 ᖺᗘᴫせࡢⓎ⾜࡟࠶ࡓࡗ࡚
 ᙜࢭࣥࢱ࣮ࡣࠊᖹᡂ  ᖺ  ᭶࡟ᐇ㦂⏕≀ඹྠ฼⏝㒊㛛ࠊື≀ᰁⰍయඹྠ฼⏝㒊㛛ࠊ㑇ఏᏊᐇ㦂ඹྠ฼⏝㒊㛛ࡢ  㒊㛛࡛ᵓᡂࡉࢀࡿᩍ⫱◊✲ඹྠ฼⏝᪋タ࡜ࡋ࡚Ⓨ㊊ࡋࡲࡋࡓࠋྛ㒊㛛ࡣྛࠎࠊᪧᐇ㦂⏕≀ࢭࣥࢱ࣮ࠊᪧ⌮Ꮫ㒊㝃ᒓື≀ᰁⰍయ◊✲᪋タࠊᪧ㑇ఏᏊᐇ㦂᪋タ࠿ࡽᨵ⤌ࡉࢀ࡚ࡁࡓṔྐࢆࡶࡗ࡚࠾ࡾࠊ⌧ᅾࡢᙜࢭࣥࢱ࣮ࡢ୺࡞άື࡜ࡋ࡚ࠊ㏆஺⣔ࣛࢵࢺ࣭࣐
࢘ࢫࡢ⣔⤫ಖᏑࠊ⏕≀⣔ᐇ㦂ᐊ࣭㣫⫱ᐊ࣭ᐇ㦂ᶵჾ㢮ࡢ⥔ᣢ⟶⌮࡜฼⏝⪅࡬ࡢᥦ౪࡞࡝ࠊᚑ
᮶ࡢᶵ⬟ࢆ⏕࠿ࡋࡓᨭ᥼஦ᴗࢆ㐍ࡵ࡚࠸ࡲࡍࠋࡲࡓࠊᙜࢭࣥࢱ࣮ࡢ඲ᩍဨࡀ኱Ꮫ㝔⌮Ꮫ◊✲㝔⏕≀⛉Ꮫ㒊㛛࡟ᡤᒓࡋࠊࡑࡢᩍ⫱◊✲࡜ᙜࢭࣥࢱ࣮ࡢ㐠Ⴀࢆවᢸࡋ࡚⾜ࡗ࡚࠸ࡲࡍࠋᘓ≀⮬యࡣᪧ᪋タࢆ౑⏝ࡋ࡚࠸ࡿࡓࡵࠊᐇ㦂⏕≀ඹྠ฼⏝㒊㛛ࡣࢭࣥࢱ࣮ᮾᲷࠊࡑࡢ௚ࡢ  㒊㛛ࡣࢭࣥࢱ࣮すᲷࡢ  ࢝ᡤ࡟ศ࠿ࢀ࡚◊✲ᨭ᥼άືࢆ⾜ࡗ࡚࠸ࡲࡍࡀࠊ௒ᚋࡢࡼࡾຠ⋡ⓗ࡞ࢭࣥࢱ࣮㐠Ⴀࡢࡓࡵ࡟ࠊ୍య໬ࡋࡓᘓ≀࡜ᶵ⬟ࢆᵓ⠏ࡋ࡚⾜ࡁࡓ࠸࡜࠸࠺ᑗ᮶ᵓ᝿ࡀ࠶ࡾࡲࡍࠋ
 ࡉࡽ࡟ࠊᩥ㒊⛉Ꮫ┬ࡢࢧ࣏࣮ࢺ࡟ࡼࡾࠊ᪥ᮏ඲ᅜࡢ◊✲ᶵ㛵࡟ಖ⟶ࡉࢀ࡚࠸ࡿ㈗㔜࡞⏕≀㑇ఏ㈨※ࢆ⅏ᐖ࡟ࡼࡿᦆኻ࠿ࡽᏲࡿࡓࡵࠊ኱Ꮫ㐃ᦠࣂ࢖࢜ࣂࢵࢡ࢔ࢵࣉࣉࣟࢪ࢙ࢡࢺ㸦,%%3㸧ࡀ㐍ࡵࡽࢀ࡚࠾ࡾࠊᙜࢭࣥࢱ࣮ࡣ඲ᅜ࡟㸵ࡘ࠶ࡿ኱Ꮫࢧࢸࣛ࢖ࢺᣐⅬࡢ㸯ࡘ࡜ࡋ࡚ࡶࡑࡢ㔜せ࡞ᙺ๭ࢆᢸࡗ࡚࠸ࡲࡍࠋᖹᡂ  ᖺ  ᭶࡟㉳ࡁࡓᮾ᪥ᮏ኱㟈⅏࡟ࡼࡾࠊከࡃࡢ◊✲ᶵ㛵࡛⏕≀ᮦᩱ࣭㑇ఏ㈨※ࡀኻࢃࢀ◊✲ࡢ⥅⥆࡟ᨭ㞀ࢆࡁࡓࡋࡲࡋࡓࠋ,%%3 ࡣ෌ࡧࡇࡢࡼ࠺࡞ࡇ
࡜ࡀ㉳ࡇࡿ࡜ᡃࡀᅜࡢᅜ㝿ⓗ➇தຊ࡟ࡶᝏᙳ㡪ࢆ୚࠼࠿ࡡ࡞࠸࡜ࡢ༴᝹࠿ࡽࠊឡ▱┴ᒸᓮᕷࡢᇶ♏⏕≀Ꮫ◊✲ᡤࢆ୰᰾ᣐⅬ࡜ࡋ࡚㛤ጞࡉࢀࡓࣉࣟࢪ࢙ࢡࢺ࡛ࡍࠋ ࡲࡓࠊࡈᢎ▱ࡢࡼ࠺࡟᫖ᖺ  ᭶  ᪥ࡢ໭ᾏ㐨⫹᣺ᮾ㒊ᆅ㟈࡟ࡼࡾᮏᏛ࡛ࡶ኱つᶍ೵㟁ࡀⓎ⏕ࡋࠊ኱ࡁ࡞⿕ᐖࢆࡶࡓࡽࡋࡲࡋࡓࠋᙜࢭࣥࢱ࣮࡛ࡣࠊࡇࡢࡼ࠺࡞኱つᶍ⮬↛⅏ᐖ࡟ᑐࡍࡿ
Ᏻᐃࡋࡓᩍ⫱◊✲⎔ቃࢆᣢ⥆ⓗ࡟ᥦ౪ࡍࡿࡓࡵࡢᨭ᥼⤌⧊࡜ࡋ࡚ࠊࡑࡢ౯್ࡀ୍ᒙ㧗ࡲࡗ࡚ࡁ࡚࠸ࡲࡍࠋ ௒ᚋࡶᩍ⫱◊✲ࡢඹྠ฼⏝᪋タ࡜ࡋ࡚ࠊ฼⏝౯್ࡢ㧗࠸ᨭ᥼ࡢᥦ౪࡟ດࡵ࡚⾜ࡁࡲࡍࡢ࡛ࠊ
ࡈ⌮ゎ࡜ࡈ༠ຊࡢ⛬ࠊࡼࢁࡋࡃ࠾㢪࠸⏦ࡋୖࡆࡲࡍࠋ                 ௧࿴ඖᖺ  ᭶                 ኱Ꮫ㝔⌮Ꮫ◊✲㝔                      㝃ᒓࢤࣀ࣒ࢲ࢖ࢼ࣑ࢡࢫ◊✲ࢭࣥࢱ࣮㛗                                   ᒣཱྀ ῟஧㸦ᖹᡂ  ᖺ  ᭶  ᪥ࡼࡾ╔௵㸧

┠ ḟ

・

２０１９年度概要の発行にあたって・センター教職員・運営委員一覧、平成 30 年度活動報告

Ｐ １

・

実験生物共同利用部門運営状況

Ｐ ３・

遺伝子実験共同利用部門・動物染色体共同利用部門運営状況

Ｐ ３１

・

添付資料

Ｐ ４３

䝀䝜䝮䝎䜲䝘䝭䜽䝇◊✲䝉䞁䝍䞊䚷ᩍ⫋ဨ䝉䞁䝍䞊㛗䚷
䚷ᐇ㦂⏕≀ඹྠ฼⏝㒊㛛

ᩍ䚷㻌ᤵ䚷

ቑ⏣䚷㝯୍

෸ᩍᤵ䚷䚷
෸ᩍᤵ䚷䚷
ຓ㻌䚷ᡭ

໭⏣䚷୍༤
ຍ⸨䚷ᚭ
ฟཱྀ䚷ၿ⾜

ᩍ㻌䚷ᤵ
ຓ㻌䚷ᩍ

ቑ⏣䚷㝯୍
す⏣䚷⩏᠇

ຓ㻌䚷ᩍ

ྜྷ⏣䚷㑳ஓ

ᢏ⾡ᑓ㛛⫋ဨ
◊✲ᨭ᥼᥎㐍ဨ஦ົ⿵ຓဨᑠ㔪䚷ᕸᐇᏊ㻌䠄ᮾᲷ䠅ᑠᆏ䚷䛒䜖䜏㻌䠄すᲷ䠅ᒸᮧ䚷ᰤ⌮䚷㻌㻌䠄すᲷ䠅㻌㻌㑇ఏᏊᐇ㦂ඹྠ฼⏝㒊㛛㻌㻌㻌㻌㻌ື≀ᰁⰍయඹྠ฼⏝㒊㛛㻌㻌㻌㻌㻌䝉䞁䝍䞊⫋ဨ䝀䝜䝮䝎䜲䝘䝭䜽䝇◊✲䝉䞁䝍䞊㐠Ⴀጤဨ఍ጤဨ㛗 㻌㻌㻌㻌ᇼཱྀ䚷೺㞝㻌㻌㻌㻌ቑ⏣䚷㝯୍䚷㻌㻌㻌㻌ຍ⸨䚷ᚭ㻌㻌㻌㻌໭⏣䚷୍༤㻌㻌㻌㻌ຍ⸨䚷ᩔஅ㻌㻌㻌㻌㯮ᒾ䚷㯞㔛㻌㻌㻌㻌ᑠட䚷୍ᘯ㻌㻌㻌㻌ỈἼ䚷ㄔ㻌㻌㻌㻌ᒣཱྀ䚷῟஧㻌㻌㻌㻌㈗ᓥ䚷♸἞㻌㻌㻌㻌ᕝཎ䚷Ꮫᩍ㻌䚷ᤵ㻌㻌㻌㻌㻌㻌⌮Ꮫ◊✲㝔ᩍ㻌䚷ᤵ㻌㻌㻌㻌㻌㻌⌮Ꮫ◊✲㝔෸ᩍᤵ㻌㻌㻌㻌㻌㻌⌮Ꮫ◊✲㝔෸ᩍᤵ㻌㻌㻌㻌㻌㻌⌮Ꮫ◊✲㝔ᩍ㻌䚷ᤵ㻌㻌㻌㻌㻌㻌⌮Ꮫ◊✲㝔ᩍ㻌䚷ᤵ㻌㻌㻌㻌㻌㻌⌮Ꮫ◊✲㝔ᩍ㻌䚷ᤵ㻌㻌㻌㻌㻌㻌⌮Ꮫ◊✲㝔ᩍ㻌䚷ᤵ㻌㻌㻌㻌㻌㻌⌮Ꮫ◊✲㝔ᩍ㻌䚷ᤵ㻌㻌㻌㻌㻌㻌⌮Ꮫ◊✲㝔ᩍ㻌䚷ᤵ㻌㻌㻌㻌㻌㻌㎰Ꮫ◊✲㝔෸ᩍᤵ㻌㻌㻌㻌㻌㻌㎰Ꮫ◊✲㝔ᖹᡂ㻟㻜ᖺᗘ㻌㻌άືሗ࿌䞉㻌ᴫせ㻌㻞㻜㻝㻤㻌䠄㻌ᖺḟሗ࿌㻌䠅㻌㻌Ⓨ⾜㻌䠄㻌㻤᭶㻌䠅䞉㻌㐠Ⴀጤဨ఍䠄ᣢ䛱ᅇ䜚䠅㛤ദ㻌䠄㻌㻝㻝᭶㻌䠅䞉㻌ᐇ㦂⏕≀ඹྠ฼⏝㒊㛛㻌᪂つ฼⏝⪅ㅮ⩦఍㛤ദ㻌䠄㻌㻠᭶䞉㻡᭶䞉㻢᭶䞉㻝㻜᭶䞉㻝᭶䞉㻞᭶䚷ィ㻣ᅇ㻌䠅

−1−

ᮾᲷ

ᐇ㦂⏕≀ඹྠ฼⏝㒊㛛

−3−

ᐇ 㦂⏕
⏕ ≀ඹ
ඹ ྠ฼⏝㒊㛛ᙜ㒊㛛ࡣࠊ࣐࢘ࢫࠊࣛࢵࢺࠊ࢘ࢧࢠࠊࣔࣝࣔࢵࢺࡢ့ஙື≀࡟ຍ࠼ࠊ࢟ࣥ࢝ࢳࣙ࢘
ࡸ࢘ࢬࣛ➼ࡢ㫽㢮ࠊ࣓ࢲ࢝➼ࡢ㨶㢮ࠊ࢖࢝࡜࠸ࡗࡓᾏ⏘ࡢ↓⬨᳝ື≀ࠊࢥ࢜ࣟࢠࡸࢦ
࢟ࣈࣜ➼ࡢ᪻⹸ࠊࡲࡓࢩࣟ࢖ࢾࢼࢬࢼ➼ࡢᐇ㦂᳜≀࠿ࡽࠊ㔝እ᥇ྲྀࡋ࡚ࡁࡓ᳜≀ࠊࡉࡽ࡟ᶞᮌ࡟⮳ࡿࡲ࡛ࠊᵝࠎ࡞⏕≀✀ࢆ㣫⫱࣭᱂ᇵ࡛ࡁࡿࡼ࠺ᩚഛࢆ⾜ࡗ࡚࠸ࡿඹྠ฼⏝᪋タ࡛ࡍࠋ㏆ᖺࡣࠊ࣐࢘ࢫࠊࣛࢵࢺࡢࡳ࡞ࡽࡎࠊࢥ࢜ࣟࢠ࡟࠾࠸࡚ࡶ㑇ఏᏊᨵኚື
≀ࡀ㣫㣴ࡉࢀ࡚࠾ࡾࠊἲ࡟‽ᣐࡋࡓ㣫⫱タഛࡢᩚഛ࡟ࡶᚰࡀࡅ࡚࠾ࡾࡲࡍࠋࡇࢀࡽࡢ
ඹྠ฼⏝࡟ຍ࠼ࠊ࣐࢘ࢫ࡜ࣛࢵࢺࡢ㏆஺⣔ࡢ⥔ᣢ஦ᴗࡶ⾜ࡗ࡚࠾ࡾࡲࡍࠋ
ᮏᖺᗘ࡟࠾ࡁࡲࡋ࡚ࡣࠊᖹᡂ  ᖺ  ᭶  ᪥࡟Ⓨ⏕ࡋࡓ໭ᾏ㐨⫹᣺ᮾ㒊ᆅ㟈ࡀ≉グࡉࢀࡲࡍࠋᆅ㟈ࡢᦂࢀ࡟క࠺⿕ᐖࡣ࡯࡜ࢇ࡝࡞࠿ࡗࡓࡶࡢࡢࠊ㛗ᮇ࡟⥆࠸ࡓ೵㟁࡟ࡣᝎࡲࡉࢀࡲࡋࡓࠋᖾ࠸ࠊ⮬ᐙⓎ㟁ᶵࡀసືࡋࠊ㈗㔜࡞ᐇ㦂⏕≀ࡸࣇ࣮ࣜࢨ࣮ෆࡢ᳨యࢆ
Ᏺࡿࡇ࡜ࡀ࡛ࡁࡲࡋࡓࠋ⅏ᐖ᫬࡟ࡣࠊ◊✲ࣜࢯ࣮ࢫࢆᏲࡿ࡭ࡃࠊ᪥㡭࠿ࡽ⮬ᐙⓎ㟁ᶵࢆྵࡴࣂࢵࢡ࢔ࢵࣉయไࢆ୓඲࡟ᩚഛࡋ࡚࠾ࡃᚲせᛶࡀ③ឤࡉࢀࡲࡋࡓࠋ
ࡲࡓࠊᮏ᪋タࡢ❹ᕤࡣ᫛࿴  ᖺ࡛࠶ࡾࠊࡍ࡛࡟⠏  ᖺࢆ⤒㐣ࡋ࡚࠸ࡿࡇ࡜࠿ࡽࠊ
⥅⥆ⓗ࡟ᑠࡉ࡞ಟ⧋ᕤ஦ࡀᚲせ࡜࡞ࡗ࡚࠸ࡿ࡜ࡇࢁ࡛ࡍࠋᖾ࠸࡟ࡶᖹᡂ  ᖺᗘ࡟࠾ࡁ
ࡲࡋ࡚ࠊ㛵ಀྛᡤࡢࡈ༠ຊ࡟ࡼࡾࠊᬮᡣ⏝ࡢỈ⟶ࡢ୍㒊ࢆ᭦᪂ࡍࡿࡇ࡜ࡀ࡛ࡁࡲࡋ

ࡓࠋ⌮Ꮫ◊✲㝔㛗ࢆࡣࡌࡵࠊ⌮Ꮫ࣭⏕࿨⛉Ꮫ஦ົ㒊஦ົㄢࡢⓙᵝ࡟ࡣࠊࡈᨭ᥼ࢆ࠸ࡓ
ࡔࡁࡲࡋࡓࡇ࡜࡟ឤㅰࢆ⏦ࡋୖࡆࡲࡍࠋ
௒ᚋ࡜ࡶ◊✲⎔ቃࡢྥୖ࡜⥔ᣢࢆ┠ᣦࡋ࡚ࠊ◊✲ཬࡧᐇ㦂ࡢሙࢆᥦ౪ࡋ࡚࠸ࡃດຊ
ࢆ⥆ࡅ࡚ࡲ࠸ࡾࡲࡍࠋ



ᖹ ᡂ㸱
㸱 㸮ᖺ
ᖺ ᗘ ᐇ 㦂⏕⏕ ≀ඹඹ ྠ฼⏝㒊㛛 ฼ ⏝≧ἣ ᡤ ᒓ ฼ ⏝⪅ᩘᩘ኱Ꮫ㝔ᩥᏛ◊✲⛉㸯኱Ꮫ㝔་Ꮫ◊✲㝔㸯኱Ꮫ㝔ᆅ⌫⎔ቃ⛉Ꮫ◊✲㝔㸱኱Ꮫ㝔⌮Ꮫ◊✲㝔㸯㸶኱Ꮫ㝔㎰Ꮫ◊✲㝔㸰኱Ꮫ㝔ᕤᏛ◊✲㝔㸰㟁Ꮚ⛉Ꮫ◊✲ᡤ㸯໭᪉⏕≀ᅪࣇ࢕࣮ࣝࢻ⛉Ꮫࢭࣥࢱ࣮

㸯⥲ྜ༤≀㤋

㸯ᩍ⫱Ꮫ◊✲㝔

㸯

ィ

㸱㸯 



−4−

ᖹᡂ㻟㻜ᖺᗘ฼⏝⪅䛾ෆヂ㟁Ꮚ◊㻘㻌㻝

໭᪉䢈䡤䡬䢕䢀䢚㻘㻌㻝

ᩍ⫱㻘㻌㻝
ᩥᏛ㻘㻌㻝

ᾏỈ㻘㻣
့ ங 㢮 䞉㫽㢮㻘 㻝 㻜༤≀㤋㻘㻌㻝་Ꮫ㻘㻌㻝⣧Ỉ㻘㻠⌮Ꮫ㻘㻌㻝㻤

ᕤᏛ㻘㻌㻞

᪻⹸㻘㻡

㎰Ꮫ㻘㻌㻞

Ỉ⏕ື≀㻘㻞

ᆅ⌫⎔ቃ㻘㻌㻟
฼⏝⏦ㄳ⪅ᡤᒓ㒊ᒁ
42

H23

41

H24

38

H25

᳜≀㻘㻟

ᢅ䛳䛶䛔䜛⏕≀ཬ䜃Ỉ➼฼⏝䛾⏦ㄳᩘ
41
36

H26

H27

34

H28

30

31

H29

H30

฼⏝⏦ㄳᩘ᥎⛣⣔⤫ಖᏑ䝸䝇䝖
䝬䜴䝇䠄㻤⣔⤫䠅⣔⤫ྡ౪⤥ᩘ䝷䝑䝖䠄㻝㻝⣔⤫䠅౪⤥ඛ⣔⤫ྡ㻮㻭㻸㻮㻛㼏

㻭㻯㻵㻮㻭㻸㻮㻛㼏㻭㼚

㻭㻯㻵㻔㻿㻕

㻮㻷㻴

㻭㻸㻮

㻯㻮㻭

㻮㻺

㻯㻮㻭㻛㻴㻙㼀㻢

㻷㼅㻺㻯㻡㻣㻮㻸㻛㻢

㻸㻱㻭

㻯㻟㻴

㻸㻱㻶

㻸㼀㻛㻿㼢

㻺㻵㻳㻙䊢

౪⤥ᩘ

౪⤥ඛ

䗠㻣䠈䗢㻤

ி㒔኱Ꮫ䞉㻺㻮㻾㻼䛂䝷䝑䝖䛃

㻿㻰㻶
㼀㻻
㼃㻷㻭㻴

−5−

䚷

ᖹᡂ㻟㻜ᖺᗘ㻌ᐇ㦂⏕≀ඹྠ฼⏝㒊㛛䚷ಖᏲႠ⧋䝸䝇䝖

ᖹᡂ㻟㻜ᖺ

ᖹᡂ㻟㻝ᖺ

㻠᭶

ᐃᮇΎᤲ㻔ẖ᭶䠅⎔ቃ⾨⏕⟶⌮Ⅼ᳨䠄ẖ᭶䠅ᶞᮌ෤ᅖ䛔እ䛧ᴗົ㻡᭶

㻝㝵㏻㊰ㄏᑟⅉྲྀ᭰ᕤ஦ᐊᒇ᰿䝁䞊䜻䞁䜾ᕤ஦㻤᭶㻝㝵Ὑίᐊᾘẘჾᛶ⬟᳨ᰝᩚഛ䠄䜸䞊䝖䜽䝺䞊䝤䠅㻝㝵タഛᐊෆ⵨Ẽ㓄⟶ಟ⌮㠀ᖖ⏝Ⓨ㟁ᶵ⡆᫆Ⅼ᳨㻥᭶㻝㝵タഛᐊෆ⵨Ẽ㓄⟶ಟ⌮㻝㻜᭶㻞㝵᳜≀ᇵ㣴ᐊ㻔㻝㻕㻔㻟㻕᥮Ẽᡪྲྀ᭰㻝㻝᭶㻝㝵ศ㟁┙䝤䝺䞊䜹䞊ྲྀ᭰ᕤ஦㻝㻞᭶㻱㻙㻞㻙㻜㻞䚸㻱㻙㻞㻙㻜㻡ᐊᡬ⿵ಟᕤ஦
ᶞᮌ෤ᅖ䛔ᴗົ

㻝᭶㻞㝵᳜≀ᇵ㣴ᐊ㻔㻞㻕ཬ䜃ᐇ㦂ᐊ㻔㻥㻕᥮Ẽᡪྲྀ᭰䝗䝷䝣䝖䝏䝱䞁䝞䞊᤼㢼ᶵಟ⌮㻞᭶↮✺ゎయ➼䛭䛾௚ᕤ஦䖂⌮Ꮫ㒊୰ኸண⟬䛻䛶ᇳ⾜ᖹᡂ㻟㻜ᖺ 㻝㻝᭶ᖹᡂ㻟㻜ᖺ 㻝㻞᭶㻝㝵タഛᐊෆᬮᡣ⏝ Ỉ㓄⟶ᨵಟᕤ஦ᒇୖ᤼Ẽሪ㜵Ỉ⿵ಟᕤ஦−6−ᖹᡂ㻟㻜ᖺᗘ䚷䝀䝜䝮䝎䜲䝘䝭䜽䝇◊✲䝉䞁䝍䞊䚷฼⏝⪅◊✲ㄢ㢟䠄ᐇ㦂⏕≀ඹྠ฼⏝㒊㛛䠅ᡤ㻌㻌ᒓ㒊㻌㻌㛛㻌㻌䞉㻌㻌ศ㻌㻌㔝㻌㻌㻌➼

⫋㻌ྡ

ᩥᏛ◊✲⛉

ே㛫䡸䡹䡿䢍⛉Ꮫᑓᨷ䞉ᚰ⌮䡸䡹䡿䢍⛉Ꮫㅮᗙᩍ㻌ᤵ࿴⏣㻌㻌༤⨾ 䚷䝷䝑䝖䛾㉸㡢ἼⓎኌ䛻ཬ䜌䛩Ỉ㓟໬㻼㻯㻮ཬ䜃㻼㻮㻰㻱䛾ᙳ㡪་Ꮫ◊✲㝔་ṑᏛ⥲ྜ◊✲Ჷ䞉୰ኸ◊✲㒊㛛ຓ㻌ᩍᒣ㔝㎶㻌㈗ಙ 㻌㻌⚄⤒⣔䛻䛚䛡䜛᝟ሗ䜻䝱䝸䜰䜢᫂䜙䛛䛻䛩䜛ᆅ⌫⎔ቃ⛉Ꮫ
◊✲㝔ᆅ⌫⎔ቃ⛉Ꮫ
◊✲㝔ᆅ⌫⎔ቃ⛉Ꮫ
◊✲㝔⤫ྜ⎔ቃ⛉Ꮫ䞉⮬↛⎔ቃಖ඲ᩍ㻌ᤵ㟢ᓮ㻌㻌ྐᮁ 䚷ⅆᒣ䛻䛚䛡䜛ᇙᅵ✀Ꮚ㞟ᅋ䛾ᐃ㔞䛚䜘䜃⏕⫱ᐇ㦂⤫ྜ◊✲⛉Ꮫ䞉⎔ቃ㐺ᛂ⛉Ꮫᩍ㻌ᤵἈ㔝㻌㻌㱟ᩥ 䚷ᾏ⏘⸴㢮䛾஧ḟ௦ㅰ⏘≀䛻㛵䛩䜛◊✲⎔ቃ⏕≀⛉Ꮫ䞉㝣ᇦ⏕ែᏛ

෸ᩍᤵ

ᕤ⸨䚷ᓅ

⌮Ꮫ◊✲㝔

≀⌮Ꮫ䞉㔞Ꮚ≀⌮Ꮫ෸ᩍᤵᯇỌ㻌㻌ᝅ᫂ 䚷᭷ᶵᑟయ䛻䛚䛡䜛≀ᛶ⌮Ꮫ◊✲㝔≀⌮Ꮫ䞉㟁Ꮚ≀ᛶ≀⌮Ꮫᩍ㻌ᤵἙᮏ㻌㻌඘ྖ 䚷ศᏊᛶᑟయ䛾ྜᡂ⌮Ꮫ◊✲㝔⏕≀⛉Ꮫ䞉ᙧែᶵ⬟Ꮫᩍ㻌ᤵᒣཱྀ㻌㻌῟஧ 䚷㧗➼᳜≀䛾ᶵ⬟Ⓨ⌧䞉⎔ቃᛂ⟅ไᚚᶵᵓ⌮Ꮫ◊✲㝔⏕≀⛉Ꮫ䞉⾜ື⚄⤒⏕≀Ꮫ

ᩍ㻌ᤵ

ỈἼ㻌㻌ㄔ

⌮Ꮫ◊✲㝔

⏕≀⛉Ꮫ䞉⾜ື⚄⤒⏕≀Ꮫᩍ㻌ᤵᑠᕝ㻌㻌ᏹே 䚷䝣䝍䝩䝅䝁䜸䝻䜼䛾⚄⤒⾜ືᏛⓗ◊✲⌮Ꮫ◊✲㝔⏕≀⛉Ꮫ䞉⾜ື⚄⤒⏕≀Ꮫ෸ᩍᤵ໭⏣㻌㻌୍༤ 䚷᪂䛯䛺⑓ែ䝰䝕䝹ື≀䛾సฟ䛸ཎᅉ㑇ఏᏊ䛾ྠᐃ⌮Ꮫ◊✲㝔⏕≀⛉Ꮫ䞉⾜ື⚄⤒⏕≀Ꮫ෸ᩍᤵ࿴ከ㻌㻌࿴ᏹ 䚷㬆⚺㢮䜢⏝䛔䛯ⓎኌᏛ⩦䞉⏕ᡂ䛸䛭䛾⬻ෆศᏊᶵᵓ⌮Ꮫ◊✲㝔⏕≀⛉Ꮫ䞉⾜ື⚄⤒⏕≀Ꮫ෸ᩍᤵ⏣୰㻌㻌ᬸ᫂ 䚷䝠䝯䜲䜹䛾⥅௦㣫⫱ᢏ⾡䛾㛤Ⓨ⌮Ꮫ◊✲㝔⏕≀⛉Ꮫ䞉⏕ṪⓎ⏕⏕≀Ꮫᩍ㻌ᤵᒣୗ㻌㻌ṇව 䚷⬨᳝ື≀䛻䛚䛡䜛⏕Ṫ⣽⬊ᙧᡂ䛾ไᚚᶵᵓ⌮Ꮫ◊✲㝔⏕≀⛉Ꮫ䞉⏕ṪⓎ⏕⏕≀Ꮫᩍ㻌ᤵ㯮ᒾ䚷㯞㔛 㻌㻌့ங㢮䛚䜘䜃㫽㢮䛻䛚䛡䜛ᛶỴᐃ䞉ཷ⢭䛻㛵䜟䜛ศᏊ䝯䜹䝙䝈䝮䛾ゎ᫂⌮Ꮫ◊✲㝔⏕≀⛉Ꮫ䞉⏕ṪⓎ⏕⏕≀Ꮫ

෸ᩍᤵ

ᮌᮧ㻌㻌ᩔ

⌮Ꮫ◊✲㝔

⏕≀⛉Ꮫ䞉⏕ṪⓎ⏕⏕≀Ꮫ෸ᩍᤵᑠ㇂㻌㻌཭ஓ 䚷༸ẕ⣽⬊ᙧᡂ䛸ึᮇⓎ⏕䛾ศᏊᶵᵓゎᯒ⌮Ꮫ◊✲㝔⏕≀⛉Ꮫ䞉⏕ṪⓎ⏕⏕≀Ꮫ෸ᩍᤵⲶཎ㻌㻌ඞ┈ 䚷⬨᳝ື≀䛾⏕Ṫᶵ⬟䛻㛵䛩䜛◊✲⌮Ꮫ◊✲㝔⏕≀⛉Ꮫ䞉ከᵝᛶ⏕≀Ꮫᩍ㻌ᤵᑠட㻌㻌୍ᘯ 䚷⸴㢮䛾⣔⤫㐍໬Ꮫ⌮Ꮫ◊✲㝔⏕≀⛉Ꮫ䞉ከᵝᛶ⏕≀Ꮫ

෸ᩍᤵ

᯳ཎ㻌㻌ᏹ

⌮Ꮫ◊✲㝔

⏕≀⛉Ꮫ䞉ከᵝᛶ⏕≀Ꮫ෸ᩍᤵ䜶䝺䝘䞉䝣䜷䝹䝖䜳䝘䝖⌮Ꮫ◊✲㝔⏕≀⛉Ꮫ䞉ከᵝᛶ⏕≀Ꮫ

෸ᩍᤵ

ຍ⸨㻌㻌ᚭ

⌮Ꮫ◊✲㝔

⏕≀⛉Ꮫ䞉ከᵝᛶ⏕≀Ꮫㅮᖌゅ஭䚷ᩗ▱ 㻌㻌ᑠᆺ⏥Ẇ㢮䛾ᛶ䞉⏕Ṫᵝᘧ䛻㛵䛩䜛◊✲㎰Ꮫ◊✲㝔ᇶ┙◊✲䞉⏕≀㈨※⛉Ꮫᩍ㻌ᤵ⛅ඖ㻌㻌ಙ୍ 䚷䝃䝑䝫䝻䝣䜻䝞䝑䝍䛾㓄അ⾜ື䛸⏕Ṫ㝸㞳ᶵᵓ䛻㛵䛩䜛◊✲㎰Ꮫ◊✲㝔ᇶ┙◊✲䞉␆⏘⛉Ꮫ

෸ᩍᤵ

ᕝཎ㻌㻌Ꮫ

䚷䝬䜴䝇⬇యእ᧯స䛻䜘䜛ಶయⓎ⏕䛻㛵䛩䜛◊✲ᕤᏛ◊✲㝔ᛂ⏝≀⌮Ꮫ䞉㔞Ꮚ≀ᛶᕤᏛ

ᩍ㻌ᤵ

୹⏣㻌㻌⪽

䚷పḟඖᑟయ䛾㟁Ꮚ≧ែ䛾◊✲ᕤᏛ◊✲㝔䡰䢄䢕䡳䢚䡬⎔ቃ䡸䡹䡿䢍䞉ᛂ⏝䡰䢄䢕䡳䢚䡬䡸䡹䡿䢍 ≉௵ᩍᤵẶ㻌㻌ྡ◊㻌㻌✲㻌㻌ㄢ㻌㻌㢟㻌㻌ྡ䚷䝬䝒䝶䜲䜾䝃ᒓ᳜≀䛾㏦⢊⏕ែᏛ䛻㛵䛩䜛◊✲ᐃ䚷᪻⹸䛾Ꮫ⩦䛾⚄⤒ᶵᵓ䚷့ங㢮䛾⏕Ṫ䛻䛛䛛䜟䜛䝀䝜䝮ᶵ⬟䛻㛵䛩䜛◊✲䚷ᾏ⏘↓⬨᳝ື≀䛾⣔⤫ศ㢮Ꮫⓗ◊✲䚷䜴䝭䜴䝅㢮䛻䛚䛡䜛ᤕ㣗⪅䛻ᑐ䛩䜛໬Ꮫ㜵ᚚ䛾◊✲䚷䝅䝵䜴䝆䝵䜴䝞䜶䛚䜘䜃᳜㣗ᛶ䝔䞁䝖䜴䛾㐍໬Ꮫⓗ◊✲㏆ஂ㻌㻌Ṋ⨾ 䚷⇞ᩱ㟁ụෆỈศᣲືཬ䜃෾⤖⌧㇟䛾ྍど໬ほᐹ㟁Ꮚ⛉Ꮫ◊✲ᡤ 㝃ᒓ♫఍๰㐀ᩘᏛ◊✲䡺䢙䡼䡬䞉ே㛫ᩘ⌮◊✲෸ᩍᤵ㟷἟㻌㻌ோᚿ 䚷䜽䝻䝁䜸䝻䜼䛻䛚䛡䜛㐺ᛂ⾜ື䛾ᐇ᫬㛫ᛶ䜢ไᚚ䛩䜛⚄⤒ᶵᵓ䛾◊✲໭᪉⏕≀ᅪ䝣䜱䞊
⏕≀ከᵝᛶ㡿ᇦ䞉ᾏ⏘⸴㢮㐺ᛂᶵ⬟䝹䝗⛉Ꮫ䝉䞁䝍䞊

෸ᩍᤵᅄ䝒಴㻌඾⁠ 䚷䝁䞁䝤㢮䛻䛴䛔䛶䚸ᇵ㣴ಖᏑἲ䛾☜❧䛸✀ⱑసฟᢏ⾡䛾㛤Ⓨㅮ㻌ᖌ㜿㒊㻌㻌๛ྐ 䚷⣚⸴䝋䝌ᒓ䛚䜘䜃㏆⦕ᒓ䛻㛵䛩䜛✀⏕≀Ꮫⓗ◊✲⥲ྜ༤≀㤋◊✲㒊䞉㈨ᩱᇶ♏◊✲⣔ᩍ⫱Ꮫ◊✲㝔

ᩍ⫱Ꮫ䞉೺ᗣయ⫱Ꮫ෸ᩍᤵ ᒣ௰䚷ຬ஧㑻 䚷㻌⏕≀᫬ィ䛻䜘䜛⾜ື䝸䝈䝮䛾ᨭ㓄ᵝᘧ䛸⾜ື䝸䝈䝮䛛䜙୰ᯡ᫬ィ䜈䛾䝣䜱䞊䝗䝞䝑䜽ᶵᵓ䛾ゎ᫂

−7−⏕ࡌࢀࡤ௰㛫࡜ពᛮ␯㏻ࢆᅗࡿࡇ࡜ࡀ࡛ࡁ࡞ࡃ࡞
ࡾࠊಶయࡢ⏕Ꮡࡸ⩌ࢀࡢ⥔ᣢࡀ⬣ጾ࡟ࡉࡽࡉࢀࡿ
࡜ண᝿ࡉࢀࡿࠋ
᪉ἲ㸸 ዷፎࣛࢵࢺࢆ  ⩌࡟ศࡅࠊ࢔ࣝࢥ࣮ࣝ㸦࢚኱Ꮫ㝔ᩥᏛ◊✲⛉
ࢳࣝ࢔ࣝࢥ࣮ࣝ㸧ࢆΰධࡋࡓ㣧ᩱỈࢆ୚࠼ࡓࠋ࢔
ே㛫ࢩࢫࢸ࣒⛉Ꮫᑓᨷ ᚰ⌮ࢩࢫࢸ࣒⛉Ꮫㅮᗙࣝࢥ࣮ࣝ⃰ᗘࡣ㧗⃰ᗘ⩌ 㸣ࠊప⃰ᗘ⩌ 㸣ࠊ⤫࿴⏣ ༤⨾ไ⩌ 㸣ࠊᢞ୚ᮇ㛫ࡣዷፎ  ᪥┠࠿ࡽ  ᪥┠࡛࠶ࡗࡓࠋ࡞࠾࢔ࣝࢥ࣮ࣝ⃰ᗘࡣᚎࠎ࡟㧗ࡃࡋࡓࠋ⏕◊✲ࢸ࣮࣐㸯ᚋ  ᪥┠࡟௘ࣛࢵࢺࢆ㞳ஙࡉࡏࠊಶูࢣ࣮ࢪ࡛㣫⮯⣲⣔㞴⇞๣ࢹ࢝ࣈࣟࣔࢴࣇ࢙ࢽ࣮࢚࣮ࣝࢸࣝ
㸦%'(㸧㸧ࡀࣛࢵࢺࡢ㉸㡢Ἴࢥ࣑ࣗࢽࢣ࣮ࢩࣙࣥ ⫱ࡋࡓࠋ⏕ᚋ  ᪥┠࠿ࡽ  ᪥㛫ࠊྠᛶ␗⭡ࡢࣛࢵࢺ  ༉ࢆࣉࣛࢫࢳࢵࢡ〇ࣂࢣࢶ࡟୍⥴࡟ධࢀࠊ
ࡢⓎ㐩࡟ཬࡰࡍᙳ㡪ࡢ◊✲⮯⣲⣔㞴⇞๣ %'( ࡣྍ⇞≀ࢆ⇞࠼࡟ࡃࡃࡋࠊ ศ㛫ࡢ㥆໬ࡢᚋ࡟  ศ㛫ࡢ㜚த㐟ࡧࢆ⾜ࡗࡓࠋ㜚த㐟ࡧࡢ㛫ࠊ㉸㡢ἼⓎኌࢆグ㘓ࡋࡓࠋᘏ↝ࢆ㜵ࡄࡇ࡜ࡀ࡛ࡁࡿࠋࡑࡢ཯㠃ࠊ⏥≧⭢࣍ࣝ⤖ᯝ࡜⪃ᐹ㸸 ᢞ୚⩌ࡣ 㹼N+] ࡢ㉸㡢ἼⓎኌࣔࣥ㜼ᐖ࡞࡝ࡢ᭷ᐖస⏝ࡀᠱᛕࡉࢀ࡚࠸ࡿࠋᮏ◊ࡀ⤫ไ⩌ࡼࡾቑ኱ࡋࡓࠋࡇࢀࡽࡢⓎኌࡣ୙Ᏻࠊᜍ
✲ࡣዷፎࣛࢵࢺ࡟ %'( ࢆᢞ୚ࡋࠊฟ⏕ࡋࡓ௘
ᛧࠊⱞ③࡜࠸ࡗࡓࢿ࢞ࢸ࢕ࣈ࡞ឤ᝟≧ែࢆ཯ᫎࡋ
ࣛࢵࢺࡢ㉸㡢Ἴࢥ࣑ࣗࢽࢣ࣮ࢩࣙࣥ࡬ࡢᙳ㡪ࢆ᥈࡚࠸ࡿࠋࡉࡽ࡟ᢞ୚⩌࡛ࡣ࣏ࢪࢸ࢕ࣈ࡞ឤ᝟≧ែࡿࡇ࡜࡟ࡼࡾࠊ᭷ᐖస⏝ࡢ୍➃ࢆ᫂ࡽ࠿࡟ࡍࡿࡇ
ࢆ཯ᫎࡍࡿ 㹼N+] ࡢ㉸㡢ἼⓎኌࡀῶᑡࡋࡓࠋ
࡜ࢆ┠ⓗ࡜ࡋࡓࠋ
㟷ᖺᮇࡢࣛࢵࢺ࡟࡜ࡗ࡚ࠊ㜚த㐟ࡧࡣᮏ᮶ᴦࡋ࠸
᪉ἲ㸸 ዷፎࣛࢵࢺࢆ  ⩌࡟ศࡅࠊ%'( ࢆᢞࡶࡢ࡛࠶ࡿࠋ࡟ࡶ࠿࠿ࢃࡽࡎࠊᢞ୚⩌࡛ࢿ࢞ࢸ࢕
୚ࡋࡓࠋᢞ୚㔞ࡣ㧗⃰ᗘ⩌ PJNJGD\㸦Q 㸧ࠊࣈ࡞㉸㡢ἼⓎኌࡀቑຍࡋ࣏ࢪࢸ࢕ࣈ࡞㉸㡢ἼⓎኌప⃰ᗘ⩌ PJNJGD\㸦Q 㸧ࠊ⤫ไ⩌ PJNJGD\
ࡀῶᑡࡋࡓࠋࡇࡢࡇ࡜࠿ࡽ⫾ඣᮇࡢ࢔ࣝࢥ࣮ࣝ᭚
㸦Q 㸧࡜ࡋࡓࠋ%'( ࢆ⦎ங JNJGD\ ࡟㟢ࡀࢿ࢞ࢸ࢕ࣈ࡞ឤ᝟ࢆஹ㐍ࡋࠊࣛࢵࢺࡢ♫఍ᛶΰධࡋࠊ⮬⏤ᦤ㣗ࡉࡏࡓࠋᢞ୚ᮇ㛫ࡣዷፎ  ᪥㹼Ⓨ㐩࡟ᙳ㡪ࡍࡿ࡜࠸࠺᪂▱ぢࡀᚓࡽࢀࡓࠋ
ฟ⏘ᚋ  ᪥┠࡛࠶ࡗࡓࠋ⏕ᚋ  ᪥┠࡟௘ࣛࢵࢺ ࢆ㞳ஙࡉࡏࠊಶูࢣ࣮ࢪ࡛㣫⫱ࡋࡓࠋ⏕ᚋ  ᪥┠
࠙ㄽᩥⓎ⾲ࠚ࠿ࡽ  ᪥㛫ࠊྠᛶྠ⭡ࡢࣛࢵࢺ  ༉ࢆࣉࣛࢫࢳࢵ
1. Shahrier M. A. and Wada H. (2018) Effects of perinatal
ࢡ〇ࣂࢣࢶ࡟୍⥴࡟ධࢀࠊ ศ㛫ࡢ㥆໬ࡢᚋ࡟ 
ethanol exposure on acoustic characteristics of ultrasonicศ㛫ࡢ㜚த㐟ࡧࢆ⾜ࡗࡓࠋ㜚த㐟ࡧࡢ㛫ࠊ㉸㡢Ἴ
vocalization in rat pups. Neurotoxicology, 69: 29-36,Ⓨኌࢆグ㘓ࡋࡓࠋ
2018.
⤖ᯝ࡜⪃ᐹ㸸 㹼N+] ࡢ㉸㡢Ἴࡣ୙Ᏻࠊᜍᛧࠊ
2. Wada H. and Qi Y. (2018) Effects of decabromodiphenyl
ⱞ③࡜࠸ࡗࡓࢿ࢞ࢸ࢕ࣈ࡞ឤ᝟≧ែࡢࣛࢵࢺࡀⓎ
ether (BDE-209) on ultrasonic communication in matingኌࡍࡿࠋ␗⭡ࡢ┦ᡭ࡜㜚த㐟ࡧࢆ⾜࠺᫬࡟ฟ⌧ࡍ
behavior of rats. Organohalogen Compounds (in press).
3. Qi Y. and Wada H. (2018) Effects of decabromodiphenylࡿࡀࠊྠ⭡ࡢ┦ᡭ࡜⾜࠺࡜ࡁ࡟ࡣฟ⌧ࡋ࡞࠸ࠋ௒
ether (BDE-209) on ultrasonic communication duringᅇࡣྠ⭡ࡢ┦ᡭ࡜㜚த㐟ࡧࢆ⾜ࡗࡓࡀࠊᢞ୚⩌ࡣfighting of male adult rats. Organohalogen Compounds㹼N+] ࡢ㉸㡢Ἴࢆ┒ࢇ࡟Ⓨኌࡋࡓࠋᢞ୚⩌ࡀ(in press).
ࢿ࢞ࢸ࢕ࣈ࡞ឤ᝟≧ែ࡟࡞ࡗ࡚࠸ࡓࡇ࡜ࢆ♧ࡋ࡚  ࠸ࡿࠋ୍᪉ࠊ⤫ไ⩌࡟ࡣࡇࡢࡼ࠺࡞Ⓨኌࡣ඲ࡃ࡞
࠙Ꮫ఍Ⓨ⾲ࠚ
࠿ࡗࡓࠋ%'( ࡣࢿ࢞ࢸ࢕ࣈ࡞ឤ᝟ࢆஹ㐍ࡍࡿ
1. Wada H. and Qi Y. (2018) Effects of decabromodiphenyl࡜࠸࠺᪂▱ぢࡀᚓࡽࢀࡓࠋ
ether (BDE-209) on ultrasonic communication in◊✲ࢸ࣮࣐㸰mating behavior of rats. Dioxin 2018. Krakow,
࿘⏘ᮇࡢ࢔ࣝࢥ࣮ࣝ᭚㟢ࡀࣛࢵࢺࡢ㉸㡢Ἴࢥ࣑ࣗ
Poland. August 28.
2. Qi Y. and Wada H. (2018) Effects of decabromodiphenylࢽࢣ࣮ࢩࣙࣥࡢⓎ㐩࡟ཬࡰࡍᙳ㡪
ether (BDE-209) on ultrasonic communication during ዷፎ୰ࡸᤵங୰ࡢ㣧㓇ࡀᏊ౪ࡢᡂ㛗࣭Ⓨ㐩ࢆ㜼fighting of male adult rats. Dioxin 2018. Krakow,ᐖࡍࡿࡇ࡜ࡣᗈࡃ▱ࡽࢀ࡚࠸ࡿࠋࡋ࠿ࡋࣄࢺࡀᡂ
Poland. August 27.
⇍ࡍࡿ࡟ࡣ  ᖺ㏆࠸ṓ᭶ࢆせࡋࠊࡑࡢ㛫࡟ᵝࠎ࡞
3. Shahrier M. A. and Wada H. (2018) Acoustic characteristics
せᅉࡀ஺⤡ࡋ࡚࢔ࣝࢥ࣮ࣝࡢᙳ㡪࡞ࡢ࠿ࢃ࠿ࡽ࡞
of ultrasonic vocalizations in rat pups prenatally exposedࡃ࡞ࡿࠋᮏ◊✲࡛ࡣࣛࢵࢺࢆ⏝࠸ࡿࠋࣛࢵࢺࡣ 
to ethanol. International Conference on Alcohol and㐌㛫⛬ᗘ࡛ᡂ⇍ࡋࠊ࢔ࣝࢥ࣮ࣝ௨እࡢせᅉࢆ⤫ไDrugs. Osaka, Japan. October 11.
ࡍࡿࡇ࡜ࡀ࡛ࡁࡿࠋࣛࢵࢺࡣ㉸㡢Ἴࢆ฼⏝ࡋ࡚ࢥ࣑ࣗࢽࢣ࣮ࢩࣙࣥࢆ⾜ࡗ࡚࠾ࡾࠊ㉸㡢Ἴ࡟ኚ␗ࡀ

Мဇ‫إ‬ԓᴾᴾ

−8−኱Ꮫ㝔་Ꮫ◊✲㝔㝔་ṑᏛ⥲ྜ◊✲Ჷ ୰ኸ◊✲㒊㛛ᒣ㔝㎶ ㈗ಙ ࢫࣃ࢖ࢡิࡢ࡝ࡢ⤫ィ㔞ࡀ᝟ሗ࢟ࣕࣜ࢔࠿᫂ࡽ
࠿࡜࡞ࡗ࡚࠸࡞࠸ࠋ᫖ᖺ࡜ྠᵝࠊࡇࡢ᝟ሗ࢟ࣕࣜ
࢔ࡢၥ㢟࡟ྲྀࡾ⤌ࢇࡔࠋ⚄⤒ᅇ㊰⌮ㄽ࡟ࡼࢀࡤ⣲Ꮚࡢฟຊ㛵ᩘࡢ≉ᛶ࡟౫Ꮡࡋࠊ᝟ሗ࢟ࣕࣜ࢔ࡀỴࡲࡿ㸦౛࠼ࡤࠊࢩࢢࣔ࢖ࢻ㛵ᩘ࡛࠶ࢀࡤ UDWHFRGHࠊࢫࢸࢵࣉ㛵ᩘ࡛࠶ࢀࡤ WHPSRUDOFRGH ࡞࡝㸧ࠋࢫࣃ࢖ࢡࡣ⚄⤒⣽⬊ࡀ㐣Ώ≧ែ࡟࠶ࡿ࡜ࡁ࡟⏕ࡌࡿࡓࡵࠊࢫࣃ࢖ࢡ⏕ᡂࡣ㐣ཤࡢ⚄⤒άື࡟౫Ꮡࡍࡿࡇ
࡜ࡀሗ࿌ࡉࢀ࡚࠸ࡿ㸦Segundo et al.Neuroscience
1994, Yamanobe et al. Biosystems 1998㸧ࠋࡲࡓࠊ⚄⤒⣽⬊ࡣࠊᣑᩓ㐣⛬࡛㏆ఝࡉࢀࡿ࢖࢜ࣥࢳࣕࢿࣝࣀ
࢖ࢬࠊࢩࢼࣉࢫᑠ⬊ࡢ⮬Ⓨⓗᨺฟ࡟ࡼࡿࢪࣕࣥࣉ
ࣀ࢖ࢬ࡞࡝ࢆᣢࡘ☜⋡ⓗ࡞ᣲືࢆࡍࡿ⣲Ꮚ࡛ࡶ࠶ࡿࠋࡇࢀࡽࡢࣀ࢖ࢬ࡟ࡼࡾ⚄⤒⣽⬊ࡢᛂ⟅≉ᛶࡀኚࢃࡿࡇ࡜ࡶண᝿ࡉࢀࡿࠋࡑࡇ࡛ࠊྛ⚄⤒⣽⬊࡟࠾࠸࡚㐣Ώᛂ⟅≉ᛶ࡜ࣀ࢖ࢬࡢᙳ㡪ࢆ཯ᫎࡋࡓฟ
ຊ㛵ᩘࡢ≉ᛶࢆᐇ㦂ⓗ࡟ồࡵࡿࡇ࡜ࡀࠊ᝟ሗ࢟ࣕ
ࣜ࢔ࢆゎ᫂ࡍࡿࡓࡵ࡟ᚲせ࡛࠶ࡿ࡜⪃࠼ࡽࢀࡿࠋ
ᐇ㦂࡛⚄⤒⣽⬊ࡢฟຊ㛵ᩘࢆㄪ࡭ࡿ᪉ἲࢆᵓ⠏ࡍࡿࡓࡵࠊ☜⋡ⓗ⚄⤒⣽⬊ࣔࢹࣝࡢ⤫ィⓗ኱ᇦᣲືࢆㄪ࡭ࡿࡇ࡜ࡀ㔜せ࡛࠶ࡿࠋᮏ◊✲࡛ࡣࡇࡢࡼ࠺
࡞⫼ᬒࡢࡶ࡜ࠊୗグࡢ㸱ࡘࡢࢸ࣮࣐࡟㛵ࡋ࡚◊✲ࢆᐇ᪋ࡋࡓࠋ
㸯㸬ࡇࢀࡲ࡛⾜ࡗ࡚ࡁࡓࠊࢩࢼࣉࢫࣀ࢖ࢬ࠾ࡼࡧ
࢖࢜ࣥࢳࣕࢿࣝࣀ࢖ࢬࢆᣢࡘ☜⋡ⓗ᣺ືᏊ㸦⚄⤒⣽⬊ࡢࢲ࢖ࢼ࣑ࢡࢫࡶࡇࡢ᣺ືᏊ࡛㏆ఝࡉࢀࡿሙ
ྜࡀ࠶ࡿ㸧ࡢࢲ࢖ࢼ࣑ࢡࢫ࡟㛵ࡍࡿ◊✲ࢆㄽᩥ࡜ࡋ࡚ฟ∧ࡋࡓࠋㄽᩥ୰࡛♧ࡋࡓ୺せ࡞㡯┠ࡣ௨ୗ
ࡢ㏻ࡾ࡛࠶ࡿࠋ
ձ☜⋡ⓗ᣺ືᏊࡀ୍ពゎࢆᣢࡘࡇ࡜ࢆド᫂ࡋࡓࠋ
ղ☜⋡ⓗ᣺ືᏊࡢ≧ែࡀࠊ࠶ࡿ᫬㛫ᚋࠊ࡝ࡢ≧ែ࡟᥎⛣ࡍࡿ࠿ࢆ⾲ࡍ᥎⛣☜⋡ᐦᗘࡢ₞㏆ᒎ㛤ࡍࡿࡓࡵࠊࡑࡢ๓ᥦ᮲௳࡜࡞ࡿᐦᗘ㛵ᩘࡢ㠀㏥໬ᛶࢆ♧ࡋࡓࠋճ᥎⛣☜⋡ᐦᗘ㛵ᩘࡢ₞㏆ᒎ㛤ࢆィ⟬ࡋࡓࠋࡇࡢ
ࡼ࠺࡟ࡋ࡚ᑟ࠿ࢀࡓ᥎⛣☜⋡ᐦᗘࡢ₞㏆ᒎ㛤ࡢ⢭ᗘࢆࣔࣥࢸ࢝ࣝࣟἲ࡛ィ⟬ࡉࢀࡿ᥎⛣☜⋡ᐦᗘ㛵ᩘ࡜ẚ㍑ࡋ᳨ドࡋࡓࠋմࢪࣕࣥࣉࡢᩘࡀ㸯ᅇࡲ࡛࡛࠶ࡿࡀࠊᩘ್✚ศ࡟ᡂຌࡋࠊ᥎⛣☜⋡ᐦᗘࡢ≉ᛶࢆㄪ࡭ࡓࠋ
ᮏᖺᗘࡣࡉࡽ࡟ࠊࢪࣕࣥࣉ㸯ᅇࡲ࡛ࡋ࠿ィ⟬࡛ࡁ࡞࠿ࡗࡓ᥎⛣☜⋡ᐦᗘ㛵ᩘࢆࢪࣕࣥࣉࡀ」ᩘᅇධ
ࡿࡲ࡛ᣑᙇ࡛ࡁࡿࡼ࠺ᕤኵࡋࠊࡑࡢᕤኵ࡟ࡼࡾ࡝
ࡢࡄࡽ࠸ࡢ⢭ᗘ࡛᥎⛣☜⋡ᐦᗘ㛵ᩘࢆィ⟬࡛ࡁࡿ࠿᳨ウࡋࡓࠋ
㸰㸬ࡇࢀࡲ࡛ࡢ◊✲ᡂᯝ࡟ࡼࡾࠊ☜⋡ⓗ⚄⤒⣽⬊ࣔࢹࣝࡢ᣺ࡿ⯙࠸ࡣ⥺ᙧస⏝⣲㸦↓㝈ḟඖࡢ⾜ิ㸧࡛グ㏙࡛ࡁࡿࡇ࡜ࡀศ࠿ࡗ࡚ࡁࡓࠋᮏᖺᗘࡣࠊࢩ
ࢼࣉࢫࣀ࢖ࢬࠊ࢖࢜ࣥࢳࣕࢿࣝࣀ࢖ࢬࡀຍࢃࡿሙ
ྜࡢ☜⋡ⓗ⚄⤒⣽⬊ࣔࢹࣝࡢ⥺ᙧస⏝⣲࡟ᑐࡋࠊࡑࡢᅛ᭷್ࠊᅛ᭷㛵ᩘࢆィ⟬ࡍࡿ࡜ࡁࠊࣇ࣮࢚ࣜ
⣭ᩘᒎ㛤ࡸ᫖ᖺࡲ࡛◊✲ࡋ࡚࠸ࡓࢹ࣮ࢱࡢ␯ᛶ
㸦ࢮࣟࡲࡓࡣࢮࣟ࡜ぢ࡞ࡏࡿ㒊ศࡀከ࠸ࡇ࡜㸧࡟ᇶ࡙ࡃ⥺ᙧస⏝⣲ࡢᒎ㛤ࡀ౑࠼࡞࠸࠿᳨ウࢆ⾜ࡗࡓࠋ㸱㸬࢖࢜ࣥࢳࣕࢿࣝࡢ☜⋡ⓗ㛤㛢ࡢࡓࡵࣀ࢖ࢬࡀ
⏕ࡌࡿࡀࠊࡇࡢࣀ࢖ࢬࡀ࢖࢜ࣥࢳࣕࢿࣝᩘࡢቑຍ
࡜࡜ࡶ࡟࡝ࡢࡼ࠺࡞☜⋡㐣⛬࡛㏆ఝࡉࢀࡿ࠿ࡣࠊ
⚄⤒⣽⬊ࣔࢹࣝࡢᘧࡢ࡝ࡇ࡟࡝ࡢࡼ࠺࡞ࣀ࢖ࢬࢆ
ຍ࠼ࢀࡤⰋ࠸ࡢ࠿࡜࠸࠺ၥ㢟࡜㛵㐃ࡋ࡚࠾ࡾࠊ⚄⤒⣽⬊ࣔࢹࣝࡢᣲືࡢゎᯒ࡟ᚲせ࡞᝟ሗ࡛࠶ࡿࠋ
⚄⤒⣽⬊࡟Ꮡᅾࡍࡿ㟁఩౫Ꮡᛶ࢖࢜ࣥࢳࣕࢿࣝࡣ
ࡑࡢᣲືࡀ࣐ࣝࢥࣇ㐃㙐ࢆ⏝࠸⾲ࡉࢀࡿሙྜࡀ࠶
ࡿࠋࡇࡢሙྜ࡟ࡘ࠸࡚☜⋡ゎᯒࡢᡂᯝ࡛࠶ࡿὶయ
ຊᏛᴟ㝈ࡢࢸࢡࢽࢵࢡࢆ⏝࠸ࠊ࢖࢜ࣥࢳࣕࢿࣝࣀ
࢖ࢬࡀ࠶ࡿ☜⋡㐣⛬࡛㏆ఝࡉࢀࡿ࠿௚ࡢ◊✲⪅࡟ࡼࡾ♧ࡉࢀࡓࠋࡇࡢ㏆ఝࡢጇᙜᛶࢆㄪ࡭ࡿࡓࡵ࡟ࠊ
࣐ࣝࢥࣇ㐃㙐࡟ᇶ࡙ࡃ࢖࢜ࣥࢳࣕࢿࣝࣔࢹࣝࡢཝ
ᐦ࡞ᩘ್ࢩ࣑࣮ࣗࣞࢩࣙࣥࢆ⾜࠸᳨ドࡋࡓࠋⓎ⾲ㄽᩥ
1. Ishikawa, Y., Yamanobe, T., (2018) Asymptotic
expansion of a nonlinear oscillator with ajump-diffusion process, Japan Journal of Industrial
and Applied Mathematics, 35, 969-1004
Ꮫ఍Ⓨ⾲
1. Yamanobe, T., (2018) Reduction of the dynamics
of stochastic neuronal models by sparse discretecosine transform, 11th FENS Forum of Neuroscience,
July 10, Belin㻌
2. Yamanobe, T., Reduction of a Markov operator
representing the dynamics of stochastic neuronalmodel by sparse discrete cosine transform, ➨ ᅇ᪥ᮏ⏕≀≀⌮Ꮫ఍ᖺ఍㸪 ᖺ  ᭶  ᪥㹼 ᭶  ᪥㸪ᒸᒣ኱Ꮫὠᓥ࢟ࣕࣥࣃࢫ㸪ᒸᒣ

኱Ꮫ㝔ᆅ
ᆅ⌫⎔
⎔ቃ⛉Ꮫ◊
◊✲㝔
㝔
⤫ྜ⎔
⎔ቃ⛉
⛉Ꮫ㒊㛛 ⮬↛⎔ቃಖ඲ศ㔝㟢ᓮ ྐᮁ฼⏝ : Lea Végh࣭ᙇಙ⇩࣭࿘⅋⅋ᮏ ᖺ ᗘ ࡣ ࠊ (1) ࢫ ࢼ ࢦ ࢣ (Racomitrium japonicum
Dozy et Molk.)ᡂ⫱࡟ᑐࡍࡿࣜࢱ࣮ࡢᙳ㡪࡟㛵ࡍࡿᐇ㦂࡜ࠊ(2) ᭷⌔ᒣ࡟࠾ࡅࡿ᳃ᯘࢱ࢖ࣉẖࡢᇙᅵ✀Ꮚ㞟ᅋᵓ㐀ࡢᐃ㔞໬࡟㛵ࡍࡿᐇ㦂ࢆ⾜ࡗࡓࠋ(1) ࢫࢼࢦࢣᡂ⫱࡟ᑐࡍࡿࣜࢱ࣮ࡢᙳ㡪࡟㛵ࡍࡿ

−9−

ᐇ㦂
ⅆᒣᄇⅆᚋࡢ⏕ែ⣔࡛ࡣࠊᆅ⾲㠃࡟ࢥࢣࡀඃ༨
ࡍࡿࡇ࡜ࡀከ࠸ࠋ୍᪉ࠊඃ༨ࡍࡿ⥔⟶᮰᳜≀࠿ࡽ౪⤥ࡉࢀࡿࣜࢱ࣮ࡣࢥࢣࡢศᕸࣃࢱ࣮ࣥࢆ኱ࡁࡃつᐃࡋ࡚࠸ࡿࠋࡑࡇ࡛ࠊࣜࢱ࣮ࡢ㉁(✀)࡜㔞ࡀࢥࢣࡢᡂ⫱࡟୚࠼ࡿᙳ㡪ࢆㄪ࡭ࡿࡓࡵ࡟ࠊΏᓥ㥖ࢣᓅ࠿ࡽࠊ⹊ⱏ㢮ࡢ୰࡛ඃ༨ࡍࡿࢫࢼࢦࢣࠊⅆᒣᄇฟ≀ ( ㍍ ▼ ࣭ ⅆ ᒣ ⅊ ) ࠊ ࢝ ࣛ ࣐ ࢶ (Larix kaempferi(Lambert) Carrière)࡜࣑ࢿࣖࢼࢠ(Salix reinii Franch.
et Savat. ex Seemen)ࡢࣜࢱ࣮ࢆ᥇ྲྀࡋࠊ ᐊ᮲௳ୗ࡟࡚ࣜࢱ࣮✀ẖ࡟ࢫࢼࢦࢣࡢᡂ⫱ᐇ㦂ࢆ⾜ࡗࡓࠋࡲࡎࠊ࣏ࢵࢺ࡟ⅆᒣᄇฟ≀ࢆワࡵࠊࡑࡢୖ࡟ࢫࢼ
ࢦࢣࢆ⛣᳜ࡋࡓࠋࡑࡢୖ࡟ࠊ࣐࢝ࣛࢶࣜࢱ࣮ࡲࡓ
ࡣ࣑ࢿࣖࢼࢠࣜࢱ࣮ࢆ⨨ࡃ࠿↓ฎ⌮(ࣜࢱ࣮࡞ࡋ)ࡢ 3 ฎ⌮ࢆ⾜ࡗࡓࠋྛฎ⌮࡛ 15 ཯᚟ࢆ⏝ពࡋྜィ45 ࣏ࢵࢺࢆ౑⏝ࡋࡓࠋ࡞࠾ࠊࢫࢼࢦࢣࡣᖖ⥳ᛶ࡛஝⇱⪏ᛶࡀ㧗࠸ࡇ࡜ࡀ▱ࡽࢀ࡚࠸ࡿࠋ ᐊᐇ㦂ࡣ
6 ᭶࠿ࡽ 10 ᭶ࡲ࡛⾜࠸ࠊẖ᭶ࠊNDVI(ṇつ໬᳜⏕ᣦᩘ)ࢆ ᐃࡋࡓࠋNDVI ࡣගྜᡂάᛶ࡜┦㛵ࡀ࠶ࡿࡇ࡜ࡀ▱ࡽࢀ࡚࠸ࡿࡓࡵࠊNDVIᐃ࡟ࡼࡿ㠀◚ቯⓗ࡞ᡂ⫱ ᐃྍ⬟ᛶࢆ᳨ウࡍࡿࡇ࡜ࡶ┠ⓗࡢ୍ࡘ࡛࠶ࡿࠋᐇ㦂⤊஢᫬࡟ࢥࢣࢆᅇ཰ࡋࠊࢩ࣮ࣗ
ࢺఙ㛗ࠊⴥ⥳⣲(a, b)ࢆᐃ㔞ࡋࡓࠋࡇࢀࡽࡢ⤖ᯝࢆࠊ㔝እᐇ㦂ࡢ⤖ᯝ࡜ẚ㍑ࡋࡓࠋࡑࡢ⤖ᯝࠊ ᐊ࡛ࡣ
NDVI ࡣ 8 ᭶࡟ࣆ࣮ࢡࢆ♧ࡋࠊࡑࡢᚋࠊῶᑡࡋࡓࠋ
ࢥࢣࡢ NDVIࠊⴥ⥳⣲⃰ᗘࠊఙ㛗㔞ࡣࠊࣜࢱ࣮ฎ
⌮㛫࡛ࡣᕪࡀ࡞࠿ࡗࡓࠋ୍᪉ࠊ㔝እ࡛ࡣ NDVIࠊⴥ⥳⣲⃰ᗘࠊఙ㛗㔞ࡣฎ⌮㛫࡛ᕪࡀ࡞࠿ࡗࡓࠋ୍
᪉ࠊ㔝እ࡛ࡣࠊఙ㛗㔞ࡣ࣐࢝ࣛࢶࣜࢱ࣮ࡢཌࡉࡀ
୰⛬ᗘࡢ࡜ࡇࢁ࡛㧗࠿ࡗࡓࠋࡲࡓࠊ㔝እ࡛ࡣ࣑ࢿ
ࣖࢼࢠࣜࢱ࣮ࡢཌࡉ࡜ࢥࢣࡢఙ㛗㔞࡟ࡣ㈇ࡢ┦㛵ࡀ࠶ࡗࡓࠋ⌧ᅾࠊᏳᐃ⎔ቃୗ࡛࠶ࡿᐊෆᐇ㦂࡜⎔ቃኚືࡢ኱ࡁ࡞㔝እ࡛ࡢ⤖ᯝࡢ┦㐪࡟㛵ࡋࡓせᅉࢆ᳨ウࡋ࡚࠸ࡿࠋ
(2) ᭷⌔ᒣ࡟࠾ࡅࡿ᳃ᯘࢱ࢖ࣉẖࡢᇙᅵ✀Ꮚ㞟ᅋᵓ㐀ࡢᐃ㔞໬ᐇ㦂ⅆᒣ㑄⛣࡟࠾࠸࡚ᇙᅵ✀Ꮚ㞟ᅋࡢⓎ㐩ᵝᘧࡣࠊ㑄⛣ࡢ᪉ྥࢆỴࡵࡿୖ࡛㔜せ࡛࠶ࡿࠋࡑࡇ࡛ࠊ᭷
⌔ᒣ࡟࠾ࡅࡿ 4 ࡘࡢ᳃ᯘࢱ࢖ࣉ࠿ࡽ 2017 ᖺ୰࡟῝
ࡉ 5 cm ࡲ࡛ࡢᅵተࢆᐜ㔞 100 cm3 ࡢ᥇ᅵ⟶ࢆ⏝࠸࡚᥇ྲྀࡋ෭ ᐊಖ⟶ᚋ࡟ࠊ ᐊ࡟࡚ 2018 ᖺ 1 ᭶
18 ᪥࠿ࡽ 9 ᭶ 27 ᪥ࡲ࡛Ⓨⱆᐇ㦂ࢆ⾜ࡗࡓࠋⓎⱆࡣࠊẖ㐌ࠊ᭶࣭Ỉ࣭㔠᭙᪥࡟ࢳ࢙ࢵࢡࡋࡓࠋ᪂ࡓ
࡟Ⓨⱆࡋࡓಶయ࡟ࡣࠊ␒ྕࢆグࡋࡓ᪝ࢆ㏆ࡃ࡟❧࡚࡚ಶయ㆑ูࢆ⾜ࡗࡓࠋྠᐃࡀᅔ㞴࡞ᐇ⏕࡟ࡘ࠸࡚ࡣࠊ኱ᆺ࣏ࢵࢺ࡟⛣ࡋࠊྠᐃྍ⬟࡞ࢧ࢖ࢬ࡟࡞
ࡿࡲ࡛ᡂ㛗ࡉࡏࡓࠋ඲࡚ࡢⓎⱆࡣࠊ3 ᭶୰࡟኱༙
ࡀ⤊஢ࡋࡓࠋࡑࡢ⤖ᯝࠊ඲࡚ࡢᐇ⏕ࡢྠᐃࡀ⾜࠼ࠊ඲య࡛ 73 ಶࡢᐇ⏕ࡀ☜ㄆࡉࢀࡓࠋ඲యࢆ㏻ࡋ࡚ࡢඃ༨✀ࡣࠊࣄ࣓ࢫࢤ(Carex oxyandra (Franch. et
Savat.) Kudo)ࠊ࢔࣑࢜ࢬ(Pilea mongolica Weddell)ࠊ

ࢥࣁࢥ࣋(Stellaria media (L.) Villars)ࠊ࢜࢜ࣚࣔࢠ
(Artemisia montana (Nakai) Pamp.)࡛࠶ࡗࡓࠋࡇࢀࡽ
ࡢ࠺ࡕࠊࣄ࣓ࢫࢤࡣ஝⇱ࡋࡓ᳃ᯘࠊ࢔࣑࢜ࢬࡣ‵ᛶᯘࠊࢥࣁࢥ࣭࣋࢜࢜ࣚࣔࢠࡣࠊ(༙)ேᕤᯘෆ࡛ࡼ
ࡾከࡃㄆࡵࡽࢀࠊྛ᳃ᯘᆺ࡟࠾࠸࡚≉ᚩࡢ࠶ࡿᇙ
ᅵ✀Ꮚ㞟ᅋࡢⓎ㐩ࣃࢱ࣮ࣥࡀ♧ࡉࢀࡓࠋㄽᩥ
Holle MJM & Tsuyuzaki S. (2018) The effects of shrub
patch sizes on the colonization of pioneer plants on
the volcano Mount Koma, northern Japan. Acta
Oecologica 93: 48-55
Shishir, S. & Tsuyuzaki, S. (2018) Hierarchicalclassification of land use types using multiple
vegetation indices in relation to urbanization.
Environmental Monitoring and Assessment 190:
article 342
Ꮫ఍Ⓨ⾲Végh L, Tsuyuzaki S. (2019) The fate of pioneers differences in growth of woody plants on Mount
Usu. ➨ 66 ᅇ᪥ᮏ⏕ែᏛ఍(⚄ᡞ)
㟢ᓮྐᮁ. (2019) ᭷⌔ᒣ 1977-78 ᖺᄇⅆᚋ 40 ᖺ㛫ࡢ᳜⏕ኚ໬. ➨ 66 ᅇ᪥ᮏ⏕ែᏛ఍(⚄ᡞ)ᶓᒣ ග࣭ᒸ⏣ ᘯ࣭㟢ᓮྐᮁ࣭୕ᯇ୕ᮁ࣭㜿㒊⚽ᙪ࣭Ṋᕝṇே࣭ᒸ㔝 ῟࣭▼஭ ඘࣭㕥ᮌΎ㝯࣭㧗ᶫၨ௓࣭኱ᶫு௓. (2018) ᅜ❧බᅬෆ࡟࠾ࡅࡿ
ⅆᒣ⅏ᐖ㑇ᵓࡢಖ඲ࢩࢫࢸ࣒ࡢᵓ⠏ -Ὕ√†᭷⌔ᒣࣘࢿࢫࢥୡ⏺ࢪ࢜ࣃ࣮ࢡࡢヨࡳ-. ᆅ⌫ᝨᫍ⛉Ꮫ㐃኱఍ (༓ⴥ)⤫ྜ⎔ቃ⛉Ꮫ㒊㛛 ⎔ቃ㐺ᛂ⛉Ꮫศ㔝Ἀ㔝 㱟ᩥ㸦ᾏỈ฼⏝㸧ㄽᩥⓎ⾲Kumagai, M., Nishikawa, K., Matsuura, H., Umezawa,T., Matsuda, F. and Okino, T. (2018) Antioxidants fromthe brown alga Dictyopteris undulata, Molecules, 23:
1214.

⎔ቃ⏕≀⛉
⛉Ꮫ㒊㛛 㝣⏕⏕ែᏛศศ㔝ᕤ⸨ ᓅ
฼⏝≧ἣ࣭◊✲ᡂᯝ㢧ⰼ᳜≀ࡢ⣙  ๭ࡣ㏦⢊ࢆື≀࡟㢗ࡗ࡚࠾ࡾࠊ㏦⢊ື≀ࢆຠ⋡ⓗ࡟ㄏᘬࡍࡿࡓࡵ࡟ⰼࡢⰍ࣭ᙧ࣭㤶ࡾ࡞࡝ࡢಙྕࢆ౑ࡗ࡚࠸ࡿࠋࡇࢀࡽࡢಙྕࡣ㉁
ⓗ࣭㔞ⓗ࡟㛤ⰼᮇ㛫୰୍ᐃ࡛࠶ࡿ࡜ࡣ㝈ࡽࡎࠊ㛤ⰼᮇ㛫࣭᫨ኪ㛫࡛ኚ໬ࡍࡿࡇ࡜ࡀሗ࿌ࡉࢀ࡚࠸ࡿࠋ
ࡑࡇ࡛ࠊᮏ◊✲࡛ࡣ㏆⦕✀࡛᫨ኪ㛫࡛ࡢ㏦⢊⪅ㄏ− 10 −

ᘬᙧ㉁ ⰼ⻤⢾ᗘ࠾ࡼࡧⰾ㤶㔞 ࡢኚ໬ࡀሗ࿌ࡉࢀ
࡚࠸ࡿ࣓࣐ࢶࣚ࢖ࢢࢧ 2HQRWKHUDELHQQLV ࢆᑐ
㇟࡟ࠊ㏦⢊⪅ᒙࡢኚ໬࡜⦾Ṫᡂຌᗘࡢ᫨ኪ㛫࡛ࡢ㐪࠸ࢆ᫂ࡽ࠿࡟ࡍࡿࡇ࡜ࢆ┠ⓗ࡜ࡋ࡚  ✀ࡢᐇ㦂ࢆ⾜ࡗࡓࠋࢤࣀ࣒ࢲ࢖ࢼ࣑ࢡࢫࢭࣥࢱ࣮ࡢᅡሙ࡟
࡚ࠊ ᖺ  ᭶࡟໭ᾏ㐨▼⊁ᕷࠕࡣࡲ࡞ࡍࡢୣබ
ᅬࠖ࠿ࡽࡢ᥇ྲྀಶయࢆ⛣᳜ࡋࡓࠊேᕤಶయ⩌ࢆ౑⏝ࡋࡓࠋձ ࣓࣐ࢶࣚ࢖ࢢࢧ࡟ゼⰼࡍࡿ᪻⹸┦ࡢ᫨ኪ㛫ẚ
㍑
 ᖺ  ᭶࠿ࡽྠᖺ  ᭶࡟࠾࠸࡚ࠊேᕤಶయ
⩌ࢆᑐ㇟࡟ኪ㛫࣭᫨㛫ࡑࢀࡒࢀࡢゼⰼ᪻⹸ほᐹࢆ⾜ࡗࡓࠋࡑࡢ⤖ᯝࠊኪ㛫ࡢほᐹ࡛ࡣ㫣⩟┠ ✀ࠊ᫨㛫ࡢほᐹ࡛ࡣ཮⩟┠  ✀࡜⭷⩟┠  ✀ࡀ☜ㄆࡉࢀࠊ᫨ኪ㛫࡛ࡢゼⰼ᪻⹸ࡢ㐪࠸ࡀほᐹࡉࢀࡓࠋࡲࡓࠊゼⰼ᪻⹸┦࡟ࡣᏘ⠇ᛶࡀᏑᅾࡋࡓࠋ
ࡇࡢࡇ࡜࠿ࡽࠊ࣓࣐ࢶࣚ࢖ࢢࢧ࡟࠾࠸࡚ࡶ᫨ኪ
㛫ࡢ㏦⢊⪅ㄏᘬຠ⋡࡟㐪࠸ࡀ࠶ࡿࡇ࡜ࡀ♧၀ࡉࢀࡓࠋղ ᧯సᐇ㦂࡟ࡼࡿ࣓࣐ࢶࣚ࢖ࢢࢧࡢ✀Ꮚ⏕⏘㔞
ࡢẚ㍑
࣓࣐ࢶࣚ࢖ࢢࢧ࡟࡜ࡗ࡚ࠊ᫨ኪ㛫࡛ࡢ㏦⢊⪅ㄏᘬࡢ㐪࠸ࡀ࡝ࡢࡃࡽ࠸⦾Ṫᡂຌ࡟ᙳ㡪ࡍࡿ࠿ࢆ᫂ࡽ࠿࡟ࡍࡿࡓࡵࠊࢥࣥࢺ࣮ࣟࣝࠊ㝖㞝᫨㛤ᨺࠊ㝖㞝ኪ㛤ᨺࠊ㝖㞝඲᪥ゎᨺࠊ⿄᥃ࡅࡢ ฎ⌮ࢆ⾜࠸ࠊ⤖ᯝ⋡ࠊ⤖ᐇ⋡ࠊ✀Ꮚ஝⇱㔜㔞ࢆẚ㍑ࡋࡓࠋࡇࡢ⤖ᯝࠊ⤖ᯝ⋡ཬࡧ✀Ꮚ஝⇱㔜㔞࡟ࡘ࠸࡚ࡣฎ⌮㛫ࡢᕪࡣぢࡽࢀ࡞࠿ࡗࡓࠋ⤖ᐇ⋡࡟ࡘ࠸࡚ࡣࠊ⿄᥃ࡅฎ⌮ࡢ⤖ᐇ⋡ࡀᏳᐃࡋ࡚㧗ࡃࠊ㝖㞝ฎ⌮ࢆࡍࡿ࡜⤖ᐇ⋡ࡀపࡃ࡞ࡿࡇ
࡜ࡀศ࠿ࡗࡓࠋࡉࡽ࡟ࠊࠕ᫨㛫ࡢࡳࠖࠕኪ㛫ࡢ
ࡳࠖࡢ㏦⢊࡟ࡼࡿ⤖ᐇ⋡࡟ࡣ㐪࠸ࡀぢࡽࢀ࡞࠿ࡗࡓࠋճ Ⓨⱆᐇ㦂࡟ࡼࡿ⦾Ṫᡂຌࡢẚ㍑
ୖグղࡢᐇ㦂࡟࠾࠸࡚ࠊ✀Ꮚᙧᡂᵝᘧ࡟ࡘ࠸࡚
᫂ࡽ࠿࡟࡞ࡗࡓࠋḟ࡟ࠊᙧᡂࡉࢀࡓ✀ᏊࡢⓎⱆ⬟ࡣ⮬Ṫ✀Ꮚ࡜௚Ṫ✀Ꮚ࡛␗࡞ࡿࡢ࠿ࢆ᳨ド
ࡋࡓࠋⓎⱆᐇ㦂ࡣࠊ┤ᚄ PP ࡢࣉࣛࢫࢳࢵࢡ
ࢩ࣮ࣕࣞ࡟సᡂࡋࡓᐮኳᇵᆅ࡟ࠊୖグղࡢᐇ㦂࡛ᚓࡽࢀࡓ✀Ꮚࢆ  ⢏ࡎࡘ᧛✀ࡋࠊΥࡢ୰᪥᮲௳࡛࠾ࡇ࡞ࡗࡓࠋⓎⱆほᐹࡣⓎⱆࡀᐃᖖ࡟㐩ࡍࡿ  ᪥㛫ࡲ࡛ẖ᪥⾜ࡗࡓࠋࡇࡢ⤖ᯝࠊ⮬Ṫ✀ᏊࡢⓎⱆ⋡ࡀᏳᐃࡋ࡚㧗࠸ࡇ࡜ࡀ᫂ࡽ࠿
࡟࡞ࡗࡓࠋմ ⰼࡢ㤶ࡾࡢ᫨ኪ㛫ࡢ㐪࠸ ⥅⥆୰ ࣐ࢶࣚ࢖ࢢࢧᒓࡢ᳜≀࡛ࡣⰼࡢ㤶ࡾ࡟᫨ኪ㛫࡛ࡢᕪࡀぢࡽࢀࡿ࡜࠸࠺ሗ࿌ࡀ࠶ࡿࠋࡑࡇ࡛ࠊ
ᐇ㦂ಶయ⩌࡟࠾࠸࡚࡝ࡢ⛬ᗘࡢⰼࡢ㤶ࡾࡢ㐪࠸ࡀ࠶ࡿࡢ࠿ࢆ᫂ࡽ࠿࡟ࡍࡿࡇ࡜ࢆ┠ⓗ࡜ࡋ࡚ⰼࡢ㤶ࡾࡢࢧࣥࣉࣜࣥࢢࢆ⾜ࡗࡓࠋኪ㛫ࡢ㤶ࡾ࡜ࡋ࡚᪥ἐᚋ  ᫬㛫ࠊ᫨㛫ࡢ㤶ࡾ࡜ࡋ࡚᪥ฟᚋ  ᫬㛫᥇ྲྀࡋࡓࠋࡇࡢࢧࣥࣉࣝࢆ *&06࡟࠿ࡅ࡚ゎᯒࡍࡿணᐃ࡛࠶ࡿࠋ
ࡇࢀࡲ࡛ࡢゎᯒ࠿ࡽࠊ࣓࣐ࢶࣚ࢖ࢢࢧࡣ✀Ꮚᙧᡂࡢከࡃࢆ⮬Ṫ࡛ࡲ࠿࡞࠺ࡇ࡜ࡀ࡛ࡁࡿࡇ࡜ࠊࡲࡓ
௚Ṫ࡟ࡘ࠸࡚ࡢ᫨ኪ㛫࡛ࡢ⦾Ṫᡂຌᗘࡣྠ⛬ᗘ࡛࠶ࡾࠊ᫨㛫ࡢ㏦⢊⪅ࡶ㔜せ࡛࠶ࡿࡇ࡜ࡀ♧၀ࡉࢀࡓࠋᏛ఍Ⓨ⾲኱ᔱᕼ⨾㸪ᕤ⸨ᓅ࣓࣐ࢶࣚ࢖ࢢࢧ࡟࠾ࡅࡿ⦾Ṫ≉ᛶ࡜⦾Ṫᡂຌᗘࡢ᫨ኪ㛫ẚ㍑᪥ᮏ⏕ែᏛ఍➨ ᅇ኱఍㸪 ᖺ  ᭶  ᪥㸪⚄ᡞᅜ㝿఍㆟ሙ㸦⚄ᡞᕷ㸧㸬኱Ꮫ㝔⌮Ꮫ◊✲㝔≀⌮Ꮫ㒊㛛 㔞Ꮚ≀⌮Ꮫศ㔝ᯇỌ ᝅ᫂
㸦⣧Ỉ฼⏝㸧Ꮫ఍Ⓨ⾲
Hiroyoshi Nobukane
Fractional Chern invariant in a Sr2RuO4 nanofilm,
Research Seminar@Japanese-Russian JSPS-RFBR projectFebruary 20 2019, Hokkaido University, Japan
H. Nobukane, K. Yanagihara, Y. Kunisada, Y. Ogasawara,
K. Isono, K. Nomura, T. Nomura, T. Akiyama, Y. Asanoand S. Tanda,
High-Tc superconductivity in a ruthenate nanofilm
12th International Conference on Materials andMechanisms of Superconductivity and High Temperature
Superconductors (M2S-2018)
August 19-24, 2018 Beijing, China
N. Matsunaga, T. Minamidate, T. Wada, S. Kawaguchi,
A. Kawamoto, K. Nomura
Universal Phase Diagram of the Ȣ,Ȣ̓and Ȣ̓̓SaltsInternational Conference on Science and Technology
of Synthetic Metals 2018 (ICSM 2018)
July 1-6, 2018, Busan, Korea
S. Kawaguchi, T. Wada, T. Minamidate, N. Matsunaga,
A. Kawamoto, K. Nomura
Electric and Magnetic Properties of λ’-(BEDT-STF)2GaBr4
International Conference on Science and Technology
of Synthetic Metals 2018 (ICSM 2018)
July 1-6, 2018, Busan, Korea
T. Minamidate, T. Wada, N. Matsunaga, A. Kawamoto,and K. Nomura
Magnetic Properties in OrganicS- dsystem OO’ andN-(STF)2FeX4(X = Cl, Br)
International Conference on Science and Technology
of Synthetic Metals 2018 (ICSM 2018)− 11 −

29

July 1-6, 2018, Busan, Korea

Si-NMR. Journal of the Physical Society of Japan 87:
114707.

ᘏවၨ⣧ࠊᰗཎ⯆ୡࠊᅧ㈆㞝἞ࠊᑠ➟ཎඃோࠊ♒
㔝⩧ࠊ㔝ᮧ୍ᡂࠊ⬟ᮧ㈗ᏹࠊ⛅ᒣ཭ᏹࠊὸ㔝Ὀᐶࠊ୹⏣⪽ࣝࢸࢽ࣒࢘㓟໬≀࡟࠾ࡅࡿ㉸ఏᑟ㸫⤯⦕య㌿⛣࡜⮫⏺ᣦᩘ᪥ᮏ≀⌮Ꮫ఍  ᖺ⛅Ꮨ኱఍ࠊྠᚿ♫኱Ꮫி⏣㎶࢟ࣕࣥࣃࢫࠊ
 ᖺ  ᭶  ᪥㹼 ᪥ᘏවၨ⣧ࠊ⏣⏿⿱୍ࠊ㯮⃝ᚭࠊ㓇㒊኱ᶞࠊ୹⏣⪽㕲ࢻ࣮ࣉ 1E6 ࡟࠾ࡅࡿ㏆⸨ຠᯝ࡜ࢫࣆࣥࢢࣛࢫ
ࡢඹᏑ᪥ᮏ≀⌮Ꮫ఍  ᖺ➨  ᅇᖺḟ኱఍ࠊ஑ᕞ኱
ఀ㒔࢟ࣕࣥࣃࢫࠊ
 ᖺ  ᭶  ᪥㹼 ᪥ᕝཱྀᝅྖ࿴⏣኱㜿༡⯓ᏕுᯇỌᝅ᫂Ἑᮏ
඘ྖ㔝ᮧ୍ᡂ
ȢཬࡧȢ̓ሷࡢ⤫୍┦ᅗ᪥ᮏ≀⌮Ꮫ఍  ᖺ➨  ᅇᖺḟ኱఍ࠊ஑ᕞ኱
ఀ㒔࢟ࣕࣥࣃࢫࠊ
 ᖺ  ᭶  ᪥㹼 ᪥≀⌮Ꮫ㒊㛛 㟁Ꮚ≀ᛶ≀⌮Ꮫศ㔝Ἑᮏ ඘ྖ
㸦⣧Ỉ฼⏝㸧
ㄽᩥ
Fukuoka, S., Sawada, M., Minamidate, T., Matsunaga,N., Nomura, K., Ihara, Y., Kawamoto, A., Doi, Y.,
Wakeshima, M., and Hinatsu, Y. (2018) Multistep
development of the hyperfine fields in O-(BEDT-STF)2FeCl4
studied by Mössbauer spectroscopy. Journal of the
Physical Society of Japan 87: 093705.
Ohnuma, A., Taniguchi, H., Takahashi, Y., and Kawamoto,A. (2018) Density wave state in new D-type organic
conductor D-(BEDT-TTF)2MHg(XCN)4 (M = NH4, X= Se): Material for universal phase diagram of X = S
and X = Se systems. The Journal of Physical
Chemistry C 122(42): 24321-24328.
Saito, Y., Minamidate, T., Kawamoto, A., Matsunaga,N., and Nomura, K. (2018) Site-specific 13C NMR
study on the locally distorted triangular lattice of theorganic conductor N-(BEDT-TTF)2Cu2(CN)3. PhysicalReview B 98: 205141.
Tabata, C., Ihara, Y., Shimmura, S., Miura, N., Hidaka, H.,
Yanagisawa, T., and Amitsuka, H. (2018) Uncompensatedantiferromagnetic ordering of UAu2Si2 studied by

Fukuoka, S., Fukuchi, S., Akutsu, H., Kawamoto, A.,
and Nakazawa, Y. (2019) Magnetic and electronic
properties of S-d interacting molecular magnetic
superconductor N-(BETS)2FeX4 (X = Cl, Br) studied
by angle-resolved heat capacity measurements.
Crystals 9(2): 66.
Ꮫ఍Ⓨ⾲
ᅜ㝿Ꮫ఍
Fukuoka, S.: Low temperature magnetic state of
O-(BEDT-STF)2FeCl4.
International Symposium of Structural Thermodynamics
for Young Thermodynamicists: ISST-YT, June 8 2018,኱㜰኱Ꮫ㸦㇏୰ᕷ㸧.
Arashima, K., Ihara, Y., and Yoshida, H.: 7Li NMR
study of kagome antiferromagnet, Li2Cr3SbO8㸬
The 16th International Conference on Megagauss
Magnetic Field Generation and Related Topics,
September 27-28 2018, ᮾி኱Ꮫ᯽ࡢⴥ㥐๓࢟ࣕࣥࣃࢫ㸦᯽ᕷ㸧㸬
Ihara, Y., Arashima, K., Yoshida, H., Matsui, K., Kindo, K.,
and Kohama, Y.: Development of NMR spectrometerfor the flat-top pulse magnet.
The 16th International Conference on Megagauss Magnetic
Field Generation and Related Topics, September 27-28
2018, ᮾி኱Ꮫ᯽ࡢⴥ㥐๓࢟ࣕࣥࣃࢫ㸦᯽ᕷ㸧㸬ᅜෆᏛ఍஭ཎ៞ᙪ, ⲨᓥὯᶞ, ᯇ஭୍ᶞ, 㔠㐨ᾈ୍, ᑠ℈ⰾඔ㸸࣑ࢽࣇࣛࢵࢺࢺࢵࣉࣃࣝࢫ☢ሙࢆ⏝࠸ࡓ NMRᐃᢏ⾡㛤Ⓨ㸬
᪥ᮏ≀⌮Ꮫ఍  ᖺ⛅Ꮨ኱఍㸪 ᖺ  ᭶  ᪥㸪ྠᚿ♫኱Ꮫ㸦ி⏣㎶ᕷ㸧㸬ⲨᓥὯᶞ஭ཎ៞ᙪྜྷ⏣⣫⾜㸸/L105 ἲ࡟ࡼࡿప☢ሙ࡟࠾ࡅࡿ /L&U6E2 ࡢ☢Ẽ≧ែࡢ◊✲㸬᪥ᮏ≀⌮Ꮫ఍  ᖺ⛅Ꮨ኱఍㸪 ᖺ  ᭶  ᪥㸪ྠᚿ♫኱Ꮫ㸦ி⏣㎶ᕷ㸧㸬⃝⏣㈼ᚿ, ᩧ⸨ὒᖹ, ⚟ᒸ⬶ᖹ, Ἑᮏ඘ྖ㸸 13C
NMR ࡟ࡼࡿO-(BEDT-TTF)2GaCl4 ࡢᅽຊୗࡢ㟁Ꮚ≀ᛶࡢ◊✲㸬
᪥ᮏ≀⌮Ꮫ఍  ᖺ⛅Ꮨ኱఍㸪 ᖺ  ᭶  ᪥㸪ྠᚿ♫኱Ꮫ㸦ி⏣㎶ᕷ㸧㸬⚟ᒸ⬶ᖹ, ༡⯓Ꮥு, ᯇỌᝅ᫂, 㔝ᮧ୍ᡂ, ஭ཎ៞ᙪ, Ἑᮏ඘ྖ㸸 13C NMRᐃ࡟ࡼࡿS-d ⣔≀㉁
O-(BEDT-STF)2FeCl4 ࡢప ☢Ẽ≧ែࡢ◊✲㸬− 12 −

᪥ᮏ≀⌮Ꮫ఍  ᖺ⛅Ꮨ኱఍㸪 ᖺ  ᭶  ᪥㸪ྠᚿ♫኱Ꮫ㸦ி⏣㎶ᕷ㸧㸬஭ཎ៞ᙪ, ⲨᓥὯᶞ, ᯇ஭୍ᶞ, 㔠㐨ᾈ୍, ᑠ℈ⰾඔ㸸ࣃࣝࢫᙉ☢ሙ୰ NMR ᐇ㦂࡟ࡼࡿ᰾ࢫࣆࣥ᱁
Ꮚ⦆࿴᫬㛫 ᐃ.ᮾ໭኱Ꮫ㔠ᒓᮦᩱ◊✲ᡤ◊✲఍  ᖺ  ᭶ ᪥ᮾ໭኱Ꮫ ௝ྎᕷ ⲨᓥὯᶞ, ஭ཎ៞ᙪ, బࠎᮌᏕᙪ, ྜྷ⏣⣫⾜㸸࢝ࢦ࣓཯ᙉ☢ᛶయ Ca ࢝࣌ࣛࢧ࢖ࢺ࡟࠾ࡅࡿ☢Ẽບ㉳
ࡢ 35Cl NMR ศගἲ࡟ࡼࡿ◊✲.ᮾ໭኱Ꮫ㔠ᒓᮦᩱ◊✲ᡤ◊✲఍  ᖺ  ᭶ ᪥ᮾ໭኱Ꮫ ௝ྎᕷ ⲨᓥὯᶞ, ஭ཎ៞ᙪ, బࠎᮌᏕᙪ, ᖹ⏣೔ၨ, ྜྷ⏣⣫⾜㸸7Li NMR ἲ࡟ࡼࡿ㧗☢ሙ࡟࠾ࡅࡿ Li2Cr3SbO8
ࡢ☢Ẽ≧ែࡢ◊✲㸬᪥ᮏ≀⌮Ꮫ఍➨  ᅇᖺḟ኱఍㸪 ᖺ  ᭶  ᪥㸪஑ᕞ኱Ꮫ㸦⚟ᒸᕷ㸧㸬஭ཎ៞ᙪ, ⲨᓥὯᶞ, ᯇ஭୍ᶞ, 㔠㐨ᾈ୍, ᑠ℈ⰾඔ㸸ࢫ࣮ࣟࣉࢺࢵࣉࣃࣝࢫ☢ሙࢆ⏝࠸ࡓ☢ሙᤲᘬ
NMR ࢫ࣌ࢡࢺࣝ ᐃ࡟ࡼࡿ Li2Cr3SbO8 ࡢ☢Ẽᵓ㐀ࡢ◊✲㸬᪥ᮏ≀⌮Ꮫ఍➨  ᅇᖺḟ኱఍㸪 ᖺ  ᭶  ᪥㸪஑ᕞ኱Ꮫ㸦⚟ᒸᕷ㸧㸬⚟ᒸ⬶ᖹ㸪༡⯓Ꮥு㸪ᯇỌᝅ᫂㸪㔝ᮧ୍ᡂ㸪஭ཎ៞ᙪ㸪Ἑᮏ඘ྖ㸸O-(BEDT-STF)2FeCl4 ࡢ☢Ẽ⛛ᗎ࡟㛵ࡍࡿ⇕ⓗ◊✲.
᪥ᮏ≀⌮Ꮫ఍➨  ᅇᖺḟ኱఍㸪 ᖺ  ᭶  ᪥㸪஑ᕞ኱Ꮫ㸦⚟ᒸᕷ㸧㸬㎷᫭ᖹ, ᑠᯘᣅ▮, Ἑᮏ඘ྖ㸸᭷ᶵ㉸ఏᑟయO-(BETS)2GaCl4 ࡢ 69/71Ga-NMR ἲࢆ⏝࠸ࡓ㟁Ⲵ/ࢫ
ࣆࣥ␗ᖖࡢ㑅ᢥⓗ᳨ฟ㸬᪥ᮏ≀⌮Ꮫ఍➨  ᅇᖺḟ኱఍㸪 ᖺ  ᭶  ᪥㸪஑ᕞ኱Ꮫ㸦⚟ᒸᕷ㸧㸬㒊㛛 ᙧែᶵ⬟Ꮫศศ㔝⏕≀⛉Ꮫ㒊ᒣཱྀ ῟஧1) ⢾ࡣ⏕≀ࡢᇶᖿ௦ㅰࢆᨭ࠼ࡿᰤ㣴⣲࡛࠶ࡾ㸪ࡲࡓ⏕≀ಶయࡢᡂ㛗ࡸⓎ㐩ࢆไᚚࡍࡿ㔜せ࡞ࢩࢢࢼࣝศᏊ࡜ࡋ࡚ࡶᶵ⬟ࡍࡿࠋࡋ࠿ࡋ㸪ࡑࡢ」㞧ࡉ࠿ࡽ㸪⢾ࢩࢢࢼࣝఏ㐩⣔ࡢヲ⣽࡞ศᏊᶵᵓࡣ౫↛࡜ࡋ࡚୙᫂࡞Ⅼࡀከ࠸ࠋᮏ◊✲࡛ࡣ㸪ࣔࢹ᳜ࣝ≀ࢩࣟ࢖ࢾࢼࢬࢼࢆ⏝࠸࡚㸪᪂つࡢ⢾ᛂ⟅ᛶ㌿෗ᅉᏊࢆ᥈⣴ࡋ㸪ᶵ⬟ᮍ▱ࡢ㌿෗ᅉᏊ bZIP3 ࢆྠᐃࡋࡓࠋ

㑇ఏᏛⓗゎᯒ࠿ࡽ㸪bZIP3 㑇ఏᏊⓎ⌧ࡀ⢾࡟ࡼࡗ࡚ᢚไࡉࢀࡿࡇ࡜㸪⢾ࢩࢢࢼࣝఏ㐩⤒㊰ࡢ୰᰾ࢆᢸ࠺ SnRK1 ࢟ࢼ࣮ࢮ㸦့ங㢮 AMPK ࣍ࣔࣟࢢ㸧ࡢไᚚୗ࡟࠶ࡿࡇ࡜ࡀศ࠿ࡗࡓࠋࡉࡽ࡟㸪⯆࿡῝࠸⾲⌧ᆺ࡜ࡋ࡚㸪bZIP3 ኚ␗ᰴ࡛ࡣᏊⴥࡢ཯ࡾ㏉ࡾ࡜࠸ࡗࡓ㸪ⴥࡢᙧែ࡟␗ᖖࡀぢࡽࢀࡓࠋࡇࢀࡣ㸪⢾ᛂ⟅ᛶ㌿෗ᅉᏊ bZIP3 ࡀ᳜≀ࡢ⢾ࢩࢢࢼࣝ࡜ᙧ
ែᙧᡂࢆࡘ࡞ࡄ᪂ࡓ࡞ᅉᏊ࡛࠶ࡿྍ⬟ᛶࢆ♧၀ࡋ࡚࠸ࡿࠋ [Sanagi et al, Plant Biotechnology 2018]2) ᳜≀ࡢ⎔ቃ㐺ᛂせᅉ࡜ࡋ࡚ࡢ C/N ࣂࣛࣥࢫࡢ
ゎᯒࢆ┠ⓗ࡟㸪␗࡞ࡿ❅⣲※ୗ࡛⏕⫱ࡉࡏࡓࢩࣟ
࢖ࢾࢼࢬࢼࢆ⏝࠸࡚㸪ග໬Ꮫ⣔ϩຠ⋡㸪ྛ✀ྍ⁐ᛶ⢾ྵ㔞㸪◪㓟㑏ඖ㓝⣲άᛶ࡟ࡘ࠸᳨࡚ウࢆຍ࠼ࡓࠋࡑࡢ⤖ᯝ㸪⢾㐣๫࡜ప❅⣲≧ែࡢ㛫࡟ࡣࢡࣟࢫࢺ࣮ࢡࡀᏑᅾࡋ㸪◪㓟㑏ඖ㓝⣲άᛶࢆไᚚࡍࡿ
ࡇ࡜࡛ୖグࢫࢺࣞࢫࡢᅇ㑊ࢆᅗࡿࡇ࡜ࡀド᫂ࡉࢀࡓࠋ[Huarancca Reyes et al, Plant and Cell Physiology
2018]ࠋ3) ⢾࡜❅⣲ࡢࣂࣛࣥࢫࠕC/Nࠖࡣ᳜≀ࡢ⏕⫱ࢆ኱ࡁࡃᕥྑࡍࡿᰤ㣴ࢩࢢࢼࣝ࡜࡞ࡿࠋᙜ◊✲ᐊ࡛ࡣ㸪᳜≀ࡢ C/N ᛂ⟅࡟ࡘ࠸࡚⊂⮬ࡢᐇ㦂⣔ࢆ☜❧ࡋ㸪ゎᯒࢆ⾜ࡗ࡚ࡁࡓࠋᮏⴭ᭩࡛ࡣ㸪᳜≀ࡢ⪁໬ゎᯒ࡟㛵ࡋ࡚᭷⏝࡞኱Ẽ୰ CO2 ⃰ᗘ࡜Ỉ⪔᱂ᇵ࡛ࡢ᰿
ᅪ❅⣲⎔ቃࡢไᚚ࡟ࡼࡿ C/N ฎ⌮ᐇ㦂ᡭἲࡢヲ⣽࡟ࡘ࠸࡚ᇳ➹ࡋࡓࠋ[Aoyama et al, in͆Plant Senescence
Methods and Protocols " 2018]ࠋ
㸺ཎⴭㄽᩥ㸼
Sanagi, M., Lu, Y., Aoyama, S., Morita, Y., Mitsuda, N.,
Ikeda, M., Ohme-Takagi, M., Sato, T. and Yamaguchi, J.
(2018) Sugar-responsive transcription factor bZIP3 affects
leaf shape in Arabidopsis plant. Plant Biotechnology. 35:
167-170 [10.5511/plantbiotechnology.18.0410a]
Huarancca Reyes, T., Scartazza, A., Pompeiano, A., Ciurli,A., Lu, Y., Guglielminetti, L. and Yamaguchi, J. (2018)
Nitrate reductase modulation in response to changes in
C/N balance and nitrogen source in Arabidopsis. Plant
and Cell Physiology, 59:1248-1254 [10.1093/pcp/pcy065]
Aoyama, S., Yamaguchi, J. and Sato, T. (2018) Methods
for Elucidation of Plant Senescence in Response to
C/N-Nutrient Balance in “Plant Senescence Methods
and Protocols " (edited by Y Guo), Methods in Molecular
Biology, vol. 1744, Springer Protocol, Humana Press,
New York, NY, pp. 151-159㻌 (DOI 10.1007/978-1-4939-7672-0;
ISBN 978-1-4939-7670-6)⏕≀⛉Ꮫ㒊㒊㛛 ⾜ື⚄⤒⏕≀Ꮫศศ㔝ỈἼ ㄔ ࣇࢱ࣍ࢩࢥ࢜ࣟࢠ࡟࠾ࡅࡿ )R[3 ㌿෗ᢚไᅉᏊ
ࡢ㛗ᮇグ᠈ᙧᡂ࡬ࡢ㛵୚ࡢゎ᫂)RUNKHDG%R[3 )R[3 ࡣ⬨᳝ື≀࡜⠇㊊ື≀࡞࡝− 13 −

ࡢ⣔⤫㛫࡛ಖᏑࡉࢀࡓ㌿෗ᢚไᅉᏊ࡛࠶ࡿࠋ့ங
㢮ࡸ㫽㢮࡛ࡣ )R[3 ࡣ⚄⤒⣔ࡢⓎ⏕ࠊච␿⣔ࡢㄪᩚ࡞࡝࡟㛵ࢃࡿ࡜࡜ࡶ࡟ࠊࣄࢺ࡛ࡣゝㄒ࡟ࠊ㫽࡛ࡣḷᏛ⩦࡟㛵ࢃࡿࡇ࡜ࡀ▱ࡽࢀ࡚࠸ࡿࠋ᭱㏆࣑ࢶࣂࢳࡸࢩࣙ࢘ࢪࣙ࢘ࣂ࢚࡟࠾࠸࡚ )R[3 ࡣ୺࡟⬻࡛
Ⓨ⌧ࡍࡿࡇ࡜ࡀሗ࿌ࡉࢀࠊ᪻⹸ࡢ⬻࡟࠾ࡅࡿࡑࡢ
ᶵ⬟ࡀὀ┠ࡉࢀࡿࠋ ᮏᖺᗘࣇࢱ࣍ࢩࢥ࢜ࣟࢠࢆ⏝࠸࡚ 51$L ࡟ࡼࡾ)R[3 Ⓨ⌧ᢚไᐇ㦂ࢆ⾜࡞ࡗࡓ⤖ᯝࠊ)R[351$L ಶయ࡛ࡣໝ࠸ࡢ㛗ᮇグ᠈ࡢᙧᡂࡀ㜼ᐖࡉࢀࡿࡇ࡜ࡀ
᫂ࡽ࠿࡟࡞ࡗࡓࠋໝ࠸Ꮫ⩦ࡢ⋓ᚓࡸ୰ᮇグ᠈ࡢᙧᡂࠊࡲࡓໝ࠸Ꮫ⩦࡟ᚲせ࡞ឤぬᶵ⬟ࡸ㐠ືᶵ⬟࡞࡝࡟ࡣ㞀ᐖࡣぢࡽࢀ࡞࠿ࡗࡓࠋᮏ◊✲ࡣ )R[3 㑇ఏᏊࡀ㛗ᮇグ᠈ᙧᡂ࡟ᚲ㡲࡞㑇ఏᏊ࡛࠶ࡿࡇ࡜ࢆึ
ࡵ࡚᫂ࡽ࠿࡟ࡋࡓࡶࡢ࡛࠶ࡿࠋ௒ᚋࡑࡢ࣓࢝ࢽࢬ
࣒ࡢゎ᫂ࢆ㐍ࡵ࡚࠸ࡁࡓ࠸ࠋ ࣇࢱ࣍ࢩࢥ࢜ࣟࢠ࡟࠾ࡅࡿ⩦័グ᠈ࡢࢥࣥࢸ
࢟ࢫࢺ౫Ꮡᛶ࡜ᾘཤカ⦎⪏ᛶࢥ࢜ࣟࢠࡢໝ࠸㸦᮲௳่⃭㸧࡜Ỉ㸦↓᮲௳่⃭ࠊࡲࡓࡣሗ㓘㸧࡜ࡢྂ඾ⓗ᮲௳௜ࡅ࡟࠾࠸࡚ࠊ㏻ᖖ
ࡢカ⦎᮲௳࡛ࡣࠊ᮲௳௜ࡅᚋ࡟ࢥ࢜ࣟࢠ࡟Ỉ㸦ሗ㓘㸧ࢆ༑ศ࡟㣧ࡲࡏ࡚ሗ㓘ࡢ౯್ࢆᘬࡁୗࡆࡿ࡜ࠊ
ໝ࠸࡬ࡢ᮲௳཯ᛂࡣ㉳ࡇࡽ࡞ࡃ࡞ࡿࠋ࡜ࡇࢁࡀ㐣๫࡟カ⦎ࢆ⧞ࡾ㏉ࡍ࡜ࠊỈࢆ㣧ࡲࡏ࡚ሗ㓘ࡢ౯್ࢆᘬࡁୗࡆ࡚ࡶໝ࠸࡬ࡢ᮲௳཯ᛂࡀ㉳ࡇࡿࡼ࠺࡟
࡞ࡿࡇ࡜ࢆ᭱㏆⚾ࡓࡕࡣ᫂ࡽ࠿࡟ࡋࡓࠋ㢮ఝࡢ⌧㇟ࡣࡇࢀࡲ့࡛ங㢮ࡢ࢜࣌ࣛࣥࢺ᮲௳௜ࡅ࡛ࠕ⩦័ᙧᡂࠖ࡜࿧ࡤࢀ࡚࠾ࡾࠊ⚾ࡓࡕࡢⓎぢࡣࠕ⩦័ᙧᡂࠖࡀྂ඾ⓗ᮲௳௜ࡅ࡛ࡶ㉳ࡇࡿࡇ࡜ࢆึࡵ࡚
᫂ࡽ࠿࡟ࡋࡓࡶࡢ࡛࠶ࡗࡓࠋ့ங㢮ࡢ࢜࣌ࣛࣥࢺ
᮲௳௜ࡅ࡛ᙧᡂࡉࢀࡿ⩦័グ᠈ࡣࠊึᮇグ᠈ࡼࡾࡶࢥࣥࢸ࢟ࢫࢺ౫Ꮡᛶࡀ㧗࠸ࡇ࡜ࠊࡲࡓᾘཤカ⦎ࡢຠᯝࡀప࠸ࡇ࡜ࡀ▱ࡽࢀ࡚࠸ࡿࠋ ࡑࡇ࡛ᮏᖺᗘࡣࢥ࢜ࣟࢠࡢྂ඾ⓗ᮲௳௜ࡅ࡟ࡼ
ࡾᙧᡂࡉࢀࡿ⩦័グ᠈ࡀྠᵝ࡞ᛶ㉁ࢆᣢࡘ࠿࡟ࡘ
࠸࡚ㄪ࡭ࡓ࡜ࡇࢁࠊ㐣๫カ⦎ᚋࡢ⩦័グ᠈ࡣࠊึᮇグ᠈ࡼࡾࡶࢥࣥࢸ࢟ࢫࢺ౫Ꮡᛶࡀప࠸ࡇ࡜ࠊࡲࡓᾘཤカ⦎ࡢຠᯝࡣึᮇグ᠈࡜ྠ⛬ᗘ࡛࠶ࡗࡓࠋ
ࡇࢀ࡟ࡼࡾࢥ࢜ࣟࢠࡢྂ඾ⓗ᮲௳௜ࡅ࡟ࡼࡾᙧᡂ
ࡉࢀࡿ⩦័グ᠈ࡢᛶ㉁ࡣࠊ့ங㢮ࡢ࢜࣌ࣛࣥࢺ᮲
௳௜ࡅ࡛ᙧᡂࡉࢀࡿ⩦័グ᠈࡜ࡣ㠀ᖖ࡟␗࡞ࡿᛶ㉁ࢆᣢࡘࡇ࡜ࡀ᫂ࡽ࠿࡟࡞ࡗࡓࠋࡇࡢࡼ࠺࡞ᛶ㉁ࡢ㐪࠸ࡀఱ࡟㉳ᅉࡍࡿࡢ࠿ࠊ௒ᚋ᫂ࡽ࠿࡟ࡋ࡚࠸
ࡁࡓ࠸ࠋ

ཎⴭㄽᩥ
1) Hamanaka Y. and Mizunami M. (2018) Tyrosinehydroxylase-immunoreactive neurons in the mushroom
body of the field cricket, Gryllus bimaculatus. Cell &Tissue Research https://doi.org/10.1007/s00441-018-2969-9.
2) Nishino H, Iwasaki M, Paoli M, Kamimura I,

Yoritsune A, Mizunami M. (2018) Spatial receptive
fields for odor localization. Current Biology 28:1-9.
3) Koizumi M, Shigeno S, Mizunami M, Tanaka N. K.
(2018) Calcium imaging method to visualize thespatial patterns of neural responses in the pygmy
squid, Idiosepius paradoxus, central nervous system.
Journal of Neuroscience Methods 294: 67-71.
ⴭ᭩1) ỈἼㄔ (2018) ື≀ࡢ஦඾ࠊᮎᯇ௚⦅ࠕឤぬჾࠖࢆᢸᙜࠊᮅ಴᭩ᗑ㸦༳ๅ୰㸧2) ỈἼㄔ (2018) ື≀Ꮫࡢⓒ⛉஦඾ࠊࠕ᪻⹸ࡢᚤ
ᑠ⬻ࠖࢆᢸᙜࠊ୸ၿࠊ
⥲ㄝ1) Mizunami M, Terao K, Alvarez B (2018) Applicationof a prediction error theory to Pavlovian conditioningin an insect. Frontiers in Psychology 9, 1272.
2) Matsumoto Y, Matsumoto CS, Mizunami M (2018)
Signaling pathways for long-term memory formationin the cricket. Frontiers in Psychology 9, 1014.

⏕≀⛉Ꮫ㒊㒊㛛 ⾜ື⚄⤒⏕≀Ꮫศศ㔝ᑠᕝ ᏹேᖹᡂ  ᖺᗘ◊✲ᡂᯝᐇ㦂⏕≀ඹྠ฼⏝㒊㛛࡛㣫⫱ࡋࡓࣇࢱ࣍ࢩࢥ࢜ࣟ
ࢠ㸦*U\OOXVELPDFXODWXV㸧ࢆᮦᩱ࡜ࡋ࡚㸪⚄⤒⏕≀Ꮫⓗᐇ㦂ࢆ⾜ࡗࡓࠋ ࢥ࢜ࣟࢠࡣ㸪⭡㒊ᮎ➃ࡢᑿⴥ࡜࿧ࡤࢀࡿᶵᲔឤぬჾᐁ࡛✵Ẽὶ᣺ືࢆཷᐜࡋ㸪▷࠸Ẽὶ่⃭࡟ᑐࡋ࡚⣲᪩ࡃ㐲ࡊ࠿ࡿṌ⾜⾜ືࢆ♧ࡍࠋࡇࡢ⾜ືࡣẼὶࢆᤕ㣗⪅ࡢ᥋㏆࡜ᤊ࠼㸪ᤕ㣗࠿ࡽ㏨ࢀࡼ࠺࡜
ࡍࡿ㏨㑊⾜ື࡛࠶ࡿ࡜⪃࠼ࡽࢀ࡚࠸ࡿࠋᡃࠎࡣࡇ
ࢀࡲ࡛࡟㸪Ẽὶ่⃭᪉ྥ᝟ሗฎ⌮ࡢ⚄⤒ᶵᵓ࡜ẼὶㄏᑟᛶṌ⾜㐠ືࡢไᚚࢩࢫࢸ࣒࡟㛵ࡍࡿ⚄⤒⏕≀Ꮫⓗ◊✲ࢆ⾜ࡗ࡚ࡁࡓࠋࡲࡓ㸪ᖹᡂ 30 ᖺᗘࡣ᪂ࡓ࡟࢜ࢫࢥ࢜ࣟࢠࡢⓎࡍࡿㄏᘬḷ࡟ᑐࡍࡿ࣓ࢫࡢ
᥋㏆⾜ື(㡢※ᐃ఩)ࡢ㐠ືゎᯒࢆ⾜ࡗࡓࠋࡑࡢ◊✲ᡂᯝࡣ୺࡟ḟࡢ㸰Ⅼ࡛࠶ࡿࠋ㸯㸬Ẽὶᛂ⟅ᛶᢞᑕࢽ࣮ࣗࣟࣥ࡟࠾ࡅࡿ่⃭ᙉᗘ౫Ꮡⓗ࡞᪉ྥ㑅ᢥᛶಟ㣭ࡢゎᯒࢽ࣮ࣗࣟࣥࡢ่⃭ࣃ࣓࣮ࣛࢱ࡟ᑐࡍࡿ㑅ᢥᛶࢆ⾲ࡍࢳ࣮ࣗࢽࣥࢢ≉ᛶࡣࠊឤぬ᝟ሗࡢࢥ࣮ࢹ࢕ࣥ
ࢢ⢭ᗘ࡟኱ࡁ࡞ᙳ㡪ࢆཬࡰࡍࠋࡋ࠿ࡋࠊಶࠎࡢ⚄⤒⣽⬊࡟࠾ࡅࡿࢳ࣮ࣗࢽࣥࢢ≉ᛶࡢ่⃭ᙉᗘ౫Ꮡⓗ࡞ኚ໬ࡀࠊࢥ࣮ࢹ࢕ࣥࢢ⢭ᗘ࡟࡝ࡢࡼ࠺࡟ᙳ㡪ࡍࡿ࠿ࡣ୙࡛᫂࠶ࡾࠊࢳ࣮ࣗࢽࣥࢢ≉ᛶࡢ᫬㛫ኚ໬ࡸ่⃭ᙉᗘ౫Ꮡⓗኚ໬ࡀಶࠎࡢ⣽⬊ᶵ⬟࡟࡝ࡢࡼ࠺࡟㛵ಀࡍࡿࡢ࠿ࡶࡲࡔ᫂ࡽ࠿࡟࡞ࡗ࡚࠸࡞࠸ࠋ
ࡇࢀࡽࡢၥ㢟࡟ྲྀࡾ⤌ࡴ࡟ࡣࠊྠࡌฎ⌮ẁ㝵࡟࠾

− 14 −

࠸࡚ಶࠎࡢࢽ࣮ࣗࣟࣥࡀྠᐃྍ⬟࡛࠶ࡾࠊ࠿ࡘࡑ
ࢀࡽࡢᛂ⟅≉ᛶࡀከᵝ࡛࠶ࡿࡼ࠺࡞⚄⤒ࢩࢫࢸ࣒ࡀ⌮᝿ⓗ࡛࠶ࡿࠋᮏ◊✲࡛ࡣࠊࡑࡢࡼ࠺࡞ᐇ㦂ࣔࢹࣝ࡜ࡋ࡚ࠊẼὶࡢࢲ࢖ࢼ࣑ࢡࢫࢆ᳨ฟࡍࡿࢥ࢜
ࣟࢠẼὶឤぬ⣔ࢆ⏝࠸ࡓࠋࢥ࢜ࣟࢠẼὶឤぬ⣔࡛ࡣࠊẼὶࡢ᪉ྥ᝟ሗࡣ᭱⤊⭡㒊⚄⤒⠇㸦TAG㸧ෆࡢᒁᡤᅇ㊰࡟ࡼࡗ࡚ฎ⌮ࡉࢀࠊTAG ෆ࡟ᶞ≧✺㉳ࢆᣢࡘᕧ኱௓ᅾࢽ࣮ࣗࣟࣥ㸦GIs㸧ࢆྵࡴୖ⾜ᛶᢞᑕࢽ࣮ࣗࣟࣥ࡟ࡼࡗ࡚⬻࡟ఏ㐩ࡉࢀࡿࠋࡇࢀࡲ࡛
࡟ࢥ࢜ࣟࢠ࡛ࡣ 8ࠥ10 ᑐࡢ GIs ࡀྠᐃࡉࢀ࡚࠾ࡾࠊ
ࡑࢀࡒࢀࡀẼὶࡢ᪉ྥ࡟ᑐࡋ࡚␗࡞ࡿ㑅ᢥᛶࢆ♧ࡍࡇ࡜ࡀศ࠿ࡗ࡚࠸ࡿࠋࡑࡇ࡛ᡃࠎࡣࠊ␗࡞ࡿ㸶᪉ྥ࠿ࡽᵝࠎ࡞㏿ᗘࡢẼὶ่⃭ࢆ୚࠼ࠊࡑࢀ࡟ᑐ
ࡍࡿಶࠎࡢ GI ࡢᛂ⟅ࢆࠊ⣽⬊ෆグ㘓ἲࢆ⏝࠸࡚ᐃࡋࡓࠋࡑࡢ⤖ᯝࠊ่⃭᫬㛫୰ࡢ⥲ࢫࣃ࢖ࢡᩘ࡟ᇶ࡙ࡃ
᪉ྥ㑅ᢥᛶᣦᩘ㸦DSI㸧࡜ࠊ▐㛫ⓗ࡞᭱኱Ⓨⅆ㢖ᗘ࡟ᇶ࡙ࡃ DSI ࡛ࡣࠊࡑࡢ่⃭ᙉᗘ౫Ꮡᛶࡀ␗࡞ࡗ
࡚࠸ࡓࠋࡇࢀࡣ᪉ྥ᝟ሗࢆㄞࡳฟࡍ㝿ࡢ᫬㛫ᖜ࡟ࡼࡗ࡚ࠊ่⃭ᙉᗘ࡟ᑐࡍࡿࢥ࣮ࢹ࢕ࣥࢢ⢭ᗘࡢ㡹೺ᛶ㸦ࣟࣂࢫࢺࢿࢫ㸧ࡀ␗࡞ࡿࡇ࡜ࢆព࿡ࡍࡿࠋ
ࡉࡽ࡟ࠊẼὶ㏿ᗘ࡟క࠺᪉ྥࢳ࣮ࣗࢽࣥࢢ≉ᛶࡢ
ኚ໬ࡣࠊGI ࡢ✀㢮࡟ࡼࡗ࡚␗࡞ࡗ࡚࠸ࡓࠋ
౛࠼ࡤࠊ
GI 10-2 ࡢ DSI ್ࡣࠊẼὶ㏿ᗘࡀ㏿ࡃ࡞ࡗ࡚ࡶኚࢃ
ࡽ࡞࠿ࡗࡓࠋ୍᪉࡛ࠊGI 9-1 ࡛ࡣࠊ⥲ࢫࣃ࢖ࢡᩘ
࡟ᇶ࡙ࡃ DSI ್ࡣ่⃭ᙉᗘࡀ኱ࡁࡃ࡞ࡿ࡟ᚑࡗ࡚
ῶᑡࡋࡓࡀࠊ᭱኱Ⓨⅆ㢖ᗘ࡟ᇶ࡙ࡃ DSI ್ࡣኚ໬ࡋ࡞࠿ࡗࡓࠋࡇࡢ⤖ᯝࡣࠊࢥ࢜ࣟࢠẼὶឤぬ⣔࡟࠾ࡅࡿ GIs ࡢ่⃭᪉ྥࡢࢥ࣮ࢹ࢕ࣥࢢ࡟࠾࠸࡚ࠊ
่⃭ᙉᗘ࡟ᑐࡍࡿ㡹೺ᛶࡀ⣽⬊✀㛫࡛␗࡞ࡿࡇ࡜
ࢆ♧၀ࡋ࡚࠸ࡿࠋࡇࡢࢥ࣮ࢹ࢕ࣥࢢ⢭ᗘࡢ㡹ᙉᛶࡣࡑࢀࡒࢀࡢ GIs ࡢ⾜ືᏛⓗᶵ⬟࡜㛵㐃ࡍࡿࡢ࠿ࡶࡋࢀ࡞࠸ࠋ
㸰㸬ࢥ࢜ࣟࢠࡢ㡢※ᐃ఩⾜ື࡟࠾ࡅࡿ⾜ືᡓ␎ື≀ࡣどぬ㸪⫈ぬ㸪Ⴅぬ㸪☢Ẽឤぬ࡞࡝ᵝࠎ࡞ࣔࢲࣜࢸ࢕ࡢឤぬ᝟ሗࢆ⏝࠸࡚㸪┠ⓗᆅ࡬ࡢ⛣ື㸪ࡍ࡞ࢃࡕࢼࣅࢤ࣮ࢩࣙࣥࢆ⾜࠺ࠋ⛣ື⬟ຊࡢ㧗࠸᪻⹸ࡣඃࢀࡓࢼࣅࢤ࣮ࢱ࣮࡛࠶ࡾ㸪࣑ࢶࣂࢳࡸࢧ
ࣂࢡ࢔ࣜ࡞࡝ࡢࢼࣅࢤ࣮ࢩࣙࣥ⾜ື࡟㛵ࡍࡿ◊✲ࡀ⾜ࢃࢀ࡚ࡁࡓࠋࡑࡢ⤖ᯝ㸪ᙼࡽࡀኴ㝧᪉఩ࡸ೫ගࢥࣥࣃࢫ㸪ࣛࣥࢻ࣐࣮ࢡ㸪⤒㊰✚⟬ࢩࢫࢸ࣒ࢆ
⤌ࡳྜࢃࡏ࡚㧗ᗘ࡞ࢼࣅࢤ࣮ࢩࣙࣥࢆᐇ⌧ࡋ࡚࠸ࡿࡇ࡜ࡀศ࠿ࡗ࡚ࡁࡓࠋࡋ࠿ࡋ㸪่⃭※᪉఩ࡢ᳨ฟ࠿ࡽ㐠ືไᚚ࡟⮳ࡿࢼࣅࢤ࣮ࢩࣙࣥ඲యࢆ⤫ᚚࡋ࡚࠸ࡿ⚄⤒ᇶ┙ࡢ඲ᐜࡣ౫↛࡜ࡋ࡚୙࡛᫂࠶ࡿࠋ
୍᪉㸪࣓ࢫࢥ࢜ࣟࢠࡀ࢜ࢫࡢㄏᘬḷ࡟ᑐࡋ࡚᥋㏆ࡍࡿ㡢※ᐃ఩⾜ືࡣ㸪⚄⤒⾜ືᏛࡢྂ඾ⓗࢸ࣮࣐࡜ࡋ࡚◊✲ࡉࢀ࡚ࡁࡓࠋࡇࢀࡲ࡛࡟㡢※ᐃ఩࡟㔜せ࡞ㄏᘬḷࡢㄆ㆑ࡸⓎ㡢࣓࢝ࢽࢬ࣒࡟ࡘ࠸࡚ࡢሗ
࿌ࡣከ࠸ࡀ㸪ࢼࣅࢤ࣮ࢩࣙࣥࡢ᰿ᖿ࡜࡞ࡿ㡢※᪉
఩ࡢㄆ㆑ࡸࡑࢀࢆ⮬ᕫ㐠ື࡟ኚ᥮ࡍࡿ⚄⤒ᶵᵓࡣ᫂ࡽ࠿࡟࡞ࡗ࡚࠸࡞࠸ࠋࡑࡇ࡛ᡃࠎࡣ㸪㡢※ᐃ఩୰ࡢࢥ࢜ࣟࢠ࠿ࡽ⬻⚄⤒άືࢆィ ࡋ㸪㡢※᪉఩ࡢ⚄⤒⾲⌧ࡸᐃ఩㐠ືไᚚᶵᵓࢆ᫂ࡽ࠿࡟ࡍࡿࡇ
࡜ࢆ┠ⓗ࡜ࡋࡓ◊✲ࢆ㛤ጞࡋࡓࠋࡇࢀࡲ࡛ࡢࢥ࢜ࣟࢠࡢ◊✲࡛ࡣࢺࣞࢵࢻ࣑ࣝࢆ
⏝࠸ࡽࢀ࡚ࡁࡓࡀࠊࡇࡢࡼ࠺࡞⎔ቃ࡛ࡣ㡢ཎ఩⨨ࡸ㡢ᅽࡣࢥ࢜ࣟࢠࡢ⛣ື࡟కࡗ࡚ኚ໬ࡋ࡞࠸ࡓࡵ㸪ᐇ㝿ࡢ㡢㡪⎔ቃ࡟࠾ࡅࡿ㡢※ᐃ఩⾜ືࡢヲ⣽ࡣ୙࡛᫂࠶ࡗࡓࠋࡑࡇ࡛ࡲࡎࠊ࢔࣮ࣜࢼෆࢆ⮬⏤Ṍ⾜ࡍࡿࢥ࢜ࣟࢠࡀ㡢※࡬฿㐩ࡍࡿࡲ࡛ࡢ⾜ື㐣⛬ࢆㄪ࡭ࡓࠋࡑࡢ⤖ᯝ㸪᭱ึࢥ࢜ࣟࢠࡣ⊃࠸⠊ᅖࢆ᥈⣴ࡋ࡚࠸ࡿࡀࠊ✺↛㡢※࡬ᛴ㏿࡟᥋㏆ࡋ࡚࠸ࡃࡇ
࡜ࡀࢃ࠿ࡗࡓࠋᡃࠎࡣࡇࡢ᥋㏆ࣇ࢙࣮ࢬࢆᢳฟࡋ㸪ࡑࡢ๓ᚋ࡛㐠ືෆᐜࢆẚ㍑ࡋࡓ࡜ࡇࢁ㸪᥋㏆ࣇ࢙࣮ࢬ࡛ࡣṌ⾜㏿ᗘࡸṌ⾜᫬㛫ࡀቑ኱ࡋ㸪㡢※࡬ᣦྥࡍࡿ㐠ືࡀቑ࠼ࡿࡇ࡜ࢆࡳ࠸ࡔࡋࡓࠋࡲࡓ㸪ࡇࡢ᥋㏆ࣇ࢙࣮ࢫࡢ㏵୰࡛㡢่⃭ࢆᾘኻࡉࡏࡓ࡜ࡇ
ࢁ㸪Ṍ⾜ࡣ⥅⥆ࡍࡿࡀ㡢※࡬ࡢᣦྥᛶࡀῶᑡࡋࡓࠋ
ࡋࡓࡀࡗ࡚᥋㏆ࣇ࢙࣮ࢬ୰ࡶࢥ࢜ࣟࢠࡣᖖ࡟㡢※᪉఩ࢆㄆ㆑ࡋ⥆ࡅ࡚࠸ࡿࡇ࡜ࡀ♧၀ࡉࢀࡓࠋࡉࡽ
࡟㸪᥋㏆ࣇ࢙࣮ࢬ㏵୰࡛㡢※఩⨨ࢆ✺↛⛣ືࡉࡏࡓ࡜ࡇࢁ㸪ࢥ࢜ࣟࢠࡣࡍࡄ࡟⛣ືඛࡢ㡢※࡬ᐃ఩ࢆࡣࡌࡵࡿࡀ㸪ㄏᘬḷࢆᥦ♧ࡋ࡚࠿ࡽ᭱ึࡢ㡢※࡬ࡢ᥋㏆ࣇ࢙࣮ࢬࢆ㛤ጞࡍࡿࡲ࡛ࡢ᫬㛫ࡼࡾࡶ㸪㡢※఩⨨ࢆኚ໬ࡉࡏ࡚࠿ࡽ➨㸰ࡢ㡢※࡬ࡢ᥋㏆ࣇ࢙࣮ࢬࢆ㛤ጞࡍࡿࡲ࡛ࡢ᫬㛫ࡢ᪉ࡀ▷࠿ࡗࡓࠋࡍ
࡞ࢃࡕ㸪୍ᗘ᥋㏆ࣇ࢙࣮ࢬ࡟ධࡿ࡜㸪㡢※఩⨨ࡀኚ໬ࡋ࡚ࡶࡍࡄ࡟ḟࡢ㡢※࡬ᐃ఩ࢆࡣࡌࡵࡸࡍࡃ
࡞ࡗ࡚࠸ࡿࡽࡋ࠸ࠋࡋ࠿ࡋ㸪㡢่⃭ࢆᥦ♧ࡏࡎ㸪⮬Ⓨⓗ࡟Ṍ⾜ࡋ࡚࠸ࡿࢥ࢜ࣟࢠ࡟ᑐࡋ࡚㸪㐠ື୰࡟ㄏᘬḷࢆ⫈࠿ࡏࡓሙྜ࡟ࡶ㸪⣲᪩ࡃ᥋㏆ࣇ࢙࣮ࢬ࡟ධࡗࡓࡇ࡜࠿ࡽ㸪㡢※ᐃ఩࠿࡝࠺࠿࡟㛵ࢃࡽࡎ㸪Ṍ⾜㐠ື୰ࡣ᥋㏆ࣇ࢙࣮ࢬ࡟ධࡾࡸࡍ࠸࡜⪃࠼ࡽࢀࡿࠋ௒ᚋࡣࢥ࢜ࣟࢠࡢ㐠ື࡟కࡗ࡚㡢่⃭ࡢᥦ♧ࡸ఩⨨ࢆኚ໬ࡉࡏ㸪ࡉࡽ࡟㡢※ᐃ఩⾜ື࡟࠾ࡅࡿ⫈ぬ᝟ሗࡢ౑ࢃࢀ᪉ࢆ᫂ࡽ࠿࡟ࡋ࡚㸪㡢※࡬ࡢࢼࣅࢤ࣮ࢩࣙࣥࡢィ⟬ᶵᵓ࡟㏕ࡾࡓ࠸ࠋⓎ⾲ㄽᩥ1) Sakuma, T., Nishi, K., Kishimoto, K., Nakagawa, K.,
Karasuyama, M., Umezu, Y., Kajioka, S., Yamazaki,
S.J., Kimura, K.D., Matsumoto, S., Yoda, K., Fukutomi,
M., Shidara, H., Ogawa, H., Takeuchi, I. (2019)Markerless visual servo control of a servosphere forbehavior observation of a variety of wandering
animals. Advanced Robotics, 33: 134-152.
2) Iwatani, Y., Ogawa, H., Shidara, H., Sakura, M.,
Sato, T., Hojo, M.K., Honma, A., Tsurui, K. (2019)Markerless visual servo control of a servosphere forbehavior observation of a variety of wandering
animals. Advanced Robotics, 33: 183-194.
3) Someya, M. and Ogawa, H. (2018)
Multisensory enhancement of burst activity in an− 15 −

insect auditory neuron. Journal of Neurophysiology,
120㸸139-148.
ᅜ㝿఍㆟Ⓨ⾲
1㸧 Ogawa, H., Tanaka, K., Someya, M., and Shidara,
H. (2018)
Stimulus-intensity impacts on direction encoding
and its temporal dynamics in insect mechanosensory
projection neurons.
48th Annual Meeting of Society for Neuroscience,
San Diego Convention center㸦San Diego㸧
2㸧 Shidara, H., Hommaru, N., Ogawa, H. (2018)
Phonotactic behaviors in freely-moving female
crickets.
48th Annual Meeting of Society for Neuroscience,
San Diego Convention center㸦San Diego㸧
3㸧 Sato, N., Shidara, H., Ogawa, H. (2018)
Trade-off between speed, directional accuracy andbehavioral flexibility in action selection of theescape behavior in the cricket.
48th Annual Meeting of Society for Neuroscience,
San Diego Convention center㸦San Diego㸧ᅜෆᏛ఍Ⓨ⾲ᑠᕝᏹே
1㸧 ᯘ⏣㈼ᶞ㸪タᵹஂᚿ㸪⏥ᩫຍᶞ᮶㸪ᑠ㸦2019㸧
ࢥ࢜ࣟࢠ⬻ෆࡢከឤぬࢽ࣮ࣗࣟࣥࡢ␗✀่⃭࡟ᑐࡍࡿ᪉ྥ㑅ᢥᛶࡢゎᯒ᪥ᮏື≀Ꮫ఍➨ 63 ᅇ໭ᾏ㐨ᨭ㒊኱఍㸪໭ᾏ㐨኱Ꮫ㸦ᮐᖠᕷ㸧
ᑠᕝᏹே㸦2019㸧2㸧 ᮏ୸ᑦே㸪タᵹஂᚿ㸪ᑠࢥ࢜ࣟࢠࡢ㡢※ᐃ఩⾜ື࡟࠾ࡅࡿ⾜ືᡓ␎᪥ᮏື≀Ꮫ఍➨ 63 ᅇ໭ᾏ㐨ᨭ㒊኱఍㸪໭ᾏ㐨኱Ꮫ㸦ᮐᖠᕷ㸧
3㸧 ᑠᕝᏹே㸦2019㸧ࢥ࢜ࣟࢠ㡢※ᐃ఩ࢼࣅࢤ࣮ࢩࣙࣥ࡟࠾ࡅࡿ⾜ືせ⣲ᢳฟ࡜⎔ቃ᧯స➨ 66 ᅇ᪥ᮏ⏕ែᏛ఍኱఍㸪⚄ᡞᅜ㝿఍㆟ሙ㸦⚄ᡞᕷ㸧
4㸧 ᑠᕝᏹே㸪⚟ᐩཪ୕㑻㸪ᰁ㇂┿⍆㸦2019㸧ࢥ࢜ࣟࢠࡢ⫈ぬ่⃭࡟ࡼࡿ⏕ᚓⓗ⾜ືࡢಟ㣭࡜ࡑࡢ⚄⤒ᇶ┙
᪥ᮏ㡢㡪Ꮫ఍ 2019 ᖺ᫓Ꮨ◊✲Ⓨ⾲఍㸪㟁Ẽ㏻ಙ
኱Ꮫ㸦ㄪᕸᕷ㸧5㸧 タᵹஂᚿ, ᮏ୸ᑦே, ᑠᕝᏹே㸦2018㸧ࢥ࢜ࣟࢠ㡢※ᐃ఩⾜ື࡟࠾ࡅࡿࢼࣅࢤ࣮ࢩࣙࣥ
ᡓ␎
6㸧 ⏣ ୰ ᑑ ᕼ , ᰁ ㇂ ┿ ⍆ , タ ᵹ ஂ ᚿ , ᑠ ᕝ ᏹ ே㸦2018㸧
ࢥ࢜ࣟࢠẼὶᛂ⟅ᛶᢞᑕࢽ࣮ࣗࣟࣥ࡟࠾ࡅࡿ่
⃭㏿ᗘ౫Ꮡⓗ࡞᪉ྥ㑅ᢥᛶಟ㣭ࡢゎᯒ
Neuroscience 2018—➨ 41 ᅇ᪥ᮏ⚄⤒⛉Ꮫ኱఍—ࠊ⚄ᡞᅜ㝿఍㆟ሙ㸦⚄ᡞᕷ㸧ᑠᕝᏹே㸦2018㸧
7㸧 ᰁ㇂┿⍆㸪ᑠࢥ࢜ࣟࢠឤぬ⣔௓ᅾࢽ࣮ࣗࣟࣥࡢከឤぬ⤫ྜ࡟࠾ࡅࡿ⥺ᙧᛶࡣάືࣞ࣋ࣝ࡟౫Ꮡࡍࡿ
Neuroscience 2018—➨ 41 ᅇ᪥ᮏ⚄⤒⛉Ꮫ኱఍—ࠊ⚄ᡞᅜ㝿఍㆟ሙ㸦⚄ᡞᕷ㸧8㸧 ⚟ᐩཪ୕㑻, タᵹஂᚿ, ᑿཎ࿴ஓ, す࿴ᘺ, బஂ㛫ᣅே, ๓ᕝ༟ஓ, ➉ෆ୍㑻, ᑠᕝᏹே㸦2018㸧ᶵᲔᏛ⩦࡟ࡼࡿ⏕యࢼࣅࢤ࣮ࢩࣙࣥࡢ㌶㊧ศᯒ㸸ࢥ࢜ࣟࢠ㏨㑊⾜ື࡟࠾ࡅࡿ᫬⣔ิࢹ࣮ࢱゎᯒ
Neuroscience 2018—➨ 41 ᅇ᪥ᮏ⚄⤒⛉Ꮫ኱఍—ࠊ⚄ᡞᅜ㝿఍㆟ሙ㸦⚄ᡞᕷ㸧㒊㛛 ⾜ື⚄⤒⏕≀Ꮫศศ㔝⏕≀⛉Ꮫ㒊໭⏣ ୍༤
6SHHG\5LQJR ࣇ ࢓ ࣑ ࣜ ࣮ ࢱ ࣥ ࣃ ࢡ ㉁ ࡣ&\FOLQGHSHQGHQWNLQDVH &'. ࢱࣥࣃࢡ㉁࡜⤖ྜࡋࠊࢧ࢖ࢡࣜࣥࢱࣥࣃࢡ㉁ࣇ࢓࣑࣮ࣜ࡜ࡣูಶ࡟
&'. ࢆไᚚࡍࡿࡇ࡜ࡀ▱ࡽࢀ࡚࠸ࡿࠋ࣐࢘ࢫࢤࣀ࣒
࡟࠾࠸࡚ࡣࠊ6SG\D6SG\H6SG\H6SG\I ࡢ ࡘࡢ࣍ࣔࣟࢢࡀࢥ࣮ࢻࡉࢀ࡚࠸ࡿࠋࡇࢀࡽࡢࣇ࢓࣑
࣮ࣜࢱࣥࣃࢡ㉁ࡣࠊࡼࡃಖᏑࡉࢀࡓ 6SHHG\5LQJRࢻ࣓࢖ࣥࡀྵࡲࢀ࡚࠾ࡾࠊఝࡓᶵ⬟ࢆⓎ⌧ࡍࡿࡶࡢ࡜᝿ീࡉࢀ࡚ࡁࡓࠋࡋ࠿ࡋࠊ୍᫖ᖺ࡟⊂❧ࡋ࡚  ࢢ࣮ࣝࣉࡀ 6SG\D ࣀࢵࢡ࢔࢘ࢺ࣐࢘ࢫࡢሗ࿌ࢆࡋ࡚
࠾ࡾࠊ㞤㞝࡜ࡶ࡟୙ዷ࡛ῶᩘศ⿣೵Ṇࡀほᐹࡉࢀࡓࠋ
ࡍ࡞ࢃࡕࠊ6SG\D6SG\H6SG\H6SG\I ࡣᶵ⬟ศ໬ࡋ࡚࠸ࡿࡇ࡜ࡀ♧၀ࡉࢀࡓࠋ
ࡑࡇ࡛ࠊ௒ᖺᗘ࡟࠾࠸࡚ࡣ 63'<) ࢱࣥࣃࢡ㉁ࡢᢠయ
࡜ᑐ↷࡜ࡋ࡚ 63'<$ ࢱࣥࣃࢡ㉁ࡢᢠయࢆసฟࡋࠊࡑ
ࢀࡒࢀࡢᢠཎࡢศᕸ࡟㐪࠸ࡀࡳࡽࢀࡿ࠿ྰ࠿ࢆ᳨
ウࡋࡓࠋ63'<$ ࢱࣥࣃࢡ㉁ࡣࣃ࢟ࢸࣥᮇࡢࢸ࣓ࣟ࢔
࡟ᒁᅾࡋ࡚࠾ࡾࠊࡍ࡛࡟  ࢢ࣮ࣝࣉ࠿ࡽሗ࿌ࡉࢀ࡚
࠸ࡿ⤖ᯝࡀ㏣ヨࡉࢀࡓࠋ୍᪉ࠊ63'<) ࢱࣥࣃࢡ㉁ࡣ
୺࡟ࣃ࢟ࢸࣥᮇࡢᖖᰁⰍయ࡛  ࠿ᡤ࡟ࠊ;<ERG\
࡛ࡣᰁⰍయ඲య࡟ᒁᅾࡋ࡚࠾ࡾࠊ63'<$ ࢱࣥࣃࢡ㉁ࡢࡑࢀ࡜ࡣ␗࡞ࡗ࡚࠸ࡓࠋࡋࡓࡀࡗ࡚ࠊ63'<$ ࢱࣥ
ࣃࢡ㉁࡜ 63'<) ࢱࣥࣃࢡ㉁ࡢࣜࣥ㓟໬ࢱ࣮ࢤࢵࢺ
ࢱࣥࣃࢡ㉁ࡶ␗࡞ࡗ࡚࠸ࡿࡇ࡜ࡀ᝿ᐃࡉࢀࡿࠋ௒ᚋࠊ
0/+ ࢱࣥࣃࢡ㉁➼ࡢᢠయ࡜ඹᰁⰍࡍࡿࡇ࡜࡟ࡼࡾࠊ
63'<) ࢱࣥࣃࢡ㉁࡜ඹᒁᅾࡍࡿࢱࣥࣃࢡ㉁ࢆྠᐃࡋ
࡚ࠊᶵ⬟ゎᯒࢆᐇ᪋ࡋࡓ࠸ࠋ

⏕≀⛉Ꮫ㒊㒊㛛 ⾜ື⚄⤒⏕≀Ꮫศศ㔝࿴ከ ࿴ᏹ
◊✲⤖ᯝ㡢ኌⓎኌᏛ⩦ࡢ⮫⏺ᮇไᚚཬࡧࠊ✀≉␗ⓗ࡞Ⓨኌࣃࢱ࣮ࣥ⏕ᡂ࡟㛵ࢃࡿ㑇ఏᏊ⩌ࢆ᫂ࡽ࠿࡟ࡋࠊ
ࡑࡢ⬻ෆศᏊᶵ⬟ࢆ⚄⤒⾜ືᏛⓗ࡟᳨ドࡍࡿࡇ࡜ࢆ┠ⓗ࡜ࡋ࡚◊✲ࢆ㐍ࡵ࡚࠸ࡿࠋࡑࡢࡓࡵ࡟ື≀ࣔࢹࣝ࡜ࡋ࡚㬆⚺㢮ࢯࣥࢢࣂ࣮ࢻࢆ⏝࠸ࡓ◊✲ࢆ᪋⾜ࡋ࡚࠸ࡿࠋࣄࢺࡢゝㄒࡸᑠ㫽ࡢḷࡣࠊぶ࡞࡝௚ಶయࡢⓎኌࣃࢱ࣮ࣥࢆࡲࡡࡿࡇ࡜࡛ᚋኳⓗ࡟⋓ᚓࡉࢀࡿࠋࡇࡢⓎኌᏛ⩦࡟ࡣࠊᏛ⩦ࡀຠ⋡ࡼࡃ㐍ࡴ᫬ᮇࠊࡍ࡞ࢃࡕᏛ⩦⮫⏺ᮇ㸦ឤཷᛶᮇ㸧ࡀ࠶ࡿ− 16 −

ࡇ࡜ࡀ▱ࡽࢀ࡚࠸ࡿࠋࡋ࠿ࡋ㸪⬻ෆ࡛ⓎኌᏛ⩦ࡢ⮫⏺ᮇࡀ⤊஢ࡍࡿ࣓࢝ࢽࢬ࣒ࡣṤ࡝᫂ࡽ࠿࡟ࡉࢀ
࡚࠸࡞࠿ࡗࡓࠋ
㬆⚺㢮ࢫࢬ࣓ள┠ࡢ࢟ࣥ࢝ࢳࣙ࢘ࡣ㸪Ꮴ໬ᚋ 㹼 ᪥ࡢ⣙  ࢝᭶ࡢ㛫㸦Ꮫ⩦⮫⏺ᮇ㸧࡟  ᪥ᩘⓒ
ᅇ௨ୖࡢⓎኌ⦎⩦ࢆ⧞ࡾ㏉ࡍࡇ࡜࡟ࡼࡾ⮬ศࡢḷࢆ᏶ᡂࡉࡏࠊ᏶ᡂࡋࡓḷࡣࡑࡢᚋ୍⏕ᾭ⥔ᣢࡉࢀࡿࠋ᫖ᖺᗘࡢ◊✲ࢆྵࡵࡇࢀࡲ࡛࡟ࠊࡇࡢᏛ⩦⮫⏺ᮇࡀ༢࡟⏕ࡲࢀ࡚࠿ࡽࡢ᪥ᩘ㸦᪥㱋㸧࡛Ỵࡲࡿࡢ࠿ࠊࡑࢀ࡜ࡶⓎኌ⦎⩦⾜ືࡢ✚ࡳ㔜ࡡ࡟ࡼࡾไ
ᚚࡉࢀ࡚࠸ࡿࡢ࠿ࢆㄪ࡭ࡿࡓࡵࠊᏛ⩦⮫⏺ᮇ୰ࡢ⮬Ⓨⓗ࡞Ⓨኌ⦎⩦ࢆ㜼ᐖࡍࡿᐇ㦂ࢆ᪋⾜ࡋࡓࠋࡑࡢ⤖ᯝࠊⓎኌ⦎⩦ࢆ㜼ᐖࡋࡓ㫽ࡣᮏ᮶࡛࠶ࢀࡤᏛ
⩦ࡀ࡛ࡁ࡞ࡃ࡞ࡗ࡚࠸ࡿࡣࡎࡢᡂ㫽࡟࡞ࡗ࡚ࡶࠊ
ᗂ㫽ࡢࡼ࠺࡞ᮍ⇍࡞ḷࢆฟࡋࠊࡉࡽ࡟ࡑࡢ᫬Ⅼ࠿
ࡽ࡛ࡶⓎኌᏛ⩦ࡀ࡛ࡁࡿࡇ࡜ࡀ᫂ࡽ࠿࡟࡞ࡗࡓࠋ
ḟ࡟ࠊࡇࡢⓎኌ⤒㦂࡟ࡼࡿᏛ⩦⬟ຊࡢኚ໬ࡀ⬻ෆ࡛࡝ࡢࡼ࠺࡞ศᏊ࣓࢝ࢽࢬ࣒࡛ไᚚࡉࢀ࡚࠸ࡿࡢ
࠿ࢆㄪ࡭ࡓࠋ⬻ෆ࡛ㄞࡳࡔࡉࢀ࡚࠸ࡿ㑇ఏᏊ⩌ࢆḟୡ௦ࢩ࣮ࢡ࢚ࣥࢫ࡟ࡼࡾ඲ࢤࣀ࣒࡛ࣞ࣋ࣝㄪ࡭
ࡓ⤖ᯝࠊ⬻ෆࡢⓎኌᏛ⩦࡟㛵ࢃࡿ⚄⤒ᅇ㊰࡟࠾࠸࡚ࠊⓎኌ⦎⩦᫬࡟ࡔࡅㄞࡳࡔࡉࢀࠊⓎኌ⦎⩦ࡢ✚ࡳ㔜ࡡ࡟ࡼࡾㄞࡳࡔࡉࢀ࡟ࡃࡃ࡞ࡗ࡚࠸ࡃ㑇ఏᏊࡢ୍⩌ࢆⓎぢࡋࡓࠋࡇࢀࡽࡢ⤖ᯝࡣࠊⓎኌ⦎⩦⾜ືࡀࡇࢀࡽࡢ㑇ఏᏊࡢ࿧ࡧฟࡋㄪ⠇ࢆ௓ࡋ࡚ࠊᏛ
⩦⮫⏺ᮇࢆไᚚࡋ࡚࠸ࡿྍ⬟ᛶࢆ♧ࡋ࡚࠸ࡿࠋㄽᩥⓎ⾲
Hayase S, Wang H, Ohgushi E, Kobayashi M, Mori C,
Horita H, Mineta K, Liu WC, Wada K. (2018) Vocal
practice regulates singing activity-dependent genes
underlying age-independent vocal learning in songbirds.
PLoS Biology 16:e2006537.
Asogwa NC, Mori C, Sánchez-Valpuesta M, Hayase S,
Wada K. (2018) Inter- and intra-specific differences inmuscarinic acetylcholine receptor expression in theneural pathways for vocal learning in songbirds.
Journal of Comparative Neurology 526:2856-2869.
Hayase S, Wada K. (2018) Singing activity-driven Arc
expression associated with vocal acoustic plasticity injuvenile songbird. European Journal of Neuroscience
48: 1728-1742.
Mori C, Liu WC, Wada K. (2018) Recurrent developmentof song idiosyncrasy without auditory inputs in thecanary, an open-ended vocal learner. Scientific Reports
8:8732.
Ꮫ఍Ⓨ⾲
࿴ከ࿴ᏹ㬆⚺㢮ࢯࣥࢢࣂ࣮ࢻࡢḷᏛ⩦ಶయᕪࢆࡘࡃࡿ⏕ᚓⓗᏛ⩦ࣂ࢖࢔ࢫ᪥ᮏ㐍໬Ꮫ఍➨ ᅇ኱఍ ᖺ  ᭶  ᪥ᮾி኱Ꮫ㥖ሙ࢟ࣕࣥࣃࢫ
Sánchez-Valpuesta Miguel, Yukino Shibata, Asogwa
Chinweike Norman, and Kazuhiro Wada: Cell-type
specific investigation of cortical striatal premotor
neurons in song learning. Society for Neuroscience
meeting, 2018, November 5, San Diego, USA.  㒊㛛 ⾜ື⚄⤒⏕≀Ꮫศศ㔝⏕≀⛉Ꮫ㒊⏣୰ ᬸ᫂
㸦ᾏỈ฼⏝㸧
ᮌෆ࿴⚽ࠊ⏣୰ᬸ᫂㸸ࣄ࣓࢖࢝ࡢ⭎ෆ㒊ࡢ⚄⤒ᵓ㐀ゎᯒ➨㸯ᅇ࢖࢝ࢱࢥ◊✲఍ ᖺ  ᭶  ᪥㞃ᒱ⮫ᾏᐇ㦂ᡤ
ᑠἨඖẎࠊ⏣୰ᬸ᫂㸸ࣄ࣓࢖࢝ࡢ⬻ࡢᵓ㐀࡟ࡘ࠸࡚➨㸯ᅇ࢖࢝ࢱࢥ◊✲఍ ᖺ  ᭶  ᪥㞃ᒱ⮫ᾏᐇ㦂ᡤ
Tanaka N㸸Three-dimensional brain atlas of pygmy
squid, Idiosepius paradoxus, revealing the largest
relative vertical lobe system among the cephalopods.
Janelia Conference "Neuro-evo: A Comparative
Approach to Cracking Circuit Function II" 2018 ᖺ
5 ᭶ 8 ᪥. Janelia Farm (USA).

⏕≀⛉Ꮫ㒊㒊㛛 ⏕ṪⓎ⏕⏕≀ᏛᏛศ㔝ᒣୗ ṇව⬨᳝ື≀࡟࠾ࡅࡿ⏕Ṫ⣽⬊ᙧᡂࡢไᚚᶵᵓ
ࢮࣈࣛࣇ࢕ࢵࢩࣗ༸ẕ⣽⬊࡟࠾ࡅࡿ F\FOLQ%
P51$ࡢᚤ⣽ᵓ㐀ࣄࢺࢆྵࡴከࡃࡢ⬨᳝ື≀࡟࠾࠸࡚ࠊῶᩘ➨୍ศ⿣๓ᮇࡢ༸ẕ⣽⬊㸦ᮍᡂ⇍༸㸧ࡣ༸ᡂ⇍ㄏ㉳࣍ࣝࣔࣥࡢ่⃭ࢆཷࡅ࡚➨஧ศ⿣୰ᮇ࡟㐩ࡋࠊཷ⢭ྍ⬟࡞ᡂ⇍༸࡜࡞ࡿࠋࡇࡢ㐣⛬ࡣ༸ᡂ⇍࡜࿧ࡤࢀࠊ
༸ࡀཷ⢭࣭Ⓨ⏕⬟ࢆ⋓ᚓࡍࡿ㔜せ࡞㐣⛬࡛࠶ࡿࠋゐ፹ࢧࣈࣘࢽࢵࢺࡢ &GF ࢱࣥࣃࢡ㉁࡜ࡑࡢไᚚ
ࢱࣥࣃࢡ㉁࡛࠶ࡿ &\FOLQ% ࡀ⤖ྜࡋࡓ༸ᡂ⇍ಁ㐍ᅉᏊ㸦03)㸧ࡀ༸ẕ⣽⬊ෆ࡛ᙧᡂ࣭άᛶ໬ࡉࢀࡿ
ࡇ࡜࡛༸ᡂ⇍ࡣㄏ㉳ࡉࢀࡿࡀࠊከࡃࡢ⬨᳝ື≀ࡢᮍᡂ⇍༸࡟ࡣ &GF ࢱࣥࣃࢡ㉁ࡣᏑᅾࡍࡿࡀࠊ&\FOLQ% ࢱࣥࣃࢡ㉁ࡣᏑᅾࡏࡎࠊࡑࡢP51$ ࡀ⩻ヂᢚไࡉࢀࡓ≧ែ࡛Ꮡᅾࡋ࡚࠸ࡿࠋ༸ᡂ⇍ㄏ㉳࣍ࣝࣔࣥ࡟ࡼࡗ࡚ F\FOLQ%P51$ ࡢ⩻ヂᢚไࡀゎ㝖
ࡉࢀ࡚ &\FOLQ% ࢱࣥࣃࢡ㉁ࡀ᪂つྜᡂࡉࢀࡿ࡜ࠊ
᪤Ꮡࡢ &GF ࢱࣥࣃࢡ㉁࡜」ྜయࢆᙧᡂࡋ࡚ 03) ࡜− 17 −

࡞ࡿࠋ03) ࡣࡑࡢᚋࠊࣜࣥ㓟໬࡟ࡼࡗ࡚άᛶ໬ࡉ
ࢀࠊ༸ᡂ⇍ࢆᘬࡁ㉳ࡇࡍࠋࢮࣈࣛࣇ࢕ࢵࢩࣗᮍᡂ⇍༸୰࡛ࠊF\FOLQ%P51$
ࡣື≀ᴟ┤ୗࡢ⣽⬊㉁࡟⩻ヂࢆᢚไࡉࢀࡓ≧ែ࡛ᒁᅾࡍࡿࠋF\FOLQ%P51$ ࡢ⩻ヂࡢሙᡤ࡜ࢱ࢖࣑
ࣥࢢࡢཝᐦ࡞ไᚚࡀࠊṇᖖ࡞༸ᡂ⇍୪ࡧ࡟ῶᩘศ
⿣ࡢ㐍⾜࡟୙ྍḞ࡛࠶ࡿࠋP51$ ࡢᒁᅾࡸ⩻ヂ᫬ᮇࡢไᚚ࡟ࡣ⣽⬊㦵᱁࡜」ᩘࡢ⩻ヂไᚚࢱࣥࣃࢡ㉁ࡀ㛵୚ࡋ࡚࠸ࡿ࡜⪃࠼ࡽࢀࡿࡀࠊࡑࡢヲ⣽ࡣᮍࡔ୙࡛᫂࠶ࡿࠋࡇࢀࡲ࡛ࠊࢮࣈࣛࣇ࢕ࢵࢩࣗ༸ẕ⣽⬊࡟࠾ࡅࡿ F\FOLQ%P51$ ⩻ヂࡢ᫬✵㛫≉␗ⓗไᚚࡢศᏊᶵᵓ࡟ࡘ࠸࡚ࠊ୺࡟⏕໬Ꮫⓗᡭἲࢆ⏝࠸ࡓ◊✲ࡀᮏ◊✲ᐊ࡟࠾࠸࡚㐍ࡵࡽࢀࠊ3XPLOLRࠊ
,03ࠊ6WDXIHQ ➼ࡢ 51$ ⤖ྜࢱࣥࣃࢡ㉁ࡢᶵ⬟ࡀ᫂ࡽ࠿࡟ࡉࢀ࡚࠸ࡿࠋ୍᪉ࠊF\FOLQ%P51$ ࡢᙧែ࡟ࡘ࠸࡚ࡣගᏛ㢧ᚤ㙾ࣞ࣋ࣝࡢほᐹࡋ࠿⾜ࢃࢀ࡚࠾ࡽࡎࠊࡑࢀ࡟ࡼࡿ࡜ࠊᮍᡂ⇍༸࡛⩻ヂᢚไࡉ
ࢀࡓ F\FOLQ%P51$ ࡣ㢛⢏≧ᵓ㐀ࢆ࡜ࡾࠊ⩻ヂゎ㝖࡜ྠ᫬࡟㢛⢏ᵓ㐀ࡀᾘኻࡍࡿࡇ࡜ࡀ♧ࡉࢀ࡚࠸ࡿࡔࡅ࡛ࠊ⩻ヂ㛤ጞ๓ᚋ࡟࠾ࡅࡿ㟁Ꮚ㢧ᚤ㙾ࣞ࣋ࣝࡢヲࡋ࠸ᵓ㐀ኚ໬ࡣ᫂ࡽ࠿࡟ࡉࢀ࡚࠸࡞࠸ࠋ
ࢮࣈࣛࣇ࢕ࢵࢩࣗ༸ẕ⣽⬊ࡢື≀ᴟ┤ୗ࡟Ꮡᅾࡍࡿ F\FOLQ%P51$ ࡢᚤ⣽ᵓ㐀ࢆࠊ㟁Ꮚ㢧ᚤ㙾
LQVLWXK\EULGL]DWLRQ㸦(0,6+㸧ἲࢆ⏝࠸࡚㏱㐣ᆺ㟁Ꮚ㢧ᚤ㙾࡛ほᐹࡋࡓࠋࡑࡢ⤖ᯝࠊື≀ᴟഃ⣽⬊㉁࡟ࡣ༸㯤ࡸ⾲ᒙ⬊࡜ࡣ␗࡞ࡾࠊཎᙧ㉁⭷࡛ᅖࡲࢀ࡚࠸࡞࠸ᑠࡉ࡞㢛⢏≧ࡢᵓ㐀㸦ᚤ㢛⢏ᵓ㐀㸧ࡀぢࡽࢀࠊࡑࡇ࡟ F\FOLQ%P51$ ࡀᏑᅾࡍࡿࡇ࡜
ࡀ᫂ࡽ࠿࡜࡞ࡗࡓࠋࡲࡓࠊ(0,6+ ἲ࡜㟁Ꮚ㢧ᚤ㙾ච␿⤌⧊໬Ꮫἲࢆే⏝ࡋࠊึᮇ༸ẕ⣽⬊࡟࠾ࡅࡿ
F\FOLQ%P51$ ࡜ 6WDXIHQ ࢆྠ୍ษ∦ୖ࡛ྠ᫬࡟᳨ฟࡋࡓ⤖ᯝࠊF\FOLQ%P51$ࡀᏑᅾࡍࡿᚤ㢛⢏ᵓ㐀ࡢእഃ࡟ 6WDXIHQ ࡀ㓄⨨ࡍࡿࡇ࡜ࡀ᫂ࡽ
࠿࡜࡞ࡗࡓࠋ6WDXIHQ ࡢࡼ࠺࡞⩻ヂไᚚࢱࣥࣃࢡ㉁ࡣ P51$ ࡢ  㠀⩻ヂ㡿ᇦ  875 ࡟⤖ྜࡍࡿࡇ࡜ࡀከ࠸ࡇ࡜࠿ࡽࠊF\FOLQ%P51$ ࡣᚤ㢛⢏ෆ࡛
 875 ࢆእഃ࡟ྥࡅ࡚㓄⨨ࡉࢀ࡚࠸ࡿࡇ࡜ࡀ♧၀ࡉࢀࡓࠋࡑࡇ࡛ࠊ 875ࠊ⩻ヂ㡿ᇦ 25) ࠊ 875ࢆࡑࢀࡒࢀㄆ㆑ࡍࡿ 51$ ࣉ࣮ࣟࣈࢆస〇ࡋࠊᚤ㢛⢏ෆ࡟࠾ࡅࡿ F\FOLQ%P51$ ࡢ㓄⨨ࢆほᐹࡋࡓ⤖ᯝࠊ25) ࡣᚤ㢛⢏ࡢ୰ኸࠊ 875 ࡜  875 ࡣᚤ㢛⢏ࡢእഃ࡟Ꮡᅾࡍࡿࡇ࡜ࡀ᫂ࡽ࠿࡜࡞ࡗࡓࠋ
௨ୖࠊࢮࣈࣛࣇ࢕ࢵࢩࣗ༸ẕ⣽⬊࡟࠾ࡅࡿ
F\FOLQ%P51$ ࡢᚤ⣽ᵓ㐀࡜⩻ヂ㛤ጞ๓ᚋࡢᵓ㐀ኚ໬ࡢほᐹࡀྍ⬟࡟࡞ࡗࡓࠋᚑ᮶࠿ࡽ⵳✚ࡉࢀ࡚࠸ࡿ⏕໬Ꮫⓗ࡞ゎᯒ࡟ࡼࡿ⩻ヂไᚚᶵᵓࡢศᏊࣞ
࣋ࣝࡢ⌮ゎ࡟ຍ࠼ࠊᚤ⣽ᙧែᏛⓗഃ㠃࠿ࡽࡶF\FOLQ%P51$ࡢ⩻ヂไᚚᶵᵓࢆ⪃ᐹ࡛ࡁࡿ㐨ࡀ㛤࠿ࢀࡓࠋࡇࢀࡽศᏊᙧែᏛⓗゎᯒ࡟ࡼࡾࠊࢮࣈ
ࣛࣇ࢕ࢵࢩࣗ༸ẕ⣽⬊࡟࠾ࡅࡿ F\FOLQ%P51$ ࡢ⩻ヂᢚไ࡜ࠊ่⃭࣍ࣝࣔࣥ࡟ࡼࡿゎ㝖ᶵᵓࡢヲ⣽

࡟㏕ࡿࡇ࡜ࡀ࡛ࡁࡿ࡜ᮇᚅࡉࢀࡿࠋㄽᩥⓎ⾲Takahashi, K., Ishii, K., and Yamashita, M. (2018)
Staufen1, Kinesin1 and microtubule function in cyclin
B1 mRNA transport to the animal polar cytoplasm ofzebrafish oocytes. Biochemical and BiophysicalResearch Communications 503: 2778-2783.
Ꮫ఍Ⓨ⾲ⴗཎⲁ㸪Ⲷཎඞ┈㸪ᒣୗṇව㸪⾜ಇ⏤ோ㸪㧗ᶫᏕ⾜࣓ࢲ࢝᤼༸๓ℐ⬊࡟࠾ࡅࡿ*$3⤖ྜࡢ᤼༸࡟ᑐࡍࡿᙺ๭᪥ᮏື≀Ꮫ఍➨  ᅇ኱఍㸪 ᖺ  ᭶  ᪥㸪ᮐᖠࢥࣥ࣋ࣥࢩࣙࣥࢭࣥࢱ࣮
㸦ᮐᖠᕷ㸧
బ⸨❳୍㸪ᒣୗṇවࢧ࢖ࢺ࢝࢖ࣥⓑ⾑⑓ᢚไᅉᏊ㸦/LI㸧ࡢ࣓ࢲ࢝⢭ᕢ࡟࠾ࡅࡿⓎ⌧ゎᯒ࡜⣽⬊ᇵ㣴⣔ࢆ⏝࠸ࡓᶵ⬟ゎᯒ᪥ᮏື≀Ꮫ఍໭ᾏ㐨ᨭ㒊➨  ᅇ኱఍㸪 ᖺ  ᭶  ᪥㸪໭ᾏ㐨኱Ꮫ⌮Ꮫ㒊㸦ᮐᖠᕷ㸧ⴭ᭩ᒣୗṇව㸦㸧ື≀ࡢⓎ⏕㓄അᏊᙧᡂ࣮༸
࡜⢭Ꮚ⏕࿨ࡢ㐃⥆ᛶ࡜ከᵝᛶࡢ※ࠕື≀Ꮫࡢⓒ⛉
஦඾ࠖSS୸ၿฟ∧㒊㛛 ⏕ṪⓎ⏕⏕≀Ꮫ
Ꮫศ㔝
⏕≀⛉Ꮫ㒊
㯮ᒾ 㯞㔛
ᡃࠎࡀ◊✲ᑐ㇟࡜ࡋ࡚࠸ࡿ࢔࣐࣑ࢺࢤࢿࢬ࣑
(Tokudaia osimensis) ࡜ ࢜ ࢟ ࢼ ࣡ ࢺ ࢤ ࢿ ࢬ ࣑(Tokudaia muenninki) ࡣࠊࡑࡢᛶᰁⰍయ࡜ᛶỴᐃࡢศᏊ࣓࢝ࢽࢬ࣒࡟ࣘࢽ࣮ࢡ࡞≉ᚩࢆࡶࡘࠋࡑࡇ࡛ࠊ
ࢺࢤࢿࢬ࣑ᒓࡢᛶᰁⰍయ㐍໬ࡢ㐣⛬࠾ࡼࡧᛶỴᐃศᏊ࣓࢝ࢽࢬ࣒ࢆ᫂ࡽ࠿࡟ࡍࡿࡇ࡜ࢆ┠ⓗ࡜ࡋ࡚ࠊࢺࢤࢿࢬ࣑ᒓ㸰✀ࡢ BAC ࣛ࢖ࣈ࣮ࣛࣜࢆస〇ࡋࠊࢤࣀ࣒すᲷ࡟࡚ಖ⟶ࠊ฼⏝ࡋ࡚࠸ࡿࠋಖ⟶ࡋ࡚࠸
ࡿ BAC ࣛ࢖ࣈ࣮ࣛࣜࡣࠊ࢔࣐࣑ࢺࢤࢿࢬ࣑ࡀ 384
࢙࢘ࣝࣉ࣮ࣞࢺ 255 ᯛࠊ࢜࢟ࢼ࣡ࢺࢤࢿࢬ࣑ࡀ 384
࢙࢘ࣝࣉ࣮ࣞࢺ 420 ᯛ࡛࠶ࡿࠋࡇࢀࡽࡢ BAC ࣛ࢖ࣈ࣮ࣛࣜࡣ⊂❧⾜ᨻἲே ⌮໬Ꮫ◊✲ᡤ ච␿࣭࢔ࣞࣝࢠ࣮⛉Ꮫ ⥲ྜ◊✲ࢭࣥࢱ࣮㸦⚄ዉᕝ┴ᶓ὾ᕷ㸧࡛సᡂࠊಖᏑࡉࢀ࡚࠸ࡓࡶࡢ࡛࠶ࡿࠋᕼᑡ࡞
ࢧࣥࣉࣝࢆ᪥ᮏྛᆅ࡟ศᩓࡉࡏࠊ⅏ᐖ᫬ࡢᦆኻࢆ
᭱ᑠ㝈࡟ᢚ࠼ࡿ┠ⓗ࠿ࡽࡶࠊᙜࢭࣥࢱ࣮࡟࡚ಖ⟶ࡋ࡚࠸ࡿࠋ Y ᰁⰍయࢆኻ࠸ࠊ㞤㞝࡜ࡶ࡟ XO ᆺࡢ࢔࣐࣑ࢺ
ࢤࢿࢬ࣑࡛ࡣࠊᡃࠎࡢඛ⾜◊✲࡟࠾࠸࡚ࠊY ᰁⰍయࡢ୍㒊ࡢ㑇ఏᏊࡀ X ᰁⰍయ㛗⭎ᮎ➃㒊࡟㌿ᗙࡍࡿࡇ࡜࡟ࡼࡾࠊ㞤㞝ࡢࢤࣀ࣒୰࡟ṧࡉࢀ࡚࠸ࡿࡇ

− 18 −

࡜ࡀࢃ࠿ࡗ࡚࠸ࡿࠋࡑࡇ࡛ࠊX ᰁⰍయ࡟Ꮡᅾࡍࡿ
ඖ Y ᰁⰍయ㡿ᇦࡢ඲ࢤࣀ࣒㓄ิࢆ᫂ࡽ࠿࡟ࡍࡿࡓ
ࡵ࡟ࠊඖ Y 㡿ᇦࡢ㓄ิࢆྵࡴ BAC ࢡ࣮ࣟࣥࢆࢫ
ࢡ࣮ࣜࢽࣥࢢ࡟ࡼࡾᚓ࡚ࠊࢤࣀ࣒㓄ิࢆỴᐃࡋࡓࠋࡑࡢ⤖ᯝࠊඖ Y 㡿ᇦ࡟ࡣ 6 ✀㢮ࡢ Y 㑇ఏᏊ࠾ࡼࡧ
㸯␒ᰁⰍయࡢ୍㒊ࡢ㓄ิࡀྵࡲࢀ࡚࠸ࡿࡇ࡜ࡀ᫂
ࡽ࠿࡜࡞ࡗࡓࠋࡇࢀࡽࡢ⤖ᯝࡣࠊ6 ✀㢮ࡢ Y 㑇ఏᏊࡣࠊ⢭Ꮚᙧᡂ➼࡟ാࡁࠊ㞝ᛶ࡟ᚲ㡲ࡢ㑇ఏᏊ࡛࠶ࡿࡇ࡜ࠊࡉࡽ࡟ࠊ1 ␒ᰁⰍయࡢ୍㒊ࡢ㓄ิࡀྵࡲࢀ࡚࠸ࡿࡇ࡜࠿ࡽࠊY 㡿ᇦࡢ X ᰁⰍయ࡬ࡢ㌿ᗙࡣࠊ௚ࡢᰁⰍయࢆᕳࡁ㎸ࡴ」㞧࡞ᵓ㐀ኚ໬࡛࠶ࡗ
ࡓࡇ࡜ࡀ♧၀ࡉࢀࡓࠋ ࡲࡓࠊ့ங㢮ࡢ Y ᰁⰍయୖࡢ㑇ఏᏊࡣࠊ࢜ࢫࡢ
⬻࡛ࡶⓎ⌧ࡋ࡚࠸ࡿࡀࠊࡑࡢᶵ⬟ࡸ㔜せᛶ࡟ࡘ࠸࡚ࡣ୙᫂࡞Ⅼࡀከ࠸ࠋࡑࡇ࡛ࠊ⡿ᅜ࣑ࢬ࣮ࣜ኱Ꮫ
ࡢ◊✲ࢢ࣮ࣝࣉ࡜ࡢඹྠ◊✲࡟ࡼࡾࠊ࢔࣐࣑ࢺࢤ
࣑ࡢ㞤㞝ࡢ⬻࡛Ⓨ⌧ࡋ࡚࠸ࡿඖ Y 㑇ఏᏊࡢ✀㢮ࠊ
Ⓨ⌧㔞ࢆ RNA-seq ࡟ࡼࡾ☜ㄆࡋࡓࠋ ࡲࡓࠊ㫽㢮࡟࠾ࡅࡿ⏕ṪⓎ⏕Ꮫ◊✲ࢆᒎ㛤ࡍࡿࡇ࡜ࢆ┠ⓗ࡜ࡋ࡚ࠊࢤࣀ࣒ᮾᲷ࡟࡚ࢽ࣍ࣥ࢘ࢬࣛ
(Coturnix japonica) ࢆ㣫⫱ࡋ࡚࠸ࡿࠋᮏᖺᗘࡣࠊCRISPR/Cas9 ࡢ⣔ࢆ⏝࠸ࡓࢽ࣍ࣥ࢘ࢬࣛ࡟࠾ࡅࡿ
ࢤࣀ࣒⦅㞟ᢏࢆ☜❧ࡍࡿࡓࡵ࡟ࠊ㸯⣽⬊ᮇཷ⢭༸࠾ࡼࡧ⬇┙ⴥࢆᑐ㇟࡜ࡋࡓᐇ㦂ࢆ⾜ࡗࡓࠋࡉࡽ࡟ࠊ
ᮏ✀࡟࠾ࡅࡿᛶỴᐃ࠾ࡼࡧᛶศ໬㛵㐃㑇ఏᏊࡢྠᐃࠊྛ㑇ఏᏊࡢⓎ⌧ᵝᘧࡢ☜ㄆࢆ⾜ࡗࡓࠋࡲࡓࠊ
」ᩘࡢⓎ⏕ẁ㝵࡟࠶ࡿึᮇ⏕Ṫ⭢࡟࠾࠸࡚RNA-seq ゎᯒࢆᐇ᪋ࡋࠊᚓࡽࢀࡓࢹ࣮ࢱࢆ㞤㞝㛫࡛ẚ㍑ࡍࡿࡇ࡜࠿ࡽࠊᛶỴᐃ࡟㛵ࢃࡿ W ᰁⰍయୖࡢ᪂つ㌿෗⏘≀ࡢྠᐃࢆ⾜ࡗࡓࠋㄽᩥⓎ⾲࣭Ꮫ఍Ⓨ⾲➼
1. Aduma, N., Izumi, H., Mizushima, S., Kuroiwa,
A. (2018) Knockdown of DEAD-box helicase 4(DDX4) decreases the number of germ cells inmale and female chicken embryonic gonads.
Reprod Fertil Dev. RD18266.
2. Kuroiwa, A. (2018) Sex determination anddifferentiation in birds. “Reproductive andDevelopmental Strategies; the Continuity ofLife” (eds: Kobayashi K, Kitano T, Iwao Y,
Kondo M), Springer Japan (Tokyo).
3. Ortega, M.T., Johnson, S.A., Bivens, N., Jogahara, T.,
Kuroiwa, A., Givan, S.A., Rosenfeld, C.S. Sexual
dimorphism in brain transcriptomes of Amami
spiny rat (Tokudaia osimensis): a rodent species
where males lack the Y chromosome. BMC
Genomics. 20:87.
4. Hiyama, G., Mizushima, S., Matsuzaki, M., Tobari,Y., Choi, J.H., Ono, T., Sugita, S., Sasanami, T.
(2018) Female Japanese quail visually differentiate
testosterone-dependent male attractiveness formating preferences. Sci Rep. 8:10012.
− 19 −

5. 㯮ᒾ㯞㔛. (2018) Y ࢆࡶࡓ࡞࠸୙ᛮ㆟࡞့ங
㢮—ࢺࢤࢿࢬ࣑㸬㐃㍕ࠕ⚾ࡢᐇ㦂ື≀ࠊࡸࡗࡥࡾಶᛶὴ࡛ࡍ㸟ࠖᐇ㦂་Ꮫ 36:1938-1941.
6. Kuroiwa A: Sex chromosome evolution andsex-determining mechanism in SRY-absent XO/XO
mammals, genus Tokudaia. 8th Internationa
symposium on the biology of vertebrate sex
determination,
19th
Apr,
2018,
King
Kamehameha’s Kona Beach Hotel, Kailua-Kona,
Hawaii. (Invitation speaker)
7. Kuroiwa A: Unique sex chromosome andsex-determining mechanism in Japanese native
mammals, genus Tokudaia. Genetic Society ofAustralia 2018 and 6th Asia-Pacific Chromosome
Colloquium (GSA2018_APCC6), 5th Jul, 2018,
University of Canberra, Canberra. (Keynote lecture)
8. 㯮ᒾ㯞㔛㸸ࢺࢤࢿࢬ࣑ᒓ࡟࠾ࡅࡿᛶᰁⰍయ࡜
ᛶỴᐃ࣓࢝ࢽࢬ࣒ࡢ㐍໬㸪➨ 31 ᅇࣔࣟࢩࢾࢫ
◊✲఍㸪2018 ᖺ 6 ᭶ 22 ᪥㸪໭ᾏ㐨኱Ꮫ㸦ᮐᖠᕷ㸧㸬㸦ᣍᚅㅮ₇㸧
9. Ỉᓥ⚽ᡂ㸸࢘ࢬࣛ༸ࡢཷ⢭ᡂ❧࡟㛵ࡍࡿศᏊᶵᵓ㸪➨ 3 ᅇ᪥ᮏ␆⏘Ꮫ఍ⱝᡭ௻⏬ࢩ࣏ࣥࢪ࣒࢘㸪2018 ᖺ 8 ᭶ 8 ᪥㸪ᗈᓥ኱Ꮫ㸦໭ᗈᓥᕷ㸧㸬
㸦ᣍᚅㅮ₇㸧
10. ᯇᓮⱆ⾰㸪Ỉᓥ⚽ᡂ㸪ᗈᶫᩍ㈗㸪➲ᾉ▱ᏹ㸸⢭Ꮚ㈓ⶶ⟶ୖ⓶⣽⬊⾲㠃ࡢ⢾㙐ࡀ⢭Ꮚ౵ධ࡬ཬࡰࡍᙳ㡪㸦ཱྀ㢌Ⓨ⾲㸧㸪᪥ᮏᐙ⚺Ꮫ఍⛅Ꮨ኱఍㸪2018 ᖺ 9 ᭶ 6 ᪥㸪ᮾ໭኱Ꮫ㸦௝ྎᕷ㸧㸬
11. 㡲⏣༓ᬗ㸪Ỉᓥ⚽ᡂ㸪㯮ᒾ㯞㔛㸸CRISPR/Cas9ࢩࢫࢸ࣒ࢆ฼⏝ࡋࡓ࢘ࢬࣛࢳࣟࢩࢼ࣮ࢮ㑇ఏᏊࣀࢵࢡ࢔࢘ࢺಶయసฟࡢヨࡳ㸦࣏ࢫࢱ࣮Ⓨ⾲㸧㸪᪥ᮏື≀Ꮫ఍➨ 89 ᅇᮐᖠ኱఍㸪2018ᖺ 9 ᭶ 13 ᪥㸪ᮐᖠࢥࣥ࣋ࣥࢩࣙࣥࢭࣥࢱ࣮㸦ᮐᖠᕷ㸧㸬
12. ᕤ⸨൉㤿㸪Ỉᓥ⚽ᡂ㸪㯮ᒾ㯞㔛㸸ࢺࢤࢿࢬ࣑
ᒓ࡟࠾ࡅࡿ X ᰁⰍయ୙άᛶ໬ᶵᵓ㸦࣏ࢫࢱ࣮Ⓨ⾲㸧㸪᪥ᮏື≀Ꮫ఍➨ 89 ᅇᮐᖠ኱఍㸪2018ᖺ 9 ᭶ 13 ᪥㸪ᮐᖠࢥࣥ࣋ࣥࢩࣙࣥࢭࣥࢱ࣮㸦ᮐᖠᕷ㸧㸬
13. బ⸨ᮃ㸪Ỉᓥ⚽ᡂ㸪㯮ᒾ㯞㔛㸸ࢽ࣍ࣥ࢘ࢬࣛ
ึᮇ⬇࡟࠾ࡅࡿጞཎ⏕Ṫ⣽⬊ࡢᒁᅾゎᯒ㸦࣏ࢫࢱ࣮Ⓨ⾲㸧
㸪᪥ᮏື≀Ꮫ఍➨ 89 ᅇᮐᖠ኱఍㸪2018 ᖺ 9 ᭶ 13 ᪥㸪ᮐᖠࢥࣥ࣋ࣥࢩࣙࣥࢭࣥ
ࢱ࣮㸦ᮐᖠᕷ㸧㸬
14. ᐑᮏ῟ኴ㑻㸪Ỉᓥ⚽ᡂ㸪ዟ㔝ᮍ᮶㸪ఀ⸨Ṋᙪ㸪㯮ᒾ㯞㔛㸸ࢽ࣍ࣥ࢘ࢬࣛ࡟࠾ࡅࡿᛶศ໬㛵㐃㑇ఏᏊࡢゎᯒ㸦࣏ࢫࢱ࣮Ⓨ⾲㸧㸪᪥ᮏື≀Ꮫ఍➨ 89 ᅇᮐᖠ኱఍㸪2018 ᖺ 9 ᭶ 13 ᪥㸪ᮐᖠࢥࣥ࣋ࣥࢩࣙࣥࢭࣥࢱ࣮㸦ᮐᖠᕷ㸧㸬
15. 㯮ᒾ㯞㔛㸸ࢺࢤࢿࢬ࣑ᒓ࡟࠾ࡅࡿ SRY 㑇ఏᏊࡢᶵ⬟࡜㐍໬㸪➨ 41 ᅇ᪥ᮏศᏊ⏕≀Ꮫ఍ᖺ఍࣮࣡ࢡࢩࣙࢵࣉࠕ⬨᳝ື≀ࡢᛶỴᐃࠊᛶศ໬ࡢศᏊᶵᵓࠖ㸪2018 ᖺ 11 ᭶ 29 ᪥㸪ࣃࢩࣇ࢕ࢥᶓ὾㸦ᶓ὾ᕷ㸧㸬㸦࣮࢜࢞ࢼ࢖ࢨ࣮㸪ㅮ₇㸧
16. ᒸ㔝┿భ㸪ዟ㔝ᮍ᮶㸪Ლ㇂ᕊ㸪㯮ᒾ㯞㔛㸪ఀ⸨Ṋᙪ㸸Y ᰁⰍయࢆࡶࡓ࡞࠸࢔࣐࣑ࢺࢤࢿࢬ
࣑ࡢ㑇ఏᏊ㓄ิࢆ⏝࠸ࡓ㐍໬Ꮫⓗゎᯒ㸦࣏ࢫࢱ࣮Ⓨ⾲㸧㸪➨ 41 ᅇ᪥ᮏศᏊ⏕≀Ꮫ఍ᖺ఍㸪2018 ᖺ 11 ᭶ 30 ᪥㸪ࣃࢩࣇ࢕ࢥᶓ὾㸦ᶓ὾ᕷ㸧㸬17. ᕤ⸨൉㤿㸪Ỉᓥ⚽ᡂ㸪㯮ᒾ㯞㔛㸸ࢺࢤࢿࢬ࣑
ᒓ࡟࠾ࡅࡿ X ᰁⰍయ୙άᛶ໬ᶵᵓ㸦ཱྀ㢌Ⓨ⾲㸧㸪᪥ᮏື≀Ꮫ఍➨ 89 ᅇᮐᖠ኱఍㸪2018 ᖺ
9 ᭶ 13 ᪥㸪ᮐᖠࢥࣥ࣋ࣥࢩࣙࣥࢭࣥࢱ࣮㸦ᮐᖠᕷ㸧㸬
18. 㡲⏣༓ᬗ㸪Ỉᓥ⚽ᡂ㸪㯮ᒾ㯞㔛㸸CRISPR/Cas9ࢩࢫࢸ࣒࡟ࡼࡿࢤࣀ࣒⦅㞟࢘ࢬࣛࡢసฟ࡟
㛵ࡍࡿ◊✲㸦ཱྀ㢌Ⓨ⾲㸧㸪᪥ᮏື≀Ꮫ఍໭ᾏ㐨ᨭ㒊➨ 63 ᅇ኱఍㸪2019 ᖺ 3 ᭶ 23 ᪥㸪໭ᾏ㐨኱Ꮫ㸦ᮐᖠᕷ㸧㸬
19. Ỉᓥ⚽ᡂ㸪㯮ᒾ㯞㔛㸸㫽㢮ࡢཷ⢭ᡂ❧࡟㛵ࡍࡿ᰾ศゎ㓝⣲ࡢᙺ๭㸦ཱྀ㢌Ⓨ⾲㸧㸪᪥ᮏື≀Ꮫ఍໭ᾏ㐨ᨭ㒊➨ 63 ᅇ኱఍㸪2019 ᖺ 3 ᭶ 23᪥㸪໭ᾏ㐨኱Ꮫ㸦ᮐᖠᕷ㸧㸬20. Ỉᓥ⚽ᡂ㸪➲ᾉ▱ᏹ㸪ᑠ㔝⌔எ㸪㯮ᒾ㯞㔛㸸࢘ࢬࣛࡢ㓄അᏊ࡟Ⓨ⌧ࡍࡿ DNase I ࡢཷ⢭࡟࠾ࡅࡿᶵ⬟㸦ཱྀ㢌Ⓨ⾲㸧㸪᪥ᮏᐙ⚺Ꮫ఍᫓Ꮨ኱఍㸪2019 ᖺ3 ᭶30 ᪥㸪㯞ᕸ኱Ꮫ㸦┦ᶍཎᕷ㸧㸬21. ᯇᓮⱆ⾰㸪ᗈᶫᩍ㈗㸪㒔⠏ᨻ㉳㸪๓⏣↷ኵ㸪Ỉᓥ⚽ᡂ㸪➲ᾉ▱ᏹ㸸࢘ࢬࣛ⢭Ꮚ㈓ⶶ⟶࡬ࡢ⢭Ꮚ౵ධ࡜⢭Ꮚ➇த࡜ࡢ㛵ಀ㸦ཱྀ㢌Ⓨ⾲㸧㸪᪥ᮏᐙ⚺Ꮫ఍᫓Ꮨ኱఍㸪2019 ᖺ 3 ᭶ 30 ᪥㸪㯞ᕸ኱Ꮫ㸦┦ᶍཎᕷ㸧㸬⏕≀⛉Ꮫ㒊㒊㛛 ⏕ṪⓎ⏕⏕≀Ꮫศ㔝ᮌᮧ ᩔ
㸺◊✲ㄢ㢟ྡ㸼့ங㢮ࡢ⏕Ṫ࡟㛵ࢃࡿࢤࣀ࣒ᶵ⬟࡟㛵ࡍࡿ◊✲㸺฼⏝≧ἣ࣭ᡂᯝ㸼ᙜ◊✲ᐊ࡛ࡣࠊ့ங㢮ࡢ⏕Ṫ࡟࠾ࡅࡿࢤࣀ࣒
ᶵ⬟ࢆゎ᫂ࡍࡿࡇ࡜ࢆ┠ⓗ࡜ࡋ࡚ࠊ⏕Ṫჾᐁ࡟࠾ࡅࡿ㑇ఏᏊⓎ⌧ㄪ⠇ᶵᵓࢆㄪᰝࡋ࡚࠸ࡿࠋලయⓗ࡟ࡣ༸ᕢࠊ⢭ᕢࠊ⫾┙࡛Ⓨ⌧ࡍࡿࡉࡲࡊࡲ࡞㑇ఏᏊࡀ࡝ࡢࡼ࠺࡞࣓࢝ࢽࢬ࣒࡛ไᚚࡉࢀ࡚࠸ࡿࡢ࠿
ࢆศᏊ࡛ࣞ࣋ࣝㄪ࡭࡚࠾ࡾࠊᚲせ࡟ᛂࡌ࡚㑇ఏᏊࡢᶵ⬟ゎᯒࡶ⾜ࡗ࡚࠸ࡿࠋ௒ᖺᗘࡣ⢭ᕢ≉␗ⓗ࡞long noncoding RNA㸦lncRNA㸧࡛࠶ࡿ Tesra ࢆⓎぢࡋ࡚ᶵ⬟ゎᯒࡋࡓ◊✲࡟ࡘ࠸࡚ࠊㄽᩥⓎ⾲ࢆ⾜ࡗࡓࠋ့ங㢮ࡢࢤࣀ࣒ࡣࢱࣥࣃࢡ㉁ࢆࢥ࣮ࢻࡍࡿ㓄ิࡀ 1㹼2㸣ࡋ࠿࡞࠸୍᪉࡛ࠊࡑࡢ኱㒊ศࡀ lncRNA࡜ࡋ࡚㌿෗ࡉࢀ࡚࠸ࡿࠋ⢭ᕢࡣࠊ့ங㢮ࡢ⤌⧊ࡢ୰࡛ࡶ≉࡟ከࡃࡢ lncRNA ࢆ㌿෗ࡍࡿჾᐁ࡛࠶ࡾࠊ
⢭Ꮚᙧᡂ࡟࠾ࡅࡿ㔜せᛶࡀᣦ᦬ࡉࢀ࡚࠸ࡿࡀࠊᶵ
⬟ゎᯒࡣ㐍ࢇ࡛࠸࡞࠸ࠋᡃࠎࡣࠊ⢭ᕢ≉␗ⓗ࡞ࢭࣜࣥࣉࣟࢸ࢔࣮ࢮࢆࢥ࣮ࢻࡍࡿ㑇ఏᏊࡀ 3 ࡘ୪ࢇࡔࠊ࣐࢘ࢫ 9 ␒ᰁⰍయୖࡢ Prss/Tessp 㑇ఏᏊᗙ࠿ࡽ᪂つࡢ lncRNA ࢆⓎぢࡋ࡚ Tesra ࡜ྡ࡙ࡅࡓࠋࢡ
࣮ࣟࢽࣥࢢ࡜Ⓨ⌧ゎᯒࡢ⤖ᯝࠊTesra ࡣ඲㛗 4435
ሷᇶࡢ⢭ᕢ≉␗ⓗ࡞ lncRNA ࡛࠶ࡾࠊ⢭ཎ⣽⬊ࠊ୍ḟ⢭ẕ⣽⬊ࠊ෇ᆺ⢭⣽⬊࡛ࡣ᰾࡜⣽⬊㉁࡟ࠊఙ
㛗⢭⣽⬊࡜ࣛ࢖ࢹ࢕ࢵࣄ⣽⬊࡛ࡣ୺࡟⣽⬊㉁࡟ᒁ
ᅾࡋࠊ⢭ᕢ࢚ࢡࢯࢯ࣮࣒㸦⣽⬊እᑠ⬊㸧୰࡟ࡶᏑᅾࡍࡿࡇ࡜ࡀࢃ࠿ࡗࡓࠋ୍ḟ⢭ẕ⣽⬊࡛ࡣከࡃࡢ㔜せ㑇ఏᏊࡀ㌿෗άᛶ໬ࡍࡿࡇ࡜࠿ࡽࠊᡃࠎࡣ᰾
࡟ᒁᅾࡍࡿ Tesra ࡀࡇࢀࡽࡢ㌿෗ㄪ⠇࡟㛵ࢃࡿྍ⬟ᛶࢆ᳨ドࡋࡓࠋࡲࡎ ChIRP ゎᯒ࡜࠸࠺᪉ἲ࡛
Tesra ㌿෗⏘≀ࡀ࡝ࡢࢡ࣐ࣟࢳࣥ㡿ᇦ࡟⤖ྜࡋ࡚࠸ࡿ࠿ࢆㄪ࡭ࡓ⤖ᯝࠊTesra ࡣ࣐࢘ࢫ⢭ᕢ⏕Ṫ⣽⬊࡟࠾࠸࡚ Prss42/Tessp-2 㑇ఏᏊࡢࣉ࣮ࣟࣔࢱ࣮㡿ᇦ࡟⤖ྜࡋ࡚࠸ࡿࡇ࡜ࡀࢃ࠿ࡗࡓࠋࡑࡇ࡛ in vitro⣽⬊ᇵ㣴⣔࡟࠾࠸࡚ Tesra ࢆ㐣๫Ⓨ⌧ࡉࡏࡓ࡜ࡇ
ࢁࠊෆᅾࡢ Prss42/Tessp-2 㑇ఏᏊࡢⓎ⌧㔞ࡀୖ᪼ࡋࡓ࠺࠼ࠊ࣏࣮ࣞࢱ࣮㑇ఏᏊ࡟ࡘ࡞࠸ࡔ Prss42/Tess-2
ࣉ࣮ࣟࣔࢱ࣮ࡢάᛶࡶୖ᪼ࡋࡓࠋࡋࡓࡀࡗ࡚ Tesra ࡣ
Prss42/Tessp-2 㑇ఏᏊࡢ㌿෗άᛶ໬࡟ᶵ⬟ࡍࡿࡇ࡜ࡀ᫂ࡽ࠿࡟࡞ࡗࡓࠋࡇࡢ Prss42/Tessp-2 㑇ఏᏊ࡟ࡘ࠸࡚ࠊᡃࠎࡣࡍ࡛࡟࢚ࣥࣁࣥࢧ࣮ࡢᏑᅾࢆ᫂ࡽ
࠿࡟ࡋ࡚࠸ࡓࡢ࡛ࠊTesra ࡀࡇࡢ࢚ࣥࣁࣥࢧ࣮࡜༠
ാⓗ࡟ᶵ⬟࡛ࡁࡿ࠿ࢆ᭱ᚋ࡟᳨ドࡋࡓࠋࡑࡢ⤖ᯝࠊ࢚ࣥࣁࣥࢧ࣮࡜ Tesra ࡣ஫࠸࡟㠀౫Ꮡⓗ࡛ࡣ࠶ࡿࡶࡢࡢࠊ┦ຍⓗ࡟ Prss42/Tessp-2 㑇ఏᏊࡢ㌿෗άᛶ໬ࢆ⾜࠺ࡇ࡜ࡀࢃ࠿ࡗࡓࠋPrss42/Tessp-2 㑇ఏᏊࡣ⢭Ꮚᙧᡂ࡟࠾࠸࡚ῶᩘศ⿣ࡀ஧ḟ⢭ẕ⣽⬊࠿ࡽ⢭⣽⬊࡬࡜㐍⾜ࡍࡿ㝿࡟ᚲ㡲ࡢᙺ๭ࢆᣢࡘࡇ࡜࠿ࡽࠊࡑࡢ㌿෗άᛶ໬ࢆᢸ࠺ Tesra ࡶῶᩘศ⿣ࡢ㐍⾜࡟㔜せ࡛࠶ࡿ࡜⪃࠼ࡽࢀࡿࠋ௨ୖࡢᡂᯝࡣࠊ⢭ᕢ
lncRNA ࡢ᪂ࡓ࡞ᶵ⬟ࢆゎ᫂ࡋࡓࡔࡅ࡛࡞ࡃࠊ㏆ᖺ኱ࡁ࡞ㄢ㢟࡜࡞ࡗ࡚࠸ࡿlncRNA ࡜࢚ࣥࣁࣥࢧ࣮ࡢ
㛵ಀ࡟ࡘ࠸࡚ࡶ㔜せ࡞▱ぢࢆ୚࠼ࡿࡶࡢ࡜࡞ࡗࡓࠋ
㸺ㄽᩥⓎ⾲㸼
1. Maruyama Y. and Kimura A.P. (2019) A molecularmechanism of mouse placental spongiotrophoblast
differentiation regulated by prolyl oligopeptidase.
Zygote (in press)
2. Satoh Y., Takei N., Kawamura S., Takahashi N.,
Kotani T., and Kimura A.P. (2019) A novel
testis-specific long noncoding RNA, Tesra,
activates the Prss42/Tessp-2 gene during mouse
spermatogenesis. Biol. Reprod. (in press)
3. Takei N., Nakamura T., Kawamura S., Takada Y.,
Satoh Y., Kimura A.P., and Kotani T. (2018)− 20 −

High-sensitivity and high-resolution in situ
hybridization of coding and long non-coding
RNAs in vertebrate ovaries and testes. Biol.
Proced. Online 20: 6.
㸺Ꮫ఍Ⓨ⾲㸼
1. బ⸨ඃ⾰ࠊṊ஭ኟᾏࠊᕝᮧ⩧ᖹࠊ㧗ᶫఙᙪࠊᑠ
㇂཭ஓࠊᒣᮏ㞝ᗈࠊΏ㎶೺ᏹࠊᯇཎఙࠊబ➉⅖ࠊᮌᮧᩔࠕ࣐࢘ࢫ⢭ᕢ≉␗ⓗ lncRNA Tesra ࡢⓎ⌧ࣃࢱ࣮ࣥ࡜㌿෗άᛶ໬࡟࠾ࡅࡿᶵ⬟ࠖ➨ 41ᅇ᪥ᮏศᏊ⏕≀Ꮫ఍ᖺ఍㸦ࣃࢩࣇ࢕ࢥᶓ὾ࠊ
2018 ᖺ 11 ᭶ 28㹼30 ᪥㸧
2. 㓇஭཭㔛ࠊ㓇஭⩏ᓅࠊబ⸨ඃ⾰ࠊᮌᮧᩔࠕ࣐࢘
ࢫ⢭ẕ⣽⬊࡛ᶵ⬟ࡍࡿ dual promoter‒enhancerࡢస⏝࣓࢝ࢽࢬ࣒ࠖ➨ 41 ᅇ᪥ᮏศᏊ⏕≀Ꮫ఍ᖺ఍㸦ࣃࢩࣇ࢕ࢥᶓ὾ࠊ2018 ᖺ 11 ᭶ 28㹼30 ᪥㸧
3. ๓⏣┤ᶞࠊ୸ᒣඃᶞࠊᮌᮧᩔࠕ࣐࢘ࢫ༸ᕢ㢛⢏⭷⣽⬊࡟࠾ࡅࡿ LH ཷᐜయ㐣๫Ⓨ⌧⣽⬊ᰴࡢᶞ❧࡜ゎᯒࠖ➨ 43 ᅇ᪥ᮏẚ㍑ෆศἪᏛ఍኱఍㸦ᮾ໭኱Ꮫ㟷ⴥᒣࢥࣔࣥࢬࠊ2018 ᖺ 11 ᭶ 10 ᪥㸧
4. Bandara A.M.T.K., Matsubara S., Shiraishi A.,
Satake H., and Kimura A.P. ࠕ Transcriptional
regulation by genome regions with H3K4me1 andH3K4me3 marks in mouse spermatocytesࠖ➨ 43
ᅇ᪥ᮏẚ㍑ෆศἪᏛ఍኱఍㸦ᮾ໭኱Ꮫ㟷ⴥᒣࢥ
ࣔࣥࢬࠊ2018 ᖺ 11 ᭶ 10 ᪥㸧
5. ࠙ᣍᚅㅮ₇ࠚᮌᮧᩔࠕ࣐࢘ࢫ⢭Ꮚᙧᡂࡢ୍ḟ⢭ẕ⣽⬊࡟࠾ࡅࡿ㌿෗άᛶ໬࣓࢝ࢽࢬ࣒ࠖ᪥ᮏື
≀Ꮫ఍➨ 89 ᅇ኱఍ࢩ࣏ࣥࢪ࣒࢘㸸➨ 8 ᅇ࣓࣍࢜ࢫࢱࢩࢫࣂ࢖࢜ࣟࢪ࣮ࢩ࣏ࣥࢪ࣒࢘ࠕ⬨᳝ື≀ࡢ㓄അᏊᙧᡂ࡟࠾ࡅࡿᜏᖖᛶ⥔ᣢࡢࡓࡵࡢ
㌿෗࣭⩻ヂㄪ⠇ࠖ㸦ᮐᖠࢥࣥ࣋ࣥࢩࣙࣥࢭࣥࢱ
࣮ࠊ2018 ᖺ 9 ᭶ 15 ᪥㸧⏕≀⛉Ꮫ㒊㒊㛛 ⏕ṪⓎ⏕⏕≀ᏛᏛศ㔝ᑠ㇂ ཭ஓ
༸ẕ⣽⬊ࡣࡑࡢᡂ㛗㐣⛬࡟࠾࠸࡚ከࡃࡢ㌿෗⏘≀ࢆ⵳࠼ࡿࠋ࣐࢘ࢫ࡜ࢮࣈࣛࣇ࢕ࢵࢩࣗࢆ⏝࠸ࡓ◊✲࠿ࡽࠊࡑࡢᩘࡣ  ୓✀㢮ࢆ㉸࠼ࡿࡇ࡜ࡀ᫂ࡽ
࠿࡜࡞ࡗ࡚ࡁࡓࠋࡇࢀࡽࡢከࡃࡣ⩻ヂࢆᢚไࡉࢀ
༸⣽⬊㉁࡟Ꮡᅾࡍࡿࡓࡵࠊࢱࣥࣃࢡ㉁ࢆྜᡂࡋ࡞
࠸ࠋ᭱㏆ࡢ⥙⨶ⓗ࡞ゎᯒ࠿ࡽࠊࢮࣈࣛࣇ࢕ࢵࢩ࡛ࣗࡣ  ✀㢮㏆ࡃࡢ P51$ ࡀཷ⢭ᚋ࡟⩻ヂࡉࢀࡿࡇ࡜ࡀ♧ࡉࢀࡓࠋࡋ࠿ࡋࠊࡇࢀࡽ P51$ ࡀ༸ᙧᡂ㐣⛬ࡢ࠸ࡘ࠿ࡽ㌿෗ࡉࢀ࡝ࡢࡼ࠺࡞ᶵᵓ࡛⩻ヂࢆᢚไࡉࢀࡿࡢ࠿ࠊཷ⢭ᚋ࡟࡝ࡢࡼ࠺࡞ᶵᵓ࡛⩻ヂࡀ㛤ጞࡉࢀࡿࡢ࠿ࡣ࡯࡜ࢇ࡝ศ࠿ࡗ࡚࠸࡞࠸ࠋ
ᮏ◊✲ࡣ༸ᙧᡂ࡜⬇Ⓨ⏕࡟࠾ࡅࡿ㑇ఏᏊⓎ⌧ᶵᵓ࡟㏕ࡿࡇ࡜ࢆ┠ⓗ࡟ࠊ3RXI 㑇ఏᏊࡢⓎ⌧ࢆヲ⣽࡟ゎᯒࡋࡓࠋ3RXII ࡢⓎ⌧ࢆࣀࢵࢡ࢔࢘ࢺࡋ

ࡓ࣐࢘ࢫࡣ⬇┙⬊ᮇ࡛Ⓨ⏕ࢆ೵Ṇࡋ⮴Ṛ࡜࡞ࡿࠋ⯆࿡῝࠸ࡇ࡜࡟ࠊཷ⢭༸࡟࠾ࡅࡿ㐣๫Ⓨ⌧ࡣྠᵝ࡟⬇
┙⬊࡛ࡢⓎ⏕೵Ṇࢆᘬࡁ㉳ࡇࡍࠋ3RXI ࢆᣢࡓ࡞࠸ࢮࣈࣛࣇ࢕ࢵࢩࣗ༸ẕ⣽⬊ࡣࠊཷ⢭ᚋ࡟ෆ⬇ⴥࡢศ໬࣭ཎ⭠ᙧᡂ࡞࡝࡟␗ᖖࢆࡁࡓࡍࠋࡋࡓࡀࡗ࡚ࠊ
༸ᙧᡂ࠿ࡽึᮇⓎ⏕࡟࠾ࡅࡿ 3RXI ࢱࣥࣃࢡ㉁ࡢ
Ⓨ⌧ࡣཝᐦ࡟ไᚚࡉࢀࡿᚲせࡀ࠶ࡿࠋ࣐࢘ࢫ࡟࠾ࡅ
ࡿ࣏ࣜࢡ࣮ࣟࢼࣝᢠయࢆ⏝࠸ࡓ◊✲࠿ࡽࠊ3RXIࢱࣥࣃࢡ㉁ࡣ༸ᙧᡂ㐣⛬ࡢึᮇ࠿ࡽ᰾࡟Ⓨ⌧ࡋࠊ༑ศ࡟ᡂ㛗ࡋࡓ༸ẕ⣽⬊࡛ࡶ㧗ࣞ࣋ࣝࡢⓎ⌧ࡀ⥔ᣢࡉࢀࡿ࡜ࡉࢀ࡚ࡁࡓࠋࢱࣥࣃࢡ㉁ࡀ㧗ࣞ࣋ࣝ࡟᳨ฟ
ࡉࢀࡿ࡟ࡶ㛵ࢃࡽࡎࠊ3RXI ࡢ㌿෗⏘≀ࡀ༸ᙧᡂ㐣⛬ࡢ࡝ࡢ᫬ᮇ࠿ࡽⓎ⌧ࡍࡿࡢ࠿ࡣ୙࡛᫂࠶ࡗࡓࠋ
ᮏ◊✲࡛ࡣึࡵ࡟ࠊᡃࠎࡀ⊂⮬࡟☜❧ࡋࡓ⤌⧊࣭ჾᐁ࡟࠾ࡅࡿ 51$ ࡢ㧗ឤᗘ᳨ฟᢏ⾡ࢆ⏝࠸ࠊ3RXI㌿෗⏘≀ࡢⓎ⌧ࢆゎᯒࡋࡓࠋ3RXIP51$ ࡣཎጞ༸⬊ࡢ༸ẕ⣽⬊࡛ࡣⓎ⌧ࡀぢࡽࢀࡎࠊ୍ḟ༸⬊ࡢ༸ẕ⣽⬊࡛ึࡵ࡚Ⓨ⌧ࡀ᳨ฟࡉࢀࡓࠋࡇࢀࡣࠊ࣏ࣜࢡࣟ
࣮ࢼࣝᢠయࢆ⏝࠸ࡓࢱࣥࣃࢡ㉁ࡢ᳨ฟࡼࡾࡶ㐜࠸
᫬ᮇ࡟ P51$ ࡀⓎ⌧ࡍࡿࡇ࡜ࢆ♧ࡍࠋ
P51$ ࡢⓎ⌧ࡣࠊࡑࡢᚋࡢ༸ᡂ㛗㐣⛬࡟࠾࠸࡚ᖖ࡟⣽⬊㉁᳨࡛ฟࡉ
ࢀࡓࠋ༸ᙧᡂ㐣⛬࡜ึᮇⓎ⏕࡟࠾ࡅࡿ 3RXI ࢱࣥ
ࣃࢡ㉁ࡢⓎ⌧ࢆࠊࣔࣀࢡ࣮ࣟࢼࣝᢠయࢆ⏝࠸෌᳨ドࡋࡓࠋᡃࠎࡢ࢙࢘ࢫࢱ࣭ࣥࣈࣟࢵࢺࡢ⤖ᯝࡣࠊࡇࡢ
ᢠయࡀ 3RXI ࢱࣥࣃࢡ㉁ࡢࡳࢆㄆ㆑ࡍࡿᴟࡵ࡚≉␗ᛶࡢ㧗࠸ᢠయ࡛࠶ࡿࡇ࡜ࢆ♧ࡍࠋከࡃࡢ◊✲࡜ྠ
ᵝ࡟ࠊ3RXI ࢱࣥࣃࢡ㉁ࡢ㧗ࣞ࣋ࣝࡢⓎ⌧ࡀ⬇┙⬊ࡢෆ㒊⣽⬊ሢ࡟᳨ฟࡉࢀࡓࠋ୍᪉ࠊཷ⢭ᚋࡢ  ⣽⬊ᮇ࠿ࡽపࣞ࣋ࣝࡢⓎ⌧ࡀ᳨ฟࡉࢀࡓࠋࡋ࠿ࡋࠊ༸
ᙧᡂ㐣⛬ࡢ༸ẕ⣽⬊࡟࠾࠸࡚ࢱࣥࣃࢡ㉁ࡢⓎ⌧ࡣ᳨ฟࡉࢀ࡞࠿ࡗࡓࠋ௨ୖࡢ⤖ᯝࡣࠊ3RXI 㑇ఏᏊࡣ༸ᙧᡂࡢึᮇ࡟㌿෗ࡉࢀࡿࡶࡢࡢࠊࡑࡢ⩻ヂࡣᢚไࡉࢀ࡚࠾ࡾࠊཷ⢭ᚋࡢ  ⣽⬊ᮇ࠿ࡽ  ⣽⬊ᮇ࡟࠿ࡅ࡚⩻ヂࡀ㛤ጞࡍࡿࡇ࡜ࢆ♧၀ࡍࡿࠋࡇࢀࡣࠊ ⣽⬊ᮇ࡟࠾ࡅࡿ 3RXI ࢱࣥࣃࢡ㉁ࡢ㐣๫Ⓨ⌧ࡀⓎ⏕೵Ṇࢆᘬࡁ㉳ࡇࡍ⤖ᯝ࡜୍⮴ࡍࡿࠋࡑࡢ⩻ヂไᚚࡢゎ᫂ࡣࠊ3RXI ࢱࣥࣃࢡ㉁ࡢⓎ⌧㔞ࢆཝᐦ࡟ไᚚ
ࡍࡿ௙⤌ࡳࡢゎ᫂ࡢࡳ࡛࡞ࡃࠊ༸ᙧᡂ࠿ࡽⓎ⏕࡟࠾ࡅࡿ㑇ఏᏊⓎ⌧ᶵᵓࡢゎ᫂࡟ᴟࡵ࡚㔜せ࡛࠶ࡿࠋ
Ⓨ⾲ㄽᩥ
Saitoh, A., Takada, Y., Horie, M., Kotani, T. (2018)
Pumilio1 phosphorylation precedes translational activationof its target mRNA in zebrafish oocytes. Zygote, 26:
372-380.
Satoh, Y., Takei, N., Kawamura, S., Takahashi, N., Kotani,T., Kimura, A.P. (2019) A novel testis-specific longnoncoding RNA, Tesra, activates the Prss42/Tessp-2 geneduring mouse spermatogenesis. Biology of Reproduction,in press.

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Ꮫ఍Ⓨ⾲ᑠ㇂཭ஓ㸸ࢮࣈࣛࣇ࢕ࢵࢩࣗ࡜࣐࢘ࢫ༸ᙧᡂ࡟࠾
ࡅࡿ⩻ヂㄪ⠇ᶵᵓ. ➨ 89 ᅇ᪥ᮏື≀Ꮫ఍ᮐᖠ኱఍,2018 ᖺ9 ᭶15 ᪥, ᮐᖠࢥࣥ࣋ࣥࢩࣙࣥࢭࣥࢱ࣮㸦ᮐᖠᕷ㸧㧗⏣⿱㈗, ᑠ㇂཭ஓ㸸࣐࢘ࢫ༸ᙧᡂ࠾ࡼࡧ⬇Ⓨ⏕࡟࠾ࡅࡿ Pou5f1/Oct4 Ⓨ⌧ゎᯒ. ➨ 89 ᅇ᪥ᮏື≀Ꮫ఍ᮐᖠ኱఍, 2018 ᖺ9 ᭶13 ᪥, ᮐᖠࢥࣥ࣋ࣥࢩࣙ
ࣥࢭࣥࢱ࣮㸦ᮐᖠᕷ㸧㧗⏣⿱㈗, ᑠ㇂཭ஓ㸸့ங㢮ࡢ༸ᙧᡂ࡜⬇Ⓨ⏕࡟࠾ࡅࡿ Pou5f1/Oct4 ࡢⓎ⌧ゎᯒ. ➨ 63 ᅇ᪥ᮏື≀Ꮫ఍໭ᾏ㐨ᨭ㒊኱఍, 2019 ᖺ 3 ᭶23 ᪥, ໭ᾏ㐨኱Ꮫ⌮Ꮫ㒊 5 ྕ㤋పᒙᲷ 2 㝵㸦ᮐᖠᕷ㸧㒊㛛 ⏕ṪⓎ⏕⏕≀Ꮫ
Ꮫศ㔝
⏕≀⛉Ꮫ㒊
Ⲷཎ ඞ┈⬨᳝ື≀ࡢ⏕Ṫჾᐁࡢᶵ⬟࡟㛵ࡍࡿ◊✲㸦◊✲┠ⓗ㸧⬨᳝ື≀ࡢ⏕Ṫάືࡣ㸪どᗋୗ㒊㸦⬻㸧㸫⬻ୗᆶయ㸫⏕Ṫ⭢࠿ࡽ࡞ࡿ⏕ṪෆศἪ⣔࡟ࡼࡾㄪ⠇ࡉࢀ㸪༸ᕢ࡛ࡘࡃࡽࢀࡿ༸࡜⢭ᕢ࡛ࡘࡃࡽࢀ
ࡿ⢭Ꮚࡢྜయ㸦ཷ⢭㸧࡟ࡼࡾ᪂ࡋ࠸ಶయࡀ࡛ࡁࡿࠋ
ᙜ◊✲ᐊ࡛ࡣ㸪༸ᕢ㸪⢭ᕢ➼࡟࠾ࡅࡿᵝࠎ࡞⌧㇟ࢆศᏊ࡛ࣞ࣋ࣝゎᯒࡋࠊࡑࡢ࣓࢝ࢽࢬ࣒ࢆゎ᫂ࡍࡿࡇ࡜ࢆ┠ⓗ࡟◊✲ࢆ㐍ࡵ࡚࠸ࡿࠋ⌧ᅾࡣࠊ⬨᳝ື≀ࡢ༸ᕢᶵ⬟㸦≉࡟༸Ꮚᙧᡂࡸ᤼༸㸧࡟㛵㐃ࡍࡿᮍゎ᫂࡞ㄢ㢟࡟ྲྀࡾ⤌ࢇ࡛࠸ࡿࠋ
㸦⤒㐣࣭⤖ᯝ㸧 ᙜ◊✲ᐊ࡛ࡣ㸪ᐇ㦂ᮦᩱ࡜ࡋ࡚࣓ࢲ࢝ࡸ࣐࢘ࢫ
ࢆ⏝࠸࡚◊✲ࢆ⾜ࡗ࡚࠸ࡿࠋ࣓ࢲ࢝ࡣࠊ᤼༸ᐇ⾜࡟ᚲせ୙ྍḞ࡞㓝⣲㸦᤼༸㓝⣲㸧ࡀࡍ࡛࡟ྠᐃࡉ
ࢀ࡚࠾ࡾࠊࡉࡽ࡟⏕యእ࡛᤼༸⌧㇟ࢆほᐹ࡛ࡁࡿᇵ㣴⣔ࡀ฼⏝ྍ⬟࡞ࡇ࡜࠿ࡽࠊ᤼༸◊✲࡟㐺ࡋࡓᐇ㦂ື≀࡛࠶ࡿࠋ࣓ࢲ࢝ࢆ⏝࠸ࡓ᤼༸㛵㐃◊✲࡜ࡋ࡚௨ୗࡢ 2 ࢸ࣮࣐࡟ࡘ࠸࡚◊✲ࢆ⾜ࡗ࡚࠸ࡿࠋ
ࡲࡓࠊ࣐࢘ࢫࢆ⏝࠸ࡓ◊✲ࢸ࣮࣐࡜ࡋ࡚௨ୗࡢ◊✲ࢆ⾜ࡗ࡚࠸ࡿࠋ
㸦1㸧᤼༸㛵㐃㑇ఏᏊḞᦆ࣓ࢲ࢝ࡢస〇ᙜ◊✲ᐊࡢࡇࢀࡲ࡛ࡢ◊✲࡟ࡼࡾࠊ᤼༸┤๓࡟ᛴ⃭࡟ㄏᑟࡉࢀࡿ MT2-MMP ࡀ᤼༸ࡢ࣮࢟㓝⣲࡜ࡋ࡚ᶵ⬟ࡍࡿࡇ࡜ࡀ᫂ࡽ࠿࡜࡞ࡗ࡚࠸ࡿࠋࡲࡓࠊ
MT2-MMP ࢆྵࡴᵝࠎ࡞᤼༸㛵㐃㑇ఏᏊࡢⓎ⌧ㄏᑟ࡟᰾ෆࣉࣟࢤࢫࢳࣥཷᐜయ㸦nPR㸧ࡀ㛵୚ࡍࡿࡇ࡜ࡶ᫂ࡽ࠿࡜࡞ࡗ࡚࠸ࡿࠋࡑࡇ࡛ࠊࡇࢀࡽ᤼༸࡟㔜せ࡞ᅉᏊࡢḞᦆ࣓ࢲ࢝ࢆ CRISPR-Cas9 ࢩࢫࢸ࣒ࢆ⏝࠸࡚స〇ࡋ࡚࠸ࡿࠋMT2-MMP ࡟ࡘ࠸࡚ࡣ࣊
ࢸࣟࡢసฟࡲ࡛㐍ࢇ࡛࠸ࡿࠋ୍᪉ࠊnPR ࡟ࡘ࠸࡚ࡣࢤࣀ࣒⦅㞟࡟ᡂຌࡋࡓ⛶㨶ࡀᚓࡽࢀ࡚࠾ࡾࠊࡇ
ࢀ࠿ࡽ࣊ࢸࣟࡢసฟ࡟ྲྀࡾ᥃࠿ࡿணᐃ࡛࠶ࡿࠋ
㸦2㸧༸ẕ⣽⬊࡜ℐ⬊⣽⬊ࡢ㐃㛵ᛶ༸ẕ⣽⬊ࡣℐ⬊࡜࠸࠺ᵓ㐀యࡢ୰࡛ᡂ㛗ࡍࡿࠋ
ℐ⬊ࡣ༸ẕ⣽⬊࡜ࡑࡢ࿘ࡾࢆᅖࡴℐ⬊⣽⬊ᒙ࠿ࡽᡂࡿࡀࠊ୧⪅ࡢ㛫࡟ࡣࢥ࣑ࣗࢽࢣ̿ࢩࣙࣥࡀᏑᅾ
ࡍࡿࡇ࡜ࡀ့ங㢮࡟࠾࠸࡚ሗ࿌ࡉࢀ࡚࠸ࡿࠋࡑࡇ
࡛ࠊ၏୍ࠊ⬨᳝ື≀࡟࠾࠸࡚᤼༸ࡢศᏊᶵᵓࡀ᫂
ࡽ࠿࡜࡞ࡗ࡚࠸ࡿ࣓ࢲ࢝ࢆ⏝࠸࡚༸ẕ⣽⬊࡜ℐ⬊⣽⬊ᒙ㛫࡛ࡢࢥ࣑ࣗࢽࢣ࣮ࢩࣙࣥࡢ᭷↓ࢆㄪ࡭ࡓࠋ
༸ᕢ࠿ࡽ༢㞳ࡋࡓℐ⬊ࢆࡑࡢࡲࡲᇵ㣴ࡍࡿ࡜᤼༸
ࡢ┤๓࡟ MT2-MMP ࡀㄏᑟࡉࢀࡿࡀࠊℐ⬊ࢆ₽ࡋ࡚༸ẕ⣽⬊ࢆ◚ቯᚋ࡟ᇵ㣴ࢆ⾜࠺࡜ MT2-MMP
ࡣ᤼༸ࡢ࠿࡞ࡾ๓ࡢ᫬Ⅼ࡛ㄏᑟࡉࢀࡿࡇ࡜ࡀุ᫂
ࡋࡓࠋࡇࡢ⤖ᯝࡣࠊ༸ẕ⣽⬊࠿ࡽศἪࡉࢀࡿఱࡽ
࠿ࡢᅉᏊ࡟ࡼࡾ MT2-MMP ࡢⓎ⌧ࡀㄪ⠇㸦ᢚไ㸧ࡉࢀ࡚࠸ࡿࡇ࡜ࢆ♧၀ࡋ࡚࠸ࡿࠋ့ங㢮ࢆ⏝࠸ࡓ◊✲࠿ࡽࠊGDF9ࠊBMP15 ࡞࡝ࡀ MT2-MMP ࡢⓎ⌧ไᚚ࡟㛵ࢃࡗ࡚࠸ࡿྍ⬟ᛶࡀ࠶ࡿࡇ࡜ࡀ⪃࠼ࡽࢀࡓࡢ࡛ࠊࡇࢀࡽࡢⓎ⌧ࢆㄪ࡭ࡓ࡜ࡇࢁࠊ࣓ࢲ
࢝᤼༸๓༸ẕ⣽⬊࡟Ⓨ⌧ࡋ࡚࠸ࡿࡇ࡜ࡀ☜ㄆ࡛ࡁࡓࠋ௒ᚋࡣࠊࡇࢀࡽࡢᅉᏊࡀ MT2-MMP ࡢⓎ⌧ไᚚ࡟㛵ࢃࡗ࡚࠸ࡿ࠿࡝࠺࠿ㄪ࡭ࡿணᐃ࡛࠶ࡿࠋ
㸦3㸧ℐ⬊㑅ᢥᶵᵓࡢゎ့᫂ங㢮ࡢ༸ᕢ࡟ࡣࠊཎጞℐ⬊࡜ࡼࡤࢀࡿℐ⬊ࡀከᩘࢫࢺࢵࢡࡉࢀ࡚࠸ࡿࠋ࠶ࡿ่⃭࡟ࡼࡾࡇࡢཎ
ጞℐ⬊ࡢ୍㒊㸦ᩘ༑ಶ㹼ⓒಶ⛬ᗘ㸧ࡀᡂ㛗ࢆ㛤ጞࡋࠊ᭱⤊ⓗ࡟㸯㹼10 ಶ⛬ᗘࡢ༸Ꮚࡀ᤼༸ࡉࢀࡿࠋ
ᡂ㛗ࢆ㛤ጞࡋࡓℐ⬊ࡢ኱㒊ศࡣࠊ㏵୰࡛ᡂ㛗ࢆ೵Ṇࡋࠊ࢔࣏ࢺ࣮ࢩࢫ࡟ࡼࡾṚ⁛ࡍࡿࠋࡇࡢ㐣⛬ࡀ
ℐ⬊㑅ᢥ࡛࠶ࡿࡀࠊᮏ◊✲࡛ࡣ࡝ࡢࡼ࠺࡟ࡋ࡚⏕ࡁṧࡿℐ⬊ࡀ㑅ࡧฟࡉࢀࡿࡢ࠿ࠊࡑࡢศᏊᶵᵓࢆ
᫂ࡽ࠿࡟ࡍࡿࡇ࡜ࢆ┠ⓗ࡜ࡋ࡚࠸ࡿࠋ
㏻ᖖࠊPMSG/hCG ➼࡛㐣᤼༸ࢆㄏᑟࡍࡿ࡜
10-20 ಶ⛬ᗘ㸦࣐࢘ࢫࡢሙྜ㸧ࡢ༸ࡀ᤼༸ࡉࢀࡿ
ࡀࠊ࠶ࡿ⸆๣ࢆ⏝࠸ࡿ࡜᤼༸ᩘࡀ 100 ಶ๓ᚋ࡜࡞
ࡿࡇ࡜࠿ࡽࠊࡇࡢ⸆๣ࡣℐ⬊㑅ᢥࢆᢚไࡋ࡚࠸ࡿ
ࡢ࡛ࡣ࡞࠸࠿࡜⪃࠼ࡓࠋࡑࡇ࡛ࠊࡲࡎ PMSG ࡜ࡑࡢ⸆๣࡛ฎ⌮ࡋࡓ༸ᕢ࡛ࡢ㑇ఏᏊⓎ⌧ࣃࢱ࣮ࣥࢆ
RNA-seq ࡛ゎᯒࡋࠊⓎ⌧㔞࡟ᕪࡢ࠶ࡿ㑇ఏᏊࢆ㑅ูࡋࡓࠋࡑࡢ⤖ᯝࠊⓎ⌧㔞࡟ᕪࡢ࠶ࡿ㑇ఏᏊࡀᩘከࡃ㑅ูࡉࢀࡓࡀࠊ⌧ᅾ࡝ࡢ㑇ఏᏊࡀℐ⬊㑅ᢥ࡟㛵୚ࡋ࡚࠸ࡿ࠿ࢆㄪ࡭࡚࠸ࡿẁ㝵࡛࠶ࡿࠋㄽᩥTakahashi T., Hagiwara A., Ogiwara K. (2018) Follicle
rupture during ovulation with emphasis on recent
progress in fish models. (review) Reproduction 157,
issue 1 DOI: 10.1530/REP-18-0251.
Takahashi T., Hagiwara A., Ogiwara K. (2018) The− 22 −

roles of prostaglandins in vertebrate ovulation. (Minireview) Grobal J Reprod. Medicine 4, DOI:
10.19080/GJORM.2018.04.555639.
Takahashi, T. Hagiwara, A. Ogiwara, K. (2018)
Prostaglandins in teleost ovulation: A review of theroles with a view to comparison with prostaglandins inmammalian ovulation. Mol. Cell. Endocrinol. 461:236-247.
Ꮫ఍Ⓨ⾲ⴗཎⲁࠊⲶཎඞ┈ࠊᒣୗṇවࠊ⾜ಇ⏤ோࠊ㧗ᶫᏕ⾜㸸࣓ࢲ࢝᤼༸๓ℐ⬊࡟࠾ࡅࡿ GAP ⤖ྜࡢ᤼༸࡟ᑐ
ࡍࡿᙺ๭➨ 89 ᅇ᪥ᮏື≀Ꮫ఍ᮐᖠ኱఍ࠊᖹᡂ 30 ᖺ 9 ᭶ 13-15
᪥ࠊᮐᖠࢥࣥ࣋ࣥࢩࣙࣥࢭࣥࢱ࣮㸦ᮐᖠᕷ㸧Ⲷཎඞ┈ࠊΏ㑔ᘺஓࠊ୰₲┤ᕫࠊ➉ᑿ㏱㸸࣐࢘ࢫ༸ᕢ࡟࠾ࡅࡿℐ⬊㑅ᢥ࡟㛵୚ࡍࡿᅉᏊࡢ⥙⨶ⓗゎᯒ➨ 89 ᅇ᪥ᮏື≀Ꮫ఍ᮐᖠ኱఍ࠊᖹᡂ 30 ᖺ 9 ᭶ 13-15
᪥ࠊᮐᖠࢥࣥ࣋ࣥࢩࣙࣥࢭࣥࢱ࣮㸦ᮐᖠᕷ㸧㒊㛛 ከᵝᛶ⏕≀Ꮫศศ㔝⏕≀⛉Ꮫ㒊ᑠட ୍ᘯ
㸦ᾏỈ฼⏝㸧Ⓨ⾲ㄽᩥ
Masakazu Hoshino, Shozo Ishikawa and Kazuhiro
Kogame. 2018. Concordance between DNA-based
species boundaries and reproductive isolating barriersin the Scytosiphon lomentaria species complex(Ectocarpales, Phaeophyceae). Phycologia: 57: 232–242.
Suttikarn Sutti, Masaya Tani, Yukimasa Yamagishi,
Tsuyoshi Abe, Kathy A. Miller and Kazuhiro Kogame.
2018. Neochondria gen. nov. (Rhodomelaceae,
Rhodophyta), a segregate of Chondria, including N.
ammophila sp. nov. and N. nidifica comb. nov.
Phycologia 57: 262–272.
Masakazu Hoshino, Tatsufumi Okino and Kazuhiro
Kogame. 2019. Parthenogenetic female populations in thebrown alga Scytosiphon lomentaria (Scytosiphonaceae,
Ectocarpales): decay of a sexual trait and acquisition ofasexual traits. Journal of Phycology 55: 204-213.
Agnieszka P. Lipinska, Martha L. Serrano-Serrano,
Alexandre Cormier, Akira F. Peters, Kazuhiro Kogame,
J. Mark Cock and Susana M. Coelho. 2019. Rapid
turnover of life-cycle-related genes in the brown algae.
Genome Biology 20:35.

࣮ࢥ࣮ࢹ࢕ࣥࢢ㸪᪥ᮏ⸴㢮Ꮫ఍➨ 43 ᅇ኱఍㸪2019ᖺ 3 ᭶ 16㸪17 ᪥㸪ி㒔኱Ꮫ㸦ி㒔ᕷ㸧ᫍ㔝㞞࿴㸪⚄㇂඘ఙ㸪ᑠட୍ᘯ㸸〓⸴࢝ࣖࣔࣀࣜ
ࡢ༢Ⅽ⏕Ṫ⣔⤫ࡢ㉳※࡟ࡘ࠸࡚㸪᪥ᮏ⸴㢮Ꮫ఍➨
43 ᅇ኱఍㸪2019 ᖺ 3 ᭶ 16㸪17 ᪥㸪ி㒔኱Ꮫ㸦ி㒔ᕷ㸧ᫍ㔝㞞࿴㸪⏣୰ཌᏊ㸪ୖ஭㐍ஓ㸪⚄㇂඘ఙ㸪ᖹᒸ㞞つ㸪ᑠட୍ᘯ㸸㞃ⶸ✀ࢆྡ௜ࡅࡿ㸸᪥ᮏ⏘〓⸴࢝ࣖࣔࣀࣜࢆ6✀࡟ศࡅࡿ㸪᪥ᮏ⸴㢮Ꮫ఍➨43ᅇ኱
఍㸪2019ᖺ3᭶16㸪17᪥㸪ி㒔኱Ꮫ㸦ி㒔ᕷ㸧ᫍ㔝㞞࿴㸪⩚⏕⏣ᓅ᫛㸪ᑠட୍ᘯ㸸᪥ᮏ⏘〓⸴࢘ࢫ࢝ࣖࣔ㸦࢝ࣖࣔࣀࣜ⛉㸧ࡢศ㢮ࡢ෌᳨ウ㸪᪥ᮏ⸴㢮Ꮫ఍➨ 43 ᅇ኱఍㸪2019 ᖺ 3 ᭶ 16㸪17 ᪥㸪ி㒔኱Ꮫ㸦ி㒔ᕷ㸧⏕≀⛉Ꮫ㒊㒊㛛 ከᵝᛶ⏕≀Ꮫศศ㔝᯳ཎ ᏹ
㸦ᾏỈ฼⏝㸧
ụỌ₶ᖹ, Ἴࠎ఑㒊ኟ⨾, ᖾሯஂ඾, ྜྷ⏣ Ꮫ, ᯳ཎ ᏹ㸸୕ᓮ⏘࢖ࢯࣄ࣒ࣔࢩಶయ⩌࡟ࡣ࢜ࢫࡋ࠿࠸࡞࠸㸽 㸦⣣ᙧື≀㛛␗⣣⹸㢮㸧. ᪥ᮏື≀ศ㢮Ꮫ఍➨ 54 ᅇ኱఍, 2018 ᖺ 6 ᭶ 10 ᪥,㮵ඣᓥ኱Ꮫ㸦㮵ඣᓥᕷ㸧.
Kajihara, H. and Hookabe, N.: Why do certain speciesof heteronemerteans get so long? ➨ 9 ᅇ⣣ᙧື≀Ꮫᅜ㝿఍㆟, 2018 ᖺ 8 ᭶ 14 ᪥, ࢔ࣝࣇ࣮ࣞ
ࢺ࣭ࣦ࢙࣮ࢤࢼ࣮ᴟᆅᾏὒ◊✲ᡤࣦ࢓ࢵࢸࣥ
ᾏᐇ㦂ᡤ㸦ࢻ࢖ࢶ࣭ࢬࣗࣝࢺ㸧.
㛗㇂ᕝᑦᘯ, ᯳ཎ ᏹ㸸࢝ࢱ࣐࣒ࣜࣛ࣎ࣖ㸦࣍ࣖ⥘࣐࣎ࣖ┠ࢩࣟ࣎ࣖ⛉㸧ࡢ⣔⤫ศ㢮Ꮫⓗ◊✲.
᪥ᮏື≀ศ㢮Ꮫ఍➨ 54 ᅇ኱఍, 2018 ᖺ 6 ᭶ 9᪥, 㮵ඣᓥ኱Ꮫ㸦㮵ඣᓥᕷ㸧.
Ἴࠎ఑㒊ኟ⨾, Megan L. Schwartz, ᯳ཎ ᏹ, Jon L.
Norenburg㸸ࢡࣟࣄ࣒ࣔࢩ㸦⣣ᙧື≀㛛␗⣣⹸㢮㸧ࡢศ㢮Ꮫⓗ෌᳨ウ. ᪥ᮏື≀ศ㢮Ꮫ఍➨54 ᅇ኱఍, 2018 ᖺ 6 ᭶ 9 ᪥, 㮵ඣᓥ኱Ꮫ㸦㮵ඣᓥᕷ㸧.
Hookabe, N., Schwartz, M. L., Kajihara, H., andNorenburg, J. L.: Molecular systematics of theheteronemertean genus Dushia (Nemertea,
Pilidiophora). ➨ 9 ᅇ⣣ᙧື≀Ꮫᅜ㝿఍㆟, 2018
ᖺ 8 ᭶ 15 ᪥, ࢔ࣝࣇ࣮ࣞࢺ࣭ࣦ࢙࣮ࢤࢼ࣮ᴟ
ᆅᾏὒ◊✲ᡤࣦ࢓ࢵࢸࣥᾏᐇ㦂ᡤ㸦ࢻ࢖ࢶ࣭ࢬࣗࣝࢺ㸧.

Ꮫ఍Ⓨ⾲㛵ᮏ⍠⳯㸪ᑠட୍ᘯ㸸〓⸴㢮ࡢᚤᑠ⸴యࡢ COI ࣂ− 23 −

⏕≀⛉⛉Ꮫ㒊㒊㛛 ከᵝᛶ⏕≀Ꮫศศ㔝ຍ⸨ ᚭ
ᙜ◊✲ᐊ࡛ࡣ㸪㔝እ࠿ࡽ᥇㞟ࡋࡓ✀ࠎࡢࢩࣙ࢘ࢪ
ࣙ࢘ࣂ࢚ࢆ⣔⤫Ꮫ◊✲ࡸ㞟ᅋ㑇ఏᏛ◊✲࡟⏝࠸ࡿ࡜࡜ࡶ࡟㸪㣫⫱ྍ⬟࡞ࢩࣙ࢘ࢪࣙ࢘ࣂ࢚࡟ࡘ࠸࡚
ࡣ㸪⥅௦㣫⫱࡟ࡼࡗ࡚⣔⤫ࢆ⥔ᣢࡋ࡚࠸ࡿ㸬ࡲࡓ㸪௒ᖺᗘࡣ㸪ࢩࣙ࢘ࢪࣙ࢘ࣂ࢚௨እ࡟ࡶ㸪㔝እ࠿ࡽ࣐ࢨࢺ࣒࢘ࢩࢆ᥇㞟ࡋ࡚㣫⫱ࡋ㸪㞝ࡢ஧ᆺ࡜㓄അ⾜ືࡢ㛵ಀࢆㄪᰝࡋࡓ㸬1) ྠᡤⓗ࡟ศᕸࡍࡿ㏆⦕ࢩࣙ࢘ࢪࣙ࢘ࣂ࢚ 2 ✀ࡢ㈨※฼⏝ࡢ㐪࠸
㸸Lordiphosa collinella ࡣᮾ࢔ࢪ࢔࡟ᗈࡃศᕸࡍ
ࡿⲡᮏ㣗ᛶࡢࢩࣙ࢘ࢪࣙ࢘ࣂ࢚࡛㸪ࢽࣜࣥࢯ࢘
࡞࡝ࡢ᫓᳜≀ࢆ୺࡞⦾Ṫ㈨※࡜ࡋ࡚฼⏝ࡍࡿ㸬୍᪉㸪໭ᾏ㐨࡟ࡣ㸪L. collinella ࡜ᙧែⓗ࡟㓞ఝࡍࡿ L. sp. aff. collinella ࡀྠᡤⓗ࡟ศᕸࡍࡿࡀ㸪ࡇࡢ✀ࡣᖖ⥳పᮌ࡛࠶ࡿࣇࢵ࢟ࢯ࢘ࡢ⩌ⴠ࠿ࡽ㧗㢖ᗘ࡛᥇㞟ࡉࢀࡿ㸬୧✀ࡢ㈨※฼⏝ࡢᵝᘧ࡟㐪࠸ࡀ࠶ࡿ࠿ࢆ᳨ドࡍࡿࡓࡵ㸪୧✀ࡀྠᡤⓗ࡟⏕ᜥࡍࡿᆅⅬ࡟࠾࠸࡚⦾Ṫ⏕ែࢆㄪᰝࡋࡓ㸬ࡑࡢ⤖ᯝ㸪L. collinella ࡣࢽࣜࣥࢯ࢘ࢆࡣࡌࡵ࡜ࡋ
࡚ẚ㍑ⓗࡸࢃࡽ࠿࠸」ᩘ✀ࡢ᳜≀࡛⦾Ṫࡀ☜ㄆࡉࢀࡓࡀ㸪L. sp. aff. collinella ࡣᑓࡽࣇࢵ࢟ࢯ࢘
ࡢ⭉ࡗࡓⴥࡸⱼࢆ⦾Ṫ࡟฼⏝ࡋ࡚࠸ࡓ㸬ࡲࡓ㸪ᐇ㦂ᐊ࡛ࢽࣜࣥࢯ࢘࡜ࣇࢵ࢟ࢯ࢘ࢆ୚࠼࡚㞤ᡂ⹸ࡢ⏘༸㑅ዲᛶࢆㄪ࡭ࡓ⤖ᯝ㸪 L. collinella ࡣࢽࣜࣥࢯ࢘࡜ࣇࢵ࢟ࢯ࢘ࡢ୧᪉ࢆ㑅ࡧࡘࡘࢽࣜࣥࢯ ࢘ ࢆ ዲ ࡴ ഴ ྥ ࢆ ♧ ࡋ ࡓ ୍ ᪉ 㸪 L. sp. aff.
collinella ࡣࣇࢵ࢟ࢯ࢘࡬ࡢ㢧ⴭ࡞㑅ዲᛶࢆ♧ࡋࡓ㸬ࡇࢀࡽࡢ⤖ᯝࡣ㸪๓⪅ࡣ」ᩘ✀ࡢ᳜≀ࢆ⦾Ṫ࡟฼⏝ࡍࡿࡀ㸪ᚋ⪅ࡣࣇࢵ࢟ࢯ࢘ࡢ฼⏝࡟≉໬ࡋ࡚࠸ࡿࡇ࡜ࢆ♧၀ࡍࡿ㸬2) ࢟ࣀࢥ㣗ᛶࢩࣙ࢘ࢪࣙ࢘ࣂ࢚⩌ࡢ⣔⤫࡜㐍໬ࢩࣙ࢘ࢪࣙ࢘ࣂ࢚⛉ࡣ⣙ 4000 ✀ࡀグ㍕ࡉࢀ࡚࠾ࡾ㸪ࡑࡢ㣗ᛶ࡜ࡋ࡚ᶞᾮ㣗㸪ᯝ≀㣗㸪ⲡᮏ㣗㸪࢟ࣀࢥ㣗࡞࡝ࡀ▱ࡽࢀ࡚࠸ࡿ㸬ࡑࡢ࠺ࡕ㸪࢟ࣀࢥ㣗ࡣࢩࣙ࢘ࢪࣙ࢘ࣂ࢚࡟࠾࠸࡚ࡣẚ㍑ⓗ⌋ࡋ࠸㣗ᛶ࡛࠶ࡾ㸪୍㒊ࡢࢩࣙ࢘ࢪࣙ࢘ࣂ࢚⩌㸦 Hirtodrosophila ᒓ 㸪 Mycodrosophila ᒓ 㸪
Zygothrica ᒓ㸪࠾ࡼࡧ Drosophila ளᒓࡢ quinaria✀⩌㸧࡟೫ࡗ࡚ࡳࡽࢀࡿᙧ㉁࡛࠶ࡿ㸬ࡋ࠿ࡋ࡞ࡀࡽ㸪ࡇࢀࡽ࢟ࣀࢥ㣗ᛶࢩࣙ࢘ࢪࣙ࢘ࣂ࢚ࡢ⣔⤫㛵ಀ࡟ࡘ࠸࡚ࡣ◊✲ࡀ㐍ࢇ࡛࠾ࡽࡎ㸪ࡑࡢᙧ㉁ࡀࢩࣙ࢘ࢪࣙ࢘ࣂ࢚ࡢ㐍໬ࡢ㐣⛬࡛࡝ࡢࡼ࠺࡟⋓ᚓࡉࢀࡓࡢ࠿ࡣ୙᫂࡞ࡲࡲ࡛࠶ࡿ㸬ࡑࡇ࡛ୖグศ㢮⩌ࡢࢩࣙ࢘ࢪࣙ࢘ࣂ࢚ࢆᑐ㇟࡟㸪40௨ୖࡢ㑇ఏᏊࢆ⏝࠸࡚㧗ゎീᗘࡢศᏊ⣔⤫ᶞࢆᵓ⠏ࡍ࡭ࡃ㸪PCR ቑᖜ࡜ሷᇶ㓄ิࡢỴᐃࢆ㐍ࡵ࡚࠸ࡿ㸬3) ࣐ࢨࢺ࣒࢘ࢩ࡟࠾ࡅࡿ㞝ࡢ஧ᆺ࡜㓄അ⾜ືࡢ㛵ಀ࣐ࢨࢺ࣒࢘ࢩ㸦Phalangium opilio㸧࡛ࡣ㸪㞝ࡢゅࡢᙧែ࡟஧ᆺࡀᏑᅾࡍࡿࡇ࡜ࡀ♧၀ࡉࢀ࡚࠸ࡿ㸬ࡑࡇ࡛㸪ᮐᖠᕷཌู༊࡟⏕ᜥࡍࡿ࣐ࢨࢺ࣒࢘ࢩࡢ㞝ࡢᙧែࢆㄪᰝࡋࡓ࡜ࡇࢁ㸪㗑ゅࡢඛ➃ࡢゅࡢ㛗ࡉ࡜ゐ⫥⭣⠇㛗࡟஧ᆺࡀᏑᅾࡍࡿࡇ࡜ࡀ☜ㄆࡉࢀࡓ㸬ࡲࡓ㸪࣐ࢨࢺ࣒࢘ࢩࡢ㓄അ⾜ືࢆࣅࢹ࢜᧜ᙳࡋ࡚
ほᐹࡋࡓ⤖ᯝ㸪㞝ಶయࡀ㞤ಶయ࡟ࡲ࡜ࢃࡾࡘࡃ⾜ືࡀほᐹࡉࢀࡓࡇ࡜࠿ࡽ㸪ࡑࡢᅇᩘࢆ஧ᆺ㞝㛫࡛ẚ㍑ࡋࡓ࡜ࡇࢁ㸪ゅࡀ኱ࡁ࠸㞝ࡢ᪉ࡀ㸪ࡼࡾከࡃ㞤࡟ࡲ࡜ࢃࡾࡘࡃഴྥࡀほᐹࡉࢀࡓ㸬ࡇࡢ⤖ᯝࡣ㸪ࡇࢀࡽ஧ᆺ㞝ࡢ㛫࡛ࡣ㓄അᵝᘧ࡟ࡶ㐪࠸ࡀ࠶ࡿྍ
⬟ᛶࢆ♧၀ࡍࡿ㸬ㄅୖⓎ⾲
1. ຍ⸨ᚭ㸪୕ࢶᶫᆂ㸪Ἠ㇂ὒஅ㸪ᡞ⏣ṇ᠇㸦2019㸧ྠᡤⓗ࡟⏕ᜥࡍࡿ㏆⦕ࢩࣙ࢘ࢪࣙ࢘ࣂ࢚ 2 ✀ࡢ㈨※฼⏝࡜㑇ఏศ໬㸬᪻⹸࡜⮬↛ 54:30-33㸬⏕≀⛉Ꮫ㒊㒊㛛 ከᵝᛶ⏕≀Ꮫศศ㔝ゅ஭ ᩗ▱฼⏝≧ἣ࣭◊✲ᡂᯝᮏᖺᗘࡣ㸪◊✲ㄢ㢟ྡࠕᑠᆺ⏥Ẇ㢮ࡢᛶ࣭⏕Ṫᵝᘧ࡟㛵ࡍࡿ◊✲ࠖ࡜ࡋ࡚㸪ࢤࣀ࣒ࢲ࢖ࢼ࣑ࢡࢫ◊✲ࢭࣥࢱ࣮ᐇ㦂⏕≀ඹྠ฼⏝㒊㛛㸦ᐇ㦂ᐊ㸦㸰㸧㸧ࡢ฼⏝⏦ㄳࢆ⾜࠸㸪ࢱࢼ࢖ࢫ┠࡜࠸࠺ᑠᆺ⏥Ẇ㢮ࡢ୍⩌࡟㛵ࡍࡿ◊✲ࢆ⾜ࡗࡓ㸬୍࢝᭶㛫㸦2018 ᖺ
9 ᭶㸧ࡢ฼⏝ᮇ㛫୰࡟㸪ᚸ㊰‴࡟࡚᥇㞟ࢆ⾜ࡗࡓࢼ࣑ࢱࢼ࢖ࢫᒓࡢ 1 ✀㸦Zeuxo sp.㸧ࡢ㉮ගᛶࡢ᭷↓ࢆ☜ㄆࡍࡿᐇ㦂ࢆᐇ᪋㸪ᮏ✀ࡀṇࡢ㉮ගᛶࢆ♧ࡍࡇ࡜ࢆ᫂ࡽ࠿࡟ࡋࡓ㸬࡞࠾ᮏ◊✲ࡣ㸪ࢱࢼ࢖ࢫ┠⏥Ẇ㢮࡟࠾࠸࡚㸪㉮ගᛶࡢᏑᅾࢆᐇ㦂ⓗ࡟♧ࡋࡓึࡢ◊✲࡛࠶ࡿ㸬ᮏ◊✲ࡣ⚾ࡢᣦᑟᏛ⏕࡛࠶ࡿᒸᮏᬸ㌍࡟ࡼࡿ⏕≀⛉Ꮫ◊✲ᐇ⩦ࡢ୍⎔࡜ࡋ࡚ᐇ᪋ࡋ㸪◊✲ᐇ⩦Ⓨ⾲఍࡟࠾࠸࡚࣏ࢫࢱ࣮Ⓨ⾲ࢆ⾜ࡗࡓ㸬ᴗ⦼ࣜࢫࢺ㸦ㄽᩥⓎ⾲࣭Ꮫ⾡Ⓨ⾲➼㸧㸦༞ᴗ࣭ಟኈ࣭༤ኈ◊✲Ⓨ⾲㸧
Nobuya Okamoto: Taxonomic and behavioral studies
of Zeuxo sp. (Crustacea: Tanaidacea) from Oshoro Bay,
Japan㸦ᚸ㊰‴⏘ࢼ࣑ࢱࢼ࢖ࢫᒓࡢ 1 ✀ࡢศ㢮Ꮫⓗ࣭⾜ືᏛⓗ◊✲㸦⏥Ẇள㛛ࢱࢼ࢖ࢫ┠㸧㸧. ᖹᡂ
30 ᖺᗘ⏕≀⛉Ꮫ⛉㸦⏕≀㸧◊✲ᐇ⩦Ⓨ⾲㸪࣏ࢫࢱ࣮␒ྕ 4㸪2019 ᖺ 2 ᭶ 1 ᪥㸪໭ᾏ㐨኱Ꮫ⌮Ꮫ㒊㸦ᮐᖠᕷ㸧㸬

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኱Ꮫ㝔㎰Ꮫ◊✲㝔ᇶ┙◊✲㒊㛛 ⏕≀㈨※⛉Ꮫศ㔝⛅ඖ ಙ୍
▷⩟ᛶ࡛⛣ື⬟ຊ࡟ஈࡋ࠸㸪ࢧࢵ࣏ࣟࣇ࢟ࣂࢵࢱ
Podisma sapporensis ཬ ࡧ 㸪 ࣑ ࢝ ࢻ ࣇ ࢟ ࣂ ࢵ ࢱParapodisma mikado ࢆᮦᩱ࡟⏝࠸㸪㞟ᅋ㛫ࡢ㓄അ⾜ືࡢศ໬ࡢ㐍໬ࣉࣟࢭࢫࡸ㸪⏕Ṫ㝸㞳ᶵᵓࡢゎ᫂࡟ྲྀࡾ⤌ࢇ࡛࠸ࡿ㸬≉࡟ࢧࢵ࣏ࣟࣇ࢟ࣂࢵࢱࡣ㸪ࡇࢀࡽࡢ◊✲ࡢᮦᩱ࡜ࡋ࡚㸪௨ୗࡢ 2 Ⅼ࡟࠾࠸࡚㠀ᖖ࡟≉␗ⓗ࡛࠶ࡿ㸬1㸧ᛶỴᐃࢩࢫࢸ࣒ࡀ␗࡞ࡿ 2 㞟ᅋࡀ㸪␗ᡤⓗ࡟Ᏻᐃⓗ࡟Ꮡᅾࡋ࡚࠸ࡿ㸬໭ᾏ㐨す㒊࡟ࡣ㸪࢜ࢫ XO㸪࣓ࢫ XX ࡛ᛶỴᐃࡍࡿ XO ࣮ࣞࢫ㞟ᅋࡀ㸪໭ᾏ㐨ᮾ㒊࡟ࡣ࢜ࢫ neo-XY㸪࣓ࢫ neo-XX࡛ᛶỴᐃࡍࡿ neo-XY ࣮ࣞࢫ㞟ᅋࡀศᕸࡋ㸪୧࣮ࣞࢫ㛫࡟ࡣ㸪⏕Ṫⓗ㝸㞳ࡀᏑᅾࡍࡿ࡜࠸࠺ሗ࿌ࡀ࠶ࡿ㸬
2㸧ࡇࢀࡲ࡛ࡢ◊✲࡛㸪ࢧࢵ࣏ࣟࣇ࢟ࣂࢵࢱࡣ஺ᑿ
⾜ື࡟࠾࠸࡚㐨ෆࡢᆅᇦࡈ࡜࡟┦㐪ࡀᏑᅾࡍࡿࡇ
࡜ࡀ᫂ࡽ࠿࡟࡞ࡗ࡚࠸ࡿ㸬ᮏ✀ࡢ࣓ࢫࡣ㸪࢜ࢫ࡟ᑐࡋ࡚஺ᑿᣄྰ⾜ືࢆ♧ࡍࡀ㸪஺ᑿᣄྰࡢᙉࡉࡀᆅᇦ
㞟ᅋ࡟ࡼࡗ࡚␗࡞ࡗ࡚࠾ࡾ㸪㞟ᅋ㛫࡛஺ᑿࡉࡏࡿ࡜㸪஺ᑿ㢖ᗘࡣࣛࣥࢲ࣒࡛ࡣ࡞ࡃ㸪͆㠀ᑐ⛠࡞஺ᑿ͇ࡀ
ぢࡽࢀࡿ㸬ࡍ࡞ࢃࡕ㸪஺ᑿᣄྰຊࡢᙉ࠸㞟ᅋࡢ࣓ࢫࡣ㸪௚㞟ᅋࡢ࢜ࢫࢆ࡞࠿࡞࠿ཷࡅධࢀ࡞࠸ࡀ㸪஺ᑿᣄྰຊࡢప࠸㞟ᅋࡢ࣓ࢫࡣ௚㞟ᅋࡢ࢜ࢫࢆཷࡅධ
ࢀ㸪ᐜ᫆࡟஺ᑿࡀほᐹ࡛ࡁࡿ㸬ࡲࡓ㸪࣓ࢫࡢ஺ᑿᣄ
ྰຊࡀᙉ࠸㞟ᅋࡢ࢜ࢫࡣ஺ᑿάຊࡀᙉࡃ㸪࣓ࢫࡢ஺ᑿᣄྰຊࡢప࠸㞟ᅋࡢ࢜ࢫࡣ஺ᑿάຊࡀᙅ࠸㸬ࡑࡢ
Ⅽ㸪஺ᑿάᛶࡀᙉ࠸㞟ᅋࡢ࢜ࢫࡣ஺ᑿᣄྰຊࡀᙅ࠸
㞟ᅋࡢ࣓ࢫ࡜㞟ᅋ㛫஺㓄ࢆ⾜࠺࡜㸪஺ᑿᅇᩘࡀከࡃ
࡞ࡿ㸬ᮏᖺᗘࡣ㸪᫖ᖺࡢ᪂ࡋ࠸㞟ᅋࢆ⏝࠸ࡓ㞟ᅋ㛫஺㓄ࡢணഛᐇ㦂࡟࠾࠸࡚㸪஺ᑿᡂຌ⋡ࡀ㧗࠸㸪༶ࡕ஺ᑿάᛶࡀ㧗࠸࡟ࡶ㛵ࢃࡽࡎ㸪ㄪᰝ᫬㛫࠶ࡓࡾࡢ஺ᑿᅇ
ᩘ⮬యࡣከࡃࡣ࡞࠸࡜࠸࠺㸪ࡇࢀࡲ࡛ᐇ㦂࡟⏝࠸ࡽࢀ࡚ࡁࡓ㞟ᅋ࡜ࡣࡸࡸ஺ᑿഴྥࡀ␗࡞ࡿ㞟ᅋࢆ⏝࠸࡚㸪஺ᑿᐇ㦂㸪⾜ືᐇ㦂ࢆ⾜ࡗࡓ㸬ࡇࡢ㞟ᅋࡢ≉ᚩ࡜ࡋ࡚㸪ᮏ✀ࡢ࡯࡜ࢇ࡝ࡢᆅᇦ㞟ᅋࡀ⥳Ⰽࢆ࣮࣋ࢫ࡜ࡋࡓయⰍࢆࡋ࡚࠸ࡿࡢ࡜␗࡞ࡾ㸪㯮Ⰽࡀ㠀ᖖ࡟ከ࠸≉␗࡞Ⰽᙬࢆࡋ࡚࠸ࡿ㸬ࡇࡢⰍᙬኚ␗ࡀ⥔ᣢࡉࢀ࡚࠸ࡿ࣓࢝ࢽࢬ࣒ࡀ㸪஺ᑿ⾜ືࡢ㐪࠸࠿ࡽ᮶࡚࠸
ࡿྍ⬟ᛶࢆ᳨ドࡋࡓ㸬ࡲࡎ᳨ドᐇ㦂࡟⏝࠸ࡿࡢ࡟㐺ࡋࡓ㞟ᅋࢆ㑅ᐃࡍࡿⅭ㸪⥳Ⰽᆺࡢ㞟ᅋ࡜㯮Ⰽᆺࡢ㞟ᅋࡀ᭱ࡶ㏆࠸ሙᡤ࡟ศᕸࡍࡿᆅᇦࢆ᥈⣴ࡋࡓ㸬ㄪᰝࡢ⤖ᯝ㸪㯮Ⰽᆺࡣ㸪ࡇࢀࡲ࡛ࡈࡃ⊃࠸⠊ᅖ࡛⏕ᜥࡋ࡚࠸ࡿ࡜⪃࠼ࡽࢀ࡚࠸ࡓᆅᇦ࠿ࡽ㸪ᑡ࡞ࡃ࡜ࡶ໭ࡣ
45 ࢟ࣟ㸪ᮾࡣ 27 ࢟ࣟ࡟ཬࡪ࠿࡞ࡾࡢᗈ⠊ᅖ࡛⏕ᜥࡋ࡚࠸ࡿ஦ࡀ᫂ࡽ࠿࡟࡞ࡗࡓ㸬ࡋ࠿ࡋ࡞ࡀࡽ㸪⥳Ⰽᆺ࡜㯮Ⰽᆺࡢ୰㛫ᆺࢆ♧ࡍࡼ࠺࡞ಶయࡸぢࡽࢀࡎ㸪ࡲࡓ㯮Ⰽᆺࡢศᕸᇦ࡟ࡣ⥳Ⰽᆺࡢ㞟ᅋࡣぢࡽࢀ࡞࠿ࡗࡓ㸬஺ᑿᐇ㦂㸪⾜ືᐇ㦂ࢆ⾜ࡗࡓ⤖ᯝ㸪୧Ⰽᙬᆺ㛫࡟஺㓄๓㝸㞳ࡣᏑᅾࡋ࡞࠿ࡗࡓ㸬௒ᚋ㸪⥳Ⰽᆺ࡜㯮Ⰽᆺ㛫ࡢ஺㓄࡛ᚓࡽࢀࡓ㉺෤༸࠿ࡽ㸪ཷ⢭⋡࡜
F1 ࡢⰍᙬࢆㄪ࡭㸪㯮Ⰽᆺ㞟ᅋࡀࡇࢀࡲ࡛ࡢ▱ぢࡼ
ࡾࡶᛴ㏿࡟ศᕸᇦࢆᣑ኱ࡋࡘࡘ࠶ࡿྍ⬟ᛶࡢ᳨ドࡶྵࡵ㸪㯮Ⰽᆺࡀ⥔ᣢࡉࢀ࡚࠸ࡿ࣓࢝ࢽࢬ࣒ࡢゎ᫂
ࢆ┠ᣦࡍ㸬


ศ㔝
ᇶ┙◊✲㒊㛛 ␆⏘⛉Ꮫศᕝཎ Ꮫ
့ங㢮࡟࠾࠸࡚ࠊ༸ẕ⣽⬊ࡣ⢭Ꮚ࡜ẚ㍑ࡋ࡚᥇
ྲྀࡀᅔ㞴࡛࠶ࡾࠊཷ⢭ᚋࡢึᮇ⬇Ⓨ⏕ࡢᡂྰࡣ༸
ẕ⣽⬊ࡢⓎ⏕ᨭᣢ⬟࡟኱ࡁࡃ౫Ꮡࡋ࡚࠸ࡿࠋ㑇ఏⓗ࡟ከᵝ࡛࠶ࡿᐙ␆࡞࡝ࡢື≀✀࡛ࡣಶయᕪࡶ኱
ࡁࡃ඲࡚ࡢ༸ẕ⣽⬊ࡀ༑ศ࠿ࡘᆒ୍࡞Ⓨ⏕ᨭᣢ⬟ࢆ᭷ࡋ࡚࠸ࡿࢃࡅ࡛ࡣ࡞࠸ࡓࡵࠊⓎ⏕ᨭᣢ⬟ࡢప࠸༸ẕ⣽⬊ࡢ฼⏝ᛶࡢྥୖࡣ㎰Ꮫࠊື≀Ⓨ⏕ᕤᏛ࡞࡝ࡢศ㔝࠾࠸࡚኱ࡁ࡞ၥ㢟࡜࡞ࡿࠋ୍᪉ࠊ༸ẕ⣽⬊ࡢⓎ⏕ᨭᣢ⬟ࡢపୗ࡟ࡘ࠸࡚ࡣࠊ᤼༸ᚋ࡟య
እ࡛㛗᫬㛫㟼⨨ࡋࠊཷ⢭ࡢ㐺ᮇࢆேⅭⓗ࡟㐜ᘏࡉࡏࡓሙྜ࡟ࡶ㢧ⴭ࡛࠶ࡿࠋᡃࠎࡣ㐣ཤ࡟ࠊ᤼༸ᚋ
ࡢ⤒㐣᫬㛫ࡀ༸ẕ⣽⬊ࡢⓎ⏕⬟࡟ཬࡰࡍᙳ㡪࡟ࡘ࠸࡚ㄪ࡭ࠊ12 ᫬㛫⤒㐣ࡋࡓ༸ẕ⣽⬊(12 ᫬㛫༸Ꮚ)࡛ ࡣ య እ ཷ⢭ (IVF) ᚋ࡟ ཷ ⢭ ⋡ ࡀప ୗ ࡋ (8.0 s1.1㸣)ࠊ⬇┙⬊ᮇࡲ࡛ࡢⓎ⏕ࡀ㉳ࡇࡽ࡞ࡃ࡞ࡿࡇ࡜ࠊ
ࡲࡓࠊ24 ᫬㛫⤒㐣ࡋࡓ༸ẕ⣽⬊(24 ᫬㛫༸Ꮚ)࡛ࡣ
༸๭⮬య㉳ࡇࡽ࡞࠸ࡇ࡜ࢆ☜ㄆࡋࡓࠋḟ࡟ࠊࡇࡢ
ࡼ࠺࡞Ⓨ⏕ᨭᣢ⬟ࢆᦆ࡞ࡗࡓ➨஧ῶᩘศ⿣୰ᮇ(MII ᮇ)༸ẕ⣽⬊ࡢ᰾࡜ᑡ㔞ࡢ⣽⬊㉁ࢆ྾ᘬࡋࠊ㝖᰾ࡋࡓ᪂㩭༸ẕ⣽⬊㉁࡜⼥ྜࡉࡏࡓ MII ᮇ༸ẕ⣽⬊᰾⨨᥮ࢆヨࡋࡓ࡜ࡁࡢ෌ᵓ⠏༸ẕ⣽⬊ࡢ IVF ᚋࡢⓎ⏕⬟ຊࢆㄪ࡭ࡿ࡜࡜ࡶ࡟ࠊಶయࡲ࡛ࡢⓎ⏕⋡ࠊ࠾ࡼࡧࠊฟ⏕ᚋࡢቑయ⋡ࡸ㣫ᩱ᥇㣗㔞࡟ࡘ࠸࡚ㄪ
࡭ࡓࠋᐇ㦂ࡢ⤖ᯝࠊ12 ᫬㛫༸Ꮚ᰾⏤᮶ࡢ IVF ⬇࡛ࡣࠊཷ⢭⋡ࡀ 84.6 s 14.1%ࠊ⬇┙⬊ᮇࡲ࡛ࡢⓎ⏕⋡ࡀ 53.8 s 17.9%࡛࠶ࡾࠊ㏻ᖖࡢ IVF ⬇(ཷ⢭⋡:8.0 s 1.1㸣; ⬇┙⬊ᙧᡂ⋡: 8.0 s 1.1㸣)࡜ẚ㍑
ࡋࡓ࡜ࡇࢁࠊ᭷ព࡞ᕪࡣࡳࡽࢀ࡞࠿ࡗࡓࠋࡇࡢ࠺
ࡕ 23 ಶࡢ⬇┙⬊ᮇ⬇ࢆ⬇⛣᳜࡟౪ࡋ࡚ಶయࡲ࡛
ࡢⓎ⏕⬟ࢆㄪ࡭ࡓ࡜ࡇࢁࠊ10 ༉ࡢ᪂⏕Ꮚࢆ☜ㄆࡋࡓࠋࡲࡓࠊᚓࡽࢀࡓ࣐࢘ࢫࡢ᥇㣗㔞࠾ࡼࡧቑయ⋡ࢆ㏻ᖖࡢ IVF ⬇⏤᮶࣐࢘ࢫ࡜ẚ㍑ࡋࡓ࡜ࡇࢁࠊ㢧ⴭ࡞ᕪࡣࡳࡽࢀ࡞࠿ࡗࡓࠋࡇࢀࡽࡢ⤖ᯝ࡟ࡼࡾࠊ
ཷ⢭⬟ࡸⓎ⏕ᨭᣢ⬟ࡀⴭࡋࡃపୗࡋࡓ 12 ᫬㛫༸Ꮚࡢ᰾࡛ࡶࠊMII ᮇ᰾⨨᥮ࡋ IVF ࡟ࡼࡾቑయࡸ᥇
㣗㔞࡟࠾࠸࡚㏻ᖖ IVF ⬇⏤᮶࡜ྠ➼ࡢ࣐࢘ࢫࡀస
ฟྍ⬟࡛࠶ࡿࡇ࡜ࡀ♧ࡉࢀࡓࠋࡉࡽ࡟ࠊ24 ᫬㛫༸Ꮚ࡛ࡶྠᵝࡢヨ㦂ࢆ⾜࠸ࠊಶయࡲ࡛ࡢⓎ⏕ᨭᣢ⬟ࢆㄪ࡭ࡓ࡜ࡇࢁࠊ8 ༉ࡢ᪂⏕Ꮚࢆసฟࡍࡿࡇ࡜ࡀ

− 25 −

࡛ࡁࡓ(34.8 r 25%)ࠋࡇࡢࡇ࡜࠿ࡽࠊ᏶඲࡟ཷ⢭⬟ࡸⓎ⏕⬟ຊࡀኻࢃࢀࡓ༸ẕ⣽⬊࠿ࡽ࡛ࡶ᰾⨨᥮᧯సࡍࡿࡇ࡜࡟ࡼࡾ㧗⋡࡟⏘Ꮚࢆసฟࡍࡿࡇ࡜ࡀ࡛
ࡁࡿࡇ࡜ࡀุ᫂ࡋࡓࠋࡋ࠿ࡋ࡞ࡀࡽࠊ24 ᫬㛫༸Ꮚ᰾⏤᮶ࡢ࣐࢘ࢫࡣ᥇㣗㔞ࡸయ㔜ࡀ㏻ᖖࡢ IVF ⬇⏤᮶⏘Ꮚࡼࡾࡶ᭷ព࡟ቑຍࡍࡿࡇ࡜࠿ࡽࠊ㐣ᗘ࡟⪁໬ࡋࡓ༸Ꮚ࡛ࡣ᰾⨨᥮ࢆࡶࡗ࡚᤼༸┤ᚋࡢ༸Ꮚ⣽⬊㉁࡟⛣᳜ࡋࡓ࡜ࡋ࡚ࡶࠊฟ⏕ᚋ࡟␗ᖖࡀ⌧ࢀࡿࡇ࡜ࡀ᫂ࡽ࠿࡟࡞ࡗࡓࠋࡇࢀࡽࡢᐇ㦂⤖ᯝࡣࠊ༸ẕ⣽⬊᰾⨨᥮ἲࡢྍ⬟ᛶ࡜㝈⏺ࢆ♧ࡋ࡚࠸ࡿࡇ࡜
࠿ࡽࠊ࣐࢘ࢫ௨እࡢ௚✀ື≀࡟࠾ࡅࡿᛂ⏝฼⏝ࡣៅ㔜࡟⾜࠺ᚲせࡀ࠶ࡿࠋㄽᩥⓎ⾲
1. Overgrowth of mice generated from postovulatory
aged oocyte spindles. Kouki Shiina1, Masaya
Komatsu, Fumi Yokoi, Hanako Bai, Masashi
Takahashi, and Manabu Kawahara. FASEB
BioAdvances, 2019, IN PRESS.

኱Ꮫ㝔ᕤ
ᕤᏛ◊✲㝔

Kawamoto: "STM/STS on E”-(BEDT-TTF)4[(H3O)Ga(C2O4)3]
࣭C6H5NO2", 4th International Conference on Functional
Materials Science 2018 (4th ICFMS 2018) inconjunction with 2nd RIKEN Symposium, November
13-16, 2018, Bali, Indonesia.
Harison Rozak, Wan Nurfadhilah Zaharim, Issei
Miyazaki, Nur Eliana Ismail, Siti Nuramira Abu Bakar,
Daruliza Kernain, Razip Samian, Koichi Ichimura,
Mohamed Ismail Mohamed-Ibrahim, Shukri Sulaiman,
Isao Watanabe: "The Study of the Structure of 12-Mer
Single-Stranded Polyadenine and Its Relationship with
Electrical Conductivity", 4th International Conference
on Functional Materials Science 2018 (4th ICFMS
2018) in conjunction with 2nd RIKEN Symposium,
November 13-16, 2018, Bali, Indonesia.
୹⏣⪽㸪୰ὠᕝၨ἞㸪㓇㒊኱ᶞ㸪๽ᓽ㸪ᮎỌ࿴▱㸸༢ᒙࠊ2 ᒙࠊ3 ᒙࡢ 1T-TaS2 ࡢ┤᥋ほᐹ㸬᪥ᮏ≀⌮Ꮫ఍➨ 73 ᅇᖺḟ኱఍㸪2018 ᖺ 3 ᭶ 22-25 ᪥㸪ᮾி⌮⛉኱Ꮫ㔝⏣࢟ࣕࣥࣃࢫ㸦㔝⏣ᕷ㸧㸬㯮⃝ᚭ㸪㧗ရ༤ග㸪㏆⸨ᗈᗣ㸪ᕷᮧ᫭୍㸪᱈㔝┤ᶞ㸪ྜྷ⏣⣫⾜㸪ᑠ⏣◊㸪ఀᅵᨻᖾ㸪୹⏣⪽㸸STM/STS࠿ࡽࡳࡓ 1T-TaS2 ࡢ㟁Ꮚ≧ែ II㸬᪥ᮏ≀⌮Ꮫ఍➨73 ᅇᖺḟ኱఍㸪2018 ᖺ 3 ᭶ 22-25 ᪥㸪ᮾி⌮⛉኱Ꮫ㔝⏣࢟ࣕࣥࣃࢫ㸦㔝⏣ᕷ㸧㸬ᛂ⏝≀⌮Ꮫ㒊㛛 㔞Ꮚ≀ᛶᕤᏛศ㔝୹⏣ ⪽
㸦⣧Ỉ฼⏝㸧
Ꮫ⾡ㄽᩥ
Nomura, A., Yamaya, K., Takayanagi, S., Ichimura, K.,
Matsuura, T., and Tanda, S. (2018) "Effect of Cu
Doping on Superconductivity in TaSe3: Relationshipbetween Superconductivity and Induced Charge
Density Wave", Europhysics Letters 124: 67001-1-7.
Ꮫ఍Ⓨ⾲
Koichi Ichimura, Hiroki Kokubo, Satoshi Tanda, Tohru
Kurosawa, Hiroyuki Yoshida, Migaku Oda, Masayuki
Ido, Hiroki Honma, Noriaki Matsunaga, Kazuto Moribe,
Yoshihiko Ihara, Atsushi Kawamoto: "STM/STS on theCharge Ordering State in E”-(BEDT-TTF)4[(H3O)Ga(C2O4)3]
࣭C6H5NO2", International Conference on Science andTechnology of Synthetic Metals 2018, July 1-6, 2018,
Busan, Korea.
Syuma Yasuzuka, Shinya Uji, Shiori Sugiura, Taichi
Terashima, Yoshio Nogami, Koichi Ichimura, Satoshi
Tanda: "In-Plane Anisotropy of Upper Critical Field in
Layered Transition Metal Dichalcogenide NbSe2",
International Conference on Science and Technology
of Synthetic Metals 2018, July 1-6, 2018, Busan,
Korea.
Koichi Ichimura, Hiroki Kokubo, Satoshi Tanda, Tohru
Kurosawa, Migaku Oda, Hiroki Honma, Noriaki
Matsunaga, Kazuto Moribe, Yoshihiko Ihara, Atsushi㏆⸨ᗈᗣ㸪୰ᮧ୍㸪ᘏවၨ⣧㸪୰ὠᕝၨ἞㸪ᕷᮧ᫭୍㸪㯮⃝ᚭ㸪୹⏣⪽㸸1T-TaS2 ⷧ⭷ヨᩱࡢ☢Ẽ᢬ᢠ࡜࣮࣍ࣝ᢬ᢠ ᐃ㸬᪥ᮏ≀⌮Ꮫ఍➨ 73 ᅇᖺḟ኱఍㸪2018 ᖺ 3 ᭶ 22-25 ᪥㸪ᮾி⌮⛉኱Ꮫ㔝⏣࢟ࣕࣥࣃࢫ㸦㔝⏣ᕷ㸧㸬୰ᮧ୍㸪ᘏවၨ⣧㸪㏆⸨ᗈᗣ㸪ᕷᮧ᫭୍㸪୹⏣⪽㸸࡬ࡁ㛤ⷧ⭷໬ࡋࡓᒙ≧㖡㓟໬≀⤯⦕యࡢ㍺㏦⌧㇟㸬᪥ᮏ≀⌮Ꮫ఍➨ 73 ᅇᖺḟ኱఍㸪2018 ᖺ 3 ᭶ 22-25᪥㸪ᮾி⌮⛉኱Ꮫ㔝⏣࢟ࣕࣥࣃࢫ㸦㔝⏣ᕷ㸧㸬୰ὠᕝၨ἞㸪⸨஭ᩄஅ㸪୹⏣⪽㸸ࢹࢥࣄ࣮ࣞࣥࢫ
࡟ࡼࡿ Quantum Time Crystal㸬᪥ᮏ≀⌮Ꮫ఍➨ 73ᅇᖺḟ኱఍㸪2018 ᖺ 3 ᭶ 22-25 ᪥㸪ᮾி⌮⛉኱Ꮫ㔝⏣࢟ࣕࣥࣃࢫ㸦㔝⏣ᕷ㸧㸬ᮏ㛫ᘯᶞ㸪ᘏවၨ⣧㸪ᕷᮧ᫭୍㸪Ἑᮏ඘ྖ㸪ᯇỌᝅ᫂㸸ᨃ୍ḟඖ᭷ᶵᑟయ(TMTTF)2X ࡢ STM ศග㸬2018 ᖺ 3 ᭶ 22-25 ᪥㸪ᮾி⌮⛉኱Ꮫ㔝⏣࢟ࣕࣥࣃࢫ㸦㔝⏣ᕷ㸧
㸬Ᏻሯ࿘☻㸪Ᏹ἞㐍ஓ㸪ᮡᾆᰭ⌮㸪ᑎᔱኴ୍㸪㔝ୖ⏤ኵ㸪ᕷᮧ᫭୍㸪୹⏣⪽㸸2H-NbSe2 ࡟࠾ࡅࡿୖ㒊⮫⏺☢ሙࡢ㠃ෆ␗᪉ᛶ㸬2018 ᖺ 9 ᭶ 9-12 ᪥㸪ྠᚿ♫኱Ꮫி⏣㎶࢟ࣕࣥࣃࢫ㸦ி⏣㎶ᕷ㸧㸬− 26 −㔝ᮧ 㸪ᒣ㇂࿴ᙪ㸪㧗ᰗ⁠㸪ᕷᮧ᫭୍㸪ᯇᾆᚭ㸪୹⏣⪽㸸Cu ࢻ࣮ࣉࡋࡓ TaSe3 ࡟࠾ࡅࡿ㟁ⲴᐦᗘἼ(CDW)ࡢฟ⌧㸬2018 ᖺ 9 ᭶ 9-12 ᪥㸪ྠᚿ♫኱Ꮫி⏣㎶࢟ࣕࣥࣃࢫ㸦ி⏣㎶ᕷ㸧㸬㔝ᮧ 㸪ᒣ㇂࿴ᙪ㸪㧗ᰗ⁠㸪ᕷᮧ᫭୍㸪୹⏣⪽㸸Cu ࢻ࣮ࣉࡋࡓ TaSe3 ࡟࠾ࡅࡿ㉸ఏᑟ㸸㉸ఏᑟ࡜ฟ⌧ࡋࡓ㟁ⲴᐦᗘἼ(CDW)ࡢ㛵ಀ㸬2018 ᖺ 9 ᭶ 9-12᪥㸪ྠᚿ♫኱Ꮫி⏣㎶࢟ࣕࣥࣃࢫ㸦ி⏣㎶ᕷ㸧㸬ᕷᮧ᫭୍㸪ᑠஂಖᏹᶞ㸪୹⏣⪽㸪㯮⃝ᚭ㸪ᑠ⏣◊㸪ᮏ㛫ᘯᶞ㸪ᯇỌᝅ᫂㸪᳃㒊୍ᩯ㸪஭ཎ៞ᙪ㸪Ἑᮏ඘ྖ㸸E”-(BEDT-TTF)4[(H3O)Ga(C2O4)3]࣭C6H5NO2
ࡢ STM/STS㸬2018 ᖺ 9 ᭶ 9-12 ᪥㸪ྠᚿ♫኱Ꮫி⏣㎶࢟ࣕࣥࣃࢫ㸦ி⏣㎶ᕷ㸧㸬ᐑᔘ୍ㄔ㸪ᕷᮧ᫭୍㸪୹⏣⪽㸪㯮⃝ᚭ㸪ᑠ⏣◊㸸
FeTe ࡢ STM/STS㸬2018 ᖺ 9 ᭶ 9-12 ᪥㸪ྠᚿ♫኱Ꮫி⏣㎶࢟ࣕࣥࣃࢫ㸦ி⏣㎶ᕷ㸧㸬
㏆⸨ᗈᗣ㸪㧘ရ༤ග㸪ᘏවၨ⣧㸪୰ᮧ୍㸪୰ὠᕝၨ἞㸪ᕷᮧ᫭୍㸪㯮⃝ᚭ㸪୹⏣⪽㸸1T-TaS2 ⷧ⭷ヨᩱࡢ☢Ẽ᢬ᢠ࡜࣮࣍ࣝ᢬ᢠ ᐃ II㸬
2018 ᖺ 9 ᭶ 9-12᪥㸪ྠᚿ♫኱Ꮫி⏣㎶࢟ࣕࣥࣃࢫ㸦ி⏣㎶ᕷ㸧㸬Harison Rozak, Fadhilah Zaharim, Fahmi Astuti,
Eliana Ismail, Amira Abu Bakar, Issei Miyazaki,
Shukri Sulaiman, Mohamed Ismail Mohamed-Ibrahim,
Koichi Ichimura, Isao Watanabe 㸸 PSR Studies on
Electron Motion in DNA. 2018 ᖺ 9 ᭶ 9-12 ᪥㸪ྠᚿ♫኱Ꮫி⏣㎶࢟ࣕࣥࣃࢫ㸦ி⏣㎶ᕷ㸧㸬࢚ࢿࣝࢠ࣮⎔ቃࢩࢫࢸ࣒㒊㛛ᛂ⏝࢚ࢿࣝࢠ࣮ࢩࢫࢸ࣒ศ㔝㏆ஂ Ṋ⨾㸦⣧Ỉ฼⏝㸧Ⓨ⾲ㄽᩥAoyama, Y., Tabe, Y., Nozaki, R., Suzuki, K., Chikahisa, T.,
and Tanuma, T. (2018) Analysis of Water Transport
inside Hydrophilic Carbon Fiber Micro-Porous
Layers with High-Performance Operation in PEFC.
Journal of The Electrochemical Society 165(7):F484-F491.
Tabe, Y., Sakaida, S., and Chikahisa, T. (2018) Scalemodel experiments for evaluation of liquid watertransport in the gas diffusion layer of PEFCs.
Journal of Thermal Science and Technology
13(2): 1-15.
Onishi, F., Tabe, Y., and Chikahisa, T. (2018) Experimental
Study on the Balance between Microscopic Water
Production and Temperature Rise during Cold
Startup in PEFC. ECS Transactions 86(13): 89-96.

Iiri, T., Tabe, Y., and Chikahisa, T. (2018) Experimental
Analysis of Oxygen Transport Resistance forDifferent Types of Ionomer in PEFC Catalyst.
ECS Transactions 86(13): 141-150.
Satake, T., Tabe, Y., and Chikahisa, T. (2018) Analysisof Oxygen Transport Resistances in the Catalyst
Layers with Different Carbon Supports in PEFC.
ECS Transactions 86(13): 171-178.
Ꮫ఍Ⓨ⾲㔝ᓮᾴ㸪⏣㒊㇏㸪㏆ஂṊ⨾㸸PEFC ࡢప ࣭㧗 ᮲௳ୗ࡟࠾ࡅࡿゐ፹ᒙෆእࡢ㓟⣲㍺㏦᢬ᢠゎᯒ㸬➨ 55 ᅇఏ⇕ࢩ࣏ࣥࢪ࣒࢘㸪2018 ᖺ 5
᭶ 29 ᪥㸪ᮐᖠࢥࣥ࣋ࣥࢩࣙࣥࢭࣥࢱ࣮㸦ᮐᖠᕷ㸧㸬㣤฼ᣅᐇ㸪⏣㒊㇏㸪㏆ஂṊ⨾㸸ᅛయ㧗ศᏊᙧ⇞ᩱ㟁ụࡢゐ፹ᒙෆ࢖࢜ࣀ࣐࣮࠾ࡼࡧ࣮࢝࣎ࣥ
✀㢮࡟ᑐࡍࡿ㓟⣲㍺㏦᢬ᢠẚ㍑㸬➨ 55 ᅇఏ⇕ࢩ࣏ࣥࢪ࣒࢘㸪2018 ᖺ 5 ᭶ 29 ᪥㸪ᮐᖠࢥࣥ࣋ࣥࢩࣙࣥࢭࣥࢱ࣮㸦ᮐᖠᕷ㸧㸬኱すྐே㸪⏣㒊㇏㸪㏆ஂṊ⨾㸸PEM ᆺ⇞ᩱ㟁ụࡢᗘୖ᪼㐣⛬࡟࠾ࡅࡿịⅬୗ㉳ື≉ᛶ㸬➨55 ᅇఏ⇕ࢩ࣏ࣥࢪ࣒࢘㸪2018 ᖺ 5 ᭶ 29 ᪥㸪ᮐᖠࢥࣥ࣋ࣥࢩࣙࣥࢭࣥࢱ࣮㸦ᮐᖠᕷ㸧㸬బ➉Ꮥಖ㸪⏣㒊㇏㸪㏆ஂṊ⨾㸸PEFC ࢝ࢯ࣮ࢻゐ፹ᒙ࡟࠾ࡅࡿ࣮࢝࣎ࣥᢸయᵓ㐀ࡀ㓟⣲㍺㏦࡟
ཬࡰࡍᙳ㡪ゎᯒ㸬➨ 55 ᅇఏ⇕ࢩ࣏ࣥࢪ࣒࢘㸪2018 ᖺ 5 ᭶ 29 ᪥㸪ᮐᖠࢥࣥ࣋ࣥࢩࣙࣥࢭ
ࣥࢱ࣮㸦ᮐᖠᕷ㸧㸬
Kumar, A., Iiri, T., Satake, Y., Tabe, Y., and Chikahisa,
T. Trial of applying graphene nano platelet tocatalyst layers of a PEFC. The 16th
International Heat Transfer Conference(IHTC-16), 2018 ᖺ 8 ᭶ 10 ᪥-15 ᪥, China
National Convention Center (Beijing, China)㸬
ၿᙜဴஓ㸪⏣㒊㇏㸪㏆ஂṊ⨾㸸ࣞࢻࢵࢡࢫࣇ࣮ࣟ
㟁ụᛶ⬟࡟ᑐࡍࡿᵓ㐀࣭㐠㌿᮲௳ᙳ㡪ࡢ㞟⣙୺せᅉᏊ㸬᪥ᮏᶵᲔᏛ఍ 2018 ᖺᗘᖺḟ኱఍㸪2018 ᖺ 9 ᭶ 11 ᪥㸪㛵す኱Ꮫ༓㔛ᒣ࢟ࣕࣥࣃࢫ㸦྿⏣ᕷ㸧㸬ὸᕝኟ㉺㸪⏣㒊㇏㸪㏆ஂṊ⨾㸸ᑠᆺࢥࢪ࢙ࢿ࣮ࣞ
ࢩࣙࣥᗫ⇕฼⏝෭ᡣࡢࡓࡵࡢάᛶⅣ-࢚ࢱࣀ࣮ࣝ྾╔≉ᛶ࡟㛵ࡍࡿ◊✲㸬᪥ᮏᶵᲔᏛ
఍⇕ᕤᏛࢥࣥࣇ࢓ࣞࣥࢫ㸪2018 ᖺ 10 ᭶ 20᪥㸪ᐩᒣ኱Ꮫ஬⚟࢟ࣕࣥࣃࢫ㸦ᐩᒣᕷ㸧㸬Can, E.M., Tabe, Y., Chikahisa, T., and Sakaida, S.㸸
Effect of wettability design on water transport
in GDL of PEFCs analyzed by scale modelexperiment and LBM simulation㸬᪥ᮏᶵᲔᏛ
఍➨ 9 ᅇ࣐࢖ࢡ࣭ࣟࢼࣀᕤᏛࢩ࣏ࣥࢪ࣒࢘㸪2018 ᖺ 10 ᭶ 31 ᪥㸪ᮐᖠᕷẸ஺ὶࣉࣛࢨ㸦ᮐᖠᕷ㸧㸬− 27 −

(2019) Defecation initiates walking in the cricket
Gryllus bimaculatus. J. Insect Physiol. 112: 117-122.

㟁Ꮚ⛉Ꮫ◊✲ᡤ㝃ᒓ♫఍๰㐀ᩘᏛ◊✲ࢭࣥࢱ࣮」㞧⣔ᩘ⌮◊✲ศศ㔝㟷἟ ோᚿ
ࣟ࣎ࢵࢺࡢࡼ࠺࡞ேᕤ≀ࡣ㸪୍้้࡜ኚ໬ࡋ㸪ண ࡀᅔ㞴࡞↓㝈ᐃ࡞⎔ቃ࡛ࡢືసࡣ㠀ᖖ࡟ᅔ㞴࡛࠶ࡿ㸬⏘ᴗ⏝ࡢࣟ࣎ࢵࢺࡣ㸪୍ぢ㎿㏿࡛ࡋ࡞ࡸ࠿࡞ືసࢆࡋ࡚࠸ࡿࡀ㸪ᐇࡣ㸪㝈ࡽࢀࡓ⎔ቃࡢ୰ࡔࡅ࡛࠺ࡲࡃືసࡍࡿࡇ࡜ࡀ࡛ࡁ㸪ࡦ࡜ࡓࡧ↓㝈ᐃ࡞⎔ቃ࡟ᣢࡕฟࡉࢀࡓࡽ㸪ࡓࡕࡲࡕṆࡲࡗ࡚ࡋࡲ࠺㸬ࣟ࣎ࢵࢺࡣ㸪࿘ࡾࡢ⎔ቃࢆほ ࡋ㸪ほ ⤖ᯝ࡟ᛂࡌ࡚ືసฟຊࢆỴࡵ࡚࠸ࡿࡓࡵ࡛㸪ィ 㝈⏺ࢆ㉸࠼ࡓ⎔ቃୗ࡛ࡣືస࡛ࡁ࡞࠸࠿ࡽ࡛࠶ࡿ㸬୍᪉㸪ື≀ࡣ㸪↓㝈ᐃ࡞⎔ቃࡢ୰࡛ࡶ㐺ᛂⓗ࡟᣺ࡿ⯙ࡗ࡚࠸ࡿ㸬᪤▱ࡢ⎔ቃࡢ୰࡛ࡋ࠿ேᕤ≀ࢆไᚚࡍࡿ᪉ἲࡋ࠿ᣢࡕྜࢃࡏ࡚࠸࡞࠸ᡃࠎ࡟࡜ࡗ࡚
ࣇ࣮࣒ࣞၥ㢟࡟ᑐฎࡍࡿ࡟ࡣఱࡽ࠿ࡢᡭᮏࡀᚲせ࡛࠶ࡾ㸪ࡑࢀࡀ⏕≀࡛࠶ࡿ㸬ື≀ࡣ㸪้ࠎ࡜ኚࢃ
ࡿ≧ἣ࡟ᛂࡌ࡚ᐇ᫬㛫࡛㐺ᛂⓗ࡟㐠ືࡸ⾜ືࢆⓎ⌧ࡋ࡚࠸ࡿ㸬
ᡃࠎࡣ㸪ࡇࡢᵝ࡞㐺ᛂⓗ࡞㐠ືࡸ⾜ືࡢᇶ┙࡜
࡞ࡿ㌟యࡸ⬻ࡢ࣓࢝ࢽࢬ࣒ࢆ⌮ゎࡋ㸪᭦࡟ࡑࡇ࠿
ࡽ᪂ወ࡞ேᕤ≀ࡢไᚚㄽࡢ☜❧ࢆ┠ᣦࡋ࡚࠸ࡿ㸬ࡑࡢࡓࡵ࡟㸪ಶయࢆྲྀࡾᕳࡃ⎔ቃࡸಶయࡢ㌟యࡢ
ືసࢆ᧯సࡍࡿࡇ࡜࡛㐺ᛂⓗ࡞㐠ືࡸ⾜ືࡢᇶ┙࡜࡞ࡿไᚚᶵᵓࢆ࠶ࡪࡾฟࡋ⌮ゎࡋࡼ࠺࡜ࡋ࡚࠸
ࡿ㸬᪻⹸ࡢࢥ࢜ࣟࢠ(G. bimaculatus)࡛ࡣ㸪Ṍ⾜ࡢㄏⓎ࡟㛵ࢃࡿ୰ᯡ⚄⤒⣔ࡢാࡁ࡟ࡘ࠸࡚ㄪ࡭ࡓ㸬㏿࡟㸪᤼ἥ⾜ືᚋ࡟ㄏⓎࡍࡿṌ⾜࡟╔┠ࡋ㸪᤼ἥ࡟క࠸ᮎᲈ࠿ࡽࡢୖ⾜ᛶࡢಙྕ࡟ࡼࡾṌ⬮ࡢ㐠ືࡀㄏⓎࡉࢀ㸪ࡑࡢᚋ⬻࠿ࡽࡢୗ㝆ᛶࡢಙྕ࡟ࡼࡾ
ࣜࢬ࣑ࢵࢡ࡞Ṍᐜࣃࢱࣥࡀ⏕ᡂࡉࢀࡿࡇ࡜ࡀࢃ࠿
ࡗࡓ㸬࢔ࣜ(O. kuroiwae)࡛ࡣ㸪➽㦵᱁⣔ࡢᶵᵓ࡜㐠ືไᚚ࡟ࡘ࠸࡚X⥺࢖࣓࣮ࢪࣥࢢἲࢆ⏝࠸࡚ㄪ࡭ࡓ㸬ࡑࡢ⤖ᯝ㸪㦵᱁ࡢᙎᛶኚᙧࢆ฼⏝ࡋࡓ㧗㏿㐠ືࡢⓎ⏕࣓࢝ࢽࢬ࣒ࡢゎ᫂࡟㏆࡙ࡅࡓ㸬ࢡࣔࣄࢺ
ࢹ(O. incrassate)࡛ࡣ㸪㌟యᵓ㐀ࡢኚ໬࡜ࣜࢬ࣑ࢵ
ࢡ࡞㐠ືࣃࢱࣥࡢ⏕ᡂ࣓࢝ࢽࢬ࣒࡟ࡘ࠸࡚㸪ᩘ⌮ࣔࢹࣜࢆᵓ⠏ࡋࡓ㸬ࡑࡢ⤖ᯝ㸪㌟యࡢᵓ㐀࡜⏕ᡂࡉࢀࡿࣜࢬ࣑ࢵࢡ࡞㐠ືࣃࢱࣥࡢ⏕ᡂ࡟ࡘ࠸࡚⌮ゎࡀ῝ࡲࡗࡓ㸬࣒࢝ࢹ(S. rubiginosus㸪S. subspinipes
mutilans)࡛ࡣ㸪⎔ቃ࡟ᛂࡌࡓṌᐜኚ໬࡟ࡘ࠸࡚⬮㛫༠ㄪࡢ࣓࢝ࢽࢬ࣒࡟ࡘ࠸࡚ㄪ࡭ࡓ㸬ࡇࢀࡽᵝࠎ࡞ᐇ㦂ື≀ࢆ⏝࠸ࡓ◊✲࠿ࡽ㸪⎔ቃ࡜㌟య࡜⚄⤒⣔ࡢ┦஫స⏝࠿ࡽ⏕ࡌࡿ㐺ᛂᶵ⬟ࡢⓎ⌧࣓࢝ࢽࢬ࣒࡟ࡘ࠸࡚⪃ᐹࡋ㸪㐺ᛂⓗ࡞᣺ࡿ⯙࠸ࡀ๰Ⓨࡉࢀࡿタィ࡜ࡑࡢไᚚᵓ㐀ࡢ⌮ゎࡀ῝ࡲࡗࡓ㸬ㄽᩥⓎ⾲Naniwa K., Sugimoto Y., Osuka K. and Aonuma H.

Clark E.G., Kanauchi D., Kano T., Aonuma H., Briggs
D. and Ishiguro A. (2019) The function of the
ophiuroid nerve ring: How a decentralized nervous
system controls coordinated locomotion. J. Exp. Biol.
doi: 10.1242/jeb.192104
Watanabe T., Ugajin A. and Aonuma H. (2018)
Immediate-early promoter-driven transgenic reporter
system for neuroethological researches in ahemimetabolous insect. eNeuro. 5(4): e0061-18.
Aonuma H., Totani Y., Sakakibara M., Lukowiak K.
and Ito E. (2018) Comparison of brain monoaminecontent in three populations of Lymnaea that correlateswith taste-aversive learning ability. Biophys. Physicobiol.
15: 129-135.
DiRienzo, N. and Aonuma H. (2018) Plasticity inextended phenotype increases offspring defense despite
individual variation in web structure and behavior.
Animal Behav. 138: 9-17.
Momohara Y., Aonuma H. and Nagayama T. (2018)
Tyraminergic modulation of agonistic outcomes incrayfish. J. Comp. Physiol. 5:465-473.
Aonuma H., Totani Y., Kaneda M., Nakamura R.,
Watanabe T., Hatakeyama D., Dyakonovae V. E.,
Lukowiak K. and Ito E. (2018) Effects of 5-HT andinsulin on learning and memory formation in
food-deprived snails. Neurobiol. Learn. Mem., 148: 20-29.
Ꮫ఍Ⓨ⾲㟷἟ ோᚿ* : ࠕ᪻⹸࡛ぢࡽࢀࡿ㞟ᅋࢧ࢖ࢬ࡟ᛂࡌ
ࡓᨷᧁᛶࡢࣔࢹࣝ໬ࠖ
ࠊ㟁Ꮚ᝟ሗ㏻ಙᏛ఍ NC㸪NLPྜྠ◊✲఍ࠊ໭ᾏ㐨኱ᏛࠊJapan (2019-01)⬥⏣ ኱㍤*ࠊᾉⰼ ၨྑࠊ᪩℩ ཭⨾஀ࠊ㟷἟ ோᚿ :
ࠕAutonomous decentralized coordination in the diskof the green brittle Starࠖࠊィ ⮬ືไᚚᏛ఍➨ 31 ᅇ⮬ᚊศᩓࢩࢫࢸ࣒ࢩ࣏ࣥࢪ࣒࢘ࠊᅜ❧Ẹ᪘༤≀㤋ࠊ
Japan (2019-01)ᾉⰼ ၨྑ*ࠊᮡᮏ 㟹༤ࠊ኱㡲㈡ බ୍ࠊ㟷἟ ோᚿ :ࠕ᤼౽࡟క࠸ㄏⓎࡍࡿࢡࣟࢥ࢜ࣟࢠࡢᐃᆺⓗ⾜ືࠖࠊィ ⮬ືไᚚᏛ఍➨ 31 ᅇ⮬ᚊศᩓࢩࢫࢸ࣒
ࢩ࣏ࣥࢪ࣒࢘ࠊᅜ❧Ẹ᪘༤≀㤋ࠊJapan (2019-01)
A. Matsuda*, H. Aonuma, K. Naniwa and S. Kaneko :“Image-based Measurement of Ultra-fast Movement of
Mandible in Trap-Jaw Ants”, 12th France-Japan and10th Europe-Asia Congress on Mechatronics, ⟃Ἴ,− 28 −

Japan (2018-09)໭᪉⏕≀ᅪࣇ࢕࣮ࣝࢻ⛉Ꮫࢭࣥࢱ࣮᪩℩ ཭⨾஀*ࠊ⬥⏣ ኱㍤ࠊ㟷἟ ோᚿ : ࠕࢡࣔࣄࢺࢹ࡟ぢࡽࢀࡿ࣏ࣥࣆࣥࢢࢲ࢖ࢼ࣑ࢡࢫࠖ
ࠊ᪥ᮏ≀⌮Ꮫ఍ࠊྠᚿ♫኱ᏛࠊJapan (2018-09)
D. Owaki*, Y. Sugimoto, A. Ishiguro and H. Aonuma :“Change in Electromyographic Patterns After Leg
Amputation in the Cricket”, International Congress for
Neuroethology, Brisbane, Australia (2018-07)
H. Aonuma*, K. Naniwa, K. Kagaya and K. Osuka :“Latching mechanisms to generate ultrafast movementof the trap jaw in the ant Odontomachus kuroiwae”,
International Congress for Neuroethology, Brisbane,
Australia (2018-07)㧗ཎ ⪽*ࠊ㟷἟ ோᚿࠊ㔠Ꮚ ಇ୍ : ࠕࢡࣔࣄࢺࢹ
ᶍೌࣟ࣎ࢵࢺタィ࡟ྥࡅࡓ X ⥺࣐࢖ࢡࣟ CT ࡟ࡼࡿ㦵᱁ᙧ≧ᢳฟࠖ
ࠊ➨ 24 ᅇ⏬ീࢭࣥࢩࣥࢢࢩ࣏ࣥ
ࢪ࣒࢘ SSII2018ࠊࣃࢩࣇ࢕ࢥᶓ὾࢔ࢿࢵࢡࢫ࣮࣍ࣝࠊJapan (2018-06)ᯇ⏣ ᮅ㝧*ࠊ㟷἟ ோᚿࠊᾉⰼ ၨྑࠊ㔠Ꮚ ಇ୍ :ࠕ࢔ࢠࢺ࢔ࣜࡢ㢡ࡢ㉸㧗㏿ᅇ㌿㐠ືࡢᶵᵓࣔࢹࣝ
ᵓ⠏ࡢࡓࡵࡢ⏬ീࣃࢱࣥィ ࠖࠊ➨ 24 ᅇ⏬ീࢭࣥࢩࣥࢢࢩ࣏ࣥࢪ࣒࢘ SSII2018ࠊࣃࢩࣇ࢕ࢥᶓ὾࢔
ࢿࢵࢡࢫ࣮࣍ࣝࠊJapan (2018-06)
D. Wakita*, Y. Hayase and H. Aonuma : “Rhythmic
movement of disk in the green brittle star Ophiarachna
incrassata”, 16th International Echinoderm Conference,
Nagoya Univ., Japan (2018-05 ࠥ 2018-06)
T. Kano*, D. Kanauchi, H. Aonuma and A. Ishiguro :“Decentralized control mechanism underlying interand intra-arm coordination the brittle star's locomotion”,
16th International Echinoderm Conference, Nagoya Univ.,
Japan (2018-05 ࠥ 2018-06)⏕≀ከᵝᛶ㡿ᇦ ᾏ⏘⸴㢮㐺ᛂᶵ⬟ศ㔝ᅄࢶ಴ ඾⁠㸦ᾏỈ฼⏝㸧Ⓨ⾲ㄽᩥ
Kawai, T., Galanin, D., Tskhay, Z., Kroupnova, T., andYotsukura, N. (2018) Present status of saccharinian
kelp in southern Primorye and Sakhalin, far-east
Russia, and Northern Hokkaido, Japan, in Sea of Japan.
Algal Resources 11: 33-42.
Kawai, T., Galanin, D., and Yotsukura, N. (2018) Chocolate
and tea made from kelp. Algal Resources 11: 49-51.
Zhang, J., Yao, J., Min Hu, Z., Jueterbock, A., Yotsukura,N., Krupnova, T., Nagasato, C., and Duan D. (2019)
Phylogeographic diversification and postglacial range
dynamics shed light on the conservation of the kelp
Saccharina japonica. Evolutionary Applications: doi:
10.1111/eva.12756
Kawai, T., Klochkova, N., Kogame, K., Abe, T., Galanin,D., and Yotsukura, N. (2019) Costal marine flora innorthern Hokkaido and southern Sakhalin along thecoast of the Sea of Japan. Rishiri Studies 38: 95-98.
Boonprab, K., Matsui, K., Akkabe, Y., Yotsukura, N., andKajiwara, T. (2019) 11-Hydroperoxide eicosanoid-mediated2(E), 4(E)-decadienal production from arachidonic
acid in the brown algae, Saccharina angustata. Journalof Applied Phycology (accepted).
Ꮫ఍Ⓨ⾲ᯘ⿱୍, ⬟Ⓩ㇂ṇᾈ, ᅄࢶ಴඾⁠㸸ᓥ᰿┴㞃ᒱࡢᓥἢᓊࡢࢶࣝ࢔࣓ࣛ 2 ᙧࡢ⩌ⴠᵓ㐀. ᪥ᮏᛂ⏝⸴㢮Ꮫ఍➨ 15 ᅇ኱఍, 2018 ᖺ 5 ᭶ 12 ᪥, ᮾிᾏὒ኱Ꮫ
㸦ᮾி㒔 ༊㸧ᅄࢶ಴඾⁠㸸ࢥࣥࣈ㈨※ࡢಖ඲㸫⎔ቃኚື࡟ࡼࡿ
ࣜࢫࢡ࡜ࡑࡢᑐᛂ㸫. ᖹᡂ 30 ᖺᗘ᪥ᮏỈ⏘Ꮫ఍໭ᾉⰼ ၨྑ*ࠊ㟷἟ ோᚿࠊᮡᮏ 㟹༤ࠊ኱㡲㈡ බ୍ :ᾏ㐨ᨭ㒊኱఍බ㛤ࢩ࣏ࣥࢪ࣒࢘ࠕ⎔ቃኚືࢆ⪃៖ࠕ⬻࡟ࡼࡿ୰ኸไᚚ⣔ࢆ᤼㝖ࡋࡓࢥ࢜ࣟࢠࡢࡩࡿࡲ࠸ ࠖࠊࣟ࣎ࢸ࢕ࢡࢫ࣭࣓࢝ࢺࣟࢽࢡࢫㅮ₇఍ 2018ࠊ ࡋࡓ໭ᾏ㐨ࡢỈ⏘ቑṪࡢᒎᮃࠖ, 2018 ᖺ 11 ᭶ 23
᪥, 㔲㊰ᕷ⏕ᾭᏛ⩦ࢭࣥࢱ࣮㸦㔲㊰ᕷ㸧⚟ᒸࠊJapan (2018-06)
H. Aonuma* : “Oscillator model to understand groupsize dependent behavior in the cricket”, The 3rd A3
International Workshop for Mathematical and Life
Sciences, Hiroshima Univ., Japan (2018-05)− 29 −

すᲷ㑇ఏᏊᐇ㦂ඹ
ඹྠ฼⏝㒊㛛
ື≀ᰁⰍయඹ
ඹྠ฼⏝㒊㛛

− 31 −㑇 ఏᏊᐇ㦂ඹྠ฼⏝㒊㛛㛛  ᙜ㒊㛛࡛ࡣࠊ㑇ఏᏊ⤌ࡳ᥮࠼ᐇ㦂࡟ᚲせ࡞ࢡ࣮ࣜࣥ࣋ࣥࢳࡸᏳ඲࢟ࣕࣅࢿࢵࢺࠊ኱
ᆺ࣮࢜ࢺࢡ࣮ࣞࣈ➼ࡢ≉Ṧタഛࢆഛ࠼࡚࠾ࡾࠊ⥅⥆ⓗ࡟ά⏝ࡉࢀ࡚࠸ࡲࡍࠋ㑇ఏᏊ⤌᥮࠼ᐇ㦂ࡀ୍⯡໬ࡋࡓࡇ࡜࠿ࡽ≉Ṧタഛ฼⏝௨እࡢ฼⏝⪅ࡣῶᑡࡋࡲࡋࡓࡀࠊ◊✲㐣⛬୍࡛ᐃᮇ㛫୍᫬ⓗ࡟ᚲせ࡟࡞ࡿࡶࡢࡢ◊✲ᐊ࡛‽ഛࡍࡿ࡟ࡣ㈇ᢸ࡜࡞ࡿࡼ࠺࡞ᶵჾ
ࡸࠊ◊✲ᐊࡢ❧ࡕୖࡆ᫬࡟ᶵჾࡀᩚ࠺ࡲ࡛ࡢᮇ㛫ࠊࡶࡋࡃࡣᨾ㞀ᶵჾࡢ⿵ಟ᏶஢ࡲ࡛
࡟฼⏝࡛ࡁࡿྛ✀㑇ఏᏊゎᯒᶵჾࢆ඲Ꮫ࡟ᥦ౪ࡋ࡚࠸ࡲࡍࠋࡋ࠿ࡋࠊ඲యⓗ࡟タഛ➼ࡢ⪁ᮙ໬ࡀ㐍ࡳ㸪ࡉࡽ࡟ᖹᡂ  ᖺ  ᭶࡟ࡣ኱つᶍ࡞₃Ỉ஦ᨾࡀⓎ⏕ࡋ࡚࠾ࡾࠊࡑࡢᑐ
⟇ࡀᛴົ࡛ࡍࠋ ᖹᡂ  ᖺ  ᭶࡟㉳ࡁࡓᮾ᪥ᮏ኱㟈⅏࡟ࡼࡾࠊ◊✲ᶵ㛵࡛ከࡃࡢ⏕≀ᮦᩱ࣭㑇ఏ㈨※ࡀኻࢃࢀ◊✲ࡢ⥅⥆࡟ᨭ㞀ࢆࡁࡓࡋࡲࡋࡓࠋ෌ࡧࡇࡢࡼ࠺࡞ࡇ࡜ࡀ㉳ࡇࡿ࡜ᡃࡀᅜࡢ
ᅜ㝿ⓗ➇தຊ࡟ࡶᝏᙳ㡪ࢆ୚࠼࠿ࡡ࡞࠸࡜ࡢ༴᝹࠿ࡽࠊ᪥ᖖⓗ࡟◊✲⏝⏕≀㑇ఏ㈨※ࡢಖ⟶ࢆ⾜࠺ᩥ㒊⛉Ꮫ┬ࣉࣟࢪ࢙ࢡࢺࠕ኱Ꮫ㐃ᦠࣂ࢖࢜ࣂࢵࢡ࢔ࢵࣉࣉࣟࢪ࢙ࢡࢺ㸦IBBP㸧ࠖࡀឡ▱┴ᒸᓮᕷࡢᇶ♏⏕≀Ꮫ◊✲ᡤࢆ୰᰾ᣐⅬ࡜ࡋ࡚㛤ጞࡉࢀࠊᙜࢭࣥࢱ࣮
ࡣ໭ᾏ㐨ᆅ༊ࡢࢧࢸࣛ࢖ࢺᣐⅬ࡜ࡋ࡚ࡇࡢࣉࣟࢪ࢙ࢡࢺ࡟ཧ⏬ࡋ࡚࠸ࡲࡍࠋື ≀ᰁⰍయඹྠ฼⏝㒊㛛㛛 ື≀ᰁⰍయඹྠ฼⏝㒊㛛࡛ࡣ⣽⬊ᇵ㣴タഛ࡜ࡋ࡚ࠊࢡ࣮ࣜࣥ࣋ࣥࢳࠊ&2 ࢖ࣥ࢟ࣗ࣋
࣮ࢱ࣮ࠊ㐲ᚰᶵ࡞࡝ࢆࠊࡲࡓ⣽⬊ಖᏑタഛ࡜ࡋ࡚ࡣࠊᑓ⏝ࡢΥࢹ࢕࣮ࣉࣇ࣮ࣜࢨ࣮
࠾ࡼࡧ኱ᆺᾮయ❅⣲෾⤖ಖᏑᐜჾࢆഛ࠼࡚࠸ࡲࡍࠋᙜ㒊㛛࡛ࡣࠊ⣔⤫ศ㢮Ꮫࡸ㐍໬Ꮫⓗ࡟฼⏝౯್ࡀ㧗࠸࡜ᛮࢃࢀࡿึ௦ᇵ㣴⣽⬊㢮㸸့ங㢮㸦 ┠  ⛉  ✀  ள✀㸧ࠊ㫽㢮㸦 ┠  ⛉  ✀㸧ࠊ∐⹸㢮㸦 ┠⛉✀㸧࡜ࠊⓎ⏕Ꮫࡸ⏕໬Ꮫࠊ⣽⬊㑇ఏᏛ࡞࡝ࡢᐇ㦂࡟⏝࠸ࡽࢀࡿྛ✀ᇵ㣴⣽⬊ᰴࢆ෾⤖ಖᣢࡋ࡚࠾ࡾࠊᚲせ࡟ᛂࡌ࡚ෆእࡢ◊✲⪅࡬ࡢศ୚ࢆ⾜ࡗ࡚࠸ࡲࡍࠋࡇࢀࡽࡢ⣽⬊ࡢ࠺ࡕከࡃࡢࡶࡢࡣᪧ⌮Ꮫ㒊㝃ᒓື≀ᰁⰍయ◊✲᪋タ࡟࠾࠸࡚ᶞ❧ࠊ࠶ࡿ࠸ࡣ཰㞟ࡉࢀࡓࡶࡢ࡛࠶ࡾࠊྠ᪋タࡢ⤫ᗫྜᚋࡣࠊ୺࡟す⏣༓㭯Ꮚඛ⏕ࡢࡈດຊ࡟ࡼࡗ࡚⥔ᣢࡉࢀ࡚ࡁࡓࡶࡢ࡛ࡍࠋࡇࡢࡼ࠺࡞Ṕྐⓗ࡟㈗㔜࡞⣽⬊ࢆ௒ᚋࡶ౪୚ྍ⬟࡞≧ែ࡛⥔ᣢࡋ⥆ࡅࡿ࡜࡜ࡶ࡟ࠊᏛෆ࡛ࡢඛ➃ⓗ࡞◊✲࡟⏝࠸ࡽࢀࡓྛ✀ᇵ㣴⣽⬊㢮ࡸ⏕≀ヨᩱࢆᆅ㐨࡟཰㞟ࠊಖᏑࡋ࡚࠸ࡃࡇ࡜ࡶ௒ᚋࡢㄢ㢟࡛ࡍࠋ ᖺ  ᭶ࡢ໭ᾏ㐨⫹᣺ᮾ㒊ᆅ㟈࡟క࠺㟁※႙ኻ࡛ࡣࠊᏛෆࡢከࡃࡢ◊✲ᐊ࡛㈗㔜࡞ヨᩱࡀኻࢃࢀࡲࡋࡓࠋ⏕యヨᩱࡢಖᏑ࡜౪⤥࡟㛵ࢃࡗ࡚ࡁࡓᙜ㒊㛛࡜ࡋ࡚ࡣࠊ
ヨᩱಖ⟶࡟㛵ࡍࡿࣂࢵࢡ࢔ࢵࣉሙᡤࡢᥦ౪ࡸ⥭ᛴ᫬࡟࠾ࡅࡿヨᩱࡢ㑊㞴ሙᡤࡢᥦ౪࡞
࡝࡟ࡘ࠸࡚⪃࠼࡚࠸ࡃᚲせࡀ࠶ࡿ࡜ᛮࢃࢀࡲࡍࠋ

− 32 −

ඹྠ฼⏝ᶵჾ㻌䞉㻌タഛ䝯䞊䜹䞊ྡ䞉ᆺ␒➼タഛ䛚䜘䜃ᶵჾྡ
ᇵ㣴ᐊ㻌䠄㻝㻜䟝䠅ᇵ㣴ᐊ㻌䠄㻝㻠䟝䠅㻼㻝ᐇ㦂ᐊ㻼㻞ᐇ㦂ᐊ㻌䠄Ᏻ඲䜻䝱䝡䝛䝑䝖䠅పᐇ㦂ᐊ㻌䠄㻠䉝䠅

⣽⬊ಖᏑᐊᾮయ❅⣲ᘧ෾⤖ಖᏑᐜჾኴ㝧᪥㓟㻌䞉㻰㻾㻙㻝㻜㻜㻜㻾㻵㻌䞉⺯ග䝞䜲䜸䜲䝯䞊䝆䝱䞊㻲㼁㻶㻵㻌㻲㻵㻸㻹㻌䞉㻲㻸㻭㻙㻣㻜㻜㻜

໬ᏛⓎග᧜ᙳ⿦⨨㻭㼀㼀㻻㻌䞉㻱㼆㻙㻯㼍㼜㼠㼡㼞㼑䝀䝹᧜ᙳ⿦⨨

㻭㼀㼀㻻㻌䞉㻭㻱㻙㻢㻥㻟㻞䝀䝹᧜ᙳ⿦⨨

㻭㼀㼀㻻㻌䞉㻭㻱㻙㻢㻥㻞㻜㑇ఏᏊᑟධ⿦⨨㻌䠄䜶䝺䜽䝖䝻䝫䝺䞊䝍䞊䠅㻮㼕㼛㻙㻾㼍㼐㻌䞉㻳㼑㼚㼑㻌㻼㼡㼘㼟㼑㼞㑇ఏᏊᑟධ⿦⨨㻌䠄䜶䝺䜽䝖䝻䝫䝺䞊䝍䞊䠅㻿㻴㻵㻹㻭㻰㼆㼁㻌䞉㻳㼀㻱㑇ఏᏊᑟධ⿦⨨㻌䠄㑇ఏᏊ㖠䠅㻮㼕㼛㻙㻾㼍㼐㻌䞉㻴㼑㼘㼕㼛㼟㻌㻳㼑㼚㼑㻌㻳㼡㼚䝹䝭䝜䝯䞊䝍䞊

㻭㼀㼀㻻㻌䞉㻭㻮㻙㻞㻞㻜㻜᳜≀ᇵ㣴䝏䝱䞁䝞䞊㻌䠄䝁䜲䝖䝖䝻䞁䠅䠄㻟ྎ䠅㻷㻻㻵㼀㻻㻌䞉㻴㻺㻸㻙㻝㻜㻭䠄㻞ྎ䠅㻘㻌㻴㻺㻮㻙㻝㻜㻭䠄㻝ྎ䠅᳜≀䜲䞁䜻䝳䝧䞊䝍䞊䠄ேᕤẼ㇟ჾ䠅㼀㻻㻹㼅㻌䞉㻯㻸㻱㻙㻟㻜㻡኱ᆺᜏ᣺䛸䛖ᇵ㣴ᶵ㻌䠄䝞䜲䜸䝅䜵䞊䜹䞊䠅㼀㻭㻵㼀㻱㻯㻌䞉㻮㻾㻙㻟㻜㻜㻸㻲኱ᆺᜏ᣺䛸䛖ᇵ㣴ᶵ㻌䠄䝞䜲䜸䝅䜵䞊䜹䞊䠅㼀㻭㻵㼀㻱㻯㻌䞉㻮㻾㻙㻝㻤㻜㻸㻲ᑠᆺᜏ᣺䛸䛖ᇵ㣴ᶵ㻌䠄䝞䜲䜸䝅䜵䞊䜹䞊䠅㻌䠄㻞ྎ䠅㼀㻭㻵㼀㻱㻯㻌䞉㻮㻾㻙㻞㻟㻲㻼ᑠᆺ㉸㐲ᚰᶵ

㻮㼑㼏㼗㼙㼍㼚㻌䞉㼀㻸㻙㻝㻜㻜ᑠᆺ㉸㐲ᚰᶵ

㻴㼕㼠㼍㼏㼔㼕㻌䞉㻴㼕㼙㼍㼏㻌㻯㻿㻝㻜㻜㉸㐲ᚰᶵ㻌䠄㻞ྎ䠅㻴㼕㼠㼍㼏㼔㼕㻌䞉㻡㻡㻼㻙㻣㻞㧗㏿෭༷㐲ᚰᶵ㻌䠄㻟ྎ䠅㼀㻻㻹㼅ᚤ㔞㧗㏿෭༷㐲ᚰᶵ㻌䠄㻡ྎ䠅

㼀㻻㻹㼅

㻼㻯㻾⿦⨨

㻭㻿㼀㻱㻯㻌䞉㻼㻯㻣㻜㻝㻼㻯㻾⿦⨨㻭㻿㼀㻱㻯㻌䞉㻼㻯㻣㻜㻤㻼㻯㻾⿦⨨㼀㼍㼗㼍㼞㼍㻌䞉㼀㼔㼑㼞㼙㼍㼘㻌㻯㼥㼏㼘㼍㼞㻌㻰㼕㼏㼑㻌㼀㼛㼡㼏㼔ᐃ㔞㻼㻯㻾⿦⨨㼀㼍㼗㼍㼞㼍㻌䞉㼀㼔㼑㼞㼙㼍㼘㻌㻯㼥㼏㼘㼍㼞㻌㻰㼕㼏㼑㻌㻾㼑㼍㼘㻌㼀㼕㼙㼑㻌㻿㼥㼟㼠㼑㼙㻌䊡䝟䝹䝇䝣䜱䞊䝹䝗㟁ẼὋື⿦⨨㻮㼕㼛㻙㻾㼍㼐㻌䞉㻼㼡㼘㼟㼣㼍㼢㼑㻣㻢㻜䝤䝻䝑䝔䜱䞁䜾⿦⨨㻌䠄Ὃືᘧ䞉䝉䝭䝗䝷䜲ᘧ䠅䠄㻞ྎ䠅㻭㼀㼀㻻㻌䞉㻭㻱㻙㻢㻢㻣㻜㼁㼂䜽䝻䝇䝸䞁䜹䞊㻲㼡㼚㼍㼗㼛㼟㼔㼕㻌䞉㻲㻿㻙㻤㻜㻜ศගගᗘィ㻿㻴㻵㻹㻭㻰㼆㼁㻌䞉㼁㼂㻙㻝㻢㻜ศගගᗘィ㻮㼑㼏㼗㼙㼍㼚㻌䞉㻰㼁㻙㻢㻡ᚤ㔞ศගගᗘィ㼀㼔㼑㼞㼙㼛㻌㻿㼏㼕㼑㼚㼠㼕㼒㼕㼏㻌䞉㻺㼍㼚㼛㻰㼞㼛㼜㻌㻞㻜㻜㻜䠄ỈᖹẼὶᆺ䠅㻌䜽䝸䞊䞁䝧䞁䝏㻌䠄㻞ྎ䠅㻿㻴㻻㼃㻭㻌䞉㻿㻙㻝㻟㻜㻜䠄ỈᖹẼὶᆺ䠅㻌䜽䝸䞊䞁䝧䞁䝏㻴㼕㼠㼍㼏㼔㼕㻌䞉㻼㻯㻴㻙㻝㻟㻜㻟㻮㻿ᐇయ㢧ᚤ㙾㻻㼘㼥㼙㼜㼡㼟㻌䞉㻿㼆㻢㻝㉸㡢Ἴ◚○⿦⨨㼀㻻㻹㼅㻌䞉㼁㻰㻙㻞㻜㻝኱ᆺ䜸䞊䝖䜽䝺䞊䝤㼀㻻㻹㼅㻌䞉㻲㻸㻿㻙㻝㻜㻜㻜୰ᆺ䜸䞊䝖䜽䝺䞊䝤㼀㻻㻹㼅㻌䞉㻸㻿㼄㻙㻡㻜㻜ᑠᆺ䜸䞊䝖䜽䝺䞊䝤㼀㻻㻹㼅㻌䞉㻮㻿㻙㻞㻠㻡ᑠᆺ䜸䞊䝖䜽䝺䞊䝤㻿㻭㻺㼅㻻㻌䞉㻹㻸㻿㻙㻞㻠㻜㻜஝⇕⁛⳦ᶵ㻌䠄㻟ྎ䠅㼅㼍㼙㼍㼠㼛㉸ప㻿㻭㻺㼅㻻෭෾ᗜ㻌䠄㻙㻤㻜䉝䠅䠄㻠ྎ䠅䝗䝷䝣䝖䝏䝱䞁䝞䞊㻰㻭㻸㼀㻻㻺㻌䞉㻰㻲㻙㻯䠄≉䠅ᰯṇ⏝ศ㖡ෆⶶᆺ㟁Ꮚኳ⛗㻭㻒㻰㻌䞉㻲㼆㻙㻞㻜㻜㻜㼕− 33 −

䚷䚷ᖹᡂ㻟㻜ᖺᗘ㻌㑇ఏᏊᐇ㦂ඹྠ฼⏝㒊㛛㻌䞉㻌ື≀ᰁⰍయ◊✲㒊㛛䚷ಖᏲႠ⧋䝸䝇䝖

ᖹᡂ㻟㻜ᖺ

ᖹᡂ㻟㻝ᖺ

㻠᭶

ᐃᮇΎᤲ㻔ẖ᭶䠅䝪䜲䝷䞊ᐊ⵨ẼᏳ඲ᘚྲྀ᭰

㻡᭶✵ㄪᶵჾ䝅䞊䝈䞁䜲䞁Ⅼ᳨㻢᭶㻝㝵㼃㻙㻝㻙㻜㻞ᐊ䝇䝔䞁䝺䝇ὶ䛧ྎ᤼Ỉワ䜎䜚ಟ⌮ᕤ஦㻝㝵㼃㻙㻞㻙㻜㻣ᐊὶ䛧ྎ᤼Ỉ⟶ಟ⌮㻣᭶㻝㝵㼃㻙㻝㻙㻜㻝ᐊ௚㟁Ẽタഛᕤ஦㻤᭶㻝㝵ዪᏊ౽ᡤᵝᘧ኱౽ჾ䝣䝷䝑䝅䝳䝞䝹䝤₃䜜ྲྀ᭰䛭䛾௚ಟ⌮ᕤ஦㻥᭶₃Ỉ⿵ಟ➼䛭䛾௚ᕤ஦㻝㻝᭶㻝㝵㼃㻙㻝㻙㻜㻞ᐊ㟁ᶵቑタᕤ஦㻝᭶⋞㛵㟁Ẽ㘄ྲྀ᭰ᕤ஦
✵ㄪᶵჾ䝅䞊䝈䞁䜸䝣Ⅼ᳨− 34 −

ᖹᡂ㻟㻜ᖺᗘ䚷䝀䝜䝮䝎䜲䝘䝭䜽䝇◊✲䝉䞁䝍䞊䚷฼⏝⪅◊✲ㄢ㢟䠄㑇ఏᏊᐇ㦂ඹྠ฼⏝㒊㛛䞉ື≀ᰁⰍయඹྠ฼⏝㒊㛛䠅◊㻌㻌✲㻌㻌ㄢ㻌㻌㢟㻌㻌ྡᡤ㻌㻌ᒓ㒊㻌㻌㛛㻌㻌䞉㻌㻌ศ㻌㻌㔝㻌㻌➼⫋㻌ྡ㻌᝟ሗ⛉Ꮫ◊✲⛉⏕࿨ே㛫᝟ሗ⛉Ꮫᑓᨷ䞉䝞䜲䜸䜲䞁䝣䜷䝬䝔䜱䜽䝇ㅮᗙᩍ㻌ᤵ㐲⸨㻌㻌ಇᚨ 㻌㻌䝠䝵䜴䝍䞁䛾ศᏊ⣔⤫ゎᯒ⌮Ꮫ◊✲㝔໬Ꮫ䞉᭷ᶵ䞉⏕࿨໬Ꮫᩍ㻌ᤵᆏཱྀ㻌㻌࿴㟹

⌮Ꮫ◊✲㝔⏕≀⛉Ꮫ䞉ᙧែᶵ⬟Ꮫᩍ㻌ᤵຍ⸨㻌㻌ᩔஅ 㻌㻌䝅䝻䜲䝚䝘䝈䝘䛻䛚䛡䜛ᙧែᙧᡂ㛵㐃㑇ఏᏊ䛾ゎᯒ⌮Ꮫ◊✲㝔⏕≀⛉Ꮫ䞉⾜ື⚄⤒⏕≀Ꮫ

ᩍ㻌ᤵ

ỈἼ䚷ㄔ

⌮Ꮫ◊✲㝔

⏕≀⛉Ꮫ䞉⾜ື⚄⤒⏕≀Ꮫ෸ᩍᤵ໭⏣㻌㻌୍༤ 㻌㻌᪂䛯䛺⑓ែ䝰䝕䝹ື≀䛾సฟ䛸ཎᅉ㑇ఏᏊ䛾ྠᐃ⌮Ꮫ◊✲㝔⏕≀⛉Ꮫ䞉⾜ື⚄⤒⏕≀Ꮫ෸ᩍᤵ࿴ከ䚷࿴ᏹ 䚷㬆⚺㢮䜢⏝䛔䛯ⓎኌᏛ⩦䞉⏕ᡂ䛸䛭䛾⬻ෆศᏊᶵᵓ⌮Ꮫ◊✲㝔⏕≀⛉Ꮫ䞉⏕ṪⓎ⏕⏕≀Ꮫᩍ㻌ᤵᒣୗ㻌㻌ṇව 㻌㻌⬨᳝ື≀䛻䛚䛡䜛⏕Ṫ⣽⬊ᙧᡂ䛾ไᚚᶵᵓ⌮Ꮫ◊✲㝔⏕≀⛉Ꮫ䞉⏕ṪⓎ⏕⏕≀Ꮫᩍ㻌ᤵ㯮ᒾ㻌㻌㯞㔛

XOᆺ䝖䝀䝛䝈䝭䛻䛚䛡䜛䝀䝜䝮ゎᯒ⌮Ꮫ◊✲㝔⏕≀⛉Ꮫ䞉⏕ṪⓎ⏕⏕≀Ꮫ

෸ᩍᤵ

ᮌᮧ㻌㻌ᩔ

㻌㻌့ங㢮䛾⏕Ṫ䛻䛛䛛䜟䜛䝀䝜䝮ᶵ⬟䛻㛵䛩䜛◊✲⌮Ꮫ◊✲㝔⏕≀⛉Ꮫ䞉⏕ṪⓎ⏕⏕≀Ꮫຓ㻌ᩍྜྷ⏣㻌㻌㑳ஓ

့ங㢮୙άᛶXᰁⰍయ䛾෌άᛶ໬ᶵᵓ䛾ゎᯒ⌮Ꮫ◊✲㝔⏕≀⛉Ꮫ䞉ከᵝᛶ⏕≀Ꮫᩍ㻌ᤵቑ⏣㻌㻌㝯୍ 㻌㻌့ங㢮䛚䜘䜃㫽㢮䛾ศᏊ㐍໬䛸㑇ఏⓗከᵝᛶ䛻㛵䛩䜛◊✲⌮Ꮫ◊✲㝔⏕≀⛉Ꮫ䞉ከᵝᛶ⏕≀Ꮫᩍ㻌ᤵᑠட䚷୍ᘯ 䚷⸴㢮䛾⣔⤫㐍໬Ꮫඛ➃⏕࿨⛉Ꮫ ⏕࿨ᶵ⬟⛉Ꮫ◊✲䞉⏕≀᝟ሗゎᯒ⛉Ꮫ◊✲
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Ụ⏣㻌㻌┿Ẏ 㻌㻌⌧⏕䛚䜘䜃㑇㊧ฟᅵ䛾ື≀㉁㈨ᩱ䛾ሷᇶ㓄ิ䛾Ỵᐃ◊✲㒊䞉㈨ᩱ㛤Ⓨ◊✲⣔Ặ㻌㻌ྡᖹᡂ㻟㻜ᖺᗘ䚷฼⏝≧ἣᡤ㻌㻌ᒓ㻌㻌

฼㻌⏝㻌⪅㻌ᩘ኱Ꮫ㝔᝟ሗ⛉Ꮫ◊✲⛉㻝኱Ꮫ㝔⌮Ꮫ◊✲㝔㻝㻝኱Ꮫ㝔ඛ➃⏕࿨⛉Ꮫ◊✲㝔

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− 35 −⒴ᢚไ䝍䞁䝟䜽㉁p53䛚䜘䜃㛵㐃ᅉᏊ䛾ᵓ㐀ᶵ⬟┦㛵㻌㻌䝣䝍䝩䝅䝁䜸䝻䜼䛾Ꮫ⩦䛚䜘䜃ᛶỴᐃ䛾◊✲

㻌㻌⺮ⓑ㉁䛾❧యᵓ㐀ゎᯒ䛻㛵䛩䜛◊✲Мဇ‫إ‬ԓᴾᴾ኱Ꮫ㝔᝟᝟ሗ⛉Ꮫ◊✲⛉⏕࿨ே㛫᝟ሗ⛉Ꮫᑓᨷࣂ࢖࢜࢖ࣥࣇ࢛࣐ࢸ࢕ࢡࢫㅮᗙ
㐲⸨ ಇᚨ
㢟ྡ㸸ࣄࣙ࢘ࢱࣥ Lagenaria siceraria ࡢศᏊ⣔⤫ゎᯒ࡜ึᮇఏ᧛⤒㊰ࡢゎ᫂ ࣄࣙ࢘ࢱࣥࡢ㉳※࡜ఏ᧛⤒㊰ゎ᫂ࡢࡓࡵࡢศᏊ
⣔⤫ゎᯒࢆ๓ᖺ࠿ࡽ⥅⥆ࡋ࡚⾜ࡗࡓࠋࣄࣙ࢘ࢱࣥ
ࡣ✀Ꮚᙧែ࠿ࡽ኱ࡁࡃ࢔ࣇࣜ࢝ᆺ࡜࢔ࢪ࢔ᆺ࡟ศ
㢮ࡉࢀࠊ㉳※ࡣ࢔ࣇࣜ࢝࡜⪃࠼ࡽࢀ࡚࠸ࡿࡀࠊ✀Ꮚᙧែࡣከᵝᛶࡀ㧗ࡃࠊ࢔࣓ࣜ࢝኱㝣ࡢఏ⤫ⓗ᱂ᇵ✀ࡀ࡝ࡕࡽ࡟ᒓࡍࡿ࠿ࡀㄽத࡟࡞ࡗ࡚࠸ࡿࠋ࢔
ࢪ࢔࠿ࡽࡢே㢮⛣ఫ᫬ᣢ㎸ㄝ࡜࢔ࣇࣜ࢝኱㝣࠿ࡽࡢ⮬↛⁻╔ㄝࡀ࠶ࡿࠋᚋ⪅ࡣⴥ⥳యࢤࣀ࣒ゎᯒ࠿ࡽᨭᣢࡉࢀ࡚࠸ࡿ㸦Kistler et al. 2014㸧ࡀࠊ㓄ิࡢ≉ᚩ࡜ศᒱᖺ௦࡟୙⮬↛࡞Ⅼࡀ࠶ࡿࡓࡵ᳨ドࢆ㐍ࡵࡓࠋᮏᖺᗘࡣࠊ๓ᖺ࡟ NGS ゎᯒࢆ⾜ࡗࡓ᱂ᇵᰴ 3 ✀㢮㸦࢔ࢪ࢔ᆺ (Laos #S1)ࠊ࢔ࣇࣜ࢝ᆺ (Namibia#348)ࠊ࢔࣓ࣜ࢝ (Mexico #600)㸧࠾ࡼࡧࠊ㔝⏕㏆
⦕✀ L. sphaerica (#362), L. abyssinica (#451), L.
breviflora (#381)ࡢࣂ࢖࢜࢖ࣥࣇ࢛࣐ࢸ࢕ࢡࢫゎᯒࢆ㐍ࡵࠊ3 ኱㝣ࡢ᱂ᇵᰴ࡜࢔ࣇࣜ࢝ࡢ㔝⏕✀ 3 ✀ࡢ㑇ఏᏊࢫ࣌ࢡࢺࣝẚ㍑⤖ᯝࢆᚓࡓ㸦ᅗ 1㸧ࠋࡇࡢ⤖ᯝࠊ766 ࡢ㔝⏕✀ᅛ᭷㑇ఏᏊ࡜ 401 ࡢ᱂ᇵ✀ᅛ᭷㑇ఏᏊࡀぢࡘ࠿ࡗࡓࠋࡇࡢ࡯࠿࢔ࢪ࢔ရ✀࡟ᅛ᭷ࡢ㓄ิࡀ 299 ಶྠᐃࡉࢀࡓࠋ⊂⮬ࡢ㑇ఏᏊᑟධࡀ࠶ࡗࡓྍ⬟ᛶࡀ࠶ࡿࠋ

ᅗ 1 ࡢゎᯒ࡛ࡣࠊSwissProt ࢹ࣮ࢱ࡜ࡢ┦ྠᛶࣞ࣋ࣝ 90㸣௨ୖࡢ㓄ิ࡟ࡘ࠸࡚ẚ㍑ࢆ⾜ࡗࡓࡓࡵࠊཧ
↷㓄ิࡀື≀ࡢ㑇ఏᏊࡢሙྜ࡞࡝ࡣゎᯒ࠿ࡽእࢀ
࡚ࡋࡲ࠺ሙྜࡀከ࠸ࠋࡇࡢࡓࡵ┦ྠᛶࣞ࣋ࣝࢆ
70㸣ࡲ࡛ୗࡆ࡚ྠᵝࡢゎᯒࢆ⾜ࡗࡓ࡜ࡇࢁࠊඹ᭷
㑇ఏᏊ 8531ࠊ㔝⏕✀ᅛ᭷ࡀ 1572ࠊ᱂ᇵ✀ᅛ᭷ࡀ
1238 ࡜࡞ࡗࡓ࡯࠿ࠊ࢔ࢪ࢔ᰴᅛ᭷ࡢ㓄ิࡣ࡯ࡰ300 ࡢࡲࡲ࡜࡞ࡗࡓࠋ㑇ఏᏊ㔜」ศࡀ⦰㔜ࡋ࡚࠸ࡿࡇ࡜ࢆ⪃࠼ࡿ࡜ࠊࡇࡢ⤖ᯝࡣࣄࣙ࢘ࢱࣥࡢࢤࣀ
࣒ࡢᐇ㝿ࢆ཯ᫎࡋࡓࡶࡢ࡜⪃࠼ࡽࢀࡿࠋࡇࢀࡽࡢ
⤖ᯝ࠿ࡽࠊ᱂ᇵ✀ࡢࣄࣙ࢘ࢱࣥࡣ㔝⏕✀ࢤࣀ࣒ࡢ
10㸣⛬ᗘࡢ㑇ఏᏊࢆኻ࠸ࠊྠ⛬ᗘࡢ᱂ᇵ✀ᅛ᭷㑇ఏᏊࢆᚓࡓ࡜⪃࠼ࡽࢀࡿࠋࡲࡓࠊ࢔ࢪ࢔ᰴࡣ⊂⮬ࡢᅛ᭷㓄ิ 300 ⛬ᗘࢆ⋓ᚓࡋ࡚࠸ࡿྍ⬟ᛶࡀ♧၀ࡉࢀࡓࠋ௒ᚋࠊࡼࡾ㧗⢭ᗘࡢࢤࣀ࣒㓄ิࢆ⏝࠸᳨࡚ドࡋࡓᚋࠊ⤖ᯝࢆㄽᩥ໬ࡍࡿணᐃ࡛࠶ࡿࠋᏛ఍ⓎⓎ⾲➼Dai Watabe, Hiroshi Yuasa, Naoki Osada, Kazuhiro
Satomura, and Toshinori Endo
Hybrid type of nuclear and plastid DNAs suggests the
hypothesis for ancient propagation of American bottle
gourd (Lagenaria siceraria), Society of Molecular
Biology and Evolution, Yokohama, 2018.
㐲⸨ ಇᚨ ྂ௦⎭⟸ࡢ㌶㊧ࢆ㎺ࡿ ➨ 72 ᅇ᪥ᮏே㢮Ꮫ఍኱఍ බ㛤ࢩ࣏ࣥࢪ࣒࢘, ୕ᓥ, 2018

ᅗ 1 ᱂ᇵ✀ 3 ᰴ࡜ L. sphaerica ࡢ㑇ఏᏊࢫ࣌ࢡࢺ
ࣝẚ㍑㸦ཧ↷㓄ิ 90㸣୍⮴ࣞ࣋ࣝ㸧
SwissProt ࢹ࣮ࢱ࡬ࡢ┦ྠᛶ 90㸣ࡢ㓄ิ࡟ࡘ࠸࡚
ࡢẚ㍑ࠋ
− 36 −

኱Ꮫ㝔⌮Ꮫ◊✲㝔໬Ꮫ㒊㒊㛛 ᭷ᶵ࣭⏕࿨໬Ꮫศ㔝ᆏཱྀ ࿴㟹㸦ᶵჾ฼⏝㸧
‫ۑ‬ㄽᩥⓎ⾲ 
Ogasawara S., Chuman Y., Michiba T., Kamada R.,
Imagawa T. and Sakaguchi K. (2018)
Inhibition of protein phosphatase PPM1D enhances
retinoic acid-induced differentiation in human embryoniccarcinoma cell line. J. Biochem., DOI 10.1093/jb/mvy119.
‫ۑ‬Ꮫ఍Ⓨ⾲௒ᕝᩄ᫂ࠊ㭯ᒸᶞࠊ୰ᒣ⤮⨾㔛ࠊᡞཱྀౡࠊ㙊⏣⍠Ἠࠊᆏཱྀ࿴㟹㸸ࢩࣥࢢࣝࢭࣝゎᯒ࡟ࡼࡿ⣽⬊࿘ᮇ࡜ࡀࢇᢚไࢱࣥࣃࢡ㉁ p53 ౫Ꮡⓗ㌿෗άᛶ࡜ࡢ┦㛵.
➨ 91 ᅇ᪥ᮏ⏕໬Ꮫ఍኱఍, 2018 ᖺ 9 ᭶ 24 ᪥, ᅜ❧ி㒔ᅜ㝿఍㤋㸦ி㒔ᗓி㒔ᕷ㸧㭯ᒸᶞࠊ୰ᒣ⤮⨾㔛ࠊ㙊⏣⍠Ἠࠊ௒ᕝᩄ᫂ࠊᆏཱྀ࿴
㟹㸸ࢩࣥࢢࣝࢭࣝゎᯒ࡟ࡼࡿ⒴ᢚไࢱࣥࣃࢡ㉁ p53
ࢆ௓ࡋࡓ⣽⬊ࢫࢺࣞࢫᛂ⟅ከᵝᛶⓎ⌧ᶵᵓゎ᫂.
➨ 50 ᅇⱝᡭ࣌ࣉࢳࢻኟࡢຮᙉ఍, 2018 ᖺ 8 ᭶ 6᪥, ⮫῭᐀᪉ᗈᑎὴ኱ᮏᒣ᪉ᗈᑎ㸦㟼ᒸ┴὾ᯇᕷ㸧
Tatsuki Tsuruoka, Rui Kamada, Toshiaki Imagawa andKazuyasu Sakaguchi㸸Heterogeneity in the cellular stressresponse mediated by tumor suppressor protein p53
using single cell analysis.
The 37th Sapporo International Cancer Symposium,
2018 ᖺ 7 ᭶ 18 ᪥, ࣟ࢖ࢺࣥᮐᖠ㸦໭ᾏ㐨ᮐᖠᕷ㸧㭯ᒸᶞࠊ୰ᒣ⤮⨾㔛ࠊ㙊⏣⍠Ἠࠊ௒ᕝᩄ᫂ࠊᆏཱྀ
࿴㟹㸸ࢩࣥࢢࣝࢭࣝゎᯒ࡟ࡼࡿ⒴ᢚไࢱࣥࣃࢡ㉁p53 ࢆ௓ࡋࡓ⣽⬊ࢫࢺࣞࢫᛂ⟅ከᵝᛶゎᯒ.
᪥ᮏ⏕໬Ꮫ఍➨ 55 ᅇ໭ᾏ㐨ᨭ㒊౛఍, 2018 ᖺ 7 ᭶
13 ᪥, 㓗㎰Ꮫᅬ኱Ꮫ㸦໭ᾏ㐨Ụูᕷ㸧⏕≀⛉Ꮫ㒊㒊㛛 ⏕ṪⓎ⏕⏕≀Ꮫศศ㔝ྜྷ⏣ 㑳ஓ ᮏᖺᗘࡣࠊື≀⣽⬊ᇵ㣴ᐊࠊཬࡧࠊὙίᐊࢆ౑
⏝ࡉࡏ࡚㡬ࡁࠊ௨ୗࡢ◊✲ࢆ⾜ࡗࡓࠋ౑⏝ࡋࡓඹ⏝ᶵჾࡣ3&5⿦⨨ࠊศගගᗘィࠊᚤ㔞㐲ᚰᶵࠊᜏᵴࠊࢤࣝ᧜ᙳ⿦⨨ࠊ෗┿᧜ᙳ⿦⨨ࠊ(]&DSWXUH࡞࡝࡛࠶ࡿ㸸 ㌿෗ᅉᏊ(VUUE࡟ࡼࡿ(&⣽⬊㞟ᅋ࡛ࡢᖿ⣽⬊ᵝ๓㥑⣽⬊ࡢฟ⌧࡜㠀ᑐ⛠ศ⿣ࡢไᚚ㸸0&⣽⬊㞟ᅋ୰࡟☜⋡ⓗ࡟ฟ⌧ࡍࡿᖿ⣽⬊ᵝ๓㥑⣽⬊㸦7*⪏ᛶ㸧ࡀ㠀ᑐ⛠ศ⿣࡟ࡼࡗ࡚+$7⪏ᛶ⣽⬊࡜๓㥑⣽⬊⮬㌟ࢆ⏕ࡳฟࡍ⌧㇟࡟㛵ࡋ࡚ࠊ๓ᖺ
ᗘࡲ࡛ࡢ◊✲⤖ᯝ࠿ࡽ๓㥑⣽⬊ࡢฟ⌧࡟ࡣ(VUUEࡢ㐺ษ࡞Ⓨ⌧㔞ࡀᚲせ࡛࠶ࡿྍ⬟ᛶࡀ♧ࡉࢀ࡚࠸ࡓࠋ௒ᖺᗘࡣࠊ๓ᖺᗘࡢ⤖ᯝ࡟ຍ࠼࡚ࠊ㐣๫࡟Ⓨ⌧ࡋࡓ(655%ࡣ+$7⪏ᛶ⣽⬊ࡢฟ⌧ࢆጉࡆࠊ⤖ᯝ࡜ࡋ࡚ᑐ⛠ศ⿣࡟ࡼࡗ࡚㸰ಶࡢ๓㥑⣽⬊ࡀ⏕ࡳฟࡉࢀ࡚࠸ࡿࡇ࡜ࢆࠊ'(6ฎ⌮ࡋࡓ(655%㧗Ⓨ⌧ᰴH
H࡛᫂ࡽ࠿࡟ࡋࡓࠋ'(6ฎ⌮࡟ࡼࡗ࡚+$7⪏ᛶࢥࣟࢽ࣮ࡢฟ⌧⋡㸦㸻๓㥑⣽⬊ࡢฟ⌧⋡㸧ࡀⴭࡋࡃୖ
᪼ࡍࡿࡇ࡜ࢆ)OXFWXDWLRQWHVWࢆ⏝࠸࡚ヲ⣽࡟ゎᯒࡋࡓࠋ(655%Ⓨ⌧㔞ࡀ㠀ᖖ࡟ከ࠸(6⣽⬊ࡸึᮇ⬇
ࡢ,&0࡛ࡣᖿ⣽⬊ࡀᑐ⛠ศ⿣ࢆ࠾ࡇ࡞࠺ࡀࠊࡇࡢᑐ
⛠ศ⿣࡟ࡣ(VUUEࡀ㛵୚ࡋ࡚࠸ࡿࡇ࡜ࢆ♧ࡍ⤖ᯝ࡜࡞ࡗࡓࠋ,&0࡛ࡣࡇࡢࡼ࠺࡞ᖿ⣽⬊ࡢᑐ⛠ศ⿣ࡣ୍㐣ⓗ࡞≧ែ࡛࠶ࡾࠊࡸࡀ࡚㠀ᑐ⛠ศ⿣ࢆ௓ࡋ࡚
୕⬇ⴥ࡬ࡢศ໬ࡀ㉳ࡁࡿࠋึᮇ⬇࡟࠾ࡅࡿᖿ⣽⬊ࡢᑐ⛠ศ⿣࠿ࡽ㠀ᑐ⛠ศ⿣࡬ࡢኚ໬ࡣࠊ(VUUEࡢⓎ⌧㔞࡟ࡼࡗ࡚ไᚚࡉࢀ࡚࠸ࡿྍ⬟ᛶࡀ௒ᅇࡢᐇ㦂࠿ࡽ♧၀ࡉࢀࡓࠋࡲࡓࠊ(VUUEࡢⓎ⌧㛤ጞ࡟㛵ࢃࡿș࢝ࢸࢽࣥࡢ᰾⛣⾜࡟ࡘ࠸࡚ࡣࠊᚑ᮶⪃࠼࡚࠸ࡓࡼ࠺࡞⣽⬊㉁࡛ࡢ኱ᖜ࡞㔞ⓗኚ໬ࡣᚲせࡀ↓ࡃࠊࢃࡎ࠿࡞㔞ࡢኚື࡟ࡼࡗ࡚㉳ࡁᚓࡿࡇ࡜ࡶ᫂ࡽ࠿
࡟࡞ࡗ࡚ࡁࡓࠋș࢝ࢸࢽࣥࡢ⣽⬊㉁ෆ࡛ࡢ┦ᑐⓗ࡞⃰ᗘࢆኚ࠼ࡿせᅉ࡟ࡘ࠸࡚ࡣࡉࡽ࡟ゎᯒࢆ⥆ࡅࡿᚲせࡀ࠶ࡿࠋࡉࡽ࡟୍㐃ࡢࢫ࢖ࢵࢳࡀാ࠸࡚
(655%ࡢⓎ⌧ࡀ㛤ጞࡋࡓᚋ࡟ࠊ๓㥑⣽⬊ࡀฟ⌧ࡍࡿ࡟ࡣᩘୡ௦௨ୖࡢ⣽⬊ศ⿣ࡀᚲせ࡛࠶ࡿࡇ࡜ࡶ᫂
ࡽ࠿࡟࡞ࡗ࡚ࡁࡓࠋࡇࡢᩘୡ௦ࡢศ⿣୰࡟ఱࡀ㉳ࡁ࡚๓㥑⣽⬊ࡢฟ⌧࡟⮳ࡿࡢ࠿࡜࠸࠺ၥ㢟ࡶࠊ௒ᚋࡢ㔜せ࡞ㄢ㢟࡛࠶ࡿࠋ

⏕≀⛉Ꮫ㒊㒊㛛 ከᵝᛶ⏕≀Ꮫศศ㔝ቑ⏣ 㝯୍◊✲ࢢ࣮ࣝࣉ㸸⌮Ꮫ◊✲㝔⏕≀⛉Ꮫ㒊㛛ࠊࢤࣀ࣒ࢲ࢖ࢼ࣑ࢡࢫ◊✲ࢭࣥࢱ࣮࣭ቑ⏣㝯୍⌮Ꮫ◊✲㝔⏕≀⛉Ꮫ㒊㛛ࠊࢤࣀ࣒ࢲ࢖ࢼ࣑ࢡࢫ◊✲ࢭࣥࢱ࣮࣭す⏣⩏᠇ᖹᡂ 30 ᖺᗘ࡟ࡣ୺࡟௨ୗࡢᡂᯝࡀᚓࡽࢀࡓࠋ
(1) ࣮ࣘࣛࢩ࢔኱㝣࡟࠾ࡅࡿࢩ࣋ࣜ࢔࢖ࢱࢳࡢᆅ⌮ⓗኚ␗ ᮏ◊✲࡛ࡣࠊࣟࢩ࢔࡜ࡢඹྠ◊✲࡜ࡋ࡚ࠊ࣮ࣘ
ࣛࢩ࢔ࡢᗈ⠊ᅖ࡟⏕ᜥࡍࡿࢩ࣋ࣜ࢔࢖ࢱࢳ࡟ࡘ࠸࡚ࠊヲ⣽࡞⣔⤫ᆅ⌮ⓗゎᯒࢆ⾜ࡗࡓࠋᮾ࢔ࢪ࢔࠾
ࡼࡧࣟࢩ࢔ࡢᵝࠎ࡞ሙᡤ࡛᥇㞟ࡉࢀࡓಶయࢆ୰ᚰ
࡜ࡋࠊྂ࠸ẟ⓶ࢆྵࡵࡓከᩘࡢࢧࣥࣉࣝ࠿ࡽ DNA
ࢆᢳฟࡋࠊ࣑ࢺࢥࣥࢻࣜ࢔ DNA ࡢࢥࣥࢺ࣮ࣟࣝ㡿ᇦ㸦D-loop 㡿ᇦ㸧ࡢሷᇶࢆỴᐃࡋࠊศᏊ⣔⤫ゎᯒࢆ⾜ࡗࡓࠋࡑࡢ⤖ᯝࠊmtDNA ⣔⤫ࡣ኱ࡁࡃ㸱ࡘࡢࢢ࣮ࣝࣉ࡟ศࡅࡽࢀࡓࠋࡉࡽ࡟ࠊ⣔⤫࡟ࡼࡾከᵝᛶࡢ㐪࠸࠾ࡼࡧศᕸᣑᩓࡢ㏿ᗘࡀ␗࡞ࡿࡇ࡜ࡀ᥎

− 37 −

ᐃࡉࢀࡓ㸦Ꮫ఍Ⓨ⾲㸸▼ᕝ࡯࠿㸦2018㸧㸧ࠋ(2) ∗⣔㑇ఏᏊࡢከᵝᛶ࠿ࡽࡳࡓ࢔࢝ࢠࢶࢿࡢ⣔⤫ᆅ⌮ Y ᰁⰍయୖࡢ 2 ࡘࡢ࣐࢖ࢡࣟࢧࢸࣛ࢖ࢺ㑇ఏᏊᗙࡢከᆺᛶ࡟ᇶ࡙ࡁࠊ࣮ࣘࣛࢩ࢔ࡢᗈᇦ࡟ศᕸࡍ
ࡿ࢔࢝ࢠࢶࢿࡢ⣔⤫ᆅ⌮ゎᯒࢆ⾜ࡗࡓࠋ࣐࢖ࢡࣟ
ࢧࢸࣛ࢖ࢺࣔࢳ࣮ࣇࡢ⧞ࡾ㏉ࡋᅇᩘ࡜ሷᇶ㓄ิ࡟ᇶ࡙ࡁࣁࣉࣟࢱ࢖ࣉࢆỴᐃࡋࡓ⤖ᯝࠊ୺࡟ 3 ࡘࡢ⣔⤫ࢢ࣮ࣝࣉ࡟ศࡅࡽࢀࡓࠋྛ⣔⤫ࡢᆅ⌮ⓗศᕸ࡟ࡣ኱㝣ࣞ࣋ࣝ࠾ࡼࡧ᪥ᮏิᓥࡢᓥ㛫࡛≉ᚩࡀぢࡽࢀࡓࠋࡇࡢࢹ࣮ࢱ࡜ࠊᚑ᮶ࡢẕ⣔㑇ఏᏊ㸦mtDNA㸧ࢹ࣮ࢱࢆྜࢃࡏࡿࡇ࡜࡟ࡼࡾࠊ࢔࢝ࢠࢶࢿࡢື≀ᆅ⌮ᏛⓗṔྐࢆࡼࡾヲ⣽࡟⪃ᐹࡍࡿࡇ࡜ࡀ࡛ࡁࡿ
ࡇ࡜ࡀุ᫂ࡋࡓ㸦Ꮫ఍Ⓨ⾲㸸⧊⏣࡯࠿㸦2018㸧㸧ࠋ(3) ໭ᾏ㐨ࡢ࢟ࢱ࢟ࢶࢿ࡟࠾ࡅࡿ୺せ⤌⧊㐺ྜ㑇ఏᏊ」ྜయ㸦MHC㸧ࡢከᵝᛶ ࢟ࢱ࢟ࢶࢿ㞟ᅋ࡟ࡘ࠸࡚ࠊMHCࢡࣛࢫII DRB㑇ఏᏊࡢ㞟ᅋ㑇ఏᏛⓗゎᯒࢆ⾜ࡗࡓࠋࡑࡢ⤖ᯝࠊ㐨༡㞟ᅋࡣ௚ࡢᆅᇦ㞟ᅋ࠿ࡽ㑇ఏⓗ࡟ศ໬ࡋ࡚࠸ࡿ
ࡇ࡜ࠊ㑇ఏⓗከᵝᛶࡀపୗࡋ࡚࠸ࡿࡇ࡜ࡀㄆࡵࡽ
ࢀࡓࠋࡉࡽ࡟ࠊDRB㑇ఏᏊࡀṇࡢ㑅ᢥࢆཷࡅ࡚㐍໬ࡍࡿ࡜࡜ࡶ࡟ࠊᖹ⾮㐍໬࡟ࡼࡿ✀ࢆ㉸࠼ࡓከᆺ㸦trans-species polymorphism㸧ࢆ♧ࡍࡇ࡜࡞࡝ࡶ᫂
ࡽ࠿࡜࡞ࡗࡓ㸦Amaike et al. 2018㸧ࠋ(4) ࣐ࢲࣛ࢖ࢱࢳࡢ୺せ⤌⧊㐺ྜ㑇ఏᏊ」ྜయ
㸦MHC㸧㑇ఏᏊࡢከᵝᛶ ࣐ࢲࣛ࢖ࢱࢳࡣ༡࣮ࣚࣟࢵࣃ࠿ࡽ୰ᅜす㒊࡟࠿ࡅ࡚ศᕸࡍࡿ࢖ࢱࢳ⛉࡟ᒓࡍࡿᕼᑡ✀࡛ࠊIUCN ࡢ
Red List ࡛ࡣ VU㸦༴ᛴ㸧࡟ศ㢮ࡉࢀࡿࠋᮏ✀ࡢ㐺ᛂ㐍໬࡟㛵ࡍࡿᇶ♏ⓗ▱ぢ࡞࡝ࢆᚓࡿࡓࡵࠊච␿⣔࡛㔜せ࡞ᶵ⬟㑇ఏᏊࡢ୍ࡘ࡛࠶ࡾࠊMHC ࢡࣛࢫ II ࡟ᒓࡍࡿ DRB 㑇ఏᏊࡢሷᇶ㓄ิከᵝᛶࢆᣦᶆ࡜ࡋࡓศᏊ㐍໬Ꮫⓗゎᯒࢆ⾜ࡗࡓࠋࣈࣝ࢞ࣜ࢔㞟ᅋࡢ 10 ಶయࢆヨᩱ࡜ࡋ࡚ᚓࡿࡇ࡜ࡀ࡛ࡁࠊࡇࢀࡽ
ࡼࡾ 9 ✀㢮ࡢᑐ❧㑇ఏᏊࡀ᳨ฟࡉࢀࡓࠋᗈᇦศᕸ
ࡍࡿࡶࡢ࡜≉ᐃᆅᇦ࡛ࡢࡳぢࡽࢀࡿࡶࡢࡀᏑᅾࡋ
ࡓࡇ࡜࠿ࡽࠊࡇࡢ✀ࡢ DRB 㑇ఏᏊࡀᢠཎࡢᙳ㡪ୗ࡛ከᩘࡢᑐ❧㑇ఏᏊࡀ㞟ᅋ୰࡛⥔ᣢࡉࢀࡿ
pathogen-driven balancing selection ࢆཷࡅ࡚㐍໬ࡋ࡚ࡁࡓࡇ࡜ࡀ♧ࡉࢀࡓࠋࡲࡓࠊᚓࡽࢀࡓሷᇶ㓄ิࢹ࣮ࢱ࡟ᇶ࡙ࡃ⤫ィⓗゎᯒ࠿ࡽࠊࡇࡢ㑇ఏᏊࡢ㐍໬࡟ࡣṇࡢ⮬↛㑅ᢥࡸ㑇ఏᏊ⤌᥮࠼⌧㇟ࡶ㛵୚ࡋ࡚ࡁࡓ࡜⪃࠼ࡽࢀࡓࠋࡉࡽ࡟࣋࢖ࢬἲ࡟ࡼࡿศᏊ
⣔⤫ゎᯒ࡟ࡼࡾࠊ࢖ࢱࢳ⛉࡟ᒓࡍࡿ௚✀ࡢ DRB ᑐ❧ 㑇 ఏ Ꮚ ࡜ ࡢ ✀ ࢆ ㉸ ࠼ ࡓ ከ ᆺ 㸦 trans-species
polymorphism㸧ࡀぢ࠸ࡔࡉࢀࠊᮏ✀ࡢ㐍໬ྐࡢ୍ࠋ➃ࡶ♧ࡉࢀࡓ㸦Nishita et al. 2018㸧
Ⓨ⾲ㄽᩥ
1. Nishita, Y., Abramov, A.V., Murakami, T., andMasuda, R. (2018) Genetic diversity of MHC

2.

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Low genetic diversity in an isolated red fox(Vulpes vulpes) population on Mt. Hakodate,Japan, revealed by microsatellite analyses of fecal
samples. Mammal Study 43: 141-152.
Amaike, Y., Nishita, Y., Uraguchi, K., and Masuda, R.
(2018b) Genetic diversity of MHC class II DRB1
exon 2 in the red fox (Vulpes vulpes) on Hokkaido,
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Saka, T., Nishita, Y., and Masuda, R. (2018) Low
genetic variation in the MHC class II DRB geneand MHC-linked microsatellites in endangeredisland populations of the leopard cat (Prionailurus
bengalensis) in Japan. Immunogenetics 70: 115-124.
Omote, K., Surmach, S.G., Kohyama, T.I., Takenaka,T., Nishida, C., and Masuda, R. (2018) Phylogeographyof continental and island populations of Blakiston’s
fish-owl (Bubo blakistoni) in northeastern Asia.
Journal of Raptor Research 52: 31-41.
Abramov, A.V., Puzachenko, A.Yu., and Masuda, R.
(2018) Cranial variation in the Siberian weasel
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ቑ⏣㝯୍ (2019) ့ங㢮ࡢ⣔⤫ᆅ⌮ࢆ᥈ࡿࠥࣄࢢ࣐ࡢ⛣ືྐ㸬⏕≀⛉Ꮫ 70: 25-30㸬ቑ⏣㝯୍ (2018) 㑇ఏᏊࡀゎࡁ᫂࠿ࡍ໭ᾏ㐨ࡢ⏕≀ᆅ⌮㸬ࣅ࢜ࣄࢫࢺ࣮ࣜ 30 ྕ: 12-15㸬ቑ⏣㝯୍ (2018) ࢩ࣐ࣇࢡࣟ࢘ࡢ㑇ఏⓗከᵝᛶ࡜ᮍ᮶࡬ࡢᒎᮃ㸬㔝㫽 8 ᭶ྕ㸦827 ྕ㸧: 6-9㸬ⴭ᭩
1. ቑ⏣㝯୍㸦2019㸧࣮ࣘࣛࢩ࢔ື≀⣖⾜, ᒾἼ᪂᭩㸬
2. ቑ⏣㝯୍ ⦅ⴭ㸦2018㸧᪥ᮏࡢ㣗⫗㢮㸸⏕ែ⣔ࡢ㡬Ⅼ࡟❧ࡘ့ங㢮㸪ᮾி኱Ꮫฟ∧఍㸬
3. Masuda R. (2018) Status and perspective of the
population based on genetic diversity: Introduction.
In “Biodiversity Conservation Using Umbrella
Species. Ecological Research Monographs (F.
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4. Omote K., Kohyama T.I., Nishida C., Takenaka T.,
Surmach S.G., and Masuda R. (2018) Status andperspective of the Blakiston’s fish owl population,
based on genetic diversity. In “Biodiversity
Conservation Using Umbrella Species. EcologicalResearch Monographs (F. Nakamura, eds)”, pp.
135-141, Springer, Singapore.
5. Akiyama T., Kohyama T.I., Nishida C., and Masuda
R. (2018) Status and perspective of the red-crowned
crane based on genetic diversity. In “Biodiversity
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Research Monographs (F. Nakamura, eds)”, pp.
143-148, Springer, Singapore.
Ꮫ⾡ㅮ₇࣭Ꮫ఍Ⓨ⾲
1. ቑ⏣㝯୍㸸“㑇ఏᏊ࠿ࡽ᥈ࡿື≀ࡢ࣮ࣝࢶ࡜ከ
ᵝᛶ”㸪➨56ᅇ໭ᾏ㐨㧗➼Ꮫᰯᩍ⫱◊✲኱఍࣭⌮⛉㒊఍࣭⏕≀ศ⛉఍ㅮ₇㸪2019ᖺ1᭶10᪥㸪ᮐᖠᖹᓊ㧗ᰯ㸦໭ᾏ㐨ᮐᖠᕷ㸧㸬
2. ▼ᕝᜨኴ㸪Alexei V. Abramov㸪す⏣⩏᠇㸪ቑ⏣㝯୍㸸Phylogeography of the Siberian weasel(Mustela sibirica), based on mitochondrial DNA
analysis㸦࣑ࢺࢥࣥࢻࣜ࢔DNAゎᯒ࡟ࡼࡿࢩ࣋
ࣜ࢔࢖ࢱࢳ(Mustela sibirica)ࡢ⣔⤫ᆅ⌮࡟ࡘ࠸࡚㸧㸬᪥ᮏື≀Ꮫ఍➨89ᅇ኱఍㸪2018ᖺ12᭶9᪥㸪ᮾி኱Ꮫ㸦ᮾி㒔㸧㸬ࠕ࣏ࢫࢱ࣮ࠖ
3. ⧊⏣ ᮍᕼ, Alexei V. Abramov, Evgeniy Raichev,
Pavel A. Kosintsev, Alexey P. Kryukov, ᾆཱྀᏹ஧, す⏣⩏᠇, ቑ⏣㝯୍㸸YᰁⰍయୖࡢ࣐࢖ࢡ
ࣟࢧࢸࣛ࢖ࢺࡢከᵝᛶ࠿ࡽࡳࡓ࢔࢝ࢠࢶࢿ
(Vulpes vulpes) ࡢ⣔⤫ᆅ⌮㸬᪥ᮏື≀Ꮫ఍➨89ᅇ኱఍㸪2018ᖺ12᭶9᪥㸪ᮾி኱Ꮫ㸦ᮾி㒔㸧㸬ࠕ࣏ࢫࢱ࣮ࠖ
4. Nishita, Y.: Genetic variation of the MHC class II
DRB genes among three weasel species: Japanese
weasel (Mustela itatsi) and Siberian weasel (M.
sibirica) and least weasel (M. nivalis), Seminar at
University of Helsinki, February 25, 2019, Finland.

኱Ꮫ㝔ඛඛ➃⏕࿨⛉Ꮫ◊✲㝔

Shimono, K., Mizuno, M., Kojima, K., Tsukamoto, T.,
Kandori, H., Mizutani, Y., Wada, A., Sudo, Y. (2018)
Production of a Light-Gated Proton Channel by
Replacing the Retinal Chromophore with Its Synthetic
Vinylene Derivative : The Journal of Physical Chemistry
Letters 9(11) : 2857-2862
Shionoya, T., Mizuno, M., Tsukamoto, T., Ikeda, K.,
Seki, H., Kojima, K., Shibata, M., Kawamura, I., Sudo,
Y., Mizutani, Y. (2018) High Thermal Stability of
Oligomeric Assemblies of Thermophilic Rhodopsin in
a Lipid Environment : The Journal of Physical Chemistry
B 122(27) : 6945-6953
Kawagoe, S., Nakagawa, H., Kumeta, H., Ishimori, K.,
Saio, T. (2018) Structural insight into proline cis/transisomerization of unfolded proteins catalyzed by thetrigger factor chaperone. : Journal of Biological Chemistry293(39) : 15095-15106
Fujisawa, T., Abe, M., Tamogami, J., Kikukawa, T.,
Kamo, N., Unno, M. (2018) Low-temperature Raman
spectroscopy reveals small chromophore distortion inprimary photointermediate of proteorhodopsin : FEBS
Letters 592(18) : 3054-3061
Stewart, N.J., Kumeta, H., Tomohiro, M., Hashimoto, T.,
Hatae, N., Matsumoto, S. (2018) Long-range heteronuclear
J-coupling constants in esters: Implications for 13C
metabolic MRI by side-arm parahydrogen-induced
polarization. : Journal of Magnetic Resonance 296 : 85-92
Tsukamoto, T., Kikuchi, C., Suzuki, H., Aizawa, T.,
Kikukawa, T., Demura, M. (2018) Implications for the
impairment of the rapid channel closing of Proteomonas
sulcata anion channelrhodopsin 1 at high Cl−
concentrations : Scientific Reports 8 : 13445⏕࿨ᶵ⬟⛉Ꮫ◊✲㒊㛛⏕≀᝟ሗゎᯒ⛉Ꮫ◊✲ศ㔝ฟᮧ ㄔ
㸦ᶵჾ฼⏝㸧
࠙Ꮫ⾡ㄽᩥࠚ
ࠚ
Tamogami, J., Kikukawa, T., Ohkawa, K., Ohsawa, N.,
Nara, T., Demura, M., Miyauchi, S., Kimura-Someya,T., Shirouzu, M., Yokoyama, S., Shimono, K., Kamo,
N. (2018) Interhelical interactions between D92 andC218 in the cytoplasmic domain regulate proton
uptake upon N-decay in the proton transport of
Acetabularia rhodopsin II : Journal of Photochemistry
and Photobiology B: Biology 183 : 35-45
Penkhrue, W., Sujarit, K., Kudo, T., Ohkuma, M.,
Masaki, K., Aizawa, T., Pathom-aree, W., Khanongnuch,
C., Lumyong, S. (2018) Amycolatopsis oliviviridis sp.
nov., a novel polylactic acid-bioplastic-degrading
actinomycete isolated from paddy soil : International
Journal of Systematic and Evolutionary Microbiology
68(5) : 1448-1454
Takayama, R., Kaneko, A., Okitsu, T., Tsunoda, S.P.,

Sénéchal, H., Keykhosravi, S., Couderc, R., Selva,
M.A., Shahali, Y., Aizawa, T., Busnel, J.M., Arif, R.,
Mercier, I., Pham-Thi, N., Charpin, D.A., Poncet, P.
(2018) Pollen/Fruit Syndrome: Clinical Relevance of
the Cypress Pollen Allergenic Gibberellin-Regulated
Protein : Allergy, Asthma & Immunology Research11(1) : 143-151
Hasemi, T., Kikukawa, T., Watanabe, Y., Aizawa, T.,
Miyauchi, S., Kamo, N., Demura, M. (2018)
Photochemical study of a cyanobacterial chloride-ion
pumping rhodopsin : Biochimica et Biophysica Acta –Bioenergetics 1860(2) : 136-146
Takahashi, H., Yoshino, M., Morita, K., Takagi, T.,
Yokoyama, Y., Kikukawa, T., Amii, H., Kanamori, T.,
Sonoyama, M. (2018) Stability of the two-dimensionallattice of bacteriorhodopsin reconstituted in partiallyfluorinated phosphatidylcholine bilayers : Biochimica− 39 −

et Biophysica Acta – Biomembranes 1861(3) : 631-642
Geng, X., Dai, G., Chao, L., Wen, D., Kikukawa, T.,
Iwasa, T. (in press) Two Consecutive Polar Amino
Acids at the End of Helix E are Important for Fast
Turnover of the Archaerhodopsin Photocycle :Photochemistry and Photobiology
࠙Ꮫ఍Ⓨ⾲ࠚ
ࠚᏳ㒊ಟᖹ㸪⳥ᕝᓠᚿ㸪⏣ẕ⚄῟㸪ሯᮏ༟㸪┦ἑᬛᗣ㸪ຍⱱ┤ᶞ㸪ฟᮧㄔ㸸Cl-࣏ࣥࣉ halorhodopsin ࡢ
H+࣏ࣥࣉ໬࡬ࡢヨࡳ㸬➨ 45 ᅇ⏕యศᏊ⛉Ꮫウㄽ఍㸪
2018 ᖺ 6 ᭶ 22 ᪥㸪኱㜰ᕷ❧኱Ꮫ⏣୰グᛕ㤋㸦኱㜰ᕷ㸧㸬ᮧ㒊ᆂ♸㸪⳥ᕝᓠᚿ㸪ሯᮏ༟㸪┦ἑᬛᗣ㸪ฟᮧㄔ㸸࢖࢜ࣥ㑅ᢥ⭷ࢆ⏝࠸ࡓ Na+࣏ࣥࣉᆺࣟࢻࣉࢩࣥࡢ
ᶵ⬟ゎᯒ㸬➨ 45 ᅇ⏕యศᏊ⛉Ꮫウㄽ఍㸪2018 ᖺ 6
᭶ 22 ᪥㸪኱㜰ᕷ❧኱Ꮫ⏣୰グᛕ㤋㸦኱㜰ᕷ㸧㸬ಖᮏ ⨾✑Ꮚ, ᖹᓠ 㔛⳯, Ᏽ 㞵㐜, బ⸨ ඃḟ, ሯᮏ ༟, ⳥ᕝ ᓠᚿ, ฟᮧ ㄔ, ୰ᮧ බ๎, ⥤㒊 ᫬ⰾ, ┦ἑ ᬛᗣ㸸ᑒධయඹⓎ⌧ࢆ฼⏝ࡋࡓ኱㔞Ⓨ⌧⣔࡟࠾࠸࡚ᢠ⳦࣌ࣉࢳࢻ cryptdin family ࡢ⏕⏘ᛶࢆỴᐃࡍࡿᅉᏊࡢゎᯒ㸬➨ 18 ᅇ᪥ᮏ⺮ⓑ㉁⛉Ꮫ఍ᖺ఍㸪2018 ᖺ 6 ᭶ 27 ᪥㸪ᮒ㮛࣓ࢵࢭ㸦᪂₲ᕷ㸧㸬
Takashi Tsukamoto 㸸 Photochemical Properties ofMicrobial Rhodopsin Pumps and Channels. Asia
Pasific Society for Biology and Medical Sciences(APSBMS) 2018 Annual Meeting, 2018 ᖺ 7 ᭶ 21 ᪥,
Sapporo Convension Center㸦ᮐᖠᕷ㸧㸬ụ⏣ ⳹Ꮚ, ✄ᮧ ຬ㞞, ஂ⡿⏣ ༤அ, ⇃ᮌ ᗣ⿱,኱す ♸Ꮨ, ሯᮏ ༟, ⳥ᕝ ᓠᚿ, ฟᮧ ㄔ, ᒸᯇ⁠⨾, ௰᐀᰿ ㇏୍, ఀᮾ ᫀ❶, ┦ἑ ᬛᗣ㸸NMR࣓ࢱ࣮࣒࣎ࣟࢆ⏝࠸ࡓ᪻⹸㣗࡟㛵ࡍࡿ◊✲㸬 ➨57 ᅇ NMR ウㄽ఍(2018)㸪2018 ᖺ 9 ᭶ 20 ᪥㸪ᮐᖠࢥࣥ࣋ࣥࢩࣙࣥࢭࣥࢱ࣮㸦ᮐᖠᕷ㸧㸬ຍ⸨ ㈗⣧, ኱἟ ᖾᬦ, ▼⏣ ༤᫛, ሯᮏ ༟, ⳥ᕝᓠ ᚿ , ฟ ᮧ ㄔ , Hans J. Vogel, ┦ ἑ ᬛ ᗣ 㸸calmodulin ⼥ྜⓎ⌧⣔ࢆ⏝࠸ࡓᢠ⳦࣌ࣉࢳࢻ኱㔞Ⓨ⌧ࡢ NMR ゎᯒ࡬ࡢᛂ⏝㸬➨ 57 ᅇ NMR ウㄽ఍(2018)㸪2018 ᖺ 9 ᭶ 20 ᪥㸪ᮐᖠࢥࣥ࣋ࣥࢩࣙࣥࢭ
ࣥࢱ࣮㸦ᮐᖠᕷ㸧㸬໭⏣ ┤ஓ, ᒣ㔝 ࡵࡄࡳ, ✄ᮧ ຬ㞞, ஂ⡿⏣ ༤அ, ⇃ᮌ ᗣ⿱, ኱す ⿱Ꮨ, ሯᮏ ༟, ⳥ᕝ ᓠᚿ,ฟᮧ ㄔ, ┦ἑ ᬛᗣ㸸NMR ࣓ࢱ࣑࣎ࣟࢡࢫ࡟ࡼࡿ
ࣈࢱ⏕⫱≧ែࡢゎᯒ㸬➨ 57 ᅇ NMR ウㄽ఍(2018)㸪
2018 ᖺ 9 ᭶ 20 ᪥㸪ᮐᖠࢥࣥ࣋ࣥࢩࣙࣥࢭࣥࢱ࣮㸦ᮐᖠᕷ㸧㸬ໟ ඞ㠀, ໭⏣ ┤ஓ, ✄ᮧ ຬ㞞, ⇃ᮌ ᗣ⿱, ኱す⿱Ꮨ, ሯᮏ ༟, ⳥ᕝ ᓠᚿ, ฟᮧ ㄔ, ୰ᮧ බ๎,⥤㒊 ᫬ⰾ, ୰ᮧ ᖾᚿ, ⋢⭜ ᬡᏊ, ┦ἑ ᬛᗣ, ᒣ㔝 ࡵࡄࡳ, ᒣᮧ ෽኱㸸Optimized Sample Storage
Strategy for Metabolic Profiling of Human Feces. ➨57 ᅇ NMR ウㄽ఍(2018)㸪2018 ᖺ 9 ᭶ 20 ᪥㸪ᮐᖠࢥࣥ࣋ࣥࢩࣙࣥࢭࣥࢱ࣮㸦ᮐᖠᕷ㸧㸬ຍ⸨ ᮅஓ, ᮧ㒊 ᆂ♸, ⳥ᕝ ᓠᚿ, ሯᮏ ༟, ┦ἑᬛᗣ, ฟᮧ ㄔ㸸Photochemical analysis of sodium
ion-pumping rhodopsin from Indibacter alkaliphilus㸬
➨ 56 ᅇ᪥ᮏ⏕≀≀⌮Ꮫ఍ᖺ఍㸪2018 ᖺ 9 ᭶ 16 ᪥㸪ᒸᒣ኱Ꮫ ὠᓥ࢟ࣕࣥࣃࢫ㸦ᒸᒣᕷ㸧㸬
Takashi Tsukamoto㸸Anion Concentration Dependency
on the Photocycle of PsuACR1: Implications for the
Impairment of its Fast Channel Function. 18th
International Conference on Retinal Proteins, 2018 ᖺ 9
᭶ 25 ᪥㸪Hockley Vallay Resort㸦ࣔࣀ࣭࢝ࢼࢲ㸧㸬
Takashi Kikukawa㸸Functional importance of trimer
formation of light-driven H+ pump Gloeobacter rhodopsin㸬
18th International Conference on Retinal Proteins,2018 ᖺ 9 ᭶ 25 ᪥㸪Hockley Vallay Resort㸦ࣔࣀ࣭࢝ࢼࢲ㸧㸬Yuga Inamura, Megumi Yamano, Naoya Kitada, Hiroyuki
Kumeta, Yasuhiro Kumaki, Yuki Onishi, Takashi
Tsukamoto, Takashi Kikukawa, Makoto Demura, Naoki
Fukuma, Yuki Murazumi, Satoshi Koike, Yasuo Kobayashi,
Tomoyasu Aizawa㸸Identification of ruminal bacterium
interactions by NMR-based metabolomics. 6th International
Life-Science Symposium (6th ILSS)㸪
2018 ᖺ 11 ᭶ 19 ᪥㸪
Hokkaido University’s Conference Hall㸦ᮐᖠᕷ㸧
Yue Zhao, Hiromu Suzuki, Md. Ruhul Kuddus,
Farhana Rumi, Takashi Tsukamoto, Takashi Kikukawa,
Makoto Demura, Tomoyasu Aizawa㸸Construction ofover-expression system for peamaclein, a peachallergenic peptide by Pichia pastoris. 6th International
Life-Science Symposium (6th ILSS)㸪2018 ᖺ 11 ᭶ 19
᪥㸪Hokkaido University’s Conference Hall㸦ᮐᖠᕷ㸧Tomoya Kato, Keisuke Murabe, Takashi Kikukawa,Takashi Tsukamoto, Tomoyasu Aizawa, Makoto Demura㸸 Na+-transport reactions of Indibacter alkaliphilus
rhodopsin probed by transient absorption spectroscopy.
6th International Life-Science Symposium (6th ILSS)㸪
2018 ᖺ 11 ᭶ 19 ᪥㸪
Hokkaido University’s ConferenceHall㸦ᮐᖠᕷ㸧
Naoya Kitada, Megumi Yamano, Yuga Inamura,
Hiroyuki Kumeta, Yasuhiro Kumaki, Yuki Onishi,− 40 −

Takashi Tsukamoto, Takashi Kikukawa, Makoto
Demura, Tomoyasu Aizawa 㸸 NMR metabolomic
profiling of health condition of Piglet. 6th International
Life-Science Symposium (6th ILSS)㸪2018 ᖺ 11 ᭶ 19
᪥㸪Hokkaido University’s Conference Hall㸦ᮐᖠᕷ㸧௨ୗࡢ」ᩘ㒊㛛฼⏝ࢢ࣮ࣝࣉࡢሗ࿌ࡣࠊᐇ㦂⏕≀ඹྠ฼⏝㒊㛛ࢆཧ↷ࠋ኱Ꮫ㝔⌮Ꮫ◊✲㝔Ύᓥ✑㸪ሯᮏ༟㸪⳥ᕝᓠᚿ㸪┦ἑᬛᗣ㸪ฟᮧㄔ㸸ග㥑ືᆺ⣽⬊ෆྥࡁࣉࣟࢺࣥ㍺㏦ࢱࣥࣃࢡ㉁ࡢ๰
ᡂ࡜ศᏊᶵᵓࡢゎ᫂㸬➨ 4 ᅇ໭ᾏ㐨኱Ꮫ㒊ᒁᶓ᩿ࢩ࣏ࣥࢪ࣒࢘㸪2019 ᖺ 1 ᭶ 25 ᪥㸪໭ᾏ㐨኱Ꮫ་Ꮫ㒊Ꮫ཭఍㤋ࣇࣛࢸ࣮࣍ࣝ㸦ᮐᖠᕷ㸧㸬

⥲ྜ༤≀㤋
◊✲㒊
 ㈨ᩱ㛤Ⓨ◊✲⣔Ụ⏣ ┿Ẏ㸦ᶵჾ฼⏝㸧࠙ⴭ᭩ࠚࠚỤ⏣┿Ẏ㸦2018㸧ࠕఀ㇋ㅖᓥ࣭㫽ᓥࡢࣇ࢕࣮ࣝࢻㄪ
ᰝ࡜໭ᾏ㐨࣭♩ᩥᓥࡢ㑇㊧㈨ᩱࡢศᯒ࠿ࡽᑤ
㛶ㅖᓥࡢ࢔࣍࢘ࢻࣜࢆ᥈ࡿࠖỈ⏣ᣅ࣭㧗ᮌṇ⯆ࠗᓥࡢ㫽㢮Ꮫ―༡すㅖᓥࡢ㫽ࢆࡵࡄࡿ⮬↛ྐ―࠘
㸪76-94㸪ᾏ​⯋㸬ࠚ࠙Ꮫ఍Ⓨ⾲ࠚ
⏣୰ᮃ⩚㸪ᑠᯘᛌḟ㸪Ụ⏣┿Ẏࠕ໬▼࣡ࢽ㢮࡬ࡢᛂ⏝ࢆ┠ⓗ࡜ࡋࡓ⌧⏕࣡ࢽ㢮ࡢࢥ࣮ࣛࢤࣥ࡟
ࡼࡿࢱࣥࣃࢡศᯒࠖ᪥ᮏ㐍໬Ꮫ఍➨ 20 ᅇ኱఍㸪2018 ᖺ 8 ᭶ 22 ᪥㸪ᮾி኱Ꮫ㸦ᮾி㸧㸬Ụ⏣┿Ẏ㸪Ἠ ὒỤ㸪ᕝୖ࿴ே㸪Ἀ⏣⤮㯞ࠕࠕ㬼ࢆᢪࡃዪࠖࡀᢪࡃ㫽ࡣఱ࠿㸽ࢥ࣮ࣛࢤࣥศᯒ࠿ࡽࡢ᳨ウࠖ᪥ᮏᩥ໬㈈⛉Ꮫ఍➨ 35 ᅇ኱఍ࠊ2018 ᖺ 7 ᭶ 7 ᪥㸪ዉⰋዪᏊ኱Ꮫ㸦ዉⰋᕷ㸧㸬− 41 −⏕≀⛉Ꮫ㒊㛛 ⾜ື⚄⤒⏕≀Ꮫศ㔝ỈἼ ㄔ
⏕≀⛉Ꮫ㒊㛛 ⾜ື⚄⤒⏕≀Ꮫศ㔝໭⏣ ୍༤
⏕≀⛉Ꮫ㒊㛛 ⾜ື⚄⤒⏕≀Ꮫศ㔝࿴ከ ࿴ᏹ
⏕≀⛉Ꮫ㒊㒊㛛 ⏕ṪⓎ⏕⏕≀Ꮫศศ㔝ᒣୗ ṇව㒊㛛 ⏕ṪⓎ⏕⏕≀Ꮫศศ㔝⏕≀⛉Ꮫ㒊㯮ᒾ 㯞㔛
㒊㛛 ⏕ṪⓎ⏕⏕≀Ꮫศศ㔝⏕≀⛉Ꮫ㒊ᮌᮧ ᩔ
⏕≀⛉Ꮫ㒊㒊㛛 ከᵝᛶ⏕≀Ꮫศศ㔝ᑠட ୍ᘯ

㈨ ᩱ࣭ࢭࣥࢱ࣮ ฼⏝ᡭ⥆ࡁࣇ࣮ࣟࢳ࣮ࣕࢺ㸦ᖹᡂ 23 ᖺᗘ௨㝆㸧࣭໭ᾏ㐨኱Ꮫ኱Ꮫ㝔⌮Ꮫ◊✲㝔㝃ᒓࢤࣀ࣒ࢲ࢖ࢼ࣑ࢡࢫ◊✲ࢭࣥࢱ࣮ᐇ㦂⏕≀ඹྠ฼⏝㒊㛛 ᪋タ฼⏝ࡢᡭᘬࡁ㸦ู⾲ 1 ඹྠ฼⏝ᶵჾࣜࢫࢺ ᖹᡂ 25 ᖺ 4 ᭶௨㝆㸧㸦ู⾲ 2 ᐇ㦂⏝ື≀౪⤥⣽๎㸧࣭໭ᾏ㐨኱Ꮫ኱Ꮫ㝔⌮Ꮫ◊✲㝔㝃ᒓࢤࣀ࣒ࢲ࢖ࢼ࣑ࢡࢫ◊✲ࢭࣥࢱ࣮つ⛬࣭໭ᾏ㐨኱Ꮫ኱Ꮫ㝔⌮Ꮫ◊✲㝔㝃ᒓࢤࣀ࣒ࢲ࢖ࢼ࣑ࢡࢫ◊✲ࢭࣥࢱ࣮㐠Ⴀጤဨ఍ෆつ࣭໭ᾏ㐨኱Ꮫ኱Ꮫ㝔⌮Ꮫ◊✲㝔㝃ᒓࢤࣀ࣒ࢲ࢖ࢼ࣑ࢡࢫ◊✲ࢭࣥࢱ࣮㛗㑅⪃ෆつ− 43 −

ࢭࣥࢱ࣮฼⏝ᡭ⥆ࡁࣇ࣮ࣟࢳ࣮ࣕࢺձ ஦๓࡟฼⏝ྍ⬟ᶵჾࡸࢫ࣮࣌ࢫࡢ✵ࡁ≧ἣࢆ☜ㄆࡋ࡚ୗࡉ࠸ࠋ
ࢭࣥࢱ࣮࢙࢘ࣈࢧ࢖ࢺ    https://www.sci.hokudai.ac.jp/gdynamics/ၥ࠸ྜࢃࡏඛ㸦ࢭࣥࢱ࣮஦ົ㸧e-mail : cepa@sci.hokudai.ac.jp (ෆ⥺ 3580)

Ўղ ฼⏝⏦ㄳ᭩࣭฼⏝⪅ྡ⡙ࢆࢭࣥࢱ࣮࢙࢘ࣈࢧ࢖ࢺࡼࡾࢲ࣮࢘ࣥࣟࢻࡍࡿࠋ
㸦඲㒊㛛ඹ㏻ࣇ࢛࣮࣐ࢵࢺ࡛ࡍࠋ㸧ὀ㸧ᐇ㦂⏕≀ඹྠ฼⏝㒊㛛㸦ᮾᲷ㸧฼⏝⪅ࡢ࠺ࡕ᪂つࡢ᪉࡜ࠊ⥅⥆฼⏝࡛࠶ࡿࡀ⏦ㄳෆᐜ࡟ኚ᭦ࡢ࠶ࡿ᪉ࡣࠊୖグࡢ௚ࠕᮾᲷ฼⏝⏦ㄳ᭩ࠖࡢᥦฟࡀᚲせ࡛ࡍࠋЎճ ྛᵝᘧ࡟ᚲせ஦㡯ࢆグධࡍࡿࠋЎմ グධ῭ࡳࡢྛᵝᘧࢆ⤒㈝ࡢᨭᡶ㈐௵⪅ࡢ e-mail ࢔ࢻࣞࢫ࠿ࡽࢭࣥࢱ࣮஦ົ
e-mail ࡬ࡢῧ௜࡟࡚ᥦฟࡍࡿࠋ
ὀ㸧ᡤᒓᶵ㛵࣭㒊ᒁ࡟࡚ྲྀᚓࡋࡓ࣓࣮ࣝ࢔ࢻࣞࢫ࡛࠶ࡿࡇ࡜ࠋЎյ ฼⏝チྍ᭩ࢆཷࡅྲྀࡿࠋЎն ฼⏝㛤ጞࠋ
ὀ 1㸧ධ㤋Ⓩ㘓ࡢࡓࡵࠊಶே ID ࡶูᵝᘧ࡟࡚ᥦฟࡋ࡚࠸ࡓࡔࡁࡲࡍࠋ
ὀ 2㸧ᐇ㦂⏕≀ඹྠ฼⏝㒊㛛㸦ᮾᲷ㸧฼⏝⪅ࡣࠊ஦๓࡟ࠕᮾᲷ᪂つ฼⏝⪅ㅮ⩦఍ࠖࡢཷㅮࡀᚲせ࡛ࡍࠋ− 44 −

໭ᾏ㐨኱Ꮫ኱Ꮫ㝔⌮Ꮫ◊✲㝔㝃ᒓࢤࣀ࣒ࢲ࢖ࢼ࣑ࢡࢫ◊✲ࢭࣥࢱ࣮
ᐇ㦂⏕≀ඹྠ฼⏝㒊㛛㸦ᮾᲷ㸧᪋タ฼⏝ࡢᡭᘬࡁ
㸯㸬฼⏝⏦ㄳ㸦㸯㸧 ᙜ㒊㛛㸦ᮾᲷ㸧㣫⫱ᐊ࣭ᐇ㦂ᐊ➼ࡢ฼⏝ࢆᕼᮃࡍࡿ᫬ࡣࠊ㟁ヰ࠿㟁Ꮚ࣓࣮࡛ࣝᚲࡎ஦๓࡟
ࡈ㐃⤡ࡃࡔࡉ࠸⅛
㸦㸰㸧 ◊✲ᐊࡈ࡜࡟฼⏝⏦ㄳ᭩࠾ࡼࡧ฼⏝⪅ྡ⡙࡬ᚲせ஦㡯ࢆグධࡋࠊᥦฟࡋ࡚࠸ࡓࡔࡁࡲࡍࠋ
฼⏝チྍ᭩㸦༢ᖺᗘ㸧ࢆ⤒㈝ࡢᨭᡶ㈐௵⪅ᐄ࡟Ⓨ⾜࠸ࡓࡋࡲࡍࠋ㸦㸱㸧 㑇ఏᏊ⤌᥮࠼ᐇ㦂ࡸື≀ᐇ㦂ࢆ⾜࠺᪉ࡣᏛෆࡢ࣮ࣝࣝ࡟ᚑࡗ࡚⏦ㄳࢆ⾜࠸ࠊࡑࡢチྍ␒ྕࡲࡓࡣᢎㄆ␒ྕࢆୖグ㸦㸰㸧฼⏝⏦ㄳ᭩࡟グ㍕ࡍࡿᚲせࡀ࠶ࡾࡲࡍࠋ㸦㸲㸧 ᪂つ฼⏝⪅ࡀᙜ㒊㛛ࡢධ㤋ࢩࢫࢸ࣒࡬Ⓩ㘓ࢆᕼᮃࡉࢀࡿሙྜࡣࠊ⌮Ꮫ◊✲㝔ࡢᐃࡵࡿ᭩ᘧ
࡛ࡢ⏦ㄳࡶ࠾㢪࠸ࡋࡲࡍࠋࡲࡓࠊᙜ㒊㛛୺ദࡢ฼⏝⪅ㅮ⩦఍ࡢཷㅮࡶ⩏ົ௜ࡅ࡚࠾ࡾࡲࡍࡢ
࡛ࠊᛀࢀࡎ࡟ཷㅮࡋ࡚ࡃࡔࡉ࠸ࠋ
㸦㸳㸧฼⏝⏦ㄳᡭ⥆ࡁࡣ㝶᫬ཷࡅ௜ࡅࡲࡍࠋ
㸦㸴㸧⥅⥆ࡋ࡚฼⏝ࡉࢀࡿሙྜࡢᖺᗘẖࡢ᭦᪂ᡭ⥆ࡁࡣ୙せ࡛ࡍࠋࡓࡔࡋࠊୖグ㸦㸰㸧࡟࠾࠸࡚
ᥦฟࡋࡓෆᐜ࡟ኚ᭦ࡸ㏣ຍࡀ⏕ࡌࡓሙྜ࡟ࡣᚲせ᭩㢮ࡢ෌ᥦฟࡋ࡚࠸ࡓࡔࡁࡲࡍࡢ࡛ࠊ⫋ဨࡢᣦ♧ࢆ௮࠸࡛ࡃࡔࡉ࠸ࠋ
㸦㸵㸧ୖグ㸦㸴㸧࡟࠾࠸࡚฼⏝⪅ྡ⡙࡟᪂ࡓ࡟㏣ຍࡉࢀࡓ⪅ࡣ㸦㸲㸧࡟ヱᙜࡋࡲࡍࡢ࡛ࠊྠᵝࡢᡭ⥆ࡁཬࡧ฼⏝⪅ㅮ⩦఍ࡢཷㅮࢆ࠾㢪࠸ࡋࡲࡍࠋ
㸦㸶㸧฼⏝ࢆ⤊஢ࡉࢀࡿሙྜ࡟ࡣᥦฟ᭩㢮ࡣ࠶ࡾࡲࡏࢇࡀࠊ᧔㏥ࡉࢀࡿ㸰㐌㛫๓ࡲ࡛࡟㐃⤡ࢆ࠾㢪࠸ࡋࡲࡍࠋ฼⏝ᩱ㔠ࡣ᭶༢఩࡛࠶ࡿⅬ࡟ࡈὀពࡃࡔࡉ࠸ࠋ
㸰㸬ࡆࡗṑ㢮㣫⫱ᐊ㸦㸯㸧ࡆࡗṑ㢮㣫⫱ᐊ࡟ධᐊࡢ㝿ࡣࠊ㜵␿ୖࠊᚲࡎⓑ⾰ࢆ╔⏝ࡋᑓ⏝ࡢୖᒚࡁ࡬ᒚࡁ᭰࠼࡚ࡃࡔ ࡉ࠸ࠋ㸦㸰㸧ࡆࡗṑ㢮㣫⫱ᐊ࡬ࡢື≀ࡢᦙධ࣭ᦙฟࡣࠊ㣫⫱㒊ᢸᙜ⫋ဨࡢᣦ♧࡟ᚑࡗ࡚ࡃࡔࡉ࠸ࠋ
㸦㸱㸧㸦ᐇ㦂⏝ື≀ࡢ౪⤥㸧  ᙜ㒊㛛࡛ࡣࠊࣛࢵࢺ࣭࣐࢘ࢫ➼ࡢᐇ㦂⏝ື≀ࡢ౪⤥ࢆ⾜ࡗ࡚࠾ࡾࡲࡍࠋ౪⤥ᕼᮃ⪅ࡣࠊู⾲㸰ࡢࠕᐇ㦂⏝ື≀౪⤥⣽๎ࠖ࡟ᚑࡗࡓᚲせᡭ⥆ࢆ῭ࡲࡏ࡚ࡃࡔࡉ࠸ࠋ
㸦㸲㸧㸦ື≀ࡢ㣫⫱㸧− 45 −  ື≀ࡢ⤥㣵࣭⤥Ỉ࣭ᗋᩜࡢ஺᥮ࡣࠊ฼⏝⪅ࡀ⾜ࡗ࡚ࡃࡔࡉ࠸ࠋ
㸦㸳㸧㸦࢚ࢧࠊᗋᩜ㸧 㣫⫱࡟ᚲせ࡞࢚ࢧࡣ฼⏝⪅ࡀ⏝ពࡋ࡚ࡃࡔࡉ࠸ࠋࡓࡔࡋࠊᙜ㒊㛛࡛ࡶࡆࡗṑ㢮⏝ࡢ࢚ࢧ㸯⿄  ਻ࢆᐇ㈝࡛࠾ศࡅ࡛ࡁࡲࡍࠋ㣫⫱࡟ᚲせ࡞ᗋᩜࡣࡇࡕࡽ࡛⏝ពࡋࡓࡶࡢࢆ౑⏝ࡍࡿࡇ࡜ࡶྍ⬟࡛ࡍࠋ
㸦㸴㸧㸦ᚤ⏕≀ࣔࢽࢱࣜࣥࢢ⾲ࡢᥦฟ㸧  㣫⫱ᐊ࡟࠾ࡅࡿఏᰁ⑓⶝ᘏࢆண㜵ࡍࡿࡓࡵࠊእ㒊ࡢᶵ㛵࠿ࡽᦙධࡋࡓື≀ࢆ㣫⫱ࡍࡿሙྜ
ࡣࠊᦙධࡍࡿື≀ࡀᙜ㒊㛛ࡢࠕᶆ‽సᴗᡭ㡰᭩ࠖ࡟つᐃࡍࡿ⑓ཎయ࡟ឤᰁࡋ࡚࠸࡞࠸ࡇ࡜ࢆ
ド᫂ࡍࡿ᭩㢮㸦ᚤ⏕≀ࣔࢽࢱࣜࣥࢢ⾲㸧ࢆணࡵᥦฟࡋ࡚ࡃࡔࡉ࠸ࠋࡓࡔࡋࠊᦙධ๓㸯ᖺ௨ෆ࡟ྠࡌᶵ㛵࠿ࡽྠ✀ࡢື≀ࢆᦙධࡋࡓ㝿࡟ᚤ⏕≀ࣔࢽࢱࣜࣥࢢ⾲ࢆᥦฟࡋࡓሙྜ࡟ࡣࠊᨵࡵ
࡚ᥦฟࡍࡿᚲせࡣ࠶ࡾࡲࡏࢇࠋ
㸦㸵㸧㸦ࡆࡗṑ㢮ฎ⨨ᐊࡢ฼⏝㸧  ࡆࡗṑ㢮㣫⫱ᐊࡢ฼⏝⪅ࡣࠊᙜ㒊㛛ෆ࡛㣫⫱ࡍࡿື≀࡬ࡢᐇ㦂ⓗฎ⌮ࢆ⾜࠺ࡓࡵ࡟ฎ⨨ᐊࢆ฼⏝ࡍࡿࡇ࡜ࡀ࡛ࡁࡲࡍࠋ฼⏝ࡋࡓ࠸ሙྜࡣ஦๓࡟✵ࡁ≧ἣࢆᢸᙜ⪅࡟☜ㄆࡋࠊ฼⏝ᕼᮃ᪥᫬࡜ฎ⨨ෆᐜࢆᢸᙜ⪅࡬࠾ఏ࠼ࡃࡔࡉ࠸ࠋ
㸱㸬࢞ࣛࢫᐊ࣭ᅡሙ
㸦㸯㸧ᘓ≀㸯㸰㝵࡟௜ᒓࡋࡓ࢞ࣛࢫᐊࠊ࠾ࡼࡧᒇእ࡟ᅡሙࡀ⏝ពࡉࢀ࡚࠸ࡲࡍࠋ฼⏝ࢆᕼᮃࡍࡿ
㝿࡟ࡣ஦๓࡟ᢸᙜ⫋ဨ࡜฼⏝᪉ἲࢆᡴࡕྜࢃࡏࡓ࠺࠼࡛฼⏝⏦ㄳࢆࡋ࡚ࡃࡔࡉ࠸ࠋ
㸦㸰㸧฼⏝༊ᇦࡢ㝖ⲡࡸΎᤲࡣ฼⏝⪅ྛ⮬࡛⾜ࡗ࡚ࡃࡔࡉ࠸ࠋ㎰ල㢮ࡢ㈚ฟࡶ⾜ࡗ࡚࠸ࡲࡍࡢ࡛ࠊ
฼⏝ࡢ㝿࡟ࡣ⫋ဨ࡬⏦ࡋฟ࡚ࡃࡔࡉ࠸ࠋ
㸦㸱㸧᱂ᇵᗫᲠ≀ࡣᏛෆࡢᗫᲠ≀ฎ⌮᪉ἲ࡟ᚑࡗ࡚ฎศࡋ࡚ࡃࡔࡉ࠸ࠋ
㸲㸬Ỉᵴᐊ
㸦㸯㸧Ỉᵴᐊ࡟ࡣࠊ ᗘࢥࣥࢺ࣮࣮ࣟࣛ௜ࡢỈᵴࡀ⏝ពࡉࢀ࡚࠸ࡲࡍࠋࡲࡓࠊ㣫⫱Ỉ࡜ࡋ࡚஭Ỉ㸦㞧⏝Ỉ㸧࡜ᾏỈࢆ฼⏝࡛ࡁࡲࡍࠋ
㸦㸰㸧฼⏝⪅ࡣࠊ༙ᖺ࡟୍ᗘࠊỈᵴ࡜᤼Ỉ⁁ࡢᤲ㝖୪ࡧ࡟ℐ㐣◁ࡢὙίࢆ⾜ࡗ࡚ࡃࡔࡉ࠸ࠋࡲࡓࠊ
Ỉᵴࡢ฼⏝ࡀ⤊஢ࡋࡓሙྜࡶྠᵝ࡛ࡍࠋ㸳㸬ඹྠ฼⏝㸦㸯㸧฼⏝⪅ࡣࠊᙜ㒊㛛ࢆ฼⏝ࡋ࡚⾜࠺ᐇ㦂࣭◊✲ࡢ㐙⾜ୖࠊᚲせ࡜ㄆࡵࡽࢀࡿసᴗ࡟ࡘ࠸࡚ࠊ

− 46 −

ඹྠ฼⏝ᶵჾ㸦ู⾲㸯ཧ↷㸧ࢆ฼⏝ࡍࡿࡇ࡜ࡀ࡛ࡁࡲࡍࠋᕼᮃ⪅ࡣ⏦ㄳ᭩ࠕ฼⏝ᶵჾࠖḍ࡟௜グࡋࠊ฼⏝᪉ἲࢆ⫋ဨ࡟☜ㄆࡋ࡚࠿ࡽࡈ฼⏝ࡃࡔࡉ࠸ࠋ
㸦㸰㸧㒊㛛ෆ࡛ࡢ㣫⫱࣭ᐇ㦂ࡢࡓࡵ୍᫬ⓗ࡟㣫ᩱㄪᩚᐊࢆ฼⏝ࡍࡿࡇ࡜ࡀ࡛ࡁࡲࡍࠋ౑⏝ᚋࡣᤲ㝖࣭ࢦ࣑ฎ⌮➼ࢆ⾜࠸ࠊඖࡢ≧ែ࡟ᡠࡋ࡚ࡃࡔࡉ࠸ࠋ
㸦㸱㸧㣫⫱࣭ᐇ㦂࡟⏝࠸ࡓ⬨᳝ື≀ࡢṚయࡣࠊᑓ⏝ࣇ࣮ࣜࢨ࣮࡟ᗫᲠࡋグ㘓⡙࡬ᩘ㔞➼ࢆグධࡋ
࡚ࡃࡔࡉ࠸ࠋ
㸦㸲㸧฼⏝⪅ࡀ㝶᫬฼⏝ሙᡤࢆΎᤲ࡛ࡁࡿࡼ࠺ࠊᤲ㝖㐨ලࡢ㈚ࡋฟࡋࢆ⾜ࡗ࡚࠾ࡾࡲࡍࠋ⫋ဨ࡟ࡈ┦ㄯ࠸ࡓࡔࡃ࠿ࠊ㸰㝵ᗯୗ࡟࠶ࡿࣟࢵ࣮࢝ࡢᤲ㝖㐨ලࢆࡈ౑⏝ࡃࡔࡉ࠸ࠋ
㸦㸳㸧Ᏻ඲࢟ࣕࣅࢿࢵࢺࠊ&2 ࢖࣮ࣥ࢟ࣗ࣋ࢱ➼ࡀタ⨨ࡋ࡚࠶ࡿඹྠ฼⏝ᐊࢆ฼⏝ࡍࡿሙྜࠊ฼⏝⏦ㄳ᭩࡬ࡢグ㍕ࡀᚲせ࡛ࡍࠋᐇ㦂࡟ᚲせ࡞ᾘ⪖ရࡣ฼⏝⪅ࡀ⏝ពࡋ㐺ษ࡟ᗫᲠࡋ࡚ࡃࡔࡉ࠸ࠋタ⨨ᶵჾ㢮ࡢ฼⏝᫬㛫ࡸሙᡤ➼࡟㛵ࡋ࡚ࡣ฼⏝⪅ྠኈࡀヰࡋྜ࠸ㄪᩚࡋ࡚ࡃࡔࡉ࠸ࠋ᪂ࡓ࡟
ᶵჾ➼ࢆ฼⏝ᐊ࡟ᣢࡕ㎸ࡴሙྜࠊ๓ࡶࡗ࡚⫋ဨࡲ࡛࠾▱ࡽࡏࡃࡔࡉ࠸ࠋ
㸴㸬✵ㄪ➼ࡢ␗ᖖ࣭ࢺࣛࣈࣝ
㸦㸯㸧ྛᐊ࡟࠶ࡿ✵ㄪไᚚ┙ࡣᡤᐃࡢタᐃ್࡟ࢭࢵࢺࡉࢀ࡚࠸ࡲࡍࡢ࡛ࠊኚ᭦ࢆᕼᮃࡍࡿሙྜࡣ ᚲࡎ⏦ࡋฟ࡚ࡃࡔࡉ࠸ࠋ
㸦㸰㸧ྛᐊࡢ㆙ሗタᐃ ᗘ㸦ୖ㝈࣭ୗ㝈㸧ࡣᙜ㒊㛛⫋ဨࡢ᪉࡛タᐃࡋࡲࡍࡀࠊ฼⏝ࡋ࡚࠸ࡿ㒊ᒇ࡟ࡘ࠸࡚≉࡟ᕼᮃࡀ࠶ࡿሙྜ࡟ࡣࡈ┦ㄯࡃࡔࡉ࠸ࠋ
㸦㸱㸧ᐊࡢ␗ᖖࢆ▱ࡽࡏࡿ㆙ሗ┙ࡀࠊ㸯㝵⋞㛵࣮࣍ࣝ࡜㸰㝵ᮾഃᗯୗ୰ኸ࠾ࡼࡧ㸰㝵ᮾഃᗯୗࡢ໭➃࡟ഛࢃࡗ࡚࠸ࡲࡍࠋ㆙ሗࡀ㬆ࡗ࡚࠸ࡿ࡜ࡁࡣ⫋ဨ࡬㐃⤡㢪࠸ࡲࡍࠋ
㸦㸲㸧⫋ဨ୙ᅾࡢ᫬㛫ᖏ࡟㆙ሗࡀ㬆ࡗ࡚࠸ࡓࡾࢺࣛࣈࣝ࡟㐼㐝ࡋࡓࡾࡋࡓ㝿ࡣࠊ⌮Ꮫ㒊㸰ྕ㤋ࡢ㜵⅏ࢭࣥࢱ࣮㸦ෆ⥺ 㸧࡬ఏ࠼࡚ࡃࡔࡉ࠸ࠋ㸵㸬೵㟁㸦㸯㸧ᙜ㒊㛛࡟ࡣ೵㟁࡟ࡼࡿ⏕≀ཬࡧᐇ㦂⣔࡬ࡢᙳ㡪ࢆ᭱ᑠ㝈࡟㣗࠸Ṇࡵࡿࡓࡵࠊ✵ㄪ࡞࡝࡬ࡢᚲせ᭱ప㝈ࡢᶵჾࡢ㐠㌿ࢆಖドࡍࡿ⮬ᐙⓎ㟁⿦⨨ࡀഛࢃࡗ࡚࠸ࡲࡍࠋ
㸦㸰㸧⮬ᐙⓎ㟁⿦⨨ࡣ୍᫬ⓗ࡞㟁ຊ౪⤥ࡋ࠿࡛ࡁࡲࡏࢇࡢ࡛ࠊ஦๓࡟೵㟁ࡀ▱ࡽࡉࢀ࡚࠸ࡿሙྜ
ࡣ೵㟁๓࡟ᚲࡎྛᶵჾࡢ㟁※ࢆษࡗ࡚ࡃࡔࡉ࠸ࠋ
㸦㸱㸧೵㟁࡟ࡣ᩿Ỉࡀక࠺ሙྜࡀ࠶ࡾࡲࡍࠋ⤥Ỉ࣭⤥‫ࡈࡣ⏝౑ࡢࣞ࢖ࢺࡧཬ‮‬㐲៖ࡃࡔࡉ࠸ࠋ
㸦㸲㸧⥭ᛴ࡟೵㟁ࡀⓎ⏕ࡍࡿ࡜ᩘ⛊ᚋ࡟⮬ᐙⓎ㟁⿦⨨ࡀసືࡋࡲࡍࠋᩘศ⤒ࡗ࡚ࡶ㏻㟁ࡋ࡞࠸➼− 47 −

ࡢࢺࣛࣈࣝࡀ࠶ࡗࡓሙྜ࡟ࡣ⮳ᛴ⫋ဨ࡬㐃⤡㢪࠸ࡲࡍࠋ㸦⫋ဨ୙ᅾ᫬ࡣ㸴㸬㸦㸲㸧࡜ྠᵝ࡟ᑐᛂࡋ࡚ࡃࡔࡉ࠸ࠋ㸧㸶㸬⣧Ỉࠊ㉸⣧ỈࠊᾏỈ㸦㸯㸧ᙜ㒊㛛࡛ࡣࠊ⣧Ỉ㸦52㸧ࠊ㉸⣧Ỉ㸦࣑ࣜ 4㸧ࠊᾏỈࡢࡳࡢᥦ౪ࡶ⾜ࡗ࡚࠸ࡲࡍࠋᕼᮃ⪅ࡣ౯᱁ࢆ☜ㄆࡋࠊ฼⏝⏦ㄳ᭩ࢆᥦฟࡋ࡚ࡃࡔࡉ࠸ࠋ
㸦㸰㸧౑⏝ࡢ㝿࡟ࡣᚲせ㔞ศࡢᐜჾࢆᣢཧࡋࠊᡤᐃࡢ⏝⣬࡟グධࡋ࡚ࡈ฼⏝ࡃࡔࡉ࠸ࠋ
㸷㸬⤒㈝
㸦㸯㸧✵ㄪࠊࡑࡢ௚ㅖタഛࡢ⥔ᣢ࣭⟶⌮ࡢࡓࡵࡢㅖ⤒㈝ࡢ୍㒊࡜ඹ㏻ᾘ⪖ရࡢ㈝⏝࡜ࡋู࡚࡟ᐃ
ࡵࡿᩱ㔠ࢆ฼⏝⪅࡟㈇ᢸࡋ࡚࠸ࡓࡔࡁࡲࡍࠋ࡞࠾ࠊ฼⏝ᩱ㔠ࡣᖺᗘጞࡵ࠿ࡽ㸱ࣨ᭶ẖ࡟ᚩ཰ࡉࡏ࡚࠸ࡓࡔࡁࡲࡍࠋ
㸯㸮㸬ࡑࡢ௚
㸦㸯㸧ධ㤋ࡍࡿ㝿࡟Ⓩ㘓 ,& ࣮࢝ࢻࢆᚲࡎᦠᖏࡋ࡚ୗࡉ࠸ࠋ
㸦㸰㸧ᙜ㒊㛛ෆࡣᅵ㊊⚗Ṇ࡛ࡍࡢ࡛ࠊᑓ⏝ࡢୖᒚࡁࢆࡈ฼⏝ࡃࡔࡉ࠸ࠋ
㸦㸱㸧฼⏝⪅ࡀⓎὀࡋࡓᶵჾಟ⧋࣭≀ရࢆ┤᥋ᙜ㒊㛛࡬⣡ရࡍࡿ㝿ࡣࠊ஦๓࡟⣡ရᴗ⪅࡜⣡ရ᪥ࢆ⫋ဨ࡬㐃⤡ࡋࠊᦙධࡸసᴗࡢ᪥᫬ࡢㄪᩚࢆ⾜ࡗ࡚ࡃࡔࡉ࠸ࠋ㸦㸲㸧 ᙜ㒊㛛ෆ࡛ࡢ≀ရࡢᢅ࠸࡟ࡘ࠸࡚ࡣࠊ௨ୗࡢⅬ࡟ᚚὀពࡃࡔࡉ࠸ࠋ∝ ≀ရ࡟ࡣ඲࡚ᡤ᭷⪅࣭ᡤᒓㅮᗙ࡞࡝ࢆศ࠿ࡾࡸࡍ࠸ሙᡤ࡟᫂グࡋ࡚ࡃࡔࡉ࠸ࠋ
∝ ഛရ␒ྕࡀࡘ࠿࡞࠸ჾල࣭ᶵჾ࣭ᾘ⪖ရ࡟ࡘ࠸࡚ࡶᡤᒓࡀศ࠿ࡿࡼ࠺࡟ㅮᗙྡ➼ࢆࡈグධࡃࡔࡉ࠸ࠋ
∝ ኱ࡁ࡞㈨ᮦࡸ≀ရࢆฟࡋධࢀࡍࡿ㝿࡟ࡣࠊ஦๓࡟⫋ဨ࡬㐃⤡ࡋࠊ㐺ᐅ⏦ㄳ᭩ࡢ᭩ࡁ᥮࠼
ࢆ⾜ࡗ࡚ࡃࡔࡉ࠸ࠋ∝ ⏦ㄳࡢ㝿࡟฼⏝ሙᡤ࡜ࡋ࡚᫂グࡉࢀ࡚࠸࡞࠸ሙᡤ㸦౛࠼ࡤ⮬ศࡢ౑⏝ሙᡤ௨እࡢ㒊ᒇ࣭ᗯୗ࣭⋞㛵➼㸧࡟୍᫬ⓗ࡟࡛ࡶ≀ရࢆタ⨨ࡍࡿሙྜ࡟ࡣࠊ⫋ဨࡲ࡛ࡈ㐃⤡ࡃࡔࡉ࠸ࠋ
㸦㸳㸧฼⏝ࢆ⤊஢ࡍࡿ㝿࡟ࡣࠊ㒊ᒇ࣭ᶵჾ➼ࢆඖࡢ≧ែ࡟ᡠࡋࠊ⫋ဨࡢ☜ㄆࢆཷࡅ࡚ࡃࡔࡉ࠸ࠋ
㸦㸴㸧ᙜ㒊㛛㐠Ⴀ㈝ࡢ୍㒊ࡣࠊⓙᵝ࠿ࡽࡢ฼⏝ᩱ㔠࡟ࡼࡗ࡚㈥ࢃࢀ࡚࠸ࡲࡍࠋග⇕Ỉᩱࡣ㐠Ⴀ㈝⥲㢠ࡢከࡃࢆ༨ࡵ࡚࠾ࡾࠊᙜ㒊㛛ࢆ㐠Ⴀࡍࡿ࠺࠼࡛㈇ᢸ࡜࡞ࡗ࡚࠾ࡾࡲࡍࠋᙜ㒊㛛ࢆඹྠ฼⏝᪋タ࡜ࡋ࡚ࡼࡾ୍ᒙά⏝ࡍࡿࡓࡵ࡟ࡶࠊ㟁Ẽ࣭Ỉ㐨➼ࡢ↓㥏౑࠸ࢆ࡞ࡃࡋࠊ┬࢚ࢿࣝࢠ࣮໬ࢆᚰ᥃ࡅ࡚ࡃࡔࡉ࠸ࠋ

− 48 −ู⾲㸯ඹྠ฼⏝ᶵჾ
࣮࢜ࢺࢡ࣮ࣞࣈ㸦ඹྠᐇ㦂ᐊ㸧   ஝⇕⁛⳦ᶵ㸦ඹྠᐇ㦂ᐊ㸧    〇ịჾ㸦㣫ᩱㄪᩚᐊ㸧&2 ࢖࣮ࣥ࢟ࣗ࣋ࢱ㸦ඹྠᐇ㦂ᐊ㸧Ᏻ඲࢟ࣕࣅࢿࢵࢺ㸦ඹྠᐇ㦂ᐊ㸧㸫Υࣇ࣮ࣜࢨ࣮㸦୰ᗯୗ㸧




ู⾲㸰
ᐇ㦂⏝ື≀౪⤥⣽๎㸬 ᐇ㦂⏕≀ඹྠ฼⏝㒊㛛࡟࠾ࡅࡿ࣐࢘ࢫ࣭ࣛࢵࢺࡢ౪⤥࡟ࡘ࠸࡚ࡣࠊࡇࡢ⣽๎ࡢᐃࡵࡿ࡜ࡇࢁ࡟ࡼࡿࠋ㸬 ᐇ㦂⏝ື≀ࡣࠊཎ๎࡜ࡋ࡚◊✲ᐇ㦂⏝࡟౪⤥ࡍࡿࠋ㸬 ᐇ㦂⏝ື≀ࡢ㣫⫱⤒㈝ࡢ㈇ᢸࡣࠊḟࡢ࡜࠾ࡾ࡜ࡍࡿࠋ
ࣁࢶ࢝ࢿࢬ࣑  ⣔⤫ࠊᛶูࠊ㐌㱋ࡢ㸦࣐࢘ࢫ㸧ᣦᐃ᭷ࡾ  㢌ᙜࡓࡾ   ෇    ᣦᐃ↓ࡋ  㢌ᙜࡓࡾ   ෇ዷፎಶయ  㢌ᙜࡓࡾ  ෇ࢻࣈࢿࢬ࣑  ⣔⤫ࠊᛶูࠊ㐌㱋ࡢ㸦ࣛࢵࢺ㸧 ᣦᐃ᭷ࡾ  㢌ᙜࡓࡾ  ෇     ᣦᐃ↓ࡋ  㢌ᙜࡓࡾ   ෇ዷፎಶయ  㢌ᙜࡓࡾ  ෇ 㸬 ౪⤥ࡢ⏦㎸ࡳࡣࠊணࡵࡆࡗṑ㢮㣫⫱㒊ࡢᢸᙜ⫋ဨ࡜ᡴྜࡏࢆ⾜࠸ࠊᡤᐃࡢ⏝⣬࡟ᚲせ஦㡯ࢆグධࡢୖࠊ⾜࠺ࡇ࡜ࠋ㸬 㸬࡟つᐃࡍࡿ㈝⏝ࡢㄳồࡣࠊᐇ㦂⏕≀ඹྠ฼⏝㒊㛛ࡢ฼⏝ᩱࡢㄳồ࡟ྜࢃࡏ࡚
⾜࠺ࠋ౪⤥࡟㝿ࡋ࡚ᚲせ࡜࡞ࡿ㈝⏝࡟ࡘ࠸࡚ࡣࠊ⏦㎸⪅ࡀ㈇ᢸࡍࡿࠋ
− 49 −

໭ᾏ㐨኱Ꮫ኱Ꮫ㝔⌮Ꮫ◊✲㝔㝃ᒓࢤࣀ࣒ࢲ࢖ࢼ࣑ࢡࢫ◊✲ࢭࣥࢱ࣮つ⛬ᖹᡂ㸰㸮ᖺ㸯㸯᭶㸯᪥ᾏ኱㐩➨㸯㸳㸮ྕ 㸦㊃᪨㸧➨㸯᮲ ࡇࡢつ⛬ࡣ㸪ᅜ❧኱Ꮫἲே໭ᾏ㐨኱Ꮫ⤌⧊つ๎㸦ᖹᡂ㸯㸴ᖺᾏ኱㐩➨㸱㸯ྕ㸧➨㸰㸵᮲ࡢ㸵➨㸲㡯ࡢつᐃ࡟ᇶ࡙ࡁ㸪໭ᾏ㐨኱Ꮫ኱Ꮫ㝔⌮Ꮫ◊✲㝔㝃ᒓࢤࣀ࣒ࢲ࢖ࢼ࣑ࢡ
ࢫ◊✲ࢭࣥࢱ࣮㸦௨ୗࠕࢭࣥࢱ࣮ࠖ࡜࠸࠺ࠋ
㸧ࡢ⤌⧊ཬࡧ㐠Ⴀ࡟ࡘ࠸࡚ᐃࡵࡿࡶࡢ࡜ࡍࡿࠋ 㸦┠ⓗ㸧➨㸰᮲ ࢭࣥࢱ࣮ࡣ㸪໭ᾏ㐨኱Ꮫࡢ◊✲⪅࡟ᑐࡋ࡚㸪㑇ఏᏊཬࡧᰁⰍయ࡟㛵ࡍࡿ◊✲ࢆ⾜࠺ࡓࡵࡢ᪋タཬࡧタഛࢆᥦ౪ࡍࡿ࡜࡜ࡶ࡟㸪ື≀㸪᳜≀ࡑࡢ௚⏕≀ᮦᩱࡢ౪⤥ࢆ⾜࠺ࡇ࡜࡟ࡼࡾ㸪⏕≀⛉Ꮫศ㔝ࡢ◊✲ࡢ㐍ᒎ࡟ᐤ୚ࡍࡿࡇ࡜ࢆ┠ⓗ࡜ࡍࡿࠋ 㸦ඹྠ฼⏝㒊㛛㸧➨㸱᮲ ࢭࣥࢱ࣮࡟㸪ḟ࡟ᥖࡆࡿඹྠ฼⏝㒊㛛ࢆ⨨ࡃࠋ  ື≀ᰁⰍయඹྠ฼⏝㒊㛛  㑇ఏᏊᐇ㦂ඹྠ฼⏝㒊㛛  ᐇ㦂⏕≀ඹྠ฼⏝㒊㛛 㸦⫋ဨ㸧➨㸲᮲ ࢭࣥࢱ࣮࡟㸪ࢭࣥࢱ࣮㛗ࡑࡢ௚ᚲせ࡞⫋ဨࢆ⨨ࡃࠋ 㸦ࢭࣥࢱ࣮㛗㸧➨㸳᮲ ࢭࣥࢱ࣮㛗ࡣ㸪໭ᾏ㐨኱Ꮫ኱Ꮫ㝔⌮Ꮫ◊✲㝔ࡢᑓ௵ࡢᩍᤵཪࡣ෸ᩍᤵࢆࡶࡗ࡚඘࡚ࡿࠋ㸰 ࢭࣥࢱ࣮㛗ࡣ㸪໭ᾏ㐨኱Ꮫ኱Ꮫ㝔⌮Ꮫ◊✲㝔㛗㸦௨ୗࠕ◊✲㝔㛗ࠖ࡜࠸࠺ࠋ㸧ࡢ┘╩ࡢୗ࡟㸪ࢭࣥࢱ࣮ࡢᴗົࢆᤸ⌮ࡍࡿࠋ㸱 ࢭࣥࢱ࣮㛗ࡢ௵ᮇࡣ㸰ᖺ࡜ࡍࡿࠋ㸲 ࢭࣥࢱ࣮㛗ࡣ㸪෌௵ࡉࢀࡿࡇ࡜ࡀ࡛ࡁࡿࠋ㸳 ࢭࣥࢱ࣮㛗ࡢ㑅⪃ࡣ㸪◊✲㝔㛗ࡀ᥎⸀ࡍࡿೃ⿵⪅࠿ࡽ㸪⥲㛗ࡀ⾜࠺ࠋ㸴 ࢭࣥࢱ࣮㛗ೃ⿵⪅ࡢ㑅⪃࡟ࡘ࠸࡚ࡣ㸪◊✲㝔㛗ࡀู࡟ᐃࡵࡿࠋ 㸦㐠Ⴀጤဨ఍㸧➨㸴᮲ ࢭࣥࢱ࣮࡟㸪ࢭࣥࢱ࣮ࡢඹྠ฼⏝࡟㛵ࡍࡿ஦㡯ࡑࡢ௚ࡢࢭࣥࢱ࣮ࡢ㐠Ⴀ࡟㛵ࡍࡿ㔜せ஦㡯ࢆᑂ㆟ࡍࡿࡓࡵ㸪㐠Ⴀጤဨ఍ࢆ⨨ࡃࠋ㸰 㐠Ⴀጤဨ఍ࡢ⤌⧊ཬࡧ㐠Ⴀ࡟ࡘ࠸࡚ࡣ㸪◊✲㝔㛗ࡀู࡟ᐃࡵࡿࠋ 㸦㞧๎㸧➨㸵᮲ ࡇࡢつ⛬࡟ᐃࡵࡿࡶࡢࡢ࡯࠿㸪ࢭࣥࢱ࣮ࡢ⤌⧊ཬࡧ㐠Ⴀ࡟㛵ࡋᚲせ࡞஦㡯ࡣ㸪◊✲㝔ࡢᩍᤵ఍ࡢ㆟ࢆ⤒࡚㸪◊✲㝔㛗ࡀู࡟ᐃࡵࡿࠋ   㝃 ๎㸯 ࡇࡢつ⛬ࡣ㸪ᖹᡂ㸰㸮ᖺ㸯㸯᭶㸯᪥࠿ࡽ᪋⾜ࡍࡿࠋ㸰 ࡇࡢつ⛬ࡢ᪋⾜ᚋ㸪᭱ึ࡟௵࿨ࡉࢀࡿࢭࣥࢱ࣮㛗ࡢ௵ᮇࡣ㸪➨㸳᮲➨㸱㡯ࡢつᐃ࡟࠿࠿ࢃࡽࡎ㸪ᖹᡂ㸰㸰ᖺ㸱᭶㸱㸯᪥ࡲ࡛࡜ࡍࡿࠋ   㝃 ๎ ࡇࡢつ⛬ࡣ㸪ᖹᡂ㸰㸯ᖺ㸲᭶㸯᪥࠿ࡽ᪋⾜ࡍࡿࠋ   㝃 ๎ ࡇࡢつ⛬ࡣ㸪ᖹᡂ㸰㸵ᖺ㸲᭶㸯᪥࠿ࡽ᪋⾜ࡍࡿࠋ− 50 −

໭ᾏ㐨኱Ꮫ኱Ꮫ㝔⌮Ꮫ◊✲㝔㝃ᒓࢤࣀ࣒ࢲ࢖ࢼ࣑ࢡࢫ◊✲ࢭࣥࢱ࣮㐠Ⴀጤဨ఍ෆつ 㸦㊃᪨㸧
➨㸯᮲ ࡇࡢෆつࡣ㸪໭ᾏ㐨኱Ꮫ኱Ꮫ㝔⌮Ꮫ◊✲㝔㝃ᒓࢤࣀ࣒ࢲ࢖ࢼ࣑ࢡࢫ◊✲ࢭࣥࢱ࣮つ⛬㸦ᖹᡂ  ᖺᾏ኱㐩➨  ྕ㸧➨㸴᮲➨㸰㡯ࡢつᐃ࡟ᇶ࡙ࡁ㸪໭ᾏ㐨኱Ꮫ኱Ꮫ㝔⌮Ꮫ◊✲㝔㝃ᒓࢤࣀ࣒ࢲ࢖ࢼ࣑ࢡࢫ◊✲ࢭࣥࢱ࣮㐠Ⴀጤဨ఍㸦௨ୗࠕጤဨ఍ࠖ࡜࠸࠺ࠋ㸧ࡢ⤌⧊ཬࡧ㐠Ⴀ࡟ࡘ࠸࡚ᐃࡵࡿࡶࡢ࡜ࡍࡿࠋ 㸦ᑂ㆟஦㡯㸧➨㸰᮲ ጤဨ఍ࡣ㸪໭ᾏ㐨኱Ꮫ኱Ꮫ㝔⌮Ꮫ◊✲㝔㝃ᒓࢤࣀ࣒ࢲ࢖ࢼ࣑ࢡࢫ◊✲ࢭࣥࢱ࣮㸦ḟ᮲➨㸯㡯࡟࠾࠸࡚ࠕࢭࣥࢱ࣮ࠖ࡜࠸࠺ࠋ
㸧ࡢඹྠ฼⏝࡟㛵ࡍࡿ஦㡯ࡑࡢ௚ࡢࢭࣥࢱ࣮ࡢ㐠Ⴀ࡟㛵ࡍࡿ㔜せ஦㡯ࢆᑂ㆟ࡍࡿࠋ 㸦⤌⧊㸧
➨㸱᮲ ጤဨ఍ࡣ㸪ḟ࡟ᥖࡆࡿ⪅ࢆࡶࡗ࡚⤌⧊ࡍࡿࠋ ࢭࣥࢱ࣮㛗 ࢭࣥࢱ࣮ࡢᴗົࢆවົࡍࡿ໭ᾏ㐨኱Ꮫ኱Ꮫ㝔⌮Ꮫ◊✲㝔ࡢᩍᤵཬࡧ෸ᩍᤵ㸦ᅜ❧኱Ꮫἲே໭ᾏ㐨኱Ꮫ≉௵ᩍဨᑵᴗつ๎㸦ᖹᡂ㸯㸶ᖺᾏ኱㐩➨㸱㸳ྕ㸧➨㸱᮲➨㸰ྕ࡟ヱᙜࡍࡿ≉௵ᩍဨࢆྵࡴ㸧 ࡑࡢ௚⌮Ꮫ◊✲㝔㛗ࡀᚲせ࡜ㄆࡵࡓ⪅ ⱝᖸྡ㸰 ๓㡯➨㸱ྕࡢጤဨࡣ㸪⌮Ꮫ◊✲㝔㛗ࡀጤკࡍࡿࠋ 㸦௵ᮇ㸧
➨㸲᮲ ๓᮲➨㸯㡯➨㸱ྕࡢጤဨࡢ௵ᮇࡣ㸪㸯ᖺ࡜ࡍࡿࠋࡓࡔࡋ㸪⿵Ḟࡢጤဨࡢ௵ᮇࡣ㸪๓௵⪅ࡢṧ௵ᮇ㛫࡜ࡍࡿࠋ㸰 ๓㡯ࡢጤဨࡣ㸪෌௵ࡉࢀࡿࡇ࡜ࡀ࡛ࡁࡿࠋ 㸦ጤဨ㛗㸧
➨㸳᮲ ጤဨ఍࡟ጤဨ㛗ࢆ⨨ࡁ㸪⌮Ꮫ◊✲㝔ࡢ๪⌮Ꮫ◊✲㝔㛗ࡢ࠺ࡕ࠿ࡽ㸪⌮Ꮫ◊✲㝔㛗ࡀᣦྡࡍࡿ⪅ࢆࡶࡗ࡚඘࡚ࡿࠋ
㸦ጤဨ఍ࡢᣍ㞟ཬࡧ㆟㛗㸧➨㸴᮲ ጤဨ㛗ࡣ㸪ጤဨ఍ࢆᣍ㞟ࡋ㸪ࡑࡢ㆟㛗࡜࡞ࡿࠋ㸰 ጤဨ఍ࡣ㸪ጤဨ㛗ࡀᚲせ࡜ㄆࡵࡓሙྜ࡟ᣍ㞟ࡍࡿࡶࡢ࡜ࡍࡿࠋ㸱 ጤဨࡢ㸱ศࡢ㸯௨ୖࡢồࡵࡀ࠶ࡿሙྜ࡟ࡣ㸪ጤဨ㛗ࡣጤဨ఍ࢆᣍ㞟ࡋ࡞ࡅࢀࡤ࡞ࡽ࡞࠸ࠋ 㸦㆟஦㸧
➨㸵᮲ ጤဨ఍ࡣ㸪ጤဨࡢ㐣༙ᩘࡢฟᖍࡀ࡞ࡅࢀࡤ㆟஦ࢆ㛤ࡁ㸪㆟Ỵࡍࡿࡇ࡜ࡀ࡛ࡁ࡞࠸ࠋ㸰 ጤဨ఍ࡢ㆟஦ࡣ㸪ฟᖍጤဨࡢ㐣༙ᩘࢆࡶࡗ࡚Ỵࡍࡿࡶࡢ࡜ࡍࡿࠋ 㸦ጤဨ௨እࡢ⪅ࡢฟᖍ㸧➨㸶᮲ ጤဨ఍ࡀᚲせ࡜ㄆࡵࡓ࡜ࡁࡣ㸪ጤဨ఍࡟ጤဨ௨እࡢ⪅ࡢฟᖍࢆồࡵ㸪ㄝ᫂ཪࡣព
ぢࢆ⫈ࡃࡇ࡜ࡀ࡛ࡁࡿࠋ

− 51 − 㸦ᗢົ㸧➨㸷᮲ ጤဨ఍ࡢᗢົࡣ㸪⌮Ꮫ࣭⏕࿨⛉Ꮫ஦ົ㒊࡟࠾࠸࡚ฎ⌮ࡍࡿࠋ 㸦㞧๎㸧
➨  ᮲ ࡇࡢෆつ࡟ᐃࡵࡿࡶࡢࡢ࡯࠿㸪ጤဨ఍ࡢ㐠Ⴀ࡟㛵ࡋᚲせ࡞஦㡯ࡣ㸪ጤဨ఍ࡀู࡟ᐃࡵࡿࠋ   㝃 ๎ ࡇࡢෆつࡣ㸪ᖹᡂ㸰㸮ᖺ㸯㸰᭶㸲᪥࠿ࡽ᪋⾜ࡍࡿࠋ   㝃 ๎ࡇࡢෆつࡣ㸪ᖹᡂ㸰㸱ᖺ㸲᭶㸯᪥࠿ࡽ᪋⾜ࡍࡿࠋ− 52 −

໭ᾏ㐨኱Ꮫ኱Ꮫ㝔⌮Ꮫ◊✲㝔㝃ᒓࢤࣀ࣒ࢲ࢖ࢼ࣑ࢡࢫ◊✲ࢭࣥࢱ࣮㛗ೃ⿵⪅㑅⪃ෆつ


㸦㊃᪨㸧
➨㸯᮲ ࡇࡢෆつࡣ㸪໭ᾏ㐨኱Ꮫ኱Ꮫ㝔⌮Ꮫ◊✲㝔㝃ᒓࢤࣀ࣒ࢲ࢖ࢼ࣑ࢡࢫ◊✲ࢭࣥࢱ࣮つ⛬㸦ᖹᡂ㸰㸮ᖺᾏ኱㐩➨㸯㸳㸮ྕ㸧➨㸳᮲➨㸳㡯ࡢつᐃ࡟ᇶ࡙ࡁ㸪໭ᾏ㐨኱Ꮫ኱Ꮫ㝔⌮Ꮫ◊✲㝔㝃ᒓࢤࣀ࣒ࢲ࢖ࢼ࣑ࢡࢫ◊✲ࢭࣥࢱ࣮㛗ೃ⿵⪅㸦௨ୗࠕೃ⿵⪅ࠖ㸧ࡢ㑅⪃࡟㛵ࡋ㸪ᚲせ࡞஦㡯࡟ࡘ࠸࡚ᐃࡵࡿࡶࡢ࡜ࡍࡿࠋ
㸦ೃ⿵⪅ࡢ㑅⪃㸧➨㸰᮲ ೃ⿵⪅ࡢ㑅⪃ࡣ㸪ḟࡢ࠸ࡎࢀ࠿࡟ヱᙜࡍࡿሙྜ࡟⾜࠺ࠋ ࢭࣥࢱ࣮㛗ࡢ௵ᮇࡀ‶஢ࡍࡿ࡜ࡁࠋ ࢭࣥࢱ࣮㛗ࡀ㎡௵ࢆ⏦ࡋฟ࡚㸪⥲㛗ࡀㄆࡵࡓ࡜ࡁࠋ ࢭࣥࢱ࣮㛗ࡀḞࡅࡓ࡜ࡁࠋ
㸦ೃ⿵⪅ࡢ㑅⪃㸧➨㸱᮲ ೃ⿵⪅ࡣ㸪⌮Ꮫ◊✲㝔ࡢᑓ௵ࡢᩍᤵཬࡧ෸ᩍᤵࡢ࠺ࡕ࠿ࡽ㸪⌮Ꮫ◊✲㝔㛗ࡀᣦྡࡍࡿ⪅ࢆࡶࡗ࡚඘࡚ࡿࠋ
㸦ᩍᤵ఍ࡢሗ࿌㸧➨㸲᮲ ⌮Ꮫ◊✲㝔㛗ࡣ㸪๓᮲ࡢつᐃ࡟ࡼࡾᣦྡࡉࢀࡓೃ⿵⪅࡟ࡘ࠸࡚㸪໭ᾏ㐨኱Ꮫ኱Ꮫ㝔⌮Ꮫ◊✲㝔ᩍᤵ఍࡟ሗ࿌ࡍࡿࡶࡢ࡜ࡍࡿࠋ㝃 ๎ࡇࡢෆつࡣ㸪ᖹᡂ㸰㸮ᖺ㸯㸰᭶㸲᪥࠿ࡽ᪋⾜ࡍࡿࠋ㝃 ๎ࡇࡢෆつࡣ㸪ᖹᡂ㸰㸵ᖺ㸲᭶㸯᪥࠿ࡽ᪋⾜ࡍࡿࠋ− 53 −