a -lf'ii-~~~~-~-!~--f
Transactions and Proceedingsof thePalaeontological Society of Japan
New Series

No. 56

Palaeontological Society of japan
Dec. 20th, 1964

CONTENTS
TRANSACTIONS
Page

477.

Bryozoa of Akiyoshi. Part 2.

Lower Carboniferous Bryozoa from the

Uzura Quarry .......................................... Sumio SAKAGA!III 295
478.

Some Solitary Corals from off Aomori Prefecture, japan ........ Kei !\IoRI 309

479.

A Rudistid from the Cretaceous Deposits of Ryoseki, Kochi Prefecture,japan ...................................................... Yasuji SAITO 317

480.

Yabeiceras (Cretaceous Ammonites) from Futaba, Northeast japan ..... .
. . . . Tatsuro MATSUMOTO, lkuwo OBATA, Shiro MAEDA and Teruo SA To 322

481.

On some Lower and Middle Triassic Ammonoids from japan .. Yuji BANDO 332

PROCEEDINGS ................................................................ 3,16
Systematic Index.................................................. •............ 3,17

Index of Families, Genera and Species ......................................... . 349

President: Teiichi KOBAYASHI
Councillors:

Kiyoshi ASANO (Editor of "Fossils"), Riuji ENDO, Haruyoshi FUJIMOTO.

Shoshiro HANZAWA, Tetsuro HANAI (Secretary), Wataru HASIIIMOTO (Treasurer).
Kotora IlATAI, lchiro HAYASAKA. Hiroshi OZAKI (Planning), Teiichi KOBAYASHI,
Tatsuro l\1ATSUMOTO (Editor of Special Publications). Masao l\IINATO, Tokio
Sill KAMA (1\fembership), Fuyuji T AKAI (Editor), Ryuzo TORIYAMA
Assistant Secretary:

Takeo ICHIKAWA

All Communications relating to this journal should be addressed to the
PALAEONTOLOGICAL SOCIETY OF JAPAN
Geological Institute, Faculty of Science, University of Tokyo, japan

Trans. Proc. Palaeont. Soc. Japan. K. S .. 1'-io. 56. 295-308. pis. 44. 45. Dec 20, 1964

477.

BRYOZOA OF AKIYOSHI. PART 2. LOWER CARBONIFEROUS
BRYOZOA FROM THE UZURA QUARRY
SCi'vliO SAKAGA\1!
Department of Geology, Hokkaido Gakugci University-F:J\sQ)l\i!U, -t;-0) 2, ? _;.( 71*ti~i!EF5C1=i~~ll\!R: '7 ;.( 7t~1:i~tJ. G:!*tf!l- ::'-:.:rilfZ-B!flQ)f.ll!lHt.:.r:i 8 .hlS 13 fill (6 ~fifi. -1 ~:JE;;Fk§till. 1 ll!t~till· 2 mfl.:ttm) :a-~~~na~ Lt.:, .:.
Q)JW£1!,1i!IJ~Hti:t :1 ill! 1- Jv-'\" 7- ;$1 :/$1!!1.JQ)Jii]~f~ff.!il.llf.!:: ~ft!t.d'tt{l.:ltl:a--1·; L. <:I.' 7.:>tJ'., ? 7: =j;H;~ Feneslel/a Mil:~f,(Jt.fl~t'L.fLi!. li!:I-:.-'CQ)rnliilif!:Q):Jjj(;l: 7.:> ~ ,!:;Q:Jtjjf~~J'LQ,
lli 1: ill cK
Introduction and Acknowledgements

was supported by a grant from the Scientific Research funds of the i\linistryof Education of the japanese Government.
All of the specimens described in thepresent paper are preserved in the collection of the Department of Geology.
Hokkaido Gakugci University. Hakodate.

Since the discovery of the Mil/erellazone in the Akiyoshi limestone by TORIYAl\IA. the fauna from the Uzura Quarry
located about 1 km east of Yobara on
the Ofuku plateau has been studied by
MUI<ATA (fusulinids, undescribed, 1961).
YA:-.:AGmA (brachiopods. 1962) and YAMAGI\\'A and 0TA (corals, 1963).
The
writer found some bryozoans in the limestone blocks collecled from the Uzura
Quarry and there is the scope of thepresent article.
The writer is indebted to Drs. juichi
YANAGIDA of the Kyushu University
and Masamichi 0TA of the Akiyoshi-Dai
Science Museum for their kind guidance
to the Uzura Quarry. and to Professor
Kotora 1-lATAI of the Tohoku University
for his kindness in reading this manuscript. The writer also wishes to express his thanks to Drs. G. G. ASTROVA
and N. A. SHJSHOVA of the Paleontological Institute. Academy of Science.
U.S. S. R. for their kind advices on the
Rhabdomesonid Bryozoa. This study

Remarks on the Bryozoan Fauna

The bryozoans from the Uzurn Quarry
were found in association with abundantspecimens of brachiopods. whereas rarely
with corals and others.
The following 13 species of Bryozoa\Vhich are distributed among eight
genera were discriminated from the col-lection.
Fistulipora sp. indet. A
Fistulipora sp. indet. B
Pseudobatostomella cf. P. granulosa(NIKIFOROVA)
Stenopora pusilimoni/a SAKAGAMI,n. sp.
Leioclema uzuraensis S.\KAGA).Il.
n. sp.
Saffordotaxis yanagidae S:\KACAl\fl,* Received July 2,1. 1964; read Sept. 20.
at Hiroshima 1964.
2%

Snmio S.4K.c1GAMI

2fH1

n. sp.

Streblascopora antiqua SAKAGAMT,
n. sp.

Streb/otrypella astrovae SAKAGAMJ,
n. sp.

Septopora waberi NIKIFOROVA
Pe1111i retepora ojukuensis SAKAGAMI,n. sp.

Penniretepora cf. P. fragilis CROCK·
FORD

Pemriretepora sp. indet.
.4canthocladia ? sp. indet.
Saffordotaxis yanagidae SAKAGA\11, n.
'Sp. is very similar to 5. incrassata(ULRICH) which was reported from the
Keokuk group of the United States ofAmerica by ULRICH (1890) and the lower
Visean of U.S. S. R. by NEKHOROSHEV(1953) and TrmNA (1958) but is not identical as mentioned in the description.
.Streb/otrype/la astrovae SAKAGAi\11, n. sp.
resembles Hyphasmopora mukhini NIKI·FOIWVA which was reported from thelower Carboniferous of Turkestan. Penniretepora ofulwensis SAKAGA\H, n. sp.
may be related to P. cf. jlexicarinnata·(YOUl'G & You:-.:G) which NIKIFOROVA
recorded from the lower Carboniferous·of Turkestan. The other species des·cribed as new can be distinguished fromthe previously known forms. Septopora
waberi NIKIFOROVA described in thisarticle coincides with the Russian specimens which were described originallyfrom the lower Carboniferous to Turkestan and later from the Taruskian horizon ~Ctr 1 : upper Visean), about 300 km
south of Moscow by SIIISHOVA (1952).
Pseudobatos/ome/la cf. P. granulosa(NIKIFOROVA) and PenniretejJOra cf. P.
Jragilis CROCKFORD described in thiswork m:1y be identical with the original
species which were reported from thelower Carboniferous of Turkestan andthe upper Visean of the Riverleigh lime-stone of Queensland. respectively, but
specific comparisons in dclail were notpossible. The other four indeterminable
forms are compared with known forms
but are not named because of their stateof preservations or bad orientation of
the specimens.
NIKIFOROVA (1926, 1933) described andillustrated 67 species of Bryozoa among\Vhich 33 species belong to the Family
Fenestellidae from the lower Carboniferous of Turkestan. She (1933) mentioned that the Turkestan fauna, especially in the cryptostomatous Bryozoa.
is closely related to the lower Carboniferous fauna of North America. England,
and the Kuznetzk and Donetz basins in
U.S.S.R. Although the fenestellid bryo·zoan fauna could not be found from thel.' zura Quarry. a considerable number of
species of the present fauna are considered to be very intimate or to haveclose relation with the Turkestan species
whereas there arc only a few species
resembling those of Australia and theUnited States of America. The writeris in the opinion that the closer resemblance is to be expected between
the Uzura fauna and the Turkestan
fauna. should fenestellid bryozoans befound from the Uzura Quarry.
YANAGIDA (1962) described and illust·
rated 13 species of brachiopods and hestated that it constitutes a unique faunalassemblage allied to the upper Viseanbrachiopods of Europe and to those ofsouthern Asia and Australia.
The geological age of the bryozoan
fauna described in the present article
was determined to be the same as concluded from the brachiopods by Y Ar\AGI·
D.-\.

Systematic Descriptions
Genus Fistulitwra l\'i'CoY 1850

,.

477.

Bryozoa. of A.kiyoshi.

Fistulipora sp. indet. A
Plate 44, Figure 2.

Zoarium attached to foreign substance.
.-encrusting, composed of t\'.·o or threelayers. each layer varying in thickness
from 0.5 mm to 1.0 mm. Zooecial tube
straight from the proximal to distal
ends. small and circular in tangential
section. Diameter of zooecial tube about·0.16 mm to 0.19 mm. There are usually·6 zooecia in 2 mm in diagonal direction.
Lunarium may be developed but indistinct. Interspaces between zooecial tubes·0.10 mm to 0.16 mm and filled with welldeveloped and fine vesicular tissue whichis regularly arranged in longitudinalrows. Detailed structures of vesicular
tissue unobservable. In tangential section. vesicular tissue rounded polygonal:and regular in size. and two to threerows of vesicles in interspaces between
zooecial tubes. Diaphragms unobserv:able.
Remarks :-Although rather well oriented sections were obtained, detailed microscopic observation could not be done
because of deposition of secondary
materials. The comparison with thepreviously described species as well as
naming are reserved until better preserved specimens are found.
Reg. nos. 13015-A. 13017.

Fistulipora sp. indet. B
Plate 44. Figures 1, 3.

A longitudinal section of a single fragment. Zoarium encrusting. 1.6 mm in
the thickest part. Zooecial tube straight,
becoming gradually larger from inner
to outer in diameter, about 0.19 mm to·0.29 mm. Diaphragm lacking. Vesicular.tissue well developed through the inner

Pco·t 2297

to outer. Vesicules small, fish-scale likeand arranged irregularly. Interspaces
between tubes at zoarial surface concave.
Stereom not developed entirely.
Remark.~:- The present form seems toresemble ()'clotrype conferta PERWI and
GUTSCIIICK which was described fromthe Amsden formation (upper \lississippian). southwest l'>lontana, U.S. A. inthe tangential section. However, thesingle fragmentary longitudinal sectionat hand is insufficient for specific decision.
Reg. no. 13024.
Genus Pseudobatostomellal\IOROZOVA, 1960

P:;eudobatoslomel/a cf. P. granulosa
INIKIFOIWVA)
Plate 4-l. Figures 8. 10. 11.
Compare.1926. Baloslomella granulosa NIKIFOI~O\'A,
hr. Ceo[. Kom .. Tom XLV. 110. 3. pp. 188,
189, pl. 4, fig. 2; pl. 5. figs. 16-18.

A single thin section showing both
typical transverse section and a longi·
tudinal section. Zoarium a cylindricalstem, 1.3 mm in diameter measured ontransverse section.
Thin sections :-Zooecial tube circularor oval in tangential section, its shorter
diameter 0.160 mm to 0.208 mm. !vleso·pores present but rather rare, subangular
to circular, their diameter 0.064·mm to0.080 mm. Thickness of zooecial wall
0.0,18 mm to 0.0&1 rum. Acanthopore present, but its details unobservable. In
the longitudinal section. zooecial tube
curved gradually outward from imma·
ture region. Wall of immature region
very thin and becoming gradu:illy thickin mature region. l'vloniliform poorly
developed at inner part of mature regionSmnio SAKAGAMI

of wall. Diaphragms usually arrangedin irregular interval in zooecial tube. theirintervals ranging from 0.05 mm to 0.16
mm. about 2 to 4 in mature region of a
tube. but in some tubes, diaphragms
lacking entirely. Diaphragms in meso·
pores probably present but indistinct.
Remarks :-The present form coincides·with Pseudobatostomella granulosa which
NIKIFOROVA describEd under the genericname of Batostomella from the lower
Carboniferous of Turkestan in the gener·a! characters. The detailed comparison
between them. however. is not adequate
because there is only one thin sectionat hand and NIKIFOROVA's original illu·
strations of the thin sections were drawn
by hand.
Reg. no. 13011.
Genus Stenopora LO);SDALE, 18•11
Stenopora pusilimonila SAKAGA(I.II.
n. sp.
Plate 44. Figures 4. 5, 9.

Two longitudinal but partly tangential
sections. Zoarium a cylindrical stem .
.~Jeasurements (in mm) :-Tangential sections:- Zooecial tube
oval or rounded polygonal in mature
region. but angularly polygonal in im·mature region. Diameter of zooecium
0.194 mm in average. ranging from0.14<1 mm to 0.224 mm 130 measurementson two specimens). l\lcsopore rather
rare. subangular to circular. its average
diameter 0.083 mm, ranging from 0.048.
mm to 0.112 mm {10 measurements on
two specimens). Acanthopores rather
small, usually disposed at each point ofintersection of zooecial wall but occasionally disposed at edge of zooecial walland surrounded by concentric fibrous.
tissue. their inner diameter 0.006 mm.
Longitudinal sections :-Zooecial tube·
nearly parallel to growth trend in im·mature region. curved rapidly outward
at the boundary between immature andmature regions. and straight and perpendicular to the surface in mature
region. Moniliform poorly developed.
Zooecial wall very thin in immature
region and becoming thick in mature
region. Usually one diaphragm disposed
at proximal end of mature tube. but insome tubes no diaphragms observable.

Specimen

1:13018-A
2.1

Diameter of zoarium

0.160-0.224

Diameter of zooeciurn..

0. 0·18-0. 096

Diameter of rnesopore

I

13032

I

0. 141-0. 22<1;

1.6

0. 064-0. l\2

i

0.006

Diameter of acanthopore
Thickness of zooecial wall
No. of diaphragm:; in zooecial tube

II

0.004-0.008

0.028-0.068

0.020-0.060

0-1

0-1

4-6
No. of acanthopore:; per zooecial tube
4-6!....---------------------------- - - - - - - -Remarks:-There are some questions
as to whether the present form can be
included in the genus Stenopora because

it has a diaphragm which is usuallydeveloped at the proximal end of maturetube, and poorly developed moniliform.

477.

Bryozoa. of Akiyoshi..

The present form differs from the al·
ready known species of Ste1wpora and its
similar genera in the features described.above.
Reg. nos. 13018-A (holotype), 13032.

l'm·t 2usually between zooecial tuhes, theiraverage diameter 0.064 mm. ranging from
0.048 mm to 0.112 mm (9 measurementson specimen No. 13022). Acanthopores
rather small. usually disposed at eachpoint of intersection of zooecial wall.
but occasionally disposed at edge ofzooecial wall and projected to inner sideof zooecial tube. and surrounded by darkcolored concentric fibrous tissue. Their
inner diameter less than 0.006 mm.
Longitudinal section :-Zooecial tube
straight from immature to m.1ture regions. \Vall of immature region verythin and becoming gradually thick inmature region. Length of mature regionof tube 0.32 111111 to 0.6'1 mm rneasurefl on
specimen No. 13013. .Moniliform unob·
servable. Diaphragms arranged irregularly in zooecial tubes, their interval
about 0.05 mrn to 0.14 nun, 3 to ,1 inmature region of a tube. Diaphragms
in mesopores uncertain.

Genus Leioclema ULRICH, 1882
Leioclema? uzuraensis SAK:\GA;-.11, n. sp.
Plate 44, Figures 6, 7.

Zoarium lamellate, probably thin but
its thickness unknown.
Tangential section :-Zooecia circularor rounded polygonal in mature region,
but angularly quadrate, pentagonal orhexagonal in immature region. Diameterof zooecia 0. I90 mm in average. ranging
from 0.160 rnm to 0.221 mm (32 measure·ments on typical tangential section of
specimen No. 1:~022). ~lesopores com·monty present. one to two disposed
A!easurelllent8 (in 111111) : --Specimen!

Diameter of zoo ecial tube
Diameter of me $Opore----------

1302 2

1:~0~7-A

0.144-0.244

0. 160-0. 224

0. 2ll8 0. 2-10-

0. 048 0. 112

0. 06,1-0. 112

0. 00.

0.006

13013

I

- -

Diameter of aca nthopore

0.006' Thickness of zo oecial wall

I ~o. ~f zooecia per 2 mm

II

-

I'

-

----

0. 024-0. 032-

1\o. of diaphragms in zooecial tube

No. of _acanthopores per zooecial tube
!____

-----

-

3-4

..i

4-6

10

I_ _ _ __-i__[_-_'----~

Remarks:-The acanthopores disposed
usually at each point of intersectionof the zooecial wall, diaphragms irregu·
larly arranged in mature ref.{ ion of zoo·
ecial tubes and less developed moniliform may be the important charactersof the present species.
The present
form can be distinguished from the

?
-?--

I

.!::6- -1

-

-

--

- -

previously described species by thefeatures mentioned above.
Reg. nos. 13013, 13022, 13027-A (holo·type).
Genus Sajfordotaxis l3ASSI.ER. 1952
Sajfordotaxis yanagidae SAKAGAJI,JJ, n. sp.

300

Swnio S;lKJlGAMI

-M

0

"'0

~
~

0

....;

Plate 45.1 Figure:; I '1.

000"'..........

0

0

-

.....

"'
"'
0

~

.....
0

:::l

~

0

0

;:::!

a.

0

00

M

a.

0

-I
..,. 1

~

a.
0

0

0

"'.....

00
0

0I

0
<6

<C

00

-

I
I

I'-

0

I~ I

~

6

0
00
0

00

0

0'

M

6

J

a.
0

0
~

0

- "'

a.

M

~

0

'

....;

0

I

"'
"' ...,.

M

0

c<')

~

0

.0

ttl
~

::::
0

M

00
0

a.
0

Ol

0

"'0

~

g

0

0

'

0

6

0

tn

,::;

I

<')

I

0

..0

0

"'

"'

"" ""~

0

8M

.....

0

0

00
00

0

0

0

0

i

I {',,>.,
I

t-

I

~

0

0

-

----"'~
~

0
0

:::l

.....
.....

a.

'
a.
0

....;

~

0
---

0

0

..0

"'

0

' ,---..

,__, I ~I ... I ,---..
OJ I c.... I !:£.... I
I
c
~

~

-"'

Ill
...._,

-

~

E

v:

ro

::;;

....

~

~

c

v:

E

Q)

<::

·;:;

"'""

0-

~

Q)

(/)

I
I E::;;
u

Q)

0

0
0

'0

.....0

N

E

~

~

Q)

'

I

;:,:

'?
0O'l

~

~

I

L-,
"

0

~

0

0

....Q)

N

...

c:

!:£ I

-0

c:
0

E

I

u

!E

::;;

Q)

0

~

~

0

I E

'-

0

""'--

'--

0.

a:;

~
Q)

~

E

E

E

Q)

ttl

i5"' i5"' c

Q)

ttl

;:;
<lJ

c
c

N

- ------

I

-"'
~
~

c

'-OJ

~~
~ u

~~
~~

~.,

.; ttl.§c::l "'

I0

Zoarium a cylindrical stem. attached'on foreign substance such as pebblesand shells. Diameter of zoaria aboutl mm in average, ranging from 0.8 mm
to 1.1 mm (7 specimens).
Lo11gitudinal and fallgelltial sections:Zooecial tube arises from a central axis
at angle of about 30• to ,10• and curved
gradually outward with outer surface of
zoarium at an angle of about 70• to go•_Zooecial aperture regularly arranged
both in longitudinal and diagonal series~
6.5 to 8 zooecia in 2 mm longitudinal
distance. Zooecial tube in tangential
section near surface oval and its shorter·
diameter excluding zooecial wall ranging
from 0.080 mm to O.l\2 mm and longer
diameter 0.144 mm to 0.208 mm. One
kind of acanthopores (megacant hopores)
surrounding zooecial tube regularly arranged in one row but occasionally arranged irregularly and their outside diameter ranging from 0.04 mm to 0.06mm.
About 0.1 mm near surface of maturezone covered by dark and well laminatedlayer. ivlesopore. diaphragm and hemiseptum lacking .
Remarks :-The present form is mostsimilar to the type species of the genus:
Saffordotaxis incrassata which was described originally under the generic nameof Rhombopora from the Waverly groupof Ohio. United States of America by
ULRICH 11888\, and later it was recorded
by ULRICH (18901 from the Keokuk groupof Kentucky, United States of America.
by NEKHOROSIIEV ll953l as a confer form
from the lower Visean of Kazakhstan
and by TRIZl'A (19581 from c;·F (lower
Visean) of the Kuzunetz basin. The
present form, however, can be distinguished from S. incrassala by the smaller
diameter of the stem (0.8-1.1 mrn insteadof 1.2-1.8 mm), larger number of zooecial~~·~"'I I .

Bryozoa of Akiyoshi.

aperture in longitudinal series (6.5-8 mm
instead of 5) and fewer megacanthopores
which surround zooecial tube.
The specific name is dedicated to Dr.
Juichi YANAGIDA of the Kyushu Uni·
versity who studied the brachiopod
fauna of the Uzura Quarry.
Reg. nos. 13006, 13010, 13014, 13016,
13021. 13028 (holotype). 13031.

Part 2

301

.I

M

M

0
M

......

......
0

_;

""

0

-

"'......

Genus Streb/ascopora BASSLER, 1952

Strehlascopora antiqua SAKAGAl\11.
n. sp.
Plate 45. Figures 5. 6. 13.

Zoarium a cylindrical stem. Diameterof zoarium about 0.9 mm in average,
ranging from 0.8 mm to 1.2 mm (6 specimens). Diameter of central bundle 0.21
mm in average, ranging from 0.16 mm
to 0.32 mm (6 specimens). Ratio of zo·
aria! diameter to central bundle about
4.5: 1 in average.
Longitudinal and tangential seclions:Zooecia1 tube arises from central bundleat an angle of 20" to 25,, straight inimmature region and curved gradually
outward in mature region. Number oftubes in central bundle 3 to 4 counted
on 3 longitudinal sections. In tangential
section of mature region. zooecial tube
oval and its shorter diameter excluding
zooecial wall 0.079 mm in average, rang·
ing from 0.072 mm to 0.092 mm (12measurements on one specimen) and its
longer diameter excluding zooecial wall
0.147 mm in average, ranging from 0.128
mm to 0.176 mm (12 measurements on
one specimen). Zooecial apertures regu·
Jarly arranged and about 6 per 2 rnm oflongitudinal series. Mesopores of twoor three rows with 2 to 3 in each row
longitudinally. Total number of meso·
pores disposed between zooecial tubes in

80

-

M

8M"'

-

..,.

......

"'

I

M

~')

0
'
~

"'
0
"'

- I I

I

I

c
'""'

3
2;

J.

~

'-"

- i

c.;

~·

..s........

ci
c

c

....c
E

E

c

<.>

E

302

Snmio S.4.101GAJ11

one series varying from 5 to 8. Diameterof mesopore 0.016 mm to 0.021 mm.
Superior hemiseptum poorly developed
at posterior end of mature region. andinferior hemiseptum also poorly developed at opposite side of inner part ofsuperior hemiseptum. Diaphragm occasionally located at lower part of immature zone of tube.
Remarks: - Streb[a!;cojwra seems tooccure more commonly in the Permianand Carboniferous rocks of Japan thanin those of foreign countries. About 19
species including the present new species
may belong to the genus. and of them
7 species (3 are from the Carboniferous,and 4 are from the Permian) were known
from japan. The present form is moresimilar to S. delicatula which SAKAGAMI
described from several localities of thejapanese Permian than the Carboniferous
species. However. the present form canbe distinguished from S. delicatula by
the simpler centrnl bundle.
Reg. nos. 13001. 13002 (holotype). 13008,
13012-C, 13018-B. 13033.

Strehlolrype/la as/rovae SAKAGA!\11,
n. sp.
Plate 15. Figures 7. 8.

Three sections which may be fromone colony. Zoarium c~•lindrical and its
diameter 1.1 mm in average. no branchingobserved.
Thin sections :-Zooecial apertures may
be regularly arranged in longitudinal
series. Zool:cial tube arises from central
axis and elliptical in longitudinal section near surface. with distinct peristomc
and its shorter and longer diameter
0.064 mm to 0.080 mm and 0.144 mm to0.160 mm. respectively. Number of zooecia in 2 mm of longitudinal row 6 to
6.5 near surface. Interspace between
zooecial tubes near surface in longitudinal series usually 0.11 mm to 0.23 mm.
Wall of immature region thin. but becoming rapidly thick at inner part ofmature region. Width of immature andmature regions 0.48 mm and 0.32 mm,
respectively.
1\lesopores arranged inlongitudinal series but not so regularand the inner diameter variable, ranging
from 0.004 mm to 0.020 mm. Diaphragmand hemiseptum unobservable.

Genus Streblotrypellu NtKlFOROVA. 19·18:\leasurements (in 111111) : ------Specimen

I13026-A

l3026 B

l.2

l.O

0.06·1-0. 080

0.080

-~-

Diameter of zoarium------------

Diameter of zooecium (shoncr)

---

- :1

0. lH-0. 160

Diameter of zooecium (longer)
No. of zooecia per 2 mm of longitudinal row '

-

~

I

I

6-6.5

~-o~oo.t::D.020j-- _0·_0_1_6 _ _ 1===========-!

Inside diameter of mesopore
Thickness of mature zone

I

Interspace between zooecial tubes

___

.. -~

--

0.11-0. 23

?

1

0. 32

-----1-------

477.

B-ryozoa 4 Akiyoshi.

Ne111arks :- The present species resembles llyphasmopora muk/zini NIKIFOROVA (1933) \'ihich was described fromthe lower Carboniferous of Turkestan in
the measurements except for the numberof zooecia in 2 mm of longitudinal row(6-6.S instead of •1.5 of the Russianform). Later. JI mukhini was describedunder the generic name of Nikifonwellaby NEKHOROSIIEV (in NIKIFOROVA's
paper in 19·18, personal communicationfrom G. G. ASTIWVA) and was also illustrated in the text-figure by SHULGA·NESTERE:\KO et a[. (1960). The present
species may be referred to the genus
Streblotrype//a by the zoarium having
central axis and the arrangement of
meso pores.
The specific name is dedicated to Dr.
G. G. AsTROVA, a Russian Paleozoic
Bryozologist who kindly helped me during my study.
Reg. nos. 13026-A (holotype). 13026-8,13026-C.

I'ctrt 2

Genus Septopora PROUT, 1859

Septopora zcaberi NIKIFORO\'.-\
Plate 45. Figure 14.

1926. Se/Jiopora waberi:\IKIFOROVA,/l;tt.

Geol Kom .. Vol. ·15. No. 3. pp. 183.
18•1. pl. ·1. fig. 9.
1952. Septopora tmberi SIIISHO\'A. Trud)'
Paleont. Inst. ;1./V., SSSR .. pp. 170. 171.
pl. 5, fig. 1, text figs. 13. 14.

A single fragmentary tangential section. Zoarium composed of straightbranches with dissepiments which usually bear 2 or 3 zooecia. Branches 0.10 mm
to 0.45 mm wide and probably about 10
in 10 mm horizontal. Width of dissepiment 0.21 mm to 0.32 mm. Fcncstrules
elongated rectangular with rounded
corners, their width and length about
0.40 mm and 1.12 mm tone measurement),
respectively. and probably about 7 to 8
in 10 mm vertical length. Zooecial tube
elongated pentagnonal at lower levc:l of
branch, kidney shaped at middle andTable :-Showing the measurements of Scptopora lt·aberi for comparisonof the present form with the l~ussian forms.
ljThe present form

Specimens-----1,\STRO\' \'s

1

' ·
----

· ·

13-1·1

11

7-8

ll

11

15

18

17-18

0. ·100-0. ·1•18

0. 35-0. •10

0. 112-0. 128

0. 12-0. 14'.Vidth of dissepiment0.240-0.320

0.20 0.25

Width of fenestruleabt. 0. 500

0.30-0. :>0

l No. of fenestrule in 10 mm \'erticalI

No. of zooecia in 5 111111
Width of branch

I,___
Diameter of zooecium
.

1

--

' _ _ __ _-----0. 70-0. 75

Length of fenestrulc
Number of zooecia t>cr fene:;trule3

3

No. of zooecia in dissepiment

2-3

2-3

Distance between zooecia

form 'i~~10

No. of branches in 10 mm horizontaiii

1

N!Kll-'oRO\~'\·:iform

0. U.l-0. 160

-I
I
-I
I

I

I

-I

0.25-0.30
0. 13
0.25-0.30
0.50 0.63
0.60-0.6'>
32-3

0.13-0.15

I

I

I

304

Sumio SA!\AG.{Mlcircular at upper levels. its diameterReg. no. 13004.

0.112 mm to 0.128 mm; about 15 in 5 mm
length of one row. usually 3 apertures

Genus Penniretepora D'ORBIGl\Y, 1849

per fenestrule. Occasionally, very large
tubercles disposed in longitudinal interspace between zooccial tubes. Carina
well developed, nearly straight. Nodes
may be present but indistinct. Stereom
covering the reverse side, developed
rather fine striations along the length ofbranch.
Meshwork formula :-abt. 10/7-8//15.
Remarks :-Septopora u·aberi was described originally from the lower Carboniferous of Turkestan by NJKIFOIWVA(1926) and later recorded from the Taruskian horizon (C," : upper Visean) of
the Nerutsch River. about 300 km southof Moscow by SHISHOVA (19521. The
present form coincides \Vith the Russian
forms especially in the internal structure
but the meshwork formulae are slightly
different. The formula of the present
form is. however, inaccurate in measurements because of there being only one
ill-preserved specimen at hand.

Penniretepora ofukuensis SAKAGA~II.
n. sp.
Plate -15. Figure 10

Two tangential sections. Zoarium consists of broad, straight main branch,about 0.8 mm in width. Lateral branchlets 0.32 mm to 0.38 mm in width, extending alternately at an angle of 65'
to 700 and intervals of 0.32 mm to 0.48
mm. with 6.5 to 7 branchlets in 5 mm.
Zooecial tubes arranged in alternating
longitudinal series. triangularwithrounded corners at middle level of tan·gential section, about 13 to 14 in 5 mm
length of one row, and spaced regularly
in pairs per interval between lateralbranchlets. Stereom covering the reverse
side, developed rather coarse fibrous
striations along the length of branch.
Interspaces between striations 0.01 mm
to 0.05 mm.

A1easurements (in mm) :13007
Specimen
Width of main _b_r_a_n_cl_l_ _ _ _ _ _ _ _ Il _ _ _t_l.-8(-1Width of lateral branchlet---1I________

abt. 0. 80

0. 320-0. 3&1

1:

Angles of later-al-b-ra_n_c_h_le_t_t_o_m_a_i_n_b_r_a_n(-.h-~Intervals between lateral branchlets

13023

65'-70<indL;tinct
abt. 70'I

0.320 0.480

indistinct

J

-·

i

3.5-7

6.5-7

I

Diameter of zooeciu m near surface

abt. 0. 100

0.120

I

No. of zooecia in one row of 5 mm

14

13 1-1

No. of zooecia per lateral branch

22

No. of branches it 5 mm

Remarlls :-In the meshwork (numberof lateral branchlets in 5 mmjnumber ofzooecia in one row I /number of zooccia

I--I

per lateral branch lets). the present form
is nearest toP. cf. _flexicarinnata (You:-:G& YOUNG) in NIKIFOROV:\'s paper (1933),

477.

Dryozon of Akiyoshi.

namely the meshwork formula of thepresent form is 6.5-7/13-U//2 and
NIKIFor<OVA 's specimen is 6-6.5/14//2.
However, the present form can be dis·tinguished from P. cf. _tlexicarinnata by
the wider main branch and lateralbranchlets (about two times of the latter
form, respectively) and narrower inter·vals between the lateral branches.
Reg. nos. 13007 (holotype), 13023.

Penniretepora cf. P. fragilis CROCKFORD
Plate 45. Figure 9.
Compare:1947 Penniretepora fragilis CROCKFORD.
Proc. Li1111. Soc. N. S. TV.. Vol/. lxxii.
Pt. /-2. p. 21. pl. 3. fig. 3, text-fig. 19.

Part 2:105

A single tangential section. Zoarium
consists of rather slender. straight mainbranch. about 0.4 mm in width. Lateralbranchlet about 0.24 mm to 0.29 mm inwidth. extending alternately at angle of
70' and at intervals of 0.64 mm to 0.80
mm. with 5 branchlets in 5 mm. Zooecial
tubes arranged in alternating longitudi·
nal series. somewhat elongated triangularat lower and middle levels of tangential
section, about 14 in 5 mm length of one
row, and spaced regularly in pairs perinterval between lateral branchlets. Diameter of zooecial tube near surface about
0.10 mm. Stereom covering the reverse
side. composed of very fine fibrous tissue
and two conspicuous striations observed
along the length of branch.

Table :-Showing the measurementa of the pre3ent form
and P. jragilis CrwcKFOJ~D for compari.;on.

S[>ccimcn

1_ _ _ _ i[_·_r_he_·_P_r_es_e_n_J_f_o_r_m_[ _~_P_._f_r_ag_,_·t_is~~i_\_v_i_d_tl_l_o_r_n__l_a_in~b-r_a_n_cl_l~-------~~~~
1
Width of lateral branchletI

0. 2-10-0. 288

Angles of lateral hranchlet to main branch

i,

70°

<15• -75°

0.640-0.800

0. 71-l. 68

5

-1.25

0. 10

0. 10

14

1,1

3

2-5

Intervals between lateral branchlets
No. of branches in 5 mm

II
II

Diameter of zooecia near surface

0. 37

0. 28-0.50-

0, 25-0. ,10

---

l

---

I

I No. of zooecia in one row of 5 mm

I

..

!
!

Xo. of zooccia per lateral branch

Remarks :-All measurements of thepresent form are covered by those of
Penniretepora f ragili s which CROCKFORD(194i) described from the Riverleighlimestone. the age of which has been
determined from the coral fauna to be
upper Visean or probably slightly
younger. of Queensland, Australia. However, specific comparison of the internal
structure is not possible because the-present material consists of only a single
tangential section.
Reg. no. 13010.

Penniretepora sp. indet.
Plate 45. Figure ll.

A single tangential section. Zoarium
consists of straight main branch. about

:~()()

O.lmm in width. Lateral branchlets
about 0.4.mm in width. extending alternately at an angle of 60' and at intervals of OA8mm to 0.56 mm. with about
5 branchlets in 5 mm. Zooecial tubes
arranged in 2 rows of slightly alternating intersection. elongated kidney-likeshaped at lov,·er and middle levels oftangential section. About 10 zooecia in
5 mm length of one row, and probably
spaced in pairs per interval between
lateral branchlets. Diameter of zooecial
tube near surface 0.11 mm. Stereom
covering the branch composed of very
dark. rather coarse fibrous tissue. andconspicuous striations and granules notobserved.
Remarks :-Because of the badly oriented specimens and insufficient material
at hand, the species was left unnamed.
Reg. no. 13012-A.
Genus Acau/hodadia K1~c. 18,19:lea nthocladia? sp. in del.
Plate 45. Figure 1~.

Text-tigure. Diagramatic schemeof the branching system of Acalllhoclcu/ia? sp. indet.
A. B, C: Angles of zigzag mainbranch. about 1:~oo to 150'.
a, b. c: Angles of lateral branchlets to main branch. about 90° to 100•.

Four fragmentary tangential sections.
Zoarium consists of zigzag main branch

Explanation of Plate ,111, 3. Fistulipora sp. indet. B
1. oblique tangential section. x20, Reg. no. 13015-A, 3, longitudinal section. x20, Reg.
no. 13017.
Fig.
2. Fistulipora sp. indet. A
Longitudinal section, x20, Reg. no. 13024.
Figs. ·1, 5. 9. Stenopm·a pusilimonila SAKAGA~II. n. sp.
5, longitudinal but partly tangential section (holotype), x20, Reg. no. 13018-A. -1. enlarged mature region of Fig. 5, 9, enlarged part of Fig. 5, showing the zooecial tube intangential section, x60, respectively.
Figs. 6, 7. Leioclema uzuraensis SAKAGA~II. :1. sp.
6. typical tangential section (paratype). x60. Reg. no. 13022. 7, oblique longitulinal :;ection (holotype), x20, Reg. no. 13027-A.
Figs. 8, 10. 11. Pseudoba/ostomella cf. P. granulosa (NIKIFORO\'A)
10. longitudinal and transverse section (holotype). x20, Reg. no. 13011, 8, enlarged partof Fig. 10, showing the mode of zooecial tube in the transverse part, 11, enlarged partof Fig. 10, showing the mode of zooecial tude in the longitudinal part, x60, respectively.

Fig3.

SAKAGAMI:

Bryozoa of Akiyoshi

Plate 44

11

SI\1\AGAMI photo477.

Eryozoct of Akiyoshi.

l'm-t 2307

Measurements (ill mm) : 13009

Specimen

13025

13029

---i-----------------------

abt. 1.00\Vidth of zoarium

abt. 1. 00

LOO--Xo. of row of zooecia
Diameter of zooecia near surface

-·

4

5

0.160-0.176

0. 114 -0. 160

4

0. 160-0. 176-~o.

of zooecia in 5mm longitudinal

11-12

n

12-13
--

Diameter of spicule?
-(about 130·-150•) and about 1 mm wide.
Width of lateral branchlets about aswide as in main branch, extending alternately at angle of about S0'-100'. Its
diagramatic scheme is shown in the textfigure. Zooecial tube arranged in alternating longitudinal series, usually 4 rows
on both main branch and lateral branchlet. rhomboidal at lower level. becoming
oval to circular at middle and upper
levels of tangential section. about 11 to
13 zooecia in 5 mm length of row.
Stereom composed of less developedcoarse fibrous striations and upper one
rather light. coarse granules arranged
in regular spaces. Inner diameter of
granules about 0.006 mm and interspace
between them about 0.02 mm.
Remarks :-So far as the present thinsections are observed, the present form
may be included in the genus Acanthocladia. However, there is some doubt
as to the generic identification because
the thin sections at hand are fragmentary and the growth form of the zoarium
seems to be similar to that of the genus!lnastomopora (=Neteporidra).
Reg. nos. 13009-A, 13009- U, 13025,
13029.
Heferences
B... ssLER. R. S. (1953): Bryozoa. in 1\-looRE's"Treatise on lnYertebrate Paleontologr ··.
Part G. Geol. Soc. America and Kansas

0.06

I

0.06-0.08

Unit•. Press. p. GI-G~53.
CROCKFORD, J. (1947): Bryozoa from theLower Carboniferous of New South \Valesand (Jucen:.land. Proc. Linn. Soc. N. S. W ..
Vol. 72. pt. 1-2. p. 1-48. pl. 1-6.
Mt·R.\TA. M. (1961):
On the Geological
Structure of the Akiyoshi Plateau. Inst.
Geo/. l'alaeo1tt. Tohoku Unit· .. :Vo. 53. p.
1-46.
KEKHOIW!-'IIEV. V. P. (1953):
Niznekamennougolnye l'\Lhanki Kazakhstana. Akad.
Nauk SSSR .. Trudy VSEGEI. 235p .. 25pl.
- - (1956): Niznekamennougolnye Mshanki
Altay a i Sibili. Ibid .. Tom 13. 'H9p .. 57
pl.
NIKIFOROVA. A. I. (1926) : Lower Carboniferous Bryozoa of Turkestan. lzt•. Geo/.
Kom .. Tom. XLV. :Vo. 3. p. 175-192. pl.
4-6.
- - (1933) : The Carboniferous Deposits ofCentral Asia.
Contributions to the
Knowledge of the Lower Carboniferous
Bryozoa of the Turkestan. Trans. United
Geo/. Prospecting Sen•. USSR, Fasc. 207,
p. 1-77. pl. 1-12.
PERRY. T. G. and GuTsCHICK. R. C. (1959):
Bryozoans from the Amsden Formation,
Southwest Montana. four. Paleont., Vol.
33, No. 2, p. 313-322, pl. 46-38.
SAKAGAt\11, S. (1961):
japanese Permian
Bryozoa.
Palaeont. Soc. fczpan, Spec.
Pap .. No. 7. 58p .. 30 pl.
SHISHOV!I. N. A. (1952):
Podmoskovnie i
Dono-medvedichkie
Kamennougolnyemshanki roda Septopora. Akad. NaukSSSR. Trudy. Paleont. Inst. XL. p. 159-176.
pl. 1-5.
Slll'LGA-NESTERE:"Ko, M. I. et a/. (1960):

Sumio s~tKAGAMI

308

Otrjad Crypto:;tomata. Osnovi Paleont.
l\·l.;hanki i Brachiopodi. lzd· Vo AN. SSSR ..
p. 72 92
ToRIYA:\IA. R. (1958): Geology of Akiyo:,hi:Pan 3. Fu:;ulinids of Akiyo.;hi.
Mem.
Fac. Sci. Kyuslm Unit .. (U). Vol. 7. p.
1-264. pl. l-48.
TRIZ:-;A, V. B.
(1958) :Rannekamennougolnye l\hhanki Kuznechkoi Kotloviny.
Trudy Vi\'JGRJ. 122. 435 p .. 64 pl.
YA:\IAGIWA. N. and 0TA, M. (1963): Fauna:;
and Correlation of .. Uzura.. Quarry.

Akiyoshi, Southwest Japan. Part I. Corals.
Bull. Akiyos!Ji-Dai Sci. Mus. No. 2, p.
87-93, pl. 1-2.
Y,\:-;AGIDA.
J.
(1962):
Carboniferous
Brachiopods from Akiyoshi. Southwest
Japan. Part 1. Mem. Fac. Sci .. Kyuslw
Univ. (D). Vol. 12. No. 1. p. 87-127. pl.
14- 21.
ULRICII, E. 0. (1890): Palaeozoic Bryozoa.
Gcol. Surv. Ill .. Vol. 8, p. 285-688, pl.
29-78.

Explanation of Plate -!5
Saffordotaxis yanagidae S.\KAGA:\11, n. sp.
4. longitudinal but partly tangential section (holotype). x20, Reg. no. 13028, l, 3. showing
the arrangements of acanthopores, x60. Reg. nos. 13014, 13028, respectively, 2. showing
the zooecial tube and wall in longitudinal section. x60. Reg. no. 13010-A .
.Figs. 5. 6, 13. Streblascopora antiqua SAKAGAMI, n. sp.
5, typical longitudinal section (holotype). x20, Reg. no. 13002. 6. enlarged part of Fig. 5,
showing the mode of wall in mature region. x60. 13, transverse section, x20, Reg. no.
13012-C.
Figs. 7. 8. Streblotrypella astrovae SAKAGA:\11. n. sp.
7, oblique section (holotype), x20. Reg. no 13026-A. 8. enlarged part of Fig. 7, showing
the arrangements of zooecial tubes and mesopores, x60.
Fig.
9. Penniretepora cf. P. fragilis CRoCI,FORD
Tangential section, x20. Reg. no. 13010-B.
Fig. 10. Penniretepora ofukuensis SAKAGA:\11, n. :;p.
Tangential section (holotype). x20. Reg. no. 13007.
Fig. 11. Penniretepora sp. indet.
Tangential section, x20. Reg. no. 13012-A.
Fig. 12. A.canthocladia? sp. indet.
Tangential section. x20, Reg. no. 13029.
~Fig.
14. Septopora u·aberi NIEIFOROL\.
Tangential section, x20. Reg. no. 13004
Figs.

1-·L

SAKAGAMI:

Bryozoa of Akiyoshi

SAT<AGAM I photo

Plate 45

Trans. Proc. Palacont. Soc. japan. ~. S.. ~o. 56. pp. 309-316. pl. 46. Dec. 20. 1961

-l78.

SO:\IE SOLITARY CORALS FR0:\·1 OFF

AOMORI PREFECTURE. JAPANKEI ~lOR!
Institute of Geology and Paleontology, Tohoku University·------~

Introduction
The present work treats the Recent
simple corals dredged from the conti·
nental shelf of the Pacific coast of
Aomori Prefecture. Japan. The species
here treated are all well known in Japan,
and their dis! ributions are also wide.
Concerning 1he Recent simple corals
from the continental shelf bordering
Japanese Islands; Honshu. Shikoku andKyusbu, many works have been pub·lished. Especially YABE and EGUCHI(1942) described the materials dredgedby the surveying ship "Soyo-maru .. of
the Imperial Fisheries Institute. Tokyo
during 1922-1930. Their monographic
work is an important one.
The study of the Recent simple corals
is important for the interpretation of
the fossil faunal composition, paleoecolo·gy and other features. Taking this
oppotunity the writer treats the systema·tics of the simple corals dredged fromthe continental shelf, off Shimokita
Peninsula, Aomori Prefecture as funda·
mental for further study.
The materials upon which this study
is based were collected by the membersof the Tohoku Regional Fisheries Re·
search Laboratory, Fisheries Agency of-~Received Julr 27, 196·1: read june 7. l<J6•lat Kyoto.

japan during their surveys on the fish·
ing ground in the summer of 1954. Unfortunately, whether the specimens werealive or dead when they were collected,
is not known. but it is thought that the
majority lived in the area where they
were dredged. On the other hand, neither holotype nor topotype specimens arc
accessible for direct comparison.
Concerning the four species and one
subspecies here reported an attempt ismade to compare them wilh the materi·als reported from other regions by Y ABE
and EGUCHI (1932. 1936. 1937. 19-H, 1942)and others.
Acknowledgements
The writer takes this opportunity to
express his deep gratitude to Professor
Kotora HATAI of the Institute of Geology
and Paleontology. Tohoku University for
his kind suggestions and critical readingof the manuscript. Acknowledgement
is due to Professor 1\lotoki EGUCHI of
the Department of 1\lining Engineering,
Faculty of Technology, Tohoku Uni·
versity for his kind guidance and helpful
criticism on the hexacorals. Deep appreciation is due to Professor Ikuzo
HAMAl of Faculty of Fisheries, Hokkaido Universit)' for his donation of theimportant materials and valuable sugges-

T\ei MORI

310
tions.

Hetel·ocyatlws sp.

He is indebted to !vir. 1\imiji

KU;\!AGAI

for his photographic works.

Studied Materials andCollected Localities

The distribution of the stations andthe data for the coral-bearing dredgings
arc given in Fig. l and Table 1, respectively.

The stations where dredge operations
w€re undertaken amount to 14, among
which the hexa·corals here treated occurcd at five stations of which the depth
ranges from 80 to 160 meters. Fromthese stations, four species and one subspecies distributed among three genera
were discriminated: they are:

0
Q)

00

c
0

..,a.
0

Flabellum transZJersale 1-lOSELEY
Flabellum transz:ersale conicun1
Y ABE and EGUCHI
Deltocyathus orienta/is DUNCAN
Heterocyathus japonicus (VERRILL)

Fig. 1. Location of dredgings on thecontinental shelf of the Pacific coast. off
Cape Shiriya. Aomori Prefecture.

Table l. Station data for the coral-bearing dredging
StationNo.

Date in
1954

I Depthin ~---Surface
metersten:p.

-------;.1

BottomSolitary corah;temp.

l====i====t====---=====·1===~=======-1

1.

Augu:;t 25j

80

21.60

15.07

Flabellum transl·ersa/eF. transpersale conicum
Deltocyatlms orienta/is
Heterocyat!ms japonicus
H. sp.

I

1--------t--------·---------ll---------!--------i----------------------l

1----1_0_.__ _._A._u_g_u_st_2:>_- 1--16_0___ ____
?0_._79_._ __ 13. -~6 _1
1
1
August
25
70
20.70
11.
16. 0()---~I August 25

90

20.60

_l_)_el_t_oc_y_a_t_h_us_o_r_ie_n_ta_l_is__ _1

1-l. 72

Flabellum transrersa/eFlabellum trans1·ersale
Deltocyatltus or·ientalis

1--------1--------1----------1

13.

August 23

140

21.80

9.40

Deltocyat!ms ol"ientalis

1---------r---------f,----------1---------·I-----------------------------"--14.
August 23
90
21.80
13.65
Deltocyat!ms orienta/isi'--------'---------- -·--------'----------'---------------'

Distribution of Species
and Bionomic Note

All the species reported in this article
arc well known from the seas surrounding japan. Some of them are widespreadand have been recorded from the Riukiu

Islands. Australia and East Indies. etc.
They have also been reported as fossil
from the Pliocene and Pleistocene rocksof japan" The distributions of eachspecies are given in the systematics.
The habitat of simple corals is controlled by the ecologic factors: temper-

478.

;~ l lSolitary Co·~·als o:tr Aomori Pnfectun

ature. depth, substratum, etc.
However. they are Jess restricted environmentally than
the reef-building hermatypiccorals.
As indicated in Table 1, thesimple corals were dredged
from an area where the surface-water temperature ranges
from 20.60 to 2l.80"C and thebottom-water temperature from
9.40 to 16.o6·C. On the other
hand, the distribution of thesame species from the seassurrounding japan were inareas having the following
bottom - watertemperature
range CYABE and EGUCHI. 1942).

ABCDEFABCDEFABCDEFSTATION I

STATION lO

STATION ll

J..ll

ABCDEF~ABCDEFABCDEFSTATION 12

STATION 13

STATION 14

Flabellum transrersale:
L4-l7.8·c
F. transt•ersale conicum:7.0-17.0"C
Deltocya/hus orienta/is: :!.3-23.3"CHe/erocya/hus japonicus: l2.Q .. 16.0"C

As mentioned, the range of temperature is fairly large. but the maximum
de...-elopment of these species in japan
is found in areas where the temperature
is between 12"C and 19"C.
The depth from which the ahermatypic corals were dredged is in the rangeof 80-160 meters, off Shimokita Peninsula. Aomori Prefecture.
In the seasaround japan, on the other hand, thedepth range of each species is as follows.
Flabellum transrersale: 17-344 meters
F. /ransuersa/e coni cum: 80-219 meters
Deltocyathus orienta/is: 59-3411 meters
Heterocyathus japonicus: 55-188 meters
The most favorable bathyenvironmentof the four mentioned simple corals is
between 80-200 meters in japan, butabroad. the greatest known depth fromwhich the simple corals have been recorded is 4914 meters for Dc/tocyathus

Fig. 2. Diagram showing the grain size ofthe sediments of each station. (!".>)
A: 0.05 (mm). B: 0.20-0.05. C: 0.50 0.20.
D: 1. 00-0. 50, E: 3. 00-1. 00. F: 3. 00.

orienta/is and 522 meters for Flabellum
transt·ersale.
The histograms (Fig. 21 represent thevarious types of sediment from where
the simple corals were dredged. Although the number of stations and coral
specimens are insufficient to permit ananalysis of grade preference, it is problematic whether the samples were deador alive at the time of collection, butseveral inference can be drawn; there
is a marked preference by Flabellum
transz·ersa/e and Deltocyathus orientalis
to fine to medium grained sand or shells
and for substrates where fine to medium
grained sand and shells are the dominantconstituents.
The silt and gravel-grade facies show
smaller distribution compared with thatof the sand and shells where Flabellum
trans~·ersale and Deltocyathus orienta/is
were found.
Concerning Flabellum
transz·ersale conicum and Helerocyatlws

Kei il!ORI"312

japonicus the data at hand are insufficient for drawing conclusions.
Among the illustrated three species
and one subspecies, Flabellum transversale, Deltocyathus orienta/is and
1Ieterocyat hus japonicus are well known
as fossils from the Pliocene and Pleisto-cene rocks in japan. The data presented
in the present article are important for
an interpretation of the paleoecology of
the Pliocene and Pleistocene fossils ofsolitary corals, but detail discussions are
not given in this work. because thetreated materials are few and restricted.
However, this is an important subject
for further study on the solitary corals
in japan.
Systematics
Family Flabellidae BOURNE, 1905

Genus Flabellum LESSON, 1831
Flabellum fransz;ersale !\loSELEY
Pl. 46. fig.;. 6. 7.

Flabellum transoersa/e 1\IosELEY, 1881. Sci.
Exped. H. i\1. S. Clwllenge,·. Rep., Zoo/.,
2. Vol. 7. p. 174. pl. 6, figs. 6-6a: YABE
and EGUCIII. 1932. Imp. Acad .. Proc.,Vol. 8. No.9, p. 4<13; YAIIE and EGUCI!I,
19-12, Tohoku Imp. Unir•. Sci. Rep .. 2ndSer. (Geol.). Vol. 22. No. 2. pp. 99-100.
pl. 6, fig. 11: pl. 7. figs. 1-6. 8-10, 12:
YAilE and EGt;CHI, 1942, Ibid. Vol. 22.
No.2. pp. 134-135, pl. 11, figs. 16-17.
Flabellum dens ALCOCK, 1902. Siboga Exped.
Monogr .. 16a. p. 32. pl. '1. fig.>. 30-30a.
Flabellum inconstans MAREI':lELI.ER. 1904,
Wiss. Ergeb. deutsch. Tiefsee-Exped .. Bd.
7, p. 277. pl. 17, fig. 11.
Flabellum llarmeri GARDI:-!ER, 1929. Brit.
Mus., Nat. llist. Rep. Zoo/., 5. No. 4, p.
122, pl. 1, figs. 19-20.

Dimensious {in mm) :Height of corallum
Longer diameter of calice
Shorter diameter of calice
Edge angle
Facial angle
Number of septa

1
20.0
20.0
10.0
41>
23•
96

Remarks:- The present species was
first recorded from Bass Straits, Australia by MoSELEY in 1881; it also occurs
in the seas around japan. Flabellum
dens ALCOCK (ALCOK. 1902) originallyrecorded from the East Indies, Flabellum
inconstans MARENZELLE}{ (MARENZELLER,
1904) which was first reported from
Australia and Flabellum harmeri GARDINER have been included into Flabellum-fl'{fllSVersafe MOSELEY by YABE and
EGUCHI (1942). This view is accepted-by the writer.
Among the specimens here treated,
three specimens have the septa of the-5th cycles complete and in regular2
28.6:~2. 0
11. 5
60•
21 c
120

3
22.0
29.0
10.0
62•
23•
96

-I
29.0
25(?)
13( ?)
-16•
26•
96

5
27(?)
27(?)
11(?)
53'
23•
100

hexameral plan. In this respect, they
resemble Flabellum transversale conicum
YABE and EGUCHI (1942), but can be
distinguished from the latter by having
more compressed corallum. They are
considered to be of forms intermediate
between the typical species and thesubspecies conicum.
The present specimens are more or lesssimilnr to Flabellum rubrum (Quoy andGAil\IARD). but the latter has morecharacteristic crescent-shaped basal scar.
Recently. Donald F. SQUIRES (1963) published a detailed study concerning Flabellum rubrum and in it he included
various species into the synonymy of it.

478.

Solita.ry Cora[.'! o.ff' Aomori J>n~(ectnre

But at present, the writer accepts theview of Y ABE and EGUCHI (1942) con·.cerning the relation between Flabellum
transversale and Flabellum rubrum.
Although the specimens measured
above are not excellently preserved andsomewhat different from the typical one,their features fall in the range of vari·ation of Flabellum tnlllsversale MOSELEY.
Occurrence :-St. 1. 80 m : St. 11. 70 m ;St. 12, 90 m.

Geographical dis I ribution: - This is-one of the widely distributed species.
Besides the Japanese seas, it has beenrecorded from the East Indies and theBass Strait, Australia. As fossil. thespecies has been recorded from the
Pliocene and Pleistocene sediments of
Chiba, Shizuoka and Kanagawa Prefectures. The chronological range of thespecies may be from Pliocene to Recent.

:~ 1:~CHI (1942) the most characteristic feature
distinguishing Flabellum transversale
conicum from Flabellum transversale isthat the septa are of the regular hexameral arrangement in Flabellum trans·
rerSltle conicum, whereas they are irregular in Flabellum I ransversale.
One of the specimens here mentioned
has 56 septa and is more or less similar to
Flabellum transuersale, but the generalshape of the corallum closely resembles
this subspecies. This may be a typetransitional between Flabellum transversale and transrersale conicum.
Occurrence :-St. 1, 80 m.
Geographical dislribulion :-This subspecies has been known from the seasaround Japan and the distribution isrelatively restricted compared with Flabellum transuersale.
Family Caryophylliidae GRAY, 1847

Flabellum transuersale conicum

YABE and EGUCHI

Genus lleterocyathus Milne
Eow A 1ms and liAil\m, 1848

Pl. 46. fig 3.

.Flabellum

I ransz•ersale conicum YARE and
EcucHt. 1942. Toltoku Imp. Unil' .. Sci.
Rep .. 2nd. Ser. (Ceo/.). Vol. 22. No. 2.
pp. 100-101. pl. 7. figs. 8. 10: YARE and
EcucHt. 1942, Ibid .. Vol. 22. No. 2. p.
135, pl. 11. fig. 18.

Ileterocyal hus japonicus (VERRILL)
Pl. 46. fig. 4.

Stepltanoseris japonica VERRILL, 1866. Essex
lnst .. Proc .. Vol. 6. pl. 47 (fide YABE
and EGt;CIII, 1942)

Dimensions (in mm):Height of corallum
Longer diameter of calice
Shorter diameter of calice
Edge angle
Facial angle
Number of :;epta

Subfamily Caryophylliinae GRAY, 1847

Stephanoseris lamellosa VERRILL, 1866. Ibid ..
11. 5
11. 5
5.5
60•
24•

56

9.2
7.0
5. 7

28'
48

Remarks :-Of two measured specimens
the smaller one agrees well with thetype of Flabellum transversale conicum,
but the other (larger one) is somewhat
different. According to Y ABE and EGU-

p. 51 (fide YABE and EGUCHI. 1942)

Steplwnoseris cartlzausi FELIX. 1913, Palaeontogr .. Vol. 60. p. 32. pl. 27. figs. 9-9h:
C\IAKIYA~IA, 1926. Ceo/. Soc. Tokyo. jour ..
Vol. 33. p. 5.
Hetcrocyatlms(Stephanoseris)
carthausi
FELIX. SAKAKt;R.-\. 1935, Ceo/. Soc. To·
kyo. jour .. Vol. 42. p. 189.
Heterocyatlms japonicus (VERRJt.L). YABE
and EGUCIII, 1942. Tohoku Imp. Univ ..
Sci. Rep., 2nd Ser .. Vol. 22. No. 2. p.
127, pl. 11. fig. 6.

Kei ll!ORI

314

Remarlls :-This species was first described by VERRILL as Stephanoseris
japonica from Kagoshima Bay, Kyushu,
Japan. The present species is more orless similar to Ileterocyatlzus aequicoslalus :tv!ilne EDWARDS and ]. HAIME in
general appearance, but differs in having
indistinct pali, the costae alternating insize, and the thinner costae are coloreddarker than the thicker one. Becauseof the abmre mentioned characters andby having perforated septa, the present
specimen is identical with Ileterocyatlzus
japonicus (VERRILL).
Occurrence :-St. 1, 80 m.
Geographical distribution:- This species has been reported from the seassurrounding Japan and as fossil it is
known from Chiba Prefecture, Taiwan
and Java. This species and Heterocvatlzus aequicostatus are the most widelydistributed species in the Genus f!eterocyatlzus in Japan.
Heterocyatlws sp.
PI 46. fig. 5.

Remarks:- The general outer shapeof the corallum of the present specimen
resembles I!eterocyathus japonicus (VER·
RILL), but it cannot be identified with it,
because the calice of the specimen exa·mined is ill preserved.
Occurrence :-St. 1, 80 m.
Genus Deltoryathus Milne EDWARDS
and HAI~IE. 18,18

Deltocyatlzus orienta/is Dm>CAN

Proc .. Vol. 8. No. 8, p. 388: YABE and
EcucHJ, 1932, Ibid .. Vol. 8, No.9, p. 442:
YABE and EGt;cur. 1937, Toltoku Imp.
Uni1•. Sci. Nep .. 2nd Ser. (Ceo/.), Vol.
19 . .1\ro. 1. pp. 131-135, pl. 20. figs. 1-10;
YALlE and EcL'CHI. 1942, Ibid .. Vol. 22.
No. 2. pp. 125-126.
Deltocyathus lens ALcocK. 1902. Siboga Exped .. 1Honogr .. J6a. pp. 19-20, pl. 2, figs.
16-16a; U~rBGROVE, 1925, Geological andPetrological and Palaeontological Results.
of Exploration carried from September
1917 till June 1919 in the Island of
Ceram by L. RuTTE:-1 and \V. Hou:,2nd. Ser .. Paleont .. No. 1, p. 4. pl. 1,.
figs. 8-10.

Remarks:- Fifty-eight specimens of
this species were examined. They arein varied stages of growth and vary insize; the largest specimen at hand is.
8.3 mm in diameter. 5.8 mm in heightand has 48 septa, and the smallest one
is 11.0 mm in diameter. 1.9 mm in heightand 36 septa.
This species is one of the well know·n
deep water corals from Japan. and among
the coral specimens here reported. thoseof the present species are most common.
Occurrence :-St. 1. 80 m; St. 10. 160·m; St. 12, 90 m: St. 13, 140m: St. 111.
90 m.
Geographical distribution:- This species is well known from the seas around.lnpan, the Philippines, East Indies andthe :Mediterranean Sea. Among the solitary corals recorded from Japan, this is.
one of the predominant species. As.
fossil, this species has been recorded
from the Pliocene and Pleistocene rocks.
of Chiba and Kanagawa Prefectures andof the Riukiu Islands.

Pl. 46. figs. 1, 2.
Dellocyatlms orienta/is Du:->cAJS, 1876. Zoo/.
Soc. Lo11don, Proc., p. 431, pl. 38, figs. 47; YAnE and EGL'CIIr, 1932, Imp. Acad..

References
ALcocK. A. (1902): Report on the deep-sea!vladreporaria of the Siboga Expedition_

-'178.
Siboga-l~X/){'(/itie.

Sohlar!} Corals o.tr .tlomo1·i FrefectnJ·eMon., 16a, 51 pp., 5 pis.
Notices of some deep·sea and liTtoral corals from the Atlantic
Ocean, Caribbean. Indian. New Zealand.
Persian Gulf and Japanese seas. Zoo/.
Soc. London. Pro<·.. ,'-lo. 29. pp. 428-442.
4 pis.
EGUCIJI, M. (l!l:~9): On the coral fauna of
the Toyama·Bar, Japan. Trans. Bioi. Soc.
Manclzoukou. Vol. 2. No. 3. pp. 41-47.
- - (1941): On two species of simple corals
from Kagoshima-ken. Kyushu. Geol. Soc.
japan, Jour., Vol. 48. No. 575, pp. 418420.
FELIX. ]. (1913): Die fossilen Anthozo;;naus der Umgegend \'On Trinil. Palaeonlogr .. Vol. 60, pp. 311-371. Taf. 24-27.
GARDT:-;ER. j. S. (1929): Coelenterata. pt. 4
l\tadreporaria. /Jril. Mus. Nat !fist .. Rep ..
Zoot .. !'o/. !i, No, 4. pp. 121-130.
HATAt. K. and IIAYASAK,;. S. (1955): Significance of the Pleistocene Marine faunafrom the .Jizodo Formation, Chiba Prefecture.
Tokyo 1\yoiku Daigaku. Sci.
Rep .. Sa. C. Vol ..J. No. 21i. !>P· 1-6.
MAKIYA~IA. J. (1!126):
On three simple
corals in Commensalism. Geo/. Soc. Tollyo, jour .. Vol . .'13. pp. 1-13.
1\IL-\RE:'-IZELJ.Io:Tl. E. von. (190·1): Steinkorallen.
Wiss. Ergeb. deulsch. Ticjsee-Exped. 18981890. 7. pp. 2ti]-31!l. pls. H-18.
MooRE. R. C. (1956): Treatise on Invertebrate Paleontology. Pt. F. Coelenterata.
MosELEY. II.!><. (1881):
Deep-sea .Madreporaria.
Exped. H . .\/. S. "Challenger"
Rep. Sci .. Zoo/.. 2. pp. 127-208. pi:-;. 1-16.
·QlioY. ].R.C. and GAT:.TARD, ]. P. (1833):.. Voyage de !'Astrolabe". Zoo/ .. Fol. 4.
p. 188.
SAKAKURA. K. (1935): Les Heterocyathus
fo:;siles de Ia Prefecture de Tiba. Japon.
Geo/. Sue. japan. jour .. Vol. 42. pp. 182191.
SQGTRES. D. F. (1963): Flabellum rubrum(Quoy and GAT MATH>) . .'\'. z. Dep. Sci.
Indus/r. Res. /Jull.. 15·1. pp. 1-43.
Tohoku Regional Fisheries Research Laboratory Hachinohe Branch (1956): Bottom
Fic;h. Annual l<cp. on Lhe Fish Resources
DUi'\CA:". P. :vi. (1876):

in Oct. 1953-Sept. 1954. Sec ..J.
R. F. (1921) Report on Pleisto·
cene and Pliocene Corals from Ceram.
Geological, P.:lrographical and Palaeontological Re:;ult.; of Explorations carried
from September 1917 till June 1919 in
the Island of Cerarn by L. RuTTE:-.1 andW. HoLz. 2nd Ser .. Palaeon/., No. 1. pp.
1-22.
V.\liGHA:-<. T. W. and WELLS. J. W. (1943):
Revision of the suborders, familie~. andgenera of the Scleractinia. Geol. Soc.
Amer., Spec. Paper. No . .J.J. pp. 1-363.
pis. 1-51. text.figs. 1-39.
YABE, H. and EGt;CIII. M. (1932): A studyof the recent deep-water coral fauna ofJapan. Imp ..·lead.. Proc .. Vol. 8. A'o. 8.
pp. 387-390.
- - and - - (1932) : Deep-water cor:tbfrom the Riukiu Lime:>tone of Kikai·jirna,
Riukiu Islands. Ibid., Vol. 8, No. 9. pp.
442-145- - and - - (1932) : A new species ofEndopachys. Endo/Hlchys jafJOnicus. froma younger Cenozoic dcpo~it of japan.
jap. jour. (;eo/. (;eof!r .. V o/. 10. Nos. 12. pp. 11-17.
- - and - - (1936) : Deep·watcr coral.>from off Owasi, J\Iie Prefecture. Jmp.
;lead., ?roc., Vol. 12. No. 6. pp. 1G7-168,
3 figs.
- - and - - (1937): Notes on /Jc/tocyathus
and Discolrochus from japan. Tohoku
Imp. Unit' .. Sci. Rep., 2nd Sa. (Geol.).
Vol. 19. 1\lo. 1. pp. 127-147.
- - and - - (1941) : Simple corals fromthe Sumagui formation. the Philippine
Islands. Imp. Acad. Tokyo. Proc., l'ol.
17. No. 6. pp. 210-215.
--and (1941): On some simple corals
from the )\eogene of Java. Ibid .. Vol.
17, No. 7, pp. 269-273.
- - and - - (19-12): Fo3sil and Recent
F'/abellum from .Iapan.
Tohoku Imp.
U11i1' .. Sci. Rep., 211d Ser., Vol. 22, No.
2. pp. 87-103. pis. S-8.
- - and - - (1942): Fossil and I~ecent
simple corals from Japan. Ibid., Vol. 22,
No. 2. pp. 105--178. pis. 9-12.

u~TnGROVE.

Kei ilfORI

316

Shiriya

.....

0?.

Explanation of Plate 16(All figures in X 2.3, a:

calicular vew, b: lateral view. c:edgewise view)Figs. 1-2: Deltocyathus orienta/is DL•:-;cAN. Loc. St 1 IGPS coli. cat. no. 85718.
Fig. 3: Fla!Je/lum tranSI'ersale conicum YAnr: and EGliCIII. Loc. St. I. IGPS coli. cat. no. 85717.
Fig. 4: Heterocyal/ws japonicus (VERRILL). Loc. St. 1. IGPS coli. cat. no. 85719.
Fig. 5: Heterocyatlms sp. Loc. St. I. IGPS coil. cat. no. 85B67.
Figs. 6-7: Flabellum transversale MosELEY. Loc. St. I, 12. IGPS coli. cat. no. 85716.

Plate 46
MORI:

Sirnp~e Cora~s

3c

lb

3a

2a

4b

KuMAGAIphoto

Trarn. Proc. Palaeont. Soc. Japan, N. S., No. 56. pp. 317-321. pl. 17. Dec. 20, 1964

47!1.

A RUDISTID FROM THE CRETACEOUS DEPOSI'fS
OF RYOSEKI. KOCHI PREFECTURE. JAPANYASUJ! SAITO
Institute of Geology and Paleontology. Tohoku University

i\'li~I!l.l:llfWQ)J'JIIli*C~tH?Q)

Rudistid:

l'iS~~B!tlll{h'if~f:Lmfdlf'iffl>Jo."'Q) l:ift;n·~~V.

L t-:.J•,(j,jq··-·_f';(H Monople11ra sp. i.:.--:>l ·-c;lctU_J-:.,

:tf $

m-

men.
Here the writer wishes to express hi&
sincere gratitude to Professor Kotora
IIATAI of the Institute of Geology andPaleontology, Faculty of Science, Tohoku
University for his kind guidance andadvice throughout this work. He also
thanks Dr. Kunihiro ISHIZAI\1 of thesame Institute for information with regard to the stratigraphy of the area.
Thanks are expressed to Mr. 1\imiji
KUC\IAGAI for taking the necessary photographs.

Introduction and Acknowledgements

Rt1distids which are an aberrant typeof sessile pelecypods are common andwidely distributed in late Mesozoic deposits of the world. They have various
shapes and generally are con11ned to thetropical and semi-tropical facies of theCretaceous. These highly specialized oraberrant bivalves have a short verticalrange and a wide distribution, therefore
they may render valuable aid in thecorrelal ion and determination of stratigraphic units in the Cretaceous. Rudistids. however. are extremely rare in
the Cretaceous rocks of the japaneseIslands and the species hitherto known
are only two, l)raecaprotina yaegashii(YEHAR/d and Toucasia carinata orientalis NAGAO. The writer describes one
rudistid which was collected by Dr.
Kunihiro ISIII7.AKI from the Cretaceous
deposits of Ryoseki. Kochi Prefecture,
Shikoku during his geological survey in1958.
Since no rudistids have been
known from the area and the genus
discussed in present work may be thefirst record from the Cretaceous of japan,
it is thought to be of no small importance to describe the present speci-

Geological and Paleontological Notes

According to PALMER (1928) the distribution of rudistids is represented witha band that extends almost around theearth and that with the exception of the
detour around northern Africa, roughly
parallels the present equator and lies in
the area of warm water. Although thejapanese Islands are geographically remote from the northern limit of distribution. there have been reported the occurrences of rudistids by Y ABE and YEHARA(1913), YEHARA (1920), YABE and NAGAO(1926) and NAGAO (1932, 1933).
The species already described are
Praecaprotina yaegashii (YEHARA) andToucasia carina/a orienta/is NAGAO.

Received August 6, 1964; read june 7.
196·1 at Kyoto.

317

1' a.suji SA.! TO

318

The former was 11rst discovered by YEfrom the Cretaceous deposits ofl\1 iyako. I wate
Prefecture, northeast
japan and regarded as belonging to
Plagioplyc/ws (YABE and YEHARA, 1913).
In a later publication with its detailed
description YEHARA (1920) assigned it to
Horiopleura and called it H. yaegashii.
Later, Y ABE and NAGAO (1926) proposeda new generic name and ascribed it totheir Praecaprolina n. gen. The latter
is from the thin lens of the Orbitolina
Limestone, exposed along the IshikariRiver, between Ponmoshiri and Shimanoshita, Province of Ishikari, Hokkaido
and its detailed description was given
by NAGAO (1932. 1933).
Concerning the classification of therudistids many workers have studied toarrange them into sections. FISCHER(1887) divided them into five families;
Chamidae, ~lonopleuridac, Caprinidae,
Hippuritidae, and Radiolitidae.
Thisclassification was modified by later
IIARA

workers and two families were added;
Diceratidae, and Caprotinidae. When
PAL:\IER made a study on the rudistidsof southern IVlexico in 1928. he summarized them into five groups in so far
as resemblance is evidence of relationship. They are Diceratidae-ChamidaeGroup,
Monopleuridae-Caprotinidae
Group. Caprinidae, Radiolitidae, andHippuritidae.
Recently. DECHASEAUX(1952) divided the rudistids except
Diceras into the type fixed by the left
valye and the type fixed by the rightvalve. The former is represented mainlyby the Diceratidae and includes Ilelerodiceras, Requinia, Toucasia, etc. The
latter is divided into five forms, which
are namely, the ancient type without
accessary cavities. the type with accessary cavities, the type with palliar canals,
I-lippuritidae. and l~adiolit idae.
The specimen treated in this article
was collected by Dr. Kunihiro lSHIZAKI
of the Institute of Geology and Paleont-N---Fault""'·~

Limeatone

Serpentine
Fig. I.

Index map showing the fossil locality (after lsJIIZt\1<1. !9GO)

479.

319

11 Rnclistid from Ryo..;;eki

ology, Faculty of Science. Tohoku Uni·
versity, from the Cretaceous deposits at
Okuminodani, west of Ryoseki, Kureta·
Cho. Nangoku City, Kochi Prefecture.
Shikoku (fig. 1).
According to lSH!ZAKI (1960) the SO·
called Chichibu terrain in the area. although mainly composed of Paleozoic
formations, also includes Nlesozoic sedi·ments mainly of the lower Cretaceous.
These Cretaceous deposits are classified
into three formations. namely, the Ryo·seki, the Nishinotani, and the Hibihara inascending order. The present specimen
was collected from the upper part of theKureta sandstone member of the Hibihara formation which is of !vliyakoan
age. From the same locality. he collected
such fossils as Trigonia sp .. Exogyra sp.,
Astarte sp .. Cerithium sp., and an Echi·
noid spine.
The specimen at hand can be easily
distinguished from the two genera ofrudistids already reported from \liyako
and lshikari in only superficial aspect.
So far as the external characters areconcerned. the present specimen resem·
bles the species of Ilippurites, Radiolites,
Durania, and Monopleura in having theoperculate upper valve and much deeper
lower valve ornamented with verticalribs. I-Iow·eyer. the present specimen
differs from Hippurites in the absenceof two deep siphonal grooves in addition
to the ligamenta! groove which are wellshown on the surface or in cross section.
The present specimen also differs from
Ilippuritcs, Radiolites. and Durania inthe ligament.
Although those three
genera are characterized by the ligament
being shifted to a subcentral position,and by the elongate teeth being modified
for vertical motion of the upper valve,on the contrary the specimen at handis considered to have a ligament situatedalong the hinge. The present specimen
has no accesssary cavities and as statedbelow. many of its features are in com·
mon with the genus Monopleura MAT·
HERO~. Although the specimen in poorly
preserved and the literature on Mono·
pleura not abundant, the present speci·
men may be considered to be new toscience.

Description
Family lvlonopleuridae FISCHER. 1887
Genus Monopleura 1\IATHERON, 18-!2,\/onopleura sp.
Pl. '17. figs. 1-7.

DescrijJiion :-Two valves very inequi·valve. and unfortunately broken away
at attachment of the lower valve. The
lower valve may have been conical inshape. being much deeper than the upper,
and in cross section from transversely
ovate to rounded subquadrate. Shell·
wall rather thick especially on the dorsal
side. Exterior of lower valve withlongitudinal ribs at regular intervals
approximately of 1.7;) mm apart, ornamented by fine vertical striae. besides
fine growth lines appear as rings They
are more or less distinct. Ligament
internal. ligamenta! groove not seen ex·
ternally but in cross section appearing
as a shallow concavity. Upper valve
ovate in outline. some\vhat convex or.
operculate with eccentric umbo situatedon the dorsal margin. Surface with
distinct concentric growth lines. Shellstructure obscure. Body cavity occupies
most of the interior. Verlical tooth andmuscle scars not visible. Hinge plate
known only by inference, but no elongate
teeth modified for vertical motion. No

320

Ya.suji SAITO

accessary cavities or no vertical canals.
Dimensions:- The present specimen isabout 65.0 mm long and with the dia·meter of 35.0 mm to 44.5 mm .. but was
originally much longer.
Nemarlzs :-The present material isreferred to the genus Monop!eura because of its general resemblance to thatgenus. though a well developed tooth ofthe lower valve or two teeth of theupper valve could not be observed. This
specimen belongs to the lVIonopleuriclaeCaprotinidae Group of PALl\IER (1928)
which is characterized by the generalcone or funnel shape of the lower
attached valYe, the presence of one
vertical groove (the ligamenta! groove)on the surface. and by the comparatively
small upper valve which may be flat or
somewhat convex. This group is usually
ornamented with vertical ribs and in·eludes the genera. Caprotina, Chaperina,
Polyconites, lloriop/eura, Monop/eura,etc. Monopleura, as pointed out by
DECIIASEAUX (1952). differs from theother genera in the :-~bsence of accessary
cavities.
Comparing this specimen with Mono·
pleura lri/oba/a (o'OHBIGNY) in the collec·
tion of the Institute of Geology andPaleontology. Tohoku University, it isfound that the former is distinguishable
from the latter by the vertical ribs of
the lower valve and by the lack of ribson the upper valve that radiate fromthe ligament. This specimen is similar
to .\lmwpleura salazari PAL~!ER from
Soyatl:-~n de Adentro, jalisco. Mexico inoutline of the shell. but differs from thelatter in the absence of deep horizontal
growth stages. Monop!eura imbricata
1\lATIIERO:\ which is another species re-

Explanation of Plate 47
Figs. 1-7: .Monopleura sp. (IGPS coli. cat. no. 85769) xI. Loc. Okuminodani. west of Ryoseki.
Kureta-Cho. Nangoku City. Kochi Prefecture.
1. Posterior ,·iew. a: Lower valve. b: Com·ex lenticular operculate upper vah·e.
2. Side view.
3. Surface of lower valve ornamented with vertical ribs.
4. Upper valve ornamented with concentric ribs.
5. Area of attachment of the lower valve.
6-7. Cross section of the specimen. cut slightly below upper valve.
(view of interior of the lower \'alve).
L: Ligamenta! groove.
G: Body cavity.
Figs. 8-9: Touetzsia carina/a orienta/is NAGAO. A topotypc specimen collected by \V. I!ASIII·~tOTo (IGPS coli. cat. no. 59592). X I
8. Lateral view.
9. View from bottom or of lower valve.
Figs. 10-12: Praecaprotina yaegaslzii (YEHARA).
10. Polished specimen of lower val\·e previously described by YABI' and NAGAO (1926)
from the Orbitolina Limestone of the lower course of the Sorachi-River. Sorachi-gun.
Hokkaido (IGPS coli. cal no. 22324). xl
11. Lateral view of a specimen collected by ~r. I·!ASIII~IOTO from Shimanoshita. Ashi·
betsu. Sorachi-gun. Hokkaido (IGPS coli. cat. no. 58586). x0.7
12. An upper valve collected by F. SAITO from Iliraiga, Miyako, Iwate Prefecture (IGPS
coli. cat. no. 35H2). x0.5

SAITO : Rudistid

KUMAGA I photo

Plate 47

479.

A Rudistid .frorn Ryoseki

sembling the present one superficiallyand with the lower valve ornamented
by vertical ribs. is different in having
radial ribs on the upper valve and a
few deep horizontal growth stages. The
present specimen lacks the radial ribs
usually found on the surface of theupper valve of the Genus :Honopleura.
As the cardinal teeth of the present
specimen remain unknown, further comparison is not possible.
Occurrence:- The upper part of theKureta sandstone member of the Hibihara Formation at Okuminodani, KuretaCho, Nangoku City. Kochi Prefecture(IGPS coli. cat. no. 85769).
References
CHUBB, L. J. (1956): Some Rarer Rudists
from Jamaica, B. W. I. Paleontogr. Amer..
Vol. 4. No. 26. pp. 1--24. pis. 1-5.
FISCHER. P. (1887): Manuel de Conchyliolo·
gie et de Paleontologic Conchyliologique.
pp. 1044-1068.
lsJJJZAKJ, K. (1960) : On the Geology and
Tectonics in the Area northeast of Kochi
City. Shikoku-On the Unconformable
Relationship between the Upper Paleozoic
and the Lower Cretaceous. jour. Ceo/.
Soc. japan. Vol. 66. pp. 553 565 (inHibihara
KuretaNishinotani
Okuminodani

13

J.t

~

.fL 01

illi

/

~~

tt-

~-?;/f't

321

Japanese).
Kl'JJ:'-1. 0. (1941) : Morphologisch-anatomische
Untersuchungen an Rudisten. Zentral·blatt f. Min. etc., Abt. B. l•ir. 12. S. 362-

371.
MATJJERON. P. (1842): Difference3 qui existent entre les Hippurites et les Radiolites.
Bull. Soc. Ceo/. Fr .. Vol. 13, p. 520.
NAGAO. T. (1932) : Occurrence of Toucasia
in the Lower Cretaceous of Japan. Proc.
Imp. llcad .. Vol. 8. No. 2. pp. 51l-5U.
- - (1933) : A new variety of Toucasia
carina/a (MATJJERO:->) from the Lower
Cretaceous of Japan. jour. Fac. Sci.,
Hokkaido Imp. Unit'., Ser. 4. Vol. 2. No.
2. pp. 16:{-167. pis. 21. 22.
PAL~tER, R. H. (1928):
The Rudistids ofSouthern Mexico. Occ. Pap. oj the California .4cad Sci. l.J. pp. 1-96. pl. 7.
DECHASEALiX. C. (1952): Lamellibranches in
PlVETEAU';; Traite de Paleontologic II,
pp. 323 .. 364
YABE. H. and NAGAO. T. (1926) : Praecaprotina. nov. gen. from the Lower Cretaceous of Japan. Sci. Rep. Tohoku Imp.
Ullit• .. 21ld Ser .. Vol. 9. pp. 21-24. pl. 7.
YAnE. H. and YEHARA, S. (1913) : The
Cretaceous Deposits of Miyako. Ibid.
Vol. 1. pp. 9-23.
YEHARA. S. (1920) : A Pachyodont Lamellibranch from the Cretaceous Deposits of
Miyako in Rikuchu. jour. Ceo/. Soc.
Tokyo. Vol. 27. pp. 39-43, pis. 11. 12.

Ponmoshiri
RyosekiShimanoshita

jlf-

il

~li

$4;

8{.

T1
/

r:

Trans. Proc. Palaeont. Soc. japan. N. S .. No. 56, pp. 322-331, pl. 48. Dec. 20, 196~

480.

LlBEICER:18 (CRETACEOUS Al\L\IONITES)FROM FUTABA. KORTHEAST JAPAN*
TATSURO ~IATSVivlOTO
Department of Geolog)'. Kyushu University

IKU\VO OBATA
Department of Palaeontology. National Science J\luseurnand

SHIRO :viAEDA and TERUO SATO
Geological Institute, Chiba University

Jttit

-v-"

~~ *K~~,m
1 -1:: 7"' (L'Iilli~c 7 / -r: :~·-f 1-) : l:fflliJ'lli*i'fii''rttlf Hfl;ll~H;:.J~-t 7.:>
JX~~ Hfl;ll)~'t':H'hH?~L-t.: Yabeiceras orientale ToKu:--:AGA and S111:-11Zli 11)1t;t!i(Ar.:i*l:t· ]RJJ(~~Q)*I.::~~f.cf.l: I?~~ Lt.: c ~t.c ~;(L o, ;c.:: 'L'lii'G!gJ1l!f.l• ~;;I? t.::.i\'[*a-A'I.t :> t~1JVc~t.:.. r<¥l!trf:.•Ni!.!:t.•J·~JQ)i/Jrta-A'I·n·t.:. :lfl:i!IlifJa=i(1)51.\f.:L-t.:llK!lft1if<rr~t.:.. {~llit./~l.:. ~ 1J2 W·1i!!:Q)fl::ti~l*!t~ l L .: t.:.l? ;.:.-?~ ·"C <*" L-~ ·rJijo~~f:l'flt.!~Hl't.c~ ··}'abeiceras Q)MU\i~f*i:.~::>l ··C:iill-:1"7.:>. Jlij~Q)i!J;f1f,!';;t, Forresteria i.::.~.:J.t.:;A\t;~;Joo, it.:. Eboroceras 13.\SSE. 1946 i:t. Y'abeiceras ToKt.:N1\li and SIII:>!IZU. 1926 Q)-/ / = J, ·c,!;, 7.>, F. t E. ~ 1t'l:;l~ Barroisicerali nae (.::.)\;(1 ~h "( l · ;,5, :it2~ L t.:l:l'f*lt.fLffl"*~;-:l;:.f*n'-to,
1965 ~fi.:.9;:~'11JH~i\:#:l;!:.fl:x.Q) 88 Jiltl.::.t.cl?it'!"t • .;'\l..t.'L':t.;~£t · L . .::Q)•J·~'i':it~l.:.
~~ m; · 'l' .'1'. m; 1o · !iii 1::!:1 11t1 ru; · fl.: iji til! 'A*

mJ:tf :t-t.

Introduction

ToKUI\AGA and Sm:-.nzu (1926) des(;ribed a number of Cretaceous fossils
from the Futaba Formation in the Joban
coalfield, about 200 km to the north·
northeast of Tokyo. This was an admirable contribution to our kno\vledgc inthat the Upper Cretaceous deposits wererecognized on the evidence of fossils
underneath the coal-bearing Tertiary of
the joban area and that some of thedescribed species were entirely new toscience. Yabeiceras. an interesting ammonite genus. was established on that
• Received August 20. 1964: read Sept.
20. 196-1.

occasion.
The original type specimens describedby ToKUNAGA and SIIIMIZU (1926), mostof which must have been preseryed at
the \Vaseda University, Tokyo, together
with other fossils from the Joban area,
are judged to have been burnt up by afire of World War II, 19H. except for a
few specimens w·hich have been for somereason deposited at the Tohoku Uni·
versity, Sendai. This judgement is basedon the replies given to the writers fromthe Faculty i\lembers of Waseda ljni·
versity, Tohoku University and the Uni·
versity of Tokyo and also Dr. H. Y ABE.
The holotype, by monotypy, of Vabei·
ceras orientale TOKUNAGA and SHIMIZU

:322

480.

Yabr~iceras(Cretaceous Ammonites)(1926. p. 201. pl. 22, fig. I: pl. 27, fig. lac), the type-species of Yabeiceras TOKU·NAGA and SHii\IIZU, 1926 (originally de·signated in p. 199), and the type-specimens of other two species, }·. koloi and
V. himuroi. of the same genus were
probably destroyed. This regrettablesituation has given to palaeontologists
some difficulty in the appropriate deter·
mination of the systematic position of
the genus as well as in the correct
identification of the species.
The two senior authors (T. ·M. andI. 0.), therefore. have endeavoured to get
specimens of J'aheiceras from the type·
locality. upper reaches of Sakurazawa
in Oriki, where Lower l'vlember of the
Futaba Formation is exposed. Although
bivalves occur there in abundance. ammonites are quite rare. Only a small.
probably immature specimen of Yabei·cera.'> orientale and a few heteromorpha
have been collected there.
In summer 1963, the third author (S. tv!.\
conducted a departmental excursion to
the Futaba area. After six days fieldwork by a group of thirteen students.
an adult ammonite which was unfamiliar
to them was finally acquired at the same
locality by the last author (T. S.). a
student. The specimen was brought tothe Department of Palaeontology. National Science Museum. Tokyo and shown
to the second author (I. 0.1. who told
him its significance. It was developed
from the rock matrix and cleaned in alaboratory of Chiba University and thendonated to Kyushu University for fur·ther palaeontological study.
This paper contains the description
and illustration of the t\VO specimens.
13efore going further the writers wish
to acknowledge the debt of gratitude toPrefessor Emeritus Hisakatsu YABE.
Professors Kotora HATAI, Sadajiro Tsu-

323

CHID A and Dr. l'vlasafumi MURAT A forgiying them information about whereabouts of TOKUNAGA and SHIJ\IIZU's
types. and to tvlr. C. \V. WRIGHT in
London for his critical reading of thetypescript. Dr. I taru I-lA YA~II took pho·
togr::~phs at Kyushu University and !vliss
Tomoko l'viJY AZAKI ::~ssisted in preparing
the manuscript.

Description}·abeiceras orientale TOKl!1\ AGA
and SHIMIZU
Pl. 48. Figs. la-c. 2a-e:
Text-figs. la-d. 2a-e, 3.

Yabeiceras orientale TOI<U:-<AGA andfour. Fac. Sci .. Imp. Uniz·. Tokyo. Sect. II. Vol. 1. Pt. 6, p. 201. pl.
22. fig. 7; pl. 27. fig. la-c.
1926. Yabeiceras oz·ientale. SHE\IIZl'. Proc.
Imp. Acad.. japan. Vol. 2, No. 10. p.

1926.

SHIMIZC,

547.

1)'pe :-The holotype is the specimen
described and figured by ToKt]NAGA andSHIMIZU (1926. indicated above), which\\·as almost certainly destroyed in 1944.
A neotype has to be officially proposed
and its validity should be acknowledged.
.Material:-The two specimens to be
described in this paper are GK. H5556,
an internal mould of an adult shell(CoiL S. l'viAEDA and T. SATO). and GK.
H5557, that of an immature shell (CoiL
I. OBATA). They are preserved in the
Type-Specimen Room, Departmet of
Geology, Kyushu University. Fukuoka.
The rock matrix is made up of calcareous sandstone. Therefore the fossils aresecondarily little deformed. On somepart an inner layer of the test is preserved. The innermost nuclear part anda half of the body-whorl of the first
specimen arc not preserved. The un-Tat.-;Hro M.4.TSUMOTO et al .

.324

illustrated side of the shell is dissolved
and hardly separable from the rockmatrix in which shells of Apiotrigonia,

Glycymeris and other bivalves are accumulated.

1\1easuremen/s (in millimeters):Diameter

Umbilicus

Height123.0(1)
GK. H5556
52.0(1).. [3/2 yo!. early].. [ 2 vol.
J
-to. 0(1)]c .
.. [ 3 vol.
29.6(1)
GK. II5557].. [ t vol.
20.4(1).. [ ! vol. ,. ]Holotype(T. & Sn.) 62

59.4(0.48)
23. 0(0. -1-!)

39.6
17. 3(0. 33)
15.5
12.5(0.31)
12. 0(0 . .JO)
10.6
8. 8(0. 43)
22(0. 35)

Specimen [Part]..

16.0(0.4)
9.5(0.32)
6.4(0.31)
26(0.-11)

DescrijJtion :-The characters of theshell of this species change to a large
extent with growth, as has been alreadymentioned by TOI<U:-\AGA and SHIMIZU(1926) and Sllli\IIZU (1926). The change
may be shown more clearly, at least insome respects, by the present two speci·
mens than by the holotype. There may
be also a certain extent of variation
within a species. Even if the shells of
the same diameter are brought into
comparison, they may be somewhat dis·similar.
The smaller specimen, GK. H5557 (Pl.
48, Fig. la-c; Text.fig. la-d), probably
represents a young shell. It is about
30 mm. in diameter and is wholly septate.
It has an umbilicus of moderate size
(about 31 percent of the diameter). It
is moderately involute, about a half of
the inner whorl being overlapped by theouter. The whorl of this stage is nearlyas high as broad and subcircular in theintercostal section, but it is somewhat
broader than high and rather polygonalin costal section. The ornament in thisstage consists of fairly strong tubercles
at about the middle of the flank (i. e.
slightly below the mid-height), about
eight in a half volution, simple, blunt

Breadth(Costal) (lnterc.)
15. 8x2
27. 5(0. 53) 25. 5(0. ·19)
20.0
15.5
16. 2(0. -!0) 13.2(0.33)
u 5(0. 48) 12.3(0.41)
12.3
10.8
10.8(0.53) 9. ·1(0. 46)
26 (0.42)

B.;H.
(C.) (I.)
0.8
1.58 1. ·17
1. 29 1.0
1. 29 1. 05
1. 20 1. 02
1.16 1. 01
1. 22 1. 06
1.18

ribs extending radially from the tubercle
to near the umbilical m:J.rgin without
umbilical tubercles. secondary ribs bifur·
eating from the mediolateral tubercles
with a moderate projection on the venter,
and weak tubercles at the ventrolateralshoulder on the secondary ribs. A low
siphonal keel is weakly undulated : each
undulation corresponds in number andposition with one of the secondary ribs
which form obtuse chevrons on theventer. The lateral tubercles on theanterior part of the preserved last whorlof this specimen are rather mammillate
and on the rest part they are bullate,although this is the character observedon the internal mould.
On the next inner whorl of the same
specimen distant, weak radial ribs, which
are broadened and elevated at the medio·
lateral tubercles are observable. The
tubercles look subangular and in contact
with the steep umbilical wall of theouter whorl. A siphonal keel. lateral
tubercles and ribs seem to appear atabout the shell diameter of 9 mm. The
flanks are somewhat flattened and theventer is low. roof-shaped in this stage,
although the details of the venter are
not clearly seen. The characters of still

480.

Yaheiceras (C1·etaceous Am.m.o11ite.~)Fig. la-d. }'abeiceras orientale TOKU>'AGA and SIII~IIZU. A
diagrammatic sketch (partly restored) of a young shell. GK. H555i.
Lateral (a). ventral (b) and sectional (c) views; External suture(d) at lhe stage marked as s in Fig. la. T. M. deli11.

inner whorl are nol well shown.
The suture at whorl-height of 10 mm.
on the outer whorl is illustrated (Text·
fig. ld). The heads of the saddles arearranged on a descending line. The
saddle between E and L is high andasymmetrically divided; the outer larger
branch is further subdivided. L is deepand has fairly deep and narrow. four,
asymmetrically disposed branches. E isnearly as deep as or somewhat larger
than L in this stage. Other auxiliaryelements (U 2 . adjacent saddles, subdividedU,) are smaller. All the clements aremoderately incised, ins pi te of the small.
immature state of the shell.
The other. larger, specimen. GK. 1-15556(Pl. 48 Fig. 2a-e; Text-figs. 2a-e, 3lshows slowly growing three outer whorls.
Its inner nuclear p:1rt as large as thepreceding specimen is not well preserved
for some reason. The observable young·est whorl, with height of 12.5 mm. andintercostal breadth of 13.2 mm., approxi·
mately represents the stage which
follows that of the smaller specimen
described above. It has a similarly sub·circular intercostal section and the same
type of ornamentation as that of thesmaller specimen. The lateral, rounded
tubercles become increasingly stronger.
There are fourteen tubercles in thewhorl of about 10 mm. in diameter.
Faint secondary ribs spring in pairs
from the lateral tubercles and they areprovided with faint outer ventrolateralor rather to say ventral tubercles on aline fairly close to the mid-venter. The
siphonal keel is low and faintly undu·
lated.
The succeeding whorl is unusuallydepressed and rather coronate in cross·section, with a much larger proportionof breadth to height (see Measurements).
It has a sleep umbilical wall and a32<1

Tatsu1·o M1l TSUMOTO et al.

/-

I

1

I
I

I

L_/
I'
\
\

c

30mm

Fig. 2a-e. Yabeiceras orientale ToKI:NAGA and Sm:-.11zu. A sketch of an adultshell, GK. H5556. Lateral (a) and sectional (b) views; a ventral view of the inner
whorl is shown in Fig. 2b. Ventral view (partly restored) (c), cross-section (d)and suture (e) of a part of the last second whorl. which is removed and only indicated by broken lines in Fig. 2a. The illustrated suture is at the stage marked
as s in fig. 2a. T. M. de/in.

rounded umbilical shoulder. It embraces
about one third of the inner whorl. The
lateral tubercles are strong, rounded at
the base and mammillate. The radial
rib which extends from the lateral
tubercle to the umbilical margin is bluntin this stage. The secondary. projected

ribs which spring outward and forwardfrom the lateral tubercles are also blunt,fading away on the venter, without
showing clearly the paired state. Thereremain, however, scarcely perceptible
elevations on the lines parallel to andnot far from the siphonal keel. They

480.

Yabeiceras (Cretacemts A'l'mnoni.tes)

look as if they formed very faint sidekeels. The siphonal keel itself is also
weakened and smoothed, although it ismore distinct than the side keels.
The last suture is exposed on the latepart of the tuberculate whorl. i. e. thelast second whorl (see Pl. 48. Fig. 2a ;Text-figs. 2a, 3J. Beyond the last suture
the flank of the whorl is less inflated
and the lateral tubercles are rapidly
weakened.
The suture in the stage of the strongly
tuberculate, depressed whorl is illustrated
in Text-fig. 2e. It consists of the ele·ments of E. L, U~. u. (=S). U1 • I. They
are in general moderately deeply incised.
E is broad and nearly as deep as L. L
is longer than broad and has narrow anddeep, apparently trifid branches, but its
stem is not much narrowed. U:: ismoderately broad and its minor incisions
are not very deep. The internal elements(I and U1) are narrow. The summits ofthe saddles are arranged on a line
descending to the umbilical seam. The
saddle between E and L is considerably
deeply incised and its stem is narrowed.
Its head is asymmetrically bifid, with anarrower and somewhat oblique branchon the side of L. The saddle between\\.\ . J%?\
_

.--

__

...._

..,

I

'··-"

I
I

5mm
__,

I

I

Fig. 3. Yabeiceras orientale Toand Sm:-.Hzu. Exposed partof the last suture on the last second
whorl of GK H5556 (see a sketch inFig. 2). Broken line indicates the umbilical margin of the outer whorl; thedotted line the mediolateral tubercle.
T. M. de/in.
Kt::"AGA:127

L and Ue is moderately broad. being
situated on the row of lateral tubercles.
It is as a rule divided but the outline
in detail may be modified. probably due
to the relation with the large tubercle.
The saddle between Ue and U3 is rather
unusually deeply divided.
The living chamber occupies more thana full volution in this specimen. A
posterior half of the last whorl is un·
fortunately missing. Presumably it had
been destroyed and gone before thedeposition, as is suggested by the dis·position of associated bivalved shells. As
far as the observable part is concerned,
the last whorl is quite dissimilar to theseptate one. It grows slowly and over·laps about one third of the inner whorl.
The umbilicus is wide, occupying wellover 40 percent of the entire shell dia·meter. The umbilical wall is steep andsomewhat excavated; the umbilicalshoulder is subrounded to subangular.
The whorl is considerably higher thanbroad, with a proportion of 8: 10 between
breadth and height. It is broadest between the umbilical shoulders; the flanks.
are convergent and only gently convex;
the venter is fiat or slightly arched. withextremely obscured side keels at theventrolateral shoulders and a faint remnant of the mid-ventral keel. The whorl·
section is, thus. roughly trapezoid. The
surface is smooth : tubercles and ribs,
with which the inner whorls are characteristically ornamented, completely dis·appear from the last whorl.
The character of the whorl changes
little within the last half volution. The
diameter of the shell at the last end of
the observable part is 123 mm. The
apertural margin is not preserved in thepresent specimen.
Occurrence:- The two specimens were
found in nodular calcareous sandstoneembedded in the Lower Conglomerate

~28

Tatsuro MATSUMOTO et a(.

J\Jember of the Futaba Formation, in theupper reaches of the Sakura-zawa. Oriki.
I-lirono-mura, Futaba-gun, Fukushima
Prefecture. tvlany bivalves of shallow
sea environments such as Glycymeris,.4piolrigonia, etc. are associated with theammonites.
Inoceramus uwajimet1.~is
YEH:\RA. an index of the Lower Uraka·wan (Coniacian), occurs abundantly inthe same member.

Remarks
Comments are given here mainly on
two points.-(1) Are the two described
specimens from Futaba really of identi·
cal species with the holotype of }'abei·
ceras orientale? t2) '\Vhat do the observed facts suggest about the affinities
of Yabeiceras?
Although the holotypc of Y. orientale
is at present inaccessible to the writers.
its characters are understandable to aconsiderable extent from the description
and illustration of ToKU~AGA and SHI~liZU (1926, p. 201, pl. 22, fig. 7 ; pl. 27.
fig. la-c). In the essential points thetwo specimens under consideration wellagree with the holotype.
The smaller specimen, GK. H5557. isapparently more narrowly umbilicate
than the larger one. This is probably
ascribed. at least partly, to the difference
between immature and grown whorls,
because the width of umbilicus in proportion to the shell diameter gradually
increases, as shown by the figures in themeasurements. There can be also slightdifference between individuals. The dimensions· of the holotype, as measured
by ToKU:\AGA and SHI:-.uzu, indicate asome\vhat smaller proportion of umbilicus to diameter than the present larger
specimen. GK. 1-15556. The difference is.
however. by no means great and canwell be regarded as being within theextent of variation.
This species exhibits a peculiar typeof change with growth in the shape of
the whorl. In the relatively early growthstage the whorl is nearly as high as
broad; the flanks arc rather flat when
the shell is 5 to 15 mm. in diameLer. andthen become rounded !in costal section).
In the middle growth-stage, with diameters over 30 mm. up to 60 mm. or so.
the whorl is broader than high and theflanks are inflated. The most depressed
whorl in the specimen GK. H5556. at adiameter of 52 mm., has the proportionof 1.47 bet ween breadth (intercostal) andheight. This seems to be somewhat
larger as compared with 1.18 at a diameter of 62 mm. in the holotype. The
exact comparison is, howeyer, difficult,without seeing the actual specimen. since
the change of dimensions with growth
was not indicated by figures in the ori·gina! description. The last whorl ishigher than broad and has smooth,
flattened flanks.
In this point thewriters' specimen agree with the holo·
type. although the ventral part of thelast whorl of the latter is much damaged.
In the diagnostic change of ornamentation the two specimens described above
essentially agree with the holotype. The
ribs which extend from the lateral tuber·clcs to the umbilical margin look moreblunt in GK. l-15556 than in the holotype.
The lateral tubercles in the present two
specimens seem to be situated on a line
somewhat lower than that in the holo·
type. The differences are again rather
slight and these characters change withgrowth even in one and the same individual. TOKUNAGA and SHIMIZU did notdescribe the faint elevations, i. e. thereduced outer ventrolateral tubercles,
which should be discernible on the480.

Yabeiceras tCretaceous Ammonites)

secondary, ventral ribs on the inner
whorls. This character is, however.
shown on the photographs (ToKUNAGA
and Sm~nzu. 1926, pl. 27. fig. I and alsopl. 16, fig. 15 [ V. lwtoi]).
The suture changes with growth and·can be modified in correlation with theminor details of a whorl-shape and also
strength and relative position of a tubercle. In the illustrated suture of thepresent larger specimen the saddle between L and U2 is broader than that of
the holotype. In the smaller specimen
it is as narrow as the latter. but thesaddle between E and L is comparatively
broad. This and other minor differencesin the suture between the present specimens and the holotype are not significant
enough to separate them into different
species. The general similarity is more
remarkable than the minor differences.
To sum up the specific identity of thepresent two specimens with the holotype
of Yabeiceras orientale is warrantable.
granted that there is a certain entent
of variation between individuals.
As regards the affinities of }·abeicerasorientale, Cenomanian Schloenbachia
NEUMA YR, 1875, was at first consideredby TOKlJ~AGA and SHI~f!ZU (1926, p. 201).
SHLMIZU (1926, p. 548) on another occasion
mentioned a similarity of Vabeiceras inshell-form and ornamentation to " thegroup of Barroisiceras desmo/insi." He(193-l. p. 78) ascribed Y a/Jeiceras to thefamily Priontropidae as understood atthat date. For about two decades nocomments had been given to the systematic position of the genus. until WRIGHT(1957. p. L ,129) placed it in the subfamily
Peroniceratinae of the family Collignoniceratidae. MATSUMOTo's opinion (personal communication in 1954) that Yabeiceras seemed to be somewhat similar to
Gauthiericcras was taken into consideration by WRIGHT.

:3:29

As has been described above, }'abeiceras orientale shows so particular typeof changes w·ith growth that it may be
a puzzle to tell its true systematic position.
In the evolute, widely umbilicate. andtricarinate (although very indistinctly so)shell. Yabeicems could be taxonomically
included in the subfamily Perniceratinae.
In fact the suture of Yabeiceras considerably resembles that of Peroniceras DE
GROSSOUVRE, 1894. The carinate andarched venter of the septate whorls of
Yabeiceras is similar to that of Gauthiericeras DE GROSSOUVRE. 1894, andCiryella WIEm.IA:-\N, 1959. The mediolateral tubercles are developed in the
middle growth-stage of Gauthiericeras
margae. the type-species of Gauthiericeras, if not so strongly as in the adolescent stage of Yabeiceras orientale. But
such an entirely smooth last whorl asthat of Y. orientale is not seen in any
well known examples of Peroniceralinae.
Some species of Peroniceras attain alarge size, as has been exemplified bythose described by DESIO (1936) and VA:t\
HOEPEN (1955). The last whorl in thosespecies is distinctly costate and tricarinate.
In Yabeiceras orientale the mode oflife must have been considerably different between the animal of the strongly
ornate and inflated whorls of the middle
ages and that of the smooth and compressed one of the adult to senile ages.
although the writers have not got enoughevidence in the mode of occurrence ofthe fossils.
In view of the moderate involution,
polygonal costal section. and a particular
type of ornamentation, the inner whorlsof Yabeiceras resemble those of Forresteria REESIDE, 1931. of the subfamily
Barroisiceratinae. In Forresleria not
only the mediolateral but also the ventro-

:t10

Tatsllro MllTSUJIOTO et al.

lateral and ventral tubercles are strong,forming angular shoulders, while in}·abeiceras mediolateral tubercles arevery strong but the ventrolateral andventral tubercles are weak and finallylost as the shell grows and it has noangular ventrolateral shoulders. On theouter, adult whorl of Forresteria themediolateral tubercles normally fuse withthe exaggerated ventrolateral ones (insubgenus Forresteria s. s.), or to the umbilical ones (in subgenus Reesideocaas),or disappear (in subgenus Harleites).
Har/eites is however, much cotupresscd
and involute, and has angular ventrolateral shoulders on which fine clavaeremain. Thus, }·abeiccras is related to
but is distinguishable from any subgenera of Forresleria.
As far as the described species areconcerned, Yabeiceras most closely resembles the subgroup of Forresteria
ampozaloalwense BASSE (1947, p. 131. pl.
9, fig. 9), in which the ornaments gradually fade away on the outer whorl.
BASSE's species is evidently more involute than }'. orieutale. It probably
needs a new subgeneric name.
The suture of the Barroisiceratinae isfairly variable, as has been demonstrated
by BASSE (1947) and others. One of theillustrated sutures of Forresteria forresteri REESIDE (1931, p. 17, pl. 5, fig. 7)
considerably resembles that of Yabei-cera.~

orientale (Text-fig. ld in this.
paper), although the former has broader
saddles than the latter. In general thesuture of Fabeiceras in the late growthstage is more deeply and finely incisedand has a narrower first lateral saddle
than that of Forresteria.
It should be noted that Eboroceras
magnumbilicat um BASSE (1946, p. 73. pl.
2, fig. 2a, b), from the Coniacian of
I\ladagascar, closely resembles Yabeicerasorientale. The former has weak umbilical tubercles in addition to the strong
mediolateral tubercles. This difference is
specific, for Yabeiceras koloi ToKUKAGA
and SHI!IHZU also has small umbilicaltubercles on the inner whorl. Eboroceras
BASSE, 1946, is certainlysynonym of
Yabeiceras TOKUNAGA and Sunnzu, 1926.
Further discussions and the final con·elusion about the systematic position of
Yabeiceras are to be reserved until more
examples of l"abeiceras and related
genera from llokkaido are described by
one of the writers (T. M.).

aReferences Cited
BAssi', Eliane (1946) : Sur deux ammonites
nouvelles du Coniacien du sud-ouest de
Madagadcar: Subbarroisiceras n. g. mahafa/ense n. sp. et Eboroceras n. g. magnumbilicatum n. :;p. Bull. Soc. Ceo/. France.
5 ser .. tJo/. 16. p. 71-76. pl. 2.

Explanation of Plate 48
Figs. la-c, 2a-e. Yabeiceras orientale ToKUNAGA and Stlt~IIZU.
1. A young shell, GK. H5557, from the upper reaches of lhc Sakurazawa. Oriki. Hironomura. Futaba-gun. Fukushima Prefecture. Lower member of the Futaba Formation.
Two lateral (a, b) and ventral (c) views, x3/2.
2. An adult shell. GK. H5556, from the same locality and member. Lateral (a) and ventral(b) views of the main part. Natural cross-sections and ventral views of inner whorls(c). Lateral (d) and ventral (e) Yiews of a part of the last second whorl. which is removed in Fig. 2a: a-e all in natural size.
(Photos by I. HAYAMI. without whitening)MATSUMOTO et al.:Yabeiceras

Plate 48

480.

Yabeicera.s (Cretaceous Ammon·ites)- - (19•17): Paleontologie de Madagascar
N. xxvi-Lcs peuplements Malgaches deBarroisiceras. l~evision du genre Barroisiceras DE GRoss. .4nn. Pa/eo11t .. vol.
33, p. 100-190, pis. 7-15 [1-9].
DI·:sJO, A. (1920): La creta nel bacino di.Firenze. l'aleontogr. Jtalica. Nil. 26, p.
187-243. pl. 12-19 [1-8].
·GRossouVJIE, Albert DE (189-1): Recherches
sur Ia Craie superieure. II Paleontologic.
Les ammonites de Ia Craie superieure.
Mbn. Carte geol. del. France [1893]. 264
p., 39 pis.
HOEPDI. E. C. N. \'A~ (1955): Turonian-Coniacian ammonites from Zululand. South
African jour. Sci., 1'0/. 51. no. 12, p.
361-377. includ. figs. 7-31.
NEUMAYJI, M. (1875): Die Ammoniten der
Kreide und die Systematik der Ammonitiden. Zeitschr. Deutsch. Ceo/. Gesell.,t•ol. 27, p. 85•1·892.
REESIDE. J B. jr. (1931): The Upper Creta·
ceous ammonite genus Barroisiceras in
the United States. IJ. S. Geol. Sul'l·., Prof.
Paper. 170-/1, p. 9-29. pis. 3-10.
SJJJ~II7.U, Saburo (1926):
Three interesting
Cretaceous ammonites recently acquired
from llokkaido and Saghalien. Proc.
Imp. :lead. (Japan), 1'01. 2. no. 10. p. 5•17550.

- - (1934): Ammonoidea. /11 SJJJ~JJZU, Saburo and 0BATA. Tadahiro(HJ3-l):Cephalopoda. IH,anami's Ser. Ceo/. & Pa·
laeont .. 137 p .. Tokyo (in Japanese).
ToKt::-iAtiA. Shigeyasu and SIJJ~JJZt:. Saburo(1926): The Cretaceous formation of
Futaba in Iwaki and its fossils. jour .
Fac. Sci .. Imp. Uni1·. Tokyo. sect. 2, vol.
1, pt. 6. p. 181-212. pis. 21-27.
WJED~JA:-;:-;. Jost (1959): Le Cretace super·
ieur de t·Espagne et du Portugal et ce~
cephalopodes. S.Je Congres des Soc. Sat•·antes. 1959. p. 709-764. includ. pis. 1-8.
WRIGHT. C. W. (1957) In MooRE. R. C. [Editor] (1957) : Treatise on !lwertebrate
Paleontology. Pari L. Mollusca. Cephalo·
podea. Ammonoidea. p. L1-L·190, Geol.
Soc. Amer. & Univ. Kansas Pre3s.

Locality Guide
Upper reaches of Lhc Sal<urazawa, Oriki.
Ilirono-mura,Futaba-gun. Fukushima
Prefecture(Approximately 1-10•57'·15'' East Long .. 37" 12'
10 11 North Lat.)l'Mh'li!IIUJOll1HIIIAll!f.f'HJl'*ti(iRJ-.mc (IJil U'l!!l~fli:Ji: 5
li'if(f) 1 Jt!!lfW<l~~i f'llliliJJ {f)Jfill!lllllb·l?~t•1050 rn (lij ·1250 m GI)Jt!!l.r:)Trans. Proc. Palaeont. Soc. Japan. N. S .. !\o. 56. pp. 332-344. pl. 49. Dec. 20. 196.!

481.

ON SOME LOWER AND MIDDLE TRIASSIC
A:\I::\IONOIDS FROl\I JAPAN*
YUJI BANDO
Department of Geologr. Kagawa Uniyersitya*~· 1ft • -r;;4::=:·1!H-c7 :/ .;:7·1 1·1.:."?\·-c: r b4C·~J:e 7:/ -r:7·1 1-.:: '- -c~!@!f!.Jtf':{:'f!llll~.lhl(Jli~IJ#.U~. lllf!!il:fJ!UJif!li!l~rf. Hemiprioniles !~<1)-·~ff.JU: !'arananni/es f~t::.=..

:2(1)7:/-t::J-11·. rp,'if>.:_J~:;f:C7:/-t:-t-1 r.!:L.-crti&9~;:ftJ::iliJU!. ·,~;~~:;J>:;trt1Uktft111Jffrll.!.iJiJ(l)fJrl!iiJiilffl. :t,; .t lfflllfliti:ltJJ(I)fJJif.f~b ·I? iff. if\ L.t..:l&t.f!(l) t (I).!:, [.TLJIH (1)/t!Jii;!J;!;JJil~IDU::;IJf,'Z'{IItJ*'t<:· 1LllE1i(I)[:IW~~~!IiJ·I?;lb 1? t.:I.:.Jff:/1\ L.t..: 7 =- ;;-- .7 !1 !l'J 7 :/ -t: -r 1 1-. Ptychites sp.
;t.o .t 0: Beyrichil es cf 1·ot ellijorm is (MEn:) (l)__:.j:.ji :a-:Tc!f\H-& ;•;· L. n "?l~l~~,li[j(l)~f\:;;.:.
0t,-c7.::.::,.-.7;.-i.:.:h.t ..;;nrugti:(I);Jr,~.:..!::a-,ifilct.:,
JJ.i #!*IT TiJ

Abstract

The Lower Triassic Anasibirites fauna(Ilemiprionites shimizui BAJ\DO. n. sp.,
l111asibirite.~ sp .. Hemiprionites sp.) described in this paper from the Tao For·mation at Tao (Taho) near Uonashi,
Ehime Prefecture, Shikoku, including
those previously mentioned (BA:-<oo, 1961.
1964'!, belongs to the Upper Owenitan
and is represented by the Anasibirites
mult~formis zone. Some !\Iiddle Triassic
ammonites (Balatonites cf. gottschei(MOJSISOVICS), Ptychites sp .. Paracerati/es sp. and Beyrichites sp.) are also described in this paper. The t-.-liddle Trias·
sic ammonites are from the lsatomae
Formation (Anisian) in the Kital<ami
Massif and the Zohoin Formation (Ladinian) of eastern Shikoku. The ammonites
from the Zohoin Formation at Gorodani,
Nagayasu. Kaminaka·cho, Naka-gun,
Tokushima Prefecture in eastern Shi·kol<u, suggest that the age of the Zoho·
in formation may be Anisian to Ladini·an.
* Received August 24. 1961: read Sept.
20. 1964. at Hiroshima.

Geological and chronological reviewof the fossil localities

The Triassic formations of the Kita·
kami Massif are included into twogroups, i.e. the lnai and the Saragai.
The Inai Group consists of four forma·tions the Hiraiso. Osawa. Fukkoshi andthe lsatomae in ascending order. The
first two formations are regarded as of
the Scythian stage and the later two asof the Anisian stage. On the other hand,
the Saragai Group is Upper Triassic inage and belongs to the upper Carnian to
Rhaetian?.
The Anisian ammonites from the Isatomae Formation have been known since
many years ago and are characterized
by and called the Hol!andites fauna. In
the southern part of the Kitakami ~vlassif
the Triassic beds show monoclinal structure along the Paci fie coast whereas thegeneral structure is of large synclinalform \liith NNE-SSW trend of axis which
The
plunges SSW with low angle.
general order is normal, but the ScythoAnisian beds and Aniso- Ladinian beds
show different distribution, and forma-481.

Lo·wer and Middle T·riassic Ammonoids

333

T.
Fig.!. MAP

OF JAPANSHOWING

LOCALITY

iCc, r

OF

~r

"""

TRIASSIC

'I

~

10'

AMMONITES 4 4 - - - - + - - - ;\'---~

/'.

~'(
v__,_

138'

~

'I
~KITAKAMIRIFUH--··;.<·· 38'

\
I

130'

IWAI

-if1
LEGEND.t.

Upper Triassic ammonites

X Middle Triassic ammonites
•

Lower

Triassic ammonites

Fig. 1.

tions correlative with upper Ladinian tolower Carnian are missing in this district.
On the synclinal axis. the lJpper Triassic
Saragai Group and lhe Lower Jurassic

formations rest on the Middle Triassic
lnai Group, but their distributions are
restricted to the eastern wing of the
syncline. No Lower Triassic ammonite

Yuji BANDO

Fig.2.1nde)( map show ng
localities of described sp
ecimens from Eest.Shikoku
SHOBU

t

HAIGYU.

0Fig.

has been recorded from the Hiraiso Formation, but the Osawa Formation yieldeda single characteristic ammonite, Subcolumbites cf. pcrrini-smilhi ARTIIABER(BA:>~no, 1%1).
The locality of Paraceralites sp. and Baratonites cf. gottschei(MOJSISOVJCS), both described in thispaper, is about 300 meters stratigraphically above the locality which yieldedthe Lower Triassic ammonite, and its
geological horizon corresponds to the
middle part of the lsatomae Formation.
The Triassic formations in the Naka·
gawa region, Tokushima Prefecture,eastern Shikoku are distributed in theso called Chichibu Terrane with aschuppe structure and with a generalstrike of ENE- WSW trend similar to the
Paleozoic and tvlesozoic strata. The for·mation of the locality of Ptychites sp.
and Beyrichiles sp. is the Zohoin Forma·

2.

tion and the age of this formation isconsidered to be upper Ladinian. Recent·ly NAKAZA WA (1964) described someAnisian ammonites i.e. Ussurites yabei
DIENER, Ilollandites sp. and Balatonites
sp. from the Zohoin Formation(?) at Uonashi in western Shikoku. From the Zohoin
Formation at Gorodani in eastern Shikoku, the occurrence of a Ptychites likeammonite was reported by KosA YASHI
and ICHIKAWA (1951, p. 93) and the material described in the present paper may
probably be of the same species. How··ever, the Zohoin Formation at Annamota of Nagayasu yielded upper Ladinian ammonite, Protrachyceras cf. p.~eudo­archelaus (IAUHE), (BANIJO, 1963, 19(}1.l.
Accordingly the writer considers that
the Zohoin Formation may range fromthe Anisian to upper Ladinian, although
no lower Ladinian formations or beds

~Rl.

Lo·wer and Middle Triass·ic Ammonoids

correlative with the Rifu Formation havebeen discovered in any known outcropsof the Zohoin Formation in Shikoku.
However the material at hand is remark·able because the specimen of Ptychites
sp. contains a single fragmental specimenof Daonella shell. Consequently, the paleontologic evidence shows that the geologic horizon of the formation whichyielded Plychiles and Beynchites may
not be Anisian but should be taken as
the lower Ladinian. Consequently, theformation which yielded those ammonites
may be correlated with the Rifu Formation of the southern part of the Kita·
kami r..Iassif in Northeast japan.
Acknowledgements

The writer expresses his cordial thanks
to Dr. Hisakatsu YARE. professor Emeri·tus of the Tohoku University, for hiskind guidance and valuable suggestions
and to Professor 1\otora HATAI of theInstitute of Geology and Paleontology of
the Tohoku University for his continuous
guidance. The writer also thanks Pro·
fessor Yoshio ONUKI of the Departmentof Geology, Faculty of Education, To·hoku University, for his encouragements
in many ways. Concerning the present
work, acknowledgements are also due toAssistant Professor Kazumi SUYARI, De·partment of Geology of the Tokushima
University, his valuable suggestions and!VIr. Kagetoshi H:\SHDIOTO, the collectorof the fossils.
Finally the writer his indebtedness tothe ~Jinistry of Education. for a grantfrom the Science Expenditure Fund.
Description of Ammonoid fossils

Family Prionitidae HYATT, 1900
Genus Ilemiprionites SPATH, 1929

Ifemiprivnites :;himizui BANDO, n. sp.
Pl. 49, Figs. 1, 2

Description :-Shell involute, laterallycompressed, discoidal, rather deeply embracing, umbilicus narrow. Side slightly
convex from umbilical shoulder to ven·
tral shoulder. Venter tabulated. neither
keels nor furrows. Outer whorl indented
at about one-fourth height by inner
whorl. Height of whorl a little less thanhalf of total diameter of shell, shellwidth about two-thirds of shell height.
Umbilicus slightly opened, umbilicalwidth a little narrower than one-fourthof total diameter of shell. Sides ornamented with radial folds, which aremost elevated at half height of shelland diminish on umbilical margin. about
7-8 ribs in quarter part of outer whorl.
Sides gently convex at a little below
one-half height of \Vhorl, where is maxi·
mum thickness. Venter tabulated andventral edge slightly bicarinated. peripheral margin of side abruptly becoming
flattened.
Septa caratitic. consists of roundedsaddles and serrated lobes; peripheral
lobe rather deep, divided by small angu·
Jar saddles into four narrow entire lobes,
medial lobe, shallower than peripheral
one, divided into three lobes, wedge·
shaped and terminates in two points.
Umbilical lobe shallowest, separated into
three to four lobes and three smallrounded saddles. External saddle higher
than first and second saddles, and roundly
arched without serration, and second
saddle lowest.
Remarlzs :-Two specimens \vere examined. This new· species resembles
llemiprionites tahoensis (YEHARA) in thewhorl shape, umbilicus and septal features, but can be distinguished from itby having distinct radial ribs on the

Ynji. BANDO:\t/easurements (in mm.) :-I
IGPS •, col\. cat. no. 45158
GLKU••-Cl58

D

H.

H/D

\V.

27.5

13.0
17.0

0.47

8.0
9.0

0.62
0.53

U.

U/D

6.5

0. 2·1• IGPS-Abbreviation for the Institute of Geology and Paleontology, Tohoku
University. Sendai. Japan. •• GLKU-Abbreviation for the Department of Geology.
Kagawa University. Takamalsu. Japan.

surface. abruptly rounded umbilicalshoulder, and higher external and firstlateral saddles. The present species issimilar to H. morianus (YEIIARA) andthe typical form of H shilwkuensls (SHI·MIZU and ]IMBO) from lhe same locality
at Tao in the septal feature and umbilicus form, but differs in the shell ornamentation. The latter species has asmoother surface without radial ribs anda narrow umbilicus than that of thepresent species. On the other hand, thepresent new species is similar to H. typus(WAAGE:-<), which was described fromthe Upper Ceratite Limestone of Chideru,
Sail Range, by WAAGEN (1895), in the
septal feature and shell form, but differs
in the shell sculpture.
Occurrence and geological horizon:Argillaceous limestone of the Tao Formation at Tao near Uonashi, Shirokawacho, Higashiuwa·gun, Ehime Prefecture.
Upper Scythian. Anasibiriles mull(formis
Zone of the upper Owenitan ammonite
stage.
Reposilory:-lGPS, coli. cat. no. 45158
and GLKU-C158 Coli. H. YABE, T. SUGI·Y:UIA, S. SIII:\IIZU. N. Jnmo and Y.
BAN DO

1/rmipronites sp.
Pl. 49. Fig. 4

DescrifJ/ion :-Shell laterally compress·
ed, involute. discoidal. with small umbilicus. Outer whorl indented at about
one-third height by inner whorl. Venter

narrowly tabulated with sharply angularventral shoulder. Umbilicus very small,with abruptly rounded umbilical shoulder.
General shape of umbilicus a funnel
shape. Shell surface ornamented with
faint falcoid ribs which diminish on
peripheral and umbilical sides: mostprominent on two-thirds height of outerwhorl. and each ribs feeble on the urn·
bilical side. Height of shell about three·fifths of total diameter of shell. Widthof outer shell about three-sevenths ofheight. \Vidth of umbilicus about 1/25
of total diameter of shell and almost
closed.
Septa ccratitic, with rounded entire
saddles and denticulated lobes. Siphonal
lobe narrow, siphonal saddle narrowly
arched; external lobe shallower thanfirst lateral one, denticulated into threelobes by two narro\v saddles: external
saddle higher than other saddles; firstlateral lobe deepest, widest, and corru·
gated by small saddles into three lobes;
first lateral saddle roundly arched: second
lateral lobe slightly corrugated into twopoints; auxiliary series consists of lower
and wider saddle than any other saddles,
umbilical lobe slightly serrated (See Fig.
3l.

+\,I\~J'\_-~ I
0Fig. 3:

0.5

I em.

Suture line of l!t•miprionites
sp. GLKU-Cl5<1. X~

481.

Lower and Middle Triassic Ammonoids

Measurements (in 111111.) :--GLKU-Cl5.t

D.

II.

H/D

w.

W;H

u.

U;D

25. 0(?)

14.0

0. 56(?)

6.0

0.43

1.0

0.04

Remarl?s :-A single specimen was examined. A part of the venter is broken
off. The features of the septal element,
umbilicus width. and whorl shape resemble with those of Hemiprionites
kuharanus iyonus BA1\DO, but the ma·
terial is distinguished therefrom by thefeatures of the lateral lobes and the shellornamentation. The general character ofthe septa is of the Ilemiprionites type.
Occurrence and geological horizon:Argillaceous limestone of the Tao Formation at Tao near Uonashi, Shirokawacho, Higashiuwa-gun. Ehime Prefecture .
.-l.nasibirites multzformis Zone (upper
Owenitan). upper Scythian.
Repository:- GLKU-Cl54. Coli. Y.
BAN DO

J\na.-;ibirites sp.
PI. -19. Figs. 3a-c

Description :-Shell laterally compresseel. evolute, umbilicus slightly opened,surface smooth with fine radial striations.
Umbilicus rather small and narrow, butslightly opened. Ventral part broken.
Umbilical shoulder abruptly rounded andof rather funnel shape. Total diameternot measured because of broken ventral
part Septa well preserved, of ceratitic
form, with rounded entire saddle andwell serrated lobes. Ventral suture unknown, bul auxiliary series well preserved.
External saddle very high,rounded and broader than other saddles :first lateral saddle rounded and shorter
than external one: second lateral saddle
lowest and narrower than other lateral
ones. External lobe well serrated byconsiderably deep narrow lobes into very
high, narrow saddles. about 8 saddles;
first lateral lobe well corrugated to near
top of first lateral saddle by narrow lobes
into 6 narrow saddles. Auxiliary lobes
rather smooth with about 3 very small
lobes. Second lateral saddle on umbilicalshoulder and auxiliary series on umbilicalwall.

Measurements (in mm.) : D.

GLKU-CIS5
-----

j_

H.

H/D

22. 0( ?)Remar!?s :-A single specimen was examined. A part of the ventral portionand half of the outer volution of thespecimen is broken, but the sept:tl featun:'s are well preserved and differ fromany known Anasibirites species. The
shell ornamentation is similar to Ana.~i-

w.

\V/H

12-0

0.55

u.

u;o

7.0

--------

birites onoi (YEIIAI~A) and i1. archiperi·
phera.'i BANilO. but is distinguished fromthem by the septal features. judging
from the character of the ventral formand shell ornamentation, the present
specimen belongs to the genus Anasibirites. The general character of thel'n)'i BANDO
Pl. 49, Figs. 10, 11

material is most allied to A. archipcri·
pheras, but the present specimen shows
deeper corrugation in lateral lobes than
the above species. The feature of the
septa is not identical with any known
species of Anasibirites and it seems to
be of a special type.
Occurrence a11d geological horizon:Argillaceous limestone of the Tao For·mation at Tao near Uonashi, Shirokawa·cho, Higashiuwa·gun, Ehime Prefecture,
western Shikoku. Anasibirites multi·/armis Zone (upper Owenitan1, upper
Scythian.
Repository:- GLKU-C155. Coli. Y.

Description :-Shell form involute. subglobose, slightly depressed, with broadly
arched venter. Sides smooth, with a
few radial irregular folds. Umbilicus
very small, funnel shaped, nearly closed.
Inner whorl of globose form. Venter
broad and gently arched.
Suture lines ceratitic, with broadly
serrated ventral lobes, and large external
and internal saddles. Ventral lobe slight·
ly corrugated, external lobe serrated into
five points. Auxiliary series not observed because of poor preservation (SeeFig. 4).

BAN DO

Family Paranannitidae SPATH, 19:30?

Subfamily Paranannitinae SPATH, 1930

0.____ _ _0.5.......__ _ _..JI em.

Genus Paranan11ites HYATTand Sl\IITH, 1905

Fig. 4:

Paranannites sp.

Suture line of Paranannites
sp., GLKU-Cl56. x4

Measurements (in 111111.) : -GLKU-Cl56

D.

II.

H/D

\V.

Wfii

u.

U/D

10.0

6.0

0.6

8.0

1.3

1.:3

0.15

Nemarlls :--Two specimens \Vere examined. The materials were collectedfrom a horizon slightly above the Anasi·
birites-Hemiprionites beds (about 5-7 m.
above) of the Tao Formation at Tao.
The present materials belong to thegenus Paranannites and show the char·actenst1c form of the whorl and septal
features. Especiall)'. the present form
resembles Paranannites gracilis KIPARI·SOVA (KIPARISOVA. 1954, p. 21, fig. 1;
1961, p. 125. pl. 28, figs. 3-4), which was
described from the Olenekian beds ofRussky Island in Eastern Siberia, in theshell form and sutures. On the other

hand, in the septal feature, the present
form resembles P. aspensis HYATT and
SMITH (1905, p. 81, pl. 78, figs. 1-3. figs.
4-30; pl. 8, figs. 1-15; SPATH, 1934. p.
190, pl. 14, figs. 6a-c. text·figs. 5a, h),from North America. But the specificidentification with them cannot be done
because of the poor preservation of theauxiliary series; the present materials
are young forms.
Occurrence and geological horizon:Argillaceous limestone of the Tao Formation at Tao near Uonashi. Shirokawa·cho, 1-ligashiuwa-gun, Ehime Prefecture.
Anasibiriles muliiformis Zone (upper

481.

Lowe1· and Middle Triassic Ammonoid.~

Owenitan), upper Scythian.
Repository:- GLKU-C156.

Coli.

Y.

HAl'\ DO

Family Balatonitidae SPATH 1951
Genus Balatonites JvloJsrsovrcs. 1879
Balatonites cf. gottschei(MOJS!SOVICS)

339

ing. The present form is also alliedwith B. kitakamicus (DIENER~. but in theshell ornamentation it is more similar toB. gottschei than B. llitakamicus.
Occurrence and geological horizon:Paraceratites trinodosus Zone? of thelsatomae Formation at lsatomae. Utazucho, Motoyoshi-gun. l\1iyagi Prefecture.
Repository: - GLKU-C152 Coil. T.
KAl\IAOA. 1962

Pl. 49, Fig. 8
Cf.

Arpadites gottschei. MoJSISOVICS. 1888.
Ueber einige japanischen Trias-fossilien,
Beitr. z. Paliion/. Oest.-Ung. zt. d. Orients,
Bd. 7. Heft 2. p. 170 (10). pl. 2, figs.
2a-b.
Balatonites gottschei (MoJsiso\·cs), SHI·:\IIZU, 1930. On Some Anisic Ammonites
from the Hol/andites Bed.; of the Kitakami Mountainland, Sci. Rep., Tolloku
Imp. Uniz•., 2nd Ser. (Ceo/.), Yo/. l.J,
no. 1, p. 64.

Description :-Shell evolute, laterallycompressed, with ornamented surface.
Sides almost flattened, with numerous
radial ribs. Umbilicus and venter indis·
tinct. Shell ornamentation of numerous
radial ribs composed of primaries andsecondaries, latter start from ventral
shoulders and die out on middle portionof flanks. Ventral end of each ribs elevated and with a node. Middle portionof flanks with small tubercles arranged
in concentric line, about 9 nodes on onethird portion of outer whorl, and situatedon primary rib. Radial ribs inserted inrather deep radial furrows. About three
to four ribs to each furrow.
Remarks:- The present specimen re·sembles Balatonites gottschei (JiviOJSI·
sovrcs) from the lsatomae Formation
at Minato near Inai. lnai-mura, Ojikagun. Miyagi Prefecture (MOJSISOVICS,
1888) in the shell ornamentation, but
strict identification is impossible because
the ventral part of the specimen is miss-

Family Ptychitidae .MoJsrsovtcs. 1882
Genus Ptyclzites 1\IOJSISOVICS, 1875
Ptychiles sp.
Pl. 49, Fig. 7

Description :-Shell laterally compressed, discoidal. involute. with narrow
funnel shaped umbilicus. Shell with almost smooth surface. without ribs orfolds. Venter roundly arched in shape.
Thickness of shell greatest at umbilicalshoulder which is abruptly rounded:their wall of funnel shaped. Umbilicus
considerably narrow and deep. Heightof shell about two-fifths of total diameter of shell and width about one-thirdof height. Umbilicus about one-seventhof total diameter of shell. Septa unknown.
I

I

''\

5

Cl'lt

'

\
'
,.:'
I

t\:

:'

'
'
''

Fig. 5. Ptychites sp. Cross-section of theouter whorl. Specimen. GLKU-Cl50. x 5

340

Yuji RANDO
1\leasurements (in mm.):-

GLKU-Cl50

D.

H.

H/D

65.0

28.0

0.-13

Remarks:- The present material showsa Ptychitid shell form in the generalshell shape, whereas the ornamentationof the shell resembles the Arcestid or
Joannitid type rather than the Ptychitid
ammonite. Not with standing the miss·ing of the septa and constrictions or stri·ations, which are characteristic of theshell ornamentation of the genera Arces·
tes and foannites, the umbilicus widthand form of its shoulder most resemble
those of the genus Ptychites. But specificidentification is reserved due to the poor
preservation of the material.
Occurrence and geological horizon:-

w.

WfH

u.

U/D

9.0

0.13

Dark gray calcareous fine sandstone of
the Zohoin f-ormation at Gorodani Naga·yasu, Kaminaka·cho, Naka·gun, Toku·shima Prefecture. i'vliddle Triassic, upper
Anisian?.
Repository: - GLKU-Cl50 Coli. K.
HASHI:\!OTO

Family Ceratitidae lvloJSISOVICS, 1879
Genus Paraceratites HYATT. 1900
Paraceratites sp.
Text-figs. 6a-b: Pl. 49, Fig. 6

6 a

Figs. 6a, b. Paraceratites sp.
6a. External mold and gumtype of fig. 6b, GLKU-Cl51. x.J/5.
6b. External mold, GLKU-Cl51, X·l/5.
Loc. L;atomae. Utazu-cho. 1\lotoyoshi-gun. Miragi Prefecture.
Isatomae formation of the Inai group. Anisian.

Lower w11cl :1·hddle Tri.assic i11n-nwnoids

481.

Description:- Laterally compressed,
evolute, discoidal shell, with shallow umbilicus and well sculptured shell surface.
Whorls high, rather deeply embracing
and increasing rapidly in height. !Ieightof whorl a little greater than half oftotal diameter of shell. Diameter ofumbilicus about one-third of diameter ofshell. Surface of shell ornamented withfine flexuous ribs and comparatively
broad radial ribs, which start from umbilical shoulder and end at half of heightwith lateral tubercles. Tubercles consistof umbilical, lateral and marginal ones:
first one weaker than other tubercles
and lateral one most prominent. Lateral
tubercles prominent at inner-half of outer
volution. about 6 to 10 in outer volution,
but diminished on body w·horl. l'v!arginal
tubercles about 20 to a revolution andtheir position correspond to marginalend of ribs. Yen tral part not preserved.
Septa unknown.

Measurements (in mm.) :----~~~=~~-

w.

D.

II.

HJD

79. oc ?):lo. oc ?)o. 38(?)

Remarks:- The present material, unfortunately, lacks its ventral part andsepta. In the shell sculpture and umbili·
cus, the material resembles Paraceratites
trinodosus (1\loJISISOVJCS), bul differs in
the whorl indentation and umbilicus
width. The same may be said between
the material and Paraceratites clarkei(S~IITII) and moreover. it is more laterallycompressed than those two forms. Paraceatites erich SC\'IITJJ (1914. p. 87, pl. -17,
figs. 19-24) is also similar to the present
material in the shell sculpture. especiallyin the arrangement of the tubercles andribs, but specific identification is reserved
because of the poor preservation of thekeel-like ridge, which characterizes P.
cricki.
The three species of Paraceratiles
from Japan are from the Rifu Formation:
they are P. aff. trinodosus (MoJsrsovrcs).
P. cf. wanli (S~IITH) and P. orienta/is(YARE and SHil\IIZU), which were described by YARE and SAI~llZU (1927) and
SHn1JZU !1930). On the other hand,recently, the present writer described
three species of Paraceratites, i. c. P. cf.

W/H

u.

U/D

29.0

0.36

clarkei (S~llTH 1, P. orienta/is (YABE andSHI~IIZU) and P. cf. trinodosus (!\1oJSISOVICS), from the

Rifu Formation of thesouthern part of the Kitakami t\1assif
fBA:-;no. 1964, Triassic stratigraphy andammonite fauna of Japan). Recently
NAKAZA\\'A (1963) reported on the occurrence of Paraceratites sp. with Ifollandites sp. and Ussurites vabei from theAnisian calcareous sandstone at 500 m.
west of Uonashi-bashi, Shirokawa-cho,Higashiuwa-gun, Ehime Prefecture. western Shikoku. The present material is
the first occurrence from the type locality of the Isatomae Formation in thesouthern part of the Kitakami :\lassif.

Occurrence and geological horizon:Paraceratites trinodosus Zone? of the
Jsatomae Formation at Isatomae, Utazucho. Ivlotoyoshi-gun. i\liyagi Prefecture.
Repository :-GLKU-Cl51 Coli. T. KA·
l\IAKURA. 1962
Family Beyrichitidae SPATII. 1931
Genus Beyrichites WAAGE=--. 1895

Beyrichi tes cf. rotelliformis (t\lEEK)Yuji RAlVDO~42

Pl. 49, Fig. 5

Cf.

the Hritish Museum (Nat. llist.). Pt. 4.
The Ammonoidea of the Trias, p. 422.

Gymnoloceras rotelliformis, MEEK. 1877.
U.S. Ceo(. Expl. 40th Par.. l'o(. 4. p.
111, pl. 10. figs. 9-9a.
Beyrichites rotelliformis (MEEK), S~ttTII.
1904, The comparative stratigraphy ofthe marine Trias of western America.
Proc. Calif. Acad. Sci .. 3rd ser .. Geol .. 1•ol.
I. ITO. 10. p. 379. pl. 45. fig. 5: pl. 43.
figs. 13-14: HYATT and S~IITtl. 1905.
The Triassic cephalopod genera ofAmerica, U.S. G. S. Prof. Paper .JO, p.
155, pl. 23. figs. 1-11: pl. 58. ftgs. 1-15:
S:o..IITII, 1914. The middle Triassic marine
invertebrate faunas of North America.
U.S. G. S. Prof. Paper 83, p. 118, pl. ·1.
figs. 1-7: pl. 8. figs. 1-15: pl. 4. fig. 9:
pl. 31, figs. 1-6: pl. 91. figs. 1-10: Dmi':ER.
1915, Fossilium Catalogus. 1. Pt. 8. Cephalopoda triadica. p. 70: SPA Til. 1934.
Catalogue of the Fossil Cephalopoda in

Description :-Shell involute, discoidal,
laterally compressed. Umbilicus very
narrow and shallow. Venter slightly
rounded. Sides almost flattened andsmooth with falcoid radial folds. Heightof whorl about half of outer revolution.
Umbilicus width about one-fifth of total
diameter of shell. Shell width about
two-fifths of height.
Surface with numerous simple, fine,
flexuous or falcoid ribs and radial striaeof growth. with falcoid curve on flanks
sharp bend foreward just below ventral
shoulders. Each rib strongest on flanks
and becomes weaker near umbilical andventral shoulders. About 6 to 7 ribs inquarter part of outer whorl. Septa unknown.

Measurements (in mm.) :-GLKU-C153

D.

H.

H!D

w.

W/H

u.

U/D

31.0

15.0

0.48

6. 0( ?)

0. 4( ?)

7.0

0.22

Remarks:- The present material, unfortunately, lacks a part of the outerwhorl and the preserved volution is the
body whorl. The present specimen isvery similar to Beyrichites rotelliformis(MEEK), which was described from the•· Daonella dubia zone" of North America
by MEEK (1877). SMITH (1904). HYATTand SMITH (1905) and S~I!TH (1905) and
Sl\HTH (1914), in the shell sculpture andumbilicus form. From the American B.
rotelliformis the present one may be
distinguished by its more compressed
whorl. The present material also resembles B. falcifonnis S~IITH, which was
first described by Sl\HTH (1914) from theMiddle Triassic ·• Daonella dubia zone··(Paraceratites trinodosus subzone) of
Fossil Hill. West Humboldt Range, Ne-vada, but the present form differs fromit in its more compressed whorl. The
genus Beyrichites from the Zohoin For·mation of the outer zone of southwest
Japan is the first occurrence, and moreovere, this specimen suggests the occurrence of the Anisian beds in the Chichibu terrane of eastern Shikoku. In western Shikoku, NAKAZAW A (1963) discovered deposits of Anisian age andsome characteristic ammonite fossils,
Ussuriles, Paraceratites and !Iollandites,from the calcareous sandstone of the
Zohoin Formation at Uonashi, Shirokawacho, II igashiU\va·gun, Ehime Prefecture.
Occurrence and geological horizon:Anisian? beds at Gorodani. Nagayasu,
Kaminaka-cho. Naka-gun, Tokushima
Prefecture.

Lcncer a11d klidd/e Triassic Ammonoids

481.
Repository: -GLKC-C153

Coil.

K.

IIASHil\IOTO

Beyrichl/es sp.
Pl. 49. Figs. 9a-b

Description :-Shell laterally compressed. involute. discoidal, with funnel shaped
small umbilicus and narrowly rounded
venter. Surface with flexuous falcoid
Height of lastribs and striations.

whorl about a half of total diameter ofshell. Width of outer whorl about one·
thirds of height. \\1idth of umbilicus.
about one-sixth of total diameter ofshell. Umbilicus wall of funnel shape,.
its shoulder abruptly rounded. Venterof narrowly arched shape, its shoulder
rounded or slightly acuted. Width ofshell largest at a little below middle
height. Surface ornamented with flexuous, falcoid striations and foldings. Septa
unkO\Vn.

Measurements (in mm.) :-CLKU-C157

D.

H.

H;D

w.

w;H

u.

U;D

64.0

33.0

0. 51

11.0

0.33

7.0

0.11

Remarks:- The present material unfortunately lacks a half of the outer
volution. but the features of the umbilicus, shell sculpture. and venter are wellpreserved. Considering from the whorl
shape and shell ornamentation the pre·sent material belongs to the genus Bey·richites, but specific identification is re·served because of their septal elements
missing and due to the compressed de·formation. The shell sculpture of thepresent specimen most resembles that of
Beyrichites falciformis SMITH (19U. p.
116. pl. 91. figs. 11-13, type species).
which was described from the l\liddle
Triassic " Daonella dubia zone " of Fossil
Hill, ·west Humboldt Range, Nevada. in::---Jorth America, but the present specimen differs from the American species in
the umbilicus width and the width of
the shell.
Occurrence and geological horizon:Dark gray calcareous sandstone of the
Rifu Formation at Hamada, Shiogama
City, 1\liyagi·gun, l\liyagi Prefecture.
Northeast japan. Upper Anisian Paraceratites lrinodosus Zone?

Repository :-GLKU-C157 Coli. Y. BA:-;noReferences
BA:-JDo, Y. (1961): Biostratigraphical Correlation of the Lower and the Middle Triassic Formations by Ammonoid Fossils.
from Japan. }. Geol. Soc. japan. t•ol. 67,
no. 789, pp. 323-334 (in Japanese withEnglish abstract).
- - (l964a) : Study on the Triassic Ammo-.
noids and Stratigraphy of Japan-Part l :.
Lower Triassic. with a note on the bound-.
ary between the Permian and Triassic
Systems in Japan. j. Geo/. Soc. japan,
110/. 70. no. 825- - (196•1b): The Triassic Stratigraphy andAmmonite Fauna of japan. Sci. R11p.,
Tohoku U11i1'., 2nd. Su. (Ceo/.). z•ol. 36.
no. 1
DJEKER. C. (1915): Fossilium Catalogus. Pt ..
8. Cephalopoda triadica. pp. 1-369. Ber·lin.
HYATT. A. (1900): Cephalopoda in ZittelEastman·, Textbook of Palaeontology. pp.
502~604. London.
and S~IITll. ]. P. (1905): The Triassic
Cephalopod Genera of America. U. S,

Yuji BANDO

Explanation of Plate 49
Figs. 1, 2. Hemiprionites shimizui BA:-JDo, n. sp.
1a, b. Holot~•pe, IGPS, coli. cat. no. ·15158, xl. 5.
2a, b. Crushed outer whorl. paratype. GLKU-C158, X 1. 5.
Loc. Tao near Uonashi. Shirokawa·c.ho, Higashiuwa-gun, Ehime Prefecture. Tao formation,
upper Scythian.
Figs. 3a-c. Anasibirites sp.
Specimen, GLKU--C155, Figs. 3a. b, x 1; Fig. 3c, x 1. 3.
Loc. Same as figs. 1 and 2.
Fig. 4. l!emiprionites sp.
Specimen. GLKU-C154. x 1. 8.
Loc. Same as figs. 1 and 2.
Fig. 5. Beyric!Jites cf. rotellifarmis (;'1.1J.:EK)
External mold of outer whorl. GLKU-Cl53, x2.
Loc. Gorodani. Nagayasu. Kaminaka-cho. Nakagun, Tokushima Prefecture. Zohoin formation. Aniso-Ladinian.
Fig. 6. Paraceratites sp.
External mold. GLKU-Cl5l. x3j5.
Loc. Isatomae. Utazu-cho. Motoyoshi-gun, l\liyagi Prefecture. Isatomae formation. Anisian.
Fig. 7. . Piychites sp.
External mold, GLKU-Cl50, x l.l.
Loc Same as fig. 5.
Fig. 8. Balatonites cf. gottscllei (.MOJSJSO\'Jcs)
External mold. GLKU-C152. x3
Loc. Same as fig. 6.
Figs. 9a. b. Beyricltites sp.
Specimen. GLKU-Cl57, xl.
Loc. Hamada along the Senseki Electric Car Line, Shiogama City, Miyagi Prefecture. Rifu
formation, Aniso·Ladinian.
Figs. 10. 11. Paranannites sp.
lOa. b. Complete specimen. GLKU-C156. x2. I.
lla. b. Crushed specimen of outer whorl. GLKU-Cl59. x 2.
Loc. Same as figs. 1 and 2.
The illustrated specimen of fig. 1. is kept in the Institute of Geology and Paleontology,'Tohoku University. Sendai. Japan. The other specimens are preserved in the Department of-Geology, Kagawa University, Takamatsu. Japan.

RANDO:

Triassic Ammonites from Japan

Plate 49

~81.

Lower a.nd .1liddle Triassic Ammo-noids

Ceo/. Sun•. Prof. Paper -10. pp. 1-3~}1, pls.
1-85.
1-.:HIKAWA. K. (1951): Triassic System in
the Southern Part of the Kitakami
l\hssif. in the Triassic Stratigraphy ofJapan. Ceo[. Sun•. Japan, Rep. Spec.
Numb .. pp. 7-26 (in Japanese).
KIPARtsovA. L. D. (1961): Palaeontological
Basis of the Stratigraphy of the Triassic
Formations in Ussuriland. Part 1. Cephalopoda. Trudi. /Jses. Nauts.-Jssred.,
Ceo/. /lrst. (VSECEJ), 110. 48. pp. 1-278,
pls. 1-38 (in Russian).
KoBAYASHI. T. and lciiiK!I.WA. K. (19i>l):
Triassic Formations in the SakuradaniTenkaizan area in Tokushima Prefecture,in the Triassic Stratigraphy of Japan.
Ceo/. Sun·. japan, Rep. Spec. Numb .. pp.
89-95 (in Japanese).
MEEK, F. B. (1877): Palaeontology. ll. S.
Ceo/. Explor. 40th Parallel, CJol ..J. pt. 1.
pp. 99-129, pls. 10-11 (Trias).
:V1oJsiSOVICS. E. v. (1888): Ueber einige
japanische Triasfossilien. Beitr. Paliiont.
Oest.-Ung. u.d. Orients. Bd. 7. Heft 2.
pp. 163-178, pis. 1-4.
NAIC-\ZAWA, K. (1964): Discovery of theAnisian Fauna from Shikoku, Southwest
Japan and its Geological !\leaning. Alent.
Coli. Sci., Kyoto Unit•.. Ser. B, r•ol. 30,
no. •1. Ceo/. and Mineral .. Art. 2. pp. 7·
19, pls. 1-2.

Annomoto
Mt / Jt:
Fukkoshi~~
Gorodani
E.. Rli 1'1Hamada~fJI
Higa.;hiuwa-gun JR~fp.flf,
Hirai so
>¥
111
lnai
11: I
fill
lsatomaefJt JR rJiJ
Kaminaka-cho
J:JJlljltlllf
Minato~Miyagi-gun
.!l£m

s

3-15

S!lnii7.U, S. (1930): On Some Anisic Ammonites from the Hollandites Beds of theKitakami 1\lountainland. Sci. Rep .. Tohoku Imp. Unit•., 2nd. Ser. (Geol.), t·ol.
1-1, no. 1. pp. 63-74. pl. 24.
- - and ]I~IBO. N. (1933): On the Triassic.-\mmonites from Tao, Uonashi-mura(Shirokawa-cho). Higashiuwa-.;;un, Ehime
Prefecture. Chikyu (The Globe). vol. 19.
no. 1. pp. 10-31 (in Japanese).
S~IITII. ]. P. (191 J): The Middle Triassic
Invertebrate Fauna of North America.
U.S. Ceo!. Sun·. Prof. Paper. 83. pp. 1254, pis. 1-99.
SPATII, L. F. (1930): The Eotriassic Inver·
tcbrate Fauna of East Greenland. Med.
om Crnland. no. 83, pp. 1-90, pls. 1-12.
- - (1934) : The Ammonoidea of the Trias
1: Catalogue of the Fossil Cephalopoda
in the Briti3h Museum (Nat. Hist.). pt.
4. pp. 1-521, pl3. 1-18.
W.\AGEN. \V. (1895): Fos:;ils from the Cera·tite Formation. Pal. Indica. 13th Ser.
Salt Range Fossil,; 2, pp. 1-323. pls. 1-·10.
Yt\llE, II. and SIII:-.IIZt:. S. (1927): The Triassic Fauna of Rifu near Sendai. Sci. Rep ..
Toltoku Imp. Uui1"' .. 2nd Ser. (Geo/.), 1•ol.
11. no. 2. pp. 101-136. pis. 10-14.
YEII.\R.~. S. (1927):
The Lower Triassic
Cephalopod and Bivalve Fauna of Shi ·koku. japan. ]. Ceo/. Cl'ogr., z•ol. 5, pp.
135-172. pis. 13-17.

:'vlotoyoshi-gun
Nagayasu
Naka-gunOjika-gun
Rifu
Saragai
Shiroka wa-cho
Tao (Taho)
Uonashi
Utazu-cho
Zohoin

hnt
:R
'/z:.
WJ'nlt\u Jj[ tH:

*

flj
Hil

I{J

~ fojtlllf

ll.
lJJt Ill lllJ
JII
lJ~
I~
:i!J.

i!< ?* itX

PROCEEDINGS OF THE PALAEONTOLOGICAL SOCIETY
OF JAPAN

B*~~-~~-~~~~~~~r~~,·~
ill[fCJiiJ#Jft:t:itl'f.CO~.i'ai_ll.:.-:>l ·"'(COy:/ :f- ~ '7 ./...;!

1964 ~ 9 Jl 20 B (B) 11\li!i*''f:J!l!~/lf,t.:.:t<H ·"'tf,fHll! ~ *1. t.:

0

The Miocene Noroshi Flora of 1\oto, Japan.
. . . . . . . .. . .. . . . . . . . . . . . . . .. . Shiro lsiiJOA
On the Shell Wall of Triticites . ............ .
.......................... Kimiyoshi SADA
Re\·ision of the Takagami Conglomerate.
Ch6shi Peninsula. and Descriptions of thePermian Bryozoa from Its Limestone
Pebbles ................ Sumio SAKACA~·II
Bryozoa of Akiyoshi. Part L. Permian Bryo.
zoa from the Shigeyasu Quarry ......... .
........................ Sumio SAKAGA~Il
Ditto. Part 2. Lower Carboniferous Bryozoa
from the Uzura Quarry .. Sumio SAKM;A~llThree Carboniferous Species of Bryozoa
from Khao Noi. Central Thailand ....... .
. . . . . . . . . . . . . . . . . . . . . . . . Sumio SAKAGA~Il
On the Molluscan Fossils Collected from the
Ryiikyu Limestone of Okinoerabu Island.......................... Katsura 0YA~IA
Semifossils from the Silica Sand Deposits ofThui Tri. Vietnum ...... Katsura OYAMA
Disco\·ery of Marine Shells from the Kunohe
Tarrace Deposits. Kuji City, Iwate Pre.
fecture ................................. .
...... Kotara HAT AI and Rikizo li.IAIZLT:-.II.Monotis (E~ttomonotis) from Kyushu ....... .
. . . . . . . . . . . . . . . . . . . . . . . . Minoru T A~IVRA
On the Corbiculidae from the Lower Neo-com ian Ryoseki Series. japan ........... .
.......................... Yoshihisa 0TA
Nautilus pompilius Drift on the West Coastof Thailand .............. Ryuzo TOHIYA·
MA. Tadashi SATO and Takashi HA:\IAUA
Notes on the Drifted Nautilus in Thailand ..
. ....................... Takashi K-\:o.IADA
Yabeiceras (Cretaceous Ammonites) from
Fu taba, Northeast Japan ............... .
. . . . . . . . . . . . Tatsuro MATSUMOTO, lkUWO·
0BATA, Shiro MAEDA and Teruo SATO
On some Lower and :\fiddle Triassic Ammonoids from japan .......... Yuji BA:-;uoSome Cephalopods in the Permian Faunuleof Takakurayama. Fukushima Prefecture.
japan .................. lchiro HAYASAKA
Fossil Echinoids in the Museum of Sbirahama. \Vakayama Prefecture. Japan ..... .
........................ Akira MORISHITA
Preliminary Report of the Ordovician andSilurian Conodonts from the Langkawi
Island, Malaya ......................... .
Hisayoshi leo and To.;bio KOIKE~JijllliJ!fti.::.Mt r.,fil!COMI&: ... · ·. · ·. · .... Jllli1l'iftl~l~::- :/ :r. '/? J...: '/"' 7 · F!!II!~W!~ft:U$0)~.ifL,; , 7. s!ffi~O)wtu=tutco~.i!l! ...... ~*B ;.f,:J:'Jilli~=fiJJi. 7*-

rn* --r~o:.-:>~ .·c .... >t>!llf:it:~fi

8!Hi~:Cll!i!l~Jfi7:J·i.::. --:>~ ·"'( ........ W:;f;;.ffil~l~>P'!:ft711.z.~M7*--rr.:.-:>~·"'t .... tfflJ! ?...

SYSTIMATIC INDEX(New Series No. 49~No. 56)

PALA EOZOOLOG Y
Protozoa
Page
AOKI. N.: ;\liocene Foraminifera from the Kiyosumi Formation, Boso Peninsula .......... 163- - - : Noles on the Foraminiferal Genus Pseudocibicidoides . . . . . .
.........
195
Islllz.\Kl. K.: Verbeekininae from the Inferred Upper \Volfcampian Limestone in the Westof Ryoseki. Kochi Prefecture . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 51
Upper Carboniferous Fusulinids Similar to Fusulinel/a pseudobocki (LEE andCHE~) .... ''. ' ..... ' ' .... ' .. ' . .
. ............. ' ... ' .......... '' . .
102

Coelenterata
HAsHI~IoTo.

\\'. and NAG.-\o, S.: A Find of Spongiomorphoids from the Cretaceous System
of Rebun Island and the Esashi Group of lhe Esashi Mountains. Hokkaido....
271
IGo. H.: Diphyphyl/um from Itoshiro. Fukui Prefecture. Central Japan .
liO
MaRl, K.: Some Solitary Corals off Aomori Prefecture, Japan
309
SAKAr;t;cHI. S. and YA:-IAGI\1':\, N.: Additionai Notes on Late Paleozoic Corals found in
the southern Part of the Tamba District.. .. .. .. . . .. . . .. .. .. . . ..... . ..
9

Bryozoa
SAKAGA~II. S.: Lewer Carboniferous Bryozoa from the Omi Limestone, japan. Part 2
15- - - : Some Permian Rryozoa from the Yamanba Limestone in Sakawa Basin. Shi·koku. Japan.
.. .. . . . ... . . .. .... . .. . .. .. . .. . .. . .. . .. . . .. ..
155- - - : Bryozoa of Akiyoshi. Part 2. Lower Carboniferous Bryozoa from the Uzura
Quarry... . .. .... ... .
.....................
295

Brachiopoda
HAYASAK.-\. I.: Orthotichia japonica liAY.'\s.-u;A (1933) and Orthotichia magnifica GRABAU(1936) ... '.. ' .. ' .. ' ' .. ' ' ..... ' .... ' ' . . .
15·1
MoRI. K. · Brachiopods from the Late Jura~sic Nakanosawa Formation in Fukushima
Prefecture. Japan. .. . ... ... .. . . .. .. . . .
. . . . . . . . . -11:'1-lollusca
BA:-;no. Y.: A Dibranchiate Cephalopod from the Rifu Formation (Triassic) near Hamada, Shiogama City. l\liyagi Prefecture. Japan..........
46- - - : On some Lower and i\liddle Triassic Ammonoids from Japan...
332
leo. 11.: On the Occurrence of Gonialites (s. s.) from the IIida iVIassif. Central Jap::tn
23-1
h\'.>.sAKI. Y.: Pseudamiantis. a Pelecypod Genus..............
.........
91
Kt:ROIH. H. and UR \T.\. II.: Discovery of a Fossil Perotrochus in the i\liike Coal-field.

31~

Systematic lnde:t:

Kyushu. Japan ................ .
263:
S.: Some Jurassic Trigonids from the Tetori Group in the Kuzuryu District.
Central Japan................
... .. ... .. .. .. .. . . .. . ... . . .. .. . .. . .... .... .. .. .
1
Trigonioides from the Late Mesozoic Tetori Group. Central japan . . . . . . . . . . . . 79·
MAst;oA, K.: Remarks on some Fossil Dosinia of japan.................................. 29·- - - - : The So-called Patinopecten of Japan........... . . . . . . . . . . . . . . . . . . . . . . . .
1·15
MATOBA. Y.: A Cretaceous Trigonid from the Miocene Misaki Formation in the Miura
Peninsula. Kanagawa Prefecture . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
. ... 256·
MATSli!'.IOTo. T. and A\IA:->:0, 1\·1.: Notes on a Cretaceous Nautiloid from Kyushu ........ 173.
- - - - . 0BATA. I.. M.\[,;DA. S. and SATO. T.: Yabeiceras (Cretaceous Ammonites) from
Futaba. Northeast Japan. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . ......... 322
MuRAT:\, M.: Some Middle Permian Aviculopectinidae from the Kitakami Massif. Northeast Japan .......................................................................... 215·
NAGASAWA, J.: Significance of the Variation of Fossil Batillaria cumingi (CRossE) from
Quaternary Deposits of South Kanto. japan...............
115
l\ATORI, H.: Some Molluscan Fossils from the Tertiary Muro Group in the Kii Peninsula,
Japan...............
. . . . . . . . .. . . . .
. ............ 217
SAITO. Y.: A Rudistid from the Cretaceous Depo:;its of Ryoseki. Kochi Prefecture, Japan 317
SATO, T. et KANIE. Y.: Lilloetia sp. (Ammonite callovienne) de Mitarashi au Bassin deTetori.... .. . . .. .. .. .. .. ... .. . .. . . .. ..
..........
8·
SHUTO, T.: Naticid Gastropods from the Miyazaki Group. (Palaeontological Study of the
Miyazaki Group- X) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
. .......... 281
UFDA, Y.: Two New Species of Opis from the Cretaceous of japan...................... 70·
MAEtM.•

Arthropoda
Ihr, Chung-Hung: Some Lower Ordovician Trilobites from Flank lin Mountain;;, Texas . . 86·- - - : The Dimorphism anti Ontogenr of Norwuodella halli RESSER
12g.

Vertebrate
IL\TAI. K. and KoT''"''· T.:

New Species of Gadus from the Pliocene of Jat>an . . . . .

25-

PALAEOBOTANY
E:-.oo. S.: On the Genus Acer with Description of ~ew Species ........ .
65·- - - : On the Genus Platanus from Hokkaido, Japan ................... .
133
0YA:.L<\, T. and MATSUo. H.: :\otes on Palmaean Leaf from the (larai Flora (Upper
Cretaceous). 6arai machi. Ibaraki Prefecture. Japan ................................ 211
TAKAIIASIII. K.: Pollenformen aus den !Iitomaru-Schichiten in der Yuya.,Van Gegend
120
Microplankton from the Asagai Formation in the Joban Coal-field.....
201
TA~-:,\1. T. and SuzuKI. :\.: On the Genus Ailanthus from the Tertiary of Japan . . .
135·

MISCELLANEA
lGo. H. and KoiKE. T.: Carboniferous Conodonts from the Omi Limestone. Niigata Prefecture, Central Japan (Studies of Asian Conodonts. Part I) ........................ 179·
Mt'LLER. K.].: Disco\·ery of Conodonts and Horothurian Sclcrites in Japan .............. 29-1

INDEX OF F AM ILlES, GENERA AND SPECIES
Notes: Words listed below are names of families. genera and species, which are either
described or illustrated in the Transactions and Proceedings of the Palaeontological Societyof Japan. New Series. Nos. 49-56: words in heavy type arc names of new genera and species.

A
Acant hocladia? sp.
306
Acanthopecten cf. coloradoensis .......... 225
222- - onukii ...... .
225- - spinosus .................. .
Acer arcticum ..
66
67- - micranlhum
67- - proforufinerFe .............. .
Ailanthus altissima .... .
1H- - yezoense ...................... .
111
126
Alnipollenites eminens
337
Anasibirites SJ> ...
226
Annuliconcha ..
227- - kituknmie11RiR
Apon if eus . . .
. ........ .
87--tutus .......... .
~7212
Apteodinium sp.
47
AI ractites ..
48- - lwtai ...
At·iculopecten lwyasakui . . . . . . . . . . . . .
219
221- - hatuii ..................... .
218- - swwkii . .. .

B
Balalonites cf. go/tschei ............... . ;{{9206Ballisphaeridium (?) polyceratum
207- - sp . .\ ........ .
207- - sp. B ... .
208- - sp. C
118
Bat it/aria cun1ing1........ .
/Jeyrichites d. rotellijormis .. .
3·11
3-!3
- - s p .....
166
Bolirinila ![IWdrilalero ...... .
62
Hret•axina lrataii
166
Bulimina st,·ialuc"Callisla" (Pseudamiantis) crenulatus
Cassidulina cf. subglobosa ...

94
169? Cical ricosisporiles sp.
. ....... .
Cymatiosphaera g/ohulosu .......... .
- - reticulosa ...................... .

123
210

210

D
Delfocyathus orienta/is ........ .
Diphyphyllum cfr. delicutum
Dosinia akaisiana ...
(Piwcosoma) hataii .......... .
(--) japonica .............. .
(Kaneharaia) kaneharai ...... .
(--) karmoi ............. .
(Pharosoma) lugaruana

31-1

171
31
34
31

38
35.

32

E
Eoschubertella obscura"Eponides" umbonatus ..
Euprioniodina dentata

106
167
185.

F
Fissurina 1/lltllllla
Fistulipora sp. A .
- - sp. B ............. .
Flabellum I ran st•ersate- - - - conirum
Fortipectininae ........ .
Fusulina :;p ..... .
Fusu!ine/la aff. e/egantula- - jamesensi., ................. .
- - dr. rlwm/Joides

164
297
297
312
313

150
109
107
109
108

G
26
Gadus cltikagau·aensis ..... .
- - luscus ............................. . 28
- - macrocephalus . . .
. ......... .
28
27- - mnsm/ai ....... .
- - morshua ....................... .
28
Gnathodus opimus............... .
189

Index of Families. Genera. nnd Swcies
192- - sp.

236·Gonialites sp ......
89
Goniotelina cf. brighti
16i·Gyroidina komu/Hili ................ .
- - prof11nda ......................... . 16i

IIHayasakapora erectoradiala/lemiprionites t~himiz11i ...
- - sp ......... .
/Jeritschioides sp. . ............... .

\58

335
336
12

Heterocyalhus japonicus ............ .

313- - sp.

314······················

Hindeode/la ltHiulica......... .
- - parudelicaiula ...
- - t~akcrgamii . . . . . ........... .
Huangia sp. . ................... .
Hysl riclwsplweridiu 111 sp ............. .

lbotrigonia lelrwiensiH
ldiognat/wdus loyaslrii
lnaperturapol/enites pseudodu/Jius[soleloides fmnklinensis

183

13

209

·I

188
126
88

- - sp.

·······················

Ligo nodi na hmwii .. .
Ulloetia sp. . ........ .
Lonchodina? nipponica
Lunatia pal/ida ........ .

3
.J
157

299
209

.................. .
.............. .
................... .

206
206
206
206

? Microreticulatisporites sp ........ .
55
Mise/lin a a/iciae........... .
- - cylindrica ...
56- - aff. ibukiensis
59- - subelleiptica ..
61.llizuhopecten . ...
151- - yessoensis ......................... . 152? .\Jonocolpopollenites kyuslwensis ..... . 126
Monopieura sp .......................... . 219
Mugil cephalus ........ .
28
Myophorella (Promyophorella) orienta/is
6

183
184

L
Latilrigonia lwrii ....
- - kwwii ...
Leioclema nmwkae ...
--? IIZIII'fii'IIHiH
Leiofusa sp. . ........... .
Leiosphaeridia /aet•iuata .. .
- - cf. laerigaltl ........ .
- - rugulata .. ........... .

- - sp. A- - sp. H- - sp. C

NNeoplatycephalus macrodon ....... .
28· Nodosaria" longi.~wta
165
Non ion cf. pa1·kerae................. . 168
Nont·oodella haiti ............ .
129
hal!i 0 ..
. .............. .
130t:~o--haiti~..
. .......... .

0Upis (Opis) amakusensis- - (--) lwkkaidoenRis ...
01·thomurphina obitsuensis
Orthotichia japoniw .............. .
- - magnijica
Ozarkodina orienlnle

i3
7,1

165
154
15-1

187

211

p

21-!
211
212
186
8
186

Paraceralites sp.
3-10
Paracymatoceras
1i5- - tzmglraicum
175
Parajissurina kiyoRumienRiR ............. . 1&1
Paranannites sp. . .
. ................ . 338
Patino pecten caurinus ................ . 150- - healeyi .......................... . 152
Penniretepora cf. fragi/is ............... . 305
16- - lrigaslriyamensiH ................ .
16- - cf. irregularis .................. .
- - ofukuensiH.
30·1
17- - reuularis

288

28i
,\/ammila mtlltra ..
286- - melanostoma... . ........ .
157
Aleekopora? sp ......... .
28
Merluccius r•u/garis ........ .
Micrhystridium a:;agaienst' . . . . ........ . 203
205--fragile .......... .
203---minllH ... .
20,1- - misentm ........ .
204- - - (?) Hpinuliferum ..... .

- - sp.

18- - sp.

305
126

Periporopollenites asiaticus ........... .
- - porulosus .....
Perot roclws eocenicus
Pilar clarki

126
267
25--1

lndc:;: q( Famihe.'-1. (ifmera and Speeics- - hataii- - kotoi- - kyushuensis
Platanus aceroides yubariensis

250
252
25-!

133
13·1
Pleurostomelfa alternans ............... . 167- - sp.

Poli nices (Glossaulax) hagenoshitensis .. 28-1(--) h/III{Jensis ................. . 282(--) aff. reiniana ............... . 285
281(Nn•erita) sagamiensis ........ .
278
Polyphy/loseris? cf. iwalensis ......... .
............... . 15
Polypora fujilae
126
Polyporopollenite.- grandis
320
Praecaprotina yaegashii
89
Protopliomerella conlracla
18
l'rotoretepora hayasakae .......... .
Prolrilicites ........................... . 110
110- - nakahalenRiR
9:~l'seudamiantis
96- - pingui.~95- - tauyensis .............. .
~97
Pseudobatostomella cf. granulosa
198
Pseudocibicidoides
199- - katasensis ...
262
Plerotrigonia /lokkaidoana259- - cf. hokkaidoana262- - pocilliformis.
339
Ptychites sp.
88
Plyocephalus dclit·at us

88- - sp.

R
19
158- - sp.
··················
Nlwmbopora sp. . ...................... . 19
Rhabdomeson 11abei ..... ..... .

l~obulus pseudorutulat us........ . 16,1
Roundya suhacodus ..................... . 186s
Sabatites oortmiensis:243Saj]'ordotaxis JllliWgidae. . . . . . . . . . . . 299
Septopora tcaberi..................
303
Solemya (tlclzarax) dalli . . . . . . . . . . . .
254- - (--) muroensis . . . ............. 249
Spathognathodus echigoensis
187
Spongiomorpha asiatica...
278
Stenopora pusilimonila . . . . ............ :!98
Sli/ostomella ketienziensis
16oStreblascopora amabilis............
20

:351

--anliqua ......... .
:~o1- - grossa.....
. .............. .
21
St reblochundria millallloriensis229
Sl rcblot rype/la astrovae.
. ........ .
302- - ? sp .................. .
21
Streplognathodus expansus ..
189- - japonicus . . . . . . .
. ............. . 188
Sf romalomorpha Tl'llllllellsis ............. . 276
10Stylidophyllum kinkiense
SubtrifJOropo/leniles wnsimilis ..
126
Sulcorelepora complicafa ............... . 22
- - nipponica ......................... . 159
23- ? sp. ···············........ .
Synprioniodina collinsoni ........... .
185
181- - microden/aT
Tectonalica jant/wslomoides

289- - sp.
·························.. Terebrat uta" /wlaii.
.. - - " iLvaii

290
·12

· · - - " sp ................... .

·13

43

28
Theragra chalcogramma
166
Tostria hanzawai ...... .
Toucasia carina/a orienta/is .......... . 320
126
Tricolpopofleniles micro ref icula/us
126- - l'rtlgaris . . . . . . .
. .......... .
121
Tricolporopollenit es hilomaruensis
Trigonioides
80- - kifadmziensis .
83
81- - leforiensis ..
Tuberculalisporiles parl'ierinaceus ..... 126- - sp.

································ 126

126
Ulmipollenites undulosus
28
Uroplrycis braziliensis ...... .
Urigerina cf. akitaensis............ . 166
166- - proboscideavllaugonia kuzuryuensis.

w
Waagenophylhmr /ambenseIIy}'abeicerlls orientale
323
Yabepecfen .
...........
1-!9- - lokunagai............... 150

a -lt"'~±~~~-~-~~fTransactions and Proceedingsof thePalaeontological Society of Japan~ew

SeriesPalaeontological Society of japan

The heading in japanese commemorates the handwriting of Prof. M. YoKOYA~lA,
father of Japanese Palaeontology, who was Professor of Stratigraphy and Palaeontology at the Geological Institute, Imperial University of Tokyo.
The fossil on the front page is Helicoprion bessonowi KARPI:'\SKY, 1899.

CONTENTS

Number :19 (Published April 10, 1963)
Transactions
Article

442.
443.

444.
445.
446.

Page

Shiro l\IAEDA: Some jurassic Trigonids from the Tetori Groupin the Kuzuryu District, Central japan.......................... 1-7
Shigeo SAKAGUCHI and Nobuo YAMAGIWA: Additional Notes on
Late Paleozoic Corals found in the Southern Part of the Tamba
District . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 9-14
Sumio SAKAGAJ\11 : Lower Carboniferous Bryozoa from the Orni
Limestone, japan. Part 2 ...................................... 15-24
Kotora HATAI and Tamio KoTAKA: ~ew species of Gadus fromthe Pliocene of Japan .......................................... 25-28
Koichiro l\IASUDA: Remarks on Some Fossil Dosinia of Japan .. 29-38

Plat~

1

2
3-•1
5
6

Short Note10.

Tadashi SATO et Yasumitu KA:\IE: Lilloelia sp. (Ammonite call·ovienne) de tvhtarashi au Bassin de Tetori.................... . .
8Proceedings ...................... 39-·tO

Number 50 (Published June 10, 1963)
Transactions'1<17.
448.

4:19.
450.

451.

Kei MoiH: Brachiopods from the Late Jurassic Nakanosawa Formation in Fukushima Prefecture, japan ........................ ·11-45
Yuji BANDO: A Dibranchiate Cephalopod from the Rifu Formation (Triassic) near Hamada, Shiogama City, Miyagi Prefecture.
japan . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 46-50
Kunihiro ISHIZAKI: Verbeekininae from the Inferred Upper Wolfcampian Limestone in the West of Ryoseki. Kochi Prefecture .. 51-64
Seido EN no: On the Genus .·leer with Description of New Spe·cies ............................................................ 65-69
Yoshiro UEDA: Two New Species of Opis from the Cretaceousof japan ........................................................ 70-78

7

8
9
10

11

II

Number 51 (Published September 10, 1963)
Transactions

452.
453.
'15·1.
455.
456.

457.

Shiro MAEDA: Trigonioides from the Late ~lesozoic Tetori Group,
Central japan .................................................. 79-85
12
Chung-Hung Hu: Some Lower Ordovician Trilobites from Franklin Mountains. Texas ............................................ 86-90
13
Yasuhide IWASAKI: Pseudamiantis, a Pelecypod Genus ........ 91-101 14-15
Kunihiro ISIIIZAKI: Upper Carboniferous Fusulinids Similar to
Fusu/ine/La pseudobocki (LEE and CHEN) ........................ 102-114
16
joji NAGASAWA: Significance of the Variation of Fossil Hatillaria wmingi (CROSSE) from Quaternary Deposits of South Kanto,
Japan .......................................................... 115-119
17
Kiyoshi T AKAHASIII: Pollenformen aus den Hitomaru-Schichiten
in der Yuya-Wan Gegend................................. 120-127
18
Proceedings......................
128

Number 52 (Published December 10, 1963)
Transactions

458.
459.
c!60.
461.
462.

Chung-llung Hu: The Dimorphism and Ontogeny of Norwoodellaha/li RESSER .................................................... 129-132
19
Seido ENDO: On the Genus Platanus from Hokkaido, Japan .... 133-13-1
20
Toshimasa TANAI and Nobuo SUZUKI: On the Genus Ailanthusfrom the Tertiary of Japan .................................... 135-1,14
21
Koichiro rvlASUDA: The So-called Patinopecten of japan ........ 145-153 22-23
Sumio SAKAGAMI: Some Permian Bryozoa from the Yamanba
Limestone in Sakawa Basin, Shikoku, .Japan .................... 155-160
24
Short Note

11.

lchiro HA YASAKA: Orthotichia japonica H.-\ YASAKA !1933) andOrtlzotichia magnifica GRARAU (1936)............................
Proceedings . . . . . . . . . . . . . . . . . . . . . .

154
161

Number 53 (Published April 10, 1964)
Transactions

463.
461.

Naoaki AoKI: ~liocene Foraminifera from the Kiyosumi Formation, Boso Peninsula ............................................ 163-169
Hisayoshi !Go: Diphyphyl/um from ltoshiro, Fukui Prefecture,

25

iii

465.
466.

Central japan .................................................. 170-172
Tatsuro ~dATSll!\IOTO and \lasahisa A:\IA~o: Notes on a Cretaceous Nautiloid from Kyushu .................................. 173-178
26
Hisayoshi !Go and Toshio KOIKE: Carboniferous Conodonts fromthe Omi Limestone, Niigata Prefecture, Central japan (Studies
of Asian Conodonts, Part I) .................................... 179-193 27-28
19·1
Proceedings . . . . . . . . . . . . . . . . . . . . . .
Number 51 (Published june 30, 1961)
Transactions

467.
468.
469.
470.

Naoaki AOKI: 2'{otes on the Foraminiferal Genus Pseudocibicido29
ides ............................................................ 195-200
Kiyoshi TAKAHASlll: ~licroplankton from the Asagai Formationin the .Joban coal-field .......................................... 201-214 30-33~vlasafumi l\IURATA:
Some l\liddle Permian Aviculopectinidae
from the Kitakami ~lassif. Northeast Japan .................... 215-233 34-35
Hisayoshi IGo: On the Occurrence of Goniatitcs .s. s.· from the
Hida Massif, Central japan .................................... 234-238
239
Proceedings . . . . . . . . . . . . . . . . . . . . . .
Number :55 (Published September 20, 1964)
Transactions

471.

472.
473.
474.
,175.

·176.

Toshiji 0YA!\IA and Hidekuni ~viATSUO: Notes on Palmaean Leaffrom the Oarai Flora (Upper Cretaceous'. Oarai machi. lbaraki
Prefecture, japan .............................................. 211-246
1-liro'o NATOI<I: Some l\lolluscan Fossils from the Tertiary l\luro36
Group in the Kii Peninsula. japan .............................. 247-255
Yasumochi MATOHA: A Cretaceous Trigonid from the i\liocenc
37:'vlisaki Formation in the r--.liura Peninsula, Kanagawa Prefecture .. 256-262
Hidetaka KURODA and !I ideo URATA: Discovery of a Fossil
Perotrochus in the l'vliike Coal-field. Kyushu. japan .............. 263-270
38
Wataru HASHIJ\IOTO and Sutekazu 1\.o\GAO: A Find of Spongiomorphoids from the Cretaceous System of Rebun Island and theEsashi Group of the Esashi JVIountains. Hokkaido .............. 271-280 39-41
Tsugio SHllTO: Naticid Gastropods from the l'vliyazaki Group.
(Palaeontological Study of the Miyazaki Group-X) ............ 281-293 42-43
Short Note

12.

Kbus ]. MULLER:

Discovery of Conodonts and llorothurian

iv

Sclcrilcs in japan ............................................. .
Number 56 (Published December 20, 1964)
Transactions

477.
478.
479.
480.

481.

Sumio SAKAGAJ\II: Bryozoa of Akiyoshi. Part 2. Lower Carboniferous Bryozoa from the Uzura Quarry .................... 295-308
Kei rviORI: Some Solitary Corals off Aomori Prefecture. japan .. 309-316
Yasuji SAITO: A Rudistid from the Cretaceous Deposits of Ryoseki. Kochi Prefecture, Japan .................................. 317-321
Tatsuro iviATSU!\!OTO_. lkuwo 0BATA, Shiro ~viAEDA and Teruo
SA TO: }·ebeiceras (Cretaceous Ammonites) from Futaba, Northeast japan ...................................................... 322-331
Yuji BANDO: On some Lower and !vliddle Triassic Ammonoids
from japan .................................................... 332-3,15
Proceedings . . . . . . . . . . . . . . . . . . . . . .
346
Systematic Index ................ 3,17-3,18
Index of Families,
Genera and Species ........ 3·19-351

47

48

49

I
.'

•

Ill! II~ l1l!
--------

L965q:.~~~ • 9'·~

---~

----

I ~IJ-"LH?~~ftl"i
I

llll

flli

------.mitlt$~~WEI

1:1
----

1965 ~f- 1 J1 23. 2·1o1964 ~ 12 11 20 El<ID:ili'iA. Jo~~tr

pq.)

News

O tp; 3 l•Jitl4~tfWt.f~lilf~~J,\l (International Association for Quaternary Research Vll International Congress) ti 1965 ~ 8 11 30 El .t 1J 9 Jl 5 El rc..ht.: IJ 7-" 11 71 ~~114 ::z 1=2 7 r ffi0)7' :r'.7"7-crf-?,~.-~-"t'IHlill!;;n.:;,, .::O)~illQtc..f1~~ ·iWiliHH~~y;mjH~iY.J~fl'O)ftJ!. B-0~QO)~. 3
6 '00)-:/ :r ,f.:/? Al.l'·.iE:Jll!i ;;n -c:i<i iJ. 3l.!llf5'f-O)lilfrJe~{t >b IIHil·:tL.Q • .:: n~O)~-frO) 5""J0)5H-+~.

t, IS'~$b~K.reU~~I*~.:;, >b O)t.:.lt. ~~~!UcO)Jtl!~l¥1t.~:!J:Vt&O:/ff!Et.:.mJ-t .QJ}fH~-c. IS'~~.!: rtt·~~.J1l!JII!~ • 'ff!1ti!J~&o:~fl'ittJliJ4tO)iS'~#r~ IS'i!YJ4tit!!.fll!'lt · :ttw~r: c 71 'J ~ -7t.:. .1: .:;,~f~iR
Si!t.~: .!:.' ii':J.&f&iJ·:bh.Q.::}:: l.:.ts:..., -c:t.; IJ' ~~Jtlj~1;_f~O)Jri:~.!: ""'- 1) :/ p'0)~1H~K.!Ml-t 9-:/ :/ ;f- :/ rycA >bllM·tt9,< :bL< li.'fil'i!-".i!l!fli-t.Q.!: .t 1.',

Dr. Gerald M. RicliMO:-iD. Secretary General VII INQUA Congress, Building 25,
Denver Federal Center, Denver, Colorado, U.S. A.

1964 ~12J-jl5(31 9 6 4 ~ 12 J-j 20 13

a* i1~4t~41:¥tH'.r · ~*Tr J; m56-f}
40 0 Fl:!ti:Ji\:k~l.!l!~fl!Hl!!'t1t~~~F"J
D*i!i~4t~~
if:j

m

*

m

~

13

ii
<Wt ~ IJ ~)I!}( 84780 •>E!J .fliU ;ti Jt! .3?.: ~ ~ Iii z Jt rHiJ 2 / 13~l«irr<!ilr:tJBO~~~.!l: $II1
5[:

Regulations for Publication in Transactions and Proceedings of
the Palaeontological Society of Japan(Jan. 15, 1963)
1. Manuscripts considered for publication should have been read at the General Meeting or
the Ordinarp meeting of the Palaeontological Society of Japan.
2. Manuscripts shall be written in European language, they should be typed (Pica) on oneof standard-size (22.5x 27.5 em) paper and clouble.spaced throughout. Biological names
should be in italics and be undealined by the author.
3. Manuscripts (including of text-figu1·es, maps and tables) will be limited to 24 printed
pages (less than 54 type.written pages).
4. Illustrations will be limited to two plates (14.2x20.0 em).
5. Text-figures will be limited to 10 in number, not exceeding 60 tsubo in total area (1 tsubo
is 1 square sun). They should be drawn carefully on white paper with drawing or Indianink, letters used in the figures should be either printed or typed letters pasted-in. Figures
may be reduced, so authors are requested to carefully select the size and thickness of thelines or letter used.
6. Maps should be accompanied with scale, fractions should not be used.
7. The author is requested to pay for any cost extending beyond the above stated regulations.
In short manuscripts not exceeding 4 printed pages, the author is requested to pay thecost of all plates and of text-figures exceeding 2 in number or 12 tsubo in total area.
8. Manuscripts should have the title and a brief abstract in Japanese, (such will be addedfor persons not familiar with Japanese language).
9. Literature cited or referred to should be listed at the end of the manuscript in the formof bibliography. Bibliography should be arranged in alphabetic order of author and by.year. The order will be, Author, Year, Titleof Paper, Name of Journal, Volume, Page,
Plate, Figure, Map, Table.
10. The author's official address should be given below his name. under the title.
11. Palaeontological notes which can be fitted into less than one printed page (including
figures, maps, tables) will be published in the order recieved as space becomes available.
12. The desired number of reprints should be indicated on the right corner of the front pageof the manuscript. 100 reprints without cover, but will reference to volume, number andyear will be furnished free of charge to the author (if more than one author is involved
they shall be divided). Additional reprints will be furnished at the printers rate.

Editorial Regulations
l. The Editorial Staff will transact, preserve and edit the manuscripts.
2. When the Editorial Staff transacts a manuscript, a notification with date of acceptancewill be sent to the author, if the manuscript is clear, and abides with the regulations.
3. Acceptance or non-acceptance of manuscripts will be decided by the Edtorial Council.
4. Manuscripts not accepted for publication will be returned to the author will notificationfrom the Editor of the reason(s) for its rejection.
5. Manuscripts accepted will be published in the order received with the date of accetanceindicated thereon.
6. Manuscripts whose contents are altered by the author after being accepted for publication,
will have their date of acceptance changed.
7. Proof reading will be done under the responsiblity of the Publication Committee.