fJ -lt"~±.~~~~.:k· t~tTransactions and Proceedingsof thePalaeontological Society of Japan
New Series

No. 6

Palaeontological Society of JapanMay 31, 1952

CONTENTS
TRANSACTIONS
213. On Some Species of Cyclina from Japan and Korea ... Yasuhiko KAMADA 167
214.

Two New Triassic Estherians from Province of Nagato in West Japan.................. ··························· ········· ······ ................. . Tciichi KoBAYASHI 175

215.

A Miocene Crab, l/yas tsuclzidai n. sp. from the Wakkanai Formationof Teshio Province, Hokkaido ....................................... Rikizo IMAIZU!\11 179

216. Descriptions of the Paleogene Molluscs from Sakito, Nagasaki Prefec-

ture, Japan ............................................................... Atsuyuki MizuNo 185
Publications received............................................................................

lH

Memorial to Thomas Wayland V AUGIIAN ... .. . ... .. . .. . .. . .. . .. . .. . .. . ... ... .. . ... ... ...

184

PROCEEDINGS ............................................... .

194

President:

Hisakatsu YABE

Councillors:

Kiyoshi AsANo, Ryuji EN no,

ENoo, Haruyoshi

FuJIMoTo,(Busii~ess

Shoshiroand Accountant), Seido

HANZAWA, lchiro

HAYASAKA,

Koichiro kHII\AWA, Teiichi KoRAYASIII (Editor), Jiro MAI\IYAMA, Tatsuro
MATSUMoTo, Masao MlNATo, Tsunctcru

OINOMII\ADo(Publication),

Tokio SHIKAMA, Fuyuji TAKA! (General Affairs), Hisakatsu YABE.

All Communications relating to this Journal should be addressed to the
PALAEONTOLOGICAL SOCIETY OF JAPAN
Geological Institute, Faculty of Science, University of Tokyo, Japan

Trans. Proc. Palawnt. Soc. Japan. N.S., No. 6, pp. 167-173, pl. 15, May 30, 1952.

213. ON SOME SPECIES OF CYCLINA FROMJAPAN AND KOREN)Y ASUHIKO KAMADA
Tohoku University, Sendai

a*JlrFttlUim:::.=:<'J Cyc!itiO 1"::-::>v•'L: ffl.ft~ Cyclina orietllalis (SOWERRY) l-{l::Pfl
C. japonicl KAMADA n. sp. )).rF C. asagaiensis KAMADA n. sp. a:flC.l'i l..., Cyclitwrbis
MAKIYAMA ~~IJli! L...'L.jlj.~~Lt;: 0 C.joponica JkrF C.ltmulata 17)ti~:%\H:~~tl:tc,j,lj:J!1:, :tl?)jl;.Plfl <'Jtf'jt;&· !':>, !fllifiltl?)lf~h ·iil!.Y> 'Ltfv•lfit:: flth- '"Cv• :tc·~;ll,EJ !':> »· t:: l'.l: !J, .?OfHfU: Vicarya~ !J,tri&r.tll@.;aii:l?);f, o ]JJli}'IJ ~ 0 C. asagaiensis l;t'iJi'~-'*111itw.1!!-ff RIFt J: !J ~ L.. ftl!.fl! ,1-~t..:: !J~ll¥1i:ilEr¥t::t~h- -~1(· tc.!: nu:~ ~.n.. ~ oilui•~

lntrodttdion :-The genus Cyclina is
distributed from Cochin China to North
Japan and includes the following recent
species:ment, Shimane University, Matsue, for
his encouragement. Thanks are also due
to Messers. F. UEDA, T. AKuTsu andK. MASUDA for kindly donating their
specimens for this study.

Cyclina sinensis (GMELI:--;) ... Cochin China,
China, Philippine Islands, Ryukyu.
Cyclina j/Jvida DESI!AYES ... China, Korea,
japan.
Cyclitw orienta/is (SoWERBY) .. .Japan.

Systematic Descriptions
Family Veneridae LEACH
Genus Cyclitta DEsHA YES, 18,19

As fossil the genus has been reported
from the Paleogene of northern Kyusyu(Cyclina (?) 1zodai NAGAo and Cyclina
compressa NAGAo), Hokkaido ( Cyclina?
shirokia11a YoKOYAMA) and from thereon
through the Cenowic formations of Japan.
During my study on the specimens of
this genus, stored in the Institute ofGeology and Paleontology, Tohoku University, I have found two undescribed
forms which are dealt with herein.
Before going to the description, I wish
to offer my thanks to Prof. S. HM-'ZA wA
and Dr. K. HATAI of the Institute ofGeology and Paleontology, Tohoku University, Sendai for their supervision andfor reading the manuscript and to Mr.
S. NISIY AMA of the Geological Depart-

1849. Cyclitla DESHA YES, Trait. Element. Conchy., Vol. 1. fasc. 2. pp. 623-626.
1791. Genotype:- Vet1us sinensis GMELIN, Recent, China Sea.
Subgenus Cyclina s. s.

Cyclina ( Cyclitza) orienta/is(SoWERBY)
Plate 15, Figures. 7a-b.
1855. Artemis orietJia/is SoWERRY, Thes. Colicit. Vol. 2, p. 661, pl. 144, fig. 79.
1869. Cyclina orienta/is PFEIFFER in MARTINI
u. CHEMNJTZ, Syst. Conch. Cab., Vol.
11, pt. 1, p. 113, pl. 28, figs. 7-9.;
DUNKER, 1882, Ind. Moll. fap., p. 205.
1920. Cyclitza chi11eusis YoKoYAMA, ]ot4r. Coli.

Sci. Imp. Univ. Tokyo, Vol. 39, art. 6,
pp. 119-120, pl. 11, figs. 7, 8.

1) Read June 30, 1951; received Dec. 3, 1951

167

168
1936.

Yasuhiko KAMADA

Cycli11n sinensis NoMURA and HATAI,
Saito Ho-011 Kai Mus. Res. Bull. No.
10, p. 128. (with full reference prior to1936; specimens not well preserved.)
1950. Cyclinn orienta/is Qy AMA. Miner. &Gcol. Vol. 3, no. 6, p. 2.

SoWERny's original description is asfollows:" Art. testa rotundata, ventricosa, con~entrice et radiatim ad latera vix stri·ata, pallide aurantift. prope marginem
ventralem rubecente; intus margine
crena to; lunul<i nulla. "This species has a shell which is round·er and more solid than Cyclitza sine11sis( GMELI N ). The shell sculpture consistsof both concentric and irregular striations, which are somewhat raised over·the surface and become more elevatedtoward the extremity ; there are crossedwith fine impressed radiating striae.
Surface covered with nidus and fawn-colored epidermis with a beautiful violet
band only near the border. Of the threecardinals in the right valve, the middle
is the stoutest, the anterior one is short-est and thin, ar.d the largt:st or posterior
one is slightly arched and bifid. In theleft valve, the posterior is longest andthin, the middle and anterior ones areslightly bifid.
As Recent this species has been re-ported as Cyclitza sinensis (GMELIS)
from many localities in Japan and Korea
and as fossil it has been recorded fromDi IIICIISiOtiS(in mm.)the Pliocene, Pleistocene and the raisedbeach deposits of Kanagawa and Chiba
Prefectures and the environs of Tokyo.
Fossil specimens in the Pliocene Dainenji
formation from Aoba-yama and Mukaiyama both in the western part of Sendai
City are also identified with C. orienta/is.
Living distribution:From Mutsu Bay, Aomori Prefecture,southwards along the Pacific coast to
Kagoshima and Nagasaki Prefectures inKyusyu along the Tung-hai, and along
the Japan Sea northwards to Mikata,
Fukui Prefecture. Also west coast of
Korea.

Cyclina ( Cyclina) japonica
KAMADA, n. sp.
Plate. 15, Figures. la-b, 2, 4.
1926.

Cycliun chinensis YoKOYAMA. ]our. Fac.
Sci. Imp. Univ. Tokyo, Ser. 2, vol. 1,
p. 222, pl. 28, fig. 7 (not of GMELIN,
1791).

Shell equivalvc, inequilateral, very gibbous, a little higher than long. Anterodorsal margin nearly straight. Posterodorsal border arched, posterior marginsubtruncate, ventral border rounded.
Beaks bluntly pointed. Sculpture withconcentric, fine growth lines, crossedwith numerous impressed, fine radiating
striae. Pallial sinus triangular. Inner
margin of the shell crenulated.

IGPS• coiLcat. no.

LengthHeightWidth

Holotype
Para type 1
2
3

72952
7110

-li.7
48.0

30.9

52776

432
44.8

50.2
!:0.3
45.0

4

72954

36.0

C. (C.) japonica---··------

Both valve

49.0

30.1
28.5
16.0

Left valve

36.9

22.4

lloth valve.

* Abbreviation for Institute of Geology and Paleontology, Sendai.

On Some Species of Cyclina from Japan and Korea

Remarks :-This new species is distinguished from C. orienta/is by the more
ventrally sloping antero-dorsal margin,
by the less incurved beaks, by having
less angular corner between the posterndorsal margin and posterior border, andby the more acute apex of the pallial
sinus.
It is also distinguished from
Cyclina takayamai 0YAJ\1A from the Syobara Miocene (0TUKA, 1938, pp. 32-33)
by the large size, more inequilateral
aspect of the valves and by the more
sloping antero-dor,;al margin of the shell.
Localities and formation :-IGPS loc.
no. Iw-3; 200m west of Tokunari, Machino-machi, Fugeshi-gun, Ishikawa Prefecture. (Horyuzan ). Lat. 37° 21' 36'' N.,
Long. 137° 05' 20" E. Type locality;
Higashi-Innai formation, I\Iiocene. Holotype.
IGPS loc. no. Gi-4; Shobasamahora,
1 km. west of Tsukiyoshi, Aki-mura,Toki-gun, Gifu Prefecture. (Mino-Ota)
Lat. 35° 221 48'' N, Long. 137~ 13' 36'' E.
Tsukiyoshi formation, I\iiocene. IGPS
coli. cat. no. 7110. Paratype.
IGPS loc. no. Sh-1 ; Pass between Ayukawa, Ayukawa-mura and Kurokawa.
Yamauchi-murd both of Koga-gun, Shiga
Prefecture. (Kamiyama) Lat. 34' 56' 0:1''
N., Long. 136° 20' 43'' E. Kaminohira
formation, Miocene. IGPS coil. cat. no.
52776. Paratype.
IGPS loc. no. Fs-13; In the 1st pit of
the Taisho Coal Mining Co., Kamiyamada, Yamada-mura, Iwaki-gun, Fukushima Prefecture. (Onahama). Lat. 36°
57' 07" N., Long. 140° 57' 10'' E. Yamada
formation, Miocene. IGPS coil. cat. no.
72954. Paratype.
Cyclina ( Cyclina) asagaiensis
KAMADA, n. sp.
Plate. 15, Figures. 3a-c.

169

Shell rather thin, suborbicular excepting the umbonal region, gibbous,equivalve, nearly equilateral, a little
higher than long. Antero-dorsally slightly concave, postero-dorsal margin shortand convex, ventral margin rounded.
Umbonal region narrO\\", projecting, beaks.
situated almost nearly central, small,pointed and inclined inwards. Surface
with concentric ribblets crossed by many
radial ribs broader than the interspaces.
There is a lunule-like area bordered by
the narrow groove which extends fromthe beaks to the postero-ventral corneron the inner side of the shell. Pallial
sinus rather small, deep, sinusly ascending, rounded. Inner margin crenulated.
Length of holotype 37.0 mm., height
40.8 mm., width of intact shells 26.6 mm.
Holotype :-IGPS coil. cat. no. 72955.

Remarks :-Cyclina (Cyclina) japonica
n. sp. resembles the present n~w species,
but can be distinguished by the smaller
and thinner 'shell, the more projecting
umbonal region and by the position of
the beaks. The radial ribs of the present
new species are stronger than the impressed radial striae of japonica or ori-

mllalis.
Localities and formation :-IGPS Joe.
no. Fs-11; In the Omachi abandoned
shaft of the Joban Coal Mining Co.,
north of Takasaka, Uchigo-machi, Iwakigun, Fukushima Prefecture. (Taira) Lat.
37° 03' 07 11 N., Long. 140° 51' 04'' E.
Type locality. Asagai formation, Oligocene (Rupelian). Holotype.
IGPS loc. no. Fs-12 ; 500 m. north of
Yumoto railway station on the Jobanline, Mukaida, Yumoto-machi, Iwaki-gun,
Fukushima Prefecture. (Taira) Lat. 37°
00' 24'' N., Long. 140° 51' 55'1 E. Asagai
formation. IGPS coli. cat. no. 72956.
Paratype.

Y asuhiko KAMADA

1-70

Subgenus Cyclinorbis MAKIY AMA,

1926. emend.
Type: Cyclina lunulata MAKIYAMA, 1926.

Cyclinorbis was proposed by J. MAKI·~AMA(1926, p. 158) as a new section of

Cyclina, based on CycUna lrmulata MAKI·
YAMA from the Miocene Heirokudo for·mation of North Korea.
Subsequently in 1936 (p. 212), he statedthat the features upon which he founded·Cyclinorbis are also found on some ex:·amples of C. sinensis, and therefore
Cyclinorbis may be a synonym of Cycli·na s.s.
My study of numerous Cyclina bothfossil and recent leads to that Cyclinorbis
may be retained as of subgeneric value
and can be defined as follows :Shell suborbicular, a little longer thanhigh, anterior end produced, not so swollen as in Cyclina s. s., the umbones are
small, the beaks are posterior. Sculpture
with fine concentric lines and no radialstriae. Crenation of inner margin only
<m antero- and dorsal borders. Of threerather thin cardinals, only the posterior{)ne in the right valve is bifid.

Cyclina ( Cyclinorbis) lmmlata
MAKIYAMA
Plate. 15, Figures. 5-6.
Cyclina (Cyclinorbis) lmwlafa MAKI·
YAl\1A, Mem. Coli. Sci. Kyoto Imp. Univ.,
Ser. B. vol. 2, 110. 8, art. 8. p. 158, pl.
13, fig. 1.
1936. Cycli11a (s. s.) lunu/ata 1\IAKIYA:\tA. ibid.,
vol. 11, no. 4, art. 8. p. ·212.
1938. Cycli11a lunulata NoMURA and HATAI,Japan. ]our. Geol. Geogr .. Vol. 16, nos.

"Shell suborbicular, a little longer thanhigh, inequilateral, moderately convex, thickc:oncentrically finely ribbed. Beaks not muchraised, a little convex, acute, approximate,inclined forward, located at about the posterior
third. Anterior end produced, broadly rounded,
the dorsal margin long, arcuate, slowly declivate ; posterior end broader. more elevated,
regularly round ; the dorsal margin high, shortarcuate: ventral margin semicircular. Lunulevery small, concealed below the beaks, super·
ficial ; there is defined a large oval lunule-likearea bordered by a distinct line. Sculpture
consisting of fine dense, regular concentric
riblets, which are sharpened and elevated at theextremities. Hinge-plate moderately broad,arcuate, with 3 divergent, subvertical cardinals,
the right po3terior bifid. Posterior dorsal
margin crenate within, the other part of theinterior margin S!1100th. Pallial sinus long,subvertical. Height, 47 mm.; length, 49 mm.;diameter, 27 mm. "Remarks :-In this description,]. MAKI·
YAMA described that the posterior dorsal
margin is crenate within but on examination of topotype specimens of thisspecies, it is found that crenations on
the margin are restricted to the antero·dorsal border while the other parts aresmooth.
C. lrmulata is characteristic in its
moderate thickness of the shell, which
is a little longer than high, produced
anterior end, rather small convexity andconcentric riblets. By the said features,
this species can be distinguished from

1926.

1-2, p. 5.

]. MAKIYAMA's original description is
as follows:I

C. ( Cycli1zorbis) luzmlata

I
I
I

Length ~ Height WidthI

Dimensions(in mm.)

48.0

49.5

30.2

46.2

45.5

41.3
40.2

39.2
39.5

263
21.1
21.5

I 33.2I

307

160

Topot}'pe
I(IGPS coli. cat. I
1

No. 64665)

I

On Some Species of Cyclina from Japan and Korea

the other species of Cyclina s. s. This
~pecies resembles the Recent C. jlavida
DESHAYES (Pl. 15, fig. 8), but can distinguished from it by having a smooth
inner ventral and posterior margin andby the subvertical sinus.
Associated fauna and the ecologicalsignificance of Cyclina :Generally, the Recent Cyclhza is foundliving in bays rather than in regions
directly in the influence of open seas.
In bays, the genus is found flourishingin regions of either brackish-water orpure marine conditions, thus it may besaid that the animal is capable of enduring such diverse conditions as inwaters-of inlets or lagoons or in shallow
waters of estuaries where the salinity isrelatively dilute.
For example, Matsukawa-ura (Matsukawa inlet) in Fukushima Prefecture, isa typical brackish-water inlet isolated
from the open sea (Pacific Ocean) by awell developed sand-bar. Here Cyclitza
orienta/is (SoWERBY) is found from the
muddy bottom of this inlet in association
with the following molluscs;
Ostrca (Crassostrea) gigas THUMBERG
Meretrix meretrix (LINNE)
VeneruPis (Amygdala) philippinarum (ADAMSet REEVE)
Gastrmw ym1taiensis CROSSE et DEBAUXSolen gouldi CoNRAD
Mactra t•eneriformis RF.EVE
M_va (Are11omya) japonica ]AY
Batilloria mztltiformis (LISCHKE)
Cerithidea rhizoporarum (A. ADAMS)

It is remarkable that the paleo-ecological conditions of Cyclitta lzmulata
and C. japonica have close relationshipwith the genus Vicm·ya as already described by H. YABE and K. HATAI (1938,
pp. 161-164).

In the Meisen district of North Korea,

C. lzmulata was found from the Heiro-

171

kudo formation in association with Batil-

laria yammzarii MAKIYAMA, B. tateiwai
MAKIYAMA, Cerithidea kanpokttensis MA·
KIYAMA, Anadara daitokudoemis MAKI·
1 AMA and Vicarya japmzica YABE andHATAI.
From this assemblage it isreadily noticed that the Heirokudo formation was deposited in a bay.
In Tsukiyoshi, Akiyo-mura, Toki-gun,
Gifu Prefecture, Cyclina japonica occurs
abundantly from the lower part of Tsukiyoshi formation in association withVicarya yokoyamai TAKEYAMA and V.
martini YABE and HATAI. The othermolluscs occurring in association are,according to the recent studies of K.
HuziTA and S. OGoSE (1950. p. 488) as
follows:Vicaryclla ishiiana (YoKOYA~IA)
Cerithium kaneltarai HUZITA and OGOSE" Pseudomurex" minoensis (OYAMA and
SAKA)

Trapezium modiolaeforme OYA:'.IA and SAKA
Cleme1ztia "papyracea (GRAY)"
Dosinia japonica? REEVE
Dosinia anguloides NOMURA
Sanguinolaria minocnsis (YOKOYAMA)
Katelysia nakamurai !KEBE
Lucina yokoyamai OTUKA

The fossils above enumerated do notsuggest a typical brackish water fauna,
although there seems to be little doubt
as to the embayment nature of thesea. The lower part of the Tsukiyoshi
formation consists chiefly of tuffaceous
fine-grained or silty sandstones. Conesof the pinetree or leaves and carbonaceous materials are frequently foundin the formation in association with themolluscs. Judging from the paleontological evidences just mentioned, the paleoecological nature of the Tsukiyoshi formation is "one of a sandy or muddy
enviroment, probably in the reach ofbrackish-waters" as already mentioned

172

Yasuhiko KAMADA

by H. YA13F. and K. HATAI (1938, p. 162).
In the eastern part of Koga-gun, Shiga
Prefecture, Cyclina japonica occurs fromthe Kaminohira formation. From thelocality where Cyclina is found, N. IKF.RE(1934, pp. 114-115) listed the following
molluscs, namely:Anadara cf. valentula YoKOYAl\1A
Trapezium kurodai lKE13E
Dosinia (Phacosoma) japonica troscheli LISCIIKE

Solctellina miuoensis YoKoYAMA
Aloides ta11abensis? (YoKOYAMA)
Batillaria sp.
·vicarya yokoyamai TAKEYAMA
Vicaryel/a bacula (YoKOYAMA)The Kaminohira fauna contains somebrackish-water molluscs such as the genera Trapezium and Batillaria.
In Tokunari, Machino-machi, Fugeshi·gun, Ishikawa Prefecture, Mr. K. MAsuDA collected Cycli11a japonica in association with the following molluscs fromhis Higashi-Innai formation:Anadara daitokudocnsis 1\IAKIYA:\tA
Anadara kakebatensis HATAI and NISIYAMA
Lucinicus yokoyamai OTUKA
Phaxus izumoensis (YoKOYAMA)
Taras ferruginafa r..I.-'t.KIYAMA
Vicarya callosa ]E:-;KI:\S
Vicaryclla becula (YoKOYAMA)The fossils from the Higashi·Innai formation suggest that the formation was
deposited in shallow water under warm
temperate thermal conditions. The occurrence of brackish water genera as
Vicarya and Vicaryella is of interest
because it suggests that the conditions
then prevailing may not have been puremarine.
From the features described above, it
is noticed that both Gyclt"na ltemtlata
and japonica flourished in an embayment
analogous to the Recent Cyclitza. No
boreal or cold water species are foundin association with either of the twomentioned cylinids although warm temperate to subtropical genera as Vicarya
and Phaxus are common. This fact
leads to the conclusion that these fossil
cyclinids flourished under conditions similar to the two genera mentioned.
CycHna asagaiensis is one element of
the Asagai fauna in the Joban coal-field.
Its occurrence is very rare and is foundonly from the Uchigo and Yumoto areas.
The Asagai formation contains many
molluscs and foraminifers which arecharacteristic of boreal or cold shallow
water as already described by K. HATAI
and myself (1950, p. 67) and K AsANo(1949, p. 474). Accordingly C. asagaiensis
may be a cold water species of theneritic zone.
References
ASA:-iO, K. (1949). Foraminifera from the
Asagai Formation (Tertiary) of Fuku·
shima Prefecture. Japan. ]our. Paleont.,
vol. 23. tzo. 5. pp. 473-478.
DALL, W. H. (1902), Synop;is of the Family
Vemridae and of the North American
Sp..:cies. U. S. Nat. Mtts., vol. 26, p. 348.
HATf'd, K. and KAMADA. Y. (1950), Fossil
Evidence for the Geological Age of the
Uchigo Group. Joban Coal-field. Shortpapers IGPS. 110. 2, pp. 58-74, 2 text-figs.
HUJI"rA, K. and OGoSE, S. (1950), Lithologic
Classification of the Cenozlic Strata inthe :'\orthern Area of Mizunami-machi,
Toki-gun. Gifu Prefecture. jour. Geol.
Soc. japan .. Vol. 56, no. 662, pp. 481-192.
1!\EBE. N. (193<1), Miocene Series of EasternPart of Koga.gun, Shiga Prefecture. Chi·kyu, vol. 22, 110. 2, pp. Jl0-123. 3· text-figs.
MAKIYAMA, J. (1926). Tertiary Fossils from
North Kankyo-do, Korea. frfem. Co!l. Sci.
Kyoto Imp. Univ., Ser. B Vol. 1 I, no. 3,
art. B. pp. 143-160. 2 pis.
- - (1936), The Meisen 1\liocene of North
Korea. Mem. Col!. Sci, Kyoto Imp. Utliv.,
Ser. B, vol. 11, 110. 4, art. 8, pp. 193-228,

KAMAD1\ : Cyclina from Japan and Korea

Photo. by KUMAGAI

Pr.Nm 15

On Some Species of Cyclina from Japan and Korea
2 pis .• 2 maps.
NAGAo, T. (1928). Paleogene Fossils of theIsland of }(yfishfi, Japan. Part II. Sci.
ReP. Tohoku Imp. Ur.iv. 2nd Ser. (Geology), val. 12. 110. 1, pp. 11-140, 17 pis.
NoMURA, S. and HATAT, K. (1936), Fossils
from the Tanagura Beds in the Vicinityof the Town Tanagura, Hukushima-ken,
Northeast Honsyu, Japan. Saito Ho-on Kai
Mus. Res. Bull., tzo. 10. pp. 109-155, 5
pis. (with bibliography).
·OTUKA, Y. (1938), Mollusca from the Mioceneof Tyugoku, Japan. jour. Fac. Sci. Imp.
U1ziv. Tokyo. Vol. 5, pt. 2, pp. 21-45, 2
text-figs, 4 pis.
-DYAMA, K. (1950), Remarks on Japanese Fossil l\Iolluscan name. fl.fi1zer. & Ceo/. Vol.
3. 110. 6, pp. 1-4.
PFEIFFER, L. in MARTINI und CHEM~ITZ

173(18~.9), Systematiches Chonchylien-Cabinet. ~·of. 11, pt. I, Die Families der Ven ·usmuscheln, Veneracea. V. Cyclina pp.
110-117.
YABE, H. and HATAI. (1938), On the JapaneseSpecies of Vicarya. Sci. Rep. Tohoku Imp.
Ut:iv., Sendai. 2nd Scr. (Geology), val. 1.9,
110. 2, pp. 149-172, 1 pl.
YoKoYAMA, 1\1. (1920), Fossils from the Miura
Peninsula and its Immediate f\orth. ]our.
Col/. Sci. Tokyo Imp. Univ., val. 39, art.
6. pp. 1-193, 19 pis., 1 map.
- - ( 1926), Molluscan Fossils from the Tertiary of Mino. ]our. Foe Sci. Univ. Toky•o,Ser. 2, val. 1. pt. 7, pp. 213-227, 1 p!.
- - (1932) Tertiary Mollusca from the Coal·
field of Uryu, Ishikari. jour. Foe. Sci.
Imp. Univ. Tokyo, Ser. 2, val. 3. pt. 6,
pp. 221-247, 4 pis.

Explanation of Plate 15
Figs. la-b, Cyclhza ( Cyclina) japonica KAMADA, n. sp. Holotype. IGPS coil. cat. no. 72952. a. Side
view of right valve, b. Anterior view of the same. Loc. IGPS Icc. no. lw-3, 200m. west of
Tokunari, Machino-machi, Fugeshi-gun, Ishikawa Prefecture. Higashi-Innai formation, Miocene.
Fig. 2, Cyclina (Cyclina) japonica KAMADA, n. sp. Paratype. IGPS coil. cat. no. 7110. Internalview showing the pointed apex of the pallial sinus. Loc. IGPS Joe. no. Gi-4. Shobasamahora,
1 km. west of Tsukiyoshi, Akiye·mura, Toki-gun, Gifu Prefecture. Tsukiyoshi formation,
1\·iiocene.
Figs. 3a-c. Cyclina (Cyclitw) asagaiensis KA'YIADA, n. sp. Holotype. IGPS coli. cat. no. 72955. a.
Side view of left valve, b. side view of right valve, c. anterior view of the s:~me: Lee. IGPS
Icc. no. Fs-11. In the Omachi abandoned shaft of the ]oban Coal Mining Company, north ofof Takasaka. Uchigo-machi, Iwaki-gun, Fukushima Prefecture. Asagai formation, Oligocene.
Fig. 4. Cyclina (Cyclina) japonica KAMADA, n. sp. Paratype. IGPS coli. cat. no. 72954. Loc. IGPS
Joe. no. Fs-13. From the 1st pit of the Taisho Coal Mining Company, Kamiyamada, Yamada·mura, Iwaki-gun, Fukushima Prefecture. Yamada funnation Miocene.
Figs. 5-6. Cyclir.a (CJclinorbis) ltmulata ].\{AKIYAMA. Topotype. IGPS coil. c.• t. no. 64665. 5. Side
view of left valve, 6. internal view showing vertically pallial sinus and anterior crenulation.
Loc. Nanseki, l'vJeisen-gun, Kanky6-d6, North Korea. Heirokudo formation, Miocene.
Figs 7a-b, Cyclina (Cyclina) orienta/is (SoWERBY). IGPS coli. cat. no. 72685. Loc. Matsuka.wa-ura,Nakamura-machi. Soma-gun, Fukushima Prefecture. Recent.
Fig. 8. Cyclina jlavida DESHAYES. IGPS coil. cat. no. 8810. Loc. Mokpo, Zenra-nando, South Korea.
Recent.

174

Proceedings of the Palaeontological Society of Japan

PUBLICATIONS RECEIVED

No. 878. Er.usoN. S. P.: Subsurface Woodford
Black Shale. West Texas and Southeast New Mexico. Bureaz4 Econ. Geol.,
Univ. Texas, Rep. Invest., No. 7, 1950.
No. 879. BARNES, V. E.: Utilintion of Texas
Serpentine. Utziv. Texas Pub/., No.
5020, 1950.
No. 880. VAS HoEPEN, E. C. N.: The So-called
Old Mouth-edges of the Ammonite
Shell. Paleontologiese Navorsing, Na.
sionale Museum, Bloemfontein, Dl. 1.
St. 8, 1951.
No. 881. VAN HoEPE:o-;, E. C. N.: Pervinquier·idae, Arestoceratidae, Cainoceratidae.
!hid., Dl. 1, St. 9, 1951.
No. 882. VAN HOEPEN, E. C. N.: A Remarkable Dcsmoceratid from the South
African Albian. Ibid., D!. 1. St. 10,
1951.
No. 883. Rocznik Polskiego Towarzystwa Geo·
logicznego, Vol. 20, Nos. 1-2, 1951.
No. 8&t. Ministero Agricultura, Notas Prelim·
inares estudos. No. 50, 1949.
No. 885. Ibid., No. 51, 1950.
No. 886. Minisfero Agricultura, Boletim. No.
132, 1949.
No. 887. Ibid., No. 133. 1949.
No. 888. Ibid., No. 84, 1949.
No. 889. Ibid., No. 86, 1950.
No. 890. Mario da Silva Pinto, Atividades do

Departmento Nacional da Producao
Mineral, em 1g.l8, 1950.
No. 891. JI!~1&'1Ut~lti (1fi 1 f"lEil.W3r).
No. 892. )A.~*.~ . .lll1.~lUF9l~-'5-. No. 1, 1951.
No. 893. :Ri!!iiif>tfil, No. 22, 1951.
No. 894. i't~lilf1tfll. No. 23, 1951.
No. 895. Proc. Liverpool Geol. Soc. Vo•. 20.
Pt. 3, 1950.
No. 896. EJFI'ER, G. K. Jr.: Geology of the
B:1rrilla Mountains, Texas. Bureaz4

Econ. Geo/., Ut~iv. Texas, Rep. Invest.,
No. 8, 1950.
No. 897. FI.AWN, P. T.: Pegmatites of the
Van Horn Mountains, Texas. Ibid.,

No. 9, 1951.
No. 898. HERAI.D, F. A. : Occurrence of Oiland Gas in Northeast Texas. Univ.
Texas Pub/., No. 5116, 1951.
No. 899. REINHART, R. H.: A New Shark of
the Family Ptychodontidae fromSouth America. Univ. Calif. Pub/.,
Bull. Ceo/. Sci., Vol. 28, No. 8, 1951.
No. 900. REINHART, R. H. : A New Genus of
Sea Cow from the 1\ficcene of Colombia. Ibid., Vol. 28, No. 9, 1951.
No. 901. SAVAGE, D. E.: Late Cenozoic Vertebrates of the San Francisco Bay
Region. Ibid., Vol. 28, No. 10, 1951.
No. 902. STIRTo:--<, R. A. : Ceboid Monkeys
from the :1\Iiocene of California. Ibid.,
Vol. 28, No. 11, 1951.
No. 903. Rocznik Polskiego Towarzystwa Geo.
logicznego, Vol. 20, No. 3, 1951.
No. 904. Ibid., Vol. 20, No. 4, 1951.
No. 905. HoFMANN. F.:
Zur Stratigraphie
und Tektonik des st. gallischthurgaui·
schen Miozans. ]afzrb. St. Ga/lischen
Naturw. Gese/ls. Bd. 74, 1951.
No. 906. BiiCHI, U. P. und WEI.TI, G.: Zur
Geologie der siicllichen mittellanclischen Molasse der Ostschweiz zwischen Goldingertobel und Toggenburg.
Eclog. Geol. Helv., Vol. 44, No. 1,
1951.
No. 907. STAUB, R.: Uber die Beziehungenzwischen Alpen und Apennin und die
Gestaltung der alpinen Leitlinien
Europas. Ibid., Vol. 44, No. 1, 1951.
No. 908. HIBBARD, C. W.: A New Mastodon,
Serridentim4s meadensis, from theMiddle Pliocene of Kansas. Conlr.
Mus. Paleont., Ut;iv. Michigan, Vol. 9,
No. 6, 1951.
No. 909. HIBBARD, C. W.: Vertebrate Fossils
from the Pleistocene. Stump Arroyo
Member, Meade County, Kansas. Ibid.,
Vol. 9, No. '(. 1951.

Trans. Proc. Palaeont. Soc. Japan N.S., No. 6, pp. 175-178, May 30, 1952.

214. TWO NEW TRIASSIC ESTHERIANS FROM PROVINCE
OF NAGATO IN WEST JAPANTEIICHI KOBAYASHI
Geological Institute, University of Tokyo:&r'JJt. <0-"'4Hc !'l~:U 2 *ill!:- J!/.i\l;tif...t.!ili§TJi<O Estherites atsue11sis, Estherites 11akazawai
t:ft;ltl..., k<O EstheriinatiOOOf;f.t<:-:::>~t•"Cp~t~-J-.,

The occurrence of Estherites coreanica(OzAWA and WATANABE) and Estherites
kawasakii (OZAWA and WATANABE) at
Hikiji in the Asa area in Prov. Nagato(Yamaguchi Prefecture) in the lower
Noric Kamosho stage of the Mine series
has recently been reported (KnRAYASHI,
1951). Two new sp~cies, Estlzerites a·
tsttensis and l!..stherites nakazawai, described here, were subsequently discovered by N,u:AzAWA in a shale bed of.
the Atsu series at Hamada of Tsubuta
in the same district. This cstherianhorizon is located between the coal measures of Tsubuta and Akaiwa and lies
below the Ha/obia shale containing Ha/obia aotii KoBAYASIII and IcHIKAWA (1950),(i.e. H alobia multistriata KoaA YASH! andAoTI, 1943). Therefore the age of theestherians must be either Ladino-Carnicor early Carnic In other words they
are the oldest estherians in Japan andprobably the oldest among the Mesozoic
ones so far known in Eastern Asia.
My thanks are due to Mr. Keiji
NAKAZAWA for the privilage of describing
them and to Prof. S. MATSUSHITA and
Mr. F. KATo for the facility and asRead Feb. 10. 1952; received Jan. 19, 1952.
175,j,~jt-sistance of this study in my visit to the
Kyoto University. The type spc!cimens
are kept in the Geological Institute of
Kyoto University.

Estherites atsuensis KonA YAS III,
new species
Plate 16, figures 1-7

Description :-Carapace valve subelliptical but truncated at about threequarters of the short diameter of theellipse by the straight dorsal margin andnearly equilateral, but generally theanterior margin is somewhat more broadly rounded than the posterior marginwhich is in turn more gently curving on
the ventral than on the dorsal side;height corresponds about three-quarters
the length; dorsal margin occupies aboutfour-fifths the length of the carapace;umbo terminal ; umbonal half or so of
the valve more convex than the peripheral zone; growth lines very weak or
absent in the umbonal quarter or onethird; in the rest they are distinct, butvery fine, widely apart from one anotherand countable about 12 in the full grownstage; their intervals ornamented byradial lirae.

Teiichi KoBAYASHI

176
Specimen

I

No. 1
-

No. 2
~-

~~-

I
I~H_I_ ~M
3.75
2.80
2.20
L

I

-

3.65

2.70

2.20

3.10

2.50

~~-

No. 4

3.90

No. 12I 2.60 I 2.50
s.. to I (3.30) us())

No. 14----3.90(in em.)L:

lengthII:heightDM : length of dorsal margin

Observation :-Specimens Nos. 1 and 2(figs. 1 and 2), both right valves, areselected respectively for the holo- andpara-type of this species. Their dorsal
margin is straight and as long as fourfifths of the valve; the maximum height-of the valve runs through the middle ofthe margin and a little anterior to the
middle of the carapace-length.
Theumbonal half of the valve is slightly
depressed in the holotype and somewhat~onvcx in the paratype.
Six or sevenlines of growth are distributed equidistantly in the more distal part.
In a slab (fig. 9) three valves in thesan1.e orientation are all equally depressed dorso-ventrally, while one disposed
diagonal to them is compressed obliquely.
If the secondary deformation of theoutline to this extent is brought into
consideration, there is no doubt that ataller right valve (specimen No. 4, fig.
.3) and a broader left valve (specimen
No. 12, fig. 4) belong to this species,
because they agree with the types inmost other aspects. They are not muchdifferent from the types in height-length
proportion, nor the proportional length~f the dorsal margin to the valve.
Growth lines are countable six in the

peripheral zone in the specimen No. 4.
These lines are usually very narrow, if
compared to their intervals which arenearly equal in breadth.
It is certainly a remarkable fact thatthe umbonal part is strongly depressedand deformed secondarily in most specimens at hand. This is probably because
the part has been more convex thanthe peripheral zone. Specimen No. 13(fig. 5) is an external mould of a rightvalve on which a part of the left valve
lies near the umbo. It has seven linesof growth. The umbonal part less than
one-third the height is practically smooth. Furthermore it is noted that thevalve becomes somewhat more convex
in the umbonal half and especially so in
the umbonal quarter. Such a swelling
tendency can be seen also in the umbonat one-third of the specimen No. 14(fig. 6) which is the largest. There no
growth line is discernible. As the secondary reduction of the convexity is
generally stronger in shales, the swellmust have been much more prominent.
Because the strongly deformed part ofthe preceding specimens corresponds to
the S\vell in the last two specimens, thepart of the former may be interpreted
to have been depressed, causing a cracknear the margin of the swell in many
specimens, i.'-lo. 12 for example. If so
deformed, the position of the umbo isindeterminable, but it can be ascertained
to be terminal in the latter specimens.
In the specimen No. 5 (fig. 7) lirae areseen to radiate, occasionally joining or
branching, in the intervals near the
periphery of the valve.
In summarizing these observations, thedistinguishing characteristics of thespecies are the smooth umbonal swell,paucity and fineness of growth lines, theirintervals of uniform breadth, the radiallirae on them and the subelliptical out-

Two New Triassic Estherians from Province of Nagato in West japan

line truncated by the dorsal margin.
ComParison :-It is certainly interestIng to see such an umbonal outgrowthof the carapace as seen in Estheriitza
JoNES (1897), because, as suggested by
Jor-:Es for his genus, it shows probably
the lateral outgrowth of the soft part in
the immature stage. In the Bahian estherians from Brazil, however, thegrowth lines are usually distinct on theumbonal swell but obsolete or muchfiner and denser in the peripheral zone.
The change between the two portions ismore abrupt and distinct in them thanin this species.
Jmms referred Estlteria limbata GoLD·EKBERG and Cardinia freysteitzi GEIKITZ
from the Carboniferous of Germany to
Estheriina, although RAYMoND (1946)
considers the latter to be a member of
his PseutkJestlzeria. In the case of limnic-creatures as estherians it is improbable,
I think, that the Carboniferous species
in Germany are linked with the Cretaceous ones in Brazil through the Triassic
ones in Japan. These three groups ofestherians can be distinguished from one
another in their carapaces. More precisely, the flat peripheral zone is muchnarrower in E. limbata and E. freysteitzi
than in E. bresiliensis, E. expansa and E.
astarloides. While the growth lines are
more developed on the swell in all ofthem than in the peripheral zone, thereverse is the case of Estheriles atsuensis
as well as Estlzerites nakazawai. Nevertheless, it is not improbable that theumbonal swelling shows similar development in the immature stage. Therefore
it is my opinion that the three groups
are off-shoots in the similar trend of
evolution, but derived intermittently
from the main stock of Estlzerites atdifferent times and places. If the subgeneric distinction from Estherites sen.
str. bears propriety, each of the three177

groups must be distinguished on the basisof the size of the swelling and the surface ornaments. A new subgenus is,however, deferred to establish for theJapanese species until more is known of
the umbonal swell.
Finally the types of this species coincides with Estheria ovata LEA in JoNEs,
1862, in fig. 28, pl. 2, in outline almost
perfectly, but the surface aspect is totally different.
Occurrence :-Hamada of Tsubuta in
Habu-town, Asa-gun, Yamaguchi Prefecture ; in a shale in the upper part of theAtsu series, which is black coloured, but
becomes white, upon weathering.

Estherites 11akazawai KoBAYASHI,
new species
Plate 16, figure 8

The specimen No. 7 (fig. 8) on which
this species is founded has an ovate outline, expanding posteally and attaining
the maximum height at about one-thirdfrom the posterior end. But otherwise
it agrees well with the preceding species.
The type left valve is 4.45 mm. long and3.3 mm. high. Although the posterodorsal part is brocken off, its dorsal
margin is estimated to be about 3.2 rom.
Namely it is a little longer than in theproceding species. Because the heightlength proportion is about the same asthat of the specimen No. 12, the differencein outline cannot be attributed to secon·
dary deformation.
Among the Asiatic species it is notedin Estlzerites sinkiangettsis (CHI), (1931)
that the umbonal one-sixth is smooth
and about 10 concentric lines outside of
it are, like in this species, separated bywide intervals. Its outline is obliquelyovate, but if compared with the preceding, more slender, less expanding antero-

Teiichi KoBAYASIII

ventrally as well as postero-dorsally.
Furthermore no mention is given by
C!u of the swelling of the umbonal
part.
Occurrence :-Same place as the preceding, but very rare.
·---------

Reference
CHI. Y. S. (1931). On the Occurrence of Fossil
Estheria in China and its geological Significance. Bull: Ceo/. Soc. China, t•ol. 10.
JoNES, T. R. (1968), A Monograph of theFossil Estheriae. Palaeontol. Soc.
- - (1897), On fossil Entemostraca from
Brazil. Ceo/. Mag. N. S. Dec. 4, val. 4.
KoBAYASHI, T. (1951), Older ;\lesozoic Estherites from Eastern Asia. ]our. Fac. Sci.
Unit•. Tokyo, sect. 2. vo/. 7, pt. 10.
- - and K. AoTI (19-l3), Haboliae in Jap:m.
]onr. Shigenkagaku Kenkyusho, vol. 1, no.
2.
--and K. ICIIIKAWA (19-1,9), Myophor_ia andother Upper Triassic Pelecypods from the
Sakawa Basin in Shikoku. fa pan. f OM.
Ceo/. Ceogr. rol. 21.
RAYMOND, P. E. (1946), The Genera of Fossil Conchostraca. Bull. Mus. ComP. Zoo/.
vol. 96, 110. 2.

Explanation of Plate 16

Estlzerites atsuensis KoBAYASHI, new species
FigureFigureFigureFigureFigure1. Holotype right valve; specimen No. 1.
2. Paratype right valve; specimen No. 2.
3. Specimen No. ·1·: a right valve.
4. Specimen No. 12; a left valve.
5. Specimen No. 13; an external mould of a right valve with a p'lrt of a left valve near
the umbo.
Figure 6. Sp!"cimen No. 14 ; a left valve and a part of a right valve.
Figure 7. Radial lirae in the inten·ais among lines of growth near the periphery: specimen No.
5. x40.7

Estlterites nakazawai KosAYASIII, new species
Figure 8.

Holotype left valve, specimen No. 7.

Esthcrian shale of Hamada near Tsubuta
Figure 9.

A sla!J showing the mode of deformation of four v:tlves aligned in two different
directions; specimen No. 10. x 4.5
All figures ten times magnified unless noted.
All specimens are kept in the Geological Institute, Kyoto University.

PLAT!!. 16

KosA Y ASH! : Triassic Estherians from Nagato

3
1

5

7

6

Trans. Proc. Palaeont. Soc. Jap:m. N.S., No. 5, pp. 179-183, 1 text-fig. May 30, 1952.

215.

A MIOCENE CRAB, HYAS TSUCHIDAI N. SP. FROM
THE W AKKAN AI FORMATION OF TESHIOPROVINCE, HOKKAIDORIKIZO IMAIZUMI
lst college of Arts and Sciences. Tohoku University, SendailiJi-IIH~t"-1-~"' t "fjf.::. =::Jti)iJ!Xlji~~itffi1:~ai'ii~;1J,~=-' 7=--:" D --:·/ J111lutmJ¥.

§Zu-99- 2~~~m~v-~w~v±~2*~K~D·~~~~'l1-~/t"f'ff.::.tt$~­
vm'l:.fi!i Jlyas coarctatus LEACH ~~::~~U:fjTQ 0 ~*Austria, Preding V Helvetian ~ !JHyas meridio11alis GLAESS:-.lER, 1928 :D: Algeria. Oran v Sahelian ~ 9 Hyas orane11sis VAN
STRAELEN. 1936 ;6:ft{fr;Ht."C~·~o A,&.1J~
The fossil crab described herein wascollected by Mr. T. TsucHIDA of the
Teikoku Oil Company from the upper
Miocene Wakkanai formation at Saro,
Toyotomi-mura, Teshio Province, Hokkaido, along the south bank of the
Sarubetsu, a tributary of the Pankeepekorobetsu and kindly submitted byhim to the writer for study.
The writer proposed a new specificname Hyas tsuchidai for this fossil form,
the specific name was dedicated to thecollector of the specimens.
All the recent species of the genus
Hyas are the inhabitants of the Arctic
circle, except for only one subspecies,
Hyas coarcta!tts ursitms RATHB:JS, which
is now known to be distrib:J.ted fromthe Okhotsuk Sea to Amoy along theSouth China Sea via the Japan Sea, as
stated below again.
The exceptional
distribution just mentioned is evidently
controlled by the cold Liman stream.
Therefore, the occurrence of the fossil
crab suggests that the W akkanai formation which has yield this species was

deposited in the same ecological condi~tion as the Recent species of the genus
are governed. The first described fossil
species Hyas meridionalis G1.AESSNER
was found in the Helvetian stage ofW enzeldorf, Preding, Austria. This fossil record from Austria seems to me toindicate the influence of a cold current
in that region during that period.
The writer wishes to express his sincer
thanks to Mr. T. TsucmoA for his kindoffer of the fossil crab, and also to Dr.
H. Y ABE, Prefessors S. I-lANZA wA and K.
AsA::-<o and Dr. K. HAT AI of the Instituteof Geology and Paleontology, TohoktL
University, for their encouragement.

1) Read June 30. 1951;
1952.

Preservation :-one carapace and its
impression preserved in a gray tufface-

Family Majidae Ar.cocK
Subfamily Hyasteniitzae BAI.ss
Genus Hyas LEACH 1814
Genotype, Hyas araneus (LINNAEUs).
RATHBUN, M. ]., 1925, U. S. Nat. l\1us.
Bul. 129, Smith. lnst., p. 252.

Hyas tsuchidai IMAIZUl\11, n. sp.

received Feb. 5,

179

180

Rikizo IMAIZUMI-ous fine sandstone. It is compressed
towards the right frontal direction but
exposes the left lateral side and the-orbital cavity as a result of the left
·supra-orbital eave and the left postocular cup being shoved towards theirontal region. Right branchial regionswith numerous wrinkles extending fromthe posterior margin to the anterolateral,-due to subsequent pressure.
The color of the fossil is brownish
sepia. Convex parts of the carapace and·the tops of the tubercles are worn, but
the concave surface of the impression iswell preserved. Appendages lo~t.
Descriptiotz :-carapace broad, subtriangular of lyrate in shape, but narrower than the ·post frontal length.
Rostrum triangular, flat, bifid, a narrow
cleft between rostral horns, the tips
more or less separated ; tips of horns·subacute. Cavity of orbit oblong-spheroidaL Post·orbital and hepatic region
not dilated laterally, antero-lateral margin of hepatic region sharp with arounded postero-lateral angle. Posterolateral margin of carapace rounded.
Cervical groove distinct, marginal line
interrupted by a small and shallow
sinus between hepatic and branchialregions. Surface uneven, ornamented
with granules and granular tubercles,
especially on median gastric area orproto- and meso-gastric regions and in
an oblique row on branchial region.
Meso-gastric region with two large median tubercles. Proto-gastric region with
two inner and three outer tubercles arranged side by side: the inner two tubercles in proto-gastric region continue
with smaller ones to frontal region.
Pustules on inner side of outer surfaceof horns of rostrum decrease in size
anteriorly.
Row of tubercles along'boundary between meso- and metabranchial regions distinct. Lateral mar-gin of carapace tuberculate behind
hepatic rigion. Outer surface of rostrum
densely granulate. Granules on gastric,
cardiac and intestinal region also densely arranged. Epi- and meta-branchialregions distinct.
Dimensions :-carapace, length 16mm. width
13mm.
rostrum, width 2.5mm.
proto- and meso-gastric region,
width 5mm.

lfolotype :-IGPS con. cat. no. 74001.
Locality and geological horizon :-IGPS
loc. no. Te-10. Under the derick of theboring "Toyotomi" Well no. 2 of the
Teikoku Oil Co. at Saro on the bank of
the Sarubetsu, a tributary of the Pankeepekorobetsu-gawa, Sarobetsu, Saro,
Toyotomi-mura, Teshio Province, Hokkaido, (Toyotomi sheet), Lat. 45°41 43"
N., Long. 141°50' 10."4 E., Wakkanai
formation, upper Miocene. T. TsucmoA
wlL 1950.
Geographical distribution of the Recent
species of the genus Hyas :-According
to RATHB~N (1925, pp. 252-281) and
SAKAI <1934, p. 295, 1935, p. 94), thisgenus is now living in Arctic Ocean
with the exception of the Siberian coast
between Long. 147°E. and the Kara Sea,
south of Novaya Zemlya and of theAmerican coast between Baffin Ray andLangton Bay, North West Territories;
Bearing Seas in the Arctic circle, besides, North Pacific Ocean southward
along the Asiatic side to Korea andperhaps Amoy, and on the Americanside to the state of Washington ; North
Atlantic, Cape Hatteras, North Carolina,
and on the European side to the English
channeL
The geographical distribution of thespecies.
1. Hyas araneus (LINNAEUS), (RATH-A Miocene Crab. Hyas tsr1chidai n. sp. from the Wakkanai

181

Text-fig. 1. Hyas tsuchidai1MAIZUMl n. sp.
Holotype, x 1.
Text-fig. 2.
Impression ofholotype, x 1.
Text-fig. 3. The same specimen as text-fig. 2,
enlarged to show
details, x 5.
Text·fig. 4. Holotype restored,
x2.
Text-fig. 5. A copy of Hays
meridio11alis
from
GLAESSSER's fig.l5,
Taf. III.

A Miocene Crab, H.vas Tsuclridai n. sp. from the
Wakkanai formation of Teshio Province. Hokkaido.
BUN, 1925, p. 258), type-locality, North

Sea.
West coast of Greenland from Lat. 69°151 to&!0 111 N., Labrador at Hebron, about 60>E.,
in Kara Sea,? Atka, Aleutian Isis.
The species ranges from tidal zone to 500
meters in depth with a temperature gradientof 29. 0 7-64.0 5 F., (RATHBUK, 1925. pp. 255257, 1925, p. 270). The bottom materials from
where the species has been collected comprises
sands. pebbles and gravels and broken shells,
rarely silts (RATHBUS, 1925, pp. 255-257).

2. Hyas coardatus LE.\Cll, (RATHBUN,
1925, pp. 259-268), type locality, North
Sea.
From West (Lat. 70°251 N.) and East Greenland (Lat. 60°N.) to Hudson Strait and Bay,thence southward via Kamtchatka and Sakha·
lin Is!. to Yeso Strait and through the Sea ofJapan to Korea (Lat. 37°02 1 N.) ; subspecies.
11rsinus RATHBUN, Shangh:ti, Amoy (L"It.
24°30 1 N.)

Iceland, Arctic coast of Europe to Lon~.
49"30' N.
Corst of Siberia and northward, as far west
as Bennett lsi. (about 147"E.) as north as
76°501 N. to East Cape.
The species ranges from low water mark to
682 meters in depth with an exceptional recordof 1656 meters in depth (W. Atlantic). The
temperature ranges from 29. 0 8·52. 0 9 F. (RATHBUN, 1925, pp. 260-268). The bottom materials
comprises silts, sands, pebbles, gravels andshells (RATHBUN, 1925. pp. 260-268).
T. SAKAI (1934, p. 295) reported two female
specimens in the collection of the surveying
ship Oshioro-maru of the Fisheries Departmentof the Hokkaido University from the East
China Sea (loc. unknown). subspecies, altaceus
BRASDT. T. SAKAl states that Hyas coarctatus
has never been recorded from the Pacific sideof Honshu of Japan.

3. Hyas lyratus DAKA, (RATHBUN,
1925, p. 275), type locality, Oregon coast.
Bearing Sea, from a line connecting Bearing
Is!., Commander Isis., Siberia, with Pribilof

Rikizo lMAlZUMI

182

1sl. and Bristol Bay, Alaska (greatest Lat. 58°
38130" N.), southward to Admiralty Inlet,
Washington (about Lat. 48°N.).
This species ranges from 9 meters to 4~0
meters depth (RATHBUN, 1925, pp. 271-275).
"The bottom materials comprises silts, sands,
pobbles, gravels and broken shells (RATHBUN,
1925, pp. 271-275).

Geological and geographical distribu1:ion of the fossil species of the genus
Hyas ever recorded.
1. Hyas meridiottalis M. F. GLAESSSER,
1928, (text-fig . .!.: ), type locality,
Wenz2ldorf of Preding, Austria, sandy silt,
Helvetian, and Gamlitz, torton Leithakalk,~aden, Korallenkalk of Rauchstallbrun, coli.,
Joanneum Graz.

2. Hyas oranensis V. VAN STRAELE:-1,
1936, type locality, Oran,
Algeria, limy rock, Sahelian. depository, l\Iu·-seum national d'Histoire naturelle, Paris.

Remarks :-According to M. F. GLAEs"SENR's description of Hyas meridiotzalis,... Die Verwandten der Ubringen Tertiarfauna unseres Gebietes finden sich dagegen grosstenteils in tropischen Meeren,.zum geringeren Teil ins Mittelmeer."
The specimen of Hyas meridionalis isincomplete, its carapace is very convex,-especially in the gastric region, thehepatic region is dilated laterally but
less weiter than llyas coardatus. The-surface of Hyas meridionalis is covered
llyas araneus

Hyas coarctatus

Hyas lyratus

with granules and pustules, the pustuleson the cardiac region being very distinct.
Hyas tsuchidai is distin~ished from
Hyas meridiottalis by the ornamentati<;mof the carapace. The hepatic region islarger than the intestinal region in Hyastsuchidai but the rever3e in Hyas meri·diotzalis and from Hyas ormzensis by theornamentations of the gastric and cardiac
regions. It is also distinguishable fromthe Recent species of the genus Hyas asstated below, the rostrum of llyas araneus resembles Hyas tsuchidai but theone of Hyas tsuchidai is broader and theposterior angles of the hepatic projectionof the former species subacute but thelater has broadly rounded posterior
angles of the hepatic region. The ornamentation of the surfaces of the carapace
differs in both species.
From Hyas coarctatus, the shapes of
the carapace, the rostrum and the hepatic region and the ornamentation of
the carapace serve to separate Hyastsuchidai. The shapes of the proto- andthe meso-gastric regions are similar inboth Hyas lyratus and llyas tsuchidai•
Hyas tsuclzidai resembles Hyas araneus in the shapes of the gastric regionsand Hyas lyratus in the shape of therostrum.
Dimensions of the carapace of Recent
species are as follows,(RATHBUN, 1925, p. 258)
femalemale
male, Iceland(RATHBUN, 1925, p. 269)
male, altaceus, Grande Bandsmale, west Greenland
the largest specimen of America, malethe largest specimen known of the typical form,
male, Murman Sea(RATHBUN, 1925, p. 275)
male(length)72.4mm.
94mm.
UOmm.

(width)
54mm.
72mm.
86mm.

80mm.
99mm.
30.2mm.

64.5mm.
74mm.
197mm.

51.5mm.

35mm.

lOSmm.

80mm.

A Miocene Crab, Hyas lsuclzidai n. sp. from the Wakkanai

The length of the carapace of Hyas
cra11e11sis from the Sahelian of Oran,
Algeria is more than 30mm. and its'width more than 34mm. The fossil
species from the Helvetian of Austria
and from the upper Miocene of Japanare both small in size, and they may be.considered as dwarf species.

References
GLAESSNER, M.

F. (1928) Die Dekapoden
fauna des Ost. Jungertiiirs. ]alzrbuch. d.
Geol. Bundesamt. 78, pp. 198-200, Taf.
III, fig. 15.
- - (1929) Fossilium Catalogus, I: Animalia,

183

pars 41, Crustacea Decapoda. pp. 224, 225.
RATIIBUK, M. ]. (1925) The Spider Crabs ofAmerica. U. S. Nat. Mus. Bull. 129. pp.
252-275, J/yas ara11eus (L.), pis. 92, 93,
Hyas coarctalus LEACH, pis. 94-97, Hyaslyratus DAKA, pl. 235.
SAKAl, T. (1934) Brachyura from the Coastof Kyushu, Japan. Sci. Rep. Tokyo Bunrika
Daigaku, Sec. B, no. 25, p. 295.
(1939) Crabs of Japan, p. 94.
VAN STRAELE~. V. (1936) Sur quelques crabs
du Sahflien moyen des Environs d'Oran(Algerie), Bull. Soc. Geo!. FraiiCe, 5 ser.
t. VI, pp. 474, 475, pl. XXXIJI, fig. 2.
The Tempoku Coal-Field Hokkaido. ]oum.
Coal-Mines, Coal Assoc., Hokkaido (1950),
vol. 1. no. 1, pp, 15, 16.

184

Proceedings of the Palaeonto!ogical Society of Japan

Memorial to Thomas Way land VA UGH AN
By Shoshiro HANZA W A
Dr. Thomas Wayland VAUGHA~, oneof America's most distinguished scientists passed away on January 16, 1952
at the age of 81 in Washington, D. C.
He was born in Jonesville, Texas on
September 20, 1871.
After he received a B. S. degree from
Tulane Universit\' in 1889 and a l\I.A and a
Ph. D degn•es ai Harvard L'niversity in 189l
and 1903, r~sp..-ctively, he served as geologist
in the U.S. l;eological Survey from 1907 to
1939. He was also attached as paleontologist
to the National Museum, Washington. D.C.
from 1924 to 1Y42. He was Director of Scripp::
Institution of Oceanography. Cniversitr ofCalifornia at La Jolla, California in 1924-1936.
and was ;q>pointed Director Emeritu;' thereafter.

Dr. VAl'GHAS was devoted to thestudy of the Cretaceous and Tertiarylarger Foraminifera and Corals fromsouthern states of North America, Central and South Americas and the West
Indies, and of the Recent corals of
Hawaii and Torres Strait. He extended
his research to problems relating to con·figuration of the ocean bottom, marine
sediments, and ecology of marine organisms, and made a great contribution to
the coral reef problems.
He was the author of many new
species and several ne\'1 genera of
Cretaceous and Tertiary Foraminifera
and corals from the aforementioned
regions. In recognition of his great
paleontological achievement, his namewas dedicated to many foraminiferal
and coral species and genera by various
authors.
For his brilliant accomplishment in thescientific world, he was awarded an LL. D. by
the University of British Columbia in 1933 andby the University of California in 1936, anhonorary D. Sc. by Tulane University in 1944,
Agassiz medal of the National Academy ofAmerica in 19:{5, the Mary Clark Thompson
medal in 1945, and the Penrose medal of theGeological Society of America in 1946. Healso served as President of the GeologicalSociety of America in 1936.

His personal contact with the Japanesescientists begun in 1920 when he wasone of the American delegates to theFirst Pan-Pacific Science Congress, Honolulu. Hawaii. He came twice to Japan.
His first visit to Japan as one of theAmerican delegates to the Third Pan·
Pacific Science Congress. Tokyo was in1926. During his second visit to Japanon his trip around the world in 1933, he
was gracefully granted by Emperor
Hirohito of Japan in private audience.
He \vas awarded by the Japanese Government the Order of the Rising Sun, thirdclass and a beautiful vase cloissonne.
He was well acquainted with the historyof Japan and loved Japanese customs,
arts and the Chinese poems, moreover,
he ahvays endevored to understand theoriental people, and mastered the sixth
grade Japanese reader when he was
Director of the Scripps Institution of
Oceanography at La Jolla. He hadmany friends among the Japanese geologists, paleontologists, biologists, andoceanographers who all enjoyed his
friendship until his death. He wasactive in his field until 1947.

Trans. Proc. Palaeont. Soc. Japan N.S., No.5 pp. 185-193, pl. 17, May 30, 1952.

216. DESCRIPTIONS OF THE PALEOGENE MOLLUSCS FROMSAKITO, NAGASAKI PREFECTURE, JAPANn
ATSUYUKI MIZUNO
University of Tokyo"'*

Jij,~W:~?ffli!i: l'i~=Mfiit;{ l'l{t:Off K?lt· L: ft~~~Filll]'u.J'ili:Cf.> tlti1'Jlft.RottU·&.tL
1dl 1t:rll\fQ)-$ a: >l=i!:tt-r o MsH: I!Jt= Q)-<:. c. c. Kk t1Jt'i~IU: 'ffi',!f t... ~Q) 5 ~t~Q)xlt a:rr-:::>

t= 0 · Cra$sate/lites iesakai, Crassatellites clongatus. Ver.ericardia vestioides, Ve1lerica,·dia injlata,
Vcnericordia j"ponica. 7.1<lJ!f.liHr

Long ago a number of Paleogene mol1uscs of Northern Kyushu had been studied by M. YoKoYAMA (1911 and 1927)
and T. NAGAO (1928 a, b), but their
paleontological studies have not been somuch performed as compared with the-stratigraphical studies of this area, except the studies on fresh-water molluscs
by both T. UEJI (1934 a, b) and K. SuzuKI(1941).
For the following reasons, the writer
has considered some years that it must
be necessary to make clear the charactersof the fossils of this area ;1) The Japanese Paleogene strata
being typically distributed in the coal
fields of Northern Kyushu.
2) Many fossils, which were recently
collected from strata of "Paleogene " atseveral localities of Japan, being compared with the fossils from Northern
Kyushu.
3) The fossils of Nortl:ern Kyushu
being given very little attention except
by a few investigators since their pioneer works by both YoKoY Al\IA and

NAGAo and being never compared withrhe Paleogene species abroad.
4) Accordingly the analytical studiesof these fossils being very significant inmaking clear the characters of the Japanese Paleogene faunas and in establishing the biochronological studies of theJapanese Paleogene.
Through the efforts of Mr. Sadao
IEsAKA, a geologist to the Sakito Coalmine, Mitsubishi Mining Company, thewriter had a chance to inspect a part ofthe marine molluscan fossils collectedfrom the Nishisonoki group near the
Sakito Coal-mine. Here the writer explains some of the descriptions of them,but their ecological or chronologicalstudies will be given in future days.
The writer wishes to express manythanks to Mr. S. IESAKA for his offeringof the specimens and data. Thanks arealso due to Professor Takao SAKAMoToand Assistant professor Fuyuji TAKA! inthe Geological Institute, University of
TokYt> for looking over this manuscript.
According to H. l\iATsUsHITA (MATSUSHITA, 1949, p. 20), the stratigraphicalsuccession of the Paleogene near Sakito
is, in descending order, as follows;

1) Read Feb. 9. 1952; Received March 7,
1952.
185

186

Atsuyuki MIZUNo '1)
2)
Kurose group ............... Ushikubi formation
4)
i'hhima formation
3)
5)
Nishisonoki group ...... Kakino~)a formation{
Itanoura formation
8){Sakito coal-bearing
7)
formation
Matsushima group ......
9)
Nakado formation............ Unconformity ........... .
10)

Terashima group ......... Terashima formation
11)
12)
Akazaki group ............ Yobikonose formation. . . . . . . . . . . .Unconformity ........... .
Crystalline schists and granites

By T. SEKI's oral communication the
Oshima formation is subdivided into six
members; namely Higire13> alternationof sandstone and shale, Lima sandstone.
Tonool41 alternation of sandstone andshale, Tsurusakil5> sandy shale, Okuural6> shale and Fukuural71 tuff, in descending order.
Both the Nishisonoki and Matushimagroups are abundantly fossiliferous, thefossils being partly listed by H. MATSU·SHITA (MATSUSHITA, 1949, p. 21.) and thepresent ones were gathered from the
Oshima and Itanoura formations .
The specific names here determined
are shown in the following table.

List of Species ( x ...... Abundant)
x Turritella (s. str.) karatsuemis NAGAO ........................................................... 2 3
x Turritella (Ilaustator) sakitoe11Sis NAGAO .........................................................
3
Scala sp .................................................................................................... , ..... 2
Polinices sp. .................
.. ................................................................ 1
Neplunea (s. str.) chikuzenensis (NAGAO) ...................................................... 1
Dentalium sp .................................................................................................... 2
Barbatia sp ....................................................................................................... 2
Glycymeris (s. str.) sp .................................................................................. 1
Cre11ella subfornicata NAGAO ........................................................................... 2
xSeptijer (s. str.) nagaoi OYAMA ........................................................................ 2
Chlamys (Coraliclzlamys?) rutteni (MARTI!-<) ...................................................... 2
Rece11 (s. str.) sP.............................................................................................. 2
xCrassatellites (Eucrassatella?) iesakai 1\IJZUNo, sp. nov ........................................ 2
Crassatellites l Eucrassatella ?) elongatus MIZUNO, sp. nov. .. ............................... 2
Crassatellites (Eucrassatella) yabei NAGAO ......................................................... 2x Venericardia (Cyclocardia) vestitoides MIZUNo. sp. nov ....................................... 2
x Ve11ericardia (Cyclocardia) injlata MIZUKo, sp. nov. . ........................................ 2Ve~~ericardia (Cyclocardia) siogamensis No:.IURA ................................................ 2x Venericardia (Megacardita) jaPo11ica lVIJZUNo, sp. nov ........................................ 2
Cal/ista {s. str.) hmzzawai (NAGAo) .................................................................. 2
Pilar (s. str. ?) matszmzotoi (NAGAO) .............................................................. 2
Venerupis (Amigdala) sp ............................................................................... 2(18)
1 Hirabara, Kakinourashima, Sakito-machi.. ................... Okuura shale, Oshima formation.
(19)
(20)
2 Takamine and Yokoura, Sakito-machi ....................... Fukura tuff, Oshima formation.
( 21)
3 Nakado, Sakito·machi ........................
..lt'lnoura formation.
1) ~iii
9) !Jl.F16) ~ltll*·fa3) jl!.j:f&:ft
4)5) Milli6) iliiili7) t,;.fft
2) -'f-11
12) ll'f7·lt.it13) !HJJ
14) ;!§~
11) i'O;~~t10) 'WEb
19) iWiflt
20) Uiiili21) IJ1 J=i
17) ijii\iiifi18) lf-lhl

8)

*ljp

15) - -

Descriptions of the Paleogene Molluscs ficim SaKito, Na~asaki Prefecture, Japan
Amorig the fossils listed above, thespecifically identified ones are fifteen in
number consisting of five new species,
an upper Eocene one of Java, a Miocene
one of Japan, and the remaining eightthat are common with the " Ashiya fauna''. In most cases, shells are irregularly
deposited, detached and much crowded
in strata, especially in the Fukuura tuff.

187

mation. The former (fig. 5b) agrees
well with the holotype, while the latter(fig. Sa) is distinguished from it, inbeing lower in height, and having fewer
carinae on each whorls.
Occurrence : Itanoura formation andFukuura tuff, Oshima formation.
Buccinidae
Genus Neptzmea (BoLTEN) RooiNG,

Descriptions of some fossils

1798 (part)

GASTROPODA

Nept:mea (s. str.) clzikuzenettsis

Turritellidae(NAGAO)

Genus Turritella LAMARCK, 1799

1928.

Tzerritella (s. str.) sakitoensis NAGAo

Chrisodomus chikuzenemis, NAGAo: Sci.
Rep., Toltoku Imp. Univ., 2nd. Ser., Vol.
Xll, No. 1, p. 108 (98), pl. 18, figs.
10-12.

Plate 17, Figure 4.

Only one specimen, much deformed.
was obtained.
Occttrrence : Okuura shale.

1928.

Turritella sakitoensis, NAGAO: Sci.
Rep., Tohoku Imp. Univ., 2nd. Ser., Vol.
XII, No. 1, p. 101 (91), pl. 15, Figs.

1950.

Turritella (s. str.) s::zkitoensis, KoTAKA:
Short Paper. IGPS., No. 1, p. 33, pl.

BIVALVIA

5, Fig. 14.

Mytilidae

34·40.

Genus SePtifer RECLUZ, 1848

Several imperfect specimens were ob·tained.
Occurrence: Itanoura formation.

Septijer {s. str.) nagaoi OYAMA
Plate 17, Figures la-d.

Turritella (/laustator) karatsuensis
NAGAO
Plate 17, Figures Sa, b.
1928.

Turritel/a karatsuensis, NAGAO: Sci.
Rep., Tohoku Imp. Univ., 2nd. Ser. Vol.
Xll, No. 1, p. 99 (89), pl. 15, Figs.

23-26.
1950. Turritella( llaustator)

karatstlCIISis.

KOTAKA: Short, Paper, !CPS, No. 1,
p. 33, pl. 5, Fig. 15.
Specimens were mostly obtained fromthe Itanoura formation and only one
from the Fukuura tuff, Oshima for-

1927.

Mytilus ltirsutus, YoKOYAMA: ]our.
Fac. Sci., Imp. Univ. Tokyo, Soc. II, Vol.

If. p. 187, pl. 50, figs. 3, 4,
1928. Septifer (?) sp., NAGAO:

Sci. Rep.,
Tohoktt Imp. Univ., 2•zd. Ser., Vol. X II,
No. 1, p. 44 (34 1, pl. 1, figs. 7, 8.
1951. Septifer 11agaoi. OYAMA: Mi11. Geol.,Vol. IV, Nos. 1, 2, p, 56.

Description : Shell moderate or large.
obliquely elongate-trigonal, acute atanterior, rounded in posterior part;ventral margin broadly arcuated ; dorsal
margin nearly straight, shorter than thepreceding, making an obtuse angle with

Atsuyuki MIZUNO

188

broadly arcuated posterior margin on
about 1/2 of length of shell. Beaks
acute, terminal, situated at the anterior·extremity of shell. Surface sculptured
with radial riblets and concentric growthlines. the former somewhat wrinkled,diverged near the ends. Inner margin
finely crenulated.
Dimensio11s: Holotype (designated by
X. OYAMA) ... Length, 56.5 mm ; Height,
26.5mm.
Specimens at hand-

i
2
3

4

L. mm.
56.0
48.8
46.5
39.0

H. mm.
31.5
2o.3
29.0
30.6

ComParison : As described by K.
OYAMA S. nagaoi is similar to S. bilocularis (REEVE, 1858, pl. IX, fig. 42. ), a
living species, but they are different toeach other both in the surface-sculpture
and outline; the former having convex
ventral margin, the latter having nearlystraight or concave one.
The present species was first describt..<>dby M. YoKoYAMA from Hako-jima, Nishisonoki-gun being referred to Tricomyahirsuta (LAMARCK) and in the next year
by T. NAGAo to Septifer (?) sp. Recently,
K. Ov AMA gave a new name of S. nagaoi
to the specimens figured by them, designating the specimen shown by l\L
YoKoYAM,\ to the holotype. Several
specimens at hand all resemble th<:>m in.outline.
Observations on all specimens showthat the present species has two different
forms which can be clearly recognizedin the mature stage of growth ; all theimmature individuals show nearly sameform, while the mature individuals differcnciate into two fom1s as mentioned
above ; one is figured in Figs, 1a, lb.
which represents majority of the specimens, having slight variations in it andhaving a broader postero-dorsal part ;another is figured in Figs. 1c, ld, which
represents a few specimens, having anarrower one, as is shown by the halotype. They owe these variations to thedifference of the rate of growth on thepostero-dorsal part of shell.
Occurrence: Fukuura tuff.
Genus Crenella BRoWN, 1827

Crenella subfornicata NAGAo
1928.

Crmella subfornicata, NAGAo: Sci. Rep.,
Tohoku Imp. U11iv., 2nd. Ser., Vol. XII,
No. 1, p. 46 (36), pl. 5, figs. 8, 9.

Only one specimen, very small andill-preserved.
Occurrence: Fukuura tuff.
Pectinidae
Genus Clzlamys RoniNG, 1798

Chlamys ( Coralichlamys ?) mttmi
MARTIN
Plate 17, Figure 6.
1914.

Ch/amys (s. str.) rutteni. l\lARTIN:
Samml. Ceo/. Rcichs.-Mus. Lsiden, 11. f.,
Bd. II, HI. 4. p. 182, taf. 7 .. figs. 188,188a.

Only one outer impression of rightvalve which is broken in the anteriorpart, measuring 12.0 mm. in length and12.5 mm. in height.
C. rutte11i was described by K. MARTIN
from the upper Eocene Nanggulan formation in Java, and is distinguishedfrom the other species of Ch/amys inhaving a smaller shell and narrower
radial ribs which are twenty-seven in
number. The ribs are faintly nodose
regularly arranged, and separated bybroader vaileys. The present specimen
has the above-mentioned characters,Descriptions of the Paleogene Molluscs from Sakito. Naga>aki Prefecture, Japan

though larger than the holotype.
Occurrence: Fukuura tuff.
Crassatellitidae
Genus Crassatellites KRiiGER, 1823

Crassatellites (Eucrassatella? J iesakai
Mtzut-:o, sp. nov.
Plate 17, Figures 2a-d, Text-figure A.

DescriPtion : Shell small, somewhat
compressed, transversely oblong. inequilateral; ventral margin slightly arcuated, the anterior narrowly rounded;
posterior somewhat subtruncated ; antero- and postero-dorsal margins almost
symmetrically descending from the beak,making an angle of about 120" to eachother, the former nearly straight, thelatter faintly concave, shorter than the

Dimensions
Len~th(mm.)
Height (mm.)
Thickness (mm.)

former. Beaks not prominent, pointed ..
lying at about anterior 3/7 of shelllength. Surface sculpturd with coarse
concentric ribs, about 20, forming striking ridges.
Hinge relatively large,
triangular: teeth on the left valve as
follows; 2a thin, oblique ; 2b thin, prominent and nearly perpendicular ; clarge,
trianguiar, relatevely shallow, separated
into two areas by obsolete ridge on theposterior part of c. Inner margin ofshell not crenulated.

~- ..·· ...
~:·~c 2b 2 a
Text-figure A. Sketch showin~ thehinge of Crassalel/iles iesakai.

Paratype 1

Holotype

Paratype 2

P. 3

24.0
16.0

19.0
12.0
6.0

18.0
12.5
6.0

18.0
12.3
5.0

Comparison: C. iesakai is similar to
C. bellula (A. All.) (Cr.Rsst:-.:, 1886, p. 35,
taf. 6, fig. 6) and C. 11ames (An. etRvE.) (CLESS!N, 1886. p. 15. taf. 6. fig.

3) which are living now, but differs in
the following respects; C. namts having
a smaller and higher shell, and C. bellula
having finer growth-lines and the bothhaving crenulated inner margins. In
comparison with C. pauxi/la (YoKoYA~1A)(YoKoYAMA, 1925, p. 122, pl. 15, figs.
8-11.), described from the Tertiary of
Chichibu, Japan, the new species has
finer growth-lines, though they resembles
with one another in outline. C. yabei
NAGAo, the important species in the"Ashiya fauna ", can be clearly disting-

189

I
I

P. 4
16.-l
11.2
50

uished from the new species, in having
larger and more crooked shell and moresmooth surface.
Many specimens at hand are slightly
variable in outline. The phenomenon,
that is unrecognizable on a younger
shell. seems to be due to the differenceof rate of growth on some parts ofshell when individuals become adult, asin the case of Septifer 1zagaoi. The
specimen figured by T. NAGAO (NAGAo,
1928b, pl. 3, fig. 19. ), which was referred
to C. yabei by him, is considered as amature form of C. iesakai.
The name is dedicat~d to Mr. S.
IESAKA, who sent the specimens to thewriter.

190..

Atsuyuki· MIZUNO

Occurrence : Fukuura tuff.
Crassatellitcs ( Eucrassatella ?)elongatus MIZUNO, sp. nov.
Plate 17. Figure 3.

DescriPtion : Shell small, inflated,transversely elongated, ovate-trigonal;
ventral margin broadly arcuated, ob·liquely ascending to narrowly rounded
anterior margin, and rectangu.Jarly cut
by sharply truncated posterior margin ;Antero-dorsal margin concave, on about
1/2 of postero-dorsal margin in length,
the latter nearly straight, obliquely descending behind beak; from beak strik·
ing ridge running to postero-ventral
corner. Beaks prominent, pointed, incurved, slightly prosogyrate, lying at
anterier 1/3 of shell. Surface sculptured
with concentric ribs, equally coarse andstriking as those of the preceding species.
Lunule broad, deep and lanceolate ; escutcheon narrow, long. Inner margin
not crenulated.
Dimensiotzs: Holotype-Lentgh, 225
mm. ; Height, 13.0 mm. : Thickness,
8.6mm.

Comparison : C. elongatus is somewhat
similar to C. iucompiszms NAGAo fromthe Paleogene of Northern Kyushu, but
the new species is more inflated andmore oblique in form and more prosogyrate in beaks. C. inconspicuus (?)
NAGAo figured by T. NAGAO, (NAGAo,
1928, pl. 3, fig. 24) is referred to C.
~longatus.

Crassatellitcs ( Eucrassatella)ya.bei NAGAo
1928. Crassalellilcs yabei, NAGAO:

Sci. Rep.,
Tohokl4 Imp. Univ., 2nd Ser., Vol. XJJ,
No. 1, p. 48 (38), pl. 2, figs. 17-20,
23 ·28. pl. 3, figs. 5, 6, 10-12, 23-24.

Few imperfect specimens are at hand.
In comparison with the holotype they
are all smaller and have coarser ribs.
Occurrence: Fukuura tuff.

Carditidae
Genus Vcneri'cardia LAMARcK, 1801

Venericardia (Cyclocardia)
vcstitoides MIZU!'\o,sp. nov.
Plate 17, Figures 7a-c.

Description: Shell small in size, roundtrigonal, longer than high, inequilateral,
inflated at umbonal area ; ventral margin strongly arcuated ; antero-dorsal
margin slightly concave, shorter thanthe postero-dorsal, the latter somewhat
convex, gradually continuing to ventral
margin, and making an angle of about120" with the antero-dorsal at umbonal
area. Beaks large, prominent, pointed,
slightly prosogyrate, situated at anterior
2/5 of length of shell. Surface sculptured
with radial strong ribs and concentric
growth-lines. the former about 15, strongly noded, and separated by narrowerand deep valleys. Lunule short, deep;
e!'cutcheon not apparent.
Dimcmions: Holotype-Length, 28.0
mm. ; Height, 23.1 mm. ; Thickness (single
valve), 5.0mm. Paratype 1-Length, ?;Height, 23.5 mm; Thickness (s. v.),
5.0 mrn. Paratype 2- Length, 21.5 mm. ;Height, 19.0 mm. ; Thickness (s. v.),
3.5mm.
Comparison : V. vestitoides resembles
V. vestita (DESH.) (C!.ESSI!':, 1888, p. 32,
taf. 12, fig. 8, 9.), now living in the
boreal sea, in outline, but differs fromit in having a larger shell and fewer
ribs. Also it differs from V. matherotti(1\L>\YER) (CossMANN et PAYRoT, p. 60,

Descriptions of tbe Paleogene Molluscs from Sakito, Nagasaki Prefecture, Japan

pl.A, figs. 9-12.), the Helvetian species
in France, in having fewer ribs, though
they are similar to each other in. outline.
In comparison with V. mandaica (YoKoYA~IA) (YoKOYAMA, 1911, p. 9, pl. 2,
figs. 8-11.) from the lower Paleogeneof Northern Kyiishii, the present new
species is larger in apical angle, fewer
in number of ribs and more elongatedin its postero-ventral part.
Several specimens were obtained.
Occurrence : Fukuura tuff.

Venericardia ( Cyc!ocardia)
in/lata Mizm-:o, sp. nov.
Plate 17, Figures Sa. b.

Description : Shell, inflated, trigonal,
inequilateral ; ventral margin convex,
antero-dorsal margin o~?liquely descending from beak to make a small semicircle with ventral margin, the posterodorsal slightly convex, making an angleof about 90° with the antero-dorsal.
Beaks prominent, acute, incurved. Surface sculptured with radial ribs andconcentric growth-lines, the former 15
or 16 in number. nearly straight, separated by narrower valleys.
Dimensions: Holotype-Length, 21.4mm.; Height, 20.5 mm,; Thickness (single valve), 5.0mm.
Comparison : The new species resembles V. Velti1la (SMITH.) (CLESSW,
1888, p. 12, pl. I, figs. 7, 8.), a living
species near Mexico, in the strong inflation and number of ribs, but distinguished from it in more irregularoutline. V. siogamensis NoMURA is alsosimilar to V. in/lata, but the latt€r has
more trigonal form and fewer ribs.
A few specimens were obtained.
Occurrence : Fukuura tuff.

191

Vmericardia ( Cyclocardia)
siogamensis NoMURA
Plate 17, Figure 9.
1935.

Venericardia siog:mumsis, NoMURA :Saito Hoon-Kai Mus. Res. Bull., No. 6,
p. 212, pl. 17 (2), figs. 8-ll.

Several specimens at hand, which are
identified with V. siogamensis, are moreor less variable in outline, but resembles
the holotype in having a trigonal formand 18-20 ribs, though differs from it in
the convex postero-dorsaf margin. The
type specimen of the species was described from the ·• Upper shell-beds" inthe vicinity of Shiogama, Miyagi Pref.,Japan.
Occurrence: Fukuura tuff.

Venericardia ( Megacardita)japonica 11tzuNo, sp. nov.
Plate 17, Figures lOa, b, Text-figure B.

Description: -Shell small, moderately
inflated, transversely elongated, ovate,
inequilateral ; ventral margin broadly
arcuated, gradually ascending to anterior
and posterior margins, both being n~r­
rowly rounded nearly in the same degree; antero-dorsal margin very short, thepostero·dorsal very long, slightly convex.
Beaks prominent, pointed, incurved, prosogyrate, lying at anterior 1/4 of shell.
Surface sculptured with radial ribs 18
or 19 in number, separated by narrower
valleys. Hinge relatively small ; teeth
on right valve as follows; 3a approaching to lunule border, slightly oblique,small, short; 3b very oblique, elongatedback\vards, strongly prominent, somewhat acute at the posterior end.

192

Atsuyuki MIZUNoVery deformed specimens were obtained.
Occurrence: Fukuura tuff.

3a 3b
Text-figure B. Sketch showing thehinge of Vt11ericardia japo11ica.

Dimensions :-Holotype ...... Length, 25.6
mm.; Height, 18.5mm.; Thickness (single
valve), 6.0mm.
Comparison :-V. japonica differs fromV. ferruginosa in having longer form
and much ribs, and from V. laticostata(REEVE, 1845, pl. 7, figs. 36a--d.) in
being longer. V. jouamzeti dertolonga
SAcco, (SAcco, 1899, p. 11, tav. 3, figs. 15,16 ), the Tortonian and upper Helvetianspecies of Italy, resembles the new
species in outline, but the former has
broader and fewer ribs.
Observations of the holotype and many
paratype show that the young individalsof the species are shorter, more trigonalin outline and less inflated and thegrowth of shell accompanies the elonga·tion, rounding and inflation of shell.
The holotype is maturely grown form.
V. (M.) 11. sP. reported from the Kami·
yokose formation in Chichibu, Japan, by
K. WATANABE etc. (WATAKABE, ARAI &HAYASIII, 1950, p. 87. pl. 2. fig. 10) isthought as the similar species to the new
species.
Occurre11ce: Fukuura tuff.
Veneridae
Genus Cal/ista Pou, 1791

Cal/ista (s. str.) hanzawai (NAGAo)
1928. Macrocallista ha11zawai, NAGAO: Sci.
ReP .. 1olroku Imp. U11iv., 211d. Ser., Vol.
XII, No. 1. p. 69 (59), pl. 11, figs. 16.
18, 21-24, pl., 13, figs. 7, 8, 16 (?) 17.

Genus Pitar RoMF.R, 1857

Pitar (s. str.) matsumotoi (NAGAo)
1928. Pitaria m1tsumotoi. NAGAo: Sci. Rep.,
Tohokl4 Imp. Ut1iv. 2nd. Ser., Vol. XII,
No. 1, p. 73, (6), pl. 11, fig. 17, pl. 13.
figs. 4-6.

Several specimens at hand are scarcely
distinguished from the holotype of thepresent species shown by T. NAGAO.
Occurrence : Fukuura tuff.

References
CLESSIN, S. (188~) Die Gattung Crassatella
LAM .. Syst. Conch .. Cab.
- - (1888) Die Familie der Carditaceen., Syst.
Conch.·Cab.
CoSSMANN, M. & PEYROT, A. (1909) Conchologie Neogenique, de L'Aquitaine.
Tome II.
MATSUSHITA, H. (1949) Stratigraphical Studies
on the Paleogene in Northern Kyi:ishu.
Sci. ReP.• Fac. Sci, Kyushu Univ. Geo/.,Vol. Ill, No. 1.
NAGAo, T. (1928a) Paleogene Fossils of theIsland of Kyli>hil, Japan. Part 1. Sci.
Rep., Tohoku Imp. U11iv., 2nd. Ser., Vol.
9, No.3.
'1\AGAO, T. (1928b) Paleogene Fossils of theIsland of Kylishil, Japan, Part 2. Sci. Rep.,
Tohoku Imp. Uuiv., 2nd. Ser., Vol. 12,
No.1.
REEVE, L. A. (1845)Mon ograph of the Genus
Cardita. Conch. lc:>nica.
REEVE, L. A. ( 1858) Monograph of the Genus
Afytilus. Conch. lconica.
SAcco, D. F. (1599) I 1\Iolluschi dei Terreni
Terziarii del Piemonte e della Liguria Pt.
XXVI!.
SUZUKI, K. (19,U) The Paleogene Corbiculidsof North.western Kyi:ishii ]our. Fac. Sci.,

M IZllNo : Palaeogene Molluscs from Nagasaki

PLATE 17

2 d

5 a

5 b

9

8 b

Descriptions of the Paleogene Mollusc5 from Sakito, Nagasaei Prefecture, Japan

Imp. UttiV. TOkyO', Sec. II, Vol. VI, Pt. 3.
UEJI, T. (1934a) Lamprotula nojimemis n. sp.,a Fossil Unionid from the Kitamatsuura
Coal-Field. Northern Kyushu, Japan.

Ve11us Vol. IV.
- - (l934b) Fresh-Water Fossil Molluscs from
Kitamatsuura Coal-Field, Northern Kyushu. Japan, Venus Vol. IV.
WATANABE, K.. ARAI, J. & HAYASHI, T.
(1950) Tertiary Geology of the Chichibu
Basin. Bull. Chichibu Mus. Nat. Hist.

193

No.1.
YOKOYAMA, M. (1911) Some Tertiary Fossils
from the J\Iiike Coal Field. ]our. Col/.

Sci., Imp. Univ. Tokyo. Vol. XXVII. Art.
20.
- - (1925) Mollusca from the Tertiary Basinof Chichibu. ]our. Fac. Sci., Imp. Univ.

Tokyo, Se..:. II, Vol. II. Pt. 3.
- - (1927) Teritary Fossils from Western
Hizen. ]our. Foe. Sci., Imp. Univ. Tokyo,
Sec. II, Vol. II, Pt. 4.

Explanation of Plate 17
Figs. 4a-d. Septifer (s. sir.) nagaoi OYAMA
a. Internal mould of the right valve, x 1.
b. Internal mould of the right valve. x 1.
c. lntermal mould of the left valve, x 1. (Holotype designated by K. OYAMA, reproduced
from YOKOYAMA's figure, 1927, Pl. L, Fig. 4.)
d. Internal mou'd of the left valve. x 1.
Figs. 2a·d. Crassateflites (Eucrassatella ?) iesakai MIZUNo, sp. sp. nov.
a. Internal mould of the left valve, x 1.5.
b. lntenal mould of the right valve, x 1.5, the holotype.
c. Internal mould of the right valve. x 1.5.
d. Internal mould of the right valve, x 1.5.
Fig. 3. Crassafellifes (Eucrassatella ?) elot~gatus MIZUNO, sp. nov.
Internal mould of the left valve, x 1.5, the holotype.
Fig. 4. Tz4rritella (Hausiator) sakitoet1sis NAGAO
Internal mould.
Figs. Sa, b. Turritel/a (s. sir.) karatsuensis NAGAO
a. External cast from the Fukuura tuff. x 1.
b. A fragment of shell from the Itanoura formation. x 1.
Fig. 6. Chtamys (Coralichlamys? rutteni MARTI!'<
External cast of the right valve, x 1,5.
Figs. 7a-c. Vettericardia (Cycloc'lrdia) vestitoides l\-liZUNo, sp. nov.
a. Internal mould of the left valve, x 1.5.
b. Internal mould of the right vale, x 1, the holotype.
c. Internal mould of the left valve, x 1.
Figs. Sa. b. Veneric1rdia (Cycolcaadia) infiata MIZUNO, sp. nov.
a. Internal mould of the left valve, x 1.5.
b. Internal mould of the left valve, x 1.5, the holotype.
Fig. 9. Venericardia (Cyclocardia) siogammsis NoMURA
Internal mould of the right valve, x 1.5.
Figs. lOa, b. Vcnericardia (Megacardita) japonica Mizu:-;o, sp. nov.
a. Internal mould of the left valve, x 1.5, the holotype.
b. Internal mould of the left valve, x 1.5.
All the specimens are kept in the Geological Institute, University of Tokyo.

194

Proceedings of the Palaeontological Society of Japari

PROCEEDINGS OF THE PALAEONTOLOGiCAL SOCIETY
OF JAPANra *"i!i1~!1o/.~~~ 49 IIDf91J~J l!l'ifn 27 "1'2 H 9 13 ;fL1i1:k~~~1tltJ:tl!.r'{~:tli:.t·H: 1k"L ~1UI-.t <~~X- 14 45)o ~'Yiiiii1{·J!ftl::~/~iiillm~::X
O)iJl ~0On the Two New Species of Fossil
juglOtiS .............................. Seido ENDO
l,1JUIY1~3Mtfti~H~ii-3 (ft.m) .........

................................................ Jt£fi!il-~*FtiJilf!lfi{JfL.!It;'/!I(K-?v•L: ............ mJ/i~~
New Boullonia and other Fusulinids associated with Calcareous Algae from
Northern Tai (Siam~ (ft~) .......... ..
...... .. . .. ..... .... .... .... .. .. .... Kenji KoNISHI
The Lower Carboniferous Kakisako
formation of Southern Kyushu, witha Note on Some New Corals andFusulinids ............ Kametoshi KANMERA
Pennsylvanian Fusulinids from the
Akiyoshi Limestone ...... Ryuzo ToRIYAMAHalysites shikokuensis, sp. nov. from
Yokokurayama, K&hi-ken ... Mitsuo NoDA
A New Tertiary Species of Echinara.
chnius from Hokkaido (f\';~) .........
..................... lchiro HAYASAKA andMatsutaro SHIBATA,\'b}J;(~J:,*ttM.it:1l-{t?:it:: 8.-tv· -c.........
.. . .. ........................... :MH\ t-t • ~ J:IE pg~i!tii\!UUI'J~ o * ?7 -r ti--! f-lo 1tii ( ft~) ........................... ~ft JfJ • *~'H'Hri
The Discovery of New Miocene Fauna
in the Northern Part of Nishitagawa
Coal-Field, Yamagata Prefecture, japan( f\:J/0 ............ Toshimasa TA~AI and
Kenzo 0GASA WARA

Some Non-marine Fossils from thelower Cretaceous Formations of the
Yatsushiro District, Kyushu ............ ·.......................... Tatsuro MATSUMOTO
Belemnites from Lower Cretaceous
Deposits of Miyako District (f\';~) ...
.. . ...... .... .. ... ..... ............. Tetsuro HAN AI
Bu11kokuda {~/iii) .!o Bunkokudidae(f.lrf-1-) t;:-::>v•L: (f\:i'J,!) ........... .tJ';ft;Jl(J~J11li'fi<7.J~~*t!Ef.IHi-=~F1 (f\:.i'l\!) tJ-;ft;t'f.m!UI.Ml!o~l7)fiJifjjj&UCfi#.tl~t;:.l: o ~ft

(1-\:li'J\!) .................... tl,*+'J.lt-·flliJ! t-t
On the Two Fossil Elephants from
Kyushu Island, Japan ..................... .
............ Fuyuji TAKA & Masaaki INOUEr B *.11!!.~!¥~~ 591F~-~I::jR-tj 7.> B *il·~!lm~~~i*i~IWi~J l!g*.IJ 21 (.f. 4 n 6 B~i-*~!¥~~~iii\1::1ft-c!»ltll! :t 0 ~~iiii~·]ftl::ff~iiit:Hill ;:}( O)f.lfl ~ o*~~7¥iitfllf..i¥itt.&rFJ!'t.f¥t.lllt-=aNt o :f.iR!k~o;lffifl ..................... ~.~iE~ • 'f':i!!!7i~~~J:IlJJt!!A!i*iR..&U:JJf-kiR 17)tfjit!k.. ...........................................

......
~JII~$!iH'-~ 'JafJIJ~.* f'Ftl:ft!!/517) W14!!kit:-l'i t;:

R..tv· -c ................................... -TW:iftii!.;
Astriclypeus K:V,h· -c .................. ~1' ,11,~~if!Jit'tf.t Yoldia Mr.:N..t"·'t:' ............. ..
.. .... ...... ...... ............ J,l..!ti-AA • f! ft t1\Lebens spuren 17)if~ (!~o l) ..... JijllHi!~
Porltmites t. p)UJij- OvaliPes ......... -q-*:bil:idfl~ti1§$l: !J }f.lfi L1tiffftlt~

Slego.

do11 KN'.tv•'t:' ............ i!i~~-= • :#fJ:lE!ig
7 ;;r. -<: ;;r. ~ Jl.- ;;r. om:~(~ 2 'ft!).. ..........

~$k~-~~~-=-~miE=·~oo~~~~--~-~. ~W*&DoU~fiDK-?v· ·c (1t.l%) .......................... J.f<!J!f.l!Htna!~~IJim 17)flt~1~::u~e~"· 1:

Discovery of Cretaceous Ammonites
from the undivided Mesozoic Complex
o! Shikoku ...... Tatsuro MATSUMOTO,
Toshio KIMURA & jiro KATTO¥~~MU.l:!J~~-tfti~re~~U:·A~:'ti:i~~lri'/-Ofii~ff......... ~ni~~........................ ?kfti!IJIJ

1Xwo~Ju ~Q')~n*ll~hl!l!oJ{t::Ut::ftv· -c............... liiiiBP!!~.................... .

Announcement
Bibliography of Japanese Palaeontology and Related Sciences

1941-1950
including papers published by Japanese authors as well as thoseby others which concern Japan and its surroundingsCompiled by

Ryuji Endo, D. Sc.
Pr~fessor of Geology and Palaeontology
Saitama University, Urawa, Japanjust published as a non-periodical publication of the Society.
Foreign subscribers free; foreign non-subscribers US $1.50
Domestic subscribers Yen 300; domestic non-subscribers Yen 400(Mailing charges inclusive in all cases)
Orders to be addressed to

the Palaeontological Society of Japan
Geological Institute, Faculty of Science, University of Tokyo, Japan.
~---

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1952 ~- s fJ 31 a~ft

JE fl <~fUf-O)/i4:51Xl:.J.R lJ) 1 t~i) 200 !LU
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CONSTITUTION

of the
PALAEONTOLOGICAL SOCIETY OF JAPANARTICLE 1. Name
The Society shall be known as the Palaeontological Society of Japan. The Society is a sectionof the Geological Society of Japan.
ARTICLE 2. Object
The object of the Society shall be to promote the study of palaeontology and related sciences.
ARTICLE 3. Achievement
The Society in order to execute Article 2 shall ( a) issue the Society journal and other publications, (b) hold or sponsor scientific lectures and meetings, and (c) sponsor collecting or
field trips, and lcctm es.
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The Society shall be composed of persons who are active of interested in palaeontology or
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ARTICLE 6. Administration
The Society shall have the following organizations for its administration.
(a) General meeting. The general meeting shall be composed of the Society members. Morethan one tenth of regul:!r members shall be present to hold general meetings. Administrative :~ffairs shall he decided during the general meeting.
(b) President. The president shall be elected from among the regular members. The president
shall represent the Society and supervise its business matters.
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(e) Officers shall be elected by vote of returned mail ballots, as a general rule.
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By-Laws and Administration
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Addendum
ARTICLE 1. There shall he four business councillors for the present.
ARTICLE 2. The Society journal shall be is3ued twice a year for the present.