ᒣ⏣ࢩ࣏ࣥࢪ࣒࢘᥼ຓ୍ぴ⾲⤌⧊ጤဨ㛗໭㝣ඛ➃⛉Ꮫᢏ⾡኱Ꮫ㝔኱Ꮫ⏣୰ ᏹ࿴㞟఍
➨ 5 ᅇᒣ⏣ࢩ࣏ࣥࢪ࣒࢘ࠕ⮬↛⾜ື୰ࡢ⬻ᶵ⬟࢖࣓࣮ࢪࣥࢢࠖ
Neuroimaging of Natural Behaviors఍ᮇ㛤ദᆅ

201710/2~10/5ᮾி㒔┠㯮༊㸯௳ ᥼ຓ㔠 8,000,000 ෇ᡂᯝሗ࿌
➨ 5 ᅇᒣ⏣ࢩ࣏ࣥࢪ࣒࢘ࠕ⮬↛⾜ື୰ࡢ⬻ᶵ⬟࢖࣓࣮ࢪࣥࢢࠖ
Neuroimaging of Natural Behaviors
2017 ᖺ 10 ᭶ 2 ᪥㹼5 ᪥ ᮾிᕤᴗ኱Ꮫ ⶶ๓఍㤋໭㝣ඛ➃⛉Ꮫᢏ⾡኱Ꮫ㝔኱Ꮫ ⏣୰ᏹ࿴ ➨ 5 ᅇ ᒣ ⏣ ࢩ ࣥ ࣏ ࢪ ࢘ ࣒ ࠕ ⮬ ↛ ⾜ ື ୰ ࡢ ⬻ ᶵ ⬟ ࢖ ࣓ ࣮ ࢪ ࣥ ࢢ ࠖ (Neuroimaging of NaturalBehaviors) ࡀࠊ2017 ᖺ 10 ᭶ 2 ᪥࠿ࡽ 5 ᪥ࡲ࡛ࡢ 4 ᪥㛫ࠊᮾிᕤᴗ኱Ꮫ኱ᒸᒣ࢟ࣕࣥࣃࢫⶶ๓఍㤋࡟࡚㛤ദࡉࢀࡲࡋࡓࠋᚑ᮶ࡢ⬻ᶵ⬟࢖࣓࣮ࢪࣥࢢศ㔝࡟࠾࠸࡚ࠊᐇ㦂ᐊ᮲௳ୗࡢ⤫ไᐇ㦂࡛⬻ᶵ⬟࡟㛵ࡍࡿከࡃࡢ▱ぢࡀᚓࡽࢀࡓ୍᪉ࠊ㌟యࡣᣊ᮰ࡉࢀឤぬධຊࡣ༢⣧࡞ࡶࡢ࡟㝈ᐃࡉࢀ࠾ࡾࠊᡃࠎࡀ᪥ᖖ⾜࠺ືసࡸぢ⪺ࡁࡍࡿឤぬධຊࢆ⬻ࡣ࡝ࡢࡼ࠺࡟ฎ⌮ࡍࡿࡢ࠿࡟㛵ࡋ࡚ࠊᮍࡔከࡃࡢㅦࡀṧࡉࢀ࡚࠸ࡲࡍࠋ㏆ᖺࠊ⬻Ἴ㟁ᴟ࡞࡝ࡢࣁ࣮ࢻ࢙࢘࢔ࠊ⊂❧ᡂศศᯒ࡞࡝ࡢಙྕฎ⌮ࠊ⮬↛ᫎീ࡞࡝ࡢᐇ㦂ࣃࣛࢲ࢖࣒ࠊࡑࡋ࡚⬻ෆィ⟬ฎ⌮ࡢࣔࢹࣝ໬࡜࠸ࡗࡓ㐍ᒎ࡟ࡼࡾࠊ⮬↛࡞᮲௳ୗ࡛ࡢ⬻ᶵ⬟࢖࣓࣮ࢪࣥࢢࡀྍ⬟࡟࡞ࡾࡘࡘ࠶ࡾࡲࡍࠋᮏࢩ࣏ࣥࢪ࣒࡛࢘ࡣࠊ⾜ື୰ࡢ㌟య㐠ື࣭⬻Ἴྠ᫬ィྠ᫬ィࠊᐇ⎔ቃ࡛ࡢࣈࣞ࢖࣭ࣥࢥࣥࣆ࣮ࣗࢱ࣭࢖ࣥࢱ࣮ࣇ࢙࢖ࢫࠊ⮬↛ᫎീど⫈ࠊ」ᩘே

࡟ᇶ࡙ࡃ♫఍⬻⛉Ꮫ࡞࡝࡜࠸ࡗࡓ⮬↛࡞≧ἣୗ࡛ࡢ⬻ᶵ⬟࢖࣓࣮ࢪࣥࢢ࡟㛵ࡋ࡚ࠊୡ⏺

୰ࡢ◊✲⪅ࡀ୍ᇽ࡟఍ࡋ࡚ṇ࡟ୡ⏺ࢺࢵࣉࣞ࣋ࣝࡢㅮ₇࡜㆟ㄽࡀ⧞ࡾᗈࡆࡽࢀࡲࡋࡓࠋᾏእㅮ₇⪅ࡣ࢔࣓࣭ࣜ࢝ࢻ࢖ࢶ࣭࣮࢜ࢫࢺࣛࣜ࢔࣭ࣇࣛࣥࢫ࣭ྎ‴࣭࢖ࣥࢻ࣭ࣟࢩ࢔࠿ࡽィ 12 ྡࠊᅜෆㅮ
₇⪅ࡣ 20 ྡ࡛ࠊྜィ 32 ௳ࡢㅮ₇ࡀ࠶ࡾࡲࡋࡓࠋ4 ᪥㛫ࡢࢩ࣏ࣥࢪ࣒࢘ࡣࠊ௨ୗࡢ 11 ࡢࢭࢵࢩࣙ
ࣥ࡟ศ࠿ࢀ࡚࠾ࡾࡲࡋࡓࠋ⬻Ἴࡢࣁ࣮ࢻ࢙࢘࢔㛤Ⓨࡸಙྕゎᯒ࡜࠸ࡗࡓᇶ♏ᢏ⾡࠿ࡽࠊ⚄⤒⛉Ꮫⓗࣜࣁࣅࣜࢸ࣮ࢩࣙࣥࡸࣈࣞ࢖࣭ࣥࢥࣥࣆ࣮ࣗࢱ࣭࢖ࣥࢱ࣮ࣇ࢙࢖ࢫ࡜࠸ࡗࡓᛂ⏝ศ㔝࡬ࠊࡑࡋ࡚♫఍⚄⤒⛉Ꮫࡸ㡢ᴦ࡜࠸ࡗࡓேᩥศ㔝ࡲ࡛ࢆಠ▔ࡍࡿࠊࡲࡉ࡟Ꮫ㝿ⓗ࡞఍ྜ࡛࠶ࡾࡲࡋࡓࠋ
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Neurorehabilitation
Music and Rhythm
Brain-computer interfaces
Locomotion and balance
Clinical translational neuroscience
Mobile brain/body imaging (I/II)
Social neuroscience
Brain networks
Natural visual experiences
Advanced signal processing

Opening Address
I would like to welcome all of you to the 5th Yamada
Symposium on Neuroimaging of Natural Behaviors. It is my great
pleasure and honor to welcome all and to give an opening addresson this symposium on behalf of the Yamada science foundationỤᑼ⌮஦㛗(YSF).

YSF was established in 1977 by Mr. Kiro Yamada, the former president of the Rohto
Pharmaceutical Company Ltd. on the basis of his personal funds to promote basic science in Japan.
The YSF supporting programs include: i) basic science research projects; ii) international
conferences/symposia and workshops; and iii) overseas exchanges of young scientists. The sciencefields to be covered are physics, chemistry and life science. YSF puts emphasis on original ideas andchallenges to develop new science frontiers.
The Yamada symposium started in 2003 on the occasion of the YSF 25th anniversary. It aims todevelop new science fields through active discussions among active scientists over various fields andgenerations.
The present symposium is just what we would like and be pleased to extend our support to. Thissymposium puts emphasis on brain mechanisms of natural behaviors, innovative fields of neuroimaging, new measurements and analyses of neuroimaging, new opportunities to study brains,synergistic effects for future progresses, and vital discussions among various fields. These aresubjects of frontiers of the fields, interdisciplinary researches and new developments. The subjects
and the ways of the discussions in the present symposium are just what YSF would like to appreciateand encourage.

Hiroyasu Ejiri
President of the Yamada Science Foundation,
Professor Emeritus of Osaka University

Preface
Neuroimaging has revolutionized our understanding of thefunctions and structures of the living human brain especially since
the introduction of functional magnetic resonance imaging in early
1990s, furthering our knowledge beyond the confines of clinical⏣୰⤌⧊ጤဨ㛗observations and animal experiments. Neuroscience has since

expanded its territories from low-level sensory processing and motor control into high-level cognitive
functions specific to humankind such as memory, language, and executive functions. Until recently,
however, human neuroimaging has been restricted to a laboratory setting where stimuli are artificiallysimplistic and body movements are strictly controlled, so our understanding of brain functions innatural environments is limited. Human intelligence and creativity are best understood in naturalenvironments where humans interact with natural objects and communicate with other humans tosolve problems in need.
Recent developments in measurement hardware, signal processing, and experimental designs haveopened new opportunities to overcome the movement restriction of fMRI and to investigate thehuman brain in action. The core concept is Mobile Brain/Body Imaging (MoBI) advocated by Drs.
Scott Makeig and Klaus Gramann. MoBI is a new neuroimaging modality, integrating mobile brain
imaging methods and other data streams such as motion captures and eye tracking to investigate brain
activity while participants actively move in and interact with their environment. The newdevelopments have promise for new insights into human brain functions interacting with physical andvirtual environments.
The 5th Yamada Symposium focuses on innovative fields of neuroimaging dedicated tounderstanding brain mechanisms of natural behaviors. The Symposium also covers novel researchareas that are awaiting for new possibilities for human neuroimaging. I hope the Symposium willprovide a stimulating and synergistic opportunity for forthcoming innovations. Finally, on the behalfof the organizing committee, I would like to express sincere gratitude for the generous support fromthe Yamada Science Foundations.

Hirokazu Tanaka
Organizing Committeeࡲࡎᒣ⏣⛉Ꮫ᣺⯆㈈ᅋ⌮஦㛗ࡢỤᑼᏹὈඛ⏕࠿ࡽ㈈ᅋࡢࡈ⤂௓ࠊࡑࡋ࡚ᒣ⏣ࢩ࣏ࣥࢪ࣒࢘ࡢ୺᪨࡛࠶ࡿ⛉Ꮫࡢ㐍ᒎࡢࡓࡵࡢ⊂๰ᛶ࡜Ꮫ㝿ᛶ࡟㛵ࡋ࡚ࡈㄝ᫂࠸ࡓࡔࡁࠊᮏࢩ࣏ࣥࢪ࣒࢘ࡣᖥࢆ㛤ࡅࡲࡋࡓࠋࡘ࡙࠸࡚ᇶㄪㅮ₇⪅ࡢ୍ே࡛࠶ࡿᕝேග⏨ඛ⏕࠿ࡽࠊ⬻ᶵ⬟࢖࣓࣮ࢪࣥࢢࡢඛ➃ᡭἲ࡛࠶ࡿࢹࢥ࣮ࢹ࢕ࣥࢢἲ࡟ࡼࡿ⬻ෆ᝟ሗࡢゎㄞ࡜ࠊࡑࡢゎㄞࡉࢀࡓ᝟ሗࢆᇶ࡟⬻ࢆカ⦎ࡍࡿヨࡳ࡟㛵ࡋ࡚ࠊ⚄⤒⛉Ꮫⓗࣜࣁࣅࣜࢸ࣮ࢩࣙࣥࡢ᭱ඛ➃ࡢࡈ◊✲ࢆ⤂௓࠸ࡓࡔࡁࡲࡋࡓࠋ᪥ᮏⓎࡢ⊂๰ⓗ࡞◊✲࡟ࠊᅜෆࡢ⫈⾗ࡣࡶࡕࢁࢇࠊᾏእㅮ₇⪅࠿ࡽࡶάⓎ࡞㉁␲ࡀᕳࡁୖࡀࡾࡲࡋࡓࠋᾏእ࠿ࡽࡢᇶㄪㅮ₇⪅࡛࠶ࡿ࢝ࣜࣇ࢛ࣝࢽ࢔኱Ꮫࢧࣥࢹ࢕࢚ࢦᰯࡢ
Scott Makeig ඛ⏕࠿ࡽࡣࠊ⮬↛⾜ື୰ࡢ⬻Ἴィ ࣃࣛࢲ࢖࣒࡛࠶ࡿࣔࣂ࢖ࣝ⬻㌟య࢖࣓࣮ࢪࣥࢢἲ࡟㛵ࡋ࡚ࡈㅮ₇ࢆ㡬ࡁࡲࡋࡓࠋࣁࣥࢫ࣭࣮࣋ࣝ࢞࡟ࡼࡿ⬻ἼࡢⓎぢࡢṔྐ࠿ࡽࠊ
Makeig ඛ⏕ࡈ⮬㌟ࡢᴗ⦼࡛࠶ࡿ⊂❧ᡂศศᯒ࡟ࡼࡿ⬻Ἴゎᯒࡲ࡛ࢆಠ▔ࡍࡿࠊࡲࡉ࡟Ṕྐⓗ࡞ㅮ₇࡛ࡋࡓࠋMakeig ඛ⏕࡟ࡣ࡯࠿࡟ࡶከᩘࡢ㢧ⴭ࡞ᴗ⦼ࢆᣲࡆ࡚࠾ࡽࢀࡲࡍࡀࠊࡉࡽ࡟᪂ศ㔝ࢆ㛤ᣅࡉࢀࡼ࠺࡜ࡋ࡚࠸ࡿጼࡣᅜෆࡢ◊✲⪅࡟ᙉⅯ࡞༳㇟ࢆṧࡋࡓ࡟㐪࠸࠶ࡾࡲࡏࢇࠋᒣ⏣ࢩ࣏ࣥࢪ࣒࢘ࡢࡶ࠺୍ࡘࡢ୺᪨ࡣᏛ㝿ᛶ࡛ࡍࠋࡼࡾ⮬↛࡞≧ἣ࡛ࡢ⬻ᶵ⬟࢖࣓࣮ࢪࣥࢢࡀྍ⬟࡟࡞ࡗࡓࡇ࡜࡟ࡼࡾࠊᚑ᮶ࡣ⬻⛉Ꮫ࡜↓⦕࡛࠶ࡗࡓศ㔝࡜ࡢ㐃ᦠࡀᛴ㏿࡟㐍ࢇ࡛࠸ࡲࡍࠋࢩ࣏ࣥࢪ࣒࡛࢘ࡣࠊࢪࣕࢢࣜࣥࢢࡸࣇࣛ࢖ࢺࢩ࣑࣮ࣗࣞࢱ᧯స୰࡜࠸ࡗࡓ୍ேࡀ⾜࠺⾜ື࠿ࡽࠊ㡢ᴦࡸ♫఍ᛶ࡜࠸ࡗࡓே㛫ᅛ᭷ࡢ」ᩘே༠ㄪ⾜ືࡲ࡛ࠊᩘᖺ๓࡛ࡣ⪃࠼ࡽࢀ࡞࠿ࡗࡓ⬻ᶵ⬟࢖࣓࣮ࢪࣥࢢᐇ㦂ࡀከᩘሗ࿌ࡉࢀࡲࡋࡓࠋ⬻ᶵ⬟࢖࣓࣮ࢪࣥࢢࡢ㐍Ṍ࡟ࡼࡾࠊ⬻⛉Ꮫࡣ⬻ࡔࡅ࡛࡞ࡃࠊே㛫඲యࠊࡑࡋ࡚♫఍඲యࢆᑐ㇟࡜ࡍࡿ⛉Ꮫ࡛࠶ࡿ࡜࠸࠺ࣃࣛࢲ࢖࣒ᨵ㠉ࡀ࠾ࡇࡾࡘࡘ࠶ࡾࡲࡍࠋࡇࡢࡼ࠺࡞⬻ᶵ⬟࢖࣓࣮ࢪࣥࢢࡢᏛ㝿ⓗ࡞◊✲఍ࡣᾏእ࡛ࡶ౛ࡀᑡ࡞ࡃࠊᅜෆ࠿ࡽࡢཧຍ
⪅ࠊ≉࡟Ꮫ⏕࡟࡜ࡗ࡚ࡣከࡃࡢ่⃭ࢆᚓࡿᶵ఍ࡔࡗࡓࡇ࡜࡜ᛮ࠸ࡲࡍࠋㅮ₇ࡢ㛫࡟ࡣ࣏ࢫࢱ࣮ࢭࢵࢩࣙࣥࢆ㛤ദࡋࠊࢥ࣮ࣄ࣮ࢆᴦࡋࡳ࡞ࡀࡽ࣏ࢫࢱ࣮ࡢ๓࡛ᅜෆእࡢ◊✲⪅ࡀ⇕ࡃ㆟ㄽࢆ⧞ࡾᗈࡆ࡚࠸ࡿᵝᏊࡀぢࡽࢀࡲࡋࡓࠋᏛ⏕ࡢ࣏ࢫࢱ࣮Ⓨ⾲ࡶከࡃࠊỴࡋ࡚ὶᬸ࡜ࡣゝ࠼࡞࠸ⱥㄒ࡛ᾏእࡢㅮ₇⪅࡟⮬㌟ࡢ◊✲ࢆㄝ᫂ࡋࡓ⤒㦂ࡣࠊᏛ⏕࡟ྲྀࡗ࡚ᛀࢀࡽࢀ࡞࠸⤒㦂࡛࠶ࡗࡓࡇ࡜࡛ࡋࡻ࠺ࠋࡇࡢࡼ࠺࡞⮬↛⾜ື୰ࡢ⬻Ἴィ◊✲࡟㛵ࡋ࡚ࡣࠊᾏእㅮ₇⪅ࡢ୍ே࡛࠶ࡿ Klaus Gramann ඛ⏕ࡀ 2013 ᖺ 9 ᭶࡟ࢻ࢖ࢶ࡛㛤ദ࡟ࡋࡓࡢࡀึࡵ࡚࡛ࡍࠋᐇࡣ⚾ࡣࡑࡢ➨ 1 ᅇࡢ◊✲఍࡟ᣍᚅࡉࢀࡓࡇ࡜࡛ࠊࡇࡢศ㔝࡟῝ࡃ㛵ࢃࡿࡇ࡜࡟࡞ࡾࡲࡋࡓࠋ௒ᚋࡣᒣ
⏣ࢩ࣏ࣥࢪ࣒࢘ࢆዎᶵ࡟ࠊከࡃࡢᏛ⏕ࡀ᪂ࡋ࠸ศ㔝࡟ᣮᡓࡉࢀࡿࡇ࡜ࠊࡑࡋ࡚᪥ᮏࡀࡇࡢศ㔝ࢆ࣮ࣜࢻࡋ࡚࠸ࡃࡇ࡜ࢆ㢪ࡗ࡚࠸ࡲࡍࠋࡇࡢࡼ࠺࡞◊✲ୖࡢ㆟ㄽ࡟ຍ࠼࡚ࠊᾏእㅮ₇⪅࡜ᅜෆ◊✲⪅ࡀಶேⓗ࡞஺ὶࢆ῝ࡵࡓࡇ࡜ࡀⰋ࠿ࡗࡓ࡜ឤࡌ࡚࠸ࡲࡍࠋࢩ࣏ࣥࢪ࣒࢘ࡢࣉࣟࢢ࣒ࣛࡣࢱ࢖ࢺ࡛ࡋࡓࡀࠊึ᪥ኪࡢ᠓ぶ఍࡟ຍ࠼ࠊࢩ࣏ࣥࢪ࣒࢘ᚋࡣኪ㐜ࡃࡲ࡛ᾏእㅮ₇⪅࡜ኤ㣗ࢆඹ࡟ࡋࠊᪧ஺ࢆ ࡵࠊࡲࡓ᪂ࡓ࡞཭ேࡀከࡃ࡛ࡁࡲࡋࡓࠋࢩ࣏ࣥࢪ࣒࢘ࡢ‽ഛࡣ୍ᖺ௨ୖ๓࠿ࡽ⾜ࡗ࡚ࡁࡲࡋࡓࡀࠊࢩ࣏ࣥࢪ࣒࢘ࡀᡂຌ⿬࡟⤊ࢃࡾࠊ‽ഛ࡟᥃ࡅࡓປຊࡶሗࢃࢀࡲࡋࡓࠋ᭱ᚋ࡟࡞ࡾࡲࡋࡓࡀࠊࡇࡢࡼ࠺࡞᭷ព⩏࡞ࢩ࣏ࣥࢪ࣒࢘ࢆ㛤ദࡍࡿᶵ఍ࢆ୚࠼࡚ࡃࡔࡉࡾࠊከ኱࡞ࡈᨭ᥼ࢆ㡬ࡁࡲࡋࡓᒣ⏣⛉Ꮫ᣺⯆㈈ᅋ࡜㛵ಀ⪅ࡢⓙᵝ࡟ࠊᚰࡼࡾᚚ♩ࢆ⏦ࡋୖࡆࡲࡍࠋ࠙⤌⧊ጤဨ఍ጤဨࠚ⤌⧊ጤဨ㛗㸸 ⏣୰ᏹ࿴㸦໭㝣ඛ➃⛉Ꮫᢏ⾡኱Ꮫ㝔㸧⤌⧊ጤဨ㸸⚄ཎ⿱⾜㸦ᮾிᕤᴗ኱Ꮫ㸧ࠊᐑ⭜ㄔ㸦࢝ࣜࣇ࢛ࣝࢽ࢔኱Ꮫ㸧
࠙ᣍᚅㅮ₇⪅㸦ᾏእ㸧ࠚ
Scott Makeig (University of California San Diego), Tzyy-Ping Jung (University of California SanDiego), Irina N. Beloozerova (Barrow Neurological Institute), Lewis Chuang (Max Planck Institute,Tubingen), Luca Cocchi (Queensland Institute of Medical Research, Australia), Klaus Gramann (UTBerlin), J. Swaroop Guntupalli (Dartmouth College), Stefan Haufe (Columbia University), JohnIversen (University of California San Diego), Sandra Loo (University of California Los Angeles),Andrew Zalesky (University of Melbourne)
࠙ᣍᚅㅮ₇⪅㸦ᅜෆ㸧ࠚᕝேග⏨ (ᅜ㝿㟁Ẽ㏻ಙᇶ♏ᢏ⾡◊✲ᡤ)ࠊᐃ⸨つᘯ (⏕⌮Ꮫ◊✲ᡤ)ࠊ✄㑚ဴஓ (ᅜ❧᝟ሗᏛ◊✲ᡤ)ࠊ∵ሙ₶୍ (៞ᛂ⩏ሿ኱Ꮫ)ࠊ㤶ᕝ㧗ᘯ (ឡ▱ᕤᴗ኱Ꮫ)ࠊᕝ㘠୍᫭ (ᅜ㝿㟁Ẽ㏻ಙᇶ♏
ᢏ⾡◊✲ᡤ)ࠊ㕥ᮌ㝯ᩥ (⬻᝟ሗ㏻ಙ⼥ྜ◊✲ࢭࣥࢱ࣮)ࠊ୰㞍ඞᕫ (ᒾᡭ་⛉኱Ꮫ)ࠊ୰㔝⌔ᐇ (኱㜰኱Ꮫ)ࠊすᮏఙᚿ (⬻᝟ሗ㏻ಙ⼥ྜ◊✲ࢭࣥࢱ࣮)ࠊ㔝⃝Ꮥஅ (ᮾிᕤᴗ኱Ꮫ)ࠊᖹ㈡⍠⨾ (⟃Ἴᢏ⾡኱Ꮫ)ࠊྂᒇ᫴୍ (ࢯࢽ࣮ࢥࣥࣆ࣮ࣗࢱࢧ࢖࢚ࣥࢫ◊✲ᡤ)ࠊᰡ⃝⌶ྐ (኱㜰኱Ꮫ)ࠊ
ᒣୗᐂே (ᅜ㝿㟁Ẽ㏻ಙᇶ♏ᢏ⾡◊✲ᡤ)ࠊྜྷᮧዉὠỤ 㸦ᮾிᕤᴗ኱Ꮫ㸧ࠊDaniel Callan (⬻᝟ሗ㏻ಙ⼥ྜ◊✲ࢭࣥࢱ࣮)ࠊJason Palmer (኱㜰኱Ꮫ)ᒣ⏣ࢩ࣏ࣥࢪ࣒࢘ࡢ࣏ࢫࢱ࣮࡜ᅄྡࡢᇶㄪㅮ₇⪅㸦 % 6FRWW0DNHLJ ඛ⏕ࠊ & 7]\\
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