Huffman, Michael A.

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1

CURRICULUM VITAE

Michael A Huffman
Google Scholar Citations (last updated 2022.3.1)
All Since 2017
Citations
10070
3516
h-index56
31
i10-index
157
111

Personal:
Born 1958, Denver, Colorado
Mailing Address:

Inuyama Campus
Kyoto University41-2 Kanrin
Inuyama, Aichi 484
JAPAN
Tel: +81(568) 63-0354
Fax: +81(568) 63-0085
email: huffman.michael.8n@kyoto-u.ac.jp

Home page:https://www.wrc.kyoto-u.ac.jp/en/members/HuffmanMA.html

Current Status:
Associate Professor (2022.4.1~) Wildlife Research Center (Inuyama Campus),
Kyoto University.
Associate Membership to the School of Graduate Studies, University of Lethbridge, Lethbridge,
Canada (April 2017~)
Adjunct Professor, Universita degli Studi di Scienze Gastronomiche, Masters Course of
Gastronomy "Food Ecologies and Sovereignty", Pollenzo, Italy (2017~ current)
Scientific advisory board member, Austrian Research Center for Primatology, Ossiach, Austria(2016~ current)
PhD Co-Supervisor, Keble College, Oxford University, Oxford, UK.
(2021, March~ current).
Previous affiliations:
Associate Professor (2001.9.16 – 2022.3.31), Section of Social Systems Evolution, PRI, Kyoto
University
Director (2021.4.1 – 2022.3.31), Center for International Collaboration and Advanced Studies inPrimatology (CICASP), PRI, Kyoto University (2021.4.1~ 2023.3).
Adjunct Faculty, National Institute of Advanced Studies, Indian Institute of Science Campus,
Bangalore India (from September 2011~2012)
Director, Japanese Macaque Field Research Centre, Primate Research Institute, Kyoto
University (2005.4~ 2007.3)
Visiting Lecturer, University of Toyama, School of Humanities (2002.10.09~2003.3.31).
Visiting Lecturer, Hamamatsu University School of Medicine (2000.4.1~2003.3.31).
Adjunct Professor, Section of Behavioral Development, Center for Human Evolution ModelingResearch, Primate Research Institute, Kyoto University (1999.5 ~ 2001.8).

Huffman, Michael A.

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Visiting Lecturer, Hiroshima City University, Ecological Anthropology & Primatology (1996.9~ 1999.6)
Adjunct Research Associate Professor, University of Colorado-Denver, Department of
Anthropology (1993.3~2001.9)
Education:
1979 (Japanese studies), Kansai University of Foreign Language Studies, Hirakata, Osaka Japan
1983 B.Sc. (Biology), Fort Lewis College, Colorado
1986 M.Sc. (Zoology), Kyoto University, Japan
1989 D.Sc. (Zoology), Kyoto University, Japan
Languages Spoken: English (native), Japanese (professional fluency), Swahili (basic working
fluency)
Professional Organizations and Duties:
American Primatological Society (Associate Editor 2004~)
Primate Society of Japan (Editorial Secretary 2004~2015)
PRIMATES Associate Editor (2002~2005; 2015~)
Advisory Board (1999~2001)
Scientific Reports (Editorial Board 2012~2021)
African Primates (Editorial Board 2013~)
Asian Primate Journal (Editorial Board 2013~)
Conference Papers in Biology- Zoology Section (Editorial Board)
Anthropologial Science (Associate Editor 2012~)
Journal of Ethnobiology and Ethnomedicine (Associate Editor)
Vidyodaya Journal of Humanities and Social Sciences (Editorial Board, 2020.2~)
Borneo Journal of Medical Sciences (Editorial board (2021~2023)
IUCN /SSC & Re-introduction Specialist Group (RSG): (Serving member-consultant)
IUCN SSC Primate Specialist Group - Africa Section: (Serving member-consultant 2014~)
IUCN SSC Primate Specialist Group – South Asia, South East Asia Sections: (Serving
member-consultant 2021~)
International Primate Society (Lifetime Member)Royal Asiatic Society of Sri Lanka (Lifetime Member 2011.9~)
International Society of Ethnopharmacology (Editorial Consultant)Societe Francaise d'Ethnopharmacologie (Scientific advisory board)
Kyoto University English Newsletter ‘Rakuyu’ (Senior Editor - 2007-2009)
Society for Wildlife and Nature (S.W.A.N. Taiwan) International AdvisorAwards & Recognition:
1993 Primate Society of Japan's Award for Overseas Research
1997 Frucht Memorial Lecturer (Key-Note Speaker), University of Alberta
1999 Primate Society of Japan's “Takashima Prize”
2000 ‘The Asia 500’ Baron’s Special 2000 Award for Global Achievement
2001 Presidential Award ‘500 Great Minds’ Barons Who’s Who (USA)2007 ~ Lifetime Honorary member of Associazone Primatologica Italiana
2013 Primate Most Cited Paper Award “Monkeys in the middle”,
2016 Primates Most-Cited Paper Award: “The effect of urban and rural habitats andresource type on activity budgets of commensal rhesus macaques (Macaca mulatta) in
Bangladesh. Primates 54:49–59. MF (Jaman and M. A. Huffman)2018 Primates Social Impact Award “Beneficial effects of hot spring bathing on stress levels inJapanese macaques. 59:215-225. (Takeshita, RSC, Bercovitch FB, Kinoshita, K, Huffman
MA).
Journals refereed for:

Huffman, Michael A.

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3African Journal of Plant Science, American Journal of Physical Anthropology, American
Journal of Primatology, American Institute of Biological Sciences, ANIMAL, Animal Behavior,
Anthropological Science, Appetite, Journal of Archaeological Science, Asian Journal ofPrimatology, Behaviour, Behavioral and Brain Science, Behavioral Processes, Behavior andPhysiology, Biology Letters, Biotropica, Brazilian Journal of Pharmacognosy, Chemical
Ecology, Child Development Research, Current Anthropology, Current Biology, Current
Clinical Pharmacology, EcoHealth, Emerging Infectious Diseases, Ethology, Evolution &Human Behavior, Evolutionary Anthropology, Folia Primatologica, Frontiers in Ecology andthe Environment, Helminthologia, International Journal of Comparative Psychology,
International Journal of Primatology, International Journal of Parasitology, Journal of
Ethnobiology, J.S. Guggenheim Memorial Foundation-Canada, Journal of Ethnopharmacology,
L.S.B. Leakey Foundation, Journal of Malaysian Science, Journal of Threatened Taxa (JoTT),
Mammalia, Mammal Study Natural Sciences and Engineering Research Counsel (Canada),
Molecular Ecology Resources, National Science Foundation (USA), Neuroscience, Parasite,
Parasitology, Philosophical Transactions of the Royal Society B, PloSOne, Primates,
Primatologie, Proceedings of the Royal Society B: Biological Sciences, Range Ecology andManagement, Scientific Research and Essays, Scientific Reports, The Natural Environment
Research Council (UK), International Journal of Nutrition and Metabolism, Trends in
Evolutionary Anthropology, Wildlife Conservation- Japan, Journal of Virology, International
Journal of Physiological Anthropology,
Diploma & Graduate Degree Students, External PhD Advisor:
1. Teruyoshi Takagaki (Masters in Agricultural Sciences, 1991) "Chemical constituents ofVernonia amygdalina, a possible medicinal plant used by wild chimpanzees", Faculty ofAgriculture, Kyoto University [collaborating advisor]
2. Mitsuo Jisaka (Doctor of Agricultural Sciences, 1992) "Bioactive Constituents of Vernonia
amygdalina, a possible medicinal plant used by wild chimpanzees.", Faculty of Agriculture,
Kyoto University [collaborating advisor]
3. Daisuke Izutsu (Masters in Agricultural Sciences, 1993) "Steroids, their related compounds
and antiparasitic activity of Vernonia amygdalina, a medicinal plant used by chimpanzees in
the wild" , Faculty of Agriculture, Kyoto University. [collaborating advisor]
4. Gail Renehan (VM, Tufts University, Boston, U.S.A.) "Evaluation of the potential for the useof medicinal plants in the treatment of parasite infections in captive chimpanzees of African
Zoos and Sanctuaries, 1995, 1996" [external advisor]
5. Michelle G. Burrows (DVM, Glasgow University, U.K.) "Investigation of the intestinal
parasites of chimpanzees in the Budongo Forest, Uganda- 1996" [external advisor]
6. Jessica Zippin (Masters in Geology, 1999) "Geochemistry of termite mound soils ingested bychimpanzees of the Mahale Mtns.- 1995, 1996", York University, Canada. [co-advisor]
7. Laura Ketch (Masters in Botany, 2000) "Microbiology of termite mound soils ingested bychimpanzees of the Mahale Mountains. - 1996", University of Toronto, Canada. [co-advisor]
8. Linda Turner (MSc 1994) "Laterality in Activities of the Japanese Macaques of Arashiyama,(Ph. D received March 2000) “Feeding Ecology of chimpanzees in the Mahale Mountains,
Tanzania 1994 - 1997.” Dept. of Zoology, Kyoto University. [co-advisor]
9. James Wakibara (MSc 2000) (PhD. 2003) The adaptive significance of Geophagy in Japanese
Macaques of Arashiyama-; Seed dispersal of chimpanzees in the Mahale Mountains.” Dept. of
Zoology, Kyoto University. [co-advisor]
10. Ivona Foitova (PhD 2003): “Host-parasite relationship and self-medication in the wild

Huffman, Michael A.

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4orangutan of Northern Sumatra”, Brno University, Brno, Czech Republic [external advisor]
11. Sylvia K. Vitazkova (PhD 2005): “Ethnobotany, zoopharmacognosy of wooly monkeys in
Belize.” Columbia University, NY [co-advisor, examiner]
12. Amani Kitagile (MSc 2005, University of Dar es Salaam, Dept. Zoology): Ecology andconservation of colobus and guenons of the Amani Nature Reserve, Tanga, Tanzania.
[external advisor]
13. Liza R. Moscovice (PhD 2005, University of Wisconsin-Madison) “Ecology and Behaviorof wild chimpanzees on Rubondo Island, Tanzania”,. [collaborator, co-advisor with Chuck
Snowden]
14. Tom Gillespie (PhD 2005 Univ. Florida-Gainesville) Colobine and Guenon parasite
infections, foraging behavior and anthelmintic properties of the diet.”, University of Florida,
Gainesville [external advisor and external examiner]
15. Sabrina Krief (PhD 2005, Museum of Natural History, Paris) “Kibale chimpanzee health,pharmacognosy of diet and self-medication in the Kibale Forest, Uganda” [collaborator andco-advisor]
16. Charmalie A.D. Nahallage (MSc 2005, PhD 2008, Kyoto University, Primate ResearchInstitute,) “Stone handling behavior in Japanese macaques: biological, environmental andsocial perspectives of a cultural behavior” Principal advisor
17. Jared Bakuza (MSc 2006, University of Dar es Salaam, Dept. Zoology) “Ecologicalparasitology of chimpanzees in Gombe National Park, Tanzania.” [external advisor]
18. M. Firoj Jaman (MSc March 2006, PhD 2010, Kyoto University, Primate ResearchInstitute,) "The Affect of Environmental Differences on Behavioral Budgets and the Feeding
Strategies of Captive Japanese Macaques (Macaca fuscata): A Social-Ecological Approach toAssessing the Contribution of Natural Vegetation as Social and Nutritional Enrichment."
Principal advisor
19. Christine J. Rheinhart (PhD 2008, University of Lethbridge, Alberta CA) “Cooperation andcompetition during play fighting in tonkean and Japanese macaques: an examination ofjuvenile behaviour within egalitarian and despotic social systems” [external examiner]
20. Damiana FC Ravasi (PhD 2009, University of Cape Town, Cape Town SA)“Gastrointestinal parasite infection in chachma baboons (Papio h. ursinus) of Cape Peninsula,South Africa: the influence of individual, troop and anthropogenic factors. [external examiner]
21. Andrew J.J. MacIntosh (PhD 2011, Kyoto University, Primate Research Institute) “Healthmaintenance, disease ecology, and the potential for self-medication in Macaca fuscata yakui”.
Principal advisor
22. Jaime Castillo Garcia (MSc 2011, University of Malaya) The Malasian slow loris(Nyctocebus coucang coucang) in the wild: A study combining radio telemetry and molecularbiology. [external examiner]
23. Paula Anne Pebsworth (PhD 2012, Kyoto University, Primate Research Institute) “An
investigation of geophagic and ranging behavior of chacma baboons (papio hamadryasursinus) living in a human-modified habitat. Principal advisor
24. Ilona Profusova (MVS 2012) Department of Parasitology, University of Veterinary andPharmaceutical Sciences, Brno, Czech Republic) Biology and Ecology of TroglodytellaHuffman, Michael A.

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5abrassarti in the common chimpanzee. [external examiner]
25. Rafaela Sayuri Cicalise Takeshita (M.Sc. 2013, Kyoto University, Primate ResearchInstitute) Non-invasive analysis of adrenal hormones in captive Japanese macaques. Principal
advisor w/ Fred Bercovitch
26. Fany Brotcorne (PhD 2014, Behavioral Biology Unit, University of Liege, Belgium)
Human-primate interface, long-tailed macaque behavioural ecology and parasitology.
[External committee, defense jury member, defended and examined on 2014.9.23]
27. Yena Kim (PhD Kyoto University, Primate Research Insitute) Prosocial behavior in captive
orangutan. (Co-advisor, defense committee member, 6.2017).
28. Josué Samuel Alejandro Pastrana (MSc, Kyoto University, Primate Research Institute)
Behavioral assessment of the effect of different environments on the stress levels male
Japanese macaques. (Principal advisor)
29. Azucena G. Enríquez Magaña (Masters candidate, Centro de Estudios e Investigaciones en
Comportamiento CUCBA-Universidad de Guadalajara, Mexico) Fur rubbing behavior in
captive spider monkeys in the Guadalajara Zoo. [co-supervisor with Prof. Gerardo A Ortiz
Rueda]
30. Himani Nautiyal (M.Sc. 2015.5 Wildlife Biology, A.V.C College Mayiladuthurai, Tamil
Nadu, India) Life High Up In: The Himalayas: Factors Affecting The Activity Budgets And
Diet Of Village And Forest Dwelling Central Himalayan Langurs (Semnopithecus
schistaceus) In The Kedarnath Wildlife Sanctuary, Uttrakhand India. [External advisor]
31. Mwanahamisi Issa (Ph.D. 2015, Institute of Vertebrate Biology Academy of Sciences of the
Czech Republic) Diversity and host specificity of malaria agents in African primates [external
examiner, successfully completed November 2015]
32. Lena Pfluger (PhD 2016, Department of Anthropology, University of Vienna, Austria)[external examiner, successfully completed December 16, 2016]33. Sofia Kaliope Bernstein (PhD 2016, Kyoto University, Primate Research Institute) The
Vocal Repertoire of Tibetan Macaques: A Quantitave Analysis, Congeneric Comparisons, andSpecies-Specific Calls. [Principle academic advisor, co-advisor Koda]
34. Wei Yi Amanda Tan (PhD 2016, Nanyang Technological University, School of Humanities
and Social Sciences, Psychology) Behavioral processes and social influences on thedevelopment of stone-tool use in long-tailed macaques. [External examiner, successfully
November 2016]
35. Nelson Broche Jr. (2019 MSc), Kyoto University, Primate Research Institute) Developmentof a non-invasively determined measurement of stress and ‘pleasure’ in Japanese macaques
using hormones excreted in saliva (Principal advisor)
36. Josué Samuel Alejandro Pastrana (PhD candidate, Kyoto University, Primate ResearchInstitute) Behavioral and hormonal assessment of the effect of different environments on the
stress levels male Japanese macaques. (Principal advisor)
37. Milena Canzler (Masters in Environmental Biology, Animal Behavior, University of Utrecht,
The Netherlands) On the potential use of stimulant drugs by wild animals. [Supervisor ofthesis writing in partial requirement for the degree]
38. K. Santhosh (2017, PhD, Bharathiar University, Coimbatore, India) Study of population,

Huffman, Michael A.

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6ecology and conservation of lion-tailed macaque Macaca silenus in Sirsi-Honnavara, western
Ghats, Karnataka. [external examiner, successfully completed January 9, 2017]
39. Renata Mendonca (PhD 2017, Kyoto University, Primate Research Insitute) Mother-infant

behaviour in wild orangutans (Pongo pygmaeus morio), Danum Valley, Sabah,
Borneo. Secondary advisor w/ Tetsuro Matsuzawa, Misato Hayashi.

40. Victoria Pallante (2017, PhD, Universita Degli Studi Firenze, Italy) Strategies to maintaingroup cohesion: embracing, agonistic support and post-conflict affiliation in two tolerant
species (Theropithecus gelada AND Macaca Tonkeana) (External examiner)
41. Y. Kurihara (2017, PhD Kyoto University, Primate Research Institute). Feeding competitionin Japanese macaques in Yakushima: effects of intergroup hostility and group size. [PhDevaluation committee member]
42. Benjamin Siart (PhD, 2017, Department of Anthropology, University of Vienna, Austria),
Social status, prosocial behavior and measurements of endocrine and protein stress markers inhumans. [external examiner, 2017]43. Jade Burgunder (PhD 2018, Department of Botany and Zoology, Faculty of Science,
Masaryk University, Czech Republic) Complexity in behavioural organization: a novel
approach to assessing parasite infection. [external examiner, October 2018].
44. Rafaela Sayuri Cicalise Takeshita (PhD 2018), Kyoto University, Primate ResearchInstitute) Factors regulating steroid hormones in Japanese macaques and orangutans(Principal advisor w/ Fred B Bercovitch)
45. Cecile Sarabian (PhD, 2019), Kyoto University, Primate Research Institute) On the origins
of disgust: Hygiene, infection-risk avoidance and health maintenance in Papionini and
Hominini [PhD evaluation committee member]
46. Liesbeth Frias (PhD, 2018), Kyoto University, Primate Research Institute) Host –parasite
community interactions in a human-modified habitat [PhD evaluation committee member]
47. Bidyut Sarania (PhD 2019) Tezpur University, Assam, India. Population Distribution,
Behavioural Ecology and Morphometry of Macaca munzala, Arunachal Pradesh, India. [PhDevaluation committee member]
48. Himani Nautiyal (PhD 2020), Kyoto University, Primate Research Institute, Behavioral
ecology of the Central Himalayan Langur (Semnopithecus schistaceus) in a human dominatedlandscape: Multi-species interactions and conservation implications [primary advisor,
committee chair].
49. Raquel Costa (PhD 2020), Primate Research Institute, Kyoto University) ‘How young
gorillas see the world: assessing the impact of ecotourism in Bwindi Impenetrable National
Park, through the eyes of the youngsters’ [Co-academic advisor, with/ M Tomonaga, M
Hayashi].
50. Rosemary Blersch (PhD 2020~202?), University Lethbridge, Canada, Parasitological studyof primates in South Africa. [PhD committee member]
51. G. Vanaraj (PhD 2021), Bharathiar University, Coimbatore, Tamil Nadu, India, Population
Status, Distribution, Activity and Dietary Budgets of Tufted Grey Langur (Semnopithecus
priam priam) at the Kalakkad-Mundanthurai Tiger Reserve, Southern Western Ghats, Tamil
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52. Roy Hammer (Masters, 2022), Utrecht University. Medical School in the Animal
Kingdom? The role of innate propensities and social learning in self-medicative behaviour inanimals. Thesis in partial fulfillment of the Master’s Degree. [thesis co-supervisor with Anne
Marijike Schel]
53. Anna Zanoli (PhD 2022), University of Turin, Unimodal and multimodal communication ofemotional states in human and non-human primates. [external reviewer; academic advisor
Prof. Ivan Norscia]
54. Elodie Freymann (PhD candidate, Institute of Cognitive and Evolutionary Anthropology
Keble College, Oxford University), Intergenerational Transmission of Self-Medicative
Knowledge in Chimpanzees? [co-advisor with Susana Carvalho]
Hosted postdoctoral research fellows, visiting scholars & undergraduate internships
1. Jean-Babtiste Leca (2003-2005, 18 months) Lavoisier Grant, “Ethnographic approach tostone handling, a behavioral tradition in 10 groups of Japanese macaques, Macaca fuscata:Environmental, demographic, and social factors.”
2. Klara Petrzelkova (2004.6.1 – 8.29) JSPS short term-visiting researcher. “Feeding behavior
and possible medicinal plant use in chimpanzee populations on Rubondo Island National Park,
Tanzania.”
3. Cecile M. Garcia (2006.8.16 - 2007.12.20, 18 months) Lavoisier Grant, "Social stress,
nutritional state and reproduction in female Japanese macaques (Macaca fuscata)
4. Alexander Hernandez (2006.8.29 – 2008.9.27, 24 months), JSPS Post-Doctoral Fellowship,“Structure and energetics of the Yakushima Island food web sustaining Japanese macaques(Macaca fuscata) and their helminth parasites.”
5. Jean-Babtiste Leca (2007.11.28 – 2009.11.27, 24 months) JSPS Post-Doctoral Fellowship,“Japanese macaque behavioral traditions: demographic and social factors influencing diffusion
and maintenance”
6. Broox Boze (2008.6.24 – 8.20) NSF-JSPS Summer Internship Program “Behavioral
responses of a coprophagous beetle to a nematode parasite infecting Japanese macaques on
Yakushima Island, Japan”.
7. Cecile Garcia (2008.8.29 - 2008.11.28, 2 months) JSPS Post-Doctoral Short-term
Fellowship, "Social stress, nutritional state and reproduction in female Japanese macaques(Macaca fuscata)."
Professional Consulting Activities
Scientific Advisor to:
BBC Natural History Wildlife Unit, Bristol U.K. (1992- )National Geographic Society, Washington D.C. (1993- )ANC Productions, Tokyo, Japan (1989- )NHK, Tokyo, Osaka, Japan (1983- )University of the Air, Tokyo, Japan (1998- )Natural History New Zealand Ltd. (2001- )Discovery Channel, CANADA (2001- )K Production, FRANCE (2013) for ARTE International
Research Activities(see publications for details)

Huffman, Michael A.

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81977 (1 month) Observation of free-ranging vervet monkeys on St. Kitts Isl., Northern Antilles,
Caribbean (International Expedition Training Institute).
1979-80, 1983- to present, Ecological and sociological studies of the Japanese monkey, Japan(Arashiyama, Takasakiyama, Funekoshiyama, PRI captive colonies)
1983 (1 month) behavioral study of feral cattle on Kuchinoshima Isl., in the Tokara Archipelago,Japan
1985 (6 months) Sociological study of the M Group chimpanzees in the Mahale Mountains
National Park, Tanzania
1987 (6 months) Socio-ecological study of the M Group chimpanzees in the Mahale Mountains
National Park, Tanzania
1989 (6 months) Ecological and parasitological study of the M Group chimpanzees at Mahale
Mountains National Park, Tanzania
1991-92 (4 months) Ethological, phytochemical, and parasitological study of the M Groupchimpanzees in the Mahale Mountains National Park, Tanzania
1991- current, Principle organizer of The C.H.I.M.P.P. Group (Chemo-ethology of theInteractions of Hominoids with Medicinal Plants and Parasites).
1993-94 (4 months) Ethological, phytochemical and parasitological study of the M Groupchimpanzees in the Mahale Mountains National Park, Tanzania
1995-96 (4 months) Ethological, phytochemical and parasitological study of the M Groupchimpanzees in the Mahale Mountains National Park and Gombe Stream National Park,
Tanzania
1996 (1 month) Ethological, microbiological and parasitological study of the M Groupchimpanzees in the Mahale Mountains National Park and Gombe Stream National Park,
Tanzania
1997 (1 month) Ethological, botanical and parasitological study of the Bossou Group of
chimpanzees in Guinea West Africa.
1998 (21 days) Botanical survey and plant collection in Tanzania and pharmacological study of
medicinal plant activity on swine parasites at the Danish Center for Experimental Parasitology,
Copenhagen Denmark.
1998 (1 month) Ethological, botanical and parasitological study of the Sonso Group, Budongo
Forest, Western Uganda; collaborative exchange visit for pharmacological study of medicinal
plant activity at the laboratories of Homeopathy & Pharmacognosy, Parasitology, Universite dela Mediterranee Aix-Marseille, Marseille France; University of Leiden, Leiden, The
Netherlands; Danish Center for Experimental Parasitology, Copenhagen Denmark.
1999 (1 month) Phytochemical study of the Mahale M group chimpanzees in the Mahale
Mountains National Park, Tanzania (October)2000 (1 month) Ethological, phytochemical and parasitological study of the M Groupchimpanzees in the Mahale Mountains National Park, Tanzania. Preliminary site survey of
Rubondo Island National Park, TZ (September)Huffman, Michael A.

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2001- 2009: Principle investigator of Rubondo Island Chimpanzee Development Initiative:
Integrating Research and Conservation. Long-term research project for work on Rubondo Island
National Park
2002 (January - February) Site survey of Rubondo Island National Park, TZ:
SIV cpz sample, general survey of the M Group chimpanzees in the Mahale Mountains National
Park and Rubondo Island National Park, TZ
2002 (June) Ethological study, SIVcpz sampling of the M Group chimpanzees in the Mahale
Mountains National Park, Tanzania, TZ.
2003 (February) Ecological survey of chimpanzee habitat on Rubondo Island National Park,TZ; 3 day visit to Mahale.
2003 (September) Behavioral study on the acquisition and transmission of leaf swallowing
behavior in three captive groups of chimpanzees at Parco Natura Viva-Safari Park,
Bussolengo-Verona Italia.
2003 (December) Ethological, phytochemical and parasitological study of the M Groupchimpanzees in the Mahale Mountains National Park, Tanzania.
2004 (October) Primate observations in the wet and dry zone regions of central (Kurunegala,Kandy area) and northern (Anuradhbura) parts of Sri Lanka.
2005 (January) Short survey of Rubondo Island National Park and Mahale Mountains National
Park chimpanzees, TZ.
2005 (March) Site visit and discussions with collaborators in primate foraging research program.
National University of Taiwan, Institute of Ecology and Evolutionary Biology, Taipei & Fushan
Research Field Station and Botanical Gardens, Ilan County
2006 (January) Primate observations in the wet and dry zone regions of southwestern(Sinharaja), central (Kandy area) and northern (Polonnaruwa) parts of Sri Lanka.
2006 (April-May) Visit, lectures and discussions with collaborators in primate research. Instituteof Wildlife Conservation, National Pingtung University of Science and Technology (Pingtung),
Kaoushiung Medical University (Kaohsiung), Kenting National Park, southern Taiwan
2007 (February) Survey of primates in the southern and central regions of Sri Lanka. Visit toBandipur National Park and Mysore region, southern India to observe bonnet macaques andhanuman langurs.
2007 (March) Visit, lectures and feeding experiments on captive orangutans with collaboratorsin primate research at the Institute of Wildlife Conservation, National Pingtung University of
Science and Technology (Pingtung), Kaoushiung Medical University (Kaohsiung) Taiwan.
2007 (September-October) Survey of primates in Karnataka (Bandipur NP), Kerera (Sharavathi
Wildlife Sanctuary) and Tamil Nadhu (Indira Ghandi Wildlife Sanctuary, Western Ghats,
Anaimalai hills, Valparai) states in southern India to observe bonnet macaques, lion-tailed
macaques, nilgiri langurs, slow loris and hanuman langurs. Based out of National Institute forAdvanced Studies, Bangalore.
2007 (November 4-12) Five-part lecture series: Fundamentals of Primatology, at the
Behavioral Research Institute in the University of Guadalajara-Mexico.
2008 (May 7-12) "The animal roots of herbal medicine- People learning from animals aboutHuffman, Michael A.

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10

ways to cure disease. Another reason why we need to conserve plant and animal bio-diversity"
Lectures at the Institute of Wildlife Conservation, National Pingtung University of Science andTechnology (Pingtung), Kaoushiung Medical University (Kaohsiung) Taiwan. Collaboration on
civet leaf swallowing and parasite expulsion.
2008 (August 14-28) Indonesia, Bali, Ubud temple long-tail macaque stone handling study.
2009 (February 23 – March 24) Sri Lankan Primate Surveys; India New Delhi, brief
observations of urban rhesus monkey group.
2009 (October) Sri Lankan Primate Surveys in Western Province.
2010 (March 8 - 24) Sri Lankan Primate Survey in Western Province, examination of extent andfossil primates, participated in cultural anthropological field trip to interview EasternVedda inBatticaloa Provence, lectures at Sri Jayawardenepura University and Post-graduate Institute ofArchaeology, visit to Potana and Anuradhapura archaeological sites.
2010 (March 24-27) Singapore; Seminar at Nanyang University, observation of longtailed
macaques.
2010 (August 17-29) Vietnam, Khan Vinh Province, Khanh Phu Commune, Primate survey andfecal sample collection for the study of monkey malaria (Plasmodium knowlesi) transmission in
the Raglai indigenous human population in a mountainous area of south central Vietnam.
2010 (December 7-23) Sri Lanka Primate Surveys in Central Provence.
2011 (January 10–23) Observation of wild chacma baboons in the Western Cape of South
Africa.
2011 (February 20-March 7) Vietnam, Khan Vinh Province, Khanh Phu Commune, Primate
survey and fecal sample collection for the study of monkey malaria (Plasodium knowlesi)transmission in the Raglai indigenous human population in a mountainous area of south central
Vietnam.
2011 (November) Sri Lanka Primate Surveys in Western, Central and Subaragamuwa Provinces.
2012 (February 25 – March 9) Vietnam, Cuc Phuong National Park & Khan Vinh Province,
Khanh Phu Commune, Primate survey and fecal sample collection for the study of monkey
malaria (Plasodium knowlesi) transmission in the Raglai indigenous human population in a
mountainous area of south central Vietnam.
2012 (June 3 – 22 / 23 -30) Sri Lanka primate surveys / and a week at National Institute ofAdvanced Studies in Bangalore, India
2012 (November 6 – 22) Sri Lanka primate survey and sample collection.
2013 (January 12- 27) Sri Lanka primate survey and sample collection.
2013 (March 3-13) Khan Vinh Province, Khanh Phu Commune, Primate survey and fecal
sample collection for the study of monkey malaria (Plasodium knowlesi) transmission in the
Raglai indigenous human population in a mountainous area of south central Vietnam. Intestinal
parasite survey of Monkey Island macaques.
2013 (June 16-July 1) Tanzania, Mahale Mountains National Park, behavioral ecology study andfilming of program ‘Animal Doctors’, K productions for ARTE TV.

Huffman, Michael A.

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112013 (August 17- / ) Sri Lanka primate survey and sample collection.
2013 (October 13-18 India/ 19-25 Nepal/ 26-31 Vietnam) Visit to NIAS Bangalore/ Primate
survey and sample collection Nepal/ Khan Vinh Province, Khanh Phu Commune, Primate
survey and fecal sample collection for the study of monkey malaria (Plasmodium knowlesi)transmission in the Raglai indigenous human population in a mountainous area of south central
Vietnam.
2014 (June 12-26 Sri Lanka/ 27- July 4 China) Sri Lanka primate survey and sample collection(Western Province-Wilpattu NP, Northern Province to Jaffna).
2015 (January 20- February 8 Sri Lanka) Sri Lanka primate survey, sample collection and DNA
extraction (Mihintale NCP, South, Eastern SL).
2015 Taiwan
2015 (June 4-29) Field site visit to see Central Himalayan langurs in Mandal village and Chopta,
Uttarakhand, India.
2015 (July 31-August 17) Field trip. ITP2015 training program in Sri Lanka (Participantssponsored by PWS), Field trip co-organizer, resource person.
2015 (October 24 – November 3) Khan Vinh Province, Khanh Phu Commune, Monkey Island,
Pet monkey survey and fecal/blood sample collection for the study of monkey malaria(Plasodium knowlesi).
2015 (December 20 – 2016 January 11) Sri Lanka primate survey, behavioral observation,
sample collection and DNA extraction.
2016 (May) Lectures at Taipei National University, Kaohsiung Medical University, National
Pingtung University of Science and Technology.
2016 (June) Paper presentation at Oxford University: International Symposium on Primate
Archeology, visit to Trinity University, Dublin Ireland.
2016 (July 23~30) “Animal self-medication: ethnoveterinary medicine without human cultural
bias?” Joint Natural Products Conference 2016 Copenhagen, Copenhagen, Denmark,2016 (August 7~24) Field site visit to see Central Himalayan langurs in high altitude alpine
meadows of Rudranath, Uttarakhand, India.
2016 (October 14~27) Co-organizer of 5th Asian Primate Symposium. Site visits to Pollonaruwa,
Nachimale temple, Dombagaskanda Forest Reserve, Sri Lanka (transit visit to Bangkok).
2017.3.8~27 Primate survey of montane and arid areas (Knuckles, Nuwara Eliya, Mannar
Island).
2018.2.18~3.9 Primate survey of montane, dry zone, and wet areas (Knuckles, Mihintale,
Kotiyakumbura).
2018.08.28~.09.16 Primate survey of dry zone langurs and human-primate conflict of theWestern Province.
2018.11.20~.11.29. International Training Program in Terrestrial and Marine Ecology, Field trip,
Co-organizer, resource person.

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2018.12.25-26. Stone handling behavior in free-ranging Taiwanese macaques of Mt. Shoushan(Kaohsiung) and in the Animal Rescue Centre (Pingtung National University of Science andTechnology).
2019.3.9-24. Primate survey of dry zone grey langurs in Mannar, Jaffna, Mulaitivu, Trincomaleeand Mihintale.
2019.5.23, 25-26. Site visit and discussion for field study on self-medicative behavior in wild
horses and free-ranging Maremmena cattle and pigs. San Rossore State Park and Tunata
Paganico organic farm. Tuscany, Italy.
2019.6.18~14. Site visits and discussion for field study of monkey malaria (P. knowelsi) in
Sabah State Malaysia (Borneo).
2019.12.11~20. Site visit and review of data being collected on stone-handling in Taiwanesemacaques at the Wildlife Rescue Center, National Pingtung University of Science andTechnology, Nepu (Pingtung), southern Taiwan.
2020.3.5-19 Sri Lanka primate survey. Knuckles Range.
2021.3~ COVID-19
Public Education, Popular Science Activities
A few textbooks, technical reference citations
1. The Plant-Book. A portable dictionary of the vascular plants. 2nd Edition. (1997). D. J.
Mabberley, Cambridge University Press, Cambridge. Vernonia amygdalina description
includes use of chimpanzees when sick and bioactivities properties of pith. (p. 744).
2.

Parasitology & Vector Biology 2nd Edition (2000). William C. Marquardt, Richard S.
Demaree, Robert B. Grieve, Harcourt Academic Press, San Diego. “Self-treatment forparasites” (text box p. 390).

3.

The Natural History of Medicinal Plants (2000). Judith Sumner, Timber Press, Portland.
“Chimpanzees and self-medication” (pp. 149-150).

4.

Natures Medicine Plants that Heal (2000) Joel L. Sherdlow, National Geographic Society,
Washington, D.C. “Vernonia amygdalina, includes description of medicinal uses forhumans and chimpanzees. (p. 388). On animal self-medication (pp. 176-177, 180).

5.

Parasite Rex. Inside the world of nature’s most dangerous creatures (2000). Carl Zimmer,
The Free Press, New York (pp. 201-202).

6.

Medicine Quest. In search of Nature’s Healing Secrets (2000). Mark J. Plotkin, Viking
Penguin Press, New York, “Plants of the Apes” (pp. 163-165, 167-169).

7.

Wild Health. How animals keep themselves well and what we can learn from them (2002)
Cindy Engel, Houghton Mifflin Company, Boston, “chimpanzees” (pp. 46, 200), parasites(pp. 139, 140, 142-143, 146), “self-medication, determination of” (p. 37). 「動物たちの
自然健康法— 野生の知恵に学ぶ」 （２００３） シンディ エンジェル（羽田節子訳）紀伊
國屋書店

8.

Primate Behavioral Ecology 1st ~ 4th Editions (2000, 2003, 2007, 2011) Pearson PrenticeHall; 5th Edition (2017), Routledge. Karen Strier, Allyn and Bacon, Boston.; “Box 6.1
Forest Pharmacy”.

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Parasites and the Behavior of Animals (2002). Janice Moore, Oxford Series in Ecology
and Evolution, Oxford University Press, Oxford. “Self-medication” (pp. 135-137).

10. Modern Biology (2005~ Editions) Holt, Rinehart and Winston, Austin. Science in Action
Feature “Do animals self-medicate? (p. 598).
Some recent selected on-line interviews & articles“Monkey Culture” The Washington Post, December 10, 1997, Lucille Craft.
https://www.washingtonpost.com/archive/1997/12/10/monkey-culture/7d0918a9-22da-4e06-92
b7-e9546789fe69/“How man apes animal medicine” The Telegraph, Aisling Irwin. Dec. 14, 2000
http://www.telegraph.co.uk/news/science/science-news/4757357/How-man-apes-animal-medici
ne.html“Apes were first to get high on drugs” The Telegraph 26 December 2001. Roger Highfield
http://www.telegraph.co.uk/news/worldnews/1366329/Apes-were-first-to-get-high-on-drugs.ht
ml“Hot tub monkeys” offer eye on nonhuman “culture” National Geographic News, February 6,
2004. Bijal P. Trivedi
http://news.nationalgeographic.com/news/2004/02/0206_040206_tvmacaques.html
Nature's Pharmacy: Ancient Knowledge, Modern Medicine: Really Wild Remedies. Universityof Iowa Hospital & Clinics, Medical Museum.
http://www.uihealthcare.com/depts/medmuseum/galleryexhibits/naturespharmacy/remedies/wild.html“Self-medicating animals” The Scientist, October 23, 2012
http://www.the-scientist.com/?articles.view/articleNo/32957/title/Self-Medicating-Animals/“Natural-born doctors” The Scientist, October 23, 2012
http://www.the-scientist.com/?articles.view/articleNo/32966/title/Natural-Born-Doctors/
Herbal Medicina came from animals. The Hindu, June 27, 2013.
http://www.thehindu.com/sci-tech/health/medicine-and-research/herbal-medicine-came-from-an
imals/article4853660.ece?utm_source=MostCommented&utm_medium=Homepage&utm_camp
aign=WidgetPromo“Wild Medicine- Some animals have learned how to use plants to heal themselves” January 19,
2014. https://student.societyforscience.org/article/wild-medicine“Study notes widening disease transmission by monkeys” Sci Dev Net August 20, 2014.
http://www.scidev.net/asia-pacific/malaria/news/study-notes-widening-disease-transmission-bymonkeys.hAnimaux médecins : La science inspirée par Curious George
https://www.actualitte.com/insolite/animaux-medecins-la-science-inspiree-par-curious-george-5
2415.html“Animals Hack Nature’s Medicine Cabinet – We Should Too” Discover, Science for the Curious.
December 22, 2014

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http://blogs.discovermagazine.com/crux/2014/12/22/animals-hack-natures-medicine-cabinet/#.
VJhLo0CwIA“News Features: Animals that self-medicate.” Joel Shurkin December 9, 2014 PNAS 111(49):17339-17341. http://www.pnas.org/content/111/49/17339.full

How animals SELF-MEDICATE: Dogs, elephants, chimps and even parrots use natural
remedies to treat digestive problems and even induce birth. Mail Online December 10,2014http://www.dailymail.co.uk/sciencetech/article-2868348/Do-animals-SELF-MEDICATE-Dogselephants-chimps-parrots-use-natural-remedies-treat-digestive-problems-induce-birth.html
Wild medicine "On an expedition early in his career, Michael Huffman was watching a groupof chimpanzees in an East African jungle when the primate researcher made" January 17, 2014 |US Science News (United States)

Automedicación animal
http://www.acercaciencia.com/2015/01/23/automedicacion-animal/
Is self-medication the future of pharmacy?" March 9, 2015 | Deccan Herald (India)
http://www.deccanherald.com/content/464489/is-self-medication-future-pharmacy.html
用天然材料做药？动物也会 2014.12.16, Chinese introduction of Joel Shurkin article above.
http://www.guokr.com/article/439671/
Dr. Zoo: Radio interview on the field of animal self-medication (German).

The Self-Medicating Animal: What can we learn from chimps and sheep and maybe
even insects that practice medicine on themselves? The New York Times On-line. A
version of this article also appears in print on May 21, 2017, on Page MM68 of the
Sunday Magazine with the headline: Doctor Who?
By MOISES VELASQUEZ-MANOFFMAY 18, 2017https://www.nytimes.com/2017/05/18/magazine/the-self-medicating-animal.html?rref=collectio
n%2Fsectioncollection%2Fscience&action=click&contentCollection=science&region=stream&
module=stream_unit&version=latest&contentPlacement=3&pgtype=sectionfront
Press conference relaease. 野生ニホンザル 温泉でストレス解消か 京大グループ
https://www3.nhk.or.jp/news/html/20180404/k10011390021000.html 日本語
https://www3.nhk.or.jp/nhkworld/en/news/20180404_06/

EnglishHot Springs Lower Stress in Japan’s Popular Bathing Monkeys
James Gorman New York Times:
https://nyti.ms/2Gvp3JB

Japanese macaques relieve stress in the hot springs. Ein heißes Affenbad zum
Stressabbau (Ärzte Zeitung online, 04.04.2018)https://www.aerztezeitung.de/panorama/article/961013/japan-heisses-affenbad-stressabbau.html

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The New Evolutionary Enlightenment: Animal self-medication (Written interview toquestions on animal self-medication; in Spanish and English; uploaded March 2021)
https://ilevolucionista.blogspot.com/2021/02/automedicacion-animal-entrevista.html

Podcasts and Webinar

The PrimateCast #42: An Interview with Dr. Michael Huffman
April 29th 2016
http://www.cicasp.pri.kyoto-u.ac.jp/news/podcasts/primatecast-42-interview-dr-michael-huffman

TEDxOsaka Animal Self-Medication
https://www.youtube.com/watch?v=WNn7b5VHowM“The evolution of animal self-medication and lessons for the development of medicine
and new medicines”
Plenary Lecture featured at the 64th International Congress and Annual Meeting of the Society
for Medicinal Plant and Natural Product Research (GA). Innsbruck, Austria. Sept. 1-5, 2019https://www.ga-online.org/webinars-workshops
Association of Indian Primatology, livestream Webinar. 2020.08.13
The evolution of self-medication in primates and the origin of human medicinal plant use.

YouTube https://www.youtube.com/watch?v=li5ZsOBpPP0
Facebook https://www.facebook.com/IndianPrimates/videos/632989237336466/
BBC New World Service Crowd Science
Do animals use medicine?
March 26, 2021

CICASP International Primatology Lecture Series, The past, present and the future
June 16, 2021 Lecture #1 Huffman, M.A. Huffman ' Learning to become a monkey andother lessons for becoming a primatologist'
YouTube https://youtu.be/0euvFq7gVkM

Selected public science television programs, radio interviews and consultation

1. 新ニホンザル学−２１世紀への科学 １９８４. "New perspectives in Japanese macaquestudies - Science in the 21st Century" (1984) NHK. Featuring with. The history of Japanese
primatology is presented by one of the pioneers in the field, Prof. Junichiro Itani, and features
some recent findings by two of his students Yukio Takahata and Michael A Huffman (Kyoto
University primatologists) in this NHK special program (in Japanese). This video was
uploaded strictly for non-profit, educational, or personal use only.
https://www.youtube.com/watch?v=VvEosziekbY
2. ウォッチング「熱中 石で遊ぶニホンザル」1985 年 6 月 26 日：NHK 総合テレビ司会者 タモリ；
ゲスト：河合雅雄（京都大学霊長類研究所教授）；マイケル・ハフマン（京都大学動物学教室）
Watching “Feverish stone handling Japanese monkeys. ”NHK Sougo TV, 1985 June 26. Host Tamori,guests, Prof. Kawai, Masao (Primate Research Institute), Michael A Huffman graduate student (Kyoto
University, Department of Zoology)

3. Chimpanzees use Medicine: ‘New Station’ Asahi TV evening program, aired in Feb. 1992.
A popular evening news program of Asahi TV aired between 1985 to 2004. The programHuffman, Michael A.

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presented on national TV for the first time, news about recent the research findings
demonstrating the anti-parasitic properties of medicinal plants by chimpanzees in the Mahale
Mountains of Tanzania. Featuring researchers Michael A Huffman and Koichi Koshimizu of
Kyoto University, in Japanese. This video was uploaded strictly for non-profit, educational, or
personal use only. https://www.youtube.com/watch?v=tRNdhZLfUb0
4. University of the Air (放送大学): A multi-part special series called “Human - In Search of
the Roots of Humankind". This was originally produced by the University of the Air inJapanese as video course material. Later the scripts were translated into English and narratedby Michael A Huffman (Kyoto University) in collaboration with Professor Itani to share theinformation with an international audience. The species presented in this series represent
some of the many important field sites featuring a diverse range of primate species around theworld pioneered by Japanese primatologists of Kyoto University. Prof Itani, featured here
toward the end of his career introduces each site and sets the stage for the on-site presenter, aprimatologist responsible for the long-term work conducted at each site by his students over
the years. These videos have been uploaded for educational purposes only with the intent topublicly archive its valuable information.: The Society of Japanese macaques, Social Groupsof Prosimians, Gorilla Society, The Society of Chimpanzees.
Available on Primate I/O:
https://www.youtube.com/channel/UCRb9cKNQvqlWtVcPy5qghlQ
The New World Monkeys of Macarena, Colombia
https://www.youtube.com/watch?v=m3Lm95BPOwI
5. “Wisdom of the Wild” (2000), Argo Productions Washington D.C. for Public Broadcasting
System (USA). A national and later internationally televised documentary with special featureof chimpanzee self-medication research project at Mahale. (Appearance, interview, scientificadvisor)6. “Magic, murder, and medicine”, BBC Radio 4 (2001), Interview for 2 part series on
medicinal plants. (Interview, scientific advisor)
7. “Nature’s Secrets” (2002), Discovery Channel, Toronto. An internationally televised
documentary with special feature of chimpanzee self-medication research at Mahale.
(Appearance, interview, scientific advisor)
8. “Dr. Animal” (2003), Tele Image Paris for Discovery Channel. An internationally televised
documentary, with special feature of chimpanzee self-medication research at Mahale.
(Appearance, interview, scientific advisor)
9. “Animal self-medication”, BBC Radio 4 (2003), A special program on animal
self-medication. (Interview, scientific advisor)
10. “Animal Medicine” (2004), St. Thomas, Prod., Aix en Provence, France for National
Geographic Television Documentary on self-medicative behavior in the animal kingdom.
(Appearance, interview, scientific advisor)
11. Animals Like Us- Culture (2009), National Geographic TV (St. Thomas, Prod., Aix en
Provence, France). An internationally televised documentary with special feature ofchimpanzee self-medication research at Mahale and Stone handling, a behavioral tradition in
Japanese macaques. (Appearance, interview, scientific advisor)
12. Animals Like Us- Medicine (2009), National Geographic TV (St. Thomas, Prod., Aix en
Provence, France). An internationally televised documentary on self-medication in animals
with special feature of chimpanzee self-medication research at Mahale. (Appearance,interview, scientific advisor)Huffman, Michael A.

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13. Animal Doctors (2014), ARTE television (Produced and Directed by R Productions,
Jacques Mitsch), An internationally televised documentary on self-medication in animals withspecial reference to the history of self-medicative research by MA Huffman on chimpanzee
self-medication research at Mahale. (French, German, English)
14. Life Story: Episode ‘Courtship’ (2014) BBC Wildlife History Unit (Narrated by David
Attenborough), Scientific Advisor for Japanese macaque part of Episode ‘Courtship’, on-site
at Arashiyama Japan.
15. Animals at Play (2020), BBC 2 part series on play in animals. Scientific, interviewee forsequence on stone handling in Japanese macaques of Arashiyama.
16. 新説! 所ジャパン #063 (2020) Shinsetsu! Tokoro Japan, Hot spring monkeys of Shiga
Heights, Nagano. March 2, 2020 Fuji TV program (Appearance).
Museum exhibits
1. United States Botanic Garden, Washington D.C., Film and text of research included in‘Navipass Multimedia Tour handheld devise for self-guided educational narration of the
botanical gardens. Used to describe use of chimpanzee medicinal plants in the gardens.
May 2005 (Scientific advisor)
2. Ibaraki Prefectural Nature Museum, “Medicinal Plants: A Boxful of Gifts from Mother
Nature”, 33rd Exhibition, Special section on animal self-medication, with a focus on
chimpanzees. On displayed from March 19-June 19, 2005. (scientific advisor, provided
exhibit materials). ミュージアムパーク茨城県自然博物館「第３３回企画展 YAKUSO 野山
は自然のくるりばこ」、２００５年３月１９日〜６月１９日開催
3. The Kyoto University Museum, Kyoto, Special Exhibition “The Natural History ofMedicine” On display from June 1 to September 1, 2003. (scientific advisor in charge of‘Animals and Medicine’ section). 京都大学総合博物館、平成１４年特別企画展「薬と自然詩」、２００３年６月１日〜９月１日開催