Bull. Natl. Mus. Nat. Sci., Ser. A, 33(4), pp. 153–174, December 21, 2007

Eight New Spiders of the Families Hahniidae, Theridiidae, Linyphiidae
and Anapidae (Arachnida, Araneae) from Japan
Hirotsugu Ono
Department of Zoology, National Museum of Nature and Science,
3–23–1, Hyakunin-cho, Shinjuku-ku, Tokyo, 169–0073 Japan
E-mail: ono@kahaku.go.jp

Abstract Eight new species of spiders of the families Hahniidae, Theridiidae, Linyphiidae andAnapidae (Arachnida, Araneae) are described from Japan: Hahniidae: Cryphoeca shinkaii sp. nov.
(from Nippara, Okutama-cho, Nishitama-gun, Tokyo, Honshu) and Cryphoeca shingoi sp. nov.
(from Mt. Odaigahara-zan, Kami-kitayama-mura, Yoshino-gun, Nara Prefecture, Honshu);
Theridiidae: Pholcomma tokyoense sp. nov. (from Akasaka, Minato-ku, Tokyo, Honshu) andAllothymoites kumadai gen. et sp. nov. (from Kouchi, Geinou-cho, Age-gun, Mie Prefecture, Honshu); Linyphiidae: Walckenaeria iriei sp. nov. (from the bank of Honmyou-gawa River, Isahaya-shi,
Nagasaki Prefecture, Kyushu), Kagurargus kikuyai gen. et sp. nov. (from Mt. Kagura-yama, Saekishi, Oita Prefecture, Kyushu), and Meioneta tenuipes sp. nov. (from Aso, Taiki-cho, Watarai-gun,Mie Prefecture, Honshu); Anapidae: Mysmenella ogatai sp. nov. (from Ikegane-cho, Okazaki-shi,
Aichi Prefecture, Honshu). Two new genera are established both by monotypy: Theridiidae:
Allothymoites gen. nov. (type species: Allothymoites kumadai sp. nov.); Linyphiidae: Kagurargus
gen. nov. (type species: Kagurargus kikuyai sp. nov.).
Key words : Taxonomy, Araneae, Hahniidae, Theridiidae, Linyphiidae, Anapidae, Japan.

Although the spider fauna of Japan has beenquite well investigated in comparison with thoseof other Asian countries (F. and J. Murphy,2000), many species are left unstudied in somegroups, especially those with small body size.
The present report deals with the result of ataxonomical study on eight species of small
spiders (body length less than 3 mm) of thefamilies Hahniidae, Theridiidae, Linyphiidae
and Anapidae from Japan. The material used
for the study was selected from the specimens
contributed to the arachnid collection of the
National Museum of Nature and Science, Tokyo,
by many arachnologists and friends of thepresent author.
Cryphoeca Thorell, 1870, is a small groupcomposed of ten species mainly from the
Palearctic region and two species of these havebeen recorded from Japan, that is, C. silvicola (C.
L. Koch, 1834) and C. angularis S. Saito, 1934.

Palearctic Cryphoeca silvicola was recorded forthe first time by Yaginuma and Nishikawa (1969)
from Mt. Ontake, 2500 m alt., Gifu Prefecture,
Honshu, Japan, while C. angularis was describedby Saito (1934) from Sapporo, Hokkaido. However, the latter species has never been recognizedsince its original description. In the present material from Japan, further two new species of thegenus has been discovered.
Pholcomma Thorell, 1869 (Theridiidae) is alsoa small group including several species from Europe, North America and Australia. Since all the
known members of the genus from Japan weretransferred to other genera, it is though that thegenus does not occur in Japan (Yoshida, 2003).
During faunal researches in the Akasaka Imperial
Gardens, Tokyo, made by the National Museumof Nature and Science, some specimens of an
unknown spider were obtained, which seem toreally belong to the genus in question (Ono, 2005).

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Hirotsugu Ono

A peculiar theridiid collected by Mr. Kenichi
Kumada from Mie Prefecture has been entrusted
to the present author for study. The spider is regarded to belong to a new genus standing closeto Thymoites Keyserling, 1884.
Although Mysmenella jobi (Kraus, 1967)(Anapidae, Mysmeninae) was long known to be
the only species of the genus in Japan (Yaginuma, 1986; Shinkai, 2006), another species of thegenus was recently found in Aichi Prefecture by
Mr. Kiyoto Ogata. This spider lives between
grasses, while Mysmenella jobi spins her web inthe vacancy of the ground. After a careful examination of the specimens the present author recognized that the newly found Japanese spider resembles to Mysmenella gongi Yin, Peng et Bao,
2004, described from China, rather than to M.
jobi, but differs in details from the Chinese
species.
Linyphiidae is the largest family in spiders,containing more than five hundred genera and six
thousand species mainly in Northern Hemisphere. Of these, about three hundred species of
98 genera are known in Japan (Ono, Kamura andNishikawa, 1999), but there is a strong possibility that more new species and records should be
added to these numbers (Saito and Ono, 2001;
Ono and Saito, 2001). A new genus and three
new species are selected from unclassified specimens preserved in the collection of the National
Museum and featured herein.
Thus, two new genera and eight new species of
spiders are described in this paper.
The abbreviations used are as follows: ALE,
anterior lateral eye; AME, anterior median eye;
PLE, posterior lateral eye; PME, posterior median eye. The type specimens of the new species
are deposited in the collection of the Department
of Zoology, National Museum of Nature and Science, Tokyo (NSMT-Ar).
Taxonomy
Family Hahniidae
Genus Cryphoeca Thorell, 1870

Cryphoeca shinkaii sp. nov.
[Japanese name: Musashi-sumi-tanagumo](Figs. 1–15)

Diagnosis. This new species can be easily
distinguished from all other congeners by theshort embolus of male palp. In general appearance, it strikingly resembles the next new
species, Cryphoeca shingoi sp. nov., but differsfrom the latter in details of the male palp, especially in the shape of embolus (cf. Figs. 13, 26),
and the structure of female genitalia (cf. Figs.
5–7, 17–18).
Material examined. Holotype: male from
Nippara, 1025 m alt., Okutama-cho, Nishitamagun, Tokyo, Japan, 6–IV–2006, S. Hatsushibaleg. (NSMT-Ar 6958); allotype: female, same locality and collector as for the holotype,
13–IV–2007 (NSMT-Ar 6959); paratypes: 5 females and 4 males, same data as for the allotype(NSMT-Ar 6971–6978).
Description (holotype and allotype). Measurements: Body length female 2.75 mm, male 2.72mm; prosoma length female 1.24 mm, male 1.50m, width female 0.80 mm, male 0.92 mm; opisthosoma length female 1.42 mm, male 1.20 mm,
width female 1.01, male 0.82 mm; lengths oflegs [total length (femurpatellatibiametatarsustarsus)]: female I 3.02 mm (0.810.380.78
0.680.37), II 2.88 mm (0.810.360.670.67
0.37), III 2.58 mm (0.720.330.520.66
0.35), IV 3.36 mm (0.900.360.790.890.42),
male I 3.84 mm (1.050.410.970.860.55), II
3.55 mm (0.980.400.84+0.850.48), III 3.18
mm (0.880.360.670.830.44), IV 4.21 mm(1.130.380.991.110.60).
Prosoma (Figs. 1, 8): Carapace much longer
than wide (length/width female 1.55, male 1.63
in ratio), flat, with some hairs, median furrow
distinct. Eyes compactly set (Figs. 2, 9), anterior
and posterior eye rows procurved in frontal view,straight in dorsal view, PMEPLEALEAME
in female (3.7 : 3.5 : 3.5 : 1 in diameter), PLE
PMEALEAME in male (3 : 3 : 3 : 1 in diameter), AME-AMEAME-ALE (4 : 5 in female, 3: 5 in male), PME-PMEPME-PLE (1 : 1 in fe-New Spiders from Japan

155

Figs. 1–7. Cryphoeca shinkaii sp. nov. [1–3 & 5–7, female (allotype: NSMT-Ar 6959), 4, female (paratype:
NSMT-Ar 6976)]. — 1, Body, dorsal view; 2, eyes, frontal view; 3, chelicera, ventral view; 4, opisthosoma,
dorsal view; 5, epigynum, ventral view; 6, inner organ of female genitalia, dorsal view; 7, same, ventral view.
[Scales: for Figs. 1 & 4, 0.5 mm, for Fig. 2, 0.2 mm, for Figs. 3, 5–7, 0.1 mm.]

male, 2 : 1 in male), median ocular area slightly
wider than long (length/width female 0.94, male0.92), wider behind than in front (anterior
width/posterior width female and male 0.34),
clypeus much shorter than width of eye area.

Chelicerae (Figs. 3, 10) with three teeth on promargin of fang furrow and five teeth on retromargin, labium wider than long (length/width female
0.66 male 0.70), sternum with long hairs, slightly
longer than wide or as long as wide in female

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Figs. 8–15. Cryphoeca shinkaii sp. nov., male (holotype: NSMT-Ar 6958). — 8, Pro- and opisthosomata (appendages omitted), dorsal view; 9, eyes, frontal view; 10, chelicerae, ventral view; 11, femur, patella and tibiaof leg I, prolateral view; 12, palpal organ, ventral view; 13, same, retrolateral view; 14, tibia of palp, dorsal
view; 15, embolic division, prolateral view. [Scales: for Fig. 8, 0.3 mm, for Figs. 9–11, 0.2 mm, for Figs.
12–15, 0.1 mm.](length/width 1.07 in female, 1.00 in male).
Legs: Conspicuous trichobothria on tibiae,
metatarsi and tarsi, tarsal claws of legs with twoor three small teeth. Spiniformation: Female:femora I–IV dorsally 0–1–1–1, I prolaterally
0–0–0–1; patellae I–IV dorsally 1–0–1 (apical);tibiae I–IV dorsally 1–1, prolaterally 0–1–0–1(I), 0–1–1–0 (III–IV) or 0–1–1–1 (II), retrolaterally 0–1–1 (I, III–IV) or 0–1–0 (II), ventrally
2–2–2–0 (I–II), 0–1–2–0 (III) or 1–1–2 (IV);metatarsi I–IV prolaterally 0–1–0–1 (I), 0–1–1–1(II) or 0–1–1–0 (III–IV), retrolaterally 0–0–0–1

New Spiders from Japan(I), 0–1–0–2 (II–III) or 1–0–1–2 (IV), ventrally
2–2–0 (I–III) or 2–2–1 (IV); male (Fig. 11):femora I–IV dorsally 0–1–1–1 (I–III) or 0–1–1–2(IV); patellae I–IV 1–0–1 (apical); tibiae I–IV
dorsally 1–0–1, prolaterally 1–1, retrolaterally
1–0 (I–II) or 1–1 (III–IV), ventrally 2–2–2–2(I–II), 2 (III) or 1–1–2 (IV); metatarsi I–IV prolaterally 1–1–1–1 (I–II) or 1–2–0–2 (III–IV),retrolaterally 0–1–0–1 (I), 0–1–0–2 (II), 1–1–0–2(III) or 0–2–0–2 (IV), ventrally 2–2–0 (I), 2–2–1(III) or 2–2–2 (II, IV). Leg formula: IV–I–II–III.
Male palp (Figs. 12–15): Tibia with strongly
sclerotized apophyses: the retrolateral one withlarge tooth (Fig. 13), dorsal one distinct (Fig.
14). Cymbium simple with sclerotized, retrolateral margin, palpal organ large with expanded
tegulum (Fig. 12) and spiniform tegular apophysis (Figs. 12–13), embolus short, with sharply
narrowed tip (Fig. 15) and conducting groove.
Opisthosoma (Figs. 1, 4, 8): Ovate, longer thanwide (length/width female 1.41, male 1.46), its
dorsum wholly covered with long hairs. Spinnerets short and cylindrical, colulus present.
Female genitalia (Figs. 5–7): Genital field relatively large, epigynum (Fig. 5) with median septum and well-visible inner organ. Paired genital
openings situated at posterior part of epigynum.
Spermathecae reniform, intromittent canal very
short, large club-shaped process present (Figs.
6–7).
Coloration and markings: Female and male:
carapace blackish brown with yellow sides andblack head. Chelicerae yellowish brown, maxillae, labium and sternum light yellow, legs andpalps yellowish brown. Opisthosoma dorsallydark gray, with white markings (indistinct inholotype), venter light gray, spinnerets light yellowish brown.
Variation. Body length of the paratypes: females 3.00–3.18 mm, males 2.70–2.83 mm. The
markings of opisthosoma are variable: typical is
the pattern as given in Fig. 1 in females andmales, but those of some males are dark andwithout distinct white markings (Fig. 8) and contrary to this, the opisthosoma of some females
are light-colored as in Fig. 4.

157

Distribution. Japan (at present known only
from the type locality).
Etymology. The species is dedicated to Mr.
Eiichi Shinkai, an excellent spider researcher ofTokyo.
Cryphoeca shingoi sp. nov.
[Japanese name: Oodai-sumi-tanagumo](Figs. 16–28)

Diagnosis. See the diagnosis of the former
new species.
Material examined. Male holotype from Mt.
Oodaigahara-zan, 1500 m alt., Kami-kitayamamura, Yoshino-gun, Nara Prefecture, Honshu,
Japan, 22–VI–2005, S. Hatsushiba leg. (NSMTAr 6810); female allotype: same data as for theholotype (NSMT-Ar 6811).
Description (holotype and allotype). Measurements: Body length female 2.43 mm, male
2.80 mm; prosoma length female 1.23 mm, male
1.43 mm, width female 0.77 mm, male 0.84 mm;
opisthosoma length female 1.06 mm, male 1.31
mm, width female 0.86, male 0.90 mm; lengthsof legs [total length (femurpatellatibia
metatarsustarsus)]: female I 3.13 mm (0.88
0.380.820.680.37), II 3.10 mm (0.800.38
0.860.680.38), III 2.49 mm (0.670.31
0.530.620.36), IV 3.36 mm (0.940.39
0.790.810.43), male I 3.68 mm (0.990.38
0.940.840.53), II 3.47 mm (0.980.390.84
0.810.45), III 3.00 mm (0.800.340.68
0.750.43), IV 4.15 mm (1.120.351.03
1.090.56).
Prosoma (Figs. 16, 19): Carapace much longer
than wide (length/width female 1.76, male 1.70),
flat (Fig. 20), with some hairs, median furrow
distinct. Eyes compactly set (Figs. 21), anterior
and posterior eye rows procurved in frontal view,recurved or straight in dorsal view, PME
PLEALEAME in female (4 : 4 : 3.5 : 1 in diameter), PMEPLEALEAME in male (6 :5 : 4 : 1 in diameter), AME-AMEAME-ALE(5 : 6 in female, 3 : 4 in male), PME-PMEMEPLE (1 : 1 in female, 6 : 5 in male), median oculararea as long as wide (length/width female and158

Hirotsugu Ono

Figs. 16–18. Cryphoeca shingoi sp. nov., female (allotype: NSMT-Ar 6811). — 16, Pro- and opisthosomata(appendages omitted), dorsal view; 17, epigynum, ventral view; 18, inner organ of female genitalia,
dorsal view. [Scales: for Fig. 16, 0.2 mm, for Figs. 17–18, 0.1 mm.]male 1.00), wider behind than in front (anterior
width/posterior width female 0.27, male 0.30),
clypeus much shorter than width of eye area.
Chelicerae (Fig. 23) furnished with two teeth(with a indistinct denticle) on the promargin offang furrow and five teeth on retromargin, labium
wider than long (length/ width female 0.65 male0.67), sternum with long hairs, longer than wide(length/width 1.25 in female, 1.18 in male).
Legs: Conspicuous trichobothria on tibiae,
metatarsi and tarsi, tarsal claws of legs with twoor three small teeth. Spiniformation of legs: Female: femora I–IV dorsally 0–1–1–1, I prolaterally 0–0–0–1; patellae I–IV dorsally 1–0–1 (apical); tibiae I–IV dorsally 1–1, prolaterally 0–1–
0–1, retrolaterally 0–1–0–1 (I–II), 0–0–1 (III) or
0–1–1 (IV), ventrally 2–2–2–2 (I), 2–2–2 (II), 1–

1–2 (III) or 1–3–0 (IV); metatarsi I–IV prolaterally 0–1–0–1 (I–II), 1–2–2 (III) or 0–2–2 (IV),retrolaterally 0–1–0–1 (I–II), 0–2–2 (III) or 1–1–
2 (IV), ventrally 2–2–2 (I–II) or 2–2–1 (III–IV);
male: femora I–IV dorsally 0–1–1–1, I and IV
prolaterally 0–0–0–1, IV retrolaterally 0–0–0–1;
patellae I–IV 1–0–1 (apical); tibiae I–IV dorsally
0–1–0–1, prolaterally 0–1–0–1 (I–II) or 1–1 (III–IV), retrolaterally 0–1–0–1 (I), none (II), 0–1(III) or 0–1–1 (IV), ventrally 2–2–2–2 (I–II), 3(III) or 1–1–2 (IV); metatarsi I–IV prolaterally
0–1–1–1 (I), 0–1–0–1 (II) or 1–2–0–2 (III–IV),retrolaterally 0–1–1 (I), 0–1–0–2 (II), 1–1–0–2(III) or 1–1–1 (IV), ventrally 2–2–2 (I–II), 2–2–1(III–IV). Leg formula: IV–I–II–III.
Male palp (Figs. 25–28): Tibia with strongly
sclerotized apophyses: the retrolateral one with

New Spiders from Japan

159

Figs. 19–28. Cryphoeca shingoi sp. nov., male (holotype: NSMT-Ar 6810). — 19, Pro- and opisthosomata (appendages omitted), dorsal view; 20, carapace, lateral view; 21, eyes, frontal view; 22, maxillae, labium andsternum, ventral view; 23, chelicerae, ventral view; 24, spinnerets, ventral view; 25, palpal organ, ventral
view; 26, palp, retrolateral view; 27, tibia of palp, ventral view; 28, patella and tibia of palp, dorsal view.
[Scales: for Figs. 19, 22, 0.2 mm, for Figs. 20–21, 23–28, 0.1 mm.]

digitiform tooth (Fig. 27), dorsal one distinct andvery large (Figs. 27–28). Cymbium simple withsclerotized, retrolateral margin, palpal organ
large with expanded tegulum (Fig. 12) and without tegular apophysis (Figs. 25–26), embolus

short, with truncate tip (Fig. 25), conducting
groove large.
Opisthosoma (Figs. 16, 19): Ovate, longer thanwide (length/width female 1.23, male 1.45), its
dorsum wholly covered with long hairs. Spin-

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Hirotsugu Ono

Figs. 29–31. Pholcomma tokyoense sp. nov., male (holotype: NSMT-Ar 6981). — 29, Pro- and opisthosomata(appendages omitted), dorsal view; 30, palpal organ, ventral view; 31, same, retrolateral view. [Scales: forFig. 29, 0.5 mm, for Figs. 30–31, 0.15 mm.]nerets short and cylindrical, colulus present (Fig.
24).
Female genitalia (Figs. 17–18): Genital fieldwider than long, epigynum (Fig. 17) with a median septum indistinct and well-visible inner organ.
Paired genital openings situated at middle ofepigynum. Spermathecae reniform with very
short intromittent canal and with small, granulate
gland (Fig. 18).
Coloration and markings: Female: carapace
beige with white sides and black head. Chelicerae light yellowish brown, maxillae and labium
light yellowish brown, sternum, legs and palps
milky white. Opisthosoma dorsally gray without
distinct markings, light gray ventrally, spinnerets
darker. Male: carapace blackish brown withwhite sides and black head. Chelicerae yellowish
brown, maxillae and labium yellow, sternum dull
yellow, and marginated with black, femur andpatella of palp reddish brown, tibia and tarsus
yellowish brown, legs yellowish brown. Opistho-soma dark gray dorsally, with distinct white
marking, light gray ventrally, spinnerets yellowish white.
Distribution. Japan (at present known only
from the type locality).
Etymology. The species is dedicated to thecollector of the type specimens.
Family Theridiidae
Genus Pholcomma Thorell, 1869
Pholcomma tokyoense sp. nov.
[Japanese name: Toukyou-kabuto-himegumo](Figs. 29–31)
Pholcomma sp.: Ono, 2005: 451.

Diagnosis. This new species resembles thetype species of the genus, Pholcomma gibbum(Westring, 1851), known from Europe, but canbe distinguished from the latter by the structureof male palpal organ, especially the shorter em-

New Spiders from Japanbolus (cf. Figs. 30–31 and Wiehle, 1937, p. 218,
fig. 277 or Levi, 1957, p. 109, figs. 15–16).
Material examined. Holotype: male from the
Akasaka Imperial Gardens, Minato-ku, Tokyo,
Japan, 27–II–2004, S. Nomura leg. (NSMT-Ar
6981); paratypes: two males, same data as for theholotype (NSMT-Ar 6982–6983).
Description (holotype). Measurements: Body
length 1.23 mm; prosoma length 0.56 mm, width
0.48 mm, height 0.30 mm; opisthosoma length
0.79 mm, width 0.59 mm; lengths of legs [total
length (femurpatellatibiametatarsustarsus)]: I 1.57 mm (0.490.190.350.270.27),
II 1.26 mm (0.380.170.280.220.21), III
1.18 mm (0.350.150.240.220.22), IV 1.55
mm (0.480.170.370.250.28).
Prosoma: Carapace longer than wide (length/
width 1.17), relatively high, with several longsetae at the middle, median furrow absent. Eyes:
PMEPLEALEAME (6 : 5 : 5 : 3 in size), theanterior eye row recurved, the posterior one
procurved in dorsal view, lateral eyes close toeach other, AME-AMEAME-ALE (7 : 2),
PME-PMEPME-PLE (7 : 3), median oculararea wider than long (length/width 1.75), wider
behind than in front (anterior width/posterior
width 0.65), clypeus longer than AME-AME (3 :1). Chelicerae with two small teeth on promarginof fang furrow; labium not fused with anterior
margin of sternum, triangle in shape, wider thanlong (length/width 0.85); maxillae short, as longas labium, without marginal hairs; sternum
slightly wider than long (length/width 0.94), its
posterior margin truncated and wide. Legs relatively short and thick, without spines; leg formula: I–IV–II–III.
Male palp (Figs. 30–31): Tibia simple andshort, with long spines and trichobothrium retrolaterally. Cymbium an oval cup, distally modified
with digitiform process (paracymbium); tegularapophysis wide plate with a small tooth apically,
median apophysis spiniform, conductor with distinct tooth; embolus long spiniform and winding,
without any projection at base.
Opisthosoma: Ovate, longer than wide (length/
width 1.34), dorsally forming sclerotized plate,161with many hairs. Colulus: Large, anterior spinnerets the largest, thick and short, posterior ones
developed and conical.
Coloration and markings (Fig. 29): Carapace
yellowish brown, ocular area blackish, chelicerae, maxillae and labium yellowish brown, sternum yellowish brown without any markings, legs
pale yellow. Opisthosoma shiny black dorsally,
mat blackish brown ventrally.
Distribution. Japan (at present known only
from the type locality).
Etymology. The specific epithet is derived
from the name of the city.
Remark. Female is unknown.
Genus Allothymoites nov.
[Japanese name: Harabiro-sasahimegumo-zoku]

Type species. Allothymoites kumadai sp. nov.,
by monotypy.
Diagnosis. The new genus belongs to thesubfamily Therdiinae referring to the conditionof cymbium and should be standing close to thegenus Thymoites Keyserling, 1884, in body size
less than 1.5 mm and in the ratio of the length ofpatella and tibia of the first leg to the length ofcarapace 1.03–1.14, as well as the dark colored
opisthosoma without distinct markings. However,the new genus is recognized as an independent
genus by the combination of following characteristics: Carapace with warty clypeus, but without
distinct projection, extremely large eyes, chelicera with two teeth on promargin, presence oflarge tegular apophysis and hamulus-shaped median apophysis of male palp, opisthosoma triangle-shaped, epigynum with strongly sclerotizedhole of genital opening with expanded posterior
margin, spermathecae paired with long fertilization tubes, booklungs presumably without function, and absence of colulus.
Etymology. The generic name is formed by a
combination of a Greek word allos meaning another and the name of the existent genus, Thymoites, which is Greek thymos-ites and meansa creature with warty excrescence. The gender ismasculine.

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Hirotsugu Ono

Figs. 32–39. Allothymoites kumadai gen. et sp. nov. [32–34, female (allotype: NSMT-Ar 6986), 35–39, male(holotype: NSMT-Ar 6984)]. — 32, 35 Pro- and opisthosomata (appendages omitted), dorsal view; 33, 37,
prosoma, frontal view; 34, opisthosoma, ventral view; 36, prosoma, lateral view; 38, chelicera, ventral view;
39, maxillae, labium and sternum, ventral view. [Scales: for Figs. 32–37, 39, 0.2 mm, for Fig. 38, 0.1 mm.]

Allothymoites kumadai sp. nov.
[Japanese name: Harabiro-sasa-himegumo](Figs. 32–44)

Diagnosis. See the above generic diagnosis.
Type specimens. Holotype: male from Hounami, Kouchi, Geinou-cho, Age-gun (at present
Tsu-shi), Mie Prefecture, Honshu, Japan, 20–
IV–2001, K. Kumada leg. (NSMT-Ar 6984); allotype: female from Inoue, Ureshino-cho, Ichishi-

gun, Mie Prefecture, 7–III–2002, K. Kumada leg.
(NSMT-Ar 6986); paratypes: one female, samedata as for the allotype (NSMT-Ar 6987) and one
male, same data as for the holotype (NSMT-Ar
6985).
Description (holotype and allotype). Measurements: Body length female 1.32 mm, male
1.25 mm; prosoma length female 0.60 mm, male
0.61 mm, width female 0.52 mm, male 0.55 mm;
opisthosoma length female 0.74 mm, male 0.65

New Spiders from Japan

163

Figs. 40–44. Allothymoites kumadai gen. et sp. nov. [40–41, female (allotype: NSMT–Ar 6986), 42–44, male(holotype: NSMT-Ar 6984)]. — 40, Epigynum, ventral view; 41, inner organ of female genitalia, dorsal view;42, palpal organ, ventral view; 43, same, retrolateral view; 44, embolic division, prolateral view. [Scales:
0.1 mm.]mm, width female 0.82 mm, male 0.67 mm;
lengths of legs [total length (femurpatella
tibiametatarsustarsus)]: female I 1.83 mm(0.630.200.420.330.25), II 1.65 mm (0.550.190.360.300.25), III 1.16 mm (0.38
0.160.210.210.20), IV 1.40 mm (0.440.19
0.290.230.25), male I 2.08 mm (0.720.20
0.500.370.29), II 1.84 mm (0.610.190.42
0.350.27), III 1.30 mm (0.430.150.26
0.240.22), IV 1.54 mm (0.490.170.320.29
0.27).
Prosoma: Carapace slightly longer than wide(length/width female 1.15, male 1.11), with long
hairs standing on tubercle, respectively (Figs. 32,
35–36). Median furrow absent, surface of carapace strongly sclerotized, with many folds forming reticulation. Eyes (Figs. 33, 37) very large
and almost same in size except for female PLE
little larger and the male ALE smaller than others, anterior eye row recurved, posterior one

straight in dorsal view, AME-AMEAME-ALE(6 : 1 in female and male), PME-PMEPMEPLE, median ocular area longer than wide(length/width female 1.14, male 1.13), wider infront than behind or as long as wide (anterior
width/posterior width female 1.00, male 1.09),
clypeus longer than AME-AME, remarkably tuberclate. Chelicerae (Fig. 38) with two large teethon the promargin of fang furrow, labium triangle
and fused with anterior margin of sternum, wider
than long (length/ width female 0.71 male 0.62),
maxillae twice as long as labium, anterior marginof labium sclerotized and sharpened, sternum
strongly sclerotized, with long hairs, slightly
longer than wide (length/width female and male1.05), conspicuously marginated and posteriorly
truncated (Fig. 39). Female palp furnished withclaw. Legs slender and hairy, femur I: pro- andretrolateral sides with line of long hairs each on
tubercle, patellae curved ventrad. Leg formula:

164

Hirotsugu Ono

I–II–IV–III.
Male palp (Figs. 42–44): Femur relatively
short, as long as total length of the other segments. Cymbium long, palpal organ compactly
set in distal part of cymbium, paracymbium without hook; tegular apophysis wide plate and large,
median apophysis spiniform, curved apicallywith sharp angle, conductor indistinct; embolus
spiniform and winding, without any projection atbase.
Opisthosoma (Figs. 32, 34–35) wider in front
than behind and forming an equilateral triangle,
slightly wider than long (length/width female
0.90, male 0.97), its dorsum smooth and wholly
covered with white, long hairs. Booklungs ineffective presumably, without opening. Anterior
spinnerets and posterior lateral spinnerets thickand conical, posterior median spinnerets small,colulus absent.
Female genitalia (Figs. 40–41): Genital fieldwider than long, with sclerotized plate, and internal structure not visible through plate. Genital
openings situated at anterior margin of large holeof epigynum, the posterior margin of hole expanded ventrad. Intromittent canal thick andshort, spermathecae globular and close to eachother, fertilization tube long and curved.
Coloration and markings (Figs. 32, 35): Female and male: carapace wholly dark reddish
brown, chelicerae, maxillae and labium yellowish
brown, sternum reddish brown, palps and legs
yellowish brown, tibiae, metatarsi and tarsi oflegs ventrally black. Opisthosoma dorsally blackor gray, with some white, thin lines and dots,
ventrally gray with some white patches, spinnerets yellowish white.
Variation. Body lengths of the paratypes: female 1.38 mm, male 1.17 mm. Prosoma of themale paratype is blackish brown.
Distribution. Japan (at present known only
from Mie Prefecture).
Etymology. The species is dedicated to thecollector of the type specimens.
Family Linyphiidae

Genus Walckenaeria Blackwall, 1833
Walckenaeria iriei sp. nov.
[Japanese name: Kyushu-tsuno-nukagumo](Figs. 45–50)

Diagnosis. The present new species is closely related to Walckenaeria keikoae Saito, 1988,
distributed in eastern and northern Honshu, but
can be separated from the latter by the shape ofmale palpal tibia (cf. Fig. 50 and Saito, 1988, p.
20, fig. 9). The presence of additional prolateralapophysis (Fig. 49) of this new species is characteristic. These two species have peculiar projection on their ocular area.
Material examined. Holotype: male from Isahaya-shi, along the bank of Honmyou-gawa River,
Nagasaki Prefecture, Kyushu, Japan, 8–VIII–1998,
leg. T. Irie (NSMT-Ar 6988); paratype: one male,same data as for the holotype (NSMT-Ar 6989).
Description (holotype). Measurements: Body
length 2.17 mm; prosoma length 1.12 mm, width
0.69 mm; opisthosoma length 1.05 mm, width
0.75 mm; lengths of legs [total length (femur
patellatibiametatarsustarsus)]: I 2.91 mm(0.840.250.750.620.45), II 2.66 mm (0.780.220.670.570.42), III 2.23 mm (0.66
0.220.530.490.33), IV 2.93 mm (0.850.22
0.770.680.41).
Prosoma (Figs. 45–46): Carapace longer thanwide (length/width 1.62), with peculiar projection with long hairs on eye field, superficial median furrow and some radial lines present. Eyes
compactly set around projection, almost same insize, ALE and PME slightly larger, anterior eye
row procurved in frontal view, posterior row
procurved in dorsal view, all eyes close to eachother, clypeus almost as long as width of eye
field. Chelicerae (Fig. 47) with four teeth on promargin of fang furrow and three teeth on retromargin, distinct stridulating files present, labium
fused with anterior margin of sternum, wider
than long (length/width 0.56), maxillae long, distally with hair tuft, sternum longer than wide(length/width 1.07), posteriorly truncated. Distinct spines not recognizable on tibiae of legs except for tibia III proximally with one weak spine

New Spiders from Japan165

Figs. 45–50. Walckenaeria iriei sp. nov., male (holotype: NSMT-Ar 6988). — 45, Pro- and opisthosomata (appendages omitted), dorsal view; 46, carapace and chelicera, lateral view; 47, chelicerae, ventral view; 48, palpal organ, retrolateral view; 49, tibial apophysis, prolateral view; 50, tibia of palp, dorsal view. [Scales: forFig. 45, 0.3 mm, for Figs. 46–50, 0.1 mm.]

166

Hirotsugu Ono(2–2–1–1 is typical for this genus), distance between coxae IV less than their diameter, claws oflegs with three or four long teeth, Tm I 0.49, leg
formula: IV–I–II–III.
Male palp (Figs. 48–50): Ratio of length offemur to that of patella 2 : 1, tibia as long as
patella, dorsally with three trichobothria, without
long spines, dorsal tibial apophysis large, very
wide at base and sharp at peak, additional apophysis present prolaterally, which is wide and depressed with tip branched four times (Fig. 49).
Cymbium short, paracymbium small and curved,
embolic division not very large, embolus filiform, winding, with small projection at base.
Opisthosoma: Ovate, longer than wide (length/
width 1.40), colulus present.
Coloration and markings (Fig. 45): Carapace
orange, ocular area blackish, chelicerae, maxillae, labium and sternum light orange, legs yellowish brown, opisthosoma dorsally light graywithout any marking, ventrally relatively lighter.
Variation. The single male paratype has
body length 2.25 mm.
Distribution. Japan (at present known only
from the type locality).
Etymology. This species is dedicated to thecollector of the type specimen.
Remark. Female unknown.

one of Micrargus. In the male palpal organ, tibiaof Kagurargus is more complicated than that ofMicrargus and has a strong spine dorsally, which
is absent in the latter. In the shape of male palpal
tibia the new genus relatively resembles thegenus Oedothorax Bertkau in Förster et Bertkau,
1883, but the tibia of the latter possesses a dentate apophysis instead of the spine as in the new
genus. Although the structure of female genitalia
of Kagurargus is closer to that of Oedothorax
than to Micrargus, general characters of Oedothorax are far apart from those of the other two
genera as Tm I larger than 0.50, Tm IV present
and the body size more than 2 mm. The presenceof very long and sclerotized conductor along thefiliform embolus and a strong spine on male palpal tibia are the key characters of this new genus.
Etymology. The generic name is formed by a
combination of the Japanese word ‘kagura’
meaning sacred music and dance derived fromthe name of the mountain in the type area and the
Greek ‘Argos,’ the name of a being in mythology
having a hundred eyes taken from the name of
the related genus Micrargus. The gender is masculine.
Kagurargus kikuyai sp. nov.
[Japanese name: Kagura-gomagumo](Figs. 51–61)

Genus Kagurargus nov.
[Japanese name: Kagura-gomagumo-zoku]

Type species. Kagurargus kikuyai sp. nov., bymonotypy.
Diagnosis. The new genus stands close to
the genus Micrargus Dahl, 1886, in some important characters for linyphiids, especially in Tm I
0.41–0.42 in ratio, the absence of Tm IV, tibial
spines 2–2–1–1 in order, and the small body size
less than 2 mm. However, Kagurargus differsfrom the latter by the shape of male carapace andthe genital morphology. The male head of the
new genus has no sulcus and pit behind each
PLE, but has a deep furrow and five pairs of peculiar setae behind ocular area. The eye arrangement of male is also different from the typical

Diagnosis. See the above generic diagnosis.
Material examined. Holotype: male and allotype: female from Mt. Kagura-yama, 300–400 m
alt., Aoyama, Saeki-shi, Oita Prefecture, Kyushu,
Japan, 8–XII–2006, leg. N. Kikuya (NSMT-Ar
6999–7000).
Description (holotype and allotype). Measurements: Body length female 1.33 mm, male 1.26mm; prosoma length female 0.65 mm, male 0.60mm, width female 0.48 mm, male 0.44 mm;
opisthosoma length female 0.77 mm, male 0.67mm, width female 0.50 mm, male 0.49 mm;
lengths of legs [total length (femurpatella
tibiametatarsustarsus)]: female I 1.48 mm(0.450.160.320.260.29), II 1.36 mm (0.400.150.300.240.27), III 1.16 mm (0.32

New Spiders from Japan

0.150.230.230.23), IV 1.59 mm (0.53
0.150.380.290.24), male I 1.37 mm (0.45
0.150.300.240.23), II 1.29 mm (0.380.15
0.300.230.23), III 1.06 mm (0.300.12
0.220.210.21), IV 1.35 mm (0.410.15
0.320.250.22).
Prosoma: Carapace longer than wide (length/
width female 1.35, male 1.36), with some radiallines, its surface forming minute reticulation,
median furrow absent, cephalic part sexuallydimorphic: male carapace with deep furrow behind eye field and five pairs of peculiar setae(Figs. 54–56). Eyes relatively small, ALEPLE
PMEAME (5 : 5 : 3 : 3) in female (Fig. 51),
ALEPLEPMEAME (5 : 5 : 4 : 3) in male,167

the anterior eye row recurved in female and male,the posterior one procurved (female) or straight(male) in dorsal view, AME-AMEAME-ALE,
PME-PMEPME-PLE (female 1 : 1, male 4 : 1),
median ocular area longer than wide in female,wider than long in male (length/width female
1.38, male 0.63), wider behind than in front (anterior width/posterior width female 0.77, male0.44), clypeus almost as long as width of oculararea. Chelicerae (Figs. 52, 57) with six teeth on
the promargin of fang furrow, five teeth on retromargin, labium fused with anterior margin ofsternum, wider than long (length/width female
0.57 male 0.62), maxillae twice as long as labium, its anterior margin sclerotized and sharp-

Figs. 51–57. Kagurargus kikuyai gen. et sp. nov. [51–52, female (allotype: NSMT-Ar 6999), 53–57, male (holotype: NSMT-Ar 7000)]. — 51, 56, Eyes, dorsal view; 52, 57, chelicera, ventral view; 53, pro- and opisthosomata (appendages omitted)], dorsal view; 54, carapace and chelicerae, frontal view; 55, same, lateral view.
[Scales: for Figs. 51–52, 56–57, 0.1 mm, for Figs. 53–55, 0.2 mm.]

168

Hirotsugu Ono

ened, sternum slightly longer than wide (length/
width female and male 1.05), female palp without claw. Legs relatively robust, tibial spines oflegs 2–2–1–1, distance between coxae IV wider
than their diameter, Tm I female 0.41, male 0.42,
Tm IV absent, leg formula: IV–I–II–III infemale, I–IV–II–III in male.
Male palp (Figs. 59–61): Femur much longer
than patella (5 : 3), tibia as long as patella, dorsally with spiniform apophysis and long spine (Fig.
61). Cymbium short, without notch, paracymbium simple and curved, with digitiform tip, embolus filiform, long and winding, with developed
conductor.

Opisthosoma: Longer than wide (length/width
female 1.54, male 1.37), its dorsum smooth withshort hairs.
Female genitalia (Fig. 58): Genital field muchwider than long, without scape, but square plate.
Intromittent canals short, long spermathecae visible through integument.
Coloration and markings (Fig. 53): Female andmale: carapace yellowish brown, chelicerae,
maxillae and labium light yellowish brown, sternum yellowish brown (female) or yellowish
white (male), palps and legs light yellowish
brown, without annulation. Opisthosoma white(female) or yellowish white (male) dorsally,

Figs. 58–61. Kagurargus kikuyai gen. et sp. nov. [58, female (allotype: NSMT-Ar 6999), 59–61, male (holotype:
NSMT-Ar 7000)]. — 58, Epigynum, ventral view; 59, palpal organ, ventral view; 60, palp, retrolateral view;
61, tibia of palp, dorsal view. [Scales: for Fig. 58, 0.05 mm, for Figs. 59–61, 0.1 mm.]

New Spiders from Japan

lighter ventrally.
Distribution. Japan (at present known only
from the type locality).
Etymology. The species is dedicated to thecollector of the type specimens.
Genus Meioneta Hull, 1920
Meioneta tenuipes sp. nov.
[Japanese name: Ashiboso-keshigumo](Figs. 62–66)

Diagnosis. Although Meioneta spiders are
not popular in Japan, more than 150 species ofthis genus have been described from the Northern Hemisphere. Of these, the new species seems
to be closely related to Meioneta falcata Li et
Zhu, 1995 described from Shennongjia Forests,
Hubei Province, China (Song, Zhu and Chen,
1999). Both the species have a similar construction of male palpal organ and female genitalia.
However, the new species can be distinguishedfrom the Chinese one by the details of lamella
and embolus of male palp and the condition of
intromittent canals of female genitalia (cf. Figs.
64–66 and Li and Zhu, 1995, p. 43, fig. 3 h, k) as
well as by the coloration of the opisthosoma.
Material examined. Holotype: male and allotype: female from Aso, 500–600 m alt., Taikicho, Watarai-gun, Mie Prefecture, Honshu,
Japan, 27–VII–2003, leg. K. Kumada (NSMT-Ar
6990–6991).
Description (holotype and allotype). Measurements: Body length female 1.95 mm, male 1.46mm; prosoma length female 0.78 mm, male 0.61mm, width female 0.60 mm, male 0.48 mm;
opisthosoma length female 1.08 mm, male 0.76mm, width female 0.75 mm, male 0.50 mm;
lengths of legs [total length (femurpatella
tibiametatarsustarsus)]: female I 3.92 mm(1.070.221.050.980.60), II 3.35 mm (0.960.200.830.830.53), III 2.52 mm (0.75
0.190.590.600.39), IV 3.27 mm (0.940.19
0.850.810.48), male I 3.45 mm (0.920.19
0.890.890.56), II 2.88 mm (0.780.19
0.730.710.47), III 2.11 mm (0.620.150.47
0.510.36), IV 2.69 mm (0.780.150.68

169

0.660.42).
Prosoma: Carapace simple, longer than wide(length/width female 1.30, male 1.27 in ratio),with sparse hairs, a triangle patch at the middle
and radial lines, median furrow indistinct. No
projection on male carapace. Eyes relatively
large and almost same in size except for the ALE
and PME larger than the others, the anterior eye
row recurved, the posterior one straight in dorsal
view, AME-AMEAME-ALE, PME-PME
PME-PLE (5 : 4 in diameter) in female and male,
median ocular area almost square (length/width
female 0.87, male 1.00), wider behind than infront (anterior width/posterior width female 0.70,
male 0.78), clypeus longer than AME-AME.
Chelicerae with five teeth on the promargin offang furrow, three teeth on retromargin (Fig. 63),
labium fused with anterior margin of sternum,
wider than long (length/width female 0.53 male0.61), its anterior margin incrassated, maxillae
twice as long as labium, swollen, its anterior
margin sclerotized and sharpened, sternum
slightly longer than wide in female and male(length/width 1.11). Legs very slender, tibial
spines of legs 2–2–2–2 in order, distance between coxae IV less than their diameter, Tm I female 0.15, male 0.18, Tm IV absent, leg formula:
I–II–IV–III.
Male palp (Fig. 66): Tibia longer than patella,dorsally with two trichobothria, but without distinct apophysis and long setae. Cymbium with alarge notch retrolaterally, paracymbium thick andstrongly sclerotized, with digitiform tip, embolus
short and spiniform, curved apically.
Opisthosoma: Longer than wide (length/width
female 1.44, male 1.52), its dorsum smooth withshort hairs.
Female genitalia (Figs. 64–65): Genital fieldmuch wider than long, with indistinct scape.
Winding intromittent canals well visible throughintegument, spermathecae reniform.
Coloration and markings (Fig. 62): Female andmale: carapace wholly blackish brown, chelicerae, maxillae, labium and palps dark brown, sternum blackish brown, legs light yellowish white,their femora, patellae and tibiae with black lines

170

Hirotsugu Ono

Figs. 62–66. Meioneta tenuipes sp. nov. [62–65, female (allotype: NSMT-Ar 6991), 66, male (holotype: NSMTAr 6990)]. — 62, Pro- and opisthosomata (appendages omitted), dorsal view; 63, chelicerae, ventral view;
64, epigynum, ventral view; 65, female genitalia, posterior view; 66, palpal organ, retrolateral view. [Scales:
for Fig. 62, 0.5 mm, for Figs. 63–66, 0.1 mm.]pro- and retrolaterally. Opisthosoma milky whitedorsally, blackish distally, light gray ventrally,
spinnerets darker.
Distribution. Japan (at present known only
from the type locality).
Etymology. The specific epithet of this new
spider is formed by a combination of the Latin
tenuis (thin, delicate) and pes (foot, leg), and isderived from the condition of its legs.
Family Anapidae
Subfamily Mysmeninae
Genus Mysmenella Brignoli, 1980
Mysmenella ogatai sp. nov.
[Japanese name: Yamato-kotsubugumo](Figs. 67–75)

Diagnosis. This new species seems to beclosely related to Mysmenella gongi Yin, Peng et
Bao, 2004, described from Daoxian County,
Hunan Province, China, but can be distinguishable from the latter by the shape of male palp andthe structure of female genitalia. In Mysmenella
ogatai the distal part of embolus has a larger seculiform process and the female genitalia ismuch wider (cf. Figs. 70, 75, and Yin et al.,2004, p. 81, figs. 4, 8). Mysmenella jobi (Kraus,1967) widely distributed from Europe to Japandiffers from these two species by the absence ofthe round accessory organ at the spermatheca
and the longer scape of epigynum as well as details of male palpal organ.
Material examined. Holotype: male, allotype: female from Ikegane-cho, about 200 m alt.,
Okazaki-shi, Aichi Prefecture, Honshu, Japan,

New Spiders from Japan

29–VI–1995, K. Ogata leg. (NSMT-Ar 7376–
7377); paratypes: two females and two males,same data as for the holotype (NSMT-Ar 7378–
7381); three males (non type), from Itayama-cho,Handa-shi, Aichi Prefecture, Japan, 13–V–1993,
leg. K. Ogata (NSMT-Ar 7382).
Description (holotype and allotype). Measurements: Body length female 1.39 mm, male 1.14mm; prosoma length female 0.54 mm, male 0.52mm, width female 0.46 mm, male 0.50 mm, heightfemale 0.30, male 0.55; opisthosoma length
female 0.90 mm, male 0.67 mm, width female
0.76, male 0.54 mm; lengths of legs [total length(femurpatellatibiametatarsustarsus)]: female I 1.67 mm (0.530.210.350.290.29),

171

II 1.50 mm (0.450.200.320.250.28), III
1.09 mm (0.340.150.200.180.22), IV 1.43
mm (0.460.170.310.230.26), male I 1.80
mm (0.570.210.370.300.35), II 1.48 mm(0.450.200.310.240.28), III 1.07 mm (0.350.140.180.170.23), IV 1.34 mm (0.42
0.160.290.230.24).
Prosoma: Carapace slightly longer than wide(length/width female 1.17, male 1.04), very highin male (height /width female 0.65, male 1.10),
highest at the ocular area, with several setae at
middle, median furrow present, cervical bald
spots absent, the surface of carapace normal.
Head not modified, both eye rows recurved in
dorsal view, ALEPMEPLEAME in female

Figs. 67–70. Mysmenella ogatai sp. nov., female (allotype: NSMT-Ar 7377). — 67, Pro- and opisthosomata (appendages omitted), dorsal view; 68, same, lateral view; 69, epigynum, ventral view; 70, inner organ of
female genitalia, dorsal view. [Scales: for Figs. 67–68, 0.2 mm, for Figs. 69–70, 0.1 mm.]

172

Hirotsugu Ono(6 : 7 : 7 : 6 in diameter), AMEALEPME
PLE in male (10 : 8 : 7 : 6 in diameter), AMEAMEAME-ALE (2 : 1) in female and male,
PME-PMEPME-PLE in female and male, ALE
and PLE close to each other, median ocular area
almost square in female, as long as wide andwider in front than behind in male (anterior
width/posterior width 1.16), clypeus long, extremely in male, much longer than AME-AME.
Chelicerae with two teeth on the promargin offang furrow, labium wider than long (length/

width female and male 0.50), furnished with twopairs of setae, anterior margin sclerotized andsharpened, sternum covered with hairs, slightly
longer than wide (length/width female 1.05, male1.03), female palp present (Fig. 68).
Legs: Femora without spines, patellae I–IV
apically with long, dorsal spine in female, 1–0–1
spines in male, tibiae I–IV dorsally with longspine at middle, tibia I of male distally with twostrong, lateral spines, metatarsus I of male with astrong spine curved (Fig. 71), tarsus almost as

Figs. 71–75. Mysmenella ogatai sp. nov., male (holotype: NSMT-Ar 7376). — 71, Pro- and opisthosomata, dorsal view; 72, same, lateral view; 73, palp, retrolateral view; 74, palpal organ, ectal view; 75, embolic division,
dorsal view. [Scales: for Figs. 71–74, 0.2 mm, for Figs. 75, 0.1 mm.]

New Spiders from Japan

long as wide in all legs, teeth on claws of the legs
indistinct. Leg formula: I–II–IV–III.
Male palp (Figs. 73–75): Femur simple, without apophysis, much longer than patella; patella
not modified, without apophysis, tibia short, withsome long setae. Cymbium proximally simple
and small, distally with sclerotized plate (paracymbium) furnished with small tooth at middle(Fig. 74), palpal organ large and expanded fromcymbium, the embolic division bare, embolus filiform and winding, with securiform process large(Fig. 75).
Opisthosoma: Globular, longer than wide(length/width female 1.18, male 1.24), slightly
projected behind and the posterior part forming
oval plate with transverse folds. Anterior spinneret and posterior-lateral spinneret conical, posterior-median one small, colulus present withpair of hairs.
Female genitalia (Figs. 69–70): Genital fieldwider than long with small scape. Genital openings situated at the scape, internal organs not visible through integument. Intromittent canals thin
and winding, spermathecae reniform with an accessory process round.
Coloration and markings (Figs. 67–68, 71–72):
Female and male: carapace yellowish brown,with some radial markings in black, around both
AMEs black, chelicerae, maxillae, labium andpalps yellowish brown, sternum yellowish white,legs yellowish brown with dark annulations.
Opisthosoma dorsally grayish beige or gray, withmore than three pairs of white spots, posteriorly
with a pair of longitudinal white markings andfive transverse black lines, ventrally gray.
Variation. Body length of paratypes: females
1.27–1.53 mm, males 1.11–1.20 mm. A female
has larger white spots of opisthosoma, the markings of opisthosoma are not distinct in two males.
Distribution. Japan (at present known only
from Aichi Prefecture).
Etymology. The species is dedicated to thecollector of the type specimens.

173

Acknowledgements
The present author wishes to express his sincere thanks to Dr. Narayoshi Kikuya, Oita, Mr.
Teruo Irie, Kumamoto, Mr. Eiichi Shinkai,Tokyo, Mr. Hiroshi Saito, Yamanashi, Mr. Kiyoto
Ogata, Aichi, Mr. Kenichi Kumada, Mie, Mr.
Shingo Hatsushiba, Tokyo, and Dr. Shuhei Nomura, Tokyo, for their kind advice and for offering interesting material. This study was supported in part by the Grant-in-aid No. 16540431 for
Scientific Research by the Ministry of Education,
Science, Sports and Culture, Japan.
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