Bull. Tohoku Univ. Museum, No. 15, pp. 9–106, 2016
© by The Tohoku University Museum

A catalogue of name-bearing type specimens of fossil
Bivalvia (Mollusca) registered in the Tohoku University
MuseumShin’ichi Sato1)*, Tomoki Chiba2), Takaki Yamanaka2), Jun Nemoto2), Masanori Shimamoto3) and
Takashi Matsubara4)
1)

The Tohoku University Museum, Aoba 6-3, Aramaki, Aoba-ku, Sendai 980-8578, Japan      *Present address: Institute of Geosciences, Shizuoka University, Oya 836, Shizuoka 422-8529, Japan <sato.shinichi.c@shizuoka.ac.jp>
2)
Institute of Geology and Paleontology, Graduate School of Science, Tohoku University, Aoba 6–3, Aramaki, Aoba-ku, Sendai
980-8578, Japan
3)
Techno Hase Co., Ltd., Hasekura 2-10, Aoba-ku, Sendai 980-0824, Japan4)
Laboratory of Earth Sciences, Faculty of Education, Hokkaido University of Education at Kushiro, Shiroyama 1-15-55, Kushiro
085-8580, Japan
Abstract: Since 1911, more than 110,000 fossil specimens have been studied and registered in theInstitute of Geology and Paleontology, Tohoku University, Sendai (IGPS). This catalogue includes
452 name-bearing type specimens (i.e. holotype, lectotype, syntype and neotype) of fossil Bivalvia,
registered in the Tohoku University Museum. Each taxon data includes the following items: 1)
species name with its author name(s) and date of publication, 2) bibliographic data of the originaldescription, 3) kind of type specimen(s) and present condition of specimen(s), 4) type locality, 5)
stratigraphic unit, 6) geologic age. This database and color pictures of the existing type specimens
are searchable on the website of the Tohoku University Museum (http://webdb2.museum.tohoku.
ac.jp/t_bivalve/). The lectotype of Thyasira tokunagai Kuroda and Habe, 1951 is designated herein.
Key words: database, fossil bivalves, name-bearing type specimens, the Tohoku University Museum

Introduction
Tohoku University was established in 1907 as the third
Imperial University of Japan, and the Institute of Geology
and Paleontology (abbreviated as IGPS) was founded
in 1911 with Hisakatsu Yabe as the first professor. Since
then, the institute has produced a number of distinguishedmolluscan paleontologists, such as Ichiro Hayasaka,
Takumi Nagao, Sitihei (or Shichihei) Nomura, Saburo
Shimizu, Kotora Hatai, Tamio Kotaka, Yasuhiko Kamada,Kôichirô Masuda, Masafumi Murata, Hiroshi Noda, Kenshiro
Ogasawara, and Yutaka Honda, and until today, a lot of newmolluscan taxa have been described by the professors andstudents of the institute.
The type and referred molluscan fossil specimens had
been deposited in the institute with the IGPS registration
numbers. In 1998, more than 110,000 paleontological
specimens deposited in the institute were moved to the
Tohoku University Museum, which was newly established inthis year. The details of each IGPS molluscan type specimen

can be seen in the past checklists and monographs such as
Hatai and Nisiyama (1952), Oyama et al. (1960), Masuda
and Noda (1976) and Ogasawara (2001, 2002) for Cenozoic;
Hayami (1975), Tanaka and Toshimitsu (2003), Toshimitsu
and Tanaka (2003) for Mesozoic; and Nakazawa andNewell (1968), Hayami and Kase (1977) and Kase (2004)
for Paleozoic; in addition to Hanzawa et al. (eds., 1961).
However, no institutional or museum catalogue of the IGPS
molluscan collection has been published, and the present
condition of the specimens has not been clear. Moreover,
we can not find the precise morphological characters of thetype specimens from some original descriptions, because of
the indistinct pictures. Therefore, the thoroughly curatorial
survey of the IGPS paleontological collection is needed.
As a first step of the making the complete catalogue ofthe IGPS paleontological collection, we herein present
a catalogue of fossil Bivalvia, which occupies the large
amount of the collection. The present catalogue includes all
information about the name-bearing type specimens of fossil
Bivalvia in the IGPS collection with photographs.

10

Shin’ichi Sato, Tomoki Chiba, Takaki Yamanaka, Jun Nemoto, Masanori Shimamoto and Takashi Matsubara

Methods
On the basis of Ogasawara (2001, 2002), Matsuoka andUgai (2001), Toshimitsu and Tanaka (2003), Tanaka andToshimitsu (2003) and Kase (2004), we compiled a basic
checklist of name-bearing type specimens of fossil Bivalviapreserved in the Tohoku University Museum. Several
species-group names missing in these previous catalogues
were added with their name-bearing type specimen data.
All data were rechecked by the bibliographic survey ofthe original descriptions. To confirm the present condition(existing or missing) of the name-bearing type specimens,
we compared the candidate type specimens and theirregistration numbers with figures and registration numbers
shown in the original description. We also checked forconsistency of taxon name, locality, lithostratigraphic unit
(or formation name), geologic age between label attachedto the specimens and original description. Recent Bivalvia
was excluded from the present catalogue. The photographsof all existing name-bearing type specimens were taken to
make database, and they were illustrated in the plates of thepresent catalogue.

Format
The name-bearing type specimens were arrangedalphabetically in the genus- and species-group names
to make easy for reference. Each taxonomic entry in thepresent catalogue is composed of the following data as given
in descending order: 1) original spelling of the taxon name,together with author name(s) and publication date, 2) plate
number and figure number(s) in the plates of the present
catalogue, 3) bibliographic data of the original description,
including abbreviation of the publication, volume, page,
plate number(s) and figure number(s), 4) category of thename-bearing type specimen (holotype, lectotype, syntype
or neotype), together with IGPS (Institute of Geology
and Paleontology, Faculty of Science, Tohoku University,
Sendai) registration number and present condition (existingor missing), 5) type locality given as names of prefecture,city, town, village and more details both in the way they
appeared in the original description and currently known,
6) stratigraphic unit (formation name) from which the typespecimen was obtained, 7) geologic age, 8) taxonomic
remarks including synonymous status of a given taxon andcorrection for original spelling and registration number.
The name-bearing type specimen shifted from DGS(abbreviation of the Department of Geology, Faculty of
Education, Tohoku University, Sendai; abolished in 1994) to
IGPS, the previous DGS registration number was indicatedin round brackets behind the IGPS registration number. The
literature in which the lectotype or neotype was designated
was also added in round brackets. We indicated current

names of city, town and village in square brackets, as wellas correction of them from the original description. The
updated information on stratigraphic unit and geologic age
were also added in square brackets.
The photographs of existing name-bearing typespecimens were illustrated in the plates at the end of thepresent catalogue.
The database and color pictures of the existing typespecimens are also searchable on the website of the Tohoku
University Museum (http://webdb2.museum.tohoku.ac.jp/t_bivalve/).

Abbreviation of publications
Bull. Biogeogr. Soc. Japan: Bulletin of the BiogeographicalSociety of Japan.
Bull. Tohoku Univ. Mus.: Bulletin of the Tohoku University
Museum.
Geol. Palaeont. Southeast Asia: Geology and Palaeontologyof Southeast Asia.
Illust. Cat. Jpn. Shells: Illustrated Catalogue of Japanese
Shells.
Japan. Jour. Geol. Geogr.: Japanese Journal of Geology
and Geography.
Jour. Fac. Sci. Hokkaido Imp. Univ., Ser. 4: Journal of theFaculty of Science, Hokkaido Imperial University. Series
4, Geology and Mineralogy.
Jour. Fac. Sci., Imp. Univ. Tokyo, Sec. 2: Journal of theFaculty of Science, Imperial University of Tokyo, Section
2, Geology, Mineralogy, Geography, Seismology.
Jour. Geol. Soc. Japan: Journal of the Geological Society ofJapan.
Jour. Paleont.: Journal of Paleontology.
Jub. Publ. Commem. Prof. H. Yabe 60th Birthday: Jubilee
Publication in the Commemoration of Professor H.
Yabe, M.I.A. Sixtieth Birthday.
Mem. Coll. Sci. Univ. Kyoto, Ser. B: Memoirs of the Collegeof Science, Kyoto University, Series B.
Miner. Geol.: Kôbutu-to-Tisitu (Mineral and Geology).
Paleontol. Res.: Paleontological Research.
Palaeont. Soc. Japan, Spec. Pap.: Palaeontological Societyof Japan, Special Papers.
Proc. Imp. Acad.: Proceedings of the Imperial Academy.
Prof. S. Kanno Mem. Vol.: Professor Saburo Kanno
Memorial Volume.
Rep., Geol. Surv. Japan: Report, Geological Survey ofJapan.
Res. Rep. Kochi Univ., Nat. Sci.: Research Reports of the
Kochi University, Natural Science.
Saito Ho-on Kai Mus. Nat. Hist. Res. Bull.: Saito Ho-on Kai
Museum of Natural History Research Bulletin.
Saito Ho-on Kai Mus. Res. Bull.: Saito Ho-on Kai Museum
Research Bulletin.

11

Catalogue of type specimens of fossil Bivalvia

400

Missing
Existing

350
300
250
200
150
100

NeotypeSyntype0

Lectotype50
Holotype

1XPEHURIVSHFLHVíJURXSQDPHV

As a result of the present survey, it became obvious that452 name-bearing type specimens of fossil Bivalvia are

Fig. 1. Total number of species-group names in eachcategory of the name-bearing type specimens of fossil
Bivalvia deposited in the Tohoku University Museum andtheir present conditions.

Details on geologic ages are as follows (Fig. 2):
Cenozoic:366
Quaternary:
71
Holocene:
3
Pleistocene:
68
Neogene:
191
Pliocene:
35
Miocene:
156
Paleogene:
104
Oligocene:
58
Eocene:46
Mesozoic:
60
Cretaceous:
52
Jurassic:
1
Triassic:
7
Paleozoic:
23
Permian:
20
Carboniferous:
1
Devonian:
0
Silurian:
0Ordovician:	
2
Unknown:
3.

160
140

Missing
Existing

120
100
80
60
40
20
0

Holocene
Pleistocene
Pliocene
MioceneOligocene
EoceneCretaceous
Jurassic
Triassic
Permian
Carboniferous
Devonian
SilurianOrdovician
Unknown

Summary

registered in the Tohoku University Museum. Among them,
412 specimens are still existing.
Details of the type of the name-bearing types are as
follows (Fig. 1):
Holotypes:
381 specimens
Lectotypes:	   36 specimens
Syntypes:
34 specimens
Neotype:1 specimen.

1umberofsSeciesígrouSnames

Saito Ho-on Kai Spec. Pub., no. 2 (Prof. T. Kotaka
Commem. Vol.): Saito Ho-on Kai Special Publication,
no. 2 (Professor Tamio Kotaka Commemorative Volume
on Molluscan Paleontology).
Sci. Rep., Fac. Arts and Liter., Nagasaki Univ.: Science
Reports of the Faculty of Arts and Literature, Nagasaki
University.
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.): Science
Reports of the Tôhoku Imperial University, Second
Series (Geology).
Sci. Rep. Tohoku Univ., 2nd Ser. (Geol.): Science Reports of
the Tohoku University, Second Series (Geology).
Sci. Rep. Tôhoku. Univ., 2nd Ser. (Geol.), Spec. Vol.:
Science Reports of the Tôhoku University, Second
Series (Geology), Special Volume.
Sci. Rep. Tohoku Univ., 2nd Ser. (Geol.), Spec. Vol.: Science
Reports of the Tohoku University, Second Series(Geology), Special Volume.
Short Pap. Inst. Geol. Paleont., Tôhoku Univ.: Short Papers
from the Institute of Geology and Paleontology, Tôhoku
University, Sendai.
Trans. Proc. Palaeont. Soc. Japan, N.S.: Transactions andProceedings of the Palaeontological Society of Japan,
New Series.
U.S. Geol. Surv., Prof. Pap.: United States Geological
Survey Professional Paper.
Venus: Venus (Japanese Journal of Malacology).

Fig. 2. Total number of species-group names in eachgeologic age, from which the name-bearing type specimensof fossil Bivalvia deposited in the Tohoku University Museum
were obtained.

12

Shin’ichi Sato, Tomoki Chiba, Takaki Yamanaka, Jun Nemoto, Masanori Shimamoto and Takashi Matsubara

Fig. 3. Total number of species-group names in eachchronological period, in which new species-group namesof fossil Bivalvia were proposed and their name-bearing
type specimens were deposited in the Tohoku University
Museum.

40
30
20
0

Others10
T. Kotakaí

DQGODWHUí

í

í

í

í

í

í

í

50

J. Akutsu

60

I. Hayasaka70Y. Kamada

80

H.Yabe

0LVVLQJ([LVWLQJ

90

K. Hatai100

H. Noda0LVVLQJ([LVWLQJ

S. Nomuraí

1XPEHURIVSHFLHVíJURXSQDPHV

These results suggest that 1) 91.2% of the name-bearing
type specimens of fossil Bivalvia of the IGPS collection are
still existing in the Tohoku University Museum, 2) 81.0% ofthem were obtained from Cenozoic, and 34.5% were from
Miocene, and 3) 25.4% and 28.3% of them were proposed
in 1921­–1930 mainly by I. Hayasaka, T. Nagao and H. Yabe
and in 1961–1970 by K. Masuda, M. Murata and H. Noda,
respectively.
A total of 33 researchers proposed new species-groupnames of fossil Bivalvia as the first author, and depositedthe name-bearing type specimens in IGPS. Of them, T.
Nagao described 76 new species-group names as thefirst author during 1928 and 1943, and S. Nomura and K.

T. Nagao

6
115
87
23
42
128
25
15
9
1
 1.

K. Masuda

1911–1920:	
1921–1930:	
1931–1940: 
1941–1950:
1951–1960: 
1961–1970:	
1971–1980: 
1981–1990: 
1991–2000: 
2000–2010:	
2011 and later:

Masuda described 63 species-group names in 1932–1939
and 1952–1994, respectively (Fig. 4). H. Yabe and H. Noda
described 40 new species-group names in 1926–1942 and
1962–1971 respectively, and K. Hatai also described 32
species-group names in 1940–1974.
Among the name-bearing type specimens of fossil
Bivalvia in the IGPS collection, 26 specimens were thoseof members of the genus Anadara, followed by those of
the genus Chlamys (22 species-group names) and Pecten(20 species-group names) (Fig. 5). Nineteen new speciesgroup names of Anadara were described by H. Noda, and
20 of Chlamys were described by K. Masuda. In the genus
Glycymeris, Patinopecten and Venericardia, 13 new speciesgroup names and their name-bearing type specimens werealso proposed and registered in IGPS.
According to Ogasawara (2004), 68 species-group names
described under the genus Chlamys and 54 species-groupnames of Anadara were described from Japanese Islands
and its adjacent areas during the 20th Century. Among
their name-bearing type specimens, 32.3% of Chlamys and48.1% of Anadara were deposited in the Tohoku University
Museum.
These results suggest that many researchers of Tohoku
University especially proposed new species-group namesof Anadara, Chlamys and Pecten for their studies on thetaxonomy, biostratigraphy, paleobiogeography, evolutionary
paleobiology and paleoenvironmental analyses. In particular,a number of new species-group names were proposed
during the two "golden ages", namely 1921–1940 and 1961–
1970 (Fig. 3), in this university.

1XPEHURIVSHFLHVíJURXSQDPHV

The chronologic succession of the number of the bivalve
studies, in which new species-group names were proposed
and the name-bearing types were reposited at IGPS, are as
follows (Fig. 3):

Fig. 4. Total number of species-group names in each author,
who proposed new species-group names of fossil Bivalvia
as the first author and deposited their name-bearing typespecimen in the Tohoku University Museum.

0LssLQJ
ExLstLQJ

25

400
350
300

20

250

15

200
150

10

100
5

OthersNuculaArca

Cardium

Tellina

Venericardia

Patinopecten

PectenGlycymeris

50
Anadara0

0

1XPEer Rf specLesíJrRXp QDPes(Others cDteJRry)

30

Chlamys

1XPEerRIsSeFLesíJrRXSQDPes
(Except fRr Others cDteJRry)

Catalogue of type specimens of fossil Bivalvia

Fig. 5. Total number of species-group names in each genusof the name-bearing type specimens of fossil Bivalvia
deposited in the Tohoku University Museum.

Catalogue
Acanthopecten onukii Murata, 1964
Plate 1, Figure 1
Trans. Proc. Palaeont. Soc. Japan, N.S., no. 54, p. 222,
pl. 35, fig. 1a–­b. Holotype: IGPS no. 85745 (Existing).
Type locality: IGPS loc. no. Mi 117 (Murata, 1964), Shigejizawa, Kamiyasse, Kesennuma City, Miyagi Prefecture.
Stratigraphic unit: Lower part of Kanokura Formation.
Geologic age: Middle Permian.
Acanthopecten spinosus Hayasaka, 1925b
Plate 1, Figure 2
S ci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 8,
no. 2, p. 10, pl. 8, fig. 6. Lectotype: IGPS no. 22388(designated by Murata, 1964; Existing). Type locality:
Imô, Yahagi-mura, Kesen-gôri, prov. Rikuzen [currently
Imo, Yahagicho, Rikuzentakata City, Iwate Prefecture].
Stratigraphic unit: Lower part of Kanokura Formation.
Geologic age: Lower Permian [Middle Permian].
Acila divaricata chitosensis Noda, 1962b
Plate 1, Figure 3
Sci. Rep. Tohoku Univ., 2nd Ser. (Geol.), vol. 34, no. 3, p.
227, pl. 16, fig. 9. Holotype: IGPS no. 79060 (Existing).
Type locality: Loc. No. 344 (Noda, 1962b), cliff of the
Shibumi River, just under the Chitose Bridge, Chitose,
Matsudai-machi, Higashikubiki-gun [currently Tokamachi
City], Niigata Prefecture. Stratigraphic unit: Higashigawa
Formation. Geologic age: “Middle” Pliocene.

13

Acila (Acila) kiiensis Masuda and Katto, in Katto and
Masuda, 1978
Plate 1, Figures 4a–b
Res. Rep. Kochi Univ., Nat. Sci., vol. 27, p. 104, pl. 3,
figs. 1a–3b. Holotype: IGPS no. 96071 (Existing). Type
locality: Loc. no. 2 (Katto and Masuda, 1978), Tanozaki,
about 1 km SWS of Tanami, Kushimoto-cho, Nishimurogun [currently Higashimuro-gun], Wakayama Prefecture.
Stratigraphic unit: Tanami Formation. Geologic age:
Oligocene [Early Miocene].
Aequipecten matsunagiensis Masuda, 1966b
Plate 1, Figure 5
Trans. Proc. Palaeont. Soc. Japan, N.S., no. 64, p. 323,
pl. 35, figs. 4–6. Holotype: IGPS no. 90089 (formerly
DGS no. 4506; Existing). Type locality: No. 32 (Masuda,
1966a), small road side exposure, about 200 m S of the
Matsunagi Primary School, [Matsunagi-machi], Suzu
City, Ishikawa Prefecture. Stratigraphic unit: Higashi-innai
Formation. Geologic age: Miocene [latest Early Miocene].
Amussiopecten akiyamae Masuda, 1962aPlate 1, Figure 6
Sci. Rep. Tohoku Univ., 2nd Ser. (Geol.), vol. 33, no. 33, p.
224, pl. 27, figs. 1–3. Holotype: IGPS no. 90658 (formerly
DGS no. 3854; Existing). Type locality: Sea cliff at Fudôiwa,Kanaya, Amaha-machi, Kimitsu-gun [currently Futtsu City],
Chiba Prefecture. Stratigraphic unit: Inagozawa Formation[Nokogiriyama Formation]. Geologic age: Miocene [latest
Late Miocene–earliest Early Pliocene].
Anadara (Scapharca) akitaensis Noda, 1966aPlate 1, Figures 7a–d
Sci. Rep. Tohoku Univ., 2nd Ser. (Geol.), vol. 38, no. 1, p.
106, pl. 5, figs. 9, 10, 19, 20. Holotype: IGPS no. 16330(Existing). Type locality: Tayazawa, Wakimoto-mura,
Minamiakita-gun [sic, Wakimoto, Oga City], Akita Prefecture.
Stratigraphic unit: Sasaoka Formation. Geologic age:
Pliocene [latest Late Pliocene–early Early Pleistocene].
Anadara (Anadara) amicula elongata Noda, 1966aPlate 1, Figures 8a–b
Sci. Rep. Tohoku Univ., 2nd Ser. (Geol.), vol. 38, no. 1, p.
84, pl. 5, figs. 2–7. Holotype: IGPS no. 85907 (originallystated IGPS no. 62435; Existing). Type locality: Upstream of
Kanakusare [sic, Kanakusari] River, Makimachi, Kanazawa
City, Ishikawa Prefectrure. Stratigraphic unit: Omma
Formation. Geologic age: Pliocene [Early Pleistocene].
Anadara (Anadara) arasawaensis Noda, 1966aPlate 1, Figures 9a–b
Sci. Rep. Tohoku Univ., 2nd Ser. (Geol.), vol. 38, no. 1,

14Shin’ichi Sato, Tomoki Chiba, Takaki Yamanaka, Jun Nemoto, Masanori Shimamoto and Takashi Matsubara

p. 86, pl. 4, figs. 13, 15–17. Holotype: IGPS no. 90046(Existing). Type locality: Arasawa, Gomyojin-mura [sic,Omyojin], Shizukuishi-machi [sic, Shizukuishi-cho], [Iwategun], Iwate Prefecture. Stratigraphic unit: Sakamotogawa
Formation. Geologic age: Miocene.
Anadara (Anadara) gentaroensis Noda, 1966aPlate 1, Figure 10
Sci. Rep. Tohoku Univ., 2nd Ser. (Geol.), vol. 38, no. 1, p.
87, pl. 7, figs. 19–21. Holotype: IGPS no. 86411 (Existing).
Type locality: Gentaro, Kozai-mura [sic, Kosai], Ouchimachi [sic, Marumori-machi], Igu-gun, Miyagi Prefecture.
Stratigraphic unit: Yoshizawa Formation [sic, Yoshigasawa
Formation]. Geologic age: Miocene [latest Early–earliest
Middle Miocene].
Anadara (Anadara) hataii Noda, 1966aPlate 1, Figures 11a–c; Plate 2, Figures 1a–b
Sci. Rep. Tohoku Univ., 2nd Ser. (Geol.), vol. 38, no. 1, p.
88, pl. 6, figs. 1, 4–7. Holotype: IGPS no. 73210 (Existing).
Type locality: Nishigoto [sic, Nishigodo], Hanawamachi, Higashishirakawa-gun, Fukushima Prefecture.
Stratigraphic unit: Tanagura Formation [=Kubota
Formation]. Geologic age: Miocene [early Late Miocene].
Anadara (Anadara) hokkaidoensis Noda, 1966aPlate 2, Figures 2a–c
S ci. Rep. Tohoku Univ., 2nd Ser. (Geol.), vol. 38, no.
1, p. 90, pl. 4, figs. 18, 19, pl. 5, fig. 14, pl. 8, figs. 1,
8. Holotype: IGPS no. 86408 (Existing). Type locality:
Upstream of the Ishii-sawa [sic, Ishiizawa], Atsunai,
Horonobe-machi [sic, Urahoro-cho], Tokachi-gun, Tokachi
Province, Hokkaido. Stratigraphic unit: Chokubetsu
Formation [=Ishiizawa Formation]. Geologic age: Miocene[early Middle Miocene].
Anadara (Scapharca?) iwashibaraensis Noda, 1965
Plate 2, Figure 3
Trans. Proc. Palaeont. Soc. Japan, N.S., no. 59, p. 104,
pl. 10, fig. 15. Holotype: IGPS no. 29050 (Existing).
Type locality: Iwashibara, Kamitareki-mura, Ogasa-gun[sic, Yashibara, Shimotaruki, Kakegawa City], Shizuoka
Prefecture. Stratigraphic unit: Dainichi Formation.
Geologic age: Pliocene [Early Pleistocene].
Anadara (Anadara) iwatensis Noda, 1966aPlate 2, Figures 4a–b
Sci. Rep. Tohoku Univ., 2nd Ser. (Geol.), vol. 38, no. 1, p.
91, pl. 4, figs. 21, 22. Holotype: IGPS no. 90048 (Existing).
Type locality: Arasawa, Myojin-mura, Shizukuishimachi [sic, Omyojin, Shizukuishi-cho], Iwate-gun, Iwate
Prefecture. Stratigraphic unit: Sakamotogawa Formation.

Geologic age: Miocene.
Anadara (Anadara) iwatonoensis Noda, 1966aPlate 2, Figures 5a–b
Sci. Rep. Tohoku Univ., 2nd Ser. (Geol.), vol. 38, no. 1, p.
92, pl. 9, fig. 21, pl. 11, fig. 19. Holotype: IGPS no. 63342(Existing). Type locality: Jakotsuzawa, Saruhashi, KitaTsuru-gun [sic, Saruhashimachi, Otsuki City], Yamanashi
Prefecture. Stratigraphic unit: Iwatono Formation.
Geologic age: Miocene.
Anadara (Anadara) kakehataensis Hatai and Nisiyama,
1949
Plate 2, Figures 6a–d
Jour. Paleont., vol. 23, no. 1, p. 88, pl. 23, figs. 8–10.
Holotype: IGPS no. 72510 (Existing). Although Ogasawara(2001) stated IGPS no. 72511 as the holotype, it is theholotype of Anadara (Anadara) kurosedaniensis Hatai
and Nisiyama, 1949. Type locality: West cliff, about 50
m S of the bridge at Kakehata, Unohana-mura, Nei-gun[currently Yatsuomachi, Toyama City], Toyama Prefecture.
Stratigraphic unit: Susahara Formation [=Kurosedani
Formation]. Geologic age: Miocene [latest Early Miocene].
Remarks: The present species is a type species of
Hataiarca Noda, 1966a (by original designation).
Anadara (Hataiarca) kogachiensis Noda, 1971
Plate 2, Figures 7a–c; Plate 3, Figures 1a–b
Trans. Proc. Palaeont. Soc. Japan, N.S., no. 81, p. 37, pl.
6, figs. 1–5, 8–17. Holotype: IGPS no. 86757 (Existing).
Type locality: Loc. no. 109 (Noda, 1971), west of Kogachi,
Haneji-son, [Kunigami-gun] [currently Kogachi, Nago
City], Okinawa-jima, Okinawa Prefecture. Stratigraphic
unit: Kogachi Member of Haneji Formation. Geologic age:
Pliocene [Pleistocene].
Anadara (Anadara) kurosedaniensis Hatai and Nisiyama,
1949
Plate 3, Figures 2a–b
Jour. Paleont., vol. 23, no. 1, p. 89, pl. 23, figs. 11, 12.
Holotype: IGPS no. 72511 (Existing). Type locality:
West cliff, about 50 m south of the bridge at Kakehata,
Unohana-mura, Nei-gun [currently Yatsuomachi, Toyama
City], Toyama Prefecture. Stratigraphic unit: Susahara
Formation [=Kurosedani Formation]. Geologic age:
Miocene [latest Early Miocene].
Anadara makiyamai Hatai and Nisiyama, 1939 [“1938”]
Japan. Jour. Geol. Geogr., vol. 16, nos. 1–2, p. 143–144,
pl. 9, fig. 7. Holotype: IGPS no. 62430 (Missing). Type
locality: Nanseki, Meisen-gun, Kankyô-dô, North Tyôsen[currently Myongchon-gun, Hamgyongpuk-to, North

Catalogue of type specimens of fossil Bivalvia
Korea]. Stratigraphic unit: Heiroku Beds of Meisen Series[Heiroku Formation of Meisen Group]. Geologic age:
Miocene [latest Early–earliest Middle Miocene]. Remarks:
This species is a junior synonym of Anadara abdita(Makiyama, 1926).
Anadara (Scapharca) masudai Noda, 1966aPlate 3, Figures 3a–b
Sci. Rep. Tohoku Univ., 2nd Ser. (Geol.), vol. 38, no. 1, p.
118, pl. 5, figs. 16–17, pl. 13, figs. 2, 4. Holotype: IGPS
no. 51550 (Existing). Type locality: Subsurface about 4–5
m below the Star Hotel, Honmoku, [Naka-ku], Yokohama
City, Kanagawa Prefecture. Stratigraphic Unit: Tokyo
Formation. Geologic age: Pleistocene [Late Pleistocene].
Anadara (Anadara) naganoensis Noda, 1966aPlate 3, Figure 4
Sci. Rep. Tohoku Univ., 2nd Ser. (Geol.), vol. 38, no. 1, p.
95, pl. 3, fig. 8. Holotype: IGPS no. 86410 (Existing). Type
locality: Stream cliff of a tributary of the Saikawa, Fukudo,
about 10 m E of the Fukudo Bridge, Shinanoshin-machi[sic, Shinshushin-machi], Kamiminochi-gun [currently
Shinshushinmachi, Nagano City], Nagano Prefecture.
Stratigraphic unit: Gonda Formation. Geologic age: Late
Miocene.
Anadara (Tegillarca) obessa Kotaka, 1953
Plate 3, Figures 5a–b
Trans. Proc. Palaeont. Soc. Japan, N.S., no. 10, p. 35, pl.
4, figs. 1a–6c. Holotype: IGPS no. 66536 (Existing). IGPS
no. 66356 given by original description as the holotype
is incorrect. Type locality: Coast near Nago-machi [sic,Nago-cho], Kunigami-gun, [Government of the Ryukyu
Islands] [currently Nago City, Okinawa Prefecture].
Stratigraphic unit: Raised beach deposits. Geologic age:
[Late Pleistocene or Holocene].
Anadara (Scapharca) omaruensis Sasaki, 1991
Plate 3, Figures 6a–b
Trans. Proc. Palaeont. Soc. Japan, N.S., no. 161, p. 710,
figs. 11.2a–b, 12.5a–12.8b. Holotype: IGPS no. 101325(Existing). Type locality: Iwasebashi, Matsubara, [Heida],
Kawaminami-machi [sic, Kawaminami-cho], Koyu-gun,Miyazaki Prefecture. Stratigraphic unit: Koyu Formation.
Geologic age: Pliocene [late Late Pliocene–early Early
Pleistocene].
Anadara (Hataiarca) pseudosubcrenata Ogasawara, 1977
Plate 3, Figures 7a–b
Sci. Rep. Tohoku Univ., 2nd Ser. (Geol.), vol. 47, no. 2, p. 92,
pl. 6, figs. 1–2b, 5a–6. Holotype: IGPS no. 95070 (Existing).
Type locality: Loc. no. KO-01 (Ogasawara, 1977), river floor

15

of Sai-gawa, 350 m SE of Okuwa Bridge, Okuwa, Kanazawa
City, Ishikawa Prefecture. Stratigraphic unit: Omma
Formation. Geologic age: Pliocene [Early Pleistocene].
Anadara (Scapharca?) shizuokaensis Noda, 1965
Plate 3, Figures 8a–b
Trans. Proc. Palaeont. Soc. Japan, N.S., no. 59, p. 103, pl.
11, figs. 7, 8. Holotype: IGPS no. 78919 (Existing). Type
locality: Hosoya, Kitaogasa-mura, Ogasa-gun [currently
Kakegawa City], Shizuoka Prefecture. Stratigraphic unit:
Nango Sandstone and Mudstone Alternation. Geologic
age: Pliocene.
Anadara (Scapharca) taiwanica Noda, 1966aPlate 3, Figure 9; Plate 4, Figures 1a–c
Sci. Rep. Tohoku Univ., 2nd Ser. (Geol.), vol. 38, no. 1,
p. 112, pl. 10, figs. 1, 4–6. Holotype: IGPS no. 42357(Existing). Type locality: About 1,450 m W of Hoko
[sic, Hokkô, Ôbokô], Byoritsu-gun, Shinchiku-shu[currently Yamukeng, Jinshih Village, Hsihu Township,
Miaoli County], Taiwan. Lithostratigraphic unit: Tokazan
Formation [=Toukoushan Formation]. Geologic age:
Pliocene. Remarks: Anadara taiwanica Hu, 1992, from theraised coral reefs in Taiwan, is a junior homonym (Huang
and Masuda, 2001).
Anadara (Scapharca) takanabensis Sasaki, 1991
Plate 4, Figures 2a–b
Trans. Proc. Palaeont. Soc. Japan, N.S., no. 161, p. 710,
figs. 11.4a–b, 12.1a–12.4b. Holotype: IGPS no. 101327(Existing). Type locality: Toriyama, [Heida], Kawaminamimachi [sic, Kawaminami-cho], Koyu-gun, Miyazaki
Prefecture. Stratigraphic unit: Koyu Formation. Geologic
age: Pliocene [late Late Pliocene–early Early Pleistocene].
Anadara (Hataiarca) takayamai Noda, 1966aPlate 4, Figures 3a–b
S ci. Rep. Tohoku Univ., 2nd Ser. (Geol.), vol. 38, no.
1, p. 120, pl. 2, figs. 18, 22, pl. 13, figs. 5, 6, 17, 18.
Holotype: IGPS no. 86403 (Existing). Type locality:
Kubusu River cliff, Kakehata, Yatsuo-machi, Nei-gun[currently Yatsuomachi, Toyama City], Toyama Prefecture.
Stratigraphic unit: Kurosedani Formation. Geologic age:
Miocene [latest Early Miocene].
Anadara (Anadara) tanakuraensis Noda, 1966aPlate 4, Figures 4a–b
Sci. Rep. Tohoku Univ., 2nd Ser. (Geol.), vol. 38, no. 1, p.
100, pl. 8, figs. 11, 12. Holotype: IGPS no. 28404 (Existing).
Type locality: Kamitoyonosawa, [Nagare], Tanaguramachi, Higashishirakawa-gun, Fukushima Prefecture.
Stratigraphic unit: Tanagura Formation [=Kubota

16

Shin’ichi Sato, Tomoki Chiba, Takaki Yamanaka, Jun Nemoto, Masanori Shimamoto and Takashi Matsubara

Formation]. Geologic age: Miocene [early Late Miocene].
Anadara (Tosarca) tosaensis Noda, 1965
Plate 4, Figure 5
Trans. Proc. Palaeont. Soc. Japan, N.S., no. 59, p. 105, pl.
11, figs. 11–13. Holotype: IGPS no. 54609 (Existing). Type
locality: Oono, Yasuda-mura [sic, Yasuda-machi], Aki-gun,
Kochi Prefecture. Stratigraphic unit: Ananai Formation.
Geologic age: Pliocene [late Late Pliocene–early Early
Pleistocene]. Remarks: The present species is a typespecies of Tosarca Noda, 1965 (by original designation).
Anadara (Anadara) tsudai Noda, 1966aPlate 4, Figures 6a–c
S ci. Rep. Tohoku Univ., 2nd Ser. (Geol.), vol. 38, no.
1, p.103, pl. 2, figs. 19–21. Holotype: IGPS no. 86396(Existing). Type locality: Hirabayashi, Yatsuo-machi,Nei-gun [currently Yatsuomachi, Toyama City], Toyama
Prefecture. Stratigraphic unit: Joyama Formation.
[=Higashibessho Formation]. Geologic age: Miocene[latest Early–early Middle Miocene].
Anadara (Hataiarca) yatsuoensis Noda, 1966aPlate 4, Figures 7a–c
Sci. Rep. Tohoku Univ., 2nd Ser. (Geol.), vol. 38, no. 1,
p. 122, pl. 13, figs. 1, 3, 9, 16, 19–20. Holotype: IGPS
no. 86402 (Existing). Type locality: Kubusu River cliff,
Kakehata, Yatsuo-machi, Nei-gun [currently Yatsuomachi,
Toyama City], Toyama Prefecture. Stratigraphic unit:
Kurosedani Formation. Geologic age: Miocene [latest
Early Miocene].
Angulararca yubaensis Noda, 1966b
Saito Ho-on Kai Mus. Res. Bull., no. 35, p. 16, figs. 7, 8,
17, 18. Holotype: IGPS no. 72884 (Missing). Type locality:
Road side cliff, west of Yuba, Rifu-mura [currently Sugaya,
Rifu-cho], Miyagi-gun, Miyagi Prefecture. Stratigraphic
unit: Aoso Formation. Geologic age: Miocene [LateMiocene]. This species is the type species of Angulararca
Noda, 1966b.
Angulus maximus submaximus Mizuno, 1964
Plate 5, Figures 1a–b
Rep., Geol. Surv. Japan, no. 204, p. 60. Syntypes: IGPS
no. 36452 (Existing) [=paratypes of Tellina maxima Nagao,
1928b figured in Nagao, 1928b, pl. 4, figs. 8, 11]. Type
locality: Taya, [Yamaga], Ashiya-machi, Onga-gun, Provinceof Chikuzen [Fukuoka Prefecture]. Stratigraphic unit: Yamaga
Beds [Yamaga Formation]. Geologic age: Oligocene.
Anisocorbula ohiroi Masuda, 1966b
Plate 5, Figures 2a–c

 rans. Proc. Palaeont. Soc. Japan, N.S., no. 64, p. 329,
T
pl. 35, figs. 35a–36b. Holotype: IGPS no. 90734 (formerly
DGS no. 4546; Existing). Type locality: Loc. No. 36(Masuda, 1966a), river cliff near small road, about 600
m SSE of Kakuma, [Otanimachi], Suzu City, Ishikawa
Prefecture. Stratigraphic unit: Higashi-innai Formation.
Geologic age: Miocene [latest Early Miocene].
Annuliconcha kitakamiensis Murata, 1964
Plate 5, Figure 3
Trans. Proc. Palaeont. Soc. Japan, N.S., no. 54, p. 227, pl.
34, fig. 12a, b. Holotype: IGPS no. 22389 (Existing). Type
locality: IGPS loc. no. It 044 (Murata, 1964), Imô, Yahagimachi, Rikuzen-takada City [currently Imo, Yahagicho,
Rikuzentakata City], Iwate Prefecture. Stratigraphic unit:
Lower part of Kanokura Formation. Geologic age: Middle
Permian.
Anomia pseudotruncata Yabe and Nagao, in Yabe,
Nagao and Shimizu, 1926
Plate 5, Figures 4–5
S ci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 9,
no. 2, p. 62, pl. 12, figs. 26, 27, pl. 13, figs. 26, 27, 36,
37. Syntypes: IGPS no. 22522 (Existing). Type locality:
Bômekizawa, Ôhinata[-mura], [currently Ohinata, Sakuhomachi], Minamisaku-gun, Province of Shinano [Nagano
Prefecture]. Stratigraphic unit: Ishidô Group [Shiroi
Formation]. Geologic age: Cretaceous [Early Cretaceous].
Arca (Arca) andoi Nomura, 1933
S ci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol.
16, no. 1, p. 33, pl. 3, fig. 8a–b. Holotype: IGPS no.
48383 (Missing). Type locality: Station no. 42 (Nomura,1933), 550 m E of Sankwakô, Tsûshô-shô, Byôritsugun, Shinchiku-shû [currently Sanwo-kou, Tongwan
Village, Tongsiao Township, Miaoli County], Taiwan.
Lithostratigraphic unit: Byôritsu Beds [Miaoli Formation].
Geologic age: Pliocene [Pleistocene].
Arca (Arca) kikaizimana Nomura and Zinbô, 1934
Plate 5, Figures 6a–b
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 16, no. 2,
p. 152 (44), pl. 5 (1), figs. 4a–5b. Holotype: IGPS no. 50198(Existing). Type locality: Plateaux area near Kamikatetu,
Kikai-zima, [Kikai-cho, Oshima-gun], Kagoshima Prefecture.
Lithostratigraphic unit: “Ryûkyû Limestone”. Geologic age:
Pleistocene. Remarks: This species is the type species of
Kikaiarca Noda, 1966a (by original designation).
Arca (Arca) miurensis Noda, 1966aPlate 5, Figures 7a–b
Sci. Rep. Tohoku Univ., 2nd Ser. (Geol.), vol. 38, no. 1, p.

Catalogue of type specimens of fossil Bivalvia
57, pl. 12, figs. 11, 12, 14, 15. Holotype: IGPS no. 23846(Existing). Type locality: Urago, Taura-machi, Miuragun [currently Uragocho, Yokosuka City], Kanagawa
Prefecture. Stratigraphic unit: Miyata Formation. Geologic
age: Pliocene [Pleistocene]. Remarks: Because Noda
(1966) thought Arca miyatensis Oyama, 1951b to be
unavailable, he proposed Arca (Arca) miyatensis for thesame taxon. However, A. miyatensis Oyama, 1951b isavailable, because Oyama (1951b) proposed this newname in association with a bibliographic refecence to Arca
kobeltiana Pilsbry sensu Yokoyama (1920). Habe (1967)
considered these two species are hardly distinguishedfrom the Recent Arca boucardi Jousseaume, 1894.
Arca (Noetia) pondaungensis var. transversa Nagao, 1928b
Plate 17, Figure 13
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 12,
no. 1, p. 26, pl. 6, figs. 8–10. Holotype: IGPS no. 36012(Existing). Type locality: Hôshuyama Mine, Hôshuyamamura [currently Tôhô-mura], Asakura-gun, Provinceof Chikuzen [Fukuoka Prefecture]. Stratigraphic unit:
Doshi Beds [=Doshiyama Formation]. Geologic age:
Eocene [Late Eocene]. Remarks: A new replacement
name, Noetia nagaoi MacNeil, 1938, was proposed forthis taxon since it is a primary junior homonym of Arcatransversa Say, 1822.
Arca sakamizuensis Hatai and Nisiyama, 1952
Plate 5, Figure 8
Sci. Rep. Tôhoku Univ., 2nd Ser. (Geol.), Spec. Vol., no. 3, p.
30. Holotype: IGPS no. 35996 (Existing). Type locality: Beachrocks along the sea coast [about 500 m N of] Sakamizu,
Shimago-mura, Onga-gun [currently An’ya, Wakamatsuku, Kitakyushu City], Fukuoka Prefecture. Stratigraphic unit:
Sakamizu Formation. Geologic age: Oligocene. Remarks:
This species was proposed as “n. sp.” without description,
but is regarded to be elected as “nom. nov.” for Arca (Arca)
sp. a of Nagao (1928b: Sci. Rep. Tôhoku Imp. Univ., 2nd Ser.
(Geol.), vol. 12, no. 1, p. 27, pl. 3, figs. 9, 9a).
Arca shinanoensis Yabe and Nagao, in Yabe, Nagao andShimizu, 1926
Plate 5, Figures 9–11
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 9, no.
2, p. 42, pl. 13, figs. 33–35. Syntypes: IGPS no. 22521(Existing). Type locality: Bômekizawa, Ôhinata[-mura],[currently Ohinata, Sakuho-machi], Minamisaku-gun,
Province of Shinano [Nagano Prefecture]; and Shiroi,[Narahara], Ueno-mura, Tano-gun, Province of Kôzuke
[Gumma Prefecture]. Stratigraphic unit: Ishidô Group [Shiroi
Formartion]. Geologic age: Cretaceous [Early Cretaceous].

17

Arca (Arca) sokeishiensis Nomura, 1933
Plate 5, Figure 12
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 16, no. 1,
p. 38, pl. 2, fig. 10a, b. Holotype: IGPS no. 47770 (Existing).
Type locality: Sôkeishi, Kwanden-shô, Sobun-gun, Tainanshû [currently Shuangsizih, Daci Village, Kuantien District,Tainan City], Taiwan. Lithostratigraphic unit: Byôritsu Beds
[Miaoli Formation]. Geologic age: Pliocene [Pleistocene].
Arca (Arca) takaoensis Nomura, 1933
Plate 5, Figure 13
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 16, no. 1,
p. 39, pl. 3, figs. 2–5. Holotype: IGPS no. 37444 (Existing).
Type locality: Shinsui, Ensô-shô, Okayama-gun, Takaoshû [currently Shengshui, Shengshui Village, Yangchao
District, Kaohsiung City], Taiwan. Stratigraphic unit: “KaizanBeds”. Geologic age: Miocene [Late Miocene].
Arca (Barbatia) yokoyamai Nomura, 1933
Plate 5, Figures 14a–e
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 16, no. 1,
p. 41, pl. 1, fig. 3a–d. Holotype: IGPS no. 42588 (Existing).
Type locality: Station no. 38 (Nomura, 1933), Wangwa,
Kôryû-shô, Chikunan-gun, Shinchiku-shû [currently Wanwa,
Jhonghe Village, Houlong Township, Miaoli County],
Taiwan. Lithostratigraphic unit: “Byôritsu Beds”. Geologic
age: Pliocene [Pleistocene]. Remarks: This species isthought to be a junior synonym of Striarca interplicata(Grabau and King, 1928) (Habe, 1958; Noda, 1966a).
Astarte minor Nagao, 1934
Plate 17, Figures 12a–b
Jour. Fac. Sci. Hokkaido Imp. Univ., Ser. 4, vol. 2, no.
3, p. 220, pl. 28, figs. 5–10. Lectotype: IGPS no. 66425(designated by Hayami, 1965b; Existing). Type locality:
Southern coast of Hiraiga Inlet [Wano, Tanohata-mura,
Shimohei-gun, Iwate Prefecture]. Stratigraphic unit:
Hiraiga Sandstone [Hiraiga Formation]. Geologic age:
Cretaceous [Early Cretaceous]. Remarks: The speciesname is preoccupied by Astarte sulcata var. minor
Jeffreys, 1864 [“1863”]. See Nicaniella (Trautscholdia)
nagaoi Matsubara, 2016.
Astarte miyakoensis Nagao, 1934
Jour. Fac. Sci. Hokkaido Imp. Univ., Ser. 4, vol. 2, no. 3,
p. 218, pl. 30, fig. 8, pl. 32, figs. 1, 3–5. Lectotype: IGPS
no. 7105 (designated by Hayami, 1965b; Missing). Type
locality: Southern coast of Hiraiga [Raga, Tanohatamura, Shimohei-gun, Iwate Prefecture]. Stratigraphic unit:
Hiraiga Sandstone [Hiraiga Formation]. Geologic age:
Cretaceous [Early Cretaceous].

18

Shin’ichi Sato, Tomoki Chiba, Takaki Yamanaka, Jun Nemoto, Masanori Shimamoto and Takashi Matsubara

Astarte shinanoensis Yabe and Nagao, in Yabe, Nagao
and Shimizu, 1926
Plate 5, Figure 15
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 9, no.
2, p. 47, pl. 13, figs. 29, 30. Lectotype: IGPS no. 22544(designated by Hayami, 1965b; Existing). Type locality:
Ishidô, near Ôhinata[-mura] [currently Ôhinata, Sakuhomachi], Minamisaku-gun, Province of Shinano [Nagano
Prefecture]. Stratigraphic unit: Ishidô Group [IshidôFormation]. Geologic age: Cretaceous [Early Cretaceous].
Astarte subomalioides Nagao, 1934
Plate 5, Figure 16
Jour. Fac. Sci. Hokkaido Imp. Univ., Ser. 4, vol. 2, no.
3, p. 219, pl. 27, figs. 3, 4. Lectotype: IGPS no. 66446(designated by Hayami, 1965b; Existing). Type locality:
Southern coast of Hiraiga Inlet [Wano, Tanohata-mura,
Shimohei-gun, Iwate Prefecture]. Stratigraphic unit:
Hiraiga Sandstone [Hiraiga Formation]. Geologic age:
Cretaceous [Early Cretaceous].
Astarte subsenecta Yabe and Nagao, in Yabe, Nagao
and Shimizu, 1926
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 9, no. 2,
p. 47, pl. 13, figs. 14–16, pl. 14, fig. 11. Lectotype: IGPS no.
22534 (designated by Hayami, 1965b; Missing). Type locality:
Ishidô, near Ôhinata[-mura] [currently Ishidô, Ôhinata,
Sakuho-machi], Minamisaku-gun, Province of Shinano[Nagano Prefecture]. Stratigraphic unit: Ishidô Group [IshidôFormation]. Geologic age: Cretaceous [Early Cretaceous].
Astarte subsenecta var. costata Yabe and Nagao, in
Yabe, Nagao and Shimizu, 1926
Plate 5, Figure 17
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 9, no. 2,
p. 48, pl. 14, fig. 10. Holotype: IGPS no. 22483 (Existing).
Type locality: Ôze, near Kagahara, Nakazato-mura [currently
Kanna-machi], Tano-gun, Province of Kôzuke [Gumma
Prefecture]. Stratigraphic unit: Kawarazawa Group [Ishido
Formation]. Geologic age: Cretaceous [Early Cretaceous].
Aviculopecten hataii Murata, 1964
Plate 5, Figures 18a–b
Trans. Proc. Palaeont. Soc. Japan, N.S., no. 54, p. 221, pl.
34, fig. 10a, b. Holotype: IGPS no. 85732 (Existing). Type
locality: IGPS loc. no. Mi 116 (Murata, 1964), Toya-zawa[Toyasawa], Kamishishiori [Shiraishi], Kesennuma City,
Miyagi Perfecture. Stratigraphic unit: Shigeji-zawa Member,
Kanokura Formation. Geologic age: Middle Permian.
Aviculopecten hayasakai Murata, 1964
Trans. Proc. Palaeont. Soc. Japan, N.S., no. 54, p. 219, pl.

34, fig. 4a, b. Holotype: IGPS no. 85737 (Missing). Type
locality: IGPS loc. no. Mi 116 (Murata, 1964), Toya-zawa[Toyasawa], Kamishishiori [Shiraishi], Kesennuma City,
Miyagi Perfecture. Stratigraphic unit: Shigeji-zawa Member,
Kanokura Formation. Geologic age: Middle Permian.
Aviculopecten minoensis Hayasaka, 1925b
Plate 5, Figure 19
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 8, no. 2,
p. 5, pl. 8, fig. 1. Lectotype: IGPS no. 22412 (designatedby Nakazawa, 1967; Existing). Type locality: Kinshôzan,Akasaka-machi, Province of Mino [currently Akasakacho,
Ogaki City, Gifu Prefecture]. Lithostratigraphic unit: theFusulina limestone [Akasaka Limestone]. Geologic age:
Middle Permian. Remarks: The name of species was
originally spelled as minöensis, and is corrected.
Aviculopecten? onukii Murata, 1969
Plate 5, Figures 20a–b
Saito Ho-on Kai Mus. Res. Bull., no. 38, p. 17, pl. 3, figs.
1a–d, 2a, b. Holotype: IGPS no. 91380 (Existing). Type
locality: Senmatsu, [Okago], Fujisawa-cho, HigashiIwai-gun [currently Fujisawacho, Ichinoseki City], Iwate
Prefecture. Stratigraphic unit: Upper part of Toyoma
Formation. Geologic age: Late Permian.
Aviculopecten reticularis Hayasaka, 1925b
Plate 5, Figure 21
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 8, no.
2, p. 6, pl. 8, fig. 2. Holotype: IGPS no. 22227 (Existing).
Type locality: Kinshôzan, Akasaka-machi, Province ofMino [currently Akasakacho, Ogaki City, Gifu Prefecture].
Lithostratigraphic unit: the Fusulina limestone [Akasaka
Limestone]. Geologic age: Middle Permian.
Aviculopecten sasakii Murata, 1964
Plate 5, Figures 22a–b
Trans. Proc. Palaeont. Soc. Japan, N.S., no. 54, p. 218, pl.
34, fig. 1a, b. Holotype: IGPS no. 85730 (Existing). Type
locality: IGPS loc. no. Mi 117 (Murata, 1964), the uppercourse of the Shigeji-zawa, Kamiyasse, Kesennuma City,
Miyagi Prefecture. Stratigraphic unit: Kanokura Formation.
Geologic age: Middle Permian.
Azorius philippianus Kotaka and Noda, 1977
Plate 5, Figure 23
Geol. Palaeont. Southeast Asia, vol. 18, p. 142, pl. 25,
figs. 9, 10. Holotype: IGPS no. 95087 (Existing). Type
locality: Road-side cliff, about 500 m S of Amuntay[Tagkawayan Municipality, Province of Quezon], westernpart of the Bondoc Peninsula, Philippines. Stratigraphic
unit: “Gumaca Formation”. Geologic age: Miocene.

Catalogue of type specimens of fossil Bivalvia
Barbatia (Acar) hayasakai Noda, 1966aPlate 5, Figures 24a–b
Sci. Rep. Tohoku Univ., 2nd Ser. (Geol.), vol. 38, no. 1, p.
67, pl. 4, figs. 10, 11. Holotype: IGPS no. 76249 (Existing).
Ogasawara (2001) erronerously cited IGPS no. 76429 as
the holotype. Type locality: Ogi, Nishikoshi-mura [currently
Izumozaki-machi], Santo-gun, Niigata Prefecture.
Stratigraphic unit: Funabashi Sandstone [=Haizume
Formation]. Geologic age: Late Pliocene [Pleistocene].
Barbatia kanezawaensis Akutsu, 1964
Plate 5, Figure 25
Sci. Rep. Tohoku Univ., 2nd Ser. (Geol.), vol. 35, no. 3, p.
282, pl. 59, fig. 2. Holotype: IGPS no. 85501 (Existing).
Type locality: About 1.8 km upstream from Kanezawa, theFukasawa valley, Shiobara-machi, Shioya-gun [currently
Nasushiobara City], Tochigi Prefecture. Stratigraphic unit:
Kanomatazawa Formation. Geologic age: Miocene [LateMiocene].
Barbatia (Acar) numaensis Noda, 1966aPlate 5, Figures 26a–b
Sci. Rep. Tohoku Univ., 2nd Ser. (Geol.), vol. 38, no. 1, p. 68,
pl. 5, figs. 11–12. Holotype: IGPS no. 86137. (Existing) Type
locality: Numa, Tateyama City, Chiba Pefecture. Stratigraphic
unit: Numa Coral Bed. Geologic age: Holocene.
Barbatia (Pugilarca) tsurushizakiensis Noda, 1966aPlate 5, Figure 27
Sci. Rep. Tohoku Univ., 2nd Ser. (Geol.), vol. 38, no. 1, p.
70, pl. 10, figs. 9, 10. Holotype: IGPS no. 17271 (Existing).
Type locality: Sea cliff, Tsurushizaki [sic, Tsurushu-misaki],[Higashicho], Hitachi City, Ibaragi [sic, Ibaraki] Prefecture.
Stratigraphic unit: Hatsuzaki Formation [=Hitachi
Formation]. Geologic age: Pliocene [Early Pliocene].
Barbatia (Barbatia) uetsukiensis Hatai and Nisiyama, 1949
Plate 5, Figure 28
J our. Paleont., vol. 23, no. 1, p. 89, pl. 23, figs. 6, 7.
Holotype: IGPS no. 72522 (Existing). Type locality: Road
side cuting about 100 m NE of the shrine at Dainichi-saka,
Uetsuki-mura [sic, Uetsuki-son] [currently Sho’o-cho],
Katsuta-gun, Okayama Prefecture. Stratigraphic unit:
Uetsuki Formation [=Yoshino Formation]. Geologic age:
Miocene [latest Early Miocene].
Barbatia (Pugilarca) yabei Noda, 1966aSci. Rep. Tohoku Univ., 2nd Ser. (Geol.), vol. 38, no. 1, p.
71, pl. 2, figs. 14, 15. Holotype: IGPS no. 25787 (Missing).
Type locality: Sea cliff, Tsurushizaki [sic, Tsurushu-misaki],[Higashicho], Hitachi City, Ibaragi [sic, Ibaraki] Prefecture.
Stratigraphic unit: Hatsuzaki Formation [=Hitachi

19

Formation]. Geologic age: Pliocene [Early Pliocene].
Bassina (Callanatis) hayasakai Kotaka, 1977
Geol. Palaeont. Southeast Asia, vol. 18, p. 128, pl. 22,
figs. 3–15. Holotype: IGPS no. 94933 (Missing). Type
locality: Yuanli Shell Mound, near Miaoli, [currently
Yuanli Township, Miaoli County], Taiwan. Geologic age:
Holocene.
Batissa nagaoi Suzuki, 1941
Plate 5, Figures 29a–b; Plate 6, Figure 1
Jour. Fac. Sci., Imp. Univ. Tokyo, Sec. 2, vol. 6, no. 3, p.
46, text-figs. 1a–3b, pl. 1, figs. 1a–2. Holotype: IGPS no.
35777 (Existing). Type locality: Kakize Mine, Takashima,
Takashima-machi [sic, Takashima-cho], Nishisonogigun [currently Takashimacho, Nagasaki City], Nagasaki
Prefecture. Stratigraphic unit: Hashima Formation. Geologic
age: Eocene [Middle Eocene]. Remarks: This species was
proposed as nom. nov. for Cyrena (Batissa) ponderosa
Nagao, 1928a, non Cyrena ponderosa Prime, 1860.
Batissa taiwanensis Nomura, 1933
Plate 6, Figures 2a–b
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 16,
no. 1, p. 68, pl. 1, figs. 1a–d. Holotype: IGPS no. 45087(Existing). Type locality: Sôkeishi, Kwanden-shô, Sobungun, Tainan-shû [currently Shuangsi, Daci Villege,
Kuantien District, Tainan City], Taiwan. Lithostratigraphic
unit: Byôritsu Beds [Miaoli Formation]. Geologic age:
Pliocene [Pleistocene].
Brachidontes takiensis Kamada, 1962
Plate 6, Figure 3
Palaeont. Soc. Japan Spec. Pap., no. 8, p. 73, pl. 3, figs.
15, 16. Holotype: IGPS no. 79380 (Existing). Type locality:
Sorida, Shimotaki, Tonomachi, Iwaki City, Fukushima
Prefecture. Stratigraphic unit: Iwaki Formation. Geologic age:
Oligocene [latest Late Eocene–earliest Early Oligocene].
Callista chinensis takagii Masuda, 1955
Plate 6, Figure 4
Trans. Proc. Palaeont. Soc. Japan, N.S., no. 20, p. 121,
pl. 19, fig. 7. Holotype: IGPS no. 90885 (formerly DGS no.
2501; Existing). Type locality: Tokunari, Machino-machi,
Fugeshi-gun [currently Wajima City], Ishikawa Prefecture.
Stratigraphic unit: Higashi-innai Formation. Geologic age:
Miocene [latest Early Miocene].
Callista pseudoplana Yabe and Nagao, 1925
Plate 6, Figure 5
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 7, no. 4,
p. 120, pl. 28, figs. 9, 10, pl. 29, figs. 2, 4. Lectotype: IGPS

20

Shin’ichi Sato, Tomoki Chiba, Takaki Yamanaka, Jun Nemoto, Masanori Shimamoto and Takashi Matsubara

no. 8553 (designated by Ichikawa and Maeda, 1963;
Existing). Type locality: The upper course of Ponnebets[u],a tributary of the Horomui, province of Ishikari [Manji,
Kurisawacho, Iwamizawa City], Hokkaido. Stratigraphic
unit: The Trigonia Sandstone [Middle Yezo Group in Manjiarea]. Geologic age: Cretaceous [Cenomanian–Turonian].
Callista sekiyaensis Akutsu, 1964
Plate 6, Figure 6
S ci. Rep. Tohoku Univ., 2nd Ser. (Geol.), vol. 35, no.
3, p. 285, pl. 59, figs. 8, 12. Syntypes: IGPS no. 85506(Existing). Type locality: Cliff of the Hoki River, about 100
m down stream from Daikoku-iwa, Sekiya, Shiobaramachi, Shioya-gun [curretly Nasushiobara City], Tochigi
Prefecture. Stratigraphic unit: Kanomatazawa Formation.
Geologic age: Miocene [Late Miocene].
Cardiomya (Cardiomya) kotakai Honda, 1989
Plate 6, Figure 7
Sci. Rep. Tohoku Univ., 2nd Ser. (Geol.), vol. 60, no. 1,
p. 90, pl. 5, fig. 6. Holotype: IGPS no. 99278 (Existing).
Type locality: NB-17 (Honda, 1989), riverside cliff along
the lowerstream of the Sakudano-sawa, a tributary of
the Chambetsu-gawa, Ombetsu-machi [sic, Ombetsucho], Shiranuka-gun [currently Onbetsu-cho, Kushiro
City], Hokkaido. Stratigraphic unit: Nuibetsu Formation.
Geologic age: Oligocene.
Cardita katsumatai Nagao, 1928b
Plate 6, Figures 8a–b
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 12, no.
1, p. 54, pl. 9, fig. 17. Holotype: IGPS no. 36331 (Existing).
Type locality: Namazuta Mine near Iizuka-machi, Kahogun [currently Iizuka City], Province of Hizen [Fukuoka
Prefecture]. Stratigraphic unit: Namazuta Fossil Bed[=Honso Formation]. Geologic age: Eocene.
Cardita kondoi Nagao, 1928b
Plate 6, Figure 9
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 12,
no. 1, p. 53, pl. 9, figs. 4–6. Holotype: IGPS no. 36407(Existing). Type locality: Tokuman, Ôshima, Provinceof Hizen [currently Oshimacho, Saikai City, Nagasaki
Prefecture]. Stratigraphic unit: Kakinoura Beds [Kakinoura
Formation]. Geologic age: Oligocene [latest Late Eocene–
earliest Early Oligocene].
Cardium (Acanthocardia) cancellatum Nomura, 1933
Plate 29, Figures 12a–b
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 16, no. 1,
p. 81, pl. 3, fig. 9a, b. Holotype: IGPS no. 46500 (Existing).
Type locality: Station no. 14 (Nomura, 1933), exposure,

550 m SE of Jô-tsûshô-wan, Tsûshô-shô, Byôritsu-gun,Shinchiku-shû [currently Shangtongsiaowan, Tongwan
Village, Tongsiao Township, Miaoli County], Taiwan.
Lithostratigraphic unit: Byôritsu Beds [Miaoli Formation].
Geologic age: Pliocene [Pleistocene]. Remarks: A new
replacement name Trifaricardium nomurai Kuroda andHabe, 1951 was proposed for this species, as it is a primary
junior homonym of Cardium cancellatum Gmelin, 1791.
Cardium (Trachycardium) hanpeizanense Nomura, 1933
Plate 6, Figure 10
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 16, no. 1,
p. 77, pl. 1, figs. 7a–8b, pl. 2, figs. 8, 9. Holotype: IGPS no.
37455 (Existing). Type locality: Hanpeizan, Okayama-gun,Takao-shû [currently Ban Pin Shan, Kaohsiung City], Taiwan.
Lithostratigraphic unit: “Riukiu Limestone” [correlative of the
Ryukyu Limestone]. Geologic age: Pleistocene.
Cardium (Cerastoderma) hanzawai Nomura, 1933
Plate 6, Figures 11a–b
S ci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol.
16, no. 1, p. 79, pl. 3, figs. 18a–19b. Holotype: IGPS
no. 46803 (Existing). Type locality: Station no. 25(Nomura, 1933), exposure, 1,050 m E of Hakushaton,Kôryû-shô, Chikunan-gun, Shinchiku-shû [currently
Baishaton, Tongsiao Township, Miaoli County], Taiwan.
Lithostratigraphic unit: Byôritsu Beds [Miaoli Formation].
Geologic age: Pliocene [Pleistocene].
Cardium (Cerastoderma) hizenense Nagao, 1928b
Plate 6, Figure 12
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 12,
no. 1, p. 61, pl. 10, figs. 15–17. Holotype: IGPS no.
36369 (Existing). Type locality: Iô-jima, off Nagasaki,[=Iôjima-mura, Nishisonogi-gun][currently Iôjimamachi,
Nagasaki City], Province of Hizen [Nagasaki Prefecture].
Stratigraphic unit: Iô-jima Beds [=Funazu Formation].
Geologic age: Eocene [Late Eocene].
Cardium (Trachycardium) infantile Nomura and Zinbô, 1934
Plate 6, Figures 13a–b
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 16,
no. 2, p. 156 (48), pl. 5 (1), fig. 17a, b. Holotype: IGPS
no. 50386 (Existing). Type locality: Plateaux area near
Kamikatetu, Kikai-zima, [currently Kikai-cho, Oshima-gun],Kagoshima Prefecture. Lithostratigraphic unit: “Ryûkyû
Limestone”. Geologic age: Pleistocene.
Cardium ishidoense Yabe and Nagao, in Yabe, Nagao
and Shimizu, 1926
Plate 6, Figure 14
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 9, no.

Catalogue of type specimens of fossil Bivalvia

212, p. 48, pl. 12, figs. 9, 16, 18. Lectotype: IGPS no. 22553(designated by Hayami, 1965b; Existing). IGPS no. 22533
cited by Tanaka and Toshimitsu (2003) is incorrect. Type
locality: Ishidô, near Ôhinata [currently Ishidô, Ôhinata,
Sakuho-machi], Minamisaku-gun, Province of Shinano[Nagano Prefecture]. Stratigraphic unit: Ishidô Group[Ishidô Formation]. Geologic age: Cretaceous [Early
Cretaceous].

Chlamys (Chlamys) akutsui Masuda, 1962aPlate 7, Figure 1
Sci. Rep. Tohoku Univ., 2nd Ser. (Geol.), vol. 33, no. 2, p.
160, pl. 18, fig. 29. Holotype: IGPS no. 28293 (Existing).
Type locality: River cliff of the Naka-gawa, south of
Ogawa-machi [currently Ogawa, Nakagawa-machi],Nasu-gun, Tochigi Prefecture. Stratigraphic unit: Ogane
Formation. Geologic age: Middle Miocene [Late Miocene].

Cardium (Cerastoderma?) kishimaense Nagao, 1928b
Plate 6, Figures 15a–b
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 12,
no. 1, p. 63, pl. 10, figs. 23, 25, 25a–b. Holotype: IGPS
no. 36367 (Existing). Type locality: Hanjô, Asahi-mura,
Kishima-gun [currently Takeo City], Province of Hizen[Saga Prefecture]. Stratigraphic unit: Kishima Beds
[Kishima Formation]. Geologic age: Oligocene [latest Late
Eocene–earliest Early Oligocene].

Chlamys chinkopensis Masuda and Sawada, 1961
Plate 7, Figure 2
Japan. Jour. Geol. Geogr., vol. 32, no. 1, p. 21, pl. 4, figs.
6, 7. Holotype: IGPS no. 90595 (formerly DGS no. 3896;
Existing). Ogasawara (2001) erroneously cited IGPS no.
90965 as the holotype. Type locality: Right river cliff ofthe Toshibetsu-gawa, about 1.5 km SE of Chinkope-toge,[=Hanaishi-toge], Imagane-machi [sic, Imakane-cho],
Setana-gun, Shiribeshi Province, Hokkaido. Stratigraphic
unit: Setana Formation. Geologic age: Pliocene [Early
Pleistocene].

Cardium miikense Nagao, 1928a
Plate 6, Figure 16
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 9, no.
3, p. 111, pl. 18, figs. 5, 5a–b. Lectotype: IGPS no. 35733(designated by Hatai and Nisiyama, 1952; Existing). Type
locality: Ôura [sic, Ôuramachi], Ômuta City, Fukuoka
Prefecture. Stratigraphic unit: “Upper Orthaulax japonicus
Zone” [=Nanaura Formation]. Geologic age: Eocene[Middle Eocene].
Cardium (Nemocardium) torii Nomura, 1933
Plate 6, Figures 17a–b
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 16,
no. 1, p. 79, pl. 2, fig. 6a, b. Holotype: IGPS no. 49000(Existing). Type locality: Fûsuirei, Station no. 26, Sachinshô, Shinkwa-gun, Tainan-shû [currently Fengchuiling,
Jhongjheng Village, Tsuochen Township], Taiwan.
Lithostratigraphic unit: Byôritsu Beds [Miaoli Formation].
Geologic age: Pliocene [Pleistocene].
Caryocorbula saikawai Kotaka, 1955
Plate 6, Figure 18
S aito Ho-on Kai Mus. Res. Bull., no. 24, p. 27, pl. 2,
figs. 5, 6. Holotype: IGPS no. 74010 (Existing). Type
locality: IGPS loc. no. Ao-15 (Kotaka, 1955), upper courseof the Isomatsu-gawa, Wakimoto-mura, Kitatsugarugun [currently Wakimoto, Goshogawara City], Aomori
Prefecture. Stratigraphic unit: Isomatsu Formation.
Geologic age: Oligocene [Early Miocene]. Remarks: The
present species is a junior synonym of Nipponomarcia
nakamurai (Ikebe, 1941) (Matsubara, 1995).

Chlamys cosibensis hanzawae Masuda, 1959
Plate 7, Figure 3
Trans. Proc. Palaeont. Soc. Japan, N.S., no. 35, p. 125, pl.
13, figs. 10a–15. Holotype: IGPS no. 90648 (formerly DGS
no. 3690; Existing). Type locality: Ukibuta, Higashiyurimura, Yuri-gun [currently Oikata, Higashiyuri, Yurihonjô
City], Akita Prefecture. Stratigraphic unit: Sugota Formation.
Geologic age: Early Miocene [early Middle Miocene].
Chlamys daishakaensis Masuda and Sawada, 1961
Plate 7, Figure 4
Japan. Jour. Geol. Geogr., vol. 32, no. 1, p. 23, pl. 4, figs.
8, 9. Holotype: IGPS no. 90708 (formerly DGS no. 3880;
Existing). Type locality: Right stream cliff, about 2.3 km N
of Daishaka Station along the Ou Line, Namioka-machi,
Minamitsugaru-gun [currently Namioka, Aomori City],
Aomori Prefecture. Stratigraphic unit: Daishaka Formation.
Geologic age: Pliocene [Early Pleistocene]. Remarks:
This species is a junior synonym of Chlamys (Leochlamys)
tanassevitschi (Khomenko, 1934) (Amano, 1994).
Chlamys (Chlamys) hanaishiensis Masuda, 1962aPlate 7, Figure 5
Sci. Rep. Tohoku Univ., 2nd Ser. (Geol.), vol. 33, no. 2, p.
166, pl. 22, figs. 1, 2. Holotype: IGPS no. 90565 (formerly
DGS no. 3935; Existing). Type locality: Left floor of the
Toshibetsu River, about 1 km SW of Pirika Station of the
Setana Line, Imagane-machi [sic, Imakane-cho], Setanagun, Shiribeshi Province, Hokkaido. Stratigraphic unit:
Setana Formation. Geologic age: Early Pliocene [Early
Pleistocene].

22

Shin’ichi Sato, Tomoki Chiba, Takaki Yamanaka, Jun Nemoto, Masanori Shimamoto and Takashi Matsubara

Chlamys (Chlamys) hasimotoi Masuda, 1962aPlate 7, Figure 6
Sci. Rep. Tohoku Univ., 2nd Ser. (Geol.), vol. 33, no. 2, p.
167, pl. 19, figs. 1, 2. Holotype: IGPS no. 90527 (formerly
DGS no. 3867; Existing). IGPS no. 90600 cited as theholotype by Ogasawara (2001) is the paratype. Type
locality: Natsukawa-en [=Honbetsu Koen], upstream of
the Honbetsu River [Higashihonbetsu], Honbetsu-machi[sic, Honbetsu-cho], Nakagawa-gun, Tokachi Province,
Hokkaido. Stratigraphic unit: Rawan Conglomerate ofOkuashiyoro Formation [=Honbetsu Formation]. Geologic
age: Early Pliocene.
Chlamys hataii Masuda and Akutsu, 1956
Plate 7, Figure 7
Trans. Proc. Palaeont. Soc. Japan, N.S., no. 21, p. 130,
pl. 20, figs. 1a–6b. Holotype: IGPS no. 90726 (formerly
DGS no. 1370; Existing). Type locality: Niiya, Tawara,
Kawachi-mura, Kawachi-gun [currently Ishii machi,
Utsunomiya City], Tochigi Prefecture. Stratigraphic unit:
Nagaoka Formation. Geologic age: Early Miocene [early
Middle Miocene]. Remarks: The present species issynonymous with Chlamys (Nomurachlamys) meisensis(Makiyama, 1926) (Kurihara, 2010).
Chlamys (Chlamys) hatakeyamae Masuda, 1962aPlate 7, Figure 8
Sci. Rep. Tohoku Univ., 2nd Ser. (Geol.), vol. 33, no. 2,
p. 168, pl. 18, figs. 18–20. Holotype: IGPS no. 27563(Existing). Type locality: Ikatsuchi-zawa at Kanayama,
Higashi-Takizawa, Yuri-mura, Yuri-gun [currenlty Ikazuchi,
Yurihonjô City] Akita Prefecture. Stratigraphic unit: Sugota
Formation. Geologic age: Early Miocene [Middle Miocene].
Chlamys imanishii Masuda and Sawada, 1961
Plate 7, Figures 9a–b
Japan. Jour. Geol. Geogr., vol. 32, no. 1, p. 25, pl. 4, figs.
10a–11. Holotype: IGPS no. 72555 (Existing). Type locality:
Sea cliff at Hamada, Yokohama-machi, Kamikita-gun,
Aomori Prefecture. Stratigraphic unit: Hamada Formation.
Geologic age: Early Pliocene [Early Pleistocene].
Chlamys (Chlamys) ishidae Masuda, 1962aPlate 7, Figure 10
Sci. Rep. Tohoku Univ., 2nd Ser. (Geol.), vol. 33, no. 2,
p. 171, pl. 18, figs. 21–24. Holotype: IGPS no. 90676(formerly DGS no. 3909; Existing). Type locality: Left cliffof the Do River, Tsuzara, Osawano-machi, Kaminikawagun [sic, Kaminiikawa-gun] [currently Tsuzuhara, Toyama
City], Toyama Prefecture. Stratigraphic unit: Kashio
Alternation of Kurosedani Formation. Geologic age: Early
Miocene [latest Early Miocene].

Chlamys (Chlamys) itoigawae Masuda, 1962aPlate 7, Figure 11
Sci. Rep. Tohoku Univ., 2nd Ser. (Geol.), vol. 33, no. 2,
p. 172, pl. 18, figs. 25, 26. Holotype: IGPS no. 90538(formerly DGS no. 3819; Existing). Type locality: Floor of
the Toki River, about 300 m upstream of the Relay Stationof Wireless Telegraph at Kamado [Kamadocho], Mizunami
City, Gifu Prefecture. Stratigraphic unit: Shukunohora
Formation. Geologic age: Early Miocene [latest Early–
earliest Middle Miocene].
Chlamys kitamurai Kotaka, 1955
Plate 7, Figure 12
Saito Ho-on Kai Mus. Res. Bull., no. 24, p. 26, pl. 2, fig. 2.
Holotype: IGPS no. 74009 (only a plastic cast is existing).
Type locality: IGPS loc. no. Ao-15 (Kotaka, 1955),
upper course of the Isomatsu-gawa, Wakimoto-mura,
Kitatsugaru-gun [currently Wakimoto, Goshogawara City],
Aomori Prefecture. Stratigraphic unit: Isomatsu Formation.
Geologic age: Oligocene [late Early Miocene].
Chlamys (Chlamys) kotakae Masuda, 1962aPlate 8, Figure 1
Sci. Rep. Tohoku Univ., 2nd Ser. (Geol.), vol. 33, no. 2, p.
175, pl. 20, figs. 8, 9. Holotype: IGPS no. 90646 (formerly
DGS no. 3832; Existing). Type locality: Cliff of the left side
valley of Kamifutamata-zawa, upper course of Biu River,about 5.5 km NE of Biu, Niikappu-machi [sic, Niikappu-cho],
Niikappu-gun, Hidaka Province, Hokkaido. Stratigraphic
unit: Noya Formation. Geologic age: Early Miocene.
Chlamys kumanodoensis Masuda, 1953aPlate 8, Figure 2
Trans. Proc. Palaeont. Soc. Japan, N.S., no. 12, p. 85, pl.
8, figs. 9a–12b. Holotype: IGPS no. 90522 (formerly DGS
no. 1049; Existing). Type locality: Hill side about 500 m Wof the Kumano Shrine, Kumanodô, Takadate-mura, Natorigun, [currently Takadate, Natori City], Miyagi Prefecture.
Stratigraphic unit: Moniwa Formation. Geologic age: Early
Miocene [early Middle Miocene].
Chlamys (Chlamys) matsunoi Masuda, 1962aPlate 8, Figure 3
Sci. Rep. Tohoku Univ., 2nd Ser. (Geol.), vol. 33, no. 2, p.
176, pl. 19, fig. 20, pl. 21, figs. 2–4. Holotype: IGPS no.
90627 (formerly DGS no. 3831; Existing). Type locality:
Cliff of the left side valley of Kamifutamata-zawa, uppercourse of Biu River, about 5.5 km NE of Biu, Niikappumachi [sic, Niikappu-cho], Niikappu-gun, Hidaka Province,
Hokkaido. Stratigraphic unit: Noya Formation. Geologic
age: Early Miocene.

Catalogue of type specimens of fossil Bivalvia

23

Chlamys (Chlamys) nagaoi Masuda, 1962aPlate 8, Figure 4
Sci. Rep. Tohoku Univ., 2nd Ser. (Geol.), vol. 33, no. 2, p.
179, pl. 20, figs. 4–6. Holotype: IGPS no. 36440 (Existing).
Type locality: Beach, west of the Hachiman-zaki [sic, Yahatamisaki], about 300 m N of Wakita [sic, Waita], Wakamatsu
City [currently An’ya, Wakamatsu-ku, Kitakyushu City],
Fukuoka Prefecture. Stratigraphic unit: Wakita [sic, Waita]
Formation. Geologic age: Late Oligocene.

Chlamys shitakaraensis Honda, 1980
Plate 8, Figures 9a–c
Trans. Proc. Palaeont. Soc. Japan, N.S., no. 117, p.
258, pl. 30, figs. 1, 2, 5, 6. Holotype: IGPS no. 95439(Existing). Type locality: Loc. B-06 (Honda, 1980), a small
northeastern tributary of the Urahioro-gawa, about 3.8 km
N of Rushin, Urahoro-machi [sic, Urahoro-cho], Tokachigun, Hokkaido. Stratigraphic unit: Shitakara Formation.
Geologic age: Oligocene [Middle–Late Eocene].

Chlamys (Chlamys) niikappuensis Masuda, 1962aPlate 8, Figure 5
Sci. Rep. Tohoku Univ., 2nd Ser. (Geol.), vol. 33, no. 2,
p. 180, pl. 21, fig. 5. Holotype: IGPS no. 90536 (formerly
DGS no. 3835; Existing). Type locality: Cliff of the left side
valley of the Kamifutamata-zawa, upper course of the
Biu River, about 5.5 km NE of Biu, Niikappu-machi [sic,Niikappu-cho], Niikappu-gun, Hidaka Province, Hokkaido.
Stratigraphic unit: Noya Formation. Geologic age: Early
Miocene.

Chlamys tamurae Masuda and Sawada, 1961
Plate 8, Figure 10
Japan. Jour. Geol. Geogr., vol. 32, no. 1, p. 27, pl. 4, figs.
12a–15. Holotype: IGPS no. 90550 (formerly DGS no.
3862; Existing). Type locality: Small hill-side exposure at
Maruyama [Shinsei], Kitahiyama-machi [sic, Kitahiyamacho], Setana-gun [currently Kitahiyama-ku, Setana-cho,
Kudo-gun], Shiribeshi Province, Hokkaido. Stratigraphic
unit: Setana Formation. Geologic age: Pliocene [Early
Pleistocene].

Chlamys oidensis Hatai, Masuda and Noda, 1974
Plate 8, Figures 6a–b
S aito Ho-on Kai Mus. Res. Bull., no. 43, p. 36, pl. 4,
figs. 1a–2. Holotype: IGPS no. 64419 (Existing). Type
locality: Oido, Motowakuya, Wakuya-machi, Toda-gun,
Miyagi Prefecture. Stratigraphic unit: Oido Formation.
Geologic age: Early Miocene [Middle Miocene]. Remarks:
The present species is a junior synonym of Chlamys(Leochlamys) arakawai (Nomura, 1935b) (Matsubara, in
Matsubara et al., 2014).

Circe (Circe) triangulus Kotaka and Noda, 1977
Plate 8, Figure 11
Geol. Palaeont. Southeast Asia, vol. 18, p. 137, pl. 25,
figs. 5, 6, text-fig. 2-2. Holotype: IGPS no. 95085 (Existing).
Type locality: Road-side cliff, about 500 m S of Amuntay,[Tagkawayan Municipality, Province of Quezon], westernpart of the Bondoc Peninsula, Philippines. Stratigraphic
unit: “Gumaca Formation”. Geologic age: Miocene.

Chlamys otukae Masuda and Sawada, 1961
Plate 8, Figures 7a–b
Japan. Jour. Geol. Geogr., vol. 32, no. 1, p. 19, pl. 4,
figs. 1a–5. Holotype: IGPS no. 90607 (formerly DGS no.
2616; Existing). Type locality: Road-side exposure at
Oido, Motowakuya [currently Kozuka], Wakuya-machi [sic,Wakuya-cho], Tôda-gun, Miyagi Prefecture. Stratigraphic
unit: Oido Formation. Stratigraphic unit: Oido Formation.
Geologic age: Early Miocene [Middle Miocene].
Chlamys (Chlamys) setsukoae Masuda, 1962b
Plate 8, Figures 8a–b
Saito Ho-on Kai Mus. Res. Bull., no. 31, p. 20, pl. 1, figs.
1a–8, pl. 2, figs. 6–8. Holotype: IGPS no. 90477 (formerly
DGS no. 4230; Existing). Type locality: Road-side cutting,about 900 m WSW of the Ken’yoshi Station of the Tohoku
Main Line, Ken’yoshi, Nagawa-machi [currently Nanbucho], Sannohe-gun, Aomori Prefecture. Stratigraphic unit:
Togawa Formation. Geologic age: Early Pliocene.

Clementia japonica Masuda, 1955
Plate 8, Figure 12
Trans. Proc. Palaeont. Soc. Japan, N.S., no. 20, p. 121,
pl. 19, fig. 8. Holotype: IGPS no. 90405 (formerly DGS no.
1382; Existing). Although Ogasawara (2001) stated IGPS
no. 90880 as the holotype, it is a paratype. Type locality:
Tokunari, Machino-machi, Fugeshi-gun [currently Wajima
City], Ishikawa Prefecture. Stratigraphic unit: Higashi-innai
Formation. Geologic age: Miocene [latest Early Miocene].
Clementia (Clementia) nakosoensis Kamada, 1962
Plate 8, Figures 13a–b
Palaeont. Soc. Japan Spec. Pap., no. 8, p. 119, pl. 13, figs.
15a, b. Holotype: IGPS no. 79385 (Existing). Type locality:
Kokozura, Nakoso City [currently Nakosomachi, Iwaki
City], Fukushima Prefecture. Stratigraphic unit: Kokozura
Formation. Geologic age: Miocene [latest Early Miocene].
Clinocardium asagaiense arakawae Kamada, 1962
Plate 9, Figures 1a–b
Palaeont. Soc. Japan Spec. Pap., no. 8, p. 105, pl. 10,
figs. 15–17. Holotype: IGPS no. 79383 (Existing). Type

24

Shin’ichi Sato, Tomoki Chiba, Takaki Yamanaka, Jun Nemoto, Masanori Shimamoto and Takashi Matsubara

locality: Mukaida, Yumoto-machi, Joban City [currently
Joban-Yumotomachi, Iwaki City], Fukushima Prefecture.
Stratigraphic unit: Asagai Formation. Geologic age:
Oligocene [Early Oligocene].
Clinocardium asagaiense makiyamae Kamada, 1962
Plate 9, Figures 2a–b
Palaeont. Soc. Japan Spec. Pap., no. 8, p. 104, pl. 10, figs.
18–21. Holotype: IGPS no. 15800 (Existing). Type locality:
Nabezuka [sic, Nabetsuka], [Kamikitaba], Hirono-machi,
Futaba-gun, Fukushima Prefecture. Stratigraphic unit: Asagai
Formation. Geologic age: Oligocene [Early Oligocene].
Clinocardium chikagawaense Kotaka, 1950
Plate 9, Figures 3a–b
Short Pap. Inst. Geol. Paleont., Tôhoku Univ., no. 2, p. 46,
pl. 5, figs. 1–6. Holotype: IGPS no. 72999 (Existing). Type
locality: IGPS loc. no. Ao-51 (Kotaka, 1950), a sea cliff at
the outlet of the Chikagawa River at Chikagawa, Tanabumachi, Shimokita-gun [currently Okunai, Mutsu City],
Aomori Prefecture. Stratigraphic unit: Hamada Formation.
Geologic age: Pliocene [Early Pleistocene].
Clinocardium hataii Hayasaka, 1956
Plate 9, Figure 4
Saito Ho-on Kai Mus. Res. Bull., no. 25, p. 18, pl. 2, fig.
3a, b. Holotype: IGPS no. 77375 (Existing). Type locality:
Loc. No. 1 (Hayasaka, 1956), cliff of the Takasegawa
River west of Takakura, [Ide], about 5 km SW of the
Namie-machi Railroad station on the Joban Line, Namiemachi, Futaba-gun, Fukushima Prefecture. Stratigraphic
unit: Ishiguma Formation. Geologic age: Pliocene.
"Clinocardium" nomurai Hayasaka, 1956
Plate 9, Figure 5
Saito Ho-on Kai Mus. Res. Bull., no. 25, p. 18, pl. 2, fig. 4a,b. Holotype: IGPS no. 77376 (Existing). Type locality: Loc.
2 (Hayasaka, 1956), path side cutting at Onoda, Namiemachi, Futaba-gun, Fukushima Prefecture. Stratigraphic
unit: Ishiguma Formation. Geologic age: Pliocene.
Clinocardium omagariense Honda, 1981b
Plate 9, Figures 6a–c
Trans. Proc. Palaeont. Soc. Japan, N.S., no. 122, p. 130,
pl. 15, figs. 1–13. Holotype: IGPS no. 95740-1 (Existing).
Type locality: Loc. no. OM-32 (Honda, 1981b), riverside
cliff along the Urahoro-gawa, about 1,250 m NNE from thejunction between the Urahoro-gawa and the Tokomurogawa, Urahoro-machi [sic, Urahoro-cho], Tokachi-gun,
Hokkaido. Stratigraphic unit: Omagari Formation. Geologic
age: Oligocene.

Codakia kitamurai Hatai and Nisiyama, 1949
Plate 9, Figure 7
J our. Paleont., vol. 23, no. 1, p. 91, pl. 24, figs. 5, 6.
Holotype: IGPS no. 72526 (Existing). Type locality:
Shukunohora, Hiyoshi-mura, Toki-gun [currently
Hiyoshicho, Mizunami City], Gifu Prefecture. Stratigraphic
unit: Hiyoshi Formation [=Shukunohora Member of Akeyo
Formation]. Geologic age: Miocene [latest Early–earliest
Middle Miocene].
Codakia (Jagonia) okinawazimana Nomura and Zinbô, 1936
Plate 9, Figures 8a–b
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 18,
no. 3, p. 241, pl. 11, figs. 9a–b. Holotype: IGPS no.
51319 (Existing). Type locality: Gabusoga, Haneji-mura,
Kunigami-gun [currently Nago City], Okinawa Prefecture.
Stratigraphic unit: Simaziri Beds [=Haneji Formation].
Geologic age: Early Pliocene [Early Pleistocene].
Codakia semipolita Nomura, 1933
Plate 9, Figures 9a–b
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 16, no.
1, p. 73, pl. 1, figs. 9a–10b. Holotype: IGPS no. 45086(Existing). Type locality: The upper course of Sairyôkyô,Sachin-shô, Shinkawa-gun, Tainan-shû [currently Yseiliao
Bridge, Yseiliao, Ronghe Village, Tsuochen District, Tainan
City], Taiwan. Lithostratigraphic unit: Byôritsu Beds [Miaoli
Formation]. Geologic age: Pliocene [Pleistocene].
Conularia rectangularis Hayasaka, 1920
Plate 9, Figures 10a–c
J our. Geol. Soc. Japan, vol. 27, no. 327, p. 87, textfigs. 1–3. Lectotype: IGPS no. 7320 (designated by
Sugiyama, 1942; Existing). Type locality: Iimô, Yahagimura, Kesen-gôri, prov. Rikuchyû [sic, Rikuzen] [currently
Imo, Yahagicho, Rikuzentakata City, Iwate Prefecture].
Stratigraphic unit: Lower part of Kanokura Formation.
Geologic age: Middle Permian. Remarks: This species
was originally thought a conulariid (Cnidaria), and agenus Neoconularia Sugiyama, 1942 was proposed with
Conularia rectangularis Hayasaka, 1920 as the typespecies. However, it is referable to Aviculopinna Meek,
1864, and thus Neoconularia Sugiyama, 1942 is a juniorsynonym of Aviculopinna (Nakazawa and Newell, 1968;
Hayami and Kase, 1977).
Corbicula (Corbicula) kotakai Honda, 1981aPlate 9, Figures 11a–b
Trans. Proc. Palaeont. Soc. Japan, N.S., no. 121, p. 22,
pl. 2, figs. 6, 9, 10, 13–17. Holotype: IGPS no. 96758(Existing). Type locality: Loc. no. YB-10 (Honda, 1981a),river side cliff along the Rubeshube-zawa, a tributary

Catalogue of type specimens of fossil Bivalviaof the Chokubetsu-gawa, Urahoro-machi [sic, Urahorocho], Tokachi-gun, Hokkaido. Stratigraphic unit: Yubetsu
Formation. Geologic age: Oligocene [Middle Eocene].
Corbicula (Veloritina?) sanchuensis Yabe and Nagao, in
Yabe, Nagao and Shimizu, 1926
Plate 9, Figures 12–15
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 9, no.
2, p. 53, pl. 12, figs. 8, 8a, pl. 13, figs. 8–10, 17, 17a.
Syntypes: IGPS nos. 22449, 22467 (Existing). Type
locality: Bômeki-zawa, Ôhinata[-mura], [currently Ôhinata,
Sakuho-machi], Minamisaku-gun, Province of Shinano[Nagano Prefecture]. Stratigraphic unit: Shiroi Group [Shiroi
Formation]. Geologic age: Cretaceous [Early Cretaceous].
Corbicula (Corbicula) sachalinensis Suzuki, 1943
Plate 9, Figures 16a–b
Venus, vol. 12, nos. 3–4, p. 163–165 (in Japanese), p.
167–169 (in Germany), pl. 8, figs. 1–3, 5, 6. Holotype:
IGPS no. 8353c (Existing). Type locality: Mittellauf desTuimis, Nord-Sachalin, USSR [currently Middle reachesof T'ym River in Tymovskoe or Nogliki District, Sakhalin
Oblast, Russia (precise locality unknown)]. Stratigraphic
unit: Mãch-Gruppe. Geologic age: earliest Miocene(Aquitanian).
Corbicula (Corbicula) shimizui Suzuki, 1943
Plate 9, Figures 17a–b
Venus, vol. 12, nos. 3–4, p. 161–162 (in Japanese), p.
166–167 (in Germany), pl. 7, pl. 8, figs. 4, 7. Holotype:
IGPS no. 8353a (Existing). Type locality: Mittellauf desTuimis, Nord-Sachalin, USSR [currently Middle reachesof T'ym River in Tymovskoe or Nogliki District, Sakhalin
Oblast, Russia (precise locality unknown)]. Stratigraphic
unit: Mãch-Gruppe. Geologic age: earliest Miocene(Aquitanian).
Corbicula takasago Nomura, 1933
Plate 9, Figure 18
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 16, no. 1,
p. 69, pl. 2, figs. 1a–2b. Holotype: IGPS no. 45079 (Existing).
Type locality: Station no. 34 (Nomura, 1933), Wangwa,
Kôryû-shô, Chikunan-gun, Shinchiku-shû [currently Wanwa,
Jhonghe Village, Houlong Township, Miaoli County], Taiwan.
Lithostratigraphic unit: Byôritsu Beds [Miaoli Formation].
Geologic age: Pliocene [Pleistocene].
Corbula (Cunaecorbula) kyushuensis Nagao, 1928a
Plate 9, Figure 19
S ci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 9,
no. 3, p. 116, pl. 21, fig. 16. Lectotype: IGPS no. 35834(designated by Hatai and Nisiyama, 1952; Existing). Type25

locality: Akase, Ôda-mura, Uto-gun, Province of Higo[currently Akasemachi, Uto City, Kumamoto Prefecture].
Stratigraphic unit: “Lower Orthaulax japonicus Zone”[=Fukuregi Formation]. Geologic age: Eocene [early
Middle Eocene].
Corbula (Corbula) subtumida Nagao, 1928a
Plate 9, Figure 20
S ci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 9,
no. 3, p. 115, pl. 22, fig. 6. Lectotype: IGPS no. 35828(designated by Hatai and Nisiyama, 1952; Existing).
Type locality: Takesaki, Kôyagi-jima, [Koyagi-mura],
Nishisonogi-gun, [currently Koyagimachi, Nagasaki City],
Province of Hizen [Nagasaki Prefecture]. Stratigraphic
unit: “Lower Orthaulax japonicus Zone” [=Futagojima
Formation]. Geologic age: Eocene [Middle Eocene].
Corbula (Corbula) taiwanensis Nomura, 1933
Plate 9, Figures 21a–b
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 16,
no. 1, p. 106, pl. 3, figs. 10a–11b. Holotype: IGPS no.
48966 (Existing). Type locality: Station no. 18 (Nomura,1933), Wangwa, Kôryû-shô, Chikunan-gun, Shinchiku-shû[currently Wanwa, Jhonghe Village, Houlong Township,
Miaoli County], Taiwan. Lithostratigraphic unit: Byôritsu Beds
[Miaoli Formation]. Geologic age: Pliocene [Pleistocene].
Crassatellites asakuraensis Nagao, 1928b
Plate 9, Figure 22
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 12, no.
1, p. 49, pl. 2, fig. 22. Holotype: IGPS no. 36258 (Existing).
Type locality: Hôshuyama Mine, Hôshuyama-mura [currently
Tôhô-mura], Asakura-gun, Province of Chikuzen [Fukuoka
Prefecture]. Stratigraphic unit: Doshi Beds [Doshiyama
Formation]. Geologic age: Eocene [Late Eocene].
Crassatellites formosanus Nomura, 1933
Plate 9, Figure 23
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 16, no. 1,
p. 66, pl. 1, figs. 14, 15. Holotype: IGPS no. 37441 (Existing).
Type locality: About 1,500 m NNW of the Police station of
Shinsui, Ensô-shô, Okayama-gun, Takao-shû [currently
Shengshui, Shengshui Village, Yangchao District, KaohsiungCity], Taiwan. Lithostratigraphic unit: Byôritsu Beds [Miaoli
Formation]. Geologic age: Pliocene [Pleistocene].
Crassatellites inconspicuus Nagao, 1928b
Plate 10, Figures 1a–b
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 12, no.
1, p. 50, pl. 2, figs. 14, 14a. Holotype: IGPS no. 36308(Existing). Type locality: Tokuman, Ôshima, [Kurose-mura,Nishisonogi-gun], Province of Hizen [currently Oshimacho,

26

Shin’ichi Sato, Tomoki Chiba, Takaki Yamanaka, Jun Nemoto, Masanori Shimamoto and Takashi Matsubara

Saikai City, Nagasaki Prefecture]. Stratigraphic unit:
Kakinoura Beds [Kakinoura Formation]. Geologic age:
Oligocene [latest Eocene–earliest Oligocene].
Crassatellites matsuraensis Nagao, 1928b
Plate 10, Figures 2a–c
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 12, no.
1, p. 50, pl. 3, figs. 20, 20a. Holotype: IGPS no. 36310(Existing). Type locality: Ôbô, near Arita-machi [currently
Arita-cho], Nishimatsura-gun [sic, Nishimatsuura-gun],
Province of Hizen [Saga Prefecture]. Stratigraphic
unit: Kishima Beds [Kishima Formation]. Geologic age:
Oligocene [latest Eocene–earliest Oligocene].
Crassatellites tosanus Nomura, 1937
Plate 10, Figures 3a–b
Japan. Jour. Geol. Geogr., vol. 14, nos. 3–4, p. 81, pl. 6,
fig. 8a, b. Lectotype: IGPS no. 54638 (designated by Hatai
and Nisiyama, 1952; Existing). Type locality: Tônohama,Yasuda-mura [sic, Yasuda-cho], Aki-gun, Kôti [=Kochi]
Prefecture. Stratigraphic unit: Tosa Pliocene [=Ananai
Formation]. Geologic age: Pliocene [latest Late Pliocene–
early Early Pleistocene].
Crassatellites yabei Nagao, 1928b
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 12, no.
1, p. 48, pl. 2, figs. 19, 19a, b. Holotype: IGPS no. 36388(Missing). Type locality: Iwaya, Shimagô-mura, [Ongagun] Province of Hizen [currently Arige, Wakamatsu-ku,
Kitakyushu City, Fukuoka Prefecture]. Stratigraphic unit:
Wakita Beds [=Waita Formation]. Geologic age: Oligocene.
Crassostrea gravitesta eoilensis Kim, Noda and Yoon,
1974
Plate 10, Figures 4a–b
Trans. Proc. Palaeont. Soc. Japan, N.S., no. 93, p. 274,
pl. 38, fig. 15, pl. 39, fig. 1a–c. Holotype: IGPS no. 92938(Existing). Type locality: Loc. no. 1-1 (Kim et al., 1974), S
of Songjeon-ri, Yangbug-myeon, Weolseong-gun [currently
Gyeongju City], Gyeongsangbuk-do, Korea. Stratigraphic
unit: Eoil Formation. Geologic age: Miocene [latest Early–
earliest Middle Miocene]. Remarks: This “subspecies”
is thought an ecophenotype of Crassostrea gravitesta(Yokoyama, 1926) (Matsubara et al., 2011).
Crassostrea sunakozakaensis Ogasawara, 1976
Plate 10, Figures 5a–b
Sci. Rep. Tohoku Univ., 2nd Ser. (Geol.), vol. 46, no. 2,
p. 45, pl. 11, figs. 19–21, pl. 12, figs. 15, 20. Holotype:
IGPS no. 95018 (Existing). Type locality: Loc. no. Su01 (Ogasawara, 1976), river side cliff of Asano-gawa at
Higashi-Ichise [sic, Higashi-ichinosemachi], Kanazawa

City, Ishikawa Prefecture. Stratigraphic unit: Sunakozaka
Formation. Geologic age: Miocene [latest Early Miocene].
Crenella (Megacrenella) nuibetsuensis Honda, 1989
Plate 10, Figure 6
Sci. Rep. Tohoku Univ., 2nd Ser. (Geol.), vol. 60, no. 1, p.
60, pl. 4, figs. 4, 17. Holotype: IGPS no. 97096 (Existing).
Type locality: Loc. NB-08 (Honda, 1989), Muri-kawa,
Ombetsu-machi [sic, Ombetsu-cho], Shiranuka-gun [currently
Onbetsucho, Kushiro City], Hokkaido. Stratigraphic unit:
Nuibetsu Formation. Geologic age: Oligocene.
Crenella (Megacrenella) shitakaraensis Honda, 1989
Plate 10, Figure 7
Sci. Rep. Tohoku Univ., 2nd Ser. (Geol.), vol. 60, no. 1,
p. 61, pl. 4, fig. 18. Holotype: IGPS no. 97111 (Existing).
Type locality: Loc. SK-46 (Honda, 1989), upstream of
Chokubetsu-gawa, Urahoro-machi [sic, Urahoro-cho],
Tokachi-gun, Hokkaido. Stratigraphic unit: Shitakara
Formation. Geologic age: Oligocene [Middle–Late Eocene].
Crenella striatocostata Nagao, 1928b
Plate 10, Figure 8
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 12, no. 1,
p. 46, pl. 6, figs. 20, 21. Holotype: IGPS no. 36291 (Existing).
Type locality: Kojôbaru, Hôshuyama-mura, Province of
Chikuzen [currently Tôhô-mura, Asakura-gun, Fukuoka
Prefecture]. Stratigraphic unit: Kawamagari Beds [Kawamagari
Formation]. Geologic age: Eocene [Middle Eocene].
Crenella subfornicata Nagao, 1928b
Plate 10, Figures 9a–c
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 12,
no. 1, p. 46, pl. 5, figs. 8–9b. Holotype: IGPS no. 36288(Existing). Type locality: Taya, [Yamaga], Ashiya-machi,
Onga-gun, Province of Chikuzen [Fukuoka Prefecture].
Stratigraphic unit: Yamaga Beds [Yamaga Formation].
Geologic age: Oligocene.
Crenella tomiyaensis Hatai and Nakamura, 1940
Plate 10, Figure 10
B ull. Biogeogr. Soc. Japan, vol. 10, no. 7, p. 113, fig.
1. Holotype: IGPS no. 61354 (Existing). Type locality:
Tomiya, Tomiya-mura [currently Tomiya-machi], Kurokawagun, Miyagi Prefecture. Stratigraphic unit: Nanakita Beds
[Nanakita Formation]. Geologic age: Miocene [late Late
Miocene].
Crenipecten kesenensis Hayasaka, 1925b
Plate 10, Figure 11
S ci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 8,
no. 2, p. 11, pl. 8, figs. 8, 8a. Holotype: IGPS no. 22383

Catalogue of type specimens of fossil Bivalvia(Existing). Type locality: Imô, [Yahagi-mura], Kesen-gôri,prov. Rikuzen [currently Imo, Yahagi-cho, Rikuzentakata
City, Iwate Prefecture]. Stratigraphic unit: Kanokura
Formation. Geologic age: Middle Permian. Remarks: The
name of genus was originally spelled as Cenipecten, andis corrected.
Ctena hataii Masuda, 1966b
Plate 10, Figures 12a–b
Trans. Proc. Palaeont. Soc. Japan, N.S., no. 64, p. 325,
pl. 35, figs. 15–20. Holotype: IGPS no. 90729 (formerly
DGS no. 4524; Existing). IGPS no. 90865 (=formerly DGS
no. 4523) cited as the holotype by Ogasawara (2001)
is the paratype. Type locality: Loc. No. 30 (Masuda,
1966a), road side cutting near Koeiji Temple, Otani [sic,Otanimachi], Suzu City, Ishikawa Prefecture. Stratigraphic
unit: Higashi-innai Formation. Geologic age: Miocene[latest Early Miocene].
Ctenodonta manchuriensis Endo, 1935
Plate 10, Figures 13a–b
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 16,
no. 4, p. 197, pl. 14, figs. 7–8. Holotype: IGPS no. 55582(Existing). Type locality: In the black banded limestone
at the northern foot of San-len hill, near the Wu-hu-tsui
colliery, [Fuzhou County, Fengtian Province], Liaotung,Manchoukuo [currently Wafangdian, Dalian City, Liaoning
Province, China]. Stratigraphic unit: Kangyao Formation.
Geologic age: Early Ordovician.
Ctenodonta takahashii Endo, 1935
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 16, no.
4, p. 196, pl. 14, figs. 1–6. Syntype: IGPS no. unknown(Missing). Type locality: In the black banded limestone
at the northern foot of San-leng hill, near the Wu-hu-tsui
colliery, [Fuzhou County, Fengtian Province], Liaotung,Manchuokuo [currently Wafangdian, Dalian City, Liaoning
Province, China]. Stratigraphic unit: Kangyao Formation.
Geologic age: Early Ordovician.
Cucullaea delicatostriata Yabe and Nagao, 1925
Plate 10, Figure 14
S ci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 7,
no. 4, p. 113, pl. 28, fig. 1. Lectotype: IGPS no. 8555
(designated by Hayami, 1975; Existing). Type locality:
The Cape Khoi Bed, exposed along the sea-cliff 1 km S ofCape Khoi, near Alexandrovsk, north Saghalin [currently
Alexandrovsk-Sakhalinsky District, Sakhalin Oblast,Russia]. Stratigraphic unit: Werblud Group ("Cape Khoi
beds") in Alexandrovsk area. Geologic age: Cenomanian–Turonian, Cretaceous.

27

Cucullaea ezoensis Yabe and Nagao, 1928
Plate 10, Figure 15
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 9, no.
3, p. 81, pl. 16, figs. 1–3. Syntypes: IGPS nos. 22601,
22611 (Existing). Type locality: Futamatano-sawa, Miruto,[Kurisawa-mura], Sorachi-gun, province of Ishikari[currently Kurisawacho, Iwamizawa City, Hokkaido].
Stratigraphic unit: Mikasa Formation. Geologic age:
Cenomanian–Turonian, Cretaceous.
Cucullaea iriomotensis Masuda and Sato, 1988
S aito Ho-on Kai Spec. Pub., no. 2 (Prof. T. Kotaka
Commem. Vol.), p. 442, pl. 1, figs. 3–7. Holotype: IGPS
no. 99713 (Missing). Type locality: Sea cliff at Nishizaki,
about 1 km NW of Nakano, Taketomi-cho, Iriomote-jima,[Yaeyama-gun], Okinawa Prefecture. Stratigraphic unit:
Iriomote Formation. Geologic age: early Middle Miocene[latest Early–earliest Middle Miocene].
Cucullaea nipponica Nagao, 1928b
Plate 11, Figures 1a–c
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 12, no.
1, p. 25, pl. 6, figs. 1, 1a, b. Holotype: IGPS no. 35999(Existing). Type locality: Western sea cliff, about 400 m
W of the primary school at Ipponmatsu, Iô-jima, Iô-jimamura, Nishisonogi-gun [currently Nagasaki City], Nagasaki
Prefecture. Stratigraphic unit: Funatsu Formation.
Geologic age: Eocene [Late Eocene].
Cultellus? brevis Nagao, 1928a
Plate 11, Figure 2
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 9, no. 3,
p. 114, pl. 22, fig. 1. Holotype: IGPS no. 35788 (Existing).
Type locality: Koyagi-jima, [Koyagi-mura], Nishisonogigun, Province of Hizen [currently Koyagimachi, Nagasaki
City, Nagasaki Prefecture]. Stratigraphic unit: “Upper
Orthaulax japonicus Zone” [=Futagojima Formation].
Geologic age: Eocene [early Middle Eocene].
Cultellus? leguminoides Nagao, 1928b
Plate 11, Figure 3
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 12, no.
1, p. 84, pl. 4, fig. 21. Holotype: IGPS no. 36447 (Existing).
Type locality: Iô-jima, off Nagasaki [Iojima-mura],
Nishisonogi-gun [currently Iojimamachi, Nagasaki City],
Province of Hizen [Nagasaki Prefecture]. Stratigraphic
unit: Iôjima Beds [Funatsu Formation]. Geologic age:
Eocene [Late Eocene].
Cyclina (Cyclina) asagaiensis Kamada, 1952
Plate 11, Figures 4a–b
Trans. Proc. Palaeont. Soc. Japan, N.S., no. 6, p. 169,

28

Shin’ichi Sato, Tomoki Chiba, Takaki Yamanaka, Jun Nemoto, Masanori Shimamoto and Takashi Matsubarapl. 15, fig. 3a–c. Holotype: IGPS no. 72955 (Existing).
Type locality: IGPS loc. no. Fs-11, in the Ômachi
abandoned shaft of the Joban Coal Mining Co., N of
Takasaka, Uchigô-machi, Iwaki-gun [currently Uchigotakasakamachi, Iwaki City], Fukushima Prefecture.
Stratigraphic unit: Asagai Formation. Geologic age:
Oligocene [Early Oligocene].
Cyclina compressa Nagao, 1928b
Plate 11, Figure 5
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 12, no. 1, p.
76, pl. 10, fig. 11. Holotype: IGPS no. 36415 (Existing). Type
locality: Ôbô, Arita-machi [currently Arita-cho], Nishimatsuuragun, Province of Hizen [Saga Prefecture]. Stratigraphic unit:
Kishima Beds [Kishima Formation]. Geologic age: Oligocene[latest Late Eocene–earliest Early Oligocene].
Cyclina (Cyclina) japonica Kamada, 1952
Plate 11, Figures 6a–b
Trans. Proc. Palaeont. Soc. Japan, N.S., no. 6, p. 168, pl.
15, figs. 1a–b, 2 [non fig. 4; =Cyclina (Cyclina) jobanica(Yokoyama, 1924), fide Kamada, 1962]. Holotype: IGPS
no. 72952 (Existing). Type locality: IGPS loc. no. Iw-3
(Kamada, 1952), 200 m W of Tokunari, Machino-machi,
Fugeshi-gun [currently Machinomachi, Wajima City],
Ishikawa Prefecture. Stratigraphic unit: Higashi-innai
Formation. Geologic age: Miocene [latest Early Miocene].
Cyclina? nodai Nagao, 1928b
Plate 11, Figures 7a–b
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 12,
no. 1, p. 75, pl. 7, figs. 19, 20, 26. Holotype: IGPS no.
36442 (Existing). Type locality: The Namazuta Mine near
Iizuka-machi, Kaho-gun [currently Iizuka City], Provinceof Chikuzen [Fukuoka Prefecture]. Stratigraphic unit:
Namazuta Fossil Bed [=Honso Formation]. Geologic age:
Eocene [Middle Eocene].
Cyclina (Cyclina) umbonata Kotaka and Noda, 1977
Plate 11, Figure 8
Geol. Palaeont. Southeast Asia, vol. 18, p. 141, pl. 25, fig.
15. Holotype: IGPS no. 95090 (Existing). Type locality:
Road-side cliff, about 500 m S of Amuntay, [Tagkawayan
Municipality, Province of Quezon], western part ofthe Bondoc Peninsula, Philippines. Stratigraphic unit:“Gumaca Formation”. Geologic age: Miocene.
Cyclocardia fujinaensis Ogasawara and Nomura, 1980
Plate 11, Figure 9
P rof. S. Kanno Mem. Vol., p. 90, pl. 9, figs. 11a–17.
Holotype: IGPS no. 96024 (Existing). Type Locality: Loc.
no. Fj-10 (Ogasawara and Nomura, 1980), road sidecliff about 400 m E of Shinji Station of National Railway,
Shinji-cho, Yatsuka-gun [currently Shinjicho, Matsue City],
Shimane Prefecture. Stratigraphic unit: Fujina Formation.
Geologic age: Miocene [late Middle Miocene].
Cyrena mirabilis Nagao, 1928b
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 12, no.
1, p. 52 (42), pl. 7, fig. 27. Holotype: IGPS no. unknown(Missing). Although Ogasawara (2001) cited the holotype
as IGPS no. 36264, it is the paratypes. Type Locality:
Iwaya Mine, Kiuragi-mura, Higashimatsura-gun [sic,Higashimatsuura-gun] [currently Iwaya, Kyuragimachi,
Karatsu City], Saga Prefecture. Stratigraphic unit:
Yoshinotani Beds [Yoshinotani Formation]. Geologic age:
Oligocene [Late Eocene].
Cyrena otsukai Yabe and Nagao, in Yabe, Nagao andShimizu, 1926
Plate 11, Figures 10–13
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 9, no.
2, p. 50, pl. 13, figs. 20–24. Syntypes: IGPS nos. 22453,
22476 (Existing). Type Locality: Bômekizawa, Ohinata[mura] [currently Sakuho-machi], Minamisaku-gun, Provinceof Shinano [Nagano Prefecture]; Kagikake, Ohinatamura [currently Kagikakezawa, Ohinata, Sakuho-machi],Minamisaku-gun, Province of Shinano [Nagano Prefecture];
and Shiroi, Ueno-mura, Tano-gun, Province of Kôzuke
[Gumma Prefecture]. Stratigraphic unit: Shiroi Group [Shiroi
Formation]. Geologic age: Cretaceous [Early Cretaceous].
Cyrena (Batissa) ponderosa Nagao, 1928a
Plate 5, Figures 29a–b; Plate 6, Figure 1
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 9, no.
3, p. 108, pl. 20, figs. 10, 30, pl. 21, fig. 21, pl. 22, figs.
16–19. Syntypes: IGPS nos. 35685, 35691, 35708, 35709,
35777, 35811, 35812 (Existing). Type locality: Urakoshi,
Oniki-mura, Amakusa-Shimo-shima [currently Onikimachi, Ushibuka City, Kumamoto Prefecture]; Kakize
Mine, Takashima off Nagasaki, province of Hizen [currently
Takashimacho, Nagasaki City, Nagasaki Prefecture];Kôyagi-jima off Nagasaki, province of Hizen [currently
Kouyagi-machi, Nagasaki City, Nagasaki Prefecture].
Stratigraphic unit: Toishi Beds [Toishi Formation]; Hashima
Beds [Hashima Formation]. Geologic age: Eocene [early
Middle Eocene]. Remarks: Suzuki (1941) proposed Batissa
nagaoi as nom. nov. for Cyrena (Batissa) ponderosa
Nagao, 1928a because the latter name is a primary juniorhomonym of Cyrena ponderosa Prime, 1860.
Cyrena radiatostriata Yabe and Nagao, in Yabe, Nagao
and Shimizu, 1926
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 9, no.

Catalogue of type specimens of fossil Bivalvia
2, p. 51, pl. 12, figs. 29–35. Lectotype: IGPS no. 35523(designated by Hayami, 1965b; Missing). Type locality:
Hachimanzawa [currently Kagahara, Kanna-machi], Tanogun, Province of Kôzuke [Gumma Prefecture]. Stratigraphic
unit: Shiroi Group [Sebayashi Formation]. Geologic age:
Cretaceous [late Early Cretaceous (Aptian–Albian)].
Cyrena shiroiensis Yabe and Nagao, in Yabe, Nagao andShimizu, 1926
Plate 11, Figures 14–16
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 9, no. 2,
p. 52, pl. 14, figs. 4–6, 19, 20, 22, 25. Syntypes: IGPS no.
22451 (Existing). Type locality: Bômekizawa, Ohinata[-mura],
Saku-machi [currently Sakuho-machi], Minamisaku-gun,[Nagano Prefecture], Province of Shinano; and Shiroi, Uenomura, Tano-gun, Province of Kôzuke [Gumma Prefecture].
Stratigraphic unit: Shiroi Group [Shiroi Formation]. Geologic
age: Cretaceous (early Early Cretaceous].
Cyrena shiroiensis var. alata Yabe and Nagao, in Yabe,
Nagao and Shimizu, 1926
Plate 11, Figures 17–19
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 9, no.
2, p. 53, pl. 13, fig. 26, pl. 14, figs. 15, 28. Syntypes: IGPS
nos. 22443, 22447 (Existing). Type locality: Shiroi, Uenomura, Tano-gun, Province of Kôzuke [Gumma Prefecture];
and Bômekizawa, Ohinata[-mura], Saku-machi [currently
Sakuho-machi], Minamisaku-gun, Province of Shinano[Nagano Prefecture]. Stratigraphic unit: Shiroi Group[Sebayashi and Shiroi formations]. Geologic age:
Cretaceous [Early Cretaceous].
Daonella densisulcata Yabe and Shimizu, 1927
Plate 11, Figure 20
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 11,
no. 2, p. 124, pl. 12, fig. 9a. Holotype: IGPS no. 7571
(Existing). Type locality: Zôhôin near Sakawa, Provinceof Tosa [currently Ko, Sakawa-cho, Takaoka-gun, Kochi
Prefecture]. Stratigraphic unit: Zohoin Group [KochigataniGroup]. Geologic age: Ladinian, Triassic.
Daonella densisulcata var. subquadrata Yabe andShimizu, 1927
Plate 11, Figure 21
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 11,
no. 2, p. 124, pl. 12, fig. 8. Holotype: IGPS no. 7890
(Existing). Type locality: Zôhôin near Sakawa, Provinceof Tosa [currently Ko, Sakawa-cho, Takaoka-gun, Kochi
Prefecture]. Stratigraphic unit: Zohoin Group [KochigataniGroup]. Geologic age: Ladinian, Triassic.

29

Daonella kotoi var. alta Yabe and Shimizu, 1927
Plate 11, Figure 22
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 11,
no. 2, p. 122, pl. 12, fig. 10. Holotype: IGPS no. 5298
(Existing). Type locality: Zôhôin near Sakawa, Provinceof Tosa [currently Ko, Sakawa-cho, Takaoka-gun, Kochi
Prefecture]. Stratigraphic unit: Zohoin Group [KochigataniGroup]. Geologic age: Ladinian, Triassic.
Daonella kotoi var. multistriata Yabe and Shimizu, 1927
Plate 11, Figures 23–24
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 11, no. 2,
p. 123, pl. 11, figs. 12, 14, pl. 13, fig. 11. Syntypes: IGPS nos.
22998, 7892, 35278 (Existing). Type locality: Railway cuttings
NE of the Rifu Station, Okinoi (IGPS no. 22998), Yawata,
Tagajô-mura [currently Tagajo City]; NE of Rifu Station (IGPS
no. 7892) and Hamada (IGPS no. 35278), Rifu-mura [currently
Rifu-cho], Miyagi-gun, Miyagi Prefecture. Stratigraphic unit:
Rifu Formation. Geologic age: Ladinian, Triassic.
Diplodonta confusa Nagao, 1928b
Plate 11, Figure 25
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 12, no.
1, p. 60, pl. 10, figs. 8, 10, 10a. Holotype: IGPS no. 36351(Existing). Type locality: Asakawa, Shimagô-mura, Ongagun [currently Yahatanishi-ku, Kitakyushu City], Privince of
Chikuzen [Fukuoka Prefecture]. Stratigraphic unit: Yamaga
Beds [Yamaga Formation]. Geologic age: Oligocene.
Diplodonta? problematica Nagao, 1928a
Plate 11, Figure 26
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 9, no.
3, p. 110, pl. 22, figs. 4, 4a. Lectotype: IGPS no. 35827(designated by Hatai and Nisiyama, 1952; Existing).
Type locality: Mirokudake [=Jiromaru-dake], AmakusaKamishima, Amakusa-gun [currently Matsushimamachi,
Kamiamakusa City], Kumamoto Prefecture. Stratigraphic
unit: “Lower Orthaulax japonicus Zone” [=Fukuregi
Formation]. Geologic age: Eocene [early Middle Eocene].
Dosinia chikuzenensis Nagao, 1928b
Plate 11, Figure 27
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 12,
no. 1, p. 64, pl. 10, fig. 5. Holotype: IGPS no. 36277(Existing). Type locality: Sakamizu, Shimagô-mura, Ongagun [currently Wakamatsu-ku, Kitakyushu City], Province of
Chikuzen [Fukuoka Prefecture]. Stratigraphic unit: Sakamizu
Beds [Sakamizu Formation]. Geologic age: Oligocene.
Dosinia ettyuensis Hatai and Nisiyama, 1939 [“1938”]
Plate 11, Figure 28
J apan. Jour. Geol. Geogr., vol. 16, nos. 1–2, p. 150,

30

Shin’ichi Sato, Tomoki Chiba, Takaki Yamanaka, Jun Nemoto, Masanori Shimamoto and Takashi Matsubara

pl. 9, fig. 4. Holotype: IGPS no. 62429 (Existing). Type
locality: A cliff of the Zintu-gawa [=Jintsû-gawa] at Zyônô[=Jônô], Sugihara-mura, Nei-gun, Ettyû Province [currently
Jônô, Yatsuomachi, Toyama City, Toyama Prefecture]
Stratigraphic unit: Zyônô Beds [=Otogawa Formation].
Geologic age: Pliocene [Late Miocene].
Dosinia (Phacosoma) hataii Masuda, 1963
Plate 12, Figures 1a–c
Trans. Proc. Palaeont. Soc. Japan, N.S., no. 49, p. 34, pl.
6, figs. 1a–2. Holotype: IGPS no. 72476 (Existing). Type
locality: Hattomaki, Hanawa-machi, Higashishirakawagun, Fukushima Prefecture. Stratigraphic unit: Kubota
Formation. Geologic age: Middle Miocene [Late Miocene].
Dosinia (Kaneharaia) kaneharai fujinaensis Masuda, 1967
Plate 12, Figure 2
Saito Ho-on Kai Mus. Res. Bull., no. 36, p. 24, pl. 1, fig.
9, pl. 2, fig. 3. Holotype: IGPS no. 90888 (Existing). Type
locality: S of Morinoko [sic, Mominoko], Fujina, Tamayumachi [sic, Tamayu-cho], Yatsuka-gun [currently Tamayucho,
Matsue City], Shimane Prefecture. Stratigraphic unit: Fujina
Formation. Geologic age: Middle Miocene.
Dosinia (Kaneharaia) kannoi Masuda, 1963
Plate 12, Figures 3a–b
Trans. Proc. Palaeont. Soc. Japan, N.S., no. 49, p.
35, pl. 6, figs. 6a–b, 7a–b. Holotype: IGPS no. 64682(Existing). Type locality: Kinseidô, Kinshôdô, Eihokumen, Kisshu-gun, Kankyô-hokudo [currently Yonpuk, Kiljugun, Hamgyongpuk-to], North Korea. Stratigraphic unit:
Heiroku Formation. Geologic age: Early Miocene [latest
Early–earliest Middle Miocene].
Dosinia (Phacosoma) okinawaensis Masuda and Sato, 1988
S aito Ho-on Kai Spec. Pub., no. 2 (Prof. T. Kotaka
Commem. Vol.), p. 446, pl. 3, figs. 12–17. Holotype: IGPS
no. 99716 (Missing). Type locality: Sea cliff at Nishizaki,
about 1 km NW of Nakano, Taketomi-cho, Iriomote-jima,[Yaeyama-gun], Okinawa Prefecture. Stratigraphic unit:
Iriomote Formation. Geologic age: early Middle Miocene[latest Early–earliest Middle Miocene].
Dosinia tatunokutiensis Nomura, 1938
Plate 12, Figures 4a–b
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 19, no.
2, p. 256, pl. 34, figs. 2–6, pl. 35, fig. 11a, b. Holotype:
IGPS no. 15944 (Existing). Type locality: Gôroku cliff along
the right bank of the Hirosegawa-River, western borderof Sendai [Gôroku, Hirose-mura, Miyagi-gun] [currently
Gôroku, Aoba-ku, Sendai City], Miyagi Prefecture.
Stratigraphic unit: Tatunokuti Shell Bed [=Tatsunokuchi

Formation]. Geologic age: Early Pliocene [latest Late
Miocene–earliest Early Pliocene].
Ennucula praenipponica Kamada, 1962
Plate 12, Figures 5a–c
Palaeont. Soc. Japan Spec. Pap., no. 8, p. 42, pl. 1, figs.
8–11. Holotype: IGPS no. 79375 (Existing). Type locality:
Hieda, Shimokajiro, Ena-machi [currently Nagasaki], Iwaki
City, Fukushima Prefecture. Stratigraphic unit: Honya
Formation. Geologic age: Miocene [Early Miocene].
Entomonotis kurosawai Sakaguti, 1939
Jub. Publ. Commem. Prof. H. Yabe 60th Birthday, vol.
1, p. 229, pl. 15, figs. 1–6. Syntypes: IGPS no. unknown(Missing). Type locality: Nisi-Togaki, Hosoura [currently
Shizugawa, Minamisanriku-cho], Motoyosi-gun, Miyagi
Prefecture. Stratigraphic unit: Saragai beds [Saragai
Group]. Geologic age: Norian, Triassic. Remarks: The
name of genus was originally spelled as Entomotis, and iscorrected.
Ezocallista kurodae Kamada, 1962
Plate 12, Figures 6a–b
P alaeont. Soc. Japan Spec. Pap.,, no. 8, p. 110, pl.
12, figs. 1a–3. Holotype: IGPS no. 79384 (Existing).
Type locality: Karasawa, Kuroda, Tabito-mura [currently
Tabitomachi, Iwaki City], Fukushima Prefecture.
Stratigraphic unit: Iwaki Formation. Geologic age:
Oligocene [latest Late Eocene–earliest Early Oligocene].
Fimbria yaeyamaensis Masuda and Sato, 1988
S aito Ho-on Kai Spec. Pub., no. 2 (Prof. T. Kotaka
Commem. Vol.), p. 445, pl. 3, figs. 7, 8. Holotype: IGPS
no. 99714 (Missing). Type locality: Sea cliff at Nishizaki,
about 1 km NW of Nakano, Taketomi-cho, Iriomote-jima,[Yaeyama-gun], Okinawa Prefecture. Stratigraphic unit:
Iriomote Formation. Geologic age: early Middle Miocene[latest Early–earliest Middle Miocene].
Fortipecten kuroishiensis Kotaka and Noda, 1967
Plate 13, Figure 1
Saito Ho-on Kai Mus. Res. Bull., no. 36, p. 40, pl. 1, fig.
21. Holotype: IGPS no. 90767 (Existing). Type locality:
Loc. no. 10 (Kotaka and Noda, 1967), downstream of the
Karasuzawa Dam of Nakano-gawa, [Okawara], Kuroishi
City, Aomori Prefecture. Stratigraphic unit: Ogawara
Formation. Geologic age: Middle Natotian [Late Miocene].
Gafrarium (Circe) hanzawai Nomura and Zinbô, 1936
Plate 13, Figures 2a–b
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 18,
no. 3, p. 245, pl. 11, figs. 14a–15b. Holotype: IGPS no.

Catalogue of type specimens of fossil Bivalvia
51348 (Existing). Type locality: Gabusoga, Haneji-mura,
Kunigami-gun [currently Nago City], Okinawa Prefecture.
Stratigraphic unit: Simaziri Beds [=Haneji Formation.
Geologic age: Early Pliocene [Early Pleistocene].
Gari (Gari) pitogoensis Kotaka and Noda, 1977
Plate 13, Figure 3
Geol. Palaeont. Southeast Asia, vol. 18, p. 145, pl. 25,
fig. 8. Holotype: IGPS no. 95086 (Existing). Type locality:
Road-side cliff, about 500 m S of Amuntay, [Tagkawayan
Municipality, Province of Quezon], western part ofthe Bondoc Peninsula, Philippines. Stratigraphic unit:“Gumaca Formation”. Geologic age: Miocene.
Gervillia shinanoensis Yabe and Nagao, in Yabe, Nagao
and Shimizu, 1926
Plate 13, Figure 4
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 9, no. 2, p.
59, pl. 14, figs. 17, 18. Syntypes: IGPS no. 22507 (Existing).
Type locality: Kagikake and Bômekizawa, Ohinata[-mura][currently Ohinata, Sakuho-machi], Minamisaku-gun,
Province of Shinano [Nagano Prefecture]. Stratigraphic
unit: Shiroi Group [=Ishido Formation]. Geologic age:
Cretaceous [early Early Cretaceous].
Glans naomiae Masuda, 1966b
Plate 13, Figures 5a–b
Trans. Proc. Palaeont. Soc. Japan, N.S., no. 64, p. 325,
pl. 35, figs. 13a–14. Holotype: IGPS no. 90181 (formerly
DGS no. 4517; Existing). Type locality: Loc. No. 35(Masuda, 1966a), sea coast, about 500 m N of the outletof the Fushimi-gawa, Konami, [Misakimachi], Suzu City,
Ishikawa Prefecture. Stratigraphic unit: Higashi-innai
Formation. Geologic age: Miocene.
Gloripallium izurense Masuda, 1958aPlate 13, Figure 6
Trans. Proc. Palaeont. Soc. Japan, N.S., no. 31, p. 227,
pl. 32, figs. 4–5b. Holotype: IGPS no. 90541 (formerly
DGS no. 3577; Existing). Type locality: Izura, Ôtsumachi [sic, Otsucho], Kitaibaraki City, Ibaraki Prefecture.
Stratigraphic unit: Kokozura Formation. Geologic age:
Miocene [latest Early Miocene]. Remarks: The name of
species was originally spelled as izurensis. However, thecorrected original spelling should be izurense because
Gloripallium is a neuter noun.
Glycimeris altoumbonata Nagao, 1928b
Plate 13, Figure 7
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 12,
no. 1, p. 28, pl. 3, fig. 14. Holotype: IGPS no. 36013(Existing). Type locality: Hôshuyama Mine, Hôshuyama-

31

mura [currently Tôhô-mura], Asakura-gun, Province of
Chikuzei [Fukuoka Prefecture]. Stratigraphic unit: Doshi
Beds [=Doshiyama Formation]. Geologic age: Eocene.
Remarks: The original spelling of the genus name is anerror for Glycymeris.
Glycimeris amakusensis Nagao, 1930
Plate 13, Figures 8–10
Jour. Fac. Sci. Hokkaido Imp. Univ., Ser. 4, vol. 1, no. 1, p.
15, pl. 2, figs. 4–7. Syntypes: IGPS no. 42727 (Existing).
Type locality: Wadanohana, Takado-mura, Amakusa-Kamishima [Amakusa-gun] [currently Takado, Ryugatakemachi,
Kamiamakusa City, Kumamoto Prefecture]. Stratigraphic
unit: Himenoura Group. Geologic age: Coniacian–Santonian, Cretaceous. Remarks: The original spelling of
the genus name is an error for Glycymeris.
Glycimeris cisshuensis var. compressa Nagao, 1928b
Plate 13, Figures 11a–b
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 12, no.
1, p. 30, pl. 3, figs. 25, 25a–c. Holotype: IGPS no. 36001(Existing). Type locality: Sakamizu, Shimagô-mura, Ongagun, Province of Chikuzen [currently An'ya, Wakamatsuku, Kitakyushu City, Fukuoka Prefecture]. Stratigraphic
unit: Wakita Beds [Waita Formation]. Geologic age:
Oligocene. Remarks: The original spelling of the genus
name is an error for Glycymeris. Although this name isa secondary junior homonym of Glycymeris compressa(Weller, 1907), a new replacement name is unnecessary,
because it is synonymous with Glycymeris (Veletuceta)
cisshuensis (Makiyama, 1926). Glycymeris compressa
Kanno, 1956 [non Weller, 1907, nec Nagao, 1928b], wasrenamed as Glycymeris kannoi Matsukuma, 1979.
Glycymeris gorokuensis Nomura, 1938
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 19, no.
2, p. 246, pl. 33, figs. 12–14. Holotype: IGPS no. 16142(Missing). Type locality: Gôroku cliff along the right bankof the Hirosegawa-River, [Gôroku, Hirose-mura, Miyagigun] [currently Gôroku, Aoba-ku, Sendai City], Miyagi
Prefecture. Stratigraphic unit: Tatunokuti Shell Bed[=Tatsunokuchi Formation]. Geologic age: Pliocene [latest
Late Miocene–earliest Early Pliocene].
Glycymeris hanzawai Nomura and Zinbô, 1934
Plate 13, Figures 12a–b
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 16, p.
152 (44), pl. 5 (1), fig. 3a, b. Holotype: IGPS no. 50196(Existing). Type locality: Plateaux area near Kamikatetu,
Kikai-zima, [Kikai-cho, Oshima-gun], Kagoshima
Prefecture. Lithostratigraphic unit: “Ryûkyû Limestone”.
Geologic age: Pleistocene.

32

Shin’ichi Sato, Tomoki Chiba, Takaki Yamanaka, Jun Nemoto, Masanori Shimamoto and Takashi Matsubara

Glycymeris iriomotensis Masuda and Sato, 1988
S aito Ho-on Kai Spec. Pub., no. 2 (Prof. T. Kotaka
Commem. Vol.), p. 443, pl. 1, figs. 8–14. Holotype: IGPS
no. 99712 (Missing). Type locality: Sea cliff at Nishizaki,
about 1 km NW of Nakano, Taketomi-cho, Iriomote-jima,[Yaeyama-gun], Okinawa Prefecture. Stratigraphic unit:
Iriomote Formation. Geologic age: early Middle Miocene[latest Early–earliest Middle Miocene].
Glycymeris nakosoensis Hatai and Nisiyama, 1949
Plate 13, Figures 13a–b
Jour. Paleont., vol. 23, no. 1, p. 88, pl. 25, figs. 15, 16.
Holotype: IGPS no. 72502 (Existing). Type locality: A
small cliff in front of the dormitory of the Nippon Coal
Mining Company, Nakoso-machi, Iwaki-gun [currently
Nakosomachi, Iwaki City], Fukushima Prefecture.
Stratigraphic unit: Iwaki Formation. Geologic age:
Oligocene [latest Late Eocene–earliest Early Oligocene].
Glycymeris nozokiensis Hatai and Nisiyama, 1951
Plate 13, Figure 14
Saito Ho-on Kai Mus. Res. Bull., no. 21, p. 1. Figs. 1, 2.
Holotype: IGPS no. 72883 (Existing). Type locality: Road
side cutting on the main highway about 500 m S of the
Nozoki Primary School, Nozoki-mura [currently Nozoki,
Mamurogawa-machi, Mogami-gun], Yamagata Prefecture.
Stratigraphic unit: Nozoki Formation. Geologic age:
Miocene [early Middle Miocene].
Glycymeris ogawaraensis Kotaka and Noda, 1967
Plate 13, Figure 15
Saito Ho-on Kai Mus. Res. Bull., no. 36, p. 38, pl. 2, fig.
1. Holotype: IGPS no. 90739 (Existing). Type locality:
Loc. no. 10 (Kotaka and Noda, 1967), down stream of
the Karasuzawa Dam of the Nakanogawa, Ogawara [sic,Okawara], Kuroishi City, Aomori Prefecture. Stratigraphic
unit: Ogawara Formation. Geologic age: “Middle Natorian”[Late Miocene].
Glycymeris oshimaensis Noda, 1962b
Plate 13, Figure 16
Sci. Rep. Tohoku Univ., 2nd Ser. (Geol.), vol. 34, no. 3, p.
229, pl. 16, fig. 8. Holotype: IGPS no. 79057 (Existing).
Type locality: Loc. No. 924 (Noda, 1962b), road sidecliff, about 300 m E of the pass between Oshima andHosono, Matsunoyama-machi [sic, between Oshimamura and Yasuzuka-machi], Higashikubiki-gun [currently
between Kamitate, Oshimaku and Hosono, Yasuzukaku,
Joetsu City], Niigata Prefecture. Stratigraphic unit: Kubiki
Formation. Geologic age: Middle Miocene.

Glycymeris rhynconelloides Nomura and Hatai, 1939[“1938”]
Plate 13, Figure 17
Japan. Jour. Geol. Geogr., vol. 16, nos. 1–2, p. 5, pl. 1,
fig. 6. Holotype: IGPS no. 60022 (Existing). Type locality:
Husina [sic, Huzina][=Fujina], Tamayu-mura Yatuka-gun[=Yatsuka-gun], Izumo [currently Tamayumachi, Matsue
City, Shimane Prefecture]. Stratigraphic unit: Izumo
Neogene [=Fujina Formation]. Geologic age: Middle
Miocene. Remarks: Matsukuma and Okamoto (1986)
regarded this species as nomen dubium because theholotype is poorly preserved.
Glycymeris subpectiniformis Nomura and Zinbô, 1934
Plate 13, Figures 18a–b
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 16, p.
151 (43), pl. 5 (1), fig. 2a, b. Holotype: IGPS no. 50200(Existing). Type locality: Plateaux area near Kamikatetu,
Kikai-zima, [Kikai-cho, Oshima-gun], Kagoshima
Prefecture. Lithostratigraphic unit: “Ryûkyû Limestone”.
Geologic age: Pleistocene.
Glycymeris yamaguchii Hayasaka, 1956
Plate 13, Figures 19a–b
Saito Ho-on Kai Mus. Res. Bull., no. 25, p. 16, pl. 2, figs.
1a–2c. Holotype: IGPS no. 77373 (Existing). Type locality:
Loc. No. 1 (Hayasaka, 1956), cliff of the Takasegawa
River west of Takakura, about 5 km SW of the Namiemachi Railroad Station on the Joban Line, Namie-machi,
Futaba-gun, Fukushima Prefecture. Stratigraphic unit:
Ishiguma Formation. Geologic age: Pliocene.
Grammatodon yokoyamai Yabe and Nagao, in Yabe,
Nagao and Shimizu, 1926
Plate 13, Figure 20
S ci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 9,
no. 2, p. 44, pl. 12, figs. 12–13, 25. Lectotype: IGPS no.
22555 (designated by Hayami, 1965a; Existing). Type
locality: Ishidô, near Ohinata[-mura] [currently Ishidô,Ôhinata, Sakuho-machi], Minamisaku-gun, Nagano
Prefecture. Stratigraphic unit: Ishidô Group [IshidôFormation]. Geologic age: Cretaceous [late Early–early
Late Cretaceous].
Halicardia akitaensis Ogasawara and Takayasu, 1982
Plate 13, Figures 21a–b
Venus, vol. 41, no. 3, p. 199, pl. 1, fig. 1a–f. Holotype:
IGPS no. 96013 (Existing). Originally cited as “monotype”.
Type locality: A road-side cliff about 500 m downstream
from Fudotaki (water-fall), Iwami-Sannai, Kawabe-machi,
Kawabe-gun [currently Kurosawa, Taihei, Akita City], Akita
Prefecture. Stratigraphic unit: Sunakobuchi Formation.

Catalogue of type specimens of fossil Bivalvia
Geologic age: late Early–early Middle Miocene.
Halicardia miyagiensis Fujiwara, 1992
Plate 13, Figures 22a–b
Saito Ho-on Kai Mus. Nat. Hist. Res. Bull., no. 60, p. 9,
figs. 1–4. Holotype: IGPS no. 101555 (Existing). Type
locality: Nishino-iri, Tomiya-cho [sic, Tomiya-machi],
Kurokawa-gun, Miyagi Prefecture. Stratigraphic unit:
Nanakita Formation. Geologic age: late Late Miocene.
Homomya? dubia Yabe and Nagao, in Yabe, Nagao andShimizu, 1926
Plate 14, Figures 1–3
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 9, no. 2.
p. 55, pl. 12, figs. 11, 39, pl. 14, fig. 1. Syntypes: IGPS no.
22505 (Existing). Type locality: Hachimanzawa [currently
Kagahara, Kanna-machi], Tano-gun, Province of Kôzuke
[Gumma Prefecture]. Stratigraphic unit: Shiroi Group[Sebayashi Formation]. Geologic age: Cretaceous [lateEarly Cretaceous (Albian–Aptian)].
Horiopleura yaegashii Yehara, 1920
Jour. Geol. Soc. Japan, vol. 27, no. 321, p. 41, pl. 1, figs.
1–3, pl. 2, figs. 1–3. Syntypes: IGPS nos. 22348, 22349(Missing). Type locality: Moshi, Iwaizumi [currently Omoto,
Iwaizumi-cho] and Hiraiga, Tanohata-mura, Shimoheigun, Iwate Prefecture. Stratigraphic unit: Miyako Group.
Geologic age: Aptian, Cretaceous.
Inoceramus balticus var. toyajoanus Nagao andMatumoto, 1940
Plate 14, Figure 4
Jour. Fac. Sci. Hokkaido Imp. Univ., Ser. 4, vol. 6, no. 1,
p. 20, pl. 9, fig. 3. Lectotype: IGPS no. 4540 (designatedby Matsumoto in Takai and Matsumoto, 1961; Existing).
Type locality: Toyazyô, Province of Kii [currently Toyajo,
Aridagawa-cho, Arida-gun, Wakayama Prefecture].
Stratigraphic unit: Toyajo Formation. Geologic age:
Santonian–Camapanian, Cretaceous.
Inoceramus concentricus var. nipponicus Nagao andMatumoto, 1939
Jour. Fac. Sci. Hokkaido Imp. Univ., Ser. 4, vol. 4, nos.
3–4, p. 267, pl. 24, fig. 2, pl. 25, figs. 1–6. Lectotype: IGPS
no. 58017? (designated by Pergament, 1966; Missing).
Type locality: Ugoizawa, a tributary of the Naibuti [sic,Ugui-zawa, Naibuchi] [Ochiai-machi, Sakaehama-gun,
Karafuto Prefecture], south Sakhalin, [currently Nayba
River, Bykov, Dolinsky District, Sakhalin Oblast], Russia.
Stratigraphic unit: Middle Yezo Group. Geologic age:
Cenomanian–Turonian, Cretaceous.

33

Inoceramus pedalionoides Nagao and Matumoto, 1939
ex Inai (MS)Plate 14, Figure 5
Jour. Fac. Sci. Hokkaido Imp. Univ., Ser. 4, vol. 4, nos. 3–4,
p. 277, pl. 26, figs. 8, 9. Syntype: IGPS no. 22720 (Existing).
Type locality: Pombetu, [Mikasayama-mura, Sorachi-gun],
Province of Ishikari [currently Mikasa City], Hokkaido.
Stratigraphic unit: Middle Yezo Group to lowest part of theUpper Yezo Group. Geologic age: Turonian, Cretaceous.
Inoceramus yabei Nagao and Matumoto, 1940
Jour. Fac. Sci. Hokkaido Imp. Univ., Ser. 4, vol. 4, nos. 3–4,
pl. 34, figs. 5–7 [nomen nudum]; Jour. Fac. Sci. Hokkaido
Imp. Univ., Ser. 4, vol. 6, no. 1, p. 1, pl. 1, figs. 1–6.
Lectotype: IGPS no. 22685 (designated by Matsumoto andHarada, 1964; Missing). Type locality: Ikusyunbetu district,
Province of Isikari [currently Ikushumbetsu, Mikasa City],
Hokkaido. Stratigraphic unit: Mikasa Formation. Geologic
age: Cenomanian–Turonian, Cretaceous.
Isognomon (Isognomon) hataii Noda and Furuichi, 1972
Plate 14, Figure 6
Venus, vol. 31, no. 3, p. 120, text-fig. 1. Holotype:
IGPS no. 91766 (Existing). Type locality: W of Abuzaki,[Teshima], Tonosho-cho, Shozu-gun, Kagawa Prefecture.
Stratigraphic unit: Teshima Formation. Geologic age:
Miocene [Late Eocene].
Katelysia (Nipponomarcia) endoi Hatai and Kotaka, 1952
Plate 14, Figure 7
Short Pap. Inst. Geol. Paleont., Tôhoku Univ., no. 4, p.
84, pl. 7, fig. 25. Holotype: IGPS no. 74341 (Existing).
Type locality: Paiponchon, Shinsoruton, San-u-nanmyon,
Myonchon District, Hamukyon-pukuton, Korea [currently
Hwasong-gun, Hamgyongpuk-to, North Korea].
Stratigraphic unit: Heiroku Formation Geologic age: Early
Miocene [latest Early–earliest Middle Miocene]
Leptochondria? hataii Murata, 1973
Plate 14, Figure 8
Sci. Rep. Tohoku Univ., 2nd Ser. (Geol.), Spec. Vol., no.
6 (Prof. K. Hatai Memorial Vol.), p. 273, pl. 29, figs. 1–3.
Holotype: IGPS no. 92661 (Existing). Type locality: Hikado,
Motoyoshi-cho, Motoyoshi-gun [currently Motoyoshicho,
Kesennuma City], Miyagi Prefecture. Stratigraphic unit:
Osawa Formation. Geologic age: Scythian, Triassic.
Leukomoides nipponicus Ogasawara, 1976
Plate 14, Figure 9
Sci. Rep. Tohoku Univ., 2nd Ser. (Geol.), vol. 46, no. 2, p.
52, pl. 11, figs. 3, 4. Holotype: IGPS no. 95190 (Existing).
Type locality: Loc. no. Su-3 (Ogasawara, 1976), small

34

Shin’ichi Sato, Tomoki Chiba, Takaki Yamanaka, Jun Nemoto, Masanori Shimamoto and Takashi Matsubara

road side cutting, about 1000 m W from Futamata-machi,
Kanazawa City, Ishikawa Prefecture. Stratigraphic unit:
Sunakozaka Formation. Geologic age: Miocene [latest Early
Miocene]. Remarks: The present species is a type speciesof Leukomoides Ogasawara, 1976 (by original designation).
Lima amaxensis var. kumasoana Nagao, 1928a
Plate 14, Figures 10a–b
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 9, no.
3, p. 104, pl. 20, figs. 31, 31a, 32. Lectotype: IGPS no.
35808 (designated by Hatai and Nisiyama, 1952; Existing).
Type locality: Amakusa-Shimoshima, precise locality
being unknown (probably cliff behind houses immediately
W of the main road, about 450 m NW of the contact
point of the main road and the road at Imada, Itchodamura, Amakusa-gun [currently Kawauramachi, Amakusa
City], Kumamoto Prefecture: Hatai and Nisiyama, 1952).
Stratigraphic unit: Sakasegawa Formation. Geologic age:
Eocene [Late Eocene].
Lima eocenica Nagao, 1928a
Plate 14, Figure 11
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 9, no.
3, p.105, pl. 21, figs. 1–3. Lectotype: IGPS no. 35817(designated by Hatai and Nisiyama, 1952; Existing). Type
locality: Cliff behind the house immediately W of the mainroad about 450 m NW of the contact point of the mainroad at Imada, Itchoda-mura, Amakusa-Shimoshima,
Amakusa-gun [currently Kawauramachi, Amakusa City],
Kumamoto Prefecture. Stratigraphic unit: Sakasegawa
Formation. Geologic age: Eocene [Late Eocene].
Lima (Limatula) ishidoensis Yabe and Nagao, in Yabe,
Nagao and Shimizu, 1926
Plate 14, Figure 12
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 9, no.
2, p. 60, pl. 14, figs. 7, 16. Lectotype: IGPS no. 22539(designated by Hayami, 1965a; Existing). Type locality:
Ishidô, near Ôhinata[-mura] [currently Ishidô, Ôhinata,
Sakuho-machi], Minamisaku-gun, Province of Shinano[Nagano Prefecture]. Stratigraphic unit: Ishidô Group.
Geologic age: Cretaceous [Early Cretaceous].
Lima (Limea) limopsis Nomura and Zinbô, 1934
Plate 14, Figures 13a–b
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 16,
no. 2, p. 154 (46), pl. 5 (1), figs. 11a–12b. Holotype:
IGPS no. 50368 (Existing). Type locality: Plateaux area
near Kamikatetu, Kikai-zima, [Kikai-cho, Oshima-gun],Kagoshima Prefecture. Lithostratigraphic unit: “Ryûkyû
Limestone”. Geologic age: Pleistocene.

Limatula sakoi Masuda and Katto, in Katto and Masuda,
1978
Plate 14, Figure 14
R es. Rep. Kochi Univ., Nat. Sci., vol. 27, p. 104, pl.
5, fig. 13. Holotype: IGPS no. 96073 (Existing). Type
locality: Loc. no. 9 (Katto and Masuda, 1978), sea coastof Uematsu, Kushimoto-cho, Nishimuro-gun [currently
Higashimuro-gun], Wakayama Prefecture. Stratigraphic
unit: Uematsu Formation. Geologic age: Miocene [latest
Early–early Middle Miocene].
Limipecten bandoi Murata, 1969
Plate 14, Figures 15a–b
Saito Ho-on Kai Mus. Res. Bull., no. 38, p. 18, pl. 4, fig.
1a–c. Holotype: IGPS no. 91384 (Existing). Type locality:
Nagahata [sic, Nakahata], Towa-cho, Tome-gun [currently
Tome City], Miyagi Prefecture. Stratigraphic unit: The lower
part of Toyoma Formation. Geologic age: Late Permian.
Lioconcha tosana Nomura, 1937
Plate 15, Figures 1a–b
Japan. Jour. Geol. Geogr., vol. 14, nos. 3–4, p. 82, pl. 6,
fig. 14a–b. Holotype: IGPS no. 54674 (Existing). Type
locality: Tônohama, Yasuda-mura [sic, Yasuda-cho], Akigun, Kôti [=Kochi] Prefecture. Stratigraphic unit: "Tosa
Pliocene" [=Ananai Formation]. Geologic age: Pliocene[latest Late Pliocene–early Early Pleistocene].
Lithophaga otukai Nomura and Hatai 1936
Saito Ho-on Kai Mus. Res. Bull., no. 10, p. 211, text-figs.
1–3. Holotype: IGPS no. 62437 (fide Hatai and Nisiyama,
1939 [“1938”]; (Missing). Type locality: Southeast valleyof Siratori, Nisatai-mura, Ninohe-gun, Mutu [=Mutsu]
Province [currently Shiratori, Ninohe City, Iwate
Prefecture]. Stratigraphic unit: Lower Kadonosawa Series[=Tate Sandstone Member of Kadonosawa Formation].
Geologic age: Miocene [latest Early Miocene].
Loripes tonohamanus Nomura, 1937
Plate 15, Figures 2a–b
Japan. Jour. Geol. Geogr., vol. 14, nos. 3–4, p. 82, pl.
6, fig. 5a, b. Holotype: IGPS no. 54645 (Existing). Type
locality: Tônohama, Yasuda-mura [sic, Yasuda-cho], Akigun, Kôti [=Kochi] Prefecture. Stratigraphic unit: "Tosa
Pliocene" [=Ananai Formation]. Geologic age: Pliocene[latest Late Pliocene–earliest Early Pleistocene]. Remarks:
Although the name of species was originally spelled as
tonohamana, it should be corrected as tonohamanus
because Loripes is a musculine noun.
Lucina kunigamiensis Nomura and Zinbô, 1936
Plate 15, Figures 3a–b

Catalogue of type specimens of fossil Bivalvia35

 ci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 18,
S
no. 3, p. 241, pl. 11, figs. 10a–11b. Holotype: IGPS no.
51350 (Existing). Type locality: Gabusoga, Haneji-mura,
Kunigami-gun, Okinawa-jima [currently Nago City], Okinawa
Prefecture. Stratigraphic unit: Simaziri Beds [=HanejiFormation]. Geologic age: Early Pliocene [Early Pleistocene].

(Existing; broken). Type locality: Station no. 12 (Nomura,1933), 700 or 800 m NE of Hakushaton, Koryu-sho,
Chikunan-gun, Shinchiku-shu [currently Baishaton,
Baidong Village, Tongsiao Township, Miaoli County],
Taiwan. Lithostratigraphic unit: Byôritsu Beds [Miaoli
Formation]. Geologic age: Pliocene [Pleistocene].

Lucinoma otukai Hatai and Nisiyama, 1949
Plate 15, Figures 4a–b
Jour. Paleont., vol. 23, no. 1, p. 91, pl. 24, fig. 10.
H olotype: IGPS no. 72501 (Existing). Type locality:
F u k u d a , S e k i m o t o - m u r a , Ta g a - g u n [ c u r r e n t l y
Sekimotocho, Kitaibaraki City], Ibaraki Prefecture.
Stratigraphic unit: Kamenoo Formation. Geologic age:
Miocene [Early Miocene]. Remarks: This species is ajunior synonym of Lucinoma kamenooensis (Otuka, 1943).

Macoma hokiensis Akutsu, 1964
Plate 15, Figure 9
Sci. Rep. Tohoku Univ., 2nd Ser. (Geol.), vol. 35, no. 3, p.
287, pl. 60, fig. 8. Holotype: IGPS no. 85512 (Existing).
Type locality: Along the Hôki River, Sekiya, Shiobaramachi, Shioya-gun [currently Nasushiobara City], Tochigi
Prefecture. Stratigraphic unit: Kanomatazawa Formation.
Geologic age: Miocene [Late Miocene]. Remarks: This
species is synonymous with Rexithaerus sectior (Oyama,1950) (Amano et al., 1999).

Lucinopsis? boshihoensis Nomura, 1933
Plate 15, Figures 5a–b
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 16,
no. 1, p. 98, pl. 2, fig. 17a, b. Holotype: IGPS no. 48007(Existing). Type locality: Station no. 13 (Nomura, 1933),
Bôshiho, Shiko-shô, Byôritsu-gun, Shinchiku-shû [currently
Maozihbu, Wuhwu Village, Shihwu Township, Miaoli
County], Taiwan. Lithostratigraphic unit: Byôritsu Beds
[Miaoli Formation]. Geologic age: Pliocene [Pleistocene].
Lucinopsis kosuiensis Nomura, 1933
Plate 15, Figure 6
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 16,
no. 1, p. 99 (no indication). Holotype: IGPS no. 48001(Existing; broken). Type locality: Kôsui, Hokuseikwa,
Tsûshô-shô, Byôritsu-gun, Shinchiku-shû [currently
Peishihwo, Fulong Village, Tongsiao Township, Miaoli
County], Taiwan. Lithostratigraphic unit: Byôritsu Beds
[Miaoli Formation]. Geologic age: Pliocene [Pleistocene].
Macoma (Macoma) atsunaiensis Honda, 1988
Plate 15, Figures 7a–b
S aito Ho-on Kai Spec. Pub., no. 2, (Prof. T. Kotaka
Commem. Vol.), p. 360, pl. 2, figs. 1–6. Holotype: IGPS
no. 97106-2 (Existing). Type locality: Loc. no. AN-20(Honda, 1988), about 4.5 km NNE from Onbetsu-machi,
upper stream of the Shakubetsu-gawa, Onbetsu-machi[sic, Onbetsu-cho], Shiranuka-gun [currently Onbetsucho,
Kushiro City], Kushiro Province, Hokkaido. Stratigraphic
unit: Atsunai Formation. Geologic age: Middle Miocene.
Macoma hakushatonensis Nomura, 1933
Plate 15, Figures 8a–b
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 16,
no. 1, p. 103, pl. 1, fig. 6a, b. Holotype: IGPS no. 48589

Macoma yamadai Nagao, 1928b
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 12,
no. 1, p. 81, pl. 6, fig. 3. Holotype: IGPS no. unknown(Missing). Type locality: Third Namazuta Mine near Iizukamachi, Kaho-gun [currently Iizuka City], Province of Hizen[Fukuoka Prefecture]. Stratigraphic unit: Namazuta Fossil
Bed [=Honso Formation]. Geologic age: Eocene [Middle
Eocene].
Macrocallista ariakensis Nagao, 1928a
Plate 15, Figure 10
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 9, no.
3, p. 112, pl. 18, fig. 16, pl. 20, figs. 26–29. Lectotype:
IGPS no. 35758 (designated by Hatai and Nisiyama,
1952; Existing). Type locality: Akase, Ôda-mura, Utogun, Province of Hizen [currently Akasemachi, Uto
City, Kumamoto Prefecture]. Stratigraphic unit: “Lower
Orthaulax japonicus Zone” [=Fukuregi Formation].
Geologic age: Eocene [early Middle Eocene].
Macrocallista hanzawai Nagao, 1928b
Plate 15, Figure 11
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 12, no.
1, p. 69, pl. 13, fig. 7. Holotype: IGPS no. 36236 (Existing).
Type locality: Meinohama[-machi], Sawara-gun [currently
Nishi-ku, Fukuoka City], Province of Chikuzen [Fukuoka
Prefecture]. Stratigraphic unit: Meinohama Sandstone[Meinohama Formation]. Geologic age: Oligocene.
Macrocallista kahoensis Nagao, 1928b
Plate 15, Figures 12a–b
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 12, no.
1, p. 67, pl. 11, fig. 8, 8a, b. Holotype: IGPS no. 36389(Existing). Type locality: Third Namazuta Mine, near Iizuka-

36

Shin’ichi Sato, Tomoki Chiba, Takaki Yamanaka, Jun Nemoto, Masanori Shimamoto and Takashi Matsubara

machi, Kaho-gun [curently Iizuka City], Province of Chikuzen[Fukuoka Prefecture]. Stratigraphic unit: Namazuta Fossil
Bed [=Honso Formation]. Geologic age: Eocene. Remarks:
Original incorrect spelling kahöensis is corrected.
Macrocallista matsuraensis Nagao, 1928b
Plate 15, Figures 13a–c
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 12,
no. 1, p. 66, pl. 11, figs. 7, 7a, b. Holotype: IGPS no.
36209 (Existing). Type locality: Pass-side cutting on theboundary between Oyama-mura and Arita-machi [currently
between Hiroseyama and Obo in Arita-cho], about 500
m NW of the shrine at Obo, Arita-machi [currently Aritacho], Nishimatsuura-gun, Saga Prefecture. Stratigraphic
unit: Kishima Formation. Geologic age: Oligocene [latest
Eocene–earliest Oligocene].
Mactra (Spisula) asperaeformsis Nomura and Zinbô,
1934
Plate 15, Figures 14a–b
S ci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol.
16, no. 2, p. 156 (48), pl. 5 (1), fig. 20a, b. Holotype:
IGPS no. 50396 (Existing). Type locality: Plateaux area
near Kamikatetu, Kikai-zima, [Kikai-cho, Oshima-gun],Kagoshima Prefecture. Lithostratigraphic unit: “Ryûkyû
Limestone”. Geologic age: Pleistocene.
Mactra nakayamaensis Kamada, 1962
Plate 15, Figures 15a–b
Palaeont. Soc. Japan Spec. Pap. no. 8, p. 123, pl. 14,
figs. 1, 2. Holotype: IGPS no. 79386 (Existing). Type
locality: W of Tatsuzawa, Iino, Taira City [currently Iwaki
City], Fukushima Prefecture. Stratigraphic unit: Nakayama
Formation. Geologic age: Miocene [latest Early Miocene].
Mactra sulcataroides Akutsu, 1964
Plate 15, Figure 16
Sci. Rep. Tohoku Univ., 2nd Ser. (Geol.), vol. 35, no. 3, p.
286, pl. 59, fig. 5. Holotype: IGPS no. 85509 (Existing).
Type locality: Shimotokurazawa, Sekiya, Shiobaramachi, Shioya-gun [currently Nasushiobara City], Tochigi
Prefecture. Stratigraphic unit: Kanomatazawa Formation.
Geologic age: Miocene [Late Miocene].
Margaritifera owadaensis Noda, 1970
Plate 15, Figures 17a–b
Trans. Proc. Palaeont. Soc. Japan, N.S., no. 77, p. 240,
pl. 25, fig. 3a–c. Holotype: IGPS no. 86893 (Existing).
Type locality: Loc. No. 1 (Noda, 1970), exposure along the
National Road side near Owada, Rumoi City, Hokkaido.
Stratigraphic unit: Owada Formation. Geologic age:
Eocene.

Meretrix deguchii Hayasaka and Hayasaka, 1960
Plate 15, Figures 18a–b
Trans. Proc. Palaeont. Soc. Japan, N.S., no. 38, p. 267,
pl. 31, fig. 8a–b. Holotype: IGPS no. 77512 (Existing).
Type locality: Tungyüping in the Penghu Islands,[Dongping Village, Wang'an Township, Penghu County],
Taiwan. Lithostratigraphic unit: “Sandstone bed”. Geologic
age: Pleistocene.
Meretrix meretrix ninohensis Hatai, 1940
Plate 15, Figures 19a–b
Bull. Biogeogr. Soc. Japan, vol. 10, no. 9, p. 128, pl. 1,
fig. 1. Holotype: IGPS no. 61351 (Existing). Type locality:
Yazawa, Nisatai-mura, Ninohe-gun [currently Nisatai,
Ninohe City], Iwate Prefecture. Lithostratigraphic unit:
Shiratori Beds [=Tate Sandstone Member of Kadonosawa
Formation]. Geologic age: Miocene [latest Early Miocene].
Meretrix pseudomeretrix Nagao, 1928b
Plate 15, Figure 20
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 12,
no. 1, p. 65, pl. 9, figs. 7, 7a. Holotype: IGPS no. 36361(Existing). Ogasawara (2001) erronously stated IGPS no.
35381 as the holotype. Type locality: Wakita, Shimagômura, Onga-gun [currently Wakamatsu-ku, Kitakyushu
City], Province of Chikuzen [Fukuoka Prefecture].
Stratigraphic unit: Wakita Beds [=Waita Formation].
Geologic age: Oligocene.
Meretrix tochigiensis Akutsu, 1964
Plate 15, Figures 21a–b
S ci. Rep. Tohoku Univ., 2nd Ser. (Geol.), vol. 35, no.
3, p. 285, pl. 60, figs. 1, 2. Holotype: IGPS no. 85507(Existing). Type locality: Cliff of the Hoki River, about 100
m down stream from Daikoku-iwa, Sekiya, Shiobaramachi, Shioya-gun [currently Nasushiobara City], Tochigi
Prefecture. Stratigraphic unit: Kanomatazawa Formation.
Geologic age: Miocene [Late Miocene].
Miyagipecten matsumoriensis Masuda, 1952
Plate 16, Figures 1a–b
Trans. Proc. Palaeont. Soc. Japan, N.S., no. 8, p. 252,
pl. 24, figs. 4a–7b. Lectotype: IGPS no. 90690 (formerly
DGS 1015; designated by Masuda, 1957; Existing).
Type locality: The foot of the dam of the water reservoir
at Dôgasawa, about 700 m N of Matsumori, Nanakitamura, Miyagi-gun [currently Izumi-ku, Sendai City],
Miyagi Prefecture. Stratigraphic unit: Nanakita Formation.
Geologic age: Middle Miocene [late Late Miocene].
Remarks: This species is a type species of Miyagipecten
Masuda, 1952 (by original designation).

Catalogue of type specimens of fossil Bivalvia
Mizuhopecten kimurai kagaensis Ogasawara, 1976
Plate 16, Figure 2
Sci. Rep. Tohoku Univ., 2nd Ser. (Geol.), vol. 46, no. 2,
p. 43, pl. 14, figs. 10, 17, pl. 15, figs. 8, 9, 12, 16, 18.
Holotype: IGPS no. 95012 (Existing). Type locality: Riverfloor of Sai-kawa, about 1000 m upstream, Omma Bridge[sic, Okuwa Bridge], Omma [sic, Okuwa], Kanazawa
City, Ishikawa Prefecture. Stratigraphic unit: Saikawa
Formation. Geologic age: Miocene [Middle Miocene].
Mizuhopecten paraplebejus murataensis Masuda andTakegawa, 1965
Plate 16, Figures 3a–b
Saito Ho-on Kai Mus. Res. Bull., no. 34, p. 10, pl. 1, figs.
7–10. Holotype: IGPS no. 90826 (formerly DGS no. 4681;
Existing). Type locality: Loc. no. 17 (Masuda and Takegawa,1965), road cliff at Nanamagari Pass, about 250 m SE of thejunction of two main roads at the boundary of Murata-machi
and Kawasaki-machi, Shibata-gun, Miyagi Prefecture.
Stratigraphic unit: Fukuda Member of Kanagase Formation.
Geologic age: Middle Miocene [late Late Miocene].
Modiola? ishidoensis Yabe and Nagao, in Yabe, Nagao
and Shimizu,1926
Plate 16, Figure 4
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 9, no.
2, p. 63, pl. 13, fig. 42. Holotype: IGPS no. 8570 (Existing).
Tanaka and Toshimitsu (2003) erroneously stated IGPS
no. 8750 as the holotype. Type locality: Ishidô, nearÔhinata[-mura] [currently Ishido, Ohinata, Sakuho-machi],Minamisaku-gun, Province of Shinano [Nagano Prefecture].
Stratigraphic unit: Ishidô Group [Ishido Formation].
Geologic age: Cretaceous [late Early Cretaceous].
Modiolus ezoensis Yabe and Nagao, 1928
Plate 16, Figures 5a–b
S ci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 9,
no. 3, p. 89, pl. 17, fig. 11. Holotype: IGPS no. 22627(Existing). Type locality: Washino-sawa, Horomui [currently
Kurisawacho, Iwamizawa City], Ishikari Province,
Hokkaido. Stratigraphic unit: Mikasa Formation. Geologic
age: Cenomanian, Cretaceous.
Modiolus wanizakiensis Masuda, 1966b
Plate 16, Figures 6a–b
Trans. Proc. Palaeont. Soc. Japan, N.S., no. 64, p. 324,
pl. 35, figs. 8a–9. Holotype: IGPS no. 90180 (formerly
DGS no. 4510; Existing). Although Ogasawara (2001)
stated IGPS no. 90876 as the holotype, it is the paratype.
Type locality: Loc. No. 33 (Masuda, 1966a), road cuttingat Wanizaki, Suzu City, Ishikawa Prefecture. Stratigraphic
unit: Higashi-innai Formation. Geologic age: Miocene.

37

Modiolus (Modiolatus) yasuhiroi Kamada, 1962
Plate 16, Figures 7a–b
Palaeont. Soc. Japan Spec. Pap., no. 8, p. 72, pl. 3, figs.
13–14b. Holotype: IGPS no. 79379 (Existing). Type locality:
Numanouchi Harbor, Toyoma-machi, Taira City [currently
Numanouchi, Taira, Iwaki City], Fukushima Prefecture.
Stratigraphic unit: Numanouchi Formation. Geologic age:
Miocene [latest Early–earliest Middle Miocene].
Monia denselineata Hatai, Masuda and Suzuki, 1961
Plate 16, Figures 8a–d
Saito Ho-on Kai Mus. Res. Bull., no. 30, p. 26, pl. 2, figs.
1a–3. Holotype: IGPS no. 90505 (Existing). Type locality:
Loc. no. M 1 (Hatai et al., 1961), left cliff of Maekawa River,near the outlet of Maekawa, Chikagawa [sic, Nakanosawa],
Mutsu City, Aomori Prefecture. Stratigraphic unit: Hamada
Formation. Geologic age: Pliocene [Early Pleistocene].
Mya arenaria kitahukuokaensis Hatai, 1940
Plate 16, Figure 9
B ull. Biogeogr. Soc. Japan, vol. 10, no. 9, p. 131, pl.
1, fig. 9. Holotype: IGPS no. 61363 (Existing). Type
locality: Anaushi, Kitahukuoka-mati [sic, Ishikiridokoromura], Ninohe-gun [currently Ishikiridokoro, Ninohe City],
Iwate Prefecture. Stratigraphic unit: Kadonosawa Beds
[Kadonosawa Formation]. Geologic age: Miocene [earliest
Middle Miocene].
Mya convexa Noda, 1992
Plate 16, Figure 10
Sci. Rep. Tohoku Univ., 2nd Ser. (Geol.), vol. 62, nos. 1–2,
p. 90, pl. 7, figs. 1–9. Holotype: IGPS no. 100877 (Existing).
Type locality: Loc. no. SK11 (Noda, 1992), middle streamof the Chepotsunai River [=Chiebotsunai River], a tributaryof the Kotanbetsu River, Haboro-machi [sic, Kogawa,Tomamae-cho], [Tomamae-gun], Rumoi Province,
Hokkaido. Stratigraphic unit: Lower part of Sankebetsu
Formation. Geologic age: Early Miocene [Late Eocene].
Remarks: The species name is preoccupied by Mya
convexa Wood, 1815 (Matsubara, 2016).
Myadora okadae Hatai and Masuda, 1960
Plate 16, Figures 11a–b
Saito Ho-on Kai Mus. Res. Bull., no. 29, p. 33, figs. 1, 2.
Holotype: IGPS no. 90184 (Existing). Type locality: Road
side cutting south of Takada about 1 km NW of the junctionof the two roads at Moniwa, [Taihaku-ku], Sendai City,
Miyagi Prefecture. Stratigraphic unit: Moniwa Formation.
Geologic age: Early Miocene [early Middle Miocene].
Myadora suzuensis Masuda, 1966b
Plate 17, Figure 1

38

Shin’ichi Sato, Tomoki Chiba, Takaki Yamanaka, Jun Nemoto, Masanori Shimamoto and Takashi Matsubara rans. Proc. Palaeont. Soc. Japan, N.S., no. 64, p. 324, pl.
T35, figs. 10, 11. Holotype: IGPS no. 90776 (formerly DGS
no. 4513; Existing). IGPS no. 90777 cited as the holotype
by Ogasawara (2001) is the paratype. Type locality: Loc.
No. 24 (Masuda, 1966a), river cliff about 500 m E of Fujio,
Suzu City, Ishikawa Prefecture. Stratigraphic unit: Higashiinnai Formation. Geologic age: Early Miocene [latest Early
Miocene].
Myoconcha hamadaensis Yabe and Shimizu, 1927
Plate 17, Figure 2
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 11, no.
2, p. 134, pl. 13, figs. 13–16. Holotype: IGPS no. 35286(Existing). Type locality: Hamada, Rifu-mura [currently
Rifu-cho], Miyagi-gun, Province of Rikuzen [Miyagi
Prefecture]. Stratigraphic unit: Monophyllites Zone in Rifu
Formation. Geologic age: Ladinian, Triassic.
Myophoria japonica Hayasaka, 1925b
Plate 17, Figure 3
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 8, no. 2, p.
16, pl. 8, fig. 14. Lectotype: IGPS no. 22422 (designated by
Hanzawa et al., 1961; Existing). Type locality: Kinshô-zan,
near Akasaka-machi, prov. Mino [currently Akasakacho,
Ogaki City, Gifu Prefecture]. Horizon: Kuro zone [Akasaka
Limestone]. Geologic age: Permian [Middle Permian].
Myophoria subelegans var. tobai Hayasaka, 1924
Plate 17, Figures 4–7
Japan. Jour. Geol. Geogr., vol. 2, no. 4, p. 109, pl. 15, figs.
4–8. Syntypes: IGPS nos. 8403–8407 (Existing). Type
locality: Yahagi-mura, Kesen-gun, prov. Rikuzen [currently
Yahagicho, Rikuzentakata City, Iwate Prefecture].
Stratigraphic unit: Upper Productus-limestone [Lower partof Kanokura Formation]. Geologic age: Lower Permian[Middle Permian]. Remarks: The species name wasoriginally spelled as sub-elegans.

erroneously cited IGPS no. 722503 as the holotype. Type
locality: Stream exposure slightly W of Ogawa, Kawabemura, Ishiki-gun [sic, Iwaki-gun] [currently Kawabemachi,
Iwaki City], Fukushima Prefecture. Stratigraphic unit: Iwaki
Formation. Geologic age: Oligocene [latest Late Eocene–
earliest Early Oligocene].
Nemocardium (Keenaea) yoshidai Masuda andMiyasaka, 1994
Plate 17, Figure 10
Saito Ho-on Kai Mus. Nat. Hist., Res. Bull., no. 62, p. 1, pl.
1, figs. 1a–3b. Holotype: IGPS no. 102544 (Existing). Type
locality: Cliff along a small stream at Takinosawa, Osawagoshuku, Nishisenboku-machi, Senboku-gun [currently
Osawagoshuku, Daisen City], Akita Prefecture. Stratigraphic
unit: Tentokuji Formation. Geologic age: Pliocene.
Neoburmesia iwakiensis Yabe and Sato, 1942
Plate 17, Figure 11
P roc. Imp. Acad., vol. 18, no. 5, p. 251, text-figs. 1–3.
Holotype: IGPS no. 65274 (Existing). Type locality: 1.5 km
W Koike, Kami-mano-mura, Soma-gun [currently Koike,
Kashima-ku, Minamisoma City], Hukushima [Fukushima]
Prefecture. Stratigraphic unit: Torinosu limestone[Nakanosawa Formation in Soma area]. Geologic age: Upper
Jurassic [Kimmeridgian (Upper Jurassic–Lower Cretaceous)].
Nicaniella (Trautscholdia) nagaoi Matsubara, 2016
Plate 17, Figures 12a–b
Bull. Tohoku Univ. Mus., no. 15, p. 107. Holotype: IGPS
no. 66425 (Existing). Type locality: Southern coast ofHiraiga Inlet, Tanohata-mura, Shimohei-gun, Iwate
Prefecture. Stratigraphic unit: Hiraiga Formation. Geologic
age: Early Cretaceous.

Mytilus haboroensis Noda, 1992
Plate 17, Figure 8
Sci. Rep. Tohoku Univ., 2nd Ser. (Geol.), vol. 62, nos.
1–2, p. 61, pl. 3, figs. 12a–13. Holotype: IGPS no. 100727(Existing). Type locality: Loc. no. SS2 (Noda, 1992), upperstream of the Mosetakinai River, Haboro-machi [sic, Sakae,
Shosanbetsu-mura], Tomamae-gun, Rumoi Province,
Hokkaido. Stratigraphic unit: Lower part of Sankebetsu
Formation. Geologic age: Miocene [Late Eocene].

Nipponopagia ommaensis Ogasawara, 1977
S ci. Rep. Tohoku Univ., 2nd Ser. (Geol.), vol. 47, no.
2, p. 126, pl. 15, figs. 4, 6, 8, pl. 16, fig. 3. Holotype:
IGPS no. 95072 (Missing). Type locality: Loc. no. KO-22(Ogasawara, 1977); cliff, distant from the road, 300 m W
of Kaminakamachi, Kanazawa City, Ishikawa Prefecture.
Stratigraphic unit: Omma Formation. Geologic age:
Pliocene [Early Pleistocene]. Remarks: The present
species is a type species of Nipponopagia Ogasawara,
1977 (by original designation). However, it is thought to bea junior synonym of Macoma (Macoma) middendorffi Dall,
1884 (Petrov, 1982; Amano, 1996).

Mytilus ogawaensis Hatai and Nisiyama, 1949
Plate 17, Figure 9
J our. Paleont., vol. 23, no. 1, p. 89, pl. 24, fig. 17.
Holotype: IGPS no. 72503 (Existing). Ogasawara (2001)

Noetia nagaoi MacNeil, 1938
Plate 17, Figure 13
U.S. Geol. Surv., Prof. Pap., 189-A, p. 30, pl. 4, figs. 19–21.
Holotype: IGPS no. 36012 (Existing). Type locality: The

Catalogue of type specimens of fossil Bivalvia
Hoshuyama Mine (about 200 m S of the bridge at E of
Kawamagari, and about 600 m W of the village office) at
Daigyoji, Hoshuyama-mura [currently Toho-mura], Asakuragun, Fukuoka Prefecture. Stratigraphic unit: Doshi Formation.
Geologic age: Upper Eocene. Remarks: Proposed as nom.
nov. for Arca (Noetia) pondaungensis var. transversa
Nagao, 1928b, non Arca transversa Say, 1822.
Nucula fukasawaensis Akutsu, 1964
Plate 17, Figure 14
Sci. Rep. Tohoku Univ., 2nd Ser. (Geol.), vol. 35, no. 3, p.
282, pl. 59, fig. 1. Holotype: IGPS no. 85500 (Existing). Type
locality: Fukasawa, about 1.8 km upstream from Kanezawa,
Shiobara-machi, Shioya-gun [currently Nasushiobara City],
Tochigi Prefecture. Stratigraphic unit: Kanomatazawa
Formation. Geologic age: Miocene [Late Miocene].
Nucula hizenensis Nagao, 1928b
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 12,
no. 1, p. 19, pl. 7, figs. 18, 18a. Holotype: IGPS no.
35985 (Missing). Type locality: Okino-shima, Iôjima-mura,Nishisonogi-gun [currently Iojimamachi, Nagasaki City],
Province of Hizen [Nagasaki Prefecture]. Stratigraphic
unit: Okino-shima Beds [Okinoshima Formation]. Geologic
age: Eocene [late Middle Eocene].
Nucula (Acila) hokkaidoensis Nagao, 1932
Jour. Fac. Sci., Hokkaido Imp. Univ., Ser. 4, vol. 2, no.
1, p. 28, pl. 5, figs. 17–18. Holotype: IGPS no. 6421
(Missing). Type locality: A point about 100 m S of thejunction of the Abeshinai with its tributary Sakai River[currenly Nakagawa-cho, Nakagawa-gun], Teshio
Province, Hokkaido. Stratigraphic unit: Upper Ammonite
Beds [Middle Yezo Group and Upper Yezo Group in
Teshio area]. Geologic age: Upper Cretaceous [Turonian–Campanian [Santonian], Cretaceous].
Nucula ishidoensis Yabe and Nagao, in Yabe, Nagao andShimizu, 1926
Plate 17, Figures 15a–b
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 9, no.
2, p. 41, pl. 13, figs. 46, 47. Syntypes: IGPS no. 7125
(Existing). Type locality: Ishidô, near Ôhinata [currently
Ishidô, Ôhinata, Sakuho-machi, Minamisaku-gun, Provinceof Shinano [Nagano Prefecture]. Stratigraphic unit: IshidôGroup [Ishidô Formation]. Geologic age: Cretaceous [Early
Cretaceous (late Neocomian)].
Nucula karatsuensis Nagao, 1928b
Plate 17, Figure 16
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 12, no.
1, p. 20, pl. 7, fig. 17. Holotype: IGPS no. 35988 (Existing).

39

Type locality: Obô near Arita-machi, Nishimatsura-gun [sic,Nishimatsuura-gun], Province of Hizen [Saga Prefecture].
Stratigraphic unit: Kishima Beds [Kishima Formation].
Geologic age: Oligocene [latest Eocene–earliest Oligocene].
Nucula kokozuraensis Hatai and Nisiyama, 1949
Plate 17, Figures 17a–b
J our. Paleont., vol. 23, no. 1, p. 87, pl. 25, figs. 1–5.
Holotype: IGPS no. 72507 (Existing). Type locality: A
small cliff in front of the grocery store at the northern
entrance of Kokozura, Nakoso-machi, Ishiki-gun [sic,Iwaki-gun] [currently Nakosomachi, Iwaki City], Fukushima
Prefecture. Stratigraphic unit: Kokozura Formation.
Geologic age: Miocene [latest Early Miocene].
Nucula (Acila) mirabilis var. ashiyaensis Nagao, 1928b
Plate 17, Figures 18a–c
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 12,
no. 1, p. 21, pl. 7, figs. 6–7 , 10 [non figs. 8, 9; =Acila(Truncacila) nagaoi Oyama and Mizuno, 1958]. Holotype:
IGPS no. 35981 (Existing). Type locality: Taya, Ashiyamachi, Onga-gun, Province of Chikuzen [Fukuoka
Prefecture]. Stratigraphic unit: Yamaga Beds [Yamaga
Formation]. Geologic age: Oligocene.
Nucula (Ennucula) omagariensis Honda, 1989
Plate 17, Figure 19
Sci. Rep. Tohoku Univ., 2nd Ser. (Geol.), vol. 60, no. 1, p.
58, pl. 1, figs. 5, 7–9, 11, 12. Holotype: IGPS no. 955761 (Existing). Type locality: OM-6 (Honda, 1989), roadsidecliff along the Taron-no-sawa, a tributary of the Rushingawa, Urahoro-machi [sic, Urahoro-cho], Tokachi-gun,
Hokkaido. Stratigraphic unit: Omagari Formation. Geologic
age: Oligocene.
Nucula (Nucula) shiroyamaensis Noda, 1962b
Sci. Rep. Tohoku Univ., 2nd Ser. (Geol.), vol. 34, no. 3,
p. 226, pl. 16, fig. 7. Holotype: IGPS no. 79061 (Missing).
Type locality: Loc. No. 480 (Noda, 1962b), upstream of
the Shirono-sawa, a tributary from the Shibumi River,facing the northern slope of the Uragawara-mura [sic,Matsudai-machi], Higashikubiki-gun [currently Murono,
Tokamachi City], Niigata Prefecture. Stratigraphic unit:
Nitta Formation. Geologic age: Early Pliocene.
Nuculana (Nuculana) karihaensis Hatai and Nisiyama, 1949
Plate 17, Figures 20a–c
J our. Paleont., vol. 23, no. 1, p. 87, pl. 24, figs. 3, 4.
Holotype: IGPS no. 72633 (Existing). Type locality: Bridge
leading to Takayanagi, Ishiguro-mura, Kariha-gun [sic,Kariwa-gun] [currently Ishiguro, Takayanagicho, Kashiwa
zaki City], Niigata Prefecture. Stratigraphic unit: Ishiguro40Shin’ichi Sato, Tomoki Chiba, Takaki Yamanaka, Jun Nemoto, Masanori Shimamoto and Takashi Matsubara

Formation. Geologic age: Miocene [Late Miocene].
Nuculana (Nuculana) matsukuchiensis Noda, 1962b
Plate 17, Figure 21
S ci. Rep. Tohoku Univ., 2nd Ser. (Geol.), vol. 34, no.
3, p. 227, pl. 16, figs. 4a–5. Holotype: IGPS no. 79059(Existing). Type locality: Loc. No. 80 (Noda, 1962b),
Bed of the Esudo River, about 400 m N of the bridge
between Mioke and Matsukuchi, Matsunoyama-machi,
Higashikubiki-gun [currently Matsunoyama, Tokamachi
City], Niigata Prefecture. Stratigraphic unit: Nitta
Formation. Geologic age: Early Pliocene.
Nuculana sanchuensis Yabe and Nagao, in Yabe, Nagao
and Shimizu, 1926
Plate 17, Figures 22–24
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 9, no.
2, p. 42, pl. 12, figs. 21–23. Syntypes: IGPS no. 7115
(Existing). Type locality: Ishidô, near Ôhinata [currently
Ishidô, Ôhinata, Sakuho-machi, Minamisaku-gun],
Province of Shinano [Nagano Prefecture]. Stratigraphic
unit: Ishidô Group [Ishidô Formation]. Geologic age:
Cretaceous [Early Cretaceous].
Nuculites kimurai Hayasaka, 1925a
Plate 17, Figures 25a–c
Japan. Jour. Geol. Geogr., vol. 3, no. 2, p. 49, pl. 6, figs. 4–7.
Holotype: IGPS no. 22418 (Existing). Type locality: Myôjinyama, Ogachi, Jyûgohama-mura, Monô-gun, prov. Rikuzen[currently Ogatsucho, Ishinomaki City, Miyagi Prefecture].
Stratigraphic unit: Ogachi slate [Toyoma Formation].
Geologic age: Upper Paleozoic [Upper Permian].
Nuttallia uchigoensis Kamada, 1962
Plate 17, Figures 26a–b
Palaeont. Soc. Japan Spec. Pap., no. 8, p. 127, pl. 14,
fig. 4a, b. Holotype: IGPS no. 79387 (Existing). Type
locality: Goten, Uchigo City [currently Uchigo, Iwaki
City], Fukushima Prefecture. Stratigraphic unit: Asagai
Formation. Geologic age: Oligocene [Early Oligocene].
Ostrea cassis Nagao, 1928b
Plate 18, Figure 1
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 12, no.
1, p. 32, pl. 5, figs. 6, 6a, b. Holotype: IGPS no. 36031(Existing). Type locality: Kôyagi-jima, [Kôyagi-mura,Nishisonogi-gun] [currently Koyagimachi, Nagasaki City],
Province of Hizen [Nagasaki Prefecture]. Stratigraphic
unit: Okinoshima Beds [Okinoshima Formation]. Geologic
age: Eocene [Middle Eocene].

Ostrea kahoensis Nagao, 1928b
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 12, no. 1, p.
33, pl. 6, fig. 2. Holotype: IGPS unregistered (Missing). Type
locality: The Tadakuma Mine near Iizuka-machi, Provinceof Chikuzen [currently Iizuka City, Fukuoka Prefecture].
Stratigraphic unit: Namazuta Fossil Beds [=Honso
Formation]. Geologic age: Eocene [Middle Eocene].
Ostrea lunaeformis Nagao, 1928b
Plate 18, Figure 2
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 12,
no. 1, p. 35, pl. 4, fig. 28. Holotype: IGPS no. 36039(Existing). Type locality: Yanagi, Taira-mura, Nishisonogigun [currently Taira, Oosetocho, Saikai City], Province ofHizen [Nagasaki Prefecture]. Stratigraphic unit: Kakinoura
Beds [Kakinoura Formation]. Geologic age: Oligocene[latest Late Eocene–earliest Early Oligocene].
Ostrea sakitoensis Nagao, 1928b
Plate 18, Figures 3a–b
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 12, no.
1, p. 34, pl. 5, figs. 17, 17a. Holotype: IGPS no. 36053(Existing). Type locality: Kaminoura-jima [sic, Kakinourashima], Province of Hizen [currently Kakinourago,
Sakitocho, Saikai City, Nagasaki Prefecture]. Stratigraphic
unit: Itanoura Beds [Itanoura Formation]. Geologic age:
Oligocene [latest Late Eocene–earliest Early Oligocene].
Ostrea (Ostrea) yokoyamai Kamada, 1962
Plate 18, Figure 4
Palaeont. Soc. Japan Spec. Pap., no. 8, p. 68, pl. 4, fig.
1. Holotype: IGPS no. 79378 (Existing). Type locality:
In the Yumoto-Gokô, Joban Coal-mine, Yumotomachi,
Joban City [currently Joban-Yumotomachi, Iwaki City],
Fukushima Prefecture. Stratigraphic unit: Iwaki Formation.
Geologic age: Oligocene [latest Late Eocene–earliest
Early Oligocene]. Remarks: This species is considered
as a junior synonym of Ostrea eorivularis Oyama andMizuno, 1958 (Honda, 1989).
Oxyperas okinawaensis Masuda and Sato, 1988
S aito Ho-on Kai Spec. Pub., no. 2 (Prof. T. Kotaka
Commem. Vol.), p. 446, pl. 4, figs. 6–14. Holotype: IGPS
no. 99715 (Missing). Type locality: Sea cliff at Nishizaki,
about 1 km NW of Nakano, Taketomi-cho, [Yaeyama-gun],Iriomote-jima, Okinawa Prefecture. Stratigraphic unit:
Iriomote Formation. Geologic age: early Middle Miocene[latest Early –early Middle Miocene].
Palaeoneilo ogachiensis Hayasaka, 1925aJapan. Jour. Geol. Geogr., vol. 3, no. 2, p. 51, pl. 6, figs.
8–12. Syntypes: IGPS unregistered (Missing). Type

Catalogue of type specimens of fossil Bivalvia
locality: Myôjin-yama, Ogachi, Jyûgohama-mura, Monôgun, prov. Rikuzen [currently Ogatsucho, Ishinomaki
City, Miyagi Prefecture]. Stratigraphic unit: Ogachi slate[Toyoma Formation]. Geologic age: Upper Paleozoic[Upper Permian].
Panomya izumo Nomura and Hatai, 1939 [“1938”]
Plate 18, Figures 5a–b
Japan. Jour. Geol. Geogr., vol. 16, nos. 1–2, p. 6, pl. 1, fig.
2a, b. Holotype: IGPS no. 51884 (Existing). Type locality:
Husina [sic, Fujina], Tamanoyu-mura [sic, Tamayumura], Yatuka-gun [=Yatsuka-gun], Izumo [currently
Fujina, Tamayumachi, Matsue City, Shimane Prefecture].
Stratigraphic unit: Izumo Neogene [=Fujina Formation].
Geologic age: Middle Miocene.
Panope kanomatazawaensis Akutsu, 1964
Plate 18, Figure 6
S ci. Rep. Tohoku Univ., 2nd Ser. (Geol.), vol. 35, no.
3, p. 288, pl. 60, figs. 3a–5. Holotype: IGPS no. 85513(Existing). Type locality: A tributary of the Kanomatazawa
valley, cliff of the Hoki River, opposite side of the
Hokigawa Electric Power Station, Sekiya, Shiobaramachi, Shioya-gun [currently Nasushiobara City], Tochigi
Prefecture. Stratigraphic unit: Kanomatazawa Formation.
Geologic age: Miocene [Late Miocene].
Panope kanomatazawaensis fudozawaensis Akutsu, 1964
Plate 18, Figure 7
Sci. Rep. Tohoku Univ., 2nd Ser. (Geol.), vol. 35, no. 3, p.
288, pl. 60, fig. 7. Holotype: IGPS no. 85514 (Existing).
Type locality: Fudozawa, about 2 km upstream from theentrance of the valley, Fukuwata, Shiobara-machi, Shioyagun [currently Nasushiobara City], Tochigi Prefecture.
Stratigraphic unit: Kanomatazawa Formation. Geologic
age: Miocene [Late Miocene].
Panopea nomurae Kamada, 1962
Plate 18, Figure 8
Palaeont. Soc. Japan Spec. Pap., no. 8, p. 135, pl. 16,
figs. 9a–12. Holotype: IGPS no. 79395 (Existing). Type
locality: Numanouchi Harbor, Toyoma-machi, Taira City
[currently Numanouchi, Taira, Iwaki City], Fukushima
Prefecture. Stratigraphic unit: Numanouchi Formation.
Geologic age: Miocene [latest Early–earliest Middle
Miocene]. Remarks: The present species is a juniorsynonym of Panopea tyugokuensis (Otuka, 1941)(Matsubara, 2011).
Paphia (Paphia) euglypta ohiroi Masuda, 1966b
Plate 18, Figure 9
Trans. Proc. Palaeont. Soc. Japan, N.S., no. 64, p. 326,

41

pl. 35, figs. 22, 23. Holotype: IGPS no. 90735 (formerly
DGS no. 4527; Existing). Ogasawara (2001) erroneously
cited IGPS no. 90736 as the holotype. Type locality: Loc.
no. 38 (Masuda, 1966a), small exposure near the stream,
about 300 m S of Sotonoyama [currently Sotoyama,Otanimachi], Suzu City, Ishikawa Prefecture. Stratigraphic
unit: Higashi-innai Formation. Geologic age: Early
Miocene [latest Early Miocene].
Paphia shimotsukensis Akutsu, 1964
Plate 18, Figure 10
Sci. Rep. Tohoku Univ., 2nd Ser. (Geol.), vol. 35, no. 3, p.
286, pl. 59, fig. 3. Holotype: IGPS no. 85508 (Existing).
Type locality: Cliff of the Hoki River, Sekiya, Shiobaramachi, Shioya-gun [currently Nasushiobara City], Tochigi
Prefecture. Stratigraphic unit: Kanomatazawa Formation.
Geologic age: Miocene [Late Miocene].
Paphia (Paphia) suzuensis Masuda, 1966b
Plate 18, Figure 11
Trans. Proc. Palaeont. Soc. Japan, N.S., no. 64, p. 327, pl.
35, figs. 24, 25. Holotype: IGPS no. 90086 (formerly DGS
no. 4530; Existing). Type locality: Loc. no. 30 (Masuda,
1966a), road side cutting near Koeiji Temple, Otani[machi],Suzu City, Ishikawa Prefecture. Stratigraphic unit: Higashiinnai Formation. Geologic age: Early Miocene [latest Early
Miocene].
Parallelodon obsoletiformis Hayasaka, 1925b
Plate 18, Figures 12a–b
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 8, no. 2,
p. 18, pl. 8, figs. 15, 16, 16a. Lectotype: IGPS no. 22392(designated by Hanzawa et al., 1961; Existing). Type
locality: Kinshô-zan, Akasaka-machi, prov. Mino [currently
Akasakacho, Ogaki City, Gifu Prefecture]. Stratigraphic
unit: Kuro zone [Akasaka Limestone]. Geologic age:
Permian [Middle Permian].
Parapholas satoi Nagao, 1928b
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 12, no.
1, p. 88, pl. 6, figs. 6, 6a, 7. Holotype: IGPS unregistered(Missing). Type locality: Meinohama, Meinohama-machi,
Sawara-gun [currently Nishi-ku, Fukuoka City], Provinceof Chikuzen [Fukuoka Prefecture]. Stratigraphic unit:
Meinohama Beds [Meinohama Formation]. Geologic age:
Oligocene.
Patinopecten ibaragiensis Masuda, 1953b
Plate 19, Figure 1
Short Pap. Inst. Geol. Paleont. Tohôku Univ., no. 5, p.
44, pl. 5, figs. 1–5, pl. 6, figs. 1–5. Holotype: IGPS no.
90698 (formerly DGS no. 1031; Existing). Type locality:

42

Shin’ichi Sato, Tomoki Chiba, Takaki Yamanaka, Jun Nemoto, Masanori Shimamoto and Takashi Matsubara

Cliff below the Sukegawa Gas Company, [Higashicho],
Hitachi City, Ibaraki Prefecture. Stratigraphic unit: Hitachi
Formation. Geologic age: Late Miocene [Early Pliocene].
Remarks: This species is synonymous with Mizuhopecten
planicostulatus (Nomura and Niino, 1932) (Tanaka andAmano, 1997).
Patinopecten kagamianus moniwaensis Masuda, 1958b
Plate 19, Figure 2
Trans. Proc. Palaeont. Soc. Japan, N.S., no. 32, p. 276,
pl. 41, figs. 3–6b. Holotype: IGPS no. 90700 (formerly
DGS no. 3506; Existing). Type locality: Moniwa, [Taihakuku], Sendai City, Miyagi Prefecture. Stratigraphic unit:
Moniwa Formation. Geologic age: Early Miocene [early
Middle Miocene].
Patinopecten kimurai nakosoensis Masuda, 1960
Plate 19, Figure 3
Trans. Proc. Palaeont. Soc. Japan, N.S., no. 38, p. 256, pl.
29, figs. 2, 3. Holotype: IGPS no. 90581 (formerly DGS no.
1930; Existing). Type locality: Road-side cliff at Kokozura,
Nakoso City [currently Nakosomachi, Iwaki City], Fukushima
Prefecture. Stratigraphic unit: Kokozura Formation. Geologic
age: Middle Miocene [latest Early Miocene].
Patinopecten kimurai yudaensis Masuda, 1960
Plate 19, Figure 4
Trans. Proc. Palaeont. Soc. Japan, N.S., no. 38, p. 256,
pl. 30, figs. 1, 2. Holotype: IGPS no. 90661 (formerly
DGS no. 1936; Existing). Type locality: Right river sideof Mabechi-gawa at Yuda, Kintaichi-mura, Ninohegun [currently Kintaichi, Ninohe City], Iwate Prefecture.
Stratigraphic unit: Shiratori Formation [=Tate Sandstone
Member of Kadonosawa Formation]. Geologic age: Early
Miocene [latest Early Miocene].
Patinopecten kobiyamai Kamada, 1954
Plate 19, Figures 5a–b
Trans. Proc. Palaeont. Soc. Japan, N.S., no. 15, p. 174, pl.
23, figs. 1a–3b. Holotype: IGPS no. 72963 (Existing). Type
locality: IGPS loc. no. Fs-23, about 500 m E of Nakayama,
Ono-mura, Iwaki-gun [currently Yotsukuramachi, Iwaki
City], Fukushima Prefecture. Stratigraphic unit: Kabeya
Formation [=Honya Formation]. Geologic age: Miocene[late Early Miocene].
Patinopecten (Kotorapecten) naganoensis Masuda, 1962aPlate 19, Figure 6; Plate 20, Figure 1
S ci. Rep. Tohoku Univ., 2nd Ser. (Geol.), vol. 33, no.
2, p. 219, pl. 25, fig. 1a, b. Holotype: IGPS no. 90547(formerly DGS no. 3888; Existing). Type locality: Floor of asmall tributary of the Dojiri River, Shiroshita [sic, Joshita],

Nakajo-mura, Kamiminochi-gun [currently Nakajo, Nagano
City], Nagano Prefecture. Stratigraphic unit: Shigarami
Formation. Geologic age: Late Miocene [Early Pliocene].
Patinopecten nakajimai Masuda, 1954b
Plate 20, Figures 2a–b
Trans. Proc. Palaeont. Soc. Japan, N.S., no. 15, p. 159,
pl. 21, figs. 1a–5. Holotype: IGPS no. 90580 (formerly
DGS no. 1063; Existing). Ogasawara (2001) erroneously
cited the holotype as IGPS no. 95080. Type locality:
River cliff of Natori-gawa near the Akiu Car-line Station at
Kita-Akaishi, Oide-mura, Natori-gun [currently Moniwa,
Taihaku-ku, Sendai City], Miyagi Prefecture. Stratigraphic
unit: Moniwa Formation. Geologic age: Early Miocene[early Middle Miocene]. Remarks: This species is ajunior synonym of Nipponopecten akihoensis Saga, in
Matsumoto, 1930 (Sato, 1991; Matsubara et al., 2014).
Patinopecten poculum tsudae Noda, 1962b
Plate 20, Figures 3a–b
Sci. Rep. Tohoku Univ., 2nd Ser. (Geol.), vol. 34, no. 3, p.
228, pl. 16, figs. 1–3. Holotype: IGPS no. 79062 (Existing).
Type locality: Loc. No. 1061-c (Noda, 1962b), cliff of theHigashi River, about 250 m S of a junction with Etsudo
and Higashi Rivers, Matsunoyama-machi, Higashikubikigun [Matsunoyama, Tokamachi City], Niigata Prefecture.
Stratigraphic unit: Higashigawa Formation. Geologic age:
Middle Pliocene.
Patinopecten (Patinopecten) tokyoensis hokurikuensis
Masuda, 1962aPlate 20, Figure 4
Sci. Rep. Tohoku Univ., 2nd Ser. (Geol.), vol. 33, no. 2,
p. 211, pl. 25, fig. 6, pl. 27, figs. 6, 7. Holotype: IGPS no.
13430 (Existing). Type locality: Road-side cliff near the
Sai River, about 400 m SE of the contact point of the tworoads at Onma-machi [currently Okuwamachi], Kanazawa
City, Ishikawa Prefecture. Stratigraphic unit: Onma [Omma]
Formation. Geologic age: Pliocene [ealry Pleistocene].
Remarks: This subspecies is a junior synonym ofPatinopecten (Patinopecten) tokyoensis hokurikuensis
Akiyama, 1962, as noted by Masuda (1962a: p. 238).
Patinopecten yamasakii kintaichiensis Masuda, 1958cPlate 20, Figure 5
Saito Ho-on Kai Mus. Res. Bull., no. 27, p. 47, pl. 4, figs.
3a–4. Holotype: IGPS no. 90715 (formerly DGS no. 3610;
Existing). Type locality: Entrance to the eastern valley
running parallel with the Kamiyama-zawa, Kintaichimura, Ninohe-gun [currently Kintaichi, Ninohe City],
Iwate Prefecture. Stratigraphic unit: Suenomatsuyama
Formation [Tomesaki Formation]. Geologic age:

Catalogue of type specimens of fossil Bivalvia
Late Miocene [early Middle Miocene]. Remarks: The
present species is a junior synonym of Masudapecten
iwasakiensis (Nomura, 1935) (Masuda et al., 1994).
Patinopecten yamasakii ninohensis Masuda, 1954a
Plate 20, Figure 6
Saito Ho-on Kai Mus. Res. Bull., no. 23, p. 13, figs. 1a–3.
Holotype: IGPS no. 90684 (formerly DGS no. 2532;
Existing). Type locality: River floor of Shiratorizawa [sic,Shiratorigawa] in Fukuoka-machi, Ninohe-gun [currently
Fukuoka, Ninohe City], Iwate Prefecture. Stratigraphic
unit: Suenomatsuyama Formation. Geologic age: Late
Miocene [earliest Middle Miocene]. Remarks: IGPS no.
90683 given by Ogasawara (2001) is incorrect.
Patinopecten (Patinopecten) yessoensis
nakatonbetsuensis Masuda, 1962aPlate 20, Figure 7
Sci. Rep. Tohoku Univ., 2nd. Ser. (Geol.), vol. 33, no.
2, p. 214, pl. 24, fig. 4. Holotype: IGPS no. 90662(formerly DGS no. 3899; Existing). Type locality:
Asahidai, Nakatonbetsu-machi [sic, Nakatonbetsu-cho],Esashi-gun, Kitami Province, Hokkaido. Stratigraphic
unit: Nakatonbetsu Formation. Geologic age: Late
Miocene [Middle Miocene]. Remarks: This taxon isa junior synonym of Patinopecten (Patinopecten)
nakatombetsuensis Akiyama, 1962 ex Masuda (MS),as noted by Masuda (1962a: p. 238).
Patinopecten (Patinopecten) yessoensis yokoyamae
Masuda, 1962aPlate 21, Figure 1
Sci. Rep. Tohoku Univ., 2nd Ser. (Geol.), vol. 33, no. 2, p.
215, pl. 25, figs. 2, 3, pl. 26, fig. 7, pl. 27, fig. 12. Holotype:
IGPS no. 90543 (formerly DGS 3836; Existing). Type locality:
Left cliff of the Chikagawa River, about 200 m from the sea
shore at Chikagawa [sic, Nakachikagawa, Nakanosawa],
Mutsu City, Aomori Prefecture. Stratigraphic unit: Hamada
Formation. Geologic age: Pliocene [Ealry Pleistocene].
Pecten (Pseudamusium) akihoensis Saga, in
Matsumoto, 1930
Plate 21, Figures 2a–b
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 13,
no. 3, p. 106, pl. 40, figs. 7, 8. Neotype: IGPS no. 90586(formerly DGS 1001, designated by Masuda, 1952;
Existing). Matsumoto’s (1930) syntypes are missing(Masuda, 1952). Type locality: 100 m W of the Kumanodoshrine, Takadate-mura [currently Takadate-Kumanodo,
Natori City], Miyagi Prefecture. Stratigraphic unit: Moniwa
Formation. Geologic age: Late Miocene [early Middle
Miocene]. Remarks: Originally spelled as akihoënsis

43(Matsumoto, 1930).
Pecten (Chlamys) ashiyaensis Nagao, 1928b
Plate 21, Figure 3
S ci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol.
12, no. 1, p. 39, pl. 9, figs. 2, 2a. Holotype: IGPS no.
36448 (Existing). Type locality: Yamaga, Ashiya-machi,
Onga-gun, Province of Chikuzen [Fukuoka Prefecture].
Stratigraphic unit: Yamaga Formation. Geologic age:
Oligocene [latest Early–earliest Late Oligocene].
Pecten (Chlamys) ashiyaensis var. denselineatus Nagao,
1928b
Plate 21, Figure 4
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 12, no.
1, p. 40, pl. 8, fig. 9. Holotype: IGPS no. 36437 (Existing).
Type locality: Kaminoura-jima, [Sakito-mura, Nishisonogigun], Province of Hizen [currently Kakinourago, Sakitocho,
Saikai City, Nagasaki Prefecture]. Stratigraphic unit:
Itanoura Beds [Itanoura Formation]. Geologic age:
Oligocene [latest Late Eocene–earliest Early Oligocene].
Pecten (Vola) byoritsuensis Nomura, 1933
Plate 21, Figure 5
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 16,
no. 1, p. 57, pl. 1, figs. 4a–5b. Holotype: IGPS no. 45001(Existing). Type locality: The upper course of Sairyôkyô,Sanchin-shô, Tainan-shu [currently Yseiliao Bridge,
Yseiliao, Ronghe Village, Tsuochen District, Tainan City],
Taiwan. Lithostratigraphic unit: Byôritsu Beds [Miaoli
Formation]. Geologic age: Pliocene [Pleistocene].
Pecten (Propeamussium) cowperi var. yubarensis Yabe
and Nagao, 1928
Plate 21, Figure 6
Sci. Rep. Tôhoku Imp. Univ., vol. 9, no. 3, p. 88, pl. 16,
figs. 17–19. Holotype: IGPS no. 22599 (Existing). Type
locality: Pankemo-yubari, Oyubari, Yubari-gun [currently
Pankemoyuparo River, Kashima-shirogane, Yubari City],
Hokkaido. Stratigraphic unit: Middle Yezo Group andUpper Yezo Group in various areas of Hokkaido. Geologic
age: Cenomanian–Campanian, Cretaceous.
Pecten (Aequipecten) gabusogaensis Nomura andZinbô, 1936
Plate 21, Figure 7
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 18, no. 3, p.
236, pl. 11, figs. 3a–4b. Holotype: IGPS no. 51321 (Existing).
Type locality: Gabusoga, Haneji-mura, Kunigami-gun[currently Nago City], Okinawa-jima, Okinawa Prefecture.
Stratigraphic unit: Simaziri Beds [=Haneji Formation].
Geologic age: Early Pliocene [Early Pleistocene].

44

Shin’ichi Sato, Tomoki Chiba, Takaki Yamanaka, Jun Nemoto, Masanori Shimamoto and Takashi Matsubara

Pecten (Decadopecten) izuensis Nomura and Niino, 1932
Plate 21, Figures 8–12
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 15, no.
3, p. 179, pl. 11, figs. 9–13. Syntypes: IGPS no. 44583(Existing). Type locality: Nagata, Shirahama-mura, Kamogun, Province of Izu [currently Shirahama, Shimoda
City, Shizuoka Prefecture]. Stratigraphic unit: Shrahama
Shell-Beds of Shirahama Group [Shirahama Formation].
Geologic age: Miocene [Pliocene].
Pecten (Chlamys) kagamianus miyagiensis Nakamura,
1940a
Plate 21, Figure 13
J apan. Jour. Geol. Geogr., vol. 17, nos. 1–2, p. 10,
pl. 2, fig. 4. Holotype: IGPS no. 61336 (Existing).
Type locality: Ôtutumi [=Otsutsumi], Miyatoko-mura[currently Miyatoko, Taiwa-cho], Kurokawa-gun, Miyagi
Prefecture. Stratigraphic unit: Ôtutumi Beds [=Otsutsumi
Formation]. Geologic age: Miocene [early Late Miocene].
Remarks: The present subspecies is a junior synonymof Nanaochlamys notoensis otutumiensis (Nomura andHatai, 1937) (e.g. Masuda, 1962a; Sato, 1991).
Pecten (Pecten) kakisakiensis Nomura and Niino, 1932
Plate 21, Figures 14–15
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 15,
no. 3, p. 177, pl. 12, figs. 3, 4. Syntypes: IGPS no. 44582(Existing). Type locality: Kakisaki, Hamasaki-mura [sic,Hamazaki-mura], Kamo-gun, Province of Izu [currently
Kakisaki, Shimoda City, Shizuoka Prefecture]. Stratigraphic
unit: Shirahama Shell-Beds of Shirahama Group [Shirahama
Formation]. Geologic age: Miocene [Pliocene]. Remarks:
The present species is synonymous with Mimachlamys
satoi (Yokoyama, 1928) (e.g. Masuda, 1962a).
Pecten (Aequipecten) kikaiensis Nomura and Zinbô, 1934
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 16,
no. 2, p. 153 (45), pl. 5 (1), fig. 9a, b. Holotype: IGPS
no. 50357? (Missing). Type locality: Plateaux area
near Kamikatetu, Kikai-zima, [Kikai-cho, Oshima-gun],Kagoshima Prefecture. Lithostratigraphic unit: “Ryûkyû
Limestone”. Geologic age: Pleistocene.
Pecten (Patinopecten) kimurai matumoriensis
Nakamura, 1940a
Plate 22, Figures 1a–b
Japan. Jour. Geol. Geogr., vol. 17, nos. 1–2, p. 13, pl.
1, figs. 1, 2, pl. 2, figs. 1–3. Syntypes: IGPS no. 61335(Existing). Type locality: Matumori [=Matsumori], Nanakitamura, Miyagi-gun [currently Matsumori, Izumi-ku, Sendai
City], Miyagi Prefecture. Stratigraphic unit: Nanakita Beds
[Nanakita Formation]. Geologic age: Miocene [late Late

Miocene].
Pecten kyushuensis Nagao, 1928b
Plate 22, Figure 2
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 12,
no. 1, p. 37, pl. 8, figs. 1, 1a. Holotype: IGPS no. 36459(Existing). IGPS no. 36450 cited by Ogasawara (2001) isincorrect. Type locality: Wakita, Shimagô-mura, Onga-gun[currently Wakamatsu-ku, Kitakyushu City], Province of
Chikuzen [Fukuoka Prefecture]. Stratigraphic unit: Wakita
Beds [=Waita Formation]. Geologic age: Oligocene.
Pecten (Swiftopecten) nanakitaensis Nakamura, 1940b
Plate 22, Figure 3
Jour. Geol. Soc. Japan, vol. 47, no. 561, p. 264, text-fig. 1.
Holotype: IGPS 61334 (Existing). Type locality: Southernslope of Dôniwayama, Nenosiroisi-mura [=Nenoshiroishimura], Miyagi-gun [currently Nenoshiroishi, Izumiku, Sendai City], Miyagi Prefecture. Stratigraphic unit:
Nanakita Beds [=Nanakita Formation]. Geologic age:
Miocene [late Late Miocene]. Remarks: This species is ajunior synonym of “Gloripallium” crassivenium (Yokoyama,1929) (Masuda, 1958a).
Pecten (Patinopecten) planicostulatus Nomura andNiino, 1932
Plate 22, Figure 4
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 15,
no. 3, p. 177, pl. 11, figs. 2–5. Syntypes: IGPS no.
44584 (Existing). Type locality: Ichiyama, Yugashimamura [sic, Kamikano-mura], Tagata-gun, Province of Izu[currently Izu City, Shizuoka Prefecture]. Stratigraphic unit:
Shirahama Shell-Beds of Shirahama Group [Shirahama
Formation]. Geologic age: Miocene [Pliocene].
Pecten (Chlamys) sakitoensis Nagao, 1928b
Plate 22, Figure 5
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 12,
no. 1, p. 37, pl. 17, figs. 37, 38. Holotype: IGPS no.
36438 (Existing). Ogasawara (2001) erroneously cited
IGPS no. 3638 as the holotype. Type locality: Itanoura,
Seto-mura, Nishisonogi-gun [currently Itanourago, Seto,
Osetomachi, Saikai City], Province of Chikuzen [Nagasaki
Prefecture]. Stratigraphic unit: Itanoura Formation.
Geologic age: Oligocene [latest Late Eocene–earliest
Early Oligocene].
Pecten (Pecten) shirahamaensis Nomura and Niino,
1932
Plate 22, Figures 6a–b
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 15, no.
3, p. 175, pl. 12, figs. 5, 5a. Holotype: IGPS no. 44587

Catalogue of type specimens of fossil Bivalvia

45(Existing). Type locality: Nagata, Shirahama-mura, Kamogun, Province of Izu [currently Shirahama, Shimoda
City, Shizuoka Prefecture]. Stratigraphic unit: Shirahama
Shell-Beds of Shirahama Group [Shirahama Formation].
Geologic age: Miocene [Pliocene].

no. 3, p. 82, pl. 17, fig. 22. Holotype: IGPS no. 22613(Existing). Type locality: Ponhorokabetsu [Ponporokabetsu
River], Yubari-machi, Yubari-gun [currently Yubari City],
Hokkaido. Stratigraphic unit: Mikasa Group in Yubari area.
Geologic age: Cenomanian–Turonian, Cretaceous.

Pecten (Pecten) subsquamatus Nomura, 1933
Plate 22, Figures 7a–b
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 16,
no. 1, p. 53, pl. 1, fig. 11a, b. Holotype: IGPS no. 44702(Existing). Type locality: Station no. 57 (Nomura, 1933),
550 m E of Jô-tsûsho-wan, Tsûshô-shô, Byôritsu-gun,Shinchiku-shû [currently Shangtongsiaowan, Tongwan
Village, Tongshao Township, Miaoli County], Taiwan.
Lithostratigraphic unit: Byôritsu Beds [Miaoli Formation].
Geologic age: Pliocene [Pleistocene].

Pectunculus sachalinensis Yabe and Nagao, 1925
Plate 23, Figures 3a–b
S ci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 7,
no. 4, p. 112, pl. 29, figs. 7, 8. Syntypes: IGPS no. 8554
(Existing). Type locality: Sea cliff 1 km S of Cape Khoi [nearAlexandrovsk, north Saghalin] [currently AlexandrovskSakhalinsky District, Sakhalin Oblast, Russia].
Lithostratigraphic unit: Cape Khoi bed. Geologic age:
Cretaceous [Cenomanian–Turonian, Cretaceous].

Pecten (Patinopecten) taiwanus Nomura, 1933
Plate 22, Figure 8
S ci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol.
16, no. 1, p. 56, pl. 2, fig. 5a, b. Holotype: IGPS no.
45007 (Existing). Type locality: Station no. 35 (Nomura,1933), Wangwa, Kôryû-shô, Chikunan-gun, Shinchikushû [currently Wanwa, Jhonghe Village, Houlong
Township, Miaoli County], Taiwan. Lithostratigraphic
unit: Byôritsu Beds [Miaoli Formation]. Geologic age:
Pliocene [Pleistocene]. Remarks: The present species issynonymous with Mizuhopecten tokyoensis (Tokunaga,
1906) (e.g. Masuda and Huang, 1994).
Pecten (Decadopecten) tayamai Nomura and Niino, 1932
Plate 22, Figures 9–11
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 15,
no. 3, p. 178, pl. 11, figs. 6–8. Syntypes: IGPS no. 44581(Existing). Type locality: Nagata, Shirahama-mura, Kamogun, Province of Izu [currently Shirahama, Shimoda
City, Shizuoka Prefecture]. Stratigraphic unit: Shirahama
Shell-Beds of Shirahama Group [Shirahama Formation].
Geologic age: Miocene [Pliocene].
Pecten (?Amussiopecten) yabei Nomura, 1933
Plate 23, Figure 1
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 16,
no. 1, p. 59, pl. 2, figs. 3, 4. Holotype: IGPS no. 25265(Existing). Type locality: Near Rahau, Banchi, Kagi-gun,Tainan-shû [currently Arishan Township, Chiayi County],
Taiwan. Stratigraphic unit: Kaizan Beds. Geologic age:
Miocene [Late Oligocene].
Pectunculus hokkaidoensis Yabe and Nagao, 1928
Plate 23, Figure 2
S ci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 9,

Pedalion tomiyasui Nagao, 1928a
Plate 23, Figure 4
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 9, no.
3, p. 103, pl. 20, figs. 5, 5a. Lectotype: IGPS no. 35686(designated by Hatai and Nisiyama, 1952; Existing). Type
locality: Kôyagi-jima, [Kôyagi-mura], Nishisonogi-gun,
Province of Hizen [currently Koyagimachi, Nagasaki City],
Nagasaki Prefecture. Stratigraphic unit: Lower Orthaulax
japonicus Zone [=Futagojima Formation]. Geologic age:
Lower Eocene [Middle Eocene].
Periploma fujikuraense Noda, 1962b
Plate 23, Figure 5
S ci. Rep. Tohoku Univ., 2nd Ser. (Geol.), vol. 34, no.
3, p. 229, pl. 16, fig. 6. Holotype: IGPS no. 79056(Existing). Type locality: Loc. no. 1300 (Noda, 1962b),
upstream of the Higashi River, about 400 m S of Fujikura,
Matsunoyama-machi, Higashikubiki-gun [currently
Matsunoyama, Tokamachi City], Niigata Prefecture.
Stratigraphic unit: Nitta Formation. Geologic age: Early
Pliocene. Remarks: Although the name of species was
originally spelled as fujikuraensis, it is corrected herein as
fukuraense because Peliploma is a neuter.
Periploma pulchellum Hatai and Nisiyama, 1949
Plate 23, Figures 6a–b
Jour. Paleont., vol. 23, no. 1, p. 90, pl. 23, figs. 17, 18.
Holotype: IGPS no. 72508 (Existing). IGPS no. 72507
given by original description as the holotype is incorrect.
Type locality: A small cliff in front of the grocery store
at the northern entrance to Kokozura, Nakoso-machi,
Ishiki-gun [sic, Iwaki-gun] [currently Nakosomachi, Iwaki
City], Fukushima Prefecture. Stratigraphic unit: Kokozura
Formation. Geologic age: Miocene [latest Early Miocene].

46

Shin’ichi Sato, Tomoki Chiba, Takaki Yamanaka, Jun Nemoto, Masanori Shimamoto and Takashi Matsubara

Perna sanchuensis Yabe and Nagao, in Yabe, Nagao andShimizu, 1926
Plate 23, Figures 7a–b
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 9, no. 2, p.
57, pl. 12, figs. 1–4. Lectotype: IGPS no. 22457 (designatedby Hayami, 1965a; Existing). Type locality: Hachimanzawa[currently Kagahara, Kanna-machi], Tano-gun, Provinceof Kôzuke [Gumma Prefecture]. Stratigraphic unit: Shiroi
Group [Sebayashi Formation]. Geologic age: Cretaceous[late Early Cretaceous (Aptian–Albian)].
Peronidia ochii Kamada, 1962
Plate 23, Figure 8
Palaeont. Soc. Japan Spec. Pap., no. 8, p. 133, pl. 16,
figs. 6–8. Holotype: IGPS no. 79388 (Existing). Type
locality: Yumoto colliery, Joban Coal-mine, Yumotomachi,
Joban City [currently Joban-Yumotomachi, Iwaki City],
Fukushima Prefecture. Stratigraphic unit: Iwaki Formation.
Geologic age: Oligocene [latest Late Eocene–earliest
Early Oligocene].
Pholadomya brevitesta Nagao, 1943
Plate 23, Figures 9a–c
Jour. Geol. Soc. Japan, vol. 50, no. 596, p. 157, pl. 13 (10),fig. 3. Holotype: IGPS no. 72710 (Existing). Type locality:
Koikorobe, Tanohata-mura, Shimohei-gun, Province of
Rikutyû, [Iwate Prefecture]. Stratigraphic unit: precise
horizon uncertain [?Miyako Group in Tanohata area].
Geologic age: Early Cretaceous.
Pholadomya kazusaensis Nagao, 1943
Plate 23, Figures 10a–c
Jour. Geol. Soc. Japan, vol. 50, no. 596, p. 154, pl. 12 (9),figs. 2, 2a, 3, 3a. Holotype: IGPS no. 19534 (Existing).
Type locality: A point between Seki and Minato, Provinceof Kazusa [currently Futtsu City, Chiba Prefecture].
Stratigraphic unit: Nokogiri Bed [=Nokogiriyama
Formation]. Geologic age: Miocene [late Late Miocene–
earliest Early Pliocene].
Pholadomya? miyamotoi Nagao, 1943
Plate 23, Figure 11
Jour. Geol. Soc. Japan, vol. 50, no. 596, p. 158, pl. 12(9), figs. 8, 9. Lectotype: IGPS no. 72737 (designated byHayami, 1966; Existing). Type locality: Haipe, Tanohatamura, Shimohei-gun, Iwate Prefecture. Stratigraphic unit:
Hiraiga Sandstone [=Hiraiga Formation] in Tanohata area.
Geologic age: Early Cretaceous (Aptian).
Pholadomya takasimensis Nagao, 1943
Plate 23, Figures 12a–c
Jour. Geol. Soc. Japan, vol. 50, no. 596, p. 156, pl. 13 (10),fig. 1, 1a, b. Holotype: IGPS no. 72706 (Existing). Type
locality: Okinoshima, in the Takashima Coal-field, Provinceof Hizen [currently Iojimamachi, Nagasaki City, Nagasaki
Prefecture]. Stratigraphic unit: Okinoshima Bed [=Okinoshima
Formation]. Geologic age: Eocene [Middle Eocene].
Pinna asakuraensis Nagao, 1928b
Plate 23, Figures 13a–b
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 12, no.
1, p. 31, pl. 8, figs. 20, 20a. Holotype: IGPS no. 36017 or
36018 (Both numbers are written on the same specimen;
Existing). Type locality: Doshi, Hôshuyama-mura [currently
Tôhô-mura], Asakura-gun, Province of Chikuzen [Fukuoka
Prefecture]. Stratigraphic unit: Kawamagari Beds
[Kawamagari Formation]. Geologic age: Eocene.
Pitar hataii Natori, 1964
Plate 23, Figure 14
Trans. Proc. Palaeont. Soc. Japan, N.S., no. 55, p. 250, pl.
36, fig. 3a, b. Holotype: IGPS no. 85728 (Existing). Type
locality: Motomiya, Motomiya-cho [sic, Hongu, Hongucho], Higashimuro-gun [currently Hongucho, Tanabe City],
Wakayama Prefecture. Stratigraphic unit: Ukekawa-Muro
Formation. Geologic age: Oligocene or Early Miocene.
Pitar hokkaidoensis Nomura, 1935a
Plate 24, Figures 1a–b
S ci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol.
18, no. 1, p. 35, pl. 4, figs. 1, 2. Holotype: IGPS no.
56371 (Existing). Type locality: At the junction of the
Ponrurumoppe-gawa and the Rumoi-gawa, Rumoi-gun,Teshio [River cliff about 650 m NW of Owaga Station at
the junction of the Ponrurumoppe-gawa and the Rumoigawa, Rumoi-machi, Rumoi-gun, Teshio Province][currently Togeshita, Rumoimura, Rumoi City], Hokkaido.
Stratigraphic unit: Takikawa Formation [correctly Togeshita
Formation: Hayasaka and Uozumi, 1954]. Geologic age:
Pliocene [late Middle or Late Miocene].
Pitar kotoi Natori, 1964
Plate 24, Figure 2
Trans. Proc. Palaeont. Soc. Japan, N.S., no. 55, p. 252, pl.
36, fig. 4a, b. Holotype: IGPS no. 85729 (Existing). Type
locality: Motomiya, Motomiya-cho [sic, Hongu, Hongucho], Higashimuro-gun [currently Hongucho, Tanabe City],
Wakayama Prefecture. Stratigraphic unit: Ukekawa-Muro
Formation. Geologic age: Oligocene or Early Miocene.
Pitar sendaica monstrosa Nomura, 1938
Plate 24, Figures 3a–b
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 19,
p. 259, pl. 35, figs. 8a–9b. Holotype: IGPS no. 16131ACatalogue of type specimens of fossil Bivalvia(Existing). Type locality: Gôroku cliff along the rightbank of the Hirosegawa-River, western border of Sendai[Gôroku, Hirose-mura, Miyagi-gun] [currently Gôroku,
Aoba-ku, Sendai City], Miyagi Prefecture. Stratigraphic
unit: Tatunokuti Shell Bed [=Tatsunokuchi Formation].
Geologic age: Early Pliocene [latest Late Miocene–earliest
Early Pliocene].
"Pitar" shiobarensis Akutsu, 1964
Plate 24, Figure 4
Sci. Rep. Tohoku Univ., 2nd Ser. (Geol.), vol. 35, no. 3, p.
284, pl. 59, fig. 13 Holotype: IGPS no. 85505 (Existing).
Type locality: Along the Hoki River, opposite the ElectricPower Station at Sekiya, Shiobara-machi, Shioyagun [currently Nasushiobara City], Tochigi Prefecture.
Stratigraphic unit: Kanomatazawa Formation. Geologic
age: Miocene [Late Miocene].
Pitar sunakozakaensis Ogasawara, 1976
Plate 24, Figure 5
Sci. Rep. Tohoku Univ., 2nd Ser. (Geol.), vol. 46, no. 2,
p. 50, pl. 12, figs. 17, 18, 22. Holotype: IGPS no. 95028(Existing). Type locality: Loc. no. Su-01 (Ogasawara, 1976),
river side cliff of Asano-gawa at Higashi-Ichise [sic, HigashiIchinosemachi], Kanazawa City, Ishikawa Prefecture.
Stratigraphic unit: Sunakozaka Formation [=Sunagozaka
Formation]. Geologic age: Miocene [latest Early Miocene].
Pitaria? altoumbonata Nagao, 1928b
Plate 24, Figure 6
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 12, no.
1, p. 75, pl. 11, fig. 15. Holotype: IGPS no. 36445 (Existing).
Type locality: Shinyama, Ômachi-mura, Kishima-gun,
Province of Hizen [currently Omachi, Omachi-cho,
Kishima-gun, Saga Prefecture]. Stratigraphic unit: Kishima
Beds [Kishima Formation]. Geologic age: Oligocene [latest
Late Eocene–earliest Early Oligocene].
Pitaria ashiyaensis Nagao, 1928b
Plate 24, Figure 7
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 12,
no. 1, p. 72, pl. 13, fig. 1a, b. Holotype: IGPS no. 36434(Existing). Type locality: Iwaya, Shimagô-mura, Onga-gun[currently Wakamatsu-ku, Kitakyushu City], Province of
Chikuzen [Fukuoka Prefecture]. Stratigraphic unit: Yamaga
Beds [Yamaga Formation]. Geologic age: Oligocene.
Remarks: Original spelling ashiyaënsis is corrected.
Pitaria kyushuensis Nagao, 1928b
Plate 24, Figures 8a–b
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 12,
no. 1, p. 70, pl. 12, figs. 1, 1a. Holotype: IGPS no. 36238

47(Existing). Type locality: Western coast of Okinoshima,
Iôjima-mura, Nishisonogi-gun [currently Iojimamachi,
Nagasaki City], Province of Hizen [Nagasaki Prefecture].
Stratigraphic unit: Okinoshima Beds [Okinoshima
Formation]. Geologic age: Eocene [Middle Eocene].
Pitaria matsumotoi Nagao, 1928b
Plate 25, Figure 1
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 12,
no. 1, p. 73, pl. 13, figs. 4, 4a. Holotype: IGPS no. 36241(Existing). Type locality: Sea cliff facing the mouth of
Ongagawa, about 400 m NW of Yamaga, Ashiya-machi,
Onga-gun, Fukuoka Prefecture. Stratigraphic unit:
Yamaga Formation. Geologic age: Oligocene.
Pitaria takashimaensis Nagao, 1928b
Plate 25, Figure 2
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 12,
no. 1, p. 72, pl. 12, figs. 11, 11a–b. Holotype: IGPS no.
36300 (Existing). Type locality: Abô, Koyagi-jima, Koyagimura, Nishisonogi-gun, Province of Hizen [currently
Koyagimachi, Nagasaki City, Nagasaki Prefecture].
Stratigraphic unit: Okinoshima Beds [Okinoshima
Formation]. Geologic age: Eocene [Middle Eocene].
Pitaria yokoyamai Nagao, 1928b
Plate 25, Figures 3a–b
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 12, no.
1, p. 74, pl. 11, figs. 10, 10a. Holotype: IGPS no. 36443(Existing). Type locality: Sea coast S of Tokuman, Ôshima,
Kurose-mura, Nishisonogi-gun [currently Oshimacho, Saikai
City], Province of Hizen [Nagasaki Prefecture]. Stratigraphic
unit: Kakinoura Beds [Kakinoura Formation]. Geologic age:
Oligocene [latest Late Eocene–earliest Early Oligocene].
Placopecten nomurai Masuda, 1953aPlate 25, Figure 4
Trans. Proc. Palaeont. Soc. Japan, N.S., no. 12, p. 83, pl.
8, figs. 1a–7. Holotype: IGPS no. 90553 (formerly DGS
no. 1041; Existing; broken). Type locality: Stream floor ofsmall tributary of the Natori River at Moniwa, Oide-mura,
Natori-gun [currently Moniwa, Taihaku-ku, Sendai City],
Miyagi Prefecture. Stratigraphic unit: Moniwa Formation.
Geologic age: Early Miocene [early Middle Miocene].
Placopecten wakuyaensis Masuda, 1956
Plate 25, Figures 5a–b
Saito Ho-on Kai Mus. Res. Bull., no. 25, p. 23, pl. 3, figs.
1a–4. Holotype: IGPS no. 90596 (formerly DGS no. 3003;
Existing). Type locality: Oido, Wakuya-machi [sic, Wakuyacho], Tôda-gun, Miyagi Prefecture. Stratigraphic unit: Oido
Formation. Geologic age: Early Miocene [Middle Miocene].

48

Shin’ichi Sato, Tomoki Chiba, Takaki Yamanaka, Jun Nemoto, Masanori Shimamoto and Takashi Matsubara

Plicatula tuberculosa Nomura, 1933
Plate 25, Figures 6a–b
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 16, no.
1, p. 62, pl. 1, fig. 2. Holotype: IGPS no. 45030 (Existing).
Type locality: Mud-volcanoes at Konsuihei, Kyoshito,
Okayama-gun, Takao-shu [currently Kunshuiping Mud
Volcano, Yanchao Township, Kaohsiung County], Taiwan.
Geologic age: Unknown (ejecta of mud volcano).
Pododesmus (Monia) noharai Noda, 1971
Plate 25, Figure 7
Trans. Proc. Palaeont. Soc. Japan, N.S., no. 81, p.
39, pl. 7, fig. 17. Holotype: IGPS no. 86762 (Existing).
Type locality: Loc. no. 109 (Noda, 1971), W of Kogachi,
Haneji-son [currently Nago City], Okinawa-jima, Okinawa
Prefecture. Stratigraphic unit: Haneji Formation. Geologic
age: Pliocene [Early Pleistocene].
Polynemamussium masagoi Kotaka and Noda, 1967
Plate 25, Figures 8a–c
Saito Ho-on Kai Mus. Res, Bull., no. 36, p. 39, pl. 1, figs.
1–6, 8, 11. Holotype: IGPS no. 90755 (Existing). Type
locality: Loc. no. 8 (Kotaka and Noda, 1967), upstream of
the Moriai-zawa [sic, Moriya-zawa], about 800 m from thejunction with the Nakano-gawa, Ogawara [sic, Okawara],
Kuroishi City, Aomori Prefecture. Stratigraphic unit: Ogawara
Formation. Geologic age: Middle Natorian [Late Miocene].
Polynemamussium yasudae Masuda, 1962aPlate 25, Figure 9
Sci. Rep. Tohoku Univ., 2nd Ser. (Geol.), vol. 33, no. 2,
p. 156, pl. 18, figs. 9a–11. Holotype: IGPS no. 90602(Existing). Type locality: Road-side cutting, about 500 m Nof the Kogane shrine, N of Koganebasama, Motowakuya,
Wakuya-machi [sic, Wakuya, Wakuya-cho], Toda-gun,
Miyagi Prefecture. Stratigraphic unit: Oido Formation.
Geologic age: Early Miocene [Middle Miocene].
Portlandia (Portlandella) enaensis Kamada, 1962
Plate 25, Figures 10a–b
P alaeont. Soc. Japan, Spec. Pap., no. 8, p. 51, pl. 1,
figs. 28a–30. Holotype: IGPS no. 79377 (Existing). Type
locality: Hieda, Shimokajiro, Ena-machi [sic, Ena], Iwaki
City, Fukushima Prefecture. Stratigraphic unit: Honya
Formation. Geologic age: Miocene [Early Miocene].
Protocardia ibukii Nakazawa and Murata, 1966
Plate 25, Figure 11
Mem. Coll. Sci. Univ. Kyoto, Ser. B, vol. 32, no. 4, p. 314,
pl. 4, fig. 7a, b. Holotype: IGPS no. 85765 (Existing). Type
locality: Kanayama-zawa near the Omine mine [currently
Kamigocho, Tono City], Iwate Prefecture. Stratigraphic

unit: Kamihei Group in Kamihei area. Geologic age:
Neocomian, Cretaceous.
Psammocola kazusensis atsumiensis Hayasaka, 1961
S ci. Rep. Tohoku Univ., 2nd Ser. (Geol.), vol. 33,
no. 1, p. 54–55, pl. 7, figs. 1–2. Holotype: IGPS no.
78419 (Missing). Type locality: Loc. no. SC132–SC144(Hayasaka, 1961), Takamatsu coast, Takamatsu,
Akabane-cho, Atsumi-gun [currently Takamatsucho,
Tahara City], Aichi Prefecture. Stratigraphic unit: “Tonna
bed” in Toshima Sand, Middle Pleistocene.
Psammocola sekiyaensis Akutsu, 1964
Plate 25, Figure 12
Sci. Rep. Tohoku Univ., 2nd Ser. (Geol.), vol. 35, no. 3, p.
287, pl. 60, fig. 6. Holotype: IGPS no. 85511 (Existing).
Type locality: Sekiya along the Hoki River, Shiobaramachi, Shioya-gun [currently Nasushiobara City], Tochigi
Prefecture. Stratigraphic unit: Kanomatazawa Formation.
Geologic age: Miocene [Late Miocene].
Pseudoactinodontophora yabei Murata, 1971
Plate 25, Figures 13a–eTrans. Proc. Palaeont. Soc. Japan, N.S., no. 82, p.
112, pl. 14, figs. 1a, b, 4. Holotype: IGPS no. 86885(Existing). Type locality: Motoiwazawa, SW of Kawaguchi,
Sumita-cho, Kesen-gun, Iwate Prefecture. Stratigraphic
unit: Lower part of Motoiwazawa Sandstone Member,
Sakamotozawa Formation. Geologic age: Early Permian.
Pseudogrammatodon pacificus Nomura and Zinbô, 1934
Plate 25, Figures 14a–b
S ci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol.
16, no. 2, p. 153 (45), pl. 5 (1), figs. 6a–7b. Holotype:
IGPS no. 50341 (Existing). Type locality: Plateaux area
near Kamikatetu, Kikai-zima, [Kikai-cho, Oshima-gun],Kagoshima Prefecture. Lithostratigraphic unit: “Ryûkyû
Limestone”. Geologic age: Pleistocene.
Pteria sunakozakaensis Ogasawara, 1976
Plate 25, Figure 15
Sci. Rep. Tohoku Univ., 2nd Ser. (Geol.), vol. 46, no. 2, p.
40, pl. 11, figs. 15, 16. Holotype: IGPS no. 95008 (Existing).
Type locality: Loc. no. Su-01 (Ogasawara, 1976), riverside cliff of Asano-gawa at Higashi-Ichise [sic, HigashiIchinosemachi], Kanazawa City, Ishikawa Prefecture.
Stratigraphic unit: Sunakozaka Formation. Geologic age:
Miocene [latest Early Miocene].
Rexithaerus shiratoriensis Matsubara, 1994
Plate 25, Figure 16
Saito Ho-on Kai Mus. Nat. Hist. Res. Bull., no. 62, p. 24,

Catalogue of type specimens of fossil Bivalvia
pl. 1, figs. 1, 2, 3a–5b, pl. 2, figs. 1a–4. Holotype: IGPS
no. 102562 (Existing). Type locality: A small tributary of the
Shiratorigawa River, south of Shiratori, Ninohe City, Iwate
Prefecture. Stratigraphic unit: Kadonosawa Formation.
Geologic age: latest Early Miocene.
Rochefortia obsoletoradiata Nomura and Zinbô, 1936
Plate 26, Figures 1a–b
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 18, no.
3, p. 242 (14), pl. 11 (1), fig. 12a, b. Holotype: IGPS no.
51351 (Existing). Type locality: Gabusoga, Haneji-mura,
Kunigami-gun, Okinawa-jima [currently Nago City], Okinawa
Prefecture. Stratigraphic unit: Simaziri Beds [=HanejiFormation]. Geologic age: Early Pliocene [Early Pleistocene].
Saccella confusa toyomaensis Kamada, 1962
Plate 26, Figure 2
Palaeont. Soc. Japan Spec. Pap., no. 8, p. 50, pl. 2, figs.
1–5. Holotype: IGPS no. 79376 (Existing; broken). Type
locality: Numanouchi Harbor, Toyoma-machi, Taira City
[currently Numanouchi, Taira, Iwaki City], Fukushima
Prefecture. Stratigraphic unit: Numanouchi Formation.
Geologic age: Miocene [early Middle Miocene].
Saccella konnoi Hatai and Masuda, 1962
Plate 26, Figure 3
Trans. Proc. Palaeont. Soc. Japan, N.S., no. 46, p. 259,
pl. 40, figs. 3–5. Holotype: IGPS no. 90196 (formerly
DGS 4235; Existing). Type locality: Loc. no. 4 (Hatai
and Masuda, 1962), river cliff, about 600 m W of Gôdo,
Higashimatsuyama City, Saitama Prefecture. Stratigraphic
unit: Tokigawa Formation. Geologic age: Late Miocene.
Saccella saikaiensis Masuda, 1966b
Plate 26, Figures 4a–c
Trans. Proc. Palaeont. Soc. Japan, N.S., no. 64, p.
322, pl. 35, figs. 1a–3. Holotype: IGPS no. 90088(formerly DGS no. 4494; Existing). Type locality: Loc.
no. 23 (Masuda, 1966a), river cliff, about 1 km SES of
Mukaiyama, [Niemachi], Suzu City, Ishikawa Prefecture.
Stratigraphic unit: Higashi-innai Formation. Geologic age:
Miocene [latest Early Miocene].
Sanguinolites bisectus Hayasaka, 1925b
Plate 26, Figures 5a–c
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 8, no.
2, p. 13, pl. 8, fig. 10. Holotype: IGPS no. 22414 (Existing).
Type locality: Hikoroichi-mura, Kesen-gôri, prov. Rikuzen[currently Hikoroichicho, Ofunato City, Iwate Prefecture].
Stratigraphic unit: Dark grayish sandy shale. Geologic
age: Early Carboniferous.

49

Schizothaerus nuttalli kissyuensis Hatai, 1941
Plate 26, Figure 6
Bull. Biogeogr. Soc. Japan, vol. 11, no. 13, p. 109, pl. 3,
fig. 9. Holotype: IGPS no. 64751 (Existing). Type locality:
Kinseidô, Eihoku-men, Kissyû-gun, Kankyô-hoku-dô,
North Tyôsen [currently Kilju-gun, Hamgyongpuk-to, North
Korea]. Stratigraphic unit: Banko Beds [Banko Sandstone].
Geologic age: Miocene [early Middle Miocene].
Septifer sinelnikovae Noda, 1992
Plate 26, Figure 7
Sci. Rep. Tohoku Univ., 2nd Ser. (Geol.), vol. 62, nos.
1–2, p. 62, pl. 2, figs. 4a–6. Holotype: IGPS no. 100734(Existing). Type locality: Loc. no. CS7 (Noda, 1992), upperstream of the Shosambetsu River, Haboro-machi [sic,Haboro-cho] [Goryo, Shosambetsu-mura], Tomamae-gun,Rumoi Province, Hokkaido. Stratigraphic unit: Basal partof Chikubetsu Formation. Geologic age: Miocene [lateEarly–early Middle Miocene].
Serripes hataii Noda, 1962a [“1961”]
Plate 26, Figure 8
Sci. Rep. Tohoku Univ., 2nd Ser. (Geol.), Spec. Vol., no. 5(Prof. E. Kon’no Memorial Vol.), p. 224, pl. 37, fig. 3. Holotype:
IGPS no. 74593 (Existing). Type locality: Iwaigawa [sic, Kubo
River], Kamikurosawa, Hagihana-mura [sic, Haghisho-mura],
Nishiiwai-gun, [currently Sakainokami, Hagisho, Ichinoseki
City], Iwate Prefecture. Stratigraphic unit: Shimokurosawa
Formation. Noda and Tada (1968) incorrectly cited the“Nishikurosawa Formation” as the horizon of the type locality.
Geologic age: Middle Miocene.
Serripes japonica Noda, 1962a [“1961”]
Plate 26, Figure 9
Sci. Rep. Tohoku Univ., 2nd Ser. (Geol.), Spec. Vol., no.
5 (Prof. E. Kon'no Memorial Vol.), p. 225, pl. 39, fig. 4.
Holotype: IGPS no. 78680 (Existing). Type locality: Mukai,
Sakekawa [sic, Sakegawa-mura], Mogami-gun, Yamagata
Prefecture. Stratigraphic unit: Sakekawa Formation.
Geologic age: Late Miocene [Pliocene].
Serripes makiyamai nigamiensis Noda, 1962a [“1961”]
Plate 26, Figure 10
Sci. Rep. Tohoku Univ., 2nd Ser. (Geol.), Spec. Vol., no.
5 (Prof. E. Kon'no Memorial Vol.), p. 227, pl. 39, fig. 1a–c. Holotype: IGPS no. 78684 (Existing). Type locality:
Nigami, Ooshima-mura, Higashikubiki-gun [currently
Oshima-ku, Joetsu City], Niigata Prefecture. Stratigraphic
unit: Shiiya Formation. Geologic age: Late Miocene.
Serripes muraii Noda and Tada, 1968
Plate 26, Figure 11

50

Shin’ichi Sato, Tomoki Chiba, Takaki Yamanaka, Jun Nemoto, Masanori Shimamoto and Takashi Matsubara rans. Proc. Palaeont. Soc. Japan, N.S., no. 69, p. 202,T
pl. 22, fig. 22. Holotype: IGPS no. 88058 (Existing). Noda
and Tada (1968) erroneously stated IGPS no. 88059 as
the holotype in the text. Type locality: Loc. no. 2 (Nodaand Tada, 1968), small tributary of the Kakkonda River,about 4 km NNW of the Takinoue Spa, Shizukuishimachi [sic, Shizukuishi-cho], Iwate-gun, Iwate Prefecture.
Stratigraphic unit: Yamatsuda Formation. Geologic age:
Miocene [early Late Miocene].
Serripes shiobarensis Noda, 1962a [“1961”]
Plate 26, Figure 12
Sci. Rep. Tohoku Univ., 2nd Ser. (Geol.), Spec. Vol., no.
5 (Prof. E. Kon'no Memorial Vol.), p. 228, pl. 39, fig. 5.
Holotype: IGPS no. 78687 (Existing). Type locality: Cliff facing
the Hokigawa Electric Power Station along the Hoki River,
Sekiya, Shiobara-machi, Shioya-gun [currently Nasushiobara
City], Tochigi Prefecture. Stratigraphic unit: Kanomatazawa
Formation. Geologic age: Middle Miocene [Late Miocene].
Solemya (Acharax) bosoana Hatai and Koike, 1957
Plate 26, Figure 13; Plate 27, Figure 1
Japan. Jour. Geol. Geogr., vol. 28, nos. 1–3, p. 86, pl. 4,
fig. 1. Holotype: IGPS no. 94774 (Existing). Type locality:
Loc. no. 9 (Hatai and Koike, 1957), Ôkuzure, Katsuyamamachi [currently Kyonan-machi], Awa-gun, Chiba
Prefecture. Stratigraphic unit: Ôkuzure Conglomerate[Ôkuzure Formation; contained in a concretion reworked
from the underlying Hota Group]. Geologic age: Oligocene[latest Late Oligocene–Early Miocene].
Solemya (Acharax) muroensis Natori, 1964
Plate 27, Figure 2
Trans. Proc. Palaeont. Soc. Japan, N.S., no. 55, p. 249,
pl. 36, fig. 1. Holotype: IGPS no. 85727 (Existing). Type
locality: Motomiya, Motomiya-cho [sic, Hongu, Hongucho], Higashimuro-gun [currently Hongucho, Tanabe City],
Wakayama Prefecture. Stratigraphic unit: Ukekawa-Muro
Formation. Geologic age: Oligocene or Early Miocene.
Solen connectens Oyama, 1951a
Plate 27, Figure 3
Miner. Geol., vol. 4, nos. 1–2, p. 56. [nom. nov. for Solen
intermedius Nagao, 1928b, non Koch, in Philippi, 1843].
Holotype: IGPS no. 36435 (Existing). Type locality:
Yamaga, Ashiya, Ashiya-machi, Onga-gun, Fukuoka
Prefecture. Stratigraphic unit: Yamaga Formation. Geologic
age: Oligocene [latest Early–earliest Late Oligocene].
Solen intermedius Nagao, 1928b
Plate 27, Figure 3
Sci. Rep. Tôhoku Imp. Univ., 2nd. Ser. (Geol.), vol. 12,

no. 1, p. 83, pl. 4, figs. 5, 6. Holotype: IGPS no. 36435(Existing). Type locality: Yamaga, Ashiya-machi, Onga-gun,
Province of Chikuzen [Fukuoka Prefecture]. Stratigraphic
unit: Yamaga Beds [Yamaga Formation]. Geologic age:
Oligocene [latest Early–earliest Late Oligocene]. Remarks:
Because this species is preoccupied by Solen intermedius
Koch in Philippi, 1843, a new replacement name Solen
connectens was proposed by Oyama (1951a).
Solen shitakaraensis Honda, 1989
Plate 27, Figure 4
S ci. Rep. Tohoku Univ., 2nd Ser. (Geol.), vol. 60, no.
1, p. 84, pl. 8, figs. 7–10. Holotype: IGPS no. 951291 (Existing). Type locality: Loc. no. B-08 (Honda, 1989),
uppermost stream of Yukirubeshubeno-zawa a tributary of
the Tokomuro-gawa, Rushin, Urahoro-machi [sic, Urahorocho], Tokachi-gun, Hokkaido. Stratigraphic unit: Shitakara
Formation. Geologic age: Oligocene [Middle–Late Eocene].
Solenomorpha elegantissima Hayasaka, 1925b
Plate 27, Figures 5–7b; Plate 28, Figure 1
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 8, no.
2, p. 21, pl. 9, figs. 1, 1a, 2, 2a, 3–7. Syntypes: IGPS nos.
8203, 22228, 22229 (Existing). Type locality: Kinshô-zan,
Akasaka-machi, prov. Mino [currently Akasakacho, Ogaki
City, Gifu Prefecture]. Stratigraphic unit: Kuro zone [Akasaka
Limestone] Geologic age: Permian [Middle Permian].
Spisula (Cymbophora) ezoensis Yabe and Nagao, 1928
Plate 28, Figure 2
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 9, no. 3, p.
91, pl. 16, figs. 5–10. Holotype: IGPS no. 22614 (Existing).
Type locality: The upper course of the Ponnebets, Manji[currently Nishimanji, Kurisawacho, Iwamizawa City],
Ishikari Province, Hokkaido. Stratigraphic unit: Trigonia
sandstone [Middle Yezo Group in Manji area]. Geologic
age: Cretaceous [Cenomanian–Campanian, Cretaceous].
Spisula iwatensis Hatai, 1940
Plate 28, Figure 3
Bull. Biogeogr. Soc. Japan, vol. 10, no. 9, p. 130, pl. 1,
fig. 7. Holotype: IGPS no. 61360 (Existing). Type locality:
Siratori [=Shiratori], Nisatai-mura, Ninohe-gun [currently
Shiratori, Ninohe City], Iwate Prefecture. Stratigraphic unit:
Siratori Beds [=Tate Sandstone Member of Kadonosawa
Formation]. Geologic age: Miocene [latest Early Miocene].
Spisula (Mactromeris) polynyma nagakoensis Hatai andNisiyama, 1949
Plate 28, Figure 4
J our. Paleont., vol. 23, no. 1, p. 92, pl. 24, fig. 18.
Holotype: IGPS no. 72506 (Existing). Type locality: Small

Catalogue of type specimens of fossil Bivalvia
exposure in drainage W of Nagako, Nishiki-mura, Iwakigun [currently Nishikimachi, Iwaki City], Fukushima
Prefecture. Stratigraphic unit: Asagai Formation [correctly
Iwaki Formation]. Geologic age: Oligocene [latest Late
Eocene–earliest Early Oligocene].
Spisula shimotsukensis Akutsu, 1964
Plate 28, Figure 5
Sci. Rep. Tohoku Univ., 2nd Ser. (Geol.), vol. 35, no. 3, p.
287, pl. 59, fig. 11. Holotype: IGPS no. 85510 (Existing).
Type locality: Upstream of the Shimotokurazawa
valley, Sekiya, Shiobara-machi, Shioya-gun [currently
Nasushiobara City], Tochigi Prefecture. Stratigraphic unit:
Kanomatazawa Formation. Geologic age: Miocene [LateMiocene].
Strebrochondria miyamoriensis Murata, 1964
Plate 28, Figure 6
Trans. Proc. Palaeont. Soc. Japan, N.S., no. 54, p. 229, pl.
35, fig. 14a, b. Holotype: IGPS no. 85733 (Existing). Type
locality: Southeastern slope of Hiryu-yama, Miyamorimura, Kamihei-gun [currently Miyamoricho, Tono City],
Iwate Prefecture. Stratigraphic unit: Hiryuyama Formation.
Geologic age: Middle Permian.
Tapes nagahamaensis Saito, Bando and Noda, 1970
Plate 28, Figure 7
Trans. Proc. Palaeont. Soc. Japan, N.S., no. 77, p. 282,
pl. 31, figs. 1–4, 22. Holotype: IGPS no. 78707 (Existing).
Although Ogasawara (2001) cited IGPS no. 86743 fromthe Teshima Formation as the holotype, the specimens
with this reg. no. are figured specimens by Saito et al.
(1970). Type locality: Nagahama, Tonoshô-cho, Shodo-gun[sic, Shôzu-gun], Kagawa Prefecture. Stratigraphic unit:
Shikai Formation. Geologic age: Miocene [Late Eocene].
Taras millepustulata Nomura, 1933
Plate 28, Figure 8
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 16, no.
1, p. 75, pl. 3, fig. 16. Holotype: IGPS no. 45096 (Existing).
Type locality: Hanpeizan, Okayama-gun, Takao-shû[currently Ban Pin Shan, Kaohsiung City], Taiwan.
Lithostratigraphic unit: "Riukiu Limestone" [correlative of
the Ryukyu Limestone]. Geologic age: Pleistocene.
Tellina (Pharaonella) akiana Nomura, 1937
Plate 28, Figure 9
Japan. Jour. Geol. Geogr., vol. 14, nos. 3–4, p. 83, pl. 6,
fig. 4. Holotype: IGPS no. 54691 (Existing). Type locality:
Tônohama, Yasuda-mura [sic, Yasuda-cho], Aki-gun, Kôti[=Kochi] Prefecture. Stratigraphic unit: Tosa Pliocene[=Ananai Formation]. Geologic age: Pliocene [latest Late

51

Pliocene–early Early Pleistocene].
Tellina equideclivis Nagao, 1928b
Plate 28, Figure 10
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 12, no.
1, p. 79, pl. 4, fig. 29. Holotype: IGPS no. 36457 (Existing).
Type locality: Abô, Kôyagi-jima, Kôyagi-mura, Nishisonogigun, Province of Hizen [currently Koyagimachi, Nagasaki
City, Nagasaki Prefecture]. Stratigraphic unit: Okinoshima
Beds [Okinoshima Formation]. Geologic age: Eocene[Middle Eocene].
Tellina hamadai Masuda, 1955
Trans. Proc. Palaeont. Soc. Japan, N.S., no. 20, p. 122,
pl. 19, figs. 10, 11. Holotype: IGPS no. 90415 (formerly
DGS no. 1657; Missing). Type locality: Tokunari, Machinomachi, Fugeshi-gun [currently Wajima City], Ishikawa
Prefecture. Stratigraphic unit: Higashi-innai Formation.
Geologic age: Early Miocene [latest Early Miocene].
Tellina kikaizimana Nomura and Zinbô, 1934
Plate 28, Figures 11a–b
S ci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol.
16, no. 2, p. 157 (49), pl. 5 (1), fig. 19a, b. Holotype:
IGPS no. 50398 (Existing). Type locality: Plateaux area
near Kamikatetu, Kikai-zima, [Kikai-cho, Oshima-gun],Kagoshima Prefecture. Lithostratigraphic unit: “Ryûkyû
Limestone”. Geologic age: Pleistocene.
Tellina kuntsuipinensis Nomura, 1933
Plate 28, Figures 12a–b
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 16,
no. 1, p. 99, pl. 4, fig. 10a, b. Holotype: IGPS no. 48595(Existing). Type Locality: Ejecta of mud-volcanoes at
Konsuihei [currently Kunshuiping Mud Volcano, Yanchao
Township, Kaohsiung City], Taiwan. Geologic age:
Unknown (ejecta of mud volcano)Tellina maxima Nagao, 1928b
Plate 28, Figure 13
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 12,
no. 1, p. 80, pl. 4, figs. 8–11. Holotype: IGPS no. 36412(Existing; broken). Type locality: Shinyama, Ômachimura [currently Omachi-cho], Kishima-gun, Province ofHizen [Saga Prefecture]. Stratigraphic unit: Kishima Beds
[Kishima Formation]. Geologic age: Oligocene [latest Late
Eocene–earliest Early Oligocene]. Remarks: See also
Angulus maximus submaximus Mizuno, 1964.
Tellina notoensis Masuda, 1955
Plate 28, Figure 14
Trans. Proc. Palaeont. Soc. Japan, N.S., no. 20, p. 122,

52

Shin’ichi Sato, Tomoki Chiba, Takaki Yamanaka, Jun Nemoto, Masanori Shimamoto and Takashi Matsubarapl. 19, fig. 12. Holotype: IGPS no. 90451 (formerly DGS
no. 2502; Existing). Type locality: Tokunari, Machinomachi, Fugeshi-gun [currently Wajima City], Ishikawa
Prefecture. Stratigraphic unit: Higashi-innai Formation.
Geologic age: Early Miocene [latest Early Miocene].
Tellina prototenuilirata Nomura, 1933
Plate 28, Figure 15
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 16, no.
1, p. 100, pl. 4, figs. 7a–9b. Holotype: IGPS no. 48192(Existing; broken). Type locality: Futoko (?), Bôshiho,
Shiko-shô, Byôritsu-gun, Shinchiku-shû [currently
Maozihbu, Wuhwu Village, Shihwu Township, Miaoli
County], Taiwan. Lithostratigraphic unit: Byôritsu Beds
[Miaoli Formation]. Geologic age: Pliocene [Pleistocene].
Tellina scabricostulata Nomura and Zinbô, 1936
Plate 28, Figures 16a–b
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 18,
no. 3, p. 246, pl. 11, figs. 16a–17b. Holotype: IGPS no.
51333 (Existing). Type locality: Gabusoga, Haneji-mura,
Kunigami-gun [currently Nago City], Okinawa Prefecture.
Stratigraphic unit: Simaziri Beds [=Haneji Formation].
Geologic age: Early Pliocene [Early Pleistocene].
Tellina tricarinata Nagao, 1928b
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 12, no. 1,
p. 78, pl. 3, fig. 17. Holotype: IGPS no. unknown (Missing).
Type locality: Chôgiri, Kiuragi-mura, Higashimatsuuragun [currently Ouchicho, Karatsu City], Privince of Hizen[Saga Prefecture]. Stratigraphic unit: Kiuragi Beds [Kiuragi
Formation]. Geologic age: Eocene [Late Eocene].
Tellina umedairensis Shikama, 1951
Plate 28, Figures 17a–b
Trans. Proc. Palaeont. Soc. Japan, N.S., no. 1, p. 15,
text-figs. 1, 2. Holotype: IGPS no. 90404 (Existing). Type
locality: Umedaira, Yaegôchi-mura [sic, Yaekawachimura], Shimoina-gun [currently Yaegochi, Minamishinano,
Iida City], Nagano Prefecture. Stratigraphic unit: Wada
Formation. Geologic age: Oligocene [Early Miocene].
Tellina venulosa zyonoensis Hatai and Nisiyama, 1939[“1938”]
Plate 28, Figure 18
Japan. Jour. Geol. Geogr. vol. 16, nos. 1–2, p. 150, pl. 9,
fig. 3. Holotype: IGPS no. 62424 (Existing) Type locality:
A cliff of the Zintu-gawa [=Jintsu-gawa] at Zyônô [=Jônô],Sugihara-mura, Nehi-gun [sic, Nei-gun], Ettyû [=Etchû]Province [currently Jônô, Yatsuomachi, Toyama City, Toyama
Prefecture]. Stratigraphic unit: Zyônô Beds [=Otogawa
Formation]. Geologic age: Pliocene [Late Miocene].

Teredo matsushimaensis Hatai, 1951
Plate 28, Figure 19
Short Pap. Inst. Geol. Paleont., Tôhoku Univ., no. 3, p. 30,
pl. 5, figs. 1–5. Lectotype: IGPS no. 73697 (designatedby Hayami, 1966; Existing). Type locality: IGPS loc.
no. It-4 (Hatai, 1951), Matsushima, Taro-mura [sic,Taro-cho], Shimohei-gun [currently Taro, Miyako City],
Iwate Prefecture. Stratigraphic unit: Hiraiga Sandstone
Formation [Hiraiga Formation]. Geologic age: Early
Cretaceous [Aptian].
Thetironia affinis var. japonica Yabe and Nagao, 1928
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 9, no. 3, p.
89, pl. 17, figs. 14, 15. Holotype: IGPS no. 22605 (Missing).
Type locality: Ponbets [sic, Pombetsu River], a tributaryof the Ikushumbets [sic, Ikushumbetsu River], [currently
Mikasa City, Hokkaido]. Stratigraphic unit: Mikasa Formation.
Geologic age: Cenomanian–Turonian, Cretaceous.
Thracia hataii Kamada, 1955
Plate 28, Figures 20a–b
S ci. Rep., Fac. Arts and Liter., Nagasaki Univ., no.
4, p. 10, pl. 1, figs. 5, 6. Holotype: IGPS no. 72960(Existing). Type locality: IGPS loc. no. Fs-37 (Kamada,1955), about 600 m N of Ena-machi water-reservoir,
Igamesaku, Nagasaki, Ena-machi [currently Ena, Iwaki
City], Fukushima Prefecture. Stratigraphic unit: Nakayama
Formation. Geologic age: Miocene [latest Early Miocene].
Thracia kamayasikiensis Hatai, 1940
Plate 28, Figures 21a–b
Bull. Biogeogr. Soc. Japan, vol. 10, no. 9, p. 123, pl. 1,
fig. 2. Holotype: IGPS no. 61349 (Existing). Type locality:
Kamayasiki [=Kamayashiki], Tomai-mura, Ninohe-gun[currently Shimotomai, Ninohe City], Iwate Prefecture.
Stratigraphic unit: Suenomatuyama Beds [=Tomesaki
Formation]. Geologic age: Pliocene [Middle Miocene].
Thracia kidoensis Kamada, 1955
Plate 28, Figures 22a–b
Sci. Rep., Fac. Arts and Liter., Nagasaki Univ., no. 4, p.
11, pl. 1, figs. 1–2b. Holotype: IGPS no. 72958 (Existing).
Type locality: IGPS loc. no. Fs-38, in the tunnel, W of theIriumi mineral spring, Kobansaku, Kido-mura [currently
Naraha-machi], Futaba-gun, Fukushima Prefecture.
Stratigraphic unit: Asagai Formation. Geologic age:
Oligocene [Early Oligocene].
Thracia kurosawaensis Hayasaka, 1957
Plate 28, Figures 23a–b
Saito Ho-on Kai Mus. Res. Bull., no. 26, p. 28, text-fig.
1a–c. Holotype: IGPS no. 77497 (Existing). Type locality:

Catalogue of type specimens of fossil Bivalvia
Loc. no. 6 (Hayasaka, 1957), river side cliff at the westernentrance of the railway tunnel between the Iwateyuda and
Kawajiri Stations, Yuda-mura [currently Nishiwaga-machi],Waga-gun, Iwate Prefecture. Stratigraphic unit: Kurosawa
Formation. Geologic age: Miocene [Middle–Late Miocene].
Thracia (Thracia) shitakaraensis Honda, 1989
Plate 28, Figure 24; Plate 29, Figure 1
Sci. Rep. Tohoku Univ., 2nd Ser. (Geol.), vol. 60, no. 1, p.
89, pl. 6, figs. 2, 5, pl. 7, figs. 1–3, 5–7. Holotype: IGPS
no. 95428-4 (Existing). Type locality: Loc. no. B-03 (Honda,
1989), tributary of Rushin River, Rushin, Urahoro-machi[sic, Urahoro-cho], Tokachi-gun, Hokkaido. Stratigraphic
unit: Shitakara Formation. Geologic age: Oligocene[Middle–Late Eocene].
Thracidora nishizawaensis Akutsu, 1964
Plate 29, Figure 2
Sci. Rep. Tohoku Univ., 2nd Ser. (Geol.), vol. 35, no. 3, p.
283, pl. 59, fig. 4. Holotype: IGPS no. 85502 (Existing).
Ogasawara (2001) erroneously cited IGPS no. 85504 as
the holotype. Type locality: Nishizawa, Uzuno [sic, Utsuno],
Shiobara-machi, Shioya-gun [currently Nasushiobara City],
Tochigi Prefecture. Stratigraphic unit: Kanomatazawa
Formation. Geologic age: Miocene [Late Miocene].
Thyasira bisecta var. nipponica Yabe and Nomura, 1925
Plate 29, Figures 3a–c
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 7, no.
4, p. 84, pl. 23, figs. 3a–b, pl. 24, figs. 2a–4b. Lectotype:
IGPS no. 7485 (designated by Hatai and Nisiyama,
1952; Existing). Type locality: Môrai, Atsuta-mura, Atsutagun [currently Atsuta-ku, Ishikari City], Ishikari Province,
Hokkaido. Stratigraphic unit: Kawabata Series [=Morai
Formation]. Geologic age: Tertiary [Late Miocene].
Thyasira inflata Yabe and Nomura, 1925
Plate 29, Figure 4
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 7, no. 4,
p. 93, pl. 23, fig. 5a–b. Holotype: IGPS no. 17251 (Existing).
Type locality: Sea cliff S of Tsurushizaki, Hitachi-mura,
Taga-gôri, province of Hitachi [currently Higashicho, HitachiCity, Ibaraki Prefecture]. Stratigraphic unit: not cited [Hitachi
Formation]. Geologic age: Tertiary [Early Pliocene].
Thyasira quadrata Yabe and Nomura, 1925
Plate 29, Figures 5a–c
S ci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 7,
no. 4, p. 92, pl. 23, figs. 1a–b. Holotype: IGPS no. 8188
(Existing). Type locality: “Japan” (exact locallity unknown).
Stratigraphic unit: unknown. Geologic age: Tertiary [precise
age unknown].

53

Thyasira subexcavata Yabe and Endô, in Yabe andNomura, 1925
Plate 29, Figure 6
S ci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 7,
no. 4, p. 93, pl. 24, figs. 1a–b. Holotype: IGPS no. 6973
(Existing). Type locality: Near Takai, Sekimoto-mura,
Taga-gôri [currently Sekimotomachi, Kitaibaraki City],
Ibaraki Prefecture. Stratigraphic unit: not cited [Kokozura
Formation or Taga Group]. Geologic age: Tertiary[Miocene].
Thyasira tokunagai Kuroda and Habe, 1951
Plate 29, Figures 7a–b
Illust. Cat. Jpn. Shells, no. 13, p. 86 [“nom. nov.” forThyasira gouldii (Philippi) sensu Yabe and Nomura, 1925,
p. 94, pl. 23, figs. 6a–b (not of Philippi, 1845)]. Lectotype:
IGPS no. 13786 (designated herein) (Existing). Type
locality: Shinagawa, near Tôkyô [=Shinagawa-machi,
Ebara-gun, Tokyo Prefecture] [currently Shinagawa-ku,
Tokyo; precise locality unknown]. Stratigraphic unit: Tôkyô
Beds [Tokyo Formation]. Geologic age: Pliocene [Late
Pleistocene].
Trapezium (Neotrapezium) ichinohense Matsubara, 1995
Plate 29, Figures 8a–b
Trans. Proc. Palaeont. Soc. Japan, N. S., no. 180, p.
328, figs. 4-1, 4-7a, b, 4-8a–c. Holotype: IGPS no.
102605 (Existing). Type locality: Loc. no. 18 (Matsubara,
1995), upper reaches of the Nesori River about 3 km E
of Nosokei [sic, Nosoke], Ichinohe-machi, Ninohe-gun,
Iwate Prefecture. Stratigraphic unit: Yotsuyaku Formation.
Geologic age: late Early Miocene.
Trapezium (Neotrapezium) isoharense Kamada, 1962
Plate 29, Figure 9
P alaeont. Soc. Japan Spec. Pap., no. 8, p. 91, pl. 8,
figs. 4–6. Holotype: IGPS no. 79382 (Existing). Type
locality: 100 m upstream from the Futatsujima mineralspring, Isoharamachi, Kitaibaraki City, Ibaraki Prefecture.
Stratigraphic unit: Kunugidaira Formation. Geologic age:
Miocene [Early Miocene].
Trapezium isomatsuense Kotaka, 1955
Plate 29, Figure 10
Saito Ho-on Kai Mus. Res. Bull., no. 24, p. 28, pl. 2, fig.
1. Holotype: IGPS no. 74006 (Existing). Type locality:
IGPS loc. no. Ao-15 (Kotaka, 1955), upper courseof the Isomatsu-gawa, Wakimoto-mura, Kitatsugarugun [currently Wakimoto, Goshogawara City], Aomori
Prefecture. Stratigraphic unit: Isomatsu Formation.
Geologic age: Oligocene [Early Miocene].

54

Shin’ichi Sato, Tomoki Chiba, Takaki Yamanaka, Jun Nemoto, Masanori Shimamoto and Takashi Matsubara

Trapezium jobanicum Hatai and Nisiyama, 1949
Plate 29, Figure 11
J our. Paleont., vol. 23, no. 1, p. 90, pl. 24, figs. 8, 9.
Holotype: IGPS no. 72505 (Existing). IGPS no. 72506
given by original description is incorrect. Reg. no. of theholotype by Ogasawara (2001) is identical that in theoriginal description. Type locality: A small cliff in front of
the dormitory of Nippon Coal Mining Company, Nakosomachi, Ishiki-gun [sic, Iwaki-gun] [currently Nakosomachi,
Iwaki City], Fukushima Prefecture. Stratigraphic unit: Iwaki
Formation. Geologic age: Oligocene [latest Late Eocene–
earliest Early Oligocene].
Trifaricardium nomurai Kuroda and Habe, 1951
Plate 29, Figures 12a–b
Illust. Cat. Jpn. Shells, no. 13, p. 86. [nom. nov. for
Cardium (Acanthocardia) cancellatum Nomura, 1933
non Gmelin, 1791] Holotype: IGPS no. 46500 (Existing).
Type locality: Station no. 14 (Nomura, 1933), exposure,
550 m SE of Jô-tsûsho-wan, Tsûshô-shô, Byôritsu-gun,Shinchiku-shû [currently Shangtongsiaowan, Tongwan
Village, Tongshao Township, Miaoli County], Taiwan.
Lithostratigraphic unit: Byôritsu Beds [Miaoli Formation].
Geologic age: Pliocene [Pleistocene]. Remarks: This
species is a type species of Trifaricardium Kuroda andHabe, 1951 (by monotypy). See Cardium (Acanthocardia)
cancellatum Nomura, 1933 [non Gmelin, 1791] also.
Trigonarca? obsoleta Yabe and Nagao, in Yabe, Nagao
and Shimizu, 1926
Plate 29, Figure 13
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 9, no. 2,
p. 43, pl. 12, fig. 24. Holotype: IGPS no. 22538 (Existing).
Type locality: Ishidô, near Ôhinata[-mura] [currently Ishidô,Ôhinata, Sakuho-machi], Minamisaku-gun, Province ofShinano [Nagano Prefecture]. Stratigraphic unit: IshidôGroup [Ishidô Formation]. Geologic age: Cretaceous [lateEarly–early Late Cretaceous (late Neocomian or Aptian).
Trigonia ainuana Yabe and Nagao, 1928
Plate 30, Figure 1
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 9, no. 3, p.
84, pl. 16, fig. 20. Holotype: IGPS no. 22591 (Existing). Type
locality: Ponbets [=Pombetsu River] (near the Ikushumbets[=Ikushumbetsu] coal-mines), Sorachi-gun [currently Mikasa
City], Hokkaido. Stratigraphic unit: Trigonia sandstone[Mikasa Formation]. Geologic age: Cretaceous [early Late
Cretaceous (Cenomanian–Turonian)].
Trigonia brevicula Yehara, 1915
Plate 30, Figure 2
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 2, no.

2, p. 42, pl. 2, figs. 18, 19. Lectotype: IGPS no. 4329
(designated by Nakano and Numano, 1961; Existing).
Type locality: Ikushumbets [=Ikushumbetsu], some 3
km above the Ikushumbets [=Ikushumbetsu] coal mines[currently Mikasa City], Hokkaido. Stratigraphic unit: Thetis
aff. affinis zone [Middle Yezo Group (Mikasa sandstone)].
Geologic age: Cretaceous [early Late Cretaceous(Cenomanian–Turonian)].
Trigonia datemasamunei Yehara, 1915
Plate 30, Figures 3–4
S ci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 2,
no. 2, p. 38, pl. 2, figs. 13, 14. Syntypes: IGPS no. 4331
(Existing). Type locality: Raga II, Miyako District [Southerncoast of Hiraname, northeast of Raga, Tanohata-mura,
Shimohei-gun, Iwate Prefecture]. Stratigraphic unit:
Orbitolina sandstone [=Miyako Group]. Geologic age:
Cretaceous [late Early Cretaceous (Aptian–Lower Albian)].
Trigonia hokkaidoana Yehara, 1915
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 2, no. 2,
p. 39, pl. 1, figs. 1–8. Lectotype: IGPS no. 4224 (designatedby Kobayashi and Nakano, 1957; Missing). Type locality:
Tokuzo, southern coast of Hiraiga inlet, Tanohata-mura,
Shimohei-gun, Iwate Prefecture. Stratigraphic unit: Miyako
Group (Hiraiga Formation). Geologic age: late Early
Cretaceous (Aptian–Lower Albian).
Trigonia pustulosa Nagao, 1930
Plate 30, Figures 5a–8
Jour. Fac. Sci. Hokkaido Imp. Univ., Ser. 4, vol. 1, no.
1, p. 17, pl. 3, figs. 9–12. Syntypes: IGPS nos. 35603,
35608, 35609, 35610, 35612, 35613 (Existing). Type
locality: Goshonoura-jima [sic, Goshoura-jima], Amakusagun, Province of Higo [At the seashore of Enokuchi,
Goshonoura [currently Goshoura, Goshouramachi],Amakusa City, Kumamoto Prefecture]; Shishi-jima,
Nagashima-mura, Province of Satsuma [currently
Nagashima-cho, Izumi-gun, Kagoshima Prefecture].
Stratigraphic unit: Goshonoura Group. Geologic age:
Cretaceous [Albian(?)–Cenomanian, Cretaceous].
Trigonia yokoyamai Yehara, 1915
Plate 30, Figures 9–10
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 2, no. 2,
p. 41, pl. 2, figs. 15–17. Syntypes: IGPS nos. 4366, 4367
(Existing). Type locality: Tokuzo, southern coast of Hiraiga
inlet, Tanohata-mura, Shimohei-gun, Iwate Prefecture.
Stratigraphic unit: Miyako Group (Hiraiga Formation).
Geologic age: Upper Neocomian(?)–Aptian, Cretaceous.

Catalogue of type specimens of fossil Bivalvia
Vasticardium arenicoloides Akutsu, 1964
Plate 30, Figure 11
Sci. Rep. Tohoku Univ., 2nd Ser. (Geol.), vol. 35, no. 3, p.
284, pl. 59, figs. 6, 7. Syntypes: IGPS no. 85504 (Existing).
Type locality: Upstream of the Shimotokurazawa valley,
Sekiya, Shiobara-machi, Shioya-gun [currently Nasushiobara
City], Tochigi Prefecture. Stratigraphic unit: Kanomatazawa
Formation. Geologic age: Miocene [Late Miocene].
Vasticardium shimotokuraense Akutsu, 1964
Plate 30, Figure 12
Sci. Rep. Tohoku Univ., 2nd Ser. (Geol.), vol. 35, no. 3, p.
283, pl. 59, figs. 9, 10. Syntype: IGPS no. 85503 (Existing).
Type locality: Shimotokurazawa, Sekiya, Shiobaramachi, Shioya-gun [currently Nasushiobara City], Tochigi
Prefecture. Stratigraphic unit: Kanomatazawa Formation.
Geologic age: Miocene [Late Miocene]. Remarks:
Although the name of species was originally spelled as
shimotokuraensis, it is corrected as shimotokuraense
because Vasticardium is a neuter.
Vasticardium teshimaense Saito, Bando and Noda, 1970
Plate 30, Figure 13
Trans. Proc. Palaeont. Soc. Japan, N.S., no. 77, p. 283, pl.
31, figs. 5–7. Holotype: IGPS no. 86745 (Existing). Type
locality: Loc. no. 1 (Saito et al., 1970), (about 500 m SW
of Abuzaki), Teshima Island, Tonosho-cho, Shôdo-gun[sic, Shôzu-gun], Kagawa Prefecture. Stratigraphic unit:
Teshima Formation. Geologic age: Miocene [Late Eocene].
Venericardia crenulicostata Nomura, 1933
Plate 30, Figures 14a–b
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 16, no.
1, p. 69, pl. 2, figs. 12a–16b. Holotype: IGPS no. 46221(Existing). Type locality: Station no. 18 (Nomura, 1933),
650 m SE of Jô-tsûshô-wan, Tsûshô-shô, Byôritsu-gun,Shinchiku-shu [currently Shangtongsiaowan, Tongwan
Village, Tongsiao Township, Miaoli County], Taiwan.
Lithostratigraphic unit: Byôritsu Beds [Miaoli Formation].
Geologic age: Pliocene [Pleistocene].
Venericardia (Cyclocardia) enaensis Kamada, 1962
Plate 30, Figure 15
Palaeont. Soc. Japan Spec. Pap., no. 8, p. 87, pl. 7, figs.
7, 8. Holotype: IGPS no. 79381 (Existing). Type locality:
Igamesaku, Nagasaki, Ena-machi [currently Ena, Iwaki
City], Fukushima Prefecture. Stratigraphic unit: Honya
Formation. Geologic age: Miocene [late Early Miocene].
Venericardia (Cyclocardia) ferruginea complexa
Ogasawara, 1977
Plate 30, Figures 16a–b

55

 ci. Rep. Tohoku Univ., 2nd Ser. (Geol.), vol. 47, no. 2,
S
p. 107, pl. 11, fig. 3a, b, pl. 12, fig. 4. Holotype: IGPS
no. 95127 (Existing). Type locality: Loc. no. KO-22(Ogasawara, 1977), cliff, distant from the road, 300 m W
of Kaminakamachi, Kanazawa City, Ishikawa Prefecture.
Stratigraphic unit: Omma Formation. Geologic age:
Pliocene [Early Pleistocene].
Venericardia granulicostata Nomura, 1933
Plate 30, Figures 17a–e
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 16,
no. 1, p. 70, pl. 2, fig. 7a–d. Holotype: IGPS no. 46136(Existing). Type locality: 1,450 W of Hokkô, Ôbokô, Station
14, Shiko-shô, Byôritsu-gun, Shinchiku-shû [currently
Yamukeng, Jinshih Village, Hsihu Township, Miaoli
County], Taiwan. Lithostratigraphic unit: Byôritsu Beds
[Miaoli Formation]. Geologic age: Pliocene [Pleistocene].
Venericardia millegrana Nomura and Zinbô, 1934
Plate 30, Figures 18a–b
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 16,
no. 2, p. 154 (46), pl. 5 (1), figs. 13a–14b. Holotype:
IGPS no. 50375 (Existing). Type locality: Plateaux area
near Kamikatetu, Kikai-zima, [Kikai-cho, Oshima-gun],Kagoshima Prefecture. Lithostratigraphic unit: “Ryûkyû
Limestone”. Geologic age: Pleistocene.
Venericardia nagaoi Mizuno, 1964
Plate 30, Figure 19
R ep., Geol. Surv. Japan, no. 204, p. 57. Holotype:
IGPS no. 36314 (Existing). [=Venericardia nipponica
Yokoyama, sensu Nagao 1928b, pl. 13, fig. 11]. Type
locality: Nakayama Pass, Hôshuyama-mura, Asakuragun, Fukuoka Prefecture [currently Tôhô-mura, Asakuragun, Fukuoka Prefecture]. Stratigraphic unit: Kawamagari
Formation. Geologic age: Eocene.
Venericardia okinoshimaensis Mizuno, 1964
Plate 30, Figure 20
Rep., Geol. Surv. Japan, no. 204, p. 57. Holotype: IGPS no.
36371 (Existing) [=Venericardia nipponica Yokoyama, sensu
Nagao, 1928b, pl. 13, figs. 15, 15a]. Type locality: Abo,
Kôyagi-jima, Kôyagi-mura, Nishi-Sonogi-gun, Nagasaki
Prefecture [currently Abo, Kôyagi-machi, Nagasaki City,
Nagasaki Prefecture]. Stratigraphic unit: Okinoshima
Formation. Geologic age: Eocene [Middle Eocene].
Venericardia (Megacardita) ommaensis Ogasawara, 1977
S ci. Rep. Tohoku Univ., 2nd Ser. (Geol.), vol. 47, no.
2, p. 106, pl. 10, figs. 2, 5, pl. 13, fig. 3. Holotype:
IGPS 95122 (Missing). Type locality: Loc. no. KO-15(Ogasawara, 1977), river-side cliff of Asano-gawa, 400

56

Shin’ichi Sato, Tomoki Chiba, Takaki Yamanaka, Jun Nemoto, Masanori Shimamoto and Takashi Matsubaram W of Tagamihonmachi, Kanazawa City, Ishikawa
Prefecture. Stratigraphic unit: Omma Formation. Geologic
age: Pliocene [Early Pleistocene]. Remarks: Trominina
japonica (Takeda) from the Oligocene Charo Formation isregistered in the same number as the holotype.
Venericardia onukii Masuda and Takegawa, 1965
Plate 31, Figures 1a–b
Saito Ho-on Kai Mus. Res. Bull., no. 34, p. 12, pl. 1, figs.
15–22. Holotype: IGPS no. 90828 (formerly DGS no.
4683; Existing). Type locality: Loc. no. 13 (Masuda andTakegawa, 1965), river cliff about 200 m downstream ofDam, Taniyama, NW of Ashitate, Murata-machi, Shibatagun, Miyagi Prefecture. Stratigraphic unit: Fukuda Memberof Kanagase Formation. Geologic age: Middle Miocene[late Late Miocene].
Venericardia quadriangulata Nomura and Zinbô, 1934
Plate 31, Figures 2a–b
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 16,
no. 2, p. 155 (47), pl. 5 (1), figs. 15a–16b. Holotype:
IGPS no. 50385 (Existing). Type locality: Plateaux area
near Kamikatetu, Kikai-zima, [Kikai-cho, Oshima-gun],Kagoshima Prefecture. Lithostratigraphic unit: “Ryûkyû
Limestone”. Geologic age: Pleistocene.
Venericardia ryukyuensis Nomura and Zinbô, 1936
Plate 31, Figures 3a–b
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 18, no.
3, p. 240 (12), pl. 11 (1), figs. 7a–8b. Holotype: IGPS no.
51344 (Existing). Type locality: Gabusoga, Haneji-mura,
Kunigami-gun [currently Nago City], Okinawa Prefecture.
Stratigraphic unit: Simaziri Beds [=Haneji Formation].
Geologic age: Early Pliocene [Early Pleistocene].
Venericardia subnipponica Nagao, 1928b
Plate 31, Figures 4a–c
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 12, no.
1, p. 55, pl. 14, fig. 19, 19a. Holotype: IGPS no. 36405(Existing). Type locality: Taya, Ashiya-machi, [Onga-gun],
Province of Chikuzen [Fukuoka Prefecture]. Stratigraphic
unit: Yamaga Beds [Yamaga Formation]. Geologic age:
Oligocene.
Venericardia yoshidai Nagao, 1928b
Plate 31, Figure 5
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 12,
no. 1, p. 57, pl. 12, fig. 14. Holotype: IGPS no. 36327(Existing). Type locality: Ôbô, near Arita-machi [currently
Arita-cho], Nishimatsuura-gun, Province of Hizen [SagaPrefecture]. Stratigraphic unit: Kishima Beds [Kishima
Formation]. Geologic age: Oligocene [latest Late Eocene–

earliest Early Oligocene].
Veniella? japonica Nagao, 1930
Plate 31, Figures 6–7
Jour. Fac. Sci. Hokkaido Imp. Univ., Ser. 4, vol. 1, no. 1,
p. 21, pl. 2, figs. 1, 3a–c. Syntypes: IGPS nos. 35578,
44306, 44307 (Existing). Type locality: Goshonourajima [sic, Goshoura-jima], Province of Higo [currently
Goshoura, Goshouramachi, Amakusa City, Kumamoto
Prefecture]. Stratigraphic unit: Goshonoura Group in
Goshonoura area. Geologic age: Cretaceous [Albian(?)–
Cenomanian, Cretaceous].
Venus (Chione) byoritsuensis Nomura, 1933
Plate 31, Figure 8
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 16,
no. 1, p. 84, pl. 3, fig. 7a, b. Holotype: IGPS no. 48061(Existing). Type locality: Station no. 48 (Nomura, 1933),
Chû-tsushô-wan, Tsûshô-shô, Byôritsu-gun, Shinchiku-shû[currently Shangtongsiaowan, Tongwan Village, Tongsiao
Township, Miaoli County], Taiwan. Lithostratigraphic unit:
Byôritsu Beds [Miaoli Formation]. Geologic age: Pliocene[Pleistocene].
Venus (Chione) taiwanensis Nomura, 1933
Plate 31, Figure 9
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 16, no.
1, p. 86, pl. 1, figs. 12a–13b. Holotype: IGPS no. 47771(Existing). Type locality: Hanpeizan, Okayama-gun, Takaoshû [currently Ban Pin Shan, Kaohsiung City], Taiwan.
Lithostratigraphic unit: “Riukiu Limestone” [correlative of
the Ryukyu Limestone]. Geologic age: Pleistocene.
Venus (Chione) yabei Nomura and Zinbô, 1934
Plate 31, Figures 10a–b
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 16,
no. 2, p. 156 (48), pl. 5 (1), fig. 18a, b. Holotype: IGPS
no. 50651 (Existing). Type locality: Plateaux area near
Kamikatetu [=Kamikatetsu], Kikai-zima, [Kikai-cho,Oshima-gun], Kagoshima Prefecture. Lithostratigraphic
unit: “Ryûkyû Limestone”. Geologic age: Pleistocene.
Volsella yokoyamai Hatai and Nisiyama, 1949
Plate 31, Figures 11a–b
Jour. Paleont., vol. 23, no. 1, p. 90, pl. 23, figs. 13, 14.
Holotype: IGPS no. 72504 (Existing). Type locality: Near
the Mimatsu Colliery, Ogawa, Kawabe-mura, Ishiki-gun[sic, Iwaki-gun] [currently Kawabemachi, Iwaki City],
Fukushima Prefecture. Stratigraphic unit: Iwaki Formation.
Geologic age: Oligocene [latest Late Eocene–earliest
Early Oligocene].

57

Catalogue of type specimens of fossil Bivalvia
Waagenoperna elongata Nakazawa and Murata, 1966
Plate 31, Figure 12
Mem. Coll. Sci. Univ. Kyoto, Ser. B, vol. 32, no. 4, p. 312,
pl. 4, fig. 1a, b. Holotype: IGPS no. 85764 (Existing). Type
locality: Kanayama-zawa, near the Omine mine, Tono
City, Iwate Prefecture. Stratigraphic unit: Kamihei Groupin Kamihei area. Geologic age: Neocomian, Cretaceous.
Yabepecten ogasawarai Matsubara, 2003
Plate 31, Figures 13a–b
Paleontol. Res., vol. 7, no. 2, p. 169, Figs. 2, 3, 4.2, 4.4,
4.6a, b. Holotype: IGPS no. 98911 (Existing). Type locality:
Bed of the Sagae River, about 250 m downstream of"Uwano-o-hashi" (Uwano big bridge) over the Sagae Riverand about 500 m S-SE of the hamlet of Uwano, [Shiraiwa],
Sagae City, Yamagata Prefecture. Stratigraphic unit: Oya
Tuffaceous Sandstone Member of the Hongô Formation.
Geologic age: Late Miocene.
Yoldia iwatensis Hatai, 1940
Plate 31, Figures 14a–b
Bull. Biogeogr. Soc. Japan, vol. 10, no. 9, p. 121, pl. 1,
figs. 5, 6. Holotype: IGPS no. 61348 (Existing). Type
locality: Kadonosawa, Nisatai-mura, Ninohe-gun [currently
Fukuoka, Ninohe City], Iwate Prefecture. Stratigraphic
unit: Kadonosawa Beds [=Kadonosawa Formation].
Geologic age: Miocene [latest Early–earliest Middle
Miocene].
Yoldia scaphoides Nagao, 1928b
Plate 31, Figures 15a–c
Sci. Rep. Tôhoku Imp. Univ., 2nd Ser. (Geol.), vol. 12, no.
1, p. 23, pl. 2, figs. 29, 29a. Holotype: IGPS no. 35987(Existing). Type locality: Taya, Ashiya-machi, Onga-gun,
Province of Chikuzen [Fukuoka Prefecture]. Stratigraphic
unit: Yamaga Beds [Yamaga Formation]. Geologic age:
Oligocene.
Zirfaea hataii Masuda and Noda, 1969
Plate 31, Figure 16
Trans. Proc. Palaeont. Soc. Japan, N.S., no. 75, p. 133, pl.
14, figs. 3, 4. Holotype: IGPS no. 86742 (Existing). Type
locality: Gôroku cliff along the right bank of the Hirosegawa, near the waterway tunnel of the Sankyozawa
Electric Plant, in the western border of Sendai City [sic,Gôroku, Miyagi-machi, Miyagi-gun][currently Aobaku, Sendai City], Miyagi Prefecture. Stratigraphic unit:
Tatsunokuchi Formation. Geologic age: Early Pliocene[latest Late Miocene–earliest Early Pliocene].

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and Ogasawara, K., eds., The Database of Japanese Fossil
Type Specimens Described During the 20th Century (Part 1), p.
221–373, Palaeontological Society of Japan, Tokyo.
Ogasawara, K., 2002: Cenozoic Bivalvia (Supplement). In, Ikeya,
N., Hirano, H. and Ogasawara, K., eds., The Database ofJapanese Fossil Type Specimens Described During the 20th
Century (Part 2), p. 558–566, Palaeontological Society ofJapan, Tokyo.
Ogasawara, K., 2004: Outline of the database of Japanese fossil
type specimens described during the 20th Century (Part 1–3)
toward a practical use. Fossils, no. 75, p. 24­–29. (in Japanesewith English abstract)
Ogasawara, K. and Nomura, R., 1980*: Molluscan fossils fromthe Fujina Formation, Shimane Prefecture, San-in district,Japan. In, Igo, H. and Noda, H. eds., Professor Saburo Kanno
Memorial Volume, p. 79–98, pls. 9–12. Memorial Association ofProfessor Saburo Kanno’s Retirement, Tsukuba.
Ogasawara, K. and Takayasu, T., 1982*: Fossil Halicardia from
the Miocene of Akita Prefecture and the Pleistocene of Chiba
Prefecture, Japan. Venus (Japanese Journal of Malacology),

vol. 41, no. 3, p. 199–216.
Otuka, Y., 1941: Fossil Mollusca from Tazima, Hyogo Prefecture,
Japan. Japanese Journal of Geology and Geography, vol. 18,
nos. 1–2, p. 21–24.
Otuka, Y., 1943: Miocene Mollusca from Iisaka and Manseioozi,
Hukusima Prefecture, Japan. Journal of the Geological Societyof Japan, vol. 50, no. 592, p. 221–227, pl. 2. (in Japanese withEnglish abstract)
Oyama, K., 1950: Remarks on Japanese fossil molluscan name.
Kôbutu-to-Tisitu (Mineral and Geology), vol. 3, no. 6, p. 1–4. (inJapanese)
Oyama, K., 1951a*: Three new fossil shells from Palaeogene ofKyûshû. Kôbutu-to-Tisitu (Mineral and Geology), vol. 4, p. 56. (inJapanese)
Oyama, K., 1951b: A preliminary taxonomy of the Neogene
Taxodonta of Japan. Kôbutu-to-Tisitu (Mineral and Geology),
vol. 4, p. 146–156, pl. 6. (in Japanese)
Oyama, K. and Mizuno, A., 1958: On the new forms of Paleogene
molluscs in Japan. Bulletin of the Geological Survey of Japan,
vol. 9, no. 9, p. 1–18, pls. 1–4.
Oyama, K., Mizuno, A. and Sakamoto, T., 1960: Illustrated
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Geological Survey of Japan, Kawasaki.
Pergament, M. A., 1966: Zonalnaya stratigrafiya i inotseramy
nizhnej chasti verkhnego mela Tikhookeanskogo poberezhya
SSSR. [Zonal stratigraphy and Inoceramus of the lower-mostUpper Cretaceous on the Pacific coast of the USSR]. Trudy
Geologicheskii Institut, Academiia Nauk SSSR, vol. 146, p.
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Tikhogo okeana [Marine mollusks of the Anthropogene fromthe northern region of the Pacific]. Trudy Geologicheskii Institut,
vol. 357, p. 1–144, pls. 1–24. (in Russian)Philippi, R. A., 1843: Solen Tab. I. In, Philippi, R. A., Abbildungen
und Beschreibungen neuer oder wenig gekannter Conchylien,
unter Mithülfe mehrerer deutscher Conchyliologen. Erster
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Philippi R. A., 1845: Bemerkungen über die Mollusken-Fauna von
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Prime, T., 1860: Descriptions of new species of Cyrena andCorbicula in the cabinet of the Academy of Natural Sciences ofPhiladelphia. Proceedings of the Academy of Natural Sciencesof Philadelphia, vol. 12, p. 80.
Saito, M., Bando, Y. and Noda, H., 1970*: Fossil molluscs from
Teshima, Shôdo-gun, Kagawa Prefecture, Southwest Japan.
Transactions and Proceedings of the Palaeontological Societyof Japan, New Series, no. 77, p. 276–289, pl. 31.
Sakaguti, S., 1939*: A new species of Entomotis from the upper
Triassic of the Kitakami mountainland. In, Jubilee Publicationin the Commemoration of Professor H. Yabe, M.I.A. Sixtieth
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Kanreki Kinen-Kai, Sendai.
Sasaki, O., 1991*: Discrimination of two new Anadara species usingshell growth parameters and Fourier descriptors. Transactions
and Proceedings of the Palaeontological Society of Japan,
New Series, no. 161, p. 697–713.
Sato, Y., 1991: Paleontological study of molluscan assemblagesof the Miocene Moniwa Formation, northeast Japan anddescription of their Pectinidae. Report, Geological Survey ofJapan, no. 272, p. 1–249.
Say, T., 1822: An account of some of marine shells of the United

Catalogue of type specimens of fossil Bivalvia
States. Journal of the Academy of Natural Sciences ofPhiladelphia, vol. 2, p. 221–248, 257–276.
Shikama, T., 1951*: Paleogene Mollusca from south Nagano-ken.
Transactions and Proceedings of the Palaeontological Societyof Japan, New Series, no.1, p. 13–16.
Sugiyama, T., 1942: Studies on the Japanese Conularida. Journal of
the Geological Society of Japan, vol. 49, no. 589, p. 185–194,
pl. 15.
Suzuki, K., 1941*: The Palaeogene corbiculids of northwesternKyûshû. Journal of the Faculty of Science, Imperial Universityof Tokyo, Section 2, Geology, Mineralogy, Geography,
Seismology, vol. 6, pt. 3, p. 39–62, pls. 1–4.
Suzuki, K., 1943*: Zwei neue tertiäre Corbicula-Arten aus NordSachalin (Materialien zur Monographie der ostasiatischen
Corbiculiden 3). Venus, vol. 12, nos. 3–4, p. 159–171, pls. 7–8.
Takai, Y. and Matsumoto, T., 1961: Cretaceous-Tertiary unconformity
in Nagashima, southwest Kyushu. Memoirs of the Faculty of
Science, Kyusyu University, Series D, Geology, vol. 11, no. 2, p.
257–278, pls. 11–12.
Tanaka, T. and Amano, K., 1997: Taxonomy and Distribution ofMizuhopecten planicostulatus (Nomura and Niino). Bulletinof the National Science Museum. Series C, Geology andPaleontology, vol. 23, nos. 3–4, p. 101–111.
Tanaka, H. and Toshimitsu, S., 2003: Cretaceous Bivalvia. In,
Ikeya, N., Hirano, H. and Ogasawara, K., eds., The Databaseof Japanese Fossil Type Specimens Described During the20th Century (Part 3), p. 192–276, Palaeontological Society ofJapan, Tokyo.
Tokunaga, S., 1906: Fossils from the environs of Tôkyô. Journal of
the College of Science, Imperial University of Tokyo, vol. 21,
art. 2, p. 1–96, pls. 1–6.
Toshimitsu, S. and Tanaka, H. 2003: Triassic and Jurassic Bivalvia.
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Database of Japanese Fossil Type Specimens Described
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Weller, S., 1907: A report on the Cretaceous paleontology of New
Jersey, based upon the stratigraphic studies of George N.
Knapp. New Jersey Geological Survey Paleontological Series,
vol. 4, p. 1–106.
Wood, W., 1815, General Conchology; or, A Description of Shells,
Arranged According to the Linnean System, and Illustrated with
Plates, Drawn and Coloured from Nature, Vol. 1. 7 + 4 + lxi +246 p., 59 pls. J. Booth, London.
Yabe, H. and Nagao, T., 1925*: New or little-known Cretaceous
fossils from North Saghalin (Lamellibranchiata and
Gastropoda). Science Reports of the Tôhoku Imperial

63

University, Second Series (Geology), vol. 7, no. 4, p. 111–124,
pls. 28–29.
Yabe, H. and Nagao, T., 1928*: Cretaceous fossils from Hokkaido:
Annelida, Gastropoda and Lamellibranchiata. Science Reportsof the Tôhoku Imperial University Second Series (Geology),
vol. 9, no. 3, p. 77–96, pls. 16–17.
Yabe, H., Nagao, T. and Shimizu, S., 1926*: Cretaceous Mollusca
from the Sanchû-Graben in the Kwantô Mountainland, Japan.
Science Reports of the Tôhoku Imperial University, Second
Series (Geology), vol. 9, no. 2, p. 33–76, pls. 12–15.
Yabe, H. and Nomura, S., 1925*: Notes on the Recent and Tertiary
species of Thyasira from Japan. Science Reports of the
Tôhoku Imperial University, Second Series (Geology), vol. 7,
no. 4, p. 83–95, pls. 23–24.
Yabe, H. and Sato, S., 1942*: A new bivalve from the Jurassic
Torinosu series of the Abukuma mountainland. Proceedings ofthe Imperial Academy, vol. 18, no. 5, p. 251–254.
Yabe, H. and Shimizu, S., 1927*: The Triassic fauna of Rifu, near
Sendai. Science Reports of the Tôhoku Imperial University,
Second Series (Geology), vol. 11, no. 2, p. 101–136, pls. 10–
14.
Yehara, S., 1915*: The Cretaceous Trigoniae from Miyako and
Hokkaido. Science Reports of the Tôhoku Imperial University,
Second Series (Geology), vol. 2, no. 2, p. 35–44, pls. 1–2.
Yehara, S., 1920*: A pachyodont lamellibranch from the Cretaceous
deposits of Miyako in Rikuchu. Journal of the GeologicalSociety of Japan, vol. 27, no. 321, p. 39–44, pls. 1–2.
Yokoyama, M. 1920: Fossils from the Miura Peninsula and its
immediate north. Journal of the College of Science, Imperial
University of Tokyo, vol. 39, art. 6, p. 1–193, pls. 1–20.
Yokoyama, M., 1923: On some fossil Mollusca from the Neogeneof Izumo. Japanese Journal of Geology and Geography, vol. 2,
no. 1, p. 1–9, pls. 1–2.
Yokoyama, M., 1926: Fossil Mollusca from the oil-fields of Akita.
Journal of the Faculty of Science, Imperial University of Tokyo,
Section 2, Geology, Mineralogy, Geography, Seismology, vol. 1,
pt. 9, p. 377–389, pls. 44–45.
Yokoyama, M., 1928: Mollusca from the oil-field of the island of
Taiwan. Imperial Geological Survey of Japan, Report, no. 101,
p. 1–112, pls. 1–18.
Yokoyama, M., 1929: Pliocene shells from near Nanao, Noto.
Imperial Geological Survey of Japan, Report, no. 104, p. 1–7,
pls. 1–6.
Note: Bibliography with asterisk (*) behind the date of publication
includes the original description(s) of listed taxa.

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Shin’ichi Sato, Tomoki Chiba, Takaki Yamanaka, Jun Nemoto, Masanori Shimamoto and Takashi Matsubara

Explanation of Plates

All figures in natural size, unless otherwise stated
Plate 1
1 . Acanthopecten onukii Murata, 1964. IGPS no. 85745(Holotype). 2. Acanthopecten spinosus Hayasaka,
1925b. IGPS no. 22388 (Lectotype). 3. Acila divaricata
chitosensis Noda, 1962b. IGPS no. 79060 (Holotype). 4a–b. Acila (Acila) kiiensis Masuda and Katto, in Katto and
Masuda, 1978. IGPS no. 96071 (Holotype). 5. Aequipecten
matsunagiensis Masuda, 1966b. IGPS no. 90089 (Holotype).
6. Amussiopecten akiyamae Masuda, 1962a. IGPS no.
90658 (Holotype). 7a–d. Anadara (Scapharca) akitaensis
Noda, 1966a. IGPS no. 16330 (Holotype). 8a–b. Anadara(Anadara) amicula elongata Noda, 1966a. IGPS no. 85907(Holotype). 9a–b. Anadara (Anadara) arasawaensis Noda,
1966a. IGPS no. 90046 (Holotype). 10. Anadara (Anadara)
gentaroensis Noda, 1966a. IGPS no. 86411 (Holotype).
11a–c. Anadara (Anadara) hataii Noda, 1966a. IGPS no.
73210 (Holotype).
Plate 2
1a–b. Anadara (Anadara) hataii Noda, 1966a. IGPS no.
73210 (Holotype). 2a–c. Anadara (Anadara) hokkaidoensis
Noda, 1966a. IGPS no. 86408 (Holotype). 3. Anadara(Scapharca?) iwashibaraensis Noda, 1965. IGPS no.
29050 (Holotype). 4a–b. Anadara (Anadara) iwatensis
Noda, 1966a. IGPS no. 90048 (Holotype). 5a–b. Anadara(Anadara) iwatonoensis Noda, 1966a. IGPS no. 63342(Holotype). 6a–d. Anadara (Anadara) kakehataensis Hatai
and Nisiyama, 1949. IGPS no. 72510 (Holotype). 7a–c.
Anadara (Hataiarca) kogachiensis Noda, 1971. IGPS no.
86757 (Holotype).
Plate 3
1a–b. Anadara (Hataiarca) kogachiensis Noda, 1971.
IGPS no. 86757 (Holotype). 2a–b. Anadara (Anadara)
kurosedaniensis Hatai and Nisiyama, 1949. IGPS no. 72511(Holotype). 3a–b. Anadara (Scapharca) masudai Noda,
1966a. IGPS no. 51550 (Holotype). 4. Anadara (Anadara)naganoensis Noda, 1966a. IGPS no. 86410 (Holotype). 5a–b. Anadara (Tegillarca) obessa Kotaka, 1953. IGPS no.
66536 (Holotype). 6a–b. Anadara (Scapharca) omaruensis
Sasaki, 1991. IGPS no. 101325 (Holotype). 7a–b. Anadara(Hataiarca) pseudosubcrenata Ogasawara, 1977. IGPS
no. 95070 (Holotype). 8a–b. Anadara (Scapharca?)
shizuokaensis Noda, 1965. IGPS no. 78919 (Holotype). 9.
Anadara (Scapharca) taiwanica Noda, 1966a. IGPS no.
42357 (Holotype).
Plate 4
1a–c. Anadara (Scapharca) taiwanica Noda, 1966a.

IGPS no. 42357 (Holotype). 2a–b. Anadara (Scapharca)
takanabensis Sasaki, 1991. IGPS no. 101327 (Holotype).
3a–b. Anadara (Hataiarca) takayamai Noda, 1966a.
IGPS no. 86403 (Holotype). 4a–b. Anadara (Anadara)tanakuraensis Noda, 1966a. IGPS no. 28404 (Holotype). 5.
Anadara (Tosarca) tosaensis Noda, 1965. IGPS no. 54609(Holotype). 6a–c. Anadara (Anadara) tsudai Noda, 1966a.
IGPS no. 86396 (Holotype). 7a–c. Anadara (Hataiarca)
yatsuoensis Noda, 1966a. IGPS no. 86402 (Holotype).
Plate 5
1a–b. Angulus maximus submaximus Mizuno, 1964. IGPS
no. 36452 (Syntype). 2a–c. Anisocorbula ohiroi Masuda,
1966b. IGPS no. 90734 (Holotype). 3. Annuliconcha
kitakamiensis Murata, 1964. IGPS no. 22389 (Holotype).
4–5. Anomia pseudotruncata Yabe and Nagao, in Yabe,
Nagao and Shimizu, 1926. IGPS no. 22522 (Syntypes). 6a–b. Arca (Arca) kikaizimana Nomura and Zinbô, 1934. IGPS
no. 50198 (Holotype). 7a–b. Arca (Arca) miurensis Noda,
1966a. IGPS no. 23846 (Holotype). 8. Arca sakamizuensis
Hatai and Nisiyama, 1952. IGPS no. 35996 (Holotype).
9–11. Arca shinanoensis Yabe and Nagao, in Yabe, Nagao
and Shimizu, 1926. IGPS no. 22521 (Syntypes). 12.
Arca (Arca) sokeishiensis Nomura, 1933. IGPS no. 47770(Holotype external mold). 13. Arca (Arca) takaoensis
Nomura, 1933. IGPS no. 37444 (Holotype). 14a–e. Arca
(Barbatia) yokoyamai Nomura, 1933. IGPS no. 42588(Holotype). 15. Astarte shinanoensis Yabe and Nagao,
in Yabe, Nagao and Shimizu, 1926. IGPS no. 22544(Lectotype). 16. Astarte subomalioides Nagao, 1934. IGPS
no. 66446 (Lectotype). 17. Astarte subsenecta var. costata
Yabe and Nagao, in Yabe, Nagao and Shimizu, 1926. IGPS
no. 22483 (Holotype). 18a–b. Aviculopecten hataii Murata,
1964. IGPS no. 85732 (Holotype; b. external mold). 19.
Aviculopecten minoensis Hayasaka, 1925b. IGPS no. 22412(Lectotype). 20a–b. Aviculopecten? onukii Murata, 1969.
IGPS no. 91380 (Holotype). 21. Aviculopecten reticularis
Hayasaka, 1925b. IGPS no. 22227 (Holotype). 22a–b. Aviculopecten sasakii Murata, 1964. IGPS no. 85730(Holotype; b. external mold). 23. Azorius philippianus
Kotaka and Noda, 1977. IGPS no. 95087 (Holotype). 24a–b. Barbatia (Acar) hayasakai Noda, 1966a. IGPS no. 76249(Holotype). 25. Barbatia kanezawaensis Akutsu, 1964. IGPS
no. 85501 (Holotype). 26a–b. Barbatia (Acar) numaensis
Noda, 1966a. IGPS no. 86137 (Holotype). 27. Barbatia(Pugilarca) tsurushizakiensis Noda, 1966a. IGPS no. 17271(Holotype). 28. Barbatia (Barbatia) uetsukiensis Hatai
and Nisiyama, 1949. IGPS no. 72522 (Holotype). 29a–b.
Cyrena (Batissa) ponderosa Nagao, 1928a. IGPS no. 35777(Syntype) [non Prime, 1860; =Batissa nagaoi Suzuki, 1941.
IGPS no. 35777 (Holotype)].

Catalogue of type specimens of fossil Bivalvia
Plate 6
1. Cyrena (Batissa) ponderosa Nagao, 1928a. IGPS no.
35777 (Syntype). [non Prime, 1860; =Batissa nagaoi
Suzuki, 1941. IGPS no. 35777 (Holotype)]. 2a–b. Batissa
taiwanensis Nomura, 1933. IGPS no. 45087 (Holotype).
3. Brachidontes takiensis Kamada, 1962. IGPS no. 79380(Holotype). 4. Callista chinensis takagii Masuda, 1955.
IGPS no. 90885 (Holotype). 5. Callista pseudoplana Yabe
and Nagao, 1925. IGPS no. 8553 (Lectotype). 6. Callista
sekiyaensis Akutsu, 1964. IGPS no. 85506 (Syntype).
7. Cardiomya (Cardiomya) kotakai Honda, 1989. IGPS
no. 99278 (Holotype). 8a–b. Cardita katsumatai Nagao,
1928b. IGPS no. 36331 (Holotype). 9. Cardita kondoi
Nagao, 1928b. IGPS no. 36407 (Holotype). 10. Cardium(Trachycardium) hanpeizanense Nomura, 1933. IGPS
no. 37455 (Holotype). 11a–b. Cardium (Cerastoderma)hanzawai Nomura, 1933. IGPS no. 46803 (Holotype). 12.
Cardium (Cerastoderma) hizenense Nagao, 1928b. IGPS
no. 36369 (Holotype). 13a–b. Cardium (Trachycardium)
infantile Nomura and Zinbô, 1934. IGPS no. 50386(Holotype). 14. Cardium ishidoense Yabe and Nagao,
in Yabe, Nagao and Shimizu, 1926. IGPS no. 22553(Lectotype). 15a–b. Cardium (Cerastoderma?) kishimaense
Nagao, 1928b. IGPS no. 36367 (Holotype). 16. Cardium
miikense Nagao, 1928a. IGPS no. 35733 (Lectotype). 17a–b. Cardium (Nemocardium) torii Nomura, 1933. IGPS no.
49000 (Holotype). 18. Caryocorbula saikawai Kotaka, 1955.
IGPS no. 74010 (Holotype).
Plate 7
1. Chlamys (Chlamys) akutsui Masuda, 1962a. IGPS no.
28293 (Holotype). 2. Chlamys chinkopensis Masuda and
Sawada, 1961. IGPS no. 90595 (Holotype). 3. Chlamys
cosibensis hanzawae Masuda, 1959. IGPS no. 90648(Holotype). 4. Chlamys daishakaensis Masuda and Sawada,
1961. IGPS no. 90708 (Holotype). 5. Chlamys (Chlamys)
hanaishiensis Masuda, 1962a. IGPS no. 90565 (Holotype).
6. Chlamys (Chlamys) hasimotoi Masuda, 1962a. IGPS no.
90527 (Holotype). 7. Chlamys hataii Masuda and Akutsu,
1956. IGPS no. 90726 (Holotype). 8. Chlamys (Chlamys)hatakeyamae Masuda, 1962a. IGPS no. 27563 (Holotype).
9a–b. Chlamys imanishii Masuda and Sawada, 1961. IGPS
no. 72555 (Holotype). 10. Chlamys (Chlamys) ishidae
Masuda, 1962a. IGPS no. 90676 (Holotype). 11. Chlamys(Chlamys) itoigawae Masuda, 1962a. IGPS no. 90538(Holotype). 12. Chlamys kitamurai Kotaka, 1955. IGPS no.
74009 (Plastotype).
Plate 8
1. Chlamys (Chlamys) kotakae Masuda, 1962a. IGPS no.
90646 (Holotype). 2. Chlamys kumanodoensis Masuda,
1953a. IGPS no. 90522 (Holotype). 3. Chlamys (Chlamys)

65

matsunoi Masuda, 1962a. IGPS no. 90627 (Holotype). 4.
Chlamys (Chlamys) nagaoi Masuda, 1962a. IGPS no. 36440(Holotype). 5. Chlamys (Chlamys) niikappuensis Masuda,
1962a. IGPS no. 90536 (Holotype). 6a–b. Chlamys oidensis
Hatai, Masuda and Noda, 1974. IGPS no. 64419 (Holotype).
7a–b. Chlamys otukae Masuda and Sawada, 1961. IGPS
no. 90607 (Holotype). 8a–b. Chlamys (Chlamys) setsukoae
Masuda, 1962b. IGPS no. 90477 (Holotype). 9a–c. Chlamys
shitakaraensis Honda, 1980. IGPS no. 95439 (Holotype; a–b. external molds of both valves, c. cast from the holotype).
10. Chlamys tamurae Masuda and Sawada, 1961. IGPS no.
90550 (Holotype). 11. Circe (Circe) triangulus Kotaka andNoda, 1977. IGPS no. 95085 (Holotype). 12. Clementia
japonica Masuda, 1955. IGPS no. 90405 (Holotype). 13a–b. Clementia (Clementia) nakosoensis Kamada, 1962. IGPS
no. 79385 (Holotype).
Plate 91a–b. Clinocardium asagaiense arakawae Kamada, 1962.
IGPS no. 79383 (Holotype). 2a–b. Clinocardium asagaiense
makiyamae Kamada, 1962. IGPS no. 15800 (Holotype).
3a–b. Clinocardium chikagawaense Kotaka, 1950. IGPS
no. 72999 (Holotype). 4. Clinocardium hataii Hayasaka,
1956. IGPS no. 77375 (Holotype). 5. "Clinocardium"nomurai Hayasaka, 1956. IGPS no. 77376 (Holotype).
6a–c. Clinocardium omagariense Honda, 1981b. IGPS
no. 95740-1 (Holotype). 7. Codakia kitamurai Hatai andNisiyama, 1949. IGPS no. 72526 (Holotype). 8a–b. Codakia(Jagonia) okinawazimana Nomura and Zinbô, 1936. IGPS
no. 51319 (Holotype). 9a–b. Codakia semipolita Nomura,
1933. IGPS no. 45086 (Holotype). 10a–c. Conularia
rectangularis Hayasaka, 1920. IGPS no. 7320 (Lectotype).
11a–b. Corbicula (Corbicula) kotakai Honda, 1981a. IGPS
no. 96758 (Holotype). 12–14. Corbicula (Veloritina?)
sanchuensis Yabe and Nagao, in Yabe, Nagao andShimizu, 1926. IGPS no. 22467 (Syntypes). 15. Corbicula(Veloritina?) sanchuensis Yabe and Nagao, in Yabe, Nagao
and Shimizu, 1926. IGPS no. 22449 (Syntype). 16a–b.
Corbicula (Corbicula) sachalinensis Suzuki, 1943. IGPS no.
8353c (Holotype). 17a–b. Corbicula (Corbicula) shimizui
Suzuki, 1943. IGPS no. 8353a (Holotype). 18. Corbicula
takasago Nomura, 1933. IGPS no. 45079 (Holotype). 19.
Corbula (Cunaecorbula) kyushuensis Nagao, 1928a. IGPS
no. 35834 (Lectotype). 20. Corbula (Corbula) subtumida
Nagao, 1928a. IGPS no. 35828 (Lectotype). 21a–b. Corbula(Corbula) taiwanensis Nomura, 1933. IGPS no. 48966(Holotype). 22. Crassatellites asakuraensis Nagao, 1928b.
IGPS no. 36258 (Holotype). 23. Crassatellites formosanus
Nomura, 1933. IGPS no. 37441 (Holotype). 22a–b.
Crassatellites inconspicuus Nagao, 1928b. IGPS no. 36308(Holotype). 23a–c. Crassatellites matsuraensis Nagao,
1928b. IGPS no. 36310 (Holotype). 24a­–b. Crassatellites

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Shin’ichi Sato, Tomoki Chiba, Takaki Yamanaka, Jun Nemoto, Masanori Shimamoto and Takashi Matsubara

tosanus Nomura, 1937. IGPS no. 54638 (Lectotype).
Plate 10
1a–b. Crassostrea gravitesta eoilensis Kim, Noda andYoon, 1974. IGPS no. 92938 (Holotype). 2a–b. Crassostrea
sunakozakaensis Ogasawara, 1976. IGPS no. 95018(Holotype). 3. Crenella (Megacrenella) nuibetsuensis
Honda, 1989. IGPS no. 97096 (Holotype). 4. Crenella(Megacrenella) shitakaraensis Honda, 1989. IGPS no.
97111 (Holotype). 5. Crenella striatocostata Nagao, 1928b.
IGPS no. 36291 (Holotype). 6a–c. Crenella subfornicata
Nagao, 1928b. IGPS no. 36288 (Holotype). 7. Crenella
tomiyaensis Hatai and Nakamura, 1940. IGPS no. 61354(Holotype). 8. Crenipecten kesenensis Hayasaka, 1925b.
IGPS no. 22383 (Holotype). 9a–b. Ctena hataii Masuda,
1966b. IGPS no. 90729 (Holotype). 10a–b. Ctenodonta
manchuriensis Endo, 1935. IGPS no. 55582 (Holotype). 11.
Cucullaea delicatostriata Yabe and Nagao, 1925. IGPS no.
8555 (Lectotype). 12. Cucullaea ezoensis Yabe and Nagao,
1928. IGPS no. 22611 (Syntype). 13. Cucullaea nipponica
Nagao, 1928b. IGPS no. 35999 (Holotype).
Plate 11
1a–b. Cucullaea nipponica Nagao, 1928b. IGPS no. 35999(Holotype). 2. Cultellus? brevis Nagao, 1928a. IGPS no.
35788 (Holotype). 3. Cultellus? leguminoides Nagao,
1928b. IGPS no. 36447 (Holotype). 4a–b. Cyclina (Cyclina)
asagaiensis Kamada, 1952. IGPS no. 72955 (Holotype).
5. Cyclina compressa Nagao, 1928b. IGPS no. 36415(Holotype). 6a–b. Cyclina (Cyclina) japonica Kamada,
1952. IGPS no. 72952 (Holotype). 7a–b. Cyclina? nodai
Nagao, 1928b. IGPS no. 36442 (Holotype). 8. Cyclina(Cyclina) umbonata Kotaka and Noda, 1977. IGPS no.
95090 (Holotype). 9. Cyclocardia fujinaensis Ogasawara
and Nomura, 1980. IGPS no. 96024 (Holotype). 10–13.
Cyrena otsukai Yabe and Nagao, in Yabe, Nagao andShimizu, 1926. IGPS no. 22476 (Syntypes; external andinternal molds). 14–16. Cyrena shiroiensis Yabe andNagao, in Yabe, Nagao and Shimizu, 1926. IGPS no. 22451(Syntypes). 17–19. Cyrena shiroiensis var. alata Yabe andNagao, in Yabe, Nagao and Shimizu, 1926. IGPS no. 22443(Syntypes). 20. Daonella densisulcata Yabe and Shimizu,
1927. IGPS no. 7571 (Holotype). 21. Daonella densisulcata
var. subquadrata Yabe and Shimizu, 1927. IGPS no. 7890
(Holotype). 22. Daonella kotoi var. alta Yabe and Shimizu,
1927. IGPS no. 5298 (Holotype). 23. Daonella kotoi var.
multistriata Yabe and Shimizu, 1927. IGPS no. 35278(Syntype). 24. Daonella kotoi var. multistriata Yabe andShimizu, 1927. IGPS no. 22998 (Syntype). 25. Diplodonta
confusa Nagao, 1928b. IGPS no. 36351 (Holotype). 26.
Diplodonta? problematica Nagao, 1928a. IGPS no. 35827(Lectotype). 27. Dosinia chikuzenensis Nagao, 1928b.

IGPS no. 36277 (Holotype). 28. Dosinia ettyuensis Hatai
and Nisiyama, 1939 [“1938”]. IGPS no. 62429 (Holotype).
Plate 12
1a–c. Dosinia (Phacosoma) hataii Masuda, 1963. IGPS
no. 72476 (Holotype). 2. Dosinia (Kaneharaia) kaneharai
fujinaensis Masuda, 1967. IGPS no. 90888 (Holotype).
3a–b. Dosinia (Kaneharaia) kannoi Masuda, 1963. IGPS
no. 64682 (Holotype). 4a–b. Dosinia tatunokutiensis
Nomura, 1938. IGPS no. 15944 (Holotype). 5a­­–c. Ennucula
praenipponica Kamada, 1962. IGPS no. 79375 (Holotype).
6a–b. Ezocallista kurodae Kamada, 1962. IGPS no. 79384(Holotype).
Plate 13
1. Fortipecten kuroishiensis Kotaka and Noda, 1967. IGPS
no. 90767 (Holotype). 2a–b. Gafrarium (Circe) hanzawai
Nomura and Zinbô, 1936. IGPS no. 51348 (Holotype). 3.
Gari (Gari) pitogoensis Kotaka and Noda, 1977. IGPS no.
95086 (Holotype). 4. Gervillia shinanoensis Yabe andNagao, in Yabe, Nagao and Shimizu, 1926. IGPS no. 22507(Syntype). 5a–b. Glans naomiae Masuda, 1966b. IGPS no.
90181 (Holotype). 6. Gloripallium izurense Masuda, 1958a.
IGPS no. 90541 (Holotype). 7. Glycimeris altoumbonata
Nagao, 1928b. IGPS no. 36013 (Holotype). 8–10.
Glycimeris amakusensis Nagao, 1930. IGPS no. 42727(Syntypes). 11a–b. Glycimeris cisshuensis var. compressa
Nagao, 1928b. IGPS no. 36001 (Holotype). 12a–b.
Glycymeris hanzawai Nomura and Zinbô, 1934. IGPS no.
50196 (Holotype). 13a–b. Glycymeris nakosoensis Hatai
and Nisiyama, 1949. IGPS no. 72502 (Holotype). 14.
Glycymeris nozokiensis Hatai and Nisiyama, 1951. IGPS no.
72883 (Holotype). 15. Glycymeris ogawaraensis Kotaka andNoda, 1967. IGPS no. 90739 (Holotype). 16. Glycymeris
oshimaensis Noda, 1962b. IGPS no. 79057 (Holotype).
17. Glycymeris rhynconelloides Nomura and Hatai, 1939[“1938”]. IGPS no. 60022 (Holotype). 18a–b. Glycymeris
subpectiniformis Nomura and Zinbô, 1934. IGPS no. 50200(Holotype). 19a–b. Glycymeris yamaguchii Hayasaka, 1956.
IGPS no. 77373 (Holotype). 20. Grammatodon yokoyamai
Yabe and Nagao, in Yabe, Nagao and Shimizu, 1926.
IGPS no. 22555 (Lectotype). 21a–b. Halicardia akitaensis
Ogasawara and Takayasu, 1982. IGPS no. 96013 (Holotype;b. cast from the holotype). 22a–b. Halicardia miyagiensis
Fujiwara, 1992. IGPS no. 101555 (Holotype).
Plate 14
1–3. Homomya? dubia Yabe and Nagao, in Yabe, Nagao andShimizu, 1926. IGPS no. 22505 (Syntypes). 4. Inoceramus
balticus var. toyajoanus Nagao and Matumoto, 1940.
IGPS no. 4540 (Lectotype). 5. Inoceramus pedalionoides
Nagao and Matumoto, 1939 ex Inai (MS). IGPS no. 22720

Catalogue of type specimens of fossil Bivalvia(Syntype). 6. Isognomon (Isognomon) hataii Noda andFuruichi, 1972. IGPS no. 91766 (Holotype). 7. Katelysia(Nipponomarcia) endoi Hatai and Kotaka, 1952. IGPS no.
74341 (Holotype). 8. Leptochondria? hataii Murata, 1973.
IGPS no. 92661 (Holotype). 9. Leukomoides nipponicus
Ogasawara, 1976. IGPS no. 95190 (Holotype). 10a–b. Lima
amaxensis var. kumasoana Nagao, 1928a. IGPS no. 35808(Lectotype). 11. Lima eocenica Nagao, 1928a. IGPS no.
35817 (Lectotype). 12. Lima (Limatula) ishidoensis Yabe
and Nagao, in Yabe, Nagao and Shimizu, 1926. IGPS no.
22539 (Lectotype). 13a–b. Lima (Limea) limopsis Nomura
and Zinbô, 1934. IGPS no. 50368 (Holotype). 14. Limatulasakoi Masuda and Katto, in Katto and Masuda, 1978. IGPS
no. 96073 (Holotype). 15a–b. Limipecten bandoi Murata,
1969. IGPS no. 91384 (Holotype; a. external mold, b. castfrom the holotype).
Plate 15
1a–b. Lioconcha tosana Nomura, 1937. IGPS no. 54674(Holotype). 2a–b. Loripes tonohamanus Nomura, 1937.
IGPS no. 54645 (Holotype). 3a–b. Lucina kunigamiensis
Nomura and Zinbô, 1936. IGPS no. 51350 (Holotype). 4a–b. Lucinoma otukai Hatai and Nisiyama, 1949. IGPS no.
72501 (Holotype; external molds of both valves). 5a–b.
Lucinopsis? boshihoensis Nomura, 1933. IGPS no. 48007(Holotype). 6. Lucinopsis kosuiensis Nomura, 1933. IGPS
no. 48001 (Holotype; broken). 7a–b. Macoma (Macoma)
atsunaiensis Honda, 1988. IGPS no. 97106-2 (Holotype).
8a–b. Macoma hakushatonensis Nomura, 1933. IGPS no.
48589 (Holotype; broken). 9. Macoma hokiensis Akutsu,
1964. IGPS no. 85512 (Holotype). 10. Macrocallista
ariakensis Nagao, 1928a. IGPS no. 35758 (Lectotype). 11.
Macrocallista hanzawai Nagao, 1928b. IGPS no. 36236(Holotype). 12a–b. Macrocallista kahoensis Nagao,
1928b. IGPS no. 36389 (Holotype). 13a–c. Macrocallista
matsuraensis Nagao, 1928b. IGPS no. 36209 (Holotype).
14a–b. Mactra (Spisula) asperaeformsis Nomura andZinbô, 1934. IGPS no. 50396 (Holotype). 15a–b. Mactra
nakayamaensis Kamada, 1962. IGPS no. 79386 (Holotype).
16. Mactra sulcataroides Akutsu, 1964. IGPS no. 85509(Holotype). 17a–b. Margaritifera owadaensis Noda, 1970.
IGPS no. 86893 (Holotype). 18a–b. Meretrix deguchii
Hayasaka and Hayasaka, 1960. IGPS no. 77512 (Holotype).
19a–b. Meretrix meretrix ninohensis Hatai, 1940. IGPS no.
61351 (Holotype). 20. Meretrix pseudomeretrix Nagao,
1928b. IGPS no. 36361 (Holotype). 21a–b. Meretrix
tochigiensis Akutsu, 1964. IGPS no. 85507 (Holotype).
Plate 16
1a–b. Miyagipecten matsumoriensis Masuda, 1952.
IGPS no. 90690 (Lectotype). 2. Mizuhopecten kimurai
kagaensis Ogasawara, 1976. IGPS no. 95012 (Holotype).

67

3a–b. Mizuhopecten paraplebejus murataensis Masuda
and Takegawa, 1965. IGPS no. 90826 (Holotype). 4.
Modiola? ishidoensis Yabe and Nagao, in Yabe, Nagao andShimizu,1926. IGPS no. 8570 (Holotype). 5a–b. Modiolus
ezoensis Yabe and Nagao, 1928. IGPS no. 22627 (Holotype).
6a–b. Modiolus wanizakiensis Masuda, 1966b. IGPS no.
90180 (Holotype). 7a–b. Modiolus (Modiolatus) yasuhiroi
Kamada, 1962. IGPS no. 79379 (Holotype). 8a–d. Monia
denselineata Hatai, Masuda and Suzuki, 1961. IGPS no.
90505 (Holotype). 9. Mya arenaria kitahukuokaensis Hatai,
1940. IGPS no. 61363 (Holotype). 10. Mya convexa Noda,
1992. IGPS no. 100877 (Holotype). 11a–b. Myadora okadae
Hatai and Masuda, 1960. IGPS no. 90184 (Holotype).
Plate 17
1. Myadora suzuensis Masuda, 1966b. IGPS no. 90776(Holotype). 2. Myoconcha hamadaensis Yabe andShimizu, 1927. IGPS no. 35286 (Holotype). 3. Myophoria
japonica Hayasaka, 1925b. IGPS no. 22422 (Lectotype).
4. Myophoria subelegans var. tobai Hayasaka, 1924. IGPS
no. 8405 (Syntype). 5. Myophoria subelegans var. tobai
Hayasaka, 1924. IGPS no. 8404 (Syntype). 6. Myophoria
subelegans var. tobai Hayasaka, 1924. IGPS no. 8406
(Syntype; external mold). 7. Myophoria subelegans var.
tobai Hayasaka, 1924. IGPS no. 8407 (Syntype; externalmold). 8. Mytilus haboroensis Noda, 1992. IGPS no. 100727(Holotype). 9. Mytilus ogawaensis Hatai and Nisiyama,
1949. IGPS no. 72503 (Holotype). 10. Nemocardium(Keenaea) yoshidai Masuda and Miyasaka, 1994. IGPS no.
102544 (Holotype). 11. Neoburmesia iwakiensis Yabe andSato, 1942. IGPS no. 65274 (Holotype). 12a–b. Astarte
minor Nagao, 1934. IGPS no. 66425 (Lectotype) [=Nicaniella(Trautscholdia) nagaoi Matsubara, 2016. IGPS no.
66425 (Holotype)]. 13. Arca (Noetia) pondaungensis var.
transversa Nagao, 1928b [=Noetia nagaoi MacNeil, 1938].
IGPS no. 36012 (Holotype). 14. Nucula fukasawaensis
Akutsu, 1964. IGPS no. 85500 (Holotype). 15a–b. Nucula
ishidoensis Yabe and Nagao, in Yabe, Nagao and Shimizu,
1926. IGPS no. 7125 (Syntype). 16. Nucula karatsuensis
Nagao, 1928b. IGPS no. 35988 (Holotype). 17a–b. Nucula
kokozuraensis Hatai and Nisiyama, 1949. IGPS no. 72507(Holotype). 18a–c. Nucula (Acila) mirabilis var. ashiyaensis
Nagao, 1928b. IGPS no. 35981 (Holotype). 19. Nucula(Ennucula) omagariensis Honda, 1989. IGPS no. 955761 (Holotype). 20a–c. Nuculana (Nuculana) karihaensis
Hatai and Nisiyama, 1949. IGPS no. 72633 (Holotype). 21.
Nuculana (Nuculana) matsukuchiensis Noda, 1962b. IGPS
no. 79059 (Holotype). 22–24. Nuculana sanchuensis Yabe
and Nagao, in Yabe, Nagao and Shimizu, 1926. IGPS no.
7115 (Syntypes). 25a–c. Nuculites kimurai Hayasaka,
1925a. IGPS no. 22418 (Holotype). 26a–b. Nuttallia
uchigoensis Kamada, 1962. IGPS no. 79387 (Holotype).

68

Shin’ichi Sato, Tomoki Chiba, Takaki Yamanaka, Jun Nemoto, Masanori Shimamoto and Takashi Matsubara

Plate 18
1. Ostrea cassis Nagao, 1928b. IGPS no. 36031 (Holotype).
2. Ostrea lunaeformis Nagao, 1928b. IGPS no. 36039(Holotype). 3a–b. Ostrea sakitoensis Nagao, 1928b.
IGPS no. 36053 (Holotype). 4. Ostrea (Ostrea) yokoyamai
Kamada, 1962. IGPS no. 79378 (Holotype). 5a–b. Panomya
izumo Nomura and Hatai, 1939 [“1938”]. IGPS no. 51884(Holotype). 6. Panope kanomatazawaensis Akutsu, 1964.
IGPS no. 85513 (Holotype). 7. Panope kanomatazawaensis
fudozawaensis Akutsu, 1964. IGPS no. 85514 (Holotype).
8. Panopea nomurae Kamada, 1962. IGPS no. 79395(Holotype). 9. Paphia (Paphia) euglypta ohiroi Masuda,
1966b. IGPS no. 90735 (Holotype). 10. Paphia
shimotsukensis Akutsu, 1964. IGPS no. 85508 (Holotype).
11. Paphia (Paphia) suzuensis Masuda, 1966b. IGPS
no. 90086 (Holotype). 12a–b. Parallelodon obsoletiformis
Hayasaka, 1925b. IGPS no. 22392 (Lectotype).
Plate 19
1. Patinopecten ibaragiensis Masuda, 1953b. IGPS
no. 90698 (Holotype). 2. Patinopecten kagamianus
moniwaensis Masuda, 1958b. IGPS no. 90700 (Holotype).
3. Patinopecten kimurai nakosoensis Masuda, 1960.
IGPS no. 90581 (Holotype). 4. Patinopecten kimurai
yudaensis Masuda, 1960. IGPS no. 90661 (Holotype). 5a–b. Patinopecten kobiyamai Kamada, 1954. IGPS no. 72963(Holotype). 6. Patinopecten (Kotorapecten) naganoensis
Masuda, 1962a. IGPS no. 90547 (Holotype).
Plate 20
1. Patinopecten (Kotorapecten) naganoensis Masuda,
1962a. IGPS no. 90547 (Holotype). 2a–b. Patinopecten
nakajimai Masuda, 1954b. IGPS no. 90580 (Holotype).
3a–b. Patinopecten poculum tsudae Noda, 1962b. IGPS
no. 79062 (Holotype). 4. Patinopecten (Patinopecten)
tokyoensis hokurikuensis Masuda, 1962a. IGPS no. 13430(Holotype). 5. Patinopecten yamasakii kintaichiensis
Masuda, 1958c. IGPS no. 90715 (Holotype). 6.
Patinopecten yamasakii ninohensis Masuda, 1954a. IGPS
no. 90684 (Holotype). 7. Patinopecten (Patinopecten)
yessoensis nakatonbetsuensis Masuda, 1962a. IGPS no.
90662 (Holotype).
Plate 21
1. Patinopecten (Patinopecten) yessoensis yokoyamae
Masuda, 1962a. IGPS no. 90543 (Holotype). 2a–b.
Pecten (Pseudamusium) akihoensis Saga, in Matsumoto,1930. IGPS no. 90586 (Neotype). 3. Pecten (Chlamys)ashiyaensis Nagao, 1928b. IGPS no. 36448 (Holotype). 4.
Pecten (Chlamys) ashiyaensis var. denselineatus Nagao,
1928b. IGPS no. 36437 (Holotype). 5. Pecten (Vola)
byoritsuensis Nomura, 1933. IGPS no. 45001 (Holotype).

6. Pecten (Propeamussium) cowperi var. yubarensis Yabe
and Nagao, 1928. IGPS no. 22599 (Holotype). 7. Pecten(Aequipecten) gabusogaensis Nomura and Zinbô, 1936.
IGPS no. 51321 (Holotype). 8­–12. Pecten (Decadopecten)
izuensis Nomura and Niino, 1932. IGPS no. 44583(Syntypes). 13. Pecten (Chlamys) kagamianus miyagiensis
Nakamura, 1940a. IGPS no. 61336 (Holotype). 14­– 15.
Pecten (Pecten) kakisakiensis Nomura and Niino, 1932.
IGPS no. 44582 (Syntypes).
Plate 22
1a–b. Pecten (Patinopecten) kimurai matumoriensis
Nakamura, 1940a. IGPS no. 61335 (Syntype). 2. Pecten
kyushuensis Nagao, 1928b. IGPS no. 36459 (Holotype).
3. Pecten (Swiftopecten) nanakitaensis Nakamura,
1940b. IGPS 61334 (Holotype; external mold). 4. Pecten(Patinopecten) planicostulatus Nomura and Niino, 1932.
IGPS no. 44584 (Syntype). 5. Pecten (Chlamys) sakitoensis
Nagao, 1928b. IGPS no. 36438 (Holotype). 6a–b. Pecten(Pecten) shirahamaensis Nomura and Niino, 1932. IGPS no.
44587 (Holotype). 7a–b. Pecten (Pecten) subsquamatus
Nomura, 1933. IGPS no. 44702 (Holotype). 8. Pecten(Patinopecten) taiwanus Nomura, 1933. IGPS no. 45007(Holotype). 9–11. Pecten (Decadopecten) tayamai Nomura
and Niino, 1932. IGPS no. 44581 (Syntypes).
Plate 23
1. Pecten (?Amussiopecten) yabei Nomura, 1933. IGPS
no. 25265 (Holotype). 2. Pectunculus hokkaidoensis Yabe
and Nagao, 1928. IGPS no. 22613 (Holotype). 3a–b.
Pectunculus sachalinensis Yabe and Nagao, 1925. IGPS
no. 8554 (Syntype). 4. Pedalion tomiyasui Nagao, 1928a.
IGPS no. 35686 (Lectotype). 5. Periploma fujikuraense
Noda, 1962b. IGPS no. 79056 (Holotype). 6a–b. Periploma
pulchellum Hatai and Nisiyama, 1949. IGPS no. 72508(Holotype). 7a–b. Perna sanchuensis Yabe and Nagao,
in Yabe, Nagao and Shimizu, 1926. IGPS no. 22457(Lectotype). 8. Peronidia ochii Kamada, 1962. IGPS no.
79388 (Holotype). 9a–c. Pholadomya brevitesta Nagao,
1943. IGPS no. 72710 (Holotype). 10a–c. Pholadomya
kazusaensis Nagao, 1943. IGPS no. 19534 (Holotype).
11. Pholadomya? miyamotoi Nagao, 1943. IGPS no.
72737 (Lectotype). 12a–c. Pholadomya takasimensis
Nagao, 1943. IGPS no. 72706 (Holotype). 13a–b. Pinna
asakuraensis Nagao, 1928b. IGPS no. 36017 or 36018(Holotype). 14. Pitar hataii Natori, 1964. IGPS no. 85728(Holotype).
Plate 24
1a–b. Pitar hokkaidoensis Nomura, 1935a. IGPS no. 56371(Holotype). 2. Pitar kotoi Natori, 1964. IGPS no. 85729(Holotype). 3a–b. Pitar sendaica monstrosa Nomura, 1938.

Catalogue of type specimens of fossil Bivalvia
IGPS no. 16131A (Holotype). 4. "Pitar" shiobarensis Akutsu,
1964. IGPS no. 85505 (Holotype). 5. Pitar sunakozakaensis
Ogasawara, 1976. IGPS no. 95028 (Holotype). 6. Pitaria?
altoumbonata Nagao, 1928b. IGPS no. 36445 (Holotype).
7. Pitaria ashiyaensis Nagao, 1928b. IGPS no. 36434(Holotype). 8a–b. Pitaria kyushuensis Nagao, 1928b. IGPS
no. 36238 (Holotype).
Plate 25
1. Pitaria matsumotoi Nagao, 1928b. IGPS no. 36241(Holotype). 2. Pitaria takashimaensis Nagao, 1928b. IGPS
no. 36300 (Holotype). 3a–b. Pitaria yokoyamai Nagao,
1928b. IGPS no. 36443 (Holotype). 4. Placopecten nomurai
Masuda, 1953a. IGPS no. 90553 (Holotype; broken). 5a–b. Placopecten wakuyaensis Masuda, 1956. IGPS no.
90596 (Holotype). 6a–b. Plicatula tuberculosa Nomura,
1933. IGPS no. 45030 (Holotype). 7. Pododesmus (Monia)
noharai Noda, 1971. IGPS no. 86762 (Holotype). 8a–c. Polynemamussium masagoi Kotaka and Noda, 1967.
IGPS no. 90755 (Holotype). 9. Polynemamussium yasudae
Masuda, 1962a. IGPS no. 90602 (Holotype). 10a–b.
Portlandia (Portlandella) enaensis Kamada, 1962. IGPS
no. 79377 (Holotype). 11. Protocardia ibukii Nakazawa
and Murata, 1966. IGPS no. 85765 (Holotype; externalmold). 12. Psammocola sekiyaensis Akutsu, 1964. IGPS
no. 85511 (Holotype). 13a–e. Pseudoactinodontophorayabei Murata, 1971. IGPS no. 86885 (Holotype; a. internalmold, b. external mold, c–e. casts from the holotype). 14a–b. Pseudogrammatodon pacificus Nomura and Zinbô, 1934.
IGPS no. 50341 (Holotype). 15. Pteria sunakozakaensis
Ogasawara, 1976. IGPS no. 95008 (Holotype). 16.
Rexithaerus shiratoriensis Matsubara, 1994. IGPS no.
102562 (Holotype).
Plate 26
1a–b. Rochefortia obsoletoradiata Nomura and Zinbô,1936. IGPS no. 51351 (Holotype). 2. Saccella confusa
toyomaensis Kamada, 1962. IGPS no. 79376 (Holotype;broken). 3. Saccella konnoi Hatai and Masuda, 1962. IGPS
no. 90196 (Holotype). 4a–c. Saccella saikaiensis Masuda,
1966b. IGPS no. 90088 (Holotype). 5a–c. Sanguinolites
bisectus Hayasaka, 1925b. IGPS no. 22414 (Holotype). 6.
Schizothaerus nuttalli kissyuensis Hatai, 1941. IGPS no.
64751 (Holotype). 7. Septifer sinelnikovae Noda, 1992.
IGPS no. 100734 (Holotype). 8. Serripes hataii Noda,
1962a [“1961”]. IGPS no. 74593 (Holotype). 9. Serripes
japonica Noda, 1962a [“1961”]. IGPS no. 78680 (Holotype).
10. Serripes makiyamai nigamiensis Noda, 1962a [“1961”].
IGPS no. 78684 (Holotype). 11. Serripes muraii Noda
and Tada, 1968. IGPS no. 88058 (Holotype). 12. Serripes
shiobarensis Noda, 1962a [“1961”]. IGPS no. 78687(Holotype). 13. Solemya (Acharax) bosoana Hatai and69

Koike, 1957. IGPS no. 94774 (Holotype).
Plate 27
1. Solemya (Acharax) bosoana Hatai and Koike, 1957.
IGPS no. 94774 (Holotype). 2. Solemya (Acharax)
muroensis Natori, 1964. IGPS no. 85727 (Holotype). 3.
Solen intermedius Nagao, 1928b [=Solen connectens
Oyama, 1951a]. IGPS no. 36435 (Holotype). 4. Solenshitakaraensis Honda, 1989. IGPS no. 95129-1 (Holotype).
5. Solenomorpha elegantissima Hayasaka, 1925b. IGPS
no. 8203 (Syntype). 6a–c. Solenomorpha elegantissima
Hayasaka, 1925b. IGPS no. 22228 (Syntype). 7a–b.
Solenomorpha elegantissima Hayasaka, 1925b. IGPS no.
22229 (Syntype).
Plate 28
1. Solenomorpha elegantissima Hayasaka, 1925b. IGPS
no. 22229 (Syntype). 2. Spisula (Cymbophora) ezoensis
Yabe and Nagao, 1928. IGPS no. 22614 (Holotype). 3.
Spisula iwatensis Hatai, 1940. IGPS no. 61360 (Holotype).
4. Spisula (Mactromeris) polynyma nagakoensis Hatai andNisiyama, 1949. IGPS no. 72506 (Holotype). 5. Spisulashimotsukensis Akutsu, 1964. IGPS no. 85510 (Holotype).
6. Strebrochondria miyamoriensis Murata, 1964. IGPS no.
85733 (Holotype). 7. Tapes nagahamaensis Saito, Bandoand Noda, 1970. IGPS no. 78707 (Holotype). 8. Taras
millepustulata Nomura, 1933. IGPS no. 45096 (Holotype). 9.
Tellina (Pharaonella) akiana Nomura, 1937. IGPS no. 54691(Holotype). 10. Tellina equideclivis Nagao, 1928b. IGPS no.
36457 (Holotype). 11a–b. Tellina kikaizimana Nomura andZinbô, 1934. IGPS no. 50398 (Holotype). 12a–b. Tellina
kuntsuipinensis Nomura, 1933. IGPS no. 48595 (Holotype).
13. Tellina maxima Nagao, 1928b. IGPS no. 36412(Holotype; broken). 14. Tellina notoensis Masuda, 1955.
IGPS no. 90451 (Holotype). 15. Tellina prototenuilirata
Nomura, 1933. IGPS no. 48192 (Holotype; broken). 16a–b.
Tellina scabricostulata Nomura and Zinbô, 1936. IGPS no.
51333 (Holotype). 17a–b.Tellina umedairensis Shikama,
1951. IGPS no. 90404 (Holotype). 18. Tellina venulosa
zyonoensis Hatai and Nisiyama, 1939 [“1938”]. IGPS no.
62424 (Holotype). 19. Teredo matsushimaensis Hatai,
1951. IGPS no. 73697 (Lectotype). 20a–b. Thracia hataii
Kamada, 1955. IGPS no. 72960 (Holotype). 21a–b. Thracia
kamayasikiensis Hatai, 1940. IGPS no. 61349 (Holotype).
22a–b. Thracia kidoensis Kamada, 1955. IGPS no. 72958(Holotype). 23a–b. Thracia kurosawaensis Hayasaka,
1957. IGPS no. 77497 (Holotype). 24. Thracia (Thracia)
shitakaraensis Honda, 1989. IGPS no. 95428-4 (Holotype).
Plate 29
1. Thracia (Thracia) shitakaraensis Honda, 1989. IGPS no.
95428-4 (Holotype). 2. Thracidora nishizawaensis Akutsu,

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1964. IGPS no. 85502 (Holotype). 3a–c. Thyasira bisecta
var. nipponica Yabe and Nomura, 1925. IGPS no. 7485
(Lectotype). 4. Thyasira inflata Yabe and Nomura, 1925.
IGPS no. 17251 (Holotype). 5a–c. Thyasira quadrata Yabe
and Nomura, 1925. IGPS no. 8188 (Holotype). 6. Thyasirasubexcavata Yabe and Endô, in Yabe and Nomura, 1925.
IGPS no. 6973 (Holotype). 7a–b. Thyasira tokunagai
Kuroda and Habe, 1951. IGPS no. 13786 (Lectotype,designated herein). 8a–b. Trapezium (Neotrapezium)
ichinohense Matsubara, 1995. IGPS no. 102605 (Holotype).
9. Trapezium (Neotrapezium) isoharense Kamada, 1962.
IGPS no. 79382 (Holotype). 10. Trapezium isomatsuense
Kotaka, 1955. IGPS no. 74006 (Holotype). 11. Trapezium
jobanicum Hatai and Nisiyama, 1949. IGPS no. 72505(Holotype). 12a­–b. Cardium (Acanthocardia) cancellatum
Nomura, 1933 [=Trifaricardium nomurai Kuroda and Habe,
1951]. IGPS no. 46500 (Holotype). 13. Trigonarca? obsoleta
Yabe and Nagao, in Yabe, Nagao and Shimizu, 1926. IGPS
no. 22538 (Holotype).
Plate 30
1. Trigonia ainuana Yabe and Nagao, 1928. IGPS no. 22591(Holotype). 2. Trigonia brevicula Yehara, 1915. IGPS no.
4329 (Lectotype). 3–4. Trigonia datemasamunei Yehara,
1915. IGPS no. 4331 (Syntypes). 5­a–b. Trigonia pustulosa
Nagao, 1930. IGPS no. 35612 (Syntype; a. external mold,b. cast from the syntype). 6a–b. Trigonia pustulosa Nagao,
1930. IGPS no. 35613 (Syntype; a. external mold, b. castfrom the syntype). 7–8. Trigonia pustulosa Nagao, 1930.
IGPS no. 35613 (Syntypes; 7a. external mold, 7b–8.
casts from syntypes). 9. Trigonia yokoyamai Yehara,
1915. IGPS no. 4367 (Syntype). 10. Trigonia yokoyamai
Yehara, 1915. IGPS nos. 4366 (Syntype). 11. Vasticardium
arenicoloides Akutsu, 1964. IGPS no. 85504 (Syntype).
12. Vasticardium shimotokuraense Akutsu, 1964. IGPS no.
85503 (Syntype). 13. Vasticardium teshimaense Saito,

Bando and Noda, 1970. IGPS no. 86745 (Holotype). 14a–b. Venericardia crenulicostata Nomura, 1933. IGPS no.
46221 (Holotype). 15. Venericardia (Cyclocardia) enaensis
Kamada, 1962. IGPS no. 79381 (Holotype). 16a–b.
Venericardia (Cyclocardia) ferruginea complexa Ogasawara,
1977. IGPS no. 95127 (Holotype). 17a–e. Venericardia
granulicostata Nomura, 1933. IGPS no. 46136 (Holotype).
18a–b. Venericardia millegrana Nomura and Zinbô, 1934.
IGPS no. 50375 (Holotype). 19. Venericardia nagaoi
Mizuno, 1964. IGPS no. 36314 (Holotype). 20. Venericardia
okinoshimaensis Mizuno, 1964. IGPS no. 36371 (Holotype).
Plate 31
1a–b. Venericardia onukii Masuda and Takegawa,
1965. IGPS no. 90828 (Holotype). 2a–b. Venericardia
quadriangulata Nomura and Zinbô, 1934. IGPS no. 50385(Holotype). 3a–b. Venericardia ryukyuensis Nomura
and Zinbô, 1936. IGPS no. 51344 (Holotype). 4a–c.
Venericardia subnipponica Nagao, 1928b. IGPS no. 36405(Holotype). 5. Venericardia yoshidai Nagao, 1928b. IGPS
no. 36327 (Holotype). 6a–b. Veniella? japonica Nagao,
1930. IGPS no. 44307 (Syntype). 7. Veniella? japonica
Nagao, 1930. IGPS no. 44306 (Syntype). 8. Venus (Chione)
byoritsuensis Nomura, 1933. IGPS no. 48061 (Holotype).
9. Venus (Chione) taiwanensis Nomura, 1933. IGPS no.
47771 (Holotype). 10a–b. Venus (Chione) yabei Nomura
and Zinbô, 1934. IGPS no. 50651 (Holotype). 11a–b.
Volsella yokoyamai Hatai and Nisiyama, 1949. IGPS no.
72504 (Holotype). 12. Waagenoperna elongata Nakazawa
and Murata, 1966. IGPS no. 85764 (Holotype). 13a–b.
Yabepecten ogasawarai Matsubara, 2003. IGPS no. 98911(Holotype). 14a–b. Yoldia iwatensis Hatai, 1940. IGPS
no. 61348 (Holotype; b. external mold). 15a–c. Yoldia
scaphoides Nagao, 1928b. IGPS no. 35987 (Holotype). 16.
Zirfaea hataii Masuda and Noda, 1969. IGPS no. 86742(Holotype).

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Indexainuana, Trigonia akiana, Tellina (Pharaonella) akihoensis, Pecten (Pseudamusium) akitaensis, Anadara (Scapharca) akitaensis, Halicardia akiyamae, Amussiopecten 
akutsui, Chlamys (Chlamys) alata, Cyrena shiroiensis var alta, Daonella kotoi var 
altoumbonata, Glycimeris 
altoumbonata, Pitaria? amakusensis, Glycimeris 
andoi, Arca (Arca) arakawae, Clinocardium asagaiensearasawaensis, Anadara (Anadara) arenicoloides, Vasticardium 
ariakensis, Macrocallista 
asagaiensis, Cyclina (Cyclina) asakuraensis, Crassatellites 
asakuraensis, Pinna ashiyaensis, Nucula (Acila) mirabilis var ashiyaensis, Pecten (Chlamys) ashiyaensis, Pitaria asperaeformsis, Mactra (Spisula) atsumiensis, Psammocola kazusensis atsunaiensis, Macoma (Macoma) bandoi, Limipecten 
bisectus, Sanguinolites boshihoensis, Lucinopsis? bosoana, Solemya (Acharax) brevicula, Trigonia 
brevis, Cultellus? brevitesta, Pholadomya 
byoritsuensis, Pecten (Vola) byoritsuensis, Venus (Chione) cancellatum, Cardium (Acanthocardia) cassis, Ostrea chikagawaense, Clinocardium 
chikuzenensis, Dosinia chinkopensis, Chlamys 
chitosensis, Acila divaricata complexa, Venericardia (Cyclocardia) ferruginea compressa, Cyclina compressa, Glycimeris cisshuensis var 
confusa, Diplodonta connectens, Solen convexa, Mya 
costata, Astarte subsenecta var 
crenulicostata, Venericardia daishakaensis, Chlamys 

p. 54
p. 51
p. 43
p. 13
p. 32
p. 13
p. 21
p. 29
p. 29
p. 31
p. 47
p. 31
p. 16
p. 23
p. 13
p. 55
p. 35
p. 27
p. 25
p. 46
p. 39
p. 43
p. 47
p. 36
p. 48
p. 35
p. 34
p. 49
p. 35
p. 50
p. 54
p. 27
p. 46
p. 43
p. 56
p. 20
p. 40
p. 24
p. 29
p. 21
p. 13
p. 55
p. 28
p. 31
p. 29
p. 50
p. 37
p. 18
p. 55
p. 21

datemasamunei, Trigonia deguchii, Meretrix delicatostriata, Cucullaea denselineata, Monia denselineatus, Pecten (Chlamys) ashiyaensis var densisulcata, Daonella 
dubia, Homomya? elegantissima, Solenomorpha elongata, Anadara (Anadara) amiculaelongata, Waagenoperna enaensis, Portlandia (Portlandella) enaensis, Venericardia (Cyclocardia) endoi, Katelysia (Nipponomarcia) eocenica, Lima eoilensis, Crassostrea gravitesta equideclivis, Tellina 
ettyuensis, Dosinia ezoensis, Cucullaea ezoensis, Modiolus ezoensis, Spisula (Cymbophora) formosanus, Crassatellites 
fudozawaensis, Panope kanomatazawaensis 
fujikuraense, Periploma 
fujinaensis, Cyclocardia fujinaensis, Dosinia (Kaneharaia) kaneharai
fukasawaensis, Nucula gabusogaensis, Pecten (Aequipecten) gentaroensis, Anadara (Anadara) gorokuensis, Glycymeris 
granulicostata, Venericardia haboroensis, Mytilus hakushatonensis, Macoma 
hamadaensis, Myoconcha 
hamadai, Tellina hanaishiensis, Chlamys (Chlamys) hanpeizanense, Cardium (Trachycardium) hanzawae, Chlamys cosibensis
hanzawai, Cardium (Cerastoderma) hanzawai, Gafrarium (Circe) hanzawai, Glycymeris hanzawai, Macrocallista hasimotoi, Chlamys (Chlamys) hataii, Anadara (Anadara) hataii, Aviculopecten hataii, Chlamys hataii, Clinocardium 
hataii, Ctena hataii, Dosinia (Phacosoma) hataii, Isognomon (Isognomon) hataii, Leptochondria? 

p. 54
p. 36
p. 27
p. 37
p. 43
p. 29
p. 33
p. 50
p. 13
p. 57
p. 48
p. 55
p. 33
p. 34
p. 26
p. 51
p. 29
p. 27
p. 37
p. 50
p. 25
p. 41
p. 45
p. 28
p. 30
p. 39
p. 43
p. 14
p. 31
p. 55
p. 38
p. 35
p. 38
p. 51
p. 21
p. 20
p. 21
p. 20
p. 30
p. 31
p. 35
p. 22
p. 14
p. 18
p. 22
p. 24
p. 27
p. 30
p. 33
p. 33

Catalogue of type specimens of fossil Bivalvia
hataii, Pitar 
hataii, Serripes hataii, Thracia hataii, Zirfaea hatakeyamae, Chlamys (Chlamys) hayasakai, Aviculopecten hayasakai, Barbatia (Acar) hayasakai, Bassina (Callanatis) hizenense, Cardium (Cerastoderma) hizenensis, Nucula hokiensis, Macoma hokkaidoana, Trigonia hokkaidoensis, Anadara (Anadara) hokkaidoensis, Nucula (Acila) hokkaidoensis, Pectunculus hokkaidoensis, Pitar hokurikuensis, Patinopecten (Patinopecten) tokyoensis 
ibaragiensis, Patinopecten 
ibukii, Protocardia ichinohense, Trapezium (Neotrapezium) imanishii, Chlamys 
inconspicuus, Crassatellites 
infantile, Cardium (Trachycardium) inflata, Thyasira intermedius, Solen iriomotensis, Cucullaea iriomotensis, Glycymeris 
ishidae, Chlamys (Chlamys) ishidoense, Cardium 
ishidoensis, Lima (Limatula) ishidoensis, Modiola? ishidoensis, Nucula isoharense, Trapezium (Neotrapezium) isomatsuense, Trapezium 
itoigawae, Chlamys (Chlamys) iwakiensis, Neoburmesia 
iwashibaraensis, Anadara (Scapharca?) iwatensis, Anadara (Anadara) iwatensis, Spisula 
iwatensis, Yoldia 
iwatonoensis, Anadara (Anadara) izuensis, Pecten (Decadopecten) izumo, Panomya 
izurense, Gloripallium 
japonica, Clementia japonica, Cyclina (Cyclina) japonica, Myophoria japonica, Serripes japonica, Thetironia affinis var japonica, Veniella? jobanicum, Trapezium 
kagaensis, Mizuhopecten kimurai

p. 46
p. 49
p. 52
p. 57
p. 22
p. 18
p. 19
p. 19
p. 20
p. 39
p. 35
p. 54
p. 14
p. 39
p. 45
p. 46
p. 42
p. 41
p. 48
p. 53
p. 22
p. 25
p. 20
p. 53
p. 50
p. 27
p. 32
p. 22
p. 20
p. 34
p. 37
p. 39
p. 53
p. 53
p. 22
p. 38
p. 14
p. 14
p. 50
p. 57
p. 14
p. 44
p. 41
p. 31
p. 23
p. 28
p. 38
p. 49
p. 52
p. 56
p. 54
p. 37

kahoensis, Macrocallista kahoensis, Ostrea kakehataensis, Anadara (Anadara) kakisakiensis, Pecten (Pecten) kamayasikiensis, Thracia kanezawaensis, Barbatia 
kannoi, Dosinia (Kaneharaia) kanomatazawaensis, Panope karatsuensis, Nucula karihaensis, Nuculana (Nuculana) katsumatai, Cardita kazusaensis, Pholadomya kesenensis, Crenipecten kidoensis, Thracia kiiensis, Acila (Acila) kikaiensis, Pecten (Aequipecten) kikaizimana, Arca (Arca) kikaizimana, Tellina kimurai, Nuculites 
kintaichiensis, Patinopecten yamasakii kishimaense, Cardium (Cerastoderma?) kissyuensis, Schizothaerus nuttalli kitahukuokaensis, Mya arenaria
kitakamiensis, Annuliconcha kitamurai, Chlamys kitamurai, Codakia kobiyamai, Patinopecten kogachiensis, Anadara (Hataiarca) kokozuraensis, Nucula kondoi, Cardita 
konnoi, Saccella kosuiensis, Lucinopsis kotakae, Chlamys (Chlamys) kotakai, Cardiomya (Cardiomya) kotakai, Corbicula (Corbicula) kotoi, Pitar kumanodoensis, Chlamys kumasoana, Lima amaxensis var 
kunigamiensis, Lucina 
kuntsuipinensis, Tellina 
kurodae, Ezocallista kuroishiensis, Fortipecten kurosawaensis, Thracia kurosawai, Entomonotis kurosedaniensis, Anadara (Anadara) kyushuensis, Corbula (Cunaecorbula) kyushuensis, Pecten kyushuensis, Pitaria leguminoides, Cultellus? limopsis, Lima (Limea) lunaeformis, Ostrea makiyamae, Clinocardium asagaiense 

103
p. 35
p. 40
p. 14
p. 44
p. 52
p. 19
p. 30
p. 41
p. 39
p. 39
p. 20
p. 46
p. 26
p. 52
p. 13
p. 44
p. 16
p. 51
p. 40
p. 42
p. 21
p. 49
p. 37
p. 16
p. 22
p. 24
p. 42
p. 14
p. 39
p. 20
p. 49
p. 35
p. 22
p. 20
p. 24
p. 46
p. 22
p. 34
p. 34
p. 51
p. 30
p. 30
p. 52
p. 30
p. 14
p. 25
p. 44
p. 47
p. 27
p. 34
p. 40
p. 24

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makiyamai, Anadara manchuriensis, Ctenodonta masagoi, Polynemamussium 
masudai, Anadara (Scapharca) matsukuchiensis, Nuculana (Nuculana) matsumoriensis, Miyagipecten matsumotoi, Pitaria matsunagiensis, Aequipecten matsunoi, Chlamys (Chlamys) matsuraensis, Crassatellites matsuraensis, Macrocallista matsushimaensis, Teredo 
matumoriensis, Pecten (Patinopecten) kimuraip 
maxima, Tellina miikense, Cardium millegrana, Venericardia millepustulata, Taras 
minoensis, Aviculopecten 
minor, Astarte 
mirabilis, Cyrena 
miurensis, Arca (Arca) miyagiensis, Halicardia miyagiensis, Pecten (Chlamys) kagamianus miyakoensis, Astarte miyamoriensis, Strebrochondria miyamotoi, Pholadomya? 
moniwaensis, Patinopecten kagamianus 
monstrosa, Pitar sendaica 
multistriata, Daonella kotoi var muraii, Serripes murataensis, Mizuhopecten paraplebejusmuroensis, Solemya (Acharax) nagahamaensis, Tapes nagakoensis, Spisula (Mactromeris) polynyma naganoensis, Anadara (Anadara) naganoensis, Patinopecten (Kotorapecten) nagaoi, Batissa nagaoi, Chlamys (Chlamys) nagaoi, Nicaniella (Trautscholdia) nagaoi, Noetia 
nagaoi, Venericardia 
nakajimai, Patinopecten 
nakatonbetsuensis, Patinopecten (Patinopecten) yessoensis
nakayamaensis, Mactra 
nakosoensis, Clementia (Clementia) nakosoensis, Glycymeris nakosoensis, Patinopecten kimurai nanakitaensis, Pecten (Swiftopecten) naomiae, Glans nigamiensis, Serripes makiyamai niikappuensis, Chlamys (Chlamys) ninohensis, Meretrix meretrix

p. 14
p. 27
p. 48
p. 15
p. 40
p. 36
p. 47
p. 13
p. 22
p. 26
p. 36
p. 52
p. 44
p. 51
p. 21
p. 55
p. 51
p. 18
p. 17
p. 28
p. 16
p. 33
p. 44
p. 17
p. 51
p. 46
p. 42
p. 46
p. 29
p. 49
p. 37
p. 50
p. 51
p. 50
p. 15
p. 42
p. 19
p. 23
p. 38
p. 38
p. 55
p. 42
p. 43
p. 36
p. 23
p. 32
p. 42
p. 44
p. 31
p. 49
p. 23
p. 36

ninohensis, Patinopecten yamasakii nipponica, Cucullaea nipponica, Thyasira bisecta var nipponicus, Inoceramus concentricus var nipponicus, Leukomoides 
nishizawaensis, Thracidora 
nodai, Cyclina? noharai, Pododesmus (Monia) nomurae, Panopea 
nomurai, "Clinocardium" nomurai, Placopecten 
nomurai, Trifaricardium 
notoensis, Tellina 
nozokiensis, Glycymeris nuibetsuensis, Crenella (Megacrenella) numaensis, Barbatia (Acar) obessa, Anadara (Tegillarca) obsoleta, Trigonarca? obsoletiformis, Parallelodon 
obsoletoradiata, Rochefortia ochii, Peronidia ogachiensis, Palaeoneilo 
ogasawarai, Yabepecten ogawaensis, Mytilus ogawaraensis, Glycymeris ohiroi, Anisocorbula ohiroi, Paphia (Paphia) euglypta oidensis, Chlamys 
okadae, Myadora okinawaensis, Dosinia (Phacosoma) okinawaensis, Oxyperas okinawazimana, Codakia (Jagonia) okinoshimaensis, Venericardia omagariense, Clinocardium 
omagariensis, Nucula (Ennucula) omaruensis, Anadara (Scapharca) ommaensis, Nipponopagia ommaensis, Venericardia (Megacardita) onukii, Acanthopecten 
onukii, Aviculopecten? onukii, Venericardia oshimaensis, Glycymeris 
otsukai, Cyrena 
otukae, Chlamys 
otukai, Lithophaga otukai, Lucinoma 
owadaensis, Margaritifera pacificus, Pseudogrammatodon pedalionoides, Inoceramus 
philippianus, Azorius 
pitogoensis, Gari (Gari) planicostulatus, Pecten (Patinopecten) 

p. 43
p. 27
p. 53
p. 33
p. 33
p. 53
p. 28
p. 48
p. 41
p. 24
p. 47
p. 54
p. 51
p. 32
p. 26
p. 19
p. 15
p. 54
p. 41
p. 49
p. 46
p. 40
p. 57
p. 38
p. 32
p. 16
p. 41
p. 23
p. 37
p. 30
p. 40
p. 24
p. 55
p. 24
p. 39
p. 15
p. 38
p. 55
p. 13
p. 18
p. 56
p. 32
p. 28
p. 23
p. 34
p. 35
p. 36
p. 48
p. 33
p. 18
p. 31
p. 44

Catalogue of type specimens of fossil Bivalvia
ponderosa, Cyrena (Batissa) praenipponica, Ennucula 
problematica, Diplodonta? prototenuilirata, Tellina 
pseudomeretrix, Meretrix 
pseudoplana, Callista 
pseudosubcrenata, Anadara (Hataiarca) pseudotruncata, Anomia pulchellum, Periploma 
pustulosa, Trigonia quadrata, Thyasira 
quadriangulata, Venericardia radiatostriata, Cyrena rectangularis, Conularia 
reticularis, Aviculopecten rhynconelloides, Glycymeris 
ryukyuensis, Venericardia sachalinensis, Corbicula (Corbicula) sachalinensis, Pectunculus saikaiensis, Saccella saikawai, Caryocorbula sakamizuensis, Arca sakitoensis, Ostrea sakitoensis, Pecten (Chlamys) sakoi, Limatula sanchuensis, Corbicula (Veloritina?) sanchuensis, Nuculana sanchuensis, Perna sasakii, Aviculopecten satoi, Parapholas scabricostulata, Tellina scaphoides, Yoldia sekiyaensis, Callista 
sekiyaensis, Psammocola semipolita, Codakia 
setsukoae, Chlamys (Chlamys) shimizui, Corbicula (Corbicula) shimotokuraense, Vasticardium 
shimotsukensis, Paphia shimotsukensis, Spisula shinanoensis, Arca shinanoensis, Astarte shinanoensis, Gervillia 
shiobarensis, "Pitar" 
shiobarensis, Serripes shirahamaensis, Pecten (Pecten) shiratoriensis, Rexithaerus 
shiroiensis, Cyrena shiroyamaensis, Nucula (Nucula) shitakaraensis, Chlamys shitakaraensis, Crenella (Megacrenella) shitakaraensis, Solen 

p. 28
p. 30
p. 29
p. 52
p. 36
p. 19
p. 15
p. 16
p. 45
p. 54
p. 53
p. 56
p. 28
p. 24
p. 18
p. 32
p. 56
p. 25
p. 45
p. 49
p. 21
p. 17
p. 40
p. 44
p. 34
p. 25
p. 40
p. 46
p. 18
p. 41
p. 52
p. 57
p. 20
p. 48
p. 24
p. 23
p. 25
p. 55
p. 41
p. 51
p. 17
p. 18
p. 31
p. 47
p. 50
p. 44
p. 48
p. 29
p. 39
p. 23
p. 26
p. 50

shitakaraensis, Thracia (Thracia) shizuokaensis, Anadara (Scapharca?) sinelnikovae, Septifer sokeishiensis, Arca (Arca) spinosus, Acanthopecten 
striatocostata, Crenella subexcavata, Thyasira 
subfornicata, Crenella submaximus, Angulus maximus
subnipponica, Venericardia 
subomalioides, Astarte subpectiniformis, Glycymeris 
subquadrata, Daonella densisulcata var 
subsenecta, Astarte subsquamatus, Pecten (Pecten) subtumida, Corbula (Corbula) sulcataroides, Mactra 
sunakozakaensis, Crassostrea sunakozakaensis, Pitar 
sunakozakaensis, Pteria 
suzuensis, Myadora 
suzuensis, Paphia (Paphia) 
taiwanensis, Batissa 
taiwanensis, Corbula (Corbula) taiwanensis, Venus (Chione) taiwanica, Anadara (Scapharca) taiwanus, Pecten (Patinopecten) takagii, Callista chinensis takahashii, Ctenodonta takanabensis, Anadara (Scapharca) takaoensis, Arca (Arca) takasago, Corbicula 
takashimaensis, Pitaria takasimensis, Pholadomya takayamai, Anadara (Hataiarca) takiensis, Brachidontes tamurae, Chlamys 
tanakuraensis, Anadara (Anadara) tatunokutiensis, Dosinia tayamai, Pecten (Decadopecten) teshimaense, Vasticardium 
tobai, Myophoria subelegans var tochigiensis, Meretrix 
tokunagai, Thyasira 
tomiyaensis, Crenella tomiyasui, Pedalion 
tonohamanus, Loripes 
torii, Cardium (Nemocardium) tosaensis, Anadara (Tosarca) tosana, Lioconcha 
tosanus, Crassatellites 
toyajoanus, Inoceramus balticus var 

105
p. 53
p. 15
p. 49
p. 17
p. 13
p. 26
p. 53
p. 26
p. 16
p. 56
p. 18
p. 32
p. 29
p. 18
p. 45
p. 25
p. 36
p. 26
p. 47
p. 48
p. 37
p. 41
p. 19
p. 25
p. 56
p. 15
p. 45
p. 19
p. 27
p. 15
p. 17
p. 25
p. 47
p. 46
p. 15
p. 19
p. 23
p. 15
p. 30
p. 45
p. 55
p. 38
p. 36
p. 53
p. 26
p. 45
p. 34
p. 21
p. 16
p. 34
p. 26
p. 33

106

Shin’ichi Sato, Tomoki Chiba, Takaki Yamanaka, Jun Nemoto, Masanori Shimamoto and Takashi Matsubara

toyomaensis, Saccella confusa transversa, Arca (Noetia) pondaungensis var triangulus, Circe (Circe) tricarinata, Tellina 
tsudae, Patinopecten poculum 
tsudai, Anadara (Anadara) tsurushizakiensis, Barbatia (Pugilarca) tuberculosa, Plicatula uchigoensis, Nuttallia uetsukiensis, Barbatia (Barbatia) umbonata, Cyclina (Cyclina) umedairensis, Tellina 
wakuyaensis, Placopecten wanizakiensis, Modiolus yabei, Barbatia (Pugilarca) yabei, Crassatellites yabei, Inoceramus 
yabei, Pecten (?Amussiopecten) yabei, Pseudoactinodontophora yabei, Venus (Chione) 

p. 49
p. 17
p. 23
p. 52
p. 42
p. 16
p. 19
p. 48
p. 40
p. 19
p. 28
p. 52
p. 47
p. 37
p. 19
p. 26
p. 33
p. 45
p. 48
p. 56

yaegashii, Horiopleura 
p. 33
yaeyamaensis, Fimbria 
p. 30
yamadai, Macoma 
p. 35
yamaguchii, Glycymeris 
p. 32
yasudae, Polynemamussium 
p. 48
yasuhiroi, Modiolus (Modiolatus) p. 37
yatsuoensis, Anadara (Hataiarca) 
p. 16
yokoyamae, Patinopecten (Patinopecten) yessoensis  p. 43
yokoyamai, Arca (Barbatia) 
p. 17
yokoyamai, Grammatodon 
p. 32
yokoyamai, Ostrea (Ostrea) 
p. 40
yokoyamai, Pitaria 
p. 47
yokoyamai, Trigonia 
p. 54
yokoyamai, Volsella 
p. 56
yoshidai, Nemocardium (Keenaea) 
p. 38
yoshidai, Venericardia 
p. 56
yubaensis, Angulararca 
p. 16
yubarensis, Pecten (Propeamussium) cowperi var p. 43
yudaensis, Patinopecten kimurai 
p. 42
zyonoensis, Tellina venulosa 
p. 52