ᕝ༤⣖せࠉྕࠉ㸸ᇸ⋢┴す㒊ᆅᇦ࡛ᤕ⋓ࡉࢀࡓ࢔ࣛ࢖ࢢ࣐࡜࢔ࢼࢢ࣐ࠊ࠾ࡼࡧᇸ⋢┴❧ᕝࡢ༤≀㤋࡜ᇸ⋢┴❧⮬↛ࡢ༤≀㤋ࡢ࣊ࣅ㢮ࡢእ㒊ᐤ⏕⹸ㄪᰝ㧗ᶫࠉᏲ࣭୕ゅோᏊ㸦ᇸ⋢་⛉኱Ꮫ㸧ࠊ㤿ሙ⿱⨾㸦ᮾ໭኱Ꮫ㸧ࠊዟᮧࡳ࡯Ꮚ㸦ᇸ⋢┴❧⮬↛ࡢ༤≀㤋㸧ࠊ⸨⏣ᏹஅ㸦ᕝࡢ༤≀㤋㸧ࡣࡌࡵ࡟ྠ୍ࡢ⑓ཎయ࡟ࡼࡾࠊࣄࢺ࡜ࣄࢺ௨እࡢ⬨᳝ື≀ࡢ཮᪉ࡀ⨯ᝈࡍࡿឤᰁ⑕ࡀே⋇ඹ㏻ឤᰁ⑕࡛ࠊ࣌ࢫࢺࡸ᪥ᮏ⬻⅖࡞࡝ࡣࡼࡃ▱ࡽࢀ࡚࠸ࡿ⑌⑓࡛࠶ࡿࠋ⌧ᅾࠊࣄࢺ࡬ࡢឤᰁࡀሗ࿌ࡉࢀ࡚࠸ࡿᚤ⏕≀ࡢ⣙㸣㸦⣙✀㸧ࡀື≀࡟⏤᮶ࡍࡿ࡜⪃࠼ࡽࢀ࡚࠾ࡾࠊே⋇ඹ㏻ឤᰁ⑕㸦ື≀⏤᮶ឤᰁ⑕㸧ࡢ㔜せᛶࡀㄆ㆑ࡉࢀ࡚ࡁ࡚࠸ࡿ㸦ᅜ❧ឤᰁ⑕◊✲ᡤ㸪㸧ࠋ့ங㢮ࡸ∐⹸㢮࡟ࡣࠊࣄࢺ࡟ே⋇ඹ㏻ឤᰁ
⑕ࡢ㉳ᅉ࡜࡞ࡿྛ✀⑓ཎయࢆ፹௓ࡍࡿࢲࢽࡸࣀ࣑㢮࡞࡝ࠊᵝࠎ࡞⠇㊊ື≀ࡀᐤ⏕ࡋ࡚࠸ࡿࠋ
ࡋ࠿ࡋ㏻ᖖࠊእ㒊ᐤ⏕⹸ࡢ✀㢮ࡣࠊᐟ୺ື≀ࡢ✀㢮ࡸ⏕ᜥሙᡤࠊࡉࡽ࡟Ꮨ⠇࡟ࡼࡾ␗࡞ࡿሙྜࡀ࠶ࡿࡓࡵࠊᝈ⪅ࡀⓎ⏕ࡋࡓ㝿࡟ࠊ፹௓✀ࢆ᥎ᐃࠊ࠶ࡿ࠸ࡣỴᐃࡍࡿࡓࡵ࡟ࠊࡑࢀࡽࡢእ㒊ᐤ⏕⹸┦ࢆศ㢮Ꮫⓗ࡟᫂ࡽ࠿࡟ࡋ࡚࠾ࡃࡇ࡜ࡣ኱ኚ㔜せ࡛࠶ࡿࠋࡑࡋ࡚ᚓࡽࢀࡓእ㒊ᐤ⏕⹸ࡀࠊ⑓ཎᚤ⏕≀ࢆಖ᭷ࡋ࡚࠸ࡿ࠿࡝࠺࠿ࡢ᳨⣴ࡀ࡛ࡁࢀࡤࠊ⏦ࡋศ࡞࠸ࠋᮏሗ࡛ࡣᇸ⋢┴す㒊ᆅᇦ࡛ᤕ⋓ࡉࢀࡓ࢔ࣛ࢖ࢢ࣐㸦Procyon lotor㸧࡜࢔ࢼࢢ࣐㸦Melesanakuma㸧ࠊ࠾ࡼࡧᇸ⋢┴❧ᕝࡢ༤≀㤋཰㞟࠾ࡼࡧᇸ⋢┴❧⮬↛ࡢ༤≀㤋ᡤⶶ࣊ࣅ㢮ࡢእ㒊ᐤ⏕⹸ࡢㄪᰝࢆ⾜ࡗࡓࡢ࡛ሗ࿌ࡍࡿࠋㄪᰝ᪉ἲᇸ⋢┴❧ᕝࡢ༤≀㤋࡟෭෾ಖᏑࡉࢀ࡚࠸ࡓ࢔ࣛ࢖ࢢ࣐࡜࢔ࢼࢢ࣐ࡣࠊゎ෾ᚋ඲㌟ࢆ⫗║ⓗ࡟⢭ᰝࡋࡓࠋࡲࡓᇸ⋢┴❧ᕝࡢ༤≀㤋ࡀ཰㞟ࡋࡓᶆᮏ࡜ᇸ⋢┴❧⮬↛ࡢ༤≀㤋ᡤⶶ࣊ࣅ㢮㸦෭෾ࡲࡓࡣ࢚ࢳࣝ࢔ࣝࢥ㸫ࣝᾮᾐᶆᮏ㸧ࡣࠊᐇయ㢧ᚤ㙾ୗ࡛⢭ᰝࡋࡓࠋࡑࡢ⤖ᯝࠊㄆࡵࡽࢀࡓእ㒊ᐤ⏕⹸㸦ࣀ࣑㢮ࠊ࣐ࢲࢽ㢮㸧ࡣࡍ࡭࡚㸣࢚ࢳࣝ࢔ࣝࢥ㸫ࣝ࡟ಖᏑࡋࡓࠋࡑࡢᚋࣀ࣑㢮ࡢ඲࡚࡜࣐ࢲࢽ㢮ࡢ୍㒊ࡣࠊ㸣Ỉ㓟໬࣒࢝ࣜ࢘Ỉ⁐ᾮ࡟㹼᪥㛫ᾐࡋ࡚ෆᐜ≀ࢆ⁐࠿ࡋࡓᚋࠊ᪂ࡋ࠸Ỉ࡟⛣ࡋࠊị㓑㓟୍⁲ࢆຍ࠼࡚୰࿴ࡋࡓᚋࠊ෌ᗘ᪂㩭࡞Ỉ࡟᪥㛫ᨺ⨨ࡋࡓࠋࡑࡢᚋ࣐ࢲࢽ㢮ࡣࢫࣛ࢖ࢻࢢࣛࢫୖ࡟࣍࢖ࣖ㸫Ặᾮ㸦ࢣࢽࢫᰴᘧ఍♫ࠊ኱㜰㸧ࢆ㸯㹼㸰⁲⁲ୗࡋ࡚ࠊ࢝ࣂ㸫ࢢࣛࢫᑒධᶆᮏࢆస〇ࡋࡓࠋࣀ࣑㢮ࡣ㸣࢚ࢳࣝ࢔ࣝࢥ㸫ࣝ࡟㸯᫬㛫ධࢀࡓᚋࠊ㸪㸪㸪࡟ྛ㸯᫬㛫ධࢀ࡚⬺Ỉࡋࠊࢫࣛ࢖ࢻࢢࣛࢫୖ࡟ࣘࣃ ࣛ ࣝ ⁐ ᾮ㸦Euparal Solution, WALDECKGmbH & Co. KG, DIVISION CHROMA㸧ࢆ୍⁲⁲ୗࡋࡓ୰࡟ࣀ࣑ࢆධࢀࠊ࢝ࣂ㸫ࢢࣛࢫ࡛ᑒධᶆᮏ࡟ࡋࡓࠋࡇࢀࡽࡢᑒධᶆᮏ࡜࣐ࢲࢽ
ࡢᾮᾐᶆᮏࡣࠊࡑࢀࡒࢀ㏱㐣ᆺගᏛ㢧ᚤ㙾
Nikon Eclipse E600 DIC ࡜ ᐇ య 㢧 ᚤ 㙾㸦Nikon SMZ-U㸧࡛ᙧែほᐹࡋࠊ࣐ࢲࢽ㢮ࡣYamaguti, et al.㸦㸧Voltzit and Keirans ⸨⏣࣭㧗⏣㸦D㸧㧗⏣࡯࠿ ࠊࣀ࣑㢮ࡣSakaguti㸦㸧࡟ᚑࡗ࡚✀ࡢྠᐃࢆ⾜ࡗࡓࠋ⤖ᯝ࡜⪃ᐹϨ㸬࢔ࣛ࢖ࢢ࣐࡜࢔ࢼࢢ࣐ࡢእ㒊ᐤ⏕⹸ㄪᰝ⤖ᯝ㸦⾲㸯㸧࢔ࣛ࢖ࢢ࣐ࡣ㸵㢌ࡢ࠺ࡕ㸴㢌࡟࣐ࢲࢽࡢᐤ⏕ࡀࡳࡽࢀࡓࡀࠊ࢔ࢼࢢ࣐࡟ࡣᐤ⏕ࡋ࡚࠸࡞࠿ࡗࡓࠋᚓࡽࢀࡓ࣐ࢲࢽࡣࢱ࢝ࢧࢦ࣐࢟ࣛࣛ
ࢲࢽAmblyomma testudinariumಶయ㸦඲࡚ⱝ⹸㸧ࠊ࠾ࡼࡧ࢟ࢳ࣐ࢲࢽ+DHPDSK\VDOLVÀDYDࠉಶయ㸦ᡂ⹸㞝ࠊᡂ⹸㞤㸯ࠊⱝ⹸㸧ࡢྜィ㸰✀ಶయ࡛࠶ࡗࡓࠋࡲࡓࣀ࣑ࡣ࢔ࣛ࢖ࢢ࣐㸵㢌ࡢ࠺ࡕ㸳㢌࠿ࡽParaceras melis sinensisಶయ㸦㞝㸰ࠊ㞤㸧࡜Chaetopsylla mikadoࡢ㞤㸯ಶయࡢྜィ㸰✀ಶయࡀᚓࡽࢀࡓࡀࠊ࢔ࢼࢢ࣐࡟ࡣᐤ⏕ࡋ࡚࠸࡞࠿ࡗࡓࠋ௨ୗࠊศ㢮⩌ࡈ࡜࡟ศࡅ࡚グࡍࠋ㸯ࠉ࣐ࢲࢽ㢮ᚓࡽࢀࡓ㸰✀ࡢ࣐ࢲࢽࡢᴫㄝ࡜ᇸ⋢┴࡟࠾ࡅࡿ࣐ࢲࢽ㢮ࡢグ㘓ࢆ௨ୗ࡟♧ࡍࠋ ࢱ ࢝ ࢧ ࢦ ࢟ ࣛ ࣛ ࣐ ࢲ ࢽ Am blyom ma
testudinarium Koch,㸦ᅗ㸯㸧ᖺ᭶᪥࡟࡜ࡁࡀࢃ⏫᪥ᙳ࡛ᤕ⋓ࡉ㸫  㸫㧗ᶫࠉᏲ࣭୕ゅோᏊ࣭㤿ሙ⿱⨾࣭ዟᮧࡳ࡯Ꮚ࣭⸨⏣ᏹஅᇸ⋢┴す㒊ᆅᇦ࡛ᤕ⋓ࡉࢀࡓ࢔ࣛ࢖ࢢ࣐࡜࢔ࢼࢢ࣐ࠊ࠾ࡼࡧᇸ⋢┴❧ᕝࡢ༤≀㤋࡜ᇸ⋢┴❧⮬↛ࡢ༤≀㤋ࡢ࣊ࣅ㢮ࡢእ㒊ᐤ⏕⹸ㄪᰝ⾲㸯ࠉ࢔ࣛ࢖ࢢ࣐࡜࢔ࢼࢢ࣐ࡢእ㒊ᐤ⏕⹸ㄪᰝಶయ✀ྡ᥇㞟ᖺ᭶᪥ᤕ⋓ሙᡤ␒ྕ,' ࢔ࣛ࢖ࢢ࣐  ᇸ⋢┴ẚ௻㒆࡜ࡁࡀࢃ⏫᪥ᙳ,' ࢔ࣛ࢖ࢢ࣐  ᇸ⋢┴ẚ௻㒆࡜ࡁࡀࢃ⏫᪥ᙳ,' ࢔ࣛ࢖ࢢ࣐  ᇸ⋢┴ẚ௻㒆࡜ࡁࡀࢃ⏫᪥ᙳ,' ࢔ࣛ࢖ࢢ࣐  ᇸ⋢┴ẚ௻㒆࡜ࡁࡀࢃ⏫᪥ᙳ ࢔ࣛ࢖ࢢ࣐  ᇸ⋢┴ᮾᯇᒣᕷᒾẊ ࢔ࣛ࢖ࢢ࣐ ᇸ⋢┴ẚ௻㒆࡜ࡁࡀࢃ⏫᪥ᙳ ࢔ࢼࢢ࣐ ᇸ⋢┴ẚ௻㒆࡜ࡁࡀࢃ⏫᪥ᙳ ࢔ࣛ࢖ࢢ࣐ ᇸ⋢┴ẚ௻㒆࡜ࡁࡀࢃ⏫᪥ᙳ*M: male, F: female, N: nymph
A. test : Amblyomma testudinarium ࢱ࢝ࢧࢦ࣐࢟ࣛࣛࢲࢽ+ÀDYD : +DHPDSK\VDOLVÀDYD ࢟ࢳ࣐ࢲࢽ
P. m. sinensis : Paraceras melis sinensis
C. mikado : Chaetopsylla mikado

య㔜 NJ








࣐ࢲࢽ
A. test
5N*
2N
2N
3N
-+ÀDYD1N
2N
3M, 1F, 4N
9M, 7N-ࣀ࣑
P.m.sinensis C. mikado
1M*,14F*
5F
2F
1M, 11F3F1F-ᖺࡢᐤ⏕౛࡛࠶ࡗࡓࠋࡑࢀࡲ࡛ࡣ⣙㢌௨ୖࡢ⭾኱࡞ᩘࡢᐟ୺ື≀ࢆㄪ࡭࡚ࡶࠊ඲ࡃᐤ⏕౛ࡀ࡞࠿ࡗࡓࡇ࡜࠿ࡽ㸦⸨⏣࡯࠿㸪㸹Takahashi and Sekine, 㸹 ⸨ᮏ࡯࠿㸪㸹㧗ᶫ࡯࠿㸪㸧ࠊ㏆ᖺఱࡽ࠿ࡢ࠿ࡓࡕ࡛㐠ࡤࢀࡓಶయࡀᐃ╔ࡋࠊ⦾Ṫࡋࡓࡢ࠿ࡶࡋࢀ࡞࠸ࠋᮏ✀ࡣ༡᪉⣔ࡢ✀࡛࠶ࡿࡓࡵࠊ⦾Ṫ࡛ࡁࡿ᮲௳ࡢ୍ࡘ࡟ࡣ ᬮ໬ࡀ⪃࠼ࡽࢀࡿࡀࠊࡑࢀࡀ஦ᐇ࠿࡝࠺࠿ࡣ௒ᚋࡢㄢ㢟࡛࠶ࡿࠋ୍᪉ࠊศᕸࡢᣑ኱࡟ࡣ࢖ࣀࢩࢩࡢศᕸ࡟ྠㄪࡍࡿഴྥࡀ࠶ࡿࡇ࡜ࡀᣦ᦬ࡉࢀ࡚࠸ࡿࡓࡵᅗ㸯ࠉࢱ࢝ࢧࢦ࣐࢟ࣛࣛࢲࢽ㸦ⱝ⹸㸧$㸪⫼㠃㸹%㸪⭡㠃ࠋྑୖࡢᯟෆ࡟➨㸯⬮ᇶ㒊࡟࠶ࡿ㸰ᮏࡢ≉ᚩⓗ࡞ 㸦Kodama et al., 㸧ࠊ௒ᚋࡢㄪᰝ࡟ࡼࡾࠊᲲ㸦DE㸧ࢆ♧ࡍࠋࡉࡽ࡞ࡿศᕸࡀ☜ㄆࡉࢀࡿྍ⬟ᛶࡀ࠶ࢁ࠺ࠋ࡞࠾ࠊศᕸࡋ࡚࠸ࡿ࡜࠸࠺ሙྜࠊ༢࡟᥇ࢀࡓ㸲㢌ࡢ࢔ࣛ࢖ࢢ࣐࡟ࡣᐤ⏕ࡋ࡚࠸ࡓࡀࠊ㞟ࡉࢀࡓ࠿ࡽࡑࡇ࡟ศᕸࡋ࡚࠸ࡿࢃࡅ࡛ࡣ࡞ᮾᯇᒣᕷᒾẊ࡛ᖺ᭶᪥࡟ᤕ⋓ࡉࢀࡓ㸯ࡃࠊ∵ࡸ㤿࡞࡝ࡢᨺ∾ᆅ࡛ᮏ✀ࡀᐤ⏕ࡋ࡚࠸㢌࠾ࡼࡧᖺ᭶᪥࡟࡜ࡁࡀࢃ⏫᪥ᙳ࡛ᤕࡓሙྜ࡞࡝ࡣࠊ௚ᆅᇦ࡛ᐤ⏕ࡋࡓಶయࡀᣢࡕ⋓ࡉࢀࡓ㸯㢌࡟ࡣᐤ⏕ࡋ࡚࠸࡞࠿ࡗࡓࠋࡇࡢ㎸ࡲࢀࡓྍ⬟ᛶࡶ࠶ࡿࡢ࡛ὀពࢆせࡍࡿࠋศࡇ࡜࠿ࡽࠊ࡜ࡁࡀࢃ⏫᪥ᙳ࡟ᐃ╔ࡋ࡚࠸ࡓಶᕸࢆᐃ⩏ࡍࢀࡤࠊࡑࡢሙᡤࡲࡓࡣࡑࡢᆅᇦ࡛య࡛ࡣ࡞ࡃࠊู࡞ሙᡤ࡛ᐤ⏕ࡋࡓಶయ࡜⪃࠼⦾Ṫࡋࠊ⏕ά⎔ࡀࡲࢃࡗ࡚࠸ࡿ࡜⪃࠼ࢀࡤࡼࡽࢀࡓࠋࡉࡽ࡟ᐤ⏕ಶయᩘࡀᑡ࡞࠸ࡇ࡜࡜ࠊ࠸ࡔࢁ࠺ࠋᡂ⹸࡜ᗂ⹸ࡢᐤ⏕ࡀぢᙜࡓࡽ࡞࠿ࡗࡓࡇ࡜࠿ᮏ✀ࡣ࢔ࢪ࢔኱㝣ࡢ᪥ᮏ࠿ࡽ࢖ࣥࢻࡸ࢖ࣥࡽࠊ୍㒊ࡢ㝈ࡽࢀࡓᆅᇦ࡟ศᕸࡋ࡚࠸ࡓࡶࡢࢻࢿࢩ࢔ࡲ࡛ࡢ ᖏ࠿ࡽ⇕ᖏᆅᇦࡲ࡛ࠊᗈ⠊࡜ᛮࢃࢀࡓࠋᅖ࡟ศᕸࡋ࡚࠸ࡿ Petney and Keirans, ;≉࡟௒ᅇࠊ࡜ࡁࡀࢃ⏫᪥ᙳ࡛ᤕ⋓ࡉࢀࡓ࢔
Kolonin, ; Guglielmone et al. ; Petneyࣛ࢖ࢢ࣐࠿ࡽグ㘓ࡉࢀࡓࢱ࢝ࢧࢦ࣐࢟ࣛࣛࢲ
etal. ࠋᅜෆ࡛ࡣ⚟ᓥ┴㸦ቨ㇂࡯࠿㸧ࠊࢽ࡟㛵ࡋ࡚ࠊᮏ✀ࡣᖺ᭶᪥࡟ẟ࿅ᒣ⏫
㟷 ᳃ ┴ ᑎ ⏣ ࡯ ࠿ ࠊ ໭ ᾏ 㐨㸦Nakao et࡛ᤕ⋓ࡉࢀࡓ࢖ࣀࢩࢩ࡟ᐤ⏕ࡋ࡚࠸ࡓಶయࡀࠊal., 㸧࠿ࡽࡶ᩿∦ⓗ࡞グ㘓ࡀ࠶ࡿࡶࡢࡢࠊ
ᇸ⋢┴࡛ࡢึグ㘓࡛࠶ࡗࡓ㸦Takahashi et al.,ࡑࢀࡽࡢᆅᇦ࡛ᐃ╔ࡋ⦾Ṫࡋ࡚࠸ࡿドᣐࡀ㸧ࠋ㉺⏕⏫࡛ࡢேయ่ည౛㸦ᖺ᭶ᚓࡽࢀ࡞࠸ࡓࡵࠊ୍㐣ⓗ࡟᥇㞟ࡉࢀࡓྍ⬟᪥㸧㸦㧗ᶫ㸪㸧ࢆྵࡵࡿ࡜ࠊᮏ✀ࡣ௒ᛶࡀ㧗࠸ࠋࡑࢀࡺ࠼ࠊᮏ✀ࡢศᕸࡢ໭㝈ࡣࠊᅇࡢグ㘓࡟ࡼࡾࠊす㒊ᆅ༊ࡢேᐙ࡟㏆࠸ୣ㝠௒ࡢ࡜ࡇࢁ㛵ᮾᆅ᪉࡜࠸࠺ࡇ࡜࡟࡞ࢁ࠺ᖏ㸱⟠ᡤ࡛☜ㄆࡉࢀࡓࡇ࡜࡟࡞ࡾࠊࡋ࠿ࡶ㏆㸫  㸫ᕝ༤⣖せࠉྕࠉ㸸

Takahashi et al.,  ᅜෆ࡛ࡢᐟ୺ࡣࠊࣄࢺࢆྵࡴ့ங㢮ࠊ㫽㢮ࠊ∐⹸㢮ࠊ୧⏕㢮࡞࡝ከᒱ࡟ࢃࡓࡿࠋ≉࡟ᡂ⹸ࡣࠊ୺࡟࢖ࣀࢩࢩ࡞࡝ࡢ኱࣭୰ᆺ့ங㢮࡟ᐤ⏕ࡋࠊᗂ࣭ⱝ⹸ࡣ∐⹸㢮ࢆྵࡴ୰࣭ᑠᆺື≀࡟ࡳࡽࢀࡿ㸦㧗⏣࡯࠿ ; Takahashi et al.,㸧ࠋ་Ꮫⓗ࡟ࡣ⣚ᩬ⇕⩌ࡢRickettsia tamuraeࢆ≉␗ⓗ࡟㧗⋡࡟ಖ᭷ࡋࠊࣄࢺ࡬ࡢឤᰁ౛ࡶሗ࿌ࡉࢀ࡚࠸ࡿ Ᏻ⸨࣭⸨⏣㸹▼␇࡯࠿
 ; Imaoka et al., ; 㧗 ᶫ ࠋ ࡲ ࡓ㏆ᖺࡣ࢘࢖ࣝࢫឤᰁ⑕ࠕ㔜⑕⇕ᛶ⾑ᑠᯈῶᑡ⑕ೃ⩌㸦6)76㸧ࠖࡢ፹௓✀ࡢ୍✀࡛࠶ࡿྍ⬟ᛶ ࡶ ᣦ ᦬ ࡉ ࢀ ࡚ ࠸ ࡿ ᅜ ❧ ឤ ᰁ ⑕ ◊ ✲ ᡤ ࠋᮏ✀ࡢᡂ⹸㞤ࡀ༑ศ࡟྾⾑㸦㣬⾑㸧ࡍࡿ࡜ࠊయ㛗PPࠊయ㔜J࡟ࡶ࡞ࡾࠊ⏘༸ᩘࡣ㸰୓ಶࢆ㉸࠼ࡿ㸦㧗ᶫ࡯࠿ᮍⓎ⾲㸧ࠋࣜࢣࢵࢳ࢔ࡸ࢘࢖ࣝࢫࡀࠊࡇࡢ㣬⾑ᡂ⹸࠿ࡽ⤒༸ఏ᧛࡟ࡼࡾᗂ⹸࡟ཷࡅ⥅ࡀࢀࠊ⑓ཎయಖ᭷ಶయࡢⴭࡋ࠸ቑຍ࡟ຍ࠼ࠊࣄࢺ่ညᛶࡀ࠶ࡿࡓࡵࠊ་Ꮫⓗព⩏ࡀ㧗ࡲࡗ࡚࠸ࡿ㸦Ἀ㔝࡯࠿ᅜ❧ឤᰁ⑕◊✲ᡤ㸹㧗⏣࡯࠿㸪㸧ࠋᅗ㸰ࠉ࢟ࢳ࣐ࢲࢽ㸦ⱝ⹸㸧$⫼㠃㸹%⭡㠃㸹&㢡య㒊⭡㠃ࠋཱྀୗ∦ࡢṑิ㸦▮༳㸧ࡣ≉ᚩⓗ࡞㸹'㢡య㒊⫼㠃ࠋ▮༳ࡣゅ≧య࡛ࠊ௚✀࡜ࡢ༊ู࡟౑࠺㸹(➨㸯⬮ᇶ㒊ࡢᲲ ࢟ ࢳ ࣐ ࢲ ࢽ Haemaphysalis flava
Neumann, 㸦ᅗ㸰 㸲㸧ࠉᮏ✀ࡣ࡜ࡁࡀࢃ⏫᪥ᙳ࡛ᖺ᭶࡟ᤕ⋓ࡉࢀࡓ࢔ࣛ࢖ࢢ࣐㸲㢌ࡢ࠺ࡕ㸰㢌࡟ࠊྜィ㸱ಶయࡢⱝ⹸ࡀᐤ⏕ࡋ࡚࠸ࡓ㸬ࡲࡓᅗ㸱ࠉ࢟ࢳ࣐ࢲࢽ㸦㞝㸧ᖺ᭶࡜᭶࡟࡜ࡁࡀࢃ⏫᪥ᙳ࡜ᮾᯇ
$⫼㠃㸹%⭡㠃㸹&ᕥഃࡢ➨㸯㹼㸲⬮ᇶ⠇ࠊ≉࡟➨㸲⬮ᇶ⠇࡟࠶ࡿ㸯ᮏࡢ⣽㛗࠸Ჲࡀ≉ᚩⓗࠋᒣᕷᒾẊ࡛ᤕ⋓ࡉࢀࡓྛ㸯㢌ࡢ࢔ࣛ࢖ࢢ࣐࡟ࠊࡑࢀࡒࢀಶయ㸦ᡂ⹸㞝㸷㸪ⱝ⹸㸵㸧㸪㸶ಶయ㸦ᡂ⹸㞝㸱㸪ᡂ⹸㞤㸯㸪ⱝ⹸㸲㸧ࡢᐤ⏕ࡀぢࡽࢀࠊ࢔ࣛ࢖ࢢ࣐ࡀ⏕ᜥࡍࡿୣ㝠ᆅ࡟ᗈࡃศᕸࡋ࡚࠸ࡿࡶࡢ࡜ᛮࢃࢀࡓࠋᮏ✀ࡣᇸ⋢┴ෆࡢᒣᯘࡢ࠶ࡿᖹᆅࡲࡓࡣୣ㝠ᖏ࠿ࡽࠊᶆ㧗⣙㹫ࡢᒣᆅᖏୖ㒊࡟ࡲ࡛ᖜᗈࡃศᕸࡋ࡚࠾ࡾࠊಶయᩘࡣୣ㝠ᖏࡸᒣᆅᖏୗ㒊࡟ከ࠸ࠋࡑࢀࡣ୺࡞ᐟ୺࡛࠶ࡿࢱࢾ࢟ࠊ࢟ࢶࢿࠊࣀ࢘ࢧࢠࠊࢽ࣍ࣥࢪ࢝࡞࡝ࡢ୰ᆺ㹼኱ᆺ့ங㢮ࡀࠊẚ㍑ⓗከࡃ⏕ᜥࡋ࡚࠸ࡿࡓࡵ࡛࠶ࢁ࠺㸦⸨⏣࡯࠿㸪㸧ࠋ࢔ࣛ࢖ࢢ࣐ ᅗ㸲ࠉ࢟ࢳ࣐ࢲࢽ㸦㞤㸧ࠋ$⫼㠃 %⭡㠃㸹&ᕥഃࡢ➨㸯㹼㸲⬮ᇶ⠇ࠊྛᇶ⠇ࡢࡑࢀࡒࢀ࡟ࡣࡀࡇ࠺ࡋࡓୣ㝠ᆅࡢேᐙ࡟㏆࠸ᒣᯘࢆ⏕ᑠࡉ࠸Ჲࡀ㸯ᮏ࡙ࡘ࠶ࡿ㸦▮༳㸧ࠋᜥᇦ࡟ࡋ࡚࠸ࡿࡇ࡜࠿ࡽࠊᮏ✀ࡀᐤ⏕ࡋ㸫  㸫㧗ᶫࠉᏲ࣭୕ゅோᏊ࣭㤿ሙ⿱⨾࣭ዟᮧࡳ࡯Ꮚ࣭⸨⏣ᏹஅᇸ⋢┴す㒊ᆅᇦ࡛ᤕ⋓ࡉࢀࡓ࢔ࣛ࢖ࢢ࣐࡜࢔ࢼࢢ࣐ࠊ࠾ࡼࡧᇸ⋢┴❧ᕝࡢ༤≀㤋࡜ᇸ⋢┴❧⮬↛ࡢ༤≀㤋ࡢ࣊ࣅ㢮ࡢእ㒊ᐤ⏕⹸ㄪᰝ࡚࠸ࡓࡶࡢ࡜ᛮࢃࢀࡓࠋᮏ✀ࡢᡂ⹸࡜ⱝ⹸ࡢᐤ⏕ಶయᩘࡣࠊ࠸ࡎࢀࡶᇸ⋢┴࡛ࡣ㹼᭶ࡢᖺ㸯ᅇࡢࣆ࣮ࢡࡀぢࡽࢀ㸦⸨ᮏ࡯࠿㸪㸧ࠊ௒ᅇࡢ᥇㞟᭶ࡶ㹼᭶୍࡛⮴ࡋࡓࠋᮏ✀ࡣ໭ᾏ㐨࠿ࡽἈ⦖ࡲ࡛ࠊ඲ᅜ࡟ศᕸ
ࡍࡿᬑ㏻✀࡛࠶ࡿ㸦⸨⏣࣭㧗⏣㸪E㸧ࠋ་Ꮫⓗ࡟ࡣ᪥ᮏ⣚ᩬ⇕ࡢ⑓ཎ⳦࡛࠶ࡿ
Rickettsia japonicaࡸࠊ ⣚ ᩬ ⇕ ⩌ ௨ እ ࡢ
Rickettsia canadensisࡢศ㞳౛ࡀ࠶ࡿ 㧗⏣࡯࠿㸪 ࠋࡲࡓ6)76࢘࢖ࣝࢫࡢ㑇ఏᏊࡀ᳨ฟࡉࢀ࡚࠾ࡾࠊࣄࢺ่ည౛ࡣᚲࡎࡋࡶ
ከ࠸ࢃࡅ࡛ࡣ࡞࠸ࠋࡋ࠿ࡋ㌟㏆࡞࡜ࡇࢁ࡟ ᅗ㸳ࠉ3DUDFHUDVPHOLVVLQHQVLV $㞤ࡢ㢌㒊%㞤ࡢ⏕Ṫჾ&㞝ࡢ㢌㒊'㞝ࡢ⏕Ṫჾከᩘ⏕ᜥࡋ࡚࠸ࡿࡢ࡛ὀពࢆせࡍࡿ Ἀ㔝, Takahashi et al.,   㸧ࠊ࡯࠿㸪Ᏻ⸨࣭⸨⏣㸹ᅜ❧ឤᰁ⑕◊ࡲࡓ Ya m a g u t i e t a l.  ࡢグ㘓࡜ࡋ࡚✲ᡤ ࠋHaemaphysalis campanulata ࡜ Ixodes
acutitarsusࠊࡲࡓᒣᮏ࡯࠿㸦㸧ࡢグ㘓࡜㸦㸧ᇸ⋢┴࡟࠾ࡅࡿ࣐ࢲࢽࡢグ㘓ࡋ ࡚Haemaphysalis kitaokai࡜H. hystricisࢆ ྵᇸ⋢┴࡟࠾ࡅࡿ࣐ࢲࢽࡢ✀㢮࡜ศᕸ࡟㛵ࡋࡵࡿ࡜㸳ᒓ✀࡟࡞ࡿ㸦⾲㸰㸧ࠋ࡚ࡣࠊᖺ࠿ࡽᖺࡲ࡛ࠊᮏ┴ࡢᖹ㔝㒊࠿ࡽ㧗ᒣᖏ࡟⏕ᜥࡍࡿ⛉✀ࡢ့ங㢮㸦㕥ࣀ࣑㢮ᮌ㸧ࡢ࠺ࡕࠊᮍⓎ⾲ศࢆྵࡵࡿ࡜ࠊ࢜ࣀ࣑ࡣ྾⾑࡟ࡼࡾࠊᐟ୺࡟┤᥋ⓗ࡞⿕ᐖࢆࢼ࢞ࢨࣝ⛉ࡢࢽ࣍ࣥࢨࣝࢆ㝖ࡃ⛉✀ࠊ⣙୚࠼ࡿࡔࡅ࡛࡞ࡃࠊ࣌ࢫࢺࠊⓎ⑈⇕ࠊ㔝ඡ⑓ࠊ㢌ࠊ 㫽 㢮⛉ ⣙⩚ ࢆ ㄪ ࡭ ࡓ ⤖ ᯝࠊⓎ⑈ࢳࣇࢫ࡞࡝ࠊከࡃࡢே⋇ඹ㏻ࡢྛ✀⑓㸳 ᒓ✀㸦 ⸨ ⏣ ࡯ ࠿  Takahashi andSekine⸨ ᮏ ࡯ ࠿ 㧗 ᶫ ࡯ ࠿ཎయࢆ፹௓ࡍࡿࡇ࡜ࡀ▱ࡽࢀ࡚࠾ࡾࠊ་Ꮫⓗ ࡟ 㔜 せ ࡞ ୍ ⩌ ࡛ ࠶ ࡿ㸦 B i b i k o v a⾲㸰ࠉᇸ⋢┴࠿ࡽグ㘓ࡉࢀ࡚࠸ࡿ࣐ࢲࢽ㢮Cavanaugh and WilliamHopla
GenusࠉArgasࠉLatreille, 1796ࠉࣄ࣓ࢲࢽᒓ
ࠉA. japonicus <DPDJXWL&OL൵RUGDQG7LSWRQࠉࢶࣂ࣓ࣄ࣓ࢲࢽ
Traub et al., 㸧ࠋࡶࡕࢁࢇ඲࡚ࡢࣀ࣑ࡀࠊ
GenusࠉAmblyommaࠉKoch, 1844ࠉ࢝ࢬ࢟ࢲࢽᒓ
ࣄࢺ࡬ࡢ⑓ཎయࡢ፹௓࡟ಀࡿ࡜ࡣᛮ࠼࡞࠸ࠋ
ࠉA. testudinarium Koch, 1844 ࢱ࢝ࢧࢦ࣐࢟ࣛࣛࢲࢽ
Genus Dermacentor Koch, 1844 ࢝ࢡ࣐ࢲࢽᒓࡋ࠿ࡋࣀ࣑ࡣỗᐟ୺ᛶࡢ✀ࡀከࡃࠊ࡝ࡢ✀ࡶ
ࠉD. bellulus (schulze, 1935) ࣋ࣝࣝࢫ࢝ࢡ࣐ࢲࢽ⑓ཎయࢆ፹௓ࡍࡿྍ⬟ᛶࢆ୍ᴫ࡟ࡣྰᐃ࡛ࡁ
Genus HaemaohysalisࠉKoch, 1844ࠉࢳ࣐ࢲࢽᒓࠉ
࡞࠸ࡇ࡜࠿ࡽࠊྲྀࡾᢅ࠸࡟ࡣὀពࡋࡓ࠸ࠋ
ࠉH. campanulata Warburton, 1908ࠉࢶࣜ࢞ࢿࢳ࣐ࢲࢽ
ࠉ+ÀDYD Neumann, 1987ࠉ࢟ࢳ࣐ࢲࢽ
௒ᅇᚓࡽࢀࡓ㸰✀ࡢࣀ࣑ࡢᴫㄝ࡜ᇸ⋢┴࡟
ࠉH. hystricis Supino, 1897ࠉ࣐ࣖ࢔ࣛࢩࢳ࣐ࢲࢽ
࠾ࡅࡿࣀ࣑㢮ࡢグ㘓ࢆ௨ୗ࡟♧ࡍࠋ
ࠉH. japonica Warburton, 1908ࠉ࣐ࣖࢺࢳ࣐ࢲࢽ
ࠉH. kitaokai Hoogstraal, 1969ࠉࣄࢤࢼ࢞ࢳ࣐ࢲࢽ
ࠉH. longicornis Neumann, 1901ࠉࣇࢱࢺࢤࢳ࣐ࢲࢽ
Genus Ixodes Latreille, 1795 ࣐ࢲࢽᒓ
ࠉI. acutitarsus (Karsch, 1880)ࠉ࢝ࣔࢩ࣐࢝ࢲࢽ
ࠉI. angustus Neumann, 1899ࠉࢺ࣐࢞ࣜࢲࢽ
ࠉI.columunae Takada anf Fujita, 1992ࠉࣁࢩࣈࢺ࣐ࢲࢽ
ࠉI. monospinosus Saito, 1967ࠉࣄࢺࢶࢺࢤ࣐ࢲࢽ
ࠉI. nipponensis Kitaoka and Saito, 1967ࠉࢱࢿ࢞ࢱ࣐ࢲࢽ
ࠉI. ovatus Neumann, 1899ࠉ࣐ࣖࢺ࣐ࢲࢽ
ࠉI. persulcatus Shulze, 1930ࠉࢩࣗࣝࢶ࢙࣐ࢲࢽ
ࠉI. simplex simplex Neumann, 1906ࠉࢥ࣐࢘ࣔࣜࢲࢽ
ࠉI. tanuki Saito, 1964ࠉࢱࢾ࣐࢟ࢲࢽ
ࠉI. turdus Nakatsuji, 1942ࠉ࢔࢝ࢥࢵࢥ࣐ࢲࢽ
ࠉI. LC by Kitaoka, 1979

 Paraceras melis sinensis㸦Liu,  㸦ᅗ㸳㸧࢔ࣛ࢖ࢢ࣐㸵㢌ࡢ࠺ࡕ㸳㢌࡟ᐤ⏕ࡀぢࡽࢀࡓࠋࡇࡢ㸳㢌࡜ࡶ࡜ࡁࡀࢃ⏫᪥ᙳ࡛ᤕ⋓ࡉࢀࡓಶయ࡛ࠊᮾᯇᒣᕷ࡛ᤕ⋓ࡉࢀࡓಶయ࡟ࡣᐤ⏕ࡋ࡚࠸࡞࠿ࡗࡓࠋࡑࡢࡓࡵศᕸ࡟∦ᐤࡀ࠶ࡿ࠿ࡶࡋࢀࡎࠊ௚ᆅᇦࡢಶయ࡜ࡢẚ㍑࡞࡝ࠊ௒ᚋࡢㄪᰝ࡛᫂ࡽ࠿࡟ࡋࡓ࠸ࠋᮏ✀ࡣࢱࢾ࢟ࡸ࢔ࢼࢢ࣐ࠊࢸࣥ࡞࡝ࡢ୰ᆺ့ங㢮࠿ࡽࡢグ㘓ࡀከ࠸ࡶࡢࡢࠊ㢖ᗘࡣ㸫  㸫ᕝ༤⣖せࠉྕࠉ㸸ప ࠸ ࡀ ࢶ ࢟ ࣀ ࣡ࢢ࣐࠿ࡽࡶ グ 㘓 ࡉ ࢀ ࡚ ࠸ ࡿ㸦Sakaguti Sakaguti
and Jameson, 㧗ᶫ࡯࠿㸹ᒣෆ࣭Ụⲡ
ᒣෆ࡯࠿㸧ࠋᇸ⋢┴ෆࡢ⏕ᜥᇦࡣࠊᶆ㧗㹼Pࡢୣ㝠ᖏ࡜ᒣᆅᖏ࡛ࠊேᐙ
࡟᥋ࡍࡿୣ㝠ᖏ࡟ẚ㍑ⓗከ ᅗ㸴ࠉ&KDHWRSV\OODPLNDGR $㞤ࡢ㢌㒊%㞤ࡢ⏕Ṫჾ࠸ഴྥࡀࡳࡽࢀࠊࢱࢾ࢟ࡸࢀࡓࠋᇸ⋢┴ෆ࡛ࡣࢱࢾ࢟ࢆ➹㢌࡟ࠊ࢟ࢶࢿࠊ࢔ ࢼ ࢢ ࣐ ࡢ ⏕ ᜥ ᇦ ࡟ ୍ ⮴ ࡍ ࡿ㸦 㧗 ᶫ ࡯࢔ࢼࢢ࣐ࠊࢸࣥࠊ࢖ࢱࢳࠊ࣒ࢧࢧࣅ࡞࡝ࡢ့࠿㸧ࠋ࡜ࡁࡀࢃ⏫᪥ᙳࠊࡲࡓࡣࡑࡢ࿘㎶ங㢮࠿ࡽグ㘓ࡉࢀ࡚࠸ࡿࠋࡲࡓᇸ⋢┴࡛ࡢᮏ࡟⏕ᜥࡋ࡚࠸ࡿ࢔ࣛ࢖ࢢ࣐ࡢಶయ㛫ࠊ࠶ࡿ࠸✀ࡢ⏕ᜥᇦࡣࠊᶆ㧗㹼Pࡢୣ㝠ᖏ࡜ᒣࡣࢱࢾ࢟࡞࡝ࡢ୰ᆺ့ங㢮࠿ࡽࡢⴠୗಶయࡢ
ᆅᖏ࡛ࠊP. m. sinensis࡜ྠᵝ࡟ࠊୣ㝠ᖏ࡟ẚ෌ᐤ⏕࡟ࡼࡾࠊศᕸࡀᣑ኱ࡍࡿྍ⬟ᛶࡶ࠶ࡾࠊ
㍑ⓗከ࠸ഴྥࡀࡳࡽࢀࠊࢱࢾ࢟࡞࡝ࡢ୰ᆺ့
ࡑࡢືྥ࡟ὀពࡋࡓ࠸ࠋᮏ✀࠿ࡽఱࡽ࠿ࡢ⑓ங㢮ࡢ୺࡞⏕ᜥᇦ࡜୍⮴ࡍࡿ㸦㧗ᶫ࡯࠿㸪ཎయࡢሗ࿌ࡣࡲࡔ࡞࠸ࠋ㸧ࠋ㏆ᖺࠊ࢔ࣛ࢖ࢢ࣐ࡣᮾᯇᒣᕷࢆ୰ᚰ࡜ࡋࡓ
 Chaetopsylla mikado Rothschild, ẚ௻ᆅᇦࡸ⛛∗ᆅᇦ࡛ࡢ⏕ᜥಶయᩘࡀከࡃࠊࠉࠉ㸦ᅗ㸴㸧㞄᥋ࡍࡿ኱㔛ᆅᇦࡸධ㛫ᆅᇦ࡟ࡶ᪤࡟┦ᙜᩘ㸯ಶయ㸦㞤㸧ࡢࡳࡀᖺ᭶᪥࡟ࠊ࡜ࡁ
ࡀ⏕ᜥࡋࠊࡑࡢ௚ࡢᆅᇦ࡟ࡶᛴ㏿࡟ศᕸࡀᣑ
ࡀࢃ⏫᪥ᙳ࡛ᤕ⋓ࡉࢀࡓ࢔ࣛ࢖ࢢ࣐࠿ࡽᚓࡽ኱ࡋ࡚࠸ࡿ ᇸ⋢┴ ࠋ࢔ࣛ࢖ࢢ࣐ࡢ⏕⾲㸱ࠉᇸ⋢┴࠿ࡽグ㘓ࡉࢀ࡚࠸ࡿࣀ࣑㢮ᜥᇦࡀࠊࢱࢾ࢟࡞࡝ࡢ୰ᆺ့ங㢮ࡢ⏕ᜥᇦ࡜␒ྕ✀ࠉࠉࠉࠉྡ
㔜࡞ࡿࡓࡵࠊ௒ᚋ࢔ࣛ࢖ࢢ࣐࡬ࡢᮏ✀ࡢᐤ⏕ Ctenocephalides canis (Curtis, 1826)ࡀ☜ㄆࡉࢀࡿ㢖ᗘࡀ㧗ࡲࡿࡶࡢ࡜ᛮࢃࢀࡿࠋ
 Ctenocephalides felis felis (Bouche, 1835)
 Chaetopsylla (Chaetopsylla) mikado Rothschild, 1904ᅜෆ࡛ࡣࠊᮏ✀࠿ࡽఱࡽ࠿ࡢ⑓ཎయࡢሗ࿌
 Chaetopsylla (Chaetopsylla) jamesoni Sakaguti, 1958ࡣࡲࡔ࡞࠸ࡀࠊ୰ኸ࣮ࣚࣟࢵࣃࡢࢫࣟࣂ࢟࢔
 Stivalius aestivalis Jameson and Sakaguti, 1954 Hystrichopsylla (Hystroceras) ozeana Nakagawa and Sakaguti, 1959ඹ࿴ᅜ࡟⏕ᜥࡍࡿ࢔࢝ࢠࢶࢿ㸦Vulpes vulpes㸧 Stenoponia montana Darskaya, 1949࡟ᐤ⏕ࡋ࡚࠸ࡿChaetopsyllaᒓࡢࣀ࣑࠿ࡽࡣࠊ Stenoponia tokudai Sakaguti and Jameson, 1959
 Rhadinopsylla (Actenophthalmus) japonica Sakaguti and Jameson, 1956≟ ࡢ ឤ ᰁ ⑕ ࡢ ㉳ ᅉ ⳦ ࡜ ᛮ ࢃ ࢀ ࡿBartonella
 Rhadinopsylla (Actenophthalmus) attenuate Jameson and Kumada, 1954
spp.ࡸTheileria sp.࡞࡝ࡢ⑓ཎ㑇ఏᏊࡀ᳨ฟࡉ
 Rhadinopsylla (Actenophthalmus) alphabetica Jameson and Kumada, 1959ࢀ࡚࠸ࡿ Víchová et al., 㸧ࠋࡑࢀࡺ࠼᪥
 Rhadinopsylla (Actenophthalmus) ohnoi Nakagawa, 1957
 Palaeopsylla nippon Jameson and Kumada, 1953ᮏᅜෆ࡛ࡢྠᵝࡢឤᰁ⑕ࡢⓎ⏕ࡢሙྜ࡟ࡣࠊ
 Palaeopsylla miyama Sakaguti and Jameson, 1959ᮏ✀ࡀࡑࡢ፹௓✀ࡢ୍✀࡟࡞ࡿྍ⬟ᛶࡀ⪃࠼
 Palaeopsylla mogura Sakaguti and Jameson, 1960
 Ctenophthalmus (Euctenophthalmus) congener congeneroides Wagner, 1930
ࡽࢀࡿࠋ














Doratopsylla coreana Darskaya, 1949
Neopsylla sasai Jameson and Kumada, 1953
Ischnopsyllus (Ischnopsyllus) elongatus (Curtis, 1832)
Peromyscopsylla udagawai Sakaguti and Jameson, 1959
Frontopsylla (Frontopsylla) nakagawai Kumada and Sakaguti, 1959
Paraceras melis sinensis (Liu, 1935)
Monopsyllus argus (Rothschild, 1908)
Monopsyllus anisus (Rothschild, 1907)
Monopsyllus indages lis Ono, 1958
Ceratophyllus farreni chaoi Smit and Allan, 1955
Ceratophyllus gallinae dilatus Dudolkina, 1946
Ceratophyllus garei Rothschild, 1902
Malaraeus (Amalaraeus) andersoni (Rothschild, 1908)㸦㸧ᇸ⋢┴࡟࠾ࡅࡿࣀ࣑ࡢグ㘓ࣀ࣑ࡣ⠇㊊ື≀㛛᪻⹸⥘ࣀ࣑┠㸦㞃⩟┠㸧࡟ᒓࡍࡿ᪻⹸࡛ࠊᅜෆ࡟ࡣ✀ࡀグ㘓ࡉࢀ࡚࠸ࡿ㸦SakagutiSakaguti and Jameson Takahashi et al.,  ࠋᇸ⋢┴࡛ࡢㄪᰝࡣࠊ့ங㢮࠾ࡼࡧ㫽㢮࡟ᐤ⏕ࡍࡿ࣐ࢲࢽ࡞࡝ࡢእ㒊ᐤ⏕⹸ㄪᰝࡢ୍⎔࡜ࡋ࡚ࠊᖹ㔝㒊࠿ࡽ㧗ᒣᖏࡲ࡛ㄪᰝࢆ⾜࠸ࠊࡇࢀࡲ࡛࡟ᒓ㸫  㸫㧗ᶫࠉᏲ࣭୕ゅோᏊ࣭㤿ሙ⿱⨾࣭ዟᮧࡳ࡯Ꮚ࣭⸨⏣ᏹஅᇸ⋢┴す㒊ᆅᇦ࡛ᤕ⋓ࡉࢀࡓ࢔ࣛ࢖ࢢ࣐࡜࢔ࢼࢢ࣐ࠊ࠾ࡼࡧᇸ⋢┴❧ᕝࡢ༤≀㤋࡜ᇸ⋢┴❧⮬↛ࡢ༤≀㤋ࡢ࣊ࣅ㢮ࡢእ㒊ᐤ⏕⹸ㄪᰝ⾲㸲ࠉᇸ⋢┴❧ᕝࡢ༤≀㤋཰㞟࣊ࣅ㢮ࡢእ㒊ᐤ⏕⹸ㄪᰝಶయ␒ྕ✀ࠉྡ࢔࢜ࢲ࢖ࢩࣙ࢘ࣄࣂ࢝ࣜࢩ࣐ࣟࢲࣛࢪ࣒ࢢࣜࢽ࣐࣒࣍ࣥࢩࢩ࣐࣊ࣅࢽ࣐࣒࣍ࣥࢩ࢔࢜ࢲ࢖ࢩࣙ࢘ࢩ࣐ࣟࢲࣛ࢔࢜ࢲ࢖ࢩࣙ࢘ࢱ࢝ࢳ࣍࣊ࣅ࢔࢜ࢲ࢖ࢩࣙ࢘ࢱ࢝ࢳ࣍࣊ࣅࢩ࣐࣊ࣅࢱ࢖࣡ࣥࣁࣈࢩ࣐ࣟࢲ࣐ࣛࣖ࢝࢞ࢩࢩ࣐࣊ࣅࣄࣂ࢝ࣜࢽ࣐࣒࣍ࣥࢩ࢔࢜ࢲ࢖ࢩ࣐ࣙ࢘ࣖ࢝࢞ࢩࣄࣂ࢝ࣜ࢔࢜ࢲ࢖ࢩࣙ࢘ࢩ࣐ࣟࢲࣛࢩ࣐ࣟࢲࣛ࢔࢜ࢲ࢖ࢩࣙ࢘ᤕ⋓᪥୙᫂୙᫂୙᫂୙᫂୙᫂୙᫂⾲㸳ࠉᇸ⋢┴❧⮬↛ࡢ༤≀㤋ᡤⶶ࣊ࣅ㢮ࡢእ㒊ᐤ⏕⹸ㄪᰝ㈨ᩱ␒ྕ
✀ྡ
5H  ࣄࣂ࢝ࣜ5H  ࢱ࢝ࢳ࣍࣊ࣅ
5H  ࢩ࣐࣊ࣅ5H  ࣄࣂ࢝ࣜ5H  ࢩ࣐࣊ࣅ5H  ࢔࢜ࢲ࢖ࢩࣙ࢘
5H  ࣄࣂ࢝ࣜ5H  ࢩ࣐ࣟࢲࣛ
5H  ࢔࢜ࢲ࢖ࢩࣙ࢘
5H  ࢪ࣒ࢢࣜ5H  ࢩ࣐࣊ࣅ5H  ࢪ࣒ࢢࣜ࡞ࡋ ࡞ࡋ ࣐࣒ࢩ᥇㞟ᖺ᭶᪥୙᫂୙᫂ࡁ࡟ࡣᐤ⏕ࡋ࡚࠸࡚ࡶࠊࢲࢽ㢮ࢆ୺࡜ࡍࡿእ㒊ᐤ⏕⹸ࡢከࡃࡣࠊᐟ୺ࡢṚᚋ▷᫬㛫࡛ᐟ୺࠿ࡽ㞳⬺ࡍࡿࠋࡑࢀࡺ࠼ࠊ஺㏻஦ᨾṚࡋࡓಶయࡸࠊᬻࡃࡢ㛫㣫⫱ࡋࡓᚋ࡟ಖᏑࡉࢀࡓಶయ࡛ࡣࠊእ㒊ᐤ⏕⹸ࢆ᥇㞟ࡍࡿࡇ࡜ࡣ㞴ࡋ࠸ࠋࡋ࠿ࡋࠊ
㏻ᖖ᪂㩭࡞࣊ࣅ㢮ࢆㄪ࡭࡚ࡶࠊᐤ⏕⋡ࡣ኱ኚప࠸ࡢࡀ⌧≧࡛࠶ࡿࠋࡉࡽ࡟࠶ࡿ⛬ᗘࡢᩘࡢ࣊ࣅ㢮ࢆ᥇㞟ࡍࡿࡇ࡜⮬యࡀ㞴ࡋࡃࠊࡑࡢ࠺࠼࣐࣒ࢩࠊࣁࣈࠊ࣑࢘࣊ࣅ㢮࡞࡝ࡢẘ⺬࡟ᑐᛂࡋᚓࡿุ᩿ࡸㄆ㆑ࡶᣢࡗ࡚࠸࡞ࡅࢀࡤ࡞ࡽࡎࠊ⇍⦎⪅௨እ࡛ࡣᤕ⋓ࡀᅔ㞴࡞ሙྜࡀከ࠸ࠋ௒ᅇࡣᕝࡢ༤≀㤋࡜⮬↛ࡢ༤≀㤋࡟ಖᏑࡉࢀ࡚࠸ࡿಶయࢆㄪ࡭ࡓࡀࠊṧᛕ࡞ࡀࡽእ㒊ᐤ⏕⹸ࡣᚓࡽࢀ࡞࠿ࡗࡓࠋᅜෆࡢ࣊ࣅ㢮࡟ᐤ⏕ࡍࡿࢲࢽ㢮ࢆ୺࡜ࡍࡿእ㒊ᐤ⏕⹸ࡢ᝟ሗ㔞ࡣᑡ࡞࠸ࡢࡀ⌧≧࡛ࠊࡇࢀࡲ࡛ࡢㄪᰝ࡛ࠊ࣐ࢲࢽ㢮ࡣࢱ࢝ࢧࢦ࣐࢟ࣛࣛࢲࢽ㸦A. testudinarium㸧ࠊ࣓࣐࢝࢟ࣛࣛࢲ
ࢽ㸦A. geoemydae㸧࠾ࡼࡧ࣑࢘࣊ࣅ࣐࢟ࣛࣛ
ࢲࢽ㸦A. nitidum㸧ࡢ㸱✀ 㛗㇂ᕝ໭ᒸ࣭㕥 ᮌ Kitaoka  㕥 ᮌ ࡯ ࠿ Takahashi, et al 㸪㸪
Kodama et al  ࠊࢶࢶ࣒࢞ࢩ㢮ࡣ⫵ᐤ⏕ࢆྵࡵࡿ࡜㸵✀㸦㧗ᶫ࡯࠿㸪ᮍⓎ⾲㸧ࡢᐤ⏕ࡀ☜ㄆࡉࢀ࡚࠸ࡿࠋࡇࢀࡽࡢㄪᰝሗ࿌࠿ࡽࠊ௒ᅇ࢔ࣛ࢖ࢢ࣐࠿ࡽᚓࡽࢀࡓࢱ࢝ࢧࢦ࣐࢟ࣛࣛࢲࢽࡣࠊ㝣ୖࡢ࣊ࣅ㢮࡟ࡶᐤ⏕ࡍࡿࡇ࡜ࡀࢃ࠿ࡗ࡚ࡁࡓࠋࡋ࠿ࡶᐤ⏕ࡋ࡚࠸ࡿࡢࡣⱝ⹸ࡔࡅ࡛࠶ࡗࡓࠋ࡞ࡐⱝ⹸ࡋ࠿ᐤ⏕ࡋ࡚࠸࡞࠸ࡢ࠿ࠋ࠶ࡿ࠸ࡣ࡞ࡐ㔝እ࡟ᡂ⹸ࡸᗂ⹸ࡀከᩘ⏕ᜥࡋ࡚࠸ࡿ࡟ࡶ࠿࠿ࢃࡽࡎࠊ㔝⏕ಶయ࡟ᐤ⏕ࡋ࡚࠸࡞࠸ࡢ࠿ࠋ୍᪉ࠊࢶࢩ࣐࣐࣒ࢩ࡛ࡣࠊᐤ⏕ࡋ࡚࠸ࡿⱝ⹸ࡢ඲࡚ࡀᐤ⏕ᚋ㸯㐌㛫௨ෆ࡟Ṛஸࡍࡿ⌧㇟ࡀࡳࡽࢀ࡚࠾ࡾࠊᐟ୺ഃࡢ࣐ࢲࢽᐤ⏕࡬ࡢ㜵ᚚ཯ᛂࡀ㔝⏕ࡢ࣊ࣅ࡛ࡣࠊୡ⏺࡛ึࡵ࡚☜ㄆࡉࢀࡓ㸦Kodama et al 㸧ࠋ࣊ࣅ㢮ࠊࡦ࠸࡚ࡣ∐⹸㢮ࡢእ㒊ᐤ⏕⹸ࡢㄪᰝࡣࠊ༢࡟ᐤ⏕ᤕ⋓ሙᡤ⤖ᯝᇸ⋢┴኱㔛㒆ᐤᒃ⏫ᑠᅬࠉᕝࡢ༤≀㤋࡞ࡋᇸ⋢┴ධ㛫㒆ẟ࿅ᒣ⏫୰ኸ࡞ࡋᇸ⋢┴኱㔛㒆ᐤᒃ⏫ᑠᅬࠉᕝࡢ༤≀㤋࡞ࡋᇸ⋢┴኱㔛㒆ᐤᒃ⏫ᑠᅬࠉᕝࡢ༤≀㤋࡞ࡋᇸ⋢┴኱㔛㒆ᐤᒃ⏫ᑠᅬࠉᕝࡢ༤≀㤋࡞ࡋ⩌㤿┴㧗ᓮᕷྜྷ஭⏫ୖዟᖹ࡞ࡋᇸ⋢┴㣤⬟ᕷ㣤⬟࡞ࡋᇸ⋢┴኱㔛㒆ᐤᒃ⏫ᑠᅬ࡞ࡋᇸ⋢┴኱㔛㒆ᐤᒃ⏫ᑠᅬ࡞ࡋᇸ⋢┴኱㔛㒆ᐤᒃ⏫ᑠᅬࠉᕝࡢ༤≀㤋࡞ࡋ୙᫂࡞ࡋᇸ⋢┴኱㔛㒆ᐤᒃ⏫ᑠᅬࠉᕝࡢ༤≀㤋࡞ࡋᇸ⋢┴኱㔛㒆ᐤᒃ⏫⛅ᒣ࡞ࡋᓥ᰿┴ฟ㞼ᕷୖᓥ⏫
࡞ࡋ୙᫂࡞ࡋᇸ⋢┴ධ㛫ᕷ௖Ꮚ࡞ࡋᓥ᰿┴ฟ㞼ᕷୖᓥ⏫
࡞ࡋᇸ⋢┴ᆏᡞᕷ㛗ᒸ࡞ࡋᇸ⋢┴኱㔛㒆ᐤᒃ⏫㔠ᑿ࡞ࡋ୙᫂࡞ࡋ୙᫂࡞ࡋᇸ⋢┴ධ㛫㒆㉺⏕⏫㯮ᒣ࡞ࡋᒣᙧ┴す⨨㈷㒆ᑠᅜ⏫㊊㔝Ỉ࡞ࡋᇸ⋢┴኱㔛㒆ᐤᒃ⏫ᑠᅬࠉᕝࡢ༤≀㤋࡞ࡋᇸ⋢┴ẚ௻㒆࡜ࡁࡀࢃ⏫᪥ᙳ࡞ࡋᇸ⋢┴ධ㛫㒆ẟ࿅ᒣ⏫㙊໭࡞ࡋᮾி㒔ᑠ㔠஭ᕷ㈏஭໭⏫ࠉᮾிᏛⱁ኱㎰ᅬụࠉࠉ࡞ࡋ᥇㞟ሙᡤ୧ୖᮧ㛗™⏫㔝ୖⓙ㔝⏫᪥㔝ἑⓙ㔝⏫᪥㔝ἑඣ⋢㒆⚄ἨᮧⳭⵦ⏫ኳ⋤ᒣሯྂቡ༡ᇸ⋢㒆Ⳮⵦ⏫
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Bibikova, V. A.   Contemporary views on the
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human and animal diseases. Ann. Rev. Entomol
Cavanaugh, D. C. and Williams, J. E.   Plague:
Some ecological interrelationships. In: Traub, R. andStarcke, H. ed  Fleas: SS A. A.
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Petney, T.N., Estrada-Peña, A. and Horak, I.G.  The hard ticks of the world Acari: Ixodida:Ixodidae SS., Springer, Dordrecht, Heidelberg,
New York and London㛗㇂ᕝⱥ⏨㸦㸧୚㑣ぞᓅ⏘ࣜࣗ࢘࢟ࣗ࢘࢔࢜࣊
ࣅ࡟ᐤ⏕ࡋ࡚࠸ࡓ࣐࢟ࣛࣛࢲࢽᒓ࣐ࢲࢽ࡟ࡘ࠸࡚㸬AKAMATAHopla, C. E.  ࠉFleas as vectors of Tularemia inAlaska. In: Traub, R. and Starcke, H. ed  Fleas:SS.A. A. Balkema, Rotterdam, Netherlands
Imaoka K, Kaneko S, Tabara K, Kusatake K, MoritaE  7KH¿UVWKXPDQFDVHRIRickettsia tamurae
infection in Japan. Case Reports in Dermatology  ̽▼␇ࠉྐ㸪⸨⏣༤ᕫ㸪ᖹ㔝ᫎᏊ㸪▮㔝Ὀᘯ㸪㧗⏣ఙᘯ㸦㸧⚟஭┴࡟࠾ࡅࡿ᪥ᮏ⣚ᩬ⇕ࡢឤᰁ⎔ㄪᰝ㸬⚟஭┴⾨⎔◊ᖺሗ㸪㸸㸬ቨ㇂ᫀᙪ㛛㤿┤ኴ⇃⏣♩Ꮚ᰿ᮏࠉ၏㰻⸨᲍⤮௰ᮧࠉ✲㔠ගᩗ஧⸨⏣༤ᕫ㸦㸧⚟ᓥ┴࡟࠾ ࡅ ࡿ 㔝 ⏕ ື ≀ ᐤ ⏕ ࣐ ࢲ ࢽ 㢮 ࡢ ㄪ ᰝ⾨ ⏕ ື≀㸸໭ᒸⱱ⏨㸪㕥ᮌࠉ༤㸦㸧༡すㅖᓥ࡟࠾ࡅࡿ་ື≀Ꮫⓗ◊✲㸬዆⨾኱ᓥ༡㒊ࡢ࣐ࢲࢽ࡜Ꮨ⠇ᾘ㛗㸬⾨⏕ື≀㸪㸸
Kitaoka S.   Ticks Ixodoidea  In: Sasa etal. ed , Animals of medical importance in the Nansei
Islands in Japan. SS.  Shinjuku Shobo,Tokyo. Japan.
Kodama, T., Takahashi, M. and Mori, A.   Host
resistance to ticks Acari: Ixodidae in a pit viper,
Gloydius tsushimaensis, Reptilia: Squamata:Viperidae observed in the field. Int. J. Acarol., 
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Kolonin, G.V.  Fauna of ixodid ticks of the world.
http://www.kolonin.org/ August    Last
accessed  Jun Nakao R, Shinjo K, Sakiyama T, Ogata S, Kusakisako K,Kinoshita G, Naguib D, Chatanga E, Mohamed
WMAM, Moustafa MAM, Matsuno K, Ito T, Nonaka
N, Sashika M, Tsubota T, Shimozuru M. Amblyomma testudinarium infestation on a brownbear Ursus arctos yesoensis captured in Hokkaido,
a northern island of Japan. Parasitology International,
Ἀ㔝ဴஓ㸪ᚋᕝࠉ₶㸪ⓗሙஂ⨾Ꮚ㸪ึ㮵ࠉ஢  ࠉᮏ㑥࡟࠾ࡅࡿ࣐ࢲࢽேయᐤ⏕౛ࡢᴫほ̿ᩥ⊩ⓗ⪃ᐹ̿㸦㸧ࢱ࢝ࢧࢦ࣐࢟ࣛࣛࢲࢽ่⑕౛ᕝᓮ
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ࢽ่⑕౛ᕝᓮ་఍ㄅ㸸
Petney, TN. and Keirans, JE.  Ticks of the genera
Amblyomma and Hyalomma from South East Asia.
Trop. Biomed., ̽
Petney TN, Saijuntha W, Boulanger N, Chitimia-Dobler㸫  㸫㧗ᶫࠉᏲ࣭୕ゅோᏊ࣭㤿ሙ⿱⨾࣭ዟᮧࡳ࡯Ꮚ࣭⸨⏣ᏹஅᇸ⋢┴す㒊ᆅᇦ࡛ᤕ⋓ࡉࢀࡓ࢔ࣛ࢖ࢢ࣐࡜࢔ࢼࢢ࣐ࠊ࠾ࡼࡧᇸ⋢┴❧ᕝࡢ༤≀㤋࡜ᇸ⋢┴❧⮬↛ࡢ༤≀㤋ࡢ࣊ࣅ㢮ࡢእ㒊ᐤ⏕⹸ㄪᰝ/3IH൵HU0(DPXGRPNDUQ&$QGUHZV5+$KDPDG
M, Putthasorn N, Muders SV, Petney DA, Robbins
RG.㸦㸧Ticks (Argasidae, Ixodidae) and tickborne diseases of continental Southeast Asia. Zootaxa,
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Sakaguti, K.  A monograph of the Siphonaptera ofJapan, plate , The Nippon Printing andPublishing Co., Ltd., Osaka, Japan.
Sakaguti, K. and Jameson, E. W.   The
6LSKRQDSWHUDRI-DSDQ3DFL¿F,QVHFWV0RQRJUDSK SS., Entomology Department, Bernice P. Bishop
Museum, Honolulu, Hawaii, U. S. A.
㕥ᮌࠉἨ㸪ᒣᮏṇᝅ㸪ሷᒣ㝧Ꮚ㸪⸨⏣༤ᕫ㸦㸧ᐑᓮ┴ෆࡢ㸯ᆅᇦ࡟࠾ࡅࡿ࣊ࣅ㢮ᐤ⏕࣐ࢲࢽ࡟㛵ࡍࡿㄪᰝ㸬⾨⏕ື≀㸪 6XSSO 㕥 ᮌ Ḡ ྖ㸦㸧 ᇸ ⋢ ࡢ ့ ங 㢮㸬 ᇸ ⋢ ┴ ື ≀ ㄅࠉᇸ⋢┴ᩍ⫱ጤဨ఍㸪SS㧗⏣ఙᘯ ⸨⏣༤ᕫ 㧗ᶫࠉᏲ ኟ⛅ࠉඃ㸦㸧࣐ࢲࢽᡂ⹸ࡢᒓ✀࡬ࡢ᳨⣴࡜ゎㄝࠉ㧗⏣ఙᘯ㸦⦅㸧㸪་ࢲࢽᏛᅗ㚷 SS ໭㝯㤋 ᮾிࠉ㧘ᶫࠉᏲ ᒣᮏ㈆ྖ ᩧ⸨ࠉ㈗ ⏫⏣࿴ᙪ㸦㸧ᇸ⋢࠾ࡼࡧ⩌㤿┴ୗ࡟࠾ࡅࡿ့ங㢮࠾ࡼࡧ㫽㢮࡟ᐤ⏕ࡍࡿࣀ࣑㢮 ࠉ୰࣭኱ᆺ့ங㢮࡟ᐤ⏕ࡍࡿࣀ࣑㢮ࠉ኱ཎᖺሗ㸸
Takahashi, M. and Sekine, M.  Ectoparasite fauna
on mammals and birds in Saitama and Gunma
Prefectures, Central Japan. Part  Ixodid ticks(Acarina; Ixodoidea). Bull. Kawagoe Senior HighSchool. 
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Takahashi, M., Misumi, H., Kaito,T., Kuriyma, T. andFujita, H.   First record of Amblyomma
geoemydae tick on the snake Dinodon semicarinatum,with records of ticks on snakes in Japan. Annu. Rep.
Ohara Hosp. Takahashi M, Kuriyama T, Fujita H.㸦㸧 Ticks(Acari: Ixodidae) on terrestrial snakes in the Izu
Peninsula, Japan. Annu. Rep. Ohara Hosp. 
Takahashi, M., Misumi, H., Kawai, K. and Sato, M.
 The first finding of a bat flea Myodopsylla
trisellis Siphonaptera: Ischnopsyllidae on Myotis
gracilis Chiroptera: Vespertilionidae in Japan. Med.
Entomol. Zool., ̼Takahashi, M., Misumi, H. and Tamura, H.  First
record in Japan of tortoise tick Amblyomma
geoemydae Cantor Acari: Ixodidae
parasitizing Pryer’s keelback snake Hebius pryeri
Boulenger,  Reptile: Colubridae  Int. J.

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Takahashi, M., Misumi, H., Natsume, M., Machida, K.,
Kazama, K., Kazama, M., Kawai, K., Iwasawa, D.,
Sato, M. and Fujita, H.  Preliminary record ofIxodes simplex simplex Neumann,  Acari:Ixodidae  on bats Chiroptera: Vespertilionidae inHokkaido, Aomori, Gunma, Saitama, and Gifu
Prefectures, Japan. Annu. Rep. Ohara Hosp., 㧗ᶫࠉᏲ   㛵ᮾࡢࢶࢶ࣒࢞ࢩ⑓ࡢ⌧ἣࠊຍ࠼࡚࣐ࢲࢽ㢮ࡢࢺࣆࢵࢡࢫ 6$', ࢽ࣮ࣗࢫ㸦ᖺ ᝟ ሗ ஺ ᥮㸧 http://www.sadi-web-site.
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 First record of the hard tick Amblyomma
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