Transactions and Proceedingsof thePalaeontological Society of Japan
New Series

No. 88Palaeontological Society of Japan
December 20, 1972

---------------------·-·
Editor:Takashi HAMADA

Associate Editor: Yasuhide IWASAKI

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Trans. Proc. Palaecint. Soc. Japan, N. S., No. 88, pp. 447-457, pl. 54, December 20, 1972

607.

UPPER TRIASSIC CEPHALOPODS FROM THE TANOURA
DISTRICT, KUMAMOTO PREFECTURE, JAPAN*
T AKESHI ISHIBASHI
Department of Geology, Faculty of Science, Kyushu University

Jm;fs:~llliinit!!JrRO)J:ll!I.:::.~*~~.:Em = lm**~HI\.

mim!i!Wll::~:tn-t7.> .=:.t~l±liilll!

:t,;.tCJ'!liili"'JI.t IJ imllit...t-:. \!J:.lEID~#it-tt..~ifi*· ;;t-l,~W nt. ~:Pii stm~~ll'.t..
t-:., ? ~~1i 1 f!H1¥rf<1l't'<l:>oo .:.tt,~O)iji.lEmri'l'ft~'t'li<I:>.Oil:, 8*0).:::.:\ilf..t 1HJJII.>"Cffl'.!i' <Ht.7.> ~lim t <!:> 1J, i!ti.lEfflitJ#JtlfO)lVf~i;:l;tiJtli:"t' <1; .o. .:.tt. 'f "t':i':l::::. =tHHt:ril:t c-? ~~~~ttn:~n.: Mt. -c~.o,t~n:. .:. .:. "t'iii.lEmft1i~:. J: .o IF.fttl::::. ··:w-cfi=f~ t:.. llliitl/1
O)J~t'lKiiHil-~7:--'l::i~-t.ORfi'mM:~mmit...t=,

:Pt~

*:1

upper members. The lower member
yields cephalopod fossils, which werecollected from the following localities
denoted by numbers** :Introduction and aclmowledgments

The cephalopods described in this ar·
ticle were collected from the Upper
Triassic Tanoura (TAMURA et a!., 1958)
and Takagochi (ORITA, 1962) Formations
distributing in the Tanoura district,
Kumamoto Prefecture. TAMURA (1959)
and ORITA (1962) studied the stratigra·
phy and molluscan fossils of the Tanoura Formation. Almost all the specimens used in this study were collectedby them, except for a few which werecollected by myself.
ORITA (1962) listed several ammonoids
which were preliminarily examined by
MATSUMOTO. Based on the bivalves andammonoids, the Tanoura Formation wasreferred to as the Carnian and the Ta·
kagochi Formation as the Carnian toNorian in geologic age (TAMURA 1959;
0RITA. 1962).
According to ORITA (op. cit.), the Tanoura Formation is about 250m in thick·ness and is divided into the lower and· Loc. 85 Buchites kumamotoensis n. sp.
Loc. 43 Phormedites sp.
Loc. 46 Rhacophyllites sp.
Loc. 25 Buchites kumamotoensis n. sp.
Arcestes sp. cf. A. (Proarcestes)ausseeanus (HAUER)
Ussuritidae or Discophyllitidae
gen. et sp. indet.
Loc. 44 Holconautilus sp.
The Takagochi Formation is about
400 m in thickness (ORITA, op. cit.), andyields molluscan fossils through out theformation. Ammonoid was found in its
middle part (Joe. 52 and 55), and is represented by only a single species, Piacites sp. aff. P. oxyphyllus (MOJSISOVICS).
This ammonite is associated with Pleuromysidia kanmerai TAMURA, Paleoneil~
iwaiensis lCHIKA wA, Halobia sp ., etc .
Pleuromysidia kanmerai TAMURA and* Received May 25, 1972; read Jan. 23,
1972 .at Chiba.

•• The numbers of locality correspond withthose of 0RITI. (1962).

447

448

0

Takeski ISHIBASHI

3KM

Text-fig. 1.

Location of samples collected.

Halobia (?) sp. occur in the lower part(loc. 54), whereas 1Honotis (Entomonotis)
sp. cf. M. (E.) typica (KIPARISOVA) is
known from the upper part (toe. 56).
On the basis of the molluscan fossils
ORITA (op. cit.) concluded that the upper
part was early Norian, and the middle
and lower parts were Carnian in age.
NAKAZAWA (1964b) also considered thatthe middle horizon (locs. 52, 55) of Takagochi Formation was referable to theLate Carnian.
I investigated the cephalopod fossils
collected from the Tanoura and Takagochi Formations, and identified one
nautiloid species and five ammonoid species, of Which one is new. On the basisof this ammonoid faunule the lower
member of the Tanoura Formation iscorrelated with the L~te Carnian. Accordingly the upper member probably
extends to the Early Norian. Placites
sp. aff. oxyphyllus from the middle partof Takagochi Formation is suggestiveof the Norian, conservatively the CarnoNorian age.
The cephalopod fauna of the present
study generally resembles that of Saragai Group of Northeast Japan which includes species of Placites, Rhacophyllites
and Arcestes (SHIMIZU and MABUCHI,
1932; NAKAZAWA, 1964a).
ISHIBASHI(1970) reported numerous Carnian ammonites from the Okinawa-jima, butlittle genera are common between the
Tanoura district and the Okinawa-jima,
except for Arcestes. Two species of
Buchites and Phormedites to be describedin this article are known from the Ja·
panese Islands for the first time.
I wish to express my sincere thanks
to Professor Tatsuro MATSUMOTO of
Kyushu University for his arrangementsof the material. criticisms and readingof the manuscript, and to Professor
Ryuzo TORIY AMA of the same university

607.

Triassic cephalopods from Tanoum"'for kindly reading preliminary draft and-encouragement. Thanks are also due to. Professor Minoru TAMURA of Kumamo··to University for the offer of his ammonoid collections. I am obliged to Miss
:Seiko 1-IAYAKAWA for typewriting the.manuscript.
Repository.-All specimens illustrated··in this article are deposited· in the De_partment of Geology, Kyushu University·with the registered number using a:symbol of GK. F.
Systematic paleontology

Class Cephalopoda CUVIER, 1797
:Subciass Ammonoidea AGASSIZ, 1847
Order Ceratitida HYATT, 1884
Superfamily Clydonitaceae
MOJSISOV!CS, 1879
Family Buchitidae HYATT, 1900
Genus Buchites MOJSISOVICS, 1893

Type-species.-Ceratites (Buchites) al·
cdrovandii MOJSISOVICS, 1893.
Remarks.-The genus Buchites was·originally established by MOJSrsovrcs•(1893) as one of the subgenera of Cera·
tites. The species belonging to Buchites. are known only from the Carnian toNorian of the Tethys region, such as·the Alps, Greece, Sicily, Himalayas and·Timor. Buchites has flexuous ribs which:are projected at the ventrolateral shoul·-der and gradually disappear on the·rounded venter but occasionally cross·over it in the last whorl of a mature·specimen. Whorls are moderately involute, increasing its height at a mo··derate rate. Ribs are distinct and in··.tercalated with the short ones.

449

Buchites kwrlamotoensis ISHIBASHI,
sp. nov .
Pl. 54, Figs'. 1-9, 10? and 11?
1962. Helictites? sp., ORITA, Sci. Rept., Fac.
Sci., Kyushu Univ., Vol. 6, No. 1, p. 4,
listed .

Material.-The holotype (GK. F 458)
and some of paratypes (GK. F 459-462),
collected by Dr. TAMURA from the Uminoura area (loc. 25), Kumamoto · Prefec·
ture. Two others (GK. F 374, 375) listed
already as Helictites ? sp. collected by
Mr. ORITA and the rest (GK. F 463-466)
by myself.
Description.-Shell considerably involute and compressed, increasing gradually its height.
Whorl embracing
about a half or one-third of the inner
whorl; umbilicus narrow and moderately
deep; ribs strong, rounded, flexiradiateon the flank and projected on the ven·
trolateral part, with irregularly inter·calated secondaries, without crossing
over the venter ; venter round, smoothwhere ribs gradually disappear ; suture
ceratitic. with serrated lobes.
Remarks.-The present species is some·what similar to Helictites beneckei re·ported by MOJSISOvrcs (1893) from theAlps in lateral view, but the ribs of thelatter cross straight over venter .
The present new species rather resem·
bles Buchites aldrovandii {MOJSISOV!CS)
from the Alps but the whorl of thelatter is more evolute than that of theformer.
The other known species from theAlps and Himalayas are characterized by
the appearance of the flexuous shortlateral ribs at the ventrolateral area. B.
kumamotoensis has bifurcating ribs someof ~hich diverge from the main ribs ator near the umbilical shoulder, and its
suture-line has the serrated accessary

450'7 alcesh'i [SffiBASfll
1lobes. The suture:iine of B. kumamotoensis is rather· similar to B. modestus(BUCH) .from the Alps, but the latter isevidently different in that the ribs sli·
.ghtly cross over the venter and has a·broad U:rhbilicus. B. emersoni and B. cf.
hilaris described by DIENER (1906) haveevolute whorls and primitive sutures.
Buchites heteroplichus reported by
GEMMELLARO (1904) is very similar tothe present new species with respects
to only the whoi! volution. but is clif·
ferent from the latter in the characteristics of ribs, whorl size, venter, etc. It
is resonable to establish a new species
in the genus Buchites for the present
form.
Occurrence.-Locs. 25 and 85. With Arcestes sp. cf. A. (Proarcestes) ausseeanus(HAUER) and Halobia ? sp. (Loc. 25). ani:!
Pleuromysidia kanmerai TAMURA (Loc.
85).
Genus Phormedites MOJSISOVICS, 1893
Type-species.-Ceratites (Phormedites)
juvavicus MOJSISovrcs, 1893

Remarks.-MOJSrsovrcs (1893) establi·
shed the subgenus Phormedites in thegenus Ceratites of the family Ceratitidae.
ZITTEL (1900). GEMMELLARO (1904) and
DIENER (1906) also treated it as a sub·genus of Ceratites. DIENER (1923) ranked
it as a genus of the same family. but
SPATH (1951) included the genus Phormedites in the family Buchitidae. SPATH's
classification seems to be reasonable atpresent. The species referable to thisgenus, including the type-species designated by DIENER (19.15). have been
known from the Carnian and Norianages. The Carnian species. Ph, pygmaeus and Ph. schopeni were describedby GEMMELLARO (1904) from Sicily, andPh. aff. pygmaeus was reported by PAKU·CKAS (1928) from Timor. On the ·other
hand, the Norian one, Ph. juvavicus was~described . by MOJSisovrcs (1893) from,the Alps and by PAKUCKAS (1928) from
Timor. Ph. fascia/us was reported bY'
MOJSISOVICS (1893), DIENER (1906) ana
PAKUCKAS (1928) from the Alps, Hima-layas and Timor, respectively.
Based on the original descriptions of'these species, species of Phormedites are"divided into the following two groups:.
1. The group of Ph. schopeni-Ph.
juvavicus· Shell is consideral::lly involute with.
fine, prorsiradiate ribs. The ribs cross ..
over the venter. The venter is round ..
The type-species, Ph. juvavicus, is about.
28 mm in diameter.
2. The group of Ph. pygmaeus-Ph.
fasciatus
The shell is evolute with fine, prorsi-radiate ribs. The ribs are bundled. The'
venter is roundt:d with a faint. groove ..
Ph. fasciatus is about 32 mm in diameter ..
The characteristics of this group arevery similar to those of the genus DaPhinites as pointed out by MOJSISOVICS.
(1893).
Phormedites sp. indet.
Pl. 54, Figs. 13 and 14
1962. Phormedites (?) sp .. OR ITA, Sci. Rept ., .

Fac. Sci., Kyushu Univ., Vol. 6, No. 1,
p. 4, listed.

Material.-A part of outer whorl of'
rubber cast which was made from anexternal mould (GK. F 377a), and a fragmental part of ventral part (GK. F 377b).
These specimens were collected by Mr.
ORITA.
Descriptive remarlzs.-The shell is
considerably involute and compressed.
Judging from outer whorl. it is at least
about 60 mm in diameter, though the607.

Triassic cephalopods [rom Tanotwa,umbilical part is l.ost.. The venter is·narrowly arc,!}ed. The l~teral surface·of sh6ll .has, numerous, g~ntly fiexira·diate· ri~~',which: are· ~t'Qji~te.d· sharply
for~war\f.r at. the~ ventrolateral shoulder.
The rib.$. are: sl.ender, keep, almost the·same ~i'dth fro'm:'·the umbilital shoulder·to the. ve.qtral· part, and· cross over theventer. ·The i~tercostal space is v.ery
narrow·near: u.mbilical shoulder. These· ch~ra.cters ·s\ligest that the present spe·
· cim.ens .beJon'g- to the genus Plzormedites.
1\;Tiq~·g: the sp.ecies of Phormedites,-only two, Ph. pygmaeus and Ph. schopeni,
have been known from the Carnian of:Sicily. Others hav.e been reported. from·the Norian of the Alps, Himalayas and·Timor. As mentioned above, all these·species are smaller than the species
from Tanoura, being one·third or a halfin diameter. There are many differencesin respect to ribs, volution, size andventer between the species from theAlps, Sicily and Japan.
The specimens at hand are so poor·that the present species has been left
unnamed until better preserved material:becomes available.
Occurrence.-Loc. 43. Shale bed ofupper Lower Member of Tanoura For·mation. With Halobia sp., Halobia mo·.lukkana WANNER, and Pleuromya fors.bergi (BOHM).
Superfamily Arcestaceae
MOJS!SOV!CS, 1875
Family Arcestidae MoJsrsovrcs,
1875
Genus Arcestes SUESS, 1865
1ype-species.-Ammonites galeiformis
HAUER, 1850
Subgenus Proarcestes MOJS!SOVICS, 1893

4,51

Type-species.-Arcestes bramantei
MOJSISOVICS, 1869.

Arcestes (Proarcestes) sp. cf. A. (P.)
ausseeanus (HAUER)
Pl. 54, Fig. 15, Text-figs. 2 and 3

Compars1847. Ammonites

ausseeanus HAUER, Hai·
dinge1·s Nat. Abhandl., I, p. 268. pl. 8,
figs. 6-8.
1875. llrcestes (Proarcestes) ausseeanus HA·
UER: MoJSISOVICS, ]ahrb. Geol. Rei·chsanst., Wien, No. 6, Heft. 1, pl. 51,
figs. 1 and 4, pl. 53, figs. 28 and 31.
1907. Arcestes (P.) ausseeanus HAUER: FRE·CH, N. ]ahrb. f. Min. Festbd., p. 20. pl.
4. fig. 5.
1914. Arcestes (P.)
bicarinatus-ausseeanus
HAUER: WELTER, Paliiont. von Timor.
I, p. 180.
1927. Arcestes (P.) ausseeanus HAUER: ARTHABER, ]ahrb. Mijnw. Nederl. Ind ..
Vol. 55. No. 2, p. 51, pi 5, figs. 1-6.

Material.-One external mould (GK. F
467) and its plaster cast (GK. F 468) areexamined here. The external mould
was collected by Dr. M. TAMURA of
Kumamoto University.
Description.-Shell, involute. globose,
about 30 mm in diameter ; whorl. broad,
24 mm in width deeply embracing andindented by the inner volutions ; helmet·
shape in section : surface with threeoblique, periodic constrictions cross overthe ventral part, no striae and ribs;
venter narrow. its shoulder abruptly
rounded; suture, ammonitic, deeply di·gitate, having numerous lobes and saddles, with accessary lobes.
Remarfls.-Arcestes (Proarcestes) aus·
seeanus has been known from the Car·nian of the Northern Alps, Timor.
Greece, Himalayas. etc. This species
was originally described by HAUER(1847). MOJSISOV!CS (1873) included thisTakeshi ISHIBASHI

452

species into the group of A. (P.) bicari·
natus (MUNSTER), and RENZ (1910) and
WELTER (1914) reported ausseeanus as avariety of A. (P.) bicm·inatus. A. (P.)
ausseeanus has three constrictions on
the lateral part. but the specimens described by MOJS!SOVICS (1875, pl. 51. figs.
4a-b) have evidently four constrictions.
ARTHABER (1927) described A. (P.) aus·
seeanus from Timor. One of the Timor
specimens (pl. 5, fig. 5) is immature andhas two constrictions, but the othermature ones have four constrictions. A.
(P.) hanieli, which was originally described by WELTER (1914) from Timor,
was also reported by ARTHABER (1927)
from the same region. A. (P.) hanieli
described by ARTHABER closely resem·
bles A. (P.) ausseeanus, though the !at·ter has a higher whorl than the former.
SMITH (1927) described several new
species of Proarcestes from the Carnianof California. of which A. (P.) pacificus
is similar to the present specimen inwhorl section, but the former has asmaller shell and a simpler, flared suture·line in which the accessary lobes. are notdeveloped.
The following species of arcestids havebeen known from the Japanese Islands:_...- -.__

/

/

'

I

I
I
\
\

I

... 't"',....'\ ', __
\
\

' ,_
0

10

- - - - - mm

'Text-fig. 2. Whorl section of Arcestes(Proarcestes) cf. ausseeanus (HAUER).

Text-fig. 3. Suture-line preserved atthe external mould of Arcestes (Proarce ..
stes) cf. au sse anus (HAUER). x 5.

Arcestes (A.) aff. oligosarcus MOJSISOVICS ..
SIIIMIZU and MABUCHI (1932) (Saragail
Group, Kitakami)
Arcestes sp. NAKAZAWA (1964) (Saragai~
Group, Kitakami)
Arcestes (Stenarcestes) sp. NAKAZAWA (1959)·(Jito Formation, Nariwa, Okayama)
Arcestes (Proarcestes) aff. hanieli WELTER~SHIMIZU (1931) (Proarcestes Bed, !CHI~
KAWA (1951))
Arcestes (P.) aff. bicarinatus (MUNsTER)SHIMIZU (1931) (Kochigatani Group, Shi-koku)

The first two species came from the,Norian strata, whereas the last thre~from the Carnian. The present species.
is easily distinguished from these species by whorl section, suture-line ancl
other characteristics.
To sum up the present specimen is.
best comparable to and may be identified with A. (P.) ausseeanus with respects to whorl section, ornamentation,.
suture-line and other characteristics.
Occurrence.-Loc. 25. Shale bed ofupper Lower Member of Tanoura Forma-

607.

Triassic cephalopods from Tanoura. . ·.; '· .

;

.

;.,

.

453

.

. \

O__________________________
IOmm

Text-fig. 4.

Suture-line of Placites aff. oxyphyllus (MOJSISOVICS).

tion, where Buchites kumamotoensis IsruBASI·II and Halobia ? sp. are associated.
Superfamily Pinacocerataceae
MOJSISOV!CS, 1879
Family Pinacoceratidae
MOJS!SOV!CS, 1879
Genus Placites MOJSISOVICS, 1896
Placites sp. aff. P. oxyphyllus(MOJSISOV!CS)Pl. 54, Figs. 12a-c and Text-fig. 4
1962. Placites? sp.; ORITA, Sci. Rept., Fac.
Sci., Kyushu Univ., Vol. 6, No. 1, p. 9,
listed.

Material.-One individual formed ofthree parts (GK. F 379) is here examined.
They are two external moulds and aninternal mould collected by Mr. ORITA.
Descriptive remarks.- The shell is com·pressed with a narrowly arched venter,rather involute and smooth on· the surface. The essential part of the suture
is well preserved on the present specimen, showing the typical pattern ofPlacites, which consists of numerous,
deeply incised elements (Text-fig. 4).
The present species closely resembles
PL. oxyphyllus (MOJSISOV!CS, 1873), fromthe Norian of the Alps and Sicily, inthat its external saddle has shallow lobules. Placites perauctus (MOJSISOVICS,
1873), from the Norian of the Alps, has
deep lobules on the external saddle but
differs from Pl. oxyphyllus by the deep
lobules in the lateral saddles.
Two ammonoids, Placites aff. oxyphllus
and Arcestes aff. oligosarcus have been
listed by SHIMIZU and MABUCHI (1932)
from the Norian Saragai Group, Kita·
kami. but their specimens have been lost
(NAKAZAWA, 1964a p. 25). So it is impossible to compare the present specimen with them. The Takagochi Formation, in which the present specimen J
occurs. has been considered by ORIT A

454

Takeshi ISHIBASHI(1962) as the Carno-Norian in age. Thereis a possibility that the bed containing
Placites aff. oxyphyllus belongs to the
Norian age, though the present speci·
men is not so well preserved that the
specific name has not been determinedwith certainty.
Occurrence.-Loc. 52. Shale in the
middle part of Takagochi Formation,
Kumamoto Prefecture. It occurs with
Pleuromysidia kanmerai TAMURA, Paleoneilo iwaiensis ICHIKAwA, Halobia spp.,
Camptonectes sp., Oxytoma sp., etc. according to ORITA (1962).

Order Phyltoceratida ARKELL, 1950
Superfamily Phyttocerataceae
ZITTEL, 1884
Family Discophyllitidae SPATH, 1927
Genus Rhacophyllites ZITTEL, 1884
Rhacophyllites sp. indet.
Pl. 54, Fig. 16 and Text-fig. 5
1962. Rhacophyllites? sp.; ORITA, Sci. Rept.,
Fac. Sci., Kyusltu Univ., Vol. 6, No. 1,
p. 4, listed.
1964. ·Rhacophyllites sp.; NAKAZAWA, Mem.
Col!. Sci., Univ. Kyoto, .Vol. 30, No. 4,
p: 29, 4th line from the top.

JV!aterial.-One fragmental specimen(GK. F 371) is examined here, which wascollected by Mr. ORIT A.
Descriptive remarks.-The shell is in·volute with a rounded venter. The urn·
bilical part is almost lost. The sutureis exposed on the ventrolateral part ofthe shell, in which the first lateral sad·dle shows a diphyllitic type (Text-fig. 5).
The phylloid leaf is slender for the
genus, probably because of the immaturestate of the shell. Rhacophyllites is closeo_________;_:IO mm
Text-fig. 5. Suture-line of Rhacophyllites sp.

to Discophyllites, in the whorl shape andornamentation. The difference of thetwo genera is mainly in the pattern of
the sutures. Namely Rhacophyllites isdiphyllitic or triphyllitic, whereas Dis·
cophyllites is monophytlitic. The speciesof Rh. are known from the Upper Carnian to Norian in the various part of
the world.
When NAKAZAWA (1964a) described
Rhacophyllites sp. from the Saragai
Group, the present specimen was examined by him. He said that the characters of Rh. sp. of the Saragai Group
resembled especially those of the Car·nian Rhacophyllites sp. of the Tanoura
Formation rather than those of Norianspecies such as Rh. debilis and Rh. neo·jurensis and those of Carnian species
reported by TRECHMANN {1917) as Discophyllites cf. ebne1·i.
As mentioned above, the species of
Rhacophyllites from the Tanoura Formation and the Saragai Group certainly
resemble each other in the character ofsuture. The sutural patterns of these
two specimens are also quite similar tothat of Rh. ·· neojurensis" illustrated by607.

Triassic cephalopods from Tanoura

WELTER (1914, pl. 30, fig. 6*, text-fig. 72)
from the Norian of Timor which was·excluded by WIEDMANN (1970) from the·synonym list of the true neojurensis of
WELTER. I would support the opinion·of WIEDMANN. Unfortunately the spe··cific name was not given by WIEDMANN
for the Timor specimens at that time.
On the basis of suture-line, the present·specimen may be suggestive of the Early
Norian in age.
Occurrence.-Loc. 46. Shale of the,upper Lower Member of Tanoura For·mation. With Pleuromysidia kanmerai"TAMURA.
.Subclass Nautiloidea AGASSIZ, 1847
Order Nautilida AGASSIZ, 1847
Superfamily Tainoceratacea
HYATT, 1883
Family Tainoceratidae HYATT, 1883
cGenus Holconautilus MOJSISOVICS, 1902
Type-species.-Nautilus semicostatus
8EYRICH, 1867
Holconantilus sp. i ndet.
Pl. 54, Fig. 17

Material.-One specimen (GK. F 378)
·collected by Mr. ORITA, in 1962. is exa·mined here.
Descriptive remarlzs.-Only outer whorl{ca. 60 mm in diameter) is preserved.
The shell is very evolute.and presumably
widely umbilicate. The whorl is nearly.as high as broad, increasing its heightgradually. The rectiradiate, ·distant ribs

* WIEDMANN did not exclude the figs. 5;and 7 in the same plate but the suture of.fig. 6 seemed to be drawn from the specimen•Of figs. 5 and 7.

455

develop on the flank. The venter isround and smooth. The suture-line runs
almost parallel to ribs and shows one
lateral lobe. ~The siphonal position is
not observabie in the present specimen.
On the grounds of the observed cha·racters the present specimen is referable
to the genus Holconautilus. The speciesof Holconautilus are known from theAnisian to Carnian of the Alps, Ger·
many, the Meditteranean region, Timor.
and Japan. The present specimen isclosely similar to H. semicostatus (BEY·RICH) in volution. ribs and suture. The
other species of Holconautilus are dif·ferent from the present species in thecharacters of whorl. The whorls in theformer species increase height quicker
than H. semicostatus and the present
species, and sometimes have a ventro·
lateral tubercle on the rib.
BANDO (1964) described Pleuronautilus(Holconautilus) yabei BANDO from theLadinian Rifu Formation of Northeast
Japan. The present specimen is evi·
dently distinguished from it in that it
has a quadrate whorl section, with aflattened venter, two spiral rows ofnodes on the venter and less prominent
ribs.
Pleuronautilus(?) sp. reported by YABEand SHIMIZU (1927) without text-figure
is ornamented with distant umbilicaltubercles.
Occurrence.-Loc. 44. Shale bed of theLower Member of Tanoura Formation
with Freguelliella (l(umatrigo11ia) tanourensis TAMURA, Balcevelliid sp., Costatoria (?) sp.
References cited
ARTHABER, G. von (1927) : Ammonoidea
Leiostraca aus der oberen Trias vonTimor. 2 Nederl. Timor Expeditie 19~6
onder Ieiding van Dr. H. G. ]ONK~R :·

~

. .

. '. \ ;~ :·. ~...~. . :. ,..

456

Talceshi ISHIBASHI

Utigegeven soor Dr. H. A. BROUWER, 4,
jaarb. Mijnw. Nederl. Ind., Vol. 55, Pt.
2, pp. 1-174, pis. 1-20 (1926)
BANDO, Y. (1964): On some Middle Triassic fossil cephalopods from Japan, witha note on the Middle Triassic formationsin Japan. japan. jour. Geol. Geogr .. Vol.
35, Nos. 2-4, pp. 122-137, pl. 5.
DIENER, C. (1906) : Fauna of the Tropites·
Limestone of Byans. Palaeont Indica,Ser. 15, Vol. 5, No. 1, pp. 1-201, pis. 1-17.
- - (1915) : Cephalopoda triadica. Fossilium
Catalogus, Altimalia I, Pars 8, pp. 1-369,
Junk, Berlin.
- - (1920) : Die Ceratitoidea der karnischnorischen Mischfauna des Feuerkogels
bei Aussee.
Sitzungsber. Akad. Wiss.
Wien, Vol. 129, pp. 589-618, pis. 1-3.
- - (1923) :Ammonoidea Trachyostraca
aus der mittleren und oberen Trias vonTimor. jaarb. Mijnw. Nederl. Ind., Vol.
49, pp. 73-276, pis. 1-23.
FRECH, F. (1907):
Die Hallstlitter Kalke
bei Epidauros (Argolis) und ihre cephalo·
poden. Neues ]ahrb. Min. Geol. Paliiont.
(Fest Bd.), pp. 1-32, pis. 1-6.
GEMMELLARO, G. G. (1904) : I cefalopodi del
Trias superiore della regione occidentaledella Sicilia. Giornale Sci. Nat. Econ.,
Palermo, Vol. 24, pp. 1-319, pis. 1-30.
I-lAUER, F. von (1847): Neue Cephalopoden
aus dem rothen Marmor von Aussee.
/laidinger's Naturwiss. Abhandl., I, pp.
257-277, pis. 7-9.
IcHIKAWA, K. (1951): Triassic formation at
Taho district, Ehime Prefecture. in the

Triassic stratigraphy of Japan. GeoL
Surv. japan, Rep Spec. No., pp. 108-110.
(in Japanese)
ISHIBASHI, T. (1970) : Upper Triassic Ammonites from Okinawa-jima, Part 1. Mem.
Fac. Sci., Kyushu Univ., Ser. D, Geo/.,.
Vol. 22, No. 2, pp. 195-223, pis. 26-29.
MoJSISOV.ICS, E. von (1869) : Beitrlige zur
Kenntniss der Cephalopoden-fauna desc
alpinen Muschelkalks (Zone des Arceste.s:
studeri). jahrb. Geol. Reichsanst., Wien,.
Vol. 19, pp. 567-594, pis. 15-19.
- - (1873) : Die l\4ollusken-Faunen der ZJa.
mbach· und Hallstatter-Schichten. Abhandl. Geol. Reichsanst., Wien, Vol. 6,
No. 1, pp. 1-82, pis. 1-32.
- - (1893) : Die Cephalopoden der Hallstatter Kalke. Abhand/. Geol. Reichsanst.,
Wien, Vol. 6, Pt. 2, pp. 1-835, pis. 71200.
NAKAZAWA, K. (1959): Two cephalopod
species from the Norian Nariwa GrouP'in Okayama Prefecture, West Japan.
japan. jour. Geol. Geogr .. Vol. 30, pp.
127-133, pl. 11.
- - (1964a) : On· the Monotis typica Zone inJapan. Mem. Col/. Sci., Univ. Kyoto, Ser.
B, Vol. 30, No. 4, pp. 21-39, pis. 3-5.
- - (1964b) : On the Upper Triassic Monotis
Beds, especially, on the Monotis typica
Zone. jour. Geol. Soc. japan, Vol. 70,
No. 829, pp. 523-535. (in Japanese withEnglish abstract)
ORITA, Y. (1962): Upper Triassic Series in
the Tanoura district, Kumamoto Prefec·
ture. Sci. Rep. Fac. Sci., Kyushu Univ.,

Explanation of Plate 54
Figs. 1-11. B11chites kumamotoensis IsHIBASHI, n. sp.
Fig. 1 GK. F 463: Fig. 2 GK. F 375 ventral part x2: Fig. 3 GK. F 459 x2: Fig. -i
GK..F 465: Fig. 5 GK. F 464: Fig. 6 GK. F 462: Fig. 7 GK. F 460: Fig. 8 GK. F 458,
Holotype: Fig. 9 GK. F 461 : Fig. 10 GK. F 374: Fig. 11 GK. F 466 x 2.
Figs. 12 a-c. Plactes sp. aff. P. oxyphyllus (MOJSISOVICS) GK. F 379 X 2.
Figs. 13 and 14. Phormedites sp. Fig. 13 GK.F 377b ventral part: Fig. 14 GK.F 377a.
Fig. 15. .4rcestes sp. cf. A. (Proarcestes) ausseeanus (HAUER) GK. F 468.
Fig. 16. Rhacophyllites sp. GK. F 371 ventral part.
Fig. 17. llolconautiltts sp. GK. F 378.
(Figures in natural size except as indicated)

ISHIBASHI :Upper Triassic cephalopods

Plate 54

607.

Ser. Geol., Vol. 6, No. 1, pp. 1-13, 2 maps,(in Japanese with English abstract)
PAKUCKAS, C. (1928) : Nachtrag zur mittel·
und obertriadischen Fauna der Ammonea
trachyostraca, C. DIENER's aus Timor,
mit Einleitung und stratigraphischer Zu·
sammenfassung von G. von ARTHABER.
jaarb. Mijnw. Nederl. Ind., Vol. 56. pp.
143-218, pls. 1-2.
RENZ, C. (1910) : Die Mesozoischen Faunen
Griechenlands. Theil 1. Die Triadischen
Faunen der Argolis. Palaeontographica,Vol. 58, pp. 1-104, pls. 1-7.
SHIMIZU, S. (1931):
Notes on two Carnic
species of Proarcestes from Shikoku.
japan. jour. Geol. Geogr., Vol. 8, No. 3,
pp. 117-121, pis. 12-13.
- - and MAsucHI, S. (1932): Upper Triassic in the Kitakami Massif. jour. Geol.
Soc. japan, Vol. 39, No. 465, pp. 313-317(in Japanese).
SMITH, ]. P. (1927) : Upper Triassic marine
invertebrate faunas of North America.
U.S. Geol. Surv., Prof. Pap. 141, pp. 1261, pis. 1-71.
SPATH, L. F. (1951): The Ammonoidea of theTrias (II). Cat. Fossil Cephalopoda Brit.
Mus. (Nat. Hist.), Pt. 5, 209 p.
On Kumatrigonia, a
TAMURA, M. (1959):
new subgenus of Frenguelliella, and a]ito
Kitakami
Kochigatani
NariwaOkinawa-jima
Ota

457

T?·iassic cepha~opods jrom Ta•noura

j1J!.

iiJi

:lt

J::

i"II*W~$.
~~i'P*'l.l%~[lJ

Tosapecten from the Carnic Tanoura formation in Kyushu, Japan. Mem. Fac.
Educ., Kumamoto Univ, Vol. 7, pp. 212224, pl. 2 .. :- - (1960) : Stratigraphical study on theSakamoto Group in Kyushu. jour. Geol.
Soc. japan, Vol. 66, No. 777, pp. 371-383(in Japanese with English abstract).
- - , KANMERA, K., AKAZU, K. and YAMA·SHITA, A. (1958): Notes on some newfinds on the Mesozoic strata in the Kuma.
and Yatsushiro Districts. jour. Geol Soc.
japan, Vol. 64, p. 370 (in Japanese).
TRECHMANN, C. T. (1917): The Trias of·
New Zealand. Quart. jour. Geol. Soc.
London, Vol. 73, pp. 165-245, pis. 17-25.
WELTER, 0. A. (1914): Die obertriadischen.
Ammoniten und Nautiliden von Timor.
Paliiontologie von Timor, Vol. 1, pp. 1-·
258, pls. 1-36.
WIEDMANN, J. (1970) : Uber dem Ursprung
der Neoammonoideen-Das Problem einer·
Typogenes. Eclogae Geologicae Helvetiae,.
Vol. 63, No. 3, pp. 923-1020, pis. 1-10.
YABE, H. and SHIMIZU, S. (1927) : TheTriassic Fauna of Rifu near Sendai.
Tohoku Imp. Univ., Sci. Rep., 2nd, :Ser ..
Geol., Vol. 11, No. 2, pp. 101-136, pls.
10-14.

Rifu
SaragaiTakagochi
TanouraUminoura

li'U

lf.f

1D1

~

Al iiif r"l
IE
im
iii
iili

"Trans. Proc. Palaeont. Soc. Jagan,. N. S.., No .. 88 .. pp. 458-461, pl. 55 .. December 20, 1972

-608 .. A FOSSIL WOEM TRAIL FROM THE UPPER CRETACEOUS
NAKAMINATO FORMATION, 1BARAKJ". PREFECTURE*
KOTORA HAT AI and HIROSHI NODA
Institute of Geology and Paleontology, Faculty of Science,
Tohoku University, Japan$C!JI.i.)~U.:lltll3.illi*n11Jii.l~mffii ;:A,ii:-ff.JI::.:O~C."?v''L: ~!J£1'iHJFIJii.IIJfm 1HI!tiiUI;i!l: 5Hrd-~.1·.:111!3!!~!*JK>Jii.l~m!fltlil<7)lf~il!T$/11i'€> ilE!±ll...t.: liv'i!JFft~lifil~:iXQ)R:~*"l 29 em. m

1.2-2.0 em Q)flji!llllilfi~ 1..-bb !..-, f~ 3 em Q)ll!ISJ'I;:r.J 8 JlJQ)!J~J:f*ti'(;;?: Ji!.€>:11.. Jf?-~(~l:::tn{J.:.l-n, Nentes Q){ljlfh,(J).t? tJ:t\':IA:$1~fffi(J)tl/i:iX~Wl~JJ!i~1:ffl:l/:-ff.t.!tt?:h.tJ:It'. -:J:t:. U'l~J:f*[!li): ~n!J,;j" '.) Arenicola, Balanoglossus, Chaetopterus c <7) lt~~iit M'TfJ:-:> t,:;?:, .I!'~
P1Jffll~ffi{Q)t11,.,~{1:t.il1 Hitachia nakaminatoensis fJ:'.}~~. f/i~"eif.,'.);: c ~~~t.:.
~ ; 1:::. *ll:.:r:ii!E!fll;: J:: ~ lf-~tl!IRhf:*ilmJtffl~Q)ffii~rf.HJJilJIQ)lGi¥ifii~Q)JftfJ'H~l? ~ '.>;:

c '.i::~~l..-t.:.

WJ:J:t-1J'11f. g!fiiJfc!iffi

comprises chiefly dark gray sandstone
with interbedded graded sandstone layers." The Hiraiso-machi Member yieldedto SAITO (1961, p. 112), Didymoceras awajiense (Y ABE), D. nakaminatoense (SAITO),
Nipponaster nakaminatoensis SAITO, besides several dther 'fossil molluscs. And
the discovery of the fossil worm trail
verifies that the Hiraiso-machi Member
was deposited in shflllOw water.
The worm trail uppon which the pre·sent article is based was collected by
the junior writer during the writer's
field trip to the Nakaminato area withthe students majoring in geology of theFaculty of Education, lbaraki University,
guided by Associate Professor Toshio
SAITO and Research Associate Haruyuki TAKAHASIII of the same University.
At this place the writers express their
deep appreciation to Associate Professor
Toshio SAITO and Research Associate
Haruyuki TAKAHASHI for their kind
guidance to the Cretaceous deposits dis·tributed along the coast of Hiraiso, Na·

Introduction and acknowledgments

Exposed on t.he surface of a weather·-ed sandstone at the sea-side of Hiraiso.
Nakaminato City, lbaraki Prefecture,
was found a trail of a worm, probably·of marine origin. This trail, impressed·on the sandstone of the Hiraiso-machi
Member of the Nakaminato Formationis the first discovery of a trail fossil
from the member and is thus considered
important for interpretation of the con··ditions under which that part of theformation was deposited. The Hiraisomachi Member occupies the middle part·of the Nakaminato Formation of Upper·Cretaceous age.
According to SAITO (1961) the type
locality of the Hiraiso·machi Member of
the Nakaminato Formation is "at thesea coast of Hiraiso-machi where it at·
tains about 510 meters in thickness and* Received June 3, 1972; read June 3, 1972,.at Utsunomiya.

458

608.

Upper ·cretaceous worm trail

Tertiary System................... unconformity ...................................................................................~

Nakaminato Formation

Oarai Formation

lsoai Member, 730 m thick. Graded sandstone and siltstone.
Inoceramus cf. shikotanensis, Nipponaster nakaminatoensis,
Baculites inornatus, etc.
Hiraiso-machi Member, 510 m. thick. Siltstone with inter·ca:tated sandstone. Didymoceras awajiense, Pravitocerassp., etc.
Chikko Member, 35m thick. Granule ·sandstone in lower and.coarse to fine grained sandstone in upper .
\........................... conformity

I1030 m thicl<. Conglomerate intercalated with thin sandstonelayers, coal seams, and plant fossils.
· Base not exposed ·······--··-········--··························

kaminato City, and also for their information on the geology of the area.
Thanks are due to Mr. Kimiji KUMAGAI
for photographic work and to Mrs. Kimiko SHIBUYA for typing the manuscript.
Stratigraphic position of theworm trail
The stratigraphic position of theHiraiso-machi Member in the geologicalcolumn of the Cretaceous deposits distributed along the sea coast of Nai<aminato City, is, according to SAITO(1961), as follows.
The fossil traH forming the subjectof the present article was found in theupper part of the Hiraiso-machi Member.
and represents the first discovery of a
trail fossil from the Member. All of
the three members of the Nakaminato
Formation are conformable with one
another, unconformable with the superjacent Tertiary System, and also con·formable with the underlying Oarai
Formation.
The worm trail
The worm trail, which consists of asingle specimen, lacks both anterior andposterior extremities, and thus its original length remains unknown. But, as.
shown in the annexed figure, features.
suggesting the original segm·ents of thebody, and the approximate thickness of
the rounded .body are still preserved_
Since the sandstone ·layer in Which the.
worm trail is .preserved .is not overturned, the exposed q10rphological features represent the original .ventral -sur·face of the ·worm.
The worm trail measures. about 29 emin length measured along its longitu·
dina! middle part and the width, al·though variable at places according to.
the state ·Of preservation, measures from.
12 to 20 mm in minimum and maximum
widths, and there are about 8 segments.
within a distance of about 30 mm. The
body as can be judged from the preserved outline is rounded elongate. No·
longitudinal mesial depression or podialike structures are observed and features.
suggesting Nereites affinity could not be
observed.
Since the preserved segment·struc·tures are interpreted to represent theoriginal ventral surface of the worm,.
affinity is found with such living genera.
of marine worms as Arenicola, Balan'O·
glossus and Chaetoptents, all , of whichhave well developed segments. 'Among.

'

460

....... ".._ .. . -:;\Kotora HATAI and Hiroshi NODA

-the mentioned genera, the present fossil-worm trail shows resemblance with the
Recent species Arenico/a crista/a STIM·· PSON, now living along the coast of.Japan. That species, according to OKU·
DA and I!ZUKA (1949) attains about 60"90 mm in length and about 18 mm inwidth. Crista/a lives in sandy mud or
muddy sand in shallow depths along the"coast of Japa_n.
Arenicola claparedii
LEVINSON (OKUDA and IIZUKA, 1949),
which attains about 100-200mm in length:also lives in sandy mud or muddy sandin shallow water, and is distributed from·Central Japan to the Kurile Islands. Of·these two Recent species, the present·fossil worm impression seems to be
more similar to A. cristata than to A.
-claparedii, although differing from both.
Among the fossils reported by HO··WELL (1962), none expect possibly the·_genus Tosalorbis proposed by KATTO"(1962) based upon specimens from the
Eocene Muroto Formation in Kochi Pre·fecture, Shikoku, Japan, seem to be com··parable with the present fossil. How. ever, Tosalorbis differs from the pre··sent specimen in having closer set seg. ments and much shorter body length.
:Specimens comparable with the present. one were not noticed in the recently
published work "Trace Fossils" edited:by CRIMES and HARPER (1970).
Under the circumstances mentioned•above it seems that a new name should_given to the present interesting worm
-trail, the first of its kind from the Cre-taceous of Japan, in order that its usage
in biostratigraphy, paleoecology andtaxonomic classification be emphasized.
Genus Hitachia HATAI & NODA, n. gen.
Type species :-Hitachia nakaminatoensis HATAI and NODA, n. gen. et n. sp.
Type locality, formation, geological age,and depository:-Upper Part of the Hiraiso-machi Member of the Nakaminato
Formation exposed along the sea coast
at Hiraiso, Nakaminato City, lbaraki
Prefecture, Cretaceous. Preserved in thecollection of the Institute of Geology andPaleontology, Faculty of Science, Toho·
ku University (Reg. No. 91765).
Diagnosis :-Body rounded· elongate.
measuring .12-20 mm in minimum andmaximum widths, and 29 em in length,
segments well developed. strong, numbering 8 in a distance of 30 mm. Without longitudinal mesial depression or
podia-like structures.
Hitachia nakaminatoensis HA TAl
and NODA, n. sp.
Pl. 55

Description:-Worm trail impression
elongate, winding, 29 em in length; bodyrounded, thicker at middle part than atextremities, measuring 12 to 20 mm inminimum and maximum widths; seg·ments well developed, strong, number
8 within a distance of 30 mm, broader
than interspaces, indistinct at or near
extremities; external morphology areni·
coloid.
Remarks :-Comparison and affinity of

Explanation of Plate 55~Hitachia

nakaminatoensis HAT AI and NODA, natural size. Locality: Sea coast near Hiraiso,
Nakaminato City, Ibaraki Prefecture, Upper Cretaceous Hiraiso-machi Member of the Nakaminato Formation, IGPS coli. cat. no. 91765.

HAT A! and NODA:

K. KuMAG A I photo.

Upper Ctetaceous worm trail"Plate 55

!

608.

Upper C1·etaceous worm. trail

the present worm trail with other spe-cies was given above and here it is tobe said that its similarity with the spe-cies of the genus Arenicola suggeststhat it may have led a similar life. It
is judged that the fine-grained somewhat muddy sandstone of the upper
part of the Hiraiso-machi Member inwhich the present worm trail was found,
and where some ammonite and pelecypod
fossils have been found, was depositedunder a shallow water, open sea en·
vironment, and further it is inferredthat the water temperature was mo<lerate to warm.
It is also thought that the Hiraisomachi Member just mentioned which is:in conformable stratigraphic relation
with the underlying terrestrial Oarai
Formation, represents an early phase inmarine transgression and an environ·ment transitional from terrestrial to·shallow marine conditions.
References cited·CRIMES, T. P., and HARPER, J. C.,
1970.
Trace fossils. Geol. jour. Sp6c. Issue,
no. 3, 547 pp., See! House Press, Liverpool.
HowELL., B. F., in R. C. MooRE, 1962. Trea-

ChikkoHirai so
Isoai

461

tise on Invertebrate Palaeontology, PartW, Worms, p. 144-177, figs. 85-108.
KATTO, J., 1960. Some Problematica fromthe so-called unknown Mesozoic strata of
the southe~n part of Shikoku, Japan. Sci.
Rep. Tohoku Univ., 2nd Ser., Spec. Vol.,(HAI'ZAWA Memorial Volume), no. 4, p.
323-334, 2 pis.
MACSOTAY, 0., 1967. Huellas problematicas
y suvalor paleoecologico en Venezuela.
Geos. Escuela de Geologica, Minas y Metalurgiu, Univ. Vent. Venezuela, no. 16, p.
7-38, 18 pis.
OKADA, S., and liZUKA, K., 1949. Polychaeta
in lllustrated Encyclopedia of the Fauna
of Japan. Hokuryu-Kan, Pub/., 1339 pp.,3794 figs.
PERDIGAO, J. C., 1961.
Nereites do Baixo
Alentejo.
Comunicacoes dos Servicos
Geologicos de Portugal, tom. 45, p. 339363.
SAITO, T., 1958. Notes on some Cretaceous
fossils from the Nakaminato Formation,
Nakaminato City, lbaraki Prefecture,
Japan. Bull. Fac. Arts and Sci., lbaraki
Univ., Nat. Sci., no. 8, p. 83-94, 5 pls.
- - , 1959. Ditto, Part 2. Ibid., no. 9, p. 7985, 2 pis.
- - , 1961. The Upper Cretaceous System of
Ibaraki and Fukushima Prefectures, Japan. (Part 1). Ibid., no. 12, p. 103-144,
4 text-figs., 10 tabs.
- , 1962. Ditto, (Part 2). Ibid., no. 13, p.
51-87, 8 pis.

MurotoNakaminato
Oarai

~

p

»11 Ji!I ~

*

i')l;

~.

Trans. Proc. Palaeont, Soc. Japan, N. S .. No. 88, pp. 462-471, pl. 56 December 20, 1972

609.

AMMONITES FROM THE MESO,ZOIC Y AMABUFORMATION, KYUSHUMASAYUKlNODA
Wasada Junior High School, Oita

.7L.fflllHlllliM •€> Hii:U L.t.: 7 :/ ~ 7-1 1- 1::-::> "' '"C : JL~1j0) ~5C~fl;:lfli Jtp':JI.::J'll:f1-n, S!lli~iifl't~O)-·-::>-e ~ Q llJffil/lf';l:.: tt. ~ -c: ~~ifU/ltO) ~~{:ti*lt e: lj. t.t: ~ tt. ~ t:vi, Ul>t: rJ.7cO)~Ul;~J: IJ.::f!lii7J7:/~'}-1'J-il:/*~~ttf.:. --::>f';l: 3 - P ou'ti!i~!tl!~O)-'!J7'/7:/::a:ij'j7f-i" Q Berriasella "('. ftl!.i';l:J{ 7 :/-¥ :=. 7 :/ J:.ffil ::a:t.\'~-?tt Q Pset~doosterella l:tt·~ f...

-c

~QtO)~~~ • .:~~J:?-c~ma*~•~t~J:.ili!tl!O)•~·w•~·~•mw!tl!O)~

tLJIIliii::~Ht ~,? ~ .-.; ... 7;Y:I.::tt-~-::>?<imnts!\liJi~O):f-Ft:E-tQ.: c.v~ Tttim~•ftt..:. *~-c
l:t. :::.ttl? 7 ;/~ r1 ,. mO)iJcl\:1.::1JIJ.::t:c W$/IO)it!!JZ!P.itt~e-::>~t•-c:pp!t~ t... .f:'O)f!Jl.Wi:5.~¥Hit17J~l/Iic L.. '"C ~~II IJ 1.:: Q c c t lc.. :Jl!:aij. fil!ttll.llt.t: I? rJ.I.::.'I'llJill/!~ t ~U>'"C.
;:O)f-:rJ.17J~.lM~ .-.;.,. 7*S!li!~17J.t;!fi.I:OO-tQ IE~I¥Jr,mml::~tt..-c6atJg -~1.::-::>~t•'t'-'f-:rg~~~A-tQ,•mm~

and assigned the sequence to the Lower
Cretaceous Ryoseki Series.
Further·more, he pointed out that the forma·tions described above were all of bra·ckish water or non-marine environment.
Since then, MATSUMOTO (1954, 1967)
regarded this formation as the RyosekiGroup, and he and TERAOKA (1970) as·
signed it to the Kochian Series.
Anyhow, the Ryoseki Group thus un·derstood was assigned to the lowest partof the Cretaceous deposits of Japan, andapproximately correlated to the Wealdenof Europ3.
On December 29th, 1970. the author
unexpectedly collected several specimens of ammonites from the black shaleintercalated in the basal conglomerate,of which the one is referable to a species closely allied to Berriasella patulafrom the various localities of France.
There is furthermore another specie3.
This paper deals with the paleontolo·
gical descriptions of the two species ofIntroductoin
The Mesozoic Yamabu formation is
exposed in a narrow area of eastern
Kyushu, which belongs to the Chichibu
belt of the Outer Zone of Southwest
Japan. It lies unconformably on the
Paleozoic strata in the western part ofthe southern margin and is demarcatedby faults in the other sides, occupying
the area of about 6 km from east to
west and about 1.2 km from north tosouth. Its sequence is about 450 m inthickness.
The stratigraphy and the structure ofthis formation were previously reportedby FUJII (1954) and TERAOKA (1970).
FUJII considered that the basal to middle
member of the Yamabu formation might
be correlated to the Ryoseki formationof south Shikoku, and also to the Kawa·
guchi formation of the Yatsushiro area,
western Kyushu, for the reasons of theresemblance in fossils and lithofacies,
462

609.

463

Ammonites front Yamabu .formation
EXPLANAliON

shalt_./ laullsandston•fossil locality (animal)

co119lomoroto ()tos<il locality (plant)

2km

Text.fig. 1. Geological map of the Yamabu area, showing lithostratigraphic
division and distribution (adopted from TERAOKA, 1970).

ammonites from the Yamabu formation
with a remark on the geological age andalso some discussions on the problem ofJurassic-Cretaceous boundary in ] a pan.
Before going further, I express mysincere thanks to Professor Tatsuro
MATSUMOTO of Kyushu University for
his kind guidance and supervision ofthis study and also to Dr. Tadashi SA TO
of the University of Tokyo for his
useful suggestion about the identification of the species. Acknowledgements
are also due to Dr. Yoji TERAOKA of theGeological Survey of Japan for his kindness of permission to adopt his geological map in this paper.

Stratigraphy of the Yamabu formation

The Yamabu formation structurallyshows the syncline with axial trend fromeast to west. In its sequence four minor
sedimentary cycles are distinguished.
(1) Basal member. About 20 m-90 m
in thickness. Mainly exposed on thesouthern synclinal wing and partly inthe northern area, generally thickened
westwards and northwards. Its lower
part consists of ill-sorted conglomeratewith intercalation of black shale about
20 m in thickness near Shinkai, Honjo
Village, Minamiamabe County, Oita Pre-

Masayulci NODA

464

fecture, and generally becomingirich insandstone upwards. The upper partconsists of thin bedded and moderately
thickened alternation of sandstone andshale. Th'e··conglomerate is made up ofwell-rounded boulders, cobbles and pebbles of granite, granite-porphyry, aplite,
diorite, quartz-porphyry, porphyrite,
sandstone, slate, chert and green-schist,commonly with sandy matrix and partlywith muddy one. Sandstone is wellsorted and medium-grained. Thin bedded shale inserted in sandstone bears
carbonaceous or poorly preserved plantfossils.
'(2) Lower member. About 80 m" inthickness. Its lower part consists ofwhite or grayish white feldspathic-quartz-sandstone, well-bedded about 1 m inthickness. The main part consists ofhard, fine to medium-grained sandstone,occasionally inserted with thin layers of
shale, and gradually becoming morefine-grained upwards.

(3) Middle member. Ab.out 200m inthickness. Its lower part consists ofconglomerate about 10-15 m in thickness.
The main,·middle part is fine to mediumgrained sandstone and shale; the upper
part. sandy shale. The features of thisconglomerate is similar to those of thebasal one in grading, and kinds ofgravel but distinguishable in its more
muddy matrix. Occasionally '·it is sorich in matrix that it can be called thepebbly mudstone. In the main part, thefollowing brackish water mollusks of
the Ryoseki type are contained in theblack. shale inserted in feldspathicquartz-sandstone (FUJII, 1954; TERAOKA,
1970).
Brotiopsis kobayashii SUZUKI
Polymesoda (Isodomella) otsukai (Y ABE
and NAGAO)
Protocyprina cf. naumanni (NEUMA YR)
Eomiodon cf. sakawanus (KOBAYASHI
and SUZUKI)

...r
.0

EXPLANATION>-·r------

z

0

i=
<
~
a:
~

M

.0

>

~

'18 301•nal•

Elillillilll00N

sandston•

-l-.

:::>
CD

<

::L

~

_..
~
conglomfl'Qto

E

p

0

--+-------.0>

Paleozoic~

0

VB 102plant fossil bod

E___...
L_ _______
0

it '18 101xanimal fossil bod

Text-fig. 2. Schematic columnar section of the Yamabu formation.

609.
()strea465

Ammonites from Yamabu formationryosekiensis KOBAYASHI andSUZUKI

Aloides sp.
(4) Upper member. More than 100m
'in thickness, mainly occupying the axial:area of· the syncline.
Its lower part-consists of conglomerate. The mainpart is dark gray coarse-grained sandstone and thin bedded alternation ofsandstone and shale. Upper limit isuncertain becaus.e of the synclinal structure.
Sandstone is commonly graywacke with hard matrix of aphanitic
silica.
The geological map and stratigraphic--column of this formation are shown in·Text-figs. 1 and 2,

Paleontological descriptions

Family Berriasellidae SPATH, 1922
·Subfamily Berriasellinae SPATH, 1922
Genus Berriasella UHLIG, 1905

Berriasella sp. aff. B. patula SCHNEID
PL 56, Fig. la, b.

·Compare.1915. Berriasella patula ScHNEID, Ceo/. Palaeont. Abh. 13, (5), p. 66, pl. 7, fig. 51939. Berriasella patula,
MAZENOT, Mem.
Soc. Geol. France, N. S. N•. 41, pp. 60,
61, pl. 4, fig. 5a, 5b.
..1962. Berriasella sp. aff. patula, CoL.LIGNON,
Atlas des fossils Caracteristiques deMadagascar, p. 6, pl. 117, fig. 772

Material.-One specimen from toe.
YB101. the basal member of the Yamabu
formation, Oita Prefecture, Kyushu, collected by M.: NODA (1970) is concernedwith the description.
Descriptive remarks.-The specimen isso crushed secondarily that the original
proportion of breadth and height is uncertain. The umbilical wall is probably
steep, forming a rounded shoulder withthe flank. The whorl is fairly evolute,
probably less than one fourth of inner
whorl being embraced by the outer one,increasing comparatively slowly inheight. The original proportion of theumbilicus and the diameter is not precisely known owing to the effect of
deformation, but it is probably larger
than the measured value. The flanks
are rather flat. on which, especially on
the outer two whorls, the ornamentation
is fairly well-preserved.
On innerwhorls the ornamentation is unfortunately indiscernible because of the poor
preservation. In later stages more ~han
33 mm in diameter, the whorls are orna·
mented with sharp, distinct and some·what sigmoidal ribs. They begin at theumbilical margin with conspicuous elevation and bifurcate at the middle orat the point a little below the middleof the flank, curving considerably for·ward on the ventrolateral part. There
are 41 ribs in the last whorl which ismore than 116 mm in diameter. On thelast whorl. the ribs are as wide as interspaces and on the inner half of theflank, the interspaces are fairly wide

Measurements.Diameter

(1)
.(2)

Umbilicus

U!D

(1)

105.1 mm

41.4mm0.40(2)85.4mm

37.1mm

0.42

At the end of visible whorl.
About 65' back from (1).

466

Masayuki NODA
Table 1.
speciesmaterial

B. priversensis
B. priversensis
B. priversensis
B. sp. aff. B. priversensis
B. collistoides
B. callistoides
B. patulaB. patula

MAZE NOTMAZE NOTMAZE NOTMAZE NOTMAZE NOTMAZE NOTMAZE NOTScHNElD

present specimen

and fiat or broadly concave. The suturelines are indiscernible.
These characteristics are comparablewith those of Berriasella patula SCHNEID,
1915 from the Upper Tithonian of Neuburg and the Berriasian of Saint-Julienen Bochaine, Berrias and Bournet,
France. But there are some differences
between the specimens concerned with
the SCHNEID's original description (1915)
and the photograph of the same species
by MAZENOT (1939). That is, the MAZENOT's specimen has more rapidly enlarging whorls and a less wide umbilicus
than the SCHNEID's. The Yamabu specimen is rather closer to the MAZENOT'sin this respect, if the effect of deformation is taken into consideration.
The present specimen also resembles
B. priversensis (PICTET, 1867) and B.
callistoides (8EHRENDSEN, 1891), but differs from them in the following characters. (1) The umbilicus of the present
specimen as shown in Table 1, is fairly
wide in proportion to diameter. (2) The
ribs are flexuous in the present species
but nearly straight in B. priversensis.
(3) The ribs bifurcate near the middleof the flank in the present species but
at much higher point up about a fourthof the flank in B. priversensis.

pl. 2, fig. 3
pl. 2, fig. 4
pl. 2, ftg. 6
pl. 2, fig. 5
pl. 7, fig. 1
pl. 7. fig. 2
pl. 5, fig. 5
pl. 7, fig. 5

diameter umbilicusj

u;o

58mm
56mm
42mm
38mm
76mm
81 mm
73mm
74mm

2lmm
23mm
16mm
12mm
22mm
24mm
33mm
36mm

0.36
0.40
0.38
0.32
0.29
0.30
0.45
0.48

93. 5mm

38.8mm

0.42-

B. morti MAZENOT, 1939 from the·Upper Tithonian of Vogue, also resembles this specimen but differs from thelatter in more crowded ribs which bifurcate at somewhat higher point thanthe middle of the flank, and absence ofthe conspicuous elevation at the starting·point of the ribs.
Occurrence.-Locality YB101, at Yamabu, Honjo Village, Minamiamabe
Location:
County, Oita Prefecture.
Long. 131 "38'33"8E, Lat. 32.55'5"7N ..
The specimen occurred in the blackshale inserted in the basal conglomerate.
of the Yamabu formation.

Text-fig. 3.

Locality map.

609.

Ammonites from Yamabu formation

467~Family Oosterellidae BREISTROFFER, 1940is associated with Berriasella aff. patula.

Genus Pseudoosterella SPATH, 1924

On the geological age of theYam~bu formation

Pseudoosterella sp.
PI. 56, Figs. 2-4

Material.-Three specimens from toe.
YB101. the basal member of the Yamabu
formation, Oita Prefecture, Kyushu,·(coli. M. NODA) are concerned with~ the·description.
Descriptive remarks.-Although the·specimens are all· very incomplete fragments. the original. characteristics arepreserved in the convexity of whorl and-ornamentation. The specimen of NODA
collection JG. I-11003 is an adult indivi·dual about one sixth of whorl with 19.3mm in height and other two specimens·(JG. I-11004 and JG. I-11005) are both im~
mature, of which the one, JG. I-11004, isabout one fourth of whorl and the otheris about one sixth . of whorl.
.
The umbilical wall is fairly steep,
forming a round shoulder with the flank.
The whorl is compressed probably in~·Creasing slowly in height. The flank is'broadly convex and forms a subangularshoulder at the ventrolateral part. Thereis a finely serrate keel at the mid-venter.
The simple ribs bigin at the umbilicalmargin, running rectiradially on theflank. and sharply projected at the ventrolateral shoulder. They are as wide as
the concave interspaces, and regular in~strength and distance. The rib density
ranges from 8 to 9 per one sixth ofwhorl and 10 per one fourth of whorl.
In these features, this species is referable to Pseudoosterella. The specificidentification remains uncertain because-of the incomplete materials.
Occurrence.-Loca!ity, YB10l, at Ya·
mabu, Honjo Village, Minamiamabe·County, Oita Prefecture. The specimen

The Yamabu formation has been considered as the non-marine deposits. because of the occurrence of Protocyprina
cf. naumanni in its middle member. It has
been set to the lowest part of the Lower
Cretaceous in Japan, probable correlativeof the Wealden of England (see remarksin the introduction). The discovery ofammonites from the basal member.
however, indicates that the early stageof sedimentation of the Yamabu formation is probably of marine origin andthe occurrence of Berriase/la probably
Accordingindicates Berriasian age.
to MAZENOT (1939), Berriasella patula
occurs not only in the Lower Tithonianof Nefiburg (SCHNEID regarded the same
locality as the Upper Tithonian) but also
from the Berriasian of Saint-Julien-en·
Bohain, Berrias and Bournet of Southeast France. Although there are somequestions about the range of that species
described in MAZENOT, the occurrenceof· Berriasella aff. patula from the Yamabu formation. at least, indicates the.
geological age corresponding to theBerriasian.
SATO (1958, 1961) previously describedsome Berriasian and Valanginian ammonites from the Lower Cretaceous deposits which comformably overlie theJurassic strata in the Kitakami and the
Abukuma massifs. They are Berriasella
akiyamae SATO, Thurmanniceras isokusensis KOBAYASHI and FUKADA from the
Isokusa formation, Berriasella sp. ex gr.
B. berthei TOUCAS, Spiticeras (Spiticeras)
cf. binodige1· UHLIG. Olcostephanus sp.
and Kilianella juv. sp. from the Naga·saki formation and Parakilianella umazawensis SATO, Thurmanniceras sp. and468

Masayuki NODA

Berriasella sp. from the Koyamada formation. Based on the assemblage of
species he considered that there aresome marine deposits in Japan which
are contemporaneous with or somewhat
older than the non-marine Ryoseki Series
and that they probably correspond to
the Berriasian in the international scale,
regardless the occurrence of Olcostephanus sp., the genus from the Upper
Valanginian of Europe, South Africa,
Madagascar, Pakistan. Mexico and Peru,
being later than Berriasella in theseareas.
The basal member of the Yamabu
formation, therefore, may be correlated
to those formations of Northeast Japan.
But there is a question that Pseudoosterella sp. which should normally characterize the Upper Valanginian occurs
from the same bed with B. aft. patula.
This is similar to the case of the Nagasaki formation.
At present, it is uncertain whether or
not the association of the Berriasian andthe Valanginian species is the exceptional case found only in Japan. The
solution of this problem is importantfor more precise world-wide correlation.
The upper part of the middle memberof the Yamabu formation is characterized by brackish water mollusks listed
in p. 464, common in various localitiesof the Ryoseki Series in Japan. The
similar examples in the Lower Cretacous, i. e. the marine deposits under the
non-marine ones are also found in the Uminoura formation, Kumamoto Prefecture(TAMURA, 1960b; SATO, 1961 and MATSUMOTO. 1962) and the Kawaguchi formation, Kumamoto Pref. (MATSUMOTO, personal information, 1971). However, due
to the insufficient data, more precise
correlation of the Yamabu formationand those of Kumamoto Pref. is left
unsolved. I must investigate it as a,,__.,.

...... """"BERIIAS

-

L-o<

Up;tK

AHCllOCERATI DAE

- --

-,--

Vb'go.toaplncllna•

lo..

~:l!

Dctsofi'Qnilina•~

Vigcllll"'l•~A•ph:toc•rotll'lato

I

I

'~

CICIIII•d•l•nae
Gorni~trturallnae

- --

I

--

-

r--

i

follllna.
~----

---~

Splllootollnat

I~

Polyptrc:hllina•fOlc.otl~tphclnlna~e1-,&.rrloteiiJMe

I

HlmQICififlncJit_T_

NltOcmlllfi.Git

f----

I

r-

OOSlfAEUIDAE·t:: ?:;:.:

I''"""""""

z

'n
'::!

SiMOCII't'GtlM•

Hoploc:tralldcle
Clpp•IIICia.

!il

."

i=
1'ir'~1·m

DESWOC£JL\TAC£AE

I

Text-fig. 4. Range chart of ammonites at
Jr/Cr boundary (after WIEDMANN, 1967).

further problem.
The international problem of the

Jr/Cr b9undary
The geological age of the Yamabu•.
formation has some relations to the international problem of the Jr/Cr boundary, because there is no agreed opinion,.
at present, for the border-line by meansof paleontological evidence and sedimentary structure. The geological divisions of the Upper Jurassic and the Lower
Cretaceous are interpreted differently by
various researchers as shown in Table 2..
The PERGAMENT's division is based.
on paleontological evidence, notwith-·
standing it differs from that of HAUG,TOUCAS and WIEDMANN. He has regarded the zones of Subthurmannia boissieri and Riasanites rjasanensis as the·lowest limit of the Cretaceous. under the·name of Valanginian and he excluded
the name of Berriasian in his division.
In other words. the zones characterized.
by BerriastJlla spp. are included into the:609.
Table 2.

Ammonites from Yamabu formation

Geological division of the Upper Jurassic and theLower Cretaceous .(thickness not to scale).

.,

COOUAHDPICTET

1887

1871
toUAH
1910
MAZENOTBREIST~OFF.

CA.SE't
1953

Hl39

4Ll£N
1955

1964

PERGJ\MENT1965

TOUC4S1890

HAJJ()
1898

WIEDM.AHN
196~

BOSNARDO
HI 55

ValanQinlcn

1-:slt.

Wealdtn(CO...!.AIIE a.
~LLJi"SllU

Yalo.n.glnian

Bll'rriaskln(ODOl 115-~·

Ceooulllelf'/11

Infra
Yatanglnianc ~ ll!,t,lll ttl ~I

469

Upo"PDuris ton•~

Valc.nglnla.nVutonginlan$.,:Sit,S..Sfr,tl

Volanginl.ul)

SATO
HIS&

19U

Valanglnlo.n
Ryosei:l(URltl'fl

Valonglnla.n

c

"'

1917

MATSUMOtO

ValanginianW•1ld•n

u

i.

YABE

Kochic.nRyczanlan

B.rrlaslon

IIOGC!l.OIISIUIJ..s.lr.

111S)(WAfSUICI.Oit4ll

Purb.cldan(lfOIOIUtf

un1

Bds

hriaslan

UpJM-rTithonian
Tithonian

Lowe-r Purbtdl:(Ard•sdan)Upp•rVolglan(Lulworth hds)

J,.J.Ir

Tlthor.ion

Tithonian(Porllandlari crouc.u t.lto..J:s.l.
Volglan(~III.IWI 'NlJ

(Volglan)

Portlandian Portla.ndkln

r.tori~jlr~'fSOI(tOICQtN"'-ItM.

(TLthontan) (TIIhonoan)

Low.,(!M'I'nTU! I

Porlla.ndians. ~tr.

Tlthonlan

Lowu
Volglans. sl.~

TithonianTotlnosu(twi&Oilltol(Donuboan)(IIOUJUISO.I

I

Volgian. While, TOUCAS and others
included the two zones in the Tithonian,hence the top of the Tithonian in theirdefinition corresponds to the border-lineof 'Berriasian and Valanginian of MAZENOT. BUSNARDO, BREISTOROFFER and
KILIAN.
WIEDMANN's division is ba&ed on a
range chart of ammonites (Text-fig. 4).
For the Cretaceous of Japan the division proposed by YABE (1927) had beenused. He intended to use the term Ryoseki Series as a time-stratigraphic unit
but gave a definition that it is a series,
mostly of fresh and brackish water
deposits, which is the product of a timeof marine regresssion and that it is approximately correlated with the Wealden, the time of transition Jrom theJurassic Period to the Cretaceous in theEuropean standard. Actually, what had
been recognized as the Ryoseki in various areas of Japan was rather a facies·
stratigraphic unit and accordingly MA-

TSUMOTO (1943, 1954) proposed the Kochian ( =Kotian) for the lower series of
the tripartite Lower Cretaceous in Japan.
He attempted to define it by the available marine fossils (trigonians), but ourknowledge of the taxonomy and stratigraphic .occurrence of the trigonians atthat time was not. sufficient. SATO's(1958, 1961) discovery of ammonites fromthe lsokusa and the Nagasaki formations
has made it possible to correlate theJapanese scale with the international
one. From the fact that the ammonites
of Upper Valanginian aspect are associated with Berriasian species, it isbetter to ascribe these formations to theLower Cretaceous Berriasian as definedby MAZENOT and BUSNARDO rather than
to the Tithonian. The facts newly observed in the Yamabu formation also
support this.
The apparent intermingling of Valanginian aspect ammonites with the undoubtedly Berriassian ones at several

609.
Table 2.

Ammonites from Yamabu formation

Geological division of the Upper Jurassic and theLower Cretaceous .(thickness not to scale).

COQUA!IO--~

1111
KILIANPfCTET

18117

469

tltO

CASEY
1963

MAZENOT
IUQ

8R£1 ST AOFP'.
1964

AU£N
19115

PEAG.UCNT

1885

J

YABE
1U7

TOUC45

HAUG
1188

BUSIWIOO
1865

1110
WIEDioiANN

MA~SUI«lTO

1987,

SATO
1956

liU

Vq1a.noinianR)'oukl(r..a£ 1tvt

Volcnglnlon 1 - - - - + - - - - 1

(OU~

UHI

Low., Purbtdl(l.utwo.rth Beds )Tithonian(ArdneiCW'I)

Upp~r(for!~~ft lfOUC.AS ltM.I--i..-~---1

VololonJ,lfr

nthonlon

Porllcnd\Qn Potttan41Gn

lllhonlon

Yotgfan,..... "".'''(Vo\gian)r.torl-(O'o.o.rrli1GI

1---~(TIIhonlonJ (TIIhonlon)

l'tOICR'MUIIMI

Low«(~11UI

Portlandian.s.~lr,TithonianTorlnos-u

Lowor
Yal;lonTlth!M'an(Oorublun)ti'WU.OI. ' " ' '.l.Jir.

IIIOLUU1tOII

'

I
I
~

f·

'

Volgian. While, TOUGAS and others
included the two zones in the Tithonian,hence the top of the Tithonian in theirdefinition corresponds to the border-lineof 'Berriasian and Valanginian of MA·
ZENOT. BUSNARDO, BREISTOROFFER and
KILIAN.
WIEDMANN's division is ba~ed on a
range chart of ammonites (Text-fig. 4).
For the Cretaceous of Japan the division proposed by YABE (1927) had beenused. He intended to use the term Ryoseki Series as a time-stratigraphic unit
but gave a definition that it is a series,
mostly of fresh and brackish water
deposits, which is the product of a timeof marine regresssion and that it is ap·
proximately correlated with the Weal·den, the time of transition from theJurassic Period to the Cretaceous in theEuropean standard. Actually, what had
been recognized as the Ryoseki in various areas of Japan was rather a faciesstratigraphic unit and accordingly MA·

TSUMOTO (1943, 1954) proposed the Ko·
chian ( =Kotian) for the lower series of
the tripartite Lower Cretaceous in Japan.
He attempted to define it by the avail·able marine fossils (trigonians), but ourknowledge of the taxonomy and strati·
graphic .occurrence of the trigonians atthat time was not_ sufficient. SATO's(1958, 1961) discovery of ammonites fromthe Isokusa and the Nagasaki formations
has made it possible to correlate theJapanese scale with the international
one. From the fact that the ammonites
of Upper Valanginian aspect are associated with Berriasian species, it isbetter to ascribe these formations to theLower Cretaceous Berriasian as definedby MAZENOT and BUSNARDO rather than
to the Tithonian. The facts newly observed in the Yamabu formation als()
support this.
The apparent intermingling of Valanginian aspect ammonites with the un·doubtedly Berriassian ones -at several

Masayuki NODA

470

localities in Japan is unusual and left
to be solved in the future.
To sum up, the discovery of new data
in Japan as described in this paper, notonly enables us to attempt the world·
wide correlation but also may give some
contribution to the international problemof the Jr/Cr boundary.

Reference cited ·CASEY, R., 1963. The dawn of the Creta·
ceous Period in Britain. SE Union Sci.
Soc. Bull. no. 117, pp. 1-15.
Fuj(l, K., 1954. Stratigraphy and geologicalstructure of the Usuki area, Oita Prefecture, Kyushu (1). j~ur. Geol. Soc.
japan, vol. 60, pp. 413-427. (in Japanesewith English abstract).
KAMBE, N. and TERAOKA, Y., 1968. Geologyof the Usuki district. Quad. Ser. 1: 50000,
Fukuoka (14), no. 88, Geol. Surv. japan.
(in Japanese).
MATSUMOTO, T., 1943. Fundamentals in the
Cretaceous stratigraphy of Japan, Parts
II & III. Mem. Fac. Sci., Kyushu Univ.,
Ser. D (Geol.), vol. 2, pp. 97-237.
- - (ed.), 1954. The Cretaceous System inthe japanese Islands. Japan Sci. Promortion Soc., Tokyo. (for 1953).
- - and KANMERA, K., 1952. The lower
valley of the Kuma. Guidebook of thegeological excursions. Dep. Geol., Fac.
Sci., Kyush1t Univ. (in Japanese).
- - and - - , 1964. Hinagu, Kagoshima-49.
Explanatory text of the geological map ofjapan. 1: 50000, Geol. Surv. japan. (inJapanese).
MATSUMOTO, T., NooA, M. and M!YAHISA,
M., 1962. Regional Geology of Kyushu.
Asakura, Japan. (in Japanese).

MATSU~tOTO, T. and others, 1967.

Historicalgeology. Asakura, Japan. (in Japanese).
MAZE NOT, G., 1939. Les Palaeohoplitidae
Tithoniques et Berriasiens du Sud-Est
de Ia Fr~nce. Mem. Soc. Geol. France,N•. 41, pp. 1-303, pls. 1-40.
MOORE, R. C. (ed.), 1957. Treatise on. in·vertebrate paleontology, Mollusca 4. Geol.
Soc. America, Univ. Kansas Press.
PERGAMENT, M.A., 1965. Zonal scale of theCretaceous in the North-east· of Asia andits correlation with the American andEuropean scales. Report to the II I symposium on the development of oil and gas
resources in ECAFE countries, Tokyo.
1965.
SATO, T., 1958. Presence du Berriasien dans
Ia stratigraphie du plateau de Kitakami.
Boll. Soc. Geol. France, ser. 6, vol. 8, pp.
585-599, pl. 28.
- - , 1961. La limit Jurassico-Cretacee dans
Ia stratigraphie Japonaise. japan. jour.
Geol. Geogr., vol. 32, pp. 533-541, pl. 12.
- - , 1961. Faune Berriasienne et TithoniqueSuperieure Nouvellent Decouverte au
Japon. Ibid., pp. 543-551, pl. 13.
ScHNEID, T., 1915.
Die Ammoniten fauna
der obertithonischen Kalke von Neuburg
A. D. Geol. Palaeqnt. Abh., N. F. Bd. 13,
pp. 305-413, pis. 17-29.
TAMURA. M .. 1960a. A stratigraphic studyof the Torinosu group and its relatives.
Mem. Fac. Educ. Kumamoto Univ., vol. 8,
Supplement, pp. 1-40. (in Japanese).
- - , 1960b. Stratigraphical study on the Sakamoto group in Kyushu. four. Geol.
Soc. japan, vol. 66, pp. 371-383. (in Japanese with English abstract).
TERAOI<A, Y., 1970. Cretaceous formationsin the Onogawa Basin and its vicinity,
Kyushu, Southwest Japan. Rept. Geol.
Surv. japan, No. 237, pp. 1-87, pis. 1-18.
(in Japanese with English abstract).

Explanation of Plate 56
Fig. la, b. Berriasella sp. aff. B. patula ScHNEID, natural size. Loc. YBlOl, Shinkai, Honjo
Village, Minamiamabe County, Oita Prefecture. (Coil. NooA, 1970)
Figs. 2-4. Pseudoosterella sp. 2. natural size, 3, 4. x 1.5. Loc. YBlOl. (Coil. Noo·A, 1971) J<:NODA:

A1nmonites j?·om Yamabtt fonnation

Plate 56

609.

471

Ammonites from Yamabu formationJ., 1968.
Das Problem stratigraphischer Grenzziehung und die Jura/
Kreide-Grenze. E'cologae Geologicae Helvetiae, vol. 6l, pp. 321-386.

'WIEDMANN,

Abu kuma~iiJ ~AI!
Honjo
[!r.
lsokusa
1ii
Til
Kawaguchi)II
IJ
Kitakami;f~..!=.
Koyamada,j, LlJ 1B
Minamiamabe iliiliJllll*

Y ABE, H., 1927. Cretaceous stratigraphy of
the Japanese Islands Sci. Rep., Tohoku
Imp. Unilj., 2nd ser., vol. 11, pp. 27-100,
pis. 3-9. 'Nagasaki
RyosekiShinkai
Uminoura
YamabuYatsushiro

-Bi:
~

Uf
:fi

1m
i1il I iili
LlJ
all
i\
tt

tiT

Trans. Proc. Palaeont, Soc. Japan, N. S., No. 88, pp. 472-484, pl. 57, December 20, 1972

610.

SOME FOSSIL PTEROPODA FROM MIYAZAKI AND OKINAWA
PREFECTURES, SOUTHWEST JAPAN*
HIROSHI NODA
Institute of Geology and Paleontology, Faculty of
Science, Tohoku University

i!Yi~IFJ *i<i'kil! :tJ J:: D''f..iUf!~ITidll'~~f!i(t:fiiC-?It '"C : M'liii*£!Jrtlf¥j$!C~i!!T ~ J-.!JK4l¥r
1!!:.E.i-JliiiJlUt'i c r~:MUiiUt:t'frJ!~ :t~ .t tJ' }'fi~H~Iflf.fl\1~5Htl-i- ~ 4l/fllr~1Wfi!t~i~ifirfff10)JJIJ]iJ,'<J\~ilJi:fl',O)l}.'0E~fHt1i~~H L t:. IJ(.il;';~)IIJJiHi!l~il·l? C/eodora hataii, ffr!I!Jfi J:: F) Cleodora
okinawana, Cavolinia okinawana, Atlanta okinawana O)J!9lJifl1i~~tr 7 ~ 8l•T!O)RC!li£'d::ft~?t:,

firo~m

According to the authorities just mentioned the Pteropoda are cosmopolitan.
in distribution.
Fortunately the writer collected someMiocene and Pliocene species from Miyazaki and Okinawa Prefectures. Southwest Japan. These records are important for regional correlation and·reconstruction of the paleo-environmentof the deposits.

Introduction
There are only few studies on thefossil pteropods in and outside Japan,
probably because of their rare occurrence due to thin and fragile shell.
Although the Pteropoda range fromthe Paleogene (HARRIS, 1894; CURRY,1965) to Recent, in Japan the oldest
record is a Miocene species. Cavolina
raritatis NOMURA and ZINBO, 1935, fromthe Yanagawa Shell Bed in Fukushima
Prefecture and several Pleistocene species from the Semata Shell Bed by YAMAKAWA and ISHIKAWA (1912a, b), NoMURA and HATAI (1936) and KOBAYASHI(1956), whereas there are no records
from the Pliocene in Japan. The systematic classification of the shell bearing
pteropods has been based mainly on thesoft parts (RANG, 1825 ; PELSENEER,
1888b : TESCH, 1904; WENZ, 1960; TAYLOR
and SOHL, 1962; SPOEL, 1967; HYMAN,
1967; BOL TOBSKOY, 1971 and others).

Acknowledgments
The writer wishes to express his deep·
gratitude to Professor Kotora HATAI of
the Institute of Geology and Paleontology, Faculty of Science. Tohoku Uni··
versity, for his continuous encouragement and supervision during the present
study. Acknowledgments are due to·
Associate Professor Tamio KOTAKA,.
Tohoku University; Associate Prof. Tomohide NO!-IARA, Ryukyu University;:Drs. Taijiro KONISHI, Taisuke SUZUKI and
Kazuo MORI, Geological Survey of Japan.
for their kind information and sugges-·• Received June 20, 1972; read June 3,
1972, at Utsunomiya.

472

610.

Fossil Pteropoda, Miyazaki and Okinawations on the stratigraphy of Okinawa-ji·
rna. Thanks are also due to the membersof the Research Group of Cenozoic Mol·!uses (Profs. Koichiro MASUDA, Miyagi
University, Kotora HATAI, Tohoku Uni·
versity, Tokio SHIKAMA, Yokohama Uni·
versity, Shozo HAYASAKA, Kagoshima
University, Associate Professors Tamio
KOTAKA, Tohoku University, TakehikolWAI, Hirosaki Unive·rsity, Taisuke TA·KAYASU, Akita: University, Saburo KAN·
NO, Tokyo University of Education, Ki·
yotaka CHINZEI. Tokyo University, Hiro·yuki OTSUKA, Kagoshima University,
Junji ITOIGAWA, Nagoya University,
Sakae OIIARA, Chiba University, Drs.
Yasuhide IWASAKI, Tokyo University,
Kazuo OKAMOTO, Hiroshima University,
Messrs. Kimihiko OKI, Kagoshima Uni·
versity, and Kenshiro OGASAHARA, To·hoku University) for their kind informa·tion on the biostratigraphy of the Miya·zaki Basin and discussions in the field.
Thanks are expressed to Mr. Michio
KATO, Tohoku University for picking
up the fossil pteropods under the mic·roscope and to Mr. Kimiji KUMAGAI for
the photographic work.

Stratigraphic occurrence offossil Pteropoda
The fossil pteropods treated in thisarticle were collected from the two iso·lated areas (Miyazaki and Okinawa) as
described below.
A. Okinawa District.
The fossil pteropods were collectedfrom some localities in the area of dis·tribution of the Shinzato and Yonabaru
formations (Text-fig. 2) of the ShimajiriGroup (HANZAWA, 1935). The classifica·tion of the Neogene stratigraphy of the
Okinawa-jima proposed by HANZAWA(1935) and MACNEIL (1960). has recently

47~

been revised by KONISHI et al. (1970.
Accord·ing to them, the stratigraphical sequenceof the sou~h to central parts of Okina·
wa-jima is:1971) and FUKUDA et al. (1970).

Ryukyu Group

Naha Limestone

Chinen Sand \Kunigami Gravel
Shinzato Formation
ShimajiriGroup

Yonabaru Formation

Upper
Middle{LowerNaha Formation
Oroku Sand
Base unexposed - - - - -The lower part of the Yonabaru For·mation (FUKUDA et al.. 1969, 1970) is
nearly the same as the lower part of
the Yonabaru Clay of MACNEIL (1960)in the western part of island. The for·mation dips eastwards with low dipsand is composed mainly of massive darkgray to brownish gray clayey siltstonesometimes intercalated with fine-grained
tuffaceous sandstone layers which yiel·
ded Cavolinia telemus LINNAEUS, Cleodora okinawana n. sp., and Creseis acicula RANG. The first mentioned species.
is rather common but the other two arerather rare in occurrence. These pteropods are sometimes associated withmolluscs, foraminifers and fish otoliths.
The Shinzato Formation, first proposed by MACNEIL (1960) for the white
acidic tuffs intercalated in the massive
clayey siltstone around Shinzato, Sashi·ki·son in the eastern part of island.
under the name of the Shinzato Tuff
Member, also yielded fossil pteropods.
The formation was revised by FUKUDA
et al. (1970), and stated to be composed
mainly of massive dark gray to brownish gray siltstone intercalated withvery fine, layered acidic tuffs and partly-Hiroshi NODA

474

sandy siltstone. The formation is distributed mainly on the eastern side of
the island and in a small isolated area
<lf the Katsuren Peninsula (HAYAMI,
1971) and Miyagusuku-shima (Text-fig.!).
The Shinzato Formation near the type
locality yielded Cavolinia telemus. Cleodora okinawana and Creseis acicula. The
same formation around Yagena andTobaru, composed mainly of massive
tuffaceous sandy siltstone intercalatedwith white very fine acidic tuffs, yielded
Atlanta okinawana, Hyalocylix striata,
Cleodora okinawana and Diacrina bisulcata commonly. There is another isolated area around Yonagusuku where
the Shinzato Formation yielded somepteropods.
The Shinzato Formation, according to
FUKUDA et al. (1970) and the field survey
by the writer, changes gradually fromthe Y onabaru Clay (MACNEIL, 1960) andt

Yonabaru Formation (FUKUDA et al.,1970).
B. Miyazaki District.
Only one new species Cleodora hataii,n. sp. was found from the bluntly laminated pale gray to brownish gray, hard,
medium to fine-grained sandstone of the
Kawabaru Member. of the Koyu Formation (SHUTO, 1961). The fossil locality
is near to the localities nos. 46 and 47
of SHUTO (1961). According to SHUTO(1961), the Kawabaru Member occupies
the lowest part of the Koyu Formation.
The formation was subdivided by SHUTO(1961) into three members in his area of
Aoshima Facies. which is distributed
mainly along the western part of the
Miyazaki Basin. SHUTO (1961) statedthat the formation is Middle Miocene inage based upon the molluscan and plan·ktonic foraminiferal fossils. Cleodora
hataii is noteworthy for occupying alower stratigraphic position in Japan. An
allied species was described from thetype Langhian Cessole Formation (Italy)
by ROBBA (1971).

Remarks on the Pteropoda records

Text-fig. 1.

Map showing fossil localities.

As already stated by PELSENEER (1888b)
the Pteropoda differ from the Cephalopoda, Tentaculites, Conularia and Hyolithes though the external sculptures andmorphology are somewhat similar. PELSENEER {1888b), TESCH (1904), CHEN andB~ (1964), HYMAN (1967) and others havemade a historical review of the Pteropoda and further reworks are not necessary. At this place, remarks are
given on the first records from theMiyazaki and Okinawa Prefectures, southwest Japan because of their being
first from the Japanese Pliocene and· thesecond from the Miocene. As known atpresent the earliest record of the shell-

610.

Fossil Pteropoda, Miyazaki and Okinawa

bearing Pteropoda is from the Eoceneof England (HARRIS, 1894; CURRY, 1965).
The Koyu Formation in Miyazaki Prefecture is Middle Miocene in age according to SHUTO (1961), who studied themarine molluscan fossils from the formation. SHUTO (1961) subdivided the formation into three members, among which
the lowest or the Kawabaru Member
yielded Cleodora hataii, n. sp., a species
resembling Clio pulcherrina reported
from the Cessole Formation (Langhian).
Italy by ROBBA (1971). This Middle
Miocene record is the oldest in theJapanese Neogene.
The Shimajiri Group originally described by HANZAWA (1935) was revised
by MACNEIL in 1960. Recently, FUKUDA
et al. (1969, 70) published a geological mapof the Shimajiri Group distributed in
the central southern part of the Okinawajima from the view point of the natural
gas resources. The Yonabaru Formation. the middle part of the group. according to FUKUDA et al. (1969, 1970)
underlies the Shinzato Formation with
conformity and yielded Amussiopecten

praesignis, Hawaiarca uwaensis, Limopsis tajimae, L. tokaiensis, Glycymeris
pilsbryi and many other molluscan species besides Cavolinia telemus which was
originally described from the Mediterranean Sea and also reported from the
Miocene Bowden and Pliocene Manchioneal beds of Jamaica and Jacnel
Pliocene Formation in Tahiti. The geological age of the Yonabaru Formation
may be Upper Miocene from the macrofossils cited above. The Shinzato Formation yielded seven pteropod species
among which three are new species, andDiacrina bisulcata, Cavolinia telemus andCreseis acicula have been recorded from
the Middle Miocene Bowden of Jamaica,
Santa Rosa. Vera Cruz, Mexico andVirginia. All of the species mentioned

475

above are known to be distributed in
the subtropical to temperate seas asstated by PELSENEER (1888a, b), TESCH(1904), PILSBRY (1922) TOKIOKA (1955,
1960 in OKADA et al.), CHEN and BE(1964) and others. From the occurrenceof the pteropod species the Shinzato
Formation is inferred to have been deposited under an open sea condition but
whether of rather deep water analogous
to the depth of pteropod ooze may bequestionable. The correlation of separated -geological formations by thepteropod fossils is at present difficult
owing to the few occurrences, but should
their records be increased their value inbiostratigraphic correlation and paleobiogeographic analysis may become important.
~ystematic

description

Family Cavolinidae FISCHER, 1833
Genus Cleodora PERON and
LESUEUR, 1810

Cleodora hataii NonA. n. sp.
Pl. 57, figs. 9-11

Type locality :-Road side cliff, eastof the tunnel northeast of Yamaji, Saito
City, Miyazaki Prefecture, Kawabaru
Member of the Koyu Formation, Miocene. IGPS coli. cat. no. 92579.
Description .·-Shell very thin, rather
large for the genus, triangular, somewhat inequilateral in form. Ventral
side convex with a single broad, slightly
elevated median ridge with concentric
equi-spaced growth bands. Dorsal sideslightly convex with medium rib near
apex, branching into three longitudinalribs ; middle rib narrower than theothers with concentric growth lines.
Both sides of radial ridge on both sides

Text-fig. 2.

~~~~ ShinzatoE!'ormation

p: .::q Chinen Sand

8...

'\!EJ : Fossil locality

li?77:1;1 Naha Limestone. ~~') km(c)

Fossil localities of three separated areas (A; Naha district, B; Gushikami district,
Peninsula district). Geological map from FUKUDA et a/., 1970.

E:"SS'-';1 Naha Formation

•

Upper
}
MiddleYonabaru
Key BedFormation
Lower

buni

( 8)

C; Chinen

~

:to

~
1::::1

""·

~

0

~
"'i

-.::1

m

610.

Fossil Pteropoda, Miyazaki and Okinawa

477

Dimension:Length
Length

19.0 mm
20.6 mm

Width
Width

12.0mm
17.9mmHeightHeight

4.2 mm (92579)-mm (92580)

Iof shell become very fiat. Apex verysmall, pointed. Aperture wide, narrow,
middle part rather wide, both ends ofaperture narrowly fiat.
Comparison and affinities:- The present species resembles Cleodora bowdenensis COLLINS originally describedfrom the Middle Miocene of Jamaica(COLLINS, 1934) in having radial ridgeon the shell surface and in shell form
but differs ~rom the latter by the small
width for shell· length, distinct ·radial
ridges from near a'pex to aperture andrather distinct concentric growth lines
on the external surface. Cleodora balantium RANG illustrated by DALL (1878)
unqer the name of Balantinum recurvum
CHILDREN from the Recent sea of Australia resembles the present species in
the undulated ridged growth lines, andthree longitudinal ridges but the former
differs from the latter in the U -shaped
shell except for the sharp apical spine.
The latter species has a V-shaped trigonal shell form.
Clio pulcherrina(MAYER) described recently by ROBBA(1971) from the Langhian Cessole Formation in Italy resembles the present
new species in the shell form and undulated growth lines crossing the longitudinal ridges bu~ the former differsfrom the latter by having five longitudinal ridges.
Remarks:- At present, the present
species is the earliest record of Pteropoda in Japan.

Cleodora okinawana NODA, n. sp.
Pl. 57, figs. 6, 12, 13

Type locality :-Near the type localityof the Shinzato Formation, south cliffof Shinzato, Sashiki-son, Okinawa Prefecture, Shinzato Formation, Pliocene.
IGPS colt. cat. no. 92581.
Description :-Shell very fragile. moder·ate in size, and of lozenge-form. Dorsalside of shell ·sculptured with medium
rather narrowly elevated, longitudinal
rib which becomes divided into threesimilar longitudinal ribs. Interstices
flat, wider than longitudinal ribs
sculptured with very fine concentric
growth striations.
Aperture sharply
triangular in form. Lateral sides ofshell . very. thin rather fiat. Lateralcorner also sharp. Apical end prominentand spinose.
Dimension :-(Holotype) Height 12.8mm, width 6.0 mm.
Comparison and affinities:- The species
is characterized by its rather distinct
three longitudinal ridges on the apertural margin and its lozenge-form.
Cleodora bowdenensis COLLINS, 1934
originally described from the Middle
Miocene Lower Zone at Bowden in
Jamaica by COLLINS (1934), resembles thepresent new species but the former
differs from the latter in having straight,very prominent spinose apical end, andthree distinct radial ridges on the dorsalside of the shell. The latter species ischaracterized by its wider aperturallength, rather slight lateral elevation ofmedial ridge and recurved apex. Cleodora
pyramidata LINNAEUS (fide COLLINS, 1934,
p. 201) is another species allied with thepresent one but differs in having rather
inflated shell and distinct median ridge.
Cleodora hataii, n. sp. from the Miocene
Koyu Formation in Miyazaki Prefecture
resembles the present new species but

478

Hiroshi NODA

differs in the large size, inflated shell,
elevated medial part with three distinct
longitudinal ridges on the dorsal sideand distinct growth lines. Cleodora
australis o'ORBIGNY illustrated by DALL
(1878) resembles the present species in
the pyramidal shell form but the former
differs from the latter in having nointerstitial folds between the central
costae.
,Locality and formation :-Loc. no. 13,
prefectural road (Route 44) side cliff
near Kamima, Haebaru-son, Okinawa
Prefecture, Middle Part of Yonabaru
Formation, Miocene, IGPS col!. cat. no.
92582; Loc. no. 15, south of Shinzato,
Sashiki-son, Okinawa Prefecture, Shinzato Formation, Pliocene. IGPS coil.
cat. nos. 92581, 92582 ; Loc. no. 12-06, seaside cliff, west of Tobaru, Yonagusukuson, Okinawa Prefecture, Shinzato
Formation, Pliocene, IGPS coil. cat. no.
92596.
Genus Hyalocylix H. FOLKSKAL, 1875
Hyalocylix striata (RANG, 1828)
Pl. 57, figs. 7, 8

Creseis striata RANG, 1828. Ann. Sci. Nat.,Tom. 13, p. 315, pl. 17, fig. 7 (fide LAMARCK, 1836).
Cleodora striata RANG: LAMARCK, 1836, Hist.
Nat. Animaux, Verteb .. Tom. 7, p. 433.
Creseis striata RANG: DALL, 1878, pl. 5, figs.
30a, b. (non vidi).
Clio (Hyalocylix) striata (RANG) : PELSENEER,
1888a, Rept. Sci., Voy. H.M.S. ChalleHgel",vol. 23, p. 54, pl. 2, fig. 3.
Clio (Hyalocy/ix) striata (RANG) :TESCH, 1904,
Siboga Exped. Monogr., vol. 52, p. 27, 28,
pl. 1, figs. 16-17.
Hyalocy/ix striata (RANG) : TOK!OKA, 1955,
Publ. Seto Mar. Bioi. Lab., vol. 5, no. 1,
p. 65, pl. 9, fig. 21.
Hyalocylix striata (RANG) : TOK!OKA in OKA·
DA et al .. 1960, Encyclop. Zoo/. lllust.

Col .. Vcl. 3, p. 171, fig. 642.
Hyalocylix striata (RANG) : CHEr>: and BE~
1964, Bull. Mar. Sci. Gulf; Carrib., vol.
14, no. 2, p. 193, fig. 3g.
Hyalocylis striata (RANG): KEEl\, 1971, p ..
805, fig. 2285.

Description :-Shell very thin, fragile,.
small, conical tube, nearly straight but
slightly curved at end of apex. Aperture round in cross section. Apex
narrow and rather spinose. Shell surface
sculptured with equispaced distinct
annulations.
Dimension :-Length 6.2mm and width
2.5mm.
Comparison and affinities:- The present species is characterized by its
slightly curved apex and distinct externalannulations.
This species resembles
Hyalocylix haitensis COLLINS, 1934 inhaving similar external annulations but
the former has finer and denser crenuJations on the external surface and theapical end is more spinose, comparedwith the latter.
Remarks :-The present species, known
as a Recent form of wide distribution
in the warm to subtropical sea realm,
has no previous record as fossil. The
present discovery extends the geologicalrange of this species back to the Pliocene.
Locality and formation :-Loc. no. 1206, sea side cliff, west of Tobaru,
Miyagusuku-shima,
Yonagusuku-son,
Okinawa Prefecture, Shinzato Formation,
Pliocene, IGPS coil. cat. no. 92583.

Genus Diacria GRAY, 1847
Diacria bisulcata GABB, 1873
Pl. 57, fig. lB.

Diac1·ia bisnlcata GABB, 1873, Trans. Amer.
Phi los. Soc., n.s., vol. 15, p. 200. (non
vidi)

610.

Fossil Pteropoda, Miyazaki and Okinawa

CavoliiiQ (Diacria) bisulcata (GABB) : DALL,
1892, Trans. Wagner lnsl. Philad .. vol.
3, pt. 2, p. 430. (non vidi).
Diacria bisulcata GABB: PILSBRY, 1922, Proc.
Acad. Nat. Sci., Phila, vol. 73, p. 309,
fig. 4.
Diacria bisulcata GABB: WooDRING, 1928,
Carnegie Inst. Washington, no. 385, p. 116,
pl. 1, figs. 14-17.
Diacrina bisulcata (GABB) : CoLLINS, 1934,
john Hopkins Univ. Geol., no. 11, p.
197-200, pl. 10, figs. 1-3, pl. 9, figs. 13-

479

by WOODRING (1928, pl. 1, figs. 16-17)
and COLLINS (1934, pl. 10, figs. 1-3) re·semble Diacrina trispinosa s. s. by pos·sessing five longitudinal' ridges on thedorsal surface.
For this reason thespecies group should be re-examined.
Locality and formation :-Loc. no. 1206, sea side cliff, west of Tobaru,
Miyagusuku-shima,
Yonagusuku-son.
Okinawa Prefecture, Shinzato Formation,
Pliocene, IGPS coil. cat. no. 92584.

20.

Description :-Shell small. equilateral,
somewhat lozenge in form. Aperturesmoothly rounded, both sides broadly
angulated, its external ends rather sharp,
not spinose, apical area narrow and long.
External sculpture with slightly elevatedlongitudinal ridge (medial) and secondary
ridges, outer sides become fiat, without
striations beside growth lines. Ventral
side unexamined at present.
Height
4.8 mm and width 3.8 mm.
Comparison and affinities :-The pre·sent species, only one shell showing thedorsal side, resembles Diacria mbaensis
LADD originally described by LADD (1934)
from the Miocene Suva Formation, Fiji
Island in the external shell form but thelatter is characterized by five subequalrounded longitudinal ribs on the dorsal
surface, a character different from thepresent species.
Diacrina trispinosa(BLAINV!LLE) illustrated by KURODA,
HABE and OYAMA (1971) from Sagami
Bay, Kanagawa Prefecture (1971 in Bioi.
Lab. Imp. Household) resembles the pre·sent species by having rounded aperture,
sharply spinose ends and longitudinalridges on the dorsal surface but thepresent species differs from the former
by its low apertural margin, narrow
concaved sides and distinct longitudinalridges on the dorsal surface. Some
specimens identified as Diacrina bisulcata

Genus Cavolinia ABILDGAAD, 1791

Cavolinia telemus LINNAEUS, 1758
Pl. 57, figs. 14-17, 19-20, 22-26, 29-31

Cavolinia telemus LINNAEUS, 1758, Syst. Nat.
ed. 10, p. 365, (fide CoLLINS, 1934).
Cavolina telemus (LINN~) : WooDRING, 1928,
Carnegie /nst. Washington, no. 385, p. 10,
p. 113, pl. 1, figs. 6-7.
Cavolina telemus (LINNE) : CoLLINS, 1934,
john Hopkins Univ., Stud. Geol., no. 11,
p. 188-190, pl. 8, figs. 4-7.
Cavolina (Cavolina) telemus (LINNE) : WENZ,
1959, p. 52, fig. 174.
Cavolinia telemus (LINNE) : KuRODA, HABE
and OYAMA in Biol. Lab. Imp. Household,p. 300, pl. 64, fig. 12.

Description :-Shellratherlarge,
fragile, inequivalve and equilateral.
Ventral valve much swollen, particularly
on apertural margin. External surface
rather smooth, with very fine concentricstriations conspicuous near aperture andslightly bent backward medially. Dorsal
valve rather flat. external surface sculptured with three distinct elevated longitudinal ribs; one of them distinct andsituated on middle part of shell, othertwo slightly weaker. Apertural margin
with elevated rim, overhanging ventral
part of shell, rather smoothly rounded.
Posterior horn spinose, lateral ends notspinose but shouldered. Best preserved

480· Hiroshi NODA

and largest specimen is 12.9 mm in lengthand 10.5 mm in width.
Comparison and affinities:- The pre·sent species resembles Cavolinia globulosa(GRAY) and Cavolinia uncinata (RANG) in
having similar shell form and growthlines but the latter two differ from thepresent species by their .short andslightly incurved posterior horn andwide apertural~side. Cavolinia tridentata_which was described from the Semat"'
Pleistocene Formation by Y AMAKAWA
and ISHIKAWA (1912b) may be identified
with the present species in being charac·terize~ by one medial longitudinal ridgewith two on the dorsal shell, the shellform and posterior and lateral ends.
Remarks :-The present species is. very
common in tuffaceous medium to finegrained sandstone and tuffaceous siltstone of the Shinzato Formation. The
species has been discussed in detail by
COLLINS .(1934) who listed its many syno·
nyms. The generic names Cavolinia andCavolina were treated by PELSENEER(1888a) who used Cavolinia in sa·me sense
as Cavolina. Many previous authorities
used the generic name Cavolina butrecently SPOEL (1967) and KURODA, HABE
and OYAMA (1971) used Cavolinia for
Cavolina. The nomenclature of thegeneric name should be subjected tofurther consideration.
Locality and jormati011 :--'-Loc. no. 42,
east of
Nesabe,Tomigusuku-son,
Okinawa Prefecture, lower part ofYonabaru Formation, Miocene, IGPS,
coil. cat. no. 92585 : Loc. no. 15, southof Shinzato, Sashiki-son, Okinawa Pre·
fecture, Shinzato Formation, Pliocene,
IGPS, coli. cat. no. 92586 ; Loc. no. 123,
small road side cliff, north of Hanagu·
suku, Gushikami-son, Okinawa Prefec·
ture, Shinzato Formation, Pliocene, IGPS,
coli. cat. hot. 92587; Loc. no. 126, smallroad side cliff, north of Hanagusuku,

Gushikami-son, Okinawa . Prefecture,
Shinzato Formation, Pliocene, IGPS, coiLcat. no. 92588; Loc. no. 129, west sidecliff of Route 46 (prefectural road),
north of Gushikami, Gushikami-son,
Okinawa Prefecture, Shinzato Fo-rmation,
Pliocene, IGPS, coli. cat. no. 92589; Loc.
no. 12-06. sea side cliff, we~t of Tobaru,
Miyagusuku'shima,
Yonagusuku-soh,Okinawa Prefecture, Shinzato Formation,
Pliocene, IGPS, coli. cat. no. 92590.
Cavolinia okinawana NODA, n. sp.
Pl. 57_, fi_gs. 27-28

Type locality :-Loc. no. 12-06, sea si.decliff, west of Tobaru, Miyagusuku·shima,
Yonagusuku-son, Okinawa Prefecture,
Shinzato Formation, Pliocene.
Description :-Shell fragile, rather smallin size, subrounded, _inequiv'alve, equilaleral in shell form. Ventral shellinflated, external surface smooth, covered
with dorsal apertufe. Dorsal valve notso swollen, sculptured with faint, ele·
vated radial fold on apertural marginand indistinct on middle to posterior
side., Both sides. narrowly depressedwith small elevated ridge. Aperture
narrow, quadrate, apertural plate obli·
quely positioned with smooth surface ..
Lateral sides quadrate but not spinose.
Posterior end small and spinose.
Dimension :-Height 5.9 mm and width
5.9mm.
Compa,·ison and affinities:- The pre·sent species resembles Cavolinia tridentata (FORKSKAL) . which is characterized
by faint radial ribs on the dorsal surface
but differs from the latter in having
narrow radial depressed area, and smal!
elevated ridge on the lateral side, andfaint radial fold at the apertural margin
on dorsal surface. Cavolinia telemus
LINNAEUS is considered a synonym of610.

Fossil Pteropoda, Miyazaki and Okinawa

Cavolinia tridentata by KURODA, HABE
:and OYAMA (in Bioi. Lab. Imp. House·
hold, 1971) but the present writer con··siders that both species are valid be·cause the characteristics of the dorsal
surface are different from each other.as mentioned by TOKIOKA (1960) and
YAMAKAWA and ISHIKAWA (1912b). Cavolinia telemus differs from the present·species in having only one mid-radial
ridge on the apertural margin and nar··row apertural shell band.
Locality and formation:- Loc. no. 12·06, sea side cliff, west of Tobaru, Miya_gusuku·shima, Yonagusuku-son, Okinawa Prefecture, Shinzato Formation, Plioeene, IGPS coil. cat. no. 92591.

1

Genus Creseis RANG, 1828
Creseis acicula (RANG, 1828)
Pl. 57, figs. 1-5.
·Clio acicula RANG, 1828, Ann. Sci. Nat. Ser.,vol. 13, p. 318, pl. 17, fig. 6, (fide PEL·SENEER, 1888a)·Clio acicula RANG : PELSENEER, 1888a, C/rallenger Rep., vol. 23, p. 51-53.
··Clio (Creseis) acicula RANG: YAMAKAWA
and IsHIKAWA, 1912a, jour. Geol. Soc.
Tokyo, vol. 19, p. 2-3, pl. 1, figs. la-b.
-Creseis acicula (RANG) : CoLLINS, 1934, john
Hopkins Univ. Geol.. no. 11, · p. 207-208,
pl. 9, figs. 6-7, pl. 13, figs. 7-8.
-Creseis acicu{a (RANG): ABBOTT, 1954, p.
294, fig. 64-n.
·Creseis acicu{a (RANG) : TOKIOI<A, 1955, Seto
Mar. Bioi. Lab., Publ., vol. 5, p. 64.
-Crr1seis acicula (RANG) : TOKIOKA, 1960, in
OKADA et a{., p. 170, fig. 1.
·.Creseis acicula (RANG) : CHEN and B€:, 1964,
Bull. Mar. Sci., Gulf Carrib., vol. 14,
no. 2, p. 216, fig. 3-e.
·Creseis acicula (RANG) : KEEN, 1971, p. 805,
fig. 2281.

Description :-Shell very small, long481

tuberous in form and circular in crosssectiol).. Aperturai part widest, posterior
end spinose put not curved. Shell sur·face smooth but with transverse stria·
tion.
Comparison and affinity:-The present
species resembles Creseis virgula RANG .
Both Creseis acicula and C. virgula are
subdivided into subspecies, the former
into two and the latter into four sub·species according to the morphologicai
features of the posterior end and trans·verse striation (s). The present species
differs from the Creseis virgula groupin having straight. posterior end without
expansion near the posterior e'nd likea knob. A subspecies, Creseis acicula
clava RANG, has a shorter tuberous shell
but resembles the· species.
Locality and formation :-Loc. no. 12,
west of Y agena Harbour, Y agena, Yo·nagusuku-son, Okinawa Prefecture, Shi·nzato Formation, Pliocene,· IGPS,' coil.
cat. no. 92592; Loc.' no. "15, south ofShinzato, Sashiki-son, Okinawa Pre·
fecture, Shinzato Formation, Pliocene,
IGPS, coil. cat. no. 92593.

Family Atlantidae
Genus Atlanta LESUEUR, 1817
Atlanta okinawana NODA, n. sp.
Pl. 57, fig. 21

Type locality :-Loc. no. 12-06, seaside cliff, west of Tobaru Miyagusukushima, Yonagusuku-son, Okinawa Prefecture, Shinzato Formation, PI'iocene,
IGPS, coil. cat. no. 92594. ·Description :-Shell small, fragile andplainly spiral. Three spirals with very
thin, fragile plate on outer margin ofshell, distinct on outer whorl and fainton inner. Shell surface smooth except

482

Hiroshi NODA

for fine growth lines. Shell characterized by similar sculpture and height.
Aperture elongated fusiform, apertural
slit not observable.
Dimension:-Width 3.4 mm and thickness 0.3 mm.
Comparison and affinities:-The pre·sent species is characterized by its flatshell with thin marginal plate without
apertural slit. Atlanta Lesueur SOULEYET
resembles the present new species inouter view of the shell but the former
has the apertural slit not so deep.
Atlanta Peroni LESUEUR differs from thepresent sp_ecies in having distinct keelon the marginal whorl".
TOK!OKA (1960, in OKADA et al.) illustrated some species allied to the present
species. Among them, Atlanta lesueriof TOKIOKA resembles the new species
except for the apertural slit. Atlanta
gaudichaudi SOULEYET is another allied
species in having low-spiral whorls with
marginal keel but has a relatively higher
shell compared with the present new
species. The present species resembles
the American fossil species Atlanta
diamesa WOODRING originally describedfrom the Miocene Bowden Formation(WOODRING, 1928, p. 133, pl. 2, figs. 2325) in having similar number of coiled
whorls and marginal keel, but the former
differs from the latter in having low·
elongated profile of apertural form
(latter one is trigonal) and the keel
appears from inside of the suture.
Remarks :-Only two specimens of thisspecies were collected from the Shinzato
Formation. This is the first record of
the genus as fossil.
Locality and formation :-Loc. no. 1206, sea side cliff, west of Tobaru, Miya·gusuku·shima, Yonagusuku ·son, Okinawa
Prefecture, Shinzato Formation, Pliocene,
IGPS, coli. cat. no. 92594.

References cited
ABBOTT, R. T., 1954, American seashells_
D. Van Nostrand Camp. Inc., Princeton_
541 pp., 40 pis., 97 figs.
Biological Laboratory, Imperial Household,.
1971, The sea shells of Sagami Bay.:
Maruzen Co., Tokyo. p. 1-489 (in English)~p. 1-741 (in Japanese), pis. 1-120.
BOL.TOVSI<OY, D., 1971, Pteropods thecosomados del Atlantica sudocidental. Malacologia, vol. 11, no. 1, p. 121-140, pl. 1.
fig. 1, 1 table.
CHEN, C., and B~. A. W., 1964, Seasonal
distributions of euthecosomatous pteropods in the surface waters of five stations in the western north Atlantic.
Bull. Mar. Sci., Gulf and Carribean, vol·.
14, no. 2, p. 185-220, figs. 1-12.
CoL.L.INS, R. L., 1934, A monograph of theAmerican Tertiary pteropod mollusks_
john Hopkins. Univ. Studies in Geology,.
no. 11, p. 137-234, pis. 7-14.
CuRRY, D., 1965, The English Paleogene
pteropods.
Proc. Malac. Soc. London~
vol. 36, pt. 6, p. 357-371, pis. 16-17, figs.
1-20.
FUKUDA, 0., et al., 1969,
Natural gas re··sources of the Ryukyu Islands-Preliminary report by the 4th phase survey
team of G. S. ].-Bull. Geol. Surv. japan,.
vol. 20, no. 2, p. 101-124, figs. 1-24.
- - , et al., 1970, Natural gas resources of·
the Ryukyu Islands-Preliminary reportby the 5th phase survey team of G. S. J ..
-. Ibid., vol. 21, no. 11, p. 627-672, figs.
l-30, tabs. 1-20.
HANZAWA, S., 1935, Topography and geology of the Ryukyu Islands. Sci. Rept.,.
Tohoku Imp. Univ., 2nd Ser., vol. 17, P1-61, pis. 1-15, figs. 1-7, chart 1, maps.
1-5.
HAI~RIS, G. F., 1894,
On the discovery of apteropod in British Eocene strata, with.
a description of a new species. Proc.
Malac. Soc. London, vol. 1, p. 61-62.
HAYAMI, T., 1971, Some Neogene Cheilosto-~mata (Bryozoa) from Okinawa-jima.
Trans. Proc. Palaeont. Soc. japa11, N.S.,.
no. 82, p. 73-92. pis. 10-12, figs. 1-2.

610.

483

Fossil Pte·ropoda, Miyazaki and Okinawa

LHYMA:-1, L. H .. 1967, The invertebrates. vol.
1. Mollusca I. MacGratu Hill Book Co.,
N.Y. 792 pp .. 249 figs.
KEEN, A.M., 1971, Sea shells of tropicalwest America. Stanford Univ.. Press ..
Stanford. 1064 pp., 22 pis.
KoBAYASHI, T .. 1956, A paleo-meteorologicalinterpretation to the occurrence of the
Argonautinae in province Kaga, central
Japan. japan. jour. Geol. Geogr., vol.
27, nos. 2-4, p. 93-104, pl. 6, figs. a-d.
KoNISHI, T., et a{., 1970,
Reports on thedevelopment of underground water re·sources. nos. 1-3, Geol. Surv. japan, 77
pp., 27 figs.
- - . et al., 1971, Ditto .. no. 4, Geol. Surv.
japan, 70 pp., 15, figs. 9 tabs.
LAI)D, H. S., 1934, Geology of Viti levu, Fiji.
Bernice P. Bishop Mus., Bull .. 119, p. 1263, pis. 1-44, figs. 1-11, tabs. 1-7.
LAMARCK, Par ]. B. P. A., 1836,
Historie
Naturelle des Animaux sans Vertebres.
Tom. 7, p. 1-735.
MAcNEIL, F. S., 1960, Tertiary and Quaternary Gastropoda of Okinawa. U.S. Geol.
Surv. Prof. Paper, 339, p. 1-148, pis. 121, figs. 1-17.
NoMURA, S. and ZINBO, N., 1935, Mollusca
from the Yanagawa shell-beds in the
Hukusima Basin, Northeast Honshu,
Japan. Saito Ho-on Kai Mus., Res. Bull.,no. 6, p. 151-192, pl. 15.
NoMURA. S. and HATAI, K., 1937, Somefossil marine invertebrates from the
Semata shell-beds. Semata, Itihara-gun,Kazusa Province, Central Japan. Bull.
Biogeogr. Soc. japa11, vol. 7, no. 6, p.
63-66, figs. 1-5.
PELSE:-IEER, P., 1888a, Report on Pteropoda
collected by H. M.S. Challenger during
the years 1893-1876, II. Thecosomata.
Rept. Sci. Result, Voyage H.M.S. Challenger, Zool., vol. 23, pt. 65, p. 1-132,
pis. 1-2.
- - . 1888b, Ditto, III Anatomy. Ibid., vol.
23, pt. 66, p. 1-97, pls. 1-5.
Pn.sBRY, H. A .. 1922, A revision of W. M.
GABB's Tertiary Mollusca of Santa Domingo. Proc. Acad. Nat. Sci. Phi/ad.,vol. 73, pt 2, p. 305-435, pis. 16-47, figs.
1-48.
RANG, S., 1825, Description de deux generes
nouveaux ~ppartenant Ia classe des pteropodes. Ann. d. Sci. Nat., Tom. 13,(non vidi).
REEVE, L., 1878, Monograph of the genus
Pteropoda. Conch. Cab., vol. 20. pis. 1-6.
ROBBA, E., 1971, Associazioni a pteropodi
della formazione di Cessole (Langhiano).
Riv. /tal. Paleont., vol. 77, no. 1, p. 19126, pls. 1-5, fig. l-5.
SPOEL, S. van der, 1.967, Eutheocosomata.
A group with remarkable development
stages. (Gastropoda- Pteropoda).
Zoo/.
Mus., Amsterdam, ]. Noorduijin en Zoo11,
N. V., p. 1-376. (fide Ov AMA, K., 1970,
Venus, vol. 29, no. 1, p. 12).
TAYLOR, D. and SoHL, N., 1962, An outlineof gastropod classification. Malacologia,
vol. 1, no. 1, p. 7-32.
TESCH, ]. J .• 1904, The Thecosomata andGymnosomata of the Siboga Expedition.
Siboga Exped. Mo11ogr., vol. 52, p. 1-92,
pis. 1-6.
Toi<IOI<A, T .. 1955, On some planktonic animals collected by Syuntoku-maru in MayJune, 1954. IV. Thecosomatous pteropods. Pub/. Seto Mar. Bioi. Lab., vol.
5, no. 1, p. 59-74.
- - . 1960, (in OKADA. K., et al.), Encyclopedia Zoologica illustrated in colours,
vol. III. HollUryu-Kan, Pub/. Co., 200 pp.,pis. 1-91.
WENZ, W., 1959, Gastropoda, Teil 2, Euthyneura. Gebr. Born .• 834 pp., 2515 figs.
WOODRING, W. P., 1928, Miocene mollusks
from Bowden, Jamaica. Pt. 2, Gastropods and discussion of results. Carnegie/nst. Washington, Pub/. no. 385, p. 1-564,
pis. 1-40, figs. 1-3.
YAMAKAWA, G. and IsHIKAWA, M., 1912a,
Some pteropods from the Neogene of
Semata. jour. Geol. Soc. Tokyo. vol. 19,
no. 222, p. 1-8, pl. 1.
- - and - - , 1912b, Some pteropods fromthe Neogene of Semata. Ibid., vol. 19,
no. 223, p. 17-24, pl. 6.

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Explanation of Plate 57
Figs. 1-5, Creseis acicula RAr.;G, x 20, Loc. no. 15, Pliocene Shinzato Formation, IGPS coiLcat. no. 92593.
Figs. 6, 12, 13, Cleodora okinawana NOD A, n. sp., x 3, figs. 6, 12, Loc. 15, IGPS·, coli. cat:nos. 92582, and 92581, Fig. 13, Loc. 12-06, Pliocene Shinzato Formation, IGPS coli. cat.
no. 92596.
Figs. 7, 8, Hyalocylix striata (RANG), x 5, Loc. no. 12-06, Pliocene Shinzato Formation, IGPS
coli. cat. no. 92583.
Figs. 9-11, Cleodora hataii NooA, n. sp., x2, Miocene Kawabaru Member of the Koyu Formation, IGPS coli. cat. no. 92579.
Figs. 14-17, 19-20, 22-26, 29-31, Cavolinia telemus LINNAEUS, Figs. 14-16, 19--20, Loc. 123, x 2,.
IGPS coli. cat. no. 92587; fig. 17, Loc. 129, x3. IGPS coli. cat. no. 92589,; fig. 22, Loc.
123, x 2, IGPS coli. cat. no. 92587; fig. 24, Loc. 123, X 3, IGPS coli. cat. no. 92789; fig. 23,
x2, Loc. ~5. IGPS coli. cat. no. 92586; figs. 25, 26, x3, fig.27, x5, fig.30-31, x3,allfronlc
Loc. 12-06, IGPS colt. cat. no. 92590; fig. 29, Loc. no. 126, IGPS colt. cat. no. 92588. All,.
Pliocene Shinzato Formation.
Fig. 18, Diacrina bisulcata GABB, x 5, Loc. 1206, Pliocene Shinzato Formation, IGPS coli. cat ..
no. 92584.
Fig. 21, Atlanta okinawana NooA, n. sp., x 8, Loc. 12-06, Pliocene Shinzato Formation, IGPS:
coil. cat. no. 92594.
Fig. 28, Cavolinia okinawana NoDA, n. sp., x3, Loc. 12-06, Plioc~ne Shinzato Formation, IGPS
coli. cat. no. 92591.

NODA.:

Fossil Ptero1;oda, Miyazaki and Okinawa

Plate 57

3

7

5

8

13
11

10

14

JS

16

17

18

19

20

21

24

28

25

26

2\J

30

23

22

27

31

PROCEEDINGS OF THE PALAEONTOLOGICAL SOCIETY
OF JAPAN

r-1*i!ig:_:f'/1J~~m no IID"J~r:t. 1972 ~ 10 FJ
28 13 (±) ~~:k~Illl~§l\l;:::t;lt•"(f}~fil!~t'l.t.:.
<~1J!J~ so ,g). flf!~ 29 a <s) 1::1:t. racQ)i~ ~Q))lli~~fi-?t.:, (#1m~ gs,g).

1;'f Jlll ff1i i1il
Paleontologic potpourri of the Pacific Coastof North America ........ DANNER, W:R:
(British Columbia Univ.)
i~AaRiiit

Corallinaceae Q)$7"J!l'H;:: -?It •""C -:/liili:Q)=:~Q)·~~il::-?lt•"(Q)j"ifrj!.-............. .=r;IIJ l!ll

Upper Devonian plants from Yokokura-yama, Kochi Prefecture, Japan ......... .
.............. ASAMA, K. & HIRATA, M.
Cor1iopteris (Dicksoniaceae) species from the
Itoshiro Sub-group, the Tetori Group,
Central Honshu, Japan ...... KIMUI~A. T.
ltt•fi/iillf~Q) lobed white oak ft:r:ill.:-?lt•""C. . . . . . . . . . . . . . . . . . . . . . . -11tl*ltm · tMill •~~,fi1~Q) 2· 3 Q)q:s'"iJtti1#1Jil:. -?It•"( ....... .

. . . . . . . . . . . . . . . . . . . . . . . .11tJ:J:I:i$(fft .

**

~~-j

Early and Middle Pennsylvanian fusulinidsof southern British Columbia. Canada
and northwestern Washington, U.S. A.
.............. SAOA, K. & DAJ."'NER, w. R.
Early Permian Parafusulina and Pseudofusulinella from the Chilliwack Group,southwestern British Columbia ....... .
. . . . . . . . . . . . . . SAOA, K. & DANNER, w. R.
Late Lower Carboniferous Eostaffella andHexaphyllia from Central Oregon, U.S. A .
. . . . . . . . . . . . . . SADA, K. & DANNER, w. R ..
Lower Permian fusulines from Terbat,
Sarawak; East Malaysia (Preliminary
report) .............. : ........... TORI·
YAMA, R., YANAGIDA, J. & IsHIBASHI, T.
ti~R~!Ih~t:Jl~I:::.Mt Q ft.:01&nzffiffl/¥1::-? It' -c................................. -3KUl :llr'''Ui1iH.Jl!WJ-c ~*• ~ n t.: ~ #il ~q:s Q)1f1L !:R
IIUU:::.-?It•-c ..................... ·*ltm ;I.)
Some Late Palaeozoic compound corals
from Portuguese Timor.-Palaeontological study of Portuguese Timor(VI)- .................... YAMAGJWA, N.
· Lower Carboniferous Visean faunas discovered from Mitsusawa. southeasternpart of the Kwanto massif, Japan. Pt.
2, Rugosa ................ YAMAGIWA, N.
Lower Carboniferous Visean faunas discovered from Mitsusawa, southea'sternpart of the Kwanto massif, Japan. Pt.
3, brachiopods .............. YANAGIDA, J.
1!9001!97Ji-JIIJiif~Q)H:>~Qfljl:t:l: II mf.1 / -i! 7

A ;r.~::-?lt•-c ...................... !l!fm~~z

Upper Cretaceous pelecypods from theHimenoura Group, Kyushu, Japan. Part
1, Orders Nuculoida and Arcoida (ftlf)!)............................ TASHIRO, M .
Upper Triassic ammonites from Okinawajima. Part 4 .............. ISHIBASHI) T .
Palaeobiogeography of Turonian ammonites.......................... MATSUMOTO, T.
;faM~:ll'I¥~tP t::>l-J.t.: Inoceramus (Sphenoceramus) naumanni fii*!I¥Q)g:_ffi.ji'bt;Q)Iflj
r.n~1l:ft ........................ liJ!llJI-nt
Miocene molluscs from the Bihoku Groupat Miyauchi (Shinchiku)-cho, Shobara
city, Southwest Japan ................. .
.......... OKAMOTO, K. & TERAUCHI, M.
ft:fi7)·t;,~t.:7l<ill~:r?*fli;'* ........ mm

10 Fl 29 s (A)) i!!li¥'i13*q:s'*t~iiH"·!A.nfiAF (ttit*'Wt) ~:ii::ti!!OJ~f.t c =,gJiHt(~~.

1:il*~~l*l~ lk#i!!i=:JJI!

SYSTEMATIC INDEX(New Series No. 81-No. 88)

PALAEOZOOLOGY
Protozoa
Page
CHOI, Dong Ryong: Colania douvillei (OZAWA), a fusulinid Foraminifera, from the Northern Kitakami Mountains, NE Japan................................................
KuRIHARA, Kenji: Foraminifera from the. Hayama Group, Miura Peninsula . . . . . . . . . . . .
MATSUMARU, Kuniterui!' 1 The genera Nephrolepidina and Eulepidina from New Zealand..
OKIMURA, Yuji: Permo-Carboni·ferous endothyraceans from Japan, Part 1. Biseriamminidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
SADA, Kimiyoshi: Fusulinids of the Profusulinella zone of the Taishaku limestone.
(Studies of the stratigraphy and microfossil faunas of the Carboniferous and Permian
Taishaku Limestone in West Japan, No. 2) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
YAMAGIWA, Nobuo and SAKA, Yukiyasu: On the Lepidolina zone discovered from the
Shima Peninsula, Southwest Japan....................................................

369
131
179
414

436
260

Coelenterata
KATO, Makoto:

J. FLEMING's species of British Lower Carboniferous corals............

1

Bryozoa
HAYAMI, Tomoko: Some Neogene Cheilostomata (Bryozoa) from Okinawa-jima . . . . . . . .
- - - - : On some Bryozoa from near Namioka-cho, Minami-Tsugaru-gun, Aomori Prefecture, Japan....... . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
SAKAGAMI, Sumio: The Triassic Bryozoa from Kusaka, Sakawa basin, Shikoku, Japan..

73
196
275

Mollusca
IsHIBASHI, Takeshi: Upper Triassic cephalopods from the Tanoura district, Kumamoto
Prefecture, Japan . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
MAsUDA, Koichiro: On some Pathtopecten from North America . . . . . . . . . . . . . . . . . . . . . . . .
- - - : Amussiopecten from North America and northern South America......... . . . . .
- - - : Swiftopecten of the Northern Pacific . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
MATSUMOTO, Tatsuro, MURAMOTO, Tatsuo and !NOMA, Akitoshi: Two small desmoceratid ammonites from Hokkaido (Studies of the Cretaceous ammonites from Hokkaido
and Saghalien-XXIV) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
MuRATA, Masafumi: Fusulinid biostratigraphy and molluscan fauna from the uppermost
part of the Sakamotozawa formation, and the pre-Kanokura unconformity in the southern part of the Kitakami massif, North-east Japan . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .

486

447
166
205
395

377

93

Systematic Index'NooA, Hiroshi: New anadarid and associated molluscan fauna from the Haneji Formation, Okinawa-jima, Ryukyu Islands . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
- - - : Some fossil Pteropoda from Miyazaki and Okinawa Prefectures, Southwest
Japan . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
NooA, Masayuki: Ammonites from the Mesozoic Yamabu formation, Kyushu . . . . . . . . . .
·SATO, Tadashi: Some Bajocian ammonites from Kitakami, Northeast Japan . . . . . . . . . . . .
TASHIRO, Masayuki: Upper Cretaceous glycymerids in Japan . . . . . . . . . . . . . . . . . . . . . . . . . .
- - - : On the surface ornamentation of the pennatae trigoniids, and on three new
species of the trigoniids from the Himenoura Group, Kyushu, Japan . . . . . . . . . . . . . . . .

487
27
472
462
280
225
325

Arthropoda
Hu, Chung-Hung: Ontogeny of three Cedaria zone trilobites from Upper Cambrian, Montana.................................................................................. 245- - - & TAN, Li-Lin: Ontogenies of two Upper Cambrian trilobites from northernBlack Hills, South Dakota . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
61

Vertebrata
HAT AI, Kotora and NooA, Hiroshi: A Problematica from the Mizuho-To of Niigata Prefecture . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .

319

Miscellanea
HATAI, Kotora and KoTAI<A. Tamio: Some coprolites from Wakayama Prefecture......
52
HAT AI, Kotora, NooA, Hiroshi and OGASAHARA, Kenshiro: Scaphopoda-like fossils fromthe Udo formation (Miocene) of Miyazaki Prefecture, Japan . . . . . . . . . . . . . . . . . . . . . . . . 409
HAT AI, Kotora and NooA, Hiroshi: A fossil worm trail from the Upper Cretaceous Nakaminato formation, Ibaraki Prefecture . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 458

PALAEO BOT ANY-(Megafossils)
KIMURA, Tatsuaki and SEKIDO, Shinji: The discovery of the cycad-like leaflets withtoothed margin from the Lower Cretaceous Itoshiro Sub-group, the Tetori Group,
Central Honshu, Japan................................................................ 190--- &: Ctenis species from the Itoshiro Subgroup (Lower Cretaceous), theTetori Group, Central Honshu, Japan . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 360(Microfossils)
BuRCKLE, Lloyd H.: Correlation of Late Cenozoic marine sections in Japan and theEquatorial Pacific . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 117
FUJI, Norio: Fossil spores and pollen grains from the Neogene deposits in Noto Peninsula, Central Japan-IV. A palynologiccal study of the Late Miocene Tsukada Member . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
. . . . . . . . . . . . . 295
Ko!ZUMI, Itaru: Marine diatom flora of the Pliocene Tatsunokuchi Formation in Fukushima Prefecture . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 340
NisHIDA, Shiro: Nannofossils from Japan IV. Calcareous nannoplankton fossils fromthe Tonohama Group, Shikoku, Southwest Japan ................................ -... 143

488

Nos. 81,.._88 (1971, 1972)

TAKAHASHI, Kiyoshi: Microfossils from the Pleistocene sediments of the Ariake· Sea
area, west Kyushu. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .. . . . . . . . . . . . . . . . . . . . . . .
llt
TAKAYAMA, Toshi<iki: A note on the distribution of Braarudosphaera bigelowi (GRAN& BRAARUD) DEFLANDRE in the bottom sediments of Sendai Bay, Japan............ 429·

PSEUDOFOSSIL
HAT AI, Kotora and NooA, Hiroshi: Peculiar markings on a sandstone layer of the Ilagino Formation, Nagano Prefecture..................................................

162:

489>

INDEX OF FAMILIES, GENERA AND SPECIES
Notes: Words listed are names of families, genera and species, which are either described
or illustrated in the Transactions and Proceedings of the Palaeontological Society of Japan,.
New Series, Nos. 81-88; words in heavy type are names of new genera and species.

A
316
Abies sp.
Actinocyclus ellipticus ................. . 350
elongatus ....... .............. . 350
Adeonellopsis arculifera ............... . 87
- - - sttbleres ....................... . 87
- - - yarrae11sis . . . . . . . . . . . . . . . . . . . . . . 87
Amussiopectelt... . . . . . . . . . . . . . . . . . . . . . . . 209
Amussiopecten antiguensis. . . . . . . . . . . . . . 211- - - churuguarensis................ 213- - - - harrisi . . . . . . . . . . . . . . . . . . . . . . . . 213- - - preglyptus . . . . . . . . . . . . . . . . . . . . 212- - - sp ............................. 215- - - va11vlecki...................... 210- - - wooclringi . . . . . . . . . . . . . . . . . . . . 214
Anadara (Hataiaraca) kogachiensis . . . . 37
Antropora tinct a . . . . . . . . . . . . . . . . . . . . . . . . 77
Aphelaspis walcotti. . . . . . . . . . . . . . . . . . . . . . 62
Apiotrigonia... . . . . . . . . . . . . . . . . . . . . . . . . . 331- - - minor . . . . . . . . . . . . . . . . . . . . . . . . 326- - - obsoleta . . . . . . . . . . . . . . . . . . 326, 331- - - cf. obsoleta.................... 326- - - postonodosa.................... 326- - - undulosa . . . . . . . . . . . . . . . . . . . . . . 326- - - utoensis . . . . . . . . . . . . . . . . . . . . . . 333
Arcestes (Proarcestes) cf. ausseeanus . . 451
Asteromphalus darwinii . . . . . . . . . . . . . . . . 350
Astrononion ltayamaense . . . . . . . . . . . . . . 140
Atlanta rniyagusukuana . . . . . . . . . . . . . . 481
Aviculopecten sp. . . . . . . . . . . . . . . . . . . . . . . 110? sp. . . . . . . . . . . . . . . . . . . . . . . . . . . 111

B
Bacteriosira fragilis. . . . . . . . . . . . . . . . . . . . 350
Baltag nostus beltensis. . . . . . . . . . . . . . . . . . 254
Baltisphaeridium sp. . . . . . . . . . . . . . . . . . . . . 20
? sp. . . . . . . . . . . . . . . . . . . . . . . . . . . 21
Batillaria zo1talis........................ 45
Bellerophon (Bellerophon) kitakamiensis 105
Biedulphia aurita . . . . . . . . . . . . . . . . . . . . . . 350

Braarudosphaera bigelowi . . . . . . . . . . 155, 43(}
Buchites. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 449- - - kumamotoensis . . . . . . . . . . . . . . 449·
Bulimina striata....... . . . . . . . . . . . . . . . . . 142.

cCaberea hataii . . . . . . . . . . . . . . . . . . . . . . . . . . 8(}
Carya sp. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 312
Aff. Carya sp ...... :. . . . . . . . . . . . . . . . . . . . 314
Caryophy/lea (sic) a/finis . . . . . . . . . . . . . . . . 4- - - (sic) duplicata . . . . . . . . . . . . . . . . . . ~(sic) fasciculata . . . . . . . . . . . . . . . . 4
Catapsydrax dissimilis . . . . . . . . . . . . . . . . . . 14Z
Cavolinia okinawana . . . . . . . . . . . . . . . . . . 480- - - - telemus . .'...................... 479·
Cellaria punctata . . . . . . . . . . . . . . . . . . . . . . . . 80·
Ceratolithus cristus . . . . . . . . . . . . . . . . . . . . 155- - - aff. tricorniculatus . . . . . . . . . . . . 15&
Cigclisula occlusa........................ 83.
Cladog ramma californica . . . . . . . . . . . . . . 350
Cleidochasma bassleri ................... . _ 82
- - - biavicularia . . . . . . . . . . . . . . . . . . . . 81
- - - aff. fa/lax... . . . . . . . . . . . . . . . . . . . 82
- - - granifera ...................... 82
Clementia (Clementia) vatheleti . . . . . . . . 42
Cleodora ltataii . . . . . . . . . . . . . . . . . . . . . . . . 475.
- - - okinawana . . . . . . . . . . . . . . . . . . . . 477
Coccolithus pelagicus . . . . . . . . . . . . . . . . . . 149Codakia (]agonia) okinawazimana . . . . . . 41
Colania douvill ei . . . . . . . . . . . . . . . . . . . . . . 371
Conescharellina cf. breviconica . . . . . . . . . . 89concava . . . . . . . . . . . . . . . . . . . . . . . . 8~
Copidozoum planum . . . . . . . . . . . . . . . . . . .
78.
Coscinodiscus lineatus . . . . . . . . . . . . . . . . . . J.50.
nodulifer . . . . . . . . . . . . . . . . . . . . . . 350·
- - - pustulatus . . . . . . . . . . . . . . . . . . . . 35(}
Costazia radiata . . . . . . . . . . . . . . . . . . . . . . . . 89_ _ _ rota . . . . . . . . . . . . . . . . . . . . . . . . . . 201
Cranosina coronata . . . . . . . . . . . . . . . . . . . . . . 78.
Creseis acicula . . . . . . . . . . . . . . . . . . . . . . . . 481

490

Nos. 81,.._88 (1971, 1972)

Cricolithus sp .......................... . 149
Ctenis ..... ............................ . 360- - - kaneharai ................... . 364- - - nipponica ................... . 361- - - s p ............................ . 365
Cyclammina orbiwlaris ............... . 142
Cyclococcolithus leptoporus . ............ . 149
Cymatiosphaera globulosa . .............. . 21
- - - reticulosa ..................... . 21
Cysti sella ? americana . ................ . 201- - - midwaym1ica ................. . 200

D
Denticula kamtschatica- - - seminae ..................... .
Desmoceratinae ....................... .
Diacria bisulcata ..................... .
Diatosula marionense ................. .
Dis coaster brouweri ................... .
- - - brouweri rutellus ............. .
- - - challengeri ................... .
- - - dilatus ....................... .
- - - kugleri . ...................... .
- - - pentaradiatus ................. .
- - - stellulus ..................... .
- - - surwlus ..................... .
Discolithina sp. A ........... : ......... .
Ditrupa ............................... .
------,. miyazakiensis

350
350
378
478

200

423
142
226
226
239
229
228
232
239
234
234
235
235
233
233
234
236
228

154

154

H

154

Hanaia . . . . . . . . . . . . . . . . . . . . . . . . . . . . .. . . 233
Helicoponlosphaera kamptneri . . . . . . . . . . 152
Hemicystodinium cf. zoharyi . . . . . . . . . . . . 18
Hemidiscus cuneiformis . . . . . . . . . . . . . . . . 351- - - weissftogi. . . . . . . . . . . . . . . . . . . . . . 351
Heterotrigonia . . . . . . . . . . . . . . . . . . . . . . . . . . 334- - - diversicostata . . . . . . . . . . . . . . . . 332- - - himenourensis . . . . . . . . . . . . . . . . 334- - - Sltbovalis . . . . . . . . . . . . . . . . . . . . . . 332
Hippoporella spinigera . . . . . . . . . . . . . . . . . . 83
Hitachia . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 460- - - nakaminatoensis.............. 460
Hoeglundina elegans. . . . . . . . . . . . . . . . . . . . 1•12
Holconautilus sp. . . . . . . . . . . . . . . . . . . . . . . 455
Holoporella palmata . . . . . . . . . . . . . . . . . . . . 88
- - - (?) sp. . . . . . . . . . . . . . . . . . . . . . . . . 89
Hyc1locylix striata . . . . . . . . . . . . . . . . . . . . . . 478
Hyalodiscus obsoletus . . . . . . . . . . . . . . . . . . 351
Hystrichokibotium sp. . . . . . . . . . . . . . . . . . . 19
Hystrichosphaeridium ct. ferox . . . . . . . . . . 17
- - - sp. a .......................... 18- - - sp. b .......................... 18
cf. tiara . . . . . . . . . . . . . . . . . . . . . . I 7

154

154
154

155
155
149
412
412

E
Encephalartites leipzigii . . . . . . . . . . . . . . . .
Eulepidina dilatata dilatata . . . . . . . . . . . .

- - - sp. B..........................
Globoquadrina dehiscens . . . . . . . . . . . . . . . .
Glycymeris ........ : . . . . . . . .. . . . . . . . . . . .
Glycymeris (Glycymeris) amaktisensis . .
(Glycymerita) aff. apletos......
--- () himenourensis . . . .
--- () japonica . . . . . . . . . .
--- () multicostata........
--- () veatchii .. ·. .. . .. .. ..
Glycymeris (Hanaia) densilineata . . .
- - - ( - - - ) haipensis . . . . . . . . . .
--- () hokkaidoensis . . . . . .
--- () katsurazawensis . .
--- () matsumotoi . . .
--- () salida . . . . . . . . . . . . . .
--- () sublaevis .. .. .. .. ..
(Pseudoveletuceta) mifunensis
Glycymerita . . . . . . . . . . . . . . . . . . . . . . . . .

192

184

F
Fagus sp ................... ''I'. . . . . . . . . . 312
Fragilariopsis cyclindrus .... :.... . . . . . . . 350
Fulvia sp. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 41

GGlaphyraspis parva . . . . . . . . . . . . . . . . . . . . . . 66
Globigerinatella insueta . . . . . . . . . . . . . . . . 142·Globiualvulina..... . . . . . . . . . . . . . . . . . . . . . 417- - - cf. gracea . . . . . . . . . . . . . . . . . . . . 421- - - granulosa compressa . . . . . . . . . . 420- - - kamensis . . . . . . . . . . . . . . . . . . . . . . 418- - - cf. kantharensis . . . . . . . . . . . . . . 421- - - mosquensis.................... •119
- - - 1·egularis ..................... 422- - - sp. A . . . . . . . . . . . . . . . . . . . . . . . . 423

I
Cf. llex sp. . . . . . . . . . . . . . . . . . . . . . . . . . . . . 314
Inapertipollent ites sp. . . . . . . . . . . . . . . . . . . 312

Jfuglans sp .....

312

491

Index of Families, Genera and Species
K
Kitchinites (Neopuzosia) haboroensis . . 384
Knightites (Retispira?) hanzawai . . . . . . 106
Kubikichtlrys . . . . . . . . . . . . . . . . . . . . . . . . . . 322- - - rm·is . . . . . . . . . . . . . . . . . . . . . . . . . . 322

L
Laevicardium sp. . . . . . . . . . . . . . . . . . . . . . . . . 41
Leiosphaeridia glolmlifera . . . . . . . . . . . . . . 22
Leopecten . . . . . . . . . . . . . . . . . . . . . . . . . . . . 170- - - bakeri . . . . . . . . . . . . . . . . . . . . . . . . 171- - - bakeri diazi .. .. .. .. .. . .. .. ... 172
marquerensis . . . . . . . . . . . . . . . . . . 173
Lepitlolina kumaensis . . . . . . . . . . . . . . . . . . 267- - - multiseptata gigantea
265
multiseptata multiseptata . . . . . . 266
Limopsis kogata . . . . . . . . . . . . . . . . . . . . . . . . 237
Lithostrotion jloriforme . . . . . . . . . . . . . . . . . . 3
marginatum . . . . . . . . . . . . . . . . . . . . 3
striatum . . . . . . . . . . . . . . . . . . . . . . . . 2
Lunella coronatus granulatus . . . . . . . . . . . . 44
- - - sp ............................. 44

Navicula jamalinensis . . . . . . . . . . . . . . . . . .
Neozamitcs . . . . . . . . . . . . . . . . . . . . . . . . . . . .
- - - elongata . . . . . . . . . . . . . . . . . . . . . .
Nephrolepidina hornibrooki . . . . . . . . . . . .
howchini . . . . . . . . . . . . . . . . . . . . . .
orakeiensis . . . . . . . . . . . . . . . . . . . .
Neopuzosia . . . . . . . . . . . . . . . . . . . . . . . . . . . .
Nitzschia cf. exline/a . . . . . . . . . . . . . . . . . .
- - - fossilis........................
- - - - reinholdii . . . . . . . . . . . . . . . . . . . . .
Nixonel/a . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
- - - montanensis . . . . . . . . . . . . . . . . . .
Nodosaria longiscata . . . . . . . . . . . . . . . . . .
Normannites (Itinsaites) cf. itinsae . . . .
Nuphar (?) sp. . . . . . . . . . . . . . . . . . . . . . . . .

351

191
192
186
187
1.85

384
351
352
351
250
251
142
283

314

0
Osmund a sp. . . . . . . . . . . . . . . . . . . . . . . . . . . 314
Ostrea (Ostrea) denselamellosa. . . . . . . . . . 40
Otoites sp. . . . . . . . . . . . . . . . . . . . . . . . . . . . . 28~
Ovoidites cf. microligneolus . . . . . . . . . . . . 2~
- - - ellipsoideus.................... 22M

p

Macoma (Macoma) praetexla ........... . 42
Martinottiella communis . .............. . 142
Melonis pacificus ..................... . 142
albicans ..................... . 351
Mesopuzosia cf. pacifica ............... . 391
Metadoliolina gravitesla . .............. . 270
Micrhystridium ar·iakense ............. . 19---densum ....................... . 20
Microdesmoceras ..................... . 378- - - tetragonum ................. . 378
Microporina articulata ............... . 198
Microtrigonia ......................... . 335
amanoi ....................... . 326- - - imutensis ................... . 335
Millerella sp. B ....................... . 437
Mizuhopecten ......................... . 170- - - warreni ..................... . 170
Modiolus sp ............................ . 38
Mucronella sp .......................... . 84
"Murchisonia" sp. . . . . . . . . . . . . . . . . . . . . . 107
Myriozoum coarctatum ................ . 202- - - subgracile ................... . 202

272
Pachyphloia sp.
Parasmittina aviculoumbonata ......... . 84
- - - peristoaviculata ............... . 84
- - - spathulata ................... . 84
- - - trispinosa ..................... . 85
Patinopecten . .......................... . 169
Patinopecten (Patinopecten) skonunensis 169
Pelekodites cf. pelekus ............... . 282(Spatulites) spatians ......... . 281
Phormedites ........................... . 450

NNankinella yokoyamai . . . . . . . . . . . . . . . . 441
Nassarius (Zeuxis) caelatus . . . . . . . . . . . 46

- - - sp ............................ .
Picea sp .............................. .

450

316

Placites ............................... . 45~
Planktoniella sol ..................... . 352
39
Pododesmus (Monia) noharai .......... .
Polinices cumingianus madioenensis ... . 46
Pontosphaera ~alboranensis ............. . 152- - - japonica ..................... . 152- - - sp. ···························

153- - - cf. vadosa ................... .
Porosira glacial is ..................... .
Profusulinella fusiformis ............. .
toriyamai......................
Pseudoactinodontophora . . . . . . . . . . . . . .
- - - yabei . . . . . . . . . . . . . . . . . . . . . . . .
Pseudobatostomella kobayashii . . . . . . . . . .

152

352
439
43&
111
112
275

492

Nos. 81,-...88 (1971, 1972)

Pseudopodosira elegans ............... . 352
Pseudostaffella taishakuensis ......... . 440
Pseudoosterella sp. . .................. . 467
Pseudoueletrtceta ..................... . 236
Pteria cf. coturnix ..................... . 38
Pullenia bulloides ............ : ........ . 142
Puzosiinae ........................... . 383
Pyxidicula weyprechtii ..... ............ . 352

Q·Quercus sp.

284
Stephanoceras sp.
Stephanopyxis lwrridus ............... . 348- - - schenckii ..................... . 353
Striarca interp/ic~ta ................... . 36
Strigoceras cf. languidum ............. . 285
Swiftopecten ..... ...................... . 398- - - swi/tii ....................... . 399- - - swiftii parmeleei ............. . 401
Syracosphaera pulchra ................. . 153
Syspacheilus dunoirensis ............. .. . 246

314

T

R
Rhabdosphaera claviger . . . . . . . . . . . . . . . . 157
Rhacophyllites sp. . . . . . . . . . . . . . . . . . . . . . 454
Rhaphoneis ischaboensis . . . . . . . . . . . . . . . . 352
margaritalimbata . . . . . . . . . . . . . . 352- - - - tatsunokuchiensis . . . . . . . . . . . . 349
Rhizosolenia bergonii . . . . . . . . . . . . . . . . . . 353
Rhynchozoon aff. grandice/la . . . . . . . . . . . . 86
sp ............................. 86(?) sp .........................

86s.Salix sp. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 312
Schisrnopora chrysalis . . . . . . . . . . . . . . . . . . 202.Schizobrachiella subhexagona . . . . . . . . . . 200
Schizomavella ovoidea . . . . . . . . . . . . . . . . . . 81
- - - sp ............................. 81
Schizoretepora tumescens . . . . . . . . . . . . . . 201
Schyphosphaera apsteini . . . . . . . . . . . . . . . . 157
Siphonodendro11 affine . . . . . . . . . . . . . . . . . . . . 5
Smiltina sp. . . . . . . . . . . . . . . . . . . . . . . . . . . . . 85
.Smittoidea acaroensis. . . . . . . . . . . . . . . . . . . . 85
.Sphaeroidinlllla seminulina . . . . . . . . . . . . 142
Spiniferites ramosus . . . . . . . . . . . . . . . . . . . . 16
Spirosigmoilinella r:ompressa . . . . . . . . . . 142
Sta.ffella akagoensis . . . . . . . . . . . . . . . . . . . . 443
Steganoporella magnilabris:. . . . . . . . . . . . . . 79

Tainoceras kitakamiense......... . . . . . . . 108
Thalamoporella novae·hollandiae . . . . . . . . 79
Thalassiosira antiqua . . . . . . . . . . . . . . . . . . 353
convex a . . . . . . . . . . . . . . . . . . . . . . 353
gravida..... . . . . . . . . . . . . . . . . . . . 353
hyalina... . . . . . . . . . . . . . . . . . . . . . 353
kryophila . . . . . . . . . . . . . . . . . . . . . . 354
nidulus ................ , . . . . . . . 354
nordenskioldi . . . . . . . . . . . . . . . . . . 354- - - zabelinae . . . . . . . . . . . . . . . . . . . . . . 3S4
Tilia sp. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 312
Towapteria nipponica . . . . . . . . . . . . . . . . . . 109
Tricellaria sp...... . . . . . . . . . . . . . . . . . . . . . 199
Tricolpropollenites sp. . . . . . . . . . . . . . . . . . . 316

uUmbilicosphaera cricota . . . . . . . . . . . . . . . . 153
Umbonium (Suchium) moniliferum
decoratum ..... .'. . . . . . . . . . . . . . . . . . . . . . 43.

w
Waagenoperna (Permoperna) hayamii . .

109·

yYabeina columbiana....................
aff. globosa....................

268
269

Transactions and Proceedingsof thePalaeontological Society of Japan
New Series
No. 81 ....... No. 88
1971-1972

Palaeontological Society of Japan

The heading in Japanese commemorates the handwriting of Prof. Matajiro·
YOKOYAMA, father of Japanese palaeontology, who W~S a professor of stratigraphy
and palaeontology at the Geological Institute, Imperial University of Tokyo.

Fossil on the cover is left lower M2 of Palaeoloxodon naumamti (MAK!Y AMA,
1924) from the uppermost part of the Tokyo formati'on (Upper Pleistocene) at Ike·
bukuro, Tokyo.

CQNTEN.TS
I
;

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I

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J!f;or' l • t; ...,'J

1• .... ,-,!

j,

Page

Plate

1-10

1

11-26

2-5

27-51

6-7

I

.,

',/

Number 81 (Pubiish~d· April20, 1971)
Transactions

Article574. KATO, Makoto: J. FLEMING's species of British Lower Carboniferous corals .................................................... .
575. TAKAHASHI, Kiyoshi: Microfossils from the Pleistocene sediments of the Ariake Sea· area, west Kyushu ................... .
576. NODA, Hiroshi: New anadarid and associated molluscan faunafrom the Haneji Formation, Okinawa-jima, Ryukyu Islands . ... .
'..
577. HATA!, Kotara and KOT:"-KA, Tamio: Some coprolites from Wakayama Prefecture ......................................· . . . . . . . . .
Proceedings

52-58
59-60

Number 82 (Published June 30, 1971).
Transactions
578.

Hu, Chung-Hung and TAN, Li-Lin: Ontogenies of two Upper
Cambrian trilobites from northern Black Hills, South Dakota . . 61-72
8-9
579. HAY AMI, Tomoko: Some Neogene Cheilostomata (Bryozoa) from
Okinawa-jima ..........................·........................ 73-92 10-12580. MuRATA, Masafumi: Fusulinid biostratigraphy and molluscan
faj.lna from the uppermost p~u·t of the Sakamotozawa formation,and the pre-Kanokura unconformity, in the so-uthern part of theKitakami massif, Northeast japan .......... .' ................... 93-116 13-14
581. BURCKLE, Lloyd H. : Correlation of Late Cenozoic marine sectionsin Japan and the Equatorial Pacific ............................ 117-128

Number 83 (Published September 20, 1971)
Transactions
582. KURIHARA, Kenji : Foraminifera from the Hayama Group, Miura
Peninsula ........................................................ 131-142

15

ii

583.

NISHIDA, Shiro: Nannofossils from Japan IV. Calcareous nannoplankton fossils from the Tonohama Group, Shikoku, Southwest
Japan ........................_... , ... , ....... , ..................... 143-161 16-lS.
jt~584. HATAI, Kotora and NODA, Hiroshi: Peculiar markin~s on a sandstone layer of the Hagino Formation, Nagano Prefecture ........ 162-165
f"

o

j;

I

'-

'!

I

•

585. MASUDA, K6ichir6: On some Patinopecten .. from No~th America............................... ·..... ·: ........................... 166-178 19-21

Number 84 (Published December 30, 1971)
Transactions
586. MA TSUMARU, Kuniteru : The genera Nephrolepidina and EulePi·
dina from New Zealand ........................................ 179-189 22-23
587. KIMURA, Tatsuaki and SEKIDO, Shinji : The discovery of thecycad· like leaflets with toothed margin from the Lower Cretaceous
ltoshiro Sub-group, the Tetori Group, Central Honshu, Japan .. .' .190-195· 24588. HAY AMI, Tomoko: On some Bryozoa from near Namioka-ch'o,
Minami·Tsugaru:gun, Aomori Prefecture, Japan ................ 196-204
589. MASUDA. K6ichir6: Amussiopecten from North America and nor·them South America ............................................ 205-224 25-26
590. TASHIRO, Masayuki: Upper Cretaceous glycrmerids in Japan .. 225-242 27-28Proceedings ...................... 243-244

Numper 85 (Published April 20, 1972)
Transactions . . . .
591. Hu, Chung· Hung: Ontogeny of three Cedaria zone trilobitesfrom Upper Cambrian, Montana ................... ·............. 245-259 29-30
592. YAMAG!WA~ Nobuo and SAKA, Yukiyasu: Ori the Lepidolina zohe
discovered fromthe Shima Peninsula, Southwest Japan ... ·..... 260-274
31-32
.
.
593. SAKAGAMI, Sumio: The Triassic Bryozoa from Kusaka, Sakawa
basin, Shikoku, Japan ........................ : ................. 275..:279
33
594. SATO, Tadashi: Some Bajocian ammonites from Kitakami Nor·theast Japan ............................. : ...................... 280-292
34
Proceedings .......................... 293

iii
! •

Number 86. (Pt.Jl>lished june 30,1972)

:595.

'596.
.'597.

:598.
599.

·600.

Transactions.
. . . _. ·I :FUJI, Norio: Fossil spores and· pollen grains from the Neogene
deposits in Noto Peninsula, Central· Japan-IV. A palynologicalstudy of. the Late Miocene 1.'!!ukada
Memper ......... , ...
._ ..... 295-318
35-38
'
. '
..
HAT AI,
Kotoraand
NODA,
Hiroshi:A
Problematica
from
the
...
.
.
.
Mizuho·To of Niigata Prefecture ............................ : ... 319-324,
39
TASHIRO, Masayuki: On the surface ornamentation of the pen·natae trigoniids, and on three new species of the trigoniids fromthe Himenoura Group, Kyushu, Japa_n ... : ...................... 325-339 40-41
KOIZUMI, Itaru: Marine diatom flora of the Plioce~e Tatsuno·
kuchi·Formation,in Fukushima Pr~fecture ...................... 340-359 41-43
KIMURA, Tats~aki and SEKIDO, Shi~ji : Ctenis species from theltosh\ro Sub-group (Lower Cretaceous), the Tetori Group, Central
Honshu, japan .................................................. 360-368 44-45
CHOI, Dong Ryong: Colania douvillei (OZAWA), a fusulinid Fora·
minifera, from the Northern Kitakami Mountains, NE japan .... 369-374
46~

Number 87 (Published September 30, 1972)
Transactions
·601.

·602.
·603.

·604.
·605.

-606.

MATSUMOTO, Tatsuro, MURAMOTO, Tatsuo and lNOMA, Akitoshi :Two small desmoceratid ammonites from Hokkaido (Studies of
the Cretaceous ammonites from Hokkaido and Saghalien-XXIV) .. 377-394
47
MASUDA, Koichiro: Swiftopecten of the Northern Pacific ...... 395-408 48-49
HAT AI, Kotora, NODA, Hiroshi and OGASAHARA, Kenshiro: Sea·
phopoda·like fossils from the Udo formation (Miocene) of Miyazaki Prefecture, Japan .......................................... 409-413
OKIMURA, Yuji:
Permo-Carboniferous endothyraceans fromJapan, Part 1. Biseriarnminidae ................................ 414-428 50-51
TAKAYAMA, Toshiaki: A note on the distribution of Braarudosphaera bigelowi (GRAN & BRAARUD) DEFLANDRE in the bottom
sediments of Sendai Bay, Japan ................................ 429-435
SADA, Kimiyoshi : Fusulinids of the Profusulinella zone of the
Taishaku limestone. (Studies of the stratigraphy and microfossil
faunas of the Carboniferous and Permian Taishaku Limestone in

iv

West japan, No. 2) .............................................. 436-445 52-53:-::1'•!··-..;.:•:!t.•·'Number 88 (Published December 20, 1972)· F:~:-tt•".::/! ,d r r, • ·:,,fl ·~. :1': ., • f,:·
!1. ··.··t :•. , • ,. •._. T..ransactio.n...s._ , _ ..•· ... .
__
j

·;

, , , ... : :·,•• r •••• ,;~ ( 1r; ': _:6o{'· I§ms.\sHr, Takeshi: Upper Triassic cephalopods'from the·Tanouta·'· ·, .. :- :..
district, 'Kum3:il1b'to Prefecture; Japan··:.'.~· .. .'.: ·.. ·. ::.' .. ·..... ·, :·! ·:: :447'-457'608. 1-IATAI.' Kotora and NODA, Hiroshi: A fossil ·worin trail ·from the ·Upper Cretaceous Nakaminato formati.on, lbaraki Prefecture .... 458-461
609. · NODA, Masayuki' :· Ammonites from the Mesozoic .Yamabu forma'· · tion, Kyushu . : .... : .. : ............ , ......... ·................... :461-471
610. NODA, Hiro'shi': Some fossil Pteropoda from Miyazaki and Oki ..
n~rwa Prefectures, Southwest Japan ........................... .472-484
Proceedings . : . ......................... 485
Systematic Index ..................... .486-488
Index of Families, Genera and Species .. 489~492•'' ·;

' ·54

·55
5657

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Transactions and Proceedings of the PalaeontologicalSociety of Japan

New Series No. 88

December 20, 1972

CONTENTS
TRANSACTIONS

607. lSHIBASIU, Takeshi: Upper Triassic cephalopods from the Tanoura district,
Kumamoto Prefecture, Japan. . ....................................... 447
608. HATAI, Kotara and NODA, Hiroshi: A fossil worm trail from the Upper
Cretaceous Nakaminato formation, Ibaraki Prefecture................. 458
609. NODA, Masayuki: Ammonites from the Mesozoic Yamabu formation, Kyu·
shu .................................................................. 462
610. NODA, Hiroshi : Some fossil Pteropoda from Miyazaki and Okinawa Pre·
fectures, Southwest Japan ............................................ 472
PROCEEDINGS ................................................................ 485
Systematic Index .............................................................. 486
Index of Families, Genera and Species. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 489

Editor: Takashi HAMADA
Associate Editor: Yasuhide IWASAKI

Officers for 1971 - 1972
President:

Tokio SHIKAMA

Councillors (* Executives) : Kiyoshi ASANO*, Kiyotal<a CHINZEI*, Takashi HAMADA",
Tetsuro HANAI*, Kotora HAT AI, ltaru HAY AMI, Koichiro ICHIKAWA, Taro
KANAYA, Kametoshi KANMERA, Tamio KOTAKA, Tatsuro MATSUMOTO*, Hiroshi
OZAKI*, Tokio SHIKAMA*, Fuyuji TAKAI*, Yokichi TAKAYANAGISecretaries: Wataru HASHIMOTO, Saburo KANNOExecutive Committee
General Affairs : Tetsuro HANAI, Naoaki AOKIMembership: Kiyotaka CHINZEI, Toshio KOIKE
Finance: Fuyuji T AKAI, Hisayoshi !GO
Planning : Hiroshi OZAKI, Kazuo ASAMA
Publications
Transactions:
Special Papers:

Takashi HAMADA, Yasuhide IWASAKI
Tatsuro MATSUMOTO, Tomowo OzAwA" Fossils" : Kiyoshi ASANO, Toshiaki TAKAYAMA---------------- - ---All communications relating to this journal should be addressed to the
PALAEONTOLOGICAL SOCIETY OF JAPAN
c/o Business Center for Academic Societies, Japan
Yayoi 2-4-16, Bunkyo-ku, Tokyo 113, Japan.
Sole agent:

University of Tokyo Press, Hongo, Tokyo