ᕝ༤⣖せࠉྕࠉ㸸ⲨᕝỈ⣔㧗㯇ᕝᨭὶࡢᯘ㐨⬥ࡢ‪ỈỈ⁀ࡾ࡛ࡢࢺ࢘࢟ࣙ࢘ࢧࣥࢩࣙ࢘࢘࢜ࡢ⦾Ṫ⾜ື┿㔝༤࣭἟⃝㤶⧊࣭἟⃝ࠉᾴ࣭἟⃝ࠉⅉ࣭἟⃝Ὧᚿ࣭┿㔝ᶞᏊ㸦ᇛす኱Ꮫ㧗㯇ᕝ࠿ࢃ࣮࢞ࣝ㸧࣭⸨⏣ᏹஅ㸦ᕝࡢ༤≀㤋㸧ࡣࡌࡵ࡟ࢺ ࢘ ࢟ ࣙ ࢘ ࢧ ࣥ ࢩ ࣙ ࢘ ࢘ ࢜ Hynobius
tokyoensisࡣࠊ⚟ᓥ┴ࡢኴᖹὒᓊࡢ୍㒊ࡢᆅᇦࠊ⩌㤿┴ࢆ㝖ࡃ㛵ᮾᆅ᪉ࡢ㸯㒔㸳┴ࡢẚ㍑ⓗ⊃࠸⠊ᅖ࡟⏕ᜥࡋ࡚࠸ࡿ㸦ⲡ㔝㸧ࠋᇸ⋢┴࡛ࡣࠊ⛛∗͌ධ㛫͌ẚ௻͌኱㔛ᆅ᪉ࡢపᒣᖏ࠿ࡽྎᆅ͌ୣ㝠ᖏ࡟࠿ࡅ࡚⏕ᜥࡋ࡚࠸ࡿ㸦ᇸ⋢┴⎔ቃ㒊ࡳ࡝ࡾ⮬↛ㄢ㸧ࠋࢺ࢘࢟ࣙ࢘ࢧࣥࢩࣙ࢘࢘࢜ࡣࠊᖹ㔝㒊࡜ᒣᆅࡀࡪࡘ࠿ࡿᶆ㧗P⛬ᗘࡲ࡛ࡢୣ㝠ᆅᖏ࡟⏕ᜥࡋࠊᒣ㛫ࡢỈ⏣ࡸ‪ࡁỈࡢ⁀ࡲࡗࡓỈ⁀ࡾ࡜ࠊࡑࡢ࿘
ᅖࡢ᳃ᯘࡢᯘᗋࡀ୺せ࡞⦾Ṫࡸ⏕άࡢሙᡤ࡟࡞ࡗ࡚࠸ࡿ㸦ⲡ㔝㸧ࠋࢺ࢘࢟ࣙ࢘ࢧࣥࢩࣙ࢘࢘࢜ࡣࠊ㸯㢌ࡢ࣓ࢫࡀ㸯ᑐࡢࢡࣟ࣡ࢵࢧࣥᆺࡢ༸ᄞࢆỈ୰ࡢᯤࢀᯞࠊⴠࡕⴥࠊ▼➼࡟⏘ࡳࡘࡅࠊ࢜ࢫࡀᨺ⢭ࡋ࡚యእཷ⢭ࡀ⾜ࢃࢀࡿࠋࡋ࠿ࡋࠊ⏘༸⾜ືࡣࠊኪ㛫⾜ࢃࢀࠊࡲࡓࠊほᐹࡋࡸࡍ࠸ࡼ࠺࡞⃈ࢇࡔỈ୰࡛⾜ࢃࢀࡿ࡜ࡣ㝈ࡽ࡞࠸ࡓࡵ⏘༸⾜ືࡢほᐹሗ࿌ࡣᑡ࡞࠸ࠋᮏ✏࡛ࡣࠊ㧗㯇ᕝᨭὶࡢᯘ㐨⬥ࡢ‪ỈỈ⁀ࡾࡢⴠࡕⴥࡢୗ࡛⾜ࢃࢀࡓ⦾Ṫ⾜ືࢆグ㘓ࡍࡿࠋㄪᰝᆅⅬࡢᴫἣ࡜ㄪᰝࡢ᪉ἲࠉձࠉㄪᰝᆅⅬࡢᴫἣ᪥㧗ᕷෆࡢ㧗㯇ᕝᨭὶ࡟ἢࡗࡓᯘ㐨ࡢᶆ㧗⣙PࡢᆅⅬ࡛グ㘓ࡋࡓࠋ‪Ỉࡢᮾഃࡀ㧗ࡃ࡞ࡗ࡚࠾ࡾࠊᶆ㧗P⛬ᗘࡢ᳜ᯘ࡟ࡼࡿ㔪ⴥᶞᯘ࡛ࠊⱝ࠸ᗈⴥᶞࡶ࠶ࡿࠋ‪Ỉࡣࠊ⣙P࡟ࢃࡓࡗ࡚Ỉ⁀ࡾ࡜Ỉ㊰࡟࡞ࡗ࡚࠸ࡿ㸦෗┿㸯㸧ࠋ ᭱ ࡶ 㧗 ఩ ࡟ ఩ ⨨ ࡍ ࡿ Ỉ ⁀ ࡾ FPFP ࡣẚ㍑ⓗ῝࠸Ỉ῝࡛ࡑࡇ࠿ࡽP⛬ᗘ㠀ᖖ࡟ὸ࠸Ỉ㊰࡟ࡘ࡞ࡀࡾࠊFPFP⛬ᗘࡢỈ῝ࡢὸ࠸Ỉ⁀ࡾࡀ㸱ࡘ⧅ࡀࡗ࡚࠸ࡿࠋࡑࡢୗὶࡣ⣙Pࡢ㠀ᖖ࡟ὸ࠸Ỉ㊰࡜࡞ࡗ࡚࠸ࡿࠋ㝆㞵࡟ࡼࡗ࡚ᯘ㐨ࢆὶࢀࡓ㞵ỈࡣࠊỈ⁀ࡾࡸỈ㊰ࡢ཯ᑐഃࢆὶࢀࡿࡓࡵࡇࡢỈ⁀ࡾࡸỈ㊰࡟኱ࡁࡃᙳ㡪ࡋ࡞࠸ࠋᩘᖺ㛫ࠊᮏᆅⅬ࡛ࢺ࢘࢟ࣙ࢘ࢧࣥࢩࣙ࢘࢘࢜ࡢ༸ᄞࢆほᐹࡋ࡚࠸ࡓࠋղ ㄪᰝ᪉ἲᖺ᭶ᮎ࠿ࡽ᭶ᮎࡲ࡛ᮏỈ㊰࡛⏘༸ࡉࢀ ࡓ ༸ ᄞ ࢆ ᩘ ࠼ ࡓࠋ ࡲ ࡓࠊ᭶㸦ࠊࠊࠊࠊࠊࠊࠊࠊࠊࠊ᪥㸧ࡢኪ㛫 㹼  ࡟㉥Ⰽࣛ࢖ࢺࢆ⏝࠸࡚ࢺ࢘࢟ࣙ࢘ࢧࣥࢩࣙ࢘࢘࢜ࡢ⦾Ṫ⾜ືࢆほᐹࡋࡓࠋ⦾Ṫ⾜ື͌⏘༸⾜ືࡣࠊ࢜ࣜࣥࣃࢫ20'(0O
0DUN ,,,ࢆ ౑ ⏝ ࡋ ࡚5$:ᙧ ᘧ ࡛ ᧜ ᙳ ࡋࠊ2O\PSXV:RUNVSDFH࡛࣮࢝ࣛ໬ࡋࡓࠋ⤖ᯝᖺࡢ༸ᄞࡢグ㘓ᩘࢆ⾲㸯࡟♧ࡍࠋ࡞࠾༸ᄞ☜ㄆ᪥ࡣࠊ⏘༸᪥࡜୍⮴ࡋ࡞࠸ࠋᖺࡢᮏᆅⅬࡢ⏘༸༸ᄞᩘࡢࣆ࣮ࢡࡣ᭶ࠊࠊ᪥࡛࠶ࡗࡓࠋࡲࡓ⾲୰࡛ࠊ᭶᪥࡟ᑐ࡜࠶ࡿࡢࡣࠊ༸ᄞ㸯ಶࡀᾘኻࡋ࢔ࣛ࢖ࢢ࣐➼ࡢ⿕ᐖ࡟࠶ࡗࡓ࡜⪃࠼࡚࠸ࡿࠋࡲࡓࠊ᭶᪥࡟㢌㒊࡟㗦࠸∎࡛ษࡽࢀࡓࡼ࠺࡞യࡀ࠶ࡿࢺ෗┿㸯㸸⏘༸⾜ືࢆほᐹࡋࡓỈ⁀ࡾࡢᵝᏊ㸫  㸫┿㔝ࠉ༤࣭἟⃝㤶⧊࣭἟⃝ࠉᾴ࣭἟⃝ࠉⅉ࣭἟⃝Ὧᚿ࣭┿㔝ᶞᏊ࣭⸨⏣ᏹஅⲨᕝỈ⣔㧗㯇ᕝᨭὶࡢᯘ㐨⬥ࡢ‪ỈỈ⁀ࡾ࡛ࡢࢺ࢘࢟ࣙ࢘ࢧࣥࢩࣙ࢘࢘࢜ࡢ⦾Ṫ⾜ືࢩࣙ࢘࢘࢜ࡀ༸ᄞ࠿ࡽ㞳ࢀࠊ⏘༸⾜ືࡀ⤊஢ࡋࡓ㸦෗┿㸵㸧ࠋศ㛫⛬ᗘࡢ⏘༸⾜ື࡛࠶ࡗࡓ㸦⾲㸰㸧ࠋ⏘ࡳࡘࡅࡽࢀࡓ┤ᚋࡢ༸ᄞࡣ⭾₶࡛ࡁ࡚࠸࡞࠿ࡗࡓࡀࠊ⩣᭶᪥᫬ศࡢ᫬Ⅼ࡛ࡶ༸ᄞࡣ⭾₶ࡀ㏵୰࡛࠶ࡗࡓࡀࠊᚋ᪥᏶඲࡟⭾₶ࡋࡓࠋࡲࡓࠊᮏ༸ᄞࡣⴠࡕⴥ࡟௜╔ࡋ࡚࠸ࡓ㸦෗┿㸶㸧ࠋ⾲㸯ࠉᮏほᐹᆅⅬ࡟࠾ࡅࡿࢺ࢘࢟ࣙ࢘ࢧࣥࢩࣙ࢘࢘࢜ࡢ༸ᄞᩘ㸦ᑐ㸧ᖺ᭶᪥᭶᪥᭶᪥᭶᪥᭶᪥᭶᪥᭶᪥᭶᪥᭶᪥᭶᪥᭶᪥᭶᪥༸ᄞ㸦ᑐ㸧࢘࢟ࣙ࢘ࢧࣥࢩࣙ࢘࢘࢜ࡢ࢜ࢫࡢᡂయ㸰㢌ࡢṚ㧁ࢆ☜ㄆࡋࡓࡀࠊ࢔ࣛ࢖ࢢ࣐➼࡟ᤕ⋓ࡉࢀࡑ࠺࡟࡞ࡗࡓ㝿࡟㢌㒊࡟ᦆയࢆཷࡅ⮴࿨യ࡟⮳ࡗࡓࡇ࡜ࡀ᥎ ࡉࢀࡿ㸦෗┿㸰㸧ࠋᮏỈ㊰௜㏆࡛࢔ࣛ࢖ࢢ࣐ࡢ㊊㊧ࡣ☜ㄆ࡛ࡁ࡞࠿ࡗࡓࡀࠊᮏỈ㊰࠿ࡽPୖὶࡢ㇂ᡞ࡟タ⨨ࡋ࡚࠶ࡿࢺࣞ࢖࣓ࣝ࢝ࣛ࡟᭶᪥᫬ศ࡜᭶᪥᫬ศ࡟࢔ࣛ࢖ࢢ࣐ࡀグ㘓ࡉࢀ࡚࠸ࡓࠋ
⏘༸⾜ືࡀほᐹ࡛ࡁࡓࡢࡣࠊᅇࡢほᐹࡢ࠺ࡕ㸯ᅇࡢࡳ࡛࠶ࡗࡓࠋࡑࢀ௨እࡢᅇࡣࠊ࣓ࢫࢆỈ୰࡛ᚅࡘᩘ㢌ࡢ࢜ࢫࡢࢺ࢘࢟ࣙ࢘ࢧࣥࢩࣙ࢘࢘࢜ࢆほᐹࡍࡿࡢࡳ࡛࠶ࡗࡓࠋ࢜ࢫࡣỈ୰ࡢᯞ࡟ࡘ࠿ࡲࡗ࡚࠸ࡓࡾࠊⴠࡕⴥࡢୗࡸỈ୰࡛㟼Ṇࡋ࡚࠸ࡓࡾࠊ⾜ື㔞ࡣẚ㍑ⓗᑡ࡞࠿ࡗࡓࡀࠊ㸱ࡘࡢ⧅ࡀࡗࡓỈ⁀ࡾࡢ㛫ࢆ᫬㛫ࢆ࠿ࡅ࡚ࠊ⛣ືࡍࡿࢺ࢘࢟ࣙ࢘ࢧࣥࢩࣙ࢘࢘࢜ࡢᵝᏊࢆほᐹࡋࡓࠋ⏘༸๓ࡢ⭡ࡢ኱ࡁ࡞࣓ࢫࡢࢺ࢘࢟ࣙ࢘ࢧࣥࢩࣙ࢘࢘࢜ࢆほᐹ࡛ࡁࡓࡢࡣୗグࡢ⏘༸⾜ືࢆほᐹࡋࡓ࡜ࡁࡢࡳ࡛࠶ࡗࡓࠋ⏘༸⾜ືࢆ၏୍ほᐹ࡛ࡁࡓࡢࡣࠊ᭶᪥࡛࠶ࡗࡓࠋ᭶᪥᫬ศ࡟⭡㒊ࡢ኱ࡁ࡞㸯㢌ࡢ࣓ࢫࡢࢺ࢘࢟ࣙ࢘ࢧࣥࢩࣙ࢘࢘࢜ࢆ☜ㄆࡋࡓ㸦෗┿㸱㸧ࠋࡑࡢ㝿ࠊ㸱㢌ࡢ࢜ࢫࡣỈ୰࡟࠸ࡓࠋ᫬ศ࡟ࡇࡢ࣓ࢫࡀỈ୰ࠊ㝣ୖ࡜┒ࢇ࡟⛣ືࡋጞࡵࡓ ෗┿㸲 ࠋ᫬ศࡇࢁ෗┿㸯ࡢỈ⁀ࡾࡢ୰ኸ㒊ศ࡛᭱ࡶỈ῝ࡀὸ࠸ሙᡤࡢⴠࡕⴥࡢୗ࡟࣓ࢫ࡜」ᩘࡢ࢜ࢫࡀࡶ
ࡄࡾࠊ⏘༸⾜ືࢆ㛤ጞࡋࡓ㸦෗┿㸳 ࠋ㸯㢌ࡢ࣓ࢫ࡜」ᩘࡢ࢜ࢫࡀ⤡ࡳྜ࠺⏘༸⾜ືࢆほᐹࡋࡓࠋ㸯㢌ࡢࢺ࢘࢟ࣙ࢘ࢧࣥࢩࣙ࢘࢘࢜ࡣ
༸ᄞࢆᘬࡁฟࡋ࡚࠸ࡿᵝ࡟ࡶぢ࠼ࡓ㸦෗┿㸴㸧ࠋ᫬ศ࡟ࡑࢀࡒࢀࡢࢺ࢘࢟ࣙ࢘ࢧࣥ⪃ᐹᮏᆅⅬ࡛ࠊ⏘༸⾜ືࢆほᐹࡋ࡚࠸ࡓᮇ㛫୰ᑐࡢ༸ᄞࢆ☜ㄆࡋࡓࡀࠊ⏘༸⾜ືࢆほᐹ࡛ࡁࡓࡢࡣ㸯ᅇ࡛࠶ࡗࡓࠋࡑࡢࡓࡵࠊ⏘༸⾜ືࡣࠊᮏㄪᰝࡢほᐹ᫬㛫௨እ࡛⾜ࢃࢀࡓ࡜⪃࠼ࡿࠋࡲࡓࠊ㏆ᖺࢺ࢘࢟ࣙ࢘ࢧࣥࢩࣙ࢘࢘࢜ࡣ
⏕ᜥ⎔ቃࡢᝏ໬࡞࡝࡛ᛴ⃭࡟ಶయᩘࡀῶᑡࡋ
࡚࠸ࡿࡀࠊㄪᰝᆅⅬࡣⰋዲ࡞⏕ᜥ⎔ቃࡀಖࡓࢀ࡚࠸ࡿ࡜⪃࠼ࡽࢀࡿࠋ
ࢺ࢘࢟ࣙ࢘ࢧࣥࢩࣙ࢘࢘࢜ࡢ⏕ែ࡟ࡘ࠸࡚ࡣ⦾Ṫᖺ㱋ࡸ༸ࡢ⏕Ꮡ⋡ࡢඛ⾜◊✲࡜ࡋ࡚
Kusano㸦  ࡀሗ࿌ࡋ࡚࠸ࡿࠋ⏘༸⾜ື࡟ࡘ࠸࡚ࡣࠊⲡ㔝㸦㸧࡟࡚ヲ⣽࡟グࡉࢀ࡚࠸ࡿࡀࠊᯤࢀᯞ࡟⏘༸ࡍࡿᵝᏊ㸦㛵㸪㸧ࡀ෗┿࡟࡚⤂௓ࡉࢀ࡚࠸ࡿࠋࡋ࠿ࡋࠊ࠸ࡎࢀࡶᮏ✏ࡢᵝ࡟ὸ࠸Ỉ῝࡛ⴠࡕⴥ࡟⏘ࡳࡘࡅࡿᵝᏊࡢሗ࿌࡛ࡣ࡞࠿ࡗࡓࠋࡲࡓࠊ኱ࡁ࡞㞟ᅋ࡛ࡢ⏘༸⾜ືࡀࡳࡽࢀࡿྠᒓࡢࢡࣟࢧࣥࢩࣙ࢘ ⮻⏣ ࣭⮻⏣ ࣭⸨⏣ ࡸ࢜࢜࢖ࢱࢧࣥࢩࣙ࢘࢘࢜㸦ᮎྜྷ࣭୵㛫㸪㸧࡞࡝࡟ࡳࡽࢀࡿࡼ࠺࡞኱ࡁ࡞㞟ᅋࢆᙧᡂࡋࡓ⏘༸⾜ື࡛ࡣ࡞࠿ࡗࡓࠋࢡࣟࢧࣥࢩ࡛ࣙ࢘ࡣ࢜ࢫࡀ࣓ࢫࡢ⥲᤼ἥ⭍࠿ࡽ༸ᄞࢆᘬࡁฟࡍຓ⏘⾜ື ⮻⏣ ࡀ▱ࡽࢀ࡚࠸ࡿࡀࠊ௒ᅇࡢほᐹ࡛ࡣࡑࢀࡽࡋࡁ⾜ືࡀࡳࡽࢀࡓࡶࡢࡢࠊ☜ᐃࡍࡿ࡟⮳ࡽ࡞࠿ࡗࡓࠋࡲࡓࠊ࢜ࢫࡣ࣓ࢫࡢ⥲᤼ἥ⭡⭍ࢆ㏆࡙ࡅ࡚ᨺ⢭ࡍࡿ࠿ྰ࠿࡞࡝ࡣ☜ㄆ࡛ࡁ࡞࠿ࡗࡓࠋࡋ࠿ࡋࠊᮏ✏࡛ࡣࠊ⏘༸ࡉࢀࡓ༸ᄞ࡟᭱ᚋࡲ࡛ᐤࡾῧ࠺ࡼ࠺࡞ᙧ࡛࢜ࢫ࡜⪃࠼ࡽࢀࡿಶయࡢᑿࡀほᐹ࡛ࡁࡓࠋࡲࡓࠊᮏ✏ࡣࠊࢺ࢘࢟ࣙ࢘ࢧࣥࢩࣙ࢘࢘࢜ࡀὸ࠸Ỉ῝ࡢỈ⁀ࡾࡢⴠࡕⴥࡢ୰࡛⏘༸ࡀ࠾ࡇ࡞ࢃࢀࡓグ㘓࡜࡞ࡗࡓࠋࢺ࢘࢟ࣙ࢘ࢧࣥࢩࣙ࢘࢘࢜ࡀ࢔ࣛ࢖ࢢ࣐ࡢ⿕ᐖࢆཷࡅ࡚࠸ࡿࡇ࡜ࡣࡼࡃ▱ࡽࢀ࡚࠸ࡿࡀ㔠⏣࣭ຍ⸨࣭㔠⏣ ࠊᇸ⋢┴ෆ࡛ࡶ⿕ᐖࡀⓎ⏕ࡋ࡚࠸ࡿ㸦ᇸ⋢┴⎔ቃ㒊ࡳ࡝㸫  㸫ᕝ༤⣖せࠉྕࠉ㸸෗┿㸰㸸࢜ࢫࡢࢺ࢘࢟ࣙ࢘ࢧࣥࢩࣙ࢘࢘࢜ࡢṚ㧁෗┿㸴㸸࢜ࢫࡀ༸ᄞࢆᘬࡁฟࡋ࡚࠸ࡿᵝ࡟ぢ࠼ࡿ෗┿㸱㸸⏘༸⾜ືࡢ๓࡟㝣ୖࢆṌࡃ࣓ࢫ㸦ྑ㸧࡜Ỉ୰࡛ᚅࡘ࢜ࢫ㸦ᕥ㸧 ෗┿㸵㸸⏘༸⤊஢ ෗┿㸲㸸Ỉ୰࡛⏘༸ሙᡤࢆ᥈ࡍ࣓ࢫ㸦ୖ㸧࡜࢜ࢫ㸦ୗ㸧෗┿㸶㸸⩣᪥᪩ᮅࡢ༸ᄞࡢ≧ែ෗┿㸳㸸Ỉ῝ࡢὸ࠸ⴠࡕⴥࡢୗ࡛⏘༸ ࡾ⮬↛ㄢ 㸧ࠋᮏᆅⅬ࡛ࡶࢺ࢘࢟ࣙ࢘ࢧࣥࢩࣙ࢘࢘࢜ࡢᡂయࡸ༸ᄞࡀ࢔ࣛ࢖ࢢ࣐➼࡟ࡼࡿ⿕ᐖࢆཷࡅࡓ࡜⪃࠼ࡽࢀࡿࠋ≉࡟௒ᅇぢࡽࢀࡓࡼ࠺࡞ᡂయࡢᤕ㣗⿕ᐖࡣࠊᡂ㛗ࡀ㐜ࡃᑠつᶍ࡞ಶయ⩌࡛ୡ௦஺௦ࢆ⧞ࡾ㏉ࡍᑠᆺࢧࣥࢩࣙ࢘࢘࢜㢮࡟ࡣ኱ࡁ࡞ࢲ࣓࣮ࢪ࡜࡞ࡿࠋ௒ᚋᛴ⃭࡞ಶయᩘࡢῶᑡ࡟ࡣせὀព࡜⪃࠼ࡿࠋ୍᪉ࠊᮏᆅⅬࡣ୍⯡ⓗ࡟ࡣᯘ㐨⬥ࡢỈ⁀ࡾ࡟ࡋ࠿ぢ࠼࡞࠸ሙᡤࡢࡓࡵࠊᯘ㐨౑⏝⪅ࡀࢱࣂࢥࡢ྾࠸Ẇ➼ࢆᢞᲠࡋ࡚࠸ࡓࠋࡉࡽ࡟ࠊ㏆㸫  㸫┿㔝ࠉ༤࣭἟⃝㤶⧊࣭἟⃝ࠉᾴ࣭἟⃝ࠉⅉ࣭἟⃝Ὧᚿ࣭┿㔝ᶞᏊ࣭⸨⏣ᏹஅⲨᕝỈ⣔㧗㯇ᕝᨭὶࡢᯘ㐨⬥ࡢ‪ỈỈ⁀ࡾ࡛ࡢࢺ࢘࢟ࣙ࢘ࢧࣥࢩࣙ࢘࢘࢜ࡢ⦾Ṫ⾜ື⾲㸰ࠉࢺ࢘࢟ࣙ࢘ࢧࣥࢩࣙ࢘࢘࢜ࡢ⏘༸⾜ືᖺ㸱᭶᪥ࢺ࢘࢟ࣙ࢘ࢧࣥࢩࣙ࢘࢘࢜ࡢ⾜ື᫬ศ㸱㢌ࡢ࢜ࢫࡀỈ୰࡛ᚅᶵࠋ᫬ศ⭡㒊ࡢ኱ࡁ࡞㸯㢌ࡢᢪ༸࣓ࢫࢆỈ୰࡛☜ㄆࠋ᫬ศ࣓ࢫ࡜࢜ࢫࡀỈ୰࡛ࡺࡗࡃࡾ⛣ືࡋ࡞ࡀࡽᩘFPࡢ㊥㞳ࡲ࡛㏆ᐤࡿࠋ࣓ࢫࡶ࢜ࢫࡶỈ୰࡛⛣ືࡋ⥆ࡅࡿࠋ᫬ศ࣓ࢫࡀ㝣ୖ࡟⛣ືࡋࠊ㸯㢌ࡢ࢜ࢫࡣ࣓ࢫࢆ㏣࠸࠿ࡅࡿࠋ᫬ศ௚ࡢ㸰㢌ࡢ࢜ࢫࡣỈ୰࡛ᚅᶵࡋࡓࡲࡲࠋ᫬ศ࣓ࢫࡀỈ῝ࡢὸ࠸ⴠࡕⴥࡢୗ࡟₯ࡾ㎸ࡴࠋྠ᫬้㸱㢌ࡢ࢜ࢫࡣ㡰ḟ࣓ࢫࢆ㏣ࡗ࡚ⴠⴥࡢୗ࡟₯ࡾ㎸ࡴࠋ㸯㢌ࡢ࢜ࢫࡣ⭡࡜ᑿࢆ⣽࠿ࡃ᣺ືࡉࡏ࡚࠸ࡓࠋ࢜ࢫ࣭࣓ࢫ㸲㢌ࡀ⤡ࡳࡣࡌࡵࡿࠋ᫬ศ༸ᄞࡀฟ࡚࠸ࡿࡇ࡜ࢆ☜ㄆࡋࡓ㸦⏘༸㛤ጞ㸧ࠋ᫬ศ࢜ࢫ㸯㢌ࡀ༸ᄞࢆཱྀ࡛ဃ࠼࡚ᘬࡁฟࡍᵝᏊࢆ☜ㄆࠋ᫬ศ༸ᄞࢆ୰ᚰ࡟࢜ࢫ࣭࣓ࢫ㸲㢌ࡣ⤡ࡳྜ࠸⥆ࡅࡿࠋ᫬ศ㸯㢌ࡢ࢜ࢫࡀ⥲᤼ฟ⭍ࢆ༸ᄞ࡟ᢲࡋࡘࡅᑿࢆ⣽࠿ࡃ᣺ࡿࠋࡑࡢᚋࠊ༸ᄞ࠿ࡽ㞳ࢀࡣࡌࡵࡿࠋ᫬ศ༸ᄞ࡜㸰㢌ࡢᑿࢆ☜ㄆࠋ᫬ศ༸ᄞ࡜㸯㢌ࡢᑿࢆ☜ㄆࠋ᫬ศ඲࡚ࡢಶయࡀ༸ᄞ࠿ࡽ㞳ࢀࡿࠋᖺࡢ኱㞵࡟ࡼࡿ⿕ᐖ࡜ᯘ㐨ᩚഛ࡟ࡼࡾࠊỈ⁀ࡾࡸỈ㊰ࡢ≧ἣ࠿ࡽ⏕ᜥ⎔ቃࡢᝏ໬ࡀᠱᛕࡉࢀࡿࠋⰋዲ࡞ࢺ࢘࢟ࣙ࢘ࢧࣥࢩࣙ࢘࢘࢜ࡢ⏕ᜥᆅࠊ⏘༸ሙ࡜ࡋ࡚ࠊᮏᆅⅬࢆ⏕ᜥ⎔ቃಖ඲࡜࡜ࡶ࡟ኳᩛ࡛࠶ࡿ౵␎ⓗ࡞እ᮶✀࠿ࡽᏲࡿࡓࡵࡢఱࡽ࠿ࡢ᪉⟇ࡀᚲせ࡛࠶ࡿ࡜⪃࠼ࡿࠋᘬ⏝ᩥ⊩⸨⏣ᏹஅ㸦 ᭶ᒣᒣ㯄ࡢすᕝ⏫᭶ᒣἑ࡟࠾ࡅࡿࢡࣟࢧࣥࢩࣙ࢘࢘࢜⏘༸ࡢほᐹ஦౛ᐮἙỤᕝὶᇦ⮬↛ྐ◊✲㸦㸧㔠⏣ṇே͌ຍ⸨༟ஓ㸦㸧እ᮶⏕≀࢔ࣛ࢖ࢢ࣐࡟⬣࠿ࡉࢀࡿ∐⹸୧⏕㢮∐⹸୧᳇㢮Ꮫ఍ሗ㸦㸧㸸㔠⏣ṇே͌ᒣ㷂ᩥᬗ͌⚄ᒣዉ⏤Ꮚ  እ᮶⏕≀࢔ࣛ࢖ࢢ࣐ࡢᾘ໬⟶ෆᐜ≀࡜ࡋ࡚ぢࡘ࠿ࡗࡓࢺ࢘࢟ࣙ࢘ࢧࣥࢩࣙ࢘࢘࢜༸ᄞ∐⹸୧᳇㢮Ꮫ఍ሗ㸦㸧
Kusano, T.  Breeding and egg survival ofa population of a salamander, Hynobius
nebulosus tokyoensis TAGO. Res. Popul. Ecol.
21: 181-196.
Kusano, T.   Postmetamorphic growth,survival, and age at forst reproduction of thesalamander, Hynobius nebulosus tokyoensis
TAGO in relation to a consideration on theoptimal timing of first reproduction. Res.
Popul. Ecol. .

ⲡ㔝ࠉಖ㸦 ࢺ࢘࢟ࣙ࢘ࢧࣥࢩࣙ࢘࢘࢜᪥㧗ᩄ㝯㸦┘ಟ㸧᪥ᮏື≀኱ⓒ⛉ ➨㸳ᕳ୧⏕㢮͌∐⹸㢮͌㌾㦵㨶㢮ᖹซ♫ᮾிSSⲡ㔝ࠉಖ࣭ᕝୖὒ୍࣭ᚚᡭὙᮃ㸦⦅㸧  ࢺ࢘࢟ࣙ࢘ࢧࣥࢩࣙ࢘࢘࢜㸸㛗ᮇㄪᰝ࡛ศ࠿ࡗࡓಶయ⩌ࡢ⾶㏥࡜⤯⁛ᮾி㒔ከᦶᆅ༊࡟࠾ࡅࡿᖺᗘ⏕ᜥ≧ἣㄪᰝሗ࿌᭩ᇸ⋢┴⎔ቃ㒊ࡳ࡝ࡾ⮬↛ㄢ ⦅   ᇸ⋢┴ࣞࢵࢻࢹ࣮ࢱࣈࢵࢡື≀⦅ 㸦➨㸲∧㸧ᇸ⋢┴㛵ៅኴ㑻㸦㸧ᯇ஭ṇᩥ㸦┘ಟ㸧㔝እほᐹࡢࡓࡵࡢ᪥ᮏ⏘୧⏕㢮ᅗ㚷 ➨㸱∧⥳᭩ᡣᮾிSSᮎྜྷ㇏ᩥ࣭୵㛫◊அ㸦㸧࢜࢜࢖ࢱࢧࣥࢩࣙ࢘࢘࢜ࡢ⏘༸⾜ືᐑᓮ┴⥲ྜ༤≀㤋◊✲⣖せ㸦㸧㸸⮻⏣ࠉᘯ  ࢡࣟࢧࣥࢩࣙ࢘࢘࢜ࡢ⦾Ṫ⾜ືࠊ≉࡟㞝ࡢຓ⏘⾜ື࡟ࡘ࠸࡚∐⹸୧᳇㢮Ꮫ㞧ㄅ㸦㸧⮻⏣ࠉᘯ  ࢡࣟࢧࣥࢩࣙ࢘࢘࢜ࡢ⦾Ṫ⾜ື࡟࠾ࡅࡿಶయࡢἼࢆ㉳ࡇࡍ⾜ື࡜Ἴ࡬ࡢ཯ᛂ∐⹸୧᳇㢮Ꮫ㞧ㄅ㸦㸧㸫  㸫