〔AREIPGR Vol. 37: 68-82, 2021〕
doi:10.24514/00007159
Original Paper

Exploration and Collection of Plant Genetic Resources
in Northeastern Cambodia, 2021
Sreynech OUCH 1), Dolla ROS 1), Sophany SAKHAN 1), Vathany THUN 1),
Makara OUK 1), Katsunori TANAKA 2), Yoichi KAWAZU 3)
1) Cambodian Agricultural Research and Development Institute , National Road 3, Prateahlang, Dangkor, P.O.
Box 01, Phnom Penh, Cambodia
2) Faculty of Agriculture and Life Science, Hirosaki University , 3 Bunkyo, Hirosaki, Aomori 036-8561, Japan
3) Institute of Vegetable and Floriculture Science , National Agriculture and Food Research Organization (NARO),
360 Kusawa, Ano, Tsu, Mie 514-2392, Japan
Communicated by H. OKUIZUMI (Research Center of Genetic Resources, NARO)
Received Aug. 19, 2021, Accepted Dec. 6, 2021
Corresponding author: Y. KAWAZU (e-mail: ykawazu@affrc.go.jp)

Summary
The Cambodian Agricultural Research and Development Institute (CARDI) conducted an exploration andcollection of plant genetic resources in northeastern Cambodia in 2021 in cooperation with the National Agriculture andFood Research Organization (NARO), Japan. The exploration was conducted within the framework of the Plant Genetic
Resources Asia (PGRAsia) project funded by the Ministry of Agriculture, Forestry, and Fisheries, Japan. The teamsurveyed mainly cucurbitaceous vegetables in two provinces of northeastern Cambodia (Ratanakiri and Stung Treng)and collected 70 samples, including 34 pumpkins (Cucurbita moschata ), 23 melons (Cucumis melo ), five cucumbers(Cucumis sativus ), two watermelons (Citrullus lanatus ), four eggplants (Solanum melongena ), and two chili peppers(Capsicum sp.). Seeds were divided between the CARDI and the Genetic Resource Center, NARO, using the standard
material transfer agreement (SMTA).
KEY WORDS: pumpkin, melon, cucumber, genetic resource, Cambodia

Introduction
It is important to collect new plant genetic resources
to develop crop varieties with advantageous traits, suchas resistance to pests or diseases, high quality, andhigh yield. With this goal in mind, the Plant Genetic
Resources Asia (PGRAsia) project, started in 2014
and funded by the Ministry of Agriculture, Forestry
and Fisheries, Japan, promotes the collection of plant
genetic resources. The objectives of the project are
to collect, characterize, evaluate, and develop plant
genetic resources for food and agriculture (PGRFA), in

collaboration between Asian countries and Japan, andestablish open databases for the effective use of PGRFA.
Here, we report the results of our surveys of mainly
cucurbitaceous vegetables, such as pumpkin, melon,
cucumber, and watermelon, in northeastern Cambodia.
Cucurbitaceous vegetables were collected from westernand northwestern Cambodia in 2014 (Matsunaga et al.
2015) and 2018 (Yashiro et al. 2019); northern Cambodia
in 2016 (Tanaka et al. 2017) and 2018 (Kondo et al.
2019); southern Cambodia in 2017 (Tanaka et al. 2019)and 2019 (Sudasinghe et al. 2020); and eastern Cambodia- 68 -

in 2015 (Tanaka et al. 2016), 2016 (Tanaka et al. 2017),
2017 (Matsushima et al. 2018), and 2019 (Kawazu etal. 2020). The 2014 to 2019 collection sites of melons,
pumpkins, and cucumbers have been summarized
by Kawazu et al. (2020). Surveys of cucurbitaceous
vegetables in northeastern Cambodia were conducted in2015 (Tanaka et al. 2016) and 2017 (Matsushima et al.
2018). The 2016 collection sites reported in Tanaka et al.
(2017) also included northeastern Cambodia. Therefore,
the study locations of the present study avoided thosereported by Tanaka et al. (2016), Matsushima et al.
(2018), and Tanaka et al. (2017). Due to COVID-19
regulations, the present study could not be conducted incollaboration with our Japanese colleagues. Therefore,doi:
the details of exploration and collection were discussed

by e-mail, and only Cambodian scientists conductedsurveys on plant genetic resources in this study.
Methods
The surveys were initiated on January 7, 2021,
and lasted for 10 days. With a Toyota Land Cruiser(Table 1, Fig. 1), the team travelled from Phnom Penhthrough Mondulkiri and spent four days collecting in
Ratanakiri. We left Ratanakiri to Stung Treng on January
12 where we conducted four days of sample collections.
The team left Stung Treng on January 16 and arrived at
Phnom Penh in the evening. Fruit and seed samples werecollected from farmers’ backyards or roadsides. In thecase of fruit samples, the team removed the seeds fromthe fruits, washed the seeds with tap water, and air-dried

TableÊ1.ÊItineraryÊofÊtheÊexplorationÊandÊcollectionÊofÊplantÊgenetic
Table
1. Itinerary of the exploration and collection of plant genetic
resourcesin northeastern Cambodia, 2021
resourcesÊinÊnortheasternÊCambodia,Ê2021
Date

Day Itinerary

Stay

1/7

Thu PhnomÊPenhÊ-ÊMondolkiriSenÊMonorom

1/8

Fri

MondolkiriÊ-ÊRatanakiri

Ratanakiri

1/9

Sat

Ratanakiri

Ratanakiri

1/10

Sun Ratanakiri

Ratanakiri

1/11

Mon Ratanakiri

Ratanakiri

1/12

Tue RatanakiriÊ-ÊStungÊTreng

StungÊTreng

1/13

Wed StungÊTreng

StungÊTreng

1/14

Thu StungÊTreng

StungÊTreng

1/15

Fri

StungÊTreng

StungÊTreng

1/16

Sat

StungÊTrengÊ-ÊPhnomÊPenh

(monthÊ/Êday)

doi:

47-55

69-70

57-64

56
46-49

65-68

41-45

13-18
12
1-7

8-11

22-40
19-21

Fig.Fig.
1. Sitesin northeastern
Cambodia
wherewhereplant plant
genetic
resourceswere collected.
1. Sitesin Northeastern
Cambodia
genetic
resources
wereSamples156
werecollected
inRatanakiri,
and
57
70
were
collected
inTreng.
collected. Collection numbers 1-56 were collected in Ratanakiri, and Stungcollection
numbers 57-70 were collected in Stung Treng.

- 69 -

them in fine-mesh nets. Most farmers stored the seeds ofdifferent vegetables together, and in this case, the teamseparated the seeds on site. The passport data of eachsample were recorded, including the local vegetable
name, vegetable status, sowing season, harvest season,
usage, and cultivation methods. The team also recorded
information regarding the collection sites, including
place name, latitude, longitude, and altitude, using a
Garmin Foretrex 401 (Garmin International Inc., Olathe,
KS, USA).

previous collections in Cambodia (Okuizumi et al. 2017;
Matsushima et al. 2018; Kawazu et al. 2020). Seed andfruit samples were collected from four different ethnic
groups, Tumpoun, Kroeung, Leav, and Charay, eachreferring to pumpkins by their local name. In Tumpoun,
pumpkins are called “ropeuy”; in Kroeung, “ropov”;
in Leav, “ma-euk” or “lpov”; in Charay, “plery” or“yatol”; and Khmer, also “lpov.” The Tumpoun groupmostly grows pumpkin in slash-and-burn fields withother cereal crops, such as rice and sesame. Pumpkinseeds were sown at the beginning of the wet season(June), and fruits were harvested at the end of the wet
season (October). In Cambodia, most farmers do notuse fertilizers and chemicals in pumpkin cultivation.
The pumpkin collected in this study was grown without
irrigation, fertilizer, or chemical application.

Results and Discussion
The team collected 70 samples, including 34
pumpkins (Cucurbita moschata ), 23 melons (Cucumis
melo ), five cucumbers ( Cucumis sativus ), twowatermelons (Citrullus lanatus ), four eggplants (Solanum
melongena ), and two chili peppers (Capsicum sp.)
(Table 2). Table 3 contains detailed information for all
the samples. None of the farmers used any fertilizers
or chemicals during the entire growing season, and all
harvests were kept for subsistence only. The seeds wereequally divided between the CARDI and the Genetic
Resource Center, NARO, based on the standard materialtransfer agreement (SMTA).

Melon
Twenty-three melon samples were collected.
Farmers mostly cultivate melons with rice during thewet season, in upland areas or slash-and-burn fields.
The local people believe that slash-and-burn practices
provide fertile soil, with no pests or diseases. Melon
cultivation was for subsistence only, and fertilizers andchemicals were not applied. Some farmers in Cambodia,
use fertilizers and chemicals when growing melons forthe market. Most farmers in this study stored their seeds
in nets or plastic bags. Only one farmer, who provided
20-025 and 20-026, stored seeds in a hollow bottle
gourd and a metal box with mixed pumpkin, melon, andwatermelon seeds (Photo 1). Sample 20-028 was also
stored with other crop seeds. In these cases, the seeds
had to be separated on site (Photo 2). The collected
melon seeds varied in length (mean of 7.9 mm, ranging
from 5.5 mm [Sample 20-007] to 11.7 mm [Sample 20043]).

PumpkinAll pumpkin samples belonged to Cucurbita
moschata (Table 3). Nine fruit samples were collected,
and the other 25 samples were collected as seeds. The
average weight of the collected fruits was 1.5 kg, of
which 1.9 kg was the heaviest individual fruit (Sample
20-042) and 1.2 kg was the lightest (Sample 20-009 and20-055). The fruit shapes of samples 20-027, 20-042,
20-054, and 20-055 were flat, while that of sample 20009 were elongated; 20-010 were pyriform; and 20-011,
20-022, and 20-051 were narrow pyriform. Different
pumpkin fruit shapes have also been observed in
doi:

Table
ofthe
thegenetic
geneticresources
resources
collected
in northeastern
Cambodia,
Table 2.
2. A
A summarysummary ofcollected
in northeastern
Cambodia,20212021
Data

Province

District

Altitude

Cucurbita

Cucumis

CucumisCitrullus

Solanum

Capsicum(m)moschata

melosativuslanatusmelongena

sp.

1/8

Ratanakiri

Lampat131 - 147

6

2

2

1/8

Ratanakiri

Borkeo140 - 172

4

2

2

1/9

Ratanakiri

O'Yadav

197 - 240

9

8

1

1/9

Ratanakiri

Ondoung Meas

112 - 11522Veoun Sai

95 - 102

6

3

1

10

3

1

7

1

1/10 Ratanakiri

1/11 Stung Treng Thala Borivat

65 - 792

1/11 Stung Treng Sesan

49 - 55

3

1/11 Stung Treng Seim Bouk

30 - 55

1

2

1/11 Stung Treng Thala Borivat
Total

60

1

1

34

23

1

1

Total

1
1

12
9
19

1

5
2

- 70 -

4
2

5

2

4

2

70

Eggplant
The team collected four eggplant samples (two from
Ratanakiri and two from Stung Treng). Three of themwere collected as seeds and the other as fruit. Farmers
grow several plants in the backyard for household
consumption and harvest them year-round. Kroeung
and Leav call eggplant “trob,” as in Khmer, while in
Tumpoun eggplant is called “prao.”
Chili pepper
Two chili pepper samples were collected from
Ratanakiri. One was collected from a Tumpoun farmer as
seeds, and the other from a Charay farmer as fruits. Chili
is called “hang” by Charay and “mates,” as in Khmer, by
Tumpoun.

Photo 1. A fruit of bottle gourd (circle) and a metal box(rectangle) are used to store seeds.

Acknowledgements
This work was supported by a MAFF commissioned
project study on “A Collaborative Research Projecton Characterization and Evaluation of Plant Genetic
Resources for Food and Agriculture (PGRAsia)” Grant
Number JPJ007117.
Photo 2. Seed separation in the farmer’s front yard.

References
Kawazu Y, Kuzuya M, Ouch S, Sakhan S and Ouk M(2020). Collaborative exploration of Cucurbitaceae
genetic resources in eastern Cambodia, 2019.
AREIPGR 36: 92-111.
[Genebank, NARO], [JaLC]
Kondo F, Layheng S, Tokuda M, Rathnayaka RMSMB,
Sophany S and Matsushima K (2019). Collaborative
exploration of plant genetic resources in northCambodia, 2018. AREIPGR 35: 162-184.
[Genebank, NARO], [JaLC]
Matsunaga H, Matsushima K, Tanaka K, Theavy S,
Layheng S, Channa T, Takahashi Y and Tomooka N(2015). Collaborative exploration of the Solanaceae
and Cucurbitaceae vegetable genetic resources in
Cambodia 2014. AREIPGR 31: 169-187.
[Genebank, NARO], [JaLC]
Matsushima K, Layheng S, Hatakeyama K, Kurumada S
and Sophany S (2018). Collaborative exploration ofplant genetic resources in eastern Cambodia, 2017.
AREIPGR 34: 118-136.
[Genebank, NARO], [JaLC]
Sudasinghe SP, Leakhena M, Bando K, Yamaguchi
K, Sakhan S, Ouk M and Matsushima K (2020).
Collaborative exploration of plant genetic resourcesin southern Cambodia, 2019. AREIPGR 36: 128 147.
[Genebank, NARO], [JaLC]

CucumberFive cucumber seed samples were collected from
Ratanakiri. These samples were provided by Tumpoun
and Charay farmers. According to the farmers, theyusually buy cucumber seeds at markets as they mostly
harvest young cucumber fruit that lack seeds. Only some
farmers that preferred preserving their landraces saved
seeds for the next season.
A farmer who provided us with seeds reported
that these landraces bear large fruits weighing up to
0.5 kg. Cucumber is cultivated in the wet season withupland rice, similar to pumpkins and melons. Fruit were
harvested for household consumption only. As part ofa traditional cultivation technique, farmers do not use
fertilizers or chemicals.
Watermelon
Two samples of watermelon were collected from
Charay farmers in Ratanakiri, whom called it “ov leuk,”in Khmer. The cultural practice of watermelon is likepumpkin, melon, and cucumber. The farmer that provided
watermelon seed 20-024 reported that this landrace wasresistant to pests, but that the flesh sweetness was very
low. The fruit of this landrace bears so many seeds that
seeds from one fruit were enough for the next season.
The seeds of the two watermelon samples had similarcolors (light brown).

- 71 -

Tanaka K, Duong TT, Yamashita H, Lay Heng S,
Sophany S and Kato K (2016). Collection of cucurbit
crops (Cucurbitaceae) from eastern Cambodia, 2015.
AREIPGR 32: 109-137.
[Genebank, NARO], [JaLC]
Tanaka K, Shigita G, Sophea Y, Thun V, Sophany
S and Kato K (2017). Collection of melon andother cucurbitaceous crops in Cambodia in 2016.
AREIPGR 33: 175-205.
[Genebank, NARO], [JaLC]
Tanaka K, Shigita G, Dung TP, Sophea Y, Thun V,
Sophany S and Kato K (2019). Collection of melon
and other cucurbitaceous crops in Cambodia in 2017.
AREIPGR 35: 121-146.
[Genebank, NARO], [JaLC]- 72 -

Yashiro K, Tanaka K, Sophea Y, Thun V, Sophany S
and Kato K (2019). Collaborative exploration of
Cucurbitaceae vegetable genetic resources in westernand northwestern Cambodia in 2018. AREIPGR 35:
147-161.
[Genebank, NARO], [JaLC]

カンボジア北東部における植物遺伝資源の探索・収集，2021 年
Sreynech OUCH 3)・Dolla ROS 1)・Sophany SAKHAN 1)・Vathany THUN 1)・
Makara OUK 1)・田中 克典 2)・川頭 洋一 3)1) カンボジア農業開発研究所
2) 国立大学法人 弘前大学農学生命科学部
3) 国立研究開発法人 農業・食品産業技術総合研究機構 野菜花き研究部門
和文摘要
カンボジア農業開発研究所（CARDI）は国立研究開発法人 農業・食品産業技術総合研究機構（農研機構）と連携して，2021 年にカンボジア北東部において植物遺伝資源の探索・収集を実施した．この探索は，農林水産省委託プロジェクト研究「海外植物遺伝資源の民間等への提供促進」（PGRAsia プロジェクト）の予算により実施された．本探索ではカンボジア北東部の 2 つの州（Ratanakiri, Stung Treng）を訪問し，合計 70 点の遺伝資源を収集した．その内訳はニホンカボチャ（Cucurbita moschata ）が 34 点，メロン（Cucumis melo ）が23 点，キュウリ（C. sativus ）が 5 点，スイカ（Citrullus lanatus ）が 2 点，ナス（Solanum melongena ）が 4 点，
トウガラシ（Capsicum sp.）が 2 点である．収集された遺伝資源の種子の半分は CARDI に保管され，残りの半分は標準材料移転契約（SMTA）に基づいて農研機構遺伝資源センターに送付された．

- 73 -

Table 3. Passport data of the genetic resources collected in northeastern Cambodia in 2021
JP No.

- 74 -

Coll. Crop Name Species
ProvinceNo.
273988 20-001 Pumpkin
Cucurbita moschata Ratanakiri
Ratanakiri
273989 20-002 Cucumber Cucumis sativus
Cucumis melo
Ratanakiri
273990 20-003 Melon
Solanum melongena Ratanakiri
273991 20-004 Eggplant
Ratanakiri
273992 20-005 Cucumber Cucumis sativus
Ratanakiri
273993 20-006 Chili pepper Capsicum sp.
Cucumis melo
Ratanakiri
273994 20-007 Melon
Cucurbita moschata Ratanakiri
273995 20-008 Pumpkin
Cucurbita moschata Ratanakiri
273996 20-009 Pumpkin

District
LampatLampatLampatLampatLampatLampatLampatLampatLampat273997 20-010 Pumpkin

Cucurbita moschata Ratanakiri

Lampat273998 20-011 Pumpkin

Cucurbita moschata Ratanakiri

LampatCucurbita moschata Ratanakiri
273999 20-012 Pumpkin
Cucurbita moschata Ratanakiri
274000 20-013 Pumpkin
Cucurbita moschata Ratanakiri
274001 20-014 Pumpkin
Cucumis melo
Ratanakiri
274002 20-015 Melon
Cucurbita moschata Ratanakiri
274003 20-016 Pumpkin
Cucumis melo
Ratanakiri
274004 20-017 Melon
Ratanakiri
274005 20-018 Chili pepper Capsicum sp.
Cucurbita moschata Ratanakiri
274006 20-019 Pumpkin
Ratanakiri
274007 20-020 Cucumber Cucumis sativus
Cucumis melo
Ratanakiri
274008 20-021 Melon
Cucurbita moschata Ratanakiri
274009 20-022 Pumpkin
Cucumis melo
Ratanakiri
274010 20-023 Melon
Ratanakiri
274011 20-024 Watermelon Citrullus lanatus
Cucumis melo
Ratanakiri
274012 20-025 Melon
Cucurbita moschata Ratanakiri
274013 20-026 Pumpkin
Cucurbita moschata Ratanakiri
274014 20-027 Pumpkin
Cucumis melo
Ratanakiri
274015 20-028 Melon
Cucumis melo
Ratanakiri
274016 20-029 Melon
Cucurbita moschata Ratanakiri
274017 20-030 Pumpkin
Cucurbita moschata Ratanakiri
274018 20-031 Pumpkin
Cucumis melo
Ratanakiri
274019 20-032 Melon
Cucurbita moschata Ratanakiri
274020 20-033 Pumpkin
Ratanakiri
274021 20-034 Cucumber Cucumis sativus
Cucurbita moschata Ratanakiri
274022 20-035 Pumpkin
274023 20-036 Cucumber Cucumis sativus
Ratanakiri
Cucurbita moschata Ratanakiri
274024 20-037 Pumpkin
Cucurbita moschata Ratanakiri
274025 20-038 Pumpkin

LampatBorkeoBorkeoBorkeoBorkeoBorkeoBorkeoBorkeoBorkeoBorkeoO'YadavO'YadavO'YadavO'YadavO'YadavO'YadavO'YadavO'YadavO'YadavO'YadavO'YadavO'YadavO'YadavO'YadavO'YadavO'YadavO'Yadav

CommuneVillageNorth
East
Altitude Tribe
Collection Local name Remarks
Latitude Longitude(m)methodSeda
Patok
13 29 51 107 06 46
139 Tompoun Single fruit ropeuy
Slash-and-burn field. Sowing: June. Harvest: Oct.
Seda
Patok
13 29 51 107 06 46
139 Tompoun Bulk fruit ya-dauch
Slash-and-burn field. Sowing: June. Harvest: Oct.
Seda
Patok
13 29 51 107 06 46
139 Tompoun Bulk fruit pe-poong Slash-and-burn field. Sowing: June. Harvest: Oct.
Seda
Patok
13 29 51 107 06 46
139 Tompoun Bulk fruit prao
Slash-and-burn field. Sowing: June. Harvest: Oct.
Seda
Thmey
13 30 35.7 107 08 22.7 146 Tompoun Bulk fruit ten-pia
Slash-and-burn field. Sowing: June. Harvest: Oct.
Seda
Thmey
13 30 35.7 107 08 22.7 146 Tompoun Bulk fruit mates
Grow in farm yard. Sowing: all year around.
Seda
Thmey
13 30 35.8 107 08 22.8 147 Tompoun Bulk fruit tro sok srov Slash-and-burn field. Sowing: June. Harvest: Oct.
Seda
Thmey
13 30 42.7 107 20 28.3 131 Tompoun Bulk fruit ropeuy
Fruit: 1.8 kg. Sowing: June. Harvest: Oct.
Seda
Thmey
13 30 42.7 107 20 28.3 131 Tompoun Single fruit ropeuy
Fruit: 1.2 kg. Fruit length: 36 cm, Fruit width: 8 cm.
Sowing: June. Harvest: Oct.
Seda
Thmey
13 30 42.7 107 20 28.3 131 Tompoun Single fruit ropeuy
Fruit: 1.7 kg, Fruit length: 19 cm, Fruit width: 15 cm.
Sowing: June. Harvest: Oct.
Seda
Thmey
13 30 42.7 107 20 28.3 131 Tompoun Bulk fruit ropeuy
Fruit: 1.6 kg, Fruit length: 30 cm, Fruit width: 14 cm.
Sowing: June. Harvest: Oct.
Seda
Samut Kroam 13 33 04.5 107 10 23.7 145 Khmer Bulk fruit lpov
Intercrop with sugar cane. Sowing: June. Harvest: Oct.
SeoungSormorl
13 34 29.7 107 14 05.3 172 Tompoun Bulk fruit ropeuy
Sowing: June. Harvest: Oct.
SeoungYasoum
13 34 54.9 107 16 31.7 151 Khmer Bulk fruit lpov
Sowing: June. Harvest: Oct.
SeoungYasoum
13 34 54.9 107 16 31.7 151 Khmer Bulk fruit tro-sok
Sowing: June. Harvest: Oct.
SeoungYasoum
13 35 02.7 107 16 57.9 149 Jaray
Bulk fruit plery
Sowing: June. Harvest: Oct.
SeoungYasoum
13 35 02.7 107 16 57.9 149 Charay Bulk fruit mon
Sowing: June. Harvest: Oct.
SeoungYasoum
13 35 02.7 107 16 57.9 149 Charay Bulk fruit hang
Sowing: June. Harvest: Oct.
Seoung
Takok Jray
13 33 37.2 107 18 32.5 140 Charay Bulk fruit ya-tol
Grow with rice. Sowing: June. Harvest: Sep.
Seoung
Takok Jray
13 33 37.2 107 18 32.5 140 Charay Bulk fruit mon
Sowing: June. Harvest: Sep.
Seoung
Takok Jray
13 33 37.2 107 18 32.5 140 Charay Bulk fruit tro-sok srov Sowing: June. Harvest: Sep.
YatongPak
13 40 14.0 107 23 59.8 197 Charay Single fruit tol
Sowing: June. Harvest: Oct.
YatongPak
13 40 04.6 107 23 57.6 207 Charay Bulk fruit tro-sok srov Sowing: June. Harvest: Oct.
YatongPak
13 40 04.6 107 23 57.6 207 Charay Bulk fruit ov leuk
Sowing: June. Harvest: Sep.
YatongPak
13 40 04.4 107 23 58.1 211 Charay Bulk fruit chhel-mon Grow with rice. Sowing: June. Harvest: Sep.
YatongPak
13 40 04.4 107 23 58.1 211 Charay Bulk fruit lpov
Sowing: June. Harvest: Oct.
YatongTen
13 40 31.2 107 25 09.0 233 Khmer Single fruit lpov
Sowing: June. Harvest: Oct.
YatongTen
13 40 31.2 107 25 09.0 233 Khmer Bulk fruit tro-sok srov Sowing: June. Harvest: Sep.
YatongTensos
13 40 24.1 107 25 18.5 226 Charay Bulk fruit tro-sok srov Grow with rice. Sowing: June. Harvest: Sep.
YatongTensos
13 40 24.1 107 25 18.5 226 Charay Bulk fruit lpov
Grow with rice. Sowing: June. Harvest: Oct.
YatongTensos
13 40 30.8 107 25 06.9 228 Charay Bulk fruit lpov
Grow with rice. Sowing: June. Harvest: Oct.
YatongTensos
13 40 30.8 107 25 06.9 228 Charay Bulk fruit tro-sok srov Grow with rice. Sowing: June. Harvest: Oct.
YatongTen Ngol
13 40 57.1 107 25 26.6 228 Charay Bulk fruit lpov
Sowing: June. Harvest: Oct.
YatongTen Ngol
13 40 57.1 107 25 26.6 228 Charay Bulk fruit tro-sok srov Sowing: June. Harvest: Oct.
Saom Thom Saom Trok Jas 13 42 49.1 107 24 15.6 240 Charay Bulk fruit lpov
Sowing: June. Harvest: Oct.
Saom Thom Saom Trok Jas 13 42 49.1 107 24 15.6 240 Charay Bulk fruit tro-sok
Sowing: June. Harvest: Oct.
Saom Thom Saom Trok Jas 13 42 49.1 107 24 15.6 240 Charay Bulk fruit lpov
Sowing: June. Harvest: Oct.
Pok NhaiJas
13 44 26.6 107 26 12.7 210 Charay Bulk fruit lpov
Sowing: June. Harvest: Oct.

Table 3. (Continued).
JP No.

Coll. Crop Name Species
ProvinceNo.
274026 20-039 Melon
Cucumis melo
Ratanakiri
Cucumis melo
Ratanakiri
274027 20-040 Melon
Cucurbita moschata Ratanakiri
274028 20-041 Pumpkin

District

CommuneVillageNorth
East
Altitude Tribe
Latitude Longitude(m)Jas
13 44 26.6 107 26 12.7 210 Charay
Jas
13 44 26.6 107 26 12.7 210 Charay
Ka Jout kroam 13 52 24.9 107 18 10.6 112 Khmer- 75 -O'Yadav Pok Nhai
O'Yadav Pok Nhai
Ondoung Nhang
Meas274029 20-042 Pumpkin
Cucurbita moschata Ratanakiri Ondoung Nhang
Ka Jout kroam 13 52 25.0 107 18 09.5
Meas
274030 20-043 Melon
Cucumis melo
Ratanakiri Ondoung Nhang
Ka Jout kroam 13 52 24.7 107 18 11.3
Meas
274031 20-044 Melon
Cucumis melo
Ratanakiri Ondoung Nhang
Ka Jout kroam 13 52 24.7 107 18 11.3
Meas
274032 20-045 Watermelon Citrullus lanatus
Ratanakiri Ondoung Nhang
Ka Jout kroam 13 52 24.7 107 18 11.3
Meas
274033 20-046 Pumpkin
Cucurbita moschata Ratanakiri Veoun Sai Kajoun
Vong vay
13 44 25.3 106 59 08.4
Cucumis melo
Ratanakiri Veoun Sai Kajoun
Vong vay
13 44 25.3 106 59 08.4
274034 20-047 Melon
Cucumis melo
Ratanakiri Veoun Sai Kajoun
Vong vay
13 44 25.3 106 59 08.4
274035 20-048 Melon
Cucumis melo
Ratanakiri Veoun Sai Kajoun
Vong vay
13 44 25.3 106 59 08.4
274036 20-049 Melon
Cucurbita moschata Ratanakiri Veoun Sai Kajoun
Kajoun
13 57 23.1 106 53 14.4
274037 20-050 Pumpkin
Cucurbita moschata Ratanakiri Veoun Sai Kajoun
Kajoun
13 58 11.0 106 53 43.8
274038 20-051 Pumpkin
Solanum melongena Ratanakiri Veoun Sai Koh peakKoun
274039 20-052 Eggplant
Cucurbita moschata Ratanakiri Veoun Sai
Pong
13 58 23.1 106 49 55.5
274040 20-053 Pumpkin
Cucurbita moschata Ratanakiri Veoun Sai Veoun Sai
Veoun Sai
13 58 43.8 106 48 39.3
274041 20-054 Pumpkin
Cucurbita moschata Ratanakiri Veoun Sai Veoun Sai
Veoun Sai
13 58 43.8 106 48 39.3
274042 20-055 Pumpkin
Cucurbita moschata Stung Treng Thala
Hang ko ban Sakhoun
13 53 54.1 106 53 42.0
274043 20-056 Pumpkin
Borivat
274044 20-057 Pumpkin
Cucurbita moschata Stung Treng Thala
Hang ko ban Sakhoun
13 33 45.5 106 01 43.9
Borivat
274045 20-058 Pumpkin
Cucurbita moschata Stung Treng Sesan
Som khouy Srae tapan
13 33 48.5 106 02 32.5
Cucumis melo
Stung Treng Sesan
Som khouy Srae tapan
13 33 48.5 106 02 32.5
274046 20-059 Melon
Cucumis melo
Stung Treng Sesan
Som khouy Srae tapan
13 33 48.5 106 02 32.5
274047 20-060 Melon
Cucumis melo
Stung Treng Sesan
Som khouy Srae tapan
13 33 48.5 106 02 32.5
274048 20-061 Melon
Cucurbita moschata Stung Treng Sesan
Som khouy Srae tapan
13 33 48.5 106 02 32.5
274049 20-062 Pumpkin
Cucurbita moschata Stung Treng Sesan
Som khouy Srae tapan
13 34 13.6 106 03 32.8
274050 20-063 Pumpkin
Solanum melongena Stung Treng Sesan
Som khouy Srae tapan
13 34 13.6 106 03 32.8
274051 20-064 Eggplant
Cucumis melo
Stung Treng Seim
Koh
Koh Sampeay 13 26 43.9 105 56 56.7
274052 20-065 Melon
Bouk
Sampeay
274053 20-066 Melon
Cucumis melo
Stung Treng Seim
Koh
Koh Sampeay 13 25 22.6 105 56 49.3
Bouk
Sampeay
274054 20-067 Eggplant
Solanum melongena Stung Treng Seim
Koh
Koh Sampeay 13 25 22.6 105 56 49.3
Bouk
Sampeay
274055 20-068 Pumpkin
Cucurbita moschata Stung Treng Seim
Koh
Koh Sampeay 13 24 55.7 105 56 31.8
Bouk
Sampeay

Collection
methodBulk fruit
Bulk fruit
Bulk fruit

Local name Remarks
tro-sok srov Sowing: June. Harvest: Oct.
tro-sok srov Sowing: June. Harvest: Oct.
lpov
Sowing: June. Harvest: Oct.

115

CharaySingle fruit lpov

Sowing: June. Harvest: Oct.

115

CharayBulk fruit tro-sok srov Sowing: June. Harvest: Oct.

115

Khmer

Bulk fruit tro-sok srov Sowing: June. Harvest: Sep.

115

CharayBulk fruit ov-leouk

102
102
102
102
102
96
97
95
95
79

Kreoung Bulk fruit ropov
Sowing: June. Harvest: Oct.
Kreoung Bulk fruit ro-dob
Sowing: June. Harvest: Oct.
Kreoung Bulk fruit ro-dob
Sowing: June. Harvest: Oct.
Kreoung Bulk fruit tro-sok srov Sowing: June. Harvest: Oct.
Tompoun Bulk fruit lpov
Sowing: June. Harvest: Oct.
Tompoun Single fruit lpov
Kreoung Bulk fruit trob
Sowing: all year around.
Leav
Bulk fruit lpov
Sowing: June. Harvest: Oct.
Leav
Bulk fruit lpov
Sowing: June. Harvest: Oct.
Leav
Bulk fruit lpov
Sowing: June. Harvest: Oct.
Leav
Bulk fruit lpov
Sowing: June. Harvest: Oct.

65

Leav

Bulk fruit ma-euk

55
55
55
55
55
49
49
55

Leav
Leav
Leav
Leav
Leav
Leav
Leav
Khmer

Bulk fruit
Bulk fruit
Bulk fruit
Bulk fruit
Bulk fruit
Bulk fruit
Bulk fruit
Bulk fruit

30

Leav

Bulk fruit tro-sok srov Sowing: June. Harvest: Oct.

30

Leav

Bulk fruit trob

Sowing: June. Harvest: Oct.

47

Khmer

Single fruit lpov

Sowing: June. Harvest: Oct.

Sowing: June. Harvest: Sep.

Seeds from Banlong. Sowing: June. Harvest: Oct.

lpov
Sowing: June. Harvest: Oct.
tro-sok srov Sowing: June. Harvest: Oct.
tro-sok srov Sowing: June. Harvest: Oct.
tro-sok srov Sowing: June. Harvest: Oct.
lpov
Sowing: June. Harvest: Oct.
lpov
Sowing: June. Harvest: Oct.
trob
Sowing: June. Harvest: Oct.
tro-sok srov Sowing: June. Harvest: Oct.

Table 3. (Continued).
JP No.

Coll. Crop Name Species
Province
DistrictNo.
274056 20-069 Pumpkin
Cucurbita moschata Stung Treng Thala
Borivat274057 20-070 Melon
Cucumis melo
Stung Treng Thala
BorivatCommuneVillageO'trelNorth
East
Altitude Tribe
Latitude Longitude(m)
13 32 09.2 105 55 58.6
60
Leav

Collection Local name Remarks
methodBulk fruit lpov
Sowing: June. Harvest: Oct.

Thala
Thala

O'trel13 32 09.2 105 55 58.6

Bulk fruit tro-sok srov Sowing: June. Harvest: Oct.

60

Leav

- 76 -

Photos of collected samples

Sample Photo 1.
20-001 Cucurbita moschata

Sample Photo 2.
20-002 Cucumis sativus

Sample Photo 3.
20-003 Cucumis melo

Sample Photo 4.
20-004 Solanum melongena

Sample Photo 5.
20-005 Cucumis sativus

Sample Photo 6.
20-006 Capsicum sp.

Sample Photo 7.
20-007 Cucumis melo

Sample Photo 8.
20-008 Cucurbita moschata

Sample Photo 9.
20-009 Cucurbita moschata

Sample Photo 10.
20-010 Cucurbita moschata

Sample Photo 11.
20-011 Cucurbita moschata

Sample Photo 12.
20-012 Cucurbita moschata- 77 -

Sample Photo 13.
20-013 Cucurbita moschata

Sample Photo 14.
20-014 Cucurbita moschata

Sample Photo 15.
20-015 Cucumis melo

Sample Photo 16.
20-016 Cucurbita moschata

Sample Photo 17.
20-017 Cucumis melo

Sample Photo 18.
20-018 Capsicum sp.

Sample Photo 19.
20-019 Cucurbita moschata

Sample Photo 20.
20-020 Cucumis sativus

Sample Photo 21.
20-021 Cucumis melo

Sample Photo 22.
20-022 Cucurbita moschata

Sample Photo 23.
20-023 Cucumis melo

Sample Photo 24.
20-024 Citrullus lanatus- 78 -

Sample Photo 25.
20-025 Cucumis melo

Sample Photo 26.
20-026 Cucurbita moschata

Sample Photo 27.
20-027 Cucurbita moschata

Sample Photo 28.
20-028 Cucumis melo

Sample Photo 29.
20-029 Cucumis melo

Sample Photo 30.
20-030 Cucurbita moschata

Sample Photo 31.
20-031 Cucurbita moschata

Sample Photo 32.
20-032 Cucumis melo

Sample Photo 33.
20-033 Cucurbita moschata

Sample Photo 34.
20-034 Cucumis sativus

Sample Photo 35.
20-035 Cucurbita moschata

Sample Photo 36.
20-036 Cucumis sativus- 79 -

Sample Photo 37.
20-037 Cucurbita moschata

Sample Photo 38.
20-038 Cucurbita moschata

Sample Photo 39.
20-039 Cucumis melo

Sample Photo 40.
20-040 Cucumis melo

Sample Photo 41.
20-041 Cucurbita moschata

Sample Photo 42.
20-042 Cucurbita moschata

Sample Photo 43.
20-043 Cucumis melo

Sample Photo 44.
20-044 Cucumis melo

Sample Photo 45.
20-045 Citrullus lanatus

Sample Photo 46.
20-046 Cucurbita moschata

Sample Photo 47.
20-047 Cucumis melo

Sample Photo 48.
20-048 Cucumis melo

- 80 -

Sample Photo 49.
20-049 Cucumis melo

Sample Photo 50.
20-050 Cucurbita moschata

Sample Photo 51.
20-051 Cucurbita moschata

Sample Photo 52.
20-052 Solanum melongena

Sample Photo 53.
20-053 Cucurbita moschata

Sample Photo 54.
20-054 Cucurbita moschata

Sample Photo 55.
20-055 Cucurbita moschata

Sample Photo 56.
20-056 Cucurbita moschata

Sample Photo 57.
20-057 Cucurbita moschata

Sample Photo 58.
20-058 Cucurbita moschata

Sample Photo 59.
20-059 Cucumis melo

Sample Photo 60.
20-060 Cucumis melo

- 81 -

Sample Photo 61.
20-061 Cucumis melo

Sample Photo 62.
20-062 Cucurbita moschata

Sample Photo 63.
20-063 Cucurbita moschata

Sample Photo 64.
20-064 Solanum melongena

Sample Photo 65.
20-065 Cucumis melo

Sample Photo 66.
20-066 Cucumis melo

Sample Photo 67.
20-067 Solanum melongena

Sample Photo 68.
20-068 Cucurbita moschata

Sample Photo 69.
20-069 Cucurbita moschata

Sample Photo 70.
20-070 Cucumis melo

- 82 -