;mesonotal praescutum with the disk chiefly black, the fourstripes being divided only by paler gray interspaces; ventral
pleurites darkened; legs with the basitarsiblack, the tips notor but narrowly ri'hitened,tarsal segmentstwo and three snowy
white; male hypopygium with the tergal lobesproducedcaudad
into small slenderpoints; lateral tergal arms almost glabrous;
dististylestwo, there being a small cylindrical style or lobe at
the baseof the major one; phallosomewithout blackenedparts,
the outer lateral anglesproduced into obtusehairy lobes.

Feb.,7s47 Bulletin of the Brooklyn EntomologicalSociety

23Male.-Length, about 11-13 mm.; wing 7-8 mm.; antenna
about G8 mnr.
Fenrale.-Length, about 1f13 mm.; wing 7-9 mrn.
Frontal prolongation of head yellow, the basal portion silvery ; palpi brown basally,passinginto black. Antennaeblack
throughout. Head behindblack, gray pruinose.
Pronotum very restricted,pale yellow. Mesonotalpraescutum with the disk chiefly black, producedby four conspicuous
stripes and only slightly paler gray interspaces;humeral andlateral regions pale yellow, sparselypruinose; scutum paleyellow, each lobe with two separateblack areas,the posteriorone very small; posterior scleritesof notum yellow. Pleurasilvery, the ventral sternopleuriteand meron restrictedly
brownish black; a rl1oreor less distinct darkenedarea on theanepisternurn,obscuredby pruinosity; in cases,thesometinres
mesopleuraeven more extensivelydarkened. Halteres pale,knobsweakly infuscated. Legs with the coxaeand trochanters
yellow; femora brownish yellow, the tips passinginto black;tibiae dirty whitish, the tips narrowly infuscated; basitarsi
brownish black,the tips very narrowly to scarcelywhitened;tarsal segmentstwo and three snowy-white,four and five black.
Wings rvith a faint grayish tinge, unpatterned;veins brown,
thoseat extremebasemore yellowed. Venation: r-nl, variable
in position,from shortly beforethe fork of Rs to about an equal
distancebeyondon Rn*r,'Rsvariablein length,in casesonly as
nearly twice this vein.
long as r-in, in other speeimens
Abdomen of both sexesbrownish black, in male the subterminal segmentsa trifle paler. Male hypopygium with thelobesof the tergiteproducedcaudadinto slenderspinouspoints;
lateral tergal arms almost glabrous,with only a few long setae
just beforethe acuteapicalspine. Dististylestwo, there beinga small cylindrical style or lobe at the baseof the major one.
What appears.torepresentan interbaseis a slendercurved
horn, the apicalhalf very attenuated. Phallosomestout, withpoints; outer anglesproducedlateradinto obtuse
out selerotizedhairy lobes;apex obtuse.
In sochenii,the lobesof the tergite are low and obtuse,hairy,
not produced; lateral tergal arms relativelystout, with scatteredsetaeover the entire length, more concentratedabout the acuteblackterminalspine. A singledististyle,providedwith numerous setae,broadestat base, narrowed outwardly. The supposed interbasehas the basal half thickened,the apical spine

24Bu,tletinof the Brooklyn EntontologicalSociety vot'xLIr

nearly straight. Phallosomewith heavily blackened,sclerolobe with coarse
tized armature,the long simpleunblaekened
setae.
Habitat-Northwestern North Arnerica(Vancouveran).
Holoty7e,3, PeavineRidge,near N{cNlinnville,Oregon,Station
3 A, M"y 15,1946(K. M. Fender). Allotyltc,?, Albright'sRanch,Dayton,Oregon,September19,1946(K. M. Fender). Paratopotypes,4&3, Stations3 ancl3 A, M"y 5-26, 1945,August 22, 1946,
Septenrber17, 1946 (K. M. Fender); ltaratyltes,| 3, with theallotype; | 3, Massett,QueenCharlotteIsland, British Columbia,BritishColurn1898(J.H.Keen);19, StanleyPark,Vancouver,bia, Septernber
3, 1930(H.8. Leech);d?, Ashford,Washington,August 18,1940(H. & M. Townes); d, Keyport,Washington,J,tly
1905 (R. W. Doane); Lewis and Clark State Park, Washington,September28, 1946 (K. M. Fender). Certainof the aboveparatypeswereearlier(Bull. BrooklynEnt. Soc.,38:41;1943) recorded
sacheniiand the changeshouldbe noted.
as beingBittacomorphella
Mr. Kenneth M. Fender,keen studentof the Canthariclbeetles,
first called to llty attention the fact that there were two distinct
occurringat his study stationson Peaspeciesof.Bittacornorphellavine Ridge,near N{cMinnville,Oregorr. From a stuclyof the tnale
genitalia there is no questionbut that trvo distinct speciesare
involved. I am rnost pleasedto nanle this new speciesfor Mr.
Fender,in appreciationof invaluableco-operationin the study of
our western Tipuloidea. Von Roder's clescriptionof sackanii(Wiener Entonrcl. Zeitung,9: 230; 1890), for a copy of u'hich I
anr indebtedto Mr. GeorgeGyrisko, is short but quite sufficierrt
for purposesof identificationof the species. It is eviderrtthat
sackeniiis a somewhatmore southernspecies,its kuowtr raltge
includingNevadaand California,as rvell as Washingtonand Oregon, as far north as Mount Rainier, Washington. The type wasfrom the Sierra Nevada,in Nevada,presumablyfrom the Lake
irr theTahoesection,takenby Herbert K. Morrison,who collectecl
state irr 1878and again in 1884. Aldrich (Psyche,7 , 20V201 ;what he consideredto be sachenia(from Lake1895) re-described
Llnion, Seattle,Washington,August-septernberL894, John M.
Alclrich) but which is very evidentlythe new speeies,
f enderiana.