ᕝ༤⣖せࠉྕࠉ㸸ᇸ⋢┴す㒊ᆅᇦ࡛ᤕ⋓ࡉࢀࡓ୰ᆺ့ங㢮ࡢእ㒊ᐤ⏕⹸ㄪᰝ㸦ᖺᗘ㸧㧗ᶫࠉᏲ࣭୕ゅோᏊ㸦ᇸ⋢་⛉኱Ꮫ㸧࣭⸨⏣ᏹஅ㸦ᕝࡢ༤≀㤋㸧ࡣࡌࡵ࡟ࠉ㔝⏕ື≀ࠊ࡜ࡃ࡟ᜏ ື≀࡛࠶ࡿ့ங㢮ࡣ⾜ື⠊ᅖࡀᗈࡃࠊே⋇ඹ㏻ឤᰁ⑕㸦ື≀⏤᮶ឤᰁ⑕㸧ࢆ፹௓ࡍࡿࢲࢽࡸࣀ࣑㢮࡞࡝ࠊᵝࠎ࡞⠇㊊ື≀ࡀᐤ⏕ࡋ࡚࠸ࡿࠋࠉᮏሗ࡛ࡣ᫖ᖺᗘࡢሗ࿌࡟⥆ࡁ㸦㧗ᶫ࡯࠿ࠊ㸧ࠊᇸ⋢┴ࡢす㒊ᆅᇦ࡛ᤕ⋓ࡉࢀࡓ࢔ࣛ࢖
ࢢ࣐㸦Procyon lotor㸧ࠊࢽ࣍ࣥ࢔ࢼࢢ࣐㸦Melesanakuma㸧ࠊࣁࢡࣅࢩࣥ㸦Paguma larvata㸧࠾ࡼࡧ໭㒊ᆅᇦ࡛ᤕ⋓ࡉࢀࡓ࣍ࣥࢻࢸࣥ㸦Martes
melampus㸧ࡢእ㒊ᐤ⏕⹸ࡢㄪᰝࢆ⾜ࡗࡓࠋࡲࡓẟ࿅ᒣ⏫࡛ᤕ⋓ࡉࢀࡓ࢔ࣛ࢖ࢢ࣐࡟ከᩘࡢࢶࢶ࣒࢞ࢩ࡜࣐ࢲࢽࡀᐤ⏕ࡋ࡚࠸ࡓࡓࡵࠊࡑࡢᤕ⋓ሙᡤ࿘㎶࡛ࡢࢶࢶ࣒࢞ࢩ࡜࣐ࢲࢽࡢ᳜⏕ୖࡢணഛⓗㄪᰝࡶేࡏ࡚⾜ࡗࡓࡢ࡛ሗ࿌ࡍࡿࠋㄪᰝ᪉ἲ 㸧እ㒊ᐤ⏕⹸ࡢ᥇㞟ࠉ᭷ᐖ㫽⋇㥑㝖࡜ࡋ࡚ࠊẟ࿅ᒣ⏫࡛ᤕ⋓ࡉࢀࡓ࢔ࣛ࢖ࢢ࣐࡜ࣁࢡࣅࢩࣥࡢ࠺ࡕࠊ୺࡟᫓࡜⛅࡟ᤕ⋓ࡉࢀࡓಶయࡢ୍㒊ࢆㄪᰝ࡟౪ ࡋ ࡓࠋ ㄪ ᰝ ࡣ ᠱ ᆶ ἲ ࢆ  㹼  ᪥ 㛫 ⥅ ⥆ࡋࠊỈࢆᙇࡗࡓ࢔࣑ࣝᶆ‽ྜ㔠ࣂࢵࢺ  ྕ㸦xxੈ , ㇂ཱྀ㔠ᒓᕤᴗ㸧࡟ⴠୗࡋࡓእ㒊ᐤ⏕⹸ࡣࡍ࡭࡚㸣࢚ࢳࣝ࢔ࣝࢥ࣮ࣝ࡟ಖᏑࡋࡓࠋࡲࡓᇸ⋢┴❧ᕝࡢ༤≀㤋࡟෭෾ಖᏑࡉࢀ࡚࠸ࡓಶయࡣࠊゎ෾ᚋ඲㌟ࢆ⫗║ⓗ࡟⢭ᰝࡋࠊ๓㏙ྠᵝ࡟እ㒊ᐤ⏕⹸ࢆ᥇㞟ࡋಖᏑࡋࡓࠋ࡞࠾ᇸ⋢┴໭㒊ࡢ⛛∗㒆ᑠ㮵㔝⏫୕ᒣ⏣அ㢌ᆅෆ࡛ᤕ⋓ࡉࢀࡓ࣍ࣥࢻࢸࣥ  㢌ࡣࠊ␾Ⓞ⨯ᝈಶయ࡛ࠊ㧗ᶫ๤〇ᗑ࡟ಖ⟶ࡉࢀ࡚࠸ࡓಶయࡢᥦ౪ࢆཷࡅࡓࠋ 㸧᳜⏕ୖࡢ࣐ࢲࢽ࡜ࢶࢶ࣒࢞ࢩࡢ᥇㞟ࠉẟ࿅ᒣ⏫ⴱ㈏࡜⠼࿴⏣࡛ᤕ⋓ࡉࢀࡓ࢔ࣛ࢖ࢢ࣐࡟ࠊᩘ༓ಶయࡢࢱࢸࢶࢶ࣒࢞ࢩࡢᐤ⏕ࡀࡳࡽࢀࡓࡢ࡛ࠊࡇࢀࡽࡢ࢔ࣛ࢖ࢢ࣐ࡢ⾜ື⠊ᅖࡢ࡝ࡇ࠿࡟ࢱࢸࢶࢶ࣒࢞ࢩࡢࢡࣛࢫࢱ࣮ࡀ࠶ࡿ࡜⪃࠼ࡽࢀࡓࠋࡑࢀࡺ࠼ணഛㄪᰝ࡜ࡋ࡚ࠊᤕ⋓ሙᡤࢆ୰ᚰ࡟ࠊ࢔ࣛ࢖ࢢ࣐ࡢ㏻ࡾ㐨࡜ᛮࢃࢀࡿሙᡤ࡛ࠊ㯮ᕸἲ࡟ࡼࡾࢶࢶ࣒࢞ࢩࡢ᥇㞟ࢆヨࡳࡓࠋࡇࢀࡣ㯮ᕸ㸦x੉㸧ࢆⴠࡕⴥࡸᯤࢀᯞ࡞࡝ࡢୖ࡟ᩘ⛊㛫ᢲࡋ࠶࡚ࡿ࡜ࠊ༸࠿ࡽࡩ໬ࡋࡓᗂ⹸㸦ᮍ྾╔ࡢࢶࢶ࣒࢞ࢩ㸧ࡀ㯮ᕸ࡟௜╔ࡍࡿᛶ㉁ࢆ฼⏝ࡋࡓ᥇㞟ἲ࡛࠶ࡿ㸦ෆᕝ࡯࠿ࠊ㸧ࠋ௜╔ࡋࡓᗂ⹸ࡣ྾ᘬჾ࡛྾࠸ྲྀࡾࠊᐇ㦂ᐊ࡟ᣢࡕᖐࡗࡓᚋࠊ㸣࢚ࢳࣝ࢔ࣝࢥ࣮ࣝ࡟ಖᏑࡋࠊࣉࣞࣃ࣮ࣛࢺᶆᮏ࡟ࡋࡓࠋࡲࡓࡇࡢ୧ᆅᇦ࡛ᤕ⋓ࡉࢀࡓ࢔ࣛ࢖ࢢ࣐࡟ࡣࠊከᩘࡢ࣐ࢲࢽࡶᐤ⏕ࡋ࡚࠸ࡓࡓࡵࠊ᳜⏕ୖ࡟ࡶྠ✀ࡢ࣐ࢲࢽࡀ࠸ࡿࡢ࠿࡝࠺࠿ࢆ☜ㄆࡍࡿࡓࡵࠊࣇࣛࣥࢿࣝἲ㸦ᅗ 㸧࡛࣐ࢲࢽࡢ᥇㞟ࡶヨࡳࡓࠋࡇࢀࡣ ੉ ੉ࡢⓑⰍࡢࣇࣛࣥࢿࣝࢆ⏝࠸࡚ࠊᯘ⦕ࡸᯘෆࡢⴠࡕⴥࡢୖࢆࡺࡗࡃࡾ࡜࡞ࡒࡿࡇ࡜࡛ࠊࡑࡇ࡟࠸ࡿ࣐ࢲࢽ࡞࡝ࡀ௜╔ࡍࡿ᥇㞟ἲ࡛᪝᣺ࡾἲ࡜ࡶ࠸࠺㸦㧗⏣㸪㸧ࠋࠉㄪᰝࢆ⾜ࡗࡓẟ࿅ᒣ⏫ⴱ㈏࡜⠼࿴⏣ࡢ⮬↛⎔ቃࡣ௨ୗࡢࡼ࠺࡛࠶ࡗࡓࠋẟ࿅ᒣ⏫㈏ⴱ㸦ᅗ  㸧㸸ᖺ᭶᪥࡟ᤕ⋓ࡉࢀࡓ࢔ࣛ࢖ࢢ࣐㸦ಶయ␒ྕ㸧࡟ࠊಶయࡢࢱࢸࢶࢶ࣒࢞ࢩࡀᐤ⏕ࡋ࡚࠸ࡓሙᡤ࡛࠶ࡿࠋࢲ࢖ࢥࣥࡸⓑ⳯࡞࡝ࡢ㔝⳯⏿࡛ࠊ㢖⦾࡟࢔ࣛ࢖ࢢ࣐ࡢ౵ධࡀࡳࡽࢀࡓࠋ⏿ࡢ࿘ᅖࡣⴠⴥᗈⴥᶞࡢ࢚ࣀ࢟ࡀᩓᅾࡍࡿ࡜࡜ࡶ࡟ࠊ࢔ࢬ࣐ࢿࢨࢧࡀᐦ⏕ࡋ࡚࠾ࡾࠊ࢔ࣛ࢖ࢢ࣐ࡢ㏻ࡾ㐨࡜ᛮࢃࢀࡿࢺࣥࢿࣝࡀఱ࠿ᡤࡶ࠶ࡗࡓࠋࢺࣥࢿࣝෆࡢᆅ⾲㠃ࡣⴠࡕⴥࡸᯤࢀࡓ➲ࡢⴥ࡛ᩜࡁࡘࡵࡽࢀ࡚࠾ࡾࠊࡇࡢࡼ࠺࡞ሙᡤ࡛㯮ᕸ࡟ࡼࡿ᥇㞟ࢆ⾜ࡗࡓࠋẟ࿅ᒣ⏫⠼࿴⏣㸦ᅗ  㸧㸸ⴱ㈏࠿ࡽ໭࡟⣙  ੊㞳ࢀࡓ఩⨨࡟࠶ࡾࠊᖺ᭶᪥࡟Ẹᐙࡢᗞ࡛ᤕ⋓ࡉࢀࡓ࢔ࣛ࢖ࢢ࣐㸦ಶయ␒ྕ㸧࡟ࠊಶయࡶࡢከᩘࡢࢱࢸࢶࢶ࣒࢞ࢩࡀᐤ⏕ࡋ࡚࠸ࡓࠋࡇࡢẸᐙࡢ⿬ࡣୣ㝠ᆅ࡟࡞ࡗ࡚ᅗ  ࠉࣇࣛࣥࢿࣝἲ࡟ࡼࡿ࣐ࢲࢽ᥇㞟㸫㸫$㸪ᆅ⾲㠃ࡸ᳜⏕ୖࢆࡺࡗࡃࡾ࡞ࡒࡿ㸬%㸪ࣇࣛࣥࢿࣝୖࡢ࣐ࢲࢽ㧗ᶫࠉᏲ࣭୕ゅோᏊ࣭⸨⏣ᏹஅࠉᇸ⋢┴す㒊ᆅᇦ࡛ᤕ⋓ࡉࢀࡓ୰ᆺ့ங㢮ࡢእ㒊ᐤ⏕⹸ㄪᰝ㸦ᖺᗘ㸧࠾ࡾࠊࣄࣀ࢟ࡸࢫࢠࡀ᳜ᯘࡉࢀࠊࡑࡢ࿘ᅖ࡟ࡣࢥࢼࣛࡸࢩࣛ࢝ࢩ࡞࡝ࡢⴠⴥᗈⴥᶞࡀぢࡽࢀࡓࠋᯘᗋࡣẚ㍑ⓗ᫂ࡿࡃ࢔ࢬ࣐ࢿࢨࢧࡸࢡ
࣐ࢨࢧࡀⅬᅾࡋࠊⴠࡕⴥࡸᯤⴥࡀᩜࡁワࡵࡽࢀ࡚࠾ࡾࠊ࢔ࣛ࢖ࢢ࣐ࡢ㏻ࡾ㐨࡜ᛮࢃࢀࡿ㋃
ࡳ㊧ࡀぢࡽࢀࡓࠋࡇࡢࡼ࠺࡞ሙᡤࢆ୰ᚰ࡟㯮ᕸ࠾ࡼࡧࣇࣛࣥࢿࣝἲ࡟ࡼࡿㄪᰝࢆ⾜ࡗࡓࠋㄪᰝࡣ㸰ࣨᡤ࡜ࡶᖺ᭶  ᪥࡟ᐇ᪋ࡋࠊㄪᰝ᫬㛫ࡣࠊ࠸ࡎࢀࡶ㯮ᕸἲࡀ  ᫬㛫ࠊࣇࣛࣥࢿࣝἲࡀศ࡛࠶ࡗࡓࠋࠉᤕ⋓ࡉࢀࡓື≀ࡸ᳜⏕ୖ࠿ࡽ᥇㞟ࡉࢀࡓእ㒊ᐤ⏕⹸ࡢྲྀࡾᢅ࠸࡜ྠᐃࡣࠊ᫖ᖺᗘ࡜ྠᵝ࡟ࡋ࡚⾜ࡗࡓ㸦㧗ᶫ࡯࠿ࠊ㸧ࠋᅗ  ࠉẟ࿅ᒣ⏫ⴱ㈏ࡢㄪᰝᆅ
$㸪㔝⳯⏿࡜㸪ࡑࡢ࿘㎶ࡣ࢔ࢬ࣐ࢿࢨࢧ࡜࢚ࣀ࡛࢟ᅖࡲࢀ
࡚࠸ࡿ㸹%㸪࢔ࢬ࣐ࢿࢨࢧࡀᐦ㞟ࡋ࡚࠾ࡾ㸪࠸ࡃࡘࡶࡢ࢔
ࣛ࢖ࢢ࣐ࡢࢺࣥࢿࣝࡀ࠶ࡿ㸹&㸪ࢺࣥࢿࣝෆ࡟㯮ᕸࢆ⨨ࡃ㸹'㸪㯮ᕸୖࢆ㏺࠺ࢱࢸࢶࢶ࣒࢞ࢩ㸹(㸪ⓒ෇⋢ࡢୖ࡟⨨࠸ࡓࢶࢶ࣒࢞ࢩࡢ኱ࡁࡉ࡟ὀ┠㸹)㸪య㛗⣙ੈࡢࢱࢸࢶࢶ࣒࢞ࢩᮍ྾╔ᗂ⹸㸬6FDOHEDUȣP⤖ᯝ࡜⪃ᐹࠉእ㒊ᐤ⏕⹸ࡢ᳨ᰝࢆ⾜ࡗࡓ࢔ࣛ࢖ࢢ࣐㢌ࠊࣁࢡࣅࢩࣥ  㢌ࠊࢽ࣍ࣥ࢔ࢼࢢ࣐  㢌࠾ࡼࡧ࣍ࣥࢻࢸࣥ  㢌ࡢྜィ  ✀㢌ࡢ୰ᆺ့ங㢮࠿ࡽ  ✀ಶయࡢ࣐ࢲࢽࠊ  ✀ಶయࡢࢶࢶ࣒࢞ࢩࠊ࠾ࡼࡧ  ✀ࡢࣀ࣑ಶయࡀ᥇㞟ࡉࢀࡓࠋࡲࡓ᳜⏕ୖ࠿ࡽࡣࢱࢸࢶࢶ࣒࢞ࢩ  ✀ࡢࡳಶయ࡜  ✀ࡢ࣐ࢲࢽಶయࡀ᥇㞟ࡉࢀࡓࠋヲ⣽ࢆ௨ୗ࡟グࡋࡓࠋϨ㸬࢔ࣛ࢖ࢢ࣐ࡢእ㒊ᐤ⏕⹸㸦⾲  ࠊᅗ  㸧ࠉ᳨ᰝࡋࡓ࢔ࣛ࢖ࢢ࣐㢌ࡢ඲࡚࡟ఱࡽ࠿ࡢእ㒊ᐤ⏕⹸ࡢᐤ⏕ࡀぢࡽࢀࡓࠋ≉࡟ẟ࿅ᒣ⏫
ⴱ㈏࡜⠼࿴⏣࡛ࠊ⛅ᮇ࡟ᤕ⋓ࡉࢀࡓ࢔ࣛ࢖ࢢ࣐࡟ᩘ༓ಶయࡢࢱࢸࢶࢶ࣒࢞ࢩࡢᐤ⏕ࡀぢࡽࢀࡓࡇ࡜ࡣࠊ௒ᚋࡘࡘࡀࡴࡋ⑓ࡀⓎ⏕ࡋࡓ࡜ࡁࡢᅗ  ࠉẟ࿅ᒣ⏫⠼࿴⏣ࡢㄪᰝᆅ$㸪ࢫࢠ㸪ࣄࣀ࢟ࡀ᳜ᯘࡉࢀ࡚࠸ࡿୣ㝠ᆅ㸹%㸪᫂ࡿ࠸ᯘෆ㸹&㸪ᯘෆࡢⴠࡕⴥࡢୖ࡟㯮ᕸࢆᩘ⛊㛫⨨ࡃ㸹'㸪㯮ᕸୖ
ࢆ㏺࠸ࡲࢃࡿయ㛗ੈࡢࢶࢶ࣒࢞ࢩࡢᮍ྾╔ᗂ⹸⾲  ࠉ࢔ࣛ࢖ࢢ࣐ࡢእ㒊ᐤ⏕⹸ಶయ␒ྕID ID ID ID ID ✀ࠉྡ᥇㞟ᖺ᭶᪥ᛶู య㔜 (kg)ᤕࠉ⋓ࠉሙࠉᡤ࢔ࣛ࢖ࢢ࣐   ᇸ⋢┴ẚ௻㒆࡜ࡁࡀࢃ⏫᪥ᙳ࢔ࣛ࢖ࢢ࣐   ᇸ⋢┴ẚ௻㒆࡜ࡁࡀࢃ⏫᪥ᙳ࢔ࣛ࢖ࢢ࣐   ᇸ⋢┴኱㔛㒆ᐤᒃ⏫ᑠᅬ࢔ࣛ࢖ࢢ࣐   ᇸ⋢┴⛛∗㒆㛗™⏫ୗ㔝ୖ࢔ࣛ࢖ࢢ࣐   ᇸ⋢┴ධ㛫㒆ẟ࿅ᒣ⏫኱㢮࢔ࣛ࢖ࢢ࣐   ᇸ⋢┴ධ㛫㒆ẟ࿅ᒣ⏫኱Ꮠୗᕝཎ࢔ࣛ࢖ࢢ࣐   ᇸ⋢┴ධ㛫㒆ẟ࿅ᒣ⏫㜿ㄶゼ࢔ࣛ࢖ࢢ࣐   ᇸ⋢┴ධ㛫㒆ẟ࿅ᒣ⏫༡ྎ࢔ࣛ࢖ࢢ࣐   ᇸ⋢┴ධ㛫㒆ẟ࿅ᒣ⏫ᕝゅ࢔ࣛ࢖ࢢ࣐   ᇸ⋢┴ධ㛫㒆ẟ࿅ᒣ⏫ᕝゅ࢔ࣛ࢖ࢢ࣐  ᇸ⋢┴ධ㛫㒆ẟ࿅ᒣ⏫኱Ꮠすᡞ࢔ࣛ࢖ࢢ࣐  ᇸ⋢┴ධ㛫㒆ẟ࿅ᒣ⏫す኱ஂಖ࢔ࣛ࢖ࢢ࣐  ᇸ⋢┴ධ㛫㒆ẟ࿅ᒣ⏫㜿ㄶゼ࢔ࣛ࢖ࢢ࣐  ᇸ⋢┴ධ㛫㒆ẟ࿅ᒣ⏫ⴱ㈏࢔ࣛ࢖ࢢ࣐  ᇸ⋢┴ධ㛫㒆ẟ࿅ᒣ⏫ⴱ㈏࢔ࣛ࢖ࢢ࣐  ᇸ⋢┴ධ㛫㒆ẟ࿅ᒣ⏫ⴱ㈏࢔ࣛ࢖ࢢ࣐  ᇸ⋢┴ධ㛫㒆ẟ࿅ᒣ⏫ⴱ㈏࢔ࣛ࢖ࢢ࣐  ᇸ⋢┴ධ㛫㒆ẟ࿅ᒣ⏫ⴱ㈏࢔ࣛ࢖ࢢ࣐  ᇸ⋢┴ධ㛫㒆ẟ࿅ᒣ⏫⠼࿴⏣࢔ࣛ࢖ࢢ࣐  ᇸ⋢┴ධ㛫㒆ẟ࿅ᒣ⏫ⴱ㈏࢔ࣛ࢖ࢢ࣐  ᇸ⋢┴ẚ௻㒆࡜ࡁࡀࢃ⏫᪥ᙳྜࠉࠉࠉࠉࠉィ*M: male, F: female, N: nymph, L: larva
A. test: Amblyomma testudinariumࠉࢱ࢝ࢧࢦ࣐࢟ࣛࣛࢲࢽ
+ÀDYD+DHPDSK\VDOLVÀDYDࠉ࢟ࢳ࣐ࢲࢽ+ORQJ+DHPDSK\VDOLVORQJLFRUQLVࠉࣇࢱࢺࢤࢳ࣐ࢲࢽ
I. nippo: Ixodes nipponensisࠉࢱࢿ࢞ࢱ࣐ࢲࢽ
I. ova.: Ixodes ovatusࠉ࣐ࣖࢺ࣐ࢲࢽ
I. tan.: Ixodes tanukiࠉࢱࢾ࣐࢟ࢲࢽ

M
M
M
M
FFM
FM
FFM
MM
FM
FFFF࣐ࠉࠉࠉ ࢲࠉࠉࠉ ࢽA. test+ÀDYD

N

N
 N, L

M, NM, N





H. longM, F, N
M, F, N
M, FN, LLM, F, N
M, F, N
M, F, N
M, F, N
MFL
N, L M, F, N, L
N, LM, F, N, L
N, L
N, LM, F, N, LM, FNL
M, N, L
N
N
N

F, NM, F, N,
L


F, N

N, Lࢶࢶ࣒࢞ࢩ
I. ova.

I. tan.

L. scut.

L. pall.

ࣀࠉࠉ࣑C. mik

F

P.m.sinen.

C. fellisFM, FFM, F

M
F

N


F

M, FM, FM, FFF
M, F

L. scut.: Leptotrombidium scutellareࠉࢱࢸࢶࢶ࣒࢞ࢩ
L. pall.: Leptotrombidium pallidumࠉࣇࢺࢤࢶࢶ࣒࢞ࢩ&PLN&KDHWRSV\OODPLNDGR
P. m. sinen.: Paraceras melis sinensis&IHOOLV&WHQRFHSKDOLGHVIHOLVIHOLV㸫㸫I. nippo.

FFNFM, FM, Fᕝ༤⣖せࠉྕࠉ㸸ᅗ  ࠉ࢔ࣛ࢖ࢢ࣐ࡢయ⾲࡟ᐤ⏕ࡋ࡚࠸ࡿ࣐ࢲࢽᅗ  ࠉ࢟ࢳ࣐ࢲࢽ㸦ᗂ⹸㸧D㸪࢟ࢳ࣐ࢲࢽ㸦㞝㸧
㸹E㸪࠿࡞ࡾ྾⾑ࡋࡓ࢟ࢳ࣐ࢲࢽ㸦㞤㸧㸹
F㸪ࢱ࢝ࢧࢦ࣐࢟ࣛࣛࢲࢽ㸦ⱝ⹸㸧᭷ຊ፹௓✀࡜ࡋ࡚⪃࠼࡚࠾ࡃᚲせࡀ࠶ࢁ࠺ࠋ 㸧࣐ࢲࢽ㢮㸦ᅗ  㹼㸧ࠉ࢔ࣛ࢖ࢢ࣐㢌ࡢ඲࡚࡟࣐ࢲࢽࡢᐤ⏕ࡀࡳࡽࢀࡓࠋ᥇㞟ࡉࢀࡓ࣐ࢲࢽ㢮ࡢ✀㢮ࡣࠊࢱ࢝ࢧࢦ࣐࢟ࣛࣛࢲࢽ Amblyomma testudinariumࠊ࢟ࢳ࣐ࢲࢽ +DHPDSK\VDOLVÀDYDࠊࣇࢱࢺࢤࢳ
࣐ࢲࢽ +DHPDSK\VDOLVORQJLFRUQLVࠊࢱࢿ࢞ࢱ࣐ࢲ ࢽ Ixodes nipponensisࠊ ࣖ ࣐ ࢺ ࣐ ࢲ ࢽ Ixodesovatusࠊ ࢱ ࢾ ࢟ ࣐ ࢲ ࢽ Ixodes tanuki ࡢ  ✀ಶయ࡛࠶ࡗࡓࠋࡇࡢ࠺ࡕ࢟ࢳ࣐ࢲࢽࡀಶయ㸦㸣㸧࡛᭱ࡶከࡃࠊḟ࠸࡛ࢱ࢝ࢧࢦ࣐࢟ࣛࣛࢲࢽಶయ㸦㸣㸧ࠊࣇࢱࢺࢤࢳ࣐ࢲࢽ ಶయ㸦㸣㸧ࡢ㡰࡛ࠊࢱࢿ࢞ࢱ࣐ࢲࢽࠊ࣐ࣖࢺ࣐ࢲࢽࠊࢱࢾ࣐࢟ࢲࢽࡣ࠸ࡎࢀࡶ㸣࡛ᑡ࡞࠿ࡗࡓࠋࡇࢀࡽࡢᑡᩘಶయࡢ࣐ࢲࢽࡣࠊ௒ᅇㄪᰝࡋࡓ࢔ࣛ࢖ࢢ࣐ࡸࣁࢡࣅࢩࣥࡀࠊ࣐ࢲࢽࡢᮏ᮶ࡢᐟ୺࡛࡞࠸࠿ࠊ࠶ࡿ࠸ࡣ⏕ᜥᩘࡀᑡ࡞࠸ࡢ࠿ࡶࡋࢀ࡞࠸ࠋ࡞࠾ࠊࣇࢱࢺࢤࢳ࣐ࢲࢽ  ಶయࡣ⛛∗㒆㛗™⏫࡛ᤕ⋓ࡉࢀࡓ࢔ࣛ࢖ࢢ࣐࡟ᐤ⏕ࡋ࡚࠸ࡓಶయ࡛࠶ࡿࠋ

$㸪⫼㠃㸹%㸪㢡య㒊⫼㠃㸬ゅ≧య㸦▮༳㸧ࡣ୕ゅᙧ㸹&㸪➨ ⬮ᇶ⠇ࡢෆᲲ㸦D㸧ࡣᑠࡉࡃ㸪➨  ⬮࡛ࡣࡉࡽ࡟ᑠࡉ࠸㸦E㸧㸬ᅗ  ࠉࣇࢱࢺࢤࢳ࣐ࢲࢽ㸦ⱝ⹸㸧$㸪⫼㠃㸹%㸪⭡㠃㸬➨  ⬮ᇶ⠇ࡢෆᲲࡣ➨  㹼  ⬮ᇶ⠇ࡢෆᲲ㸦▮༳㸧ࡼࡾࡸࡸ㛗࠸㸹&㸪ཱྀୗ∦ࡢṑิࡣࡀ≉ᚩⓗ㸬6FDOHEDUȣPᅗ  ࠉࢱࢿ࢞ࢱ࣐ࢲࢽ㸦㞤㸧ᅗ  ࠉࢱ࢝ࢧࢦ࣐࢟ࣛࣛࢲࢽ㸦ᗂ⹸㸧$㸪⫼㠃㸹%㸪⭡㠃㸹&㸪⭡㠃ྑ➨  㹼  ⬮ᇶ⠇㸬➨  ⬮ᇶ⠇ࡢෆᲲ㸦D㸧ࡣ㛗࠸㸬➨  㹼  ⬮࡟࠶ࡿእᲲ㸦DEF㸧ࡣ▷࠸㸬6FDOHEDU  ੈᅗ  ࠉ࣐ࣖࢺ࣐ࢲࢽ㸦㞤㸧$㸪⫼㠃㸹%㸪ཱྀୗ∦㸹&㸪⭡㠃ྑഃࡢ➨  ⬮ᇶ⠇㸬ෆᲲ㸦D㸧ࡣᑠࡉࡃእᲲ㸦E㸧ࡣ኱ࡁࡃ᫂ࡽ࠿㸬6FDOH EDU ȣP$㸪⫼㠃㸹%㸪⭡㠃ྑഃࡢ➨  㹼  ⬮ᇶ⠇㸬➨  ⬮ᇶ⠇ࡢୗ᪉༙ศࢆⓑ࠸⭷≧ࡢ≀㉁࡛そࢃࢀࠊ⫧ཌࡋ࡚࠸ࡿࡢࡀ≉ᚩ㸦▮༳㸧㸬㸫㸫㧗ᶫࠉᏲ࣭୕ゅோᏊ࣭⸨⏣ᏹஅࠉᇸ⋢┴す㒊ᆅᇦ࡛ᤕ⋓ࡉࢀࡓ୰ᆺ့ங㢮ࡢእ㒊ᐤ⏕⹸ㄪᰝ㸦ᖺᗘ㸧࠾ࡼࡧ &WHQRFHSKDOLGHVIHOLVIHOLV㸦ᅗ㸧ࡢ  ✀C.f.felisಶయ࡛࠶ࡗࡓࠋࡇࡢ࠺ࡕࡀಶయ㸦㸣㸧࡛᭱ࡶከࡃࠊḟ࡟ P.m.sinensis ࡀಶయ㸦㸣㸧ࠊC.mikado㸦㸣㸧ࡢ㡰࡛࠶ࡗ
ࡓࠋC.f.felis ࡀ᭱ࡶከ࠿ࡗࡓࡢࡣࠊ࢔ࣛ࢖ࢢ࣐ࡀᤕ⋓ࡉࢀࡓሙᡤࡀேᐙ㏆ࡃࡢ⏿ࡸᒣᯘ࡛࠶ࡗࡓࡓࡵࠊ㣫࠸⊧ࡸ㔝⏕໬ࡋࡓ⊧࠿ࡽࡢⴠୗಶయࡀ࢔ࣛ࢖ࢢ࣐࡟⛣⾜ࡋࡓࡶࡢ࡜ᛮࢃࢀࡿࠋᮏ✀ࡣࠊࡇࢀࡲ࡛࡟ࢽ࣍ࣥ࢖ࢱࢳࠊࢩ࣋ࣜ࢔࢖ࢱࢳࠊ࢖ࢾࠊࢱࢾ࢟ࠊࣄࢺ࡞࡝࠿ࡽ
ࡶグ㘓ࡉࢀ࡚࠸ࡿ㸦Sakagutiࠊ㸹Sakaguti
and Jameson, 㸹㧗ᶫ࡯࠿ࠊ㸧ࠋᅗࠉࢱࢾ࣐࢟ࢲࢽ㸦ⱝ⹸㸧$㸪⫼㠃㸹%㸪㢡య㒊⭡㠃㸬ཱྀୗ∦ࡢඛ➃ࡣࡸࡸ୸࠸㸹&㸪⭡㠃ᕥഃࡢ➨  㹼  ⬮ᇶ⠇㸬➨  ⬮ᇶ⠇ࡢෆᲲ㸦D㸧ࡣእᲲ㸦E㸧ࡼࡾ㛗࠸ࠋ➨  ⬮ࠊ  ⬮ࡢእᲲ㸦F G㸧ࡣᑠࡉ࠸㸬
6FDOHEDUȣP 㸧ࢶࢶ࣒࢞ࢩ㢮㸦ᅗ㸧ࠉࢶࢶ࣒࢞ࢩࡣ  㢌࡟ᐤ⏕ࡋ࡚࠾ࡾࠊᐤ⏕⋡ࡣ㸣㸦/㸧࡛࠶ࡗࡓࠋࠉ᥇㞟ࡉࢀࡓࢶࢶ࣒࢞ࢩ㢮ࡣࠊࢱࢸࢶࢶ࣒࢞
ࢩ Leptotrombidium scutellare ࡜ࣇࢺࢤࢶࢶ࣒࢞ࢩ Leptotrombidium pallidum ࡢ  ✀ಶయ࡛ࠊࢱࢸࢶࢶ࣒࢞ࢩࡀಶయ࡛㸣ࢆ༨ࡵ࡚࠸ࡓࠋࣇࢺࢤࢶࢶ࣒࢞ࢩࡣ  㢌࠿ࡽࠊࢃࡎ࠿ಶయࡋ࠿᥇㞟࡛ࡁ࡞࠿ࡗࡓࠋࡇࢀࡣ࢔ࣛ࢖ࢢ࣐ࡢ⪥ࡢ࿘⦕࡟ᐤ⏕ࡋ࡚࠸ࡓಶయ࡛ࠊ⪥ࡢ஝⇱ࡀⴭࡋࡃࠊከᩘࡢ஝⇱ࡋࡓࢶࢶ࣒࢞ࢩࡀ☜ㄆ࡛ࡁࡓࡢ࡛ࠊ⢭ᰝࡍࢀࡤࠊࡉࡽ࡟᥇㞟࡛ࡁࡓࡶࡢ࡜ᛮࢃࢀࡿࠋᅗࠉẟ࿅ᒣ⏫ⴱ㈏࡛ᤕ⋓ࡉࢀࡓ࢔ࣛ࢖ࢢ࣐࡬ࡢࢶࢶ࣒࢞ࢩᐤ⏕౛$㸪ࢶࢶ࣒࢞ࢩࡀᐤ⏕ࡋ࡚࠸ࡓ࢔ࣛ࢖ࢢ࣐㸹%㸪⪥ࡢ࿘⦕㒊࡟ᐤ⏕ࡋ࡚࠸ࡓࣇࢺࢤࢶࢶ࣒࢞ࢩ㸦D㸧࡜ࢱࢸࢶࢶ࢞
࣒ࢩ㸦E㸧
㸹&㸪ࢱࢸࢶࢶ࣒࢞ࢩᗂ⹸ࡢ⫼㠃㸹'㸪⫼⏥ᯈ㸹(㸪ࣇࢺࢤࢶࢶ࣒࢞ࢩᗂ⹸ࡢ⫼㠃㸹)㸪⫼⏥ᯈ㸬6FDOHEDUȣP 㸧ࣀ࣑㢮ࠉ ࣀ ࣑ 㢮 ࡢ ᐤ ⏕ ⋡ ࡣ㸣㸦/㸧 ࡛ࠊ&KDHWRSV\OODPLNDGRࠊParaceras melis sinensisϩ㸬ࣁࢡࣅࢩࣥࠊࢽ࣍ࣥ࢔ࢼࢢ࣐࠾ࡼࡧ࣍ࣥࢻࢸࣥࡢእ㒊ᐤ⏕⹸㸦⾲  㸧ࠉࣁࢡࣅࢩࣥࡣᖺ  ᭶࡜  ᭶࡟ᤕ⋓ࡉࢀࡓ  㢌ࢆㄪ࡭ࡓ࡜ࡇࢁࠊ  㢌࠿ࡽࡣእ㒊ᐤ⏕⹸ࡀ᥇㞟ࡉࢀࡓࡀࠊẟ࿅ᒣ⏫኱㢮ࡢ  㢌㸦ಶయ␒ྕ㸧࡟ࡣᐤ⏕ࡋ࡚࠸࡞࠿ࡗࡓࠋࡲࡓࢽ࣍ࣥ࢔ࢼࢢ࣐࡜࣍ࣥࢻࢸࣥࡣࠊࡑࢀࡒࢀ 㢌ࡎࡘࢆㄪ࡭ࠊ࠸ࡎࢀࡶ࣐ࢲࢽ࡜ࣀ࣑ࡢᐤ⏕ࡀぢࡽࢀࡓࠋㄪ࡭ࡓ඲㢌࡟ࢶࢶ࣒࢞ࢩࡢᐤ⏕ࡣぢࡽࢀ࡞࠿ࡗࡓࠋࠉօࠉ࣐ࢲࢽ㢮ࠉࣁࢡࣅࢩࣥ࡟ᐤ⏕ࡋ࡚࠸ࡓ࣐ࢲࢽ㢮ࡣࠊࢱ࢝ࢧࢦ࣐࢟ࣛࣛࢲࢽࡢ  ಶయ㸦㞝  ࠊ㞤  ࠊⱝ⹸  ࠊᗂ⹸  㸧࡜࢟ࢳ࣐ࢲࢽ  ಶయ㸦㞤  ࠊⱝ⹸  ࠊᗂ⹸  㸧ࡢ  ✀ಶయ࡛࠶ࡗࡓࠋࡇ
ࡢ࠺ࡕ࢟ࢳ࣐ࢲࢽࡣ  㢌ࡢ࠺ࡕ  㢌࡟ᐤ⏕ࡋ࡚࠾ࡾࠊᐤ⏕⋡㸣࡛࠶ࡗࡓࠋࡋ࠿ࡋ࢔ࣛ࢖ࢢ࣐࡬ࡢᐤ⏕ᩘࡼࡾࡣ኱ኚᑡ࡞࠿ࡗࡓࠋࣁࢡࣅࢩࣥࡣẟ࡙ࡃࢁ࠸࡟ࡼࡗ࡚✚ᴟⓗ࡟ࢲࢽࢆྲྀࡾ㝖ࡃ⾜ືࡀࡳࡽࢀࡿࡀࠊࡑࡢࡓࡵ⏕ែⓗࢺࣛࢵࣉ࡜ࡋ࡚స⏝ࡍࡿྍ⬟ᛶࡀ࠶ࡾࠊᐤ⏕ᩘࡀᑡ࡞࠸࡜⪃࠼ࡽࢀࡿ㸦Doi at al., 㸧ࠋࠉᖺ᭶࡟ᤕ⋓ࡉࢀࡓ஧࣍ࣥ࢔ࢼࢢ࣐ࢆ⢭ᰝࡋࡓ࡜ࡇࢁࠊయ⾲ࡣ኱ኚࡁࢀ࠸࡛ࠊ⪥௓࡟࢟ࢳ࣐ࢲࢽ  ಶయ㸦㞝  ࠊ㞤  㸧ࡢᐤ⏕ࡀぢࡽࢀࡓ࡟ࡍࡂ࡞࠿ࡗࡓࠋ୍᪉ࠊ⛛∗㒆ᑠ㮵㔝⏫ࡢ࣍ࣥࢻࢸࣥ࡟ࡣ࢟ࢳ࣐ࢲࢽ  ಶయ㸦㞝  ࠊ㞤 ࠊⱝ⹸  㸧࡜ࣇࢱࢺࢤࢳ࣐ࢲࢽࡢⱝ⹸  ಶయࡀᐤ⏕ࡋ࡚࠸ࡓࠋᮏ✀ࡣẚ㍑ⓗ᫂ࡿ࠸⎔ቃࢆዲࡳࠊᕷ⾤ᆅࡢⲡᆅࡸ㎰⪔ᆅ࿘㎶ࡢⲡᆅ࠿ࡽࡋࡤࡋࡤ᥇㞟ࡉࢀࡿࡇ࡜࠿ࡽ㸦㧗⏣࡯࠿ࠊ㸧ࠊ࡞ࡐẟ࿅ᒣ⏫ࡢᕷ⾤ᆅ࡟ฟἐࡍࡿ࢔ࣛ࢖ࢢ࣐ࡸࣁࢡࣅࢩࣥ࠿ࡽ᥇㞟ࡉࢀ࡞࠿ࡗࡓࡢ࠿ࠊ௒ᚋࡢㄪᰝ࡛᫂ࡽ࠿࡟ࡋࡓ࠸ㄢ㢟࡛࠶ࡿࠋ㸫㸫ᕝ༤⣖せࠉྕࠉ㸸⾲  ࠉࣁࢡࣅࢩࣥࠊࢽ࣍ࣥ࢔ࢼࢢ࣐࠾ࡼࡧ࣍ࣥࢻࢸࣥࡢእ㒊ᐤ⏕⹸ಶయ␒ྕID ✀ࠉࠉྡ᥇㞟ᖺ᭶᪥ᤕࠉ⋓ࠉሙࠉᡤࣁࢡࣅࢩࣥ  ᇸ⋢┴ධ㛫㒆ẟ࿅ᒣ⏫ᒾ஭ᮾࣁࢡࣅࢩࣥ  ᇸ⋢┴ධ㛫㒆ẟ࿅ᒣ⏫኱㢮ࣁࢡࣅࢩࣥ  ᇸ⋢┴ධ㛫㒆ẟ࿅ᒣ⏫ᕝゅࣁࢡࣅࢩࣥ  ᇸ⋢┴ධ㛫㒆ẟ࿅ᒣ⏫⟪࿴⏣ࣁࢡࣅࢩࣥ  ᇸ⋢┴ධ㛫㒆ẟ࿅ᒣ⏫༡ྎࢽ࣍ࣥ࢔ࢼࢢ࣐  ᇸ⋢┴ẚ௻㒆ᑠᕝ⏫ୖྂᑎ࣍ࣥࢻࢸࣥ ᇸ⋢┴⛛∗㒆ᑠ㮵㔝⏫୕ᒣ*M: male, F: female, N: nymph, L: larva
A. test: Amblyomma testudinariumࠉࢱ࢝ࢧࢦ࣐࢟ࣛࣛࢲࢽ
+ÀDYD+DHPDSK\VDOLVÀDYDࠉ࢟ࢳ࣐ࢲࢽ
+ORQJ+DHPDSK\VDOLVORQJLFRUQLVࠉࣇࢱࢺࢤࢳ࣐ࢲࢽ

࣐ࢲࢽ+ÀDYD
F

ᛶูయ㔜
㸦਻㸧

M
FF
F
M
MMF, NN MFNL N, LM, FM, F, N

A. testH. long

C. mikFNࣀ࣑
P.m.sinenM, FM, F
F&PLN&KDHWRSV\OODPLNDGR
P. m. sinen: Paraceras melis sinensisࠉֆࠉࣀ࣑㢮ࠉࣁࢡࣅࢩࣥ  㢌࠿ࡽ C. mikado  ಶయ㸦㞤㸧࡜ P.m.sinensis  ಶ య㸦  㞝  㞤 㸧ࠊ஧࣍ࣥ࢔
ࢼࢢ࣐࠿ࡽ P.m.sinensis  ಶయ㸦㞝  ࠊ㞤  㸧ࡀᚓࡽࢀࠊ࣍ࣥࢻࢸࣥ࠿ࡽ P.m.sinensis ࡢ㞤 ಶయࡀᚓࡽࢀࡓࠋ࠸ࡎࢀࡶࡇࢀࡽࡢ୰ᆺ့ங
㢮࡟ᐤ⏕ࡍࡿᬑ㏻✀࡛࠶ࡿ㸦Sakagutiࠊ㸹
Sakaguti and Jameson, 㸹㧗ᶫ࡯࠿ࠊ㸧ࠋ䊢䠊 ᇸ⋢┴ධ㛫㒆ẟ࿅ᒣ⏫ⴱ㈏䛸⠼࿴⏣䛷䛾䝒䝒䜺䝮䝅䛸䝬䝎䝙ㄪᰝ 䠄⾲ 3 䠅ࠉ୧ᆅᇦ࡜ࡶࠊ࠾ࡶ࡟࢔ࣛ࢖ࢢ࣐ࡢ㏻ࡾ㐨࡜ᛮࢃࢀࡿሙᡤ࡛ㄪᰝࢆᐇ᪋ࡋࡓ࡜ࡇࢁࠊࢱࢸࢶ࣒࢞ࢩࡀⴱ㈏࡛ಶయࠊ⠼࿴⏣࡛ಶయࡀ᥇㞟ࡉࢀࡓࡀࠊᩘⓒಶయࡀᐦ㞟ࡍࡿᆅⅬ㸦ࢡࣛࢫࢱ࣮㸧ࢆぢࡘࡅࡿࡇ࡜ࡣ࡛ࡁ࡞࠿ࡗࡓࠋࡋ࠿ࡋ࿘ᅖࡀᒣ࡛ᅖࡲࢀ࡚࠸ࡿୣ㝠ᆅ࡛ࡶࠊ᪥ᙜࡓࡾࡀࡼ࠸༡ᩳ㠃ࡸࠊ㐺ᗘ࡟ⴠࡕⴥࡀሁ✚ࡋ࡚࠸ࡿሙᡤ࡞࡝ࠊᐟ୺ື≀ࡀዲࡴࡼ࠺࡞ሙᡤ࡟ᮏ✀ࡀศᕸࡋ࡚࠸ࡿࡇ࡜ࡀ☜ㄆ࡛ࡁࡓࠋࠉ࣐ࢲࢽ㢮ࡣẟ࿅ᒣ⏫ⴱ㈏࡛ࡣ࢟ࢳ࣐ࢲࢽࡢⱝ⹸  ಶయࠊࡲࡓẟ࿅ᒣ⏫⠼࿴⏣࡛ࡣࢱ࢝ࢧࢦ࣐࢟ࣛࣛࢲࢽࡢᗂ⹸  ಶయࠊ࢟ࢳ࣐ࢲࢽࡢⱝ⹸  ಶయ࡜࢔࢝ࢥࢵࢥ࣐ࢲࢽ Ixodes turdusࡢⱝ⹸  ಶయ㸦ᅗ㸧ࡀᚓࡽࢀࡓࠋ࢟ࢳ࣐ࢲࢽࡣㄪᰝࡋࡓ့ங㢮ࡍ࡭࡚࡟ᐤ⏕ࡋ࡚࠾ࡾࠊᆅ⾲㠃ࡢಶయࡣᐤ⏕ࡍࡿᶵ఍ࢆᚅࡗ࡚࠸ࡿಶయ࡛࠶ࢁ࠺ࠋࡲࡓࢱ࢝ࢧࢦ࣐࢟ࣛࣛࢲࢽࡢᮍ྾╔ᗂ⹸  ಶయࡀᚓࡽࢀࡓࡇ࡜ࡣࠊࡇࡢሙᡤ࡛⦾Ṫࡋ࡚࠸ࡿࡇ࡜ࢆ♧ࡍ᭷ຊ࡞ドᣐ࡜⪃࠼ࡽࢀࡓࠋ࢔࢝ࢥࢵࢥ࣐ࢲࢽࡣࠊࡑࡢ඲Ⓨ⫱ᮇࡀ㫽㢮࡟ぢࡽࢀࠊ㫽Ⴔዲᛶࡀᙉ࠸ࡶࡢࡢࠊ᳜⏕ୖ࠿ࡽ᥇㞟ࡉࢀࡿࡇ࡜ࡶ࠶ࡾࠊ⛥࡟ࡣ㔝㰡࠿ࡽࡢ᥇㞟౛ࡶ࠶ࡿ㸦Yamaguti et al., 㸹㧗⏣࡯࠿ࠊ㸧ϫ㸬እ㒊ᐤ⏕⹸ࡢศ㢮⩌ࡈ࡜ࡢᴫせࠉ࣐ࢲࢽ㢮ࡣ  ✀ࡀ᥇㞟ࡉࢀࡓࡀࠊẚ㍑ⓗᐤ⏕ᩘࡢከ࠿ࡗࡓࢱ࢝ࢧࢦ࣐࢟ࣛࣛࢲࢽ࡜࢟ࢳ
࣐ࢲࢽ࡟ࡘ࠸࡚ࠊ⏕ែࡢ୍㒊ࢆグࡋࡓࠋ௚ࡢ ✀ࡣ᥇㞟ᩘࡀᩘಶయ࡛࠶ࡗࡓࡓࡵࠊᮏ᮶ࡢᐟ୺࡜ࡣ␗࡞ࡿ࡜ᛮ࠼ࡓࡢ࡛┬␎ࡋࡓࠋࡲࡓࣀ࣑㢮࡟ࡘ࠸࡚ࡣࠊ᫖ᖺᗘ࡜࡯ࡰྠࡌ✀ࡀᐤ⏕ࡋ࡚࠸ࡓࡓࡵ㸦㧗ᶫ࡯࠿ࠊ㸧ࠊࡑࢀࡒࢀࡢእ㒊ᐤ⏕⹸ࡢ㡯┠࡟グࡋࡓࠋᅗࠉ࢔࢝ࢥࢵࢥ࣐ࢲࢽ㸦ⱝ⹸㸧$㸪⫼㠃㸹%㸪ཱྀୗ∦ࡢඛ➃ࡀᑤࡗ࡚࠸ࡿ㸹&㸪⬗㒊࡟⏕࠼ࡿከᩘࡢ㛗࠸ẟࡀ≉ᚩ㸦▮༳㸧㸹'㸪⭡㠃ᕥഃࡢ➨  㹼 ⬮ᇶ⠇㸬➨  ⬮ᇶ⠇ࡢෆᲲ࡜እᲲ㸦▮༳㸧ࡣ᫂ࡽ࠿㸬➨ 㹼  ⬮ᇶ⠇ࡢእᲲࡣ᫂ࡽ࠿㸦▮༳㸧ࡔࡀ㸪ෆᲲࡣ࡞࠸࠿ᑠࡉ࠸㸬ᅗࠉ&WHQRFHSKDOLGHVIHOLVIHOLV$㸪㞤ࡢ㢌㒊㸹%㸪㞤ࡢ⏕Ṫჾ㸹&㸪㞝ࡢ㢌㒊㸹'㸪㞝ࡢ⏕Ṫჾ㸫㸫㧗ᶫࠉᏲ࣭୕ゅோᏊ࣭⸨⏣ᏹஅࠉᇸ⋢┴す㒊ᆅᇦ࡛ᤕ⋓ࡉࢀࡓ୰ᆺ့ங㢮ࡢእ㒊ᐤ⏕⹸ㄪᰝ㸦ᖺᗘ㸧⾲  ࠉᇸ⋢┴ධ㛫㒆ẟ࿅ᒣ⏫ࡢ  ࠿ᡤ࡛ࡢࢶࢶ࣒࢞ࢩ࡜࣐ࢲࢽㄪᰝሙࠉࠉᡤㄪᰝ᪥㯮 ᕸ ἲL. scut.

ࣇࣛࣥࢿࣝἲẟ࿅ᒣ⏫ⴱ㈏

:㸫:㸸㸸ẟ࿅ᒣ⏫⠼࿴⏣

:㸫:㸸㸸

A. test.

+ÀDYDI. turd.

NL

N

N

N: nymph, L: larva,ÀDYD+DHPDSK\VDOLVÀDYDࠉ࢟ࢳ࣐ࢲࢽ
I. turd.: Ixodes turdusࠉ࢔࢝ࢥࢵࢥ࣐ࢲࢽ

L.scut.: Leptotrombidium scutellareࠉࢱࢸࢶࢶ࣒࢞ࢩ
A. test: Amblyomma testudinariumࠉࢱ࢝ࢧࢦ࣐࢟ࣛࣛࢲࢽ ࠉ࣐ࢲࢽ㢮ࠉօࠉࢱ࢝ࢧࢦ࣐࢟ࣛࣛࢲࢽࠉ  㢌ࡢ࢔ࣛ࢖ࢢ࣐㸦ᐤ⏕⋡㸣㸧࠿ࡽྜィಶయ㸦ⱝ⹸ ᗂ⹸㸧ᚓࡽࢀࡓࠋᏘ⠇ⓗ࡟ࡣ᫓㸦  ࠊ  ᭶㸧࡟ᤕ⋓ࡉࢀࡓ㢌ࡢ࢔ࣛ࢖ࢢ࣐ࡢ࠺ࡕࠊ  㢌࠿ࡽྜィಶయ㸦ⱝ⹸ ᗂ⹸㸧ࡀᚓࡽࢀࡓࡀࠊᡂ⹸ࡢᐤ⏕ࡣࡳࡽࢀ࡞࠿ࡗࡓࠋ⛅㸦ࠊ᭶㸧࡟ᤕ⋓ࡉࢀࡓ 㢌࡟ࡣ඲ࡃᐤ⏕ࡀࡳࡽࢀࡎࠊࢃࡎ࠿࡟᭶ ᪥࡟ᤕ⋓ࢃࢀࡓ  㢌࡟ࠊ  ಶయࡢⱝ⹸ࡀᐤ⏕ࡋ࡚࠸ࡓ࡟ࡍࡂ࡞࠿ࡗࡓ㸦⾲  㸧ࠋᮏ✀ࡣẟ࿅ᒣ⏫ࡢ㞄⏫࡛࠶ࡿẚ௻㒆࡜ࡁࡀࢃ⏫࡛ࠊᖺ  ᭶᪥࡟ᤕ⋓ࡉࢀࡓ࢔ࣛ࢖ࢢ࣐ 㢌ࡢ඲࡚࠿ࡽࠊྜィಶయࡢⱝ⹸ࡢࡳࡀᚓࡽࢀ࡚࠸ࡿ㸦㧗ᶫ࡯࠿ࠊ㸧ࠋࡲࡓᖺ ᭶᪥࡜᪥࡟ẟ࿅ᒣ⏫࡛ᤕ⋓ࡉࢀࡓ 㢌ࡢ஧࣍ࣥ࢖ࣀࢩࢩ㸦Sus scrofa㸧࠿ࡽࡣ඲
࡚ࡢⓎ⫱ᮇࡢᐤ⏕ࡀࡳࡽࢀࡓ㸦ᡂ⹸㞤ಶయࠊ ᡂ ⹸ 㞝ಶ యࠊ ⱝ ⹸ಶ యࠊ ᗂ ⹸  ಶయ㸧㸦Takahashi et al.,㸧ࠋ௨ୖࡢࡇ࡜࠿ࡽ᫓ᮇ࡟ࡣࠊ㉺෤ࡋࡓᗂ⹸ࠊⱝ⹸࠾ࡼࡧᡂ⹸ࡀᐤ⏕ࡋ࡚࠾ࡾࠊ  ᭶ୗ᪪࡟ࡣ㣬⾑ᗂ⹸ࡀ⬺⓶ࡋ࡚ⱝ⹸࡟࡞ࡗࡓಶయࡀᐤ⏕ࡋ࠺ࡿ࡜⪃࠼ࡽࢀࡓࠋࠉࡲࡓ  ᭶ୗ᪪࡟ᐤ⏕ࡋ࡚࠸ࡓⱝ⹸ࡣࠊ  ᭶࡟ࡣ⬺⓶ࡋ࡚ᡂ⹸࡟࡞ࡾࠊኟᮇ㸦    ᭶㸧࡟ࡣᡂ⹸ࡢᐤ⏕ࡀぢࡽࢀࡿ࠿ࡶࡋࢀ࡞࠸ࠋ㛗ᓮ┴ᑐ㤿ᕷࡢ࢖ࣀࢩࢩ࡟ࡣࠊኟᮇ࡟ከࡃࡢᡂ⹸ࡢᐤ⏕ࡀぢࡽࢀ࡚࠾ࡾࠊࡇࢀࡽࡢᡂ⹸ࡣ๓ᖺᗘ࠿ࡽࡢ㉺෤ᡂ⹸࡜ࠊ㉺෤ⱝ⹸࠿ࡽ⬺⓶ࡋࡓᡂ⹸ࡶᐤ⏕ࡋ࡚࠸ࡿࡶࡢ࡜⪃࠼ࡽࢀࡓࠋᑐ㤿࡛᥇㞟ࡋࡓ㣬⾑ᡂ⹸ࢆࠊᕝ㉺ᕷࡢ㔝እࡢ⮬↛᮲௳ୗ࡛ᐇ㦂ⓗ࡟㣫⫱ࡋ࡚ࡳࡿ࡜ࠊୖ㏙ࡋࡓࡼ࠺࡞⏕άྐࡢ୍㒊ࡀ☜ㄆࡉࢀ࡚࠸ࡿ㸦㧗ᶫ࡯࠿ࠊᮍⓎ⾲㸧ࠋࠉ࡞࠾ࠊᮏ✀ࡢẟ࿅ᒣ⏫ࡢ࢔ࣛ࢖ࢢ࣐࡬ࡢᐤ⏕⋡ࡀࠊ᫓ᮇ㸦    ᭶㸧࡟ࡣ㸣㸦/㸧࡜㧗࠸ࡇ࡜࠿ࡽࠊ㏆㞄ᆅᇦࡣࡶ࡜ࡼࡾࠊ࠿࡞ࡾᗈ⠊ᅖ࡟ศᕸࡀᣑ኱ࡋ࡚࠸ࡿࡶࡢ࡜᥎ᐃࡉࢀࡿࠋࠉᮏ✀ࡣᖺ  ᭶  ᪥࡟ẟ࿅ᒣ⏫࡛ᤕ⋓ࡉࢀࡓࢽ࣍ࣥ࢖ࣀࢩࢩ࡟ᡂ⹸ࡀᐤ⏕ࡋ࡚࠸ࡓࡇ࡜ࠊ⩣ᖺ  ᭶᪥࡜᪥࡟ᤕ⋓ࡉࢀࡓ஧࣍ࣥ࢖ࣀࢩࢩ࠿ࡽࡣ඲Ⓨ⫱ᮇࡀ᥇㞟ࡉࢀࡓࡇ࡜࡟ࡼࡾࠊᮏ✀ࡢᇸ⋢┴࡛ࡢศᕸࡢึㄆ౛࡟࡞ࡗࡓ㸦Takahashi et al㸬, 㸧ࠋࠉ་Ꮫⓗ࡟ࡣ⣚ᩬ⇕⩌ࣜࢣࢵࢳ࢔ࡢ୍✀࡛࠶ࡿ Rickettsia tamurae ࢆ≉␗ⓗ࡟㧗⋡࡟ಖ᭷ࡋࠊࣄࢺ࡬ࡢឤᰁ౛ࡀ࠶ࡿࡇ࡜㸦Ᏻ⸨࣭⸨ ⏣ࠊ㸹 ▼ ␇ ࡯ ࠿ࠊ㸹Imaoka etal㸬㸹 㧗ᶫࠊ㸧ࠋࡲࡓ࢘࢖ࣝࢫឤᰁ⑕ࡢ㔜⑕⇕ᛶ⾑ᑠᯈῶᑡ⑕ೃ⩌㸦SFTS㸧ࡢ፹௓✀ࡢ୍✀࡛࠶ࡿྍ⬟ᛶ㸦ᅜ❧ឤᰁ⑕◊✲ᡤ㸧ࠋࡉࡽ࡟⑓ཎయ࡜ࡢ㛵ಀࡣ୙᫂ࡔࡀࠊᮏ✀ࡢ่⑕࡟క࠺㐟㉮ᛶ⣚ᩬ Tick-associatedrash illness㸦TARI㸧㸦ኟ⛅࡯࠿ࠊ㸧࡞࡝ࡀ࠶ࡿࡓࡵࠊࡑࡢ་Ꮫⓗព⩏ࡣ㧗࠸ࠋࠉֆࠉ࢟ࢳ࣐ࢲࢽࠉㄪᰝࡋࡓ୰ᆺ့ங㢮  ✀㢌ࡢ࠺ࡕ㢌㸦㸣㸧࡟ᮏ✀ࡢᐤ⏕ࡀぢࡽࢀࠊࡋ࠿ࡶ඲Ⓨ⫱ᮇࡀ᥇㞟ࡉࢀࡓࠋ௒ᅇㄪᰝࡋࡓ  ✀ࡢ့ங㢮ࡢ඲࡚ࡀୣ㝠ᆅ࠿ࡽᒣᆅᖏୗ㒊࡟࠿ࡅ࡚⏕ᜥࡋ࡚࠾ࡾࠊࡇࢀࡽࡢᆅᇦ࡟ᗈࡃศᕸࡋ࡚࠸ࡿࡶࡢ࡜ᛮࢃࢀࡿࠋ௚ࡢ୰ᆺ࠿ࡽࢩ࢝ࡸ࢖ࣀࢩࢩ࡞࡝ࡢ኱ᆺ့ங㢮ࡢከࡃࡶࠊୣ㝠ᆅ࠿ࡽᒣᆅᖏ࡟࠿ࡅ࡚⏕ᜥࡋ࡚࠾ࡾࠊࡲࡓᮏ✀ࡀᐟ୺ࢆ࠶ࡲࡾ㑅ࡤ࡞࠸࡛ᐤ⏕ࡍࡿࡇ࡜࠿ࡽࠊࡇࢀࡽࡢᆅᇦ࡛ࡢ᭱ඃ༨✀࡟࡞ࡗ࡚࠸ࡿ࡜ᛮࢃࢀࡿ㸦⸨ᮏ࡯࠿ࠊ㸹⸨⏣࡯࠿ࠊ㸧ࠋࡓࡔᑠ့ங㢮ࡢࢧࢻ࢔࢝ࢿࢬ࣑ࡸ࢚ࢰࣖࢳࢿࢬ࣑࠿ࡽࡢグ㘓ࠊࡲࡓ㫽㢮࠿ࡽࡢグ㘓ࡶ࠶ࡿࡀ㸦Saito㸹 ኱㔝ࠊ㸹Yamaguti et al., 㸧ࠊࡇࢀࡽࡣ኱ኚ⛥࡞ᐤ⏕౛࡛࠶ࡿࡇ࡜࠿ࡽࠊᮏ✀ࡢᮏ᮶ࡢᐟ୺࡛ࡣ࡞࠸࡜ᛮࢃࢀࡿࠋࠉ ࢔ ࣛ ࢖ ࢢ ࣐㢌 ࡢ ࠺ ࡕ㢌㸦㸣㸧 ࡟ᮏ✀ࡢᐤ⏕ࡀぢࡽࢀࡓࡢ࡛ࠊ᫓ᮇ㸦᭶㸧࡜⛅ᮇ㸦᭶㸧࡛ᐤ⏕≧ἣࢆẚ㍑ࡋ࡚ࡳ

㸫㸫ᕝ༤⣖せࠉྕࠉ㸸ࡓࠋ᫓ᮇ࡟ࡣྜィಶయᚓࡽࢀࠊࡑࡢⓎ⫱ᮇࡈ࡜ࡢ᥇㞟ᩘࡣࠊ㞝㸦  㢌࠶ࡓࡾࡢᐤ⏕ᩘ㸧ࠊ㞤㸦㸧ࠊⱝ⹸㸦㸧ࠊᗂ⹸㸦㸧ࠊ⛅ᮇ࡟ࡣྜィಶయᚓࡽࢀࠊ㞝㸦㸧ࠊ㞤㸦㸧ࠊⱝ⹸㸦㸧ࠊᗂ⹸㸦㸧࡛࠶ࡗࡓࠋ᫓ᮇ࡜⛅ᮇ࡛ࡣࠊᡂ⹸࡜ᗂ⹸ࡣ⛅ᮇࡢ᪉ࡀࡸࡸከࡃࠊⱝ⹸ࡢᐤ⏕ᩘ࡟ࡣᕪࡀ࡞࠸ࡼ࠺࡟ᛮ࠼ࡓࠋ⸨ᮏ࡯࠿㸦㸧ࡣࠊᇸ⋢┴༡す㒊ࡢపᒣᖏ࠿ࡽள㧗ᒣᖏ࡟࠿ࡅ࡚ࠊࣇࣛࣥࢿࣝἲ࡟ࡼࡿ᳜⏕ୖ࡛ࡢᮏ✀ࡢᏘ⠇ⓗᾘ㛗ࢆㄪ࡭ࠊᡂ⹸ࡣ᫓ᮇ࡟᳜⏕ୖ࡬ࡢฟ⌧ᩘࡀ᭱ࡶከࡃࠊኟᮇ࡟ࡣ᭱ప࡟࡞ࡾࠊ⛅ᮇ࡟෌ࡧฟ⌧ᩘࡀከࡃ࡞ࡗࡓࠋⱝ⹸ࡶࠊᡂ⹸࡯࡝࡛ࡣ࡞࠸ࡀࠊ࡯ࡰྠࡌഴྥࢆ♧ࡋࡓࡶࡢࡢࠊᖺ㛫ࢆ㏻ࡌ࡚ከᩘಶయࡀぢࡽࢀ࡚࠸ࡿࠋ୍᪉ࠊᗂ⹸ࡣኟ࠿ࡽ⛅࡟࠿ࡅ࡚᫂░࡞ࣆ࣮ࢡࢆᣢࡘ  ᓠᛶࡢᏘ⠇ᾘ㛗ࢆ♧ࡋࡓࠋ㔝⏕ື≀ࡢయ⾲ୖࡢᐤ⏕ᩘ࡜᳜⏕ୖ࡛ࡢ᥇㞟ᩘࡣࠊᚲࡎࡋࡶ୍⮴ࡍࡿࢃࡅ࡛ࡣ࡞࠸ࡀࠊ࢔ࣛ࢖ࢢ࣐࡛ࡢ᫓ᮇ࡜⛅ᮇ࡛ࡢᐤ⏕ᩘࡣࠊ⸨ᮏ࡯࠿㸦㸧࡟ࡼࡿ᳜⏕ୖ࡛ࡢฟ⌧ᩘ࡜࡯ࡰྠࡌഴྥࢆ♧ࡋ࡚࠸ࡓࠋ௒ᚋࡣᖺ㛫ࢆ㏻ࡋ࡚ࠊື≀ࡢయ⾲ୖ࡜᳜⏕ୖࡢฟ⌧ᩘࡀ୍⮴ࡍࡿࡢ࠿࡝࠺࠿ࠊ࠶ࡽࡓࡵ࡚ㄪ࡭࡚ࡳࡿᚲせࡀ࠶ࢁ࠺ࠋ ࠉࢶࢶ࣒࢞ࢩ㢮ࠉẟ࿅ᒣ⏫ⴱ㈏࡛ᤕ⋓ࡉࢀࡓ  㢌ࡢ࢔ࣛ࢖ࢢ࣐࠿ࡽࠊࢱࢸࢶࢶ࣒࢞ࢩࡀಶయ࡛᭱ࡶከࡃ᥇㞟ࡉࢀࡓࠋࡲࡓẟ࿅ᒣ⏫⠼࿴⏣࡛ᤕ⋓ࡉࢀࡓ  㢌࠿ࡽࡶಶయࡀ᥇㞟ࡉࢀࡓࠋࡲࡓẟ࿅ᒣ⏫ⴱ㈏࡛ᤕ⋓ࡉࢀࡓ  㢌࡟ࡣࠊࡑࢀࡒࢀಶయࠊಶయࠊಶయ࡜ᑡ࡞࠿ࡗࡓࡇ࡜ࡣࠊᮏ✀ࡢศᕸ࡟∦ᐤࡀ࠶ࡿࡓࡵ࡛࠶ࢁ࠺ࠋࡋ࠿ࡋࡑࡢཎᅉ࡟ࡘ࠸࡚ࡣࠊࢶࢶ࣒࢞ࢩࡢ㣵ࢆྵࡵࡓᚤ⎔ቃࡀᙳ㡪ࡋ࡚࠸ࡿ࡜ᛮࢃࢀࡿࡀࠊලయⓗ࡞ཎᅉ࡟ࡘ࠸࡚ࡣ௒࡞࠾୙࡛᫂࠶ࡿ㸦㧗ᶫ࡯࠿ࠊ㸧ࠋࠉᮏ✀ࡣᇸ⋢┴ෆ࡛ࡣࠊධ㛫ᕝࡸⲨᕝࡢἙᕝᩜ࡟ᬑ㏻࡟ศᕸࡋ࡚࠸ࡿࡀ㸦㧗ᶫ࣭㛵᰿ࠊ㸧పᒣᖏ ࡛ ࡢ ᥇ 㞟 グ 㘓 ࡣᖺ᭶᪥࡟ࠊẚ௻㒆࡜ࡁࡀࢃ⏫ࡢᶆ㧗⣙㹫ࡢ┴㐨ୖ࡛஦ᨾ࡟࠶ࡗࡓ࣍ࣥࢻࢱࢾ࢟࠿ࡽ᥇㞟ࡉࢀ࡚࠸ࡿ㸦㧗ᶫࠊ㸧ࠋ࡞࠾ࠊᮏ✀ࡢศᕸ໭㝈ࡣࠊࡇࢀࡲ࡛ࡢ࡜ࡇࢁᒣᙧ┴࡛࠶ࡿࡀࠊࡑࡢศᕸ࡟ࡣ⃰ῐࡀࡳࡽࢀࡿࠋ≉࡟ከ࠸ࡢࡣ㮵ඣᓥ┴࡛ࠊᮡᯘࡢᯘᗋ㒊࡟ಶయࢆ㉸࠼ࡿࢡࣛࢫࢱ࣮࡟ࠊࡋࡤࡋࡤ㐼㐝ࡋࡓࡇ࡜ࡀ࠶ࡿࠋࣄࢺ่ညᛶࡀ࠶ࡾࠊࢶࢶ࣒࢞ࢩ⑓ࡢ፹௓✀࡛ࡶ࠶ࡿ㸦బࠎࠊ㸹㫽ᒣ࡯࠿㸪㸹Misumi et al㸬, 㸹 Takahashi etal㸬㸧ࠋࠉᖺ᭶᪥࡟ẟ࿅ᒣ⏫ⴱ㈏࡛ᤕ⋓ࡉࢀࡓ࢔ࣛ࢖ࢢ࣐࠿ࡽࣇࢺࢤࢶࢶ࣒࢞ࢩಶయࡀ᥇㞟ࡉࢀࡓࠋ⪥ࡢ࿘⦕㒊࡟ᐤ⏕ࡋ࡚࠸ࡓࡶࡢ࡛ࠊ࡯࡜ࢇ࡝ࡢಶయࡀ஝⇱ࡋ࡚ᖸ࠿ࡽࡧ࡚࠸ࡓࠋࡇࢀ࡛ࡣ࠸ࡃࡽᠱᆶἲ࡛᥇㞟ࢆヨࡳ࡚ࡶᅇ཰࡛ࡁࡎࠊࡲࡓయ⾲࡟྾╔ࡋ࡚࠸ࡿ⏕ࡁࡓಶయ࡛ࡶయ㛗⣙ੈ࡜ᑠࡉ࠸ࡓࡵࠊయ⾲ࢆ⢭ᰝࡋ࡚᥈ࡋฟࡍ࡟ࡣ㝈⏺ࡀ࠶ࡿࠋᮏ✀ࡣࢱࢸࢶࢶ࣒࢞ࢩ࡜ࡣࠊⱝᖸࡢⓎ⏕ࣆ࣮ࢡ࡟ᕪࡀ࠶ࡿࡀࠊ኱ࡁ࠸ἙᕝࡢἙᕝᩜ࡛ࠊࢱࢸࢶࢶ࣒࢞ࢩ࡜ྠ᫬࡟᥇㞟ࡉࢀࡿࡇ࡜ࡀከ࠸ࠋἙᕝᩜ࡟㝈ࡽࡎࠊᒣ㛫ᆅࡢỈ⏣ࡢ␏㐨ࡸࠊୣ㝠ᆅࡢᩳ㠃࡞࡝࡟Ⓨ⏕ࡍࡿࡇ࡜ࡶ࠶ࡿࠋࢱࢸࢶࢶ࣒࢞ࢩ࡜ྠᵝࠊ⛅ᮇ࡟࠾ࡅࡿࢶࢶ࣒࢞ࢩ⑓ࡢ୺せ፹௓✀࡛࠶ࡿ㸦㧗ᶫ࡯࠿ࠊ㸹Takahashiet al.㸧ㅰࠉࠉ㎡ࠉ࢔ࣛ࢖ࢢ࣐ࢆ୺࡜ࡍࡿ୰ᆺ့ங㢮ࡢእ㒊ᐤ⏕⹸ࡢㄪᰝ࡟㝿ࡋࠊከࡃࡢ᝟ሗࢆᥦ౪ࡋ࡚ୗࡉࡾࠊࡲࡓᤕ⋓࡟ࡈ༠ຊࡋ࡚ୗࡉࡗࡓୗグࡢಶே࠾ࡼࡧᶵ㛵ࡢⓙᵝ࡟ࠊㅽࢇ࡛ឤㅰࡢពࢆ⾲ࡋࡲࡍࠋᕷཎᖖḟࠊ㧗ᶫࠉ❶ࠊᖹᒣⰋ἞ࠊᏳ㇂῟ಙࠊᇸ⋢┴㎰ᴗᢏ⾡◊✲ࢭࣥࢱ࣮㸦ᩗ⛠␎㸧ᘬ⏝ᩥ⊩Ᏻ⸨⚽஧㸪⸨⏣༤ᕫ㸦㸧ᅜෆ࡟࠾ࡅࡿ⣚ᩬ⇕⩌ࣜࢣࢵࢳ࢔⑕ࢆ፹௓ࡍࡿ࣐ࢲࢽ㢮࡜⑓ཎయ࡜ࡢከᵝ࡞㛵ಀ㸬⾨⏕ື≀㸪㸸㸬
Doi K., Kono M., Kato T., Hayama S.㸦㸧
Ecological traps and boosters of ixodid ticks:
the differing ecological roles of two sympatricintroduced mammals. Ticks Tick Borne Diseases doi: /j.ttbdis
⸨⏣༤ᕫ㸪㧗ᶫࠉᏲ㸪ᒣᮏ㈆ྖ㸪ᩪ⸨ࠉ㈗㸪⏫
⏣࿴ᙪ㸦㸧ᇸ⋢࠾ࡼࡧ⩌㤿┴ୗ࡟࠾ࡅࡿ့ங㢮࡜㫽㢮࡟ᐤ⏕ࡍࡿ࣐ࢲࢽ㢮㸬   ᐟ୺㛵ಀ㸪ᆅ⌮ⓗ࡞ࡽࡧ࡟ᆶ┤ศᕸ㸪࠾ࡼࡧࡑࡢ་Ꮫⓗព⩏㸬኱ཎᖺሗ㸪㸸㸬⸨ᮏ࿴⩏㸪ᒣཱྀࠉ᪼㸪㧗ᶫࠉᏲ㸦㸧࣐ࢲࢽ㢮ࡢ⏕ែᏛⓗ◊✲  㸬ᇸ⋢┴༡す㒊పᒣᖏ࡟࠾ࡅࡿ᳜⏕ୖ࡜㔝⏕ື≀ୖࡢ࣐ࢲࢽ㢮㸬⾨⏕ື≀㸪㸸㸬⸨ᮏ࿴⩏㸪ᒣཱྀࠉ᪼㸪㧗ᶫࠉᏲ㸦㸧࣐ࢲࢽ㢮ࡢ⏕ែᏛⓗ◊✲  㸬ᇸ⋢┴༡す㒊࡟࠾ࡅ㸫㸫㧗ᶫࠉᏲ࣭୕ゅோᏊ࣭⸨⏣ᏹஅࠉᇸ⋢┴す㒊ᆅᇦ࡛ᤕ⋓ࡉࢀࡓ୰ᆺ့ங㢮ࡢእ㒊ᐤ⏕⹸ㄪᰝ㸦ᖺᗘ㸧ࡿ  ✀ࡢ࣐ࢲࢽ㢮㸪࢟ࢳ࣐ࢲࢽ㸪࣐ࣖࢺ࣐ࢲࢽ㸪ࢱࢿ࢞ࢱ࣐ࢲࢽࡢᏘ⠇ⓗᾘ㛗ࡢẚ㍑㸬
⾨⏕ື≀㸪㸸㸬
Imaoka K, Kaneko S, Tabara K, Kusatake K, MoritaE. () The first human case of Rickettsia
tamurae infection in Japan. Case Reports inDermatology. (): ▼␇ࠉྐ㸪⸨⏣༤ᕫ㸪ᖹ㔝ᫎᏊ㸪▮㔝Ὀᘯ㸪㧗⏣ఙᘯ㸦㸧⚟஭┴࡟࠾ࡅࡿ᪥ᮏ⣚ᩬ⇕ࡢឤᰁ⎔ㄪᰝ㸬⚟஭┴⾨⎔◊ᖺሗ㸪㸸㸬㫽ᒣࠉྐ㸪㕥ᮌࠉ༤㸪⏣୰㇏㐨㸬㸬ࢱࢸࢶࢶ࣒࢞ࢩࡢ่ည࡟ࡼࡿ⓶⭵⑕≧ؐ⮫ᗋ౛࡜ேయ྾╔ᐇ㦂⤒㐣࡟ࡘ࠸࡚㸬⓶⭵⮫ᗋ㸪㸸㸬ᅜ❧ឤᰁ⑕◊✲ᡤ㸦㸧㔜⑕⇕ᛶ⾑ᑠᯈῶᑡ⑕ೃ⩌㸦SFTS㸧㸪ᖺ  ᭶⌧ᅾ㸬⑓ཎᚤ⏕≀᳨ฟ᝟ሗ㸪㸸㸬
Misumi, H., Takahashi, M., Kadosaka, T., Urakami,H., Takahama, M., Lerdthusnee, K., Misumi, M.
and Matsumoto, I. () Comparative study ofhuman dermatitis caused by the bites of unfedlarval trombiculid mites, Leptotrombidium
pallidum and L. scutellare (Acari: Trombiculidae).
Med. Entomol. Zool., : ኟ⛅ඃ㸪㧗⏣ఙᘯ㸪ᕝ➃ᐶᶞ㸪బ⸨ࠉᲈ㸪㧗㔝ࠉឡ㸦㸧ࢱ࢝ࢧࢦ࣐࢟ࣛࣛࢲࢽ่⑕
࡟క࠺㐟㉮ᛶ⣚ᩬ㸸Tick-associated rash illness(TARI). Med. Entomol. Zool. Vol Nop኱㔝ၿྑ㛛㸦㸧㸸໭ᾏ㐨࡟࠾ࡅࡿ㔝㰡ᐤ⏕࣐ࢲࢽ㢮࡟ࡘ࠸࡚㸬໭ᾏ㐨⾨◊ᡤሗ㸪㸸㸬
Saito, Y. (): Studies on ixodid ticks , part I. On
ecology, with reference to distribution and seasonaloccurrence of ixodid ticks in Niigata Prefecture,Japan. Acta Med. Biol., : 
Sakaguti, K. () A monograph of the Siphonapteraof Japan plate  The Nippon Printingand Publishing Co., Ltd., Osaka, Japan.
Sakaguti, K. and Jameson, E. W. (). The
6LSKRQDSWHUDRI-DSDQ3DFL¿F,QVHFWV0RQRJUDSK pp., Entomology Department, Bernice P. BishopMuseum, Honolulu, Hawaii, U. S. A.
బࠎࠉᏛ㸦㸧᜔⹸࡜᜔⹸⑓㸪pp㸪་Ꮫ᭩㝔㸪ᮾி㸬㧗⏣ఙᘯ㸦㸧࣐ࢲࢽࡢ᥇㞟࡜ᶆᮏసᡂἲ㸬㧗 ⏣ ఙ ᘯ㸦 ⦅ 㸧㸪 ་ ࢲ ࢽ Ꮫ ᅗ 㚷㸬pp
㸪໭㝯㤋㸪ᮾி㸬㧗 ⏣ ఙ ᘯ㸪 ⸨ ⏣ ༤ ᕫ㸪 㧗 ᶫࠉ Ᏺ㸪 ኟ ⛅ࠉ ඃ㸦㸧࣐ࢲࢽᡂ⹸ࡢᒓ✀࡬ࡢ᳨⣴࡜ゎㄝ㸬㧗 ⏣ ఙ ᘯ㸦 ⦅ 㸧㸪 ་ ࢲ ࢽ Ꮫ ᅗ 㚷㸬pp
㸪໭㝯㤋㸪ᮾி㸬㧗ᶫࠉᏲ㸦㸧ᇸ⋢┴す㒊࡟࠾࠸࡚ࡣࡌࡵ࡚

ぢ࠸ฟࡉࢀࡓࢱࢸࢶࢶ࣒࢞ࢩ Leptotrombidium
scutellare Nagayo, Miyagawa, Mitamura, Tamiya etTenjin, ࡟ࡘ࠸࡚㸬ᇸ⋢⏕≀㸪㸸㸬㧘 ᶫࠉ Ᏺ㸪 ᒣ ᮏ ㈆ ྖ㸪 ᩧ ⸨ࠉ ㈗㸪 ⏫ ⏣ ࿴ ᙪ㸦㸧ᇸ⋢࠾ࡼࡧ⩌㤿┴ୗ࡟࠾ࡅࡿ့ங㢮࠾ࡼࡧ㫽㢮࡟ᐤ⏕ࡍࡿࣀ࣑㢮ࠋࠉ୰࣭኱ᆺ့ங㢮࡟ᐤ⏕ࡍࡿࣀ࣑㢮㸬኱ཎᖺሗࠊ㸸㸬㧗ᶫࠉᏲ㸪㛵᰿୓㔛Ꮚ㸦㸧ᇸ⋢┴ᕝ㉺ᕷࡢධ㛫ᕝἙ␁࡟࠾ࡅࡿ㔝㰡ࡢእ㒊ᐤ⏕⹸㢮Ϫ 㸪ࢶࢶ࣒࢞ࢩᗂ⹸ࡢᏘ⠇ᾘ㛗㸬ື≀࡜⮬↛㸪㸸㸬㧗ᶫࠉᏲ㸪ᮧ⏣㐨㔛㸪⏫⏣࿴ᙪ㸪ᇼࠉᰤኴ㑻㸦㸧⛛∗ᕷ⨺ᒣᆅ༊ࡢࢶࢶ࣒࢞ࢩᗂ⹸ࡢㄪᰝ࡜ࣜࢣࢵࢳ࢔ࢶࢶ࣒࢞ࢩࡢศ㞳  㸬ᇸ⋢་኱ㄅ㸪㸸㸬Takahashi, M., Urakami, H. and Misumi, H. ()Trombidiosis-dermatosis induced by the biteof larval trombiculid mite, Leptotrombidium
scutellare (Nagayo, Miyagawa, Mitamura, Tamiyaet Tenjin, ) (Prostigmata: Trombiculidae).
Ann. Rep. Ohara Hosp., : Takahashi, M., Misumi, H., Urakami, H., Nakajima,S., Furui, S., Yamamoto, S., Furuya, Y., Misumi,
M. and Matsumoto, I. () Mite vectors (Acari:Trombiculidae) of Scrub typhus in a new endemic
area in Northern Kyoto, Japan. J. Med. Entomol.,: 㧗ᶫࠉᏲ㸪୕ゅோᏊ㸪㧗⏣ఙᘯ㸦㸧ࢱࢸࢶࢶ࣒࢞ࢩࡢ⏕ᜥᚤ⎔ቃ࡜ྎ㢼࡟ࡼࡿὥỈୗ࡛ࡢ⏕Ꮡᡓ␎㸬Med. Entomol. Zool (Suppl.)㸸㸬㧗ᶫࠉᏲ㸦㸧㛵ᮾࡢࢶࢶ࣒࢞ࢩ⑓ࡢ⌧ἣ㸪ຍ࠼࡚࣐ࢲࢽ㢮ࡢࢺࣆࢵࢡࢫ㸬SADI ࢽ࣮ࣗࢫ㸦ᖺ ᝟ ሗ ஺ ᥮㸧㸸㸦http://www.
sadi-web-site.com/SADInews.pdf㸧
Takahashi, M., Misumi, H., Baba, Y., Fujita, N.
and Fujita, H. () First record of the tick
Amblyomma testudinarium Koch,  (Acari:Ixodidae) in Saitama Prefecture, Japan. Bull.
Saitama Mus. Nat. Hist. [N. S], : 㧘ᶫࠉᏲ㸪୕ゅோᏊ㸪㤿ሙ⿱⨾㸪ዟᮧࡳ࡯Ꮚ㸪⸨⏣ᏹஅ㸦㸧ᇸ⋢┴す㒊ᆅᇦ࡛ᤕ⋓ࡉࢀࡓ࢔ࣛ࢖ࢢ࣐࡜࢔ࢼࢢ࣐㸪࠾ࡼࡧᇸ⋢┴❧ᕝࡢ༤≀㤋࡜ᇸ⋢┴❧⮬↛ࡢ༤≀㤋ࡢ࣊ࣅ㢮ࡢእ㒊ᐤ⏕⹸ㄪᰝ㸬ᕝ༤⣖せ㸪㸸㸬ෆ ᕝ බ ே㸪 ᕝ ᳃ ᩥ ᙪ㸪 ᕝ ྜ Ύ ஓ㸪 ⇃ ⏣ ಙ ኵ㸦㸧ࢱࢸࢶࢶ࣒࢞ࢩᗂ⹸ࡢ㔝እࢧࣥࣉࣜࣥࢢ࡟ຠ⋡ⓗ࡞㕥ᮌࡢぢྲྀࡾἲ㸬᪥ᮏࢲࢽᏛ఍ㄅ㸪㸸㸬Yamaguti N, Tipton VJ, Keegan HL, Toshioka S.
() Ticks of Japan, Korea, and the Ryukyu
Islands. Brigham Young University ScienceBulletin Biological Series. : i-iv, 㸫㸫