ȹ൉࿾ત‫ݮ‬ᅔඦྎඏ࣢ѝмуѽభ౞ຊ࿼೮ɇ‫؞‬ɇȹ̵൸భઊ‫ލ‬ᆪѝмуѽ‫ސ‬ᅔଝњಊვࠫᔔଝ̵ȹ৶ȹ ៈȹ ଴ȹ ଧ̞ჵȹ ᇮȹ ྔȹ ੌȹȹȹȹɚʊʎɺʍʂɼȹɟɺʎʇɺȹʂʇȹʍʁɾȹɧɺʄɺʈȹɝʂʌʍʋʂɼʍȹʂʇȹɤɺʄʎɅȹɨʂʍɺȹɜʂʍʒɇȹ ‫؞‬ɇȹ̵ȹ ɢʇʏɺʌʂʏɾȹɺʇɽȹɞʇɽɺʇʀɾʋɾɽȹɬʉɾɼʂɾʌȹʈɿȹɟʋɾʌʁȹɰɺʍɾʋȹɦʈʅʅʎʌɼʌȹ ̵ȹȹɭɚɤɚɡɚɦɚɅȹɡʂɽɾʄʂȹɺʇɽȹɤɚɝɨɰɚɤɢɅȹɦʂʇɺȹȹȹȹȹȹȹȹȹȹȹȹȹȹȹȹȹ൉࿾൉޸ࢯ‫گ‬࿳ળ‫޸݌‬࿨ड़ࢂࡆᅃȹ ൊ 33 ߪൊ 2 ৺2011 ໧ 10 ॆȹ သੰ
Reprinted From
THE RESEARCH BULLETIN OF THE FACULTY OFEDUCATION AND WELFARE SCIENCE,OITA UNIVERSITY
Vol. 33, No. 2̜October 2011
OITA, JAPANᄢಽᄢቇᢎ⢒⑔␩⑼ቇㇱ⎇ⓥ♿ⷐ㧔Res. Bull. Fac. Educ.䋧Welf. Sci., Oita Univ.䋩

125ᄢಽᏒ⾐᧪ਛየ࿾඙ߦ߅ߌࠆ᳓↢േ‛⋧㧚Τ㧚̆᷆᳓↥⽴㘃ߦ߅ߌࠆᄖ᧪⒳ߣ⛘Ṍෂᗋ⒳̆
㜞 ự ⑲ ᮸㧖࡮ 㐷 ⣁ ⟤ ⩿㧖㧖‫ ⷐޣ‬ᣦ‫  ޤ‬ᄢಽᏒㄭ㇠ߩ㉿࿾࡮㉿ጊߢ޽ࠆ⾐᧪ਛየ࿾඙ߢ㧘᳓↰ߣߘߩ๟ㄝ᳓〝ߦ↢ᕷߔࠆ᷆᳓↥⽴㘃ߦߟ޿ߡ⺞ᩏߒ㧘ᄖ᧪⒳ࠬࠢࡒ࡝ࡦࠧࠟࠗ
㧔Pomacea canaliculata㧕ߣ⛘Ṍෂᗋ⒳ߩ↢ᕷ⁁ᴫࠍ᣿ࠄ߆ߦߒߚ‫ࡒࠢࠬޕ‬࡝ࡦࠧࠟࠗߩಽᏓߢߪ㧘ૐ࿾ߦ޽ࠆ᳓↰ߣ๟ㄝ᳓〝㧔ૐ࿾ᐔမㇱ᳓↰ 4 ඙ၞ㧕ߪߔߴߡଚ౉↰ߢ޽ࠅ㧘↢ᕷኒᐲߪ 1.5 ୘૕㧛ট㧔r0.4㧕ߢ޽ߞߚ‫⼱ޕ‬㑆ߦ޽ࠆ᳓↰ߣ๟ㄝ᳓〝㧔⼱㑆᫜⁁ㇱ᳓↰㧟඙ၞ㧕ߢߪ㕖ଚ౉↰ߢ޽ߞߚ‫ߚ߹ޕ‬㧘ૐ࿾ߣ⼱㑆ߩਛ㑆ߦ૏⟎ߔࠆ᫜⁁ߩ᳓↰㧔ਛ㑆᫜⁁ㇱ᳓↰ 1 ඙ၞ㧕ߢߪ㧘ଚ౉↰ߣ㕖ଚ౉↰߇⷗ࠄࠇߚ‫ޟޕ‬Ḱ⛘Ṍෂᗋ⒳‫ߦޠ‬ᜰቯߐࠇߡ޿ࠆࠝࠝ࠲࠾ࠪ
㧔 Cipangopaludina japonica 㧕㧘ࡑ࡞࠲࠾ࠪ㧔 Cipangopaludina chinensis

laeta㧕㧘ࡑࠪࠫࡒ㧔Corbicula leana㧕ࠍ᳓↰ߩ๟ㄝ᳓〝ߢ⏕⹺ߒߚ߇㧘↢ᕷ႐ᚲߪ㒢ቯ⊛ߢ޽ߞߚ‫ޕ‬‫  ޤ࠼࡯ࡢ࡯ࠠޣ‬᷆᳓↥⽴㘃 ࠬࠢࡒ࡝ࡦࠧࠟࠗ㧔Pomacea canaliculata㧕ᄖ᧪⒳ ⛘Ṍෂᗋ⒳ ᄢಽᏒㄭ㇠ߩ㉿࿾࡮㉿ጊߪߓ߼ߦᣣᧄߩ᳓↰ߪ㧘㉿㧔㉿࿾࡮㉿ጊ㧕ߦ޽ࠆㄘ᧛᥊ⷰࠍᒻᚑߔࠆⷐ⚛ߩ৻ߟߢ޽ࠅ㧔ⅣႺ⋭⥄ὼⅣႺዪ✬㧘2008㧕㧘ᄙߊߩ↢‛ߦߣߞߡ᳓↰߅ࠃ߮ߘߩ๟ㄝߩ᳓〝߇↢‛ߩ↢ᕷߦ㊀ⷐߥḨ࿾ߣߥߞߡ޿ࠆ㧔ᢧ⮮߶߆㧘1988㧧⷏ၔ㧘2001㧧Yamazaki et al., 2004㧧ጤ↰࡮⮮ጟ㧘2006㧕‫ޕ‬ߒ߆ߒ㧘ㄘᬺࠍᎼࠆ⁁ᴫߩᄌൻ㧔ᷫ෻߿⠹૞⠪㜞㦂ൻߦࠃࠆ᡼᫈᳓↰ߩჇട㧘࿛႐ᢛ஻ߦࠃࠆ᳓♽ᢿ⛘㧘ੇ↰ൻߦࠃࠆᓇ㗀㧕߆ࠄ㧘㉿࿾࡮㉿ጊߦ޽ࠆ᳓↰ࠍ↢ᕷ႐ᚲߣߒ㧘᥉ㅢߦ⷗ࠄࠇߡ޿ߚࡔ࠳ࠞ߿࠼࡚ࠫ࠙ߥߤߩ㝼㘃߿࠲ࠟࡔ߿ࠥࡦࠧࡠ࠙ߥߤߩ᣸⯻߇㧘ᣣᧄฦ࿾ߢ↢ᕷᢙࠍᷫߓ㧘࿾ၞߦࠃߞߡߪ⛘Ṍ߇ෂᗋߐࠇࠆ⒳ߦ᜼ߍࠄࠇߡ޿ࠆ㧔ⅣႺ⋭㧘2003㧕‫৻ޕ‬ᣇߢ㧘ᣣᧄߩ᳓↰߅ࠃ߮๟ㄝߩ᳓♽ⅣႺߦㆡᔕߒߚ࠙ࠪࠟࠛ࡞߿ࠕࡔ࡝ࠞࠩ࡝ࠟ࠾ߥߤᄖ᧪⒳߇ଚ౉࡮❥ᱺߒ㧘↢‛ᄙ᭽ᕈߩૐਅ߿ᣣᧄ࿕᦭⒳ߩ୘૕ᢙᷫዋࠍ߽ߚࠄߔሽ࿷ߣߥߞߡ޿ࠆ㧔⷏ේ߶߆㧘
2006㧧ᏒᎹ㧘2008㧕‫ޕ‬ࠬࠢࡒ࡝ࡦࠧࠟࠗ㧔 Pomacea canaliculata 㧕ߪධࠕࡔ࡝ࠞේ↥ߩ᷆᳓ߦ↢ᕷߔࠆᏎ⽴ߢ㧘ᐔᚑ 23 ᐕ 5 ᦬ 31 ᣣฃℂ㧖ߚ߆ߪ߹࡮߭ߢ߈ ᄢಽᄢቇᢎ⢒⑔␩⑼ቇㇱ↢‛ቇᢎቶ㧖㧖߆ߤࠊ߈࡮ߺߥ ᄢಽᄢቇᢎ⢒⑔␩⑼ቇㇱੱ㑆⑔␩⑼ቇ⺖⒟↢ᵴⅣႺ⑔␩ࠦ࡯ࠬⅣႺಽ㊁

126㜞 ự࡮㐷 ⣁1964 ᐕߦᣣᧄߦೋ߼ߡ߽ߚࠄߐࠇ㧘1981 ᐕߦ㘩↪ߣߒߡบḧ⚻↱ߢౣᐲ⒖౉ߐࠇ㧘ฦ࿾ߢ㙃ᱺ߇ⴕࠊࠇߚ‫ߩߘޕ‬ᓟ㧘㙃ᱺߩ஗ᱛߣߣ߽ߦ㊁↢ൻߒ㧘ᣣᧄߩ᷆᳓ⅣႺ㧘․ߦ᳓↰ߦㆡᔕߒ㧘⷏ᣣᧄࠍਛᔃߦᕆㅦߦ↢ᕷၞࠍ᜛ᄢߒߡ޿ࠆ㧔⩽⫱㧘1996㧧ᣣ㣔㧘1998㧧Wada, 2004㧕‫ޕ‬Ⓑ૞߳ߩⵍኂ߇ႎ๔ߐࠇ㧔ᐔ੗㧘1989㧕㧘‫ⷐޟ‬ᵈᗧᄖ᧪↢‛࡝ࠬ࠻‫ޠ‬㧔3㧕‫ޟ‬ㆬቯߩኻ⽎ߣߥࠄߥ޿߇ᵈᗧ༐⿠߇ᔅⷐߥᄖ᧪↢‛‫ߦޠ‬ᜰቯߐࠇߡ޿ࠆ㧔ⅣႺ⋭⥄ὼⅣႺዪ㧘2011㧕‫ޕ‬ᄢಽᏒ⾐᧪ਛየ࿾඙ߪ㧘Ꮢⴝ࿾߆ࠄࠊߕ߆ᢙࠠࡠࡔ࡯࠻࡞ߩ〒㔌ߦ޽ࠆ㓸⪭ߢ㧘㉿࿾࡮㉿ጊࠍᒻᚑߒߡ޿ࠆ‫ߩߎޕ‬࿾඙ߪᐔᚑ 20 ᐕᐲࠃࠅㄘᨋ᳓↥⋭ߩ‫ޟ‬ㄘ࿾࡮᳓࡮ⅣႺ଻ోะ਄ኻ╷‫ޠ‬ߩᵴേ࿾඙ߦᜰቯߐࠇ㧔ㄘᨋ᳓↥⋭㧘2010㧕㧘࿾඙ߦ↢ᕷߔࠆ↢‛ߩ଻⼔߽ⴕࠊࠇߡ޿ࠆ‫ޕ‬ᚒ‫ޘ‬ߪ㧘ਛየ࿾඙߆ࠄߩଐ㗬ࠍฃߌ㧘࿾඙ౝߩ᳓♽ߦ↢ᕷߔࠆ᳓↢േ‛⋧ߩ⺞ᩏߒߚ㧔㜞ự߶߆㧘
2011㧕‫ߩߎޕ‬࿾඙ߢ㧘ࠥࡦࠫࡏ࠲࡞ߥߤ㉿࿾࡮㉿ጊߦ⷗ࠄࠇࠆ᥉ㅢߦ⷗ࠄࠇࠆ⒳ߣࠦࠟ࠲ࡁࠥ
ࡦࠧࡠ࠙ߥߤ⛘Ṍ߇ෂᗋߐࠇࠆᏗዋ⒳ߩ↢ᕷࠍ⏕⹺ߒߚ߇㧘޿ߊߟ߆ߩᄖ᧪⒳ߩଚ౉߽␜ߐࠇ
ߚ‫ޕ‬ᄖ᧪⒳ߩ߁ߜࠬࠢࡒ࡝ࡦࠧࠟࠗߩ❥ᱺ߇ᦨ߽㗼⪺ߢ㧘ઁߩ᳓↢േ‛ߦਈ߃ࠆᓇ㗀ߪᄢ߈޿ߣᕁࠊࠇࠆ‫ߒ߆ߒޕ‬㧘ߎߩ࿾඙ߦ߅ߌࠆࠬࠢࡒ࡝ࡦࠧࠟࠗࠍߪߓ߼᷆᳓↥⽴㘃ߦߟ޿ߡߩ⹦⚦ߥ⎇ⓥߪߥ޿‫⎇ᧄߢߎߘޕ‬ⓥߪ㧘ᄢಽᏒ⾐᧪ਛየ࿾඙ߩ᳓↰ߣߘߩ๟ㄝ᳓〝ࠍኻ⽎ߦ᷆᳓↥⽴㘃ߦߟ޿ߡ⺞ߴ㧘ᄖ᧪⒳ߦࠃࠆ↢ᘒ♽߳ߩᓇ㗀ߦߟ޿ߡ⠨ኤߒߚ‫ޕ‬᧚ᢱߣᣇᴺ
ᄢಽᏒ⾐᧪ਛየ࿾඙ߢ㧘2010 ᐕ↰ᬀ߃ᓟ㧔6́7 ᦬㧕ߦ㧘᷆᳓↥⽴㘃ߩ↢ᕷ⁁ᴫߦߟ޿ߡ㧘ߔߴߡߩ᳓↰ߣߘߩ๟ㄝ᳓〝ࠍ⋡ⷞߢ⺞ᩏߒߚ‫ߪߡ޿ߟߦࠗࠟࠧࡦ࡝ࡒࠢࠬޕ‬㧘ᴡᎹ࡮᳓〝㧘᮸ᨋ߅ࠃ߮㆏〝ߢ㓒ߡࠄࠇߡ޿ࠆ㧤▎ᚲࠍ⺞ᩏ඙ၞ㧔A-H㧕ߣߒߡ⸳ቯߒ㧔࿑㧝㧕㧘ฦ⺞ᩏ඙ၞ߆ࠄ᛽಴ߒߚ᳓↰㧔5㧙10 ╩㧕ߢ㧘᳓↰㧝╩ߩో๟࿐ߩ⇜߆ࠄ㧞m એౝߦ↢ᕷߔࠆᲖ㜞 10 চએ਄ߩ୘૕ᢙࠍ⸘᷹ߒ㧘᳓↰ౝߩ↢ᕷኒᐲ㧔᳓↰ 1 ট޽ߚࠅߩᐔဋ୘૕ᢙrᮡḰ஍Ꮕ㧕ࠍ▚಴ߒߚ‫ߩઁޕ‬᷆᳓↥⽴㘃ߪ㧘࿾඙ౝߩ᳓↰ߣߘߩ๟ㄝ᳓〝ߢ㧘࠲ࡕ✂ߦࠃࠆយ޿ขࠅߢ᝝₪ߒ㧘⒳ࠍหቯߒ㧘↢ᕷ⁁ᴫࠍ⏕⹺ߒߚ‫ޕ‬⚿ᨐ‫ޣ‬ᄖ᧪⒳ࠬࠢࡒ࡝ࡦࠧࠟࠗߩ↢ᕷኒᐲ‫ޤ‬ᄢಽᏒ⾐᧪ਛየ࿾඙ߦ޽ࠆ᳓↰ߣߘߩ๟ㄝ᳓〝 8 ඙ၞߢ㧘ࠬࠢࡒ࡝ࡦࠧࠟࠗߩ↢ᕷ⁁ᴫߣ↢ᕷኒᐲࠍ⴫㧝ߦ␜ߔ‫ߩߎޕ‬࿾඙ߩ᳓↰ߣߘߩ๟ㄝ᳓〝ߪ㧘᳓↰߇૏⟎ߔࠆ࿾ᒻ߆ࠄૐ࿾ᐔမㇱ㧔࿑㧞㧕㧘ਛ㑆᫜⁁ㇱ㧔࿑㧟㧕㧘⼱㑆᫜⁁ㇱ㧔࿑㧠㧕ߩਃߟߦಽߌࠄࠇࠆ‫ߩߎޕ‬࿾඙ߢߪ㧘Ⓑ૞
ߦᔅⷐߥ᳓ߪห৻ߩ᳓♽ࠍ᳓Ḯߣߒ㧘ਛፉᎹߣḳᳰ߆ࠄ⼱㑆᫜⁁ㇱ㧘ਛ㑆᫜⁁ㇱࠍ⚻ߡૐ࿾ᐔမㇱߦᵹࠇߡ޿ࠆ‫↢ߩࠗࠟࠧࡦ࡝ࡒࠢࠬޕ‬ᕷ⁁ᴫߪߎߩਃߟߢ⇣ߥߞߡ޿ߚ‫ޕ‬ૐ࿾ᐔမㇱߦ޽ࠆ᳓↰㧔⺞ᩏ඙ၞ A-D㧕ߪ㧘ૐ࿾ߢ㜞ૐᏅߩዋߥ޿ᐔမߥ႐ᚲߦ޽ࠅ㧘᳓↰ߩᄢㇱಽࠍභ߼ߡ޿ࠆ‫ޕ‬ૐ࿾ᐔမㇱߦ߅ߌࠆ᳓↰ౝߩᐔဋ↢ᕷኒᐲߪ 1.5 ୘૕㧛ট㧔r0.4㧕ߢ㧔࿑㧡㧕㧘⺞ᩏ඙ၞ㑆ߢߩ㆑޿ߪ⹺߼ࠄࠇߥ߆ߞߚ‫ޕ‬ૐ࿾ᐔမㇱ᳓↰ߢߪ㧘⺞ᩏ඙ၞౝ߅ࠃ߮⺞ᩏ඙ၞએᄖߢ߽㧘ߔߴߡߩ᳓↰ߣ๟ㄝ᳓〝ߢ୘૕ߩଚ౉߇ⷰኤߐࠇ㧘ଚ౉↰ߢ޽ߞߚ‫৻ޕ‬ㇱߩ᳓↰ߢ㧘⧣ߩ㘩ኂ߽⹺߼ࠄࠇߚ㧔࿑㧢㧕‫ޕ‬ਛ㑆᫜⁁ㇱ᳓↰㧔⺞ᩏ඙ၞ E㧕ߪ㧘㓸⪭߇޽ࠆ㜞บߩᢳ㕙ߦ૏⟎ߒ㧘ૐ࿾ᐔမㇱߩ⺞ᩏ඙ၞ D ߣ㓞ធߒߡ޿ࠆ‫ޕ‬ૐ࿾ᐔမㇱߣ㓞ធߔࠆ 3 ╩ߩ᳓↰ߢ㧘ኒᐲߪ㜞ߊߥ޿127ᄢಽᏒ⾐᧪ਛየ࿾඙ߦ߅ߌࠆ᷆᳓↥⽴㘃߽ߩߩ㧘୘૕ߩ↢ᕷ߇⏕⹺ߐࠇߚ‫ߒ߆ߒޕ‬㧘ࠃࠅ㜞޿૏⟎ߦ޽ࠆ᳓↰ 7 ╩ߢߪ㧘୘૕ߩଚ౉ߪ⏕⹺ߐࠇߥ߆ߞߚ‫⼱ޕ‬㑆᫜⁁ㇱ㧔⺞ᩏ඙ၞ F-H㧕ߩ᳓↰ߪ㧘࿾඙ౝߦ޽ࠆ᮸ᨋ࿾ߩ⼱㑆ߦ૏⟎ߒ㧘ઁߩ඙ၞߣ㔌ࠇ㓒⛘ߐࠇߡ޿ࠆ‫ߪߢߎߎޕ‬㧘ߔߴߡߩ᳓↰ߢࠬࠢࡒ࡝ࡦࠧࠟࠗߩ↢ᕷߪ⏕⹺ߐࠇߕ㧘㕖ଚ౉↰ߢ޽ߞߚ‫ޕ‬⾐᧪Ꮉ

NBD

ᄢಽ Ꮢ⾐᧪ਛየ࿾඙㓸⪭ਛየḳ ᳰAHECਛፉᎹ

GFૐ࿾ᐔမㇱ᳓↰ਛ㑆᫜⁁ㇱ᳓↰100m᮸ᨋ࿾䊊䉴↰⼱㑆᫜⁁ㇱ᳓↰ਥⷐ㆏〝᳓♽࿑㧝 ᄢಽᏒ⾐᧪ਛየ࿾඙ߩ᭎⇛࿑㧚࿑ਛߦ⺞ᩏ඙ၞ㧔A-H㧕ߣ᳓↰ߩ⁁ᘒࠍ␜ߔ㧚࿑㧞㧚ૐ࿾ᐔမㇱ᳓↰㧔⺞ᩏ඙ၞA㧕ߩ౮⌀㧚ᅏߪ㓸⪭߇޽ࠆ㜞บ㧚ߔߴߡߩ᳓↰ߣ๟ㄝ᳓〝ߦࠬࠢࡒ࡝ࡦࠧࠟࠗ߇ଚ౉㧚࿑㧟㧚ਛ㑆᫜⁁ㇱ᳓↰㧔⺞ ࿑㧠㧚⼱㑆᫜⁁ㇱ᳓↰㧔⺞ᩏ඙ၞE㧕ߩ౮⌀㧚ᅏߪ㓸 ᩏ඙ၞG㧕ߩ౮⌀㧚ਔ஥ߪ⪭ߦ㓞ធߔࠆ᮸ᨋ࿾ߢ㧘ᢳ ᮸ᨋ࿾ߣߥߞߡ޿ࠆ㧚᳓↰㕙ߦ᳓↰߇᫜⁁ߦ૏⟎ߔࠆ㧚 ߪ᫜⁁ߢ㧘ࠬࠢࡒ࡝ࡦࠧࠟࠬࠢࡒ࡝ࡦࠧࠟࠗߪᚻ೨ߩ ࠗߩଚ౉ߪ⷗ࠄࠇߥ޿㧚᳓↰㧔⍫ශ㧕ߦଚ౉㧚࿑㧡㧚ૐ࿾ᐔမㇱ㧔⺞ᩏ඙ၞD㧕ߢࠬࠢࡒ࡝ࡦࠧࠟࠗߩ↢ᕷኒᐲ߇㜞޿᳓↰ߩ౮⌀㧚࿑㧢㧚ૐ࿾ᐔမㇱ᳓↰㧔⺞ᩏ඙ၞD㧕ߢࠬࠢࡒ࡝ࡦࠧࠟࠗߦࠃࠆ㘩ኂ߇⷗ࠄࠇߚ᳓↰ߩ౮⌀㧚࿑㧣㧚ࠝࠝ࠲࠾ࠪߩ౮⌀㧚⼱㑆᫜⁁ㇱ᳓↰㧔⺞ᩏ඙ၞH㧕ߩ๟ㄝ᳓〝ߦ↢ᕷ㧚128㜞 ự࡮㐷 ⣁⴫㧝 ᄢಽᏒ⾐᧪ਛየ࿾඙ߩ᳓↰ߦ߅ߌࠆࠬࠢࡒ࡝ࡦࠧࠟࠗߩ↢ᕷ⁁ᴫૐ࿾ᐔမㇱਛ㑆᫜⁁ㇱ⼱㑆᫜⁁ㇱ⺞ᩏ඙ၞA↢ᕷ⏕⹺᳓↰ᢙ䋯⺞ᩏ᳓↰ᢙ㪍㪆㪍⺞ᩏ᳓↰㕙Ⓧ䋨䋛䋩㪈㪎㪏㪐↢ᕷᢙ䋨✚ᢙ䋩㪉㪋㪈㪊↢ᕷኒᐲ䋨୘૕ᢙ䋯䋛䋬ᐔဋ㫧ᮡḰ஍Ꮕ䋩 㪈㪅㪋㫧㪈㪅㪈⺞ᩏ඙ၞE↢ᕷ⏕⹺᳓↰ᢙ䋯⺞ᩏ᳓↰ᢙ㪊㪆㪈㪇⺞ᩏ᳓↰㕙Ⓧ䋨䋛䋩㪊㪎㪎↢ᕷᢙ䋨✚ᢙ䋩㪊㪇㪌↢ᕷኒᐲ䋨୘૕ᢙ䋯䋛䋬ᐔဋ㫧ᮡḰ஍Ꮕ䋩 㪈㪅㪇㫧㪇㪅㪐⺞ᩏ඙ၞF↢ᕷ⏕⹺᳓↰ᢙ䋯⺞ᩏ᳓↰ᢙ㪇㪆㪏⺞ᩏ᳓↰㕙Ⓧ䋨䋛䋩㪏㪈㪏↢ᕷᢙ䋨✚ᢙ䋩㪇↢ᕷኒᐲ䋨୘૕ᢙ䋯䋛䋬ᐔဋ㫧ᮡḰ஍Ꮕ䋩㪇B㪏㪆㪏㪉㪊㪊㪊㪉㪍㪏㪈㪈㪅㪉㫧㪈㪅㪎C㪍㪆㪍㪈㪊㪈㪋㪉㪊㪍㪐㪈㪅㪏㫧㪈㪅㪋G㪇㪆㪌㪎㪎㪈㪇㪇H㪇㪆㪐㪈㪈㪎㪍㪇㪇

D㪍㪆㪍㪈㪊㪋㪌㪉㪋㪎㪇㪉㪅㪉㫧㪉㪅㪏‫ߩઁޣ‬᷆᳓↥Ꮞ⽴‫ޤ‬ᄖ᧪⒳ࠨࠞࡑࠠࠟࠗ㧔Physa acuta㧕ߪ㧘ૐ࿾ᐔမㇱߩ᳓↰㧔⺞ᩏ඙ၞ A㧙D㧕ߢⷰኤߐࠇߚ‫ޕ‬․ߦࡂࠬ↰ߦ㓞ធߔࠆ඙ၞ C ߢߪᄙߊߩ᳓↰߿๟ㄝ᳓〝ߢ⹺߼ࠄࠇߚ‫ޕ‬ਛ㑆᫜⁁ㇱ߿⼱㑆᫜⁁ㇱߩ᳓↰ߢߪ⹺߼ࠄࠇߥ߆ߞߚ‫ޕ‬
࿷᧪⒳ࠝࠝ࠲࠾ࠪ㧔Cipangopaludina japonica㧕㧔࿑㧣㧕ߪ㧘ḳᳰߦ㓞ធߔࠆ⼱㑆᫜⁁ㇱ Hߩ᳓↰ߩ๟ㄝ᳓〝ߢߩߺⷰኤߐࠇ㧘᳓↰ౝ߿ḳᳰߢߪ⷗ࠄࠇߥ߆ߞߚ‫ޕ‬࿷᧪⒳ࡑ࡞࠲࠾ࠪ
㧔Cipangopaludina chinensis laeta㧕ߪ㧘ࡂࠬ↰ߦ㓞ធߔࠆૐ࿾ᐔမㇱ C ߣ⼱㑆᫜⁁ㇱ H ߩ᳓↰ߩ๟ㄝ᳓〝ߣࡂࠬ↰ౝߢⷰኤߐࠇ㧘᳓↰ౝߢߪోߊⷰኤߐࠇߥ߆ߞߚ‫ޕ‬࿷᧪⒳ࡑࠪࠫࡒ
㧔Corbicula leana㧕ߪ㧘⼱㑆᫜⁁ㇱ H ߩ᳓↰ߩ๟ㄝ᳓〝ߣૐ࿾ᐔမㇱ C ߩ᳓↰ߩ๟ㄝ᳓〝ߢⷰኤߐࠇ㧘᳓↰ౝߢߪⷰኤߐࠇߥ߆ߞߚ‫ޕ‬࿷᧪⒳ࡅࡔࡕࡁࠕ࡜ࠟࠗ㧔Austropeplea ollula㧕ߪ㧘ૐ࿾ᐔမㇱ㧔⺞ᩏ඙ၞ C㧕߿ਛ㑆᫜⁁ㇱߩ৻ㇱߩ᳓↰߅ࠃ߮๟ㄝ᳓〝ߢⷰኤߐࠇߚ‫ޕ‬࿷᧪⒳ࠞ
ࡢ࠾࠽㧔Semisulcospira libertina㧕ߪ㧘࿾඙ౝߩ᳓↰๟ㄝ᳓〝ߢᲧセ⊛ᄙߊ⹺߼ࠄࠇ㧘᳓↰ౝߢߪኒᐲߪૐ޿߽ߩߩᄙߊߩ႐ᚲ㧔⺞ᩏ඙ၞ A㧘B㧘C㧘E㧘F㧘G㧕ߢ⏕⹺ߐࠇߚ‫ޕ‬⠨ኤᧄ⎇ⓥߪ㧘ᄢಽᏒ⾐᧪ਛየ࿾඙ߩ᳓↰߅ࠃ߮๟ㄝ᳓〝ߦ↢ᕷߔࠆ᷆᳓↥⽴㘃ߩ⺞ᩏࠍⴕ޿㧘
ᄖ᧪⒳ࠬࠢࡒ࡝ࡦࠧࠟࠗ߇㧘ૐ࿾ߦ޽ࠅ㜞ૐᏅߩዋߥ޿᳓↰ߢߪߔߴߡߦଚ౉ߒߡ޿ࠆߎߣ㧘⼱㑆ߢ㓒⛘ߐࠇߡ޿ࠆ᫜⁁᳓↰ߢߪ㕖ଚ౉ߢ޽ࠆߎߣ㧘ߘߩ㑆ߦ૏⟎ߔࠆਛ㑆᫜⁁ㇱ᳓↰ߢߪଚ౉↰ߣ㕖ଚ౉↰ߩႺ⇇߇޽ࠆߎߣࠍ␜ߒ㧘᳓↰߇૏⟎ߔࠆ⁁ᘒߢಽᏓߦ㆑޿߇޽ࠆߎߣࠍ᣿ࠄ߆ߦߒߚ‫ߚ߹ޕ‬㧘᳓↰๟ㄝ᳓〝ߢ⛘Ṍෂᗋ⒳ߢ޽ࠆࠝࠝ࠲࠾ࠪ㧘ࡑ࡞࠲࠾ࠪ㧘ࡑࠪࠫࡒߩ↢ᕷࠍ⏕⹺ߒߚ‫ޕ‬ࠬࠢࡒ࡝ࡦࠧࠟࠗߪ㧘1981 ᐕߩౣዉ౉એ᧪਻Ꮊߦ߅޿ߡ߽ᐢߊಽᏓߒ㧘ᄢಽ⋵ౝߢ߽ಽᏓࠍ᜛ᄢߒߡ޿ࠆߣ⠨߃ࠄࠇࠆ‫ޕ‬ᄢಽᏒ⾐᧪ਛየ࿾඙ߢߩࠬࠢࡒ࡝ࡦࠧࠟࠗߩ↢ᕷኒᐲߪ㧘ઁߩ࿾ᄢಽᏒ⾐᧪ਛየ࿾඙ߦ߅ߌࠆ᷆᳓↥⽴㘃129ၞߢߩ⺞ᩏߣᲧセߒߡૐ޿୯ߢߪߥߊ㧔Ꮢἑ㧘2002㧕㧘❥ᱺ߇ㅴⴕߒߡ޿ࠆߎߣ߇␜ߐࠇߚ‫ޕ‬ᾢᧄ⋵⷏วᔒ↸ߢߩಽᏓ⺞ᩏߢߪ㧘ਐ㧘ᴡᎹᴪ޿ߥߤ᳓↰ߩ޽ࠆ႐ᚲߦࠃࠅಽᏓኒᐲ߇⇣ߥࠆߎߣ߇᣿ࠄ߆ߦߐࠇߡ޿ࠆ㧔Ꮢἑ㧘2002㧕‫⎇ᧄޕ‬ⓥߦ߅޿ߡ߽ಽᏓߩ௑ะߪห᭽ߢ޽ࠆߎߣ߇
␜ߐࠇߚ߇㧘⼱㑆ߩ᫜↰ߦߪଚ౉ߒߡ޿޿ߥ޿ߎߣ߇ಽ߆ߞߚ‫ߚ߹ޕ‬㧘ଚ౉↰ߣ㕖ଚ౉↰ߩႺ⇇߇޽ࠆਛ㑆᫜⁁᳓↰ߩࠃ߁ߥ႐ᚲߪ㧘ಽᏓ᜛ᄢߩㄝ✼ㇱߣ⠨߃ࠄࠇߚ‫ߩߎޕ‬࿾඙ߢ␜ߐࠇࠆ
ࠃ߁ߥಽᏓߩ․ᓽ߇㧘ᄢಽᐔ㊁߿ᄢಽ⋵ౝߢ౒ㅢߒߡ޿ࠆ߆ߤ߁߆ߪਇ᣿ߢ޽ࠆ‫ޕ‬ࠝࠝ࠲࠾ࠪ㧘ࡑ࡞࠲࠾ࠪ㧘ࡑࠪࠫࡒߩ 3 ⒳ߪ㧘↢ᕷⅣႺߩᖡൻ߆ࠄ୘૕ᢙࠍᷫߓ㧘‫ޟ‬Ḱ⛘Ṍෂᗋ⒳‫ߦޠ‬ᜰቯߐࠇߚ㧔ⅣႺ⋭⥄ὼⅣႺዪ㧘2008㧕‫⎇ᧄޕ‬ⓥߢ㧘ߎߩ࿾඙ߦߎࠇࠄ 3 ⒳߇↢ᕷߒߡ޿ࠆߎߣ߇⏕⹺ߐࠇߚ߇㧘ḳᳰ㓞ធߔࠆ᳓↰๟ㄝ᳓〝ߣࡂࠬ↰ߦ㓞ធߔࠆ᳓↰๟ㄝ᳓〝ߣߦ㒢ቯߐࠇߡ޿ࠆߎߣ߇ಽ߆ߞߚ‫⛘ࠄࠇߎޕ‬Ṍෂᗋ⒳߇↢ᕷߔࠆ᳓〝ߪ㧘౻ቄߦ߅޿ߡ߽᳓߇Ḗ߃ࠄࠇ㧘ቯᦼ⊛ߦ⛽ᜬ▤ℂߐࠇߡ޿ࠆ႐ᚲߢ޽ࠆ‫‛↢ޕ‬ᄙ᭽ᕈ⛽ᜬߩ㕙ߢ㧘ࡂࠬ↰ߦߪ᳓↰ߣߪ⇣ߥࠆᓎഀ߇␜ߐࠇߡ޿ࠆ㧔ጤ↰࡮⮮ጟ㧘2006㧕‫ߩߎޕ‬࿾඙ߩ᳓↰ⅣႺ߇ࡂࠬ↰߿ḳᳰߦ㓞ធߒߡ޿ࠆߎߣߢ㧘Ꮧዋߣߥߞߚ࿷᧪⒳߇ᄖ᧪⒳ߣ౒ሽߢ߈㧘↢‛ᄙ᭽ᕈߩ⛽ᜬߦ⦟޿ⅣႺ
ࠍ߽ߚࠄߒߡ޿ࠆߣᕁࠊࠇࠆ‫ޕ‬Ḱ⛘Ṍෂᗋ⒳ࠝࠝ࠲࠾ࠪ㧘ࡑ࡞࠲࠾ࠪ㧘ࡑࠪࠫࡒߩ 3 ⒳߇↢ᕷߔࠆ⼱㑆⁁᫜↰᳓↰๟ㄝߩ᳓〝ߢߪࠬࠢࡒ࡝ࡦࠧࠟࠗߩଚ౉ߪߥ޿߇㧘ࡑ࡞࠲࠾ࠪߣࡑࠪࠫࡒ߇↢ᕷߔࠆૐ࿾ㇱߩࡂࠬ↰ߦ㓞ធߔࠆ᳓↰๟ㄝ᳓〝ߢߪᐢ▸࿐ߦࠊߚࠆଚ౉߇⷗ࠄࠇߡ޿ࠆ‫ޕ‬᳓↰ⅣႺߦㆡᔕߒߚᄖ᧪⒳ࠍ
㚟ㅙߔࠆߎߣߪኈᤃߢߪߥ޿㧔⩽⫱㧘1996㧧ᣣ㣔㧘1998㧧Wada, 2004㧕‫┹ޕ‬วߩ⚿ᨐ㧘⛘Ṍ߇ෂᗋߐࠇࠆ⒳߇㚟ㅙߐࠇࠆ ᔨ߽޽ࠆ߇㧘⃻⁁߇⛽ᜬߐࠇࠇ߫↢ሽߩน⢻ᕈ߽ᱷߐࠇߡ޿ࠆ‫ޕ‬⃻࿷଻ߚࠇߡ޿ࠆ↢‛ᄙ᭽ᕈߩ᳓Ḱࠍ⛽ᜬߔࠆߦߪ㧘᳓↰㧘ࡂࠬ↰㧘ߚ߼ᳰߩ⛽ᜬ▤ℂ߇ᔅⷐਇนᰳߢ㧘ߎߩ࿾඙ߩ૑᳃ߦࠃࠆ଻ో߳ߩᗧ⼂ߩ㜞ߐߦࠃߞߡᡰ߃ࠄࠇߡ޿ࠆ‫ޕ‬‫ޟ‬ㄘ࿾࡮᳓࡮ⅣႺ଻ోะ਄ኻ╷‫⽷ߥ߁ࠃߩޠ‬᡽⊛ᡰេ߽ᔅⷐߢ޽ࠆ‫ޕ‬⻢ㄉᄢಽᏒ⾐᧪ਛየ࿾඙ਛፉᎹߦ߅ߌࠆ↢‛⺞ᩏߦ޽ߚࠅ㧘࿾඙ߩ૑᳃ߩᣇ‫⎇ᧄߪߦޘ‬ⓥ߳ߩߏ
ℂ⸃ߣߏදജࠍ⾦ࠅ㧘⻯ࠎߢᗵ⻢ߩᗧࠍ⴫ߔࠆ‫ޕ‬ෳ⠨ᢥ₂1㧕ⅣႺ⋭⥄ὼⅣႺዪ✬㧔2008㧕㧦 ㉿࿾㉿ጊ଻ోౣ↢⸘↹૞ᚑߩᚻᒁ߈㧘p1-46㧚
2㧕ᢧ⮮ᙗᴦ࡮ ㊁ୃ࡮ዊᴰ㗼㓶㧔1988㧕㧦᷆᳓㝼ߩ᳓↰๟ㄝߦ߅ߌࠆ৻ᤨ⊛᳓ၞ߳ߩଚ౉ߣ↥ෆ㧚ᣣᧄ↢ᘒቇળ⹹㧘╙ 38 Ꮞ㧘35-47㧚
3㧕⷏ၔᵗ㧔2001㧕㧦ፉᩮ⋵ߩ᳓↰ߣḳ߼ᳰߦ߅ߌࠆ᳓↢᣸⯻ߩቄ▵⊛ᶖ㐳ߣ⒖േ㧚ᣣᧄ↢ᘒቇળ⹹㧘╙ 51 Ꮞ㧘1-11㧚
4㧕Yamazaki, M., Hamada, Y., Kamimoto, N., Momii, T., and Kimura, M. (2004): Habitats ofaquatic organisms in a paddy field during flooding period. Edaphologia, No.74, 1-10.
5㧕ጤ↰᮸࡮⮮ጟᱜඳ㧔2006㧕㧦ࡂࠬ↰ߣࠗࡀ↰ߦ߅ߌࠆ౻ᦼḖ᳓ߩ᦭ή߇૞‛ᚑ㐳ᦼߩ᳓↢േ‛⋧ߦਈ߃ࠆᓇ㗀㧚଻ో↢ᘒቇ⎇ⓥ㧘╙ 11 Ꮞ㧘94-104㧚
6㧕ⅣႺ⋭㧔2003㧕㧦ᡷ⸓࡮ᣣᧄߩ⛘Ṍߩ߅ߘࠇߩ޽ࠆ㊁↢↢‛ 4㨇ᴁ᳓࡮᷆᳓㝼㘃㨉㧚⥄ὼⅣႺ⎇ⓥ࠮ࡦ࠲࡯㧚7㧕⷏ේ᣹๋࡮⧛ㇱᴦ♿࡮㣐⼱޿ߕߺ㧔2006㧕㧦᳓↰ߦ↢ᕷߔࠆࠥࡦࠧࡠ࠙㘃ߩ⃻⁁ߣ଻ో㧚଻ో↢

130㜞 ự࡮㐷 ⣁ᘒቇ⎇ⓥ㧘╙ 11 Ꮞ㧘143-157㧚
8㧕ᏒᎹᙗᐔ㧔2008㧕㧦㉿࿾ߩ᳓↢᣸⯻ߩ⃻⁁ߣ଻ో㧚╙ 19 Ꮞ㧘47-50㧚
9㧕⩽⫱ା৻㇢㧔1996㧕㧦ࠬࠢࡒ࡝ࡦࠧࠟࠗߩ↢ᘒߣ㒐㒰㧚ᬀ‛㒐∉㧘╙ 50 Ꮞ㧘211-217㧚
10㧕ᣣ㣔৻㓷㧔1998㧕㧦᳓↰ߦ߅ߌࠆ↢‛ᄙ᭽ᕈ଻ోߣⅣႺୃᓳဳㄘᴺ㧚ᣣᧄ↢ᘒቇળ⹹㧘╙ 48 Ꮞ㧘
167-178㧚
11㧕Wada, T. (2004): Strategies for controlling the apple snail Pomacea canaliculata (Lamarck)(Gastropoda: Ampullariidae) in Japanese direct-sown paddy fields. JARQ, No.38, 75-80.
12㧕ᐔ੗೰ᄦ㧔1989㧕㧦ࠬࠢࡒ࡝ࡦࠧࠟࠗߩ⊒↢ߣಽᏓ᜛ᄢ㧚ᬀ‛㒐∉㧘╙ 43 Ꮞ㧘498-501㧚
13㧕ⅣႺ⋭⥄ὼⅣႺዪ HP㧔2011㧕㧦‫ⷐޟ‬ᵈᗧᄖ᧪↢‛࡝ࠬ࠻‫ޠ‬㧔3㧕ㆬቯߩኻ⽎ߣߥࠄߥ޿߇ᵈᗧ༐⿠߇ᔅⷐߥᄖ᧪↢‛㧚http://www.env.go.jp/nature/intro/1outline/caution/index.html.
14㧕ㄘᨋ᳓↥⋭✬㧔2010㧕㧦ㄘ࿾࡮᳓࡮ⅣႺ଻ోะ਄ߩߚ߼ߦ㧘p1-36㧚
15㧕㜞ự⑲᮸࡮ᄢୖ㋕਽࡮㤥ᧁ⟤㉿㧔2011㧕㧦ᄢಽᏒ⾐᧪ਛየ࿾඙ߦ߅ߌࠆ᳓↢േ‛⋧㧚Σ㧚̆᥉ㅢߦ⷗ࠄࠇࠆ⒳ߣ⛘Ṍෂᗋ⒳̆㧚ᄢಽᄢቇᢎ⢒⑔␩⑼ቇㇱ⎇ⓥ♿ⷐ㧘╙ 33 Ꮞ㧘╙ 1 ภ㧘51-58㧚
16㧕Ꮢἑస਽㧔2002㧕
㧦ࠬࠢࡒ࡝ࡦࠧࠟࠗߩಽᏓ᜛ᄢߦ෸߷ߔ࿾ᒻߩᓇ㗀㧚਻Ꮊㄘᬺ⎇ⓥ㧚╙ 64 ภ㧘
84㧚
17㧕ⅣႺ⋭⥄ὼⅣႺዪ HP㧔2008㧕㧦ⅣႺ⋭↢‛ᄙ᭽ᕈᖱႎࠪࠬ࠹ࡓ㧘⛘Ṍෂᗋ⒳ᖱႎᬌ⚝㧘ᡷ⸓࡟࠶࠼࡝ࠬ࠻㧚http://www.biodic.go.jp/rdb/rdb_f.html.

Aquatic Fauna in the Nakao District in Kaku, Oita City. Τ.
̆ Invasive and Endangered Species of Fresh Water Molluscs ̆
TAKAHAMA, Hideki and KADOWAKI, Mina

Abstract
We investigated fresh water molluscs in the Nakao district in Kaku,
Oita City.

The density of apple snails (Pomacea canaliculata) in thesurrounding rice fields was estimated and the presence of three
endangered species was confirmed.

All paddy fields situated in the flatand low area were invaded by the snails and the average density was 1.5/
ট (r0.4), but they had not invaded the terraced paddy fields.

In theintermediate area between them, the boundary of invaded andnon-invaded rice fields was observed.
(Cipangopaludina

Three endangered species

japonica, Cipangopaludina chinensis laeta, andCorbicula leana) were confirmed only in the water ways in this district.

䇼Key words‫  ޤ‬fresh water shells, apple snail (Pomacea canaliculata),
invasive species, endangered species, Satochi-Satoyama near the urbanarea