a .ft~-1;-±~1'~•*·ic.f
Transactions and Proceedingsof thePalaeontological Society of Japan
New SeriesNo. 20Palaeontological Society of Japan

December 30, 1955

CONTENTS
TRANSACTIONS

Page274. Notes on Cambrian Fossils from Yentzuyai, Tawenkou, in Shantung.............................................................................. Teiichi KonAYAsm 89
275. Cretaceous Echino:d, Niponaster from the Island of Awazi, Japan ......
Akira MoRISHITA 99
276. Amussiopecten from the Miyazaki Group, Miyazaki Prefecture, Japan(Paleontological Study of the Tertiary Miyazaki Group) ... Tsugio SnuTo 101
277. Les Ammonites recueillies dans le Groupe de Kuruma, Nord du Japon
central ........................................................................... Tadashi SATo Ill
278. Miocene Mollusca from Nota Peninsula, Japan. Part 1, (l) .............. .
... ...... ... ... ... ... ... ... ... ... ... ... ... ... ... ... ... ... ... ... ... ... ... ... ... Ko:chiro MASUDA 119
PROCEEDINGS .......................................................................................... 128

President:

Hisakatsu YAOE

Councillors:

Kiyoshi AsAr-:o, Riuji E:-.oo (Business and Accountant), Seido

ENoo, Haruyoshi FuJIMoTo (Publication), Shoshiro HANZAWA, Kotora
IIATAI, lchiro HAYASAI<A, Nobuo limBE, Teiichi KoBAYASHI (Editor), Jiro
MAKIYAMA, Tatsuro MATSUMoTo, Masao MINATO, Tokio SHIKAMA, Fuyuji
TAKAI (General Affairs), Hisakatsu YABE.

All Communications relating to this Journal should be addressed to the
PALAEONTOLOGICAL SOCIETY OF JAPAN
Geological Institute, Faculty of Science, University of Tokyo, Japan

Trans. Proc. P.1laeont. Soc. Japan, N. S., No. 20, pp. 89-98, pl. 14, Dec. 30, 1955

274.

NOTES ON CAMBRIAN FOSSILS FROM YENTZUYAI,TAWENKOU, IN SHANTUNG*TEIICHI KOBAYASHI
Geological Institute, University of Tokyollf~~:kP( llitirffii!}iQ)~JiUe.f~l:. -?I.'-(-: KEEGENBERG O:>!*~r\i."'(:> MoNKE Q) )I -1 7"
Z.liJf~ L -c

Damesellidae Q)~1.:~. Damesella paronai "'(:> Ampholon deois Q)flJ-i"Q)fthl:. -?I. '-c if

4Ji~Z..Il!.HI Lt.:o MoNKE Q)!Jj{tf/,l:f:t;:. Myoua Q)?J"iit."CI.'~~i:,~llU>~~~t L-c~ftiii~ L-(~.><

0

1' u ~{ -1Yentzuyai or Yentsyyai near Tawenkou
on the Chinp'u railway line is a famous
fossil locality in Shantung, China andCambrian trilobites procured therefrom
by F. Kom~FER were thoroughly investigated by ·Dr. H. MoNJm. A revision is,
however, desirable to be done in thelight of modern advancements, as it
was about half a century ago. As aresult of such a study on a part of
MoNKE's types in addition to KEEGEN·
BERG's collection from Tawenkou, sundry
hotes are given here as follows :6. Hypostoma of Damesella paronai(AIRAGIII).

7. Hypostoma of Amplwlmz deois(WALCOTT).

8.

AmPhoton (Smtill) tel/us(WAL·

COTT).

9. E11torachis, a new subgenus of
Psilaspis RESSER and ENilO.
10. J\l[yona flabelliformis KonAYASHI.
For the privilage of studying these
specimens I am very grateful to Prof.
H. 0. ScmNDEWOLF of the University at
Tiibingen, Germany.

1. Taxonomic Notes on the Trilobitesfrom Yentsyyai.
2. Immature forms of Miomeric
Trilobites.
3. Early Ontogeny of the Damesllidae.
4. Telnistimz magna KoBAYASHI, new. species.
5. Steplwnocare (?) bergermzi KoBA·

1. Taxonomic Notes on the Trilobitesfrom Yentsyyai.
Putting aside some bypostomata andlarval forms of doubtful position MoNKE(1903) described eight species of trilobites as follows :Agtzostus koerferi MoNKE=Pseudagttostus douvil/ei. (BRRGEJWN ), 1899.
Liosft·acina krausei MoNKr.:

YASIJI.

Teinisti'<m la11si MoNKE
Teinistio11 /ansi MoNKE= {·Teinistion monlwi KoBAYASHI, new species. See page 93.

* Read at the 61th l\lecting of the Palaeontological Society of Japan, June 18th, 1955, at Kyoto;received June Ill, 1955.
89

Teiichi KoBAYMIIT

90Blackwelderia ex. sinensis (BERGERoN), 1899
Teinistion sodeni MoNKE= {Blackwclderia paronai (ATRAGlll), 1903
an arcuate anterior margin and divided
Drepanura premesnili BERGERoN
Drepanu.,.a kettele1·i MaNKE
into three subequal lobes by axial furStephanocare sincmis BERGERON
rows and into some 9 segments by
Stephanocare (?) sp.=B!aclnllelde1·ia
transverse furrows ; axial lobe distinctlymonkei (WALCOTT), 1911.
elevated above the gently convex pleural
Agnostus koerferi is inseparable fromones ; some axial rings bearing relatively
Agtwstus douvil/ei. Agnostus Pii AIRAlarge median tubercles ; marginal border
GIII, 1902, is probably a small form ofabsent or unpreserved.
the same species in poor preservation.
All of them occur in the Kushan stage
3. Early Ontogeny of the Dameselin Shantung and belong to Pseudagnoslidae.
tus (KonAYASlll, 1939). As noted elseThere are several metaprotaspids ofwhere, (19,!2), the cranidia on whichpolymeric trilobites, but they are mostly
MoNI<E founded Tei11isti01z sodeni appear
ill-preserved. Two best ones illustrated
to be those of Blackwelderia ex. sinensis
in figs. 14 and 15 are similar to MoNKE'sin immature stages. ' MaNKE's pygidia(Taf. 6, Fig. 16; pl. It Fig. d), but disof the same species may be those ofagree with his primarily in the posteriorBlackwelderia paronai in the stage
outline. These two larvae are not muchbefore the sixth pair of short spines
different in size of the dorsal shield or
appear. MaNKE's Stephanocare (?) sp.
in proportion between the head and taiL
represents Blackwelderia monlwi.
The divisional line and occipital furrow
are fairly distinct. The axis is more or
2. Immature Forms of Miomericless fusiformed in the A form (0.79 x
Trilobites.
0.64 mm), while it narrows regularly
Among MnNK!i's larval shields the
backward in the B form (0.78x 0.65 mm).
smallest and simplest is an elliptical
In both of them the dorsal furrow is
one about 0.5 mm. long. (Taf. 6, Fig. 15).
deep aQd expanded into a triangular
It is truncated by a straight articulating
depression at the anterior extremity.
margin whence a broad axial lobe tapers
There the periphery of the shield isdistally, forming a cone. It is quitetransversal or broadly arcuate. The
segmentation becomes indistinct in the
different from any larval form of polydistal sides of the axis. No eye-band ismeric trilobites, but is simila.r to an
seen. The axial lobe of the pygidium
immature pygidiurn of Peronopsis intiger( BARRANDE), 1852, although the marginalis apparently composed of 2 rings anda terminal lobe. Pleural furrows areborder is not shown.
On the slabs I examined none looks
indiscernible. The posterior margin of
the pygidium is nearly transversal andlike that larva, but there are a few
there appears to be a tiny process at
small pygidia which are suggestive ofthe lateral end.
Eodiscidian affinity. They resemble at
W ALcoll''s trisegmented metaprotaspid
the same time the pygidium of W AL·
corr's Slmmardia (?) sp. (1913) fromwhich was procured from Yenchuang;the Changhia limestone in Shantung.
(C6). Shantung, in association with
The shield is long, ovate, truncated by
Aguostus koerferi, Shantzmgia spimfera

Notes on Cambrian Fossils from Yentzuyai, Tawenlwu, in Shantung

and Siepllanocare. ricltthofeni is similar
to the larva MaNKE illustrated, but theglabella is very broad in WALcoTT's.
A prec~se comparison among thosefour larvae shows some differences apart of which is attributable to thematter of preservation and another partto growth change among rnetaprotaspids.
Still another part depends upon taxAgnostus koerjeri ......... ..... .46
Liostracina krausei ............ 52
Teinisfi011 {ansi ............... ... 72 )Drepanura premesnili ......... 79
Drepanura kctteleri ............ 9
Stephatzocare richthojeni 155}
Steplzanocare (?) sp. . ..... 1
Hypostomata ............... 20

91

onomic difference. Nevertheless they areso similar to one another that they arepresumed to belong to the same family.
For the purpose to determine theirtaxonomic position the frequency ofoccurrences is now brought into consideration. The number of dismemberedcarapaces counted on MaNKE's Tafel 9
is tabulated below.

Agnostidae .............. 46
Liostracinidae ......... 52
Dorypygellinae ...... 160}

Jlam,~llid"

316

Damesellinae ......... 156

Thus it is evident for the larvae thatthe probability of their reference tothe Damesellidae is so high that the
Liostracinidae can be ignored. It isthen extraordinarily interesting to findtwo kinds of metaprotaspids with andwithout a pair of lateral processes on
the pygidium. The spiniferous form
must be the larva of the Dorypygellinae.
assuming the process to be the progenitor of a long spine. The other would
be of _Stephatzocare, and if so, of Ste-

Phatwcare richtlzofeni.
In 1939 ENno has illustrated an excellent ontogenetical series of Blacllwdderia
tJttadrata which is intimately related toE. paronai. if not identic1l. There are,however, some gaps in the series between the protaspid and meraspid stages(Figs. a-c. e-g). His latest anaprotaspids(1939 B, pl. l, figs. 1(}--12) have transitory
pygidia or protopygidia. no mention
being, however, given of their segmentation. The proportional size of thepygidium to the cephalon is much smaller in them than in the metaprotaspids
above referred to. Supplemented withthem, the early ontogeny of the Dame-sellidae can now be figured out completely.
The late anaprotaspid of B. quadrata
has the eye- band extending from tl:eneck of the frontal lobe laterally andthen posteriorly along the lateral margin.
In this respect it is similar to tl:e middle
metaprotaspid of Redlichia chinensis
and the late anaprotaspid of Paradoxides
pimts (?), although the cheeks are notsegmented as in R. chi11e11sis and thefrontal lobe of the glabella not bilobed
as in P. pinus.
It is allied also to Palaeolenus deprati
MANstrv in the anaprotaspid stage. but· in the latter the preocular area issmaller and narrower ; the glabella contracted at the fifth lobe (Lu. 1942). The
cephalon of the latter. 0.7 mm long, isquite distinct from that of B. qzeadrata
in the metaprotaspid stage or meraspid
stage, but very similar to the cephalon(fig. le) in the nepionic stage which
WALCOTT (1913) referred to Ampltoton

deois.
In the anaprotaspid stage the Damesellidae are quite distinct from theOlenellidae and closer to the Redlichidae,

92

Teiichi KoBAYASHI

Dolerolenidae and Paradoxidae than to
the Olenidae, Andrarinidae (Liostracidae)
and Asapbiscidae (Blainia ?).
The metaprotaspids of the Damesellinae are quite different from those of
the Olenellidae and Paradoxidae. In
the Damesellidae there is no genal
spine as seen in Olcmrs, Sao or Dalmanititza. Furthermore it is remarkablethat the eye-band is indiscernible andeither the frontal lobe or the occipital
one is no more expanded than the threeintermediate lobes.
The glabella isgently tapering back at the beginning,
but later it becomes subcylindrical androunded in front. On each side of thefrontal lobe there is a triangular depression. It becomes larger and united
with its cotmter in a lunate depressed
frontal border in the meraspid stagesof Blackwelderia quadrata. This is anincipient preglabellar depression whichis quite developed in the cranidium of
Bla~kwclderia paronai in the full grownstage.
Eye bands exist on the meraspids of
B. quadrata. By their inward migration
there appear free cheeks. Still later
the eyes become smaller and eye-ridges
are effaced as can be seen in the adultof Blackwelderia.
The early meraspid cephalon of B.
quadrata is somewhat similar to that
W Ar.coTT ( 1913) refen·ed to Amp/wtotz
deois (fig. 1d) or Fuclwuia manclmriensis.
Similar larvae are, however,found in the east of Liaoyang in aTangshihan shale with Proasaphiscus
yabei but without Amplwton or Fuclloui'a.
MoNK F.'s meraspid cephalon ( Taf. 6, Fig.
17) is different from the precedings in
the absence of eye-bands. Whether it
belongs to the Damesllidae or the Liostracinidae is a question.
The early holaspid cephalon of Liosft·acitza krausei (Taf. 3, Fig. 13) can

easily be distinguished from those of
Stephanocare richthofetti (Taf. 7, Fig.
6), Teinistion lattsi (Taf. 4, Fig. 6) andDrepamera premesnili (Taf. 5, Fig. 9)
by the short glabella and obscurity ofthe eyes. The eyes are small andlocated far anteriorly in D. premesnili,
while they are large in Teinisti01t andStep!wnocarc and located in the middle
in S. t·ichthofeni and in the posterior in
T. /ansi. Thus these cranidia are already not essentially different from theiradults.

4. Teinistion magna KonAYASIII,
new species
Pl. 14. Fig. 12.

This cranidium from Tawenkou is thelargest of the genus I ever saw. It
measures 14.5 mm in length.
Glabella long, conical, provided withan occipital and two lateral furrows:
fixed cheek narrow ; eyes fairly large,
opposed in the posterior of the cranidium: eye-ridge quite oblique; two weakoblique folds seen behind the ridge;
anterior rim straight and remarkably
erected: a groove between the frontal
limb and rim very broad and concave.
A further note given in the description of the genus.
Genus Teini'stion MaNKE, 1903
1903.

Teinistio11 MaNKE, fahrb. ktitJigl. Preuss. Geol. Landesanst. 11. Bergakad. Bd.
28, Hjt. 1, p. 117.

Teinistio11 WALCOTT, Cambrian Faunas
oj China, p. 109.
1935. Teinistion KoBAYASHI, four. Fac. Sci.
Imp. Ut1iv. Tokyo, Sec. 2, Vol. 4. Pt.
2, p. 254.

1913.

Diagrzosi's:-Dorypygellinae with broad
cephalon, cylindrical or truncato-conical.
narrow glabella, eyes of medium size
located posteriorly and oblique eyeridges.

Notes on Cambrian Fossils from Yentzuyai, Tawenkou, in ShantungType species :-Tei11istion lansi MoNK!\,
1903.

93

Specific list :-The present determina"tion is cited on the right side.

Tcinistion /ansi 1\loNKE, 1903, in fig. 1, pl. I only. Teinistion monkei KonAYASH!, new species
Blackwelderia ex. sinensis (BERGERON)
Tcitzistion sodeni 1\lo:--:KE, 1903{
B/ackwelderia paronai (AIRAGHI)
Tcinistion a/con (WALCOTT), 1913
Dorypyge//a a/COli WALCOTT, 1905
Teinistion subconica SuN, 1924
ditto
Teinistion truncatus ENDO, 1937
ditto.
Tcinisiiotl sulcatus ENno, 1937
Doryj>yge/la ? sulcata (ENoo)
Tcinistio11 obtusus ENDO, 1937
Dorypyge/la ? obtusa (ENDO)
Teinistiotl lanci{orme ENDO, 1944
Teinistion ? lanciforme ENDo

Remarks :-I once considered that
Teinistion is a member of the Emmerichellidae, but as discussed later (1941),
it is a close relative of Dorypygella,
as WALCOTT has synonymized it with
Tcittislimz. In MaNKE's ontogeneticalseries of T. la11si the youngest cranidiurn(Taf. 4, Fig. 6) looks more similar to
Dorypygella than Teiuistion in the adultstage. It has a conical unfurrowed
glabella and large eyes, but no obliquefolds on fixed cheeks. Diminution of
the eyes, development of eye-ridges andoblique folds and the sinuation of theanterior border are shown by MoNKE to
be morphic changes through growth.
According to his ontogenetical seriesof T. latzsi, the cylindrical glabella isintroduced from the conical one. The
cranidium in Taf. 4, Fig. 2 is assigned
here as the lectotype of Teinistion !ansi.
Because it is quite improbable thatthe conical glabella appears again in astill later stage, Teinistiou monkei is
proposed for the cranidium in Taf. 4,
Fig. 1, MoNKE, 1903, having a truncatoconical glabella.
There are, in addition. a few otherspecies having this kind of glabellae.
ENno's trrmcatus is different frommonkei in the absence of the obliquefolds on the fixed cheeks.
SuN's co11ica has pustulose test, while
all others of this genus have smoothones. The development of the preglabellar area is the distinct feature ofof T. magna.
JJistribution:-Kushan stage of Eastern
Asia except for T. (?) lanczformes fromthe Taitzuan of Hualienchai, Southmanchuria.

5. Step/wnocare (?) bergeroni
KouAYASIII

Pl. 14, Fig. 9.

Damesellid pygidia have spines in 5
to 8 pairs one or two pairs of which
are often longer than the others. Fiveor eight pairs of spines are, however,met with not so commonly as 6 or 7
pairs. Step/zmwcare (?) bergcrmzi fromthe Drepanura zone of South Korea issuch an uncommon form having 5 pairsof spines on the pygidium.
A pygidiurn from Yentsyyai, fits with
the South I\orean one nicely. 1he lackof an intramarginal groove in the
Shantung sr:ecimen does not mean any
taxonomic distinction, because thegroove in the Korean specimen must bea surface impression of the doublure
which came out secondarily.

6.

Hypostoma of Damesel!a paronai(AIRAGill)

Pl. 14, Fig. 8.

All of detached carapaces on ·a marlTeiichi KoBAYAsHI

94

specimen from Tawenkou belong exclusively to Cheirurus parouai ArRAGHI,
i.e. Damesella blackwcldcri WALCOTT.
Among three hypostomata in the smallslab a \vell preserved one reveals agreat similarity to WALCOTT's (1913, pl.
10, fig. 13). but the posterior outline isremarkably expaned laterally.
The central body is subelliptical, but
the posterior ridge and groove are fairly
acute. Surrounding its middle and posterior parts and separated from it by arelatively deep groove, the flat borderis expanded laterally. The posteriorborder which forms a shallow sinuation
is narrow. The antero-lateral wing is
undeveloped.
7.

Hypostoma of Am jJlwtou dcuis(WALCOTT)
Pl. 14, Figs. 13a-b.

A hypostoma on a limestone slab
from Tawenkou (?) is referable to AmPiwlotz deoi's, because all other dismembered carapaces on the slab belong to
the .species, if Pcrouopsis chineusis isoverlooked. It resembles wALCOTT's
hypostoma of the species (1913, pl. 22.
fig. 1c), but by cleaning the matrix, theanterior wings are found much larger.
The outline of the hypostoma is
subtriangular ; central body subovate,strongly vaulted, provided with a low
lunate band on the rear side and surrounded by a narrow rim: anterior.margin slightly arcuate; antero-lateralwing very large, trigonal, depressed anddistinctly slant laterally.
This is somewhat similar to the hypostoma of Ampholon (Suuia) typica
KoBAYASHI (1912, pl. 1, fig. 15), butevidently more triangular in outline, inwhich respect the hypostoma of Allwbaskia belus (WALCOTT), 1916, may be
Comparable to it.

Amphoton (Stmia) tel/us (WALCOTT)

8.

Pl. 14, Figs. 10-11.

LonclwcePfwlus tellus was proposedby WALCOTT (1913) for a poorly preserved cranidium from Yenchuang,
Shantung. Though not much better
preserved, some specimens from Tawenkou allow me to figure out its cephalon.
Glabella subquadrate. a little longer
than broad, gently convex and unfurrowed except a shallow cccipital furrow ;a long spine protruded from a small
subtriangular neck ring ; eyes large, setclose to the glabella ; frontal border
fairly large, more or less elevating upand for-ward; free cheek provided witha genal spine; facial suture anterior to
the eye subvertical, but a little bowing
laterally.
Such a nuchal spine is seen in Annamilia, Redliclzaspis, Loncl?occplzalus,
Saratogia, ldahoia and a few othergenera, but they have frontal limbs ofdifferent size. In my opinion this ismost probably a Stmia having a shorter
glabella than that of Stmia typica.
9.

Etztorac/u's, a new subgenus of
Psila!'pis RESSER and ENno

Psilaspis typified by Psilaspis manclzuriensis Rr:ssER and ENuo has aconical glabella. Entorachis is proposed
here to include Psilaspis. but having
a parallel-sided glabella. A11omocare
alc'ione WALCOTT, 1913, is the ·type of
this subgenus and Ptychoparia (Conocep/wlites) mcmor REED, 1910, can safely
be referred to it.
The glabella is also parallel-sided in
Hmziwa and some species of Eymckops,
but the eye- band· is much ·larger inthem than in Entorachis. The posterior
end of the eye is located close to theglabella and the frontal margin of thestraight and transversal· in Ilaniwa,

Notes on Cambrian Fossils from Yentzuyai, Tawenkou, in Shantung

95

whereas the margin is rounded and theend of the eye apart from the glabella
in Ettioraclzis.
Insofar as the cephalon is concerned.
the most resembling may be Wilbemia
WALCOTT, 1924, but it is distinct fromEntorachis in two pairs of lateral furrows distinctly impressed on the glabellainternally. They are far from confusion,
if their pygidia are compared. Pleuralfurrows are quite pronounced in WAr.co·n's Ptychoparia ? diadcmata (1899)
for which RESSER (1937) proposed Withernia lmdsonensis. The furrows arewell developed also in WILSoN's Wilbemia sp. indet (1954).
The distribution of the subgenus is
re~ricted to the Middle Cambrian in
Eastern and Southern Asia.

Compared to Entoraclzis alcione theglabella is more slender in this species.
In alcione there is a narrow intramarginal ridge which is most probably theimpre...~ion of the inner edge of the
doublure. It is abruptly bent inward
near the genal angle. Entorachis memor
REED (1910). especially the cranidium
in fig. 4, pl. 4, 1oo1's similar to thisspecies. but distinct in the strong occipital furrow. According to REED the
occipital ring carries a median tubercle:
axial furrow pitted at the anterior end
and the test granulose. Furthermore
the marginal furrow appears more distinct, although it is said indistinct andshallow.
Occurreuce :- Tawenkou, Shantung.

Psilaspis ( Eutorachis) shautuugettSis
KoBAYASHI, new species

1'1. 14, Figs. 5-7.

Pl. 14, Fig. 1-4.

Cephalon exclusive of genal spines
semicircular. Cranidium as long as
broad; glabella outlined by dorsal furrows long, cylindrical, rounded in front,
gently convex and devoid of lateralfurrows ; fixed cheek relatively broad:eyes fairly large, semicircular, opposed
a little posterior to the middle of theglabella and connected with it by anoblique and straight narrow eye-ridge;
frontal limb gently inclined forwardand merges with a flat or a little concave frontal border. Free cheek with agenal spine. In the internal view the
doublure . is remarkably broadened at
the genal spine. Facial sutures shortand diagonal behind the eyes and lessaivergent in their anterior.
Pygidium semicircular; axial lobe
delimited by a deep axial furrow oneach side and elevated above the broadgently convex pleural lobes ; marginalborder concave.

10.

Myotza jlabclliformis KoBAYASHIThis is an enigmatic creature possiblyof ArUu·opodan nature. Jt was·· firstfound in the Drepamera zone of the
Seison slate in South Korea which belongs to the Kushan stage. It is extraordinarily interesting to find the samekind of fossils in the same stage at
Yentsyyai in Shantung.
With the Korean specimens it was
ascertained that the carapace is equivalved and almost equilateral. The
median part of the dorsal margin appears somewhat protruded above in formof an umbo, but there is no hinge apparatus. Therefore it is probable thatthe two valves were fused along thedorsal margin.
The Shantung specimens are internalmoulds of detached valves. Because
they are detached from each other, it
is presumable that the two valves wereconnected along the dorsal margin notin the whole length but only in theumbonal pol'tion. The· carapace valve
is taller on one side than on, . the other,

96

Teiichi KoRA y As Ill

but it is difficult to answer which is thefront or back. It is liat but abruptly
turns inward near the periphery.
Internally there are numerous radial
ridges which are separated into an innerand outer series by a deep and broadconcentric sulcus. This aspect of internal relieves reminds me of the diver·
ticula of the digestive tube in Burgcssia
bella WALCOTT. The carapace is, however, not truncated in the rear part asin that species.
Because there is no trace of trilobation, Myona may be a member of the
PEeudostraca or Hymenocarina. Theinternal structure prevents me to refer
Myoua to the Ostracoda. Ribeirida or
Bradorina. It may be combined with
Septadella STUBBLEFIELD. 1933. in an
unnamed group of the Pseudocrustacea.
The lack of the concentric sulcus is thedistinction of that genus from !11yo1la.
Finally it is noted that !11yona flabel!Tformis occurs at Tayiiyaotze, Tsinghuiho region, Inner Mongolia ( Wi7Kil'J1fr~
tr7.(~:g-r) in the Dre pamwa zone together with Drepamtra premesnili,
Blackwelderia pm·onai, B. sinensis,
Stephanocare richthtofeni, Lioslracina
krausei, Ordosia fimbricauda and Loreuzella sp. (Lu, 1954).
References
AIRAGHI, C. (1902), Di Alcuni Trilobiti dellaCina. Atti della Soc. Italimza di Sci. Nat.
Vol. 41.
BARRANDE, ]. (1852), Systeme Silurien duCentre de Ia Boheme, Vol. 1.
ENDO, R. (1939A), The Post-Embryonic Development of Blackwelderia quadrata ENDo.
Proc. Imp. Acad. Tokyo, Vol. 15.
(1939B), Description of Blackwelderia
quadrata, n. sp. and its post-embryonic
Development.
Bull. Central Nat. Mus.
Manchoukuo, No. 1.
(1944). Restudies on the Cambrian For-mations and Fossils in Southern Manchoukuo. Ibid. No. 7.
and RESSER, C. E. (1937), The Sinianand Cambrian Formations and Fossils ofSouthern Manchoukuo. Manchuria11 Sci.
Mus. Bull. 1.
KOBA Y ASH!, T. (1935 ), Ti1e Cambro-Ordovician Formations and Faunas of South
Chosen, Palaeontology, Pt. 3, Cambrian
Faunas of South Chos~n with a special
Study on the Cambrian Trilobite Generaand Families. ]our. Fac. Sci. Imp. Ut1iv.
Tokyo, Sect. 2, Vol. 4, ·Pt. 2.
(1939), On the Agnostids, (pt 1). Ibid.
Sec. 2, Vol. 5, Pt. 5.
- - (1941-42), Studies on Cambrian Trilobite
Genera and Families (1-4). ]apa11. four.
Ceo/. Ceogr. Vol. 18.
(1942), On the Dolichometopinae. four.
Ji'ac. Sci. Imp. U11iv. Tokyo, Sec. 2, Vol. 6.
Pt. 10.
- - (1944), On the Eodiscids. Ibid. Sec. 2,
Vol. 7, Pt. 1.
--and KATO, F. (1951), On the Ontogeny
and the Ventral Morphology of Redlichia
chinensis with Description of Alutella tlakamurai, new gen. and sp. Ibid. Sec. 2, Vol.
8. Pt. 3.
Lu, YEN HAo (1942), Some Lower Cambrian
Trilobites from Chintingshan. N. Kueichou.
Bull. Ceo/. Soc. China, Vol. 22.
- - (1953), Two new Trilobite Genera of theKushan Formation. Acta Palaeontol. Si11ica,
Vol. 2, No. 4.
MoNJ<E, H. (1903), Beitrage zur Geologie vonSchantung, 1. Obercambrische Trilobiten
von Yen-tsy-yai. ]ahrb. kti'nigl. Preuss, Ceol.
Lat1desa11st. rmd Bergakad. fur 1902, Bd.
23. lift. 1.
RESSER, C. E. (1937), Third Contribution to
Nomenclature of Cambrian Trilobites.
Smithson. Misc. Coli. Vol. 95, No. 22.
REED, F. R. C. (1910), The Cambrian Fossilsof Spiti. Palaeontol. I11dica, Ser. 15, Vol.
7, 111em. No. .1.
SMITH, S. and STt:BBLEFIELD, C.]. (1933),
On the Occurrence of Tremadoc Shales in
the Tortworth Inlier (Gloucestershire) withNotes on the Fossils. Quart. four. Ceo/. Soc;
Lot1do11, Vol. 89.

Notes on Cambrian Fossils from Yentzuyai, Tawenkou, in Shantung
STj6RMER, L. (19·12). Studies on Trilobite
Morphology, Pt. 2. The Larval Development,
the Segmentation and the Sutures and their
Bearing on Trilobite Classification. Norsk
geo/. Tidsskr. 21.
- - (1944), On the Relationship and Phylogeny of fossil and recent Arachnomorpha.
Skr. Vid.-Akad. i Oslo, 1, Math-l'>taturv. Kl.
19·1, No.5.
SllN, Y. C. (1924). Contributions to the Cambrian Faunas of North China. Palaeo/Ito/.
Siuica, Ser. b, Vol. 1, Fasc. 4.
WALCOTT, C. D. (189!!), Cambrian Fossils in
Geo:ogy of the YelloW:ltone National Park.
U. S. Ceo/. Surv. ltio11ogr. 32.
- - (1911), ?>Iiddle Cambrian Merostomata.

91

Smiths. Misc .. Col/. Vol. 57, No. 2.
- - (1912), Middle Cambrian Branchiopoda ..
Malacostraca, Trilobita and Merostomata.
Ibid. Vol. 57, No. 5.
- - (1913), The Cambrian Faunas of China.
Research i11 China, Vol. 3.
- - (1916). Cambrian Trilobites, Cambrian
GeoloJ.,<y and Paleontology, 3. Smiths. Misc.
Coli. Vol. 64, No. 5.
- - (1924), Cambrian and Lower Ozarkian
Trilobites. Ibid. Vol. 75, No. 2.
(1925), Cambrian and Ozarkian Trilobites. ibid. Vol. 75. No.3.
WILSON, J. L. (1954), Late Cambrian and
Early Ordovician Trilobites from the Marathon Uplift, Texa>'. four. Pal. Vol. 28.

98

Teiichi KOBAYASIIl

Explanation of Plate 14
Psi/aspis (Eilloraclzis) slzantrmgensis KonAYAsHI, new species ............................................. p. 95
Figure I. Holotype cranidium, x 1
Figures 2-4. Cranidium, free cheek and pygidium, x 1
Myona flabe/lifonnis KoBAYASHI . . . . . . . . . . . . . . . . . . . . . .. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . p. 95
Figures 5-7. a-c valves, x 4
Damesella parotwi (AI RAG 111) .......................................................................................... p. 93
Figure B. Hypostoma. x 1.5
Steplzanocare (?) bergeroni KoRA Y AS HI . . . . . . . . . . . . .. . .. . . . . . . . . . . . . . . . . . . . . . . . . . .. . . . . . . . . . .. . . . . .. . . . . . . . . . . . . . .. . p. 93
Figure 9. Pygidium, x 1.5
Amplzoton (Sunia) tel/us (WALCOTT) ............................................................................ p. 94
Figures 10-11. Cranidium and free cheek, x 1.5.
Tei11istion magtra KYBA Y AS II I, new species . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .. .. .. . . . . . p. 92
Figure 12. Cranidium, x 1.5
Amphototr deois (WALCOTT) .......................................................................................... p. 94
Figures'13 a-b. Ventral and lateral views of a hypostoma, x 3
Metaprotaspids of Dorypygellinae . . . . . . . . . . .
. . . . .. . . .. . .. .. .. . .. . . . . . . . .. . .. .. .. . . . . .. . . . . . . . . .. .. . . . . . . . . .. . p. 90
Figures 14 and 14a. B form, x ca. 30
Figures 15 and 15a. A form, x ca. 30
Early ontogeny of Damesellidae . . . . . . . . . . . . .. . . .. . .. .. . . . . . . . . . .. . . . . ... . .. . .. .. . .. . . .. . . . .. .. . .. . .. . ... .
p. 90
Figures a-c. Early, middle and late anaprotaspids of Blackwe/deria quadrata ENDO
respecJively in Figs. lb ( x 36). 3h ( x 10) and 5b ( x 20) in EN no (1939A).
Figure cf Metaprotaspid of Damescllinae or Steplzanocare in MoNKE, 1903, Taf. 6,
Fig. 16. X 15.
Figures e-g. Early, middle and late meraspid cephala of B/ackwe/derin quadrala,respectively in Figs. 6b ( x 15), 7h ( x 14) and 9b ( x 10) in ENDO (1939A).

T. KoBAYASHI: Cambrian Fossils from Yentzuyai

14

PLA TE

7

8

ll

''

15b

12

®
J3a

13b

b

15a

c

e

r1r

g

Trans. Proc. Palaeont. Soc. Japan, N: S., No. 20, pp. 99-100, Pl. 15, Dec. 30, 1955

275. CRETACEOUS ECHINOID, NIPONASTER FROM
THE ISLAND OF AW AZI, JAPAN*
AKIRA MORISHITA
Geol. and l\-lineral. Institute, Univ. of Kyoto

QQ)ailli*h·~~HI Lt.: Nipouaster 1:.-01. ·-c: Niponaster /iltli.UiiiJI.:~tlllJJll'iil)~t!ffi*il•l?ilfi
HI Lt.: Anancltylinarmn sp. J.:')c1 L J. LAMBERT iJ:.g.;t t.:.fiO{#~"t:!;.'!>iJ;, ffil;l[\~iJ;ftjjl)i~. lf'!il.: rffii<1)/j!ij:fj&iJ;Ii ~ c~ I?~ vel. ·t~1. 'o ali:C~H14-Iill. ~MI-:iJ·I?im IIJ Lt.:"-:;t.':i.: -01. '"C. I., <-0:fJ•il)Wf L1. ·lf'!i~~fifiJEJfll.:ii'iltu Lt..:off .F d1c

One specimen of echinoid from theIzumi Group of Awazi was just examinedby the author. The occurrence of somefossil echinoids was reported by SAsAI
and others together with several Ammonites, Belemnites, Iuoccramus etc. (jour.
Gcogr., vol. 6, p. 109, 1894; jour. Geol.
Soc. japa~t. vol. 43, p. 600, 1936) SAsAI
referred an European species, Ananchytes
ovatus (LESKE) to one of his material
without giving any palaeontological note.
This form to which the present specimen under examination belongs is
identical with Niponaster Jwkkaide11sis
LAMBERT of Hokkaido. As LAMBERT's
description of this species is not very
precise; a supplementally diagnostic
features of Niponaster is given in thefollowing lines.

Description :-Tl:e test is large, highand round in outline. The abactinal
surface is arched from the ambitus to
the apex. The actina! surface is flat orslightly concave and sunken only adorally. The posterior interambulacrum ·issomewhat elevated. There is no frontal
depression.
The apical system is small and at
about the centrq. The genital pores areindistinct.
All the petaloidal ambulacra are nearly equal in its shape, namely the odd
one is not well differentiate from theothers. The ambulacra are not depressed. open and nearly reach to themargin of the test. They are straightand gradually widened towards the distal
ends. The poriferous zones are broad
and the interporiferous narrow ( widtl1
at the distal end: 5.8 mm. 2.3 mm). The
ambulacral and interambulacral plates
are larger at the lower part than at theupper. The tubercles are indistinct.
The peristome is at the 1/4 diameterfrom the anterior ambitus, depressed
and transversely oval. The periproct
is oval and inframarginal PO!?terjorly.
The posterior interambulacrum forms
sternum and amphisternous.
The marginal fascicle is indistinct:

Description of Species
Genus Ni'Po1zaster LAMBERT 1920
Niponaster lwkkaidcnsis LAMBERT
Pl. 15, figs. 1-3

1894. Anancllyti11amm sp. indet., K. ]!MBO,
p. 45, Taf. IX, fig. 8.
1950. Nipo11aster lzokkaideiiSiS Til. MORTEK·
SEN. p. 170, fig. 132.

* Read June 18, 1955; received June 27, 1955.
99

100

Akira MORISIIITA

Measurements:Longitudinal diameter: 78 mm (deformed)
Transverse diameter
87 mm (deformed)
Height
44 mm (deformed)

Observations and Remarks : -This
deformed specimen seems to have teen
originally higher and longer than theabove measurements. In view of the
shape of abactinals. the ambulacra IV
is in the front. But the positions ofperistome and periproct in the actinalside clearly show the exact anteroposterior direction. On the abactinal
surface, the left half is narrow comparedwith the widened right. Presumably J.
LAMBERT proposed the genus Niponasterwithout the specimens at his hand.
Therefore, Jn.IBo's poor description, ofAua11chytinarum sp. indet. seems to be
only one to show certain characters of
NipotZaster. ]IMBo shortly mentions asfollows: "Zwei sehr zerquetschte Stuckeliegen als einzige Echiniden-Reste vor.
Die Schale ist sehr abgerieben. tmd vonder Unterseite ist nur der Rand
theilweise zu beobachten. l'vlund- und
Afteroffnungen sind nicht erhalten. Die
Kc>rner auf der Obert1ache sind nur an
der Unterseite deutlich untl mit H<>fenversehen. Ambulacra einfach, mit schmalen porenstreifen, breiter gegen den
unteren Theil, aber auf der Unterseite
nicht mehr vorhanden. Die porenpaare
stehen entfernt und einreihig.
Das
Scheitelschild fast compact, aber nicht
gut erhalten.

Fundorte: Mergelknollen von Urokobets und dem oberlauf des Yubari, naheder Miindung des Pankemoyubari."With this description. the actina! feature is not known. The present observations clarified the amphistemouscharacter of the plastron that havenever been accepted by Th. l'vloRTENSEN.
Accordingly the Genus Nipouaster is notallied to Stenonaslcr or Memethiaster(Meridosternata), but to Arnphisternata.
Matrix :-Shale
Geological Horizon:- Minato Shale,
upper part of Izumi Group (up.
Cretaceous)
Locality :-Near Minato-machi, Mihara-gun, Hyogo Prefecture
Acknowledgements. - The acknowledgements must be indebted to Mr. Y.
TucHIGA and Mr. T. YASUDA for theirkindnesses offering the valuable specimen, Mr. Y. MAEDA for his kind suggestion on the geological age and Professor J. MAKIYA!\IA of Kyoto University
for reading over the manuscript. This
specimen is due to be deposited in Mr.
TucmGA's hand.
References]1!\IBO. K. (1894). Beitrage zur

Kenntniss derFauna der Kreideformation von Hokkaido,
Palaeontologische Abhandlzmgcu, N. F., VI.
MoRTENSEN, Th. (1950), A Monograph of
the Echinoidea, v,, Spatangoida 1.
SA~Al, H. (1936), The Izumi Sandstone Series
in Awazi, four. Geol. Soc. Japan, J?ol. 43,
No. 515.

Explanation of Plate 15(All figures in natural size)

Niponaster hokkaidensis LAMBERT
Fig, 1. Abactinal side. Upper part of the figure shows the front.
Fig. 2. Posterior profile. Two ambulacra show the posterior paired ambulacra.
Fig. 3. Actina) side. Upper part of the figure shows the front. Peristome and the amphisternous
character of plastron are distinct.

A. MoR l SHIT A : Niponaster

PLATE

15

Trans. Pro:. Palaeont. Soc. Japan, N. S., No. 20, pp. 101-110, Pis. 16-17, 4 text-figs, Dec. 30, 1955_

AMUSSIOPECTEN FROM THE MIYAZAKI GROUP,MIYAZAKI PREFECTURE, JAPAN,(Paleontological Study of the Tertiary Miyazaki Group)*276.

TSUGIO SHUTO
Department of Geology. Faculty of Science, Kyushu University.

B'-..1o!illf~ Amussiopecten: Amussiopecten 1tJt*J:fl'l'JTtltQ)/Y'i, "'(:~ .Qil;. E ;;j;: C. ~Q)ili~l't!!lJJJ:Q).;z). ·c V l.-? -h'il~.mff!:Q)mi\1~ftli C. L -c-:EIP?h. Jfl:WJ'!YI'J•l? (;t. (:t.-?~ IJ Lt..:fi!i1'1¥&·;'c; ~:h."CI. ·f.t:l. '·~fUit:t.a-~ml!f»)J:.ffilil•l?!f.tfrtltfi A. praesignis~Q)Jfl'i;fifU:fll!(l)~ru.rt~'U:~a-~Y>-cilc.~-t.Q.

z.. rffllil· G~ h t 1t5.1Uf11JiQ).fll!aot'J·fJ: fJ ~~-!*~ Lt..:.

1i ~ (!~ !73

T,,-o forms have been considered as
being closely related to this genus,
Flabellipecten SAcco 1897 and Oopectetz
SAcco 1897 (" Gruppe der Pecte1z
rotmzdatus LAMARCK" of PHILIPPI or"groupe du Chlamys mtmzdata (LA1\IARcK)" of Jean RoGER). These forms
represent the transitional position between Amussi'opecten and Pecten or
Chlamys. Therein H. E. PHILIPPI supposed that Amussiopecten and theallied two genera had differentiatedfrom " normal Aequipectett " in theenvironment of the tropic water in acertain age of older Tertiary. He
furthermore assumed that Amusium
belonged to a quite different lineage
from that of Amussiopecten. However
Amussiopecten actually resembles Amusium RuMrmus 1711 em. Kr.F.IN 1752
in general feature except that thelatter is provided with equally convex
valve and more prominent interior ribs
and is devoid of exterior ribs. On thefoundation of the comparative anatomy
modern taxonomists set Amusium andAmussiopecten in subfamily Amusiinae

Introduction

The purpose of this brief note ismainly to describe two species of Amussiopecten from the Miyazaki group of
Miyazaki Prefecture, south east Kyushu,
with some discussion on their bearing
in biostratigraphy. Before entering into
the subject the definition and thetaxonomic position of the genus arehere remarked.
Genus Amussiopecten, well known insouth Europe as an index genus of
Miocene, was established by SAcco in1897 on the basis of Pecten burdigalensis LAMARcK. The generic diagnosis
may be described as follows : convex
right valve and plano-covex left halve ;both valves provided with numerous
exterior radial ribs, which are d·istinct
near the beak and diminish as shell
grows, and with interior ribs reaching
the ventral margin.

* Read at the Meeting of the Nishi-Nippon
Division of the Geological Society of Japan at
Fukuoka, Oct. 15, 1954; received July 14, 1955.
101

102

Tsugio SHUTO

and Pecten and C!zlamys in subfamily
Pectininae. The present writer adopts
this classification.

Text fig. 1
Distributed Area of the Miyazaki Group.

Various species of Amussiopecteu
have been reported from the Mediterranian basin and the Pannoic basin
in Europe. Th,eir geological ranges
are generally limited in BurdigalianandVindovonian,especiallyinHelvetian, with an exception of
A. burdigalemis, which begins to appear in upper Aquitanian. The genus
occurs, outside Europe, in Egypt, Iranand Indonesia and its age is again
Miocene. However in Japan and adjacent islands it has been regarded as
an index fossil of Early Pliocene rather
than Miocene. Of the three species of
the genus which have been reported inthis country", the most familiar species,
A. praesignis (YoKoYAMA), occurs frequently in the Lower Pliocene on the
Pacific side of southwest Japan anc.J also
in that of Ryukyu and Formosa. Of* Amussiopecten praesig11is (YOI<OYAMA), A.
? yabei NOMURA and A. sp. of 0TUKA.

course, Amussiopecie1t is not utterly
absent in the Miocene strata; it has
been reported from the Sagara group,
the Sizul<awa beds and the Siroyama
sandstone in Sizuoka Prefecture andfrom the Kaisan beds in Formosa.
However, its occurrence in Miocene ismuch less common than in Pliocene,and some paleontologists doubt thatcertain Miocene specimen which wereidentified as A. praesignis do not
actually belong to that species.
The writer has collected a considerable number of specimens belonging to
two species of this genus in the Miyazakigroup. One is A. praesi'gnis (YoKoYAMA)
and found in the Takanabe member.
The other is a new species and found in
the Boroishi, Tano and Takaoka members. As the writer has already described(Smno, 1952) the Miyazaki group
ranges from late Middle Miocene tolate Early Pliocene in age. The Takanabe member occupies its upper partand is assigned to Lower Pliocene.
Therefore the stratigraphic occurrenceof A. praesigms in this group is quite
in harmony with that in other groUJ:S
of the Pacific side of southwest Japan.
The Boroishi and Tano members are
the basal conglomerates and the Takaaka member is the lower middle partof the group ; they are Upper Miocene
in age. Thus in Miyazaki Prefecture,
Amussiopecletz not only occurs in Lower
Pliocene but also in Upper Miccene:
Yet the genus has not been known
from Middle and Lower Miocene.
The writer wishes to eJ\.']Jress hishearty thanks to Dr. Katura Ov AMAof the Geological Survey of Japan forhis helpful suggestions and criticisms
about the present subject. He is alsoindebted to Professor Tatsuro MATsuMoTo of the Kyushu University for his
guidance, criticisms and encouragement

Amussiopecten from the Miyazaki Group, Miyazaki Prefecture, Japanin the· course of this study and for103

reading over the typescript.

Miyazaki Group

Tsuma facies

Miyazaki facies

Aoshima facies!!!

~·~~~-----.-x-----------------.::l:~Takanabe M.
\ Sadowara 111.

Aoshima F-.

\ Orodo M.

.,

~

:::>

~

:X:\

~

.g

i\- - - - - - 1

Tsuma M.

-

Kurokita JVJ.

Kuraoka M.

\- - - - - - - 1\ Togo sub!'.

I\ Takaoka M.

\

Text fig. 2.

X

Stratigraphic Division of the Miyazaki Group.
M ....... member, F ....... formation.
X...... .locality of Amussiopectetl

Systematic Description of Species

K I Y AMA, pp. 34-36.
1928. Pectetl praesignis, YOKOYAMA, p. 335.

1929.

Family Pectinidae Subfamily Amusiinae

Pecten (Amusium) praesignis,

YoKo-

YAMA. p. 11.

1930.

Pecten (Amussiopecten) praesigtlis, OTV·

Genus Amu.~siopecten SAcco 1897
Amussiopectetz praesi"gnis (YoKOYAMA)

1931.

Amussiopecten praesigtlis, MAKIYAMA,

Pl. 17, figs. 1 and 3

1932.

Pecten praesignis, NoMURA and NllNo,

1922.

Pecten praesignis YoKoYAMA, pp. 1-2,

1926.

Pecten praesigtlis,

KA, pp. 507-509.

pp. 12-29.
p. 180.
1933. Pecten (Anmssiopecletl) praesigt1is, NoMf-JRA. p. 60.

pl. 5, figs. 1-3.
YoKoYAMA, p.

357,

pl. 40, figs. 1-2, pl. -t1, fig. 1.
1927.

Pecten (Amussiopecten) praesignis, MA·

1934.

Amussiopecten praesigtzis,

OTUKA,

p.

567.
1936. Pecten (Anmssiopecten) praesignis, No-

Tsugio SHUTO

104

Table 1. Measurements. on the selected specimens of A,mussiopecten praesign_is (YoKOYAMA)I

1Registe r
Numbe
GK-L-rl

Locality

MI-

(mm)

-

r-

..

.

..

..

..
..

1

(mm) (mm)l Line I(%) (%)(mm)______

106.4
4370 :5595 Toriyama 116.4
12.21
10.9
109. 1
98.9
4371
7.3
73.6
76.3
4372
"
"
I
90.0±
4424
"
110. 0±
4430
"
20.7
19.0
4678
"
"
30.4
30.4
4679
"
30.5
33.7
4681
"
4.2
29.7
28.9
4682
"
"
3.5
44.8
44.7
4683
"
31.4
33.5
46!!4
"
54.4
57.0±
4685* "''
106. 0±
111.8
4686

..

Length I

I Len!,.Tth Height Depth.1 H~ge H;L: D/L

..

..

..

I

I
ApicalAngle Valve
Hin••'
LINRibs....,(degree)(%)- - - · -49.8
91.3 10.4
50.7
87.5 10.0
37.0
96.3 9.6
40.0±
53.4
12.6
91.7
16.7 100.0
16.5± 90.0
16.0± 97.1 14.2i
22.0± 99.5 7. 8·
17.7 93.61
30.0±195.0
54.1
94.0
-

42.8
44.8
48.4
44.4
48.6
60.9
54.9
49.0
54.0
49.0
52.7
53.0
48.2

16
15
18
17
18
16
16
18
18
14
18
15
15

--133
130
123
125
131
113
114
113

114

114

115
124

R
L
R
R
R
R
R
R
R
L
R
L
R

I

* triple ridges on the interior ribs
1\IURA and ZINBO, p. 247.
1937. Amusium praesignis, NoMURA, p. 71.
1938. Amussiopectell praesignis, OTUKA, pp.
6-7. pl. 1, fig. 2.
1943. Amussiopecten praesiguis, SHIKA~IA, p.
241, pl. 39, fig. 3.
1952. Amussiopectell praesignis. SHUTO, p. 25.

Material.-GK-1 4370-72, 4424, 4427.
4430. 4678-79, 4681-86. All the specimens came from Toriyama, Kawaminami Mura, Koyu Gun*, :Miyazaki
Prefecture (Joe. no~ Ml-5595).
)'vieasu1·e1ire1zts.-Sli_own in the table 1.
Remm·lls.-The holotype (Yo_KoY ,, M A,
1922, pl.. 5, figs. 1-3) and other specimens from the Kakegawa group, fromwhich the holotype came, have thecharacteristic ribs which splits on themarginal part. but the present specimens have the ribs splitting ·not onlyin the adult stage but also in theadolescent. This splitting character ofthe ribs seems to be markedly variable,

though original author enumerated it
as one of the specific characters. The
specimens of the writer's collection
from the Tonohama group of Kochi
Prefecture are quite identical to theoriginal form, but thosa from Shimajirigroup of Ryukyu at the writer's disposal at the Department of Geology,
Kyushu University show the flat ribs
without showing any sign of splitting.
Another peculiarity is found in thewriter's collection. Though the interior
rib of this genus generally: has thepaired ridges on itself, one specimen
before the writer (GK-L 4685) has theinterior rib with triple ridges. However
the available materiai is not sufficientenough to clarifY the meaning of this]:ecularity. For the time being it isregarded as one of variations.
Horizon.-Upper horizon .of the Takac
m.be member (Lower Pliocene).
.) ~

...

Amussiopecfell from the Miyazaki Group, Miyazaki Prefecture, japan

Amussiopecten hyugaen.sis n. sp.
Pl. 16, figs. 1-5, Pl. 17. figs. 2, 4
and 5, text fig. 3

Material.-Holotype-GK-L 4368. Joe.
MI-3592: paratypes-GK-L 4150-{)3, 4155.
4159, 4172, 4364. 4418. 4674, 4676 and 4695.
Measurenzents.-Shown in the table 2.
Diagosis.-Shell medium to large insize, rather solid, compressed, orbicularin outline, somewhat broader than high,slightly inequivalve. Right valve a little
convex, slightly curved near the beak.
provided with radial ribs; exterior ribs
about 11, low, broad. flat-topped, straight.
becoming gradually lower towards
the ventral and lateral margin : interstices narrower than ribs themselves;interior ribs wide and flat at the begining, later becoming concave to haveprominent ridges on both sides, andinterstices becoming wider near theventral margin. Concentric lines ofgrowth weak but distinct. E:us subequal : anterior portion being somewhat
wavy at anterior border with acuteR~

Text fig. 3.

Amussiopeclen hyugaensis

n. sp. Lateral view and rib form incross section.
L. .... .left valve, R. ..... right valve

105

corner and with weak but distinct
byssal notch. Left valve planoconvex :ornamented with exterior and interiorribs and growth lines ; exterior ribs
rather round-topped, as wide as interstices ; interior ribs like those on theright valve. Ears equal. Hinge line
horizontal, about fourty percent of shelllength in the adult stage.
Comparisom.- The present species isclosely allied to A. praesignis (YmmYAMA), but the former has smaller andthicker shell than the latter and isornamented with less numerous ribs
which never split into riblets.
A. ? yabci NoMURA. which may be
included in A. praesigms, is also allied
to this new species, but the former has
more numerous ribs and large apicalangle than the latter.
Amussiopecteu
jlavellum
UGoLINI( Flavellipecleu pasiuii MR:-~F.GIII KI),
reported from Sardaique, closely resembles the young specimens of thepresent new species in the generaloutline and in size, but the former has
somewhat more numerous ribs, which
are narrower and slightly weaker thanthe latter .
.. Pecten (Vola)" sinkirensis MARTIN
has much more numerous ribs, which
are rather roof-shaped on the left valve.
Oopecten 1·otmtdatus (LAMARCK) is
similar to the present species, but isdistinguished by its larger size. moreconvex valves and its ribs are slightly
narrower than the interstices. Furthermore the internal ribs are not prominent in the former.
Variatio11. -As indicated in table 3,
this new species is remarkably Yariablein the size. depth of the valve, number
and strength of the ribs etc. The
specimens from the lowest horizon
attain to 70 mm, those from the middle
horizon are less than 50 mm, and those

106

Tsugio SIIUTO

from the highest horizon are 100 mm ormore in the length of the shell. It is
neccessary to determine whether or not
the larger specimens in each locality
are really the adult shells. For thediscrimination of this point the reliefof the ribs may be useful. The exteriorribs of Amussiopecteu are distinctly highin relief in young and become lower
and obscure as the shell grows. Tho
growth-stage of each individual is notwell indicate:! by size of the shell butby the relief of the ribs at the margin.
The ribs are generally almost equal
in relief at the shell-margin among thespecimens of equal size. regardless ofthe horizon of the specimens. ln otherwords in the present new species there
may be merely the difference in growthstage. Therefore the fact that the size
is dissimilar among different localities
or horizons may be due to such factors
as ecological and sedimentary environments, mode of preservation and orothers. although the number of theadult shells are not so numerous as toconfirm this interpretation.
Conceming the ecology of the following fact should be noted. Apart fromthe above mentioned difference in size,
some other characters also vary byhorizon. For instance the specimensfrom the basal part of the Tano andthe Boroishi member show the wide
variation in the number and the strengthof the ribs, the depth of therightvalve and the ratio of height
to width.
Those from the1upper part of these members
left( C!ementia-Paphia horizon) arerather invariable, the shell of about
50 mm in length, being provided
2
with 11 to 12 ribs of moderate relief
and with rather deep valves.
Furthermore the shells from thehighest horizon attain large size
3020
and have the tl1ick shell, tl1e deep
111111valves and the ribs of rather variable number. The lithology of tlle
4lower horizon is silty sandstone,coarse sandstone or granulebearing sandstone and remarkably
357
emvariable laterally and vertically,
Text fig. 4. Relation between the relief of theand that of the middle horizon
rib and the size of the shell. Relief is shown in
is rather monotonous massive
cross section. Hatched part indicates the shellsilty sandstone or sandy siltstone.
matter. Note that any essential difference canIn the upper horizon the lithology
not be observed among three forms.
is again coarse sandstone but is1-3, Amussiopecten h_vugaensis n. sp.
less variable. It has been well
-1, A. praesignis (YoKOY.HIA)
known
that many Pectinids are
Size of the shell is represented by thetheactiveswimmer and accorddistance frorn the beak (scale below).
ingly
necessitatemuch oxigen.
Relief of the rib is mea-,urable by the~ca\e above.
Thereupon they favour the coarse

I il
I I l \

'
I I !

I

I i

\

\

I

Amussiopecte11 from the Miyazaki Group, Miyazaki Prefecture, Japan

sandy, gravelly or rocky bottom,
where water is generally agitatedand contains plenty free oxigen. If
Amussiopecten has similar habit, it isreasonably explained that the large andsolid specimens occur in the coarse
sediments and the small specimens in
the muddy sediments.
Much moreevidence is required to settle the quesTable 2.
Register
I!GK-L-.
,Number

1
1

107

tion.
The variations in the importantcharacters range as follows.
1) The ratio of height to width ofthe valve varies between 89 and 102
percent and most frequently between
96 and 99 percent.
2) The ratio of depth of the valve
to width varies between 11 and 19 per-l\'ieasurements on the selected specimens of Anwssiopectem hyugae11sis n. sp.
.cl~oo'e :I:"'
t:::_,., I >-l,., I.Jv~~-~!
.~-& ~...
, Cl"" i!!'a Is::>:>:: j o. s:: to~~

QJ,........rLocallity

s::

~ E

MI-

I_
4150
1620 Yusunokihashi 55.4
54.6 12.4
4151
903 Kariyabaru
3.356.0± 58.4
4152
8118 Inohaye
47.5± 45.5±1 3.5
4153
8118 lnohaye
39.7:19.0 1 7.4±4154
8118 Inohaye
20.0±1' 19. 0± 3.4±
4155•
903 Kariyabaru
50.5± 47.5
33.7 . 33.5
4156 I 2470 Maruno
4. 7
4157
3189 Obira
36.0± 33.0± 4.1
4159
3189 Obira
43.0
41. 0±1
ao. 3 3.2
4161
826 Horiguchi
4162
826 Horiguchi
24.0± 23.5
4164
157 Kano
20.5
20.0± 3.3
4168
2698 lyeichigo
17.4
17.0±12.0
4169
3121 Hokobo
18.5± 17.0±
4170
2610 Kagamisu
22.3
21.4
3.2
4176
743 Yanaze
74.0±
4177
1615 Yusunokihashi 85.0±
4178
437 .Mokudo ·44.0±
437 .Mokudo
4180. 56:0± 54.0±
2610 Kagamisu
4181
32.2
28.6
525 Morinaga
4182(i0.0±
4184
1620 Yusunokihashi 27. 8
27.3 '
839 Horiguchi
27.4
4185
24. 8 I •1. 8
4186
161 Akatani
28. 4
25.6
33. 1
4187
3121 Hokobo
4189
903 Kariyabaru
23. 4
23. 7
4191
903 Kariyabaru
24. 5
22. 0±
4192
903 Kariyabaru
22.2
22. 0±
4194
903 Kariyabaru
23. 2
22. 4
1. 9
19.6
19. 1
4195
3121 Hokobo
4349
1334 Haigano
14.2
1:~. 9· 43511334 Haigano
14.3
13.9
4364
3592 Oyamada
30. 9
30. 1
4. 2
4365
3592 Oyamada
42. 3
41. 3
7. 9
4366
3592 Oyamada
39.7
2.1
4368
3592 Oyamada
103. 0± 101.7 19. O**·
4369
3592 Oyamada
100. 0 ± 99. 0 ±4"421
3121 Hokobo
19.4
19.3
4671
3592 Oyamarla
30. 1
27. 7
3. 9
58.0 ± 55. 4
5. 4
4672
3592 Oyamada
35. 1
32. 3
6. 9
4675
3592 Oyamada
4676
3592 O~oamada
24. 4 . 22. 93. 9108. 0± 107. 0±46773592 Oyamada. . . .I~

~ ........

......- I

'-'

.
~

.1"""".,.. .z'Ol

1

1- - - - - - - - -----"--* conjoined and deformed.

1~ ~ OJ-o'-'---·--~----

22 ± 98
2<1 ± 102
24. 4
96
98
95
94
99
15.2
92
96

4
6

40
42
51

7

19
17
I

98
97
98
92
96

11

11
10
11

11
11
I 14
42 13
12
10

14
11

.

1

(UQJCIJ·~ '-'· :::

11

16
12

115
103
117
117
108
118
115
115
116
115±
111
112
112
108
111

12
10
11
14
11
14 I
129
13
14
ll5
96
113
10
89
11
! 15
98
' 11 I 114
91
17 I
14 1 115
90 I
12
112
17 .4 I
52 11
117
101
11
112
90
13
100
12
13 .!J
96
60 11
8
Ill
97
12
107
98
11
113
I
10! 10897
16.7 I 97
13 \54 13
117
19
97
11
117
I
I
19.3
11
119
48 .0± !)9122
18**1 47 11
99
11
127
}')
99
114
13 . 1
13
98
110
12
24 .5
96
9
42 11
118
I
I 12 115
92
19 I
12 .4
94
Ill
16 1 51 i 11
99
11
125

1

----'--~* thickness of conjoined

I

14·1

I

valves.

i

1

~I

>i
-L
L
L

I

R
R
L

R
R

L
R
R

R
R

L
R·
R:

R
R
R

L
L
R
R

R
L
L
L
L
I.
L
L
L
R
R

L
L
L
L
R

L
R
R

L

I

Tsugio SIIUTO

cent, and most frequently between 12
and 17 percent in the right valve; between 4 and 8 percent in the lett.
3) The ratio of hinge-line to shelllength varies between 40 and 50 percent
in the adolescent and adult shells, but
the young specimens show larger value,
for example. it is 60 percent in a specimen whose shell-length is 23.2 mm.
4) Apical angle varies between 103
and 129 degrees and most frequently
between 110 and 117 degrees in theadolescent shells and between 122 and
127 in the adult.
5) The number of the exterior ribs
varies between 10 and 15, and mostfrequently between 11 and 12.
llorizmz.-Lower part of the Miyazaki group (late Middle Miocene to
middle Upper Miocene).
Locality.-Waritsuke and Mokudo,
Aya Machi*; Morinaga, Honjo Machi**;Yanaze, Akatani and Kano, Takaoka
Machi*u; Yusunokihashi and Oyamada,
Mukasa Mura**~·*; Kariyabaru, Haigano,
Hokobo. Horiguchi and Obira, Tano
Machi.*****; lyeichigo, Maruno andKagamisu, Miyazaki City******; Inohaye, Kitago Mura*******.
Works Cited
DEPF.RF.T, Ch. et ROMAN, F. (1905), Monographic des Pectinides Neogenes de!'Europe et des Regions Voisines-ll

Flavellipeclm.

Afem. Soc. Ceo/. Fra11ce
13, Fas. 2, pp. 105-136.

Paleot~lologie Tm.

- - et - - (1910), Monographic des Pectinides Neogenes de J'Europe et des Regions
Voisines-Il Flavellipecten. ibid. Tm. 18,
Fas. 2, pp. 137-139. pis. 8-13.
- - et (1912), 1\Ionographie des Pectinides Neogenes de I'Europe et des Regi----------

ons Voisines-II Flave/lipectell. ibid. Tm.
19, Fas. 1, pp. 139-168, pis. 1-6.
MARTIN, K. (1910), Die Fossilien von Java.
Bd. 1, Abt. 2. Heft. 1, Moll. pp. 333-356,
Taf. 46-50.
MAKIYAMA, Jiro (1927), Mollus:an Fauna of
the Lower Part of the Kakegnwa Series
in the Provjnce of Totomi. Mem. Col/.
Sci. Ryoto. Imp. U11iv. Ser. B, Vol. 3. No.
1, Art. 1, pp. 1-147. pis. 1-6.
- - (1931 ), Stratigraphy of the Kakegawa
Pliocene in Totomi. ibid. Vol. 7, No. 1,
Art. 1, pp. 1-55, pis. 1-2.
(1932). Wf~:=_tc ~i'!U~~ Neogene Tertiary. lwanami-Koza (Geology and Paleontology) pp. 1-57, pis. 1-2.
NOMUHA, Shichihei (1933), Catalogue of the
Tertiary and Quntery Mollusca of Taiwan. Sci. Rep. Tohokr1 Imp. Univ. 2ndSer., Vol. 16, pp. 1-108, pis. 1-4.
- - (1937), The Molluscan Fauna from the
Pliocene of Tosa. ]apa11. four. Ceo/. Ceogr. Vol. 14, No. 3-4, pp. 67-90, pl. 6.
- - and NIINo, Hiroshi. (1932), Fos3il Mollusca from Izu and Ilakone. Sci. Rep. To-

hoku Imp. U11iv. 2nd Scr., Vol. 15, No.3,
pp. 169-192, pis. 11-12.
- - and ZINno, Noboru (1936), Molluscan
Fossils from the Shimajiri Beds of Okinawazima. ibid. Vol. 18, No. 3, pp. 229265, pl. 11.
OTUKA, Yanosuke (1930), ~ltf!Ji!;~mPIJ~ftili~
O~a~r.lJM Geologic Problems in the
Takanabe District, Miyazaki Preft>cture(in Japanese). Ceogr. Review, Vol. 6, No.
7, pp. 1048-1074.
(1934), ~~~fi(O)f~:fi H!'im.ljlJIJI{ij&'(fl~Jl1t:Ei On Some Fossils from the Northern Foot of Mount Minobu and from the
Hayakawa Tuffite of llakone (in Japanese). ]our. Gcol. Soc. Tokyo. Vol. 51, No.
492, pp. 562-568.
- - (1938), Neogene Fossils of the lhara
District, Sizuoka Prefecture, Japan. jour.
Fac. Sci. Imp. U11iv. Tokyo Ser. 2, Vol.
5, pt. 1, pp. 1-19, pis. 1-2.
PHILIPPI, H. E. (1898), Beitriige zur MorphoJogie und Phylogenie der Lameliibranchier.
Zeitsch. Deut. Ceo/. Geselsch. Bd. 50, pp.
597-622.
- - (1900). Beitriige zur Morphologic unci

Amussiopecte11 from the Miyazaki Group, 1\liyazaki Prefecture, Japan
Phylogenie der Lamellibranchier. Bd. 52,
pp. 64-117.
RoGgR, Jean (1939), Le Genre Ch/amys dans
Ies Formations Neogenes de !'Europe.

Mem. Soc. Geo/. Fra11ce. Nouvelle Ser. N
40, pp. 1-294, pls. 1-28.
SHUTO, Tsugio (1952), ~-~"ifrFI?)i!l!.~.I.YF:l'l':Jlilf~: Stratigraphical Study of the 1\liya?..aki Group (in japanese). Sci. Rep. Fac.
Sci. Kyuslm U11iv. Geol. Vol ..J, No. 1, pp.
1-40.
YoKOYAMA, 1\Iatajiro (1922). On a New Species of Pecten from the Neogene of Japan.

109

jour. Geol. Soc. Tokyo. Vol. 29, No. 530,
pp.1-2, pl. 5.
- - (1926), Tertiary Mollusca from the Southern Totomi. jour. Fac. Sci. Imp. U11iv.
Tokyo Ser. 2. Vol. 1, pt. 9, pp. 313-364,
pls. 38-41.
- - (1928), Plio:ene Shells from Hyuga. ibid.
Vol. 2, pt. 7, pp. 331-350, pls. 66-67.
- - (1029-a), Mollusca from the Oil-Field of
the Island of Taiwan. Imp. Geol. Surv.
japan, RejJ. No. 101, pp. 1-112, pis. 1-18.
- - (1929-b). I'lioccne Shells from Tonohama,Tosa. ibid. No 104, pp. 9-17, pis. 7-8.

110Tsugio SHIJTO

Explanation of Plate 16
Figs. 1·5. ( x 1), Amussiopecte11 hyugaensis n. sp.
1, right valve, paratype, GK-L 4163, Joe. l\ll-157.
2, left valve, paratype, GK-L 4696, Joe. Ml-903.
3. left valve, holotype, GK-L 4368, loc. Ml-3592.
4, left valve, paratype, GK-L 4151, Joe. Ml-903.
5, left valve, paratype, GK-L 4152, Joe. Ivll-1620.

Explanation of Plate 17
Fgis. 1 and 3. Amussiopecten praesigtzis (YOKOYAMA)
I, (x 1), right valve, GK-L 4682, Joe. Ml-5595.
3, ( x 5/6), left valve, GK-L 4371, Joe. MI-5595.
Figs. 2, 4 and 5. ( x 1), Amussiopcclen hyugaCIIsis n. sp.
2, right valve, paratype, GK-L 4364, Joe. MI-3592.
4, left valve, GK-L 4152, loc. l\U-8118.
5, left valve, inner cast, GK-L 4159, loc. Ml-2198.

T. SHUTo: Amussiopecten

Photos by T. SHUTo

PLATE 16

T. SHUTe: Amussiopecten

Phot:os by T. SHuTo

PLATE

17

Trans. Proc. Palaeont. Soc. japan, N. S. No. 20, pp. 111-118, Pl. 18, Dec. 30, 1955

277.

LES AMMONITES RECUEILLIES DANS LE GROUPE DE
KURUMA, NORD DU JAPON CENTRAL*TADASHI SATO
L'Institut de Geologie, Universite de Tokio*·~~f,f.t IJ)$ftl Lt..:~:fi;:.'0~··C:*·~I\'!1JI¥0)11lfft(tll{jj:j!I:.~.\?G;h"C~·t.:il•"':>f..:il;, j;:.If!JII
3!tliftO)=lf.ft-il· G Amaltlzeus sp. indet., Aff. Deroceras sp. indt•t., Ca11amria sp. ex gr. geyeria11a(HAAs) a-rlliHl Lf..:O)·~. -$(t Domerian -c:~.;,.:. til;ft"?f..:.og: iii IE

Ia Prefecture de Toyama, au Nord duJapon central. Les Ammonites ont ete
recoltees dans une a~sise assez mince, etsont rcmarquablement concentrees.
Toute b gratitude de !'auteur va en
particulierl\I. le Professeur T. KoBA·
YASIII qui, le premier, a fait lui connaitre
le gisement cl'Ammonites dans cetteregion et le surveille pendant son etude.
L'auteur remercie M. le Docteur P. L.
MAuBF.UCE qui l'a donne beaucoup decritiques et a corrige son manuscrit, etaussi M. le Professeur T. MATSUMOTo,
Universite de Kyushu. qui l'a enseigne
des taita sur le Groupe de Toyora. MM.
K. KoNISHI, M. TAMUJ{,\ et G. MoRI ont
n~colte beaucoup de sp(·cirnens dans Ia
vallee de Teradani avec !'auteur,qui
ses remerciernents vont particulierement.

Introduction

La serie de Kuruma s'etcnd dans la
Nord partie du Japon central, notamrnent
sur Ia region frontiere des Prefecturesde Niigata ct de Toyama. mais la succe~sion des couches de cette si·rie etait
peine etablie, puisque lcs fossiles
indicateurs et les assises caracteristiques
n'y ont pas ete decouverts. Toutefois,
plusieurs auteurs ont explore cette region
et n~colte beaucoup de fossiles paraliques
et neritique::, qui sont tres abondamment
concentres dans quelques horizons. rnais
ne sont pas i-tudies en detail. l.es fossiles vegetaux ont aussi ete trouves;!'age rhetien est indique par cette ftore.
Cependant, l'on ne pom·ait pas exactement preciser I' age de cette serie d'apres
ces seuls fossiles.
Dans une periode
ancienne. quelques geologues ont trouve
Ia riviere de Daira des Ammonites qui
sont en tout cas incompletes et non
identifiable generiquement.
En 1952-54, beaucoup de specimensd'Ammonites ont ete trouves dans Ia
vallee de Teradani, une branche de Ia
Daira, qui est situee a Ia partie Est de

a

a

a

Geologie en sommairea

Le Groupe de Kuruma est compose decouches tres epaisses, contenant plusieurs
assises de gres, d'argiles et de conglomerats, clont l'epaisse:.Ir est d'environs
6.000-7.000 metres. A Ia partie presquemediane de cette serie se trouvent les
couches marines, d'ott les Ammonites
proviennent. Elles sont plus ou moins
minces, composees d'argiles massives et* Read Oct. 29, 1955, received July 23, 1955.
111

112

Tadashi SATO

de gres aussi massifs, riche en fossiles
marins.
L'assise dans laquelle Ies Ammonites
et les Bivalves marins ont cte trouves,affieure dans Ia vallee de Teradani, ou
on constate trois zones d'Ammonites ou
plus, et aussi dans la vallie de Kanayamadani.
Cette assis:! s'appele la"Formation de Teradani ". !:extension
de cette assise est reconnu dans Ia valleed'Itsumba vers l'Ouest, et vers l'Est dansla vallee de Kanayamaclani, Prefecturede Niigata, mais il est impossible dereconnaitre sa limite Ouest.
Les Ammonites, differentes de celles
de la Formation de Teradani. ont aussi
ete trouvees dans Ia vallee d'Otakidani,mais elles sont trop mal conservees pourune identification.
Age

La faune de Teradani est composeedes Ammonites des genres: Amaltlteus,
Deroceras, Cauavaria et P/Jy/loceras etde Pelecypodes marins. Amaltheus setrouve dans le Domerien nord-europeen,
et notre espece est tres proche d'une
forme d'Amalt/u:us, qui est connu au
Lias delta inferieur, c'est-a-dire Ia zone
a depressus, dans l'Allemagne S-E et au
Pliensbachien superieur de I'Angleterre.
Done, l'age de cette assise semble unpeu plus ancien que le Dorni·rien. Amaltheus se cantonne dans Ie Lias delta
des regions europcennes.
Deroceras est malheureusement recueilli en debris. dont Ia roche est peutetre originaire d'une zone independante.
II se trouve dans !'Europe au Pliensbachien inferieur. Canavaricz est tresabondamment recueilli dans cette localite, et joue un role tres important dansla cotTelation stratigraphique: il setrouve . dans Ie Domerien de la region
mediterraneenne. Done, Ia Formationade Teradani, une pattie de Ia serie deKuruma, ont cte deposees pendant le
P!iensbachien (superieur ?).
Descrit>tion des Especes
Canavaria fit>. ex gr. geyeriana (HAAs)
1913.

1931.

Harpoccras (Arieticeras) retrorsicosta
OrPEL var. gcyerimza, 1-IAAs, Lias vo11
Ballino, p. 6:!, Tav. 11. Figs. 10, 12 non
11.
Cawwaria gcycriaua, FUCINI, Foss.
domer. di11tomi Taormi11a, p. 154, Pl. 16,
Figs. 11-13.

Diagnose :-Coquille discoide, sa section est quadrangulaire chez Ia spirejeune, puis rnoins epaisse chez l'adulte(stade d'environ 10 mm de la hauteur
du tour). devenant plus elliptique. Tours
nombreux, peu embrassants Cl/5 du tour
precedant), Ia croissance du tour tm peu
rapide. Chute du tour sur l'ombilic
inclinee et convexe ; Ie bord ombilicalest arroncli mais !'aspect de cette region
est peine reconnue chez les jeunes denotre collection.
L'ornementation varie avec l'acroissement de Ia spire. Jusqu'a la hauteur
d'environ 2 mm de tour. le ventre est
orne de la carene arrondie et obtuse.
Au Stade de 2 mm
7 mm environ de
Ia hauteur de tour. Ia carene ventrale
haute, epaisse, mais arrondie est bordeede sillons assez profonds et tres etroits.
Les c{)tes sur lc ftanc sont droites,
elevees, et arrondies, se terminant aux
tubercules qui sont petits et absents au
stade adulte, ou detormes sur quelquesechantillons. De ces tubercules, naissent
les ci>tes accessoires qui sont tres faibles,
dirigees en avant, sont inclinees de fa~on
abrupte, et quelquefois une sorte defaible carene pres de sillons. Ce Stadese continue en principe au diametre de
5-6 mm, mais c'est plus ou moins variable. Dans !'autre cas, Je sillon n'est pasaaI.es Ammonites recueillies dans Ie Groupe de Kuruma, Nord du japon central

ne, mais Ia carcne se distingue nettement de Ia region ventrale. Au Stadesuivant. qui est de 7-15 mm environ de
Ia hauteur de tour, le sillon a disparu,mais Ia carene reste arrondie et epaisse,
depassant nettement de Ia region ventrale ; mais dans quelques cas. Ia carime
est encore bordee du sillon rudimentaire.
Les ct'ltes sont invariablement droites.
terminees aux petits tubercules qui sootdisparus au stade adulte. Section dutour plus comprimee et elliptique. Le
ventre semble generalement arrondi.
Quand les tours dcviennent plus grands,le ventre est completement arrondi.
Mais il est impossible de constater lecaractere adulte, parce que !'on ne peut
pas trouver un bel exemplaire dans notre
collection.
Bouche inconnue.
Ligne
suturale inconnue.

113

que cette variation n'est pas visible sur
plusieurs des atitres echantillons. elle nesemble peut-etre pas reprcsenler une
variation.
AJ}inite et Comparaison :-Cette forme
provenant abondamment de !a Formation de Teradani est aparemment prochede Cmuwaria GF.MMEr.r.AR<>, jugee parson ornementation de Ia spire. La
carene assez obtuse mais haute, les deux
sillons pres de celle-d sur Ia region
ventrale, qui soot peu profonds maisbien distincts de Ia region ventrale, lestul::ercules de chaque cl>te sur Ia region
periphero-laterale, les inflexions descotes vers !'avant sur Ia region peripheroIaterale : tout cela indique les caracteresde Cauavaria. Bien que les cloisons nesoient pas reconnues, !'auteur n'hesite
pas Ia rapporter a Cauavaria.

Dimensions :-en mm.
Num{,rosT5303-12
T5303-36
1'5303-,15
T5303-46
1'5303-79

Diametre
237
68,3
ca 75
62,5
36,0

Diametrede J'ombilic
10.5
30,5
31,5
27,5
17,0

Observation : -L'ornementation de Ia
coquille change d'autant plus que lestade d'acroissernent s'avance. En particulier, les tubercules ont disparu a un
stade remarquablement different, avec
celui !'absence du sillon. Les tubercules
ne soot pas visibles sur toutes Ies formesde plus 20 mm de hauteur du tour, mais
il y a un groupe ott ils manquent au
stade moins haut. Au tour haut de 78 mm, les tubercules peuvent etre nettement reconnus ; le stade de disparition
des tubercules est tardif (a environ 10mm de Ia hauteur). Sur les tours,
T5303-37, T5303-45, T5ao3-54. le ventre
est aigu. depourve de carcnc. Sur une
spire d'un specimen (T5303-45) les cbtes
sont plus serrees et plus faibles. Parce

Hauteur
8,2
21.5
25,0
20,5
12,0

Epaisseur
?

6,5
6,5
10,3

Les deux spires de tubercules ont
donne naissance sur Ia partie Ia plus
interne et sur Ia partie periplH~rique descotes chez les plusieurs especes deCanavaria ; mais il y a un groupe
pourvu d'une ligne de tubercules, (C.
geyeriana represente ce groupe), auquelnotre forme peut etre rapportee.
Dans la region mediterraneenne il ya plusieurs genres tres semblable a cegenre : Emaciaticeras en est surtout leplus proche. II est orne des ci'Jtes
similaires, contenant des esp£.-ces que!-·
quefois pourvues de tubercules petits
sur Ia partie peripherique au st.:'lde jeune.
qui tend a disparaitre pluti>t. Mais
FuciNI ne precisa pas les distinctions
pour .Ia creation de ce genre et ·laTadashi SATO

114

naissance de tubercules comme caractere
generique n'est pas non plus donnee;
mais cela suggere l'affinite tres intime
de deux genres. Seguentia. indus parmi
Arieticeras par les auteurs anciens, semble proche de cette espece de Teradani.
Mais il est aisement distinguable par Ia
reunion des cotes sm le ftanc interne,
par Ia. disparition de l'ornementation sur
la partie interieure du ftanc. Arieticems,
un representant de ce groupe d'Harpoceratide, en differe par les details suivants : les d>tes ftexeuses, Ia carcne tres
haute et mince et la disposition plus
espacee des cotes sm le llanc.
Position stratigraphique:- C. geycriana
a recueilli au Domerien de Kratzalpe etde Taormina. Mais les formes douteusessont citees a Medolo, Brianza, Spezia etdans l'Appenin central dans le Lias
moyen d'apres HAAs. Mais dans cetteregion, les zones d'Ammonites n'ont pas
ete precisees et !'on ne peut pas reconnaitre le niveau correct de cette forme:
par consequent, !'auteur veut le rapportera rage domerien ou au Lias tnoyen.
Aff. Deroceras sp. indct.

Diagnose :-Coquille discoide, peu embrassante ; spire croissant plus ou moins
lentement, composee de 4 tours.
Iasection quadrangulaire et convergente.
au ftanc trcS etroit e(; un pelt renfte, ~1]a region ventrale tres etendue et arrondie, depourvue de carene. Ombilic
tres ouvert, large : chute du tour sur
I'ombilic tres inclinee et son bord arrondi.
2 spires les plus internes liEses etanondies. Troisicme tour orne des c6tes
sur le !lane tres obtuses et assez distantes, qui ont donne naissance au bordombilical et deviennent plus saillantes
contre Ia region yentrale. et pourvues
de tubercules tres epineux, qui tendentaa prolonger dans Ia direction du rayon.
Ces tubercules sont disposes de Ia meme
maniere sur deux tlancs, et Ia region
ventrale est ornee des ct>tes plus fines
et plus nombreuses que celle sur le ftanc.
Cettes cotes obtuses et fines sont disposees asymetriquement de telle sorteque deux epines disposes sur Ia region
peripherale sont retmies par les diverses
c6tes. Ligne suturale inconnue.
Dimension :Diametre, 15,7 mm; Diametre de l'ombilic,
7,9 mm; Hauteur, 4,3 mm; Epaisseur, 4,6 mm

Affinite :-Nous avons un seul echantillon. II semble qu'il appartient
Deroceras. bien qu'il ne presente qu'unstade tres jetme. II est tres proche de
Ia serie de Btfericeras.Deroceras. mais!'auteur ne pcut pas le rapporter siirement Bzfcricems, p:irce que la designation originelle ne cite pas l'ornementation ventrale de Bifericeras. Jugee
sur les quelques figures citees, elle est
formee des cfltes inclivisibles, au contraire
du groupe de ptauicosta qui a l'ornementation des c6tes bifurquees sur la region
ventrale: done, on peut Ie rapporter plus
aisement cornmc un membre d'une
brancl:c de Deroceras. On ne pent pas
decider son espece puisqu'il manque lestours adultes. Mais il faut remarquer
les deux ou trois tours qui sont tous
l'isses et arrondis. II faut aussi noter
Ia ressemblance pour le stade jeune
ci'Arietites coregonensis WAH;>;ER, maiscelui-ci a les tubercules allonges radialement; en particulier, il y a une tendance~l l'abaissement des cotes vera la suture
ombilicale chez ce specimen recueilli
Teradani. Parce qu'il rnanque les tours
adultes, il est preferable de ne pas
conclure sur sa position systematique.

a

a

a

Amaltlleus sp. indet. ·Diagnose :-Coquille discoide, pluS. ou

Les Ammonites recueillies dans le Groupe de Kuruma, Nord du japon centralmoins plate, assez profondement embrassante. Tours section elliptique, mais
tres plate, avec rt~paisseur maxima sur
Ie milieu du flanc. Ombilic tres etroit
et bas. Chute du tour sur J'ombilic
inclinee chez Ia spire jeune, mais verticale chez J'adulte. Ventre plus ou moins
aigu. et pourvu de Ia carene.
C<>tes tres obtuses mais assez eievees
sur le ftanc, disposees radialement, lesplus elevees pres du bord ombilical, puiss'abaissant sur Ia region ventro-laterale,
pour se diriger en avant. Far consequent, la region ventro-laterale est
presque Iisse, surtout sur Ia spire assez
agee. Les Ctltes, moins elevees et plus
nombreuses que celles sur le ftanc,passent sur Ie ventre aminci, sans aucuneinterruption et sans aucun faiblissement
au milieu du ventre, en formant Iacar(me creneJ(·e qui n'est pas distinguee
nettement de Ia region ventro-laterale.
Quelques cotes continuent de Ia suture
ombilicale Ia region ventrale.
Ligne suturale profondement decoupee.
Lobe ventral bifide et pourvu de deux
lobules latt~raux. Selle ventrale n·es
haute, vide, etroite a Ia base, et divisee
par les trois lobules. Lobe lateral superieur plus profond que le ventral. plus
ou moins vide, et profondement bifurque:Ia branche interne est toujours plus
longue et plus nettement decoupee quel'exteme. Lobes et selles lateraux inferieurs et auxiliaires sont tous petits
et, graduellement, decroissent contre lebord ombilical.
_ Observation :-Nous avons 6 ecbantillons trouves dans Ia meme localite, mais
trois sont si mal conserves qu'ils nepouvent pas ctre utilises pour une identi-aaDimcnsimzs :-en mm.
Numeros
T5303-l1'5303-laDiametre
35,3
43,0

115

fication. Tous nos echantillons sontplus ou moins detormes par Ia pression
de Ia roche d'oi1 ils proviennent (excepte
un exemplaire) et sont ainsi aplatis.
Sur le moule interne de la spire duspecimen typique reste un fragment dela coquille, qui est orne de stries fines
dirigees radialement. Ce caractcre est
reconnu sur le moule interne d'autres
echantillons. Sur le specimen typiqueles cotes ne continuent pas de Ia region
ombilicale
la region ventrale, mais
elles le font souvent sur les autres
echantillons. II apparait que I'on peutreconnaitre une carene vraisemblable
sur quelques parties de Ia spire. Maisl'absence de la car(me vraie est prouvee
par l'ornementation de Ia region ventralechez les stades adultes du specimen
typique. Ce caractere est visible chez
quelques exemplaires d'Amaltlzeus, d'apres WRIGHT et FRI>NTZEN.
ComParaiso1l et Affinite :-Cetle espece
apparait Ctre tres proche d' Amalthcus
depressus. La disposition des ctltes etl'involubilite de la spire de cette espece
suggere l'affinite du stade compressus
d'apres FREKTZEN (tandis que le genre
Amalthcus par FRENTZEN comprend Ies
especes qui appartiennent obligatoirement
Paltoplcuroceras). mais on nepeut pas negliger des caracteres assez
importants qui different entre les deux
formes. Quelques-uns des exemplaires
cites dans les planches de FRENTZEN,
1937, Taf. I, Fig. 28; Taf. II, Fig. 17,
presentent le meme aspect de Ia region
ventrale, tel que la carene ne se distingue pas de la region ventrale ; tandis
qu'elle n'existe qu'a quelques stades deIa croissance, I'on ne peut p: s reconnaitre un beau dcveloppernent de Ia

Diametrede J'ombilic
9,5
10,5

a

a

Hauteur
15,6
20,5

Epaisseurca 6,5?

116

Tadashi SATO

carene chez quelques-uns de nos echantillons. Ainsi, I'aspect de Ia merne
manicre est trouve sur Amaltlzeus
clevela11dicus (figure par BucKMA!';;
Type Ammonites, Pl. CIX, Figs. 1, 2)
rnais il differe de notre forme tres
nettement par l'ornementation beaucoup
plus forte sur Ie ftanc. Ce caracteres'accompagne de Ia disparition des ornements sur le ftanc chez les specimens
de l'Allemagne SO, au stade d'engclhardti d'apres FRENTZEN, Taf. II, Fig.
19 ; en outi'e, notre specimen montre
que les ornements sur le ftanc tendent
disparaitre avec plus de retard que 1acarene.
De plus. WRIGHT a repn~Eente la forme
ornee de la nH~me maniere sur Ia region
ventral e. Taf. LVI. Fig. 15, sans donnerIa vue peripherique. Cependant, !'auteur
ne peut pas reconnaitre les spires internes et les tubercules generalement apparus chez Ie stade jeune. II semble
que les formes de notre collection. cellesde FRENTZEN et de WRIGHT, tandis que
proche au stade comparable, se differencient assez distinctement. La ligne
suturale est moins decoupee que lesformes citees par FRENTZEN et m1tres.
Le lobe lateral superieur n'y est pas
bifurque, et le tronc en est assez profondet vide; Ie lobe lateral inferieur y est
plus petit que celui de cette espece.
Mais son arrangement est fait en principede la meme maniere que dcpressus, etle degre d'incision est tres re3semblant.
L'auteur veut decrire cette espece comme
une affinite d'Amaltlieus dcpressus.
BucKMAN a decrit quelques especes
d'Amaltlzeus d:ms "Type Ammonites";
Amaltheus sedwickii (Pl. CXXV) et A.
depress us (PI XXV, Figs. l. 2) sont tres
ressemblants a notre espece. 1\Iais l'on
ne peut pas constater les cloisons etl'aspect ventral de celui-la, et les cloisonsde celui-ci; done, !'auteur ne pouvait pasaexactement les rapporter a notre forme,
tandis qu'ils semblent tres proche denotre forme en general.
II est necessaire de remarquer que
Proamaltlzeus etabli par Lange en 1932
est ressemblant a cette espece, mais il
en differe par ]'absence de cotes aussi
fortes que celle de notre espece, et parla presence des fines stries sur Ia region
ventrale.
Cette espece a une ressemblance plus
ou moins nette avec quelques formes
du genre Acgoceras, qui a la section
generalement arrondie. Par exemple,
Discamphiceras est ressemblant a notre
espece (VVXIINER, Taf. XXV, Figs. 6a, b,c); mais les cotes ventrales tendent a
s'y abaisser au milieu de la region
ventrale. Cette tendance represente sonallinite avec Schlotheimia. Et sa ligne
suturale est tres differente, qui a 4 lobes
et selles laterales inferieures et auxiliaires; et elles decroissent beaucoup plus
rapidement contre Ie bord ombilical.
Position stratigraphique :-Amaltlteus
n'est trouve qu'au Lias moyen, notamment Lias delta. dans Ia region europeenne et mediterraneenne. Notre forme
semble aussi domerienne. A Nagato, M.
le Professeur MATSUMoTo a recueilli un
specimen ressernblant a notre collection,
avec association d'un individu ressemblant notre Canavaria (Communication
personelle). Ils sont tres fragmentaires,
tels que la determination est impossible.
Si les formes sont aussi Canavaria et
Amaltheus, !'age de la Formation de
Higashinagano de laquelle ils proviennent, est problematique, puisque une
assise susajacente est deterrninee commedu Domerien par M. Ie Professeur MATSUl\toTo.

aOutre cettes especes. quelques exemplaires tres fragmentaires et mal conserves, qui sont vraisemblablement

Les Ammonites recueillies dans le Groupe de Kuruma, Nord du Japon centralproche des Phylloceratides et des Lytoceratides, ont ete recueillis dans la mcme
localite de Teradani. Ces exemplaires
seront decrits apres la recolte de meilleurs specimens.
Bibliographie
BUCKMAN, S. S. (1909-30), Type Ammonites
DACQUE, E. (1933), Leitfossilien Wirbellosen
des Jura
DoNOVAN, D. T. (1954), Synoptic Supplement
to WIHGHT's ".Monograph on the Lias
Ammonites of the British Islands", Palaeonl.
So,;., London
FRENTZEN, K. (1934), Der Lias delta (Amaltheen-Schichten) im Gebiete zwischen Aselfingen und Aalen, Sitz. Heidelb. Akad. lViss,
Math.-Nat. Ktasse, 2 Abh.
- - (1937), Ontogenie, Phylogenie und Systematik der Amaltheen des Lias delta
Siidwestdeutschlands, Abh. Heide/b. Akad.
Wiss., Math.-Nat. K/asse, 23 Abh.
FuciNI, A. (1903), Cefalopodi liassice dellVlonte Cetona, Pt. III- V, Pa/eont. Ita/., Vol.

IX
- - (1929-31), Fossili domeriani dei dintorni

117

di Taormina, pts. 3·4. Ibid. Vols. 29-30
HAAS, 0. (1913). Die Fauna des mittleren Liasvon Ballino in Siidtirol. pt. 2, Beitr. Pal.

Oesterr. Ung. Orients, Vol. 26- - (1951), Some Comments on VECCHIA,"Su Alcuni Nomi Generici di Ammoniti
Liassici ", ]oum. Paleont.. Vol. 25, No. 1
LANGE, W. (1932), Ueber ein Hammatoceras
und einen Amaltheenvorlaufer (Proamaltheus
wertheri n. gen .. n. sp.) aus dem Lias delta
und gamma von 'Werther in Westfalen,
Zeitsch. deut. geol. Ges.. 84
1VloNE5TIER, ]. (1928). Recherches sur le
Polymorphisme et Ia Phylogenie des AmaJ.
theides domeriens d'aprc~ les materiaux
aveyronnais, ltfem. Soc. Lett. Sc. At·ls
d'Aveyron. 22 (L'auteur ne pouvait pas
trouver.)
QURNSTEDT, F. A. (1883-87), Die Ammoniten
des Schwiibischen Jura, Bd. I-ll
WAHNRR, 0. (1894), Reitrage zur Kenntnis
der tieferen Zonen des unteren Lias in dem
nordosterr!:'ichen Alpen, VI-Vll Teiles, Beitr.

Ceo/. Pa/ti'ont. Oesterr. lfng. Orients, Vols.
8-9
WRIGHT, T. (1878-86), Monograph on theLias Ammonites of the British Islands,
Palaeont. Soc .. London

118

Tadashi SATO

Explication de Ia Planche 18
Amaltheus sp. indet. ....................................................................................................... p. 114
Fig. 1a. T5303-1. vue laterale, Loc. 5803, x 1
Fig. I b. Vue peripherique du meme specimen que Ia, x 1
Fig. 2. T5303-1a, vue laterale, Loc. 5303, x 1
Aft. Derocerns sp. indet .................................................................................................. p. 114
Fig. Sa. Vue Jaterale. Teradani (dans Je debris), x 2
Fig. 3b. Vue periplu!rique du me me specimen (jlle 3a, X 2
Crmavaria sp. ex gr. geyeriaua (HAAs)........................ . ................................................ p. 122
Fig. 4. T530a-46, vue laterale, Loc. 5303, x I
Fig. 5. 1'5303-36. vue Iaterale, Loc. 5303, x 1
Fig. 6. T5303-81. vue periphero-laterale, Loc. 5303, x 1
Fig. 7. T5303-l2. vue Jaterale, Loc. 5303, x 1
Fig. 8. T5303 ·82. vue peripheriQue, Lac. 5303, x 1
Fig. !l. T5303- 25, vue laterale, Lac. 5303, x 1
Fig. 10. 1'5303-45, vue laterale Loc. 5303, x 1
Fig. 11. T5303-80. vue peripherique, Lac. 5303, x 1
Fig. 12a. T5303 79. vue laterale, specimen typique, Lac. 5303, x 1
Fig. 12b. Vue periphcrique du mcme specimen que 12a, X 1

T. S ATO : Ammonites de Kuruma

Planche 18

3a

la
2

ll

3b

Trans. Proc. Palaeont. Soc. Japan, N. S., 1\o. 20, pp. 119-I27, Pi. 19, Dec. 30, 1955

278. MIOCENE MOLLUSCA FROM NOTO PENii~SULA, JAPAN.
PART 1.

(I)*

KOICHIRO MASUDA
Departm~nt

of Geology, College of Education, Tohoku Uni\·ersity, Sendai. Japan1}1§:(f*-%i;Q)*fJflll:illl\i*lliiJ~; .:f:-Q) I, (I). *1il:l:Jl!tl:. @~2j"'.);;;;.ftfalC7)1:jJII!Ji~~!il;rtriltms · !~3lmll!J!I!fPIJ. i!111DlM~It:ilCt:j£~ t...-c'- '-!>mm=*Q)11J!'fi:l: 01. '-t:*lli'f u..:t~;. 4-ITII. Jttt:PI"'kiin· t-.tlm t...
t..:Jl. ft£1:--Jl. ''C~~ L.. t..:o -t_-Q)~51l:. J~U~!lrt-3fi,'?iil;ifll!itJ~7kV:rii'J<~Q)JifJH!l.l: '' t.&. 7,;,:::. c. Z.[lAf.l.l: L..
t.:.o il!I:IBJ'J.fPIJjg/fU !J !*m L..t..:{t£Q)J:ft, 12 .flfNl t. I WT.!lliMa-:?clX L..t.:.il:. ~roiQ)t&fr~'t!i:t.
=ttll 4 !!i!C.~j\ 1 fiJiQ)ili3lXHf!Jift..:o :!flEE~-·$

University, for his l<ind supervision. andto Dr. Kiyoshi AsA!'>n of the Instituteof Geology and Paleontology, Faculty of
Science, Tohoku University, for his encouragement and advice. Thanks arealso due to Mr. Akio I·IA;-.-tADA, teacherof the Najimi Middle School at Najimi,
Wajima-shi, for his assistance in collecting specimens.

Introduction and Acknowledgements

In the present article, are treated the
Miocene molluscan fossils collected fromthe Higashi-lnnai formation developedin Machino-machi and Yanaida-mura.
Fugeshi-gun, and Najimi. Wajima-shi.
Ishikawa Prefecture in the northern partof Noto Peninsula, Japan. The majorityof the specimens studied are now preserved in the collection of the Department of Geology, College of Education,and the minority in the collection of
the Institute of Geology and Paleontology, Faculty of Science, both of the
Tohoku University in Sendai. The
Cenozoic Mollusca in the collection of
the Institute of Geology and Paleontology. Tohoku University and of the
Saito Ho-on Kai Museimi in Sendai wereused for comparative work with thefossils from this region.
Acknowledgements are due to Dr.
Kotora HATAI of the Dep:1rtment of
Geology, College of Education, Tohoku

Geological Notes

The stratigraphy of the Tertiary strata
developed in this region was previously
published by the writer 0954). wl10
recognized the following formations, inascending order:
K<mosuyama volcanics :Andesite,basalt, dacite and their agglomerates.
and diorite. Part of the riiorite isyounger than the volcanics. Their
stratigraphic sequence is unknown.
-(stratigraphic relation uncertain)Tokunari formation: Pumiceous tuff
and brecciated tuff with lenticular layersof tuffaceous sandstone, tuffaceous mudstone and conglomerate. These rocks* Read June 28,.1955; received July 20, 1955.
119

Koichiro MASUDA

120

are intercalated between the flows ofandesite. basalt and dacite. This formation ·probably interfingers with theK6nosuyama volcanics, though theircontact was not seen.
( tmconformity)
Higashi-Innai formation: Mudstone,siltstone, san'dstone and conglomeratefrom the upper to the lower, in whichlignite is intercalated. Mollusca andForaminifera abundant.
.Awagura formation and Iwakurayama
rhyolite : The former comprises finely
stratified white or light green coloredpumiceous tuff. brecciated tuff and sandy
tuff. The Iwakura.yama rhyolite eruption took place during deposition of the
Awagura and is responsible for the large
quantities of tuff.
Najimi formation: Tuffaceous mudstone and tuffaceous siltstone. locallywith pumiceous layers and laminatedtuffaceous sandstone layers. Molluscan
shells and sponge spicules occur throughout the formation. In the northem partof the area. Radiolaria and Globigerina
are common and Cmtclwcele is the outstanding megafossil. In the southernpart of the area, Foraminifera are abundant and Globigerina is also found. The
common occurrence of Globigeritza suggests a free connection with the opensea.
·(unconformity)
Okawa terrace deposits: Sand. gra\rel
and clav. Fossils not found.
The fossils dealt with in this article
were collected from the middle or lower
part of the Higashi-Innai formation.
From the fossil locality of Tokunari, themost important of the Higashi-Innai

formation. the writer has identified 37
species in total among which 13 species
are new to· science. Further. several
genera are recorded for the first timefrom the Miocene or Cenozoic rocks ofJapan.
The Higashi-Innai formation shows
lateral change in lithological facies andcontains rocks of marine and brackishwater facies. Although there are many
fossil localities of this formation. only
those from the sandy mudstone containing much carbonaceous matter at
Tokunari. Macbino-machi, Fugesbi-gun(lat. 37°24'36"N., long. 137°05'20"E.) are
well preserved : these are described inthis mticle.
Notes on the Paleontology

The fossil fauna in the vicinity ofTokunari compared with those fromother localities consists of specimens
with intact valves. which are well preserved and little water-wom shells were
found. This fact indicates that thefauna probably lived in the environment
in which it was buried, at least it canbe inferred that it was not subject totransportation from a remote place.
Therefore, the said fauna may represent
a biocoenose. It is also worthy to mention that the brackish water genera as
Vicarya, Vicaryel/a, Cerithidea and
Batillaria do not occur in association
with Operculina, Miogypsitza and smaller
foraminifers, although they can be inferred to the same stratigraphic horizon.
These facts suggest that during deposition of the formation, both pure marine
and brackish water environments existed
at the same time but at different places.
and that the latter environment may
have been one of an embayment.
From the faunal elements of theHigashi-Innai formation it is evident

Miocene l'vlollusca from Noto Peninsula, japan.

that deposition proceeded under the
influence of warm and shallow waterconditions. It is also inferred from thestratigraphic relationship of the megafossils to the smaller foraminifers previously reported by K. AsAtm (1953), thatduring the initial stage the Higashi-Innai
formation was under the influence ofshallow waters, but by progressive development, the seas gradually became
deeper towards the upper part of theformation.
The occurrence of Vicarya, Vicaryclla
and other fossils given in the list indicate that the age of the fauna of theHigashi-Innai formation is Early Miocene, more precisely lower Middle Miocene in age.
The distribution of the respective
fossils given in the list show that theHigashi-Innai formation can be considered to be a correlative of the fauna of
the Meisen formation of Korea, theFujina of Shimane Prefecture. the Yatsuo
of Toyama, the Nishikurosawa of Akita,
the Shobara of Hiroshima, the Tsuyama
of Okayama, and the Tsukiyoshi formation of Gifu Prefectures. From such a
correlation it is evident that the warm
seas of that age were rather extensive
in distribution.
Description of New Species

Family Veneridae
Genus Callista Por.1, 1791
Callista chinensis takagii MASUDA,
n. subsp.
Pl. 19, fig. 7.

Shell small in size, rather thin, moderately inflated, transversely elongated ;surface ornamented with fine. numerous
concentric growth lines ; beak moderately swollen, bluntly pointed and situatedat about posterior one-third of shelllength. Antero-dorsal margin gently

121

sloping into rounded anterior side; postero-dorsal margin longer than anterior,gently sloping into rounded posteriorside ; ventral margin broadly rounded.
Dimensions (in mm.) :-Height 13.0, length21.5 (holotype), height 12.0, length 19.3
(paratype).
Occurreucc :-Few.
Depository:- DGS*, Reg. No. 2501
(Holotype).
Remarks-The specimens are characterized by their little inflated shell,rounded anterior end and narrow posterior end, and the position of the beak.
Callista chi1tensis (HoLTEN) (HIRASE,
1951, pl. 34, fig. 7), a common Recent
shell of Japan, is distinguishable fromthe present ~pecies by the less inflated
beak. broader anterior side, longer postero-dorsal margin, higher shell andmore compressed aspect.
This species is named in honor of Mr.
:tvlagokichi TAKAGI who helped thewriter in the field.
Genus Clementia GRAY, 1847
Clemeutia japonica MASUDA, n. sp.
Pl. 19, fig. 8.

Shell moderate in size, thin. transversely elongated, inequilateral ; posterodorsal margin nearly straight, posterior
end sharply rounded ; antero-dorsal
margin shorter than the posterior,forming concave arc with beak, its end
regularly rounded ; ventral margin
broadly rounded. Beak moderately inflated, directed forwards, small, nottouching.
Surface sculptured withcoarse concentric waves and fineconcentric threads, both irregularly
spaced: wave~ tend to become obsolete* DGS=abbreviation for Department of Geology, College of Education. Tohoku University, Sendai, japan. This abbreviation will
be used throughout in tlus paper.

K6ichiro l\'1AsuDA

122

near the margin. F.scutcheon obscure.
Characteristics of hinge unknown. Dimensions (in mm.) :-Height 34. length
44, depth 13, apical angle 120°(holotype).
Occurrence :-Rare.
Depository:- DGS, Reg. No. 1382
(Holotype).
Remarks :-This species is characterized by its transversely elongated shell,irregularly spaced concentric undulations
and fine concentric threads, and large
apical angle.
. Clementia (Egesta) ·mart iui (Cr.ARI.;)(WooDRING, 1926, p. 40, pl. 15. fig. 3)
from the Miocene San Pablo formation.
Catra Coasta County, California, resembles this species in having a transversely
elongated shell. rather large apicalangle, and in the sculpture of the external surface, but it is distinguishable
from the present one by the proportionof length and height, details of theposterior margin, and by the shallowly
concaved anterior margin. Clementia
nalummrai Orr..:KA (OTVKA, 1938, p. 14,
pl. 1. figs. 7, 11). a Miocene species ofJapan differs from the present one in
the nearly quadrate shell outline.
Family Tellinidae
Genus Tellina LIN:-:AEUS, 1758
Tellina !wmadai MAsvuA, n. sp.
Pl. 19, figs. 10, 11.

Shell small in size. thin, compressed.
transversely elongated; surface finely
and concentrically sculptured : beaks
small, approximate, pointed, and situatednear middle of shell length. Anterodorsal margin rather sharply sloping
into narrowly rounded anterior side;
postero-dorsal margin a little longer
than anterior. nearly straight above andmerging into rounded posterior side :ventral margin very broadly rounded ornearly straight ; anterior and posterior

ends rounded. the latter more broadly
than the former which is more abrupt.
Escutcheon narro\v and short, slightly
clepreswd. Surface sculpture consists ofnumerous, regular, close-set and fineconcentric lines, and with faint foldfrom beak to anterior end. Interior
characters inaccesible. Dimensions (inmm.) :-Height 8.6, length 14.4, thickness
3.3 (holotype), height 5.5, length 9.5(paratype).
Occurreuce :-Common.
Depositmy :- DGS, Reg. No.· H157
(Holotype).
Rcmarl~s :-This species is characterized by its small. transversely elongatedand compressed shell with fine, numerrous concentric lines on the surface andfaint fold which extends from beak toanterior end. Among the paratype specimens there is one having the anterior
end bent to the right.
This species more or less resembles
Telliua iridal/a 1VIARTENS (YoKOYAMA,
1924, p. 41, pl. 5, fig. 23) which is a
Pleistocene and Recent species of Japan,
but that species has a beak situated moreposteriorly than the present species.
Thi~ species is named in honor of Mr.
Akio I-IAMAln who helped the writer inthe field.

Tcllina 110/oeusis 1\L-\scmA. n. sp.
Pl. 19, fig. 12.

Shell small. thin, comprtssed. transversely elongated, nearly equilateral:
surface was probably finely and concentrically ornamented (original shellmaterial not preserved): beaks small.
pointed, and situated nearly at middle
part of shell length.
Antero-dorsal
margin nearly straight. gently sloping
anteriorly into well rounded anterior
side: postero-dorsal margin nearlystraight: anterior and posterior ends

Miocene lVIolluscn from Noto Peninsula, japan.

regularly rounded : ventral margin verybroadly rounded, or nearly straight.
Escutcheon very narrow. depressed.
Surface with weak fold extending frombeak to anterior end. Pallial sinus very
deep. Interior characters inaccesiblc.
Dimensions (in mm.) :-Height 12.5. length23.0 (holotype).
Occurre11ce :-Rare.
Depository : - DGS. Reg. No. 2502
(Holotypc).
Remarl~s-A single specimen lacking
the original shell matter is in the col·lection. This species is distinguishable
from Te/lina hamadai described in thisarticle by the larger size of the shelland nearly equilateral outline.
Family Skeneidae
Genus TeitUJstoma A. Ao.\MS, 1853
Te:'11ostoma yabei MASUDA, n. sp.
Pl. 19, figs. 16·18.

Shell very small in size. lenticular inform; surface rather smooth, somewh~t
glassy, ornamented with microscopic
network due to the crossing of spiraland longitudinal threads, making interspaces appear as elongated pits. vVhorls
about 4. nearly nat. rapidly enlarging,separated from each other by rather

123

distinct, impressed sutures. Body whorl
ventricose, with regularly rounded periphery; base a little concave; imperforate. Aperture nearly circular; outer
lip thin, simple ; inner lip stout, withwidely spread. smooth callosity. Dimension (in mm.) :-Height 1.8, maximum
diameter 3.G (holotype). height 1.9, diameter 4.1, height 1.5, diameter 2.8, height
1.7, diameter 3A (paratypes).
Occurrence :-Abundant.
Dl'pository : - DGS. Reg. No. 1537
(Holotype).
Rt'IIWrks :-This is a rare genus in the
Cenozoic formations of Japan, and thepresent record may be the first.
TcitUJstoma smikron GARD:\ER (GARD·
NER, 19,18, p. 187, pl. 25, figs. 18-20) fromthe Pliocene Waccamaw formation,
Neills Eddy Landing, Cape Fear River,
Columbus County, North Carolina, resembles this new species. but it differsfrom the present one by the callosity
which is slightly sunken below thesurrounding surface and the feeble
sculpture.
This species is named in honor of Dr.
Hisakatsu YAnE, Emeritus Professor of
the Tohoku University. who founded theInstitute of Geology and Paleontology.
Faculty of Science, Tohoku University,
Sendai, Japan.

Emmcula sp.

I I i_ _ _ I
I Barbatitl cf. uelsukic11sis HATAI and Nls!y,~MA--;-- - · - - -~ F _ _ _ jI

1

I

ll'ucula11a sp.

,----1 Anadara(Anadara)kakehataemisHATAlandNISIYAMA•'R~~ar~ad~) lm~ct1si~ a~\~~~IA-------------------~--

1

A

I

=-l=--l_A- -~-~ -~1

Koichiro MAsuDA

124

List of Fossi8 (* illustrated)'1-~ -~~

Genus and ~;:i:;------

Localities

I

I

NislYA~fA

2

A_muJara cf. makiyamai HATAl and
1-G_/_y-cy_m_e_r-is-c-f.-r-l-,y-u-c-h-ol-le-l-lo-id_e_s_N_o_M_U_R_A_a_n_d_li_A__l-.A-l--,134LithojJhaga sp.
ch/amys tlisataieiiSis OTIII\A

--

5

6

7

8

-- - - -~:- -~-,-1 C-------------1-:-

Brachidontes sp---------- -- --II]

IC;A F-~F'-F~--~l

----

-

c.-1---___ · _ . _ _ _ _ I_______ I_ !-~--~FF I:---F-~-~~-=

I~ =--i ~ j=-1=1- ~=- - - - - - - - - - ----1- · - - - - 1-1::::::::::: ::mmi

Placopecten sp.
Mania radiala(YoKoYAMA)___ _ _ _____(DEsHAYF.S)R

R•1 ___

1! _---~-~--.-_1 R , F
l1•

1_o__s_tr_e_a_c_f_.g_,_·e_as_T_H_u_N_B_E_·,_w_._____-_~~ --=~-~- _[=~~~~- -~~~-_:_:_:r_r:_:_:_::_~-v-it-es-ta_Y_<_JK-·o_Y_A_M~A~~~-~~==------------~--~~---•_
~~~

~I

1A

Fl

lR' f~-F ~~I---:;;~as sp.

--,- --! -- :--- - - R~---------~-- - - --~--F·--~--~-CI.-. _._f_, ___ _~-~---]ommisiella meisensis 1\-lAKIYAMA*-1

Luci11isca k-hataii (OTUKA)*------------------------Luci11isca sp.

F

F

Papyridea sp.

--i->

Clinocardi11m sp.

- - - - - - - - -----Fulvia sp.

F :!F i :III

I--C~y~c-li-~_w___j=ap-o~ll_l_.c_l_I-~---A~!'.-1-,_\-D-A-_-_-_-_-----~~------~~~----; _ ~-~=~-~ A ~=~=-~---~
Pilar sp.
·
' F I
' .
I

Callist-;;;h;,~ t~;i~;;iM~\,n. suusp.*

Dosinia cf. nagaii OTUK A

Dosit~a sp. ___ _ _ _ _

_ _

-

-=

=~=j==~=~
\

I

F

I

,

__ _ __

F ._ F
1

!.~

: R

R

~~.~--.

1

1

' F

t

: _ _ _ _ . _ ,.

1

1

F·

---~--l _ _ ,_ _ _ j_

Protothaca tateiwai J\-1AK I YA1\IA

Katelysia sp.
Clemmtia japo11ica MASl'l>A, n. sp...

I= -c=l
1

I

1

1

.

1

I

-~--~--- --.-1 R~--- -- --~
1

I

125

Miocene Mollusca from Noto Peninsula, Japan.
Ust of Fossils (* illustrated)-;:n~s__:~t~d -Si>:_:~:_:c~i-e'-'-s'- - _-_-_-_-_-_--~---------_-_-_-___L_o_c:_~_it_~_:-=-=~1--1-'-[_2_:~ ~ ~ ~~ 7 8
Clementia 11akamurai OTllKA

[

F

F

I

F--~----FI

Clementia sp.

___ _- - - - - - - - - - - - - - - - - - - - - - - - - - ! - - - - --~- ' - - - - Paplzia cf. lzirabayasii OTUKA
c
I
R

I

-

Cardilia cf. yudae11sis OTUKA------1--1-----

Soletellina mi11oensis

Tellina lzamadai 1\IAstiDA, n. sp.*

--

-

1

-- - - -R:-i(YoKOYAMA}"

R

-~-~-

-- c:

1-

-,-

Tellina ttoloet~~i~--~1-A--st_J_D_A_.-n--.s-p-...- - - - - - - - - - ·~~~-=!= =~ R . __- ---~-1

1__
1'<_e_ll_in_a_!lP_·_ _ _ _ _ _ _ _ _ _ _ _ _ _ _ _ _ _ _ _ 1__ I__ I _ _~I__

Macoma sp.
I
I
1_S_o_l_e_11_c_f_._R_o_u-ld_i_C_c_>_N_R_A_I_)*_______ - - - - - - - --~-- -_- ------------~----

___ _

R- R-

!F

---~-

--A----~-

Phaxas cf. izumoensis (YoKoYAMA)*---------I----~- [ {--Thracia sp.

-i----------------------------------------·1-CryfJfomya lmsoensis YoKoYAMA-----------------------------------1--Denlalium cf. wcinkaujii DuNKER[

R
1 - - - - - - - - - - - - - - - - - - 1 - - -~- Denlalium sp.
R
I__

--v-1

Margarilcs sp.

--

-~--

1_c_a_u_io-~-,o-,-,-a-<-J-..,-~~-,,-o-,)-,,-a-,-,,-,c-,-za-k_u_e_n_s_is_H_A_T_~-~_a_n_d_I_{_O_T_>_\K___A_*~I-- F

==

---

I= =I r=-~--~R-1-

Troc/ws sp.

c__ R _-_A ~~--I

Calliosloma (Calotropis) simane NoMl'RA and HATAI--

Teinosloma yabei 1\[ASUDA, n. sp. •Turritella sp.

---------~~--F I! _ _ F-1-_!1. c

Vicaryella noloensis 1\·IAstJDA, n sp.*=r

.1:.-_ ___-,--_---

Vicarya callosa japonica YARE and HATAI*A

Vicaryelln sp.
F1--------------------------1-. --cCcrillzidea kmrpokuensis (MAK 1YAI\IA)*~~-·--RI

1------------------------1------ --1

A: - -

Cerithidea toktmariensis MASUDA, n. sp.*

- - - - - - - - - - - - - - - - - - - - - - - - - - 1 - - - - - - -- Ceritlzidea sp.

F--

Batillaria cf. yama11arii MAKIYA:VIA
1

I__ :_I_Z
1

Koichiro MASUDA

126

List of Fossils (* illustrated).---------------------------------------------

-

----

-

-

-

-

-

-

Localities

~-

-

8

--

Pmclava sp.
Amalhina sp.

===
=
=
1
R

--_--_--==~ F [~Poliuices (Euspira) meisensis MAKIYAMA_*_______
Polinices (Euspira) olukai l\IASVDA, n. sp.•

- - - - - - - ----------

Polinices sp.

F--~p-~- - ~ ------s~,-,_i_c_,-,_,-.,=e=ia___y~o_-k_o_ya~-,_a-i~C-N_'_o_~_n_J_R_A-ancl _H_A_T_A__I_)_______---~-~--- F ~~- ___Bursa sp.

-------------------~-----J---j--.1_

Chicoreus asanoi MA<>rDA. n. sp.*--------

-

R

-~--

A

- - - - - - - - - - - - - - - - - - - - - - - ---- - - ----

Chicoreus 110/oensis MAsUDA, n. sp.*------------ - - - - - - -F------------ - - - - - --I~----

Chicoreus lateiwai HATAl and Kn'I'AKA

F----------

Neplrmea sp.

---------Pltos sp.

R

1 - - - - - - - - - - - - - - - - - - - - - - 1 - - - 1 - - --~-Nassarius notoe11sis MAsUDA, n. sp.*A

Mitra sp.

- - - - - - - - - - - - - - - - - - - - 1 - - - - - - - - - -------- -Turbonilla sp.

----------------1------Pyramidella halaii MASUDA, n. sp.*

R

R------·-

Acteocina hamadai MASUDA, n. sp''
Coplolhyris grayi (DAVIDSoN)--------Loc. 1:

R--~--A------------------~-~-~-~-~~-~----

Nishi-Innai. Najimi, Wajima-shi, Ishikawa Prefecture. Higashi-Innai formation (finesandstone).
Loc. 2: Higashi-Innai, Najimi,Wajima-shi. Higashi-lnnai formation (fine sandstone).
Loc. 3: Konishiyama, Najimi, Wajima-shi. Higashi-Innai formation (very fine sandstone).
Loc. 4: Tannoji, Onishiyama, Najimi, Wajima-shi. I ligashi-Innai formation ·(medium sandstone).
Loc. 5: Tokunari, l\1achino-machi, Fuge5hi-gun. Higashi-Innai formation (sandy mudstone
containing much carbonaceous matter).
Loc. 6: Hizumewaki, Yanaida-mura, Fugeshi·gun. Higashi-Innai formation (very fine sand·stone).
Loc. 7: Ishii, Yanaida-mura, Fugeshi-gun. Iligashi-Innai formation (tuffaceous very fine sandstone).
Loc. 8: Wasumi, Yanaida-mura, Fugeshi-gun. Higashi-Innai formation (tuffaceous fine sandstone).
Abbreviation: A-abundant, more than 10 specimens; C-common, more than 5 specimens;
F-few, more than 2 specimens; R-rare, only one specimen.

K. MASUDA : Miocene Mollusca

K. KuMAGAI photo.

PLATE 19

Miocene l\Iollusca from Noto Peninsula, Japan.

.127--------

Explanation of Plate 19
Barbatia cf. uetsukiensis IIATAl and NlSlYAMA, DGS, !{eg. No. 1387, Right valve, x 1.
Loc. Tokunari, ~vlachino-machi, Fugeshi-gun, Ishikawa Prefecture.
Figs. 2a-b. Anadam (Anadara) kakehataensis HATAl and NlSIYAMA, DGS, Reg. No. 1383. a,Right valve, x 1. b, Posterior view of beaks and area of 2a, x 1. Loc. Same as above.
Fig. 3. Anadara (Anadara) kurosedaniensis HATAI and NlslYAMA, DGS, Reg. No. 1386. Leftvalve, x 1. Loc. Same as above.
Fig. 4. ]ommisiella meise11sis MAKIYAMA, DGS, Reg. No. 1193. Left valve, x 1. Loc. Same asabove.
Figs. 5, 6. Lucinisca k·hataii (OTUKA), DGS, Reg. No. 1116. 5, Left valve, x 1. 6, Right valve,x 1. Loc. Same as above.
Fig. 7. Cal/ista chinensis takagii MAsUDA, n. subsp. Holotype. DGS, Reg. No. 2501. Right valve,x 1. Loc. Same as above.
Fig. 8. Clementia japonica MAstJDA, n. sp. Holotype, DGS, Reg. No. 1382. Right valve, x 1.
Loc. Same as above.
Fig. 9. Soletellina minoensis (YoKOYAMA). DGS, Reg. No. 1114. Internal mould of rigut valve,x 1. Loc. Same as above.
Fig. 10. Tellina hamadai MAstii>A, n. sp. Ho:otype, DGS, Reg, No. 1657. Right valve, x 1. Loc.
Same as above.
Fig. 11. Tellina hamadai MAslJI>A, n. sp. Paratype, DGS, Reg. No. W57. Right valve, x 1. Loc.
Same as abQve.
Fig. 12. Tellina notoemis MAslli>A, n. sp. llolotype, DGS, Reg. No. 2502. Internal mould of rightvalve, x 1. Loc. Same as above.
Fig. 13. Solen cf. gouldi CONRAil, DGS, Reg. No. 2499. x 1. Loc. Same· as above.
Figs. 14a-h. Phaxas cf. izumoensis (YoKoYAMA), DGS, Heg. No. 11 12. a, Umbonal view. x 1. b,
Left valve, x I. Loc. Same as above.
Figs. 15a-b. Calliostoma 11amuclzakuensis IIATAl and KoTAKA, DGS, Reg. :r\o. 1110. a, Aperturalview, x 3. b, Dorsal view, x 3. Loc. Same as above.
Figs. 16a-c. Tcinostoma yabci MAstrDA, n. sp. Ilolotype, DGS, Reg. No. 1537. a, Apical view, x 8.
b, Umbilical view, x 8. c, Apertural view. x 8. Loc. Same as above.
Figs. 17a-c, 18a-c. Tei11ostoma yabei MAsUDA. n. sp. Paratypes, DGS, Reg. No. 1537. a, Apicalview, x 8. b, Umbilical view, x 8. c, Apertural view, x 8. Loc. Same as above.
Fig. 1.

128

PROCEEDINGS OF THE PALAEONTOLOGICAL SOCIETY
OF JAPANra *"6£E#J~~m 61 mJWIJ~J 1955 ~ 6 n
1s n J.i!:frll k~JJI!et.$ttl!Wq';tv;#Jt¥.¥t3f :-r::: h":V>'LimtUH.-tc (~~off 23 :fi)o ~lliii.ii~~v:
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-······-·····-·································- ~::{tre=n~'¥lWJ~~ii*Rf.'&~i¥.J~V:1.:.1E:j&i~(J(jfiJf:1E........ .
············ ······ ······ ······ ···· .............. X/;~B.:::~B

On the " Retral Processes " in Some Elphidium(ft~) .................................... Hiroshi UJIIEOn the Variation of Elphidium crispum s~nsu
Jato (ft;~) ........................ Yukinori Ft'JITA
Notes on the Osobudani Conglomerate of the
Fukuji Ditrict, With Reference to Some Per·
mian Fusulinids of its Limestone Pebbles(11::iflt) ................................. Hisayoshi lGo
On Some New Species of Genus Rauserella
from lbuld-yama, Shiga Prefecture, Central
Japan ( ft~) ............... 1\fanabu KonA YASH I'-'"C

1.:. "":)........................................... ~ F"'1

On the Form Variation of Lima goliath andits Allid Species ..................... Shigcru AoKI
On Some Miocene Species of Dosinia fromJapan (~ib't) ........................ Saburo KANNOOn Luci11oma aokii, A New species from the
Pleistocene Deposits of Chiba Prefecture,
Japan (ftlft) ............... Katsumi HIRAYAMA
Notes on Molluscan Fossils from Problematical Horizon of the Tomioka District,
Northern Joban Coal-field, Japan (ft~) ...
................................ Katsumi HIRAYAMA,On the Molluscan Fauna found from theNorthern District of Tanzawa Mountain·
land (ftm'i!) .................. Matsutaro SHIBATA
2, 3 "' G!ycymeris 1.:.--:)

15'MJ•F'3m=*l11i"'................................................ fMiJIIW.:::.
( i-"'I) Ctt!D'l). ................................... ttm~-n6

fJ13ft~f..JJ;:it.fflS "'rf:t~i t!tiM·f*lWJ~t::. -:)I,, -c1L}'I·I~I:i!!iJ51H::.:tult7.>. '-'h\<J>7.>~Wi~J'J.1t.;f.it'(l~t,¥~t 1\~) ... .................. -........... 7kJrnlrr

ft!Rklll.W.~i:tcl11iaNfti!~?tfiJ · B~T{I%i~iittl•l?ilflHI Lt-:.~Hl}U':C.Ifijlf'il Echit~ocorys

1 1 1MF'il'~~='.:*.imo:J Lucinidae 1.:.--:)I.''C ....... .

1.,-c .......................................... .ff:.f(IJIJ~.;cFounding of Problematic Fauna From the
Abukuma Mountainland (ftitr~) .............. .
............ 1\lasae OMoRI and Abukuma grouprm!ill~f:ll*h'9llfi.!ii=ttst-~::. 01. ,-c
.:t:~

1:

-'(-Q) 2:

Taxodonta lkV: Prionodonta
Periplomya e/!iptica NAGAo &0TATSVJ\IE Q)pj.~;M ................. .

.................. :rtJJillili-·~~ . niJB:ll*1<:
Ober die Plcurogrammiltodontinae, eine Jiingmesozoische Prionodonte Unterfamilie ..... .
................................. Koichiro ICHIKAWA
Yebisites, a i\ew Lower Jurassic Amr:nonite
from J;~pan (ftif\'l) ...... Tatsuro MATSUMOTO
On the Damesellidae and Kaolishaniinae withNotes on Parakoldi11ioidia and the so-called
Wuting Trilobites ( ftitr~) ....................... .
............ , ....... , ............... Teiichi KonAYASill
Notes on the Cambrian Fossils from Yentzuy<ti, Tawenkou, Shantung (fUM) ........ .
.................................... Teiichi KoBAYASHIElectron-microscopic Fine Structure of Fossil
Diatoms IV ........................... Haruo OKUNOA New Species of Damesitcs from Cenomanianof llokkaido, Japan ( ftim) ................... ..
..... Rinji SAITO and' Tatsuro MATSUMOTOOn Myoplzoria goldfussi var. kobayashii KAMBE ................................. Keiji NAI\A7.AWA
qm!fl · l'·li6::=11*"' Bakevel!ia j::."":)~,,-c ...

................................................... !f:tjR~:.:
Pseudorause1·ella, a New Permian Fusulinid
Genus from the Kuma Massif, Kyushu,
Japan (1\ilfl) ............ Kametoslii KANMERA

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CONSTITUTION
of the
PALAEONTOLOGICAL SOCIETY OF JAPANARTICLE 1. Name
The Society shall be known as the Palaeontological Society of Japan. The Society is a sectionof the Geological Society of Japan.
ARTICLE 2. Object
The object of the Society shall be to promote the study of palaeontology and related sciences.
ARTICLE 3. Achievement
The Society in order to execute Article 2 shall ( a) issue the Society journal and other publi·
cations, ( b ) hold or sponsor scientific lectures and meetings, and ( c) sponsor collecting or
field trips, and lectures.
ARTICLE 4. Membership
The Society shall be composed of persons who are active of interested in palaeontology or
related sciences, and shall be known as regular members, honorary members, and patrons.
ARTICLE 5. The members of the Society shall be obliged to pay annual dues to the Society, forwhich they shall enjoy the privilege of receiving the Society's journal and of submitting
papers which have been read and discussed at the meetings for publication in the Society's
journal.
ARTICLE 6. Administration
The Society shall have the following organizations for its administq.tion.
(a ) General meeting. The general meeting shall be composed of the Society members. Morethan one tenth of regular members sh:tll be present to hold general meetings. Administrative affairs shall be decided during the general meeting.
(b) President. The president shall be elected from among the regular members. The president
shall represent the Society and supervise its business matters.
(c) Council. The council shall be composed of councillors who are elected from among theregular members. The council shall discuss administrative affairs.
(d) Business council. The business councillors shall be elected from among the council members, and shall administer business affairs.
(e) Officers shall be elected by vote of returned mail ballots, as a general ntle.
ARTICLE 7. Amendments to the constitution shall be by decision of the general meeting.

By-Laws and Administration
ARTICLE 8. The Society's journal shall be issued quarterly.
ARTICLE 9. Regular members shall be persons who have knowledge, experience, or interest inpalaeontology or related sciences.
ARTICLE 10. Patrons shall be selected individuals or organizations who give special support to theobjectives of the Society.
ARTICLE 11. Honorary members shall be persons of distinguished achievement in palaeontology.
The council shall nominate honorary members for decision by the general meeting.
ARTJCLE 12. Applicants for membership to the Society shall submit their full name, mailing address, date of birth, occupation, and name of school from which they graduated.

Dues
ARTICLE 13. Rates for annual dues of the Society shall be decided during the general meeting.
Annual dues for regular members are Yen 600.00 (domestic members) and U.S. $3.00 (foreign
members). Patrons are individuals or organizations donating more than Yen 15,003.00 annua!Iy. Honorary members are free from obligations.
ARTICLE 14. The Society income shall be from membership dues and bestowals.
ARTICLE 15. The Society shall have one chairman, fifteen councillors, and several business councillors, whose term of office shall be two years. They may be re·elected.

Addendum
ARTICLE 1. There shall be four business councillors for the present.