a -tt~~i.MJ~~~~·ic.lJ
Transactions and Proceedingsof thePalaeontological Society of Japan
New Series

No. -!1

Palaeontological Society of japan

April 10, 1961

CONTENTS
TRANSACTIONS
Page

39·1. The Miocene Molluscan fossils from the Area Southwest of Ilirosaki City,
Aomori Prefecture, Japan ................................ Takehiko lwAI
395.

1

A New Palaeogene Species of the Genus t:ucomma from Hokkaido, Japan. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1\.azuo HuzioKA

93%.

A Subfossil Cuon from Ang Thong. Thailand ................ Fuyuji TAKA I 13

397.

Foraminifera from the lmozawagawa Formation in Sendai, Japan ......... .
. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Naoaki AoKr

398.

15

Mesozoic Plants from the ltoshiro Sub-Group, the Tetori Group, Central
Honshu, Japan ........................................ Tatsuaki KIMURA

~1

399.

T\'·/O new Species of the Parafusulina yabci type from Tomuro, Totigi Pre33

400.

fecture. Central Japan ...... Rukuro J\1oRrKA "'" and Yoshinari TAKAOKAA New Species of Miocene Gastropoda (iVIollusca) from the Ginzan HotSpring, Yamagata Prefecture .......... Kotora l-lATAI and Tamio KoTAKA

41

Address to the Twenty-fifth Anniversary of the Society . . . . . . . . . . . . . . . . . . . . . . . .
PROCEEDINGS . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .

45
47

Honorary President : 1-Iisakatsu Y AilE
President : Teiichi KonA Y AS III
Councillors:

Kiyoshi AsA!':o (Editor of " fossils"), Riuji ENDO (Planning), 1-Iaruyoshi

FuJr~IoTo,

Shoshiro H.\:-<ZAWA. Wataru HAsHIMoTo (Treasurer), Kotora HATAI,

lchiro IL-"YASAI,,\, Koichiro lciiilc\wA, Nobuo II\EI3E, Tatsuro MATsUMoTo,
Masao Mr:-;.\To, Tokio S1111'AMA (General Affairs), Teiichi KollAYASIII, Fuyuji
T.\K,\1 (Editor). Ryuzo ToRIY,\:O.L\
Secretary:

Tctsuro I lANAI

Assistant Secretary : Takeo Icm KAw A

The fossil on the front page IS Xenvxylon latiporosum (CRAMER) GoTIIAN 1910

All Communications relating to this Journal should be addressed to the
PALAEONTOLOGICAL SOCIETY OF JAPAN
Geological Institute. Faculty of Science, University of Tokyo, Japan

Trans. Proc. Palaeont. Soc. japan. N. S., No. 41. pp. 1-8. pl. 1. Apr. 10. 1961

:39-!.

THE MIOCENE MOLLUSCAN FOSSILS FRO~I THE AREA
SOUTHWEST OF HIROSAKI CITY. AO?IIORIPREFECTURE. JAPAN*
TAKEHIKO IW AI
Department of Geology. Faculty of Education, llirosaki University'i!.iiiimffii!!!ffi:i:.Jei! L- -n -.ofiP'!JilRt <<t>l'IT~) •=4-lillf1-Ht.011)£.Jt!!l1)·)t:, 13 llilifflb•0 35 ~52 f\1!
(f)il\tt¥mb~a-J:l'(m-t .o ~ e: b:t±l*t-:11)-e. -t:IT)fJJ~fif(f)JtHrt~ e: IT)i)g*. P¥f\:~~:&V'.Mlta­
rr-?t.:...
::5 :Jt: .u~ ~'i!.iiiiml~ilf!ffll(f)<Pttft!!:W:tUu~ftG:•t~.oe:.:.-?•.:.Mt:ob:f!;ttt·n-7.>.

Here the writer wishes to express hishearty thanks to Professors Gunjiro
SAKAr of the Hirosaki University in
Aomori Prefecture and Kotora HATAr of
the Tohoku University in Miyagi Prefecture. for their constant ad\·ice andsupervision during the course of thepresent work. Acknowledgements arealso due to T\Iessers. Koichiro l\L-\SL'DA.
Shozo 1-L\YASAKA and Tetsuo SuroBARA.
and Drs. Taro KA:--;AYA and Tamio Ko1AKA. all of the Tohoku University for
their support in many ways.

Introduction andAcknowledgements
The marine molluscan fossils uponwhich the present article is based comprises the collect ion of the present writerduring his geological surveys in l%!1 to
1958 in the area southwest of Hirosaki
City. Aornori Prefecture. where l\'eogene
deposits are \'ery well developed but
hitherto little studied. The treated fossils are from the Ainaigawa formation(T. K\:--;.\YA. l~H9) and comprise the collection made from a total of 1:1 new
fossil localities of the formation.
In this article the writer. from thedetermined fossils of 13 fossil localities
and stratigraphic position of the stratigraphic unit which haYe yielded 1hem,
attempts correlation of them with previously known geological formations ofJapan. their bearing on the geologicalage detenninat ion. and a short discussion
is presented concerning the Ainaigawa
formation.

Notes on the Geology of the
Ainaigawa Formation
The Ainaigawa formation. first introduced by T. KA:--;,\ Y _\ '.19-19 as a stratigraphic unit. has its type locality at
Ainaigawa. Soma-mura. 1\aka-Tsugarugun. Aomori Prefecture. where the rocks
are well developed. This designatedlocality lie~ approximately in the central
part of distribution of the formation.
With regard to the formation. T. KA:--;AYA (1949) described as follows.
The Ainaigawa fom1ation is the lowest:
part of the Keogene Tertiary strata developed in the southwestern part of the* Received Jan. 23. 1960: read at the Annual lVIceting of the Society at Sendai Jan.
17. 1960.
1

Takelziko I w:. u
Hirosaki structural Basin. and comprises
mainly green. bluish gray and darkcolored tuffs and tuff breccia. The upper part of the Ainaigawa gradua!iychanges from the lower into fine grainedmaterials (light blue. grayish yellow andgray tuffaceous shale). and the lower
part is dominantly of conglomeratic
sandstone (granule size pebbles derived
from andesite and Paleozoic rocksl. In
the eastern part of the Tochinai-gawano volcanic materials are found. The
western extension of the formation ischaracterized with basaltic andesite flows
with minor amounts of conglomerate andsandstone. The thickness of the formation is estimated at 300 to 100 meters intotal.
The Ainaigawa formation unconfor-

Text-figure 1.

mably overlies the pre-Tertiary basementof granodiorite and Paleozoic rocks.
The upper part of the formation gradually changes into the superjacent Owasawa
formation, which comprises an alternation of green tuff and hard shale.
The Ainaigawa formation, from thepresent writer's observation, can be subdivided into two parts. The upper partis composed mainly of sandstone andconglomerat.; which yield molluscan
fossils from the middle part. The basal
part has yielded fish remains probablyof freshwater origin. plant fossils andlignite. The lower part of the formation consists of green tuff. green tuff
breccia and andesite lava flo\vs at its
lowermost. No marine fossils have beenfound from this part.

Index 11ap of the Fossil Localities

Collecting Localities andMolluscan Fauna

The specifically determined molluscan
fossils, collected by the writer from 13
fossil localities within the southwesternpart of the Hirosaki Basin, are from theupper part of the Ainaigawa formation.
and these are abridged in Table 1. The
fossil localities, their respective elements.
and detail situations are given in thefollowing list. Table I, and Text-fig. 1,
respectively.
The fossil localities are as follows, andthe numbers before tlwm refer to those

394.

The Miocene 111oliuscan.Fossils from the Area Southwest of Hirosaki
Table 1.

Faunal List of the Ainaigawa Formation
Loc. No.

Species :\"ame-1-~Vumla sp~---- - - - - ~~3

-11 Clz/(u,-'·SOkitalul(YoKO\'.:X:\IA). 1926~(Fig.-27)_ _ ,_ _12 Chlamys nisataie11sis OTUKA, 1943
13 Chlliiii)'S s p.
14 PatinoPecten kimurai (YOKOY.-UI.\), 192fi* *15 Patinopectel/ ilimurai ugowsis 1-IATA 1 and

.j

5 6 7

8

9 10 11 12 13----~~-..*

2 Acila divarirata (H1l'\Ds). 1843 (Fig. 6)
3 Acila sp.
4 Portlandia scapha (YoKOY.-\~1A). 19211 (Fig. 2)
5 Yoldia tokunagai YoKOYA:\1A, l92:i (Fig. 3)
6--G(vcymeris-i)/~i(i/tyei-No~1CRA, 193ii (Fig. 23)
7 Gfycymeris sp. 'A
8 Glycymeris sp. B
9 Limopsis :;p.
10 ,\/ytilus sp.

N1~1YA:\1A,

:3

*
*

*
*
*
*
*

*

.. * *

*

*

* *
*

1934

16-PatinopiCtcu YanzOsakTnu·asalilcllsi~ (No\tl:RA)-.1935 (Fig. 26)
17 Patinopecil'll sp.
18 Venericartlia siogamensis :\ o~n- R ", 1935
19 Venericartlia sp.
* ..
20 Corbicula sp.
--21 Lucinonla andiHneatum (Cc}:,-RAI-,), 1849 (Fig. 10)
22 Pillucina yolw_vamai (OTVKA), 193·1 (Fig. 9)
23 Vasticardium n. sp. (C. arenicola (REEV1-:)-type)
24 Laez,icardium sp. (Fig. 17)
25 Pap_rridea sp. (P. cf. kurodai HATAI andNISIYA~IA, 1953)'i6C!inocartlitmz/ial·usmcaense-NO.\Wl~A. 1935
27 Clinocardium cf. shinjiense (YoKOYA:\IA), 1923
(Fig. 18)
28 Nemocardium samarangae ~L-\I<lYA.\1A. 1932
29 Dosinia ka11eharai YOKOYA:\1A, 1926 (Fig. 1)
30 Dosi11ia ciiilwzenensis XAGAO, 1928 (Fig. 8)
31 Dasinia akaisiww l\o.\lUR~l935- (Fi-g,-7'')~--32 Dosinia odosensis NO\lVI~A. 1935 (Fig. 11)
33 Dosinia lllftantana Noxll:RA, 1935 (Fig. 4)
3,1 Dosinia sp.
35 (..)•di11a jaj>onicu ICniADA. 1952 (Fig. 16)-36Venus cCtoreunla(GOl: LD f1850~---~~37 Venus n. sp. A (Fig. 12)
38 Venus n. sp. B (Fig. 13)
39 J!ercenaria cf. cltitaniana (YOKOYA:\IA), 1926
(Fig. 5)40 Ve11erupis n. sp.
4fclenumtia (?)-s:c-p_--42 Macoma optim (YOKOYA~1A). 1923 (Fig. 21)
43 Macoma spp. (Figs. 20, 22)*
44 Falmlina n. sp. (F cf. nitidu/a (DU:-<KER), 1860)
45 Peronidia cf. protovemtlosa (Nol-.Wf{A). 1935--;r6-ciiTtfilTitslzililii,IJnsls-'roK.oy A~1.C 1923-<Fig. 13)47 Solen cf. gouldi CO:-IRAD. 1867
48 Patzomya _,imatomensis 0Tt..:KA. 1934 (Fig. 15)
49 PmwPe japo11ica (A. ADAl-.15), 184!:1 (Fig. 14)
50 Turrite/la l!:lwlaii :\"Q)I.Il'RA. 1935 (Fig.__2_;;L___ __ 1_ _ _51 Nept111wa cf. lmkusimensis anomalia Nm.tt..:RA and i'
HATAI, 1936
52 Babylonia d. kozaiense l'0\1URA, 1939 (Fig. 24)
53 Balanus sp.

*

*

*

*

*
*

*

*
*

*

*
*

cf

*

*
* *
*
*
* *

..*

*

*

*

*
*

*
*

*

*
*

*

*
*

*
*
*
*
*

*

4

Takehiko lW.4!

in Text-fig. 1.
Loc. 1: Upper stream of the Inakari-zawa,
l'vlatsukitai. Hirosaki City.
Loc. 2: About one kilometer south of Owa.
sawa along the Miyarna.zawa. Hiro·saki City.
Loc. 3: Under the bridge at the south endof lchi-no.watari, llirosaki City.
Loc. 4: About one kilometer northeast of.Mt. Kudoji, Sakamoto. Hirosaki City.
Loc. 5: Upper ::;t ream of the Tochinai-gawa.
Soma-mura. 1\'aka·Tsugaru-gun.
Loc. 6: A right side cliff of the Ainai·gawa.
Sorna-mura, Naka·Tsugaru-gun.
Loc. i: The Funauchi !\line. Soma-mura,Naka. Tsugaru.gun.
Loc. 8 : Left side of the I waki River near
the Kawara-tai. 1\ishi-meya-mura.
Naka- Tsugaru-gun.
Loc. 9: Junction of the lwaki Ri\·er and the0-kawa. Kawara-tai. 2'\ishi-meyarnura. Naka- Tsugaru .gun.
Loc. 10: Middle stream of the Onikawabezawa, Nishi.mcya-mura. l'\aka-Tsugaru-gun.
Loc. 11 : Head of the stream of the Onikawabe-zawa, l'\ishi-meya.mura, 1\akaTsugaru-gun.
Loc. 12: About one kilomtter northWl'st of
the Anmon Fall. 1'\ishi-meya-mura.
:'\aka-Tsugaru-gun.
Loc. 13: Upper stream of the l'\akamuragawa (Nagamatsu-no-sawa). Ajigasawa-machi. :\ishi•Tsugaru-gun.

Hemarks on the Molluscan Fauna
The 13 fossil localities according tolithology can be classified into fourgroups of A. B. C and D. A is characterized with bluish gray, greenish blue
or greenish gray medium grained sandstone with or without pebble-bearing
sandstone in association. Group B includes gray siltstone which becomesbrown on weathered surface. Group C
consists of pebble to granule conglomerate and brown tuffaceous sandstone. andis associated with muddy sandstone.
Group D comprises gray to bluish graycoarse grained sandstone or greenish
gray medium grained sandstone. The
respective fauna of these groups may
be summarized as follows :Group A: Patinopecten kimu;ai, Venericardia
sp., :\'uct1/a sp.. ~llacoma sp., Acila dh•aricala,tlcila sp .. C:linocardium cf. shinjiense, Venus
sp., Cultel/us izumoensis, Solen cf. gouldi,( :tycymeris sp. and Venaupis sp.
Group B: Portlandia scapha, Yoldia tokunagai. Clzlamys sp.. Lucinoma acutilineatum.
Laez•icardium sp .. Clinocardizmz cf. narusawaense,/!osinia kaneharai, D. tugaruana. Venus cf.
loreunw. V. sp .. Jlacoma optit•a. 1\1. sp .. Permzidia cf. Protm·emtlosa, Panope jaPmtica.
Group C: G/ycymeris oinouyei, Jlyti/us sp.,l'atinopecten kimurai. Pat. yamasakii iwasakicnsis. nosinia kanelwmi. lJ. akaisiana. D. Ill·,;aruana. Fobulina sp .. Panomya simotomensis,
Turritella S·hataii. .Veptunea cf. lmkusimaensis.
Group D: Patinopecterr kimurai. JVemocardium samarangae. Pillut'ina yokoyamai. Dosiniasp., Cyc!ina japonica.

From the assemblages tentatively considered as related with bottom sediments
in which the infauna Jived. it seems that
there existed some differences in theirrespective environments. Leaving aside
the swimming forms (Patiopeclt'll. Ozlamys'. and indeterminable specimens whose
individual number is very few ( Veuericardia, Nunt!a, 11frtilus. Fabulina, Venerupis),the following remarks may be presented
with concern to the relationship between
lithofacies and fauna.
The conditions under \Vhich Group A
were buried may ha\'e been a very sh<ll·
low sea not far from the strand line
and of a depth less than about ten met crs. This may be upheld by tll,e pre·sence of burrowing forms as Cultellus
and Solen. two genera which occur insuch regions in the present day seas

394.

The lvfiocene Afol/uscan Fossils from the Area Southwest of Hirosaki

surrounding central to southern Japan.
The presence of either Arila or Clinocardittm does not hinder such a view.
In the case of Group B. the conditions
appear to have been different from thatof Group A. because from the occurrenceof Purtlandia, Yuillilr. Lucinoma. a widely
open embayment or open sea em·ironment is more favorable to their flourishing than a closed bay open to the sea
only with a narrow strait. The abundantoccurrence of specimens of Dosinia andVenus together with the presence ofdeeply burrowing forms as PanojJe andshallowly burrowing ones as Clinocardium.
Dosinia and Peronidia. it is suggested
that the sea may not have been very
deep. although deeper than that of Group
A. Genera as Lucinoma, Dosinia andcockles are usually found on beaches
where rather strong waves prevail andsuch may ha\'e been the em·ironment of
the present group.
(;roup C. \Vhirh is characterized by abundant species of Doshia. burrowing pelecypods as Pmzomya and G~rrymeris together with epifauna gastropods as Turritrl·
Ia and .1\ieptun.ea. evidently points to an
environment locally governed br conditions different from those of Group A
and B. Because of the epifauna. it isconsidered that no strong waves inlluE:nced this area. although it is notnecessary that the region was sheltered.
Group D which has only three specifically determined species at present. isdifficult to judge so far as the environment is conerned. However. from thepresence of C~w:!i11a japonica. a genus
which thrives in brackish to very shallow waters of sheltered areas. it may bethat the present area was also of sucha place.
Definite conclusions regarding the environmental conditions of the four
lithologically distinguished groups are5

withheld until further specimens arecollected and until the details of thegeology are;worked out.
The number of species in common
between each of the 13 fossil localities
is small. but Patinopecten kimurai (YoKOY:\:.tA) occurs most widely.
The few
number of species in common between
each of the fossil localites may be explained by the local differences in eco·logical conditions. difficulty in collecting
well preserved materials. bottom controlof the sediments, situations within thesedimentary basin. and not by the difference in stratigraphic horizon because
detail sections were taken and stratigraphic positions were determined. However. the occurrences of the fauna of each
respective locality are considered important in proving the distribution of theneritic faunae within the said formation.
and in extending the distibution of themwithin the Miocene deposits of Japan.
The molluscan fossils from the Ainaigawa formation. when compared withthose reported from the Tanosawa formation tS. No ... a·t~A. 1935). reveal important characteristics. which indicate thatthe two may be contemporaneous ingeological age. It is thought that thissimilarity between the Ainaiga\va andTanosawa format ions will be further
strengthener} by the discovery of morematerials from the Ainaigawa.
The molluscan fossils which are considered to be characteristic species in
the japanese Neogene and in common
between the Ainaigawa and Tanosmva
formations are:(;~vcymeris
ohzouyei
No~tt·t~A. Patinopel"len kimurai (YoKOYA\tA l. Patinopecten ilwsakiensis ! :\m.tt:RA).
Veuericarrlia sio}.[amensis NoML'RA. Luciuoma acutilinealum (CoNH,·\1>), Dosinia
tugaruana Nn\n: •~A. Dosinia odosensis:\ O\ll'RA.
Dosinia akaisiana :\o:-.tuRA,
Cult ell us izumoensis YOKOY /\ \t". Pan ope

Takehik_o :!W!I!
japonica (A. ADA~ls) and Panomya simo·
tomensis On~KA. Of these mentioned
species. Dosinia /ugaruaua. D. odosensis
and D. akaisiana all of No:~-1uv\ 11935)
may yet prove to be Dosinia chikuzenensis
NAGAo (1928).
However, if such becomesthe case. then the distribution of N A<.Ao's
species will be extended and the geolo·
gical significance strengthened.
That the Ainaigawa formation .is near·ly equivalent to the Tanosawa formation
can readily be recongnized by the mutual
species. and further. it may be addedthat the Shiogama formation in Miyagi
Prefecture. whose fossils were describedand illustrated by S. No:-.Jci~A (19351 isalso approximately correlatable with the
Ainaigawa.
The molluscan species
mutual between the Ainaigawa and Shio·gama formation are : J!cila dil'aricata(Hc•ms), Patinopecten kimurai (YolWY.-\·l.tA). Luciuoma acutilineatum (Co:-.:RAD).
Vonericardia siogamensis No:I.IL'I~A. Cultel·
Ius izumoensis YoKOYAi\IA, PanojJe japonica(A. ADA~Is) and Turritella s·hataii Nol\ll'·
RA. The last mentioned species is considered to be a good guide fossi I in theearly ~·Iiocene deposits of Japan.
Important in determination of the geological age and correlation, and occur·
ring in the Ainaigawa formation are suchspecies as Dosinia (Kaneharaia) kanelwrai
YoKOYA~tA. Cyclina japonica KA~IADA and
Mamma optiva (YoKoYA~IA), all of whichhave rather widespread distribution andrather short geological range.
From the extinct characteristic molluscan fossils occurring from the Ainaigawa formation, it is thought that theformations to be mentioned are its cor·
relatives because of the mutual yield ofidentical species and stratigraphic realtionship of those formations to subjacentand superjacent ones in the areas inwhich they develop. These formations
are: Sugota in Akita Prefecture. Higa-. , ·~·.

shi Innai in Ishikawa Prefecture, Kadonosawa and l\arusawa -in Iwate Prefecture. Moniwa and Shiogama in Miyagi
Prefecture. and the Tanagura formationin Fukushima Prefecture.
The stratigraphical evidences exhibitedby the Ainaigawa formation as the nonmarine sediments of its lower part. theterrestrial conditions of its lower upper
part and marine fossil bearing sedimentsof its upper part, indicate that although
gradual subsidence was taking place,marine waters did not invade the present
area until the middle upper part of theformation. This marine inYasion whichbrought many warm thermal mollusca
to their site of burial is considered toindicate the first stage of marine transgression of the Neogene in Aomori Prefecture. This mentioned phenomenon isa characteristic of the early Miocene ofJapan. and may be recongnized in many
areas outside of the present one.

Heferences
Aomori Prefectural GoYernment (1949), Underground Resources of Aomori Prefec1ure. Part 3. Naka-Tsugaru District. Pub!.
Aomori Pref. Gol'cnt.
K. and NtsiYA~IA, S. (1949). New
Tertiary Mollusca from Japan. four.
l'a/eont .. l'ol. 23. No. I. pp. 87-94, 2 pis.
KA!\1.\DA. Y. (1952), On Some Species ofC.rdiua from japan and Korea. Trans.
/'ror. Palaeont. Soc. japan, N. S., 1'\o. 6.
pp. 161-173. 1 pl.
KAKAYA. T. (1948), The Geology of thesouthern Part of the Ilirosaki Basin.
Report of PEAC (i\IS).
- - (19-19). The Gcologr of the Southern
Part of the Hirosaki Basin. ]our. Geol.
Soc. japan. Vol. 55, Nos. 648-649, pp.
HAT,\1.

181-182.
MASL'DA, K. (1955). Miocene Mollusca fromi\"oto Peninsula, Japan. Part 1. (I), Tra11s.
/'roc. Palneont. Soc. ] apnn. N. S .. No. 20,

394.

The Miocene Molluscan Fossils from the Area Southwest of Hirosaki

pp. 119-127. 1 pl.
- - (1956), Ibid. Part. 1. (II). Ibid. iVo. 21.
p(>. 161-167. 1 pl.
NAGAO. T. (1928). Palaeogene Fossils of theIsland of Kyushu. japan. Part 2. Sci.

Rep .. Tolwku imp. Uuil• .. Ser. 2. Vol. 12.
No. 1. pp. 11-1,10, 17 pis.
Noi'>JllRA. S. (1935). lVliocene iVIollusca fromthe ~isi-Tugaru District. Aomori-1\en.
Kortheast Honsyli. japan. Saito /Jo-011
Kci .\Ius .. Res. Bull., _\To. 5, pp. 19-74. 7
pis.
- - (1935), On Some Tertiary Mollusca from
Northeast Honsyu. japan. Part 1, Fossil
Mollusca from the Vicinity of the Narusawa !lot Spring. Northeastern Part of
the Kurikoma Vokano. h\·ate-ken. ibid.
l\io. 5. pp. 11-100. 2 pis.
- - (1!:135), Miocene Mollusca from Siogama.
Northeast 1-lonsyli, japan. Ibid. No. ti,
pp. 193-234, 2 pis.
- - (1940). 1\lolluscan Fauna of the ~VIoniwa
Shell Beds exposed along the ~atori­
Gawa in the Vicinity of Sendai. !VIiyagi
Prefecture. Sci. Rep., Tolwku Imp. Uni11.,,)'1'1'. 2. ·vol. 21, No. 1, pp. 1-46. 3 pis.
- - and HATAI. K. (1936). Fossils from the
Tanagura Beds in the Vicinity of the7

Town Tanagura, llukusima-Ken N~lrth­east llonsyu. Japan. Saito Ho-110 1\aiMus .. Res. Bull .. l·•·o. 10. pp. 109-155. 5 pis.
--and - - (1936). On Some Fossils from
Akita Prefecture. Northeast Honsyu.
Japan. ] ap. ]our. Geol. Geogr.. ·vol. 16.
Nos. 1-6. pp. 39-64, 5 text-figs.
- - and ZJ:sso, N. (1935). l'v!ollusca from the
Yanagawa Shell-Beds in the Ilukusima
Basin, :\'ortheast Honsyu. japan. Saito
llo-ou 1\ai 1llus.. Res. B.-1!1., cVo. 6. pp.
151-192, 1 pl.
OTUKA, '{. (1934). Tertiary Structure of thel\orthwest End of the 1\.itakami Moun·tainlancl. Iwatc Prefecture, Japan. Bull.
Earllu1. Res. lust .. Vol. 12, Pt. 3. pp. 5666~8. 1 map. 8 pis.
S,\KAJ. l;, (1954). Report oi Investigation ofUnder flow \Vater along the course of
Owasawa l~iver. Pub!. Aomori Gt1t·ern .. pp.
1-103.
- - (1959), "Sunakose-Buraku" (~lem. Lect.
for erection of the l\leya-Dam). Pub/.
llirosaki Jlunicipal OJ]ice.
YoKOYAMA. l\1. (1925). Molluscan Remains
from the Uppermost Part of the .To-Ban
Coal- Field. ]our. Col/. Sci.. Im.b. Unit·.
Tokyo, Vol. 45, Art. 5, pp. 1-34. 6 pis.

Takehiko I w:.4I

8

Explanation of Plate 1(All figures in natural size except Fig. 25)
Fig.
Fig.

L
2.

/Josiuia (1\.auelwraia) kaneharai 'YoKOYAMA. Loc. No.8. External cast of a left valve.
l'orllmulia scapha (YOKOYA~IA). Loc. No. 8. External cast of a left valve.

Fig.

3.

Voldia tofumagai YoKOYAMA.

Fig.

4.

/losiuia tugaruana No:-.IURA. Loc. No. 7. Internal mould of a right valve.
Merrenaria cf. cltilaniana (YoKOY/;:\1.-\). Loc. Ko. 8. External cast of a leit valve.
Arila dimricala (Hir-;os). Loc. 5. External cast of a right valve.
/Josiuia akaisiaua No:\ll'RA. Loc. No. 7. Internal mould of a left valve.
/Josiuia chifwzrmsis NAGAO. Loc. No. 13. External cast of a right valve.
Pilludna yokoyamai (OTt:KA). Loc. No. 10. External cast of a left valve.
Luciuoma actilineatum (Co:-;R.';D). Loc. No. 10. Internal mould of a left valve.
f)osiuia odoseusis No:-.n.:RA. Loc. No. 7. External cast of a left valve.
Vem1s n. sp. A. Loc. No. 8. External cast of a right valve.
Vetms n. sp. B. Loc. ~o. 5. External cast of a left val\·e.
Pauope japouica (A. ADA:\Is). Loc. No. 8. Internal mould of a left valve.
Pauomya simo/omensis 0Tt::KA. Loc. No. 7. Rubber cast of a right valve.
Cyclitut japo11ica KA:\IADA. Loc. Ko. 10. External cast of a right valve.
Lacl'icartlium sp. Loc. ~o- 8. External cast of a left valve.
Clilwwrclium cf. shiujiensis (YOKOYA~IA). Loc. No. 8. External cast of a left valve.
Cultellus izumoensis YoKOYAMA. Loc. No. 5. Internal mould of a left valve.
Macoma sp. Loc. No. 8. Rubber cast of a right valve.
Macoma optit·a (YoKoY,\:\IA). Loc. No. 8. External cast of a right valve.
,)/acoma sp. Loc. No. 8. External cast of a right \'alve.
Gl_n:rmeris oinouyei No:-.ICRA. Loc. No. 7. Internal mould of a left \'alve.
Babylouia cf. kozaiensis xo~IURA. Loc. No. s. External cast. showing front part view.
Turritdla s-lwtaii No~IL'RA. x:l. Loc. No. 7. Rubber cast of spire whorls.
i'alinopedett yamasakii i1casakie11sis (NO:\IUI~A). Loc. l'\o. 7. Rubber cast of a left valve.
Chlamys akilmw (YoKoY,\~IA). Loc. No. 13. Left Yalve of a younger shelL

Fig. 5.
Fig. 6.
Fig. 7.
Fig. 8.
Fig.

9.

Fig. 10.
Fig. 11.

Fig. 12.
Fig. 13.

Fig. 14.
Fig. 15.
Fig. 16.
Fig. 17.
Fig. 18.
Fig. 19.
Fig. 20.

Fig. 21.
Fig. 22.
Fig. 23.

Fig. 24.
Fig. 25.
Fig. 26.
Fig. 27.

Loc. No. 8.

External cast of a left valve.

[ WAJ:

Miocene Mollusca

Plate 1

Trans. Proc. Palaeont. Soc. Japan, N. S., Ko. 41, pp. 9-12, pl. 2. Apr. 10. 19ol

:i95.

A i\EW PALAEOG!:.:NE SPECIES OF THE GENUS
EUCOJ.VJMIA FR0:\1 HOKKAIDO. JAPAi\*KAZUO HUZIOKA

Institute of 11ining Geology, Mining College, Akita University;ftifa.i1i:i¥fi!·ta:::::fCO) Eucommia jTfifi: m*iF..f.Rfi!;J:: I) tJ•:fl;.fdft!(;t;:l*~ Lt-:: Eucomn:ia
Kobayashii HuzloKA (ilil11D ~;rt:~Lt-::. E11commia Jtijl'£ 11iJin:•f1~;:.5}::(Ji-f7.JO)k"(·;{;~n:. rt:ott:lt-'I"-£RO)~'E* .t ':> 3 fil!;;-o Gtt·n -~ M: ~~-lJi',Q)fflf~ll'1.t 'J ::::i'.r~:l::n::IJalll! E.
ulmoides OLIVER 0)~111 ~{'H1lf Lt.:, .:. O)~mHtiA:J.:~-ettii!£-~~~~·O)i!i~::=:*Cfii!"C' ~ 7.> o
0

i1i

IMJ

!J}

deposits of Montana. l\IJKI (1941) reported the occurrence of Eucommia-fruit,
which is indistinguishable from the liv·ing E. ulmoides. from the Lower Pliocene
Pinus trifolia Bed of Central Japan. The
valid fossil species hitherto recorded are
four as mentioned above. and to them E.
Kobayashii HtJzJoKA, sp. nov. has been
added.
The four species of Eucommia and theiroccurrences are enumerated in the table
and in the map (Text-fig. 1).
All fossil species hitherto known to us,
as well as a new Japanese species. were
found only on fruits without leaves andother organs. As far as fruit is con·cernerl. all Neogene species of Europe
and that of Japan are quite similar tothe living E. ulmoides. but the Palaeogene forms including the present new
species are considerablly remote in similarity from the living. It is believed thatthe appearance of the genus Eucommia
may have been at latest in the Early
Palaeogene Tertiary epoch and spreadwidely in the Northern Hemispherethrough the Neogene until it was extinct
from the most part of the area excepting the Yantze Valley region of China
where E. ulmoides is seen existing.

Lately, a new species of the genus
Eucommia Oun:R has been collected by
Mr. ?-.1. KoBAYAY.·\SHI in the Eocene Yu·bari coal-bearing formation at the Shimizusawa coal-mine of Yubari in lshikari
coal-field of Hokkaido.
The monon·pic species of the modern
Eucrmunia. E. ulnwides Or. I VEl~. now grows
only in China its distribution confined toprovinces of Hupeh. Szechwan, Anhwei
and Chekiang t,H. H. Hu and W. H. CHuN,
19:21). This deciduous large tree bears
the novel fruit characteristic to the genus
as shown in Pl. .t3. Fig. 1-b.
The fossil Euwmmia was first reported
by C1.. and E. M. REID (191)7) as an undefinable fruit. which was later named
Eucommia europaea ~l~DI.Eil 11939.1. fromthe Pliocene lignite-bearing deposits ofFrankiurt am i\!ain in Germany. Next.
BERI~Y !1930) described some fruit-specimens of the Wilcox flora in Tennessee
under the names of Simarubites e01:enicus
and Carpo!itlms banisteroidcs. both ofwhich were corrected by BRow:--: (Hl40)
to Eucommia eocellif'a (BERRY). Anotherfossil speci.::s of 0;orth America is E.
montmw BRow:-~ (1940) from the Oligocene* Read June ~0. 1956: received Apr. 30. 1960.
!I

Kazuo HUZIOK.4

10

Species of Eu('(Jmmia and their occurrences
Species

E. ulmoides
OI.IVERI

RecentPliocene

MioceneJapan (Gihul Olig~ne

Szechwan)-------·-- --(Tegelen and I
Brunssum) ,·Hungary(Mizerma);
Germany(Franldurt
a ..\I.)

France(Coiron)u.s.:\.

E. montana------(Grant,l'vlonta na)

Br~owN

E. eocenica(BERHY) BROWL"

Eocenepref.) ; Poland
IChina(1-lupeh.
(Rostov);
Chekiang <tnd Holland

E. l'ltropaea
f\t!;',nt.ER
~~

I

I

- E. Kobayashii
III.:ZIOKA

I

1-----1'~---------~--"-----------

L.S. A.
(Holly Springs,
Tennessee)Japan(Hokkaido)- - --------'--------

Text-fig. 1. Map showing the distribution of Eucomn:ia.
1.
2.
3.

E. eocenica (BERRY) BROW:o\ .. Eocene
E. montana BRow:-: ........ Oligocene
E. Kobayashii HuzroKA ...... Eocene-1.

E. europaea MADI.ER and E. ulmoides

5.
G.

OLIVER ........ Pliocene and i\liocene
E. ulmoides OLIVER ........ Pliocene
E. ulmoides OLJ VEH . . . . . .... Recent

395.

A New Palaeogene Species of tlze Genus Eucommia

l•,'ucommia 1\ohayasllii, sp. nov. is theearliest species of the genus in the Old
W c. rid being the ftrst and only Palaeogene species.
The writer wishes to express his cordial thanks to l'vlr. Masao KoHA L\SHt of
the IIokkaido Tanko-Kisen Co. Ltd. whocollected the specimens and sent themto him for study. and to Prof. S. :\ltKt
of Osaka City Cniversity and !VIr. S. lsHtDA of Kyoto University who kindly gaveassistance to him in the determination of
Eucommia.
Description of Species

Family Eucommiaceae
Genus Fucommia 01.1\'ER
Eucommia Ko/1(/yashii HuztoKA. sp. nov.
1'1. ::!. Figures 5· 10.

Dcscriptiou:- Fruit samaroid, coriaceous
wing surrounding the nutlet. Samara
with rigid peduncle 5-8 mm long, compressed. elliptic to oblong in outline. 2030 mm long and 8-12 mm broad. inequilateral. rounded or obtuse at the apex. narrowly cuneate at the base. entire at themargin. markedly notched at a little below the apex. Nutlet oblong to lanceolate. 10-15 mm long and 3-4 mm wide. 1seeded. suported by a stout rib from theapical notch. very fine fibrous membrane
derived from the nutlet to the marginof samara.
The fruit-specimens are not so rare in
the Yubari coal-bearing formation at theShimizusawa coal-mine. but leaf-specimens of Eucommia have never been obtained in spite of the careful collection
during past ten years. Fruit-specimens
are represented mostly by distinct impressions on dark-coloured shale, butrarely by depressed remains. One of the11remains. a nutlet covered by fibrous
membrane, is shown in Pl. 43, Fig. 10.
Comparison :-Compared to /~'ucommia
ulmoides OLI\'ER (Pl. 43. Fig. l·b). which
is a monotypic species living only in
China. the present fossil fruit is very
similar in the essential characters, butis generally smaller in size and distinctly inequilateral. the apical notch being
situated at the lateral side. The known
fossil species of Eucommia are E. em·euica(BERRY) BRow"' i. Pl. 43. Fig. 2a-b' 1BERRY.
1930. p. 94. Pl. 44. Figs. 15 and 16, as
Simarubites eoceuicus: p. 134. Pl. 33. Figs.
5 and 6. as Carpolithus bauisteroides:
BRow~. 1940, p. 349; from the Wilcox
flora in Tennessee. F.. montana Btww~(Pl. 43. Fig. 3) (BRoil'="· 1940. p. 349. Textfig. 3) from the Oligocene deposits ofMontana, E. europaea i\L\IJt.Eil (Pl. 43.
Figs. -la-b) (l\hDLER, Ul39. p. 10a. Pl. 8.
Figs. 29-31 : Pl. 9. Figs. 9-JJ, from the
Pliocene lignite-bearing format ion at
Frankfurt am Main in Germany. and E.
ulmoides OuYEt~ (Mtl\t, 1~41, p. 277. Pl.
6E. Fig. 14F) of the Pliocene Pinus trifo!ia Bed at Ichinohara in (;ihu prefecture of Central japan. E 1\obayasliii iseasily distinguishable from all of themby the situation of the apical notch. In
the new species. fruit is notched at theapical part of the lateral side as seen in
the fruit of Euptelea polyandra StEB. etZL'CC. (Pl. 43. Fig. lll.
Remarks:-The associated genera with
Eucommia Kobayashii in the A plantzonule of the type locality are as follows:
Ginkgo, Jletasequoia. G/yptostrobus. Jugfans, Alnus. Betula. Cmpinus. Castanea,
Quercus. D(mplzyllum. Ulmus. Zelkmm.
Planera. Cercidiplzyllum. 1\-Iagnolia. llamame/is, Fotlzergi/la, Liquidambar. P/almm~.
Crataegus. Jlfallotus. Rims. Celastrus ..-leer.
Paliums, Afar/ea. Fra.xinus. and etc. Judging from these generic association. theforest in which F. l\.obayasl1ii existed is12

Kazuo HUZJOKII

suposed to be the deciduous temperate
forest which is closlely related to themodern forest in Central China where
E. u/moides now grows.
Occurrellce :-The type locality: Shimizusawa coal-mine, Yubari city, lshikari
coal-field, llokkaido: the A fossil-plants
zonule of the Yubari coal-bearing formation : Eocene.
Coli. l\lasao Kos.>. Y ASH 1.
Holotype : Fruit-specimen figured in
Pl. -13. Fig. 5.
Depository: Institute of ~lining Geo·logy. Mining College. Akita University.
References
BERRY. E. W. (1930). Revision of the Lower
Eocene Wilcox Flora of the SouthwesternStates: with Description of ~ew Species.
chiefly from Tennessee and Kentucky. U.

S. G. S. Prof. Paper 15().
BRowN, R. \V. (1940).
New Species andChanges of Name in Some American Fossil Floras. TFashinglfm Acad. Sci., ]ounz.,
Vol. 30, No. 8.
GRA:-;GE0:-1, P. (1958). Contribution a !'etudede Ia paleontologic vegetale du Massif
du Coiron (Ardeche). Mhn. Soc. d'Jlist.
Nalur. d'Aut•ergne, No. fl.
Hu H. I I. and W. Y. CHu:-; (1927). leones Plantarum Sinicarum. Fasc. I.
~liKI. S. (1941). On the Change of Flora in
Eastern Asia since Tertiary Period. (I)The Clay or Lignite Beds Flora in Japan
with Special Reference to the Pinus trifolia Beds in Central Hondo. ]ap. ]ounz.
Botany, Vol. 11.
:\1:\m.EJ{, K. (1939). Die plit>z:inc Flora vonFrankfurt am Main.
Abh. Senckenb.
Naturf. Ges .. ;1M. 446.
SZAFI·:R, W. (1951). The Family Eucommiaceae in the Tertiary of Europe. Odbitka
1\osmosu, Ser. A. Vol. 66.

---------------

Explanation of Plate 2
Figs. la-b.

Eucommia ulmoides 0J.IVER (living leaf and fruits for comparison)la Leaf x1/2. lb fruits.
Figs. 2a-b. Eucommia eoce11ica (BERRY) BROW:-> (for comparison)
Type specimen. copy from BERRY (1930).
Fig. 3.
Eucommia montana BIWWi'> (for comparison)
Type specimen. copy (rum BROWN (1940).
Figs. ·la-b. Eucommia europaea M;\DI.ER (for comparison)
Type specimens (Fig. -Ia is the holot~·pc). copy from l\L\DLER (1939).
Figs. 5-10. T:uwmmia Kobayashii lhJZIOKA, sp. nov.
Fig. 5 I-Iolotype: Fig. 10 a nutlet remain covered by fibrous membrane. enlarged.
(Loc. Shimizusawa coal-mine, Yubari. Ishikari coal-field. Hokkaido; Yubari coal·bearing formation: Eocene).
Fig. 11.
1-.'uptelea po/ywulra SII·:n. ct Zucc. (living fruits for comparison) X 2.

A Palaeogene Species oj Eucommia!iUZ/0!\A :Plate 2

lb

x3

2b

\
2a

11

10

x2

x2

9

5

K. SHIBATA photo.

67

Trans. Proc. P a laeont. Soc. Japan, N. S., No. 41, pp. 13-14, Apr. 10, 1961

A SUBFOSSIL CUON FROM ANG TH0 2\G. THAlLA D*

306.

FUYUJI TAKAI
Geologica l Instit ute, University of T okyo

YMJ;~

Ang Thong 0¥. Cuon c7)=Ffl:: b :... ~ /:.1 :; !/c7),lbfJ 100 km Ang T hong M.ifi:

7J -<:0 J.1ffi IJ /11 ;>:;: ;f t/:: Cuon c7) ,/'{1::.1:1~:;l: ·r 7- -t: l) r; ; . Jfu:lj f:: JJL1=. L-:- v<:;, Cuon a/pinus infusms
P ococK f:: J'i.iJ7E 6t1.;,;, ,it'i oif .'f,=:

alluv ia l deposits underlying the g reatcentral valley of the Chao Phraya.
The specimen is a left lo\\'er ja\Y ofyoung individua l and bears only two
premolars (P3 and P4). from the alveolar aspect Ml belongs to the carnasial
tooth, but M2 never appear. judging
from the number of molar and t he reduction in size of l\II:Z, the specimen
clearly belongs to the wild dog, Cuou

A subfossil of W ild dog was offeredfrom Mr. Saman B ~; Ru\·A s, Chief, Geolog ical Survey Di vision, Royal Bureau of
M ines, Bangkok, T hai land, to me byfavor of Prof. T. K osAYAS I I I in order to
ident ify it. T he specimen has beenunearthed f rom a river sand near Ang
Thong situating about 100 km northfrom Bang kok. J udging from its presen ·at ion, it mu st be obtained from an

Text- fig ures l a, b, c . Left ramus of Curm a/jJi111ts i 1(,:usnts P o c n c 1'
a, upp!.! r \·iew; b, outer ,·icw; c, inner ,·iew.

* Read at th <:> an ual m e ::!ti Pg of th:'! Soc i ~ty a t T okyo, De c. 6, 19fif:; r e ce inO> d M J ~- ll. 1%0.
1•_)
•)

1-l

Fuyuji TAKA!

a/pinus PALLAs which lives in the continental Asia southward of north latitude
50' and eastward of east longitude /o'·and the adjacent islands except Ceylon,
Borneo, Philippines. Formosa and theJapanese islands. The following elevengeographical races are now discrimatedby several anthors, namely Curm a!pinus
a!pinus PALLAs (Amurland. Manchuria.
and Sakhalin), javanicus DEsl.JAREST(Java). sumatrensis lL'\RDWJCKI~ (Sumatra
and ~Ialay Peninsula). duklumeusis SYKEs(Peninsular India). primaezms Hoocso:--~(Nepal and Bhutan). !eptums HEL'DE(Kiangsi Province. China). hesperius
AFA:-::\SII"~ and ZoLoTARE\' (Eastern Rus·sia), infusws PococK (North Tenasserium
and ? Annam), fumosus PococK (West
Szechuan, China\ /aniger PococK lKash·mir and South Tibet). and adistus PococK(Cpper Rurma and Indochina).
According to the report from theSiamese l\linistry of Commerce and Com·
munications... Cuon mtilans (S. tVll'LLER) ··which is a synonym of Cuon a/pinus
jamnicus. is distributing widely in thewhole territory of Thailand. On theother side C. B. KLoss has reported two
occurrences of .. Cyon javanicus" from
Bang Nara near Patani at PeninsularThailand (Siam) and Lat Bua Kao at
Korat Plateau.. As the locality of typesof jamniws ·and rutilans is java, theseoccurrences of Javan race in Thailand

is very questionable. I suppose that the
Siamese specimens including this subfossil. must be rather assigned to infuscus
from Moulmein in north Tenasserium
and Kontaum in Annam than to javam·cus from Java in view of their geographical distribution.
Cumz a/pinus infuscus PococK

Locality of type:- Moulmein
Distribution :-Some glass land along
the edge of jungle at North Tenasserium
and possibly Annam.
Measurements of this subfossil (mm.) :Length of the ramus measured from thesymphysis to the condyle
127
Length of the ramus measured from thesymphysis to the angular process 124
Height of the ascending ramus-l7
Depth of the ramus at Pi
17
Thickness of the ramus at Pl
9
Depth of the ramus at Ml
21.5
Thichness of the ramus at l'vll
9.5
Transverse length of the condyle
20.5
Length of P3
9.7
Width of P4
4.5
Length of P4
10.7\Vidth of P4;-,.oFinally I express my cordial thanks
to l'vlr. S. BvRUVAS and prof. T. KoBAYASHI.

Trans. Proc. Palaeont. S:lc. Japan, N. S .. No. -H. pp. 15-20. pl. 3. Apr. 10. 1961

:i!ll.

FORA:\UNIFERA FROM THE E\WZA WAGA W A FORMATIO)lIN SENDAI. JAPAN*
NAOAKI AOKI
lnstitut"' of Geology and Ylineralogy. Tokyo University of Educ::-tion'g".JJkViHtli-ami!!!ffil;=.ft;fJl-t -9 ·· ~¥fr~ .. O)'fi~JJI/WI.I· G.
z- c .:, t.c; fi··ll!P.. (t:fillf.~.:Z..~ ·fC. L. t.:, :lf!
1Jfl!t1JJJ!Il0) Anstelian 7 ;~·- :r!=.MW. L. -c. .:.(1)i1J!mxO).I!'Hii'-!l.!:~1!tO)l.Jt9f~ ~ &?.9J::l.:.OO
ll:f:n:0.9,
rr rnt ag
ftl!f?O)'¥/f\111.~1.1· ;.)O)tr:iL£a:

E!phididla lwmwi C: f.1phidium d. oregrmense*

tuffaceous mudstone. fine to medium
grained sandstone and small-pebble
conglomerate. intercalating a few lignite
seams. In the lower half horizon, two or
three layers (0.5-l meter thick) in which
crowded the molluscan fossils. chiefly
composed of the large shells of Ostrea
gigas THUI\IBERG, are well deYeloped andcontinued laterally with some extents atthe north of Ayashi. west of SendaiCity.
All fossil assemblages and species offoraminifera recorded here are derived
from these oyster-bed of the Imozawagawa. The seven samples had beencollected from the following localities in
Miyagi-mura. west of Sendai City, Miyagi
Prefecture.

Introduction
In the course of the foraminiferal
study of the so-called "Pliocene" formations distributed in Sendai District.
Pacific side of Northeast Japan, such
peculiar fauna as has not· been previously
unknown in Japan. was found and is.briefly reported in this paper. for it isof great interest to age-determination.
Before going. the writer wishes to express his thanks to all Professors of theInstitute of Geology and Mineralogy.
Tokyo University of Education for theirkind guidances and encouragements. andto Dr. Hiroshi UJrrrt for many valuablesuggestio:1s en the classification of foraminifera.

Three adjacent localities at Hongo,
Ayashi. (Sample 1-3)
Hill-side cliff at Ogasso, Ayashi. (Sample

Localities

4)
River-side cliff of the Hirose River at
Otake. Ochiai. (Sample 5)
Two adjacent outcrops in a stream at
Yokote. Imozawa; about 4 km. northof Ayashi. (Sample 6. 7)

The lmozawagawa formation (about 40
meters in thickness) is characterized bv
bluish-gray mudstone and also compris~s* Read and received May 21. 1960.
15

Hi

Naoaki AOKI
Table 1.

--

Foraminifera from the lmozawagawa formation in s~ndai(Number of specimens m 10 grs. of sediment)-

SampleSpecies

Buccella frigida (CUSI!~1AI'\)
F./phidium dtwatum CUSHI\1:\:"
F.!phidiella hannai CL'SHMA:X and GRAl'\T
E/phidium subgramtlosum As A :xol\'onioll pcmcilocu/um CusH~tA:"
Buliminel/a e/eganlissima (L>'ORBIG:XY)
Elphidium subgramtlosmn aureum AoKI
Elphidium cf. oregmumse Cusm.tA:" and GR.'\:-;TKonion 2 spp.
Pseudononion cf. /redecum ASA:'\0
Discorbis globularis (o'OR B IG:XY)
Discorbis cf. p/aua HERO:X·ALLE:X and EARi.A:"\0Fall'lllineria aff. polila PARR
Globigeriuoitlcs I riloba immatura LERoY
Globigerinu pachytlerma (EllRENBERG)

1

2

3

4

5

6

7

175
103

171
31
1<1
1

161
538
107
25

220
119
15

2

3

8

1

5

128
13
12
2
2

25
13
17
3
3

1

4

6

4

11

Fauna

Total number of foraminiferal tests
taken out of 10 grammes of each rocksample, varies from thirty to more thanseveral hundreds. In spite of the abundance of specimens, the examined species
are only 16 in total number including a
few indeterminate:d forms and the twoof the planktonic.
The present fossil assemblage is very
simple and made of primarily the three
species: that is. Buccella frigida tCcsH:vtA:-;). f:!phidium c/cwatum Cusu:-.JA:-; andElphidiella hamzai (CusHMA:-1 and GRAKT)
are extr~mely rich and take more than
90% of the population. In addition tothem. EIP!lidium subKranulosum AsA:"O.
Nmtiou paucilocu!ttm Cvsll~l."' ~. Bulimiuella ele!!altlissima (o'ORBIGNY) and Elphiclium d. oregonense CusHMA!\ and GRA:"\T
are rarely occurred in several samples
as accessory elements. The other re-

1<1
3

?2

?2

3

6

4

I

7

31

75

3

2

1

297

Total

:1

5

23-!

841

310

161

mainders are very restricted in occurrence and are represented by a few
specimens at hand. In general, thespecies belonging to both I he families
Nonioniidae and Elphidiidae are prominent in the present fauna.
Ecology

The aboYe-cited species are seemed to
be all important as the characteristics
for analysis of the ecological conditions.
Nearly all of the foraminiferal constituents haYe been mostly recorded in
the Pliocene to Recent sediments of cold
water in the temperate and the higher
latitude regions. 1\foreover. some are
typically living in the arctic region.
An arctic species. E!phidium d. oregonense Ct·sii\1:\N and G,~,,;-.;,., lhou~h a few
and small-sized specimens obtained. areassociated with a rather high percentage('1-20+%) of Dj>lticlicl!a lunmai (CPsu:-.tA:"i397.

Foraminifera from the Imozawagau.1a Formation in Sendai

and GRANT) which closely related to
Elplzidiella arrtiw (PARKER and Jo=--Es).
So. the present fauna can be stated toshow distinctly a cold water one and it
suggests that the temperature of seawater of the Imoza\vagawa would belower than that of the Tatsunokuchi
which has also been thought to beaffected by a cold current (NoMURA,
1938 and T.u;AYA:-iAGI, 1950).
Considering from the lithology. paleogeography and the molluscan fossil faunaof the formation. the depositional site ofthis oyster-bed is inferred to be a central
or an interior part of an inlet : in otherwords, a .. littoral-brackish environment.''
The high frequency of Buccella frigida(CcsHMA:-i) and the existence of shallow
water elements would support this interpretation. It is notable, however. Rotalia
beccarii (LI:-.:-;~) and its varieties that
occur everywhere and commonly fromthe littoral environment are entirely
absent in the fauna.
Age
There is no other example in Japanthat the foraminiferal composition similar to the lmozawagawa fauna is foundeither in fossil or living. But, it calls
our attention that the Amstelian faunain Netherlands reported by vA:-. VooRTHt:YsE:-; (l!:l58) has a several points ofresemblance to the present one. The
composition of that lower Pleistocene
fauna is such as follows :E!p/;idium oregonense Ct:siHIA:>:and GilA:-;T
E!p!Jidie!/a sp. cf. E. arctica(PARI'I"l and }ONES)(Juinquelocu!i Ita semi nulum(LINN!~)

Alphidium spp.
Streblus beccarii (LI:-;:-;£:)
Eponides frigidus (CCSH:O.JA:-;)others

57
27

5
3
1
4

17

Besides, the similar assemblages are
known from the Pleistocene of Cape
Blanco in Oregon (8.-\:"DY. 1950l. and of
a submarine beach in Alaska ICcsH:-.IA:>;,
1941).

On the Netherland fauna. ,.,,:-.; YooR(1953) remarked its prominent
character of the abundant occurrence ofHlphidiella cf. antica (PARKER and jo--.;Es)
together with E!phidium oregonense
CL:sH:-.tAN and GRA:>;T and he thoughtthat the co-existence of the two conspicuous forms of arctic is to be an .. important climatic indicator" and a good.. guidemarker" to show the first refrigeration of sea-water in the Glacialage. Therefore. the oyster-bed yielding
these two indices may be considered aswell to represent the basal part of the
Pleistocene period.
The Imozawaga\va formaticn had beenthought to be equivalent to the so-called"Lower Pliocene" Tatsunokuchi formation in Sendai, and is mistaken for apart of the latter. Recently, the writer(1960) could make clear that the former
is lied on the latter interposed by thenon-marine deposit of the Shimoda Tuff
between the two. Thus, it comes to beof middle or upper Pliocene age. according to usage. which does not coincidewith the above supposition on age fromthe view-point of the foraminiferal
studies.
·
Tllt:YsE:-;

Remarks on Species
All specimens of the species listed
below are deposited in the collections of
the Institute of Geology and 1\lineralogy,
Tokyo University of Education.

Nonio11 pauciloculum CusH:-.IA:-;. 1944
Pl. 3, figs. 2a, bThis is found rarely from the oyster-Naoaki AoJa

18

bed of the Imozawagawa formation andcharacterized by the seven chambers in
the last whorl and the strongly curved
sutures on which whitish and opaquefine pustules are deposited. l'Vlost of our
specimens are somewhat different fromthe type-figure by its Jess circular out·line.

-17) from the Pliocene Wakimoto formation in the Oga Peninsula. Akita Prefecture. may be clcsely related to thepresent species.
Elp/1idium subgramtlosum aureum
Aoro, 1960
Pl. 3. figs. 7a. b

Vonion 2 species indet.
The two small-sized forms belonging
to the genus Nonio11 could not be determined their specific names. Both havethe umbilicus and sutures covered withsecondary shell materials and slighly
resemble to 1Vonirm pauciloculum CusH·~~,,,,N in general appearances.
Pseudononio11 cf. tredecum AsA:-.:o. 1936
Pl. 3, figs. lla. b

Two specimens at hand have more
limbated sutures compared \Vilh thetype form recorded from the PI iocene
Setana formation in Hokkaido, andresembles to Nonio11ella umekae HL·sEzr:o.rA and l\hRuH,\Sr from the Pliocene
Haizume formation and also to Pseudommimzjaponiwm A sA:-.:o from the Pleistocene Naganuma formation. This form
seems to be rather widely distributed, atleast, over the northern half of japan.

This subspecies was originally described from the recent sediments inTokyo Bay., , li is characterized by afewer number or chambers and theinflated last-formed chamber.
Hlp/iidium cf. oregonense Cusrr11.1A:-.:and GRANT, 1927
Pl. 3. figs. 4, Ga. b

Twelve specimens came from the
Imozawagawa formation had been examin<!d. They have all small-sized tests
and 1-1-15 chambers in the last whorl
and may be of juvenile stage of thisspecies. In japan. the present species
characteristic in cold water had beensometimes reported under the different
name, "Elp/iidium (or Cribroelp/iidium)
ezoense AsANo ··. Diameter of the largest
one is 0.37 mm.
Elpliidium subgramtlosum AsA:-.:o, 1938
Pl. 3. figs. 9a, b

Dphidium clavalmn Cr.;smJAN, 1930
Pl. 3. ligs. Sa, b

It shows a considerable variation in theumbilical and sutural characters. It isabundant in some localities, particularlyrich in the sample from Ogasso, and has
been recorded especially from the Pleistocene to Recent sediments of the cold
water regions. .. Elphidium lzughesi foraminosum" of AsA:-<o (1950, p. 8, figs. 46,

This coarsely perforated form, has
somewhat inflated chambers and a depressed umbilical region without a boss.
Elp/iidiel!a liamzai CusmrANand GRANT, 1927
Pl. 3. figs. lOa. b

The large and rather well preserved
specimens of this species are commonly397.

Foraminifera from the bmzawagawa Formation in Sendai

found in the lmozawagawa formationand is also rarely in the Tatsunokuchi.
Our specimens are all characterited by
the ''narrow sutures", the smooth andhighly polished surface. and the finegranules on a few early chambers of
the last whorl and the apertural face. It
is identical to E!pliicliella nitida Cus11~1AN.
which has been placed in synonym with
E!pllidiella lunmai by BAxov {1950) andit may be also synonymous with .. F!PIIidi'!lla cf. arclica .. of \'AX VooRTIHiYSEN
from the ~orthwestern Europe. In ourmaterials. there is a variety which has
a round test in equatorial outline andcorresponds to .. B-type" of "E. cf.
ardica" of \'A=' Voownn;YsEN (1953),
who well summarized the distribution of
the species in Recent and Pleistocene
age. This is a first record in Japan asfossil Elp!tidie/fa lumnai Cl:smi.•\N and
GRA:-·n, but the another form allied to E.
arctica has been recorded as E!phidie//anagaoi As.\No from the Pliocene depositsof Japan.

19

p/awL in general shape. but. owing to

the lack of the mature specimen. thecomplete determination could not bedone at present. The type was fromthe East Falkland Islands.
Bucce//a frigida ICL·smTAx), 192~
Pl. 3. figs. Sa-c

This familiar species had been recently
re-defined and fully described by ANDERSEN tl952l in the monographic work of
the genus Bucce!la. It is most predominant in our materials and occurred frequently from the Tatsunokuchi.
This
species inhabits in shal!O\v and brackish
waters of cold.
Vah•ulint:ria aff. po!ita PATH{. 19;i0

Only one specimen is found from
Hongo. It is somewhat resembled to
Valvu!ineria polita PARR described fromoff Tasmania, hut differs from the latterin having more inflated and Jess chambers.

Buliminella eleganfissima(o'0RBIGYf},

1839

Pl. 3, fig. 3This widely distributed species were
occurred in the lmozawagawa samples
collected from the environs of Ayashi
and also in the Tatsunokuchi.
Discorbis itlobu/aris (o.ORBIG:"'Y), 1826

Three specimens are all of small size.
Discorbis cf. plana HEiwx-Au.1·:r-:and EARLAND. 1932
Pl. 3, figs. la-cA single and juvenile specimen from
Yokote is somewhat similar to Discorbis

Globigerina pachyderma(EHR E;>;DF.I~C), 1861

A single specimen.

Globi:.;erinoides triloba immatura
L~-:Rov, 1939

The larger one is 0.28 mm
largest diameter.

in theneferencesAol\1, X. (1960), On the stratigraphy of the

Tertiary System in Sendai, :!\ortheast
Japan. (in Japanese with English ab·stract). jour. Geol. Soc. Japan,. rot. 66.
110. 77 2, pp. 17-26. 1 fig.
AsA:-<0. K. (1950). Illustrated catalogue ofJapanese Tertiary smaller Foraminifera.

Naoaki Aol\1

20

Part 1 : Nonionidae. Tokyo. Japan. 12 pp.,
73 text-figs.
BA:SDY, 0. L. (1950). Some later Cenozoic
Foraminifera from Cape Blanco. Oregon.
four. Paleont.. vol. 2·1. uo. 3. pp. 2ti9-2Sl.
pis. '11-42.
CusH~tA!':. ]. A. (1941). Some fossil
Foraminifera from Alaska. Cantr. Cushman
Lab. Foram. Res .. 1'0/. II, pt. 2. pp. 33-38.
pl. 9.
- - (1948}. Arctic Foraminifera. Cushman
Lab. Foram. Res.. SjJec. Pub .. No. 23. 79
pp .. 8 pis.
DA~I. A. TE:-; and TH. REI>:IIOLD (19-11). Die
stratigraphische Gliederung des ~ieder­
Hindischen Plio-Pleistozans nach Foraminifercn. !lied. Geol. Stichting. Serie CV-No. I. 66 pp .. 6 pis .. 5 charts.
HA>:ZA\\'A, s .. K. HAT.\1, J. l\\'AJ. ~- KIT:\·:.HJRA and T. SHJl!ATA (1953). The geology of Sendai and its enYirons. Sri. Rep.
,Tolwlm Uui1• .. 2nd ser .. l'ol. 25. 50 pp .. 4
charts, 2 maps.
Lm:sucH. A. R. and II. TAPPA:--> (1953}.
Studies of Arctic Foraminifera. Smithsonian Misc. Colt .. rot. 121. no. 7, 150 pp ..
24 pis.
No~ll.iRA. S. (1938), Ivlolluscan fossils fromthe Tatunokuli shell bed exposed at
Goroku cliff in the western border of
Sendai. Sci. Rep. Toltokt1 Unir., 2nd ser.,r·ol. 18, 110. 2, pp. 238-275. pis. 33-36.
TAJ,AYANAGI. Y. (1950). Pliocene smaller
Foraminifera from western Sendai. ShortP.zpers I. G. P. S .. No. 2. pp. 23-28:
VooRTHL:YsE:--:. J. H. \'A:S (1949). Elphidiellaarctica as a climatic indic:ttor· for the
Quaternary. Geol. en Mijinb .. lie faarg.,
no. 3. pp. 127-130. (cited in VA:>: VooHTHUYsE:s. 1953)- - (1950). The quantitative distribution of
the Plio-Pleistocene Foraminifera of aboring at the Hague (Netherlands). Med.
Geol. Sticllting. N. S .. 110 ..J, pp. 31-49. pis.
1-4. 1 chart.
- - (195~). Elphidium oregonense CvsH:.IA~and GRA:-->T. a possible marker for Amstclian (Lower Pleistocene) in North America and northwestern Europe. Contr.

Cushman Found. Foram. Res., 1'0/. 3, pt. I,
pp. 22-23.
- - (1953). Some remarks about the PlioPleistocene microbiostratigraphy in north·western Europe and in North America.
Jour. Paleonl .. rot. 2(. no. -1. pp. 601-604.

-----------

Explanation of !'late 3(All figures are x 100 unless otherwise marked. a. side or
dorsal view: b. edge view: c. ventral view.)
Fig. 1. Discorbis cf. ptana (D'ORBJG!'\Y). Reg. no. 51208. Lac. Yokote. Imozawa (Sample 7).
Fig. 2. Nonion pauciloculmn Cusw.JA!':, Reg. no. 5119~1. Loc. Hongo, Ayashi (Sample 3).
Fig. 3. Buliminella elegantissima (o'OtwtG>:Y), Reg. no. 51207. Lac. Otake. Ochiai (Sample 5).
Fig. 4. Etphidium cf. oregonense CusH~IA:S and GRA:-IT. Reg. no. 51203, Lac. Yokote, Imozawa(Sample 6).
Fig. 5. Bucce/la frigida (Cusl!MA:>:). Reg. no. 51209, Loc. 1-longo. Ayashi (Sample 2).
Fig. 6. E!Phidium cf. oregrmeuse CUSII~JA!' and GRANT, Reg. no. 51204. Loc. llongo, Ayashi(Sample 3).
Fig. 7. Elphidil4m subgramtlosum aureum AOKJ. Reg. no. 51202. Lac. Yokote, Imozawa (Sample 7).
Fig. 8. Elphidium clal'tltum Cusu~tA:-->, x70, Reg. no. 51201. Lac. Ogasso. Ayashi (Sample ·l).
Fig. 9. Hlpllidium subgrmmlosum AsANo. Reg. no. 51205, Loc. Bongo, Ayashi (Sample 3).
Fig. 10. E/jJhidiella hamwi (CcsH:>.IA!'\ and GJU:"T). x7o. Reg. no. 51206. Loc. llongo, Ayashi(Sample 3).
Fig. 11. Pseudononio11 cf. tredecum AsA!':O, Reg. no. 51200. Loc. Hongo. Ayashi (Sample 2).

Aora: Foraminifera from Sendailblc

Plate 3

2b

3

5b

6b

8a

5c

Trans. l'roc. Palaeont. Soc. Japan, N. S .. No. 41. pp. 21-32, pis. 4-6, Apr. 10. 1961

2!18.

l\IESOZOIC PLA::--rTS FRO:\I THE rrr:osHIRO SUB-GHOUP.

THE TETORI GROUP. CENTRAL HONSHU ..JAPAN.

PART 2*TATSUAKI KHvlCRA\Lguro Gakuen=fl[:{ll'flff.£i!IRI':llllif\"Jflfc1)'fl'+.ftilllfu: ,H<:ill ~~f!tl::t. r>!l.P~~U::<ft.c.-~ L -c:riJli~M~ 11ft~~. II Hi"tJ\tl:m:.:.f'Stll-t 7.> Eff!is:m;~fifil• btti a? L ~'ll\"t.cllli~ft:fi~ft-~ Lt.:. :. .::.:.;\C~-t' Q0:>!1-i-c1)-$"L'. --.:. 7-7 .A~;:.lll.*·"!:":M .f::., ::::
Dictyozamites ::t.l"l·:irM~
g;.:.t"'1ii.c:I!!S"'C'S.Q.
;t N ~ 19]

<

*

<::.

The present writer wishes. furthermore. to express his thanks to themembers of the Conducting Commissionof the 1\omatsu City Museum~ 1\Iessrs.
A. l-I.-\:\1.-\SAJC\. M. SEKIDO, I. Kn~AKA. S.
:1\,,K.-\YA~'"· Y. SHIROSAJ(I and Y. Y.-\:\JAzAKt for their great help in collecting
the fossil plants described here and else'vhere. and Mr. I. KH>oci'CHI for hiskind help in furnishing lodgings. Finally,
particular thanks are due to the Komatsu
Board of Education and the Komatsu
City l\h1seum !president: Mr. 0!. (kAwA)
for the grant in aiel which has rendered
possible the collection of the Mesozoic
plants described here and later on.

PrefaceThis study deals with some l\Iesozoic
plants collected by l'vlr. Shinji SEKIDo
and the members of the Komatsu City
~vluseum. lshika\\'a Prefecture in 19i1G.
1957 and 19:i!l, from the alterne>.tion ofsandstone and black shale members at
the most upper course of the Mekkcdani.
Ozo. Oguchi-mura. Ishikawa prefecture.
which are presumed to he an equivalent
in geological age to the Kuwashima formation. the ltoshiro sub-group (1\cchian ~the lowest of the japanese triple Lo\ver
Cretaceous).
It is a pleasure to record here writer's
debt of gratitude to the following persons
and organs for their help and support
extended to him in conducting the present
work. The present writer first expresses
his sincere gratitude for the help of Dr.
Seido E:-;oo who kindly guided him in
the prosecution of this study. The chiefcollector Shinji SEKIDO owed much toProf. Z. TsliK:\:-.:o, Fukui Cniversity and:\Ir. II. i\L,Tseo, Kanazawa University.
both of whom kindly guided him in the
study of fossil plants.

Description of Species
BE:\~ETTITALESGenus Dictyozamiles Ot.DliA~I
This genus is one of the most importantand characteristic genera in consideringthe ~Iesozoic floral provinces of the japanese Islands (proposed by T. K1~H%\, 1959
l\1S) which are believed to have existed
in ge.ological age from the Domerian toat least the Albian~ especially in consider.i}lg the Inner side (the llida) Floral
Province.

* Head ~ovembcr 21. 1959 and January 17,
1960; received June 7. 1960.
21

Tatsuaki KIMUR.4
be2n described or recorded. Their vertical as well as horizontal distribution isshown in Table L.

In the japanese Islands and South
Korea. five distinct species and three
undetermined specimens have already

T::.ble 1. Th? Occurrcnc: of Diclyozamiles-------------Eiornl Pro\'inccs I

Outer side--

Province

Geological Ag~

ToyoraFloralProvince

florC~!

Naktong
FloralProvince

a a

ci.
Ul

Q

.::::

0

-.

0

0

0

0

0

0

0

Inner side
(Hida) Flcral
Provincec::i0(I\ibu f.)(Kiycsuc f.):'o.Ialm (Upp:!r Jurassic)

Dogger (Middle Jurassic)(Utano f.)I-- - - - - - - - --~----------(* s~ems to b;; p~culi~r l~ each Flor::~lThis genus has so far bten considered
to be an impt;lrtant ~lement of the ~1iddle
jurassic floras as sta'i:'ed by SEwArw (l!H7J
and also by AR:-.:ow (19-171. But in thejapanese Islands and Korea Peninsula.
most of all the species belonging to thisgenus occur from th~ Upper Jurassic
and the Lower Cretaceous strata. especially from the latter.
In 1936. OrsHt stated ··The geologicalage of both the Tetor~ and the Naktong
Series corresponds approximately to the
Lpper jurassic in the European standard.
Therefore. all of 1he japan:'-se species ofthis genus are the youngest record!-\ inthe world". But the Koch:an (the lowest of the japanese triple Lower Creta·
ceous) age in which the main part ofProvir;cc)

both series (or groups) was formed is
now accepted by most stratigraphers andalso palaecntologists.
It is a striking fact that no species
belonging to this genus has yet beenrecorded from the Outer side Floral Province. This fact shows the differenceof climate between the Outer and theInner side Floral Provinces bounded by
the median ridge. The Toyora and the
Naktong Floral Provinces were the intermingled area of both florai elements. andthe mingling be might explained by thepalaeogeographical circumstances in bothareas in those days.
It is also noteworthy that .. the !Jictyozami!es does not seem lo have beenyielded from the continental Mesozoic

398.

Mesozoic Plants from the Itoshiro Sub-group. the Tetori Groupstrata of Asia. i.e .. Siberia. l\Ianchuria,
China and so on.
The imperfect specimen formerly described by the writer (1958) from thedark greenish shale exposed on themountainsides (Kuzuryu sub-group?)
near Lake Kizaki. Nagano prefecture.
under the name of Dictyozamites sp., now
can be identified as a well-known species
called Dictyozmnites falcatus (:.VI oRr! 1s)
0LDHA~t by its o:.~tward appearance.
ARr-:ow (19471 stated that this genus
probably belongs to the Wiliiamsoniaceae.
hut no evidence for testifying the above
statement has yet been fou11d in spite of23

the writer's strenuous efforts.
In describing the Japanese and Korea
species belonging to this genus thewriter had so far been compelled todepend only on the ill-preserved materials.
The specimens he had obtained. however.
were fortunately so good in preservationthat the writer v.·as strongly tempted todescribe and i11ustrate them as follows.

Dirlyozamites imamurae OtsHt emend.
Pl. 5. fig. 1 : Text·fig. 1· E'36.

lJiclyozamiles imamurae OiSHI: p. 27.
Pl. IX. rigs. 2. 2a: Text·lig. 1.
1940. fJictyozamites imamurae
OISHI: p. 326.

Mesozoic Floral Provinces{Domer ian -Albian)After K ~JURA.T., 19 59 l\1 S

ToyoraFloral c:>
Province
Index:e0<D

Korhian and !\lonobe~awan Floras
Upper Jurassic Flor'dS
llpprr L.i.tssic and Middle Juras:-.ic Floras

Text-fig. 11. i\lcsozoic (Domcrian·Aibian) Floral ProYincesof the Japanes'= Islands and Korea Peninsti.la.
(After T. Kt:o.lt'I~A. 1959 l\1S)

Emended Diagnosis: Frond.
pinnate. large. more than 20
em. long and 10 em. wide
probably
oblanceolate to
obovate in outline, contracting gradually towards thebase and more or less abrupt·
ly to\vards the apex. The
detail of the apical portion
has not been made clear yet.
Rachis slender 2-3 mm. thickmeasured on impression andalternately sending off the
pinnae at a wide angle.
Pinnae set closely together.
touching or overlapping eachother laterally: linear. slight·ly falcate. parallel-sided withthe rounded apex and seemto be convex in their uppersurface: the base asymmetri·cal. sessil and attached tothe upper surface of rachis
by a portion about 5-7 mm.
wide at the central portion:
upper half cf base •uriculated and lower base contracted:
typical pinnae. 5.5 em. long

24

Tatsuaki KIMURA

and 2 em. wide. :\"erves very crowded,
divergent, anastomosed. the reticulum
consisting of meshes ; meshes in thecentral median region about 1 em. long,the density being about 25 per em. andbecome abruptly shorter and narrower
towards the margin.
Description of specimens: !'vlany finespecimens in good preservation wereobtained. One of them is shown in Pl.
5, fig. 1 on which the present emended
diagnosis is mainly based. The rachis
seems to be buried into the rock material. The lower pinnae are strongly
rolled backwards as shown in the lefthand-side of the rachis. Text-fig. 1
shows the outline of the pinnae.
Remarlls: This species is well characterized by the more or less linear
pinnae provided \vith rounded apex andby having asymmetrical and auriculatedbase.
Dictyozamites cfr. imamurae describedby the writer in the present papar. thewriter can not identify thoroughly theformer with the present species owing
to both disagreeable characters of obtusely pointed apex and of the convex upper
margin in the former.
Dictyozamites tateitcae originally described by OisHI from the :\Taktong groupis an allied form to the present species.
But as formerly stated by OisHI (1936),
in the former species the pinnae are
elongated triangular in form and theapex of them is obtusely pointed insteadof being parallel-sided and rounded respectively, and the nervation is less
crmvded.
Occurrence: Abundant.
Reg. No. : K.\il-598lll.

!vlany specimens were obtained. Pl. 4,
fig. 1 shows a Dictyozamites frond. morethan 14 em. long and 10 em. wide. The
rachis slender. about 2 mm. across measured on impression and sending off
alternately the pinnae at a wide angle,
approximately 2.5 ern. distant on eachside. The pinnae set closely together,touching or overlapping each otherlaterally; linear. lateral margin beingnearly parallel-sided and apex obtusely
pointed or rarely rounded. The basal
part of pinnae is apparently sessil. auriculated and asymmetrical as seen insome of Otozamites pinnae. and attachedto the upper surface of the rachis by
the middle portion of the base. Nerves
are very crowded. radiated and often reticularly anastomosed and end in the wholeof the outer margin. The meshes in thecentral median region are approximately
1.0 em. long. the density being about 2528 per em. and become shorter and narrower towards the marg!n.
Pl. 5. fig. 2 ::;hows two pinnae belong·
ing to Dict_rozamites. They well repres;:;nt the reason that the writer can notthoroughly identify such many finespecimens with Dictyozamites imamurae
newly emended in this paper; pinnae
set closely, nearly parallel-sided and end
obtusely pointed apex: both margin,
especially in the upper margin, areslightly convex. This specimen is aportion especially characterized, then it
might be possible to distinguish so iar
as such a specimen from Dictyozamites
imamurae or the other known species.
Text-fig. 2 shows an outline of pinnai 11 ustrated here.
Occurrence: Abundant.
Reg. :Vos.: KM-59006. KIVI-598112.

Dictyozamites cfr. imamurae 0Isiii
Pl. .J. fig. I : Pl. 5. fig. 2:
Text-fig. 2

Dictyozamites lwwasakii TATEIWA
Pl. .J. fig. 2: Text-figs. 7-8

398.
1929.
1929.
1936.
1940.
1950.

Mesozoic Plants from the Itoshiro Sub-group. the Tetori Group

Dictyo:wmiles kawasakii
TATEIWA:
Plate. figs. 6a-b.
!Jiclvozamiles katcasakii TATE IIV A var.
grossillt'l'l'is T:\TEIWA: p. 2.
Dictvozamiles katcasakii OISHI: p. 27,
Pl. ix. fig. 4: Text·fig. 2.
Diclyozamiles katmsakii OISHI: p. 325,
Pl. XXIX, figs. •l-5.
Dict\·ozamiles kawasakii T.-\1-;AIIASIII et
NAI~O: p. 1811, Text· fig. 1.

A striking good specimens which agree
all with TATEiwA's original specimens
were obtained. Frond pinnate. large
more than 15 em. long and 8.5 em. wide.
Rachis probably slender. Pinnae alternate. reniform in outline, about 4.5 em.
long and 2.5-3 em. \Vide, with rounded
apex. and attached by a narro\v portion
about 4 mm. wide in the middle of thebase to the upper surface of therachis.
Nerves numerous. divergent,
anastomosed. I he reticulum consisting ofpolygonal meshes. Pl. '1, fig 2 shows agood preserved specimen on which theabove description was mainly based. The
rachis seems to be buried in the rockmaterial. Text-fig. 8 shows the outlineof pinnae of this species which is to be
distinguishable from Didyozamites reuiformis described in the present paper.
Text-fig. 7 might show an anterior portion of the frond. The only outline ofeach pinna impression reminds us cf
that of Podozamites or Otozamites leaves.
but its nervation is apparently divergent
and reticulately anastomosed as well as
the specimens described above.
In 19:10. T,\KAII:\SHI and ~AlTO reported the occurrence of this species fromthe Lpper most of Jurassic. Kiyosue formation. According to them. their specimens \vere not distinguishable from
Dictyozamites J:au:asakii and D. reniformis.
and these two forms were quite continuous and they could be safely united into
one species. D. hmmsakii.
The present writer thinks that the25

above statement is possibly natural. In
the writer's collection of a large quantity,
however, there have yet been found noevidence showing that the two iorms
belong to the same species. So now thewriter deals with them as distinct species respectively.
Occurrence: Common.
Reg. cVos.: Kl\1-57001. KM-5700~.

Dictyozamites reniformis 01s11t
Text·fig. 3---1

Diclyozamites ren~formis OISHI : p. 29.
Pl. IX. ligs. 1. 1a.
1940. Dictyozamiles ren~formis 015111: p. 327.
1950. ?Diclyozamites kawasakii TAKAHASHI ct
NAITO: p. 188, Text-fig. 2.
1936.

The writer's specimens. though they
are not so good in preservation, agree
all with Otstn's characteristic species
derived from Yambara (the upper courseof R. Kuzuryu, Fukui prefecture) of the
Itoshiro sub-group.
Pinnae reniform in outline. distinctly
short stalked, set closely 1ogether andfilled with reticulum consisting of finemeshes radiating from the top of thestalk and distributed in almost equal
density over the lamina.
Occurrence: Not rare.
Reg. .Vos.: K.iVI-51003. K.M-59010.
Genus Utozamites F. W. BRAt:-;In the Japanese Islands. this genus
seems to be an Older l\Iesozoic element.
flourishing in Jurassic and becoming to
decline in the early stage of Cretaceous.
Eight distinct species and several undetermined specimens have already been
described; 0. lmzismme. 0. lawifolius
and 0. molinianus from the Saragaian(mainly Norian in European standard)
strata. 0. fujimoloi and 0. mo!iniamts
from the Liassic (Lower-Middle) strata,

26

Tatsuaki KIMURA

0. beani from the Middle to Upper
jurassic strata in the North-East andthe Hida Floral Provinces respectively.

As to the Cretaceous species, only 0.
k/ipstein.ii has been known commonlyfrom both Outer and Inner side Floral

4

3

9

2

~

Text-figures 1-10

10

398.

Mesozoic Plants from the Itoshiro Sub-group. the Tetori Group

Provinces.
The specimens described by OISHI(1932) from the Shi!:aka Coal Field. Kyoto
prefecture. under the name Otozamites
k/ipstei11ii are so imperfect that his
identification has need of reexamination
about them.
.·Except some exceptional cases. it
might be admitted to consider that eachspecies belong to this genus is respectively peculiar to a certain floral province.
Otozamites sp. described by the present
writer (1958) from the Kuzuryu subgroup in Hida massif is a characteristic
form. but its specific identification has
not been done yet owing to its ill-preservation.
Otozamites dr. beaui (LI:-;ot.F.Y et
1-lu·I"I'O:-;) Blw:-;c::-<1.\RT
Text-fig. 5Comparable specimens :1940.

Olozamill·s lwanip.

OISHI:

3:l8.

Pl.

XXIX. lil{s. 7. Sa.

19-l•l.

Otozamiles l!t•tmilA-C, IE.

IIAt<IHS:

p. 419. figs.

~~

rounded or obtusely pointed apex. The
bases. though the detail is obvious be·cause of ill-preservation of material. aresomewhat asymmetrical, but not sostrongly auriculated. 1\erves probably
originating from the point of attach·ment, fine. often forking. straight anddiverging to the margin of the lamina .
The writer's specimen closely reseJ!1·
bles in general habit those described by
01sH1 from the Kiyosue formation andalso by HARRis from the Gristhorp Bed(l\liddle Estuarine, Bajocian) under thename of Otozamites beani.
The writer. howeYer. hesitates to
identify the present specimen with theEuropean species because he concerns
anxiously himself about the following
reasons. i.e.. the present specimen isimperfect in preservation and furthermore. in Otozamites. it is common thatthe outline of pinnae widely ,·aries ac·
cording to the position of a frond. then
only a part of the frond makes it identi·
fication impossible or very difficult.
Occurrence: Rare.
Reg. No. · KlVI-59005.
Otozamites endoi Kt~IL:RA sp. r;ov.

Text-fig. fi shows an imperfect portion of the frond. Rachis obscure owing
to being covered with closely set pinnae.
Pinnae broad deltoid in outline with

Pl. 5. fig. 4; Text·fig. 6

Descriptiou of species: Frond pinnate.
unknown size. more than 12 em. long

Explanation of Text-figures
Text-fig. 1: Dic/yozamites imamurae OISHI
emend: (Reg. ~o. KM-598111) x~
Text-fig. 2: Dictyozamites dr. imamurae
OISHI : (J{eg. No. KM-598112) x!
Text-fig. 3: Dictyoz'lmite.~ ren~formis OISHI:(Reg. No. KM-57003) X~
Text-Jig. ·1: Diclyozamites ren(formis OISHI:(Reg. No. KM-59010) x!
Tc:o:t-tig. 5: Utowmifes cfr. beani (LI:->I>LEYet Hurro;-.;) BIHJXG.,IART; (Reg. No.
KM-59005) X~

Text-fig. 6: Otozamites endoi K1~tt:RA sp.
nov.: (Reg. No. KM-59011) X~
Text-fig. 7': Dictyozamifes kawosakii TATE·IWA: (Reg. No. KM-57002) x~
Text-fig. 8: Dictyozamites lwl!'osakii TATE·
1\L\; (Reg. No. KM-5i00l) X~
Text. fig. Y: Otozamiles sp.: (Reg. No.
KM-59002) X!
Text-fig. 10: Otozamifes sp.: (i{eg. No.
Kl\1-59001) X~

Tatsuaki KIMURA

28

and 5 em. wide. Rachis slender about
2 em. across measured on impression.
Pinnae elongated OYal in shape, approxi·
mately 5 mm. long and 1-1.3 em. wide at
the widest portion, apex rounded andbase contracted but the upper baseslightly auriculated; attached to theupper surface of the rachis "\vith thelower half of base and strongly directed
forward; set closely together. overlapping each other laterally. Nerves fineand delicate, originating from the pointof attachment. parallel each other near
the base and radiating to the whole ofmargin. not anastomosed.
Pl. 5, fig. 4 shows a specimen on
which above diagnosis was based andText-fig. G shO\vs the outline of pinnaof this characteristic species.
Remarks and Compm'ison: A single
figured specimen has been examined.
The writer's species is well characterizedby having elongated pinnae strongly
directed forward. and there is no comparable species of this genus. though
the specimen is not a satisfactory one.
An anterior portion of 0/ozamites graphicus lLECKDiBY) figured by HARRIS(19-19. p. 561. Fig. 2A) somewhat resembles the present specimen in outline of
pinnae, but smaller and more strongly
auriculated than the latter.
Otozamites lancifolius originally clescribecl by OisHI from the Saragaian
Nariwa group is allied form to the present specimen, but the former is different in having acutely pointed apex andrepresenting coarse nervation from thelatter.
It might be possible to obtain further

good specimens than the present one
from the same locality sometime infuture.
Occurrence: Probably rare.
Reg. No.: lGvl-59011.

Otozamites sp.
Text-figs. 9-10
Large specimens. though they are imperfect, were obtained as shown in Textfig. 10. Frond long and narrow. nearlyparallel-sided, more than 30 em. long andprobably 7 em. wide, both of apical andbasal portion missing. Rachis considerably thick, 3 mm. across measured on
impression. Pinnae alternate, oblong oroval in shape, apex bluntly rounded,
about 3 em. long and 1.5 em. wide at
the widest portion: bases almost symmetrical and not auriculated and attachedto the upper surface of the rachis by
their middle portion. Text-fig. 9 shows
probably an anterior portion of thefrond: pinnae seem to be short petioled.
Nerves indistinct because of ill-preservation of material, but seem to be forking near the broad point of attachment,
nearly parallel. ending in the outermargin and considerably coarse in density.
The present specimens closdy resemble Otozamites kondoi described by 0Is!IIfrom the Mone formation (Upper Jurassic) at the northern Pacific coast of theJapanese Islands, in the shape of frond
and also of pinnae, mode of attachment
to the rachis and the nervation. But
the former species is approximately
t\Yice as large as the latter.

Explanation of Plate 4
Fig. 1.
Fig. 2.

Dictyozamites cfr. imamurae 015111; (Reg. No. KM-591:1112)
fwwasakii TATr:I\\'A; (Reg. No. KM-51001)

Dic 1yaz:ml;!e.~KrMUJ?A :Mesozoic Pla;1i$
Plate 4

398.

Mesozoic Plants from the Itoslziro Sub-group. the Tetori Group

Otozamites molinimms Z1c:w• is an another allied form to the present specimens. But in the former species. thebase of pinna is slightly auriculated
instead of symmetrical and the size of
pinna is smaller than the latter.
It also can be expected to obtain
more good specimens than the present
ones from the same locality.
Occurrence: N'ot rare.
Reg. l'los.: KlVI-59001. KM-59002.
CYCADOPHYT A
Cycadales
Genus :Vi!ssouia (BRo:->G:-:IART)
Jl.'ilssonia koloi (Y oKaY A~IA) 01sm

2~J

ed surface of the rachis to its medianline.
This species closely resembles the
specimens described by Yo~<oY .\:O.IA andYABE and 0Isu•
under the name of
Nilssonia sinensis from the Jurassic
strata of Manchuria (Sha-ho-tsu andWei-chia-pu-tzu, Liaoning). and Jurassic
strata of Ussuriland and \·ladiYostok(KRYSHTOFO\'JCH: 1916, 1923L
It is the writer's preliminary opinion
that these Japanese and the continental
specimens might possibly belong to asame species. This species has net yet
been recorded from the Outer side Floral
Province of the Japanese Islands in thosedays.
Occurrence: i'\ot rare.
Reg. No.: KM-51111.

Pl. 5. fig. 3
p. ,1~. Pl.
VII. figs. la-c. c: Pl. XIV. fig. 14.
1905. Diurmit"s ? sp. YM3E: p. H. Pl. III.
fig. 7.
1905. Cteuophyllum ? sp. YABE: p. 15. Pl. IV.
fig. 7.
19-lO. .Yilssonia koloi 0ISIII: p. 302. Pl. XXV.
figs. 3. 3a: Pl. XLIV. fig. 3b.

1889. Dioonites kotoi

Genus Ctenis LI:->OLF.Y et I·h-TTo:->

YoKOY:\~IA:

The writer's specimens are closely referable in general feature to those originally described by YoKOYAMA from
Kuwashima (Shimamura) and Tanimuraof the Itoshiro sub-group. Frond simple
and delicate. l~achis slender. being l
mm. wide measured on impression andnarrowly grooved longitudinally on theupper surface. Lateral lamina deeply
cut up to the rachis into long and narrow segments; segments opposite oralternate, lightly curved and directed
forward. l\'erves considerably tine. mostly simple or rarely once forking near
the base. parallel to each other and to
the upper margin of the segment. running obliquely downwards in the groov-

Ctenis lumeharai YoKoYA:<.IA
Pl. 6, figs. 1-2

1906.

Ctenis kmu:harai Ym,oYA~IA: p. 2~1. Pl.

1933.

Clenis kaneharai YABE et O!Sifl: p. 32.
Ctenis kaneharai OISHI: p. 256, Pl.

IX. figs. 1, la.
19JO.

1950.

XXIV, fig. 1.
Ctenis kanelwrai HARRIS: p. 1001. Text·
figs. lA-C. 2A-C, 3A-B.

Description of sjJecimem: Frond large,
unknown size. pinnate. Rachis attaining
a width of 0.6 em. across measured on
impression. the ornamentation of its
Pinnae apparently
surface not clear.
subopposite. attached to the lateral sidesof the rachis, either in contact or slightly
separated in the lower part of the frond,
becoming more widely spaced in the
middle and upper part of the frond.
Pi1:nae arising nearly at right angle to
the rachis. Pl. 6. fig. 1 shows the middle or upper part of the frond which is
closely referable to the specimens des-:_\(1

Tatsuaki KIMURA.

cribed originally by Yo.-oYAM.\ from~ien-tzu-kou Cml-field (l\liddle Jurassic?). L!9.o;::ng, ;\Ianchuria, by OrsH:
fro:n the Kuwashima formation and by
HARRIS from the Lower Estuarine; pinnae
elongated, somewhat inclined forward,
mostly parallel-sided, contracted at theb3.se. never expanded; the length of pinnae
is not known, as they are not wholly preserved, but the width in one of thE:m is
30 mm. which diminishes to 1i mm. atbase.
Owing to its intimate occurrence withthe spedmen described above, Pl. 6, fig.
2 seems to show the IO\ver part cf
the frond of the same species, thoughsuch a type of pir:nae has not been recorded so far; pinnae set closely together,
9.5-10.5 em. long and 3 em. wide at the
middle portion. tapering to an acuteapices and more or less contracted atbase. :\'erves strong near the base. becoming comparatively fine to\vards theapex, often dichotomizing and also uniting so as to form elongated meshes.
Density 9-10 per em. at the base, rather
more crowded, 13-1ii per em. near theapex.
Remarks: In this species, the apicalnatur-e of pinnae was more cleared upby the present specimens. and still more,
it might be possible to obtain the complete specimens from the same locality
in future.
In 1950, HARRrs emended the diagnosisof this species adding the detail of its
cuticle. In the Japanese specimens, it isregretted that an anatomical study has

not yet succeeded on account of thestate of preservation. in spite of thewriter's exertion.
In the japanese Islands, this species
has only been known from the Kochianof the Inner side Floral Province.
Occurrence: ?\ot rare.
Reg. Nos.: KM-59101, KM-57101.
GI~KGOPHYTA

Genus Phoenicopsis HEER
This Ginkgoalean genus is considered
to b;:: a conb:2ntzl one which has been
widely known from the l\11esozoic strata
of Siberia, tvianchuria. the Northern andWestern part of China, Korea and theArctic region, whi Ie it has been rare in
the Japanese Mesozoic.
In the Japanese Islands, Phoenicopsis
spp. have been described by OisHI (193~)
from the Norian Nariwa group and also
by Otsllr (1931) and Kr~tl:RA (1959) respectively from the Lower-Middle Liassic Kuruma group. These were all
impressions of incomplete and detachedleaves. The following genera, i.e .. Czelummcs!?ia, Hartzia, TVindwardia, Torrellia
and Phoenicopsis. possess long, slender,
ribbon-like leaves growing in clusters
upon scaly dwarf shoots. Without researting into the epidermal and stomal
characters, it is very difficult to distinguish one from another in connection\vith the above genera, while dealing
only with incomplete or detached leaves.
The generic name of Pfwenicopsis being

Explanation of Plate 5
Fi~. 1. Dict_vozamites imamurae 0!5111 emend; (Re~- No. K.M-598111)
Fig. 2. Dictyozamifes dr. imamurae OiSHI; (Reg. Xo. KM-598113)
Fig. 3. 'Vilssonia kotoi (YoKOYAMA) Orsr11: (Reg. ~o. Kl\l-51111)
Fig. -1. Otozamites e11doi Kr~u;RA sp. nov.: (Reg. No. KM-5901 L)

Plate 5
KIMURA :Mesozoic Plants

398.

Afesozoic Plants from the Jtoshiro Sub-group. the Tetori Groupgiven to the japanese specimens alreadydescribed. they are probably referable
either to the well-known species in theJurassic of Eastern Asia, PltoenicoPsis
rmf.;ustifolia HEEl~ or to P. mandmris
originally described by Y.\BE and 01s11J.

PlwenicoPsis ? sp. cfr. P. sPeciosa HEEl{
Pl. 6. ftg. 3

Only a specimen shovm in Pl. 6. iig.
3 was obtained. This is represented bycluster of detached leaves attaining 13
em. long and 0.6-l.Ocm. wide. narrowing
very gradually into slender basal portion.
ap:::x is nussmg. Nerves are distinct.
parallel. about 15 in number and faintly
marked interstitials bet\veen each pairof them being able to seen.
In the present specih1en. it is regreted
that both the apical and the basal portions are broken away but the general
habit of the leaves reminds us of Plwenicopsis. especially Phoenicopsis sPeciosa
originally described by HEER from theJurassic of Siberia.
The specimens described by KAwAsAKI(1925) from the Daido Series of Korea
under the name of Plzocmicopsis speciosa
is an allied form to the present specimen.
The designation titled above being precisely given to it. the present specimen
is the youngest record of this genus in
Eastern Asia. At any rate. this genus
is a continental element and rare in the
Jap:inese Islands and it has not yielded
yet in the Outer side Floral Province.
Occurrence: Very rare.
Reg. !Vo.: K!VI-57008.
Addendum

The writer has been conducting researches on the occurrence of fossil
plants. especially on the relation between
the stratigraphical horizon and its bear-:31

ing species.
As to the o::currence of fossil plants, wehave more or less known that for instance.
tree or shrub species was dominant in acertain horizon, while grass species was
dominant in another horizon. According
to the writer's experience. good example
showing the above fact can be seen in
the Iwamuro format:on (Lower-Middle
Liassic) and also in the lloshiro subgroup studied in detail.
So far as at least the sphere of geological unit of a m~mber or a fo:-mationis concerned. the d:stinction of theyielded species among various horizons
should be explained so much by thevertical range of plant evolution as by
the rise and fall of the sedimentary
ba6in having much to do with the veget'ltion and its variation growing aroundthe basin.
It is interesting to note that tree andshrub species are dominant in the present
locality, though such is not the case in
its lower horizons. The detail will be
stated by the writer sometime in future.
References
AIC\<>LD. C. A. (19·!7). An Introduction to
Paleobotany. Nell' J'ork.
HARIHS, T. M. (19-14). Notes on the Jurast;ic
Flora of Yorkshire. 10-12. ..Inn. & .Uag.
Nat. !li$1 .. Ser. 11, Vol. Sf.
- - (1949), Notes on the Jurassic Flora ofYorkshire. ·10-42. Ibid., Ser. 12. Vol. Il.
- - (1950), Notes on the Jurassic Flora ofYorkshire. 46-48. Ibid .. Ser. 12. Vol. Ill.
XA\\.\SAKI. S. (1925). Some Older Mesozoic
Plants in Korea.
Geol. Sttrt'. Cho$ell(Korea), Vol. IV. Pt. 1.
Kl~llli~A. T. (1958). Mesozoic Plants from the
Kizaki District. Nagano Prefecture. Japan.
}ubil. Pub/. Comm. Prof. FuJNtoro, Tokyol..·nit•. Educ.
- - (1958). On the Tetori Flora (Pt. 1).
~lesozoic Plants from the Kuzuryu Sub·

Tatsuald KIMURA.

32

Group. Tetori Group. Japan. Bull. Seu
1/igh Sch. Tok.m U11it•. Educ .. No. Il-2.
- - (1959). Mesozoic Plants from the Kotaki
Coal-Field. the Kuruma Group. Central
Honshu. Japan. Ibid., No. Ill.
- - (1959). Preliminary Notes on the Liassic
Floras of the Japanese Islands. Ibid.
- - (1959). On the Tetori Flora (Pt. 2).
Addition to the Mesozoic Plants fromthe Kuzuryu Sub·Group, Tetori Group.
japan. Ibid.
- - (1959), On the I\'lesozoic Floral Provinces
Preliminarily Found in the Japan.::seIslands. Tokyo (MS).
KRYSHTOFO\'ICH. A. (1916). Materials from
Jurassic Flora of Ussuriland. Trar. Mus.
Geol ..\lin .. SI.·PNerslm1·g, Fol. II.
- - (1923), Equivalent of the Lower Jurassic Beds of Tonkin, near Vladivostok.
Rec. Geol. Comm. Russ. Far East, Xo. 2.
0JSIII. S. (1931). Mesozoic Plants from KitaOtari. Prov. Shinano. Japan. Tour. Fac.
Sci., Hokkaidu Imp. Unit•.. Ser. I 1·. Vol. [,No.2.

- - (1932). The Rhaetic Plants from the
Nariwa District, Okayama Pref., Japan.
Ibid., Vol. II. ..Vos. 3-4.
- - (1932), The JurassiC Plants from Shitaka(the Maizuru Coai·Field). Prov. Tango(Kyoto Pref.). japan. Ibid., Vol. II. Nv. 1.
- - (1936). On the Japanese Species ofDictyozamites. fap. four. Geol. & Geogr.,

Vol . .Yill. Nos. 1-2.
- - (1!'40). The 1\·lesozoic Floras of Japan.
jour. Fac. Sci.. Hokkaido Imp. Uniz· .. Ser.
I v; Vol. V. Sos. 2-4.
SEWARD. A. C. (1917). Fossil Plants. Vol. Ill.
Cambridge.
- - (Hll9). Fossil Plants. Vol. IV. Ibid.
TAKAIIASlll. E. and \'AlTO, G. (1950), Dictyo·
zamites from the Jur~ssic Toyopjshi
Serbs of Prov. ~agato. (Yamaguchi Pref.).
four. Geol. Soc. }open. Vol. 56. Z1lo. 188.
T ..\TEIWA. I. (1929). Geological Atlas ofC,hoscn (Korea). \'o. 10, Keishu·Eisen·
Taikyu and Wakwan Sheets. Geo/. Sun•.
Chosen (Korea).
YAnE, H. (1905). Mesozoic Plants from Korea.
four. Coli. Sci .. Imp. Uuit•. Tok.vo. Vol. XX,
tlrl. 8.
YABE, H. and OISHI. S. (1?29). Xotes on Some
Fossil Plants from Korea and China Be·
longing to the Genera 1\'ifssmtiu and Ptero·
phyllum. fap. jour. Geol. & Geogr., T'of.
VI ..Nos . .1-.J.
- - (l!l33). Mesozoic Plants from Manchuria.
Sci. Rep.. Tolwku ln:p. l.!nir.. Sec. Ser.
(Geol.). Fol . .\II . .Vo. 2B.
Ym;oYA,..,tA·, M. (1889). Jurassic Plants from
Kaga. llida and Echiz~n. four. Col!. Sci.,
Tokyo Imp. l..'nit•.. Vol. Ill. Art. 1.
- - (1906): Mesozoic Plants from China. !hid..
Vol. XXI. Art. 9.

Explanation of Plate 6
Fig. 1. Ctenis kamltarai YoKoYAli!A: (Reg. No. KM-59101)
Fig. 2. Ctenis kane!Tarai YoKOYA~tA; (Reg. ~o. KM-57101)
Fig. 3. Phoenicopsis ? sp. cfr. P. speriosa HEER: (Reg. No. Kl\1-59112)(All specimens illustrated here are in natural size)
Photogr. by S. AoKI
All spJcimens described here are deposited in Department of Geolog ,. of the Komatsu Cityl'vluseum. Ishikawa Prefecture.

{(fMUJV J. :Mesozpic Plants

Plate 6

Trans. !'roc. Palacont. Soc. Japan, N. S .. No. 41. pp. 33-40. pis. 7-8. Apr. 10. l!llil:~9!1.

TvVO NEW SPECIES OF THE PARAFUSULINA YABEITYPE FROM TOl\HlRO. TOTIGI PREFECTURE.
CENTRAL JAPA:\ROKURO MORIKAWA
Department of Earth Sciences. Saitama UniYersity
and

YOSHINARI TAKAOKA
Toho Gakuen

1/jj*g,u::;:¥ .J: I) Parafusuliua ?rJQ) 2 ~1i:

Wj;;f:~lj!.~g:~ Parafusulina yabei WJ.Q) 2
Parafumlina tomuroensis, P. nabeyamensis Q) 2 Jrf!f!~~cJt!lt. #-?"( IJ•iKtJ~!>K.~' .J: IJ
Parafusulina kaerimizensis .!::: L "(jS~,ilQ) '(, Q)~¥it'i!r L t~t;:.:: .:h-lt Parafusulina tomuro·ensis "t'iif.t:~·IJ•c.fl!:l.') ~ >;.:.:.;.f>;:ml.:.:t;;~·-cifl:ll~n-ct-·~ Parafusuliua
2 -?Q):/If..-;/'. P. yabei 1\1!. P. japonica ~l.:.:btt. P. yabei W!J.r -1 -;/'Q) Parafusuliua t.:.~_:.t-n ·~-:.!:::
a-~ ct.: o~ Jll ;'"; I~> • i:'Ji IMJ ,¥f 1111:
~[IIi

0

ll is well known that Tomuro, Totigi
Prefecture, yields abundant fusulinids andmany authors have described them. Y ABE (189\l) first described Fusulina japonica
Gc:-.JBEL var. Subsequently. HA:-.zAwA(1942) restudied it and proposed for it a
new specific name. Parafusulina yabei.
OzAWA (1925) described Parafusulina
kaerimizeusis from the same locality, including the specimens of P. lwerimizensis
from Kaerimizu. Recently ToRIY,\~JA. in
his monograph of the Fusulinids from
Akiyoshi. made a restudy on P. kaerimizensis in detail. According to him P.
kaerimizensis reported by OzAWA fromTomuro differs from the holotype of
Kaerimizu.
The specimens from Kaerimizu resemble the ones from Tomuro in the elongate
form, size of proloculus. rate of volutionexpansion, but compared the latter. theformer has heavier axial filling and. lessintense ftut ing of septa. esp~cially in thepolar regions. As above cited. they area-

not conspecific with each other. We
should like to propose a new· specificname. Parafusuliua !omuroeusis for Parafusulina kaerimizensis reported by OzAwA
from Tomuro. Parafusuliua lomuroensis
rather resembles Parafusu!illa yabei inits strong and regular fluting of septa.
and their thin septa. especial!~· in thepolar regions and in that the chamberlets
reach the next spirotheca. but the former
is more elongate. and has a smaller proloculus than the latter.
We discovered another new species
described in this paper as Parafusulino
11abeyamensis: this is inflated fusifom1,
and some of the species sho\\· distorted
fluting of the septa. its fluting resembles
that of Parafusulina tnwcata.
Parafusulina yabei. Paraf. tomuroensis
and Paraf. nabeyamensis arc closely allied
to Parafusulina n·ordensis from North
America in their thin spirotheca andsepta. especially thin in the polar region.
and strong septal fluting and in that the

:34

Rokuro MoRIKtlTV:l and Yoshinari T-lK:lOKAchamberlets reach the next spirotheca.
Beside the above cited species, thefollowing species from Japan resemble
Parafusulina wordensis in their import·
ant characters.
Parafusulina yabei HA:'\ZA\1".\, HA:'\Z.-\WA 19•12.
Loc. Kuzu.
Parafusulina yahei IIA:"ZAWA, MORIKAWA
1898. Loc. Akasaka.
Parafusulina HA~ZAWA, !Go, 1959. Loc. Hi·
rayu.

Parafusulina kiuosakii (i\foRtKA\YA). MRtKA·\\"A 1958.
Loc. Akasaka.
Parafusulina tnmcata (OZA ,,.A). leo et Oc;A w ,\1958. Loc. Funafuseyama.
Parafusulina japmzica var. tnmcata (0ZA WA),OzAWA 1927. Loc. Akasaka.
Parafusttlina japouica Yar. tnmcata. leo 1959.
Loc. Hirayu.

Parafusulina truncata (OzAWA). leo ct 0GA·
w .... 1958. Loc. Funafuscyama.
Parafusuliua tnwcala (OZAW."\). l'vloRtK."\WA1958. Loc. Akasaka.
Pamfusulina uakamigawai l\foRtKAWA andHoRIGUCHI. 1956.
Loc. Kuzu.
Parafusulina 11afwmigmmi !Go and OG.-\WA
1957.

Loc. Funafuseyama.

Parafusulina iisakai leo and OGAWA. lc;o and
OGAWA 1957. Loc. Funafuseyama.
Para[usuli11a amzamitheca (DEPRAT) 1914. Loc.
Akasaka.
Parafusulina uenoensis KoBAYo\SIII, KonAYA·
SHI

1957.

Loc. Hirayu.

Sclucagerina giganlojapmzica KoBA YASflt. Ko·
DAYASHI 1957. Loc. Jbuki.
Sc/umgerina ilmlliensis KOBAYASHI. KOBA YASlll
1957. Loc. lbuki.
Parafu~ulina hirayuensis !Go, Jt;o 1959. Loc.
Hi;ayu.

Pamfusulina ? sp. !Go. 1959. Loc. Hirayu.
Parafusulina sp. A. KOBAYASHI 1957. Loc.
lbuki.
Pamfusutina sp. leo and 0GA w A 1958. Lee.
Funafuseyama.

Para[usuli11a ambigua (DEPRAT), KonAYASHI
1957. Loc. lbuki.
Sclumgerina sp. aff. S. japonictl (GUMBEL)
var. chzcta (REICHEL) leo 1959. Loc.
Hirayu.

Pseudofusulina crassitectoria (DV:-\BAR ct SKI:-\·:'\ER). KonA YASHI 1957. Loc. Ibuki.
Parafusulina ozau:ai (ltSAKA). ltSAKA 1932.
Loc. Nyukawa.

Parafusulina nakamigaU"ai is longer
than Parafusulina tomuroensis and resem·
bles Parafusulina umrdensis. but has nostrong axial tilling.
Parafusulina tnmcata resembles Para·
fusulina nabeyamensis but is shorter thanthe latter. Parafusulina kinosakii has asmaller proloculus than Parafusulina
yabei. Parafusu!i na annamitlzeca (notoriginal) from Akasaka described by
DEPRAT has stronger axial filling, and alarger proloculus than Parafusulina kinosakii. Parafusulina uudata from Akasaka
resembles Parafusulina yabei. but has alarger and more inflated fusiform shelland thick spirotheca.
Parafusulina uenoensis and Parafusuliua
gigantojaponica from Ibukiyama may bereferable to Parafusulina yabei in theirimportant characters. Parafusu!ina ibukiensis is more inflated fusiform and has
a larger proloculus than Parajusulina
yabei. Parafusulina !1irayuensis is elongate fusiform and resembles Parafusulina
iisalwi. Parafusulina /i[t)'ashii has loosely
fluting septa and its chamberlets arerough if compared with Pamfusulina
yabei. Parajusulina? sp. described by!Go from llirayu is a so ill-oriented section that its specific position can not be
determined.
However it very closely
resembles Parafusulina hirayuensis andPamfusulina iisakai.
Parafusulina japonica is one of themost common species of Schwagerinids
in Japan. It differs from Parafusulina
yabei and its allied species in its thickand less intensely fluted septa and thickspirotheca. The following species inJapan resemble Parafusulina japonica intheir important characters.

339.

Ttvo new species of tlze Parafusulina yabei type from Tomuro

P.uafusulina japonica (GC~IUEL). Sell II' ACE R
1883. Loc. Akasaka.
Para.fusuliilajaprmica (GU~tllEL). OZAWA 192:5.
Loc. Akiyoshi.
P.trafusulinajapmzica (GO~tREt.). OZAWA 1927.
Loc. Akasaka.
Para.fusu/ina japanica (Gt:~tBI".). FL'JI~IOTo
1936. Loc. Kanto.
Para.fusulina japonica (Gt:~tBEl.), l\'ioRIKAIVA
1958. Loc. Akasaka.
Parafusulina japnnica (G L'~IUEI.). KollA YASHI
1957. Loc. lbuki.
Parafusulina d. japouica (GU~tllEL), SAIC\·GA~II 1959. Loc. Tama.
Sdm•agerina japonica \'ar. lwyasakai (LEE).
FuJt~IOTo 1936. Loc. Kanto.
Schwagerina gigantojaponica KooA L\SHI, :\Io·
RIKAWA 1955. Loc. Akasaka.
Parafusulina tomeganensis :\loR 1K ,, w A. :\fr,R I·
KAWA 1958. Loc. Akasaka.
Parafusulina kaerimizensis (OzAWA), KA"t;:O.IA
1959. Loc. Oppara.
Parafusulina mizu/anii MoRII\AII'A, MoRIKAWA
1958. Loc. Akasaka.
Parr(fusulina laniyashikiensis Mo11 1IC\ w .\. :\!IoRIK.-\1\'A 1958. Loc. Akasaka.
Parafusulina japonica var. kinsyoensis MoRI·
KA\1'.-\, lVIoRIKAWA 1958. Loc. Akasaka.
Parajusulina lutugini (SCHELt.II'IE:-1). OZAWA
1925. · Lee ..-\kiyoshi.
Parafusulina /utugini ToRI'> ·\~L·\ 1958. Loc.
Akiyoshi.
Parafusulina sapperi (STAFF). KoKAY.-\SHI
19;>7. Loc. !buki.
Parafusulina gig::s KonAYA~lll, KollAYASHI
1957. Loc. lbuki.
Parc~fusuliua ambigua (DE PH AT). l\loRI "" WA1958. Loc. Akasaka.
P<trtzfusulina ambigua (DEPJ~AT). OzAwA 1925.
Loc. Akiyo;;hi.
Parafusulina ambigua (DEPRAT), OZAWA 1927.
·Loc. Akasaka.
Parafusulina ambigua (DF:PR.·\T), FUJI:-IOTO
1936. Loc. Kanto.
Parcfusulma am!tigua (DEPRAT). KA:"l:~IA1959. I.oc. Oppara.
Pttrt{fusu/ina exifis (SCHWAGER). SCIIWAGER
1883. Loc. Akas1ka.
Parafusulina exilis (SCIIWAGEII), Dt-:PRA'< 1914.
Loc. Akasaka.

:35

Parafusulina exilis (SCII\\'AGER), OzA\1'.\ 192:5.
Loc. Akiyoshi.
Parafusulina exilis (SCfiWAGER). :\IORIKAII'A
1958. Loc. Akasaka.
Parafusulina exilis var. takeii :\loRIK.'!. 11'.-\.
:\loR 1K A w A 1958. Loc. Akasaka.
Schwagerina gw·mbeli var. Pseudoregularis(De:xA.-\R and SKI:":XER) KoB.-\YASHI 1957.
Loc. lbuki.
Parafusulina ;.;uembeli var. pseudoregularis
KOBAYASIII 1957. Loc. lbuki.
Para[usulina l'il'iiihofmzi (ScHWAGER). Oz ... IVA
1927. Loc. Akasaka.
Para[usulina cf. kallaensis (SCH\YAGER), OzAW.\ 1925.
Loc. Akiyoshi.
Parafusulina subobsoleta var. okuboensis (OzAWA). OzAWA 1927. Loc. Akasaka.
Parafusulina okuboe11sis (Oz.\ w A) emend MoRIKAWA. :\1oll!KAWA 1958. Loc. Akasaka.
Para,litsulina sp. a. KoBAYASHI 1957. Loc.
Ibuki.
Para.fusulina sp. a. ToHtYA~IA 1958. Loc. Akiyoshi.
Para.fusulina KAl'~IEH:\ 1954. Loc. Kumamoto.
Parafusulina sjJ/cmtlens (DU:'\13AR and SKt:"!'lE R),:VIoRIKAII'A 1958. Loc. Akasaka.
Parc~fusulina untlata MoRIKAWA, :VloRIKAWA
1958. Loc. Akasaka.
Parafusulina gigantea (DEPRAT) ?, TORIYA:\IA
1958. Loc. Akiyoshi.
Parafusulina ecloensis (OZAWA), OzA\\'A 1925.
Loc. Akiyoshi.
Parafusulina ecloensis (OZ.-\1\'A), TORIYA:'>IA
1958. Loc. Akiyoshi.
Parafusuliua kaerimizensis (OZAWA). OZAWA
1925. Loc. Akiyoshi.
Para,(usulina kaerimizensis (OzAwA), FL'Jt~ro­
TO 1936. Loc. Nyukawa.
Parafusuiinrt kaerin;izensis (OzAwA). ToRIYAI'-IA 195S. Loc.;. Akiyoshi.
Parafusuliua kaerimizensis (Oz_.\ w A). .\loRIKA \\'A 1955. Loc. Shomaru.
Para.fusulina kaerimize11sis (OZA 11· A). KoB.-\YASHI 1957. Loc. lbuki.
Parafusulina paraguembeli (.:\ioRIK.\ w A). l\IoRIKAW:\ 1955.
Loc. Shornaru.
Para[usulina odakaii MORIKAWA 1955. Loc.
Shornaru.
Pr.rafusulina gifuensis :\loR IKA IL\. :\1oR 1K:\WA 1958.
Loc. Akasaka.

3G

Rokuro MaRl K -l WA and Yoslzinari T.li\AU/\.1

Parafusulina cayeuxi (DEI'RAT). l\loRrr<AWA
191:8. Loc. Akasaka.
Para,titsulina sp. B. ? !Go IY:iY. Loc. Hirayu.
Pamfusulitta matsubuishi F! · J r:o.roTo 19:C6. Loc.
Kitakami.

We tentatievly divided the above listed
species into two groups. typified by
Pamjitsu!iua yabei and J>arafusu!ina japonica. Parafusu!iua ya/Jei and its allied
species resemble Parafusufina t!'ordensis.
the type species of Parafusu!ina. These
two groups are considered to have been
derived from different stocks phylogenetically. namely, the P:tr(Jfusulina japoniw
group is thought to have come from
Pseudofusulina, whereas, the Parafusu·/ina yabei group from Quasifusu/ina.
japanese students on fusulinids seemto emphasize too much importance ofthe strong septal liuting and apparently
do not recognize the value of their thinspirotheca and septa. Accordingly, it ismost desirable to restudy Prtrafusu!ina
with concern to the abo\'c mentioned
characters.
Here we express our thanks to Prof.
Haruyoshi Ft.:JI~IOTD of the Yamagata
Uni\·ersity for his kind guidance. Our
thanks are due also to Dr. l\'Iosaburo
KAc-;c~rA. As.<>istant Professor of the Tokyo Gakugei Daigaku, !'vir. Hisayoshi !Goand Saburo 1\r.~AGt of the Tokyo Uni\'ersity of Education for their encouragement. Their thanks also due to Prof.
Kotora HATAI of the Department of Geology, Tohoku University for reading of
this manuscript.
Parafusufina tomuroensis l\IoRrr' ,\ w A
and T.·\KADKA, n. sp.
Pl. 7, Figs. 1-7.

Descriptioll .-Shell large, su bcylindricalfusiform. with nearly straight to slightly
curved axis of coiling, rounded poles, andslightly convex lateral surfaces. Large

specimens of seven to eighi volutions(commonly eight1 <:re 9.0 to 12.5 111111. longand 3.8 to 45 mm. wide. giving form
ratios of 3.3 to 4.1. The first volution
is subcylindrical fusiform. and the shell
maintains closely similar shape throughgrowth.
Proloculus of medium size. with outside diameter of about 0.2 to 0.4 mm.
averaging 0.3 mm. for six specimEns.
Shell rem<.~ins rather loos~ly coiled
throughout. Chambers are about uniform
in height in a central third of the shell
but become slightly higher as poles areapproached. Average height of chambc:rs
above the tunnel in the first to seventh
Yolution in live specimens is 0.08, 0.16.
0.20, 0.26. 0.30. 0.32 and 0.36 mm respectively.
Spirotheca thin and finely alveolar inall volutions. Spirotheca of same thickness throughout its length. Average
thickness of the spirotheca in the firstto seventh volution in five specimens is
0.02. 0.03. 0.04, 0.05. 0.05, 0.06 and 0.06 nun
respectively.
Septa closely spaced and highly fluted
throughout the length of shell. Fluting
brings the septa in contact with eachother for about half their height. Septal
intervals in outer volutions are 0.8 mm.
Tunnel narrow with slightly irregularpath. Intense lluting of the septa maketunnel sides dlllicult to identify in allparts of all specimens.
Chomata indistinct.
Axial filling commonly fills completely
the chambers along the axis from near
the middle to the ends.
~leasurements of this species are given
in Table I.
Comparison:-Parafusulina kaerimi zen sis
which was reported by OzAwA in 1925
from Tomuro is a different species
from 1\aerimizu, Akiyoshi. according to
ToRrYA~''' tl~l!iS\ and may be an elongate

339.

Two new species of the Parajusulina yabei type from TonUtro

form of Parafusulina tomuroensis. Pamf.
tomuroensis resembles Parafusulina yabei
but is distinguished from the latter by
the smaller proloculus and more elongate
subcylindrical form. Although it is alsosimilar to Parafusulina 1z.abeyamensis it
is distinguished therefrom by its elongate subcylindrical form.
Occurrence :-Okada Quarry, Nabeyama.
Parajusulina nabeyamensis lVloR rK A w A
and T AKAOKA, n. sp.
Pl. 8, Figs. 1-9.

Description :-Shell large, inflated fusiform, with nearly straight to slightly
curved axis of coiling, round poles, andslightly concave lateral surface. Large
specimens of seven to eight volution are
6.5 to 8.0 mm. long and 3.3 to 4.5 mm.
wide, giving form ratios of 1.8 to 1.9.
First volution fusiform and the shell
grows gradually throughout its growth.
Proloculus of medium size with outside diameter of about 0.24 to 0.36 mm.,averaging 0.34 mm. far seven specimens.
Shell regularly coiled throughout
growth. The chambers expand slowly
in height but become rapidly higher
along the axis. Average height of chambers above tunnel in the first to seventh
specimens is 0.06, 0.14, 0.22, 0.26, 0.30,
0.32, and 0.32 mm. respectively.
Spirotheca thin and fine alveoli develops through all volutions, its thickness from first to eighth volution is 0.04
0.04, 0.03, 0.04, 0.05, 0.08 and 0.08 mm.
respectively.
Septa thin and reach outer spirothecaand some of them have distorted fluting,
septal fluting strong and regular. The
intense fluting of the septa makes thetunnel side difficult to identify in allparts of all specimens. Septal intervals
about 0.8 mm in outer volutions. Chomata
indistinct and secondary deposit fills

37

from first to fourth volution.
Measurements of this species are given
in Table 2.
Comparismz :-This species is charac·tcrized by the large shell inflated fusiform. thin septa and spirotheca and axial
deposit which fills both sides of the proloculus as in Parafusulina yabei. Rut the
septa in some specimens are distorted to
one direction and the septal fluting isstrong. The present form differs from
Parafusulina yabei in its inflated fusiform
and smaller proloculus.
Occurrence :-This is abundant in Tomuro.
References(1914), Etude des Fusulinides
du Japon, de Chine et d'Indochine et classification des calcaires a fusulines (III.Ivlemoire). Etude comparative des fusul·
inidcs d' Akasaka (Japon) et des Fusulinides de Chine et d'lndochinc. Indochine
Gcol. Mtfm., vol. 3, fasc. 1, pp. 1-45, pis.
1-8.
Du"'>HAR, C. 0. and SKI:":"I::R, j. W. (1937),
New Fusulinids genera from the Permianof west Texas. Am. four. Sci., 5th Ser.,
l'Of. 22, pp. 252-268, pl. 1-3.
FUJI:'>IOTO II. (1936), Stratigraphical and Pal·
aeontological studies of the Titibu system of Kwanto Mountainland. Part II.
Paleontology. Tokyo Btmrika Univ., Sci.
Rep., Sec. C. I, pp. 29-125, pis. 1-26.
ll.o\:"ZAWA, S. (1924), Para_lilsztlina yabci n. sp.
from Tomuro, Shimotuke province, Japan.
fap. ]our. Geol. Geog., val. 17, No. 4, pp.
127-131, pis. 13-14.
leo, H. (1959), Some Permian Fusulinids fromthe Hirayu District, SouthcasHr.l part of
the Ilida Massif. Central japan. Tokyo
Univ. Education, Sci. Rep .. val. 6, Sec. C.,
No. 56, pp. 231-254, pis. 1-4.
and OGAWA K. (1958), Fusulinids fromthe Funafuseyama Limestone, Part I. (Onsome Interesting Parajusulitw from the
Funafuseyama Limestone). Jubilee P11b.
Comm. Prof. H. Fujimoto, 60th Birthday.

DEPIUT, }.,Rokuro MORIKAWA and Yoshinari TAKAOKA

38

Table 1.

Measurements of Parafusulina

I
I
I,
Height of Voiutions
Slide No.
L.
W.
F. R.' N. V. D.P.,
II!
12
314
5'6
7
- 1 1-1 1 I KZ 6-7- 113.5 3. 3-~-7- --~--~- -- - --~--~-- - - -1. I '. 25
08 . 15 . 15 . 20 . 20
25 I • 25
2 ~-1
2 !a-6.:.::16 13.sl 3.81 3.51_7_ ~35-~~~~~~~-:-25 i--.30'-I

I

SP., Pl. Fig

I

I

I

I

'

IKz

I •

1

3
4

1 -~

i1

6-=!4 12. 0 i 3. 6 I 3. 3 I

4 I KZ 6-37

I 3. 5 I

7

_· 5~

I •

I=-=: . I·
15 i--l5~~ j . 10

7

Table 2.

SP., Pl. Fig. Sljde No.

L.

W.

25 ____.__:~ . 35-. 14 ! . 16 I • 22 I . 30 I • 30
Measurements of Parafusulina

Height of Volutions
F. R. N. V. D.P.'~----.-~----~,_I___
2_j_3 ___<1__5 _6__ ,_i________,____
8_-~:___:_ 11 KZ 15-21~~-,j-s_I_L~_I_8_~1~~~--~ .28 :_4o~~~---1~~­_:_i_:__ __: KZ 15-18a[~ ~ 1. 91_8 _ ---~ ' . 10 . 20 I . 24 ~' . 26 ~;__- 261 . 26
1

1--~ KZ 15-16bl~ ~ - 2_-_ 1 I_ 6 '_2'1_1 . 1_2_ ~I .20 . ~~~ __-__:: - I _4 __:__ 4 KZ 15-22JJ~~ _ 3_-_~- ~ _8_ _._32 l_-___o_6 ~~- ~~ ~-2~! .28 ~ . 32 ~ __3 -2

_ 5 _2_6

2

72--~- KZ 15-16c___ ~7

KZ 15-23ai_ _8__

3. 6-S- KZ 15-i ~-- U

8-

pp. 49-57, pls. 1-2.
l!sAKA, C. (1932), On a new species of Schell·
wienia, Sclz. ozawai. from the Mt. Funafuse
district, Province Mino, Japan. ]om·. Geol.
Soc. Tokyo, t•ol. 39, pp. 1--1, pl. 4.
KA:--:~IERA, K. (1954), Fusulinids from theupper Permian Kuma formation, Southern
Kyushu. Japan-with special reference tothe Fusulinid Zone in the upper Permianof Japan. JV!em. Fac. Sci .. Kyuslm Univ.,
Ser. D, vol. 4. No. 1. pp. 1-38, pis. 1-6.
KoBAYASt-JL M. (1957), Paleontological studyof the Ibukiyama Limestone, Shiga Prefecture, Central Japan. Tokyo Unit•. Edu·
cation, Sci. Rep., vo/. 5, Sec. C, No. -18, pp.
247-311, pis. 1-10.
MoRIKAWA, R. (1955), Schwagcrininae in thevicinit}r of the Shomaru pass. Easternpart of Kwanto Mountainland, central
Japan. Sci. Rept., Saitama Unit•., Ser. B,

11~ ~]_._r_:_[ .20 ~ -~~!~~ ---~~~
J

.

34

. 06

. 141. 221 . 36

. 32l

-8-~-- 32 ~ ~ ~~~ ~~~ ~~-:--36
vol. II, No. 1, pp. 45-114, pis. 5-15.
- - (1956), Fusulinids from the Akasaka
Limestone (Part 1). Saitama Unit•. Sci.
Rep., t•ol. III, No. 1, pp. 93-130, pis. 21-26.
- - and HoRIGUCHI, M. (1956), Pan{fusulina
nakamigawai n. sp. from the Adoyaina
formation in the Neighbourhood of Kuzu
city, Tochigi-Prefecture. Saitama. Utziv.,
Sci. Rep., Sec. 13, 2, pp. 261-264, pl. 35.
OzAwA, Y. (1925), Paleontological and Strati·
graphical Studies of the Permo-Carboni·ferous Limestone of Nagato. Part II. Paleontology. Jour. Coil. Sci., Imp. Univ. To·
kyo, vol. 45, Art. 6, pp. 1-90, p:s. 1-14.
(1927), Stratigraphical Studies of theFusulina Limestone of Akasaka, Provinceof Mino. j(mr. Fac. Sci., Imp. Univ. Tokyo,
Sec. 2, vol. II, Part 3, pp. 121-162, pis.
33--!5.
ScHWAGER, (1883), Die Karbonische Foraml·

339.

Two new species of the Parafu.sulina. yabei type from T01mtro

39

tomuroensis MoRIKAWA and TAKAOKA n. sp.

Septal countThickness of Spirotheca

1

2

I

~o3~1

3

I 4 ! 5

.04.1

.o4

6

SP. __
1_1_2_ _ _3 ___4_~1_6_

7.o6 ~~oil~~~~~~~· .06 .061
~ 041~1~--:10,---:u;--.

---:os~ -:o4T~~o4i~o5 --:oG ---:os-

~

4

I 16

10

24

I

~ 31 I 36

26

L-Length
N. V.-Number of volution\\<'-Width
D. P.-Diamctcr of proloculus
F. R.-Form ratio *-half length or width

nabeyame11Sis Mor<IKAWA and TAKAOKA n. sp.

Thickness of Spirotheca

Septal count

4

5

16

_5_~·~_2o 1 31

34

37

SP.

1

6

1

13 ' 23
1

L-Length
W-WidthF. R.-Form
ratio

nifera aus China und Japan. RICHTHOI'ENS
Chiua, t•ol. 4. pp. 106~159, pis. 15~18.
TORIYA~IA, R. (1958). Geology of Akiyoshi.
Part III, Fusulinids of Akiyoshi. Mem.
Fac. Sci., Kyuslut Univ., Sec. D, Ceo/., vol.

2

11

3

29

1

34

17
N. V.-Number of volution
D. F.-Diameter of proloculus*-half length or width

7, Nn. 1, pp. 1-264, pis. l-48.
H. (1899). On Fusutina japmzica (SCH·
WAGER) from Tomuro, prov. Schimotsuke. ]our. Geol. Soc. Tokyo, vot. 6, No.
68, pp. 189-194, Text-figs. 1-4.

YAH!-:,

40

Rokuro MORIKAWA and Yoshinari TAKAOKA

Explanation of Plate 7
Figs. 1-7. Parafusuliua tomuroeusis MoRIKAWA and T AKAOKA n. sp.
1. Axial section of the holotype, x 10.
2, 3. Axial sections. xI 0
4. Sagittal section. X 10
Loc. Okuda Quarry, Nabeyama.

Explanation of Plate 8
Figs. 1-9. Parajusuli1UJ 11abeyamensis MoRIKAWA and TAKAOKA n. sp.
1. Axial section of the holotype. x 10
2-4. Axial sections. X 10
5. Axial section of young stage. X 10
6-8. Sagittal sections. x 10
9. Oblique section. X 10

M ORIKAWA and T AKAOI\A:

Parafusutin:.t

Plate 7

3

2

MOR.Tl\AWA and T A.Jt;AOI\A :Pamfusulina

2

Plate 8Trans.

-100.

Proc. Palacont. Soc. Japan. N. S .. No. -11. pp. 41-.W, Apr. 10. 19til

A l\E'\:1.' SPECU:S OF L\UOCENE GASTROPODA (l\IOLLUSCAJFROM THE GINZAN HOT-SPRING. YA:\IAGATA PREFECTURE,~ORTHEASTHONSHU. JAPAN*

KOTORA HATAl and TAL\110 KOTAKA
Jnscitute of Geologv and Paleontology, Tohoku University

lliC9M:aJlf!ffil.!Rilf.m~tt: ~Htli~f!l.r;tt.i!f:U)•rm*'i.t 'J @:-t.01!iHU) 1 llrfll!~::tC.iW\t... ~IV:'Etm£t(J1]:.f'l(:.:.-?t··nn:U):'!f~~i>..:t.c,~L
M ~1: 'J· J1f.. • 'J· t"li r~ ~In the previous paper (1959). the writers
described three new species of Gastropoda based upon the collection of Mr.
Tsunemasa SAITO of the Institute of Geology and Paleontology. Tohoku University
from near the Ginzan Hot-Spring in
Yamagata Prefecture. In that paper it
was stated that anolher new species was
also discriminated among his collection,but owing to the unfavorable preservation ·it was not described but merely
left in inanuscript form under the nameof Jlonodonta (?) saitoi I h T AI and KoTAKA C\IS), and this name was refered
to by T. SAITo (1960) in his descriptionof the geology of the area in which the
Ginzan Hot-Spring is situated.
Fortunately the junior writer during his
visit to the type locality of the gastropods described in the previous article(HATAI and KorAKA, 1959), collected additional specimens of the gastropod
species left in manuscript form. The
description of th.e species based upon
the additional material is given in thisarticle.

* Received Oct. 1. 1960; read at 76th
meeting of the Society at Matsue. Sept. 2·1.
1960.

Family Trochidae n'ORBIG:>;Y, 1837
Subfamily Margaritinae STot.tczKA. 1868
Genus Turcica A. AnAMs. 185-!
Turcica A. ADA~IS, 1854, Proc. Zool. Soc., London. pt. 22. p. 37. Type species (lVlonotypc) : Turcica monilifera A. ADA:O.IS. 1854.
Generic Diagnosis: Shell conoidal, thin. subdiaphanous, imperforate, whorls withtransverse series of granules, the lastrounded at the peripherr; columellasolid, spirally twisted at the upper part,
ending below or anteriorly in an obtuseprominent point; outer lip thin, simple,
acute. (A. An,\Ms. 1854. p. 37)

Turcica saitoi HATAt and
KoTAKA, n. sp.
Text-figs. 1-·5

Description: Shell solid and heavy,
spire broadl}' conical. Apex lacking,body \vhorl slightly convex and the
younger ones nearly flat in profile.
Whorls sculptured with five prominentspiral cords. subequal in strength andbreadth, abapical one usually covered
with succeeding \vho:·Js except on body·I •)

Kotora HAT.-11 and Tamio KoTAK.4

whorl. Spiral cords form rows of elongated beads. interspaces between eachspiral rather narrow and smooth, about
one third of spiral in breadth. deeply
excavated. Suture distinct. narrow anddeep. Base of body whorl convex andornamentated with three distinct andone obsolete spiral cords, also form rowsof elongated beads.
Aperture round and inclined posteriorly. outer lip thin and sharp, composedof lateral and basal lips. inner peripherysmooth in lateral lip. with prominentcrest in basal lip: parietal and columellar lips form inner lip and covered withthin callus, the former indistinct andflatly covers ba::;e of whorls, the latter
distinct and with two prominent teeth.
Columella straight with two folds ofwhich adapical one rather prominent

3

and abapical one very weak, On columellar lip or at terminals of columellar
folds. two prominent teeth preserved.
Dimensions (in mm.):Holotype (IGPS coli. cat. no. 77799:
apex lacking, three whorls preserved),
I Ieight approximately 40. width of bodywhorl 41. diameter of aperture approximately 30, pleural angle about 70. Paratypes (IG I 'S coli. cat. ·no 77800. not measured because of preservation not beinggood).
Type locality a11d horizon: Upstreamof the Okamami-zawa. Taniano area,
Obanazawa-machi, Kita-Murayama-gun,
Yamagata Prefecture. The Kaminohata
member of the Ginzan formation. Miocene.
Remarks: The specimens examinated
all lack their apex. thus the true shape:

4Text-fiigs. 1-4. Turcim saitoi HATAI and KoTAKA. n. sp. Front, top. lateral ;md basal Yiewsof the holotype specimen in natural size. IGPS coiL cat. no. 77799.
Text-fiig. 5. Turcica saitoi H,\TAI and KoTAKA, n. sp. Columellar folds of paratype specimen
natural size. IGPS coli. cat. no. 77800.

4 0 0.

A 1Vew Species of Miocene Gastropoda (Mollusca lof the younger whorls or spire remains
unknown. However. the spire seems tohave attained considerable height judging from the rate in increase of thesucceding whorls. The general featuresof the shell. especially the shape of thespire so far as preserved and the characters of the aperture agree well withthe generic diagnosis given by A. AoA:-.ts(185.!). According to \V. W E"z (1938. p.
271). the apertural characters of thegenus Turcica are:"~abel durch eine Schwiele bedeckt;!Vh.idung schief. rundlich: Aussenranddtinn. scharf. Basalrand bogig ausgeschnitten: Spindel verdickt, oben mit einer
Zahnfalte. darunter ein krattiger Zahn ".
Certain trochid shells resembling inshape of spire and surface sculptures tothe present form are distinguished from
Turcic-a in having distinct umbilicus
and rather flat side and base of thewhorls.
Some species of the genera !11onodonta
and Clanculus have whorls and spiralsculptures resembling the present new
species. but differ in their distinct
dentitions along the inner periphery of
the aperture. and the genus Clancufus
has a prominent umbilicus.
The genus Ca!liostoma is also more orless similar to the prc:sent new gastropcd in the type of spiral cording, but
has no columellar folds nor teeth on
the columellar lip of the aperture.
Turcica sai toi resembles T monilifem
A. AoA:-.Is. the type species of the genus
in the apertural characters as above-mentioned. but the former is distinguishedfrom the latter in having a rather low
spire. heavy shelL subequal five spiralheads. rather narrow interspaces between
each spiral. and by the less number ofbc:sal spirals of the whorls.
Turcica imperialis A. Ao ..-.~ts ( = Turcica
coreensis PEASE) figured by G. \V. TRYo:-.-(1889, p. 414. pl. 63, fig. 30) is also similar to the present new species. but may
be distinguished from it by the rather
thick shell. prominent cords and thestate of the callus on the parietal lip.
The gastropod fauna derived from the
Ginzan formation is interesting in comprising such species as. Apo!len sazanami
H:\TAI and KoTAK.\, J.Yucella freycincti
saitoi HATAt and KoTAKA. Ocenebra
aduuca protoaduuca H.\TA! and KoTAI(A.
and Turcica saitoi HATA! and KoTAt'.A.
besides Olive!la itcakiensis No:o.JL'RA andHATAI. Ca!liostoma sp.. Lemiutina sp.,and Neverita cf. didyma. This assemblage is of particular interest because thelisted gastropods are those which arecommonly found in the inner neritic
zone crawling over rocky places on
which sea-weeds are growing (0/irella.
Calliostoma), on rocky shores close to
the tidal zone !Ocenebrai. attached to
rocky bottoms. rocky shores aba\'e low
tide level or on fore:ign materiab (Lemintiual. sandy bottoms ( Neveritai. tofairly deep water (Apu!!on. Turcinn.
Since none cf the specimens show abrasion of their shells it may be inferredthat they were quickly covered withsediments to become \Vel! preserved andnot transported from a distance. All of
the genera can be found in beach collections along the seas of southwest Japan,
which suggests that they can inhabit
shallow waters \'ery near to the strand
line. This may also be supported fromthe occurrence of pelecypods belonging
to such genera as Anadam. Volsella.
Ostrea. Callista. Pilar. Dosinia. Spisula.
Macoma and Sole11 as well as of Mya.
All of these pelecypod genera are common in shallow waters and can often hefound just below the low tide level oreven above it (Ostrea. Volsella).
From the giyen evidence it ts thoughtthat the shore line must have been Yery

44

Kotora !L-!TAI and Tamio KoTAKA

close to the place where they wereburied and that the depth estimatedfrom the bathymetrical ranges of thegenera mentioned above in the seas ofJapan can be, at the minimum. placednear the strand line. and at the maximum
at depths shallo\\·er than 20 meters.
Should further data accumulate. it may
be possible to judge the position of theancient shore line.
References Cited
AoA~Js.

A. (1854). Further Contributions
towards the Natural History of the Tro·chidae. with the fksc:ription of a New
Genus and of Several Species. from theCumingian C:Jllection. Proc. Zoo!. Soc.,
London. pt. 2~. pp. 37-H. moll. pl. 27.
- - (1863). Description of a xew Genus andof Twelve New Species of .Mollusca. Proc.
Zoot. S'oc .. Lmulrm. 1863, pp. 506-508.

liATAJ. K.

and KoTAKA, T. (1959), Some :\'ew
Miocene Gastropods from ncar the Ginzan
Hot-Spring. Yamagata Prefecture. Saito
Ho·on 1\ai Mus .. Res. Bull.. No. 28. pp. 611. figs. 1-6.
PEASE, \V. H. (1860). Dzsc·riptions of Thrc..:
Species of Marine Shells from the Pacific
Ocean. Proc. Zoo/. Soc., London, pt. 28,
pp. 189-196, moll. pl. 51.
S.\JTO. T. (1960). Tertian· Stratigraphy of
the Ou Backbone Range bordering Yamagata and Miyagi Prefectures-Pt. l. The
Western Border (in japanese). four. Geol.
Soc. fapau. val. 66, no. 774. pp. 157-169,
4 text-figs .. 3 tables.
Trno:-.:, G. W. (1889). l'vlanual of Conchology,
vol. 11. Trochidae, Stomatiidae. Pleurotomariiclac. Haliotidae, pp. 1-519. 67 pis.
WE:-.:z. \V. (1938). Gastropoda. Teil 2. Handbuch der PaHior.oologie. Bel. 6. Gebriider
Bortraeger, Berlin. pp. 241-482, figs. 47212H.

Trans. l'roc. Palaeont. Soc. Japan. \:. S., Xo. 41. pp. 45-4~. Apr. lU, 1'-'til

ADDHESS TO THE TWENTY-FIFTH ANNIVERSARY OF
THE P~-\l..AEONTOLOGICAL SOCIETY OF JAPANTokyo. Japan (:\·Ieiji ZG. 1893l were successively founded more than half acentury ago.t> These are senior societies.
but there are some juniors. the Societyof ~lining Geologists of Japan (1950 andthe :\lineralogical Society of japan (1952).
for example.
The oldest among the societies of thisscience is the Palaeonlographical Society
which was instituted in 18.ti for publishing monographs of British fossils. Since
then the societies were founded in
Switzerland (1874). the United States(1909), Germany (l\H4), Russia (1917.1.
Austria (1933) and so forth.") llowever.
even in the United Kingdom it was in1957 that the Paleontological Association
which holds meetings and demonstrations
beside the publication of its journal has
The Palaeontologicalbeen founded.
Society of China was founded in 1929.
but its activity has not been commenced
before 1948. 3 ' Lately more companion
societies were founded in Argentina ll955).
India (1956) and Brazil (1958'. In looking
through them this society may be said
middle among the ages of foundation.
Retrospecting these 25 years. the history of this society is not simple. In
late spring. 1935. the general assembly

It is our great pleasure to have had
the twenty-fifth anniversary of thissociety last spring. Its celebration was.
however. postponed for seYeral months
for preparation. On behalf of the society
I wish to express at the outset. themost cordial thanks to the l\linistry of
Education. Geological Society of Japanand other related societies and mining
and industrial companies as well.as sustaining persons for their warm assistancesand cooperation which we have received
since the foundation of this society.
'Jl1e llistmy of Pala('ontolugy of Japan
will be detailed in the writing no\v inpreparation by the Honorary President
II. YAnE with a few joint authors. At
any rate the history of research does
not go as far back as a century, putting
aside the study of medicine stones andthe knowledge of stone curiosity. During the Showa epoch (1926- J the numberof palaeontologists was gradually increased as much as a society can be established. Stimulated by the constitution of
the International Palaeontological Union.
1933. this society was established at
length on the 29th of June. 1935.
Among the societies of natural history
in Japan the oldest is the IlakubutsuTomo-Non Kai (Meiji 11, 1878). It was
the progenitor of the botanical and zoological societies (1882. 1885). in addition
to the geological society or Chigaku-Kai(1882-1892) which was. however. laterannexed with the Tokyo Geographic
Society (lRScl). Not only these but also the
Anthoropological Society of Tokyo. Xippon (1884) and the Geological Society of1) Geological Society of japan (1953).
History of the Geological Society of Japan,
Commemoration of its Sixtieth Anniversary.
2) ScHIXDEWOLF, 0. H. (1956). A Note on
Palaeontological Societies. jour. l'al. Soc.
l11dia, Vol. 1. j\io. 1.
3) CHAO. C. K. (19·18). l'oundatiotl andHistory of the Palaeontological Society of
China. Bull. Pal. Soc Chiua .•No. 1.

45

Teiichi KoBA.YASHI
was held at the University of Tokyo
for its foundation as a section of theGeological Society of Japan: the firstmeeting held at the Geological Instituteof the university at the end of :\ovember.
The papers were published in the sectionof the society in the journal of the Geelogical Society. Their reprints werecompiled quarterly for distribution toits members. Thus the Transactions
and Proceedings of the PalaeontologicalSociety of Japan has been published 21
numbers by 1941. This dual publieation.
however. became impossible during theSecond Great War. Consequently, Trans·
actions :\umbers 123-191 were printed
only in the journal.
This society has held in each year
four meetings at various towns. but thisactivity ceased with the 38th meetingat Fukuoka, February, 1945. The meet·ings were. however, resumed from June.
1948. Since 1954 one general assembly
and three local meetings were held ineach year. Thus 77 meetings and 6
assemblies are countable by the end oflast year.
According to Dr. YAHE's suggestion
this society followed the Paleontological::)ociety in its relation to the GeologicalSociety of America. The PaleontologicalSociety, however. became independent
from the Geological Society and since
1926 it began to publish ]ounwl ofPaleontology jointly with the Associa·
tion of Economic i\lineralogists andPaleontologists. Similarly this society
became independent from the GeologicalSociety of japan in 1957. This parallel·
ism is noteworthy because it shows thegeneral tendency for the differentiationof sciences through their development.
New Series of Transactions and Pro·
ceedings and Special Papers were firstpublished in 1951. They are mostly
written in English.
The former isquarterly. while the latter is occasionaland 6 numbers have been already published. In adclition, the initial numberof Fossil or Ktwseki which is in Japanese, has appeared last year. It comprises the symposium on the fusulinids.
Beside them. Catalogue of Type Specimensof Fossils in Japan will soon come qut
as a Twenty-Fifth Anniversary Publi·
cation. The other new activities are
special lectures and symposia at the local
meetings and general assemblies.
~eedless to say, palaeontology is in·separably related to neontology on one
side and to stratigraphy on the ether.
Therefore this society comprises geologists as well as biologists as its mem·bers. It is not a big society but hears
some international aspects. One of themis its foreign members which attain
more than 10 percents of the total. Is
it accidental that this percentage is ap·
proximately the same with that of the
Paleontological Society occupied by themembers resident outside North Amer·
ica? The Paleontological Society combined with A. E. lVI. P. is, however, con·siclerably greater. having several times
more members than this society.
Recent Volumes of loumal of Paleon·
tology each consists of about 1150 pages,
while 40 numbers of New Series in past
10 years total about 1450 pages. This
relation roughly corresponds to theannual membership due which is about
ten times greater for the former thanthe latter}> Likewise. the Penrose be·quest" for the former may be comparedto the publication subsidy to the latterfrom the goverment.
As shown by the history of thesocieties. palaeontography must precede4) This means the domestic membershipdue which excludes the cost of Special Paper,
whereas the foreign membership due is inclusive of the cost.

Address to the Tzoenty-fiflh Anniversary of the Society
palaeontology. because the latter does
not exist without the former. Therefore
101 new species. 12 ne\Y genera and 5
ne\v families and 293 new species, 28
new genera and 2 new families established respectively through the old andnew series of Transactions and Proceedings must be highly evaluated. If thenon-serial part of the publication in 19411950 is brought into account. the total
number of new species, subspecies.
genera and subgenera described in thisjournal attain respectively 420. 42. c13
and 5. It means that about one-tenthof the total new species and subspecies
so far described by Japanese palaeontologists made their debuts in this journal.
~aturally various groups of fossils in
the fields of palaeobotany. vertebrate andinvertebrate palaeontology and micropalaeontology are dealt with in 393 transactions. Many of these works are notmerely descriptive. but various problems
are discussed from different points ofviews. namely, morphology. anatomy,
taxonomy. terminology, phylogeny. ontogeny. ecology. ichnology. speleology, biostratigraphy, palaeogeography. palaeoclimatology. biometry. fossilization. applied

17

palaeontology and so forth. One ofthem is dealt with ancient documentsof the so-called dragon's skeleton. Most
fossils were collected from Japan oreastern Asia. but some papE:rs concern
various fossils from the Philippines.
Viet-Nam. Thailand. Timor. Java. Nicebar. the Americas and other continents.
I appreciate a few papers which haYe
been contributed by foreign members.
Here I have tried to orientate thesociety in the spheres of palaeontology
and also of nat ural history of Japan andto summarize the past activities of thissociety. because the history and thepresent status are two important ba;:,isfor the future planning. Through themost difficult time the society has
made slow but steady advancements.
It is still only 25 years old. and its
future improvement depends principallyon the efforts and cooperation of its
members.
Finally. as an executive councillor
since the day of its foundation, I amvery happy to have had the honour topresent this address on this memorial
occasion and thank you all.
T. KoBAYAsHI, President

PROCEEDINGS OF THE PALAEONTOLOGICAL SOCIETY
OF JAPANs *!!11:.~Cf:~~ 17 @lf71)~:~ 1960 1F 11 JJ 19
a. iJ~~Jt!r.:.f:~c~JW.-c.>ill!!-~#t:;:.:to~ ·"COOt!¥.
;Htt.:. ($hll1i 26 iJL f§JJ~K.:tolt~~ii:it-"J·~r.v:\.:.~WI~ II fiJUD)!li I)"(~ :0 ~. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . {£EE.t::iffWf~

Metadolioliua 1.::.-:>~ ·"C ............... .

. . . . . . . . . . . . . . . . . . . . . . lit!'~- . IJ!f l:Jit~

5]1)[I *<J)~t'tl!.t)..J~ii.::.-:>~, "(Projusulina t7) wall structure 1.::. -:>~ •"(( 7. 71' F tl:#l)...... .. .. .... .. ...... .. . i'&i#:S!-=-A Faraminiferal sequence of the Yurakucho
Formation (Holocene) in Tokyo City ..
Try to represent the Foraminiferal Zon·

48
ation stalistically .......... !Iiroshi UJIJ£
B1·achiopoda of the upper Permian Gujo
Formation. Kyoto Pr-efecture. Southwest
Japan ........................ ii!JJF::ktff$
On the Occurrence of Ordovician Nautiloids
in North Thailand .... Teiichi KoBAYASHI
Ordovician nautiloid distributio~ of Nor.th
China .......................... l}ifl.t.*A±r6. Sepia (l)fl:fii.:.-::>t ·"C (t'C~) ....... .
........................ j\!!j;.t.: §: . 'i'f!f=.f!l>:!tifil.IHJ: IJ Oxytropidoceras (l)~j! ((i::~) ....

....................... ' !iii* ][ . mra,IY}~

t~re=:fi!~~iQ

c &iJikf.'t(l) JU'lHtH::. -?l •-c _ .. :fi

i!l.f/J~llf" (l)if1:~~1¥J!Vf~- ((i::~) ....

................................ l.K!J.!fmrr~me*~••~Q)~m=enmw(/)•M<R~) .............................. ;t-[l!J~fi

11 *tli~£.¥Q!'f:~f!B:fU 35 ipfft'<f~l1 1961 {f 1 fJ
15 B 1!DIC:k~:JlW'7:ffl)J1!!71!:-'f:fxii[I::.~J.. ·-c!mll!! ~:ht~ (~~:-{.·57 1!), 1f~;0;.:.Mt.Q~M:ii!t1'\·)IY-0:i.:.JI>jitillliW :1(XQ)}jfi I) -c;~;;,

0}]I]~ 5 lffill'IIJf.iill~'.¥-a"f:.i!ll{t 111.0:m 21 juJ7i(;§l.111!'t!I"f:~,ljlli>:lfii~L. -c......... ' ..... tJ-1i:J.li.-

Umbilicated Pseudoschwagerinu from Japan. . . . . . . . . . . . . . . . . . . . . . . . . . Ilisayashi !GOStratigraphical occurrence of the genus.tlcerroschwagerina ........ Ilisayashi !GoOn the variation of Parafusulina yabei HA:-o:ZA ,,. A . . . . . . . . . . . . . . . . . . . . Hisayashi !GO
On the specific variation of Neoschwageriuamargarita DEPRAT and Yabeina globosa
YAilE .................. Kimiyoshi s_. . o_...
Uralian Trilicites from the Atetsu lime·stone. Okayama Prefecture ........... .
. . . . . . . . . . . . . . . . . . . . . . . . Kimiyoshi SAD>\{ci':l-lis'JtiJ,;B~Ifk Cancel/ina 1.:.-?J..•<:: .... ~JIIhfif;
IL1Z.Jii-1Ji1,,:':j~li di .!lf1i (I)~~!U;.:. "'J~ • -c

· ·· ·· · · · · ·..... · ................. ~JIIhRf.; · =~*-

Planktonic Foraminifera from the Nobori
Formation . . .
. .. Yokichi
TAKA YANAG I and Tsunemasa SA ITO
Re~·ision of the Terebridae from JapaneseTertiary ................ 1\atsura 0YA~IA

ljJJi\pHlifllfei\:(1) Vc11cricardia Ja.i(I)~Hat~f:flti!Vf~­.. ~r-"JifliJ~lif .. <•!;·~.=*C) Q):$}ltl~I¥Jlilf~-t (/) 1 ( f~f.i'~) . ......... A'!llfJ.iHr · 11'-l:iEJIH
Jjl)f;I*JJ\~Bf.llff(}) Crassatellitidae :l<; J: V: Tellini·dae Q)jffiJii::.-?t. · -c -" ijHJill/J~!If" (I)~J'li~(l(_jlilf~ -l-(1):! (fi::V\:) ................ ..

.... · ............... · WfiEIIS · l.Kff~fi
On some Miocene Mollusks from the l\-fizunami Group, Japan ............ *ff.tlll~=
Scutellidae (I) lunule 't slit Q):f,li .... ~r a"'t
17900'¥;f!J~@:~~*Cilltllllft'E Hnr.iaster. -~r db
A new Genus of the Phillipsidae from Thailand ................ Teiichi KOBAYASHI
Equid fossils from Hiramaki Formation _.
.... Shinji YOSIIIDA and Tokio SHIKA~IA
On the Neomeris (A :t- ~ !1) from Onsen-gun.
Ehimc Prefecture .............. ft.f:i.Jfl~fll

mt~:m(/)? ' =- 7-fU1 t !\11&t (/) l~ff: 1.::. :v.t ~ --c............... " ........ " .. .. . :~kiJU1{~
Ecological and Geographical Distributionof Jurassic Pelecypods in Japan ..... .
. . . . . . . . . . . . . . . . . . . . . . . . . . Itaru 1-I.... YA~Il
Note on Heterolrigouiu .'Ubm·tr!is (J 1~lBO) ..
................... Mitsuo NAKA!'>O
On some Gyliakian Pterotrigouia from Kyushu and Hoklwido ...... rvlitsuo K·\1\Al'O
On some consideration of the molluscan
fauna from the Asagai formation, Fuku·
shima Prefecture ........ Saburo KA!':'!':oOn the Asagaian Cardiidae (Pelecypoda)
and its geological distribution in the
Joban Coal-field ...... Yasuhiko KA:>IATA
Pala:oosynecology of the Younger Tertiary
Deposits in the 1\Iabuchi River Yalley.
:Xorthcast Japan ...... Kiyotaka Clli:O.ZET
Ammonites Oxfordiennes de l'ile de Mindoro. Philippines .......... T~·.dashi SATO::t A!- 7 "-~·(I) Dcsm~d~nt c Lophodcnt

OJt!.!lff: ........... ' .............. :(f::Jf:~.P.F>
Lias J: IJ Ichthyosaurus ji_({1lf(l) ~ji! ..
....................... '~~ jf:~= . {Iiili lE;-;!.;~JII

Fossil Felis from the Maru-hcn Quarry in
the northeastern part of the ~likata­bara fan deposits .. T. SHIKAMA, Y. HAsEGA\L\ and Tokai-Gakuen Chigaku Club
Eugollcphyl!um. a new Permo-Pennsylvanianalgal genus ......................... .
...... Kenji 1\oi'\JSIII and John L. \\'RAY4!)Equisetiles Kitamurae KoN'NO, n. 5p. fromthe lower division of the TaishU Groupin Tsushima, japan ........ Enzo Ko:-.·:·w
Two types of Equisetacem1 fructifications
from C.unic formation in Yamaguchi
Prcfcctuae, Japan .......... Enzo Kol'O'No

A Miocene Problematica from Wakayama
Prefecture .... K. HAT AI and T. KoTAKA
On the Cenozoic floras of Japanese Islands.............................. Seido E:-.noill): Jit9,URII!Jff£ Dcsmostyl id IJ)1Ht/tll)l!ll!Q::[)~i~J:.

/f>:i:'ti.W.H'n 35 {f- 6 f.l ~9 fla-t~ "'Cf{IJ.lz: 25 I.'J
1H:· .i!l!X. f.; IJ)"(.:: it a- ~c :&': L.. "( F;itl()) .::'.!: <oiLL~~·ii"tJ;, -tiJ)J.\;~~t.d·. 1.\i~Wf:V?ft.iitJ'.J;,?

;JC%~~rr~f.:"
1Ri'i0 · ;'l!.l,{!:,j~jjl{~ · i!!C~~Iii!Hftt361F I H
1-1 PPIII"f:±~~-c-rmfllt ~.tti,;1Juit3'-~:t)
a- ;t "(~*"'(';ji;,? f.;;, ~())11\~lti:k()).ilf!IJ '"C';li;.

a(n~, tJH'i~ IJIJ~IJ)ff · :ftJ~~H~ · *'i'if5l;T;•J ( r~*Jm1'1+~ ~~t-=r-IJJY~N.: .t I) ·F,1.31l) .t :> f.cm,,;.J tJ:;li;. ~f.:) · tJI. iliV.k. 0: t51. ·~ Wi ~li (International
Palaeontological Union ()) C. PoULSEN 0*=.J: I) IJ):t.;tJI.~ ·()) 'f~~u. t:.6b t'i\GIJ)3-ffil+~ . fJF'll.:m- . MAt.c c." .J: IJ NJ.il . ·~ 'i[tJ;;li;. ~f.;;) • ~"f:~1tr Jl',t.-JII'H.k (~$:lii:%f,-'j-~_HI:.+i~:fJ:IHJ--::>~.

-c .......................... llt£r..,u~·~"(f*J~iiJ)J~j£tJ;Ji> I)' -tJt.tJ;ijl~ ·"'Ci-.J'.Iki~'Eii51J)(nJl:l.:.t'::.b't:--:> ;t, ()) c 5~· x.. tAd -t

0

i!Vi. J'< k rrr~~~.

T};.H~':I!.fW-t!:t" » :/7- I·IJ)rrlJ-c;, .lt!ft"i.-:(;IJ),i~f\::11:1.:.

--::>~·"(, JH!IH..:~~-Itl?h"t:too~Ht. *~~ifl'i~~-K~~5}K~~~IJ)::c!::"C~IJii",
H~'{:IJ))Cji;;t~!lR'"C'J;,~.::~.~ ·a-fH.. ft!!IJ)

cK..

H:;.f:WIJ)f~iic!::.!:: t1..:. i"'~M\'LIJ)i~l ldH!!H:.l£<.
*''f:.i';())JCIJ~a-:toM 1J L... t.lt7'1iill),wc~·t.:L..i-t.

IIWfll 36 -¥ 1 H 14 aMIJ).J:5~,.,¥~~M~~.tt~~·+&~~&~-'J. (:(E}I'iJillf} · ~tt:;.t.l'!li · ,h:·~~~ · 9Jfi/X11

IJ)~a~~~-~*~•~~.tt~)·••~~~'.}<{{¢ (tflllllti\!JUtl:. ~$.5:!JU.k.V:Si2-f.ttJ::t.; I?
:td.:) · flfl~IJ)~t, trl~~.iPfM:0 r'c1~f.'d 7 :/ ·L: -r
1 I> IJ)jifl:l..:"?~ ·"'C <~*~l~IH) · i'JlVd'HHl<!RII::U c fi<llt~Xflt Ux!ff rtf:'f.t) · 5tf!i .!:jilt (Jiii
JII:::C~:RL

l!t~~(;I} !J 7*,._.=-7*~'1:())

R. w. CIIANI·:y f'IJ±. 1 !) / 1 ffl:10t!!N.111HtmIJ) M. L. THOMPSON j•!J±:k:MIJ)fJ!!f';~i~~Ul'·'~ L...lll'i~-c: ;ji;,? f.;;)0*

El 151:.~ ~-·(:01J)JIIJ.ll: 25 J,',j ;~'-;lrl.ft ~II:. "1 f.: I)'*~IJ)~~H.ltlfi1~ ·f~ L.. i-t" "l"~ftJEIJ) l¥li~ 11.
ffl'cilll\%!t~iR.!:: ici'b.tt.

:: IJ)ff!tf l:Ji-11. ttJI[ijiJ)~
i.i!i.:. .t ~.::.!:: tJ:, ~ <. rJ;. <f!I:IHJI..:ll,g~-!t I?~;;, .t:) 1..: t.c 1J i L.. L ~ L.. 'F I?. t~d~i V? }£11,$~ ~ t.c-t t ())
li.. }}5~H?f:'"C~ 1Ji-t" f*{,j, tHfiict!J-a:.tt"ti'i* This commemorates also the centenaryof the birth of Prof. M. YoKOYAMA, father
of Japanese Palaeontology.

The rapid development of Palaeontologicalresearch in Japan and the cxccllcnl achieve·
ments of Japanese Palaeontolo~-:isls arc rc·
cognized in all countries as contributions ofworld-wide importance.
It is lherefore apleasure to participate in the celebration of
the 25th anniversary of the PALAEONTO·
LOGICAL SOCIETY OF JAPAN. On beh:llf
of the INTERNATIONAL PALAEONTO·
LOGICAL UNION I welcome this opportunity
to send our congratulations and expressionof our most cordial wishes for the growthand prosperity of your Society. I am con·lidcnt that you will contribute much during
the years to come to the advancement ofpalaeontology and related sciences. and thatJapanese palaeontological res~rch will con·tinue to be of great importance not only to
Japan but to the entire scientilic world.
Nov. 23, 1960
Christian PouLSE:O:, PresidentInternational Palaeontological Union

50;: Q)ft!! The Palaeontological Society Q) N.D.
NEWELl. ;f';J~.

The Palaeontolos:ical Society ofIndia (J) M. R. SAHNI ~!~. Tubingen -).::'f.Q)
0. H. SCBlNilEWOLF fl~. -f?j'l'j:;k"f:Q).I.!HPIH&~- Societe Geologique de France (J) A. F. DE
LAPPARENT 0R. The Palaeontological Institute of the Academy of Sciences of USSR(})7 71 :r, -~n Y. A. ORLov,
SocieteGi~ulogiquc de Pologne Q) H. SwmztNSKI 0}~. The Palaeontological Association Q) U. M.

w±.

B. BULMAN ~ R· ~tj~:;k~Q)I*'Bj(j;li~t.cm;:,;r · ~·ldJ~<l!>-:>t.:.

c: .t I)~~~~~.tiJ~~~h~~~~-ftfiftK.t

1J rcatalogue of Type Specimens of Fossils
in JapanJ n:Ulllfi~.ht-:. 5Efllli 1800 P], ;tl;Q)ft!!i\c~111~ L. "( 1800 11'(\;l;::tf:lJrei ~h~i'J:Jll! (Gt>Y·

c

I.ER,

NAU~lA:-.IK,

MoJstsov tcs

NATIIORST,SCHWAGER,

t.c l:::) [gljJiQ)ljj'Mt.c E t =r-~~ L. "C~,,;;,.

I'I•l::.•~~t'i:J!.:J..i!$t\'Q) rw;t::&V':It·,- ,~. 1J 71f'l'lfife~:ric1)filf'Jl:J :-::k·lL+J.IIll!iUIIll.t 'HlliiJitJ:tz~~;ht.:.
1'; ft l\1. L. Thomp:;')n :!lH:.i~~'- I JJ tl •ii] 7 ,~.- 7' 'J 1 ~ 3CW!f,{~1~*~*- "CM;j~ L. L

il:M.:k~!f'f'FAnJL!llt~~~!,~ .t '> f,{f11:ft'1Hd~-Y- ~;Itt-:.
~ 4+J9~i'Ei 1-t"J' r • JillJt!! 1i (J)Jt!!1~'¥:ft'~Wf1e"JnliJIY.c I?V: i.::.Jl!!'Cf:·l'Y .U.l:G')i~~M•-=- f-:1..,. L "C• I•J.t;l:lfiUII~.t 'J

w.

o:J•Ehl)( ft·N:v:~~ ~nt-:.

News
1~;1i!(J)lXll.::.~fltJta 3 ~t/·f!l:lJ!.~h£H~r~r'?E1'mc1) ·! ·t4-h:.
<-.h.t.: t ~a- J<~-r 7.>.: e: n:tf> 7.>n:. ;:;h.lt:;f;:.ttl.:.l~f~JUJ<a- L t-: c. c G'),:b'!> ,V;;.::~N 1>f.:l ,c_ ~.O:JII.il: ~~ ,Cf)"( E L. E L. *ittl.:.!aJill!:f~r~ Mli~mt-ti.'L'ii"om! :11!!

IJIJ
~ 78

:r; 79 l"l ~~ ~~j§tJ ~

I :h

iH.

** I

:IIi *-'+

ii/1\' &l El• ;i6 l'(! YJ 1-1

II

{lli

1961 if 5 J1 13, 1-1 D

tk Ill "Af:~~ ~tt1 80 l"l

I
o

'"'

IJtl

1961 iF 911 23,24

1961 ::1:. •1 .H 30 IIn1001 1f. 8

I 1961 if 11 }] 18 13

n 31 n1961 if. 10 )1 25 II1961 ::r 5 JJ 13 fl <J:) l.::.t;;m-c:tlllllli ~nZ>:;$:i';~ 78 @!~~~.:.~ t,. ii!Jflli!11:.~~-~~.:.M-t oJ.t.iQJ0tJ:~tlii•1Mvn·o. ~tli~U\':flti~!af~;r,. . -:1,1~ 14 B (13) 1.::. •· 13*QJrfr.$Jil!tM~ff" 1.::.1~-totr.J',il~~~~hljrjt; -{:QJ:/P ?'7 .t..l1~i';QJ7'P ?"7 .l. cftl.l~i.:.:t<;?:n ~-l!'L i-t. lli:if.S~ iU!f/llJ-,J · iJl1l
o~FtBm.

o 1961 1f. 9 n 23 n <±) '..:.~!R-c:r.nrm~:no*~fR 79 (il]~fti.::.ttia<drlt'i~'f: <all· 1i1Kaic:tt)
i.:.M-toiM.MS::~;.j·pl~•l•-c:-t.

lli:.lli~

rllJII ilL~JliJj,; J: V:lli~-~IUEZl'~

1961 {J'. 1 Jl 15 II
7 l.JltJ:!fJ:~ ~tt.f.:.

.!l£5J{*.c'J:-c:I»Jh·.ttt.:.l3 *~!!1:.~~~fM~-c:(kQJta:J <~ffiJiJI1 3 :!f;:.QV:llill!(!IJ.5E 2. 3,

L 0JIIJ;n 3 1(:QJ!n 4 lJl c t, 'L('XQJ.!JHl ~7t!IJ11i" o.
4. f~f:a!QJflJ:JYJ • ~WJ:t,;J:V:IilfJl:X~Htf.!:~V:r.:.~f;hi.:.~.J'i"Z>!/J:lJ'QJJ.(~.:Cd)(l!\:;(~ 2 ;1~r1.)11(1'Jilil&:i.::.
ift-to.:. c.
2. ¥..tfMl;iE 2, 3, 7 .IJH:-lXQJtJ'H.:.~~i" o,
2.

3.

7.

/Ji{~~~:t.~:)(I.:.~'J-UJ;; 1:171;; -JH~

(22.5x27.5cm) 1.::.-:1 1J'f:H;i..:.-c 1 (f:t;~!..:.19JI[C(;.::.;,r
1:171l·L, f:1Ji11!J V -;;t'l.:.~?il'l.J?m)E-tZ>. [7-:;,.~-'J1~Lt..:i':I:.7}11Y{:ioHd•fiJ...

i"o.J/Jl(ft:i <f•Pii<l· Jl!!~~l • HJ<~~tr) 11 12 FmliiJU (tt 1771 ;,r -Jil~t.J 30 tlc) [JJl:.i:!UI!IJii:1
8 l'll.liliJJ.'( (!11771 ;,r -Jfl~f.J 18 &)J ~ml!te:-to.
tJ,J:QJjil{)f£~m;:t 7.,.ij,jftlt *:fitJ'·..f:"QJ f't}fl~ 1\lli-t Z,{ll LI~~IC "'=>~ '-cl:t. mlf',J? 01/· :1;!.;1) ;._:,tJ,j1'\-l;t. 'li~JlGIJ1'(QJiJ.b'JI.:. 1 ~lflfi~bu;:t7.>C.cn:-c:~o. [7;.;f.l'-71-~-L-t~8j;})'QJ~{>I•J~J.f•

A. -to ,J1960 if. 1 Jj 17 ll.t IJ 1961 iF ,t fl 3 A~-c:r1.)~Jll'$lll!Jiti'XV).iffiiJ-c:.i;o.
A.§;~

IJ-'Ifh1J11H·

MD: !,Jt:k
i!!i!.::li:N:WII~rr

IJ•~IJ;(,Il::::r1*1!1 ¥.o q: 1'<~;1'lr r.tl

tit*llfUlt;~:5-.:t~~«:iii~lfJ*~:~-fff:t'~Jii¥~Ff IIJ1
jfj Ullin:.mJI' ~WO~;­

Jt:L:!tt=(!.:ltfiirlo~

r:JJ

SIJ

50

J.tt~:k'!!f..Jm'!!f.tm.lt!!t'!-:_f.f!{'jip1fi

13*1!11:.~'+~

til {~' · K:C*m ~ :tt~

f~IIJ',ji~~j:JJl.lE=

fi

~

1f

il'
Jll

m

~-

rut

~~~

(.l~

Uf41-l}2 5o I'J

ff- Ralph W. Chaneyilmm..

Winn mx
MHii.lEFti
H. G. Schcnck(9tL:)~C!J.lE:X(9tL:)

1 9 6 1 if. 4 JJ 1 11
1 9 6 1 if. 4 )j 10 IIli*

W.iilimrm IJIJ~

J!f[!J f.'i'rrj

r=u Jill:tt*

ti n 1'¥. :it! ji•: 8 ·17 8 0 fft)
Jlf /ill ~~ g z: Jt t"J HiJ 2 / 1:1~-~illma~~~~

~

w

~

(1961, 1. 15)~ 1 ~*~r±E*ti~f!d!'F~ct'·L

1n 2 ~*~!irti~~A~J!~"J: 0: ULI<:OO!x;<b ~ :iff~j~.flO)Jlft!!i::toJ: O:t!ilk ~~t~O)~ bl ff~*~liff,2$0)fJifH·.i!i!'"t7.>t.:~~qyJi~~B?oW, 3 1:

L ~~H O)fi:f.~>O):J\~il?aO)~fJ'"

c Too

2. ~Wil.i\Jf~i00)0llltllo;:f,6~

3. ';YfJ:.ZO)t_:~O)f*!f!~ • ;i,IJilii~-fO)fftJ>O)Ul1{1!L,l. liJFJ~c:Dl!l:!IJJ · ~11lb:t.> .t t)(/~l·:l\:X~8ltJ: t;,ry;.::.0fj)t-=:.X-t-to~J'Jic:D~tPcO)fiJ!tn 2 ~c:D Bii'Jili!lill:
rcfi-to:::c,::<f.=~O)JJ(f~~&-r 7.> t.:~J~~O);t~H'f'l;;$:~!<:~fillO)IiJf~~a~~ll!l <c:::. ciJ;-c ~ 7.>o;,}:01iri~ffd~j':HJ: 0: L. ft!<:l.\(1~£b ~ ;t/f"f:FH<:~iij:H!f-?~ffJ.-c*-tl~T oo~~~]}WC ,u;-Jm~.W. • t~1JIJ~fl· *IYJ~f.l.t;;J:Uiltl~J~cT 7.>o;:f,7$;:f,S$

t!flJiliZHfi;;$:~1<: 10 :!'j:fJ. ti~JJ.I:'<b ~ ~!:1::~~1<:-?t 'T~~O)ib 7.> t 0)1:'. ~5JU~J'J. 5 :gO)fffi

ffll*
m5~

tt-.illi~dJ.IiJfr:JEO)A.~rt'.i6:;tl=HltlW vt_:~·f<:-? ~ ll'l':lnif:l.~O);lDii<:J:-:> ··ot:~ 7.> o

J.l'!'O)ib-:> t.: t O)IZ:.-? ~ ;N1 .llit.l.~O)~IZ:.J:-:> T~~ 7.> o~t!!b~Alitll 2 $0) t::HfH:· tttUb'"t oW. A. -c;w:~&fl.~O)tffnmrz:.J: oo~!!H~v.l.fib~l3.7~.f:lt:.-?t 'Tll1i~=t,(!1J*IJ.O);h 7.>#1r..-? ~;lfJD!J;;!~tJqf{HiiJLt, ~~ 0) ~~ ll..J:-:>"lJE~) Oo

m121:~.flt.itjj 12 $1:~~ .; :tt t.: ~Vl HlJ~ t.;. t}~lli t.;. Gt.H 'o ~~~i~;ltO)Nd/p ~~f:tili 3 *:ll..mJt v t.: ~H~IZ:.~:!Ju-r 1.. L. c iJ>-c ~ 'So
01~0)1!Z'/mlimi~li::.1h Tit~;:,, ~·~::J:ttiffif<R:If 600 f1J, ti'i.llU:tR~ 1. ooo fiJ, M!!IJ:tf!
~ 10. ooo JLJUJ:c -t ~ ~ ii U~W±itl'.ttfjA.0)1il'%iJ>t.r~'o t£)'1-0)~J;l.liiP !It c Lt~lts:toJ:
U!f.J~J!JUlllfi!fmO)IlliJ:(Ii ~:;!:f:J" 1.> o*:t'O)I'H-~a~~~ ·?iffi~ • tifJ!YJtl>.t.rC:I;:J: ~o~1~~ 1 7if.f.I.J:il'utfi Ltt.:#~OJ: U;,}:~O)i; ~>etTj'"t{ic.}O)ib-:> t.:·tnt.. ;Ji!~f:l.~O).lj~~Tiij;~T~ L. tiJ;-c ~ oo~15~*~O)~Dtl~GI~. ~-n~~tv.

5&~~ii~mm•ac.-r;:,oeMua-r2~tv

ti.fi:l!lH.fjtft.r.t',t~RO)~ll!li!Z:..J: ~ *:t:lL~'n:toJ: V"~:iG:tiT-:7. ~ill< L. iJ:-c ~ 7.> o
1iffh:ffi!fHiiW,l&Ji.l.±:z:.~Jt 'Tlii.m ~tt i5 o J'l!~.8.1i!J'MJU~f.l.0!fliJ> l?>:t~ 0) imi~.Q~P1- IC.J:-:> T.i:MIU ~tt;:,"~ ~I:J:tHJUt;f{O)rj1iJ> G.W~t:l.~lt~;;t 'Ti:l!llll ~ ~t. ;;$:t;>e f-t~ v~f&~l[fl!l!'"t oo~~IOfi,l&<b ;:,t~{;IJ:01H>~rt!jl<::f~l!l!~~lllliT 'Sa;4>.:~t;::t;t~~0.lH:·f/'!t< ~ CIJ~~~ ¢, ~:ft11fH.tilf~ftfM:tJEJ!!H. ~~ 0) ik~K .t -?"LJE"'->¢, i5tH~R:Iiil¥~~H~i;::~jJU-to;:: ciJ:~~ ¢,fl118~;,}:01i1iJ~-ful/'.tf71JW~~g~< -'(O)~i'tfZ:.Ii~~iJ>~ IJ ;;$:~.i!J!~O)~;;$:;fj~~tl(Jt-r 7.>oo*£i';O)~~~:tJ¥~.CH~iJ:etl£T;:, o0Nii£,J!!iJ.>J.J 1.> c ~!0 J..H]IillliH!!W~~Hm-t oo •:tlit;~O)+J}O)-f.J.J:O) Ul!r~ ~ t--::>-r~:i'l:-9'5,i';l:H1~ft0)=:.0)5}-£J.l:O):fi·t;:k~O) I::Jil':Jt_: 7.> '.IJlJUo.J: UfJ~O)J!l!I:H~ItH~ L..t.:.Jf
Tk~~~E!I!OJ~?J;.R~~U f.:JJ,\frl.il&lilt!ft@~~ fJ!f!T 1.> ot~~IZ:.U\1,'?; vt.H '~Hii!il!O)!f.\i,'{,:ftHIZ:.-'CO)~#tf'mO)fi~~~ ffT 7.> L.
~·lt0J.~O)l!lifJ;tf1tiO)ftfili 1 A 1 i;n:Jn~olin <a-t-?

c iJ>-c ~ ~o {fl (..,, 'x.

fn20~

tit~O);~&iJ~l:J:v r<tkr:.l: ~ .

m21*~:IH>J:U;if-;i,&W.ri;W.~W.~~t.lll~v. k~~0#t;~&r:J: i5t.1i-*1J~f·KVft'.i!l!'l:t!1l!J1l ~ 'li'f.ll#tlt

m22~

1~ f6~.li'~l:l'ii~f.6ti!fl0Hfi~ vlW.mt:l.~O)iJ~.J&r<::.Jk:::it 'T~f.6~t4nT o"*~O)~;tf?rJJUi !J.iiJ! 1 FJ 1 EJ rc.!lll ;t ~ 12 ff 31 u rr..~ o 0*~~nu~t.:!l.!-r;;, 'c::tf.l;;fzl: It.:cqJUI\'i):tft.O)=::J.tO)=£J.J:O)fill

j'ff23~£H 2,1 tt.:

UJ lliri1J&O)Ii'!ili.l&!iiJ> c:::. ~t<a-t!(~;:,,-r 1..o.14v.

f.(t 'o{i.IIIJ1) il'H&.W.£:CT;;;Dit).!I:J:~TJH€Jo1Hit\'JH::.J: ~,m~fiF"tt:tt:i '" ~;