3?f
ARKIV FOR ZOOLOGII]TGIYET ,{YK. SVENSKA VETENSKAPSAKADBMIEN
BeNo 27 L.

N:o lZ.

SCHWEDISC}I-CHINESISCI{E.tryIs
SEN.
SCI-I
AF]'I,[C}[EEXPEDI]'ION
NA(IHDEN
IiORD}\TESTLICHEI{PROYINZEI{CHINASUNTBR I,EITUNG'\ION DR. SYEN HEDITYUND PROF. ST7PING.CIIAN(;

INSI'I{TEN( i t . t s , \ \ t \ i l .'.tt' \ o l l s ( . F I \ \ ' 1 . ; t ) t s ( : i l |- .\ t \t z ' 1 I. ) I i t t t i x t , t . : t ) t . t . t ( ) N
I ) t r . I ) ; t t 'I I ) 1 i l ' , .^vt I I , : t!.1 f 2 ; - l 1 ) : J { }32I)IPTI'RAi.

'rIPT]I,IDAE
BY

CTIAITLESP. ALEXANDER

STOCKHOLM
Ar.MeYlS't.c{\\-IKSI.ILLS BOKTRYCKERI_A. -B.

BERLINPARIS

R. TBIEDLANnT:n &, soux
I I cenr,srRAsaE

trBnArRrE c. KLTNCKsTEoKI I nuu Drq r,rr-r_E

I9 3 4

ARKIYTORZOOLOGI.
BAND 27 A.

N:o L7.

Schweilisch'chinesische
wissenschaftlicheExpe.
dition nachden nordwestlichenProvinzen Chinas,
unter Leitung vonf)r. Sven Hetlin und Ilrof . Sii Pirrg'ehflng.
Insekteng e srmn re l t vo m schwedisch Ar zt der lixpedition

Dr. David Hunrnrel 1928-1930.

32.
Diptera.
7. Tipulidae.
tll'

CHARLES P. ALEXANDER.
\\' ith 3 plutt^s.
C o u r u t t r t ti e a te tl()e to b e r 2 5 th l9:13 lr v Y. S.liist' Bur autl U. - ( tr xsr o.

The ver.y interesting crane-fly rnaterial collected in theProvince of Kansu, northwest China, by Dr. Dlvro I{uuunl,
while acting as physician for the Svnw lluorlv-Expedition of
1928-1930, has been submitted to me for examination b.yProf. Yuavn S.riisrno'r. Virtually all of this material wascollected ut the following two stations both in Kina, Kansu:
Drakana, altitude about 3000 rneters,mid.-August.
Kung-tze-tagga, Tsaluk valley, Min-shan, alt'itude about
3028 meters, July 20, 1930 (ut light).
The types and most duplicates of the speciesdiscussed
have been returned. to ProfessorSrtisrnor, for deposition in
Arki,a lor :oologi. Bd 27 A.

N..o 17.

2

An,Krv t'iitt, zool,ocr.

IJAND 27 L.

N:o 17.

the Naturhistoriska Riksmuseum,Stockholm. Duplicate specimens where available are preseryedin my collection. I amvery deeply ind.ebted.to Dr. Iluuunl, and to Dr. S.rtisrnnr
for the opportunity of studying this rich and instruetive seriesof Central Asian Tipulidae.

Tipulin&e.
Tipula (Yestiplex) subearinat& Al,nxeNDER.
1921. Tipula suhcarinaftt AT,ExANDER;Itull. Mus. Hist. Nat.
1921: 347-348.

l'aris.

Originally described frour a small series of _specirnens
taken i; Mongolia. Two additional specimensin the present
collection, one from Southern Mongolia, 1927; the second.fronr
Tabo-al, China (Awounssou).
The present specimensare a, little srnaller than the types(MaIe. -: Length,13-14 mm.; wing, 14-15 mm.; antenna,"
about 6 mm.). Nasus long and cottspicuous.Antennae (male)
relatively long, the basal three segmentsyellow, the_rerainderof flagellurn black: segmentsexceedingin length-the longest
verticils. Wings (Fig. 1) with a pale brown and white pattem.
the white color in ceII R-" continued to the rving-apexor
nearly so. The male hypopygium has been describetlbut notpreviously figured (Figs.-13-Lb). The most conspicuousfeaio"e* lie-in the pale tergite. g f, with the caudalmargin broadly
emarginate; basistyle, b, produced, at apex on_proxirnal facewith a small blackenedtooth, less distinct in the presentspecimens.
The speeies falls in that group of the srtbgenushaving
the posterior portion of the basistr-leunarttretlwith a spineor lobe.
1t'ipultl( \'estiplex) ltultrrneli sp. n.
Mesouotal pra,escutum l'ellowish gray, with foul darker'
gray
stripes thal are narrowly bordered by blacliish. fonning';
median black vitta; posterior sclerites of notutu sraJ'. rvitha capillar.,v d.ark vitta; apices of knobs of htr,ltel'es1-ellorv; Iegs
chiefly biack, the femorul bases narrorvlr' -r'ellorv: rvinss palecream-yellorv, with a diffuse pale and. rlarker ltrorvtr pattem,
the darker brown areas includ.ing the bases of cells -R and J1.
Length, about 20 nlrn.; $'ing, 22 lnm.
Female.
Frontal proloneution of heacl llrorvnish black, heavilr'
pruinose; oas..s stout ; palpi black. Antennae rvittr setll)oCHAIi,L}]SP. AT,EXAI.iI)ER,32.

THI.] 1'II'UI,II)A}.].

3

brownish yellow; peclicelclear light yellorv; flagellum uniformly
brown; flagellar segmentssimple, the verticils slightly exceeding
the segments. Head graf, more yellowish gra"y on front andanterior vertex; vertieal tubercle with a, conspicuorls d.ark
brown median stripe that ends at near midlength of vertex.
Mesonotal praescutum light yellowish graI, with four
clarker gray stripes that are narrowly but conspicuously bordered bv blackish, the internal borders of the intermediate
stripes confluent or virtually so, forming a conspicuous blackrnedian vitta; internal borders of lateral praescutal stripes
rnore evidentlv clarkeneclthan the lateral bord ers ; scutal lobes
vellowish graJ, variegated with brown; scutellum and rnediotergite gra)', with ^ narrow blackish median vitta. Pleura,s'ra,y, slightly varieEatecl by more .yellowish gra-y; tlorsopleuralregion buffy. Ilalteres yellow, the basal half of knob clark
browrr. Legs with the coxae gra.v; troehanters .yellorv; femora
almost entirely blackened.the basesnarrolvlv and conspicuously
light yellorv, the amount suberlual on all legs; tibiae and.tarsi
brownish black to black. Wirrss (FiS. 2) with the ground.color chiefly crearn-rellow, the prearcular and costal iegions
extensively clear bright 1'ellorv; an extensive pale brown anda much more restricted d.arker brown pattern, the pale brown
including most of the wing-clisk, restricting the sround to an
incomplete fascia bevond. eord., a pre-stigmal brightening, Llarge area near outer end of cell lJ[, ancl a broken fascia,
across the basal fifth of wing, beeinning in cell l?; the clarker
brown areas include the stigrna and eonfluent darkenings onanterior cord, the stigrna not involvinq the costal cell exeept
ttt free tip of Str; zl,srnall cloucl at oliq'in of /is; ltasesof
cells It trnd J[,, itnrl cell (' on both sirlesof arcnlus,ilarkened;
less eviclent infttseations along vein ('rr. r't'ins lellorv ancl
brorvn. the outer radial veins errtirelr lellow'. the outer rnedial
veins brorvn n'ith their apic'espaling' to vr.llow. llaerotrichia
trf veins bevond eorcl snrall and sl)arse; r'eins ICr+:, J'lzt:t;-R.3
and tt A without trichia ; Ii 1.;, ,l[1. Jf ,,, ftu and tips of ]UI.L
ancl errt I with a fen' seatterecltrichia; tlistal sectioir of Cu,
rvith relatively abundant triehia on clistal three-fourths. Verration: Il rl , entire but pale: petiole of cell ,l[r subequal toor shorter than m; nt -('t( on ,l[r shortlv bevond the perpen,licular base.
Abdorninal tergites obscure ;'ellorv basallv, the outer l)ortiorts rnore clarkenecl zurcl pruinose; a conspieuolls rneclianblaekish stripe; lateral borclers of tergites nal.l'ou'lr-Pale; stern ites gra)-. Ovipositor broken.
I{rlotrypr:.
?. Kina. June l9 (D. }{r'rrrrr:r,.)

AR,Krv riin zoor.,ocr. B A N D 2 7 A .

N:o 17.

This splenclid fly is namecl in honor of the collector ofthis rich series of crane-flies,Dr. Devro Huuunr,. The nearest
ally is Tiltula (l'-estiltler) grahami Al,nx.l.wDER(Szechwan-Tibet
Border, &t high altitudes) which differs in many details ofcoloration of the body, legs and. wings, most evidently in theyellow femora with narrowl.y blackened tips, and the muchd,arker, more contrasted.wing-pattern. In the present uniquetype the region of the scuturn is damaged and it cannot beaffirmed that the median black notal vitta is eontinuous over'the entire thoracic dorsum, though this is very probably thecase.
Tipula (Vestiplex) hetlini sp. n.
Allied. to su,bcentralis;antennae with basal three seErnents
yellow, the reuraining segments almost uniformly blaekened.
bnly the basal segments vaguely bicolorous; mesoxotal _praescutum graI, with four gra-v stripes that are narrowly borderedby darker; wings brown, variegated. by darker brown andwhite, the pattern highly eontrasted; outer half of cell Ri
pale; male hypopygium with basistyle unarmed; outer distist.yle
broad.ly spatulate; intrer dististvle biclentate.
L e n g t h , a b o u t 1 8 - 2 0 n l l r r .; r t ' i t t g ' , I ! f - 2 ( ) l l l l l l . :M ale.
antenna, about 5,;'t-5,.; lttul.
Lengt[, about 25 lllnl.; rri1g, !2 lrllll.
Fernale.

Frontal prolongation of heacl brownish yellow. clarker
laterally beneath; nasus short and stout; palpi dark brown.
Antennae with basal three segments ;'ellow; succeedingsegments very weakly bicolorous, the basal enlargements black,the apical portions-uniformlyof the segrnents dark brown; outer !egblackened; longest verticils subequalto
ments moreor a trifle shorter than the segments. Head gray, with a,brown, capill&ry, median vitta.
Mesonotal praeseutumlight gra)', with four slightl-v briglrter
gra.y stripes that are naruowl.y bordered by brown; scututn
ivith mediat. area buff.y .9'ral, the lobes with two gray areasthat are encircled.b"v brown; posterior scleritesof rtotum gr&\-1
with a, capillar.v bi'own rnedian vitta. Pleura gl'a)'. -vaguelv
marked" wilh paler gra)'; dorso-pleural reqion ltuffr. _Halteresyellow, the knobs dartr brorvn. Legs rvith the fenror&brorvnish
so-olryellow, the tips rveakly darkenecl. less conspicuously_
posterior legs bt type; libiae obscure_r'ellorv.-the -ti1r1darker;
larsi passing into darker brown. ln fetnale, !!u l.g!- areshorter, storiter and generally darker in color. Wings (FiS. 3)
with the ground.-color relatively dark brorvn, triglly eontrasted'with whilish areas that include the distal half of cell lir;oHAlir-,Es r'. AL,ExANDER, 32.

THn rIl,trr.,IDAI.r.

;)

conspicuous crossband.sbefore and. be.-voncl
corcl, the fo'uer
broader; cell Llu chiefly pale; base of cell lrt ,!, vel..)-extentt^tu} pal_e;darker brown areasat stigma, anterio" .o"ii, origin
of .Bs and enclof ^5'c;cells /? aud It not darkeneclbasallj veins;brown. fn female, the pattern is still clarker and roo". eontrasted. Venation: )[r+1 sub€r1ual
to or exeeecling
basal sectio'of I[r.
Abdomen rvith ba,sal tereites yellow, narrowly trivittate
with dark brown, the median stripe narrower and ciarkerthan
the laterals, interrupted at eaudal bord.ersof segrnents;lateralrnargins of tergites pale; outer se_gments
nrore .Lifo"t1i,v clarkened; sternites obscure vellow, t[e outer segnrentsclarkelecl.
In fenrale, sternites more brownish yellow. il{ale h.r,pop.}'gi*rn
mnch resembling .cubcentrali,,s;
tergal saueer (FiS. r i;, li i')'.orr_
spicuous, the cauclal rnargin broacll.yernarginerte,at base of
erna_rginationwith a further U-shapeclnotc|: trvo Slaekened.
teeth on either side. tht: rnore cephalieacute; posterior5ord.erof saueersrnooth. Basistvleunarrirecl.Outer distist.ylen broad
spatulate blarde. Intter,iittirttl. consliicnouslybicleltate, theirrner point heavilr- blackenecl(Fig. I (j; orl. ttl).
Holotulte,d, Fu-shu-shi.Tien-shan,Aug'st
1g2g(D. Huuo
MEL.)All,otl1pe,?, Fu-shu-shan, Tien-shan, altitucle 1g00-2000
rneters(D. Huurrr:r,). P?l1tJ p_es,2 dc, rvith allot.ype,altitucle
2650 rneters,June 1928 (D. Huulrnr_,).
Tipula ( I'9sti1tler.)hedi,niis respectfull.vcleclicatecl
to Dr.
Svnx Hunrx, distinquisheclexplorer-scieltistof Central Asia.
Tlre species is nost g'enerall1sirnilar to 'l'. ( l-. ) .*ttbcetttrrtli*
A r,uxANDER(Kaurtchatka). tliftering. consPicuonsl.vin the
structure of the st.vli of the tnale h1-popvr.l'i.,ttr
aucl less evielently in cletailsof coloration, as the ,rro"siririfonrrlr-blackenecl
antennae. The course of vein fI, of the lvings is clilterentjn
tlre two flies. in ,srrDrentrrtl,a.r
being verv slightly'convex,whereas
irr heilini it is erluallv gentlv cottve*.
Tiprrla ( A r.rrtipula)pertinax sp. n .
General coloration clark; basal three segmentsof altenlae
lellow; wings with the grouncl-color gra)-, sparsely variegatedl]fPulg brorvn and rvhitish, the pattern p"l.; basal foul abtlotninal segrnentschiefl.y.yellorv,th-e remairicler brorvnish blacl<;
rnale h:'popJ'giuP rvith the median lobe of tere.ite relatively
broad, its apex bifid; inner clististyle with the oiiter lobe com_
Pressed, tenlrinating in trvo slencler, spinous poirrts tltat aredecussate.

(iARI(IY riin, zooLocir.

r r A N D2 ? A .

N:o 17.

Length, 14-15 mm.; wing, 15-16 mln.; anMaIe.
tenna, about 3,5 mln.
Describedfrom alcoholic specimens'
Frontal prolongation of [ead tiSU-t brown; nasus l9"S
palpi brown, the incisures of intermediate
and conspicobo*;-pale,
the terminal segment chiefly pale. Antennae
segments
reiatively short; basal three segments yellow, the sueceeding
segmentsweakly bicolorous,the basal enlargementsblack, thesegthrough brown_ish
a,pex passrng- yellow to brown; outer^uniformly
darkened. Head. brownish gray.
ments
Mesonotal praescutum apparently-almost uniform g?y or
brownish gray, &nJ stripes if normull present Iittle evid'ent;
poste"ior rZt."ii.r of notum almost uniformlv darkened. Pleura
sternopleuritevarie,gated.
lellow, the anepisternum and ventralof the knobs*itn grayish b""o*n. Halteres brown, the_ar,pices,o-*'ihul paler. Legs with tlre coxae yellow, rrarrorvlr. darkened basaily; trochlnters yellow; femora brown, '&rroyly*o"-e blackened,;tibiae brown, darker'
i;i.- basalll:, the tipl
i,pically; tri'*i black. Wings with ? pale gr?y ground',sparsell'whitish aod pale brown, the latter color inoi,"i.gut.a by"stigrna,bases of outer raclial field ancl the usual
cludiig thei; cell Cui the white inclucles^, conspicuousobliterative;ilsir.eak before cord.,the narrow basesof cells ^lIr. .!"'t-)[ " atrrl
I[r; pale areas on either sirle of the duskl- cubital t'loucl iurtl" ,pot in cell ft at near trvo-thirclsthe length; veins brorvn.
Venation: Petiole of cell JI '1iuf. in ilre obliterative areas.
variable in length, in casessubequalto m. in other specimens
to one-half longer than this element'
Abdomen j'e11ow on basal four segrnents, the tergites
including hypooug...ly trilineate with darker; outer segment-s,
brownish black, the st.yli of the hyPoplg.tunr pale.
lr;Ai#,
with the rnediair lobe of tergilg (fis. l7 , I t\
MaIe nypopi;i.r*'u"Zadbased.,a little narrowed.outwardly-,the caudal
Arp"g;la,ma"gio witS a ll-shapecl notch, the lobules thus formed u'ith
small blackened spinbus points. Outer dististvle- relativelv(
narrow, the basal' portion narrorved. Inner clististr-le Fig'.
ii, ;rt) of characteristic form, tttu outer blaclebroatllv eornp"*rr.a, terminatinE in two slender _acutespines thrrt are tlt'cussatb; outer group of setae numberi'g_ about 3,r to 40;
basal lobe of stl'le d arkened.,lvith a, lateral caritttt.
Yalle-y,Holotype, alcoholic &, Kung-tze-tirg.ga.
_- Tsultrk
Min-shan,'altitudeabout 3028 toe[ett, Julv 20, 193(),at light(D- Huuunr,). Paratopo!)'pes, 3 alcololic JJ.
(Amti
The nearest allies' of ihe present fl1' are Tiltul'a*, (sze-chwan-TibetBorder, &t higfr
tntta) ilesid,iosaAr,uxANDE(EasternSibiria).\'hiehAr,r:xAr(Drrtuttitoaer) urr,l 7'. ( -1) t'((n(tCHARLES l). ALEXANDldri, 32.

'r,HB I'It,ur,rD,\H.

I

have a somewhat similar pale wins-pattern but have thestructure of the male h.ypopygiumquite distinct.
Tipula (Acutipula ) eaptiosa sp. 11.
Male.
Length, about 17 mm; wing, I7,; mm.
Yery closely allied and generally similar io 1'. (A ) pertinaa
sp. n., difrering especially in the structure of the male hypopygium, notably the unusually slender, simple, med.ianlobe of fhe
ninth tergite and the structure of the inner dististyle. Despite
the similarity of the two flies, they fall in two distinct groupsof the subgenus, based on the simple or bifid.nature of the median lobe of the ninth tergite of the male hypopygium.
Size larger than in pertin&r. Male h.ypopygiurn with the
med.ian lobe of the tergite (F'ig. lfl, 9 t) broacl-based,rapidly
narrowed. outwardl.)-,the apex subaeute,densel.yset with acuteblackened points or spicules; basal portion of meclian lobe
and ahnost all of dorsal surface of tergal plate with very
abund.ant setae that are longer arncl coarser on the lateral
portions of eaudal marsin. Inner distist.vle(Fig. lg,id) withthe spines of outer lobe wiclel.ysepara,ted,the setaesurrounding
them reduced in number to I or 10.
rr o I o ty p e, alcoholic d , Kung-t ze-taggu. Tsaluk Yalley,
Min-shan, altitude about 3028 meters, Julv 20, 1930, at lieht(D. I{urvrrrrnr,).
'fipult (Yarnatotipulrr,)
nrorrgolieaAr,r,;xrNr)r.rR.
1 9 2 1 . l I ' i p u l r t t t t r n t g o l i e c i.,\ t . r : x . \ \ r ) r t R ; l ' , u l l . \ l u s .
l 92l : 31t)-:i50.

Hist. \rrt. I'itris,

One broken sltecirnen.Hut.-iertu(iol. Southern llongolia.
The fly is rnost closel.yallied to 'f'. ( ,-.) m()ri(tet'rr
Ar,rxex(Eastern
DER
Siberia).rvhieh nrar-prove t, lre onll a race ofthe present speeies.
Tipula rninensissp. n.
Allied to uariipetiolaris Ar,nxrn Dtr;R, but rnuch larger(wing, &, 2+ mm.); antennae with scape ancl pedicel brorvnish
vellow, flagellum black, the basal segment paler; fenrora black;wings brown, streaked with whitish; abdomen .yellow. thetergites narrowly trivittate with dark brown; h.l'poPyeium
eenerally darkenecl.
MaIe. -- Length, about 23 rnrn.; rving. 24 nrn.; rr,ntenna,
about 5 nrm.
Fore leg, fernur, 13 rnnl.; tibia, 16 rnnr.; tarsus 2I mm.

AriKrv riin zoor.,ocr. BAND 27 L.

N:o 17.

Midclle l.g, femur, LA,smm.; tibia, 14 rnm.; tarsus, 22 mm.
Ilind ltg, femur, 15 rnm.; wing, 20 mm.; tarsus, 27,5mm.
Described from an alcoholic specimen.
Frontal prolongation of head obscure yellow; nasus lacking
or represented only by a low tubercle; palpi brown. Antennae
with the basal two segments brownish yellow; first flagellar
segment brown, the remaind.er of organ passing into black;verticils subequal in length to the segments; terminal segrnent
relativel.y long, nearly one-half the penultimate. Head bro'wn,brighter on front.
Mesonotal praescutum pale brown, with three darker stripes,
the center of the rnedian stripe paler; posterior sclerites ofnotum brown, the scutal lobes variegated with darker; it cannot
be stated as to whether or not the thorax is lrruinose. Pleura
chiefly brownish yellow, the ventral sternopleurite rlarker
brown ; remainder of pleura, including the tlorsopleural regiott,rnore or less variegated with dusky. Halteres pale, the knobs
chiefly dark brown. Legs with coxae d.arkened basally, theapices paling to yellow; trochanters brownish yellow; femora
black ; tibiae obscure y ellow, the tips n amowly infuscated;tarsi brownish black. lVings (Fig. 4) brown, sparsely variegatecl with darker brown and. conspieuouslv streakecl longitudinally with whitish; the clarlier areas inelucle tltt' stis'rntr
and' a confluent seam on atrterior eortl' origi' of /i'*' attd YerYnarrow brown seams alouq' Yeius ('tt, Iit- ,' outer rnetlialveinsand 2ttd A; the whitish areas include cell )[, bases of AnaI
cells; a conspicuous white streak involving cells 7't JIz, bases
or Mr, )u,I lfz and lfr, together with the outer two-thirds of
R r,, other brightened areas before cord. and beyond stigma ;in cells Il, JL, Jfr. J[, and. 1tt A, the Eround is rnore or lessconfined to the centers of the cells, the adioining veins being
broadly bordered by white; veins brown Venation : Rs long;
Rr+,t entire i Ms++ shorter than basal section of Mr; nt, suberlual to petiole of cell trf ,, this character probably variable,
as irr the alliecl rariipetiolcu'i,-:?tt--cu at fork of JIt+t.
Abdonrinal tergites vellow., narrowly but consltieuouslr
trivittate rvith dark brorvn.,the sublateral stripes a little n'icler
than the rnedian; lateral borders of segrnents narrorvlv pale;sternites obscure vellorv, with a broatl dark tneclian stripe;
hypopygium generalh clarkenecl. Male hypoprgiuttr rvith thebasistyl-e and terEite entirelr rlistinct frotu sternite. Caudal
margin and lobes of tergite blacliened.the c'onfonnation muchas in aariipetiolaris. Outer clistist.vle(FiS. 2ll, otl\rnore elonsate
than in uariipetiolaris; the inner dististvle (rrl) of slightl.y different structure; setiferous tubercles on outer margin at nearrnidleneth unusually eonspieuotls. Eiqhth sternite (fig. 2(), ,s's)

C'II^\IiLES P. TILEX,\I{DER,,

32.

THI.] TII'ULIDAE.

rvith the eaudal margin narrorved. ancl mod.erately sheathing,
the apex with a tT-shaped med.ian notch, on either side of
rvhich is a flattened brush of lons yellow setae that are decussate aerossthe midline; above the notch lies a very flattened,
rlepressed.,rned.ian lobe, pale in color, its apex expanded.and.
rveakly emarginate.
alcoholic J, Kung-tze-tagga, Tsaluk valley,
4olotype,
Min-shan, altitude about 3()28 rneters, Julv 20, I gg0, at light(D. Huuunr).
-Ttr- p_resent.fly _is . closel.v relatetl to the comparatively
srna_llTfincln rariiltetiolaris Ar,nxlNDnri (Szechwan-TibetBorder,
at high altitucles), difrering in the rnaior size, black femora
ancl tletails of the male hypop.ygiurn.
T i p u l n d o l o s as p . n .
Geueral coloratiou brorvn; antennae (c/) r.eliltivel.r'long,
basalthree seg'rnents
1'ellow.the renrainclerblack: n,i1Er .l"oogiyinfunrecl. variegated bv restrictecl darker brown at a whitish
etreas, the latter including a, conspicuousobliterative streak
before corcl and. a post-stigmal whitenirg ; Ii , *, entire ; basal
abdominal segments yellow, beyond the ntn bhckenecl; malehypopyglum with the ninth tergite notcheclmediall.l'; basist.yle
entire, its caudal entl llrod.ueeclinto a flattened obtuse tobeor blade.
Male.
LenEth, about 20 nlm. ; lving, 18 mrn., anteppa,
about 6,8 mm.
Describeclfronr nlr. alcoholic speeirnen.
Frontal prolongation of head brorvn. nasusellnEate; palpi
1rale. Antennae (rnale) relativelv elt'rrrgate.if bc-nt backward
t'xtending' to beyontl rving-root' basal three segrnents1-ellow,
the apex of third clarkened,tlre succeeclingsegments-black;
3 tnicroscopic button-like 14th seEnrent; verticils subetlual inleneth to seEtnents. Head brown, the vertical tubercle entire.
Mesonotum chiefly brown, &DI praescutal stripes normallypresent little evident in alcohol. Pleura dark, the ventral
pleurotergite conspicuousl.v .yellow; meral region and ventral
pteropleurite rnore obscure yellow. Halteres pale, the knobs
infuseated. Legs rvith the coxae obscure yellorv; trochanters.-vellow;rernaind.erof legs brown, the tips of fernora ancl outertar,sal seEments darker. Wings (Fis. b) with ilre pr.earculara,ncl costal regions -vellorvi ground-color of wings strongly
infumed. variegatecl bv darker brown and conspicuouswhitish
areas, the former incluclinE the stigma, anterior eord, nt-cu,
ir,nrl Qtttr; the pale areas includ.e large markings before cord,
extending from before stisrna into baseof cell ftr a similar post-

10

Alr,KIv riitt'/,ooLoL+t. BAND27 A.

N:o l7 .

stigmal brightening, in cells /1,,, ll, and liu; veirts brou'tt,
whltish in the obliterative areas. Macrotrichia on veins beyoncl
cord relatively abund.ant. Venation : ftr * s entire ; ft.s elongate.
approximately twice ??t-cu; cell 7n M, IarEe, its outer end'
more or less pointed; petiole of eell lWt shorter than m; n?-(uon nf n iust beyond base.
Abflomen with basal five segmentsYellow, the termina'lsegments, including hypopygiurn, black. Male h.ypop.ygium(Fig. 2l) with the tergite, I t, entirely separate from the ster'nite; basistyle large, complete. Ninth tergite with ^ deep tlshaped median notch, the lateral lobes obliquel-vflat_tened,
when viewed. from the sid.eappearing slightlv decurved'to a'small point; d.orso-med.ianregion of tergite narrowly me_-:branous for its entire length. Basistvle, b. rvith the caud'al
end. extended into a, flattenedlobe, as in polleu ventral portionof basist.ylea, Iittle protuberant, clothecl with tr sroup of longsetae. Outer dististyle a little flattened but still narron'.
Inner dististyle, id, with a, cephalic heavily blackenedportion
cut off from main rnass by white membrane. Eighth sternite,
rg.\.,rvith the cauclal margin broacl, nearly transverse, without
lobes or emargina,tions,but with ^ wide frinEe of lonE vellorv
setae.
Holotype, alcoholic 4 . Kurrg-tze-tuq'gtt.Tsa,luli Vallq'.
Min-shan, altitude about 30?8 rneters,Julr- 2o. t 9iitl. at light(D. Htruunr,).
has a hrpop.rgiunr that is sugrestive of that
Tiptila tl,olosu,
Siberia), but the apicallr'of T. iibit'iertsisAr,nxeNDEn,(Eastern'I'.
(Japan).
poller AT,uxANDER
produced basistyle is more as in
The ver.y different wing-pattern readih' serves to separatethe
spec.iesfrom all sueh forms havirrg this general tfpe of hvpollYElum.

Tipula {initirna sp. n.
Generli coloration bron'n; scutellum J-ellon' rvith tt brou'tt
lredian yitta: rnesepistemnm dark, the mesellirneroll rnore
vellowish; rvinEs strongly tingecl with brownish vellorv. -oblilerative areas t'e".I restricted; basal abdominal seqlnents lellorvthe outer segments bla,ckenecl;rnale hvpolr.vqiuttt rvith c'attdal
margin of ninth tersite tritobed. the obtusell- trunctr.tecllaterallobes blackened.; basist.vle produced. into a blaekenecl point.
Lelgth, about 15 lnm.: wirtq.1{ mm.: antenna,
Male.
about 5,5 mm.
Described fronr an alcoholic specimen.
tr'rontal prolongation of heatl brorvtr; rrelsuslgtg and slettfler : palpi brorvn. Antennae of rnoderate leneth; scape ancl

c H A R , L l t sr ' . A L U x A N D I ; r I , i J z .

THE TIpuLrDAn.

11

ltedicel yellow; flagellurn chiefl.t' tlark brown, the basal enlargements of the segments a, little darker than the rernainder;
longest verticils a trifle shorter than the seglrnents. Heacl
brownish gray; vertical tubercle relativel.r'low. entire.
Mesonotal praescutum ancl scutum chiefly brown (in alcohol), the median region of the latter palei; scutellurn obscure yellow with a, med.ian brown vitta; mecliotergite chiefl1''darkened. Pleura rvith anepisternurn,sternopleuriteand meron
darkened; pteropleurite and pleurotergite yellowish. Halteres
obscure yellow, the knobs more tlarkened, especially theirbases. Legs with the fore and middle coxae darkened, theposterior coxae .yellow, darkened. onl.y at bases;trochanters
yellow; - femora, brown, the bases restrictedly yellorv, the tips
gradually d.arker brorvn; remaind.er of legs passinq.throughbrown to brownish blaclr. Wings relatively broad, stronglrtinged with brownish yellow, the prearcular and costal regions
more saturated.; stignra brown; obliterative areas before cord
lttd be.yond.stigrna verr srnall ancl restricted; r'eins t'ellowish
brorvn. Macrotrichia of veins bevonclcorclrelativelv abundant.
Venation: /1,+..rentire ; petiole of cell II, exceeding nt ; tr[,r+,about one-half the basal section of I[r.
Abd.omenchieflv yellow, the tersites indistinctl,vtrivittate
with brown, the stripes ver.l' nanow and interrupted; sternites
with a, narrow dark line before the equall.vnarrolv pale caudal
bord.er; outer segments,including h.ypopl'giurn,brownish black.
Male hypopygium (Fig. 22) relatively small; tergite entirel.v
separated from sternite; basistvle entire. Ninth tergite, g t,
lonEer than wicle, the cephalic-lateralportions extend.ingbeneath the eiehth tergite; lateral lobes heavill- blackenecl,obtuselv truneatecl: dorsomedian area of tere'ite nAl.l.on,l\'llale
and rnembrallous;a pale rrreclian
tonq'ne-likelolte iutting caudarl
betrveenthe lateral lobes. Basistvle, lt. rvith cauclirlend l)rotrudinE as a small blackenecllobe. the tip alriculate. Ninthsternite with a conspicuousblaekenedlobe orr either sicleof
midline, this clothed rvith short erect setulue. Eighth sternite.
8 s, with the eaud.alrnarEin gentll conrex, clothed rvith lonq
puia setae.
Holot.ype, alcoholicJ, Kansu: Drakana, altitucle about3000 meters, mid-August (D. Huuunr,).
The nearest regional ally of the present fl.r' is 'l'ilnrkr
tloltt.susp. n., which tliffers conspicuousl.yin the u'ing-patterna,nd in the structure of the male hypopt'giuur. especiallr-of
the tergite and basist"vle. The dististvles ancl eiehth sternite
are rrrther sirnila,rin the trvo flies.

r2

aRKrv r,iin zoor,ocr. BAND 27 A.

N : o 1 7.

'Iipula a,bseonsasp. n.
General coloration of head. and thorax brownish gray;autennae blaek, the basal two segmentsa little paler; wings withthe ground.-color pale brown, sparsely variegatecl by pale anddarker brown areas; basal abdorninal segmentsyellow, narrowly
striped. medially with brown, the outer segments darkenecl;
male hypopl'gium with the eighth sternite unarmed with lobes
or setae.
Male.
Lenqth, about 14 rnm.; wing,,13 mrn.; antenna,
about c mrn.
Described from An alcoholic specimen.
Frontal prolongation of head yellowish brown; nasus conspicuous; palpi brown, the second.and third segments paler
at tips. Autennae with scape and perJicelbrown, flaqellurn
black; flagellar segments elongate, rnoderatell-incisecl: r'erticils
much shorter than the segments; outer flagellar segments
broken. Head brownish gray.
Mesonotal praescutum and scutum chiefly brorvnish graf,
rvitltout clistinct pattern (itt alcohol): scutellurn somewhat paler
brown; scutellum and midline of scutunr rvith a clelicatecapillar; brorvn line; mediotergite chiefly ,ilarkened. Halteres pale,the knobs infuseated. Legs with the eoxaebrorvn: troc.hanterssomelvhat paler; fernora pale brown. the tips nArrorvlv l-rrorvnish black; tibiae .yellou'ishbrorvn. the tips niu'ro\\'ll blrrckened;tarsi passing into black. Wings rvith a pale brorvn ground,the prearcular and eostal regions rnorerellorvish: stigma slightly
clarker brown, especiall.y its posterior borcler: pale creamcolored. areas that are but little eviclent, including a, poststigmal brightening in bases of cells ft, and 1?r;very restricted
obliterative areas at the cord. and brightened. spots before andbe.yond rnidlength of cell II, adjoining vein Cu; veins brorvn.
Macrotrichia of veins beyond. cord relatively abunclant but notincluding an)' section of Mt+t or the basal fourth of R n rt.
Venation: lir-r-: entire; petiole of cell 1l[, about one-half lonser
than m.
Abclornen with basal five or six segments yellon'. bothtergites ancl sternites rvith a clelicate median Lrrorvn line thatis slightly interruptecl at the sutures: lateral tersal stripes
obsolete; terminal abdonrinal seEnrentsblownish black. Malelrypopygium (Fig. 23) with the tergite entirelv separate fromsternite; basistlle, b, large, the suture sepirrating it from sternite slightlv interruptecl; ventro-cauclirl portion of basistylervith nurnerous yellow setae, but the apieal portion unproduced.
Ninth tergite, I t, rvith the eaudal marqin shallowlv emarginate.
thc rned.ian reEion rvith a further small U-shapeclincision:

cHARLlrs r'. ar,ExANDER,32.

T H t r ir r p r r r , r D A n .

18lateral lobes thus formed broad, obliquel,l' truneated, the outerapical angles a trifle produced; ventrad. of this point a seeond.
small lobule; mid-line of tergite narrowly but entirely divjded
by pale membrane. Inner distist.yle, id, deeply ineised on
posterior margin. Ventral portion of ninth sternlte , g,e, rvitha stout pendulous lobe, clothed with conspicuousyellow setae.
Eighth sternite, 8 $, with caudal margin truncate or ver.y feebl,v
emarginate, without marginal setae.
Holotype, alcoholic &, Kansu: Drakana, altitude about3000 meters, mid-August (D. Hurvrunr,).
_ Despite the somewhat different general appearance ancl
details of structure of the ma,le hypopygiurn, as tne unannecl
eighth sternite, I believe the nearest allies of the present fl.v
to be Tipula dolosasp. n. and T. finititna sp. n., rvhich diffei.
conspicuously in the wing-pattern and strueture of the hvllop_ygium. The irrter-relationships of these three slrecies a"eshown by the sotnewhat peculiar structure of the inner clistist;'le, rvhich is deeply incised on lrosterior nrargin to eut off
a large pale basal lobe.
Tipula maearta sp. n.
- Be.Iongsto the eru,ciatagroup, allied to tttultittettsis;generalcoloration of head and thorax dark; abdomeri with baJal fiveseg_mentsyellow, rvith a narrow median stripe on both tergites
and sternites; outer sesments,including hypobygiurn, darkened;wings rvith
at or close to fork of fii; gieral colorationof wings brown, varieeated b.r- rvhitish falciae. including avirtrrall)- complete band ber ond corcl; aPex of x,inE uniforiirlv
darkenetl; urale hvpopvEium n'ith outer. busal lobe of innerdistist;'le a broad flattenetl structure. its'tip one or tn'o spinotrs
points.
Male.
L e n g t h , a b o u t 1 3 - - 1 + r n r l r . :r v i n g . l 3 - l + l n n l .
Female.
L e n g t h , a b o u t 1 5 - 1 7 r n m . ;r v i n s tl' 4. --lir,': mnr.
Describeclfrom alcoholic specimens.
Allied and generally similar to 'l'ipulctntupinel/,r?.\.,
differins.
especially in the structure of the inn-er dististyle of the malehypopygium.
Frontal prolongation of head brownish 5'ellorv. Antennae
relatively short, the flagellar segments weaklr- bieolorous.
Head dark.
Thorax chiefly dark, probably pruinose in living speci*g-ns, the praescutum presumably with darker stripes. Femo"ayellow to obscure yellow, the tips narrowly dark 6"on*'n;tibiae
yellow, the tips narrowly darkened; tarsi brownish black.
Wings (Fig. 6) brown, handsomelv varieEated.bv rvhitish, the14.\IiKrv l.'iin zool,ocl.

r r A N D2 7 A . N : o 1 7 .

Iatter including broad transverse fasciae before and beyond.
cord, the former comprising a, crescentic mark that extends
from before stigrna into the entirely pale ceILCu; band beyond
cord cornplete or virtually so; cells C and Sc more yellowish
brown; stigma and a cloud on anterior cord. darker brown.
Venation: Rr+, entire; r-tllx'ioining Rs at fork, rrrore rarely
before or beyond this point; cell 7* M, small, pentagonal.
Abdomen with basal five segmentsyellow, both tergites andsternites with a, narrow dark brown median stripe that is interrupted by the pale eaudal marsins of the segments;outer segrnents, including hypopygium, uniforml.r' blackened. Male hypopygium (FiS. 24) with the tergite and basistyle entirely separate
from sternite. Ninth terEite, I t, extensive,Eradually narrowed.
outwardly beyond.midlength; caudal margin with a very shallow
U-shaped emargination, with a, small rned.ianpoint; eclgeof
emarglnationn&rrowly but heavilv blackenedand polished;clorsal
surface of tergite with scattered,delicate, pale setae; as in manyother allied species, the tergite is composedof two closelv
applied plates, each having a, quite identical caudal border.
( iuler dististyle slend.er,sinuous at base,the distal end.dilated,
obliquely truncated.ancl providecl with long eonspicuoussetae.
fnner dististvle rvith the basal lobe broarl, the apical angle
into one or two spinous points (irl). Eighth sternite,
lrroclueed.
portiorr,9,ermoderately sheathing,the slightlr- producetltrtecliatr
with a, fringe of long yellorn-setae.
Holotype,, alcoholic J ,, Kung-t ze-tasga, Tsaluk Yalley,
Min-shan, altitude aborrt 3028 meters, July 2(1, 1930, at light
1 C , numerous(O. Huuunr.,). Allotoptotype,9. Paratopohlpes,??. Paratgpres,4 99, Tsang, Drakana, altitud.e about 3000
rneters,rnid,-August(D. Huuunl).
T'ipula, ntaiarta is most nearly allied to T. _nrupinensis
Ar,nxarlDnn ancl 'f'. pedi,cella,risAr,nxa,xDnti, both from the
Szechwan-Tibet Borcler, at high altitudes. It differs tnost
conspicuously in the structure of the basal lobe of the iturer
in both of the sperlistiitvle of the rnale hypopl-gium,which 'l'.
ct'ttciatrtIh)\YAItI)scies above listed is a slenderacnte'spine.
(sihkirn) is cluite rlistinet in the rnore variegateclu'irtg-patternuncl in the struetureof the ttttrle hr'1ro1,'t'g'iutrt.

Nephrotolllit tlrakitlta' slt. ll.

antennae rvitlr scap(' tt,ttclpedieel
Allietl to ltir,r-uticuu(lct:rlark brown; herrrl rvith a blaek ttrerlian stlipe extending fromn'ith three black stripes.
pr?rescuturrr
vertical tubercle to occiput;-celrhalicends turned. Iaterad; posterior'the outer ones withselerites of loturrr n'ith a rrrerliarrltlacl< r'itta: pleurer eottsl-ri-rJcHARLES t,. ALExANDEII,32.

'rHIt r.rpuLrDAE.

15

cuously variesated with black; wings subhyaline;stigma small,pale brown;-abdomen(male)almost entirely black; male hypopygium with the eighth sternite deeply incised.and provided
u'ith abundant elongate setae.
MaIe.
Length, about 12 mm.; wing, g rnm.
FemaIe. - Length,about 13-14 mm.; wing, l1_-12 rnnr.
Described from alcoholic specimens.
X'rontal prolongation of head blackish d.orsallv. morebrownish yellow on sides. Antennae with scape and ped.icel
dark brown; flagellum black, the segments moderatel.yincised.
-Head yellow, with a, continuous black median streak extending frorn summit of vertical tubercle to vertex, narrowed
to a capil]ary line behind the tuberele, on vertex expanded.
into an elongate shield-shaped.area; a srnall black spot on
orbits at narrowest part of vertex.
Pronotum mediall-v obsenre vellow. blackened on sides.
Mesonotal praescutum obseurevellow rvith three polished black
stripes, the outer pair rvith elnterior t nds directed laterad, incases with this portion paler in color; scutal lobes extensively
blackened; scutellum and nrediotergite brownish .vellow, with?t median blackish vitta that is expanded on posterior borderof the latter sclerite. Pleura browrrish yellorv, with corrspicuous black areas on propleura, ventral anepisternum, vential
sternopleurite, cephalic portion of pteropleurite, ventral meronand less distinctly on pleurotergite. Halteres obscure brownish
vellow, the base of knob more blackened. Lregswith the coxae
blaek, the nrid-coxae a trifle paler ut apicesl trochanters obseure yellow; fernora vellow, the tips narrowl.y blackenerl:
tibiae obscure lellow, the tips slightlr rnore blackenecl; tarsiblack. Wings subhvaline; stigrna.small, pale brou'n; cell ,5'(.
light r-ellorv: ir sr.areelv inclicatecl lrrou'n eloucl on anter.ior
cord: veins brotvtt. ,Fischieflr- lrirle trnrl sulrobliterrrterl.
Yenation : Cell il.,, rather narron lr- sessile.
Abdomen, including lt v po1,r'giunr.t.hieflr' trlirck. the subla,teral portions of tergites olle unrl tn'o iL little paler. Irr
female, the stenrites are sonrervhat lraler sublaterirlll-. Abtlomen (rnale) rvith the ventral surface of sternites rrilh con.
spictto_userect setae, beeorninE yerv long'oll tlre lrrofouncllyincised..eighth sternite. Male hr-pol- rvs'iurrr rvith the tereite(I'iq. 2b) cleeply notchecl rnediallr.
H rlot'u1tt:.alcoholic J ,, Kansu: Drakana. ultitutle tr,bout;](xx) nreters. micl-AuEust (D. Ilulr:rrnr,). ^llloto1irttry1tr,.
?. parato1iotry1r:.
?.
fefltrrttoma rl,t'uA,anri'is allied to ^\-. hir.stttiettuda
AL,nxerI)l.lR (Norther.l- Japan), the Eelrera,lap;reetranceof the hypo-

1(iA R , K\Ir r i i n z o o L o c r .

BAND 27 L

N:o l?.

pygia of the two being very similar. The coloration of thepresent flr- is verv different frorn that of hirsutir'auda'
Nephrotoma Perobliqua sP. 11.
General coloration yellolv, the praescutum and' scuturrt
variegatecl with polished black; antennal scape and peclicel
yuttoi', the flageilurn-black; p-ostnotal mediotergite wilh cgli"al portion pitu ; pleura yello*,- scarcel.y variegated'^with- wings'pale ybllolu, tire cord. and outer section of veindarker;Cir, togethEr ivith the wing-tip, pale brown; stigma. d'arker
brown;1rr-ru long and.onosoallv oblique in position; abdolnerr
with a rnedian dark striPe that is interyello*, the tergitls"sutures;sternites with three sueh stripes, t\eiuptetl' at thelateral ones paler; terminal segmentsmore unifonnlv blaekened:male hypopygium with the ninth sternite bearinE a small[obl,_ the lip pale T4 obtuse'pendulous'
MaIe. - Length; about 12 mm'; wing, 11,r rnm'
Described from an alcoholic specimen'
Frontal prolongation of head obscure yellow, the rnidd.orsal resion ou""odlv darlrened.;nasus black; palpi bl-a-ck..A.ntennae n,ith scape ur..l pedicel light vellow; flagellurn black. theproximal end. of the first seqtteut a little Pttler':flag-ellarsegments mocleratelv incised. Head I'ellorv, the occiPita,Ibrantl
brownish black, triangular, sencline,, ttuttou' da,rkline -cephaladto summit of vertica"I tubercle; genae more or less clarkened.
Pronotum light ,vellow med.ialll', more -darkened on sicles.
Mesonotal p"u.r.-otum .yellow with ihree polished black stripes,
the lateral pai" straight or with only- a weak brown suffusion
Iaterad of iUrir ant6rior ends; scutal lobes conspicuouslyvariegated with black; remainder of mesonotum yellow, the-mepale yellow- mediallv' more infuscated on
dio-tergite broaclly
either side. pr.i,ra yeilow, onl.v vaguely rnarked with darker
along the suture between the mesopleural episternurn antl
epiniron and. on the ventral rneron. ilalteres pale, the knobs
bicolored, lrartlv clarkened, partlr rvhitish. Le-gs rvith $e
coxae and iroc6anters vellowi f.*o"u obscureyellow. tlte tips
tibiae brownish-rellorv'
or""o*ly but conspicuousl.l-black_erred;
the tips blackenedl tarsi black. Wings (Fig.,7) pale lellowish,
the piearcular and costal region,s n;e"1-.pale I!|lor1 ; stigmaoval, clark brown; wing-tip b"roadlvbut iirsensiblr darkened;narrow brown clouds on uni.rior cord, nt--(tt and d'istal sectionof Cur; veins brown. Venation : Cell 7n l[Z of rnod'eratesize;
ceLIlltrver.y narrowly sessile;nt--a{ unusually l^olg and oblique'its caud.al' eod lving some distanee basad of level of oriEi'
of J?s.

c H A R r . , E Sp . A L E x A N D E I i . S 2 .

17

THE'frpuLrDAE.

Abdominal tergites yellow, with a conspicuousblack meclian
stripe that is narrowl.y interrupted at the incisures; lateral
stripes lacking; sternites yellow, narrowly and interruptedl"_v
trivittate with black, the lateral stripes paler brown and morebroken; segmentsbeyond the sixth, including the hypop,ogium,
black or brownish black, only the dististyles yellowish. MaIe
hypopygium (F'iS. 27) with the tergite (FiS. 26, I t) black, thecaudal margin with a rounded U-shaped.notch, each inner angle
prod,uced.
into a small blackenedspine; viewed laterally, below
the apical spine is a more slender one that is directed.more
ventrad. Ninth sternite, I s, rvith a median pendulouslobe,
slenr:ler,darker at base than at the white obtuse tip, the lobe
directecl ventrad and slightly cephalad; in a, position of rest,
this lobe lies in the small median emargination of the eighthsternite, the latter with only inconspicuous and relatively
sparse setae.
Holotupe,,alcoholic C, Kung-tze-tagga,Tsaluk Yalle.v,Minslran, altitucle about 3()28 nreters.Jul.y 20, 1930, at light (D.
Huulrnr,).

The pi'esent fly, while similar in certain regards to species
sueh AS Nephrotoma bispinosct,Ar,nxlNDER,, is quite distinct
from all regional species so far made known.

Lirnoniinae.
Limonia (Lirnonia) quadrinotata(Mnrcnx).
1818. Lim,nobia quaclrinotataIIErcinx; S;'st. Ileschr., l: 1{{.

Two rnales, Kung-tze-tagga,, Tsaluk Valley, Ifin-shan.
altitucle about 3028 meters, July 20, 1930, at light (D. Huuunr,). I c&n find no differences between these specimens ancl
abundant European material. Moreover, the Nearctic L. (L)solitariq, (Osten Sacken) is verv closely allied but mav perhaps
be maintained as a, valid subspeciesor very nearlv allied vicarious species.

Limonia (Dicranornyia)kinensis sp. rr.
Male.
Length, 6,b--T rnm.; wing, ?,;-tl rnnr.
Fernale.
Length, 8,J-9 mm.: wing, 8,;-10 lnln.
Described from alcoholic specimens.
Belongs to the tristis group; most nearly allied to L. (D.)
tc'istoidesAr,nxeNDER (Manchuria), differing especially in the
Iarger size and structure of the male hypopl'gium.
Arkit, 16r zoologi. Bd 27 A.

N.'o 17.

,

18

ABKrv niin zooloer.

BAND 27 A.

N:o 17.

General coloration of entire body dark brown, und.oubtedly
gray pruinose in fresh speeimens, the ventral dististyles ofthe male hypopygium light yellow. Antennae and legs black.
Ilalteres pale, the knobs dusky. Wings (Fig. 8) whitish subhyaline, the surall stigma darker; in somespecimens,especiallyfemales, the cord and outer end. of ceII lt' M, are narrowly
seamed with darker; veins much darker and more distinct
than in tristoides. Yenation: Sc, ending a short distancebeyond the origin of fis, Sc, shortly remoYed.from its tip;cell 1't Xf, subequal to or longer than vein Mr+2 beyond it;,ttt-l,tr, at or close to fork of l)I.
Male hypopygium (Fig. 28) chiefly black. Ninth tergite
transverse, as in tristoides, the caudal margin very gently
emarginate. Basistyle with two small accessorylobules andhair-pencils, ad,ditional to the usual ventro-mesallobe, rvhich
is provid.ed with setae at apex but otherwise unnrodified..
Dorsal dististyle entirely blackened,the apex acute. Ventral
d.ististyle, ud, higher at basethan in tristoides,,narrowed gradualty to the intensely blackened rostral prolongation, the latterool sud.denly narrowed at point of insertion of the rostralspines, as in tristoides; spines with basesless elongate thanin tristo'ides. Gonapophyses,g, with the rnesal-apicallobe
and more strongl.y curvecl.
IongerHolotype,alcoholic d, Kung-tze-taqga,Tsaluk Yallu.F,I\Iinshan, altitude about 3028 meters, July 20, 1930, at light (D.
several cfg.
Huunnr). Allotopotype,9. Paratoltotypes,
The distinctions between the presentspeciesand tristoide,'
have been given above. It may be noted further that the
male hypopygium in tristoidesis yellow, whereasin the preseut
fl.v only the ventral dististyle is brightened.
I-,imoni:t(Dieranornyia)sjiistedti sp. n.
General coloration (itr alcohol) clark, probably pruinosein
antennae dark throughout; femora brownish
fresh specimens;
yellow, the tips narrowly clark brown; wings whitish, with a'*putre brown pattern; Scr Yery long, exceeding-Bs;nrale_llpo.
notched rnetliallr';
pygium with the ninth tergite conspicuo-usly"ost"al prolongation of ventral d.ististyle suddenlr- nanowed,
the two rostral spines placed near its tip.
Length, 6-6,5 mm.; wing, 7 - 7 . s r l t l l l .
MaIe.
L e n g t h , 7 , 5 - 8 , 5 m m . ; w i n E , 8-8, s mrrr.
Female.
Described from alcoholic specimens.
Rostrum brownish black; palpi black. Antennae withscape and pecLicelbrownish black ; flagellum black; flagellar
seginents oval, the outer segmentsbeeoming more elongate-

cHARLES p. Al,ExaNDEri, 32.

THE TrpuLrDAE.

lgoval, the terminal segment elongate, exceedingthe penultimate;verticils shorter than the segments. Head dark, probably
pruinose in fresh specimens.
Mesothorax uniformly blackened, presumably pruinose in
fresh specimens. Ilalteres pale, the knobs weakly infuscatecl.
Legs with the eoxae pale, the outer faces darker basally;trochanters yellow; femora brownish yellow, the tips narrowly
dark brown; tibiae brownish yellow, the tips narrowly darkbrown; tarsi passing into black. Wings (Fig. 9) whitish, with asparsebrown pattern that is much heavier in the female sex;thedark areas are distributed as follows: Stigma and a, confluent
searn at fork of fis; Srr; origin of fts; cord and outer end ofcell 7o Mr; seam along vein Cu; in the more heavily-patterned
individuals, the tips of the AnaI veins and the axillary region
are likewise darkened.; veins brown, the basal half of trt yellow.
Yenation: Sc, ending opposite origion of 11.s,.9c, far from thetip of ^5'c1,the latter alone about one-thircl longer than Bs;
l1t-ctc at or before fork of Llt.
Abclornen dark brown, the ventral dististyle of male hypopvgium pale. Male hypoposium (Fig. 29) with the tergite
ver.\- d.eeply notched. medially, the emargination narrowly Ushaped, or, on slides, even appearing V-shaped. Yentromesal
lobe of basistyle, 1,, relatively large but unmodified. Dorsal
dististyle pale, strongly curved., nanowed to the acute tip.
Ventral dististyle, ud, with the rostral prolongation abruptl.v
narrowed., with two slightly unequal spines close to tip, these
not arisinE from basal tubercles. Mesal-apical lobe of gonapophvseslong and. narrow, pale.
Holohlpr:,alcoholic €, KunE-tze-tagea,Tsaluk Yallev, IIinshatr. altitucle about 3f)28 rneters,July 20, 193().at lisht (D.
Ht'nrrul,). ^lllotoprtttlltr',
several 4?.
9. Purutoltotrype,*.
I take verr great llleasurein narningthis distinct Limottiu
in honor of Dr. Yxcvr,: S;liislnnr, to n'horn I arn highly indebted for mueh kindlv co-operation in stud.l'ing the Tipulidae
of various regions. The speciesis readily told frorn all regionalsirnilar fornrs b.y the characters diagnosed above, especiallythe structure of the male hypopl giunr, as the very d.eepllemarginate ninth tergite, the clistal position of the rostralspines ancl the long slender apical lobe of the gonapophysis.
Antoeha (Antoeha) nebulipennisAr,r:xANDr.rR.
1s31. Antocha (Antochct)nebuliltennisAT,ExANDER;
I'hilippine .Joum.
-1{:;ti2-353, pl. 1, fig. 9 (venation);pl. 2, tig.+t ,.d hypopygium).
Sei.,

Described from Mupin, Western Szechwan, altitude 3500
feet, June 3(), I 929, collecteclbv ReverendD.q.vrnC. Gnl HAnr.

20

AriKrv riin zoor,ocr.

BAND 26 A.

N:o 17.

fn the present series are a few specirnensof both sexes,Kungtze-tagga, Tsaluk VaIIey, Min-shan, Kansu, altitude about 3028
meters,Jul-v 20, 1930, at light (D. Huuunr,).
Ifeolimnophila lirseoeubitalissp. n.
General coloration gray; praescuturnrnore yellowish grav,
with three brown. stripes, the median one often dividedt b.y a,capillary pale vitta; wings whitish, the stigma and a brbad
conspicuous seam along vein Cu,brown; costal border not or
scarcely darkened; nz close to fork of lt.
Length, 7-8 mm.; wing, 8-9 mm.
Male.
Length, 7,5-8 lnm.; rving, 7,5-8,5 nun.
Female.
Rostrum and palpi gray. Antennae with scal)eand pedicel
black; flagellum brownish black. Heacl gra)'.
Mesonotal praescutum yellowish gray with three or four
brorvn stripes, the intermediate pair in easesunited into asingle vitta or nearly so; posterior sclerites of notum clearergra). Pleura gray. I{alteres pale, the knobs weakly darkened.
Legs with the coxae pruinose; trochanters brownish yellow;
remaind.er of legs black, the extreme basesof fernora ^ trifle
paler. Wings (Fig. 10) whitish, the stisma ancl a broad conspicuous sealn along vein (.'u brorvn; costal borclel not or
scarcely clarker than the rernainclerof clisli. in eusesrvith iL
slight clarkening in base of cell ,5c ancl at h; in heavilrpatterned specimells, the anterior cord. is narrowly clarkened
but the posterior cord is uncloudecl; veins dark brown, Xf
chiefly pale yellow, especially at base. Venation: Rz close tofork of Rz13
*4, usually just beyond; r-t?t somedistancebeyond.
fork of lls; tn-cu approximately its own length beyond.fork
of II.
Abdomen gralr rnore brownish in female. Male hylropygiurn with two interbasal spines on basistyle, these much
shorter than in alticola and almost equal to one another inlength and d.iameter.
Holofu1pe,&, Yu-Long-Si, Szechwan-TibetBorder, altitude
15,600feet, July 28, 1930 (D. C. Gnrnru); in U. S. National
72 d?, Julr' 2EMuseum. Atlotoltotupe,9. Pat'atopotglpe-*,August 7, 1930 (D. C. Gnrnru). Parahlpes.??. Yin-KuanTsai, altitude 13,000-15,000feeb,July 2-D,193o (D. C. Gn,eulu); 15 alcoholic9?, Kung-tze-tagga,'Isaluk Yallev, Ifin-shan,
Kansu, altitude about 3028 meters, Julr' 9(). 1930, at light(D. Huuunr.,).
I have had this speciesbefore nre fronr the United States
National Museum and take the opportuuit.y of describing theis most nearlrflv at this time. Neolintnopltila.f:u.*t'0t'ttbita,lis

cHARLEs r'. AI,ExANDER,32.

THE TrpuLrDAE.

2L

allied to I[. fu,sc'itteruisEowenus (Yunnan., Szechwan-Tibet
Border) which is distinguished. by the much heavier wingpattern, the eostal region being darkened.,the entire corcl andouter end of cell 7't lL conspicuously seameclwith brown,
and the posterior borcler of wing narrowly but entirely borderedby clusky. The present fly has the posterior corcl, outer encl.
of ceII l-utnfz and the costal and posterior wing-bord.ersclear
or virtually so. The general effect is thus one of ^ wing
nnrnarkecl except for the stigma and a, conspicuousdark seamzr,Iongvein Cu. One specimen of the type-serieshas both ,nt,
and the basal section of ftu lacking in otre winq. only.
Drioptera (Psiloeonopa)tritlentieulata sp. n.
General coloration yellorv, variegated by clarli brown; antennae with scapeyellorv, peclicelancl flagellurn brownish black;mesonotal praeseutum with three confluent striltes; thoracic
pleura stripecl, more heavilv so ou ventral portion; legs black;wings vellorvish rvhite. rvith verJ'restric'teclpale brown seams
olr. certain of the v eitrs; ,Scei ust be,vonclorigin of -Il.s; veins
beyond. cord long ancl parallel; cell 1't JI2 closed; Anal veinsdivergent; male hypopygiurn rn'ith outer dististyle bearing threespines at and near tip.
MaIe.
Length, about 3,i-3,T mm.; wins'' {-4,2 nr.rn.
Described from alcoholic specimens.
Rostrum and palpi brown. Antennae short; scape lieht
vellorv,the remaining segmentsbrownish black; flaqellar seg'ments
oval, rvith verticils that are subequalto the seEmentsor netrrl.yso. Heatl clark brown, possibly pruinose in fresh speeinrens.
Pronotum yellorv, narrorvlv brorvrr nrecliallv. Ilesonotum
rellorv, conspicuousl.vvatiegatetl lr.t rlarli brorvn; praescutum
rvith three confluent strilles. the rneditur one rnore or lessclivid.eclbv a caPillar.v clar*er vitta that is best inclicated in
front; posterior sclerites of notunr chiefly darkened,the meclian
region of scutum, rnost of scutellunr and antero-lateral angles
of mediotergite more vellowish. Pleura yellow, variegated byd.ark brown, that is clistributed.more or less as stripes; themore dorsal stripe very narrow and interruptecl, on the suture
between anepisternurn and sternopleurite with posterior end.
arcuated dorsad, continued. eaudad even less evidently onto
the pteropleurite; ventral darlrening much wider and rnore
conspicuous,including the sternopleurite and meron. Ilalteres
generallv pale, the knobs weakl.r darkenecl. Legs with thecoxae obscure yellow, the outer faces more or less infumed;
trochanters vellow; remainclerof leqs black throughout. Wings(Fig. I t ) rellorvish white, rvith very restrieted pale brown

22

A R K r v n i i n z o o r , o c r . B A N D2 7 L .

N:o 17.

se&ms on cord. and as marginal spots on Rr+, and Rr; veinsbrown, the bases of M and.tt A fading to white and becoming
almost invisible, Macrotrichia abundant and well-distributed onveins beyond.cord., basad of cord more restrictecl; basal sectionof M and 7st A glabrous. Yenation: Sc, 'iust beyond originof fts, Sc, thus very long, exceeding fis; Rz and Rr+t subequal; veins beyon,clcord. unusually straight and parallel, thecells cleep; cell 7't I[, shorter than any of the veins issuingfrom it; ?t1straight to strongly arcuated; m-cu nearly its own
length before fork of llI; Anal veins diverging, E*t A gentl.yconvex toward. 7rt A.
Abdominal tergites chiefly brown, the pleural region andsternites more yellowish; hypopygium obscure yellow. Male
hypopygium (F.ig. 30) with the two dististyles terminal; outerstyle, 0d, blackened, terminating in two equal spines, with athird similar spine on outer face 'iust back from apex; surfaceof style without punctures; inner distist.yle, id, subequal inlength to outer, a, trifle dilated ab outer end., the surface withseveral microscopic punctures. Gonapophyses,9,, appearing as
pale flattened blades, the mesal-apical lobe a gentl.y curved.
blackened spine.
Holotype, alcoholic d, Kunq-tze-tage'a,Tsaluk Yalle.v,Minshan. altitucle about :]028 rneters,Julv 20, 1930. at lisht (D.
Huuunr,). Paratoltotglte,d .
There is no near regional all-v of the present fl)', rvhich
is read,ily told by the venation and structure of the malehypopygiuur. The venation reminds one strongly of Helobict
and allies, and perhaps, the present species would be better
placed.in this complex. Ifowever, it is becoming inereasinglv
appareut that aII of the genera of Eriopterini with pot-belliedmeron are elosely inter-related and it is ver.y possible that
Helobia ma]' later on be placecl as a subgenus of Erioptera.
llrioptera (Bmpeila) prelata, sp. rr.
Size ver)' large (wing, female, 6,8 mm.); general color:ttion
dark; halteres pale rellon'; leEs black tlrrouqhout; u'ings paleyellowish gral ; stigma faintlv indicatecl; ,SclonE, Jl, rtot t'ottspicuouslv oblique, cell Mt open lr)- atrophv of tn.
F e r n a l e . L e n g t h , a b o u t ( j , ; m l n . ; r v i n E . ( ; . 8X 2 . t ; l t r l l r .
Described.from an aleoholic specirnen.
Rostrum and. palpi black. Antennae black throughout,Head brorvnish black.
Thorax clark brown, the dorso-pleura,lregion paler; infresh specimens, the coloration is ver.) probably pruinose.
Halteres pale rellow. Legs black throughout, in fresh spe-

cHARLEs p. AT,ExANDEB.32.

THE TrpuLrDAE.

23

cimens with the coxae probabl.y pruinose. Wings (Fig. L2)
pale yellorvish graI, the prearcular and, costal regions clear
light yellow; stigma very pale brown; veins pale brown. Yenation: 8c relatively long, Sc, ending about opposite threefourths fts, Sc, a, short distance from its tip; -Iluoblieuo, subequal to Rt+t+ni Rr+2,and /lr+n subequal;veins nB and n4
only slightly divergent, cell R, at margin being nearly two-thirds
ftr; cell fI, open by atrophy of m; nt-cu shortly before forkof M; vein ?ndA divergine from 7't A throughout its entire
lengthAbdomen clark brown, the pleural membrane paler. Ovipositor with cerci elongate, gently upcurved..
Holotgpe,alcoholic ?, Kung-tze-tagga,Tsaluk Yalley, Minshan, altitude about 3028 meters, July 20, 1930, at light (D.
If uurvrnr,).
The present fly is well-d.istinguishecl
frorn all allies in therelatively great size, the wing-area of the fernale being about
trvice that of the nearest all.v, Ilrioptera (Iinryetla) ,japonica
Ar,nxaxDnn. This latter specieshas the trichiae of the wingveins longer ancl more conspieuous,ancllvith certain details ofvenation quite distinct, as the shorter frs, much longer andrnore strongly arcuated. Ru, and. strongly curved.Mn, which atnear midlength bends markedly towarcl the wing-tip. fn thepresent fl.y, n[, is nearly straight throughout its length. The
regional E. (8.) c*aillantiAr,uxLNDEn(Mongolia)is well-distinsnished.by the short S'c and closed.cell 7't II.

Hxlllanatiolr of ltlates.
I'late I,
Fig. l. Tipulu r l-estiplet')subcitrinutu-\r,nx-.rxDEn;uirrg.
,
2. Tipu,la ( l-estiplt,r) ltunr,mclisp. u.; rving.
ril. Ti'pula (Vestiplefl heclinisp. n.; rving.
,
1. Tiltula nfinensissp. n.; rving.
,
5. Tipula dolosasp. n.; wing.
>i g. T'ipula ntacarta sp. n.; wing.
,
7. Nephrotoma,
pet'obliquasp. n.; rving.
g. Li,nrcnia (Dicranonty'ia) kinensiasp. n.; rving.
r"
9. Limonia (DicranomEia) s.jdstedtisp. n.; wing.
, 10. Ieoli,mnophilafuscocubital'issp. n.; rving.
11. Erioptera (Psiloconopa)tridenticulatctsp. n.; u'ing.
12. Erioptera (Empeda) prrelata sp. n.; rving.

Plate II.
Fig. 13. 'I'iltulu (Vestipler) subcarinata Ar,rx,rNDER; urale hypopygiun,lateral aspeet.
"
1 4 . Tipula ( Irestipler) subearinuta AnnxANDITR;the s&me, dorsal.

2+

ABKrY rtin zool,oer. B A N D 2 7 L .

N:o l?.

Fig. 1D. Tipula (Vesti'pler) subcarinata Ar,nxaxDEB; the same, ventral.
,
1 6 . Tiputa (Irestipter) hedini sp. n.; male hypopygium, details.

,
,
,

>

L 7 . Tipula (Acuti'pu,la) pertinan sp. n.; t_hesame.
1 8 . Ti,pula ( Acu,tipttla) eaptiosasp. n.; the same' ninth tergite.
1 9 . Tipula Qcutipt3tla) captiosa sp. n.; the s&me, inner dististyle.
20.

Ti,pula mi,nensissp. n.; the s&me, details.
Plete III.

Fig. 2L. Tipula dolosa sp. n.; male hypopygium, details.
,
22. Tipula finitima sp. n.; the same.
" 23. Tipwla absconsasp. n.; the same.
,, 24. Tipula macarta sp. n.; the s&me.
" 26. Nephrotoma drakan@ sp. n.; the s&me,ninth tergite.
, 26. Nephrotomaperobliqua sp. n.; the same.
,
2 7 . Nelphrotomaperobliqua !p_.n.; the same' lateral aspect.,
28. L'o:monia(Dicranomaia) ki,nensissp. n.; male hypopygium') 29. Li,monia (Di,cranomyia) si\stedti sp. n. ; the sane',
30. Eri,optera (Psilieonopa) triilenticulata sp. n.; the same.

Explanation of Symbols.
(Hypopl'gial Chnracters.'e, :D:
ild, :
.q:
icl :
od :
p :
s:
fad :

ndeagus.
basistyle.
dorsal dist'istyle.
gonapophysis.
inner dististyle.
outer tlististyle.
phallosome.
Bternites.
torgites.
ventral tlististyle.

Tryckt den 22 februari 1934.

Upprult 1934. Almqvist & Wlkrolls Boktryckeri-A'-B'

A r r , K r vr i i n , z o o l , o c r .

BAND 27 A.

Itig. | - I2.

N:o 17.

PIJ.

I.

ABKry rtin zooloor.

BAND 27 L.

N:o 17.

PII.

II.

);;\,4

-\

n

IE

20
F'ig. 13-20.

AriKrv riin zoor,oer. BAND27 L.

N:o 17.

PL. III.

26

-t
vi

Itig. 21-30.