DIWPA: DIVERSITAS in the Western Pacific and Asia

March, 2017

DIWPA News Letter

No.36

Ofﬁce:CenterforEcologicalResearch,KyotoUniversity,Otsu,Japan

Message from the Chairperson

We, Center for EcologicalResearch (CER), haveconcluded the Memorandum ofUnderstanding (MoU) for Academic
and Research Cooperation with
National Institute of Ecology (NIE),
South Korea on December 1, 2016.
Shin-ichi Nakano
NIE was established in 2013, tobe an institution for ecology and biodiversity, andhas been recognized as one of the most importantresearch institutes for ecology in Asia now. Prof. Jae
C. Choe, the former NIE President, and I had many
discussions about the MoU through e-mail since
January 2016. Then Prof. Jeong-Kyu Kim, Head at
Bureau of Ecological Research, Dr. Chulgoo Kim
and Dr. Eunha Ko visited CER on October 18, 2016
to have a discussion about the MoU and a joint
symposium between NIE and CER. On December 9,
2016, six CER professors together with Dr. Tsubaki
the former CER Director visited NIE, and we held
the symposium with many people celebrating thesigning of our MoU. Presentations by both NIE
and CER researchers were very excellent and I wasvery much impressed with the intense and insightful
discussions during coffee-breaks
We will collaborate in the fields of ecology,
biodiversity and environmental sciences thatmutually benefit both institutions, so as torecognize the importance of academic andresearch collaboration, as well as to enhancemutual understanding and establishing long-termcooperation between the two institutions.
CER has 25 years of history with many
trials and tribulations, and is looking forward to
having more events in future. NIE is a fresh andenergetic institution, and is expected to have somefuture challenges. CER would like to provide its
experience for development of NIE, with hope forthe formation of Asian base for ecology, biodiversity
and environmental sciences as mutual collaboration
between NIE and CER. South Korea and Japan areclose in terms of climate, culture and geographicaldistance. Such scientific exchanges allow us toshare the variety of histories for ecology, limnology,
mathematical biology, microbial ecology and so on.
Through the act of MoU, we will exchange scientificmaterials, information, faculty members, post-docs
and students, and conduct joint research activities, inhope of the development of the future researches in
Asia and all over the world. We will provide regular
updates on the activities through the MoU. If you areinterested in our activities, please join us.

International Field Biology Course (IFBC) in 2017

IFBC will be conducted in Japan in the summer of 2017, where graduate and undergraduate
studentsintheWesternPaciﬁcandAsiaregionswillstayatastationtogetherwith
and learn in the ﬁeld of terrestrial cosystems. Motivated students, who belong towhose supervisors are the DIWPAmembers, are highly welcome for applying.The details will
be announced in the spring on our HP. Please check our HP. (http:/diwpa.ecology.kyoto-u.
ac.jp/index.html)

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DIWPA News Letter No.36

Report 1

2016 DIWPA International Field Biology Course, Kiso River,
Nagano, Japan
Dwi Atminarso
Research Institute for Inland Fisheries, Ministry of Marine Affairs and Fisheries(Indonesia)

D

IVERSITAS in the Western Pacific and Asia tickets with vending machines for trains and buses(DIWPA) is a non-governmental and non- when I had to take transportation to a hotel by myself.
political-regional organization financially supported Before the departure, I looked up for information on
by the Center for Ecological Research (CER), Kyoto how to use vending machines and transport paths I
University. DIWPA focuses on the integration of the would take, and I read carefully the instructions given
efforts and resources of scientists towards the study bytheDIWPAoffice.
of the unique ecosystems of the Western Pacific andI arrived in Osaka on 15 August from theAsia including the terrestrial Green Belt from Siberia Palembang-Jakarta-Tokyo which indeed was a very
to New Zealand and the marine Blue Belt along its east pleasant for me to enjoy flying with one of the best
border. The secretary office of DIWPA is launched at airlines in the world, All Nippon Airways, with first
CER. One of the main functions of DIWPA is “capacity class service and food, and typical Japanese travel
building of scientists in particular young scientists companions. At 10:00 local time, the plane landedfrom developing countries”. In the summer of 2016, smoothly at Itami-Osaka airport. I had to use two
the International Field Biology Course (IFBC) took different types of transportations, buses and trains, toplace in Kiso River, Nagano, Japan, where the graduate reach the hotel booked. I had no problem buying a bus
students in the Western Pacific and Asia regions ticket to Shin-Osaka station because their customer
stayed at a station together with professors and learned service supported me. A 25-minute travel to Shinstream ecology including basic limnology in streams, Osaka was smoothly without any traffic jam at all.
fundamental ecological and/or biodiversity studies on After arriving at Shin-Osaka station I had to buy asessile algae, benthic invertebrates, freshwater fish, ticket to Seta station, the nearest station to CER. I
data analysis and database preparation.
wanted to purchase a ticket with a vending machine,
I am a researcher from Research Institute for Inland but I did not find the English language version on it.
Fisheries, The Republic of Indonesia. My institution Finally, I asked the staff for assistance. It was veryis responsible for conducting research studies on difficult to understand Japanese train systems
freshwater fishery resources, environment and the information is displayed only in Japanese language.
study of fish stock assessment throughout Indonesia. I finally reached to Seta station at 13:00 after relying
IFBC appealed to me because Japan is a modern on the direction provided via the e-mail from DIWPA
country, which produces superior fisheries research secretary. It surprised me that almost all trains come on
result, so I believe I get a lot of experience and skills time in Japan. The hotel that was reserved by DIWPA
there. My chief and deputy recommended me to submit is only 300 m far from Seta station. The hotel is clean
an application to DIWPA. I was very grateful when and modern and have nice scenery. It is also convenient
I got an email from DIWPA office that I am one of for shopping. Although the room was not spacious, it
the winners to join the short course. This is my first was very comfortable
opportunity to go overseas. It was fortunate that Japan,
Not long after I arranged clothes in the room anda developed country, was my first overseastook
countrya bath, tothe phone in my room rang from Professor
visit. I had a lot of pre-trip preparation works before Nakano made the call from the hotel lobby. It wasgoing to Japan. I have started preparing my training heartwarming moment to meet Prof. Nakano again
equipment a month before my departure, such as after we first met at the inland fisheries seminar inbuying swimsuits, waterproof shoes, luggage, portable Palembang, Indonesia, five years ago. He did notirons and so on. Using technological electronics in recognize me at the time we met in the hotel lobby
Japan made me a bit nervous, especially purchase of but he was shocked after I told him about our times in2

DIWPA News Letter No.36

Report 1

Fig. 1. Center for Ecological Research (CER)

Fig. 2. Genetic laboratory

Palembang and he gave me a hug. After the greeting the authorities to do research and take samples on the
he took me to his office at the Center for Ecological river. On the third day, a lecturer taught us the sampling
research (CER), Kyoto University with the public techniques for macroinvertebrate and chlorophylltransportation bus taking an approximately 30-minute using a brush, plastic containers, markers and a grid ofdrive and 25-minute walk (Fig. 1). View of the city rubber. For chlorophyll sampling, we chose green or
was very charming, clean, orderly and organized, and greenish-black rocks with a smooth surface. Then thethe traffic rules are obeyed. Prof. Nakano showed
professor me
showed us how to use the grid, brush stones,
their laboratories. Their equipment are very modern and to rinse the brush with water in plastic containers.
and fully maintained. He also introduced me to the For sampling macroinvertebrate, we placed sybernet
researchers and employees in CER. I am very interested at the bottom of the shallow water and then made thein the genetic laboratory with the best and latest clay mouth to affix to the sybernet so as to facilitate
equipment which range from every stage of the genetic insects to enter. After completing the chlorophyll andanalyzes (Fig. 2). These equipment are of domestic macroinvertebrate sampling, all samples were labeledproductions. Besides the genetic laboratory, they also according to the corresponding location (latitude andhave other laboratories that facilitate researchers to longitude), collector and the sampling time. After that,
do any research related to marine and inland water all samples were inserted into the cold box that has
biodiversity. Many international students from different been filled with gel ice to maintain the temperature
countries were awarded scholarship to do research prior to further analysis. Before the samples wereusing CER facilities every year. After that, Prof. taken to the laboratory, a lecturer taught us the way of
Nakano invited me to a famous Japanese restaurant in good sorting using the proper equipment. After that,
Seta to have dinner together. I am very interested in a lecturer, Dr. Jeremy Jay Piggot from New Zealand,
Japanese cuisine served in the restaurant because it is taught us how to use the identification book to best
myfirsttimetryingJapanesefood.
identify macrozoobenthos.
On the second day, we drove 7 hours to Kiso
On the fourth day, a lecturer taught me about basic
biological station in Nagano prefecture. 10 participants limnology, which he covered how to measure dissolve
including 4 teachers and 6 students shared two CER oxygen, acidity, and conductivity in situ using digital
official cars. I was the only foreign studentequipment.
taking thisHe also showed me how to measuretraining course. We arrived at Kiso biological station chlorophyll using two techniques, fluorometer andat 15:00. Only one person is in charge of the building spectrophotometer technique. After that, Prof. Keisukeand provided meals during our stay in Nagano. Here Koba taught us how to measure nitrate in the water.
wehadtobeself-sufficient;cleaningroomsand
toilets,
He showed
me to measure nitrate step by step. And
preparing the beds, and washing our own clothes. After then, Dr. Yoshikuni Hodoki taught us how to use DNA
finished tidying up the bedroom, we visited
Kurakawa
techniqueto identify cyanobacteria. In the afternoon,
River to check the possibility of conducting a training Prof. Nakano taught us the identification of periphof sampling techniques and requested permission from in the river. He also taught us how to use the book to3

DIWPA News Letter No.36

Report 1

identify periphyton well and make observations using
a microscope with thoroughness. In the evening, Dr.
Jeremy Jay Piggot, a researcher and a lecturer of Otago
University, gave a presentation on his research project
conducted in Germany and Australia, which have goodresults for the management of the river at the location.
He is a researcher who has numerous publications
in journals with high grade impact factor. Following
the presentation, a documentary film associated withstream ecology was shown to us. The film contained
words of an expert limnology of Hokkaido University,
Dr. Shigeru Nakano, who has contributed significantly
Fig. 4. Sorting samples
to the development of limnology before he passedaway.
2. EPT index was 45t/25/30% in site 1 and 58/24/18%
On the fifth day, Dr. Yoshikuni Hodoki taught in site 2. Ephemeropterans dominated in both two
us laboratory techniques for the genetic analysis of sites, but superiority of plecopterans and tricopteranscyanobacteria. He covered laboratory techniques different in two sites (Fig. 4). I have presented myranging from DNA extraction to the Polymerase results in a seminar which all participants and lecturers
Chain Reaction. Each participant was responsible were present. There were many questions asked, andfor one sample. After completion of the DNA lab, discussions and suggestions were given to improve myProf. Koba and Prof. Nakano taught us the data paper.
calculation of chlorophyll analysis results. EachOn the sixth day, we cleaned up the station andparticipant worked on the calculation for the respective prepared for going back to Seta. We departed at 9:00
samples we took. During the day, all participants and arrived in Seta at 15:00. I still had one day in Seta
were given two hours to work on our own research. and one day in Osaka before going back to my country.
I decided to conduct individual research with title Finally, I expressed my gratitude to DIWPA (Diversity“Diversity of Macrozoobenthos and EPT index at in West Pacific and Asia) for making my dream of
Kurokawa Station”. I wanted to know how many going abroad come true, to concentrate on the studying
macrozoobenthos species live at Kuro kawa station of stream ecology. Beside that, I also had many great
and the percentage of ephemeropterans, plecopterans, encounters with good scientists, lecturers and students
and trichopterans. I took two samples from two from different countries. Japan is a great country withsites in indifferent water discharge and vegetation many excellent lecturers and scientists in all research(Fig. 3). Site 1 was characterized by slow current areas especially in fisheries. I hope I can continue mwater and surrounding of aquatic plants. Site 2 was study in Japan under the Masters degree scholarshipcharacterized by rapid current water and surrounding program.
of many stones. From our analysis, I found six families
macroinvertebrate and 102 individuals in site 1 and 4
families macroinvertebrate and 114 individuals in site

Fig. 3. Collecting samples at Kuro-kawa research station

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DIWPA News Letter No.36

New Site 1

Transformation of tropical peatland ecosystems in Indonesia
Satomi Shiodera
Center for Southeast Asian Studies (CSEAS), Kyoto University(Japan)Tropical peatlands exist mainly in Southeast Asia
and South America. The total area is 0.44 million
2
km with distribution in Southeast Asia (0.25 million
km2, 56 %) and most of them are found in Indonesia(47 %), Malaysia (6 %) and Papua New Guinea (3 %),with small pockets and remnants in Brunei, Myanmar,
the Philippines, Thailand and Vietnam (Page et al.
2011). They are distributed in low lying areas andriversides, where the forest floor would be flooded
during the rainy season (Fig. 1). Plant residues suchas fallen leaves, branches and trunks are soaked in thewater, and the activities of the soil microorganisms aresuppressed. As a result, plant residues accumulate aspeat soil. This tropical peat is called “woody peat”, andcan be distinguished from the peat in the high latitudes
that is formed by sphagnum or grass litters due to lowtemperature and under water logged condition. The
organic materials in the soil are maintained as a peat insteady state, and tropical peatlands store large amountsof carbon, where it was said about 20 percent of global
soil carbon (89 Pg) might be accumulated in tropicalpeatlands (Page et al. 2011). Due to their particularcharacteristics, such as their tendency to be flooded
seasonally and low soil nutrients, tropical peatlands
have been neglected from development for a long
period. Thus, forests in tropical peatlands (tropical peatswamp forests) abound with many endemic species
adapted to its specific environment and have served
as habitats of the endangered species (animals such as
the orangutan and tree cat, and rare tree species such
as Gonystylus bancanus and Meranti). As the forests
are also a treasure-trove of indigenous species, theyplay an important role in the nurturing biodiversity andin preserving rare species (Anderson 1963, Whitmore
1984, Page et al. 1999, Simbolon and Mirmanto 2000,
Yule 2010, Mirmanto 2011, Rahajoe et al. 2016).
However, for these two decades, the deforestationand degradation of tropical peatlands due toanthropogenic disturbance have caused various
environmental issues. In Indonesia, the large-scale
agricultural land development project, which called“mega rice project” had been conducted in Central

Fig. 1. Peat swamp forest in the rainy season

Kalimantan Province in 1990s. On the other hand,development of the large scale plantation of fastgrowing trees and oil palm made great advancementin Sumatra Island. Infrastructure for these large-scale
agricultural land and plantation development, such as
roads and canals, enabled more people to enter the area,transforming the peat swamp forests into a frontier
for companies and migrants alike. Drainage canal,
which had been established, had led to a decrease ofgroundwater levels and drying of peatlands, resulting in
an increase of emissions of greenhouse gases (GHG),
especially carbon dioxide (CO2) by peat decomposition,and frequent fires (Hirano et al. 2009, 2012, 2014).
Furthermore, the smoke hazards (haze) from the peatfire have become more serious issues in recent years(Fig. 2). The biodiversity of the tropical peat swamp

Fig. 2. Smoke hazards (haze) in Riau Province

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DIWPA News Letter No.36

New Site 1

Knee roots and pneumatophores help to enhance gas
exchange in the waterlogged substrate (Fig. 3). They
also adapt to lower nutrients to develop the root mat
system to enhance uptake of nutrients since they are
more abundant in the surface layers (Posa et al. 2011).
However, after peat fire disturbance, the environment
will be totally changed from the pristine peat swamp
forests. Although the area suffers flooding and low
nutrients, it will change from the closed forest to theopen area, and suffers from severe dry situation in thedry season. Once the delicate balance between variousorganisms is destroyed by disturbance, it is very
difficult to recover in a short period. In the trop
is well known that Alang-alang (Imperata cylindrica,
Poaceae)grasslandwillbeestablishedafteralar
However, in the peatlands, the vegetation seen after thefire is grassland of various ferns. In peat fires, becau
not only forest trees but also the peat soil itself wereburnt, the ground becomes uneven and there will bemany ponds filled with rain water in rainy season.Th
circumstance suppresses the germination and growthof plant seedlings there, and only plants which have a
Fig. 3. Pneumatophores of treestrong resistance to this environment can grow up. For
forests have also been threatened by these disturbances this reason, groundwater level and water chemistry(Shiodera et al. 2016). In 2015, in 2.1 million ha of can be important factors to define the ecosystems offorests was burned, and most of them were peatlands in disturbed peatlands. We research on the kind of species
Indonesia.
that can grow and the factors that can regulate the
The research site of our group is in the Sumatra development of vegetation after severe fire distIsland, Indonesia, which is one of the hot spots of in the sight of ecology and hydrology. While a large
environmental issues in peatlands. Our objectives are to number of forests continue to be lost due to rapid
understand the mechanisms and environmental factors anthropogenic forest disturbances, clarifying theto construct ecosystems of peat swamp forests, to detect mechanisms of remaining forests and recovery systems
the change of biotic and abiotic (that is climate and for degraded forests remains an important task.
geology, etc.) factors after anthropogenic disturbance,
Our group consists of researchers from various
and to clarify the possibility of forest recovery after the backgrounds, which is hydrology, biogeochemistry,
disturbance. Peat swamp forests are the specificbiology, forests
ecology and atmosphere chemistry. Ourthat have been established under seasonal flooding research targets also include various biotic and abiotic
over many years. Fundamentally, the inputs of water components in pristine and disturbed tropical peatand nutrients to the ecosystems are only from rain swamp forests. We deal with not only all importantwater, and the nutrients are fixed with undecomposedfeatures of ecosystems such as soil, water, air, tree
conditions as peat soil, or leach to the groundwater. and insects, but also microclimate and geologicalThus, soil nutrient level and pH is extremely low. This information. For example, understanding the material
situation has been maintained under the suppressed cycling in peat swamp forests, especially carbon
activity of microorganisms by flooding for a long and nitrogen cycling, is important to recognize theperiod of time. In peat swamp forests, adaptation to mechanisms of the emission of GHG from disturbed
the extreme environment can be an important factor peatlands. For this object, many kinds of simultaneousof survival of the plant species. Plants have various observations are needed to carry out. Here, we also
special organs or traits for it. They have buttress and research about 1) the carbon storage in peat soil, 2)
stilt roots to provide stability in the soft peat substrate. changes in water quality and quantity which determine

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DIWPA News Letter No.36

New Site 1
the amount of carbon flow in water, 3) CO 2 andmethane (CH 4) emission rates from soil surface tothe atmosphere and also 4) wood or soil decomposers(such as termites and ants) activities to understandcarbonflowinthewholeecosystem.
In Center for Southeast Asian studies, we conducted
the project entitled “In Search of Sustainable
Humanosphere in Asia and Africa” (the JSPS Global
COE Program, project reader: Prof. Kaoru Sugihara,
CSEAS) in tropical peatlands in Riau Province,
Indonesia from 2007 to 2011. We will also start a new
project, which targets the whole tropical peatland
beyond Indonesia, named “Toward the Regenerationof Tropical Peatland Societies: Establishment of anInternational Research Network and Proposals fora Better Future” sponsored by Research Institute of
Human and Nature (RIHN) (project reader: Prof.
Kosuke Mizuno, CSEAS). In this project, we willanswer how we can control and manage tropicalpeatlands under corroboration with local societies. Our
research project will conduct transdisciplinary researchon the social-ecological system of tropical peatlands tounderstand and address their vulnerabilities. Throughcollaboration with local stakeholders, the project will
integrate scientific findings with local knowledges
to establish mitigation and adaptation strategies inorder to achieve sustainable (low carbon) use andconservation of tropical peatlands.

and vegetation in a tropical peat swamp forest.
Philosophical Transactions of the Royal Society of
London Series B-Biological Sciences 354(1391):
1885-1897.
Page S.E. et al. 2011. Global and regional importanceof the tropical peatland carbon pool. Global Change
Biology 17: 798-818.
Shiodera S. et al. 2016. Impact of peat-fire disturbanceto forest structure and species composition in tropicalpeat forests in Central Kalimantan, Indonesia, In
Tropical Peatland Ecosystems (Mitsuru Osaki andNoriyuki Tsuji Eds.), pp 197-212, Springer Tokyo,
Tokyo, Japan.
Simbolon H. and E. Mirmanto. 2000. Checklistof Plant Species in the Peat Swamp Forests ofCentral Kalimantan. Indonesia. Proceedings of theInternational Symposium on Tropical Peatlands: 179190.
Whitmore T.C. 1984. Tropical rainforests of the Far
East, 2nd edn. Oxford. Clarendon Press.
Yule C.M. 2010 Loss of biodiversity and ecosystem
functioning in Indo-Malayan peat swamp forests.
Biodiversity and Conservation 19: 393-409.

Message from the Secretary GeneralW

e held an International Field
Biology Course (IFBC) at a
research station in a stream of Kiso
River in Japan in August, 2016. The
report from the attended student ispublished in this volume.
In February 2017, we conducteda
symposium
at the Ogasawara
Atsushi Ishida(Bonin) islands, which is one ofWorld Natural Heritage sites, with general public.
The several reports were written in the past DIWPA
Newsletters. In the summer of 2016, the Ogasawara
islands were besieged by relatively prolonged drought
and some drought-adapted endemic trees died. In the
symposium, physiological mechanisms of droughtinduced tree die-offs were reported. The reports of thesymposium will be published in the next volume.
In Thailand, severe drought occurred in the dry
season from 2015 to 2016 and some canopy trees died.
In January 2017, severe flooding occurs especiallyin southern part of Thailand. These climatologicalphenomena may be related to global climate change,and have affected severe influence not only to humanlife but also to ecosystems.
We have to enhance the ability of predictionof ecosystem change in future and to consider its
adaptation. We hope that DIWPA becomes a good
platform to exchange our opinions. Please send theinformation of the situations in your countries to
DIWPA Newsletter (diwpa@ecology.kyoto-u.ac.jp).

References
Anderson J. 1963. The flora of the peat swamp forestsof Sarawak and Brunei. Including a catalogue of allrecorded species of glowering plants, ferns and fernallies. Gard Bull Singap 29: 131 228.
Hirano T. et al. 2009. Controls on the Carbon Balance ofTropical Peatlands. Ecosystems 12: 873-887.
Hirano T. et al. 2012. Effects of disturbances on the
carbon balance of tropical peat swamp forests. Global
Change Biology 18: 3410-3422.
Hirano T. et al. 2014. Carbon dioxide emissions throughoxidative peat decomposition on a burnt tropicalpeatland. Global Change Biology 20: 555-565.
Joeni Setijo Rahajoe et al. 2016, Floristic diversity in thepeatland ecosystem in Central Kalimantan Mitsuru
Osaki and Noriyuki Tsuji Eds.), pp 167-196, Springer
Tokyo, Tokyo, Japan.
Mirmanto E. 2011. Spatial distribution, dynamics andrecovery of wetland forest in Central Kalimantan,
Hokkaido University, Sapporo, Japan.
Page S.E. et al. 1999. Interdependence of peat

7

CALL FOR NEW MEMBER OF DIWPA
We are now calling for membership of DIWPA. Membership fee is no charge. When you become amember, you can;
1. Receive the DIWPA News Letter
You can receive the News Letter by post or e-mail. News Letter contains various information of
biodiversityresearchinAsiaPacificarea,especiallyreportsoneveryprojectandactivitiesofAP-BON(Asia–Pacific Biodiversity Observation Network) as well as J-BON (Japan Biodiversity Observation
Network), IPBES (Intergovernmental Science and Policy Platform on Biodiversity and Ecosystem)and new big project which is “Observation, Evaluation and Prediction of Biodiversity in Asia” by theEnvironment Research and Technology Development Fund. All you can get above information is
ONLYinDIWPANewsLetterexceptthatAP-BONprovidestheminitsownwebsite.
2.ApplyfortheFieldBiologyCourse
You can apply for the Field Biology Course sponsored by DIWPA. The chosen candidate would
participate the Field Biology Course which is organized by Kyoto University. There are few
international initiatives which conduct educational activities such as DIWPA even though some of theinitiatives carry out biodiversity research.
3. Run your articles regarding your biodiversity activities in DIWPA News Letter
DIWPA introduces our member’s various activities in DIWPA News Letter. Your activities would spreadthroughout the world, and you may receive more information about biodiversity conservation as wellas the supports.
4.Buildupacircleoffriendswithinbiodiversityresearch
Ask DIWPA when you want some information of foreign countries regarding your biodiversity
research. DIWPA can introduce people who have information you want. More than 400 members in
4 countriesbelongtoDIWPAnetwork.
IfyouwouldliketojoinDIWPA,pleasecontactto"DIWPA@ecology.kyoto-u.ac.jp".

DIWPA OFFICE
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Secretary
GeneralSecretary
F o r a f r e es u b s c r i p t
tiot
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addresschange,pleasee- mailtothe
DIWPA
OFFICE
diwpa@ecology.kyoto-u.ac.jp
We a r er e a dt
y od e l i vaenre l e c t r oc noip yc t oyou. Thankyou foryourcooperation
inpromoting
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Also,we alwayswelcomeyourarticles,
newsorany
information
relatedtobiodiversity
forDIWPA
News
Letter.
Pleasesendthemtous.

Shin-Nakano
ichi
Atsushi
IshidaShigeoYachi
Sakai
Yoshikuni
Hodoki
Assistant
Secretary
Akiko
Seki

CenterforEcologicalResearch
Kyoto University,3-905chome,-2

Hirano,JAPANOtsu,31 2-0 5
Tel & Fax: 832-9457 18+
E-mail: diwpa@ecology.kyoto-u.ac.jp
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