a it" -li- i. ~ ~ ~-~·ic.f

Transactions and Proceedingsof thePalaeontological Society of Japan
New Series

No. 64

Pa laeontolo gical Society of Japan
Dec. 15th, 1966

CONTENTS
TRANSACTIONS
Page

512.

Molluscan Fauna of the Higashi-lnnai Formation of Noto Peninsula,japan- II; Remarks on Molluscan Assemblage and Description ofSpecies .............................................. Koichiro lVIASUDA

513.

317

Fusulinids from the Funafuseyama Limestone in Yamagata-Gun. Gi fu
Prefecture, japan ................................ Kuniteru l\lATSUMARli514.

On a Reptilian Skeleton from the Palaeozoic Formation of San Paulo,

515.

Notes on Ammonites bravaisianus n'ORBJGNY from the Cretaceous ofBrazil .............................. Tokio SHlKAM.\ and Hiroshi OzAKI

338
351

France ...................... Tatsuro rvlATSUMOTO and l\lasayuki NUDA

359

PROCEEDINGS . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .

366

Systematic Index . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .

367

Index of Families, Genera and Species . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .

369

President:
Councillors:

Kiyoshi ASANO
Kiyoshi AsANO (Editor of ··Fossils"), Haruyoshi FUJJ!\IOTO, Tetsuro!·lANAI (Editor of Transactions and Proceedings), Wataru IIASIIJMOTO (Treasurer), Kotora liATAI, Ichiro HAYASAKA. Koichiro ICIIIKAWA. Taro KANAYA,
Teiichi KOBAYASHI. Tatsuro MATSUMOTO (Editor for Special Papers), Masaol\IINATO, Hiroshi OZAKI (Planning), Tokio SHIKAMA (Membership), Fuyuji
T AKAI (Chairman of Executive Councillors' Committee), Ryuzo ToRIYA:\IA
Secretaries:

Takashi

HAMADA.

Hisayoshi!Go, Yokichi TAKAYANAGI, Yasuhide

IWASAKIAll Communications relating to this journal should be addressed to the
PALAEONTOLOGICAL SOCIETY OF JAPAN
Geological Institute, Faculty of Science, University of Tokyo, japan

Trans. Proc. Palaeont. Soc. Japan. N. S., No. 61. pp. 317-337. pis. 35, 36, Dec 15, 1966

512. :MOLLUSCAN FAUNA OF THE HIGASHI-INNAI FORMATIONOF NOTO PENINSULA. JAPAN-II: REMARKS ON MOLLUSCANASSEMBLAGE AND DESCRIPTION OF SPECIESKOICHIRO rvfASUDA
Department of Geology. Miyagi University of Education. Sendai

1i~3t.>J':,I.',) J.HnlkJR'!OJJ!i,k{>jqJlfJ~!IJ ft:Wl\f-ll; fl.!t1illrlfU: N\OJili.~4i~: *~t.il->:l·"i 11-illi!l\1·.'11~
OJrlirnJl, 11i!t!!mlOJ ~i'lii \? ~~ L L i: t.: H H:.:Pflf.OJ u.}ft:t.;:~t(l.'lrfii:1JJ!t1P5:7kl·;::, ~ 'i' UJJ G:t.; ·1;:: Lt.:, ~ &r: 11 ~~H ~:Jc~£ t..t:ftl!. s=rOJt\:t;alf!l!OJl'H~rr~rrf.~:-:;~t.:,ll'J m ?J: -- .M

Concluding Remarks on
Molluscan Assemblage.)

The stratigraphical and geographicaldistributions of the molluscan shells andthe other fossils from the Higashi-Innai
Formation are shown in Tables 1-4.
For the sake of conyenience the strati·
graphical and geographical changes ofthe molluscan <>.ssemblages will be de·scribed from the lower to the upperhorizon.
(a) First horizon
During this stage the environmental
conditions may not have been favourable
to the molluscs because of the rapid
deposition of the sediments and unstablesedimentary basin. Howeyer, gradually
the conditions changed and the environment became favourable to the molluscs.
The molluscan assemblages are generally
characterized by the few number of
species and individuals and by suchbrackish water genera as Ostrea, Vicarya.
Vicaryella. Cerithidea, Batillaria. 1Vas·sari us, etc.
* Recei\'ed February 10, 1966: read Sept.
25, 1965 at Nagasaki.

317

From the little change in the assemblages from the different localities andtheir rather uniform and extensive distributions in the present field it is inferred that the molluscs were able tosurvive the environmental conditions involving variable and extreme salinities
and temperatures. The environmental
conditions of the first horizon may havebeen a shallow embayment (Fig. 1).
(b) Second horizon
With the opening of this stage theseas continued to transgress towards thesouth from the north.
The faunal assemblage of this stage
is varied, being composed of besides themolluscs, corals. echinoids, larger foraminifera and others. The molluscan species
become very numerous in both species
and individuals compared with the firsthorizon. The distributions of the molbscs are shown in Figs. 2 and 3. The
distributions of the hard bottom dwellingmolluscs coincide with the distributionsof such fossils as corals. bryozoans, calcareous algae, etc.
From the figures and tables it is noticed that the molluscan species show
wide variation in the assemblage geo-

Koichiro ATAs u oil

318

D
~
~ ~&Citlilll ••trJII flll'w'~OI'Y[NTS

Fig. 1.

Paleogeographical map of the First horizon.

·•

V'1c......-a,..

t~M~N,.

•

v.c.~rru..

/1101(KN$1S'

Fig. 2.

' ...

Distributions of Vicarya callosa japonica YABE and HAT AI. Vicaryel/a ishiiana
and Vicaryella no/oensis MASUDA (large circle: abundant to common occur.
renee of that species: small circle: few to rare occurrence of that species).
(YoKOYAMA)

'il2.

Fig. 3.

!llolluscan Fau.na of the Hiuashi-Innai

Relative nbundancy of hard bottom dwelling molluscs (dark colored).

D"··

fig. 4,§...

~IIUUJ5.!-I

c

z

t

J

liUlVtH•"tfr.AGNWEHTS.,~,~,,

P;lleol!"eographical m~.p of the Second horizon.

Koichiro MASUDA

~20

0

2

j

)

......

Fig. 5. Paleo6eographical map of the Third horizon.

!

<.::::

I

--

a;)
.\

~

-·

D ...,v...

~~DII~~SNWJ.Tl'lt~..f"Tt

.

'

I

'

·~

.41l

~
Fig. 6.

Paleogeographical map of the Fourth horizon.

•

l512.

Moll7tscan Fmma o/ the Higashi-lnnni

graphically. Among the molluscs there
are some good ecologic indicators as Vicarya callosa japonica YABE and I I AT AI.
l'icarye/la notoensis i\IASUDA. ,Vassarius
noloensis ;v!ASUDA. Rae/a sp.. Cyclina
japonica KAMADA, etc. These are good
indicators of brackish water ecologicalconditions. The occurrence of allopatricforms as Vicaryella notoensis !vlASUDA.
V. ishiiana (YOKOYAMA). Nassarius nota·
en sis :MASUDA, N. minoensis ITOIGAW A.
Cyclina japonica KAMADA and C. lzwulala~vlAKIY AMA are important for consideration of the paleoenvironmental condi·
tions during deposition.
From the fossils, it is inferred thatthe brackish environmental conditions
existed in the southern area near Kanagura. in the vicinity of Awagura and in
the vicinities of Kobunayama and Fujio.
Else\vhere the conditions are inferred tohave been marine. From the lithologic
characters and the thickness variation in
the vicinity of Kakuma there may haveexisted a channel extending from Otani
to near Fujio through Kakuma (Fig. 4).
(c) Third horizon
This stage is represented by only a
few fossil localities. The molluscan species from the uppermost part of Loc. No.
24 comprise brackish water forms as
Vicarya callosa japo11ica YABE and HA·TAl, Vicaryella notoensis l\lASUDA, etc.
Those from other localities are few inspecies and individuals but consist of amarine assemblage.
Judging from the lithological characteristics. fauna and distribution it isthought that there was some fluctuation
during the marine transgression of thisstage or at least some differential movements. The map of this stage is shownin Fig. 5.
(d) Fourth horizon
The seas of this stage extended to thesouthern area and the carbonaceous

321

sandy siltstone near Kyuden yielded
many molluscan shells but the fossil
molluscs are contained in the glauconite
bearing coarse-grained sandstone at otherlocalities. The molluscan fauna from
Kyuden (Loc. No. 22) consists of Raeta
sp .. Saxolucina k-hataii (OTUKA), Cerith·idea sp., etc. It seems that there may
have been a small embayment in the\'icinity of Kyliden opened towards thenorthwest. The paleogeographical mapof this stage is shown in Fig. 6.
(e) Conclusion
From the faunal elements and thepaleogeography it is considered that theHigashi-Innai Formation \Vas depositedunder the influence of a warm and shallow sea as shown by the occurrence oftropical to subtropical molluscs as Ctena,Katelysia, Huchelus, Monilea, Turbo. Nerila, Littorinopsis, Rissoina, Architectonica,
Vicarya, Vica1·yella, Pachycrommium, Cypraea, Apollon. Oliva. Vexillum. Mitra,
Philberlia, Cythara, Conus, etc. Thisview is upheld by the associated occurrence of reef building corals and by theassemblages of the smaller foraminifera
as pointed out by ASANO (1953).

Geological Age and Correlation
The Higashi- Innai Formation is characterized with such fossil molluscs as
Anadara makiyamai HATAI and NrsrYAMA, A. kurosedaniensis IIATAI andNISIY AMA, Chlamys ishidae MASUDA.
C. iwamure11sis ITOIGAWA, Crassatellites
suyamensis 0!::\0~HKADO. Fe/anie/la ferruginata (MAKIYA"-IA), ]oannisiel/a meisensis l'viAKIYAMA, Saxolucina k-hataii(OTUKA). Pillucina yokoyamai (OTUKA),
Clinocardium ogurai (OTUKA). Cyclina
japonica KA~IADA, C. lunulata i\IAKIYAMA. !Josinia akaisiana NOMURA. D.
tugaruana No~HJRA. Protothaca tateiwai(.MAKIY A~IA), l'itar itoi UviAKIY MtA),

Koichini iVASUDA
1\atelysia nakamurai II~EJIE, Soletel!ina
minoensis YoKo\' A«IA. Cultellus izumoensis YoKOYAMA. Turbo ozawai OTllKA.
l.ittorinopsis miodelicatula (hA~!A. Architectonica kurodac TsurH. Cerithidea lwnpokuensis l\IAKIYANIA. Vicarya cal/ostl
japonica Y ABE and HATM. Vicaryel/aishiiana (YoKOYA~IA), V. notoensi.~ J\IASUDA. Cerithium ancisum (YOKOY Ai\lA).
Calyptraea tubura OTL!KA, Polinices meisensis l'v1AK!YAMA. Neverita coticazae(\1AKIYAl\IA). Pachycrommium japonicum
KA~l\o. Sinum inefJ/um (YOI-:oYAMAl.
.·lpol/on osawanoensis TSUDA. Chicoreus
asanoi lviASUDA. Conus tokunagai OTUK.\,etc. Among the molluscan species the
Recent species or the ones referable to
the Recent species are only nine such as
Lima cf. goliath SoWERBY. Ostrea cf. denselame/losa LISC!IKE. Phvlyctiderma cf.
japonica (P!LSRR\'), l.ucinoma acutilinealum (CO!':RAD). Cryptomya busoensis Yo1\0YA:-.IA. ]oannetia cf. Clllllillgi (SoWER·
BY). Solen cf. gouldi Du:\KER. Dentalium
cf. hexagonum GoULD and D. weinlwuffi
DU:--IKER.
This formation is also characterizedwith larger foraminifers as Miogypsi1w/wtoi 1-IANZAW A and Opercu/ina complana/a japonica IIANZAWA and with
the Orito Flora (ISHIIlA. 196,1) which corresponds to the Daijima Flora (HUZIOKA,
191!9).
From the above mentioned accounts
the geological age of the Higashi-lnnai
fauna is considered to be Early Miocene(MASUDA. J962a, 1963).
From the molluscan species given in
the faunal lists the 1-Iigashi-lnnai For·mation is considered to be a correlativeof the faunas of the Heirokud6. I\leisen
and Ennichi Formations in Korea. theKurosedani Formation of Toyama Prefecture. the Uchiura and J<unimi of
Fukui. the Kawai of Shimane, the Orito
and Tsugawa ·of Niigata. the Nishi-

kurosawa of Akita, the Ainaigawa andTanosawa of Aomori. the Takinoue ofllokkaido, the Nlizunami of Gifu. theTsuzuki of Kyoto. the Fujishima of
Wakayama. the Tsuyama of Okayama.
the Sh<)bara of Hiroshima. the Kaisekizan of Mie. the Shiratori of Iwate. theKunugidaira of Fukushima and the Ajiri
of l\Iiyagi Prefectures.

Description of New Species andRemarks on Some Species
Family Nuculanidae
Genus Sacce/la WooORI:-\G, 1925
Saccel/a sailzaiensis MASUDA. n. sp.
Pl. 35. figs. la. b. 2, 3

1/ololyj>e:-DGS"', Reg. No. 4494.
/Jescription :-Shell small, moderately
thick, moderately inflated, transversely
elongated. longer than high: anterodorsal side slightly convex, passing
gradually into rounded anterior margin;postero-dorsal side nearly straight. descending abruptly into broadly roundedventral margin: posterior margin bluntly
pointed. Beak somewhat curved, pointed.
situated at nearly half of disc length.
Surface with much curved ridge extending from beak to posterior end. sculp·
tured with conspicuous, coarse. smoothconcentric lines nearly equal to theirinterspaces in width. Escutcheon narrow. oblong, lanceolated. defined byridge: lunule narrow, sharply definedby ridge from beak to near anterior
end: interior character unknown.

* Abbre\-iation for the D.::partmenl of Geology. Faculty of Education. Tuhoku Univcr
sity. Sendai.

512.

322

:l1ollnscan Pnuna. of the Hignshi.-Tnnni.

Dimensions (in mm) :Height
Length

7.8·
12.9

Depth

5**

7.8
13.4
3. 0

7.4
13. 2
ca. 3. 0

12.0
2.8

6. 3
12.0
2. 2

6. 0
ca. 9. 5
2. 2

5.8

10.2
2.5

*-holotype : *•-breadth

Remarks:- The present species resem·
bles Saccella minoensis ITOIGAW A (1960.
p. 263, pl. 1. fig. 1) from the rvlizunami
Group, Gifu Prefecture. but is distinguishable therefrom by its smaller shell.
coarse concentric lines on surface, beakat nearly central part of disc, one radi·
ating ridge extending from beak to posterior end and sharply defined lunule.
Saccel/a gordonis (YOKOYAi\·IA) (1920. p.
177, pl. 19. figs. 4, 5,1 from the Pliocene
Naganuma Formation, Kanagawa Prefecture is distinguished from the pre·sent one by its sharply pointed posterior
end. two radiating ridges and tine concentric lines. Also distinguishable from
Saccella confusa toyomaensis KAMADA(1962. p. 50. pl. 2, figs. 1-5) from thel'vliocene Numanouchi Formation. Fuku·
shima Prefecture by its rather large
and short shell.
Type locality :-Loc. No. 23.
Occurrence:- Type loc.-C; Loc. No.3
-C; Loc. No. 4-F.

19. ele\·ated. rather squarish. round-topped radial ribs and fine concentric growthlines; radial ribs nearly equal or a little
narrower than their interspaces, sometimes with faint, fine, shallow furrows
on surface near ventral margin; interspaces flat-bottomed: auricles medium
in size; anterior auricle larger than posterior one. with prominent byssal notch,
sculptured with a few radial threads andconcentric lines: posterior one truncatedbehind at about right angle. with sculpture similar to anterior one: hinge with
distinct cardinal crura. shallow and narrow resilial pit with short, distinct lateralridges. widely open nne! dislinct ctenoIium. Left yalve with sculpture nearly
equal to right valve. Interior surface
rather distinctly folded, with flutingsnear ventral margin.
Dimensions (in mm) :Valve
HeightLengthDepthRight*
21. 0
18.5

Right

14.0
12.0

5.0

Right
8.5
7. 8

Left

10.0
9.0
2.5

Family Pectinidae"-holotypc

Genus Aequipecten FISCHER. 1887

Remarll., :-This species is distinguished from Aequipecten yanagawaen.sis (No~1URA and ZINBO) (1936. p. 337. pl. 20. figs.
2a-b) from the Miocene Yanagawa Formation, Fukushima Prefecture by its
subequilateral shell. smaller apical angle.
fewer roundtopped radial ribs and nointercalary threads.
Chlamy.s ishidae
r-.L\suni\ (1962a. p. 171. pl. 18. figs. 21-24)
from the J'vliocene Kurosedani Formation.
Toyama Prefecture is distinguished fromthe present one by its larger number of

Aequipec/en matsunagiensis:'viASUDA, n. sp.
Pl. 35.

fig~.

•l-6

Holotype :-DGS, Reg. No. 4506.
Description :-Shell small. moderately
inflated, higher than long, nearly equilateral except ror auricles. nearly equivalve: valves radiately ribbed, apicalangle about 90". Right valve with about

K&i,chiru JIIASUDA

324

rounded radial ribs, intercalary threads
and characters of marginal serration,
Type locality:-Loc. No. 32.
Occurrence:- Type loc.-R; Loc. No.
3-R; Loc. No. 36-C.

Family t-.'lyochamidae
Genus ;\.f.radvra GJ~AY. 1840

Jlyadvra suzuensis l'viASUI>A, n. sp.
Pl. 35. figs. 10, 11

Family \-lytilidae
Genus Modiolus LA~IARCK, 1799,\lodiolus wanizakiensis l\IASUDA. n. sp.
Pl. 35. figs. Sa-b. 9

Holotype :-DGS, Reg. No. 4510.
Description :-Shell rather small, verythin. elongated, expanded posteriorly.
moderately inflated; antero-dorsal side
short. concave. rather abruptly passing
into bluntly pointed anterior end: pos·tero-dorsal side long. nearly straight nearbeak, rounded near posterior end, pas·
sing gradually into rounded poste_rior
margin; ventral margin nearly stra1ght
except for rounded antero- and postero·ventral margins. Beak rather low. situ·
ated near anterior end. Surface withvery faint, fine concentric growth lines
slightly folded near margin. Interior
characters unknown.

Dimensi01ts (in mm) :Height
LengthBreadth

18•
30.5
10.5

21
35
ca. 11.0

Holotype :-DGS. Reg. No. 4513.
Descnption :-Shell medium in size.
rather thin. triangularly elongate ovalin shape, longer than high: apical angle
about 120· : antero-dorsal margin nearlystraight, passing gradually into broadly
rounded ventral margin; postero-dorsal
margin gently curved downwards, passing abruptly into posterior margin andtruncated behind; ventral margin broadly rounded. Right valve somewhat inflated; sculptured with very coarse,
rather irregular, smooth concentric lines
which are narrower than their interspaces and rather well-developed keel
extending from beak to posterior corner,
making elongated posterior border whichis sculptured with broad, smooth concentric lines. Beak situated at nearly
half of disc length. Left valve nearly
fiat. with sculpture nearly equal to rightvalve. Interior margin smooth; hingewith elongated, rather wide excavation
bounded by distinct ridge: other interior
characters unknown.

·)_;)

Dimensions (in mm) :15.8•-holotype

Remarks:- This species resembles
Modiolus yasuhiroi KA~IADA (1962. p. 72.
pl. 3, figs. 13, H) from the Numanouc_hi
Miocene. Fukushima Prefecture but d1f·
fers by its small, protruded anterior end
and rounded posterior corner.
Type locality :-Loc. No. 33.
Occurrence :-Type loc.-C.

ValYe
HeightLengthDepthRight•
10.0
13.2
3.2

Right
10.2

Left
6.2

7.5
ca. 3. 5•-holotypc

Remarks:- The present new species isdistinguished from Myadora okadae
HATAI and MASUDA (1960, p. 33, figs. 1,2) from the l\liocene Moniwa lVIember ofthe Hatatate Formation, Miyagi Prefecture by its elongated shell, broad, smooth

51::2.

.Molluscan ]•'mow of the Hiyashi-lnnai

concentric lines and rather well developed keel. J!yadora ikebei HABE {1950, p.
27, pl. ,1. figs. 4-6), a Pleistocene andRecent species of japan, can be distinguished by its ill-developed keel on thesurface and regularly spaced coarse concentric lines.
Type locality :-Loc. No. 24.
Occurrence:- Type loc.-F; Lac. 33-F.
Family Carditidae
Genus Glans i'viEGERLE. 1811

Glans naomiae l'vlASUDA. n. sp.
Pl. 36, figs. 13a, b. 14

Holotype :-DGS. Reg. No. 4517.
/Jescription :-Shell small, moderately
thick, subovate, moderately inflated;
antero-dorsal side short. concave in frontof beak. passing gradually into roundedventral margin ; postero-dorsal side long,rounded. passing gradually into ventral
margin: beak pointed. curved forwards,
situated anteriorly at about 2/3 of shelllength: lunule small. sharply defined.
Surface with about 22, elevated. conspicuously imbricated radial ribs and fineconcentric growth lines: radial ribs somewhat broader than their interspaces;
scales on radial ribs disc-like, rather flaL.
becoming closely packed and imbricatedtowards ventral margin and inclined
apically. Interior margin with distinct
coarse dentation: muscle scars faintly
impressed: anterior cardinal and posterior cardinal teeth of left valve somewhat
developed; hinge characters of rightvalve unknown.
Oimensions (in mm} :- Height 9.6,
length 10.1. breadth 6.0 (holotype·l: height
7.0. length 7.6 (paratype).
Nemarks :-The present genus is rather
rare in the Tertiary strata of japan andmay be the first record.

Glans Jzirasei (DALL) (KJRA. 1960. p. 130,
pl. 52, fig. 20), a Recent species of japan,
can be distinguished from the present
one by the large and subquadrate shell.
Clans sagamiensi.~ KURODA and HABE
(HABE, 196L p. 122, app. 37. pl. 55. fig.
H). a f{ccent species of japan, also canbe distinguished from the present one
by its larger and subquadrate shell,
nearly equal radial ribs and interspaces.
The description of the present species
is based upon a single cojoined shell anda single small. isolated shell; they may
represent rather young shells. The newname is taken after the name of mydaughter. Naomi :VIASUDA.
Type localily:-Loc. No. 35.
Occurrence:- Type loc.-R; Loc. No.
23-R.

Family Lucinidae
Genus Ctena l\tl<)RCII. 1861
Ctena hataii ~IASUDA. n. sp.
Pl. 35, !igs. 15-20

Holotype :-DGS, Reg. No. ·1521.
Description :-Shell rather small. moderate in thickness, inequilateral. longer
than high. slightly inflated: antero-dorsalside rather long, somewhat concave infront of beak. passing gradually into
broadly rounded \'Cntral margin; posterodorsal side short, rounded, merging into
ventral margin: beak pointed, somewhat curved forwards. situated posteriorly at about one-third of shell length:
lunule small, oblong, rather distinct.
Surface with radial ribs. intercalarythreads and concentric growth lines:
radial ribs ten or a little more. somewhat
broader than their interspaces, divided
into two parts near beak and further
bifurcate at about half of disc height:
intercalary threads between radial ribs

:32li1\oichiro MASUDA

usually appear at upper half of disc andsometimes bifurcate ventrally: secondary intercalary threads between radialribs and primary intercalary threads ap-

pear near ventral margin: concentric
growth lines distinct.
Lateral teeth
rather large and distinct ; inner margin
finely crenulated.

Dimensions (in mm) :Height
Length
Depth

9.8*
11. 1
3.0

11.2
12.4
3.5

9.6
10.4
3.0

9.0
10.0
2.:>

9.0
10.0

7. 5

8.2
2.0

6.0
6.2

*-holotype

Remarks:-This new species is named
in honor of Professor Kotora HATAJ of
the Tohoku University.
The present species resembles Ctena
minoensi.~ iTOIGAW A (1960, p. 269. pl. 2.
fig. 6) from the ~liocene 1\lizunami groupof Gifu Prefecture and Ctena delicatula(P!LSBRY} (1904, p. 555. pl. '11. figs. 15, 16).
a Recent species of Southern Japan. But
the former is distinguished from hataii
by its small shell, coarse radial ribs andno intercalary threads. and the latter by
its large shell and rather simple surface
sculpture.
Type locality:-Loc. No. 30.
Occurrence:-Type loc.-C: Loc. No.
25-F; Loc. No. 32-R: Loc. No. 35-F:
Loc. No. 36-R.
Family Veneridae
Genus Cyclina DESHAYES. 1849
Subgenus Cyc/inorbis MAKIY Ai\IA. 1926
Cyclina (Cyclinorbis) lunulata

Holotype :-Geological Survey of Chosen (Korea). Reg. No. 32.
Remarks :-In 1952 KAMADA described
Cvclina japonica, n. sp. based upon thespecimens from the Higashi-Innai Formation at Tokunari (Loc. No. 12) in thepresent field and discussed on the Recent
and fossi I species of the genus Cyclina
of Japan and Korea.
Among the specimens from the Higashilnnai Formation, the ones referable to
Cyclina japonica KAMADA occur only at
Tokunari (Loc. No. 12), the type localityof Cyclina japonica. The specimens fromthe localities other than just mentioned
are referred to Cyclina lwwlata MAKIYA:\IA.
From the morphological characters it
is evident that Cyclina japonica KA?IIADA
is closely related to Cyclina lunulala
MAKIYAI\IA.
Type localily :-Nanseki, ~leisen District. North Korea.
Occurrence :-Loc. No. 23-F: Loc. No.
2c!-F; Loc. No. 3-F.

MAKIYAMA, 1926
Pl. 35. fig. 33
1926. Cyclina (Cyclinorbis) lrmulata JVIAKIYA:O.IA, jour. Coli. Sci., /mfJ. Univ. Kyoto.
Ser. B. Vol. 2. No.3. p. 158, pl.l::l. tig.l.
1952. Cyclina (Cyctinorbis) lunulata MAKIYA~I.\. KA~IADA, Trans. Proc. Pa/aeon/.
Soc. japan, N. S .. No. 6, p. 170, pl. 15.
figs, 5, 6.

Genus Paphia RtioJ:\G, 1798
Subgenus Paphia s. s.
Paphia (s. s.) euglypta ohiroi MASUDA,
n. subsp.
Pl. 35. figs. 22. 23

lioloiype :-DGS, Reg. No. 4527.

Molluscan Fn·una. of the fhyashi-fnnai

512.

Description :--Shell rather large, transyersely elongated, moderately inflated;
antero-dorsal side short, broadly concave.
passing gradually into rounded anteroventral margin; postero-dorsal side long.
slightly convex but rounded near posterior end and merging into rounded postero-ventral margin: ventral margin

nearly straight : beak situated anteriorly
at about 2/3 of shell length. Surface withvery coarse, round-topped concentriclines; concentric lines rather fine near
beak but tend to become coarse and tostrengthen with growth. Interior characters unknown.

Dimensions (in mm) :Height
Length
Depth

29.0•
53.0
9 0

23.5
-18.0

27. 5
ca. '18. 0

29.5

52.0

23.5
43.0
7.;)

20.5
ca. 36. 0
7.0*-holotypc

Remarks :-This new subspecies ismuch related to J>aphia euglypta (PHILIPPI)(1848. p. 76. pl. 7, fig. 3), a Recent speciesof Southern japan, but it is distinguishable therefrom by its oblong shell, broadly concaved antero-dorsal side and very
coarse concentric lines. This species canbe distinguished from Paphia exil/istakaokaensis SIIUTO (1957. p. 1-15. pl. 2.
figs. 5-7. 21-22) from the l\liocene l\liyaz:.~ki group of l'vliyazaki Prefecture by
its large shell and very coarse concentriclines. Paphia pseudoliratus VRrmE:'\·
BURG (1928. p. :IS7. pl. 31. figs. 2-5) fromthe !VIiocene l'vlekran Series of India isdistinguished from the present one byits concentric folds which are broader
than their intercalary furrows.
This new subspecies is named inhonor of ~vir. Suyft OHIRO who helped thewriter in the field.
Type /ocali/y .-Loc. No. 38.
Occurrence·- Type loc.-F: Loc. No.

37-C: Loc. No. 39-R.

Paphia 1,s. s.:' mzuensis ::v!ASUDA. n. sp.
Pl. 35, figs. 2J. ~5

Ho/otype :-DGS. Reg. No. 15:30.
Descrijllion.·-Shell of medium size.
moderately inflated. transversely elongated ; antero-dorsal side short. slightly
concave. passing gradually into rather
acutely rounded anterior margin: posterodorsal side long. nearly straight. passing
gradually into rather acutely rounded
posterior margin: ventral margin broadly
rounded; beak pointed. situated anteriorly about one-third of shell length: escutcheon narrow. rather sharply defined.
Surface with rather low. irregular. somewhat lamellated, close-set, coarse concentric lines: concentric lines at posterodorsal side somewhat more prominent
than those of other parts. broader thantheir interspaces.
Interior characters

Dimensions tin mm) :-Length

25.3·
13.0

Depth

s.;:;

Height21. 2
3tl. 5
7.5

18.0
29.0

*-holotype

1-t.O

12.5

22.5
5.5

20.0
:{ 5

8.5
12.5
3.0

N:oichiro MASUDA
unknown.
Remarks:- The present species can be
distinguished from Paphia hirabayashii
0TUKA ll938. p. 12. pl. 1, fig. 9, pl. 2,
figs. 16, 19) from the .t\liocene Shiroyama
Formation, Shizuoka Prefecture by its
much elongated shell and finely lamel·
lated coarse concentric lines. Papl:ia
grata tsumaensis SHUTO (1957, p. 153, pl.
12. figs. 9-11) from the Ivliyazaki Miocene.
Miyazaki Prefecture resembles the pre·sent species. but SHuTo's species is dis·tinguishable from suzuensis by its fineconcentric lines. This species can be distinguished from Paphia pseudoliratus
VREDENBURG (Op. cit.) by the shell formand surface sculpture.
Type locality :-Loc. No. 30.
Occurrence:- Type loc.-F: Loc. \"o.

23-F.
Genus Venaupis LA!\IARCK. 1818
Subgenus Amygdala ROMER. 1857
Venerupis (Amygdala) takagii(MASUDA).

1955

Pl. 35, figs. 28a, b, 29
1955. Callista

chi111msis

takagii

MASUDA,

Trans. Proc. Palaeont. Soc. japan, N. S ..
No. 20, p. 121. pl. 19. fig. 7.

Holotype :-DGS, Reg. No. 2501.
Remarks :-Cal/ista chineusi.~ lakagii
was described by the writer from thelligashi-Innai Formation at Tokunari(Loc. No. 12) based upon two. small. unfavourably preserved specim~ns. Numerous rather well preserved specimens
referable to /a/wgii were collected fromthe Higashi-Innai Formation at different
localities. These specimens show thattakagii should be refered to the genus
Venerupis and subgenus _,lmygdala andthat the type specimens may be youngshells.
The newly collected specimens takethe following description: Shell medium
to small. rather thin, moderately inflated,transversely elongated ; antero·dorsalside short. slightly rounded, passing
gradually into rounded anterior margin;postero-dorsal side long. nearly straight,
grading into rounded posterior margin;
ventral margin nearly straight to slightly rounded at central part of shell but
tend to become rounded at both anteroand posterior ends; beak pointed, situated anteriorly at about 3/5 of shelllength, escutcheon narrow. rather distinctly defined. Surface sculptured withnumerous. close-set. rater lo\v, fine radial
threads and fine concentric growth lines:
radial threads nearly equa I to theirinters paces; concentric lines rather more
distinct at anterior part than at posterior
part and appear somewhat decussated.
Pallial sinus large. deep, nearly horizontal; upper line subparallel withlower, broadly rounded at end; interior
margin smooth ; hinge characters unknown.
The present species resembles llenerupis (Amygdala) japonica (DESIIAYES)(KIRA. 1960, p. 1-14, pl. 56. fig. 23), a Recent species of japan. but it is distinguishable therefrom by its rather small,oblong shell. nearly straight ventral
margin. rather low, close-set radial
threads. faint. fine concentric growthlines and large pallial sinus. Anotherclosely related species is Veuerupis(_,lmygdala) uariegn/n (HANLEY) (SOWERBY, 1855, p. 696. pl. 151, figs. 133-138),
a Recent species of Southern japan. butrariegata has a rather large shell. somewhat rounded postero-dorsal side andshallow pallial sinus.

512.

Mollu.<;·:an Fa11nn of the Higashi-Innni

Dimensions (in mm) :Height
LengthDepth

11. 5•
19.2

18.5

16.8

13.5

13.4

27.8
9. 8**

2S.3

21.0

21.3

5.0

1!.0

6.5
11. 2
2.3

* .Jwlotype: "'-breadthType locality:-Loc. No. 12.
Occurrence:-Type Loc.-F; Loc. No.
33-R; Loc. No. 35-C.

Family Erodonidae
Genus .:\nisocorbula IREDALE, 1930
Anisocorbula oltiroi l\•IASUDA. n. sp.
Pl. 35, figs. 35a-b. 36a-b

Jlolo/ ype :-DGS. Reg. No. 4546.
Description :-Shellrathersmall,
moderately inflated. oblong. inequilateral;right valve slightly larger than left
valve at ventral margin; antero·dorsalside shorter than postero-dorsal side;slightly rounded but somewhat concavein front of beak. passing gradually into

broadly rounded ventral margin: posterodorsal side somewhat rounded, passing
abruptly into ventral margin. Beaksmall, pointed. situated somewhat anteriorly. Right valve \Vith numerous.
subequal, rather coarse. low. smooth concentric lines which are narrower thantheir interspaces, well developed keel
extending from beak to postero-ventral
corner. posterior border elongated withcoarse concentric lines and ill developed
keel from beak to postero-dorsal end.
Left valve smaller than right. with surface sculpture equal to right valve except for obsolete keel from beak topostero-dorsal end.
Interior surface
smooth: hinge with small but somewhat
distinct resilial pit; muscle scars rather
distinct.

Dimensions (in mm) :Height
Length
Depth

6.5•
9.5
·l. 5**

7.5
10.2
:~. 0

5.2

4.5

2.1

8.2
2.0

6. 2

3.6

2.0
3.5

*-holotypc: ••-breadth

Remarks:- This species is distinguishable from Anisocorbula peregrina (YOKO·
YAMA) (1923. p. 55, pl. 6. figs. 1-t. 14a)from the \liocene Fujishima Formation.
Wakayama Prefecture and Anisocorbula
osawanoensis TsUDA (1960. p. 79. pl. 3.
figs. lOa-c) from the Kurosedani !vliocene.
Toyama Prefecture, in having small,rather thin shell. numerous. rather coarse
but low. smooth concentric lines andless distinct and small resi lial pit.
TyjJl' localily:-Loc. No. 36.
Occurrence :-Type Loc.- F: Loc. No.
24-F: Loc. No. 30-F: Loc. No. 32-F;

Loc. No. 33-R.
Family Haliotidae
Genus Ha/iolis LJNNAEUS, 1758
Hafiotis notoensis MASUDA, n. sp.
Pl. 36. fig. 1

Holotype :-DGS, Reg. No. ,1551,
Description :-Shell small, thin, auriform. low: whorls low. about four innumber. separated by rather indistinct
sutures; about two younger whorls

Kuichiro MilSUDA
smooth. Body whorl very large. low.
somewhat shouldered. -...vith a little morethan ten small openings on shoulder
and with somewhat elevated. curved.
coarse longitudinal threads and num·erous, unequal. rather fine spiral threads;
longitudinal threads narrower than theirinterspaces, more elevated than spiralthreads and somewhat granulated; spiralthreads at periphery under shoulder
nearly equal, smooth. much more distinct
than those on surface, about !he in
number, narrower than interspaces.
Dimensions (in mm) :-Height ca. -18.
width 14.0 x 19.5 (holotype) ; height 5.0.
width 29.0x20.0; width 22.0x 17.0 (para·type).
Remarks :-From Haliol is diuersicolor
REE\"E (1846, pl. 12. sp. 39), a Recent
species of Southern japan. the present
new species can be distinguished by its
strong longitudinal threads. fewer spiralthreads on periphery and nne spiralthreads on surface.
Type locality :-Loc. No. 24.
Occurrence:·- Type loc.-R; Loc. ~o.
10-F: Loc. No. 30-F: Loc. :-.Jo. 32-R.
Family fo'issurellidae
Genus Tugali GRAY, 1843

Tugali no/oensis ;\IASUDA. n. sp.
Pl. 36. figs. 2a-b

Holotype ·-DGS, Reg. No. 4S52.
Description :-Shell rather small. low·
conic; apex prominent, directed posteriorly. situated at about 3/4 of shell length;
anterior surface somewhat concaye;
posterior surface broadly rounded. Sur·face with numerous longitudinal ribletsand concentric lines, making reticula·tion: riblets rather low. unequal. somewhat broader than their interspaces;
concentric lines rathr conspicuous at an·

terior margin: slit fasciole a little elevated. Aperture elongate oval. tapering
posteriorly; interior margin finely denticulated.
/Jimensions (in mm):- Height 7.5,
length of aperture ca. 22.0. breadth ofaperture 13.5.
Remarks:- This new species is distinguished from Tuga/i decussatoides(NOMURA and 1-IATAI) (1936b, p. l18. pl.
17. fig. ll) from the l\Iiocene Kubota
Formation. fukushima Prefecture by
the position of its apex. broadly rounded
posterior surface and more ri blets.
Type locality :-Lac. No. 24.
Occurrence:- Type loc.-R: Loc. No.
30-R.
Family Troc.hidae
Genus Euchelus PJJILIPI'l. 1847

Euchelus noloc11sis 1viASUDA. n. sp.
Pl. 3G. figs. ·1a··c. 5a-c. Ga-c

l!olotype :-DGS, Reg. No. 4554.
/Jescription :-Shell rather small to
medium in size. turbinate conical. withfour post nuclear whorls and one and ahalf nuclear whorls: spiral angle about90". ·whorls rounded. shouldered, separated by distinctly channeled sutures:
nuclear whorls smooth. rounded; surface
with several, conspicuous. fine granulated. unequal spiral threads and somewhat oblique, fine longitudinal threads;
two spiral threads on first post nuclear
whorl. tend to increase their number byintercalation downwards. nearly equal
to their interspaces: interspaces withoblique. fine threads.
Base of bodywhorl with fine, distinct. finely granulated. nearly equal spiral c01·ds which
are subequal to their interspaees. Aperture rather wide. subo\·al; outer lip
rather thick, denticulated within; inner512.

.l!ollHscan Fa11nn of the Hiua.sh·i-lnna.i.

lip sharply defined; umbilicus rather
narrow. deep. crack-like; columela near-:131

ly straight. subvertical.

Dinu>nsions (in mrn) :Height
Diameter

5. 8•
7. 0

ca. 9. 6
ca. 10. 0

5.0
G.O

4.3

.t.O

~.4

4.8

3.-t
·1. 0

*-holotype

Remarks:- The present species is dis·tinguished from Euc!udus ornatissimus
YOKOYAMA (1926b, p. 374. pl. 43. figs. 7.
8) a Pleistocene species of Atsumi Peninsula. Aichi Prefecture by its rather low
whorl and distinct. deep umbilicus.
Euchclus minocnsis ITOJc;AwA (1960. p.
275, pl. 3. fig. 10) from the l\liocene
Mizunami Group. Gifu Prefecture resembles the present new species but differsin having rather higher whorls sculptured with rather less prominent granulated spirals. three post nuclear whorls
and no dentation within the outer lip.
Type /ocality:-Loc. No. 24.
Ocwllence:-Type loc.-F; Loc. No.
30-F; Loc. No. 35-R.

thin: columella oblique.
Dimensions (in mm) :-Height 4.6. diameter 3 (holotype).
Remarks:- This new species resembles
Calliostoma lwrodai IKFBE (19~2. p. 275.
pi 28 figs. 13a-c). a species living inToyama Bay. japan, but kurodai is distinguishable from otaniensis in having
larger shell, a little more spiral angle
and less distinct shoulders.
Type locality :-Loc. No. 30.
Ocwrrence :-Type loc.-R.

Genus Cal/iostoma SWAINSON, 1840

Hololype :-DGS, Reg. No. 4556.
Description :-Shell medium in size.
conical. with two nuclear and four post
nuclear whorls; spiral angle about 90".
Whorls moderately high. rounded, separated by somewhat channeled sutures:
nuclear whorls low. smooth. rounded:surface with several. unequal. rather
low. more or less lamellated. smoothspiral threads and oblique. faint. fineincremental lines: spiral threads three
on first post nuclear whorl and increase
their number by intercalation towards
base. Base of body whorl moderately
rounded. with several (about seven), low.
flat-topped. smooth spiral cords which
are much broader than their interspaces.
Aperture subovate, oblique; outer lip
thin. simple: inner lip rather thick withweak denticles; umbilicus medium,

Calliostoma otaniensis l\1ASUDA. n. sp.
Pl. 36. figs. 7a-b

Holotype :-DGS, Reg. No. 1406.
Description :-Shell small, conical. with
about six whorls of which one and ahalf are nuclear whorls; spiral angle
about 55<. Whorls shouldered. separated
by distinct. impressed sutures: sculptured with rather strong. granulated
spiral threads which are broader thantheir interspaces; three primary and one
secondary spiral threads in penultimate
whorl and three primary and two secondary spiral threads on body whorl.
Rase of body whorl \Vith about nine.
fine, smooth spiral threads. Aperture
subquadrate: outer lip simple; inner lip

Genus ,\lonilea Swi\11\SO:'\, 1&.10
Alonilea yoshioi 1\IASUDA, n. sp.
Pl. 36. figs. Sa-c. 9a-c. lOa-b

332

f{l.ichiro AI ASUDA

rather deep : columella oblique.

Dimensions (in mm) :Height
Diameter

\J.O*
11. s

9.ii

9.0

8.0

7.5

10.5

10.5

10.0

9.0

7.0

7.2
7.6

8. ·I

*-holotype

Renwrks :-Monilea ringulata I\·lAKI·
YAl\IA (1927. p. 60, pl. 3. figs. 3, '1) fromthe Pliocene Oainichi Formation. Shizuoka Prefecture resembles the present
new species, but it can be distinguishedfrom yosllloi by its larger shell, larger
number of post nuclear whorls andlarger number of spiral threads. Homa·
lopoma hidaensis lTOIGAWA (1960, p. 277,
pl. 3. figs. 12a-b) and Homalopoma tulii·
yosiensis Qy MIA and SAKA (1944, p. 138.
pl. U. figs. 1a-c), both from the 1\liocene!vlizunami Group. Gifu Prefecture also
resemble the present species, but bothdiffer by the shape and height of thespire. strength of spiral threads andaspect of the umbilicus.
The specific name is given to the nameof my son, Yoshio ~·IASt:PA who wasborn recently.
Type locality :-Loc. No. 39.
Occurrence:- Type loc.-A; Loc. No.
30-C.

Monilea hamadae l\IASUPA n. sp.
Pl. ~6. figL lla-c, 12a-c. 13a-b. 14a-c

Jfolotype :-DGS. Reg. No. '1558.
Description :-Shell rather small, rather
thin. conical. with two nuclear and four
post nuclear whorls ; spiral angle a little
less than right angle. Whorls shoulder·eel. somewhat rounded. separated by dis·tinctly channeled sutures; nuclear whorls
low. smooth. rounded : surface with two,strong. elevated spiral threads, onerather weak subsutural thread and faint,
fine. oblique longitudinal threads, rarely
with a few. fine intercalary threads:
spiral threads much narrower than theirinterspaces. Base of body whorl round·ed. with about seven, rather low. smoothspiral cords which are nearly equal totheir intcrspaces. Aperture subovate:outer lip thin: umbilicus very wide, deep.
with fine spiral cords.

!Jimensions (in mm) :Height
Diameter

5.0*
5.2

5. 8
5.8

5.~

5.4

4.8
·t 8

4.5

4.8

4.0
4.2

*-holotype

Remarks:- The present new species isdistinguished from Monilea yoshioi ~IA ·SUIJA described in this article by its
smaller and somewhat higher shell.
shouldered whorls. t\VO conspicuous
spiral threads and \·ery wide umbilicus.
Homalopoma tukiyosiensis OYAMA and
SAKA (Op. cit.) resembles the present
one, but differs from the present one byits larger and higher shell. ftattish

whorls. shallow sutures and larger number of spiral threads. Monilea (Rossi·
feria) osawanoensis TsUDA (1959, p. 81.
pl. 3. figs. 15a-b) from the Kurosedani
I\liocene of Toyama Prefecture also re·sembles the present one. but it is dis·tinguishable from lwmadae by its higher
shell, slightly shouldered whorls. surface
sculpture and characters of inner lip.
Type locality:- Loc. No. 23.

512.

Mollwwan Fauna o.f the J-li[laslti:-Innai

Occurre11ce :-Type Joc.-C: Lac. No. 3-C: Lac. No. 30-I~.

Family Turbinidae
Genus Turbo LINNAEUS. 1758
Subgenus i\larmorosloma
S\\'AI!\SON. 1829
Turbo (Marmorostoma) ozazl'ai

0TUI\A, 1938
1938. Turbo (Lunel/a) ozau·ai On KA. jour.
Fac. Sci .. Imp. Univ. Tokyo, Sec. 2, Vol.
5. Pt. 2. p. 36. pl. 3. figs. 22-24.
1959. Turbo (Marmorostoma) cf. ticaonica
REic\'E, lloRJKOSFII in KoBAYASHI andHoRJKOSIII. Japan. jour. Ceo{. Geogr ..
Vol. 29, Nos. 1-3. p. 49, pl. •1. figs. la,b. text-fig. b.

Syntype :-GT*, Reg. No. 10018.

Remarks:- This species was first de·scribed by OTUKA from the Miocene Sh6·
bara Formation, Hiroshima Prefecture
based upon two opercula, thus the shellcharacters remained unknown. In 1958
llotUKOSHI illustrated Turbo (Marmoros·lama) cf. ticaonica REEVE. a Recent
species of the Philippine, from the l'vlio·
cene Uchiura Formation. Fukui Prefec·
ture based upon a single shell without
the operculum. At that time he statedthat from the characters of the oper·
culum 0TUKA's ozawru should be referr·
ed to the subgenus Marmorostoma and" if ozawai is found together with ashell of Turbo cf. ticaonica at any fossil
locality. OTUKA's denomination is vali·
dated for Turbo cf. ticaonica ".
The abundant shells and operculafrom various localities in the present
field show that the former can be referr·
ed to the HORIKOSHI's Turbo cf. ticaonica* Abbreviation for the Geological Institute.
University of Tokyo.

•>•)0t)t_hJand the latter to 0TUKA's ozawai.
The shells and opercula from the Hi·
gashi-lnnai Formation take the following
description: Shell medium. thick, ovate·
conical; whorls six of which two are
nuclear ones: spiral angle about 70'.
Nuclear whorls smooth. rounded; post
nuclear whorls moderately rounded, moreor less shouldered. separated by im·pressed sutures; surface with several,elevated, strong. unequal, more or lessrounded spiral cords, fine interstitial
threads and fine but rather distinct in·cremental Jines: first post nuclear whorlwith three. unequal, strong spiral cords
which are much narrower than theirinterspaces and three, weak. fine spiralthreads. Body whorl with six, strong,unequal spiral cords, a fine interstitial
spiral thread between strong spiral cords
and rather distinct oblique incremental
lines: spiral cords forming shoulder andoccasionally more or less granulated at
upper part of whorl; base rounded. withseven to eight, strong, more or lessgranulated, unequal revolving threads
and rather distinct incremental lines.
Aperture subovate; outer lip rather
thin: inner lip rather sharply definedby thin callus; columella somewhat con·cave. Operculum slightly elongate-oval.
much inflated, asymmetrical in profile:base slightly inflated. Surface rounded,with numerous, fine, unequal, rounded
tubercles: fine tubercles sometimes con·nected with each other and with irregularly granulated lines subparallel withbase. Base more or less undulated byrather distinct coiling, with faint, fineincremental lines and very faint, fineradial threads which can be observed
only by lens.
Turbo (A!armorostoma) licaonica REEVE(1848, pl. 5. sp. 23) is distinguished fromthe present one by its larger, umbilicatedshell and larger number of spiral threads.
REEVE

J{iJi.chiru MASUDA
The present species also resembles Turbo
parvuloides NoMURA (19,10, p. 35. pl. 3.
figs. 2a-b) from the ~liocene \loniwa
l'vlember of the Hatatate Formation. 1\liyagi Prefecture and also Turbo (Mannoros/oma) minoensis lTOlGAWA (1960. p.
278. pl. I. figs. 2. :3) from the l'vliocene~·lizunami Group of Gifu Pr.:fecture.
llowever. it differs therefrom in having
larger shell and fewer coarse spiralthreads.
The shells from the Higashi-lnnai Formation show considerable variaton inmorphological features. though the opercula are similar with one another except
for the surface tubercles. That is tosay. the specimens from Loc. Nos. 24. :10
and 32 have somewhat higher shell.
somewhat fewer interstitial spiral threads
and coarse. smooth spiral cords but with

Explanation of Plate 35(All figures in natural size. unless otherwise stated). Figs. la-b. 2. 3. Saccella saikaiensis :\IASIJD.-\. n. sp. la-b. holotype. DGS. Hcg. No. 4494:
Loc. No. 23. x 2. 2. 3. paratype. DGS. Reg. No. 4495: Loc. No. 3. x 2............... p. 322
Figs. I, 5. 6. Aequipecten matsunagiensis MAstmA. n. sp. ·1. holotype. DGS. Reg. No. 4506:
Loc. No. 32. 5. paratyp:>, D~S. Reg. ~o. ·1655: Loc. No.3. 6. paratype, DGS. Reg. No.
4507 : Loc. No. 32. . . . . . . . . . . . . . . . . . . . . . . .
. .................................. p. 323
Fig. 7. Placopecten protnmnllitus (Nm!!iRA). DGS. !{eg. No. 4509: Loc. No. 32.
Figs. 8a-h. 9. Modiolus ll'flllizllkicusis lVL\suL>.\, n. sp. Ba·-b. holotype, DGS. J~eg. 'No. 4510:
Loc. No. 31. 9. para type. IJGS. Reg. No. ·1511 : Lm:. No. 31. ........................ p. 32,1
Figs. 10. 11. Myadora suzuensis MAsUDA. n. sp. 10, paratype. DGS. Reg· No. 151-1: Loc. No.
21. x2. ll. holotype. DGS. Reg. No. -!513: Loc. No. 21. x2 ......................... p. 324
Fig. 12. Crassatellites suyamensis Ot:-.o~!!KADO. DGS. Reg. No. 1415: Loc. No. 30.
Figs. 13a-b. 14. Glans naomiae MASUDA. n. sp. 13a-b. holotype. DGS. Reg. No. 4517: Loc.
No. 35. x2. 14. paratype. DGS. Reg. No. 4518; Loc. No. 23. x2 ................... p. 325
Figs. 15-20. Ctena hataii MAstll>A. n. sp. 15. holotypc. DGS. Reg. No. '1521: Loc. No. 30. x 2
16, 17, 19. p:uatype, DGS. Reg. No. ·1523: Loc. No. 30. x2. 18. 20. paratype, DGS. Reg.
No. -1525: Loc. No. 24. x 2. . ....................................................... p. 325
Fig. 21. Siratoria siratoriensis (OTUI\A). DGS. Reg. No. •1535: Loc. No. 3.
Figs. 22. 23. J'aphia euglypta ohiroi M:\SUDA. n. subsp. 22. holotypc. DGS. Reg. No. •1527:
Loc. No. 38. 23. paratypc. DGS. Reg. No. 4523: Loc. No. 39......................... p. 32()
Figs. 2t 23. Paphia suwensis MASUDA. n. sp. 21. parat)·pe. DGS. Reg. No. 4529: Loc. No. 30.
25. holotype. DGS. Reg. No. ·1530: Loc. No. 30..................................... p. 327
Figs. 26. 27a-b. Leukoma itoigawae TsuDA. DGS. Reg. No. ·1532: Loc. No. 30. x 2.
Figs. 28a-b. 29. Venerupis (Amygdala) takagii (MAsUDA). DGS. Reg. No. -1537: Loc. No. 35.
... ...... .... .. .... ...... .. .... ..
. ........................................ p. 328
Fig. 30. Clementia japonica MASUDA. DGS. Reg. No. •15·10: Loc. No. 23.
Fig. 31. Pilar iloi (MAKIYA~·IA). OGS. Reg. No. •1536; Lnc. No. 23.
Figs. 32. "Venus" sp. DGS. Reg. No. -15•15: Loc. No. 24.
Fig. 33. Cyclina (Cyclinorbis) lunulata 1\Lo\KlYA~IA. DGS. Reg. No. 4538: Loc. No. 23 ... p. 326
Fig5. 34a-b. .4nisocorbula peregrina (YOKOYA:'>.tA). DGS. Reg. ?\o. 4549; Loc. No. 24. x2.
Fig5. 35a-c. 36a-b. Anisocorbula ohiroi :\hsuoA, n. sp. 35a-c. holotype. DGS. Reg. No.
45·16: Loc. No. 2·1. x 2. 36a-b, paratype. DGS, Reg. No. 4547 : Loc. :-\o. 30. x 2. . ... p. 329
Fig. 37. Raeta sp. DGS. Reg. No. 4541: Loc. No. 12.
Figs. 38a-b. Cardilia toyamae11sis TsuDA. DGS. Reg. No. 4543: Loc. No. 35.

K. MASUDA:

Higashi-Innai Molluscan Fat~na

Plate 35
3

5

8b

10

•

;;u.

:135

.lfollusca.n Fauna of the Hiqashi-1-nnaino distinct tubercles comp3.red with theindividuals from Loc. l'\o. 35. ll'o andl<A:\IETAKA (1943) state that the development of tubercles or spines in Turbo isrelated with the environmental conditions. that is to say. the shells with illdeveloped spines are usually found inrather calm water and those with welldeveloped spines in the influence ofrough water conditions. Therefore. it

seems that from the development of th~
tubercles in ozawai from the Higashi·
lnnai Formation the eJ1\:ironmental con·ditions in the vicinity of Loc. No. 35
may haw been of rocky bottom influenced by strong waves and currents.
This yiew is supported by the hardbottom dwelling molluscan assemblage.
reef building corals. calcareous algae.
etc. and also by the lithologic facies.

Dimensions (in mml :Shell :HeightDiameter

36.5
31.0

35.0
32.0

3-1. 0Height

7.6

/.0

6.5

Maximum

1/. :{

lf\.:1

16. ·I:l2. 0

30.0

13.0

24.5

10.5

')

5.0

3. 5

lu.o

12.8

~). 2

32.0
26.0

Operculum:clianwtcr

Type locality:-Suketo. Sh6bara-machi.
Higa-gun. Hiroshima Prefecture.
Occurrence :-Loc. ~o. 1-F (shell,::Loc. No. 3-F (shell): Loc. No. 10-F
(shell and operculum) : Loc. No. 2-!. operculum-A. shell-F: Loc. No. 30-F
(shell and operculum) : Loc. No. 32. shell-C. operculum-A: Loc. No. 33-F (oper·
culum) : Loc. No. 35-A (shell and operculum).
Family Neritidae
Genus Nerita LI~N AEUS. 1758

Nerita ishidae l\IASUDA. n. sp.

'·"'

Pl. ~6. fig~. 23n, b. 24n-c. 25

Holotype :~DGS. Reg. No. c156-L
Descriptioll :-Shell of medium size.
thick, semi-globose: whorls minute: surface smooth but with faint. fine incre·
mental lines. Body whorl very large.
ventricose, with regularly rounded peri·
phery. Aperture crescent form : outer
lip sharply angulated. regularly rounded
anteriorly. somewhat projected below.
rather thin. thickened inwards and denti·
culatecl on inner margin: inner lip distinct, finely denticulated; callus thick.
makes wide. flat smooth platform andinclined inwards.

Dimensions (in mm) :Height
Diameter

10.5*
l·t. 0

18.5
21. 0

13.5

9.0

8.5

li. 5

11. 0

10. 5

4.5

*-holotype

Remarks :-Nerita lwmigiriensis lTOI·(1960. p. 140. pl. 6. figs. U, 15)from the l'vlizunami 1\liocene of Gifu
Prefecture is distinguished from the

GAW A

present new species by its several. smalL
irregular granules on the columella area.
Nerita subgranulosa ~AGAO (1928. p. 117.
pl. 12. figs. 4. 4a. pl. 19. figs. 14. l4a) from

336

Koich1ru MASUDA

the Eocene Shira taka Formation of Northern Kyushu. Kumamoto Prefecture differsfrom the present one by its surface
sculpture, angulated shoulder and small
granules on the flattened inner lip.
The denticulation of the inner marginof the outer lip can be seen in rather
large specimens but not in the younger
specimens.
This species is named in honor of Dr.
Shiro ISIIII>A of the Geological and ~Iin­eralogical Institute of Kyoto University,
who collaborated with the \\!riter in thefield.

Type locality :-Loc. No. 30.
Occurrence:- Type loc.-A; Loc. No.
1-F; Loc. No. 10-F; Loc. No. 23-R;
Loc. No. 24-R; Loc. No. 32-F; Loc.
No. 33-F; Loc. No. 35-R; Loc. No. 39-F.

Family Lacunidae
Genus Lacuna TURTON, 1827

Lacuna japonica rviASUDA, n. sp.
Pl. 36. figs. 26a-b, 27

Explanation of Plate 36
Fig. 1. llaliotis notoensis MASL'DA, n. sp. Holotype, DGS, Reg. No. 4551: Loc. No. 24. x2 ... p. 329
Figs. 2a-b. Tugali notoensis MAst;DA, n. sp. Holotype, DGS, Reg. No. 4552: Loc. No.
24. X 1. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
. ............ p. 330
Fig. 3. Col/isel/a sp. DGS, Reg. No. ·1553; Loc. No. 3. x 2.
Figs. 4a-c, 5a-c, 6a-c. Euche/us notoensis MAsllr>A, n. sp. 'la-c, 6a-c. paratypc, DGS,
Reg. No. 4555; Loc. No. 24. x 2.5. 5a-c. holotype, DGS, Reg. No. 4554 ; Loc. No. 24.
x2.5 ............................................................................... p. 330
Figs. 7a-b. Calliostoma otaniensis MAsUDA, n. sp. Ilolotype, DGS, Reg. No. 1406; Loc. No.
30. X 5. . ........................................................................... p. 331
Figs. Sa-c, 9a-c. lOa-b. Jfonilea yoshioi MASUDA, n. sp. Sa-c, holotype, DGS, Reg. No.
4556; Loc. No. 39. x 2. 9a-c, lOa-b. paratype. DGS. Reg. No. 4557: Loc. No. 39. x 2... p. 331
Figs. lla-c, 12a·-c, 13a-b, 14a-c. Monilea hamadac MAst;uA, n. sp. lla-c. holotype, DGS.
Reg. No. ·1558: Loc. No. 3. x 2.5. 12a-c. 13a-b. 14a-c, para type. DGS. Reg. No. 4559:
Loc· No. 3. x 2.5. . ................................................................. p. :332
Figs. 15a-b. So/ariella sp. DGS, Reg. No. 4560; Lac. No. I. xI.
Figs. 16a-b, 17a-b. Protorotella shukuborensis ITOIGA\1' A. DGS, Reg. No. 4561; Loc. No.
3. x2.
Figs. 18a-b, 19a-b, 20a-b, 21a-c, 22a-c. Turbo (Afarmorostoma) ozawai OTUK.'\. 18a-b,
20a-b, 21a-c. 22a-c. DGS. Reg. No. 4562 : Loc. No. 35. x 1. 19a-b, DGS, Reg. No. 4563 ;Loc. No. 32. x 1 .................................................................... p. 333
Figs. 23a-b, 24a-c. 25. Nerita ishidae MAsUDA, n. sp. 23a-b, holotype. DGS, Reg. No.
456·1 : Loc. No. 32. x 2. 2·la-c, para type. DGS, Reg. No. 1648: Loc. No. 30. x 1. 25,
para type. DGS, Reg. No. 4565; Loc. No. 10. x 2..................................... p. 335
Figs. 26a-b, 27. Lacuna japonica MASUDA. n. sp. 26a-b, holotype. DGS, Reg. No. 4566:
Loc. No. 30. x 5. 27, para type, DGS, Reg. No. 4567; Loc. No. 30. x 5............... p. 336
Figs. 28a-b, 29a-b, 30a-b. Littorinopsis miodelicatula ()y AMA. 28a-b, DGS, Reg. No. <1568:
Loc. No. 24. x2. 29a-b. 30a-b, DGS, Reg. No. 4569; Loc. No. 32. x2.
Fig. 31. Pseudoliotia sp. DGS, Reg. No. 4570; Lac. No. 35. x 5.
Figs. 32a-b, 33a-b, 34a-b. Rissoina naomiae MAsUDA. 33a-b. holotype. DGS, Reg. No.
4571 ; Loc. No. 24. x 5. 32a--b. 34a-b, para type. DGS, Reg. No. 30. x 5. . ........... p. 337
Figs. 35a-c. Architectonica kuroclae TsUDA. DGS, Reg. No. 4572; Loc. No. 2<L x 1.

K. MA SUDA:

Higashi-Innai Molluscan Fauna
3'.Ia

Plate 36
5a

5b

5c

D12.

Molluscan Fauna of Uw Hiuashi-lnnm:

Holotype :-DGS. Reg. No. 4566.
Description :-Shell very small. moder·ately thick. globose. with about fivewhorls; spiral angle about go·. Whorls
roundly shouldered. separated by rather
distinct sutures; surface smooth. Aperture wide. ovate; outer lip thick. rounded; inner lip with thin callus; umbilicus
rather wide, deep and oblong.
Dimensions (in mm) :-Height 3.2. diameter 3.0 (holotype); height 2 2. diameter
2.0 (paratype).
Remarks :-This new species is distinguished from Lacuna i·hayasakai No~IURA (1938, p. 272. pl. 33. figs. •la-b)
from the Pliocene Tatsunokuchi Formation, ~liyagi Prefecture by its larger
spiral angle and rounded, thick outer lip.
Type locahly:-Loc. No. 30.
Occurrence:-Type loc.-F.
Family Rissoidae
Genus Rissoina D'OJWIG:-;Y, 1840
Rissoina naomiae MASUDA, n. sp.
Pl. 36. fig. 32a, b. 33a, b, 34a, b

l/olotype :·- DGS, Reg. No. ·1571.
De~cription :-Shell small, rather thick.
elongate-conical, imperforate, with sevenwhorls. two nuclear whorls lacl<ing:spiral angle about 300. Whorls moderately inflated. separated by somewhat
channeled sutures; sculptured with distinct longitudinal ribs and numerous
spiral threads: longitudinal ribs subYertical, rather strong, more or lessrounded, extending from summit tolower suture, about 25 on body whorl.
about 21 on penultimate whorl and tend
to decrease their number towards upper
whorls: last longitudinal rib makes aprominent. rounded varix on back ofouter lip; interspaces between longitudinal ribs broader than ribs themselves; spiral threads usually faint andfine but tend to become distinct towards
lower part of body whorl. Body whorl
contracted below : base rounded, with astrong, rounded. longitudinal costellated
spiral cord. Aperture subovate. withrather acute angle posteriorly; outer lip
prominent, marginate. anteriorly tortile:
inner lip smooth, with rather thick callus.
slightly concave.

Dimensions (in mm) :Height
Diameter

6. O•

7.5

6.8

2. 6

:{. 0

')

-·"
~

6.5
3.0

5. 2
2. ;{

5.0
2.2

•-holotype

Remarks:-The present new species
resembles Rissoina rosea ( DESIIAYESt(SOWERBY. 1878. pl. 5, sp. 42). a Recent
species of Southern japan and Rissoina
submerculialis YOKOYAl\IA (1920, p. 73,
pl. '1. fig. 15). a I' leistocene species ofCentral japan. but it differs from theformer by its numerous spiral threads

and no columella plait and from thelatter by its large and long shell. larger
number of subvertical longitudinal ribs
and rather thick callus.
Type locality:- Loc. No. 30.
Occurrence:-Type loc.-A; Loc. No.
1-F; Loc. No. 10-C: Loc. No. 2·1-F:
Loc. No. 35-F.

Trans. Proc. f>alaeont. Soc. Japan. N. S., No. 6:1. pp. 338-350, pl. 31, Dec. 15. 1966

51:1.

FUSULINIDS FR0;\1 THE FUNAFUSEYAl\IA LIMESTONEIN YAMAGATA-GUN. GIFU PREFECTURE. JAPANKUNITERU ~v!ATSU.\1!\RU
InstituLe of Geology and Paleontology, Facully of Science,
Tohoku Uniyersily, Sendai. Japan

llttU,'l.~~LIJlUll~ltHJ\Ufl.iY;'fiiJ ~o);j-Ji~!h:Jlt(!,;llrFilk:~~:·n ·!:?ffiH\-t MJi~!l~it.11fJ: 12/j:f.

43 H!n: ~IE~~·· .=tO)') -t.., llif!il>f!iFf!'t:' t, :S.

..:. :h l?([JMi~:l!:l!.fUil~n.. tJ:O) Paraschwagerilw (.-lcen•oschwagerina) illi1f.'~·~J.:ti0) }"abeina igoi !lli:;i;;t·::>;t'\U;IJ~:lt~•, :<(~~O)~:Ji~!l'.ft.:.ri

c i'iwi. w:;tU. fJ<:Y:tlJ<!J.:tttco);iJJt~iVP?-t..:..

r~ :kllil !!!!

~-~----·

Introduction~It. Funafuseyama (10·13.3 m) and its
adjacent area has been studied by manygeologists and palentologists, namely
liSAK A (1932). KonA YASlll (1951) leo and
OGAWA (1958). IsHII and his collaborators (1963, 1964), NAKA1viURA (1965j. andothers. llowever, most of the studies on
the fusulinids from this area were thedescriptions of new· and interesting
species from only several restricted localities. The Funafuseyama Limestone
is distributed about l2 km in east-west
and about 9 krn in north-south direction
in Yamagata-Gun. Gifu Prefecture.
From the extensive distribution andgood development as well as yield of
fusulinids. the limestone of this rtrea isconsidered important for establishmentof a standard biostratigraphic sequenceof the fusulinids and for a basic column
for interregional correlation.
Fortunately the writer at the adviceof NAK1UICRA \vho worked out the• Rccch·cd l\larch 19. 1966: Read June 18.
1966.

geology of the limestone was able tocollect to abundant fusulinids. Basedupon the fusulinids and the geology ofthe limestone. which may serve as astandard. the writer in this article at·
tempts interregional correlation.
Acknowledgements
The writer takes this opportunity to
express his hearty thanks to; Professorl<iyoshi AsA:\0 of the Institute of Ge·
ology and Paleontology, Faculty of
Science, Tohoku University. for his encouragement, Professor Kotora I-lATA!
of the same Institute, for his encouragement and reading of the manuscript,
Professor Rokuro MOI~IKAWA of the
Department of Earth Science. Saitama
University. for his suggestion on theclassification and zoning of the fusu·
linids, Drs. i\·lasafumi ;'viURATA andKunihiro ISHIZAI\1. both of the Instituteof Geology and Paleontology, Tohoku
University. for their kind discussions on
the fusulinids, Dr. Manjiro NAKA:\IlJIL\of the Kokan !\lining Co. Ltd., for hiskind offer of the limestone blocks fromthe Funafuseyama area and valuable
information on the stratigraphy.

Stratigraphic notes
According to NAKAMUI~A (1965j. therocks of the Funafuseyama area can beclassified into two major units. One is
the non-calcareous group and the otheris the calcareous group. The former is
subdivided into the Kanzaki and Tsubaki lithogenetic units. The Kanzaki
Formation is composed mainly of chert
intercalated with schalstein and limestone lenses. The Tsubaki Formation iscomposed mainly of sandstone, shaleand chert intercalated with schalstein
and limestone lenses.
The Funafuseyama Limestone is composed mainly of grayish white, gray,dark gray and black limestones anddolomite intercalated with schalstein.
The Kanzaki Formation of the non-cal·careous group overlies the Tsubaki
Formation in the western part of thearea with conformity but is superposedon the formation in the eastern part. inother words. they interfinger with one
another. The Funafuseyama Limestoneof the calcareous group lies upon the
Kanzaki Formation with conformity.
The succession of the strata from theolder to the younger in the non-calcareous group is from sandstone and shale
to chert in the western part and fromsandstone to shale in the eastern part.
In the calcareous group the lowest
observable strata is sandstone. and thisgrades upwards into shale through chert
to limestone. The non-calcareous groupand the calcareous group are stratigraphically contemporaneous but in different lithofacies according to NAK.'\C\!Uf{A( \96;)). The relation can also be upheldby paleontological evidence of the mutual

occurrence of fusulinids as. :Veoschwagerina margaritae. N. sp. and }'abeinaigoi, from the limestones of both groups.
On the other hand. !Go and OGAWA\1958) once stated that the stratigraphic
relationship between the Funafuseyama
Li!l1estone and the Kanzal<i Formation
is doubtful. but the larger part of theformations are contemporaneous with
one another and the lower part of thelatter may be older than that of theformer.
Paleontological remarks

1. Zoning of the Funafuseyama Limestone
The Funafuseyama Limestone can beclassilied into four zones and five subzones based upon the respective chronological ranges of the fusulinids collectedfrom the limestones of the present area.
The zones and subzones recognized in
upward sequence in the Funafuseyama
Limestone are:Pseudosc/m:agerina zone
Paraschwagai11a(Acen•oschwagerina)
subzoneParafusulina zone
Neoschwagerina zone
Neoscilll'agerina fuwensis subzone
N1!osc/ucageri110 co/aniae-rotunda subzone
N eoschtcageri rw margaritae-craticulifera
suhzone1'abeiua zone}'aiJeiua igoi subzone

In spite of the occurrence of fusulinid
species from most of the localities ofths funafuseyama Limestone, particularspecies characterizing the Parafusulina
zone were not recognized. It is. however,expected that the Parafusulina japonica
subzone can be separated from theParafusulina zonl~ by future detailed stratigraphic work and refined study of the
Parafusulina species in this area.

340]{unitent, MIITSUAiARU

From the above cited fusulinid zonesof the Funafuseyama Limestone, thetime range of this limestone is considered to be longer than hitherto believed
as once suggested by !Go and OGAWA( 1958).
2.
1)

Description of each zones
Paraschwagerina (Acervoschwa}!erina)
subzoneThis subzone is distributed mainly in
the Hatsushikadani area (refer to locality
map for all localities and locality numbers) and is characterized by the occurrences of Paraschwagarina ( Acervoschwa}!erina*) sp., Pseudofusulina lzraffti, Psf sp,, Yangchienia sp., Codonofusiella sp., and Schubertel/a sp.
This assemblage referrs them to the1/ seudoschwagerina
zone.
This fossil
zone is relatively poor in fossil yield
just as the Paraschwagerina (...lcervoschwagerina) subzone of the lbukiyama
Limestone, Gifu Prefecture, according to
KoBAY ASIII (1957). The locality numbers
are indicated in the locality map, together with the geographical or local
names of the area in which they occur.
Locality names
Locality numbers
Hatsushikadani .... , .... .401. 185. 391, 395
Kanzaki ......... , ........ 71

2)

Parafusuli 11 a zone
This fossil zone includes the Kanzaki
and Tsubaki Formations of the noncalcareous group and the Funafuseyama
Limestone of the calcareous group. The
outstanding species of this zone are
Parafusulina japonica, P. mizutanii.
Pseudodoliolina ozawai and Neosclwagerina ni pponica.
This fusulinid fauna is similar to the* The writer considers l he Acerlloschu:agerina subzone of KoHAYA:;lll (19S7) to beequivalent to the Parasclru·agerina (Acervoscluvagerina) subzone.

fusulinids of the Parafusulina japonica,
Pseudodoliolina ozawai and Neoschwagerina nipponica zones of the Akasaka
Limestone. Gifu Prefecture. Those zones
are characterized by the occurrences ofParafusulina Japonica, Pseudodoliolina
ozawai and Neoschwagerina nipponica,
respectively, as already pointed out by
MORIKAWA (1958), The following species
were discriminated from the Parafusutina zone of the Funafuseyama Limestone:
Yangcl!ienia sp,, Codonofusiella sp,, Sclrubertella sp., Parafusulina exilis, P. gifuensis,
P. frmafusensis. P. japonica. P. gigantojaponica, P, kinosakii, p, takeyamai. P. okubol'nsis. P. taniyaslrilliensis, P. mizutanii, P.
trrmcata, P. tomeganensis. P. o;:azcai. P. yabei,P, kinslroensis. P. parakinosallii. P, undata, P.
neoensis, P. takeii, P, gumbdi, P. sp .. Clrusenella douvillei, I' erbeekina sp., Misellina
claudiae, Pseudodoliolina ozawai, Neoschwagerina nipponica, N. paraniP/IOnica.

The fusulinids from the Parafusulina
zone are abundant both in individuals
and number of species.
Chusenella
douvillei from this zone of the Funafuseyama Limestone closely resembles
the same species recorded from thel'vlaokou Limestone, Kwangsi, China by
SHENG (1963). Ch usenella douFillei from
the Maokou Limestone is stated to occur
in association with Schwagerina jJseudocompac/a, S. quasibrevipola, S quasiregularis. S pingdingensis, Chusenella conicocylindrica. Pseudodoliolina pulchra and
Neosclzwagerina kn·angsiana. This fusulinid assemblage belongs to the Neoschwagerina zone of SHENG. Therefore,
it is very interesting that Chusenella
dozwillei from the Funafuseyama Limestone occurred from the Parafusulina zone
and not from the Neoschwagerina zone
as in China. From the occurrence fromthe Parafusulina zone Chusenella must
be considered to range from the Para-

:l·llfusulina zone up to the Neosclllcagerinazone.
The localities are as follows.
Locality names
Locality numbers
Funafuscyama .... 10, 14, 15
Nakagoshi ........ 416, 173.420, 415.88. 91. 92
Mizudc ............ 93, 410.413,56. 60.57
Fukase ........... .'15, 52. 118. 110, 1}(ashibaradani .... 69.63.121.107,189.191,
192,193.414
Iwatani .......... 126, 164. 162.92, 156.98,136
Nakagashi ........ 132. H2Shiraiwa .......... 139
Oiwake ............ 30, 10,1. 6. 3. 26. 5, 25, 2~
Mahira ............ 220, 221.222, 21
Takaradani ........ 381
Enbara ............ 197. 82
Kanzaki... 322
Akadaru .......... 2D7. :~51

Neoschwagerina fuwensis subzone
Neoschwageaina fuu:ensis is the characteristic species of this subzone and isassociated with rangchzenia sp., Codonofusiella sp.. Parafusulina exilis, P. japonica, P. kinosakii. P. taniyashikier1.~is,Pseudodoliolina ozawai, Neoschwagerinanipponica and other fusulinids. Thesefusulinids first appear in the upper partof the Parafusulina zone and extend
into the lower part of the Neoschwagerinazone. For such reason this subzone may
better be placed in the ParafusulinaNeoschwagerina zone as already pointed
out by MoRIKAWA (l960). However. inorder to emphasize the importance of
Neoschu.wgerina fuzl'ensis as a stratigraphic key and paleontological horizon,
the writer proposes this subzone.
The localities are as follows.
3)

Locality names
Locality numbers
Nakagashi ...................... 421. 90.91
Oiwake .......................... 27
Enbara .......................... 326. 22·I) Neoschwagerina colaniae·rolunda subzone

This subzone is characterized by Neo·schwagerina as Neoschwagerina colaniae
and N. rotunda. Those two characteris·tic species \Vere collected in association
with Vangchienia sp .. Parafusulina exi/is,
P. japo111ca. P. gigantoJaponica. P. gifu·
ensis. P. kinosaltii. P. takeyamai. P. tani·
yashikiensis. P. llinshoensis, Verbeekina
sp .. Pseudodoliolina ozau·ai. Neoschwage·
rina ni pponica, N. akasakensis. f'.:. t uwe•lsis. N. colaniae. N. rotunda, and N. sp.
from the Localities given below.
.Veoschwagerina colaniae and N. rotunda are restricted in chronological distribution to this subzone and occurred
only from the Limestone. However.
KAMMERA (1961) reported that Neosch·wagerina rotunda is a member of the:\'eosl'hwagerina craticulifera zone. which
occurs in the Kozaki Formation at
Shimofukami, in the Kuma massif.
Kumamoto Prefecture.
The localities are as follows.
Locality names
Locality numbers
Mizude........ .409. 405. <10•1
Fukasc.... • • .... ··· ........ 51. 119, 1){)... 27
OiwakcEnbara ....................... 22,328.80

Neosclul'agerina margaritae-craticulifera subzone
In this subzone. Neoschwagerina margaritae, .V. craticulifera. and N. larga are
the conspicuous species. and they areassociated with Parafusulina japonica,
P. kinshoensis. Verbeekina 1.·ebveeki, V.
sp., Pseudodoliolina ozawai and Neosch·wagerina sp. The subzone is situatedin the upper part of the Funafuseyama
Limestone.
The localities are as follows.
5)

Locality names
Locality numbers
Nakagoshi. ........................... 87
Mizude .............................. ·13. 47
Fukasc .............................. 53
Nakagashi .......................... 287

342

lvun,itent MATSUMARU

513.

PusHhn·idH from t/:.u PnnafaHeyama Limestone

Shiraiwa ............................ 2
Oiwake .............................. 101
Mahira .............................. 323}·abeina igoi subzone
This subzone is the uppermost fusulinid bearing horizon in the Funafuseyama Limestone. and is characterized
by the occurrence of Neoscll!cagerinamargaritae. N. sp., and Yabeina igoi.
l'abeina igoi, from its structure, seems
to be the most primitive form of thegenus Yabeina.
The Akasaka Limestone yielded such
fusulinids as Yabeina g!obosa. }·. lwtoi,
V. inouyei and Neoschwagerina minoensis.
This fauna is thought to represent a
horizon higher than the rabeina igoi

6)

subzone above mentioned.
The fusulinids characterizing this subzone of the Funafuseyama Limestone
were collected from the following localities.
:vlizudc ................................. .'12
Tsubaki ................................ 77

3.

Range chart of the fusulinid fossils
from the Funafuseyama Limestone
The fusulinid fossils discriminatedfrom the abundant rock samples collected from the Funafuseyarna area arelisted in the Range chart. which shows
the respective ranges of the different
species are their relationship with theestablished zones and subzones.

Table I. l~angc chart of the fusulinid fossils
from the Funafuscyama Limestone.

-·..

~'"'"* ~"""'"-;·-~~""..,..'~
01~~·

~

, ..t.. •. •t"'!hO'fl'lll!>
N/....... 0 ....11

""""

"'~''
t~ ...

~· ~ltt,us.n,.1h ooiiY!i.\.ci•tl~'!'41"••

oi!'I'D0e1o..i

-- ·-

"•
'·'•MII....a !1.-ud••eo~loC+NOZ..t.oai~""".aQMotlil•.p.-ir'!P:tonc;o.,.._._,,.,..,..
...,cufoe,..H.r•l,p,a~·~,~-.~·•;lafg.a

N ITI¥g.lt ~••I!:_~;'.

v.. ~JUGJ

4.

Correlation
From the paleontological evidence ofthe fusulinids. the writer attempted tocorrelate the zones and sub zones recognized in the Funafuseyama Limestone
with some well known areas in Central

Japan, where paleontological and geo·
logical studies have been undertaken andpublished.
1\ Correlation with the Omi Limestone
According to KAWADA (195,1), the Omi
Limestone. Niigata Prefecture ranges inknntt.eru. ili!ITSUA!ARU
geological age from the early Carboniferous (Visean- Tournaisian) to the middle
Permian (Kungrian) based upon the
paleontological evidence of the fusulinid
fossils. The fossil zones of the Permian
System of the Omi Limestone are thePseudoschwagerina-Pseudofusulina zoneof the P 1 Formation and the Neoschwagerina zone of the Pr Formation. The
P 1 Formation is overlain by the P2 Formation with conformity. The Pseudoschwagerina-Pseudofusulina
zoneischaracterized by Pseudoschu·agen'na andParaschwagerina in the lower part ofthis zone and of Pseudofusulina in theupper part. The time range of Pseudofusulina is apparently longer than thatof Pseudoschwagerina and Paraschwagerina. and /'seudofusulina is also superior in numbers and species in comparison with the latter two genera.
Therefore. l'seudofusu/ina is considered
to characterize this zone.
The limestone of the P 1 Formation
has yielded Pseudofusulina kraffti, aspecies which also occurred from theParaschwageri 11 a
(Ar:ervoschwagerina)
subzone of the Funafuseyama Limestone.
However, Pseudofusulina z;zi/garis, which
occurred from the upper part of the P 1
Formation was not found in the materials from the Paraschwagerina (llcer·
voschwagerina) subzone of the Funafuseyama Limestone. although leo and OGA·w A (1958) recognized it in the lower
part of the Funafuseyama Limestone,
which probably corresponds to the Para·
schwagerina ( ..1cervoschwagerina) subzoneof the writer.
Accordingly.
theParaschwagerina(Acen·oschwagerina) subzone of the
Funafuseyama Limestone may be correlated with this zone of the P 1 Formation of the Omi Limestone.
On the other hand, the Neoschzcagerinazone of the P, Formation can be s•.1h-divided into three fossil assemblages
based upon the outstanding species.
namely. the lower is the Schwagerina(=Parafusulina) japorzica assemblage;
the middle is the NeoschuHtgerina douuillei and N. craticulifera assemblage ;and the upper is the Neoschwageritla
margaritae assemblage.
Thus, a part of the P2 Formation may
correspond to the Neoschwagerina margaritae-craticulijera subzone of the Funafuseyama Limestone from the mutual
occurrence of I\'eoschwagerina craticulifera and N. margaritae.
The fusulinids discriminated from therolling limestone blocks of the Omi
Limestone are Sumatrina cfr. annae,
Yabeina cfr. globosa and others. Thesefusulinids belong to the }'abeina zone,
and are thought to have been derived
from the uppermost part of the Neoschwagerina part of the Neoschwagerinazone of the P, Formation. If the Yabeina zone is actualy developed in the uppermost part of the .Yeoschwagerinazone of the P 2 Formation, the Yabeina
igoi subzone of the Funafuseyana Limestone may be correlated with the part
between the lower part of the Yabeina
zone and the uppermost part of the
Neoschwagerina zone of the Omi Limestone.
2) Correlation with the Akasaka Limestone
MOR!KAW A (1958) recognized 15 fusulinid zones in the Akasaka Limestone.
This limestone is situated at about 29 km
west from the present area. The 15
fusulinid zones, in ascending order, areas follows :Pseudoschwagerina zone
Pseudofusu/ina grrmrmz-avenae zone
Parajusulina zone
Parafusulina japonica zone
Pseudodoliolina ozau·ai zone
Neosclu!'agerina nipponica zone

51-'J.

F'tumlinids from lhe Fnnafuseyama Dimestone

Neosclzu·agerina zone
Neosch1cagerina fu:t'eiiSis zone
Neoschwagcrina rotunda zone
Neoschwagerina colaniae zone
Neoschwagerina craticulifera zone
Neosclueagerina margaritae zone
Neosclm;agerina /arga zone}'abeina zone
Yabeina igoi zonel"abeina globosa zone
Yabeina kaloi zone
Yabeina inouyei zone
Neoschu·agerina minoensis zone

The Pseudojusulina gnmum-avenae
zone which is the lowest fusulinid zone of
the Akasaka Limestone is characterizedby Pseudofusulina grunum-avenae, andParafusulina tnmcata besides otherfusulinids. This zone is equivalent to
the horizon Np? (Benijima) described by
OZAWA (!927). and is considered by KoBAY ASH! (1957) to correspond to the/'arasch wageri 11 a(: !cen)(Jschwager ina)
subzone of the lbukiyama Limestone,
and also by the writer to the Paraschwagerina (Aceruoschwagerina) subzoneof the Funafuseyama Limestone. becauseof the occurrence of mutual species.
Therefore, the Pseudofusulina grunumave/zae zone of the Akasaka Limestone
can be correlated with the Paraschu,agerina (Acervoschwagersna) subzone of
the Funafuseyama Limestone.
According to the correlation table of
MrKAl\H (1965). the Akasaka Limestone
and the lbukiyama Limestone do notalways correspond to limestones of thesame names mentioned by KoBA Y ASH!.
There may be confusion in stratigraphic
terminology, subdivision of the limestone
and other problems. Thus. it seems
evident that there still remains moreroom for study concerning the correlation of those limestones.
The characteristic species discriminated from the Parafusu/ina japonica~45

zone to the Neoschwagerina larga zoneof the Akasaka Limestone. point to thatthey can be correlated with the Para.fusulina zone to the 1\ieoslzwagerina mar·garitae·craticulifera subzone of the
Funafuseyama Limestone.
The J ·abeina igoi zone of the Akasaka
Limestone which is characterized by Yabeina igoi is associated with Neosrlzwa·
gerina margaritae. Verbeekina sp .. Ya/Jeina
cfr. multiseptata and other fusulinids.
The Yabeina igoi zone of the Akasaka
Limestone can be correlated with theYabeina igoi subzone of the Funafuseyama Limestone, from the mutual occurrence of the short-ranged species. Yabeina igoi.
3) Correlation with the Ibukiyama
Limestone
The lbukiyama Limestone was subdivided by KoBAYASHI (1957) into thefollowing subzones: (in ascending order1
Pa rasclzwagerin a ( f[cerposch u•agcri 11 a)
subzonePseudofusulina ambigua subzone
Parafusulina safJperi subzone
l ..ieoschwagerina craticullfera subzone
Neosclzwagerina margarilae subzone
Yabeina subzone
The Paraschwagerina (Acervoschwagerina) and Pseudofusulina amhigua subzones which is in the lower part of the
Ibukiyama Limestone has yielded J'araschwagerina {ilcervoschzcagerina) cfr.
lwgemoriensis. Parasclzwagerina (:lcervosclzwagerina) sp., Schwagerina lzawllinsi.
Pseudofusulina ambigua. Psf. cfr. rulgaris. Psf. sp., MinoJapanella sp .. Sclzu·/Jeri ella sp. and other fusulinids.
This fusulinid fauna from the lbukiyama Limestone is quite similar to thatof the Funafuseyama Limestone, andshow that the two can be correlatedwith each other.
The Parafusulina sapperi subzone to
the Neoschwagerina margaritae subzone

Kuni.tern i1!.4TSUMARU:Wi

of the lbukiyama Limestone can be correlated with the Parafusulina zone to
the Neoschwagerina mragaritae-craliwlifera subzone of the Funafuseyama Limestone, based upon the occurrence ofmany important mutual species as Parafusulina japonica, P. gigantojaponica,
Pseududoliolina ozazcai, Neoschwagerinanipponica, .Y. craticulifera, Y margaritae
and others.
Although the correlation between thelimestones mentioned above seems to bereasonable, it should be noted that Neoschwagerina fuwensis. N. rotunda, andN. colaniae from the lower part of the
Neoschwagerina zone of the Funafu:;eyama and Akasaka Limestones have notbeen found from the lower part of the
Neosclm•agerina zone of lbukiyama Limestone, and Neoschzcagerina larga fromthe upper part of the Neoschwagerinazone of the Funafuseyama and Akasaka
Limestones has not been recorded fromthe upper part of the Neoschwagerinazone of the lbukiyama Limestone.
The Vabeina subzone of the uppermost
part of the lbukiyama Limestone which
overlies with conformity the Neoschtmgerina margaritae subzone has yielded
no definitely determined species of Vabeina, although the following ones havebeen found, namely, rabeina cfr. katoi,
Y. cfr. cascadensis, r. sp., j\leoschwagerina craticulifera, l\i. margaritae, Verbeekina, verbeeki, and Pseudodolio/ina
ozawai.
l\•loreover, from the phylogenetic relation of the subfamily Neoschwagerina
from the Akasaka Limestone, according
to MORIKAWA and SUZUKI (1961), theTable 2. Correlation between the Funafuseyama
Limc:;tonc and other areas .

.
.'"
c:
~

:or~e

!
~

subzone0":z

U.fuw.~ns•ssubt on~S~f---''"'o"-r_,_o_,N.fuweon-:.is

•

::one

~-----+---='o"'n~•- l - - - - l(P, forma.tion)

N.nipponkat•r.

ParafusuHna

0

"
~
&

zone

P.sapp::-r i

LOft~

Psdd,Qzaw,:iNn-N~•,.;

subzo·..,l---"--''0"'"'.:.•_.•~ipponica,
P. j,aponi:::;,. '
zone

zontt

Paraschwagf!rina Psf_a":.~:~:r.e Psi.:~~~~;- N p ?

• ~ce

;

"

ptAcer ... oschw

subzone!;ubzon!

zone

Pscudos.t.l-tNagerina(Benijima)-PseudolusuHna:on~( P1 Formati o-)513.

Fnsuli.nids fnnn the Funafnseymlw Limestone

range of l"abeina katoi r~pres~nts thetime range of th<! !'abeiila }!lobosa to r.
inouyei zone of the Akas:tka Limestone.
and also associated with Neoschwagerina.
Though the rabeina Sp?cies from the
Vaheina subzone of the lb:tkiyama Limestone is not identical with Vaheina lwtoi
but rather nearer to Yabeina cfr. lwtoi.
the uppermost rabeina subzone of thelbukiyama Limestone can probably becorrelated with the }'abeina lwtoi zoneof the Akasaka Limestone.
From the reasons given above. thewriter considers that the Yabeina igoi
subzone of the Funafuseyama Limestone
may be correl:lted with the lower partof the Yabr:ina subzone of the lbukiy:-tm<lLimestone.

axis of coiling. polar ends bluntly r,oint·
ed. :vlature specimens of l to 5 volutions, measure 3.5 to 4.3 mm in lengthand 1.9 to 2.3 mm in width. giving form
ratios of 1.8 to 2.0. shell gradually expands from first volution outwards. fusi·
form throughout shell growth.
Proloculus large. usually irregular.
with outside diameter of 0.37 to 0.50 mm.
averaging 0.'12 mm. Chambers expand
slowly. nearly uniform in height in central part of shell but become higher
nearer to poles.
Average height ofchambers from first to fifth volution in
adult specimens about 0.12. 0.13, 0.22.
025, 0.23 mm, respectively.
Spirotheca thin. composed of teclllm
and coarse keriotheca in outer volutions
and fine keriotheca in inner volutions.
Average thickness of spirotheca fromfirst to fifth volution in adult specimens
about 0.034. 0.035. 0.054, 0.068. 0.068 mm.
respectively.
Septa thin. closely, and regularly
fluted throughout shell. Fluting reachouter spirotheca near polar regions.
Chamberlets small and low.
Chomata and tunnel indistinct. Axial
deposits fill all volutions except central
portions.

Hystematic DescriJ)tion
Genus Parafusulina DU?\HER
and SKIN:'\F.R, 19:H
Parafusu/ina funafusen.<;isl\(ATSU:IIARU, n. sp.

Pl. 37. figs. 1-1•1
JJescription:
Shell small. fusiform.
with nearly straight to slightly convex

l'vleasuremen ts (in mm):
- -

Pl.

- - - ----··~--

---

---

Fig.

L.

w.

F.R.

1

4.3
3.6
3.5

2.3

1.9

-1-- 0.40
5

1.8

2.0

4

1.9

1.8

4

5
•I

37
37
37
37
37

-------

2
3
-1
5

?

1.8

?

3.5

1.9

1.8

?

2.3

N.V.

-1

D.P.

0. 50
0. 40
0.27
0.43
0.37

Kuniteru AfATSUMARU
Height of Volutions

Thick ness of Spi rotheca-----~-

1
2
I------0. 10
0.09
0.15
0.11
0.08
0.17
0.07
0.11
0. 14
0. 14
0.15
0.19

3

4

5

1

0.21
0.23
0.25
0.17
0.23
0.22

0.20
0. 3·1
0.20
0.23
0. 19
0.37

0.18

0.030
0. 05:~
0.017
0.027
0.033
0.0-17

I

w.

L.

Fig.

Pl.

0.29

-3
4
5
2
I - - - - - -I---0.063 ' 0.063
0.050
0.033
0.037 I o.on
0.093
I
0.030
0.050
0.050
0.087
0.027
0.053
0.077
0.040
0.050 ' 0. 057
0.0-130.036
0.067 I
F. R.

N.V.

D.P.

------

6
7

3F
3t

2.5

Height of Volutions

Thickness of Spirotheca

1

2

3

4

1

0.11
0.17

0. 18
0.33
----

0.24
0.-10

0.37
0. 37

0. 037o. o:1o--Septal count

1

2

8
7

12
12

3

I 3
0.040
0.050
0.060 I 0.067
2

CnE:>~, S.

12?

Remarks: The present form resembles Pseudofusulina houziguanica SIIE~Gin general shape but is distinguished byits highly and strongly fluting thicker
septa. This species also can be distinguished by its inlluted form and larger
proloculus from Parafusulina okubocmsis
OZAWA emend (i\fORIKAW A) Parafusulina
cayeuxi (Dr~PRAT) resembles this species,
but differs in the more elongate form
and irregularly fluting of septa.
Occurrence: The species occurs commonly in the Funafuseyama locality andis associated with Parafusulilla okuboensis, P. kinosakii, and P. sp ..
Locality: Funafuseyama Loc. No. 10,
14, 15.

40.090
0. 067--

References cited.----------16
20

0.53
0.60

(1956) : Fusulinidae of South China,
Part 2. Pal. Sinica New Ser. 13. No. 6. p.
-17. pl. 9, figs. 5-6, pl. 13, figs. 1-2.
DEP!dH, ]. (1913):
Etude des Fusulinides
de Chine et d'lndochine et classification
des calcaires (2 Memoire). Les Fusulinides des calcaire carboniferiens ct permiens du Tonkin, du Laos et du NordAnnam. Atem. Sen. Geol. d'bzdochine,Vol. 2·. fasc. l, pp. 1-74, pis. 1-10.
- - (1914) : ]~tude des Fusulinides du japan,
du Chine et d'lndochine et classification
des calcaires
fusulines (3 Memoire).
Etude
comparative des Fusulinides
d'Akasaka (Japan) et des fusulinides deChine et d'lndochine. Ibid., 3, fasc. l. pp.
1-9.
(1915): Etude des Fusulinides de Chineet d'lndochine et classification des calcaires ;\ fusulines (4 Memoirc). Lcs
Fusulinidcs des calcaires carbonifcrience
et permiens du Tonkin. du Laos et duNord-Annam. [bid .. Vol. 4, fasc. ], pp.

a

513.

FHsulinids from thf' Funafusryarna. Lim.estmze

1-3, pis. 1-3.
FUJI~IOTO,

et a/. (1962): The Upper
Paleozoic System of the Hida Massif.
Ceo/. Res. Hid a 1Hassij (in Japanese: pp.
Hida Sanchi no Chishitsu Ken Kyu), pp.
44-70. figs. 1-5, table 1.
FuJil,IOTO, H. (1936) : Stratigraphical andPaleontological studies of the Titibu
system of the Kwanto .Mountainland.
Part 2. Paleontology. Tokyo Bunrika
Daigaku, Sci. Rep/ .. Sec. C. Vol. l, No. 2,
pp. 29-125, pis. 1-26.
IIANZAWA, S. (1939) : Stratigraphic distri·bution of the Genera Pseudoschwagerinaand Paraschwagerina in Japan with Distribution of Two New Species of Pseudo·
schwageri1ra from the Kitakami Moun·tainland. Northeastern Japan. japan. jour.
Ceo!. Ceogr .• Vol. 16. :'{o. 1-2, pp. 65-73,
pl. ·1.
lc.o, H. and OGAWA, K. (1958): Fusulinids
from the Funafuseyama Limestone. part
1. jub. Pub[. Commem. Prof. II. FuJI~IOTo,
pp. 49-S<I, pis. 1-2.
leo. H. (1959): Some Permian Fusulinids
from the Hirayu District. SoutheasternPart of the Hida Massif. Central Japan.
II.

Sci. Rept., Tokyo Kyoiku Daigaku. Sec. C.
Vol. 6, No. 56. pp. 231-25•1.
- - (1964) : On some Pseudoschwagerina
and Zel/ia from Japan. J~ur. Pal., Vol.
38, No. 2, pp. 281-293, pis. 45-·16.
I rSAI<A. G. (1932) : On a new species ofSchellwiena, Sell. ozawai from the Mt.
Funafuse District, Prov. Mino, Japan.
Ceo/. Soc. Tokyo. jour. Vol. 39. p. 1-4.
ls1111. A. et al. (196:i) : Geology of the
Funafuseyama District, Gifu Pref. (inJapanese), Geol. Soc. japan, jour., Vol.
69, No. 808-819, p. 340.
Js1m, A. (19&1) : Geology of the Funafuseyama District, Gifu Pref.. (Part 2). (inJapanese) Ibid, Vol. 70. No. 826, p. 381.
KA:-I~IERA, K. (1958):
Fusulinids from the
Yayamadake Limestone of the Hikawa
Valley, Kumamoto Prefecture. Kyushu,
Japan, part 3-Fusulinids of the Lower
Permian. Kyushu Univ .. Mem. Fac. Sci.
Ser. D. Geol.. Vol. 4, No. 3. pp. 153-215.
pis. 24-35.

- - (1961): Middle Permian Kozaki Formation (in Japanese). Ibid., Vol. 5, No. 4,
pp. 196-215.
KAWADA, S. (1954): Stratigraphical andPaleontological Studies of the Omi lime·stone in the Itagami district. Niigata
Pref. (in Japanese). Sci. Nept .. Tol<yol('voiku Daigaku. No. 3. pp. 15-27.
- - (1954): Stratigraphical and Paleontological Studies of the Omi limestone in
the Mt. Kurohime District .. Niigata Prcf.
Miscellaneous Nept., Res. Ins/. Nat. Resources., No. 35, pp. 48-56.
- - (1954): Stratigraphical and Paleontological Studies of the Om i limestone in
the Mt. Myojo District, Niigata Pref.
Ibid., No. 36. pp. 39-48.
KOBAYASIII. M. (1957) : Paleontological studyof the Ibukiyama Limestone, Shiga Pre·
fecture, Central Japan. Tokyo l(l'oiku
Dai~aku, Sci. Rrpt.. Sec. C. Vol. 5. No. ·18.
pp. 247-311, pis. 1-10.
KoBAYASHI. Y. (19Sl): Geology of the Funafuscyama District, Gifu Prefecture andso·called Funafuseyama Klippe (abstract).
Geol. Soc. japan, jour .. Vol. 71, No. 841.
pp. 475-493.
1\frKA:\11, T. (1965) : The Type Sakamotozawa
Formation. (in Japanese), Geo/. Soc.
japan, jour., Vol. 71, No. 841. pp. 475-493.
figs. 1-7, tables 1-3.
MoRIKAWA. R. (1953): Some Schwagerinaiike Parafusu/ina. Saitama Unir.• .. Sci.
Rept., Vol. l, No. 1, pp. 28-3,1, pl. 2.
- - (1955) : Schwagerinidae in the Vicinityof the Shornaru Pass. Eastern Part of
K wan to Mountainland, Central Japan.
Ibid .. Vol. 2, No. I. pp. 45-IH pis. 5-15.
- - and IIORIGliCIII. M. (1956) : Parafusulina
nakamigawai n. sp. from the Adoyama
formation in the neighborhood of Kuzu
City, Tochigi Prefectun•. Ibid .. Vol. 2.
No. 2, pp. 261-264, pl. 35.
l'.IoRIKAWA. R., e/ a/. (1958): Fusulinids
from the Akasaka Limestone (Part I).
Ibid., Vol. 3, No. l. pp. 95-130, pis. 12-26.
MORIKAWA, R. (1960): Permian Fossil Zones
in Japan. (in Japanese), Fossil. No. l. pp.
51-61.
--and ISOMI. II. (1960): A New Genus,

::Go

Kuniter!l ilfATSUAIAl<U/Jiwae//a, Schwagerina-like Schuberti!l/a.
ibid .. Vol. 3, No. 3. pp. 301-305, pl. 54.
- - (1961): Studies of Permian Fusulinids
in the cast of Lake Biwa, Central Japan.
Geo/. Survey. japan, Rept .. No. 191, pp.
1-30. pis. 1-21.
- - and St'ZLIKI. Y. (1961): Fusulinids fromthe Akasaka Limestone (Part 2). Saitama
Uni11, Sci. Rept., Ser. B. No. I. pp. 43-7·!.
pis. ·1-22.
- - (1962): Solidgraphical study of Fusulinid Foraminifera. (in japanese). Geol.
Soc. japan, jour. Vol. 68. No. SO.t pp.
518-521. figs. 1-4. pl. 5.
NAKA~IURA.
l\1. (1965) :StratigraphicalStudy of the Funafuseyama Dolomite (inJapanese), Gypsum and Lime No. 76. p.
3·18-358, figs. 1-5, table 1.
OzAwA, Y. (1927): Stratigraphical Studiesof the Fusulina- Limestone of Akasaka,
Province of Mino. jour. fac. Sci., Imp.
Unit'. Tokyo, Sec. 2. Vol. 2, art 3. pp.
121-164. pis. 34-35.
SJJE:-<G, J. C. (1963) : Permian Fusulinids ofKwangsi, Kueichow and Szechuan. Pal.
Sinica. New Ser. B. No. 10, pp. 129-247,
pis. 1-35.
To~l!·soN, !\L L. (19·18): Studies of American
Fusulinids. Univ .. 1\cmsas. Pal. Contr ..
Protozoa Art. 1. pp. 1-184. pis. 1-38.
- - (1954) : American Wolfcampian Fusulinids, Ibid .• Art 5. pp. 1-226, pis. 1-52.
- - (1964): Fusulinacea, Treat. Invert. Pal ..
Part. C. Prot. 2, Vol. 1, p. C 358-C 436.
ToRIYAMA, R. (1958): Geology of Akiyoshi,
Part 3, Fusulinids of Akiyoshi. Kyushu
Univ., Afem. Fac. Sci., Ser. D., Geol., Vol.
7. p. 1-264, pis. 1-48.

----llatsushikadani f)) l.lli tiNakagoshi
~E
fill
·7]-:
MizudeII'.
Kash ibaradahi ~IJR tiFukasej;fitli
Shiraiwa.{:__j!:'!
Oiwakeif!.
51
Konzima
NfJ<7
Iwatani
'G
~
~

Iwodo
Takaradani
Nakagashi
MahiraAkadaru
EnbaraKanzakiTsubakiMatsuda

{JI

n Ji
1+

::.iC:

111

~~

r~

-'I'-

m

;A":

llJ
1'1'

/Jf
~~i

Hi

m m

-·----- - - - - - - - - - - - - - - - - - -Explanation of Plate 37
Figs. 1-1·1. Parajusulina funafusensis MATSC\tARU. n. sp.
1-Axial section. holotype, x 12. IGPS coil. cat. no. 86620-a.
2·-Axial section. paratype. x 12. IGPS coil. cat. nn. 86620-b.
5-Axial section, x 12.
3, ·l. 8, 9-0blique sections. x 12.
6, 7-Sagittal section,;. x 12.
10-Fig. 6. enlarged 60 times. to show the proloculus pore and spirothecal structure of thetectum and coarse keriothcca.
11-Fig. 1. enlarged 56 times. to show the spirothecal structure of first to fifth volutions.
12-Fig. 7. enlarged 52 times. to show the proloculus spur.
13-Fig. 4. enlarged 51 times, to show the proloculus pore and the circum-proloculus
chamber.
1·1-Fig. :~. enlarged 55 times. to show the spirothccal structure in oblique section.

MATSUMARU:

Fusulinids from the Funafuseyama Limestone

1

4

10

11

13

KuMAGAI

12

14

and M A T SUMARU photo,

Plate 37

Trans Proc. Palaeont. Soc. Japan. N. S .. :.ro. 6!. pp. 351-358. pis. 38, 39, Dec. 15, 1966

514.

ON A REPTILIAN SKELETON FRO.M THE PALAEOZOICFORMATION OF SAN PAULO. BRAZIL*TOKIO Sl-liKAMA
Geological Institute, Yokohama National University
andHIROSHI OZAKI
Xational Science Museum, Tokyof7 :hL-. "t /,: t7 ~ O)(l{ :lW•'rJI'l:l!.h 'l'TifttiT:

Brazil. San Paulo ift'i;f> Tatui O){ttlJfJf
Brazilosaurus sanpau/ oensis;~DJJJXO) E~J~·c-·1H!. t ~ t t:lre!l•. h'l'l'ttit '2 Pr~tli~ll c li- C.:.'),.-!:: ,;fifi, fic·i&L.t.:.,;:,0)·:;;-t!T:;t. -F;7Ji-:1![i'f,O> Irati /WO)/;J(~:fi/;J.:'f-iri',~·l'>)E~~:!1. tuiJj:,fiO)
fJ:r!IHJ~tJr:J: · t=t:~ r~

fri* •T(- ·t\: ::: J: -:rc~h'f J:. :.:. t t:: r:,;::: ht.:. {, O)·r_' zb ;:, ,In Autumn of 19&1, \fr. Zyuichi SAIKI
in Fukui City. formerly a member of theImperial Diet, submitted the writers a
reptilian skeleton for study. which was
donated to him by t\1r. lnosuke liA!\A·
YA:\IA in Tatui. San Paulo, Brazil. dur·ing his journey there in April of 1964.
HANAY A:\!A found the fossil in his farmin Tatui during the clearing of land.
His farm is situated 16 km NW of Tatui
City and the area is composed of late
Palaeozoic formations. From the lower
Permian Irati formation, Mesosauru s
brasUiensis ivlcGREGOR was reported in
1908, and generally speaking Mesosaurus
is rather common in the Brazilian Permian. The fossil now on hand is preserved in a light creamy to grey limestone of fine lamination. The bed whichyielded the fossil may belong to the Irati
formation.

Here the writers extend their cordial
thanks to Mr. SAIKI for his generous
offer of the specimen for study. Also
they express their hearty gratitude toProf. F. HUENE in Tiibingen, Prof. A.
ROMER of the Harvard University, Dr.
L. I. PRICE in Rio De janeiro, [vliss R.
FUSEJIMA o( the National Science ivluseum, Tokyo for their valuable suggestions and kind help during the courseof this study.

Brazil osauru s sa npaul oensis
gen. & sp. nov.
PI:;. 38, 39

Ho/otype: A skeleton belongs to Z.
SAIKI and is preserved at the National
Science Museum in Tokyo.
Locality: 1-l.>\~A YAJ\!A's farm. 16 km
NW of Tatui. San Paulo, Brazil.
Formation: Irati formation (?): lower
Permian.

- - - - - -* Recei\·ed April 19. 1966: read January.
1965.

351

352

Tokio 8FllKA.MA and Hiroshi. OZ,I/(1Description: Skull, vertebrae and costae of presacral region and fore limb
largely preserved ; skull and cervic vertebrae preserved on their left lateralside. while thoracic- and lumbnr regions
on their dorsal side, and a torsion isfound on p:>sterior part of cervic vertebrae, by which the animal much bent
its neck backw:~rd. Caudal region andhind limb almost unknown.
Animal slender, elongate and smallsized like Mesosaurus. Cervic region
very long and longer than skull. Animal
272 mm long as preserved; skull, vertebrate column of cervic region and thatof thoracic-lumbar region 46, 86 and
136 mm long respectively. Body ver~narrow, about 25 mm wide at ninth
thoracic vertebra. Fifteen cervic-. nineteen thoracic-lumbar-, one sacral- and one
caudal vertebrae are retained. Detailed
precise characlerist ics of each hone,
especially of skull and jaw. almost invisible because surface of bone detached.
Skull and teeth.
General outline of skull rather likethat of .\Iesosaurus. It is very long,
low and with strongly projected snout
and small cranium. Characteristic of
this animals. teeth short and not so muchdeveloped as in Mesosaurus. Orbit relatively large, 6x4 mm in diameters. subquadrate and situated aboye the thickest
portion of lower jaw. Premaxilla long,narrow and acutely projected forward.
Nasal also long and narrow. but maxilla
relatively thick and high compared withthat of lvfesosaurus.
Preorbital andpostorbital foramens as seen in kfesosaunts unclear in preservation. Jugal andprefrontal probably same as that of
Mesosaurus. Dorsal margin of cranium
gently cun-ed and its posterior dorsal
corner broken. Parietal detached fromskull preserved apart from it. irregurllay
quadrate and has corrugated surface.

Postorbital runs vertically. narrow, forming a posterior margin of orbit andwith relatively distinct posterior margin.
Quadratojugal relatively thick and supratemporal broad and developed. Surfaceof posterior portion of cranium corrugated. Cranium relatively larger thanthat of ,\!esosau r 11 s.

4cm......._~___._......_~

Text-fig. 1. Skeleton of Brasi/iosaurus
scwpauloensis sp. nov.

Lower ja\v long. narrow and straight,
its ventral margin not recurved as in
Mesosaurus; it is highest at a point
just below posterior margin of orbit.
Posterior dorsal corner of jaw broken.
Dentary large, long and very straight.
Suprangular and angular relatively shortand high. There are many homoeodont
teeth which are short, acutely pointed, alittle curved backward, probably conical::~nd seem te be pleurodont.

514.

:~53

l'alrwozm:c Brazilosaurus ·n. !/tit.

Po

0

1

3cm

Text-fig. 2.

Skull of /Jrasi/osaurus sanpauloensis sp. no\· .

.M: Maxilla. Pm: Premaxilla. N: Nasal. F: Frontal, 1': Parietal. Prf:
Prefrontal. J: Jugal. Po: Postorbit:tl. Qj: Quadratojugal. Q: Quadrate,!:iq: Supraquadrate,[): lkntary. A: Angular. Sq: Supraangular, At:
Atlas, Ax: Axis.

Longitudinal width

Maximum hl'ight

12. 2±
19.1±

1. 7 ± (mm)2.·1±

14. ·I±12. 4±

1.3±
1.0±

4.0
1.7±

3.2
4.3±

8.1±
6.9±

0.8±

41.9

3.2
1.9

Premaxilla
MaxillaNasal
FrontalParietal as preserved
Postorbital
Quadratojugal
Supratemporal
Lower ja\\'
Dentary
AngularSupraangular

Vertebrae
As first cost a as preserved being at·tached to sixteenth vertebrae. there
seems to be fifteen cervic vertebrae,
stout and relatively broad transversally.
Four vertebrae from fifth to eighth.
longer than broad and fifth vertebrae
longest of them. Atlas relatively shortand with well developed neural spine.
dorsal margin of which much con vex
upward and posterior dorsal corner distinctly projected aftward. Transverse
process running postero-laterally. Axis
slightly longer than atlas and has vaulted

33. 9±
16.3±
12. 2±

1.3±
1.5±

neural spine. Third cervic vertebrae
relatively short. quadrate in transverse
view. with recurved dorsal and aft margins; transverse process relatively thick.
Fourth cervic vertebra longer than third
one with relatively straight dorsal margin. Fifth and sixth vertebrae elongate
quadrate in lateral view, w·ith straightdorsal margin and thick transver~e process. Four vertebrae from seventh to:enth ones quadrate in lateral view.
Eleyenth and t wei fth cervir yertebrae
relatively short and fifteenth vertebrae
wider than long.

Dorsal side of thoracic-lumbar verte·brae preserved and centrum of them
becomes wider posteriorly from first to
eleventh vertebrae. Left lateral bordersof centrum from twelfth to ninteenthvertebrae broken, especially eighteenth
one. Centrum of thoracic-lumbar verte·brae generally wider than long, althoughthird one almost as wide as long; they
are quadrate in general outline and theirright lateral margins gently convex out·
ward. Neural spine relatively thick andprojected posteriorly as seen in fifth,
eighth- eleventh vertebrae. Left transverse process preserved on vertebrae;\·led ian longitudinal lcngl h of cen Lrum
Transverse width of ditto at middle
Maximum height

C9

C10

C11

C12

from fourth to eighth, flat, longer thanwide and its free margin irregularly
curved. Spine-like short transverse pro·
cess is found on left posterior side ofvertebrae from twelfth to fourteenth.
This may indicate that vertebrae poste·rior of eleventh belong to lumbar region.
Transverse process of eleventh andtwelfth cervic vertebrae also spiny andmuch pointed. Pelvic girdle seems tobe jointed with the twentieth vertebra
as preserved, it may be a sacral vertebra. Only one caudal vertebra preserved,
which is small sized, quadrate and alittle wider than long.
C1

C2

C:{

Ccl

C5

C6

'1. 1+

4.3-;-

5.3+

5. 7

7.5

6. 7

7.3

·1. 5

"J. 3

4.5

5. 3

5?

6. 9

6. 1

6.4

•1. 5

4.5

·1. :~

C13
5.1

6.6

7.4

8.3

10.0

9.6

8.3

6. 1

C7
C8
---

T5
6.5
7.0-8.3
TLlcl

Cl4

Cl5

Tl

T2

T3

T4

5.6

5. 7

5.8

6. 6

6.9

6.9

6.5
6.8

6.8
6.8

5. 7
6.8

T6

T7

TB

T9

T10

Tll

TL12

TL13

6.·1±
7.0-9.1

6. 7±
7.5-9. 7

7. 9±

7. 6±
8.6

6. 7±
8. 6

6.0±
8.8

6.6±
8.1±

6.3±
7.5±

TLI9

Sl

CAl

•I. •! ±6.6

5. 5

5.8 (nun)
6.6

TL15

7.0±

6.8±

8. 7::t8.4+

8.2-9.8

TL16
TL17
TL18
----6. I ±6. 7::t
6. 5±
8.6
7.2·1·
8.4+

CostaeThere are preserved above twenty
costae on right side and seven costae onleft side of the skeleton. First costa ofboth sides jointed with first thoracic- or
sixteenth vertebra from first cervic one.
Both right and left costae jointed withyertebrae on first. second. seventh. eighthand ninth ones, while right costa jointed
with vertebrae on fourth. tenth-seventeenth ones: left costa jointed with fifth

6.5

and sixth vertebrae. Isolated costae ofright and left third, left fourth. left fifth,
right sixth, sixteenth, seventeenth andeighteenth are preserved on right sideof the sl<eleton. Right sixth costa pre·served in a position nearest to skull, andsixteenth. seventeenth and eighteenthcostae arranged in distant positions.
Costa gently curved although its proximal portion strongly curved. Dista!
portion of two-third of total length be·51.4.

Pa/.acoz1Yic Braz·ilosan·rn.'-1 n. ue1i.

comes thicker gradually distal ward. l t
is not pachyostosed as seen in ;\lcsosaurus.
First right. first and second left costae
preserved turning forward: first costae
as preserved relatively short and lirstleft one preserved only proximally.
Second left costa also turned forward.
Right and left third costae preserved
appressed to each other and parallelwith vertebrate column. Right fourth
costa as preserved running between
fourth thoracic-lumbar- and sixth cervic
vertebrae: left fourth costa lies posteriorly of right fourth costa in parallelsituation. Right fifth costa as preserved
concave outward, lies posterior side ofleft fourth costa and right eighth costa.

Left fifth to ninth costae preserved intheir natural position. Right sixth costapreserved bet\veen skull and righteighth costa. concave outward and retainiug an entire outline of costa. Leftseventh-seventeenth costae preserved intheir natural position. Distal portion of
ninth-thirteenth costae closelyappressedto each other. Sixteenth and sewnteenthcostae of probably right side run
straight and detached from their natural
position. Right eighteenth costa also
detached from eighteenth thoraciclumbar vertebra and concave inward.
There are seen many fine small sized
ventral costae on ventral side of third
to eighth cervic vertebrae.

__.___.._

2,....._....._
l~
L

3
___._
R
L

6.4 24028.0 28.6
1.0 1. •I

31.2 16. I
1.8 1.8,....._.....89_..._,

R

Length as preserved along the anterior border'IVidth at distal cnd as preserved

,,
R

L

31.4

27. 1
1.5

1.4

:~5. 8

1.8

1.5
7

6
____,_
R
L

5
_._._
L
R

-----'-

L

R"L

R

_
L

R

L

3-1. 1

21. 4

1.9

1.8±

35.7
:2.0

30.2
1.6

32.9
1.8

30.4
1.5±

30.0
2. 1

31. 7
1.9

1-1
R

15

16

17
,....._....._

R

R

R

L

18
R

28.2
1.7

25 6

22.6

24. 6 (mm)

2. I

8.0
1.2

18.5

1.6

10
R

11

12

n

R

H

R

35.1±
1.8±

20. 0-l~

14. 2

23.0

1.7

1.5

1.5

Girdle and limbs
Scapula and coracoid unknown. Thereis a suboval bone of small size. attachedto the distal end of left fourth costa,
3.4x 1.8 mm, and has a corrugated surface: it may be a part of pectoral
girdle but details unknown.
Humerus rodlike, straight. becomingslightly wider distally in right one andshows distinct expansion of distal end.
Proximal margin of head slightly recurved as seen in left humerus. On
left humerus, head trigonal in lateral

37. 7
1.8

1.7

view and shaft narrowest at middle.
Distal end wider in right humerus aspreserved than in left one and posterior
margin of right one much concave backward w·hile its anterior margin almost
straight ; its distal margin also straight.
Distal end of left humerus convex distally and not so much expanded as inright one. Right humerus probablv indicates its outer side and left one its
anterior or posterior side. Head of righthumerus brol<en off.

Maximum length as pres(•rvcd
Width nf proximal margin as preserved
Ditto of distal margin

Ulna and radius of left side preserved
but ulna broken on its middle of shaft.
They are straight, slender and narrow:

Three carpal bones of left side pre·served attached to distal ends of ulna
and radius. They are subcircular andanterodislal one situated near distal
end of radius smallest. They may cor·
respond to ulnare. intermedium andradiale as seen in Mcsosaurus. Theirdimensions as follows from anterior toposterior ones: 1.6 X 2.5. 3.4 X 1.9, and
2.4 X2.0 mm. Small bones of centrale:\1aximum length as preserved
Maximum width of proximal end

Right ilium preserved detached fromskeleton and in a position apart fromleft posterior side of it. The bone large
sized, subtrigonal and with recurved andlong three margins, the upper one of

Ldt

27. l

2\l. 2 (mm)

2.2

5.2
'l. 6

8.8

both proximal and distal ends not somuch expanded.

,
Maximum length as preserved
Width of proximal end as preserved

Right

Ulna

Radius-~--~10. 5+
13. 1 (mm)2.1
1.9

unpreserved except one preserved besideright fifth metacarpus: it is suboval and
1.7 >< 1.3 mm in dimensions.
Right metacarpi of three bones fromthird to fifth preserved bet ween left
fourth and right fifth costae. They areelongate trigonal, slender. straight andnarrow; proximal margin gently curved
and distal end broken off; middle bone
as preserved longest.
III

IV

v

3.9

5. 9

5. :~ 111111

1.2

1.2

1.3

which being longest. Anterior margin
straight and short w·hile posterior one
concave aftward. Proximal margin gen·tly curved and concave upward. Acromion border convex downward.

Distance between both ends of upper border ........................ 15. '' mmWidth of acromion border . . .. . .. . . .. . . . .. . . . .. . . . .. . . . . . . . . . . . . . . . . 5. 2
Length along fore margin . . . . . . . . . . . . .
.. .. .. .. .. .. .. ..
10. 5Median height as presen·ed.......................................... 6. 4
- - - - - ·--. - - - - - -Explanation of Plate 38
Bra<ilusaurus S<llljJUulvensis SIIIKAI-.IA and Oz.\1\1. gen. & sp. nov. Holotypc, natural size.

SHIKAMA and OZAKI:

Brazilosau1·us

Plate 38

51-~.

l'alawzuic Brazilosazu us n. uc n.

Two fragments of bone preserved
scattered in a position posterior of ilium
but whether it belongs to ischium or topubis is unknown. At left side of ninth
thoracic-lumbar- and sacral vertebrae.
there are preserved rodlike bones. andat right side of them two rod like bones.
Largest bone of left side may be proximal portionofleftfemur. It is 14.6mm
long as preserved, gently cun·ed. concave
fonvard and head not so much expandedwith straight proximal margin of 4.1 mm
length. The other two bones of bothsides may be proximal portions of tibia
and fibula respectively. Tibia slender,
straight and with head not so much expanded: right one is 8.0 and left one
8.5 mm long as preserved. Fibula larger
and stouter than tibia with stout subtrigonal head : it is much expanded, -LO
and 4.5 mm in maximum width on rightand left ones respectively. Right andleft fibula 12.1 and 8.7 mm long respectively.
Diagnosis: Skull elongate, narrow andlow. with developed snout. Orbit large
and supratemporal-parietal region developed with corrugated surface. Teeth
short, many. conical. a little curved backward and may he pleurodont. Fifteen
cervic. ninteen thoracic-lumbar (probably
ten thoracic and nine lumbar), one sacralvertebrae are divided. Transverse process lanceolate on fourth to eighththoracic-lumbar vertebrae and shortspiny on twelfth to fourteenth one.
Neural spine thick and developed. Cervic region very long, longer than skull
and with spiny transverse process.
There are many line spiny ventral costae
on ventral side of cervic column. On all
thoracic-lumbar vertebrae there are
jointed costae of slender shape and gentle curvature; <'ostae not pachyostosed.
Humerus slender. rel<~tively long and its
distal portion much expanded.
Ulna,radius and fibula slender and rodlike.
Clnare. intermedium and radiale as in:\Iesosaurus. Ilium relatively large andits three margins moderately recurved.
Considerations:
In general outline
this species is rather like Mesosaurus,
but is clearly different from it by shorter
teeth. more cervic vertebrae, longer neck,
unpachyostosed costae and by morestrongly developed cranium with corrugated surface etc. In Mesosurus there
are ten cervic vertebrae. ivlany skulls
of ,\/. brasiliensis ~vlcGHF.GOR describedby j. H. \kGREGOR and F. v. 1-It;nmha\·e extremely long teeth which are
more extended than dorsal margin of
upper snout or ventral margin of lower
jaw when upper and lower jaws aredosely set to each other as seen in thisspecimen. In this specimen there is seen
no trace of long teeth. We can not accept these differences as owing to growthstages or sexual dimorphism. They are
fundamental and anatomical characteristics. U. brasiliensis is reported from thelower Permian Irati formation of Southeastern Brazil and Urguay. M. /enuidens GEH\'AIS. the type species of thegenus 1\1/esosaurus GEHVAIS. reported
from the lower Permian bed between the
Dwyka and Ecca formations in South
Africa. has also relatively large sized:;kull and twenty-nine presacral vertebrae. Hence :\Jesosaurus is safely excluded from comparison with this species.
Noteosaurus africanus BR00:\1,
regarded by HIJENE as same as Mesosaurus tenuidens. is out of comparison
owing lack of specimens compared.
Stereoslernum /umidium COI'E known
from the Permian of San Paulo, Brazil
is about 80 em long and has thirty-four
presacral (eleven cervic and twenty-threethoracic-lumbar) vertebrae. and rather
allied outline of skeleton. But il is nlso
dislinguished by relatively short neck,

;J5S

Tokio SHIKAM.-1 and Hi·rosht OZAKI

Jess number of cervic vertebrae andpachyostosed costae. Both 1Hesosaurus
and Stereosternum carry a very long tail.
and perhaps this species also has a longtail as long as presacral region. hence it
might be more or less 540 mm in bodylength, generally corresponding to theBrazilosaurus
size of Mesosaurus.
stands rather apart from the stock of
Alesosaurus and Stereostenwm fromtaxonomical points of view. In longneck it may be nearer Sauropterygia
than lchtyosauria.

References selected
Col'E. E. D. (1886) : Vertebrate Palaeontologyof Brazil.

!'roc. ,!mer. /'hit. Soc., vol. 23,

p. 7.

McGREGOR. j. II. (EIU8) :On Mesosaurus
brasiliensis n. sp. from the Permian ofBrazil. Fi11al Report Commissas de Estadias das Minas de Carvae cle f'edrtl do
Brasil. Rio de Janeiro.
HuE1\E, F. v. (1937) : Osteologic und Systematischc
StellungvonMesosaurus.
Palaeonlographica, bel. 92, abt. A. pp.

-15-58.
- - (1956) : Palaontologie und Phylogenie
der Niedcren Tetrapoden, pp. 2-13-2-15.
PI\'ETEAU. ]. (1955) : Traitc de Pah.'ontologie. Yol. 5, pp. 409-111.
Ro~tER. A. S. (1953) :Vertebrate Paleontology, Chicago.
- - (1~)56): Osteology of the Reptiles, Chi-

cago.

Explanation of Plate 39
IJrazilo.~aurus

sall/Hlllloeusis SJJIKA~JA and OzAKI. gen. & sp. nov. Holotypc, x2.

SHIKAMA

and OZA KI;

Brazilosaurus

Plr:Lte 39

Trans. hoc. Palaeont. Sue. japan. N. S .. No. ti-!. pp. 3S0-3u5, pl. .lo, Dec. 15, HJtiti

515.

NOTES ON AMMOl\liTES BRA. V.~1ISIA1\J'US D'OH.BlGNY

FROl\l THE CRETACEOUS OF FRANCE*TATSURO NIATSUNIOTO and ~viASAYUI\1 NODA**
Department of Geology. Kyushu University. Fukuoka

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-t-i.tii<'}'@JO)>E.!~H:.:;·f.: I) J: <{.-:. ·. ;j<>Ji•HJ:u)-~)'U·~;~c-'J0tt·.: 1• • ;:," 1:f.f[!;l: Subprionocyc/us nef>/zmi "(-· S. bralllleri t.: t c lJifrl L, lUI,_:, iJ ,:.::_ Subpriollocyc/us (Collignoniceratidae) l::kJ.;·j- i:, o)\>:~J!:.:.~j}jlij~·ct.:;:, ft.:fif.I'·:Jr.HI O) :;!;!llJIIIPiHn·$0_) inoceramus teshioensis ~t' (7-. : I . - P.::. 7 / J·tlil) t.l·l? 9£QS;/t,t~::. .:: n:j\i}J~ c t&:. ., ·c.
7 7 /7. i'!'fO)JJlH~.i.:'d:'~Hlt~U.:m·c ;~; ;r,t.z:. 7L1Hf~EG') t ()) :.::. >1. • ·c- li. 4-@l :t 2 1~ ~>2~111i •i• 1::. :t;,h-tt·:-ttJ<iiXi' o ;:.JJ:oo. <:1:>
L l,·~~;iilii;l:})IJfJ:~~::.:J:>f Q, *&lit 7 7 ;.-"7. ·::fi-7· =-a.::. 7 :/ J:.$0) Coilopoceras requienianum-Romaniceras deuerianum 'fidr· 0 ~·'iifu)'J::.f.it l, 7j~ilf;ft.fi<TJ 1 ·:> c ·t Q;:,. c i.J:

-c-~Q.

ri~* i'l!

~B· !!!r rn ~ z

Furthermore. we have obtained a number of specimens. from the zone of Inoceramus teshioensis, approximately Upper Turonian. in the Cretaceous depositsof the Onogawa basin. eastern Kyushu,
Japan. which are referable to the species
in question.
This species was referred to Schloenbachia by LEONHARDT (1897. p. 58. pl. 6,
fig. 3), to Gautlziericeras by GROSSOVVI~E~189-t p. 93: 1901. p. 124), to Collignoniceras ? by WRIGHT and WRIGHT (1951.
p. 38). and to Prionocyclus by BASSE andSOR:\AY (1959. p. 16. 18).
It is. thus. obviously necessary to settle the question of types, to make clear
the diagnosis and extent of variationand to know the taxonomic position ofthis species. One of us (T. l\L) haverecently had an opportunity to study
r/ORBIGC\Y's syntypes at the Institut deIntroduction
Ammonites bravaisianus was establish·
ed by n'ORBIGNY (1841, p. 308, pl. 91,
figs. 3-4) for several specimens from theCretaceous of Uchaux and Mondragon,
Vaucluse. France. On the basis of subsequent collections from the Turonianof the Uchaux basin. RO:\IAi'\ and I\lAZ·
ERAN (1913. p. 22. pl 1. figs. 13-17) redescribed the same species under thegeneric name Prionotropis. It is noticed
that the illustration of n'ORBIG:-;Y bydrawing is not quite identical with thatof RoMAi\ and MAZERAN by photographs.

* Read June 18, 1966; Received April :w
1966.

*' Masayuki :-.ioDA: Visiting Rc;;earch 1\,;sociate, july-Dec .. 1965, at the Department of
Geology. Kyushu University.

359;)l)O

Tatsuru MAT::>UMOTO and Mwwuuki 11/ovA

PalconLOlogie, l\luscum National d'llistoire Naturclle, Paris. \Ve have, furthermore, studied more carefully the specimens on Joan. This paper presents re·suits of our study. The specimens fromjapan are taken into consideration. too,
but their full description will be given
in a separate paper.
Before going further, we wish to thankDr. jacques SoRNAY. Paris. for his kind
help to this study. and Mr. C.W. WRIGHT.
London. for critical reading of the typescript. Dr. Itaru HAYA!\11, Fukuoka. has
assisted us in various ways.
Types
Twelve specimens from Uchaux ~Vau­cluse) of :lmmonites braraisianus in theCollection of n'ORBIGNY. Museum National d'l·listoire Naturelle. Paris, areavailable for study. They arc presumed
to be syntypes. They are all small.
more or less incomplete. and represented
bv internal moulds whose matrix is calc~reous sandstone, weathered to brownish. None of them agrees with n'ORBIG~Y·s illustration (1841. pl. 91, figs. 34), which must have been much restored,
perhaps synthesised from several speci·
mens and magnified (probably about
twice as large as the natural size}. AI·
though ll'ORBIGNY (1841. p. 309) indicatedthe dimensions of a representative specimen, they do not agree with those of
his illustration nor with those of anr
syntypes before us.
This measured
syntype, if it ever existed. seems to be
missing. If we temporarily put numbers
from left to right and from bottom to
top on the twelve surviving specimens
which are fixed on one tablet (figured
here in 1'1. 110, Fig. 6l. No. I specimen(Pl. .tO. J'ig. 21 is most similar to theoriginal illustration. It is. however. incompletely prcs::rvcd. without its nucleus.

and seems to represent an extreme endof the variation of the species (seell aria/ion}.
No. 2 specimen (Pl. ·10. Fig. 1) is thelargest of the twelve, comparatively better preserved, very close to what RoMAN
and MAZERAN (1913) called a typicalform. It is accordingly designated here
as the lectotype of Ammo11iles brat,aisianus n'OimiGNY. The other syntypes (ofthe twelve) are identified as probably of
the same species, except for No. 5 speci·
men. which has constrictions and nokeel nnd obviously belongs to a different
species, probably of the Puzosiinae.
Although the other specimens figured
by Rol\IAN and MAZEI~AN (1913, p. 22.
pl. I. figs. 13-17) in the collection of the
University of Lyon are not at our disposal, they are so clearly described andso finely illustrated that we can be sureof their specific identity with the lectotype. The largest of them (ROMAN and!VIAZERAN, 1913, pl. 1, fig. 17) probably
represents an adult shell. This is sup·ported by another example of a similar
size from japan.
Specific Characters
The shell is comparatively small, attaining a diameter of 60 mm or so in aprobably adult shell. It is discoidal,
having an umbilicus about 33 to 45 (?)
percent of the diameter. The whorl enlarges moderately on the average, embracing slightly less than a third toabout tHth of the inner one. It is higher
than broad. with a proportion about 10: 6
to 8. lts flanks are slightly convex or
fiat : venter narrowly arched ; umbilicalshoulder abruptly rounded; umbilicalwall low and steeply inclined.
The venter is provided with a distinct,
continuous, median keel. which may apparently look entire but is actually ser-

3Gl

Ammonites bra:vaisinnw; D'ORBJ!;NV
rate. The serrations. which corresp:>nd
in number to the ribs. c:re weak in theearly growth-stage but moderately
strong in the adult stage.
The flank, except in the very early
growth-stage (of 2-5 mm in diameten.
is ornamented with numerous. fine. somewhat prorsiradiate and gently sigmoidal
ribs, about 3:! to 56 (or perhaps morel
per whorl. which are separated by interspaces as narrow as or slightly broader
than the ribs. They are as a rule alternately long and short. The longer ribs
have a bullate. rather weak tubercle atthe umbilical shoulder. The shorter ones
normally arise from a point slightly
above and ahead of the umbilical bullaof the longer ones. but sometimes arenearly in contact with the latter, as if
branched at the bulla. Every rib isprovided with two ventrolateral tubercles. lower. (or inner) and upper (orouter). The lower one is small. at which

the rib is bent some\vhat forward. The
upper one is clavate or obliquely e!ongated. situated on a line fairly close to
the median keel. At this tubercle therib is bent strongly forward and thenweakened as it approaches the keel.
The suture consists of E. L. C,, U,,(=S). U, and l. The elements are rather
massive and not much deeply incised.
Variation
It is reasonable and natural to assume
that the lectotype and ten other syntypesof o'ORBIG'<Y represent a single variable
species. because one grades into another.
This is. furthermore. supported by the
specimens described by ROMAN and~1AZERAN. with whom we agree in admitting a considerable extent of variation of the species in question.
The following measurements illustrnte
the situation.
-----------

Specimen
Lectotype
Syntype Ko.
No. :{
No. 'I
No. 7
r>'ORBIGi"Y, p. 309* D'0RBICNY. pl. 91, f. 3-4* R.+M .. pl. I. f. 13. pl. 1. f. 14. pl. 1. f. 15. pl. 1. f. 16. pl. 1. f. 17

DiameterUmbilicus

Height15. 5(1)1·1. 3
118
15.0
12.5

5.8(0. 37)
6. 4(0. 44)
5.2(0.44)
5.2(0.35)

6. 2(0. 40)
5.0(0.3S)
4. 2(0. 36)
5. 6(0. 37)
5.3(0.42)
7 (0. :{5)

20
29 0
H.4

4. 8(0. 38)
7 (0. 35)
12. 8(0. ·14)
5.5(0.38)

clS
12.3
14.0
c60

cS (0.33)
4. 8(0. 39)
5.4(0.38)
c24 (0.4)

10. ·1(0. 36)
5.5(0.38)
6.3(0.42)
4. 7(0.38)
5. 7(0.40)
c19. 6(0. 33)

BreadthB/HRibs

3. 8(0. 3·1)

0.61
0.8
0.8

52

0. 74

34
46

.j 4(0. 30)

3. 5(0. 30)
c.4. 0(0. 27)
5 (0 ·25)
8.5(0.29)

36
42
0 " 12x3(?)
50(?)
0./1 "40-60"..

56(+rr)
50
36
36(?)
--

------* as measured from the illustration.

To know precisely the extent ofvariation we should collect sufficiently
numerous specimens from the type- andother localities. At present we are unable to do field work in France. so wedepend on the previous collections. As
far as these materials are concerned. aconsiderable number of specimens show
a form in which the whorl increasses
moderately. with its height 0.3:1 to 0.:10

362

Tatsnro MAT :UAIOTO and Mwwyuki NODA

of diameter. the umbilicus is also moderately wide. occupying about 35 to 40
percent of the diameter. and ribs are
numerous H5 to 50). comparatively fineand gently flexuous. The lectotype designated above falls within this range.
Although it has a comparatively higher
whorl, it is not at the extreme end of
the variation, since there is a form. exemplified by Ro'\!A:\ and MAZERAN's pl.
1. fig. 14, in which the whorl is higher.
more involute. more narrowly umbilicate(33 percent of diameter), and has denser
and more numerous ribs. On the other
hand syntype No. 1 (Pl. 40. Fig. 2) seems
to represent the other end of the variation. seeing that its umbilicus is fairly
wide (4·1 percent) and the ribs are coarser. less numerous and less flexuous.
D'ORBIGN r's drawing also comes close
to this extreme form. Syntype No. 3(Pl. '10, Fig. 3) is as widely umbilicate
and as thickly whorled as No. 1, but
has more numerous, denser, gently
flexuous ribs. Syntype No. 4 (Pl. 40.
Fig. 4) is as narrowly umbilicate as thelectotype but its ribs are coarser andless numerous than in the latter and aredenser and more flexuous than in No. 1.
Examples from the Onogawa Group,
japan, seem to show another aspect ofvariation in which the umbilicus is comparatively wide but the ribs are fineand numerous (Pl. 40. Figs. 7, 8J. They
could be grouped as a subspecies \\·hichis separable from the French one.
Affinities and Generic Attribution
The present species is closely allied
to Subprionocyclus neptuni (GEI~ITZ) (see.MATSU:\IOTO, 1959. p. 112; 1965, p. 52).
a world-wide Turonian species, in var·ious respects but is distinguished by its
less involute, on the ayerage more wide·ly umbilicate whorl. The former has

4S a rule regularly alternating longer
and shorter ribs which do not showsuch distinct branching as in certain
growth-stages of the !alter. S. neptzmi
has stronger but less numerous umbilicaltubercles and more frequent shorterribs. It is noted, however, that the extents of variation somewhat overlap eachother between the two species withregard to the involution of whorl, sizeof umbilicus, proportion of breadth toheight. etc. l\loreover. in some immature shells of S. neptzmi fine, numerous
ribs of alternating length occur without
branching on a part of the whorl in
addition to the paired ones. This looks
as if it foreshadows the diagnostic
character of the present species. A
specimen from Hokkaido C\IATSU~fOTO,
1965, p. 54. pl. 13. fig. 1) may exemplify
such a connecting feature. In the adolescent and adult stages of S. neptuni
ribs are coarser, much more distant andmore prorsiradiate and the tubercles andserrations are stronger, probably reflecting certain characters of the ancestral
Collignoniceras woollgari (l\IANTELL) (seeMATSUJ\101\0. 1965, p. 11).
The comparatively coarsely ribbed
variety of the present species is somewhat similar to Subprionocyclus branneri(ANDERSON) (emended by .iVIATSUl\IOTO,
1959, p. 109; 1965, p. 50). but the latterhas still coarser. stronger. more distantand less numerous ribs. in which shorter
ribs are much fewer. Its keel is morestrongly serrate and its whorl section israther rectangular, being not so slenderas that of the present species.
Some of the variable immature shellsof Collignoniceras wool/gari (l'viANTELL).
as described under subgroup D by .i\IA·
TSUl\!OTO (1965, p. 14), may resemble thepresent species. but they have more
distinctly simple. almost equally long,
Jess flexuous and more prorsiradiate

:3(13

A?WJnonifes hra·va.i.'lianus L 'ORIJII;N l'ribs. The distinction in the adolescentand adult stages is great between thetwo species.
In the present species a serration of
the keel corresponds to each rib, although the character is obliterated orapparently modified on the weatheredsurface of some specimens (including
syntypes). This species is. accordingly.
not referable to Prionocyclus in which
the serrations are fine and more numerous than the ribs, the upper ventrolateral tubercles are normally indistinct,
the lower ventrolateral tubercles areprominent and the ribs are less flexuous,
nor is it referable to Gauthiericeras in
w·hich the median keel is entire or very
finely crenate, faint side keels may develop and the ventrolateral tubercules
are not double.
To sum up ;lmmoni/es bravaisianus ismost closely allied to but distinguishedfrom Subpriollocyclus neptuni and certainly attributed to the genus Su/Jprionocyclus, as one of us (tiiATSU1\IOTO,
1965, p. 54) has already mentioned.
Occurrence
D'ORBIGNY (1811. p. 310) indicated theoccurrence of his original specimens in.. Le grcs vert superieur d'Uchaux et deMondragon, pres d'Orange (Vaucluse}."We have not examined the specimensfrom Mondragon. Gres vert d'Uchaux(Vaucluse) (n'ORBIGNY, 1841. p. 419) insoutheast France is referred to Turonian(see SORNAY, \956, p. 359).
Rm.IAN and l'viAZERA:--< (1913. p. 4) cited
liEBERT's (1875} subdivision of the Grcs
d'Uchaux and stated that the third level(from the bottom) is the principal layerfrom which came the gastropoda andcephalopoda described in their monograph, of which the species of morestratigraphical importance for correla-tion are [,ewesiceras mantelli WRIGHTand \VRICIIT (=Pachydiscus ua_iu in Ro)IA:'\ and MAZERAN, l.ewesiceras romani
SOR!\AY, 196:1) (\-VRIGHT. 1966, personalcommunication). Romaniceras deverianum(n'ORBIGNY) and Bacu/ites undulatus
n'ORBIGNY-all widespread Upper Turonian species. Subp1'ionocyrlus brauaisianus lo'OIHIIGNY) occurs in the same
fossiliferous bed along with Coi/opoceras
1equienia1111111 (o'ORRIG:->Y). This layer
is referred to Upper Turonian by BASSE
and SORNAY (1959, p. 18), too.
In Touraine. the type area of theTuronian in the northern province ofFrance. this species is recorded as occurring in the .. Craie
Bryozoaires ",Upper Turonian, along with Romaniceras
deverianum (o'ORBIGNY) and Coi/opoceras
requienianum (see GROSSOUVI<E, 1901. p.
336; BASSE and SORNAY, 1959, p. 16).
\Ve have not yet examined the specimens of this species from Touraine.
FRITSCH and Sclii.OENilACII 0892. p.
29, pl. 8. fig. 5; pl. 16. fig. :1) reported.. Ammonites bravaisianus ·· from the
Weissenberger and other beds of Bohemia, while LEONHARDT (1897, p. 58.
pl. 6, fig. 3} described "Schloenbachia
hravaisianus" from the Brongniartizoneof "Oberschlesia" (now in Poland), but
we cannot give good comments without
seeing the specimens.
In japan. as seen in the sequence of
the Upper Cretaceous of the Onogawa
basin, eastern Kyushu, probable examples of Subprionocyclus bra1'(lisianus occur in a bed above that of Subprionocyclus ncptuni (GEINITZ) and below thatof Subprionocyclus norma/is (ANDERSON).
A probable example of S. branneri has
been obtained from the same level as S.
brm;aisianus. S. neptuni occur in thezone of Inoceramus hobetsensis, approximately lower Upper Turonian. while S.
bravaisianus. S. bramzeri and S. norma/is

a

364

Talsuro MATSUMOTO and .llasa.yuki NDDA

in the zone of Inoceramus leshiocnsis.
approximately upper Upper Turonian.
A specimen described as Priono/ropis
sp. by RcniAN and MAZERAl\ (1913. p. 25.
pl. '1. fig. 18. 18a) is probably identified
with S. branneri A:\DERSO:'\ [=S. crislatus (BII.LINCHURST)]. It came from thevery layer where S. hrcmaisianus occurred more abundantly. We would liketo know whether the same succession of
the species of Subprionocyclus as that injapan is maintained or not in the Turonian sequence of France and various
other areas in the world.
P. S. C. W. WRIGHT has recently given
us information that S. neptuni, S. hitcltinensis, S. branneri and S. norma/is all
occur together at Hitch Wood in England.
He does not agree wilh us to distinguishS. brcwaisianus from S. nefJ/illli (ielter
Sept. 9. 1966).

References Cited
BAssE de Menorval. E. and SOR:-IAY, Jacques(1959) : Gcneralites sur les faunes d•ammonites du Cretace superieur fran~ais.
84'' Congr. Soc. Savantes. Dijon, 1959, 726.
FRITSCH. A. and SCilLOENBACH, U. (1872) :Cephalopodeu der bohmischen Kreideformalion. 51 p., 16 pis., Prague.
GROSSOU\RE, Albert DE (189,1) : Recherches
sur Ia Craie supericure. II. Pah!ontologie.
Les ammonites de Ia Craie superieure.
Mem. Carte Geol. Det. France [1893], 264
p., 39 pis.
- - (1901) : Recherches sur Ia Craie superieure. I. Stratigraphie generale, 1 & 2.
Ibid. [1901], 1013 p.
Ht~BERT. E. and TOUCAS, A. (1875) : Materiaux pour servir a Ia description du terrain cretace superieur en France. Description du Bassin d'Uchaux. Ann. Sci.
geol.. 6. 1-132, pis. 3-6.
LEO II AIWT. Richard (1897) : Die Fauna dcr

Explanation of Plate 40
Fig~.

1-8. Subprionocyclus brarmisianus (t>'ORBIC.!'\Y) ................................ Page 360
1. Lectotype, from Dchaux (Vaucluse). France. in D'0RBIGNY'S collection. Two lateral(a, b), ventral (c) and frontal (d) views·, x2. 5. The serrations of the ventral keel
seem to have been in part modified by weathering.
2. Syntype No. I, a widely umbilicate and coarsely ribbed variety. Lateral (a) andfrontal (b) \·iews, x 2. 5.
3. Syntypc No. 3. an example with a similarly wide umbilicus but denser ribs. Lateralview, x2.5.
•1. Syntype No. 4. an example with a narrow umbilicus and dense but coarse ribs. Lateral view. x 2. 5.
5. Syntype No. 7. an example which is fairly similarly to but smaller than the loctotype.
Lateral view, x 2. 5.
6. A set of syntypes of "Schloenbachia bravaisiana (o'ORBIGKY) ,. in D'ORBJG:-IY's collection, Museum National d'Histoirc Naturelle, Paris. No. 2 is the lectotype designatedin this paper. No. 5 should be excluded from the present species. Natural size.
7. GK. H 6975. an example from Nakahaji Shale, Onogawa Group (Coli. l\-1. NooA). Lat·era! view, x 1. 5.
8. GK. H 6976, a smaller example from Nakahaji Shale. Onogawa Group (Coli. M. NooA).
Lateral view, x 2. The last two specimens arc preserved in the Type Specimen
l~oom. Department of Geology. Kyushu University.

MATSUMOTO and NQD.A:

Ammonites bravaisianus

Plate 40

2b

5

6

8

.·lmmoniles !J·nwaigianns D'ORBIGNJ'
Kreideformation in Obserschle:;ien. Pal·
aeontographica, 49. 11-70, pis. 3-6.
MATSUMOTO, Tatsuro (1959): Upper Cretaceous Ammonites of California. Part II.
Mem. Fac. Sci .. /(yushu Univ .. CDJ. G1·ol ..
Special Vol. I. 1-172. pis. 1-41.
- - (1%5): A monograph of the Collignoni·
ceratidae from llokkaido. Pan I. A/em.
fac. Sci .. f(yuslw Uni11 .. [D]. Ceo/., 16.
(1), 1-80, pis. 1-18.
ORBIGNY. Alcide n· (18<10-42): Pa/ionlologie
Francaise, Terrains Cretace, 1. 662 p .. 1·18
pis. (p. 1-120 . 1840; p. 121-430. 18H ; p.
431-662, 1842). Paris.
Ro:-.tAK. Frederic and M.\ZERA:", Pierre(1913) : Monographic paleontologique deIa faune du Tumnien du bassin d'Uchaux

1\akaha ji

et de ses dependances. Arch. Musee Hist.
Nat. Lyon, 12. 1-137. pis. 1-11.
SoR:->AY, Jacques (Ed.] (1956):
Cretace.
Lexique Straligr. Intern .. 1. Europe, (4ab). •IO:l p., Paris.
- - (196,1) : Sur un Lewrsireras nouveau duTuronien d·Uchaux (V:wcluse).
;\1111.

Pa/ionl.. 50. (2). 1-7.
WRIGHT, C. W. and WRICIIT. E. V. (1951): A
survey of the fossil Cephalopoda of the
Chalk of Great Britain.
Primarily anomcnclatorial revison of Daniel SHARPE'~" Description of the fossil remains ofmollusca found in the Chalk of England.
Part I Cephalopoda (1853-1857). Palaeon·
to gr. Soc. (1950). 40 p.

Onogawa

PROCEEDINGS OF THE PALAEONTOLOGICAL SOCIETY
OF JAPANs;;t,:;'J'lJl.7"(;~i:fPl4fi'li91]~1t,

19661f.9 r12-1

f::l (JJ l-klll:k'f;j[/;tJJif:;}Y,~,Yf~~:~J;'-•L" l~lfll\~lt
t:.. f!''· 9 25 A Ef) ;.::ii\!'!Iii!m;<ro:>~'f:IJ~iinc:Hi:!:>*1.t:., (~Jm~ 13 iS)

Sasanohara. Southern Kilakami Mountains
Yasuji SAITO'i'-fiii@Hll <D~J$tl\%ii~Rl'll:¥.11ill-~ · tllll:JHii'Hf. · iflit~"!/t)\(RI>
iJJn !',Hlli1J:ItirlJr.l' (rrJlllrJI'l"Ji 1!1:).00 ~;I' }.'.!; J:~NI ll+ o:> llH~.fff ~"L!h i.: J: {5 5Hii'

K.f.W;J-'JJ · itJit~"!/t;.!..:P-1>
l-Tili110)Jf'lfrlll:ltf(~ftT:iK.!liiUiJ-1}} • liliitJ!i~"k!'!~fk llll1,~0)'1~·1lllifi~ll+ (I& WI •l•lfrill:)[¥.W;J-~

w*il['filA :;$:~l.jfiJti!li: Gobius lt.Ti't'~·*fi'(I\'Jl'!•ilt:;$:Ji.';J: i) 3e!i!.~hf_: Halobiidae (fl::m'.:)T=IMii~ · 'l·fl't!f-· · d·il§~=

A Lower Carboniferous Ammonoid from

[~

n~

rrJ

!JH/t · i\!~lli!l&O~W.*illll:tlo:>lti!tJi!.'.f:~Pit'

Iii! W.J-9}

SYSTEMATIC INDEX(New Series No. 57-No. 6-1).

PALAEOZOOLOGY
Protozoa
Page
HIGUCHI, Yu: On some new species of fossil Foraminifera from the cores in the gas field
in Chiba Prefecture, Japan.....................................................
HuANG, Tunyow: .4sanonella shojii n. gen .. n. sp. (Family Discorbidae) from Tokuno·
shima, Kagoshima Prefecture. Japan . .. .. .. .. .. .. .. .. .. .. .. . . .. .. .. .. .. .. .. .. .. .. .. .. .
Planktonic Foraminifera from the Somachi formation. Kikai-jima. Kagoshima
Prefecture, Japan . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
MATSUMARU, Kuniteru: Fusulinids from the Funafuseyama limestone In Yamagata.gun,
Gifu Prefecture, Japan................................................................
SADA. Kimiyoshi and YoKoYA~tA, Tsuruo: Upper Permian fusulinids from the Taishaku
limestone in West Japan ..............................................................
UJIIE, Hiroshi: Shell structure of japanese smaller Foraminifera. Part l, Ammonia
tochigiensis (Uc111o)..................................................................
- - - - : Shell sttuclure of japanese Slllallcr Foraminifera Part 2. l'ararota/ia nipponica(ASAI\0, 1936). . .
. . ' .. ' ....... ' .. ' .. ' .......................... '' ' ...... ' . . . . . . . . .

178
166
217
338
303
156
HJl

Bryozoa
SAKAGA~tl. Sumio:

Revision of the Takagami conglomerate, Cho~hi Peninsula, and des·criptions of the Permian Bryozoa from its limestone pebbles ......................... .

Brachiopoda
IIATAl. Kotora and IIAY ASAKA, Shozo: Two fossil species of Discinisca (Brachiopoda)
from North Honshu, japan . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .

172~1ollusca

AoKt . Naoaki: Molluscan fossils from the Nobori formation Shikoku. japan . . . . . . . . . . . . 251!lAsE. Kotaro: Some marine mollusca from the alluvial deposits of Yamashita-cho, Watari·gun, Miyagi Pretecture . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 67
IIATAI. Kotora and IMAtzu~tl, Rikizo: Discovery of marine shells from the Kunohe terrace deposits, Kuji City. Iwate Prefecture............................................
28
HAYA!\11, ltaru and NAKAI. !sao: On a lower Cretaceous pelecypod, "Cyrena" naumanni.
from japan . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1].1----and ICliiKAWA. Takeo: Occurrence of Nippononaia ryosekiana from the Sanchu
area, Japan . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 145
HAYASAKA. Ichiro: Some cephalopods in the Permian faunule of Takakura·yama. Fuku·
shima Prefecture, Japan (with a note on the geology of the di!'trict by Ichiro YANAGI·SAWA and Mamoru NE~toTo)
8

MASUDA. Koichiro: Molluscan fauna of the Higashi-Innai formation of Nato Peninsula,
Japan-f. A general consideration of the fauna . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
· - - - · Molluscan fauna of the Higashi-lnnai formation of Noto Peninsula, Japan-11;
Remarks on molluscan assemblage and description of species . . . . . . . . . . . . . . . . . . . . . . . .
MATSill\IOTO, Tatsuro: Notes on .4.mmonilts bourgeoisi D'0Rn!G;>;Y from the upper Crc·
taceous of France . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
· - - - - · ISIIIKAWA. 1-Iideo and YAMAI<IlCIII. Shiro: A l'vh-sozoic ammonite from AmarniOshima ..............................................................................
Notes on Ammonites jlaccidicos/a Rn~IEI~ from the Cretaceous of Texas . . . . . .
- - - : and No!>A, Masayuki: Noks on Ammonites brllllllisianus D'ORiliCI'I\' from theCretaceous of France . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
NAGASAWA, Joji: A gigantic scallop from the sea bottom off the Danjo Islands. Nagasaki
Prefecture, Japan . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
- - - - : The significance of the magnesium and calcium ratios of fossil shells ofUmbonium costa/11m (KIF.:"ER) from the deposits since the Pliocene in the southern
Kanto region. Japan . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
NAKAI, Isao and HADA, Shigeki: Discovery of Aptian nmmonites from the Shimanto
terrain. western Shikoku. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
NonA, Hiroshi: Some fossil !lnadara from Southwest Japan . . . . . . . . . . . . . . . . . . . . . . . . . . . .
OUATA. lkuwo: Allometry of Reesidilt•s minimus. a Cretaceous ammonite species........

261
317
201

234
294
359

110

207

242
92
3!!

Arthropoda
KOHAYASIII, Teiichi and HA~It\J>A. Tnkashi: An occurrence of Scutellum in the Silurianof Shikoku Island . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .

74

Echinodermata
MoRISIIITA, Akira: New species of Sa/enia from the Miocene of Japan (On some fossil
echinoids of Japan) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
TA:-->AKA, Keisaku: Cretaceous echinoids from the Sanchu graben, central Japan . . . . . . . .

64
126

Vertebrate
II ATAl, Kotora, Mllll,\TA, Masafumi and MAsUDA, Koichiro: A sting ray and eagle ray fromthe Tatsunokuchi formation (Pliocene) in Sendai City, Miyagi Prefecture, Japan . . . .
34
lluA~G. Tun yow: A new species of a whale tympanic bone from Taiwan. China........
18:~
SIIIKA\IA, Tokio and OzAKI. Hiroshi: On a reptilian skeleton from the Palaeozoic formation of San Paulo, Brazil .
351
00

00

..

00

00

00

..

00

00

..

00

..........

00

....

00

..

00

..

00

00

00

00

..

..

•

Palaeobotany
E:"\DO, Seido:

On the genus Tilia from the Woodwardia zone of IIokkaido. with description of two new species..............................................................

188

Miscellanea!Go, Hisayoshi and KOIKE, Toshio: Carboniferous conodonts from Yobara, Akiyoshi
limestone, Japan (Studies of Asiatic conodonts, part II) ... , . . . . . . . . . . . . . . . . . . . . . . . . . .

83

1NilEX OF FAMiLiES, GENERA AND SPECiES
Notes: \Vords listed below arc name~ of families. gcncr:1 and ~pecie;;, which are either
described or illustrated in the Tram;actions and Proceedings of the Palaeontological Societyof japan. New Series. No;;. 57-6-1; word::; in heavy type arc names of new genera and species.

A
70
Acmaea pallida
Aequ i peel en mal sutlagiellsis ........... . 323
Agalhice,·as cf. suessi ................. .
19
Ammonites bourgeoisi ................. . 202
Amussiopec/en j1raesignis ............. . 255
Anadara (Snrpharca) suzukii ......... . 100- - - ( - - - -?) ill'ashibaraensis ... . 10,1
makiyamai ................... . 280- - - - (Scapha rca.') s/rizuokae11sis ... . 103- - - - suzukii. . . . . . . . . . . . . . . . . . . . . . . . 254(Scapltarca) lakaoensis. . . . . . . . 101( Tosan·a) to.~aensis. . . . . . . . . . . . 105- - - - (Scaplwrca) lricenicos/a . . . . . . 102( - - - ? ) sp. . . . . . . . .. . . . . . . . 101- - - - sp. . . . . . . . . . . . . . . . . . . . . . . . . . . . . 25-1
Anisocorbula ohiroi . . . . . . . . . . . . . . . . . . . . 329
Anomia chinensis . . . . . . . . . . . . . . . . . . . . . .
68
11Phelas/er serolinus.................... 133
Archilec/onica kurodae . . . . . . . . . . . . . . . . 336.4sallollell a . . . . . . . . . . . . . . . . . . . . . . . . . . . . Hi7
167- - - slwjii

Chinocardium californiense . . . . . . . . . . . .
Chlamys nipponensis . . . . . .. . . . . . . . . . . .
Chuse11ella sp.. . . . . . . . . . . . . . . . . . . . . . . . . .
Cleme11/ia jajJonica . . . . . . . . . . . . . . . . . . . .
Collignoniceras (Selwynoceras) sp. nov.
(?) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
Collisella sp. . . . . . . . . . . . . . . . . . . . . . . . . . .
Crassa/ellitt•s suyamensis . . . . . . . . . . . . . .
Clena lwlaii. . . . . . . . . . . . . . . . . . . . . . . . . . . .
Cycliua (Cyclinorbis) lunula/a ... .......

68
68
313
334
235
336
334
325
326

D
Dasyba/us d. alwjei....................
Disci11isca..............................
- - - miuagiensis . . . . . . . . . . . . . . . . . .
- - - semlaiensis. . . . . . . . . . . . . . . . . . . .

34
174
175
174

E
Eponides CIHCITWi . . . . . . . . . . . . . . . . . . . . . . . .
Euche/us tWiflf'fiHiiJ . . . . . . . . . . . . . . . . . . . .

180
330

F
B
Babylonia cla/rl ....................... .
Ba/aelloptera ......................... .
Balaenoplera lairmnica ............... .
Bolliella adamsi . ...................... .
BrazilosauruiJ smrprruloensis . ... .

256
183
184
226
351

c
Calliosloma olcmiensis . ................ .
Candeina nil ida ....................... .
Carcharhi11us gangelicus ............. .
Cardilia loyamacnsis ................. .
Chelomceras (Citeloniceras) sp. aff.
minimum ........................... .
( - - · -) shimizui . .......... .

331

226
186
331

Felauiella u.~/aFissulina malobai ..................... .
Fistulipora sp. . ...................... .
Fulgoraria cf. cance/lata

69
180

6

256

G
Ga/eocerdo adrmcus
Glans mwmiae ....................... .
Globigeri11a aper/ura ................. .
- - - - bu/loides ..................... .
- - - decoraperla ................. .
- diJ!lostoma ................... .
---folia/a ....................... .
iiiWIIIJ!Ia.................. .
- - - jmdrydcrma ................. .

186
325

226
226
226
226
226
226

227

Jnde~~ o/ Jtamili.£~s. (;enent

227
227
G/obigerinila glulina/a ............... . 2:!.7- - - - lwmilis ....... ................ . 227
uvula ....................... . 227
227
Globigerinoiril>s conglobatus ........ .
- - - - cyclostomus ................. . 227- - - - elongatus . .................... . 227
immaturus ................... . 227
obliqzws ..................... . 227- - - - nwer . ........................ . 228- - - - sacculifer ................... . 228- - - - trilobzzs ..................... . 22B
Globorotalia crassaformis ............. . 22B- - - - hirsuta ....................... . 22B- - - - menardii menardii ........... . 228
menardii multicamera/a ..... . 22::!
menardii /umida ............. . 229/osaensis ..................... . 229
trrnzcatulinoides ............. . 22\1
G/oboquudrina duterlrei ............... . 223
Glycymais nakamurai ................. . 25·1
- - - - veil ita ....................... . 69
- - - - tJrtinqdo/ia---

tliOOtfi . . . . . . . . . . . . . . . . . . . . . . . .

69••.•..•...••.....
Gnatlwdus commutalus naga/oensis ... . 89
- - - - ofJimus ....................... . 89
Gomphina melanac>gis ................. . 69
Granulifusus dual is ................... . 256
Gyroidiua kn:wmense ................. . 181- " - - - .VI!SStll'/ISiS

H
329

and. Spel'·ies

Leuwsyriux coreanica . . . . . . . . . . . . . . . . . . 256
Leukoma itoigawae . ................. 281, 334
Ligonodina hanaii. . . . . . . . . . . .. . . . . . . . . . 86
Limo psis cumingii...................... 254- - - cf. forskalii . . . . . . . . . . . . . . . . . . 254
Lillzolhanwium (?) sp...................
71
Lillorinopsis miodelica/ula .......... 281, 336
Lonchodina akiyoshie11sis . . . . . . . . . . . . . . 87
- - - ' sp. ........... .... ...........
Lonchodus sp· . . . . . . . . . . . . . . . . . . . . . . . . . .

Lyria mizuhonica . . . . . . . . . . . . . . . . . . . . . .

,\/acoma /okyoensis ................... .
Medlicollidae' . ........................ .
Mercenaria stimpsoni ................. .
Miogypsina kotoi ..................... .
Mizuhopeclen cf. yessoensis ........... .
kumagui ..................... .
Modiolus wunizukiensis ............... .
Monilea lwmcu/ae ..................... .
- - - - yoshioi . ...................... .
Afyadora suzue11Ris ................... .
Myliobatis semlaicus

86
86
256

69

16
69
283
31
Ill

324
332
:i:H
324
35

N
Nei/onel/a coix . . . . . . . . . . . . . . . . . . . . . . . . 254
Nep/zwea arthrilica . . . . . . . . . . . . . . . . . . . . 70
Nerita ishiclae . . . . . . . . . . . . . . . . . . . . . . . . 335
Nez•erila didyma. . . . . . . . . . . . . . . . . . . . . . . . 70
Nippononaia . . . . . . . . . . . . . . . . . . . . . . . . . . 150- - - - ryosekiana . . . . . . . . . . . . . . . . . . . . 147]liou·akites )1accidicostus................ 296

Haliotis twioellsis ..................... .
Harpa losu ........................... .
Jlasligerina siphonifera ............... .
lleteraster d. nexilis ................. .
- - - - yuasensis . .................... .
Heteromacoma irus . . . . . . . . . . . . . . . . . . . .
/lindeodella sakagamii. . . . . . . . . . . . . . . . . .
1/indeodus impcrfec/us..................
Howakites. . . . . . . . . . . . . . . . . . . . . . . . . . . . . .

136

0

69
8ii
87
294

Octubronteus (?) sp.. . . . . . . . . . . . . . . . . . . . 79
Operculina comp/anala jCipvnica........ 283
Orbulina universa. . . . . . . . . . . . . . . . . . . . . . 229
Ostrea denselamel/osa . . . . . . . . . . . . . . . . . . 69
- - - - gigas. . . . . . . . . . . . . . . . . . . . . . . . . . 69
Ozarkodina orientale . . . . . . . . . . . . . . . . . . 87

Isurus lwstalis

186

p

257
229
135

L
Lacuna juponicct . . . . . . . . . . . . . . . . . . . . . . 336
5
Leioc/ema? c/w.~hie11sis ............... .

Paphia (s. s.) eugh/•la ohil'tli . . . . . . . . . . 326(s. S.) 81/ZI/I'IISiH . , . . . . . . . . . . . . 327[Jaraceltites aff. e/egans ... ,............
17
Parafusulina fruwfuHt'IIHis ............. .

lnde:t o/ l•'mwil·it;s, Genera. wnd Species
Pararotalia ........................... .
nipponica ... .................. .
Pecten cf. albicans ................... .
Perip/oma sp.
Pet rico/a dit·crf.iCIIS .............. .
Phlyctiderma japonica ................. .
Pilar itoi ............................. .
Placopecten protomollitus ............. .
Polytropa heysea111J.............. .
Popanoceras sp ........................ .
Propinacoceras sp. . .................. .
Protocyprina ......................... .
naumanni ................... .
sp............................ .

Protorotella shukuborensis ............. .
Protothaca englypta ................... .
Psendogastrioceras ................... .

195

192
2\l

2!1S
(j9

70
334
33-1

70
2:i
15
115

117
H!J
;{:Hi

70
18

18
Pseudojacobites ....................... . 2D8~

sp.

. ........................ .

- - - - te.\'C/111111 ..................... .
Pseudoliotia sp ........................ .
Pulleniatina obliquiloculata ........... .
Purpura bronni ....................... .
Pygurus (Pygurus) complmwlus . ...... .

2!l!l
3:l6

229

70
128

371

Spondy/us cruentus ................... .
Stacheoceras aff. griinu•aldti . .......... .
Strongy/ocent,·otus (?) sp .............. .
Subbotina jalconensis ................. .
Su/coretepora sp ...................... .
Synpri011iodina sp.

70
21
71

229
6

85

T
Tainoccras abukumaense ............. .
- - - - aff. unklesbayi ............... .
Terebra bifrons ....................... .
Thsarropeltella ....................... .

13

Thysanopeltidae ....................... .
Thysanopeltis ......................... .
Tilia eojaponica ..................... .
- - - japonica ..................... .
- - - - protojapunica
Tosarca ............................... .
Toxaster HWichuensiH . . . . .
. ....... .
Tugali noloensis ..................... .
Turbo ozart•ai ......................... .
- - - - (Marmorostoma) ozawai ..... .
Turborotalia acostaensis ............... .
Turritella perlerebra ................. .
T.vloncwtilus permicus..................

75

14
70
76

76
188
189
189
104
131
330

281
333
230
255
13

R
Raeta sp.
Reesidites minim us . . . . . . . . . . . . . . . . . . . .
Rissoina mwmic1e ...

334
39

Umbtmium costatum ............ .

207

v

s
Salerria nippunica .................... .
Sa.ridomus purpura/us ................. .
Saccella saikait'llsis ................... .
Swtel/um japo11icum ................. .
- - - (?) sp ........................ .
Seluynoceras . . . . . . . . . . . . . . . . . . . . . . . . . .
Siphonalia cf. sikokuana . . . . . . . . . . . . . .
Siratoria siratorie11sis. . . . . . . . . . . . . . . . . .
So/ariel/a sp. . . . . . . . . . . . . . . . . . . . . . . . . .
Spathognathodus minutus . . . . . . . . . . . . . .
Sphaeroidinella dehiscens . . . . . . . . . . . . . .
Spiloplectammia kujukurieuse. . . . . . . . . .
Spirotropis subdeclitJis..................
Spisu/a sachalinensis . . . . . . . . . . . . . . . . . .
- - - sachalirrensis . . . . . . . . . . . . . . . . . .

u

&I

70
:122

77
78
235
2!16
3:H

:B6
88

229
180
2!17
:Jl

70

Venericardia panda ................... .
Venerupis (Amygdala) takagii ....... .
Ventrico/oidea foveola/a ............... .
.. Venus" ,;p .......................... .

255
328
255
334

w
IF aagenoceras cf. dieneri richardsoni ..
Washitas/er (?) !<p ....................•
Weberopdtis

23
138

76

y
311
Yabeina elongata
minula . ...................... . 309
multiseptata multiseptata ..... . 304
shiraiwensis ......... . 307

a -lt~ -li- i.~ ~~-~·ic.f
Transactions and Proceedingsof thePalaeontological Society of Japan
New ~eries

No. 57 ,..._No. (i.J
1%5 ,..._ HlGG

l'alacontological Society of japan

The heading in japanese commemorates the handwriting of Prof. 1\1. YoKOYAMA,
father of japanese Palaeontology. who was Professor of Stratigraphy and Palaeontology at the Geological Institute, Imperial University of Tokyo.
The fossil on the front page is Pecten talwlwshii YoKOYAI\!A. 1930.

CONTENTS

Number 57 tPublished April 30. 1965;
Transactions
Article

Page

Plate

SAKAGA:O.IT, Sumio: Revision of the Takagami conglomerate.
Choshi Peninsula. and descriptions of the Permian bryozoa fromits limestone pebbles . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1-7:183. llAYASAKA. lchiro: Some cephalopods in the Permian faunule of
Takakura-yama. Fukushima Prefecture. japan (with a note on
the geology of the district by lchiro YA:-\AGISAWA and ;'vlamoru
NEI\IOTO)........................................................ 8-27
484. HAT AI, Kotora and IMAIZtnii. l~ikizo: Discovery of marine shells
from the Kunohe terrace deposits. Kuji City. lwate Prefecture .. 28-33
485. HAT AI. Kotora, 1\luRATA, \lasafumi and !viASUilA. Koichiro: A
sting ray and eagle ray from the Tatsunokuchi formation (Pliocene) in Sendai City. l\liyagi Prefecture. japan .................. 34-37
Proceedings ...................... 37-38

2-3

482.

Number 58 (Published june 30. 1965)
Transactions
0BATA. Ikuwo: Allometry of Nn:sidites minim us. a Cretaceousammonite species .............................................. 39-63
487. MORISHITA. Akira: Ne\v species of Salenia from the \liocene ofJapan (On some fossil echinoids of japan) ...................... 6'1-66
1188.
I-lASE. Kotaro: Some !\Iarine Mollusca from the alluvial deposits
of Yamashita-cho, Watari-gun. Miyagi Prefecture .............. 67-73
489. KoBAYASHI. Teiichi and HAMADA. Takashi: An occurrence of
Scutellum in the Silurian of Shikoku Island .................... 74-81
:186.

4-5

6
7

Number 59 (Published September 30. 1965)
Transactions
490.

lGo, Hisayoshi and KoiKE. Toshio: Carboniferous conodonts from
Yobara. Akiyoshi limestone. japan tStudies of Asiatic conodonts.
Part II) ........................................................ 83-91

8-9

NonA. Hiroshi: Some fossil :lnadara from Southwest Japan .... 92-109
NAGi\SAWA . .Joji: A gigantic scallop from the sea bottom off
off the Danjo Islands. Nagasaki Prefecture, Japan .............. 110-113-193. 1-lAYAMI. ltaru and NAKAI. !sao: On a lower Cretaceous pelecypod,"Cyrena" nauma1111i, from japan ................................ 11·1-125
49·1. TANAKA, Keisaku: Cretaceous echinoids from the Sanchu graben,central japan .................................................. 126-142
Proceedings . . . . . . . . . . . . . . . . . . . . . . . . . . . .
143

491.
·192.

10-11
12
13-14
15-16

Number 60 (Published December 20. 1965)
Transactions
495.

l-IAYAMI, Itaru and IcHIKAWA, Takeo: Occurrence of Nippononaia
rynsekiana from the Sanchu area, japan ........................ 145-155·196.

UJIIE, Hiroshi: Shell structure of Japanese smaller Foraminifera.
Part l. Ammonia tochigiensi:; (UCHIO, 1951) ........ ~ ............. 156-165 18-19
HUANG, Tunyow:
Asannnd/a slwjii n. gen.. n. sp. (Family
Discorbidae) from Tokunoshima. Kagoshima Prefecture, japan .. 166-171
20
HATAI, Kotora and HAYASAKA, Shozo: Two fossil species of
1Jisci11isca (Brachiopoda) from North Honshu, japan ............ 172-177
HIGUCHI, Yu: On some new species of fossil Foraminifera fromthe cores in the gas field in Chiba Prefecture. japan ............ 178-181
21
Proceedings . . . . . . . . . . . . . . .
182

497.
498.
499.

17

Number 61 !,Published April 20. 1966)
Transactions
500.
501.
502.
503.
504.

HuANC. Tunyow: A new species of a whale tympanic bone from
Taiwan, China .................................................. 183-187
ENno. Seido: On the genus Tilia from the IVoodwardia zone of
Ilokkaido. with description of two new species .................. 188-190
UJIIE. Hiroshi: Shell structure of japanese smaller Foraminifera.
Part 2. Pararotalza nipponica (ASANO. 1936) .................... 191-200
rviATSUl\IOTO, Tatsuro: Notes on Ammonites bourgeoisi o'ORBIGNY
from the upper Cretaceous of France .......................... 201-206
NAGASAWA, Joji: The significance fo the magnesium and calcium
ratios of fossil shells of Umbonizmz costatum IKIENEH) from thedeposits since the Pliocene in the southern Kauto region, japan .... 207-213
Proceedings ...................... ZH-215

22
23

24-25
26

iii

Number 62 (Published june 30. 1966)
Transactions
505.

HuANG, Tunyow: Planktonic Foraminifera from the Somachi
formation, Kikai-jima, Kagoshima Prefecture, japan ............ 217-233 27--28
506. !'v!ATSUJ\IOTO, Tntsuro, ISIIIKAWA. 1-lideo and YAMAKUCIII. Shiro:
A rvlesowic Ammonite from Amami-Oshima .................... 23-l-241
507. NAKAI, !sao and I-I ADA. Shigeki: Discovery of Aptian ammonites
from the Shimanto Terrain. Western Shikoku .................. 242-250 29-30
508. AoKI. Naoaki: Molluscan fossils from the Nobori formation,
Shikoku. japan .................................................. 251-259
31
Number 63 (Published September 30, 1966)
Transactions
509.

510.
511.

l'viASUDA, Koichiro: \lolluscan fauna of the Higashi-lnnai formation of Noto Peninsula, Japan-I. A general consideration of thefauna .......................................................... 261-293
l'vlATSUMoro. Tatsuro: Notes on Ammonite's .flaccidicosta RoMER
from the Cretaceous of Texas .................................. 29:1-302
32
SADA. Kimiyoshi and YoKOYAMA, Tsuruo: Upper Permian fusulinids from the Taishaku limestone in West .Japan .............. 303-315 33-3·1
Proceedings . . . . . . . . . . . . . . . . . . . . . .
316
Number 64 !Published December 15, 1966)
Transactions

512.

513.

MASUDA. Koichiro: !vlolluscan fauna of the Higashi-lnnai formation of Noto Peninsula. japan-!!; Remarks on molluscan assemblage and description of species ............................ 317-337 35-36

MATSUMARU, Kuniteru: Fusulinids from the Funafuseyama
limestone in Yamagata-gun, Gifu Prefecture. japan ............ 338-350
37514. SlllKA:\IA, Tokio and OzAKI, Hiroshi: On a reptilian skeleton
from the Palaeozoic formation of San Paulo. Brazil . . . . . . . ... 351-358 38-39
515. MATSUMOTO. Tatsuro and NonA, Masayuki: Notes on Ammonites bravaisianus n'ORBIGNY from the Cretaceous of France .... 359-365
40
Proceedings . . . . . . . . . . . . . . . . . . . . . .
366
Systematic Index ................ 367-368
Index of Families.
Genera and Species ........ 369-371

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Inti. Symposium on Jurassic Stratigraphy
July 17-27. Luxemburg: Dr. P. L. MAtiJEt:GE, 141 ave. Carnot, St-Max, France.
Inti. Sedimentological CongressAug. 11--15. Reading and Edinburgh: Sedimentology Re:;earch Lab., Geol. Dept., theUniversity. Whiteknights Park, Reading, England.
European Colloquium of Micropaleontology
Sept. 2-9, Poland: Polska Academia Nauk, Zklad Paleozoologii, AI. Zwirki i Wigury 6,
Warszawa, Poland.
Inti. Symposium on De\·onian
Sept. 6-8, Calgary, Alberta: Alberta Soc. Petroleum Geulogi:;ts, POB 53, Calgary,Alberta, Canada.
Inti. Congress of Carboniferous Stratigraphy and Geology
Sept. 11-16. Shefield, England: Dr. A. W. WooDLAND, Secretary-general, Geologi..:alSurvey Office, Ring Road Halton, Leeds 15, England.
Congress on Mediterranean Neogene Stratigraphy
Sept. 19-30, Bologna: Prof. R. SELLL Institute di Geologia e Paleontologia, univ. di
Bologna, Via Zamboni 63, Bologna, Italy.
Symposium on Gondowana Stratigraphy
Sept. 26-0ct. 15, Buenos Aires: Dr. E. R. PLUMSTEAD, Dept. of Geology, Univ. of
the Witwatcrsland, Johannesburg, South Africa.
Inti. Conference on Planktonic Mic1·ofossils
Sept. 27-0ct. 3, Geneva: Univ. & Museum of Nat. Hist. of Geneva, Plank toni\:
Conference, Box. 160, 1211 Geneva 4, Switzerland.
Meetings of the International Palaeontological Union, 1966. ~p: Ill•'! A ·JLit~£:Wj0;;&rl• 9 Jl 1 U
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Regulations for Publication in Transactions and Proceedings of
the Palaeontological Society of Japan(Jan. 15, 1963)
1. Manuscripts considered for publication should have been read at the General Meeting or
the Ordinarp meeting of the Palaeontological Society of japan.
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