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Jul y 30, 1984

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MATSUMOTO et ai.: Cretaceous Nautiloids - IV

Plate 60

3

23

63

6b

6e

6d

MATSUMOTO et al. : Cretaceous Nautiloids - IV

Plate 59lbTrans. Proc. Palaeont. Soc. Japan, N.S., No. 134, pp. 335-346, pis. 62-66, July 30, 1984

777. CRETACEOUS NAUTILOIDS FROM HOKKAIDO-V*
TATSURO MATSUMOTO l ), TOSHlYA MIYAUCHl 2), YASUMITSU KANIE 3>,
YUICHIRO MIYATA l ) and YOSHIRO UEDA 4)

Abstract.
This paper contains Parts 5, 6 and 7 of the Cretaceous nautiloids
from Hokkaido. Part 5 concerns with the species from the Lower Cretaceous. The
descriptions are given under 3 sections by different coauthors. Section A by T.
Matsumoto and T. Miyauchi is on Cymatoceras sp. aff. C. virgatum (Spengler) fromthe Albian of Oyubari and Soya, which differs from the holotype from India by lessinflated whorl. Section B by T. Matsumoto, Y. Kanie and Y. Miyata is on Cymatoceras sp. cf. C. sakalavum Collignon from the Albian at a locality (Text-fig. 1)
about 17 km NNW of Urakawa. Section C by T. Matsumoto and Y. Ueda is on
Cymatoceras sp. aff. C. mikado (Krenkel) from the Neocomian of Rebun Island,
which resembles C. mikado from Tanzania but differs in having subtrapezoid
instead of subcircular whorl section.
Part 6 by T. Matsumoto is on an additional species from the Santonian of
Ikushumbets based on T. Takahashi's collection. It is identified with Cymatoceras
sharpei (SchlUter), being especially similar to the Gosau form.
Part 7 is a summary of results by T. Matsumoto. (1) The described 15 species
are listed with the age of the original beds (see p. 341). (2) The phylogeny of the
Nautilidae is tentatively presented (Text-fig. 4), showing the relationships of the
newly established genus Kummeloceras with other revised genera. (3) In Hokkaido
nautiloid species are different from a stage to another. Certain species are characteristic of a particular zone. Some are well correlated with the allied ones inextra-Japanese provinces. (4) Many nautiloid genera show world-wide distribution
during the Cretaceous Period, but at specific level there are regional differences.
This is exemplified also by the species from Japan. (5) Discussion is given on thepalaeoenvironment on some species from Hokkaido.

This article consists of the following threeparts:
Pa,rt 5.
Part 6.

Part 7.

Some Nautiloids from the Lower
Cretaceous of Hokkaido
An Additional Nautiloid from theSantonian of Hokkaido
Summary of Results

PART 5. SOME NAUTILOIDS FROM THE
LOWER CRETACEOUS OF HOKKAIDO*Received January 13, 1983; read January 23,
1983 in Tokyo.
1) c/o Department of Geology, Kyushu University 33, Fukuoka 812.
2) Natural History Collection of Northern
Hokkaido, Midori 6, Wakkanai 970.
3) Yokosuka City Museum, Yokosuka 238.
4) Geochemistry & Technical Service Department, Geological Survey of Japan, Tsukuba
305.

T. Matsumoto, T. Miyauchi, Y. Kanie,
Y. Miyata and Y. Ueda
Introduction
The Lower Cretaceous deposits of Hokkaido
are not so prolific in cephalopods as the Upper
Cretaceous ones. Fortunately, however, some
335

336

Tatsuro MATSUMOTO et al.

nautiloid specimens have been obtained during
our geological field works. The persons con·cerned with the field works cooperate withthe first author in describing the species. Accordingly, this part is subdivided into three
sections, A, Band C, with the authorships
indicated in parentheses under respective subtitles.

end of the phragmocone is 110 mm in MNH. 145
and 130 mm in GK. H5726.
The whorl is well rounded in cross-section.
It is somewhat (but not much) broader thanhigh, with the maximum breadth in the lower
part. The umbilicus is almost closed in MNH.
145, which has shelly layers, and very narrowly
opened in the internal mould of GK. H5726.
The Cymatoceras type ornamentation is wellshown on the shell of MNH. 145 and partly onthat of GK. H5726. On the shell of late growthstage the so-called ribs are flat-topped andseparated by narrower groove-like interspaces.
They are somewhat sharper on the earlier whorl.
They are scarcely impressed on the internalmould. The rib bifurcation is frequent andmultiple, occurring near the umbilical margin,
again at various places on the flank and even on
the ventral part. The backward sinus of the ribson the venter form an obtuse angle of 110°130°. On the venter of the body-chamber, withheights of about 70 to 80 mm, the ribs number
5 in MNH. 145 and 6 in GK. H5726 within
the distance of 30 mm.
The septa are moderately frequent. They
number 10 in GK. H5726 and 9 in MNH. 145
in a half whorl. The suture is gently sinuous,
showing a very shallow external lobe, low andasymmetric saddle at the ventrolateral part,shallow and broad lobe on the main part of theflank, and a small and low saddle at about theumbilical shoulder.
The position of siphuncle is unknown.

A. A Nautiloid Species from Soya and Oyubari(T. Matsumoto and T. Miyauchi)
Family Cymatoceratidae Spath, 1927
Genus Cymatoceras Hyatt, 1884
Cymatoceras sp. aff. C. virga tum (Spengler)
PI. 62, Fig. 1; PI. 63, Fig. 1; PI. 64, Fig. 1
Compare:1910. Nautilus (Cymatoceras) virgatus Spengler,
Beitr. Pale/ont. Geol. Osterr.-Ungarns u. d.
Orients, vol. 23, p. 130, pI. 11, fig. 3; pI.

12, fig. 7c.
Material:-MNH. 145, obtained by H. Honmaon the eastern coast of Cape Soya, and GK.
H5726, obtained by T. Matsumoto at loco Y632
of the Oyubari area.
Description:-In MNH. 145, the left half ofthe body-chamber with the test and the preceding septate part are preserved. GK. H5726 consists mostly of the internal mould of a septate
whorl and a portion of the body-chamber, with
only a part of the test preserved.
The shell is fairly large. The diameter at theTable l. Measurements (in mm) of Cymatoceras sp. aff. C. virgatum.
Specimen

DiameterUmbilicus

MNH. 145

145

o (?)

144.0

12.0 (.08)(-180°)

GK. H5726"(-210°)*restored (47 x 2)

HeightBreadth

BIH

90

(.62)

94* (.65)

1.04

55

58

1.05

83.5 (.58)

94.0 (.65)

1.13

46.4

54.5

1.11

777. Cretaceous Nautiloids
Comparison:-With respect to the generalshell-fonn, the characteristically branching ribs
and the gently sinuous suture, the two specimensfrom Hokkaido closely resemble the holotype(by monotypy) of Cymatoceras virgatum(Spengler) (1910, p. 130, pI. 11, fig. 3; pI. 12,
fig. 7c), from the Upper Albian of southern
India. That holotype, however, has a muchbroader whorl than ours, for its B/H = 1.24 even
if B is assumed to be 8.6 as measured from his
illustration. We, therefore, hesitate to conclude
the speCific identity, unless the extents ofvariation be proved to overlap considerably
between the two provinces. According toSpengler, C. virgatum has tripartite ribs inaddition to the frequently bipartite ribs. On
MNH. 145 an apparent tripartite state is seen
where the points of bifurcation are approximated.
The holotype of C. carlottense (Whiteaves)(1900, p. 269), from the Albian Haida Fonnation of the Queen Charlotte Islands, British
Columbia (Canada), whose replica has been
gifted to Kyushu University from the Geological
Survey of Canada by courtesy of Dr. J. A.
Jeletzky, shows a similar mode of rib branching
but its ribs are narrower and more numerous
than those of our fonn and also the holotype of
C. virgatum. Furthennore, that Canadian species
has a less rounded whorl, with less convex flanks
and a broader venter. Its body-chamber has ashallow groove along the median line of theventer. This character is undevefoped in the
Indian and the Japanese species.
Occurrence:-MNH. 145 is in a calcareous
nodule of pumiceous siltstone. It was transported to loco So. 367 on the coast of Chiyenaibo, eastern side of Cape Soya, northern Hokkaido. At the same place another transported
nodule contained a fragmentary ammonite
resembling Pictetia or A rgonauticeras. As thegeologic structure is complicated in the eastern
coastal area of Cape Soya, the original bed of
MNH. 145 has not yet been detennined, although the existence of the Albian is suggested.
At the nearby localities in the south, there
occur Parajaubertella imlayi Matsumoto and

337

Desmoceras kossmati Matsumoto, suggesting theLower Cenomanian.
GK. H5726 is in a calcareous nodule from thedark grey siltstone exposed at loco Y632, on thefirst branch of the Hikageno-sawa, a tributary
to the River Shiyubari, Oyubari area, central
Hokkaido. The rock is assigned to Unit If, thetop member of the Lower Yezo Group. This
unit is Albian on the ground of stratigraphic
position and the occurrence of Tetragonites
cf. T. timotheanus (Mayor) (see Matsumoto,1942).

B. A Nautiloid Species from the Urakawa District(T. Matsumoto, Y. Kanie & Y. Miyata)
Cymatoceras sp. cf. C. sakalavum Collignon
PI. 64, Fig. 2Compare:1949. Cymatoceras sakalavum Collignon, Ann.
Geol. Servo Mines, no. 16, p. 41, pI. 6,
figs. 1, 2;pl. 21, fig. 1.
Material:-A single specimen obtained by Y.
Miyata and Y. Kanie at loco U 1092 in thefield work of 1978, now kept at the Yokosuka
City Museum, YCM. Ur 1092001.
Description:-Both the phragmocone and the
body-chamber are distorted. The diameter,
without reconstruction, is 135 mm, but it musthave been originally somewhat smaller.
The whorl is somewhat higher than broad
under the crushed condition but must have beenoriginally slightly or somewhat broader thanhigh, with more rounded outline. The umbilicus
is almost closed.
The surface of the shell is ornamented withrather flat ribs, which are separated by narrowerinterspaces. The number and coarseness of theribs are moderate for the genus, 32 being
counted on the venter of the last half whorl. The
rib branching is seen fairly frequently at or below
the middle of the flank. The ribs show someflexuosity on the flank, beginning to curve
backward at about the middle of the flank orslightly above it. The ventral sinus of the ribs
looks to be rather gentle on some part but fairly
pronounced on other parts. Probably thereTatsuro MATSUMOTO et ai.

338

is effect of secondary deformation in somedegree.
The suture is partly exposed, showing the
gentle sinuosity, but again the curvature seems tohave been modified by the secondary deformation. The siphuncle position is unknown.
Comparison:-The above described specimen
is so poorly preserved that it is hardly identified
with certainty. On the ground of the observed
characters, however, it is best comparable with
Cymatoceras sakaiavum Collignon, 1949, fromthe Albian of Madagascar.
C. carlottense (Whiteaves) (1900, p. 269),
from the Albian of the Queen Charlotte Island(British Columbia), is somewhat similar to C.
sakalavum, but has more numerous ribs whichshow a narrower angle of ventral sinus. There is ashallow depression along the median line of theventer on the living chamber of that Canadianspecies.
C. pseudoneokomiense Shimizu (1931, p. 18,
pI. 1, figs. 1-4), from the Upper Aptian of
the Miyako area (Northeast Japan), has a subtrapezoidal whorl-section and its ribs are separated by wider interspaces and show less frequent
branching than those of the present species. In
that species the ribs run almost radially from theumbilical margin to the ventrolateral shoulder,without showing such flexuosity as seen on thepresent specimen or on the typical specimens of
C. sakalavum.
Occurrence:-Loc. U 1092, on a branch
stream (called the Matsumoto-zawa) of the
Sakubai, a tributary to the River Kerimai,
Mitsuishi, Urakawa district, southern central
Hokkaido; solitarily found in massive fine-N.==: claystone
YVvV tuff:~.:::

sands tone;:::::::::::::: greens tone

42°15 'N

~
~

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T
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142 40'E-:.~\ ~)( \.
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33

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\./

Km

U1092

0

100

200m

,.

~l

Text-fig. 1. Geological route map along the Matsumoto-zawa, showing the locality (U 1092)
of the described nautiloid. Km: Kamuikotan Metamorphics, MIa, Mlb: Subdivisions of the Middle
Yezo Group (Mid-Cretaceous), T: Tertiary (Miocene). Inset at the upper right corner is the index
map.
(Y. Kanie delin.)

Explanation of Plate 62
Fig. 1. Cymatoceras sp. aff. C. virgatum (Spengler) . . . . . . . . . . . . . . . . . . . . . . . . Page 336
MNH. 145, obtained by H. Honma from loco So. 367, Chiyenaibo, on eastern coast of Cape
Soya. Lateral (a) and ventral (b) views, xL
All photos in PIs. 62 - 66 by courtesy of Dr. M. Noda.

MATSUMOTO et al. : Cretaceous Nautiloids - V

Plate 62

777. Cretaceous Nautiloids
grained sandstone of Member MIa, lower partof the Middle Yezo Group (Text-fig. 1).

339----- --....,

\

\

,
\

C. A Nautiloid Species fromthe Neocomian of Rebun Island(T. Matsumoto and Y. Ueda)

\

\
\

\

I
I

PI. 65, Fig. 1; Text-fig. 2Compare:1910. Nautilus Mikado Krenkel, Beitr. Paliiont.
Geol., Osterr.-Ungarns u. d. Orients,

vol. 23, p. 219, pI. 23, figs. 4, 5; pI. 22,
fig. 14.
Material:-GK. H5927, obtained by Y. Ueda
from Jizo-iwa, Rebun Island.
Description:-As only the right half of afragmentary whorl is preserved, the shell formcannot be exactly reconstructed. The preserved
part shows a gently inflated flank, well roundedventrolateral part and a rather flat venter. The
original whorl-section seems to be subtrapezoid
and broader than high (Text-fig. 2). The umbilicus seems to be narrowly opened.
The surface of shell is ornamented withnumerous fine ribs, which are rather flat-topped
and separated by narrower groove-like interspaces. At the whorl-heights of 65 to 75 mm 9
ribs are counted on the venter within the intervalof 30 mm. Branching of the rib takes place near
the umbilicus and also occasionally on theventrolateral part. The ribs are gently flexuous
on the ventrolateral part, forming a sinus ofmoderate sharpness (about 100-120°) on theventer.
The suture is partly exposed, showing ashallow and broad lateral lobe and an asymmetricsmall saddle near the umbilicus, passing to adescending line on the umbilical wall. The
siphuncle position is unknown.
Comparison:-The Rebun specimen is somewhat similar to C. mikado (Krenkel), from the
Neocomian (probably Barremian) of Tanganyika(Tanzania), in the numerous and fine ribbing,but it has a subtrapezoidal, rather than a semicircular, whorl-section, a flatter top of venter,

I

Cymatoceras sp. aff. C. mikado (Krenkel)

I
I

-

. '"

I

I

I

Text-fig. 2. Cymatoceras sp. aff. C.
mikado (Krenkel).
Whorl-section. Broken line: restored part.
Scale bar: 10 mm. (T. Matsumoto delin.)
and a wider umbilicus.
With respect to the shell-form, it is rather
allied to C. elegantoides (d'Orbigny) (1840, p.
69; Foord and Crick, 1890, p. 549, fig. 2),
from the Albian of England and France, but
the rib-branching is multiple in that species.
The siphuncle is subcentral and slightly
deviated to the dorsum in C. mikado, whereas
it is evidently subdorsal in C. elegantoides.
This point cannot be examined in the specimen
from Rebun.
Although some affinities and distinctions
are recognized as in the above comparison,
the present specimen is too incompletely preserved to establish a new species.
Occurrence:-From dark coloured, finegrained sandstone (wacke) exposed on a cliff,
about 100 m southeast of Jizo-iwa, on thesouthwestern coast of the island of Rebun, off
the west coast of northern Hokkaido (45°10'40"
N., 141°1'30" E.) (Text-fig. 3). The rock is
assigned to the lowest member of the 2000 m
sequence of the volcanogenic deposits called the
Rebun Group (Nagao et al., 1963). At the same
locality some ammonites, including a pulchelliid,
were found.
PART 6. AN
ADDITIONAL NAUTILOID
FROM THE SANTONIAN OF HOKKAIDO
T. Matsumoto

340

Tatsuro MA TSUMOTO et al.
study by Mr. Takemi Takahashi, to whom I am
much obliged. This has given an addition to the
Santonian nautiloids.

<>

Cymatoceras sharpei (SchlUter)
PI. 66, Fig. 1
1876. Nautilus sharpei Schliiter, Palaeontographica, vol. 24, p. 171, pI. 46, figs. 5-7.
1982. Cymatoceras sharpei (Schliiter); Immel
et al., Zitteliana, vol. 8, p. 8, pI. 1, figs.
1-2.

N

o

1km

!

Text-fig. 3. Geological outline map of thesouthern part of Rebun Island, indicating thenautiloid locality with a solid circle (e). RI-R5:
Members of the Lower Cretaceous Rebun Groupin ascending order,
T: Tertiary formations,
TV: Tertiary andesite, Q: Quaternary. Upperline of the map quadrangle: 45°20' N., left
line: 141°0' E. For the location of Rehun
Island see text-fig. 1 of Part 1 (Matsumoto,1983). [Geological outline adopted from Nagao
etal.,1963.]

After the manuscript of Part 2 was submitted to the editor of the Society, an interesting specimen was kindly provided me for thisMaterial:-A single specimen of Takemi Takahashi's Collection (TTC. 370713), from Ikushumbets.
Description:-The shell seems to be rather
small for the genus, for its diameter is about
80 mm at the end of the phragmocone andabout 110 mm at the end of the body-chamber
which is preserved for about a half whorl.
The whorl is nearly as high as broad, although
there may be some effect of secondary compression. The ventral part is moderately rounded,passing to the moderately inflated flanks. The
maximum breadth is at about the lower one
third of the whorl-height. The umbilicus is very
narrow or almost closed.
The surface of the shell is ornamented withnumerous fine ribs, which are hardly impressedon the internal mould. Rib branching is seen
frequently near the umbilical margin and occasionally on the lower part of the flank. The backward sinus of the ribs on the venter is gentle.
Suture is nearly linear, with slight sinuosity
on the main part of the flank, and descending
towards the umbilicus, showing an indistinct
saddle near the subrounded umbilical shoulder.
The septa are fairly distant.
The siphuncle position is unknown, unless
this single specimen be cut appropriately.

Explanation of Plate 63
Fig. 1. Cymatoceras sp. aff. C. virgatum (Spengler) . . . . . . . . . . . . . . . . . . . . . . . . Page 336
GK. H5726, obtained by T. Matsumoto at loco Y 632, Shiyubari, Member If of the Oyubari
sequence. Frontal (a) and lateral (b) views. Scale bar: 20 mm.

777. Cretaceous Nautiloids

341

Table 2. Measurements of Cymatoceras sharpei.

This specimen

DiameterUmbilicus

HeightBreadth

B/H

105.5

5.2 (.05)

67.0 (.64)

65.0 (.62)

0.97Comparison:-With respect to the fairly
distant, almost linear sutures and the numerous
fine ribbing, this specimen is closely allied to thelectotype* (here designated: illustrated specimenof SchlUter 1876, p. 171, pI. 46, figs. 5-7),
from the Cenomanian of West Germany. It is,however, smaller and has a less inflated whorlwith a more narrowly rounded venter. Accordingly, it is more similar to the specimens fromthe Santonian part of the Gosau Group (Austria)
described by Immel et al. (1982, p. 8, pI. 1, figs.
1-2) under C. sharpei. The difference in thewhorl-shape as compared with the lectotype
may be due partly to the secondary compression
and partly to individual variation. It could besuggested, however, that the Santonian form
might be generally smaller and less inflated
than the typical Cenomanian form.
Occurrence:-17 km point on the forestry
railway (now abandoned) along the main streamof the Ikushumbets. The strata at and aroundthis point are referred to the Santonian, as
evidenced by the co-occurrence of Hyphantoceras orientale (Yabe).

PART 7. CRETACEOUS NAUTILOIDS FROMHOKKAIDO-SUMMARYOF
RESULTS
T. Matsumoto
In this final part I attempt to summarize
concisely the results of the serial study on the
Cretaceous nautiloids from Hokkaido describedin Parts 1-6 and also Matsumoto (1967).
1. Summarized list of the described species with
their stratigraphical occurrences:Eutrephoceratinae (Nautilidae)*This was indicated as the holotype by Immel
et al. (1982, p. 8), but SchlUter evidently dealt
with at least nine specimens (i.e. syntypes).

Kummeloceras yamashitai Matsumoto(Turonian)
Kummeloceras yezoense Matsumoto(Coniacian)Kummeloceras kamuy Matsumoto et
Muramoto (Santonian)
Eutrephoceras nodai Matsumoto(Turonian)
Eutrephoceras kobayashii Matsumoto(Santonian?)
Eutrephoceras soyaense Matsumoto et
Miyauchi (Campanian)
Cymatoceratidae
Cymatoceras aff. C. mikado (Krenkel)(Barremian)
Cymatoceras cf. C. sakalavum Collignon(Albian)Cymatoceras aff. C. virgatum (Spengler)(Albian)Cymatoceras pseudoatlas (Yabe et
Shimizu) (Santonian)
Cymatoceras sharpei (SchlUter) (Santonian)
Cymatoceras pacificum Matsumoto et
Muramoto (Campanian)
Cymatoceras honmai Matsumoto et
Miyauchi (Campanian)
A nglonautilus japonicus Matsumoto et
Takahashi (Cenomanian)Anglonautilus mamiyai Matsumoto et
Miyauchi (Campanian)
2. Systematic ciassification:-The genus Kummeloceras Matsumoto has been introduced toaccommodate a group of species whose suturesare similar to but more sinuous than those of
Cenoceras but less sinuous than those of
Cimomia or Nautilus. It is regarded as representing a transitional stage in the evolutionary
development from Cenoceras to Cimomia andNautilus. It is suggested that there are several
Cretaceous (and possibly also Tertiary) species
which can be assigned to Nautilus. The typespecies (K. yamashitai) and several other exam-

Tatsuro MATSUMOTO et al.

342

think it better to rank it down to a subfamily of
the family Nautilidae, because Nautilus seems
to be intimately related to Cenoceras by way ofKummeloceras as mentioned above. Its main
stock is represented by Cenoceras and Kummeloceras, whereas Eutrephoceras (emend.) has
simplified reducing sutures. There are certain
other specialized genera (e.g., Pseudocenoceras,
Carinautilus and Obinautilus) which can be

pIes of Kummeloceras are rather high-whorled
and could possibly be regarded as directly
ancestral to Nautilus, whereas the less compressed subgroup of Kummeloceras could lead to
Cimomia. As another alternative, Nautilus
might have been derived from Kummeloceras
by way of Cimomia.
The family Eutrephoceratidae Miller, 1951
was revived in Part 1 (Matsumoto, 1983). I now
d

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NAUTILINAE

.~

+->

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I-

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_-

-- . ...... --u-- -- .. ----- ....
---"- .. _-'"--. ---.... ' " -.... ------ ... ...... .
Vl

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---- ...- .. - .----.. -_ .... ----

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Text·fig. 4.

Presumed phylogeny of the Nautilidae.

Explanation of Plate 64
Fig. 1. Cymatoceras Spa aff. C. virgatum (Spengler) . . . . . . . . . . . . . . . . . . . . . . . . Page 336
GK. H5726, ventral view. (See PI. 63 for other views.)
Fig. 2. Cymatoceras sp. cf. C. sakalavum Collignon . . . . . . . . . . . . . . . . . . . . . . . . Page 337
YCM. Ur 1092001, obtained by Y. Miyata and Y. Kanie at loco U 1092, Matsumoto-zawa,
branch of the Sakubai, a tributary to the River Kerimai, NNW of Urakawa. Lateral (a) andventral (b) views. Scale bar: 20 mm.

;::.

I

777. Cretaceous Nautiloids

included in Eutrephoceratinae in a revised sense.
Angulithes (=Deltoidonautilus) is a derivativeof Kummeloceras, acqumng more sinuous
sutures and triangular whorl-sections. K. yezoense, for example, represents a transitional statefrom Kummeloceras to Angulithes.
The sutures of the recent species of Nautilus
are as sinuous as (or somewhat more sinuousthan) those of many species of Cimomia, which,in turn, has been regarded as ancestral to Hercoglossa and other related genera. Accordingly
the Hercoglossinae falls in a synonym of Nautilinae.
The presumed phylogeny of the discussed
genera is shown in Text-fig. 4. This is tentative
and two alternative interpretations are indicatedby different lines of descent. If one of them beproved by sufficient evidence, the subdivisionof the Eutrephoceratinae and the Nautilinae
might become unwarrantable.
Some genera of the Cymatoceratidae havebeen discussed (mainly in Part 2), without
giving a great alteration. Anglonautilus is well
represented in Japan by the three species fromthe Aptian (outside Hokkaido), the Cenomanianand the Campanian respectively. Descriptions
in details have been given in Part 4 on a species
from the Lower Cenomanian of the Ikushumbets area, on the basis of well-preserved specimens of various growth-stages. It shows theintimate relationships of Anglonautilus with
Cymatoceras and Paracymatoceras.
3. Biostratigraphical implications:-So far as theavailable material from the Cretaceous of Hokkaido is concerned, every stage has species which
are particular to it. In other words, no species
from Hokkaido shows a long stratigraphicalrange which goes through more than one stage.
This might have come from the collection
failure or from the environmental conditions
and the apparent range may be local or partial.
There are, however, examples whose actual
occurrences are stratigraphically restricted andgeographically extensive. Cymatoceras pseudatlas, for instance, occurs in the Lower Santonian
Zone of Inoceramus amalwsensis in Hokkaido
and Kyushu. It may be reckoned as one of themembers in the assemblage of that zone.

343

Aside from the distribution in the Japaneseprovince, certain species are closely allied to thecontemporary species in other provinces. Cymatoceras aff. C. mikado, C. cf. C. sakalavum
and C. aff. C. virgatum, as well as C. sharpei(Gosau form) are such examples. C. aff. C.
virgatum is also somewhat similar to C. carlottense (Whiteaves), from the Albian on the
Pacific side of Canada.
4. Palaeobiogeography:-The three species fromthe Lower Cretaceous of Hokkaido (describedin Part 5) seem to have a close affinity with
Cymatoceras mikado (Krenkel), from the Barremian of eastern Africa, C. sakalavum Collignon,from the Albian of Madagascar and C. virgatum(Spengler), from the Albian of southern India,
respectively. The differences of our forms fromthe original types could possibly be proved subspecific, if sufficient material were obtained inboth provinces. The original species are all
distributed on the coastal area of the Cretaceous
Indian Ocean which had free connection withthe eastern Tethys and the western Pacific (seereconstructed palaeogeographic maps by Barron
et al., 1981).
Aside from Cymatoceras sharpei (SchlUter),eleven species from the Upper Cretaceous of
Hokkaido are all seemingly endemic, as none ofthem has been known outside the Japaneseprovince of the Late Cretaceous times.
In general, many genera of the Cretaceous
nautiloids show worldwide distribution, but at
specific level there seem to have been provincial
differences. For instance, very few species arecommon between western Europe and North
America, although the Cretaceous Atlantic
Ocean must have been much narrower than the
Recent one. This is a remarkable contrast to thefact that certain species of the Ammonoidea
had really worldwide distribution.
There is much to be worked out to understand the true implications of the above facts.
5. Palaeoenvironments:-I can state little about
the palaeoenvironments of the Cretaceous
nautiloids. Although the life history of therecent species of Nautilus is being studied intensively (e.g., Hirano et al., 1982), how to applythat knowledge to Mesozoic nautiloids may

344

Tatsuro MATSUMOTO et al.

involve another problem. Since the living Nautilus is so to speak a relict at the end of thelong-continued evolutionary history of the
Nautiloidea, it may have some peculiarity in its
mode of life. Anyhow, I give here concisely thefollowing notes through the study of the Cretaceous nautiloids from Hokkaido.
(1) The nautiloid fossils have been obtained
from the strata in which ammonites and inoceramids occur commonly.
(2) The body-chamber is preserved in manyof the studied specimens, although it may be
incomplete in some cases.
(3) A nglonau tilus, a truly ribbed nautiloid,
occurs in the sediments of near shore, shallow
sea environments. For example, numerous
specimens of A. japonicus were embedded in
the sandstone of shallow open sea facies inwhich ornated ammonites such as Mantellicerasand Hypoturrilites occurred commonly. A.
mamiyai was found in a nodule which contained
drifted fragments of land plants.
(4) Eutrephoceras soyaense was found fairly
commonly in a bed of near shore, shallow open
sea facies in which the nodules contain abundantly drifted fragments of land plants. In
North America and other regions, E. dekayi
and other related species, which are characterized
by globose shells with inflated whorls andnearly linear sutures, occur also in a shallow
shelf facies.
(5) As to Kummeloceras only the generalaccount mentioned in (1) is applicable and noparticular point has been noted.
(6) Cymatoceras has many kinds of species,
showing varieties of shell-form, ribbing andsutural sinuosity. Accordingly, there musthave been a considerable difference in the
habitat and mode of life of Cymatoceras froma species to another. Examples from Hokkaido
seem to support this general statement, but

it is still difficult to give precisely an ecologicalinterpretation on each of the described species.
Incidentally, no example of Cymatoceras has
been known from the Turonian of Hokkaido,
whereas specimens of Kummeloceras have beenobtained not rarely from that stage. Whether
this is due to collection failure and just accidental or has some ecological meaning is aquestion to be answered in the future.
In conclusion, the serial papers in Parts 1 to7 have given the present knowledge as to whatkinds of species occur in what strata of whichages in Hokkaido. This is, however, a start for
further study which should be developed by
the cooperation of zealous naturalists.
References Cited(For brevity the references listed in Parts 1-4
are not repeated here.)
Barron, E. J., Harrison, C. G. A., Sloan II, J. L.
and Hay, W. W. (1981): Paleogeography,
180 million years ago to the present. Eclogae
Geol. Helv., vol. 74, p. 443-470, pis. 1-9.
Collignon, M. (1949): Recherches sur les faunes
Albiennes de Madagascar. I.-L'Albien
d'Ambarimaninga. Ann. Geol. Servo Mines,
Madagascar, fasc. no. 16, p. 1-128, pIs.
1-22.
Foord, A. H. and Crick, G. C. (1890): A revisionof the group of Nautilus elegans J. Sowerby,
Geol. Mag., new ser., decade III, vol. 7,
p.542-552.
Hirano, H., Fukuda, Y., Tanabe, K. and Obata,I. (1982): Fundamental study on ammonite
palaeobiology Part I-Recent progress of thestudy on living Nautilus. Jour. Geol. Soc.
Japan, vol. 88, p. 725-735 (in Japanesewith English abstract).
Immel, H., Klinger, H. C. and Wiedmann, J.
(1982): Die Cephalopoden des Unteren
Senon der Gosau von Brandenberg/Tirol,
Osterreich. Zitteliana, vol. 8, p. 3-32, pIs.
1-11.

Explanation of Plate 65
Fig. 1. Cymatocems sp. aff. C. mikado (Krenkel) . . . . . . . . . . . . . . . . . . . . . . . . . Page 339
GK. H5927, obtained by Y. Ueda from Jizo-iwa, lowest member of the Rebun Group(Neocomian), Rebun Island. lateral (a) and ventral (b) views. Scale bar: 20 mm.

MA TSUMOTO et at.: Cretaceous Nautiloids - V

Plate 65

777. Cretaceous Nautiloids
Krenkel, E. (1910): Die untere Kreide vonDeutsch-Ostafrika. Beitr. Palciont. Ceol.
Osterr.-Ungarns u. d. Orients, vol. 23, p.
201-250, pis. 20-23.
Matsumoto, T. (1942): Fundamentals in the
Cretaceous stratigraphy of Japan Part 1.

345

Mem. Fac. Sci. Kyushu Imp. Uniu., Ser. D,
vol. 1, no. 3, p. 129-280, pis. 5-19.
Nagao, S., Akiba, C. and Omori, T. (1963):
Rebunto. Expl. Text Ceol. Map Japan,
scale 1 :50,000, Geol. Surv. Hokkaido, p.
1-43, 1 map (in Japanese with English
resume).

Chiyenaibo '1-=71 of, Hikage·no-sawa fl ~1i(l)iR, Ikushumbets [Ikushunbetsu] ~lf5Jlj, Jizo·iwa:It!llil¥i", Kerimai J1tl., Matsumoto-zawa t~*iR, Mitsuishi :=':5, Oyubari :;!;:671.!t, Rebun :tLx,
Sakubai -tt· (l /~ 1, Soya <;;'i:fr, Shiyubari :l::67iJN, Urakawa im{IiJ, Yezo j!j!;l9n = :/J~t1iiJ):fi!¥RiIf!*c* 1/ A 111 M- V. -t(l) 5-7 ~fHJ', -t(l) 5. "fflilsilf!*i&[l1Jmo

A.

*:frc :k!Y'lJtgjf(l) 1 f;1!:

Cymatoceras sp. aff. C. virgatum (Spengler). 1:/ 1-- (l) 7 II.-

~7:/~.~.*K.W~~~, • • OO~-thfi~hfWHb;IK~<, ~~2~_~0~I),~~b; 3 ~j8(~reOb Ght.i:~ ' 0 *671.!t(l)f:t. 7 II.- \:-7:/ ; *:fr (l) (i lI'1tL*l:Eo (t~*~~~· ~~\r"Jfit{ijIt)

B. r~jiiiJ:Illi~i.t(l) 1:flIi: Cymatoceras sp. d. C. salwlavum Collignon. !m{IiJ~tkilli 17 km(l).ra.JIIs!i::iJlE(l) 1 :It!l,~L 4'ffll=:/(l) Mia 1tH":'ito =i'7(~tf}L l:l '~b;"' 31"7l.7-.;t; II.-Jf!j7 II.- \:'7 :/it(l).8C.fll!t.::JJlfJ.Jo (t~*)l~~· MiIlift)'t· '8m!lt-~~)C. :tU(~ * ::! ~ 7:/ it 1 :fil: Cymatoceras sp. aff. C. mikado (Krenkel). 31:/ -tT =- 7(l)**::! ~ 7 :/it(l)l:.j[IH.cJ~~tJ;, ~JV:J~llfrllilb;*r;Mf1cA.-t{,J1\~'o :Illi~¥i":JIim(l):tLxfflilM:/&"f liIIfflil ito (t~*~~~· MJjrij~~)-t(l) 6. -tj-:/ I- =- 7:/ ltQ)~:l.m 1 :fill:: Cymatoceras sharpei (SchlUter). 1--1 ')I Q) -t / "' =7 :/ it~.*.t I) fttJ' L 0 * -.7-. I- ~ 7 Gosau fflilf,'f-+j-:/ I- =- 7:/ b, G:I&::JIi$IH5'Q) (, (l) v;:,.t <l.cJ~o ~~*5JIJJlIl::lJlE·tj-:/ I- =- 7:/.t I) (,ij:j1.ijji~~):\;l**o (t~*~tl~)-t(l)7. ~twjfii&S!lli;j;C*?A;t;'1m JJX:*(l)j~Hf:Jo (1)i1c~Lt.::15,flH:~~, i&t±lfflil(l)llif~~
vr~C. (p. 341). (2) Kummeloceras ~>a:~ftL, *~n'-~IH:''')~'l:1fr~J!~~~ (m4~(l)*~f!t~Jl!:\). (3)1}1lM',::'cl:.~tJ:~f:il!;/J~i&L-l::t-oI), 9=>IU:t.11:::5*mJili;:fill:Q),,( ://~-t.::l)f4~{,(l) {, i5 ~ 0 iwJi·mW:flTi c (l)M Jt (l) c' ~ ~ {, (l) {, i5 ~ 0 (4)i11:.~:It!lJlll., (5)i1~ml:' ")~, l: {"T (l)"i c Ob ~ili\.;z.,. t.::o (t~*~tlll)**'

Postscript. The generic name Kummeloceras Matsumoto, 1983 was preoccupied by Kummeloceras
Shimansky, 1967 (Bull. Palaeont. Inst. Acad. Sci. USSR, vol. 115, p. 121), which was proposed
for K. sibiricum Shimansky, from the Permian of Siberia and referred to the family Temnocheilidae
with a query. I hereby propose Kummelonautilus to replace Kummeloceras Matsumoto, 1983
(this series, part 1, p. 12). I sincerely thank Dr. V. N. Shimansky for his kind information about
the work in the Soviet Union.
(Tatsuro Matsumoto)346

Tatsuro MATSUMOTO et al.

Explanation of Plate 66
Fig. 1. Cymatoceras sharpei (SchlUter) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Page 340
TTC. 370713, collected by T. Takahashi from 17 km point along the main stream of theRiver Ikushumbets. Two lateral (a, b) frontal (c) and back (d) views, x 1.

MATSUMOTO et al.: Cretaceous Nautiloids - V

Plate 66

Trans. Proc. Palaeont. Soc. Japan, N.S., No. 134, pp. 347-360, pI. 67, July 30, 1984

778. A VISEAN BRACHIOPOD FAUNA FROM THE MANO FORMATION,
SOMA DISTRICT, ABUKUMA MOUNTAINS, NORTHEAST JAPAN*
JUN-ICHI TAZAWA, YUKIO GUNJI and KEI MORI
Institute of Geology and Palaeontology,
Tohoku University, Sendai 980

Abstract. An Early Carboniferous (Late Visean) brachiopod fauna is describedfrom the top of the Mano Formation of the Abukuma Mountains, northeast Japan.
It includes the following species: Leptagonia analoga, Rugosochonetes cf. celticus,
Yanishewskiella cf. angulata, Spirifer sp., Brachythyris sp., Syringothyris sp. andTylothyris laminosa.
The Late Visean age assignment for this fauna matches well with the stratigraphic data suggesting that the Mano Formation is overlain conformably by theUpper Visean Tateishi Formation.

Introduction
140'51'The Carboniferous brachiopods described
below were collected at a locality in Tateishi (lat.
37°42'45"N, long. 140°50'39"), Kashima-cho,Soma-gun, Fukushima Prefecture, i.e. the Soma
district in the Abukuma Mountains of northeast
Japan (Text-fig. 1). They came from sandstone
in the uppermost part of the Mano Formation.
The Mano Formation named by Sato in 1956,
was regarded as Early Carboniferous (Sato,
1956, 1974) or Tournaisian (Minato, 1960;
Konishi, 1963; Minato, Gorai and Hunahashi,
1965; Ichikawa, Fujita and Shimazu, 1970;
Yoshida, 1977; Minato and Kato, 1978; Minato
et al., 1979) in age without firm evidence.
Until now only some species of brachiopods,a species of rugose coral and a species of trilobite
were recorded from this formation (Sato, 1956,
1974; Minato et a!., 1979), but they were notdescribed or illustrated.
In this paper the brachiopods of Tateishi
are described and the age of the fauna is discussed. Stratigraphic relationship between the140'53'//,.."-"/--"/(':::"",
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HARAMACHI CITY

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U'4'140' 3Text-fig.1. Map showing the fossil locality(X) and the Soma district.

*Received April 13, 1983.
347

Jun-ichi TAZA WA et al.

348

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21

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16

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,~----~~----~'

p; __ '00
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A-__________.....---.4

0-+++++-.00-++ + ++,o~ -B
400100-++_0

Text-fig. 2. Geologic map of the Soma district. 1-4: Matsugadaira Formation (Pre-Upper
Devonian; 1: Mafic schist and phyllite; 2: Pelitic schist and phyllite; 3 : Chert; 4 : Metabasite);
5: Ainosawa Formation (Upper Devonian); 6 : Mano Formation (Lower Carboniferous); 7: Tateishi
Formation (Lower Carboniferous); 8-11 : Uwagaya Formation (Lower Permian; 8: Shale; 9:
Alternation of sandstone and shale ; 10: Sandstone ; 11: Limestone); 12-14: Oashi Formation(Middle Permian; 12 : Sandstone; 13 : Alternation of sandstone and shale; 14 : Acidic tuff); 15 :Yumiorezawa Formation (Upper Permian); 16: Conglomerate of the Uwagaya, Oashi and Yumiorezawa Formations; 17: Neogene Tertiary; 18: Granitic rocks ; 19: Dyke rocks; 20: Dip and strikeof bedding plane; 21: Dip and strike of bedding schistosity .

778.

I

Visean brachiopod Fauna from Soma

Mano Formation and the overlying Tateishi
Formation is also mentioned. This has been oneof the most important problems for the Carboniferous stratigraphy and geohistory of the
Abukuma Mountains, since Minato (1955) notedthat there is a sharp angular unconformity underthe base of the Tateishi Formation.
In the present study description of the strati·
graphy was prepared by Gunji, Mori and Tazawa
while the systematics and correlation were prepared by Tazawa. The brachiopod specimens are
stored in the Institute of Geology and Palaeontology, Faculty of Science, Tohoku University.

I
50 m

I

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Stratigraphy

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349

I: :'. ' -.1 6.......... ....
. . . . . ....

Text-fig. 3. Columnar section of
the Lower Carboniferous of the Soma
district, showing the stratigraphic position of the fossil locality shown on index
map, see Text-fig. 1. 1: Massive limestone; 2: Bedded limestone; 3: Sandy
limestone; 4: Conglomerate; 5: Sandstone; 6: Alternation of sandstone andshale; 7: Acidic tuff.

7

The Mano Formation crops out in two areas,around Tateishi and east of Oashi, both in the
Soma district (Text-fig. 2). In the former, theformation forms a synclinorium plunging with
20°-30° to the SSW. It consists of fine-grainedsandstone and shale in alternation, with several,thin, acidic tuff intercalations, about 125 m intotal thickness (Text-fig. 3). The Mano Forma·tion overlies with probable conformity the
Ainosawa Formation, and is in turn overlain
conformably by the Tateishi Formation. Fossils
are rare in the Mano Formation, except for the
top, 5 m thick, massive, dark grey, fine-grained,
calcareous sandstone, in which there are rather
numerous brachiopods, bryozoans and crinoids.
The underlying Ainosawa Formation (125 m
thick) consists mostly of tuffaceous shale andyields Cyrtospirifer. It has been correlated withthe upper part of the Tobigamori Formation(Upper Famennian) of the southern Kitakami
Mountains, northeast Japan by brachiopods(Hayasaka and Minato, 1954; Minato and Kato,
1979) .
The overlying Tateishi Formation consistingof dark-grey to black limestone, 130 m in thickness, has been correlated with the Upper Visean
Onimaru Formation of the southern Kitakami
Mountains, based on the similarities of lithofacies and the coral fauna (Sato, 1956). In thepresent work, we also collected some rugose
corals such as Lonsdaleia sp., Cyathaxonia sp.
and Dibunophyllum cf. D. biparticum (M'Coy)
from the limestone.

350

Jun-ichi TAZAWA etal.
Age of the Fauna

Subfamily Leptaeninae Hall and Clarke, 1894
Genus Leptagonia M'Coy, 1844

The brachiopods of Teteishi described herein,
and the number of specimens are shown as
follows:

Leptagonia analoga (Phillips) . . . . . . . . . 4
Rugosochonetes cf. R. celticus
Muir-Wood rev. Brand . . . . . . . . . . . . 1
Yanishewskiella cf. Y. angulata(Yanishevsky) . . . . . . . . . . . . . . . . . . 1
Spirifer sp. . . . . . . . . . . . . . . . . . . . . . 3
Brachythyris sp. . . . . . . . . . . . . . . . . .. 1
Syringothyris sp. . . . . . . . . . . . . . . . . . 5
Tylothyris laminosa (M'Coy) . . . . . . . . . 8
Among these species, L. analoga and T.
laminosa are long ranging forms; both occur inTournaisian to Visean rocks. However, thegenus Yanishewskiella is restricted to the Upper
Visean and Namurian (Winkler Prins, 1982).
Spirifer (s.s.) occurs mainly in the Visean(Thomas, 1971). Rugosochonetes is common in
the Visean to Namurian (Muir-Wood, 1962;
Afanasjeva, 1975). The Rugosochonetes from
Tateishi is close to R. celticus Muir-Wood (rev.
Brand) from the Upper Visean and Namurianof Britain. Syringothyris sp. differs from theTournaisian forms; it resembles S. fontanalis
Roberts from the Upper Visean of northwestern
Australia.
Stratigraphically, the sandstone yielding thebrachiopods is apparently lower than the limestone of the Upper Visean Tateishi Formation.
The age of the brachiopod fauna does, therefore,
not range up into Namurian.
The Tateishi fauna from the top of the Mano
Formation can be estimated to be of Visean age,probably Late Visean.

Description of Species
Order Strophomenida Opik, 1934
Suborder Strophomenidina Opik, 1934
Superfamily Strophomenacea King, 1846
Family Leptaenidae Hall and Clarke, 1894

Leptagonia analoga (Phillips, 1836)
PI. 67, Figs. 2-4

Producta analoga Phillips, p. 256, pI. 7,
fig. 10.
1842-1844 Leptaena depressa var. analoga(Phillips): de Koninck, p. 215, pI. 13,
figs. 6a-6c.
1844 Leptagonia analoga (Phillips): M'Coy,
p.117.
1861 Strophomena rhomboidalis var. analoga(Phillips): Davidson, p. 119, pI. 28, figs.
1-6,9-13.
1863 Strophomena analoga (Phillips): Davidson,
p. 173, pI. 9, fig. 18.
1872 Strophomena rhomboidalis var. analoga(Phillips): Etheridge, p. 333, pI. 18, fig. 1.
1900 Leptaena analoga (Phillips): Frech (inFrech and von Arthaber), p. 200.
1909 Leptaenarhomboidalis(Wilckens):
Sommer, p. 626, pI. 29, figs. 14, 14a.
1909 Leptaena rhomboidalis (Wilckens): Weller,
p. 292, pI. 12, figs. 2,3.
1913 Plectambonites rhomboidalis var. analoga(Phillips): Mansuy, p. 31, pI. 5, fig. 3.
1914 Leptaena analoga (Phillips): Weller, p. 49,
pI. 2, figs. 1, 2, 5-10 only.
1916 Leptaena analoga (Phillips): Frech, p.
237, pI. 2, figs. 2, 3d.
1920 Leptaena analoga (Phillips): Girty, pI. 54,
fig. 3.
1924 Leptaena analoga (Phillips): Tolmatchoff,p. 209, pI. 13, fig. 8.
1927 Leptaena analoga (Phillips): Girty, pI. 22,
figs. 6-8.
1934 Leptaena analoga (Phillips): Demanet, p.
61, pI. 5, figs. 1-14; text-figs. 11-14.
1938 Leptaena analoga (Phillips): Branson, p.
24, pI. 5, fig. 31.
1951 Leptaena analoga (Phillips): Minato, p.
361, pI. 3, fig. l.
1952 Leptaenella analoga (Phillips): Sokolskaja(in Sarytcheva and Sokolskaja), p. 36, pI.
3, fig. 18.
1954 Leptaena cf. L. analoga (Phillips): Max·
well, p. 33, pI. 4, figs. 1-3.
1957 Leptaenella cf. L. analoga (Phillips):
Campbell, p. 41, pI. 13, figs. 20,21.

1836

778.
1958

1961
1962
1963

1966

1968

1970
1971

1971

1973
1974
1974
1975

1976
1977

1978
1978
1979
1979

Visean brachiopod Fauna from Soma

Leptagonia cf. L. analoga (Phillips):
Cvancara, p. 860, pI. 110, figs. 6-13;
text-figs. 3, 4.
Leptaena analoga (Phillips): Nelson, pI. 4,
fig. 26.
Leptaena analoga (Phillips): Armstrong,
pI. 6, fig. 25.
Leptaenelia analoga (Phillips): Sokolskaja(in Sarytcheva et al.), p. 80, pI. 3, figs.
9-14.
Leptaenelia analoga (Phillips): Gretchisch·nikova, p. 94, pI. 1, figs. 19-20; pI. 2,
figs. 1-6.
Leptagonia analoga (Phillips): Brunton,
p. 29, pI. 3, figs. 26-31; pI. 4, figs. 1-9;
text-figs. 6-17.
Leptaenelia analoga (Phillips): Abramov,
p. 108, pI. 1, figs. 11, 12.
Leptagonia analoga (Phillips): Bublitschenko, p. 37, pI. 3, figs. 1-5.
Leptagonia analoga (Phillips): Thomas,
p. 30, pI. 18, figs. 1-8; text-figs. 11a11d.
Leptaenella analoga (Phillips): Nalivkin
and Fotieva, p. 20, pI. 1, figs. 9-13.
Leptagonia analoga (Phillips): Kalashnikov, p. 23, pI. 3, figs. 5a, 5b.
Leptagonia distorta (Sowerby): Jing andLiao, p. 277, pI. 142, figs. 20-22.
Leptagonia analoga (Phillips): Litvinovich,
Aksenova and Martynova, p. 53, pI. 16,
fig. 1I.
Leptagonia analoga (Phillips): Bublitschenko, p. 22, pI. 1, fig. 10.
Leptagonia analoga (Phillips): Yang, Ni,
Chang and Zhao, p. 316, pI. 131, figs.
2a,2b.
Leptagonia analoga (Phillips): Tong, p.
213, pI. 77, figs. 7a-7c.
Leptagonia distorta (Sowerby): Feng andJiang, p. 237, pI. 86, figs. 3, 4.
Leptagonia analoga (Phillips): Nalivkin, p.
18, pI. 3, figs. 1-3, 5,6.
Leptagonia distorta (Sowerby): Jing, Ye,
Xu and Sun, p. 76, pI. 22, figs. 10-12.

Material:-Four specimens: (1) external andinternal moulds of a brachial valve, IGPS colI.
cat. no. 97912; (2) external moulds of threebrachial valves, IGPS coll. cat. nos. 9791397915.

351

Description :-Brachial valve wider, trapezoidal
in outline, with the greatest width at hinge line,
flat to slightly concave on visceral disc, surrounded by a deep, rounded depression at theanterior and lateral margins of visceral disc,strongly geniculated nearly at right angles justanterior to the depression, with a trail about
10 mm long; cardinal extremities blunt, angular;
length 23 mm, width 33 mm in the best preserved specimen (IGPS coll. cat. no. 97913).
External ornament of brachial valve consisting of irregular, strong, concentric rugae andnumerous, fine capillae on visceral disc; 17
rugae and numerous capillae with a density of
18 to 20 per 5 mm at the mid portion of visceraldisc; rugae lacking on trail.
Dorsal internal structure unknown, except
for a median septum and some vascular markings
partly and faintly preserved in a specimen
numbered IGPS coll. cat. no. 97912.
Remarks:-The Soma specimens can be
identified with Leptagonia analoga (Phillips,
1836) by their medium-sized, wider trapezoidshaped brachial valves with subangular cardinal
extremities, a deep, rounded depression at theantero-Iateral margins of the visceral disc, andthe external ornament of irregular concentricrugae and numerous capillae. The dorsal depression which bounded the visceral disc wasnoted by Campbell (1957, p. 41) on the Australian specimens.
L. analoga has been reported from the Tournaisian and Visean of Europe, Asia, North
America and Australia. It seems to be worthy
to note that this species occurs commonly inthe Visean of Britain (Muir-Wood, 1948; Brunton,1968), Germany (Sommer, 1909), the Soviet
Union (Abramov, 1970; Kalashnikov, 1974),
China (Jing and Liao, 1974; Feng and Jiang,1978; Jing, Ye, Xu and Sun, 1979), Laos
(Mansuy, 1913) and eastern Australia (Campbell,
1957; Cvancara, 1958).

Suborder Chonetidina Muir-Wood, 1955
Superfamily Chonetacea Bronn, 1862
Family Chonetidae Bronn, 1862

352

Jun-ichiTAZAWA etal.

Subfamily Rugosochonetinae Muir-Wood, 1962
Genus Rugosochonetes Sokolskaja, 1950

Rugosochonetes cf. R. celticus
Muir-Wood, 1962
rev. Brand, 1970
PI. 67, Figs. 1a-1c

Compare:1962 Rugosochonetes eel tic us Muir-Wood, p.
1968
1970

68, pI. 7, figs. 3,4,5,7,8 only.
celtic us Muir-Wood:
Brunton, pI. 7, figs. 28-31; pI. 8, figs.
1-9.
Rugosochonetes celticus Muir-Wood pars:Brand, p. 108, pI. 10, figs. 12-20; pI. 13,
figs. 5, 6, 9.

Rugosochonetes

Material:-One specimen, internal mould of apedicle valve with the external mould of theposterior portion attached, IGPS colI. cat. no.
97916.
Description:-Shell large size for the genus,
semicircular in outline, with the greatest width
at hinge line; length 14 mm, width 13 mm.
Pedicle valve moderately convex, with ashallow sulcus on the anterior half of the valve.
Ventral interarea well developed, 1.2 mm high at
both sides of delthyrium; delthyrial angle 37°.
In the pedicle valve, a median septum sharp
and high in the posterior portion, rather bluntand low in the anterior portion, totalling 5.9mm long. Teeth massive, subtrigonal, demarcatedfrom interarea by grooves. A pair of smooth,elongate adductor scars at the postero-median
portion of the valve. Diductor scars large, flabellate, radially striated on the surface, occupying
half the area of visceral disc and bounded by apair of ridges of about 3.5 mm long. Papillae
strong over ear regions and the posterior halvesof lateral slopes.
Remarks:-This specimen is quite similar tothe pedicle valve of Rugosochonetes celticus
Muir-Wood, 1962 pars, figured by Brand (1970,
pI. 10, fig. 12) from the Upper Visean of Scotland, in size, shape and internal features of thepedicle valve.
R. celticus was proposed by Muir-Wood in

1962 for materials from the Upper Visean andNamurian of Britain. Subsequently, the typespecimens were examined by Brand (1970), in
which he noted that only those paratypes from
Flint, North Wales and Beith, Scotland areconsidered to be conspecific with the holotype,
and the majority of the other para types are
determined as Rugosochonetes speciosus (Cope,
1938).
Rugosochonetes sp. from the top of the
Karaumedate Formation (Upper Visean) in
the Nagasaka district, southern Kitakami Mountains, northeast Japan (Tazawa, 1980, p. 361,
pI. 41, figs. la, 1b) differs from the present
form in its more transverse shell outline andhaving a pair of prominent vascular trunks inthe pedicle valve.
In southwest Japan two Rugosochonetes
species have been known: Rugosochonetes sp.
from the Lower Visean of Hina, Okayama
Prefecture (Hase and Yokoyama, 1975, pI. 18,
figs. 2, 3) and Rugosochonetes aff. R. hardrensis(Phillips, 1841) from the Lower Namurian of
Akiyoshi, Yamaguchi Prefecture (Yanagida,
1965, p. 115, pI. 27, figs. 2-8). But the comparison to these forms is difficult, because
they are lacking information on the ventral
internal structure.
Order Rhynchonellida Kuhn, 1949
Superfamily Rhynchonellacea Gray, 1848
Family Tetracameridae Likharev, 1956
Genus Yanishewskiella Likharev, 1957

Yanishewskiella cf. Y. angulata(Yanishevsky,1910)
PI. 67, Figs. 5a, 5bCompare:1918 Goniophoria
1965
1974

angulata
Yanishevsky:
Yanishevsky, p. 67, pI. 6, fig. 27; pI. 7,
figs. 9, 10.
Yanishewskiella angulata (Yanishevsky):
McLaren, p. H589, figs. 466 2a-2c, 467.
Janishewskiella(?) angulata (Yanishevsky): Kalashnikov, p. 100, pI. 38, fig. 8.

778.

353

Visean brachiopod Fauna from Soma

Material:-One specimen, external mould of abrachial valve, IGPS coli. cat. no. 97917.
Remarks:-This specimen is characterized byits small, rhynchonelliform shell (length 21 mm,
width 16 mm) with a few, broad and subrounded
costae (7 costae are counted) on the brachialvalve.
The Soma specimen resembles shells of
Yanishewskiella angulata (Yanishevsky, 1910)
described or figured by Yanishevsky (1918),
McLaren (1965) and Kalashnikov (1974) fromthe Upper Visean and Lower Namurian of theUrals and Fergana in size, shape and external
ornament of the brachial valve. But the poor
preservation of this material makes the accuratecomparison difficult.
Yanishewskiella? sp., from the uppermost
part of the Karaumedate Formation of the Kitakami Mountains (Tazawa, 1980, p. 364, pI. 42,
figs. la-4), can be easily distinguished fromthe present form by its smaller dimensions.

Order Spiriferida Waagen, 1883
Suborder Spiriferidina Waagen, 1883
Superfamily Spiriferacea King, 1846
Family Spiriferidae King, 1846
Subfamily Spiriferinae King, 1846
Genus Spirifer Sowerby, 1816
Spirifer sp.

PI. 67, Figs. 12-14
Material:-Three specimens: (1) internalmoulds of two pedicle valves, IGPS coli. cat. nos.
97918, 97919; (2) external cast of a brachialvalve, IGPS coli. cat. no. 97920.
Description :-Shell small size for the genus,
wider sub quadrate in outline. Hinge line a little
less than the greatest width of shell, the latteroccurring at anterior to hinge line but slightly
posterior to midvalve. Cardinal extremities
rounded. Length about 30 mm, width 54 mm,
width of hinge line 44 mm in the largest specimen (IGPS colI. cat. no. 97918).
Both valves moderately convex in lateralprofile, with maximum convexity posterior tomidvalve, having a shallow sulcus and a low
dorsal fold. Pedicle valve ornamented by numerous costae and irregular, strong concentric rugae
on lateral slopes and sulcus; costae often bifurcating; 24 costae on each of lateral slopes, 12
costae on sulcus. Brachial valve with external
ornament of numerous, simple or bifurcating
costae and irregular, concentric rugae; 24 or 25
costae on each lateral slopes, 11 costae on fold.
Pedicle valve interior with a heart-shaped
muscle field situated at just anterior to theumbonal region. The muscle field radially andlongitudinally striated, divided by a medianridge (myophragm) posteriorly on two-thirdsof the length of the field. Other internal structure obscure owing to thick apical callus.
Remarks:-In the present material, the external ornament of the pedicle valve is faintly
preserved on the surface of the internal mould.
These specimens are referred to the genus
Spirifer (s.s.) Sowerby, 1816 on account of theirlarge, transverse shells with hinge line slightly
shorter than the greatest width, rounded cardinal extremities, numerous bifurcating costae
on the lateral slopes and the sulcus, and theabsence of long dental plates or a median septumin the pedicle valve.
The Soma specimens resemble shells of
Spirifer crassus de Koninck, described andfigured by de Koninck (1842-1844, p. 262,
pI. 15, fig. 5) from the Visean of Belgium, andby Davidson (1858, p. 25, pI. 6, figs. 20-22;
pI. 7, figs. 1-3) from the Visean of Ireland in
shape and external ornament, but the European
specimens are much larger in size.
Spirifer cf. S. crassus de Koninck, from the
Visean(?) of ami, central Japan (Hayasaka,
1924, p. 43, pI. 6, fig. 6), differs from thepresent form in its smaller dimensions andmore rectangular shell outline.
Family Brachythyrididae Fredericks, 1919 (1924)
Genus Brachythyris M'Coy, 1844
Brachythyris sp.

354

Jun-ichi TAZA WA et al.
PI. 67, Fig. 15

PI. 67, Figs. 6-7

Material:-One specimen, internal mould ofa brachial valve, IGPS colI. cat. no. 97921.
Description:-Brachial valve wider suboval in
outline, slightly convex except for the strongly
con vexed posterior portion and highly elevatedmedial portion. Hinge line much shorter than
the greatest width of the valve; the latter occurring at about midvalve. Cardinal extremities
rounded. Fold high and broad anteriorly. Dimensions of the brachial valve: length about 31 mm,
width 36 mm, width of hinge line 24 mm.
External ornament of the brachial valve
consisting of strong costae and very fine growthlines. Costae broad, rounded and simple; 10
costae on lateral slopes. Numerous, very fine
growth lines on the whole surface of the valve.
None of the internal structure of the brachialvalve observed.
Remarks :-This specimen is assigned to thegenus Brachythyris M'Coy, 1844 on the basis of
its suboval shell outline and broad, simple
costae on the brachial valve, but is too poorly
preserved for specific determination. In thepresent material the external ornament of thebrachial valve is impressed on the surface of
the internal mould.
In size and shape the Soma specimen resembles the brachial valve (holotype) of Brachythyris pseudovalis Campbell, 1957, figured byCampbell (1957, pI. 14, fig. 13) from the Upper
Visean of Watts, Babbinboon, New South Wales.
Brachythyris kitakamiensis Minato (1951, p.
368, pI. 3, fig. 3; pI. 4, fig. 1) from the Hikoroichi Formation of Choanji in the Hikoroichi
district, southern Kitakami Mountains differsfrom the present form in its larger, more transverse shell and more numerous costae (14 costae)on the brachial valve.

Material:-Five specimens: (1) external andinternal moulds of a pedicle valve, IGPS colI.
cat. no. 97922; (2) external moulds of two
pedicle valves, IGPS colI. cat. nos. 97923, 97924;(3) internal mould of a pedicle valve, IGPS colI.
cat. no. 97925; (4) external and internal moulds
of a brachial valve, IGPS colI. cat. no. 97926.
Remarks :-These specimens are assigned to
the genus Syringothyris on the basis of theirsimple costae, deep, smooth sulcus, the presenceof long dental plates and a syrinx, and theabsence of median septum in the pedicle valve.
The Soma species is characterized by its small
shell size (width about 60 mm in the largest
specimen, IGPS colI. cat. no. 97923) and thelow ventral interarea (about 7 mm high).
In size and shape of the shell, Syringothyris
sp. from Soma closely resembles Syringothyris
fontanalis Roberts (1971, p. 249, pI. 54, figs. 518) from the Upper Visean of the Bonaparte
Gulf Basin, northwestern Australia. But accessary costae on sulcus, a characteristic of S.
fontanalis, cannot be observed in the present
material.
The Soma species has an exterior resemblance
to Syringothyris japonica Tachibana (1969, p.
22, pI. 3, figs. 1-5; pI. 4, figs. 1-8; pI. 5, figs.
1-7; pI. 6, figs. 1-4; text-fig. 1) and Syringothyris sakamotoi Tachibana (1969, p. 24, pI. 6,
figs. 5-7) from the Karaumedate Formationof the southern Kitakami Mountains,· but it
differs from them in having less diverging dental
plates in the pedicle valve.
Syringothyris kitakamiensis Minato, 1952
from the Arisu Formation of the southern
Kitakami Mountains (Minato, 1952, p. 165, pI.
5, fig. 5) is also small in size, but is distinguishedfrom the present form by its high ventral
interarea.

Superfamily Syringothyridacea Fredericks, 1926
Family Syringothyrididae Fredericks, 1926
Subfamily Syringothyridinae Fredericks, 1926
Genus Syringothyris Winchell, 1863

Suborder Delthyrididina Ivanova, 1972
Superfamily Delthyridacea Waagen, 1883
Family Delthyrididae Waagen, 1883
Subfamily Tylothyridinae Carter, 1972

Syringothyris sp.

778. Visean brachiopod Fauna from Soma

Genus Tylothyris North, 1920
Tylothyris laminosa (M'Coy, 1844)
PI. 67, Figs. 8-11

1844

Cyrtia laminosa M'Coy, p. 137, pI. 21,

1858

Spirifera laminosa (M'Coy): Davidson, p.

1905

Syringothyris aff. S. laminosa (M'Coy):

1915

Syringothyris laminosa (M'Coy) forma 2

fig. 4.
36, pI. 7, figs. 17-19 only.
Vaughan, p. 300.

1920

1929
1937

1952
1966

1970
1979

Vaughan, p. 43, pI. 6, figs. 1-3.
Tylothyris laminosa (M'Coy): North, p.
197, pI. 13, figs. 1, 2, 12, 13, 16; textfigs. 4a, 4b.
Tylothyris laminosa (M'Coy): Dehee, p.
23, pI. 3, figs. 12-14.
Spirifer (Tylothyris) laminosus (M'Coy):
Nalivkin, p. 109, pI. 33, figs. 3, 4.
Delthyris aff. D. clarksvillensis (Winchell): Minato, p. 162, pI. 7, figs. 2, 5.
Tylothyris laminosa (M'Coy): Gretchischnikova, p. 158, pI. 16, figs. 11-14;
text-fig. 38.
Tylothyris laminosa (M'Coy): Plodowski,
p. 21, pI. 1, figs. 3-4; text-fig. 9.
Tylothyris laminosa (M'Coy): Nalivkin,p. 131, pI. 49, fig. 15.

Material:-Eight specimens: (1) externalmould of a pedicle valve and internal mouldof a conjoined valve, IGPS colI. cat. no. 97927;(2) external and internal moulds of two pedicle
valves with the external moulds of the posterior
portions attached, IGPS colI. cat. nos. 97928,
97929; (3) external mould of a pedicle valve,
IGPS colI. cat. no. 97930; (4) internal mouldsof three pedicle valves with the external mouldsof the posterior portions attached, IGPS colI.
cat. nos. 97931-97933; (5) internal mould of abrachial valve, IGPS colI. cat. no. 97934.
Description:-Shell wider trigonal in outline,
twice as wide as long; hinge line forming thewidest part of the shell; cardinal extremities
acute or slightly produced; length 20 mm,
width about 39 mm in the largest specimen(IGPS colI. cat. no. 97927).
Pedicle valve moderately and unevenly convex
in lateral profile, with the greatest convexity355

at umbo. Umbo narrow, pointed, strongly
incurved and protruding beyond interarea.
Ventral interarea apsacline, high, slightly arched;
4 mm high on a shell 17 mm long, 21 mm wide
(IGPS colI. cat. rio. 97933). Delthyrium open;
delthyrial angle 27°-33°. Sulcus deep, angular,
originating at beak and extending to the anterior
margin of the valve.
External ornament of the pedicle valve
consisting of simple, subrounded costae andregular, imbricating lamellae. Costae gradually
decreasing the size and strength towards thelateral margins of the valve, with narrow, subangular intercostal grooves; 6 to 7 costae on eachside of lateral slopes. Concentric lamellae being
with a density of 6 to 8 per 5 mm at midvalve.
In the pedicle valve, a sharp, low median
septum of one-half of the valve length occurring
between adductor scars; the posterior portionof septum completely embedded in apicalcallus. Dental plates continuing from the edgesof teeth to the valve floor. Adductor scars
smooth, narrow and elongate. Diductor scars
dendritic, ovate, situated on the flanks of sulcus.
In the brachial valve, a low median septum
extending to about one·half of the length ofthe valve. Cardinal process having about 16
sharp, very thin plates of 0.5 to 1.2 mm long.
Sockets na.'rrow. Crural plates sharp, situatedbetween cardinal process and sockets.
Remarks:-The specimens from Soma arereferred to Tylothyris laminosa (M'Coy, 1844)
because of similarities in size, shape, external
ornament and internal structure.
T. laminosa has· been described or figured
from the Lower and Upper Tournaisian ofBritain, Belgium and the Soviet Union (M'Coy,
1844; Davidson, 1858; Vaughan, 1905, 1915;
North, 1920; Dehee, 1929; Nalivkin, 1979),
the Upper Tournaisian to Lower Visean of theSoviet Union (Nalivkin, 1937; Gretchischnikova,
1966), the Lower and Upper Visean of Afghanistan (Plodowski, 1970) and the Upper Visean ofJapan (Minato, 1952).
Delthyris afi. D. clarksvillensis (Winchell,1865), described and figured by Minato (1952,
p. 162, pI. 7, figs. 2, 5) from the Lower Carbonif.

356

Jun-ichi TAZA WA et al.

erous (=Upper Visean Karoyama Formation,
after Tazawa, Itabashi and Mori, 1981; Tazawa,1981) of Okuhinotsuchi in the southern Kitakami Mountains, appears to be assigned to T.
laminosa in size, external ornament and internal
structure of the brachial valve. Ijiri and Minato(1948, p. 167) reported Tylothyris cf. T. laminosa from the same locality of Okuhinotsuchi,
although it was not described or illustrated.
Hase and Yokoyama (1975, pI. 18, figs. 2,3)
figured Tylothyris sp. from the Lower Viseanof Hina, Okayama Prefecture, southwest Japan,
but the Hina specimens are too poorly preserved
for comparison to the present form.
Tylothyris pianimedia Cvancara (1958, p.
876, pI. 112, figs. 8-10, 12; pI. 113, figs. 1-5;
text-fig. 7) differs from T. laminosa in having
more numerous costae (9 to 14 costae on eitherside of the sulcus) and a sulcus with flattenedbottom.
Tylothyris transversa Roberts (1971, p. 224,
pI. 50, figs. 1-19; text-figs. 71-73) from theLower Tournaisian of the Bonaparte Gulf Basin,
northwestern Australia can be distinguishedfrom T. laminosa by its more transverse shell
outline, lower ventral interarea and wider
delthyrial angle (50 0 -55°).
Acknowledgements
We are indebted to Mr. T. Takahashi of
Yumoto High School, Fukushima Prefecture
for information on the fossil locality, Professor
M. Kato of Hokkaido University for identification of coral fossils, Mr. S. Otomo of Tohoku
University for help with photography, Professor
P. Copper of Laurentian University, Canada forreading the manuscript, and Mr. and Mrs. Igari
of Kamitochikubo, Kashima-cho for hospitality
in the field.

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357

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Abukuma pifji,tllft Ainosawa 1t / iR. Akiyoshi fxti', Arisu ~tt, Choanji *'t(~, Fukushima m
iJh. Hikoroichi R 9'11'1). Hina 13 rn. Karaumedate r.~l1iJt1L Karoyama 1Jn:lJIU. Kashima·eho !.lILli'I,
IHJ, Kitakami ~U::. Mano J:t:'llf. Matsugadaira t'i:?- f. Nagasaka :N::[&. Oashi j:::.f'S. Okayama f,'ITJ
LU. Okuhinotsuchi ~* 0):1::.. Omi t\flliJ:. Onimaru Jt'.:h, Soma :m.~. Tateishi J'L:fi. Tobigamori~,,'f,?-.i11~,359

Uwagaya J::IJ!)', Yamaguchi I.UI=!. Yumiorezawa ~:jfriR

360

Jun-ichi TAZA WA et aZ.
~~~w~m.~.~.~~.~~-~R~.~E:~~~W~,m••• (ma.mm

l!1IHEalBJlLE) ~.~JfIMLl::'€ffl~1tY!5-iJ' i?1&JtIl ~ ~ -tlt OltrJtllE.tR*c) ~~-rR~~iHI:EiJ~14
i?ht.:o vVIE.~~~.'j!{Iliff\';lt1ltrJtllE.tR*c r 1L-;f. -tltc ~h -n't.:iJ~, j,'t.c < c U'l]JfI~~
J:'€ffllt1&!tIJ ~ ~ -tlt~.I#:flI(4o/.n:',i;Q c ~*-i?h Qo i t.:.~Jf1 c J:{ft~J:'€ffl ~ ~ -i:iff.lLE~ c~r,,'I;:'lt~ L ~ ,m#4/G~-g-iJ~fr:{£-r:Q c ~ 'bh -n ,t.:iJ~, ~~-e/G~-g-~IDE~ltWl6b i?.h 1',
LiJ,t .::. ~R~.1I::E~.t±l~;:. J: I) jj!jj~rl3'I;:.~~~h -c ~ ,t.:llifr,,'r,,'~JiHtt.c~''::' c iJ~f!ij i?iJ'l;:' t.c..., t.:o .::. ~t.: V't*~ • [l1]~ ~ht.:R~.{t(k~ 7 fiI!-c',i; Q: Leptagonia analoga (Phillips),
Rugosochonetes d. R. celticus Muir-Wood rev. Brand, Yanishewskiella d. Y. angulata(Yanishevsky), Spirifer sp., Brachythyris sp., SyringothYl'is sp., Tylothyris laminosa(M'Coy)o
EBlRMi-· ll!Ijj]$:~. ~~Explanation of Plate 67(Natural size unless otherwise indicated)
Figs.1a-1c. Rugosochonetes cf. R. celticus Muir-Wood rev. Brandla, 1b, 1c. Internal mould of a pedicle valve and the rubber cast, IGPS coll. cat. no. 97916.
Figs. 1b, 1c are x2.
Figs. 2-4. Leptagonia analoga (Phillips)
2a, 2b. Dorsal and anterior views of external mould of a brachial valve, 2c. Internal mould ofa brachial valve, IGPS colI. cat. no. 97912, 3. Rubber cast of a brachial valve exterior, IGPS
coll. cat. no. 97914, 4. External mould of a brachial valve, IGPS coll. cat. no. 97913.
Figs. 5a, 5b. Yanishewskiella cf. Y. angulata (Yanishevsky)
Rubber cast of a brachial valve exterior, IGPS colI. cat. no. 97917. Fig. 5b is x2.
Figs. 6-7. Syringothyris sp.
6. Rubber cast of a pedicle valve exterior, IGPS colI. cat. no. 97923, 7a, 7b. Ventral andposterior views of internal mould of a pedicle valve, 7c, Rubber cast of a pedicle valve
interior, IGPS coll. cat. no. 97925.
Figs. 8-11. Tylothyris laminosa (M'Coy)
8a. Rubber cast of a pedicle valve, 8b, 8c, 8d. Ventral and dorsal views of internal mould of
a pedicle valve, IGPS colI. cat. no. 97927, 9a, 9b. Rubber casts of a pedicle valve exterior
and interior, IGPS coll. cat. no. 97928, 10. Rubber cast of a pedicle valve exterior, IGPS
coll. cat. no. 97929, 11. Rubber cast of a pedicle valve interior, IGPS colI. cat. no. 97933.
Fig. 8c is x2.
Figs. 12-14. Spirifer sp.
12a, 12b, 12c, 12d. Ventral, posterior, lateral and anterior views of internal mould of apedicle valve, IGPS colI. cat. no-. 97918, 13. Internal mould of a pedicle valve, IGPS colI.
cat. no. 97919, 14. A brachial valve, IGPS colI. cat. no. 97920.
Fig. 15. Brachythyris sp.
Internal mould of a brachial valve, IGPS colI. cat. no. 97921.

TA ZA WA et al. : Visean brachiopod Fauna from Soma

Plate 67

7c

7a

8d

Trans. Proc. Palaeont. Soc. Japan, N.S., No. 134, pp. 361-373, pis. 68-70, July 30, 1984

779. OBSERVATIONS ON OCCURRENCES OF JAPANESE NEOGENE NATICIDS(GASTROPODA) BEARING CALCAREOUS OPERCULA*RYUICHI MAJIMA
Institute of Geoscience, University of Tsukuba,
Ibaraki 305, Japan

Abstract.

Twenty-four calcareous opercula of seven naticid species, including

Cryptonatica janthostoma, C. clausa, C. janthostomoides, C. adamsiana, C. ichishiana, "Naticarius" sp. and Natica vitellus, which are accompanied by their shells
are recorded from fifteen Neogene formations in Japan. The opercula of the
Miocene species Cryptonatica ichishiana, and the Miocene-Pliocene species"Naticarius" sp. (= Naticarius cf. N. niasensis Wissema of MacNeil, 1961) arerecorded for the first time in this paper.
According to the relative positions of the operculum and shell, the following
five types of preservation are recognized. Type (1): operculum is preserved at the
far inside of body whorl (18 specimens). Type (2): operculum is turned upside
down at the base of shell, namely its external surface is attached to the base (3specimens). Type (3): operculum almost coincides with the apertural margin andis only slightly recessed into the body whorl (1 specimen). Type (4): operculum
is normal to the apertural plane. The anterior half protrudes from the apertureand the internal surface is attached to the inner lip (1 specimen). Type (5): operculum is completely detached from the shell (1 specimen).
The position of head-foot mass when naticids died can be reconstructedbased on the occurrences of Types (2) and (3). Types (2) and (3) coincide, respectively, with the operculum positions of naticids that died with the head-foot mass
protruding from the shell, and those that died with it withdrawn into the shell.

The associated occurrences of fossil naticid
shells and opercula have been reported fromother countries, as follows: Natica duncani
Jenkins and Natica rostalina Jenkins from the
Miocene Subapennine Formation in Indonesia
by Jenkins (1863), Natica pachyope Cossmann
and Peyrot from the Middle Miocene (Tortonien)
in France by Cossmann and Peyrot (19171919), Natica epiglottina Lamarck and Natica
burtoni Wrigley from the Paleogene in Englandby Wrigley (1949), Stigmaulax guppiana (Toula)from the Miocene Gatun Formation in Panama
by Woodring (1957), Natica marochiensis(Gmelin) from the Upper Miocene Palau Lime-

Introduction
Naticid snails are one of the dominant groups
in the Neogene molluscan fauna of Japan (Hatai
and Nisiyama, 1952; Masuda and Noda, 1976).
The fossils of naticid opercula have been illustrated from Japanese Neogene formations by
Nomura and Hatai (1937), Kaseno and Matsuura (1965), Itoigawa et al. (1974 and 1981),
Ogasawara (1977) and Kotaka and Kato (1979),
but these opercula were not accompanied byshells.
*Received May 15, 1983;readJanuary 23, 1983,
in Tokyo.
361

Ryuichi MAJIMA

362

stone in Palau Island by Ladd (1977), andNatica (Cryptonatica) oregonensis (Conrad) fromthe Upper Miocene Empire Formation in Oregon,
U.S.A. by Marincovich (1977). In Japan, Chinzei(1959) mentioned the occurrence of naticid
opercula as "the specimens were found withcalcareous operculums" when he described
Natica (Cryptonatica) russa (Gould) from the
Pliocene Kubo Formation in Iwate Prefecture,and Itoigawa et al. (1982) described the associateoccurrence of shell and operculum of their
Cryptonatica sp. 1 from the Miocene Mizunami
Group in Gifu Prefecture.
During the paleontological study of Neogene
naticids in Japan, the present writer examined
twenty-four specimens (seven species) in association with their calcareous opercula from
fifteen Neogene formations (Text-fig. 1, Table1). It is worthwhile to record those occurrences
and to compare the operculum positions offossils with those of living naticids. Detailed
taxonomic discussions of the naticids will begiven at another opportunity.
Occurrences of Naticid Opercula in Association
with their Shells
The term "shell" is used herein in reference
to the calcareous part of a naticid specimen

Japan

except for its operculum. The localities (Textfig. 1), formations, associate species and positions of opercula for fossil naticids bearing
opercula are described according to each species,
as follows.
Cryptonatica janthostoma (Deshayes, 1839):
Pl. 68, Figs. 1a-2e, Pl. 70, Figs. 5a-b, Text-fig.
2-a.
Cryptonatica janthostoma is characterized byhaving a narrowly open umbilicus between
parietal and umbilical calluses and an almost
smoothed operculum. The shells of C. jan thostoma bearing opercula were collected from
Locs. (1) and (7).
Locality (1): A cliff on the right bank of theOshirarika River, about 1500 m east from
Misawa, Shin-totsugawa-cho, Kabato-gun, Hokkaido; Pliocene Porokaoshirarika Formation of
Kobayashi et al. (1957); PI. 68, Figs. la-2e,
Text-fig. 2-a.
Two shells of C. janthostoma bearing opercula
were obtained from tuffaceous siltstone in association with Acila gottschei (Bohm), Serripes
groenlandicus (Bruguiere), Macoma calcarea(Gmelin), Neptunea sp. and Balanus sp. (attaching to Serripes groenlandicus and Neptunea sp.).
The two shells are almost dissolved. The
shells of Figs. la-b, Ie, 2a-b, and 2e in PI. 68 areSea/"'-.-/ 8':x2:~ 1':'-0C'J~Hokkaido

X1:~;;6J5

K~ne-jima;~ ."''''
14 Okinawa- jima

Honshu~Pacific

Ocean

-10

500km
,

I

Text-fig. 1. Map showing fossil localities. Pliocene: Porokaoshirarika Formation (1), Atsuga
Formation (2), Nakanokawa Formation (3), Tomikawa Formation (4), Sunagomata Formation(5), Kobinaizawa Formation (6), Sasaoka Formation (7), Kubo Formation (8), Shibikawa Formation (9), Iioka Formation (10), Omma Formation (11), Fusakina Formation (15). Miocene:Oi Formation (12), Joyama Mudstone Member of Yatsuo Formation (13), Yonabaru Clay Member of Shimajiri Formation (14).

779.

Fossil Natic ids bearing calcareous Opercula363

Text-fig. 2. a : Occurr ences of th e shells and opercula of Cryptonatica jan thostoma, Pliocene Porokaoshirari ka Formation , Loc. 1, IGUT no. 15569-1, 2. Th eshells are rubber casts and t h e opercula rubber molds. b: Models of the specimens ofa reconstructed by Recent specimens of C. janthostoma.
represented by rubber casts. Both t h eir opercula
are well preserve d at the basal parts of the sh ells(PI. 68, Figs. 1d and 2d). Figs. 1c and 2c in PI.
68, and Text-fig. 2-a show the positions of th e
opercula represented by rubber molds. The
external surfaces of both th e opercula are
attached to th e bases of th e shells. Models ofthese occurrences are shown in Text-fig. 2-b
usin g Recent specimens of Cryptonatica jan thostoma.
Locality (7): Road side cut on th e r idge,
abou t 4 00 m northeast of Monzen, Gojome-cho,
Minami-Akita-gun, Akita Prefectur e; Pliocene
Sasaoka Formation of Honda (1978); PI. 70,
Fi gs . 5a-b.
Only one shell among the six of C. jan thostoma collected from fine to m edium grainedsandstone is accompan ied by its operculum,
which is preserved at the far inside of th e bodywhorl. At the locality, C. janthosto ma is associated with A nadara amicula elongata Noda,
Mizu hopecten poculum (Yokoyama), Turritellasaishuensis Yokoyama and Glossau lax did y ma
(R6ding) .

Cryptonatica clausa (Broderip and Sowerby ,1829): PI. 69, Figs. 1a-5b, PI. 70 , Figs. 1-4b.
Cryptonatica clausa (Broderip and Sowerby ,an umbilicus entirely closed by a semicircularumbilical callus, and an operculum with analmost flat external surface. The shells of C.
clausa bearing opercula were collected from
Locs. (2)-(6), (8) and (10).
Locality (2) : A cliff on the right bank, about
200 m u pstream of the Fupumopu-zawa, Atsugacho, Saru-gun , Hokkaido; Plioce ne Atsuga Formation of Yamaguchi (19 58); PI. 69, Figs. la-b.
One shell of C. clausa bearing an operculum
was collected from fine grai ned sandstone. The
operculum is preserved at the far inside of the
body whorl. At th e locality , C. clausa is associated with Fortip ecten takahashii (Yokoyama),
Callithaca adamsi (Reeve), Mya japonica Jay ,Turritella for tilirata Sowerby, Fusitriton oregonensis (Redfield), Buccinum sp. and Dentalium sp .
Locality (3): Small outcrop on the left bank,364

Ryuichi MAJIMA

about 1250 m upstream of the Byakutan River,
Kuromatsunai-cho, Suttsu-gun, Hokkaido; Pliocene Nakanokawa Formation of Sawada (1962):
PI. 69, Figs. 2, 4a-b, PI. 70, Fig. 1.
Four shells among the twelve of C. clausa
collected from massive silstone are associatedwith their opercula. Three of the four opercula
are preserved at the far inside of the body whorl.
The external surfaces of two opercula face theaperture (PI. 69, Fig. 2), and the other one faces
the inner wall of the body whorl (PI. 70, Fig. 1).
The remaining operculum of the four is preservedat the base of the shell and its external surface is
attached to the base (PI. 69, Figs. 4a-b). The last
occurrence almost coincides with those of thetwo specimens of C. jantohostoma (PI. 68, Figs.
1a-2e, Text-fig. 2-a) at Loc. 1.
The four shells bearing opercula mentioned
above were bored by naticids. At the locality,
C. clausa is associated with Mizuhopecten yessoensis (Jay), Euspira pi/a (Pilsbry), Mitrella sp.
and Oenopota kuromatsunaiensis Sawada. Besides the species mentioned above, Sawada(1962) reported the following species from thislocality (loc. 7 of Sawada, 1962): Acila nakazimai Otuka, Yoldia johanni Dall, Limopsis
tokaiensis Yokoyama, Macoma tokyoensis Makiyama, Turritella nipponica Yokoyama, andothers.
Locality (4): River floor, about 1200 m upstream of Hosokomata-zawa, Tomikawa-cho,
Kamiiso-gun, Hokkaido; Pliocene Tomikawa Formation of Mitani et al. (1965); PI. 69, Figs.
3a-b.
Only one shell of C. clausa bearing an operculum was obtained from conglomeratic sandstone yielding abundant molluscan fossils, suchas Limopsis tajimae Sowerby, Porterius daW(Smith), Mizuhopecten yessoensis (Jay), Swiftopecten swiftii (Bernardi), Papyridea kurodai
Hatai and Nisiyama, Pseudamiantis tauyensis(Yokoyama), Umbonium akitanum Suzuki,
Homalopoma amussitatum (Gould), Turritella
fortilirata Sowerby and Euspira pi/a (Pilsbry).
The operculum (PI. 69, Fig. 3b) was preservedat the far inside of the body whorl, but it was
detached from the shell (PI. 69, Fig. 3a) and its
posterior part was erroneously broken during
preparation.
Locality (5): A large river-side cliff, about
400 m upstream of the Chika River, Chikagawa,Mutsu City, Aomori Prefecture; Pliocene Sunagomata Formation of Imai (1961).
Among the twenty-two shells of C. clausa
collected from fine grained sandstone, twoshells have opercula at the far inside of bodywhorl. The opercula and apertural parts of thetwo specimens were conspicuously weathered at
the exposure. One shell of the two was bored bya naticid.
At the locality, C. clausa is associated with
Mizuhopecten tokyoensis hokurikuensis (Akiyama), Chlamys nipponensis Kuroda, Swiftopecten swiftii (Bernardi), Venericardia ferruginea
Clessin, Clinocardium chikagawaense Kotaka,
CaWthaca adamsi (Reeve), Turritella saishuensis
Yokoyama, Euspira pila (Pilsbry), Glossaulax
didyma (Roding) and Niotha livescens (Philippi).
Locality (6): A cliff on the left bank of the
Taneume River, Tanosawa, Futatsui-cho, Yamamoto-gun, Akita Prefecture; Pliocene Kobinaizawa Formation of Hirayama and Sumi (1963);
PI. 70, Figs. 4a-b.
Among the four shells of C. clausa collectedfrom fine grained sandstone, one shell has its
operculum at the far inside of the body whorl.
The apertural and basal parts of the specimen
were damaged during preparation.
Chinzei (1973) reported the following molluscan fossils from this locality (loc. 24 of Chinzei, 1973): Nuculana robai (Kuroda), Macoma
calcarea (Gmelin), Turritella nipponica Yokoyama, Fulgoraria prevostiana (Crosse), andothers.
Locality (8): A cliff on the right bank of the
Umiue River, Ochiai, Ninohe City, Iwate Prefecture; Pliocene Kubo Formation of Chinzei(1959);PI. 70, Figs. 2a-3b.
Three shells among the six of C. c/ausa collected from fine grained sandstone are associatedwith their opercula: two at the far inside of bodywhorl (PI. 70, Figs. 3a-b) and one completely
detached from the shell (PI. 70, Fig. 2a). The
apertural part of the last shell (PI. 70, Fig. 2c)
was damaged by weathering at the exposure.
At the locality, C. clausa is associated with
Nemocardium (Keenaea) samarangae (Makiyarna), Turritella nipponica Yokoyama, "Macron"nipponensis Chinzei and Propebela sp.
Locality (10): A quarry, about 1250 m westof Matsugishi National Railway Station, Takanocho, Choshi City, Chiba Prefecture; Pliocene

779. Fossil Naticids bearing calcareous Opercula
Iioka Formation of Ozaki (1958); PI. 69, Figs.
5a-b.
Three shells among the thirty of C. clausa
collected from siltstone have their opercula.
Two opercula of the three are preserved at the
far inside of body whorl, and the other one
almost coincides with the apertural margin (PI.
69, Figs. 5a-b). The adaxial margin of the last
operculum is at the inner lip of the aperture,
but the abaxial one is slightly recessed into thebody whorl and its external surface looks toward
the aperture. One of the former two shells andthe last were bored by naticids.
At the locality, C. clausa is associated with
Nuculana ikebei Suzuki and Kanehara, Portlandia
sp., Palliolum macrocheiricola Habe, Cardiomyasp., Euspira pallida (Broderip and Sowerby),
Ancistrolepis trochoideus (Dall), Riuguhdrillia
engonia (Watson), Aforia diomedea Bartsch,
Rectiplanes sanctiioannis (Smith) and Admete
couthonyu (Jay).

Cryptonatica janthostomoides (Kuroda andHabe, 1949): PI. 70, Figs. 6a-7b.
Cryptonatica janthostomoides is characterizedby having a shallowly and narrowly open umbilicus around the margin of the umbilical callus
in adults and its operculum sculptured with
two strong grooves at the abaxial margin of theexternal surface. One operculum in association
with a small shell obtained from the Shibikawa
Formation (Loc. 9) was fragile and the externalsurface was almost missed during preparation(PI. 70, Fig. 6b). The umbilicus is wholly coveredby the umbilical callus, which is common injuveniles. The adults from the same locality
have umbilici open shallowly and narrowly as
mentioned above (PI. 70, Figs. 7a-b).
Locality (9): Sea side cliff, about 500 m
southwest of Anden, Iriai, Oga City, Akita Prefecture; Pliocene Shibikawa Formation of
Huzioka (1959); PI. 70, Figs. 6a-b.
Among twenty-five shells of C. janthostomoides collected from fine to medium grainedsandstone, only one has an operculum at the far
inside of the body whorl. At the locality, C.
janthostomoides is associated with Acila insignis(Gould), Arca boucardi Jousseaume, Mizuhopecten yessoensis (Jay), Homalopoma amus-

365

sitatum (Gould), Turritella andenensis Otuka,
Olivella japonica (Pilsbry), Euspira pi/a (Pilsbry)
and Eunaticina papilla (Gmelin).
Cryptonatica adamsiana (Dunker, 1860): PI.
69, Fig. 6.
Cryptonatica adamsiana is characterized byhaving a deeply open umbilicus around a smallumbilical callus lobe, and an operculum sculptured with two weak grooves along its abaxial
margin.
Locality (11): A river bed of the Sai River,Okuwa, Kanazawa City, Ishikawa Prefecture;
Pliocene Omma Formation of Kaseno and Matsuura (1965);PI. 69, Fig. 6.
Only one shell among the five of C. adamsiana
collected from fine grained sandstone has an
operculum at the far inside of the body whorl.
At the locality, C. adamsiana is associated with
Acila insignis (Gould), Limopsis sp., Glycymeris
nipponica (Yokoyama), Mizuhopecten tokyoensis
hokurikuensis (Akiyama), Lucinoma
annulata (Reeve), Paphia schnelliana (Dunker),
Turritella saishuensis Yokoyama, Glossaulax
reiniana (Dunker), G. hagenoshitensis (Shuto),
Siphonalia spadicea (Reeve) and Dentalium sp.

Cryptonatica ichishiana (Shibata, 1970): PI.
69, Figs. 7-9.
Cryptonatica ichishiana is characterized byhigh spire, well swollen whorls and an umbilicus
closed by a small umbilical callus. The operculum of C. ichishiana has not previously beenfound associated with the shell. Two shells of
C. ichishiana bearing opercula were collectedfrom the type locality of the species (Loc. 12)
and from the Yatsuo Formation (Loc. 13). The
former operculum has two very weak lineations
along the abaxial margin of the external surface,
but not the latter one.
The present writer interprets the difference
between the two opercula as an intraspecificvariation because there are no morphologicaldifferences between the shells (PI. 69, Figs. 7-8).
Locality (12): The river bed of the Nagano
River, about 500 m south of Ashizaka, Iono,
Misato-mura, Age-gun, Mie Prefecture; Miocene

Ryuichi MAJIMA

366

Oi Formation of Shibata (1970); PI. 69, Fig. 9.
Fifteen shells of C. ichishiana were collectedfrom calcareous concretions in mudstone, in
assocaition with Malletia inermis (Yokoyama),
Propeamussium tateiwai Kanehara, Cardiomyasp., Spirotropis subdeclivis mitsuganoensis Shi·bata and Dentalium sp.
Only one shell among the fifteen has its
operculum at the far inside of the body whorl.
The anterior and posterior parts of the oper·
culum and apertural side of the whorls (PI.
69, Fig. 9) were erroneously broken during
preparation.
Locality (13): Small outcrop on the leftbank of the Kubusu River at Kashio, Yatsuo·cho,
Nei-gun, Toyama Prefecture; Miocene Joyama
Mudstone Member of the Yatsuo Formationof Sakamoto and Nozawa (1960); PI. 69, Fig. 7.
One shell of C. ichishiana with its operculum
was collected from pebble bearing siltstone, in
association with Portlandia watasei (Kanehara),
Ennucula osawanoensis (Tsuda), Glycymeris sp.,
Anadara makiyamai Hatai and Nisiyama, Dosinia
kawagensis Araki, Oxyperas osawanoensis Tsuda,
Protrotella yuantaniensis Makiyama, Tanea mino·
ensis (Itoigawa), Babylonia kozaiensis Nomura,
Oliva osawanoensis Tsuda, Conus sp. and Acteon
osawanoensis Tsuda.
The operculum is normal to the aperturalplane and its internal surface is attached to theinner lip. The anterior half protrudes from theaperture but was erroneously broken duringcollection. Therefore, the posterior half only ispreserved in the body whorl (PI. 69, Fig. 7).
"Naticarius" sp.: PI. 69, Figs. 1G-llb.
The present unnamed species is identified
with "Naticarius cf. N. niasensis Wissema"of MacNeil (1961) from his Miocene-Pliocene
Shinzato Tuff Member of the Shimajiri Formation in Okinawa Prefecture. According to
MacNeil (1961), "N. niasensis" was originallyproposed by Wissema in his doctor's thesis(1947) which seems to be invalid by Article 8of the International Code of Zoological Nomenclature. The present writer has unfortunately
had no chance to examine the reference, so it
should be entrusted to future work as to whether
this scientific name is available or not. The
species, therefore, is treated as "Naticarius"

sp. in the present study.
As was described by MacNeil (1961), thespecies is characterized by axial wrinkles (PI.
69, Fig. 10) which are distinct in juveniles but
die out on the central part of the body whorl
in adults. The genus Stigmaulax March, 1852,
flourishing in the tropical eastern Pacific andCaribbean seas, also has axial wrinkles, but as
mentioned by Woodring (1957) and Marincovich(1977), the external surface of the operculum
is sculptured with an elevated central rib. Though
MacNeil (1961) compared the present species
with S tigmau lax , he classified the species into
Naticarius because the operculum was unknownat that time.
The operculum of "Naticarius" sp. has two
distinct grooves along the abaxial margin of theexternal surface (PI. 69, Fig. llb), by which
the present species is distinguished from
Stigmaulax. On the other hand, this operculum
resembles that of the genus Tanea Marwick,
1931, flourishing in the tropical western Pacificand Indian Ocean, but the shell surface of
Tanea is smooth as mentioned by Oyama (1969).
Locality (14): A cliff, about 400 m east of
the Ozato Senior High School, Ozato-son,
Shimajiri-gun, Okinawa Prefecture; Miocene
Yonabaru Clay Member of the Shimajiri Formation of MacNeil (1961); PI. 69, Figs. 11 a-b.
One shell among the five of the present
species collected from siltstone has its operculumat the far inside of the body whorl. The aperturalpart of the shell was erroneously broken in thefield (PI. 69, Fig. 11a). At the locality, thepresent species is associated with Ennucula sp.,
Ancilla chinenensis MacNeil, Uromitra noharai
Noda, Asprel/a sp. and Dentalium sp.
Natica vitellus (Linnaeus, 1758): PI. 69, Figs.
12a-b.
Natica vitellus is characterized by a smalland subtrigonal umbilical callus along the posterior margin of the umbilicus. The external surface of the operculum is sculptured with twoor three distinct grooves along the abaxial
margin, and its adaxial edge is crenulated.

Table 1. Occurrences and measurements of fossil naticids bearing calcareous opercula.
Types

Species(1)

Cryptonatica
clausaC. clausa
C. clausa
C. clausa
C. clausa
C. clausa
C. clausa
C. clausa
C. janthostoma
C. clausa
C. clausa
C. janthostomoides
C. clausa
C. clausa
C. adamsiana
C. ichishiana"Naticarius" sp.
Natica vitellus
Cryptonatica
janthostoma
C. janthostoma
C.clausa
C. clausa
C. ichishiana
C. clausa

(1)
(1)
(1)
(1)
(1)
(1)
(1)
(1)
(1)
(1)
(1)
(1)
(1)
(1)
(1)
(1)
(1)
(2)
(2)
(2)
(3)
(4)
(5)

Pred.

C
C
C
C
B

B
A

?

B

B
A

Loc.

H.S.

W.S.

2

30.7

26.1+

3
3
3
4
5
5
6
7
8
8
9
10
10
11
12
14
15
1

12.6
10.3+
15.9+
11.2
10.9
16.8
14.5+
21.4
29.5
30.4
14.6
11.7
14.0
21.2
8.9+
13.7+
14.0
35.0

11.7
9.3+
14.8
9.4
10.0
15.2
15.1
22.9
26.0
25.9
13.9
10.2
12.1
19.7
7.1+
11.0+
14.0
31.3

1
3
10
13
8

44.4
20.2
12.5
17.7
24.8

33.1+
10.9
15.9
21.8

H.O.

8.3+
7.6
9.6+
5.0+
5.7+
6.8+
15.5
17.5
8.0
8.1
12.5
3.9+
9.0
8.9

27.9

7.0+
16.9

W.O.

Cat. no.

Illustrations

13.7!GUT no. 15606

PI. 69, Figs. 1a-b

5.2
4.4
6.4
3.0
2.8+
4.0+
6.2
10.0
11.8
11.2
5.0

!GUT no. 15064-2
IGUT no. 15064-3!GUT no. 15064-4!GUT no. 15065!GUT no. 15572-1!GUT no. 15572-2!GUT no. 15573!GUT no. 15574!GUT no. 15575-2!GUT no. 15575-3!GUT no. 15576-1!GUT no. 15570-2!GUT no. 15570-3
IGUT no. 15571-1
IGUT no. 15066-1!GUT no. 15067-1
GIYU no. 560!GUT no. 15569-1

PI. 69, Fig. 2
PI. 70, Fig. 1

6.6
7.2
2.8
5.7
5.2
16.8
18.1
8.0
4.5
6.7+
10.8

!GUT no. 15569-2
IGUT no. 15064-1!GUT no. 15570-1!GUT no. 15577
IGUT no. 15575-1

PI. 69, Figs. 3a-b

PI. 70, Figs. 4a-b
PI. 70, Figs. 5a-b
PI. 70, Figs. 3a-b

'I

~

~

~

:::::

~

l?PI. 70, Figs. 6a-b

~

0-

(\)

...

Q

PI. 69, Fig. 6
PI. 69, Fig. 9
PI. 69, Figs. l1a-b
PI. 69, Figs. 12a-b
PI. 68, Figs. 1a-e, Text-fig. 2
PI. 68, Figs. 2a-e, Text-fig. 2
PI. 69, Figs. 4a-b
PI. 69, Figs. 5a-b
PI. 69, Fig. 7
PI. 70, Figs. 2a-c

Types: Types of occurrences of operculum and shell. Pred.: Hole bored by naticids at ventral side of body whorl (A), left side of bodywhorl (B), between A and B (C); not bored (-), unknown (?) because of imperfect specimen. Loc.: Localities (shown in Text-fig. 1).
Measurements (in mm): H.S.= Height of shell, W.S. = Width of shell, H.O. = Height of operculum (Maximum length parallel to the axis of
adaxial margin of operculum), W.O. = Width of operculum (Maximum length perpendicular to H.O.), - = Measurement impossible. Cat. no.:Catalogued numbers of the collections of Institute of Geoscience, University of Tsukuba (IGUT) and the Geological Institute of Yokohama
National University (GIYU).

~.
~

Q

0Q
~

c

I:::

'"

~
(\)

~

I:::

is''

c.o

en
-:J

368

Ryuichi MAJIMA

Locality (15): An exposure about 200 m
northeast of Kategaru, Nakazato-son (Kumejima), Shimajiri-gun, Okinawa Prefecture; Pliocene Fusakina Formation of Makino (1975, MS);PI. 69, Figs. 12a-b.
Only one shell among the seven of N. vitellus
had its operculum at the far inside of the bodywhorl. The operculum was detached from theshell during preparation. The left side of bodywhorl was bored by a naticid.
Types of Occurrences of Naticid Opercula
with their Shells
The occurrences of naticid opercula with
their shells are classified into the following five
types (Table 1):
Type (1): This is a type in which the operculum is preserved at the far inside of the bodywhorl. Eighteen specimens of the twenty-four
belong to this type. But the positions of theopercula within this type are slightly different
in each specimen: the anterior parts of some
opercula (PI. 69, Figs. 1a-b, 6, 9, PI. 70, Figs.
3a-b, 5a--Bb) and the posterior parts of others(PI. 68, Figs. 2, lla, PI. 70, Figs. 4a-b) aredeeply recessed into the body whorl. The external surfaces of many opercula face the aperture (PI. 69, Figs. 1a-2, 6, 9, lla, PI. 70, Figs.
3a--Bb) but only one faces the inner wall of
the body whorl (PI. 70, Fig. 1).
Type (2): In this type, the operCUlum isturned upside down at the base of the shell,
namely the external surface is attached to thebase. The two specimens of Cryptonatica jan thostoma from the Porokaoshirarika Formation(Loc. 1) and one of C. clausa from the Kuromatsunai Formation (Loc. 3) belong to this type.
Type (3): This is a type in which the operculum almost coincides with the apertural

margin. The adaxial margin is at the inner lipof the aperture and the abaxial one is slightly
recessed into the body whorl. The externalsurface of the operculum faces the aperture.
One specimen of Cryptonatica clausa from the
Iioka Formation (Loc. 10) belongs to this type.
Type (4): In this type, the operculum isnormal to the apertural plane and its anterior
half protrudes from the aperture. The internalsurface is attached to the inner lip. One specimenof Cryptonatica ichishiana from the Joyama
Mudstone Member of the Yatsuo Formation(Loc. 13) belongs to this type. This type resembles one operculum belonging to Type (2),namely one specimen of C. janthostoma (PI.
68, Fig. 1c, below specimen of Text-fig. 2)
at Loc. 1, but the internal surface of the latter
faces the outer lip of the aperture.
Type (5): This is a type in which theoperculum is completely detached from its
shell. One specimen of Cryptonatica clausa
from the Kubo Formation (Loc. 8) belongs tothis type.
Summary of Observations of Naticid Opercula with their Shells
The following two distinct conditions ofliving naticid opercula can be recognized.
(a): A condition when the head-foot mass
is withdrawn in the shell. This condition, in
which the operculum closes the aperture, isshown in Text-fig. 3-a.
(B): A condition when the head-foot mass
protrudes from the shell. It is not easy to observe the operculum in this case, because theoperCUlum is entirely concealed by the headfoot mass.
When Taki (1934) illustrated an anatomical

Explanation of Plate 68
Figs. la-e, 2a-e. Cryptonatica janthostoma (Deshayes). Pliocene Porokaoshirarika Formation(Loc. 1). Figs. 1a-d, xl.O, Fig. Ie, xO.7, !GUT no. 15569-1; Figs. 2a-d, xl.O, Fig. 2e,xO.7, IGUT no. 15569-2. Figs. 1a-b and 2a-b, rubber casts. Figs. Id and 2d, opercula in
the host rock. Figs. 1c and 2c, rubber molds of opercula on the rubber casts of shells. Figs.
Ie and 2e, rubber casts of shells in the host rock.

MAJIMA: Fossil Naticids bearing calcareous Opercula

Plate 68

779. Fossil Naticids bearing calcareous Opercula

chart of Glossaulax didyma protruding its headfoot mass, the external surface of the operculum
was attached to the base of shell as shown in
Text-fig. 3-b. Schileyko (1977) illustrated
the mode of movement of the naticid operculum.
The position of the external surface of theoperculum in Schileyko's figure is identical to
Text-fig. 3-b, in which the head-foot mass
protrudes from the shell.
Condition (A) almost coincides with the fossil
operculum position of the Type (3) which isrepresented by the specimen of C. clausa (PI.
69, Figs. 5a-b) from the Iioka Formation (Loc.
10). The same occurrences of fossil naticid opercula were illustrated by Jenkins (1863 , pI. 6,
figs. 6, 8), Woodring (1957, pI. 20, fig. 18)
and Marincovich (1977 , pI. 42, fig. 9) . It is369

thought that the specimen from the Iioka wasdead under the condition (A).
Condition (B) almost coincides with thefossil operculum position of Type (2), which isrepresented by two specimens of C. janthostoma(PI. 68, Figs. la-2e, Text-fig. 2-a) from the
Porokaoshirarika Formation (Loc. 1) and oneof C. clausa (PI. 69, Figs. 4a-b) from the Kuromatsunai Formation (Loc. 3). It is thought thatthese three specimens were dead under thecondition (B).
The specimens of Type (1) are the mostfrequent in occurrence. There may be two way
to explain their operculum conditions when
they were dead (Text-fig. 3). The Type (1) is
probably traced back to the condition (A) as
shown by a-.Te in Text-fig. 3. But the displace-Text-fig. 3. Schematic positions of opercula and models of the displacements ofoperculum. Q : Position when the head-foot mass of a living naticid is withdrawn into shell(condition (A) in the text). b: Position when the head-foot mass protrudes from shell(condition (B) in the text). e : Posi t ion of fossil operculum preserved at the far inside ofbody whorl (Type (1) in the text). Q-.Te and b-.Tc-.Td-.Te: Models of operculum displacementsby sediments, showing the ex planations for the original positions of opercula of Type (1).

370

Ryuichi MAJIMA

ment of operculum shown by b~c-+d;~e in Textfig. 3 is also possible. This is due to the possibility
that the operculum may be recessed into thebody whorl by sediments displacing the soft
parts which are dissolved after the death of
the animal.
In the occurrences of the opercula classified
as Types (2) and (3), the adaxial margin of theoperculum (PI. 69, Figs. 5a-b, Type (3)) fromthe Iioka Formation (Loc. 10) is slightly recessed
into body whorl, and the posterior half of theoperculum (PI. 68, Fig. 1c, below specimen of
Text-fig. 2-a, Type (2)) from the Porokaoshirarika Formation (Loc. 1) is recessed into theaperture. These two occurrences, which differ
slightly from conditions (A) and (B), are interpreted as an intermediate step of the operculum
displacement by sediments mentioned above(Text-fig. 3). The operculum (PI. 69, Figs. 5ab) from the Iioka may have been moved by
sediments filling up the space left by soft parts
which are dissolved after the death of the animal.
The position of the operculum from the Poro-kaoshirarika (PI. 68, Fig. 1c, below specimen of
Text-fig. 2-a) almost coincides with that of
Text-fig. 3-c which shows an intermediatestep of operculum displacement by sediments(Text-fig. 3-b~-+d~e).
It is difficult to interpret the original conditions of the opercula classified to the Types(4) and (5), because their operculum positions
are far from the conditions (A) and (B).
From the considerations mentioned above,it is concluded that there are two possible
positions of the head-foot mass when naticids
die. The one state is of the head-foot mass
protruded from the shell and the other is of it
withdrawn into the shell.
It is not easy to consider the common process
of fossilization of the specimens treated hereinbecause of the following reasons. (a): There
are two modes of the occurrence of molluscan
fossils among the fourteen formations at the
localities (Loc. 15 unknown). Namely, the formations at the Locs. 4, 5, 9 and 13 yielded
abundant molluscan fossils, including the naticid

Explanation of Plate 69
Figs. la-b. Cryptonatica clausa (Broderip and Sowerby). Pliocene Atsuga Formation (Loc. 2).
x1.0, IGUT no. 15606.
Figs. 2, 4a-b. Cryptonatica clausa (Broderip and Sowerby). Pliocene Nakanokawa Formation(Loc. 3). Fig. 2, x2.0, !GUT no. 15064-2; Figs. 4a-b, x1.2, IGUT no. 15064-1.
Figs. 3 a-b. Cryptonatica clausa (Broderip and Sowerby). Pliocene Tomikawa Formation (Loc.
4). Fig. 3a, x2.0, Fig. 3b, x2.2, !GUT no. 15065. Fig. 3b, operculum preserved at the far
inside of body whorl of shell (Fig. 3a). The posterior part of operculum was erroneously
broken during preparation.
Figs. 5a-b. Cryptonatica clausa (Broderip and Sowerby). Pliocene Iioka Formation (Loc. 10).
x2.3, !GUT no. 15570-1.
Fig. 6. Cryptonatica adamsiana (Dunker). Pliocene Omma Formation (Loc. 11). x1.4, IGUT no.
15571-1.
Fig. 7. Cryptonatica ichishiana (Shibata). Miocene Joyama Mudstone Member of Yatsuo Formation (Loc. 13). x1.6, IGUT no. 15577.
Figs. 8, 9. Cryptonatica ichishiana (Shibata). Topotype. Miocene Oi Formation (Loc. 12). Fig.
8, x2.0, IGUT no. 15066-2; Fig. 9, x3.5, !GUT no. 15066-1. Fig. 8 shows the whole shellfor comparison.
Figs. 10, 11a-b. "Naticarius" sp. Miocene Yonabaru Clay Member of Shimajiri Formation(Loc. 14). Fig. 10, x1.8, !GUT no. 15067-2; Fig. lla, x2.2, Fig. lIb, x2.7, !GUT no.
15067-1. Fig. 10 shows the whole shell for comparison.
Figs. 12a-b. Natica vitellus (Linnaeus). Pliocene Fusakina Formation (Loc. 15). Fig. 12a, x2.0,
Fig. 12b, x3.0,GIYU no. 560. Fig. 12b, operculum preserved at the far inside of body whorl(Fig. 12a).

MAJIMA: Fossil Naticids bearing calcareous Opercula

Plate 69

779. Fossil Naticids bearing calcareous Opercula

specimens referred to Types (1) and (4), andmight have been deposited under high energy
conditions, but the other formations yielded
sparse molluscan fossils, including the naticid
specimens referred to Types (1), (2), (3) and(5), and might have been deposited under low
energy conditions. (b): It is well known thatnaticids live on and within soft substrate, where
they may be easily buried and preserved.
The present writer believes that the specimens
referred to Types (2) and (3), and many of thosein Type (1), might have died within soft substrates. The positions of the opercula of Type(1) and of the two opercula of Types (2) and(3) are interpreted as resulting from displacement
by sediments (Text-fig. 3). This interpretationsuggests that the specimens were almost enclosed
by sediments when they were dead.
As shown in Table 1, ten of twenty-four
specimens (43%) are bored by naticids. These
predations are the only verified cause of deathof naticid specimens treated herein. However,the correlations between predation and theoccurrence of Types (1)-(5) are uncertain because of the small number of specimens available
for study.
Acknowledgments
The present writer wishes to express his deep
appreciation to Professor Hiroshi Noda, Instituteof Geoscience, University of Tsukuba for his
valuable suggestions and discussions on thepresent study. The writer indebted to Professor
Hisayoshi Igo, University of Tsukuba and
Dr. Louie Marincovich, Jr., U.S. Geological
Survey for their careful reading of the manuscript, and to Professor Takehiko Iwai, Hirosaki
University for his kind suggestions in the fieldof Aomori Prefecture. Acknowledgments are
due to Professors Yoshikazu Hasegawa andToshio Koike, both of the Yokohama National
University for their kind arrangement to examine
the naticid specimens preserved in the University,and to Associate Professor Yutaka Honda, Mie
University for the copyright of an importantnaticid literature.

371

References Cited
Chinzei, K. (1959): Molluscan fauna of Pliocene
Sannohe Group of Northeast Honshu,
Japan. 1. The faunule of the Kubo Formation. Jour. Fac. Sci., Univ. Tokyo, Sec.
2, vol. 12, no. 1, p. 103-132, pis. 9-11.
--(1973): Omma-Manganjian molluscan faunain the Futatsui area of Northern Akita,Japan. Trans. Proc. Palaeont. Soc. Japan,
N. S., no. 90, p. 81-94, pI. 14.
Cossmann, M. and Peyrot, A. (1917-1919):
Conchologie neogenique de I'Aquitaine.
Tome 3 (Gastropodes). Actes Soc. Linn.
Bordeaux, tomes 70-71, p. "385-459,"
pis. "11-12" [po and pis. show the parts of
Naticidae] .
Hatai, K. and Nisiyama, S. (1952): Check listof Japanese Tertiary marine Mollusca.
Sci. Rep., Tohoku Univ., Spec. Vol., no. 3,
p.I-464.
Hirayama, J. and Sumi, K. (1963): Explanatory
text of the geological map of Japan. "Takanosu" sheet. Scale 1:50,000. 90p., Geol.
Surv. Japan (in Japanese with English
abstract).
Honda, Y. (1978): Molluscan fossils from the
Sasaoka Formation, Gojome area, Akita
Prefecture, Northeast Japan. Saito Ho-on
Kai Mus., Res. Bull., no. 46, p. 1-19, pIs.
1-2.
Huzioka, K. (1959): Explanatory text of thegeological map of Japan. "Toga" and "Funakawa" sheets. Scale 1 :50,000. 61p., Geol.
Surv. Japan (in Japanese with English
abstract).
Imai, I. (1961): Explanatory text of the geological map of Japan. "Chikagawa" sheet.
Scale 1:50,000. 45p., Geol. Surv. Japan(in Japanese with English abstract).
Itoigawa, J., Shibata, H. and Nishimoto, H.
(1974): Molluscan fossils of the Mizunami
Group. Bull. Mizunami Fossil Mus., no. 1,
p. 42-203, pIs. 1-63 (in Japanese).
- - , - - , - - and Okumura, Y. (1981): Miocene fossils of the Mizunami Group, Central
Japan. 2. Molluscs. Monogr. Mizunami
Fossil Mus., no.3-A, p. 1-53, pIs. 1-52
(in Japanese).
__ , __ , - - and-- (1982): Miocene fossilsof the Mizunami Group, Central Japan. 2.
Molluscs (continued). Ibid., no. 3-B, p.

372

Ryuichi MAJIMA

1-330 (in Japanese).
Jenkins, H. M. (1863): On some Tertiary Mollusca from Mount Sela, in the Island of Java.
Quart. Jour. Geol. Soc., vol. 20, p. 45-73,
pI. 4.
Kaseno, Y. and Matsuura, N. (1965): Pliocene
shells from the Omma Formation around
Kanazawa City, Japan. Sci. Rep., Kanazawa
Univ., vol. 10, no. 1, p. 27--u2, pis. 1-20.
Kobayashi, 1., Kakimi, T., Uemura, T. and Hata,
M. (1957): Explanatory text of the geological map of Japan. "Takikawa" sheet.
Scale 1 :50,000. 36p., Hokkaido Development Agency (in Japanese with English
abstract).
Kotaka, T. and Kato, H. (1979): Additional
fossil shells from the Utsutoge Formation,
Yamagata Prefecture, Northeast Japan. Saito
Ho-on Kai Mus., Res. Bull., no. 47, p. 1318, pis. 2-3.
Ladd, H. S. (1977): Cenozoic fossil mollusks
from Western Pacific Islands; Gastropoda(Eratoidae through Harpidae). U.S. Geol.
Surv., Prof Paper, 533, p. 1-84, pis. 1-23.
MacNeil, F. S. (1961): Tertiary and Quaternary
Gastropoda of Okinawa. Ibid., 339, p. 1148, pis. 1-21.
Makino, J. (1975, MS): Geology of Kume-jima,
Ryukyu Islands. Yokohama Nat. Univ.,
Grad. Thesis,· Geol. (in Japanese, title
translated).
Marincovich, L., Jr. (1977): Cenozoic Naticidae(Mollusca: Gastropoda) of the Northeastern
Pacific. Bull. Amer. Paleont., vol. 70, no.
294, p. 165-494, pis. 17-42.
Masuda, K. and Noda, H. (1976): Check list andbibliography of the Tertiary and Quaternary
Mollusca of Japan, 1950-1974. Saito Ho-on

Kai,494p.
Mitani, K., Osanai, H., Matsushita, K. and Suzuki, M. (1965): Explanatory text of the geological map of Japan. "Hakodate" sheet.
Scale 1 :50,000. 32p., Geol. Surv. Hokkaido(in Japanese with English abstract).
Nomura, S. and Hatai, K. (1937): A list of
Miocene Mollusca and Brachiopoda collectedfrom the region lying north of Rikuzen
Province, Japan. Saito Ho-on Kai Mus., Res.
Bull., no. 13, p. 121-145, pis. 17-21.
Ogasawara, K. (1977): Paleontological analysisof Omma Fauna, Toyama-Ishikawa area,
Hokuriku Province, Japan. Sci. Rep., Tohoku Univ., 2nd. Ser. (Geol.), vol. 47, no. 2,
p. 43-156, pis. 3-22.
Oyama, K. (1969): Systematic revision ofJapanese Naticidae (Preliminary report).
Japan. Jour. Malac., Venus, vol. 28, no. 2,
p. 69-88, pis. 4-5 (in Japanese with English
abstract).
Ozaki, H. (1958): Stratigraphical and paleontological studies on the Neogene and Pleistocene formations of the Tyoshi district. Bull.
Nat. Sci. Mus. (Tokyo), no. 42, p. 1-182,
pis. 1-24.
Sakamoto, T. and Nozawa, T. (1960): Explanatory text of the geological map of Japan.
"Yatsuo" sheet. Scale 1:50,000. 63p., Geol.
Surv. Japan (in Japanese with English
abstract).
Sawada, Y. (1962): The geology and paleontology of the Setana and Kuromatsunai
areas in the Southwest Hokkaido, Japan.

Explanation of Plate 70
Fig.l. Cryptonatica clausa (Broderip and Sowerby). Pliocene Nakanokawa Formation (Loc. 3).
x2.2, IGUT no. 15064-3. Operculum is turned upside down in the body whorl.
Fig. 2a--c, 3a-b. Cryptonatica clausa (Broderip and Sowerby). Pliocene Kubo Formation(Loc. 8). Fig. 2a, xl.O, Figs. 2b--c, x1.3, IGUT no. 15575-1; Figs. 3a-b, xLI, IGUT no.
15575-2.
Figs.4a-b. Cryptonatica clausa (Broderip and Sowerby). Pliocene Kobinaizawa Formation(Loc. 6). x1.7, IGUT no. 15573.
Figs. 5a-b. Cryptonatica janthostoma (Deshayes). Pliocene Sasaoka Formation (Loc. 7). xl. 7,
IGUT no. 15574.
Figs. 6a-b, 7a-b. Cryptonatica janthostomoides (Kuroda and Habe). Pliocene Shibikawa Formation (Loc. 9). Figs. 6a-b, x1.9, IGUT no. 15576-1; Figs. 7a-b, xO.9, IGUT no. 155762. Figs. 7a-b show the adult shell for comparison.

MAJIMA: Fossil Naticids bearing calcareous Opercula

Plate 70

779. Fossil Naticids bearing calcareous Opercula
Mem. Muroran Inst. Tech., vol. 4, no. 1,
p. 1-109, pIs. 1-8.
Schileyko, A. A. (1977): Data on the anatomyof naticoidea and problems of taxonomyof the subfamily (Mollusca: Mesogastro·poda). Trans. Inst. Oceanology, vol. 108, p.
77-97 (in Russian with English abstract).
Shibata, H. (1970): Molluscan fauna of thefirst Setouchi Series, Southwest Japan.
Part 1, Fauna of the Ichishi Group. Jour.
Earth Sci., Nagoya Univ., vol. 18, no. 1,
p. 27-84, pIs. 1-4.
Taki, I. (1934): On Polinices (Neverita) didyma(Bolten). Japan. Jour. Malac., Venus, vol. 4,
p. 224-234 (in Japanese).

373

Wissema, G. G. (1947): Young Tertiary and
Quaternary Gastropoda from the Island of
Nias (Malay Archipelago). Doctor's Thesis,
Rijksuniversiteit Leiden, p. 1-212, pIs. l--u
[Not seen].
Woodring, W. P. (1957): Geology and paleontology of the Canal Zone and adjoining partsof Panama. U.S. Geol. Surv., Prof. Paper,
306-A, p. 1-145, pIs. 3-23.
Wrigley, A. (1949): English Eocene and Oligocene Naticidae. Proc. Malac. Soc. London,
vol. 28, no. 1, p. 10-30.
Yamaguchi, S. (1958): Explanatory text of thegeological map of Japan. "Monzen" sheet.
Scale 1 :50,000. 20p., Geol. Surv. Japan(in Japanese with English abstract).

Age ~~, Anden ~B3, Atsuga J!1.1f., Byakutan 13m, Chikagawa ;ITJ/I, Fusakina 7""=\'-7,
Gojome :nj$;§, Hosokomata·zawa ffiII/N5!:iR, lana :n's!l!f, Iriai :nair, Kabato fli!P, Kategarulk=¥;lJj, Kobinaizawa IJ\.JtFt:JiR, Kubusu ~I!M~, Misato ~£, Monzen r~lltr, Nei ~~, Oi jdl)J,
Okuwa (Omma) :k~, Ozato :kIlL Porokaoshirarika IjlJUJn~sflj1.JJ1, Sai /lr, Saru·guB i1J;-mEflll,
Shibikawa .~.(iJ/l, Shimajiri !IbJJL, Shin·totsugawa fJj'-[-W:JII, Sunagomata 1i'Y'f- J(., Suttsu 1.fim,
Taneume fjt/ij, Umiue ifaJ::, Yatsuo j\Ji€~~.~~~#~~*nMm~e~~ff1m(R~~~.~KM~~~AI*,c~15~JmR!ltl\ I?, 2400f*~~~.~~>:r{~~ t.: ~ ~;lt 1*nMm~

m, Cryptonatica janthostoma, C.

clausa, C. janthostomoides, C. adamsiana, C. ichishiana, "Naticarius" sp. :t.; J: '(J Natica
vitellus 7 :f!1!>:r¥IH~i'~ ~ 0 *Mj!t~:fm, Cryptonatica ichishiana :t.; J: '(J MacNeil (1961) K.
J: ~ -c Naticarius cf. N. niasensis~nt.:t:pM1!ttl\ I?$fM1!t~:f!1!, "Naticarius" sp. ~c

2 flt~itfi, .: ~ffifUJt-c-, fJJ~-Cmi1i'~ n ~o~cnl!:~Ut:fi'tt::.1il:~-C, i'X~ 5 "?~f€f.~f*:(f.~ type tI;~~l?n~o Type (1): ~'i, 1*~~!1!!:t::.*:(f.~n -c~,~ (18l'-*)0 Type (2): ~H:t, nl!:f{ft::.;I;~-C.&:~L., .:t~?H'{ITiHi,
1;:!tf{ft::.iJjjL. -C~,~ (31(*)0 Type (3): ~'i, bi'tI,t::.f4;:JfiFt:Jt::.J... I),: A.,t':~1m"(', m!nl!:t:l~I::'-i!cL.-C~'~ (1~*)0 Type (4): ~'i, nl!:t:lITiit::.~ffii-c-, .:t~Htr-¥$~nl!:t:ltl\l?~
i!1±l L. -C~\~o ~~Ft:J~ITiifi, Ft:J1S'1::.ITiiL. -C~,~ (11(*)0 Type (5): ~fi, il\ttl'I?7'G~I::'/l$n-C~'~ 01(*)0cc~ ~ ;ll1 MtI;?EA.,t':II~~.liJiJe.$~~~fi, Type (2)(3) ~.~~ililit::.mJ[;~ ~.: tI;
I±l*~o Type (2) Ii, .liJi~$~'i&tI\I?~i!t':L.t.:c i!~~~~~t::., "*-t.: Type (3) fi, .liJi~$~il\tKU~~ci!~~~~~K, .:tn~n;l;-C~~l?n~o~.~-Trans. Proc. Palaeont. Soc. Japan, N.S., No. 134, pp. 374-387, pis. 71-74, July 30, 1984

780. SOME DINOFLAGELLATE CYSTS FROM THE NANGGULANFORMATION IN CENTRAL JAVA, INDONESIAKAZUMI MATSUOKA
Department of Geology, Faculty of Liberal Arts, Nagasaki University
Nagasaki, 852

Abstract. Some dinoflagellate cysts were found from the Paleogene Nanggulan
Formation distributed near Yogjakarta in Central Java, Indonesia. Thirteen species
belonging to nine genera of Gonyaulacales were described and classified under the
Lineage System. They include following four new species; G/aphyrocysta circularis
and G. dentata of Ceratioid Lineage, and Exochosphaeridium reticulatum and E.
brevispinosum of Gonyaulacoid Lineage.

Introduction

Geological Setting

Paleogene dinoflagellate microplaleontology
has been much progressed around the North
Atlantic Ocean and Europe. Now several biostratigraphic works have been published on thebasis of these dinoflagellate cysts (e.g. Bujak
et aI., 1980). On the contrary, around South
East Asia there have been only a few papers
dealt with Paleogene dinoflagellate cysts (Jainand Dutta, 1978; Kar, 1979; Kar and Saxena,
1981; Jiabo, 1978; He and Quian, 1979).
Some interesting dinoflagellate cysts andother algal fossils were preliminarily reported
from the Nanggulan Formation in Central Java,
Indonesia (Matsuoka in Saito ed., 1981). Further detailed study on these materials reveals
that this cyst assemblage contains several new
species. One of them assignable to the genus
Danea, which was temporarily identified as
Danea? sp. (Matsuoka in Saito ed., 1981) has
been already described by the writer (Matsuoka,
1983). The other dinoflagellate cysts containing four new species and two new combinations
are described in this paper.

The Nanggulan Formation is well known as
being one of the oldest formations in the Java
Island, Indonesia. This formation is mainly
exposed around Mt. Mujle located about 3.5
km northwest of the Nanggulan Village near
Yogjakarta (Text-fig. 1).
Around the Nanggulan area, following Cenozoic formations are distributed in ascendingorder; the Nanggulan Formation, Old Andesite
Formation, Jonglangan Formation, Sentlo Formation, and unsorted debris derived from the
Old Andesite Formation (Pumamaningshi andHarsono, 1981; Wartono, Sukandarrumodi and
Rosidi, 1977).
The Nanggulan Formation revised by Pumamaningshi and Harsono (1981) consists mainlyof marl, siltstone and sandstone intercalatedwith several lignite seams, nodules and pyroclastic layers. Hartono (1960, 1969) reported
on the occurrence of Hantkenina, a planktonic
foraminifera, from this formation being Late
Eocene in age. Pumamaningshi and Harsono(1981) carried out the biostratigraphic study

374

375

780. Dinoflagellate Cysts from Java
llO'E

114'E

J AVA
5 E A

8'S;-------i~--------------~--~~~i?~~~------~---+~~~~-Text-fig. la

Nt
YOGJAKARTA___ __ o~3Km'====-~==:l

1:.:':;:.:;1 1 I!; ~I 2 ~ 3 ~~::J 4 0

5 I",," 16 1:+;1 7

Text-fig.1. 1a: Index map showing Yogjakarta District. 1b: Geological sketch map showing
Nanggulan Area by Wartono, R. et aZ. with modification.
1; Nanggulan Formation, 2; Old Andesite Formation, 3; Sentlo and Jonglangan Formations,4; unsorted debris, 5; pyloclastics of Mt. Merapi, 6; Andesite, 7; Diorite.

of this formation exposed along the Kali Seputihnear Kalisongo. They recognized five planktonic
foraminiferal zones and concluded that thisformation ranges from the Middle Eocene to
Oligocene age.
The Old Andesite Formation consists ofandestic breccias, tuff, aggomerate and andestic
lava, and its geologic age is estimated to be
Late Oligocene (Purnamaningshi and Harsono,1981) or Early Miocene (Kadar, 1981).
The Jonglangan Formation is composed ofreef limestone, marls, sandstones interbedding
lignite seams, while the Sentlo Formation ismade up of marls, limestones, sandstone andconglomerate. The lower part of the Sentlo
Formation is isochronous with the Jonglangan
Formation (Kadar, 1981).
The Lithology and Geologic Age of the Nang·
gulan Formation exposed in the Kari Puru
SectionThe sampling area is located near the Kalisongo Village and on the eastern flank of the low
mountains composed of andestic rocks belonging
to the Old Andesite Formation.
On the basis of the lithologic description of376

Kazumi MATSUOKA

the Nanggulan Formation in this section carriedout by Saito et al. (in Saito, ed., 1981), following
three lithologic units are recognized; the lower
part is composed mainly of alternation of sandy
siltstone and siltstone intercalated with lignite
seams and molluscan beds; the middle part ismade up of alternation of calcarenite, sandstoneand sandy siltstone and characterized by thepresence of abundant nodules and such larger
foraminifers as Discocyclina; the upper partcomprises alternation of silty sandstone andfine sandstone, and characterized by such pyroclastic layers as lapilli tuff and tuffite, andturbidite structure.
Okada (in Saito, ed., 1981) reported theoccurrence of calcareous nannofossils from the
middle to upper parts of the Kali Puru section,and on the basis of these species association
concluded that the middle part of the Nanggulan
Formation exposed in this section ranges fromCP 13 to CP 14 (Middle Eocene) of Okada andBukry (1980), and the upper part is probably
Middle Oligocene in age.

Systematic Paleontology
The following species are described in this paper.
Baticasphaera explanata (Bujak) comb. nov.
Baticasphaera micropapillata Stover 1977
Kallospheridium sp.
Caligodinium sp. cf. C. amiculum Drugg 1970
Glaphyrocysta circularis sp. nov.
Glaphyrocysta dentata sp. nov.
Diphyes spinulum (Drugg) Stover and Evitt
1978

Exochosphaeridium reticulatum sp. nov.
Exochosphaeridium brevispinosum sp. nov.
Operculodinium uncinispinosum (De Coninck)

comb. nov.
Homotryblium sp. cf. H. oceanicum Eaton
1976

Homotryblium pallidum Davey and Williams
1966

Homotryblium plectilum Drugg and Loeblich
1966

Class Dinophyceae
Order Gonyaulacales Taylor 1980
Ceratioid Lineage

Materials and Preparation Method

Canningia GroupThe lithologic features of the samples provided for this study are recorded in Saito (ed.,1981). The dinoflagellate cysts described here
were obtained from the samples of the lower
to middle part in the Kali Puru section. Therefore, as previously mentioned, these materials
containing the dinoflagellate cysts are middle
Eocene in age on the basis of calcareous nannofossil and planktonic foraminiferal evidences.
The sample number such as NG 39 coincides
with the numbers recorded on the figures of
the columnar section and the route map of
the Kali Puru section in Saito (ed., 1981).
The palynological preparation method used
for this study is generally the same as Matsuoka(1974). The specimens illustrated in this paper
are deposited in the palynological collection of
the Department of Geology, Faculty of Liberal
Arts, Nagasaki University.

Genus Baticasphaera Drugg 1970
Type species: Baticasphera compta Drugg 1970

Baticasphera explanata (Bujak) comb. nov.
PI. 74, Fig. 81980 Chytroeisphaeridia explanata Bujak, p. 44,
pI. 13, figs. 13-14.

Description: The proximal cyst is basicallysubspherical to ovoidal in shape, but occasionallymodified by folding. The autophragm is thin,transparent and smooth or very weakly granulated. The apical archeopyle has six or sevensided principle archeopyle parasutures. Thesefeatures suggest that six paraplates and one
anterior sulcal paraplatelet might be present
in the cingular and sulcal regions of this species.
The operculum is detached.

780. Dinoflagellate Cysts from Java
Dimensions: cyst diameter 39-42 pm.
Number of specimens measured: 5.
Remarks: Bujak in Bujak et al. (1980) described this species bearing an apical archeopyle
as belonging to the genus Chytroeisphaeridea.
But according to the diagnosis of the genus
Chytroeispharidea emended by Davey (1969),
this genus is characterized by possession of a
precingular archeopyle. Therefore, this species
must be reattributed to the genus Baticasphaera.
The specimens collected from the Nanggulan
Formation are smaller than the original English
specimens in cyst diameter.
Baticasphaera micropapillata Stover 1977
PI. 74, Fig. 9

1977

Baticasphaera micropapillata Stover, p.
73, pI. 1, figs. 7-8.

Description: The small spherical cyst is
occasionally modified by folding and provides
with thin, transparent and finely granulated
autophragm. The archeopyle is a large apical
type. The operculum is detached.
Dimensions: cyst diameter 26-47 pm.
Number of specimens measured: 5.
Remarks: The morphological features of theJavanese specimens are well concordant withthe original description. This species differsfrom Baticasphaera explanata (Bujak) in possessing a finely but clearly granulated cyst surface.
The previously known age of this species is
middle Miocene in the Atlantic core hole (Stover,1977).

Genus Kallosphaeridium De Coninck 1969
Type species: Kallosphaeridium
batum De Coninck 1969brevibar-

Kallosphaeridium sp.
PI. 74, Fig. 10

Description: The proximate cyst has a subspherical to ovoidal shape. The autophragm isthin and provided with short and slender hairlike projections closely spaced and evenly dis-377

tributed. The archeopyle is an apical type andhas six to seven princeple archeopyle parasutures. The operculum attached with one or
two paraplate boundaries. JUdging from princeple and accessory archeopyle parasutures,
this species might consist of six or seven precingular paraplates including an anterior sulcal
paraplate. No other feature indicating a paratabulation is observed and any ornamentation representing the cingulum and sulcus is lacking.
Dimensions: cyst diameter 30 pm, length of
hair-like projections 2-4 pm.
Number of specimens measured: 1.
Remarks: The present species is closely
similar to Baticasphera hirsuta Stover 1975in showing halo effect of dense hair-like projections, but differs from the latter in possession of an attached operculum. Kallosphaeridium
brevibarbatum De Coninck differs from thisspecies in having a larger cyst body and a thicker
autophragm.

Genus Caligodinium Drugg 1970
Type species: Galigodinium amiculum Drugg
1970
Caligodinium sp. cf. C. amiculum Drugg 1970
PI. 74, Fig. 11

Cf. 1970 Galigodinium amiculum Drugg, p. 815,
Figs. 8A-B; 9A-E.
Description: The small proximate cyst is
broadly elongate on lateral view and composedof the autophragm only. Its surface is smoothor finely granulate. The cyst body is partiallysurrounded by a hyaline and adhesive material.
The archeopyle is an apical type, and the operculum is partially attached and composed of
one large and two smaller paraplates. Except
for the archeopyle, there is not any feature
indicating the paratabulation.
Dimensions: length of cyst 30-34 pm, width28-30 pm.
Number of specimens measured: 3.
Remarks: The present species is closely

Kazumi MA TSUOKA

378

similar to Caligodinium aniculum Drugg except
for its smaller cyst size. This species is different
from C. granulatum (Jiabo) Lentin and Williams
in having the smooth or finely punctate cyst
surface.
Cyclonepherium Group
Genus Glaphyrocysta Stover and Evitt 1978
Type species:
Glaphyroeysta
retiintexta(Cookson 1965) Stover and Evitt 1978 = eyelonepherium retiintextum Cookson 1965

Glaphyrocysta circularis sp. nov.
PI. 71, Figs. 2-5

Derivation of name: Latin, circulus (dim.
circus), a ring; with reference to the circular
trabecurae in distal margin of annulate complexes.
Diagnosis: Intermediate skolochorate cyst
subcircular to elliptical in dorso-ventral view,
and occasionally having a small antapical boss.
Processes mostly solid and slender, and occurring as annulate, soliate or linear complexes
except for a few paraplate areas. Each complex
mostly consisting of five to seven processes.
Distal part of processes furcated and united
by slender and flexous trabeculae with smoothsurface and short spines. Archeopyle apical,
tetratabular .
Holotype: Slide NG 39-5 (4xI), PI. 71, Fig. 2.
Sample no. NG 39, Nanggulan Formation,
Middle Eocene. Location; Kalisongo near Nanggulan, 20 km northwest of Yogjakarta, Central
Java, Indonesia.
Dimensions: Holotype; cyst diameter 67.2
J.1m, length of processes 20.2 J.1m. Other specimens; cyst diameter 37-68 J.1m, length ofprocesses 13-21 J.1m.
Number of specimens measured: 5.
Description: The cyst body consisting of athin autophragm is lenticular and bears occasionally a small antapical boss. The existence of anapical boss is unknown, because any complete
specimen was not encountered. The surface ofcyst shows a coarse granUlate to reticulatefeature. Processes are solid and slender, andoccur as annular, soliate and/or linear complexes.
Most paraplate areas possesses an annular or
soliate complexes made up of five to sevenflexous and slender processes, but the 3" paraplate is represented as a linear complex consisting of three slender processes. The 6", 1'"
and 6'" paraplates are often devoided of any
process, but rarely are reflected by simple
process much reduced. The antapical 1"" complex comprises about ten slender processes.
The distal extremities of processes are furcate
and united by a simple slender but flexous
trabeculae with short spines or rarely weakly
denticulate margin. Two peripheral processgroups are conspicuously developed. They
are made up of the 1", 2" and 2'" complexes,
and 4" and 4 '" complexes which joint distallywith somewhat larger trabeculae. Other processcomplexes are soliate. The cingular and sulcal
areas are lacking in any process-like ornamentation. The archeopyle is an apical type withseven princeple archeopyle sutures. The paratabulation is indicated by the complexes andthe apical archeopyle; 4'(?), Oa, 6", Oc, as,
6" ',Op, 1" ".
Remarks: This species resembles Glaphyrocysta intricata (Eaton) Stover and Evitt, G.
retiintexta (Cookson) Stover and Evitt, and?G. reticulosa (Gerlach) Stover and Evitt inhaving simple processes. G. circularis sp. nov.,
however, is distinguishable from G. retiintexta(Cookson) and G. intricata (Eaton) in constantly possessing the 3" process and two conspicuous process groups made up of jointing
trabeculae, and from ?G. reticulosa (Gerlach)in having the 3" linear complex and only one
antapical boss.

Glaphyrocysta dentata sp. nov.
PI. 71, Fig. 1

Derivation of name: Latin, dentatus, provided with teeth; with reference to the denticulate margin of trabeculae in distal part ofprocesses.

780. Dinoflagellate Cysts from Java
Diagnosis: Intermediate skolochorate cyst
subcircular to elliptical outline in dorso-ventral
view. Relatively thin autophragm reticulate.
Processes tubiform to buccinate, stout anddistally furcate, and occurring as annulate,soliate and linear complexes. Each complex
consisting of three to five processes, and distallyunited by trabeculae with deeply dentate margin.
Most complex solitary distally. Apical archeopyle
tetratabular. Operculum free.
Holotype: Slide NG 39-5 (2fIII), PI. 71, Fig.
1, sample no. NG 39, Nanggulan Formation,
Middle Eocene. Location; Kalisongo near Nanggulan, 20 km northwest of Yogjakarta, Central
Java, Indonesia.
Dimensions: Holotype; cyst diameter 64
11m, length of processes 25 11m. Other specimens;
cyst diameter 41-68 11m, length of processes
17-26 11m.
Number of specimens measured: 8.
Description: The dorso-ventrally lenticular
cyst is lacking in any apical projection. Processes
are relatively broad and tubiform to buccinate,
and occur as annular, soliate and linear complexes consisting of three to five processes.
Distal extremities are furcate and united byflexous lod-like trabeculae with deep dentate
margin at out side. Most process complexes
are distally united within a single complex,
and become soliate. The apical area has fourannulate complexes distally united with trabeculae in each other. The precingular area
bears five complexes. The 3" paraplate is represented by a linear complex made up of
three processes. The 6" paraplate is devoidedof any process. The postcingular area consistsof five complexes. The 1'" paraplate is a smaller
annulate complex of two processes. The linear
complex developed in the dorsal side reflects
the 3" paraplate. The 2", 4" and 5" paraplates
are reflected by large annular complexes. The
broad space between 1'" and 5" paraplates
might indicate the 6'" paraplate and sulcal
area. The cingular ana sulcal areas are lacking
in any process-like ornamentation, but theanterior sulcal paraplatelet is indicated by
two accessory parasu tures. The apical archeopyle379

possesses seven main archeopyle parasuturesand several accessory arheopyle parasutures.
The infered para tabulation is indicated by
the distribution of process complexes andthe apical archeopyle; 4', 6", xc+xs, 6''', xp,
1" ".
Remarks: The present new species is similar
to Glaphyrocysta circularis sp. nov., but differsfrom the latter in having process complexes
consisting of fewer (three to five) and tubiform
to buccinate processes, and deeply dentate
margin of trabeculae at out side.
Hystrichosphaeroid Lineage
Polysphaeridium Group
Genus Diphyes Cookson 1965 emend.
Davey and Williams 1966
Synonym: Lingulosphaera Drugg 1970
Type species: Diphyes colligerum (Deflandre
and Cookson 1955) Cookson 1965 = Hystrichosphaeridium colligerum Deflandre and Cookson
1955

Diphyes spinulum (Drugg 1970)
Stover and Evitt 1978
PI. 73, Figs. 4-7
1970
1978

Lingulosphaera spinula Drugg, p. 817,
figs. lOG, llD-E, 12A-B.
Diphyes spinulum (Drugg) Stover andEvitt, p. 39.

Description: The specimens of Diphyes
spinulum (Drugg) collected from the Nanggulan
Formation have a spherical to subsphericalcyst body with finely granulated periphragm.
Processes consists of two types; a single large
antapical process and many slender processes.
The antapical process basically shows a boxlike shape with four to five sided base and ishollow or closed distally. The distal extremitiesof this process bears a few tubes. Slender processes are many, hollow, mostly tapering or
long subconical but occasionally bifurcate ona half way along their length. The distal tipsof them are mostly capitate but rarely oblateor bifid. Around the apical archeopyle andKazumiMATSUOKA380

the antapical process, groups composed of
two or three slender processes occur regularly.
This feature strongly suggests that slender
processes are not nontabular but basicallyintratabular. The archeopyle is an apical typewith five to six princeple archeopyle parasutures and occasionally accessory archeopyleparasutures. The operculum is free.
Dimensions: cyst diameter 47-45 J.1m, lengthof large processes 20 J.1m, width of large process
17 J.1m, length of slender processes 10-19 J.1m.
Number of specimens measured: 12.
Remarks: Diphyes spinulum (Drugg) closely
resembles D. colligerum (Deflandre and Cook-son) in cyst diameter and morphology of antapical process. But the former is different fromthe latter in having occasionally bifurcate processes. Though some Indonesian specimens sometimes have bifid or oblate distal tips, they arealso commonly characterized by possession ofbifurcate slender processes.
Gonyaulacoid Lineage
Operculodinium Group
Genus Exochosphaeridium Davey, Downie,
Sarjeant, and Williams 1966
Type species: Exochosphaeridium phragmites

Davey, Downie, Sarjeant, and Williams 1966
Exochosphaeridium reticulatum sp. nov.

Pi. 72, Figs. 1-5?1979 Homotryblium sp. Kar, p. 35, pi. 4, figs.
73,74.
Derivation of name: Latin, re ticulatus, netlike, with reference to the reticulate surface of
cyst.
Diagnosis: Cyst subspherical to ovoidal,
composed of two layers, periphragm and endophragm appressed between processes. Periphragm clearly reticulate and endophragmsmooth. Processes fibrous and tubiform, andbasically intratabular but apparently non tabular.
Distal part of processes patilate to denticulate
and sometimes recurved. Precingular archeopylewith pentagonal margin; operculum detached.
Holotype: Slide NG 43-2 (1gIV), PI. 72,
Fig. 1, sample no. 43, Nanggulan Formation;
Middle Eocene. Location; Kalisongo, 20 km
north west of Yogjakarta, Central Java, Indonesia.
Dimensions: Holotype, length of cyst 66 J.1m,
width 54 J.1m, length of processes 13-18 J.1m.
Other specimens; length of cyst 66-88 J.1m,
width 54-81 J.1m, length of processes 13-20 J.1m.
Number of specimens measured: 10.
Description: The proximochorate cyst has asubspherical to ovoidal shape. The periphragmis relatively thick and reticulate, and the endophragm is thin and smooth. The reticulationof periphragm are clearly observed at the basal
part of processes. Processes derived from the

Explanation of Plate 71
Fig. 1. Glaphyrocysta dentata sp. nov.
a: ventral surface, b: optical corss section of dorso-ventral view showing the antapical boss,
c: dorsal surface showing the linear complex 3" paraplate.
Holotype, slide NG 39-5 (2f1II).
Figs. 2-5. Glaphyrocysta circularis sp. nov.
2a: optical cross section of dorso-ventral view, 2b: ventral surface.
Holotype, slide NG 39-5 (4xI).
3: optical cross section of dorso-ventral view.
Slide NG 39-3 (4ell).
4a: optical cross section of dorso-ventral view, 4b: lateral surface.
Slide NG 49-3 (12011).
5: free operculum showing four apical complex para plates.
Slide NG 49-3 (12xll).
All figures x 625.

Plate 71

MATSUOKA : Dinoflagellate Cysts from Java1a2a. -. , .

1b

2b

."
1c

3

G

,
4b

5

780. Dinoflagellate Cysts from Java

periphragm are fibrous, somewhat flexous andslender tubiform, and have similar length within
a single specimen. Apical and antapical processes are not differentiated and similar to otherprocesses. The distal parts of processes show
patulate to finely denticulate and occasionallyrecurved. Their distribution of processes appears
to be non tabular especially on ventral side, butaround the precingular archeopyle and on thefree operculum, processes has basically intratabular or penitabular arrangement. These
processes are sometimes reduced and become
to be fibrous, linear complex forms. The paracingulum is occasionally represented as a linear
arrangement of processes especially beneaththe precingular archeopyle, but on the ventral
side, it is unclear as well as the parasulcus. The
archeopyle formed by displacement of thethird precingular paraplate is pentagonal inshape. The free operculum provided with about
fifteen processes on it.
Remarks: Exochosphaeridium reticulatum sp.
nov. is characterized by possession of reticulate
periphragm and fibrous processes being basicallyintratabular or penitabular. This species resembles Exochosphaeridium bifidum (Clarke
and Verdier 1967) Clarke et al. 1968 in bearing
a fibro-pitted periphragm and fibrous processes,
but differs from the latter in having larger
number of processes, and lacking in conspicuously branched apical process. The processesof E. reticulatum might be hollow, but it isobscure because of slender and fibrous stalks.
Exochosphaeridium brevispinosum sp. nov.
PI. 72, Figs. 6-9

Derivation of name: Latin, brevis + spinosum;
short spine, with reference to the short processes.
Diagnosis: Cyst spherical to ovoidal andintermediate in size. Cyst wall composed of
two layers, periphragm and endophragm appressed between processes. Processes strongly
fibrous, and short and widely cylindrical. Their
distribution fundamentally intratabular, but
apparently non tabular. The wall of processes
becoming loose. Distal extremities of processes

381

dentiCUlate. Archeopyle precingular; operculum
free.
Holotype: Slide NG 33-2 (6gI), PI. 72,
Fig. 7, sample no. NG 33, Nanggulan Formation;
Middle Eocene. Location; Kalisongo, 20 km
northwest of Yogjakarta, Central Java, Indonesia.
Dimensions: Holotype; length of cyst 69 J.1.m,
width 63 J.1.m, length of processes 9 J.1.m. Other
specimens; length of cyst 53-69 J.1.m, width
53-63 J.1.m, length of processes 7-9 J.1.m.
Number of specimens measured: 5.
Description: The proximochorate cyst is intermediate in size and spherical to ovoidal in shape.
The periphragm is moderately thick and strongly
fibrous. The fibro-pitted structure is developedaround the proximal base of processes. The
endophragm is thin and smooth. The processes
composed of several strings are short and widely
cylindrical and seem to be hollow. Their walls
show a reticulate structure and their distal
extremities are apparently denticulate. The
intra tabular distribution of processes is clearly
indicated around the archeopyle. About fifteenprocesses are developed in one para plate area
based on the detached operculum. The processes
are similar in length and structure within asingle specimen. The cingulum is occasionally
represented by a linear distribution of theprocess beneath the basal parasuture of the
precingular archeopyle. Except for the archeopyle, the paratabulation is unclear. The archeopyle might be derived from loss of 3" paraplatebased on the shape of the free operculum.
Remarks: This species is similar to Exochosphaeridium reticulatum n. sp. in having afibro-pitted periphragm and fibrous processes,
but distinguishable from the latter in possessing
shorter and wider, and strongly fibrous cylindrical processes. Nevertheless the intermediate
forms were encountered in the material of the
Nanggulan Formation.
Genus Operculodinium Wall 1967
Type species: Operculodinium centrocarpum(Deflandre and Cookson 1955) Wall 1967 =Hystrichosphaeridium centrocarpum Deflandre
and Cookson 1955.

Kazumi MATSUOKA

382
Operculodinium uncinispinosum(De Coninck 1969) comb. nov.
PI. 73, Figs. 1-3
1969
1970

Cordosphaeridium uncimspmosum De
Coninck, p. 32-33, pI. IX, figs. 6-8.
C. uncinispinosum De Coninck; Gruns·
Gavagenetto, pI. 1, fig. 13.

Emended diagnosis: Intennediate to large
cyst spherical to ovoidal. Wall consisting of twolayers, fibro·pitted periphragm and endophragm
closely appressed between processes. Processes
with a smooth stalk, weakly fibrous, long,
slender, and apparently non tabular. Processes
rarely bifurcate or multifurcate on a way. Distal
extremities of processes mostly recurved. Pre·
cingular archeopyle pentagonal in shape; operculum completely free.
Dimensions: cyst diameter 75-108 p.m,length of processes 37-52 p.m, processes ratio

0.5.
Number of specimens measured: 5.
Description: The proximochorate cyst islarge to intennediate in size and subsphericalto ovoidal in shape. The periphragm is fibropitted and relatively thick, and the endophragmis thin and smooth. Processes composed ofseveral strings of the periphragm and uniform
in length, weakly fibrous, slender, flexous
and apparently nontabular, and bear a smooth
stalk. The proximal part of processes clearlyshow a pitted structure, and the distal extremities of them are apparently capitate, but
actually divided into several aculiate or recurved
strings. Some of processes are rarely bifurcate
on a way or multifurcate near the base. The
simple precingular archeopyle without any
accessory archeopyle sutures might be madeup of displacement of the third precingularparaplate. The operculum is completely detached.

Explanation of Plate 72
Figs. 1-5. Exochosphaeridium reticulatum sp. nov.
1a: ventral surface, 1 b: dorsal surface showing the precingular archeopyle and the reticulate
periphragm between processes.
Holotype: slide NG 43-2 (lgIV), x 580.
2a 0 : optical cross section of lateral view, 2b 0 : lateral surface.
Slide NG 27-4 (2oIV), x 386.
3: processes showing the recurved distal extemity and the fibro-pitted structure observed at
proximal base.
Slide NG 45-3 (6qIV), x 1400.
4: free operculum showing intratabular processes.
Slide 39-4 (6QIV), x 800.
5°: dorsal view showing the precingular archeopyle somewhat deformed, and fibrous andintratabular processes.
Slide NG 25-3 (5eIV), x 500.
Figs. 6-9. Exochosphaeridium breuispinosum sp. nov.
6ao : optical cross section showing strongly fibrous processes, 6b 0: cyst surface.
Slide 33-2 (lAIII), x 500.
7ao: dorsal view showing the precingular arch eo pyle , 7b v : optical cross section of dorsoventral view showing short and fibrous processes.
Holotype: slide NG 33-2 (6qI), x 500.
8: free operculum showing intratabular processes.
Slide NG 43-1 (7PII), x 500.
9: free operculum showing the reticulate periphragm between processes.
Slide NG 35-2 (6nIII), x 500.
0: in interference contrast

MATSUOKA: Dinoflagellate Cysts from Java

Plate 72

6b

,
..,
~

2b

W'
A

•
II

~

8

9

780. Dinoflagellate Cysts from Java
Remarks: The morphological characteristics
of Javanese specimens are well concordant withthe original description of Cordosphaeridium
uncinispinosum De Coninck 1968. But thisspecies has been kept to be a problematicalspecies of Cordosphaeridium by Stover andEvitt (1978), because its archeopyle is unclear.
The Javanese specimens have a simple precingulararcheopyle. Therefore this species is transferedfrom Cordosphaeridium to Operculodinium.
Eighteen species belonging to the genus
Operculodinium have been described. Amongthem, the following species have an intermediate
to large cyst body and relatively long slender
processes; Opercuiodinium divergens (Eisenack
1954) Stover and Evitt 1978, O. cf. hirstum(Ehrenberg 1836) Gocht 1969 and O. tiara(Klumpp 1958) Stover and Evitt 1978. The
present species differs from former two species
in having a fibro-pitted periphragm and characteristic processes with a smooth stalk andmostly recurved distal extremities, and is also
different from O. tiara (Klumpp) in bearing
processes with recurved distal tips and lacking
in thick felt-like periphragm.

Homotrybloid Lineage
Genus Homo trybliu m Davey and Williams 1966
Type species: Homotryblium tenuispinosum
Davey and Williams 1966
Homotryblium sp. cf. H. oceanicum Eaton 1976

383

denticulate or aculeate distal extremities. They,
however, slightly differs from the specimens
described from the Bracklesham Beds in southern
England in having slenderly cylindrical to somewhat tapering processes and the finely but clearly
granular periphragm between processes.

Homotryblium pallidum Davey and Williams 1966

PI. 74, Fig. 1
1966

1978
1980

Homotryblium pallidum Davey andWilliams, p. 102-103, pI. 12, figs. 4-6,
text-fig. 22.
H. pallidum Davey and Williams; Jain
and Dutta, p. 107, pI. 1-12, 13.
H. pal/idum Davey and Williams; Bujak
et aI., pI. 1, fig. 1-3.

Dimensions: cyst diameter 55-70 Jl.m, lengthof processes 20-30 Jl.m.
Number of specimens measured: 5.
Remarks: Homotryblium pallidum Davey andWilliams is different from other species in having
a thin endophragm and a finely granular tosmooth periphragm, and in possessing a circular
proximal base of slender processes with variable
distal extremities. The specimens of the Nanggulan Formation are larger in cyst diameter thanthe previously described specimens.
The previously known range of this species
is Early Eocene to Late Oligocene (Davey andWilliams, 1966; Costa and Downie, 1979; Bujak
et aI., 1980).

PI. 74, Fig. 2
Homotryblium plectilum
Drugg and Loeblich 1967

Cf. 1976 Homotryblium oceanicum Eaton, p.
268, pI. 10, figs. 5-8.
Dimensions: diameter of cyst body 54-60
Jl.m, length of processes 15-19 Jl.m, width ofprocesses 4-5 Jl.m.
Number of specimens measured: 2.
Remarks: The specimens collected from the
Nanggulan Formation are mostly incomplete
without the epicystal part. These specimens arecharacterized by the epicystal archeopyle andintratabular and hollow processes provided with

PI. 74, Figs. 3-7
1967

Homotryblium plectilum Drugg and
Loeblich, p. 184, pI. 2, figs. 1--9, text-fig.
3.

H. plectilum Drugg and Loeblich;
Williams, pI. 5, fig. 3.
1978 H. plectilum Drugg and Loeblich; Jain
and Dutta, p. 107, pI. I-II.
1980 ?Homotryblium floripes (Deflandre andCookson) Stover; Bujak in Bujak et aI.,1975

KazumiMATSUOKA384
p. 64, pI. 16, figs. 2-3.
1981

Oligosphaeridium complex (White) Daveyand Williams; Kar and Saxena, p. 115, pI.
4, fig. 79.

Dimensions: cyst diameter 65-80 Jlm, lengthof processes 19-24 Jlm.
Number of specimens measured: 7.
Remarks: The following three species of thegenus Homotryblium have characteristic intratabular processes composed of two to six tubes
closely jointing on the way;H. floripes (Deflandre
and Cookson) Stover 1975, H. plectilum Drugg
and Loeblich 1967, and H. vallum Stover 1977.
Bujak (1980) explained that H. plectilum Drugg
and Loeblich is a younger synonym of H. floripes(Deflandre and Cookson). But these two species
have clearly different character concerning aperiphragm, that is, H. {loripes (Deflandre andCookson) bears relatively thick and irregularly
scabrated, apparently granular periphragm, while

H. plectilum Drugg and Loeblich is characterized
by being thin and smooth to finely and delicately
granulated periphragm.
H. vallum Stover differs from these two
species by having smaller cyst and taeniate
processes. But Stover (1977) mentioned thatH. plectilum Drugg and Loeblich and H. vallum
Stover occur together and transitional forms
were observed.
The specimens obtained from the Nanggulan
Formation are identical to those of H. plectilum
Drugg and Leoblich owing to possession oflarger cyst body, thin and finely granular tosmooth periphragm and longer processes.
A dinoflagellate cyst described as Oligosphaeridium complex (White) by Kar and Saxena(1981) should be assignable to Homotryblium
plectilum Drugg and Loeblich on the basis of
the number of processes and its poly tubular
processes. These specimens were recovered

Explanation of Plate 73
Figs. 1-3. Operculodinium uncinispinosum (De Coninck 1968) comb. nov.
1~: oPti~al cross section. of lateral view showing the elongate cyst body, 1 b: surface of lateralvlew,lc : process showmg the slender form and the furcate form which have smooth surfaceof stalks.
Slide NG 27-5 (liIlI), la, lb x 320, 1c x 1250.
2a: dorsal surface showing the precingular archeopyle, 2b: processes showing strongly
recurved strings of distal extremities.
Slide NG 1-2 (lcIV), x 320.
3a0 : free operculum showing the fibro-pitted structure between processes 3b 0 : slender,
flexous and long processes on the free operculum.
'
Slide NG 23-5 (8mIV), x 1250.
Figs. 4-7. Diphyes spinulum (Drugg) Stover and Evitt 1978
4a 0 : apical surface of apical view showing apical archeopyle, 4b 0 : optical cross section of
apical:antapical view, 4c 0 : antapical surface of apical view showing the roundly pentagonal
antaplcal process.
Slide NG 5-5 (8LII), x 500.
5a:: optic~l cross section of polar view showing the free operculum within the endocoaI.
5b : antaplcal surface showing postcingular processes.
Slide NG 7-5 (8AIV), x 500.
6a 0 : lateral surface showing the subspherical cyst body. 6b 0 : optical cross section of lateralview showing the large antapical process.
Slide NG 5-5 (701), x 500.
7.0 : lateral view showing the much deformed apical archeopyle and basically intratabular precmgular processes.
Slide NG 7-5 (8AIV), x 1250.
0: in interference contrast

MA TSUOKA: Dinoflagellate Cys ts from Java../

Plate 73

.,

~

'.

1b

,
If

~

.... ______ 4a _____

4b4c_ _ _--'6b

780. Dinoflagellate Cysts from Java
from the Middle to Late Eocene sediments
in southern Kutch in India.
Acknowledgements
I am much indebted to Prof. T. Saito and
Dr. H. Okada of Yamagata University, andDr. D. Kadar of Geological Research and Devel·
opment Center in Bandung for their kind guidance in the field work in Indonesia. I also wish
to thank Dr. J. P. Bujak of Petro Canada for
his taxonomical suggestion, and Prof. K. Takahashi at Nagasaki University and Prof. S. Nishida
at Nara University of Education for their criticalreading of the manuscript.
The field survey in Indonesia was supported
by Japanese Ministry of Education under Grantin-Aids for Oversea Scientific Survey (Leader;Prof. T. Saito, Grant No. 504308).

References
Bujak, J. P., Downie, C., Eaton, G. L. andWilliams, G. L. (1980): Dinoflagellate
cysts and acritarchs from the Eocene ofSouthern England. Spec. Paper, Palaeontology, 24, 100 pp. 22 plates, 24 text-figures.
Costa, L. I. and Downie, C. (1979): Cenozoic
dinocyst stratigraphy of Sites 403 to 406(Rockall Plateau) IPOD, LEG 48. In
Montadert, L. & Roberts, D. C. (ed.), Initialreport of the Deep Sea Drilling Project,(U.S. Government Printing Office), Washington, 48, p. 513-529, plates 1-3.
Davey, R. J. (1976): Non·clacareous microplankton from the Cenomanian of England,
northern France and North America. Part 1.
British Museum, (Nat. Hist.) , Bull., Geol.,
17 (3), p. 103-180, plates 1-11.
and Williams, G. L. (1966): The genus
Hystrichosphaeridium and its allies: In
Davey, R. J., Downie, C., Sarjeant, W. A.
S. and Williams, G. L., Studies on Mesozoic
and Cainozoic dinoflagellate cysts. British
Museum, (Nat. Hist.), Bull., Geol. Suppl.
3, p. 53-106.
De Coninck, J. (1969): Dinophyceae et Acritarcha de L'Ypresien de Sondage de Kallo.
Belgique Inst. Royal Sci. Nat. Mem., 161,385

p.1-67,platesl-17.
Drugg, W. S. (1970): Some new genera, species
and combinations of Phytoplankton fromthe lower Tertiary of the Gulf Coast, U.S.A.
Proc. North Amer. Paleont. Convention
Part G, p. 809-843, figs. 1-19.
'
and Loeblich, A. R. Jr. (1967): Some
Eocene and Oligocene phytoplankton fromthe Gulf Coast, U.S.A. Tulane Studies
in Geology, 5 (4), p. 181-195, plates 1-3.
Eaton, G. L. (1976): Dinoflagellate cysts fromthe Bracklresham Beds (Eocene) of theIsles of Wight, southern England. British
Museum, (Nat. Hist.) , Bull. Geol., 26 (6),p. 225-332, plates 1-21.
Gruns-Gavagnetto, C. (1970): Dinophyceae,
acritarcha et pollens de la formation de
Varengeville. Rev. Micropaleontol., 13, p.
69-78, plates 1-3.
Jain, K. P. and Dutta, S. K. (1978): Lower
tertiary dinoflagellate spores and pollen
grains from Siang district, Arunachal Pradesh.
Jour. Palaeont. Soc. India, 21 & 22, p. 106111, plates 1-2.
Jiabo (1978): On the Paleogene dinoflagellates
and acritarchs from the coastal region ofBohai. Nanking Institute of Geology andPalaeontology, Acad. Sinica, p. 1-190,
plates 1-49.
Hartono, H. M. S. (1960): Hantkenina in the
Nanggulan Area. Publ. Tehnik Seri Paleontologi, no. I, p. 3-7.
- - (1969): Globigerina marls and their planktonic foraminifera from the Eocene ofNanggulan, Central Java. Cush. Found.
Foram. Res. Contr., 20 (4), p. 152-159,
plates 20-21.
He Cheng-Quan and Quian Ze-Shu (1979):
Early tertiary dinoflagellates and Acritarchs
from the Bose Basin of Guangxi. Acta
Palaeontologica Sinica, 18 (2), p. 171-188,
plates 1-2.
Kada~,
D. (1981): Planktonic foraminiferal
biostratigraphy of the Miocene-Pliocene
Sentlo Formation, Central Java, Indonesia.
In Saito, T. (ed.) Micropaleontology, petrology and lithostratigraphy of Cenozoic
rocks of the Yogjakarta region, Central Java,
p. 35-47, Special Publication of Yamagata
University.
Kar, R. K. (1979): Palynological fossils from the386

Kazumi MATSUOKA

Oligocene sediments and their biostratigraphy in the District of Kutch, western
India. The Palaeo botanist, 6, p. 16-49,
plates 1-4.
Kar, R. K. and Saxena, R. K. (1981): Palynological investigation of a bore core near
Rataria, Southern Kutch, Gujart. Geophytology, 11 (2), p. 103-124, plates 1-4.
Matsuoka, K. (1974): Some plant microfossils
from the Miocene Fujiwara Group, Nara,
Central Japan. Trans. Proc. Palaeont., Soc.
Japan, N.S. 94, p. 319-340, plates 44-46.
(1981): Dinoflagellate cysts and Pediastrum
from the Nanggulan and Sentlo Formationsin the middle Java Island, Indonesia. In

Saito, T. (ed.) Micropaleontology, petrology
and lithostratigraphy of Cenozoic rocks of
the Yogjakarta region, Central Java, p. 4852, Special publication of Yamagata University.
(1983): A new dinoflagellate cyst (Danea
heterospinosa) from the Eocene of Central
Java, Indonesia. Rev. Palaeobotan. Palynol,
40, p. 115-126.
Okada, H. (1981): Calcareous nannofossils of
Cenozoic formations in Central Java. In
Saito, T. (ed.) Micropaleontology, petrology
and lithostratigraphy of Cenozoic rocks of
the Yogjakarta region, Central Java, p. 2534, Special Publication of Yamagata Uni-

Explanation of Plate 74
Fig. 1. Homotryblium pallidum Davey and Williams 1966
a: apical surface showing the epicystal archeopyle. b: antapical surface showing postcingularand antapical processes, c: optical cross section of the taeniate process.
Slide NG 5-1 (6GI), la, 1b x 500, 1c x 1250.
Fig. 2. Homotryblium sp. cf. H. oceanicum Eaton 1976
a 0 : upper focus of ventral surface showing precingular and postcingular processes, b 0 : lower
focus of ventral surface. co: optical cross section of dorso-ventral view.
Slide NG 1-1 (5WIV), x 500.
Figs. 3-7. Homotryblium plectilum Drugg and Loeblich 1967
3a: oblique apical surface showing the epicystal archeopyle, 3b: optical cross section ofpolar view, 3c: oblique antapical surface.
Slide NG 5-4 (6cIV), x 500.
4: lateral view showing taeniate and poly tubular processes.
Slide NG 49-5 (3r11), x 540.
5: oblique apical surface showing the epicystal archeopyle.
Slide NG 5-4 (5NIII), x 500.
6: oblique apical surface showing the epicystal archeopyle.
Slide 5-4 (7011), x 500.
7: polytubular process.
Slide NG 5-4 (9cll), x 1250.
Fig. 8. Baticasphaera explanata (Bujak 1980) comb. nov.
apical surface showing the apical archeopyle.
Slide NG 5-3 (3hll), X 500.
Fig. 9. Baticasphaera micropapillata Stover 1977
a: optical cross section, b: oblique apical view showing the apical archeopyle.
Slide NG 49-1 (8DIV), X 670.
Fig. 10. Kallosphaeridium sp.
a 0 : lateral view showing the attached operculum, b 0 : optical cross section of lateral view.
Slide NG 21-2 (lkIlI), X 500.
Fig.11. Caligodinium sp. cf. C. amiculum Drugg 1970
lateral view showing the hyaline and adhesive material.
Slide NAG 25-4 (lJIII), x 500.
0: in interference contrast

MATSUOKA: Dinoflagellate Cysts from Java

Plate 74

· 1c
1a

1b

2a

2b

2c

~

.'""

..
•

.

I

•

3b

.I

-I

3c

•
",

4

5

11

780. Dinoflagellate Cysts from Java
versity.
Okada, H. and Bukry, D. (1980): Supplementary
modification and introduction of code
numbers to the low-latitude coccolith biostratigraphic zonation (Bukry, 1973; 1975).
Marine Micropaleontology, 5 (3), p. 321325.
Purnamaningshi, S. and Harsono, P. (1981):
Stratigraphy and planktonic foraminiferaof the Eocene-Oligocene Nanggulan Formation, Central Java. Publication of Geological Research and Development Center,
Paleontology series, No.1, p. 9-28.
Saito, T. (ed., 1981): Micropaleontology, petrology and lithostratigraphy of Cenozoic
rocks of the Yogjakarta region, Central
Java. Special Publication of Yamagata
University, 61 pp.

-1 :/ r* '/ 7""'$;;

387

Stover, L. E. (1977): Oligocene and early Miocene dinoflagellates from the Atlantic Core
hole 5/5B Blake Plateau. In Elisk, W. C. (ed.)
Contributions of stratigraphic palynology,
A.A.S.P. contr. Ser. no. 5A, p. 66-83,
plates 1-3.
and Evitt, W. R. (1978): Analysis of prePleistocene organic-walled dinoflagellates.
Stanford Univ., publ., Geol. Ser., 15, iii+300
pp.
Wartono, R., Sukandarrumidi and Rosidi, H. M.
D. (1977): Geological map of the Yogjakarta
Quadrangle, Java. scale 1 :100,000, Geological Survey of Indonesia.
Williams, G. L. (1975): Dinoflagellate and spore
stratigraphy of the Mesozoic-Cenozoic, offshore eastern Canada. Geol. Surv. Canada,
paper 74-30 (2), p. 107-161, plates 1-8.

l' ? £T:/ fl'- =j :/~7H)O)tlNI.,€liVi'/ 7. ,. : -1 :/ f*'/ 7<j:riffi;;l'?a;;~9;;1'~~~D~~~~~~~~~*T:/y-=j:/OO~~.~~~~~=*=j,97.
~t;:.jt~~9a13~ (?~4tfifii) ~~ttLt.:o .:ttL!?I:!:, ~=j7-?.J,,*ljIJO)Glaphyrocysta
circularis sp. nov., G. dentata sp. nov. c ":);' =* =j , 9 7.*ljUO) Exochosphaeridium reti~1lliJ~1t:
culatum sp. nov. :W~U E. brevispinosum sp. nov. -C~~oAddendum:
After this manuscript had been accepted, Chytroeisphaeridea explanata Bujak was replaced in
the genus Baticasphaera on the basis of its apical arch eo pyle by Islam (1983). Therefore, thisspecies should be described under the following name; Baticasphaera explanata (Bujak) Islam,
1983.
Reference;
Islam, M. A. (1983): Dinoflagellate cysts from the Eocene Cliff sections of the Isles of Sheppey,
southeast England. Rev. Micropaleontol., 25, p. 231-250, plates 1-4.

Trans. Proc. Palaeont. Soc. Japan, N.S., No. 134, pp. 388-392, pIs. 75, July 30, 1984

781. PRIMITIVE FUSULINACEA FROM DANGYOKEI OF TAISHAKU(STUDIES OF THE STRATIGRAPHY AND THE MICROFOSSIL FAUNASOF THE CARBONIFEROUS AND PERMIAN TAISHAKU LIMESTONE IN
WEST JAPAN, No. 5)*
KIMIYOSHI SADA, KAZUYOSHI NOMURA AND YUKIMASA OHO
Faculty of Integrated Arts and Sciences, Hiroshima University,
Hiroshima 730

Abstract. This is the fifth of the serial papers on the stratigraphy and themicrofossil faunas of the Carboniferous and Permian Taishaku Limestone in Hiroshima Prefecture of western Japan, which treats of the primitive fusulinacean
faunas in the type section of the Dangyokei Formation and its overlying limestone
beds belonging to the Eimyoji Formation at Dangyokei in the Taishaku area.

Introduction
No fusulinacean faunas have been reported
from the stratotype of the Dangyokei Formation (Yokoyama, 1957; Hase et al., 1974; Sada,
1975a, 1977) and its overlying lower part of
the Eimyoji Formation at Dangyokei in the
Taishaku Limestone upland located in Tojocho of Jinseki-gun, Hiroshima Prefecture, western Japan. However, recently we newly discovered the Eostaffella fauna in the Dangyokei
Formation and Millerella-Eostaffella fauna inthe basal part of the Eimyoji Formation at
Dangyokei. This discovery is worthy of notefor the biostratigraphic zonation and correlationof the Dangyokei Formation called by Yokoyama (1957). In this paper the faunas are discussed and Eostaffella kanmerai (Igo), whichis representative species of the faunas, is described.
The field work was financed by the grants
from the Japanese Government Ministry of
Education_

Text-fig. 1 Location of the Taishaku Limestone in Hiroshima Prefecture,
western Japan.
Stratigraphy and discussion of faunas
The Carboniferous of the Taishaku Limestone
in the central part of this upland is divided into
the Dangyokei Formation and the Eimyoji
Formation (Yokoyama, 1957; o kimura, 1966;
Hase et al., 1974; Sada, 1975a, 1977)_ Regarding the fossils from Dangyokei, in 1944 Huji-*Received May 23, 1983; read June 26, 1982
at Sapporo.
388

Fusulinacea from the Taishaku Limestone389

Text-fig. 2. Map showing the fossil locality at Dangyokei in the Taishaku Limestonearea, Hiroshima Prefecture. (Fossil locality plotted on Geographical Survey Institute ofJapan topographical map "Taishakukyo" of scale 1 :25000.)
moto reported a species of Carboniferous coral,
Nagatophyllum sp., from Dangyokei and thought
it to be of Visean age. After that, Yokoyama(1957) described Lithostrotionella taishakuensis
Yokoyama, Clisiophyllum o{ukuensis (Ozawa)and C. awa Minato from the type section of
the Dangyokei Formation and entertained theopinion that the coral fauna might be of the
Bashkirian_ The stratotype of the Dangyokei
Formation is composed of schalsteins (basaltic
lava, tuff and hyaloclastite) and an about 3 m
thick tuffaceous limestone lens which is madeof the crinoidal biosparrudite. The overlying
Eimyoji Formation consists of oolitic massive
limestone at Dangyokei.
The limestone lens at the type section (Loc.
A 78-1) of the Dangyokei Formation contains
Eostaffella kanmerai (Igo), Endothyra, smaller
foraminifers, crinoids and algae. On the other
hand, the oolitic massive limestone (Loc. A782) of the lower part of the Eimyoji Formationis characterized by Millerella sp., Eostaffellakanmerai (Igo), Endothyra, smaller foraminifers,
crinoids and algae.
The Eostaffella kanmerai fauna of the Dangyokei Formation is quite similar to the
Chesterian Eostaffella kanmerai fauna (Sada,
1969) in limestones cropping out along the
Tojo River near the town of Tojo, and also to
the Chesterian Eostaffella faunas described by
Kanmera (1952) from the Kakisako Formationin Kyushu and by Igo (1957) from the Ichinotani Formation in Gifu Prefecture_ Furthermore
the fauna can be correlated with the Russianz

g~ ~ h--'-r--'---r--l<E--A78-2 llvlillerellasp.
Eostaffella kanmerai~~

Endothyra
Smaller foramir,ifers
Crinoids

lJ.

~

~ ~

i

Algae

~

w

:i
"

z

"

"

0

t--

«
~

IX
0
lJ.

Eostaffella kanmerai

"

,. "

" " ..t!.
A 78-1 Endothyra'c" I--T...J,-;-'...l.-'r-r~0[ Smaller foraminifers

N

Crinoids
BryozoaW'"> .!2

0

<!l

z

«

0

~

Algae

~

'" "
0

lu

Schalstein""

Tuffaceous
limestone

Ooli lie
limestone

Text-fig. 3. Columnar section of the lower
part of the Taishaku Limestone exposed on thesouth side of the Taishaku Gorge in the Taishaku
area, Hiroshima Prefecture.

Kimiyoshi SADA et al.

390

Eostaffella faunas (Rosovskaya, 1963; Aisenberg
et al., 1960) and North American Eostaffella
faunas (Zeller, 1953; Anisgard and Campau,
1963; Sada and Danner, 1973). The Millerella
sp. fauna of the Eimyoji Formation resembles
in specific composition the early Pennsylvanian
Millerella faunas in Japan (Igo, 1957; Sada,
1964, 1975b, 1975c, 1980) and North America.

Description
Family Ozawainellidae
Thompson and Foster, 1942
Genus Eostaffella Rauser-Chemoussova, 1948
Eostaffella kanmerai (Igo)
PI. 75, Figs. 1-8

Millerella kanmerai Igo, 1957, p.175-177, pI.
1, figs. 20-26, pI. 2, fig. 14.

Eostaffella kanmerai, Sada, 1964, p. 230-231,
pI. 21, figs. 8, 16-17; 1967, p. 144-145,

pI. 2, figs. 1-10; 1969, p. 120-121, pI. 12,
figs. 1-13, pI. 13, figs. 1-2; Sada and Danner,
1974, p. 259-266, pI. 37, figs. 1-3, 5, 1819; Sada, 1975c, p. 35-36, pI. 8, figs. 1-5;
1980, p. 68-69, pI. 1, figs. 1-5, 11-13.

Material studied. -Description is based on 5
illustrated specimens (Rg. No. UHA 78-2Aa,
UHA78-2Ac, UHA78--2A13, UHA78-1w, andUHA78-1A and many other unillustrated
specimens from Loc. A78-1 and A78-2.
Descriptive remarks. -The shells are discoidal
in shape, having the subangular to the broadly
rounded periphery and the umbilicated poles.
The shells of the four to five volutions are 172
to 280 /1 in length and 571 to 704 /1 in width.
The form ratios range from 0.30 to 0.40. The
outer volutions are involute. The diameters of
the whorls of the 2nd to the 5th volution are
129-138, 244-266, 394-430 and 649-707 /1,
respectively. The spirotheca consists of a tectum
and inner and outer tectoria. The thickness ofTable 1. Measurements of Eostaffella kanmerai (Igo)
Specimens

UHA78-2Aa

UHA78-2Ac

UHA78-2A13

PI. , fig.

1,1

1,4

1,2

1,7

1,8

Length

0.285

0.236

0.295

0.306

0.286

UHA78-1w

UHA78-1A

Width

0.707

0.649

0.682

0.656

0.730

Form ratio

0.40

0.36

0.43

0.47

0.39

2

0.138

0.129

3

0.266

0.244

4

0.430

0.394

5

0.707

0.649

Diameter of

Vol.

whorl

Thickness of0

spiro theca

0.014

5

0.021

0.016

0.012

0.019

0.022

0.015

0.028

0.023

Measurements in rnrn.

Fusulinacea from the Taishaku Limestone

the spirotheca of the 1st to the 5th volution is14, 16-21, 12-19, 15-22 and 23-28 11, respectively.
The symmetrical chomata are low. The
present species is very poor in preservation
because of the secondary mineralization. In
the shell shape, the number of volution andthe internal biocharacteristics, the species,
however, resembles closely Eostaffella kanmerai
described by Igo (1957) from the Ichinotani
Formation in Gifu Prefecture, by Sada (1967,
1969, 1975c, 1980) from many places in westernJapan and by Sada and Danner (1974) from
British Columbia in Canada.
Occurrence. -Eostaffella kanmerai (Igo) is~~~d~~&~~raand~al~~aminifers at Loc. A78-2.
Geological age. -Chesterian to early Pennsylvanian.
References
Aisenberg, D. E. et al. (1960): Carboniferous
stratigraphy of the Donetz basin. Compte
Rendu, tome 1, p. 1-12.
Anisgard, H. W. and Campau, D. E. (1963):
Paramillerella thompso:1i, n. sp. from
Michigan and a redefinition of Paramillerella.

Contrib. Cushuman Found. Foram.
search, vol. 14, no. 3, p. 99-108.

Re-

Hase, A., Okimura, Y. and Yokoyama, T. (1974):
The Upper Paleozoic Formation in andaround Taishaku-dai, Chugoku Massif, southwest Japan, with special reference to thesedimentary facies of limestones. Geol. Rep.
Hiroshima Univ., no. 19, p. 1-39.
Hujimoto, H. (1944): Paleontological study on
Titibu-System of the Taishaku-Kyo region,
Hiroshima Prefecture. Studies Geol. Min.
Inst. Tokyo Bunrika Daigaku, no. 1, p.
1-19.
Igo, H. (1957): Fusulinids of Fukuji, southwestern part of the Hida Massif, central
Japan. Tokyo Univ. Educ. Sci., Ser. C, vol.
5, no. 47, p. 153-246, pis. 1-15.
Kanmera, K. (1952): the Lower Carboniferous
Kakisako Formation of southern Kyushu,
with a description of some corals andfusulinids. Mem. Fac. Sci. Kyushu Univ.,

391

Ser. D, vol. 3, no. 4, p. 157-177, pis. 8-l2.
Okimura, Y. (1966): Microbiostratigraphicalstudies on the foraminiferal faunas of theLower Carboniferous Formation of the
Chugoku region, Southwest Japan. Geol.
Rep. Hiroshima Univ., no. 15, p. 1-46
(in Japanese with English abstract).
Rosovskaya, S. E. (1963): The fossil types offusulinids and their ancestors. Akad. Nauk.
SSSR, Trudy Paleont. Inst., tome 97, p.
1-128.
Sada, K. (1964): Carboniferous and Lower
Permian fusulines of the Atetsu Limestone
in West Japan. Jour. Sci. Hiroshima Univ
Ser. C, vol. 4, no. 3, p. 225-269, pis. 21 ~28.
- - (1967): Fusulinids of Millerella zone of the
Taishaku Limestone (Studies of the stratigraphy and the microfossil faunas of the
Carboniferous and Permian Taishaku Limestone in West Japan, no. 1). Trans. Proc.
Palaeont. Soc. Japan, N S. no. 67, p. 139147, pis. 12-13.
--(1969): Microfossils of the lowest part of
the Taishaku Limestone (Ibid., no. 4).
Ibid., no. 75, p. 119-129, pis. 12-13.
- - (1975a): Late Mississippian and early
Pennsylvanian fusulinid faunas of the
Taishaku Limestone in West Japan. Bull.
Soc. belg. Geol. no. 1, p. 5-9.
--(1975b): Early and middle Pennsylvanian
fusulinacea from Akiyoshi. Mem. Fac.
Integr.

Arts and Sci.,

Hiroshima

Univ.

vol. 1, p. 1-29, pis. 1-7.
- - (1975c): Early and middle Pennsylvanian
fusulinacea from the Omi Limestone inJapan. Ibid., vol. 1, p. 31--39, pI. 1.
--(1977): Primitive fusulinacean faunas incentral and western Japan. Ibid., vol. 4, p.
17-25.
--(1980): Primitive fusulinacea from the
Nakamura Limestone in western Japan.
Jour. Paleontology, vol. 54, no. 1, p. 65-70,
pI. 1.
- - and Danner, W. R. (1973): Late Carboniferous Eostaffella and Hexaphyllia from
central Oregon, U.S.A. Trans. Proc. Palaeont.
Soc. Japan, NS. no. 91, p. 151-160, pis.
23-24.
--and - - (1974): Early and middle Pennsylvanian fusulinids from southern British

Kimiyoshi SADA et al.

392

Columbia, Canada and northwestern Washington, U.S.A. Ibid., no. 93, p. 249-265,
pis. 35-37.
Yokoyama, T. (1957): Notes on some Carboniferous corals from Taishaku district, Hiroshima Prefecture, Japan. Jour. Sci. Hiro-shima Uniu., Ser. C, vol. 2, no. 1, p. 73-S2.
Zeller, D. E. N. (1953): Endothyroid foraminifera
and ancestral fusulinids from the type
Chesterian (Upper Mississippian). Jour. Paleontology, vol. 27, no. 2, p. IS3-199, pis.
26-2S.

*fRltfi~,~ilfO)il'Il(!hrt~£E. (T:i J;R • =~;Y~WfRT:iFRZ·O)J\')IJ: C ~1t1:i1!iMtoM(:. '"?l '-C O)liJf'5\:. No.5): ;;'frfR1:iFR't:-I\'!MHtTj','I\O)WfrtM£WlO) stratotype /I, i? Eostaffella kanmerai([go) 1,(. 7cO)J:fl1:K-lfltJ: Q7k8ll=.'j,m{t(Tm~ J: I) E. kanmerai ([go) c Millerella sp. I,(-t<

ItIitM~~ 0) stratotype f:.Ct;:.n;l: "(*,1]}i!1£E.m/l~ i "') t.: a:g~ i?.n -C l ,tJ:/I,"') t.:O)"(', "'j-@]O);:O)*fr:tOJi!.H.[l]j stratotype 0) fI!f{-'\; i':k:tE I:' ~ l' L -C ~b~-C1lt~td:!;~I,("t,'"?"t,O)"(;0Qo JJ:.H\t}(J.EI:.{t("t,m:~"('il?Q Eastaf/ella kan11lerai ([go) 1,(;:;: 1:'~c~~L -C\,11{5 L, Itrr.fM~Jii1 0) stratotype 0)~1t1:ifJ.!.f~1,( Chesterian 0) Eastaf/ella {tf#~ (jjfjj!B. 1957;jt}J*.L~, 1952; ViII!. 1969; Rosovskaya, 1963; Zeller, 1953; Sada and Danner, 1973) Ie.n.:fh~;,&5JJj L t.:o~~~Qo~rn~ff·ffHm~·~~~~

Explanation of Plate 75
All XI00
Figs. I-S. Eostaffella kanmerai (Igo)
I-S. Axial sections. Rg. No. UHA7S-2Aa, UHA7S-2AI3, UHA7S-2AS, UHA7S-2Ac,
UHA7S-1n, UHA7S-1-3, UHA78-1w and UHA7S-1A, respectively.

SADA et al.: Fusulinacea from the Taishaku Limestone

Plate 75

393

PROCEEDINGS OF THE P ALAEONTOLOG ICALSOCIETY OF JAPAN

13 *i5~~~~~~ 133@]1YU~lillB'f059if- 6 )12413(13) ~Mm*~.W~~~OOM~hk (~~~~W~~O)~.*,*~~ih~W.~~tm~w

1S) 0

i~'ii9;rlm;qj / **i110)1{.lWN1·~J:.1\::1jldrJi:!:p..1t1j

@l

A

~

:FEt'dMJIf( 1,400 m !m@:l!mf1jIj1\<\*,H::.]l/J< jff!r1dtJ150}j:4'-H;9o)-~-.jill~ • 15~{~ ('1'¥IV - -- --.. -- ... -. -- ... -..... -- - - - iiffi. ~.Iml:· .IJtl1!IJ[i'f,'Larix remains from Pleistocene strata of Northeast Japan, with special reference to the
distribution of Larix in the later half of
the Last Glacial age .......... Suzuki, Keiji*m~jTJ+I~WLrrr"tI\::0)1l:fl.jf(Wo.jl.F,.7c0) 1 IWJH.UF~:fr~Ilifr' !b'I*J&j'f-

Miocene plants from the Koura Formation in
Shimane Prefecture ............ Onoe, Toru
H:!llf!li~/J\JII Ri')Qii.j{l(fY.l1l::P'llH::."':) ~ '-C... .rr))(,i0j~~~~K~~~7~.(7~~9mO)~~~~-.... -.................. - --.. -.. " 11i'ii"t:f1Jjj;:~tifii.lnti·:;fE:::*O)

Alangium .:f1~1l::PO)jlf~H... -_ ............. - ... - ...........
'6=FJ!ilJil!ml't!i1€1:B0)*~t "/ /.j-t1lilJj:~W

Tempskya I::'",:)l'-C ...... - ..................... - .. -pq III i'f1:K
1t1.1i!5q:,HI~t'ffffJ~I~O)tt/i*,1t1j_.......... _.. j:j:J:*:;t:.. J!t:frm·~~.-·~*W~·.W.lli·WQ~~;jt:Illi~ ........ _...........

M10)1'l-t~1l-ttO)Mlt............. .

. . . . . . . . . . . . .. *H~IJI.j· litflfJ'.[IJ • ittl*:1il*
Dipteridaceae from the Upper Triassic Mine
Group in Yamaguchi Prefecture, Japan (3). .. .. . . .. . . .. .. . . . . . . .. .. . .. Naito, Gentaro~t1fii)g !yiJ~:Illi~O) J: ~B ilE*O)j:j1..i.¥ti1::iffL!±!. ('-y-¥a) ................. _. _.. ---... -... 1fii1¥;m:;t:*Jii:Wli. E3m iHJ t(JI!;tH:i!l 0) 11\JII bfU#£ J: I) ~-t ~:::::'1;H~ Spllmellaria ...... mmlJ9#~ • illlfilli2"l!:=--Upper Triassic spicular Radiolaria .. __ ... _..
· . . .. .. . . . . .. . . .. . . .. . .. Nishimura, Harumi
Triassic Radiolaria from the Kamiaso area,
Gifu Prefecture. Part I Anisian ....... _..
· ... _. . . . . . . . . . . . . . . . . .. Nishimura, Harumi
Lower Triassic conodonts from the Kamiaso
area, Gifu Prefecture .......... _........ .
· ....... Sato, Yoshitsugu and Igo, Hisaharu*~~*~w~~mmo)~~*-~~5*,.7c0)
L '7 T 1" =- 7 ~--:h - =- 7 ~lJlW-HiU:O)::J /F' ~ r ltlfln; ...................... j!f\\Ft,n Ii&:-_fj(~ 7 Iv 7, If v :h" 1" :H (Veneridae) O)~Nlj;ili,-='"':)t·-C ... _.............. _... _.. . ,Ii!,*~!k.\Vicarya vernellili, V. callosa, t" J: u'; V.
eocenica 0)11,l;~·r,~'!*1-='"':)~ '-C . _. _.. - -;r1:;!j!f~01~
Anadara (Mollusca, Bivalvia) from the Miocene Narrow Cape Formation in Kodiak
Island, Alaska ........................... .
. . . . . . . .. . Marincovich, L. and Noda, Hiroshi
Revision of the Burmese Tertiary turrids ..
.............................. Shuto, Tsugio
Late Cretaceous (Campanian and Maastrichtian) ammonites from Awaji Island, Southwest Japan. _.......... - ........ - - . iJliif!:!7jfllj

PALAEONTOLOGICAL SOCIETY OF JAPAN
LIST OF MEMBERS
June 30, 1984

1

Patron Members
ABU DHABI OIL CO. LTD. 8-1-19,
Akasaka, Minato-ku, Tokyo 107
ARABIAN OIL CO.
LTD.,
ENG.
Dept.
3-2-3,
Marunouchi,
Chiyoda-ku, Tokyo 100
CHENGBEI (NICCHU) OIL DEVELOP.
2-2-2,
Uchisaiwai-cho,
CO.
Chiyoda-ku, Tokyo 100
DIA CONSULTANT CO.
LTD.
31631, Toshima-ku, Tokyo 171
JAPAN
PETROLEUM EXPLORATION
LTD. Otemachi Bldg., 1-6CO.
1, Chiyoda-ku, Tokyo 100
INDONESIA PETROLUM CO.
15th
16, AkefuneMori Bldg.,
7F,
Minato-ku,
cho,Nishikubo,
Tokyo 105
KANTO NATURAL GAS DEVELOPMENT
CO.
LTD. Mitsui Bldg., 2-1-1,
Muro-machi,
Nihonbashi, Chuo-

ku, Tokyo 103
KITAKYUSHU MUS.
NAT. HIST. 36-1,
Nishihon-machi,
Yahatahigashi-ku, Kitakyushu 805
MITSUI OIL EXPL. CO. LTD. 1-29, Minato-ku, Tokyo 105
NIPPON MINING CO. LTD. 2-10-1,
Toranomon,
Minato-ku,
Tokyo
105
NIPPON OIL EXPL.
CO.
LTD.,
EXPL. DEPT. 1-18-1, Shinbashi,
Minato-Ku, Tokyo 105
SG GIKEN CO. 4-2-47, Kurokami,
Kumamoto 860
SUMITOMO PETROL.
DEVELOP. 2-61,
Nishishinjuku, Shinjuku-ku,
Tokyo 160
TEIKOKU OIL CO.
LTD. 1-31-10,
Hatagaya,Shibuya-ku,
Tokyo
151

Honorary Members
ASANO,
Kiyoshi 41, Kinoshita,
Sendai 983
3-23-7,
WataruHASHIMOTO,
Nerima-ku,
Higashioizumi,
Tokyo 177HUZIOKA, Kazuo 14-20, Higashimachi,
Shinfujita, Asahikawa,Akita 010
IKEBE,
Nobuo 6-4, Hama-machi,
Ashiya 659
KOBAYASHI,
Teiichi 5-50-18,
Yoyogi, Shibuya-ku, Tokyo 151
KURODA,
Tokubei Nishinomiyakaisei Hospital,
1-4,
Ohama-

cho, Nishinomiya 662
MAKIYAMA,
Jiro
22,
Tanakaharunacho, Sakyo-ku, Kyoto 606
MATSUMOTO,
Tatsuro1-28-5,
Ohashi, Minami-ku, Fukuoka 812
OZAKI,
Hiroshi
2-5-29,Amanuma,
Suginami-ku,
Tokyo
167
TAKAI,
Fuyuji 1-9-4, Obinata,
Bunkyo-ku, Tokyo 112
Geol. ,TORIYAMA,
Ryuzo Dept.
Fac.
Sci.,
Fukuoka Uni v. ,FukuokaNanakuma,
Nishi-ku,
814

Regular Members, Fellows (0)and Foreign Members (*)
ABE,
KatsumiGeol.
Inst. ,
Fac. Sci., Univ. Tokyo, Hongo,
Bunkyo-ku, Tokyo 113
ADACHI, Shuko
c/o Mr. Kawanaka,
1239, Saiki, Sakura-mura,
Niihari-gun,IbarakiPref.
300-31AlBA,
Hiroaki Seikoso, 1-172,
Nukuikita-machi,
Koganeishi 184
AlTA,
YoshiakiInst. Geol. &Paleont.,
Fac.
Sci.,
TohokuUniv., Aoba, Sendai 980
AKAGI, Saburo 5-639-RC-1-203,
Koyamacho-minami, Tottori 680
AKAI,
Kozue
8-2,
Nakatomicho, Sakado-shi 350-02
AKAMATSU, Morio Hokkaido Kaitaku Kinenkan,
Atsubetsu-cho,
Shiroishi-ku, Sapporo 061-01°AKIBA,
Fumio
Japan
Petrol.
Exp1.
Co.
Ltd., Midorigaoka,

2

Hamura-cho, Nishitama-gun,Tokyo 190-11
AKIMOTO, Kazumi 1-10-8, Eharacho, Nakano-ku, Tokyo 163
o AKUTSU,
Jun
3-1, Asahi-cho,Nishinasuno-machi, Nasu-gun,Tochigi Pref. 329-27
AMANO, Kazutaka Joetsu Univ.
Educ., Yamayashiki, Joetsu 943
o AMANO, Masahisa
739, Deyama,
Hotakubohonmachi, Kumamoto 362* AMERICAN MUSEUM NAT. HIST.,
DEPT.
MICROPALEONT. Central
Park at 79th St., New York, NY
10024, U.S.A.
ANDO, Hisao Satsukiso 202, 133, Kaname-cho, Toshima-ku,
Tokyo 171
ANDO, Yasuji
4-15-9, Taidera,
Akashi 673
AOKI, Naoaki 6-24-5-607, Maeno-cho, Itabashi-ku, Tokyo 174
o AOSHIMA,
Mutsuharu
2149-6,
Tsuruta-cho,
Utsunomiya 320
AOYAMA, Hisamoto TakachihoHigh School, Takachiho-cho,Nishiusuki-gun, Miyazaki Pref.
882-11° ARAI, Juzo
2-2-13, Ooto, Yono-shi 338
ARAKAWA, Shinji
1385-23, Sezaki-cho, Soka-shi 340
ARAKI, Hideo
Kesennuma Library, Sasanojin, Kesennumashi 988
ARIKAWA, Ryuichi Inst. Geol. &Paleont., Fac. Sci., Tohoku
Univ., Aoba, Sendai 980
ASAGA,
Masayoshi Chiba-ken
Kyoiku Center,
Katsuragi,
Chiba 280
ASAI, Akito Fujimi Haitsu, 424-7, Horie, Urayasu-shi 27201° ASAMA, Kazuo
923-7, Mogusa,
Hino-shi 191
ASAMI, Kiyohide 445, Higashiterao, Tsurumi-ku, Yokohama
230* AVECILLA-GUADARRAMA,
CarlosDept. Bio1., Fac. Sci., Ciudad
Univ., Post No. 70-568, Mexico
20, D.F., MEXICO
BAN, ShinsukeIndonesia Petro1. Co. Ltd., Mori-Bldg. 37,

3-5-1, Toranomon, Minato-ku,
Tokyo 105;< BARKER, Reginald Wright
5120,
Palmettost,
Bellaire,
TX
77401, U.S.A.
*BURCKLE, Lioyd H. LamontDoherty, Geol. Observ., Palisades, NY 10964, U.S.A.
,,< CHANG,

Li-Sho3410, Redbud
Lane, Raleigh,NC 27607,
U.S.A.
CHIBA, Noboru 3-11-25, Hatadera, Matsuyama 790° CHIJI, Manzo 10-2, Namimatsucho, Kishiwada-shi 596° CHINZEI, Kiyotaka Geo1. Inst.,
Fac. Sci., Univ. Tokyo, Hongo,
Bunkyo-ku, Tokyo 113°CHISAKA, Takeshi 3-33-16,
Shimoigusa, Suginami-ku, Tokyo167* CHUNG, Kuang-gyi Dept. Earthsci., Natl. Univ.,
Tainan,TAIWAN
CHOE,
Kae-Lim Geo1. Inst.,
Fac. Sci., Univ. Tokyo, Hongo,
Bunkyo-ku, Tokyo 113
l(-CORGAN, James Xavier Dept.
Geogr.
& Geo1., Austin
Peay
State Univ., Clarksville, TN
37040, U.S.A.

*DANNER,
Wilbert Roosevelt
Dept. Geo1., Univ. British
Columbia, Vancouver, B.C. V6T
284, CANADA* DEHM, Richard Inst. Palaeont.
& Hist. Geol., Richard Wag ernStr., 10-11, Muenchen 2, W.
GERMANY*DEOCK, Lee Jong Dept. Earth
Sci., Jeon-Buk Natl. Univ.,
Jeonju 520, KOREA*DOWLEN, Robert John Dept.
Geo1., San Diego State Col1.,
San Diego, CA 97115, U.S.A.
;< DURHAM, John Wyatt
Mus. Paleont., Univ. California, Berkeley, CA 94720, U.S.A.
EHIRO, Masayuki
Inst. Geo1.,
Fac. Gen. Educ., Tohoku Univ.,
Kawauchi, Sendai 980
ETO, Tetsuto
Geol. Inst.,
Fac. Educ., Yokohama Natl.

3

Uni v., Toki wadai, Hodogaya-ku,
Yokohama 240
ETO, Toshiharu Shukutokugakuen, 2078, Yanaizu-cho,
Hashima-gun, Gifu Pref. 591-61° FUJI, Norio
Dept. Geol., Fac.
Educ., Kanazawa Univ., l'larunouchi, Kanazawa 920
FUJII, Eiichi Meisei HighSchool,2-9-24,
Umegaoka,
Setagaya-ku, Tokyo 154°FUJII, Shoji 161-2, Anyobo,
Toyama 930
FUJISHIMA, Yasutaka Kawasaki
Chishitsu Co., 4-8-3, Chuo,
Sendai 980
FUJITA,
Ikuo Hokkaido Rika
Kyoiku Center, Miyamori, Chuo,
Sapporo 064°FUJIYAMA, Ienori Natl. Sci.
Mus., Hyakunincho, Shinjukuku, Tokyo 160
FUKADA, Atsuo 1-15-15, Honcho,
Koganei-shi, 184° FUKUDA, Osamu 2-78, Honcho,
Kumagaya-shi 360° FUKUDA, Yoshio 1585, Yotsukaido, Yotsukaido-shi 284
FUNATSU, Hiroshi Akifuchu HighSchool, Fuchu-machi, Aki-gun,
Hiroshima Pref. 735
FUNAYAMA, Hasaaki
Inst. Geol.
& Paleont., Fac. Sci., Tohoku
Univ., Aoba, Sendai 980
FURUICHI, Hitsunobu 2269, Ido,
Miki-cho,
Kita-gun,KagawaPref. 769-23
FURUKAWA,
Takaharu 4-12-23,
Higashiikuta, Tama-ku, Kawasaki 214
FURUOYA, Hiroyuki 3-13-1-405,
Okubo, Shinjuku-ku, Tokyo 160
FURUTANI, Hiroshi Dept. Earth
Sci., Fac. Sci., Nagoya Univ.,
Chikusa-ku, Nagoya 464
FUSE, Keisuke
lnst. Geol. &Paleont., Fac. Sci., Tohoku
Univ., Aoba, Sendai 980
FUTAKAMI, Masao 15-25, Higashi-cho, Tokorozawa-shi 359* GEOLOGICALSURVEY INST. P.O.
Box 1964, Teheran, IRAN
GOKAN, Masafumi
3-2-2, Kishimachi, Urawa 336

GOTO, Hiroya 380-4, Yotsuka,
Ta tsuno-cho, Ta tsuno-shi 67941
GOTO, Masatoshi
Dept. Anatom., Tsurumi Univ., Tsurumiku, Yokohama 230
GOTO, Michiharu
Toyama City
Sci. & Cult. Center, Nishinakano-machi, Toyama 930-11
GUNJI, Yukio
lnst. Geol. &Paleont., Fac. Sci., Tohoku
Univ., Aoba, Sendai 980° HABE, Tadashige 2-26-6, Motoizumi, Komae-shi 201
HADA,
Shigeki Dept. Geol.,Fac. Sci., Kochi Univ.,
Akebonocho, Kochi 780° HAIKAWA, Takehiko Akiyoshidai
Sci. Mus., Shuho-cho, l'linegun, Yamaguchi Pref. 754-05
HAMADA, Jun-ichi
1823-3, 19isue, Tonosho-cho, Shozu-gun,
Kagawa Pref. 761-42° HAMADA, Takashi
Dept. Earth
Sci. & Astron., ColI. Arts&
Sci., Univ. Tokyo, Komaba,l'leguro-ku, Tokyo 153;~HAMLIN,
William H. Atlantic
Richfield,
Box51408,
Lafayette, LA 70501, U.S.A.
° HANAI, Tetsuro
Univ. Mus.,
Univ. Tokyo, Hongo, Bunkyo-ku,
Tokyo 113
HARA, Manabu
45-4, Shimonakaji, Yamaji, Tatsuta-mura,Ama-gun, Aichi Pref. 496
HARADA, Ken-ich Dept. Earth
Sci.,
Fac.
Sci.,
Yamagata
Univ., Koshirakawa, Yamagata
990
HARADA, Koji 9727-1, Hiromachi, Kure 737-01° HASE, Akira lnst. Geol. &Miner., Fac. Sci., Hiroshima
Univ., Higashisenda, Hiroshima 730
HASE, Kotaro
Hase Geol. Surv.
Co., 3-5-8, Hon-cho, Sendai
980
HASE, Yoshitaka
Dept. Geol.,Fac. Sci., Kumamoto Univ.,
Kurokami, Kumamoto 860
HASEGAWA, Jun-ichi
9-18, Suehirocho, Sakado-shi 350-02° HASEGAWA, Shiro lnst. Geol. &4Paleont., Fac. Sci., Tohoku
Univ., Aoba, Sendai 980
HASEGAWA, Yasuo 5-3-44, Higashihoncho, Joetsu 943
o HASEGAWA, Yoshikazu
3-16-15,
Numama, Zushi-shi 249
o HASEGAWA,
Yoshiyuki Dept.
Geol., Fac. Gen. Educ., Niigata Univ., Igarashi, Niigata
950-21o HASHIMOTO,
Isamu 6-10-29,
Nagasumi, Fukuoka 815
HASHIMOTO, Kyoichi 108-2,
Isae, Hofu-shi 747
HATTA, Akio
973-9, Shimoiino,
Futtu-shi 299-12
HATTORI, Ikuko 4012, Totsukacho, Totsuka-ku, Yokohama 244
HATTORI, Koji
c/o Mrs. Sakamaki, 3-32-8, Amanuma, Suginami-ku, Tokyo 167
HATTORI, Shuichi Tetsudokodanryo, 5-17-20, Maebaranishi,
Funabashi 274
o HAYAMI,
Itaru
Univ. Mus.,
Univ. Tokyo, Hongo, Bunkyo-ku,
Tokyo 113o HAYAMI,
TomokoDept. Geol.,Fac.
Educ.,
Ehime Univ.,
Bunkyo-cho, Matsuyama 790
o HAYASAKA,
Shozo
Inst. Earth
Sci., Fac. Sci., Kagoshima
Univ., Kurimoto, Kagoshima 890
HAYASHI, Akira 1-2-502, Yoshikawa-cho, Kitakatsushika-gun,
Saitama Pref. 342
HAYASHI, Keiichi
Fussa HighSchool, Kitaden-en Fussa-shi
197
HAYASHI, Shingo 1-1227, Omamamachi, Yamada-gun, Gumma Pref.
376-01
HAYASHI, Tadaichi 2-167, Hagiyamadai, Seto-shi 489
HAYASHI, Tokue
1091, Showamachi, Ishida-cho, lki-gun,Nagasaki Pref. 811
HIGUCHI, Yu Godo Petrol. Expl.
Co. Ltd., 2-13-12, Nihonbashi,
Chuo-ku, Tokyo 103
o HIRANO,
Hiromichi
lnst. Earth
Sci., Fac. Educ.,
WasedaUniv., Nishiwaseda, Shinjukuku, Tokyo 160
o HIRAYAMA,
Katsumi 1-33-10,
Ebara-cho, Nakano-ku, Tokyo

165
HIRAYAMA, Ren
Dept. Geol. &Miner., Fac. Sci.,
Kyoto
Univ., Sakyo-ku, Kyoto 606
HISADA,
Ken-ichiro Dept.
Geol.,
Osaka Univ.
Educ.,
Minamikawabori, Tennoji-ku,
Osaka 543
HISAMITSU,
Masao Tomita,
Koromokawa-mura, lsawa-gun,lwate Pref. 029-43
HITAKA, Minoru
170, Hoso,
Sakanoshi, Oita 870-03
HONDA, Nobuyuki lust. Geol. &Paleont., Fac. Sci., Tohoku
Univ., Aoba, Sendai 980
o HONDA,
YutakaDept. Geol.,Fac. Educ., Mie Univ., Kamihama-cho, Tsu 514
HORIGUCHI, Mankichi 6-20-7,
Kamikizaki, Urawa 338
HORIGUCHI, Toshiaki Chichibu
Cement Co.
Ltd.,
1-28-23,
Hongo, Bunkyo-ku, Tokyo 113
o HORIKOSHI, Masuoki
Ocean Res.
lnst., Univ. Tokyo, Minamidai,
Nakano-ku, Tokyo 164
HORIUCHI, Junji 2734, Izumi,
Komae-shi 182
HOUJO, Yoshio 1-15-44, Wakaba,
Yahatanishi-ku, Kitakyushu 806*HU, Chung-Hung Dept. Physics,
Taiwan Normal Univ., Taipei
117, TAIWAN* HUANG, Tun-Yow Chinese Petrol.
Corp., Chung Hwa Road, Taipei
100, TAIWAN°IBARAKI, Masako 390, Marikoshinden, Shizuoka 421-01°ICHIKAWA, Koichiro 2-10-9,
Midori-machi, Horiage, Sakai
593°ICHIKAWA, Wataru 2-28-12,
lshibiki-cho, Kanazawa 520
ICHINOSEKI, Tetsuro Japan
Petrol. Expl. Co. Ltd., Midorigaoka, Hamura-cho, Nishitama-gun, Tokyo 190-11°IGO, Hisaharu
Dept. Geol.,Tokyo Gakugei Univ., Nukuikita, Koganei-shi 184
o IGO, Hisayoshi lnst. Geosci.,
Univ. Tsukuba, Sakura-mura,
Niihari-gun, Ibaraki Pref. 305
IGUCHI, Yutaka 19-10-407,

5
Otamayashita, Sendai 980
IIJIMA,
Azuma Geol.
Inst.,
Fac. Sci., Univ. Tokyo, Hongo,
Bunkyo-ku, Tokyo 113
IKEDA, Tadashi 1-24-10-202,
Shimaizumi, Habikino-shi 583
IKEHARA, Ken 1-401-211, Azuma,
Sakura-mura,
Niihari-gun,Ibaraki Pref. 305° IKEYA, Noriyuki Geosci. Inst.,
Fac. Sci., Shizuoka Univ.,Oya, Shizuoka 422
IKUJI,
Yukio Japan Petrol.
Expl. Co. Ltd., Midorigaoka,
Hamura-cho, Nishitama-gun,Tokyo 190-11° IMAMURA, Sotoji 4-163-10, Takasu, Nishi-ku,
Hiroshima 733
IMAMURA,Tadahiko 3-138,
Nishi-machi, Seto, Kochi 78102° IMANISHI,
Shigeru 3-13-20,
Koto, Kumamoto 862
INA, Haruyuki 6-83, Ichibamachi, Tokoname-shi 479
INABA, Ryoichi Izumihaitsu, 114-7, Miyakubo, Ichikawa 272*INGAVAT, Rucha Dept. Miner.
Resouro.,
Rama
VI
road,
Bangkok, THAILANDIHOMA, Akitoshi Japan Petrol.
Expl. Co. Ltd., 1-6-1, Otemachi, Chiyoda-ku, Tokyo 100° INOUE, Eiji Geol. Surv. Japan,
Yatabe, Tsukuba-gun, Ibaraki
Pref. 305
° INOUE, Masao Fac. Educ., Iwate
Univ., Ueda, Morioka 020
IHOUE, Shigehiro 6-3-263,
Kariguchidai, Tarumi-ku, Kobe
655
INOUE, Yoko 4-6-6, Hamadayama,
Suginami-ku, Tokyo 168* INST.
DE INVESTIGACIONES,GEOLOGICAS Casilla, 10465
Santiago, CHILEINUZUKA, Norihisa Dept. Anat.,Fac. Med., Univ. Tokyo, Hongo,
Bunkyo-ku, Tokyo 113
IRYU, Yasufumi Inst. Geol. &Paleont., Fac. Sci., Tohoku
Univ., Aoba, Sendai 980°ISHIBASHI,
Takeshi Dept.
Geol.,
Fac. Sci.,
KyushuUni v., Hakozaki, Higashi-ku,
Fukuoka 812

ISHIDA, Keisuke
Dept. Geol.,Fac. Gen. Educ., Tokushima
Univ.,
Minamijosanjima,
Tokushima 770
ISHIDA, Masao Geol. Surv.
Japan, Hokkaido Branch, Chuoku, Sapporo 060° ISHIDA, Shiro 250-5, Miyakecho, Iwakura, Sakyo-ku, Kyoto
606
ISHIGA, Hiroaki Dept. Geol.,
Osaka City Univ., Sugimotocho, Sumiyoshi-ku, Osaka 558
ISHIGAKI,
Takehisa Dept.
Geol., Fac. Educ., Yamanashi
Univ., Takeda, Kofu 400
ISHIGURO, Yoshihi to 5, Ni shisakuracho, Minami-ku, Nagoya457° ISHII, Atsushi 2-14-20, Oyamadai, Setagaya-ku, Tokyo 158° ISHII, Ken-ichi 4-12-10, Aoyamadai, Suita 565
ISHII, Taizo 15-5, Hotsukodanchi, Sunagawa-shi 073-01
ISHIKAWA, Toru 1-8-1, TeraoHigashi, Niigata 950-21° ISHIWADA,
Yasufumi 3-5-9,
Matsubara, Setagaya-ku, Tokyo
156° ISHIZAKI, Kunihiro Inst. Geol.
& Paleont., Fac. Sci., Tohoku
Univ., Aoba, Sendai 980
ISODA, Kiyoshi 523, Motoshimanacho,
Takasaki 370
oISOMI,
Hiroshi 1-26-26,
Komaba, Meguro-ku, Tokyo 153
ISOZAKI, Yukio
Inst. Geol. &Miner., Fac. Sci., Yamaguchi
Univ. Yoshida,
Yamaguchi 753
ITAGAKI,
Hisaharu 5-37,
Misono-cho, Mikasa-shi 068-21
ITO, Takao Murorankonan JuniorHigh
School,Konan-cho,
Muroran 051
ITO, Tomio
Dept. BioI., Fac.
Educ.,
Shizuoka Univ.,
Oya,
Shizuoka 422
ITO, Yoshihiko
Dept. Geol. &Miner.,
Fac. Sci.,
Kyoto
Univ., Sakyo-ku, Kyoto 606° ITOIGAWA, Junji
Dept. Earth
Sci., Fac. Sci., Nagoya Univ.,
Chikusa-ku, Nagoya 464
IWAI, Jun-ichi 1-18-2, Sagigamori, Sendai 980

6°IWAI, Takehiko 13, Mori-cho,
Hirosaki 036
IWAO, Yushiro
373-8,Honjo,
Honjo-machi, Saga 840° IWASAKI, Yasuhide Dept. Geol.,Fac. Sci., Kumamoto Univ.,
Kurokami, Kumamoto 860
IYOTA, Norio Inst. Geosci.,
Univ. Tsukuba, Sakura-mura,
Niihari-gun, Ibaraki Pref. 305
JIMBO,
Noboru62,
Iizukamachi, Yamagata 990
JIMBO, Yukinori
5280-316,
Higashishinozaki, Edogawa-ku,
Tokyo 133
JITSUZAKI, Goro 426, Yayama,
Oita 870-03*JORDAN, Reiner Niedersaechs.,
Landesamt.,
Bodenforsch.,
Alfred-Benz-Haus, 3 HannoverBuchholz, W. GERMANY
JUNG, Kyu-Kui Inst. Geol. &Paleont., Fac. Sci., Tohoku
Univ., Aoba, Sendai 980
KADOTA, Masahito 458-6, Sanda,
Atsugi-shi 243-03
KAGAWA, Yoshimichi Hidari 3,
11 chome, 7 jodori, Asahikawa
070*KAHLER, Franz Linsengasse 29
A-9020, Klagenfurt, AUSTRIAKAIHO, Kunio Dept. Earth Sci.,
Fac. Sci., Yamagata Univ.,
Koshirakawa, Yamagata 990
KAMADA,
Hiroshi 2-1-2-202,
Yokodai, Isogo-ku, Yokohama
235
KAMADA, Kotaro
Dept.Geol.&
Miner., Fac. Sci., Hokkaido
Univ., Kita-ku, Sapporo 060
oKAMADA,
Yasuhiko 3-9-12,
Menoto, Nagasaki 852° KAMBE, Nobukazu 2-49-11, Chuo,
Nakano-ku, Tokyo 164° KAMEl, Tadao 18, Imakaido-cho,
Matsugasaki, Sakyo-ku, Kyoto
606
KAMEYAMA, Tokuhiko 188-36,
Kusugaura, Fukuma-cho, Munakata-gun, Fukuoka Pref. 811-32
KAMIYA, Takahiro c/o Mr. Ito,
3-17-20, Koishikawa, Bunkyoku, Tokyo 112
KANAMORI, Kunio c/o Mr. Kondo,

4-20-6, Hon-cho, Nakano-ku,
Tokyo 164
KANASUGI, Hiromi Inst. Geosci., Univ. Tsukuba, Sakuramura, Niihari-gun, Ibaraki
Pref. 305
° KANAYA, Taro 1300, Kamihatsuishicho, Nikko 321
KANDA, Kaname 1607-264, Fukuoka, Kamifukuoka-shi 356
KANEKO, Atsushi 1-15-7, Fukaehon-cho, Higashinada-ku, Kobe
658
KANEKO, Sueko 3-1-16-101,
Shinkanaoka-cho, Sakai 591° KANIE, Yasumitsu Yokosuka City
Mus., Fukada-dai, Yokosuka 238° KANMERA,
Kametoshi Dept.
Geol.,Fac.
Sci.,
KyushuUniv., Hakozaki, Higashi-ku,
Fukuoka 812° KANNO, Saburo
Joetsu Univ.
Educ., Nishishiro-cho, Joetsu
943
KANO, Akihiro
Inst. Geol. &Paleont., Fac. Sci., Tohoku
Univ., Aoba, Sendai 980
KANSHA, Yoshiyuki 1-13-5,
Tomioka, Koto-ku, Tokyo 135° KANUMA, Mosaburo
6-11-13,
Kamirenjaku, Mitaka-shi 181* KAPOOR, Hari Mohan Geol. Surv.
India, Calcutta, INDIA
KASAHARA, Yoshio 1-4-22, Kirigaoka, Obora, Gifu 501-31
KASE, Tomoki Natl. Sci. Mus.,
Hyakunincho, Shinjuku-ku,
Tokyo 160
KASE,
Yasuo Choshi HighSchool, Kasuga-cho, Choshi 288° KASHIMA, Naruhiko 4-3-1-513,
Higashinagato, Matsuyama 790
KATO, George 314-11, Higashioizumi, Nerima-ku, Tokyo 177° KATO, Makoto Dept. Geol. &Miner., Fac.
Sci., Hokkaido
Univ., Kita-ku, Sapporo 060
KATO, Matajiro 76-47, Takuhoku, Shinoro-machi, Sapporo 061° KATO, Michio Inst. Earth Sci.,
Fac. Gen., Educ., Kanazawa
Univ., Marunouchi, Kanazawa
920
KATO, Norihiko 5-13-9, Masugata, Tama-ku, Kawasaki 214
KATO, Susumu Dept. Geol., Fac.

7

Educ.,
Waseda Univ.,
Nishiwaseda, Shinjuku-ku, Tokyo 160
KATO,
Yoshijiro
3-13-1-603,
Heiwagaoka, Meito-ku, Nagoya
465
KATO, Yoshinori c/o Mr. Sato,
3-36-8, Ogikubo, Suginami-ku,
Tokyo 167
°KATTO, Jiro 8-40, Masugata,
Kochi 780'~KAUFMANN, Godfrey F. P.O. Box,643, Jensen Beach, FL 33457,
U.S.A.
KAWABATA, Jun 7-15-11, Okusawa, Setagaya-ku, Tokyo 158°KAWABE, Tetsuya Dept. Earth
Sci., Fac. Sci., Chiba Univ.,
Yayoi, Chiba 260
KAWADA, Shigema
2-3-5, Maeji,
Urawa 336
KAWAGUCHI, Ken-ichi Japan
Petrol. Expl. Co. Ltd., 3-6-8,
Hongoku-cho, Nihonbashi, Chuoku, Tokyo 103
KAWAKAMI,
Hiroshi 1049-44,
Shinmei, Ago-cho, Shima-gun,Mie Pref. 517-05
KAWAKAMI, Takeshi Iwate Pref.
Mus., Ueda, Morioka 020-01
KAWAKAMI, Toru
7-22-14,Tsuchihashi, Takatsu-ku, Kawasaki
213
KAWAMURA, Yoshinari Dept.
Geol.,
Aichi Univ.
Educ.,
Igaya-cho, Kariya-shi 448°KAWANO,
Michihiro 3-54,
Takara-machi, Yamaguchi 753
KAWASAWA, Keizo
129,Iguchicho, Kochi 780
KAWASHITA,
Yo shiro 2-Al-5,
Chiyoda, Touma tsu, Mikasa-shi
068-22KAZAMA, Satoshi 989-3, Hanamuro, Sakura-mua, Niihari-gun,Ibaraki Pref. 305
KEN, Byakumei 2-31-1-Cl14,
Ooka, Minami-ku, Yokohama 232
KIKUCHI, Hiroyuki 813, Besshomachi, Minami-ku, Yokohama 232
KIKUCHI, Yoshibumi
Inst. GeoSci., Univ. Tsukuba, Sakuramura, Niihari-gun, Ibaraki
Pret. 305
KIKUCHI, Yoshiki 3-2-2, Nishi,
Kunitachi-shi 186;~KILMER,
Frank Hale Dept.

Geol., Humboldt State ColI.,
Arcata, CA 95521, U.S.A.
'~KIM,
Bong Kyun Dept. Geol.
Seoul Natl. Univ., Seoul 151,
KOREA
*KIM, Haang Mook Dept. Geol.,
ColI. Nat. Sci., Busan Natl.
Univ., Busan, KOREA°KIMURA, Tatsuaki 2-3-6,
Minamioizumi, Nerima-ku, Tokyo
177°KIMURA, Toshio 2-25-32, Nakatehara, Kohoku-ku, Yokohama
222
KINOSHITA, Tsutomu 5-20-7,
Inokashira, Mitaka-shi 181
KISHI, Ryohei 2-4-48, Asahi,
Okegawa-shi 363
KITAMURA, Tateharu 2-5-20,
Suehiro-cho, Ome-shi 198°KITAZATO, Hiroshi Geosci.
Inst., Fac. Sci., Shizuoka
Univ., Oya, Shizuoka 422
KIYOTA, Yasuyuki 844, Magodai,
Akita-machi,
Houtaku-gun,
Kumamoto Pref. 861-52°KOBATAKE, Nobuo 2-6, Nibancho, Koshien, Nishinomiya 663°KOBAYASHI, Fumio 2-6-4-1011,
Yanagikubo, Higashikurume-shi
203
KOBAYASHI, Hiroaki Izumiso
101, 2-1-21, Kitayama, Sendai
980°KOBAYASHI, Iwao
Dept. Geol.,
Fac.
Sci.,
Niigata Univ.,
Igarashi, Niigata 950-21
KOBAYASHI, Manabu 2051-1,
Nishioizumi, Nerima-ku, Tokyo
177
KOIKE,
Hiroko Dept. BioI.,
Fac. Gen.
Educ.,
Sai-tama
Univ., Shimookubo, Urawa 330°KOIKE, Toshio
Geol. Inst.,
Fac. Educ., Yokohama Natl.
Uni v., Toki wadai, Hodogaya-ku,
Yokohama 240
KOIZUMI, Hitoshi 3-3-8-301,
Inagekaigan, Chiba 281°KOIZUMI, Itaru
Inst. Geol.
Sci., ColI. Gen. Educ., Osaka
Univ., Machikaneyama, Toyonaka
560
KOJIMA, Natsuhiko Dept. Earth
Sci.,
Fac.
Sci.,
Yamagata
Univ., Koshirakawa, Yamagata

8990
KOKAWA, Shohei Dept. Bio!.,
Osaka City Univ., Sugimotocho, Sumiyoshi-ku, Osaka 558
KOMETANI, Hideo Okinawasan-iku
Junior High Shool, Asahikawa,
Nago-shi 905-01
o KOMURA,
Seiichi 63-13-303,
Aoyagi, Sayama-shi 350-13
o KONDA, Isao 960-88, Jiyugaoka,
9-jocho, Yamatokoriyama-shi
639-11KONDO, Tasuo Geo!. Inst., Fac.
Sci., Univ. Tokyo, Hongo, Bunkyo-ku, Tokyo 113
KONDO, Toshiki 4-7-18, Hozumi,
Kitashirado, Taira, Iwaki 970
o KONISHI,
Kenji Dept. Earth
Sci.,
Fac.
Sci.,
Kanazawa
Univ., Marunouchi, Kanazawa
920
o KOTAKA,
Tamio Inst. Geo!. &Pal e.o n t., Fa c. Sci., To h 0 k u
Univ., Aoba, Sendai 980
KOTO, Jiro 4-20-10, Minaminagasaki, Toshima-ku, Tokyo
171
KOUZAI, Takeshi 1682-2-15,
Tosayamada-cho,Kami-gun,
Kochi Pref. 782
KUBO, Chikahiro 10-10-A-204,
Shinmei-cho, Koriyama 963
KUGA, Naoyuki
Dept. Geol. &Miner., Fac. Sci., Kyoto
Univ., Sakyo-ku, Kyoto 606
KUJIME, Akira
Oyo Chishitsu
Chosa Co., Osaka Branch, 2-3627, Tarumi-cho, Suita 564
o KURIHARA, Kenji 2-39-3,
Nishiikebukuro, Toshima-ku, Tokyo
171
KURIHARA, Tutaka 6-33-13,
Komagome, Toshima-ku, Tokyo
170
KURODA,Tomio Nihon Graut Kogyo Co., 15-17, Sakuragaoka,
Shibuya-ku, Tokyo 150
KUROSAWA, Toshie 8, Miyahara,
Takasaki 370-12
o KUSHIMA,
Tasuo 1039-5-83,
Oshika, Shizuoka 422
o KUSUMI, Hisashi
2-14-8,Koiosako, Nishi-ku, Hiroshima 733
KUWANO, Tukio Natl. Sci. Mus.,
Hyakunincho, Shinjuku-ku,
Tokyo 160o

KYUMA, Yuko 41,
Nagasaki 850

Furu-machi,*LEMONE, David V. Dept. Geo!.
ScL, Univ. Texas, P.O. Box 3,
EI Paso, TX 79968, U.S.A.
*LING, Ch'ao-Ch'i 3, Lane 183,
Sect 1,
Hopibg E.
Road,
Taipei, TAIWAN*LING, Hsin Yi Dept. Geol., N.
Illinois Univ., Dekalb, IL
60115, U.S.A.
*LOEBLICH, Alfred R., Jr. Dept.
Earth & Space ScL,
Univ.
California, LA, CA 90024,
U.S.A.
°HABUCHI, Sei-ichi 4-18-3,
Nukuiminami-cho, Koganei-shi
184
HAEDA, Haruyoshi Geol. Inst.,
Fac. ScL, Univ. Tokyo, Hongo,
Bunkyo-ku, Tokyo 113°MAEDA, Shiro 2-3-7, Sugano,
Ichikawa 272°MAEDA, Yasuo 45, Sakuragaokacho, Nada-ku, Kobe 657°HAIYA, Seijuro Japan Petrol.
Exp!. Co. Ltd., Midorigaoka,
Hamura-cho, Nishitama-gun,Tokyo 190-11
HAJIMA, Ryuichi Inst. Geosci.,
Univ. Tsukuba, Sakura-mura,
Tsukuba-gun, Ibaraki Pref. 305
MAKINO, Toru 3-6-3, Shichirigahama, Kamakura 248
MAKISHIMA, Kunio 410-22, Nakanomoto, Ono-cho, Ibi-gun, Gifu
Pref. 501-05
HARUYAMA,
Toshiaki Inst.
Geo!., Fac. Gen. Educ., Tohoku
Univ., Kawauchi, Sendai 980
MASATANI, Kiyoshi 461-8, Kitaakitsu, Tokorozawa-shi 359°MASUDA, Fujio Inst. GeoscL,
Univ. Tsukuba, Sakura-mura,
Niihari-gun, Ibaraki Pref. 305°HASUDA, Koichiro Dept. Geo!.,Miyagi Univ. Educ.,
Aoba,
Sendai 980°HATOBA, Yasumochi Inst. Min.
Geol., Min. CoIl.,
Akita
Univ., Tegatagakuen, Akita 010
MATSUBARA, Ke~ji 4-15-12, Akatsuka, Itabashi-ku, Tokyo 175
MATSUDA, Shinya Dept. Gen.

9
Sci.,
Fac.
Educ.,
Univ.
Ryukyu, Nishihara-cho, Nakagami-gun, Okinawa Pref. 903-01
MATSUDA, Shoji 4-31, Mitsugugawahonmachi, Kofu 400
MATSUDA. Tetsuo
Dept. Geol.,Fac. Sci., Osaka City Univ.,
Sumiyoshi-ku, Osaka 558
MATSUDA. Tomoko 1576-135-5714, Kashii, Higashi-ku, Fukuoka 813
MATSUI. Seiichiro Dept. Geol.,Fac. Educ., Utsunomiya Univ.,
Minemachi, Utsunomiya 321
a MATSUKAWA.
Masaki Dept. Earth
Sci., Fac. Sci., Ehime Univ.,
Bunkyo-cho, Matsuyama 790
a MATSUKUMA.
Akihiko Natl. Sci.
Mus., Hyakunincho, Shinjukuku, Tokyo 160°MATSUMARU, Kuniteru Dept.
Geol., Fac. Educ., Saitama
Univ., Shimookubo, Urawa 338
MATSUMOTO. Tatsuya 533, Odomocho, Jozan, Kumamoto 860
MATSUNAGA. Fumio 3-2-5, Motokitakata, Ichikawa 272
a MATSUNAGA. Takashi 76, Matsugaoka, Kanagawa-ku, Yokohama
221
a MATSUO.
Hidekuni De pt. Ea r t hSci., Fac. Sci., Ehime Univ.,
Bunkyo-cho, Matsuyama 790
MATSUO. Yasuhiro 3-16-25,
Shoan, Suginami-ku, Tokyo 167
MATSUOKA. Akira Dept. Earth
Sci., Fac. Sci., Nagoya Univ.,
Chikusa-ku, Nagoya 464
MATSUOKA.
Atsushi 8-59-8,
Minamitsukaguchi-cho, Amagasaki 661
a MATSUOKA.
KazumiDept. Geol.,Fac. Gen.
Educ.,
Nagasaki
Univ., Nagasaki 852
MATSUOKA. Keiji Dept. Earth
Sci., Fac. Sci., Nagoya Univ.,
Chikusa-ku, Nagoya 464
a MATSUSHIMA. Yoshiaki
Kanagawa
Pref. Mus.,
Minaminakadori,Naka-ku, Yokohama 231
a MATSUSHITA. Susumu 30, Yoshida, Kamiadachi-cho, Sakyo-ku,
Kyoto 606
MATSUSUE.
Kazuyuki Dept.
Geol.,Fac.
Sci.,
KyushuUniv., Hakozaki, Higashi-ku,

Fukuoka 812
MATSUURA. Nobuomi 2-7-9, Kanaiwahigashi, Kanazawa 920-03
MIEDA. Toshimitsu Satsukiso,
996, Yahagi-cho, Chiba 280
MIl. Sachiko Dept. Geol. &Miner., Fac. Sci., Hiroshima
Univ., Higashisenda, Hiroshima
730
oMIKAMI, Takahiko Shoto Chuto
Kyoiku Kyoku, Minist. Educ.,
Kasumigaseki, Minato-ku, Tokyo
100
MIKI. Akio 7-221-D-1407, Shinmatsudo, Matsudo-shi 270°MIKI.
Takashi Dept. Geol.,Fac. Sci., Kyushu Univ., Hakozaki, Higashi-ku, Fukuoka 812
MIMOTO, Kenji 720-8, Kitajinzenji, Kochi 780
MINAKI. Mutsuhiko Dept. Biol.,Fac. Sci., Osaka City Univ.,
Sugimoto, Sumiyoshi-ku, Osaka
558
oMINOURA, Koji Dept. Earth
Sci.,
Fac.
Sci.,
Hirosaki
Univ.,
Bunkyo-cho,
Hirosaki036
MIYAHARA, Akihito
3-1-15,
Yanaka, Taito-ku, Tokyo 110
MIYAMOTO, Takami
Dept. Geol.
& Miner., Fac. Sci., Hiroshima
Univ., Higashisenda, Hiroshima
730
MIYASHITA, Osamu 2000, Notsudamachi, Machida-shi 194-01
MIYATA. Yuichiro Dept. Geol.,Fac. Sci., Kyushu Univ., Hakozaki, Higashi-ku, Fukuoka 812
MIYAUCHI. Toshiya Hopposyokubutsuen, Midori, Wakkanai 097
oM I Z UNO, At s u Y u k i .1 - 4 4 - 3 0,Wada, Suginami-ku, Tokyo 166
MIZUNO, Yoshiaki 259, Nagamoricho, Moriyama-ku, Nagoya
463°MIZUTANI, Shinjiro Dept. Earth
Sci., Fac. Sci., Nagoya Univ.,
Chikusa-ku, Nagoya 464
MOMOI, Kyoko 1962, Futoo-cho,
Kohoku-ku, Yokohama 222
MORI, Gunpei 454, Tomari,
Asahi-machi, Shimoniikawa-gun,Toyama Pref. 939-07
oHORI,Kei Inst. Geol. &Paleont., Fac. Sci., Tohoku

10Univ., Aoba, Sendai 980
MORI, Shinobu 1-3, Myogan,
Narumi-cho, Midori-ku, Nagoya464° MORISHITA, Akira 1-1-1-1211,
Nishigaoka, Meitou-ku, Nagoya
465
MORITA, Koji
Nishikawaso, 528-14, Nukuikita, Koganei-shi
184
MORITA, Rihito Inst. Geol. &Paleont., Fac. Sci., Tohoku
Univ., Aoba, Sendai 980
MORIYA,
Shigehiro 344-53,
Misono-cho, Kodaira-shi 187° MOROZUMI, Yoshiro Osaka City
Mus., Nat. Hist., Nagaikouen,
Higashisumiyoshi-ku, Osaka 546*HORRIS, Robert C. Geol. Dept.,
Univ. Arkansas, Fayetteville,
AR 72701, U.S.A.
° MURAl, Sadamasa 3-5-7, Midorigaoka, Morioka 020-01
MURAMATSU, Jiro 2-18-3, Nishikubo, Musashino-shi 180
MURAMATSU,
Ken-ichi 6-1-24,
Nagata-cho, Showa-ku, Nagoya
466
oMURAMOTO, Kikuo 28-137, Yayoihanazonocho, Mikasa-shi 068-22
MURAMOTO,
Tatsuo 28-137,
Yayoihanazono-cho, Mikasa-shi
068-22° MURATA, Masafumi Dept. Geol.,Fac. Sci., Kumamoto Univ.,
Kurokami, Kumamoto 860
MUTO,Asako
9-13-19, Koganehara, Matsudo-shi 270
NAGAI,
Koichi Dept.
Geol.,
Fac. Gen. Educ., Kyushu Univ.,
Ropponmatsu, Chuo-ku, Fukuoka
810
NAGAI, Setsuji 4018-3, Dokusho, Nagiso-machi, Kiso-gun,
Nagano Pref. 399-53
NAGANO, Norihiko 4-34-6, Kamiikebukuro, Toshima-ku, Tokyo
170

NAGATA, Kyoichi Japan Petrol.
Expl. Co. Ltd., Midorigaoka,
Hamura-cho, Nishitama-gun,Tokyo 190-11
oNAITO, Gentaro 9-26, Higashimatsuzaki-cho, Hofu-shi 747
NAKAGAWA, Hiroshi 3-2-3-402,

Suwa, Tama-shi 192-02
NAKAGAWA, Takao Dept. Geol.,Fac. Gen.
Educ.,
Niigata
Univ., Igarashi, Niigata 95021
NAKAGAWA, Tomio 1-4-17, Haruyama, Fukui 910
NAKAI, Hideki c/o Mr. Sasaki,
Kita-25, Nishi-IS, Kita-ku,
Sapporo 060
NAKAI, Hitoshi 2-16-12, Takanodai, Nerima-ku, Tokyo 177
NAKAJIMA, Shuichi 2-17-5,
Koraku, Bunkyo-ku, Tokyo 112
NAKAMORI, Toru Inst. Geol. &Paleont., Fac. Sci., Tohoku
Univ., Aoba, Sendai 980°NAKAMURA, Koji Dept. Geol.,
Fac.
Gen.
Educ.,
Hokkaido
Univ., Kita-ku, Sapporo 060°NAKAMURA, Manjiro 1-21-15,
Maruyamacho, Funabashi 273
NAKANISHI, Yukari Dept. Earth
Sci., Fac. Educ.,
WasedaUniv., Nishiwaseda, Shinjukuku, Tokyo 160°NAKASEKO, Kojiro 3-18-4, Momoyamadai, Suita 565
NAKASHIMA, Koichi
12-2-301,
Chiyoda, Shinnanyo-shi 746
NAKASUJI, Haruo Dia Consultant K.K., Sendai Branch, 3-617, Hon-machi, Sendai 980
NAKAYA, Hideo
Dept. Geol. &Miner.,
Fac. Sci.,
Kyoto
Univ., Sakyo-ku, Kyoto 606
NAKAZATO, Iaoru 276-4, Okawara, Hanno-shi 357
NAKAZAWA,
Katsumi 1-490,
Wakaki, Nagano 380° NAKAZAWA, Ieiji Dept. Geol. &Miner., Fac. Sci.,
Kyoto
Univ., Sakyo-ku, Kyoto 606
NAKAZAWA, Takashi 3-3-94-301,
Hibarigaoka, Hoya-shi 202
NARUSE, Takehiko 4th Techn.
Sch., Air Self-Defence Force,
Jurokken, Kumagaya 360°NASU, Noriyuki 52-1-301, Yochomachi, Shinjuku-ku, Tokyo
162°NATORI, Hiroo
Geol. Surv.
Japan, Yatabe, Tsukuba-gun,Ibaraki Pref. 305
NEMOTO, Nagayuki 95, Sakurada,
Shimoasamikawa, Hirono-machi,

11

Futaba-gun, Fukushima Pref.
979-04* NESTELL, Merlynd K. Dept.
Geo!., Univ. Texas, Box 19049,
Arlington, TX 76019, U.S.A.
NIHONGI.
Mitsutoshi Kita-2,
Nishi-22, Chuo-ku, Sapporo 064° NIITSUMA,
NobuakiGeosci.
Inst., Fac. Sci., Shizuoka
Univ., Oya, Shizuoka 422
NIKO, Shuji Dept. Geo!. &Miner., Fac. ScL, Hiroshima
Univ., Higashisenda, Hiroshima
730
NISHI, Hiroshi c/o Mr. Maeda,
2-25-25, Chihaya, Higashi-ku,
Fukuoka 813
NISHIDA, Harufumi 1-7-26-401,
Sugano, Ichikawa 272°NISHIDA,
Shiro 1-543-53,
Shikishima-cho, Nara 631° NISHIDA, Shoichi Dept. Geol. &Miner., Fac. ScL, Niigata
Univ., Igarashi, Niigata 951° NISHIDA, Tamio 1240-13, Takagi, Takagise-cho, Saga 840
NISHIKAWA, Isao 1797, Yukiotsu, Yuki-cho, Jinseki-gun,
Hiroshima Pref. 720-18
NISHIKAWA, Makoto 3-4-33,
Minamishiro-cho, Joetsu 943
NISHIKAWA, Yasuyuki 1514-98,
Hourenkuretake-cho, Nara 630
NISHIMIYA, Katsuhiko 3-5-19,
Takeda-cho, Kofu 400
NISHIMURA, Akiko 2-7-C20-102,
Shinsenrihigashi-machi, Toyonaka 565
NISHIMURA, Akira
Geol.Surv.
Japan, Yatabe, Tsukuba-gun,Ibaraki Pref. 305
NISHIMURA,
Harumi 6-26,
Miyamae-cho, Iruma-shi 358
NISHIO,
Toshio 5-10-8,
Higashioizumi,
Nerima-ku,
Tokyo 177
NISHIWAKI, Niichi Dept. Geo!.
& Miner., Fac. ScL, Kyoto
Univ., Sakyo-ku, Kyoto 606
NISHIZAWA, Hikaru 13-2, Daikan-cho, Hiratsuka 254
NISHIZAWA, Yasuo Nakanoso, 416, Shiomigaoka-cho, Chiba 281
NISHIZONO, Yukihisa Nishinihon
Gijutsukaihatsu K.I., Sanseruko 10F,
Watanabedori,Chuo-ku, Fukuoka 810° NODA, Hiroshi Inst. GeoscL,
Univ. Tsukuba, Sakura-mura,
Niihari-gun, Ibaraki Pref. 305° NODA, Masayuki 5-kumi, Fukagafuchi, Oita 870° NODA, Mitsuo 4-29-6, Ohashi,
Minami-ku, Fukuoka 815
NODA, Yoshikazu Inst. Geo!. &Paleont., Fac. Sci., Tohoku
Univ., Aoba, Sendai 980° NOGAMI, Yasuo 5-23, Harukicho, Narutaki, Ukyo-ku, Kyoto
616° NOHARA, Tomohide Dept. Geo!.,Fac.
Educ.,
Ryukyu Univ.,
Nishihara-cho, Nakagami-gun,
Okinawa Pref. 903-01
NOKARIYA, Hiroshi 444-106,
Kogane, Matsudo-shi 270
NOMURA, Hiromi 1-5-12-201,
Hunabori, Edogawa-ku, Tokyo
132
NOMURA, Kohei 14, Matsugaoka,
Kanagawa-ku, Yokohama 221° NOMURA, Ritsuo Inst. Geo!.,Fac. Educ., Shimane Univ.,
Nishikawatsu-cho, Matsue 690
NOMURA, Takamitsu 245, Naka,
Kasumigaoka-cho, Owariasahishi 488
NUMANO, Kyoichiro 1465, Ueno,
Nada-machi, Gobo-shi 649° OBA, Tadamichi Dept. Geo!.,Fac. Gen. Educ., Kanazawa
Uni v., Marunouchi, Kanazawa
920° OBATA,
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18
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Plates 59 and 60 fMatsumoto, Takahashi, Obata and Futakami:
Cretaceous nautiloids from Hokkaido-IV]
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Transactions and Proceedings of the PalaeontologicalSociety of Japan
July 30, 1984

New Series No. 134

CONTENTS. TRANSACTIONS
777. MATSUMOTO, Tatsuro, MIYAUCHI, Toshiya, KANIE, Yasumitsu, MlYATA, Yuichiro
and UEDA, Yoshiro ~ Cretaceous nautiloids from Hokkaido-V . . . . . . . . ... 335
778. TAzAwA, Jun-ichi, GUNJl, Yukio and MORl, Kei: A Visean brachiopod fauna from
the Mano Formation, Soma district, Abukuma mountains, Northeast Japan ... 347,

779. MAJIMA, Ryuichi:

Observations on occurrence of Japanese Neogene naticids(Gastropoda) bearing calcareous opercula .. . ... . . .. . ... . . . . . . . . . . . 361
780. MATSUOKA, Kazumi:

Some dinoflagellate cysts from the Nanggulan Formation incentral Java, Indonesia . . . . . . . . . . . ... ..... .. . . . . . . . . . .. .. .... 374
781. SADA, Kimiyoshi, NOMURA, Kazuyoshi and OHO, Yukimasa: Primitive Fusulinacea
from Dangyokei of Taishaku . . . . . . . . . .. _ . . . . . . . . . . . ..... . . . . . . 388
PROCEEDINGS . . . . . . . . . . . . . . . . . . . . . .... . . . . . . . . . . . ..... .. . . . . . . . 393
PALAEONTOLOGICAL SOCIETY OF JAPAN, LIST OF MEMBERS