5lz
NEW SPECIES OF' CRANE FLIES FROM SOUTIIAMERICA. PART III.
(Family Tipulide'

Order Diptera)'
Crreurs

P. Ar.rxexoEn.

RTPAINTIID

FROI(oFl TEE ENToMorpcrcAr,

Amyars.

Vol. XXII,

SocrETy o! AuERrca,

No.4.

Coluuaus, Oaro,
December, r9e9.

tg29

NEW SPECIESOF CRANE FLIES FROM SOUTH
aMERICA. PART lrr.*f(F'amily Tipulida,

Order Diptera).

""^x'ii*,1fi:lJ"'.'
The great majority of the species discussed at this time are
ftom Argentina, collected by Messrs. Bruch, Shannon andWeiser; Paraguay, collected by Schade; and Ecuad,or, colTected,
by Tate. The interesting Tipula is frorn Colombia and wasincluded in extensive collections of crane-flies belonging to the-bv
British Museum, kindly loaned to me for studv
Mr.
Edwards. Except where stated to the contrary, the typesof the new species are preserved in my collection.

Alexander: South American Ti,ulid@

769

tibie brownish yellow, the tips very narrowly infuscated; tarsi elongate,
pale brown, gradually passing to brownish black. Wings yellowish
gray, the prearcular and costal regions bright yeilow; stigma pale
brown, a little darker than the ground-colorl veins yellowish brown.
Venation about as in other members of the macrosterna group.
Abdominal tergites reddish yellow, the intermediate segments witha shiny black median triangle; segrnents seven and eight black, to produce a subterminal ring; hypopygium large, fulvous-yellow. Male
hypopygium with the median incision of the ninth tersite relatively
deep and narrow, the lateral lobes directed caudad and mes6d, blackened.
Dististyles very complicated in structure; dorsal submedian lobe that is
chisel-shaped in trini.d.adensisis here more acutely bispinous, the ventral
spine longer and more conspicuous; the outermost style or blade isextended laterad into a conspicuousspine.
Hlerrer:
Colombia.
Holotype, d,Lake Sapatoza Region,
Chiriguana District, August-September, 1924 (C. Allen).
Typein the British Museum (Natural History) No. 1925-576.

Tipula Linnaus.

Limonia Meigen.

Tipula colombicolasp. n.

Limonia (Limonia) catamarcanasp. n.

Belongs to the macrosterna group; nearlir allied to trinid,ad,ensisAlexander; antenne more elongate, nearly one-halfthe length of the body in male; three complete thoracic stripes;
ninth tergite of male hypopygium with a quadrate mediannotch.
Male. Length about 13 mm.; wing, 12 mm.; antennaabout 6 mm.
. Frontal prolongation of head dark brorvn dorsally, heavily pruinose,
the remainder light yellow; palpi chiefly obscure-yellow.' Antenne
more elong_ate
than jn trinid,ad,-ensis,
the scapelight yellow; first flagellar
segmentblack, the distal half-vellow;remaining segmentsblack,lheir
gltt?*g tips ligh.t yellow, the outer segmentsmoretniformly darkened.*fiead_obscureyellow, the centerof the vertex conspicuouslydark brown.
- Mesonotal praescutum yellow, with three conspicuousshiny darkbrown stripes; scutal lobes ye.119y,each variegated with two iighter
brown areas; postnotum _whitish pruinose. -Pleura heaviiy irhite
pruinose, the d-orsJ'-pleuralmembrane light yellow, dusky be"forethe
wing-rootl a blackish spot_on anepisteinum. Halteres-yellow, thebase of the knobs darker. Legs with the coxa pale, weakly pruinose;
femora yellow, the tips very narrowly but conspicuouslydari brown;

General coloration brownish gray; antenne brownish black,the first segment yellow; knobs of halteres darkened; legs
chiefly obscure yellow; wings whitish subhyaline, sparsely
variegated with brown; a supernumerary crossvein in cell Sc,
close to Scz;m-cu before the fork of. M; male hypopygium withthe rostral prolongation of the ventral dististyle very high andcompressed, the two spines placed on the side near the ventral-*corriiuotion
from the Department of Entomology, Massachusetts Agricultural College.
trhe preceding parts under this general title were published *'.
in these
ANNer.sini926 (XIX: B7&S94),and lg28 iXXf r Ozb-Olf). --- '*"*"-"768r r r4 r 5 u l .

Male. Length about 5.5 mm.; wing, 6.5 mm.
Rostrum obscure yellow, the palpi darker. Anbnne with the basal
segment yeilow, the remaindei biownish black; n.s"ff"ises-""tsoval. Head chiefly brownish gray, the front and occiput-more v"liowish.
Pronotum dark medially, paling to yellow on the sides. Mesonotal
praescutum brownish gray, darker medially, the humeral region morebrightened; sctrtal lobes dark brown, the median region and*scutellum
pale y_e-llow;_postnotum dark, pruinose. Pleura variegated brownand yellow, the surface pruinose; dorso-pleural region paier. Halterespale, the lnobs darkened. Legs with the coxe-obs&re yellow, thefore coxe_darker; trochanters yellow; remainder of legs obscure yeilow,
the-tarsal- segments.passing into dark brown. wings whitisir subhy.aline,_the oval stigma darker brown; restricted dusky cloud.s at
oJrgrn of Rs,-along cord and outer end of cell .lsl M2; veins d-ark. Venation: Sc ending sorne distance beyond origin of Rs, ,Sc, at about onethird the length of Rs, Sc2some distance fiom its tip; in both wings of770

Annals Entomological Soc,iety of Arn,r,ica[Vol. XXII'

cro-ssvein
type and thus presumed to be normal,. a supernumerarytt"*-scz; Rs long, gently arcuated; ce1l'/sl Mzlong
oni:-half its length before the^iiitlne-third't6
sternites more umtergites brown, the incjsure_s-pale;with thehvpopvgrum
Milepale'.
h6"ptsi"*f"r"rfl;;ib*;
with
each
"iti"nv
low,
lbbes
the
g"ntlyeinarginate,t'"?Sit"th"tirrtgl"'"filffi-gu.i-rt't6
and
short
lobe
ventro-mesal
thewithto ten seta.
from eisht-"V;;Lat
very
prolongation
rostralthesmal1,
relativelya;st;styte
;il.
biunt
ti*n-""a tompr"ssed,'the caudal margin Seryly rounded to the protne
or
tace
the
on
placedlaterally
short,
spinestiJ: the usualtwo-;;;i'"r
Dorsal dististvle :*rved to the*uigi".
i#;il";;;'ih"
apic.allobe unusuallv
irittt.ttt".mesal
C"";poptyr".
ffi;-ffi"lip.
microscopically
notch
the
of
marginst o?t u"a intonspicuous, lhesemrlate.
Argentina. Holotype, d, Quebrada "V-olcan"'Haerrer:(V' Weiser)'
at Agua Amarillal Catamarca, January 29, 1922
regional
other
Limon'ia catamarcana is quife distinct fromisthat offly
thespecies of the subgenus. The entire aspect ofvenationthefromnthe, than a L,imonia, but
i Dirrono*yia
th" .pecies must be referred to the latter subgenus' . It seems
very possible that the presence and positio-n of the supernumerary crossveln in cell Sc is not constant for the species'

s;
ilffi
;;j-;;";;:i-r" ".iLi
of.M.
f.ork
^---eta"*i"a1

Limonia (Limonia) villarice sp' n'General coloration dark brown; knobs of halteres darkened;legs chiefly obscure yellow; wings light browl,- th-" stigma
about.opposite two-fifths the lengthaurker; Scr,"lr""f'"trditrg
distal section of Ca; male hypopythe
Lt Rr;'m-cu longer than
the rostral spines of the ventral dististyle two'
gium'with
subequal.
*
4'6 mm';-wing, 5'3 mm'
about
LengthMaIe.
one-haif the-remainderof the head, obscureii-;J;-;p;ioximately-autf.ei
palpi
brown' Antenne with the scaqal s-egapically;v"ffo*,-brownishblack flagellum; flagellar'""'""tr'tt"*o-rTfg"
piler tha" theo"tt. Head brown, paler behind'."p-""ts"*ilf".o"ot11dark'brown, the lateral margits of the prae"ti"nv
a little more reddish brown. Plerrra similarly
lo6es
scutalattd
scut-odarkreddishbrown.-Halterespale,theknobsdarkbrown'-Legsof;ih 1h; coxe brown, th" ttoctt'rnters obscure yellow; remainderWings
infuscated.
seEgnent
tarsallasti"n.- Jr""t" yellow, only theu""*" ti"i"1 ttt" o"al stigma scir.celvdarker, bar,elvindi;%;'"il;cated: ve"insdarker brJwn. Venation: Sc1ending about oppositetwoRr, it, not far from its tip.; Rs.arcuated,toweaklv;Tfi;'th;^l;"*ah;;;l-1st Mzsmall,shorterlhan the veins beyondit;;;g"t"A;ffi;iO";

1929)

Alexand,er: ..South American Ti,ulideZZln-cu a short distance before the fork of M, longer than the distal
section of Cur.
Abdominal tergites dark brown, the sternites yeilow; hypopygium,
brownish yeilow.. Male hypopygum with the-tergite iiatrrr."rr.,the caudal margin very gently imarginate, with nirmerous coarse
sete. ventral dististylefleshy, the rosiral prolongation stout basally,
narrowed to the obtuse tip; rostral spines ti'o, pliced on basal half ofthe proiongation, subequal and_placeciclose together. Dorsal dististylea relatively short curved rod, the tip suddenly narrowed into an acutespine.
..Gonapophy_seswith the mesal apicil lobe broad., narrowed
gradually to a small, slightly decurved poini, the margin irregularly andmicroscopically roughened.
Hletrel:Paraguay.
Holotype, d, Villarica, August 26,
1924 (F. Schade).
Limonia villarice is a small, insignificant species, characterised especially by its negative characters and structure of themale hypopygium.
Limonia (Dicranomyia) weiseriana sp. n.
General coloration gray; rostrum obscure yellow; wings
whitish subhyaline, clouded with brown and gray; cell lst Mz
lorrg and relatively
narrow; male hypopygiurnwith the spinesof the rostral prolongationof the ventral dististylelong andstraight, black, the outer about two-thirds of the inner: dorsal
dististyle obtuse at apex.
Male. Length about 6-6.8 mm.; wins, 7-7.8 mm.
Female. Length about 6 mm.; wing, 8.2-8.3 mm.
Rostrum obscure yellow,_the palpi 6lack. Antenne dark brown,
the scapal seggnents.somewhat pale'; flagellar segments short-oval.
Head gray, paler behind and beneath; anterior vertex broad.
Anterior, lateral,1.pretergites .yellowish. Mesonotum chiefly gray,the praescutum with th-ree darker gray stripes, the broader -meaian
stripe more distinct; m14ian region oT scutum^and the scutellum paler,more yellowish gray.
_Pleura gray, with darker longitudinal areas on
the anepisternum and ,sternopleurite. Halteres yErow, the knobs
infuscated._ Leg-swith the coxi and trochanters yeilow, the fore coxea trifle darker; femora yellow, the.tips_narrowly and weakly infumecl;
remainder of legs passinggradt'ally into brown, the tips of the inrtividual
tarsal segments narrowly blackened. v[ings whitish subhyaline, witha handsome brown ald gray clouding; sti!:rna brown, o.rlt,
o.
less connected with a large cloud at th! end*of Rs; cord atrd oni".
-or"of cel1 lst Mt more narrowly seamed with browni extensive trip.t", "rrdgray washes on the median veins beyond the cord, as a seam at near
midlength of the basal section of. cir, and the entire distal seciion ot
the same vein; similar extensive.pale gray cloud.s on both anal veins;
veins brownish yellow, darker in the ciouderl areas. venation: .gi

772

Societyof America [Vol' XXII'
Annals Entomolog'ical

short, Scr ending before origin of Rs, Sc2a short distancefrom its tip;nii"f"tir."ty sh6rt and arciated, about one-half longer than the more.ito"iiv utd".ted basal section oJ,{.+ri cel17sl Mz rclatively long andm-cu closeto the fotk of M.
narrow;-_
etaot"i"al tergites obscure yellow,.darker medially and caudally,
the surface to*"#hut pruinosel sternites brown; hJPoPYgiumlarge,
orange-yellow. Male hyprcpyguq
dististyle co-nspicuously;il; ;"tr;i*lif, itt* i"tgit" tiansvers6, the caudal margrn *i!h 3 broad.U-shaped
emareination;. Ventral diitistyle very large and 9eshy, .the.rostralproloigation stout, beyond the spinesmore narrowed,termmaLlngln a;hid";;d point; rostral spines unusually long and conspicuous,black,nearly striight and direcied strongly b?.sad;outer-sprne about twoir.itdi ttt" t"i.Eittoi ttt" inner, both Jxteeding.inlength the prolongationi[r"fi. Do'.tui dirti.tyl" strongly curved, widened at near midlength,coti"popiryse"s"with the mesal apical lobe long andiii"^lip;;fi;.slender,gently curved.
Argentina. Holotype, d, Qaebrada "Volcan,"Henrrer:
at Agua Amarilla, Catamarca, January 29, 1922-(V' Weiser)'
Altotipotype, 9. Paratopotypes, 5 6 &, with the types'
I dedicate this interesting crane-fly to the memory of thelate Engineer Vladimir Weiser, to whom I am very greatly
indebted for many Andean Tipulida, chiefly from Jujuy,
Among the described regional
Tucuman and Citamarca.
mostclosely allied to L. @')'inaalid'aisweiseriaza
species, L.
in the more extensive(Peru),
especiallydiffering
Ale*anderofstructure of the malethedetails
and
wings
the
pattern ofhypopygium.
Lirnonia (Rhipidia) inaquipectinata sp. n.
Belongs to the maculata group; antenne (d) elongate, theflagellar segments tripectinate, one of the branches being very
reiluced, the other two long and slender; antenna bicolorous;
thoracic pleura gray with two narrow longitudinal brown
stripes; *ittg. gray with a sparse brown pattern; male hypopygium with the iostral prolongation of the ventral dististyle very
Iong and slender, with two long curved spines placed far out
toward the outer end; phallosome large and complicated instructure.
Male. Length about 5 mm-; wing, 6.6 mm'
if-bent backward
an? palpi black. Antenne (d) elon-gate,
n*t*t"
extending to sorie distance beyond the base of the abdomen; scapeenlargements oJ att segments and branches black, thea*f.;-Url"tpedicelof"the flagellar segmentswith the exception.ofthe last"i""s"t";hii-";fi'i1 flagellar segmeirtwith; singlebasal branch that is about asLe2sl

Alexander: South American Tibulid@

773

long as the pedicel; flagellar segments two to eleven with two very
long basal branches, in addition to a small intermediate branch that ii
about as long as the basal enlargement; longest branches about two
and one-half times the segment, clothed with a delicate pubescence;outer face of basal enlargements of segments with very iong, conspicuous verticils that are subequal to the segmentsl terminal fligellar
segment ,e]ongate, with verticils that are a little shorter than the segments. Head dark gray.
Pronotum dark brown. Mesonotal praescutum light brown, withindications of three dark stripes near the suture; p"osterior sclerites
of mesonotum chiefly dark-colored. Pleura gray with two narrow darkbrown longitudinal stripes. Halteres yellow, the knobs dark brown.
Llgs with the coxa yellow, their bases dark brown; trochanters lightyellow; remainder of legs broken. Wings gray, with a sparse broivn
pattern;_cel1 C infuscated; cell Sc variegated with brown; stigma oval,
grayish brown; vague brown clouds at origin of Rs, confluent with aspot at end of Sc; a large areaat fork of Rs, confluent with the stigma;
pale seams along cord and outer end of cell -lsl M2;brown seams in outerends of cells Rz and R3; prearcular region white; veins dark brown,
Cu chiefly yellow. Conspicuous macrotrichia on veins beyond cord..
Venation: Sc relatively long, extending to about opposite midlength ofRs, Sczat its tip;-Rs_ angulated at origin; cell 7sl M2-sma7l;m-cu shofily
before the fork of M.
Abdominal tergites obscure yellowish testaceous, the laterai margins
na_rrowly,_ the caudal margins more broadly, infuscated; sternites
ye1low, the incisures darkened; outer segments darkened. Male
hypopygium with_the caudal margin of the teigite only gently emarginate. Basistyle with the ventro-mesal lobe long and slendei, the apex
a littie expanded, provided with long conspicuous setae; an additional
serrresof powerful seta along the margin. Ventral dististyle large andfleshy, with sparse inconspicuous seta; rostral prolongatibn ve{r longand conspicuous, the cephalic margin fimbriite
with short dens6
yellow sete; just before the apex'with two long, slender, gentiy curved
spines that are p-l.ace4so.close together as to appear almo1t ad a single
structure; immediately distad of the rostral prolongation, on the sameface, the.dististyle is produced into a pale conicaf fleshy lobe that istipped wltlr.tyo or three elongate sete. Dorsai dististyie a very long
slender chitinized rod, strongly bent near midlength, tlie tip suddenlf
nanoyg{ into a long straight spine. Gonapophyses pale,- the mesal
apical lobe elongate. Phallosome very large and complex in structure,
inciuding a p.al_efleshy central portion that is subtended by smaller
fleshy lateral lobes.
Heerrer:
Argentina.
Ifobepe,d, Tgaaza Falls, October
3-5, 1927, (R. & E. Shannon).
Type in the United States
National Museum.
The tripectinationof the flagellar segments marks theextreme tendency, so far as at present known, in the tribe

Limoniini.

1s291TT4AnnalsEntomologicalsoc'ietyoJAmer'ica[Vol'XXII'
Limonia (Rhipidia) bruchiana sp' n'Belongstothemaculatagroup;alliedLosubcostataAlexander;
mesonotum
antenne ( d) witfr eleven long-branched segments;-blackchiefly light'brown, the 6or56lp1eural region extensively
and apex of knob con;'hal-teres black, the base of stem
extensive"nedrpl""o".ty yellow; wings whitish subhyaline' with anareas;
costalgray pattern, most eilident as a series of five
M;
of
fork
i.*itftty region extensively whitened; m-cu before
rostralfrvpoiygium with ihe tergite .small, transverse;
ventral dististyle 1ong, decurved' the two-rf"
pi.f""glii#"tspines very small and stout.
mm'Male. Length about 7 mm.; wing, 6'5 mm'; antenna'about 3'3 theremainder.of
thelength'oJt-heo""-tiru
aboui
blick,
n".t*",
consprcuous'
head.; palpi brownish black' Antenna (d) long -andmeasurements;
the
by
shown
as
the
body,
of
lengthhaft tft""""tfuieg;errrs with the stemspaleyellow,the,"*'i;ffi;;:;fiilfl"c"il"rdusky on.outer half ; antenne
segirents
tLrmlnal
the6ia'nches-b1ack,
ali flagellar sesments
2*11*l;to.mula belng;;:;;'filJ;,--'thefirst flagellar segmertofto"snip;iinate;'-branches;;";;? the last
becoming graduallv
seernents
ot iucc"eaing;a;;;;Jil;A;"i^;";h;eight) more than
tos!1
se$netits(tl-otit
flagellar
io"gJr,-irt" tongesf
branchesabout
thewith
eleventwice the segmentl fl;t"11". sEgmenteyes (d) above
gray;
dark
Head
segment.
thetfri"o""-ftJfjo"g-o
for a short distance'
contiguous'""i;:;;;;
hteral pretergites restrictedlv vellow';;e- ;";;i;tbrown, theMesonotal praescutr.il--*itr ttr" stripes 1i{ht yellowish
lobes
scutalblack;
margined'with;;;"*lyintersp;;;,mar*ins andpostnotal
testaceous
mote;d#t" Ur"*r, tii" *"?ir" area-attd"scut"liumdart tro*tt, puto caudaily,. the-cephalo-lateral a.nql,?'t*ai"t"t*it"
light.siightlv prumo-se'*o.iotiiirJ'tl]i|i;,-lhe the iui{ace in an-obliqueby^ a. consplcuous
occupiedchiefly_regiondorsal
FT;;-C;;,of the stem andbasetheblai:k,ifl"t fr"Sit"a;""f ,Up".- Hiit"t"..
with the coxaLegsligh.tvellow'r."obJ^;?;rt;;;femora"oltpi""o"t1v
obscurear*"i'basa1ly] trSclanters[iJiir]rfr-V"ff"*,-yellow;a dark brown
wit!
tips
the
beforeimmediately
at
or
;;;;[
ilc;;
ring; tibie brown; i.t i a.tt brown; posterior-tarsi broken'leygnd
rJt"t tio*"ish yelbw; it-is highly probable that;ir" f;;i1.,."i,-fr.itur."r.Lg-ents of the posteriorlegsare similarly yel1ow,;ir; ;;";;di"sii" i"i,rottolt subgroup. Wings whitish subor browniih gray pattern' somewhat"."1" "ift"t--rri"*n"ts-oF'
with an oi"*i""-gtuy
hvaline,there are five major areas' thewhere.
;c';;;i;"-;;;iii:#;
more extensivethan the inter"1"G
rti**rl th.."-"t"ut
ffi';;t"g;h'"-,t"hand outer end of ce11lst Mz;
cord
the
along
da"rk.""-t;;;;i"ilarwhite;*si;" ;;;;pi;uoi.1y' veins .brown, the..subcostal;;r;i"iltlld
about
il""
ii'
;"d
veqowfh' Venation: Sc1ending
ffi;;;;;;"eM2rclaf,ivelv
tip;
cell1st
its
at
scr
Rs,
6f
thr;" nrirrrit*ffi[:"'iengtrr

Alexand.er: South American Ti|ul,id.@

I lD

smail, shorter than vein,Mn beyond it; m-cuabout one-fourth its length
before the fotk of M.
. Abdomen brown, the incisures somewhat darker. Male hypopygium with the tergite small, transverse, the caudal margin -rieariy
straight, without sete excepttwo or three on lateral portions of sclerite.
Basistyle with the ventro-mesallobe large, strongly-narrowedat apex,
terminating in a powerful seta; mesal margin of basistyle with two
powerful fasciculate seta, the more cephalic one prolonged into adelicate hair-like point. Ventral dististyle with the rostral prolongation long, gently decurved and narrowed to the apex which-bears a
powerful terminal seta; rostral spinestwo, very short and stout, slightly
separated,placed before midlength of the prolongation.
Henrrar:Argentina, Paraguay. Holotype, d, La Plata,
Argentina, August 8, 1922, in room at light (Carlos Bruch).
Allotype, 9, Villarica, Paraguay, September,1928, (F. Schade).
Paratypes, I d, 1 g , with the allotype, September-November,
1928, (F. Schade).
I take great pleasure in naming this handsorne Rhipidia
in honor of the collector, my friend and colleague, Dr. Carlos
Bruch. It is well-distinguished from subcostata,which seems
certainly to be its nearest a1ly among the described species,bythe coloration and details of structure.
Limonia (Rhipidia) pallatanga sp. n.
Belongs to the uniseriata group; general coloration yellow,
the mesonotal praescutum with a median dark brown stripe;
antenne black, the penultimate segment yellow, flagellar
segments one to ten with a single branch; pleura with two
narrow longitudinal brownish black stripes; halteres black, theapices of the knobs yellow; wings creamy-yellow, with a conspicuous brown and gray clouded pattern; abdomen yellow,
the hypopygium concolorous; male hypopygium vrith thespines of the rostral prolongation of the ventral dististyleshort and spike-like.
Male. Length about 6.5 mm.; wing, 8 mm.
Rostrum and palpi b1ack, the former about one-half the lengthof^ the head. Antenne black, the base of the first segment a liitle
brighter, the penultimate segment light yellow; axes-of the outer
segmentsbecoming paler, almost whitish; antenne w.ith ten branchedsegments,the branchesbeingsingle,the longest (about flagellarsegment
four or-five) a little longer than the segment;branch of the first segment
appearingme.relyas a stout spur that is about as long as the glJbrous
apical pedicel of. the segment. Head brownish gray, clearer gray on
the posteriororbits.

776

Annals Entomological Society of America[Vo1' XXII'

Mesonotal praescutum brownislr.yellow laterally, . the pi$ .chiefly*"""i"a tu t"ddith brown, the median stripe passinginto d-arkbrown;
pale yettow, eachscuial lobe traversedby a dark;;r"li"il;il;dthat passes-into"th"scuteilum,convergingtoward.a centp][ror*]i""postnotumwith the median area darkened,the laterai marglns
ooint;;;i;."' Fi;;;"r1o*, ihe surflrcelargely coveredby tyo narrow longiuuct stripes. Halteies black, the base of the stem
ilai;J;;;;Lrtapi"es of the knobs obscule v;11oy,- -!-:ft*y"tn tn"
p.i"'y"f.*,'ilteboxe ve1low,morebr less darkenedbasally; trochant.ersyeuow; temora
to brown, the tibie and tarsi yeliow, the terminal;il";i;ii"ilto*ni.tr"t r"g*ents darkened. Wings with the - ground-color creamy;;i1-. tn? i*t."e laieely covered"byextensivebroryn and gray clouds'of about six bf the former, thesemore extensivei";i;;id-;"*trii"ri"-rlfrr" if.,iinterspaces; broad brownish gray seamsalong the cord, outerend of cell -/sl Mz andas marginal cloidiat ends of someof the longrjttre pat"t gray areasoccup-ymost of the remaiaing surfacei"E""f;i;il;G: ""1". l;;1:"d;g-stieaks t'ear. i6e outer ends of the mqrgrnll
r"ai.i and'medial ceiiS;- axillary region Pale; veins yellow'.darker in
the infuscated areas. tenation: Scf ending just hlyond midlength of
freetip of Scza short distancenr, St, lt. tip; Rs long,gently ar-cuated;"tftr; rn-cucloseto fotk of.M.
bevond""'Al;;;i"tt
i"tsti"t brownish yellow, the sternites.b-nsfr!er.yellow'
the basal segments narrowly_ margined laterally with. black lines;
h;;";;;r*iettowish. Male hypopygium with the tergite transverse,g""ttv emarginate,the broad lobeswith conspicuous;iG"
ilJAiA;l-a;st-ififl, with"the spines of the rostral Prolongation""1*--V""ti"fbevond mlateirgtrr, small, spike-like, close together. .Dorsal
near outer end, terminating"i.""a"ai.t"i.t'f""'g""U'-."*"fr,-ote "*pattded
abruptly in an acutesPine.
Ecuad'or. Holotype, 8, Pallatanga, altitude
Heerrlt:
4400 feet, April 1, 1922 (G. H' H' Tate)'
The only^ allied regional species is Z. (R). unipectinata(Williston) *hi"f, has all details of coloration of the wings andlegs distinct.
Limonia (Rhipidia) paraguayanasp' n'Belongs to the d'omesticagroup; general coloration brownish
yellow, tte thorax variegated with -brown; postnotal medioa velvety-black triangular area; legs yellow' thei"rgit"'with
with
outlr tarsal segments darkened; wings pale yellow, dottedofthespinesthewithhypopygium
gray;
male
brown andslender'andlongdististyleventraltheofi".t""f proloig.tion
Mal.e. Length about 5.5 mm'.; wing, 6'8 mm'
putpi black. Antenne with the -scapa! segments
no.t**"ia the--flagellum extensive1y-pale yellow, the central
Uro*rri.n black,nug"1"i s-egne"t a trifle infuscated,the incisuresrhus;;;i;;;
"u"r,

1e2el

Alexander: South Arnerican Tioulide

777

remaining broadly pale; flagellar segments (d) almost simple, little
if any produced. Head dark brownish gray; anterior vertex reduced
to a linear strip.
Mesonotal praescutum chiefly pale brownish yellow, with a broad
brown median stripe that becomes darker and better delimited on theposterior half of the sclerite; indications of narrow lateral brown stripes
before the suture; scutal lobes extensively dark brown, ringed withbrownish black; median area of scutum and the scutellum pale yellowish
gray; postnotal mediotergite with a large black triangle on anterior
haif, the caudal and lateral portions of the sclerite pale. Pleura chiefly
pale brownish yellow, narrowly lined ventrally with dark brown.
Halteres pale yellow. Legs s'ith the coxe chiefly pale, the bases
traversed by a narow dark brown line; trochanters yellow; remainderof legs yellow, only the outer tarsal segments darkened. Wings paleyellow, with an abundant brown and gray dotting in all the cells; themore conspicuous brown areas are small and restricted, distributed
as follows: Above the arculus; at supernumerary crossvein in cell
Sc; origin of Rs; tip of Sc; stigma; along cord and outer end of celllst M2; the gray dots are scattered in all cells, variable in size, somevery small, others larger and tending to become confluentl veins pale,darker in the infuscated areas. Venation: A supernumerary crossveinin cell Sci ^Scrending about opposite midlength of Rs, Scz at its tip;
m-cu before the fork of .44.
Abdominal tergites chiefly brownish, the sternites paler, moreyellowish, narrowly lined laterally with black; hypopygium yellow.
Male hypooy$um with the tergite transverse but relatively wide,
narrowed outwardly, the caudal margin very gently emarginate.
Ventral dististyle of moderate size, fleshy, the rostral prolongation
long and conspicuous, with two or three long, gently curved spines, inthe type there being three such spines on one style and only two on
the other; the spines are placed shortly before the outer end of the
prolongation, the innermost a little the shorter, the longest spine
approximately two-thirds the length of the proiongation alone.

Henrrer:
Paraguay. Holotype, &,Yillarica, July 15, 1928
(F. Schade).
Z. (R.) paraguayano is most closely allied to Z. (R.) shannoni(Alexander) of Eastern North America, which differs especiallyin the structure of the male hypopygium, the spines of therostral prolongation of the ventral dististyle being short andstout.
Limonia (R"hipidia)schadei sp. n.
group; general coloration of thorax
Belongs to the d,omest'ica
dark brown, the praescutum obscure yellow with a brown
median stripe; antennal segments (a) a little produced; wings
saturated brownish yellow, sparsely patterned with brown;

778

Annals Entomolog'icalSociety of America[Vol' XXII'

le2e)

stigma pale, encircled by a dusky ring; abdominal tergites
bieolorous.
MoIe. Length about 6-6.4 mm.; wing, 7'2-7'7 mm' .
palpi black. Antenna 61ack,the apical pedicelsof
il*irt*"tia
the-flagellar t"g-"ttt. white; flagellar. segments-:tto."F1yproduced'
so the 6ase of each segmentappearstriangular' Head blackrshgray;eyes above contiguous or virtuaily so.
biack. \4esonotai praescutumobscureyellow--p-"ot"--ttol"tirfr
scutal lobesinfuscated,the center of each
stripe;with t median brown
lobe obscure yeliow; median region of scutum whitish testaceous;dusky, pale posteriorlir; postnotum dark brown. Pleura.""i.tt"dark br6vm, spaisely pruilosg, the ventral pleurites 1bru-ptly"li"nvouf".' Halteres obscure yeliow, the knobs weqkly infuscated'" Legs
yellow, the-basesdark brown;trochanters yellow; femora
ilifr tft""""evellow, with a scarcely indicated clarker subterminai ring; tibiae andyellow, the latter passing into dark brown' Wingsi"t.i"ti"*eri"wnish yellow, the outei radial cellsa little more infuscatedl."it"i"J.lis-t pale, encircied by a dusky ring; narrow or restricted brownorigin of Rs,fork of Sc,alongcor4 and outer end of cell lst' Mzl;;;;"; prEarcularcells and axillary region dusky; veins dal! brown,
oo.t"ti"tb; s;; ;"Jt of R, Cu an6 the Anal veinsexceptat tips yellow' Vcnation:
at its tip; ce17lst Mz
just beyond midlength of
S;;;;dfit-Rs-,-Sc:r
M'
just
of
f.or\
the
beyondm-cu
small;
relatively-afa,i*i"ri tergitesbicolorous,yellow, the caudaland lateral portions
dark broivn; subterminaisegmentsmore extensivelydarkened;"".ro*iy
obscure yellow; steriites chiefly pale ye11ow' Male
hwopvst*the tergite transverse,the caudal margin very gently
withi;fi;;ilirr"
Ventral aististyte with the rostral prolongation con"ir'.rii"'rt".
rpl""8"t, the two spinesunuiually long and grominent, ?.e1tly as longtfrl ptoio"gation itself, gently curved and directed slightly basad.
"!
Paraguay. Holotype, d, Santa Barbara, SepHennar:
tember 4, lg25 (F. Schade). Paratype, d, Villarica, June 30,
1925 (F. Schade).
I take great pleasure in dedicating this crane-fly to thecollector, Vt.. f\. Schade, to whose efforts we owe most of ourknowledge of the Tipulide of Paraguay. Z' (R')- schad'e'iis
allied to" d.omestica(osten Sacken) being separated especiallyon the uniformly darkened, more strongly pectinate antennal
flagellum, the bldy-coloration, and the structure of the malehypopygium.
Linonia (Geranomyia)austroandinasp. n'
Belongs to the canad,ensis group; allied to argent'inensis(AlexandJr); male hypopygium with the caudal margin of theiergite deeply notched; rostral spines of ventral dististyle

Alexand.er: South American TiDulide779

arising from very long, conspicuous swollen bases, especiallythe inner spine, the base of which is approximately
twice aslong as that of the outer spine.

'il
I

'f
.|
I'

Male. Length (excluding rostrum) about 6 mm.; wing, 7 mm.;rostrum about 4.3 mm.
Female. Length (excluding rostrum) about 7.5 mm. I wing, 6.8 mm.;rostrum about 4 mm.
Rostrum elongate, as shown by the measurements, dark brown,
more yellowish at base. Antennre with the scapal segments brownish
yellow, the flagellum dark brown. Head gray, the posterior vertex
and occiput broadly obscure yellow.
Mesonotal praescutum gray, largely covered by four darker plumbeous-gray stripes, the humeral region more yellowish; remainder ofmesonotum dark gray, the scutellum obscure yellow, the median regionof the scutal lobes obscure brownish yellow. Pleura reddish brown,
pruinose. Halteres pa1e, the knobs infuscated. Legs with the coxa
and trochanters reddish yellow; femora obscure yellow, the tips narrowly
dark brown; tibie obscure yellow, the tips even more narrowly darkened;tarsi passing into dark brown. Wings subhyaline, the oval stigma
brown; veins brown. Venation: Scr ending opposite midlength ofRs, Scsat its tip;la weak supernlrmerary crossvein in cell Sc; Rs arcuatedto weakly angulated at origin; m-cu at or before the fork of rl1, subequal
to the distal section of Cu.
Abdominal tergites dark brown, the caudal margins of the segments
narrowly and restrictedly pale; sternites more yellow, the outer segments
darkened. Male hypopyglum with the tergite transverse, the caudal
margin with a deep V-shaped median notch, the lobes with numerous
conspicuous sete. Basistyie of moderate size; the ventro-mesal lobe
large, with a smaller fleshy lobule near base. Dorsal dististyle aslender, very strongly curved hook, grad-ually narrowed into.a long
slender tip. Ventrai dististyle very large and fleshy; rostral'spines
long and conspicuous, the outer arising from a long basal tubercle,
the inner spine arising from a very long pale cylindrical tubercle thatis about twice as long as the base of the outer spine; both spines slender,
the outer a trifle the longer. Gonapophyses with the mesal apical lobe
produced into a long narrow blade that is longer than the length of thebase of the apophysis.
Heenlr:Argentina.
Holotype, d, Cienega, near Agua
Amarilla, Catamarca, altitude 3000 meters, February 12, 1923
(V. Weiser).
Allotopotype, 9, with the type.
(G.)
L.
austroand,'ina is quite distinct from the allied andgenerally similar Z. (G.) argentinensis (Alexander),in thepeculiar modifications of the male hypopygium, as described.

780

Annals Entomological Society of America

Shannonomyia

192sl[Vo1. XXII,

Alexander"

Shannonomyia argenticeps sp. n.
General coloration pale brownish yellow; antenna with thebasal three segments light yellow, the remainder dark brown;head silvery; legs yellow; wings fulvous-ye1low, with a restrictedbrown pattern appearing as dark seams to the cross-veins anddeflections; R2 at fork of Rr+n.
Female. Length about 5.5 mm.; wing, 5.8 mm.
Rostrum and palpi dark brown. Antenne ( I ) short, the first threesegments pale ye1low, the succeeding segments abruptly dark brown;
ffagellar segments oval, graduaily decreasing in size outwardly, theverticils a little longer than the segments. Head almost covered by adense silvery pruinosity, the gene more yellowish.
Mesonotum pale brownish yellow, without dark markings, thesurface very vaguely pruinose. Pleura somewhat more testaceous
yellow. Halteres obscure yellow. Legs with the coxaeand trochanters
yellow; remainder of legs yellow, only the outer tarsal segments a little
darker. Wings with a strong fulvous-yellow tinge, the prearcular andcostal regions clearer yellow; very restricted brown seams at originof Rs, on R2, along cord and outer end of cell lsl Mz; a whitish marginal
spot at apex of cell Ra; veins yellow, darker in the infuscated areas.
Venation: ,Scpale, both Scr and Scz ending before the fork of .Rs, thelatter short, angulated at origin; Rz at fork of R3*a; veins R,: and Rr
strongly diverging, cell Ra at margin more than twice Rr; celT lst Mz
long, gently widened outwardly, longer than vein M3 beyond t\ rn-cu
close to midlength of cell 1st fuIz"
Abdomen yellowish brown. Tergal valves of ovipositor long andslender, acicular, gently upcurved.

Heenet:Ecuador. Holotype, 9, Ventura, altitude 1400
feet, April 11, 1922 (G. H. I{. Tate).
Shannonomyia argenticeps rs very distinct from the numerous
regional speciesof the genus.
Shannonomyiaparaguayensissp. n.
General coloration buffy brown, sparsely pruinose; legs an'dhalteres yellow; wings yellowish, the stigma very pale brown;veins yellow; ,Scsat extreme tip of Sc'; abdominal tergites darkbrown, the ninth segment yellow; male hypopyglum with theouter dististyle unusually short and stout.
Male. Length about 5.5 mm.;wing, 6 mm.
Rostrum and palpi black, the former pruinose. Antenne with thescapal segments brownish yeilow, the basal flagellar segmentslight

i
I

I
i
l
l
rt

Alexand,er: South American Tipulid.a78rbrown, the outer segments passing into black; flagellar segments oval,
with elongate verticils. Head light gray.
Pronotum brownish gray. Mesonotum bufiy brown, the praescutum with the median region somewhat darker, the surface sparsely
pruinosel pseudosutural fovee very small, comma-shaped. Pleurabufiy, the anepisternum darker. Halteres pale. Legs with the coxe
and trochanters yellow; remainder of legs obscure yellow. only the outertarsal segments restrictedly darkened. Wings with a strong yellow
sufiusion, the oval stigma very pale brownl veins yellow. Venation:,Sc1ending about opposite two-thirds the length of Rs, Sczat its extreme
tip; Rs strongly arcuated at origin; cell /sl Mr widened outwardlyl
m-cu aboat one-half its length beyond the fork of 11.
Abdominal tergites dark brown; basal sternites obscure yellow,
narrowly infirscated laterally; outer sternites dark brown; ninth segment
yellow; remainder of hypopygrum dark brown. Male hypopygiumwith the outer dististyle unusually short and stout, widened beyond
midlength, the apex with two blackened teeth, the outer shorter andmore slender. Inner dististyle short, gently curved.

Hesnar:Paraguay. Holotype, C, Villarica, J"o1y28,1925(F. Schade).
In its general appearance, Shannonomyia paraguayensis ismost similar to S. lentina (Alexander) and S. lentoid.es (Alexander), of Middle America. It is very different in appearance
from other regional species.
Atarba Osten Sacken.
Atarba (Atarba) breviconis sp. n.
General coloration yellow; antenne ( d) relatively short,bicolorous; legs yeliow, the terminal tarsal segments darkened;wings light yellow; Sc short; m-cu before the fork of. M; subterminal segments of abdomen ( d) extensively blackened; malehypopygium witir the median region of the tergite produced
into a conspicuous depressed rectangular lobe that divides into
two divergent horns; aedeaguslarge.
Male. Length about 5 mm.; wing, 5.5 mm.
Rostrum obscureyellow, the palpi dark brown, the basal segment
pale. Antenna (c/) unusually short for this sex, if bent backward
extending about to the base of the halteresl scapal segmentsyellow;
basal flagellar segments bicolorous, the basal portion yellow, theremainder dark brownl on the first segment, the yellow equals thebrown in amount; on succeedingsegmentsthe yellow decreasesin area,
on about the eighth flagellar segment becoming obsolete, the outer
segmentsuniformly darkenedl basal flagellar segment cylindrical, theouter segmentslong-oval. Head brownish yellow.

782

of America
Annals Entomolog'i'cal Society[Vol' XXII'

whitish pruinose'
Mesonotum yellow, the postnotum .slishtlv
Halteres pale'i*ittot"'
Pleura brownish y"uo*, the suiftc" -tp"t1at'bevondtarsalsegments
Lessvellow,the ie,,,J#'frfi;;ilt;Ftt$tt"a'-in!
veinslackingl
iir"h,y"ito*; ttig-"
ll*nt:I
thJfiist darkbrown.
than 1nshorterand iessconspicuous
ieini
oifrr"roiri"fria;;ffi."'fi,;;;';;.v*"iio,,,;'"9'rffi;dt-*ii-tl;"fl"ff ;lT:;ft#"lil"il:lfftttili:*".fthe length of Rs; cell lst Mz"i'tol,l"3llit"i,'P]H;;tt'""'t*i""{-';basal
sectionof R6 straightl'.8"i'il"rrr"":r"urths.
6qua1to one-third or
ir* aistance"?'7'
mediallv;
yeltow, the tergites more or iessd.arkened six to"""fBH*#-5ilscure
involving-se*::-t:ti"g.
a broad. brownish uriiit"t'iti"t"iti^r'hvpopvgium v:i1?Y; Maleof five;eieht and tt
produced,into att'6-ttbtn"ite"u.raii;;il;or"
hipopysium with th;'##t"'t"gio"of which is
up91
ttt9.
rtb"'
conspicuous o"pr"..i"a id;:;;"t:"";str"tmoderately
distiityleO"i"th"orns'further split into t*J ai-t3tg"nt.
large
Gonapophyses
and scattered.
*lga"t*qf
slender,the teeth *i"tl""iv-i".r.il
before midtttsi-;o"ttricted
and not especiallv ffi;;J'
basad of this'Liltil;"*,i|fi ; ;ids"liik;1"u"'-i-*Ediatelf
February 12'
Paraguay' Holotype' d' Villarica'
Heener:1926 (F. Schade)'from all of the numerous
Atarba breaicornis is d'istinct
withi" btopi"tl. America' It agreesof the #;;speciesaedeagus
theof";;;;ph;i"r'AiJ";J;
in the large size
in"'ilitg'E"t better with f'ebrigi
antennotheof
but in the coloration
which
similar spJcies' from all of
Alexander tptrtg'iuii-anJoftheantenneshort
it differs most evidently in the unusuallymale sex.

small; m-cu befote trlJi"#

Atarba (Atarba) tatei sP' n'yellow; antennal flagellum
General coloration brownish
the tips of the femora
y"ilo*'
ti""tt; t"gt
uniformly Uro*ti'f'wings grayish yellow'
6U"t"t"a;narrowly r.rA
"orr.pil;"tly
Jtshtly d.arker'thanthe ground,*"ri'Jrri*;;#the stigma
color;Rs very th";;;;;;itwo-thirdsthe basalsectionof Rr;

abdominal tergites dark
ce1l /sl M, shotl-a"d ""atly square;yiakry bicolorous; hypopygium
brown; the basal G;";#
only a few coarse spines onyellow, the.outer aitiittyf" withouter margln.
6 mm'
Male. Length aborrt 5 -mm'; wing'
wing'
mm';
5'5
Ltout
-6q q*t. brown'
Friitr.-Li"gtrtdarkerttte.
y"il"*lnaibi-AnLennae
Rostrum rlgi,, itffiilil
the secondsegmentdtttt'brown'
iheof
segm#tsbasatwith the
'""p"

1e291

Alexand,er: South American Tibulid@783

paler; flagellum brownish black, the segments gradually decreasing inlength and diameter outwardly, the terminal segment shorter and paler
than the others; verticils of all except the outer segments shorter than
the segments. Head chiefly infuscated.
Mesonotum brownish yellow, unmarked, the pleura more testaceous
yeiiow. Halteres pale, the knobs slightly infuscated. Legs with thecoxa and trochanters yellow; femora yellow, the tips narrowly but
conspicuously biackened; tibie and tarsi yellow, the outer segmentsof the latter darkened. Wings grayish yellow, the costal region clearer
yellow; stigma, small oval, only a little darker than the ground-color;veins brownish yellow. Venatibn: Sc1 ending shortly before l?s, thelatter very short, about two-thirds the length of the basal section of Ra
and in alignment with it; cell -/sl Mz unusually short and nearly square;m-cu shortly beyond the fork of M
Abdomen with the basal tergites weakly bicolorous, the remainder
dark brown, especially the subterminal segments; hypopygium yellow;
basal sternites clear pale yellow. Male hypopygium with the outerdististyle unusually slender and attenuated, the spines along the outermargin large and coarse, especially the three intermediate ones; beyond
this point, the apex is slender, blackened, bearing a lateral spine close
to tip. The sternal piate is broader than long.
Hanrrar;Bcuador.

Hololype,d,Pallatanga,

altitude4400 feet, April 1, 1922 (G. H. H. Tate). Allotopotype, ?,March 26, 7922. Paratopotype, d, with the allotype.
The speciesis named in honor of the collector. By my key
to the American species of. Atarba (Ann. Ent. Soc. America,
19:171; 1926), this species runs to brunneicornis Alexander(Colombia), a very different species that has the femora uniformly yellow and the details of venation distinct.
Gnophomyia Osten Sacken.
Gnophomyiapallidapex sp. n.
General coloration black, the lateral angles of the pronotal
scutellum yeliow; wings blackish, the outer cells extensively
paler; Rr+r+, and Rz+gsubequal; m-cubeyond midlength of thelong ce1l lst Mz; abdomen, including the hypopygium, black.
Mal,e. Lengthabout 6 mm.;wing,6 mm.
Femal.e. Length abotat7 mm.; wing, 6.2 mm.
Rostrum and palpi black. Antennaeblack throughout, relatively
elongate (d), if bent backward extending to beyond the base ot theabdomen; flagellar segmentselongate-oval. Head b1ack, the anteriorvertex a little dusted with gray.
Thorax entirely black with the exceptionof the lateral anglesof the
pronotal scutellum which are yellow. Halteres and legs entireiy black.

784

19291

XXII'
Annals Entomological'Societyof America [Vo1'n'l n*'!"u"q"S nearly""d-{:*11"
parallel'-;o{-.
n'n'*';'R'
Eorrrlto or only t littl;;;h;;ilin"tt"i;'M;';16"s;;"'"ttti
mid"eitts iith m-'u bevond.{ii,proiner.,"* e.udt;d;;;;nrather elongate'cell
theitti#til;f,;;"ili=iii"g- Male hvpopvgium with
Abd.omen,rrr",..,-"iJ' rtv?tpi'gi,,i;lit"knearly transverse' iringed withthe ternte large, its iu"alt'-"tginsl"bto"t' gentlv curved tosete. Outer dististvl""i;;'t"d':i;;;;;' very tfioJ' ttt" Iip enlarged and-whichthe obtuse apex. l"i"'- di'tittyle
virtuallyt"i*' to*" of*tft'i""g^"""tpicuousstem
theobtuse,equal the stYle in length'
Barbara' October
Paraguay' Holotype' d' Santa
Heerrer:/' MolParatype'9'
17, Lg25 (F. Schad?). Alt'oto4otypa'
L925(F' Schade)"c"l igi"' i{ ovembet 9'.inasque,l"p.rt-tii'"t'""ry. distinct from the known
Gnophomyia p;Ilida4" i'itre peculiar coloration of the
Neotropicaf ,p""i"t ot*id" g"tot inwings.
GonomYia Meigen'sp' n'
Gonomyia (Progonomyia)pleurolineata
Generalcolorati onyellowish,thedorsalpleurastriped
wings grayish yellow' thelongitud.inally with daik brown;
placed justn' very fa-int to iubobsolete'stigma barelyof' M;",,'i""tlm-iu shortlv before the fork
before the fork ;i i;-';dilateddisilstyie having themale hypopvgio* *i#" tn" oot"tblackened head trilobed'
6'3 mm'Male. Length about 5'3 mm'; wing' abotte;palpi black''Antennae
p*tloie
Rostrum pur",biack' the"'"4'lLJ**tftv"''"*; t""otia segmentwith the u..ur ."ffJ#;;;;J
thatverticils*itl"loog-olr"1'tt'"-""!*entJ"-H""?
base paler; nageffum'U1a"tt''LffJ"it'of the vertex acenterthegray'
do not exceedilt"t'"tile!:Ttif",lpretergites-r':ql::tpl15l'3h#,1i"*1,:t"t'if;vellowi"sh'*"'b''oto*qore":trJffi-$:i['i*r,13iE;t*"'ltffi-"1' ttilped longitudinallvwith
r"ot'Jir*l'i' i'ffi+ettow'
especiallytrr"
I te weaka;kle+t";t stripe;sternople"
daiker,including a t?ili d-"-t1?idarkened'stighilyttnbbiHa1t-eres.ptt"',ltt"
lv darkenedvent;l]''co*e'moie darkened;
Legswith th"..co#"b"*iitft-v"frl*'-,ttt"-tote
y.e11;' the tips somet'hat darker;
trochanters y"r,o*;*i"*ora obscureattt brbwn' Wings grayish
tibie yellowi.ft t"l*i;'it*ipi'ttrttg'l"to veins brown' Venation: Sc long'
vellow. trre strgma ili"fy "Jiagnt'f long gentlv arcuated Rs' Sczsome
Sc, ending t"vo"a"'ii?ilJgth;?tl;

South American Ti'bulid@

785

distance from its tip, Sc1 alone about one-half longer than m-cu; Rzvery faint to subobsolete, placed just before the fork of R31aicell ?nd Mz
deep; m-cu shortly before the fork of M.
Abdomen yellowish brown, the hypopygium somewhat brighter'
Male hypopygium with the outer dististyle distinctly trifid, the enlarged
blackened head being divided into two pincer-like major arms, one
flattened and obtuse, the other more curyed and terminating in a
blackened subacute point; besides these, at their bases, a shorter moretriangular branch or lobe. Inner dististyle arcuated beyond base,the distal portion smooth, the apex obtusely rounded, the surface
without punctures. Apex of aedeagus weakly trifid.

thethe cells bevond the level ofWinss with a strong blackish tinge'
andcubitalthef'ln: ""1;;;;'d;;outei end of cell trt iii,"'i''**
Venation: 'iRsshort' veryveins b-lack'anal cells conspicuously'paler;t9senttvarcuated't" Jlis{;;iAlpxander:{

Holotype, d, Quebrada "Volcan,"Argentina.
HAetrer:
at Agua Amarilla, Catamarca, January 29, 1922 (V. Weiser).
Gonomyia pleurolinealo is readily told from all allied species
by the coloration of the body and wings and the structure of
The only other regional species so far
the male hypopygium.
described with a tripartite outer dististyle is G. weiseri Alexander,an otherwise very different fly.
Gonomyia (Gonomyia) catamarca sp. n.
general colorationofAllied to G. and,icola Alexander;
mesonotum yellowish brown; rostrum yellow; antenna entirely
blackened; halteres elongate, dusky; wings grayish, the stigma
poorly defined; petiole of cell R3 elongate i rn'-cu before the forkof M;ma1e hypopygium with the outer dististyle a simple rod,phallosome with the apophyses symthe apex truncated;
metrical, slender, each bifid at apex.
lfale. Length about 5 mm.;wing, 6.4 mm.
Rostrum yellow; palpi dark brown. Antenna black, the first
segment a trifle brightened at base; antenna relatively long for amember of this genus, if bent backward extending.nearly to the wingroot. Head pale, shrunken in the type, the vertex apparently infuscated.
Anterior lateral pretergites whitish yeilow. Mesonotum light yellowish brown, without distinct markings, the scutellum a trifle moreyellow. Pleura chiefly pale yellow, with a vague dusky area extending
from the propleura onto the anepisternum; sternopleurite more reddish
yellow. Halteres elongate, dusky. Legs with the coxe and trochanters paie yellow; femora pale brown, brighter basally, the tips
darker brown; tarsi dark brown. Wings rather stronglv narrowed atbase, grayish, the stigma very pale; veins brown, Sc more yellowish.
Venation: Scr ending just beyond the origin of Rs, Sczan equal distance
before this origin; .rRsalmost in alignment with Rs, the basal sectionof the iatter obliterated or nearly so; petiole of cell rRaelongate, gently
arcuated, nearly iwice vein Rs; cell 1st Mz long-rectangular, subequal tovein Ma beyond itl m-cu about one-half its length before the fork of M.

786

Annals Entomological' Societyof America

Abdominaltergitesbrownishyellow,indistinctlylinedlongitudllllY
pale yellow; hyg-oR1rSrum*itfr [t.*","'iwh"most-evident medially; sternites
medially inlo a
prodticed
tergitefhe*thnypop;;;;;il";.
sma11,
relativelyawithnisist:itefi]""t*""ii"t" iii.t ai"ia".'ii*o"outer"t*s.
pale,
structure,
fleshylong,adististytea"rr.y rou".
;jil;;;;;i
nearlythe outer sufiaceuno'ii"* *itft t"t*. _ ittttet disliityte. a sirnple,
numerouswilhmalcrnouter
thelrr" u."r tialf more dilated,
ffiil;;"d,two p"*"tt"t flattened' ones; distal portion.of style,"t*l-i""f"ai"gitt" fu"" truncated, beforethe tip with a singlegoweririi";l.ittigf;t,the aedeagusextending.slightly
phallosome
ffi-u"J.t""aet,ful seta.
elongate'
tfte sufiendittg- ;pophyses, .the latter symmetrical'
#";;
tlp'
at
spines
acute
two
into
eaih split
Argentina. Holotype, &, Bolson' Catamarca'
Ho"r"ott(V' Weiser)'altitude 2700 meters, March 7,lg24*(G') and'icolattt" only allied .p""i"t so far madg known is G'
thecoloration'
dark
Alexander (Cofot"fil), which difiers in thevenation, and structure of the male hypopygium'
Molophilus

19291[Vo1' XXII'

Curtis.

Molophilus substYlifer sP.n.
to M'. stylifer
Belongs to the plagiatus group-; closely alliedandvenation'in ihe wing-shape'
etexanaei, difiering"s|"cia$
details of structur" of ttte maie hypopygium'**'
Male. Length, excluciinghead,about 3 mm'; wing'.q'q
brown'
lightmesonotumofC",,"tat""o1oration
Head broken.
Pleurabrown'
dark
postnotum
the,ruA"-ett"a-*itft attt"t-Uio*",darkened'weakivlmobs
theyellow,;b.;"*
J*t.^'HJt";; "b;-;;dhvellow"ft.n"";;
1"116* ; trochanters vellow ; f emora
Hi"#ffipassinginto brown'
tarii
and
"
basalv.
:;;;il;";-{n"-.i"i""conspicuous,-brown;prearcular-or",r,ro..Jr;J;#;dtt;'tiflo
ifril'*rJ'*iiirif{ingi-notand costal reglons itli"-v"iitkttt.broader-than,^i1'stllifer'concave'as in the iatter
iilJ"."a-"f *u?gir, .pp;tJ irt" q1{ veins
vein ?nd A longer'"dt-t^*"*
t;liqoe-in-potition;
Venation:
soecies.
r/t-cu'venr
ot
end
about opposite the caudal
Male-hypo"ital"gAbdomen btorn,",-ii" rrypopvgi"p somewhat-brighter''
slender,
relativelybealittpyqium with-;'''-il"r.tthe u"r;rivr"'ri"fiir[".t"r*inal
dististvle with bothouter
it1't;1n'l;:;iilillateralf""gtft l"a subacute..attips; in stytiier' thearms subequal in 'hil;;;,straight
alongBasal'distist-v1-eil;".dd.
arm is much shorter?;p"ittt, ttt" short lateral spine placed at orrod that narrows t";;;;"1"riighttv beyond one-third the length'
Argentina' Holotype' d' Bolson' Catamarca'
H,tsrrer:g, lg24 (V' Weiser)'altitude 2750 metZts, March
are unusually
The antenn* oilir" male of Molophilus stylifer
unknown'is
specieselongate; the condition in the present

Alexander: South American T'ipul'ide

787

Molophilus pallatangensis sp. n.

i'1Belongs to the plagiatus group; allied and generally similar
to M. perseus Alexander, difiering especially in the coloration
and wing-pattern.
Mal,e. Length about 3.5-3.7 mm.; uring, 4.5-4.7 mm.
Rostrum and palpi dark brown. Antenne (d) of moderate length,
if bent backward extending nearly to the root of the halteresl scapal
segments dark, the flagellum brownish yellow; flagellar segments
subcylindrical, with a short dense white pubescence. Head dark gray.
Posterior pronotum pale ye1low. Mesonotum dark brownish gray,the humeral iegion of the praescutum brighter yellow. Pleura darkgray. Halteres-pale yellow, the base of the stem and the knobs lighter
yellow. Legs with the coxe and trochanters testaceous yellow; femora
darker outwardly; tibie and tarsi darker brown. WilC-s
obscure yellow,-subhyaline,
the veins darker, conspicuousiy seamed withwhitish-toproduce a more or less streaked appearance, the morebrown clouds
evident seams belng along the cord and along the main veins basad of
the cord; veins pale yellciw, somewhat darker in the clouded areas.
Macrotrichia pale brown, the long costal fringe and conspicuous patches
along the cord and on veins Cu arrd ?nd A darker brown and more
conspicuous. Venation: Almost ai in perseus; vein 2nd,.4 extending toslightly beyond m-cu..
Abdominal tergites brownish black, the sternites more obscureyellow; hypopygium dark brown. Male hlpopygium with the beak6f the basistyle simple, the base relatively stout. Basal dististyle
generally simiiar to that of perseus, appearing as a strong rod that is
extended into a powerful apical spine, the outer margin provided with aseries of ten or more spines, the basal ones shorter and appressed. theouter ones longer and more erect.
altitude
Holotype,
Ecuador.
H,lsrrar:&, Pallatanga,
Paratopotypes,
4400 feet, March 28, 1922 (G. H. H. Tate).
2 C d, March 26-28, 1922 (G. H. H. Tate).
The thorax of the type of M. perseus was described as being
light yellowish brown but a paratype female in my collection is
It seems possible that moremuch darker, more grayish brown.
than a single species is confused in the series. M. pallatangensis
differs most evidently in the conspicuously patterned wings,
with dark veins and patches of dark macrotrichia.
Molophilus paraguayanus sp. n.
Belongs to the plagiatus group; general coloration brownish
with thegray; antenne brownish black; male hypopygium
thebasal dististyle broadly expanded on basal two-fifths,

788

Annal,sEntomological,Societyof America [Vol. XXII'

mesal edge with abundant appressed spinous sete, the apicalportion abruptly narrowed into a slender spine.
Male. Length about 3.5-3.6mm.;wing, 4.3-4.4mmFernale. Length about 4.2 mm.; wing, 4'5 mm.
Rostrum and palpi black. Antenne (d) brownish black, of moderate length, if bent bickward extending about to the wing-root. Headbrownish gray.
IVleson6tum brownish gray, the anterior lateral pretergites andrestricted humeral region tight sulphur-yellow; pseudosutural foveeelonsate.dark-colored. pteura darker brown, the dorso-pleuralregion
Itatteres yellow, the knobs weakly infumed. Legs with theoale.''coxa
testaceous yellow, the fore coxe somewhat darker; tro^chanters
yellow; femora obscure yellow, tlrg guter ends darkened.; tibie andcostalregion brighter,larsi dirk brown. Wingitinged-brown, with brown, thethe macrotrichia still darker brown'
more yellowish; veins
Venation: R, lying shortly beyond the level of.r-m; vein ?nd',4 ending'iust before the outer end of m-cu.
" Abdomen dark brown, the sternites in male weakly bicolorous,
obscure yellow with the caudal ma{grns narrowly $rk }-royq, in thefemale more uniformlv darkened; hypopygium dark' Male hypopySi"- *ittr the apical Leak of the basistyle slender,blackened. Basal
Sististyle with about the basal two-fifths greatly gxqande{, the mesal
edgewith abundant appressedspinoussete, the apicai portion abruptly
nahowed into a slendii spine. Aedeagussinuous.
Henr:rer: Paraguay. Holotyp€, d, Villarica' February 1,
1925 (F. Schade). Al'lotopotype, 9, June 70, 1925. Paratopotypes,ld, with the allotype; 1 9, with the type.
-Motophilus po,raguayanus is very distinct from all species
in the structure of the male hypopygium.