Transactions and Proceedingsof thePalaeontological Society of Japan
New Series

No. 89Palaeontological Society of Japan
April 20, 1973

Editor:Hiroshi UJIIE

Associate editors:

Ikuo OBATA, Yasuji SAITO, Kazuhiko UEMURA

Officers for 1973 - 1974
Honorary President:

Teiichi KOB AYASHI

Presiden t : Tatsuro MATSUMOTO
Councillors (*Executive):

Kazuo ASAMA,* Kiyoshi ASANO, Kiyotaka CHI NZEI,* Taka-shi HAMADA,* Tetsuro HA NA I,* Kotora HATAI, Itaru HAY AMI,* Kametoshi
KANMERA,* Tamio KOTAK A, Tatsuro MATSUMOTO,* Tokio SHIKAM A, Tsugio
SHUTO, Yokichi TAKA YANAGI,* Toshimasa TA NAI, Hiroshi UJIIE*
Secretaries:

Wataru HASHIMOTO, Saburo KANNOExecutive Committee :General Affairs:
Membership:

Kiyotaka CHINZEI, Yasuhide IWASAKI

Kazuo ASAMA, Ienori FUJIY AMA

Finance:

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Planning:

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Publications
Transactions:

Hiroshi, UJIIE, Ikuo OBATA, Yasuji SAITO, Kazuhiko UEMURA

Special Papers:

Kametoshi KANMERA, Itaru HAY AMI, Tomowo OZAWA"Fossils":

Yokichi TAKAYANAGI, Toshiaki TAKAYAMA

Fossil on the cover is Bellerophon jonesianus DE KONINCK
from the Late Permian Akasaka Series in Gifu Prefecture.

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Trans. Proc. Palaeont. Soc. japan, N.S., No. 89, pp. 1-14, pis. 1-4, April 20, 197361l.

BIOSTRATIGRAPHIC STUDY OF

THE JURASSIC TOYORA G ROUP*PART II
HIROMICHI HIRANO
Department of Geology, Faculty of Science, Kyushu University.:;; ='- ;7 *cl!\WI"ii'Jil\fO)itT:1I\"ijf;~h'I"J1iJf)t - k"0) 2:
j!g:imNillfJ:: 'Jiff-r.Q 7 /-t T -1 r 0) oj'G, 3::c. L-"Cgg~I"II.iI\1t;j:"J:V':lfX!IlTffi1r}1I\O) Whitbian .toJ:V' Yeovilian ~lf,T Hildoceratidae 0) 6 ~ 11 Hne -? ~ ijr.W& L- t-:o YOKOYAMA (1904) fe J:.Q Hildoceras chrysanthemumc. H. densicostatum vi, k"O)5J~fiO)HHjI!c. ~h"Cv'.QJI1JWXfe-?~tfriML-t-:f.if*, [iij-l'illc.
'l'IJJE L- jJlf:ffO)15f!Ji~ c. -? t-:ot-:k"O) ~ i'iff: G < nl)iill"(:' ji, -? t-:;Q;, IllHHI.to J: V'/J{rJJm O)~;l'i~;Q'
G Harpoceras (s. str.) fe1'iJ}~h.Q"'~tO)C.!FUWiL-t-:o U1..c.viJjljfC, jfrt-:fe Hildoceras
felf1ti!IHl\IjJm~ftflj~~Tto)~ Hildoceras sp. c. L-"Ci-IHIj-L-t-: o fIt*' Haugia aff.japonica c.~h"Cv't-:to)fi, Haugia C:fit.J:< Phymatoceras ;Q;*~"(:·ji,.Qo Lillia toyoranac. ~ h"C v' t-: fill fe -? v'"C t ~~iO)fi'i*, Phymatocerasfefl1,T.Q to) c. 'l'UIf)i L- t-:o 3::~/!.- - r"(:'f~ Ght-:Jl:lff:!Jl-} I\'J, ggg:qLJlm, llfX!IlT I\"i 0)1'±1K!Zl C1tTiJ21:li\ml:~ {, RHm felf, L- "C ji,.Q
Jf Y!f 1Ji,. ill'*

0Introduction

Subgenus Harpoceras W AAGEN, 1869

This article is Part II of the biostratigraphic study of the Jurassic Toyora
Group. In Part I lithostratigraphy was
explained and Domerian ammonites were
described (HIRANO, 1971). In this part 6
genera including 11 species of Toarcian
Hildoceratidae and Hammatoceratidae
are described.

Harpoceras (Harpoceras) chrysanthemum(YOKOYAMA)
PI. 1, Figs. 1-3; PI. 2, Figs. 1-3;
PI. 3, Figs. 6-9.

1904. Hildoceras chrysanthemum YOKOYAMA;YOKOYAMA, Jour. Coll. Sci., Imp. Univ.
Tokyo, vol. 19, art. 20, pp. 11-12, pI. 2,
figs. 1-4.
1904. Hildoceras densicostatum YOKOYAMA;YOKOYAMA, ibid., pp. 12-13, pI. 3, figs.
1-2.
1947. Hildoceras (s.l.) chrysanthemum YOKOYAMA; MATSUMOTO in MATSUMOTO &ONO, Sci. Rep. Fac. Sci., Kyushu Univ.,
Ceol., vol. 2, no. 1, pp. 27-28.
1947. Hildoceras (s.l.) densicostatum YOKOYAMA; MATSUMOTO in MATSUMOTO &ONO, ibid., vol. 2, no. 1, p. 28.
1956. Hildoceras chrysanthemum YOKOYAMA;
ARKELL, jurassic Geology of the World,
Oliver & Boyd Ltd., p. 421.

Palaeontology (continued from Part I)
Subfamily Harpoceratinae
NEUMAYR, 1875
Genus Harpoceras WAAGEN, 1869

Type-species :-Ammonites falcifer SowERBY, 1820 (designated by ARKELL, 1951,
l. C. Z. N. Opinion 363, 1954).
• Received Feburuary 12, 1973; read jan.
23, 1971, at Tokyo.

1

2

Hiromichi HI RAND

1956. Hildoceras densicostatum YOKOYAMA;
ARKELL, op. cit., p. 421.
1962. Hildoceras chrysanthemum YOKOYAMA;
SATO, Mem. Soc. Ceol. France, no. 94,
p. 59.
1962. Hildoceras densicostatum YOKOYAMA;
SATO, ibid., no. 94, p. 59.
1963. Hildoceras chrysanthemum YOKOYAMA;TAKAI, MATSUMOTO and TORIYAMAed., Geology of Japan, Univ. Tokyo
Press, p. 83.
1963. Hildoceras densicostatum YOKOYAMA;TAKAI, MATSUMOTO and TORIYAMAed., op. cit., p. 83.

Types :-Here I designate one of theillustrated four specimens by YOKOYAMA (1904, pI. 2, fig. 1) as the lectotypeof H. chrysanthemum, MM. 7058a, and thethree other specimens as the paralectotypes (YOKOYAMA, 1904, pI. 2, figs. 2-4),
MM. 7059-7061. They were obtained at
loco 47.
At the same time I designate one of
the illustrated two specimens by YOKOY AMA (1904, pI. 3, fig. 1) as the lectotypeof H. densicostatum, MM. 7064a, and theother (pI. 3, fig. 2) as the paralectotype,
MM. 7065. The former was obtained at
Nishinagano and the latter at loco 47.
Material.·-Two hundreds and thirtyeight specimens are examined, of which
seventy-two specimens, MM. 7058-7061,
GK. G. 2431-2441, 2443-2455, 2457-2492,
2501-2508, were previously regarded as
Hildoceras chrysanthemum, eighty-eightspecimens, MM. 7064-7065, GK. G. 25512565A, B, 2566-2577, 2579-2584A, B, 25852610A, B-2633, as Hildoceras densicostatum, thirty-nine specimens, GK. G. 25092519, 2521-2548, as an intermediate form
and thirty-nine specimens, GK. G. 1131811322, 11356-11379, GYU. M. 1058-1067,
are newly examined.
Diagnosis ,'-'- A species of Harpoceras(s. str.) whose ribs have somewhat clear
relief, moderate curve and the haft aslong as inner one-third of the flank.
Description:- The whorls enlarge moderatelyand are somewhat evolute. The
umbilical periphery is more or less sharp
and the wall is steep but not so high.
The venter is unicarinate-bisulcate. The
ribs spring on the umbilical seam and go
on forward. At about one-third or twofifths of the flank the ribs sharply curve
backward. Near the ventral shoulder theribs recurve forward and more or lessprojected on the ventral shoulder but
soon fade away. The detailed curvatureof the ribs varies one by one, as illustrated by many figures in the plates.
The ribs become coarser toward outer
one-third of the flank, but they are notvery weak on the inner part of the flank.
The number of ribs per whorl varies tosome extent. Sometimes there are striae,
lirae, bifurcation and insertion. Bifurcation arises at one-third of the flankwhere the ribs curve backward. In somespecimens ribs become weaker toward
the aperture and many fine riblets are
there instead of the major ribs. In one
specimen such a change occur at 90 mm.
diameter but in another specimen alreadyat 34 mm. diameter. In a small stage(e. g., diameter 10 mm.) the curve of ribs
is generally weak and more or less similar to that of Fuciniceras nakayamenses.
In a few specimens the sutures are incompletely preserved, showing a narrowand deep lateral lobe.
Measurements :No.
MM. 7058a
MM. 7059
MM. 7060
MM. 7065
GK. G. 2552
GK. G. 2557

D.
87.0
42.8
40. 1
66.5
46. 2
39.8

U.
38.9
15.1
14.8
27.0
20.0
13. 8

H.
26.2
16.7
15. 1
22. 4
22.0
16. 4

U/D HID

Rn~

0.45
0.35
0.37
0.41
0.43
0.35

25
22
19
33
39
32

0.31
0.39
0.38
O. 34
O. 48
0.41

Statistics :-Abou t seventy specimens
are available for counting the ribs per

611.

Jurassic Toyora Group-II

whorl at a given diameter. Here theassumption is made that the sample iscollected at random from one fossil population with the normal distribution of
the characters.
The results of the chi-square test are
as follows.
Table 1. The number of ribs per half awhorl at 20 mm diameter. Data classified
for the chi-square test. The classes between 27.5 and 31.5 are evaluated as one
class.
Class

Oi

Ei

13.5-15.5
15.5-17.5
17.5-19.5
19.5-21. 5
21. 5-23. 5
23.5-25.5
25.5-27.5
27.5-29.5
29.5-31. 5

2
6
16
13
12
7
8
1
1

2.44
6.06
10. 74
14.45
13.98
9. 72
5. 13
1. 93
0.53

Total

66

64. 98

N=66, x=21.29, s=3.53, V=16.58, X2=5.53,;.:2 0 • 05 ("=5)=11.07, P»0.05, Not significant.

Table 2. The number of ribs per half awhorl at 30 mm diameter. Data classified
for the chi-square test. The two classes
between 17.5 and 21.5 are evaluated as one
class and likely two classes between 29.5
and 33.5 are done.
Class

Oi

Ei

17.5-19.5
19.5-21. 5
21. 5-23. 5
23.5-25.5
25.5-27.5
27.5-29.5
29.5-31. 5
31. 5-33. 5

2
6
6
7
8
6
2
2

1. 76
4.06
6.87
8.39
7. 86
5.34
2.64
1. 00

Total

39

37.92

N=39, x=24.95, s=3.59, V=14.39, X2 =1.28,
X20.05("=3) = 7.81, P »0.05, Not significant.
The number of ribs per half a whorl at
50 mm. diameter.
N=ll, x=30.45, s=3.45, V=11.31.

3

By the test as mentioned above nosignificant difference is found from thenormal distribution. The frequency distribution of the number of ribs at eachdiameter seems to be unimodal.
Remarks :-Hitherto several palaeontologists and biostratigraphers have had
a question concerning the taxonomic position of the present species in the generic rank. For example MATSUMOTO (in
MATSUMOTO & ONO, 1947) dealt it as
Hildoceras (s. 1.) and SATO (1967, p. 391)
suggested that it might belong to Harpoceras. In this paper the present species
is referred to the genus Harpoceras.
Hildoceras HYATT, 1867, was established on the basis of the two species, H.
bifrons and H. walcotti. Later ZITTEL(1885, pp. 458-463) dealt Hildoceras as asubgenus of Harpoceras W AAGEN, 1869
under his Arieticeras-series and from thattime the diagnoses of Hildoceras andHarpoceras become wider. HAUG (1885,
pp. 632-646) spread the scope of Hildoceras without clear definition and his
genus Hildoceras included Lillia bayle,Harpoceras levisoni, Hildaites borealis,
Harpoceras serpentinum, etc. GEMMELLARO (1885) and FUCINI (1900, 1904, 1905,
1923-28) used Hildoceras without clear
definition in much wider meaning thanthat of HYATT. FUCINI's diagnoses ofHarpoceras, Grammoceras and Hildoceras
were confusing in his articles. BETTONI(1900, pp. 53-66) also used genus Hildoceras in a wide sense. The diagnosis
given by him was different from that of
HYATT. Hildoceras chrysanthemum and
Hildoceras densicostatum were established in such an age. In the present paper
I employ the diagnosiS given by ARK ELL(1957).
The present species has a unicarinatebisulcate venter and no longitudinalgroove on the flank, and therefore it isclearly excluded from Hildoceras.

4

Hiromichi HIRANO

The present species is closely allied
to Hildaites borealis (BUCKMAN) as YoKOY AMA (1904) mentioned.
Hildaites
belongs to the subfamily Hildoceratinae
as was treated in the "Treatise" by
ARKELL (1957). The present species,
however, does not belong to Hildoceras
as mentioned above. On the other hand,the characters of the present species
are not inconsistent with the diagnosisof the Harpoceratinae, and Harpoceras
is suitable for its generic position.
The present species has been hitherto
divided into Hildoceras (s. 1.) chrysanthemum and H (s. 1.) densicostatum. YOKOYAMA (1904) described that the latter was
distinguishable from the former by thegreater number of ribs. MATSUMOTO (in
MATSUMOTO & ONO, 1947) revised partly
the diagnoses of the two species andpointed out that there are some specimens which show intermediate characters.
I have examined more than 200 specimens including type specimens of thetwo nominal species and biometricallytreated about 70 specimens. On the basisof the results of statistics as well asother observations, all the specimens of
the sample are regarded as belonging to
one species. Considering the page precedancy in YOKOYAMA'S paper, I choose
the former as the specific name.
Comparison:- The present species occurs together with Harpoceras (s. str.)
sp. cf. H. exaratum, but is distinguishable by its less strongly curved ribs.
YOKOYAMA (1904) said that the present
species was closely allied to Hildaites
borealis (BUCKMAN). Indeed in the modeof the ribs Hildaites borealis is quite
similar to the present species, but thehafts of the ribs are shorter in theformer than in the latter. The venterof H. borealis is unicarinate-tabulate andhas somewhat subangular ventro-lateralshoulders. On the other hand the venterof the present species is unicarinatebisulcate and not acute but has gentle
ventrolateral shoulders.
Harpoceras mulgravium (YOUNG &BIRD) is somewhat similar to the present species, but the former has longer
hafts of the ribs and more acute curveof ribs on the middle of the flank.
Occurrence :- Loc. 3: 113, loco 30: 29,
loco 26: 12, loco 47: 10, loco 5: 8, loco 28' :8, loco 9; 5, loco 2: 4, loco 4: 3, loco 6:
3, loco 22: 3, loco 40: 3, loco 1: 2, loco 35 :2, loco 47B: 2, loco 18E: 1, loco 41: 1, unknown loco of the Sakuraguchi-dani: 6,
Tabe district: 1, unknown loc.: 22.

Harpoceras (Harpoceras) inouyei(YOKOYAMA)
PI. 1, Figs. 4-5; PI. 3, Fig. 5.

1904. Hildoceras inouyei YOKOYAMA; YOKO·
YAMA, Jour. Coli. Sci., Imp. Univ. Tokyo, vol. 19, art. 20, pp. 13-14, pI. 2,
figs. 5-6.
1947. Hildoceras (s.l.) inouyei YOKOYAMA;
MATSUMOTO in MATSUMOTO & 01'\0,
Sci. Rep. Fac. Sci., Kyushu Univ., vol.
2, no. 1, p. 28.
1956. Hildoceras inouyei YOKOYAMA; ARKELL,
Jurassic Geology of the World, Oliver& Boyd Ltd., p. 421.
1962. Hildoceras inouyei YOKOYAMA; SATO,
Mem. Soc. Ceol. France, no. 94, p. 59.
1962. Hildoceras inouyei YOKOYAMA; T AKAI,
MATSUMOTO and TORIYAMA ed., Geology of Japan, Univ. Tokyo Press, p. 83.

Types:-As YOKOYAMA (1904) established this species on several syntypes,
one of his illustrated specimens (1904,
pI. 2, fig. 6) is here designated as thelectotype, MM. 7063, and another illustrated specimen (1904, pI. 2, fig. 5) as theparalectotype, MM. 7062. They wereobtained at loco 47.
Material :-Seventy-two specimens,

611. Jurassic Toyora Group-II
Types, GK. G. 2641-2652, 2654-2666, 26682676, 2679-2681, 2683-2685,11299,11316-7,
11342-11349, GYU. M. 1028-1047.
Diagnosis :-The diagnosis of the present species, as noted by YOKOYAMA(1904) and supplemented by MATSUMOTO(in MATSUMOTO & ONO, 1947), is: A species of Harpoceras (s. str.) which has afairly narrow umbilicus (usually lessthan 1/3 of diameter in every size) andbifurcated, indistinctly fasciculated andlor occasionally inserted ribs.
Description:- The whorls enlarge rapidly, with a fairly narrow umbilicus. The
venter is probably unicarinate-bisulcate,although the specimens with a completely preserved venter are not available.
The umbilical periphery is rounded, but
the wall is steep and low. The ribs springon the umbilical wall and go on forward. At a point one-third or two-fifthsof the flank the ribs curve backwardwith increasing coarseness, drawing arcs,
and then recurve forward near the ventral shoulder. They usually are very
weak and often inperceptible on theumbilical periphery. The bifurcation
occurs at one"third or two-fifths of theflank where the ribs curve backward.
The fasciculation is indistinctly visibleon the innermost part of the flank. Inserted ribs occasionally occur on theouter two-thirds of the flank. The lirae
and striae are rare, but, if present, they
are visible along the ribs. In some specimens the sutures are partly preserved,
but the preservation is too poor for
tracing details. An example is shownin Pl. 3, Fig. 5.
Measurements :-U.
H. UjD HID
D.
No.
MM. 7062
26.1 8. 6 10.4 0.33 0.40
39.1 11. 0 16.8 0.28 0.43
MM. 7063
GK. G. 2642 41. 0 14.3 16. 2 0.35 0.40
GK. G. 2668 42. 9 15. 4 19.2 0.36 0.45
GK. G. 2672 52.3 12.2 25. 7 0.23 0.49

Rn~

28
29
37
36
28

5

Comparison:- The present species isclosely allied to Harpoceras chrysanthemum (YOKOYAMA) in general characters,
especially in the quite similar curvatureof ribs. The former has, however, distinctly involute shell and sometimes show
fasciculation and inserted characters of
the ribs.
The present species is also very similar
to Hildaites borealis (BUCKMAN) (WRIGHT,
1883, p. 438) in the mode of ribs. The
present species, however, is characterized by the involute shell and the unicarinate-bisulcate venter.
The mode of insertion and bifurcationof the ribs is somewhat similar to thatof Lioceratoides yokoyamai (MATSUMOTO), but the latter shows more eminentand regular bifurcation and insertion.
Occurrence :-Loc. 3: 25, loco 5: 12, loco
30: 11, loco 1 : 3, loco 26: 3, loco 47: 5, loco
2: 1, loco 4: 2, loco 18D: 1, loco 18E: 1,
loco 27: 1, loco 28: 1, loco 35: 1, loco 42:
1, unknown: 4.
Harpoceras (Harpoceras) sp. d. H. (H.)
mulgravium (YOUNG & BIIm)
PI. 1, Fig. 6.

Material.·-A fragmentary specimen,
GK. G. 11355.
Description:- The whorls enlarge rather rapidly, and the umbilicus seems tobe somewhat narrow. The venter seems
to be unicarinate-bisulcate, though it ispoorly preserved. Ribs are strongly falcoid and somewhat prorsiradiate. They
are straight and inclined forward on the
inner half of the flank. At the middleof the flank they abruptly curve backward and near the ventral shoulder theyrecurve forward, moderately projecting
on the venter. They are accompanied
by lirae on the preserved last stage. The
ribs are numerous and separated by in-Hiromichi HIRANO

Fig. 1. Map showing the studied area and the fossil localities.

terspaces narrower than the ribs. They
are strong on the outer half of the flankand somewhat weak on the inner half.
Measurements :.-H: 46.1
Comparison:- The present specimen
is similar to the British specimens of
Harpoceras mulgravium (YOUNG & BIRD)(WRIGHT, 1882, p. 433, BUCKMAN, 1909,
p. 4b) in the mode of volution, venterand ribs, but it is fragmentary and its

characters of the young stage is notobservable. Therefore I tentatively describe it under Harpoceras cf. mulgravium(YOUNG & BIRD).
H. mulgravium (YOUNG & BIRD) is allied to H. falcifer (SOWERBY) (PINNA,
1968, pp. 37-39), but is distinguished in
its wider umbilicus, less involute shelland straight hafts of the ribs_
Occurrence :-Loc. 28: 1.

611. Jurassic Toyora Group-II
Harpoceras (Harpoceras) sp. cf. H. (H.)
exaratum (YOUNG & BIRD)
PI. 2, Fig. 4; PI. 3, Fig. 1.

1904. Harpoceras sp. ; YOKOYAMA, Jour. ColI.
Sci., Imp. Univ. Tokyo, vol. 19, art. 20,
p. 15, pI. 4, fig. 2.
1947. Harpoceras (s. str.) sp. aff. H. exaratum(YOUNG & BIRD); MATSUMOTO in MA·
TSUMOTO & ONO, Sci. Rep. Fac. Sci.,
Kyushu Univ., Ceol., vol. 2, no. 1, p. 28.
1962. Harpoceras sp. aff. H. exaratum (YOUNG& BIRD) ; SATO, Mem. Soc. Ceol. France,
no. 94, p. 59.

Material:-Twelve specimens, GK. G.
2711, 2716, 11336-11341, GYU. M. 10251027, MM. 7071.
Description :-The whorls enlarge rapidly and the umbilicus is narrow. The
venter bears a less stout keel. Whether
the venter has sulci or not is not confirmable. The mode of ribs is falciform
and is characteristic of Harpoceras (s.
str.). The ribs are usually simple. The
angle of the curved ribs near the middleof the flank is about 120· to 150·. The
space of the ribs varies to some extent,
being wider than the interspace in most
specimens. Much finer ribs are occasionally visible along the major ribs, and ina large fragmentary specimen numerous
fine ribs are characteristic. One specimen has its aptychus which may be referred to Cornaptychus.
Measurements :No.
D.
U.
H. UjD H/D Rn!
GK. G. 2711 ca.45 10.9 21. 1 0.24 0.47 40

Comparison: -The present specimens
have been regarded as Harpoceras (s.
str.) sp. aft. H. exaratum (YOUNG & BIRD).
Indeed the present specimens are very
similar to the European specimens of
Harpoceras (s. str.) exaratum (YOUNG &BIRD) in the dense ribbing, the mode of7

falciform ribs and the involute shell.
According to BUCKMAN (1909, p. 5b) threeontogenetic stages are discerned in H.
(H.) exaratum, of which later two stages,"(2) the single ribs, falciform stage; (3)the finer rib stage, less falciform" areconfirmed in the Toyora specimens. As
the detailed nature of venter is not
visible, I treat the present specimens
as Harpoceras (H.) sp. ct. H. exaratum(YOUNG & BIRD).
The present species is similar to Harpoceras (H.) okadai (YOKOYAMA) but isdistinguished by its wider umbilicus andstronger curve and longer hafts of theribs.
Occurrence:- Loc. 28: 3, loco 2: 2, loco
47: 3, loco 1: 1, loco 22: 1, loco 36: 1, loco
36': 1.
Subgenus Harpoceratoides
BUCKMAN, 1909
Type-species :-Ammonites alternatus
SIMPSON in BUCKMAN, 1909.
Harpoceras (Harpoceratoides) nagatoensis

sp. nov. [MATSUMOTO, MS.]
PI. 3, Figs. 2-4.

1947. Crammoceras (PleydeUia) sp.; MATSUMOTO in MATSUMOTO & ONO, Sci. Rep.
Fac. Sci., Kyushu Univ., Ceol., vol. 2,
no. 1, p. 27, pI. 1, fig. 7.
1953. Harpoceratoides nagatoensis MATSUMOTO (nom. nud.); MATSUMOTO in MATSUMOTO et al. ed., "Historical Geology" (in Japanese), Asakura Book Ltd.,
p. 359.
1962. Harpoceratoides nagatoensis MATSUMOTO; SATO (nom. nud.) Mem. Soc. Ceol.
France, no. 94, p. 59.

Types:- The specimen illustrated· by
MATSUMOTO (in MATSUMOTO & ONO,
1947, pI. 1, fig. 7) is designated here ~s theHirO?nichi HIRANO

8

Sandy shale, thick ly bedded, nol plaly,
not so liss i!£', somewhat compact.
Silt-silty shale, thickly bedded, not platy,

calcareous nodule bearing.
Sandy shal~. th ic kly bed ded,sometimes massive.
Sandy shale. bedding plane IIat.
thickly platy, sometimes massive.

U'

Silt-silty shale, !issil e,

generally platy.
Clay-silty clay shale, thinly bedded,
lis sile like paper.
Alternation of shaleand sand ston e.
Sa.ndy shale-shal)' sandsione.

Uh

Fine sandston e.

11-'"'- loc. 2&-

loe. 27-

loc.26- IDe. 25 Nm- loc. 221, 3- lot.
loc.22-- loc. 21

Nm-- loc. 20- :g~.' It- loc.17

ir~::

tOC . 16

Coarse sandstone.

ogogogo

0000

_

1

Ub

Conglomerate.

Naloc, f.!.

-_-"
"4 1

_ ' , ,· 30__ 10c:. 3.L?____

Meium san dstone.

40

Nss

- 7-

Nm

_loc.~?_

Ncs

loc.32_loc. 3L -~-loc:. 29- 11·-·.-·Nsh-_._ .-

ll ~m.
Naae

Fig . 2. Columnar sections showing loca lities. The hori zo ntal bars are the boundariesof the formations (left side of t he column) and the boundaries of the zones (right side).
The loca li ty numbers correspond to the numbers in the systematic description.
a: Column along the sequence in the Sakuraguchi-dani; b: Colum n along the Anda-d ani;c: Column along the Kogo -dani; d : Column along the Chuzankei Ravine, Higashinakayama;
e: Column along the River Era.

Exp lanation of Plate 1(Natural size)
Figs. 1- 3. Harpoceras (s. str.) chrysanthemum (YOKOY AMA)
1. MM. 7058a, Lectotype, loco 47; 2. MM. 7059, Para lectotype, loco 47; 3. MM. 7060,
Para lectot ype, loco 47.
Figs. 4-5. Harpoceras (s . str.) inouyei (YOKOYAMA)
4. MM. 7062, Lectotype, loco 47; 5. MM. 7063, Para lectot ype, loco 47.
Fig. 6. Harpoceras (s. str.) sp. d. H. muigravium (YOU N G & BIRD)
GK. G. 11355, loco 28.
Fig. 7. Dumortieria ? sp.
Rubber cast of GK. G. 2797, loco 58.
Fig_ 8. Cram11!oceras (s . str.) sp. aff. C. obesum (B UC KM AN )GK_ G. 2781, loc o 55 .

HIRANO:

Jurassic Toyora Group-II

Plate 1

611.

Jurassic Toyora Group-II

holotype, GK. G. 2007, which was obtained at loco 9. Five specimens, GK. G.
2381-2384, 2391, are designated as paratypes.
Material:- The above mentioned six
specimens.
Etymology:-"Nagato" is the old nameof the southwestern part of Yamaguchi
Prefecture.
Diagnosis :-A species of Harpocerato ides which has fine, numerous andmoderately curved ribs and proportionally wide umbilicus.
Description:- The whorls enlarge moderately or somewhat rapidly. The volutions are moderately tight in most of thespecimens and in others somewhat involute. A keel is visible on the venterand may be unicarinate. The ribs are
so fine that they look like lirae. Theircurve is falciform and is characteristicof Harpoceras. At inner one-third of theflank, they curve backward moderately.
They are numerous, crowded and fasciculated on the inner lateral and theumbilical wall. They are wider thantheir interspace. The apertural part ofa large specimen (PI. 3, Fig. 3) is covered by numerous lirae instead of theribs.
Measurements :\"0.
GK. G. 2007
GK. G. 2383
GK. G. 2382

D.

U.

26. 0
32. 4
58.6

8. 9
14. 2
22.9

H.
10.3
10.4
21. 5

U/D
0.34
0.44
0.39

Comparison :-The present species issimilar to Protograml1loceras nipponicum(MATSUMOTO) in its curvature of ribs
but the latter does not show the fasciculation of the ribs.
The present species is similar to Harpoceratoides serotinum BETTONI (1900, pp.
65-66; HAAS, 1913, pp. 106-109; MONESTIER, 1931, pp. 31-32) but is distinguishedby its more evolute shell and rather

9

weaker curve of ribs.
Occurrence :- Loc. 9: 4, loco 22: 1, loco
30: 1.
Genus Pseudolioceras BUCKMAN, 1889
Type-species:- Ammonites

compactilis

SIMPSON in BUCKMAN, 1889.

Pseudolioceras sp.
PI. 3, Figs. 10-11.

1947. Hyperlioceras sp. ; MATSUMOTO in MATSUMOTO & ONO, Sci. Rep. Fac. Sci.,
Kyushu Univ., vol. 2, no. 1, p. 29, pI.
2, fig. 2.
1956. Pseudolioceras? sp.; ARKELL, Jurassic
Geology of the World, Oliver & BoydLtd., p. 421.
1962. Pseudolioceras sp.; SATO, Mem. Soc.
Geol. France, no. 94, p. 59.
1963. Pseudolioceras sp. ; T AKAI, MATSUMOTO
and TORIYAMA ed., Geology of Japan,
Univ. Tokyo Press, p. 83.

Material :-Ten unfavourably preserved specimens, GK. G. 2012, 27722776, 2782, 2792, 2793, 2798.
Description:-The whorls enlarge very
rapidly and the umbilicus is very narrow. The venter is unicarinate-tabulate.
The umbilical wall is low and more orless steep. The ribs are moderately
falciform. They spring on the umbilicalwall and go on somewhat forward. At
the middle of the flank they curve moderately backward and then recurve forward near the ventral shoulder. The
ribs are wide and rather low. The interspaces of the ribs are also wide. In
one specimen, GK. G. 2773, lirae are present on the ribs of the last whorl.
Aleasurements :U.
D.
1l.5
GK. G. 2010
64. 1
ca.9
GK. G. 2772
44.7
GK. G. 2773 ca. 34. 7 ca. 10. 7

No.

H.
U/D
33.9
O. 18
0.20
24. 4
0.31 ca. 13.5

10Hiromichi HIRANO

Comparison:- The present species issimilar to Pseudolioceras andromacoi
FUCINI (1923, pp. 61-62) in the tight involution and falciform ribs, but is distinguished by its less dense ribs. It issomewhat similar to Pseudolioceras sublythense (HAAS) (1913, pp. 113-114), but
is distinguished by its wider ribs.
As the available specimens are unfavourably preserved, precise specificidentification is hardly done.
Occurrence :-Loc. 55: 8, loco 53: 1, loco
54: 1.

Subfamily Hildoceratinae HYATT, 1867
Genus Hildoceras HYATT, 1867
Type-species:- Ammonites bifrons BRuGUIERE, 1789 (subsequently designated by
BUCKMAN, 1889).
Hildoceras sp. aff. H. bifrons (BRUGUIERE)
PI. 4, Figs. 4-5; PI. 2, Fig. 10
in Part III.

Material.·-Five specimens, GYU. M.
1068, GK. G. 11351-11345, of incomplete
preservation.
Description:-The whorls enlarge
slowly and the umbilicus is proportionally wide. The venter is tricarinatebisulcate. The umbilical periphery isrounded but the wall is nearly perpendicular and somewhat high. Therefore
the whorl-section is roughly rectangular.
The flank is flat. The curvature of arib is falcoid and characteristic of Hildo-

ceras. The ribs curve backward at the
middle of the flank. Usually they arevery weak or sometimes imperceptible
on the inner half of the flank but stout
on the outer half. They are widely interspaced. Lirae are present on the
inner half of the flank.
Measurements :-Height of the whorlof the last stage: GK. G. 11351=28.1,
11352=32.1,11353=24.8,11354=33.2, GYU.
M. 1068=51.7.
Comparison:- The present specimens
are similar to the British specimens of
Hildoceras bifrons (BRUGUIERE) (WRIGHT,
1882, p.436, pi. 59, figs. 1-2) in the modeof volution, venter and stout ribs, but
the curvature of ribs in the former is
not so abrupt as in the latter. In thepresent specimens the inner part of theflank is not grooved but flattened andthis may be a diagnostic character ofthe present specimens. Because thecharacter of the small stage is unknown
and only the fragmentary specimens areavailable, I tentatively treat the present
specimens under Hildoceras sp. aff. H.
bifrons, although they could possibly
represent a new species.
Occurrence :- Loc. 7: 1, loco 25: 1. loco
27: 1, loco 40: 1, loco 44: 1.

Subfamily Grammoceratinae
BUCKMAN, 1904
Genus Grammoceras HYATT, 1867
Type-species :-Ammonites striatulus
SOWERBY, 1823 (subsequently designatedExplanation of Plate 2(Natural size)
Figs. 1-3. Harpoceras (s. str.) chrysanthemum (YOKOYAMA)
1. MM. 7065, loco 47; 2. MM. 7061, Para lectotype, loco 47; 3. MM. 7064, Rubber cast,
Nishinagano
Fig. 4. Harpoceras (s. str.) sp. d. H. exaratum (YOUNG & BIRD)
MM. 7071, loco 47.

HIRA NO:

J~~TaissiG

Plate 2

ToyoTa Group-II

1

611.

Jurassic Toyora Group-II

by BUCKMAN, 1890).

11

Genus Dumortieria HAUG, 1885

Subgenus Grammoceras HYATT, 1867
Grammoceras (Grammoceras) sp. aff.

Type-species :-Ammonites levesquei
O'ORBIGNY, 1884 (subsequently designated by BUCKMAN, 1890).

G. (G.) obesum (BUCKMAN)
PI. 1, Fig. 8.

Dumortieria? sp.

1947. Cf. Crammoceras sp.; MATSUMOTO in
MATSUMOTO & ONO, Sci. Rep. Fac. Sci.,
Kyushu Univ., Ceol., vol. 2, no. 1, p. 27.

PI. 1, Fig. 7.

Material :-Five specimens, GK. G. 2781,
2783, 2787, 2784, 2795.
Description:- The whorls enlarge mod-erately and the venter has a small keel.
The umbilical periphery is rounded andthe wall is low. The ribs are moderately
falcoid. They spring on the umbilicalseam and go on forward with flexuosity.
After they proceed one-fourth or onethird of the flank, they curve backward
and recurve forward near the ventral
shoulder. They are simple, somewhat
elevated and separated by wider interspaces.
Measurements :No.
GK. G. 2781

D.
59. 4

U.
20. 4

U/D
O. 34

H.
22. 7

Rn!
ca. 21Comparison:- The present species resembles Grammoceras obesum (BUCKMAN)(1904, pp. 150-151) in the mode of ribs,
but has a tighter volution and somewhat
stronger curvature of ribs.
The present species is somewhat similar to Grammoceras thouarsense (O'ORBIGNY) in the curvature of ribs, but is distinguished by its tighter volution andsparser ribbing.
As the available specimens are notnumerous and unfavourably preserved,
I provisionally treat the present species
as Grammoceras sp. aff. G. obesum (BUCKMAN).
Occurrence :- Loc. 55: 5.

1947. Dumortieria (?) sp.; MATSUMOTO in
MATSUMOTO & ONO, Sci. Rep. Fac. Sci.,
Kyushu Univ., Ceol., vol. 2, no. 1, p. 29.
1956. ? Dumortieria sp.; ARKELL, Jurassic
Geology of the World, Oliver & BoydLtd., p. 421.
1962. Dumortieria? sp. ; SATO, Mem. Soc. Ceol.
France, no. 94, p. 59.

Material:-A single fragmentary specimen, GK. G. 2797.
Description :-The whorls enlarge
somewhat rapidly and consequently theumbilicus is rather narrow. The venteris invisible. The ribs, which spring at
the umbilical seam, are somewhat flexuous, slightly falcoidal, fine, simple andwidely interspaced.
Measurements :No.
D.
GK. G. 2797 ca. 18

U.
ca. 7

U/D

Rn!

ca. 0.39

17

Comparison:- The present species
somewhat resembles Dumortieria tabulata BUCKMAN (1905, p. 185) in the modeof ribs and volution, but it has finerribs. It is similar to Dumortieria levesquei (O'ORBIGNY) (HAUG, 1885, pp. 662663) in the somewhat fine and widelyinterspaced ribs, but has more flexuous
ribs. On the other hand it is apparently
similar to Tmetoceras in its simple andfine ribs. Because only a single specimen with poorly preserved venter isavailable, a precise identification cannot
be done.
Occurrence :- Loc. 58: 1.

Hiromichi HIRANO

12

Family Hammatoceratidae
BUCKMAN, 1887
Subfamily Phymatoceratinae
HYATT, 1867
Genus Phymatoceras HYATT, 1867
Type-species:- Phymatoceras robustum
Hy A TT, 1867.

Phymatoceras toyoranum (MATSUMOTO)
PI. 4, Fig. 3.

1947. Lillia toyorana MATSUMOTO; MATSUMOTO in MATSUMOTO & ONO, Sci. Rep.
Fac. Sci., Kyushu Univ., Ceoi., vol. 2,
no. 1, p. 29, pI. 2, fig. 1.
1956. Phymatoceras toyoranum (MATSUMOTO);
ARKELL, Jurassic Geology of the World,
Oliver & Boyd Ltd., p. 421.
1962. Phymatoceras toyoranum (MATSUMOTO);
SATO, Mem. Soc. Ceoi. France, no. 94,
p. 59.
1962. Phymatoceras toyoranum (MATSUMOTO);TAKAI, MATSUMOTO and TORIYAMA ed.,Geology of Japan, Univ. Tokyo Press,
p. 83.

Types :-Here I designate one of thetwelve syntypes, GK. G. 2763, as thelectotype and others, GK. G. 2011, 2761,

2764, 2765, 2766, 2768, 2768, 2770, 2771,
2777, as paralectotypes, of which one
specimen, GK. G. 20n was previously
illustrated by MATSUMOTO (1947, pI. 2,
fig. 1).
Material :-The above mentioned
twelve specimens.
Diagnosis :-A species of Phymatoceras
which has slightly to moderately sigmoidal, widely interspaced ribs and notubercles.
Description:-The whorls enlarge
somewhat rapidly and consequently theumbilicus is narrow in the young stageof about 20 mm. in diameter. In thelater stages the whorls enlarge somewhat slowly and the umbilicus is proportionally moderate. The venter isunicarinate-bisulcate. The ribs springat the umbilical seam and go on forward.
At a point inner one-third of the flankthey curve backward and near the ventral shoulder they recurve forward.
They are, thus, slightly sigmoidal andthey become stouter toward the ventral
shoulder. They are interspaced widelyand often bifurcate at the umbilicalperiphery. In some specimens the ribs
are slightly elevated at the umbilicalperiphery where the bifurcation arises,
but the elevation is not so strong as to

Explanation of Plate 3(Natural size)
Fig. 1. Harpoceras (s. str.) sp. d. H. exaratum (YOUNG & BIRD)
Rubber cast of GK. G. 2711, loco 2.
Figs. 2-4. Harpoceras (Harpoceratoides) nagatoensis sp. nov.
2. GK. G. 2383, Para type, loco 9; 3. GK. G. 2381, paratype, loco 30; 4. GK. G. 2007, Holotype,loc. 9.
Fig. 5. Harpoceras (s. str.) inouyei (YOKOYAMA)
GK. G. 2642, loc. 3.
Figs. 6-9. Harpoceras (s. str.) chrysanthemum (YOKOYAMA)
6. GK. G. 2572, loc. 3; 7. GK. G. 2552, loc. 3; 8. GK. G. 2557, lac. 3; 9. GK. G. 250-1,
loco 3.
Figs. 10-11. Pseudoiioceras sp.
10. GK. G. 2773, loc. 55; 11. GK. G. 2012, loc. 55.

HIRANO:

Jurassic Toyora Group-II

Plate 3

611.

Jurassic Toyora Group-II

be called tubercles. One specimen, GK.
G. 2763, with a diameter of 64 mm., hassome lirae on the apertural part.
Measurements :No.
GK. G. 2763

D.
ca. 61

U.
21.6

UjD
ca. 0.35

Rn!
20

Comparison:- The present species issimilar to Phymatoceras pulcher MERLA(1933, pp. 30-31) in the mode of volution
and the curvature of ribs, but is distinguished by its stouter ribs and indistinctness of tubercles at the umbilicalperiphery.
The present species is similar to Phymatoceras malagma (DUMORTIER) (BUCKMAN, 1899, p. 21) in the curvature of ribs,
the bifurcation and the volution. The
unmistakable distinction between thetwo species is the considerably dominant
tubercles in the latter.
The present species resembles Phymatoceras obtecta (BUCKMAN) (1899, p. 21)
in the mode of ribs. The latter is distinguished in its more dominant andnumerous tubercles in younger stages
and more rapidly enlarging whorls.
The present species is also similar to
Phymatoceras narbonensis (BUCKMAN)(MONESTIER, 1931, p. 21) in the mode ofbifurcation. They are distinguished inthat the ribs are more flexuous and considerably elevated at the umbilical periphery and the whorls enlarge more
rapidly in the latter than in the former.
Occurrence:- Loc. 55: 13.
Phymatoceras sp.
PI. 4, Figs. 1-2

1947. Haugia aff. japonica (NEUMAYR); MATSUMOTO in MATSUMOTO & ONO, Sci.
Rep. Fac. Sci., Kyushu Univ., Ceol., vol.
2, no. 1, p. 29, pI. 2, fig. 5.
1956. Haugia aff. japonica (NEUMAYR); ARKELL, Jurassic Geology of the World,

13

Oliver & Boyd Ltd., p. 421.

1962. Haugia aff. japonica (NEUMAYR) ; SATO,
Mem. Soc. Ceol. France, no. 94, p. 59.

Material:-Two specimens, GK. G. 2801,
11350.
Description:- The whorls enlarge!'Ilowly and consequently the umbilicus
IS proportionally wide. The venter has
a low keel. The presence of sulci is notconfirmed. The umbilical periphery isrounded and the wall is steep. Somewhat prominent tubercles are present
between the umbilical wall and the umbilical shoulder, and two ribs springfrom every tubercle in the growth stages
less than 50 mm. in diameter. The ribs
go on backward on the flank and near
the ventral shoulder they curve forward.
In the late growth stage with diameters between ca. 50 mm. and ca. 100 mm.,the tubercles are weak or very weak,
and the twin becomes obscure. The ribs
are sigmoid, very weak on the inner
one-third of the flank and somewhat
strong on the ventral shoulder. The
curvature of the rib in this stage is thesame as that in the younger stages. In
one specimen, GK. G. 2801, lirae occur
in the part with diameters between ca.
70 mm. and ca. 85 mm. and they areprominent on the umbilical shoulder.
The tubercles and ribs become very
weak and finally fade away in the laststage over 100 mm. in diameter.
Measurements :No.
D.
D.
H.
DID Rn!
GK. G. 2801 ca.83 38. 9 ca. 23. 7 ca. O. 47 ca. 27
GK. G. 11350
54. 8
30. 9
25

Remarks:-The present species has
been considered as being allied to
Haugia japonica (NEUMAYR) from Shikoku for a long time. The taxonomic
position of Haugia japonica has so far
been discussed by many students (NEUMAYR in NAUMANN & NEUMAYR, 1890,

14

Hiromichi HIRANO

p. 35; YEHARA, 1927, pp. 37-39; SHIMIZU,
1928, pp. 219-222; KOBA YASH!, 1935, pp.
79-83) and recently ZEISS (1956, p. 76)
pointed out that it is closely allied to

Hecticoceras(Putealiceras)

mathayense(KILIAN). HAY AMI & MATSUMOTO (MATSUMOTO et al., 1963, p. 35) regarded it
as Hecticoceras? (Putealiceras) japonicum.
H.? (P.?) japonicum has less sigmoidal
ribs and tubercles at a point about onethird from the umbilical margin to theventer, but the present species has more
sigmoid ribs and tubercles on the umbilical shoulder. There are unmistakable differences between them.
Although the present species is represented only by two specimens, it is notreferred to the genus Hecticoceras by
the following ground. Ribs are strong
in Hecticoceras but moderate Or weak in
the present species. In Hecticoceras theribs end at the ventrolateral tubercles,
but such a character is not shown by
the present species. Hecticoceras has
tubercles through the growth stage, but
in the present species the tubercles are
weakened in a late stage with diameters
between 50 to 100 mm. and finally fadeaway.

The present species is not referred to
the genus Haugia, because it has a low
keel and sigmoid ribs.
Considering all the observable characters it is better to ascribe the present
species to Phymatoceras.
Comparison:- The present species issimilar to Phymatoceras pulcher MERLA(1933, pp. 30-31) in the mode of volution
and ribs, but is distinguished by its lesspersistent tubercles.
The present species is similar to Phymatoceras robustum HYATT (ARKELL etal., 1956, p. L265) in the mode of tubercles and volution. In the former, twinned ribs arises from every tubercle inthe younger stages. On the other hand,in the latter the ribs are simple, twinned or triploid from tubercles and themode is irregular.
The present species is similar to Phymatoceras toyoranum in the mode of volution and ribbing, but is distinguishedby its dominant tubercles at the umbilical shoulder in the younger stages andsomewhat weaker ribs.
Occurrence :- Loc. 56: 2.
(to be continued)

Explanation of Plate 4(Natural size unless otherwise indicated)
Figs. 1-2. Phymatoceras sp.
1. Gypsum cast of GK. G. 11350, lac. 56, x 0.8; 2. Gypsum cast of GK. G. 2801, lac. 56,
xO.8.
Fig. 3. Phymatoceras toyoranum (MATSUMOTO)
Rubber cast of GK. G. 2763, Lectotype, lac. 55.
Figs. 4-5. Hildoceras (s. str.) sp. aff. H. bi/rons (BRUGUIERE)
4. GK. G. 11351, lac. 40; 5. GK. G. 11354, lac. 27.

HIRANO.'5

J~wassic

ToyoTa G'l'"oup-III

Plate 4-

Trans. Proc. Palaeont. Soc. Japan, N.S., No. 89, pp. 15-23, pIs. 5, 6, April 20, 1973

612.

CORAL FOSSILS FROM THE KOSODE FORMATION,YAMAN ASHI PREFECTURE*
KATSUHIKO NISHIMIY A
Institute of Geosciences, Yamanashi University

andNOBUO Y AMAGIW A
Institute of Geosciences, Osaka Kyoiku University

IlI~WL IHm~I\'lO)-if' /' ::I'1I:E:

llI~g\'l;O)~t*i1ffiH;::n-1nT Q/Hdl~~ip!?:>ilIl±', L.t.: -if' /'::I'1I:Em~tUjL., 10fm~~)jIJL.1lc.iI!1(L.t.:o<c.:(-0)$ 2{11HHfrflli'"C<!;Qo ;:::h,!?:>-if'/'::I·1I:Ef,'F

O)t1lf1lX:~*0)~ fJ:, fti:JII~tl!!.~ $,C"
L.t: J~H~~llf ilJ:: ~ .~,*lWillHS~~;mO)~ 0)1;:: FnJ~t.:fJ:ili:fCJ.O)f!I!'"C<!; ~, IHm~}~HJ:J:$:1 ='- 7 i1d;::~T Q;:: f;::~Ir'fJ:tJ:Ir'o IHm~~~fJ: t:c

1rO c T Q;:: 0)j:l!!.~0):/ ='- 7*0)~pt;ilJ:: ~J:l!!.'I!i:t1lfiiil;::-::>"'Lff'1~L.t.:oi1§1.n\i:~· llIr.r,ul!;*:The coral fauna may be regarded asvery valuable in order to discuss thegeological age, correlation and palaeoenvironment of the Kosode Formation,so far as other leading fossils are indeed
lacking. Ten coral species are discriminated from the formation, and described in the present paper. In addition, the Jurassic Torinosu Series isbriefly mentioned, making special refrence to the Kosode Formation.
On publishing the present paper, thewriters wish to express their sincere
thanks to Professor Keiji NAKAZAWA
of the Kyoto University for his continuous suggestions. Thanks are due to
Mr. Toyohiko Iw AHASHI of the Osaka
Kyoiku University for his photographic
work, and also due to Mr. and Mrs.
Yasuo NOGAMI of the Kyoto University
for their helps in preparation for the
manuscript.

Introduction and acknowledgements
The Kosode Formation is a characteristic member composing the zonal structure in the Kanto-Mountainland, and isextending from Kosode toward Aoiwadani in the surveyed area, the northeastern part of Yamanashi Prefecture(Text-fig. 1). The stratigraphical andstructural studies of the formation havebeen carried out by FUJIMOTO (1939),
FUJIMOTO and SUZUKI (1969), Yamanashi
Prefecture Government (1970), and NISHIMIY A and Y AMAGIW A (1971). There has
been, however, published no palaeontological report, although corals are rather
dominant in limestone lenses at Kosodeand Aoiwadani, and occur sometimes inassociation with stromatoporoids.
* Received Oct. 20, 1972; read June 27,
1971, at Nara.

15

Katsuhiko NISHIMIYA and Nobuo Y AMAGIWA

16

N

IIiiI

\

ItXfU)\1 II
~III

~IV
~v

1._1

VI

_--

VII
VIII

o

1000

2000m

L '_ _- - ' - ,_ _- - - ' ,Kobotoke Group

Fig. 1. Geological map of the northeastern part of Yamanashi Prefecture.
A: Kumotoriyama Formation, B: Kosode Formation, C: Aoiwadani Formation, D: Kamosawa Formation, E: Oonari Formation, I: shale or slate, II: sandstone, III: conglomerate,
IV: schalstein, V: chert, VI: limestone, VII: fault, VIII: strike and dip, 1-21; Sample loco

Stratigraphy
The Jurassic strata were studied by
FUJIMOTO (1939) in the Kogochi District

and by FUJIMOTO and SUZUKI (1969) in
the Oborogawa District. They are subdivided as follows:

Kogochi District
Kogochi Group (Jurassic)

Oborogawa District
Otaki Group (Jurassic)

Takanosuyama Formation

Tochimoto Formation
Kawamata Formation
Sogoyadani Formation
Kosode Formation
Koreisan Formation

Kosode Formation
Oonari Formation
The stratigraphy of the surveyed
Aoiwadani-Kosode area is summarized,
as shown in Table 1. The Jurassic

Torinosu Series is separately distributed in three belts, namely, the Oonari
Formation in the southern belt, the612.
Table 1.

17

Corals from Kosode Formation

Stratigraphical subdivision in the surveyed area.
Aoiwadani-Kosode Area-~----- --------~,----------------------I

Cretaceous

Shimanto Series-------1--------------

Jurassic

Torinosu Series

Kobotoke Group- - - - - - - - - (fault) - - - - , - - - - - - - 1
Kosode F.

I Kamosawa F.

Oonari F.

1 - (fault) _ - '_ _ _ _ _ _~Aoiwadani Formation

Triassic?

(fault) --~-------I
PermianChichibu Series

Kumotoriyama Formation

Kamosawa Formation in the middle
belt, and the Kosode Formation in thenorthern belt. The boundaries of theseformations are bounded by fault.
The Oonari Formation: Typicallydeveloped at Oonari, well cropping out
along the middle stream of the Tabagawa River; mainly composed of sandstone, shale and slate, subordinately
made up of schalstein and chert, intercalating blackish grey limestone lenses;
about 2000 m thick.
The formation showing a generaltrend of NW -SE, dipping to northeastand southwest at angles 50-80°, generally characterized by homoclinal structure. Several faults trending to NW-SE
and NE-SW. The boundary between
the present formation and the Kamosaw a Formation in the middle belt ismarked by fault, which is cut off by
NE-SW faults.
The Kamosawa Formation: Typically
cropping at Kamosawa; mainly composed of shale and slate, intercalating
sandstone, thin limestone and chert;
about 1200 m thick.
The formation showing a strike of
N45-800W, dipping to northeast or southwest at angles 40-80°. Several faults
trending to NW-SE and NE-SW. The
boundary between the present for-mation and the presumably Triassic
Aoiwadani Formation is marked by afault, which is cut off by NE-SW faults.
The Kosode Formation: Typically
exposed along the upper stream of the
Kosodegawa River and at Aoiwadani;composed of slate (often changed into
phyllite) and sandstone, occasionallyintercalating blackish grey limestonelenses, which yield hexacorals andstromatoporoids; about 500 m thick.
The formation showing a generaltrend of NW -SE, dipping to northeast
at angles 40-70". A considerable number of faults trending to NW-SE and
NE-SW. The formation is bounded on
the north by the Permian Kumotoriyama Formation and on the south by
the Aoiwadani Formation. These formations are in contact with each otherby faults, which are cut off by NE-SW
faults.
The limestone lenses intercalated in
the Kosode Formation yield corals, as
shown in Table 2.
The fauna of corals listed in Table 2
are equivalent or similar to those fromthe Upper Jurassic in Japan, and the
Kosode Formation can be safely correlated with the Torinosu Series in the
Sagawa basin, Southwest Japan. The
Kamosawa Formation and the Oonari

Katsuhiko NISHIMIY A and Nobuo Y AMAGIWA

18

Table 2.

~
Species

List of the Jurassic corals from the Kosode Fcrrmation.
Kosodegawa

Aoiwadani

~

N

U
0
..4

~

In

"'''0

~

......

0>......

......

0
N

N

(fl-

o::l
u
u
u
u
u
U
U
.- 0
0
0
0
0
0
0
0
'<::.0[/) '-J
..4
..4
..4
..4
..4
..4
..4
- - - - - - - - - -- - - -- -x

Actinastrea? sp. indet.

Caiamophyllia sp. indet.

M"'::=lii,.....

- - - -- - - - - - - - - -- -

x

x

x

- -- - - - - - - - - - - - - x

Dimorphoseries tabayamensis- -- - - -- - - -- - - - - x

Enallhelia sp. indet.
I sastrea? sp. indet.

x- - - - - - - -- - - - - -x- -- -- -- -- -- -- -- x

Stylina kantoensis- -- -- -- -- ---Stylina? sp. indet.

x- - - - - - - -- - - - - - - Thamnasteria huzimotoixx- - - -- - - - - - - - - -- -T. aff. huzimotoi
T. sp. Axx

T. sp. B- - - - - -- -- - - - - - - - -- -- -- -- -- - - -- -x

Trocharea sp. indet.

- - - - - - - - - -- - - - - -xHexacoral gen. et sp. indet.

- - - - - - - - - -- - - -- -Formation, from which any valuablefossil has never been obtained, aretentatively regarded as the Jurassic,
judging from their lithic resemblance
to the well ascertained Torinosu Series
in the surveyed area. The details of
the correlation will be discussed by thewriters in the near future.

Systematic description

Order Scleractinia BOURNE, 1900
Suborder Astrocoeniina VAUGHAN& WELLS, 1943
Family Thamnasteriidae VAUGHAN& WELLS, 1943

x

x

Genus Thamnasteria LESAUVAGE, 1823
Thamnasteria huzimotoi EGUCHI
Plate 5, figs. 2a-c
1951.

Thamnasteria huzimotoi EGUCHI, Sci.
Rep. Tohoku Univ., 2nd ser. (Geol.),
vol. 24, p. 79, pI. 26, fig. 10.

Corallum massive and thamnasterioid.
Corallites small and confluent, without
distinct wall; central distance 2.0-4.0
mm. Septa slightly sinuous, confluent
with those of the adjacent corallites,
30-45 in number, 10-13 of them extending to center; laterally by synapticulae.
Perforations generally scarce.
Columella styliform, but in some corallites
rudimentary.

612.

Corals from Kosode Formation

Comparison: The present form isclosely allied to the type specimen ofThamnasteria huzimotoi EGUCHI (1951, p.
79, pI. 26, fig. 10) from the Torinosu
group in Itsukaichi, Kanto-Mountainlandin many common characters. However*,the former's perforations are more
scarce than the latter's. It is similar
to T. torinosuensis EGUCHI (1951, p. 67,
pI. 24, fig. 9; MORI, 1963, p. 58, pI. 22,
fig. 3) from the Upper Jurassic in the
Sagawa basin and the Soma area, but
the former differs from the latter inhaving more numerous septa. It resembles T. crateriformis GREGORY, (1900,
pp. 135-136, pI. 17, figs. 4, 5, 7) from theUpper Cutchum formation in Cutch inmany points. However, the former canbe distinguished from the latter by its
smaller coralli tes and less numerous
septa.
Occurrence: Loc. 2 at Aoiwadani and
Loc. 20 at Kosodegawa, Tabayamamura, Kitatsuru-gun, Yamanashi Prefecture.
Geological horizon: Upper Jurassic.

Thamnasteria sp. A indet.
Plate 5, figs. 3a, b

Corallum massive and thamnasterioid.
Corallites somewhat irregular in size,
withou t distinct wall. Corallite centers
3.5-6.0 mm. Septa gently flexous, somewhat perforate, confluent between corallites. Corallites 30-45 septa; 11-15 ofthem (first and second cycles) extending
to center and some of them joint with acolumella. Synapticulae present. Columella styliform, but in some corallites
rudimentary. No longitudinal section.

* According to the writers' observations,the septa of the type specimen are about 40
in number.

Compariso'n: The present fonn isvery similar to Thamnaster.ia huzimotoi
EGUCI-II (1951, p.79, pI. 26, fig. 10; NISHIMIYA & Y AMAGIWA, the present paper)from the Upper Jurassic in the KantoMountainland, but differs from thelatter in having less numerous synapticulae and longer central distance.
It resembles T. torinosuensis EGUCHI(1951, p. 67, pI. 24, fig. 9; MORI, 1963, p.
58, pI. 22, fig. 3). However, the former
has more numerous septa.
Occurrence: Loc. 2, Aoiwadani, Tabayama-mura, Kitatsuru-gun, Yamanashi
Prefecture.
Geological horizon: Upper Jurassic.

Thamnasteria sp. B indet.
Plate 5, figs. la, b

Corallum thamnasterioid. Corallites
relatively large and united by septa,
about 8.0 mm between calicural centers.
Septa slightly sinuous and less numerous (20+?); perforate, about 12 ofthem extending to center. Columella
distinct and styliform. Synapticulae
present. No longitudinal section.
Comparison: The present species
somewhat resembles Thamnasteria abukumaensis MORI (1963, p. 58, pI. 21, figs.
6, 7) from the Upper Jurassic in the
Soma area. However, the former isseparable from the latter in having
perforate septa and a styliform columella.
Occurrence: Loc. 5, Aoiwadani, Tabayama-mura, Kitatsuru-gun, Yamanashi
Prefecture.
Geological horizon: Upper Jurassic.

Family Astrocoeniidae KOBY, 1890
Subfamily Astrocoeniinae KOBY, 11890

20

Katsuhiko NISHIMIYA and Nobuo YAMAGIWA

Genus Actinastrea D'ORBIGNY 1849

Genus Stylina LAMARCK, 1816

Actinastrea ? sp. indet.

Stylina kantoensis, sp. nov.

Plate 6, fig. 6

Plate 6, fig. 1

Corallum massive.
Corallites subpolygonal in transverse section, united
by wall; generally 1.2-1.5 mm in diameter. Septa about 24, straight; 6 offirst cycle long and reach to columella,
6 of second cycle slightly shorter than
the first and reach to near the columella.
Septa of third cycle slightly developed.
No longitudinal section.
Comparison: The present form isrepresented by a single transverse section. It is similar to the species belong·
ing to the genus Stylina, but differsfrom the latter in having subpolygonalcorallites. It resembles Astrocoenia sp.
(MORI, 1963, p. 56, pI. 21, fig. 8) fromthe Upper Jurassic in the Soma area,
Northeast Japan. However, the calicuralwall of the former is subpolygonal intransverse section and the corallites ofthe former are not well preserved.
Occurrence: Loc. 3, Aoiwadani, Taba·yama-mura, Kitatsuru-gun, Yamanashi
Prefecture.
Geological horizon: Upper Jurassic.

Corallum massive. Corallites circularin transverse section, subequal in size;
generally 1.3-1.5 mm in diameter. Corallites united by coastae. Corallite centeres 2.2-3.0 mm. Septa thick, 12 in
number, arranged in two hexameral cycles; 6 of first cycles extending to nearcolumella. Columella styliform, circularin transverse section; 0.2-0.3 mm in diameter. Wall thick. Intercorallite area
well developed, formed of costae. Costae sub equal, 24 in number. No longitudinal section.
Comparison: The present form resembles Stylina sugiyamai EGUCHI (1951,
p. 60, pI. 22, fig. 7) from the Upper
Jurassic Torinosu group in the Sagawa
basin, Southwest Japan, but differs fromthe latter, having longer central distance. It can be distinguished from S.
higoensis EGUCHI (1951, p.74, pI. 12, figs.
6-9) from the Sakamoto formation in
Kuma Mountainland, southwest Japan inhaving broader corallites and longer
central distance.
Occurrence: Loc. 20, Kosodegawa,Tabayama-mura, Kitatsuru-gun, Yamanashi Prefecture.
Geological horizon: Upper Jurassic.

Family Stylinidae D'ORBIGNY, 1851
Subfamily Stylininae D'ORBIGNY, 1851

Explanation of Plate 5
All figs. x3
Fig. 1. Thamnasteria sp. B indet.
a) Transverse section, reg. no. 72092401a; b) Longitudinal section, reg. no. n092401b.
Fig. 2. Thamnasteria huzimotoi EGUCHIa) Transverse sectioh, reg. no. 72092402a; b) Longitudinal section, reg. no. 72092402b;c) Transverse (somewhat oblique) section, reg. no. 72092403.
Fig. 3. Thamnasteria sp. A indet.
a) Transverse section, reg. no. 72092404; b) Transverse section, reg. no. 72092404.

NISHI MIY A and YAMAGI WA:

Corals !1"Om Kosode Fo'rmationlb

Plate 5

612.

Corals from Kosode Formation

Subfamily Euheliinae
DE FROMENTEL, 1861
Genus Enallhelia EDWARDS& HAl ME, 1850
Enallhelia sp. indet.
Plate 6, figs. 7a, b

The present species almost agrees
with the genus Enallhelia in many
important characters. It is closely
similar to E. nipponica somaensis EGUCHI (1942, pp. 143-144, pi. 6, figs. 8, 9;
MORI, 1963, pp. 56-57, pi. 23, fig. 3) fromthe Upper Jurassic in the Soma area inmany common characters. However, it
is separable from the latter in having
a compressed columella. It is related toE. nipponica EGUCHI (1942, pp. 142-143,
pi. 6, figs. 4-7) from the Lower Cretaceous in Miyagi Prefecture, but theformer's calices are smaller than thelatter's.
Occurrence: Loc. 20 at Kosodegawa
and a boulder at Shiosawa, Tabayamamura, Kitatsuru-gun, Yamanashi Prefecture.
Geological horizon: Upper Jurassic.

21

numerous, 19 of them extending tocenter. Secondary (marginal) calice only
one. In this calice, about 12 of septa
extending to center. Septa slightly
sinuous and somewhat perforate. Distance of primary and secondary centersabout 6.5 mm. Wall relatively thin. Columella trabecular in central calice, but
rudimentary in marginal one. Synapticulae numerous and distinct.
Comparison: The present species resembles Dimorphoseris oolitic a DUNCAN(1872, pt. 3, pp. 22-23, pi. 4, figs. 1-4) fromthe Oolites series in Europe in many
common characters. However, it differsfrom the latter in having smaller corallum, trabecular columella and less numerous septa extending to columella.
Occurrence: Loc. 5, Aoiwadani, Tabayama-mura, Kitatsuru-gun, Yamanashi
Prefecture.
Geological horizon: Upper Jurassic.

Superfamily Agariciicae GRAY, 1847
Family Calamophyllidae VAUGHAN& WELLS, 1943
Genus Calamophyllia BLAINVILLE, 1830
Calamorphyllia sp. indet.

Suborder Fungiina VERRILL, 1865

Plate 6, fig. 3

Superfamily Fungiicae DANA, 1846

The present form is not well preserved. It may belong to the genus
Calamophyllia" than the genus Stylosmilia
in having no styliform columella. It
resembles Rhabdophyllia Qr Stylosmilia(EGUCHI, 1951, pi. 23, fig. 5) from theUpper Jurassic Torinosu group at Ogawa, Ogawa-mura, Takaoka-gun, Kochi
Prefecture in many respects. However,the former has somewhat larger corallites than the latter.

Family Synastreidae ALLOITEAU, 1952
Genus Dimorphoseris DUNCAN, 1872
Dimorphoseris tabayamensis, sp. nov.
Plate 6, figs. 2a, bCoralum thamnasterioid and turbinate; about 18 mm in diameter. Colony
formation by circumoral budding. Thereis a large central primary calice; septa

Katsuhiko NISHIMIYA and"Nobuo YAMAGIWA
Occurrence: ~oc. 2 and 4 at Aoiwadani ?-nd Loc. 20" ~t Kosodegawa, Tabayama-mura, Kitatsuru-gun, Yamanashi
Prefecture.
Geological horizon: Upper Jurassic.

Superfamily Poriticae GRAY, 1842
Family Microsolenidae KOBY, 1890
Genus Trocharea ETALLON, 1864
Trocharea sp. indet.

Genus Isastrea EDWARDS& HArME, 1851
I sastrea ? sp. indet.
Plate 6,fig. 4

The present specimens may belong to
the genus Isastrea in having the following characters. 1) cerioid colonies, 2)corallite wall present, but with septa
confluent partly absent, 3) monocentriccorallites, but rarely dicentric, 4) trabecular columella, but feeble. It is similar to some species of the genus Latomtwndra. "The corallites of the latter,
however, are mostly di- or tricentric.
Occurrence: Loc. 2, Aoiwadani, Tabayama-muni, Kitatsuru-gun, Yamanashi
Prefecture.
Geological horizon: Upper Jurassic.

Plate 6, fig. 5

Corallum simple. Calicural diameter
about 15mm in transverse section. Thinwall developed, although it is not recognizable in part. Septa numerous, 7080 in number. Septa slightly sinuousin part and geniculate in part, with perforations. Columella trabecular. Synapticulae present but a few. No longitudinal section.
Comparison: The present form isclosely related to Trocharea sp. (EGUCHI,
1951, pI. 25, fig. 1) from the Torinosu
group at Ogawa, Ogawa-mura, Sagawa
basin, Southwest Japan in having
many common" characters. However,the former one has less numerous septa
than the .latter.
Occurrence: Loc. 2, Aoiwadani, Tabayama -mura, Kitatsuru -gun, Yamanashi

Explanation of Plate 6
Fig. 1. Stylina kantoensis, sp. nov.
Transverse section of the holotype, x 3, reg. no. 72092405.
Fig. 2. Dimorphoseris tabayamensis, sp. nov.
a) Transverse section of the holotype, x 3, reg. no. 72092406a; b) Longitudinal sectionof the holotype, x 3, reg. no. 72092406b.
Fig. 3~ Calamophyllia sp. indet.
Transverse section, x 3, reg. no. 72092407.
Fig". 4. Isastriw? sp. indet.
Transverse section, x 3, reg. no. 72092408.
Fig~ 5. Trocharea sp. indet.
Transverse section, x 3, reg. no. 72092409.
Fig. 6. Actinastrea? sp. indet."
Transverse section, x 5, reg. no. 72092410.
Fig. 7. Enallhelia sp. indet.
a) Transverse secfion, x 5, reg. no. 72092411; b) Transverse section, x 5, reg. no. 72092412.

NISHIMIY A and YAMAGIWA:

Corals from Kosode Formation

Plate 6

612.

Corals from Kosode Formation

Prefecture.
Geological horizon:

Upper Jurassic.

All the figured specimens are collectedby Katsuhiko NISHIMIY A, and are deposited at the Institute of Geoscience,
Osaka Kyoiku University; reg. nos.
72092401 to 72092412.
References
DUNCAN, P. M. (1872): A monograph ofBritish fossil corals. Palaeontogr. Soc.,
London, Monogr., 2nd ser., pt. 3, pp. 1-24,
pIs. 1-7.
EGUCHI, M. (1924): Recent and fossil coralsof the family Oculinidae from Japan.
Ceol. Soc. japan, jour., vol. 49, no. 583,
pp. 135-l42, pI. 6 (5).
(1951): Mesozoic Hexacorals from Japan. Sci. Rep. Tohoku Univ., 2nd ser.
(Ceol.), vol. 24, pp. 1-96, pIs. 1-28.
FUJIMOTO, H. (1939): On the Torinosu Se-

Aoiwadani
Kitatsuru-gun
KosodeUshiroyamagawa
Oonari-;5- :tl~I:; 'li[1 f1l 'll1I
1"1

IJ'
MI
1:& LlJ }I[

*-

fit

23

ries of the Kwanto Mountain. jub. Publ.
Comm. Prof. Yabe's 60th Birthday, vol.
1, pp. 457-479.
- - & SUZUKI, M. (1969): Geology of thebasin of the River Oboro-gawa, a tributary of the River Arakawa. Bull. Chichibu Mus. Nat. Hist., no. 15, pp. 1-18.
GREGORY, J. W. (1900): Jurassic fauna of
Cutch. Ceol. Surv. India, ser. 9, vol. 2,
pt. 2, pp. 1-195, pIs. 2a-27.
MORI, K. (1963): Geology and Palaeontologyof the Jurassic Somanakamura Group,
Fukushima Prefecture, Japan. Sci. Rep.
Tohoku Univ., 2nd ser. (Ceol.), vol. 35,
no. 1, pp. 33-65, pIs. 21-23.
NISHlMIY A, K. & YAMAGIW A, N. (1971):
Discovery of coral fossils from the Aoyuwadani, Tabayama-mura, Yamanashi
Prefecture. Ceol. Soc. japan, Jour., vol.
77, no. 12, pp. 791-792. (in Japanese)
Yamanashi Prefecture Government (1970):
Geologic map of Yamanashi Prefectureon the scale of 1: 100,000 and its explanatory text. pp. 1-240. (in Japanese)

Kumotoriyama
Shiosawa
Tabayama-mura
Kamosawa
Tabagawa

'ill.

=

~

tj

ft tit LlJ
~

LlJ
iR

*1
iR

ft lit. }IITrans. Proc. Palaeont. Soc. Japan, N.S., No. 89, pp. 24-26, pI. 7, April 20, 1973

613.

A TRIASSIC HEPATICA FROM THE OMINE COAL-FIELD,
SOUTHWEST HONSHU, JAPAN*
KAZUO HUZIOKA
Institute of .Mining Geology, Mining College, Akita University

and
EIT ARO T AKAHASIDepartment of Mineralogical Sciences and Geology,
Faculty of Literature and Science,
Yamaguchi University*MtmIII,It::::.:llHc:aXIHr,1S: ill I::JYr'!;**J!mrl:l*Hl'fm~0)~:(,iJ\M19E J::tJID(~~nt);b -.:=. "'!l) J: fJ ffIi: t..- t-::aH!1r,:P Hepaticites oishli UfrTffi) ~~[,J~£ t..- t-:, W!lliO) Onychiopsis Series(l:.ff~:/;L :7 ~c - Tff~B!'!!!*c) 0) Marchantites yabei KRYSHTOFOVICH (Thallites yabei(KRYSHT.) HARRIS le~I6G;fc-Cv'Q)

feltt..-, NliJd*O)~i\liJ:*m<, 7J~tffl1l\f,~MTIv't..-,IlJiHi~fJIijiJ' Gj&.fRiJ;*~ t..- -C v' Q, EJ:*J!i"I5O)~Hr,:p""(" Ji, Q,±

jt{~fiiJ]-93 . ;@jt{.f,~*Jl!I\

KON'NO, Equisetostachys (Neocalamites?)pedunculatus KON'NO takahashii KON'NO,
Neocalamites carrerei (ZEILLER) HALLE,
Todites goeppertianus (MUNSTER), T.
recurvatus HARRIS, Clathropteris menisco ides BRONGNIART, C. obovata OISHI,
Dictyophyllum sp., Cladophlebis haiburnensis (LINDLEY et HUTTON), C. nebbensis(BRONGNIART), c. sp., Taeniopteris minensis OISHI, Podozamites concinnus OISHIet HUZIOKA, and P. atsuensis T AKAHASI.
The Mesozoic hepatics are rather rarein the world. Thallites yabei (KRYSHTOFOVICH) HARRIS (syn. Marchantites yabei
KRYSHTOFOVICH, 1930), an only known
species of the Mesozoic Hepaticae inJapan, is one of the most characteristic
elements in the Onychiopsis Series (Upper Jurassic to Lower Cretaceous) ofJapan (OISHI, 1940), and also in the Naktong flora (Y ABE, 1905; KRYSHTOFOVICH,

Hepaticites oishii dealt with in thispaper occurred from the Upper Triassic
Momonoki Formation of the Mine Groupat the Hagimine colliery in the Omine
coal-field, Yamaguchi Prefecture, southwest Honshu. The Mine Group, a coalbearing deposit, was subdivided by KAT AYAMA (1939) into three formations, thelower or the Hirabara Formation, themiddle or the Momonoki Formation andthe upper or the Aso FOrmation.
The undermentioned fossil plants, together with Hepaticites oishii, have been
known from the Middle Carnian Momonoki Formation at the Hagimine colliery(TAKAHASI, 1959; KON'NO, 1962): Equisetites minensis KON'NO, E. takahashii* Received Nov. 30, 1972; read June 26,
1954. at Sendai.

24

613.

Hepaticites

1930) of southern Korea and the Nikan
flora (KRYSHTOFOVICH, 1930) of the Far
East.
Several form-genera of the fossil Hepaticae, such as Thallites, Hepaticites andMarchantites, have been established by
the previous authors. According to W ALTON (1925) and HARRIS (1942), Marchantites is restricted to the hepatics agreeing with the Oligocene Marchantites sezannensis SAPORT A, the type species, inwhich marchantiaceous air chambers,
ventral scales and reproductive organs
have been demonstrated. The formgenus Thallites W ALTON (1925) is to be
used for thalloid fossils showing no character proving that they belong to the
Hepaticae rather than to the Algae. The
form-genus Hepaticites WALTON is used
for thalloid fossils showing some definite
character of the Hepaticae. In practicethis is usually the rhizoids.
The thallus of Hepaticites oishii are
well-impressed on the dark-coloured
shale, but are not sufficiently preserved
to permit their anatomical studies. This
species is the oldest representative of
the Hepaticae in japan, being of theUpper Triassic (Middle Carnian) age.
At this place the writers express theircordial thanks to Mr. N. NAMBA whocollected the materials from the Hagimine colliery of the amine coal-field.
Description of species

Class Hepaticae
Form-genus Hepaticites W ALTON, 1925

Hepaticites oishii HUZIOKA et T AKAHASI
new species
PI. 7, figs. 1, la, lb, 2, 3

Description:- Thallus regularly and dichotomously branched, narrow, 0.3-0.4cm

25

broad, composed of midrib and lamina,
repeatedly forked at angles of 30-80 degrees and intervals of 1.0-1.5 cm. Midrib
thick, distinct, ridged at under surface,with numerous rhizoids. Lamina transversally bullate, often spotted, entire towavy at margin. Rhizoids borne on
midrib, rigid, regularly and densely arranged, frequently forked.
The midrib and rhizoids are shownin fig. 1a. The spotted surface (air chamber?) and the wrinkled surface are shownin fig. lb. The thallus of this fossil
hepatica is restored as shown in fig. 3.
Comparison and remarks :-Comparedwith Thallites yabei (KRYSHTOFOVICH)
HARRIS (1942) (syn. Marchantites yabei
KRYSHTOFOVICH, 1930) which is shownin fig. 4, a well-known Upper jurassicLower Cretaceous Hepatica in eastern
Asia, this Triassic thallus are much narrower, forking at shorter intervals. HARRIS (1931) reported two species of Hepaticites, H. globosus HARRIS and H. laevis
HARRIS, from the Thaumatopteris Zoneof Scoresby Sound, East Greenland. Of
them, H. oishii closely resembles H. laevis,
but in the former species the midrib isstrongly ridged at the under-surface andbears numerous rigid rhizoids which
are often furcated.
H. oishii is somewhat similar to but
distinguishable from the jurassic H.
arcuatus (LINDLEY et HUTTON) HARRIS(1942) (syn. Marchantites erectus (LECK.)SEWARD) from England, and the Upper
Triassic H. solenotus HARRIS (1938) ofEngland, and also Thallites uralensis
KRYSHTOFOVICH et PRINADA (1933) fromeastern Urals.
Occurrence and collection :-Hagimine
colliery, amine coal-field, Yamaguchi Prefecture; Momonoki Formation (Middle
Carnian) of the Mine Group. Holotype,
AKMG-7019; paratype, AKMG-7020.Coll.
N. NAMBA.

Kazuo HUZIOKA and Eitaro T AKAHASI

26

References
HARRIS, T.M. (1931): The fossil flora of
Scoresby Sound, East Greenland. Part 1.
Medd. om Cn/mland, Bd. 85, no. 2, pp. 1102, pis. 1-18.
- - (1938): The British Rhaetic flora. Brit.
Mus. (Nat. Hist.) , London, pp. 1-84, pis.
1-5.
- - (1942): On two species of Hepatics of
the Yorkshire Jurassic flora. Ann. Mag.
Nat. Hist., ser. 11, vol. 9, pp. 393-401.
KATAYAMA, M. (1939): Stratigraphical study on the Mine Series. jour. Ceol. Soc.
japan, vol. 46, no. 546, pp. 127-141. (inJapanese)
KON'NO, E. (1962): Some species of Neocalamites and Equisetites in Japan and Korea.
Sci. Rept. Tohoku Univ., 2nd ser., Special
Volume, no. 5, pp. 21-47, pis. 9-18.
KRYSHTOFOVICH, A.N. (1930): A Liverwort
from the Middle Daido Formation of Ko-

rea and the Nikan Series of the Manchu·
rian Border. Ann. Soc. Pal. Rus., Tome8, pp. 144-147, pI. 15.
- - and V.D. PRINADA (1933): Contribution
to the Rhaeto-Liassic flora of the Cheliabinsk Brown coal-basin, eastern Urals.
Trans. United Ceol. Prosp. Servo USSR,
Fasc. 346, pp. 1-34, pis. 1-5.
OISHI, S. (1940): Mesozoic floras of Japan.
jour. Fac. Sci., Hokkaido Imp. Univ., ser.
4, vol. 5, nos. 2-4, pp. 123-480, pis. 1-48.
T AKAHASI, E. (1959): Floral changes since
the Mesozoic Age of western Honshu,
Japan. Sci. Rept. Yamaguchi Univ., vol.
10, pp. 181-237. (in Japanese)
WALTON, J. (1925) : Carboniferous Bryophyta.
1. Hepaticae. Ann. Bot., vol. 39, pp. 563572. pI. 18.
Y ABE, H. (1905): Mesozoic plants from Korea. jour. Coli. Sci., Imp. Univ. Tokyo,
vol. 20, art. 8, pp. 1-59, pis. 1-4.

Mine
Momonoki
Omine

Aso
Hagimine
Hirabara

Explanation of Plate 7
Figs. 1, la, 1b, 2, 3. Hepaticites oishii new species, from the Hagimine colliery.
1. Holotype, AKMG-7019. 1a and lb. Enlarged to show the detailed characters of thallus,
x 3 and x 2, respectively. 2. Para type, AKMG-7020. 3. A restoration of Hepaticites
oishii.
Fig. 4. Thallites yabei (KRYSHTOFOVICH) HARRIS (syn. Marchantites yabei KRYSHTOFOVICH),
from the Naktong Series of southern Korea. Reproduced for comparison; YABE, 1905,
p. 41, pI. 3, fig. 16a; originally, Sagenopteris bilobata YABE var. major YABE.

HUZIOKA and T AKAHASI:

Hepaticites

Plate 7·/~'.\(-f( \ ·U
·.\

" !

J

i

". ...'\ .f

!

.' ·1

'1 !
..
l' t

f

,'1"
4

\

\
I

I

\

\

t

/

i,

/,J

/ 1\

.- .V?i(-'
1~ L,./
i/o _cSHIBATA

photo.

Trans. Proc. Palaeont. Soc. Japan, N.S., No. 89, pp. 27-41, pI. 8, April 20, 1973

614.

V ASCOCERATID AMMONITES FROM THE TURONIANOF HOKKAIDO(STUDIES OF THE CRETACEOUS AMMONITESFROM HOKKAIDO AND SAG HALlEN-XXVI)T ATSURO MATSUMOTO
Department of Geology, Kyushu University, Fukuoka 812

j:l .::. 7 :/il!;/ A::! t: '7 Af4~::fi: Vascoceratidae i'i'f 7- AfflO)&i1iDl~lc
i:. I:.lc~tffljJ[O)-fl}11 7-;L - j:l .::. 7 :/iJ' f?
~nlcf.!.l±\~nt.: 4 tf,*a-, Vascoceras sp. aff. V. durandi (THOMAS and PERON),
Fagesia thevestensis (PERON), Fagesia sp. d. rudra (STOLICZKA) 0) 3 fllHcnrJ5E L..,
~tffljJ[ 7-;L -~,!1i:-t ~d)~,*ifi$:ti!!.~"t'·i'i ~ :h6b-C ~ nL';!i,.Q 0~~L..-C~ti-t.Qo~f?~7-;L-j:l.::.7:/M~~~.QnrJM.::fi0)~~~~m~~~a-A~,

ilifIiillia-ilit.7;..Qo Vascoceratids O)!j;f1ftJ:~:(!1vi, .lj3.IC~!t!mIl~"JtJ:~IJ;fd~I:rL'tJ: <, .;c-O)lg~t:'7!11 r iM*~.g.*Jl!, ~.Q i:.1t'IDl~, !tHfljO)~5jfr.;j: cO)~fr;'ICMrc.: L.. t.: ~filU:O)!j;f~M:a- 'h~
1.\\!:-t ~~ L', :1:.,iffiJ~rIf~~ilSfiRJ:\fct1f~l1:iJ;;!i, -;,f.:o)L'vi;!i,.Q ~ It'iJ'o 13 *IC 'hJ}l";: l..t.: 3 f<l!
vi, tU!X(j"J~~fI*Jl!iJ~fll#t"t'·, t.:~:Iv;Qfi*i:1L'~:(fjO)J1\It' flll'J1]iL';!i,-::> Lt. oj IOJ!,:hn.Qo ~iiIEO)liffi~M: 'h ~;t oj .Q iJ;, ~:(!1a-~~rr.Q Ie ~ t.: fJ, 7 :/7T~jl'tII4l-!t!HC -:J1t, -C ~QN~iJ;::fjJ/lltJ: i:. C
iJ;~MiJ,n.Qo~*

i~ ~I;

had to omit for brevity in my last paper.
Before entering in the description I
thank Dr. Hakuyu OKADA and Mr. Kazunari TANABE who found an interesting specimen in a field work with me
and provided it for this study, Mr. Hajime KOYAMA who showed me anotherinteresting specimen, and Professor Tetsuro HANAI who gave me facility to
study some specimens preserved in the
University of Tokyo. Dr. Itaru HAY AMI
and Miss Mutsuko HAYASHIDA kindly
assisted me in preparing the plates andtypescript.

Introduction

The Vascoceratidae are typically Tethyan in occurrence. This has been
evidenced by numerous works of predecessors. In my concise note on thepalaeobiogeography of the late Cretaceous Ammonoidea (MATSUMOTO, 1973), I
have mentioned that some immigrantsof the vascoceratids are found outside
the Tethys region.
In this paper I describe three speciesof the vascoceratid ammonites which
occurred rarely in the Turonian of Hokkaido. After the description supplementary remarks are given on the biogeographical distribution of the vascoceratid
ammonites, presenting a map which I

Description of Species

Family Vascoceratidae SPATH, 1925
The classification of the Vascocerati·
dae is more or less different between* Received Dec. 31, 1972; read Jan. 16, 1973
at Sendai.
27

Tatsuro MATSUMOTO

2R

authors (e. g. FUR ON, 1935, SCHNEEGANS,
1943; REYMENT, 1954, 55; BARBER, 1957 ;WRIGHT, 1957; COLLIGNON, 1958; WIEDMANN, 1964; FREUND and RAAB, 1969).
The material from Japan is too poor togive comments on the problem, but I
had fortunately an experience to lookthrough a number of specimens from
Nigeria and talked with REYMENT and
BARBER, though separately, when wewere studying Cretaceous ammonites at
the British Museum (Natural History),
1953-54. From my general impression,
I was rather inclined to agree with
SCHNEEGANS' attitude in treating of thematerial. Anyhow, it is worthwhile tonote that great many varieties of vascoceratid ammonites were differentiatedwithin a short time interval in the earlyTuronian of the Tethys region. It would
be interesting to study the mode ofevolution through examining ontogeny
and variation from population to population.
To describe a small number of incompletely preserved specimens from Hokkaido, I have to depend, for conveniencesake, on some previously proposed
framework of classification.
Genus Vascoceras CHOFF AT, 1898

Type-species :- Vascoceras gamai CHOFFAT, 1898.

Remarks:- This genus has a globose
to sub globose, involute shell, in whichumbilical tubercles and fold like costae
are distinct only in early stages andlost with growth. In Fagesia PERVINQUIERE, 1907, the shell is inflated cadicone with ornament similar to but more
persistent than that of Vascoceras. In
Plesiovascoceras SPATH, 1926, the whorl
is evolute, coronate and ornamented withstrong obtuse tubercles at the umbilicalmargin. The three genera are closely

allied to one another and have more orless similar sutures. Typically the suture of Fagesia is the most complicatedof the family. It is more deeply incisedand consists of higher saddles and narrower or deeper lobes than that of Vas-

coceras.
In Paravascoceras FURON, 1935, there
are external ribs but no umbilical tubercles. Pachyvascoceras FURON, 1935, isregarded by SCHNEEGANS (1943), BARBER(1957), and FREUND and RAAB (1969) asa synonym of Paravascoceras, because
there is a continuous series between
their type-species, V. crassum FUR ON,
1935, and V. cauvini CHUDEAU, 1909. I
would agree with FREUND and RAAB
(1969, p. 19) in regarding Broggiliceras
BENAVIDES-CAcERES, 1956, as anothersynonym of Paravascoceras. Some species which were referred to Pachyvascoceras may not automatically transferred to Paravascoceras. V. durandi (THOMAS and PERON) can be reckoned assuch an example.
Discovascoceras COLLIGNON, 1958 wascommented by WIEDMANN (1959, p. 757)
as indistinct. Its type-species, D. tesselitense COLLIGNON 1958, was regarded
by WIEDMANN as a synonym of Vascoceras (Pachyvascoceras) triangulare FARAUD, 1940 and then by FREUND and
RAAB (1969, p. 23) as that of Paravascoceras obessum (TAUBENHAUS, 1920).
COLLIGNON (1965, p. 179 [17J), however,redefined Discovascoceras, describing thechange of characters with growth andthe presence of three keels. On the
grounds of its stratigraphic occurrenceat the very base of the Turonian andalso of its morphologic characters, COLLIGNON considers to allocate Discovascoceras at the initial part where the
Vascoceratidae and the Tissotiidae aregoing to branch apart (see REYMENT,
1954, text-fig. 8).

614.

Vascoceratid ammonites

Vascoceras sp. aff. V. durandi(THOMAS and PERON)
PI. 8, Fig. 1, Text.fig. 1

Compare.1889. Pachydiscus durandi TlIm,'IAs et PlmOi':,
Moll. Foss. Tunisie, p. 27, pI. 18, figs.
5-8.
1896. Pachydiscus durandi, PERO,\, Minn. Soc.
Geo!. France, Pa!eont., Mem. No. 17, p.
44, pI. 4, fig. 1 ; pI. 5, fig. 1; pI. 17, fig. 5.
1898. Vascoceras douvillei CIIOFF AT, Faune
Cre!. Portugal, p. 59, pI. 10, figs. 3, 6;
pI. 11, figs. 2-5; pI. 21, figs. 13-16.
1907. Vascoceras durandi, PERVli'\QUIERE,
Etudes Pa!eont. Tunisienne, vol. 1, p.
332, pI. 21, fig. 1a-b; text-fig. 125.
Diameter
115mm.(1)Umbilicus
26(.23)The coiling is considerably involuteand seems to be somewhat eccentric.
The whorl is inflated on the flanks,
broader than high and broadest near
the umbilical shoulder in the lower part.
The flanks are convergent passing intoa rather flattened venter which occupies
the breadth of the external lobe. The
shell is as a whole subglobose in general
aspect.
The umbilicus is deep and surrounded
by a high and vertical wall which forms
an angular edge with the flank.
So far as the observable part is concerned, the shell is free from distinct
ornament. But a very faint buldge isdiscernible outside the umbilical edgeof the outer whorl. This is probably a
reduced umbilical tubercle, although no
distinct umbilical nodes are observed on
the visible part of the inner whorl.
From the low buldge mentioned aboveruns a very indistinct, low rib, on which
several fine rib lets and lirae are superimposed. They are somewhat prorsiradiate and gently concave on the flank.

29

1965. Vascoceras (Pachyvascoceras) aff. douvillei·durandi, COLLIGNON, Atlas Fossiles
Caracteristiques de Madagascar (Ammo.
nites), fasc. 12, p. 43, pI. 394, fig. 1676.
1969. Vascoceras durandi, FREu'>D and R,AAB,
Special Papers in Palaeont., no. 4, p. 29,
text·fig. 6h-i.

Material :-GK. H5674, from loc. R2513a,
the Obira, collected and supplied by H.
C)KADA and K. TANABE, 1971.
Description:-The specimen is fairly
well preserved on the right side but
badly crushed on the left. It is wholly
septate and the body-whorl has mostly
gone. The measurements are as follows:
I-Ieight
54(.47)

Breadth33x2(.57)

B.;H.
1. 22

The mode of ribbing, if any, on theexternal part of the inner whorl is notknown, as it is difficult to take out the
deeply overlapping outer whorl. Onlya faint umbilical buldge is perceptible.
Sutures are pseudoceratitic, with pointed lobules and phylloid folioles. The
external lobe (E) is subrectangular inrough outline, although its details are
obscured for unfavourable preservation.
The first lateral saddle between E and
L is asymmetric, steep on the side of
E and gently sloping towards L. The
first lateral lobe (L) is opened V -shapedin rough outline, moderately deeply incised by several lobules and bipartite
at the bottom. The second lateral saddle is roughly dome-shaped and shallowly incised on top, leaving phylloid folioles. The second lateral lobe (U2) ismuch narrower than L and moderately
incised, having a narrow and deeply
pointed lobule at its bottom. The saddle inside U2 is nearly as high as thesecond lateral saddle, sub quadrate inrough outl ine anr. apparently tripartite

Tats~i?"O MATSUMOTO

30

/
,/

/
I

/

/

I
{

I
\

//

\\

(

\ \ '- _ _ _ _ I

\

I

\

/

I
{

\

{

\
\
\
\

\
\
\

E

f

u.s .

<
\"

c

10 mm

,

Fig . 1. Vascoceras Sp. aff. ]I. durandi ( TH OM AS and P ERO N ) . GK . H5674, frorn loco
R2513a . Pho tog raphic late r a l v ie w (a), w ith w hit e nin g ; di agram matic fr ontal v iew ( b);external suture ( c) at s .

into folioles of unequal size, w ith thenarrowest one on the side of U2, th emoderate one in the middle and thebroadest one on the umbilical shoulder.
Narrow U3 and a part of U5 are v isibleon the umbilical wall.

ComjJarison :- A lthough the specimen
from l-lokka ido is incompletely preserved ,the olBer ve d characters su gge st that it
is very close to the extreme end of theva riation of Vascoceras durandi (THOMAS
and P ERON) (see P ERV 1NQU 1ERE, 1907),

614.

Vascoceratid ammonites

which has been known from the Lower
Turonian of Tunisia, Algeria, Portugal
and Israel. Its umbilicus is considerably
narrower than that of the measured spe·cimens from these areas. In the dimen·
sions of the umbilicus, whorl·height andbreadth in proportion to the diameter,
the present specimen is closest to thespecimen from the Turonian zone ofMammites conciliatum of Madagascar de·scribed by COLLIGNON (1965b) under thename of Vascoceras (Pachyvascoceras) aff.
douvillei·durandi. It differs from that
Madagascar specimen in that it has lessdistinct ornament and not so sub quad·rate whorl·section as the Madagascar
one. It has rather a rounded subtrape·
zoidal whorl-section. In this respect andin the general aspects in lateral view,
the present specimen is apparently similar to Choffaticeras meslei (PERON, 1897),
from the Turonian of Algeria and Israel.
It is distinguished from that species by
the absence of keels, more inflated andbroader whorls and more complex sutures which show dissimilar pattern(compare Text-fig. 1 with PERON'S pI.
17, fig. 1). The suture of the Hokkaido
specimen is close to, if not exactly identical with, that of P. durandi as illustrated by PERVINQUIERE (1907, text-fig.
125).
On account of the characteristic shellform and the indistinct, almost obsolute
ornament, I thought that the present
form could be referred to Discovascoceras COLLIGNON, 1958, which should be
provided with keels as redefined by COLLIGNON (1965b). Our species has no keel
and the lobes of its suture are moredeeply incised and less numerous on theflank than those of D. tesselitense COLLIGNON and other related species. Thus
it is not referable to Discovascoceras, if
Discovascoceras is excluded from Paravascoceras as independent genus.

31

To sum up, it is difficult to identify
precisely the present form, unless thecharacters of the early shell is more
distinctly known. It is, however, highly
possible that the described specimen isclose to the extreme end of the variation of Vascoceras durandi (THOMAS andPERON), although it could represent ageographically separated subspecies oranother closely allied species.
Occurrence :- The described specimen
was obtained at loco R2513a, from a calcareous nodule in the mudstone of the
Middle Yezo Group exposed along the
Okufutamata, a tributary of the Obira[Opirashibets], by H. OKADA and K.
TANABE, during the field work conducted by me in 1971. The mudstone of loco
R2513a is allocated at the middle of unit
Mj of TANAKA (1963). No mega-fossils
were associated with this ammonite at
this locality. In the Nakakinebetsu valley, another tributary of the Obira, about
13 km southwest from loco R2513a, aprobable extension of unit Mj and subjacent part down to the middle of unit
Mh are more prolific, where I obtained,
among others, Inoceramus (Mytiloides)
labiatus, Mammites sp., and Pseudaspidoceras sp. On the grounds of available
evidence I refer the sequence from theupper part of Mh to the top of Mj to
the Lower Turonian.
Location of R2513a is about 850 m eastand 3900 m south from the point, N 44 °10'-E 142°0', northwest corner of the
Horokanai sheet, geological map of
1: 50,000, published by the Geological
Survey of Japan (1958). It is in the
Rumoi district, Teshio province, northwest Hokkaido.
Genus Fagesia PERVINQUIERE, 1907

Type-species :-Olcostephan us superstes
KOSSMAT, 1897.

Tatsuro MATSUMOTO

32

Remarks:- This genus is well defined,
as mentioned by FREUND and RABB (1969,
p. 33). They commented, however, thatnumerous species of very wide distribution included later in this genus differ considerably from a well-definedgroup formed by F. superstes, F. thevestensis, F. rudra, F. peroni and F.
bomba, from India, Middle East, Spainand North Africa, so that in several
cases the generic position is uncertain.
In fact, Fagesia cali/ornica ANDERSON,1935 and F. shastensis ANDERSON, 1935,
from California, should be better transferred to Plesiovascoceras, as I havealready pointed out (MATSUMOTO, 1959,
p. 102).
The species from Japan described
below undoubtedly belong to the typical group of Fagesia.

Fagesia thevestensis (PERON)
PI. 8, Fig. 2, Text-fig. 21896. Mammites (?) thevestensis PERON, Mem.
Soc. Ceol. France, Pateont., Mem. No. 17,
p. 23, pI. 7, figs. 2, 3.
1904. Ammonites kotoi Y ABE, Jour. Coll. Sci.,
Imp. Univ. Tokyo, vol. 20, art. 2, p. 26,
pI. 6, figs. 3 and 4.
1907. Fagesia thevestensis, PERVINQUIERE,
Etudes Paieont. Tunisienne, vol. 1, p.
325, pI. 20, figs. 5, 6; text-figs. 123, 124.
1969. Fagesia thevestensis, FREUND and RAAB,
SPecial Papers in Paiaeont., no. 4, p.35,
text· fig. 7g.

Material :-GT. 1-3249, from loco T610bc, Saku area (colI. T. MATSUMOTO). GT.
MM 7561, from a rolled nodule of the
Yubari, the holotype of Ammonites kotoi,
is also examined.

Measurements :Specimen
GT. I-3249
GT. MM7561
PERON'S type

Diameter
80(1)100(1)
80(1)Umbilicus
20(.25)27(.27)25(.31)

Descriptive remarks :-GT. 1-3249 represents a fairly large individual, but
its outer whorl, is incomplete and shows
eroded sutures. Therefore its inner
whorl alone is measured and illustrated.
GT. 1-3249 and GT. MM7561 are complementary with each other, showing
altogether the following characteristicfeatures.
The outer whorl, of GT. 1-3249, thoughincomplete, is as large as or somewhat
larger than the large specimen of PERVINQUIERE (1907, pI. 20, fig. 6). They areboth still septate and the shell musthave reached about 200 mm., if the adultbody-whorls were preserved. PERON'S
holotype must have been an immatureshell, with which the inner whorl of GT.
1-3249 is well comparable in dimensions.

Height36(.45)44(.44)34(.42)

Breadth45(.56)53(.53)45(.56)

B./H.
1. 25
1.2
1.3

Some difference in the size of umbilicus
is probably within the variation of a
species, as FREUND and RAAB (1969, p.
35) generally admitted.
The whorl is inflated and lunate incross-section. It is broader than high,but is not so broad as in F. superstes(KOSSMA T, 1897). The measured proportion between height and breadth isof the order of that measured on the
PERON'S type, PERVINQUIERE's specimens
I and III. The large outer whorl of GT.
1-3249 seems to have broader whorl but
cannot be accurately measured. The
venter of the inner whorl is moderately rounded as in the holotype of PERON
and the smaller specimen of PERVINQUIERE (1907, pI. 20, fig. 5). The umbili·
cal wall is very steep or nearly vertical.

614.

33

Vascoceratid ammonitescF ig. 2. Fage sia the ves tensis ( P ERON) . GT. MM7561 , holot ype of Ammoni tes kotoi Y ABE,
,1904, from Yubari. Kyushu Univ. [ 1. H .] photos, with whitening.

The periumbilical tubercles are somewhat bullate or subrounded. Although
they are not completely preserved in
the Hokkaido specimens, they seem tobe similar in intensity and density tothose in the specimens described by
PERVI NQUI ERE. The ribs are also very
similar in intensity, density, forwardcurvature and mode of bifurcation to
those of the holotype and the inner
whorl of P ERV INQUIERE. In the two specimens from Hokkaido some of the ribs
are slightly more elevated than others
and some of the interspaces are somewhat deeper than others, if not forming
distinct constrictions. On the outerwhorl the ornament is much weakened.
The suture is of typical Fagesia pattern, showing three, high, deepl y incised
saddles on the side and deeply incised
lobes. The first lateral saddle is tripartite. L is nearly as deep as E andbifid at the bottom. The terminals of
the subdivided folioles are somewhat
phylloid.

To sum up the two specimens from
Hokkaido are identified with Fagesia
th evestensis ( P ERON), from the Lower
Turonian of Algeria, Tunisia and Middle East. It is still highly possible that
Fagesia thevestensis ( P ERO N, 1896) andFagesia superstes (KOSSMAT, 1897) could
be within the variation of one and thesame species, as PERON (1897, p. 84) at
first pointed out. Without treating asufficient number of specimens fromplace to place (Algeria, Tunisia, Israel
and India), I cannot decide the matter.
For the time being the two species are
distinguished as PERVI ' QUIERE did. As
there is no reason to regard the Japanese form as an independent species,
Ammonites Iw toi Y ABE, 1904, falls in thesynonym of Fagesia th evestensis ( PERO N,
1896).
Occurrence :-GT. 1-3249 was obtainedby myself from lac. T610- c, Onoderazawa, a small branch of the Sakugawa[Sakotan-gawa], unit IIc of the Middle
Yezo Group in the Saku area, Teshio

TatS'lLrO MATSUMOTO

34

province, northwest Hokkaido (for thelocation see a map of pI. 12 in MATSUMOTO, 1942). From another locality ofunit IIc about 3 km west of Saku, I haverecently obtained Inoceramus (Mytiloides)
labiatus, which indicates Lower Turonian.
GT. MM7561 was obtained by Y ABE ina rolled nodule of the Yubari, Ishikariprovince, central Hokkaido. Although I
did a field work in the same area, I was
not successful to get the same species
from this area.
Fagesia sp. cf. F. rudra (STOLICZKA)

Text-fig. 3Compare.1965. Ammonites rudra STOLICZKA, Palaeon!.
Indica, ser. 1, vol. 1, p. 122, pI. 60.
1897. Olcostephanus rudra, KOSSMAT, Beitr.
Paliiont. Oesterr. Ungarns, vol. 11. p.
29 [136J.
1907. Fagesia rudra, PERVINQUIERE, Etudes
Pateont Tunisienne, vol. 1, p. 322.
1965. Fagesia rudra, COLLIGNON, Atlas Fossiles Caracteristiques de Madagascar(Ammonites), fasc. 12, p. 48, pI. 369,
fig. 1678-A; pI. 397, fig. 1678-B.

Material :-A specimen in the private
collection of Hajime KOYAMA, Saku, Nakagawa-machi, Hokkaido.

Measurements :Specimen
KOYAMA'S
STOLICZKA'S
COLLIGNON'S

Diameter
120.0(1)170 (1)180 (1)

Umbilicus
29.5(.24)- (.23)36 (.20)

Descriptive remarks :-The last suture
is seen at 120 mm. in diameter and themain part of the body-whorl is badly
crushed. Even the septate whorl issomewhat deformed on the left side.
The above measurement of the whorlbreadth is approximately estimated fromthe little deformed right half. (see Fig.
3b).

The proportion of the dimensions of
Koy AMA's specimen is closer to STOLICZKA'S type rather than to COLLIGNON's
specimen. The shell is considerably involute and subglobose in aspect. The
whorl is about twice as broad as high,broadest between the umbilical shoulders, and semi-lunate in section. The
umbilicus is narrow and deep, with asteep and high wall and subangularshoulder. The venter is broadly convex.
The last septate whorl is nearlysmoothish, but very faint, blunt umbilical bulges are perceptible and also weak

Height50.5(.42)- (.39)95 (.53)

Breadth100(.83)
136(.76)

Bj.H.
2
2. 12
1. 43

fine ribs are discernible, showing agently forward curve on the venter.
Fine lirae may be seen on the interspaces of the ribs.
The ornament of the inner whorl is
not well seen.
The suture is of typical Fagesia pattern, showing asymmetrically tripartite,
high, lateral saddles and deeply incised
lobes.
The observed characters indicate thatthe described specimen is probably referable to Fagesia rudra (STOLICZKA),
which has been reported from the Lower
Turonian of southern India (STOLICZKA,
1865; KOSSMAT, 1897), Madagascar (COLLIGNON, 1965) and Spain (WIEDMANN,
1964, listed only).
Occurrence:-H. KOYAMA obtained thisspecimen at a locality close to my loco
T1158 or T1159, in the mudstone of unitIIc, Saku-gakko-no-sawa, Abeshinai-Saku
area, Teshio province, northwest Hok-

614.

Vascoceratid ammonites

35

E

..

"

! / ,/,/:/,/,' "
I'

:

':'

,"

:

:

,f

:, ~:"I.'". ,, ,'

•

I

;' "

c

Q

b

20mm

Fig. 3. Fagesia sp. d. F. rudra (STOLlCZKA). Diagrammatic sketch of a specimen of
H. KOYA~IA'S collection from the Saku area. Lateral view (a), whorl-section (b) and external suture (c) at S.
(T.M. delin.)kaido. Calcareous nodules in unit IIc
often shows a particular, light greenish
tint. The rock matrix of this fossil
shows the same colour. Unit lIe in thisarea contains Inoceramus (Mytiloides)
labiatus and is assigned to Lower Turonian.

Note on the Biogeographical Distributionof the Vascoceratid Ammonites

Fig. 4 is a Turonian palaeogeographic
map (without restoring the relative position of continents) in which the occurrence of selected ammonites are indi-cated. A comprehensive discussion on
the palaeobiogeography has already been
given in my last paper (MATSUMOTO,
1973). Here is given a supplementary
note.
While several Turonian genera of 'theAcanthoceratidae and the Collignon iceratidae, such as J(anabiceras, Watinoceras, Mammites, Pseudaspidoceras, Romaniceras, Collignoniceras and Subprionocyclus show world-wide distribution,ammonites belonging to the Vasco<::eratidae and also the Tissotiidae characteristically occur in the Tethys Sea region in a broad sense.
Although the generic assignment of

Tatsuro MATSUMOTO

36.. Vascoceras, Plesiova5Coceros• PuzQsia, Mesopuzosioo Mammites, Pseudaspidocera~'" Paravascoceraso Tetragoniles• Watinoceros. SumitomocerasHo Metoicaceros

t:. Fcgesia•

Collignoniceros. Selwynoceras

Nigericeras, GomheocefOS

v Neoptychites

Fig. 4.

x Hyphontoceras, Scophites, Otoscaphites? Data wanted

Turonian palaeobiogeographic map of selected ammonoids.

the higherto described species has beenrevised in the light of up-to-date classification, some personal difference in
identification may still remain. I am
afraid that this may affect the conclusion about the distribution of the vasco cera tid genera.
According to FREUND and RAAB (1969),
Paravaseoeeras (in their sense) and Paramammites prevail in the southern shelfseas of the Tethys, such as those in the
Middle East, Lybia, Tunisia, Algeria,
Niger, Nigeria, eastern Brazil and Peru,
while Vascoeeras and Plesiovaseoeeras
are common in the northern shallow
seas, such as those in Portugal, Spain,

southern France, Mexico and Texas.
The two authors state, furthermore,that Nigericeras and Gombeoceras areoften associated with the Paravaseoeeras
fauna and that Metoicoceras is so with
the Vascoceras fauna. It is noted, however, that Gombeoceras occurs in Turkestan which is certainly situated on
the north side of the Tethys, that Vascoceras is found in the Middle East andNigeria where Paravaseoeeras prevails
and that Paravascoceras is found in
Mexico and adjacent parts where Vascoeeras and Plesiovascoeeras abound. A
species of Nigericeras represented by aconsiderable number of specimens has

614.

Vascoceratid ammonites,......

foc;I'?;,'

recently been reported from southeastern Colorado (COBBAN, 1971) and anotherspecies closely allied to N. glabrum
BARBER from somewhere (locality uncertain) in North America (MATSUMOTO
and OTSUKA, 1969). Furthermore, . there
are several genera of wider distribution,such as Fagesia, Neoptychites and Thomasites, which are not restricted toeither of the northern and southernsides of the Tethys. The barrier between the northern and the southetn
provinces, if ever existent, must nothave been very rigid. It was probably
represented by a geosynclinal part of
the Tethys, which may have comprised

37

deep troughs and also rIsmg islands.
Currents may have passed across somepart of this incomplete barrier.
I do not think that Metoicoceras of the
Acanthoceratidae (Metoicoceratinae) is
so intimately associated with the Vascoceras fauna. It is most commonly
distributed in the Gulf Coast and theInterior provinces of North America,
ranging from the upper Upper Cenomanian to the lower Lower Turonian.
It is distributed fairly commonly inwestern Europe and Morocco, suggesting an intimate faunal relation on bothsides of the north Atlantic ocean, which
may have been narrow in the late Cretaceous times. The genus is rarely
found in the Middle East, southern
India, Nigeria and Madagascar (?) (thelatter two areas in Upper Cenomanianonly), but no immigrant nor floatedremnant of Metoicoceras has been foundin the Pacific region (see SASTRY and
MA TSUMOTO, 1967).
It would be desirable to examine thefaunal comparison by means of a quantitative measure, as has been demonstrated by REYMENT and T AIT (1972, p.
322). It would be likewise necessary toobserve carefully the mode of occurrence of the fossils. For instance, it
has not yet been settled whether thesmall assemblage of the vascoceratids
found at a locality in southern Montana(reported by REESIDE, 1923) represents
an ammonite faunule migrated from thesouth or merely implies an accumulationof dead cephalopod shells floated fromthe main distribution area of the vascoceratids in the south. A map ofpalaeocurrent in the early Turonianseaway in North America has alreadybeen inferred by CLOUD (1961, fig. 10).
This was hinted by the known distribution of the straight shelled ammonoid
Sciponoceras gracile (SHUMARD), but canTatsuroMATSUMOTO

also serve for explaining the dispersal
or migration of Plesiovascoceras up to
Montana and also to California.
Vascoceratid ammonites indeed seem
to l;>e generally prolific in the shelf seasof the Tethys, but they actually occur
in particular beds which occupy a restricted part of Lower Turonian sequence, as recorded precisely by FREUND
and RAAB (1969). This is also true forthe mode of occurrence of the vascoceratids in southern Montana. The factsuggests that the ammonites belonging
to the Vascoceratidae which are charact~rized by more or less pseudoceratitic sutures and reducing ornament may
have had a particular habitat and mod~of life. In this connexion it should be
noted that Neolobites of the Engonoceratidae, which are phylogeneticallyunrelated to the Acanthocerataceae but
have likewise pseudoceratitic sutures,
occurs characteristically in the late
Cenomanian shelf seas of the Tethys,i. e. generally the same region as themain distributional area of the Vascoceratidae and the Tissotiidae. It should
be furthermore noted that Fagesia which
has more deeply incised sutures thanother genera of the Vascoceratidae
shows wider distribution.
Now in Japan the ammonites belonging to the Vascoceratidae are very few
and no examples of the Tissotiidae andthe Coilopoceratidae have been found.
Only one specimen of Vascoceras sp. aff.
V" durandi, tw:o of Fagesia thevestensis
and another of Fagesia sp. cf. F. rudra
are known. from the Lower Turonian of
Hokkaido. This number of specimens
is incomparably smaller than that ofhundreds of vascoceratid ammonites
from Nigeria. On the other h~md desmoceratids and tetragonitids, which ocwr abundantly in Japan, are distributed
very sparsely or almost absent in theTuronian shelf seas of the Tethys.
As the genus Fagesia is generally
widespread, it would not be surprising
to find several fossils of this genus fromJapan. Their occurrence can be interpreted as either a rare immigrant or a
float brought by current from the south.
The known distribution of Fagesia. thevestensis is, however, much apart fromJapan, unless F. superstes is proved t9
be conspecific with F. thevestensis. A
find of Vascoceras sp. aff. V. duran# is.
rather puzzling, since V. durandi occurs
in Algeria, Tunisia, Portugal and Israel,
which are situated around the pr:esent
Mediterranean Sea. The Japanese formis, however, allied to a form from Madagascar called V. aff. choffati-durandi.
There are several species of other families which are common or closely allied'between Japan and Madagascar but not
yet found elsewhere. This is probably
caused by too poorly explored situationof the Cretaceous areas in southernAsia.
I have reported recently (MATSUMOTO,
1971) a find of Hourcquia from south
Saghalien, a specialized member of the
Vascoceratidae in late Turonian-Coniacian times. Professor W. HASHIMOTO
has shown me another specimen of
Hourcquia obtained in the AbeshinaiSaku area, Teshio province, Hokkaido.
According to HASHIMOTO (1973), it probabl)' came from the Coniacian. Although
the age is younger than Lower Turonian,
this is another example of the genus
known only from Madagascar and theJapanese biogeographic province.
To sum up, it is worthwhile to reportthat several vascoceratid ammonites do
occur, though rarely, in the Japaneseprovince. To explain this fact we needmore information about the Cretaceousammonites in southeast Asia and adjacent areas which are expected to have

614.

Vascoceratid ammonites

been the nearest source of the northward dispersal or migration of the vascoceratid ammonites up to Hokkaido
and south Saghalien.

References Cited
ANDERSON, F.M. (1935): The genus Fagesia
in the Upper Cretaceous of the Pacificcoast. Jour. Paleont-., vol. 5, no. 2, pp.
121-126, pis. 15-17.
BARBER, W. (1957) :', Lower Turonian ammonites from north-eastern Nigeria. Ceol.
Surv. Nigeria, Bull. no. 26, pp. 1-86, pis.
1-34.
BENAVIDES-CAcERES, V.E. (1956): Cretaceous system in northern Peru. Bull. Amer.
Museum Nat. Hist., vol. 108, no. 4, p. 353494, pis. 31-66.
CHOFF AT, Paul (1898): Recueil d'etudes paleontologiques sur la faune cretacique duPortugal. Vol. 1, Especes nouvelle ou peu
connues. Commisso Trav. Ceol. Portugal,
ser. 2, pp. 41-86, pis. 3-32.
CHUDEAU, R. (1909): Ammonites du Damergou (Sahara meridional). Bull. Soc. Ceol.
France" ser. 4, vol. 9, pp. 67-71, 5 pis.
CLOUD, P.E. Jr. (1961): Paleobiogeographyof the marine realm. In Oceanography,
Amer. Ass. Adv. Sci., pp. 151-200.
COBBAN, W.A. (1971): New and little-known
ammonites from the Upper Cretaceous(Cenomanian and Turonian) of the western interior of the United States. U.S.
Ceol. Surv. Prof. Paper 699, pp. 1-24, pis.
1-18.
COLLIGNON, Maurice (1958): Cephalopodes
neocretaces du Tinrhert (Fezzan). Ann.
Pateont., vol. 43 (1957), pp. 115-136 [3-24J,
pis. 16-18 [1-3].
- - (1965a): Nouvelle ammonites neocretacees Saharienne. Ibid., vol. 51 (1965),
pp. 165-202 [3-40J, pis. A-H.
(1965b): Atlas des Fossiles Caracteristiques de Madagascar (Ammonites), fasc.
12, pp. 1-82, pIs. 376-413; Servo Geol.,
Tananarive.
F ARAUD, M. (1940): Le genre Vascoceras
dans Ie Turonien du Gard. Bull. Soc.

39

Sci. Nat. Vacucluse, vol. 3, pp. 3-24, 11
pis.
FREUND, Raphael and RAAB, Menahem (1969) :Lower Turonian ammonites from Israel.
Special Papers in Palaeont., no. 4, pp. 183, pis. 1-10.
FURON, R. (1935): Le Cretace et Tertiaire
du Sahara Soudanais. Arch. Mus. Hist.
Nat., Paris, ser. 6, vol. 13, pp. 1-97, pis.
1-7.
HASHIMOTO, Wataru (1973): Hourcquia hataii HASHIMOTO, a new species of ammonite from the Upper Cretaceous Systemof the Abeshinai region, Teshio province,
Hokkaido. Professor Kotora Hatai Memorial Volume, pp. 315-318, pI. 35, Sendai.
KOSSMAT, Franz (1897): Untersuchungentiber die stidindische Kreideforma tion
(Zweiter Theil). Beitr. Paliiont. Oesterr.Ungarns u. d. Orients, vol. 11, pp. 1-46
[108-153J, pis. 1-8 [12-19J.
MATSUMOTO, Tatsuro (1942): Fundamentals
in the Cretaceous stratigraphy of Japan.
Part. I. Mem. Fac. Sci., Kyushu Univ.,
ser. D, vol. 1, no. 3, pp. 129-280, pis, 5-20.
- - (1959): Upper Cretaceous ammonites ofCalifornia. Part II. Ibid., Special vol. 1,
pp. 1-172, pis. 1-41.
- - (1971): Uncommon keeled ammonites
from the Upper Cretaceous of Hokkaido
and Saghalien. Ibid., vol. 20, no. 2, pp.
305-317, pis. 48-49.
- - (1973): Palaeobiogeography of Late
Cretaceous Ammonoidea., In HALLAM,
A. [EditorJ : Atlas of Palaeobiogeography,
Chapt. 38, Elsevier, Amsterdam.
MATSUMOTO, Tatsuro and OTSUKA, Hiruyuki(1969): Ammonite collection donated by
Yukichi OGAWA to the Museum, Bureauof Cultural Center, Kagoshima. Sci. Repts.
Dept. Ceol., Kyushu Univ., vol. 10, no. 2,
pp. 51-65, pis. 7-15 [in Japanese withEnglish abstract].
PERON, Alphonse (1889-90): Description desmollusques fossiles des terrains cretacede la region des Hauts Plateaux de laTunisie. Recueillisen 1885 et 1886 par M.
Phillipe THOMAS. Exploration scientifiquede la Tunisie, 405 pp., 35 pis., Paris.
- - (1896): Les ammonites du Cretace su-

40

Tatsuro MATSUMOTO

perieur de l'Algerie. Mem. Soc. Ceol.
France, Paleont., Mem. no. 17, vol. 6, fasc.
4, pp. 1-24, pis. 14-19 [1-6J (1896); vol.
7, fasc. 1-2, pp. 25-88, pis. 1-12 [7-18J(1897) .
PERVINQUIERE, L. (1907) : Etudes de paleontologie tunisienne. I. Cephalopodes desTerrains secondaires. Carte geol. Tunisie,
483 pp., 27 pis.
REESIDE, J.B., Jr. (1923): A new fauna fromthe Colorado Group of southern Montana.
U.S. Ceol. Surv. Prof. Paper 132-B, pp.
25-33, pis. 11-21.
REYMENT, R.A. (1954) : Some new Upper Cretaceous ammonites from Nigeria. Colonial Ceol. Min. Resources, vol. 4, no. 3,
pp. 248-270, pis. 1-5.
- - (1955): The Cretaceous Ammonoidea ofsouthern Nigeria and the southern Cameroons. Ceol. Surv. Nigeria, Bull. no. 25,
112 pp., 24 pis.
REYMENT, R.A. and TAIT, E.A. (1972): Faunal evidence for the orgin of the South
Atlantic. 24th Intern. Ceol. Congr., sect.
7, pp. 316-323, Montreal.
SASTRY, M.V.A. and MATSUMOTO, Tatsuro(1967): Notes on some Cretaceous ammonites from southern India. Part II.
Mem. Fac. Sci., Kyushu Univ., ser. D,
vol. 18, no. 1, pp. 1-5, pis. 1.
SCHNEEGANS, D. (1943): Invertebres du Cretace superieur du Damergou (Territoire
du Niger). Bull. Direc. Min. Couvern.
Cener. Afri. Ocrid. Franc., no. 7, pp. 87166, 8 pis.

SPATH, L.F. (1926): On new ammonites fromthe English Chalk. Ceol. Mag., vol. 63,
pp. 77-83.
STOLICZKA, Ferdinand (1863-66): Ammonitidae, with revision of the Nautilidae. etc.
In BLANFORD, M.F., and STOLICZKA, F.,
1861-66. The fossil Cephalopoda of theCretaceous rocks of southern India. Mem.
Ceol. Surv. India, Palaeont. Indica, vol.
1, pp. 41-216,pls. 26-94.
TANAKA, Keisaku (1963): A study on the
Cretaceous sedimentation in Hokkaido,
Japan. Ceol. Surv. Japan, Rept. no. 197,
119 pp.
T AUBENHAUS, H. (1920): Die Ammoneen der
Kreideforma tion Palastinae und Syrien.
Zeit. deutsch. Paliistina- Vereins, vol. 43,
58 pp., 9 pis.
WIEDMANN, Jost (1960) : Le cretace superieur
de l'Espagne et du Portugal et ses cephalopodes. 84 8 Congres Soc. savantes, 1959,
pp. 706-764.
- - (1964): Le cretace superieur de l'Espagne et du Portugal et ses cephalopodes.
Estudios Ceologicos, Inst. Lucas Mallada,
C.S.I.C. (Espana). vol. 20, pp. 107-148.
WRIGHT, C.W. (1957): In MOORE, R.C. [EditorJ : Treatise on Invertebrate Paleontology, Part L, Mollusca, Cephalopoda, Ammonoidea, p. Ll-L490, Geol. Soc. Amer.,& Univ. Kansas Press.
YABE, Hisakatsu (1904): Cretaceous Cephalopoda from the Hokkaido. Part II. Jour.
Coll. Sci., Imp. Univ. Tokyo, vol. 20, art.
2, pp. 1-45, pis. 1-6.

Explanation of Plate 8
Fig. 1. Vascoceras sp. aff. V. durandi (THOMAS and PERON) ........................ Page 29
GK. H5674, from loco R2513a, Member Mh of the Obira area. Lateral (a) and rear (b)views, x 0.9.
Fig. 2. Fagesia thevestensis (PERON) .................................................. Page 32
Inner whorl of GT. 1-3249, from loco T610 b-c, unit II b of the Saku area. Lateral (a),rear (b) and frontal (c) views, xO.9.
Kyushu University [I. HAYAMIJ photos, without whitening for Fig. 1 and with whitening for
Fig. 2.

MATSUMOTO:

Vascoceratid ammonites

Plate 8

614.
Abeshinai
Horokanai
Ishikari
Nakagawa-machi
N akakinebetsu
Obira
Okufutamata
Onodera-zawa

Y"':'y-j-1

~l

1m

I*J

~;t
T:i
if! )11 IllT
if! $2 ~ JJIJ
-'f
IJ'

~
Ij' !llt 1f

Ill!:
iR

Vascoceratid ammonites['tC-'f:trkJ]

41

Opirashibets:;t l:::" '7 y"':' 'Y [/J'-'f~]
Rumoi
Jlij
fl1
Saku
t!i:.
!A
Saku-gakko-no-sawa {z'c.!A,**Ji:O)iR
Saku-gawa{z'c.
!A ) II
Shakotan-ga wa{z'c. J!l H )11
Teshiojj
X.
Yubari, The-Y ~R JI I

42

PROCEEDINGS OF THE PALAEONTOLOGICAL SOCIETY
OF JAPAN

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Some Lower Cretaceous plants newly foundfrom the Ryoseki Group, Kochi Prefecture, Japan ... KIMURA, T. & M. HIRATA
Older Mesozoic plants from the Tae-Dong
Series, Korea. Pt. 1 ................. .
. .. .. .. .. .. .. .. . KIMURA, T. & B.-K. KIM
Phylogenetic system of vascular plants ..
.............................. ASAMA, K.

i'PIiUIfI< IIII~I:!Ik1.i(J) i'i J I=l}\lfl~ M:J-:" fE~f.r~l<::'"?'-"L ............................ I],JRttl-

M**~.~mMiliKft£~~~~*(J)~._

f\::1<::'-?1r'L ........................ +JR *tILower Carboniferous conodonts from Mitsusawa, north of Itsukaichi, Tokyo ..... .
................ IGo, H. & F. KOBAYASHI
Geologic age of the Mamba Group. Kanto
Massif. Pt. 1. Corals and fusulinaceans.............. .... IGo, H. & S. TAKIZAWA
Ditto. Pt. 2. Conodonts ............... .
................ TAKIZAWA, S. & H. IGO

Upper Permian fusulinaceans from Kuzuu,
Tochigi Prefecture, Japan .... H. & H. IGO
Planktonic foraminiferal sequence in apiston core from off Sanin district, Japan
Sea ............ UJIIE, H. & M. ICHIKURADistribution of the Japanese Miogypsina,
with a description of a new species................................ UJIIE, H.
~2~~R~t1:l~G~km.~.~~~U(J)mt'LI~"?'-"L ..................... Jjt,m::tfi!r.l1ll!:=r:~i!f.1:lEk T'/ J 7"7 '/!1 r '/1t1:lI<::'''?'-''

L(Tf,~).............. ~l.lJI~Hg·~I:ID. i$

Vascoceratid ammonites from the Turonianof Hokkaido ............ MATSUMOTO, T.
Evolution of Lepidolina multiseptata (Permian foraminifer) in East Asia ......... .
....... ...... ...... ...... ..... OZAWA, T.
Genetic polymorphism and geographic variation in Umbonium (Preliminary observation) ...................... OZAWA, T.
f:kFl3W,='/;j:H1;l1:(J)*~· hii'i=,!i'lFjJ!ffIJf.r..... .
.................................. ~j'W§ififrij

Growth of massive corals ...... HAMADA, T.
"Cladochonus" from the red bed of Malaya(Malaysia) .................. HAMADA, T.
On some fossil Scleractinia from Japan. ............................. EGUCHI, M .
Japanese Tertiary cassidids and their related species of the west coast of North
America ...................... KANNO, S.
"Pecten (Amusium) ocala nus Dall" and some
pectinid with internal radial ribs ..... .
............................... KANNO, S.
IJ'~J.ij( • flJ:,liMilfJfMf.r?r~(J) ";I.;;t r ~ J 71'1" 1<::.
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··································*ttfIjUffi

1=~i/]i~l'iiU1&rl~(J)Faunal analysis of the Upper Miaoli (Byoritsu) Group in the Tunghsiao area,43
Miaoli Province, Formosa, China ..... .
.............................. KOTAKA, T.
Proposed correlation of Neogene sediments
in Noto and Oga Peninsulas, Japan,
using planktonic diatoms (t-t1m) ....... .
.......................... BURCKLE, L. H.

Late Cenozoic planktonic Foraminiferafrom the oil-bearing formations of Japan(t-t~il) .. MAIYA, S., T. SAITO, & T. SAT a
Planktonic foraminifers of the Nishikurosawa Formation, northwestern Honshu,
Japan (f-tRjll) ...... SAITO, T. & S. MAIYA

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Transactions and Proceedings of the PalaeontologicalSociety of Japan

April 20, 1973

New Series No. 89

CONTENTS
TRANSACTIONS
611.

612.

613.

HIRANO, Hiromichi: Biostratigraphic study of the Jurassic Toyora Group.
Part II .. . . .... .................... . ........................... . . .. ...

1

NISHIMIYA, Katsuhiko and Y AMAGIWA, Nobuo: Coral fossils from the
Kosode Formation, Yamanashi Prefecture.......................... . .

15

HUZIOKA, Kazuo and T AKAHASI, Eitaro: A Triassic Hepatica from the
Omine coal-field, Southwest Honshu, Japan ... ... ... . ....... .. ...... . 24

614. MATSUMOTO, Tatsuro: Vascoceratid ammonites from the Turonian of
Hokkaido (Studies for the Cretaceous ammonites from Hokkaido andSaghalien-XXVI) .. ..... .. . . ........... .. ... . .... . . ........ .. . . . . . ...

27

PRODCEEDINGS . . ......... ..... . .. ...................... . . .. ...... .. . ... .....

42