Fauna Ryukyuana

ISSN 2187-6657
http://w3.u-ryukyu.ac.jp/naruse/lab/Fauna_Ryukyuana.html

First records of the Cocoa Snapper Paracaesio stonei (Perciformes: Lutjanidae)
from the Satsunan Islands, Japan
1

Harutaka Hata1, Hiroki Iwatsubo2 and Hiroyuki Motomura3

The United Graduate School of Agricultural Sciences, Kagoshima University,
1–21–24 Korimoto, Kagoshima 890–0065, Japan
2
Kagoshima Museum of Aquatic Biodiversity, Kagoshima МS Building,
11–21 Nishisengoku, Kagoshima 892–0847, Japan
3
The Kagoshima University Museum, 1-21-30 Korimoto, Kagoshima 890-0065, Japan

Abstract. Three specimens of Paracaesio stonei(Perciformes: Lutjanidae) were collected from theOsumi and Tokara islands, included in the Satsunan
Islands, Kagoshima Prefecture, southern Japan.
These represent the first records of the species from
Tokara Islands, and the Osumi specimen is thenorthernmost record for the species. A detailed
description of the three specimens is provided.
Introduction
The snapper genus Paracaesio Bleeker, 1875 iscurrently represented in Japanese waters by fivespecies, viz., P. caerulea (Katayama, 1934) “Aodai”,
P. kusakarii Abe, 1960 “Shima-aodai”, P. sordida
Abe & Shinohara, 1962 “Yogore-aodai”, P. stonei
Raj & Seeto, 1983 “Yambaru-shimaaodai” and P.
xanthura (Bleeker, 1869) “Umeiro” (Shimada 2013).
All except for P. stonei have already been recorded
from the Satsunan Islands, northern Ryukyu Islands(Ichikawa et al. 1992; Motomura et al. 2010; Meguro2013; Sakurai 2014; Eguchi & Motomura 2016).
Japanese records of P. stonei have been limited to the
Okinawa Islands and southward (Shimada 2013).
During ichthyofaunal surveys in the Tokara
Islands, conducted as a part of the Kagoshima Fish
Diversity Project, three specimens of P. stonei werecollected from Yaku-shima (Osumi Islands), andTaira-jima and Kodakara-jima (Tokara Islands). The
present study describes these specimens in detail.
This report represents the first records of P. stonei
from the Satsunan Islands, the Yaku-shima specimen
representing the northernmost record of the species.
Counts and proportional measurements follow thoseof White & Last (2012). All measurements weremade with digital calipers to the nearest 0.1 mm.
Abbreviations: SL (standard length) and KAUM
(Kagoshima University Museum, Kagoshima).
Specimens of P. kusakarii (listed below) were used
for comparative purpose to confirm validity ofcharacters of P. stonei. Japanese words “-jima” or “shima” mean “island”.

RecordParacaesio stonei Raj & Seeto, 1983
Standard Japanese name: Yambaru-shimaaodai(Fig. 1; Appendix 1)
Material examined. KAUM–I. 54128, 379.7 mm
SL, off Kodakara-jima , Tokara Islands, Kagoshima
Prefecture, Japan, 22 Apr. 2013, line-fishing(purchased at Kagoshima City Central Fish Market),
M. Matsunuma; KAUM–I. 82741, 407.8 mm SL, offTaira-jima, Tokara Islands, Kagoshima Prefecture,
Japan, ca. 100 m depth, line-fishing (purchased at
Kagoshima City Central Fish Market), 8 Dec. 2015,
H. Hata and T. Naito; KAUM–I. 200222, 220.7 mm
SL, off Yaku-shima, Osumi Islands, Kagoshima
Prefecture, Japan, line-fishing (purchased at
Kagoshima City Central Fish Market), 15 Jan. 2015,
H. Iwatsubo and T. Tanaka.
Description. Body oblong, compressed laterally,
deepest at bases of 2nd to 4th dorsal-fin spines.
Dorsal contour of body increasing from snout tip tobases of 3rd to 5th dorsal-fin spines, thereafter
decreasing steadily to caudal-fin base. Ventral profile
of body decreasing from lower-jaw tip to pelvic-fin
origin, thereafter more or less horizontal to anal-fin
origin, subsequently rising to caudal-fin base.
Uppermost and lowermost points of pectoral-fin
insertion anterior to posteriormost point of opercle
and dorsal-fin origin, respectively. Posterior tip of
pectoral fin pointed, extending beyond line verticalthrough origin of 1st dorsal-fin soft ray. Dorsal-fin
origin posterior to posteriormost point of opercle.
End of dorsal-fin base just above that of anal-fin base.
Contour of spinous portion of dorsal fin moderately
notched. Origin of pelvic fin just below origin of 2nd
dorsal-fin spine. Posterior tip of depressed pelvic finjust reaching or slightly short of anus. Anal-fin origin
between vertical lines through bases of 1st to 3rd
dorsal-fin soft rays. Caudal fin truncate, upper tip
slightly extended posteriorly. Body covered with
ctenoid scales. Anus situated just anterior to anal-fin
origin. All fins, both jaws, snout, pectoral-fin
55

Fig. 1. Fresh specimens of Paracaesio stonei (A: KAUM–I. 200222, 220.7 mm SL, Yaku-shima, Osumi Islands,
Kagoshima Prefecture, Japan; B: KAUM–I. 54128, 379.7 mm SL, Kodakara-jima, Tokara Islands, Kagoshima Prefecture,
Japan; C. KAUM–I. 82741, 407.8 mm SL, Taira-jima, Tokara Islands, Kagoshima Prefecture, Japan).
図 1. ヤンバルシマアオダイ Paracaesio stonei (A: KAUM–I. 200222, 標準体長 220.7 mm, 屋久島近海で漁獲; B:
KAUM–I. 54128, 標準体長 379.7 mm, 小宝島近海で漁獲; C: KAUM–I. 82741, 標準体長 407.8 mm, 平島近海水
深約 100 m で漁獲).
56[報告] 畑ら: ヤンバルシマアオダイの薩南諸島からの記録

Fauna Ryukyuana, 36: 55–62.

axillary region, part of area around eye and posterior
part of preopercle without scales. Eye and iris round.
Interorbital space flat, scaleless. Nostrils
paired,positioned close together, anterior to orbit.
Both nostrils oval. Dermal flap on posterior edge ofanterior nostril. Mouth terminal, small, posterior tipof maxilla extending beyond vertical line throughanterior margin of eye but not to anterior margin ofiris. Lower jaw projecting slightly beyond upper jaw.
Posterior and lower edges of preopercle serrated.
Posterior edge of opercle smooth. Single spine on
opercle. Anterior margin of scales on top of headextending forward to vertical line through iris center.
Upper and lower jaws with single outer row ofconical teeth and inner band of fine teeth. Triangular
patch of villiform teeth on vomer, fairly broad bandof villiform teeth on palatines. Tongue edentate. Gillrakers long, slender. Pseudobranchial filaments
present. Lateral line complete, running parallel todorsal body contour.
Color when fresh. Body purplish-gray with five
broad olive bands from dorsal surface to body
midline. Posteriormost band indistinct in larger two
specimens (379.7 and 407.8 mm SL). Dorsal surface
olive. Dorsal fin yellow. Upper lobe of caudal fin
grayish-white, lower lobe yellow. Pelvic and anal
fins dusky yellow. Pectoral fin semi-transparent. Iris
copper brown, pupil bluish-black (Fig. 1).
Color in alcohol. Body pale silver, dark purple
dorsally. Bands on body blackish.
Distribution. Paracaesio stonei has beenrecorded from southern Japan, southern Taiwan,
Philippines, northeastern Australia, eastern Papua
New Guinea, Vanuatu, Tuvalu, Fiji and Samoa (Raj
& Seeto 1983; Allen 1985; Chapman & Cuack 1990;
Kramer et al. 1994; Anderson & Allen 2001; Brewer
et al. 2007; Shimada 2013; this study). Previous
Japanese records have been restricted to the Okinawa
Islands and southward (Yoshino 1984; Shimada2013); the occurrence of the species at Kodakarajima and Taira-jima islands (Tokara Islands), andYaku-shima (Osumi Islands) represent new records
from each island.
Remarks. The three specimens examined in thisstudy were identified as P. stonei on the basis of thefollowing combination of characters, which closely
match the diagnostic features of P. stonei given by
Raj & Seeto (1983), Allen (1985), Anderson & Allen(2001), White & Last (2012) and Shimada (2013):
48 or 49 pored lateral-line scales; truncate caudal fin;scaleless maxilla; and body with five broad olive
colored bands from the dorsum to the body midline.
Anderson & Allen (2001) gave the number of total[Report] Hata et al.: Paracaesio stonei from Satsunan Islands

Fig. 2. Fresh specimen of Paracaesio kusakarii, KAUM–
I. 77301, 352.0 mm SL, Kuchino-shima, Tokara Islands,
Kagoshima Prefecture, Japan.
図 2. シマアオダイ Paracaesio kusakarii, KAUM–I.
77301, 標準体長 352.0 mm, 口之島北方で漁獲.

gill rakers on first gill arch of the species as 27–29,
but those of the specimens collected from theSatsunan Islands were 25–26. Ranges of the numbersof gill rakers of more than five have been reported in
the other congeners (P. kusakarii, P. sordida and P.
xanthura; Anderson & Allen, 2001). Therefore, we
judged that this difference is intraspecific variation.
The genus Paracaesio contains nine valid species(Allen 1985; Anderson et al. 1992; White & Last
2012). Although P. stonei is most similar to P.
kusakarii in sharing a similar count of pored lateralline scales, emarginated caudal fin and broad bandson the body (Allen 1985; Anderson & Allen 2001;
White & Last 2012; Shimada 2013), the former ischaracterized by a scaleless maxilla (vs. maxilla
scaled in P. kusakarii) and body with five bands, theposteriormost band indistinct in very large
individuals (vs. body with four bands; Fig. 2; Allen
1985; Anderson & Allen 2001; Shimada 2013; thisstudy).
Distributional records of P. stonei in Japanesewaters were reviewed here on the basis of publishedliterature. The first record of the species fromJapanese waters was that of Yoshino (1984), whoreported P. stonei from the Ryukyu Islands, including
a color photograph, and proposed the Japanese name“Yambaru-shimaaodai”. However, the collection siteof the photographed specimen was not referred.
Thereafter, Yoshino & Arakaki (1984) and Isa &Arakaki (1991) reported that P. stonei wasinfrequently collected in Okinawa Prefecture, again
without giving precise locality data. Shimada (2013)stated that the species is distributed in the Okinawa
Islands and southward, but no evidence was given.
No precise locality records in the Ryukyu Islands ofthe species were found during the present study.
Accordingly, the present specimens represent a57

northward extension of the distributional range forthe species.
Although large-sized lutjanid fishes including P.
stonei are important for fisheries in Kagoshima andOkinawa prefectures, detailed distributional
information for several Ryukyu Islands species
remains limited [e.g., Etelis radiosus Anderson,
1981 “Okuchihamadai”, Pristipomoides flavipinnis
Shinohara, 1963 “Kimmehimedai”, Pristipomoides
multidens (Day, 1871) “Nagasakifuedai”; Hata et al.
2015, 2016a–b]. Further taxonomic and faunal
studies on lutjanids are necessary.
Comparative material examined. Paracaesio
kusakarii: KAUM–I. 48824, 364.1 mm SL, off Sata
Point, Osumi Strait, Kagoshima Prefecture, Japan,
long-line fishing (purchased at Nishinoomote Fish
Market), 9 July 2012, M. Takayama; KAUM–I.
54120, 316.9 mm SL, off Amami-oshima, Amami
Islands, Kagoshima Prefecture, Japan, ca. 200 m
depth, line fishing (purchased at Kagoshima City
Central Fish Market), 20 Apr. 2013, M. Matsunuma;
KAUM–I. 54129, 456.5 mm SL, off Kodakara-jima,
Tokara Islands, Kagoshima Prefecture, Japan, line
fishing (purchased at Kagoshima City Central Fish
Market), 22 Apr. 2013, M. Matsunuma; KAUM–I.
55566, 292.0 mm SL, off Nakano-shima, Tokara
Islands, Kagoshima Prefecture, Japan, more than 100
m depth, line fishing (purchased at Kagoshima City
Central Fish Market), 19 July 2013, M. Matsunuma;
KAUM–I. 63111, 375.9 mm SL, south of Mageshima, Osumi Islands, Kagoshima Prefecture, Japan,
110 m depth, line fishing (purchased at
Nishinoomote Fish Market), 18 Aug. 2014, M.
Takayama; KAUM–I. 65664, 298.1 mm SL, off
Nishinoomote Port, Nishinoomote, Tanega-shima,
Osumi Islands, Kagoshima Prefecture, Japan, line
fishing (purchased at Nishinoomote Fish Market), M.
Takayama; KAUM–I. 74386, 204.2 mm SL,
KAUM–I. 74387, 202.8 mm SL, off Kumano
Fishing Port, Sakai, Nakatane, Tanega-shima, Osumi
Islands, Kagoshima Prefecture, Japan, 100 m depth,line fishing (purchased at Nishinoomote Fish
Market), 23 June 2015, M. Takayama; KAUM–I.
77301, 352.0 mm SL, north of Kuchino-shima,
Tokara Islands, Kagoshima Prefecture, Japan, line
fishing (purchased at Kagoshima City Central Fish
Market), 31 July 2015, H. Hata; KAUM–I. 91794,
259.8 mm SL, off Iloilo, Panay Island, thePhilippines (purchased at fish market in Oton), 11
Sept. 2016; KAUM–I. 200220, 218.5 mm SL, off
Tanega-shima,
Osumi
Islands,
Kagoshima
Prefecture, Japan, line fishing (purchased at
Kagoshima City Central Fish Market), 11 Oct. 2014,
H. Iwatsubo and T. Tanaka; KAUM–I.200221, 224.658mm SL, off Yaku-shima, Osumi Islands, Kagoshima
Prefecture, Japan, line fishing (purchased at
Kagoshima City Central Fish Market), 11 Oct. 2014,
H. Iwatsubo and T. Tanaka.
Paracaesio stonei: KAUM–I. 91795, 249.5 mm
SL, off Iloilo, Panay Island, the Philippines(purchased at fish market in Oton), 11 Sept. 2016.
Acknowledgements.
We are especially grateful to T. Tanaka (Tanaka
Suisan Ltd.) for collecting the specimens. We also
thank Y. Haraguchi and other volunteers, andstudents of KAUM for their curatorial assistance. We
are also grateful to G. Hardy (Ngunguru, New
Zealand), who read the manuscript and provided
help with English. Thanks also go to two reviewers
who improved our manuscript. This study was
supported in part by the Sasakawa Scientific
Research Grant from the Japan Science Society (28745); a Grant-in-Aid from the Japan Society for the
Promotion of Science for JSPS Fellows (DC2: 296652); JSPS KAKENHI Grant Numbers
JP26241027, JP24370041, JP23580259, andJP26450265; the JSPS Core-to-Core Program: B
Asia-Africa Science Platforms; the “Coastal Area
Capability Enhancement in Southeast Asia Project”
of the Research Institute for Humanity and Nature,
Kyoto, Japan; the “Biological Properties ofBiodiversity Hotspots in Japan” project of the
National Museum of Nature and Science, Tsukuba,
Japan; “Establishment of Research and Education
Network on Biodiversity and Its Conservation in theSatsunan Islands” project of Kagoshima University
adopted by the Ministry of Education, Culture,
Sports, Science and Technology, Japan; and “Guide
of Marketed Fish in Kagoshima” project ofKagoshima Museum of Aquatic Biodiversity. This
study was conducted under a Memorandum of
Agreement for joint research made by and among the
Department of Agriculture of the Republic of thePhilippines (DA), the University of the PhilippinesVisayas (UPV), the Kagoshima University Museum,the Research Institute for Humanity and Nature, andTokai University, facilitated by S. L. Sanchez[Bureau of Fisheries and Aquatic Resources (BFAR),
DA]. P. J. Alcala (DA) provided a Prior Informed
Consent Certificate, and I. P. Cabacaba and S. M. S.
Nolasco (BFAR, DA) a fish specimen Export
Certificate (No. 2016-39812). We thank the staff of
the Office of the Vice-Chancellor for Research andExtension, UPV, and UPV Museum of Natural
Sciences, College of Fisheries, UPV, including R. P.
Babaran, S. S. Garibay, U. B. Alama, V. G. Urbina,[報告] 畑ら: ヤンバルシマアオダイの薩南諸島からの記録

Fauna Ryukyuana, 36: 55–62.

L. H. Mooc, C. J. N. Rubido, E. P. Abunal, A. M. T.
Guzman, R. S. Cruz, A. C. Gaje, and R. F. M.
Traifalgar, and graduate students of the College ofFisheries, UPV for their support of this researchcollaboration.
References
Allen, G. R., 1985. FAO species catalogue. Vol. 6.
Snappers of the world. An annotated andillustrated catalogue of lutjanid species known todate. FAO Fisheries Synopsis, 6 (125): i–vi + 1–
303.
Anderson, W. D. & G. R. Allen, 2001. Lutjanidae
Snappers (jobfishes). In: K. E. Carpenter & V. H.
Niem (eds.), FAO species identification guide forfishery purposes. The living marine resources of
the western central Pacific. Vol. 5. Bony fishespart 3 (Menidae to Pomacentridae), Pp. 2840–
2918. FAO, Rome.
Anderson, W. D. Jr., P. J. Kailola & B. B. Collette,
1992. Two new snappers (Teleostei: Lutjanidae:
Apsilinae):
Paracaesio
paragrapsimodon
Anderson and Kailola from the western Pacificand P. waltervadi Anderson and Collette from thewestern Indian Ocean. Proceedings of the
Biological Society of Washington, 105 (3): 443–
461.
Brewer, D. T., D. A. Milton, G. C. Fry, D. M. Dennis,
D. S. Heales & W. N. Venables, 2007. Impacts ofgold mine waste disposal on deepwater fish in apristine tropical marine system. Marine Pollution
Bulletin, 54: 309–321.
Chapman, L. B. & P. Cusack. 1990. South PacificCommission Deep Sea Fisheries Development
Project. Report on second visit to Tuvalu 30
August–7 December 1983. South PacificCommission Noumea.
Eguchi, K. & H. Motomura, 2016. First records of
Paracaesio sordida (Perciformes: Lutjanidae)
from Tanega-shima and Amami-oshima islands,
Kagoshima Prefecture, southern Japan. Nature ofKagoshima, 42: 219–223.
Hata, H., Y. Haraguchi & H. Motomura, 2015. First
record of Etelis radiosus (Perciformes:
Lutjanidae) from the Tokara Islands in the
Ryukyu Islands, southern Japan. Nature ofKagoshima, 41: 95–99.
Hata, H. & H. Motomura, 2016a. First specimenbased records of Pristipomoides flavipinnis(Perciformes: Lutjanidae) from the Tokara andAmami islands, Japan. South Pacific Studies, 36:
103–110.
Hata, H. & H. Motomura, 2016b. First record of[Report] Hata et al.: Paracaesio stonei from Satsunan Islands

Pristipomoides
multidens(Perciformes:
Lutjanidae)
fromTanega-shimaisland,Kagoshima Prefecture, southern Japan. Nature ofKagoshima, 42: 225–229.
Ichikawa, S., S. Sunakawa & T. Matsumoto, 1992. A
general view of fishes of Yaku-shima island. In:
Team for Marine Organism Survey in Inshore ofYaku-shima island (ed.), Report on scientificsurvey of marine organisms from inshore ofYaku-shima island, Pp. 19–46. Team for Marine
Organism Survey in Inshore of Yaku-shimaisland, Yakushima.
Isa, J. & S. Arakaki, 1991. Fisher’s paradise
Okinawa. Shinsei Shuppan Public Company,
Naha.
Kramer, S. H., S. H. Kramer & S. J. Newman, 1994.
New deep-water fish records from the Great
Barrier Reef, Australia. Pacific Science, 48: 70–
79.
Meguro, M. 2013. Paracaesio sordida Abe &Shinohara, 1962. In: H. Motomura, S. Dewa, K.
Furuta & K. Matsuura (eds.), Fishes of Iou-jimaand Take-shima islands, Mishima, Kagoshima,
Japan, P. 152. The Kagoshima University
Museum, Kagoshima and the National Museumof Nature and Science, Tsukuba.
Motomura, H., K. Kuriiwa, E. Katayama, H. Senou,
G. Ogihara, M. Meguro, M. Matsunuma, Y.
Takata, T. Yoshida, M. Yamashita, S. Kimura, H.
Endo, A. Murase, Y. Iwatsuki, Y. Sakurai, S.
Harazaki, K. Hidaka, H. Izumi, & K. Matsuura,
2010. Annotated checklist of marine andestuarine fishes of Yaku-shima Island,
Kagoshima, southern Japan. In: H. Motomura &K. Matsuura (eds.), Fishes of Yaku-shima Island– A World Heritage island in the Osumi Group,
Kagoshima Prefecture, southern Japan, Pp. 65–
247. National Museum of Nature and Science,
Tokyo.
Raj, U. & J. Seeto, 1983. A new species of
Paracaesio (Pisces: Lutjanidae) from the Fiji
Islands. Copeia, 1983 (2): 450–453.
Sakurai, Y., 2014. Paracaesio caerulea (Katayama,1934). In: H. Motomura & K. Matsuura (eds.),
Field guide to fishes of Yoron Island in the middleof the Ryukyu Islands, Japan, P. 237. The
Kagoshima University Museum, Kagoshima andthe National Museum of Nature and Science,
Tsukuba.
Shimada, K., 2013. Lutjanidae. In: T. Nakabo (ed.),Fishes of Japan with Pictorial Keys to the Species.
Third edition. Pp. 913–930, 2001–2004. Tokai
University Press, Hadano. [In Japanese]
White, W. T. & P. R. Last, 2012. Paracaesio
59brevidentata n. sp., a new snapper (Lutjanidae:
Apsilinae) from Indonesia. Zootaxa, 3418: 51–60.
Yoshino, T., 1984. Paracaesio stonei Raj et Seeto.
In: H. Masuda, K. Amaoka, C. Araga, T. Uyeno& T. Yoshino (eds.), The fishes of the JapaneseArchipelago, P. 163, pl. 154-G. Tokai University
Press, Tokyo.
Yoshino, T. & S. Arakaki, 1984. Fishing of Okinawa.
Shinsei Shuppan Public Company, Naha.

薩南諸島から得られたフエダイ科ヤンバルシ
マアオダイ Paracaesio stonei
12

畑晴陵 •岩坪洸樹 •本村浩之

3

1〒890‒0065 鹿児島市郡元 1‒21‒24 鹿児島大学大学院連合農学研究科
2〒892-0847 鹿児島市西千石町 11‒21 鹿児島
MS ビル
3〒890‒0065 鹿児島市郡元 1‒21‒30 鹿児島大
学総合研究博物館要旨 . 大隅諸島屋久島, トカラ列島平島•小宝島近海においてヤンバルシマアオダイ
Paracaesio stonei が 1 個体ずつ, 計 3 個体採集された. これらは鹿児島県における本種の標本に基づく初めての記録であると同時に, 屋久島産標本は本種の分布の北限を更新する記録となる.
投稿日: 2016 年 11 月 13 日
受理日: 2017 年 5 月 18 日
発行日: 2017 年 5 月 31 日

60

[報告] 畑ら: ヤンバルシマアオダイの薩南諸島からの記録

Fauna Ryukyuana, 36: 55–62.

Appendix 1. Counts and measurements of Paracaesio stonei specimens from the Satsunan Islands, Japan.
附録 1. 薩南諸島産ヤンバルシマアオダイ Paracaesio stonei の計数•計測形質.
Yaku-shima

Kodakara-jimaTaira-jima

KAUM–I. 200222

KAUM–I. 54128

KAUM–I. 82741

220.7

379.7

407.8

Dorsal-fin spines 背鰭棘数

10

10

10

Dorsal-fin rays 背鰭軟条数

10

10

10

Anal-fin spines 臀鰭棘数

3

3

3

Standard length (mm; SL)Counts

標準体長

計数形質

Anal-fin rays 臀鰭軟条数

8

8

8

Pectoral-fin rays 胸鰭軟条数

15

16

16

Pelvic-fin spines 腹鰭棘数

1

1

1

Pelvic-fin rays 腹鰭軟条数

5

5

5

Tubed lateral-line scales 有孔側線鱗数

48

49

49

Predorsal scales 背鰭前方鱗数
Scale rows between middle of spinous dorsal-finbase and lateral line
背鰭棘部中央下の側線上方横列鱗数
Scale rows between anal-fin origin andlateral line
側線下方横列鱗数
Circumpeduncular scales 尾柄周囲鱗数

19

20

20

8

9

8

16

17

16

23

23

24

7 + 198 + 17

8 + 17

Head length 頭長

31.9

31.1

33.1

Snout length 吻長

8.4

9.3

11.0

Orbit diameter 眼窩径

9.8

8.1

7.6

Postorbital length 眼後長Distance lower jaw to isthmus
下顎先端から峡部にかけての長さ
Upper-jaw length 上顎長

14.0

14.5

16.1

11.7

11.3

14.4

12.2

12.4

14.1

Maxillary width 上顎幅

3.8

3.8

4.1

Interorbital width 眼隔域幅

11.0

11.0

12.1

Suborbital depth 眼窩骨高

2.4

3.1

4.0

Internarial distance 鼻孔間の長さDistance posterior nostril to orbit
後鼻孔から眼窩前縁までの長さ
Body depth 体高

0.8

0.6

0.9

2.4

2.9

3.1

45.5

44.8

44.5

Body width 体幅

18.1

18.6

17.4

Pre-dorsal-fin length 背鰭前長

38.6

36.8

37.9

Pre-anal-fin length 臀鰭前長

66.5

67.1

68.4

Pre-pelvic-fin length 腹鰭前長

40.7

39.7

41.6

Caudal-peduncle length 尾柄長

17.6

14.0

17.0

Caudal-peduncle depth 尾柄高

12.3

11.7

12.1

Dorsal-fin base length 背鰭基底長

53.1

50.1

49.3

First dorsal-fin spine length 背鰭第 1 棘長

7.7

8.4

brokenSecond dorsal-fin spine length 背鰭第 2 棘長

14.0

12.2

brokenThird dorsal-fin spine length 背鰭第 3 棘長

16.4

16.1

brokenLongest dorsal-fin spine length 最長背鰭棘長

17.0

15.1

15.2

Last dorsal-fin spine length 最後背鰭棘長

12.7

10.9

11.6

Gill rakers 鰓耙数Measurements 計測形質
As % SL 体各部の体長に対する割合[Report] Hata et al.: Paracaesio stonei from Satsunan Islands

61

Appendix 1. (Continiued).
附録 1. (続き).

62

Yaku-shima

Kodakara-jimaTaira-jima

KAUM–I. 200222

KAUM–I. 54128

KAUM–I. 82741

Longest dorsal-fin soft ray length 最長背鰭軟条長
Penultimate dorsal-fin soft ray length
後方から 2 番目の背鰭軟条長
Last dorsal-fin soft ray length 最後背鰭軟条長

16.2

15.1

14.6

14.7

13.5

13.7

11.8

10.9

11.1

Anal-fin base length 臀鰭基底長

21.5

19.3

20.1

First anal-fin spine length 臀鰭第 1 棘長

6.3

5.2

5.4

Second anal-fin spine length 臀鰭第 2 棘長

9.4

8.6

8.6

Third anal-fin spine length 臀鰭第 3 棘長Penultimate anal-fin soft ray length
後方から 2 番目の臀鰭軟条長
Ultimate anal-fin soft ray length 最後臀鰭軟条長

12.2

11.0

10.5

13.7

12.9

12.6

11.6

11.3

10.9

Upper caudal-fin lobe length 尾鰭上葉長

36.7

34.2

34.3

Lower caudal-fin lobe length 尾鰭下葉長

34.6

30.7

32.1

Caudal fin concavity length 尾鰭湾入長

5.1

11.0

10.9

Pectoral-fin length

40.0

36.5

36.7

胸鰭長

[報告] 畑ら: ヤンバルシマアオダイの薩南諸島からの記録