Deep-sea Fauna of the Sea of Japan, edited by T. Fujita, National Museum of Nature and Science Monographs, No. 44,
pp. 225–291, 2014

Annotated Checklist of Deep-sea Fishes of the Sea of Japan
Gento Shinohara1, Masanori Nakae1, Yuji Ueda2,
Shigeaki Kojima3 and Keiichi Matsuura1
1

Department of Zoology, National Museum of Nature and Science,
4–1–1 Amakubo, Tsukuba-shi, Ibaraki, 305–0005 Japan
E-mail: s-gento@kahaku.go.jp
2
Japan Sea National Fisheries Research Institute,
1–5939–22 Suido-cho, Chuou-ku, Niigata-shi, Niigata, 951–8121 Japan
3
Graduate School of Frontier Sciences, the University of Tokyo,
1–5–1 Kashiwanoha, Kashiwa-shi, Chiba, 277–8563 Japan

Abstract: A list of deep-sea ﬁshes from the Sea of Japan is presented on the basis of a literature
survey and specimens mainly collected in the years 2009–2013. A total of 363 species belonging to
104 families of 29 orders of deep-sea ﬁshes is listed with remarks on the literature surveys and/or
specimens.
Key words:

deep-water ﬁshes, ﬁsh fauna, distribution, Sea of Japan, voucher specimens.

Introduction
The Sea of Japan is one of the major marginal seas in the western North Paciﬁc Ocean,which is characterized by having a deep-sea ﬂoor, extending to 3,800 m depth. The sea is semienclosed, connecting to the adjacent major seas via the Tatar (10 m depth), Soya (60 m depth),
Tsugaru (140 m depth), Tsushima (120 m depth) and Korea straits (140 m depth) and also the very
narrow Kanmon Strait (46 m depth) between Kyushu and Honshu to the Seto Inland Sea (Fig. 1).
In addition, it has unique hydrographic features. The surface layer is affected by the warm Tsushima Current, a branch of the Kuroshio Current, from the south and the cold Liman Current
from the north. In depths below 200–300 m, temperatures year-around remain at 1ºC or less andare much colder than other seas around Japan and the adjacent areas (Nishimura, 1974: table 14).
This cold deep water mass (0.1–0.3ºC) with low salinity (34.0–34.1‰) and high dissolved oxygen content (5–6 mL/L) is called the Japan Sea Proper Water, which originates in the northwestern part of the sea and results from extensive cooling of the surface water during winter (Oba etal., 1991).
The Sea of Japan is the newest marginal sea around Japan. Geological studies of the Sea ofJapan revealed that the origin of the sea was a rift-valley on the eastern edge of the Asian Continent around 250 million years ago with the sea ﬂoor expanding until 15 million years ago (e.g.,
Taira, 1990). A number of environmental changes took place in the Sea of Japan during the last
85,000 years (Oba et al., 1991: ﬁg. 7). For instance, during the last glacial period (27,000–17,000
years ago), the sea was completely isolated from its neighboring seas by a lowering of the sea
level. The surface layer of the sea was also covered by a massive inﬂux of freshwater from the
Asian Continent; the development of stratiﬁcation caused severe anoxic conditions that extir-

226

Gento Shinohara, Masanori Nakae, Yuji Ueda, Shigeaki Kojima and Keiichi Matsuura

pated many deep-sea organisms (e.g., Tada et al., 1999), except for small fractions of their populations (Gorbarenko and Southon, 2000; Itaki et al., 2004).
The environmental changes and geological history in the Sea of Japan have created a uniquedeep-sea ﬁsh fauna. It is characterized by both low species diversity and relatively few endemic
species (Okiyama, 1974). The low species diversity may reﬂect the short period of time for colonization (Tyler, 2002). In addition, there are only a few oceanic deep-sea ﬁshes recorded from thesea (Nishimura, 1964; Okiyama, 1974; Honma, 1987). The endemic ﬁshes comprise coldadapted eurybathic species. Nishimura (1974: table 3) indicated 7 endemic deep-sea species/subspecies inhabited the Sea of Japan with their sister groups residing in the Sea of Okhotsk and theBering Sea.
There are several studies on ﬁsh faunas in local areas associated with or in the Sea of Japan,
e.g., Katayama (1940) for Toyama Bay, Mori (1956a) for the Oki Islands and Honma and Kitami(1967, 1970, 1980, 1995) for Sado Island. Russian researchers including G. U. Lindberg and hiscolleagues published a series of books on the ﬁsh fauna (e.g., Lindberg and Legeza, 1959, 1965;
Lindberg, and Krasyukova, 1969, 1975). Recently, Shinohara et al. (2011) reported 160 species
belonging to 18 families of 21 orders of deep-sea ﬁshes from the Sea of Japan, based on thevoucher specimens deposited in the following universities and museums: Hokkaido University
Museum, Hakodate; Kochi University, Kochi; Kyoto University, Maizuru; Osaka Museum of
Natural History, Osaka; National Museum of Nature and Science, Tsukuba; U.S. National
Museum of Natural History, Washington D.C.; Zoological Institute of Russian Academy of Sciences, Saint Petersburg.
In the following pages, a total of 363 species belonging to 104 families of 29 orders of the
deep-sea ﬁshes are listed with remarks on the basis of literature surveys and/or specimens mainlycollected in the years 2009–2013.
Materials and Methods
Collection data of specimens captured by the R/V Soyo-maru of the Fisheries Research
Agency (FRA), R/V Tansei-maru of the Japan Agency for Marine-Earth Science and Technology(JAMSTEC) and T/V Tanshu-maru of Kasumi High School are shown in Fig. 1 and Table 1. The
specimens are deposited in the National Museum of Nature and Science, Tsukuba (NSMT-P) andKyoto University (FAKU). Specimens deposited in the Sado Marine Experimental Station,
Niigata University (SMBS) were also examined (Fig. 2). Numbers of specimens are indicated inparentheses following station numbers and/or SMBS numbers.
Systematic arrangement of taxa follows Nelson (2006). Author(s) and year of each scientiﬁcname follows the Catalog of Fishes on the website of the California Academy of Sciences, USA(http://research.calacademy.org/redirect?url=http://researcharchive.calacademy.org/research/Ichthyology/catalog/ﬁshcatmain.asp).
We examined the following publications including deep-sea ﬁshes occurring below 200 m
depth; Keys to the Fishes and Fish-like Animals of Japan (Okada and Matsubara, 1938); Fish
Morphology and Hierarchy (Matsubara, 1955); The Fishes of the Japanese Archipelago, 2nd Edition (Masuda et al., 1988); The Fishes of Northern Japan (Amaoka et al., 1995); Fishes of Japan
with Pictorial Keys to the Species (Nakabo, 1993a, 2000a, 2002a, 2013a). In addition to thesemajor literature resources, revisional papers for taxonomic groups were examined to obtain
information on distribution of deep-sea ﬁshes in the Sea of Japan (e.g., Toyoshima, 1985; Kido,
1988a; Kanayama, 1991). In our literature survey, we recorded localities given with signiﬁcantAnnotated Checklist of Deep-sea Fishes of the Sea of Japan

227

Fig. 1. Study areas and collection localities in the Sea of Japan. Star, stations of R/V Soyo-maru; squares, R/V
Tansei-maru; circles, T/V Tanshu-maru.

accuracy (e.g., Sanʼin District, Toyama Bay) but excluded general locality records such as“around Japan.” In our research the Sea of Japan is delineated by the Tatar Strait in north and theTsushima and Korean straits in south.

Gento Shinohara, Masanori Nakae, Yuji Ueda, Shigeaki Kojima and Keiichi Matsuura

228

Table 1. Trawl and baited trap stations of the R/Vs Soyo-maru and Tansei-maru and the T/S Tanshu-maru in 2009–2013. BT, beam trawl; OT, otter trawl.
Station no.

Date

Gear

Position in

Position out

Depth (m)R/V Soyo-maru
SO10-C4-B4

19 July 2010

BT

39°59.20′N, 135°57.80′E

39°58.60′N, 135°57.10′E

1371–1371

R/V Tansei-maru
KT-10-08-R1
KT-10-08-R2
KT-10-08-T1
KT-11-09-E2
KT-11-09-K4
KT-11-09-M1
KT-11-09-M5
KT-11-09-M6
KT-11-09-N1
KT-11-09-N2
KT-11-09-N3
KT-11-09-N4
KT-11-09-T1
KT-11-09-T2
KT-11-09-T3
KT-11-09-T5
KT-11-09-T6

23 May 2010
23 May 2010
21 May 2010
29 May 2011
2 June 2011
27 May 2011
28 May 2011
28 May 2011
1 June 2011
1 June 2011
1 June 2011
1 June 2011
31 May 2011
31 May 2011
31 May 2011
1 June 2011
1 June 2011

BT
BT
BT
BT
BT
BT
BT
BT
BT
BT
BT
BT
BT
BT
BT
BT
BT

45°36.25′N, 140°54.73′E
45°27.25′N, 140°20.90′E
40°53.90′N, 140°00.04′E
41°50.26′N, 139°34.03′E
35°59.92′N, 134°20.58′E
44°43.78′N, 140°05.79′E
44°31.41′N, 139°47.13′E
44°16.68′N, 139°36.85′E
37°54.59′N, 136°56.92′E
37°57.06′N, 136°56.60′E
38°03.25′N, 136°53.39′E
38°08.04′N, 136°49.64′E
37°28.45′N, 137°28.15′E
37°28.77′N, 137°29.07′E
37°29.18′N, 137°33.25′E
37°18.45′N, 137°31.77′E
37°28.29′N, 137°45.89′E

45°35.66′N, 140°54.09′E
45°26.15′N, 140°21.94′E
40°53.28′N, 139°58.62′E
41°48.70′N, 139°34.18′E
35°59.61′N, 134°18.82′E
44°43.12′N, 140°05.44′E
44°29.91′N, 139°47.96′E
44°14.25′N, 139°37.25′E
37°54.51′N, 136°58.43′E
37°57.04′N, 136°58.23′E
38°03.82′N, 136°55.13′E
38°09.00′N, 136°51.20′E
37°29.16′N, 137°27.86′E
37°29.53′N, 137°28.63′E
37°29.35′N, 137°31.96′E
37°18.66′N, 137°32.85′E
37°26.70′N, 137°44.59′E

178–190402–393
350–315
388–538
611–613152–166
1035–1080
1461–1421
162–157
203–203403–414617–604
160–173
207–258
383–443794–7901488–1564

T/V Tanshu-maru
TS09-T001
TS09-T002
TS09-T004
TS09-T006
TS09-T007
TS09-T008
TS09-T010
TS09-T011
TS09-T012
TS09-T013
TS09-T016
TS09-T017
TS09-T018
TS09-T020
TS09-T023
TS09-T025
TS09-T027
TS09-T029
TS09-T032
TS09-T033
TS09-T034
TS09-T035
TS09-T036
TS09-T037
TS09-T038
TS09-T039
TS09-T047
TS09-T049
TS09-T050
TS09-T053

9 May 2009
10 May 2009
10 May 2009
9 May 2009
10 May 2009
11 May 2009
12 May 2009
9 May 2009
11 May 2009
9 May 2009
12 May 2009
12 May 2009
12 May 2009
13 May 2009
13 May 2009
13 May 2009
13 May 2009
12 May 2009
11 May 2009
11 May 2009
14 May 2009
14 May 2009
15 May 2009
15 May 200916 May 200916 May 2009
1 June 2009
2 June 2009
2 June 2009
14 May 2009

OT
OT
OT
OT
OT
OT
OT
OT
OT
OT
OT
OT
OT
OT
OT
OT
OT
OT
OT
OT
OT
OT
OT
OT
OT
OT
OT
OT
OT
OT

35°43.56′N, 131°06.41′E
35°49.86′N, 131°28.63′E
35°51.03′N, 131°34.58′E
35°53.59′N, 131°28.46′E
35°53.46′N, 131°33.78′E
35°37.75′N, 132°04.41′E
35°46.53′N, 132°02.23′E
35°27.85′N, 130°38.87′E
35°43.46′N, 132°05.83′E
35°34.34′N, 130°53.43′E
35°46.11′N, 132°13.33′E
35°39.83′N, 132°16.68′E
35°43.21′N, 132°17.38′E
35°45.09′N, 132°24.13′E
35°50.38′N, 132°23.92›E
35°49.09′N, 132°29.16′E
35°54.57′N, 132°26.46′E
35°48.05′N, 132°09.98′E
35°53.68′N, 131°54.22′E
35°51.91′N, 132°08.59′E
35°53.75′N, 132°32.75′E
36°01.72′N, 132°38.20′E
36°05.85′N, 132°40.07′E
36°08.98′N, 132°45.82′E
36°17.98′N, 132°43.84′E
36°19.53′N, 132°47.79′E
36°08.90′N, 133°46.83′E
36°09.18′N, 133°54.64′E
36°00.76′N, 133°54.48′E
36°00.39′N, 132°27.55′E

35°42.96′N, 131°04.81′E
35°50.10′N, 131°26.81′E
35°51.09′N, 131°34.19′E
35°53.50′N, 131°26.62′E
35°53.32′N, 131°34.55′E
35°37.79′N, 132°06.24′E
35°46.28′N, 132°04.04′E
35°28.90′N, 130°40.18′E
35°43.34′N, 132°07.66′E
35°34.21′N, 130°51.75′E
35°46.42′N, 132°15.13′E
35°40.62′N, 132°18.22′E
35°44.03′N, 132°18.92′E
35°46.19′N, 132°25.36′E
35°50.70′N, 132°24.22′E
35°50.23′N, 132°31.15′E
35°55.88′N, 132°27.32′E
35°48.13′N, 132°11.82′E
35°53.63′N, 131°52.99′E
35°51.75′N, 132°10.42′E
35°55.16′N, 132°33.35′E
36°00.47′N, 132°37.37′E
36°04.82′N, 132°38.73′E
36°07.92′N, 132°44.52′E
36°19.45′N, 132°44.22′E
36°21.00′N, 132°48.10′E
36°07.75′N, 133°47.49′E
36°07.75′N, 133°55.16′E
35°59.29′N, 133°54.81′E
36°01.37′N, 132°28.49′E

227–224
248–248
280–279
342–329
339–339
208–208281–281
208–211251–251
225–223
280–280
209–211234–236
215–215
272–270
230–230
280–280
321–321417–408
441–439
210–210
210–211
230–234
206–205
269–269
219–218
207–207247–247
231–231349–350

Annotated Checklist of Deep-sea Fishes of the Sea of Japan
Table 1.

229

(continued).

Station no.
TS09-T054
TS09-T056
TS09-T057
TS09-T086
TS09-T087
TS09-T088
TS09-T093
TS09-T099
TS09-T101
TS09-T102
TS09-T105
TS09-T106
TS09-T107
TS09-T110
TS09-T115
TS09-T117
TS09-T141
TS10–OW01
TS10-OW04
TS10-OW09
TS10-OW10
TS10-OW12TS10-OW13
TS10-OW18
TS10-YA05

Date

Gear

Position in

Position out

Depth (m)15 May 2009
1 June 2009
15 May 2009
3 June 2009
3 June 2009
3 June 2009
3 June 2009
5 June 2009
31 May 2009
30 May 2009
31 May 2009
30 May 2009
30 May 2009
6 June 2009
6 June 20097 June 200914 June 2009
21 Aug. 2010
21 Aug. 2010
28 Aug. 2010
28 Aug. 2010
27 Aug. 2010
27 Aug. 2010
26 Aug. 2010
29 Aug. 2010

OT
OT
OT
OT
OT
OT
OT
OT
OT
OT
OT
OT
OT
OT
OT
OT
OT
BT
BT
BT
BT
BT
BT
BT
BT

36°19.75′N, 132°42.06′E
36°17.05′N, 133°53.20′E
36°16.94′N, 132°38.50′E
35°56.27′N, 134°33.87′E
35°51.52′N, 134°43.40′E
35°49.20′N, 134°51.89′E
35°58.52′N, 134°27.70′E
36°02.18′N, 135°12.16′E
35°58.93′N, 135°30.16′E
35°53.27′N, 135°48.27′E
36°06.43′N, 135°43.39′E
36°01.78′N, 135°42.77′E
36°09.30′N, 135°54.87′E
36°10.82′N, 135°25.63′E
36°15.29′N, 135°30.95′E
36°32.51′N, 136°08.82′E
36°00.52′N, 132°33.20′E
35°40.31′N, 132°20.29′E
35°53.21′N, 132°13.76′E
35°59.00′N, 132°12.55′E
36°01.13′N, 132°13.71′E
36°09.98′N, 132°11.67′E
36°13.23′N, 132°08.69′E
36°36.78′N, 131°39.63′E
39°40.16′N, 136°01.81′E

36°21.23′N, 132°42.38′E
36°15.74′N, 133°53.96′E
36°18.42′N, 132°38.77′E
35°56.75′N, 134°35.41′E
35°51.67′N, 134°45.23′E
35°49.25′N, 134°52.56′E
35°58.98′N, 134°29.47′E
36°02.19′N, 135°14.01′E
35°58.24′N, 135°31.82′E
35°53.02′N, 135°47.72′E
36°05.13′N, 135°42.46′E
36°01.22′N, 135°44.51′E
36°08.43′N, 135°54.91′E
36°11.33′N, 135°23.88′E
36°15.67′N, 135°29.16′E
36°33.73′N, 136°09.53′E
36°01.75′N, 132°34.25′E
35°40.48′N, 132°20.66′E
35°53.29′N, 132°14.26′E
35°59.15′N, 132°13.20′E
36°01.06′N, 132°14.28′E
36°10.26′N, 132°12.21′E
36°13.20′N, 132°09.29′E
36°37.18′N, 131°40.07′E
39°40.57′N, 136°01.44′E

318–322358–352
441–434
275–274
225–225
200–200
328–330
283–284
251–250
242–242292–291
272–271
202–201
351–351
451–452
230–230
250–251
201–201504–502
1000–1011
1104–1104
1296–1300
1408–1394
1895–1894
1002–1010

Asterisks ahead of scientiﬁc names indicate records found in the publications with novoucher specimens. The species records found in the following publications are supported bytheir voucher specimens: Balushkin and Prirodina, 2008 (deposited in Zoological Institute, SaintPetersburg: ZIN); Balushkin et al., 2011 (ZIN); Kawase and Kai, 2012 (Natural History Museum
and Institute, Chiba); Matsuura et al., 2009 (National Museum of Nature and Science, Tsukuba:
NSMT-P); Nambu and Kido, 1990; Nambu et al., 1991, 1992, 1993 (Toyama Science Museum);
Sheiko and Fedorov, 2010 (ZIN); Sideleva et al., 2006a, b (ZIN); Shinohara et al., 2011 (seeIntroduction); Suzuki et al., 2000 (Osaka Museum of Natural History, Osaka); Uozu Aquarium,
1997 (Uozu Aquarium, Uozu); Voronina and Volkova, 2003 (ZIN).
Species List
Order Myxiniformes
Family Myxinidae
Eptatretus atami (Dean, 1904) [Japanese name: Kuro-nuta-unagi]
Paramyxine atami: Mori, 1952; Matsubara, 1955; Kato, 1956; Mori, 1956a; Lindberg andLegeza, 1959; Honma, 1962; Takegawa and Morino, 1970; Tsuda, 1990; Uozu Aquarium, 1997.
Eptatretus atami: Nakabo and Kai, 2013.
Eptatretus burgeri (Girard, 1855) [Japanese name: Nuta-unagi]
Eptatretus burgeri: Mori, 1928; Mori and Uchida, 1934; Okada and Matsubara, 1938; Kato,
1956; Mori, 1956a; Takegawa and Morino, 1970; Tsuda, 1990; Uozu Aquarium, 1997; Suzuki etal., 2000; Nakabo and Kai, 2013.

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Gento Shinohara, Masanori Nakae, Yuji Ueda, Shigeaki Kojima and Keiichi Matsuura

Fig. 2. Fishes deposited in the Sado Marine Biological Station, Niigata University (SMBS). A–B, dorsal andventral views of Bathyraja bergi, 217 mm TL, off Tassha, Sado Island, 13 May 1959; C, Beryx decadactylus,
185 mm SL, Ryotsu Bay, Sado Island, 4 February 1988; D, Scorpaena neglecta, 204 mm SL, off Tassha, Sado
Island; E, Icelus toyamensis, 88 mm SL, Hyōtan-se Shallow, off Sado Island, 20 September 1957; F, Cookeolus japonicus, 148–198 mm SL, off Tassha, Sado Island, 13 June 1954; G, Pterycombus petersii, 200 mm SL,
Tassha, Sado Island, 10 April 1989; H, Histiopterus typus, 74 mm SL, off Himetsu, Sado Island, 4 March
1991; I, a glass jar and Plectranthias kelloggi azumanus, 92 mm SL (2 specimens), off Aikawa, Sado Island,
23 September 1994.

Eptatretus bürgeri: Katayama, 1940.
Eptatretus okinoseanus (Dean, 1904) [Japanese name: Murasaki-nuta-unagi]
Eptatretus okinoseanus: Katayama, 1940; Tsuda, 1990; Shinohara et al., 2011; Nakabo andAnnotated Checklist of Deep-sea Fishes of the Sea of Japan

231

Kai, 2013.
Order Chimaeriformes
Family Chimaeridae
Chimaera phantasma Jordan and Snyder, 1900 [Japanese name: Ginzame]
Chimaera phantasma: Mori and Uchida, 1934; Honma, 1952; Mori, 1952; Kato, 1956; Mori,
1956a; Lindberg and Legeza, 1959; Shinohara et al., 2011; Honma, 2013; Nakabo et al., 2013a.
*Hydrolagus barbouri (Garman, 1908) [Japanese name: Kokonohoshi-ginzame]
Chimaera barbouri: Honma, 1952.
Bathyalopex barbouri: Kato, 1956.
Hydorolagus barbouri: Lindberg and Legeza, 1959; Honma, 2013; Nakabo et al., 2013a.
*Hydrolagus mitsukurii (Jordan and Snyder, 1904) [Japanese name: Aka-ginzame]
Chimaera mitsukurii: Honma, 1952.
Phasnichthys mitskurii [sic]: Kato, 1956.
Hydrolagus mitsukurii: Lindberg and Legeza, 1959; Honma, 2013; Nakabo et al., 2013a.
*Hydrolagus ogilbyi (Waite, 1898) [Japanese name: Ginzame-damashi]
Hydrolagus ogilbyi: Kato, 1956.
Order Lamniformes
Family Mitsukurinidae*Mitsukurina owstoni Jordan, 1898 [Japanese name: Mitsukurizame]
Mitsukurina owstoni: Katayama, 1940; Aonuma et al., 2013.
Scapanorhynchus owstoni: Kato, 1956; Tsuda, 1990.
Family Lamnidae
Isurus oxyrinchus Rainesque, 1810 [Japanese name: Aozame]
Isuropsis glauca: Mori and Uchida, 1934; Honma, 1952.
Isrus glauca: Mori, 1952.
Isurus glaucus: Kato, 1956; Mori, 1956a; Honma, 1956b; Lindberg and Legeza, 1959;
Honma, 1963; Tsuda, 1990.
Isurus oxyrinchus: Honma and Kitami, 1978; Uozu Aquarium, 1997; Sokolovskaya et al.,
1998; Honma et al., 2011; Honma, 2013; Yoshino et al., 2013a.
Lamna ditropis Hubbs and Follett, 1947 [Japanese name: Nezumizame]
Lamna cornubica: Katayama, 1940; Honma, 1952, 1963.
Lamna ditropis: Kato, 1956; Lindberg and Legeza, 1959; Ueno, 1971; Honma and Kitami,
1978; Honma, 1991a; Uozu Aquarium, 1997; Adrianov and Kussakin, 1998; Sokolovskaya et al.,
1998; Maeda and Tsutsui, 2003; Kitagawa et al., 2008; Honma, 2013; Yoshino et al., 2013a.
Lamma ditropis [sic]: Tsuda, 1990.
Family Alopiidae*Alopias pelagicus Nakamura, 1935 [Japanese name: Nitari]
Alopias pelagicus: Honma, 1952; Mori, 1952; Kato, 1956; Lindberg and Legeza, 1959;
Honma, 1963; Takegawa and Morino, 1970; Honma and Kitami, 1978; Kobayashi et al., 2006;
Honma, 2013.

232

Gento Shinohara, Masanori Nakae, Yuji Ueda, Shigeaki Kojima and Keiichi Matsuura

Alopias vulpes: Mori and Uchida, 1934.
Alopias vulpinus: Ueno, 1971.
*Alopias vulpinus (Bonnaterre, 1788) [Japanese name: Ma-onaga]
Alopias unlpinus [sic]: Tsuda, 1990.
Alopias vulpinus: Maeda and Tsusui, 2003; Honma, 2013; Yoshino et al., 2013b.
Order Carcharhiniformes
Family Scyliorhinidae
Cephaloscyllium umbratile Jordan and Fowler, 1903 [Japanese name: Nanukazame]
Cephaloscyllium umbratile: Mori and Uchida, 1934; Honma, 1952; Mori, 1952; Kato, 1956;
Mori, 1956a; Lindberg and Legeza, 1959; Nakaya, 1975; Tsuda, 1990; Honma, 1991a; Uozu
Aquarium, 1997; Shinohara et al., 2011; Yoshino et al., 2013a.
Cephaloscyllium isbaellum [sic]: Suzuki et al., 2000.
*Galeus eastmani (Jordan and Snyder, 1904) [Japanese name: Yamorizame]
Galeus eastmani: Kato, 1956; Tsuda, 1990.
Halaelurus buergeri (Müller and Henle, 1838) [Japanese name: Nagasaki-torazame]
Halaelurus bürgeri: Nakaya, 1975.
Halaelurus buergeri: Maeda and Tsutsui, 2003; Yoshino et al., 2013a.
Scyliorhinus torazame (Tanaka, 1908) [Japanese name: Torazame]
Catulus torazame: Mori and Uchida, 1934; Okada and Matsubara, 1938; Katayama, 1940;
Honma, 1952.
Scylliorhinus torazame [sic]: Mori, 1952.
Scyliorhinus torazame: Honma, 1955b; Kato, 1956; Mori, 1956a; Lindberg and Legeza,
1959; Honma, 1963; Ueno, 1971; Nakaya, 1975; Honma, 1977; Honma and Kitami, 1978;
Tsuda, 1990; Yoshino and Aonuma, 2000, 2002; Maeda and Tsutsui, 2003; Shinohara et al.,2011; Honma, 2013; Yoshino et al., 2013a.
Scylirhinus torazame [sic]: Suzuki et al., 2000.
SMBS uncatalogued (1).
Family Triakididae*Mustelus griseus Pietschmann, 1908 [Japanese name: Shirozame]
Cynias griseus: Mori and Uchida, 1934; Mori, 1952.
Mustelus griseus: Honma, 1955c; Honma, 1956b; Kato, 1956; Lindberg and Legeza, 1959;
Honma, 1963; Tsuda, 1990; Aonuma et al., 2013; Honma, 2013.
Mustelus manazo Bleeker, 1854 [Japanese name: Hoshizame]
Cynias manazo: Mori, 1928; Mori and Uchida, 1934; Mori, 1952.
Mustelus manazo: Snyder, 1912; Honma, 1952; Mori, 1952; Honma, 1955b; Kato, 1956;
Mori, 1956a; Lindberg and Legeza, 1959; Honma, 1963; Takegawa and Morino, 1970; Honma,
1977; Honma and Kitami, 1978; Tsuda, 1990; Uozu Aquarium, 1997; Adrianov and Kussakin,1998; Sokolovskaya et al., 1998; Suzuki et al., 2000; Maeda and Tsutsui, 2003; Kitagawa et al.,2008; Matsuura et al., 2009; Aonuma et al., 2013; Honma, 2013.
Family Carcarhinidae*Carcharhinus plumbeus (Nardo, 1827) [Japanese name: Mejirozame]

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Carcharhinus gangeticus: Kato, 1956; Ueno, 1971; Tsuda, 1990.
Carcharhinus plumbeus: Maeda and Tsutsui, 2003; Kobayashi et al., 2006; Aonuma et al.,2013; Honma, 2013.
*Rhizopriodon acutus (Rüppell, 1837) [Japanese name: Hiragashira]
Scoliodon walbeehmi: Kato, 1956.
Rhizopriodon acutus: Honma et al., 1990; Tsuda, 1990; Honma, 1991a; Aonuma et al., 2013;
Honma, 2013.
Order Hexanchiformes
Family Hexanchinidae
Heptranchias perlo (Bonnaterre, 1788) [Japanese name: Edo-aburazame]
Heptranchias perlo: Honma, 1952; Mori, 1952; Honma, 1954; Kato, 1956; Mori, 1956a;
Lindberg and Legeza, 1959; Honma, 1963; Honma and Kitami, 1978; Tsuda, 1990; Shinohara etal., 2011; Hatooka et al., 2013a; Honma, 2013.
*Hexanchus griseus (Bonnaterre, 1788) [Japanese name: Kagurazame]
Hexanchus griseus: Tsuda, 1990; Honma et al., 2011, 2012; Honma, 2013.
Order Squaliformes
Family Somniosidae*Somniosus (Somniosus) paciﬁcus Bigelow and Schroeder, 1944 [Japanese name: Ondenzame]
Somniosus paciﬁcus: Ueno, 1971; Maeda and Tsutsui, 2003.
Somniosus (Somniosus) paciﬁcus: Hatooka et al., 2013a.
Family Squalidae
Squalus acanthias Linnaeus, 1758 [Japanese name: Abura-tsunozame]
Squalus suckleyi: Mori and Uchida, 1934; Katayama, 1940; Honma, 1952; Katayama, 1952;
Ouchi, 1963.
Squalus (Acanthias) suckeryi: Mori, 1952.
Squalus acanthias: Matsubara, 1955; Honma, 1956b; Kato, 1956; Mori, 1956a; Honma,
1963; Ueno, 1971; Honma and Kitami, 1978; Tsuda, 1990; Uozu Aquarium, 1997; Adrianov andKussakin, 1998; Sokolovskaya et al., 1998; Maeda and Tsutsui, 2003; Kitagawa et al., 2008;
Hatooka et al., 2013c; Honma, 2013.
Squalus acanthias acanthias: Lindberg and Legeza, 1959.
*Squalus brevirostris Tanaka, 1917 [Japanese name: Tsumari-tsunozame]
Squalus brevirostris: Mori, 1928; Mori and Uchida, 1934; Mori, 1952.
*Squalus japonicus Ishikawa, 1908 [Japanese name: Togari-tsunozame]
Squalus japonicus: Mori, 1928; Mori and Uchida, 1934; Mori, 1952.
Squalus mitsukurii Jordan and Snyder, 1903 [Japanese name: Futo-tsunozame]
Squalus mitsukurii: Mori, 1928; Mori and Uchida, 1934; Honma, 1952; Mori, 1952; Kato,
1956; Mori, 1956a; Lindberg and Legeza, 1959; Suzuki et al., 2000; Shinohara et al., 2011;
Hatooka et al., 2013c; Honma, 2013.
Squalus mitsukuri: Tsuda, 1990.

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Order Squatiniformes
Family Squatinidae
Squatina japonica Bleeker, 1858 [Japanese name: Kasuzame]
Squatina japonica: Mori and Uchida, 1934; Katayama, 1940; Honma, 1952; Mori, 1952;
Kato, 1956; Mori, 1956a; Honma, 1956b; Lindberg and Legeza, 1959; Honma, 1963; Takegawa
and Morino, 1970; Honma, 1977; Honma and Kitami, 1978; Tsuda, 1990; Sokolovskaya et al.,
1998; Suzuki et al., 2000; Maeda and Tsutsui, 2003; Kobayashi et al., 2006; Hatooka et al.,2013c; Honma, 2013.
Squatina nebulosa Regan, 1906 [Japanese name: Korozame]
Squatina nebulosa: Katayama, 1940; Honma, 1952; Mori, 1952; Kato, 1956; Mori, 1956a;
Tsuda, 1990; Matsuura et al., 2009; Hatooka et al., 2013c; Honma, 2013.
Order Pristiophoriformes
Family Prsitiophoridae*Prsitiophorus japonicus Günther, 1870 [Japanese name: Nokogirizame]
Prsitiophorus japonicus: Mori and Uchida, 1934; Honma, 1952; Mori, 1952; Kato, 1956;
Mori, 1956a; Lindberg and Legeza, 1959; Ueno, 1971; Honma and Kitami, 1978; Tsuda, 1990;
Maeda and Tsutsui, 2003; Hatooka et al., 2013a; Honma, 2013.
Order Rhinobatiformes
Family Rhinobatidae
Rhinobatos hynnicephalus Richardson, 1846 [Japanese name: Komon-sakatazame]
Rhinobatos hynnicephalus: Takegawa and Morino, 1970; Honma et al., 1972; Tsuda, 1990;
Uozu Aquarium, 1997; Honma, 2013.
Rhinobatos schlegelii Muller and Henle, 1841 [Japanese name: Sakatazame]
Rhinobatos schlegeli: Mori, 1928; Mori and Uchida, 1934; Katayama, 1940; Honma, 1952;
Mori, 1952; Kato, 1956; Mori, 1956a; Lindberg and Legeza, 1959; Takegawa and Morino, 1970.
Rhinobatos schlegelii: Tsuda, 1990; Matsuura et al., 2009; Hatooka et al., 2013c.
Rhinobatos granulates: Honma, 2013.
Order Rajiformes
Family Torpedinidae
Narke japonica (Temminck and Schlegel, 1850) [Japanese name: Shibire-ei]
Narke japonica: Mori, 1952; Kato, 1956; Mori, 1956a; Lindberg and Legeza, 1959; Takegawa and Morino, 1970; Tsuda, 1990; Suzuki et al., 2000; Shinohara et al., 2011.
*Torpedo tokionis (Tanaka, 1908) [Japanese name: Yamato-shibire-ei]
Torpedo tokionis: Tsuda, 1990; Hatooka et al., 2013a.
Family Rajidae*Bathyraya aleutica (Gilbert, 1896) [Japanese name: Aryūshan-kasube]
Bathyraya aleutica: Maeda and Tsutsui, 2003.
Bathyraja bergi Dolganov, 1983 [Japanese name: Soko-gangi-ei] (Fig. 2AB)
Breviraja isotrachys: Honma, 1955d; Matsubara, 1955; Kato, 1956; Mori, 1956a; Honma,

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1963; Ouchi, 1963; Takegawa and Morino, 1970; Ueno, 1971; Honma and Kitami, 1978; Tsuda,
1990; Hatooka and Yamada, 1993; Hatooka et al., 2000, 2002.
Breviraja (Bathyraja) isotrachys: Ishiyama, 1967.
Bathyraja bergi: Dolganov, 1985; Amaoka et al., 1995; Adrianov and Kussakin, 1998; Sokolovskaya et al., 1998; Maeda and Tsutsui, 2003; Shinohara et al., 2011; Hatooka et al., 2013b;
Honma, 2013.
TS09-T035 (1), SMBS uncatalogued (1).
Bathyraja smirnovi (Soldatov and Pavlenko, 1915) [Japanese name: Dobu-kasube]
Raja smirnovi: Soldatov and Pavlenko, 1915 (original description).
Breviraja smirnovi: Honma, 1955a; Matsubara, 1955; Kato, 1956; Mori, 1956a, c; Lindberg
and Legeza, 1959; Ouchi, 1963; Ueno, 1971; Tsuda, 1990; Honma, 2013.
Breviraya (Arctoraja) smirnovi: Ishiyama, 1967.
Breviraja smirnovi ankasube: Honma and Kitami, 1978.
Bathyraja smirnovi: Takegawa and Morino, 1970; Ogata et al., 1973; Nakaya and Shirai,
1992; Nambu et al., 1992; Hatooka and Yamada, 1993; Uozu Aquarium, 1997; Hatooka et al.,2000, 2002; Maeda and Tsutsui, 2003; Shinohara et al., 2011; Hatooka et al., 2013b.
TS09-T006 (2).
Dipturus kwangtungensis (Chu, 1960) [Japanese name: Gangi-ei]
Raja kenojei: Mori and Uchida, 1934; Katayama, 1940; Hikita, 1950; Honma, 1952; Mori,
1952; Honma, 1955b; Kato, 1956; Mori, 1956a; Honma, 1963; Takegawa and Morino, 1970;
Ueno, 1971; Honma, 1977; Honma and Kitami, 1978; Tsuda, 1990.
Dipturus kwangtungensis: Suzuki et al., 2000; Maeda and Tsutsui, 2003; Hatooka et al.,2013b; Honma, 2013.
*Dipturus macrocauda (Ishiyama, 1955) [Japanese name: Kitsune-kasube]
Dipturus macrocauda: Hatooka et al., 2013b.
Family Urolophidae
Urolophus aurantiacus Müller and Henle, 1941 [Japanese name: Hirata-ei]
Urolophus fuscus: Honma, 1955c; Kato, 1956.
Urolophus aurantiacus: Mori, 1956a; Honma, 1963; Lindberg and Legeza, 1959; Takegawa
and Morino, 1970; Honma and Kitami, 1978; Tsuda, 1990; Suzuki et al., 2000; Shinohara et al.,2011; Honma, 2013; Yamaguchi et al., 2013a.
Family Dasyatidae
Dasyatis akajei (Müller and Henle, 1841) [Japanese name: Aka-ei]
Dasyatis akajei: Mori, 1928; Mori and Uchida, 1934; Mori, 1952; Honma, 1955b; Kato,
1956; Mori, 1956a; Lindberg and Legeza, 1959; Honma, 1963; Takegawa and Morino, 1970;
Ueno, 1971; Honma, 1977; Honma and Kitami, 1978; Tsuda, 1990; Adrianov and Kussakin,1998; Sokolovskaya et al., 1998; Suzuki et al., 2000; Maeda and Tsutsui, 2003; Matsuura et al.,2009; Honma, 2013; Yamaguchi et al., 2013b.
Order Albuliformes
Family Albulidae
Pterothrissus gissu Hilgendorf, 1877 [Japanese name: Gisu]

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Pterothrissus gissu: Tanaka, 1928; Katayama, 1940; Matsubara, 1955; Kato, 1956; Mori,
1956a; Honma, 1963; Lindberg and Legeza, 1965; Takegawa and Morino, 1970; Ueno, 1971;
Honma and Kitami, 1978; Tsuda, 1990; Aizawa, 1993; Uozu Aquarium, 1997; Aizawa, 2000,
2002; Suzuki et al., 2000; Maeda and Tsutsui, 2003; Shinohara et al., 2011; Aizawa and Doiuchi,
2013; Honma, 2013.
Pterothissus gissu [sic]: Honma, 1952.
Order Anguilliformes
Family Synaphobranchidae*Dysomma anguillare Barnard, 1923 [Japanese name: Asaba-hora-anago]
Dysomma anguillare: Hatooka, 2013.
*Meadia abyssalis (Kamohara, 1938) [Japanese name: Hirejiro-anago]
Meadia abyssalis: Hatooka, 2013.
Family Ophichthidae
Echelus uropterus (Temminck and Schlegel, 1846) [Japanese name: Hire-anago]
Echelus uropterus: Honma, 1962, 1963; Lindberg and Legeza, 1965; Honma and Kitami,
1978; Tsuda, 1990; Uozu Aquarium, 1997; Suzuki et al., 2000; Shinohara et al., 2011; Hatooka,
2013; Honma, 2013.
SMBS uncatalogued (1).
*Ophichthus urolophus (Temminck and Schlegel, 1847) [Japanese name: Suso-umihebi]
Ophichthus urolophus: Katayama, 1940; Kato, 1956; Mori, 1952, 1956a; Tsuda, 1990;
Hatooka, 2013; Honma, 2013.
Ophichthys urolophus: Lindberg and Legeza, 1965.
Ophisurus macrorhynchus Bleeker, 1852 [Japanese name: Dainan-umihebi]
Ophisurus macrorhynchus: Katayama, 1940; Honma, 1952; Mori, 1952; Kato, 1956; Honma,
1963; Mori, 1956a; Lindberg and Legeza, 1965; Takegawa and Morino, 1970; Honma, 1977;
Honma and Kitami, 1978; Tsuda, 1990; Uozu Aquarium, 1997; Suzuki et al., 2000; Matsuura etal., 2009; Shinohara et al., 2011; Hatooka, 2013; Honma, 2013.
*Pisodonophis cancrivorus (Richardson, 1848) [Japanese name: Minami-umihabi]
Pisodonophis cancrivorus: Hatooka, 2013.
Scolecenchelys fuscapenis McCosker, Ide and Endo, 2012 [Japanese name: Tsumaguro-susoumihebi]
Scolecenchelys fuscapenis: McCosker et al., 2012 (original description); Honma, 2013.
Family Congridae*Ariosoma major (Asano, 1958) [Japanese name: Ō-shiro-anago]
Arisoma majus [sic]: Hatooka, 2013.
Ariosoma meeki (Jordan and Snyder, 1900) [Japanese name: Goten-anago]
Anago anago: Tsuda, 1990.
Ariosoma meeki: Matsuura et al., 2009; Hatooka, 2013; Honma, 2013.
*Ariosoma shiroanago (Asano, 1958) [Japanese name: Shiro-anago]
Ariosoma shiroanago shiroanago: Asano, 1958; Lindberg and Legeza, 1965; Asano, 1988;
Hatooka, 1993.

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Alloconger shiroanago: Tsuda, 1990.
Ariosoma shiroanago: Hatooka, 2013.
Conger myriaster (Brevoort, 1856) [Japanese name: Ma-anago]
Astroconger myriaster: Kato, 1956.
Conger myriaster: Ueno, 1971; Tsuda, 1990; Maeda and Tsutsui, 2003; Honma, 2013.
TS09-T001 (1), TS09-T034 (1), TS09-T101 (1).
Remarks. This species occurs at depths shallower than 100 m (Hatooka, 2013) but our specimens were collected at 200–251 m depth. Shinohara and Matsuura (1997) reported this species
can be collected below 380 m depth in Suruga Bay, Japan.
*Congriscus megastomus (Günther, 1877) [Japanese name: Oki-anago]
Congriscus megastomus: Hatooka, 2013.
*Gnathophis ginanago (Asano, 1958) [Japanese name: Nise-gin-anago]
Gnathophis ginanago: Hatooka, 2013.
Gnathophis nystromi nystromi (Jordan and Snyder, 1901) [Japanese name: Gin-anago]
Rhynchocymba nystromi: Katayama, 1940; Kato, 1956.
Arisoma nystromi: Honma, 1952.
Rhynchocymba nyströmi: Mori, 1952, 1956a.
Rhynchocymba nystromi nystromi: Honma, 1956b, 1963; Lindberg and Legeza, 1965; Takegawa and Morino, 1970; Honma and Kitami, 1978.
Rhynchocymba nystromi subsp.: Honma, 1977.
Gnathophis nystromi: Honma, 1990.
Gnathophis nystromi nystromi: Suzuki et al., 2000; Shinohara et al., 2011; Honma, 2013.
Gnathophis heterognathos: Hatooka, 2013.
SMBS uncatalogued (1).
Order Argentiniformes
Family Argentinidae*Argentina kagoshimae Jordan and Snyder, 1902 [Japanese name: Kagoshima-nigisu]
Argentina elongata kagoshimae: Lindberg and Legeza, 1965.
Argentina kagoshimae: Horikawa, 1997; Hatooka, 2013.
Glossanodon semifasciatus (Kishinouye, 1904) [Japanese name: Nigisu]
Argentina semifasciata: Kishinouye, 1904a (original description), b; Katayama, 1940;
Matsubara, 1943a; Mori, 1952; Honma, 1952; Kato, 1956; Mori, 1956a; Honma, 1963; Ouchi,
1963; Lindberg and Legeza, 1965; Honma, 1977; Honma and Kitami, 1978.
Glossanodon semifasciatus: Honma and Kitami, 1967; Takegawa and Morino, 1970; Honmaand Kitami, 1980; Hatooka, 1993; Suzuki et al., 2000; Hatooka, 2000, 2002; Kitagawa et al.,2008; Matsuura et al., 2009; Shinohara et al., 2011; Hatooka, 2013; Honma, 2013.
Order Stomiiformes
Family Sternoptychidae
Maurolicus japonicus Ishikawa, 1915 [Japanese name: Kyūri-eso]
Maurolicus pennanti: Mori and Uchida, 1934.
Maurolicus japonicus: Katayama, 1940; Honma, 1952; Matsubara, 1955; Kato, 1956; Lindberg and Legeza, 1965; Ogata et al., 1973; Mori, 1956a; Honma, 1963; Takegawa and Morino,
1970; Tsuda, 1990; Maeda and Tsutsui, 2003; Kitagawa et al., 2008; Shinohara et al., 2011;

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Aizawa and Doiuchi, 2013; Honma, 2013.
Maurolicus pennanti japonicus: Honma, 1968.
Maurolicus muelleri: Honma and Kitami, 1978; Fujii, 1988; Tsuda, 1990; Aizawa, 1993;
Amaoka et al., 1995; Uozu Aquarium, 1997; Sokolovskaya et al., 1998.
Maurolicus japoicus: Aizawa, 1997.
TS09-T101 (5), TS10-OW18 (3).
*Sternoptyx diaphana Hermann, 1781 [Japanese name: Mune-eso]
Sternoptyx diaphana: Maeda and Tsutsui, 2003; Aizawa and Doiuchi, 2013.
Order Ateleopodiformes
Family Ateleopodidae
Ateleopus japonicus Bleeker, 1854 [Japanese name: Shachiburi]
Ateleopus japonicus: Kato, 1956; Lindberg and Legeza, 1965; Takegawa and Morino, 1970;
Tsuda, 1990; Amaoka and Kobayashi, 2003; Kobayashi et al., 2006; Shinohara et al., 2011;
Honma, 2013.
Ateleopus purpureus: Mori, 1956a.
Ateleopus purpureus Tanaka, 1990 [Japanese name: Murasaki-shachiburi]
Ateleopus purpureus: Kato, 1956; Tsuda, 1990; Suzuki et al., 2000; Kobayashi et al., 2006;
Aizawa and Doiuchi, 2013.
Order Aulopiformes
Family Aulopodidae
Aulopus japonicus Günther, 1877 [Japanese name: Hime]
Hime japonica: Mori, 1952; Katayama, 1940; Honma, 1952; Matsubara, 1955; Kato, 1956;
Mori, 1956a; Honma, 1963; Lindberg and Legeza, 1965; Takegawa and Morino, 1970; Honma,
1977; Honma and Kitami, 1978.
Aulopus japonicus: Tsuda, 1990; Uozu Aquarium, 1997; Suzuki et al., 2000; Matsuura et al.,2009; Shinohara et al., 2011; Honma, 2013.
SMBS uncatalogued (2).
Family Chlorophthalmidae
Chlorophthalmus albatrossis Jordan and Starks, 1904 [Japanese name: Aome-eso]
Chlorophthalmus albatrossis: Katayama, 1940; Kato, 1956; Lindberg and Legeza, 1965;
Tsuda, 1990; Shinohara et al., 2011; Honma, 2013.
Chlorophthalmus albatrosis [sic]: Mori, 1956a.
Chlorophthalmus albatrossis albatrossis: Honma et al., 1972.
Family Paralepididae
Lestidium prolixum Harry, 1953 [Japanese name: Namehadaka]
Lestidium prolixum: Suzuki et al., 2000; Shinohara et al., 2011.

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Order Myctophiformes
Family Myctophidae*Diaphus watasei Jordan and Starks, 1904 [Japanese name: Hadaka-iwashi]
Diaphus coeruleus: Kato, 1956; Mori, 1956a; Lindberg and Legeza, 1965.
Diaphus watasei: Tsuda, 1990.
*Myctophum asperum Richardson, 1845 [Japanese name: Ara-hadaka]
Myctophum asperum: Mori, 1952.
*Myctophum nitidulum Garman, 1899 [Japanese name: Susuki-hadaka]
Myctophum afﬁne: Mori, 1952.
*Notoscopelus japonicus (Tanaka, 1908) [Japanese name: Ōkuchi-iwashi]
Notoscopelus elongatus: Kato, 1956; Mori, 1956a.
Notoscopelus japonicus: Tsuda, 1990; Nakabo, 1993b, 2000b, 2002b; Nakabo and Kai, 2013.
Order Lampridiformes
Family Veliferidae
Velifer hypselopterus Bleeker, 1879 [Japanese name: Kusa-aji]
Velifer hypselopterus: Honma, 1952; Mori, 1952; Kato, 1956; Mori, 1956a; Lindberg andLegeza, 1965; Fujii, 1988; Tsuda, 1990; Honma, 1991a; Aizawa, 1993, 2000, 2002; Shinohara etal., 2011; Aizawa and Doiuchi, 2013; Honma, 2013.
Family Lampridae
Lampris guttatus (Brünnich 1788) [Japanese name: Akamanbō]Lampris regia: Katayama, 1940; Honma, 1952; Tsuda, 1990.
Lampris regius: Honma, 1955b; Kato, 1956; Mori, 1956a; Honma, 1963; Lindberg andLegeza, 1965; Honma and Kitami, 1978.
Lampris guttatus: Fujii, 1988; Aizawa, 1993; Amaoka et al., 1995; Uozu Aquarium, 1997;
Aizawa, 2000, 2002; Kitagawa et al., 2008; Aizawa and Doiuchi, 2013; Honma, 2013.
Family Lophotidae*Eumecichthys ﬁski (Günther, 1890) [Japanese name: Tengunotachi]
Eumecichthys ﬁski: Matsubara, 1955; Mori, 1956a; Tsuda, 1990; Amaoka et al., 1995;
Honma et al., 1995; Hayashi, 2000, 2002, 2013.
Eumethichthys ﬁski: Kamohara, 1942; Kato, 1956; Lindberg and Legeza, 1965; Fujii, 1988.
Eumethichthys ﬁskii: Hayashi, 1993.
Eumecichthys ﬁskii: Honma, 2013.
Lophotus capellei Temminck and Schlegel, 1845 [Japanese name: Akanamada]Lophotes capellei [sic]: Tsuda, 1990.
Lophotus capellei: Honma and Kitami, 1980; Honma and Mizusawa, 1981; Fujii, 1988;
Amaoka et al., 1995; Hayashi, 1993; Uozu Aquarium, 1997; Hayashi, 2000, 2002; Kobayashi etal., 2006; Shinohara et al., 2011; Hayashi, 2013; Honma, 2013.
Family Trachipteridae
Desmodema polystictum (Ogilby, 1898) [Japanese name: Furisode-uo]

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Trachipterus misakiensis: Matsubara, 1955; Kato, 1956; Mori, 1956a.
Trachypterus misakiensis: Lindberg and Legeza, 1965.
Desmodema polystictum: Fujii, 1988; Tsuda, 1990; Hayashi, 1993; Amaoka et al., 1995;
Tsuda, 1990; Hayashi, 2000, 2002; Kobayashi et al., 2006; Shinohara et al., 2011; Hayashi andSenou, 2013.
Trachipterus ishikawae Jordan and Snyder, 1901 [Japanese name: Sakegashira]Trachipterus sp.: Mori, 1952.
Trachipterus ishikawai: Matsubara, 1955; Kato, 1956; Mori, 1956a; Honma, 1962, 1963,
1968; Takegawa and Morino, 1970; Ueno, 1971; Honma, 1977.
Trachipterus ishikawae: Honma and Kitami, 1967, 1978; Fujii, 1988; Tsuda, 1990; Uozu
Aquarium, 1997; Hayashi, 1993; Amaoka et al., 1995; Hayashi, 2000, 2002; Maeda and Tsutsui,
2003; Kobayashi et al., 2006; Kitagawa et al., 2008; Shinohara et al., 2011; Hayashi and Senou,
2013; Honma, 2013.
Trachypterus ishikawai: Lindberg and Legeza, 1965.
TS09-T023 (1), TS09-T086 (1).
Trachipterus trachypterus (Gmelin, 1789) [Japanese name: Tengaihata]
Trachipterus iris: Matsubara, 1955; Kato, 1956; Mori, 1956a; Tsuda, 1990.
Trachypterus iris: Lindberg and Legeza, 1965.
Trachipterus trachypterus: Fujii, 1988; Kobayashi et al., 2006; Shinohara et al., 2011.
Zu cristatus (Bonelli, 1820) [Japanese name: Yuki-furisodeuo]
Trachypterus ijimae: Katayama, 1940.
Trachipterus ijimae: Honma, 1952.
Trachipterus ijimai: Matsubara, 1955; Kato, 1956; Mori, 1956a; Takegawa and Morino,
1970.
Trachypterus ijimai: Lindberg and Legeza, 1965.
Zu cristatus: Tsuda, 1990; Amaoka et al., 1995; Hayashi, 1993; Uozu Aquarium, 1997;
Hayashi, 2000, 2002; Maeda and Tsutsui, 2003; Kobayashi et al., 2006; Hayashi and Senou,
2013; Honma, 2013.
SMBS uncatalogued (1).
Family Regalecidae
Regalecus russellii (Shaw, 1803) [Japanese name: Ryūgūnotsukai]
Regalecus russellii: Okada and Matsubara, 1938; Honma, 1952; Mori, 1952; Matsubara,
1955; Kato, 1956; Mori, 1956a, b; Honma, 1963; Honma and Kitami, 1967; Lindberg andLegeza, 1965; Ueno, 1971; Honma and Kitami, 1978; Fujii, 1988; Tsuda, 1990; Hayashi, 1993,
2000, 2002; Kobayashi et al., 2006; Shinohara et al., 2011; Hayashi, 2013; Honma, 2013.
Regalecus russelli: Takegawa and Morino, 1970.
Order Polymixiniiformes
Family Polymixiniidae*Polymixia japonica Günther, 1877 [Japanese name: Ginmedai]
Polymixia japonica: Tsuda, 1990.

Annotated Checklist of Deep-sea Fishes of the Sea of Japan

241

Order Ophidiiformes
Family Ophidiidae
Hoplobrotula armata (Temminck and Schlegel, 1846) [Japanese name: Yoroi-itachi-uo]
Hoplobrotula armata: Katayama, 1940; Honma, 1952; Mori, 1952; Matsubara, 1955; Kato,
1956; Mori, 1956a; Takegawa and Morino, 1970; Lindberg and Krasyukova, 1975; Tsuda, 1990;
Uozu Aquarium, 1997; Suzuki et al., 2000; Kitagawa et al., 2008; Matsuura et al., 2009; Shinohara et al., 2011; Honma, 2013; Nakabo and Kai, 2013.
Neobythites sivicola (Jordan and Snyder, 1901) [Japanese name: Shio-itachi-uo]
Watasea sivicola: Katayama, 1940; Honma, 1952; Mori, 1952.
Neobythites sivicolus: Matsubara, 1955; Kato, 1956; Mori, 1956a; Honma, 1963; Takegawa
and Morino, 1970; Lindberg and Krasyukova, 1975; Honma, 1977; Honma and Kitami, 1978;
Tsuda, 1990; Nakabo, 2000b; Suzuki et al., 2000; Nakabo, 2002b; Honma, 2013.
Neobythites sivicola: Uozu Aquarium, 1997; Machida, 1988; Matsuura et al., 2009; Shinohara et al., 2011; Nakabo and Kai, 2013.
*Ophidion asiro (Jordan and Fowler, 1902) [Japanese name: Ashiro]
Otophidium asiro: Mori, 1956a; Kato, 1956; Fowler, 1958; Lindberg and Krasyukova, 1975.
Ophidion asiro: Tsuda, 1990; Nakabo and Kai, 2013.
*Sirembo imberbis (Temminck and Schlegel, 1846) [Japanese name: Umidojō]
Sirembo imberbis: Mori and Uchida, 1934; Katayama, 1940; Honma, 1952; Mori, 1952;
Matsubara, 1955; Kato, 1956; Mori, 1956a; Fowler, 1958; Lindberg and Krasyukova, 1975;
Tsuda, 1990; Honma, 2013; Nakabo and Kai, 2013.
Order Gadiformes
Family Moridae
Lotella phycis (Temminck and Schlegel, 1846) [Japanese name: Iso-ainame]Lotella phycis: Honma, 1952; Mori, 1952; Kato, 1956; Lindberg and Legeza, 1965; Tsuda,
1990; Uozu Aquarium, 1997; Suzuki et al., 2000; Shinohara et al., 2011; Honma, 2013.
Physiculus japonicus Hilgendorf, 1881 [Japanese name: Chigodara]
Physiculus japonicus: Katayama, 1940; Honma, 1952; Kato, 1956; Lindberg and Legeza,
1965; Tsuda, 1990; Suzuki et al., 2000; Balushkin and Prirodina, 2008; Shinohara et al., 2011;
Honma, 2013.
Family Macrouridae*Abyssicola macrochir (Günther, 1877) [Japanese name: Tenaga-dara]
Abyssicola macrochir: Honma et al., 2008; Honma, 2013.
*Coelorhynchus japonicus (Temminck and Schlegel, 1846) [Japanese name: Tōjin]Coelorhynchus japonicus: Katayama, 1940; Mori, 1952, Kato, 1956; Mori, 1956a; Lindberg
and Legeza, 1965; Tsuda, 1990; Nakabo and Kai, 2013.
Coelorhynchus multispinulosus Katayama, 1942 [Japanese name: Yari-hige]
Coelorhynchus multispinulosus: Matsubara, 1955; Kato, 1956; Mori, 1956a; Lindberg andLegeza, 1965; Okamura, 1970; Honma and Kitami, 1978; Honma et al., 1972; Tsuda, 1990;
Shinohara et al., 2011.
Coelorinchus multispinosus: Nakabo, 1993b; Okamura, 1997; Nakabo, 2000b, 2002b;
Honma, 2013; Nakabo and Kai, 2013.

242

Gento Shinohara, Masanori Nakae, Yuji Ueda, Shigeaki Kojima and Keiichi Matsuura*Coelorinchus parallelus (Günther, 1877) [Japanese name: Soroi-hige]
Coelorinchus parallelus: Kato, 1956; Lindberg and Legeza, 1965; Tsuda, 1990.
Family Bregmacerotidae*Bregmaceros japonicus Tanaka, 1908 [Japanese name: Sai-uo]
Bregmaceros japonicus: Tanaka, 1913a; Jordan and Hubbs, 1925; Okada and Matsubara,
1938; Katayama, 1940; Honma, 1952; Mori, 1952; Matsubara, 1955; Kato, 1956; Mori, 1956a;
Lindberg and Legeza, 1965; Honma, 1968; Honma and Kitami, 1978; Tsuda, 1990.
Bregmaceros nectabanus Whitley, 1941 [Japanese name: Toyama-sai-uo]
Bregmaceros nectabanus: Masuda et al., 1986; Tsuda, 1990; Nakabo, 1993b; Uozu Aquarium, 1997; Nakabo, 2000b; Shinohara et al., 2011; Honma, 2013; Nakabo and Kai, 2013.
Bregmaceros neonectabanus Masuda, Ozawa and Tabeta, 1986 [Japanese name: Kurohara-saiuo]
Bregmaceros neonectabanus: Masuda et al., 1986 (original description); Nakabo, 1993b,
2000b, 2002b.
Family Gadidae
Gadus macrocephalus Tilesius, 1810 [Japanese name: Madara]
Gadus macrocephalus: Snyder, 1912; Mori, 1928; Mori and Uchida, 1934; Okada andMatsubara, 1938; Katayama, 1940; Honma, 1952; Katayama, 1952; Mori, 1952; Matsubara,
1955; Kato, 1956; Mori, 1956a; Honma, 1957b, 1963; Ouchi, 1963; Ueno, 1971; Ogata et al.,
1973; Honma and Kitami, 1978; Tsuda, 1990; Endo, 1997; Sokolovskaya et al., 1998; Suzuki etal., 2000; Maeda and Tsutsui, 2003; Balushkin and Prirodina, 2008; Kitagawa et al., 2008; Matsuura et al., 2009; Shinohara et al., 2011; Honma, 2013.
Gadus morhua macrocephalus: Lindberg and Legeza, 1965; Adrianov and Kussakin, 1998.
KT-11-09-T1 (1), TS09-T011 (1), TS09-T038 (1).
Theragra chelcogramma (Pallas, 1814) [Japanese name: Suketō-dara]
Theragra chelcogramma: Mori, 1928; Mori and Uchida, 1934; Okada and Matsubara, 1938;
Katayama, 1940; Honma, 1952; Katayama, 1952; Mori, 1952; Honma, 1955b; Matsubara, 1955;
Kato, 1956; Mori, 1956a, c; Honma, 1963; Ouchi, 1963; Lindberg and Legeza, 1965; Ueno,
1971; Ogata et al., 1973; Honma and Kitami, 1978; Honma and Kitami, 1978; Okamura, 1988;
Tsuda, 1990; Nambu et al., 1992; Nakabo, 1993b; Endo, 1997; Adrianov and Kussakin, 1998;
Sokolovskaya et al., 1998; Nakabo, 2000b; Suzuki et al., 2000; Nakabo, 2002b; Maeda and Tsutsui, 2003; Balushkin and Prirodina, 2008; Kitagawa et al., 2008; Matsuura et al., 2009; Shinohara et al., 2011; Honma, 2013; Nakabo and Kai, 2013.
KT-10-08-R2 (2), SMBS uncatalogued (2).
Order Lophiiformes
Family Lophiidae
Lophiomus setigerus (Vahl, 1797) [Japanese name: Ankō]
Lophiomus setigerus: Mori, 1928; Mori and Uchida, 1934; Katayama, 1940; Honma, 1952;
Mori, 1952; Kato, 1956; Mori, 1956a; Honma, 1963; Ueno, 1971; Honma and Kitami, 1978;
Honma and Kitami, 1978; Tsuda, 1990; Suzuki et al., 2000; Maeda and Tsutsui, 2003; Matsuura
et al., 2009; Shinohara et al., 2011; Honma, 2013; Yamada and Yagishita, 2013.

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243

Lophius litulon (Jordan, 1902) [Japanese name: Ki-ankō]
Lophius litulon: Snyder, 1912; Mori, 1928; Mori and Uchida, 1934; Katayama, 1940;
Honma, 1952; Mori, 1952; Kato, 1956; Mori, 1956a; Honma, 1963; Honma and Kitami, 1967;
Ueno, 1971; Honma and Kitami, 1978; Honma, 1977; Honma and Kitami, 1978; Tsuda, 1990;
Suzuki et al., 2000; Maeda and Tsutsui, 2003; Matsuura et al., 2009; Shinohara et al., 2011;
Honma, 2013; Yamada and Yagishita, 2013.
TS09-T001 (1), TS09-T008 (1), TS09-T035 (1), TS09-T050 (1), TS09-T102 (1), SMBS
uncatalogued (2).
Family Chaunacidae*Chaunax abei Le Danois, 1978 [Japanese name: Midori-fusa-ankō]
Chaunax ﬁmbriatus: Katayama, 1940; Matsubara, 1955; Kato, 1956; Masuda et al., 1975;
Tsuda, 1990.
Chaunax abei: Lindberg et al., 1997; Yamada and Yagishita, 2013.
Family Ogcocephalidae
Halieutaea stellata (Vahl, 1797) [Japanese name: Akagutsu]
Halieutaea stellata: Mori and Uchida, 1934; Katayama, 1940; Honma, 1952; Mori, 1952;
Kato, 1956; Mori, 1956a; Honma and Kitami, 1978; Tsuda, 1990; Suzuki et al., 2000; Matsuura
et al., 2009; Shinohara et al., 2011; Honma, 2013; Yamada and Yagishita, 2013.
*Malthopsis lutea Alcock, 1891 [Japanese name: Fūryū-uo]
Malthopsis lutea: Mori, 1952; Kato, 1956; Mori, 1956a; Tsuda, 1990; Yamada, 2002.
Malthopsis kobayashii: Yamada and Yagishita, 2013.
Family Ceratiidae*Cryptopsaras couesii Gill 1883 [Japanese name: Mitsukuri-enaga-chōchin-ankō]
Cryptopsaras couesi: Okada and Matsubara, 1938; Matsubara, 1955.
Cryptopsaras coues [sic]: Tsuda, 1990.
Cryptopsaras couesii: Nakabo, 1993b, 2000b, 2002b.
Family Himantolophidae
Himantolophus sagamius (Tanaka, 1918) [Japanese name: Chōchin-ankō]
Himantolophus groenlandicus: Maeda and Tsutsui, 2003.
Order Beryciformes
Family Berycidae
Beryx decadactylus Cuvier, 1829 [Japanese name: Nanʼyō-kinme] (Fig. 2C)
Beryx decadactylus: Mori, 1952; Lindberg and Legeza, 1965; Honma et al., 1990; Honma,
1991a; Uozu Aquarium, 1997; Honma, 2013.
SMBS uncatalogued (4).
Beryx splendens Lowe 1834 [Japanese name: Kinmedai]
Beryx splendens: Katayama, 1940; Honma, 1952; Kato, 1956; Lindberg and Legeza, 1965;
Tsuda, 1990; Uozu Aquarium, 1997; Kitagawa et al., 2008; Honma, 2013.

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Order Zeiformes
Family Zeidae*Zenion japonicum Kamohara, 1934 [Japanese name: Soko-matōdai]Zenion japonicum: Katayama, 1940; Kato, 1956; Lindberg and Legeza, 1965; Tsuda, 1990.
Zenopsis nebulosa (Temminck and Schlegel, 1845) [Japanese name: Kagamidai]
Zenopsis nebulosa: Mori and Uchida, 1934; Katayama, 1940; Mori, 1952; Kato, 1956; Mori,
1956a; Honma, 1962, 1963; Honma and Kitami, 1967; Lindberg and Legeza, 1965; Ueno, 1971;
Honma and Kitami, 1978; Tsuda, 1990; Uozu Aquarium, 1997; Sokolovskaya et al., 1998;
Suzuki et al., 2000; Maeda and Tsutsui, 2003; Kobayashi et al., 2006; Shinohara et al., 2011;
Honma, 2013; Nakabo and Kai, 2013.
Zeus nebulosa: Honma, 1977.
TS09-T006 (1).
Zeus faber Linnaeus, 1758 [Japanese name: Matōdai]Zeus japonicus: Mori, 1928; Mori and Uchida, 1934; Katayama, 1940; Honma, 1952; Mori,
1952; Honma, 1955b; Kato, 1956; Mori, 1956a; Honma, 1963; Lindberg and Legeza, 1965;
Takegawa and Morino, 1970; Ueno, 1971; Honma, 1977; Honma and Kitami, 1978; Tsuda, 1990.
Zeus faber: Suzuki et al., 2000; Maeda and Tsutsui, 2003; Matsuura et al., 2009; Shinohara
et al., 2011; Honma, 2013; Nakabo and Kai, 2013.
SMBS uncatalogued (1).
Family Grammicolepididae*Xenolepidichthys dalgleishi Gilchrist, 1922 [Japanese name: Hishi-matōdai]Xenolepidichthys dalgleishi: Tsuda, 1990.
Order Gasterosteiformes
Family Macroramphosidae*Macroramphosus japonicus (Günther, 1861) [Japanese name: Daikoku-sagifue]
Macroramphosus japonicas [sic]: Honma, 2013
Macroramphosus japonicus: Senou, 2013.
Macroramphosus scolopax (Linnaeus, 1758) [Japanese name: Sagifue]
Macroramphosus sagifue: Katayama, 1940; Honma, 1952.
Macrorhamphosus scolopax: Kato, 1956; Mori, 1956a; Takegawa and Morino, 1970; Tsuda,
1990; Uozu Aquarium, 1997; Suzuki et al., 2000; Kitagawa et al., 2008; Honma, 2013; Senou,
2013.
Order Scorpaeniformes
Family Scorpaenidae
Helicolenus avius Abe and Eschmeyer, 1972 [Japanese name: Oki-kasago]
Helicolenus avius: Shinohara et al., 2011.
Helicolenus hilgendorﬁ (Döderlein, 1884) [Japanese name: Yume-kasago]
Helicolenus hilgendorﬁ: Matsubara, 1943b, 1955; Mori, 1956a; Takegawa and Morino, 1970;
Tsuda, 1990; Amaoka et al., 1995; Uozu Aquarium, 1997; Suzuki et al., 2000; Shinohara et al.,2011; Nakabo and Kai, 2013.
Helicolenus dactylopterus: Mori, 1952.

Annotated Checklist of Deep-sea Fishes of the Sea of Japan245

TS09-T036 (1), TS09-T037 (1), TS09-T056 (1).
Hozukius emblemarius (Jordan and Starks, 1904) [Japanese name: Hōzuki]
Hozukius emblemarius: Shinohara et al., 2011; Nakabo and Kai, 2013.
*Pontinus macrocephalus (Sauvage, 1882) [Japanese name: Hiodoshi]
Pontinus macrocephalus: Takegawa and Morino, 1970; Nakabo and Kai, 2013.
Scorpaena neglecta Temminck and Schlegel, 1843 [Japanese name: Izu-kasago] (Fig. 2D)
Scorpaena izensis: Kato, 1956; Takegawa and Morino, 1970; Tsuda, 1990.
Scorpaena neglecta: Kitagawa et al., 2008; Matsuura et al., 2009; Honma, 2013; Nakabo andKai, 2013.
SMBS uncatalogued (2).
*Scorpaena onaria Jordan and Snyder, 1900 [Japanese name: Fusa-kasago]
Scorpaena neglecta: Kato, 1956.
Scorpaena neglecta neglecta: Takegawa and Morino, 1970; Tsuda, 1990.
Scorpaena onaria: Nakabo, 1993b; Honma, 2013; Nakabo and Kai, 2013.
Sebastes baramenuke (Wakiya, 1917) [Japanese name: Bara-menuke]
Sebastes baramenuke: Matsubara, 1943b (original description); Tomiyama and Abe, 1958;
Honma et al., 1984; Lindberg and Krasyukova, 1987; Tsuda, 1990; Maeda and Tsutsui, 2003;
Kitagawa et al., 2008; Shinohara et al., 2011; Honma, 2013; Nakabo and Kai, 2013.
Sebastes (Acutomentum) baramenuke: Mori, 1952.
SMBS uncatalogued (1).
Sebastes glaucus Hilgendorf, 1880 [Japanese name: Kuro-menuke]
Sebastodes glaucus: Soldatov and Lindberg, 1930; Katayama, 1940.
Sebastodes glaucus: Katayama, 1940.
Sebastes glaucus: Kato, 1956; Ueno, 1971; Kanayama and Kitagawa, 1983; Lindberg andKrasyukova, 1987; Amaoka, 1988; Tsuda, 1990; Nakabo, 1993b; Ishida, 1997; Nakabo, 2000b,
2002b; Maeda and Tsutsui, 2003; Kitagawa et al., 2008; Shinohara et al., 2011; Nakabo and Kai,
2013.
Sebastes itinus (Jordan and Starks, 1904) [Japanese name: Yanagi-mebaru]
Sebastes itinus: Honma, 1955d; Kato, 1956; Tsuda, 1990; Suzuki et al., 2000; Maeda andTsutsui, 2003; Shinohara et al., 2011; Honma, 2013; Nakabo and Kai, 2013.
Sebastes matsubarae Hilgendorf, 1880 [Japanese name: Akōdai]Sebastodes matsubarae: Hilgendorf, 1880 (original description); Katayama, 1940.
Sebastes matsubarae: Matsubara, 1943b; Honma, 1952; Tomiyama and Abe, 1958; Uozu
Aquarium, 1997; Maeda and Tsutsui, 2003; Shinohara et al., 2011; Honma, 2013; Nakabo andKai, 2013.
Sebastes matsubarai: Kato, 1956; Mori, 1956a; Lindberg and Krasyukova, 1987; Tsuda,
1990.
Sebastes owstoni (Jordan and Thompson, 1914) [Japanese name: Hatsume]
Sebastodes ruber: Pavlenko, 1910 (original descrption); Soldatov and Lindberg, 1930.
Sebastodes owstoni: Mori and Uchida, 1934; Hikita, 1950.
Sebastes owstoni: Okada and Matsubara, 1938; Katayama, 1940; Matsubara, 1943b; Honma,
1952; Katayama, 1952; Matsubara, 1955; Kato, 1956; Mori, 1956a; Honma, 1963; Ouchi, 1963;
Honma and Kitami, 1970; Takegawa and Morino, 1970; Ueno, 1971; Ogata et al., 1973; Honma,
1977; Honma and Kitami, 1978; Kanayama and Kitagawa, 1983; Lindberg and Krasyukova,
1987; Amaoka, 1988; Tsuda, 1990; Nambu et al., 1992; Nakabo, 1993b; Amaoka et al., 1995;
Ishida, 1997; Adrianov and Kussakin, 1998; Sokolovskaya et al., 1998; Nakabo, 2000b; Suzuki

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Gento Shinohara, Masanori Nakae, Yuji Ueda, Shigeaki Kojima and Keiichi Matsuura

et al., 2000; Nakabo, 2002b; Maeda and Tsutsui, 2003; Kitagawa et al., 2008; Matsuura et al.
2009; Shinohara et al., 2011; Honma, 2013; Nakabo and Kai, 2013.
Sebastes (Hatsumeus) owstoni: Mori, 1952.
TS09-T018 (1), TS09-T034 (1), TS09-T037 (2) TS09-T039 (2), TS09-T101 (1), TS09-T102(1).
Sebastes steindachneri Hilgendorf, 1880 [Japanese name: Yanaginomai]
Sebastes steindachneri: Hilgendorf, 1880 (original description); Honma, 1952; Kato, 1956;
Ouchi, 1963; Ueno, 1971; Amaoka, 1988; Tsuda, 1990; Nakabo, 1993b; Amaoka et al., 1995;
Ishida, 1997; Adrianov and Kussakin, 1998; Sokolovskaya et al., 1998; Nakabo, 2000b, 2002b;
Maeda and Tsutsui, 2003; Kitagawa et al., 2008; Matsuura et al. 2009; Shinohara et al., 2011;
Honma, 2013; Nakabo and Kai, 2013.
Sebastodes steindachneri: Soldatov and Lindberg, 1930.
Sebastiscus albofasciatus (Lacepède, 1802) [Japanese name: Ayame-kasago]
Sebastiscus albofasciatus: Kato, 1956; Takegawa and Morino, 1970; Shimizu, 1988; Tsuda,
1990; Nakabo, 1993b; Ishida, 1997; Nakabo, 2000b, Suzuki et al., 2000; Nakabo, 2002b;
Honma, 2013; Nakabo and Kai, 2013.
*Sebastolobus macrochir (Günther, 1877) [Japanese name: Kichiji]
Sebastolobus macrochir: Mori, 1952; Honma, 1952; Kato, 1956; Mori, 1956a; Tsuda, 1990;
Sokolovskaya et al., 1998; Honma, 2013; Nakabo and Kai, 2013.
Setarches longimanus (Alcock, 1894) [Japanese name: Aka-kasago]
Setarches longimanus: Suzuki et al., 2000; Shinohara et al., 2011; Nakabo and Kai, 2013.
Family Triglidae
Chelidonichthys spinosus (McClelland, 1843) [Japanese name: Hōbō]
Chelidonichthys kumu: Snyder, 1912; Mori, 1928; Matsubara and Hiyama, 1932; Mori and
Uchida, 1934; Katayama, 1940; Honma, 1952; Mori, 1952; Honma, 1955b; Kato, 1956; Mori,
1956a; Honma, 1963; Takegawa and Morino, 1970; Honma, 1977; Honma and Kitami, 1978;
Sokolovskaya et al., 1998.
Chelidonichthys kum [sic]: Ueno, 1971.
Chelidonichthys spinosus: Tsuda, 1990; Suzuki et al., 2000; Maeda and Tsutsui, 2003;
Matsuura et al., 2009; Shinohara et al., 2011; Honma, 2013; Yamada and Yagishita, 2013.
Trigla (Chelidonichthys) kumu: Adrianov and Kussakin, 1998.
SMBS uncatalogued (1).
Lepidotrigla abyssalis Jordan and Starks, 1902 [Japanese name: Soko-kanagashira]Lepidotrigla abyssalis: Katayama, 1940; Honoma, 1952; Kato, 1956; Mori, 1956a; Takegawa and Morino, 1970; Tsuda, 1990; Suzuki et al., 2000; Matsuura et al. 2009; Shinohara et al.,2011; Honma, 2013; Yamada and Yagishita, 2013.
SMBS uncatalogued (1).
Lepidotrigla guentheri Hilgendorf, 1879 [Japanese name: Kanado]
Lepidotrigla güntheri: Mori and Uchida, 1934; Katayama, 1940; Honma, 1952; Kato, 1956;
Honma, 1963, 1977; Honma and Kitami, 1978.
Lepidotrigla guentheri: Mori, 1952, 1956a; Tomiyama and Abe, 1958; Lindberg and Krasyukova, 1987; Tsuda, 1990; Suzuki et al., 2000; Shinohara et al., 2011.
Lepidotrigla microptera Günther, 1873 [Japanese name: Kanagashira]Lepidotrigla strauchi: Snyder, 1912.

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247

Lepidotrigla microptera: Katayama, 1940; Hikita, 1950; Honma, 1952; Mori, 1952; Honma,
1956b; Kato, 1956; Mori, 1956a; Honma, 1963; Takegawa and Morino, 1970; Ueno, 1971;
Honma and Kitami, 1978; Tsuda, 1990; Adrianov and Kussakin, 1998; Sokolovskaya et al.,
1998; Suzuki et al., 2000; Maeda and Tsutsui, 2003; Matsuura et al. 2009; Shinohara et al., 2011;
Honma, 2013; Yamada and Yagishita, 2013.
SMBS uncatalogued (1).
Peristedion orientale Temminck and Schlegel, 1843 [Japanese name: Kihōbō]
Peristedion orientale: Mori, 1928; Mori and Uchida, 1934; Katayama, 1940; Mori, 1952;
Kato, 1956; Honma et al., 1972; Tsuda, 1990; Matsuura et al. 2009; Shinohara et al., 2011;
Honma, 2013; Yamada and Yagishita, 2013.
*Satyrichthys rieffeli (Kaup, 1859) [Japanese name: Iso-kihōbō]
Satyrichthys rieffeli: Kato, 1956; Tsuda, 1990.
*Scalicus amiscus (Jordan and Starks, 1904) [Japanese name: Hige-kihōbō]
Satyrichthys amiscus: Tsuda, 1990.
Family Hoplichthyidae*Hoplichthys ﬁlamentosus Matsubara and Ochiai, 1950 [Japanese name: Ito-harigochi]
Hoplichthys ﬁlamentosus: Nakabo and Kai, 2013.
*Hoplichthys gilberti Jordan and Richardson, 1908 [Japanese name: Soko-harigochi]
Hoplichthys gilberti: Nakabo and Kai, 2013.
*Hoplichthys langsdorﬁi Cuvier, 1829 [Japanese name: Natsu-harigochi]
Hoplichthys langsdorﬁi: Matsubara and Ochiai, 1950; Kato, 1956; Mori, 1956a; Takegawa
and Morino, 1970; Tsuda, 1990.
*Hoplichthys regani Jordan, 1908 [Japanese name: Harigochi]
Hoplichthys regani: Kato, 1956; Mori, 1956a; Takegawa and Morino, 1970; Tsuda, 1990.
Family Bembridae
Bembras japonicus Cuvier, 1829 [Japanese name: Akagochi]
Bembras japonica: Mori, 1956a.
Bembras japonicus: Katayama, 1940; Kato, 1956; Lindberg and Krasyukova, 1987; Tsuda,
1990; Shinohara et al., 2011; Nakabo and Kai, 2013.
Parabembras curtus (Temminck and Schlegel, 1843) [Japanese name: Ubagochi]
Parabembras curtus: Mori, 1928; Takegawa and Morino, 1970; Tsuda, 1990; Nakabo andKai, 2013.
Parabembras curta: Shinohara et al., 2011.
Family Dactylopteridae*Dactyloptena orientalis Cuvier, 1829 [Japanese name: Semihōbō]
Dactyloptena orientalis: Kato, 1956; Ueno, 1971; Honma, 2013; Tsuda, 1990.
Daicocus peterseni (Nyström, 1887) [Japanese name: Hoshi-semihōbō]Daicocus peterseni: Mori and Uchida, 1934; Katayama, 1940; Honma, 1952; Mori, 1952;
Honma, 1956b; Kato, 1956; Honma,1963; Takegawa and Morino, 1970; Honma and Kitami,
1978; Lindberg et al., 1997; Uozu Aquarium, 1997; Suzuki et al., 2000; Maeda and Tsutsui,
2003; Matsuura et al., 2009; Honma, 2013; Nakabo and Doiuchi, 2013.

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Gento Shinohara, Masanori Nakae, Yuji Ueda, Shigeaki Kojima and Keiichi Matsuura

Family Anolopomatidae*Anoplopoma ﬁmbria (Pallas, 1814) [Japanese name: Gindara]
Anoplopoma ﬁmbria: Maeda and Tsutsui, 2003.
*Erilepis zonifer (Lockington, 1880) [Japanese name: Aburabōzu]
Erilepis zonifer: Nakabo and Kai, 2013.
Family Ereuniidae
Marukawichthys ambulator Sakamoto, 1931 [Japanese name: Marukawa-kajika]
Marukawichthys ambulator: Sakamoto, 1931 (original description); Katayama, 1940;
Honma, 1955a, c; Matsubara, 1955; Watanabe, 1960; Honma, 1963; Takegawa and Morino,
1970; Lindberg and Krasyukova, 1975; Honma and Kitami, 1978; Yabe, 1988; Tsuda, 1990;
Nakabo, 1993b; Uozu Aquarium, 1997; Nakabo, 2000b, 2002b; Sideleva et al., 2006a; Matsuura
et al., 2009; Shinohara et al., 2011; Nakabo and Kai, 2013.
Morukawichthys ambulator [sic]: Kato, 1956.
Marukawichthys ambulatory [sic]: Honma, 2013.
Family Cottidae
Alcichthys elongatus (Steindachner, 1881) [Japanese name: Niji-kajika]
Alcichthys alcicornis: Kato, 1956; Watanabe, 1960; Ouchi, 1963; Tsuda, 1990; Amaoka etal., 1995; Yabe, 1997.
Alcichthys okiensis: Kato, 1956; Mori, 1956d (original description).
Alcichthys elongatus: Maeda and Tsutsui, 2003; Sideleva et al., 2006a; Kitagawa et al., 2008;
Matsuura et al., 2009; Honma, 2013; Nakabo and Kai, 2013.
Artediellus aporosus Soldatov, 1922 [Japanese name: Madara-okikajika]
Artediellus aporosus: Okada and Matsubara, 1938; Sideleva et al., 2006b; Nakabo and Kai,
2013.
*Artediellus camchaticus Gilbert and Burke, 1912 [Japanese name: Kamuchakka-kajika]
Artediellus camchaticus: Okada and Matsubara, 1938; Nakabo and Kai, 2013.
Artediellus dydymovi Soldatov, 1915 [Japanese name: Kobu-okikajika]
Artediellus dydymovi: Okada and Matsubara, 1938; Ueno, 1971; Adrianov and Kussakin,1998; Sokolovskaya et al., 1998; Maeda and Tsutsui, 2003; Nakabo and Kai, 2013.
Artediellus dydymovi dydymovi: Ouchi, 1963; Yabe, 1988; Tsuda, 1990; Amaoka et al., 1995;
Sideleva et al., 2006b; Honma, 2013.
TS09-T008 (2).
Artediellus fuscimentus Nelson, 1986 [Japanese name: Nodoguro-okikajika]
Artediellus fuscimentus: Nakabo, 1993b, 2000b, 2002b; Shinohara et al., 2011; Nakabo andKai, 2013.
TS09-T047 (3), TS09-T049 (1), TS09-T088 (1).
Artediellus ochotensis Gilbert and Burke, 1912 [Japanese name: Ohōtsuku-kajika]
Artediellus ochotensis: Okada and Matsubara, 1938; Ueno, 1971; Amaoka et al., 1995; Adrianov and Kussakin, 1998; Sokolovskaya et al., 1998; Sideleva et al., 2006b; Nakabo and Kai,
2013.
Cottiusculus gonez Jordan and Starks, 1904 [Japanese name: Oki-himekajika]
Cottiusculus gonez: Okada and Matsubara, 1938; Mori, 1952; Matsubara, 1955; Kato, 1956;

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Watanabe, 1960; Takegawa and Morino, 1970; Ueno, 1971; Honma, 1977; Honma and Kitami,
1978; Yabe, 1988; Nakabo, 1993b; Amaoka et al., 1995; Yabe, 1997; Adrianov and Kussakin,1998; Sokolovskaya et al., 1998; Nakabo, 2000b, 2002b; Maeda and Tsutsui, 2003; Sideleva etal., 2006b; Honma, 2013; Nakabo and Kai, 2013.
Cottiusculus gonez gonez: Tsuda, 1990.
KT-11-09-N2 (3), KT-11-09-T6 (1).
Cottiusculus nihonkaiensis Kai and Nakabo, 2009 [Japanese name: Nihon-kin-kajika]
Cottiusculus schmidti (not of Jordan and Starks, 1904): Kato, 1956; Watanabe, 1960; Ouchi,
1963; Ueno, 1971; Yabe, 1988; Tsuda, 1990; Adrianov and Kussakin, 1998; Sokolovskaya et al.,
1998; Maeda and Tsutsui, 2003; Kitagawa et al., 2008.
Cottiusculus schmidtii (not of Jordan and Starks, 1904): Amaoka et al., 1995 (in part).
Cottiusculus nihonkaiensis: Kai and Nakabo, 2009 (original description); Honma, 2013.
TS09-T088 (1), TS09-T107 (76), SMBS uncatalogued (2).
Enophrys diceraus (Pallas, 1787) [Japanese name: Oni-kajika]
Ceratocottus namiyæ: Snyder, 1912.
Ceratocottus namiyei: Okada and Matsubara, 1938.
Ceratocottus diceraus namiyei: Kato, 1956.
Ceratocottus diceraus: Watanabe, 1960.
Enophrys diceraus: Ueno, 1971; Tsuda, 1990; Amaoka et al., 1995; Yabe, 1997; Adrianov
and Kussakin, 1998; Sokolovskaya et al., 1998; Maeda and Tsutsui, 2003; Sideleva et al., 2006a;
Kitagawa et al., 2008; Nakabo and Kai, 2013.
Enophrys diceratus [sic]: Honma, 2013.
Gymnocanthus detrisus Gilbert and Burke, 1912 [Japanese name: Sebiro-kajika]
Gymnocanthus detrisus: Okada and Matsubara, 1938; Matsubara, 1955; Ueno, 1971; Adrianov and Kussakin, 1998; Sokolovskaya et al., 1998; Sideleva et al., 2006a; Nakabo and Kai,
2013.
*Gymnocanthus galeatus Bean, 1881 [Japanese name: Chikame-kajika]
Gymnocanthus galeatus: Nakabo and Kai, 2013.
Gymnocanthus herzensteini Jordan and Starks, 1904 [Japanese name: Tsumaguro-kajika]
Gymnocanthus herzensteini: Snyder, 1912; Hikita, 1950; Kato, 1956; Watanabe, 1960;
Ouchi, 1963; Ueno, 1971; Yabe, 1988; Tsuda, 1990; Amaoka et al., 1995; Adrianov and Kussakin, 1998; Sokolovskaya et al., 1998; Maeda and Tsutsui, 2003; Sideleva et al., 2006a; Kitagawa
et al., 2008; Honma, 2013.
Gymnocanthus intermedius (Temminck and Schlegel, 1843) [Japanese name: Ai-kajika]
Gymnocanthus ventralis: Mori, 1928; Okada and Matsubara, 1938; Mori, 1952; Matsubara,
1955; Kato, 1956; Ueno, 1971; Honma, 1977; Honma and Kitami, 1978; Tsuda, 1990.
Gymnocanthus intermedius: Mori and Uchida, 1934; Watanabe, 1960; Nakabo, 1993b; Amaoka et al., 1995; Uozu Aquarium, 1997; Yabe, 1997; Adrianov and Kussakin, 1998; Sokolovskaya et al., 1998; Nakabo, 2000b, 2002b; Sideleva et al., 2006a; Kitagawa et al., 2008; Matsuura et al., 2009; Honma, 2013; Nakabo and Kai, 2013.
Gymnocanthus pistilliger (Pallas, 1814) [Japanese name: Hage-kajika]
Gymnocanthus pistilliger: Matsubara, 1955; Kato, 1956; Ueno, 1971; Honma and Kitami,
1978; Tsuda, 1990; Nakabo, 1993b; Amaoka et al., 1995; Adrianov and Kussakin, 1998; Sokolovskaya et al., 1998; Nakabo, 2000b, 2002b; Maeda and Tsutsui, 2003; Sideleva et al., 2006a;
Kitagawa et al., 2008; Matsuura et al., 2009; Honma, 2013; Nakabo and Kai, 2013.
Hemilepidotus gilberti Jordan and Starks, 1904 [Japanese name: Yokosujikajika]

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Hemilepidotus gilberti: Okada and Matsubara, 1938; Ueno, 1971; Tsuda, 1990; Amaoka etal., 1995; Adrianov and Kussakin, 1998; Sokolovskaya et al., 1998; Maeda and Tsutsui, 2003;
Sideleva et al., 2006a; Nakabo and Kai, 2013.
*Hemilepidotus papilio (Bean, 1880) [Japanese name: Kujaku-kajika]
Melletes paciﬁcus: Ueno, 1971.
Hemilepidotus papilio: Nakabo and Kai, 2013.
Icelus cataphractus (Pavlenko, 1910) [Japanese name: Kōrikajika]
Icelus spiniger cataphractus: Kato, 1956; Mori, 1956a; Honma, 1963; Ouchi, 1963; Takegawa and Morino, 1970; Honma, 1977; Honma and Kitami, 1978; Lindberg and Krasyukova,
1987; Adrianov and Kussakin, 1998; Sideleva et al., 2006a.
Icelus spiniger: Katayama, 1940; Hikita, 1950; Honma, 1952; Mori, 1952; Kato, 1956;
Oshima, 1957; Watanabe, 1960; Ouchi, 1963; Ueno, 1971; Tsuda, 1990.
Icelus cataphract [sic]: Ueno, 1971.
Icelus cataphractus: Nakabo, 1993b; Amaoka et al., 1995; Yabe, 1997; Sokolovskaya et al.,
1998; Suzuki et al., 2000; Nakabo, 2000b, 2002b; Maeda and Tsutsui, 2003; Kitagawa et al.,2008; Matsuura et al., 2009; Shinohara et al., 2011; Honma, 2013; Nakabo and Kai, 2013.
KT-10-08-R1 (20).
Icelus gilberti Taranetz, 1936 [Japanese name: Daruma-kōrikajika]
Icelus gilberti: Taranetz, 1936 (original description); Ueno, 1971; Yabe, 1988; Sokolovskaya
et al., 1998; Sideleva et al., 2006a.
Icelus rastrinoides Taranetz, 1936 [Japanese name: Nise-matsu-kajika]
Icelus rastrinoides: Taranetz, 1936 (original description); Matsubara, 1955; Ueno, 1971;
Sokolovskaya et al., 1998; Siderleva et al., 2006a.
Icelus stenosomus Andriashev, 1937 [Japanese name: Yase-kōri-kajika]
Icelus uncinalis: Okada and Matsubara, 1938; Mori, 1952; Ueno, 1971.
Icelus uncinalis stenosomus: Kato, 1956; Honma, 1957a, 1963a; Ouchi, 1963; Honma andKitami, 1970, 1978; Adrianov and Kussakin, 1998; Sokolovskaya et al., 1998; Sideleva et al.,2006a.
Icelus stenosomus: Nakabo, 1993b; Murai et al., 1994; Nakabo, 2000b, 2002b; Honma,
2013; Nakabo and Kai, 2013.
Icelus toyamensis (Matsubara and Iwai, 1951) [Japanese name: Tomi-kajika] (Fig. 4E)
Ricuzenius (Alloricuzenius) toyamensis: Matsubara and Iwai, 1951a; Watanabe, 1960.
Ricuzenius toyamensis: Matsubara, 1955; Honma, 1957a, 1963; Ouchi, 1963; Honma, 1977;
Honma and Kitami, 1978; Lindberg and Krasyukova, 1987; Yabe, 1988; Tsuda, 1990; Uozu
Aquarium, 1997; Sideleva et al., 2006a.
Icelus toyamensis: Nakabo, 1993b, 2000b, 2002b; Shinohara et al., 2011; Honma, 2013;
Nakabo and Kai, 2013.
SMBS uncatalogued (1).
Myoxocephalus jaok (Cuvier, 1829) [Japanese name: Oku-kajika]
Myxocephalus edomius: Kato, 1956.
Myoxocephalus jaok: Watanabe, 1960; Yabe, 1988; Tsuda, 1990; Amaoka et al., 1995; Adrianov and Kussakin, 1998; Sokolovskaya et al., 1998; Maeda and Tsutsui, 2003; Sideleva et al.,2006a; Kitagawa et al., 2008; Honma, 2013; Nakabo and Kai, 2013.
*Myoxocephalus niger (Bean, 1881) [Japanese name: Ke-kajika]
Myoxocephalus niger: Watanabe, 1960; Ueno, 1971; Yabe, 1988; Adrianov and Kussakin,1998; Sokolovskaya et al., 1998.

Annotated Checklist of Deep-sea Fishes of the Sea of Japan251

Myoxocephalus polyacanthocephalus (Pallas, 1814) [Japanese name: Toge-kajika]
Myoxocephalus polyacanthocephalus: Snyder, 1912; Watanabe, 1960; Ueno, 1971; Yabe,
1988; Tsuda, 1990; Amaoka et al., 1995; Adrianov and Kussakin, 1998; Sokolovskaya et al.,
1998; Maeda and Tsutsui, 2003; Sideleva et al., 2006a; Kitagawa et al., 2008; Matsuura et al.,2009; Honma, 2013; Nakabo and Kai, 2013.
Ainocottus ensiger: Hikita, 1950; Kato, 1956; Ueno, 1971; Tsuda, 1990.
*Ricuzenius pinetorum Jordan and Starks, 1904 [Japanese name: Matsu-kajika]
Ricuzenius pinetorum: Watanabe, 1960; Ueno, 1971; Yabe, 1988; Tsuda, 1990; Maeda andTsutsui, 2003; Kitagawa et al., 2008; Honma, 2013; Nakabo and Kai, 2013.
Stelgistrum stejnegeri Jordan and Gilbert, 1898 [Japanese name: Ottosei-kajika]
Stelgistrum stejnegeri: Okada and Matsubara, 1938; Ueno, 1971; Nakabo and Kai, 2013.
Stelgistrum steinegeri [sic]: Adrianov and Kussakin, 1998; Sokolovskaya et al., 1998; Sideleva et al., 2006a.
Taurocottus bergii Soldatov and Pavlenko, 1915 [Japanese name: Kiri-kajika]
Taurocottus bergi: Okada and Matsubara, 1938; Ueno, 1971; Yabe, 1988; Amaoka et al.,
1995; Sokolovskaya et al., 1998; Maeda and Tsutsui, 2003; Sideleva et al., 2006a.
Taurocottus bergii: Nakabo and Kai, 2013.
KT-10-08-R1 (3).
Triglops jordani (Jordan and Starks, 1904) [Japanese name: Karafuto-kajika]
Triglops jordani: Okada and Matsubara, 1938; Matsubara, 1955; Ueno, 1971; Lindberg andKrasyukova, 1987; Tsuda, 1990; Pietsch, 1993; Nakabo, 1993b; Amaoka et al., 1995; Adrianov
and Kussakin, 1998; Sokolovskaya et al., 1998; Nakabo, 2000b, 2002b; Sideleva et al., 2006a;
Shinohara et al., 2011; Nakabo and Kai, 2013.
Triglops pingelii Reinhardt, 1837 [Japanese name: Hokkyoku-kajika]
Triglops beani: Okada and Matsubara, 1938; Honma, 1952; Mori, 1952; Ueno, 1971; Tsuda,
1990.
Triglops pingelii beani: Matsubara, 1955; Honma, 1963.
Triglops pingeli beani: Kato, 1956; Honma and Kitami, 1978.
Triglops pingeli: Lindberg and Krasyukova, 1987; Amaoka et al., 1995; Adrianov and Kussakin, 1998; Sokolovskaya et al., 1998; Pietsch, 1993; Nakabo, 1993b.
Triglops pingelii: Sideleva et al., 2006a; Shinohara et al., 2011; Nakabo and Kai, 2013.
Triglops pingelii pingelii: Nakabo, 2000b, 2002b; Maeda and Tsutsui, 2003.
Triglops pingeli pingeli: Honma, 2013.
Triglops scepticus Gilbert, 1896 [Japanese name: Nirami-kajika]
Triglops scepticus: Okada and Matsubara, 1938; Honma, 1952; Mori, 1952; Matsubara,
1955; Kato, 1956; Mori, 1956a, c; Honma, 1963; Ouchi, 1963; Takegawa and Morino, 1970;
Ueno, 1971; Ogata et al., 1973; Honma, 1977; Honma and Kitami, 1978; Lindberg and Krasyukova, 1987; Tsuda, 1990; Nambu et al., 1992; Pietsch, 1993; Nakabo, 1993b; Amaoka et al.,
1995; Adrianov and Kussakin, 1998; Sokolovskaya et al., 1998; Nakabo, 2000b, 2002b; Maeda
and Tsutsui, 2003; Sideleva et al., 2006a; Shinohara et al., 2011; Honma, 2013; Nakabo and Kai,
2013.
Triglops uchidai: Watanabe, 1960.
KT-10-08-R1 (15), SMBS uncatalogued (2).

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Family Hemitripteridae
Hemitripterus villosus (Pallas, 1814) [Japanese name: Kemushi-kajika]
Hemitripterus villosus: Okada and Matsubara, 1938; Mori and Uchida, 1934; Katayama,
1940; Honma, 1952; Katayama, 1952; Mori, 1952; Matsubara, 1955; Kato, 1956; Mori, 1956a;
Honma, 1963; Honma and Kitami, 1970; Takegawa and Morino, 1970; Ueno, 1971; Honma,
1977; Honma and Kitami, 1978; Nakabo, 1993b; Amaoka et al., 1995; Uozu Aquarium, 1997;
Yabe, 1997; Adrianov and Kussakin, 1998; Sokolovskaya et al., 1998; Nakabo, 2000b; Suzuki etal., 2000; Nakabo, 2002b; Maeda and Tsutsui, 2003; Mecklenburg, 2003; Sideleva et al., 2006b;
Kitagawa et al., 2008; Matsuura et al., 2009; Shinohara et al., 2011; Honma, 2013; Nakabo andKai, 2013.
Hemitripterus (americanus) villosus: Tsuda, 1990.
Nautichthys pribilovius (Jordan and Gilbert, 1898) [Japanese name: Okoze-kajika]
Nautichthys oculofasciatus: Okada and Matsubara, 1938; Matsubara, 1955; Ueno, 1971.
Nautichthys pribilovius: Adrianov and Kussakin, 1998; Maeda and Tsutsui, 2003; Mecklenburg, 2003; Balanov et al., 2006; Sideleva et al., 2006a; Nakabo and Kai, 2013.
Family Agonidae
Anoplagonus occidentalis Lindberg, 1950 [Japanese name: Nise-name-tokubire]
Anoplagonus occidentalis: Ueno, 1971; Kanayama and Maruyama, 1979; Kanayama, 1991;
Amaoka et al., 1995; Adrianov and Kussakin, 1998; Sokolovskaya et al., 1998; Kitagawa et al.,2008; Sheiko and Fedorov, 2010; Honma, 2013; Nakabo and Kai, 2013.
*Aspidophoroides monopterygius (Bloch, 1786) [Japanese name: Tate-tokubire]
Aspidophoroides bartoni: Ueno, 1971; Tsuda, 1990; Adrianov and Kussakin, 1998; Sokolovskaya et al., 1998.
Aspidophoroides monopterygius: Kanayama, 1991; Nakabo, 1993b; Amaoka et al., 1995;
Kitagawa et al., 2008; Maeda and Tsutsui, 2003.
Freemanichthys thompsoni (Jordan and Gilbert, 1898) [Japanese name: Yase-tokubire]
Podothecus thompsoni: Mori, 1928; Mori and Uchida, 1934; Okada and Matsubara, 1938;
Katayama, 1940; Honma, 1952; Katayama, 1952; Mori, 1952; Matsubara, 1955; Kato, 1956;
Mori, 1956a; Ouchi, 1963; Ueno, 1971; Lindberg and Krasyukova, 1987; Tsuda, 1990; Uozu
Aquarium, 1997; Adrianov and Kussakin, 1998; Sokolovskaya et al., 1998.
Freemanichthys thompsoni: Kanayama, 1991; Nakabo, 1993b; Amaoka et al., 1995; Nakabo,
2000b, 2002b; Maeda and Tsutsui, 2003; Sheiko and Fedorov, 2010; Shinohara et al., 2011;
Nakabo and Kai, 2013.
Freemanichthys tompsoni [sic]: Honma, 2013.
KT-10-08-R1 (2).
Hypsagonus quadricornis (Valenciennes, 1829) [Japanese name: Tsuno-shachi-uo]
Hypsagonus quadricornis corniger: Taranetz, 1933.
Hypsagonus quadricornis: Honma, 1955d; Kato, 1956; Honma, 1963; Ueno, 1971; Honma,
1977; Honma and Kitami, 1978; Tsuda, 1990; Kanayama, 1991; Nakabo, 1993b; Sheiko, 1993;
Amaoka et al., 1995; Nakabo, 2000b, 2002b; Maeda and Tsutsui, 2003; Shinohara et al., 2011;
Honma, 2013; Nakabo and Kai, 2013.
*Leptagonus frenatus (Gilbert, 1896) [Japanese name: Yase-tengu-tokubire]
Sarritor frenatus: Kato, 1956.
Leptagonus frenatus: Oshima, 1957; Kanayama, 1991; Nakabo, 1993b; Amaoka et al., 1995;

Annotated Checklist of Deep-sea Fishes of the Sea of Japan253

Nakabo, 2000b, 2002b; Nakabo and Kai, 2013.
Leptogonus frenatus [sic]: Honma, 2013.
Leptagonus leptorhynchus (Gilbert, 1896) [Japanese name: Tengu-tokubire]
Sarritor leptorhynchus knipowitschi: Lindberg and Andriashev, 1937; Honma, 1963; Takegawa and Morino, 1970; Lindberg and Krasyukova, 1987.
Sarritor leptorhynchus: Katayama, 1940; Honma, 1952; Kato, 1956; Oshima, 1957; Mori,
1956a; Ouchi, 1963; Ueno, 1971; Uozu Aquarium, 1997.
Saritor leptorhynchus [sic]: Katayama, 1952.
Sarritor leptorhynchus leptorhynchus: Matsubara, 1955; Honma, 1977; Tsuda, 1990.
Leptagonus leptorhynchus: Kanayama, 1991; Nakabo, 1993b; Amaoka et al., 1995; Sokolovskaya et al., 1998; Nakabo, 2000b, 2002b; Kitagawa et al., 2008; Sheiko and Fedorov, 2010;
Shinohara et al., 2011; Nakabo and Kai, 2013.
Sarritor knipowitschi: Ueno, 1971; Sheiko and Fedorov, 2010.
Leptogonus leptorhynchus [sic]: Honma, 2013.
TS09-T117 (3).
*Occella iburia (Jordan and Starks, 1904) [Japanese name: Saburō]
Occa iburia: Kato, 1956; Tsuda, 1990; Honma, 2013.
Occella kuronumai (Freeman, 1951) [Japanese name: Shirō]
Occa kuronumai: Freeman, 1951 (original description); Honma and Kitami, 1970; Honma etal., 1972; Honma and Kitami, 1978; Tsuda, 1990.
Occella kuronumai: Kanayama, 1991; Nakabo, 1993b; Uozu Aquarium, 1997; Nakabo,
2000b, 2002b; Sheiko and Fedorov, 2010; Nakabo and Kai, 2013.
Percis japonica (Pallas, 1769) [Japanese name: Inugochi]
Percis japonica: Tanaka, 1927; Mori, 1928; Mori and Uchida, 1934; Okada and Matsubara,
1938; Katayama, 1940; Honma, 1952; Katayama, 1952; Mori, 1952; Matsubara, 1955; Kato,
1956; Honma, 1963; Ueno, 1971; Honma and Kitami, 1978; Tsuda, 1990; Maeda and Tsutsui,
2003; Sheiko and Fedorov, 2010; Nakabo and Kai, 2013.
Percis japonicus: Kanayama, 1991; Nakabo, 1993b; Amaoka et al., 1995; Adrianov and Kussakin, 1998; Sokolovskaya et al., 1998; Nakabo, 2000b, 2002b; Kitagawa et al., 2008; Shinohara
et al., 2011; Honma, 2013.
Podothecus sachi (Jordan and Snyder, 1901) [Japanese name: Tokubire]
Podothecus xystes: Snyder, 1911 (original description); Okada and Matsubara, 1938; Kato,
1956.
Podothecus sachi: Hikita, 1950; Kato, 1956; Ueno, 1971; Tsuda, 1990; Kanayama, 1991;
Nakabo, 1993b; Amaoka et al., 1995; Adrianov and Kussakin, 1998; Sokolovskaya et al., 1998;
Nakabo, 2000b, 2002b; Maeda and Tsutsui, 2003; Kitagawa et al., 2008; Sheiko and Fedorov,
2010; Honma, 2013; Nakabo and Kai, 2013.
SMBS uncatalogued (2).
Podothecus sturioides (Guichenot, 1869) [Japanese name: Same-tokubire]
Podothecus gilberti: Kato, 1956; Ueno, 1971; Adrianov and Kussakin, 1998.
Podothecus sturioides: Nakabo, 1993b, Sokolovskaya et al., 1998; Nakabo, 2000b, 2002b;
Sheiko and Fedorov, 2010; Honma, 2013; Nakabo and Kai, 2013.
Podothecus veternus Jordan and Starks, 1895 [Japanese name: Onaga-tokubire]
Podothecus veternus: Kanayama, 1991; Adrianov and Kussakin, 1998; Sokolovskaya et al.,
1998; Sheiko and Fedorov, 2010.
Tilesina gibbosa Schmidt, 1904 [Japanese name: Oni-shachi-uo]

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Tilesina gibbosa: Okada and Matsubara, 1938; Kato, 1956; Ueno, 1971; Tsuda, 1990;
Kanayama, 1991; Nakabo, 1993b; Amaoka et al., 1995; Sokolovskaya et al., 1998; Nakabo,
2000b, 2002b; Maeda and Tsutsui, 2003; Kitagawa et al., 2008; Sheiko and Fedorov, 2010;
Honma, 2013; Nakabo and Kai, 2013.
Tilesina hubbsi: Freeman, 1951 (original description).
Family Psychrolutidae
Dasycottus setiger Bean, 1890 [Japanese name: Ganko]
Dasycottus setiger: Jordan and Evermann, 1898; Snyder, 1912; Soldatov and Lindberg,
1930; Schmidt, 1950; Oshima, 1957; Watanabe, 1960; Ueno, 1971; Yabe, 1988; Lindberg andKrasyukova, 1987; Tsuda, 1990; Nakabo, 1993b; Amaoka et al., 1995; Yabe, 1997; Sokolovskaya et al., 1998; Nakabo, 2000b; Suzuki et al., 2000; Nakabo, 2002b; Maeda and Tsutsui, 2003;
Kitagawa et al., 2008; Shinohara et al., 2011; Honma, 2013; Nakabo and Kai, 2013.
Dasycottus japonicus: Tanaka, 1914b; Okada and Matsubara, 1938; Katayama, 1940; Hikita,
1950; Honma, 1952; Katayama, 1952; Mori, 1952; Matsubara, 1955; Kato, 1956; Mori, 1956a;
Honma, 1963; Ouchi, 1963; Honma and Kitami, 1970; Ueno, 1971; Ogata et al., 1973; Honma,
1977; Honma and Kitami, 1978; Lindberg and Krasyukova, 1987.
Dasycottus sp.: Mori, 1952.
Dasycottus uchidai: Mori, 1955 (original description); Kato, 1956; Mori, 1956a, c; Takegawa
and Morino, 1970.
Dasycottus longidorosalis [sic]: Kato, 1956.
Dasycottus longipinnis: Mori, 1956a (original description).
TS09-T001 (3), TS09-T056 (1).
*Eurymen gyrinus Gilbert and Burke, 1912 [Japanese name: Yagishiri-kajika]
Eurymen bassargini: Soldatov and Lindberg, 1930 (original description); Okada and Matsubara, 1938, 1955; Watanabe, 1960; Adrianov and Kussakin, 1998.
Eurymen gyrinus: Okada and Matsubara, 1938; Yabe, 1988; Nakabo, 1993b; Amaoka et al.,
1995; Adrianov and Kussakin, 1998; Sokolovskaya et al., 1998; Nakabo, 2000b, 2002b; Maeda
and Tsutsui, 2003; Nakabo and Kai, 2013.
Gilbertidia ochotensis: Watanabe, 1960.
Eurymen gurinus [sic]: Ueno, 1971.
Malacocottus gibber Sakamoto, 1930 [Japanese name: Yamato-kobushi-kajika]
Malacocottus gibber: Sakamoto, 1930 (original description); Okada and Matsubara, 1938;
Katayama, 1940; Matsubara, 1955; Honma, 1955d; Kato, 1956; Mori, 1956a; Watanabe, 1960;
Honma, 1963; Ouchi, 1963; Takegawa and Morino, 1970; Ueno, 1971; Ogata et al., 1973;
Honma and Kitami, 1978; Tsuda, 1990; Nambu et al., 1992; Nakabo, 1993b; Shinohara et al.,
1992; Amaoka et al., 1995; Nakabo, 2000b; Suzuki et al., 2000; Nakabo, 2002b; Maeda and Tsutsui, 2003; Shinohara et al., 2011; Honma, 2013; Nakabo and Kai, 2013.
KT-11-09-T1 (1), TS09-T004 (1), TS09-T006 (2), TS09-T007 (1), TS09-T056 (1), SMBS
uncatalogued (3).
*Malacocottus zonurus Bean, 1890 [Japanese name: Kobushi-kajika]
Malacocottus zonurus: Matsubara, 1955; Tsuda, 1990; Nakabo, 1993b; Adrianov and Kussakin, 1998; Sokolovskaya et al., 1998; Nakabo and Kai, 2013.
Malacocottus zonurus microphthalmus: Mori, 1956c.
Psychrolutes paradoxus Günther, 1861 [Japanese name: Uranai-kajika]

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Psychrolutes paradoxus: Mori, 1952; Matsubara, 1955; Honma, 1957a, 1963; Takegawa andMorino, 1970; Ueno, 1971; Honma, 1977; Honma and Kitami, 1978; Yabe, 1988; Tsuda, 1990;
Nakabo, 1993b; Amaoka et al., 1995; Adrianov and Kussakin, 1998; Sokolovskaya et al., 1998;
Nakabo, 2000b, 2002b; Maeda and Tsutsui, 2003; Shinohara et al., 2011; Honma, 2013; Nakabo
and Kai, 2013.
TS09-T117 (3).
Family Cyclopteridae
Aptocyclus ventricosus (Pallas, 1769) [Japanese name: Hotei-uo]
Cyclopterichthys ventricosus: Katayama, 1940; Hikita, 1950; Honma, 1952; Oshima, 1957.
Aptocyclus ventricosus: Mori and Uchida, 1934; Mori, 1952; Matsubara, 1955; Kato, 1956;
Mori, 1956a; Honma, 1957b, 1963; Takegawa and Morino, 1970; Ueno, 1970, 1971; Ogata etal., 1973; Honma, 1977; Honma and Kitami, 1978; Lindberg and Krasyukova, 1987; Tsuda,
1990; Nambu et al., 1992; Nakabo, 1993b; Kido and Shinohara, 1996; Amaoka et al., 1995;
Uozu Aquarium, 1997; Adrianov and Kussakin, 1998; Sokolovskaya et al., 1998; Nakabo,
2000b; Suzuki et al., 2000; Nakabo, 2002b; Maeda and Tsutsui, 2003; Kitagawa et al., 2008;
Shinohara et al., 2011; Honma, 2013; Nakabo and Kai, 2013.
Pelagocyclus vitiazi: Ueno, 1970; Lindberg and Krasyukova, 1987; Kido, 1988b; Nakabo,
1993b.
TS09-T032 (1), TS09-T115 (1), SMBS uncatalogued (1).
*Cyclopterus bergi Popov, 1929 [Japanese name: Kobu-fūsen-uo]
Cyclopterus bergi: Ueno, 1970, 1971; Kido, 1988b; Nakabo, 1993b; Sokolovskaya et al.,
1998; Nakabo, 2000b, 2002b; Nakabo and Kai, 2013.
*Cyclopteropsis lindbergi Soldatov, 1930 [Japanese name: Name-fūsen-uo]
Cyclopteropsis lindbergi: Ueno, 1970, 1971; Kido, 1988b; Tsuda, 1990; Nakabo, 1993b;
Adrianov and Kussakin, 1998; Sokolovskaya et al., 1998; Nakabo, 2000b, 2002b; Honma, 2013;
Nakabo and Kai, 2013.
Eumicrotremus birulai Popov, 1925 [Japanese name: Konpeitō]
Cyclolumpus asperrimus: Tanaka, 1912 (original description); Okada and Matsubara, 1938;
Katayama, 1940; Honma, 1952; Katayama, 1952; Mori, 1952; Matsubara, 1955; Oshima, 1957;
Ouchi, 1963; Takegawa and Morino, 1970; Honma, 1977.
Cyclolumpus osperrimum [sic]: Kato, 1956.
Cyclolumpus birulai: Kato, 1956; Oshima, 1957.
Eumicrotremus birulai: Honma, 1963; Ueno, 1970, 1971; Honma and Kitami, 1978; Kido,
1988b; Nakabo, 1993b; Amaoka et al., 1995; Kido, 1997; Uozu Aquarium, 1997; Adrianov andKussakin, 1998; Nakabo, 2000b; Suzuki et al., 2000; Nakabo, 2002b; Maeda and Tsutsui, 2003;
Shinohara et al., 2011; Nakabo and Kai, 2013.
Euprotomicrus birulai [sic]: Ogata et al., 1973.
Eumicrotremus birlai [sic]: Honma, 2013.
TS09-T001 (2), TS09-T006 (18), TS09-T010 (15), TS09-T010 (15), TS09-T012 (14), TS09T102 (1), TS09-T106 (3), SMBS uncatalogued (1).
*Eumicrotremus paciﬁcus Schmidt, 1904 [Japanese name: Fūsen-uo]
Eumicrotremus paciﬁcus: Kato, 1956; Ueno, 1970, 1971; Tsuda, 1990; Amaoka et al., 1995;
Nakabo, 1993b, Adrianov and Kussakin, 1998; Sokolovskaya et al., 1998; Nakabo, 2000b,
2002b; Nakabo and Kai, 2013.

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Gento Shinohara, Masanori Nakae, Yuji Ueda, Shigeaki Kojima and Keiichi Matsuura*Eumicrotremus taranetzi Perminov, 1936 [Japanese name: Name-dango]
Eumicrotremus taranetzi: Ueno, 1970; Kido, 1988b; Amaoka et al., 1995; Nakabo, 1993b,
2000b, 2002b; Kitagawa et al., 2008; Nakabo and Kai, 2013.
Family Liparidae
Careproctus acanthodes Gilbert and Burke, 1912 [Japanese name: Toge-bikunin]
Careproctus acanthodes: Okada and Matsubara, 1938; Ueno, 1971; Kido, 1988b; Sokolovskaya et al., 1998; Chernova et al., 2004; Shinohara et al., 2011.
Careproctus batialis Popov, 1933 [Japanese name: Dattan-bukunin]
Careproctus batialis: Popov, 1933 (original description); Ueno, 1971; Pitruk, 1991; Chernova, 1998; Sokolovskaya et al., 1998; Chernova et al., 2004.
Careproctus colletti Gilbert, 1896 [Japanese name: Arasuka-bikunin]
Careproctus colletti: Okada and Matsubara, 1938; Honma, 1952; Kato, 1956; Mori, 1956a;
Lindberg and Krasyukova, 1987; Kido, 1988a, 1988b; Nambu and Kido, 1990; Tsuda, 1990;
Nakabo, 1993b; Amaoka et al., 1995; Kido, 1997; Nakabo, 2000b, 2002b; Maeda and Tsutsui,
2003; Chernova et al., 2004; Shinohara et al., 2011; Honma, 2013; Nakabo and Kai, 2013.
Careproctus puniceus: Mori, 1955 (original description); Kato, 1956; Takegawa and Morino,
1970.
Careproctus entomelas: Mori, 1956a; Tsuda, 1990.
KT-11-09-M6 (1).
Careproctus entargyreus Gilbert and Burke, 1912 [Japanese name: Sokoshiro]
Careproctus entargyreus: Ueno, 1971; Kido, 1988b; Chernova et al., 2004; Shinohara et al.,2011.
Careproctus entomelas Gilbert and Burke, 1912 [Japanese name: Sokoguro]
Careproctus entomelas: Burke, 1930; Okada and Matsubara, 1938; Kato, 1956; Kido, 1988b;
Tsuda, 1990; Chernova et al., 2004; Shinohara et al., 2011.
*Careproctus furcatus Mori, 1956 [Japanese name: Oki-bikunin]
Careproctus furcatus: Kato, 1956; Mori, 1956d (original descrpition, type lost); Chernova etal., 2004.
Careproctus notosaikaiensis Kai, Ikeguchi and Nakabo, 2011 [Japanese name: Noto-saikaibikunin] (Fig. 4A)
Careproctus notosaikaiensis: Kai et al., 2011 (original description); Nakabo and Kai, 2013.
Careproctus rastrinus Gilbert and Burke, 1912 [Japanese name: Sake-bikunin]
Careproctus pellucidus: Hikita, 1950; Katayama, 1952; Kato, 1956; Oshima, 1957; Takegawa and Morino, 1970; Ueno, 1971; Tsuda, 1990; Chernova et al., 2004.
Careproctus rastrinus: Kato, 1956; Ueno, 1971; Lindberg and Krasyukova, 1987; Kido,
1988a; Tsuda, 1990; Nakabo, 1993b; Amaoka et al., 1995; Kido, 1997; Nakabo, 2000b; Suzuki
et al., 2000; Nakabo, 2002b; Maeda and Tsutsui, 2003; Chernova et al., 2004; Kitagawa et al.,2008.
Careproctus rastrinus species complex: Honma, 2013; Nakabo and Kai, 2013.
*Careproctus rhodomelas Gilbert and Burke, 1912 [Japanese name: Bara-bikunin]
Careproctus rhodomelas: Kato, 1956; Tsuda, 1990.
*Careproctus roseofuscus Gilbert and Burke, 1912 [Japanese name: Tobi-bikunin]
Careproctus roseofuscus: Maeda and Tsutsui, 2003.
*Careproctus segaliensis Gilbert and Burke, 1912 [Japanese name: Karafuto-bikunin]

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Careproctus segaliensis: Katayama, 1940; Matsubara, 1955; Kato, 1956; Takegawa andMorino, 1970; Ueno, 1971; Lindberg and Krasyukova, 1987; Tsuda, 1990; Chernova et al., 2004.
Careproctus sinensis Gilbert and Burke, 1912 [Japanese name: Sekichiku-bikunin]
Careproctus sinensis: Gilbert and Burke, 1912 (original description); Burke, 1930; Okada
and Matsubara, 1938; Honma, 1952; Matsubara, 1955; Kato, 1956; Mori, 1956a; Oshima, 1957;
Ouchi, 1963; Takegawa and Morino, 1970; Lindberg and Krasyukova, 1987; Kido, 1988a,
1988b; Nambu and Kido, 1990; Tsuda, 1990; Nakabo, 1993b, 2000b, 2002b; Chernova et al.,
2004; Shinohara et al., 2011; Honma, 2013; Nakabo and Kai, 2013.
Careproctus trachysoma Gilbert and Burke, 1912 [Japanese name: Zara-bikunin]
Careproctus trachysoma: Gilbert and Burke, 1912 (original despcription); Burke, 1930;
Okada and Matsubara, 1938; Honma, 1952; Matsubara, 1955; Kato, 1956; Honma, 1963; Ueno,
1971; Honma and Kitami, 1978; Lindberg and Krasyukova, 1987; Kido, 1988a, b; Nambu andKido, 1990; Tsuda, 1990; Nambu et al., 1992; Nakabo, 1993b; Amaoka et al., 1995; Kido, 1997;
Nakabo, 2000b, 2002b; Maeda and Tsutsui, 2003; Chernova et al., 2004; Shinohara et al., 2011.
Careproctus rastrinus species complex: Nakabo and Kai, 2013.
KT-11-09-K4 (1), KT-11-09-N4 (2), KT-11-09-T3 (1), KT-11-09-T5 (1), TS09-T004 (2),TS09-T006 (1), TS09-T016 (2), TS09-T032 (2).
Crystallias matsushimae Jordan and Snyder, 1902 [Japanese name: Abachan]
Crystallias matsushimae: Burke, 1930; Okada and Matsubara, 1938; Katayama, 1940;
Hikita, 1950; Honma, 1952; Katayama, 1952; Mori, 1952; Matsubara, 1955; Mori, 1956a;
Oshima, 1957; Kato, 1956; Honma, 1963; Ouchi, 1963; Honma and Kitami, 1970; Takegawa andMorino, 1970; Ueno, 1971; Honma, 1977; Honma and Kitami, 1978; Kido, 1988b; Tsuda, 1990;
Uozu Aquarium, 1997; Adrianov and Kussakin, 1998; Sokolovskaya et al., 1998; Chernova etal., 2004.
Crystallias kamoae: Kato, 1956; Oshima, 1957 (original description).
Crystallichthys matsushimae: Kido, 1988a; Nambu and Kido, 1990; Nakabo, 1993b; Amaoka et al., 1995; Kido, 1997; Nakabo, 2000b; Suzuki et al., 2000; Nakabo, 2002b; Maeda andTsutsui, 2003; Kitagawa et al., 2008; Shinohara et al., 2011; Honma, 2013; Nakabo and Kai,
2013.
KT-11-09-E2 (1), KT-11-09-T2 (3). TS09-T37 (3), TS09-T088 (4).
Liparis frenatus (Gilbert and Burke, 1912) [Japanese name: Kanten-bikunin]
Cyclogaster frenatus: Gilbert and Burke, 1912 (original description).
Liparis frenatus: Okada and Matsubara, 1938; Ueno, 1971; Lindberg and Krasyukova, 1987;
Kido, 1988a, b; Nambu and Kido, 1990; Tsuda, 1990; Nakabo, 1993b; Amaoka et al., 1995;
Nakabo, 2000b, 2002b; Maeda and Tsutsui, 2003; Chernova et al., 2004; Shinohara et al., 2011;
Honma, 2013; Nakabo and Kai, 2013.
Liparis ingens (Gilbert and Burke, 1912) [Japanese name: Kōrai-bikunin]
Cyclogaster ingens: Gilbert and Burke, 1912 (original description).
Liparis ingens: Burke, 1930; Mori, 1952; Kato, 1956; Kido, 1988b; Tsuda, 1990; Adrianov
and Kussakin, 1998; Shinohara et al., 2011.
Liparis latifrons Schmidt, 1950 [Japanese name: Bōzu-kusa-uo]
Liparis latifrons meridionalis: Lindberg and Krasyukova, 1987; Chernova, 1998.
Liparis latifrons: Kido, 1988b; Nakabo, 1993b; Sokolovskaya et al., 1998; Nakabo, 2000b,
2002b; Chernova et al., 2004; Shinohara et al., 2011; Nakabo and Kai, 2013.
Liparis ochotensis Schmidt, 1904 [Japanese name: Isago-bikunin]
Liparis ochotensis: Mori, 1956c; Ueno, 1971; Lindberg and Krasyukova, 1987; Kido, 1988a;

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Nambu and Kido, 1990; Uozu Aquarium, 1997; Nambu et al., 1992; Nakabo, 1993b; Amaoka etal., 1995; Kido, 1997; Adrianov and Kussakin, 1998; Chernova, 1998; Sokolovskaya et al.,
1998; Nakabo, 2000b, 2002b; Chernova et al., 2004; Maeda and Tsutsui, 2003; Kitagawa et al.,2008; Shinohara et al., 2011; Nakabo and Kai, 2013.
Liparis rhodosoma: Takegawa and Morino, 1970; Ueno, 1971.
Liparis niger: Ogata et al., 1973; Adrianov and Kussakin, 1998.
TS09-T006 (1), TS09-T032 (1).
Liparis tanakae (Gilbert and Burke, 1912) [Japanese name: Kusa-uo]
Cyclogaster tanakae: Tanaka, 1914a.
Liparis tanakai: Okada and Matsubara, 1938; Kato, 1956; Ouchi, 1963; Ueno, 1971; Kido,
1988a; Nambu and Kido, 1990; Machida et al., 1991; Sokolovskaya et al., 1998; Maeda andTsutsui, 2003; Honma, 2013.
Liparis tanakae: Okada and Matsubara, 1938; Tsuda, 1990; Chernova et al., 2004;
Kobayashi et al., 2006; Nakabo and Kai, 2013.
TS09-T039 (1).
Liparis tessellatus (Gilbert and Burke, 1912) [Japanese name: Bikunin]
Liparis tessellatus: Burke, 1930; Okada and Matsubara, 1938; Katayama, 1940; Honma,
1952; Katayama, 1952; Mori, 1952; Matsubara and Iwai, 1954; Kato, 1956; Mori, 1956a, c;
Honma, 1963; Takegawa and Morino, 1970; Ueno, 1971; Honma and Kitami, 1978; Lindberg
and Krasyukova, 1987; Kido, 1988a, b; Nambu and Kido, 1990; Tsuda, 1990; Nakabo, 1993b;
Amaoka et al., 1995; Kido, 1997; Sokolovskaya et al., 1998; Nakabo, 2000b; Suzuki et al.,
2000; Nakabo, 2002b; Maeda and Tsutsui, 2003; Chernova et al., 2004; Kitagawa et al., 2008;
Shinohara et al., 2011; Honma, 2013; Nakabo and Kai, 2013.
Liparis brevicaudus: Mori, 1956c (original description).
KT-11-09-N1 (1), KT-11-09-T1 (1), TS09-T001 (1), TS09-T006 (7), TS09-T013 (3), TS09T046 (2), TS09-T049 (1), TS09-T101 (2), TS09-T102 (1), TS09-T105 (1), TS09-T106 (1).
Order Perciformes
Family Acropomatidae*Acropoma japonicum Günther, 1859 [Japanese name: Hotarujako]
Acropoma japonicum: Mori, 1952; Katayama, 1960; Lindberg and Krasyukova, 1969;
Takegawa and Morino, 1970.
Doderleinia berycoides (Hilgendorf, 1878) [Japanese name: Akamutsu]
Döderleinia berycoides: Mori and Uchida, 1934; Katayama, 1940; Honma, 1952; Mori,
1952; Matsubara, 1955; Kato, 1956; Mori, 1956a; Katayama, 1960; Takegawa and Morino, 1970.
Doderleinia berycoides: Katayama, 1960; Lindberg and Krasyukova, 1969; Tsuda, 1990;
Hatooka, 1993; Mochizuki, 1997; Hatooka, 2000, 2002; Kobayashi et al., 2006; Kitagawa et al.,2008; Shinohara et al., 2011; Hatooka, 2013; Honma, 2013.
*Malakichthys griseus Döderlein, 1883 [Japanese name: Ōme-hata]
Malakichthys griseus: Honma, 1952; Kato, 1956; Mori, 1956a; Katayama, 1960; Lindberg
and Krasyukova, 1969; Takegawa and Morino, 1970; Tsuda, 1990; Hatooka, 1993; Mochizuki,
1997; Hatooka, 2000, 2002, 2013; Honma, 2013.
*Malakichthys wakiyae Jordan and Hubbs, 1925 [Japanese name: Wakiya-hata]
Malakichthys wakiyai: Katayama, 1960; Lindberg and Krasyukova, 1969.
Malakichthys wakiyae: Hatooka, 2013.

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Synagrops japonicus (Döderlein, 1883) [Japanese name: Sumikui-uo]
Synagrops japonicus: Mori, 1952, 1956a; Katayama, 1960; Takegawa and Morino, 1970;
Tsuda, 1990; Suzuki et al., 2000; Hatooka, 2002; Shinohara et al., 2011; Hatooka, 2013.
Synayrops japonicus [sic]: Kato, 1956.
Family Polyprionidae
Stereolepis doederleini Lindberg and Krasyukova, 1969 [Japanese name: Ōkuchi-ishinagi]
Stereolepis ischinagi: Snyder, 1912; Mori, 1928; Okada and Matsubara, 1938; Katayama,
1940; Honma, 1952; Katayama, 1952; Mori, 1952; Matsubara, 1955; Kato, 1956; Mori, 1956a;
Katayama, 1960; Honma, 1963; Takegawa and Morino, 1970; Ueno, 1971.
Stereolepis doederleini: Lindberg and Krasyukova, 1969 (original description); Tsuda, 1990;
Hatooka, 1993; Amaoka et al., 1995; Uozu Aquarium, 1997; Sokolovskaya et al., 1998; Hatooka,
2000; Suzuki et al., 2000; Hatooka, 2002; Maeda and Tsutsui, 2003; Shinohara et al., 2011;
Hatooka and Kai, 2013.
Stereolepis ishinagi [sic]: Honma and Kitami, 1978.
Stereolepis döderleini: Honma, 2013.
*Stereolepis gigas Ayres, 1859 [Japanese name: Kokuchi-ishinagi]
Stereolepis gigas: Lindberg and Krasyukova, 1969; Hatooka, 1993; Sokolovskaya et al.,
1998; Hatooka, 2000, 2002; Hatooka and Kai, 2013.
Family Serranidae
Caprodon schlegelii (Günther, 1859) [Japanese name: Aka-isaki]
Caprodon schlegeli: Kato, 1956; Mori, 1956a; Katayama, 1960; Lindberg and Krasyukova,
1969.
Caprodon schlegelii: Mori, 1952; Tsuda, 1990; Suzuki et al., 2000; Shinohara et al., 2011;
Senou, 2013.
*Chelidoperca hirundinacea (Valenciennes, 1831) [Japanese name: Himekodai]
Chelidoperca undinacea: Uchida and Mori, 1934.
Chelidoperca hirundinacea: Mori, 1952; Kato, 1956; Mori, 1956a; Katayama, 1960; Lindberg and Krasyukova, 1969; Takegawa and Morino, 1970; Tsuda, 1990; Senou, 2013.
Epinephelus septemfasciatus (Thunberg, 1793) [Japanese name: Mahata]
Epinephelus septemfasciatus: Mori, 1928; Mori and Uchida, 1934; Katayama, 1940; Honma,
1952; Mori, 1952; Honma, 1955b; Kato, 1956; Mori, 1956a; Katayama, 1960; Honma, 1963;
Lindberg and Krasyukova, 1969; Takegawa and Morino, 1970; Ueno, 1971; Honma, 1977;
Honma and Kitami, 1978; Tsuda, 1990; Suzuki et al., 2000; Maeda and Tsutsui, 2003; Shinohara
et al., 2011; Honma, 2013; Senou, 2013.
SMBS uncatalogued (1).
Niphon spinosus Cuvier, 1828 [Japanese name: Ara]
Niphon spinosus: Cuvier and Valenciennes, 1828 (original description); Mori and Uchida,1934; Katayama, 1940; Kato, 1956; Mori, 1956a; Katayama, 1960; Honma, 1963; Takegawa andMorino, 1970; Honma and Kitami, 1978; Tsuda, 1990; Uozu Aquarium, 1997; Honma, 2013;
Senou, 2013.
Niphon spinous [laspus calami]: Shinohara et al., 2011.
Plectranthias kelloggi azumanus (Jordan and Richardson, 1910) [Japanese name: Azuma-hanadai] (Fig. 2I)

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Zallanthias azumanus: Tanaka, 1919; Katayama, 1940; Honma, 1952; Mori, 1952; Matsubara, 1955; Kato, 1956; Mori, 1956a; Katayama, 1960; Lindberg and Krasyukova, 1969; Tsuda,
1990.
Plectranthias kelloggi azumanus: Uozu Aquarium, 1997; Senou, 2013; Honma, 2013.
SMBS uncatalogued (2).
Family Callanthidae
Callanthias japonicus Franz, 1910 [Japanese name: Shikishima-hanadai]
Percanthias japonicus: Kato, 1956; Mori, 1956a; Takegawa and Morino, 1970; Honma andKitami, 1978.
Callanthias japonicus: Katayama, 1960; Lindberg and Krasyukova, 1969; Honma andKitami, 1970; Honma et al., 1972; Tsuda, 1990; Senou, 1993, 1997; Uozu Aquarium, 1997;
Senou, 2000; Suzuki et al., 2000; Kobayashi et al., 2006; Shinohara et al., 2011; Honma, 2013;
Senou, 2013.
SMBS uncatalogued (1).
Family Banjosidae*Banjos banjos (Richardson, 1846) [Japanese name: Chōsen-bakama]Banjos banjos: Mori, 1952; Kato, 1956; Mori, 1956a; Lindberg and Krasyukova, 1969;
Honma et al., 2002; Hatooka, 2013; Honma, 2013.
Family Priacanthidae
Cookeolus japonicus (Cuvier, 1829) [Japanese name: Chikame-kintoki] (Fig. 2F)
Priacanthus japonicus: Mori and Uchida, 1934.
Priacanthus boops: Katayama, 1940; Honma, 1952; Mori, 1952; Honma, 1955b; Kato, 1956;
Mori, 1956a; Honma, 1963; Lindberg and Krasyukova, 1969; Takegawa and Morino, 1970;
Honma, 1977; Honma and Kitami, 1978; Tsuda, 1990.
Cookeolus boops: Uozu Aquarium, 1997.
Cookeolus japonicus: Suzuki et al., 2000; Maeda and Tsutsui, 2003; Kitagawa et al., 2008;
Shinohara et al., 2011; Hayashi, 2013; Honma, 2013.
SMBS uncatalogued (2).
Priacanthus macracanthus Cuvier, 1829 [Japanese name: Kintokidai]
Priacanthus macracanthus: Mori and Uchida, 1934; Honma, 1952; Mori, 1952; Kato, 1956;
Lindberg and Krasyukova, 1969; Honma and Kitami, 1970, 1978; Tsuda, 1990; Uozu Aquarium,
1997; Adrianov and Kussakin, 1998; Sokolovskaya et al., 1998; Suzuki et al., 2000; Hayashi,
2002; Kitagawa et al., 2008; Shinohara et al., 2011; Hayashi, 2013; Honma, 2013.
Pristigenys niphonia (Cuvier, 1829) [Japanese name: Kurumadai]
Pseudopriacanthus niphonius: Katayama, 1940; Honma, 1952; Mori, 1952; Kato, 1956;
Mori, 1956a; Lindberg and Krasyukova, 1969.
Pristigenys niphonia: Tsuda, 1990; Uozu Aquarium, 1997; Hayashi, 2013; Honma, 2013.
Family Malacanthidae*Branchiostegus auratus (Kishinouye, 1907) [Japanese name: Ki-amadai]
Branchiostegus auratus: Mori, 1952, 1956a; Tsuda, 1990; Aizawa and Doiuchi, 2013.

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Branchiostegus japonicus auratus: Kato, 1956; Lindberg and Krasyukova, 1969; Takegawa
and Morino, 1970.
Branchiostegus japonicus (Houttuyn, 1782) [Japanese name: Aka-amadai]
Branchiostegus japonicus: Mori and Uchida, 1934; Katayama, 1940; Honma, 1952; Mori,
1952; Kato, 1956; Mori, 1956a; Honma, 1977; Uozu Aquarium, 1997; Suzuki et al., 2000;
Kobayashi et al., 2006; Kitagawa et al., 2008; Shinohara et al., 2011; Aizawa and Doiuchi, 2013.
Branchiostegus japonicus japonicus: Lindberg and Krasyukova, 1969; Takegawa andMorino, 1970.
Branchiostegus japonicas [sic]: Honma, 2013.
Family Scombropidae
Scombrops boops (Houttuyn, 1782) [Japanese name: Mutsu]
Scombrops boops: Katayama, 1940; Honma, 1952; Kato, 1956; Mori, 1956a; Lindberg andKrasyukova, 1969; Honma and Kitami, 1970; Takegawa and Morino, 1970; Ueno, 1971; Honmaand Kitami, 1978; Tsuda, 1990; Suzuki et al., 2000; Maeda and Tsutsui, 2003; Shinohara et al.,2011; Hayashi, 2013; Honma, 2013.
SMBS uncatalogued (1).
Family Bramidae
Brama dussumieri Cuvier, 1831 [Japanese name: Hime-shimagatsuo]
Brama dussumieri: Suzuki et al., 2000; Shinohara et al., 2011; Honma et al., 2012; Hatooka
and Kai, 2013; Honma, 2013.
SMBS uncatalogued (1).
Brama japonica Hilgendorf, 1878 [Japanese name: Shimagatsuo]
Brama raii: Katayama, 1940; Mori, 1952; Honma and Mizusawa, 1966; Lindberg and Krasyukova, 1969; Tsuda, 1990.
Lepidotus brama: Kato, 1956; Mori, 1956a; Takegawa and Morino, 1970.
Brama japonica: Uozu Aquarium, 1997; Suzuki et al., 2000; Maeda and Tsutsui, 2003;
Kobayashi et al., 2006; Shinohara et al., 2011; Hatooka and Kai, 2013; Honma, 2013.
Pteraclis aesticola (Jordan and Snyder, 1901) [Japanese name: Benten-uo]
Bentenia aesticola: Mori, 1952; Matsubara, 1955; Kato, 1956; Mori, 1956a.
Pteraclis velifera: Lindberg and Krasyukova, 1969.
Pteraclis aesticola: Hatooka, 1993, 2000, 2002; Shinohara et al., 2011; Hatooka and Kai,
2013.
Pterycombus petersii (Hilgendorf, 1878) [Japanese name: Ryūgūnohime] (Fig. 2G)Centropholis petersi: Matsubara, 1955; Kato, 1956; Mori, 1956a; Honma and Mizusawa,
1966; Lindberg and Krasyukova, 1969; Takegawa and Morino, 1970; Ueno, 1971; Tsuda, 1990.
Pterycombus petersii: Honma, 1991a, b; Uozu Aquarium, 1997; Suzuki et al., 2000; Maeda
and Tsutsui, 2003; Kobayashi et al., 2006; Hatooka and Kai, 2013; Honma, 2013.
Pterycombus petersi [lapsus calami]: Shinohara et al., 2011.
SMBS uncatalogued (3).
*Taractes asper Lowe, 1843 [Japanese name: Manzai-uo]
Taractes platycephalus: Mori, 1952; Honma, 1962, 1963; Lindberg and Krasyukova, 1969;
Honma and Kitami, 1978; Tsuda, 1990.
Taractes asper: Hatooka, 1993, 2000, 2002; Maeda and Tsutsui, 2003; Hatooka and Kai,

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2013; Honma, 2013.
Taractes rubescens (Jordan and Evermann, 1887) [Japanese name: Tsurugi-echiopia]
Steunegeria rubescens: Honma, 1963; Lindberg and Krasyukova, 1969; Takegawa andMorino, 1970; Honma and Kitami, 1978.
Taractes rubescens: Honma and Mizusawa, 1966; Tsuda, 1990; Uozu Aquarium, 1997;
Hatooka and Kai, 2013; Honma, 2013.
*Taractichthys steindachneri (Döderlein, 1883) [Japanese name: Hirejiro-manzai-uo]
Taractichthys steindachneri: Honma, 1963; Honma and Mizusawa, 1966; Tsuda, 1990;
Hatooka, 1993, 2000; Kobayashi et al., 2006; Hatooka and Kai, 2013; Honma, 2013.
Taractichthys longipinnis: Honma and Mizusawa, 1966; Lindberg and Krasyukova, 1969;
Honma and Kitami, 1978; Tsuda, 1990.
Family Caristiidae
Caristius macropus (Bellotti, 1903) [Japanese name: Yaegisu]
Elephenor macropus: Ueno, 1971.
Platyberyx macropus: Hatooka, 2000, 2002; Shinohara et al., 2011.
Caristius macropus: Hatooka, 2013.
Family Emmelichthyidae
Emmelichthys strusakeri Heemstra and Randall, 1977 [Japanese name: Rōsoku-chibiki]
Emmelichthys strusakeri: Hatooka, 1993, 2000; Honma et al., 2000; Suzuki et al., 2000;
Hatooka, 2002; Kitagawa et al., 2008; Shinohara et al., 2011; Hatooka and Hagiwara, 2013;
Honma, 2013.
Erythrocles microceps Miyahara and Okamura, 1998 [Japanese name: Hichibiki]
Erythrocles microceps: Hatooka, 2000; Suzuki et al., 2000; Hatooka, 2002; Shinohara et al.,2011; Hatooka and Hagiwara, 2013.
SMBS uncatalogued (2).
Erythrocles schlegelii (Richardson, 1846) [Japanese name: Hachibiki]
Erythrocles schlegeli: Mori and Uchida, 1934; Honma, 1952; Mori, 1952; Kato, 1956; Mori,
1956a; Honma, 1963; Honma and Kitami, 1967; Lindberg and Krasyukova, 1969; Takegawa andMorino, 1970; Honma, 1977; Honma and Kitami, 1978.
Erythrocles schlegelii: Uozu Aquarium, 1997; Hatooka, 2002; Suzuki et al., 2000;
Kobayashi et al., 2006; Shinohara et al., 2011; Hatooka and Hagiwara, 2013; Honma, 2013.
SMBS uncatalogued (1).
Family Lutjanidae*Etelis coruscans Valenciennes, 1862 [Japanese name: Hamadai]
Etelis carbunculus: Mori, 1952.
Family Sparidae
Dentex hypselosomus Bleeker, 1854 [Japanese name: Kidai]
Taius tumifrons: Tanaka, 1915; Mori, 1928; Mori and Uchida, 1934; Katayama, 1940;
Honma, 1952: Mori, 1952; Kato, 1956.
Dentex tumifrons: Mori, 1956a; Lindberg and Krasyukova, 1969; Tsuda, 1990; Uozu Aquar-

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ium, 1997; Suzuki et al., 2000; Shinohara et al., 2011; Honma, 2013.
Dentex hypselosomus: Hayashi and Hagiwara, 2013.
*Evynnis cardinalis (Lacepède, 1802) [Japanese name: Hirekodai]
Evynnis cardinalis: Mori, 1928; Mori and Uchida, 1934; Mori, 1952; Kato, 1956; Mori,
1956a; Lindberg and Krasyukova, 1969; Tsuda, 1990; Hayashi and Hagiwara, 2013.
Family Glaucosomatidae*Glaucosoma hebraicum Richardson, 1845 [Japanese name: Aobadai]
Glaucosoma hebraicum: Kato, 1956; Mori, 1956a; Tsuda, 1990; Hayashi and Hagiwara,
2013.
Family Pentacerotidae
Evistias acutirostris (Temminck and Schlegel, 1844) [Japanese name: Tengudai]
Evistias acutirostris: Mori, 1952; Lindberg and Krasyukova, 1969; Tsuda, 1990; Honma,
1991a; Uozu Aquarium, 1997; Suzuki et al., 2000; Maeda and Tsutsui, 2003; Kobayashi et al.,2006; Shinohara et al., 2011; Hatooka and Yagishita, 2013.
Histiopterus typus Temminck and Schlegel, 1844 [Japanese name: Kawabisha] (Fig. 2H)
Histiopterus typus: Katayama, 1940; Kato, 1956; Mori, 1956a; Lindberg and Krasyukova,
1969; Takegawa and Morino, 1970; Tsuda, 1990; Suzuki et al., 2000; Shinohara et al., 2011;
Hatooka and Yagishita, 2013; Honma, 2013.
SMBS uncatalogued (1).
*Pentaceros japonicus Steindachner, 1883 [Japanese name: Tsubodai]
Quinquarius japonicus: Kato, 1956.
Pentaceros japonicus: Tsuda, 1990; Kitagawa et al., 2008; Hatooka and Yagishita, 2013;
Honma, 2013.
Family Zoarcidae
Bilabria ornata (Soldatov, 1922) [Japanese name: Usa-genge]
Lycenchelys ornatus: Soldatov, 1922 (original description).
Bilabria ornata: Taranetz, 1937; Schmidt, 1950; Fowler, 1958; Ueno, 1971; Lindberg andKrasyukova, 1975; Hatooka, 2002.
Bothrocara hollandi (Jordan and Hubbs, 1925) [Japanese name: Noro-genge]
Allolepis hollandi: Jordan and Hubbs, 1925 (original description); Okada and Matsubara,
1938; Katayama, 1940, 1949; Honma, 1952; Katayama, 1952; Mori, 1952; Matsubara, 1955;
Kato, 1956; Mori, 1956a; Fowler, 1958; Honma, 1963; Ouchi, 1963; Takegawa and Morino,
1970; Ueno, 1971; Ogata et al., 1973; Lindberg and Krasyukova, 1975; Honma, 1977; Honmaand Kitami, 1978; Tsuda, 1990; Nambu et al., 1992; Hatooka, 1993; Nambu et al., 1993; Amaoka et al., 1995; Adrianov and Kussakin, 1998; Sokolovskaya et al., 1998; Suzuki et al., 2000;
Balanov and Solomatov, 2008; Balushkin et al., 2011.
Lycogramma crystallonota: Popov, 1933 (original description).
Allolepis nazumii: Mori, 1956c (original description).
Allolepis nazumi: Fowler, 1958; Lindberg and Krasyukova, 1975.
Bothrocara hollandi: Hatooka, 2000, 2002; Maeda and Tsutsui, 2003; Anderson andFedorov, 2004; Balanov and Solomatov, 2008; Anderson et al., 2009; Shinohara et al., 2011;

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Hatooka, 2013; Honma, 2013.
Bothrocara holladi [sic]: Kitagawa et al., 2008.
SO10-C4-B4 (1), KT-10-08-R4 (1), KT-10-08-R6 (1), KT-11-09-M5 (3), TS09-T001 (1),TS09-T004 (10), TS09-T006 (1), TS09-T010 (3), TS09-T049 (1), TS09-T053 (1), TS09-T102(2), TS09-T106 (11), TS10-OW09 (11), TS10-OW10 (2), TS10-OW12 (2), TS10-OW13 (1),TS10-OW18 (4), TS10-YA05 (7).
Bothrocara tanakae (Jordan and Hubbs, 1925) [Japanese name: Kanten-genge]
Zestichthys tanakae: Hikita, 1950; Oshima, 1957; Lindberg and Krasyukova, 1975; Tsuda,
1990.
Zestichthys tanakai: Kato, 1956; Ueno, 1971; Hatooka, 1993.
Bothrocara tanakae: Hatooka, 2000, 2002; Shinohara et al., 2011; Hatooka, 2013.
*Bothrocara zestum Jordan and Fowler, 1902 [Japanese name: Shiro-genge]
Lycogramma zestum: Katayama, 1940; Lindberg and Krasyukova, 1975.
Lycgramma zesta [sic]: Ueno, 1971.
Lycograma zosta [sic]: Tsuda, 1990.
Davidijordania lacertina (Pavlenko, 1910) [Japanese name: Naga-sarasagaji]
Davidijordania lacertina: Matsubara, 1955; Fowler, 1958; Lindberg and Krasyukova, 1975;
Hatooka, 1993, Adrianov and Kussakin, 1998; Sokolovskaya et al., 1998; Hatooka, 2000, 2002;
Anderson and Fedorov, 2004; Balanov et al., 2006; Balanov and Solomatov, 2008; Shinohara
and Kim, 2009; Balushkin et al., 2011.
Davidijordania poecilimon (Jordan and Fowler, 1902) [Japanese name: Sarasagaji]
Davidijordania poecilimon: Taranetz, 1937; Okada and Matsubara, 1938; Katayama, 1940,
1949; Honma, 1952; Matsubara, 1955; Kato, 1956; Mori, 1956a; Fowler, 1958; Honma, 1963;
Takegawa and Morino, 1970; Lindberg and Krasyukova, 1975; Honma, 1977; Honma andKitami, 1978; Tsuda, 1990; Hatooka, 1993; Nambu et al., 1993; Amaoka et al., 1995; Uozu
Aquarium, 1997; Adrianov and Kussakin, 1998; Sokolovskaya et al., 1998; Hatooka, 2000,
2002; Maeda and Tsutsui, 2003; Anderson and Fedorov, 2004; Kitagawa et al., 2008; Balushkin
et al., 2011.
Davidijordania spilota: Fowler, 1943 (original description), 1958.
Davidijordani poecilimon [lapsus calami]: Shinohara et al., 2011.
KT-11-09-N1 (1), TS09-T107 (2), SMBS uncatalogued (2).
Derjuginia japonica (Katayama, 1943) [Japanese name: Nirami-genge]
Gengea japonica: Katayama, 1943 (original description), 1949, 1952; Honma, 1955a, e;Matsubara, 1955; Kato, 1956; Mori, 1956a; Fowler, 1958; Honma, 1963; Ouchi, 1963; Takegawa
and Morino, 1970; Lindberg and Krasyukova, 1975; Tsuda, 1990; Uozu Aquarium, 1997.
Derjuginia japonica: Toyoshima, 1981; Hatooka, 1993, 2000, 2002; Maeda and Tsutsui,
2003; Balushkin et al., 2011; Shinohara et al., 2011.
Gymnelopsis ochotensis: Anderson and Fedorov, 2004; Hatooka, 2013; Honma, 2013.
TS09-T099 (2), TS09-T106 (3), TS09-T141 (2).
Krusensterniella maculata Andriashev, 1938 [Japanese name: Ai-haregaji] (Fig. 4B)
Krusensterniella maculata: Andriashev, 1938 (original description); Matsubara, 1955;
Fowler, 1958; Ueno, 1971; Lindberg and Krasyukova, 1975; Hatooka, 1993, Adrianov and Kussakin, 1998; Sokolovskaya et al., 1998; Hatooka, 2000, 2002; Maeda and Tsutsui, 2003; Anderson and Fedorov, 2004; Balushkin et al., 2011; Hatooka, 2013.
KT-11-09-M1 (1).
*Krusensterniella notabilis Schmidt, 1904 [Japanese name: Haregaji]

Annotated Checklist of Deep-sea Fishes of the Sea of Japan265

Krusensterniella notabilis: Anderson and Fedorov, 2004.
*Lycodes brevipes Bean, 1890 [Japanese name: Ashibosogenge]
Lycodes brevipes: Ueno, 1971.
Lycodes brevipes ochotensis: Lindberg and Krasyukova, 1975.
*Lycodes brunneofasciatus Suvorov, 1935 [Japanese name: Harasuji-genge]
Lycodes brunneofasciatus: Hatooka, 1993, 2000, 2002, 2013.
*Lycodes caudimaculatus Matsubara, 1936 [Japanese name: Irezumigaji]
Lycodes caudimaculatus: Tekegawa and Morino, 1970; Lindberg and Krasyukova, 1975;
Tsuda, 1990.
Lycodes japonicus Matsubara and Iwai, 1951 [Japanese name: Ashinaga-genge]
Lycodes japonicus: Matsubara and Iwai, 1951b (original description); Matsubara, 1955;
Honma, 1956a; Kato, 1956; Honma, 1957a; Oshima, 1957; Honma, 1963; Ogata et al., 1973;
Lindberg and Krasyukova, 1975; Honma and Kitami, 1978; Toyoshima, 1984; Toyoshima, 1985;
Nambu et al., 1992; Tsuda, 1990; Hatooka, 1993; Nambu et al., 1993; Uozu Aquarium, 1997;
Hatooka, 2000, 2002; Anderson and Fedorov, 2004; Balanov and Solomatov, 2008; Shinohara etal., 2011; Hatooka, 2013; Honma, 2013.
TS09-T033 (3), TS09-T53 (1), TS09-T053 (1), TS09-T057 (2), TS09-T110 (5), TS10-OW04(4).
*Lycodes macrolepis Taranetz and Andriashev, 1935 [Japanese name: Yokoshima-gaji]
Lycodes macrolepis: Katayama, 1949; Matsubara and Iwai, 1951b; Katayama,1952; Matsubara, 1955; Mori, 1956a, c; Honma, 1957a; Oshima, 1957; Fowler, 1958; Honma, 1963; Ouchi,
1963; Lindberg and Krasyukova, 1975; Honma and Kitami, 1978; Tsuda, 1990; Hatooka, 1993,
2000, 2002.
Lycodes matsubarai Toyoshima, 1985 [Japanese name: Matsubara-genge]
Lycodes yamatoi: Toyoshima, 1985 (original description); Hatooka, 1993, 2000, 2002;
Maeda and Tsutsui, 2003; Anderson and Fedorov, 2004; Balanov and Kukhlevsky, 2011; Balushkin et al., 2011; Shinohara et al., 2011; Honma, 2013.
Lycodes matsubarai: Nambu et al., 1992; Hatooka, 1993, 2000, 2002; Nazarkin and Shinohara, 2012; Hatooka, 2013; Honma, 2013.
KT-11-09- K3 (1), TS09-T004 (3), TS09-T006 (2), TS09-T013 (2), TS09-T141 (1).
Lycodes nakamurae (Tanaka, 1914) [Japanese name: Kuro-genge]
Furcimanus nakamurae: Tanaka, 1914b (original description); Jordan and Hubbs, 1925;
Okada and Matsubara, 1938; Katayama, 1940, 1949; Honma, 1952; Mori, 1952; Katayama,
1952; Mori, 1956a; Oshima, 1957.
Lycodes diapterus nakamurae: Lindberg, 1947; Tsuda, 1990.
Lycodes nakamurai: Matsubara, 1955; Kato, 1956; Honma and Sugihara, 1963; Ouchi, 1963;
Takegawa and Morino, 1970; Ogata et al., 1973; Honma, 1977; Toyoshima, 1985, 1988; Nambu
et al., 1992; Hatooka, 1993; Nambu et al., 1993; Amaoka et al., 1995; Sokolovskaya et al., 1998;
Hatooka, 2000; Suzuki et al., 2000; Hatooka, 2002; Maeda and Tsutsui, 2003; Balanov and Solomatov, 2008; Shinohara et al., 2011; Honma, 2013.
Furcimanus nakamurai: Mori, 1956c.
Lycodes diapterus nakamurai: Honma, 1963; Lindberg and Krasyukova, 1975; Honma andKitami, 1978.
Lycodes nakamurae: Ueno, 1971; Anderson and Fedorov, 2004; Balanov and Solomatov,
2008; Hatooka, 2013.
KT-11-09-N3 (2), TS09-T027 (6), TS09-T029 (2), TS09-T054 (2), TS09-T056 (2), TS09-

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T086 (1), TS09-T099 (1), TS09-T101 (2), TS09-T102 (3), SMBS uncatalogued (3).
Lycodes nishimurai Shinohara and Shirai, 2005 [Japanese name: Orochi-genge] (Fig. 4C)
Lycodes nishimurai: Shinohara and Shirai, 2005 (original description); Balushkin et al.,2011; Shinohara et al., 2011; Hatooka, 2013.
TS09-T007 (1), TS09-T033 (6), TS09-T053 (1), TS09-T057 (2).
Lycodes palearis fasciatus (Schmidt, 1904)
Lycodes palearis fasciatus: Taranetz, 1937; Lindberg and Krasyukova, 1975; Sokolovskaya
et al., 1998; Shinohara et al., 2011.
Lycodes fasciatus: Fowler, 1958.
Lycodes paucilepidotus Toyoshima, 1985 [Japanese name: Hage-mayugaji]
Lycodes paucilepidotus: Toyoshima, 1985 (original description); Hatooka, 1993; Amaoka etal., 1995; Hatooka, 2000, 2002; Maeda and Tsutsui, 2003; Hatooka, 2013.
*Lycodes schmidti Gratzianov, 1907
Lycodes palearis schmidti: Lindberg and Krasyukova, 1975; Adrianov and Kussakin, 1998;
Sokolovskaya et al., 1998.
Lycodes schmidti: Anderson and Fedorov, 2004.
Lycodes sigmatoides Lindberg and Krasyukova, 1975 [Japanese name: Esuji-gaji]
Lycodes schmidti: Kato, 1956; Ueno, 1971.
Lycodes sigmatoides: Lindberg and Krasyukova, 1975 (original description); Tsuda, 1990;
Anderson and Fedorov, 2004; Shinohara et al., 2011; Hatooka, 2013.
Lycodes sygmatoides [sic]: Sokolovskaya et al., 1998.
*Lycodes soldatovi Taranetz and Andriashev, 1935 [Japanese name: Kuro-gaji]
Lycodes soldatovi: Kato, 1956; Ouchi, 1963; Lindberg and Krasyukova, 1975.
Remarks. Hatooka (2013) mentioned the occurence from Sado Island with a question mark.
Lycodes tanakae Jordan and Thompson, 1914 [Japanese name: Tanaka-genge]
Lycodes tanakae: Jordan and Thompson, 1914 (original description); Okada and Matsubara,
1938; Katayama, 1940, 1949; Honma, 1952; Katayama, 1952; Mori, 1952; Fowler, 1958; Lindberg and Krasyukova, 1975; Tsuda, 1990; Hatooka, 2000, 2002; Maeda and Tsutsui, 2003;
Anderson and Fedorov, 2004; Balanov and Solomatov, 2008; Balushkin et al., 2011; Shinohara
et al., 2011; Hatooka, 2013; Honma, 2013.
Lycodes tanakai: Katayama, 1949; Matsubara and Iwai, 1951b; Matsubara, 1955; Kato,
1956; Mori, 1956a, c; Honma, 1963; Ouchi, 1963; Honma and Sugihara, 1963; Takegawa andMorino, 1970; Ueno, 1971; Ogata et al., 1973; Honma, 1977; Honma and Kitami, 1978; Toyoshima, 1985; Hatooka, 1993; Nambu et al., 1993; Amaoka et al., 1995; Koyanagi, 1997; Uozu
Aquarium, 1997; Sokolovskaya et al., 1998; Suzuki et al., 2000.
TS09-T002 (2), TS09-T004 (1), TS09-T006 (1), TS09-T013 (3), TS09-T049 (1), TS09-T102(6), TS09-T106 (2), TS09-T141 (1).
*Lycodes uschakovi Popov, 1931 [Japanese name: Kawari-genge]
Lycodes uschakovi: Lindberg and Krasyukova, 1975; Toyoshima, 1985, 1988; Hatooka,
1993; Sokolovskaya et al., 1998; Hatooka, 2000, 2002; Maeda and Tsutsui, 2003; Anderson andFedorov, 2004; Hatooka, 2013.
*Lycodes ygreknotatus Schmidt, 1950 [Japanese name: Waiji-genge]
Lycodes ygreknotatus: Ueno, 1971; Hatooka, 2013.
Lycozoarces regani Popov, 1933 [Japanese name: Sedaka-genge]
Lycozoarces hubbsi: Matsubara, 1955; Fowler, 1958; Ueno, 1971.
Lycozoarces regani: Popov, 1933 (original description); Lindberg and Krasyukova, 1975;

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267

Fig. 3. Zoarcid specimens of Petroschmidtia teraoi (Katayama, 1943) reported as “Sadohinagenge, Lycodes
sadoensis Matsubara and Honma (MS)” in Honma (1963), SMBS uncatalogued, 126, 127, 140 and 103 mm
SL (from top to bottom).

Hatooka, 1993; Amaoka et al., 1995; Hatooka, 2000, 2002; Maeda and Tsutsui, 2003; Andersonand Fedorov, 2004; Balushkin et al., 2011; Hatooka, 2013.
*Petroschmidtia albonotata Taranetz and Andriashev, 1934 [Japanese name: Hanagenge]
Petroschmidtia albonotata: Ueno, 1971.
Petroschmidtia teraoi (Katayama, 1943) [Japanese name: Hina-genge]
Lycodes teraoi: Katayama, 1943 (original description), 1949; Matsubara and Iwai, 1951b;
Katayama, 1952; Matsubara, 1955; Kato, 1956; Mori, 1956a; Fowler, 1958; Takegawa andMorino, 1970; Lindberg and Krasyukova, 1975; Toyoshima, 1985; Tsuda, 1990; Hatooka, 1993,
2000, 2002; Anderson and Fedorov, 2004; Shinohara et al., 2011; Hatooka, 2013.
Lycodes sp.: Kato, 1956; Honma, 1963; Ouchi, 1963; Honma, 1977.
Lycodes sadonesis [nomen nudum]: Honma and Kitami, 1978.
Lycodes sadoensis: Toyoshima and Honma, 1980 (original description); Honma et al., 1984;
Toyoshima, 1985; Hatooka, 1993, 2000, 2002; Anderson and Fedorov, 2004; Shinohara et al.,2011; Hatooka, 2013; Honma, 2013.
TS09-T001 (1), TS09-T012 (2), TS09-T013 (3), TS09-T017 (1), TS09-T018 (6), TS09-T020(20), TS09-T034 (2), TS09-T038 (1), TS09-T039 (22),TS09-T054 (1), TS09-T087 (3), TS09T099 (1), TS10-OW01 (2), SMBS uncatalogued (4).

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Remarks. Generic position and synonymies of the species follow Nazarkin et al. (2014). The
type specimens of Lycodes sadoensis were collected not from off Sado Island but off Noto Peninsula (Toyoshima and Honma, 1980). Specimens collected off Sado Island and named as Sadohinagenge in Japanese (Honma, 1963) are deposited in SMBS (Fig. 3).
Petroschmidtia toyamensis Katayama, 1941 [Japanese name: Ago-genge]
Petroschmidtia toyamensis: Katayama, 1941 (original description); Katayama, 1943, 1949;
Matsubara and Iwai, 1951b; Katayama, 1952; Honma, 1955c; Matsubara, 1955; Kato, 1956;
Mori, 1956a; Oshima, 1957; Fowler, 1958; Honma, 1963; Honma and Sugihara, 1963; Takegawa
and Morino, 1970; Ogata et al., 1973; Lindberg and Krasyukova, 1975; Honma and Kitami,
1978; Toyoshima, 1985; Tsuda, 1990; Hatooka, 1993; Nambu et al., 1993; Amaoka et al., 1995;
Koyanagi, 1997; Uozu Aquarium, 1997; Hatooka, 2000; Suzuki et al., 2000; Hatooka, 2002;
Maeda and Tsutsui, 2003; Anderson and Fedorov, 2004; Shinohara et al., 2011; Honma, 2013.
Petreoschmidtia toyamaensis [sic]: Ueno, 1971.
Lycodes toyamensis: Balanov and Solomatov, 2008; Hatooka, 2013.
KT-11-09-K4 (2), TS09-T033 (1), TS09-T106 (1).
Family Stichaeidae*Acantholumpenus mackayi (Gilbert, 1896) [Japanese name: Nuime-gaji]
Lumpenus fowleri: Kato, 1956; Fowler, 1958.
Acantholumpenus mackayi: Ueno, 1971; Tsuda, 1990; Adrianov and Kussakin, 1998; Sokolovskaya et al., 1998; Maeda and Tsutsui, 2003; Mecklenburg and Sheiko, 2004.
Anisarchus macrops (Matsubara and Ochiai, 1952) [Japanese name: Medama-ginpo]
Lumpenus macrops: Matsubara and Ochiai, 1952 (original description); Matsubara, 1955;
Honma, 1955a, e; Kato, 1956; Tomiyama and Abe, 1958; Honma, 1963, 1977; Honma andKitami, 1978.
Anisarchus macrops: Lindberg and Krasyukova, 1975; Tsuda, 1990; Hatooka, 1993; Amaoka
et al., 1995; Miki, 1997; Adrianov and Kussakin, 1998; Sokolovskaya et al., 1998; Hatooka,
2000, 2002; Mecklenburg and Sheiko, 2004; Shinohara et al., 2011; Hatooka, 2013; Honma,
2013.
KT-10-08-R1 (7), TS09-T117 (4).
Anisarchus medius (Reinhardt, 1837) [Japanese name: Nise-medama-ginpo]
Anisarchus medius: Matsubara, 1955; Lindberg and Krasyukova, 1975; Adrianov and Kussakin, 1998; Sokolovskaya et al., 1998; Mecklenburg and Sheiko, 2004; Yamanaka and Yabe,
2012.
Bryozonichthys lysimus (Jordan and Snyder, 1902) [Japanese name: Fusakake-ginpo]
Bryozonichthys lysimus: Nambu et al., 1992; Hatooka, 1993; Amaoka et al., 1995; Hatooka,
2000, 2002; Maeda and Tsutsui, 2003; Mecklenburg and Sheiko, 2004; Hatooka, 2013.
*Leptoclinus maculatus (Fries, 1838) [Japanese name: Yase-ginpo]
Leptoclinus maculatus: Ueno, 1971; Mecklenburg and Sheiko, 2004; Hatooka, 2013.
Lumpenella longirostris (Evermann and Goldsborough, 1907) [Japanese name: Nezumi-ginpo]
Lumpenella longirostris: Soldatov and Lindberg, 1930; Taranetz, 1937; Schmidt, 1950;
Ueno, 1971; Lindberg and Krasyukova, 1975; Nambu et al., 1992; Hatooka, 1993; Amaoka etal., 1995; Uozu Aquarium, 1997; Sokolovskaya et al., 1998; Hatooka, 2000, 2002; Maeda andTsutsui, 2003; Mecklenburg and Sheiko, 2004; Kitagawa et al., 2008; Shinohara et al., 2011;
Hatooka, 2013; Honma, 2013.

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269

Lumpenella nigricans: Honma, 1955a, e; Matsubara, 1955; Kato, 1956; Honma, 1963;
Honma and Kitami, 1978; Tsuda, 1990.
Lumpenopsis triocellata (Matsubara, 1943) [Japanese name: Mitsuboshi-ginpo]
Leptoclinus triocellatus: Matsubara, 1943a (original description).
Lumpenopsis triocellatus: Honma, 1963; Lindberg and Krasyukova, 1975; Honma andKitami, 1978; Tsuda, 1990; Hatooka, 1993; Miki, 1997; Hatooka, 2000, 2002; Mecklenburg andSheiko, 2004; Hatooka, 2013; Honma, 2013.
Lumpenus sagitta Wilimovsky, 1956 [Japanese name: Unagi-gaji]
Lumpenus anguillaris: Tanaka, 1913b; Okada and Matsubara, 1938; Katayama, 1940;
Honma, 1952; Fowler, 1958.
Lumpenus gracilis: Matsubara, 1955; Kato, 1956; Honma, 1963.
Lumpenus sagitta: Ueno, 1971; Lindberg and Krasyukova, 1975; Honma and Kitami, 1978;
Tsuda, 1990; Hatooka, 1993; Amaoka et al., 1995; Uozu Aquarium, 1997; Adrianov and Kussakin, 1998; Sokolovskaya et al., 1998; Hatooka, 2000, 2002; Mecklenburg and Sheiko, 2004;
Shinohara et al., 2011; Hatooka, 2013; Honma, 2013.
*Stichaeopsis epallax (Jordan and Snyder, 1902) [Japanese name: Ame-gaji]
Stichaeopsis epallax: Kato, 1956; Fowler, 1958; Ueno, 1971; Tsuda, 1990; Hatooka, 1993;
Amaoka et al., 1995; Miki, 1997; Hatooka, 2000, 2002; Maeda and Tsutsui, 2003; Mecklenburg
and Sheiko, 2004; Hatooka, 2013; Honma, 2013.
Stichaeus grigorjewi Herzenstein, 1890 [Japanese name: Nagaduka]
Stichaeus elongatus: Sakamoto, 1930 (original description); Okada and Matsubara, 1938;
Oshima, 1957.
Dinogunnellus grigorjewi: Mori and Uchida, 1934; Okada and Matsubara, 1938; Katayama,
1940; Hikita, 1950; Honma, 1952; Katayama, 1952; Mori, 1952.
Stichaeus grigorjewi: Honma, 1955b; Matsubara, 1955; Kato, 1956; Mori, 1956a; Fowler,
1958; Honma, 1963; Ouchi, 1963; Takegawa and Morino, 1970; Ueno, 1971; Lindberg and Krasyukova, 1975; Honma, 1977; Honma and Kitami, 1978; Tsuda, 1990; Hatooka, 1993; Amaoka
et al., 1995; Miki, 1997; Uozu Aquarium, 1997; Adrianov and Kussakin, 1998; Sokolovskaya etal., 1998; Hatooka, 2000, 2002; Suzuki et al., 2000; Maeda and Tsutsui, 2003; Mecklenburg andSheiko, 2004; Kitagawa et al., 2008; Shinohara et al., 2011; Hatooka, 2013; Honma, 2013.
Stichaeus nozawae Jordan and Snyder, 1902 [Japanese name: Tauegaji]
Stichaeus nozawæ: Jordan and Snyder, 1902 (original description).
Stichaeus nozawae: Okada and Matsubara, 1938; Oshima, 1957; Fowler, 1958; Honma,
1963; Lindberg and Krasyukova, 1975; Tsuda, 1990; Adrianov and Kussakin, 1998; Sokolovskaya et al., 1998; Mecklenburg and Sheiko, 2004; Hatooka, 2013.
Stichaeus nozawai: Matsubara, 1955; Kato, 1956; Ouchi, 1963; Ueno, 1971; Honma andKitami, 1978; Hatooka, 1993; Amaoka et al., 1995; Miki, 1997; Hatooka, 2000, 2002; Maeda
and Tsutsui, 2003; Honma, 2013.
Xenolumpenus longipterus Shinohara and Yabe, 2009 [Japanese name: Furisode-gaji] (Fig. 4D)
Xenolumpenus longipterus: Shinohara and Yabe, 2009 (original description); Hatooka, 2013.
Family Cryptacanthodidae
Cryptacanthodes bergi Lindberg, 1930 [Japanese name: Hadaka-ōkami-uo]
Cryptacanthodes bergi: Soldatov and Lindberg, 1930; Matsubara, 1955; Kato, 1956; Honma,
1957a; Fowler, 1958; Honma, 1963; Takegawa and Morino, 1970; Ueno, 1971; Lindberg and270

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Fig. 4. Freshly dead specimens of four species endemic to the Sea of Japan. A, Careproctus notosaikaiensis,
FAKU 135383, 160 mm SL, off Noto Peninsula; B, Krusensterniella maculata, NSMT-P 106556, 80 mm SL,
Musashi Bank; C, Lycodes nishimurai, NSMT-P 67246, 275 mm SL, Yamato Bank; D, Xenolumpenus longipterus, NSMT-P 69644, holotype, 115 mm SL, near Okushiri Island.

Krasyukova, 1975; Honma and Kitami, 1978; Tsuda, 1990; Hatooka, 1993; Amaoka et al., 1995;
Adrianov and Kussakin, 1998; Sokolovskaya et al., 1998; Hatooka, 2000, 2002; Maeda and Tsutsui, 2003; Hatooka, 2013; Honma, 2013.
Cryptacanthocephoides bergi [lapsus calami]: Shinohara et al., 2011.
SMBS uncatalogued (2).
Family Champsodontidae
Champsodon snyderi Franz, 1910 [Japanese name: Wanigisu]
Champsodon snyderi: Katayama, 1940; Honma, 1955d; Kato, 1956; Mori, 1956a; Honma,

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1963; Lindberg and Krasyukova, 1969; Takegawa and Morino, 1970; Honma and Kitami, 1978;
Tsuda, 1990; Uozu Aquarium, 1997; Suzuki et al., 2000; Shinohara et al., 2011; Hatooka andHagiwara, 2013; Honma, 2013.
Family Trichodontidae
Arctoscopus japonicus (Steindachner, 1881) [Japanese name: Hatahata]
Arctoscopus japonicus: Mori, 1928; Mori and Uchida, 1934; Okada and Matsubara, 1938;
Katayama, 1940; Honma, 1952; Katayama, 1952; Kato, 1956; Mori, 1956a, c; Honma, 1963;
Ouchi, 1963; Honma, 1968; Lindberg and Krasyukova, 1969; Honma and Kitami, 1970; Takegawa and Morino, 1970; Ueno, 1971; Ogata et al., 1973; Honma, 1977; Honma and Kitami,
1978; Tsuda, 1990; Nambu et al., 1992; Hatooka, 1993; Amaoka et al., 1995; Amaoka, 1997;
Adrianov and Kussakin, 1998; Sokolovskaya et al., 1998; Hatooka, 2000; Suzuki et al., 2000;
Hatooka, 2002; Maeda and Tsutsui, 2003; Kitagawa et al., 2008; Shinohara et al., 2011, Hatooka,
2013; Honma, 2013.
KT-11-09-M1 (2), TS09-T004 (28).
Family Uranoscopidae
Uranoscopus japonicus Houttuyn, 1782 [Japanese name: Mishima-okoze]
Uranoscopus japonicus: Mori, 1928; Mori and Uchida, 1934; Katayama, 1940; Honma,
1952; Mori, 1952; Honma, 1955b; Kato, 1956; Mori, 1956a; Honma, 1963; Lindberg and Krasyukova, 1969; Takegawa and Morino, 1970; Honma and Kitami, 1978; Tsuda, 1990; Suzuki etal., 2000; Shinohara et al., 2011; Honma, 2013; Yamada and Yagishita, 2013.
SMBS uncatalogued (3).
Xenocephalus elongatus (Temminck and Schlegel, 1843) [Japanese name: Ao-mishima]
Gnathagnus elongata: Mori, 1928.
Gnathagnus elongatus: Mori and Uchida, 1934; Katayama, 1940; Honma, 1952; Mori, 1952;
Kato, 1956; Mori, 1956a; Honma, 1963; Lindberg and Krasyukova, 1969; Takegawa andMorino, 1970; Ueno, 1971; Honma and Kitami, 1978; Tsuda, 1990.
Zenocephalus elongatus [sic]: Suzuki et al., 2000.
Xenocephalus elongatus: Maeda and Tsutsui, 2003; Shinohara et al., 2011; Honma, 2013;
Yamada and Yagishita, 2013.
SMBS uncatalogued (1).
Family Callionymidae*Foetorepus altivelis (Temminck and Schlegel, 1845) [Japanese name: Beni-teguri]
Callionymus altivelis: Mori, 1952.
Synchiropus altivelis: Kato, 1956; Mori, 1956a; Lindberg and Krasyukova, 1975; Tsuda,
1990.
Family Gempylidae*Gempylus serpens Cuvier, 1829 [Japanese name: Kurotachi-kamasu]
Gempylus serpens: Tsuda, 1990; Nakabo and Doiuchi, 2013.
Nealotus tripus Johnson, 1865 [Japanese name: Fūrai-kamasu]
Nealotus tripus: Suzuki et al., 2000; Shinohara et al., 2011; Honma et al., 2012; Honma,

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2013; Nakabo and Doiuchi, 2013.
TS09-T017 (1), TS09-T038 (1), TS09-T093 (1).
*Promethichthys prometheus (Cuvier, 1832) [Japanese name: Kuroshibi-kamasu]
Promethichthys prometheus: Kato, 1956; Mori, 1956a; Lindberg and Krasyukova, 1975;
Tsuda, 1990; Nakabo and Doiuchi, 2013.
Rexea prometheoides (Bleeker, 1856) [Japanese name: Kagokamasu]
Rexea prometheoides: Mori, 1952; Honma et al., 1990; Suzuki et al., 2000; Shinohara et al.,2011; Honma, 2013; Nakabo and Doiuchi, 2013.
Ruvettus pretiosus Cocco, 1833 [Japanese name: Baramutsu]
Ruvettus pretiosus: Suzuki et al., 2000; Shinohara et al., 2011; Nakabo and Doiuchi, 2013.
Thyrsitoides marleyi Fowler, 1929 [Japanese name: Nagatachi-kamasu]
Thyrsitoides marleyi: Honma et al., 1990; Tsuda, 1990; Uozu Aquarium, 1997; Honma,
2013; Nakabo and Doiuchi, 2013.
Family Luvaridae*Luvarus imperialis Raﬁnesque, 1810 [Japanese name: Ama-shiira]
Luvarus imperialis: Ueno, 1971; Maeda and Tsutsui, 2003.
Family Trichiuridae*Assurger anzac (Alexander, 1917) [Japanese name: Nagayume-tachimodoki]
Assurger anzac: Kobayashi et al., 2006; Nakabo and Doiuchi, 2013.
*Benthodesmus tenius (Günther, 1877) [Japanese name: Tachimodoki]
Lepidopus tenuis: Honma, 1952.
Benthodesmus tenius: Ueno, 1971; Lindberg and Krasyukova, 1975; Tsuda, 1990; Maeda andTsutsui, 2003; Honma, 2013; Nakabo and Doiuchi, 2013.
*Eupleurogrammus sp. [Japanese name: Oshiroidachi]
Eupleurogrammus muticus [sic]: Tsuda, 1990.
*Evoxymetopon taeniatus Gill, 1863 [Japanese name: Yume-tachimodoki]
Evoxymetopon taeniatus: Mori, 1952; Honma et al., 1984; Tsuda, 1990; Honma, 2013;
Nakabo and Doiuchi, 2013.
Trichiurus japonicus Temminck and Schlegel, 1844 [Japanese name: Tachi-uo]
Trichiurus haumela: Katayama, 1940; Honma, 1952; Mori, 1952.
Trichiurus lepturus: Honma, 1956b; Kato, 1956; Mori, 1956a; Honma, 1963; Takegawa andMorino, 1970; Ueno, 1971; Lindberg and Krasyukova, 1975; Honma, 1977; Honma and Kitami,
1978; Tsuda, 1990; Adrianov and Kussakin, 1998; Sokolovskaya et al., 1998.
Trichiurus japonicus: Mori and Uchida, 1934; Suzuki et al., 2000; Maeda and Tsutsui, 2003;
Shinohara et al., 2011; Honma, 2013; Nakabo and Doiuchi, 2013.
SMBS uncatalogued (2).
Family Centrolophidae
Hyperoglyphe japonica (Döderlein, 1884) [Japanese name: Medai]
Ocycrius japonicus: Honma, 1952; Mori, 1952; Kato, 1956; Mori, 1956a; Honma, 1963;
Honma and Kitami, 1970; Takegawa and Morino, 1970; Ueno, 1971; Honma, 1977; Honma andKitami, 1978.

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Hyperoglyphe japonica: Lindberg and Krasyukova, 1975; Tsuda, 1990; Amaoka et al., 1995;
Suzuki et al., 2000; Maeda and Tsutsui, 2003; Kobayashi et al., 2006; Shinohara et al., 2011;
Honma, 2013; Nakabo and Doiuchi, 2013.
TS09-T006 (4), TS09-T013 (1), TS09-T105 (1), TS09-T141 (2), SMBS uncatalogued (1).
Icichthys lockingtoni Jordan and Gilbert, 1880 [Japanese name: Kuro-medai]
Icichthys lockingtoni: Nakabo and Doiuchi, 2013.
Remarks. According to Nakabo and Doiuchi (2013), the occurrence can be conﬁrmed fromSanʼin District, based on FAKU 58768 and 58769.
Psenopsis anomala (Temminck and Schlegel, 1844) [Japanese name: Ibodai]
Psenopsis anomala: Mori and Uchida, 1934; Katayama, 1940; Honma, 1952; Kato, 1956;
Mori, 1956a; Honma, 1963; Honma and Kitami, 1970; Takegawa and Morino, 1970; Lindberg
and Krasyukova, 1975; Honma, 1977; Honma and Kitami, 1978; Tsuda, 1990; Nakabo, 1993b;
Amaoka et al., 1995; Okamura, 1997; Uozu Aquarium, 1997; Nakabo, 2000b; Suzuki et al.,
2000; Nakabo, 2002b; Maeda and Tsutsui, 2003; Shinohara et al., 2011; Honma, 2013; Nakabo
and Doiuchi, 2013.
Psenopsis anomale [sic]: Ueno, 1971.
*Psenopsis shojimai Ochiai and Mori, 1965 [Japanese name: Nise-ibodai]
Psenopsis shojimai: Nakabo, 1993b; Nakabo and Doiuchi, 2013.
Family Nomeidae*Cubiceps whiteleggii (Waite, 1894) [Japanese name: Bōzu-konʼnyaku]
Cubiceps squamiceps: Honma, 1995; Kobayashi et al., 2006; Honma, 2013.
Cubiceps whiteleggii: Nakabo and Doiuchi, 2013.
Psenes cyanophrys Valenciennes, 1833 [Japanese name: Suji-hanabira-uo]
Psenes cyanophrys: Nakabo, 1993b; Okamura, 1997; Nakabo, 2000b, 2002b; Nakabo andDoiuchi, 2013.
Remarks. According to Nakabo and Doiuchi (2013), the occurrence can be conﬁrmed from
Wakasa Bay, based on FAKU 53317, 53318, 53278 and 54787.
Nomeus gronovii (Gmelin, 1789) [Japanese name: Eboshidai]
Nomeus gronovii: Tsuda, 1990; Suzuki et al., 2000; Nakabo and Doiuchi, 2013.
Psenes pellucidus Lütken, 1880 [Japanese name: Hanabira-uo]
Icticus ischanus: Katayama, 1940.
Icticus pellucidus: Kato, 1956; Mori, 1956a; Takegawa and Morino, 1970.
Psenes pellucidus: Lindberg and Krasyukova, 1975; Honma et al., 1984; Tsuda, 1990; Uozu
Aquarium, 1997; Suzuki et al., 2000; Kobayashi et al., 2006; Kitagawa et al., 2008; Shinohara etal., 2011; Honma, 2013; Nakabo and Doiuchi, 2013.
TS09-T141 (1).
Psenes arafurensis Günther, 1889 [Japanese name: Kurage-uo]
Psenes arafurensis: Nakabo, 1993b; Okamura, 1997; Nakabo and Doiuchi, 2013.
Remarks. According to Nakabo and Doiuchi (2013), the occurrence can be conﬁrmed fromthe western Sea of Japan, based on deposited in the Seikai National Fisheries Research Institute.
Family Ariommatidae*Ariomma indicum (Day, 1871) [Japanese name: Maru-ibodai]
Ariomma indica: Nakabo, 1993b; Okamura, 1997; Nakabo, 2000b; Nakabo and Doiuchi,

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2013.
*Ariomma luridum Jordan and Snyder, 1904 [Japanese name: Ōme-medai]
Ariomma lurida: Takegawa and Morino, 1970; Ueno, 1971.
Ariomma luridum: Maeda and Tsutsui, 2003; Nakabo and Doiuchi, 2013.
Family Tetragonuridae*Tetragonurus cuvieri Risso, 1810 [Japanese name: Doku-uroko-ibodai]
Tetragonurus cuvieri: Nakabo and Doiuchi, 2013.
Family Caproidae*Antigonia capros Lowe, 1843 [Japanese name: Hishidai]
Antigonia capros: Katayama, 1940; Mori, 1952; Kato, 1956; Mori, 1956a; Tsuda, 1990;
Nakabo and Kai, 2013.
Order Pleuronectiformes
Family Citharidae
Citharoides macrolepidotus Hubbs, 1915 [Japanese name: Koke-birame]
Citharoides macrolepidotus: Mori, 1952; Matsubara, 1955; Mori, 1956a; Amaoka, 1988;
Tsuda, 1990; Nambu et al., 1991; Lindberg and Fedorov, 1993; Nakabo, 1993b, 2000b, 2002b;
Voronina and Volkova, 2003; Shinohara et al., 2011; Nakabo and Doiuchi, 2013.
Family Paralichthyidae
Paralichthys olivaceus (Temminck and Schlegel, 1846) [Japanese name: Hirame]
Paralichthys olivaceus: Snyder, 1912; Kato, 1956; Ueno, 1971; Tsuda, 1990; Nakabo,
1993b; Sokolovskaya et al., 1998; Maeda and Tsutsui, 2003; Voronina and Volkova, 2003;
Nakabo and Doiuchi, 2013; Honma, 2013.
Pseudorhombus pentophthalmus Günther, 1862 [Japanese name: Tama-ganzō-birame]
Pseudorhombus pentophalmus [sic]: Ueno, 1971.
Pseudorhombus pentophthalmus: Kato, 1956; Tsuda, 1990; Nambu et al., 1991; Maeda andTsutsui, 2003; Voronina and Volkova, 2003; Honma, 2013.
Family Bothidae*Engyprosopon maldivensis (Regan, 1908) [Japanese name: Tenaga-darumagarei]
Engyprosopon maldivensis: Nakabo and Doiuchi, 2013.
*Japonolaeops dentatus Amaoka, 1969 [Japanese name: Hina-darumagarei]
Japonolaeops dentatus: Nakabo and Doiuchi, 2013.
Laeops kitaharae (Smith and Pope, 1906) [Japanese name: Yarigarei]
Laeops lanceolata: Mori, 1952; Honma, 1962; Ochiai and Amaoka, 1962; Honma, 1963;
Takegawa and Morino, 1970; Honma and Kitami, 1978; Tsuda, 1990; Nakabo and Doiuchi,
2013.
Laeops kitaharai: Mori, 1956a; Kato, 1956; Tsuda, 1990.
Laeops kitaharae: Amaoka, 1988; Nambu et al., 1991; Lindberg and Fedorov, 1993; Nakabo,
1993b; Amaoka, 1997; Nakabo, 2000b, 2002b; Honma, 2013; Nakabo and Doiuchi, 2013.

Annotated Checklist of Deep-sea Fishes of the Sea of Japan275

Family Pleuronectidae
Acanthopsetta nadeshnyi Schmidt, 1904 [Japanese name: Urokome-garei]
Acanthopsetta nadeshnyi: Mori and Uchida, 1934; Okada and Matsubara, 1938; Mori, 1952;
Honma, 1955d; Matsubara, 1955; Kato, 1956; Mori, 1956a, c; Honma, 1963; Ouchi, 1963;
Takegawa and Morino, 1970; Honma and Kitami, 1978; Sakamoto, 1988; Tsuda, 1990; Nambu
et al., 1991, 1992; Lindberg and Fedorov, 1993; Nakabo, 1993b; Amaoka et al., 1995; Sakamoto,
1997; Uozu Aquarium, 1997; Adrianov and Kussakin, 1998; Sokolovskaya et al., 1998; Nakabo,
2000b, 2002b; Maeda and Tsutsui, 2003; Voronina and Volkova, 2003; Evseenko, 2004;
Kitagawa et al., 2008; Shinohara et al., 2011; Honma, 2013; Nakabo and Doiuchi, 2013.
Acanthopsetta nadeshyni [sic]: Katayama, 1952.
Acanthopsetta nadeshny [sic]: Ueno, 1971.
KT-10-08-R1 (2), KT-10-08-R2 (3), KT-10-08-T1 (1), SMBS uncatalogued (1).
*Atheresthes evermanni Jordan and Starks, 1904 [Japanese name: Abura-garei]
Atheresthes evermanni: Katayama, 1940; Honma, 1952; Kato, 1956; Honma, 1963; Ouchi,
1963; Ueno, 1971; Honma and Kitami, 1978; Sakamoto, 1988; Tsuda, 1990; Lindberg andFedorov, 1993; Nakabo, 1993b; Amaoka et al., 1995; Sakamoto, 1997; Nakabo, 2000b, 2002b;
Maeda and Tsutsui, 2003; Kitagawa et al., 2008; Honma, 2013; Nakabo and Doiuchi, 2013.
Clidoderma asperrimum (Temminck and Schlegel, 1846) [Japanese name: Same-garei]
Clidoderma asperrimum: Snyder, 1912; Mori, 1928; Mori and Uchida, 1934; Katayama,
1940; Honma, 1952; Katayama, 1952; Kato, 1956; Mori, 1952, 1956a; Ueno, 1971; Tsuda, 1990;
Lindberg and Fedorov, 1993; Sakamoto, 1997; Adrianov and Kussakin, 1998; Adrianov and Kussakin, 1998; Suzuki et al., 2000; Maeda and Tsutsui, 2003; Voronina and Volkova, 2003;
Ivankova, 2007; Shinohara et al., 2011; Honma, 2013; Nakabo and Doiuchi, 2013.
Dexistes rikuzenius Jordan and Starks, 1904 [Japanese name: Migi-garei]
Dexistes rikuzenius: Mori and Uchida, 1934; Mori, 1952; Honma, 1955c; Matsubara, 1955;
Kato, 1956; Mori, 1956a; Honma, 1963; Honma and Kitami, 1978; Honma and Kitami, 1978;
Tsuda, 1990; Nambu et al., 1991; Lindberg and Fedorov, 1993; Sakamoto, 1997; Uozu Aquarium, 1997; Maeda and Tsutsui, 2003; Voronina and Volkova, 2003; Shinohara et al., 2011;
Honma, 2013; Nakabo and Doiuchi, 2013.
Eopsetta grigorjewi (Herzenstein, 1890) [Japanese name: Mushi-garei]
Xystrias grigorjewi: Mori, 1928; Mori and Uchida, 1934; Katayama, 1940; Honma, 1952;
Mori, 1952, 1956a.
Eopsetta grigorjewi: Honma, 1955b; Kato, 1956; Matsubara, 1963; Takegawa and Morino,
1970; Ueno, 1971; Honma and Kitami, 1978; Tsuda, 1990; Nambu et al., 1991; Lindberg andFedorov, 1993; Nakabo, 1993b; Amaoka et al., 1995; Sakamoto, 1997; Nakabo, 2000b; Suzuki etal., 2000; Nakabo, 2002b; Maeda and Tsutsui, 2003; Voronina and Volkova, 2003; Shinohara etal., 2011; Honma, 2013; Nakabo and Doiuchi, 2013.
Eopseta grigorjwei [sic]: Sokolovskaya et al., 1998.
SMBS uncatalogued (1).
Glyptocephalus stelleri (Schmidt, 1904) [Japanese name: Hireguro]
Glyptocephalus sasæ: Snyder, 1912.
Microstomus hireguro: Tanaka, 1917.
Tanakius hireguro: Mori and Uchida, 1934.
Glyptocephalus stelleri: Katayama, 1940; Honma, 1952; Katayama, 1952; Mori, 1952;
Matsubara, 1955; Kato, 1956; Mori, 1956a, 1956c; Honma, 1963; Ouchi, 1963; Honma and276

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Kitami, 1970; Ueno, 1971; Ogata et al., 1973; Honma, 1977; Honma and Kitami, 1978; Sakamoto, 1988; Tsuda, 1990; Nambu et al., 1991, 1992; Nakabo, 1993b; Amaoka et al., 1995; Sakamoto, 1997; Adrianov and Kussakin, 1998; Adrianov and Kussakin, 1998; Nakabo, 2000b;
Suzuki et al., 2000; Nakabo, 2002b; Maeda and Tsutsui, 2003; Voronina and Volkova, 2003; ;Evseenko, 2004; Shinohara et al., 2011; Honma, 2013; Nakabo and Doiuchi, 2013.
KT-11-09-N3 (3), KT-11-09-T1 (1), TS09-T004 (1), TS09-T011 (1), TS09-T102 (5), TS09T107 (7), SMBS uncatalogued (1).
Hippoglossoides dubius Schmidt, 1904 [Japanese name: Aka-garei]
Hippoglossoides katakuræ: Snyder, 1912.
Hippoglossoides dubius: Snyder, 1912; Mori and Uchida, 1934; Okada and Matsubara, 1938;
Katayama, 1940; Honma, 1952; Mori, 1952; Matsubara, 1955; Kato, 1956; Mori, 1956a, c;
Honma, 1963; Ouchi, 1963; Takegawa and Morino, 1970; Ueno, 1971; Ogata et al., 1973;
Honma and Kitami, 1978; Sakamoto, 1988; Tsuda, 1990; Nambu et al., 1991; Nambu et al.,
1992; Amaoka et al., 1995; Sakamoto, 1997; Sokolovskaya et al., 1998; Nakabo, 2000b; Suzuki
et al., 2000; Nakabo, 2002b; Maeda and Tsutsui, 2003; Voronina and Volkova, 2003; ; Evseenko,
2004; Kitagawa et al., 2008; Shinohara et al., 2011; Honma, 2013; Nakabo and Doiuchi, 2013.
Cynopsetta dubia: Tanaka, 1913b; Katayama, 1952.
Hippoglossoides (Hippoglossoides) dubius: Lindberg and Fedorov, 1993; Adrianov and Kussakin, 1998.
SMBS uncatalogued (1).
Hippoglossoides elassodon Jordan and Gilbert, 1881 [Japanese name: Uma-garei]
Hippoglossoides elassodon: Kato, 1956; Mori, 1956a; Ueno, 1971; Tsuda, 1990; Nambu etal., 1991; Amaoka et al., 1995; Maeda and Tsutsui, 2003; Voronina and Volkova, 2003; Shinohara et al., 2011; Nakabo and Doiuchi, 2013.
Hippoglossoides (Hippoglossoides) elassodon: Lindberg and Fedorov, 1993.
Hippoglossoides pinetorum (Jordan and Starks, 1904) [Japanese name: Sōhachi]
Cleisthenes herzensteini: Mori, 1928; Mori and Uchida, 1934; Okada and Matsubara, 1938;
Katayama, 1940; Honma, 1952; Mori, 1952, 1956a; Ueno, 1971; Voronina and Volkova, 2003.
Cleithenes pinetorum: Okada and Matsubara, 1938; Kato, 1956; Takegawa and Morino,
1970; Amaoka et al., 1995; Voronina and Volkova, 2003.
Protopsetta herzensteini: Katayama, 1952.
Cleithenes pinetorum herzensteini: Matsubara, 1955; Kato, 1956; Honma, 1963; Ouchi,
1963; Honma and Kitami, 1978; Tsuda, 1990.
Hippoglossoides pinetorum: Nambu et al., 1991; Sakamoto, 1997; Nakabo, 1993b, 2000b;
Suzuki et al., 2000; Nakabo, 2002b; Maeda and Tsutsui, 2003; Kobayashi et al., 2006; Shinohara
et al., 2011; Honma, 2013; Nakabo and Doiuchi, 2013.
Hippoglossoides (Cleithenes) herzensteini: Lindberg and Fedorov, 1993; Adrianov and Kussakin, 1998; Sokolovskaya et al., 1998.
TS09-T011 (1).
Hippoglossus stenolepis Schmidt, 1904 [Japanese name: Ohyō]
Hippoglossus stenolepis: Okada and Matsubara, 1938; Katayama, 1940; Honma, 1952; Kato,
1956; Ueno, 1971; Tsuda, 1990; Nakabo, 1993b; Sakamoto, 1997; Uozu Aquarium, 1997; Sokolovskaya et al., 1998; Nakabo, 2000b, 2002b; Maeda and Tsutsui, 2003; Shinohara et al., 2011;
Honma, 2013; Nakabo and Doiuchi, 2013.
Hippoglossus hippoglosus stereolepis: Adrianov and Kussakin, 1998.
Microstomus achne (Jordan and Starks, 1904) [Japanese name: Baba-garei]

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Microstomus achne: Okada and Matsubara, 1938; Honma, 1952; Mori, 1952; Matsubara,
1955; Kato, 1956; Mori, 1956a; Honma and Kitami, 1970; Ueno, 1971; Honma and Kitami,
1978; Honma and Kitami, 1978; Sakamoto, 1988; Tsuda, 1990; Lindberg and Fedorov, 1993;
Nakabo, 1993b; Amaoka et al., 1995; Sakamoto, 1997; Adrianov and Kussakin, 1998; Adrianov
and Kussakin, 1998; Nakabo, 2000b; Suzuki et al., 2000; Nakabo, 2002b; Maeda and Tsutsui,
2003; Voronina and Volkova, 2003; Shinohara et al., 2011; Honma, 2013; Nakabo and Doiuchi,
2013.
Pleuronectes asper Pallas, 1814 [Japanese name: Kogane-garei]
Limanda asper: Tsuda, 1990; Voronina and Volkova, 2003.
Pleuronectes (Limanda) asper: Lindberg and Fedorov, 1993; Adrianov and Kussakin, 1998;
Sokolovskaya et al., 1998.
Pleuronectes asper: Maeda and Tsutsui, 2003; Shinohara et al., 2011; Nakabo and Doiuchi,
2013.
Pleuronectes bilineatus (Ayres, 1855) [Japanese name: Shumushu-garei]
Lepidopsetta bilineata: Okada and Matsubara, 1938; Ouchi, 1963.
Pleuronectes bilineata: Honma, 1952; Matsubara, 1955.
Pleuronectes bilineatus: Sakamoto, 1988; Tsuda, 1990; Nambu et al., 1991; Nakabo, 1993b;
Amaoka et al., 1995; Sakamoto, 1997; Nakabo, 2000b, 2002b; Maeda and Tsutsui, 2003; Shinohara et al., 2011; Honma, 2013; Nakabo and Doiuchi, 2013.
Pleuronectes quadrituberculatus Pallas, 1814 [Japanese name: Tsuno-garei]
Pleuronectes quadrituberculatus: Sakamoto, 1988; Nambu et al., 1991; Nakabo, 1993b;
Sakamoto, 1997; Nakabo, 2000b, 2002b; Maeda and Tsutsui, 2003; Voronina and Volkova, 2003;
Shinohara et al., 2011; Honma, 2013; Nakabo and Doiuchi, 2013.
Pleuronectes pallasii: Tsuda, 1990.
Pleuronectes (Pleuronectes) quadrituberculatus: Lindberg and Fedorov, 1993; Adrianov andKussakin, 1998.
*Pleuronectes sakhaliensis (Hubbs, 1915) [Japanese name: Karafuto-garei]
Limanda korigarei: Ueno, 1971.
Pleuronectes sakhaliensis: Nakabo and Doiuchi, 2013.
*Reinhardtius hippoglossoides (Walbaum, 1792) [Japanese name: Karasu-garei]
Reinhardtius matsuurai: Kato, 1956.
Reinhardtius matsuurae: Mori, 1956a.
Reinhardtius hippoglossoides: Ueno, 1971; Sakamoto, 1988; Tsuda, 1990; Nakabo, 1993b;
Nambu et al., 1991; Amaoka et al., 1995; Sakamoto, 1997; Nakabo, 2000b, 2002b; Maeda andTsutsui, 2003; Kitagawa et al., 2008; Honma, 2013; Nakabo and Doiuchi, 2013.
Reinhardtius hippoglossoides natsuurae: Lindberg and Fedorov, 1993.
Tanakius kitaharae (Jordan and Starks, 1904) [Japanese name: Yanagi-mushigarei]
Tanakius kitaharae: Katayama, 1940; Honma, 1952; Katayama, 1952; Mori, 1952; Matsubara, 1955; Lindberg and Fedorov, 1993; Nakabo and Doiuchi, 2013.
Tanakius kitaharai: Kato, 1956; Mori, 1956a; Ouchi, 1963; Honma and Kitami, 1970; Ueno,
1971; Honma and Kitami, 1978; Tsuda, 1990; Nambu et al., 1991; Suzuki et al., 2000; Maeda
and Tsutsui, 2003; Voronina and Volkova, 2003; Shinohara et al., 2011; Honma, 2013.
SMBS uncatalogued (1).
Verasper moseri Jordan and Gilbert, 1898 [Japanese name: Matsukawa]
Verasper moseri: Kato, 1956; Ueno, 1971; Sakamoto, 1988; Tsuda, 1990; Amaoka et al.,
1995; Sakamoto, 1997; Adrianov and Kussakin, 1998; Sokolovskaya et al., 1998; Maeda and278

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Tsutsui, 2003; Kitagawa et al., 2008; Voronina and Volkova, 2003.
Verasper variegatus (Temminck and Schlegel, 1846) [Japanese name: Hoshi-garei]
Verasper variegatus: Kato, 1956; Ueno, 1971; Sakamoto, 1988; Tsuda, 1990; Sakamoto,
1997; Adrianov and Kussakin, 1998; Sokolovskaya et al., 1998; Maeda and Tsutsui, 2003; Voronina and Volkova, 2003; Kitagawa et al., 2008; Honma, 2013; Nakabo and Doiuchi, 2013.
Family Poecilopsettidae
Poecilopsetta plinthus (Jordan and Starks, 1904) [Japanese name: Kawara-garei]
Poecilopsetta plinthus: Honma and Mizusawa, 1966; Honma and Kitami, 1978; Sakamoto,1988; Lindberg and Fedorov, 1993; Tsuda, 1990; Nambu et al., 1991; Suzuki et al., 2000; Voronina and Volkova, 2003; Shinohara et al., 2011; Nakabo and Doiuchi, 2013.
Poecilopsetta pinthus [sic]: Honma, 2013.
Family Samaridae*Samariscus latus Matsubara and Takamuki, 1951 [Japanese name: Tsumari-tsukinowa-garei]
Samariscus latus: Sakamoto, 1988, 1997; Nakabo and Doiuchi, 2013.
Family Cynoglossidae*Cynoglossus ochiaii Yokogawa, Endo and Sakaji, 2008 [Japanese name: Oki-genko]
Cynoglossus ochiaii: Yamada and Yagishita, 2013.
*Symphurus orientalis (Bleeker, 1879) [Japanese name: Azuma-garei]
Symphurus orientalis: Okada and Matsubara, 1938; Yamada and Yagishita, 2013.
Order Tetraodontiformes
Family Triacanthodidae*Macrorhamphosodes uradoi (Kamohara, 1933) [Japanese name: Fue-kawamuki]
Macrorhamphoodes uradoi: Hayashi and Hagiwara, 2013.
Triacanthodes anomalus (Temminck and Schlegel, 1850) [Japanese name: Beni-kawamuki]
Triacanthodes anomalus anomalus: Honma and Mizusawa, 1966.
Triacanthodes anomalus: Katayama, 1940; Kato, 1956;Takegawa and Morino, 1970; Tsuda,
1990; Lindberg et al., 1997; Uozu Aquarium, 1997; Suzuki et al., 2000; Kobayashi et al., 2006;
Shinohara et al., 2011; Hayashi and Hagiwara, 2013; Honma, 2013.
Family Monacanthidae
Thamnaconus hypargyreus (Cope, 1871) [Japanese name: Sarasa-hagi]
Canthrrinus tessellatus: Honma, 1952.
Cantherines tessellatus: Mori, 1952.
Novodon tessellatus [sic]: Kato, 1956.
Navodon tessellatus: Honma and Kitami, 1970; Honma and Chiba, 1978; Honma andKitami, 1978; Tsuda, 1990.
Thamnaconus hypargyreus: Suzuki et al., 2000; Hayashi and Hagiwara, 2013; Honma, 2013.
TS09-T107 (2).

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Family Ostraciidae
Kentrocapros aculeatus (Houttuyn, 1782) [Japanese name: Itomaki-fugu]
Kentrocapros aculeatus: Mori and Uchida, 1934; Kato, 1956; Mori, 1956a; Takegawa andMorino, 1970; Honma and Chiba, 1978; Tsuda, 1990; Lindberg et al., 1997; Uozu Aquarium,
1997; Suzuki et al., 2000; Shinohara et al., 2011; Hayashi and Hagiwara, 2013; Honma, 2013.
Aracana aculeata: Katayama, 1940; Honma, 1952; Mori, 1952.
Family Triodontidae
Triodon macropterus Lesson, 1831 [Japanese name: Uchiwa-fugu]
Triodon bursarius: Katayama, 1940; Mori, 1952; Kato, 1956.
Triodon macropterus: Tsuda, 1990; Lindberg et al., 1997; Uozu Aquarium, 1997; Nakabo
and Doiuchi, 2013.
Family Tetraodontidae
Sphoeroides pachygaster (Müller and Troschel, 1848) [Japanese name: Yorito-fugu]
Liosaccus cutaneous [sic]: Matsubara, 1955.
Liosaccus cutaneus: Honma and Kitami, 1970; Honma et al., 1972; Honma and Chiba, 1978;
Honma and Kitami, 1978; Tsuda, 1990.
Liosarcus pachygater: Honma, 1991a.
Sphoeroides pachygaster: Lindberg et al., 1997; Uozu Aquarium, 1997; Suzuki et al., 2000;
Kobayashi et al., 2006; Shinohara et al., 2011; Honma, 2013; Yamada and Yagishita, 2013.
SMBS uncatalogued (1).
General Remarks
In summary, a total of 363 species in 104 families and 29 orders was included in our list.
Compared to deep-sea ﬁshes of Suruga Bay (Shinohara and Matsuura, 1997: 483 spp.), Tosa Bay(Shinohara et al., 2001: 599 spp.), the Ryukyu Islands (Shinohara et al., 2005: 644 spp.) and the
Paciﬁc coast off Tohoku District (Shinohara et al., 2009: 496 spp.), the species diversity of deepsea ﬁshes is clearly lower in the Sea of Japan. Although the limited diversity of deep-sea ﬁshes ispossibly due to several factors, the extremely small number of oceanic deep-sea ﬁshes has also
contributed to the depauperate deep-sea ﬁsh fauna (e.g., Shinohara et al., 2011).
Our list revealed that relatively speciose families in the Sea of Japan are Cottidae (30 spp.),
Zoarcidae (29 spp.), Liparidae (20 spp.), Pleuronectidae (19 spp.), Scorpaenidae (15 spp.), Agonidae (13 spp.) and Stichaeidae (12 spp). The ﬁve most speciose families include about 30% of theentire species in the list. According to Shinohara and Matsuura (1997) and Shinohara et al.
(2001, 2005, 2009), the Sea of Japan is thought to be unique in having the highest number of
species in the Cottidae (30 vs. 9 in Paciﬁc off Tohoku, 0 in Ryukyu Islands, 4 in Tosa Bay and 5
in Suruga Bay), Zoarcidae (29 vs. 20, 1, 4 and 2), Liparidae (20 vs. 14, 3, 2 and 3), Pleuronectidae (19 vs. 14, 8, 14 and 7) and Stichaeidae (12 vs. 5, 0, 1 and 2). The unique fauna of the Sea ofJapan also includes 9 endemic species (Figs. 2, 4), i.e., Icelus toyamensis (Cottidae), Occellakuronumai (Agonidae), Careproctus acanthodes, C. entargyneus, C. entomelas and C. notosaikaiensis (Liparidae), Krusenterniella maculata and Lycodes nishimurai (Zoarcidae) and Xenolumpenus longipterus (Stichaeidae).

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Acknowledgments
We express our thanks to captains and all ofﬁcers and crews of the R/Vs Soyo-maru (FRA)
and Tansei-maru (JAMSTEC), T/V Tanshu-maru (Kasumi High School), Ken Fujimoto, Takahiro Kinoshita, Kunihiro Fujiwara and Ikuko Yosho (FRA), Masanori Okanishi (Kyoto University), Junko Inoue and Kei Sakuma (the University of Tokyo) and Tsunemi Kubodera, Toshiaki
Kuramochi and Toshihiko Fujita (NSMT) for their support in collecting specimens. Special
thanks go to Satoshi Awata (Niigata University) for giving us an opportunity to examine SMBS
specimens, Yoshiaki Kai (FAKU) for permission to use a color photograph of Careproctus notosaikaiensis (Fig. 4A) and Kenta Suda (Okabe Co., Ltd) for literature on the hexanchinid sharks.
Edward Murdy (George Washington University) kindly reviewed the manuscript.
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