a -lt"' ~ i. :tW ~ ~~~-i~fTransactions and Proceedingsof thePalaeontological Society of Japan
New Series

No. 44

Palaeontological Society of Japan
December 15th, 19Gl

CONTENTS
TRANSACTIONS
Page

414.

Note on Steinmamzella (Yelzarella) aimwna (Y:\BE and NAGAO) ........... .

,115.

New Subgenus of Pearl Oyster, Fopinctada, from the Cretaceous l\•lifune•116.

A new Species of the Genus Clisiophyllum from the upper Carboniferousof Ehime Prefecture, Southwest japan ............................... .

. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . i'vtitsuo NAKANO

139

Group in Kumamoto Prefecture. japan ................ Minoru T Al\IURA 117. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Ken-ichi ISHII and Nobuo YA~IAGIWA

153

417.

On the Evolution of the Osmundaceae with Descriptions of two newSpecies .................................................... Seido ENDO

157

418.

On the new Genus Metadoliolina ........ Ken-ichi ISHII and Yasuo NOGA:\11

161

419.

Spinileberis, a new Genus of Ostracoda from the Pacific .... Tetsuro HANAI

167

~120.

On the Equid Fossil from Hiramaki Formation ......................... .

421.

. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Tokio SHIKAMA and Shinji YOSIIIDA

171

On some Triassic Pelecypods from Pahang Province, l\'lalaya ............. .
. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Akira ToKUY AMA

175

SHORT NOTES
7. Further Note on a fossil Palm Trunk from Kanazawa .... Yudzma OGURA 146
8. Amygdalophyllum sp. from Hyogo Prefecture, Japan .... Nobuo YA~IAGIWA 152
PROCEEDINGS . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 182

President : Teiichi KOBA YASH!
Councillors:

Kiyoshi ASANO (Editor of .. Fossils"). Riuji ENDO (Planning), Ilaruyoshi

FUJ 1:\IOTO, Shoshiro IIA:SZA

wA. Wataru I-IASIII:\IOTO (Treasurer), Kotora HAT AI,

khiro II.\YASAKA. Koichiro lCiliKAWA, Nobuo IKEI3E, Teiichi KOBAYASIII.
Tatsuro :'vlATSmiOTO, l\Iasao MI:\ATO. Tokio SHIKAMA (General Affairs), Fuyuji
TAKA! (Editor), Ryuzo TORIYA~IA
Secretary:

Tetsuro liAi\AI

Assistant Secretary:

Takeo ICHIKAWA

All Communications relating to this Journal should be addressed to the
PALAEONTOLOGICAL SOCIETY OF JAPAN
Geological Institute, Faculty of Science, University of Tokyo, japun

Trans. Proc. Palaeont. Soc. Japan, N. S .. No. 44, pp. 13~-145. pis. 20-21. Dec. 15, 1961

414.

NOTE 01\ STEINIVIANNELLA (YEHARELLA) AINUANA(YABE AKD NAGAO)*
l'viiTSUO NAKANOInstitute of Geology and Mineralogy, Faculty of Science.
Hiroshima University

Steimnamzella ( Yeharella) ainuana (YABE and NAGAO) 1;: -?\- ·"t" :~tw:i1I::::~m~~)ii]l1f2t7) <.H'!(t7)li¥!*t.:. -:>v·~r'l4#.1~8'~iilf3'Ei!:fi.-, t.:.., -'t(7)~ !;11:. :: (7).fl!!tJ:/l\':~:il!\lfl\l.:.:t31t Q

S:ft. . .t;;J:U'I!Mrf.:a£~,(7):Wl.-v<:. b~~f!Jq~u.:.(7)"t''fWeri-.::;,

<P

~

X Em

collected numerous specimens of Stein·
11zamzella ( Yeharella) ainuana from various places. As a result. it is recognizedthat St. ( Y.) ainuana represents aremarkable change of characters in
ontogeny and this species sho\VS a certain
degree of variation.
The materials dealt here with werecollected by T. MATSUMOTO and thewriter from the Ikushumbetsu district.
Central Hokkaido and stored in the
Geological Institute of Kyushu (GK) andHiroshima (GI-l) Universities.
The writer wishes to express his
sincere thanks to Prof. Sotoji IL'.IAMURAof Hiroshima University. His thanks
are due to Prof. Teiichi KonAYASIII of
the University of Tokyo, Prof. Tatsurol\IATSUMOTO and !vir. lkuwo OnATA of
Kyushu University, JVIessrs Tomowo OSE
and Ichiro HA YASHJ of the Sumitomo
Colliery Company, and Emer. Prof. Shoshiro I-IANZA\VA of Tohoku University
for their assistances.

Introduction and Acknowledgement

Steimnamzella CRICKMA Y, approximate
to STEJ:-\;I.JA:\:-.;'s Pseudoquadratae section.
inhabited in the Indo-Pacific region from
Upper Jurassic to Upper Cretaceous. It
was synonymized with Yaadia C!UCKl\IAY
by Cox (1952) and with TransitriRonia
DIETRICH by SAVELIEV (19581. but- 1\.oBA YASHI and A\L\:-\0 ( 1955) accepted its
validity and divided into :~ subgenera.
i.e. Steimnamzella s. s., Yelzarella KOBA·
YASH I and AMA:-\0, and Setotrigouia KoBAYASHI and AMANO.
Steinmamzel!a(Yelzarella) is well characterized bv its
outline and surface costation, and ~har­acteristic in the Northern Pacific region
possibly from upper Albian to l'vlaeslrichtian.
Trigonia ainuana is a typical
Steimnmmella ( Yelwrel/a). and has beenoriginally described by YAnE and NAGAO(1928) from the " TrigoJt.ia Sandstone"(Cenomanian-Turonian) of Pombetsu in
Ikushumbetsu, ~v!ikasa-city. Central Hokkaido. Recently, the writer had an
opportunity to study the Cretaceous
rocks of the lkushumbetsu district andFamily Trigoniidae LA;o..IARCK. 1819
Subfamily Quadratotrigoniinae* RccciYed Jan. 31. 1961 : read at 78th

SAVELIEV, 1958

meeting of the Society at Akita. May 13.

Genus Steimnamu>/la CRJCK~IAY, 1930

1961.

13!)

uo

JJitsuo NAK.J.NO

Type species:- Trigonia hoiubi KITCHIN,1913. Up. Neocomian; Natal, South
Africa.
Synonym:- Trrmsitrigonia DIETRICH,
1933.
Remark~:- The writer here accepts
KOBAYASHI and Al\IANo's proposal. This
genus varies to a large extent in surface
sculpture and shell form. and comprises
3 sub~enera and a number of species andvarieties as listed by them (1955) andNAKA:\0 (1960).
Trigonia malon.eana
STOYANOW from the Albian formationsof Arizona and Texas belongs probably
to Steinmannel!a s. s.
Subgenus }'elzarelta KOBAYASHI
and A:'vL\NO, 1955

Type species:- Trigonia japonica YE·
1923. Campanian: Sanuki and!yo, japan.
!Jiagnosis:-Shell large in size, subquadrate to triangularly ovate : umbo
improminent: beak opisthogyrous andpointed anteriorly: escutcheon depressed. with transverse costellae which areeffaced later : area ornamented withtuberculate costellae at least near umbo
but in the later stage costellae become
obsolete; carinae absent but sometimes
a row of nodes aligned in place of
three carinae in the vicinity of umbo:
median furrow shallovv· and rather
indistinct. separates a broad anteal partfrom a narrO\v posteal: flank withtuberculate costae or rows of nodes
which are disposed subconcentrically or
diagonally.
Test thick; growth-lines somewhat
distinct on the whole surface.
Remar/~s :-i\s pointed out already by
KOBAYASHI and A:'IIANO. this subgenus
is easily distinguishable from the othersof .'5/eimnanne/la by the effacement of
the transverse costellae and the carinae
liARA,

on the area. The specific Iist of thesubgenus was given by them '1955) andNAKA!'\0 (1960).
The shell belonging to this subgenus
shows a remarkable morphic change
through growth.
The immature shell of this subgenus
is subcircular to ovate in outline andquite similar to that of Myophorella s. s.
in surface costation. The beak is anteromesial to subcentral. Its area is provided
with transverse to oblique costae. anddistinctly bordered by three rows ofnodes. ?vledian furrow is shallow but
distinct. Costae on the Oank are tuberculate or broken into rows of nodes.
In the later stages. the shell becomes
gradually trigonally ovate or subquadratein shape and the beak is shifted anteally.
The carinae and sculpture on the area
arc obsolete, and the median furrow is
generally indistinct. The flank is ornamented with numerous tuberculate costae
or rows of nodes which are disposed
subconcentrically or diagonally.
Growth-lines are somewhat distinct on
the whole surface through growth, especially in the posterior part of the laterstages. Internally. weak radial plications
run along the positions of the carinae
and ventral margin is smooth. As pointed out by KOBA YASIII and AMA:\0 (1955,
p. 200), in some internal moulds of theright valve (pl. 20, figs. 3 and 5) it isalso well observed that several l{rooves
and ridges behind the posterior adductor
scar and they are oblique to themargin.
The shell outline and the surface costation are fairly variable in this subgenus.
The outline is subquadrate in ainuana,
kimurai and others, but subtrigonal in_fitclzi. japonica and its variety obsoleta
etc. Leana has a subcircular shell. but
some forms of ainuana arc somewhat
trapezoidal. Costae on the t1ank are414.

Note on Steinmamzel!a I Yelzarella) aimuma ( YABE and N.-t(;.to)arranged subconcentrically in japmzica
and its ally. while they are diagonal incommon forms. Costation on the areaand escutcheon are indistinct in mostothers. but the sculpture is fairly distinct in some forms (pl. 21. fig. I) of
aiuuana. Nodose carinae developed injaponica and kimurai etc .. while theyarc obscure in most common forms. A
median furrow is often distinct. but indistinct in common forms. PACKARD's
Ieana lpl. 5. figs. I and 5) has a wide
area. almost as large as the flank. but
in most others it is nearly as wide as athird of the flank.
Distribution :-Upper Albian t?l to tvlaestrichtian in the Northern Pacific region.
As already discussed by the writer I 1958!.
in japan this subgenus appeared probabl~'in the Cenomanian of the Yezo geosynclinal region including Hokkaido andSachalin district, and flourished mainlyin the Upper Cretaceous of Southwest
japan.
Steinman nella ( Yelzarella) aiuuaua(YABE and NAGAO)
Pl. 20. Figs. 1-6; Pl. 21, Figs. 1-6.

Trigonia ainuana YABE and NACAO,
Sci. Rep. Tolwku Imp. Unit•.. 2nd Ser ..
Vol. 9. No. 3. p. 84. pl. 16, ligs. :20. 20a.
1955. Steimtzamwfla ( Yelzarel/a) aiuuana KoB.-\\'ASHI and A~tA:-;o,Japan. Jour.
1928.

1957.

Geol. Geo~-:r .. Vol. 27. :Vos. 3--J, p. 204.
Trigouia cfr. aimwna lKEG;\~11 andO~tOR t.

four. Hokkaido Gakugei Uuit•.,Sect. B. Vol. 8, :Vo. 1, pl. 3, ligs. la-c.

Alaterial :-llolotype (YARE and NAGAO's original specimen) from Pomhetsu
in lkushumbetsu, ivlikasa-city. Central
Hokkaido. The fairly well preserved
specimens which the writer has examined
are as follows :In !\L NAKA:\'O's collection. from l\1.

Ul

NAKA:>:o's horizon U,, (see 1960. p. 223.
tab. 1) of the l11oceramus lwbetsensis zone
in the upper " Trigonia Sandstone" atan old site of a quarry near the Katsurazawa-dam in lkushumbetsu. !\likasa-city(GH. Nl'vl. 1081-1095, 1097 and 1099). GH.
NM. 1096 and 1098 were collected from
~1. NAKANO's Le horizon (see 1960. p.
224. tab. 21 of the Calycocems zone 1,?)in the lower " Trigonia Sandstone"
along the Pombetsu ri·ver in lkushumbetsu, Mikasa-city.
In T. ~IATSU:O.IOTO's collection. GK. H.
6046, 60n and 6053a-b \vere collectedfrom the upper .. Trigonia Sandstone"
at T. rvL\TSU:I!OTO's Joe. IK-2015 along
the Pombetsu river in lkushumbetsu,
tvlikasa-city and GK. B. 6054 was collectedby K. SATO from the Inoceramus hobetsensis zone of the upper ·• Trigonia Sandstone" at the point !Fl of !38.60 m. in
No. 1 tunnel of the Katsurazawa-dam
in lkushumbetsu. Besides them thewriter observed numerous unregisteredmaterials from the same locality, collected by T. MATSU~VIOTO.
Description :-Shell large in size, ovately subtrapezoidal to trigonally ovate,
inequilateral. broader than high andgently convex from umbo to venter andfrom anterior to posterior but obtuse
carina! angulation developed in theboundary between the flank and thearea: antero-dorsal margin short andnearly straight or slightly convex, forming an angle of 95 to 115 degrees againstwith postero-dorsal margin ; antero\rentral rounded and transmitting gradually into broadly arched ventral ; posterodorsal nearly straight or a little convex,
about a half as long as shell; siphonal
margin gently curved and well rounded
but sometimes subangulated at junction
with dorsal or Yentral margin. Umbo
low and rather improminent : beak opisLhogyrous and pointed at about a fifthMitsuo NAK.I:YO142

to three-tenths from the front. Flank
with about 17 nodose costae or rows ofnodes which are subvertical near thearea and become thicker towards ventral
periphery where nodes are somewhat
elongated along growth-lines: umbonal
about 2 tuberculate costae concentric tosubconcenlric: succeeding some 10 nodose
costae or rows of nodes more or lessbroadly spaced and curved diagonally :last 5 or so rows of nodes. slightly
currvecl. oblique forward and arranged
diagonally. Carinae obscure, but in theearly stage a ro\\. of nodes aligned inplace of the carinae. Area smooth ex·
cept for the umbonal region with numer·
ous tuberculate costae. almost as large
as a third of the flank and distinguish·

able from the flank by an abrupt changeof curvature and absence of nodes.
!\Iedian furrow well observed in thevicinity of umbo where the area isdivided into a broader anteal and anarrower postea) part by it. but in thelater stage it becomes obscure. Escutcheon narrow. somewhat depressed, provided with numerous slightly tuberculate
costellae which are evanescent later.
Growth-lines somewhat distinct onwhole surface. especially in posterior ..
Internally. weak radial plications run
along the positions of carinae and very
thickened at the ventral periphery:
ventral margin smooth. Test thick andits thickness measures about 5 mm. incommon adult forms.

1Heasuremeuts in 111111.
Specimen
GH. :'\M. 1081
GH. I\!'.1. 1082
GIL :\::\1. 1084
GH. Nl\1. 1085
GH. :'\l\1. 1093
GH. Nl\1. 1094
GH NIVI. 1099

GK H. 605-l

Val\·eRight
Right
Right (internal)
Left (internal)
Left (internal)
Right
Left
Bivalved

Length!Ieight

L/11

/6. 0

68. ()
65.2
48.0
51. 2

1. 12
1. 14
l. 21

7-1.2

58. 3
65. 4
14.5
72. 1

62. 5
83. ·1

64.8
62.9

55. 5
68. 0

1.1-!
1. 15
114
1. 12
1. 23

Explanation of Plate 20(All natural size)
Steinmmmel!a (Ye!Jarella) ainuaua (Y.·\BE and NAGAO) .................................. p. 1-ll
Figs. la-b. Bivalved specimen (GK. H. 6054). Adult stag~. Lac. point of 138.60 m. in
No. 1 tunnel of the Katsurazawa-dam, Ikushumbctsu, .Mikasa-city. Central Hok·kaido.
Fig. 2. Modeling cast of an imperfect right ,·alve (GK. H. 6053b), showing surface
character of the area. Middle stage. Lac. T. MATst:~IOTO's loc. IK 2015 along
the Pombetsu ri,·er in Ikushumbctsu.
Fig. 3. Internal mould of a right valve (GK. H. 6047). Early adult stage. Loc. ditto.
Fig. -1. Internal mould of a right ,·alve (GH. Ni\1. 1084), showing internal character.
:\Iiddle stage. Loc. an old site of a quarry near the Katsurazawa-dam in Iku·
shumbetsu.
Fig. 5. Imperfect left valve (GH. NM. 1085). showing surface costation and coarse
growth-lines. Middle stage. Loc. ditto .
.Fig. 6. lmpcrfect right valve (GIL Nl\1. 1081a). showing internal character in the immature stage. Adult stage. Loc. ditto.

NAEANO :Steinrnannella ( Yeharella) ain11anq

flat(! 2(}414. Note on Steimn:mnella ( Yeharella) ainuana (Y-!Bf.' and N.J.GAO)
Remarks:- The ontogenetic change isremarkable in this species.
In the early stage, the shell i L=2535 mm., H=23-32 mm.l is circular to
subcircular in outline and the beaklocated at about two· fifths from thefront but sometimes subcentral. The
height-length proportion ranges I : 1.03
to l: l.lO. The surface costation of thisstage is quite similar to that of ,1Jyo·
p/wre!la s. s. Flank is sculptured with
about 11, moderately spaced. nodose
costae or rows of nodes which are arcu·
ate diagonally. Carinae are absent. buta row of nodes is aligned in place of thethree carinae. Area is as large as athird of the flank. and is ornamented
with numerous tuberculate costae which
are first transverse but oblique later.
Escutcheon is narrow. and pro\·ided withnumerous slightly tuberculate costellae.
~ledian furrow is shallow but rather
distinct (see pl. 21. fig. 6).
In the succeeding middle stage. 50 60
mm. in length and 4:-i 50 mm. in height.
the shell becomes ovate to triangularly
ovate in shape and the beak is shifted
at about three-tenths to a third fromthe anterior end. The height·length
proportion is about l : 1.20. There are
12 to J:l diagonal and arcuate nodose
costae or rows of nodes on the flank.
Carinae and sculptures on the area andescutcheon become obsolete. but thecoarse growth-lines arc well developedon the area.
Shell (L=70-85 mm.. 11=63-70 mm.)in adult is ovately subtrapezoid to trigonally ovate in outline. and ornamented
with 15 to 19 nodose costae or rows ofnodes on the ftank. The height-length
proportion is 1:1.10 to 1:1.25. and thebeak situated at about a fifth to a fourthfrom the anterior extremity.
In some internal moulds of the rightvalve (pl. 20, figs. 3 and 5), it is observed

H :~that several grooves and ridges behindthe posterior adductor scar are obliqueto the margin. This feature is sometimes
well observed in those of Steimnannella( Yelwrel/a) japonica ( 't'EIIARA) and someforms of the Quadratae. Pseudoquadratae
and other sections.
Variation :-As mentioned above. thisspecies has a certain degree of variation.
For example. the outline and surface
sculpture are not always the same evenin specimens of similar size. There is arelatively trigonally ovate form (pl. 20.
figs. la-b: pl. 21. fig. 21. while the holo·
type and lKEGA~II and O~!ORfs specimen!1957. pl. 3, figs. la-c) have the sub·trapezoidal outline. Nodose costae orrows of nodes are somewhat narrowly
disposed in the illustrated specimens on
fig. 4 in pl. 20 and on fig. 3. in pl. 21. butthey arc broadly spaced in the others
lpl. 20. figs. 1-2; pl. 21. figs. 1-2 and 5).
The rows of nodes on the ftank are wellde\'clopcd in common specimens (pl. 20.
figs. 1-2; pl. 21, figs. 2-3 and 5). On
the other hand, some specimens (pl. 20.
figs. 5·6; pl. 21. fig. l l have nodose
costae on the flank. Costation on thearea and the escutcheon are usually limiteel to the earliest stage. but rarely
obserYcd in the middle stage of somespecimen 1pl. 21. fig. 41.
Comparison :-This form is easily distinguishable from the others of Stein·mannel/a ( Yelwrel!a) by its subtrapezoidal
to triangularly ovate outline and surface
costation. As compared with Trigrmia
trausitoria STE!:\~!A~:>: from the Neocomian of South America. this form hasa smooth area and more prominent nodes
on the flank. Tri.gonia lwlubi KITCHIN
from the Neocomian of South Africa issimilar to this form. but differs by theprominent umbo and the less numerous
costae on the ftank. Trif.{onia Ieana GABB
and T. Ieana var. lcliiteavesi PACKAJW

14-1

from the Chico group of the West Coastin North America differ from this formin the more quadrate outline and thecarinae and sculpture on the area are
more obsolete. Steinmamzella ( }·ehare/laJ
jimboi KonAY ASHI and A\IAl\0 from the" Trigonia Sandstone ·· on the Pombetsu
river in. the Yubari coal-field of Central
Hokkaido has some resemblance, to somespecimens of the early adult stage ofthis species, but the former is represent·
eel by a single internal mould. In theoutline and the aspect of the area. thisspecies looks like St. ( Y.) llimurai (TOI<U·NAGA and SHIMIZU) and its ally in Honshu
from Coniacian to Companian, but the
tatters have numerous remarkable nodes
on the more arcuate and geniculate cost·ation and the area is more broader.
Occurrence :-Abundant in the lower
and the upper ·· Trigonia Sandstone"
from the Ikushumbetsu district. l'vlikasa
city, Central Hokkaido. Its range isprobably lower Cenomanian to middle
Turonian or IO\\·est upper Turonian.
Comparable specimens were reported
from the same group in the '{ubari
coal-field in Central Hokkaido. This
species is commonly associated with anumber of Ammonoids and Pelecypodsas listed by IKEGAl\11 and 0.\IORI (1957),
NAKAl\O (1960), and others.
Referencesi\xnERSON, F. M. (1958); Upper Gretaceousof the Pacific Coast. Geol. Soc. Amer.,Mem. I 1. pp. 1-378, pls. 1-75.
Cox. L. R. (1952): Notes on the Trigoniidae,
with Outlines of a Classification of the
Family. Proc. Jlal. Soc. London, Vol. 29.
Pts. 2-3, pp. 45-70. pls. 3-4.
CRICI'~IAY. C. H. (1930); Fossils from Harison
Lake Area. British Columbia. Canada
Dept. 1Uines Nat. Mus. Catrada Bull. No.
63. Geol. Ser. No. 51, pp. 31-66, pis. 1-23.
- - (1932) ; Contributions towards a Monograph of the Trigoniidae. I. Amer. jour.
Sci .. Vol. 24. No. 144. pp. 443-464. pis. 1-2.
DIETRICH. W. 0. (1933); Zur Stratigraphie
und Palaeontologie der Tendaguruschi·chten. Palemztographica, Supp/. 7, Ser. 2.
l\'o. 2. pp. 1-86, pls. 1-12.
lKEl.AMI. S. and T. 0:-.1ou1 (1957); On theso-called "Mikasa Formation" in the
Katsurazawa-dam site area near the
Ikushumbetsu River. Mikasa. Hokkaido.
jour. Hokkaido Gakugei Uuil'. Sect. B. Vol.
B. c'io. 1. pp. 70--89, pls. 1-14 (in Japanese).
KITCH IX. F. L. (1913); The Invertebrate

Explanation of Plate 21(All natural size)

Steinmamzella (Yeharel/a) ailma11a (YABE and NAGAO) .................................. p. lH
Fig. 1. Imperfect left Yalve (GH. NM. 1083). showing sculpture on the flank. Early
adult stage. Loc. an old site of a quarry near the Katsurazawa-dam in Ikushumbetsu. Mikasa-city, Central Hokkaido.
Fig. 2. Gypsum cast of a right Yalvc (GH. NM. 1082). Adult stage. Loc. ditto.
Fig. 3. Gypsum cast of an imperfect left val\·e (GH. NM. 1091). Adult stage. Loc.
ditto.
Fig. 4. Modeling cast of an imperfect right ,-aln: (GH. :\M. 1092a), showing the sculpture on the area and escutcheon. Adult stage. Loc. ditto.
Fig. 5. Gypsum cast of an imperfect right valve (Gll. NM. 1089). Adult stage. Loc.
ditto.
Fig. 6. Modeling cast of an imperfect left va!Yc (GK. H. 6046). Immature stage. Loc.
1\L>\TSl'~IOTO's Joe. 2015 along the Pombctsu rin~r in lkushumbetsu.

NA!(t1NO : Slf!fnmannel{a ( Yehar~lla) ainuanq

Plate 415

414.

Note on Steinmannella ( Yeharella) ainuana ( Y.4BE mul NAGAO)

Fauna and Palaeontological Relations of
the Uitenhagc Series. Ann. S. AfricanMus., Vol. 7, pp. 21-250, pis. 2-11.
KoBAYASHI, T. and M. A:-1.4.:-.;o (1955); On
the Pseudoquadratae Trigonians, Stein·
mmmefla. in the Indo·Pacilic Province.
japan. four. Geol. Geogr .. Vol. 27. /llos.
3-4, pp. 193-208. pis. 13-15.
MATSU~IOTO, T. (1959): Zonation of theUpper Cretaceous in Japan. i11em. Fac.
Sci., Kyus/m Unit>., Sm·. D. Vol. 9, l'lo. 2.
pp. 55-93, pis. 6-11.
NAI<Al'\0, M. (1958): On Some Upper Cretaceous Steinmannella ( Yelzarella) in Japan.
]our. Sci. Hiroshima Univ .. Ser. C. Vol. 2,
No. 2, pp. 83-88, pis. 13-14.
- - (1960) ; Stratigraphic Occurrences of theCretaceous Trigoniids in the JapaneseIslands and their Faunal Significances.
Ibid .. Vol. 3. No. 2. pp. 215-280, pis. 23-30.
PACKARD. E. L. (1921); The Trigoniae fromthe Pacific Coast of North America.
Oregon Unil•. Pub/., Vol. 1. No. 9. pp. 159. pis. 1-11.
SA \'ELIEV. A. A. (1958) : Lower Cretaceous
Trigoniids from Manghyschlack andU5\\.estern Turkmen. Leningrad. pp. 1-386.
pis. 1-58.
STEil'\:'-.IA:'>il'\. G. (1882): Die Gruppe der
Trigo nat: pseudo·q uadratae. 1\lcues ]altrb ..
Beii-Bd. 1, pp. 219-227. pis. 7-9.
SToY A:"OW. A. (1Y49) : Lower Cretaceous
Stratigraphy in Southern Arizona. Geol.
Soc. ,-.mer., Mem. 38. pp. l-156, pis. 8-26.
ToKUl'\AGA, S. and S. SHI:'-.IIZU (1926): The
Cretaceous Formation of Futaba in iwaki
and its Fossils. jour. Fac. Sci., Imp. Uuit•.
Tokyo, Sect. 2. Vol. I. Pt. 6. pp. 181-212.
pis. 21-27.
'{ABE. H. and T. NAGAO (1928): Cretaceous
Fossils from Hokkaido; Annelida, Gastropoda and Lamellibranchiata. Sci. Rep.
Tolwku Imp. Univ .. 2nd Ser., Vol. 9. No.
3. pp. 77-96. pis. 16-17
Y"HARA. S. (1923a); Cretaceous Trigoniae
from Amakusa Islands. Prov. Higo,
Kyushu. Japan. ]our. Geo!. Soc. Tokyo.
Vol. 30, pp. 1-12, pis. 4-7.
- - (1923b): Cretaceous Trigoniae fromSouthwestern Japan. ]apm1. ]our. Geo!.
Geogr .. Vol. 2, No. 3, pp. 59-84. pis. 8-13.

Trans. Proc. Palaeont. Soc. Japan. N. S .. No. H, p. 146. Dec. 15. 1961

HORT NOTE7.

FURTHER ~OTE ON A FOSSIL PAL::\1 TRUNK FROM KAJ\AZAWA*YUDZURU OGURA
Botanical Institute. University of Tokyo

The writer described in 1952 a fossil
palm trunk. situated in Kenroku Parkat Kanazawa as an ornamental stone,
called "Take·ne-ishi ". as Pa!moxy!onMaedae (0CURA, 1952). Though thisfossil was worth while in fine preservation of tissues of trunk and roots, it
was regretful that its habitat was unknown. A few years later. the writer
received a fragment of a fossil palm
trunk. found at a river running Kanazawa. Its tissue was very similar to. but
a little different from the former species,
and he described it in 1955 as Palmoxrlon
kagacnse (0GURA, 1955). This specimen
might have been drifted from the stratum. the green tuff of the Miocene age,where other fossil plants have been found.
Meanwhile another specimen of a fossil

Fi.~;.

1.

sists of a basal part of the trunk andthe root mass surrounding it, the diameter of the trunk being 35 em, andthat of the root mass 55 em (Fig. 1).
The preserYation is not very fine, but
it is enough to observe the tissue. Distribution and construction of vascularbundles. structure of sclerenchymatous
sheaths surrounding bundles (Fig. 2),
porosity of fundamental tissue, andstructure of small sclerenchymatous
strands in fundamental tissue are thesame with those of Take-ne-ishi, andthe writer intends to identify this new
specimen as Palmoxylon Maedae.
The discovery of this fossil palm isinteresting. as it suggests the habitatof Take-ne-ishi, that is. it might havebeen found near Kanazawa and may be

Fig. 2. Cross section of a vascular bundle
and is sclcrcnchmatous sheath. x35.

A new specimen of- Palmo:rylon Jt.faedae.

of a Miocene origin.

palm trunk was found at the suburbs ofKanazawa among the conglomerate, perhaps corning from the same stratum
mentioned above.** This specimen con-

ReferencesY. 1952: A fossil palm in Kenroku
Park at Kanazawa. Trans. ]'roc. Pafaeont.

Oct'RA,* Received July 2 , 1961.
** ~.if~m~~~ll!f (lB'ft-E'IIJJ!jtioUtll':/'l'JIIt-t

Soc. fapml. 8, 223-230.
- - , 1955: A fossil palm trunk from Kanazawa. ibid. 19. 85-87~) 7;:.i!l*ll.GA:1*~ (1960), :~1Et':"fl9.di!!.l'~YHf.

l·Hi

Trans. Proc. Palaeont. Soc. Japan, N.S., No. 44, pp. 147-151, pl. 22, Dec. 15, 191il-115.

NEW SUBGENUS OF PEARL OYSTER, EOPINCTADA,
FROM THE CRETACEOUS MIFUNE GROUP
I:\ KU~IAMOTO PREFECTURE, JAPAN*
MINORU TA!viURA
Faculty of Education. Kumamoto University~4;:~(1)Et~R!~~.~~!8€GT>

Pinctada (1)fjjj£~: ~!fdii!L.J:~';;j;:&U:~~illTJIIF'-;Ul llr1..'
matsumotoi TA:-.tt.:RA (~Jl!!bJi · Jfj\ft) ~ff::•
1,.. t:, *tlf!lil~tUi¥.1. -Pinctada maxima (jAi\IESO:")-i:.mli'UJt~~k't'tl: t.i!i:i't:.: hinge
structure :±:@: -IZ-t '5, J...t.i' l Y'l-tf~n::==:,ffjffl"(', {1!!(1)~ '5/i. '(,~~ 1,.. "C Eopinclada ill~~
tdlt~lt.:, C!IAVA~ I.:J::-'5 Pine/ada crassa ~Jl:f';< c Pine/ada lii.lt*<P~J~J:: IJii*H','~.n "ttJI?'f. :n:n~[iil(1):/"" 7~~M'In·•":i Pteric ~~&d-7.>-:. ~:. ~m~~-="\~fi.q)f!Mn:~z
n:lJ!I-rll~t.;.-:. 1: a:f!Ht~ ;t '5 c. ;{>;f!l!Q)w!±li± Pinctada Cl),f,mJ:JJilllt~ .:r>fH£1: .\l.'.l:l:h7.>,X01ll~i!Mlff'Fffllf~(1)~fi'l.!.l\t.!Jt1: J...'t'ii-tQ)jfgi±HtEJ:: IJ-001-:. brackish (1)!Jl'~tc!?.;,n.
i.:~;t c,.:tt. "'t'< Ln:. ;$:00(1)tf!Jifi~#JJGT>~·ft'f!l\'t!H:R::I-:~ <x~t' 7.> t<l) c .IJ!.?,(1)13lJN\~~r·f!!imi!E Pi11ctada (Eopinctada)

<,

<

III

The occurrences of pearl oyster. Pinr·!ada, from l he Terliary are exceedingly
rare and none from the Cretaceous inso·
far as the writer is aware, although
Pinctada aassa. which seems for thewriter to belong to another genus, hasbeen reported from Astartian of Calvados(CHAVAl\, 1952). In his fossil hunting,
the writer obtained many pearl oysters
from a sandstone bed in the lower for·mation of l\tiddle Cretaceous Mifune
group (\IATSU~IOTO. 1939) at Kami·
Umeki, \lifunc town and east of Kawa·
uchida, \lashiki town, both in Kumamoto
Prefecture. The lower formation of thegroup yields a rich pelecypod fauna of
which many species are coexistent with
the Goshonoura group (i\IATSlJ:MOTO,
1936). The fauna was listed by \lATSU·~IOTO (1939) but has not yet been des·cri bed except for 11Iat sumotoa japunica----------t-t~1958).
In this reporl, the writer establishedEopinctada. a new subgenus of Pindadn,
on Pinctada (Eopindada) ma/sumotoi T A·
\IURA, new species, from till' lower for·mation. Some considerations are madeas to the depositional environment of
the formation.
llere the writer expresses his sincere
gratitudes to Prof. T. KoBAYASHI of
the University of Tokyo for his constantguidance and supervision of the manuscript and to Mr. Y. TAKE\IURA for his
useful information on recent pearl
oysters.

OKADA (OKADA,

Family Pteriidae
Genus Pinctada ROEDINI;, 1798

Eopinctada TA:\IllRA, new subgenus
Type-species:- Pine/ada (Fopinrtada)matsumotoi TAMURA, new species.
Diagnosis :-Shell rather small for* Received J'vlay 17, 1961: read at 79th
meeting of l he l;Ociety at 1\anazawa, Sept.
23, 1961.
1-17

!vEnom TAivJURtl

148

Fig. 1.

Index map

A. a: Kiyama
b: Kawauchida
F: Fukuoka
h: Hetami
K: Kumamotok : Kamiumcki
I: Yuen-linc
M. m: Mifune
N: Nagasaki
r: Mifune river
y: Yokono
x : fossil locolity

Table l.

Comparison among Piltclada s. str .. Eopinctata and Pteria (Magnat•icula)

Pine/ada

genera~--Pinctada ;.~r.
Eopi;,c-ta_d_a__characters '~-- _ _ _ _ _ _ _ _ _ _ __I

- -1 Pteria (J\Iagnat•icu/a)

obliquely trig<:mal,antenor margm notproduced

obliquely trigonal
but posterior wingprominent

shape

quadrate, anterior
I margin produced

umbo

not terminal

i terminal

not terminal

ligament area

fairly wide

wide

narrow

I-~---------1------------~·---- _d_e_e_p-ly- impressed,
not dee~ly impressed. crescentic but nicked
crescentic
medially

adductor
impression

not deeply impressed.
roundedly crescentic-1-

non-nacreous
bordersurface1

wide

not present

widerugose or scaly but
not prominent inolder form

probably smoothnearly smooth

415.

JVew Subgenus of Pearl Oyster. Eopinctada

genus, depressed. nearly equivalve. in·equilateral. obliquely trigonal in outline
and higher than long. anterior and pos·terior parts not alate; umbo prosogyrate
and nearly terminal ; surface smooth;hinge area wide. obliquely divided into
three parts (anterior, median groove,
posterior) ; byssal notch fairly deep
especially in right valve; one great
crescentic adductor scar strongly im·pressed; test fairly thick and nacrewide.
Remarks :-In regard to shape of ad·
ductor scar and the oblique shell form.
Eopinctada resembles Pteria (Magnavi·cula) of which hinge structure is similar
to that of Eopinctada. but the posterior
extremity is not alate and the umbo
terminal in the former.
In the hinge structure Eopinctada
perfectly a'{rees with Pinctada s. str.
(TAKE:'.IlJRA and KAFtJKll. 1957). althoughA

Fig. 2.

B

c

Interior of Pteria and Pinctada(left valve)

A : Pteria (.\1agum·icufa) penguin(Roi~DI:-;G)

B:

X!

Pine/ada (Eopinctada) ma/sumotoi
TA~Il'llA X~

C:

Pindada maxima (J.>.~IESO~) x t

its area is wider in the former. The
non-alate posterior part of the shellagrees with that of Pinel ada. The left
valve of the l'teriidae is generally moreconvex than the right but the difference
between them is not perceived in Eo·
pine! ada as in the old shell of some

Pinctada. It is said that the non-nacreous
border of the old form of some l'inctada
as P. maxima anct P. margarit(fera is
broken and the surface is not so scaly
in younger shell.
The non-nacreous
border is not present in Eopi11clada andsurface nearly smooth. The thick test
in hinge part of Eopinctada is seen also
in old Pinctada. Fopiuctada has many
characteristics seen in old Pinctada.
The shape of the shell is a little differ·ent from that of Pinctada. The impres·sions of pallial muscles are not observable distinctly but somewhat resemble
those of Pinctada.
Pinclada martensii (DUI\I<ER) is a well·known pearl oyster and lives in theshallow sea of west japan. The pearl
culture is prosperous in Japan. The
occurrence of Pinctada, however, has
never been reported from lvlesozoic for·mation and is very rare in Tertiary.
The only one exception is Pinctada
cmssa by CHAVAN (1952) from Astartianof Calvados. His species, however. hasa prominent posterior wing and thespecimen illustrated in the plate 2 lacksof posterior part of shell. This species
seems for me to belong rather to Pteria,
probably occupying ncar position to
Jl1agnavicula. than Piuctada. But fossil
Pteria is common in the jurassic (H.A·
YAMI, 1958; TA:\IURA, 1960). Fopinctada
well agrees in many characters with
Pi11ctada s. str. and in a few others with
Pteria. Whether Pine/ada s. str. wasderived directly from either Eopinctada
or Pteria (iHagnavicu/a) is a matter ofdiscussion, but Eopinctada is most akinto Pine/ada s. str. especially Pinctada

maxima (J Al\IESON).
Distribution:- The type species occurs
in the i'vlifune group (approximately
Cenomanio·Turonian) of Central Kyushu.

150

Minoru T4!\!URA
Pinctada (Eopinctada) matsumotoi
Ti\:VIURA. new species
Pl. 22, Figs. 1-4

Description :-Sheli small for genus,
depressed; equivah·e exclusive of convexity and byssal notch. inequilatcral.
higher than long and trigonal in outline: right valve a little more convex
than left: hinge margin nearly straightand not alate posteriorly: posterior
margin nearly straight or slightly arcuate: anterior rounded but profoundly
excavated below umbo; umbo prosogyrate. terminal: surface smooth : hingearea very wide, divided into three obliquely arranged parts in which themedian part is a pit or ligament groove:
adductor impression large and deep.
situated near center. crescentic in shapeembracing large impression of retractor:
several impressions of pallial muscles
not distinct: nacre wide and non-nacreous
border probably absenl.
:lfeasurement :Holotypc (right. Ml'vl 3704)
Paratypc (left. lVB1 3705)
Paratypc (right. M.l\•1 3706)

Length
55 mm·15 mm
45 mm

Height
68 mm
nO mm
50 mm

Observation:- This species is represented by many specimens which display
all characters of both valves almost
perfectly. The byssal notch is slightly
deeper in the right valve than theother. The shell convexity is stronger
in right than in left valve. No other
difference is found between both valves.
The lower part adjacent to byssal notch
is not so strongly produced as in Pim:tadas. str. l\lorphic variation among many
individuals of this species is small.
It has been thought that the lower
formation of the Cretaceous l\lifune
group is a sediments in an embayment
under shallow marine water as c~n bejudged from stratigraphic and palaeontological evidences. No ammonite hasever been found and trigoniae andInocerami arc very rare in the formation.
IviATSUMOTO reported some clements ofbrackish inhabitants (lvlATSU:\IOTO. 1939 ;OKADA, 1960)_ Recent Pinctada is known
to exist in warm and pure marine environment. The occurrence of Pinctada(Eopindada) matsumotoi from the lower
formation of t\lifune group is an interesting fact not only in thinking thephylogeny of Pinctada bul also in thinking the environment of deposition of

Explanation of Plate 22
Pindada (Eopi11dada) malsumofoi T A:\IURA. new species

Fig. 1. Plaster cast of the internal mould of a !cit valve; showing hinge area. (.\IM3707).
Fig. 2. A broken left valve: showing exterior. (J\E\I 3708).
Fig. 3. Plaster cast of the internal mould of the holotype right valve: showing internalside. (Mi'vl 3704).
Fig. 4. Plaster cast of the internal mould of a left valve; showing interior. (MM 3705).
All are in natural size and collected at Kami-Umeki. Mifune town, Kumamoto Prefectureand stored in the Geological Institute. Universitr of Tokyo.
Pinctada maxima (JA~lESO:>:)

Figs. 5, 6.

Interior of left valve; storea in Geol.

Jnst. Univ. Tokyo; x 1/3.

T AMUI?A, :c;retaceous fear! Oyster

Plate 22

415.

New Subgenus of Pearl Oyster. Eopinctada

the lower formation of the group. Further
an echinoid fossil is reported to occur inthe formation (OKADA. 1958). So the"brackish clements ·• of the pelecypod
fauna of the formation arc necessary to
restudy more strictly.
Cumparisrm :-The wide hinge area.
non-alate posterior part and shell struc·ture without non-nacreous border arecharacteristic of the species and nocomparable species of .. Pteria" has beenreported insofar as the writer is aware.
Pindada maxima (Jameson). however.
is most akin to the species and thecomparison between Pinrlada (Eopilll:·tadai matsumotoi and P. maxima wasdiscussed already.
Orcurrence :-Sandstone (fossil bed 0.305 m thick) on the roadside of KamiUmeki. through which the road extends
from l'vlifune to (\lizukoshi.
At thislocality. specimens of this species are
crowded.
Another locality is on theroad side. cast of Kawauchida. Mashiki
town.
Several thin (0.1-0.3 m thick)\ayers of sandstone and shale yield
numerous small pelecypods. Only two
fragments of this species were obtained
there.

li)l

References cited
CnAY.-\:-i. A. (1952). Les Pelecypodes dessables astartiens de Cordebugle (Calvados). ilfem. Suisses Palaeouto .. Vol. 6.9.
HAYA:-.11. I. (1958). Liassic Volsella. Mytifus
and some other Dysodont species inJapan. Tra11s. Proc. Palaenl. Soc. japan,:V. S .. No. 29.
MATSll~JOTO, T.
(1938). Geology of the
Goshora Island. Amakusa. with Special
Reference to the Cretaceous Stratigraphy.
four. Geol. Soc. japan. Vol. 45. No. 532.
- - (1939), Geology of the 1\lifune district.
Kumamoto Prefecture.
Ibid.. ~ 'ol. 46.
No. 544.
OKADA. H. (1958),' JHatsumoloa: A new Priono·
dont Pelecypod Genus from the Cretaceous Mifune Group. Kyushu. Japan.
Jlem. Fac. Sci.. l(ruslm Unit•.. Ser. D.
Geolo![y, Vol. 8. No. 2.
- - (1960). Sandstones of the Cretaceous
1\lifune Group. Kyushu. Japan.
;llem.
Fac. Sci.. Kyus!tu Uuit·.. Ser. D. Geology.
Vol. 10. No. 1.
TAKDJt'RA, Y. and KAFGI<l:. T. (1957).
Anatomy of the Silver-Lip Oyster
Pi11clada maxima (Jameson).
Tokaiku
Fishery ExPerimental Station. A . .Vo. 44.
TA:'-.JUHA. M. (1960). Gpper Jurassic Ptcriacea
from the Soma Group. Fukushima Prefecture. Japan. Trans. Pror. Palamut.
Sol'. japan. N.S .. cVo. 31.

Trans. Proc. Palaeont. Soc. Japan, N. S., No. '1·!. p. 152. 0-:-c. 15, 1\ltil

SHORT NO; E
8.

AMYGDALOPHYLLUM SP. FROJ\1 HYOGO PREFECTURE. JAPAN*NOBUO YAMAGIWA
Osaka University of Liberal Arts and Education

The present writer has discovered,lm.vgdalophyllum sp. from a crystalline
limestone lens within a schalstein mem·ber at ~Iannotoge. Kozuki-cho. Sayogun. Hyogo Prefecture. Japan. This
locality lies at the southwestern part of
the Tamba terrain. and near the \tai·zuru zone. Gon> 0960) recently studied
the Palaeozoic formations in this district.
and discovered the Lower & !vliddle
Permian.**.·lmygda!ophyllum sp. resembles A.. nao·soidea ~viiNATO in many respects. The
latter one was alreadly reported by~11:-IATO & KATO (1957) from the lower
Upper Carboniferous of the Akiyoshi
district. Besides. A. cfr. naosoidea wasmore rccc•ntly discovered by the writer(unpublished) from the zones of Profusulinel/a-Fusulinella o[ the Atetsu district. Therefore, the rock containing
the present form may be lower or middle
Upper Carboniferous in age.
Acknowledgements are due to Drs. S.
SAKAGUCIII & K. NAKAZAWA for thecritical reading of the manuscript, to:\tr. K. OsAFLT:"E. the collector of thisspecimen.

major septa and as many for minor
ones. Major ones rhophaloid in shape,
most of them reach the axial structure.
:-.linor ones thinner and shorter thanthe major. Sometimes major and minor
septa of continuous type in distal part.
:'-Jo external wall and peripheral part ofsepta is seen. Axial structure oval inshape and solid. Longitudinal section
unobservable.·lmyl(da/ophyllum sp. indet.

GoTo, H. (1960) : The Mesozoic & P;.r laeozoic
Formations in the Harima district. Hyogochigaku, no. 9.
J\IJ:-;ATO. :\1. (1955): japanese Carboniferous
and Permian Corals. four. Fac. Sci. Hokkaido Unil·., ser. 4, 1'0/. 9, no. 2.
MI="ATO. M. & K.no. M. (1957): On the

Text·fig. 1

Corallum simple. Calicural diametermore than 13 mm in transverse section.
Septa of two orders, major and minor,
alternating. 1\umbering about 41 for* R;;cci vcd Sept. ·1. 1961.

** These formations lie southwest of thePermbn formations in the Tamba district
studied by SA".\(;UCIII (1961), and are in nocontact with the latter. because of ha,·ing
igneous rocks between them.

Text-lig. 1. .·lmygda/opltyllum sp. imlet.
Tramwersc sect ion ( x 1.8)
Heg. no. 61001.

Comparison: This form is related toAm)'l(da!op!tyl!um naosoidea :\IINATO (1955)
in many respects, but differs from thelatter in having numerous septa.

References

Carboniferous Coral Zones in the Akiyoshi
Plateau, Southwest japan. PI'(Jc. Japan
Acad., z·ol. 33.
SAKAGUCHI, S. (1961): On the Palaeozoic
Formations "Tamba Croup" dc-.•elopedin the South Tamba District. Jour. Geo!.
Soc. ]apa11, t•ol. 67, 110. 190.

Trans. Proc Palacont. Soc. Japan, N. S., No. 44. pp. 153-156. pl. 23. Dec. 15, 1961

HG.

A ~EW SPECIES OF THE GENUS CLISIOPHYLLUM
FROM THE UPPER CARBONIFEROUS OF EHIMEPREFECTURE, SOUTHWEST JAPAN*
KEN-ICI-Il ISHII & NOBCO YNviAGIWA
Osaka City University & Osaka Gakugei University·~~!li!-J::ffi'Em.~@.cn Clisiophytlum \1Jl(l)-jf1lfj: ·~~l]jl,J:If':i':fJlm,!T.HlU!Jiftil! i::tJll!.~J::
l'!1:~9!-*<P$t;lfmi1Jj~R!Jllf&v<Fn.lt-H&lllilllit!!l!t((l)'!iZJ&JII!Mlll<Pn· G Clisiophyllum ~(1)-Jflli:
C. chimense ~~iH!.. ffc~L. C. au·a N.,'{} C. atcJ atetsue11se ,!:(1)~~~.:.-:;>t,•'Cilit:.t:,'6 # ~ - . Jll ~ ~~ :*:The species described in this article
was collected by K. lSIIII from a reddish
limestone lens, intercalating tuff breccia.
within the Ryuzen group at Omodani.
Kurosegawa village. lligashiuwa-gun,
Ehimc Prefecture. Southwest Japan andfrom a limestone within the ltadorigawa
group at ltadorigawa of the same village.
The Ryuzen group was recently studied
by NAKAGAWA, SUYAHI. lCIIIKAWA, lSllll& YATIIASHITA (1959).
According tothem. this group is composed of basic
tuffs and volcanic rocks. intercalating
black mudstone and limestone lenses.
Fusulinid fossils such as Fusuline/la fiuxa(LEE et CHEx). Fusuline/la cfr. pseudobocki !LEE et CHEN) and Fusulinella spp.
were discovered by them from the limestone lens of this group at Omodani.
One of the present writers (ISHll. 1956,
1958. 1961) recently studied in detail the
Itadorigawa group. According to him.
this group is compo~ed of limestones,
intercalating mudstones and sandstones.
Also he divided this group into three~~------* Received June 1, 1961; read at the 78th
meeting of the Society at Akita, ~lay 13,
1961.

fossil zones (lt 1 lte and lt 3 ) on the basisof fusulinid fossils: the lt 1 fossil zone
is characterized by Fusulillella itadorigazcensis Is~m (l\lS) and Fusuliuella simp!icata TORIYAl\IA, the It" fossil zone by
Fusulina kamnerai ISHII. Fusulina shikolmensis ISHII. etc., and the lt 3 fossil zone by
Fusuline/la bocki MOLLER. One specimenof the coral species described in thispaper. was discovered from the lt 2 fossil
zone.
judging from the above-stated associations of fusulinids, the present writersconsider that the rocks containing thepresent coral species are the middle
Upper Carboniferous in age.
The present coral species closely resembles Clisiop/iyllum mea C'vliNATO)from the Upper Carboniferous of several
districts in Southwest japan: namely,
che Fusulillella zone of the Omi district~~111\A TO. 1955). the Clisiophyllum awa
zone of the Akiyoshi district (l\l!NATO& KATO, 1957). the zone of Nagatophyllum
fauna of the Taisyaku district (i\lE\ATO,
1951; YoKOYAMA, 1957. 19;}9) and thelowermost Upper Carboniferous of the
Atetsu district CfAl\IAGIWA. (unpublishr:·~
l ·lu

15-1

Keu-ichi Ismi and Nobuo Y.-lMAGITVAed))* It is also related to Clisiop/zylhtm
mea atetsuense (lVli.i'\ATO et N AKAZA w,\)from the lowermost Upper Carboniferousof the Atetsu district (1\II:>;ATO & NAK."'·
ZAWA, 195i: YAl\IAGIWA (unpublished)).
Systematic description
Genus Clisiop/zyllum DA:>;A. 18c16**

Clisiophyllum ehimense sp. nov.
Plate I. Figures 1. 4. 5

Corallum simple. large in size. Cali·
cular diameter usually 20-22 mm intransverse section. External wall rela·
tively thin. Dissepiments mostly ar·ranged concentrically or angulo-concent·rically. Septa of two orders. major andminor, alternating. 1\Iajor ones usually
32-38 in number. In mature stage, the;·reach near the axial structure, somewhat
rhopaloid except in outer area \vhere
they are more or less sinuous.
Inmature stage, minor ones more or lesssinuous, relatively thin and usually 1/22/3 the length of the major. However,the former usually 2/3-3/4 the lengthof the major in full mature stage. Cardi·
nal septum slightly shorter than othermajor ones. Axial structure composedof a short median plate, less numerous
septal lamellae and convex subvesicles.
concentrically arranged with com·ex
sides facing outwards.
In longitudinal section, broad peri·* 0KJ~JCHA (1958) considered the age ofthe rocks containg Clisiophytlum au:a (M17'A·TO) and Clisiophyllum azca alelsuense (i'VlJKAToet NAKAZAWA) from the Atetsu district.
Southwest Japan to be Lower Carboniferous(:\'amurian).
** Uisiupllyflum DA:"A, 1846 [ =Ciisaxophy!lum
GHABAt.: in CHI. 1931] (See WELLS & HJt.L(MooRE, ed.) l!l56. F. :!86).

pheral area composed of about 8 rows
on nearly equal vesicles, their convex
sides facing inwards as well as upwards
or inwards. Tabulae incomplete, widely
spaced and sloped up to the axial stru~­
ture. their convex sides facing upwards
and outwards. Tabellae show dome-likestructure. Tabularium occupies 3/5 entire
space of the corallite. Axial structure
occupies 2(5 the width of tabularium.
Materia/.·-Holotype. Reg. no. PC 2000a.
b. Another specimen, PC 2001. These
specimens are deposited in the Divisionof Geoscience. Fac. Sci .. Osaka City Univ.
Ocrunence ·-The holotype occurs fromthe Ryuzen group of Kurosegawa·\'illagc,
and another specimen occurs from theIt" fossil zone of the ltadoriga\\·a groupat ltadorigawa. Kurosegawa·village, Higashiuwa-gun. Ehime Prefecture.
Comparison :-This new species isclosely related to Clisiophy/lum au:a
( !-II."<ATO, 1951. p. 5, figs. cjl-d/3: 1955.
pp. 137, 138, pl. 5, fig. 1' pl. 17. fig. 7,
pl. 33, fig. 9. pl. 36. fig. 8. pl. 37. fig. 11.
text-fig. 13. figs. c/1-d/3: SAKAGUCHI &Y AMAGIWA. 1958. pp. 168. 169, pl. 1, figs.
1-31 in many respects. It differs, ho\vever, from the latter (Pl. 23. fig. 2) inhaving septal lamellae and a less prominent median plate in axial structurein transverse section. It resembles Cli.~io­
p/iy!lum mea alelsuense (1\lrNATO & NAKA·
ZAWA. 1957, p. 18-20, pl. 3, figs. 3. 4) inmany respects. but differs from thelatter (Pl. 23, fig. 3) in having septal
lamellae and a less prominent medianplate in axial structure in transYerse
section and no trabecular-like ridge inaxial structure. It is related to somespecies of the genus Dilmnop/iyllum inhaving less numerous septal lamellae inaxial structure in trans\·erse section,
but differs from the latter in having ashort median plate and longer minor
septa.

416.

A New Species of the Genus Clisiophylium

Acknowledgements
The \\Titers wish to express theirhearty thanks to Assist. Prof. Koichiro
ICHIKAWA (Osaka City Univ.) and Assist.
Prof. Shigeo SAKAGUCHI (Osaka Gakugei
Univ.) for effective counsel and readingof the manuscript. Thanks are also due
to Assist. Prof. Keiji NAKAZAWA (Kyoto
Univ.) who lent them a part of lheholotype specimen of Clisiop!zyllum awa
atetsuense C\II~:\TO et NAKAZAWA) for
the sake of comparison.

ReferencesK. (l!J56):
Itadoriguwa group in
western Shikoku. -,On some problem
conc,;rning the :Vliddle Pennsylvanian ofJapan-, jour. Geol. Soc. Japan. Pol. 62.
1w. 724. pp. 20-29. [In Japanese withEnglish resume].
- - (1958): Fusulinids from the Middle
Cpper Carboniferpus ltadorigawa Group
in '\Vestern Shikoku. Japan, Part I. Genus
Fusulina. four. Ins!. Polytec. Osaka City
Unir., ser. G. m/. .J. pp. 1-28. pls. 1-5.
- - (1961): Fusulinids from the Middle
Upper Carboniferous Itadorigawa Groupin Western Shikoku, Japan. Part Ill.
Stratigraphy and Concluding Remarks.
Ditto .. 1'0/. 5. no. 1. pp. 31-52.
Mtr-.;ATO. l\1.. (1951) : Some Carboniferous
Corals from Southwestern Japan, Trans.
Proc. P..:it:eont. Soc. japan. N. S .. no. 1. pp.
1-5.
- - (195S): japanese Carboniferous andPermian Corals. jour. Fac. Sci. Holdwido
Unit: .. ser..f. 1'0!. 9. 110. 2. pp. 1-197. pis.
1-43.
- - & M. K.no (1957) : On the Carboni·f;:rous Coral Zones in the Akiyoshi
Plateau. Southwest japan, Proc. japan
Acad.. rot. 38. 110. 9, pp. 547-552.
- - & K. X."\1(,\lAWA (1957): Two Carboni·ferous Corals from Okayama Prefecture,
Trans. Proc. Palaeo11t. Soc. Japan, N. S.,

ls1111.

1551)0. 25. pp. 17-20. pl. 3.
MooRE, R. C.. Editor (1956) : Treatise on
Invertebrate Paleontology. Part F. Coelenterata. Gent. Soc. America & Utzit•.
Kmtsas Press, F. 1-498.
NAt<ACA\\'A, C .. K. StJYARI. K. lctttt->AWA, K.
Isttti & N. YAi\tASHJTA (1959): Geologyof Kurosegawa District. Ehime Prefec.
turc. (Studies on the Chichibu-Terrainin Shikoku. IV). four. Gakugei. Tokushima
Ut~it• .. Nat. Sci .. 1'0/. 9. pp. 33-58. [InJapanese with English resume).
0KJMl'RA. Y. (1958) : Biostratigrayhical and
Palaeontological Studies on the Endothy·roid Foraminifer<~ from the Atetsu lime·stone Plateau. Okayamu Prefecture.
japan. four. Sci. Himshima lJniz• .. ser. C.
1'01. 2. no. 3. pp. 235-264, pls. 32-36.
SAKAGUCHI, S. & N. YA:\IAGIWA (1958): Thelate Palaeozoic Corals from the southernpart of the Tanba district, Jllem. Osaka
Unit!, Lib. Art. & Educ., B. Nat. Sci., uo.
7. pp. 163-178. pis. 1-5.
YoKOYA:\tA. T. (1957): ~ote on some Carboniferous Corals from Taisyaku district.
Hiroshima Prefecture, Japan. jour. Sci.
Hiroshima Unit• .. ser. C. mi. 2. 110. 1. pp.
73-82. pls. 10-12.
- - (1959) : The Geology of the Taisyaku·
kyo district. publication of lliroshima-ken.
pp. 29-·12 pls. 12. 13. [In Japanese).

Postscript: More recently, K,\:-::<.IERA re·ported a new species of the genus Clisiophyllum (C. subramosum) from the middle
lipper Carboniferous formation (the Reedeina[=" Fusu!im1 "] zone) of the Yayamadakelimestone of Kyushu (KA-.).!ERA 1961. Upper
Carboniferous Corals from the Yayamadake
Limestone, Kyushu. JUem. Fac. Sci .. Kyuslm
U11il' .. sa. D. Geol. 1•o/. 10. no. 2. pp. 211-214.
pl. 14. figs. 1-12. pl. 15. Jig. 11. text-fig. 3).
The present new species resembles C.
subnmzosum KA:->;o.tERA in general features.
especially in the axial structure and themode of dissepments. But the former canbe distinguished from the latter in having
a simple corallum and dome-like axial tabellae in longitudinal section.

156

Ken-ichi IsHII and Nobuo YrL1HGIWA.

Explanation of Plate 23
Figs. 1, 4, 5. Clisiophyllum ehimmse ISHII ct Y A~IA<iJW.\ (sp. nov.)
1. Transverse section in full mature stage. Reg. no. PC 2001. Loc. a limestone lens of
Jtadorigawa. Kurosegawa·villagc. Higashiuwa-gun, Ehime Prefecture .... lt 2 fossil zoneof the Itadorigawa group. x4.
4. Transverse section in mature stage. Holotype (Reg. no. PC 2000a), Loc. Omodani,
Yusukawa of Kurosegawa-village. Higashiuwa-gun, Ehimc Prefecture .... Ryuzen
group. x.t.
5. Longitudinal (somewhat oblique) section in mature stage. llolotype (Reg. no. 2000b),
Loc. Ibid. X 4.
Fig. 2. Axial structure in transverse section of Clisiophyllum awa (MI:-;ATO). 1951, (after
MI:>;ATO. 1951, p. <1, fig. Cl)
Fig. 3. Axial structure in transverse section of the holotype of C/isiophyllum awa atetsuense
MI:>;ATO et NAKAZAWA), 1957, (M. & N. 1957. pl. 3, figs. 3, 4).
This transverse
section (Reg. no. JPC-40029) is a part of the holotype, and deposited in Geol. &Min. Inst .. Univ. Kyoto. X4.2.

[ SHII and Y AMAGIW.tt:

A New Species of Ctisiophyllum

Plate 233

Trans. Proc. Palaeont. Soc. Japan, :\. S .. :\o. 44. pp. 157-160. pl. 24. Dec. 15. 1961

-HI.

ON THE EVOLT..JTIO:\ OF THE 'OS1viC);DACEAE\VTTil DESCRIPTIONS OF TWO ~EW SPECIES"'SEIDO ENDO

r..: ~, """1 J HCI)i~it::. 0~ --:-. zz.v: 2 ~fiil(/);JCJl.!: M~w;:.rr:--n~ft(1)rl~lltH.- t.. r:,.-::;:,;.::i r -1:!/-d J f4(/)ffl17./iPl'!l.l.f:j. <. li')!!l~t.J:tl·M(1)----:n:;t;,7::>:;, Jlt:f-f(/)j\Cftfr.lli;J'I;~li'i(1){:j~'l'f5f:;F!#f:.tTt.dfit!!H: IJ: t.. "C 01i>(1)-c'{f!lCIYf'l-c7)i, Cl) c t:...~T <IKY1Jt.,:t::!:l*'5 L. tv)i;-4. ftn-~:) .m. -c? ~{(/) '{, (1)n:m ~~ h -c IJ ;11; ·.-,;"-1~1! t G!: t.. -c lill"fl~ ~ -c ~ '5 (/) -cP-JIDli\f'i!!lt f,~i'tJ::;~ft(1)~~~t:... c ;~.: !: t !±I~- ZJi;i'-:Wllf(/)tii~ ~~Pi' t.iiiff\':21::. :a:H: L -c /i5''5lj\f.):liJj Gfl•::t;. ,.. t:.."-1-c. ~~CI)j~{!::CI)*kb't< J&ii;!.:!iUf(1)f.(l~f!::'ECI) 2 ~lll!~~~t..
t:.
j5r
~ iii

rr *

ns

rt

n

)*

In the above two species 10. claytonirma
and 0. bromeliar/olia). 2 or more pinnaeof lower part of the frond are always
fertile. and the others are sterile.
The external feature of the fertile
pinnae are quite different from thesterile.
4. Osmwula cinnamomea L. (fig. 7)
this now exists in Korth and SouthAmerica. East Asia. and in JapaneseIslands (including Sakhalin and Kuril Is.).
The fertile frond and the sterile frondof the species are different frond thatthese fronds arise from the same subterranean stem.
5. Osmzmda rega/is L. 1fig. 8) Osmunda
japonica TJ-IUNB. =Osmunda regalis L. var.
japo/li((/ M!LDE (fig. 5). This is almost
cosmopolitan species in geographicalrange. Commonly, the fertile frond of
the species differs from the sterile frond,
as well as Osmutula chmamomea. althoughthese fronds occur from the same subterranean stem. it is, however, the fertile
pinnae occurs. rarely. on the uppermost
part of the sterile frond.
Next, the most important geologicalOn the evolution of the vegetable
kingdom. the Osmundaceae is one of
the most interesting and importantfamily. All the species of the family
are crescentic petiolar bundle with several adaxial proroxylem. This crescentic
petiolar bundle is remarkable for its
similarity from the Palaeozoic genera tothat of the living Usmunda.
Among the family. the most important
living species are as follows:
L Todea barbara ~IOORE. (fig. {'. this
now exists in South Africa. Australia
and New Zealand etc. The fertile pinnules of the species are similar withthe sterile pinnules. and these occur on
the same pinnae of the same frond.
2. Osmmula dayto11iana L. (fig. 6), this
now exists in North America. East Asia
and Central Honshu of Japan.
3. Osmunda bromeliaefolia COPLA:'\Il,
this species now exists in Southern
Japan (Shikoku. Kyushu) Ryukyu. Bonin
Is. and Formosa.
-----~---*Received June 1. 1961; read at 78th
meeting of the Society at Akita, :\Jay U.
1961.

157

158

Seido EN!XJ

records are as follows (SEWARD, 1910):
1. Zalesskya gracilis IEtCHWAIW)
2. TlummoPteris sclzelchtendalii (EICH.)
These two species were recorded fromthe upper Permian beds in Russia, andthese are protostele in the stele structure.
3. Osmwufites kolbei SEWAIW, the spe·
cies was recorded from Cape Colony(probably Wealden in age). The fronds
with the stem were found imperfectly
preserved impressions of fronds referred
to Cladophlebis dentiwlata SEWARD. The
stem structure is mixed pith. composedof parenchyma and true tracheae.
4. Osmzmdites Dzmlopi KIDSTON andGWYNNE· VANGliAN.
This species was described from thejurassic rocks in the Otago district of
New Zealand in association with Cladoplllebis deutiwlata SEWARD. The stem
structure is similar to Todea barbara of
the recent species.
5. Osnuuu/itr:s Dmc!~eri CARRUTHERS.
This species was described by CARRU·
HIERS from the Lower Eocene beds at
Herne Bay: the pith is composed whollyof parenchyma and the xylem ring iscontinuous. From this type. by expan·
sion of the xylen ring and by the formation of overlapping leaf-gaps. the form
represented by Osmzmda regalis wasreached.
6. Osmumlites sl?idegatensis PE!\'HALWW.
This species was described by PEN·
HALLOW from the Lower Cretaceous ofCanada.
PENIIALLOW figures a fragment of aleaf bearing a superficial resemblance tothat of existing Osmunda C/aytonimza.
7. Todites Wil/iamsoni (BRO!'\Gl'\IART)The species was recorded by BRONG·
NIART, under the name Pecopteris Witliamsoui: afterwards. it was revised by
A. C. St-:WAJW. In the structure of thesporangia and in the general habit of
the fen i le fronds. these characters arequite similar to the recent species Todea
barbara. In Todites H'i/liamsmzi. thefertile pinnules and the sterile pinnules
occur on the same pinnae of the same
frond; and the fertile pinnules are quite
similar to the sterile.
The sterile pinnae of Todites ~Filliam·smzi is similar to the pinnae of Cladophlebis denticulata SEWARD.
In such a case. it is not easy to distinguish the sterile frond of Todites
IVilliam~mzi from C/adop/z/ebis de11ticulata.
As already stated; in the stele structure. Zalesskya. and Tlummopteris etc. of
the Paleozoic stems are a primitive type.. protostele ... while in the most common
recent species, Osmzmda 1·egalis. is advanced type "soleno-stele, and in the
Mesozoic stem Osmmulites Kolbei. thestele structure is intermediate type between the protostele and the solenostele;that is mixed pith, composed of parenchyma and true tracheae: st iII more. in
the Eocene species Osmwulites Dmf'lwri.
the pith is composed wholly of parenchyma and approach to recent Osmzmda
rega!ls.
The another t\1esozoic stern Osmundites
Dunlopi shows a closer approach to
Todea barbara than to recent species ofOsmunda. Therefore, the stem structureof Todea barbara is a primitive type. aswell as Mesozoic Osmzmdites Dzmlopi.
The external habit of sterile pinna
in Todites JVilliamsoni is quite similar
to Cladophlebis dellticulata, as SEWARD
says. and it is quite similar to thesterile pinnae of Osmmula claytonia11a
and the sterile frond of Osmzmda
cimzamomca.
In the habit of fertile frond, Todea
barbara is a most primitive. and Osmzmda
cimwmomea is a most evolved type.
Osmunda claytonimza and Osmunda
bromeliaefolia are intermediate type.
Generally, in the habit of fertile frond,

417. On the Evolution of the Osmzmdaceae()smzmda rega/is is in the advanced stage.
as well as Osmunda cilmamomea than.Osmunda bmmeliaefolia and Osmzmda
daytoniana; the former (0. regalis),·Occurs. rarely, fertile pinnae on theuppermost part of the same frond.
while the !alters (0. lmmzeliaefolia and0. claytoniana) occur, always, at thelowest or lower part of the pinnae inthe same frond.
The fertile pinnae of Osmunda bmme·
liaefolia occur at the lowest part oi thefrond; and it occurs, gradually, in theupper part of the frond.; in this case,the lowest pinna of the frond become
to sterile pinna; therefore, the habit.of the uppermost fertile pinnae of thefrond of Osnumda rega/is are in evoiYedstage than the common habit of Usmunda
clay/oniana or Osmwula bromeliaefolia.
Then. the fossil of Osmunda bromeliae·
folia was described by H. 1\IA'rsuo !1959)
from the 1\liocene plant beds of Noto pen in·sula. Ishikawa Prefecture. Honshu. japan.
H. ~·IATSUo described the species as
Dsmunda bromdiaefo!ioides. though it rl'·
vised erroneously to Osmunda tignitlllll
MATSUO's Osmwzda hrnnu>/iaefolioides isquite similar with the frond of living
Osmwula brmneliaefolia in its habit andspecial nervation of the pinnules. and it
can not be identified to the anotherspecies.
The frond of Osnumda bromeliae[oliahas a special and most characteristic
nervation. According to the characterof the nervation. it may be. safely,
identified with Osmtmda bromeliaefolia·COPLA:-;n, although, the fact that nosporangia have been found is a fatal·Objection to the identification. but, in
this case, it may be possible.
The
frond of Osmtmda bromcliaefolia may lle
briefly described as follows:
1) Osmwu/a bromeliaefolia 1 MATSUO\
Pl. 24. Figs. 2 and 3.

15!J

Description :-Frond bipinnate, pinnae
long lanceolate in shape, apex acute,base cuneate, length 10± ern .. maximum
width of the pinnae ca. 1.5 em. subopposite or alternate to the rachis.
Margins. deeply incised or dissected at
the middle and upper parts of the pinnae,and entire or serrated in the lower partor basal part.
Ivlidrib of Jobes (or pinnulcs) distinct,
secondary nerves simple or dichotomously branching, 2 or more in number. andit has. very often, another isolated nervein the same lobe lor pinnulel. which is
dichotomously branching or simple.
Texture coriaceous .
Remarks :-It is quite rare that thesame lobe (or pinnulel has the anotherisolated one nerve in addition to the
midrib. This is unknown to the present
writer in the frond of the filicales. ex·
cept the present species. Therefore. it
may be same species to Osmunda brome·
liaefolia.
According to l\'lATSt;O, the present
species occurs from the l\'liocene beds ofthe upper part of Nota flora formation.
Nakajima-machi, Kashima-gun. Ishikawa
Prefecture.
2) Osmum!a regalis L. var.?
Pl. 24. Fig. 1.
Desrriptioll ;-Frond bipinnate. pinnae
Janceolate in shape. apex obtuse. baseinequilateral; length 5 em.± maximum
width ca. 2 em.. arrangement of the
pinnae to the rachis. alternate or subopposite. l\1argins entire or minutely
serrated. l\lidrib of the pinnae. stout
and distinct. secondary nerves dichotom·
ously once or twice branching. rarely.
simple. abundant in number. Texture
coriaceous.
Remarks:- The present material isquite similar to the recent species. OJSIIIand HuziOKA (1941) described Osmwu.lajaponica TlltJNB. Subsp. nov. from thelGO

Seido ENtXJ

Ishikari Coal fields. It may be same spe·
cies with the present material. But thefact that no sporangia have been foundis a fatal objection to this identification.
The presem material was collectedfrom the Eocene IFoodtwrdia formationof Shimizusawa, Yubari-City, llokkaido,
Japan.
References
IVL\Tsuo, H. (1959) : A Revision of the i\-lioccne Fern of Osnmnda brome/iaefolioides

M '\T.' uo from Noto peninsula in the
Inner-side of Central japan, " The Science
Reports r~f the Kmwzazm l.iniz•ersity" Fol.
T·I. No. 2, pp. 85-90.
OtsHJ. S. and HuztoKA. K. (1941): Studies
on the Cenozoic Plants of Hokkaido andKarahuto. (1). Ferns from the Woodwardia Sandstone of Hokkaido. " Thefounzal of the Facully of Science, 1-lokkaido
Imp. Uniz•." Ser. IV, Vol. VI, 1\'o. 2. pp.
179-181. pl. xxxix, ligs. 1-8. -!a.
StC~I'ARD. A. C. (1910):
Fossil plants. vol. 2,
pp. 324-3,16.

Explanation of Plate 24
Fig. 1. Osmmzda regalis L. var.? Enlarged and Restored.
Collected from the Eocene Woodwardia zone of Jshikari Coal-field. Shimizusawa. Yubari·city. Hokkaido, Japan.
Figs. 2. 3. Osmzmda bromelial'folia CoPLA;>;D. x2.
Collected by H. ;\L\Tst.•o, from the :'lfiocenc Tajiri mudstone beds. Tsuchikawa, !\akajim·machi. Kashima-gun. Ishikawa Prefecture, Honshu. Japan.
Fig. 4. Todea barbara NlooR E(a), a part of frond (ca. 1/2 nat. size). (b). 0.1 part of ~crtilc pinnulc (enlarged). (c),Sporangium, much enlarged, after Kunze.
Fig. 5. Osmwcda regalis L. var. japonicrr MtLDE.
Showing the frond with the fertile pinnae on the upper part of the frond. Culti,·atedby S. E:-;no, much rr.>.duccd.
Fig. G. Osmunda cluytoniatta L. Much reduced. After :\r<DRI·:ws. Ancient plants l!J-17.
Fig. 7. Osmzmda cimramomea L.
Fig-. 8. Osmcmda regalis L.

E NDO :On the Evolution of the Osmundaceae

Plate 24

l

3

5

7

Trans. Proc. Palaeont. Soc. Japan. N. S., No. 4-1. pp. 161-166. pl. 25, Dec. 15. 1961

-ll8.

ON THE NE\N GEl\US METADOLIOLINA*
KE!\·ICHI ISHIIDivision Geosci., Fac. S: i .. Osaka City Cni\·.

andYASUO NOGAMI
Gcol. and Miner. lnst.. Coli. Sci .. Kyolo Univ.

m~ Metadoliolit~a :.:...-:>\·1:': ·r·'E*R*l.' Pseudodoliolina 1-:Jii!/\~it ..C\·t.:.iil!l.:,%:fffi£l'-Ji:i,, g_;,1W.>'fB'J;_:i,. ::..J:;fi,..,f:_ 2 "?I!)!J'JL-;/i):(h~:::._)::_'/j:J}.J::.(/)f.;.,. ,:,*1,&1!) 2 -:JI!) 1/ 1v- 71!)i'fi[sJJ>±~f.i.iTF;~ ''i-/J!i1i!l:~tl:c.t~7.dt~i1· G fJ. I). ft!!:fj!i.?ffl!ll~ tt:ii~:W,~ '/j:jof ~ J:; ll7t1l:~tl· G f.i..., "( ~ · G C: It· j .::. C: :::' &>, ~ ol!:I~l;t Pseudodoliolina ozawai(Pseudodoliolina l!)~:>tJi!D 't'ft*~ tt~ !f Jv-7''t'(h I). f&'l'fli Pseudodolioliua gral!i·
testa 't'ft~6.tt~ !7 1v- 7-c-&>, ~ o ~~~li:::.. At G(l)~!!ili(l)i\'4':!'i'/j:f!\:j:JU... "(. /&~Q) /lv7l.:.t.:.v·L~'!.l1; Metadoliolina '/j:fle~I<\Lt:o :l.fetado/iolina 1:1: Pseudodo/io/ina tJ·:C:>7tii:'.L

kJ::.~~~.

~

~

~

-·~

~

~

~

morphological differences. the bothgroups differ from each other in bio·
stratigraphic
occurrences.
Namely.
Pseudodoliolina pseudo!epida gravitesta ishitherto discovered only from the Upper
Permian Kuma formation in Kyushu.
the Doi group and the Ilaigyu groupin Shikoku and the 1\laizuru group in
Honshu. which are characterized by theadvanced species of Yabeina. Lepidolina.
etc. and are Late Permian (the zone of
Lepidolina) in age. The group of Pseudo·
do!iolina ozazmi is. on the other hand,reported from the upper part of theLower Permian to the .t>.liddle Permianof Akasaka. Akiyoshi. Atetsu. etc .. inassociation with abundant individuals of
Parajusu!ina and Neosclucagerina.
The above morphological and biostratigraphical consideration leads us toIn the course of our study on some
Permian fusulinids from japan. we noticed
that in the genus Pseudodoliolina ofjapan there are represented two distinct
forms. namely the group of Pseudo·
dotiolina ozmmi and that of Pseudo·
dolioli11a pseudo!epida grm:itesta. The
two groups resemble each other ingeneral morphology. Ho\Yever. the groupof Pseudodulio!ina pseudo!epida grmJitesladiffers from the group of Ps. ozmmi
mainly in having the well differentiatedlayers even in early stage of ontogeny.
Furthermore Pseudodoliolina pseudolepida
gravitesta has larger form. more slendersepta and narrower parachomata thanthe latter group. In addition to those* Received June 17, 1961: read at the 77th
meeting of the Society at Nagoya. Nov. 19.
1960.
lGl

162

Ken-ic/d ISHII and Yasuo Noc~Lllirestrict the genus Pseudodoliolina to thegroup of Pseudodoliolina oua('(li and to
propose here a new generic name. Metado/ioli71a for the group of Pseudodoliolina
pseudolepida gravitesta. This new genus
includes some species hitherto assigned
either to J>seudodoliolina or to Miselliua.

Description

Family Fusulinidae r-.lt)LI.Jm, 1878
Subfamily Verbeeekininae STAFF
et \VEDEKINO, 1910
Genus Metadoliolina ISJJIIet NOGAlV!I, new genus

Type-species :-Pseudodo/ioli11a pseudo·
Jepida gnwilesta KAX~IERA. 1954, 11iem.
Fac. Sci., Kyuslzu Univ., ser. D. (;eo/., vol.
4, no. 1, p. 12. pl. 2, figs. 1-6. (Pseudodoliolina pseudotepida gravitesta is in thispaper regarded as a distinct species.
which is generically separated from
Pseudodo/iolina pseudolepida: see below).
Diagnosis :-Shell large, elongate ellipsoidal to subcylindrical; central portion
slightly inflated to flattened; polar ends
broadly rounded: proloculus small tomoderate: shell expands slowly anduniformly: spirotheca not so thin. composed of tectum and lower less dense
layer in inner three or four volutions,and of tectum, thick diaphanotheca, andthin upper and lower tectoria in outer
volutions beyond them; in some partsof shell or in some specimens tectoria
often absent, keriotheca present; septa
comparatively slender. extended anteriorly at small angle from normal tospirotheca; parachomata numerous, nar·row and small.
Comparison :-Metadoliolina resembles
Pseudodoliolina YABE et I lANZA \VA, 1932

(type-species:
Pseudodoliolina ozmcai
Y ABE et HANZA WA, 1932) in the shellform, and the mode of parachomata andsepta. However. the two genera can be
distinguished from each other as follows:
1) The spirotheca of Mf'ladoliolina isdistinctly differentiated, and except for
inner two or three volutions it is composed of a tectum. a thick transparent
layer (diaphanotheca) and thin upperand lower tectoria. Furthermore, they
are often perforated by fine para\\e\dark line. This fine alveolar structure
is similar to the alveoli of the genus
Verheekina (Pl. 25, fig. '1). On the other
hand, l he spirotheca of Pscudodoliolina
is composed of an undifferentiated layer
Ia tectum and protheca) in almost allvolutions. In the type-species it is verythin and a compact single layer (Pl. 25.
fig. 6), and in some other species suchas Pseudodo!iolina pseudo lepida and Ps.
sp. (J'l. 25, figs. 7. 8) from Atetsu ofjapan, it is comparatively thicker thanthat of the type-species. In outermost
volutions the spirotheca of those species
is rarely differentiated into a transparent
layer and fine porous layer 'not keriotheca). 2) The differences of the twogenera in other features are not so large
as that in the spirotheca. In general
111etadolioliua has a larger size of theshell, more slender septa and smaller
parachornata than Pseudodoliolina.
Metadoliolina is allied to the genus
Verbeel~ina in the spirothecal structure.
The former can be. however. easily dis·tinguished from the latter by the ellipsoidal and subcylindrical form. and more
distinct parachomata even in the ontogenetically younger stages.
Metadolioliua can be distinguishedfrom the genus J1isellina (type-species:
Dolio/ina ova/is DEPRAT. 1915) in size.
shell form. foramina and parachomata.
Remarks:- The foreign species such as418.

On the new genus JVJetadoliolina

Sclucagerina !epida SCHWAGER. 1883.* S.
lepida e!lipsoidalis ScmVAGER. 1833. Dolio·!ina major DEPRAT. 1914 and D. dozwillei
GUBLER. 1935 may be also included within
the genus Jl.letadoliolina.
Sclncagerina lepida SCHWAGER wasreferred by THOMPSOl\ (1948) tentatively
to the genus Misellina, but he remarked
at the same time that it might not be
congeneric with the type-species. Doliolina
ovatis DEPRAT. Scliwagerina tepida haslarger size. more numerous volutions,and narrower and more numerous parachomata than the type-species of Misellina. Furthermore. in the attitude of
septa of the sagittal section Scliwagerina!epida belongs to Pseudodoliolina·type,whereas !l!isellina belongs to Cancellina·
type. In many shell features Sclncagerina
iepida resembles Pseudodo!iolina rather
than Misellina. In the presence of the
keriothecal structure. however. Schwagerina !epida cannot be referred to
Pseudodoliolina. In our opinion it should
be included within Metadolio!ina.
Doliolina major may be regarded as asynonym of Schwageriua lepida. D.
douvillei which resembles the genus
Pseudodoliolina in many shell characters,
have clear kcriothecal structure like s.
lepida. Therefore it can be included
within the genus Metadoliolina.
Distribution :-~Iiddle ? and Upper Per·
mian: japan. China and Cambodia.
Metadoliolina graritesta(KAC\~IERA). 195'1
Plate 25. Figs. 1-·!.

* Schwagerina lepida-SCH\\'AGER. 1883 is thetypc·spedes of Doliolina ScHELL\\'11·:1", 190Z,
which is however preoccupied by Doliolina
BORGERT,1894 (SCI-IEl'CK & T11o:-n•so!'\,1940). ln order to avoid CYentual misunderstanding it is h~rc mentioned that :lfisellina
is not a substitute 1:amc for Doliotina SciiEI.L·\\IE:". 1902.

1();3

Pseudodoliolina pseudolepida gralJi/esta
four. Fac. Sci., Kyuslm Unh• ..
ser. JJ, Geol .. vol. 4. no. 1. pp. 12-1•1, pl.
2. figs. 1-6.
1954. Pseudodoliolina n. sp.?, KA:-.:MF.Rt\,
IJitfo., pp. 14-15. pl. z. figs. 7. 8.
1954. Verbeekim1 ? n. sp .. KA:"~IER.·\. Ditto ..
pp. 15-16. pl. 2, fig. 9.
1960. Pseudodoliolina pseudo/epida gl'{ll'ilesta
CHISAKA, four. Coli. Arts. Sci., Clziba
Univ .. t•ol. 3. no. 2, p. 246, pl. •1. figs.
1-6.
cfr. 1956. Misellina spp. a. b, NooA, Rep.
Earth Sci., Dept. Gener. Edu .. Kmslm
U11iv., z•ol. 2. pp. 12-13. pl. 1, figs. 11,
15b, 16, pl. 3, figs. 4. 5. pl. 4, figs. la,
lc.
1954.

K.·\:'>11\IF.RA.

Description:-The size and number ofvolution of mature specimens is difficult
to decide precisely, because outer several
volutions of most specimens are often
crushed away. But mature specimens
seem to have 20 volutions or more. Shelllarge. elongate ellipsoidal to subcylind·rica!, tightly coiled, with flattened orslightly inflated central portion andbroadly rounded poles. Specimens of 20
volutions are 8.4-9.3 mm long and 4.26.1 mm wide : form ratio varies from 1.9
to 2.0.
Proloculus moderate to large. its out·side diameter 135-320 microns. Shell
expands slowly and uniformly.
Spirotheca not very thin. In inner
two or three volulions it is comparati\·elythin and is composed of tectum and lessdense layer. Beyond three or four volu·
tions it is clearly composed of thintectum. rather thick transparent layer(diaphanotheca) and very thin upper ~nd
lower tectoria. Keriothecal structure isseen in some volutions: in some speci·
mens it is observed in all volutions.
Thickness of spirothcca measures 5-10
microns in the 5th volution, 10-20
microns in the lOth. 15-30 microns in
the 15th and 30-35 microns in the 20th.

Hi-l

Ken-ichi ISHII and Yasuo NuGAMI

Septa closely spaced. and nearly normal
to the spirotheca. It is composed of adownward deflection of tectum and an·
terior and posterior downward extensionsof diaphanotheca or keriotheca of spiro·
theca.
Parachomata narrow and, in ontogenetically adult stages, low. They arehalf or more as high as chambers inearly stages. nearly half in middle stages
and less than half in adult stages.
Foramina small. circular or ellipticalin cross section.
Comparison: -llfetadoliolina gral'ilesta
was originally proposed as a subspeciesof Pscud(){iolio!ina pseudo/epida ( 1\AN:\IERA, 1954). However. the former dif·
fcrs essentially from the !at ter by theappearance of the diaphanothecal or
keriothecal structure even in the onto·
genetical early stages.
This species can be distinguished from
Metadoliolilw !cpida (SCHWAGER), including M. ll'pida el/ipsoidalis and iii major(DEPRAT). in larger and longer shell.
larger proloculus. thicker spirothcca.
clearer diaphanotheca, smaller para·chomata and smaller foramina.
Furthermore this species is easily
distinguished from Metadoliolina dmwillei(GUBLER) and Misellina major in CHE~
(1956) [ =Metatlo/iolina douvi/lei]. which
resemble .H. lepida in shell characters.
by the shell form. shell size and thickness of diaphanotheca.
iV!aterial :-Lectotype, Reg. no. 1\. u 49997, a limestone lens (Loc. 1\.u 499) of theupper coarse of the small tributary
towards the northern side of the Kawamata valley, Kawamata-mura, Yatsu·
shiro-gun. Kumamoto Pref.. Kyushu.
Other specimens. PF 1400. a limestonelens. 300m. north of the bridge at 1-laigyuguchi. Miyahama-mura, Naka·gun,
Tokushima Pref.. Shikoku: JPF -10505, alimestone lens. about 500 m, northwestof Nabae. Takahama·cho. Ooi-gun, Fukui
Pref.. Hokuriku. These specimens arc
stored in the Geological Institute, Kyu·
shu University 0\.u). in the Division ofc;eoscience. Osaka City University I.PF).
and in the Geological Institute. Kyoto
University UPF). Ku ·199-97. collected by
KAN:'>IERA, PF 1400. collected by SUYARI,.JPF-10505 collected by NAKAZAWA.
Oaurre11cc :-The lectotype and Ku
499-27 occur in the Kuma formation,
Kyushu. PF 1400, in the llaigyu group.
Shikoku and JPF-10505 in the l\laizura
group. 1-lokuriku. As known at present.
occurrence of these specimens is con·fined to the Late Permian (the :tone of
Lepidolina l.

Phylogeny of Meiadoliolina
and Pseuclocloliolina
As mentioned already, 11fetadoliolina
and l'seudndoliolina resemble each otherin general shell characters excepting the
spirot heca. However, the type-speciesof Metado/iolina. which is confined to
the Late Permian. is larger in size andhas lower and narrower parachomata
than the type-species of l'seudodoliolina.
i.e .. Pseudodoliolina ozazcai which is late
Early Permian and tvliddle Permian in age(Pl. 25. fig. 5). The mode of parachomata
and septa in the ontogenetically adultstages of Psl!udodo/io/ina nzmmi resembles
to some extent that of Metadoliolina.
Fun her more Pseudodoliolina pseudo/epida.
which is i\liddle Permian in age. ismorphologically intermediate between
them. The exact geological horizon of:Hetado/iolina /epida is unknown. but it
may be Middle Permian in age rather
than Upper Permian.
Metado!iolina
lepida resembles Pseudodolio/ina pseudo·/epida in many shell features except forthe juvenile stages and the spirothecal
structure.

418.

On tlze new genus MetadotioliuaAmong various morphological elementsof fusulinids. the spirotheca makes themost remarkable change in the courseof phylogeny. As remarked by ISHII(19:59). following eyolutionary stages ofwall are recognizable in many phylogenetic series within the fusulinids:
the undifferentiated layer or a single
layer (Eostaffel/a-type wall)-> parallel
layer [tectum plus lower dense layer--.
tectum plus transparent layer (diaphanotheca). sometimes associated with secondary layer (tectoriaj]-> appearence ofporous layer-> development of alveolar
layer (keriotheca). The wall of Pseudodo!io!ina indicates the stage of the undifferentiated layer or the primitive
parallel layer. while that of il!etadoliolina
indicates that of advanced parallel layer
or alveolar layer.
Thus. considering the mode of thedevelopment of morphological features
such as size. parachornata and spirotheca
and the geological occurrences of Pseudodoliolina and J1Ietadoliolina. we conclude
that illetado!io!ina was derived from
P:>eudodo!iolina.
Acknowledgements

We express our deep gratitude toAssist. Prof. K. KANxiERA (Kyushu
University) for the oppotunity of reexamining type-specimens of Metadolio!ina
gral!itesta, and to Dr. K. SUYArn IToku·shima University) for his kind offering
at our disposal of a rock-sample collectedby himself. We wish to express ourthanks also to Assist Prof. K. lC!l!KA WA(Osaka City Universityl for effective
counsel and reading of the manuscript.
References
Cm:;-;, S .. 1956: Fusulinids of South China.
Part II, P,zfaeontologi:l ,<:inica. new ser. /3.

lt15

no. 6, pp. 1-71. pls. 1-14.
T .. 1960: On some Permian Fusulinids from th~ Takegami Conglomerate.
ChoshiPeninsula. Chiba Prefecture,
Japan. four. Col/. :lrt. Sci. Chiba Unir ..
1'0/. 3. no. 2. pp. 235-25·1. pis. 1-9.
DEI'R.-\T. J. 1914:
Etud~ comparati\·e desFusulinidcs d'Akasaka (Japan) et desFusulinides de Chine et d'lndochine. l'vlem.
Sen·. Geol. hulodzim. !'(){. 3. ,fasc. 1. pp.
1--!5. pis. 1 8.
- - . 1915: Etude dt?s l'usulinides de Chineet d'lndochine et classification des calcaires it fusulines (IV" 1\lemoire). Les
Fusulinides des calcaires carboniferienset P•2rmiens du Tonkin. du Laos et duNord-Annam. ibid .. z.·o/. 4. fasc. I. pp. 130. pis. 1-3.
Gulll.ER. J .. 1935: Les Fusulinides du per·mien de l'lndochine. leur structure etleur classification. Soc. Glial. France. Jlem ..
11. s .. tome I 1. fc:sr . .J. 110. 26. pp. 1-173.
pis. 1 8 (18 24).
ls1111. K .. 1959:
On the Evolution of theFusulinida. Earth Science (Chik.l'/1 Ka~<aku).
no. 45. pp. 5-H (in Japanese).
K.·\:'O!ERA. K .. 195-!:
Fusulinids from theUpp-2r Permian Kuma Formation, South·
ern Kyushu, Japan with special Rder·ence to th~ Fusulinid zone in the Upper
Perm ian of Japan . .Hem. Fac Sci .. l(ruslm
Univ .. ser. D. Geo!.. vo/. •I. 110. 1. pp. 1-38,
pis. 1-6.
NooA, lVI., 1956: Stratigr~phical and P;:>.lae·
ontologic:al Studi("s of the Toman Formation in the Kaishantun and Kamisanbo
District. Rep. t.'arth ::ci. General Educ ..
Kruslm Unh· .. ro/. 2. p. 14, pl. 4, p. 1-22.
pis. 1-6.
OZ.-\\\'A. Y .. 1927: Stratigraphical studies of
the Fusulina limestone of Akasaka. Province of Mino. Tokyo Imp. Uuiz• .. Fac. Sci ..
sec. 2. val. 2. pt. 3. pp. 121-164. pis. 34-45.
SCBE:"'CK, H. G. & '\L L. THO.\IPSO". 1940:
JUisel/ina and Bren:xina. new Permian
fusulinid Foraminifera. four. Paleontolog\·,l'Ol. 14. pp. 548-589.
Sell\\' AGER.
C.. 1883:
Carbonische Foraminiferen aus China und Japan. Nm
RTrHTlloFEN'.' China. 1'0/. 4. pp. 106-159.
CIIISAKA.

1615

Ken-iclzi IsHII and Yasuo NoG!lMI

pis. 15-18.
THO:I.tPso:--;, l\t L.. 1948: Studies of American
Fusulinids. Uuiz•. Kausas P.:~le<mt. Coutrib.,
Protozoa, art. 1. pp. 1-18•1, pis. 1-38.
- - & C. L. FosTER. 1937: l'vliddle l'ermian
fusulinids from Szechuan. China. four.

Pa/eoutology, t•ol. 11, pp. 126-144, pls. 2325.
YABE, H. & S. HA~ZAWA, 1932: Tentativeclassification ~f the Foraminifera of theFusulinidac. hizp. A.cad., /'roc., l'O/. 8. pp.
-10-43.

Explanation of Plate 25
Figs. 1-4. Metadolioliua grat•itesta (l<A:-.!MERA). 195•1
1. Axial section, from the llaigyu group in Shikoku, deposited in Divtsion Geosci.,
Osaka City Univ., Reg. no. PF 1400, X ca. 10.
2. Axial section of the lectotype (K,\:\:1-tERA, 1954, pl. 2, fig. 1), from the Kuma forma·tion in Kyushu, deposited in Geol. Inst .. Kyushu Univ .. Reg. no. Ku 499-97, x ca. 10.
3. Slightly excentric sagittal section. from the Maizuru group in Hokuriku. depositedin Geol. & Miner. Inst.. Univ. Kyoto, Reg. no. JPE-10505, X ca. 10.
4. Enlargement of fig. 2. showing the wall structure, x ca. 200.
Figs. 5. 6. Pseudodoliolitla ozau·ai y,,nr. et HA:-.IZAWA. 1932
5. Axial section of the topotype, occurred almost exclusively from the Doliolina horizon(OZAWA, 1927) of the Akasaka limestone, deposited in Division Gcosci., Osaka City
Univ .. Reg. no. PF 1401. x ca. 10.
6. Enlargement of fig. 5, showing the wall structure, x ca. 200.
Fig. 7. Pseudodo/ioli11a pseudo/epida (DEPRAT). 191·1
Axial section. occurred together with J\'eoschwagerina dwni, N. margaritae etc. fromthe N. douz•i!lei-N. margarilae zone (NoGA~II. 1961) of the Atetsu limestone, depositedin Geol. & Miner. lnst. Univ. Kyoto, HCI<· no. JPF-10321. x ca. 10.
Fig. 8. l'seudodoliolina sp. indet.
Axial section, occurred together with Neoschwageritltt dorll'illei, Pseudodo/ioli11a pseudo/epida etc. from the N. doul'illei·:V. margaritae zone (NoGA:I-11, 1961) of the Atetsu limestone, deposited in Geol. & Miner. Inst., Univ. Kyoto. Heg. no. JPF-10327. x ca. 10.

/ SHIJ qnd N OGAMJ:

lsrm and NOGAM I photo

MetaC/qliQliNq

Plate ?5

Trans. I' roc. Palacont. Soc. japan, N. S., No. 44. pp. 167-170, Dec. 15, Hl6!

41 !1.

SPINIL1!.13ERIS, A NEW GEl'\ US OF OSTRACODA
FRO.M THE PACIFIC*TETSURO HANAI
Institute of Geology, University of Tokyol(rl,.!i Spini/eberis (T~·pe species: C)/there quadriaczdeata BRADY)~at:~ L t:..

.:. (!)

tti~:±m (.~'f-trJ: 'J s*c7>~;?ilil!!~::v,·:...,-ca**<?;.:~L~r{>, r:T~lin>t-;i'F"Jl"?.b;,:,.

?E#ff!lJj
It has been known since the time of
the Challenger f~cport that a species
called Cytltere quadriant!eata is distributed widely over the Pacific area. In
revising old generic identification ofjapanese Ostracoda in light of the recent
advancements of ostracod taxonomy,
the writer found that the species has
certain characters which do not permit
its assignment to any known genera orsubgenera. l\ new genus Spinileberis is.
therefore. proposed for this peculiar
form. However. the classificatory positionof the new gcn us is at present uncertain.
Gratitudes arc expressed to Professors
Teiichi KoBA YASIII and Fuyuji TAKA!
of the University of Tokyo and to Professor H. V. HowE of Louisiana State
University. Baton Rouge, Louisiana for
their continued encouragement.

the anterior margin. Carapace has amedian sulcus and four longitudinalridges. Dorsal ridge of each valve projects so as to make a v-shaped troughalong the hinge line. Caudal process
does not develop. but one posterior
radial pore canal is bifurcated. l\larginal
duplicature broad. l~adial pore canals
few. straight and regularly spaced. No
vestibule. Hinge amphidonl.
Remarks: -Cytl1ere quadriant!eata
BRADY has close resemblance in outline
to genus Neomouoceratina.
llowcver,Amphidont dentition of this species
does not permit to place this new genus
to schizodont group of Ostracoda.

Spini/eberis quadriacu/eata (BRAl>Y)
Text-figs. 1-7.
Cytlzere quadriaculeata BRADY. BH.\DY 1880,
Challenger Rept., vol. 1. pp. 86. 87, pl.
22. figs. 2a-d. pl. 25. figs. 4a-d: CIIAI'M.\:1:
1910. Jour. Linn. Soc.. Yol. 30. p. -!32 :CHAP:\IA:I: 1915. Zool. Res. Fishing Exp.
F. I. S. Endea\·our. vol. 3, p. ·13: CII.\P~I.\:1:
1916, Brit. Antarc. Exp .. Gcol.. \'OI. 2. pt.
3, p. 73, pl. 6. fig. 47: CII:\1':\IA!'i 1926.
Pmc. Rev. Soc. Victoria. vol. 38. (n. s.)
pp. 127. 128: CII;\1'1\IA:X. CRESI'IN and
KEBLE 1928. Rec. Geol. Sur\'. Victoria 5.
no. 1. p. 170.

Subfamily uncertain
Genus Spinileberis IIA;.<AJ, n. gen.

Type-species :-Cythere quadriaculeata
BRADY, 1880.
Diaguosis :-Carapace subquadrate.
Outer lip projects to make fringe along* R;!c;;ivcd July 12. 1961; read at 79th
meeting of the Society at Kanazawa. Sept.
23, 1961.
Hi I

168

Tetsuro H41 1AI
1

1

3

4

5

6

Text-figures 1~7. Spiuileberis
quadriaculeala (BR.\DY)l, lateral view of right valve (female).
2, lateral view of left valve (female).
3. internal view of left valve (female).
4, internal view of right valve (female).
5, internal view of left vah•c (male).
6, internal view of right valve (male).
7, dorsal view (female).

7

419.
Cytltereis

SpinifebEris. a new genus of Ostracoda from the Pacificquadriuculeata (BRAOY). G. W.
l!Jl2. Das Ticrreich 31. p. 357.

NlL'LLER

Figured specimens :-Geol. Inst.. Tokyo
Uni\".. CA '1995. a right valve. female.
text-figs. 1, 4, 7: CA 4996. a left valve.
female, text-figs. 2. 3. 7: CA 4997. aright valve, male. text-fig. 6: CA 4998.
a left valve. male. text·fig. 5.
Description :-Carapace irregularly subquadrate in lateral outline.
Anterior
margin obliquely and broadly rounded.
Outer lip projects to make fringe along
the anterior margin.
Dorsal contact
margin nearly straight and steeply inclined posteriorly: obscured by the development of the dorsal ridge to makea lateral outline which is slightly convex
and suddenly excavated in its posterior
end. Ventral contact margin slightly
concave obscured by the arched projection of bottom ridge in lateral outline. Posterior margin triangular nearlystraight to slightly concavecl in theupper half and slightly convexcd in thelower half.
Posterior cardinal angle
distinct. Carapace compressed latl:rally
in the anterior cardinal marginal area.
Surface has a median sulcus and ornamented with longitudinal ridges. Dorsal
ridge runs along the dorsal margin andsufficiently high enough to obscure thedorsal contact margin : the ridge curveat the anterior cardina angle. and reachthe anterior part of the central ridge.
Central ridge runs obliquely across themedian sulcus and terminates posteriorly
in a sharp spine. Bottom ridge runs
close to and parallel with the ventral
contact margin. Surface covered witha fairly coarse reticulation except in l heanterior laterally compressed area. In
dorsal view. carapace appears to bearrow-head shaped.
Dorsal ridge ofeach valve projects so as to make av-shaped trough along the hinge line.

HiH

Ventral surface roughly flattened. Central and ventral ridges give angulation
to make pentagonal appearance in anterior view.
I\'larginal duplicature broad.
Radialpore canals simple. straight and few;
about eight regularly spaced along theanterior margin.
Pseudo-radial pore
canals prescn l.
One posterior radialpore canal bifurcates. Hinge amphidont.
Anterior and antero-median element issmooth : postero-median element is crenulated and posterior element is smooth.
Snap pit and knob structure develop
weakly. Adductor muscle scar pattern
not observable.
Sexual dimorphism fairly strong: malemore elongate than female forms.
Dimensions :-Figured specimens :CA
49951 a right valve. female. length 0.61mm., height 0.37 mm .. thickness 0.15 mm.:
1CA 4996) a left valve. female. length
0.64 mm .. height 041 mm .. thickness 017
mm.: (CA ·1997) a right valve, male.
length 0.60 mm .. height 0.34 mm .. thickness 015 mn1.: (CA 49981 a left valve,male. length 0.57 mm., height 0.33 mm.,thickness 0.12 mm.
Occurrence :-The species was originally reported by BRADY (1880) from theInland Sea of japan (Setonaikai) andfrom off the reef at Honolulu, Hawaiian
Is. CHAI'MJ\ N (1910, 1915, 1916, 1926) andCliAP~IAN and CRESPI~ tl928) reported
this species from the Southern Pacificto the Antarctic Sea. Since the species
from Japan seems to be a inhabitant ofwarm water mud, the distribution ofthis species in the cold water of thesouthern hemisphere may require further
study. In Japan. this species is foundliving in the soft mud of the shallow
warm (10° -27°C) coastal water at Aburat:.ubo Cove. Abundant fossil specimens
are obtained from the lower horizon of
the middle Pleistocene Sakurai silty

170

Tetsuro JJ..tNAI

sand exposed along a cliff facing Tokyo
Bay at Sakurai. Kisarazu-shi, Chiba
Prefecture.
Remarlls :-This species is quite similar
to Neomonoceratina microretibulah· in its
outline and its ornamentations. This isprobably due to convergence, because.
in detailed shell structure. the carapaceof this species is quite different from
Neommwceratina. The frill-like projection along the anterior margin of thisspecies appears to be quite similar to
those of Nemnonoceratina. The frill of
this new genus is. however. a modification of the outer lamella. whereas thefrill of Neomonoceralina is a modificationof the calcified marginal portion of theinner lamella.
llingement of this genus is very close
to that of Neomonoceratina in its appearance. However, the anterior andantero·median element is not split (i.e.
not schizodontJ. and is identical withamphidont dentition. Posterior caudal
process of this genus is not developed
but one caudal radial pore canal of posterior extremity is always bifurcated.
r-.targinal duplicature of the new genus
is far broader than that of Nr:omono·
cera/ina. However. the radial porC' canalsof both genera are few tabout eight} innumber; straight and regularly spacedin this new genus, but more or lesstend to converge in the antero-ventral
area in lV"eomonoceratina.
Although the hinge structure is notquite the same, the species described by
KEY (1954) from Manila. Philippins. as
Paijenborcltella (Neonumocerati na) koenigs-

ll'aldi KEY may perhaps be closely related
to this ne\\' genus.
References
G. S .. 1880. Report of the scientificresults of the Voya_ge of H. M.S. Challenger. during the years 1873-1876. Zoo!.,vol. 1. pt. 3, Ostracoda. 184 pp .. 4-l pis.
CtiAP:..tA:". F., 1910, On the Foraminifera andOstracoda from Soundings (chiefly deepwater) collected round Funafuti by
H. M.S. Penguin: jour. Linn. So<·.. 1'0/.
BRADY.

30. pp. 388-·J.l.f. pis. 5·1-57.

- - . 1915. Report on the Foraminifera andOstracoda obtained by the F. I. S. Emlca·
vour from the east coast of Tasmania
and off Cape \Viles, South Australia:
Zoo/. Results of the Fishing Experime11ts
carried 011 by F. f. S. Eudeal'Our. 190 91914. l'ol. 3. pp. 1-51, pis. 1-3.

- - . 1916. Report on the Foraminifera andOstracoda from elevated deposits on the
shores of the Ross Sea: British ilntarclic
Expedition 1907-1909. Rep/. ~ci. lnl'estigations. Geot. t•ot. 2. Contribution to the
Palaeontology end Petrology of South
Victoria Land, pt. 2, pp. 25-51. 6 p/s.
- - . 1926. Geological Notes on Neumerella
and the section from Bairnsdalc toOrbost: Proc. Roy. Soc .. Victoria. 1.'0/. 38,
1/.S., pp.

125-142. pl. 10.

CtL\P~!A!",F .. CRESPI);. 1. and R. A. KERLE.
1928. The Sorrento Bore. Mornington
Peninsula. with a description of new orlittle known fossils: Rec. Geol. Sun·.,Victm·ia. MI. 5, pt. 1. pj). 1-195, 12 p/s.
1954. Some recent Ostracoda of

KEY, A.].,l\Ianila (Philippines): Pro. Kon. Ned.
llkad. Wet. Amsterdam. scr. b, z•ot. 5;', no.
3. pp. 35 I -363.

Trans. Proc. Palaeont. Soc. Japan, N. S., No . .U, pp. 171-17.t. pl. 26. Dec. 15. 1961

420.

ON A EQUID FOSSIL FROM HIRAMAKI F'ORMATIOK*TOKIO SHIKAMA
Geological Institute. Yokohama National Untversity. Yokohama

and
SIIINJI YOSHIDA
Institute of Earth Sciences. Aichi Gakugci University. Nagoya~J~tiljl,tlJYCmif!~Cf)!l.l~l.:t~l:t '.>.P~tf! • -ijlf{~""ff!litJ· '?@. Lt:..l.!~(1)""f~a:-klll/: L t:.,!IE
In April. 1958. R. HillA. a farmer in
Hiramaki, Kani city. Gifu Prefecture
found a lower jaw of Anc!zitlzerimn on avalley wall of Ohara in Hiramaki. andpossessed it as a part of dragon remains.
The junior writer. while making a fieldsurvey of this area. acquired this information and could borrow the specimen
from l-IIDA for this stud)'. From Omori,
1 km southeast of this locality, two frag·mental teeth of .·lnclzitllerium were foundand described by H..MATSU~IOTO in 1921.
The specimen described here is of thesame species as i\IATSUl\IOTO's and is in
far better preservation. The formationof the neighbourhood of the locality
consists of alternation of white tuffaceous
sandstone and black siltstone and belongs
to the uppermost part of Lower II ira·
maki formation. l'viATSUJ\IOTO described
Palaeotapirus yagii \lAT., Clzilotherium
pugnatur (i\IAT.), Ampliitragulus minoensis.MAT .. Gumphotlzerium annectens (MAT.).

r.rJ n!f* ·if m '-fr

sciurid sp. besides this species. He correlated the fauna of Hiramaki formation
with that of European Burdigalian or
Gaj-Lowcr Siwalik of India. In 1949 F.
TAKA! reported Palaeotapirus yagii and
Chilotherium pugnator from Katabira,Kani city and regarded the age of Hiramaki fauna as middle to late Miocene.
It may be significant that the species inquestion is yery much like A gobiense
CoLBERT from Tungur formation ofOuter Mongolia. Considered from thispoint of view. the uppermost part ofLower Hiramaki formation may betreated as late ~vliocene in age.
Here the writers extend their hearty
thanks to Prof. F. T AKAI, Dr. T. HAI'AI.
l'vlr. Y. HASECAWA for their valuableadvice and help during the course ofthis research and to Mr. R. HIDA for
his kindness to permit them to usc thisspecimen for their study .
Anchitherium hypoliippoides* Received July H. 19Gl; read at 77th

!V!ATSU:\IOTO. 19:21

Meeting of the Society at Nagoya. Nov. 19,

Pl. 26. ligs. 1-9

1960.

111

172
1921.

Tokio SH£KA.M.4 and Shinji YoSHIDA!:ci. Rep. Tolioku Imp.
Unit•., ser. 2. 1'01. .5, 110. 3, pp. 77-78.

MATsniOTO,

pl. 13. figs. 1-5.

Specimens:-Right and left rami of asame individual, collected and possessedby R. l-ImA ; gypsum model stored inGeological Institute. Yokohama ?>Jational
University.
Locality :-Ohata, Yamazaki. 1-liramaki.
Kani city. Gihu Prefecture.
Ilorizou :-Uppermost part of Lower
Hiramaki formation ; late .\Eocene.
Description :-Right and left horizontal
rami preserved in the same blocl( ofgreyish white hard tuffaceous sandstone,
the former retaining five cheek teeth
from Pc to ivl" and the latter live cheekteeth of P 2 , P,. l\1 1 • lVL, and rv!,.: teeth
thick black in colour and bone dar!<brown to black : left ramus broken at
hind of P 3 , detaching P 1 which is notpreserved; tip of rostrum with I and C
not preseryed. Ramus in poor preserva·tion, surface of which a little deformedby crushing: ramus straight and nothigh, lower border gradually ascending ,forwardly.
Cheek teeth moderately worn. andbrachyodont. with thick enamel walland \veil developed basal cingulum.
Ground surface eminently \V -shaped. P,much longer than wide, elongate trigonal
in upper view, becomes broader posteriorly and anterior lobe smaller thanposterior lobe: in outer view. anterior
lobe longer than posterior. subquadrate
in general outline. with obsolete basal
cingulum. straight anterior margin
vertical to alveolar border and withposterior margin bent backward : posterior lobe longer in upper margin thanin lower margin and with eminent basal
cingulum. In inner view, paraconid andmetaconid bent backward. while hypoconulid bent forward : they tip upward

and upper portion bluntly projected;
metastyl id obsolete ; posterior valley
larger than anterior. In upper view.
posterior lobe more strongly curved thananterior; hind margin gently curved ;paraconid and hypoconulid distinct while
metaconid weak in development.
P 3 subquadrate in upper view, longer
than wide, and a little broader posterior·ly: posterior lobe and valle~' larger thananterior. In outer view, anterior lobe alittle higher than posterior and slightly
bent backward. while posterior lobe
almost vertical to alveole: basal cingulum
well developed on both lobes and carries
many small tubercles: outer wall ofposterior lobe broken at left P3 . In
inner view. paraconid and metaconid
bent backward while hypoconulid verti·
cal to alveole: metastylid weaker thanmetaconid in development : paraconid
small. lowest. about half the height ofmetaconid ; hypoconulid eminent andbluntly projected upward. Protoconidand hypoconid strongly projected buccally but the former sharper than thelatter. Ground surface of hypoconulid
tolerably large and nearly quadrate.
P 4 largest and broadest of all teeth,
quadrate in upper view. with wide worn
surface which is more distinct in anterior lobe than in posterior: anterio·outer wall runs nearly straightly inanterior lobe, while gently curved inposterior lobe: posterior margin crenulated at inner side: posterior valley
larger than anterior. General aspect ofouter side is like that of P3 • but anterior
lobe almost as high as posterior: basal
cingulum thickest of all teeth and longer
in anterior lobe than in posterior. General aspect of inner side is also likethat of P 3 ; surface of enamel wallrugose: paraconid small. low and about
half the height of metaconid which islarge, eminent and higher than hypo-420. On a Equid Fossil from Hiramal<i Forma/ion
conulid: melastylid and metaconid not
well separated from each other: hypoconulid bluntly projected upward.
M1 corresponds to P, in size and general aspect and in development of basal
cingulum on outer wall, but contrary to
P •· anterior lobe a little broader thanposterior. In upper view. posterior lobe
more sharply projected outward thananterior: anterio-outer wall of posterior
lobe runs straight while that of anterior
gently curved : ground surface of anteriorvalley larger than anterior ; posterior
margin also crenulated at inner side. In
outer view, basal cingulum longer inanterior lobe than in posterior: anteriorlob~ as high as
posterior. General
aspect of inner side is like P 4 andsurface of enamel wall rugose; metaconid higher than hypoconulid. Anterior
half of inner enamel wall broken at left
lVI,. Ground surface of hypoconulicl quadrate and· as long as wide. :vletaconid
and metastylid not clearly separated.
M2 smaller than :VI,, quadrate, broader
anteriorly than posteriorly and posterior
enamel wall of anterior lobe straight
and wider than that of posterior lobe:

173

anterior lobe more sharply projected
than posterior: anterior valley as wideand long as posterior. In outer view.
both lobes bent backward and posterior
lobe higher than anterior: basal cingulum.
obsolete at posterior lobe. In inner view,
general construction is like that of M,
and metaconid higher than hypoconuli'd.
~letaconid and metastylid Yery faintly,
separated from each other.
\1 3 narrow. longest of all cheek teeth,
elongate trigonal in upper view and withsmall talonid; anterio-outer portion
ground; anterior valley larger than posterior: inner wall of hypoconid gently
sloped inward with sharp line or valley
bottom. In outer view. anterior lobe alittle longer than and as high as posterior
lobe; talonid blunt. carrying two small
accessory tubercles at its anterior basal
portion: basal cingulum obsolete.
In
inner view, general aspect of conids
except talonid is like that of M2 ; meta·
conid and metastylicl undivided; talonid
with median depression which is moderate in depth and outer wall largest;
surface of enamel wall rugose at metaconid.

j\1easurements :United length of right P 2-P4
Ditto of left M 1 -~1 3 . . . . .

•..•••••••••••.•••••••••.•. ........

53.0 mm
65.0

I\

p"
Right
Length at ground surface ...................... 20.3
Width at anterior lobe ........................ 10.8
Ditto at anterior lobe . . . . . . . . . . . . . . . . . . . . . . . . . . 13.8
Height of crown at outer side (posterior lobe).. 9.2
Ditto at inner side (metaconid) . . . . . . . . . . . . . . . . 6.8

M,
Right
20.6
15.0
1-1.1
9.0
9.0

Left
20.2

Right
21.0
15.0

10.7

13.9

15.7

10.2
6.7

10.5
8.3

---Me

Left
21.0

Right
19.7

H.O

12.3
10.8
9.9

1<1.0

9.8

Left
20.0
15.2

6.7

Right
21.2
16.3
16.1
10.0
9.7

Ms Jlolotypc

Left
Left Left P,
20.0 mm.
19.2
23.5
1-1.0
13.8
12.4
10.0
15.0
12.0
14.0
10.7
10.8
9.7+ 11.0

174

Tokio SHIKAM1l and Shinji YosHIDA

Remarks :-According to !\lA TSUMOTO's
study, the holotype consists of left P,(?) and right P• and the former is an
unworn tooth which is higher andsmaller than this specimen. lie saysthat this species is distinguished from
A. aureliane11se Cuv. of the BurdigalianVindovonian in Europe by higher basal.cingulum and from A. ezquerre V. MEYER
of the Vindovonian-Sarmal ian of Spainby its smaller size. Further. the \Vritcr
thinks that this species is distinct fromezquerre in having relatively longer andlower P., with more obsoletely separ·
ated metaconid and metastylid. When
COLBERT described Anclzitherium gobiense·CoLBERT from Tung Gur formation of·Outer i'vlongolia (\1 io- Pliocene), he reserved a comparison with this species
because it is very incomplete in materi·als. but he regarded that both are quite
near to each other. A. gobiense is alittle larger and has less developed basal
cingulum than this species. althoughboth species are quite like each other.
He regarded Pcmmclzitlzerium karpinskii

BoHISSIAK from Caucasus as belonged toParalzippzes. Hypo/zippus zitteli (SCHLOSSEr~)from the Pontian red clay of China is amore advanced form with less developed
basal cingulum of teeth and much larger
than this species.
References cited
E. (1939), A new anchitheriine
horse from the Tung Gur formation ofMongolia. Amer. 1lfus. Nov .. no. 1019.
pp. 1-9.
MATsntoro, H. (1921). Description of some
new fossil mammal from Kani District,
Province of Mino. with reYision of some
Asiatic fossil rhinocerotids. Sci. Repts.
TO/wlm Imp. U11i1• .. ser. 2. l'o/. 5. pp. 8592. pis. 13-14.
ScHLOsSER, 1\·1. (1924). Tertiary vertebrates
from Mongolia. Pal. Sin .. Ser. e. 11ol. 1,
fast:. 1, p. 68, pl. 5. fig. 10.
STtRTON. R. A. (1940). Phylogeny of North
American Equidae. Unit:. Cal. Pub. Bull.
Dept. Ceo/. Sci .. l'<Jl. 25. no. 4, pp. 165-198.
ZoA~sKY. 0. (1935). Equus und andere Perissodactyla. P.zt. Sin .. Ser. c. rot. 6. fasc.
5, pp. 17 20.

CoLBERT,

Explanation of Plate 26
Anclzitlwrium llypohippoides MATSUMOTO
Fig. 1. !Jpper side of the specimen. x 0.75
Fig. 2. Ditto of right teeth. x 1.05
Fig. 3. Outcrsidc oi ditto. x 1.05
Fig. 4. Inncrside of ditto, X 1.05
Fig. 5. Uppcrsidc of left P", Pa. xl.05
Fig. 6. Inncrsidc of ditto. X 1
Fig. 7. Outerside of left M 2 -M3 • X 1.04
Fig. 8. Uppcrside of ditto, X 1.04
Fig. 9. Innersi1le of ditto, X 1

SHIKAMA and Y OSHIDA :HASEGAWA photo

Equid Fossil

Plate 26

Trans. Proc. Palaeont. Soc. Japan. N. S .. No. H. pp. 175-181. pl. 27. Dec. 15. 1961.!21.

O.N SOrofE TRIASSIC PELECYPODS FROM
PAHA~GPROVINCE. MALAYA*AKIRA TOKUY AMA
Geological Institute. University of Tokyo-~*l¥.=tt1i'~iil!:
Pahang Q) Kuala Lipis it!!1J J: j) l£:lli L t~=tt1l-.P:.: 3 f.!!iQ):\lyoph'Jriids. :Veoschizodus /aerigatus elongatus. Alyop/wria golt{(ussi lipisensis subsp.
nov .. M. dr. costata Q)ftlJ Pteria, .. Plagiostoma" ~~!.iHIIJLf.:Q)"{';:;/1,(? ~llclli:Lt.:, ;:.ht:>i::t NEWTO:\ (1900) 7H~2li:Lt.:.l:$.::=:~;.f,IT.l Myoplwria ~!::t:l:I2UirJ~tl. . .Ptm.::=:~;f,ib'F-J-tQ)!::~;tG;fl. J::i:1;(7) NEWTON Q) t1fropyoria-Sandstone J: l);;ta_q(?IJ.:.:T{;J.Q),~ij!!(7) 'l,(7) t.;. (1)"{' • .: Q)'!fiftB~ :.:ttL. lower il!yop/wria-Sandstone Q)~ ~:f:ltPAA Lt.:., i'SJ~*:::t.~*.i!!!~it!!itiil.i!J!f.m J: 'J ·J, f;H'x~Q) r i.:l:& r) ht.:. ~ (7) "t' <b .'5,
i:f\i ill
a.J1

This is to report the result of studies
on two collections from l'vlalaya whichwere submitted to Prof. T. KoBAYASHI
from the Geological Survey. Federationof l'vlalaya. One from Lipis district 1.atlocalities between Benta and Lipis. andbetween Benta and Jerantut) inc! udes
ivlyop/wria gofttfussi !ipisensis and Neo·sclzizodus cfr .laerigafus. a pterioid. a limid
and an ·· Anodontop/wra ". The age ofthe faunule is determined at 1\liddle
Triassic rather than Upper Triassic.
The other from a branch of Ulu Sungei,
Kenong. Lipis District includes three
myophoriids and two indeterminable
shells. one of which resembles Hoemesia.
although the determination of the last
is not warranted. because the shell isstrongly compressed. Among the three
myophoriids one smooth form which ischaracterized by its posteriorly elongatedoutline and an acute carina, is identified
with Neosclzizodus laez•iJ;atus clougatus.
It is distinguishable from Upper Triassic
laevigatus from Japan i.KOBA YASIII &1949; KAl\IBE. 195]. 57) andIndochina (PATTE. 1926) in outline andcarination. The two ribbed species belong to the group of 111. goldfussi and
M. costa/a. They are distinguished fromthe NEwTo:s's (1900) Upper Triassic
species from Malaya. such as M. mala·yensis and M. inaequirostata. by the rib·bing and outline. These three myopho·
riids indicate the Triassic age and pro·bably !vlicldle Triassic rather than Upper
Triassic. Another r..liddle Triassic fossil
bed yields Anisic ammonites in the tributary of the same river (KUMMEL, 1960).
Here I record hearty thanks to Prof.
T. KOR\YASI-11 for his kind guidance
through the work. I am also thankful
to Dr . .1. B. ALEX.\:\UER. director of theGeological Survey. Federation of Malaya
and !VIessers H. SERVICE. J. R., PATON,F. Y. IIO:\C and other geologist of theof the survey who supplied the materials
and infonnations about the fossil beds.
ICHIKAWA,

Note on the Marine Triassic* Received 12th July, 1961; read at 79th

in Malaya:Meeting of the Society on 24th September.
1961 at Kanazawa.

The Triassic rocks are widespread in

175

176

Akira TOKUYAMA

southeastern Asia. Terrigenous facies
is developed on the continent but calcareous and siliceous rocks are common
in the archipelago. In the Malay peninsula marine Triassic beds have beenreported as follows :(lJ NEWTON (1900) described Ch/amys
Mlouiensis. Pleuroplwms elougatus (?},
Pteria pahaugensis. M_vtilus cfr. minutus.
Geroillia iuflata. Pteropema malayensis,.Myoplwria malayensis, ill. ornata. M. ina·
equicostata. M. sp. and Adinodesn;a bef.
lamyi from Lipis district. Comparing
this fauna to the Napeng one (HEAI.Ei'.
1908), he considered it Upper Triassic or
Rhaetic. On account of the prevalenceof J1Jyoplwria he proposed ·· Myoj>horiasandstone" for the bed.
(2) NEWTON (1923) reported 17 lamellibranchs including .M_vophoria ornata, M.
bittneri and Chlamys cfr. 11a/onieusis from
Shingapore and considered them also
Upper Triassic.
(31 Later (1925) he described Flalobia
malayeusis. H. moussoni and a halobiid
beside an indeterminable ammonite from
Kedah and Perak provinces. The Ilalobia-shale is 10\ver than the Myophoriasandstone and probably l\'liddle Triassic.
(4) WEIR (1925) reported another occurrence of the Jltlyop!wria-bed or theJ11yop!wria-sandstone in Pahang province.
(5) SCRIVE<\OR (1931) added 2 otheroccurrences of Jl.fyop/10ria-sandstcne inPahang beside an occurrence of Daonel/alike shell in a shale of Kalantan.
(6) Geologists of the Geological Survey of Malaya made a collection of myophoriids and other pelecypods from t\volocalities near Kuala Lipis. \Vhich weresent to Dr. Cox of the British tvluseum.
He compared the faunule to those of the
f't.Jyophoria-sandstone.*(7) Recently pelecypods were discovered from the head waters of theri,·er Ulu Sungei Kenong. According to

ALEXANDER** they are known from two
localities. Dr. Cox who examined a collection recognized H pelecypods including "Gervil!ia ". pterioids. Mysidioptera,
Myoplwria and others and considered thefaunule to be Upper Triassic. Anothercollection was made at a small stream
on the eastern bank of Ulu Sungei,
Kenong. 2.5 miles ss·w of the above
locality. The materials are submitted
to Prof. KOBAYASIII. They are now described.
(8) Km.IMEL (1960) described 4 ammonites (Paraceratities trinodosus. Sturia
sansol'inii. Acmclwriceras sp. and Ptycitessp.) from a branch of the river Sungei.
near Kuala Lipis. The age is determined
at Anisic.
In summary. several Triassic fossil
beds in l'vlalaya are presumably in the
succession as follows:l.

(Upper) Myop/wria-sandstone by :\Ewat sev~ral localities in Pahang andSingapore . . . . . . .
. .. Upper Triassic
Pte1·ia sandstone by Cox (MS) in Pahang .................... Upper Triassic
Halobia moussoui shale in Pahang by
NE\\'TO" .... Middle or Upper Triassic
Daonella shale in Kalantan by SC!U\'£·:--:oR ................•... Middle Triassic
Lower ;\fyop/wria sandstone in Pahang.. .. .. .. .. ..
. .. Middle Triassic
Ammonite bed in Pahang by KL;~I~IEL........................ Middle Tria~sic
TOK

2.
3.
4.
5.
6.

Lower illyoplwria sandstone

The lower 1\1yop/wria sandstone includes Neoscliizodus laevigatus elongatus.
and Myoplwria goldfussi !ipisensis. new* Personal communication from Mr. Harold
SF.R\'ICF. to Prof. KOBAYASHI. (GS 53/Q/083/
15).

** Dr. Au:x.\;o.;DER sent information ;;.nd
manuscript by Dr. Cox for reference (GS60/F /026/10).

421.

On some Triassic Pelecypods from Pahang ProtJince

subspecies. These two forms represent
a lower horizon than NEWToc-;·s Myophoria sandstone which yields J/. ina·
equicostata and allies such as i\1. malayensis, M. !tarpa and M. bittneri. The
dongatus-/ipisensis association is allied
to the !\Iiddle Triassic fauna of Yunnan,
where Loczy (1898) described .H. mdiata.
.H. szechettyii and N. cfr. !aet•igatus. and\L\:"\SUY (1912) reported N. lael'igatus and
M. "radiata" (=M. go/dfussi !ipisensis.
new subspecies. which is distinguishedfrom M. radiata s. s.). .V. elongatus isallied to the one described from I he
Dolomiten by OGIL\"IE-GORDO" 11927l.
The elongated laez•igatus \Vith an acutecarina is known in the Ladinic of south
Israel I LERL\!At\. 1960! and Trans- Jordan<Cox. 1924. 32). They nrc distinguishable
from the so-called lae~·iga:us in the Upper Triassic of Tonkin and Japan by
the outline and carination. Lipisensis
belongs to the group of goldlussi. It isdifferent from inaequicostda in the number and mode of ribbing. Because it is
known that the former flourished a lillie
earlier than the latter. the !ip:sensis·
elougatus horizon is probably older thanthose of inaequicostata and malayensis. It
me~ns that there are at least two Myop/zoria beds in the Triassic of \lalaya.
Therefore .. lower ,11yophoria sandstone"
is proposed here for the sandstone containing !ipisensis and clongatus.
According to Foo Yew HOC\G* the bed
is about 2 ft thick. steeply dips southwest (22J• /60°) and crops out at a small
stream on the eastern bank of Ulu Sungei Kenong in Yong Forest Reserve,
Lipis District.
The sandstone is dark grey. but when
weathered, it changes to white or yellowish brown. It contains sometimes* Pers>Jnal communication from Dr. ALEX·
A:"DERto Prof. KoBAY.\SIII.

177

large. smooth and rounded nodules ofargillaceous matrix. The sandy grainsconsist of rounded quartz and chert. but
almost devoid of feldspars. The marrix
is somewhat cherty. It is interesting
to see that laevigatus is somewhat ori·
entated in a certain direction. The factsuggests currents of water when thesandstone was accumulated. In Japan
Myophoria facies is more off-shore thanthe "trigon ian" facies. because the !l!yophoria sandstone is more currentecl thanthe JHinetrigonia-sandstone. It is supported by the fact that the Minetrigonia·
sandstone is found often in the paralic
Mine facies. but _il,finetrigonia is uncorr:com in the neritic Kochigatani facies(TOKlJYAMA. 1961). Myop/wria is, on theother hand. absent in the l'vline but common in the Kochigatani series. The
Myoplwria sandstone in :\lalaya is probably also neritic rather than littoral orparalic. because the sandgrains are composed or well-rounded quarz and chertand shells are controlled by oceanic currents in the thanato-history of the fossils.
The sandstone along the Ulu Sungei
is strongly deformed and somewhat
crushed (so-called B-Tektonite by \•IETZ.
1957). Fossils preserved in the sandstone
are strongly compressed.
The lower .1/_vophoria sandstone yields
the following species.
a) Ulu-Sungei collection:11yop/wria go/d[ussi lipisensis. new subspecies.
Jlyop/wria sp. ex gr. mstata ZE::->KER.Veoschizodus lmw'g:Jtus elongatus (PHI·
LIPPI)

Hoemesia (?) sp. indet.
b)Collection from a point
Benta and Kuala Lipis
Pteria sp. indet .
.. Plagiostoma" l?) sp. indet.

Afyoplwria gold[ussi !ipisensis

between

Al?ira ToKUYA.HA

178

Neoschizodus /(l(migatus e/ongatus
Anodoutop!wra cfr. trapezoidalis MANStlY
Description of Fossils
Genus Neoschizodus GIEREL.l''~leoschizodus laevil[afuselonl[alu s PHI 1.1 I' PI
Plate 27. figures 1-6
1898. Myopltoria tae11igata Yar. elmrgaia PHI·
LJI'I'I, p. 165. pl. 6. fig. -1 (in Schmidt,
1928. !>- 185, fig. 425).
1927. ;l!yophoria faer•igato var. efo11gata, OGIL·
VIE-GOLDO:". p. 33. pl. 3, fig. 3a, b.

DescrijJ!ion :-Shell medium in size.
subquadrate in outline, longer than high;anterior and ventral margins evenly
rounded; posterior margin somewhat
alated.
Umbo prominent, subangulate
and a little incurved. Carina sharp andpersistent.
Posterior area flat anddevoid of a median depression. Discsmooth. covered with concentric growthlines; a sinus weak and broad if present.
on its posterior part.
Obsemations :-Specimens are preservedin phyllitic sandstone and so strongly
deformed that it is hard to restore theoriginal form. It is however a remarl<a·
ble fact that most specimens are compressed dorso-ventrally. and more lesselongated antero-posteriorly. This suggests that the shells have been orientatednearly in the same direction before they
were compressed. If so. the orientation
may be related to bottom current. inview of the fact that myophoriid shells
are sometimes similarly orientated in the
Rot of l'vluschelkalk in Germany (V. FimY·
BERG, 1932).
Comparison:- Neoschizodus lael'il[alusis a well-known and widespread species.
In the German Trias it ranges from theRot to the upper ~vluschclkalk and its
age is nearly same in the Indo-Pacificregion. Its variation in outline is fairly
wide. It is however a general tendencyof the species that the carina, which isstrong in the Lower Triassic forms, becomes weaker in the later stages. Consequently the outline becomes higher
and more rounded. In the Lower Triassic form from Balatonsee (FRECH, 1904)
the carina is sharp and distinct and theposterior sinuation of the disc weak as
in l\lalayan form; FRECH's laevigatus has
a relatively higher outline than the
1\1alayan form. The lower Ladinic /aevi·
gatus from Trans- jordan (Cox. 1932! andsouthern Israel (LER:O.IA~. 1960) resemble
this form in the more or less elongatedoutline and alated posterior margin. although the carina is weaker than this.
Anisic /aevigatus var. e!on}!afus by PHI·
l.IPPI (1898. in SCH:\IIDT. 1928) represents
an elongated form carrying a more or less.
sharp carina. It is. however. not alated
postero-dorsally as in the T\lalayan form.
Laevigatus elougatus from the Dolomitic
series of southern Tirol (lower l\1uschelkalk) bv OGILVIE·GORDON (1927) is theclosest ·ally to the T\lalayan form. According to hEr .. elongatus" bears a moreor less sharp carina and an acute posterior angle. and elongated laevigatus
with rounded posterior angle is thought
to be an elongated variation of oJ•atus.
Similar /aevigatus is also reported fromYunnan (LoczY. 19981.
Finally. laevigatus is reported from the
Lipper Triassic of Tonkin IPATTE. 1926),
Yunmn (MA!':StlY, 19121 and Japan (KO·
B:\Y:\SHI & lCIIIKAWA. 19-19. from the
KochigaLani series: KA.\IBE, 1931. 57 from
Shidaka series! beside ~Iiddle Triassic
of japan 1N AK.\ZA \VA. 1960L Except for
the Lower and Middle Triassic forms
thev arc all rounded and belong to thehigh form of the oMfus-group.

421. On some Triassic Pelecypods from Pahang Province
Ocmrrence :-Common in the argilla·ceous sandstone at Ulu-Sungei Kenon.
Yong forest reserve. Lipis District, Pahang and rare at a point from the 3rd
milestone on the road between Kuala
Lipis and Benta. Pahang (no. 14820a).
Genus J1yophoria BRO);N

Myop/zoria gofdfussi /ipisensis.
new subspecies
Plate ~7. ligures 7. 8. 11-20
1912.

Myoplwria radiata. MA.--;scY. p. 121. pl.
22. fig. 2a--f.

Description :-Shell small. about 10-13
mm long and subtriangular : umbo stout
and prominent: carina strong and acute.
Disc covered with 6-8 primary costae,
which are weak. slender and narrowly
spaced in anterior part, but strong andwidely spaced in posterior; the most
posterior one so stout that the posterior
margin of disc becomes somewhat sinu·
ated. 1-2 secondary costae inserted inposterior 1 or 2 interspaces. Area divided
into 3 crescentic parts by a fairly distinct rib and furrow.
Obs('/'l'llfions :-Shells from Ulu Sungei,
Kenong arc strongly compressed as thepreceding, but those from Loc. No. 11836,
between Benta and Jerantut show ori·gina! outlines (figs. 8. 18;. As noted
above. shells of laez•igatus seem to beorientated in a direct ion at the locality.
but it is not the case of this occurrence.
because the shells are deformed in various ways (figs. 12-161.
juvenile forms (figs. 19. 20) have 2
primary ribs beside 3 secondry ribs. inserted in their interspaces. Ribs increase
their number by insertion. instead ofbranching. Therefore the shell grows
through the kefersteini·like stage.
Comparison :-It is identical to J1.
radiata by rviAl\SUY (1912. p. 121. pl. 22.

17~1

figs. 2a-f), but somewhat different fromradial a s. s. Loczy (1898, p. 155. pl. 9,
figs. 21. 22). Radiata is characterized by
the smooth and fiat area and sharply
edged carina. but MANSUY's bears a dis·tinctly costate area. and somewhat alated
outline.
J.V!yoplroria with a ribbed area is repre·
sented by goldfussi, which possesses
more numerous ribs on the disc and area
than this. HOHENSTEJ!'\'s goldfussi fromthe middle Muschelkalk of east Schwarz·
wald bears 11-13 primary ribs but theribs are commonly countable 15-25 ingoldfussi. i\1. inequicostata is allied tothis in the number and mode of ribbing.
but inequicostata is distinguished fromthis in having somewhat imbricated con·centric growth-lamellae, which are ab·sent in this. 111. malayensis and allies
by NEWTO~ (1900) belong to the groupof inequicostata. Finally, this is a varie·
tal form from goldfussi rather than ani nequicostata.
Ocwrrence :-Abundant at the two
localities of Lipis district.

klyoplwria sp. ex gr. costa/a ZEl\1\ER
Plate 27. ligures 9, 10

Two small specimens are distinguishedfrom the preceding by their rounded
outline and flattened area. It is not soinequicostate as the preceding. It belongs
to the group of either mstata or radialarather than the goh~russi-inequicostalrt
group. although ribs are fewer in thisthan in costa! a.
Occurrence :-Rare at the two localities
in Lipis district.
c;cnus Pteria SCOPOI.I

Pteria sp. inclet.
Plate 27. figure 23

A small left internal mould is trape·

Akira TOKUYAMA.

180

zoidal, alated posteriorly; hinge line
long and straight; umbo large and anteriorly inclined:. ventral margin somewhat quadrate. In outline and hinge it
agrees with non-ribbed Pteria common
in the Alpine Triassic such as P. /iof·
mmmi BITT:\ER (1901), although theumbo is large and not so elongated posteriorly in this as in hofmanni. Because
the specimen is juvenile. the umbo isrelatiYely large.
Ocwrrence :-Rare at Joe. 14820 bet ween
Benta and Lipis.
Genus Plagisotoma SOWERBYPlngiostomn i?) sp. indet.
Plate 27, figure 21

A small internal mould of a left valve
and a few fragmentary external moulds
are preserved in mudstone. They aresuboval, 1.5 times longer than high androundly inflated: beak fairly large androunded: anterior auricle narrow, elong·
ated trigonal, and fairly distinct: pos-terior one small, less distinct than anterior one. Shell ornamented with fineradial ribs or striae.
As commonly met with among Triassic forms, this bears a fairly large anterior auricle. The auricle agrees withthose of common limids or Limatula(TOKUYAMA, 1960), but the ornament isallied to those of Plagiostoma. Pla~tios·
toma is considered to be derived fromthe :VIiddle Triassic branch of .l!ysidioptera. which the latter has no such auricle
as this. The auricle suggests its beinga derivative from Limntula or Palneo!ima,
but in the ornaments this form disagrees
with these genera.
Occurrence :-Rare at Loc. 1-1836 near
Benta.
References cited
ALEXAXDER. ]. B.

(1956): lVlalaya in •· Lexique stratigraphique international "vol. 3.
Asie. fase. 6h. 33 pp.
HITTNER. A. (1895): Lamellibranchiaten deralpinen Trias. 1 Theil : Revision derExplanation of Plate 27
Figs. 1-6. Neoscllizodus laet•igatus elongatus (Prm.rPPr) .................................... 178
1-4: Internal moulds: Joe.: Along a small branch of Ulu Sungei, Kenong. X 1. 5: Internal mould of a right valve. from Joe. 14820 3rd milestone on the road between Kuala
Lipis and Benta. Pahang. 6: External clay cast of a left valve, from Ulu Sungei. Kenong.
Figs. 7. 8. 11-20. J/yop/wria goldfussi liPise1zsis nov ......................................... 179
fig. 12: holotype. left valve: 8: left internal mould showing interior. 12-17: show various deformed shells. 8, 18: from Joe. 14820a, others from Ulu Sungai. X 1. 5 (19: x3,
18a: x 1).
Figs. 9, 10. Jlyoplwria sp. ex gr. costata ZEXKER ...........•...•.•.............•..•.•..... 179
9: right internal mould. Joe. 14836 at the 33 1/2 milestone on the road from Benta to
Jcrantut. 10: left valve from Ulu Sungei, xz.
Fig. 21. Pfagiostoma ( ?) sp. indct ........................................................... 180
Left internal mould from Joe. 14820, x l. 5.
Fig. 22. ~l11odo11toplwra cfr. trapezoidalis MA:-!SUY . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 178
Left internal mould, from Joe. 14836. xl. 5.
Fig. 23. Pll'ria sp. indet. ................................................................... 179
Left internal mould. from Joe. 14820a, X 1. 5.

T OJWYAMA ;Myophoriids from Malaya

Plate 27

3

2

4

5
8

12

lOa

9

13

14

19

17

15

18 h

20
18a

21

22

23

421.

On some Triassic Pelecypods from Pahang Province

Lamellibranchiatcn

von

Set.

Cassian.

A.M. k.k. geol. R.·ll. /Jd. 18. Heft I. ~3li S.
~I Taf.
- - (1901):
Lamellibranchiaten aus der
Trias des Bakonyerwaldcs Res. \viss. Erf.
Balatonsees Bd. 1, Theil 7. 107 S. 9 Taf.
- - (1891): Triaspctrefacten von Balia in
K lcinasien. fb. k. ll. xeo/. R.·A. 41. Bd. 1.
Heft, S. 97-116. Taf. 1. 2.
Cox. L. R (1924): A Triassic Fauna fromthe jordan Valley. Ann ..l1ag. iVai. 1/i.~t ..
ser. Y. z•ol. 14. pp. 52-96.
- - (1932): Further Notes on the Trans·
jordan Trias. Ibid .. ser. 70. l!ol. 10. pp.
93-113. pl. 7.
FRECH. F. (1889):
Ubcr Mecyt~odoll undM)•ophoria. Z. deutsch. geol. Ges. Bd ..J 1.
S. 127-138. Taf. 11.
- - (190-1): Neue Zwcischaler und Brachia·
paden aus der Bakonyer Trias. Res. tl'iss.
H1j. Balatonsees. Paliimzlol. Abh .. I Tl., I
Bd .. 1.JO S.
Y. FREYilERG, B. (1932):
Die Rot-1\luschcl·
kalkgrenze bie Eisfcld (Thiiringen) und
im Schalkauer Muschelkalkplateau. (:eo/.
u. i'a/iioutof. Jll>h. N F. Hd. 19, S. 203-250,
Taf. 10-12.
HoHENSTEI:--1. V. (1913): Beitriige zur Ken·ntnis des mittleren 11uschelkalks unddes unteren Trochitenkalks am ostlichen
Schwarzwaldrand. Ibid .. N. F., /Jd. 12,H(if/ 2, S. 173-271. Taf. 12-20.
KA~rur·:, N. (1951):
On the myophorians
from Kyoto Prefecture (Prov. Tango).
Tr. Proc. Palaeo11l. Soc. Japan, .V. S., No.
2. pp. 49-56. pl. 4.
- - (1957): On the myophorians from theMiharai·yama group in Hyogo Pref. Rep!.
geo/. Sun•. japan, JVo. 173. 22 pp., 1 pl.
KoBAYASHI. T. & K. IcurJ.;A\\'A (19,19): Myophoria and other upper Triassic pelecypods
from the Sakawa basin in Shikoku. Japan.
]apa1t. f. Geol. Geogr., z•ol. 21, nos. 1-4. pp.
177-187. pl. 6.
Ku~r~IEL, B. (1960):
Anisian ammonoids
from .l'vlalaya. Breoiora. No. J2,J. 8 pp., 1
pl.
LER~JAI\. A. (1960):
Triassic pelecypods
from southern Israel and Sinai. Bull.
Res. Council Israel. Sec. G, rol. 9G, no. 1.

181

pp. 1-60, 4 pis.
•:. L6czY. L. (1898): Beschreibung der fcs·
silcn Saugethicr·Trilobiten und ;\lollus·
ken-Reste und die pahiontologisch-strat i·graphischen Rcsultatc der Reise des Gra·
fen Bela Szcchenyi in Ostasien. 18771880. IViss. Erg. Reis Grafeu JWa Sze·
dw11yi ill Ostasien, 3 ]Jd.. 6 Abt.. 224 S.
11 Taf.
NAKAZAWA. K. (1960):
Permian and Eo·
Triassic Myophoriidae from the Maizuru
zone, southwest Japan. ]apa11. f. Geol,
Geogr., t•ol. 31. 110. 1, pp. ,19-62. pl. 6.
NE\\'TOI\, B. (1900):
On marine Triassic
Lamcllibranchs discovered in the Malay
Peninsula. Proc. J1alac. Soc .. rol. 4. pt. 3.
pp. 130-135. pl. 12.
- - (1923): On Marine Triassic shells fromSingapore. Ann. Jlial(. Nat. Hist.. ser . .'),
t•o/. I 2. pp. 300-321. pl. 9.
- - (1925): On j\Iarinc Triassic fossils
from the Malayan Provinces of Kcdh andPerak. Geo!. Mag., t•o/. 62. pp. 76-85, pl.
3.
MA:-<suy, H. (1912): Etude geologique duYunnan oriental. Pal&ontologie. Mem.
sen•. Jdul. Indochine, t.•ol. 1,fasc. 2. 147 pp.,
25 pl.
PATTE. E. · (1926):
Etude paleontologiques
relatives ;i Ia geologie de !'est du Tonkin(Paleozoique et Trias). Bull. sen1, geol.
de l'/ndochine. z•ol. 15, jasc. 1, 204 pp., 12
pis.
SCHMIIJT, M. (1928): Die Lebewelt unserer
Trias. 461 S.
ScRIVEl'\OR, J B. (1931) : The geology of
Malaya. 217 pp.
TOKUYA~It\, A. (1960):
Late Triassic pelecypod fauna of the Aso formation inwest Japan. JaPan. f. Geol. Geogr., ool. 31,
110. 1. pp. 23-38. pl. 4.
- - (1961): Late Triassic Facies of Japan.
Ibid .. uol. 32. IW. 2. pp. 279-292.
WEIR. ]. (1925): On some specimens of fossiliferous sandstone from Pahang, Malay
peninsula. Ceo/, Mag .. r.•ol. 62, pp. 347-350.
v, WbJJR~IAl'\N (1893): Ueber die systematischc Stellung der Trigoniden und Abstammung der Xayaden. ]b. k. k. geo/. R.·.4., Bd. 63, S. 1-28. Taf. 1.

182
PROCEEDINGS OF THE PALAEONTOLOGICAL SOCIETY
OF JAPAN
13 ;f>:?r't*'~:~~ so @lf§!J~~:t. 19611¥11 fl17.
18 A:tL#I k't.I!!\:¥:Si:Jt!!'Ct~¥tf~'.:.:t,;l.- 'Lf#Jftl! ~ .t<,f.:_o (~ft1i 34

;g)
J]Jj

............ f.li:~fl':HF.
... · · · · · · ..... ~ffllft?k::fiJRQ)fjj(;f,J;li::.~-;-7.:>1 irr~

:IL:J-Iii.:.:~lt 7.:> rf 1 9.!:~~~A
Uifit!.tt!!Jifi1J ·1..-~J:lE!lffiFJ(~Q):i'{rjf.Jt!! (f-\:WJt)................................ E~l'lll~
Fusuline fauna of the Middle Permian Kozaki formation ............ K. KAK~JEJU
On some Jurassic Coelenterata from Eastern
Aomori Prefecture (ml;ft) .. M. 1\IuRATA
Additional notes on late Palaeozoic Corals
found in the Southern Part of theTamba District (f\:W'D ............... .
........ s. SAKAGt.:CHI & N. YA~IAGJWA
Lower Carboniferous Bryozoa from the
Ilikoroichi Series, japan.
Part 1:
Cyclostomata and Trepostomata (f\:~)..................... Sumio SAKAGA~JI
Lower Carboniferous Bryozoa from the
Ilikoroichi Series, Japan.
Part 2 :Cryptostomata ((\:~) .. Sumio S,\J<AGA!\!1
New Syringothyrids from the lowest
Carboniferous of the ?\agasaki District. Kitakami Mountain land ....... .
...... , ................... K. Tc\CHiflA:"A
Upper Permian Brachiopods from the Mizu·
koshi Formation ............ ]. YANAGIDA
Fossils from the Sh6zugahama formation. . . . . . . . . . . . . . . . . . . . . . . . . . Tsugio SmJTo
On the Megafossil fauna from the Environs
of Higashi-l\'fatsuyama City. Saitama
Prefecture, Japan (f\:;!)1;) ........ .
...... Kotora HAT AI & Kochiro MAsurM
Variation and Sexual Dimorphism of Certain
Non-Marine Gastropoda .......... .
.............. T. KoTAI<A & M. Uozu~!l_ucinacean Pelecypods from lhe l\IiyazakiGroup . . .
. .......... Tsugio St!liTO:lln?:m:n.111ll~Jl!!gjE:*i!¥. F.L fi:T:illf:.:. --:>1,., -c <f{;~>............................ 'l§'!ff.=:ru:

Serripes from Japan and Saghalien (ftltil:). .............................. H. XODA
Some Pelecypods from the Akaiwa subgroup. the upper division of the Tetorigroup, Central Japan (f~itfD ......... .
. . . . . . . . . . . . . . . . . . . . . . . . . . . . Shiro MAEDA
Zonal distribution of baculitid species in
the Cretaceous of Japan .............. .
. . . . limo On.'.TA & Tatsuro MATSU~tOTO
Notes on some ammonites from the Ilimenoura group . . . .
. ............. .
.. Tatsuro MATSU~toTo & Yoshiro VEDA
1-U P !!jl;Jj'(iJ:k:(iJJEft:O 7.71 .:r. t"f$il.:.--:>~ ·"L .. 1ffj..!1!, 7...
Fossil Temnopleurids from Japan ....... .
...................... Shozo NISHIYA~IA~timJ:~-t-11111Eilff-<=1 'l '1-fi::G . .JJr.-"' ¥-tl:!ffi • IMl~m~
3ttLM; ~~* J: t) Spongiomorphoides (l):J"cji!,....................... ·filii* £[. :&~.Ji!fDie sogennante .. Ficus" titiaefo!ia (AI.. BR.)
IIEEtl. ..
. .... Kiyoshi TAK:\HASHI
Pollen Grains and Erect Stumps from theSubmerged Forest of Asahi-mochi,
Toyama Prefecture. Central Japan (ft~) . . . . . . . . . .
. ...... Norio FuJI
Palynological study of the Miocene V.-'akura
diatomaceous Mudstone formation ofr\oto Peninsula. Central Japan (ft~) ..
.............................. Norio FuJ£::1 "'~"'?A

r ~~1<~I.:.OOi"oti~~~:'f:a91VfJ"EJ•••w~M~*Q)~~-~~:OR.WtQ)~~ ((\;;R) . · · . · .... ·. · · · · . · . · .... -11!li~FH~~tifi!Jt!..Pffrffl:'* m.~ Q):(E~0-tfri~~I'Vf~ .. f!i:~~n.J!!!'f'ifrlli:Q)-Il~j!fli;:j,;(t 7.ll:l(l ~J1!!1JQ)I!i Jt!! :Ill-~
I.:.t'L.r?tax~Q)J~fi!t~!~J::.--:>1,. ,-c- .. 7i!im:l!ll.

Terebratalia .~->m tbc
Oligocene Sari sandstone in the Karatsu coal field of Northwestern Kyushu.
Japan .
. .................. Eiji hou~:ttm :ltii'i rm kKm t.:.J<>!-t 7.:> l -:b~J>Zl ,j·- :1 'J ·:; !7

A new species of

7-7f\'Q).!W;*'fl:11~. . . . . . . . ...... £1f.m~~j1lj [] *.lt!!.!lilt !:.:to It -'5 It m::::~ .l't~Ji ll'jl(l)a!j-fi;it.J?!r.............................. 7J,!!!fJlHr
fr!f:0 ~xQ)1Efr}"f:lilfJct.:.:t,;;t o 1ffirn,lm ...... {~7Xlli:Jt~

Jifii3*1~:#E~~IiRmi::..li!Gh7.:>:fl&*'ffi· ,::H~

Article 19.
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Article 22.
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Addendumshall call a Special Meeting at the written request of more than one-third of
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Members unable to attend the General l\.Yeeting may give an attending member
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The President and Councillors shall compose the Council. The dicision of theGeneral 1\Ieeting concerning administration shall be considered and implemented
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The Executive Council shall carry out the decisions of the Council.
The fiscal year of the Society shail begin on the first of January ea~:h year andend on the thirtyfirst of December of the same year.
The amendments to the Constitution of the Society shall be decided at theGeneral Meeting and must be approved by more than two-thirds of those mem·hers who are in attendance.
1) Voting in the Council shall be by unsigned ballot.
(1961, Jan. 15)--------I

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CONSTITUTION
of the
PALAEONTOLOGICAL SOCIETY OF JAPAN
Article 1. The Society shall be known as the Palaeontological Society of Japan.
Article 2. The object of the Society is to promote the study and popularization of palaeon·
tology and related sciences.
Article 3. The Society. to execute Article 2. shall undertake the following business:
1. Issue the Society journal and other publications.
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