'J3Z(Rev.Zc,ol. Afc., XIV, 1.1(A paru le 15-VIII-1926.1

RECORDSAND DESCRIPTIONSOF TROPICALAFRICANCRANE-FLIES(TIPULIDAE, DIPTERA)B y CHA RLE S P . AL E X A NDE R(Massachusetts Agricultural

College, Amherst, Mass.)

PART III.
The eailier parts under this general title appearcd in the Reuue
ZoologiqueAfricaine, volume XI, in ry23. The species discussed
at this time arc all fto,m the Cameroun where they were collected by *y friend, the Rev. Jacob A. Rers. Certain of these species were included in matedal that had been sent to the Carnegie
Museum and were re-distributed to me,through the kind interest of
Mr. Huco K*ru.The types of these Speciesare indicated in the text
as being in the cullect-ion of the Carnegie Museum, the remaining
material being preserved in rny collection through the kindness of
Mr. Rnrs.In continuation of the writer's
to key the various^ttemp!.
genen in the Ethiopian fauna, a considerationof the involved genus
Trentepohliais given in this paper.
LIMONIINAE.
,'
Genus LIMONIA Mucrx.
Limonia holotricha, sp. n.
General coloratior-rdait'brbwn, the praescutal stripes palet, rcddish brown; legs dark brown; the apices of the tibiae and the tarsi

-$6obscure yellow; wings dark brown, the surface with abundant macrottichiae; Sc long; r long, arcuatedl male hypopygium with the rostralspine of the dististyle single.
Male. - Length about 8.5 mm.; wing ro.z mm.
Rostrum very short, browny palpi dark brown. Antennae darkbrown throughout; basal flagelLar segments subglobular, with shortapical pedicels; subterminal segments passing into oval, the terminal
segments elongate. Head brown, the posterior vertex more suffusedwith yellowish; anterior vertex rclatively broad, wider than the firstscapalsegment.
Pronotum dark brown. Mesonotal praescutum dark brown withthe usual sffipes paler,light reddish brownl scutal lobes reddish brown,
the remainder dark brownl scutellum and postnotum dark brown, theposterior half of the mediotergite of. the latter passing into obscureyellow. Pleura shiny dark brown, indistinctly variegated with paler
ateas. Halteres dark brown, the extreme base of the stem indistinctly
paler. Legs with the coxaeand trochantersbrownl femora dark brown,
the basesnarrowly obscure yellow; tibiae dark brown, the tips f.ading
into obscure yellow, broadest and more brightly colored on the posteior tibiae where approximately the distal half is involved; tarsi
obscure yellow, the terminal two segments infuscated; claws long,
slender, each with an acutebasal spine. Wings (Fig. r) with a strong
brown tinge, the costal and apicalregion in the ndial field more suffused; stigma lacking; veins dark brown. Abundant conspicuous macrotrichiae in all the cells of the wing except the basesof cellsRrM,
Cu, ut A and 2nd A,and the basalhalf of cell Sc. Venation ; Sc elongate, Su ending about opposite two-thirds Rs, Scz close to its tip;
Rs long, angulated and spurred at origin; r long, arcuatedl basal deflection of R4 + j very short; cell rsl Mz elongate-tectangular, nearly
as long as the longest vein issuing from it; m-cu gently sinuousrplaced shortly beyond the fork of. M,
Abdominal tergites dark brown, the basal sternites a little paleri
hypopygium dark brown. Male hypopygium (Fig. z) with the ninth
tergite short and broad, the caudal margin subtransverse to feebly
ematginate, on either side of the median arca with a group of about
15 conspicuous setae. Basistyle (b) conspicuously setiferous, theventro-mesallobe large, denselysetiferous.Ventral dististyle (vd) sub-

L

I

*16z-

Fis.z

Fig.8

E x p l i c a t i o n o f Sym b o ls: a :g:g o n a p o p h yse ; t:Fig. I.
Fig. 2. Fig. 3. Fig. 1. Fig. 5.
Fig. 6. Fig. 7. Fig. 8. -a d e a g u s; b :basi styl e; d.=. outer di sti styl e;
9 th te r g ite ; vd _ ventral di sti tvl e.

Wing of Limonia holotricha, so. n.
Male hypogygium of f,. holotricha.
Male hypogygium of L, mendica (Alexander).
Wing of Geranomgia ornatdx, sp.n.
Wing of Orimarga monilis, so. n.
Antenna of Xenolimnobiu ca^"rourens,s,g. et sp. n.
Wing of X, camerounensis.
Male hypogygium of X. camerounensis.

-168*
oval, fleshy, a littlesmaller than thebasistyle, the mesal face produced
into a long, yellow, glabrous rostrum, neat mi'dlength with a single
powerful black spine from a slightly elevated base; apex of rostrum
slightly cu,rved,the tip with two or three setae.Dorsal dististyle aslender, curved rod, gradually narrowed to the acute tip. Gonapophyses (g) broadly flattened, the apex slender, gently curved.
Hab, Cameroun.
Holotype,s', Nsi-Monden, July 3r, rg23 (I. A. Rrts).
Limonia

mendica (ArrxaNonn).

rgzr Dicranomyia mendica Amx.o,Norn; Ann. Mag. Nat. Hist.,(9) 8 : 164.
A male and a f.emale,Edea, Cameroun, May zo, rg24 (I. A. Rurs).
The male hypopygium is of very peculiar structure. The basistyles(Fig. Z, b) moderatelystout, the setaesparseand comparativelyshort;on the mesal face with scattered groups of setaeof different siZes;thelobe on the ventro-mesalface of the basistylevery long and slender,
cylindrical, tipped with elongate setae,Dorsaldististyle a small, blackened rod, the tip aclite. Ventral dististyle (vd) smaller than the basistyle, on the outer face near base with a pale clavatefleshy lobe that isof the style produced mesad into a
covered with long setae,^pexwhich is blackened and microapex
slender arm, narrowed to thescopically roughened. Gonapophyses(g) very long and powerful,blackened,the stem slender, the apexexpandedinto ahead,thelatetalangle of which is further produced into an acute point,
Genus GERANOMYIA

Hl'rtolv.

Geranomyia ornatrix, Sp. n.
GenenI coloration dark brownish gray; legs brownish black; wings
greyish subhyaline, with a handsome brown costal pattern, the maior
blotches alternating with small pa|er arcas that arc confined to the
costal and subcostal cells; Sc long; cell ut Mz lying far distad, theveins beyond it shorter thanthe celli m-cu before the fork of. M,
Female,- Length (excludingrostrum) about 4.8 mm.l wing 6 mm.;rostrum alone 2,4 mm,

-

16g -

Rostrum moderately long, brownish black throughout; palpi corr_
colorous, apparently z-segmented.Antennae brack throughout; fragellar segmentsoval. Head dark grayish brown, the very narrow anteriorvertex more silvery.
Pronotum brownish gray. Mesonotum dark brownish gray, without markings. Pleura conspicuouslyright gray pruinose, the sternopleurite and a vague more dorsal longitudinal stripe somewhat
darker. Halteres with the knobs dark brown, the base of the knobs andapicesof the stems a little paler, stems dark brown, the baseobscureyellow. Legs with the coxae and trochanters light brown;
femorabrownish black, paler at extreme base; tibiae and tarsi brownish black.
lfings (Fig. +) grayish subhyaline, with a handsome brown patternthat is in part slightly ocelliform; a prearcurar brown spot; six irregrlarly rcctangular brown areasalong the costal margin, the first immediately beyond h, the third at the origin of Rs, rhe second at middistance between these twol the above thtee areasextend from costa
to media; a larger and more htegulat blotch at end.of Scr, extending
from costa acrossthe fork of Rs into cell R; blotches two and three
have vaguely pale centers that.arc confined to cell R; arternatingwith
the four majot markings described are paler grayish brown clouds
that occupy cells C and Sc only; beyond the cord the brown paftern
is more extensive and furegularya conspicuousstigmal areaat the endof Rr crossescell R3 near its base and follows along the cord as analrow seami a blotch at end of vein Rzl3 is barely connected witha slightly larget spot in cell R5 near mid-length and,along theourer
end of cell rst Mz; a subapical blotch, heavier in cell R3, becoming
paler in cells R5 and znd Mz; veins dark brown, the distal portionofcosta more incrassatedand colored yellow and dark brown, alternately.Yenation:sc long,scr extending to shortly before the fork of
Rs, Scz at its tip; Rs elongate, arcuatedl cell rst Mz very large,longer than any of the veins issuing from it ; m-cu about one-half its
length before the fork of M, subequal to the distal section of cur;cell znd A nattow.
Abdomen black, the basal segments a little variegated.with paler.
ovipositor with the bases of the valves brackened, the valves themselves testaceoushorn-colorl sternal valves short and stout; tergal
valves slender, the tips obtuse.

- t 70-

Hab, Cameroun.
Holotype,9, Mo Tyan, August 14, rg23 (I. A, Rus).
The only other Ethiopian Geranomyiawith a wing-pattern of thenature of G, ornatrix is G. sex-ocellataArnxanorn of Cape Colony.
In the latter species,the markings are much larger, with pale centersthat encroach upon cells sc and c; the alternate paler blotches arenearly obsolete, being represented only by vagxe clouds in cell Sc
between the first and second and the second and third major areasi
dark spots present at the ends of the Anal veins.
Genus ORIMARGA
Orimarga

Osrrn S^ncxrx.

monilis, sp. n.

Genenl coloration reddish brownl antennae moniliform; wings
subhyaline, the stigma lacking; Sc long; Rs srongly angulated atorigin; r near tip of Rr, lying slightly f.arther distad than r-m1m-cu
opposite origin of Rs.
Female. - Leqgth about 4 mm.; wing 4.7 mm,
Rostrum reddish brown, the palpi a little darker. Antennae pale
brown throughout, moniliform, the segments subglobular, the terminal segment a ttifle larger than the penultimate. Head dark brown.
Thorax reddish brown, discolored in the type. Halteres broken.
Legs with the coxae and trochanters reddish brownl femora reddish
brown, the rcmainder of the legs paler. lfings (Fig. l) subhyaline,
the stigma obsolete;veins pale brown. Venation I Scr qnding shortly
before the fork of Rs, Scz a short distance from its tip; Rs very strongly
angulated at odgin; r about its own length from the tip of Rr and on
Rz -l Z at neat one-fifth th elength;terminal section of Rz ].3bent gradually toward the wing-tip; basal deflection of R4* j strongly angulated; r-m lying immediately proximad of the level of r; petiole ofcell M3 a tifle shorter than the cell; m-cu opposite the origin of Rs.
Abdomen reddish brown, the pleural region narrowly darker.
Ovipositor with the valves short and stout, the tergal valves gently
upcurved to the acute tips.
Hab. Cameroun.
Holotype, Q, Nsin Monden, July 3r, rg4 U. A. Rrrs).

- T7T-orimarga monilis is the first species of the genus to be describedfrom the African continent. It differs from the seychelles o. scotti
Edwards and Aldabran o. fryeri Edwards in the venation, as the rclative positions of cross-veins r and r-m, the length of the basal deflection of R+ * I and the length of cell M3 in relation to its petiole.
XENOLIMNOBIA,

g. n.

Antennae (Fig. 6) with the scapal segments enlarged.,especiallythe second; flagellum abruptly smaller than the scape; basal four flagellar segmentsshort and crorn'ded,the first largest,narrowed at base,these basal flagelLarsegmentsdensely covered with coarsewhite setae;
beyond the fourth the flagellar segments pass into elongate-cylindrical and are provided with very long verticils in addition to theshorter and more sparsepale setae.Thorax without tuberculate pits
or pseudosutural foveae. Middle and posterior coxae approximated,the meron small. Tibiae unspurredl tarsal claws rong, slender. lfings(Fig. Z) broad; Sc of moderatelength, Scr extending to about oppo_
site the fork of Rs, sca relatively indistinct, removed to a short$istance from the tip of scr, thelatter alonea rittre shorter than m-cu|
Rs relatively short and straight, shorrer than Rz lZ; Rz*3 long,straight, much longer than R3 alone; what appearsto be the compos_
ite r and baseof Rz is a long, gently sinuous vein placed almost at the
tip of Rr, the caudalend of the vein lying slightry f.artherdistad thanthe cephalic endl inner ends of cells R3, R5 and rct Mz in ahgnment;
cell Mr lacking; cell rs/ Mz open by the atrophy of m; m-cu neat
mid-length of vein Mz i +; cell M3 wide, more rhan rwice the lengthof its petiole; the semiatrophied Cuz distinct to opposite m-cu; Analveins long, vein znd A neady straight to near its outer end. Trichiation : Macrotrichiae of cells rclatively largel n:acrotrichiae of veins
occurring on all the longitudinal veins beyond the cord and on thefollowing veins basad of the cord : on sc for its whole rength; distal
fifth of Rs; distal two-thirds of. M; extreme end of basal section of
Cutl enthe length of rst .4; sub-distal third of vein znd A; on thesinuous oblique composite vein r plus base of Rz are four rarge trichiae, distributed over the whole length. Male hypopygium (Fig. g)with the ninth tetgite (t) deeply notched medially, the lateral lobes

-r72thus formed conspicuous.Basistyles(b) stout, with conspicuoussetae,
some of those on outer face of unusual length. Outer dististyle (d)a rclatively slender, nearly straight blade, the apex produced into aslightly curved spine, before the apex on inner margin with a few
microscopic spinulae; inner margin of style with a few setael innerdististyle broad-based,narrowed to the slender, cylindrical apex, thetip obtuse. Gonapophyses(g) appeaing as flattened,btackened plates,
the outer margin produced laterad into a conspicuous spine, the apex
with about three appressedspinulae.
Genotype. - Xenolimnobiacamerounensis,
sp. n. (Western Ethiopian Region).
The tiny fly that has been describedabovestill further complicatesthe aheady very involved classification of the Tipulidae. It shouldbe placedin the Hexatomoid Eriopterini but the condition of the radial
field of the wing is highly peculiar. The long, sinuous vein that appears to be the radial cross-vein is almost cefiainly-a composite vein,the cephalic portion being r, the posterior portion the basal sectionof Rz, the distal section of the latter vein being entirely obliteratedn
A somewhat comparablecondition is found in the subgeneraPlesiomongomaBrunetti and AnchimongomaBrunetti of the genus Trentepohlia. In Xenolimnobia, if the branch Ra was complete, cell Rzwould necessarilybe very tiny, as in Rhabdomastixand Gonomyia,
Xenolimnobia camerounensis, sp. n.
General coloration dark brown; wings broad.
Male, - Length about 2,5 mm.; wing 2,7 mm.
Rostrum and palpi dark brown, Antennae dark brown throughout,
its structure as describedunder the genus. Head dark brown.
Mesonoturn and pleura uniformly dark brown. Halteres short, darkbrown, the baseof the stem a very little paler. Legs with the coxaeandffochantersbrownish yellow; femora brownish yellow, the tips narrowly darkened, the segments with suberect setae; tibiae longer than thefemora, with relatively long setae, pale brown throughouti tarci pale
brown, the basitarsi exceeding in length the combined rcmaining
segments.Wings short and broad, strongly tinged with browni stig-

l--

I

-r73ma tackingl veins darker brown. Venation as discussed under the.generic chatacteization.
Abdomen dark brown, including the hypopygium, the structure
ofthe lattet as discussed under the genus.
Hab. Camercun.
Holotype,6:rEdea, September :^3,rgzS (|. A.Rrrs).

sg' B,
IstA iu,

sorR, l,

&
/'&*'

il,*,

R.
iRo*,
[|n

t+2

F is. l l.
Figures to illustrate venation of subgenera of Ttentepohlia.
- Explanation of venational
symbols : A :Anal veins; Cu :Cubitus; M --- Media; R:
Radius: Sc :Subcosta.
Fig. 9. Fig. 10.
Fig. ll.
-

Venation of Paramo4goma (nigeriensis Alex.).
Venation of Mongoma (reisi Alex.).
Venation of Trentepohlia (hgalina Alex.).

Genus TRENTEPOHLIA

Brcor.

The tropicopolitan genus Trentepoftlia is one of the most common
and omnipresent genera throughout tropical Africa. It has been sub-

-r74

-

divided into six stbgenera. - Neomongoma AtnxtNDsR, Paramon'goma BnuNnrrt, Mongoma Wnsrwooo, PlesiomongomcBnuNETTr,
TrentepohliaBrcor and AnchimongomaBnuNrrrr. These occur inthe tropics of all the geographicaldivisions of the World, the Oriental
being richest with all the groups present except Neomongoma.The
Ethiopian Region has three (Paramongoma,Mongoma and Trent'
epohlia),the Australasiantwo (Mongoma and Trentepohlia),the Neotropical two (Neomongoma and Paramongoma, the latter especiallycharacteristic).The subgenus Trentepohliahas invaded the southern
limits of the PalaearcticRegion (7. efflatounePrrnns in Egypt, T. septentrionalis Ar,nxenorn in Japan), Of the six groups, Neomongomais
the most genenllized, Paramongoma next, which would indicate an
American origin for the group despite its present very notable development in the tropics of the Old World.
The three African subgenera of. Trentepohlia all show very distinct
is the most genetvenationalpeculiarities.Of thesethree,Paramongoma
allized in the wide separationof veins Car and st A at the wingmargin.In the condition of the medial fieldrhoweverrMongoma(Figure ro) is more primitive inthat it has retained the full complementof three branches.The reduction of the medial field in Trentepohlia,
s. s. (Figure r r) is seento result from the atrophy of cross-veinm withthe f.wther elimination of both sectionsof vein M3,In Paramongoma(Figure g) the posterior elementthat closescell rsl Mz wotld seemto
be a compositevein, the anterior or cephalic portion rcpresentingm,
the posterior portion of the samevein representingthe first section ofvein M3, the outer section being entrely eliminated. In interprctrng
the medial field in this manner,one must keep in mind an alternative
view which may be correct,that is the fusion of veins R4 5 andMr -t z'
to the wing-margin, a view that is suggested by the condition in
Neomongoma.If this latter view is correct, then of the three veins issuing from cell rsl Mz in Paramongoma,the upper or first would be
the fused R4- j plus Mt lz, the second M3, the third M4.
Generalcharactersof Trentepohlia:
The legs are alwayslong and slender. The armature to be found at
the bases of the femora in the various species of. Trentepohliaand
Mongoma consists of a linear series of short, peg-like spines, appar-

-r75ently most persistent on the fore femora but in some species
occuring on all the legs. correlated with this charact", is arcmarkableotnamentation of the posterior trbiae, where, shortly before theirtips, are found a linear seriesof long, curved bristles, varyingin
number from t to 4 in different species.These bristles occur in bothsexesbut, in cefiain speciesat least,may differ in number in the two sexes.
There can be no question of the taxonomic value of these bristles
inthis involved genus but more data rcgarding the normal and sexual
variation that occurs must be forthcoming before this can be safelv
used as a ptimaty chatactet.
The following tendenciesof the venation are well-marked : sc very
long; scr and Rr approximated atthe margin;cell Re rarge andsprawly, the anterior branch, Rz, atrophied in the subgeneraplesiomongoma
and Anchimongoma;vejn R4 r j extensivelyfused with vein Mtlz,
eliminating r-m and appearingto issue from the outer cephalic endof cell st Mz; cell rsl Mz closed,with four veins issuing from it in
Neomongoma,Mongoma and plesiomongomq.
(R4 , j, .i54y+2t M3 and,
M4), closed, with three veins issuing from it in paramongoma, oropen in Trentepohliaand Anchimongomaivein cur with its ourer section always deflected strongly caudad, forming a wide angle withm-cu, in the more speciallized groups fused back from the margin
with vein ut A to c.mpletely close cell cu; this cell widely open in
Neomongoma,less so in Paramongoma,very narrowly open in Anchi_mongomq,closed in the other groups, most extensivelyso in the gracilis group of. Trentf,pohlia; vein znd A short ro very shorr.
The crane-fliesof this genusin the Ethiopian Region are as follows :SubgenusParamongoma :f , @,) mera, sp. n,'f . (P.) nigeriensisArsxA,Nonn;
Ann. Mag. Nat. Hist., (9) 6 , 43;
r92o.
T, (P.) ramisiana (Rrrorl); Voy. Anuen et
JranNEL en Afrique
Orientale, Ins. Dipt,r 3, Nemat. polyneura, pp. g5-g6, fig. rr (Mon_goma); rgr4.
Subgenus Mongoma :T. (M,) albilata Arnx.o,NosniAnn. Mag. Nat. Hist., (9) I
5 56_5g;r92o.

-176f . (M) albilatissimaAr,rxaNorn; ibid., (g) 6 : 4zi rgzo.
f . W.) dummeriArnxnworn; ibid., (g) 8 : t65-r66; ryzt.
f . (M.) fragillima (Wrsrwooo); Trans. Ent. Soc. London, r88r :Z6+-26j, pl, q, fig. t (Mongoma); r88t.
f . @.) madagascanensrs
Arrxlnorny Ann. Mag. Nat. Hist., (9) 6;
4o-4ri r92o,
f . W) metatarsatraAr,rxlwpen; ibid., (g) 6 r 4r-4zi rgzo; ibid.,(g)8:jrz;rg2r.
f . @.) niveipesAr,sxA,Norn;ibid,, (d Z :3rg3zoi rgzr,
f . W.) pennipes(Osrnw Slcrnn); Berl. Ent. Zeitschr,, 3r ,2o4
(Mongoma)i 1887,
f . @,) re6r ArrxaNonnl Ann. Mag. Nat. Hist., (9) 5:58-59; tgzo.
Subgenus Trentepohlia :T. Q,) alluaudi Aruxmtoen; Bull. Mus. Hist. Nat., Paris, rg2o ,2r9, r92o.
f , Q ) aurantiaAtrxnworn; Ann. Mag. Nat. Hist., (9)5 i 47ri rgzc.
T. (f ) breuisectorAr,rxlt.Iorn; Rev. Zool. Africaine, XI, fasc.4:
374-375t1923.
T. g.) curtipennis(Snrrsrn); Berl. Ent. Zeitschr., 5z : 175-136,(Mongoma); ryo8,
f . Q) exornata Bencnonr; Ent. Tidskrift, 1888 : ry5-ry7, fig.
3; 1888.
T. V,) fuscoapicalisAr.sxlNorn; Ann. Mag. Nat. Hist. (9) 5 :348; ryzo.
T. Q) gracilisENornrnrn; Zool, Jahrb., Syst., 3z t 6r-62, fig. Mr;
T.gr2.
T. g.) g, continentalfuAlsxnNorn; Ann. So. Afr. Mus. 18 ,2o22o3i rg2r.
f . (f .) humeralisArBxlnronn; ibid., 18 : zor-aozi rg2r.
f . Q) hyalina Arnxlxorn; Ann. Mag. Nat. Hist., (9) 8 : r68-i69;
r92r.
f , Q) inflata Ar,exaNoBn;ibid., (g) 6 : 43-44, rg2o.
f . Q.) nigricolorAr.rxanprn; ibid., (g) 8 : 166-167; ryzr.
f . Q) nox Alsxaxorn; ibid., (g) 8 : r67-t68i tgzr.
f . Q) pallidipleura, sp. n.
T. (7.) pomeroyi ArBxlnonn; Ann. Mag. Nat. Hist., (9) 8 : 169rToi r92r..

-r77T, (7.) reuercalis,
sp. n.
T. Q) speiseriEow.o,nosl
Ann. Mag. Nat. Hist., (8) rz:zo4,fig.;
r913.
f , Q) ugandaeAr.rxa,xornl ibid., (g) 6 ; 96337i rgzo,
f , Q.) zambesiae(Arrxlworn); C,an.Ent., 44 : 86-88 (Mongoma)i
rgr.2.
Besides the above 3o speciesfrom the Ethiopian Region, T. e.)efflatouni Prsnnr (Bull. Soc. Roy. Ent. Egypte, rgz2 . 83-85, figs.;ry23) has recently been described from Egypt. It is very closely allied
to r. (r.) zambesiae,differing in slight details of. size and coloration.
The lawae were found invegetable ddbris in earth by M. Errraroulr.
The speciesis not included in the following key sinceit is extra-Ethiopian.
A KEY TO THE AFRICAN SPECIESOF THE GENUS
TRENTEPOHLIA BIGOT.
r. Three branchesof M rcach the wing-margin (SubgenusMongoma
Wrsrwooo; text-figure ro).
z
Two branchesof.M reachthe wing-margin.
roa. Wings heavilyspottedwith brown. (Madagascar).
madagascariensr
Arrx.
Wings unspotted(exceptfor the stigmalspot, when present).
jBasitarsi
proximalwithends
black;
wings
andtheir veins yellowish.
3.
(Nigeria-East Africa).
metatarsatra Atnx,
Basitarsi white, concolorous with the remainder of tarsi.
4
4. Femon brown throughout; mid-tibiae slightly dilated at apex and here
fringed with conspicuous white setae (Seychelles Is.- Oriental
Region).
pennipes(O. S.).
Tips of the femora more or less whitened; mid-tibiae not ornamented
as above.
5
5. Femoral tips narrowly and indistinctly pale; size small (wing underniveipes Atux.
7 rnm.) (Cameroun).
Femoral tips abruptly and conspicuously white; size larger (wing over6
7 and usually over 8 mm.).
6. Tibiae dark brown, only the narrow tips white. (Cameroun)..
.
relsr ALsx.
Tibiae largely white; a brown band beyond the base followed by a broad
white apex that includes approximately one-third or more of thesegment.

-t7 8 -7. General coloration piceous-black; (tibial brown ring occupying onehalf the segment) (West Africa).
fragillima (Wnsrw.)
General coloration pale brown to yellowish brown.
8a Pale tibial apices occupying a little less than one-third the length of thesegment. (Uganda).
dummeri Atnx.
Pale tibial apicesoccupying the terminal half or more of the segment.99. Antennal scapelargely dark brown; tibiae with the brown ring narrower
than the pale base; pale tibial apices occupying about the distal
two-thirds of the entire segment, (Gold Coast). albilatissimaArrx.
Antennal scapeyellow; tibrae with the brown ring more extensive,being
about twice the pale base; pale tibial apices occupying about the
distal half of the entire segment.(Cameroun).
albilata Arnx.
ro. Veins Cu and rct A entfuelyseparateat wing-matgin; cell ut Mzclosed (SubgenusParamongomaBrunetti;text-figure 9).
rr
Veins Car and rsl ,4 fused for a varying distance back from the wingmatgin; cell rsl Mz open (Subgenus Trentepohlia Bigot; textfi g u re rr).
13
rr. General coloration light yellow; (femora pale brown, the tips broadly
white). (Cameroun).
mera.,sp. n.
General coloration brownl (femora pale brown in nigeriensis,or coloration unknown, in ramisiana),
I2rz, General coloration brownish; plewa grayish; wings weakly tinged
with yellowish. (British E. Africa, coastal).
ramisiana(Rrro).
Genercl coloration uniform pale brown, the pleuta more yellowish;
nigeriensisAt-xx,
r3. Wings with the costal region before mid-length of wing conspicuously
expanded. (Nigeria).
inllata Arcx.
\)flingswith the costalregion normal, straight.
r4
14. Wings subhyaline; fusion of veins Cu and tst A ex tensive, longer
than the distal section of Cur alone.
r5
Wings subhyaline or (usually) variegated with brown; fusion of veins
Cul,and rsl .Aslight, lessthan the distal section of.Clul'alone,17
15. Abdomen of normal length (about j mm);wings broad. (Portuguese
East Africa).
zambesiae(Arnx) .
Abdomen very long and slender (over 7 mm.); wings unusually narrow,
16. lfings broader (J, about:. B by r"r mm,). (Madagascar)
grccrlls Eno.
Wings narrower (ol, about 6, I by r.o5 mm.). (Transvaal).
gracilis continentalis Atnx.

-r7917. Wings hyaline or approximately so, unmarked except for na*ow
vague
seamsalong certainof the longitudinal veins, al the cells clear. rg
lfings with some of the cells suffusedwith brown or yellow.
rg
18. Wings quite hyaline; Rs elongate, fully twice the basal deflection
of
Rq+S; second section of vein R2+3 shorter than r; cell znd A
narrow, (Cameroun).
hyalina Atrx.
wings with a vague dark clouding along vein cu and, ress evidently
along vein R++S; Rs short, about equal to the basal deflection of
R++S; second section of vein Rz*3long, only a little shorter
than the first section and longer than Rs; cell znd A broad..
(Dahomey).
breuisector Atsx.
19. wings hyaline, cell R3 and the stigmal region yellowish. (cameroun).

curtipennis(Snrrs.).
Wings with distinct brownish markings.

20

eo. rving-markings virtually confined to the broad brown apex; (femoral
tip paling into white). (Cameroun).
fuscoapiialis'A:lix,
wings with a broad seam along the cord in addition to the apicar area,
2T

zr. Tips of the femora abruptly black or dark brown.
22
Femora beyond base either uniformly colored or the tips paling into
whitish.
2+
zz. Tips of the tibiae darkened; (wing-pattern very pale), (Madagascar).
alluaud.i Arxx.
Tips of the tibiae not darkened.

23
23. Thorux, including the pleura, shiny brownish black; Rs long, fully
four times the length of the second section of Rz_l3i wing-pattern comparatively heavy, cerl Ra being uniformry darkened.
(Cameroun).
nox Arxx.
Thoracic dorsum brown, the humeral region and pleura testaceousyellow; Rs relatively short, less than twice the length of the second
section of Rz*3iwing-patterncomparatively light, the centerof cell Rz extensively paler. (Cameroun). pallidipteura, Sp. n.
24, Tips of the femora narrowly and abruptly whitened.
Femora beyond the base uniformly colored.

25z6

25. Abdominal sternites yellowish, the apical third of the individual segments dark brown. (Nigeria).
pomeroyi Atnx.
Abdominal sternires dark brown on basal half or less, the distal portion of the individual segmenrs obscure yellow. (Cameroun).
reversalis,sp, n.

-

r8o

e6. Colorationof the body shiny black,the humer:,lregionof praescutum
not brightened.(Cameroun)
nigricolorAt-nx.
Colorationof the body varying from yellow to dark brown, in the lattercase with the humeral region of praescutum further bigtened.
27
27. Sizevery large (9, wing about 8.5 mm.); (mesonotalpraescutumteddish brown with three darketbrown sripes; posteriortibiaewith
ugandaeAtnx.
4long bristlesat tip). (Uganda).
Sizesmaller(9, wing under 7 mm,),28
e8. Brown markingat cord with a subhyalinecenter.(South-east
and East
Africa).
exornataBnncn.
Brown matkingat cord solid or virtually so.
29
e9. Mesonotalpraescutumlight orange-yellowwith a delicatereddish
brown medianline (Reunion).
aurantiaAtnx,
Mesonotalpraescutumnot coloredas above.
30(Cameroun,
Thoracic
dorsumreddishyellow.
brown;
legs
light
East
3o.
Africa-OrientalRegion).
speiseriEow.
Thoracic dorsumbrownishblack, the humeralregionsof the praescutum reddish;legsdark brown.(Transvaal).
humeralisArnx.
Of the speciesincluded in the above key, the writer has seen thetypes or authentic specimensof all except the following : T. fragillima(Wnsrw.), 7". ramisiana(Rtno.) and T.curtipennis(Snrrs.).Their position in the key is basedon the published descriptions alone,except in
the caseof fragillima where unpublished notes on the type arc given,
submitted to me through the kindnessof Mr. Eowlnos.
Trentepohlia(Paramongoma)rrtera, sp. n.

Genenl coloration light yellowl femora pale brownish testaceous,
the tips white; tibi.ae and tarsi white; wings uniformly pale yellow, theveins a little darker.
Female, - Length about J,z mm.; wing 4.j mm,
Rostrum and palpi brownish yellow. Antennae relatively short,brown, theflagellar segmentsoval. Head gtayrmote infuscatedmedially; vertex very narrow.
Pronotum obscure yellow. Mesonotal praescutum obscure yellowish fulvous, without markings, the pleura clearcr yellow. Halteres
pale brownish yellow. Legs with the coxae and trochanters yellow;
f.emorabrownish testaceous,becoming a little darker toward the tips,

-r8rthe latter suddenly snowy white (about o.6 mm. in length); tibiae and,tarsi snowy white, only the terminal segmentsof the latter infuscated;no outstanding spines or setaeon legs. wings uniformly pale yelrow,the veins a little darker than the ground-color. venation : very similar to T, (P.).nigeriensr(shown tntext-figwe); m-cu more than one
half its length before the fork of.M; distancebetween tips of veins car
and rs/ A along the wing-margin greatet than the outer section ofvein Cut alone.
Abdomen brownish yellow, the genitar segment clearer yellow.
ovipositor with the valves long and slender, especially the gently
upcurved tergal valves.
Hab. Cameroun.
Holotype, Q, Edea, March 26, rgzz (J. A. Rrrs),
Type in the collection of the Carnegie Museum.
Trentepohlia (Trentepohlia) pallidipleura, sp. n.
Allied to T, (7.) nox, At-nxANDERi
mesonotum dark brown, theanterior and latetal portions of the praescutum andthepleura yellowish testaceous;legs with the femora yellow, narrowly tipped withdark brown; posterior tibiae bef.oreapex with two long, slender setaej
wing-pattern pale; Rs less than twice the length of the second section of Rz+3.
Female. - Length 6 mm; wing 5 mm.
Rostrum and palpi brown. Antennae brown, the terminal segmens
a little paler. Head light brown, the vertex narrow.
Pronotum yellow. Mesonotal praescutum dark brown, the broad"
humenl region and na'ower lateral margins right yellow, the palecoloration including the anterior margin of the sclerite; remaining
sclerites of mesonotum dark brown, except the postnotal pleurotergite which is concolorouswith the testaceousyellow pleura. Halteres{ark brown, the base of the stem light yellow. Legs with the coxae
and trochanterspale testaceousyellow; femora yellow, the tips narrowly and abruptly dark brown; tibiae and tarsi obscure whitish, the tetminal segmentsof the latter a trifle darkened; fore femora with a basal
seriesof about five short spinousbristlesthat becomelargeroutwardly;

-r8zposterior femora with a more diffuse group of. neafly a dozensmaller
subequal spines; posterior tibiae with two long, srendersetae before
apex, these separetedfrom one another by a distance about one-halflonger than the length of one, the outer bristle about two and onehalf times its length from the apex of the tibia. wings whitish subhyaline, the stigma oval, brown; diffuse pale brown croudsalong veincur, at origin of Rs, along the cord and the wing-apex, the center ofcell Rz rcmaining palei the genenl effect produced is of.a pale brownpatterni veins c, sc, R and the second section of Rz |3 yellow, thercmaining veins dark brown. venation : Rs arcuated; basal sectionof Rzl 3 shorter than the second sectionl basal deflection of R+
rsabout equal to the petiole of cell R5 and a rittle morc than one-halfthe second section of. Rz+3,
Abdominal tergites dark brown, the sternites conspicuously lightyellow, the genital segment yellow; valves of ovipositor dark horncolor.
Hab, Camercun.
Holotype, e,Edea, July 28, rgz? (J, A. Ruls).
Trentepohlia(Trentepohlia)

reversalis, sp. n.

Allied to T. (7.) pomeroyiArrx^o,r,mnni
mesonotum dark brown, thehumenl region of the praescutum extensivelyobscure yellow; femora yellowish brown, the tips narrowly but conspicuously whitened;wings subhyaline, the pattern distinct but restricted; Rs one-halflonger than the basal section of Rz i3; abdominalsternites bicolorous, the basal half or less of each segment brown, the apical portion
brownish yellow.
MaIe. -

Length 8 mm.; wing 7.5 mm.

Female. - Length B-9 mm.; wing 7-7,5 mm.
Rostrum and palpi dark brown. Antennae slender, dark brown.
Head brown.
Pronotum dark brown. Mesonotum dark brown, the humenl regionof the praescutum extensivelyobscure yellow. pleura obscure yellow,
the mesopleura more infuscated; pleurotergite with a row of threelong setae. Halteres dark brown, the extreme base obscure yellow,
Legs with the coxae and trochanters yellowish testaceousl femora

-r83yellowish brown, the tips narrowry (on fore legs) to more broadly (onhind legs) obscure whitish; tibiae brownish white, the distal portion
broadly whitened; terminal tarsal segments a trifle darkenedl basesof all the femora with a longitudinal series of short, spinous setaei
posterior tibiae with a row of three or four long, black, spinous
setae(a) or with two such set,e (Q). wings subhyaline, with a restricted
but distinct brown pattern, this including clouds in the basal halfofcell M; along vein Cui inbases of cells Ca and.st A; alongRs andthe basal section of Rz*3 to the stigma; alongthe cord to the posterior margin; alongthe petiole of cell R5 and vein R4+ j to thewing_apex exteflsively infusc ateid,the center of cell Rz uniformly dark, inrare instances with the central portion pale? stigma oval, darker
brown than the other brown markingsyveins dark brown. venation :Rs elongate, one-half longer than the basal section of Rz 13 andabout two and one-half.times the length of the second sectionypetioleof cell R5 longer thanthe secondsection of. Rz*3but only about twothirds the first section.
Abdominal tergites dark brown, the hypopygium concolorousl
sternites bicolorous, the basal half or less of each segmentbrown, theremaining apical portion brownish yellow.
Hab. Cameroun.
Holotype,qf, Son Ngen, July 3o, rg4 U, A. Rus).
Allotopotype, Q.
Paratypes,3 QQ,Edea, June zz, rgzz, Octoberg, rgz4 (J. A. Rus).
Genus CLYDONODOZUS
Clydonodozus fulvithorax,

ENornrrrr.

sp. n.

General coloration fulvous, the mesonotum unstriped; femora yellow, the tips narrowly dark brownl wings broad, yellowish, sparsely
vafiegated with brown.
MaIe. -

Length r4.j mm; wing ro.5 mm.
Rostrum light fulvous brown, the reduced palpi brownish black.
Antennae with the elongate basal segment light brown beneath, theupper surface almost black; second segment dark brown, paler api_

-r84cally; flagellum black, the verticils elongate. Head dark brown, pollinose, especially anteriorlY.
pronotum light brown, darker medially. Mesonotum uniformly
fulvous, unmarkedl pseudosututal foveaesemicircular, black. Pleura
brownish yellow, the pteroplewite more infuscated. Halteres pale
brown. Legs with the coxae infuscated externally; trochanters obscurebrownish yellow; femora yellow, the tips narrowly but conspicuously
dark brown, the posterior femora more brownish yellow but with thetips still conspicuously contrasting; subterminal region of femora
indistinctly bdghtened; tibiae dark brown, the extteme base pale, theextreme tips blackened, with a f.aintly indicated paler subterminal.ingi tarsal segmentsone to threelight brown with dark 6ips,the terminal tarsal segmefitsuniformly darkened.Wings broad, with a strong
yellow tinge, the costal region mofe saturatedl small brown clouds at
odgin of Rs; in cell R near mid-length of R; at Scz; along the cord
but not including the fork of.M;atir1y cloud at fork of Mr1.ztveinsbrownish yellow, darker in the infuscated areas,Venation : Rs tather
short, strongly arcuated at oigini ce|| rst Mz shoft, as in c. brevicellula Ar.nxnNosn.
Abdomen yellow, the extreme latenl margins and a median line
the posterior margins of theon the sternitesblackened,interrupted^tsegments;subterminal sternites infuscated.
Hab, Camercun,
Holotype,6', Edea, March 27, rg22 (I. A. Rus).
Type in the collection of the Catnegie Museum'
By the author's key to the African species of. Clydonodozus(Rev'
Zool. Afui.caine,XI, fasc. I: ro-rr; 1923),the present fly would run
to C. angustifasciatusAtexlnorn, differing in the broad wings,immaculate mesonotum and sparse wing-pattetn.