ZOOTAXA
1418

Checklist of catfishes, recent and fossil(Osteichthyes: Siluriformes),and catalogue of siluriform primary types
CARL J. FERRARIS, JR.

Magnolia Press
Auckland, New ZealandCARL J. FERRARIS, JR.
Checklist of catfishes, recent and fossil (Osteichthyes: Siluriformes), and catalogue of siluriform
primary types(Zootaxa 1418)
628 pp.; 30 cm.
8 March 2007
ISBN 978-1-86977-058-7 (hardback)ISBN 978-1-86977-059-4 (Online edition)

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Zootaxa 1418: 1–628 (2007)www.mapress.com / zootaxa/

ISSN 1175-5326 (print edition)

Copyright © 2007 · Magnolia Press

ISSN 1175-5334 (online edition)ZOOTAXA

Checklist of catfishes, recent and fossil (Osteichthyes: Siluriformes),and catalogue of siluriform primary types
CARL J. FERRARIS, JR.
2944 NE Couch Street, Portland, Oregon, 97232, U.S.A. E-mail: carlferraris@comcast.net
Research Associate, National Museum of Natural History, Smithsonian Institution, Washington
Research Associate, American Museum of Natural History, New York
Research Associate and Honorary Fellow, California Academy of Sciences, San Francisco

Table of contents
Abstract ..................................................................................................................................................................................................... 4
Introduction .......................................................................................................................................................................... 4
Methods and conventions..................................................................................................................................................... 6
Acknowledgments.............................................................................................................................................................. 10
Checklist.............................................................................................................................................................................. 11
Siluriformes................................................................................................................................................................. 11
Akysidae...................................................................................................................................................................... 12
Amblycipitidae............................................................................................................................................................ 17
Amphiliidae................................................................................................................................................................. 20
Anchariidae ................................................................................................................................................................. 29
Andinichthyidae .......................................................................................................................................................... 30
Ariidae......................................................................................................................................................................... 30
Aspredinidae ............................................................................................................................................................... 57
Astroblepidae .............................................................................................................................................................. 63
Auchenipteridae .......................................................................................................................................................... 68
Austroglanididae ......................................................................................................................................................... 81
Bagridae ...................................................................................................................................................................... 81
Callichthyidae ........................................................................................................................................................... 108
Cetopsidae................................................................................................................................................................. 130
Chacidae.................................................................................................................................................................... 135
Clariidae .................................................................................................................................................................... 136
Claroteidae ................................................................................................................................................................ 153
Cranoglanididae ........................................................................................................................................................ 164
Diplomystidae ........................................................................................................................................................... 165
Doradidae .................................................................................................................................................................. 166
Heptapteridae ............................................................................................................................................................ 180
Hypsidoridae ............................................................................................................................................................. 203
Ictaluridae.................................................................................................................................................................. 203
Lacantuniidae ............................................................................................................................................................ 216
Loricariidae ............................................................................................................................................................... 216
Malapteruridae .......................................................................................................................................................... 300
Mochokidae............................................................................................................................................................... 303
Nematogenyiidae ...................................................................................................................................................... 323
Pangasiidae................................................................................................................................................................ 324

Accepted by L. Page: 4 Oct. 2006; published: 8 Mar. 2007

3

Pimelodidae............................................................................................................................................................... 329
Plotosidae .................................................................................................................................................................. 345
Pseudopimelodidae ................................................................................................................................................... 352
Schilbidae.................................................................................................................................................................. 356
Scoloplacidae ............................................................................................................................................................ 366
Siluridae .................................................................................................................................................................... 367
Sisoridae.................................................................................................................................................................... 382
Trichomycteridae ...................................................................................................................................................... 402
Unplaced names ........................................................................................................................................................ 427
Incertae sedis............................................................................................................................................................. 428
Inquirendae................................................................................................................................................................ 431
Names wrongly treated as Siluriformes .................................................................................................................... 432
Appendix 1. Siluriform genera and species named from otoliths .................................................................................... 434
Appendix 2. Museum codes ............................................................................................................................................. 436
Appendix 3. Taxonomic publications on Siluriformes issued in 2006 ............................................................................ 440
References ........................................................................................................................................................................ 445
Index................................................................................................................................................................................. 549

Abstract
A checklist of Recent and fossil catfishes (Order Siluriformes) is presented, summarizing taxonomic literature publishedthrough 2005. From 4624 nominal species group names and 810 genus group names, 3093 species are recognized asvalid, and are distributed among 478 genera and 36 families. Distributional summaries are provided for each species, andnomenclatural synonymies, including relevant information on all name-bearing types, are included for all taxa. One newname is proposed herein: Clariallabes teugelsi, as a replacement for Clarias (Allabenchelys) dumerili longibarbis David& Poll, 1937, which is preoccupied by Clarias longibarbis Worthington, 1933, but has been treated as a valid species ofClariallabes by Teugels. Acrochordonichthys melanogaster Bleeker, 1854, is designated as type species of Acrochordonichthys Bleeker, 1857, inasmuch as no earlier valid designation has been found. A new genus Pseudobagarius, is proposed for the “pseudobagarius group” of species formerly placed in Akysis. The status of 228 species group names
remains unresolved and 31 names based on otoliths ascribed to catfishes are listed but not placed into the checklist. The
current emphasis given to catfish taxonomy at present is likely to result in a dramatic increase in the total number of valid
taxa as well as major changes in the membership of some of the higher level taxa recognized here.
Key words: Nematognathi, taxonomy, nomenclature, biodiversity, freshwater fishes

Introduction
Osteichthyian fishes of the Order Siluriformes, known by the English common name of catfishes, form a welldiagnosed natural group of primarily freshwater fishes. Most catfishes are apparently tolerant of only fresh
water but the salinity tolerance of several species extends to, or is even limited to, estuarine or even oceaniclevels of salinity. Thus, while most catfishes are recorded from inland waters, the distribution of the order also
includes coastal regions of continents and nearby islands. Catfishes often have large, heavy bones that lendthemselves to fossilization and, comparatively large otoliths. As such, a large number of species of catfishes
have been named from complete or partial skeletal fossils or even from only otoliths.
A checklist of named Recent and fossil catfishes is presented herein. The checklist is thought to be complete through the end of 2005 for available names proposed for catfishes and literature that treats taxonomic
issues involving catfish taxa. From 4620 nominal species group names and 805 genus group names, 3088 species are recognized as valid, and are distributed among 477 genera and 36 families (Table 1). In addition, 228
species group names are of uncertain validity and are in need of further study. Of the valid taxa, 19 genera and72 species are based exclusively on fossil remains. Additionally, 2 genera and 31 species names that are based
exclusively on otoliths are listed herein but are not included in the above mentioned totals.

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FERRARISTABLE 1. Numbers of valid genera and species of catfishes (Order Siluriformes) and unplaced species-group names, arranged byfamily. Fossil taxa included in generic and species totals and also listed separately. Otolith based names are not included in any of thesummary statistics.

GeneraSpeciesSpecies inquirendae

Fossil genera

Fossil species

Akysidae

5

47

0

0

0

Amblycipitidae

3

27

3

0

0

Amphiliidae

12

66

0

0

0

Anchariidae

2

5

0

0

0

Andinichthyidae

3

3

0

3

3

Ariidae

26

133

56

0

13

Aspredinidae

12

36

0

0

0

Astroblepidae

1

54

0

0

0

Auchenipteridae

20

94

12

0

0

Austroglanididae

1

3

0

0

0

Bagridae

23

189

33

4

11

Callichthyidae

8

194

0

0

1

Cetopsidae

5

41

0

0

0

Chacidae

1

3

0

0

0

Clariidae

17

113

12

0

2

Claroteidae

17

83

6

1

3

Cranoglanididae

1

3

0

0

0

Diplomystidae

2

6

0

0

0

Doradidae

30

77

4

0

0

Heptapteridae

24

189

3

0

0

Hypsidoridae

1

2

0

1

2

Ictaluridae

7

64

5

1

14

Lacantuniidae

1

1

0

0

0

Loricariidae

96

716

3

0

0

Malapteruridae

2

19

0

0

0

Mochokidae

10

188

3

0

0

Nematogenyiidae

1

2

0

0

1

Pangasiidae

5

30

2

1

2

Pimelodidae

29

93

16

0

4

Plotosidae

10

35

2

0

0

Pseudopimelodidae

5

29

1

0

0

Schilbidae

14

62

1

0

0

Scoloplacidae

1

4

0

0

0

Siluridae

12

94

28

0

3

Sisoridae

22

168

8

0

1

Trichomycteridae

41

207

1

0

0

Incertae sedis

8

13

29

7

10

Total valid taxa

478

3093

18

70

2

31

Total species inquirendae
Otolith-based names

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The primary objective of compiling this checklist is to provide catfish taxonomists with a framework for
future studies by summarizing published information on the taxonomy and nomenclature to date, while at thesame time highlighting problematic names and taxa that deserve special attention. Valid species that are notplaced in genera or, in some cases, families are indicative of broader problems with current taxonomy of thegroup into which theses taxa belong. Species that have been variously placed in one of two or three genera
suggests that one or more of the genera are likely not natural and, therefore, a situation that is in clear need offurther study. Catfish taxonomy has long suffered from having many monogeneric, or even monotypic, families. These taxa may indicate highly distinctive taxa but say nothing about the relationship of these “families”
within catfishes. Although there is no real distinction between recognizing monotypic families and treating
the included species as Incertae sedis in the Siluriformes, named families that have been treated in recent
times continue to be listed as such here. Names that have not been treated in recent years as either valid taxa orsynonyms, and therefore placed as herein species inquirendae, have in some cases simply been overlooked
and likely to be easily associated with a species after study. Others suffer from poor published descriptions,with or without an unambiguous type specimen, and may not be readily linked with any species. Such names
may need to be eliminated as available names in the future, but it is too early to do so at this time. In mostcases, such names appear to be part of particularly problematic families that need more attention, which may
shed light on the appropriate disposition of at least some of those names.
By their very nature, checklists can only represent the state of the taxonomy of the included group for abrief period of time and are out of date soon thereafter. As such, checklists represent a snapshot of a continually evolving attempt to document the biological diversity of a group of organisms. The ephemeral nature ofchecklists is especially true for this work because of the relatively large amount of taxonomic research that iscurrently being conducted on catfishes. This is due, at least in part, to the ongoing All Catfish Species Inventory (ASCI), which has focused attention on catfishes and encouraged the publication of reports on the discovery of previously unnamed species as well as revisionary studies. As an example of this emphasis on catfish
taxonomy, for 2005 alone 86 publications are listed herein, which include the descriptions of more than 100
newly-named species of catfishes and eight revisionary studies. It is likely that comparable numbers of papers
and newly discovered catfish taxa will be published over the next several years, which translates to an approximate increase of up to10% in number of recognized species during the next few years. The impact of ASCIon the rate of publication of catfish taxonomic studies is likely to extend years into the future, which suggeststhat a future catfish checklist will be larger and, quite possibly, substantially different in arrangement than the
one presented here. Nonetheless, publication of this checklist at this time is considered appropriate to bring toattention taxa in special need of study, and all of the names that need to be treated in these studies, so that
future checklists will have fewer such unresolved problems.

Methods and conventions
Valid taxa. Taxa are hierarchically arranged in alphabetical order at the level of families, genera and species,
with valid names highlighted in bold font. Intermediate levels (i. e., subfamilies, tribes, subgenera) and subspecies are not used. Taxa generally accepted in the literature as valid subspecies are listed as valid species
here. Names treated variably as subspecies and as synonyms of a species are placed as synonyms here.
After each valid name is the author and year of publication of the information necessary to make the nameavailable. Additional information that facilitates linking the name to the publication in which is was madeavailable will be found in the appropriate entry in the synonymy that follows each valid name.
Species that are treated in recent literature as valid, but of uncertain generic assignment are listed at the
beginning of the Family to which they belong, under the heading Incertae sedis, with the originally proposedscientific name. These species are considered valid, but not in the genera in which they were described and,

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possibly, not in any of the listed genera. Valid catfish genera of uncertain family placement, and any species
associated with that generic name, are listed in chronological order after the last family account under theheading of Siluriformes Incertae sedis.
A number of available species names and generic names could not be placed either as a valid name or as asynonym of another name. Often, these names were overlooked in previous taxonomic studies, or proposedwith descriptions that were not sufficiently precise to allow the assignment the name to a particular species
and the type or types are either missing or otherwise unavailable for study. Generally, such names have notbeen treated in recent literature or treated as a name of uncertain status. Herein, these names are included at
the end of the taxon to which their assignment could be confidently made, under the heading Species
inquirenda (Species inquirendae, pl.) for species names and Genus inquirendum (Genera inquirendae, pl.) for
generic names. Names of uncertain species status that also cannot be assigned to any genus are listed at theend of the family account, and names not assignable to any family are listed at the end of the checklist as Siluriformes Inquirendae immediately after Siluriformes Incertae sedis.
Valid names are corrected, as necessary, to conform to the spelling conventions specified in The International Code of Zoological Nomenclature (ICZN, 1999). That is, species-group names that were originally proposed with an initial capital letter and any name that included a diacritic mark have been corrected to a lowercase initial letter or appropriately modified spelling, respectively. Names based on fossils, both valid and junior synonyms, are preceded by a dagger (†).
For each species, a statement of the geographical distribution is provided. Generally, the statement refers
to one or more river basins, but lakes, or land masses are sometimes listed as the best summary of the distribution. Distribution statements are generally based on recently-published accounts of the most comprehensive
recorded distribution but in some cases reports from two or more sources were cited. For some species, thedistribution is based on personal observation or just the locality of the type(s). Geographical names are in
English and the spellings are the preferred spelling in Webster’s Geographical Dictionary.
Family names treated here as valid are those in current use by practicing catfish taxonomists. In somecases, those names appear not to correspond with the oldest available name for the family, as shown by thesynonymies presented under each valid family name. It was decided not to strictly adopt the senior synonyms
for families at this time, inasmuch as the names listed here (derived from Ferraris & de Pinna, 1999) might not
represent the earliest usage. Family-group names have in some cases been casually proposed in early literatureand subsequent usage usually did not include an indication of the source of the names. Thus, is it still possiblethat names adopted here as valid are indeed older than some that appear in the list of synonyms to have been
proposed earlier.
Synonymies. In the synonymies, data are presented in the following sequence. For species group names: Species name; Author(s) and date of publication of the name; Citation to source publication of the name, if different from that of the name’s author; Page(s) in which the name is made available; Figure(s) of the name’stype(s) in the publication; Type locality; Information on the identity of the primary type(s), including, whenappropriate, citation to the restriction of the primary type to a single specimen; Expanded description of thespecies by the name’s author; and Published illustration(s) of the primary type specimen (or a syntype) in subsequent literature.
For generic names: Genus or subgenus name; Author(s) and date of publication of the name; Citation tosource publication of the name, if different from that of the name’s author; Page(s) in which the name is madeavailable; Type species; Source for type species’ designation; Gender of name; and, for names originally proposed as a subgenus, the generic name within which the subgenus was originally proposed.
In synonymies, the spelling of a name is reported as it appears in the original description, including diacritic marks and upper case initial letters in species group names. If more than one spelling is found in theoriginal description, the variant spellings are noted and the valid spelling is listed at the head of the synonymy.

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A citation is provided for the first reviser action that provided the basis for the decision on which spelling istreated as valid.
The date of publication of names listed herein represents the best available information of the year that the
publication containing the new name, or that portion of the publication in which elements necessary to makethe new name available, was actually issued. When necessary, the cited date is followed by the range of dates
within which the publication was issued, to facilitate bibliographic access to the publication in question.
The type locality is that which was reported by the author of the name, either in the account of the species
or elsewhere in the publication. In some species accounts, the type locality is erroneously reported, place
names are misspelled or the locality information is brief or ambiguous. Sometimes, additional informationthat clarifies a vaguely-expressed type locality is added in square brackets. All stated localities of a syntype
series are listed and, when appropriate, the locality of the selected lectotype is noted. When a neotype wasselected to replace a missing primary type, the stated type locality is that of the neotype and not the originaltype(s). If the originally-cited type locality was found to be incorrect, the corrected type locality is reported inbrackets with supporting documentation.
Page numbers provided for each proposed new name represent the page of the publication in which thename is first proposed as new and for which information is provided that makes the name available. In a few
cases, two page numbers are listed, because the information needed to make the name available is divided
between those two pages. In some 19th and early 20th Century journals, offprint of articles were issued withpagination that differs from that of the article in the journal. In general, the page number cited herein is thatfound in the journal and not that of the offprint. However, if an offprint was issued in advance of the journaland does not include the pagination of the journal, the listed page number is that of the offprint. In a few cases,
cited publications were issued without pagination. New names described in such publications are listed with apage number enclosed in square brackets, with the assumption that the article containing the new name began
on page 1. Cuvier & Valenciennes' (1828–1850) Histoire naturelle des Poissons was issued in two, apparently
simultaneous, editions. One edition, called the Regular Edition, consists of 22 volumes in octavo format. The
other, called the Strasbourg Deluxe Edition, consists of the same number of volumes, but in a slightly larger,
royal octavo, format. Although the contents of each volume the two editions are apparently identical, the pagination of the volumes differs, with fewer pages in the Strasbourg deluxe edition. Although the existence ofthe two editions is well known (e. g., Dean, 1916: 287), most authors have access to only one of the editions(usually the Regular Edition, which is more widely available), which has resulted in a certain amount of confusion about the correct page of a new name. Herein, the page number of the Regular Edition is listed, with thepagination of the Deluxe edition in square brackets.
Museum codes that are listed for primary type specimens used here are primarily those found in Eschmeyer (1998). A few additional codes were added, based on information provided by the author of a publicationin which a new code was used. A list of the codes used herein, with the name and city and country of the institution, is provided in Appendix 2, which precedes the list of References. Museum names for which no code
was created, or for which a published code conflicts with one used in Eschmeyer (1998), are written out infull.
Only primary type specimens are reported here, including literature citations to specimens that must betreated as part of a syntype series. When reported in the original description, the number of syntypes that comprise the type series is listed in after the word ‘Syntypes’ and before the colon (:). If the number of syntypes
included within a single catalog number is known, that number if listed in parentheses immediately following
the catalog number. In some cases, only a single syntype has been identified, and the actual number of syntypes is not known. That specimen is identified as a syntype (singular) in order to assure that it is not treated asa lectotype designation. For holotypes or syntypes that have not been found, or for which there is some question of the validity of the listed specimen (s), the size or range of sizes of the specimen(s) reported in the original description is listed, as stated by the author of the name, to assist in the verification process. Catalog or

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register numbers follow the numbering system used by the museum, whenever possible. The numbering system generally follows that which is listed in a museum’s on-line catalog. For example, MNHN now uses aprefix consisting of either a small letter “a” or “b”, or a four digit number that represents the year of accession(or 0000 for unknown year), followed by a unique four-digit number for each prefix, separated from the prefix
by a dash. Other published versions of the catalog numbers for this museum’s holdings variously replace thedash by a period, using capital letters rather than small letters in the prefix, and dispensing with the “0000”
prefix or any leading zeros in the four-digit number. The Stanford University collection code (SU) is sometimes replaced by CAS-SU, indicating that the material is currently at the California Academy of Sciences. In
addition, the Stanford specimen catalog numbers are sometimes replaced by an equivalent catalog number of
the CAS numbering system, which was generated by adding 100,000 to the SU catalog number, and replacing
SU with CAS. For example, SU 1040 may be seen as written CAS-SU 1040, or CAS 101040. Herein, theoriginal numbering system, with the SU museum code, is followed.
When catalog numbers of types were found to differ from those reported in recent literature, the discrepancy was noted in order to indicate that the numbers reported herein were not the result of typographicalerrors. One class of error that requires explanation relates to a portion of the Bleeker collection that was registered at the Natural History Museum in London. That collection was registered with the prefix of BMNH
1863.12.4.XXX, but the jar labels for those specimens often listed the prefix as BMNH 1863.12.11.XXX.
These incorrect registration numbers were the basis for several published reports of the types, and possible
types, of Bleeker catfish names. Herein, published reports of the incorrect number are noted in the remarks ofthe species account to prevent future use of those same numbers.
The following terms are used in cases where the type specimen(s) are not listed: No types known.– apparently types were never saved, because the descriptions may have been made from specimens that were subsequently discarded, or from illustrations for which the underlying specimens were not retained; Whereabouts
unknown.– type specimens were apparently preserved and deposited in a collection but the current location is
not known. This does not preclude the possibility that the specimen is in the stated collection (if recorded), but
it has not been recognized as a type; At [Museum code].– there is good reason to believe that the specimen
was deposited in the collections of the museum listed, but the specimen has not been recognized as a type, orthe catalog number of the specimen was not yet found.
Within the synonymies, names are arranged chronologically. Multiple synonyms proposed in a single
publication are listed alphabetically by name. If two or more junior synonyms were published in the same year
by different authors, the synonyms are ordered alphabetically by author, except when one name is the valid
name, in which case that name is listed first.
Type localities for species names are localities as written in the original descriptions, when possible. Type
localities from publications written in alphabets other than Roman were written in a transliterated form. Whenthe publication containing an original description was not examined, the type locality was taken from that
listed in the Catalog of Fishes (Eschmeyer et al., 1998).
Classification. By their nature, checklists can only provide a single classification of the included taxonomic
diversity. This becomes problematic in situations where recently published literature includes competing ideason the phylogeny and/or classification of one or more taxonomic groups. This was indeed the case for several
groups of catfishes and I was forced to make decisions on how best to summarize the diverse conclusions on
catfish taxonomy. In some cases, the classification that was grounded on a hypothesis of phylogenetic relationships was used over another that lacked such grounding. However in other cases, none of the recently proposed classifications were based on a phylogenetic study or, alternatively, more than one phylogeneticallybased study disagreed on some aspects of the resulting classification. When left with no single classification
to adopt, I attempted to integrate available phylogenetic information together with historically accepted studies to provide a classification that appears to be best supported. Studies that were used to form the final classi-

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fication are cited immediately after the synonymy of the taxon, with comments listed in a remarks section.
Undoubtedly, this classification will not please all catfish taxonomists and even those whose studies I drew
upon for my classification may feel that their results were misinterpreted. It must be recognized by anyone
who uses this checklist that the classification represents my attempt to interpret recent taxonomic literatureand may not accurately reflect the ideas of the specialists of the various catfish groups.
Relevant literature. Following the family and Generic synonymies, literature citations are listed that wereconsulted in making the taxonomic decisions used herein, and were the source of many of the listed distributional summaries. The literature citations are subdivided into categories that indicate the nature of the underlying studies, using the following categories: Taxonomic summary.— listing of valid taxa with nomenclaturalsummary, without diagnoses or characters for identification; Checklist.— listing of valid taxa without nomenclatural summary, without diagnoses or descriptions; Identification guide— diagnoses of taxa, usually species, generally with illustrations; Revision.— taxonomic summary of all included taxa, with descriptions of all
valid species; Review.— Regionally based taxonomic summary, generally with nomenclatural summary andredescriptions of included taxa; Fauna.— regionally based summary, with diagnoses, but usually not withcomplete nomenclatural summary or redescriptions; Phylogeny.— character based interpretation of phylogenetic history; and Monophyly.— characters presented that demonstrate the monophyly of the group, but interrelationships within the group were not discussed.
References. Literature citations are arranged alphabetically by author(s) and then chronologically by year.
When multiple publications were issued within a single year by a single author, or combination of authors, thepublications are arranged alphabetically by journal title and, if needed, by pagination within a journal, followed by books or chapters in books, rather than attempting to arrange them by more precise publication date.
These publications are noted by lower case letters following the year of publication in the text as well as theReferences section. In some cases, the use of this convention results in the listing of a publication annotatedwith a lower letter of the alphabet than a paper published earlier. The difficulty in determining the precise dateof publication, or even the relative dates of publication of two papers issued in the same year, made it unnecessarily complicated to attempt to arrange publications chronologically within a year. When considered relevant to the resolution of a issue of priority, more precise dates of publication are cited in brackets at the end ofcitations. More precise publication dates may be obtained in some cases by consulting the bibliography in
Eschmeyer (1998).
Otolith-based names. Names that were proposed based only on otoliths for species that were included in catfish genera, or otherwise hypothesized to be included in the Siluriformes, are listed in a separate section ofthis publication. The names are listed in chronological order, and no attempt was made to distinguish between
valid names and junior synonyms or to place the names into the classification, In most cases, names based on
otoliths have been interpreted as belonging to catfishes of the family Ariidae.

Acknowledgments
Completion of this checklist required access to an immense amount of literature. Examining this literature wasaccomplished by traveling to institutions in several continents, obtaining copies of publications from colleagues and, for more recent publications, accessing on-line sources. These various activities were accomplished with the assistance of several institutions, colleagues and friends, and without this assistance I cannot
imagine how I could have been completed this project.
The following institutional and personal libraries were consulted during research visits: The Natural His-

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tory Library, the Cullman Library, the Vertebrate Paleontology Library, the Division of Fishes Library, and thepersonal library of Dr. Richard Vari, at the National Museum of Natural History, Smithsonian Institution,
Washington; The Research Library, the Dean Memorial Library, the Vertebrate Zoology Library and the personal library of Dr. John Maisey at the American Museum of Natural History, New York; The Vertebrate
Zoology Library and the personal library of Dr. Sven Kullander, at the Swedish Museum of Natural History,
Stockholm; The main library at The Natural History Museum, London; the Maillard Library and the Ichthyology Department Library at the California Academy of Sciences, San Francisco.
Remote access to the Zoological Record and electronic access to journals was arranged by the staffs of theAmerican Museum of Natural History library and the Maillard library of the California Academy of Sciences(CAS). Interlibrary loans were arranged through the CAS library, with the generous assistance of Annie Malley, Larry Currie and, especially, Patty Shea-Diner. Richard Vari, Heok Hee Ng and John Friel located, andprovided me with, copies of publications that I was otherwise unable to find. Years ago, Dr. James Atz generously gave me much of his reprint library that dealt with catfish taxonomy and, similarly, the late Mel Rosengave me relevant portions of the library of her husband, the late Dr. Donn Rosen.
Dr. Maurice Kottelat generously shared with me information on Southeast Asian catfish nomenclature andtaxonomy that he has been compiling. Dr. Ralf Britz examined specimens and literature at the Natural History
Museum, London, that was otherwise unavailable to me. Todd Stevenson assisted in preparing the Literaturecited, with financial support from the All Catfishes Species Inventory. Portions of this checklist werereviewed by Drs. John Lundberg, John Friel, and Mark Sabaj. The assistance of all of the people named above
was greatly appreciated, inasmuch as each contributed valuable insights that improved the quality of thechecklist.
In preparing this checklist, I drew heavily on the information and interpretations of several previous publications, most notably the Catalog of Fishes (Eschmeyer, 1998), CLOFFA (Daget et al., 1986), and CLOFFSCA (Reis et al., 2003). The efforts of the authors and editors of these publications, as well as numerous
others that cannot all be listed here, to sort out the taxonomic and nomenclatural issues the relate to catfishes
is gratefully acknowledged.
Financial support for trips to the visited institutions was provided by the FishBase Project Sweden, theNeotropical Lowlands Project of the National Museum of Natural History, and the All Catfish Species Inventory. Richard & Ann Vari, Carol Prince & Jens Vindum, Diana Shih & Benoit Jadoul, and Ricardo Castro all
generously provided me with accommodations during one or more visits to institutions near their homes. Publication support for this checklist was provided by the All Catfish Species Inventory, which received a generous grant from the United States National Sciences Foundation (DEB-0315963).
And, finally, I thank my wife, Susana Ferraris, for tolerating the piles of books, papers, and computer
hardware that have been scattered across the house over the past few years, as well as for her patience at thelack of progress on the many household projects that were delayed while this checklist was being prepared.

SILURIFORMES Cuvier, 1816
Taxonomic overview: Burgess (1989), Teugels (1996), Teugels (2003a), Nelson (2006).
Fauna: Eigenmann (1920b, 1922b, 1927, western South America); Herre (1926, Philippines); Hora & Gupta (1941,
Malay peninsula); Kottelat et al., (1993, western Indonesia); Kottelat (2001a, Vietnam); Kottelat (2001b, Laos);Rainboth (1996, Cambodian Mekong); Skelton (1993, 2001, southern Africa); Nichols (1943, China); Chu et al.
(1999, China); Fowler (1951a, 1954, Brazil); Le Bail et al. (2000, French Guiana); Jayaram (1978a, 1981, 1999,
southern Asia); Tan & Ng (2000, central Sumatra), Page & Burr (1991, North America), Miller et al. (2005,
Mexico).
Review: Day (1877a, India); Misra (1976, India and South Asia); Jayaram (1977d, various families); Menon (1999,
India); Gayet & Meunier (2003, fossil taxa).
Phylogeny: Chardon (1968), Mo (1991), Arratia (1992), de Pinna (1998a), Diogo (2003a, 2004 a, b); Hardman (2005);
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de Pinna (1996, several Asian families); Howes (1983b, loricarioid families), Arratia & Huaquin (1995, loricarioid
families); Peng et al. (2005, Chinese species).
Checklist: Kottelat (1985, Cambodia); Kottelat (1989a, Indochina); Mirza (1980, Pakistan and Azad Kashmir); Gosline(1942, 1945, Neotropics); Reis et al. (2003, Neotropics).

AKYSIDAE Gill, 1861
Akyses Gill, 1861c: 52. Type genus: Akysis Bleeker, 1858.
Parakysidae Roberts, 1989a: 141. Type genus: Parakysis Herre, 1940.
Acrochordonichthyini de Pinna, 1996: 61. Type genus: Acrochordonichthys Bleeker, 1857.
Monophyly and phylogenetic position: de Pinna (1996).
Identification guide: Jayaram (1977d), South Asia.
5 genera, 47 species; no named fossil taxa.
ACROCHORDONICHTHYS Bleeker, 1857
Acrochordonichthys Bleeker, 1857b: 473. Type species: Pimelodus melanogaster Bleeker, 1854. Type by present
designation. Gender: Masculine.
Sosia Vaillant, 1902: 81. Type species: Sosia chamaeleon Vaillant, 1902. Type by monotypy. Gender: Feminine.
Revision: Ng & Ng (2001).
Remarks: Earlier statements (e. g., Eschmeyer, 1990, 1998; Ng & Ng, 2001) indicating that the type of Acrochordonichthys as A. platycephalus Bleeker, 1858, are in error. They were based on a type designation in Bleeker(1862-63) which was of a species that was not among the available named species when Acrochordonichthys
was proposed. Apparently, no valid designation of a type species have been made before now.
Acrochordonichthys chamaeleon (Vaillant, 1902)
Sosia chamaeleon Vaillant, 1902: 82, figs. 19–21. Type locality: Borneo: Raoen/Mandai. Lectotype: RMNH 7849,
designated by Ng & Ng (2001: 406).
Distribution: Kapuas River basin, western Borneo (Ng & Ng, 2001).
Acrochordonichthys falcifer Ng & Ng, 2001
Acrochordonichthys falcifer Ng & Ng, 2001: 408, figs. 5–6. Type locality: Borneo: Sabah, Sungai Segama. Holotype: ZRC 43321.
Distribution: Kinabatangan and Segama River basins, northeastern Borneo (Ng & Ng, 2001).
Acrochordonichthys guttatus Ng & Ng, 2001
Acrochordonichthys guttatus Ng & Ng, 2001: 394, figs. 6–8, 10. Type locality: Borneo: Kalimantan Tengah, Barito
River drainage, Sungai Barito at Muara Laung. Holotype: MZB 9301.
Distribution: Barito River basin, southern Borneo (Ng & Ng, 2001).
Acrochordonichthys gyrinus Vidthayanon & Ng, 2003
Acrochordonichthys gyrinus Vidthayanon & Ng, 2003: 2, fig. 1. Type locality: Thailand: Phitsanulok Province, Yom
River at Primpiram; the Inland fishery Station of Phitsanulok Province. Holotype: NIFI 2645.
Distribution: Yom River, Chao Phraya basin, Thailand (Vidthayanon & Ng, 2003).
Acrochordonichthys ischnosoma Bleeker, 1858
Acrochordonichthys ischnosoma Bleeker, 1858b: 232. Type locality: Java (Parongkalong provinciae Preanger), influmine Tjitarum. Holotype (115 mm TL): BMNH 1863.12.4.151, illustrated in Bleeker (1862–63: pl. 84[=Silur. pl. 36], fig. 4), and Ng & Ng (2001: fig. 4).
Distribution: Sumatra and Java (Ng & Tan, 2002).
Remarks: Redescribed in Ng & Tan (2002). Catalog number of holotype of Acrochordonichthys ischnosoma incorrectly reported as BMNH 1863.12.11.151 in Eschmeyer et al. (1998) and Ng & Ng (2001).
Acrochordonichthys mahakamensis Ng & Ng, 2001
Acrochordonichthys mahakamensis Ng & Ng, 2001: 400, figs. 6–7, 11. Type locality: Borneo: Kalimantan Timur,
Mahakam River, boulders upstream of Melak (0º12'S, 115º47'E). Holotype: MZB 5952.
Distribution: Mahakam River, eastern Borneo (Ng & Ng, 2001).

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FERRARISAcrochordonichthys pachyderma Vaillant, 1902
Acrochordonichthys pachyderma Vaillant, 1902: 66, figs. 11–13. Type locality: Bloeoe, Bornéo central. Holotype:
RMNH 7843 (1).
Distribution: Kapuas, Mahakam and Kinabatangan River basins, Borneo (Ng & Ng, 2001).
Remarks: RMNH 7560, from the Bo River, is incorrectly listed as a syntype in Ng & Ng (2001) and Eschmeyer etal. (1998).
Acrochordonichthys rugosus (Bleeker, 1847)
Pimelodus rugosus Bleeker, 1847b: 11 (and 1847a: 168). Type locality: Java. Possible syntypes: BMNH
1863.12.4.150 (1), BMNH 1863.12.4.161 (1), RMNH 6883 (2).
Pimelodus melanogaster Bleeker, 1854a: 89. Type locality: Sumatra orientalis, provinc. Palembang, ubi confluunt
flumina Lamatang et Enim. Holotype (110 mm TL): possibly BMNH 1863.12.4.160 (see Ng & Ng: 2001: 413);
holotype illustrated in Bleeker (1862–63: 67, pl. 84 [= Silur. 36], fig. 3) as Acrochordonichthys melanogaster.
Pimelodus pleurostigma Bleeker, 1854b: 442. Type locality: Batavia, in fluviis. Syntypes (8, 48–76 mm TL):
BMNH 1863.12.4.161 (1), NMV 45993 (1), RMNH 4188 (1), RMNH 6879 (4).
Pimelodus zonatus Bleeker, 1854b: 444. Type locality: Batavia, in fluviis. Syntypes (9, 54–80 mm TL): BMNH
1863.12.4.175 (1), NMV 45992 (1), RMNH 6880 (5). One syntype illustrated in Bleeker (1862–63: 69, pl. 84[=Silur. 36], fig. 5), as Acrochordonichthys zonatus.
Acrochordonichthys platycephalus Bleeker, 1858b: 224. Type locality: Sumatra (Palembang) in fluviis. Holotype(97 mm TL): BMNH 1863.12.4.152; illustrated in Bleeker (1862–63: pl. 84 [= Silur. 36], fig. 2).
Sosia chamaeleon pallida Vaillant, 1902: 87, figs. 22–24. Type locality: Kapoeas, la plupart du haut du Fleuve(Sous-bassins du Mandaiet du Sibau), Bornéo central. Lectotype: RMNH 7848, designated by Ng & Ng (2001:
406). Originally Sosia chamaeleon var. pallida.
Acrochordonichthys Büttikoferi Popta, 1904: 188. Type locality: le Bongan et le Bo, Bornéo central. Syntypes:
RMNH 7557 (1), RMNH 7558 (2); described in more detail, with an illustration of one of the syntypes, in Popta(1906: pl. 3, figs. 12a, 12b).
Acrochordonichthys obscurus Popta, 1904: 187. Type locality: le Bongan, Bornéo central. Holotype: RMNH 7556;
described in more detail, with an illustration of the holotype, in Popta (1906: 58, pl. 3, figs. 11a, 11b).
Acrochordonichthys varius Popta, 1904: 189. Type locality: le Bo, Bornéo central. Syntypes: RMNH 7559 (2);described in more detail, with an illustration of one of the syntypes, in Popta (1906: 63, pl. 4, figs. 13a, 13b).
Distribution: Solo, Ciliwung, Citarum River basins, Java; Barito, Kapuas, Mahakam and Rajang River basins,
Borneo; Bernam, Terengganu, Mae Nam Sungai Kolok and Pattani River basins, peninsular Malaysia and Thailand; Musi and Tulangbawan River basins, Sumatra (Ng & Ng, 2001).
Remarks: See comments in Ng & Ng (2001: 413) regarding the difficulty in identifying types of species named by
Bleeker. Register number of BMNH types of Pimelodus rugosus, P. pleurostigma, P. melanogaster, P. zonatus,
and A. platycephalus incorrectly reported in Eschmeyer et al. (1998) and Ng & Ng (2001).
Acrochordonichthys septentrionalis Ng & Ng, 2001
Acrochordonichthys septentrionalis Ng & Ng, 2001: 401, figs. 6, 12. Type locality: Thailand: Kanchanaburi Province, Kwae Noi River. Holotype: NIFI 3128.
Distribution: Mae Khlong River basin, Thailand, and Pahang River basin, Malaysia (Ng & Ng, 2001).
Acrochordonichthys strigosus Ng & Ng, 2001
Acrochordonichthys strigosus Ng & Ng, 2001: 403, figs. 5–6, 8, 13. Type locality: Borneo: Kalimantan Barat, Sungai Sibau, Mouth of Sungai Putan to 3 km downstream (1º02'28.0"N, 112º59'58.6"E). Holotype: MZB 9333.
Distribution: Kapuas River basin, western Borneo (Ng & Ng, 2001).
AKYSIS Bleeker, 1858
Akysis Bleeker, 1858b: 204, 234 (and 1858d: 419). Type species: Pimelodus variegatus Bleeker, 1846. Type bymonotypy. Gender: Masculine.
Reviews: Ng & Kottelat (1998c, Indochina, with key to species); Ng (1999e, Myanmar).
Akysis brachybarbatus Chen, 1981
Akysis brachybarbatus Chen, in He & Chen, 1981: 210, fig. 2. Type locality: Nanrui River, Mekong River system,CHECKLIST OF CATFISHES, RECENT AND FOSSIL

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Manglian Co., Yunnan, China. Holotype: KIZ 787162.
Distribution: Lancangjiang [= Mekong River] tributaries, China (Chu et al., 1999).
Akysis clavulus Ng & Freyhof, 2003
Akysis clavulus Ng & Freyhof, 2003: 312, figs. 1–2. Type locality: Vietnam: Khanh Hoa province: Song Duc My
about 10 km west of Ninh Hoa; 12º32.22'N, 109º01.38'E. Holotype: ZFMK 27159.
Distribution: Cai and Duc My rivers, central Vietnam (Ng & Freyhof, 2003).
Akysis clinatus Ng & Rainboth, 2005
Akysis clinatus Ng & Rainboth, 2005: 34, fig. 1. Type locality: Cambodia: Stung Chhay River, Tok Kong village onroad from Phnom Penh to Skhanoukville, 10º56'37.8"N 113º47'47.4"E. Holotype: ZRC 47288.
Distribution: Stung Chhay River, a Gulf of Thailand draining basin, Cambodia (Ng & Rainboth, 2005: 35).
Akysis ephippifer Ng & Kottelat, 1998
Akysis ephippifer Ng & Kottelat, 1998c: 1078, fig. 11. Type locality: Cambodia: Stung Santréa, 40 km from Pomnac. Holotype: MNHN 1977-0034.
Distribution: Xe Kong basin, Lower Mekong drainage, Laos and Cambodia (Ng & Kottelat, 1998c).
Akysis fuliginatus Ng & Rainboth, 2005
Akysis fuliginatus Ng & Rainboth, 2005: 35, fig. 4. Type locality: Cambodia: Stung Treng province, Mekong Riveron W edge of Kaoh Han, 16 km NE of Stung Treng, 13º38'N 106º3'E. Holotype: UMMZ 241338.
Distribution: Lower Mekong River basin, Cambodia (Ng & Rainboth, 2005: 37).
Akysis hendricksoni Alfred, 1966
Akysis hendricksoni Alfred, 1966: 467, fig. 1. Type locality: Terengganu River at Kuala Brang, Terengganu, Malaya.
Holotype: ZRC 1414.
Distribution: Sungai Golok, Sungai, Terengganu, Sabak Bernam, and Endau basins, peninsular Malaysia and Thailand (Ng & Tan, 1999).
Akysis heterurus Ng, 1996
Akysis heterurus Ng, 1996: 4, figs. 1–2. Type locality: Sumatra: Jambi, Sg. Alai at 19.5 km Muara Bungo-Muara
Tebo road. Holotype: ZRC 34194.
Distribution: Batang Hari River basin, eastern Sumatra (Ng, 1996).
Akysis maculipinnis Fowler, 1934
Akysis maculipinnis Fowler, 1934a: 97, fig. 46. Type locality: Chantaboon, Southeast Siam. Holotype: ANSP 59366.
Distribution: Rivers of southeastern Thailand (Ng & Kottelat, 1998c).
Akysis microps Ng & Tan, 1999
Akysis microps Ng & Tan, 1999: 359, fig. 5. Type locality: Malaysia: Johor, Sungai Kahang and tributary ca. 44.4
km from Mersing turnoff to Kluang just before side road to Endau Rompin Taman Negara, km 96 from Mersing
to Batu Pahat (2º3.93'N, 103º31.58'E). Holotype: ZRC 42596.
Distribution: Endau River basin, southeastern Malay Peninsula (Ng & Tan, 1999).
Akysis pictus Günther, 1883
Akysis pictus Günther, 1883: 138. Type locality: Tenasserim [Burma]. Syntypes: BMNH 1880.12.1.25–26 (2); illustrated in Ng (1999e: 542, fig. 1).
Distribution: Southeastern Myanmar, in former Tenasserim district; known only from types (Ng, 1999e).
Remarks: Redescribed in Ng (1999e).
Akysis prashadi Hora, 1936
Akysis variegatus variegatus Prashad & Mukerji, 1929: 180, pl. 8 (figs. 1–2). Type locality: South end of Indawgyi
Lake, along sw. shore, Myitkyina Dist., Upper Burma. Holotype: ZSI F10873/1. Preoccupied in Akysis byPimelodus variegatus Bleeker, 1846; replaced by Akysis prashadi Hora, 1936.
Akysis prashadi Hora, 1936a: 200, figs. 1, 2 (b–c). Type locality: South end of Indawgyi Lake, along sw. shore,
Myitkyina Dist., Upper Burma. Holotype: ZSI F10873/1.
Distribution: Indawgyi Lake and northern Irrawaddy River basin, Myanmar (Ng, 1999e).
Remarks: Redescribed in Ng (1999e).
Akysis recavus Ng & Kottelat, 1998
Akysis recavus Ng & Kottelat, 1998c: 1090, fig. 16. Type locality: Thailand: Western Chao Phraya basin. Holotype:

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FERRARISZRC 40716.
Distribution: Western Chao Phraya basin, Thailand (Ng & Kottelat, 1998c).
Akysis variegatus (Beeker, 1846)
Pimelodus variegatus Bleeker, 1846: 177. Type locality: Batavia. Syntypes: BMNH 1863.12.4.187 (1), possibly
NMV 46004 (1), RMNH 6881 (16).
Distribution: Java.
Remarks: Register number of syntype of Akysis variegatus incorrectly reported as BMNH 1863.12.11.187 in
Eschmeyer et al. (1998).
Akysis varius Ng & Kottelat, 1998
Akysis varius Ng & Kottelat, 1998c: 1093, fig. 18. Type locality: Xe Bangfai, ca 3 km upriver of Pan Pakphanang,
Khammouan Prov., Laos. Holotype: ZRC 41015.
Distribution: Mekong River basin, Laos, Cambodia and Thailand (Kottelat, 2001).
Akysis vespa Ng & Kottelat, 2004
Akysis vespa Ng & Kottelat, 2004: 194, fig. 1. Type locality: Myanmar: Kayin State, stream “Chon Son” between
Kyondaw and Phadaw, about 20 km NW of Payathouzu (at border with Thailand), 15º25'N, 98º15'E. Holotype:ZRC 46423.
Distribution: Ataran River basin, Myanmar (Ng & Kottelat, 2004).
BREITENSTEINIA Steindachner, 1881
Breitensteinia Steindachner, 1881a: 46. Type species: Breitensteinia insignis Steindachner, 1881. Type by monotypy.
Gender: Feminine. Also reported as new in Steindachner (1881e: 213).
Revision: Ng & Siebert (1998).
Breitensteinia cessator Ng & Siebert, 1998
Breitensteinia cessator Ng & Siebert, 1998: 647, fig. 1. Type locality: Borneo: Kapuas mainstream 58 km NE of
Sintang and 1 km downstream from Sebruang, (0º25.5'N, 111º52.5'E). Holotype: MZB 3693.
Distribution: Batang Hari and Tulangbawan basins, Sumatra, and Kapuas River basin, western Borneo (Ng & Siebert, 1998).
Breitensteinia hypselurus Ng & Seibert, 1998
Breitensteinia hypselurus Ng & Seibert, 1998: 650, figs. 3, 4a. Type locality: Borneo: Sanggau. Holotype: RMNH
16048.
Distribution: Kapuas River basin, western Borneo (Ng & Siebert, 1998).
Breitensteinia insignis Steindachner, 1881
Breitensteinia insignis Steindachner, 1881a: 46. Type locality: Teweh [= Moara Teweh, Barito River, Borneo]. Holotype: NMW 55042, illustrated in Steindachner (1881f: pl. 1, fig. 2) and described in more detail in Steindachner(1881e: 213).
Distribution: Barito River basin, southern Borneo (Ng & Siebert, 1998).
PARAKYSIS Herre, 1940
Parakysis Herre, 1940: 11. Type species: Parakysis verrucosa Herre, 1940. Type by original designation. Gender:
Masculine. Name treated by Herre as feminine, but subsequently treated as masculine (Ng & Lim, 1995), presumably following the gender of Akysis.
Revision: Ng & Lim (1995).
Parakysis anomalopteryx Roberts, 1989
Parakysis anomalopteryx Roberts, 1989a: 142. Type locality: Kapuas basin, Sungai Seriang, a forested tributary of
the Sungai Palin, 37 km W of Putussibau, Western Borneo (Kalimantan Barat, Indonesia). Holotype: MZB
3702.
Distribution: Kapuas River basin, western Borneo (Ng & Lim, 1995).
Parakysis grandis Ng & Lim, 1995
Parakysis grandis Ng & Lim, 1995: 260, fig. 7. Type locality: Sumatra: Riau Province, small stream with grassy

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banks near Pangkalankasai, 43 km. south of Rengat. Holotype: ZRC 39111.
Distribution: Kapuas and Kuching River basins, Borneo; Deli, Indragiri, and Batang Hari basins, Sumatra (Ng &Lim, 1995).
Parakysis longirostris Ng & Lim, 1995
Parakysis longirostris Ng & Lim, 1995: 262, fig. 8. Type locality: Singapore: Nee Soon swamp forest. Holotype:
ZRC 34491.
Distribution: Singapore, peninsular Malaysia and Riau Archipelago (Ng & Lim, 1995; Ng & Tan, 1999).
Parakysis notialis Ng & Kottelat, 2003
Parakysis notialis Ng & Kottelat, 2003: 48, figs. 1, 2a, 3a. Type locality: Borneo: Kalimantan Selatan, Barito Riverdrainage, area of Tamiyang Layang, 2º01'S, 115º07'E. Holotype: MZB 5994.
Distribution: Barito River basin, southern Borneo (Ng & Kottelat, 2003).
Parakysis verrucosus Herre, 1940
Parakysis verrucosa Herre, 1940: 12, pl. 6. Type locality: Mawai District, Johore, Malaysia. Holotype: SU 33009.
Distribution: Peninsular Malaysia and Riau Archipelago (Ng & Lim, 1995).
PSEUDOBAGARIUS new genus
Type species: Akysis pseudobagarius Roberts, 1989. Gender: Masculine.
Diagnosis: Akysidae with the following combination of characters: snout extending well anterior of margin of lower
jaw, rendering mouth subterminal; narial openings relatively large and closely set; and caudal fin deeply forked.
Remarks: This genus corresponds to the “pseudobagarius group” of species of the genus Akysis that was first recognized by Ng & Kottelat (1998).
Pseudobagarius alfredi (Ng & Kottelat, 1998)
Akysis alfredi Ng & Kottelat, 1998c: 1063, fig. 5. Type locality: Malaysia: Pahang, Tasek Bera, approx. 300 m E ofFort Iskandar. Holotype: ZRC 40714.
Distribution: Tasek Bera, southern Malay Peninsula (Ng & Kottelat, 1998c).
Pseudobagarius baramensis (Fowler, 1905)
Akysis baramensis Fowler, 1905: 472, fig. 4. Type locality: Baram river, Borneo. Holotype: ANSP 114887.
Distribution: Baram River basin, Borneo (Fowler, 1905).
Pseudobagarius filifer (Ng & Rainboth, 2005)
Akysis filifer Ng & Rainboth, 2005: 37, fig. 5. Type locality: Cambodia: Kandal, Tonle Sap, fishing lot 8, 22 miles
upstream from Phnom Penh, 11º44'N, 104º50'E. Holotype: UMMZ 235728.
Distribution: Tonlé Sap drainage, Mekong River basin, Cambodia (Ng & Rainboth, 2005: 39).
Pseudobagarius fuscus (Ng & Kottelat, 1996)
Akysis fuscus Ng & Kottelat, 1996: 20, fig. 1. Type locality: Borneo: Kalimantan Barat, Danau Sentarum area, Sungai Hulu Leboyan at Keluwin. Holotype: MZB 5934.
Distribution: Kapuas River basin, western Borneo (Ng & Kottelat, 1996).
Pseudobagarius hardmani (Ng & Sabaj, 2005)
Akysis hardmani Ng & Sabaj, 2005: 218, fig. 3. Type locality: Thailand: Nakhon Sawan, Mae Nam Chao Phraya andtributaries in the vicinity of Phayuka Khiri. Holotype: ANSP 178858.
Distribution: Chao Phraya River basin, Thailand (Ng & Sabaj, 2005).
Pseudobagarius inermis (Ng & Kottelat, 2000)
Akysis inermis Ng & Kottelat, 2000b: 9, figs. 1, 2a. Type locality: Laos: Attapeu province: unnamed creek entering
Xe Kaman from the north at proposed Xe Kaman dam site, 14º57'40"N, 107º09'16"E. Holotype: ZRC 45401.
Distribution: Xe Banghiang and Xe Kong basins, Mekong River drainage, southern Laos (Ng & Kottelat, 2000b).
Pseudobagarius leucorhynchus (Fowler, 1934)
Akysis leucorhynchus Fowler, 1934a: 97, figs. 44–45. Type locality: Chieng Mai, North Siam. Holotype: ANSP
59346.
Distribution: Chao Phraya basin, and rivers of southern peninsular Thailand (Ng & Kottelat, 1998c).
Pseudobagarius macronemus (Bleeker, 1860)
Akysis macronema Bleeker, 1860: 11. Type locality: Lahat [Sumatra]. Syntypes (4: 36–44 mm TL): BMNH

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FERRARIS

1863.12.4.188 (1), RMNH 6729 (2); one syntype illustrated in Bleeker (1862–63: 65, pl. 84 [= Silur. 36], fig. 1).
Distribution: Sumatra (Tan & Ng, 2000) .
Remarks: Redescribed in Tan & Ng (2000) from specimens from the Batang Hari basin of central Sumatra. Register
number of holotype of Akysis macronema incorrectly reported as BMNH 1863.12.11.188 in Eschmeyer et al.
(1998).
Pseudobagarius meridionalis (Ng & Siebert, 2004)
Akysis meridionalis Ng & Siebert, 2004: 2, figs. 1, 2a. Type locality: Indonesia, Kalimantan Tengah (Borneo); Barito
River drainage, Sungai Laung at Dessa Maruwei (0º21.986'S, 114º44.103'E). Holotype: MZB 6102.
Distribution: Barito River basin, Borneo (Ng & Siebert, 2004).
Pseudobagarius nitidus (Ng & Rainboth, 2005)
Akysis nitidus Ng & Rainboth, 2005: 40, fig. 8. Type locality: Laos: Champasak province, Mekong River at Ban
Hang Khone, just downstream from Khone falls. Holotype: UMMZ 235400.
Distribution: Mekong River basin, Laos (Ng & Rainboth, 2005).
Pseudobagarius pseudobagarius (Roberts, 1989)
Akysis pseudobagarius Roberts, 1989a: 138, fig. 107. Type locality: Bar in mouth of Sungai Melawi at Sintang,
Western Borneo (Kalimantan Barat, Indonesia). Holotype: MZB 3691.
Distribution: Moesi and Kapuas Rivers, Western Borneo (Roberts, 1989a).
Pseudobagarius similis (Ng & Kottelat, 1998)
Akysis similis Ng & Kottelat, 1998c: 1074, figs. 4e, 9, 13c. Type locality: Vietnam: Kihn Thuy Cai canal, 10 km E ofVinh Long, at mouth of canal into Mekong. Holotype: UMMZ 214902.
Distribution: Mekong River mouth, southern Vietnam (Ng & Kottelat, 1998c).
Remarks: Coloration of adults and ontogenetic change from juvenile color pattern discussed in Ng & Rainboth(2005: 41).
Pseudobagarius sinensis (He, 1981)
Akysis sinensis He, in He & Chen, 1981: 209, fig. 1. Type locality: Lancang Jiang, Xishuangbanna Daizu Zizhizhou,
Yunnan Prov., China. Syntypes: IHASW 638237 (1), IHASW 638283 (1), KIZ 735007 (1), KIZ 735008 (1).
Also as Akysis sinesis once in English summary (p. 213); first reviser apparently Eschmeyer et al. (1998).
Distribution: Lancangjiang [= Mekong River], Yunnan, China (Chu et al., 1999).

AMBLYCIPITIDAE Day, 1873
Amblycepinae Day, 1873a: cclxviii. Type genus: Amblyceps Blyth, 1858.
Identification guide: Jayaram (1977), South Asia.
Phylogeny and key to genera: Chen & Lundberg (1995).
Review: Choo (1987), Korea.
3 genera, 27 species; no named fossil taxa.
AMBLYCEPS Blyth, 1858
Amblyceps Blyth, 1858: 281. Type species: Amblyceps caecutiens Blyth, 1858. Type by monotypy. Gender: Neuter.
Branchiosteus Gill, 1861c: 52. Olyra laticeps M’Clelland, 1842. Type by original designation. Gender: Masculine.
Review: Hora (1933) south Asia; Ng & Kottelat (2000a), Indochinese species.
Remarks: Ng & Kottelat (2000a: 346) list Amblyceps inermis as a nomen nudem published first in M’Clelland(1842). Amblyceps deyi is listed in Tamang (1993: 7), as a name only and is therefore not available.
Amblyceps apangi Nath & Dey, 1989
Amblyceps apangi Nath & Dey, 1989: 2, fig. 2. Type locality: Dikrong, Arunachal Pradesh, India. Holotype: F/GUZ
1215.
Distribution: Rivers draining Arunachal Pradesh, India (Nath & Dey, 1989).
Amblyceps arnunachalensis Nath & Dey, 1989
Amblyceps arnunachalensis Nath & Dey, 1989: 3, fig. 3. Type locality: Dikrong, Arunachal Pradesh, India. Holo-CHECKLIST OF CATFISHES, RECENT AND FOSSIL

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type: F/GUZ 1213.
Distribution: Rivers draining Arunachal Pradesh, India (Nath & Dey, 1989).
Amblyceps caecutiens Blyth, 1858
Amblyceps cæcutiens Blyth, 1858: 282. Type locality: Maulmein and Pegu [Myanmar]. Syntypes: Whereabouts
unknown. Originally: Amb. cæcutiens.
Akysis kurzii Day, 1872: 703. Type locality: Pegu Yomas, Burma. Holotype: Whereabouts unknown.
Distribution: Bago River basin, Myanmar (Ng & Kottelat, 2000a).
Amblyceps carinatum Ng, 2005
Amblyceps carinatum Ng, 2005d: 243, fig. 1. Type locality: Myanmar: Kachin, Irrawaddy drainage, hillstreams at
Tonpan village, on road from Myitkyina to Tanai. Holotype: UMMZ 245588.
Distribution: Middle Irrawaddy River basin, Myanmar (Ng, 2005d).
Amblyceps foratum Ng & Kottelat, 2000
Amblyceps foratum Ng & Kottelat, 2000a: 338, fig. 4. Type locality: Malaysia: Terengganu, Sungai Brang just outside Sekayu Waterfall Park; 4º57'51"N, 102º57'46"E. Holotype: ZRC 43718.
Distribution: Northern peninsular Malaysia, southern peninsular Thailand, southeastern Thailand, and coastal Cambodia (Ng & Kottelat, 2000a).
Amblyceps laticeps (M’Clelland, 1842)
Olyra laticeps M’Clelland, 1842: 588, pl. 21 (fig. 2). Type locality: Kasyah mountains. Holotype: apparently notpreserved.
Distribution: South Asia, but distribution not well documented.
Remarks: Inclusion in Amblyceps follows Vinciguerra (1890), Hora (1936a), and Ng & Kottelat (2000a).
Amblyceps macropterus Ng, 2001
Amblyceps macropterus Ng, 2001b: 202, fig. 1. Type locality: Pakistan: Baluchistan Province: Las Bela District:
Hab River at Hinidan. Holotype: UMMZ 187411.
Distribution: Hab River, southwestern Pakistan; known only from the holotype (Ng, 2001b).
Amblyceps mangois (Hamilton, 1822)
Pimelodus mangois Hamilton, 1822: 199, 379. Type locality: Tanks of Northern Behar [India]. No types known.
Previously unpublished Hamilton illustration of a type reproduced in Hora (1929: pl. 22, fig. 4).
Distribution: South Asia; precise distribution unknown inasmuch as literature accounts of this species may refer toalmost any of the species in this genus.
Amblyceps mucronatum Ng & Kottelat, 2000
Amblyceps mucronatum Ng & Kottelat, 2000a: 340, fig. 6. Type locality: Thailand: Loei Prov.: Mekong River basin,
Huang River tributary at 17º16'0.9"N, 101º08'29.2"E. Holotype: ZRC 41323.
Distribution: Mekong River basin, Thailand, and Salween River basin, Thailand and Myanmar (Ng & Kottelat,
2000a).
Amblyceps murraystuarti Chaudhuri, 1919
Amblyceps murray-stuarti Chaudhuri, 1919: 272, pl. 22 (figs. 1, 1a, 1b). Type locality: A mountain stream in the
Putao Plains in the northern Frontier of Upper Burma. Holotype: ZSI F9736/1.
Distribution: Upper Irrawaddy River basin, Myanmar (Ng & Kottelat, 2000a).
Amblyceps platycephalus Ng & Kottelat, 2000
Amblyceps platycephalus Ng & Kottelat, 2000a: 343, fig. 8. Type locality: Thailand: Mae Hong Son Province: Nam
Mae Pai basin, Nam Mae Sa-Nga at Ban Yang Top Sole, immediately downstream of Nam Tok Pa Sua,
19º29'0"N, 97º57'50"E. Holotype: ZRC 42745.
Distribution: Salween River, Thailand, known only from the holotype (Ng & Kottelat, 2000a).
Amblyceps serratum Ng & Kottelat, 2000
Amblyceps serratum Ng & Kottelat, 2000a: 344, fig. 9. Type locality: Cambodia: Stung Treng province: shallow
channel across S end of Kaoh Han, 14 km NE of Stung Treng (13º37'N, 106º03'E). Holotype: UMMZ 234708.
Distribution: Mekong River drainage of central and southern Laos and Cambodia (Ng & Kottelat, 2000a).
Amblyceps tenuispinis Blyth, 1860
Amblyceps tenuispinis Blyth, 1860: 153. Type locality: Gházipur [India]. Holotype: Whereabouts unknown.

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FERRARISDistribution: Gházipur, India.
Amblyceps variegatum Ng & Kottelat, 2000
Amblyceps variegatum Ng & Kottelat, 2000a: 345, fig. 10. Type locality: Thailand: Kanchanaburi province, Mae
Nam Khwae Noi basin, Nam Khung, upstream of Ban Huai Pak Khung, riffles between second ford and reservoir (14º37'20"N, 98º31'24"E). Holotype: ZRC 43719.
Distribution: Mae Khlong River basin, Thailand (Ng & Kottelat, 2000a).
Species inquirendae, Amblyceps
Pimelodus indicus M’Clelland & Griffith, in M’Clelland, 1842: 584. Type locality: Loodianah. Holotype: Whereabouts
unknown, possibly at BMNH.
LIOBAGRUS Hilgendorf, 1878
Liobagrus Hilgendorf, 1878: 155. Type species: Liobagrus reinii Hilgendorf, 1878. Type by monotypy. Gender:
Masculine.
Neobagrus Bellotti, 1892: 100. Type species: Neobagrus fuscus Bellotti, 1892. Type by monotypy. Gender: Masculine.
Remarks: Systematic position discussed in Jayaram (1965b).
Review: Choo (1987), Korea.
Key: Regan (1908g).
Liobagrus andersoni Regan, 1908
Liobagrus andersoni Regan, 1908g: 61, pl. 3 (fig. 4). Type locality: Kimhoa, 65 mi. north-east of Seoul, Corea. Syntypes: BMNH 1907.12.10.63–64 (2), BMNH 1907.12.10.65 (1, skeleton).
Distribution: Chongiu, Kwangiu, Soungchon, Kosong, and Ulchin, Korea (Mori, 1952).
Liobagrus anguillicauda Nichols, 1926
Liobagrus anguillicauda Nichols, 1926b: 1, fig. 1. Type locality: Chungan Hsien, nw. Fukien Prov., China. Holotype: AMNH 8474.
Distribution: Streams of southeast China (Chu et al., 1999).
Liobagrus formosanus Regan, 1908
Liobagrus formosanus Regan, 1908e: 360. Type locality: Lake Candidius, Formosa. Holotype: BMNH
1909.4.28.23.
Distribution: Taiwan (Chu et al., 1999).
Liobagrus kingi Tchang, 1935
Liobagrus kingi Tchang, 1935a: 95, fig. 1. Type locality: Tsin-ning, Yunnan, China. Holotype: at ASIZB [=ZMFMIB 11673].
Distribution: Upper Changjiang [= Yangtze River] (Chu et al., 1999).
Liobagrus marginatoides (Wu, 1930)
Amblyceps marginatoides Wu, 1930a: 256, fig. 2. Type locality: Sichuan, China. Syntypes (2): Whereabouts
unknown.
Distribution: Upper Changjiang [= Yangtze River] (Chu et al., 1999).
Liobagrus marginatus (Günther, 1892)
Amblyceps marginatus Günther, 1892: 245, pl. 2 (fig. a). Type locality: Mountain streams running into Min River,
Szechwan [Sichuan], China. Holotype: BMNH 1891.6.13.10.
Distribution: Upper Changjiang [= Yangtze River] (Chu et al., 1999).
Liobagrus mediadiposalis Mori, 1936
Liobagrus mediadiposalis Mori, 1936: 673, pl. 24 (fig. 3). Type locality: River Rakuto at Bun-kei, Chosen [Korea].
Holotype (130 mm TL): At Preparatory Department of Keijo Imperial University.
Distribution: Rivers of Natong, Kokson, Yongsan, Kim and Tchongchin, Korea (Mori, 1952).
Liobagrus nantoensis Oshima, 1919
Liobagrus nantoënsis Oshima, 1919: 183, pl. 48 (fig. 3). Type locality: Dainansho, Nanto, Taiwan. Lectotype:

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FMNH 59080, designated by Eschmeyer (in Eschmeyer et al., 1998: 1149).
Distribution: Taiwan (Chu et al., 1999).
Liobagrus nigricauda Regan, 1904
Liobagrus nigricauda Regan, 1904a: 193. Type locality: Sea of Tien [Tien Chih], Yunnan Fu, China, elev. 6000 ft.
Syntypes: BMNH 1904.1.26.42–43 (2).
Distribution: Changjiang [= Yangtze River] basin (Chu et al., 1999).
Liobagrus obesus Son, Kim & Choo, 1987
Liobagrus obesus Son, Kim & Choo, 1987: 22, figs. 1, 5b. Type locality: K§m-river, Cho'gang-ri, Simch'4n-my4n,
Y4ngdong, Ch'ungch'4ngbug-do province, Korea. Holotype: BCUE 2101.
Distribution: Kum River basin, South Korea (Son et al., 1987).
Liobagrus reinii Hilgendorf, 1878
Liobagrus reinii Hilgendorf, 1878: 155. Type locality: Japan. No types known.
Liobagrus sugubii Regan, 1908b: 152. Type locality: Lake Biwa, Japan. Syntypes: BMNH 1907.12.23.56–59 (4).
Distribution: Honshu, Skikoku and Kyushu Islands, Japan (Matsuura et al., 2000).
Liobagrus styani Regan, 1908
Liobagrus styani Regan, 1908b: 152. Type locality: South Hupeh, China. Syntypes (2): BMNH 1902.5.30.2–3 (2,dry), BMNH 1902.5.30.4 (1, skeleton).
Distribution: Middle and lower Changjiang [= Yangtze River] (Chu et al., 1999).

Species inquirenda, Liobagrus
Neobagrus fuscus Bellotti, 1892: 101, unnumbered figure. Type locality: Jokohama [Japan]. MSNM (destroyed: Eschmeyer et al., 1998).
XIURENBAGRUS Chen & Lundberg, 1995
Xiurenbagrus Chen & Lundberg, 1995: 781. Type species: Liobagrus xiurenensis Yue, 1981. Type by original designation. Gender: Masculine.
Xiurenbagrus gigas Zhao, Lan & Zhang, 2004
Xiurenbagrus gigas Zhao, Lan & Zhang, 2004: 228, figs. 1–2. Type locality: Zhujiang River basin: HongshuiheRiver, Anyang Town (23º17'N, 106º29'E), Du’an County, Guangxi, P.R. China. Holotype: ASIZB 72944.
Distribution: Hongshuihe River, Pearl [= Zhujiang] River basin, China (Zhao et al., 2004).
Xiurenbagrus xiurenensis (Yue, 1981)
Liobagrus xiurenensis Yue, in Zheng, 1981: 185, fig. 152. Type locality: Xiuren River, Pearl River drainage, Xiuren,
Lipu, Guangxi, China, 24º28'N, 110º24'E. Holotype: ASIZB 751087.
Distribution: Zhujiang [= Pearl River] (Chu et al., 1999).

AMPHILIIDAE Regan, 1911
Amphiliidae Regan, 1911: 565. Type genus: Amphilius Günther, 1864.
Doumeinae Regan, 1911: 554. Type genus: Doumea Sauvage, 1879.
Leptoglaninae [sic, Leptoglanidinae] Roberts, 2003: 87. Type genus: Leptoglanis Boulenger, 1902.
Remarks: The name Amphiliidae takes precedence over the simultaneously-established Doumeinae by virtue of thehigher rank initially given to the former, following Art. 24 of the Code. Roberts (2003) and Diogo (2003b) independently proposed the new subfamily name Leptoglanidinae (incorrectly formed as Leptoglaninae by Roberts) forgroups of nearly identical membership. Roberts’s name was published earlier (March 14, vs. May [31]) and therefore has priority. He et al. (1999) questioned the monophyly of the Amphiliidae as used herein and, instead,
restricted the name to include only Amphilius and Paramphilus and included all other taxa in the Doumeidae.
Revision: Roberts (2003, Leptoglanidinae).

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FERRARISPhylogeny: Harry (1953); He et al. (1999); Diogo (2003b).
Taxonomic summary: Skelton & Teugels (1986); Skelton (1992, west Africa).
Key to genera: Poll & Gosse (1994).
12 genera, 66 species; no named fossil taxa.
AMPHILIUS Günther, 1864
Amphilius Günther, 1864: 115, 134. Type species: Pimelodus platychir Günther, 1864. Type by monotypy. Originally proposed as a section of Pimelodus. Gender: Masculine.
Anoplopterus Pfeffer, 1889: 15. Type species: Anoplopterus uranoscopus Pfeffer, 1889. Type by monotypy. Gender:
Masculine.
Chimarrhoglanis Vaillant, 1897: 81. Type species: Chimarrhoglanis leroyi Vaillant, 1897. Type by monotypy. Gender: Masculine.
Review: Skelton (1984, east and southern Africa).
Amphilius atesuensis Boulenger, 1904
Amphilius atesuensis Boulenger, 1904a: 17. Type locality: Atesu River, Gold Coast. Syntypes (4): BMNH
1903.4.24.74–77 (4).
Amphilius pictus Nichols & La Monte, 1933a: 1, fig. 1. Type locality: Stream near Kaleata, Liberia. Holotype:
AMNH 12313.
Distribution: Gbin River, Guinea, and Pra River, from Sierra Leone to Ghana (Skelton & Teugels, 1986).
Amphilius baudoni Pellegrin, 1928
Amphilius Baudoni Pellegrin, 1928c: 453, unnumbered figure. Type locality: rivière Passa (affluent de l’Ogoôué),Gabon. Syntypes (3): MNHN 1928-0224 (1), MNHN 1928-0225 (1), MNHN 1928-0226 (1). MRAC 20649
listed as a syntype in Eschmeyer et al. (1998), but not included in original description.
Amphilius baudoni uniformis Pellegrin, 1930: 200. Type locality: Riv. Koumbi (Ht. Ogôoué) ... Sangha. Syntypes(6): MNHN 1929-0238 (1), MNHN 1929-0239 (5). Originally as Amphilius baudoni var. uniformis.
Distribution: Ogowe River system and Sangha River, Congo basin (Skelton & Teugels, 1986).
Amphilius brevis Boulenger, 1902
Amphilius brevis Boulenger, 1902g: 268, pl. 29 (fig. 3). Type locality: Lindi River, which flows into the Congo at the
Stanley Falls. Syntypes (2): BMNH 1902.4.14.21 (1), IRSNB 11 (1).
Amphilius nigricaudatus Pellegrin, 1909a: 67. Type locality: Ngomo (Ogôoué). Syntypes (3): MNHN 1909-0014(1), MNHN 1909-0015 (1), MNHN 1909-0016 (1). Also described as new in Pellegrin (1909d: 47).
Amphilius nigricaudatus multipunctata Pellegrin, 1909a: 67. Type locality: Ngomo (Ogôoué). Holotype: MNHN
1909-0013. Originally as Amphilius nigricaudatus var. multipunctata. Also described as new in Pellegrin(1909d: 48, fig. 2).
Distribution: Congo River system and Leome, Kouliou, and Ogowe rivers (Skelton & Teugels, 1986).
Amphilius cryptobullatus Skelton, 1986
Amphilius cryptobullatus Skelton, 1986: 279, figs. 15–23. Type locality: Luongo River, Upper Zaire system, above
and at Route 74 crossing, 10º11'S, 29º43'E, Zambia. Holotype: ROM 46800.
Distribution: Luongo River, upper Congo River basin, Zambia (Skelton, 1986).
Amphilius jacksonii Boulenger, 1912
Amphilius jacksonii Boulenger, 1912a: 602. Type locality: the Hima River, eastern foot-hills of Ruwenzori, 3500
feet, flowing into Lake George (Ruisamba), Nile System. Holotype: BMNH 1912.10.15.47; illustrated in Boulenger (1916: 307, fig. 182), and Seegers (1996: 191, fig. 133).
Distribution: Widely distributed in central and eastern Africa (Seegers, 1996).
Remarks: Species redescribed in Walsh et al. (2000).
Amphilius kakrimensis Teugels, Skelton & Lévêque, 1987
Amphilius kakrimensis Teugels, Skelton & Lévêque, 1987: 94, figs. 1–2. Type locality: a small tributary of the
Kakrima River, Konkoure basin, at Kasseri (10º16'N, 12º28'W), Guinea. Holotype: MNHN 1986-0599.
Distribution: Konkoure River basin, Guinea (Teugels et al., 1987).
Amphilius kivuensis Pellegrin, 1933

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Amphilius kivuensis Pellegrin, 1933b: 172. Type locality: Boulira, Kitembo (ouest du Kivu), ... Kilembo (ouest duKivu, altitude 2000 mètres). Syntypes (8): MNHN 1932-0038 (5), MNHN 1933-0009 (1), MNHN 1933-0010(1), MNHN 1933-0011 (1).
Distribution: Rivers to west of Lake Kivu; Ruzizi River and rivers entering Lake Tanganyika (Skelton & Teugels,
1986).
Amphilius lamani Lönnberg & Rendahl, 1920
Amphilius lamani Lönnberg & Rendahl, 1920: 171, figs. 2–3. Type locality: Kingoyi, Lower Congo. Syntypes:
NRM 9910 (2).
Distribution: Lower Congo; known only from type locality (Skelton & Teugels, 1986).
Amphilius lampei Pietschmann, 1913
Amphilius lampei Pietschmann, 1913b: 190, pl. 2 (fig. 1). Type locality: Harrar, Abessynien [Ethiopia]. Holotype: at
Weisbaden Museums.
Distribution: Webi Shebeli tributaries, near Harrar, Ethiopia (Skelton & Teugels, 1986).
Amphilius laticaudatus Skelton, 1984
Amphilius laticaudatus Skelton, 1984: 61, fig. Type locality: Buzi River, at bridge on Inchope to Lourenço Marques
road (19º55'S, 34º15'E). Holotype: AMG P5815.
Distribution: Buzi River system, Moçambique (Skelton, 1993).
Amphilius lentiginosus Trewavas, 1936
Amphilius lentiginosus Trewavas, 1936: 71, pl. 2 (figs. 10–11). Type locality: a brook at Cuito, Mt. Moco, Luimbale,Cuvo River system, Angola. Syntypes: BMNH 1935.3.20.189–190 (2).
Distribution: Cuvo River system, Luachimo River and Tchimenji River, Angola (Skelton & Teugels, 1986).
Amphilius longirostris (Boulenger, 1901)
Anoplopterus longirostris Boulenger, 1901f: 447. Type locality: Hills in the Bulu country, some 75 miles in the interior from Batanga, at an elevation of between 1500–2000 feet, Cameroon. Holotype: BMNH 1901.11.12.133.
Distribution: Cameroon and Equatorial Guinea, also tributaries of Ngoko, Congo River system, in Cameroon (Skelton & Teugels, 1986).
Remarks: Specimen listed as holotype of Anoplopterus longirostris is entered in BMNH register with locality onlyas Efulen, and Boulenger (1911: 260) indicated that the locality stated in original description was near Efulen.
Amphilius maesii Boulenger, 1919
Amphilius Maesii Boulenger, 1919: 160. Type locality: Oshwe [Congo River system]. Syntypes (7): BMNH
1919.8.28.14–16 (3), MRAC 3180-83 (4).
Distribution: Congo River basin (Skelton & Teugels, 1986).
Amphilius natalensis Boulenger, 1917
Amphilius natalensis Boulenger, 1917a: 432. Type locality: Krantz Kloof, Natal. Syntypes (2): BMNH 1917.7.21.1(1).
Distribution: Ruo River, Malawi; Zambezi system of Zimbabwe, south to Umkomaas River (Natal, South Africa)(Skelton, 1993).
Remarks: Redescribed in Barnard (1943).
Amphilius opisthophthalmus Boulenger, 1919
Amphilius opisthophthalmus Boulenger, 1919: 159. Type locality: Oshwe, Lac Léopold II. Syntypes (7): BMNH
1919.8.28.17–18 (2), MRAC 3191-94 (4).
Distribution: Oshwe on the Lukenie River (Lake Mai-Ndombe) (Skelton & Teugels, 1986).
Amphilius platychir (Günther, 1864)
Pimelodus platychir Günther, 1864: 134. Type locality: Sierra Leone. Syntypes (4): BMNH 2001.12.5.574–577 (4),MNHN 1898-0121 (1).
Amphilius grammatophorus Pellegrin, 1913a: 237, fig. 1. Type locality: Rivière Kikoulo, Guinée française. Syntypes: MNHN 1913-0231 (1), MNHN 1913-0232 (1), MNHN 1913-0233 (1), MNHN 1913-0234 (1), MNHN
1913-0235 (1).
Amphilius grammatophorus brevipinna Pellegrin, 1935: 465. Type locality: Banamanan (cercle de Kissidougou), ...
Friguiagbé (près Kindia); Guinée française. Syntypes (2): MNHN 1935-0194 (1), MNHN 1935-0195 (1). Orig-

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inally as Amphilius grammatophorus var. brevipinna. nov. var.
Amphilius grammatophorus inaequalis Pellegrin, 1935: 464. Type locality: Friguiagbé (près Kindia) Guinéefrançaise. Syntypes (6): MNHN 1935-0191 (1), MNHN 1935-0192 (1), MNHN 1935-0193 (4). Originally as
Amphilius grammatophorus var. inaequalis nov.var.
Amphilius grammatophorus marmoratus Daget, 1963: 584. Type locality: Kana, alt. 600 m, Mont Nimba. Holotype:
MNHN 1960-0119.
Distribution: Sierra Leone, Guinea, Côte d’Ivoire (Skelton & Teugels, 1986).
Amphilius pulcher Pellegrin, 1929
Amphilius pulcher Pellegrin, 1929c: 640, fig. 1. Type locality: rivière Louessé (fleuve Kouilou) [Gabon]. Syntypes(3): MNHN 1929-0224 (3). MRAC 20650 listed as a syntype in Eschmeyer et al. (1998), but not listed in
description.
Amphilius pulcher ephippiata Pellegrin, 1930: 200. Type locality: Sebé (haut Ogoôué). Syntypes: MNHN 19290237 (9). Originally as Amphilius pulcher var. ephippiata nov. var.
Distribution: Ogowe and Nyanga River systems, Gabon (Skelton & Teugels, 1986).
Amphilius rheophilus Daget, 1959
Amphilius rheophilus Daget, 1959: 692, fig. 3. Type locality: Gambie à Mpantié (parc national du Niokolo-Koba);
Gambie à Banharé. Syntypes (3): MNHN 1959-0147 (2), MNHN 1960-0486 (1).
Distribution: Gambia River system (Skelton & Teugels, 1986).
Amphilius uranoscopus (Pfeffer, 1889)
Anoplopterus uranoscopus Pfeffer, 1889: 16; illustrated in Pfeffer (1896: 31, fig. 14). Type locality: Bad bei Ushonda (Ungúu), ... Bäche bei Mhonda. Lectotype: ZMH 11944, selected by, and illustrated in, Seegers (1996:
192, fig. 137).
Chimarrhoglanis Leroyi Vaillant, 1897: 82. Type locality: Zanguebar, torrent de Mrogoro, par 600 mètres d’altitude.
Holotype: MNHN 1897-0003.
Amphilius grandis Boulenger, 1905c: 63, pl. 7 (figs. 3, 3a). Type locality: Chania River of Tetse, Tana system, in
cold water at an altitude of 7000 feet. Syntypes (3): BMNH 1904.12.23.50–52 (3).
Amphilius Hargeri Boulenger, 1907c: 488. Type locality: Mlangi, British Central Africa. Holotype: BMNH
1907.10.14.8.
Amphilius krefftii Boulenger, 1911a: 356, fig. 276. Type locality: Usambara, in rivulet running down from Anani
Hills to Sigi River. Syntypes: BMNH 1909.10.19.26–27 (2).
Amphilius oxyrhinus Boulenger, 1912c: 675, pl. 80. Type locality: Eusso Mara, a swift mountain stream, tributary of
Eusso Nyiro, British East Africa. Holotype: BMNH 1912.3.22.120.
Amphilius brevidorsalis Pellegrin, 1919a: 399. Type locality: Rivière Revue (Mozambique). Holotype: MNHN
1919-0488.
Amphilius platychir cubangoensis Pellegrin, 1936a: 56. Type locality: Cubango près de Vila da Ponte [Angola].
Syntypes (9): MHNG 856.86 (1), MNHN 1936-0101 (1), MNHN 1936-0102 (1), MNHN 1936-0103 (3),
MRAC 138769, NMBA 5213 (1). Originally as Amphilius platychir var. cubangoensis.
Distribution: Okavango and Zambezi basins, eastern coastal Rivers south to the Mkuze basin, Natal, and throughout
east and Central Africa (Skelton, 1993); Lake Rukwa basin (Seegers, 1996).
Remarks: The name Amphilius transvaaliensis that was published in the caption to figure 1 in van der Horst, (1931:247) fails to qualify as an available name, following the provisions in Art. 13.1.1 of the Code. In addition, there
is uncertainty about whether the name was intended to be published, inasmuch as copies of this paper at CAS
and CU have the figure caption crossed out and replaced with the name Amphilius grandis.
Amphilius zairensis Skelton, 1986
Amphilius zairensis Skelton, 1986: 266, fig. 1. Type locality: River Lufu, about two km from confluence with Zaire
River, near Inga hydroelectric dam, 5º31.5'S, 13º37.5'E. Holotype: MCZ 50533.
Distribution: Lufu River, Congo River basin (Skelton, 1986).
ANDERSONIA Boulenger, 1900
Andersonia Boulenger, 1900d: 528. Type species: Andersonia leptura Boulenger, 1900. Type by monotypy. Gender:

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Feminine.
Slatinia Werner, 1906: 326. Type species: Slatinia mongallensis Werner, 1906. Type by monotypy. Gender: Feminine.
Andersonia leptura Boulenger, 1900
Andersonia leptura Boulenger, 1900d: 529. Type locality: a pond near Koshek, Soudan. Holotype: BMNH
1907.12.2.2545.
Slatinia mongallensis Werner, 1906: 327. Type locality: Bahar-el-Gebel, bei Mongalla. Holotype: NMW 79741.
Andersonia brevior Boulenger, 1918: 426. Type locality: Shari River, Ubanghi-Shari Colony, French Equatorial
Africa. Holotype: BMNH 1918.11.12.13.
Andersonia pellegrini Boulenger, 1918: 427. Type locality: Shari River, Ubanghi-Shari Colony, French Equatorial
Africa. Holotype: BMNH 1918.11.12.13.
Distribution: Omo, Niger and Upper Nile Rivers, Lake Chad basin (Skelton & Teugels, 1986).
Remarks: The name "Andersonia brevior sp. n." appears in the introduction of Boulenger (1918: 426) for a species
described on the next page as A. pellegrini. Harry (1953) appears to be the first to have commented on the two
names for the species and, therein, he treated A. pellegrini as valid.
BELONOGLANIS Boulenger, 1902
Belonoglanis Boulenger, 1902d: 50. Type species: Belonoglanis tenuis Boulenger, 1902. Type by monotypy. Gender: Masculine.
Belonoglanis brieni Poll, 1959
Belonoglanis brieni Poll, 1959: 107, pl. 22 (fig. 3). Type locality: Stanley Pool, Rivière N'Djili. Holotype: MRAC
118539.
Distribution: Congo River basin (Skelton & Teugels, 1986).
Belonoglanis tenuis Boulenger, 1902
Belonoglanis tenuis Boulenger, 1902d: 50, pl. 14 (fig. 4). Type locality: Ubangi à Banzyville. Holotype: MRAC
1289.
Belonoglanis nudipectus Lönnberg & Rendahl, 1920: 175, fig. 6. Type locality: Lower Congo. Holotype: NRM
16118.
Belonoglanis curvirostris Pellegrin, 1922a: 74. Type locality: Bangui (Oubanghi). Syntypes: MNHN 1921-0428 (1),
MNHN 1921-0429 (1).
Distribution: Congo River basin (Skelton & Teugels, 1986).
DOLICHAMPHILIUS Roberts, 2003
Dolichamphilius Roberts, 2003: 100. Type species: Leptoglanis brieni Poll, 1959. Type by original designation.
Gender: Masculine.
Dolichamphilius brieni (Poll, 1959)
Leptoglanis brieni Poll, 1959: 96, pl. 24 (fig. 2). Type locality: Stanley Pool, passe devant le refuge Jipo. Holotype:
MRAC 118503.
Distribution: Stanley Pool, Congo River (Roberts, 2003).
Dolichamphilius longiceps Roberts, 2003
Dolichamphilius longiceps Roberts, 2003: 100. Type locality: Republic du Congo (Kinshasa): Chutes Wagenia,
Kisangani. Holotype: MRAC 90-47-P-704-708.
Distribution: Chutes Wagenia on the Lualaba River, near Kisangani (Roberts, 2003).
DOUMEA Sauvage, 1879
Doumea Sauvage, 1879: 96.Type species: Doumea typica Sauvage, 1879. Type by original designation. Gender:
Feminine.
Doumea alula Nichols & Griscom, 1917
Doumea alula Nichols & Griscom, 1917: 722, fig. 27. Type locality: Stanleyville, Congo. Holotype: AMNH 6531.
Distribution: Congo River basin (Skelton & Teugels, 1986).

24 · Zootaxa 1418 © 2007 Magnolia Press

FERRARISDoumea angolensis Boulenger, 1906
Doumea angolensis Boulenger, 1906e: 347. Type locality: the interior of Benguella, at an altitude of 4000–5000
feet, Angola. Holotype: BMNH 1906.10.8.12.
Distribution: Quanza River system and the interior of Benguella, Angola (Skelton & Teugels, 1986).
Doumea chappuisi Pellegrin, 1933
Doumea chappuisi Pellegrin, 1933a: 114, fig. on p. 115. Type locality: Danané (Côte d’Ivoire). Syntypes (8):MNHN 1932-0302 (1), NMBA 4281-87 (7).
Distribution: Guinea, Liberia and Côte d’Ivoire; Cavally, Nuon, St. Paul, and Koumba River systems (Skelton &Teugels, 1986).
Doumea thysi Skelton, 1989
Doumea thysi Skelton, 1989: 6, fig. 3. Type locality: Cross River, 27 miles from Mamfe towards Eyomojak, Cameroon, 5º46'N, 9º03'E. Holotype: MRAC 76-32-P-2260.
Distribution: Cross River basin, Cameroon (Skelton, 1989).
Doumea typica Sauvage, 1879
Doumea typica Sauvage, 1879: 97. Type locality: Doumé [rivière Ogôoué]. Holotype: MNHN a-0965. Described in
more detail and illustrated in Sauvage (1880c: 41, pl. 3, fig. 1).
Distribution: Congo River basin and coastal rivers from Cameroon to Congo (Skelton & Teugels, 1986).
LEPTOGLANIS Boulenger, 1902
Leptoglanis Boulenger, 1902d: 42. Type species: Leptoglanis xenognathus Boulenger, 1902. Type by monotypy.
Gender: Masculine.
Remarks: Roberts (2003) removed most species from Leptoglanis and placed them, instead, into Zairichthys.
Leptoglanis xenognathus Boulenger, 1902
Leptoglanis xenognathus Boulenger, 1902d: 43, pl. 14 (fig. 1). Type locality: l’Ubangi à Banzyville. Holotype:
MRAC 1287.
Distribution: Congo basin.
PARAMPHILIUS Pellegrin, 1907
Paramphilius Pellegrin, 1907a: 23. Type species: Paramphilius trichomycteroides Pellegrin, 1907. Type by original
designation. Gender: Masculine.
Paramphilius firestonei Schultz, 1942
Paramphilius firestonei Schultz, 1942: 328, 333, pl. 36 (fig. 3). Type locality: Bromley, Harbel [Liberia]. Holotype:
USNM 118811.
Distribution: St. Paul, Du and Borlor Rivers, Liberia (Skelton & Teugels, 1986).
Paramphilius goodi Harry, 1953
Paramphilius goodi Harry, 1953: 203, figs. 1–2. Type locality: Lolodorf, Bikui River, Lokunje River system, Cameroon, West Africa. Holotype: SU 16970.
Distribution: Lokunje River basin, Cameroon (Skelton & Teugels, 1986).
Paramphilius teugelsi Skelton, 1989
Paramphilius teugelsi Skelton, 1989: 2, fig. Type locality: ruisseau affluent de Mamou (Little Scarcies River system), Guinea, west Africa, approximately 10º09'N, 11º50'W. Holotype: MNHN 1961-0601.
Distribution: Little Scarcies River basin, Guinea (Skelton, 1989).
Paramphilius trichomycteroides Pellegrin, 1907
Paramphilius trichomycteroides Pellegrin, 1907a: 24. Type locality: Ditinn (Fouta-Djalon). Holotype: MNHN 19060343.
Distribution: Senegal River headwaters, near Ditinn and Mamou, Guinea (Skelton & Teugels, 1986).
Remarks: Holotype catalog number of Paramphilius trichomycteroides incorrectly reported as MNHN 1906-0243 in
Eschmeyer et al. (1998).

CHECKLIST OF CATFISHES, RECENT AND FOSSIL

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PHRACTURA Boulenger, 1900
Peltura Perugia, 1892: 972. Type species: Peltura bovei Perugia, 1892. Type by monotypy. Preoccupied by Peltura
Agassiz, 1846, in trilobites; replaced by Phractura Boulenger, 1900. Gender: Feminine.
Phractura Boulenger, 1900d: 527. Type species: Peltura bovei Perugia, 1892. Type by being a replacement name.
Replacement for Peltura Perugia, 1892, preoccupied by Peltura Agassiz, 1846. Gender: Feminine.
Paraphractura Boulenger, 1902d: 47. Type species: Paraphractura tenuicauda Boulenger, 1902. Type by monotypy. Gender: Feminine.
Phractura ansorgii Boulenger, 1902
Phractura ansorgii Boulenger, 1902h: 623, pl. 37 (fig. 1). Type locality: Agberi [Nigeria]. Holotype: BMNH
1901.12.18.7.
Distribution: Lower Niger River (Skelton & Teugels, 1986).
Phractura bovei (Perugia, 1892)
Peltura Bovei Perugia, 1892: 972, unnumbered figures. Type locality: Congo. Holotype: MSNG 8839.
Distribution: Lower Congo River (Skelton & Teugels, 1986).
Phractura brevicauda Boulenger, 1911
Phractura brevicauda Boulenger, 1911a: 502, fig. 374. Type locality: Trib. of Lobi R., S. W. of Efulen, and Kribi R.,
South Cameroon; Abanga R., Ogowe. Syntypes: BMNH 1904.7.1.119-126 (8), BMNH 1908.5.30.176–177 (2),
BMNH 1908.5.25.118 (1).
Distribution: Lower Congo River (Skelton & Teugels, 1986).
Remarks: Illustrated specimen, from the Lobi River, may not be considered a holotype, even though labeled as“type” in figure caption inasmuch as the term type is also applied in the text to the single specimen from the
Abanga River.
Phractura clauseni Daget & Stauch, 1963
Phractura clauseni Daget & Stauch, 1963: 95, fig. 2. Type locality: Haute Comoé à Samago Iri. Holotype: MNHN
1960-0148.
Distribution: Tributaries of the Comoe and Black Volta Rivers, Upper Volta, Ogun River, Nigeria, and Upper
Benoue River, Cameroon (Skelton & Teugels, 1986).
Phractura fasciata Boulenger, 1920
Phractura fasciata Boulenger, 1920a: 34, fig. 18. Type locality: Stanley-Falls. Syntypes (4): BMNH 1919.9.10.284–
285 (2), MRAC 7241–42 (2).
Distribution: Congo River basin (Skelton & Teugels, 1986).
Phractura gladysae Pellegrin, 1931
Phractura Gladysae Pellegrin, 1931a: 209. Type locality: Haute-Louessé. Syntypes (3): MNHN 1930-0251(1),
MNHN 1930-0252 (1), MNHN 1930-0253 (1).
Distribution: Kouilou River system; known only from type locality (Skelton & Teugels, 1986).
Phractura intermedia Boulenger, 1911
Phractura intermedia Boulenger, 1911a: 503, fig. 375. Type locality: Ja R., at Bitye, Libi R. affl. of Ja R., Kribi R,
and Nong R.[South Cameroon]. Syntypes: BMNH 1903.7.28.114–116 (3), BMNH 1903.7.28.209–210 (2),
BMNH 1904.7.1.165–167 (3), BMNH 1904.7.1.213 (1), BMNH 1904.10.26.49–52 (4), BMNH 1907.5.22.187(1), BMNH 1909.4.29.103–105 (3), BMNH 1909.12.3.28 (1).
Distribution: Coastal Rivers of Cameroon, from Nyong to Kouilou (Skelton & Teugels, 1986).
Phractura lindica Boulenger, 1902
Phractura lindica Boulenger, 1902g: 268, pl. 28 (figs. 3, 3a, 3b). Type locality: Lindi River, which flows into theCongo at the Stanley Falls. Syntypes (4): BMNH 1902.4.14.25 (1), IRSNB 12 (2 or 3), MSNM 39 (1).
Phractura lukugae Boulenger, 1920c: 402, fig. 9. Type locality: Kabeke, a village 30 miles south of Tumbwe, on the
Niemba River, near the west coast of Lake Tanganyika. Holotype: BMNH 1919.7.24.32.
Distribution: Congo River basin (Skelton & Teugels, 1986).
Phractura longicauda Boulenger, 1903
Phractura longicauda Boulenger, 1903c: 439. Type locality: Kribi River, southern Cameroon. Holotype: BMNH
1903.7.28.117.

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FERRARISDistribution: Coastal rivers of South Cameroon and Equatorial Guinea, and Congo River basin (Skelton & Teugels,
1986).
Phractura macrura Poll, 1967
Phractura macrura Poll, 1967: 269, fig. 129. Type locality: Caungula, riv. Cassange, (8º07'S, 18º09'E), affl. riv.
Uamba, Angola. Holotype: MD 6272.
Distribution: Uamba and Cuilo Rivers, southern tributaries of Kasai, Congo River basin, Angola (Skelton & Teugels, 1986).
Phractura scaphyrhynchura (Vaillant, 1886)
Doumea scaphyrhynchura Vaillant, in Rivière, 1886: 18. Type locality: Ogôoué à Diélé. Holotype: MNHN 18860424.
Distribution: Ogowe and Congo River basins (Skelton & Teugels, 1986).
Remarks: Pellegrin (1900: 181), lists Phractura scaphyrhynchura as a new species (nov. sp.), but cites Vaillant’s
description in the text.
Phractura tenuicauda (Boulenger, 1902)
Paraphractura tenuicauda Boulenger, 1902d: 48, pl. 14 (fig. 2). Type locality: l’Ubangi à Banzyville. Syntypes(plusieurs): BMNH 1901.12.26.48–49 (2), MRAC 1297–1299 (3).
Distribution: Congo River basin (Skelton & Teugels, 1986).
PSAMMPHILETRIA Roberts, 2003
Psammphiletria Roberts, 2003: 101. Type species: Psammphiletria nasuta Roberts, 2003. Type by original designation. Gender: Feminine.
Psammphiletria delicata Roberts, 2003
Psammphiletria delicata Roberts, 2003: 102, fig. 17 e. Type locality: Stanley Pool. Holotype: MRAC 118455.
Distribution: Stanley Pool, Congo River (Roberts, 2003).
Psammphiletria nasuta Roberts, 2003
Psammphiletria nasuta Roberts, 2003: 101, fig. 17 a–c. Type locality: Republique Centrafricaine: Central African
Republic, sandy riffle in R. Ubangui near Banda, 72 km upriver from Bangui. Holotype: CAS 92624.
Distribution: Ubangui River above Bangui (Roberts, 2003).
TETRACAMPHILIUS Roberts, 2003
Tetracamphilius Roberts, 2003: 102. Type species: Tetracamphilius pectinatus Roberts, 2003. Type by original designation. Gender: Masculine.
Tetracamphilius angustifrons (Boulenger, 1902)
Anoplopterus angustifrons Boulenger, 1902d: 42, pl. 10 (fig. 4). Type locality: Banzyville [Ubangi River]. Syntypes(plusieurs): BMNH 1901.12.26.39–40 (2), MRAC 1313 (1).
Distribution: Ubangi River, Congo Basin (Skelton & Teugels, 1986).
Tetracamphilius clandestinus Roberts, 2003
Tetracamphilius clandestinus Roberts, 2003: 106, fig. 20 b. Type locality: Republique Centrafricaine: riffles in shallow sandy area of Ubangui near Banda. 2 km upstream from Bangui. Holotype: CAS 92653.
Distribution: Ubangui River above Bangui (Roberts, 2003).
Tetracamphilius notatus (Nichols & Griscom, 1917)
Amphilius notatus Nichols & Griscom, 1917: 715, fig. 24. Type locality: Faradje, Congo. Holotype: AMNH 6711.
Distribution: Faradje, Uele River, Congo River basin (Skelton & Teugels, 1986).
Tetracamphilius pectinatus Roberts, 2003
Tetracamphilius pectinatus Roberts, 2003: 104, fig. 18. Type locality: Republique du Congo (Kinshasa): Congobasin, tributary of R. Luala, 29 km N of Kibunzi (8 km S of turnoff to Kubunzi, Buozi dist.), 45º6.5'S, 13º48'E.
Holotype: CAS 92627.
Distribution: Ubangui River above Bangui; Luala River tributary; and Luala River of Southern Congo (Roberts,
2003).

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Zootaxa 1418 © 2007 Magnolia Press · 27

TRACHYGLANIS Boulenger, 1902
Trachyglanis Boulenger, 1902d: 48. Type species: Trachyglanis minutus Boulenger, 1902. Type by monotypy. Gender: Masculine.
Trachyglanis ineac (Poll, 1954)
Phractura ineac Poll, 1954: 62, fig. 5. Type locality: riv. Loweo, environs de Yangambi (Stanleyville). Holotype:
MRAC 77460.
Distribution: Congo River basin (Yangambi, Tohuapa) (Skelton & Teugels, 1986).
Trachyglanis intermedius Pellegrin, 1928
Trachyglanis intermedius Pellegrin, 1928b: 110. Type locality: Kasai, Congo belge. Holotype: NMBA 3755.
Distribution: Kasai River, Congo River basin (Skelton & Teugels, 1986).
Trachyglanis minutus Boulenger, 1902
Trachyglanis minutus Boulenger, 1902d: 49, pl. 14 (fig. 3). Type locality: l’Ubangi à Banzyville. Syntypes (4):
BMNH 1901.12.26.50–51 (2), MRAC 1291–1291A (2).
Distribution: Congo River basin (Skelton & Teugels, 1986).
Trachyglanis sanghensis Pellegrin, 1925
Trachyglanis sanghensis Pellegrin, 1925: 103. Type locality: Ouesso (Sangha). Holotype: MNHN 1925-0145.
Distribution: Sanga River, Congo River basin (Skelton & Teugels, 1986).
ZAIREICHTHYS Roberts, 1967
Zaireichthys Roberts, 1967: 124. Type species: Zaireichthys zonatus Roberts, 1967. Type by original designation.
Gender: Masculine.
Revision: Roberts (2003).
Zaireichthys brevis (Boulenger, 1915)
Leptoglanis brevis Boulenger, 1915: 169. Type locality: Rivière Lubumbashi, à Elisabethville. Syntypes (2): BMNH
1920.5.26.93 (1), MRAC 12702 (1).
Distribution: Congo River basin.
Remarks: Placed by Roberts (2003: 98) in the synonymy of Zairichthys rotundiceps, but comments therein indicate
only that the types are “closely similar, possibly conspecific [with Z. rotundiceps].” Therefore, Zaireichthys
brevis is retained as valid herein.
Zaireichthys camerunensis (Daget & Stauch, 1963)
Leptoglanis camerunensis Daget & Stauch, 1963: 94, fig. 1. Type locality: Bénoué à Lakdo, Cameroun. Syntypes(7): MNHN 1962-1272 (6); one specimen lost.
Distribution: Upper Benue River, Cameroon and Upper Niger River, Guinea (Roberts, 2003).
Zaireichthys dorae (Poll, 1967)
Leptoglanis dorae Poll, 1967: 211, fig. 95. Type locality: rapides de la Luachimo, dans des flaques d’eau résiduelle,à l’occasion de la mise à sec del la partie d’aval du barrage, Angola. Holotype: MRAC 161646.
Distribution: Luachimo, Congo Basin, Angola (Roberts, 2003).
Zaireichthys flavomaculatus (Pellegrin, 1926)
Leptoglanis flavomaculatus Pellegrin, 1926: 204. Type locality: Kamaiembi. Holotype: MRAC 19721.
Distribution: Lulua River, southern Congo; known only from holotype (Roberts, 2003).
Zaireichthys heterurus Roberts, 2003
Zaireichthys heterurus Roberts, 2003: 96. Type locality: Republic du Congo (Kinshasa): R. Avokoko, affluent rive
droit fleuve Zaire, km 21 route Kisangani-Wanie Rukula. Holotype: MRAC 87-42-P-1140.
Distribution: Eastern Congo River basin (Roberts, 2003).
Zaireichthys mandevillei (Poll, 1959)
Leptoglanis mandevillei Poll, 1959: 98, pl. 25 (fig. 2). Type locality: Stanley Pool (Congo R., near Kinshasa). Holotype: MRAC 118532.
Distribution: Stanley Pool, Congo River, Ubangui and Lualaba (Roberts, 2003).
Zaireichthys rotundiceps (Hilgendorf, 1905)
Gephyroglanis rotundiceps Hilgendorf, 1905: 412. Type locality: Im Bubu bei Irangi. Lectotype: ZMB 16392, des-28 · Zootaxa 1418 © 2007 Magnolia Press

FERRARIS

ignated by, and illustrated in, Seegers (1996: 196, fig. 141).
Distribution: Cunene, Okavango and Zambezi, Pungwe, Buse and Save basins, and Lake Malawi basin (Skelton,1993); Lake Rukwa basin (Seegers, 1996).
Zaireichthys wamiensis (Seegers, 1989)
Leptoglanis wamiensis Seegers, 1989: 285, unnumbered figure. Type locality: Kisangata -Bach bei Mvumi, 32 kilometer SW Kidete, am Wege nach Kimamba/Kilosa im Wami-Einzug NW Morogoro, Tanzania. Holotype:
ZFMK 15899.
Distribution: Wami River basin, Tanzania (Seegers, 1989).
Zaireichthys zonatus Roberts, 1967
Zaireichthys zonatus Roberts, 1967: 124, figs. 3–4. Type locality: Side channel of the lower rapids of the Congo
River, below Stanley Pool, at Kinsuka village, within Leopoldville city limits. Holotype: SU 64126.
Distribution: Congo River basin (Roberts, 2003).

ANCHARIIDAE Glaw & Vences, 1994
Anchariidae Glaw & Vences, 1994: 380. Type genus: Ancharius Steindachner, 1880.
Revision, with key to species: Ng & Sparks (2005).
2 genera, 5 species; no named fossil taxa.
ANCHARIUS Steindachner, 1880
Ancharius Steindachner, 1880a: 158. Type species: Ancharius fuscus Steindachner, 1880. Type by monotypy. Gender: Masculine. Also proposed as new in Steindachner (1880c: 251).
Ancharius fuscus Steindachner, 1880
Ancharius fuscus Steindachner, 1880a: 158. Type locality: Tohizona auf Madagascar. Holotype: NMW 48093. Species illustrated and described in more detail in Steindachner (1880c: 251, pl. 3, fig. 3; pl. 4).
Distribution: Widely distributed along eastern-draining rivers of Madagascar (Ng & Sparks, 2005).
Ancharius griseus Ng & Sparks, 2005
Ancharius griseus Ng & Sparks, 2005: 308, fig. 5. Type locality: Madagascar: Toliara Province, Ihazofotsy Rivernear Isalo, approximately 2-hour hike from Sakalama Village, 22º58'56"S 47º22'7"E. Holotype: UMMZ
238041.
Distribution: Onilahy River basin, western Madagascar (Ng & Sparks, 2005).
GOGO Ng & Sparks, 2005
Gogo Ng & Sparks, 2005: 311. Type species: Gogo ornatus Ng & Sparks, 2005. Type by original designation. Gender: Masculine.
Gogo arcuatus Ng & Sparks, 2005
Gogo arcuatus Ng & Sparks, 2005: 313, fig. 10. Type locality: Madagascar: Fianarantsoa province, Andriambondro
River, 16 km N of Karianga at town of Mahevelo (tributary of Rienana River) 22º21'47"S, 47º22'5"E. Holotype:
UMMZ 238042.
Distribution: Sandrananta River basin, Madagascar (Ng & Sparks, 2005).
Gogo brevibarbis (Boulenger, 1911)
Ancharius brevibarbis Boulenger, 1911a: 380, fig. 294. Type locality: Ambohimango, Madagascar. Holotype:
BMNH 1896.10.9.40. Holotype illustrated in Ng & Sparks (2005: 313, fig. 8).
Distribution: Apparently in Mananjary River basin, Madagascar (Ng & Sparks, 2005); known only from holotype.
Gogo ornatus Ng & Sparks, 2005
Gogo ornatus Ng & Sparks, 2005: 316, fig. 12. Type locality: Madagascar: Toamasina province, Mangoro River,
200 km upstream of ferry crossing on road to Marolambo. Holotype: AMNH 235213.
Distribution: Mangoro River basin, Madagascar (Ng & Sparks, 2005).

CHECKLIST OF CATFISHES, RECENT AND FOSSIL

Zootaxa 1418 © 2007 Magnolia Press · 29† ANDINICHTHYIDAE Gayet, 1988
† Andinichthyidae Gayet, 1988: 833. Type genus: † Andinichthys Gayet, 1988.
Diagnosis: Gayet & Meunier (2003).
Review: Arratia & Gayet (1995).
3 genera, 3 species; 3 named fossil taxa.
† ANDINICHTHYS Gayet, 1988
† Andinichthys Gayet, 1988: 833. Type species: †Andinichthys bolivianensis Gayet, 1988. Type by original designation. Gender: Masculine.
† Andinichthys bolivianensis Gayet, 1988
† Andinichthys bolivianensis Gayet, 1988: 834, fig. on p. 834. Type locality: Maastrichtien de la Formation El
Molino, Tiupampa (Bolivie). Holotype: MNHN Institut de Paléontologie YPFB-PAL 6178 (partial neurocranium).
Distribution: El Molino Formation, Tiupampa, Bolivia; Maastrichtian (Gayet, 1988).
† HOFFSTETTERICHTHYS Gayet, 1990
† Hoffstetterichthys Gayet, 1990: 871. Type species: †Hoffstetterichthys pucai Gayet, 1990. Type by original designation. Gender: Masculine.
Remarks: Placement in †Andinichthyidae follows Gayet (1991).
† Hoffstetterichthys pucai Gayet, 1990
† Hoffstetterichthys pucai Gayet, 1990: 871, figs. 2, 3. Type locality: Maastrichtien de Tiupampa, Province deMizqué, Département de Cochabamba (Bolivie centrale). Holotype: MHNC 6252 (partial neurocranium).
† INCAICHTHYS Gayet, 1990
† Incaichthys Gayet, 1990: 868. Type species: † Incaichthys suarezi Gayet, 1990. Type by original designation. Gender: Masculine.
Remarks: Placement in † Andinichthyidae follows Gayet (1991).
† Incaichthys suarezi Gayet, 1990
† Incaichthys suarezi Gayet, 1990: 868, figs 1, 4. Type locality: Maastrichtien de Tiupampa, Province de Mizqué,
Département de Cochabamba (Bolivie centrale). Holotype: MHNC 6253 (partial neurocranium).

ARIIDAE Bleeker, 1862
Hemipimelodinae Gill, 1861c: 46. Type genus: Hemipimelodus Bleeker, 1857.
Arii Bleeker, 1862 (in Bleeker, 1862–63): 7, 25. Type genus: Arius Valenciennes, 1840.
Batrachocephalinae Gill, 1893:132. Type genus: Batrachocephalus Bleeker, 1846.
Doiichthyidae Weber, 1913: 532. Type genus: Doiichthys Weber, 1913.
Bagreidae Schultz, 1944c: 182. Type genus: Bagre Oken, 1817.
Remarks: The classification of species within the Ariidae is arguably the most poorly resolved of any catfish family. In
recent years, there have been three independent attempts to examine the phylogeny of the family (Marceniuk, 2003;
Kailola, 2004; and Betancur et al., 2004) and produce classifications that reflect the resulting phylogenetic hypotheses. The phylogenies and their resulting classifications differ markedly from each other, which makes it quite difficult to summarize the current understanding of the taxonomy herein. Thus, many of the generic assignments andsynonymies listed here must be considered as tentative. One theme that is repeated in the various studies on the Ariidae is that the genus Arius was previously a taxonomic wastebasket and should instead be restricted to a group of
Indo-Pacific species. Thus, species that had been treated as valid in Arius from the Americas, or western Africa, arenow placed in other genera or, when no clear alternative was indicated, as Incertae sedis, herein. The family-level
name Osteogeneiosinae was proposed by Fowler (1951b: 3) in a manner that made it available when published but is
now not available (see Ferraris & de Pinna, 1999, for details). The name Arius cous, which appeared in Heckel

30 · Zootaxa 1418 © 2007 Magnolia Press

FERRARIS

(1843) and also in Hyrtl (1859) appears to have been based on Silurus cous Linnaeus (now Glyptothorax cous) andis not an available name even though it has been treated as such.
Taxonomic summaries: Taylor (1986a, 1986b, Africa); Taylor (1990, eastern tropical Atlantic); Marceniuk & Ferraris(2003, Neotropics).
Reviews: Taylor & Menezes (1978, western central Atlantic) and Acero (2002, western central Atlantic); Jayaram &Dhanze (1979, South Asia); Jayaram (1983, western Indian Ocean); Kailola (1990, freshwaters of northern New
Guinea); Daget (1992, West Africa); Kailola & Bussing (1995, Eastern Pacific); Kailola (1999, Western Pacific).
Phylogeny: Betancur-R. (2003), Betancur-R. et al. (2004, Neotropics); Kailola (2004, Indo-Pacific).
Evolution: Jayaram & Dhanze (1985, South Asia).
Identification guide: Jayaram (1982, South Asia).
Biogeography: Cione et al. (1996, southern South America).
Keys: Kailola (2004: 123, genera); Marceniuk (2005a, Brazil).
26 genera, 133 species; 13 fossil species, not including otolith-based taxa.
Incertae sedis
Arius acutirostris Day, 1877
Arius acutirostris Day, 1877 (in Day, 1875–78): 459, pl. 107 (fig. 1). Type locality: Salwein River at Moulmein in
Burma. Possible syntypes: BMNH 1870.6.14.34 (1), NMW 44358 (1), NMW 48327 (1), ZMB 2735 (1), ZSI
F.454 (1), ZSI F.500 (1), ZSI A.569 (13).
Distribution: Salween River, Myanmar (Jayaram, 1982).
Arius africana Günther, 1867
Arius falcarius africana Günther, in Playfair & Günther, 1867: 114. Type locality: Pangani River, East Africa. Syntypes: BMNH 1865.9.21.16 (1), BMNH 1867.3.7.385–386 (2, skins), BMNH 1867.3.7.499 (1). Originally as
Arius falcarius var. africana.
Distribution: Pangani River and estuaries in Madagascar (Taylor, 1986a).
Arius brunellii Zolezzi, 1939
Arius brunellii Zolezzi, 1939: 168, fig. 1. Type locality: Giuba, Somalia. Syntypes (4): Whereabouts unknown.
Distribution: Juba River, Somalia (Taylor, 1986).
Remarks: Known only from the original description.
Arius gigas Boulenger, 1911
Arius gigas Boulenger, 1911a: 386, fig. 298. Type locality: Lokoja, Upper Niger. Holotype: BMNH 1904.1.20.41.
Distribution: Volta, Niger, and other rivers of Western Africa (Taylor, 1986).
Hemiarius Harmandi Sauvage, 1880
Hemiarius Harmandi Sauvage, 1880b: 230. Isle de Phu-Quoc (golfe de Siam). Holotype: MNHN a-2390.
? Arius brevirostris Steindachner, 1901: 447. Baram-Flusse, Borneo. Holotype: at NMW.
Distribution: Gulf of Thailand and, possibly, Borneo.
Remarks: Redescribed in Kailola (1999) but not assigned to any valid genus in Kailola (2004).
Arius Heudelotii Valenciennes, 1840
Arius Heudelotii Valenciennes, in Cuvier & Valenciennes, 1840b: 73 (54 of Strasbourg deluxe edition), pl. 428. Type
locality: haut Sénégal. Syntypes: MNHN b-0689 (2).
Arius mercatoris Poll, 1949: 201, fig. 8. Type locality: Atlantic, 12º34'N, 17º09'W, 8–10 fm. Holotype: IRSNB 89.
Distribution: West coast of Africa, from Mauritania to Gabon and, possibly, to Angola; in marine waters and estuaries (Taylor, 1986).
Arius latiscutatus Günther, 1864
Arius latiscutatus Günther, 1864: 151, unnumbered figure. Type locality: Fernando Po and West Africa [restricted toFernando Po by Lectotype designation]. Lectotype: BMNH 1864.2.26.1 (skin), designated by Taylor (1990:
231).
Arius lagoensis Pfaff, 1933: 298, figs. 9–10. Type locality: Lagos [Nigeria]. Holotype: ZMUC P 29625.
Distribution: Along the western coast of Africa, from Dakar to Angola, and Bioko; sometimes in freshwaters (Tay-CHECKLIST OF CATFISHES, RECENT AND FOSSIL

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lor, 1986).
Hemipimelodus macrorhynchus Weber, 1913
Hemipimelodus macrorhynchus Weber, 1913: 549, fig. 27. Type locality: Lorentz-Fluss, van Weelskamp; LorentzFluss, “Bivak” Insel; Lorentz-Fluss, Alkmaar [New Guinea]. Syntypes: ZMA 111085 (2), ZMA 111086 (1);dorsal view of head illustrated in Desoutter (1977: 14, fig. 2).
Hemipimelodus aaldereni Hardenberg, 1936b: 367. Type locality: Middle course of Digulriver, southern New
Guinea. Holotype: ZMA 110782.
Distribution: Southern Papua (formerly Irian Jaya), New Guinea (Allen, 1996).
Remarks: Treated as valid in Hemipimelodus by Desoutter (1977), but Kailola (2004) considered the species to be
more closely related to species of Plicofollis and/or Genidens.
Arius parkii Günther, 1864
Arius parkii Günther, 1864: 154, unnumbered figure. Type locality: Lagos and River Niger [restricted to Lagos, bylectotype designation] Nigeria. Lectotype: BMNH 1865.7.4.4, designated by Taylor (1986a: 156).
Arius Capellonis Steindachner, 1867b: 64. Type locality: Monrovia in West-Afrika. Holotype: at NMW. [Description translated into English in Steindachner (1867g: 442).] Illustrated and described in more detail in Steindachner (1867e: 523, pl. 2, figs. 1–3).
Arius granulatus Peters, 1868a: 454. Type locality: Lagos, Nigeria. Holotype: ZMB 6780.
Distribution: Western Africa, from Angola to Cape Blanco and occasionally further north; in coastal waters andestuaries, sometimes in river mouths (Taylor, 1986).
Arius phrygiatus Valenciennes, 1840
Arius phrygiatus Valenciennes, in Cuvier & Valenciennes, 1840b: 79 (59 of Strasbourg deluxe edition). Type locality: Cayenne [interpreted by Boeseman (1972) as having come from Suriname]. Holotype: RMNH 3038.
Arius Dieperinki Bleeker, 1862a: 375. Type locality: Surinama. Holotype: RMNH 3038; holotype illustrated in
Bleeker (1864a: 51, pl. 10, pl. 12, fig. 3).
Distribution: Atlantic coastal rivers and estuaries of South America from Guyana to the mouth of the Amazon River(Marceniuk & Ferraris, 2003).
Remarks: Boeseman (1972) treated RMNH 3038 as the holotype of A. phrygiatus.
Arius rugispinis Valenciennes, 1840b
Arius rugispinis Valenciennes, in Cuvier & Valenciennes, 1840b: 77 (58 of Strasbourg deluxe edition). Type locality:
Cayenne. Syntypes (3): MNHN a-9403 (1).
Tachysurus atroplumbeus Fowler, 1931: 394, fig. 2. Type locality: Vessigny Beach, Brighton, Trinidad. Holotype:
ANSP 53316.
Distribution: Atlantic coastal rivers of South America from Venezuela to mouth of Amazon River (Marceniuk &Ferraris, 2003).
† Felichthys stauroforus Lynn & Melland, 1939
† Felichthys stauroforus Lynn & Melland, 1939: 14. Type locality: Calvert Formation, Maryland. Holotype: USNM
15476 (partial neurocranium).
Distribution: Calvert Formation, Zone 12, Miocene (Lundberg, 1975, as † Arius stauroforus).
AMISSIDENS Kailola, 2004
Amissidens Kailola, 2004: 125. Type species: Arius hainesi Kailola, 2000. Type by original designation. Gender:
Masculine.
Amissidens hainesi (Kailola, 2000)
Arius hainesi Kailola, 2000: 139, figs. 9–11. Type locality: Ludmilla Creek (12º27'S, 130º46'E), Darwin, Northern
Territory, Australia. Holotype: NTM S.11507-001.
Distribution: Southern coast of New Guinea and Northern Australia, between Darwin and southern Gulf of Carpentaria (Kailola, 2000).
ARIOPSIS Gill, 1861
Ariopsis Gill, 1861a: 56. Type species: Arius milberti Valenciennes, 1840. Type by monotypy. Gender: Feminine.

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Remarks: Kailola (2004) treated this genus as valid with two distinct geographic components, one distributed intropical America and the the other in the Australia-New Guinea region. Herein, the two geographic components
are treated as separate genera; the American species placed in Ariopsis and the Australia-New Guinean species
in Neoarius.
Ariopsis assimilis (Günther, 1864)
Arius assimilis Günther, 1864: 146. Type locality: Lake of Yzabal, Guatemala. Holotype: BMNH 1864.1.26.98.
Distribution: Atlantic draining rivers, from Quintana Roo, Mexico, to Honduras (Miller et al., 2005).
Ariopsis bonillai (Miles, 1945)
Galeichthys bonillai Miles, 1945: 454, figs. 4–4a. Type locality: Río Magdalena, Honda, Colombia. Holotype: atSec. Caza y Pesca, Min. Nat. Econ. Bogotá.
Distribution: Caribbean draining rivers of Colombia, including Magdalena River (Marceniuk & Ferraris, 2003).
Ariopsis felis (Linnaeus, 1766)
Silurus Felis Linnaeus, 1766: 503. Type locality: Carolina [now: Charleston Harbor, South Carolina, U.S.A. by neotype designation]. Neotype: BMNH 1985.11.11.1. Name placed on Official List in ICZN Opinion 1547, withneotype designated.
Arius Milberti Valenciennes, in Cuvier & Valenciennes, 1840b: 74 (56 of Strasbourg deluxe edition). Type locality:
New-York ... Charlestown. Syntypes: MNHN b-0593 (3).
Arius equestris Baird & Girard, 1854: 26. Type locality: Indianola, Texas. Holotype: USNM 836; holotype illustrated in Girard (1859a: 32, pl. 15).
Galeichthys guentheri Regan, 1907, in Regan, 1906–08: 124, pl. 16 (fig. 1), 19 (fig. 3). Type locality: Gulf of Mexico. Syntypes: BMNH 1855.9.19.1105–1106 (2).
Distribution: Western North Atlantic and Gulf of Mexico.
Ariopsis guatemalensis (Günther, 1864)
Arius guatemalensis Günther, 1864: 145, unnumbered figures. Type locality: Guatemala [and] Chiapan. Syntypes(2): BMNH 1853.1.11.6 (1).
Arius cærulescens Günther, 1864: 149, fig. on p. 149. Type locality: Huamuchal, Guatemala. Syntypes: BMNH
1864.1.26.208–209 (2).
Galeichthys azureus Jordan & Williams, in Jordan, 1895: 398, pl. 27. Type locality: Estuary at Mazatlán, Sinaloa, w.
Mexico. Holotype: SU 11575.
Distribution: Pacific coast and coastal rivers of North and Central America, from Gulf of California, Mexico to Honduras (Miller et al., 2005).
Ariopsis lentiginosa (Eigenmann & Eigenmann, 1888)
Tachisurus lentiginosus Eigenmann & Eigenmann, 1888b: 139. Type locality: Panama. Syntypes (2, 39 cm): MCZ
4970 (2).
Galeichthys xenauchen Gilbert, in Jordan & Evermann, 1898: 2777. Type locality: Panama. Holotype: SU 5821;
holotype figured in Gilbert & Starks (1904: pl. 4, fig. 7).
Distribution: Pacific coast in Panama (Kailola & Bussing, 1995).
Ariopsis seemanni (Günther, 1864)
Arius seemanni Günther, 1864: 147, fig. Type locality: Central America. Holotype: BMNH 1855.9.19.1107.
Tachisurus jordani Eigenmann & Eigenmann, 1888b: 142. Type locality: Panama. Syntypes (2, 25-26 cm): MCZ
4945 (2).
Galeichthys gilberti Jordan & Williams in Jordan, 1895: 395, pl. 26. Type locality: Upper part of astillero at Maz
atlán, Sinaloa, w. Mexico. Holotype: USNM 29213.
Galeichthys eigenmanni Gilbert & Starks, 1904: 21, pl. 4 (fig. 8). Type locality: Panama. Holotype: SU 6986.
Galeichthys simonsi Starks, 1906: 764, fig. 1. Type locality: Callao, Peru. Holotype: USNM 53466.
Distribution: Pacific draining rivers and estuaries of the Americas, from Mexico to Peru (Miller et al., 2005).
Species inquirenda, Ariopsis
Hexanematichthys surinamensis Bleeker, 1862a: 380. Type locality: Surinama. Holotype: RMNH 3055; holotype illus-

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trated in Bleeker (1864: 55, pl. 6, fig. 2; pl. 7, fig. 1).
Remarks: Treated as valid in Kailola (2004), but of uncertain status in Marceniuk & Ferraris (2003).
ARIUS Valenciennes, 1840
Arius Valenciennes, in Cuvier & Valenciennes, 1840b: 53 (40 of Strasbourg deluxe edition). Type species: Pimelodus arius Hamilton, 1822. Type by absolute tautonymy. Gender: Masculine.
Ariodes Müller & Troschel, 1849: 9. Type species: Bagrus (Ariodes) arenarius Müller & Troschel, 1849. Type by
subsequent designation, by Bleeker (1862–63: 8). Gender: Masculine. Originally proposed as a subgenus of
Bagrus.
Pseudarius Bleeker, 1862 (in Bleeker, 1862–63): 8. Type species: Pimelodus arius Hamilton, 1822. Type by original
designation. Gender: Masculine.
Arius arenarius (Müller & Troschel, 1849)
Bagrus (Ariodes) arenarius Müller & Troschel, 1849: 9. Type locality: China. Holotype or Syntype: ZMB 3001.
Arius fangi Chaux, in Chaux & Fang, 1949a: 194, fig. 1. Type locality: Tonkin. Holotype: MNHN 1966-0726.
Distribution: Southern China, in coastal waters.
Remarks: The synonymy here follows the proposal in Kottelat (2001a: 56).
Arius arius (Hamilton, 1822)
Pimelodus arius Hamilton, 1822: 170, 376. Type locality: estuaries of Bengal. No types known.
Arius falcarius Richardson, 1845: 134, pl. 62 (figs. 7–9). Type locality: Canton, China. Syntypes (2): BMNH (apparently lost: Whitehead, 1970: 210). BMNH has two specimens identified as Arius falcarius from the Reeves collection. One specimen preserved in alcohol, 129 mm SL (BMNH 2005.9.12.2) and the second, a mounted, left
one-half specimen, 289 mm SL (BMNH 2005.9.12.1). One or both are likely to be part of the type series.
Arius cochinchinensis Günther, 1864: 170, unnumbered figures. Type locality: Cochinchina. Holotype (skin): at
BMNH, but not found in 2005.
Arius boakeii Turner, 1867: 82. Type locality: Ceylon. Syntypes: BMNH 1866.7.11.1 (1), possibly NMSZ 1987.030(1).
Arius Buchanani Day, 1877 (in Day, 1875–78): 463, pl. 105 (fig. 6). Type locality: Hooghly at Calcutta and Burma[Irrawaddi]. Possible syntypes: ZSI 456 (1), ZSI 1302 (1, apparently lost).
Distribution: Bay of Bengal, South China Sea and, possibly, coast of Sumatra (Kailola, 1999); Pakistan, west coastof India, and Chilka Lake (Jayaram, 1982).
Remarks: See Wang et al. (2005a) for comments on the validity of this species.
? Arius bicolor (Fowler, 1935)
Hemipimelodus bicolor Fowler, 1935b: 100, fig. 23. Type locality: Bangkok, Siam. Holotype: ANSP 60777.
Hemipimelodus atripinnis Fowler, 1937: 145, fig. 34. Type locality: Bangkok, Siam. Holotype: ANSP 67906.
Distribution: Lower courses of Mekong and Chao Phraya River basins (Rainboth, 1996).
Remarks: Treated as valid in Hemipimelodus by Desoutter (1977) but as a likely synonym of Arius maculatus by
Kailola (1999).
Arius dispar Herre, 1926
Arius dispar Herre, 1926: 405, pl. 1 (fig. 6). Type locality: Paco market, Manila, Philippines. Holotype: BSMP
(destroyed).
Distribution: Philippines, Taiwan and, possibly, northern Borneo (Kailola, 1999).
Arius festinus Ng & Sparks, 2003
Arius festinus Ng & Sparks, 2003: 5, fig. 4. Type locality: Madagascar, Amboaboa River, near its confluence with
the Mangarahara River (left bank tributary of the Sofia River), 15º50'01"S, 48º42'52"E. Holotype: UMMZ
239806.
Distribution: Amboaboa River, Madagascar (Ng & Sparks, 2003).
Arius gagora (Hamilton, 1822)
Pimelodus gagora Hamilton, 1822: 167, 376, pl. 10 (fig. 54). Type locality: estuaries of Bengal. No types known.
Distribution: Bay of Bengal, from India to Thailand; in tidal portions of rivers and along the coast (Kailola, 1999).
Arius leptonotacanthus Bleeker, 1849

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Arius leptonotacanthus Bleeker, 1849: 11. Type locality: In Freto Madurae prope Kammal et Surabaya. Holotype(221 mm TL): BMNH 1863.12.4.114. Holotype illustrated in Bleeker (1862–63: 32, pl. 54 [=Silur. pl. 6], fig. 2).
Distribution: Western Central Pacific, along Gulf of Thailand and Indonesia; in coastal waters and estuaries (Kailola,1999).
Remarks: Register number of holotype of Arius leptonotacanthus listed as BMNH 1863.12.4.144 in Eschmeyer etal. (1998).
Arius maculatus (Thunberg, 1792)
Silurus maculatus Thunberg, 1792: 31, pl. 1 (fig. 2). Type locality: China, Japan. No types known.
Silurus ocellatus Bloch & Schneider, 1801: 379. Type locality: Japonia. No types known. Unneeded new name for
Silurus maculatus Thunberg, 1792.
Pimelodus thunberg La Cepède, 1803: 691, 692. Type locality: China, Japan. No types known. Unneeded new namefor Silurus maculatus Thunberg, 1792.
Bagrus gagorides Valenciennes, in Cuvier & Valenciennes, 1840a: 441 (327 of Strasbourg deluxe edition). Type
locality: [Ganges River]. Holotype: MNHN a-9330 (mounted).
Arius gagoroides Bleeker, 1846a: 168. Type locality: Batavia. Syntypes (size and number not stated): SMNS 10567(2). [See also Bleeker, 1847c.]
Distribution: Western central Pacific; in inshore coastal waters, estuaries, and lower reaches of rivers (Kailola,1999).
Arius madagascariensis Vaillant, 1894
Arius madagascariensis Vaillant, 1894a: 77. Type locality: Fl. Moroundava [Madagascar]. Holotype: MNHN 18940002.
Distribution: Western side of Madagascar, in rivers and lakes (Taylor, 1986b; Ng & Sparks, 2003).
Remarks: Redescribed in Ng & Sparks (2003).
Arius malabaricus Day, 1877
Arius Malabaricus Day, 1877 (in Day, 1875–78): 464, pl. 107 (fig. 4). Type locality: Canara [India]. Holotype: ZSI
1305.
Distribution: Peninsular India.
Remarks: See Jayaram & Dhanze (1981a) for comments on validity.
Arius manillensis Valenciennes, 1840
Arius Manillensis Valenciennes, in Cuvier & Valenciennes, 1840b: 93 (69 of Strasbourg deluxe edition). Type locality: Manille. Holotype (14 pouces): at MNHN.
Pseudarius philippinus Sauvage, 1880a: 226. Type locality: Ile de Luçon, Philippines. Holotype: MNHN a-2615.
Distribution: Philippines, in coastal waters, estuaries, and freshwater (Kailola, 1999).
Arius microcephalus Bleeker, 1855
Arius microcephalus Bleeker, 1855b: 423. Type locality: Bandjermasin, in fluviis [Borneo]. Syntypes (2, 140– 213
mm TL): BMNH 1863.12.4.149 (1, 108 mm SL, 132 mm TL), RMNH 6902 (1). One syntype illustrated in
Bleeker (1862–63:37, pl. 51 [Silur. pl. 3], fig. 1, as Pseudarius microcephalus).
? Arius sciurus Smith, 1931: 30. Type locality: Tapi River near Bandon, Peninsular Siam. Holotype: USNM 90310.
Distribution: Borneo and eastern side of Malay peninsula; in lower portions of rivers and in brackish waters(Kailola, 1999).
Remarks: Register number of syntype of Arius microcephalus incorrectly reported as BMNH 1863.12.11.149 in
Eschmeyer et al. (1998).
Arius oetik Bleeker, 1846
Arius oetik Bleeker, 1846a: 166. Type locality: Batavia. Syntype (size and number not stated): Possibly BMNH
1863.12.4.111 (1), NMV 45987 (1). Bleeker (1847: 32) changed the spelling to Arius utik, which has been usedfrequently in literature. [See also Bleeker, 1847c.]
Distribution: Gulf of Thailand and in waters around much of Malaysia and Indonesia; in coastal waters and estuaries(Kailola, 1999).
Arius subrostratus Valenciennes, in Cuvier & Valenciennes, 1840b
Arius subrostratus Valenciennes, in Cuvier & Valenciennes, 1840b: 62 (46 of Strasbourg deluxe edition). Type local-

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ity: Malabar. Holotype: MNHN 0000-1190.
Distribution: South Asia and eastward to western central (Jayaram, 1982).
Arius sumatranus (Bennett, 1830)
Bagrus sumatranus Bennett, 1830: 691. Type locality: Sumatra. Syntype: BMNH 1855.12.26.485 (1).
Distribution: Eastern Bay of Bengal to western central Pacific; in coastal waters and estuaries (Kailola, 1999).
Remarks: Treated by Kailola (1999) as a likely synonym of Arius venosus Valenciennes, 1840, but later treated asvalid (Kailola, 2004). The distribution limits stated here represent the entire distribution of A. venosus, which
may not be that of A. sumatranus.
Arius uncinatus Ng & Sparks, 2003
Arius uncinatus Ng & Sparks, 2003: 12, fig. 7. Type locality: Madagascar, Lake Andrapongy, near Antsohihy. Holotype: MHNG 2622.81
Distribution: Lake Andrapongy, Ankofia River basin, Madagascar (Ng & Sparks, 2003).
Arius venosus Valenciennes, 1840
Arius venosus Valenciennes, in Cuvier & Valenciennes, 1840b: 69 (52 of Strasbourg deluxe edition). Type locality:
Rangoon; Manille. Syntypes: MNHN 0000-1205 (2).
Bagrus Schlegelii Bleeker, 1846a: 153. Type locality: Batavia.Type(s) (size and number not stated): possibly AMS
B.8123. [See also Bleeker, 1847c.]? Arius micruropterygius Bleeker, 1847a: 165. Type locality: Batavia. Type(s): Whereabouts unknown.
Distribution: Eastern Bay of Bengal to western central Pacific; in coastal waters and estuaries (Kailola, 1999).
Species inquirendae, Arius
Arius macracanthus Günther, 1864: 167, unnumbered figure. Type locality: Siam. Holotype: BMNH 1862.11.1.222.
Arius jella Day, 1877 (in Day, 1875–78): 467, pl. 106 (fig. 3). Type locality: Coromandel Coast of India; Madras. Syntype: ZSI 1304 (1, figured specimen).
ASPISTOR Jordan & Evermann, 1898
Aspistor Jordan & Evermann, 1898: 2763. Type species: Arius luniscutis Valenciennes, 1840. Type by original designation. Gender: Masculine.
Aspistor hardenbergi (Kailola, 2000)
Arius hardenbergi Kailola, 2000: 137, figs. 7–8. Type locality: Manimeri River (2º06'S, 133º45'E), Bintuni Bay,
Papua (Irian Jaya). Holotype: WAM P.29966-001.
Distribution: Southern coast of New Guinea, from Vogelkop Peninsula to mouth of Fly River (Kailola, 2000).
Aspistor luniscutis (Valenciennes, 1840)
Arius luniscutis Valenciennes in Cuvier & Valenciennes, 1840b: 109 (82 of Strasbourg deluxe edition). Type locality: Brésil. Syntypes: MNHN a-8980 (2), MNHN b-0595 (1).
Arius physacanthus Vaillant, 1899: 155. Type locality: le Mahury [French Guiana]. Holotype: MNHN 1899-0058;
holotype illustrated in Vaillant (1900: 128, fig. 1).
Arius bonneti Puyo, 1936: 107, fig. 17 (middle). Type locality: not stated [Cayenne River, French Guiana]. Holotype: MNHN 1936-0162. Preoccupied by † Arius bonneti Priem, 1904.
Arius clavispinosus Puyo, 1936: 103, fig. 16 (left). Type locality: Tle de Cayenne (rivière de Cayenne) [French Guiana]. Holotype: MNHN 1936-0163.
Arius despaxi Puyo, 1936: 110, fig. 17 (right). Type locality: [Cayenne River, French Guiana]. Holotype: MNHN
1936-0161.
Distribution: Atlantic coastal rivers of South America from French Guiana to southern Brazil (Marceniuk & Ferraris, 2003).
Aspistor parkeri (Traill, 1832)
Silurus parkeri Traill, 1832: 377, pl. 6 (fig. 1). Type locality: Guiana. Holotype: Whereabouts unknown.
Distribution: Atlantic draining coastal rivers of South America from Guyana to northern Brazil (Marceniuk & Ferraris, 2003).

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Aspistor quadriscutis (Valenciennes, 1840)
Arius quadriscutis Valenciennes, in Cuvier & Valenciennes, 1840b: 111 (83 of Strasbourg deluxe edition). Type
locality: Cayenne ou de la Mana ... Cayenne. Syntypes: MNHN a-9402 (1), MNHN b-0596 (1), MNHN b-0613(1), RMNH 3049 (1), RMNH 3050 (1).
Distribution: Atlantic coastal rivers of South America from Guyana to northeastern Brazil (Marceniuk & Ferraris,2003).
BAGRE Cloquet, 1816
Bagre Cloquet, 1816: 52. Type species: Silurus bagre Linnaeus, 1766. Type by absolute tautonymy. Gender: Masculine. Placed on Official List (ICZN, Opinion 1402).
Bagre Oken, 1817: 1183. Silurus bagre Linnaeus, 1766. Type by absolute tautonymy. Gender: Masculine. Apparently proposed independently of Bagre Cloquet.
Glanis Spix & Agassiz, 1829: 46. Type species: Silurus bagre Linnaeus, 1766. Type by subsequent designation by
Kottelat (1988: 78). Gender: Masculine. Not preoccupied by Glanis in Rafinesque, 1818, which is not an available name.
Stearopterus Minding, 1832: 116. Type species: Stearopterus bagre Minding, 1832 [= Silurus bagre Linnaeus,
1766]. Type by monotypy. Gender: Masculine.
Breviceps Swainson, 1838: 328, 343. Type species: Silurus bagre of Bloch, 1794. Type by monotypy. Gender: Neuter. Preoccupied by Breviceps Merrem, 1820 (in Amphibia); replaced by Felichthys Swainson, 1839.
Felichthys Swainson, 1839: 305. Type species: Silurus bagre of Bloch, 1794. Type by being a replacement name.
Gender: Masculine. Replacement for Breviceps Swainson, 1838.
Ailurichthys Baird & Girard, 1854: 26. Type species: Silurus marinus Mitchill, 1815. Type by subsequent designation by Jordan & Evermann (1896: 116). Gender: Masculine. Spelled Aelurichthys in Gill (1863: 172) and elsewhere, but clearly attributed to Baird & Girard, so treated here as an unintended misspelling and not anavailable name.
Mystus Gray, 1854: 155. Type species: Mystus carolinensis Gronow, 1854. Type by subsequent designation by Jordan & Evermann (1896: 116). Gender: Masculine. Preoccupied by Mystus Scopoli, 1777.
Paradiplomystes Bleeker, 1862 (in Bleeker, 1862–63): 8. Type species: Pimelodes coruscans Lichtenstein, 1819.
Type by original designation. Gender: Masculine.
Anemanotus Fowler, 1944c: 171. Type species: Aelurichthys panamensis Gill, 1863. Type by original designation.
Gender: Masculine. Originally proposed as a subgenus of Ailurichthys.
Taxonomic overview: Acero et al. (2005).
Remarks: See Gill (1891b) for comments on the validity of Felichthys.
Bagre bagre (Linnaeus, 1766)
Silurus Bagre Linnaeus, 1766: 505. Type locality: America meridionali. On several literature accounts, including
Gronovius (1763: no. 382). Placed on Official List as type species of Bagre (ICZN, Opinion 1402).
? Pimelodes coruscans Lichtenstein, 1819: 58. Type locality: Brasilien. Holotype: Whereabouts unknown; apparently not ZMB 2978 as listed by Paepke (1999).
Felichthys filamentosus Swainson, 1839: 392. Type locality: Estuaries of rivers near Pernambuco, Brazil. No types
known.
Galeichthys Gronovii Valenciennes, in Cuvier & Valenciennes, 1840b: 40 (30 of Strasbourg deluxe edition).
Unneeded new name for Silurus bagre Linnaeus.
Bagrus macronemus Ranzani, 1842: 334, pl. 28. Type locality: Fiume Brasiliano detto di S. Francesco. Holotype:
MZUB 930. Name appears first in Ranzani (1841: 65) as a nomen nudum.
Mystus carolinensis Gronow in Gray, 1854: 156. Type locality: fluminibus majoribus Americes Meridionalis. No
types known; based on several literature sources.
Distribution: Caribbean and Atlantic coastal rivers of South America, from Colombia to the mouth of the Amazon
River (Marceniuk & Ferraris, 2003).
Bagre marinus (Mitchill, 1815)
Silurus marinus Mitchill, 1815: 433. Type locality: New York. No types known.

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Galeichthys Blochii Valenciennes, in Cuvier & Valenciennes, 1840b: 44 (33 of Strasbourg deluxe edition). Type
locality: Surinam; Bahia. Syntypes: at MNHN (2), and ZMB 2989 (Paepke, 1999: 46). Description based on
specimens and reference to Bloch (1794: pl. 365).
Galeichthys Parrae Valenciennes, in Cuvier & Valenciennes, 1840b: 33 (25 of Strasbourg deluxe edition). Type
locality: New York. No types known. Unneeded new name for Silurus marinus Mitchill.
Galeichthys bahiensis Castelnau, 1855: 37, pl. 18 (fig. 1). Type locality: Bahia [Bahia]. Holotype: MNHN b-0692.
lurichthys longispinis Günther, 1864: 178. Type locality: South America; Mexico. Syntypes (3): BMNH
1976.2.18.1 (1), BMNH 1976.2.18.2 (1).
Distribution: Coast of Gulf of Mexico, western margin of Caribbean and northern margin of South America: sometimes in rivers and estuaries (Marceniuk & Ferraris, 2003).
Bagre panamensis (Gill, 1863)
Aelurichthys panamensis Gill, 1863: 172. Type locality: Western coast of Central America. Holotype: at USNM (notfound, see Jordan & Gilbert, 1882; Ferraris & Vari, 1992).
lurichthys nuchalis Günther, 1864: 179, fig. Type locality: Pacific coast of Panama. Holotype: BMNH
1864.1.26.344; holotype illustrated in Günther (1868c: pl. 81, fig. 2).
Aelurichthys scutatus Regan, 1907, in Regan, 1906–08: 116, pl. 15 (fig. 2), 19 (fig. 2). Type locality: Panama or
Ecuador. Syntype: BMNH 1903.5.15.328.
Aelurichthys isthmensis Regan, 1907, in Regan, 1906–08: 117, pl. 15 (fig. 1), 19 (fig. 1). Type locality: Colón, Panama. Syntypes: BMNH 1877.5.31.7–8
Distribution: West coast of Americas, from off Santa Ana River in southern California, USA, to Peru. Only rarely
found north of southern Baja California (Marceniuk & Ferraris, 2003).
Bagre pinnimaculatus (Steindachner, 1876)? Galeichthys Eydouxii Valenciennes, in Cuvier & Valenciennes, 1840b: 43 (32 of Strasbourg deluxe edition). Type
locality: la rivière de Guayaquil. Holotype: MNHN 0000-1572. Name spelled Eidouxii on p. 43 and Eydouxii intable of contents (p. xviii). First reviser apparently Eschmeyer, Ferraris, Hoang & Long (1998).
Aelurichthys pinnimaculatus Steindachner, 1876: 565, pl. 8. Type locality: Panama, Altata, Westküste von CostaRica. Syntypes: MCZ 4941 (1), MCZ 23746 (3), MCZ 23747 (1), NMW 47895 (1), NMW 47897 (1), NMW
47898 (1), NMW 47899 (1), NMW 50577 (1), USNM 123011 (2).
Distribution: Pacific draining rivers of North, Central and South America, from the Gulf of California to Ecuador(Marceniuk & Ferraris, 2003).
Remarks: Species treated as valid in Kailola & Bussing (1995: 878), but with Galeichthys eydouxii as a possible
senior synonym. Therefore, the name G. eydouxii is tentatively placed here.

E

E

BATRACHOCEPHALUS Bleeker, 1846
Batrachocephalus Bleeker, 1846a:176. Type species: Batrachocephalus ageneiosus Bleeker, 1846. Type by monotypy. Gender: Masculine.
Batrachocephalus mino (Hamilton, 1822)
Ageneiosus mino Hamilton, 1822: 159, 375. Type locality: Upper part of [Ganges] estuaries. No types known.
Batrachocephalus ageneiosus Bleeker, 1846a: 176. Type locality: Batavia. Type(s) (size and number not stated):
Whereabouts unknown [See also Bleeker, 1847c.]
Batrachocephalus micropogon Bleeker, 1858b: 118. Type locality: Java (Batavia, Surabaja), in mari et aquis fluviomarinis. Syntypes (10: 180–248 mm TL): AMS B.7964, BMNH 1862.2.4.18, also on Ageneiosus mino Hamilton and Batrachocephalus ageneiosus Bleeker). One syntype illustrated in Bleeker (1862–63: pl. 17).
Distribution: Bay of Bengal, and parts of the western central Pacific, in coastal waters, estuaries, and lower reachesof rivers (Kailola, 1999); also Pakistan (Jayaram, 1982).
BRUSTIARIUS Herre, 1935
Brustiarius Herre, 1935: 388. Type species: Arius nox Herre, 1935. Type by original designation. Gender: Masculine. Originally proposed as a subgenus of Arius.
Brustiarius nox (Herre, 1935)

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Arius nox Herre, 1935: 388. Type locality: Nyaurangai, Sepik River, New Guinea. Holotype: FMNH 17195.
Distribution: Sepik River, New Guinea (Herre, 1935).
Brustiarius solidus (Herre, 1935)
Arius solidus Herre, 1935: 385. Type locality: Timbunke, Sepik River, New Guinea. Holotype: FMNH 17201.
Arius kanganamanensis Herre, 1935: 387, fig. 32. Type locality: Kanganaman, Sepik River, New Guinea. Holotype:
FMNH 17194.
Hemipimelodus bernhardi Nichols, 1940: 3. Type locality: Bernhard Camp, Idenburg River, Netherland New
Guinea. Holotype: AMNH 15039.
Arius microstomus Nichols, 1940: 2. Type locality: Bernhard Camp, Idenburg River, Netherland New Guinea. Holotype: AMNH 15041.
Distribution: Northern Papua (formerly Irian Jaya) (Allen, 1996) and Sepik River basin.
Remarks: Kailola (1990) and Allen & Coates (1990) treat Arius kangamanensis as a junior synonym of A. solidus.
As such, the earlier of these two publications serves as first reviser in establishing priority but, since the results
are identical, the question of which of these papers was published first was not pursued at this time.
CATHOROPS Jordan & Gilbert, 1882
Cathorops Jordan & Gilbert, 1882: 39, 54. Type species: Arius hypophthalmus Steindachner, 1876. Type by monotypy. Gender: Masculine. Originally proposed as a subgenus of Arius. Name dates to key on p. 39, which is partof a signature dated as 25 Sept 1882.
Cathorops agassizii (Eigenmann & Eigenmann, 1888)
Tachisurus agassizii Eigenmann & Eigenmann, 1888b: 145. Type locality: Rio Grande do Sul [Brazil]. Holotype:
MCZ 7670.
Arius pleurops Boulenger, 1897a: 296. Type locality: Magoarisinho, Cape Magoary [Marajo I., Brazil]. Holotype:
BMNH 1897.7.17.7 (plus eggs and embryos in mouth).
Distribution: Northern coast of South America including Guyana and the north and northeast Brazil (Marceniuk &Ferraris, 2003).
Cathorops aguadulce (Meek, 1904)
Galeichthys aguadulce Meek, 1904: 9, pl. 4. Type locality: Río Tesechocán, Pérez, Vera Cruz basin of the Rio Papaloapam [Mexico]. Holotype: FMNH 4678.
Distribution: Atlantic draining rivers from Panuco River basin, Mexico to Izabal Lake, Guatemala (Miller et al.,2005).
Cathorops arenatus (Valenciennes, 1840)
Arius arenatus Valenciennes in Cuvier & Valenciennes, 1840b: 106 (79 of Strasbourg deluxe edition). Type locality:
Cayenne [interpreted by Boeseman (1972) as having come from Suriname]. Holotype: RMNH 3099.
Arius fissus Valenciennes in Cuvier & Valenciennes, 1840b: 107 (79 of Strasbourg deluxe edition). Type locality:
Cayenne [interpreted by Boeseman (1972) as having come from Suriname]. Holotype: RMNH 3036.
Distribution: Atlantic coastal rivers of South America from Guyana to northeastern Brazil (Marceniuk & Ferraris,2003).
Remarks: Marceniuk & Ferraris (2003: 449) acted as first reviser in giving precedence to Arius arenatus over A. fissus.
Cathorops dasycephalus (Günther, 1864)
Arius dasycephalus Günther, 1864: 157, fig. on p. 157. Type locality: Oahu, Sandwich Islands [in Error]. Holotype:
BMNH 1855.9.19.1100.
Tachisurus longicephalus Eigenmann & Eigenmann, 1888b: 143. Type locality: Gulf of Panama, Panama. Holotype:
MCZ 4972. Name spelled Tachisurus longicehpalus in account heading, T. longicephalus in introduction (p.
119); the latter spelling is considered valid.
Distribution: Pacific coast of South and Central America, from Costa Rica to Ecuador (Marceniuk & Ferraris, 2003).
Cathorops fuerthii (Steindachner, 1876)
Arius Fürthii Steindachner, 1876: 579. Type locality: Panama. Syntypes (4): possibly MCZ 4943 (2), MCZ 4973 (3),MCZ 7691 (1), NMW 50568 (2).

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Tachysurus evermanni Gilbert & Starks, 1904: 32, pl. 5 (fig. 10). Type locality: Panama Bay. Holotype: SU 6706.
Distribution: Pacific draining rivers of the Americas from Mexico to Ecuador (Miller et al., 2005).
Cathorops hypophthalmus (Steindachner, 1876)
Arius hypophthalmus Steindachner, 1876: 581, pl. 10. Type locality: Panama. Holotype: at NMW.
Tachisurus gulosus Eigenmann & Eigenmann, 1888b: 146. Type locality: Panama. Syntypes (2): MCZ 4974 (2).
Distribution: Pacific draining rivers of Panama, in brackish and fresh water (Marceniuk & Ferraris, 2003).
Cathorops mapale Betancur-R. & Acero P., 2005
Cathorops mapale Betancur-R. & Acero P., 2005: 47, figs. 1, 2. Type locality: Ciénaga Grande de Santa Marta(CGSM), Madgalena, CO (10º59'N, 74º17'W). Holotype: INVEMAR-PEC 5333.
Distribution: Coastal Caribbean region of Colombia (Betancur-R. & Acero P., 2005).
Cathorops melanopus (Günther, 1864)
Arius melanopus Günther, 1864: 172. Type locality: Rio Motagua. Syntypes (apparently only 2): BMNH
1865.4.29.51–53 (3).
Distribution: Motagua River basin of Guatemala (Marceniuk & Ferraris, 2003).
Cathorops multiradiatus (Günther, 1864)
Bagrus arioides Kner, 1863: 227, fig. 15. Type locality: Rio Bayano im Staate Panama, Südseite. Holotype (6"): at
NMW. Also in Kner & Steindachner (1864: 47). Preoccupied by Bagrus arioides Valenciennes, 1840; replaced
by Arius multiradiatus Günther, 1864.
Arius multiradiatus Günther, 1864: 173. Type locality: Rio Bayano im Staate Panama, Südseite. Holotype: at NMW.
Replacement for Bagrus arioides Kner, 1863.
Tachysurus emmelane Gilbert, in Jordan & Evermann, 1898: 2785. Type locality: Panama. Holotype: SU 5818;
holotype illustrated in Gilbert & Starks (1904: pl. 6).
Tachysurus equatorialis Starks, 1906: 766, fig. 3. Type locality: Guayaquil, Ecuador. Holotype: USNM 53470.
Distribution: Rivers and bays of Pacific coast of Central America, from Guatemala to Panama (Marceniuk & Ferraris, 2003).
Remarks: Kailola & Bussing (1995: 881) tentatively placed the species in Cathorops, which is followed herein.
Cathorops spixii (Spix & Agassiz, 1829)
Pimelodus spixii Spix & Agassiz, 1829: 19, pl. 7 (fig. 1). Type locality: Brasilia aequatoriali. Whereabouts unknown(Kottelat, 1988).
Pimelodus albidus Spix & Agassiz, 1829: pl. 7 (fig. 1). Type locality: [Not stated]. Whereabouts unknown (Kottelat,1988). Name available from plate caption.
Distribution: Atlantic and Caribbean rivers and estuaries of northern South America from Colombia to Brazil (Marceniuk & Ferraris, 2003), but considered restricted to the Atlantic coast by Betancur-R. & Acero P. (2005).
Remarks: Taylor & Menezes (1978) indicate that two or more species may be included here. Agassiz (1831, in Spix
& Agassiz, 1829–1831: conspectus) serves as first reviser in treating Pimelodus spixii as valid over Pimelodus
albidus.
Cathorops steindachneri (Gilbert & Starks, 1904)
Tachysurus steindachneri Gilbert & Starks, 1904: 29, pl. 5 (fig. 9). Type locality: Panama Bay. Holotype: SU 7026.
Arius taylori Hildebrand, 1925: 250, fig. 10. Type locality: Río Lempa, San Marcos, El Salvador. Holotype: USNM
87224.
Distribution: Pacific draining rivers and estuaries of Central America (Marceniuk & Ferraris, 2003).
Remarks: Kailola & Bussing (1995: 882) tentatively placed the species in Cathorops.
Cathorops tuyra (Meek & Hildebrand, 1923)
Arius tuyra Meek & Hildebrand, 1923: 128, pl. 5. Type locality: Río Tuyra, mouth of Río Yape, Darien, Panama.
Holotype: USNM 79413.
Distribution: Pacific drainage rivers and estuaries of South and Central America, from Panama to Peru (Marceniuk& Ferraris, 2003).
Remarks: Placed by Kailola & Bussing (1995: 883) tentatively in Cathorops.

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Species inquirendae, Cathorops
Arius variolosus Valenciennes in Cuvier & Valenciennes, 1840b: 107 (80 of Strasbourg deluxe edition). Type locality:
Cayenne. Holotype: MNHN 0000-4169.
Arius puncticulatus Valenciennes in Cuvier & Valenciennes, 1840b: 108 (81 of Strasbourg deluxe edition). Type locality:
Buénos-Ayres. Holotype (6 pouces): at MNHN.
Arius nuchalis Günther, 1864: 171, unnumbered figure. Type locality: British Guiana. Syntypes (6): BMNH
2005.5.17.1–2 (2), possibly BMNH 1975.1.17.3–5 (3), but original registration information gives locality as West
Indies and having come from “old collection” which does not agree with information provided in Günther.
Arius laticeps Günther, 1864: 171, unnumbered figures. Type locality: British Guiana; Trinidad. Syntypes: BMNH
1863.6.18.8 (1), BMNH 1976.2.25.1 (1, 137 mm TL).
Tachysurus liropus Bristol, in Gilbert, 1897: 438. Type locality: San Juan Lagoon, near mouth of Rio Ahome, Sonora,
Mexico. Syntypes (6): SU 324 (3), USNM 47584 (2).
CEPHALOCASSIS Bleeker, 1857
Cephalocassis Bleeker, 1857b: 473. Type species: Arius melanochir Bleeker, 1852. Type by monotypy. Also published in Bleeker (1858b: 62, 98). Gender: Feminine.
Hemipimelodus Bleeker, 1857b: 473. Type species: Pimelodus borneënsis Bleeker, 1851. Type by monotypy. Gender: Masculine.
Remarks: Genus revised by Desoutter (1977) as Hemipimelodus. Kailola (2004) apparently acted as first reviser ingiving precedence to Cephalocassis over Hemipimelodus, inasmuch as no previous synonymy of the twogeneric names has been found.
Key: Desoutter (1977, as Hemipimelodus).
Cephalocassis borneensis (Bleeker, 1851)
Pimelodus borneënsis Bleeker, 1851e: 430. Type locality: Sambas, in fluviis. Holotype (110 mm TL): RMNH 6906;
dorsal view of head illustrated in Desoutter (1977: 23, fig. 5).
Hemipimelodus macrocephalus Bleeker, 1858b: 239. Type locality: Borneo (Bandjarmasin), in fluviis. Syntypes (3:125–140 mm TL): BMNH 1863.12.4.116 (1, 109 mm SL, 124 mm TL), RMNH 27618 (1). One syntype illustrated in Bleeker (1862–63: pl. 82 [= Silur. pl. 34], fig. 2).
Hemipimelodus siamensis Sauvage, 1878: 234. Type locality: Laos. Holotype: MNHN 0000-9649.
Distribution: Mekong and Chao Phraya River basins, and Sundaland (Kottelat, 2001b).
Remarks: BMNH 1863.12.4.70 (143 mm ST, ~170 mm TL), listed as type of Pimelodus borneensis, is too large tobe the type.
Cephalocassis jatia (Hamilton, 1822)
Pimelodus jatius Hamilton, 1822: 171, 376. Type locality: estuaries of Bengal. No types known.
Distribution: Rivers and estuaries of South Asia, extending well above tidal range (Jayaram, 1982); Myanmar.
Cephalocassis manillensis (Valenciennes, 1840)
Pimelodus Manillensis Valenciennes, in Cuvier & Valenciennes, 1840b: 192 (142 in Strasbourg deluxe edition).
Type locality: Manille. Holotype: MNHN 0000-1209; dorsal view of head illustrated in Desoutter (1977: 16,
fig. 3).
Distribution: Philippines.
Remarks: Inclusion in Cephalocassis is tentative, based on the treatment in Desoutter (1977) as a valid species ofHemipimelodus, and the uncertain status of the species in Kailola (2004).
Cephalocassis melanochir (Bleeker, 1852)
Arius melanochir Bleeker, 1852d: 590. Type locality: Palembang, in fluviis. Holotype (302 mm TL): RMNH 6892.
? Arius doriae Vinciguerra, 1881: 174, unnumbered fig. Type locality: Sarawak state, Borneo, East Malaysia. Syntypes: MSNG 8135 (2), RMNH 10889 (1).
Distribution: Sumatra and Borneo.
Remarks: Synonymy based on Kailola (2004).

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Species inquirenda, Cephalocassis
Hemipimelodus intermedius Vinciguerra, 1881: 178, unnumbered figure. Type locality: [Sarawak, Borneo]. Syntypes:
MSNG 14529 (2). Treated as valid in Hemipimelodus (a synonym of Cephalocassis herein) by Desoutter (1977) but
as a possible synonym of Cephalocassis borneensis by Kailola (2004).
CINETODUS Ogilby, 1898
Cinetodus Ogilby, 1898a: 32. Type species: Arius froggatti Ramsay & Ogilby, 1886. Type by original designation.
Gender: Masculine.
Pachyula Ogilby, 1898a: 33. Type species: Hemipimelodus crassilabris Ramsay & Ogilby, 1886. Type by original
designation. Gender: Feminine.
Tetranesodon Weber, 1913: 545. Type species: Tetranesodon conorhynchus Weber, 1913. Type by monotypy. Gender: Masculine.
Septobranchus Hardenberg, 1941: 223. Type species: Septobranchus johannae Hardenberg, 1941. Type by monotypy. Gender: Masculine.
Cinetodus carinatus (Weber, 1913)
Arius (Hemiarius) carinatus Weber, 1913: 537, figs. 13–14. Type locality: Lorentz-Fluss; bei Van Weelskamp; Van
der Sande-Fluss, Nebenfluss des Lorentz-Flusses [New Guinea]. Syntypes: AMNH 9265 (2), ZMA 109295 (1),
ZMA 111109 (18), ZMA 111110 (1), ZMA 111111 (1), ZMA 111112 (1).
Distribution: Southern Papua (formerly Irian Jaya), New Guinea (Allen, 1996).
Cinetodus conorhynchus (Weber, 1913)
Tetranesodon conorhynchus Weber, 1913: 546, fig. 24. Type locality: Lorentz-Fluss, v. Weelskamp [New Guinea].
Holotype: ZMA 111084.
Distribution: Southern Papua (formerly Irian Jaya), New Guinea (Allen, 1996).
Cinetodus crassilabris (Ramsay & Ogilby, 1886)
Hemipimelodus crassilabris Ramsay & Ogilby, 1886: 18. Type locality: Strickland River, New Guinea. Holotype:AMS B.9961; dorsal view of head illustrated in Desoutter (1977: 18, fig. 4).
Distribution: New Guinea (Allen, 1991).
Cinetodus froggatti (Ramsay & Ogilby, 1886)
Arius froggatti Ramsay & Ogilby, 1886: 14. Type locality: Strickland River, New Guinea. Holotype: AMS B.9936.
Septobranchus johannae Hardenberg, 1941: 223, fig. 3. Type locality: Merauke, South New Guinea [Papua, Indonesia]. Holotype (about 30 cm): Whereabouts unknown.
Distribution: Southern New Guinea and Roper River system, Northern Australia; in coastal waters, estuaries, andinto fresh water (Kailola, in Allen, 1989).
COCHLEFELIS Whitley, 1941
Cochlefelis Whitley, 1941b: 8. Type species: Arius spatula Ramsay & Ogilby, 1886. Type by original designation.
Gender: Feminine.
Remarks: Elevated from a subgenus of Arius by Kailola (2004).
Cochlefelis burmanica (Day, 1870)
Arius burmanicus Day, 1870c: 618. Type locality: Irrawaddi, Bassein district, and Salwein in the Tenasserim provinces, Burma. Possible syntypes: AMS B.7520 (1), BMNH 1870.6.14.43–44 (2), RMNH 8787 (1), ZMB 2754
(1), ZSI A.568 (8), ZSI B.292 (1, lost), ZSI 456 (1); ZSI 456 illustrated in Day (1875–78: pl. 105) (Jayaram,
1982).
Distribution: Myanmar (Jayaram, 1982).
Remarks: Placement in Cochlefelis first proposed in Kailola (2004).
Cochlefelis danielsi (Regan, 1908)
Arius (Hemiarius) danielsi Regan, 1908c: 154. Type locality: Fly River, British New Guinea. Holotype: BMNH
1905.8.15.21.
Distribution: Southern New Guinea, in river deltas and mangroves (Kailola, 1999).

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Cochlefelis spatula (Ramsay & Ogilby, 1886)
Arius spatula Ramsay & Ogilby, 1886: 15. Type locality: Strickland River, New Guinea. Holotype: AMS B.9937.
Distribution: Southern Papua (formerly Irian Jaya), New Guinea (Allen, 1996).
CRYPTARIUS Kailola, 2004
Cryptarius Kailola, 2004: 134. Type species: Arius truncatus Valenciennes, 1840. Type by original designation.
Gender: Masculine.
Cryptarius daugueti (Chevey, 1932)
Hemipimelodus Daugueti Chevey, 1932: 41, pl. 13. Type locality: Phnompenh et Snoctrou, au seuil du Grand Lac.
Type(s): at Inst. Océanogr. Indochine, Nhatrang, Viet Nam.
Distribution: Large Rivers of South East Asia.
Remarks: Kailola (2004: 135) tentatively assigned this species to Cryptarius.
Cryptarius truncatus (Valenciennes, 1840)
Arius truncatus Valenciennes, in Cuvier & Valenciennes, 1840b: 64 (48 of Strasbourg deluxe edition). Type locality:
Java. Holotype: MNHN b-0590.
Hemipimelodus cochlearis Fowler, 1935b: 101, fig. 25. Type locality: Paknam, Siam. Holotype: ANSP 60767.
Distribution: Western central Pacific; in estuaries and lower portions of rivers (Kailola, 1999); including Chao
Phraya and Mekong (Rainboth, 1996).
GALEICHTHYS Valenciennes, 1840
Galeichthys Valenciennes in Cuvier & Valenciennes, 1840b: 28 (21 of Strasbourg deluxe edition). Type species:
Galeichthys feliceps Valenciennes, 1840. Type by subsequent designation by Bleeker (1862–63). Gender: Masculine.
Galeichthys ater Castelnau, 1861
Galeichthys ater Castelnau, 1861: 62. Type locality: les mers du Cap [South Africa]. Syntypes (2, 55 cm long):
Whereabouts unknown.
Distribution: Cape of Good Hope, Southern Africa (Taylor, 1986).
Galeichthys feliceps Valenciennes, 1840
Galeichthys feliceps Valenciennes, in Cuvier & Valenciennes, 1840b: 29 (22 of Strasbourg deluxe edition), pl. 424.
Type locality: Aux environs du Cap [South Africa]. Syntypes: MNHN a-9361 (1, dry), MNHN a-9362 (1, dry),
MNHN a-9363 (1, dry), MNHN a-9364 (1, dry), MNHN a-9365 (1, dry).
Bagrus capensis Smith, 1840, in Smith 1838–49: pl. 8. Type locality: Cape of Good Hope, South Africa. Possible
holotype: BMNH 1857.6.13.147 (1, skin). Name available from plate caption.
Pimelodus Peronii Valenciennes, in Cuvier & Valenciennes, 1840b: 161 (100 of Strasbourg deluxe edition). Type
locality: la mer des Indes. Holotype: MNHN 0000-1207.
Galeichthys ocellatus Gilchrist & Thompson, 1916: 60, figure on p. 61. Type locality: Zwartkops River, Algoa Bay[South Africa]. Holotype: at PEM.
Distribution: Widely distributed in southern Africa; in freshwater, estuaries, and coastal environments (Taylor,1986).
Remarks: Kailola (1986b) treated Pimelodus peronii as a valid species in Arius, but synonymy herein follows
Kailola (2004).
Galeichthys peruvianus Lütken, 1874
Galeichthys peruvianus Lütken, 1874b: 205. Type locality: Callao. Syntypes: ZMUC P 29628 (1), ZMUC P 29629(1).
Distribution: Pacific coast of South America, from Ecuador to Peru (Marceniuk & Ferraris, 2003).
GENIDENS Castelnau, 1855
Genidens Castelnau, 1855: 33. Type species: Bagrus genidens Cuv. Val. [= Pimelodus genidens Cuvier, 1829]. Typeby absolute tautonymy. Gender: Masculine.
Guiritinga Bleeker, 1858b: 62, 67. Type species: Pimelodus commersonii La Cepède, 1803. Type by monotypy.

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Gender: Feminine.
Genidens barbus (La Cepède, 1803)
Pimelodus barbus La Cepède, 1803: 94, 102. Type locality: Amérique méridionale. No types known.
Pimelodus Commersonii La Cepède, 1803: 95, 103, pl. 3 (fig. 1). Type locality: [No locality stated]. No types
known.
Bagrus barbatus Quoy & Gaimard, 1824: 230, pl. 49 (figs. 1–2). Type locality: Rio de la Plata, South America.
Type(s): at MNHN.
Pimelodus versicolor Castelnau, 1855: 35, pl. 16 (fig. 3). Type locality: rio Araguay, dans la province de Goyaz(Brésil). Syntypes: MNHN 0000-1206 (2).
Tachisurus upsulonophorus Eigenmann & Eigenmann, 1889b: 31. Type locality: Rio Grande do Sul [Brazil]. Holotype: MCZ 23750.
Silurus 16-radiatus Larrañaga, 1923: 386. Type locality: Uruguay. No types known. Apparently an alternate namefor Silurus marinus Larrañaga.
Silurus marinus Larrañaga, 1923: 376. Type locality: Uruguay. No types known. Apparently an alternate name for
Silurus 16-radiatus Larrañaga. Preoccupied by Silurus marinus Mitchill, 1815.
Distribution: Atlantic coast of South America from the La Plata River to eastern Brazil (Marceniuk, 2005).
Remarks: Redescribed in Marceniuk (2005b).
Genidens genidens (Cuvier, 1829)
Pimelodus genidens Cuvier, 1829: 294. Type locality: Not stated. Syntypes: MNHN 0000-1213 (2), MNHN b-0678(1). Species described in more detail, as Bagrus genidens, in Cuvier & Valenciennes (1840a: 453 (335 of Strasbourg deluxe edition), pl. 419), with locality given as Rio-Janéiro.
Genidens granulosus Castelnau, 1855: 34, pl. 16 (fig. 1). Type locality: la rivière d’Araguay, dans la province brésilienne de Goyaz. Holotype: MNHN 0000-1197.
Genidens cuvieri Castelnau, 1855: 34. Type locality: la Plata. Syntypes: MNHN 0000-1213 (2), MNHN b-0678 (1).
Unneeded replacement name for “Bagrus genidens Cuv. Val.” [= Pimelodus genidens Cuvier].
Genidens valenciennesii Bleeker, 1858b: 68. Type locality: Rio-Janéiro. Syntypes: MNHN 0000-1213 (2), MNHN
b-0678 (1). Unneeded replacement name for “Bagrus genidens CV.” [= Pimelodus genidens Cuvier].
Distribution: Atlantic draining rivers of southern South America (Marceniuk & Ferraris, 2003).
Genidens machadoi (Miranda Ribeiro, 1918)
Tachysurus machadoi Miranda Ribeiro, 1918b: 108. Type locality: Macaé, Rio de Janeiro, Brasil. Holotype: MNRJ
661.
Distribution: Atlantic coastal rivers of southern South America, from Rio de Janeiro to Argentina and, possibly,
Chile (Marceniuk & Ferraris, 2003).
Remarks: Redescribed in Marceniuk (2005b). Possible inclusion of this species in the fauna of Chile is based onreport by Sielfeld (1979).
Genidens planifrons (Higuchi, Reis & Araújo, 1982)
Netuma planifrons Higuchi, Reis & Araújo, 1982: 12, fig. 1. Type locality: Lagoa dos Patos, RS, Brazil. Holotype:
MZUSP 14828.
Distribution: Laguna dos Patos drainage of Southern Brazil and nearby Atlantic Ocean (Marceniuk & Ferraris,2003).
HEMIARIUS Bleeker, 1862
Hemiarius Bleeker, 1862 (in Bleeker, 1862–63): 7, 29. Type species: Cephalocassis stormii Bleeker, 1858. Type byoriginal designation. Gender: Masculine.
Hemiarius dioctes (Kailola, 2000)
Arius dioctes Kailola, 2000: 128, figs. 1–2. Type locality: Norman River at Karumba (17º29'S, 140º50'E), Queensland,
Australia. Holotype: CSIRO C.3798.
Distribution: Southern coast of New Guinea and rivers from Kamora River to Otokwa River, and eastward to Gulf ofPapua rivers; northern coast and rivers of Australia from Adelaide to the Gulf of Carpentaria (Kailola, 2000).
Hemiarius insidiator (Kailola, 2000)

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Arius insidiator Kailola, 2000: 131, figs. 3–4. Type locality: Mouth of Wildman River, (12º26'S, 132º09'E), Northern Territory, Australia. Holotype: NTM S.11189-001.
Distribution: Southern coast of New Guinea, along the Gulf of Papua, and northern coast of Australia, east of Darwin (Kailola, 2000).
Hemiarius sona (Hamilton, 1822)
Pimelodus sona Hamilton, 1822: 172, 376. Type locality: estuaries of Bengal. No types known. Unpublished Hamilton illustration reproduced in Hora (1929: pl. 19, fig. 7).
Distribution: South Asia and southern portion of Malay Peninsula; in coastal waters, estuaries, and lower portions ofrivers (Jayaram, 1982); also Mekong River delta and possibly into Cambodia (Rainboth, 1996).
Hemiarius stormii (Bleeker, 1858)
Cephalocassis Stormii Bleeker, 1858b: 246. Type locality: Sumatra (Palembang), in flumine Mussi. Syntypes (2:415–520 mm TL): possibly BMNH 1863.12.4.65 (1, 276 mm L, ~ 315 mm TL), but apparently too short. Illustrated in Bleeker (1862–63: pl. 100 [= Silur. pl. 52], as Hemiarius Stormi).
Distribution: Rivers of Sumatra and Borneo (Ng, 2003e).
Hemiarius verrucosus (Ng, 2003)
Arius verrucosus Ng, 2003e: 3, fig. 1. Type locality: Laos, Champasak, Mekong River at Ban Hang Khone, justdownstream from Khone Falls. Holotype: UMMZ 235408.
Distribution: Middle and lower Mekong River basin (Ng, 2003e).
Species inquirenda, Hemiarius
Hexanematichthys leptocassis Bleeker, 1861: 65. Type locality: Pinang. No types known (see Bleeker, 1862–63: 30, fordetails).
HEXANEMATICHTHYS Bleeker, 1858
Hexanematichthys Bleeker, 1858b: 61, 126. Type species: Bagrus sondaicus Valenciennes, 1840. Type by monotypy. Gender: Masculine.
Hexanematichthys mastersi (Ogilby, 1898)
Arius mastersi Ogilby, 1898a: 34. Type locality: Port Darwin, Northern Australia. Holotype: AMS I.25690-001.
Arius sagoroides Hardenberg, 1941: 221, fig. 2. Type locality: Merauke, ... mouth of the Oetakwa River, New
Guinea [Papua, Indonesia]. Syntypes (3, 24–32 cm): Whereabouts unknown.
Tachysurus (Pararius) godfreyi Whitley, 1941b: 11, fig. 8 (nos. 3–4). Type locality: Port Darwin, Northern Territory,
Australia. Holotype: AMS I.5270.
Distribution: Sahul Shelf (Kailola, 2004).
Hexanematichthys sagor (Hamilton, 1822)
Pimelodus sagor Hamilton, 1822: 169, 376. Type locality: estuaries of Bengal. No types known.
Bagrus doroides Valenciennes, in Cuvier & Valenciennes, 1840a: 448 (331 of Strasbourg deluxe edition), pl. 418.
Type locality: Pondichéry ... Bengale. Syntypes: MNHN a-8668 (1), MNHN a-9349 (1).
Bagrus Javensis Valenciennes, in Cuvier & Valenciennes, 1840a: 445 (330 of Strasbourg deluxe edition). Type
locality: Java. Holotype (6 pouces): Whereabouts unknown.
Bagrus sondaicus Valenciennes, in Cuvier & Valenciennes, 1840a: 444 (329 of Strasbourg deluxe edition). Type
locality: dans le détroit de Sonde. Holotype: MNHN b-0604.
Hexanematichthys sundaicus Bleeker, 1858c: 2. Type locality: Strait of Sonde. Holotype: MNHN b-0604. Unjustified emendation of Bagrus sondaicus Valenciennes, 1840.
Distribution: Sunda shelf (Kailola, 2004); also Pakistan and west coast of India (Jayaram, 1982).
KETENGUS Bleeker, 1847
Ketengus Bleeker, 1847a:167. Type species: Ketengus typus Bleeker, 1847. Type by original designation. Gender:
Masculine.
Ketengus typus Bleeker, 1847

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Ketengus typus Bleeker, 1847b: 9. Type locality: Banjermassing, in fluviis. Syntype: BMNH 1863.12.4.112 (1, 191
mm SL).
Pimelodus pectinidens Cantor, 1849: 1243. Type locality: Pinang. Holotype (4 2/8 inches): BMNH 1860.3.19.499,
one-half skin only.
Distribution: Eastern Bay of Bengal and Malay Peninsula; in nearshore coastal waters, estuaries, and lower reachesof rivers (Kailola, 1999).
Remarks: Redescribed in Jayaram (1972b).
NEDYSTOMA Ogilby, 1898
Nedystoma Ogilby, 1898a: 32. Hemipimelodus dayi Ramsay & Ogilby, 1886. Type by original designation. Gender:
Neuter.
Doiichthys Weber, 1913: 532. Type species: Doiichthys novaeguineae Weber, 1913. Type by monotypy. Gender:
Masculine.
Remarks: Kailola (2004) considered Doiichthys novaeguineae to be the sister group to Nedystoma dayi and, on thatbasis, placed the two formerly-monotypic genera into synonymy.
Nedystoma dayi (Ramsay & Ogilby, 1886)
Hemipimelodus dayi Ramsay & Ogilby, 1886: 16. Type locality: Strickland River, New Guinea. Holotype: AMS
B.9938.
Distribution: Southern Papua (formerly Irian Jaya), New Guinea (Allen, 1996).
Nedystoma novaeguineae (Weber, 1913)
Doiichthys novae-guineae Weber, 1913: 534, fig. 12. Type locality: “Varen”-Fluss, einem Nebenfluss des LorentzFlusses [New Guinea]. Syntypes: AMNH 9482 (1), ZMA 104122 (5).
Distribution: Southern New Guinea, in estuaries and lower reaches of rivers (Kailola, 1999).
NEMAPTERYX Ogilby, 1908
Nemapteryx Ogilby, 1908: 3, 10. Type species: Arius stirlingi Ogilby, 1898. Type by original designation. Gender:
Feminine.
Nemapteryx armiger (De Vis, 1884)
Arius armiger De Vis, 1884: 454. Type locality: New Britain [in error]. Syntypes: QM I.3088 (1), QM I.3039 (1).
Arius stirlingi Ogilby, 1898b: 281. Type locality: Estuary of the Adelaide River, Northern Territory [Australia].
Holotype: SAMA F1095.
Distribution: Southern New Guinea and northern Australia; in coastal waters, estuaries, and lower reaches of rivers(Kailola, 1999).
Remarks: See Kailola (1983, 2004) for synonymy.
Nemapteryx augusta (Roberts, 1978)
Arius augustus Roberts, 1978: 36, figs. 14, 20d. Type locality: Mouth of Binge River, a large strongly flowing tributary of the Middle Fly, 675 km upriver from Toro Pass, 6º32.5'S, 140º55.0'E; Fly River, Papua New Guinea.
Holotype: AMS I.27090-001.
Distribution: Middle reaches of the Fly River (Roberts, 1978).
? Nemapteryx bleekeri (Popta, 1900)
Arius bleekeri Popta, 1900: 71. Type locality: Malay Archipelago. Syntypes: RMNH 6825 (4).
Distribution: Malay Archipelago.
Remarks: Validity and placement in Nemapteryx considered tentative by Kailola (2004).
Nemapteryx caelata (Valenciennes, 1840)
Arius cælatus Valenciennes, in Cuvier & Valenciennes, 1840b: 66 (49 of Strasbourg deluxe edition). Type locality:
Batavia ... Bombay. Syntypes: MNHN b-0589 (1), MNHN b-0614 (2).
Distribution: Widely distributed from Pakistan to Borneo (Jayaram, 1982).
Remarks: Generally treated as valid, but Kailola (1999: 1859; 2004: 139) indicated that it might, instead, be a synonym of Nemapteryx nenga (Hamilton, 1822).
Nemapteryx macronotacantha (Bleeker, 1846)

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Arius macronotacanthus Bleeker, 1846a: 159. Type locality: Batavia. Holotype (size not stated): possibly RMNH
6901 (but see BMNH 1863.12.4.59). [See also Bleeker, 1847c.]? Arius parvipinnis Day, 1877 (in Day, 1875–78): 460, pl. 113 (fig. 1). Coromandel coast of India. Syntypes: Whereabouts unknown.
Distribution: Bay of Bengal and western central Pacific; in coastal waters and estuaries (Kailola, 1999).
Nemapteryx nenga (Hamilton, 1822)
Pimelodus nenga Hamilton, 1822: 171, 376. Type locality: estuaries of Bengal. No types known.
Distribution: Eastern Arabian Sea, Bay of Bengal, and western central Pacific; in nearshore turbid waters and lower
reaches of rivers (Kailola, 1999).
NEOARIUS Castelnau, 1878a
Neoarius Castelnau, 1878a: 237. Type species: Arius curtisii Castelnau, 1878. Type by monotypy. Gender: Masculine.
Remarks: Treated as a synonym of Ariopsis in Kailola (2004); treated herein as valid for the Australia–New Guinea
species of Kailola’s Ariopsis.
Neoarius berneyi (Whitley, 1941)
Tachysurus (Pararius) berneyi Whitley, 1941b: 9, fig. 8 (no. 5). Type locality: pools of the Flinders River, near
Hughenden and Richmond, Queensland (freshwater), Australia. Holotype: AMS I.13076.
Arius cleptolepis Roberts, 1978: 37, figs. 15, 20e. Type locality: Elevala River, a major lowland tributary of theUpper Fly, 17 km E of Kiunga, 859 km upriver from Toro Pass, 6º05.7'S, 141º27.7'E; Fly River, Papua New
Guinea. Holotype: AMS I.27092-001.
Distribution: Northern Australia and central-southern New Guinea (Kailola, in Allen, 1989).
Neoarius coatesi (Kailola, 1990)
Arius coatesi Kailola, 1990: 18, figs. 5. Type locality: Market at Angoram, Sepik River, Papua New Guinea. Holotype: AMS I.25405-001.
Distribution: Sepik River basin, Papua New Guinea (Kailola, 1990).
Neoarius graeffei (Kner & Steindachner, 1867)
Arius Graeffei Kner & Steindachner, 1867: 383, pl. 4 (fig. 12). Type locality: Samoa-Inseln [in error]. Holotype:
NMW 67152.
Arius australis Günther, 1867d: 103, figure on p. 103. Type locality: Hunter River, New South Wales, near Ash
Island [Australia]. Syntypes: BMNH 1866.2.13.4 (1), BMNH 1866.6.19.7–8 (2).
Arius curtisii Castelnau, 1878a: 236. Type locality: Moreton Bay, Australia. Syntypes (several, to 15 inches):
MNHN b-0693 (1).
Distribution: Southern New Guinea and northern Australia, in rivers and along coast (Kailola, 1999).
Remarks: Synonymy follows Kailola (1983 and 2004).
Neoarius latirostris (Macleay, 1883)
Arius latirostris Macleay, 1883: 277, unnumbered figure. Type locality: Goldie River, New Guinea. Syntypes: AMS
I.9072 (1), AMS I.9073 (1), AMS I.9074 (1), AMS I.9127 (1), AMS I.13398 (1).
Arius acrocephalus Weber, 1913: 543, figs. 20–21. Type locality: Van der Sande-Fluss (Bibis-Fluss), ein Nebenflussdes Lorentz-Flusses; Lorentz-Fluss bei Regen-Insel; Lorentz-Fluss bei Alkmaar [New Guinea]. Syntypes:
AMNH 9514 (1), ZMA 111087 (1), ZMA 111088 (2), ZMA 111089 (1), ZMA 111090 (1).
Arius digulensis Hardenberg, 1936b: 369. Type locality: Middle course of Digulriver [southern New Guinea]. Holotype: ZMA 110781.
Distribution: Southern Papua (formerly Irian Jaya) (Allen, 1996), Goldie River, Papua New Guinea.
Neoarius leptaspis (Bleeker, 1862)
Hexanematichthys leptaspis Bleeker, 1862 (in Bleeker, 1862–63): 27, pl. 65. Type locality: Nova Guinea austrooccidentalis. Holotype: RMNH 3060.
Distribution: Southern New Guinea and northern Australia, in coastal waters and rivers (Kailola, 1999).
Neoarius midgleyi (Kailola & Pierce, 1988)
Arius midgleyi Kailola & Pierce, 1988: 75, figs. 1–2, 5–6. Type locality: Wickham Gorge, Victoria River, Northern

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Territory, Australia. Holotype: AMS I.20858-006.
Distribution: Victoria, Katherine, Daly, Ord, remaining Kimberley and northern River systems, Northern Australia(Kailola, 2000).
Neoarius paucus (Kailola, 2000)
Arius paucus Kailola, 2000: 141. Type locality: Flinders River near Maxwellton (20º47'S, 142º43'E), Queensland,
Australia. Holotype: QM I.12910.
Distribution: Roper and Flinders River systems, and other rivers draining into Gulf of Carpentaria, Australia(Kailola, 2000).
Neoarius pectoralis (Kailola, 2000)
Arius pectoralis Kailola, 2000: 133, figs. 5–6. Type locality: Chapman River, Queensland, 14º56'S, 141º38'E, Australia. Holotype: AMS I.27415-001.
Distribution: Southern coast of New Guinea and into estuaries and tidal rivers, and northern Australia from Darwin
to the Chapman River, Queensland (Kailola, 2000).
Neoarius taylori (Roberts, 1978)
Hemipimelodus taylori Roberts, 1978: 40, figs. 19, 20h. Type locality: Mainstream of Plamer and lower end of small
tributary about 1 km up the Palmer from Thompson Junction (mouth of Wai Mungi), 6º46'48"S, 141º36'36"E;
Fly River, Papua New Guinea. Holotype: AMS I.27087-001. Previously considered preoccupied in Arius by
Arius taylori Hildebrand, 1925; replaced by Arius robertsi Kailola, 1990 [see Remarks].
Arius robertsi Kailola, 1990: 24. Type locality: Mainstream of Plamer and lower end of small tributary about 1 km
up the Palmer from Thompson Junction (mouth of Wai Mungi), 6º46'48"S, 141º36'36"E; Fly River, Papua New
Guinea. Holotype: AMS I.27087-001. Replacement for Hemipimelodus taylori Roberts, 1978, when preoccupied in Arius by Arius taylori Hildebrand, 1925..
Distribution: Fly River, Papua New Guinea (Roberts, 1978).
Remarks: Arius robertsi Kailola, 1990, was proposed as a replacement for Hemipimelodus taylori Roberts, 1978,
which was then considered to be preoccupied in Arius by Arius taylori Hildebrand, 1925. Kailola (2004) subsequently placed A. taylori Hildebrand in Cathorops, as a junior synonym of C. steindachneri (Gilbert & Starks),and H. taylori Roberts in Ariopsis, as a synonym of Ariopsis robertsi Kailola. In adopting the interpretation of
Kailola (2004), Hemipimelodus taylori Roberts is no longer treated as congeneric with Arius taylori and, following Art. 59.4 of the Code (ICZN, 1999) H. taylori must be reinstated as valid, with Arius robertsi as an
objective junior synonym.
Neoarius utarus (Kailola, 1990)
Arius utarus Kailola, 1990: 12, figs. 4. Type locality: Murik Lakes, New Guinea. Holotype: AMS I.25406-001.
Distribution: Northern New Guinea; in freshwater and estuaries (Kailola, 1990).
Neoarius velutinus (Weber, 1907)
Hemipimelodus velutinus Weber, 1907: 225. Type locality: Sentani-see, Nord-Neu-Guinea; Mündung des Tami,
Nord-Neu-Guinea; Fluss Tawarin, Nord-Neu-Guinea. Syntypes (31): AMNH 9478 (1), FMNH 52392 (1),
NMW 8222 (1), NMW 8223 (1), RMNH 8001 (1), ZMA 112654 (3), ZMA 112655 (6), ZMA 112656 (12).
Hemipimelodus papillifer Herre, 1935: 390. Type locality: Timbunke, a village on the Sepik River, about 120 miles
from the sea, New Guinea. Holotype: FMNH 17211.
Distribution: Northern Papua (formerly Irian Jaya) (Allen, 1996) and Sepik River basin.
NETUMA Bleeker, 1858
Netuma Bleeker, 1858b: 62, 93. Type species: Bagrus netuma Valenciennes, 1840. Type by absolute tautonymy.
Gender: Feminine.
Pararius Whitley, 1940b: 409. Type species: Arius proximus Ogilby, 1898. Type by original designation. Gender:
Masculine. Originally proposed as a subgenus of Tachysurus.
Remarks: The names Catastoma (in Cuvier & Valenciennes, 1840b: 60), and Sarcogenys (in Bleeker, 1858b: 96),
were not published in a manner that made them available for purposes of zoological nomenclature and are therefore not listed above as available names; Kailola (2004) considered both to refer to alternate names for Netuma
Bleeker.

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Netuma bilineata (Valenciennes, 1840)
Bagrus bilineatus Valenciennes, in Cuvier & Valenciennes, 1840a: 434 (322 of Strasbourg deluxe edition). Type
locality: Pondichéry ... Rangoon. Syntype: MNHN a-9344 (1, mounted).
Bagrus rhodonotus Bleeker, 1846a: 157. Type locality: Batavia. Syntypes (size and number not stated): RMNH
15865 (5), SMNS 10568 (1). [See also Bleeker, 1847c.]
Netuma osakae Jordan & Kasawa, in Jordan & Hubbs, 1925: 157, pl. 9 (fig. 1). Type locality: Fish market, Osaka,
Japan. Holotype: FMNH 59388.
Arius dayi Dmitrenko, 1974: 39, figs. 1–2. Type locality: Masir Strait, Gulf of Oman, 19º47.8'N, 58º28.3'E. Holotype: Zool. Mus. Ukranian Acad. Kiev Pi 2/1–2.
Distribution: Arabian Sea, Bay of Bengal, Western Central Pacific and northern coast of Australia; from estuaries
out to continental shelf (Kailola, 1999).
Remarks: See Kailola (1986a) for discussion of validity of this species. See Wang et al. (2005b) for comments on the
distribution of this species.
Netuma proxima (Ogilby, 1898)
Arius proximus Ogilby, 1898b: 280. Type locality: Port Darwin, Northern Australia. Holotype: AMS I.25691-001.
Arius arafurensis Hardenberg, 1948: 409, fig. on p. 409. Type locality: Fish market of Dobo (Aru Islands), IndoAustralian Archipelago. Holotype (about 320 mm SL): Whereabouts unknown.
Distribution: Southern New Guinea, northern Australia and Aru Islands; in nearshore coastal waters and estuaries(Kailola, 1999).
Netuma thalassina (Rüppell, 1837)
Bagrus thalassinus Rüppell, 1837, in Rüppell, 1835–38: 75, pl. 20 (fig. 2). Type locality: Massawa, Eritrea, Red
Sea. Syntypes SMF 544 (1), SMF 5414 (1, dry), SMF 2627 (1, dry), SMF 5740 (1, skeleton).
Bagrus laevigatus Valenciennes, in Cuvier & Valenciennes, 1840a: 439 (325 of Strasbourg deluxe edition). Type
locality: la Mer Rouge. Holotype: MNHN b-0710.
Arius nasutus Valenciennes, in Cuvier & Valenciennes, 1840b: 60 (45 of Strasbourg deluxe edition). Type locality:
Java ... Malabar ... la Mer Rouge. Syntypes: MNHN a-9407 (1), MNHN a-9408 (1, dry), MNHN a-9409 (1,dry), RMNH D2390 (1).
Bagrus netuma Valenciennes, in Cuvier & Valenciennes, 1840a: 438 (324 of Strasbourg deluxe edition), pl. 417.
Type locality: Pondichéry. Syntype: MNHN a-9345 (1).
Bagrus carchariorhijnchos Bleeker, 1846b: 291. Type locality: Batavia, Java. Holotype: RMNH 6885.
Arius serratus Day, 1877 (in Day, 1875–78): 462, pl. 105 (fig. 3). Type locality: Sind [Pakistan]. Holotype: ZSI 467.
Ariodes aeneus Sauvage, 1883c: 160. Type locality: Ile Rafflès. Syntypes: MNHN a-5155 (2).
Netuma thalassina jacksonensis Whitley, 1941b: 12, fig. 8 (nos. 1–2). Type locality: Port Jackson, New South Wales[Australia]. Holotype: AMS I.10095.
Distribution: Widely distributed through Red Sea, Indian Ocean and western central Pacific; in coastal waters out tocontinental shelf margin.
Remarks: See Kailola (1986a) and Wang (2005b) for discussion of validity. See Jayaram & Dhanze (1978: 203) andKailola (1999: 1842, 1865) for comments on status of Arius serratus.
Species inquirenda, Netuma
Galeichthys stanneus Richardson, 1846: 284. Type locality: Canton, China. No types known. Appeared as ? Galeichthys
stanneus. Based solely on illustration by Reeves which was reproduced in Whitehead (1970: 210, pl. 20a).
NOTARIUS Gill, 1863
Notarius Gill, 1863: 171. Type species: Arius grandicassis Valenciennes, 1840. Type by monotypy. Gender: Masculine.
Remarks: The limits of Notarius are not clearly diagnosed. Kailola (2004) included some of the species listed here in
Aspistor.
Notarius biffi Betancur & Acero, 2004

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Notarius biffi Betancur & Acero, 2004: 3, figs. 1–3. Type locality: Bahía La Unión, El Salvador (SV) (13º20'42"N,
87º49'07"W. Holotype: STRI 6674.
Distribution: Tropical Eastern Pacific from Costa Rica to El Salvador (Betancur & Acero, 2004).
Notarius cookei (Acero & Betancur, 2002)
Arius cookei Acero & Betancur, 2002a: 134, fig. 1. Type locality: 7 km from the mouth of Rio Santa Maria, Paris,
Herrera, Panama (8º9'N, 80º33'W). Holotype: INVEMAR-PEC 3752.
Distribution: Pacific coastal rivers and estuaries of Central and South America, from Costa Rica to Colombia (Acero& Betancur, 2002).
Notarius grandicassis (Valenciennes, 1840)
Arius grandicassis Valenciennes, in Cuvier & Valenciennes, 1840b: 54 (41 of Strasbourg deluxe edition), pl. 427.
Type locality: la Guyane. Holotype: MNHN a-4608.
Arius parmocassis Valenciennes, in Cuvier & Valenciennes, 1840b: 57 (43 of Strasbourg deluxe edition). Type locality: Bahia. Syntypes: MHNG (several), MNHN a-8966.
Arius stricticassis Valenciennes, in Cuvier & Valenciennes, 1840b: 58 (44 of Strasbourg deluxe edition). Type locality: Cayenne. Syntypes: MNHN a-9406 (1), RMNH 3034 (1).
Arius vandeli Puyo, 1936: 123, fig. 21g. Type locality: [French Guiana]. Holotype: MNHN 1936-0160.
Distribution: Rivers and estuaries of northern South America from Gulf of Venezuela to the mouth of the Amazon
River (Marceniuk & Ferraris, 2003).
Notarius insculptus (Jordan & Gilbert, 1882)
Arius elatturus Jordan & Gilbert, 1882: 45. Type locality: Panama. Syntypes (3): USNM 29408 (1, missing), USNM
30995 (1 of 2).
Arius insculptus Jordan & Gilbert, 1882: 41. Type locality: Panama. Syntypes: USNM 29415 (1), USNM 30977 (2).
Betancur & Acero (2004) referred to USNM 29415 as a lectotype, but no previous lectotype designation has
been found, and indications of a lectotype in that publication do not meet the requirements of the Code for lectotype designation. Because no previously published lectotype designation was found, the specimens listed here
are considered as syntypes.
Netuma insularum Greene, in Gilbert, 1897: 439. Type locality: Galapagos Archipelago [later corrected to: Gulf ofPanama]. Holotype: USNM 47577.
Distribution: Pacific coast of Central America, from Costa Rica to Panama (Marceniuk & Ferraris, 2003).
Remarks: Jordan & Gilbert (1882) were reported to have designated a holotype for Arius elatturus by Ferraris &Vari (1992), Eschmeyer et al. (1998), Marceniuk & Ferraris (2003). However, a closer reading of the paper
shows that the specimen subsequently cited as holotype (USNM 29408) was the basis for the species description but otherwise not explicitly designated as the type. Therefore, it must instead be considered to be a syntype
as one of three “specimens at the United States National Museum” representing the new species. Betancur &Acero (2004: 7) appear to be have acted as first reviser in treating Arius insculptus as a valid species in Notarius, with Arius elatturus as a synonym.
Notarius kessleri (Steindachner, 1876)
Arius Kessleri Steindachner, 1876: 574, pl. 5. Type locality: Panama, Altata. Syntypes: MCZ 4944 (6), NMW 48112(1), NMW 48247 (1), NMW 48248 (1), NMW 48249 (2).
Distribution: Eastern tropical Pacific Ocean (Marceniuk & Ferraris, 2003).
Notarius neogranatensis (Acero & Betancur, 2002)
Arius neogranatensis Acero & Bentancur, 2002b: 7, fig. 1. Type locality: mouth of Río Tinajones medio, San
Antero, Córdoba, Colombia (9 25'N 75 48'W). Holotype: INVEMAR-PEC 3754.
Distribution: Caribbean coast of Colombia and nearby rivers mouths (Acero & Betancur, 2002).
Notarius osculus (Jordan & Gilbert, 1882)
Arius osculus Jordan & Gilbert, 1882: 46. Type locality: Panama. Holotype: USNM 29476.
Distribution: Pacific coast of Central America, from Costa Rica and Panama (Marceniuk & Ferraris, 2003).
Remarks: Placed by Kailola & Bussing (1995: 871) tentatively in Arius.
Notarius planiceps (Steindachner, 1876)
Arius planiceps Steindachner, 1876: 576, pl. 4. Type locality: Panama, Altata. Syntypes: NMW 48194 (1), NMW

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48195 (1), NMW 48196 (1), NMW 48197 (1), NMW 48198 (1), NMW 48199 (2), ZMUC P 29627 (1).
Distribution: Pacific Ocean draining rivers and estuaries of North and Central America from Mexico to Panama(Marceniuk & Ferraris, 2003).
OSTEOGENEIOSUS Bleeker, 1846
Osteogeneiosus Bleeker, 1846a:173. Type species: Arius militaris Valenciennes, 1840 (= Silurus militaris Linnaeus,
1758). Type by subsequent designation, by Bleeker (1862–63: 8). Gender: Masculine.
Osteogeneiosus militaris (Linnaeus, 1758)
Silurus militaris Linnaeus, 1758: 305. Type locality: Asia. No types known.
? Osteogeneiosus Blochii Bleeker, 1846a: 175. Type locality: Batavia. Type(s) (size and number not stated): Whereabouts unknown [See also Bleeker, 1847: 50.]
? Osteogeneiosus gracilis Bleeker, 1846a: 175. Type locality: Batavia. Type(s) (size and number not stated): Whereabouts unknown [See also Bleeker, 1847:50.]? Osteogeneiosus ingluvies Bleeker, 1846a: 174. Type locality: Batavia. Type(s) (size and number not stated):
Whereabouts unknown [See also Bleeker, 1847: 50.]
? Osteogeneiosus longiceps Bleeker, 1846a: 174. Type locality: Batavia. Type(s) (size and number not stated):
Whereabouts unknown [See also Bleeker, 1847: 50.]
? Osteogeneiosus macrocephalus Bleeker, 1846a: 173. Type locality: Batavia. Syntypes (size and number notstated): possibly BMNH 1863.12.4.51 (1, 214 mm SL, 252 mm TL), NMV 45847 (1).
? Osteogeneiosus valenciennesi Bleeker, 1846a: 175. Type locality: Batavia. Syntypes (size and number not stated):
possibly BMNH 1863.12.4.53 (1245 mm SL, 289 mm TL), NMV 46593 (1), NMV 46594 (1).
? Osteogeneiosus Cantoris Bleeker, 1854c: 120, pl. 2 (fig. 4). Type locality: Calcutta, in flumine Hooghly. Syntype(2, 199 – 225 mm): RMNH 6907 (1).
? Osteogeniosus sthenocephalus Day, 1877 (in Day, 1875–78): 469, pl. 108 (fig. 3). Type locality: Moulmein[Burma]. Holotype: ZSI 1263. Name spelled Osteogeniosus stenocephalus on plate caption; first reviser not yet
found.
Distribution: Bay of Bengal and western central Pacific, in inshore waters, river mouths, and coastal lake systems(Kailola, 1999); also Pakistan and west coast of India (Jayaram, 1982).
Remarks: At present, there is only one recognized valid species in Osteogeneiosus. On that basis, all of the othernames are tentatively placed here as synonyms of O. militaris.
PLICOFOLLIS Kailola, 2004
Plicofollis Kailola, 2004: 141. Type species: Arius argyropleuron Valenciennes, 1840. Type by original designation.
Gender: Masculine.
Plicofollis argyropleuron (Valenciennes, 1840)
Arius argyropleuron Valenciennes, in Cuvier & Valenciennes, 1840b: 104 (78 of Strasbourg deluxe edition). Type
locality: Java. Holotype: RMNH 3041.
Arius acutus Bleeker, 1846a: 167. Type locality: Batavia. Syntypes (size and number not stated): possibly BMNH
1863.12.4.67 (1), RMNH 15975 (2), SMNS 10566 (4).
Arius Hamiltonis Bleeker, 1846b: 291. Type locality: [Batavia]. Syntypes (size and number not stated): RMNH 6895
(at least part of series).
Arius macrocephalus Bleeker, 1846a: 167. Type locality: Batavia. Syntypes (size and number not stated): BMNH
1863.12.4.78 (1), RMNH 6896 (1). One syntype illustrated in Bleeker (1862–63: 39, pl. 58 [Silur. pl. 10], as
Ariodes macrocephalus.
Arius layardi Günther, 1866: 474, pl. 15. Type locality: Ceylon. Holotype: BMNH 1866.7.13.13.
Tachysurus broadbenti Ogilby, 1908: 8. Type locality: Cape York, Queensland, Australia. Syntypes (3): QM I.9745(1), QM I.9746 (1).
Hemipimelodus colcloughi Ogilby, 1910: 7. Type locality: Croker Island, N.T., Australia. Holotype: QM I.1538.
Distribution: Bay of Bengal, western Pacific Ocean, southern coast of New Guinea and northern coast of Australia;
in coastal waters and in lower portions of estuaries (Kailola, 1999).

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Remarks: Synonymy from Kailola (1999).
Plicofollis dussumieri (Valenciennes, 1840)
Arius Dussumieri Valenciennes, in Cuvier & Valenciennes, 1840b: 84 (63 of Strasbourg deluxe edition). Type locality: la côte de Malabar. Holotype: MNHN a-9401.
Arius kirkii Günther, 1864: 163, unnumbered figure. Type locality: Shupanga, Zambezi [River]. Holotype: BMNH
1864.1.9.54 (skin).
Arius goniaspis Bleeker, 1858a: 44. Type locality: Priaman [Sumatra, Indonesia]. Syntypes (2): BMNH
1863.12.4.159 (1, 71 mm SL, 84 mm TL).
Distribution: East coast of Africa, Madagascar, Arabian Sea, Bay of Bengal, and eastern Sumatra; possibly along
eastern part of Malay peninsula; in coastal waters and estuaries (Taylor, 1986; Kailola, 1999).
Remarks: Register number of holotype of Arius goniaspis incorrectly reported as BMNH 1863.12.11.159 in Eschmeyer et al. (1998).
Plicofollis nella (Valenciennes, 1840)
Pimelodus nella Valenciennes, in Cuvier & Valenciennes, 1840b: 162 (121 of Strasbourg deluxe edition). Type
locality:[Vizagapatam]. Holotype: not saved, based on Russell (1803: pl. 170). Originally as Pimelodus ? nella.
Arius leiotetocephalus Bleeker, 1846b: 292. Type locality: Batavia. Syntypes: RMNH 5277 (1), RMNH 16011 (5),SMNS 817 (2).
Bagrus (Ariodes) Meyenii Müller & Troschel, 1849: 9. Type locality: unbekannt. Holotype: ZMB 3002.
Distribution: Bay of Bengal, western central Pacific and northern coast of Australia.
Plicofollis platystomus (Day, 1877)
Arius platystomus Day, 1877 (in Day, 1875–78): 464, pl. 107 (fig. 3). Type locality: Canara [India]. Holotype: ZSI
1306.
Distribution: Kanara, Malabar and Konkan coasts, India, and Hoogly River mouth (Jayaram, 1982).
Plicofollis polystaphylodon (Bleeker, 1846)
Arius polystaphylodon Bleeker, 1846a: 172. Type locality: Batavia. Holotype or Syntype (size and number notstated): BMNH 1863.12.4.98 (1, 121 mm SL).
Distribution: Western central Pacific, eastward to northern New Guinea; in turbid coastal waters, and river mouths(Kailola, 1999).
Plicofollis tenuispinis (Day, 1877)
Arius tenuispinis Day, 1877 (in Day, 1875–78): 466, pl. 107 (fig. 5). Type locality: Bombay, and perhaps Ceylon.
Holotype: ZSI F482.
Distribution: Coast of India and eastern Africa (Jayaram, 1982).
Remarks: Redescribed in Talwar (1976), with illustration of oral tooth patterns.
Plicofollis tonggol (Bleeker, 1846)
Arius crossocheilos Bleeker, 1846a: 172. Type locality: Batavia. Syntypes (size and number not stated): RMNH
6894 (1 or more of 3).
Arius tonggol Bleeker, 1846b: 292. Batavia. Syntypes: BMNH 1863.12.4.56 (1, 183 mm SL, ~ 246 mm TL), RMNH
6894.
Distribution: Western Central Pacific, and Bay of Bengal; in estuaries (Kailola, 1999).
Remarks: The names Arius crossocheilos and A. tonggol were first made available in separate publications, but inthe same issue of Natuur-en Geneeskundig Archif voor Neerlandsch Indië. As such, they must be treated as having been published simultaneously and, therefore, in need of first reviser action to establish precedence. The
earliest first reviser action I found was Bleeker (1862 [in Bleeker, 1862–63]: 39) in which Ariodes tonggol istreated as valid with Arius crossocheilos a synonym. First included in Plicofollis by Kailola (2004), but as Plicofollis crossocheilos.
Species inquirenda, Plicofollis
Arius magatensis Herre, 1926: 396, pl. 1 (fig. 1). Type locality: Magat River at Bagabag, Nueva Vizcaya Province, Philippines. Syntypes (6): BSMP (destroyed).

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Potamarius Hubbs & Miller, 1960: 101. Type species: Conorhynchos nelsoni Evermann & Goldsborough, 1902.
Type by original designation. Gender: Masculine.
Potamarius grandoculis (Steindachner, 1877)
Arius grandoculis Steindachner, 1877b: 644, pl. 11. Type locality: Rio doce (nach Salmin), südöstlichen Brasilien.
Holotype: NMW 48272. Name spelled Arius granducolis in plate caption.
Distribution: Doce and Paraíba do Sul rivers of Brazil, at and near their mouths (Marceniuk & Ferraris, 2003).
Potamarius izabalensis Hubbs & Miller, 1960
Potamarius izabalensis Hubbs & Miller, 1960: 104, figs. 2. Type locality: Lago de Izabal, Guatemala, 1 mi. west of
El Paraíso. Holotype: USNM 134347.
Distribution: Central America: Izabal Lake and, possibly, Polochic River, Guatemala (Marceniuk & Ferraris, 2003).
Potamarius nelsoni (Evermann & Goldsborough, 1902)
Conorhynchos nelsoni Evermann & Goldsborough, 1902: 140, fig. 1. Type locality: Río Usumacinta, Montecristo,
Yucatán, Mexico. Holotype: USNM 50001.
Distribution: Usumacinta River basin, Mexico and Guatemala (Miller et al., 2005).
SCIADES Müller & Troschel, 1849
Sciades Müller & Troschel, 1849: 6. Type species: Bagrus (Sciades) emphysetus Müller & Troschel, 1849. Type by
subsequent designation by Bleeker (1862–63: 8). Gender: Masculine. Originally proposed as a subgenus of
Bagrus.
Sciadeichthys Bleeker, 1858b: 62, 66. Type species: Bagrus (Sciades) emphysetus Müller & Troschel, 1849. Type bymonotypy. Gender: Masculine.
Selenaspis Bleeker, 1858b: 62. Type species: Silurus herzbergii Bloch, 1794. Type by subsequent designation,
apparently first by Jordan & Evermann (1896: 119). Gender: Feminine.
Leptarius Gill, 1863: 170. Type species: Leptarius dowii Gill, 1863. Type by monotypy. Gender: Masculine.
Sciadeops Fowler, 1944c: 211. Type species: Sciades troschelii Gill, 1863. Type by original designation. Gender:
Masculine. Originally proposed as a subgenus of Sciades.
Sciades couma (Valenciennes, 1840)
Bagrus couma Valenciennes, in Cuvier & Valenciennes, 1840a: 459 (340 of Strasbourg deluxe edition). Type locality: Cayenne. Syntypes: MNHN a-9338 (1), MNHN a-9339 (1).
Sciadeichthys (Selenaspis) walrechti Boeseman, 1954: 60, pl. 3. Type locality: Surinam, probably neighbourhood ofParamaribo. Holotype: RMNH 2141.
Distribution: Atlantic draining rivers of South America, from the Gulf of Paria to the mouth of the Amazon (Marceniuk & Ferraris, 2003).
Sciades dowii (Gill, 1863)
Leptarius dowii Gill, 1863: 171. Type locality: Western coast of Central America. Holotype: at USNM (not found).
Arius alatus Steindachner, 1876: 569, pl. 6. Type locality: Panama. Syntype: MCZ 7725.
Distribution: Pacific draining rivers and estuaries of South and Central America, from Panama to Ecuador (Marceniuk & Ferraris, 2003).
Sciades herzbergii (Bloch, 1794)
Silurus Herzbergii Bloch, 1794: 33, pl. 367. Type locality: Surinam. Syntypes: ZMB 2962 (1), ZMB 8785 (1, skin).
Pimelodus argenteus La Cepède, 1803: 94, 102. Type locality: Surinam. Syntypes: ZMB 2962 (1), ZMB 8785 (1).
Unneeded new name for Silurus herzbergii Bloch.
Bagrus mesops Valenciennes, in Cuvier & Valenciennes, 1840a: 456 (338 of Strasbourg deluxe edition). Type locality: [Puerto Rico]. Holotype: MNHN a-9347.
Bagrus pemecus Valenciennes in Cuvier & Valenciennes, 1840a: 456 (338 of Strasbourg deluxe edition). Type locality: Cayenne. Holotype: MNHN a-9348 (dry).
Bagrus (Bagrus) coelestinus Müller & Troschel, 1849: 7. Type locality: Guiana. Syntypes: ZMB 2970 (1), ZMB
2971 (1).

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Hexanematichthys hymenorrhinos Bleeker, 1862a: 377. Type locality: [Not stated]. Holotype: RMNH 3056; holotype illustrated in Bleeker (1864: 57, pl. 11, fig. 2; pl. 13, fig. 4), with type locality stated as Guatemala.
Netuma dubia Bleeker, 1862a: 382. Type locality: Surinama. Syntypes: RMNH 3051(1), RMNH 3052 (1); syntype
illustrated in Bleeker (1864: 63, pl. 15, fig. 2; pl. 13, fig. 5).
Distribution: Caribbean and Atlantic draining rivers and estuaries of South America from Colombia to Brazil (Marceniuk & Ferraris, 2003).
Sciades passany (Valenciennes, 1840)
Bagrus passany Valenciennes, in Cuvier & Valenciennes, 1840a: 458 (340 of Strasbourg deluxe edition). Type locality: Cayenne. Holotype: MNHN a-9399.
Distribution: Atlantic draining coastal rivers from Guyana to the mouth of the Amazon River (Marceniuk & Ferraris,2003).
Sciades platypogon (Günther, 1864)
Arius platypogon Günther, 1864: 147, unnumbered figures. Type locality: Guatemala, San Jose. Syntypes: BMNH
1864.1.26.316–319 (4).
Netuma mazatlana Gilbert, 1904: 255, pl. 25. Type locality: Mazatlán, Sinaloa, Mexico. Holotype: SU 7138.
Distribution: Pacific coast of the Americas, from Mexico to Ecuador (Marceniuk & Ferraris, 2003).
Sciades proops (Valenciennes, 1840)
Bagrus proops Valenciennes, in Cuvier & Valenciennes, 1840a: 457 (339 of Strasbourg deluxe edition). Type locality: Cayenne ou de Surinam ... Porto-Rico. Syntypes: MNHN a-9331 (1), MNHN a-9332 (1), MNHN a-9333(1), MNHN b-0245 (1), MNHN b-0618 (2).
Distribution: Caribbean and Atlantic draining rivers and estuaries of South America and some Caribbean Islands
from Colombia to Brazil (Marceniuk & Ferraris, 2003).
Sciades troschelii Gill, 1863
Sciades troschelii Gill, 1863: 171. Type locality: Western coast of Central America. Holotype: USNM (not found).
Arius Brandtii Steindachner, 1876: 571, pl. 3. Type locality: Altata, Panama. Syntypes: possibly MCZ 4964 (1),
NMW 48107 (3), NMW 48110 (1).
Distribution: Pacific coast of the Americas, from Mexico to Peru (Marceniuk & Ferraris, 2003).
Species inquirenda, Sciades
Bagrus (Sciades) emphysetus Müller & Troschel, 1849: 8. Type locality: Surinam. Holotype: ZMB 2990.
Species inquirendae, Ariidae
Silurus ascita Linnaeus, 1758: 306. Type locality: India. Syntypes: NRM 72 (4). Inclusion in the Ariidae follows Fernholm & Wheeler (1983: 222).
Pimelodus gambensis Bowdich, 1825: 234, fig. 50. Type locality: Gambia River, west Africa. No types known.
Arius nigricans Valenciennes, 1835, in Valenciennes, 1835–47: pl. 3 (fig. 3). Type locality: [No locality stated, apparently La Plata River, Montevideo, Uruguay]. Holotype: MNHN 0000-0176. Name available from plate; described in
Cuvier & Valenciennes (1840b: 83, 62 of Strasbourg deluxe edition).
Bagrus albicans Valenciennes, in Cuvier & Valenciennes, 1840a: 461 (342 of Strasbourg deluxe edition), pl. 420. Type
locality: Cayenne. Holotype: MNHN a-9350.
Bagrus arioides Valenciennes, in Cuvier & Valenciennes, 1840a: 440 (326 of Strasbourg deluxe edition). Type locality:
Bengal. Holotype (dix pouces): at MNHN.
Arius Belangerii Valenciennes, in Cuvier & Valenciennes, 1840b: 71 (53 of Strasbourg deluxe edition). Type locality:
Bombay. Holotype (6 pouces): at MNHN.
Arius granosus Valenciennes, in Cuvier & Valenciennes, 1840b: 69 (51 in Strasbourg deluxe edition). Type locality:
Pondichéry. Holotype: MNHN b-0681.
Bagrus flavescens Valenciennes, in Cuvier & Valenciennes, 1840a: 462 (342 of Strasbourg deluxe edition). Type locality:
Cayenne. Holotype: MNHN a-8566 (mounted).

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Arius molliceps Valenciennes, in Cuvier & Valenciennes, 1840b: 108 (80 of Strasbourg deluxe edition). Type locality: on
en ignore l’origine. Holotype (5 ½ pouces): at MNHN.
Arius rostratus Valenciennes, in Cuvier & Valenciennes, 1840b: 63 (47 of Strasbourg deluxe edition). Type locality: dansles environs d’Alipey. Type(s): at MNHN.
Arius sinensis Valenciennes, in Cuvier & Valenciennes, 1840b: 72 (54 of Strasbourg deluxe edition). Type locality: Touraine. Holotype (4 pouces): at MNHN.
Bagrus Temminckii Valenciennes, in Cuvier & Valenciennes, 1840a: 463 (343 of Strasbourg deluxe edition). Type locality: Cayenne [apparently in error for Suriname]. Holotype: at RMNH (apparently lost). As Bagrus temminckii incontents, p. xxii, and Bagrus temminckianus on p. 463. Günther (1864: 151) used temminckii as valid while notingthat the name was spelled temminckianus in Cuvier & Valenciennes (1840a: 463) thereby acting as first reviser. Type
locality corrected by Boeseman (1972: 297).
Bagrus trachipomus Valenciennes, in Cuvier & Valenciennes, 1840a: 442 (328 of Strasbourg deluxe edition). Type locality: [Bengal]. Holotype: MNHN a-9346 (mounted).
Arius aequibarbis Valenciennes, in Cuvier & Valenciennes, 1840b: 68 (51 of Strasbourg deluxe edition). Type locality:
Rangoon; Bengal. Syntypes (2): MNHN 0000-1200 (2), MNHN b-0682 (1).
Bagrus venaticus Richardson, 1845, in Richardson, 1844–48: 33. Type locality: North-west coast of Australia. Holotype:apparently destroyed (see Günther, 1864: 174).
Bagrus vertagus Richardson, 1845, in Richardson, 1844–48: 33. Type locality: North-west coast of Australia. Holotype:apparently destroyed (see Günther, 1864: 174).
Arius caelatoides Bleeker, 1846a: 159. Type locality: Batavia. Type(s) (size and number not stated): Whereabouts
unknown.
Arius chondropterygius Bleeker, 1846a: 162. Type locality: Batavia. Type(s) (size and number not stated): Whereabouts
unknown.
Arius chondropterygioides Bleeker, 1846a: 170. Type locality: Batavia. Type(s) (size and number not stated): Whereabouts unknown.
Arius clijpeaster Bleeker, 1846a: 161. Type locality: Batavia. Type(s) (size and number not stated): Whereabouts
unknown.
Arius clypeastroides Bleeker, 1846a: 161. Type locality: Batavia. Type(s) (size and number not stated): Whereabouts
unknown.
Arius Heckelii Bleeker, 1846a: 169. Type locality: Batavia. Type(s) (size and number not stated): Whereabouts unknown.
Arius laeviceps Bleeker, 1846a: 165. Type locality: Batavia. Type(s) (size and number not stated): Whereabouts
unknown.
Arius macruropterygius Bleeker, 1846a: 164. Type locality: Batavia. Type(s) (size and number not stated): Whereabouts
unknown.
Arius manjong Bleeker, 1846a: 163. Type locality: Batavia. Type(s) (size and number not stated): Whereabouts
unknown.
Arius microgastropterygius Bleeker, 1846a: 160. Type locality: Batavia. Type(s (size and number not stated): Whereabouts unknown.
Arius micronotacanthus Bleeker, 1846a: 162. Type locality: Batavia. Type(s) (size and number not stated): Whereabouts
unknown.
Arius viviparus Bleeker, 1846a: 171. Type locality: Batavia. Type(s) (size and number not stated): Whereabouts
unknown.
Arius angulatus Bleeker, 1846a: 170. Type locality: Batavia. Type(s) (size and number not stated): Whereabouts
unknown.
Arius pidada Bleeker, 1846a: 169. Type locality: Batavia. Syntypes (size and number not stated): possibly AMS B.7965(1), BMNH 1863.12.4.57 (1), NMV 45948 (1). Type series apparently includes at least two species. In the absenceof a lectotype, the position of this name is not clear.
Pimelodus mong Richardson, 1846: 286. Type locality: Canton, China. No types known. Based solely on illustration byReeves (reproduced in Whitehead (1970: 210, pl. 20b).
Bagrus crinalis Richardson, 1846: 282. Type locality: Canton, China. No types known. Originally as ? Bagrus crinalis.

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Based solely on illustration by Reeves, reproduced in Whitehead (1970: 210, pl. 19c).
Arius melanopterygius Bleeker, 1849: 10. Type locality: In Freto Madurae prope Kammal et Surabaya [Java]. Holotype(310 mm TL): Whereabouts unknown.
Arius borneënsis Bleeker, 1851c: 67. Type locality: Bandjarmassing. Syntypes (2, 101, 170 mm): Whereabouts
unknown, perhaps at RMNH.
Arius schlegeli Bleeker, 1863d: 146. Type locality: Amoy, China. Syntypes (2): possibly AMS B.8123 (1).
Arius andamanensis Day, 1871: 699. Type locality: Andaman Is. Possible syntypes: AMS B.7931 (1), ZSI F307 (1).
Type series is apparently mixed (Kailola, 1999: 1842, 1865) and, in the absence of a lectotype, position of this nameis not clear.
Arius argenteus Lütken, 1874b: 211. Type locality: Canton, China. Holotype: ZMUC P 29624.
Arius festae Boulenger, 1898c: 5. Type locality: Narrangal, Equateur occidental. Holotype: MZUT 1479.
Arius (Hemiarius) nudidens Weber, 1913: 538, fig. 15. Type locality: Lorentz-Fluss [New Guinea]. Syntypes (4): ZMA
111507 (1), ZMA 111508 (1), ZMA 111509 (1).
Netuma aulometopon Fowler, 1915a: 204, fig. 1. Type locality: Dutch Guiana. Holotype: ANSP 8372.
Arius satparanus Chaudhuri, 1916: 430, figs. 6–8. Type locality: Channel between Satpara and Barnikuda, Chilka Lake,
Orissa, India. Holotype: ZSI F8784/1; illustrated in Jayaram & Dhanze (1981b: fig. 1).
Remarks: Possibly a synonym of Arius tenuispinis (Jayaram & Dhanze, 1981b), or Arius argyropleurodon (Valenciennes) (Kailola, 1999).
Hexanematichthys henni Fisher & Eigenmann, in Eigenmann, 1922: 30, fig. 9. Type locality: Colimes, Rio Daule, Ecuador. Holotype: CAS 60620. Authorship based on footnote on p. 27.
Netuma hassleriana Borodin, 1934: 33, fig. b. Type locality: Panama. Holotype: MCZ 33213.
Hemipimelodus sundanensis Hardenberg, 1948: 411. Type locality: Kota Agung (Sunda Strait), Sumatra. Syntypes (2,270–280 mm): Whereabouts unknown.
Unconfirmed fossil Ariidae† Arius bartonensis Woodward, 1887: 306, fig. 3. Type locality: High Cliff, Barton Hampshire, England. Upper Eocene.
Holotype: BMNH 40280 (dorsal fin spine). Originally: Arius (?) bartonensis.
† Arius egertoni belgicus Leriche, 1905. (no additional information; name from Weiler, 1956: 187).
† Arius Bonneti Priem 1904: 44 (figs. 3–8). Type locality: Bartonien superieur de Marines. Types: dans la collection Bonnet (dorsal and pectoral fin spines); also illustrated in Priem (1908: 123, figs. 63, 64, pl. 3, figs. 5–6).
† Arius Dutemplei Leriche, 1900: 181, pl. 1, figs 13–15. Type locality: Cuis, Chavot [ Marne, Lower Eocene]. Types: atlaboratoire de Géologie de l’Université de Lille (dorsal fin spine, pectoral fin spine).
Distribution: several Paris Basin localities, lowermost Eocene (Gaudant, 1981) (from Gayet & Meunier, 2003).
† Silurus Egertoni Dixon, in Dixon & Egerton, 1850: 112, 204, pl. 11, figs. 11–13. Bracklesham Bay, Sussex, Middle
Eocene. Holotype: BMNH 25612 (pectoral arch and spine).
Remarks: Redescribed as † Arius egertoni in Woodward (1887: 304, figs. 1–2).
† Arius Fraasi Peyer, 1928: 17, pl. 6, fig. 2. Type locality: Unterste Nokkattam-Stufe, Eocän, Cairo. Holotype: at Naturalienkabinett Stuttgart.
† Arius gigas [no additional information; from Gayet & Meunier (2003)].
Distribution: Chad, Mio-Pliocene (Brunet et al., 2000).
Remarks: Gayet & Meunier (2003) placed this species in the Ariidae, but without any additional information. It is
unclear whether the use of this name in Brunet et al. (2000) and Gayet & Meunier (2003) was intended to identify fossils with the Recent species Arius gigas Boulenger or this name was proposed independently of, andtherefore a homonym of, that name.
† Arius heward-belli White, 1926: 56, pl. 13, figs. 4–12. Type locality: Ameki, Nigeria, Eocene. Holotype: National Collection [Nigeria] or BMNH (pectoral spine tip).
† Arius kitsoni White, 1926: 54, pl. 13, figs. 1–2. Type locality: Ameki, Nigeria, Eocene. Holotype: National Collection[Nigeria] or BMNH (proximal half of right pectoral fin spine).
† Arius kutchensis Raghavendra Rao, 1956: 181, figs. 1–2, pl. 28. Type locality: Gypseous shale beds about two miles

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west of Nareda, Western Kutch, India; Lower Eocene. Holotype: (skull) Geological Survey of India, Calcutta 17539.
Remarks: See Sahni & Mishra (1975: 11, pl. 2, figs. 2–4) for emended diagnosis.
† Arius nucleus Weinfurter, 1952. Type locality: [Austria, Miocene]. [No other information]† Arius russi White, 1926: 58, pl. 13, fig. 3. Type locality: Oshosun, Nigeria, Eocene. Holotype: National Collection[Nigeria] or BMNH (proximal end of pectoral fin spine).
† Tachysurus landanensis [No other information, taken from Gayet & Meunier (2003)].
Distribution: Zaire, Eocene (Gayet & Meunier, 2003).
Remarks: Placement in Ariidae supported by Gayet & Meunier (2003).

ASPREDINIDAE Adams, 1854
Aspredinidae Adams, in Adams et al., 1854: 107. Type genus: Aspredo Scopoli, 1777.
Bunocephalini Bleeker, 1858b: xi, 328. Type genus: Bunocephalus Kner, 1855.
Platystacinae Eigenmann & Eigenmann, 1890: 9, 13, 20. Type genus: Platystacus Bloch, 1794.
Hoplomizoninae [sic Hoplomyzontinae] Fernández-Yépez, 1950b:113. Type genus: Hoplomyzon Myers, 1942.
Taxonomy summary: Gill (1891a); Friel (2003); Myers (1960); Mees (1987b, Aspredininae); Mees (1988b Bunocephalinae).
Keys: Mees (1988b, Aspredininae).
Revision: Stewart (1985b, Hoplomyzontinae).
Phylogeny: Friel (1994); Diogo et al. (2001).
12 genera, 36 species; no named fossil taxa.
ACANTHOBUNOCEPHALUS Friel, 1995
Acanthobunocephalus Friel, 1995: 90. Type species: Acanthobunocephalus nicoi Friel, 1995. Type by original designation. Gender: Masculine.
Acanthobunocephalus nicoi Friel, 1995
Acanthobunocephalus nicoi Friel, 1995: 90, fig. 1. Type locality: Río Sipapo, 200 m from Salto Remo, 4º34'N,
67º18'W, Venezuela. Holotype: MCNG 29000.
Distribution: Upper Orinoco River basin of Venezuela, including Sipapo, Ramoni and Casiquiare rivers (Friel,2003).
AMARALIA Fowler, 1954
Amaralia Fowler, 1954: 40. Type species: Bunocephalus hypsiurus Kner, 1855. Type by original designation. Gender: Feminine.
Amaralia hypsiura (Kner, 1855)
Bunocephalus hypsiurus Kner, 1855a: 98, pl. 1 (fig. 1). Type locality: Rio Branco [Brazil]. Syntypes: NMW 47626(2).
Distribution: Throughout the Amazon River basin (Friel, 2003).
ASPREDINICHTHYS Bleeker, 1858
Aspredinichthys Bleeker, 1858b: 328, 329. Type species: Aspredo tibicen Valenciennes, 1840. Type by monotypy.
Gender: Masculine.
Chamaigenes Eigenmann, 1910: 380. Type species: Aspredo filamentosus Valenciennes, 1840. Type by original designation. Gender: Masculine.
Revison: Mees (1987b).
Aspredinichthys filamentosus (Valenciennes, 1840)
Aspredo filamentosus Valenciennes, in Cuvier & Valenciennes, 1840b: 437 (324 of Strasbourg deluxe edition), pl.
450. Type locality: Cayenne. Syntypes: MNHN 0000-4401 (4), MNHN b-0594 (2).
Distribution: Lower portions of rivers and in coastal waters of northern South America from Venezuela to northern
Brazil (Friel, 2003).

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Aspredinichthys tibicen (Valenciennes, 1840)
Aspredo tibicen Valenciennes, in Cuvier & Valenciennes, 1840b: 438 (325 of Strasbourg deluxe edition). Type locality: Surinam. Holotype: RMNH 3111.
Distribution: Lower portions of rivers and in coastal waters of northern South America from Venezuela to northern
Brazil (Friel, 2003).
ASPREDO Scopoli, 1777
Aspredo Scopoli, 1777: 453. Type species: Silurus aspredo Linnaeus, 1758. Type by absolute tautonymy. Gender:
Masculine.
Aspredo aspredo (Linnaeus, 1758)
Silurus aspredo Linnaeus, 1758: 304. Type locality: in Americae fluviis. Syntype: ZMUU Linnaean Coll. 48 (1);also based on descriptions in Gronovius (1754: 8), which apparently is based on the same Linnaean specimen,
and on Klein (1775).
Platystacus laevis Bloch, 1794: 58. Type locality: [No locality stated]. Syntypes: ZMB 3153 (3), and several literature sources, including Silurus aspredo Linnaeus.
Aspredo sicuephorus Valenciennes, in Cuvier & Valenciennes, 1840b: 439 (326 of Strasbourg deluxe edition). Type
locality: La Mana [French Guiana]. Holotype: MNHN 0000-6362.
Aspredo batrachus Gronow, in Gray, 1854: 137. Type locality: fluminibus Americes Meridionalis. Syntype or holotype: ZMUU Linnaean Coll. 48 (1); inasmuch as Gronovius (1754: 8) based the description on an earlier Linnaean description.
Aspredo sicyephorus Günther, 1864: 269. Type locality: Mana [French Guiana]. Holotype: MNHN 0000-6362.
Unjustified emendation of Aspredo sicuephorus Valenciennes, 1840.
Distribution: Lower portions of rivers from Venezuela to northern Brazil (Friel, 2003).
BUNOCEPHALUS Kner, 1855
Aspredo Swainson, 1838: 332. Type species: Aspredo gronovii Swainson, 1838. Type by monotypy. Gender: Masculine. Preoccupied by Aspredo Scopoli, 1777; apparently not replaced.
Bunocephalus Kner, 1855a: 95. Type species: Platystacus verrucosus Bloch, 1794. Type by subsequent designation
by Bleeker (1862–63: 19). Gender: Masculine.
Platystacus Bleeker, 1858b: 329. Type species: Bunocephalichthys verrucosus Bloch, 1794. Type by subsequent
designation by Jordan (1919a: 279). Gender: Masculine.
Dysichthys Cope, 1874b: 133. Type species: Dysichthys coracoideus Cope, 1874. Type by monotypy. Gender: Masculine.
Agmus Eigenmann, 1910: 379. Type species: Bunocephalus scabriceps Eigenmann & Eigenmann, 1889. Type byoriginal designation. Gender: Masculine.
Remarks: Ferraris (1991) clarified type species of Bunocephalus, but see Mees (1997) for alternate opinion. Many of
the species listed here were placed in Dysichthys by Mees (1988b, 1989), but Dysichthys was placed in the synonymy of Bunocephalus in Friel (2003).
Review: Mees (1989, as Dysichthys, in part).
Bunocephalus aleuropsis Cope, 1870
Bunocephalus aleuropsis Cope, 1870b: 568. Type locality: Pebas, Eastern Ecuador [now Peru]. Syntypes: ANSP
8286–8288 (3); one syntype illustrated in Mees (1989: 206, fig. 4).
Bunocephalus melas Cope, 1874b: 132. Type locality: Nauta. Holotype: ANSP 21235 (in pieces).
Distribution: Amazon and Orinoco River basins (Friel, 2003).
Bunocephalus amaurus Eigenmann, 1912
Bunocephalus amaurus Eigenmann, 1912b: 126, fig. 24; pl. 2 (fig. 2). Type locality: Konawaruk, British Guiana.
Holotype: FMNH 53121.
Bunocephalus amaurus aloikae Hoedeman, 1961: 130. Type locality: Litany R., Aloiké village, French Guiana.
Holotype: ZMA 102229.
Bunocephalus amaurus sipaliwini Hoedeman, 1961: 130, fig. 1. Type locality: Sipaliwini, Suriname, 20 km from58 · Zootaxa 1418 © 2007 Magnolia Press

FERRARIS

frontier with Brazil. Holotype: ZMA 102228.
Distribution: Coastal rivers of northern South America between Orinoco to Amazon mouths (Friel, 2003).
Remarks: Redescribed in Mees (1989: 216) as a subspecies of Dysichthys coracoideus.
Bunocephalus amazonicus (Mees, 1989)
Dysichthys amazonicus Mees, 1989: 241, fig. 21. Type locality: Bolivia: creek near Todos Santos, upper course of R.
Mamoré, Cochabamba. Holotype: ZMA 109246 (37 mm SL specimen).
Distribution: Upper and middle Amazon River basin (Friel, 2003).
Bunocephalus bifidus Eigenmann, 1942
Bunocephalus bifidus Eigenmann, in Eigenmann & Allen, 1942: 86, pl. 2 (fig. 2). Type locality: creek, Yurimaguas[Huallaga River, Peru]. Holotype: CAS 35105.
Distribution: Amazon River basin of Bolivia, Brazil, Ecuador, and Peru (Friel, 2003).
Remarks: Name appeared several times (Pearson, 1924: 9; Pearson 1937: 111; Fowler, 1940c: 94) as a nomen nudum
prior to being made available in Eigenmann & Allen (1942).
Bunocephalus chamaizelus Eigenmann, 1912
Bunocephalus chamaizelus Eigenmann, 1912b: 127, fig. 24; pl. 2 (fig. 1). Type locality: Erukin, British Guiana.
Holotype: FMNH 53122.
Distribution: Essequibo River basin (Friel, 2003).
Bunocephalus colombianus Eigenmann, 1912
Bunocephalus colombianus Eigenmann, 1912a: 10. Type locality: Raspadura [Colombia]. Holotype: FMNH 56038.
Distribution: Atrato and Magdalena River basins (Friel, 2003); also Cauca, San Jorge, Sinu and Patía basins, Colombia (Maldonado-Ocampo et al., 2005).
Bunocephalus coracoideus (Cope, 1874)
Dysichthys coracoideus Cope, 1874b: 133. Type locality: Nauta [Peru]. Syntypes: ANSP 21212–21215 (4).
Bunocephalus bicolor Steindachner, 1882b: 176. Type locality: Aus dem Huallaga [Peru]. Holotype: MTD F345.
Species illustrated and described in more detail in Steindachner (1882c: 8, pl. 2, figs. 1, 1a, 1b).
Bunocephalus haggini Eigenmann & Allen, 1942: 86, pl. 2 (fig. 1). Type locality: Amazon at Iquitos [Peru]. Holotype: CAS 35107.
Distribution: Amazon River basin of Bolivia, Brazil, and Peru (Friel, 2003).
Bunocephalus doriae Boulenger, 1902
Bunocephalus doriae Boulenger, 1902b: 286. Type locality: Villa Rica, Paraguay; and Posadas, Río Parana [Argentina]. Syntypes: BMNH 1902.2.10.22–23 (2), MSNG 14274 (1), MSNG 14275 (3).
Bunocephalus retropinnis Eigenmann, in Eigenmann & Allen, 1942: 85, pl. 2 (fig. 3). Type locality: Cacequi, Brazil.
Holotype: CAS 35246.
Distribution: Paraguay-Paraná and Uruguay River basins, of Argentina, Brazil, Paraguay and Uruguay (Friel, 2003).
Bunocephalus iheringii Boulenger, 1891
Bunocephalus iheringii Boulenger, 1891: 235, pl. 26 (fig. 2). Type locality: Not stated [apparently from Rio Grande
do Sul, Brazil]. Syntypes: BMNH 1891.3.16.77–78 (2).
Bunocephalus salathei Myers, 1927: 125. Type locality: Brazil: Morro Ajudo, about 100 km from Rio de Janeiro.
Holotype: MCZ 31583.
Bunocephalus minutus Güntert, 1942: 28, figs. 1–2. Type locality: Paraguay, Dept. Villetta. Syntypes: NMBA 5296–
98 (3), NMBA 5299–5300 (2).
Bunocephalus carvalhoi Miranda Ribeiro, 1944a: 1, figure on p. 2. Type locality: Magé, Estado do Rio de Janeiro[Brazil]. Holotype: MNRJ 1060.
Distribution: Atlantic, Paraguay and Uruguay River basins of Brazil and Paraguay (Friel, 2003).
Bunocephalus knerii Steindachner, 1882
Bunocephalus Knerii Steindachner, 1882b: 176. Type locality: Canelos [Ecuador]. Syntype: NMW 47628. Species
illustrated and described in more detail in Steindachner (1882c: 9, pl. 2, figs. 2, 2a, 2b).
Distribution: Upper Amazon River basin of Colombia, Ecuador, and Peru (Friel, 2003).
Bunocephalus larai Ihering, 1930
Bunocephalus larai Ihering, 1930: 101, pl. 13 (figs. 3–3a). Type locality: Rio Piracicaba, junto ao Salto; ao 20 klm

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abaixo (ilha das Flechas) [São Paulo, Brazil]. Syntypes: Whereabouts unknown.
Distribution: Paraná River basin of Brazil (Friel, 2003).
Bunocephalus quadriradiatus (Mees, 1989)
Dysichthys quadriradiatus Mees, 1989: 244, figs. 23, 24. Type locality: Peru: Samiria, Chinguito [= Cocha Shinguita or Shirguita]. Holotype: MHNG 2157.21.
Distribution: Upper Amazon River basin of Peru (Friel, 2003).
Bunocephalus rugosus Eigenmann & Kennedy, 1903
Bunocephalus rugosus Eigenmann & Kennedy, 1903: 498. Type locality: Laguna near Arroyo Chagalalina [Paraguay]. Holotype: IU 9819 (whereabouts unknown).
Dysichthys australe Eigenmann & Ward, in Eigenmann, McAtee & Ward, 1907: 113, pl. 31. Type locality: Corumba[Paraguay]. Holotype: CAS 35240.
Distribution: Paraguay-Paraná River basin of Brazil and Paraguay (Friel, 2003).
Bunocephalus verrucosus (Walbaum, 1792)
Silurus verrucosus Walbaum, 1792: 574. Type locality: [No locality stated]. No types known. Based on Gronovius(1754, pl. 5, fig. 3).
Platystacus verrucosus Bloch, 1794: 63, pl. 373 (fig. 3). Type locality: Wahrscheinlich gerhört er in Surinam zu
Hause [= Suriname]. On Gronovius (1754: 5, pl. 5, fig. 3; and 1763: 102, no. 325).
Aspredo gronovii Swainson, 1838: 332, fig. 80. Type locality: [No locality stated]. No types known.
Aspredo verrucosa Gronow, in Gray, 1854: 137. Type locality: America Meridionali. No types known; based on
Gronovius (1763: no. 325, pl. 5, fig. 3.).
Bunocephalichthys gronovii Bleeker, 1858b: 329. Type locality: Am. Merid. No types known; based on Gronovius(1754: 153, pl. 5, fig. 3).
Bunocephalus scabriceps Eigenmann & Eigenmann, 1889b: 49. Type locality: Jutahy [Brazil]. Lectotype: MCZ
7967; lectotype illustrated and designated in figure caption of Eigenmann (1912: pl. 4) as 42 mm specimen.
Agmus lyriformis Eigenmann, 1912b: 128, fig. 25; pl. 3. Type locality: Gluck Island [Rupununi River], British Guiana. Holotype: FMNH 53120.
Distribution: Rivers of the Guianas and throughout the Amazon River basin (Friel, 2003).
DUPOUYICHTHYS Schultz, 1944
Dupouyichthys Schultz, 1944c: 244. Type species: Dupouyichthys sapito Schultz, 1944. Type by original designation. Gender: Masculine.
Dupouyichthys sapito Schultz, 1944
Dupouyichthys sapito Schultz, 1944c: 245, pl. 4 (fig. d); fig. 4b. Type locality: Río Motatán, at the bridge 22 km
north of Motatán, Venezuela. Holotype: USNM 121072.
Distribution: Magdalena River and Lake Maracaibo basins (Friel, 2003).
Remarks: Redescribed in Mojica et al. (2000).
ERNSTICHTHYS Fernández-Yépez, 1953
Ernstichthys Fernández-Yépez, 1953: 3, 4. Type species: Ernstichthys anduzei Fernández-Yépez, 1953. Type byoriginal designation. Gender: Masculine.
Ernstichthys anduzei Fernández-Yépez, 1953
Ernstichthys anduzei Fernández-Yépez, 1953: 5, fig. 1. Type locality: Río Salinas, northeast of El Baúl, Estado
Cojedes, Venezuela. Holotype: MHNLS 28.
Distribution: Orinoco River basin of Venezuela (Friel, 2003).
Ernstichthys intonsus Stewart, 1985
Ernstichthys intonsus Stewart, 1985b: 16, figs. 1f, 3, 9. Type locality: Río Napo at Añangu, Napo Prov, 0º30.8'S,76º24.0'W, Ecuador. Holotype: FMNH 94603.
Distribution: Napo River basin of Ecuador (Friel, 2003).
Ernstichthys megistus (Orcés, 1961)
Hoplomyzon megistus Orcés, 1961: 3, figs. 1–2. Type locality: Small river off of the Río Chicherota, Río Bobonaza

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FERRARISsystem, Prov. Pastaza, e. Ecuador. Holotype: Author’s personal coll. 4305 (current whereabouts unknown).
Distribution: Bobonaza and Marañon River basins of Ecuador and Peru (Friel, 2003).
HOPLOMYZON Myers, 1942
Hoplomyzon Myers, 1942: 94. Type species: Hoplomyzon atrizona Myers, 1942. Type by original designation. Gender: Masculine.
Hoplomyzon atrizona Myers, 1942
Hoplomyzon atrizona Myers, 1942: 95, fig. 3. Type locality: Brook trib. to Río Zulia, at Estacion Tachira, 60 km
north of San Cristobal, Venezuela, elev. ca. 150 m. Holotype: SU 36494.
Hoplomyzon atrizona petroleus Schultz, 1944c: 248, pl. 4 (fig. C); fig. 4a. Type locality: Río Motatán, 4 km above
Motatán, Maracaibo basin, Venezuela. Holotype: USNM 121070.
Distribution: Lake Maracaibo basin (Friel, 2003).
Hoplomyzon papillatus Stewart, 1985
Hoplomyzon papillatus Stewart, 1985b: 8, figs. 1b, 2b, 4b, 5. Type locality: Río Aguarico, 1 km upstream from confluence with Río Shushufindi, Napo Prov., 0º17'S, 76º24.5'W, Ecuador. Holotype: FMNH 94908.
Distribution: Napo and Portuguesa River basins of Ecuador and Venezuela, respectively (Friel, 2003).
Hoplomyzon sexpapilostoma Taphorn & Marrero, 1990
Hoplomyzon sexpapilostoma Taphorn & Marrero, 1990: 4, figs. 1–4. Type locality: Río Masparro at site of MasparroDam, Barinas, Venezuela, 8º50'40"N, 70º06'00"W. Holotype: MCNG 18669.
Distribution: Orinoco River basin of Venezuela (Friel, 2003).
MICROMYZON Friel & Lundberg, 1996
Micromyzon Friel & Lundberg, 1996: 642. Type species: Micromyzon akamai Friel & Lundberg, 1996. Type by original designation. Gender: Masculine.
Micromyzon akamai Friel & Lundberg, 1996
Micromyzon akamai Friel & Lundberg, 1996: 643, fig. 1. Type locality: Rio Tocantins, above confluence with Rio
Pará, 2º02'S, 49º17'W, Pará, Brazil, 10–14 m. Holotype: MZUSP 48550.
Distribution: Lower Amazon River basin (Friel, 2003).
PLATYSTACUS Bloch, 1794
Platystacus Bloch, 1794: 52. Type species: Platystacus cotylephorus Bloch, 1794. Type by subsequent designation
by Bleeker (1862–63: 19). Gender: Masculine.
Platysomatos Bloch, 1797: 115. Type species: Platystacus cotylephorus Bloch, 1794. Type by being a replacement
name. Unneeded new name for Platystacus Bloch, 1794. Gender: Masculine. See Harry (1953) for commentson this name.
Cotylephorus Swainson, 1838: 354. Type species: Cotylephorus blochii Swainson, 1838. Type by monotypy. Gender: Masculine.
Platystacus cotylephorus Bloch, 1794
Platystacus cotylephorus Bloch, 1794: 54, pl. 372. Type locality: Ostindischen Gewässer [in error; certainly fromSouth America]. Syntypes: ZMB 3157 (2), and several specimens associated with literature sources.
Silurus hexadactylus La Cepède, 1803: 58, 82. Type locality: [No locality stated]. Holotype: MNHN b-0029.
Cotylephorus Blochii Swainson, 1838: 354, fig. 90. Type locality: Ostindischen Gewässer [in error; certainly fromSouth America]. Syntypes: ZMB 3157. Unneeded new name for Platystacus cotylephorus Bloch, 1794, apparently to avoid tautonymy.
Aspredo sexcirrhis Valenciennes, in Cuvier & Valenciennes, 1840b: 441 (326 of Strasbourg deluxe edition). Type
locality: Surinam. Syntypes: MNHN a-9313 (1), MNHN b-0029 (1).
Aspredo spectrum Gronow, in Gray, 1854: 137. Type locality: in fluminibus Americes Meridionalis. Types: Whereabouts unknown; based on Gronovius (1763: no. 324), and Seba (1734–65: pl. 29, fig. 9).
Platystacus nematophorus Bleeker, 1862a: 371. Type locality: Surinama. Holotype: RMNH 3105; holotype illustrated in Bleeker (1864: 96, pl. 1, fig. 1).

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Distribution: Coastal waters and lower portions of rivers of northern South America, from Venezuela to northern
Brazil (Friel, 2003).
PTEROBUNOCEPHALUS Fowler, 1943
Pterobunocephalus Fowler, 1943c: 1. Type species: Bunocephalus albifasciatus Fowler, 1943. Type by original designation. Gender: Masculine.
Petacara Böhlke, 1959: 2. Type species: Bunocephalus dolichurus Delsman, 1941. Type by original designation.
Gender: Neuter.
Pterobunocephalus depressus (Haseman, 1911)
Bunocephalus depressus Haseman, 1911a: 319, pl. 48 (fig. 1), pl. 49. Type locality: Río Machupo near San Joaquín,
Bolivia. Holotype: FMNH 54338.
Bunocephalus albifasciatus Fowler, 1943c: 2, fig. 1. Type locality: Todos Santos, Bolivia. Holotype: ANSP 69193.
Distribution: Amazon, Orinoco and Paraguay River basins (Friel, 2003).
Pterobunocephalus dolichurus (Delsman, 1941)
Bunocephalus dolichurus Delsman, 1941: 77, fig. 12. Type locality: Trombetas-river, near Obidos [Brazil]. Holotype: IRSNB 359.
Distribution: Amazon River basin of Brazil and Peru (Friel, 2003).
XYLIPHIUS Eigenmann, 1912
Xyliphius Eigenmann, 1912a: 10. Type species: Xyliphius magdalenae Eigenmann, 1912. Type by monotypy. Gender: Masculine. Spelled two ways in original description: Xyliphius and Xiliphius, first reviser not yet identified,but may not have been formalized until noted by Eschmeyer & Bailey, in Eschmeyer (1990).
Xyliphius barbatus Alonso de Arámburu & Arámburu, 1962
Xyliphius barbatus Alonso de Arámburu & Arámburu, 1962: 219, figs. 1–2. Type locality: río Paraná en Rosario,
provincia de Santa Fe, Argentina. Holotype: MLP 6798.
Xyliphius labrosus Risso & Risso, 1964a: 11. Apparently an unneeded new name for Xyliphius barbatus Alonso de
Arámburu & Arámburu, 1962.
Distribution: Paraguay-Paraná River basin, Argentina (Friel, 2003).
Xyliphius kryptos Taphorn & Lilystrom, 1983
Xiliphius kryptos Taphorn & Lilystrom, 1983: 43, fig. 1. Type locality: Cerca del Puente de la carretera 6 sobre el
Río Aricuaisá, Estado Zulia, Venezuela. Holotype: MCNG 1224.
Distribution: Lake Maracaibo basin (Friel, 2003).
Xyliphius lepturus Orcés, 1962
Xyliphius lepturus Orcés, 1962: 50, figs. 1–2. Type locality: Cerca de Desembocadura del rio Pucayacu en el Bobonaza [Ecuador]. Holotype: Author’s personal collection 1307 (current whereabouts unknown).
Distribution: Upper Amazon and Orinoco River basins of Colombia, Ecuador and Venezuela (Friel, 2003).
Xyliphius lombarderoi Risso & Risso, 1964
Xyliphius lombarderoi Risso & Risso, 1964a: 12, pl. 1 (figs. 1–2). Type locality: el Riacho Barranqueras [Paraná,
Argentina]. Holotype: Authors’ personal collection (current whereabouts unknown).
Distribution: Paraná River basin, Argentina (Friel, 2003).
Xyliphius magdalenae Eigenmann, 1912
Xyliphius magdalenae Eigenmann, 1912a: 10. Type locality: Girardot, Colombia. Holotype: FMNH 56039; holotype
illustrated in Eigenmann (1922: 27, pl. 1, fig. 1).
Distribution: Magdalena River basin, Colombia (Friel, 2003; Maldonado-Ocampo et al., 2005).
Xyliphius melanopterus Orcés, 1962
Xyliphius melanopterus Orcés, 1962: 52, figs. 3–4. Type locality: el bajo Pucayacu no lejos de su desembocadura enel Bobonaza [Ecuador]. Holotype: Author’s personal coll. 2021 (current whereabouts unknown).
Distribution: Upper Amazon and Orinoco River basins (Friel, 2003).

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FERRARISASTROBLEPIDAE Bleeker, 1862
Argeini Bleeker, 1862 (in Bleeker, 1862–63): 15. Type genus: Arges Valenciennes, 1840.
Astroblepiformes Bleeker, 1862 (in Bleeker, 1862–63): 15. Type genus: Astroblepus Humboldt, 1805.
Cyclopidae Eigenmann, 1910: 416. Type genus: Cyclopium Swainson, 1838.
Remarks: The name Astroblepiformes takes precedence over the simultaneously published Argeini, because it was proposed at a higher taxonomic level (subfamily vs. stirps).
Taxonomic summary: Schaefer (2003b).
1 genus, 54 species; no named fossil taxa.
ASTROBLEPUS Humboldt, 1805
Astroblepus Humboldt, 1805a: 19. Type species: Astroblepus grixalvii Humboldt, 1805. Type by monotypy. Gender:
Masculine.
Cyclopium Swainson, 1838: 355. Type species: Cyclopium humboldtii Swainson, 1839. Type by subsequent monotypy. Gender: Neuter. Genus first proposed without species; species first added in Swainson (1839: 305).
Arges Valenciennes, in Cuvier & Valenciennes, 1840b: 333 (247 in Strasbourg deluxe edition). Type species: Arges
sabalo Valenciennes, 1840. Type by subsequent designation by Bleeker (1862–63: 15). Gender: Masculine.
Brontes Valenciennes, in Cuvier & Valenciennes, 1840b: 341 (254 in Strasbourg deluxe edition). Type species:
Brontes prenadilla Valenciennes, 1840. Type by monotypy. Gender: Masculine. Preoccupied by Brontes Fabricius, 1801, in Coleoptera; replaced by Strephon Gistel, 1848.
Strephon Gistel, 1848: viii. Type species: Brontes prenadilla Valenciennes, 1840. Type by being a replacement
name. Gender: Masculine. Replacement for Brontes Valenciennes, 1840, preoccupied by Brontes Fabricius,
1801, in Coleoptera.
Stygogenes Günther, 1864: 223. Type species: Cyclopium humboldtii Swainson, 1839. Type by subsequent designation of Jordan (1919b: 332). Gender: Masculine.
Astroblepus boulengeri (Regan, 1904)
Arges boulengeri Regan, 1904b: 310. Type locality: Canelos, Ecuador. Syntypes: BMNH 1890.6.13.12–13.
Distribution: Upper Amazon River basin of Ecuador (Schaefer, 2003b).
Astroblepus brachycephalus (Günther, 1859)
Arges brachycephalus Günther, 1859: 92. Type locality: Andes of western Ecuador. Syntypes: BMNH
1858.7.25.37–40 (4).
Distribution: Pacific draining rivers of Ecuador (Schaefer, 2003b).
Astroblepus caquetae Fowler, 1943
Astroblepus caquetae Fowler, 1943a: 246, fig. 23–25. Type locality: Florencia, Río Ortueguasa, Colombia. Holotype: ANSP 70506.
Distribution: Orteguaza River, upper Amazon River basin of Colombian Andes (Schaefer, 2003b).
Astroblepus chapmani (Eigenmann, 1912)
Cyclopium chapmani Eigenmann, 1912a: 13. Type locality: Boquia, Colombia. Holotype: FMNH 56071; holotype
illustrated in Eigenmann (1922: 51, pl. 6, fig.2).
Distribution: Andean streams of Cauca, Magdalena and San Juan River basins, Colombia (Maldonado-Ocampo etal., 2005).
Astroblepus chimborazoi (Fowler, 1915)
Cyclopium chimborazoi Fowler, 1915a: 241, fig. 11. Type locality: Junction of Chanchan and Chiguancay rivers.
Province of Chimborazo, Ecuador. Holotype: ANSP 43523.
Distribution: Pacific draining rivers of Ecuador (Schaefer, 2003b).
Astroblepus chotae (Regan, 1904)
Arges chotae Regan, 1904b: 313, pl. 21 (fig. 5). Type locality: Chota Valley, northern Ecuador. Holotype: BMNH
1898.11.4.22.
Distribution: Pacific draining rivers of Colombia, Ecuador, and Peru; Magdalena and Orinoco River basins(Schaefer, 2003b).

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Astroblepus cirratus (Regan, 1912)
Arges cirratus Regan, 1912b: 670. Type locality: Río Condoto, Río San Juan, S.W. Colombia. Holotype: BMNH
1912.3.2.7.
Distribution: San Juan River basin, Colombia (Maldonado-Ocampo et al., 2005).
Astroblepus cyclopus (Humboldt, 1805)
Pimelodus cyclopum Humboldt, 1805b: 24, pl. 6. Type locality: Subterranean waters in Andes of Quito (basin of Río
Esmeraldas), Ecuador. Type(s): Whereabouts unknown.
Cyclopium Humboldtii Swainson, 1839: 305. Unneeded new name for Pimelodus cyclopum Humboldt, 1805 (butlisted as Pimelodus cyclopium).
Stygogenes humboldtii Günther, 1864: 223. Type locality: Azores [in error]. Syntypes (4): BMNH 1855.6.26.14.
Distribution: Esmeraldas River and other Pacific coastal drainages of Ecuador (Schaefer, 2003b); Andean streams ofColombia (Maldonado-Ocampo et al., 2005).
Astroblepus eigenmanni (Regan, 1904)
Arges eigenmanni Regan, 1904b: 312, pl. 21 (fig. 3). Type locality: Cayambe and Machachi, Ecuador. Syntypes:
BMNH 1892.5.14.1–8 (8).
Distribution: Pacific coastal drainages of Ecuador (Schaefer, 2003b).
Astroblepus festae (Boulenger, 1898)
Arges festae Boulenger, 1898c: 9. Type locality: Vallée de Gualaquiza, Equateur Oriental. Syntypes: BMNH
1898.11.4.23–29; MZUT 1486, MZUT 1497.
Distribution: South America: Upper Marañon River of southern Ecuador (Schaefer, 2003b).
Astroblepus fissidens (Regan, 1904)
Arges fissidens Regan, 1904b: 316. Type locality: Andes of Ecuador. Syntypes: BMNH 1887.1.7.2, BMNH
1889.2.1.4397.
Distribution: Andes of Ecuador (Schaefer, 2003b).
Astroblepus formosus Fowler, 1945
Astroblepus formosus Fowler, 1945c: 10, fig. 1–2. Type locality: Acobamba, near Tarma, at 2900 meters elevation,
Río Ucayali basin, Peru, Holotype: ANSP 71647.
Distribution: Upper Ucayali River basin, Peru (Schaefer, 2003b).
Astroblepus frenatus Eigenmann, 1918
Astroblepus frenatus Eigenmann, 1918b: 676. Type locality: Quebrada de San Joaquín, Santander, Colombia. Holotype: FMNH 58384.
Distribution: Streams of Magdalena, Catatumbo and Orinoco river basins, Colombia (Maldonado-Ocampo et al.,2005).
Astroblepus grixalvii Humboldt, 1805
Astroblepus grixalvii Humboldt, 1805a: 19, pl. 7. Type locality: Río Palacé, near Popayan, Colombia. Type(s):Whereabouts unknown.
Distribution: Cauca and Magdalena River basins, Colombia (Maldonado-Ocampo et al., 2005).
Astroblepus guentheri (Boulenger, 1887)
Stygogenes guentheri Boulenger, 1887b: 348. Type locality: New Granada. Syntypes: BMNH 1880.2.26.18–25 (8).
Distribution: Andean streams of the Cauca and Magdalena River basins, Colombia (Maldonado-Ocampo et al.,2005).
Astroblepus heterodon (Regan, 1908)
Arges heterodon Regan, 1908f: 799, pl. 49 (fig. 2, 2a, b). Type locality: Jimenez, western Colombia. Holotype:
BMNH 1908.5.29.80.
Distribution: Pacific draining rivers of Colombia (Schaefer, 2003b).
Astroblepus homodon (Regan, 1904)
Arges homodon Regan, 1904b: 309, pl. 21 (fig. 1). Type locality: Villeta, Colombia, 3500 ft. Holotype: BMNH
1902.5.15.27.
Distribution: Andean streams of the Cauca and Magdalena River basins, Colombia (Maldonado-Ocampo et al.,2005).

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Astroblepus jurubidae Fowler, 1944
Astroblepus jurubidae Fowler, 1944b: 237, fig. 10–13. Type locality: clear brook, tributary to Río Jurubidá [northwestern Colombia]. Holotype: ANSP 71431.
Distribution: Pacific draining rivers of Colombia (Schaefer, 2003b).
Astroblepus labialis Pearson, 1937
Astroblepus labialis Pearson, 1937: 95, pl. 13 (fig. 4). Type locality: Balsas, Peru. Syntypes: CAS 64617 (7).
Distribution: Marañon River basin, Peru (Schaefer, 2003b).
Astroblepus latidens Eigenmann, 1918
Astroblepus latidens Eigenmann, 1918b: 674. Type locality: Piperel, Colombia. Holotype: FMNH 58366; holotype
illustrated in Eigenmann (1922: 228, pl. 6, fig. 3).
Distribution: Upper Meta River basin, Colombia (Schaefer, 2003b); and Suarez River basin, Colombia (MaldonadoOcampo et al., 2005).
Astroblepus longiceps Pearson, 1924
Astroblepus longiceps Pearson, 1924: 15, pl. 2 (fig. 5). Type locality: Río Colorado, Lower Bopi, Bolivia. Holotype:
CAS 64618.
Distribution: Upper Madeira River basin of Bolivia and Peru (Schaefer, 2003b).
Astroblepus longifilis (Steindachner, 1882)
Arges longifilis Steindachner, 1882b: 177. Type locality: Río Huambo und de Tortora [Peru]. Syntypes (4): at NMW.
Species described in more detail and illustrated in Steindachner (1882c: 19, pl. 5, figs. 3, 3a, 3b).
Distribution: Upper Huallaga River basin, Peru (Schaefer, 2003b); Magdalena and Cauaca River basins, Colombia(Maldonado-Ocampo et al., 2005).
Astroblepus mancoi Eigenmann, 1928
Astroblepus mancoi Eigenmann, in Myers, 1928: 85. Type locality: Río Comberciato, Urubamba, Peru, 1800 feet.
Syntypes: CAS 64620 (5), MCZ 30969 (11), MCZ 30970 (3).
Distribution: Upper Ucayali River basin, Peru (Schaefer, 2003b).
Astroblepus mariae (Fowler, 1919)
Cyclopium mariae Fowler, 1919a: 125, pl. 8. Type locality: Brook near the small village of Choachi, 25 kilometers
east of Bogota and at 1800 m. of altitude, Colombia. Holotype: ANSP 49368.
Distribution: Upper Meta River basin, Colombia (Schaefer, 2003b).
Astroblepus marmoratus (Regan, 1904)
Arges marmoratus Regan, 1904b: 314, pl. 21 (fig. 6). Type locality: St. Augustin, Andes of Colombia, 5000 ft. Syntypes: BMNH 1899.8.21.10–11 (2).
Distribution: Upper Orinoco River basin, Colombia (Schaefer, 2003b).
Astroblepus micrescens Eigenmann, 1918
Astroblepus grixalvii micrescens Eigenmann, 1918b: 677. Type locality: Quebrada de Agua Larga, western slope ofthe eastern Andes, north of Bogotá, Colombia. Holotype: FMNH 58376; holotype illustrated in Eigenmann(1922: 54, pl. 7, fig. 4).
Distribution: Upper Orinoco River basin (Schaefer, 2003b); widely distributed in Colombian Andes (MaldonadoOcampo et al., 2005).
Astroblepus mindoensis (Regan, 1916)
Cyclopium mindoense Regan, 1916: 80. Type locality: Mindo, western Ecuador. Holotype: BMNH 1916.8.1.1.
Distribution: Pacific coastal drainage of Ecuador (Schaefer, 2003b).
Astroblepus nicefori Myers, 1932
Astroblepus nicéfori Myers, 1932: 137. Type locality: Sonsón, Antioquia, Colombia. Holotype: SU 24796.
Distribution: Magdalena River basin (Schaefer, 2003b); Cauca River basin, Colombia (Maldonado-Ocampo et al.,2005).
Astroblepus orientalis (Boulenger, 1903)
Arges orientalis Boulenger, 1903a: 601. Type locality: Albirregas and Milla rivers above Merida, Venezuela, 3500
m. altitude. Syntypes: BMNH 1903.6.30.81–89, USNM 133142.
Distribution: Lake Maracaibo drainage (Schaefer, 2003b).

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Astroblepus peruanus (Steindachner, 1876)
Arges peruanus Steindachner, 1876: 601, pl. 9 (fig. 3–6). Type locality: Amable Maria, Peru. Syntypes: NMW
16974 (3).
Distribution: Upper Ucayali River basin, Peru (Schaefer, 2003b).
Astroblepus phelpsi Schultz, 1944
Astroblepus phelpsi Schultz, 1944c: 283, pl. 7 (fig. d). Type locality: Río Cobre above its mouth near La Grita, tributary of Río Quinta, latter tributary to Río La Grita, Catatumbo system, Venezuela. Holotype: USNM 121126.
Distribution: Lake Maracaibo drainage (Schaefer, 2003b).
Astroblepus pholeter Collette, 1962
Astroblepus pholeter Collette, 1962: 311, fig. 1–2. Type locality: Cave in Latas, 4 km north of Archidona, Napo
Prov., eastern Ecuador. Holotype: USNM 196623.
Distribution: Napo River basin, Ecuador (Schaefer, 2003b).
Astroblepus pirrensis (Meek & Hildebrand, 1913)
Cyclopium pirrense Meek & Hildebrand, 1913: 83. Type locality: Río Cana, Cana, Panama. Holotype: FMNH 7586.
Distribution: Cana River basin, Panama (Schaefer, 2003b).
Astroblepus praeliorum Allen, 1942
Astroblepus praeliorum Allen, in Eigenmann & Allen, 1942: 169, pl. 13 (fig. 6–7). Type locality: Tarma; Palca;
Oroya; Huancachupa Creek, Húanuco, Peru. Syntypes: CAS 7509 (4), CAS 16114 (82), CAS 16115 (2), CAS
47068 (10), CAS 47069 (1).
Distribution: Upper Marañon River basin, Peru (Schaefer, 2003b).
Astroblepus prenadillus (Valenciennes, 1840)
Brontes prenadilla Valenciennes, in Cuvier & Valenciennes, 1840b: 343 (255 of the Strasbourg deluxe edition), pl.
444. Type locality: des ruisseaux qui descendent du Cotopaxi [Ecuador], à 5000 mètres au-dessus dy niveau el’Ocean. Syntypes (2): MNHN a-4227 (1), MNHN a-5164 (1).
Distribution: Andes of Ecuador (Schaefer, 2003b).
Astroblepus regani (Pellegrin, 1909)
Arges regani Pellegrin, 1909c: 518. Type locality: [Río Cariyacu, Ecuador, elev. 3100 m]. Syntypes: MNHN 19030108 (1), MNHN 1903-0109 (2), MNHN 1903-0110 (3).
Distribution: Mira River basin and Pacific coastal drainage of Ecuador (Schaefer, 2003b).
Astroblepus rengifoi Dahl, 1960
Astroblepus rengifoi Dahl, 1960a: 459, unnumbered figure. Type locality: small brook, tributary to the Atrato Riveron the left side, approx. 6 ks. above the town El Carmen de Atrato, Colombia. Holotype: at ICNMHN, current
whereabouts unknown.
Distribution: Atrato River basin, Colombia (Schaefer, 2003b).
Astroblepus retropinnus (Regan, 1908)
Arges retropinna Regan, 1908f: 800, pl. 48 (fig. 2). Type locality: Jimenez, western Colombia. Syntypes: BMNH
1908.5.29.81–82 (2).
Distribution: Dagua River basin and Pacific coastal drainage of Colombia (Schaefer, 2003b).
Astroblepus riberae Cardona & Guerao, 1994
Astroblepus riberae Cardona & Guerao, 1994: 21, fig. Type locality: Cueva de Ninabamba, Peru, 06.33ºS, 78.51ºW.
Holotype: UAB unnumbered.
Distribution: Reque River basin and Pacific coastal drainage, Peru (Schaefer, 2003b).
Astroblepus rosei Eigenmann, 1922
Astroblepus rosei Eigenmann, 1922b: 55. Type locality: Río Jequetepeque, Llallán, Peru. Syntypes: CAS 64649(11), FMNH 58640 (1).
Distribution: Pacific coastal drainages of Cajamarca, Peru (Schaefer, 2003b); widely distributed in ColombianAndes (Maldonado-Ocampo et al., 2005).
Remarks: Eigenmann (1922: 55) stated that the type [= holotype] was a specimen deposited in the Indiana University collection, but eleven specimens were deposited there (now CAS 64649), and no indication was given in the
publication as to which was the type. Therefore, all eleven, plus one specimen deposited in the Carnegie

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FERRARISMuseum collection (now FMNH 58640), must be considered syntypes.
Astroblepus sabalo (Valenciennes, 1840)
Arges sabalo Valenciennes, in Cuvier & Valenciennes, 1840b: 335 (249 in Strasbourg deluxe edition), pl. 445. Type
locality: le haut Pérou. Holotype: MNHN a-4226.
Distribution: Upper Ucayali River basin, Peru (Schaefer, 2003b).
Astroblepus santanderensis Eigenmann, 1918
Astroblepus cyclopus santanderensis Eigenmann, 1918b: 675. Type locality: Quebrada de Guapota, Colombia. Lectotype: FMNH 58433, designated by Henn (1928: 82).
Distribution: Magdalena River basin, Colombia (Schaefer, 2003b); Suarez River basin, Colombia (MaldonadoOcampo et al., 2005).
Astroblepus simonsii (Regan, 1904)
Arges simonsii Regan, 1904b: 317, pl. 21 (fig. 9). Type locality: Huaras, Peru, elev. 10700 ft. Syntypes: BMNH
1900.6.20.51–55 (5).
Distribution: Santa River basin and Pacific coastal drainages of Peru (Schaefer, 2003b).
Astroblepus stuebeli (Wandolleck, 1916)
Arges stübeli Wandolleck, 1916: 158, fig. 1–4. Type locality: Lake Titicaca. Syntypes (40): MTD F977, MTD
F4639–87.
Distribution: Lake Titicaca basin (Schaefer, 2003b).
Astroblepus supramollis Pearson, 1937
Astroblepus supramollis Pearson, 1937: 94, pl. 13 (fig. 3). Type locality: Balsas, Peru. Syntypes: CAS 47012 (389),
CAS 47015 (36), CAS 47022 (31).
Distribution: Marañon River basin, Peru (Schaefer, 2003b).
Astroblepus taczanowskii (Boulenger, 1890)
Arges taczanowskii Boulenger, 1890b: 451, pl. 41 (fig. 1). Type locality: Amable Maria, Peruvian Andes. Holotype:
BMNH 1875.10.14.19.
Distribution: Upper Ucayali River basin, Peru (Schaefer, 2003b).
Astroblepus theresiae (Steindachner, 1907)
Arges theresiae Steindachner, 1907c: 228. Type locality: Hohen Anden von Cayendelet. Syntypes: NMW 48091(10), NMW 92 (7).
Distribution: Pacific coastal drainages of Ecuador (Schaefer, 2003b).
Astroblepus trifasciatus (Eigenmann, 1912)
Cyclopium trifasciatum Eigenmann, 1912a: 14. Type locality: Caldas, Colombia. Holotype: FMNH 56076; holotype
illustrated in Eigenmann (1922: 52, fig. 15; pl. 6, fig. 6).
Distribution: Dagua River basin and Pacific coastal drainages of Colombia (Schaefer, 2003b); also Magdalena andCauca River basins (Maldonado-Ocampo et al., 2005).
Astroblepus ubidiai (Pellegrin, 1931)
Cyclopium ubidiai Pellegrin, 1931b: 114. Type locality: [Lac St. Paul, near Otavalo, Imbabura Prov.,] Equateur, elev.
2600 m. Syntypes: MNHN 1930-0172 (1), MNHN 1930-0173 (1).
Distribution: Mira River basin and Pacific coastal drainages of Ecuador (Schaefer, 2003b; Vélez-Espino, 2003).
Astroblepus unifasciatus (Eigenmann, 1912)
Cyclopium unifasciatum Eigenmann, 1912a: 15. Type locality: Caldas, Colombia. Holotype: FMNH 56079; holotype illustrated in Eigenmann (1922: 52, pl. 7, fig. 2).
Distribution: Dagua River basin and Pacific coastal drainages of Colombia (Schaefer, 2003b); also Magdalena andCauca River basins (Maldonado-Ocampo et al., 2005).
Astroblepus vaillanti (Regan, 1904)
Arges vaillanti Regan, 1904b: 312. Type locality: Huamani Peak, Ecuador, elev. 10700 ft. Syntypes: BMNH
1897.2.16.2–4 (3).
Distribution: Pacific coastal drainages of Ecuador (Schaefer, 2003b).
Astroblepus vanceae (Eigenmann, 1913)
Cyclopium vanceae Eigenmann, 1913: 421, pl. 32 (fig. 2). Type locality: Small stream in highlands, southeast of

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Tarma, Peru. Holotype: FMNH 56066.
Distribution: Upper Ucayali River basin, Peru (Schaefer, 2003b).
Astroblepus ventralis (Eigenmann, 1912)
Cyclopium ventrale Eigenmann, 1912a: 15. Type locality: Caldas, Colombia. Holotype: FMNH 56074.
Distribution: Dagua River basin and Pacific coastal drainage of Colombia (Schaefer, 2003b).
Astroblepus whymperi (Boulenger, 1890)
Arges whymperi Boulenger, 1890b: 451, pl. 41 (fig. 2). Type locality: Milligalli, Andes of Ecuador. Holotype:
BMNH 1887.1.7.1.
Distribution: Esmeraldas River basin and other Pacific coastal drainages of Ecuador (Schaefer, 2003b).

AUCHENIPTERIDAE Bleeker, 1862
Euanemini Bleeker, 1858b: 49, 189. Type genus: Euanemus Müller & Troschel, 1842.
Trachelyopterini Bleeker, 1858b: 49, 250, 257. Type genus: Trachelyopterus Valenciennes, 1840.
Ageneiosi Bleeker, 1862 (in Bleeker, 1862–63): 14. Type genus: Ageneiosus La Cepède, 1803.
Astrophysi [= Asterophysi] Bleeker, 1862 (in Bleeker, 1862–63): 7. Type genus: Asterophysus Kner, 1857.
Auchenipterini Bleeker, 1862 (in Bleeker, 1862–63): 14. Type genus: Auchenipterus Valenciennes, 1840.
Centromochli Bleeker, 1862 (in Bleeker, 1862–63): 7. Type genus: Centromochlus Kner, 1857.
Pseudauchenipterini Bleeker, 1862 (in Bleeker, 1862–63): 6. Type genus: Pseudauchenipterus Bleeker, 1862.
Taxonomic summary: Britski (1972), Soares-Porto (1998, Centromochlinae), Ferraris (2003d).
Reviews: Mees (1974, Guianas).
Phylogeny: Ferraris (1988b), Curran (1989), Soares-Porto (1998, Centromochlinae), Royero (1999), Diogo et al.
(2003c).
20 genera, 94 species; no named fossil taxa.
AGENEIOSUS La Cepède, 1803
Ageneiosus La Cepède, 1803: 132. Type species: Ageneiosus armatus La Cepède, 1803. Type by subsequent designation by Eigenmann & Eigenmann (1890: 299). Gender: Masculine. Earlier type designation by Bleeker(1862–63: 14) of “Ageneiosus militaris Blkr (nec Val.) = Silurus militaris Bl.” not based on one of the originally
included species and therefore not valid.
Ceratorhynchus Spix & Agassiz, 1829: 10. Type species: Silurus militaris [= ? Silurus militaris of Bloch, nec Linnaeus]. Type by monotypy. Gender: Masculine.
Agenius Agassiz, 1846: 11. Type species: Ageneiosus armatus La Cepède, 1803. Gender: Masculine. Unjustified
emendation of Ageneiosus, therefore taking the same type species.
Davalla Bleeker, 1858b: 58, 64. Type species: Davalla schomburgkii Bleeker, 1858. Type by monotypy. Gender:
Feminine.
Pseudageneiosus Bleeker, 1862 (in Bleeker, 1862–63): 14. Type species: Ageneiosus brevifilis Valenciennes, 1840.
Type by original designation. Gender: Masculine.
Tympanopleura Eigenmann, 1912b: 203. Type species: Tympanopleura piperata Eigenmann, 1912. Type by original
designation. Gender: Feminine.
Remarks: Genus revised, with key to species, in Walsh (1990).
Ageneiosus atronasus Eigenmann & Eigenmann, 1888
Ageneiosus atronasus Eigenmann & Eigenmann, 1888b: 149. Type locality: Exact locality unknown. Holotype:
MCZ 27270.
Ageneiosus melanopogon Miranda Ribeiro, 1917: 51. Type locality: Fluvio Solimões [Brazil]. Holotype: Whereabouts unknown.
Tympanopleura nigricollis Eigenmann & Allen, 1942: 139, pl. 5 (figs. 2–3), 6 (fig. 3). Type locality: Iquitos [Peru].
Holotype: CAS 57940.
Distribution: Middle and upper Amazon River basin (Walsh, 1990).

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Ageneiosus brevis Steindachner, 1881
Ageneiosus brevis Steindachner, 1881d: 16. Type locality: Amazonenstrom bei Coary und aus dem Hyavary [Brazil]. Syntypes (4): NMW 47801 (2).
Ageneiosus rondoni Miranda Ribeiro, 1914: 12, pl. 1 (fig. 3). Type locality: Rio Negro (Manáos) [Brazil]. Lectotype: MNRJ 962A, designated by Miranda Ribeiro (1953: 394).
Ageneiosus madeirensis Fisher, 1917: 426, pl. 42. Type locality: San Joaquín, Bolivia. Holotype: FMNH 58143.
Tympanopleura alta Eigenmann & Myers, in Myers, 1928: 85. Type locality: Iquitos, Rio Marañon, Peru. Holotype:
CAS 58258.
Distribution: Upper Amazon River basin (Walsh, 1990).
Ageneiosus inermis (Linnaeus, 1766)
Silurus inermis Linnaeus, 1766: 503. Type locality: Surínami. Type (s): Whereabouts unknown.
Ageneiosus armatus La Cepède, 1803: 132, 133. Type locality: les eaux de Surinam, et peut-etre dans celles desgrandes Indes. Types: On Bloch (1794: 19, pl. 362); Silurus militaris Linné; and others.
Ageneiosus brevifilis Valenciennes, in Cuvier & Valenciennes, 1840b: 242 (180 in Strasbourg deluxe edition). Type
locality: Cayenne [but interpreted as having come from Suriname by Boeseman (1972: 303)]. Holotype: RMNH
2975 (larger of 2).
Hypothalmus dawalla Jardine, in Schomburgk, 1841: 191, pl. 9. Type locality: Junction of Rupununi and Essequibo[Guyana], and Fort St. Joaquim, in the Rio Bronco. No types known. Name spelled Hypothalmus davalla onplate; H. dawalla in text; first reviser not found.
Davalla Schomburgkii Bleeker, 1858b: 64. Type locality: Junction of Rupununi and Essequibo [Guyana], and Fort
St. Joaquim, in the Rio Bronco. No types known. Unneeded replacement for Hypothalmus dawalla Jardine,
1841.
Ageniosus axillaris Günther, 1864: 431. Type locality: Surinam. Holotype: BMNH 1864.6.2.2.
Ageneiosus sebae Günther, 1864: 192. Type locality: Suriname. Syntypes: On specimen(s) described as Ageneiosus
inermis in Cuvier & Valenciennes (1840b: 240 (178 in Strasbourg deluxe edition), pl. 440) and specimen illustrated in Seba (1734–65: pl. 29, fig. 8).
? Ageneiosus (Pseudageneiosus) therezinae Steindachner, 1909b: 341. Type locality: Rio Parnahyba und Rio Puty
bei Therezina [Brazil]. Syntypes: NMW 47840–41 (2 of “several”).
Ageneiosus ogilviei Fowler, 1914: 266, fig. 15. Type locality: Rupununi River, British Guiana ... in the highlands ofBritish Guiana, approximately secured in North Latitude 2º to 3º, and West Longitude 50º20'. Holotype: ANSP
39343.
Ageneiosus gabardinii Risso & Risso, 1964b: 12, pl. 3 (fig. 1). Type locality: Río Paraná, a la altura de la Ciudad deCorrientes [Argentina]. Holotype: Mus. Cienc. Nat. del Chaco 3, VI-63-2.
Distribution: Widely distributed in South America, east of the Andes (Walsh, 1990).
Ageneiosus magoi Castillo & Brull, 1989
Ageneiosus magoi Castillo & Brull, 1989: 73, fig. 1–2. Type locality: Los esteros de Camaguán, km 270, Estado
Guárico [Venezuela]. Holotype: MBUCV V-15666.
Distribution: Orinoco basin of Venezuela (Ferraris, 2003d).
Ageneiosus marmoratus Eigenmann, 1912
Ageneiosus marmoratus Eigenmann, 1912b: 206, pl. 22 (fig. 1). Type locality: Creek below Potaro Landing, British
Guiana. Holotype: FMNH 53245.
? Ageneiosus barranquerensis Risso & Risso, 1964b: 11, pl. 2. Type locality: frente a la desembocadura del Río
Negro, Riacho Barranqueras [Argentina]. Holotype: Mus. Cien. Nat. del Chaco 11, III-63.
Distribution: Widely distributed in South America east of the Andes mountains (Walsh, 1990).
Ageneiosus militaris Valenciennes, 1836
Ageniosus militaris Valenciennes, 1836, in Valenciennes, 1835–47: pl. 4 (fig. 1). Type locality: [not stated]. Holotype: possibly MNHN b-0690 (1), MNHN b-0691 (1), or MNHN b-3167 (1). Name available from plate (illustrated specimen is holotype, if identifiable); later described in Cuvier & Valenciennes (1840b: 233; 173 in
Strasbourg deluxe edition), with type locality as: La Plata.
Ageneiosus valenciennesi Bleeker, 1864a: 82. Type locality: La Plata. Holotype: Possibly MNHN b-0690, MNHNCHECKLIST OF CATFISHES, RECENT AND FOSSIL

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b-0691, or MNHN b-3167. New name for Ageneiosus militaris Valenciennes.
Silurus 38-radiatus Larrañaga, 1923: 386. Type locality: Uruguay. No types known.
Silurus imberbis Larrañaga, 1923: 376. Type locality: Uruguay. No types known. Either based on, or preoccupied by,
Silurus imberbis Gmelin, 1789.
Ageneiosus uruguayensis Devincenzi, 1933: 3, pl. 1. Type locality: Río Uruguay frente a Paysandú [Uruguay].
Holotype: MHNM CI 317.
Ageneiosus marquesi Risso & Risso, 1964b: 20, pl. 6 (fig. 1). Type locality: Río Paraná, frente al tanque de obras
Sanitarias de la ciudad de Corrientes, Argentina. Holotype: Mus. Cienc. Nat. del Chaco 12-V-64-1.
Distribution: La Plata River basin (Walsh, 1990).
Ageneiosus pardalis Lütken, 1874
Ageneiosus pardalis Lütken, 1874b: 190. Type locality: Caraccas [apparently in error, S. Walsh (1990, and pers.
comm.)]. Holotype: ZMUC P 29664.
Ageneiosus caucanus Steindachner, 1880b: 61, pl. 6 (figs. 1, 1a). Type locality: [Río] Cauca [Colombia]. Syntypes:
NMW 47811 (2).
Ageneiosus virgo Posada, 1909: 295. Type locality: [Río] Magdalena [Colombia]. Type(s): Whereabouts unknown.
Ageneiosus freiei Schultz, 1944c: 240, pl. 4 (fig. b). Type locality: Río Agua Caliente, 2 to 3 km above Lago Maracaibo [Venezuela]. Holotype: USNM 121260.
Distribution: Lake Maracaibo basin and Magdalena, Cauca, San Juan, and Tuira River basins (Walsh, 1990).
Ageneiosus piperatus (Eigenmann, 1912)
Tympanopleura piperata Eigenmann, 1912b: 203, pl. 20 (fig. 3). Type locality: Crab Falls [Essequibo River], British
Guiana. Holotype: FMNH 53243.
Distribution: Essequibo and upper Negro rivers near confluence of Branco River (Walsh, 1990).
Ageneiosus polystictus Steindachner, 1915
Ageneisus polystictus Steindachner, 1915b: 217. Type locality: [not stated]. Holotype: NMW 47839. Illustrated anddescribed in more detail in Steindachner (1915e: 84, pl. 7, figs. 1–3), as Agenciosus polystictus, with origin of
specimen reported as “Mündung des Rio Negro.”Distribution: Negro River and lower Branco River, Brazil (Walsh, 1990).
Ageneiosus ucayalensis Castelnau, 1855
Ageneiosus ucayalensis Castelnau, 1855: 49, pl. 17 (fig. 2). Type locality: un lac, près de l’Ucayale, et qui communiquait avec cette rivière [Peru]. Holotype: MNHN b-0611.
? Ageneiosus dentatus Kner, 1857: 441. Type locality: Surinam. Syntypes: NMW 47823 (6).
? Ageniosus porphyreus Cope, 1867: 404. Type locality: Surinam. Lectotype: ANSP 8389, designated by Fowler(1915a: 224).
? Ageneiosus parnaguensis Steindachner, 1910c: 399, figured on p. 400. Type locality: see von Lake Parnaguá inbrasílienischen Staate Piauhy [Brazil]. Syntypes: NMW 47832 (1), NMW 47837 (1), NMW 47838 (2).
? Ageneiosus guianensis Eigenmann, 1912b: 204, pl. 21 (fig. 2). Type locality: Wismar, British Guiana. Holotype:
FMNH 53247.
Distribution: Amazon and Orinoco River basins; Guianas, and upper Paraná River basin (Walsh, 1990).
Ageneiosus vittatus Steindachner, 1908
Ageneiosus vittatus Steindachner, 1908b: 64. Type locality: Rio Purus [Brazil]. Holotype: NMW 47853.
Distribution: Middle Orinoco and upper Amazon River basins (Walsh, 1990).
ASTEROPHYSUS Kner, 1857
Asterophysus Kner, 1857: 402. Type species: Asterophysus batrachus Kner, 1857. Type by monotypy. Gender: Masculine.
Astrophysus Bleeker, 1858b: 356, 358. Type species: Asterophysus batrachus Kner, 1857. Type by being a replacement name. Gender: Masculine. Unjustified emendation of Asterophysus Kner, 1857.
Asterophysus batrachus Kner, 1857
Asterophysus batrachus Kner, 1857: 403, pl. 5 (fig. 13). Type locality: Marabitanos [Brazil]. Syntypes: NMW
47515–16 (2).

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FERRARISDistribution: Orinoco and Negro River basins, Brazil and Venezuela (Ferraris, 2003d).
AUCHENIPTERICHTHYS Bleeker, 1862
Auchenipterichthys Bleeker, 1862 (in Bleeker, 1862–63): 7. Type species: Auchenipterus thoracatus Kner, 1857.
Type by original designation. Gender: Masculine.
Revision: Ferraris et al. (2005).
Auchenipterichthys coracoideus (Eigenmann & Allen, 1942)
Trachycorystes coracoideus Eigenmann & Allen, 1942: 120. Type locality: Iquitos [Peru]. Syntypes: CAS 63746(3).
Distribution: Amazon River basin and Araguaia River of the Tocantins basin (Ferraris et al., 2005).
Auchenipterichthys longimanus (Günther, 1864)
Auchenipterus longimanus Günther, 1864: 195. Type locality: River Capin, Para [Brazil]. Syntypes: BMNH
1849.11.8.11 (3).
Distribution: Lower and middle Amazon, Tocantins, and Orinoco river basins (Ferraris et al., 2005).
Remarks: Syntype register number misreported in Eschmeyer et al. (1998) as BMNH 1849.121.8.? BMNH
1849.11.8.? and Ferraris (2003d) as BMNH 1849.121.8.
Auchenipterichthys punctatus (Valenciennes, 1840)
Auchenipterus punctatus Valenciennes, in Cuvier & Valenciennes, 1840b: 219 (163 in Strasbourg deluxe edition).
Type locality: Probablement du Brésil. Holotype: MNHN b-0216; illustrated in Royero & Hureau (1996: fig. 3).
Auchenipterichthys dantei Soares-Porto, 1994: 282, fig. 3. Type locality: Brazil, Amazonas, Paricatuba, Rio Negro(3º07'S, 60º26'W). Holotype: MZUSP 43332.
Distribution: Upper Orinoco River and Upper Amazon River basins, including the Negro River (Ferraris et al.,2005).
Auchenipterichthys thoracatus (Kner, 1857)
Auchenipterus thoracatus Kner, 1857: 425, pl. 7 (fig. 22). Type locality: Rio Guaporé. Syntypes: NMW 47454 (2).
Auchenipterus thoracicus Günther, 1864: 194. Type locality: Rio Guaporé. Syntypes: NMW 47454 (2). Unjustified
emendation of Auchenipterus thoracatus Kner, 1857.
Distribution: Upper Madeira River basin (Ferraris et al., 2005).
Remarks: Syntype register number incorrectly reported as NMW 47452 in Eschmeyer et al. (1998) and Ferraris(2003d).
AUCHENIPTERUS Valenciennes, 1840
Auchenipterus Valenciennes, in Cuvier & Valenciennes, 1840b: 207 (154 in Strasbourg deluxe edition). Type species: Hypophthalmus nuchalis Spix & Agassiz, 1829. Type by subsequent designation by Bleeker (1862–63:
15). Gender: Masculine.
Euanemus Müller & Troschel, in Müller, 1842a: 203. Type species: Euanemus colymbetes Müller & Troschel, 1842.
Type by monotypy. Gender: Masculine. Also as new in Müller (1842b: 320). Priority of publication not established.
Auchenopterus Agassiz, 1846: 40. Type species: Hypophthalmus nuchalis Spix & Agassiz, 1829. Type by being a
replacement name. Gender: Masculine. Unjustified emendation of Auchenipterus Valenciennes, 1840.
Ceratocheilus Miranda Ribeiro, 1918d: 644. Type species: Ceratocheilus osteomystax Miranda Ribeiro, 1918. Typeby monotypy. Gender: Masculine. Preoccupied by Ceratocheilus Wesché, 1810, in Diptera, replaced by
Osteomystax Whitley, 1940.
Osteomystax Whitley, 1940a: 242. Type species: Ceratocheilus osteomystax Miranda Ribeiro, 1918. Type by being a
replacement name. Gender: Masculine. Replacement for Ceratocheilus Miranda Ribeiro, 1918, preoccupied by
Ceratocheilus Wesché, 1810, in Diptera.
Remarks: Revised, with a key to species, in Ferraris & Vari (1999).
Auchenipterus ambyiacus Fowler, 1915
Auchenipterus ambyiacus Fowler, 1915a: 222, fig. 6. Type locality: Ambyiacu River, Ecuador. Holotype: ANSP
21484.

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Zootaxa 1418 © 2007 Magnolia Press · 71Distribution: Upper and middle Amazon and Orinoco River basins, coastal rivers of Guyana (Ferraris & Vari, 1999).
Auchenipterus brachyurus (Cope, 1878)
Euanemus brachyurus Cope, 1878: 676. Type locality: Peruvian Amazon. Holotype: ANSP 21552.
Distribution: Upper Amazon River basin (Ferraris & Vari, 1999).
Auchenipterus brevior Eigenmann, 1912
Auchenipterus brevior Eigenmann, 1912b: 202. Type locality: Tumatumari, British Guiana. Holotype: FMNH
53249.
Distribution: Potaro River basin, Guyana (Ferraris & Vari, 1999).
Auchenipterus britskii Ferraris & Vari, 1999
Auchenipterus britskii Ferraris & Vari, 1999: 414, figs. 10–11. Type locality: Lago Janauacá, vicinity of Rio
Solimões (approx. 3º28'S, 60º17'W), Amazonas, Brazil. Holotype: MZUSP 48358.
Distribution: Middle Amazon River, Brazil (Ferraris & Vari, 1999).
Auchenipterus demerarae Eigenmann, 1912
Auchenipterus demerarae Eigenmann, 1912b: 202, pl. 21 (fig. 1). Type locality: Wismar, British Guiana. Holotype:
FMNH 53248; holotype illustrated in Ferraris & Vari (1999: fig. 12).
Distribution: Demerara, Rupununi, Essequibo and Cuyuni River basins (Ferraris & Vari, 1999).
Auchenipterus dentatus Valenciennes, 1840
Auchenipterus dentatus Valenciennes in Cuvier & Valenciennes, 1840b: 210 (156 in Strasbourg deluxe edition).
Type locality: envoyé de Cayenne [apparently originating from vicinity of Paramaribo, Suriname]. Holotype:
MNHN b-0592.
Euanemus colymbetes Müller & Troschel, in Müller, 1842: 203. Type locality: Suriname. Lectotype: ZMB 3095,
designated by Ferraris & Vari (1999). Also indicated as new, with one syntype illustrated, in Müller & Troschel(1849: 11, pl. 1, fig. 2).
Distribution: Northern coastal rivers of South America: Corantijn, Suriname, Arataye, Sinnamary and Oyapock rivers (Ferraris & Vari, 1999).
Auchenipterus fordicei Eigenmann & Eigenmann, 1888
Auchenipterus fordicei Eigenmann & Eigenmann, 1888b: 151. Type locality: Coary [Brazil]. Holotype: MCZ 7289.
Distribution: Middle Amazon River basin, Brazil (Ferraris & Vari, 1999).
Auchenipterus menezesi Ferraris & Vari, 1999
Auchenipterus menezesi Ferraris & Vari, 1999: 426, fig. 17. Type locality: Lago de Viana, Rio Pindaré, Mirim basin,
Maranhão, Brazil. Holotype: MZUSP 43591.
Distribution: Parnaíba and Pindaré-Mirim River basins of Maranhão and Piaui States (Ferraris & Vari, 1999).
Auchenipterus nigripinnis (Boulenger, 1895)
Euanemus nigripinnis Boulenger, 1895b: 524. Type locality: Paraguay. Lectotype: BMNH 1895.5.17.37, designatedby Ferraris & Vari (1999); one syntype illustrated in Boulenger (1896c: pl. 4, figs. 2, 2a).
Auchenipterus paysanduanus Devincenzi, 1933: 2. Type locality: Río Uruguay, frente a Paysandú, Uruguay. Holotype: MNHM 4889; illustrated in Ferraris & Vari (1999: fig. 20).
Distribution: La Plata River basin (Ferraris & Vari, 1999).
Auchenipterus nuchalis (Spix & Agassiz, 1829)
Hypophthalmus nuchalis Spix & Agassiz, 1829: 17, pl. 17. Type locality: Brasilia aequatoriali [now: Rio Capim,
near São Domingos do Capim, Brazil, by neotype designation]. Neotype: MZUSP 47945, illustrated in, anddesignated by, Ferraris & Vari (1999: fig. 21).
Distribution: Lower Amazon and Tocantins rivers northward to Marowijne River; possibly also in Rupununi River,
Guyana, and Negro River, Brazil (Ferraris & Vari, 1999).
Auchenipterus osteomystax (Miranda Ribeiro, 1918)
Ceratocheilus osteomystax Miranda Ribeiro, 1918d: 644, unnumbered pl. Type locality: Santa Rita das Antas, Rio
Vermelho, Goyaz [Brazil]. Holotype: MZUSP 2317.
Distribution: La Plata River basin, Tocantins River, and several rivers of lower Amazon River basin (Ferraris & Vari,
1999).

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FERRARISCENTROMOCHLUS Kner, 1857
Centromochlus Kner, 1857: 430. Type species: Centromochlus megalops Kner, 1857. Type by subsequent designation by Bleeker (1862–63: 7). Gender: Masculine.
Remarks: Included species follows Soares-Porto (1998), except for continued recognition of Gelanoglanis.
Centromochlus altae Fowler, 1945
Centromochlus altae Fowler, 1945a: 109, figs. 7–9. Type locality: Morelia, Río Caquetá basin, Colombia. Holotype:
ANSP 71700.
Distribution: Caquetá River basin, Colombia (Ferraris, 2003d).
Centromochlus concolor (Mees, 1974)
Tatia concolor Mees, 1974: 84, fig. 22. Type locality: Headwaters of Coppename Rivier [sic] (3º49'N, 56º57'W),
Suriname. Holotype: ZMA 106210.
Distribution: Coppename River basin, Suriname (Ferraris, 2003d).
Centromochlus existimatus Mees, 1974
Centromochlus existimatus Mees, 1974: 50, fig. 8 (right). Type locality: Harbour of Manaos [Brazil]. Holotype:
ANSP 73154.
Distribution: Amazon River basin (Ferraris, 2003d).
Centromochlus heckelii (De Filippi, 1853)
Auchenipterus Heckelii De Filippi, 1853: 167. Type locality: Rio Napo. Syntype: ZMUT 233.
Centromochlus Steindachneri Gill, 1870: 95. Type locality: Marañon River, upper Amazon and Napo Rivers.
Type(s): at USNM, but not found (Ferraris & Vari, 1992).
Distribution: Amazon and Orinoco River basins (Ferraris, 2003d).
Centromochlus macracanthus Soares-Porto, 2000
Centromochlus macracanthus Soares-Porto, 2000: 282, fig. 1. Type locality: Brazil: Amazonas: São Pedro, rio
Negro. Holotype: MZUSP 30620.
Distribution: Upper Negro River basin, Brazil (Soares-Porto, 2000).
Centromochlus perugiae Steindachner, 1882
Centromochlus Perugiae Steindachner, 1882b: 178. Type locality: Canelos [Ecuador]. Syntypes: NMW 47358 (3),
SMF 5355 (1). Species illustrated and described in more detail in Steindachner (1882c: 29, pl. 7, figs. 2, 2a).
Distribution: Upper Amazon River basin (Ferraris, 2003d).
Remarks: Wrongly listed as Tatia perugiae by Ferraris (2003d).
Centromochlus punctatus (Mees, 1974)
Tatia punctata Mees, 1974: 88, fig. 24. Type locality: Creeks between Kabel and Lombé [Suriname]. Holotype:
RMNH 26495.
Distribution: Coastal rivers of northern South America, above Amazon River mouth (Ferraris, 2003d).
Centromochlus reticulatus (Mees, 1974)
Tatia reticulata Mees, 1974: 90, fig. 25. Type locality: Karanambo, Rupununi, British Guiana. Holotype: BMNH
1972.7.27.702.
Distribution: South America: Rupununi River basin, Guyana (Ferraris, 2003d).
Centromochlus romani (Mees, 1988)
Tatia romani Mees, 1988a: 412, fig. 3. Type locality: Río Guarapiche, affluent of the Río San Juan, near Maturín,
Monagas [Venezuela]. Holotype: RMNH 30490.
Distribution: San Juan River, Orinoco Basin, Venezuela (Ferraris, 2003d).
Centromochlus schultzi Rössel, 1962
Centromochlus schultzi Rössel, 1962a: 27, fig. 1. Type locality: Brasilien, oberer Rio Xingu. Holotype: SMF 5462.
Distribution: Upper Xingu and Tocantins River basins (Ferraris, 2003d).
Species inquirenda, Centromochlus
Centromochlus megalops Kner, 1857: 430, pl. 8 (fig. 24). Type locality: Bogota. Syntypes: NMW 47359 (1), NMW
47360 (1).

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ENTOMOCORUS Eigenmann, 1917
Entomocorus Eigenmann, 1917a: 403. Type species: Entomocorus benjamini Eigenmann, 1917. Type by monotypy.
Gender: Masculine.
Entomocorus benjamini Eigenmann, 1917
Entomocorus benjamini Eigenmann, 1917a: 403, fig. 3, pl. 41. Type locality: San Joaquin [Bolivia]. Holotype:
FMNH 58109.
Distribution: Upper Madeira River basin, Bolivia (Ferraris, 2003d).
Entomocorus gameroi Mago-Leccia, 1984
Entomocorus gameroi Mago-Leccia, 1984: 217, figs. 1–6. Type locality: Boca del Río Apurito en el Río Apuré,cerca de San Fernando de Apuré, Guárico State, Venezuela. Holotype: MBUCV V-13808.
Distribution: Apuré River basin, Orinoco River drainage, Venezuela (Ferraris, 2003d).
Entomocorus melaphareus Akama & Ferraris, 2003
Entomocorus melaphareus Akama & Ferraris, 2003: 78, figs, 1–2. Type locality: Brazil, Pará, Monte Alegre, Costa
do Curuá, left bank of Amazon River, 2º13'46.2"S, 54º01'27.6"W. Holotype: MZUSP 76413.
Distribution: Lower and Middle Amazon River (Akama & Ferraris, 2003).
EPAPTERUS Cope, 1878
Epapterus Cope, 1878: 677. Type species: Epapterus dispilurus Cope, 1878. Type by monotypy. Gender: Masculine.
Revision, with a key to species: Vari & Ferraris (1998).
Epapterus blohmi Vari, Jewett, Taphorn & Gilbert, 1984
Epapterus blohmi Vari, Jewett, Taphorn & Gilbert, 1984: 463, fig. 1. Type locality: Laguna Los Guácimos, a broad
shallow lagoon located on Fundo Pecuario Masaguaral, Guarico State, Venezuela, 8º34'N, 67º35'W. Holotype:
MBUCV V-14100.
Distribution: Orinoco River basin and Tuy River of the Caribbean coast of Venezuela (Vari & Ferraris, 1998).
Epapterus dispilurus Cope, 1878
Epapterus dispilurus Cope, 1878: 677. Type locality: Peruvian Amazon. Lectotype: ANSP 21353, designated by
Fowler (1941b: 468, fig. 26) in caption to illustration.
Euanemus longipinnis Steindachner, 1881d: 17. Type locality: Hyavary. Lectotype: NMW 46682.1 (93 m SL), designated by Vari & Ferraris (1998: 992).
Epapterus chaquensis Risso & Risso, 1962: 5, figs. 1–3. Type locality: La laguna concida como “del Golf,” en laproximidades de la ciudad de Resistencia, Provincia del Chaco, Argentina. Holotype: Univ. Nac. del Nordeste[Argentina] 17/VII/961-1.
Distribution: Central and western parts of the Amazon River basin along and south of the main channel of the Amazon River, and Paraguay River basin in Paraguay, northern Argentina and southern Brazil (Vari & Ferraris,
1998).
GELANOGLANIS Böhlke, 1980
Gelanoglanis Böhlke, 1980: 150. Type species: Gelanoglanis stroudi Böhlke, 1980. Type by original designation.
Gender: Masculine.
Gelanoglanis nanonocticolus Soares-Porto, Walsh, Nico & Netto, 1999
Gelanoglanis nanonocticolus Soares-Porto, Walsh, Nico & Netto, 1999: 65, fig. 1. Type locality: Venezuela: Amazonas State: Río Asisa 22–30 km upstream from confluence with Río Paru, Río Ventuari drainage, Orinoco
River basin (approximately 4º33'N, 65º54'W). Holotype: MCNG 22690.
Distribution: Asisa River basin in Orinoco drainage, Venezuela, and Negro River basin, Brazil (Soares-Porto et al.,1999).
Gelanoglanis stroudi Böhlke, 1980
Gelanoglanis stroudi Böhlke, 1980: 152, fig. 1. Type locality: Colombia, Depto. Meta, Río Metica, ca. 22 km (air)SW of Puerto Lopez and 3 km SE of Hacienda Mozambique, 3º56'42"N, 73º02'23"W. Holotype: ANSP 142937.
Distribution: Meta, Apure and Masparro River basins, Colombia and Venezuela (Provenzano, 1997).

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FERRARISGLANIDIUM Lütken, 1874
Glanidium Lütken, 1874c: 31. Type species: Glanidium albescens Reinhardt, 1874. Type by monotypy. Gender:
Neuter.
Gephyromochlus Hoedeman, 1961: 135. Type species: Centromochlus (Gephyromochlus) leopardus Hoedeman,
1961. Type by monotypy. Gender: Masculine.
Glanidium albescens Reinhardt, 1874
Glanidium albescens Reinhardt, in Lütken, 1874c: 31. Type locality: in flumine Rio das Velhas cum affluentibus[Brazil]. Syntypes: MNHN 0000-9571 (1), NMW 46572 (1), ZMB 9176 (1), ZMUC P 29670–29675 (1 each).
Distribution: Das Velhas River basin, Brazil (Ferraris, 2003d).
Glanidium bockmanni Sarmento-Soares & Buckup, 2005
Glanidium bockmanni Sarmento-Soares & Buckup, 2005: 847, fig. 1. Type locality: Brazil, Minas Gerais state, Rio
São Francisco, downstream from Três Marias Dam. Holotype: MZUSP 82803.
Distribution: São Francisco River basin, Brazil (Sarmento-Soares & Buckup, 2005).
Glanidium catharinensis Miranda Ribeiro, 1962
Glanidium catharinensis Miranda Ribeiro, 1962a: 3, fig. 2. Type locality: Rio do Braço do Norte, São Ludgero,
Municipio de Tubarão, Estado de Santa Catarina [Brazil]. Holotype: MNRJ 5169.
Distribution: Tubarão River basin in Santa Catarina State, Brazil (Ferraris, 2003d).
Glanidium cesarpintoi Ihering, 1928
Glanidium cesarpintoi Ihering, 1928b: 46, fig. 1. Type locality: Cachoeira de Emas (Pirassununga), Rio Mogyguassú, Est. de S. Paulo [Brazil]. Syntypes (3): whereabouts unknown.
Distribution: Mogi-guassú River basin, São Paulo State, Brazil (Ferraris, 2003d).
Glanidium leopardum (Hoedeman, 1961)
Centromochlus (Gephyromochlus) leopardus Hoedeman, 1961: 135, fig. 3. Type locality: Litany River, village
Aloiké, sta. 295 [French Guiana]. Holotype: ZMA 102233.
Distribution: Coastal rivers of the Guianas (Ferraris, 2003d).
Glanidium melanopterum Miranda Ribeiro, 1918
Glanidium melanopterum Miranda Ribeiro, 1918d: 643. Type locality: Piquete, S. Paulo [Brazil]. Lectotype:
MZUSP 345, designated in Britski (1969).
Distribution: Piquete, São Paulo, Brazil (Ferraris, 2003d).
Glanidium ribeiroi Haseman, 1911
Glanidium ribeiroi Haseman, 1911b: 381, pl. 78. Type locality: Porto União da Victoria, Paraná, Brazil, from Rio
Iguassú. Holotype: FMNH 54253.
Distribution: Iguaçu River basin (Ferraris, 2003d).
LIOSOMADORAS Fowler, 1940
Liosomadoras Fowler, 1940a: 226. Type species: Liosomadoras morrowi Fowler, 1940. Type by original designation. Gender: Masculine.
Liosomadoras morrowi Fowler, 1940
Liosomadoras morrowi Fowler, 1940a: 226, figs. 11–14. Type locality: Ucayali River basin, Contamana, Peru.
Holotype: ANSP 68646.
Distribution: Ucayali River basin, Peru (Ferraris, 2003d).
Liosomadoras oncinus (Jardine, 1841)
Arius oncinus Jardine, in Schomburgk, 1841: 173, pl. 4. Type locality: Padauiri River. No types known. Name
spelled Arius oncina on figure caption, A. oncinus in text. Mees (1978a) served as first reviser in treating Liosomadoras oncinus as valid.
Distribution: Negro and Branco River basins, Brazil (Ferraris, 2003d); Venezuela (Taphorn et al., 1997); Orinoco
River basin (Royero, 1999).
PSEUDAUCHENIPTERUS Bleeker, 1862
Pseudauchenipterus Bleeker, 1862 (in Bleeker, 1862–63): 6. Type species: Silurus nodosus Bloch, 1794. Type by

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original designation. Gender: Masculine.
Silvaichthys Fernández-Yépez, 1973: [3]. Type species: Silvaichthys aguilerae Fernández-Yépez, 1973. Type byoriginal designation. Gender: Masculine.
Pseudauchenipterus affinis (Steindachner, 1877)
Auchenipterus (Pseudauchenipterus) affinis Steindachner, 1877b: 651. Type locality: Rio S. Matheus; Rio Mucury
bei Porto Allegre [Brazil]. Syntypes: NMW 47384 (2), NMW 47385 (2), NMW 47386 (2), NMW 47387 (2).
Distribution: Mucuri and São Mateus River basins, Brazil (Ferraris, 2003d).
Pseudauchenipterus flavescens (Eigenmann & Eigenmann, 1888)
Felichthys flavescens Eigenmann & Eigenmann, 1888b: 152. Type locality: Rio San Francisco [Brazil]. Holotype:
MCZ 7362.
Distribution: São Francisco River basin, Brazil (Ferraris, 2003d).
Pseudauchenipterus jequitinhonhae (Steindachner, 1877)
Auchenipterus (Pseudauchenipterus) Jequitinhonhae Steindachner, 1877b: 647, pl. 6 (fig. 1). Type locality: Rio
Jequitinhonha [Brazil]. Syntypes: NMW 19785–92 (8).
Distribution: Jequitinhonha River basin, Brazil (Ferraris, 2003d).
Pseudauchenipterus nodosus (Bloch, 1794)
Silurus nodosus Bloch, 1794: 35, pl. 368 (fig. 1). Type locality: Tranquebar [in error]. Lectotype: ZMB 3096, designated by, and illustrated in, Paepke (1999: 47, pl. 26, fig. 1).
Auchenipterus furcatus Valenciennes, in Cuvier & Valenciennes, 1840b: 211 (157 in Strasbourg deluxe edition).
Type locality: Cayenne ou de Surinam. Syntypes: MNHN b-0213 (1), MNHN 0000-3049 (1), MNHN a-8855(1); MNHN b-0213 illustrated in Royero & Hureau (1996: fig. 1).
Pseudauchenipterus guppyi Regan, 1906c: 387, pl. 23. Type locality: Caroni River [Trinidad]. Syntypes: BMNH
1906.6.23.49–50 (2).
Pseudauchenipterus nigrolineatus Fowler, 1911: 434, fig. 5. Type locality: Pedernales, Venezuela. Holotype: ANSP
37876.
Silvaichthys aguilerae Fernández-Yépez, 1973: [4], unnumbered figure. Type locality: Gulfo de Paria, noreste deVenezuela, frente a IRAPA, Estado Suene. Syntypes (4): originally as “Holotipo macho AFY 731.201, Holotipo
hembra AFY 731.202, y los paratipos ... AFY 731.203 (2)”; whereabouts unknown, not at ANSP as indicated in
the description.
Distribution: Lower reaches of rivers and into estuaries from Venezuela and southern Trinidad to Brazil, south atleast to Bahia (Ferraris, 2003d).
PSEUDEPAPTERUS Steindachner, 1915
Pseudepapterus Steindachner, 1915a: 199. Type species: Auchenipterus (Pseudepapterus) hasemani Steindachner,
1915. Type by monotypy. Gender: Masculine. Originally proposed as a subgenus of Auchenipterus.
Revision, with a key to species: Ferraris & Vari (2000).
Pseudepapterus cucuhyensis Böhlke, 1951
Pseudepapterus cucuhyensis Böhlke, 1951b: 38, fig. 1. Type locality: Sandbank at Cucuhy on the Columbian [sic]border, Rio Negro, Brazil. Holotype: SU 16788.
Distribution: Amazon and Negro River basins (Ferraris & Vari, 2000).
Pseudepapterus gracilis Ferraris & Vari, 2000
Pseudepapterus gracilis Ferraris & Vari, 2000: 109, fig. 5. Type locality: Venezuela, Bolivar, Caño Chuapo, approximately 20 minutes downstream from Jabillal (on opposite bank) along Rio Caura, 7º07'N, 65º00'W. Holotype:
ANSP 139502.
Distribution: Caura River basin, Venezuela; known only from holotype (Ferraris & Vari, 2000).
Pseudepapterus hasemani (Steindachner, 1915)
Auchenipterus (Pseudepapterus) hasemani Steindachner, 1915a: 199. Type locality: Pará (Belem) [Brazil]. Holotype: NMW 47397. Described in more detail in Steindachner (1915e: 82).
Distribution: Amazon River and many of its major tributary rivers, and Tocantins River basin (Ferraris & Vari,
2000).

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FERRARISPSEUDOTATIA Mees, 1974
Pseudotatia Mees, 1974: 105. Type species: Pseudotatia parva Mees, 1974. Type by original designation. Gender:
Feminine.
Pseudotatia parva Mees, 1974
Pseudotatia parva Mees, 1974: 105, fig. 29. Type locality: Joazeiro, Brazil. Holotype: FMNH 70580.
Distribution: São Francisco River basin, Brazil (Ferraris, 2003d).
TATIA Miranda Ribeiro, 1911
Tatia Miranda Ribeiro, 1911: 360. Type species: Centromochlus intermedius Steindachner, 1877. Type by subsequent designation by Jordan (1920: 545). Gender: Feminine.
Tatia aulopygia (Kner, 1857)
Centromochlus aulopygius Kner, 1857: 432, pl. 8 (fig. 25). Type locality: Rio Guaporé [Brazil]. Syntypes: NMW
47329 (2), NMW 47330 (2), NMW 47331 (3), NMW 47332 (3), NMW 47333 (5) NMW 47388 (1).
Distribution: Guaporé River basin (Ferraris, 2003d).
Tatia boemia Koch & Reis, 1996
Tatia boemia Koch & Reis, 1996: 86, fig. 2. Type locality: Rio Pelotas (tributary of the Rio Uruguai) approx.
27º47'S, 51º11'W, Rio Grande do Sul, Brazil. Holotype: MCP 17438.
Distribution: Upper Uruguay River basin (Ferraris, 2003d).
Tatia brunnea Mees, 1974
Tatia brunnea Mees, 1974: 84, fig. 21. Type locality: Compagnie Kreek [Suriname]. Holotype: RMNH 26196.
Distribution: Suriname and Marowijne River basins (Ferraris, 2003d).
Tatia creutzbergi (Boeseman, 1953)
Centromochlus creutzbergi Boeseman, 1953: 7, fig. 1c. Type locality: Djaikreek, Surinam. Holotype: RMNH 19440.
Distribution: Suriname coastal rivers and Cudajas (Ferraris, 2003d).
Tatia dunni (Fowler, 1945)
Centromochlus dunni Fowler, 1945a: 111, figs. 11–13. Type locality: Morelia, Río Caquetá drainage, Colombia.
Holotype: ANSP 71705.
Distribution: Caquetá River basin, Colombia (Ferraris, 2003d).
Tatia galaxias Mees, 1974
Tatia galaxias Mees, 1974: 86, fig. 23. Type locality: Caño de Quiribana into Río Orinoco, Venezuela. Holotype:
CAS 6567.
Distribution: Middle Orinoco River basin (Ferraris, 2003d).
Tatia gyrina (Eigenmann & Allen, 1942)
Centromochlus gyrinus Eigenmann & Allen, 1942: 118, pl. 5 (fig. 4). Type locality: brook near Rio Itaya, Iquitos[Peru]. Holotype: CAS 36979.
Distribution: Upper Amazon River basin (Ferraris, 2003d).
Tatia intermedia (Steindachner, 1877)
Centromochlus intermedius Steindachner, 1877b: 664. Type locality: Marabitanos; Pará [Brazil]. Syntypes (2):
NMW 47355 (1).
Distribution: Amazon River basin and rivers of Guianas (Ferraris, 2003d).
Remarks: Steindachner (1881d: 4) indicated that the original description was based on two specimens.
Tatia musaica Royero, 1992
Tatia musaica Royero, 1992: 301, fig. 1. Type locality: Río Atuana, Caño Cabeza de Manteco, approximadamente a
12 kms debajo del raudel Pereza, en el pidemonte del tepui Autana, Territorio Federal, Amazonas, Venezuela,
4º47'N, 67º23'W. Holotype: MBUCV V-15663.
Distribution: Atabapo and Autana River basins in upper Orinoco River drainage, Venezuela (Ferraris, 2003d).
Tatia neivai (Ihering, 1930)
Glanidium neivai Ihering, 1930: 99, pl. 13 (fig. 1). Type locality: Piracicaba, Rio Piracicaba, Est. de S. Paulo. Holotype: Whereabouts unknown.
Distribution: Piracicaba River basin in São Paulo State (Ferraris, 2003d).

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Tatia simplex Mees, 1974
Tatia simplex Mees, 1974: 90, fig. 26. Type locality: Xaventina house beach, Rio das Mortes, Mato Grosso, Brazil.
Holotype: BMNH 1971.7.29.5.
Distribution: Das Mortes River in Mato Grosso State, Brazil (Ferraris, 2003d).
Tatia strigata Soares-Porto, 1995
Tatia strigata Soares-Porto, 1995: 202, fig. 1. Type locality: Brazil, Amazonas, Municipio de Maués, IgarapéLimãozinho. Holotype: MZUSP 44065.
Distribution: Amazon River basin and upper reaches of the Orinoco River (Ferraris, 2003d).
TETRANEMATICHTHYS Bleeker, 1858
Tetranematichthys Bleeker, 1858b: 357, 359. Type species: Ageneiosus quadrifilis Kner, 1857. Type by monotypy.
Gender: Masculine.
Tetranematichthys quadrifilis (Kner, 1857)
Ageneiosus quadrifilis Kner, 1857: 442, pl. 9 (fig. 29). Type locality: Rio Guaporé. Holotype: NMW 43343.
Distribution: Upper Guaporé River; reports of specimens from other localities in the Amazon and Orinoco river
basins refer to a currently unnamed species.
TOCANTINSIA Mees, 1974
Tocantinsia Mees, 1974: 108. Type species: Tocantinsia depressa Mees, 1974. Type by original designation. Gender:
Feminine.
Tocantinsia piresi (Miranda Ribeiro, 1920)
Glanidium piresi Miranda Ribeiro, 1920: 14, pls. 15–17. Type locality: Rio S. Manoel [Mato Grosso, Brazil]. MNRJ
264.
Tocantinsia depressa Mees, 1974: 108, fig. 30. Type locality: Tocantins near Porto National, Goyaz, Brazil. Holotype: CAS 6573.
Distribution: Upper Tocantins River basin (Ferraris, 2003d).
Remarks: Synonymy follows Mees (1984).
TRACHELYICHTHYS Mees, 1974
Trachelyichthys Mees, 1974: 111. Type species: Trachelyichthys decaradiatus Mees, 1974. Type by original designation. Gender: Masculine.
Trachelyichthys decaradiatus Mees, 1974
Trachelyichthys decaradiatus Mees, 1974: 112, fig. 31. Type locality: Karanambo area, Rupununi, British Guiana.
Holotype: BMNH 1971.7.29.30.
Distribution: Rupununi River basin, Guyana (Ferraris, 2003d).
Trachelyichthys exilis Greenfield & Glodek, 1977
Trachelyichthys exilis Greenfield & Glodek, 1977: 49, fig. 2. Type locality: Río Mamón, a tributary of Río Nanay,
approximately 25 km west of Iquitos, 3º42'S, 73º16'W, Peru. Holotype: FMNH 80476.
Distribution: Nanay River basin in upper Amazon River drainage (Ferraris, 2003d).
TRACHELYOPTERICHTHYS Bleeker, 1862
Trachelyopterichthys Bleeker, 1862 (in Bleeker, 1862–63): 16. Type species: Trachelyopterus taeniatus Kner, 1857.
Type by original designation. Gender: Masculine.
Trachelyopterichthys anduzei Ferraris & Fernandez, 1987
Trachelyopterichthys anduzei Ferraris & Fernandez, 1987: 257, fig. 1. Type locality: Laguna de Carida at the mouthof Caño Carida, Río Orinoco, Terr. Federal Amazonas, Venezuela. Holotype: MBUCV V-14627.
Distribution: Upper Orinoco River basin (Ferraris, 2003d).
Trachelyopterichthys taeniatus (Kner, 1857)
Trachelyopterus taeniatus Kner, 1857: 434, pl. 8 (fig. 26). Type locality: Rio Guaporé [Brazil]. Holotype: NMW
43346.

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FERRARISDistribution: Upper Amazon River basin (Ferraris, 2003d).
TRACHELYOPTERUS Valenciennes, 1840
Trachelyopterus Valenciennes, in Cuvier and Valenciennes, 1840b: 220 (163 in Strasbourg deluxe edition). Typespecies: Trachelyopterus coriaceus Valenciennes, 1840. Type by monotypy. Gender: Masculine.
Parauchenipterus Bleeker, 1862 (in Bleeker, 1862–63): 7. Type species: Silurus galeatus Linnaeus, 1766. Type byoriginal designation. Gender: Masculine.
Review: Bertoletti et al. (1992), southeastern Brazil.
Trachelyopterus albicrux (Berg, 1901)
Trachycorystes albicrux Berg, 1901: 303. Type locality: Río de la Plata, cerca de la embocadura del Río Santiago,
Buenos Aires. Holotype: MACN 5161.
Distribution: La Plata River basin (Ferraris, 2003d).
Trachelyopterus amblops (Meek & Hildebrand, 1913)
Felichthys amblops Meek & Hildebrand, 1913: 77. Type locality: Río Tuyra, Marriganti, Panama. Holotype: FMNH
7576.
Distribution: Tuira River basin, Panama (Ferraris, 2003d).
Trachelyopterus ceratophysus (Kner, 1857)
Auchenipterus ceratophysus Kner, 1857: 427, pl. 7 (fig. 23). Type locality: Matogrosso am Rio Guaporé, Rio
Branco, und Rio Negro. Syntypes: NMW 47390–93 (4), NMW 50585 (1).
Distribution: Guaporé, Branco and Negro River basins (Ferraris, 2003d).
Trachelyopterus coriaceus Valenciennes, 1840
Trachelyopterus coriaceus Valenciennes, in Cuvier & Valenciennes, 1840b: 221 (164 in Strasbourg deluxe edition),pl. 438. Type locality: [Unknown, apparently from vicinity of Cayenne, French Guiana]. Holotype: MNHN b0242.
Trachelyopterus coriaceus maculosus Eigenmann & Eigenmann, 1888b: 157. Type locality: Porto do Moz [Pará,
Brazil]. Holotype: MCZ 7337.
Distribution: Coastal rivers of Guianas and Brazil and Amazon River basin (Ferraris, 2003d).
Trachelyopterus fisheri (Eigenmann, 1916)
Trachycorystes fisheri Eigenmann, 1916: 82. Type locality: Río Sucio [Colombia]. Holotype: FMNH 57695; illustrated in Eigenmann (1922: 48, pl. 5, fig. 1).
Distribution: Sucio River basin, Colombia (Ferraris, 2003d).
Trachelyopterus galeatus (Linnaeus, 1766)
Silurus galeatus Linnaeus, 1766: 503. Type locality: in America australi. Syntypes: On Seba (1734–65: 85, pl. 19,
fig. 7) and at least one additional specimen which has not been located.
Auchenipterus maculosus Valenciennes, 1840b: 216 (161 in Strasbourg deluxe edition). Type locality: Cayenne.
Syntypes: MNHN a-8856 (1), ?RMNH D3007; MNHN a-8856 illustrated in Royero & Hureau (1996: fig. 2).
Parauchenipterus paseae Regan, 1906c: 387, pl. 24. Type locality: Caroni River, Trinidad. Holotype: BMNH
1906.6.23.48.
Distribution: Widespread in northern South America (Ferraris, 2003d).
Trachelyopterus insignis (Steindachner, 1878)
Auchenipterus insignis Steindachner, 1878a: 89. Type locality: Magdalenen-Stromes [Colombia]. Syntypes: at
NMW. Illustrated and described in more detail in Steindachner (1879d: 35, pl. 3, figs. 2, 2a).
Auchenipterus Magdalenae Steindachner, 1878a: 89. Type locality: Magdalenen-Stromes [Colombia]. Syntypes:
NMW 47409–12 (4). Illustrated and described in more detail in Steindachner (1879d: 36, pl. 4, figs. 1, 1a).
Distribution: Lower Magdalena River basin, Colombia (Ferraris, 2003d).
Remarks: The name Auchenipterus insignis given precedence over A. magdalenae by first reviser action of Miles(1947: 73). Redescribed, as Trachycorystes magdalenae, in Schmidt (1928).
Trachelyopterus lacustris (Lütken, 1874)
Auchenipterus lacustris Lütken, 1874c: 30. Type locality: in flumine Rio das Velhas et in lacu Lagoa Santa. Syntypes: MNHN 0000-9572 (1), NMW 47402 (1), ZMB 9179 (1), ZMUC P 29665 (1), ZMUC P 29666 (1),CHECKLIST OF CATFISHES, RECENT AND FOSSIL

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ZMUC P 29667 (1), ZMUC P 29668 (1).
Distribution: Das Velhas River and Lagoa Santa basins (Ferraris, 2003d).
Trachelyopterus leopardinus (Borodin, 1927)
Trachycorystes leopardinus Borodin, 1927b: 3, fig. 2. Type locality: Rio S. Francisco, Minas Geraes [sic] Province,
Brazil. Holotype: AMNH 8637.
Distribution: São Francisco River basin, Brazil (Ferraris, 2003d).
Trachelyopterus lucenai Bertoletti, da Silva & Pereira, 1995
Trachelyopterus lucenai Bertoletti, da Silva & Pereira, 1995: 71, fig. 2–3. Type locality: Rio Jacuí at saco do Quilombo, ilha das Flores, Porto Alegre, Rio Grande do Sul, Brazil (approximately 29º58'S, 51º15'W). Holotype:
MCP 17174.
Distribution: Lowland tributaries of Uruguay River and Laguna dos Patos basins, Brazil (Bertoletti et al., 1995).
Trachelyopterus peloichthys (Schultz, 1944)
Trachycorystes insignis peloichthys Schultz, 1944c: 236, pl. 4 (fig. a). Type locality: Río Agua Caliente, 2 to 3 km
above Lake Maracaibo, Venezuela. Holotype: USNM 121281.
Distribution: Maracaibo River basin (Ferraris, 2003d).
Trachelyopterus striatulus (Steindachner, 1877)
Auchenipterus (Pseudauchenipterus) striatulus Steindachner, 1877b: 656, pl. 5. Type locality: Rio Parahyba, Rio
Doce, Rio Mucury. Syntypes: MCZ 25506 (1), NMW 10852 (1), NMW 47445 (5), NMW 47446 (2), NMW
47447 (1), SMNS 1955 (1).
Distribution: Coastal rivers in southeastern Brazil (Ferraris, 2003d).
Trachelyopterus teaguei (Devincenzi, 1942)
Trachycorystes teaguei Devincenzi in Devincenzi & Teague, 1942: 37, pl. 7 (figs. 1–2). Type locality: Río Queguay,
Uruguay. Holotype: MHNM CI 350.
Distribution: Queguay River basin in lower Uruguay River drainage (Ferraris, 2003d).
Species inquirendae, Trachelyopterus
Auchenipterus immaculatus Valenciennes, 1840b: 218 (162 in Strasbourg deluxe edition). Type locality: Cayenne. Holotype: RMNH 3008. Comments on type locality in Boeseman (1972: 302).
Auchenipterus robustus Günther, 1864: 197. Type locality: Demerara [Guyana]. Holotype: BMNH 2005.9.12.3.
Auchenipterus glaber Steindachner, 1877b: 655, footnote. Type locality: Demerara [Guyana]. Holotype: at NMW.
Auchenipterus isacanthus Cope, 1878: 677. Type locality: Peruvian Amazon. Syntypes: ANSP 21444–45 (2).
Auchenipterus brevibarbis Cope, 1878: 676. Type locality: Peruvian Amazon. Holotype: ANSP 21519.
Trachycorystes analis Eigenmann & Eigenmann, 1888b: 156. Type locality: Arary? [possibly = Lago Arari, Pará, Brazil]. Holotype: MCZ 7354.
Trachycorystes insignis badeli Dahl, 1955: 13, fig. on p. 14. Type locality: Cereté [Sinu River, Colombia]. Holotype:
Whereabouts unknown (possibly at ICNMHN).
Trachycorystes jokeannae Hoedeman, 1961: 138. Type locality: French Guiana, Ile de Cayenne, ruisseaux du Rorota.
Holotype: ZMA 102371.
TRACHYCORYSTES Bleeker, 1858
Trachycorystes Bleeker, 1858b: 200, 208. Type species: Auchenipterus trachycorystes Valenciennes, 1840. Type by
absolute tautonymy. Gender: Masculine.
Trachycorystes cratensis Miranda Ribeiro, 1937
Trachycorystes cratensis Miranda Ribeiro, 1937: 55. Type locality: Rio Granjeiro, Crato, Ceará [Brazil]. Holotype:
MNRJ 947.
Distribution: Granjeiro River basin in Ceará State, Brazil (Ferraris, 2003d).
Trachycorystes trachycorystes (Valenciennes, 1840)
Auchenipterus trachycorystes Valenciennes, in Cuvier & Valenciennes, 1840b: 214 (159 in Strasbourg deluxe edition), pl. 437. Type locality: Brésil. Holotype: MNHN a-9422; illustrated in Mees (1974: pl. 1).

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Trachycorystes typus Bleeker, 1862 (in Bleeker, 1862–63): 6. Type locality: Bresil. Holotype: MNHN a-9422.
Replacement for Auchenipterus trachycorystes Valenciennes, 1840, presumably to avoid tautonymy, therefore
taking the same type.
Auchenipterus obscurus Günther, 1863: 442. Type locality: the Essequibo, British Guiana. Syntypes: BMNH
1864.1.21.13–14 (2); one syntype illustrated in Eigenmann (1912: pl. 17, fig. 2).
Distribution: Amazon River basin, Essequibo River basin, Guyana, and Orinoco River basin, Venezuela (Ferraris,
2003d).
Species inquirendae, Auchenipteridae
Arius obesus Jardine, in Schomburgk, 1841: 174. Type locality: Rio Branco. No types known.
Pimelodus cærulescens La Cepède, 1803: 95, 103. Type locality: Cayenne. No types known. Name spelled Pimelodus
coerulescens on p. 95; Pimelodus caerulescens on 103.
Trachycorystes porosus Eigenmann & Eigenmann, 1888b: 154. Type locality: Brazil. Syntypes (2, 13–15 cm): MCZ
7351 (2).

AUSTROGLANIDIDAE Mo, 1991
Austroglanididae Mo, 1991: 160. Type genus: Austroglanis Skelton, Risch & de Vos, 1984.
1 genus, 3 species; no named fossil taxa.
AUSTROGLANIS Skelton, Risch & de Vos, 1984
Austroglanis Skelton, Risch & de Vos, 1984: 361. Type species: Gephyroglanis sclateri Boulenger, 1901. Type byoriginal designation. Gender: Masculine.
Austroglanis barnardi (Skelton, 1981)
Gephyroglanis barnardi Skelton, 1981: 220, figs. 2, 4–6, 8–21. Type locality: Noordhoeks River at roadbridge, tributary of Olifants River, Cape Province, S. Africa, 32º43'15"S, 19º03'59"E. Holotype: AMG P7647(a).
Distribution: Streams in Clanwilliam-Olifants River system, South Africa (Skelton, 1993).
Austroglanis gilli (Barnard, 1943)
Gephyroglanis gilli Barnard, 1943: 227, fig. 23 c–g. Type locality: Olifants River system, Clanwilliam Division: inan irrigation furrow on the Jan Diesels River in Bosch Kloof, Clanwilliam; Boontjes River, Citrusdal; upper
reaches of Olifants River at the farm “Keerom” and in a side tributary of the farm “Noordhoek.” Lectotype:
SAM 29231 [now at AMG], designated by Skelton (1981: 267).
Distribution: Streams in Clanwilliam-Olifants River system (Skelton, 1993).
Austroglanis sclateri (Boulenger, 1901)
Gephyroglanis sclateri Boulenger, 1901g: 228, pl. 13. Type locality: Vaal River, South Africa. Syntypes: BMNH
1901.2.11.23 (1), SAM 4708 [now at AMG] (1).
Distribution: Orange-Vaal River basin, in major tributaries and main river channel (Skelton, 1993).

BAGRIDAE Bleeker, 1858
Bagrichthyoidei Bleeker, 1858b: 49, 54. Type genus: Bagrus Bosc, 1816.
Bagrichthyes Bleeker, 1862 (in Bleeker, 1862–63): 8, 48. Type genus: Bagrichthys Bleeker, 1857.
Ritae Bleeker, 1862 (in Bleeker, 1862–63): 8. Type genus: Rita Bleeker, 1854.
Olyrinae Gill, 1893: 132. Type genus: Olyra M’Clelland, 1842.
Porcinae Fowler, 1915a: 219. Type genus: Porcus Geoffroy St. Hilaire, 1809. Based on a generic name that was placedon the Official Index of Rejected and Invalid Generic Names in Zoology, Opinion 1402 (Melville & Smith, 1987).
Batasinae Tilak, 1967: 431. Type genus: Batasio Blyth, 1860.
Pelteobagrini Jayaram, 1966b: 1071. Type genus: Pelteobagrus Bleeker, 1865.

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Bagroidinae Jayaram, 1968b: 371. Type genus: Bagroides Bleeker, 1851.
Taxonomic summaries: Risch (1986, Africa); Chu et al. (1999, China).
Reviews: Jayaram (1955a, genera); Jayaram (1966c, various Asian genera); Jayaram (1966b, African genera); Jayaram(1968b, Japanese, Chinese, Malayan and Indonesian genera); Jayaram (1978a, Chinese species of “Mystus”); Risch(1992b, West Africa); Desoutter (1975, Cambodia); Lee & Kim (1990, Korea).
Identification guides: Jayaram (1977d, 1979 South Asia).
Phylogeny: Jayaram (1976); Jayaram & Singh (1982); Mo (1991); Peng et al. (2002, East Asian species); Ng (2003j).
23 genera, 189 species; including 4 fossil genera and 11 fossil species.
Incertae sedis
Pseudobagrus kyphus Mai, 1978
Pseudobagrus kyphus Mai, 1978: 261, fig. 119. Type locality: [Ky Phu stream in Dai tu district, Thai Nguyen Province, Northern Vietnam]. Holotype: at DVZUT.
Distribution: Vu Quang, northern Vietnam (Kottelat, 2001a), Guangxi Zhuangzu Autonomous Region, China(Watanabe et al., 2002).
Remarks: Currently treated as valid as Pseudobagrus kyphus or Pelteobagrus kyphus. Redescribed in Watanabe, etal. (2002).
Pseudobagrus nitidus Sauvage & Dabry de Thiersant, 1874
Pseudobagrus nitidus Sauvage & Dabry de Thiersant, 1874: 5. Type locality: Yang-tze-kiang, China. Type(s) (130mm): at MNHN.
Distribution: from Heilongjiang [Amur River] to Minjiang [Yangtze River tributary] (Chu et al., 1999).
Remarks: Currently treated as valid as Pelteobagrus nitidus or Leiocassis nitidus.
† Brachyspondylus saropterix Marck, 1876a: 411, pl. 23, fig. 2. Type locality: Sumatra; Tertiary. Holotype: at Royal
Geol. Mus. Dresden (Posterior end of fish, without Weberian complex, pectoral girdle or head).
Distribution: Sumatra (Gayet & Otero, 1999).
Remarks: Treated as valid in the Bagridae S.L. by Gayet & Meunier (2003), but the generic name † Brachyspondylus does not have a siluriform taxon as its type; therefore this species is without a valid generic name. Sanders(1934: 28) placed the species into Aoria, but that name is preoccupied in Insecta.
BAGRICHTHYS Bleeker, 1857
Bagrichthys Bleeker, 1857b: 473. Type species: Bagrus hypselopterus Bleeker, 1852. Type by monotypy. Gender:
Masculine.
Pseudobagrichthys Bleeker, 1862 (in Bleeker, 1862–63): 9, 49. Type species: Bagroides macropterus Bleeker, 1853.
Type by original designation. Gender: Masculine.
Key: Linder & Ng (2002a).
Bagrichthys hypselopterus (Bleeker, 1852)
Bagrus hypselopterus Bleeker, 1852d: 588. Type locality: Palembang, in fluviis. Holotype (260 mm TL): possibly
RMNH 6877.
Distribution: Sumatra and Borneo (Roberts, 1989a).
Bagrichthys macracanthus (Bleeker, 1854)
Bagroïdes macracanthus Bleeker, 1854a: 88. Type locality: Sumatra orientalis, ubi confluunt flumina Lamatang et
Emin. Holotype (240 mm TL): RMNH 5216 (1 of 27); illustrated in Bleeker (1862–63: 50, pl. 67 [=Silur. pl.
19], fig. 1) as Pseudobagrichthys macracanthus.
Distribution: Mekong and Chao Phraya River basins and Sundaland (Kottelat, 2001b).
Bagrichthys macropterus (Bleeker, 1853)
Bagroïdes macropterus Bleeker, 1853b: 515. Type locality: Moara kompeh, in fluviis [Sumatra]. Holotype (256 mm
TL): Whereabouts unknown.
Distribution: Sumatra.
Remarks: As Bagroides macropterus in some recent literature.

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FERRARISBagrichthys majusculus Ng, 2002
Bagrichthys majusculus Ng, 2002a: 50, fig. 1. Type locality: Thailand: Ubon Ratchathani Province, Khong Chiam
district, Mun River at Ban Dan, 3 km upstream from Mekong River confluence. Holotype: UMMZ 214916.
Distribution: Mekong and Chao Phraya River basins (Ng, 2002a).
Bagrichthys micranodus Roberts, 1989
Bagrichthys micranodus Roberts, 1989a: 115, fig. 89. Type locality: Kapuas mainstream 58 km NE of Sintang and 1
km downstream from Sebruang, Western Borneo (Kalimantan Barat, Indonesia). Holotype: MZB 3578.
Distribution: Kapuas River, Western Borneo (Roberts, 1989a).
Bagrichthys obscurus Ng, 1999
Bagrichthys obscurus Ng, 1999f: 546, figs. 1–2. Type locality: Thailand: Roi Et Province, Lam Chi, 1.5 km below
highway 23 bridge, 4 km W. of Selaphum. Holotype: USNM 317511.
Distribution: Mekong, Bang Pakong and Chao Phraya River basins (Ng, 1999f).
Bagrichthys vaillantii (Popta, 1906)
Leiocassis macropterus Vaillant, 1902: 58, figs. 6–7. Type locality: Tepoe, bords du Mahakam, Bornéo central.
Holotype: Whereabouts unknown. Considered to have been preoccupied in Bagroides by Bagroides macropterus Bleeker, 1853; replaced by Bagroides vaillantii Popta, 1906.
Bagroides Vaillantii Popta, 1906: 228. Type locality: Tepoe, bords du Mahakam, Bornéo central. Holotype: Whereabouts unknown. Replacement name for Leiocassis macropterus Vaillant, 1902, considered preoccupied in
Bagroides by Bagroides macropterus Bleeker, 1853.
Distribution: Mahakam River basin, eastern Borneo (Ng, 2000).
Remarks: Redescribed in Ng (2000: 327).
BAGROIDES Bleeker, 1851
Bagroïdes Bleeker, 1851d: 204. Type species: Bagroides melapterus Bleeker, 1851. Type by monotypy. Gender:
Masculine.
Bagroides melapterus Bleeker, 1851
Bagroïdes melapterus Bleeker, 1851d: 204. Type locality: Bandjarmassing, in fluviis. Syntypes (2, 142, 200 mm
TL): BMNH 1863.12.4.71 (1), possibly NMV 46011 (1).
Bagroides melanopterus Bleeker, 1852b: 413. Type locality: Bandjarmassing, in fluviis. Syntypes: BMNH
1863.12.4.71 (1), possibly NMV 46011 (1). Unjustified emendation, or incorrect subsequent spelling, forBagroides melapterus Bleeker, 1851.
Distribution: Sumatra and Borneo (Roberts, 1989a).
BAGRUS Bosc, 1816
Porcus Geoffroy Saint-Hilaire, 1809: pl. 15. Type species: Silurus bajad Forsskål, 1775. Type by monotypy. Gender:
Masculine. Name placed on Official Index of Rejected and Invalid Generic Names In Zoology (ICZN, Opinion
1402).
Bagrus Bosc, 1816: 147. Type species: Silurus bajad Forsskål, 1775. Type by subsequent designation by Bailey &Stewart (1983: 168). Gender: Masculine. Placed on Official List (ICZN, Opinion 1402).
Remarks: The names Porcus and Bagrus were each proposed as new independently in several publications; see
Bailey & Stewart (1983) for details. See also Jayaram (1956a) for additional comments on the names. An earlier
type designation of Bagrus may date to Desmarest as Bagrus bayad (see Whitley, 1939: 224), but that action
may have been overridden by ICZN Opinion 1402, which listed Silurus bajad Forsskål as type.
Key (as genus Porcus): Jayaram (1966b).
Bagrus bajad (Forsskål, 1775)
Silurus bajad Forsskål, 1775: 66, xiii. Type locality: in Nilo. No types known. Placed on Official List (ICZN, Opinion 1402) as type species of Bagrus.
Porcus bayad Geoffroy Saint-Hilaire, 1809: pl. 15 (figs. 1–2). Type locality: Nil. Misspelling or unjustified emendation of Silurus bajad Forsskål.
Bagrus bayad macropterus Pfaff, 1933: 287, fig. 4. Type locality: Kabara. Holotype (47 cm): ZMUC P 28667. Orig-CHECKLIST OF CATFISHES, RECENT AND FOSSIL

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inally as Bagrus bayad var. macropterus var. n.
Distribution: Lakes Chad, Mobutu and Turkana; Nile, Niger, and Senegal rivers (Risch, 1986).
Bagrus caeruleus Roberts & Stewart, 1976
Bagrus caeruleus Roberts & Stewart, 1976: 275. Type locality: Near Inga hydroelectric dam. Lat. 5º31.5'S, Long.
13º37.5'E; lower Zaïre or Congo River. Holotype: MCZ 50582.
Distribution: Lower Congo River; known only from type series (Risch, 1986).
Bagrus degeni Boulenger, 1906
Bagrus Degeni Boulenger, 1906b: 438. Type locality: Entebbe, Lake Victoria. Syntypes (7): BMNH 1906.5.30.172(1), BMNH 1906.5.30.173–177 (5).
Distribution: Lake Victoria (Risch, 1986).
Bagrus docmak (Forsskål, 1775)
Silurus docmak Forsskål, 1775: 65, xiii. Type locality: in Nilo inferiori ad Deltam. No types known.
Bagrus koenigi Pietschmann, 1932: 92, fig. 1. Type locality: In Oberägypten im Nil. Holotype: at ZFMK.
Bagrus docmac niger Daget, 1954: 244, fig. 89. Type locality: Bamako [Niger River]. Syntypes: MNHN 1954-0004(1), MNHN 1960-0465 (2), MNHN 1960-0466 (1).
Distribution: Nile, Niger, Senegal and Volta rivers; Lack Chad basin, East African rift lakes, except Lake Kivu
(Risch, 1986).
Bagrus filamentosus Pellegrin, 1924
Bagrus filamentosus Pellegrin, 1924a: 462. Type locality: Cercle de Ségou (du Niger). Holotype: MNHN 19240215.
Distribution: Niger River system (Risch, 1986).
Bagrus lubosicus Lönnberg, 1924
Bagrus lubosicus Lönnberg, 1924: 619. Type locality: Lubosi river, a tributary to Luala River, Lower Congo. Holotype: NRM 9939.
Distribution: Lower Congo River system (Risch, 1986).
Bagrus meridionalis Günther, 1894
Bagrus meridionalis Günther, 1894: 626. Type locality: Upper Shire River [British Central Africa]. Holotype (22 ½inches): BMNH 1893.11.15.87 (skin).
Distribution: Shire River and Lake Malawi (Risch, 1986).
Bagrus orientalis Boulenger, 1902
Bagrus orientalis Boulenger, 1902d: 37. Type locality: du Pangani, Côte de Zanzibar. Syntypes (2): BMNH
1865.9.21.17 (1), BMNH 1867.3.7.488 (1). Name made available in key.
Distribution: Pangani River, Tanzania (Risch, 1986).
† Bagrus shuwaiensis Forey & Young, 1999
† Bagrus shuwaiensis Forey & Young, 1999: 125, figs. 12.6–12.16. Type locality: Late Miocene, Bahnunah Formations, west side of Shuwaihat, Abu Dhabi. Holotype: AUH 678 (posterior portion of cranium).
Distribution: Bahnunah Formations, west side of Shuwaihat, Abu Dhabi.
Remarks: Considered to be Bagridae S.L. by Gayet & Meunier (2003).
Bagrus ubangensis Boulenger, 1902
Bagrus ubangensis Boulenger, 1902d: 38, pl. 11 (fig. 1). Type locality: l’Ubangi à Banzyville. Holotype: MRAC
1295.
Distribution: Congo River system (Risch, 1986).
Bagrus urostigma Vinciguerra, 1895
Bagrus urostigma Vinciguerra, 1895: 36. Type locality: Ganana, Giuba [Somali]. Syntypes (4): MSNG 14598 (1,dry), MSNG 14599 (1), MSNG 14600 (1).
Distribution: Giuba River, Somalia (Risch, 1986).
BATASIO Blyth, 1860
Batasio Blyth, 1860: 149. Type species: Batasio buchanani Blyth, 1860. Type by original designation. Type designated as “B. Buchanani, nobis; Pimelodus batasio, B.H.” Gender: Masculine.

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Macronoides Hora, 1921a: 179. Type species: Possibly Batasio affinis Blyth, 1860. Type by subsequent designation
by Jordan (1923: 148). Gender: Masculine. Originally proposed as a subgenus of Macrones.
Revision: Hora & Law (1941).
Review: Ng & Kottelat (2001, Indochinese species)
Remarks: Systematic position reviewed in Tilak (1967) and Jayaram (1977b). Type species for Macronoides takenfrom Misra (1976: 215) although no type designation has been found; type species was not designated in Hora
& Law (1941).
Batasio affinis Blyth, 1860
Batasio affinis Blyth, 1860: 150. Type locality: Tenasserim [Myanmar]. Holotype: ZSI F7880/1 [? = A.549].
Macrones Blythii Day, 1877 (in Day, 1875–78): 445. Type locality: Tenasserim provinces. Holotype: ZSI F7880/1 [?
= A.549]. Proposed as a replacement for Batasio affinis Blyth, 1860 when thought to be preoccupied in
Macrones by Bagrus affinis Jerdon, 1849 [see Ng & Kottelat (2001: 500) for discussion].
? Leiocassis fluviatilis Day, 1888: 805. Type locality: The Anin, a stream rising near Weywoon, Wagroo, in the Tenasserim district [Myanmar]. Syntypes (4): Whereabouts unknown.
Distribution: Eastern Myanmar (Ng & Kottelat, 2001).
Batasio batasio (Hamilton, 1822)
Pimelodus batasio Hamilton, 1822: 179, 377. Type locality: River Tista [Ganges River]. No types known. Previously unpublished Hamilton illustration of a type specimen reproduced in Hora (1929: pl. 22, fig. 3).
Batasio Buchanani Blyth, 1860: 150. Type locality: River Tista [Ganges River]. No types known. Unneeded
replacement name for Pimelodus batasio Hamilton, 1822.
Distribution: Ganges River basin, India.
Remarks: Hamilton’s plate (pl. 23, fig. 60), labeled Pimelodus batasius and referred to in the account of Pimelodus
batasio, has been shown by Day (1871) and Hora (1929: 182) to fit instead the description of Pimelodus carcio
Hamilton.
Batasio dayi (Vinciguerra, 1890)
Macrones Dayi Vinciguerra, 1890: 230, pl. 7 (fig. 3). Type locality: Meetan; M.ti a Est di Toungoo (Carin Biapb),
Birmania [Myanmar]. Syntypes (6): BMNH 1893.2.16.8 (1), MSNG 14586 (1), NMW 45184 (1), RMNH
10225 (1).
Distribution: Salween and Irrawaddy River basins, Myanmar (Ng & Kottelat, 2001).
Batasio elongatus Ng, 2004
Batasio elongatus Ng, 2004c: 68, fig. 1. Type locality: Myanmar, Rakhine state: Baw Di Chaung, drainage, Baw Di
Chaung, at Baw Di bridge, 20 miles from Gwa on road to Ngathaingchaung (17º34'15"N 94º43'47"E). Holotype: NRM 40748.
Distribution: Baw Di Chaung and Kanamae Chaung basins, western Rakihine Yoma, Myanmar (Ng, 2004c).
Batasio havmolleri (Smith, 1931)
Mystus havmolleri Smith, 1931: 24, fig. 12. Type locality: Klong Thalerng near Ronpibun, Peninsular Siam. Holotype: USNM 90304.
Mystus stigmaturus Fowler, 1934a: 94, figs. 41–42. Type locality: Nakon Sritamarat, Peninsular Siam. Holotype:
ANSP 59338.
Distribution: Malay Peninsula, south of isthmus of Kra and north of Perak River basin and Terengganu River (Ng &Kottelat, 2001).
Batasio macronotus Ng & Edds, 2004
Batasio macronotus Ng & Edds, 2004: 296, fig. 1. Type locality: Nepal: Saptari/Sunsari, Kosi River drainage, purchased at Kosi barrage, 26º31'30"N 86º56'00"E. Holotype: KU 34546.
Distribution: Kosi River basin, eastern Nepal (Ng & Edds, 2004).
Batasio merianiensis (Chaudhuri, 1913)
Macrones merianiensis Chaudhuri, 1913: 253, pl. 9 (figs. 1, 1a, 1b). Type locality: Pond at Mariani junction, Assam[India]. Holotype: ZSI F7781/1.
Distribution: Assam, India.
Batasio pakistanicus Mirza & Jan, 1989

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Batasio pakistanicus Mirza & Jan, 1989: 283, figs. 1, 2. Type locality: Jinnah Barrage (River Indus) [Pakistan].
Holotype: GCM 17-F.
Distribution: Punjab, Indus River basin, Pakistan (Mirza, 2003).
Remarks: Treated as a synonym of Batasio batasio by Jayaram (1999).
Batasio sharavatiensis Bhatt & Jayaram, 2004
Batasio sharavatiensis Bhatt & Jayaram, 2004: 1339. Type locality: River Sharavati (14º14'N, 74º49'E), Joginmatha, Uttara Kannada, Karnataka [India]. Holotype: ZSI/SRS F 6419.
Distribution: Sharavati River, Uttara Kannada, Karnataka State, India (Bhatt & Jayaram, 2004).
Batasio tengana (Hamilton, 1822)
Pimelodus tengana Hamilton, 1822: 176, 377, pl. 39 (fig. 58). Type locality: Brahmaputra River. No types known.
Distribution: Brahmaputra River, India and Bangladesh.
Batasio tigrinus Ng & Kottelat, 2001
Batasio tigrinus Ng & Kottelat, 2001: 504, fig. 6. Type locality: Thailand: Kanchanaburi Province, Mae Nam
Khwae Noi basin, Huai Lia, km 49 on road from Thon Pha Phun to Sangkhla (15º04'25"N, 98º33'51"E). Holotype: ZRC 40624.
Distribution: Mae Nam Khwae Noi basin, Mae Khlong drainage, western Thailand (Ng & Kottelat, 2001).
Batasio travancoria Hora & Law, 1941
Batasio travancoria Hora & Law, 1941: 40, pl. 2 (figs. 7–9); fig. 3. Type locality: Peruntenaruvi, a tributary of the
Pamba River at Edakadathy [Travancore, India]. Holotype: ZSI F13449/1.
Distribution: Pamba, Kallada and Chittar River basins, Anaimalai Hill Range, Kerala, India (Jayaram, 1999).
CHANDRAMARA Jayaram, 1973
Chandramara Jayaram, 1973a: 816. Type species: Pimelodus chandramara Hamilton, 1822. Type by original designation. Gender: Masculine.
Chandramara chandramara (Hamilton, 1822)
Pimelodus chandramara Hamilton, 1822: 162, 375. Type locality: Atreyi River [India]. No types known. Previously
unpublished Hamilton illustration of a type specimen reproduced in Hora (1929: pl. 21, fig. 6).
Ageniosus (Silonia) diaphina Swainson 1839: 306. Type locality: Atreyi River [India]. No types known. On “Ham.
P. 162” [= Hamilton (1822: 162)]. Unneeded new name for Pimelodus chandramara Hamilton, 1822.
Distribution: Base of eastern part of Himalaya, and Sarda River basin (Tilak, 1987).
Remarks: Also treated in recent literature as valid as Rama chandramara.
COREOBAGRUS Mori, 1936
Coreobagrus Mori, 1936: 672, 675. Type species: Coreobagrus brevicorpus Mori, 1936. Type by monotypy. Gender: Masculine.
Remarks: Species diagnosed in Watanabe et al. (1992).
Coreobagrus brevicorpus Mori, 1936
Coreobagrus brevicorpus Mori, 1936: 672, pl. 24 (fig. 2). Type locality: River Rakuto at Ei-yo [but, Bun-kei elsewhere in text], Chosen [Korea]. Holotype: At Preparatory Department of Keijo Imperial University.
Distribution: Naktong River and Kim River, Korea (Mori, 1952).
Remarks: Treated in some recent literature as valid in Pseudobagrus.
Coreobagrus ichikawai Okada & Kubota, 1957
Coreobagrus ichikawai Okada & Kubota, 1957: 144, unnumbered figure. Type locality: Miya River, Mie Prefecture,
Japan. Holotype: FRLM 6230.
Coreobagrus okadai Jayaram, 1966a: 30, fig. 1. Type locality: Upper region of Suzuka River, Mie Prefecture, Honshu, Japan. Holotype: at ZSI.
Distribution: In the vicinity of Ise and Mikawa bays, Honshu, Japan (Katsuura et al., 2000).
Remarks: Treated in some recent literature as valid as Pseudobagrus ichikawai. See Watanabe (1998) for intraspecific variation, and Watanabe & Ito (1999) for comments on distribution.

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† Macronoides White, 1934: 54. Type species: † Macronoides wilsoni White, 1934. Type by original designation.
Gender: Masculine. Preoccupied by Macronoides Hora, 1921, in Recent fishes. Replaced by † Eomacrones
White, 1937.
† Eomacrones White, 1937: 144. Type species: † Macronoides wilsoni White, 1934. Type by being a replacement
name. Gender: Masculine. Replacement for † Macronoides White, 1934, preoccupied by Macronoides Hora,
1921.
Remarks: Considered to be Bagridae (sensu lato) by Gayet & Meunier (2003).
† Eomacrones wilsoni (White, 1934)† Macronoides wilsoni White, 1934: 54, fig. 13. Type locality: Gada, Nigeria, in scree; Eocene. Holotype: at
National Collection [Nigeria] or BMNH (incomplete skull).
† GOBIBAGRUS Sytchevskaya, 1986
† Gobibagrus Sytchevskaya, 1986: 90. Type species: †Gobibagrus hoerdzanicus Sytchevskaya, 1986. Type by original designation. Gender: Masculine.
† Eobagrus Sytchevskaya, 1986: 92. Type species: †Gobibagrus hoerdzanicus Sytchevskaya, 1986. Type by original
designation. Gender: Masculine. Apparently a lapsus calami for Gobibagrus.
Remarks: Type species cited as †Gobibagrus hoerdzanicus, but species account lists name as †Eobagrus hoerdzanicus. Assigned to the family Bagridae by Sytchevskaya, 1986.
† Gobibagrus hoerdzanicus Sytchevskaya, 1986
† Eobagrus hoerdzanicus Sytchevskaya, 1986: 92, pl. 15, figs. 6–10. Type locality: [Mongolia, Lower Oligocene].
Holotype: ПИН N 3874/408 (pectoral spine)
Distribution: Mongolia, upper Miocene (Gayet & Meunier, 2003: 501).
Remarks: Name as † Eobagrus hoerdzanicus in heading of species account, but † Gobibagrus hoerdzanicus on plate
caption and as type species in genus account.
HEMIBAGRUS Bleeker, 1862
Hemibagrus Bleeker, 1862 (in Bleeker, 1862–63): 9. Type species: Bagrus nemurus Valenciennes, 1840. Type byoriginal designation. Gender: Masculine. Placed on Official List (Opinion 2011; ICZN, 2002).
Macropterobagrus Nichols, 1925: 1. Type species: Hemibagrus macropterus Bleeker, 1870. Type by monotypy.
Gender: Masculine. Originally proposed as a subgenus of Hemibagrus.
Brachymystus Fowler, 1937: 148. Type species: Bagrus nemurus Valenciennes, 1840. Type by original designation.
Gender: Masculine. Originally proposed as a subgenus of Mystus.
Review: Ng & Rainboth (1999, central Indochina); Ng & Ferraris (2000, southern Asia).
Hemibagrus baramensis (Regan, 1906)
Macrones baramensis Regan, 1906b: 68. Type locality: Baram River, Sarawak state, Borneo, East Malaysia. Holotype: BMNH 1895.7.2.50.
Distribution: Baram River, Borneo (Regan, 1906b).
Hemibagrus bongan (Popta, 1904)
Macrones bongan Popta, 1904: 182. Type locality: le Bongan, Bornéo central. Syntypes: RMNH 7548 (1), RMNH
7549 (1); described in more detail, with an illustration of one of the syntypes, in Popta (1906: 35, pl. 2, fig. 5).
Distribution: Kapuas, Rajang and Baram River drainages, northern Borneo (Ng & Rachmatika, 1999).
Hemibagrus caveatus Ng, Wirjoatmodjo & Hadiaty, 2001
Hemibagrus caveatus Ng, Wirjoatmodjo & Hadiaty, 2001b: 359, fig. 1. Type locality: Sumatra: Aceh Province, Sungai Soraya, a tributary of Sungai Alas [Indonesia]. Holotype: MZB 8714.
Distribution: Alas River basin, northern Sumatra (Ng, et al., 2001b).
Hemibagrus centralus Mai, 1978
Hemibagrus centralus Mai, 1978: 253, fig. 115 [not 116]. Type locality: Northern Vietnam. Syntypes: at DVZUT.
Distribution: Northern Vietnam (Kottelat, 2001b).
Hemibagrus chrysops Ng & Dodson, 1999

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Zootaxa 1418 © 2007 Magnolia Press · 87Hemibagrus chrysops Ng & Dodson, 1999: 48, figs. 1, 2a, 3c–d. Type locality: Borneo: Sarawak, Serian wet market,
from Sadong River. Holotype: ZRC 42653.
Distribution: Sadong and Rajang River basins, northern Borneo (Ng & Dodson, 1999).
Hemibagrus filamentus (Fang & Chaux, 1949)
Macrones (Hemibagrus) filamentus Fang & Chaux, in Chaux & Fang, 1949a: 200, fig. 4. Type locality: Cambodge.
Holotype: MNHN 1966-0728.
Distribution: Middle and lower Mekong River basin of Laos, Thailand, and Cambodia (Ng & Rainboth, 1999).
Hemibagrus fortis (Popta, 1904)
Macrones fortis Popta, 1904: 185. Type locality: le Bo, Bornéo central. Syntypes: RMNH 7551 (4); described in
more detail, with an illustration of one of the syntypes, in Popta (1906: 44, pl. 2, figs. 8a, 8b).
Macrones bo Popta, 1904: 183. Type locality: le Bo, Bornéo, central. Holotype: RMNH 7549; described in moredetail, with illustration of holotype, in Popta (1906: 38, pl. 2, fig. 6).
Macrones howong Popta, 1904: 181. Type locality: le Howong, Bornéo central. Holotype: RMNH 7547; describedin more detail, with illusration of holotype, in Popta (1906: 32, pl. 1, fig. 4).
Macrones kajan Popta, 1904: 184. Type locality: le Kajan, Bornéo central. Holotype: RMNH 7550; described in
more detail, with illustration of holotype, in Popta (1906: 41, pl. 2, fig. 7). [Possibly synonym of Hemibagrus
nemurus].
Macrones fortis capitulum Popta, 1906: 48, pl. 3 (fig. 9). Type locality: le Bo, Bornéo central. Holotype: RMNH
7552. Originally as Macrones fortis var. capitulum. [First mentioned in Popta (1904: 186) as Macrones fortis
var.]Distribution: Borneo.
Remarks: Tan & Ng (2000: 271) acted as first reviser and gave precedence to Macrones fortis over M. howong, M.
kajan and M. bo.
Hemibagrus furcatus Ng, Martin-Smith & Ng, 2000
Hemibagrus furcatus Ng, Martin-Smith & Ng, 2000: 66, figs. 1, 2a, 3. Type locality: Sungai Segama, Danum Valley
Conservation Area, Lahad Datu dist., Sabah, Borneo. Holotype: SBM uncataloged.
Distribution: Segama River basin, eastern Borneo (Ng, et al., 2000).
Hemibagrus gracilis Ng & Ng, 1995
Hemibagrus gracilis Ng & Ng, 1995: 136, figs. 1–3, 4a. Type locality: Peninsular Malaysia: Johor, Ulu Endau,
Sungei Jasin. Holotype: ZRC 21484.
Distribution: Upper reaches of Sungei Endau and Sungei Kinchin, Endau-Rompin region, Peninsular Malaysia (Ng& Ng, 1995).
Hemibagrus guttatus (La Cepède, 1803)
Pimelodus guttatus La Cepède, 1803: 96, 111, pl. 5 (fig. 1). Type locality: [China]. No types known; description
based on illustration [reproduced in Kottelat et al. (1994)].
Macrones elongatus Günther, 1864: 77. Type locality: Singapore [probably in error, possibly China; see Kottelat etal. (1998: 567)]. Holotype: BMNH 1855.9.19.1099; illustrated in Jayaram (1978b: 223, fig. 2, and Kottelat etal. (1998: 567, fig. 1).
Macrones chinensis Steindachner, 1883: 196. Type locality: Kanton, China. Holotype: at NMW. Species illustrated
and described in more detail in Steindachner (1884: 1111, pl. 8). Holotype also illustrated in Jayaram (1978b:223, fig. 3).
Distribution: Qiantangjiang, Jiulongjiang, Hanjiang, Zhujiang, Yuanjiang, China (Chu et al., 1999); Nam Xam and
Nam Ma Basins, Laos, and Red River (Kottelat, 2001b).
Remarks: Synonymy follows Kottelat et al. (1998).
Hemibagrus hainanensis (Tchang, 1935)
Leiocassis hainanensis Tchang, 1935b: 175, fig. 2. Type locality: Hainan I., China. Holotype: ASIZB [= ZMFMIB
12904].
Distribution: Hainan Island, China.
Hemibagrus hoevenii (Bleeker, 1846)
Bagrus Hoevenii Bleeker, 1846a: 154. Type locality: Batavia [now Sungai Muar at Kampung Bukit Kebong, Johor,

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Distribution: Kapuas and Baram River basins, Borneo, Musi and Batang Hari River basins, Sumatra, and Johor, Paahang, and Muar River basins, western Malaysia (Kottelat & Lim, 1995).
Remarks: Redescribed in Kottelat & Lim (1995).
Hemibagrus hongus Mai, 1978
Hemibagrus elongatus hongus Mai, 1978: 252. Type locality: [Hông River and Lam River] Vietnam. Syntypes: at
DVZUT.
Distribution: Northern Vietnam (Kottelat, 2001a).
Hemibagrus imbrifer Ng & Ferraris, 2000
Hemibagrus imbrifer Ng & Ferraris, 2000: 126, fig. 1. Type locality: Mae Nam Moei at Ban Wan Le, 16º17'24"N,
98º42'21"E, Salween basin, Tak Prov., Thailand. Holotype: ZRC 45406.
Distribution: Salween River basin (Ng & Ferraris, 2000).
Hemibagrus johorensis (Herre, 1940)? Macrones bleekeri Volz, 1903a: 556. Type locality: Brackish water at Banjuasin, Palembang, Sumatra. Holotype:
NMBE 1020756. Preoccupied by Macrones bleekeri Day, 1877. Species illustrated and described in more detailin Volz (1903b: 387, pl. 26 [not 25 as stated], fig. 3).
Mystus johorensis Herre, 1940: 13, pls. 7–8. Type locality: Sungai Kayu, 16 miles north of Kota Tinggi, Johore[Malaysia]. Holotype: SU 33026.
Distribution: Peninsular Malaysia and Sumatra.
Hemibagrus macropterus Bleeker, 1870
Hemibagrus macropterus Bleeker, 1870: 257, unnumbered pl. (bottom fig.). Type locality: [Yangtze River, China].
Holotype: Whereabouts unknown. Often placed in Mystus in Chinese literature.
Distribution: From Changjiang [Yangtze River] to Zhujiang [=Pearl River], China (Chu et al., 1999).
† Hemibagrus major Roberts & Jumnongthai, 1999
† Hemibagrus major Roberts & Jumnongthai, 1999: 168, fig. 14 (tooth only). Type locality: Thailand, North-central,
Phetchabun Province, Ban Nong Pla; Miocene. Holotype: Department of Natural Resources, Thailand (DNR)
TF 5011 (incomplete specimen).
Distribution: North-central, Phetchabun Province, Ban Nong Pla, Thailand; Miocene; known only from type.
Hemibagrus maydelli (Rössel, 1964)
Mystus (Mystus) maydelli Rössel, 1964: 149, fig. 1. Type locality: Indien, Wadgaon, Bhima-Fluss. Holotype: ZMH
H 2180.
Mystus krishnensis Ramakrishniah, 1988: 139, figs. 1–2. Type locality: Nagarjunasagar reservoir, India. Holotype:ZSI FF2532.
Distribution: Middle reaches of Krishna River, peninsular India (Ng & Ferraris, 2000).
Hemibagrus menoda (Hamilton, 1822)
Pimelodus menoda Hamilton, 1822: 203, 379.Type locality: Kosi, Mahananda, and other rivers in north of Behar
and Bengal [now: Bangladesh, Suarma (Meghna) drainage, Sharighat bazaar, 22 miles NE of Sylhet on SylhetShillong hwy, by neotype designation]. Neotype: UMMZ 208726, illustrated in, and designated by, Ng & Ferraris (2000:136, fig. 5). Plate 1 of Hamilton (1822), labelled Mugil corsula, and replaced by Pimelodus in corrigenda, appears to refer to this species.
Bagrus corsula Valenciennes, in Cuvier & Valenciennes, 1840a: 408 (302 in Strasbourg deluxe edition). Type locality: Bengale [now: Bangladesh, Suarma (Meghna) drainage, Sharighat bazaar, 22 miles NE of Sylhet on SylhetShillong hwy by neotype designation]. Neotype: UMMZ 208726, illustrated in, and designated by, Ng & Ferraris (2000:132, fig. 5). Based on illustration in Hamilton (1822: pl. 1) originally labelled as Mugil corsula, butchanged in corrigenda to Pimelodus, which apparently was interpreted by Valenciennes to mean Pimelodus corsula. However, there in no justification for ascribing the name P. corsula to Hamilton, so its first use is that of
Valenciennes.
Bagrus trachacanthus Valenciennes, in Cuvier & Valenciennes, 1840a: 419 (311 of Strasbourg deluxe edition). Type
locality: Bengale. Syntype: MNHN 0000-1191 (1).

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Macrones corsula Day, 1869: 307. Type locality: Mahanuddi River at Cuttack, India. Syntypes (3): Whereabouts
unknown. Apparently intended as a new name, based in part on Hamilton (1822: pl. 1, fig. 72).
Distribution: Ganges, Brahmaputra, Mahanadi and Godavari River basins, Bangladesh and India (Ng & Ferraris,
2000).
Hemibagrus microphthalmus (Day, 1877)
Macrones microphthalmus Day, 1877 (in Day, 1875–78): 446, pl. 100 (fig. 4). Type locality: Burma, along the valleyof the Irrawaddi. Possible syntypes: AMS B.7918 (1), ZSI 2592 (1).
Distribution: Salween River, Thailand; Irrawaddy and Sittang Rivers, Myanmar; and Manipur River, India (Ng &Ferraris, 2000).
Remarks: Roberts & Jumnongthai (1999: 176) and Ng & Rainboth (1999: 576) cite ZSI 2592 as the holotype of thespecies, although Day indicated that multiple specimens were examined.
Hemibagrus nemurus (Valenciennes, 1840)
Bagrus nemurus Valenciennes, in Cuvier & Valenciennes, 1840a: 423 (314 of Strasbourg deluxe edition). Type
locality: Java. Holotype: Possibly RMNH 269 (not found) or at MNHN. Placed on Official List (Opinion 2011)(ICZN, 2002).
Bagrus Sieboldii Bleeker, 1846a: 155. Type locality: Batavia. Syntypes (size and number not stated): RMNH 15881(5), SMNS 10572 (1). Placed on Official List (Opinion 2011) (ICZN, 2002).
Mystus pahangensis Herre, 1940: 14, pl. 9. Type locality: Sungai Garam, near Karak, Pahang [Malaysia]. Holotype:
SU 33025.
Distribution: Java and likely in river basins of neighboring regions.
Remarks: This name has been applied to populations that have been subsequently determined to belong to distinct
species, but the geographic limits of Hemibagrus nemurus have not been clearly stated. Ng & Rainboth (1999:
560) treat the name Macrones luridus in Tirant (1911) as a nomen nudum and possibly intended for this species.
Hemibagrus olyroides (Roberts, 1989)
Mystus olyroides Roberts, 1989a: 124, fig. 95. Type locality: Sintang Market (reportedly caught in Sungai Kebian),Western Borneo (Kalimantan Barat, Indonesia). Holotype: MZB 3625.
Distribution: Kapuas River, western Borneo (Roberts, 1989a).
Hemibagrus peguensis (Boulenger, 1894)
Macrones peguensis Boulenger, 1894b: 196. Type locality: Sittang River near Toungoo [Burma]. Syntypes: BMNH
1894.5.21.25–26 (2).
Distribution: Irrawaddy, Sittang and Pegu River basins, Myanmar (Ng & Ferraris, 2000).
Hemibagrus planiceps (Valenciennes, 1840)
Bagrus planiceps Valenciennes, in Cuvier & Valenciennes, 1840a: 421 (311 of Strasbourg deluxe edition). Type
locality: Java. Lectotype: RMNH 2939, designated by Ng et al. (1999: 38). Roberts (1993: 30) citation of
MNHN b-0615 as holotype is incorrect (M. Kottelat, pers. comm.). Placed on Official List (Opinion 2011)(ICZN, 2002).
Bagrus anisurus Valenciennes, in Cuvier & Valenciennes, 1840a: 422 (313 of Strasbourg deluxe edition). Type
locality: Java. Lectotype: RMNH 2956; designated by Kottelat (1999: 272). Previously unpublished illustrationof lectotype reproduced in Roberts (1993: 30, fig. 66), but incorrectly listing the specimen as holotype (M. Kottelat, pers. comm.).
Bagrus flavus Bleeker, 1846a: 156. Type locality: Batavia. Holotype or syntype (size and number not stated):
RMNH 15897 (1 of 4), SMNS 10570 (1). Placed on Official List (Opinion 2011) (ICZN, 2002).
Distribution: Java.
Remarks: Bleeker (1858e: 417) acted as first reviser in giving precedence to Hemibagrus planiceps over Bagrus anisurus.
Hemibagrus pluriradiatus (Vaillant, 1892)
Macrones pluriradiatus Vaillant, 1892a: 126. Type locality: Tonkin. Holotype: MNHN 1892-0048; illustrated in
Vaillant (1904: 462, pl. 23, fig. 2) and Jayaram (1978b: 223, fig. 1).
Distribution: Yuianjiang, Hainan Island, China (Chu et al., 1999); Nam Xam basin, Laos, Red River, Viet Nam andChina (Kottelat, 2001a, b).

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Remarks: Also treated as valid in Mystus in some recent literature.
Hemibagrus punctatus (Jerdon, 1849)
Bagrus punctatus Jerdon, 1849: 339. Type locality: Cavery and its principal tributaries, southern India. No types
known.
Distribution: Cauvery River basin to Bhavani River in Tamil Nada; also rivers in Mysore, Western Ghats and adjoining hills, peninsular India (Jayaram & Anuradha, 2003).
Hemibagrus sabanus (Inger & Chin, 1959)
Mystus sabanus Inger & Chin, 1959: 294. Type locality: Kinabatangan River at Deramakot, Kinabatangan District,
North Borneo. Holotype: FMNH 68088.
Distribution: Kinabatangan River, Borneo, Malaysia (Inger & Chin, 1959).
Hemibagrus spilopterus Ng & Rainboth, 1999
Hemibagrus spilopterus Ng & Rainboth, 1999: 562, figs. 5, 7a, 8a. Type locality: Cambodia: Kandal, Bassac River
at village of Prek Chey on Vietnamese border (10º57'N, 105º06'E). Holotype: UMMZ 232611.
Distribution: Lower Mekong River basin, Cambodia (Ng & Rainboth, 1999).
Hemibagrus variegatus Ng & Ferraris, 2000
Hemibagrus variegatus Ng & Ferraris, 2000: 139, fig. 12. Type locality: Tenasserim River, Myanmar. Holotype:
BMNH 1992.11.16.11.
Distribution: Tenasserim River basin, Myanmar (Ng & Ferraris, 2000).
Hemibagrus velox Tan & Ng, 2000
Hemibagrus velox Tan & Ng, 2000: 272, figs. 2–4. Type locality: Sumatra Barat: Sungai Dareh, Pulau Punjung market, Sumatra. Holotype: MZB 9305.
Distribution: Rivers of central Sumatra (Tan & Ng, 2000).
Hemibagrus vietnamicus Mai, 1978
Hemibagrus vietnamicus Mai, 1978: 252, fig. 116 [not 115]. Type locality: [Mountain streams in North Vietnam].
Syntypes: at DVZUT. Spelled Hemibagrus vietnammicus in index (p. 337). Eschmeyer et al. (1998: 1760) acted
as first reviser in selecting vietnamicus as the correct spelling.
Distribution: Nam Ma and Nam Xam, Laos, and Northern Vietnam (Kottelat, 2001b).
Hemibagrus wyckii (Bleeker, 1858)
Bagrus Wyckii Bleeker, 1858b: 150, 156. Type locality: Java, in flumine Tjitarum, provinciae Preanger, prope vicum
Parongkalong. Syntypes (2: 320–440 mm TL): BMNH 1863.12.4.88 (1, 247 mm SL, 306 mm TL), RMNH
6897 (1). Illustrated in Bleeker (1862–63: 57, pl. 72 [= Silur. pl. 24]).
Distribution: Mekong and Chao Phraya River basins, and Sundaland (Kottelat, 2001b).
Hemibagrus wyckioides (Fang & Chaux, 1949)
Macrones (Hemibagrus) Wyckioides Fang & Chaux, in Chaux & Fang, 1949a: 199, fig. 3. Type locality: Cambodge.
Holotype: MNHN 1966-0727.
Mystus aubentoni Desoutter 1975: 449, figs. 2–3. Type locality: Stung Treng [Cambodia.] Holotype: MNHN 19740039.
Distribution: Mekong River basin, Cambodia, Laos, Thailand and China (Kottelat, 2001b).
Species inquirenda, Hemibagrus
Aoria Amemiyai Kimura, 1934: 166, pl. 5 (fig. 2). Type locality: Howchwan, Szechwan Province, China. Holotype: atLaboratory of Biological Department of the Shanghai Science Institute, Shanghai. As Aoria amemiyae on pp. 12 and
166, Aoria amemiyai on plate 5; Kottelat et al. (1998: 570) act as first revisers in selecting amemiyai as valid.
Treated as a possible synonym of Hemibagrus macropterus by Kottelat et al. (1998: 570).
HEMILEIOCASSIS Ng & Lim, 2000
Hemileiocassis Ng & Lim, 2000: 192. Type species: Hemileiocassis panjang Ng & Lim, 2000. Type by original designation. Gender: Feminine.
Hemileiocassis panjang Ng & Lim, 2000

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Hemileiocassis panjang Ng & Lim, 2000: 192, figs. 1, 2a, 3. Type locality: Indonesia: Java, Bogor. Holotype: ZMA
121529.
Distribution: Western Java (Ng & Lim, 2000).
HYALOBAGRUS Ng & Kottelat, 1998
Hyalobagrus Ng & Kottelat, 1998a: 336. Type species: Pseudobagrus ornatus Duncker, 1904. Type by original designation. Gender: Masculine.
Hyalobagrus flavus Ng & Kottelat, 1998
Hyalobagrus flavus Ng & Kottelat, 1998a: 340, fig. 6. Type locality: Indonesia: Sumatra: Jambi: Danau Arang
Arang in the vicinity of Muara Kompeh, 1º37'32.0"S, 103º47'19.0"E. Holotype: MZB 9302.
Distribution: Batang Hari drainage, Sumatra and Mentaya River basin, southern Borneo (Ng & Kottelat, 1998a).
Hyalobagrus leiacanthus Ng & Kottelat, 1998
Hyalobagrus leiacanthus Ng & Kottelat, 1998a: 341, fig. 9. Type locality: Indonesia: Borneo: Kalimantan Tengah:
Sungai Barito at Muara Laung, 0º35'34.6"S, 114º44'12.3"E. Holotype: MZB 9304.
Distribution: Kapuas and Barito River basins, central Borneo (Ng & Kottelat, 1998a).
Hyalobagrus ornatus (Duncker, 1904)
Pseudobagrus ornatus Duncker, 1904: 173, pl. 2 (figs. 13, 13a–c). Type locality: Muar Fluß bei Tubing Tinggi,
malayischen Halbinsel. Lectotype: ZMH H69, designated by Ladiges et al. (1958: 160); lectotype illustrated by
Jayaram (1968b: pl. 1) and Ng & Kottelat (1998a: fig. 10).
Distribution: Muar River drainage, southern Malay Peninsula (Ng & Kottelat, 1998a).
LEIOCASSIS Bleeker, 1857
Leiocassis Bleeker, 1857b: 416. Type species: Bagrus micropogon Bleeker, 1852. Type by monotypy. Gender: Feminine.
Liocassis Günther, 1864: 86. Type species: Bagrus micropogon Bleeker, 1852. Type by monotypy. Gender: Feminine. Unjustified emendation of Leiocassis Bleeker, 1857.
Rhinobagrus Bleeker, 1865: 7. Type species: Rhinobagrus dumerili Bleeker, 1865. Type by monotypy. Gender:
Masculine.
Nasocassis Nichols, 1925: 1. Type species: Liocassis longirostris Günther, 1864. Type by original designation. Originally proposed as a subgenus of Leiocassis. Gender: Feminine.
Dermocassis Nichols, 1925: 1. Type species: Bagrus ussuriensis Dybowski, 1872. Type by original designation.
Originally proposed as a subgenus of Leiocassis. Gender: Feminine.
Leiocassis aculeatus Ng & Hadiaty, 2005
Leiocassis aculeatus Ng & Hadiaty, 2005: 84, fig. 1. Type locality: Sumatra: Aceh province: Sungai Soraya, tributary of Sungai Alas. Holotype: MZB 8715.
Distribution: Alas River basin, northern Sumatra Ng & Hadiaty (2005).
Leiocassis crassilabris Günther, 1864
Liocassis crassilabris Günther, 1864: 88. Type locality: China. Holotype: BMNH 1864.7.9.9.
Leiocassis crassilabris macrops Nichols, 1926a: 2, fig. 2. Type locality: Min R., near Yenping (Nanping), Fukien
Prov., China. Holotype: AMNH 8445.
Distribution: from Huanghe [= Yellow River] to Zhujiang [= Pearl River] (Chu et al., 1999).
Leiocassis crassirostris Regan, 1913
Liocassis crassirostris Regan, 1913b: 552. Type locality: Kiatiang-fu, Sze Chuen, China. Syntypes (2): BMNH
1891.6.13.23.
Distribution: Sichuan, China.
Leiocassis herzensteini (Berg, 1907)
Macrones herzensteini Berg, 1907: 421. Type locality: [Onon River mouth, Russia]. Syntypes: ZIN 7092.
Distribution: Onon River basin, Russia.
Leiocassis longibarbus Cui, 1990
Leiocassis longibarbus Cui, 1990: 153, fig. 156. Type locality: Kinsha River system, Yunnan, China. Holotype: KIZ

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818719.
Distribution: Jinsha River basin [= upper Yangtze River], China (Cui, 1990).
Leiocassis longirostris Günther, 1864
Liocassis longirostris Günther, 1864: 87, unnumbered figure. Type locality: Japan [possibly in error for China].
Holotype: BMNH 1862.11.1.1.
Rhinobagrus dumerili Bleeker, 1865: 7. Type locality: China. Holotype: Whereabouts unknown.
Distribution: from Yalujiang [= Yalu River] to Minjiang [Yangtze River tributary], China (Chu et al., 1999); Seoul,
Pyongyang Kim River, Korea (Mori, 1952).
Remarks: Lee & Kim (1990: 133) state that Günther’s name appeared first; Rendahl (1927) apparently came to the
opposite conclusion and treated Leiocassis dumerili as valid. Weber & de Beaufort list the Bleeker paper as having been published in 1865, which is followed here, which results in Leiocassis longirostris as the valid name.
Leiocassis micropogon (Bleeker, 1852)
Bagrus micropogon Bleeker, 1852a: 94. Type locality: Blitong, in flumine Tjirutjup [Malaysia]. Holotype (79 mm
TL): possibly in RMNH 6873 (2).
Leiocassis chaseni de Beaufort, 1933: 34. Type locality: Ulu Jelai, Pahang, Pahang basin, Malaysia. Holotype: ZRC
290.
Distribution: Malay Peninsula, Sumatra and Borneo (Roberts, 1989a).
Leiocassis poecilopterus (Valenciennes, 1840)
Bagrus poecilopterus Valenciennes, in Cuvier & Valenciennes, 1840a: 431 (320 of Strasbourg deluxe edition). Type
locality: rivière de Hèbak à Java. Holotype: RMNH 3004.
Leiocassis regani Jayaram, 1965a: 9. Type locality: North Borneo: Sadong. Holotype: USNM 35732.
Distribution: Java, Borneo and possibly elsewhere in Southeast Asia.
Leiocassis saravacensis Boulenger, 1894
Liocassis saravacensis Boulenger, 1894a: 246. Type locality: Senah, Sarawak state, Borneo, East Malaysia. Syntypes: BMNH 1893.3.6.168–169 (2).
Distribution: Borneo.
Leiocassis tenuifurcatus Nichols, 1931
Leiocassis tenuifurcatus Nichols, 1931: 1. Type locality: Chungan Hsien, northwestern Fukien Province, China.
Holotype: AMNH 9681; holotype illustrated in Nichols (1943: 46, fig. 9).
Distribution: Fukien Province, China.
Species inquirendae, Leiocassis
Liocassis baramensis Regan, 1906b: 67. Type locality: Baram R., Sarawak state, Borneo, East Malaysia. Holotype:
BMNH 1898.11.14.1. Treated in recent literature as a synonym of either Leiocassis micropogon or Leiocassis
poecilopterus.
Liocassis doriae Regan, 1913b: 551. Type locality: Borneo. Holotype: BMNH 1868.1.28.30. Treated in recent literature
as valid or as a synonym of Leiocassis micropogon or Leiocassis poecilopterus.
Liocassis Hosii Regan, 1906b: 67. Type locality: Sibu, Sarawak state, Borneo, East Malaysia. Syntypes (6): BMNH
1906.10.29.18–22 (5). Treated as a synonym of Leiocassis micropogon or Leiocassis poecilopterus.
Liocassis merabensis Regan, 1913b: 550. Type locality: Merabeh, North Borneo. Syntypes: BMNH 1893.3.6.170–171(2). Treated as a synonym of Leiocassis micropogon or Leiocassis poecilopterus.
MYSTUS Scopoli, 1777
Mystus Scopoli, 1777: 451. Type species: Appeared first without species. Type widely listed as Bagrus halepensis
Valenciennes, 1840, with type by subsequent designation by Jordan & Evermann (1917: 21) [see Remarks].
Gender: Masculine.
Aspidobagrus Bleeker, 1862 (in Bleeker, 1862–63): 9. Type species: Bagrus gulio [= Pimelodus gulio Hamilton,
1822]. Type by original designation. Gender: Masculine.
Hypselobagrus Bleeker, 1862 (in Bleeker, 1862–63): 10. Type species: Bagrus macronemus Bleeker, 1846. Type by

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original designation. Gender: Masculine.
Heterobagrus Bleeker, 1864b: 355. Type species: Heterobagrus bocourti Bleeker, 1864. Type by original designation. Gender: Masculine.
Prajadhipokia Fowler, 1934b: 338. Type species: Prajadhipokia rex Fowler, 1934. Type by original designation.
Gender: Feminine.
Remarks: Heterobagrus is treated in some literature as valid with Prajadhipokia as a synonym. Roberts (1994: 243)
indicated that Jordan & Evermann’s (1917: 21) selection of Bagrus halepensis as type of Mystus Gronovius (aname that was ruled to be unavailable in ICZN Opinion 261 (ICZN, 1954: 281) also applied to Mystus Scopoli.
See Jayaram & Anuradha (1984) for summary of nomenclatural history of Mystus.
Revision: Jayaram & Anuradha (2003, senso lato, including Hemibagrus).
Review: Jayaram (1954, southern Asia); Roberts (1994, selected species, as Mystus sensu stricto).
Mystus alasensis Ng & Hadiaty, 2005
Mystus alasensis Ng & Hadiaty, 2005: 86, fig. 3. Type locality: Sumatra: Aceh province: Sungai Lembang, Gunung
Leuser National Park. Holotype: MZB 8704.
Distribution: Alas River basin, northern Sumatra (Ng & Hadiaty, 2005).
Mystus albolineatus Roberts, 1994
Mystus albolineatus Roberts, 1994: 245, fig. 1. Type locality: Prachinburi market, Bangpakong basin [Thailand].
Holotype: CAS 79030.
Distribution: Mekong, Bangpakong and Chao Phraya River basins (Roberts, 1994; Kottelat, 2001b).
Mystus armatus (Day, 1865)
Macrones armatus Day, 1865b: 289, fig. Type locality: Rivers and occasionally in backwaters, Cochin, India. Possible syntypes: BMNH 1865.7.17.21 (1), BMNH 1975.9.30.13 (1, dry), NMW 45165 (1), ZSI F1084 (1, lost).
Distribution: Malabar and Wayanad hills, Kerala and Manipur, India, and Mergui District, Myanmar (Jayaram &Anuradha, 2003).
Mystus armiger Ng, 2004
Mystus armiger Ng, 2004b: 883, fig. 1. Type locality: Malaysia: Kelantan, Kelantan River at Kampung Kuala Besar,
200 m from the sea (6º 12'N, 102º12'E). Holotype: CAS 218896.
Distribution: Kelantan River, Malaysia (Ng, 2004b).
Mystus atrifasciatus Fowler, 1937
Mystus atrifasciatus Fowler, 1937: 146, figs. 35–37. Type locality: Pitsanulok, Siam. Holotype: ANSP 67907.
Distribution: Mekong, Meklong and Chao Phraya River basins (Roberts, 1993; Kottelat, 2001b).
Mystus bimaculatus (Volz, 1904)
Macrones bimaculatus Volz, 1904: 466. Type locality: Sengi Si Russu and Kwantang River, near Djapura(Indragari), Sumatra. Syntypes (5): MHNG 683.28 (1), NMW 45166 (1), NMW 45167 (1).
Distribution: Sumatra (Volz, 1904).
Mystus bleekeri (Day, 1877)
Macrones Bleekeri Day, 1877 (in Day, 1875–78): 451, pl. 101 (fig. 1). Type locality: Sind, Jumna, upper waters of
the Ganges, and Burma [locality of lectotype not stated]. Lectotype: ZSI 1076, designated by Sharma & Dutt(1983: 334).
Distribution: Ganges, Indus and Mahanadi River basins (Roberts, 1994).
Remarks: Lectotype designation by Roberts (1994: 246) of RMNH 3024 is invalid because of the prior designation
by Sharma & Dutt (1983: 334). This name may be a senior synonym of Mystus rufescens (Vinciguerra), if thelectotype was selected from the Burmese specimens of Day.
Mystus bocourti (Bleeker, 1864)
Heterobagrus Bocourti Bleeker, 1864b: 355, pl. Type locality: Siam. Holotype: MNHN 0000-1553.
Prajadhipokia rex Fowler, 1934b: 339, figs. 3–4. Type locality: Bangkok, Siam. Holotype: ANSP 60179.
Distribution: Mekong and Chao Phraya River basins (Roberts, 1994; Kottelat, 2001b).
Mystus castaneus Ng, 2002
Mystus castaneus Ng, 2002d: 163, fig. 2. Type locality: Borneo: Sarawak, Serian market, from Sungai Sadong.
Holotype: ZRC 41848.

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Mystus cavasius (Hamilton, 1822)
Pimelodus cavasius Hamilton, 1822: 203, 379, pl. 11 (fig. 67). Type locality: larger freshwater rivers of the Gangetic
provinces. No types known.
Mystus mukherjii Ganguly & Datta, 1975: 293, figs. 1–2. Type locality: Subarnarekha River, below water falls at
Hundru, Ranchi District, Bihar, India. Holotype: at Fish. Lab., Dept. Zool., Univ. Calcutta.
Distribution: Ganges, Brahmaputra, Mahanadi, Subarnarekhar and Godavari river basins, India, Bangladesh and
Nepal (Chakrabarty & Ng, 2005).
Remarks: See Dutt et al. (1982) for comments on the validity of this species.
† Mystus dalungshanensis Li & Wang, 1979
† Mystus dalungshanensis Li & Wang, 1979: 48, fig. 13. Type locality: China: Guangdong, Sanshui Basin and adjacent districts; Early-Tertiary. Holotype: H-74116–1 (repository not stated).
Distribution: Guangdong, China; Eocene (Chang & Zhou, 1993)
Remarks: Apparently also described as new in Wang et al. (1981: 21, English 73).
Mystus falcarius Chakrabarty & Ng, 2005
Mystus falcarius Chakrabarty & Ng, 2005: 13, fig. 7. Type locality: Myanmar: Kachin State, Myitkyina market.
Holotype: CAS 89001.
Distribution: Irrawaddy, Salween and coastal river basins, Myanmar (Chakrabarty & Ng, 2005).
Mystus gulio (Hamilton, 1822)
Pimelodus gulio Hamilton, 1822: 201, 379, pl. 23 (fig. 66). Type locality: Higher parts of the Gangetic estuaries,
where the water is not very salt. No types known.
Bagrus abbreviatus Valenciennes, in Cuvier & Valenciennes, 1840a: 420 (311 of Strasbourg deluxe edition). Type
locality: Java. Holotype: RMNH 2942; previously unpublished illustration of holotype reproduced in Roberts(1993: 28, fig. 62).
Bagrus albilabris Valenciennes, in Cuvier & Valenciennes, 1840a: 416 (308 of Strasbourg deluxe edition). Type
locality: Calcutta ... Bengal ... Ceilan ... Pondichéry. Syntypes: MNHN 0000-4172 (1), MNHN 0000-4336 (2),MNHN a-8967 (6), MNHN a-9009 (1).
Bagrus Birmannus Valenciennes, in Cuvier & Valenciennes, 1840a: 419 (310 of Strasbourg deluxe edition). Type
locality: dans l’Irawadi. Holotype: MNHN 0000-0577.
Bagrus fuscus Valenciennes, in Cuvier & Valenciennes, 1840a: 417 (309 of Strasbourg deluxe edition). Type locality: Environs de Cananor. Holotype: MNHN 0000-0590.
Bagrus gulioides Bleeker, 1846a: 152. Type locality: Batavia. Syntypes (size and number not stated): RMNH 6862
(some of 23).
Bagrus melas Bleeker, 1846a: 152. Type locality: Java. Type(s) (size and number not stated): Whereabouts
unknown.
Bagrus rhodopterygius Bleeker, 1846a: 153. Type locality: Batavia. Type(s) (size and number not stated): Whereabouts unknown.
Bagrus Schlegelii Bleeker, 1846a: 153. Type locality: Batavia. Type(s) (size and number not stated): Whereabouts
unknown.
Distribution: Estuaries and lower portions of rivers of India, Bangladesh and Myanmar; and Java (Roberts, 1993).
Remarks: Synonymy of Bagrus birmannus based on examination of holotype; synonymy of Bagrus abbreviatus follows Roberts (1993: 28). The name Silurus porosus in Hora (1933: 133), which has often been treated as a synonym of Mystus gulio, is not available, as it was not treated as a valid species therein.
Mystus horai Jayaram, 1954
Mystus (Mystus) vittatus horai Jayaram, 1954: 536, pl. 19; text fig. 4. Type locality: River Indus, Kalabagh [Pakistan]. Syntypes (4): lost (Jayaram & Anuradha, 2003).
Distribution: Indus River (Jayaram & Anuradha, 2003; Mirza, 2003).
Mystus impluviatus Ng, 2003
Mystus impluviatus Ng, 2003a: 373, fig. 1. Type locality: Borneo: Kalimantan Timur, small river on road 12.2 km
from Sebulu crossroads towards forest and 59 km from Air Putih crossorads (0º16'S, 117º0'E). Holotype: CAS

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97049.
Distribution: Mahakam River basin, eastern Borneo (Ng, 2003a).
Mystus leucophasis (Blyth, 1860)
Bagrus leucophasis Blyth, 1860: 148. Type locality: Sitang and other Burma rivers. Possible type: AMS B.7925 (1).
Distribution: Irrawaddy and Sittang River basins, Myanmar (pers. obs.).
Mystus malabaricus (Jerdon, 1849)
Bagrus Malabaricus Jerdon, 1849: 338. Type locality: Mountain streams of Malabar [southern India]. No types
known.
Hara Malabarica Day, 1865a: 184, pl. 13 (fig. 3). Type locality: Mountain streams of Malabar, India. Possible types:
at ZSI and NMW 45228 (1). Neotype designation by Grant (1999: 170) not valid; absence of types and nomenclatural need not clearly demonstrated. Apparently a synonym, and therefore a junior homonym, of Bagrus malabaricus Jerdon, 1849.
Mystus canarensis Grant, 1999: 170. Type locality: Mountain streams of Malabar, India. Replacement name forHara malabarica Day, 1865, preoccupied in Mystus by Bagrus malabaricus Jerdon, 1849.
Distribution: Malabar Coast, Wayanad hills and Travancore hill ranges, India (Jayaram & Anuradha, 2003).
Remarks: Synonymy of Hara malabarica follows Jayaram & Anuradha (2003).
Mystus montanus (Jerdon, 1849)
Bagrus montanus Jerdon, 1849: 337. Type locality: River at Manantoddy in Wynaad, southern India. No types
known.
Macrones montanus dibrugarensis Chaudhuri, 1913: 254, pl. 9 (figs. 2, 2a, 2b). Type locality: Dibrugarh [India].
Holotype: ZSI F7795/1. Originally Macrones montanus var. dibrugarensis.
Distribution: Peninsular India, especially in the western Ghats (Jayaram & Anuradha, 2003).
Mystus multiradiatus Roberts, 1992
Mystus multiradiatus Roberts, 1992: 82, fig. 3b. Type locality: Prachinburi market [Thailand]. Holotype: CAS
76119.
Distribution: Mekong and Chao Phraya River basins (Kottelat, 2001b).
Mystus mysticetus Roberts, 1992
Mystus mysticetus Roberts, 1992: 84, fig. 3c. Type locality: Nakorn Phanom market [Thailand]. Holotype: CAS
76121.
Distribution: Mekong and Chao Phraya River basins (Kottelat, 2001b).
Mystus nigriceps (Valenciennes, 1840)
Bagrus nigriceps Valenciennes, in Cuvier & Valenciennes, 1840a: 412 (305 of Strasbourg deluxe edition). Type
locality: Java. Holotype: RMNH 2948; previously unpublished illustration of holotype reproduced in Roberts(1993: 29, fig. 64). Neotype (RMNH 3009) designation by Roberts (1993: 29) apparently invalid, following thereported rediscovery of the holotype by Ng (2002d: 166).
Bagrus micracanthus Bleeker, 1846a: 151. Type locality: Batavia. Syntypes (size and number not stated): possibly
AMS B.7574 (1), AMS B.7979 (1), BMNH 1863.12.4.77 (1, 105 mm SL, 134 mm TL), NMV 46463 (1),RMNH 5216. Neotype (RMNH 15857) designation by Roberts (1989a) regarded by Ng (2002d: 166) as unnecessary and therefore invalid.
Distribution: Java and southern Sumatra (Ng, 2002d).
Mystus oculatus (Valenciennes, 1840)
Bagrus oculatus Valenciennes, in Cuvier & Valenciennes, 1840a: 424 (314 of Strasbourg deluxe edition). Type
locality: la côte de Malabar. Holotype: MNHN 0000-1195.
Distribution: Nilgiris, Western Ghats and hills of Karnataka, Kerala and Tamil Nadu, India (Jayaram & Anuradha,2003).
Mystus pelusius (Solander, 1794)
Silurus pelusius Solander, in Russell, 1794: 210, pl. 7 (fig. 1). Type locality: Aleppo. No types known.
Bagrus Halepensis Valenciennes, in Cuvier & Valenciennes, 1840a: 413 (306 of Strasbourg deluxe edition). Type
locality: la riviere de l’Alep. Syntype: BMNH 1955.6.25.1 (1, 6 inches) Description based on illustrations in
Russell (1794: pl. 13, fig. 1) and Gronovius (1763: pl. 8a, fig. 6); BMNH specimen from collection of Russell.

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Macrones aleppensis Günther, 1864: 75. Type locality: Aleppo. No types known, based on illustrations in Russell(1794: pl. 13, fig. 1) and Gronovius (1763: pl. 8a, fig. 6). Unjustified emendation of Bagrus Halepensis Valenciennes, 1840.
Macrones colvillii Günther, 1874: 36, pl. 8. Type locality: Tigris at Baghdad. Syntypes: BMNH 1874.4.28.6–8 (3),
BMNH 1875.1.14.19–21 (3).
Mystus misrai Anuradha, 1986: 292, figs. 1–2. Type locality: Lake Antioche, Syria. Holotype: MHNG 603.95.
Distribution: Euphrates River basin (Roberts, 1994); Tigris River basin (Coad, 1995).
Remarks: Redescribed in Anuradha & Jayaram (1985).
Mystus pulcher (Chaudhuri, 1911)
Macrones pulcher Chaudhuri, 1911: 20, pl. 1 (fig. 4). Type locality: district of Bhamo close to Yunnan border[Upper Burma]. Syntypes: ZSI F4716/1 to F4719/1 (4).
Distribution: Irrawaddy basin and Sittang River, Myanmar.
Mystus punctifer Ng, Wirjoatmodjo & Hadiaty, 2001
Mystus punctifer Ng, Wirjoatmodjo & Hadiaty, 2001a: 356, figs. 1, 2a. Type locality: Sumatra: Aceh, trail in Suag
Belimbing research station, Gunung Leuser National Park [Indonesia]. Holotype: MZB 8703.
Distribution: Alas River basin, northern Sumatra (Ng et al., 2001a).
Mystus rhegma Fowler, 1935
Mystus rhegma Fowler, 1935b: 102, fig. 27. Type locality: Bangkok, Siam. Holotype: ANSP 61748.
Distribution: Mekong, Meklong and Chao Phraya River basins (Roberts, 1994; Kottelat, 2001b).
Mystus rufescens (Vinciguerra, 1890)
Macrones rufescens Vinciguerra, 1890: 226, pl. 7 (fig. 2). Type locality: Meetan, Birmania [Myanmar]. Holotype:
MSNG 14585.
Distribution: Irrawaddy, Sittang and lower Salween Rivers, Myanmar (Roberts, 1994).
Mystus seengtee (Sykes, 1839)
Pimelodus Seengtee Sykes, 1839: 164. Type locality: Deccan, India. Holotype (6 inches): possibly BMNH
1857.6.13.154 (1, dry). Also described as new in Sykes (1840: 61); illustrated and described in more detail in
Sykes (1841: 374, pl. 66, fig. 2). Illustration of type reproduced in Chakrabarty & Ng (2005: 10, fig. 6).
Distribution: Krishna and Cauvery river basins, India (Chakrabarty & Ng, 2005).
Remarks: See Chakrabarty & Ng (2005: 18) for discussion of the status of the holotype of the species.
Mystus singaringan (Bleeker, 1846)
Bagrus heterurus Bleeker, 1846a: 151. Type locality: Batavia [now: Western Java, by neotype designation]. Neotype: RMNH 2951, designated by Roberts (1994: 252).
Bagrus macronemus Bleeker, 1846a: 150. Type locality: Batavia [now: Western Java by neotype designation]. Neotype: RMNH 2951, designated by Roberts (1994: 253), but Eschmeyer et al. (1998) indicated that specimens
cataloged as SMNS 10571 may be extant syntypes. Preoccupied by Bagrus macronemus Ranzani, 1842 (now in
the Ariidae, as a synonym of Bagre bagre); apparently not replaced.
Bagrus singaringan Bleeker, 1846a: 150. Type locality: Batavia [now Western Java, by neotype designation]. Neotype: RMNH 2951, designated by Roberts (1994: 252).
Distribution: Mekong and Chao Phraya River basins, and Sundaland (Kottelat, 2001b).
Remarks: Roberts (1994: 252) serves as first reviser selecting Mystus singaringan over Bagrus heterurus; earlier
first reviser action of Bleeker (1862–63: 58), in which Hypselobagrus macronema is used as valid, with Bagrus
heterurus and B. singaringan as synonyms, is invalid because of the homonymy of Bagrus macronemus Ranzani.
† Mystus spinipectoralis Li & Wang, 1979
† Mystus spinipectoralis Li & Wang, 1979: 48, fig. 14. Type locality: China: Guangdong, Sanshui Basin and adjacent districts; Early-Tertiary. Holotype: H.74112 (repository not stated).
Distribution: China, Eocene (Chang & Zhou, 1993).
Remarks: Apparently also described as new in Wang et al. (1981: 25, English 74).
Mystus tengara (Hamilton, 1822)
Pimelodus tengara Hamilton, 1822: 183, 377, pl. 3 (fig. 61). Type locality: in the ponds of India. No types known.

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Distribution: Widely distributed in northern India, as well as Bangladesh and Pakistan (Jayaram, 1999).
Mystus vittatus (Bloch, 1794)
Silurus vittatus Bloch, 1794: 50, pl. 371 (fig. 2). Type locality: Tranquebar. Holotype: ZMB 2939.
Pimelodus Batasius Hamilton, 1822: pl. 23 (fig. 60). Type locality: Not stated [Ponds of northern parts of Bengal].
No type known. Name available from caption to plate, which corresponds to the text of Pimelodus carcio.
Pimelodus carcio Hamilton, 1822: 181, 377, pl. 23 (fig. 60). Type locality: Ponds of northern parts of Bengal. No
types known.
Bagrus Keletius Valenciennes, in Cuvier & Valenciennes, 1840a: 411 (305 of Strasbourg deluxe edition). Type locality: Pondichéry. Lectotype: MNHN a-9011, designated by Ng (2002d: 166).
Distribution: Inland and coastal rivers of Pakistan, Nepal India, Sri Lanka, and Bangladesh (Roberts, 1993); published records from Myanmar apparently in error.
Remarks: Day (1871) and Hora (1929: 182) discuss the confusion surrounding the identity of the fish illustrated in
Hamilton (1822: pl. 23, fig. 60) as Pimelodus batasius. Bagrus Keletius Valenciennes, (1840a: 411) was treatedby Ng (2002d: 165) as a synonym of either Mystus armatus (Day, 1865) or Mystus vittatus (Bloch, 1794).
Mystus wolffii (Bleeker, 1851)
Bagrus Wolffii Bleeker, 1851d: 205. Type locality: Bandjarmassing, in fluviis. Holotype (131 mm TL): possibly
AMS B.7774 (1), BMNH 1863.12.4.94 (1, 102 mm SL, 129 mm TL), NMV 46225 (1), ZMB 4794 (1).
Distribution: Large rivers from Thailand to Indonesia, in the tidal zone and brackish water (Rainboth, 1996).
Remarks: Holotype listed as BMNH 1863.12.4.94 by Desoutter (1975: 447).
Species inquirenda, Mystus
Pimelodus anisurus M’Clelland, 1842: 583. Type locality: Loodianah and Cabool River at Jullalabad [Afghanistan].
Holotype: Whereabouts unknown, possibly at BMNH. Treated in recent literature as a possible synonym of Mystus
tengara Hamilton, 1822, but treated by Günther (1864: 190) as a species of Amblyceps.
NANOBAGRUS Mo, 1991
Nanobagrus Mo, 1991: 125, 143. Type species: Akysis armatus Vaillant, 1902. Type by original designation. Gender: Masculine.
Nanobagrus armatus (Vaillant, 1902)
Akysis armatus Vaillant, 1902: 64, fig. 10. Type locality: Tepoe, bords du Mahakam, Bornéo central. Holotype:
RMNH 7844.
Distribution: Mahakam and Kapuas River basins, Borneo (Ng & Tan, 1999).
Nanobagrus nebulosus Ng & Tan, 1999
Nanobagrus nebulosus Ng & Tan, 1999: 357, fig. 3a, 4a. Type locality: Malaysia: Johor, Sungai Kahang and tributary ca. 44.4 km from Mersing turnoff to Kluang just before side road to Endau Rompin Taman Negara, km 96
from Mersing to Batu Pahat (2º3.93'N, 103º31.58'E). Holotype: ZRC 42600.
Distribution: Endau and Sedili River drainages, southeastern Malay Peninsula (Ng & Tan, 1999).
Nanobagrus stellatus Tan & Ng, 2000
Nanobagrus stellatus Tan & Ng, 2000: 278, fig. 5a. Type locality: Jambi: Sg. Alai (from aquarium collectors),
Sumatra. Holotype: MZB 9306.
Distribution: Batang Hari basin, Sumatra (Tan & Ng, 2000).
† NIGERIUM White, 1934
† Nigerium White, 1934: 49, 52. Type species: † Nigerium wurnoense White, 1934. Type by original designation.
Gender: Neuter.
Remarks: Considered to be Bagridae S.L. by Gayet & Meunier (2003).
† Nigerium gadense White, 1934
† Nigerium gadense White, 1934: 52, pl. 7 (fig. 2). Type locality: Gada, Nigeria, in scree; Eocene. Holotype: at
National Collection [Nigeria] or BMNH (incomplete skull).

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† Nigerium wurnoense White, 1934
† Nigerium wurnoense White, 1934: 49, pl. 6. Type locality: Reptile bed at Wurno, Nigeria: Eocene. Holotype: at
National Collection [Nigeria] or BMNH (incomplete skull).
Distribution: Wurno, Nigeria: Eocene.
† NKONDOBAGRUS Van Neer, 1994
† Nkondobabrus Van Neer, 1994: 108. Type species: † Nkondobagrus longisrostris Van Neer, 1994. Type by monotypy. Gender: Masculine.
† Nkondobagrus longisrostris Van Neer, 1994
† Nkondobagrus longisrostris Van Neer, 1994: 108, pl. 4. Type locality: Nkondo-Kaiso area, Nkondo Formation,
Albertine Rift Valley, Upper Miocene, Uganda. Holotype: apparently at MRAC, NK 2637'89 (anterior portionof neurocranium, with part of vomerine tooth patch).
Distribution: Uganda, upper Miocene (Van Neer, 1994; Gayet & Meunier, 2003: 498).
Remarks: Considered to be Bagridae S.L. by Gayet & Meunier (2003).
OLYRA M’Clelland, 1842
Olyra M’Clelland, 1842: 588. Type species: Olyra longicaudata M’Clelland, 1842. Type by subsequent designation,
by Gill (1861c: 51). Gender: Feminine.
Review: Hora (1936a).
Olyra burmanica Day, 1872
Olyra burmanica Day, 1872: 711. Type locality: Pegu Yomas, Burma. Lectotype: AMS B.7560; designated by
Jayaram (1980: 19).
Distribution: Pegu Yomas; known only from type (Jayaram, 1980).
Olyra horae (Prashad & Mukerji, 1929)
Amblyceps horae Prashad & Mukerji, 1929: 173, pl. 7 (fig. 1). Type locality: Indawgyi Lake along its w. shore near
Loimon village, Myitkyina District, Upper Burma. Holotype: ZSI F10854/1.
Distribution: Meghalaya, India, and Indawgyi Lake, Myanmar (Jayaram, 1980).
Remarks: Name appears as Olyra horai in recent literature, which reflects an attempt to correct the suffix to thename to the masculine form. However, the original spelling is also an acceptable form and the subsequent spelling should be regarded as a misspelling.
Olyra kempi Chaudhuri, 1912
Olyra kempi Chaudhuri, 1912: 443, pl. 41 (figs. 4, 4a, 4b). Type locality: Mangaldai (Assam-Bhutan Frontier)[North India]. Syntypes (5): ZSI F5387/1 (1).
Distribution: Mangaldai, Assam, India (Jayaram, 1980).
Remarks: Treated in some literature as a synonym of Olyra longicaudatus.
Olyra longicaudata M’Clelland, 1842
Olyra longicaudatus M’Clelland, 1842: 588, pl. 21 (fig. 1). Type locality: Kaysah Mountains. Holotype: not saved(M’Clelland, 1842: 572). Name spelled Olyra longicaudata on p. 574.
Distribution: Base of Darjelling Himalaya, Assam, India; Tenassarim, Myanmar (Jayaram, 1980).
Species inquirendae, Olyra
Heptapterus Collettii Steindachner, 1881b: 98. Type locality: La Plata [in error]. Syntypes: NMW 46191 (2). Illustrated
and described in more detail in Steindachner (1881d: 7, pl. 5, fig. 1). Shown to be a species of Olyra in Bockmann &de Pinna (2004).
Olyra elongata Günther, 1883: 140. Type locality: Tenasserim [Burma]. Syntypes: BMNH 1880.12.1.64–67 (7 ?).
PELTEOBAGRUS Bleeker, 1865
Pelteobagrus Bleeker, 1865: 9. Type species: Silurus calvarius Basilewsky, 1855. Type by monotypy. Gender: Mas-

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culine.
Remarks: Mo (1991) noted that Pelteobagrus may not be monophyletic if species placed in Pseudobagrus and Coreobagrus were excluded.
Pelteobagrus argentivittatus (Regan, 1905)
Macrones argentivittatus Regan, 1905c: 390, pl. 5 (fig. 2). Type locality: Canton, Cungkun, China. Lectotype:
MHNG 677.99, designated by Jayaram (1974).
Distribution: from Heilongjiang [Amur River] to Zhujiang [= Pearl River] (Chu et al., 1999).
Pelteobagrus brashnikowi (Berg, 1907)
Macrones (Leiocassis) brashnikowi Berg, 1907: 421. Type locality: [Amur River mouth, Russia]. Syntypes (5): ZIN
13964 (2), ZIN 13965 (2), ZIN 13966 (1), but only two specimens still at ZIN (Naseka & Bogutskaya (2004:
286).
Distribution: Amur River basin (Berg, 1907).
Remarks: Also treated in recent literature as valid in either Pelteobagrus brashnikowi or Leiocassis brashnikowi;
inclusion of species in Pelteobagrus follows Naseka & Bogutskaya (2004).
Pelteobagrus eupogon (Boulenger, 1892)
Pseudobagrus eupogon Boulenger, 1892a: 247. Type locality: Shanghai, China. Holotype: BMNH 1891.1.31.19.
Distribution: Changjiang [= Yangtze River] basin (Chu et al., 1999).
Pelteobagrus fulvidraco (Richardson, 1846)
Pimelodus fulvi-draco Richardson, 1846: 286. Type locality: Canton, China. No types known; based on illustration
by Reeves, reproduced in Whitehead (1970: pl. 1b). Originally as: Pimelodus ? fulvi-draco.
Silurus calvarius Basilewsky, 1855: 241, pl. 9 (fig. 1). Type locality: in aquis fluentibus et stagnantibus atque in sinu
Tschiliense [China]. Type(s): Whereabouts unknown.
Distribution: Widely throughout China, except the western plateau and Xinjiang (Chu et al., 1999); Nam Ma Basin,
Laos, and northern and central Vietnam (Kottelat, 2001b).
Pelteobagrus intermedius (Nichols & Pope, 1927)
Pseudobagrus intermedius Nichols & Pope, 1927: 331, fig. 5. Type locality: Nodoa, Hainan [China]. Holotype:
AMNH 8360. Also treated as valid in Pseudobagrus.
Aoria henryi Herre, 1932: 432. Type locality: Canton wholesale fish market, China. Holotype: SU 25724. Treated inrecent literature as either a synonym of Pelteobagrus intermedius or as valid as Pseudobagrus henryi.
Distribution: Hainan Island, from Minjiang to Zhujiang (Chu et al., 1999); Vietnam (Kottelat, pers. comm).
Pelteobagrus mica (Gromov, 1970)
Mystus mica Gromov, 1970: 400, unnumbered figure. Type locality: Lake Ommi, middle Amur basin, Russia. Holotype: ZIN 39434.
Distribution: Amur River basin, Russia (Gromov, 1970).
Remarks: Inclusion of species in Pelteobagrus follows Naseka & Bogutskaya (2004).
Pelteobagrus microps (Rendahl, 1932)
Leiocassis microps Rendahl, 1932: 93. Type locality: Chungking, China. Holotype: NRM 10163.
Distribution: Chungking, China.
Pelteobagrus nudiceps (Sauvage, 1883)
Pseudobagrus nudiceps Sauvage, 1883a: 145. Type locality: le Lac Biwako, Japon. Holotype: MNHN a-6525. Also
described as new (from Siam) in Sauvage (1883b). The later description has been treated as a Lapsus calami by
Smith (1934: 296; 1945: 6) and Kottelat (1984b: 811).
Distribution: Lake Biwa; central and western Honshu, eastern Shikoku, and northwestern Kyushu islands, Japan(Matsuura et al., 2000). Reported from the Lower Pliocene of Mie Prefecture, Japan, in Watanabe & Okuyama(1994).
Remarks: Occasionally treated as valid as Pseudobagrus nudiceps.
Pelteobagrus ussuriensis (Dybowski, 1872)
Bagrus ussuriensis Dybowski, 1872: 210. Type locality: Ussuri and Sungari rivers, Amur R. basin, and Khanka
Lake, Russia. Syntype(s): ZMB 7954 (1?).
Pseudobagrus emarginatus Sowerby, 1921: 1. Type locality: Mouth of the Hun Kiang, in the Yalu River on the

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South Manchurian Border. Holotype: USNM 76716.
Distribution: from Heilongjiang [= Amur River] to Zhujiang [= Pearl River], China (Chu et al., 1999); Pyeuktong,
Engpyen, Taedong River, Han River, Korea (Mori, 1952).
Remarks: Treated in recent literature as valid as either Leiocassis ussuriensis and Pseudobagrus ussuriensis; inclusion of species in Pelteobagrus follows Naseka & Bogutskaya (2004).
Pelteobagrus vachellii (Richardson, 1846)
Bagrus vachellii Richardson, 1846: 284. Type locality: Chikiang, China. Neotype: BMNH 1855.3.27.9, designatedby Jayaram (1968b: 300). Originally as “? Bagrus vachellii.”
Pseudobagrus chinensis Wu, 1930b: 53, fig. 4. Type locality: Sze-Chwan, China. Holotype: Whereabouts unknown.
Name apparently not treated since original description.
Pseudobagrus fangi Wu 1930c: 84, fig. 8. Type locality: Kiating [Szechwan, China]. Holotype: Whereabouts
unknown.
Pseudobagrus wui Miao, 1934: 210, fig. 42. Type locality: Chinkiang, southern Kiangsu, China. Holotype: Mus.
Biol. Lab. Sci. Soc. China 12495.
Distribution: from Huanghe [= Yellow River] to Zhujiang [= Pearl River], China (Chu et al., 1999); Seoul, Suwon,
Taeque, Taedong River, Korea (Mori, 1952).
Pelteobagrus virgatus (Oshima, 1926)
Aoria virgatus Oshima, 1926: 4. Type locality: Kachek River, about 40 mi. above Kachek, Hainan Island, China.
Holotype: Whereabouts unknown.
Distribution: Hainan Island, Yuanjiang (Chu et al., 1999).
Remarks: Treated as valid in recent literature as Pelteobagrus virgatus, Pseudobagrus virgatus or Leiocassis virgatus. The name Pelteobagrus virgatus vinhensis in Nguyen (1982: 26), is a nomen nudum and is not available.
PSEUDOBAGRUS Bleeker, 1859
Pseudobagrus Bleeker, 1859: 257. Type species: Bagrus aurantiacus Temminck & Schlegel, 1846. Type by monotypy. Gender: Masculine.
Fluvidraco Jordan & Fowler, 1903: 904. Type species: Pseudobagrus ransonnetii Steindachner, 1887. Type by original designation. Gender: Masculine.
Pseudobagrus adiposalis Oshima, 1919
Pseudobagrus adiposalis Oshima, 1919: 181, pl. 48 (fig. 2). Type locality: Tamusui R. near Shinten, Taiwan. Lectotype: FMNH 59079; designated by Eschmeyer (in Eschmeyer et al., 1998: 40).
Distribution: Lingjiang, Oujiang [= Wu River], Xiangjiang, Zhujiang [Pearl River], China (Chu et al., 1999); Taiwan.
Pseudobagrus albomarginatus (Rendahl, 1928)
Leiocassis albomarginatus Rendahl, 1928: 170. Type locality: Tang-tu-hsien, Anhui, China. Syntypes: NRM 10017(5).
Distribution: Anhui and Guangdong, China.
Pseudobagrus analis (Nichols, 1930)
Leiocassis (Dermocassis) analis Nichols, 1930: 4, fig. 3. Type locality: Hokou (Chaoshan), ne. Kiangsi Prov.,
China. Holotype: AMNH 9680.
Distribution: middle and lower Changjiang [= Yangtze River], China (Chu et al., 1999).
Pseudobagrus aurantiacus (Temminck & Schlegel, 1846)
Bagrus aurantiacus Temminck & Schlegel, 1846, in Temminck & Schlegel, 1843–50: 227, pl. 104 (fig. 2). Type
locality: Japan. Lectotype: RMNH 2952a, designated by Boeseman (1947: 169).
Pseudobagrus ransonnettii Steindachner, in Steindachner & Döderlein, 1887: 287. Type locality: Osaka, Japan. Syntypes (2): at NMW.
Distribution: Western Kyushu and Iki Islands, Japan (Matsuura et al., 2000).
Remarks: Redescribed in Watanabe & Maeda (1995).
Pseudobagrus brevianalis Regan, 1908
Pseudobagrus brevianalis Regan, 1908f: 151. Type locality: Lake Candidius, Formosa. Syntypes: BMNHCHECKLIST OF CATFISHES, RECENT AND FOSSIL

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1908.5.27.23–28 (6).
Distribution: Taiwan (Chu et al., 1999).
Pseudobagrus brevicaudatus (Wu, 1930)
Leiocassis brevicaudatus Wu, 1930c: 81, fig. 7. Type locality: Chungking, Szechwan, China. Holotype: Whereabouts unknown.
Leiocassis lui Tchang & Shih, 1934: 434. Type locality: valey of Lower Kialingkiang, from Chungking to Hochuanof Szechwan province [China]. Holotype: ZMFMIB 9720.
Distribution: Changjiang [= Yangtze River] basin, China (Chu et al., 1999).
Remarks: See Zhang & Wang (1996) for taxonomic discussion.
Pseudobagrus gracilis Li, Chen & Chan, 2005
Pseudobagrus gracilis Li, Chen & Chan, 2005: 50, fig. 1. Type locality: China: Guangxi Province, Guilin City,
Zhujiang basin, Lijiang drainage, 29º55'23 N, 118º28'12 E. Holotype: SCNU 0410002.
Distribution: Beijiang and Xijiang [= West River] River basins, southern China (Li et al., 2005).
† Pseudobagrus ikiensis Watanabe & Uyeno, 1999
† Pseudobagrus ikiensis Watanabe & Uyeno, 1999: 398, figs. 2–3 Type locality: Chojabaru Formation (Ika Group)
Middle Miocene, diatomite at Hachiman, Ashibe, Iki Island, Nagasaki Prefecture, Japan. Holotype: GSJ F 7735
(complete skeleton).
Distribution: Nagasaki Prefecture, Japan; Middle Miocene.
Pseudobagrus koreanus Uchida, 1990
Pseudobagrus koreanus Uchida, in Lee & Kim, 1990: 124, figs. 3b, 6. Type locality: Namwôn-shi, Sômjin R.,
Korea. Holotype: BKNU 1111.
Distribution: Sômjin River, Korea (Uchida, in Lee & Kim, 1990).
Pseudobagrus medianalis (Regan, 1904)
Macrones medianalis Regan, 1904a: 194. Type locality: Sea of Tien [Tien Chih], Yunnan Fu, China, elev. 6000 ft.
Syntypes: BMNH 1904.1.26.44–46 (3).
Distribution: Dianchi Lake, China (Chu et al., 1999).
Pseudobagrus omeihensis (Nichols, 1941)
Leiocassis omeihensis Nichols, 1941: 1. Type locality: Omeihsien, Szechwan Prov., China, elev. 1600 ft. Holotype:
AMNH 15217.
Distribution: Emei, Sichuan, China (Chu et al., 1999).
Pseudobagrus ondon Shaw, 1930
Pseudobagrus ondon Shaw, 1930: 111, figs. 1–2. Type locality: Small river at Shing-Tsong, Chekiang [Zhekjiang],
China. Holotype: ASIZB [= ZMFMIB 4449].
Distribution: Luohe, Oujiang, Linghiang, Caojiang, Huaihe, Hanshui, China (Chu et al., 1999).
Pseudobagrus pratti (Günther, 1892)
Macrones pratti Günther, 1892: 245, pl. 1 (fig. b). Type locality: Kia-tiang-fu, Szechwan [Sichuan], Prov., China,
elev. 1070 ft. Holotype: BMNH 1891.6.13.25.
Liocassis emarginatus Regan, 1913b: 553. Type locality: Kia-tiang-fu, Sze Chuen [China]. Syntypes (4): BMNH
1891.6.13.21 (1?).
Distribution: Changjiang [= Yangtze River] basin, China (Chu et al., 1999).
Remarks: Synonymy follows Watanabe (1995).
Pseudobagrus taeniatus (Günther, 1873)
Macrones (Liocassis) taeniatus Günther, 1873b: 245. Type locality: Shanghai, China. Holotype: BMNH
1873.7.30.73.
Distribution: from Changjiang [= Yangtze River] to Minjiang, China (Chu et al., 1999).
Pseudobagrus taiwanensis Oshima, 1919
Pseudobagrus taiwanensis Oshima, 1919: 180, pl. 48 (fig. 1). Type locality: Tozen R. near Taichu, Taiwan. Lectotype: FMNH 59078; designated by Eschmeyer (in Eschmeyer et al., 1998: 1646).
Distribution: Taiwan (Chu et al., 1999).
Pseudobagrus tenuis (Günther, 1873)

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Macrones (Pseudobagrus) tenuis Günther, 1873b: 244. Type locality: Shanghai, China. Holotype: BMNH
1873.7.30.72.
Distribution: from Changjiang [= Yangtze River] to Minjiang, China (Chu et al., 1999).
Pseudobagrus tokiensis Döderlein, 1887
Pseudobagrus tokiensis Döderlein, in Steindachner & Döderlein, 1887: 288. Type locality: River near Tokyo, Japan.
Possible syntypes: BMNH 1891.5.26.19–20 (2), MZS 498 (7), MZS 643 (1), ZMB 12246 (2).
Distribution: northern portions of Honshu, Japan (Matsuura et al., 2000).
Remarks: Redescribed in Watanabe & Maeda (1995).
Pseudobagrus trilineatus (Zheng, 1979)
Leiocassis trilineatus Zheng, 1979: 182, fig. 1. Type locality: Mountain stream in Luofu Shan, Boluo Xian, Guangdong Province, China. Holotype: JNU 76VII Luofu Shan 029.
Distribution: Dongjiang basin, China (Chu et al., 1999).
Pseudobagrus truncatus (Regan, 1913)
Liocassis truncatus Regan, 1913b: 553. Type locality: Kia-tiang-fu [China]. Syntypes (5): BMNH 1891.6.13.24.
Distribution: Changjiang [= Yangtze River] basin, China (Chu et al., 1999).
Species inquirenda, Pseudobagrus
Pseudobagrus changi Miao, 1934: 213, fig. 43. Type locality: Chinkiang, southern Kiangsu, China. Holotype: Mus. Biol.
Lab. Sci. Soc. China 12494. [name not in recent use.]
PSEUDOMYSTUS Jayaram, 1968
Pseudomystus Jayaram, 1968b: 359. Bagrus stenomus Valenciennes, 1840. Type by original designation. Originallyproposed as a subgenus of Leiocassis. Gender: Masculine.
Remarks: Treated in some recent literature as a synonym of Leiocassis.
Pseudomystus bomboides Kottelat, 2000
Pseudomystus bomboides Kottelat, 2000a: 77, fig. 69. Type locality: Laos: Vientiane Province, Mekong basin, confluence of Nam Leuk and Nam Ngong, 18º22'04"N, 103º05'27"E. Holotype: ZRC 45426.
Distribution: Mekong River basin and southern Thailand (Kottelat, 2001b).
Pseudomystus breviceps (Regan, 1913)
Liocassis breviceps Regan, 1913b: 551. Type locality: Deli, Sumatra. Syntypes: BMNH 1889.11.12.64–65 (2).
Distribution: Sumatra, known only from types.
Remarks: BMNH 1889.11.12.66 was incorrectly listed as holotype by Ng & Rachmatika, (1999: 178).
Pseudomystus carnosus Ng & Lim, 2005
Pseudomystus carnosus Ng & Lim, 2005: 7, fig. 5. Type locality: Sumatra: Sukadana. Holotype: RMNH 15859.
Distribution: Way Seputih River basin, southern Sumatra (Ng & Lim, 2005).
Pseudomystus flavipinnis Ng & Rachmatika, 1999
Pseudomystus flavipinnis Ng & Rachmatika, 1999: 175, fig. 5. Type locality: Borneo: Kalimantan Barat, Sungai
Embaloh at Benua Marinus. Holotype: MZB 6593.
Distribution: Kapuas River basin (Ng & Rachmatika, 1999).
Pseudomystus fumosus Ng & Lim, 2005
Pseudomystus fumosus Ng & Lim, 2005: 11, fig. 6. Type locality: Malaysia: Pahang, Taman Negara, Kuala Tahan.
Holotype: ZRC 3229.
Distribution: Pahang River basin, peninsular Malaysia (Ng & Lim, 2005).
Pseudomystus fuscus (Popta, 1904)
Leiocassis fuscus Popta, 1904: 186. Type locality: le Mahakam supérior, Bornéo central. Holotype: RMNH 7555;
described in more detail, with illustration of holotype, in Popta (1906: 53, pl. 3, fig. 10).
Distribution: Upper Mahakam River basin, Borneo, and Malay peninsula.
Pseudomystus inornatus (Boulenger, 1894)
Liocassis inornatus Boulenger, 1894a: 245. Type locality: Senah, Sarawak state, Borneo, East Malaysia. Holotype:

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BMNH 1893.3.6.179.
Distribution: Sungai Penyalin, Indonesia and Sarawak (Ng & Rachmatika, 1999).
Pseudomystus leiacanthus (Weber & de Beaufort, 1912)
Leiocassis leiacanthus Weber & de Beaufort, 1912: 536, pl. 2 (figs. 1–1a). Type locality: Sumatra (Kwantan river,Ringat and Taluk). Syntypes: ZMA 112671 (1), ZMA 112672 (1); syntype illustration reproduced in Weber &de Beaufort (1913: fig. 148).
Distribution: Sumatra; Malay Peninsula (Mohsin & Ambak, 1983).
Pseudomystus mahakamensis (Vaillant, 1902)
Leiocassis mahakamensis Vaillant, 1902: 55, figs. 4–5. Type locality: Tepoe, bords du Mahakam, Bornéo central.
Syntypes: RMNH 7838 (2).
Distribution: Mahakam River basin, Borneo.
Pseudomystus moeschii (Boulenger, 1890)
Liocassis moeschii Boulenger, 1890a: 39. Type locality: Deli River, Sumatra. Lectotype: BMNH 1889.11.12.66; lectotype designated by Ng & Lim (2005: 14).
Distribution: Deli and Banjuasin Rivers, Sumatra (Ng & Lim, 2005).
Remarks: Species redescribed in Ng & Lim (2005).
Pseudomystus myersi (Roberts, 1989)
Leiocassis myersi Roberts, 1989a: 117, fig. 92. Type locality: Small forest stream flowing into Kapuas mainstream
NE of Gunung Setunggul, 53 km NW of Sintang, Western Borneo (Kalimantan Barat, Indonesia). Holotype:
MZB 3588.
Distribution: Kapuas River, western Borneo (Roberts, 1989a).
Pseudomystus robustus (Inger & Chin, 1959)
Leiocassis robustus Inger & Chin, 1959: 290, fig. 48. Type locality: Kinabatangan River at Deramakot, Kinabatangan District, North Borneo. Holotype: FMNH 68001
Distribution: Kinabatangan River, Borneo, Malaysia (Inger & Chin, 1959).
Pseudomystus rugosus (Regan, 1913)
Liocassis rugosus Regan, 1913b: 552. Type locality: Poeh, Sarawak. Holotype: BMNH 1893.3.6.172.
Distribution: Sarawak.
Pseudomystus siamensis (Regan, 1913)
Liocassis siamensis Regan, 1913b: 550. Type locality: Bangpakong River, Siam. Holotype: BMNH 1897.10.8.126.
Leiocassis albicollaris Fowler, 1934b: 337, fig. 2. Type locality: Bangkok, Siam. Holotype: ANSP 60178.
Leiocassis bicolor Fowler, 1934a: 95, fig. 43. Type locality: Chieng Mai, North Siam. Holotype: ANSP 59284.
Distribution: Mekong and Chao Phraya River basins (Kottelat, 2001b), and peninsular Thailand.
Pseudomystus sobrinus Ng & Freyof, 2005
Pseudomystus sobrinus Ng & Freyof, 2005: 745, figs. 1, 2a. Type locality: Vietnam, Phu Yen province, stream about
15 km south of Tuy Hoa, 12º53'05"N, 109º23'70"E. Holotype: ZFMK 27167.
Distribution: Song Ba and nearby rivers, southern Vietnam (Ng & Freyhof, 2005).
Pseudomystus stenogrammus Ng & Siebert, 2005
Pseudomystus stenogrammus Ng & Siebert, 2005: 2, figs, 1, 3a [with captions identifying the species as Pseudomystus ornatus]. Type locality: Borneo: Kalimantan Tengah, Sungai Laung at Dessa Marwei. Holotype: MZB 6103.
Distribution: Barito River basin, southern Borneo (Ng & Siebert, 2005).
Remarks: The name Pseudomystus ornatus, which appears in the figure caption of the description of this species isclearly a lapsus calami for Pseudomystus stenogrammus and is thus a nomen nudum.
Pseudomystus stenomus (Valenciennes, 1840)
Bagrus stenomus Valenciennes, in Cuvier & Valenciennes, 1840a: 415 (307 of Strasbourg deluxe edition). Type
locality: Java. Holotype: RMNH 2986; previously unpublished illustration of holotype reproduced in Roberts(1993: fig. 61).
Leiocassis Ellenriederii Bleeker, 1860: 10. Type locality: Lahat, Palembang. Syntypes (3, 105–110 mm TL): Whereabouts unknown.
Distribution: South East Thailand, Kampuchea, Malay Peninsula, Sumatra, Java, Borneo.

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Pseudomystus vaillanti (Regan, 1913)
Liocassis vaillanti Regan, 1913b: 549. Type locality: Embouchure du Raoun, Kapuas basin, Borneo. Holotype:
RMNH 7840. Based on specimen identified in Vaillant (1902; fig. 8, 9) as Leiocassis moeschii; Vaillant’s illustration reproduced in Roberts (1989a: fig, 93).
Distribution: Kapuas River, known only from holotype (Roberts, 1989a).
RAMA Bleeker, 1858
Rama Bleeker, 1858b: 201, 205, 208. Type species: Rama buchanani Bleeker, 1858 (= Pimelodus rama Hamilton,
1822). Type by monotypy. Gender: Feminine.
Rama rama (Hamilton, 1822)
Pimelodus rama Hamilton, 1822: 176, 377, pl. 3 (fig. 55). Type locality: Brahmaputra River. No types known.
Rama buchanani Bleeker, 1858b: 201, 205, 208. Type locality: Brahmaputra River. No types known. Unneeded
replacement for Pimelodus rama Hamilton, 1822, apparently to avoid tautonymy.
Distribution: Brahmaputra River basin, India (Jayaram, 1999).
Remarks: Treated in some recent literature as synonym of Chandramara chandramara (Hamilton).
RITA Bleeker, 1854
Rita Bleeker, 1854c: 122. Type species: Rita buchanani Bleeker, 1854. Type by monotypy. Gender: Feminine.
Gogrius Day, 1867: 563. Type species: Gogrius sykesii Day, 1867. Type by monotypy. Gender: Masculine.
Key to species: Ferraris (1999b), Ng (2004i).
Rita chrysea Day, 1877
Rita chrysea Day, 1877 (in Day, 1875–78): 455, pl. 104 (fig. 1). Type locality: Orissa. Holotype: ZSI F498.
Distribution: Mahanadi River system, Orissa, Madhya Pradesh, India (Jayaram, 1999).
Remarks: See Jayaram (1960) for interpopulational variation.
Rita gogra (Sykes, 1839)
Phractocephalus gogra Sykes, 1839: 164. Type locality: Deccan, India. Type(s): Whereabouts unknown. Also proposed as new in Sykes (1840: 61); illustrated and described in more detail in Sykes (1841: 374, pl. 66, fig. 1).
Arius pavimentatus Valenciennes, in Cuvier & Valenciennes, 1840b: 94 (70 of Strasbourg deluxe edition). Type
locality: trouvé dans la collection de feu Jacquemont [= vicinity of Bombay, India]. Holotype: MNHN b-0684;
illustrated in Jacquemont (1835–1844: pl. 17, fig. 2).
Gogrius sykesii Day, 1867: 563. Type locality: Kurnool and the Deccan, India. Possible syntypes: ZSI 1200 (1), ZSI
3107 (1, lost), and types of “Pimelodus gogra, Sykes.” Unndeeded new name for Phractocephalus gogra
Sykes, 1839, apparently to avoid tautonymy.
Distribution: Rivers of Deccan, India, up to Krishna River basin (Jayaram, 1999).
† Rita grandiscutata Lydekker, 1886
† Rita grandiscutata Lydekker, 1886: 251, pl. 37, fig. 3. Type locality: Punjab, Siwalik Formation Lower Pliocene.
Holotype: at Indian Museum, Calcutta E. 157 (large basal bone of dorsal spine).
? † Chrysichthys (?) theobaldi Lydekker, 1886: 249, pl. 37, fig. 4. Type locality: Punjab, Siwalik Formation, Lower
Pliocene. Holotype: Indian Museum, Calcutta E. 158 (hind part of cranium).
Distribution: Punjab, Siwalik Formation Lower Pliocene.
Remarks: † Chrysichthys theobaldi was assigned to Rita by Jayaram (1955a: 121). Because the two names listed
here were applied to different parts of specimens that have been assigned to the same genus and both were
found in the same locality, I am tentatively placing the two names into synonymy and treat † Rita grandiscutata
as the valid name of the species.
Rita kuturnee (Sykes, 1839)
Phractocephalus kuturnee Sykes, 1839: 164. Type locality: Deccan, India. Type(s): Whereabouts unknown. Also
described as new in Sykes (1840: 61); illustrated and described in more detail in Sykes (1841: 372, pl. 65, fig.
3).
Arius hastatus Valenciennes, in Cuvier & Valenciennes, 1840b: 97 (72 of Strasbourg deluxe edition). Type locality:[India]. Holotype: MNHN b-0591. Illustrated in Jacquemont (1835–1844: pl. 18, fig. 2).

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Distribution: Peninsular India.
Rita macracanthus Ng, 2004
Rita macracanthus Ng, 2004i: 2, fig. 1. Type locality: Pakistan: Indus River at Attock. Holotype: UMMZ 237502.
Distribution: Indus River basin (Ng, 2004i).
Rita rita (Hamilton, 1822)
Pimelodus rita Hamilton, 1822: 165, 376, pl. 24 (fig. 53). Type locality: estuaries of Bengal. No types known.
Arius ritoides Valenciennes, in Cuvier & Valenciennes, 1840b: 92 (69 of Strasbourg deluxe edition). Type locality:
Bengale. Holotype: MNHN b-0683.
Rita Buchanani Bleeker, 1854c: 123, pl. 3 (fig. 1). Type locality: Hooghly, Calcutta. No types known. Unneeded
new name for Pimelodus rita Hamilton, 1822, apparently to avoid Stricklandian tautonymy.
Distribution: Rivers of northern India, Pakistan, Bangladesh and Nepal (Jayaram, 1999).
Rita sacerdotum Anderson, 1879
Rita sacerdotum Anderson, 1879: 864, pl. 79 (fig. 3). Type locality: Irrawaddy River at location of pagoda of Thingadow, 3rd Defile of Irrawaddy River, Burma. Holotype: BMNH 1875.8.4.7 (stuffed).
Distribution: Irrawaddy River basin, Myanmar (Ferraris, 1999b).
Remarks: Redescribed in Ferraris (1999b).
Species inquirenda, Rita
Arius cruciger Owen, 1853: 26. Type locality: Ganges. Type (s): Whereabouts unknown.
SPERATA Holly, 1939
Macrones Duméril, 1856: 484. Type species: Bagrus lamarrii Valenciennes, 1840. Type by monotypy. Gender: Masculine. Preoccupied by Macrones Newman, 1841, in Coleoptera; replaced by Aoria Jordan, 1917, and by Sperata Holly, 1939.
Aoria Jordan, 1919b: 341. Type species: Bagrus lamarrii Valenciennes, 1840. Type by being a replacement name.
Gender: Feminine. Replacement for Macrones, Duméril, 1856; preoccupied by Macrones Newman, 1841, in
Coleoptera. Aoria preoccupied by Aoria Baly, 1863, in Coleoptera; replaced by Aorichthys Wu, 1939, and byMacronichthys White & Moy-Thomas, 1940.
Sperata Holly, 1939: 143. Type species: Bagrus lamarrii Valenciennes, 1840. Type by being a replacement name.
Gender: Feminine. Holly stated that the name was intended as a replacement for Macrones, but incorrectlylisted Macrones vittata Bloch as type.
Aorichthys Wu, 1939: 131. Type species: Bagrus lamarrii Valenciennes, 1840. Type by being a replacement name.
Gender: Masculine. Replacement for Aoria Jordan, 1919, preoccupied by Aoria Baly, 1863, in Coleoptera.
Macronichthys White & Moy-Thomas, 1940: 505. Type species: Bagrus lamarrii Valenciennes, 1840. Type by beinga replacement name. Gender: Masculine. Replacement for Aoria Jordan, 1919, preoccupied by Aoria Baly,
1863, in Coleoptera.
Osteobagrus Jayaram, 1954: 529, 547. Type species: Pimelodus aor Hamilton, 1822. Type by original designation.
Gender: Masculine. Originally proposed as a subgenus of Mystus.
Remarks: Ferraris & Runge (1999: 400) concluded that Sperata has priority over Aorichthys. See also Jayaram(1968a, 1971, 1973b).
Revision: Ferraris & Runge (1999).
Sperata acicularis Ferraris & Runge, 1999
Sperata acicularis Ferraris & Runge, 1999: 403, figs. 2a, 3, 4. Type locality: Myanmar, Yangon Division, South
Oak-ka-lar-pa market (eastern Yangon). Holotype: CAS 209024.
Distribution: Irrawaddy, Bago and Tenasserim River systems, Myanmar (Ferraris & Runge, 1999).
Sperata aor (Hamilton, 1822)
Pimelodus aor Hamilton, 1822: 205, 379, pl. 20 (fig. 68). Type locality: Rivers of Bengal, and upper parts of theGangetic estuaries. No types known. [Illustration of type reproduced in Ferraris & Runge (1999: fig. 1)].
Bagrus aorides Jerdon, 1849: 336. Type locality: Cauvery River at Errode, southern India. No types known.

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FERRARISDistribution: Ganges River system and peninsular India south to Cauvery River basin (Ferraris & Runge, 1999).
Sperata aorella (Blyth, 1858)
Bagrus aorellus Blyth, 1858: 283. Type locality: Calcutta bazar. No types known.
Distribution: Ganges River, India and Bangladesh (Ferraris & Runge, 1999).
Sperata sarwari (Mirza, Nawaz & Javed, 1992)
Aorichthys aor sarwari Mirza, Nawaz & Javed, 1992: 211, fig. 1. Type locality: Ravi River at Head Balloki, Kasur
Dist., Pakistan. Holotype: GCM F-19.
Distribution: Indus River basin (Mirza, 2003).
Sperata seenghala (Sykes, 1839)
Platystoma seenghala Sykes, 1839: 164. Type locality: Deccan, India. Type(s): at BMNH. Also proposed as new inSykes (1840: 61); illustrated and described in more detail in Sykes (1841: 371, pl. 65, fig. 2) [illustration reproduced in Ferraris & Runge (1999: fig. 6)].
Bagrus Lamarrii Valenciennes, in Cuvier & Valenciennes, 1840a: 407 (302 of Strasbourg deluxe edition), pl. 415.
Type locality: [Ganges]. Holotype: MNHN a-9343 (mounted).[Illustration of holotype reproduced in Ferraris &Runge (1999: fig. 7)].
Bagrus aorinus Valenciennes, 1840, in Jacquemont, 1835–44: pl. 17 (fig. 1–1a). Type locality: India. Holotype:
Whereabouts unknown. Name available from plate caption. [Illustration of holotype reproduced in Ferraris &Runge (1999: fig. 8).]Distribution: Ganges River system and peninsular India at least as far south as the Krishna River (Ferraris & Runge,1999).
Genus inquirendum, Bagridae
Adelopeltis Dabry de Thiersant, 1872: 189, 190. Type species: Apparently not designated previously; Adelopeltis angusticeps Dabry de Thiersant, 1872, designated herein. Gender: Feminine.
Species inquirendae, Bagridae
Bagrus affinis Jerdon, 1849: 338. Type locality: Neighborhood of Madras, southern India. No types known.
Bagrus agricolus Jerdon, 1849: 339. Type locality: Ditches and inundated paddy fields in the Wynaad, southern India.
No types known.
Adelopeltis angusticeps Dabry de Thiersant, 1872: 189, pl. 47 (fig. 3). Type locality: Yang-tsee-kiang [China]. Type (s):Whereabouts unknown.
Pimelodus bilineatus Cuvier, 1829: 293. Type locality: Vishakhapatnam [Vizagapatam], India. Available from the following reference to a published illustration: “Pimel. bilineatus, Deddi-Jallah., Russel, 169 ” [= Russell (1803: pl.
169)].
Pseudobagrus eupogoides Wu, 1930b: 51, fig. 3. Type locality: Sze-Chwan, China. Holotype: Whereabouts unknown.
Name not treated in recent literature.
Pseudobagrus fui Miao, 1934: 217, fig. 45. Type locality: Chinkiang, southern Kiangsu, China. Holotype: Mus. Biol.
Lab. Sci. Soc. China 13170.
Liocassus hirsutus Herre, 1934: 285. Type locality: Market at Wuchow, Kwangsi, China. Holotype: SU 13884. Name nottreated in recent literature.
Aoria hoi Pellegrin & Fang, 1940: 341, fig. 2. Type locality: Foochow, China. Holotype: MNHN 1939-0033.
Leiocassis hwanghoensis Mori, 1933: 167, figs. 3–4. Type locality: Hwang-ho near Tsi-nan, Shantung, China. Holotype:
Whereabouts unknown.
Leiocassis kaifenensis Tchang, 1934: 41. Type locality: Kaifeng, Henan, China. Holotype: ASIZB [= ZMFMIB 8625].
Not treated since description.
Adelopeltis laticeps Dabry de Thiersant, 1872: 190, pl. 48 (fig. 5). Type locality: Yang-tsee-kiang [China]. No types
known. Not treated in recent literature.
Bagrus limbatus Richardson 1846: 283.Type locality: Canton. Holotype: at Cambridge Philosophical Inst. Sometimes

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incorrectly considered as Plotosus limbatus Valenciennes, but the description in Richardson is clearly that of abagrid.
Liocassis naso Garman, 1912: 123. Type locality: Hupei: Ichang, China. Holotype: MCZ 29847. Name not treated inrecent literature.
Bagrus (Bagrus) ramentosus Müller & Troschel, 1849: 7. Type locality: America [probably Java]. Holotype: ZMB 2949.
Treated as a synonym of Pseudobagrus aurantiacus or as valid as Pelteobagrus ramentosus.
Aoria rendahli Pellegrin & Fang, 1940: 340, fig. 1. Type locality: Ou Tchang, Hou Pé [Hubei], China. Holotype: MNHN
1939-0032. Name not treated in recent literature.
Leiocassis similis Nichols, 1926a: 1, fig. 1. Type locality: Min R., near Yenping (Nanping), Fukien Prov., China. Holotype: AMNH 8444.
Bagrus sinensis Brandt, in Mack, 1861: 195. Possibly name only and not available; from Sungatschi.[Not seen]
Leiocassis sinyanensis Fu, 1935: 178, fig. 1. Type locality: Szewangshan, Sinyang [Xinyang], s. Honan [Henan], China.
Holotype: Whereabouts unknown.
Hemibagrus taphrophilus Sauvage & Dabry de Thiersant, 1874: 6. Type locality: Canaux du Tschuang occidental, China.
Type(s) (100 mm): at MNHN.
Liocassis torosilabris Sauvage & Dabry de Thiersant, 1874: 7. Type locality: Yang-tsee-kiang, China. Holotype: MNHN
0000-5936.
Pseudobagrus wangi Miao, 1934: 215, fig. 44. Type locality: Chinkiang, southern Kiangsu, China. Holotype: Mus. Biol.
Lab. Sci. Soc. China 13173. Treated as valid in Pelteobagrus by Jayaram (1968b: 306).
Pseudobagrus wittenburgii Popta, 1911: 335, unnumbered figure. Type locality: Amur at Blagoveshchensk, Russia. Syntypes: SMNS 4390 (2).
† Aoria lacus [No other information, taken from Gayet & Meunier (2003)].
Distribution: Eocene; Hunan, southcentral China (Chang & Zhou, 1993).
Remarks: Considered to be Bagridae S.L. by Gayet & Meunier (2003).

CALLICHTHYIDAE Bonaparte, 1838
Callichthini Bonaparte, 1838: 131. Type genus: Callichthys Scopoli, 1777.
Aspidoradidi Hoedeman, 1952b: 4. Type genus: Aspidoras Ihering, 1907.
Cascaduridi Hoedeman, 1952b: 3. Type genus: Cascadura Ellis, 1913.
Corydoradinae Hoedeman, 1952b: 4. Type genus: Corydoras La Cepède, 1803.
Dianemidi Hoedeman, 1952b: 4. Type genus: Dianema Cope, 1871.
Hoplosterninae Miranda Ribeiro, 1959: 1. Type genus: Hoplosternum Gill, 1858.
Taxonomic summary: Isbrücker (2001b, Corydoradinae), Reis (2003).
Phylogeny: Reis (1998a); Britto (2003, Corydoradinae); Shimabukuro-Dias et al. (2004).
Biogeography: Reis (1998b).
8 genera, 194 species, one fossil species.
ASPIDORAS
Aspidoras Ihering, 1907: 30. Type species: Aspidoras rochai Ihering, 1907. Type by original designation. Gender:
Masculine.
Revision: Nijssen & Isbrücker (1976a).
Aspidoras albater Nijssen & Isbrücker, 1976
Aspidoras albater Nijssen & Isbrücker, 1976a: 115, fig. 6. Type locality: Brazil, Est. Goiás, Rio Tocantinzinha nearSão João da Aliança, 14º46'S, 47º30'W, Rio Tocantins system. Holotype: MZUSP 12991.
Distribution: Upper Tocantins River basin, Brazil (Reis, 2003).
Aspidoras belenos Britto, 1998
Aspidoras belenos Britto, 1998: 361, fig. 1. Type locality: Brazil: Estado do Mato Grosso, creek at Primavera do
Leste- Paranatinga road, 82 km N from Primavera do Leste, rio das Mortes basin, 15º03'S, 52º03'W. Holotype:

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FERRARISMNRJ 12433.
Distribution: Das Mortes River basin, Araguaia River drainage, Brazil (Reis, 2003).
Aspidoras brunneus Nijssen & Isbrücker, 1976
Aspidoras brunneus Nijssen & Isbrücker, 1976a: 116, fig. 7. Type locality: Brazil, Est. Mato Grosso, Serra do
Roncador, km 125 of the road Chavantina-Casximba. Holotype: ZMA 113588.
Distribution: Upper Araguaia River basin, Brazil (Reis, 2003).
Aspidoras carvalhoi Nijssen & Isbrücker, 1976
Aspidoras carvalhoi Nijssen & Isbrücker, 1976a: 117, fig. 8. Type locality: Brazil, Est. Ceará, Açude Canabrava,
Guaramiranga. Holotype: MNRJ 5230.
Distribution: Ceará State, Brazil (Reis, 2003).
Aspidoras depinnai Britto, 2000
Aspidoras depinnai Britto, 2000: 1049, fig. 1. Type locality: Brazil: Pernambuco State, creek at Amaraji-Primavera
road, rio Ipojuca basin, 08º21'S, 35º26'W. Holotype: MZUSP 56214.
Distribution: Ipojuca River basin, Brazil (Reis, 2003).
Aspidoras eurycephalus Nijssen & Isbrücker, 1976
Aspidoras eurycephalus Nijssen & Isbrücker, 1976a: 118, fig. 9. Type locality: Brazil, Est. Goiás, Rio Tocantins system, Corrego Vermelho into Rio das Almas, tributary of Rio Maranhão, about 15ºS, 49º30'W. Holotype: CAS
16010.
Distribution: Upper Tocantins River basin, Brazil (Reis, 2003).
Aspidoras fuscoguttatus Nijssen & Isbrücker, 1976
Aspidoras fuscoguttatus Nijssen & Isbrücker, 1976a: 118, fig. 10. Type locality: Brazil, Est. Mato Grosso, Rio
Paraná system, Córrego Corguinho, estrada da Três Lagoas (20º46'S, 51º43'W) — Conceição do Taboado.
Holotype: MZUSP 8573.
Distribution: Upper Paraná River basin, Brazil (Reis, 2003).
Aspidoras lakoi Miranda Ribeiro, 1949
Aspidoras lakoi Miranda Ribeiro, 1949a: 143, fig. 1. Type locality: Pequeno, córrego na floresta do Grotão, Fazenda
da Cachoeira, Município de Passos, Estado de Minas Gerais [Brazil]. Holotype: MNRJ 5292.
Distribution: Upper Paraná River basin, Brazil (Reis, 2003).
Aspidoras maculosus Nijssen & Isbrücker, 1976
Aspidoras maculosus Nijssen & Isbrücker, 1976a: 119, fig. 11. Type locality: Brazil, Est. Bahia, Rio Paiaia, into
headwaters of Rio Itapicurú, small, rocky, rapid stream from Serra Jacobina between Bom Fim and Jacobina,
about 11ºS, 40º30'W. Holotype: FMNH 54810.
Distribution: Itapicurú River basin, Brazil (Reis, 2003).
Aspidoras menezesi Nijssen & Isbrücker, 1976
Aspidoras menezesi Nijssen & Isbrücker, 1976a: 120, fig. 12. Type locality: Brazil, Est. Ceará, Rio Granjeiro at
Crato, 7º10'S, 39º25'W, tributary on left bank of Rio Salgado. Holotype: UMMZ 147336.
Distribution: Coastal rivers in Ceará State, Brazil (Reis, 2003).
Aspidoras microgalaeus Britto, 1998
Aspidoras microgalaeus Britto, 1998: 364, fig. 8. Type locality: Brazil: Estado do Mato Grosso, small tributary of
rio Culuene, km 86 of Paranatinga-Canarana road, rio Xingú basin, 14º00'S, 54º38'W. Holotype: MZUSP
51209.
Distribution: Xingu River basin, Brazil (Reis, 2003).
Aspidoras pauciradiatus (Weitzman & Nijssen, 1970)
Corydoras pauciradiatus Weitzman & Nijssen, 1970: 129, fig. 5. Type locality: Brazil, Rio Araguaia, near Aruanã,Est. Goiás (14º58'S, 51º04'W). Holotype: USNM 191625.
Distribution: Upper Araguaia River basin, Brazil (Reis, 2003).
Aspidoras poecilus Nijssen & Isbrücker, 1976
Aspidoras poecilus Nijssen & Isbrücker, 1976a: 121, fig. 13. Type locality: Brazil, Est. Mato Grosso, creek upstreamof village Porori, left bank of Rio Xingu, upper Rio Xingu. Holotype: IRSNB 560.
Distribution: Upper Xingu, Araguaia and Tocantins River basins, Brazil (Reis, 2003).

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Aspidoras psammatides Britto, Lima & Santos, 2005
Aspidoras psammatides Britto, Lima & Santos, 2005: 474, fig. 1. Type locality: Brazil, Estado da Bahia, municípiode Lençóis, rio Caldeirão, 12º39'33"S 41º22'12"W. Holotype. MNRJ 28407.
Distribution: Upper Paraguaçu River basin, eastern Brazil (Britto et al., 2005).
Aspidoras raimundi (Steindachner, 1907)
Corydoras raimundi Steindachner, 1907a: 84. Type locality: in dem Bächchen, welches bei Victoria in den Rio Parnahyba mündet [Brazil]. Lectotype: NMW 61110; designated by Nijssen & Isbrücker (1976a: 111).
Distribution: Parnaíba River basin, Brazil (Reis, 2003).
Aspidoras rochai Ihering, 1907
Aspidoras rochai Ihering, 1907: 30. Type locality: Fortaleza, capital of the state of Ceará [Brazil]. Lectotype:
MZUSP 2195; designated by Britski (1969: 206).
Distribution: Coastal rivers in Ceará State, Brazil (Reis, 2003).
Aspidoras spilotus Nijssen & Isbrücker, 1976
Aspidoras spilotus Nijssen & Isbrücker, 1976a: 123, fig. 14. Type locality: Brazil, Est. Ceará, Riacho dos Macacos,
tributary of Rio Acaráu. Holotype: ZMA 113590.
Distribution: Coastal rivers in Ceará State, Brazil (Reis, 2003).
Aspidoras taurus Lima & Britto, 2001
Aspidoras taurus Lima & Britto, 2001: 1011, fig. 1. Type locality: Brazil: Mato Grosso State, município de Alto
Garças, Fazenda Cabeceira do Itiquira, close to highway BR-364, rio Itiquira, rio Paraguay system, 16º56'S
53º32'W. Holotype: MZUSP 57154.
Distribution: Upper Paraguay River basin, Brazil (R. Reis, pers. commun., 2004).
Aspidoras velites Britto, Lima & Moreira, 2002
Aspidoras velites Britto, Lima & Moreira, 2002: 728, fig. 1. Type locality: Brazil. Estado do Mato Grosso. Município de Alto Araguaia, córrego Boiadeiro, km 487.08 of Ferronorte railroad, 17°20'01"S, 53°14'53"W. Holotype:
MZUSP 74447.
Distribution: Small tributaries of upper Araguaia River in Mato Grosso State, Brazil (Reis, 2003).
Aspidoras virgulatus Nijssen & Isbrücker, 1980
Aspidoras virgulatus Nijssen & Isbrücker, 1980c: 133, fig. 1. Type locality: Brazil: Est. Espírito Santo. Holotype:
MNRJ 5371.
Distribution: Coastal rivers in Espírito Santo State, Brazil (Reis, 2003).
CALLICHTHYS
Calichthys Scopoli, 1777: 451. Type species: Silurus callichthys Linnaeus, 1758. Gender: Masculine. Original spelling not used subsequently and spelling in prevailing usage, Callichthys, is adopted here following ICZN (1999:
Art. 33.3.1).
Callichthys Meuschen, 1778: 39. Type species: Silurus callichthys Linnaeus, 1758. Gender: Masculine.
Callichthys Linck, 1790: 32. Type species: Silurus callichthys Linnaeus, 1758. Gender: Masculine.
Cataphractus Bloch, 1794: 80. Type species: Silurus callichthys Linnaeus, 1758. Type by subsequent designation.
Gender: Masculine.
Phylogeny: Reis (1997).
Key to species: Lehmann & Reis (2004).
Remarks: Callichthys was first published by Gronow (1763: 127) in a work that was subsequently placed on the
ICZN Official Index of Rejected and Invalid Generic Names in Zoology, Opinion 261 (Melville & Smith,1987), which makes the name unavailable from that publication.
Callichthys callichthys (Linnaeus, 1758)
Silurus callichthys Linnaeus, 1758: 307. Type locality: in Americae rivulis. Syntypes: On Callichthys of Gronovius,
which may be based on BMNH 1853.11.12.194 (1, left half skin), and specimens from the Linnaean collection:
NRM 7015 (2), NRM 7018 (1), ZMUU 55 (1).
Callichthys asper Quoy & Gaimard, 1824: 232. Type locality: ruissean qui coule devant la ferme de Santa-Anna, surle chemin qui conduit de Rio de Janeiro à la colonie suisse [Brazil]. Syntypes: MNHN 0000-1951 (2).

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Cataphractus depressus Swainson, 1839: 304. Type locality: [Not stated]. Type(s): Based on Bloch (1794: pl. 377)[= Cataphractus callichthys (Linnaeus)].
Callichthys læviceps Valenciennes, in Cuvier & Valenciennes, 1840b: 309 (229 in the Strasbourg deluxe edition).
Type locality: La Mana, .... Brésil. Syntypes: MNHN 0000-1947 (1), MNHN 0000-4261 (1).
Callichthys cælatus Valenciennes, in Cuvier & Valenciennes, 1840b: 308 (229 in the Strasbourg deluxe edition).
Type locality: Bahia, .... Rio-Janéiro. Syntypes: MNHN 0000-4258 (2), MNHN 0000-4259 (1).
Callichthys loricatus Gronow, in Gray, 1854: 157. Type locality: Americes Meridionalis rivulis. Syntypes: BMNH
1853.11.12.194 (1) and several literature sources.
Callichthys kneri Gill, 1858: 394. Type locality: Trinidad. Type(s): Whereabouts unknown, apparently not at
USNM.
Callichthys tamoata Bleeker, 1864a: 22. Type locality: Surinama. Syntypes (10, 114– 250 mm): at RMNH, andmany literature sources.
Callichthys affinis Günther, 1864: 226. Type locality: Rio Grande [Brazil]. Syntypes: BMNH 1861.5.7.1–4 (4).
Callichthys arcifer Hensel, 1868: 373. Type locality: Rio de Janeiro. Holotype: ZMB 7433.
Callichthys hemiphractus Hensel, 1868: 374. Type locality: Costa da Serra [Rio Grande do Sul, Brazil]. Holotype:
ZMB 7434.
Callichthys callichthys bolteni Hoedeman, 1952b: 9. Type locality: Surinam. Holotype: ZMA 100303a.
Callichthys callichthys demararae Hoedeman, 1952b: 10. Type locality: Demarara, British Guiana. Holotype: ZMA
100304.
Distribution: Most Cis-Andean South American river drainages north of Buenos Aires (Reis, 2003).
Callichthys fabricioi Román-Valencia, Lehmann-A & Muñoz, 1999
Callichthys fabricioi Román-Valencia, Lehmann-A & Muñoz, 1999: 54, fig. 1. Type locality: Colombia, Departamento del Cauca, Municipio de Buenos Aires, Zanjón Bagazal afluente del Río Cauca 980 meters (03º03'59"Ny 76º33'55"W). Holotype: IUQ 295.
Distribution: Cauca River basin, Colombia (Reis, 2003).
Callichthys oibaensis Ardilia Rodríguez, 2005
Callichthys oibaensis Ardilia Rodríguez, 2005: 32, figs. 1–6. Type locality: [Colombia, Vereda Las Monjas, río
Oibita, trib. of río Suárez]. Holotype: CAR 251.
Distribution: Suárez River basin, Colombia.
Callichthys serralabium Lehmann & Reis, 2004
Callichthys serralabium Lehmann & Reis, 2004: 339, fig. 1. Type locality: Venezuela: Estado Amazonas: La Esmeralda: Laguna Tonina, 1 km upstream from La Esmeralda, upper Orinoco (03º08'00"N, 065º32'00"W). Holotype:
MBUCV V-30980.
Distribution: Upper Orinoco and upper Negro River basins, Venezuela and Brazil (Lehmann & Reis, 2004: 341).
CORYDORAS
Corydoras La Cepède, 1803: 147. Type species: Corydoras geoffroy La Cepède, 1803. Type by monotypy. Gender:
Masculine.
Cordorinus Rafinesque, 1815: 89. Type species: Corydoras geoffroy La Cepède, 1803. Type by being a replacement
name. Gender: Masculine. Unjustified emendation of Corydoras La Cepède.
Hoplisoma Swainson, 1838: 336. Type species: Cataphractus punctatus Bloch, 1794. Type by original designation.
Gender: Neuter.
Brochis Cope, 1871b: 112. Type species: Brochis coeruleus Cope, 1872. Type by subsequent designation by Cope
(1872a). Gender: Feminine. Appeared first without species, more fully described in Cope (1872a) with thedescription of two new species; type designated by Cope (description of Brochis dipterus includes reference to
B. coeruleus as type).
Chaenothorax Cope, 1878: 679. Type species: Chaenothorax bicarinatus Cope, 1878. Type by original designation.
Gender: Masculine.
Gastrodermus Cope, 1878: 681. Type species: Corydoras elegans Steindachner, 1877. Type by subsequent designation. Gender: Masculine. Type apparently first designated by Gosline (1940b: 10).

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Osteogaster Cope, 1894: 102. Type species: Corydoras eques Steindachner, 1877. Type by original designation.
Gender: Feminine.
Microcorydoras Myers, 1953: 270. Type species: Corydoras hastatus Eigenmann & Eigenmann, 1888. Type byoriginal designation. Gender: Masculine.
Review: Nijssen & Isbrücker (1983b, Colombia); Castro (1987, Colombia); Nijssen & Isbrücker (1986b, Peru andEcuador).
Taxonomic summary: Nijssen & Isbrücker (1970), species formerly placed in Brochis; Nijssen & Isbrücker (1980b),
Isbrücker (2001b).
Remarks: Brochis Cope was recently placed in synonymy of Corydoras by Britto (2003), while Scleromystax was
removed and treated as a valid genus.
Corydoras acrensis Nijssen, 1972
Corydoras acrensis Nijssen, 1972: 416, fig. 1. Type locality: Brazil, Est. Acre, Furo do Lago São Francisco, tributary at left bank of Rio Juruá, upstream of confluence with Mao at Cruzeiro do Sul. Holotype: IRSNB 505.
Distribution: Juruá River basin, Brazil (Reis, 2003).
Corydoras acutus Cope, 1872
Corydoras acutus Cope, 1872a: 281. Type locality: Ambyiacu River [now: Shansho Caño, Loreto, Peru, by neotype
designation]. Neotype: SU 33307, designated by Nijssen & Isbrücker (1980b: 215); original syntypes ANSP
8292–8293 lost.
Distribution: Amazon River basin in northern Peru and Ecuador (Reis, 2003).
Corydoras adolfoi Burgess, 1982
Corydoras adolfoi Burgess, 1982: 15, fig. 1. Type locality: Small tributary of the upper Rio Negro on the equator
near São Gabriel da Cachoeira, Brazil. Holotype: MZUSP 26641.
Distribution: Negro River basin, Brazil (Reis, 2003).
Corydoras aeneus (Gill, 1858)
Hoplosoma aeneum Gill, 1858: 403. Type locality: Island of Trinidad, West Indies. Lectotype: USNM 1116, designated by Nijssen & Isbrücker (1980b: 208).
Corydoras microps Eigenmann & Kennedy, 1903: 506. Type locality: From a small lagoon, half dry, near the Rio
Branco (Mato Grosso, Brazil). Holotype: CAS 20662.
Corydoras venezuelanus Ihering, 1911: 383. Type locality: Río Cabriales, Valencia, Est. Carabobo, Venezuela. Lectotype: MZUSP 146, designated by Britski (1969: 208).
Corydoras macrosteus Regan, 1912a: 219. Type locality: Rio Piracicaba, San Paulo, Brazil. Lectotype: BMNH
1905.6.9.12, designated by Nijssen & Isbrücker (1980b: 208).
Corydoras schultzei Holly, 1940: 111. Type locality: Aquarium specimen (said to be from very small water-coursesof the Amazon).
Distribution: Widely distributed in South America on the eastern side of the Andes, from Colombia and Trinidad to
La Plata River basin (Reis, 2003).
Remarks: Isbrücker (2001b) treated all of the synonyms listed above as valid, or “doubtfully valid” species, but did
not comment on the distribution of these species.
Corydoras agassizii Steindachner, 1877
Corydoras Agassizii Steindachner, 1877a: 138, pl. 12 (fig. 2). Type locality: Amazonenstrom bei Tabatinga. Lectotype: NMW 61112, designated by Nijssen & Isbrücker (1980b: 195).
Distribution: Amazon River basin near border between Peru and Brazil (Reis, 2003).
Corydoras albolineatus Knaack, 2004
Corydoras albolineatus Knaack, 2004: 73, figs. 13–17. Type locality: Bolivien, Departamento Santa Cruz, Provincia
J.M. Velasco, Raum Piso firme, System río Itenez, río Paragua (Mündungsgebiet, NW-Ufer), 13º31.989'S,
61º50.295'W. Holotype: MTD F 28470.
Distribution: Itenez River basin, Bolivia (Knaack, 2004).
Corydoras amandajanea Sands, 1995
Corydoras amandajanea Sands, 1995: 10, fig. 1. Type locality: Rio Miuã system of streams, possibly the Uarinabe
stream (Miuã system) 10–12 kilometers away from its confluence of the upper Negro River, Brazil. Holotype:

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Distribution: Upper Negro River basin, Brazil (Reis, 2003).
Corydoras amapaensis Nijssen, 1972
Corydoras amapaensis Nijssen, 1972: 417, figs. 2, 3. Type locality: Brazil, Est. Amapa, Cachoera Creek at rightbank of Río Amapari, 4 km downstream of Casa do 7. Holotype: IRSNB 476.
Distribution: Oyapock and Amapari rivers in Amapá State in Brazil and French Guiana (Reis, 2003).
Corydoras ambiacus Cope, 1872
Corydoras ambiacus Cope, 1872a: 280. Type locality: Ambyiacu River. Holotype: ANSP 8291.
Corydoras grafi Holly, 1940: 108. Type locality: Aquarium specimen (said to be from very small water-courses of
the Amazon). Holotype: Münchner Tierpark AG 92/139 (reportedly lost, but possibly NMW 84287).
Corydoras melanistius longirostris Hoedeman, 1952a: 17, fig. 13. Type locality: Amazon area. No types known.
Distribution: Upper Amazon River basin of Brazil, Colombia, and Peru (Reis, 2003).
Corydoras amphibelus Cope, 1872
Corydoras amphibelus Cope, 1872a: 282. Type locality: Ambyiacu River. Holotype: ANSP 8290.
Distribution: Upper Amazon River basin in Peru (Reis, 2003).
Corydoras approuaguensis Nijssen & Isbrücker, 1983
Corydoras approuaguensis Nijssen & Isbrücker, 1983c: 73, fig. 1. Type locality: fleuve Approuague. Holotype:
ZMA 119098.
Distribution: Approuague River basin, French Guiana (Reis, 2003).
Corydoras araguaiaensis Sands, 1990
Corydoras araguaiaensis Sands, 1990: [1], unnumbered figure. Type locality: Rio Araguaia, Brazil. Holotype: at
RMNH.
Distribution: Araguaia River basin, Brazil (Reis, 2003).
Corydoras arcuatus Elwin, 1939
Corydoras arcuatus Elwin, 1939: 126, pl. 3. Type locality: Amazon River (aquarium specimen). Holotype: BMNH
1939.3.3.1.
Distribution: Upper Amazon River basin (Reis, 2003).
Corydoras areio Knaack, 2000
Corydoras areio Knaack, 2000b: 47, unnumbered figure. Type locality: Brasilien, Mato Grosso (östlich von
Cuiaba), in Fließgewässern des Córr. Areio-Systems, in der Umgebung von Vila Nova, an der Straße 373, ca.
8.5 kilometers und ca.13.5 km von der Ortschaft aus entfernt in Richtung Poxoreo. Holotype: ZMB 33113.
Distribution: Upper Paraguay River basin, Brazil (Reis, 2003).
Corydoras armatus (Günther, 1868)
Callichthys armatus Günther, 1868a: 476. Type locality: Xeberos and Huallaga [Peru]. Lectotype: BMNH
1867.6.13.51, designated by Nijssen & Isbrücker (1980b: 194). Illustrated and described in more detail in
Günther (1868b: 231, fig. 1).
Distribution: Upper Amazon River basin, Peru (Reis, 2003).
Corydoras atropersonatus Weitzman & Nijssen, 1970
Corydoras atropersonatus Weitzman & Nijssen, 1970: 123, fig. 2. Type locality: Ecuador, Río Conambo at mouth of
Río Shione (about 2º00'S, 76º30'W), Río Tigre system, Est. Pastaza. Holotype: USNM 204359.
Distribution: Upper Amazon River basin, Ecuador and Peru (Reis, 2003).
Corydoras aurofrenatus Eigenmann & Kennedy, 1903
Corydoras aurofrenatus Eigenmann & Kennedy, 1903: 507. Type locality: Aguada, near Arroyo Trementina [Paraguay]. Holotype: CAS 20661.
Distribution: Paraguay River basin, Paraguay (Reis, 2003).
Corydoras axelrodi Rössel, 1962
Corydoras axelrodi Rössel, 1962c: 335, figs. 1, 2. Type locality: Columbien, Rio Meta. Holotype: SMF 5700.
Distribution: Meta River basin, Colombia (Reis, 2003).
Corydoras baderi Geisler, 1969
Corydoras baderi Geisler, 1969: 354, fig. 1. Type locality: Brasilien, Staat Para, Rio Paru de Oeste und Bäche bei

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der Missionsstation Tirió (2º12'N, 55º59'W). Holotype: SMF 9651.
Corydoras oelemariensis Nijssen, 1970: 29, fig. 14. Type locality: Creek at left bank of Oelemari River, near airstrip, Marowijne, Suriname. Holotype: IRSNB 472.
Distribution: Pará State in Brazil and Maroni River in Suriname (Reis, 2003).
Corydoras bicolor Nijssen & Isbrücker, 1967
Corydoras bicolor Nijssen & Isbrücker, 1967: 36, pl. 3 (fig. 3). Type locality: Sipaliwini River. Holotype: ZMA
104627.
Distribution: Coastal rivers of Suriname (Reis, 2003).
Corydoras bifasciatus Nijssen, 1972
Corydoras bifasciatus Nijssen, 1972: 420, fig. 4. Type locality: Brazil, Est. Para, creek at left bank of Río Cururu,
tributary of upper Río Tapajós near Maloca do Conzalo (about 7º45'S, 58º00'W). Holotype: IRSNB 485.
Distribution: Lower Amazon River basin, Brazil (Reis, 2003).
Corydoras bilineatus Knaack, 2002
Corydoras bilineatus Knaack, 2002a: 52, unnumbered figure. Type locality: Santa Cruz, Santiesteban, Chané, Restgewässer eines großen Überflutungs-Nährungsgebietes zwischen den Flüssen Rio Piray und Rio Yapacarie unddem Rio Grande o Guapay mit abfluss (Regenzeit) in den Arroyo Chanè. Holotype: MTD F 25925.
Distribution: Madeira River basin, Bolivia (Reis, 2003).
Corydoras blochi Nijssen, 1971
Corydoras blochi Nijssen, 1971: 92, fig. 2. Type locality: Guyana, district Essequibo, Moco Creek near Lehtem,
tributary of Rio Tacutú, Rio Branco system. Holotype: FMNH 75951.
Distribution: Venezuela, Guyana, and northern Brazil (Reis, 2003).
Corydoras boehlkei Nijssen & Isbrücker, 1982
Corydoras boehlkei Nijssen & Isbrücker, 1982: 139, fig. 1. Type locality: Venezuela, Edo Bolivar, Río Cuchime(Cusimo), about 20 miles upstream from junction of Río Caura —Río Erebato (Entre Rios) at 5º45'N, 64º24'W.
Holotype: ANSP 148097.
Distribution: Caura River basin in the Orinoco River drainage, Venezuela (Reis, 2003).
Corydoras boesemani Nijssen & Isbrücker, 1967
Corydoras boesemani Nijssen & Isbrücker, 1967: 37, pl. 4 (fig. 1). Type locality: Little tributaries of Gran-Rio
between Ligolio and Awaradam Falls [Suriname]. Holotype: RMNH 25316.
Distribution: Coastal rivers in Suriname (Reis, 2003).
Corydoras bondi Gosline, 1940
Corydoras bondi Gosline, 1940b: 20. Type locality: Rio Yuruari, 3 km east of El Callao, Venezuela. Holotype: SU
35065.
Distribution: Rio Yuruari in Venezuela, Corantijn and Rupununi River basins, Guianas (Reis, 2003).
Corydoras breei Isbrücker & Nijssen, 1992
Corydoras breei Isbrücker & Nijssen, 1992: 10, fig. 1. Type locality: Surinam, Corantijn river system, KabaleboRiver, Avanavero Falls. Holotype: RMNH 31912.
Distribution: Corantijn River basin, Suriname (Reis, 2003).
Corydoras brevirostris Fraser-Brunner, 1947
Corydoras melanistius brevirostris Fraser-Brunner, 1947: 244. Type locality: aquarium specimen [said to be fromthe Orinoco]. Holotype: BMNH 1946.10.10.1.
Distribution: Orinoco River basin, Venezuela, and coastal drainages in Suriname (Reis, 2003).
Corydoras britskii (Nijssen & Isbrücker, 1983)
Brochis britskii Nijssen & Isbrücker, 1983a: 179, fig. 1. Type locality: Brazil, Est. Mato Grosso, Lagoas Marginais,
rodovia Transpantaneira, Poconé (16º15'S, 56º37'W). Holotype: MZUSP 26811.
Distribution: Upper Paraguay River basin, Brazil (Reis, 2003).
Corydoras burgessi Axelrod, 1987
Corydoras burgessi Axelrod, 1987: 22, figures on pp. 22– 23. Type locality: reportedly from the Rio Unini, tributaryof the Rio Negro, Amazonas, Brazil. Holotype: MZUSP 37692.
Distribution: Upper Negro River basin, Brazil (Reis, 2003).

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Corydoras carlae Nijssen & Isbrücker, 1983
Corydoras carlae Nijssen & Isbrücker, 1983c: 76, fig. 5. Type locality: Petit Arroyo, affluent rive gauche du Rio
Iguazu, route no. 101 entre Porto Iguazu et Bernardino de Irigoyen, 50 km de Puerto Iguazu, bassin du Paraná.
Holotype: IRSNB 688.
Distribution: Lower Iguazu River basin, Argentina (Reis, 2003).
Corydoras caudimaculatus Rössel, 1961
Corydoras caudimaculatus Rössel, 1961: 49, pl. 5 (fig. 1). Type locality: Brasilien, oberer Rio Guaporé (=Itenez).
Holotype: SMF 5291.
Distribution: Guaporé River basin, Brazil (Reis, 2003).
Corydoras cervinus Rössel, 1962
Corydoras cervinus Rössel, 1962b: 31, fig. 1. Type locality: Brasilien, oberer Rio Guaporé (=Itenez). Holotype:
SMF 5460.
Distribution: Guaporé River basin, Brazil (Reis, 2003).
Corydoras cochui Myers & Weitzman, 1954
Corydoras cochui Myers & Weitzman, 1954: 93. Type locality: Santa Maria Nova, Rio Araguaya, State of Goyaz,
Brazil. Holotype: SU 47656.
Distribution: Upper Araguaia River basin, Brazil (Reis, 2003).
Remarks: Redescribed in Weitzman (1956).
Corydoras concolor Weitzman, 1961
Corydoras concolor Weitzman, 1961: 105, fig. 1. Type locality: Las Mangas, in a tributary to the Río Parguaza,
western part of the State of Bolivar, Venezuela, 6º20'N, 67º10'W. Holotype: SU 54131.
Distribution: Orinoco River basin, Venezuela (Reis, 2003).
Corydoras condiscipulus Nijssen & Isbrücker, 1980
Corydoras condiscipulus Nijssen & Isbrücker, 1980d: 494, fig. 1. Type locality: French Guiana, Cumuri Creek atleft bank of Oyapock River, upstream of first rapids of Grande Roche, southwest of village Oiapoque. Holotype:IRSNB 617.
Distribution: Oyapock River basin, Brazil and French Guiana (Reis, 2003).
Corydoras copei Nijssen & Isbrücker, 1986
Corydoras copei Nijssen & Isbrücker, 1986a: 70, fig. 11. Type locality: Loreto, cours inférieur du Río Huytoyacu,
près du village Nuevo Progresso, affluent de la rive droite du Rio Pastaza, environ 40 km N. du Lago Rimachi(4º13'S, 76º38'W). Holotype: ZMA 119305.
Distribution: Pastaza River basin in Loreto, Peru (Reis, 2003).
Corydoras coppenamensis Nijssen, 1970
Corydoras bondi coppenamensis Nijssen, 1970: 19, fig. 8. Type locality: Creek at left bank of Coppename River(3º52'N, 56º55'W) [Suriname]. Holotype: ZMA 105877.
Distribution: Coppename River basin, Suriname (Reis, 2003).
Corydoras coriatae Burgess, 1997
Corydoras coriatae Burgess, 1997: 140, unnumbered figure. Type locality: Río Aguaytia, a branch of the Río Ucayali, in jungles of Peru west of the city of Pucallpa. Holotype: MUSM 10701.
Distribution: Ucayali River basin, Peru (Reis, 2003).
Corydoras crimmeni Grant, 1997
Corydoras crimmeni Grant, 1997: 44, fig. 3. Type locality: Aquarium specimen, said to be from Brazil, Roraima
State, near the city of Boa Vista, possibly from the Rio Branco. Holotype: MZUSP 52490.
Distribution: Negro River basin, Brazil (Reis, 2003).
Corydoras cruziensis Knaack, 2002
Corydoras cruziensis Knaack, 2002c: 63, unnumbered figure. Type locality: kleinerer Fließgewässer in Santa Cruz,
Guarayos, A. Puente [Bolivia]. Holotype: MTD F 25930.
Distribution: Madeira River basin in Santa Cruz and Beni Departments of Bolivia (Reis, 2003).
Corydoras crypticus Sands, 1995
Corydoras crypticus Sands, 1995: 11, fig. 4. Type locality: Rio Miuã system of streams, possibly the Uarinabe

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stream, 10–12 kilometers away from its confluence of the upper Negro River, Brazil. Holotype: LIVCM
1994.4.29.
Distribution: Upper Negro River basin, Brazil (Reis, 2003).
Corydoras davidsandsi Black, 1987
Corydoras davidsandsi Black, 1987: 74, figure on p. 74. Type locality: Reportedly from the Rio Unini, Brazil. Holotype: MZUSP 38632.
Distribution: Negro River basin, Brazil (Reis, 2003).
Corydoras delphax Nijssen & Isbrücker, 1983
Corydoras delphax Nijssen & Isbrücker, 1983b: 55, fig. 3. Type locality: Guainía, Río Inírida system, Caño Bacón,
Pueblo Bretania (3º43'N, 67º59'W) [Colombia]. Holotype: NRM 26073.
Distribution: Inírida River and upper Orinoco River basin, Colombia (Reis, 2003).
Corydoras difluviatilis Britto & Castro, 2002
Corydoras difluviatilis Britto & Castro, 2002: 1007, fig. 1. Type locality: Brazil: São Paulo, Município de Santa Rita
do Passa Quatro, Parque Estadual de Vassununga, Glepa Pé-do-Gigante, córrego Paulicéia, tributary of rio
Pardo, rio Paraná system, 21º38'S, 47º38'W. Holotype: MZUSP 75268.
Distribution: Upper Paraná and Grande River basins, Paraná River drainage, and Jequitaí, Preto, and upper Da Prata
rivers, São Francisco drainage, Brazil (Reis, 2003).
Corydoras diphyes Axenrot & Kullander, 2003
Corydoras diphyes Axenrot & Kullander, 2003: 258, fig. 5. Type locality: Paraguay: Departamento Caaquazú: río
Paraná drainage, small sream (arroyo) at Estancia María Belén, 8 km from Colonel Patricio Colman,
25º40'13"S, 55º5'52"W. Holotype: MNHNP uncat (ex NRM 43483).
Distribution: Monday River and Acaray River, Paraná River basin, Paraguay (Axenrot & Kullander, 2003).
Corydoras duplicareus Sands, 1995
Corydoras duplicareus Sands, 1995: 12, fig. 6. Type locality: Rio Poranga, 8 kilometers north of the Nobua oba,
prior to where it joins a small tributary of the upper Negro River, Brazil. Holotype: LIVCM 1994.4.36.
Distribution: Upper Negro River basin, Brazil (Reis, 2003).
Corydoras ehrhardti Steindachner, 1910
Corydoras ehrhardti Steindachner, 1910a: 60. Type locality: Flußgebiet des Jaraguá [Santa Catarina, Brazil]. Lectotype: NMW 61104, designated by Nijssen & Isbrücker (1980b: 205).
Corydoras meridionalis Ihering, 1911: 380. Type locality: Colonia Hansa, Est. Santa Catharina [Brazil]. Type(s):Whereabouts unknown.
Distribution: Coastal rivers in Santa Catarina and Paraná States and Iguaçu River basin, Brazil (Reis, 2003).
Corydoras elegans Steindachner, 1877
Corydoras elegans Steindachner, 1877a: 141. Type locality: Teffé [Brazil]. Lectotype: NMW 46729, designated by
Nijssen & Isbrücker (1980b: 213).
Corydoras pestai Holly, 1940: 107. Type locality: in kleinen und kleinsten Wasserläufen der Nähe des Amazonenstromes. Holotype: Münchner Tierpark A.G. 81/1938 (destroyed).
Distribution: Upper Amazon River basin (Reis, 2003).
Remarks: Corydoras pestai Holly, 1940, treated as valid in Isbrücker (2001b: 232).
Corydoras ellisae Gosline, 1940
Corydoras ellisae Gosline, 1940b: 17. Type locality: Arroyo Poná, Sapucay, Paraguay. Holotype: FMNH 57361.
Distribution: Paraguay River basin, Paraguay (Reis, 2003).
Corydoras ephippifer Nijssen, 1972
Corydoras ephippifer Nijssen, 1972: 422, fig. 5. Type locality: Brazil, Est. Amapa, Cachoera Creek at right bank of
Río Amapari, 4 kilometers downstream of Casa do 7. Holotype: IRSNM 507.
Distribution: Amapá State, Brazil (Reis, 2003).
Corydoras eques Steindachner, 1877
Corydoras eques Steindachner, 1877a: 140, pl. 12 (fig. 3). Type locality: Amazonenstrom bei Teffé, Cudajas Brazil[restricted to Cudajas by lectotype designation]. Lectotype: NMW 61111, designated by Nijssen & Isbrücker
(1980b: 208).

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Corydoras esperanzae Castro, 1987
Corydoras esperanzae Castro, 1987: 34, fig. 2. Type locality: Caño Orocue at Orocue, Río Meta, Intendencia deCasanare (4º49'N, 71º20'W) [Colombia]. Holotype: UBJTL MM275.
Distribution: Meta River basin, Colombia (Reis, 2003).
Corydoras evelynae Rössel, 1963
Corydoras evelynae Rössel, 1963: 360, fig. 4. Type locality: Brazilien, oberer Rio Solimoes. Holotype: SMF 6107.
Distribution: Amazon River basin, Brazil (Reis, 2003).
Corydoras filamentosus Nijssen & Isbrücker, 1983
Corydoras filamentosus Nijssen & Isbrücker, 1983c: 77, fig. 6. Type locality: Bassin de la rivière Corantijn, affluent
du Sisa Creek, versant nord, environ 700 m en aval du croisement de Amatopo avec Camp Geologie Road,
03º42'N, 57º42'O [Suriname]. Holotype: USNM 225536.
Distribution: Corantijn River basin, Suriname (Reis, 2003).
Corydoras flaveolus Ihering, 1911
Corydoras flaveolus Ihering, 1911: 386. Type locality: Pequenos riachos affluentes do rio Piracicaba, acima do
Salto, Est. S. Paulo [Brazil]. Lectotype: MZUSP 424, designated by Britski (1969: 207).
Distribution: Upper Paraná River basin, Brazil (Reis, 2003).
Corydoras fowleri Böhlke, 1950
Corydoras fowleri Böhlke, 1950: 26. Type locality: Chancho Caño, near Pebas (3º10'S, 71º46'W), Peruvian Amazon, Peru. Holotype: SU 16115.
Distribution: Western Amazon River basin (Reis, 2003).
Corydoras garbei Ihering, 1911
Corydoras garbei Ihering, 1911: 382. Type locality: Rio São Francisco, Est. da Bahia [Brazil]. Lectotype: MZUSP
711, designated by Britski (1969: 207) for C. garbei Ihering, not C. garbei of Miranda Ribeiro (1937) [seeRemarks].
Distribution: São Francisco River basin, Brazil (Reis, 2003).
Remarks: An account of Corydoras garbei appeared in Miranda Ribeiro, (1937: 54) that has been treated by some as
an independent proposal of a new name for this species. Miranda Ribeiro (1953: 394) designated a lectotype forthis name and Britski (1969: 207) regarded the name as a homonym and synonym of Corydoras garbei Ihering,
1911. However, it appears that the account was a repeat of the earlier description of the species and, on thatbasis, treated by Reis (2003) as a lapsus and not a new name proposal.
Corydoras geoffroy La Cepède, 1803
Corydoras geoffroy La Cepède, 1803: 147. Type locality: Suriname, district Brokopondo, Suriname River system,
Marchall Creek, east of the road Paranam-Afobaka, 1.5 km North of Marchall village, by Neotype designation.
Neotype: ZMA 106017, designated by Isbrücker (2000: 17).
Corydoras octocirrus Nijssen, 1970: 26, fig. 12. Type locality: Marchall Creek, East of the road Paranam-Afobaka,
1.5 km North of Marchall village, Suriname River system, Suriname. Holotype: ZMA 106017.
Distribution: Coastal rivers of Suriname and French Guiana (Reis, 2003).
Remarks: Isbrücker (1999a, 2000) discussed the taxonomic history of this species.
Corydoras geryi Nijssen & Isbrücker, 1983
Corydoras bolivianus Nijssen & Isbrücker, 1983c: 75, fig. 3. Type locality: Plaine inondée, à quelques kilbmetres deTrinidad (14º46'S, 65º50'O) Bassin du Rio Mamoré [Beni Prov., Bolivia]. Holotype: MNHN 1983-0528.
Corydoras geryi Nijssen & Isbrücker, 1983c: 78, fig. 7. Type locality: Plaine inondée, à quelques kilomètres de Trinidad (14º46'S, 65º50'O), bassin du Rio Mamoré [Beni Prov., Bolivia]. Holotype: MNHN 1983-0529.
Distribution: Mamoré River basin, Bolivia (Reis, 2003).
Remarks: Isbrücker (1999b: 58) acted as first reviser in placing Corydoras bolivianus in the synonymy of C. geryi.
Corydoras gomezi Castro, 1986
Corydoras gomezi Castro, 1986b: 34, fig. 1. Type locality: Lakes of the Isla de Mocagua, Rio Amazonas/Solimões
near Leticia, Comisaría del Amazonas (3º54'S, 70º14'W) [Colombia]. Holotype: UBJTL MM536.
Distribution: Amazon River basin near the border of Colombia and Brazil (Reis, 2003).

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Corydoras gossei Nijssen, 1972
Corydoras gossei Nijssen, 1972: 424, fig. 6. Type locality: Brazil, Est. Rondonia, creek near Guajará Mirim(10º50'S, 65º21'W), Río Mamoré system. Holotype: IRSNB 487.
Distribution: Mamoré River basin, Brazil (Reis, 2003).
Corydoras gracilis Nijssen & Isbrücker, 1976
Corydoras gracilis Nijssen & Isbrücker, 1976c: 91, fig. 1 (unnumbered). Type locality: Brazil, Est. Pará, tributary of
Rio Tapajós, 66.5 kilometers west of Itaituba, 4º15'S, 55º56'W. Holotype: ZMA 114687. Type locality correctedin Nijssen & Isbrücker (1980b: 214) to Rio Jauna (= R. Juma) at Trans-Amazonica highway, about 6º09'S,
59º55'W, tributary of Rio Aripuaña, Amazonas, Brazil.
Distribution: Middle and lower Amazon River basin, Brazil (Reis, 2003).
Corydoras griseus Holly, 1940
Corydoras griseus Holly, 1940: 110. Type locality: in kleinen und kleinsten Wasserläufen der Nähe des Amazonenstromes [apparently in error, see Remarks]. Holotype: Münchner Tierpark A.G. 72/1938 (lost).
Corydoras griseus de weyeri Meinken, 1957: 5, unnumbered figure. Type locality: British-Guiana. Lectotype: ZMH
H1186, designated by, and illustrated in, Nijssen & Isbrücker (1967: 35, pl. 3, fig. 2).
Distribution: Amazon River basin, Brazil, and Guyana (Reis, 2003).
Remarks: Nijssen & Isbrücker (1980: 210) indicated that the stated type locality of Corydoras griseus was incorrect
and the correct locality was: Guyana, Essequibo, Portaro River, Kuribong trail.
Corydoras guapore Knaack, 1961
Corydoras guapore Knaack, 1961: 135, fig. 32. Type locality: Brasilien, im Hauptstrom des oberen Rio Guaporé.
Holotype: ZMB 21406.
Distribution: Guaporé River basin, Brazil (Reis, 2003).
Corydoras guianensis Nijssen, 1970
Corydoras guianensis Nijssen, 1970: 21, fig. 9. Type locality: Creek at right of Nickerie River, 12 kilometers W.S.W
of Stondansie Fall [Nickerie, Suriname]. Holotype: ZMA 105933.
Distribution: Coastal rivers of Suriname and French Guiana (Reis, 2003).
Corydoras habrosus Weitzman, 1960
Corydoras habrosus Weitzman, 1960a: 141, fig. 1. Type locality: Río Salinas, a branch of Río Pao Viejo, El Baúl,
State of Cojedes, Venezuela [8º59'N, 68º16'W]. Holotype: SU 52351.
Distribution: Upper Orinoco River basin, Colombia and Venezuela (Reis, 2003).
Corydoras haraldschultzi Knaack, 1962
Corydoras haraldschultzi Knaack, 1962: 129, fig. 1. Type locality: Brasilien, Rio Guaporé. Holotype: SMF 5475.
Distribution: Brazil and Bolivia (Reis, 2003).
Corydoras hastatus Eigenmann & Eigenmann, 1888
Corydoras hastatus Eigenmann & Eigenmann, 1888b: 166. Type locality: Villa Bella [= Parintins, 2º38'S, 56º45'W,
Amazonas, Brazil]. Lectotype: MCZ 7747, designated by Nijssen & Isbrücker (1980b: 213).
Corydoras australe Eigenmann & Ward, in Eigenmann, McAtee & Ward, 1907: 123. Type locality: Corumba[19º00'S, 57º25'W, Rio Paraguay system, Mato Grosso, Brazil]. Holotype: CAS 77305.
Distribution: Amazon and Paraguay River basins (Reis, 2003).
Corydoras heteromorphus Nijssen, 1970
Corydoras heteromorphus Nijssen, 1970: 22, fig. 10. Type locality: Creek at right bank of Coppename River(3º52'30"N, 56º53'W), Saramacca, Suriname. Holotype: ZMA 105880.
Distribution: Coppename and Nickerie River basins, Suriname (Reis, 2003).
Corydoras imitator Nijssen & Isbrücker, 1983
Corydoras imitator Nijssen & Isbrücker, 1983c: 79, fig. 8. Type locality: haut Rio Negro, localite exact inconnue[Brazil]. Holotype: MZUSP 27894.
Distribution: Upper Negro River basin, Brazil (Reis, 2003)
Corydoras incolicana Burgess 1993
Corydoras incolicana Burgess, 1993: 157, unnumbered figure. Type locality: Rio Içana, upper Rio Negro, Brazil.
Holotype: MZUSP 45717.

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FERRARISDistribution: Upper Negro River basin, Brazil (Reis, 2003).
Corydoras isbrueckeri Knaack, 2004
Corydoras isbrueckeri Knaack, 2004: 88, figs. 30–36. Type locality: Bolivien, Departamento Santa Cruz, Provincia
J.M. Velasco, System río Itenez, Nebengewässer des río Paragua (ein namenloser arroyo), 13º34.384'S,
62º03.067'W. Holotype: MTD F 28474.
Distribution: Itenez River basin, Bolivia (Knaack, 2004).
Corydoras julii Steindachner, 1906
Corydoras julii Steindachner, 1906: 480. Type locality: im Parahim, dem Ausflusse der Sess von Parnagua und an
der Ausmündung des Baches bei Victoria. Lectotype: NMW 61105, designated by Nijssen & Isbrücker (1980b:
195).
Distribution: Lower Amazon River and coastal rivers in northeastern Brazil (Reis, 2003).
Corydoras kanei Grant, 1997
Corydoras kanei Grant, 1997: 41, fig. 1. Type locality: Aquarium specimens said to be from Brazil, Roraima Statenear Boa Vista, possibly from the Rio Branco. Holotype: MZUSP 52489.
Distribution: Negro River basin, Brazil (Reis, 2003).
Corydoras lacerdai Hieronimus, 1995
Corydoras lacerdai Hieronimus, 1995: 110, fig. 5. Type locality: Brasilien, Rio Ribeira da Terra Firme, zwischen
den Städten Canavieiras und Ilheus, im Südesten des Bundesstaates Brasilien. Holotype: MZUSP 47682.
Distribution: Ribeira da Terra Firme River, Bahia State, Brazil (Reis, 2003).
Corydoras lamberti Nijssen & Isbrücker, 1986
Corydoras lamberti Nijssen & Isbrücker, 1986a: 71, fig. 16. Type locality: Loreto, cours inférieur du Rio Huytoyacu, près du village Nuevo Progresso, affluent de la rive droite du Rio Pastaza, 40 kilometers environ au N. du
Lago Rimachi (4º13'S, 76º38'W), Peru. Holotype: ZMA 119335.
Distribution: Pastaza River basin, Peru (Reis, 2003).
Corydoras latus Pearson, 1924
Corydoras latus Pearson, 1924: 19, pl. 3 (fig. 1). Type locality: Lagoons, Lake Rogoagua [Bolivia]. Holotype: CAS
36452.
Distribution: Beni River basin, Bolivia (Reis, 2003).
Corydoras leopardus Myers, 1933
Corydoras leopardus Myers, 1933: 188. Type locality: Brazil [considered by Reis (2003) as likely to have comefrom a Brazilian coastal river south of the Amazon]. Lectotype: USNM 93305, designated by Nijssen &Isbrücker (1980b: 197).
Corydoras funnelli Fraser-Brunner, 1947: 241, fig. 1. Type locality: Amazon ? (imported aquarium specimen).
Holotype: USNM 102222.
Distribution: Amazon River basin (Reis, 2003).
Corydoras leucomelas Eigenmann & Allen, 1942
Corydoras leucomelas Eigenmann & Allen, 1942: 178, pl. 12 (fig. 4). Type locality: Yarinacocha [Peru]. Holotype:
CAS 36561.
Corydoras caquetae Fowler, 1943a: 248, figs. 26–27. Type locality: Florencia, Rio Orteguasa, Colombia. Holotype:
ANSP 70509.
Distribution: Upper Amazon River basin (Reis, 2003).
Corydoras loretoensis Nijssen & Isbrücker, 1986
Corydoras loretoensis Nijssen & Isbrücker, 1986a: 68, fig. 4. Type locality: Loreto, Prov. Maynas, Río Nanay, plage
sableuge, rive droite, environ 15 minutes en amont de Santa Clara, Peru. Holotype: NRM 26079.
Distribution: Upper Amazon River basin, Peru (Reis, 2003).
Corydoras loxozonus Nijssen & Isbrücker, 1983
Corydoras loxozonus Nijssen & Isbrücker, 1983b: 67, fig. 12. Type locality: Meta, Lomalinda near Río Ariari, tributary to Río Guaviare, S.E. of Villavicencio [Colombia]. Holotype: ANSP 150170.
Distribution: Meta River basin, Colombia (Reis, 2003).
Corydoras maculifer Nijssen & Isbrücker, 1971

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Corydoras maculifer Nijssen & Isbrücker, 1971: 183, fig. 1. Type locality: Brazil, Est. Mato Grosso, Sangadina
stream, 2 kilometers W of Chavantina (14º45'S, 52º20'W), tributary of the Rio das Mortes. Holotype: BMNH
1970.10.30.3.
Distribution: Upper Araguaia River basin, Brazil (Reis, 2003).
Corydoras mamore Knaack, 2002
Corydoras mamoré Knaack, 2002d: 16, unnumbered figure. Type locality: [Bolivia]. Holotype: MTD F 26741.
Distribution: Bolivia (Knaack, 2002d).
Corydoras melanistius Regan, 1912
Corydoras melanistius Regan, 1912a: 216. Type locality: Essequibo [Guyana]. Lectotype: BMNH 1864.1.21.86,
designated by, and illustrated in, Nijssen & Isbrücker (1967: 32, pl. 2, fig. 1).
Corydoras wotroi Nijssen & Isbrücker, 1967: 44, pl. 5 (fig. 3). Type locality: Outlet of Kleine Saramacca along andbetween sand bank [Suriname]. Holotype: RMNH 25331.
Distribution: Coastal rivers of the Guianas (Reis, 2003).
Corydoras melanotaenia Regan, 1912
Corydoras melanotaenia Regan, 1912a: 217. Type locality: Honda [Río Meta system, Colombia]. Lectotype:
BMNH 1909.7.23.41, designated by Nijssen & Isbrücker (1980b: 207).
Distribution: Meta River basin, Colombia (Reis, 2003).
Corydoras melini Lönnberg & Rendahl, 1930
Corydoras melini Lönnberg & Rendahl, 1930: 1, fig. 1. Type locality: Jauareté an dem Ausfluss des Rio Papuri in
den Rio Uaupés [Brazil]. Lectotype: NRM 11091, designated by Nijssen & Isbrücker (1980b: 209).
Distribution: Upper Negro and Meta River basins (Reis, 2003).
Corydoras metae Eigenmann, 1914
Corydoras metae Eigenmann, 1914b: 230. Type locality: Barrigona, Rio Meta, Colombia. Holotype: CAS 36447;
holotype illustrated in Eigenmann (1916: pl. 14; 1922b: 227, pl. 8, fig. 5).
Distribution: Meta River basin, Colombia (Reis, 2003)
Corydoras micracanthus Regan, 1912
Corydoras micracanthus Regan, 1912a: 211. Type locality: Salta, Argentina. Lectotype: BMNH 1897.1.27.8, designated by Nijssen & Isbrücker (1980b: 206).
Distribution: Western tributaries of Paraná River in Salta Province, Argentina (Reis, 2003).
Corydoras multimaculatus Steindachner, 1907
Corydoras multimaculatus Steindachner, 1907d: 291. Type locality: Nebenarme des Rio Preto bei Sa. Rita im Staate
Bahia [Brazil]. Lectotype: NMW 46783, designated by Nijssen & Isbrücker (1980b: 196).
Distribution: Preto River basin in Bahia State, Brazil (Reis, 2003).
Corydoras multiradiatus (Orcés, 1960)
Chaenothorax multiradiatus Orcés, 1960: 3, fig. 1. Type locality: afluente occidental del río Lagartococha, cerca delpoblado de Garza-Cocha, sistema del alto Napo. Holotype: USNM 200739.
Distribution: Western Amazon River basin, Ecuador and Peru (Reis, 2003), as Brochis multiradiatus.
Corydoras nanus Nijssen & Isbrücker, 1967
Corydoras nanus Nijssen & Isbrücker, 1967: 41, pl. 5 (fig. 1). Type locality: Little tributaries of Gran-Rio between
Ligolio and Awaradam Falls [Brokopondo, Suriname]. Holotype: RMNH 25333.
Distribution: Suriname and Maroni River basins in Suriname and Iracoubo River basin in French Guiana (Reis,
2003).
Corydoras napoensis Nijssen & Isbrücker, 1986
Corydoras napoensis Nijssen & Isbrücker, 1986a: 73, fig. 25. Type locality: Napo, Lagartococha, affluent septentrional du Rio Aguarico, entre l’embouchure de la rivière (0º39'S, 75º16'W) et le village de Garzacocha (0º28'S,
75º21'W), bassin du Rio Napo [Ecuador]. Holotype: ZMA 119226.
Distribution: Western Amazon River basin, eastern Ecuador and Peru (Reis, 2003).
Corydoras narcissus Nijssen & Isbrücker, 1980
Corydoras narcissus Nijssen & Isbrücker, 1980d: 497, fig. 2. Type locality: Brazil, Est. Amazonas, Rio Purus system, creek into Rio Ipixuna, 7º31'S, 63º16'W, 30 km west of Humaitá. Holotype: ZMA 115178.

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FERRARISDistribution: Purus River basin, middle Amazon River system, Brazil, (Reis, 2003).
Corydoras nattereri Steindachner, 1877
Corydoras Nattereri Steindachner, 1877a: 143, pl. 11 (fig. 1). Type locality: Affluent of Rio Parahyba, 3 mi. up, Riode Janeiro, Brazil, by neotype designation. Neotype: MCZ 8229, designated by Nijssen & Isbrücker (1980a: 3).
Corydoras juquiaae Ihering, 1907: 36. Type locality: Juquiá River, Poço Grande, State of São Paulo [Brazil]. Lectotype: CAS 16016, designated by Ellis (1913: 403), subsequent designation by Britski (1969: 207) of MZUSP
1984 for Corydoras juquiae is invalid.
Corydoras nattereri triseriatus Ihering, 1911: 386. Type locality: Rio Doce, Est. Espírito Santo [Brazil]. Lectotype:
MZUSP 342, designated by Britski (1969: 207).
Distribution: Coastal rivers in southeastern Brazil from Espirito Santo to Paraná, Brazil (Reis, 2003).
Corydoras negro Knaack, 2004
Corydoras negro Knaack, 2004: 81, figs. 22–29. Type locality: Bolivien, Departamento Santa Cruz, Provincia Nuflode Chavez, Raum Urubicha, System río Blanco, río Negro (Hauptfluss), Restgewässer, 15º31.502'S,
62º56.208'W. Holotype: MTD F 28472.
Distribution: Blanco River basin, Bolivia (Knaack, 2004).
Corydoras nijsseni Sands, 1989
Corydoras elegans nijsseni Sands, 1989: 45. Type locality: Tributary of the Rio Negro, Brazil. Holotype: reportedly
deposited at RMNH; but reported as four syntypes in the on-line Catalog of Fishes, as RMNH 31625 and 31626.
Distribution: Negro River basin, Brazil (Reis, 2003).
Corydoras noelkempffi Knaack, 2004
Corydoras noelkempffi Knaack, 2004: 94, figs. 37–41. Type locality: Bolivien, Departamento Santa Cruz, Provincia
J.M. Velasco, System río Itenez, río San Martin, 15º23.265'S, 61º29.758'W. Holotype: MTD F 28576.
Distribution: Itenez River basin, Bolivia (Knaack, 2004).
Corydoras oiapoquensis Nijssen, 1972
Corydoras oiapoquensis Nijssen, 1972: 425, fig. 7. Type locality: French Guiana, Cumuri Creek at left bank of Rio
Oiapoque (= Oyapock River), upstream of first rapids of Grande Roche, southwest of village Oiapoque (Brazil,
03º54'N, 51º46'W). Holotype: IRSNB 489.
Distribution: Oyapock River basin, Brazil and French Guiana (Reis, 2003).
Corydoras ornatus Nijssen & Isbrücker, 1976
Corydoras ornatus Nijssen & Isbrücker, 1976b: 126, fig. 1. Type locality: Brazil, Est. Pará, Rio Tapajós, 80 km E of
Jacareacanga, 06º09'S, 58º15'W. Holotype: ZMA 114690.
Distribution: Lower Tapajós River basin, Brazil (Reis, 2003).
Corydoras orphnopterus Weitzman & Nijssen, 1970
Corydoras orphnopterus Weitzman & Nijssen, 1970: 125, fig. 3. Type locality: Ecuador, Lower Río Bobonaza
between Montalvo (2º06'S, 76º59'W) and Chicherota, Rio Pastaza system, Est. Pastaza. Holotype: USNM
204361.
Distribution: Pastaza River basin, Ecuador (Reis, 2003).
Corydoras osteocarus Böhlke, 1951
Corydoras osteocarus Böhlke, 1951a: 824. Type locality: San Fernando de Atabapo, where Rio Atabapo empties
into the Orinoco, Venezuela (at approximately 04º03' North Latitude, 67º45' West Longitude). Holotype: SU
16333.
Distribution: Orinoco River basin and coastal rivers of Suriname (Reis, 2003).
Corydoras ourastigma Nijssen, 1972
Corydoras ourastigma Nijssen, 1972: 428, fig. 8. Type locality: Brazil, Est. Acre, Rio Iquiri (= Rio Ituxi) at road Rio
Branco village —Abuna, 47 kilometers from Rio Branco village (about 10º00'S, 67º30'W), Purus River system.
Holotype: IRSNB 498.
Distribution: Purus River basin, middle Amazon system, Brazil (Reis, 2003).
Corydoras oxyrhynchus Nijssen & Isbrücker, 1967
Corydoras oxyrhynchus Nijssen & Isbrücker, 1967: 42, pl. 4 (fig. 3). Type locality: Gojo Creek above Posoegroenoe,
tributary of Saramacca River. Holotype: RMNH 25329.

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Distribution: Saramacca River basin, Suriname (Reis, 2003).
Corydoras paleatus (Jenyns, 1842)
Callichthys paleatus Jenyns, 1842 (in Jenyns, 1840–42): 113. Type locality: South America. Lectotype: BMNH
1917.7.14.18, designated by Nijssen & Isbrücker (1980b: 204).
Corydoras maculatus Steindachner, 1879a: 32. Type locality: La Plata [Argentina]. No types known.
Corydoras marmoratus Steindachner, 1879e: 26, pl. 5 (fig. 1). Type locality: Montevideo [Uruguay] und ... La Plata
innerhalb der Provinz Buenos-Ayres [Argentina]. Lectotype: NMW 5538, designated by Nijssen & Isbrücker
(1980b: 204).
Corydoras microcephalus Regan, 1912a: 211. Type locality: La Plata. Lectotype: BMNH 1890.3.12.4, designatedby Nijssen & Isbrücker (1980b: 204).
Silurus quadricostatus Larrañaga, 1923: 376. Type locality: Uruguay. No types known.
Silurus 7-radiatus Larrañaga, 1923: 385. Type locality: Uruguay. No types known.
Distribution: Lower Paraná River basin and coastal rivers in Uruguay and Brazil (Reis, 2003).
Remarks: The name Corydoras punctatus argentina first appeared in Steindachner (1879a: 26) as a name only andin synonymy of Corydoras marmoratus; as such it is unavailable from that publication and the name has apparently not been made available subsequently. Isbrücker (2001b) treated Corydoras maculatus, C. marmoratus
and C. microcephalus as doubtfully valid species, but with no additional information.
Corydoras panda Nijssen & Isbrücker, 1971
Corydoras panda Nijssen & Isbrücker, 1971: 186, fig. 2. Type locality: Peru, Est. Huanuco, Aquas Amarillas, tributary of Río Pachitea, Ucayali river system. Holotype: BMNH 1969.7.15.8.
Distribution: Upper Amazon River basin, Peru (Reis, 2003).
Corydoras pantanalensis Knaack, 2001
Corydoras pantanalensis Knaack, 2001: 59, unnumbered figure. Type locality: Bolivien, Prov. Santa Cruz, Restgewässer im nördlichen Pantanal, Überschwemmungsgebiet im Raum San Lorenzo, Flusssystem Rio Cussis
und Rio Las Petas. Koordinaten des Fundortes: 16º06' 525"S, 59º55'620"W [sic]. Holotype: MTD F24774.
Distribution: Cussis and Las Petas River basins, Bolivia (Reis, 2003).
Corydoras paragua Knaack, 2004
Corydoras paragua Knaack, 2004: 65, figs. 6–12. Type locality: Bolivien, Departamento Santa Cruz, Provincia J.M.
Velasco, río Itenez system, río Paragua,, Río Paragua, nordwestliches Ufer im Mündungsgebiet. ... 13º31.842'S,
61º90.015'W. Holotype: MTD F 28468.
Distribution: Itenez River basin, Bolivia (Knaack, 2004).
Corydoras parallelus Burgess 1993
Corydoras parallelus Burgess, 1993: 154, unnumbered figure. Type locality: Rio Içana, upper Rio Negro, Brazil.
Holotype: MZUSP 45716.
Distribution: Upper Negro River basin, Brazil (Reis, 2003).
Corydoras pastazensis Weitzman, 1963
Corydoras pastazensis Weitzman, 1963: 59, fig. 1. Type locality: Chicherata, near the mouth of the Río Bobonaza, atributary of the Río Pastaza, itself a tributary of the Río Maranon, Pastaza Province, Ecuador, about 02º33'S latitude and 76º40'W longitude. Holotype: USNM 177216.
Corydoras pastazensis orcesi Weitzman & Nijssen, 1970: 120, fig. 1. Type locality: Ecuador, Río Conambo near village of Conambo (1º55'S, 76º51'W), Río Tigre system, Est. Pastaza. Holotype: USNM 204358.
Distribution: Pastaza River basin, Ecuador (Reis, 2003).
Corydoras paucerna Knaack, 2004
Corydoras paucerna Knaack, 2004: 57, figs. 1–5. Type locality: Bolivien, Departamento Santa Cruz, Provincia J.M.
Velasco, río Itenez system, río Paragua, Nebengewässer (Oberlauf) eines namenlossen arroyos (Restgewässer):
13º34.384'S, 62º03.067'W. Holotype: MTD F 28466.
Distribution: Itenez River basin, Bolivia (Knaack, 2004).
Corydoras pinheiroi Dinkelmeyer, 1995
Corydoras pinheiroi Dinkelmeyer, 1995: 60, unnumbered figure. Type locality: Brasilien, Est. Rondonia, in einem
Nebenfluss des Rio Ribeiro bei der Ortschaft Guajará-Mirim. Holotype: MZUSP 48099.

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FERRARISDistribution: Western Amazon River basin in Rondônia State, Brazil (Reis, 2003).
Corydoras polystictus Regan, 1912
Corydoras polystictus Regan, 1912a: 216. Type locality: Descalvados, Matto Grosso [Brazil]. Lectotype: BMNH
1895.5.17.62, designated by Nijssen & Isbrücker (1980b: 197).
Corydoras virescens Miranda Ribeiro, 1912: 16. Type locality: Rio Paraguay, perto de Cáceres [Brazil]. Holotype:
MNRJ 3806.
Distribution: Paraguay River basin, Brazil (Reis, 2003).
Remarks: Publication of the name Corydoras polystictus apparently predates that of C. virescens; see Eschmeyer(1998) for reported publication dates of relevant descriptions.
Corydoras potaroensis Myers, 1927
Corydoras potaroensis Myers, 1927: 126. Type locality: Br. Guiana: Creek below Potaro Landing. Lectotype: CAS
24754, designated by Nijssen & Isbrücker (1980b: 208).
Distribution: Potaro River basin, Guyana (Reis, 2003).
Corydoras pulcher Isbrücker & Nijssen, 1973
Corydoras pulcher Isbrücker & Nijssen, 1973: 2, fig. 1. Type locality: Brazil, Est. Amazonas, Rio Purus, north of
Lábrea (7º20'S, 64º46'W), Rio Amazonas system. Holotype: SMF 9231.
Distribution: Purus River basin, middle Amazon system, Brazil (Reis, 2003).
Corydoras punctatus (Bloch, 1794)
Cataphractus punctatus Bloch, 1794: 90, pl. 377 (fig. 2). Type locality: Flüssen Surinams. Lectotype: ZMB 3149,
designated by Nijssen & Isbrücker (1975: 63). Earlier neotype designation by Nijssen & Isbrücker (1967: 28)(RMNH 25301) invalidated by rediscovery of syntype series. Placed on Official list (ICZN Opinion 1169).
Distribution: Suriname River basin in Suriname and Iracoubo River basin in French Guiana (Reis, 2003).
Remarks: Isbrücker (1999, 2000) discussed the taxonomic history of this species.
Corydoras pygmaeus Knaack, 1966
Corydoras pygmaeus Knaack, 1966a: 168, first unnumbered figure. Type locality: einem Nebenfluß des Rio
Madeira (Brasilien). Holotype: ZMB 21620. Also described as new in Knaack (1966b: 364).
Distribution: Madeira River basin (Reis, 2003).
Corydoras rabauti La Monte, 1941
Corydoras rabauti La Monte, 1941: 5. Type locality: A tributary of the Amazon River [Brazil]. Holotype: AMNH
15644.
Corydoras myersi Miranda Ribeiro, 1942: 427, fig. 1. Type locality: Rio Javari, Município de Banjamin Constant,
Estado do Amazonas, Brasil. Holotype: MNRJ 3626.
Distribution: Upper Amazon, Solimões, and Negro River basins (Reis, 2003).
Corydoras reticulatus Fraser-Brunner, 1938
Corydoras reticulatus Fraser-Brunner, 1938: 208, fig. 2 (unnumbered). Type locality: Monte Alegre River, Amazon[Brazil] (2º00'S, 54º04'W), by neotype designation. Neotype: BMNH 1926.10.27.333.
Distribution: Lower Amazon River basin, Brazil (Reis, 2003).
Remarks: Original two syntypes are considered to be lost and Fraser-Brunner (1947: 245) redescribed species with a“holotype” which Weitzman (1960b) designated as neotype (see Nijssen & Isbrücker, 1980: 197; Reis, 2003).
† Corydoras revelatus Cockerell, 1925
† Corydoras revelatus Cockerell, 1925: 398. Type locality: Sunchal, Province of Jujuy, Argentina; Tertiary. Holotype: At University of Colorado.
Distribution: Maiz Gordo Formation, Sunchal, Jujuy Province, and Salta, Argentina; late Paleocene (Reis, 1998b:
357).
Remarks: Redescribed in Bardack (1961).
Corydoras reynoldsi Myers & Weitzman, 1960
Corydoras reynoldsi Myers & Weitzman, 1960: 105, fig. 2. Type locality: Tributary to the Río Orteguaza, opposite
the town and air base known as Tres Esquinas, Caquetà Province, Colombia. Holotype: SU 52349.
Distribution: Upper Caquetá River basin, Colombia (Reis, 2003).
Remarks: The caption for figure 2 lists SU 50702 as the holotype of Corydoras reynoldsi. Apparently the holotype

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was removed from this lot and assigned a new number, and the inclusion of the original catalog number in the
figure caption is a lapsus calami.
Corydoras robineae Burgess, 1983
Corydoras robineae Burgess, 1983: 42, unnumbered figure. Type locality: Rio Aiuana, a southern tributary of the
middle Rio Negro, Amazonas, Brazil. Holotype: MZUSP 27175.
Distribution: Upper Negro River basin, Brazil (Reis, 2003).
Corydoras robustus Nijssen & Isbrücker, 1980
Corydoras robustus Nijssen & Isbrücker, 1980d: 499, fig. 3. Type locality: Brazil, Est. Amazonas, Rio Purus system,
creek into Rio Ipixuna, 07º31'S, 63º16'W, 30 km west of Humaitá. Holotype: ZMA 115179.
Distribution: Purus River basin, middle Amazon system, Brazil (Reis, 2003).
Corydoras sanchesi Nijssen & Isbrücker, 1967
Corydoras sanchesi Nijssen & Isbrücker, 1967: 43, pl. 5 (fig. 2). Type locality: Gojo Creek above Posoegroenoe,
tributary of Saramacca River [Suriname]. Holotype: RMNH 25319.
Distribution: Saramacca River basin, Suriname (Reis, 2003).
Corydoras saramaccensis Nijssen, 1970
Corydoras saramaccensis Nijssen, 1970: 38, fig. 21. Type locality: Creek at right bank of Kleine Saramacca River,
11 km E.S.E. from junction with Saramacca River, Brokopondo, Suriname. Holotype: ZMA 106018.
Distribution: Saramacca River basin, Suriname (Reis, 2003).
Corydoras sarareensis Dinkelmeyer, 1995
Corydoras saraareensis Dinkelmeyer, 1995: 61, unnumbered figure. Type locality: Brasilien, Est. Mato Grosso, Rio
Sararé, rechter Nebenfluß des Rio Guaporé. Holotype: MZUSP 48100.
Distribution: Guaporé River basin, Brazil (Reis, 2003).
Corydoras schwartzi Rössel, 1963
Corydoras schwartzi Rössel, 1963: 359, fig. 1. Type locality: Brasilien, Mündungsgebiet des Rio Purus. Holotype:
SMF 6425.
Distribution: Purus River basin, middle Amazon system, Brazil (Reis, 2003).
Corydoras semiaquilus Weitzman, 1964
Corydoras semiaquilus Weitzman, 1964: 121, fig. 5. Type locality: Igarapé Preto, a small jungle rivulet at the headwaters of the black water creeks, which empty in the upper Solimões, State of Amazonas, Brazil. Holotype: SU
55939.
Distribution: Western Amazon River basin (Reis, 2003).
Corydoras septentrionalis Gosline, 1940
Corydoras septentrionalis Gosline, 1940b: 16. Type locality: Río Piña, north of Maturín, Venezuela. Holotype: SU
35055.
Corydoras cortesi Castro, 1987: 31, fig. 1. Type locality: Río Arauca near to Arauca, Intendencia de Arauca (7º03'N,
70º46'W) [Colombia]. Holotype: UBJTL MM671.
Distribution: Orinoco River basin (Reis, 2003).
Corydoras serratus Sands, 1995
Corydoras serratus Sands, 1995: 14, fig. 7. Type locality: Rio Poranga, 8 kilometers north of the Nobua oba, where
the two streams join a small tributary of the upper Negro river, Brazil. Holotype: LIVCM 1994.4.38.
Distribution: Upper Negro River basin, Brazil (Reis, 2003).
Corydoras seussi Dinkelmeyer, 1996
Corydoras seussi Dinkelmeyer, 1996: 25. Type locality: Brasilien, Est. Rondonia, kleiner Nebenfluß des Rio Pacaás
Novos, eines rechtsseitigen Zuflusses des Rio Mamoré nahe bei Guajara-Mirim. Holotype: MZUSP 49322.
Distribution: Mamoré River basin, Brazil (Reis, 2003).
Corydoras similis Hieronimus, 1991
Corydoras similis Hieronimus, 1991: 39, fig. 1. Type locality: Brasilien, in der Nähe der Stadt Ariqumes in einem
Zufluß zum Rio Madeira, Bundesstaat Rondonia. Holotype: ZFMK 16400.
Distribution: Madeira River basin, Brazil (Reis, 2003).
Corydoras simulatus Weitzman & Nijssen, 1970

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Corydoras simulatus Weitzman & Nijssen, 1970: 126, fig. 4. Type locality: Colombia, Río Ocoá near Puerto López(04º06'N, 72º57'W), Río Meta system, Est. Meta. Holotype: USNM 197615.
Distribution: Upper Meta River, Colombia (Reis, 2003).
Corydoras sipaliwini Hoedeman, 1965
Corydoras punctatus sipaliwini Hoedeman, 1965: 87, fig. 46. Type locality: Sipalawini River [Suriname]. Lectotype: ZMA 104.278, designated by Isbrücker (2000).
Distribution: Coastal river basins of Guyana and Suriname (Reis, 2003).
Corydoras sodalis Nijssen & Isbrücker, 1986
Corydoras sodalis Nijssen & Isbrücker, 1986a: 68, fig. 6. Type locality: Loreto, Rio Yavari, Cano de Guavariba dansle Lago Matamata (04º12'S, 70º17'W), Peru. Holotype: NRM 13713.
Distribution: Amazon River of Brazil and Peru (Reis, 2003).
Corydoras solox Nijssen & Isbrücker, 1983
Corydoras solox Nijssen & Isbrücker, 1983c: 80, fig. 9. Type locality: bassin du Rio Oiapoque, 60 km S du village
Martinique (03º51'N, 51º53'O), Terr. Amapa, Brazil. Holotype: MNHN 1983-0531.
Distribution: Oyapock River basin and smaller coastal drainages in French Guiana (Reis, 2003).
Corydoras spectabilis Knaack, 1999
Corydoras spectabilis Knaack, 1999c: 74, unnumbered figure. Type locality: Brasilien, Mato Grosso, Oberlauf Rio
Guaporé im Raum Vila Bela. Holotype: ZMB 33150.
Distribution: Guaporé River basin, Brazil (Reis, 2003).
Remarks: Apparently also published as new in Knaack (1999e).
Corydoras spilurus Norman, 1926
Corydoras spilurus Norman, 1926b: 95. Type locality: Iponcin Creek, into Approuague River, French Guiana. Lectotype: BMNH 1926.3.2.738, designated by, and illustrated in, Nijssen & Isbrücker (1967: 33, pl. 2, fig. 2).
Distribution: Approuague River basin, French Guiana, and Suriname River basin, Suriname (Reis, 2003).
Corydoras splendens (Castelnau, 1855)
Callichthys splendens Castelnau, 1855: 39, pl. 18 (fig. 3). Type locality: Rio Tocantins [Brazil]. Holotype: MNHN
0000-4291.
Callichthys taiosh Castelnau, 1855: 39, pl. 19 (fig. 1). Type locality: Chiquitos [Bolivia]. Holotype: possibly MNHN
0000-4290. Castelnau indicated that he saw only a drawing of this species, but the specimen listed here wastaken by a Mr. Weddell, from the correct locality and appears to be the specimen upon which the illustration was
prepared.
Brochis coeruleus Cope, 1872a: 277, pl. 9 (fig. 3). Type locality: Tributaries of the Ambyiacu. Lectotype: ANSP
8231, designated by Nijssen & Isbrücker (1970: 157–158).
Brochis dipterus Cope, 1872a: 278. Type locality: Ambyiacu River. Holotype: ANSP 117172.
Corydoras semiscutatus Cope, 1872a: 280, pl. 6 (fig. 1). Type locality: Ambyiacu River. Holotype: ANSP 8289.
Chaenothorax bicarinatus Cope, 1878: 679. Type locality: Peruvian Amazon. Holotype: ANSP 21447.
Chaenothorax eigenmanni Ellis, 1913: 393, pl. 26 (fig. 2). Type locality: Cáceres [Mato Grosso, Brazil]. Holotype:
FMNH 54880.
Distribution: Amazon River basin, Brazil, Colombia, Ecuador, and Peru, as Brochis splendens (Reis, 2003).
Remarks: Callichthys splendens was apparently first given precedence over C. taiosh in Nijssen and Isbrücker
(1970: 157).
Corydoras steindachneri Isbrücker & Nijssen, 1973
Corydoras steindachneri Isbrücker & Nijssen, 1973: 4, fig. 3. Type locality: Brazil, Est. Paraná, Paranaguá (25º32'S,
48º36'W). Holotype: NMW 1504.
Distribution: Coastal rivers in Paraná State, Brazil (Reis, 2003).
Corydoras stenocephalus Eigenmann & Allen, 1942
Corydoras stenocephalus Eigenmann & Allen, 1942: 177, pl. 12 (fig. 2). Type locality: Yarinococha [Peru]. Holotype: CAS 36386.
Distribution: Eastern Ecuador and Ucayali River basin, Peru (Reis, 2003).
Corydoras sterbai Knaack, 1962

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Corydoras sterbai Knaack, 1962: 131, pl. 12 (fig. 2). Type locality: Brasilien. Holotype: SMF 5476.
Distribution: Central Brazil and Bolivia (Reis, 2003).
Corydoras surinamensis Nijssen, 1970
Corydoras schwartzi surinamensis Nijssen, 1970: 39, fig. 22. Type locality: Creek at right bank of Coppename
River (03º52'30"N, 56º53'00"W), Saramacca, Suriname. Holotype: ZMA 105876.
Distribution: Coppename River basin, Suriname (Reis, 2003).
Corydoras sychri Weitzman, 1960
Corydoras sychri Weitzman, 1960a: 151, fig. 7. Type locality: Tropical South America. Holotype: SU 51295.
Distribution: Nanay River basin, Peru (Reis, 2003).
Corydoras treitlii Steindachner, 1906
Corydoras treitlii Steindachner, 1906: 478. Type locality: Mündungstelle eines kleinen Baches gefangen, der bei
dem Städtchen Victoria in der Parnahyba fällt [Brazil]. Lectotype: NMW 61103, designated by Weitzman(1964: 116).
Distribution: Coastal drainages in northeastern Brazil (Reis, 2003).
Corydoras trilineatus Cope, 1872
Corydoras trilineatus Cope, 1872a: 281, pl. 6 (fig. 2). Type locality: Ambyiacu River [Peru]. Lectotype: ANSP
8294, designated by Nijssen & Isbrücker (1980b: 194).
Corydoras episcopi Eigenmann & Allen, 1942: 177, pl. 12 (fig. 3). Type locality: Rio Morona. Holotype: CAS
36448.
Corydoras dubius Nijssen & Isbrücker, 1967: 38, pl. 4 (fig. 2). Type locality: Suriname, surroundings of Paramaribo.
Holotype: ZMA 104623.
Distribution: Middle and upper Amazon River basin and coastal drainages of Suriname (Reis, 2003).
Corydoras tukano Britto & Lima, 2003
Corydoras tukano Britto & Lima, 2003: 84, figs. 1, 2. Type locality: Brazil, estado do Amazonas, rio Tiquie, comunidade de Caruru, beaches in pool below the fall, 0º16'28.9"N, 69º54'53.6"W (UTM 19N0398191/0030366).
Holotype: MZUSP 82100.
Distribution: Tiquié River basin, Negro River drainage, Brazil (Britto & Lima, 2003).
Corydoras undulatus Regan, 1912
Corydoras undulatus Regan, 1912a: 217. Type locality: La Plata. Holotype: BMNH 1912.7.10.5.
Distribution: Lower Paraná River basin and coastal rivers in southern Brazil (Reis, 2003).
Corydoras virginiae Burgess, 1993
Corydoras virginiae Burgess, 1993: 152, unnumbered figure. Type locality: Rio Ucayali, Peru. Holotype: MZUSP
45715.
Distribution: Ucayali River basin, Peru (Reis, 2003).
Corydoras vittatus Nijssen, 1971
Corydoras blochi vittatus Nijssen, 1971: 97, fig. 7. Type locality: Brazil, Est. Maranhão, tributary of Rio Itapicurú at
Caxias (4º53'S, 43º20'W). Holotype: ZMA 109990.
Distribution: Itapicurú River basin in northeastern Brazil (Reis, 2003).
Corydoras weitzmani Nijssen, 1971
Corydoras weitzmani Nijssen, 1971: 91, fig. 1. Type locality: Peru, Est. Cuzco, at Cuzco (13º32'S, 71º57'W), Vilcanota river system. Holotype: FMNH 69934.
Distribution: Western Amazon River basin, Peru (Reis, 2003).
Corydoras xinguensis Nijssen, 1972
Corydoras xinguensis Nijssen, 1972: 429, fig. 9. Type locality: Brazil, Est. Mato Grosso, Suia Missú Creek, tributary of upper Río Xingu near Posto Diauarum (about 11º15'S, 53º00'W). Holotype: IRSNB 500.
Distribution: Upper Xingu River basin, Brazil (Reis, 2003).
Corydoras zygatus Eigenmann & Allen, 1942
Corydoras zygatus Eigenmann & Allen, 1942: 175, pl. 12 (fig. 1). Type locality: Creek, Yurimaguas [Peru]. Holotype: CAS 24756.
Distribution: Western Amazon River basin (Reis, 2003).

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Dianema Cope, 1871b: 112. Type species: Dianema longibarbis Cope, 1872. Type by subsequent monotypy by
Cope (1872). Gender: Neuter. Appeared first with brief description and no species; a single species assigned tothis genus by Cope (1872a: 276).
Decapogon Eigenmann & Eigenmann, 1888b: 165. Type species: Callichthys adspersus Steindachner, 1877. Typeby monotypy. Gender: Masculine.
Phylogeny: Reis (1997).
Dianema longibarbis Cope, 1872
Dianema longibarbis Cope, 1872a: 276, pl. 7 (fig. 1). Type locality: Ambyiacu River, Peru. Syntypes: ANSP 8285
(1), ANSP 21540 (1).
Callichthys adspersus Steindachner, 1877a: 135, pl. 11 (fig. 2). Type locality: Stromgebiete des Amazonenflusses
von Santarem bis Tabatinga ... im Xingu bei Porto do Moz, im Amazonenstrome bei Cudajas und bei Tabatinga[Brazil]. Syntypes: MCZ 8223 (8), MCZ 8245 + 8265 (8), MCZ 18266 (1), NMW 47667 (2), NMW 47668 (2),
NMW 47669 (2), NMW 47672 (2), NMW 47673 (2), USNM 41518 (1).
Decapogon verissimi Miranda Ribeiro, 1911: 154, pl. 36 (fig. 1). Type locality: Pará [Brazil]. Lectotype: MNRJ
774a, designated by Miranda Ribeiro (1953: 395).
Distribution: Amazon River basin (Reis, 2003).
Dianema urostriatum (Miranda Ribeiro, 1912)
Decapogon urostriatum Miranda Ribeiro, 1912: 15. Type locality: Manáos [Brazil]. Lectotype: MNRJ 1067a, designated by Miranda Ribeiro (1953: 395). Name spelled Decapogon urotriatum in heading of account, but corrected to urostriatum in attached printed corrigendum.
Distribution: Amazon River basin, Brazil (Reis, 2003).
HOPLOSTERNUM
Hoplosternum Gill, 1858: 395. Type species: Callichthys laevigatus Valenciennes, 1836. Type by original designation. Gender: Neuter.
Cascadura Ellis, 1913: 387. Type species: Cascadura maculocephala Ellis, 1913. Type by monotypy. Gender: Feminine.
Cataphractops Fowler, 1915a: 231. Type species: Callichthys melampterus Cope, 1872. Type by original designation. Gender: Masculine. Originally proposed as a subgenus of Hoplosternum.
Ellisichthys Miranda Ribeiro, 1920: 11. Type species: Cascadura maculocephala Ellis, 1913. Type by being a
replacement name. Gender: Masculine. Proposed as a replacement for Cascadura Ellis, 1913, considered to be
an inappropriate choice for a scientific name.
Diasternum Franz, 2001: 17. Type species: Hoplosternum punctatum Meek & Hildebrand, 1916. Type by original
designation. Gender: Neuter.
Revision: Reis (1997).
Remarks: Lundberg (1997) reports a fossil identified as Hoplosternum sp. from the middle Miocene in the La Venta
formation, Magdalena River basin, Colombia.
Hoplosternum littorale (Hancock, 1828)
Callichthys littoralis Hancock, 1828: 244, pl. 32 (fig. 1). Type locality: Demerara [Guyana]. BMNH 1996.7.11.1.
Callichthys laevigatus Valenciennes, 1836, in Valenciennes, 1835–47: pl. 5 (fig. 2). Type locality: l’Ameriquedepuis Corrientes jusque dans les eaux douces des isles des Antilles, car nous l’avons de la Trinité. Holotype:possibly MNHN 0000-4271 (2), MNHN 0000-4272 (2), or MNHN 0000-4274 (1). Name available from plate,with distribution in Valenciennes (1847: 7); description in Valenciennes (1840b: 314), with location given as
Buénos-Ayres. Illustrated specimen must be treated as a holotype if it can be recognized among the listed specimens.
Callichthys subulatus Valenciennes, in Cuvier & Valenciennes, 1840b: 311 (231 in the Strasbourg deluxe edition).
Type locality: Cayenne, ... Buénos-Ayres. Syntypes: MNHN 0000-4268 (1), MNHN 0000-4269 (2), MNHN
0000-4270 (1).

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Callichthys albidus Valenciennes, in Cuvier & Valenciennes, 1840b: 316 (235 in the Strasbourg deluxe edition).
Type locality: Cayenne. Syntypes: MNHN 0000-4276 (1), MNHN 0000-4277 (1).
Callichthys chiquitos Castelnau, 1855: 38, pl. 18 (fig. 2). Type locality: la province de Chiquitos [Bolivia]. Holotype: MNHN 0000-4285.
Hoplosternum stevardii Gill, 1858: 401. Type locality: Island of Trinidad, West Indies. Syntypes (2): USNM 5944
(1).
Callichthys melampterus Cope, 1872a: 275, pl. 14 (fig. 4). Type locality: Rio Ambyiacu, Peru. Lectotype: ANSP
8318, designated by Reis (1997: 303).
Hoplosternum schreineri Miranda Ribeiro, 1911: 150, pl. 35 (fig. 3). Type locality: Amazonas, Brazil. Lectotype:
MNRJ 1051a, designated by Miranda Ribeiro (1953: 396).
Cascadura maculocephala Ellis, 1913: 387, pl. 25 (fig. 3). Type locality: Uruguayana [Rio Grande do Sul, Brazil].
Holotype: FMNH 54878.
Hoplosternum shirui Fowler, 1940a: 232, figs. 18–20. Type locality: Ucayali River basin, Contamana, Peru. Holotype: ANSP 68648.
Hoplosternum littorale daillyi Hoedeman, 1952b: 7. Type locality: Suriname (near Paramaribo). Holotype: ZMA
100277a.
Hoplosternum thoracatum cayennae Hoedeman, 1961: 130. Type locality: French Guiana: Ile de Cayenne. Holotype: ZMA 102238a.
Distribution: Most cis-Andean South American river drainages north of Buenos Aires (Reis, 2003).
Hoplosternum magdalenae Eigenmann, 1913
Hoplosternum magdalenae Eigenmann, in Ellis, 1913: 412. Type locality: Soplaviento, Colombia. Holotype: FMNH
56281; holotype illustrated in Eigenmann (1922b: 65, pl. 8, figs. 3, 4). Also described as new in Eigenmann(1913: 30).
Distribution: Magdalena and Sinu River basins, Colombia (Reis, 2003).
Hoplosternum punctatum Meek & Hildebrand, 1916
Hoplosternum punctatum Meek & Hildebrand, 1916: 264, pl. 14. Type locality: Rio Marte Arnade, 6 miles east of
Panama City, Panama. Holotype: FMNH 8943.
Distribution: Pacific coastal drainages of Panama and Atrato River basin, Colombia (Reis, 2003).
LEPTHOPLOSTERNUM
Lepthoplosternum Reis, 1997: 315. Type species: Callichthys pectoralis Boulenger, 1895. Type by original designation. Gender: Neuter.
Revision: Reis (1997).
Key: Reis & Kaefer (2005).
Lepthoplosternum altamazonicum Reis, 1997
Lepthoplosternum altamazonicum Reis, 1997: 318, fig. 14. Type locality: Peru: Pucallpa: Río Ucayali. Holotype:
MHNG 2551.01.
Distribution: Upper Amazon River basin, Peru (Reis, 2003).
Lepthoplosternum beni Reis, 1997
Lepthoplosternum beni Reis, 1997: 319, fig. 15. Type locality: Peru: Madre de Dios: Tambopata Reserve (12º15'S,
69º17'W). Holotype: USNM 329424.
Distribution: Madeira River basin of Bolivia and Peru (Reis, 2003).
Lepthoplosternum pectorale (Boulenger, 1895)
Callichthys pectoralis Boulenger, 1895b: 525. Type locality: Monte Sociedad, Paraguayan Chaco. Syntypes: BMNH
1895.5.17.57–61; syntype illustrated in Boulenger (1896c: pl. 4, fig. 4).
Distribution: Paraguay River basin (Reis, 2003).
Lepthoplosternum stellatum Reis & Kaefer, 2005
Lepthoplosternum stellatum Reis & Kaefer, 2005: 725, fig. 2. Type locality: Brazil, Amazonas, Tefé, Igarapé Remartimento, tributary of Lago Tefé, 1.5 km downstream from bridge on Tefé-Agrovila road, 03º24'28"S,64º44'10"W. Holotype: MCP 35599.

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FERRARISDistribution: Tefé Lake tributary, Upper Amazon River basin, Brazil (Reis & Kaefer, 2005).
Lepthoplosternum tordilho Reis, 1997
Lepthoplosternum tordilho Reis, 1997: 321, fig. 16. Type locality: Brazil: Rio Grande do Sul: Marginal lagoon nearsanga do Junco, on dust road between hwy BR-116 and Barra do Ribeiro. Holotype: MCP 16310.
Distribution: Northern tributaries of Laguna dos Patos in southern Brazil (Reis, 2003).
Lepthoplosternum ucamara Reis & Kaefer, 2005
Lepthoplosternum ucamara Reis & Kaefer, 2005: 728, fig. 4. Type locality: Peru, Loreto, Pacay-Samiria National
Reserve, Río Ucayali drainage, small creek into cocha Zapote, tributary to Rio Pacaya, 05º20'16"S, 74º29'10"W.
Holotype: MUSM 23000.
Distribution: Ucayali River, Peru and Japurá River mouth, Brazil (Reis & Kaefer, 2005).
MEGALECHIS
Megalechis Reis, 1997: 310. Type species: Callichthys thoracatus Valenciennes, 1840. Type by original designation.
Gender: Feminine.
Revision: Reis (1997).
Remarks: Nomenclature of species in this genus corrected by Reis et al. (2005).
Megalechis picta (Müller & Troschel, 1848)
Callichthys pictus Müller & Troschel, 1848: 630. Type locality: Britisch-Guiana. Syntype: ZMB 3151 (1).
Callichthys sulcatus Kner, 1855a: 110. Type locality: Rio branco und Marabitanas [Brazil]. Lectotype: NMW 4775,
designated by (Reis, 1997: 311). Originally Cal. sulcatus.
Hoplosternum oronocoi Fowler, 1915a: 229, fig. 8. Type locality: La Pedrita, Cano Uracoa, Venezuela. Holotype:
ANSP 37895.
Distribution: Amazon, Orinoco and upper Essequibo River basins, and coastal rivers of northern Brazil (Reis et al.,2005: 681).
Megalechis thoracata (Valenciennes, 1840)
Callichthys thoracatus Valenciennes, in Cuvier & Valenciennes, 1840b: 309 (230 in the Strasbourg deluxe edition),pl. 443. Type locality: la Mana [French Guiana]. Holotype: MNHN 0000-4266.
Callichthys longifilis Valenciennes, in Cuvier & Valenciennes, 1840b: 317 (235 in the Strasbourg deluxe edition).
Type locality: Cayenne. Holotype: MNHN 0000-4284.
Callychthys personatus Ranzani, 1841: 113. Type locality: [Unknown, apparently Brazil]. Holotype: MZUB 934.
Illustrated and described in more detail in Ranzani (1842: 322, pl. 24, fig.1).
Callichthys exaratus Müller & Troschel, 1848: 630. Type locality: Britisch-Guiana. Type (s): Whereabouts
unknown.
Hoplosternum thoracatum surinamensis Hoedeman, 1952b: 5. Type locality: Surinam. Holotype: ZMA 100292.
Distribution: Amazon, Orinoco and Paraguay River basins, and coastal rivers of the Guianas and northern Brazil(Reis et al., 2005).
SCLEROMYSTAX Günther, 1864
Scleromystax Günther, 1864: 225, 229. Type species: Callichthys barbatus Quoy & Gaimard, 1824. Type by monotypy. Gender: Masculine. Described in key (p. 225) as a subgenus of Callichthys, with species on p. 229.
Remarks: Recently removed from the synonymy of Corydoras by Britto (2003).
Scleromystax barbatus (Quoy & Gaimard, 1824)
Callichthys barbatus Quoy & Gaimard, 1824: 234. Type locality: ...petit ruisseaux qui aurosent, près de Rio de Janeiro, la ferme de Mandioca; [now: Fazenda da Japuhyba near Angra dos Reis, Rio de Janeiro, Brazil, 22º59'S,
44º17'W, by neotype designation]. Neotype: CAS 16006, designated by Nijssen & Isbrücker (1980b: 202).
Corydoras kronei Miranda Ribeiro, 1907a: 189, figure on p. 189. Type locality: buracos do leito dos Rios Una da
Aldeia e Despraiado [Iporanga, Brazil]. Lectotype: MNRJ 918A, designated by Nijssen & Isbrücker (1980b:
202).
Corydoras eigenmanni Ihering, 1907: 34. Type locality: Riverlets by Cubatão and Raiz da Serra, State of São Paulo.
Lectotype: MZUSP 153, designated by Britski (1969: 206).

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Distribution: Coastal drainages from Rio de Janeiro to Santa Catarina States, Brazil (Reis, 2003).
Scleromystax macropterus (Regan, 1913)
Corydoras macropterus Regan, 1913a: 232. Type locality: Paranagua, Brazil. Lectotype: BMNH 1913.1.1.11, designated by Nijssen & Isbrücker (1980b: 206).
Corydoras bertoni Eigenmann, in Eigenmann & Allen, 1942: 179. Type locality: Puerto Bertoni, Alto Paraná, Paraguay. Holotype (32 mm FL): IU 15441 (apparently not found at CAS).
Distribution: Coastal rivers of Brazil from São Paulo to Santa Catarina States, and some tributaries of the upper
Paraná River (Reis, 2003).
Remarks: See Buckup & Malabarba (1990) for comments on the type locality of Corydoras macropterus.
Scleromystax prionotos (Nijssen & Isbrücker, 1980)
Corydoras prionotos Nijssen & Isbrücker, 1980a: 5, figs. 6–7. Type locality: Linhares, 19º22'S, 40º04'W, Lagoa
Juparanã, Rio Doce system [Brazil]. Holotype: MNRJ 10537.
Distribution: Coastal rivers in southeastern Brazil from Espirito Santo to São Paulo, Brazil (Reis, 2003).
Scleromystax salmacis Britto & Reis, 2005
Scleromystax salmacis Britto & Reis, 2005: 482, figs. 1–2. Type locality: Brazil, Estado de Santa Catarina, Praia
Grande, arroio Molha Coco in Vila Rosa nearly 5.5 km northwest from Praia Grande, 29º10'14"S 49º58'53"W.
Holotype: MCP 38388.
Distribution: Mampituba, Araranguá and Ratones river basins, Santa Catarina State, Brazil (Britto & Reis, 2005).

CETOPSIDAE Bleeker, 1858
Cetopsini Bleeker, 1858b: 49, 250, 257. Type genus: Cetopsis Spix & Agassiz, 1829.
Helogenidae Regan, 1911: 573. Type genus: Helogenes Günther, 1863.
Taxonomic summary: Vari & Ferraris (2003).
Revision: Vari et al. (2005), Cetopsinae; Vari & Ortega (1986), Helogeninae.
Monophyly: de Pinna & Vari (1995).
5 genera, 41 species; no named fossil taxa.
CETOPSIDIUM Vari, Ferraris & de Pinna, 2005
Cetopsidium Vari, Ferraris & de Pinna, 2005: 133. Type species: Pseudocetopsis orientale Vari, Ferraris & Keith,
2003. Type by original designation. Gender: Neuter.
Cetopsidium ferreirai Vari, Ferraris & de Pinna, 2005
Cetopsidium ferreirai Vari, Ferraris & de Pinna, 2005: 133, fig. 1. Type locality: Brazil. Pará: Rio Trombetas basin,
Rio Mapuera, last cachoeira before Rio Trombetas (approximately 1º05'S, 57º01'W). Holotype: INPA 10194.
Distribution: Trombetas River basin, Brazil (Vari et al., 2005: 135).
Cetopsidium minutum (Eigenmann, 1912)
Hemicetopsis minutus Eigenmann, 1912b: 211, pl. 23 (fig. 2). Type locality: Amatuk Cataract, British Guiana. Holotype: FMNH 53262.
Distribution: Potaro and Middle Essequibo Rivers, Guyana (Vari et al., 2005: 137).
Cetopsidium morenoi (Fernández-Yépez, 1972)
Hemicetopsis morenoi Fernández-Yépez, 1972b: 19. Type locality: Río Aguaro, 1 km. abajo del paso Garcerito al
Oeste franco de Santa Rita, Estado Guárico, Venezuela. Holotype: author’s personal collection, whereabouts
unknown.
Distribution: Central and western tributaries of Orinoco River basin, Colombia and Venezuela (Vari et al., 2005:
141).
Cetopsidium orientale (Vari, Ferraris & Keith, 2003)
Pseudocetopsis orientale Vari, Ferraris & Keith, 2003: 693, fig. 1. Type locality: Suriname, Brokopondo District,
Mindrineti Kreek, close to mouth of Maykaboeka Kreek, Saramacca River basin, on Gros Rosevel Mining concession (5º07'08.8"N, 55º16'59.4"W). Holotype: MHNG 2621-040.

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Cetopsidium pemon Vari, Ferraris & de Pinna, 2005
Cetopsidium pemon Vari, Ferraris & de Pinna, 2005: 144, fig. 8–9. Type locality: Venezuela. Bolivar: Río Caroni, atmouth of Río Claro (7º54'45"N, 63º02'35"W). Holotype: MCNG 19475.
Distribution: Caroni, Caura and Meta river basins, Venezuela and Colombia, Branco River, Brazil (Vari et al., 2005:
146).
Cetopsidium roae Vari, Ferraris & de Pinna, 2005
Cetopsidium roae Vari, Ferraris & de Pinna, 2005: 147, fig. 10. Type locality: Guyana. Rupununi River, Karanambo(3º45'N, 58º18'W). Holotype: BMNH 1972.7.27.629.
Distribution: Rupununi region, Guyana (Vari et al., 2005: 149).
CETOPSIS Spix & Agassiz, 1829
Cetopsis Spix & Agassiz, 1829: 11. Type species: Silurus coecutiens Lichtenstein, 1819. Type by subsequent designation by Bleeker (1862–63: 16). Gender: Feminine.
Hemicetopsis Bleeker, 1862c: 403. Type species: Cetopsis candiru Spix & Agassiz, 1829. Type by monotypy. Gender: Feminine.
Pseudocetopsis Bleeker, 1862c: 403. Type species: Cetopsis gobioides Kner, 1857. Type by monotypy. Gender:
Feminine.
Bathycetopsis Lundberg & Rapp Py-Daniel, 1994: 382. Type species: Bathycetopsis oliveirai Lundberg & Rapp PyDaniel, 1994. Type by original designation. Gender: Feminine.
Cetopsis amphiloxa (Eigenmann, 1914)
Hemicetopsis amphiloxus Eigenmann, in Eigenmann, Henn & Wilson, 1914: 14. Type locality: Creek near San
Lorenzo, [Río] Patía basin, Colombia. Holotype: FMNH 56519; holotype illustrated in Eigenmann (1922b: 57,
pl. 3, fig. 2) and Vari et al. (2005: 151, fig. 11).
Distribution: Patía, upper San Juan and Atrato River basins, Pacific versant, western Colombia, and rivers of northwestern Ecuador (Vari et al., 2005: 152).
Cetopsis arcana Vari, Ferraris & de Pinna, 2005
Cetopsis arcana Vari, Ferraris & de Pinna, 2005: 153, fig. 13. Type locality: Brazil. Goiás: São Domingos, ParqueEstadual de Terra Ronca, Boca da Lapa de Bezerra (13º32'43"S, 46º22'35"W). Holotype: MZUSP 58637.
Distribution: Middle and upper Tocantins River basin (Vari et al., 2005:155).
Cetopsis baudoensis (Dahl, 1960)
Pseudocetopsis baudoênsis Dahl, 1960a: 452, figured on p. 453. Type locality: Quitasol, Baudó, western Colombia.
Holotype: ICNMHN 118.
Distribution: Baudó River basin, Pacific versant, Colombia (Vari et al., 2005:157).
Cetopsis caiapo Vari, Ferraris & de Pinna, 2005
Cetopsis caiapo Vari, Ferraris & de Pinna, 2005: 157, fig. 15. Type locality: Brazil: Goiás: Município de Minaçu/
Cavalcante, Rio Tocantins, at location of the future dam of the Usina Hidroeléletrica Serra da Mesa (13º50'S,
48º19'W). Holotype: MNRJ 13236.
Distribution: Tocantins River basin, Brazil (Vari et al., 2005:159).
Cetopsis candiru Spix & Agassiz, 1829
Cetopsis candiru Spix & Agassiz, 1829: 13, pl. 10 (fig. 1). Type locality: Brasiliae aequatoralis fluviis. Lectotype:
MHNG 210.05, designated by Vari et al. (2005: 160); MNHG syntype illustrated in Mahnert (1985: 12, fig. 4).
Name first appeared in Cuvier (1829) as a nomen nudum.
Cetopsis Spixii Swainson, 1839: 308. Type locality: Brasiliae aequatoralis fluviis. Syntypes: MHNG 210.05 (1),
MHNN 735–7 (3). Unneeded new name for Cetopsis candiru Spix & Agassiz, 1829.
Distribution: Amazon River basin (Vari et al., 2005: 160).
Cetopsis coecutiens (Lichtenstein, 1819)
Silurus coecutiens Lichtenstein, 1819: 61. Type locality: Brasilien. Holotype: ZMB 3093, apparently no longer
extant (Vari et al., 2005: 164).
Silurus cæcutiens Günther, 1864: 199. Type locality: Brazil. Holotype: ZMB 3093, apparently no longer extant (VariCHECKLIST OF CATFISHES, RECENT AND FOSSIL

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et al., 2005: 164). Unjustified emendation of Silurus coecutiens Lichtenstein, 1819.
Distribution: Amazon, Tocantins and Orinoco River basins (Vari et al., 2005: 164).
Cetopsis fimbriata Vari, Ferraris & de Pinna, 2005
Cetopsis fimbriata Vari, Ferraris & de Pinna, 2005: 165, fig. 20. Type locality: Colombia. Choco: Creek of upper
Río Nercua (approximately 7º01'N, 77º30'W), large tributary of Río Truando. Holotype: USNM 305348.
Distribution: Nercua River, a tributary of the Truando River which flows into the Atrato basin of the Caribbean Sea
versant of northwestern Colombia (Vari et al., 2005: 167).
Cetopsis gobioides Kner, 1857
Cetopsis gobioides Kner, 1857: 407, pl. 6 (fig. 16). Type locality: Irisanga [Brazil]. Syntypes: NMW 47378 (2).
Cetopsis chalmersi Norman, 1926a: 116. Type locality: Rio das Velhas about 32 miles north of Bello (=Belo) Horizonte, Brazil. Lectotype: BMNH 1925.10.1.4, designated by Vari et al. (2005: 171).
Distribution: Upper São Francisco, Paraná and Uruguay River basins and Juquiá River of coastal Brazil (Vari et al.,2005: 170).
Remarks: Silurus pygmaeus Natterer is an unavailable manuscript name published in the account of Cetopsis gobioides Kner (1857).
Cetopsis jurubidae (Fowler, 1944)
Pseudocetopsis jurubidae Fowler, 1944b: 235, figs. 7–9. Type locality: Clear waters of Rio Jurubidá, Nuquí [northwestern Colombia]. Holotype: ANSP 71430. Holotype illustrated in Vari et al. (2005: 173, figs. 23–24).
Distribution: Jurubidá River basin, Pacific versant, Colombia (Vari et al., 2005: 174).
Cetopsis montana Vari, Ferraris & de Pinna, 2005
Cetopsis montana Vari, Ferraris & de Pinna, 2005: 174, fig. 26. Type locality: Peru. Amazonas: 100 m upriver from
Caterpiza (approximately 3º55'S, 77º42'W). Holotype: LACM 41989-1.
Distribution: Amazon River basin of northeastern Peru and southeastern Ecuador (Vari et al., 2005: 150).
Cetopsis motatanensis (Schultz, 1944)
Pseudocetopsis plumbeus motatanensis Schultz, 1944c: 255, pl. 5 (fig. b). Type locality: Río Motatán, 4 km. above
Motatán, Venezuela. Holotype: USNM 121265.
Distribution: Lake Maracaibo basin of Venezuela and Colombia (Vari et al., 2005: 178).
Cetopsis oliveirai (Lundberg & Rapp Py-Daniel, 1994)
Bathycetopsis oliveirai Lundberg & Rapp Py-Daniel, 1994: 383, figs. 1, 3–4. Type locality: Brazil, Amazonas State,
Rio Solimões north of Ilha da Marchantaria, approximately 15 km upstream of the mouth of Rio Negro at
Manaus, approximately 15 m depth, 3º15'S, 60º00'W. Holotype: INPA 4439.
Distribution: Middle and upper Amazon River basin, Brazil and Peru (Vari et al., 2005: 180).
Cetopsis orinoco (Schultz, 1944)
Pseudocetopsis plumbeus orinoco Schultz, 1944c: 253, pl. 5 (fig. a). Type locality: Río Torbes, 1 km. above Táriba,
Orinoco system, Venezuela. Holotype: USNM 121263.
Distribution: Orinoco River basin and Aroa and Yaracuy river basins of coastal Venezuela (Vari et al., 2005: 183).
Cetopsis othonops (Eigenmann, 1912)
Hemicetopsis othonops Eigenmann, 1912a: 17. Type locality: Girardot, Colombia. Holotype: FMNH 56040; holotype illustrated in Eigenmann (1922b: 56, pl. 3, fig. 1).
Distribution: Magdalena and Sinu River basins (Vari et al., 2005); also Cauca, San Jorge, Apulo and Catatumbo
basins, Colombia (Maldonado Campo et al., 2005).
Cetopsis parma Oliviera, Vari & Ferraris, 2001
Cetopsis parma Oliviera, Vari & Ferraris, 2001: 575, fig. 1. Type locality: Peru. Departamento de Ucayali, Provincia
Coronel Portillo. Río Tambo, Río Ucayali basin, Pucallpa, Atalaya (8º23'S, 74º32'W). Holotype: MUSM 2266;
illustrated in Vari et al. (2005: 187, fig. 35).
Distribution: Upper Amazon River basin (Vari et al., 2005: 188).
Cetopsis pearsoni Vari, Ferraris & de Pinna, 2005
Cetopsis pearsoni Vari, Ferraris & de Pinna, 2005: 189, fig. 36. Type locality: Peru. Puno: Provincia de Sandia, Río
Candamo, Zona Reserva Tambopata (13º28.97'S, 69º50.08'W), 358 m elevation. Holotype: MUSM 11093.
Distribution: Upper Madeira River basin, Peru and Bolivia (Vari et al., 2005: 190).

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Cetopsis plumbea (Steindachner, 1882)
Cetopsis plumbeus Steindachner, 1882b: 178. Type locality: Canelos [Ecuador]. Lectotype: NMW 47381-1 (46 mm
SL), designated by Vari et al. (2005: 194). Species illustrated and described in more detail in Steindachner(1882c: 31, pl. 6, fig. 3).
Cetopsis macroteronema Boulenger, 1898c: 8. Type locality: Rio Zamora, Equateur oriental. Syntypes: BMNH
1898.11.4.11 (1), ZMUT 1548 (1)Distribution: Upper Amazon River basin, Ecuador and Peru (Vari et al., 2005: 193).
Cetopsis sandrae Vari, Ferraris & de Pinna, 2005
Cetopsis sandrae Vari, Ferraris & de Pinna, 2005: 195, figs. 39–40. Type locality: Brazil. Mato Grosso: Nova
Mutum, unnamed Corrego, Fazenda Buriti, approximately 1.5 km S of Fazenda (13º51'52"S, 56º11'36"W).
Holotype: MZUSP 61091.
Distribution: Arinos River, upper Tapajós River basin, Brazil (Vari et al., 2005: 150).
Cetopsis sarcodes Vari, Ferraris & de Pinna, 2005
Cetopsis sarcodes Vari, Ferraris & de Pinna, 2005: 197, fig. 42. Type locality: Brazil. Goiás: Município de Minaçu/
Cavalcante, Rio Tocantins between site of future dam of Usina Hidroeléctrica Cana Brava and Ilha do Orgulho
and Porto do Garimpo. Holotype: MNRJ 13024.
Distribution: Tocantins River basin, Brazil (Vari et al., 2005:199).
Cetopsis starnesi Vari, Ferraris & de Pinna, 2005
Cetopsis starnesi Vari, Ferraris & de Pinna, 2005: 199, fig. 43. Type locality: Bolivia. Tarija: Argentinian-Bolivian
border, Río Bermejo, 4–5 km S of Pueblo Salado, approximately 30 air km S of Bermejo (22º27'S, 64º32'W).
Holotype: USNM 314309.
Distribution: Northwestern part of La Plata River basin and upper Madeira River basin, Bolivia (Vari et al., 2005:
201).
Cetopsis umbrosa Vari, Ferraris & de Pinna, 2005
Cetopsis umbrosa Vari, Ferraris & de Pinna, 2005: 201, fig. 44. Type locality: Colombia. Meta: Río Negro, downstream from main Villavicencio to Puerto Lopez highway at La Balsa (04º04'N, 73º04'W). Holotype: ANSP
137559.
Distribution: Orinoco River basin, Colombia (Vari et al., 2005: 203).
DENTICETOPSIS Ferraris, 1996
Denticetopsis Ferraris, 1996: 162. Type species: Denticetopsis sauli Ferraris, 1996. Type by original designation.
Gender: Feminine.
Denticetopsis epa Vari, Ferraris & de Pinna, 2005
Denticetopsis epa Vari, Ferraris & de Pinna, 2005: 206, fig. 45. Type locality: Brazil. Pará: Igarapé do Limão, Rio
Tocantins, Baião (2º41'S, 49º41'W). Holotype: MZUSP 24060.
Distribution: Lower Tocantins River basin, Brazil (Vari et al., 2005: 208).
Denticetopsis iwokrama Vari, Ferraris & de Pinna, 2005
Denticetopsis iwokrama Vari, Ferraris & de Pinna, 2005: 208, fig. 47. Type locality: Guyana. Essequibo: Siparuni
River at Tumble Down Creek, Essequibo River basin (4º48'39"N, 58º51'11"W). Holotype: ANSP 177215.
Distribution: Siparuni River basin, Guyana (Vari et al., 2005: 210).
Denticetopsis macilenta (Eigenmann, 1912)
Hemicetopsis macilentus Eigenmann, 1912b: 211, pl. 23 (fig. 1). Type locality: Creek below Potaro Landing, British
Guiana. Holotype: FMNH 53260; illustrated in Vari et al. (2005: 211, figs. 48–49).
Distribution: Potaro River, Essequibo River basin, Guyana (Vari et al., 2005: 212).
Denticetopsis praecox (Ferraris & Brown, 1991)
Pseudocetopsis praecox Ferraris & Brown, 1991: 162, fig. 1. Type locality: Neblina base camp, Río Mawarinuma of
the Río Baria drainage, Territorio Federal Amazonas, southern Venezuela, 0º55'N, 66º10'W, elevation 120 m.
Holotype: AMNH 74446.
Distribution: Baria River, Upper Negro River basin, Venezuela (Vari et al., 2005: 215).
Denticetopsis royeroi Ferraris, 1996

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Denticetopsis royeroi Ferraris, 1996: 167, figs. 4, 12. Type locality: Venezuela: Estado Amazonas, Caño Chola atcrossing of San Carlos de Río Negro to Solano Road, 1º58'N, 67º00'W. Holotype: MBUCV V-26785, illustrated
in Vari et al. (2005: 216, fig. 51).
Distribution: Upper Negro River basin, Venezuela; known only from holotype (Vari et al., 2005: 216).
Denticetopsis sauli Ferraris, 1996
Denticetopsis sauli Ferraris, 1996: 164, figs. 3, 11. Type locality: Venezuela, Estado Amazonas, Outflow stream
from series of morichales, ca. 5.0 km from mouth of Río Pamoni, 2º48'N, 65º53'W. Holotype: MBUCV V20300.
Distribution: Around mouth of Pamoni River, Venezuela; known only from type series (Vari et al., 2005: 218).
Denticetopsis seducta Vari, Ferraris & de Pinna, 2005
Denticetopsis seducta Vari, Ferraris & de Pinna, 2005: 218, fig. 53. Type locality: Peru. Loreto: Caño “Sabalito,”
small creek on terra firma approximately 25 mi (=km) S of Iquitos (4º14'45"S, 73º24'57"W). Holotype: ANSP
178466.
Distribution: Western Amazon River basin, and possibly southwestern Orinoco River basin (Vari et al., 2005: 221).
HELOGENES Günther, 1863
Helogenes Günther, 1863: 443. Type species: Helogenes marmoratus Günther, 1863. Type by monotypy. Gender:
Masculine.
Leyvaichthys Dahl, 1960b: 302. Type species: Leyvaichthys castaneus Dahl, 1960. Type by original designation.
Gender: Masculine. Spelled Leyvaichths in one place, precedence established by first-reviser action of Eschmeyer & Bailey (in Eschmeyer, 1990: 221).
Revision: Vari & Ortega (1986).
Helogenes castaneus (Dahl, 1960)
Leyvaichthys castaneus Dahl, 1960b: 303. Type locality: Small pond in a brook tributary to the Guayabero River,approx. one kilometer from the mouth of the brook, and 1500 meters from the expedition Camp Nº 1, situatedon the left bank of the Guayabero about 4 kilometers from Camp Thompson, Colombia. Holotype: Whereabouts
unknown.
Distribution: Upper Orinoco, Guaviare and Meta River basins, Colombia (Vari & Ferraris, 2003).
Helogenes gouldingi Vari & Ortega, 1986
Helogenes gouldingi Vari & Ortega, 1986: 7, fig. 3. Type locality: Brazil. Amazonas: Igarapé, 15 km from Humaitá
along Rio Madeira (7º31'S, 63º02'W). Holotype: MZUSP 28854.
Distribution: Madeira River basin, Brazil (Vari & Ferraris, 2003).
Helogenes marmoratus Günther, 1863
Helogenes marmoratus Günther, 1863: 443. Type locality: Essequibo River, Guiana. Lectotype: BMNH
1864.1.21.83, designated by Vari & Ortega (1986: 16).
Helogenes amazonae Delsman, 1941: 80. Type locality: Manaos [=Manaus, Amazonas, Brazil]. Holotype: IRSNB
545.
Helogenes unidorsalis Glodek & Carter, 1978: 77, fig. 1. Type locality: Eastern Ecuador, Rio Bobonaza between
Sarayacu and Montalvo. Holotype: FMNH 80463.
Distribution: Atlantic drainages of Guianas, upper Orinoco and Negro systems and upper Amazon River basin (Vari& Ferraris, 2003).
Helogenes uruyensis Fernández-Yépez, 1967
Helogenes marmoratus uruyensis Fernández-Yépez, 1967: 166, pls. 2, 3 (fig. 2). Type locality: Río Uruyén, Venezuela. Holotype: AFY 56613 (whereabouts unknown).
Distribution: Uruyén River basin, Venezuela (Vari & Ferraris, 2003).
PARACETOPSIS Bleeker, 1862
Paracetopsis Bleeker, 1862 (in Bleeker, 1862–63): 16. Type species: Paracetopsis bleekeri Bleeker, 1862. Type byoriginal designation. Gender: Feminine.
Paracetopsis Eigenmann & Bean, 1907: 615. Type species: Cetopsis occidentalis Steindachner, 1880. Type by orig-

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inal designation. Gender: Feminine. Preoccupied by Paracetopsis Bleeker, 1862.
Cetopsogiton Eigenmann & Bean, in Eigenmann, 1910: 398. Type species: Cetopsis occidentalis Steindachner,
1880. Type by being a replacement name. Gender: Masculine. Replacement for Paracetopsis Eigenmann &Bean, 1907, preoccupied by Paracetopsis Bleeker, 1862.
Paracetopsis atahualpa Vari, Ferraris & de Pinna, 2005
Paracetopsis atahualpa Vari, Ferraris & de Pinna, 2005: 223, fig. 54. Type locality: Peru. Tumbes: Río Tumbes,
Cabo Inga (3º58'00"S, 80º24'59"W). Holotype: MUSM 11560.
Distribution: Tumbes River basin, northwestern Peru and upper Zarumilla River basin, southwestern Ecuador (Variet al., 2005: 224).
Paracetopsis bleekeri Bleeker, 1862
Paracetopsis bleekeri Bleeker, 1862 (in Bleeker, 1862–63): 16. Type locality: Not stated. Neotype: NMW 47377,
designated by Vari et al. (2005: 227).
Cetopsis ventralis Gill, 1870: 95. Type locality: Maranon or upper Amazon, and Napo Rivers [in error, actually rivers of southwestern Ecuador]. Holotype: USNM 8307.
Cetopsis occidentalis Steindachner, 1880b: 99, pl. 8 (figs. 2, 2a). Type locality: Guayaquil [Ecuador]. Holotype:
NMW 47377.
Distribution: Guayas and Santa Rosa river basins, Pacific Ocean versant of southwestern Ecuador (Vari et al., 2005:
227).
Paracetopsis esmeraldas Vari, Ferraris & de Pinna, 2005
Paracetopsis esmeraldas Vari, Ferraris & de Pinna, 2005: 228, fig. 57. Type locality: Ecuador. Esmeraldas: Río
Blanco, 4 km above junction with Río Quininde (0º20'N, 79º28'W; Río Esmeraldas basin). Holotype: MCZ
48768.
Distribution: Pacific Ocean versant rivers of northwestern Ecuador (Vari et al., 2005: 230).

CHACIDAE Bleeker, 1858
Chacini Bleeker, 1858b: 49, 310, 322. Type genus: Chaca Gray, 1831.
Revision: Roberts (1982a).
Identification guide: Jayaram & Majumdar (1964), Jayaram (1980), South Asia.
Phylogeny: Brown & Ferraris (1988).
1 genus, 3 species; no named fossil taxa.
CHACA Gray, 1831
Chaca Gray, 1831: 9. Type species: Chaca hamiltonii Gray, 1831. Type by monotypy. Gender: Feminine.
Chaca Cuvier &Valenciennes, in Valenciennes, 1832b: 386. Type species: Chaca lophioides Cuvier & Valenciennes,
1832. Type by monotypy. Gender: Feminine. Preoccupied by Chaca Gray, 1831.
Brachystacus Van der Hoeven, 1849: 280. Type species: Platystacus chaca Hamilton, 1822. Type by monotypy.
Gender: Masculine.
Chaca bankanensis Bleeker, 1852
Chaca bankanensis Bleeker, 1852c: 455. Type locality: Banka, in fluviis. Holotype (68 mm TL): RMNH 5405.
Chaca bankae Giebel, 1857: 325. Type locality: Bangka. Either an emendation of, or misspelling for, Chaca bankanensis Bleeker, 1852.
Distribution: Southern Malay Peninsula, Sumatra and Borneo (Roberts, 1982a).
Remarks: BMNH 1863.12.4.117 (113 mm SL, 131 mm TL), is listed as a type of Chaca bankanensis, but is too large
to be the holotype.
Chaca burmensis Brown & Ferraris, 1988
Chaca burmensis Brown & Ferraris, 1988: 3, fig. 1. Type locality: Sittang River, Burma. Holotype: BMNH
1891.11.30.144.
Distribution: Lower Irrawaddy and Sittang River basins, Myanmar.

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Chaca chaca (Hamilton, 1822)
Platystacus chaca Hamilton, 1822: 140, 374, pl. 28 (fig. 43). Type locality: rivers and ponds of the northeasternparts of Bengal. No types known.
Chaca hamiltonii Gray, 1831: 9. Type locality: rivers and ponds of the northeastern parts of Bengal. New name forPlatystacus chaca Hamilton, 1822, to avoid tautonymy.
Chaca lophioides Cuvier & Valenciennes, in Valenciennes, 1832b: 386, pl. 4 (fig. 2). Type locality: Bengal. New
name for Platystacus chaca Hamilton, 1822, to avoid tautonymy. Also described in Cuvier & Valenciennes(1840b: 445 [p. 330 in Strasbourg deluxe edition], pl. 451).
Chaca buchanani Günther, 1864: 29. Type locality: Ganges. No types known. New name for Platystacus chaca
Hamilton, 1822, to avoid tautonymy.
Distribution: Ganges-Brahmaputra River system, India and Bangladesh (Roberts, 1982a).

CLARIIDAE Bonaparte, 1846
Clariini Bonaparte, 1846: 5. Type genus: Clarias Scopoli, 1777.
Heterobranchoidei Bleeker, 1858b: 33, 37, 41, 333. Type genus: Heterobranchus Geoffroy St. Hilaire, 1809.
Heteropneustidae Hora, 1936a: 209. Type genus: Heteropneustes Müller, 1840.
Taxonomic summary: Teugels (1986b, Africa).
Review: Jackson (1959, Nyasaland); Skelton & Teugels (1991, southern Africa); Teugels (1992a, west Africa).
Key to genera: Poll & Gosse (1994, Africa).
Identification guide: Jayaram (1980, South Asia).
Phylogeny: Teugels & Adriens (2003); Diogo et al. (2003a), Agnèse & Teugels (2005, African species); Jansen et al.
(2005, anguilliform species).
16 genera, 113 species; including 2 fossil species.
BATHYCLARIAS Jackson, 1959
Bathyclarias Jackson, 1959: 112. Type species: Clarias longibarbis Worthington, 1933. Type by original designation. Gender: Masculine.
Remarks: Removed from the synonymy of Dinotopterus by Anseaume & Teugels (1999).
Species flock evolution: Agnèse & Teugels (2001a).
Bathyclarias euryodon Jackson, 1959
Bathyclarias euryodon Jackson, 1959: 120, fig. 4. Type locality: Nkata Bay, Lake Nyasa. Holotype: BMNH
1960.2.29.13.
Distribution: Lake Malawi (Teugels, 1986b, as Dinotopterus euryodon).
Bathyclarias ilesi Jackson, 1959
Bathyclarias ilesi Jackson, 1959: 116, fig. 2. Type locality: Nkata Bay, Lake Nyasa. Holotype: BMNH 1960.2.29.8.
Distribution: Lake Malawi (Teugels, 1986b, as Dinotopterus ilesi).
Bathyclarias longibarbis (Worthington, 1933)
Clarias longibarbis Worthington, 1933b: 309, fig. 10. Type locality: Lake Nyasa. Holotype: BMNH
1932.11.15.592.
Distribution: Lake Malawi (Teugels, 1986b, as Dinotopterus longibarbis).
Bathyclarias rotundifrons Jackson, 1959
Bathyclarias rotundifrons Jackson, 1959: 118, fig. 3. Type locality: Lake Nyasa. Holotype: BMNH 1960.2.29.11.
Distribution: Lake Malawi (Teugels, 1986b, as Dinotopterus rotundifrons).
Bathyclarias worthingtoni Jackson, 1959
Bathyclarias worthingtoni Jackson, 1959: 123, fig. 6. Type locality: [Nkata Bay] Lake Nyasa. Holotype: BMNH
1960.2.29.18.
Distribution: Lake Malawi (Teugels, 1986b, as Dinotopterus worthingtoni).

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Channallabes Günther, 1873a: 143. Type species: Gymnallabes apus Günther, 1873. Type by monotypy. Gender:
Feminine. Originally proposed as a subgenus of Gymnallabes.
Channallabes apus (Günther, 1873)
Gymnallabes apus Günther, 1873a: 142. Type locality: Interior of Ambriz, Angola. Holotype: BMNH 1873.7.28.16.
Distribution: Congo River basin (Teugels, 1986b).
CLARIALLABES Boulenger, 1900
Clariallabes Boulenger, 1900a: 136. Type species: Clarias melas Boulenger, 1887. Type by original designation.
Gender: Feminine.
Allabenchelys Boulenger, 1902e: 234. Type species: Allabenchelys longicauda Boulenger, 1902. Type by monotypy.
Gender: Feminine.
Clariallabes attemsi (Holly, 1927)
Allabenchelys attemsi Holly, 1927b: 202, fig. 2. Type locality: Kamerun. Holotype: NMW 7897.
Distribution: Cameroon, known only from an indefinite type locality (Teugels, 1986b).
Clariallabes brevibarbis Pellegrin, 1913
Clariallabes brevibarbis Pellegrin, 1913b: 272. Type locality: Ngomo (Ogôoué). Holotype: MNHN 1913-0265.
Distribution: Ogowe River, Gabon (Teugels, 1986b).
Clariallabes centralis (Poll & Lambert, 1958)
Clarias centralis Poll & Lambert, 1958: 334, figure on p. 335. Type locality: Kunungu (Congo). Holotype: MRAC
29659.
Distribution: Central Congo River system (Teugels, 1986b).
Clariallabes heterocephalus Poll, 1967
Clariallabes heterocephalus Poll, 1967: 233, fig. 108. Type locality: Poste de Cuílo, Xá-Ua, riv. Luíta, (8º02'S,
19º25'E), Angola. Holotype: MRAC 162082.
Distribution: Luita River, Angola (Teugels, 1986b).
Clariallabes laticeps (Steindachner, 1911)
Allabenchelys laticeps Steindachner, 1911d: 532. Type locality: Mawambi, Ituri–Flüss. Syntypes (9): NMW 7649
(1), NMW 47988 (1), NMW 47989 (1), NMW 479890 (1), NMW 47991 (1).
Distribution: Ituri River, Aruwimi Basin, Congo River system; known only from type locality (Teugels, 1986b).
Clariallabes longicauda (Boulenger, 1902)
Allabenchelys longicauda Boulenger, 1902e: 234, pl. 22 (figs. 1–1a). Type locality: Ja River, flowing into the Sanga,
an affluent of the right bank of the Congo, French Congo, 250 miles from the coast. Holotype: BMNH
1902.11.12.129.
Allabenchelys brevior Boulenger, 1903c: 439. Type locality: Ja River, southern Cameroon. Syntypes (2): BMNH
1903.7.28.194–195 (2).
Distribution: Central Congo River system (Teugels, 1986b).
Clariallabes manyangae (Boulenger, 1919)
Allabenchelys manyangae Boulenger, 1919: 159. Type locality: Kinkenda (Manyanga), région des Cataractes du
Bas-Congo. Syntypes (3): BMNH 1919.8.28.12 (1), MRAC 1327 (1), MRAC 1329 (1).
Distribution: Manyanga, Kinkenda; known only from type locality (Teugels, 1986b).
Clariallabes melas (Boulenger, 1887)
Clarias melas Boulenger, 1887a: 148. Type locality: Lower Congo. Holotype: BMNH 1887.1.13.6. Illustrated inBoulenger (1902e: pl. 22, fig. 2), possibly based on the holotype.
Clariallabes melas lembesseensis Fowler, 1949: 259, figs. 48–52. Type locality: Oka, Congo system. Holotype:
ANSP 71906.
Distribution: Congo River system (Teugels, 1986b).
Clariallabes mutsindoziensis Taverne & De Vos, 1998
Clariallabes mutsindoziensis Taverne & De Vos, 1998: 212, figs. 1–3. Type locality: Rivière Mutsindozi près de sonembouchure dans la Malagarasi à proximité de Gihofi, Burundi, 4º02'–4º04'S, 30º09'–30º11'E. Holotype:

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MRAC 96-31-P-53.
Distribution: Mutsindozi River, Lake Tanganyika basin, Burundi (Taverne & De Vos, 1998).
Clariallabes petricola Greenwood, 1956
Clariallabes petricola Greenwood, 1956: 556, fig. 1. Type locality: Nile, about half a mile below the Owen-falls
dam, Jinja, Uganda. Holotype: BMNH 1956.10.9.204.
Distribution: Lake Victoria and the Victoria Nile, near the Owen-falls dam (Teugels, 1986b).
Clariallabes pietschmanni (Güntert, 1938)
Allabenchelys pietschmanni Güntert, 1938: 299. Type locality: Aus dem Njong [Cameroon]. Syntypes (26): NMW
46063, ? NMW 47933 (1). Also appeared in Güntert (1939: 25).
Distribution: Nyong River; known only from type locality (Teugels, 1986b).
Clariallabes platyprosopos Jubb, 1965
Clariallabes platyprosopos Jubb, 1965: 393, fig. 1. Type locality: Upper Zambezi River, about 15 miles above theVictoria Falls, Rhodesia. Holotype: AMG P1050.
Distribution: Upper Zambezi River (Teugels, 1986b).
Clariallabes simeonsi Poll, 1941
Clariallabes simeonsi Poll, 1941a: 4, figs. 2, 5. Type locality: Uélé, rivières Mosongolia, Mekokpa, et Matanga [Precise locality of holotype not stated], Congo belge. Holotype: IRSNB 61.
Distribution: Uélé River, Congo River basin (Teugels, 1986b).
Clariallabes teugelsi nomen novum
Clarias (Allabenchelys) dumerili longibarbis David & Poll, 1937: 232, fig. 21. Type locality: Léopoldville, Congo
Belge. Syntypes: MRAC 44011–013 (3). Preoccupied by Clarias longibarbis Worthington, 1933.
Distribution: central Congo River basin (Teugels, 1986b).
Remarks: The name Clarias (Allabenchelys) dumerili longibarbis David & Poll is a junior primary homonym ofClarias longibarbis Worthington and, as such, was permanently invalid upon publication. The species wastreated as valid, as Clariallabes dumerili David & Poll, 1937, by Teugels (1986b: 67). However, Clariallabes
dumerili was not proposed as an available name in David & Poll and it also was not made available by action of
Teugels. Teugels (1986b) lists no available synonyms for this name, so the name Clariallabes teugelsi is proposed here as a replacement for Clarias (Allabenchelys) dumerili longibarbis to provide a valid name for thisspecies.
Clariallabes uelensis (Poll, 1941)
Clarias (Allabenchelys) uelensis Poll, 1941a: 14, fig. 6. Type locality: rivières Mosongolia, Mekokpa, et Matanga,
région Ibembo, Uélé [precise locality of holotype not stated.]. Holotype: IRSNB 63.
Distribution: Mosongolia, Mekokpa or Matanga Rivers, known only from types (Teugels, 1986b).
Clariallabes variabilis Pellegrin, 1926
Clariallabes variabilis Pellegrin, 1926: 201. Type locality: Dumbi ..., Kabambaie ...., Belenge ...., Tshimbu ...,
Kamaiembi; Congo Belge. Syntypes (13): MNHN 1926-0151(1), MNHN 1926-0152 (1), MNHN 1926-0153(1), MNHN 1926-0154 (1), MRAC 19689 (1), MRAC 19690 (1), MRAC 19692 (1), MRAC 19694 (1), MRAC
19696 (1), MRAC 19702 (1), MRAC 19712 (1), MRAC 19713 (1), MRAC 19714 (1).
Distribution: Congo River system (Teugels, 1986b).
CLARIAS Scopoli, 1777
Chlarias Scopoli, 1777: 455. Type species: Silurus anguillaris Linnaeus 1758. Type by subsequent designation by
Teugels & Roberts (1987). Gender: Masculine. First published without any included species; names apparently
first added by Cuvier, 1816, but spelled Clarias therein. Scopoli based his name on Clarias Gronovius, a namepublished in a work that was rejected as non binomial (ICZN, opinion 261), but with an unexplained change inspelling. Spelling in Scopoli not followed in recent literature and not used here.
`
Macropteronotus La Cepède, 1803: 84. Type species: Macropteronotus charmuth La Cepède, 1803. Type by subsequent designation, apparently by Jordan (1917). Gender: Masculine.
Clarias Cuvier, 1816: 206. Type species not clearly established and may not have been proposed inasmuch as thename has been generally attributed to Scopoli or Gronovius. Proposed originally with two species. Gender:

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Masculine.
Cossyphus M’Clelland, 1844a: 403. Type species: Cossyphus ater M’Clelland, 1844. Type by monotypy. Gender:
Masculine. Preoccupied several times, first by Cossyphus Olivier, 1791, in Insecta; replaced by Phagorus
M’Clelland, 1844.
Phagorus M’Clelland, 1844b: 225 [errata]. Type species: Cossyphus ater M’Clelland, 1844. Type by being a
replacement name. Gender: Masculine. Replacement for Cossyphus M’Clelland, 1844; preoccupied by Cossyphus Olivier, 1791.
Dinotopteroides Fowler, 1930: 41. Type species: Dinotopteroides prentissgrayi Fowler, 1930. Type by original designation. Gender: Masculine.
Prophagorus Smith, 1939: 236. Type species: Clarias nieuhofii Valenciennes, 1840. Type by original designation.
Gender: Masculine.
Anguilloclarias Teugels, 1982a: 13. Type species: Clarias theodorae Weber, 1897. Type by original designation.
Gender: Masculine. Originally proposed as a subgenus of Clarias.
Brevicephaloides Teugels, 1982a: 14. Type species: Clarias camerunensis Lönnberg, 1895. Type by original designation. Gender: Masculine. Also appeared in Teugels (1982b: 739). Originally proposed as a subgenus of Clarias.
Clarioides Teugels, 1982a: 12. Type species: Clarias angolensis Steindachner, 1866. Type by original designation.
Gender: Masculine.
Platycephaloides Teugels, 1982a: 11. Type species: Clarias platycephalus Boulenger, 1902. Type by original designation. Gender: Masculine. Originally as a subgenus of Clarias. Also appeared in Teugels (1982b: 736).
Remarks: Clarioides was first proposed as subgenus of Clarias in David & Poll (1937: 231) but without a type species and, therefore, cannot be treated as available from that publication [ICZN Art. 13.3]. Heterobranchoides
was first proposed as a subgenus of Clarias by David (1935: 82, 99), but was not assigned a type species and thename is, therefore, not available from that publication. The name appears again in David & Poll (1937: 230), but
as a synonym of Clarias and not as a valid name. It is, therefore, also not available from that reference. See Teugels (1983) for comments on the validity of Dinotopteroides as a natural subgroup of Clarias.
Review: Debouche et al. (1979), Teugels (1982c, 1986a, African species).
Clarias agboyiensis Sydenham, 1980
Clarias (Clarioides) agboyiensis Sydenham, 1980: 660, figs. 1–2. Type locality: Agboyi Creek, a ditributary of theOgun, Lagos State, south-west Nigeria, 6º36'N, 5º24'E. Holotype: BMNH 1959.8.18.66.
Clarias (Clarioides) isheriensis Sydenham, 1980: 663, figs. 3–4. Type locality: Energo Project site, Lagos-Badagry
expressway, Lagos State, Nigeria, 6º29'N, 3º15'E. Holotype: BMNH 1976.3.17.3.
Distribution: West Africa, from Ghana to Nigeria (Teugels, 1986b).
Clarias albopunctatus Nichols & La Monte, 1953
Clarias albopunctatus Nichols & La Monte, 1953: 1, fig. 1. Type locality: Kotto River, about 225 miles north of
Bangassou, eastern French Equatorial Africa. Holotype: AMNH 19840; holotype illustrated by Teugels (1986a:
94, fig. 49).
Distribution: Lake Chad, Benue River and central Congo River (Teugels, 1986b).
Clarias alluaudi Boulenger, 1906
Clarias Alluaudi Boulenger, 1906b: 437. Type locality: Entebbe and Bunuako; [and] Kavirondo Bay, Lake Victoria[restricted to Bunuako by lectotype designation]. Lectotype: BMNH 1906.5.30.159, designated by Teugels(1986a: 120); lectotype illustrated by Seegers (1996: fig. 144).
Clarias hilgendorfi Boulenger, 1911c: 54. Type locality: Lake Rukwa. Holotype: BMNH 1911.3.27.41; illustrated in
Seegers (1996: fig. 146).
Distribution: Lakes Victoria, Kyoga, Edward, Rukwa and Tanganyika (Teugels, 1986b).
Clarias anfractus Ng, 1999
Clarias anfractus Ng, 1999d: 18, figs. 1a, 2a, 3a, 4a. Type locality: Borneo: Sabah, Danum, forest stream 600 m into
conservation area, tributary of Sungai Segama. Holotype: ZRC 42598.
Distribution: Segama and Kalabakan River drainages, northeastern Borneo (Ng, 1999d).
Clarias angolensis Steindachner, 1866

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Clarias angolensis Steindachner, 1866: 766, pl. 13 (figs. 4, 7). Type locality: Angola. Holotype: NMW 46891; holotype illustrated in Teugels (1986a: 68, fig. 35).
Clarias brevinuchalis Lönnberg & Rendahl, 1922: 123, fig. 2. Type locality: Upper Congo. Holotype: NRM 9640.
Clarias confluentus Nichols & La Monte, 1933b: 3, fig. 3. Type locality: Luluabourg, Kasai District, Belgian Congo.
Holotype: AMNH 12340.
Clarias (Clarioides) curtus Matthes, 1964: 102, pl. 4a. Type locality: Rivière Kululu (mare), Zaïre. Holotype:
MRAC 138667.
Distribution: Lower and Central Congo River system (Teugels, 1986b).
Clarias anguillaris (Linnaeus, 1758)
Silurus anguillaris Linnaeus, 1758: 305. Type locality: Nile. Syntype: NRM LP 71.
Clarias Hasselquistii Valenciennes, in Cuvier & Valenciennes, 1840b: 362 (280 in Strasbourg deluxe edition), pl.
446. Type locality: Nil. Syntypes: MNHN b-0287 (1), MNHN b-0288 (2).
Clarias Senegalensis Valenciennes, in Cuvier & Valenciennes, 1840b: 376 (279 in the Strasbourg deluxe edition).
Type locality: Sénégal. Syntypes: MNHN a-8667 (1, dry), MNHN a-9429 (1, dry).
Clarias parvimanus Günther, 1864: 15. Type locality: Nile. Holotype: BMNH 1859.7.29.21.
Clarias budgetti Boulenger, 1900e: 513. Type locality: River Gambia. Syntypes (3): BMNH 1901.7.17.4–6 (3).
Clarias anguillaris nigeriensis Pellegrin, 1923a: 213. Type locality: Branche nontante de la boucle du Niger entre
Mopti et Niafunké. Holotype: MNHN 1922-0161 (skull).
Distribution: Niger, Benoue, Gambia, lower Senegal and lower and middle Nile Rivers; Chad system; coastal riversof Benin, Togo, Ghana and Côte d’Ivoire; and relict populations in Mauritania and southern Algeria (Teugels,
1986b).
Remarks: Teugels & Roberts (1987: 95) list NRM LP 71 as the holotype of Silurus anguillaris, however, the originaldescription of that species lists a range of meristic values that implies that more than one specimen contributed
to the diagnosis. Thus, the specimen mentioned by Teugel & Roberts must be considered a syntype. See Benech
et al. (1993) for characters that distinguish this species from Clarias gariepinus.
Clarias batrachus (Linnaeus, 1758)
Silurus batrachus Linnaeus, 1758: 305. Type locality: Africa, Asia. Syntypes: Whereabouts unknown [not NRM LP
71 as reported by Fernholm & Wheeler (1983: 219–220); see discussion in Teugels & Roberts (1987: 95)].
Macropteronotus jagur Hamilton, 1822: 145, 374. Type locality: in ponds and ditches [Ganges River]. No types
known.
Macropteronotus magur Hamilton, 1822: 146, 374, pl. 26 (fig. 45). Type locality: in ponds and ditches [Ganges
River]. No types known.
Plotosus (Clarias) hamiltonii Swainson, 1839: 307. Type locality: in ponds and ditches [Ganges River]. No types
known. On “Ham. 146, pl. 26, f. 45” [= Hamilton (1822: 146, pl. 26, fig. 45)]. Unneeded new name for Macropteronotus magur Hamilton, 1822.
Clarias punctatus Valenciennes, in Cuvier & Valenciennes, 1840b: 384 (285 in the Strasbourg deluxe edition). Type
locality: Java. Holotype (10 pouces): at MNHN, whereabouts unknown.
Distribution: Widely distributed in South and Southeast Asia.
Remarks: More than one species is included within what is currently called Clarias batrachus. A clarification of thesituation requires both demarcation of species and replacement of the missing primary types of Macropteronotus jagur and M. magur.
Clarias batu Lim & Ng, 1999
Clarias batu Lim & Ng, 1999: 157, figs. 1, 2, 4a, 5a, 7. Type locality: Sungai Baharu, on right side of Tekek-Juara
trail, towards Juara, Pulau Tioman, Pahang, Malaysia. Holotype: ZRC 40087.
Distribution: Peninsular Malaysia.
? Clarias brachysoma Günther, 1864
Clarias brachysoma Günther, 1864: 20. Type locality: Ceylon. Syntypes: BMNH 1859.5.31.6–8 (3).
Distribution: Sri Lanka.
Clarias buettikoferi Steindachner, 1894
Clarias bulumae Steindachner, 1894: 55. Type locality: Buluma, Liberia. Holotype: RMNH 3340. Originally as:

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Clarias bulumae n. sp. ?
Clarias Büttikoferi Steindachner, 1894: 53. Type locality: Buluma, Liberia. Holotype: registered at RMNH, but
apparently not returned by Steindachner (Teugels, 1986a: 88).
Clarias liberiensis Steindachner, 1894: 54, pl. 3 (fig. 2). Type locality: Buluma und aus dem Junk-flusse, Liberia[restricted to Junk-Flusse by lectotype designation]. Lectotype: RMNH 3342, designated by Sydenham (1978:355); lectotype illustrated by Teugels (1986a: 47, fig. 47).
Clarias guineensis Osório, 1909: 104, pl. (fig. 3). Type locality: Guinée, Bolama, Postugo. Holotype: MB(destroyed).
Clarias dorsimarmoratus Ahl, 1937a: 239. Type locality: 40–50 km landeinwärts Freetown, Sierra Leone. Holotype:
ZMB 20939.
Distribution: Comoe River, Côte d’Ivoire, to Guinea Bissau (Teugels, 1986b).
Remarks: Teugels (1986a: 88) appears to be the first to treat all three Steindachner names as synonyms and thus
acted as first reviser in selecting Clarias buettikoferi as the valid name for the species.
Clarias buthupogon Sauvage, 1879
Clarias buthupogon Sauvage, 1879: 96. Type locality: Doumé [Fleuve Ogôoué]. Holotype: MNHN a-0896; holotype illustrated by Teugels (1986a: 80, fig. 44).
Clarias megapogon Sauvage, 1880c: 39. Type locality: Doumé [Fleuve Ogôoué]. Holotype: MNHN a-0896. Either
an emendation or lapsus calami for Clarias buthupogon Sauvage.
Clarias zygouron Nichols & Griscom, 1917: 704, fig. 18. Type locality: Malela, Congo. Holotype: AMNH 6698.
Clarias lindicus Boulenger, 1920a: 26, fig. 13. Type locality: Bafwasende sur la Lindi, Congo Belge. Holotype:
MRAC 7112.
Clarias obscurus Poll, 1948: 22, figs. 8–9. Type locality: Pweto, Lac Moero. Holotype: IRSNB 86.
Chlarias angolensis macronema Fowler, 1949: 257, figs. 43–47. Type locality: Oka, 18 mi. north of Eovo, Congosystem. Holotype: ANSP 71903.
Distribution: Coastal rivers, from Nigeria to the Congo River system (Teugels, 1986b).
Clarias camerunensis Lönnberg, 1895
Clarias camerunensis Lönnberg, 1895: 182. Type locality: Cameroons. Lectotype: ZMUU 283 (88 mm TL, largest
specimen); lectotype illustrated in, and designated by, Teugels (1986a: 154, fig. 85).
Clarias walkeri Günther, 1896a: 274, pl. 14 (fig. b). Type locality: Ogowe River. Holotype: BMNH 1896.5.5.54.
Clarias breviceps Boulenger, 1900a: 135, pl. 48 (fig. 6). Type locality: Marais Scala N’toto, Mayumbe. Syntypes:
BMNH 1899.11.27.93 (1), MRAC 271 (1).
Clarias poensis Boulenger, 1908b: 1078, fig. 252. Type locality: Fernando Po. Holotype: BMNH 1904.7.23.49.
Clarias monkei Keilhack, 1910: 119. Type locality: in einem Kleinen Bache im Urwald 2 km südlich von Logobaba(4º2'N, 9º45'Ö), Kamerun. Syntypes (3): ZMB 17769 (3).
Clarias duchaillui Fowler, 1915a: 226, fig. 7. Type locality: Gaboon. Holotype: ANSP 8567 [but published as ANSP
8568].
Clarias nigeriae Popta, 1919: 4. Type locality: Wari, mouth of the Niger, West Africa. Holotype: RMNH 10332.
Clarias (Allabenchelys) submarginatus thysvillensis David & Poll, 1937: 231. Type locality: Grottes de Thysville,
Congo Belge. Holotype: MRAC 37550.
Distribution: Coastal rivers of Togo to the lower and central Congo River system (Teugels, 1986b).
Clarias cataractus (Fowler, 1939)
Phagorus cataractus Fowler, 1939: 54, figs. 1–3. Type locality: Waterfall at Trang, Siam. Holotype: ANSP 68462.
Distribution: Peninsular Thailand and, possibly, Cambodia (Rainboth, 1996).
Clarias cavernicola Trewavas, 1936
Clarias cavernicola Trewavas, 1936, 70, pl. 2 (figs. 7–9). Type locality: Aigamas Cave, north of Otavi, South-West
Africa. Lectotype: BMNH 1935.3.20.174, designated by Teugels (1986a: 134).
Distribution: Caves of south west Africa (Teugels, 1986b).
Clarias dayi Hora, 1936
Clarias dayi Hora, 1936b: 350, figs. 3, 4c. Type locality: Wynaad [India]. Holotype: at ZSI [on “Day’s Wynaad
specimen”].

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Distribution: Tamil Nadu, India (Manimekalan & Arunachalam (2002).
Remarks: Treated in current literature as valid or as a synonym of Clarias dussumieri Valenciennes, 1840.
Clarias dhonti (Boulenger, 1920)
Allabenchelys dhonti Boulenger, 1920c: 402, fig. 8. Type locality: Kabeke, a village 30 miles south of Tumbwe, on
the Niemba River, near the west coast of Lake Tanganyika. Syntypes (10): BMNH 1919.7.24.20–27 (8), MNHN
1919-0467 (1), MRAC 6991 (1). One syntype (MRAC 6991) illustrated in Teugels (1986a: 176, fig. 91) but
incorrectly listed therein as the holotype.
Distribution: Niemba River, Democratic Republic of the Congo (Teugels, 1986b).
Clarias dialonensis Daget, 1962
Clarias dialonensis Daget, 1962: 120, fig. 39; pl. 11 (fig. 26). Type locality: Bafing près Mamou (Fouta Dialon).
Lectotype: MNHN 1959-0107; lectotype illustrated in, and designated by, Teugels (1986a: 145, fig. 81) as Clarias laeviceps dialonensis.
Distribution: Fouta Dialon, Guinea, and coastal rivers of Sierra Leone (Teugels, 1986b).
Remarks: Treated by Teugels (1986a: 145; 1986b: 88) as a subspecies of Clarias laeviceps.
Clarias dumerilii Steindachner, 1866
Clarias Dumerilii Steindachner, 1866: 766, pl. 14 (fig. 5). Type locality: Angola. Neotype: BMNH 1864.7.13.42,
designated by Teugels (1986a: 148).
Clarias ngola Lönnberg & Rendahl, 1920: 170, fig. 1. Type locality: Kingoyi, Lower Congo. Holotype: NRM 9908.
Clarias lualae Lönnberg & Rendahl, 1922: 122, fig. 1. Type locality: Luala River, tributary of the Congo, near
Kinkenge, Congo basin. Syntypes: NRM 39931 (2).
Distribution: central and upper Congo River system and the Luapula system (Teugels, 1986b).
Clarias dussumieri Valenciennes, 1840
Clarias Dussumieri Valenciennes, in Cuvier & Valenciennes, 1840b: 382 (283 in the Strasbourg deluxe edition).
Type locality: Malabar, ... Mahé, ... Pondicherry. Syntypes: MNHN b-0686 (3), MNHN b-0687 (1).
Distribution: South Asia.
Clarias ebriensis Pellegrin, 1920
Clarias ebriensis Pellegrin, 1920: 118. Type locality: Eau stagnante près Bingerville, Côte d’Ivoire. Lectotype:
MNHN 1920-0037, designated by Teugels & Thys van den Audenaerde (1981: 18); lectotype illustrated by Teugels (1986a: 131, fig. 72).
Clarias dahomeyensis Güntert, 1938: 296. Type locality: Porto Novo, Dahomey. Syntypes: NMW 46922 (18),
NMW 469223 (9).
Distribution: Coastal rivers and lagoons from Nigeria to Côte d’Ivoire (Teugels, 1986b).
Clarias engelseni (Johnsen, 1926)
Allabenchelys engelseni Johnsen, 1926: 8, figs. 2–3. Type locality: Aluma, Ramadalla and Yei, Sudan. Syntypes(15): BMNH 1925.12.30.11 (1), ZMUB 4131 a–b (2), ZMUB 4132 a–e (3 + 2 skeletons).
Distribution: Sudan (Teugels, 1986b).
† Clarias falconeri Lydekker, 1886
† Clarias falconeri Lydekker, 1886: 247, pl. 37, fig. 1. Type locality: Siwalik Hills, India. Lower Pliocene. Holotype: BMNH 16402b (middle part of skull).
Distribution: Siwalik Hills, India; Lower Pliocene.
Clarias fuscus (La Cepède, 1803)
Macropteronotus fuscus La Cepède, 1803: 84, 88, pl. 2 (fig. 2). Type locality: China. No types known; based on an
unpublished painting.
Clarias pulicaris Richardson, 1845: 135, pl. 62 (figs. 5–6). Type locality: Canton, China. Holotype: BMNH
1968.3.11.21 (larger of two specimens).
Distribution: Ishigakijama, Japan; Taiwan, and south China (Masuda et al., 1984); northeastern Laos, and northern
Vietnam (Kottelat, 2001a).
Clarias gabonensis Günther, 1867
Clarias Gabonensis Günther, 1867b: 111. Type locality: River Ogowe, Gaboon. Holotype: BMNH 1867.5.22.3.
Clarias dolloi Boulenger, 1896a: 311. Type locality: Monsembe, upper Congo. Syntypes: BMNH 1896.3.9.18–20

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(3).
Clarias congicus Poll, 1942a: 6, figs. 2–3. Type locality: Lac Tumba, Congo belge. Holotype: MRAC 57759.
Distribution: Lower and middle Congo River system, including Lake Tumba and the Pool Malebo; Chiloango system and Ogowe River (Teugels, 1986b).
Remarks: Redescription and comments on status (Teugels, 1982b).
Clarias gariepinus (Burchell, 1822)
Silurus (Heterobranchus) Gariepinus Burchell, 1822: 425 (footnote), figure on p. 445. Type locality: Vaal River at
Smidtsdrift, above confluence with Riet River, Cape Province, South Africa (28º42'10"S, 24º04'29"E), by neotype designation. Neotype: SAIAB 520, illustrated in Skelton & Teugels (1992: fig. 2). Neotype designated byTeugels (1986a: 35), or Skelton & Teugels (1992: 1) [see Remarks, below].
Clarias capensis Valenciennes, in Cuvier & Valenciennes, 1840b: 377 (279 in the Strasbourg deluxe edition). Type
locality: Cap de Bonne-Espérience (in error). Holotype: MNHN a-9431 (dry).
Clarias lazera Valenciennes, in Cuvier & Valenciennes, 1840b: 372 (275 in the Strasbourg deluxe edition). Type
locality: Nil. Syntypes (3, plus one skeleton): MNHN 0000-1578 (1).
Clarias Syriacus Valenciennes, in Cuvier & Valenciennes, 1840b: 375 (275 in the Strasbourg deluxe edition). Type
locality: Syrie. Holotype: MNHN a-9426.
Clarias mossambicus Peters, 1852: 682. Type locality: Tette, Mozambique. Syntypes: possibly FMNH 54274 (1),
ZMB 3068 (1), ZMB 6591 (1), ZMB 6592 (1), ZMB 6593 (1).
Clarias macracanthus Günther, 1864: 16. Type locality: Upper Nile. Syntypes (5, one a skeleton): BMNH
1855.12.26.453 (1), BMNH 1862.6.17.51 (1), BMNH 1866.9.9.67–68 (2).
Clarias orontis Günther, 1864: 15. Type locality: Orontes. Syntype: BMNH 1855.6.25.3. On Silurus anguillaris ofRussell (1794: 217, pl. 8) and Clarias of Gronovius (1763: 100).
Clarias xenodon Günther, 1864: 16. Type locality: Senegal, West Africa. Holotype: BMNH 1863.7. 30.3.
Clarias robecchii Vinciguerra, 1893: 450. Type locality: Uebi Scebeli [Somalia]. Syntypes: MSNG 8138 (2).
Clarias smithii Günther, 1896b: 219, fig. 1. Type locality: middle course of the Shebeli Somaliland. Holotype:
BMNH 1895.12.31.65.
Clarias guentheri Pfeffer, 1896: 28. Type locality: Sansibar. Syntypes: Based on specimens identified as Clarius
gariepinus in Playfair & Günther (1867: 113), including BMNH 1867.3.9.388 (1, stuffed) and BMNH
1867.3.9.581 (stuffed, not found) and literature sources.
Clarias microphthalmus Pfeffer, 1896: 28. Type locality: Mbake, Ugogo, Bubu, Deutsch Ost-Africa. Holotype:
ZMH H477.
Clarias longiceps Boulenger, 1899b: 103, pl. 41 (fig. 1). Type locality: Boma swamps, [Congo basin]. Syntypes:
BMNH 1899.11.27.15–17 (3), MRAC 902–905 (4), MRAC 906–910 (5), MRAC 911–915 (5), MRAC 916–920(5), MRAC 921–922 (2).
Clarias moorii Boulenger, 1901d: 13. Type locality: Lake Albert. Holotype: BMNH 1905.9.7.47.
Clarias tsanensis Boulenger, 1902c: 438. Type locality: Lake Tsana, Abyssinia. Syntypes (5): BMNH
1902.12.13.441–443 (3).
Clarias vinciguerrae Boulenger, 1902c: 438. Type locality: Awhorra, Kassam River, Abyssinia. Holotype: BMNH
1902.12.13.444.
Clarias malaris Nichols & Griscom, 1917: 705, fig. 19. Type locality: Stanleyville, Congo. Holotype: AMNH 6441.
Clarias notozygurus Lönnberg & Rendahl, 1922: 125, fig. 3. Type locality: Lukosi, a tributary to Luala, Lower
Congo. Holotype: NRM 9933.
Clarias depressus Myers, 1925a: 12. Type locality: water hole of a hippopotamus, on the Albano River about 6 km.
west of Addagalla, Abyssinia. Syntypes: AMNH 8204 (2).
Clarias mülleri Pietschmann, 1939: 184. Type locality: Eritrea. Holotype: at Museo di Trieste.
Distribution: Nearly Pan African, as well as Asia Minor (Teugels, 1986b); introduced into Laos (Kottelat, 2001b).
Remarks: Teugels (1986a: 35) cites “Bruton & Teugels (1986)” as source of neotype designation of Silurus (Heterobranchus) gariepinus Burchell. That publication has not been found and may not have preceded Teugels(1986a). Tentatively, Teugels (1986a) is considered the actual source of neotype designation. Seegers (1996:206) was similarly unable to locate the Bruton & Teugels source and, instead, cited Skelton & Teugels (1992) as

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source of the actual neotype designation. See Benech et al. (1993) for characters that distinguish this species
from Clarias anguillaris.
Clarias hilli Fowler, 1936
Clarias hilli Fowler, 1936b: 299, figs. 59–62. Type locality: Saidi's Village, Belgian Congo. Holotype: ANSP
65879.
Clarias gracilis Fowler, 1936b: 303, figs. 75–78. Type locality: Saidi's Village, Belgian Congo. Holotype: ANSP
66098.
Distribution: middle Congo River system and Lake Albert (Teugels, 1986b).
Remarks: Teugels (1986b: 181) gave precedence to Clarias hilli and placed C. gracilis as a synonym, thereby acting
as first reviser.
Clarias insolitus Ng, 2003
Clarias insolitus Ng, 2003h: 2, figs. 1, 2a. Type locality: Borneo: Kalimantan Tengah, Barito River drainage, small
stream flowing into Sungai Rekut (tributary of Sungai Busang) about 1.5 km upstream from the Project Barito
Ulu base camp on Sungai Busang. Holotype: MZB 6112.
Distribution: Barito River basin, southern Borneo (Ng, 2003h).
Clarias intermedius Teugels, Sudarto & Pouyaud, 2001
Clarias intermedius Teugels, Sudarto & Pouyaud, 2001: 86, fig. 5. Type locality: Palangkaraya market, Central Kalimantan, Indonesia. Holotype: MZB 10574.
Distribution: Central Kalimantan, between the Sampit and Barito river basins, Borneo (Teugels et al., 2001).
Clarias jaensis Boulenger, 1909
Clarias jaensis Boulenger, 1909: 188. Type locality: Ja River at Bitye, South Cameroon. Lectotype: BMNH
1909.4.29.77, designated by Teugels (1986a: 55). Illustrated in Boulenger (1911a: 242, fig. 201).
Clarias cameronensis Holly, 1927c: 422, unnumbered figure. Type locality: Nachtigalschnellen des Sanagaflussses.
Holotype: NMW 13374. Considered by Pellegrin (1929b) to be preoccupied by Clarias camerunensis Lönnberg, 1895; replaced by Clarias Hollyi Pellegrin, 1929.
Clarias Hollyi Pellegrin, 1929b: 359 (footnote). Type locality: Nachtigalschnellen des Sanagaflussses. Holotype:
NMW 13374. Replacement for Clarias cameronensis Holly, 1927; considered preoccupied by Clarias camerunensis Lönnberg, 1895.
Clarias (Clarioides) aboinensis Sydenham & Olawoye, 1981: 234, fig. 1. Type locality: Ugbenu market, near
Ologbo, Bendel State, Nigeria. Holotype: BMNH 1980.5.12.1.
Distribution: southeast Nigeria, south Cameroon and Gabon (Teugels, 1986b).
Clarias kapuasensis Sudarto, Teugels & Pouyaud, 2003
Clarias kapuasensis Sudarto, Teugels & Pouyaud, 2003: 156, figs. 2a–b. Type locality: Nanga Pinoh in peat swamp
bordering the river Melawi in the upper part of the Kapuas River, West Kalimantan, Indonesia. Holotype: MZB
10965.
Distribution: Kapuas River basin, Borneo (Sudarto, Teugels & Pouyaud, 2003).
Clarias laeviceps Gill, 1862
Clarias laeviceps Gill, 1862: 139, footnote. Type locality: [probably from Liberia]. Holotype: USNM 4099; holotype illustrated by Teugels (1986a: 143, fig. 79).
Clarias kingsleyae Günther, 1903: 334. Type locality: Odumasi [and] Infoan on the R. Offim, Gold Coast. Syntypes(2): BMNH 1903.4.24.65 (1), BMNH 1903.4.24.66 (1).
Clarias vandenhoutei Poll, 1941b: 137, fig. 3. Type locality: confluence de la rivière Masei et la rivière Bon de
Boan, Côte d’Ivoire. Holotype: MRAC 66360.
Distribution: from Saint Pauls River, Liberia, to Volta River system, Ghana (Teugels, 1986b).
Remarks: Redescribed in Teugels (1981) with comments on the status of the species.
Clarias lamottei Daget & Planquette, 1967
Clarias lamottei Daget & Planquette, 1967: 279, fig. 1. Type locality: Aoué Bla, marigot affluent du Nzi, bassin duBandama près Lamto, Côte d’Ivoire. Holotype: MNHN 1966-1104; holotype illustrated by Teugels (1986a: 20,
fig. 15).
Distribution: Nzi River, tributary of the Bandama, Côte d’Ivoire (Teugels, 1986b).

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Remarks: Considered by Teugels (2003b) as likely to be an intergeneric hybrid between a species of Clarias andHeterobranchus.
Clarias leiacanthus Bleeker, 1851
Clarias leiacanthus Bleeker, 1851e: 430. Type locality: Sambas, in fluviis. Holotype (160 mm TL): possibly BMNH
1863.12.4.85 (124 mm SL).
Clarias liacanthus Günther, 1864: 20. Type locality: Sambas, in fluviis. Holotype: possibly BMNH 1863.12.4.85(124 mm SL). Unjustified emendation of Clarias leiacanthus Bleeker, 1851.
Distribution: Kapuas River (Roberts, 1989a).
Clarias liocephalus Boulenger, 1898
Clarias liocephalus Boulenger, 1898d: 496. Type locality: Lake Tanganyika. Lectotype or Holotype: BMNH
1898.9.9.70. Described in more detail and illustrated in Boulenger (1898f: 24, pl. 7, figs. 2, 2a) with locality as
Kinyamkolo. BMNH 1898.9.9.70 illustrated in Seegers (1996: 209, fig. 149) as holotype. The original description did not indicate how many specimens were included in the concept of this species. Recent authors (e. g.,
Teugels, 1986; Eschmeyer et al., 1998) have interpreted BMNH 1898.9.9.70 as a holotype, but BMNH
1898.9.9.71 was registered as the same species from the same locality. Therefore, the two specimens should
probably have been treated as syntypes. Since it was unclear whether more than one specimen was involved in
the original description, Teugels (1986) interpretation of a holotype made that specimen the lectotype.
Clarias carsonii Boulenger, 1903b: 362. Type locality: Fwambo, British Central Africa. Holotype: BMNH
1894.12.20.20.
Clarias neumanni Hilgendorf, 1905: 409. Type locality: Utim, im Guasso Nyiro in den Nguruman-Salzee fliessend,
unter 36ºÖ L., 1º50'S Br. Holotype: at Nairobi Museum, Kenya, but Seegers (1996: 214, fig. 152, bottom) illustrated BMNH 1905.7.25.49 as the holotype.
Clarias phillipsi Norman, 1925a: 189, pl. 11. Type locality: Lake Chahafi, S.O. Uganda. Holotype: MCZ 31519.
Clarias ornatus Poll, 1943b: 306. Type locality: Bururi, Haute Malagarazi, Lac Tanganika. Holotype: MRAC
56002; illustrated in Poll (1946: 230, fig. 25).
Clarias youngicus Ricardo-Bertram, 1943: 208. Type locality: Lake Young, Shiwa Ngandu, (Bangweulu region).
Syntypes: BMNH 1943.7.27.377–382 (6).
Distribution: Lakes Victoria, Edward, George, Kivu, Tanganyika, Malawi and small lakes of Uganda and Rwanda;
Kagera system, Malagarazi System, Ruzizi River, Tana River, Bangweulu-Moero system (Teugels, 1986b);
Lake Rukwa basin (Seegers, 1996).
Clarias longior Boulenger, 1907
Clarias longior Boulenger, 1907b: 51. Type locality: Kribi and Lobi Rivers, South Cameroon [restricted to Kribi
River, by lectotype designation]. Lectotype: BMNH 1904.10.26.63, designated by Sydenham (1978: 369).
Clarias melanopogon Fowler, 1936b: 301, figs. 71–74. Type locality: Thirty kilometers east of Kribi, Cameroons.
Holotype: ANSP 66075.
Distribution: Kribi, Lobi and Ntem Rivers of South Cameroon (Teugels, 1986b).
Clarias maclareni Trewavas, 1962
Clarias maclareni Trewavas, 1962: 157, fig. 3. Type locality: L. Barombi-ma-Mbu, northwestern Cameroons. Holotype: BMNH 1959.8.18.174.
Distribution: Lake Barombi-ma-Mbu; known only from the type locality (Teugels, 1986b).
Clarias macrocephalus Günther, 1864
Clarias macrocephalus Günther, 1864: 18. Type locality: Siam. Lectotype: BMNH 1862.11.1.216; lectotype designated by Teugels et al. (1999: 294).
Distribution: Mekong and Chao Phraya River basins, and possibly Philippines (Kottelat, 2001b).
Remarks: Redescribed in Teugels et al. (1999: 294).
Clarias macromystax Günther, 1864
Clarias macromystax Günther, 1864: 17. Type locality: Gambia [in error]. Holotype: BMNH 1862.5.20.4.
Distribution: Oueme River, Benin, to the Niger delta and the Niger and Benue Rivers (Teugels, 1986b).
Remarks: Teugels (1986a: 76) indicated that the stated type locality appears to be in error.
Clarias meladerma Bleeker, 1846

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Clarias meladerma Bleeker, 1846a: 178. Type locality: Batavia. Syntypes (size and number not stated): RMNH
13709 (1), RMNH 16413 (1). Illustrated in Bleeker (1862–63: pl. 97 [ Silur. pl. 49], fig. 2) as Clarias melanoderma.
Clarias melasoma Bleeker, 1852b: 427. Type locality: Prabukarta, Borneo austro-orientalis, in flumne Kusan; Palembang, Sumatrae austro-orientalis, in flumine Mussi. Syntypes (2, 170–300 mm TL): RMNH 6802 (1).
Clarias melanoderma Bleeker, 1858b: 341. Type locality: Batavia. Syntypes: RMNH 13709 (1), RMNH 16413 (1),RMNH 6802 (1). Unneeded new name for Clarias meladerma Bleeker, and Clarias melasoma Bleeker.
Distribution: Mekong River basin, and Sumatra, Borneo and Java (Kottelat, 2001b); also the Philippines (Rainboth,
1996).
Clarias microstomus Ng, 2001
Clarias microstomus Ng, 2001c: 158, fig. 1. Type locality: Borneo: Kalimantan Timur, Kayan River drainage,
Ladang near Sungai Pingai, 2º0'9.6"N, 115º9'13.8"E. Holotype: MZB 9336.
Distribution: Mahakam and Kayan River basins, eastern Borneo (Ng, 2001c).
Clarias ngamensis Castelnau, 1861
Ciarias N'gamensis Castelnau, 1861: 63. Type locality: lac N’gami, [now: Lake Ngami, S. W. Africa (17º50'W,
21º00'E), by neotype designation]. Neotype: BMNH 1910.5.31.27, designated by Teugels (1983: 17). Publishedname clearly intended as Clarias N'gamensis.
Clarias Mellandi Boulenger, 1905b: 644. Type locality: Lake Bangwelo. Holotype: BMNH 1905.11.10.7.
Dinotopterus jallae Gilchrist & Thompson, 1917: 556, fig. 165. Type locality: Sheskeke, southern Rhodesia. Holotype: SAM 14297 [now at AMG].
Dinotopteroides prentissgrayi Fowler, 1930: 41, fig. 10. Type locality: Chouzo, Quanza River, Angola. Holotype:
ANSP 51798.
Distribution: Quanza, Cunene, Okavango, Chobe, Zambezi above Victoria Falls, upper Lualaba, Luapula, Pungwe,
Buzi, Save, Limpopo, Incomati, lower Pongolo, and lower Sabi River; Lake Ngami, Moero, Bangweulu, andMalawi (Teugels, 1986b).
Clarias nieuhofii Valenciennes, 1840
Clarias Nieuhofii Valenciennes, in Cuvier & Valenciennes, 1840b: 386 (287 in the Strasbourg deluxe edition). Type
locality: Indies. Holotype: MNHN b-0300.
Clarias pentapterus Bleeker, 1851d: 206. Type locality: Bandjarmassing, in fluviis. Holotype (350 mm TL): Whereabouts unknown.
Clarias gilli Smith & Seale, 1906: 74, figs. a–b. Type locality: Rio Grande, Mindanao, Philippines. Holotype:
USNM 55620.
Distribution: Sumatra, Kalimantan, India, Thailand and the Philippines (Sudarto et al., 2004); Coastal and, probably,
Mekong side of Cardamom Range, Cambodia (Rainboth, 1996).
Clarias nigricans Ng, 2003
Clarias nigricans Ng, 2003g: 393, figs. 1, 2, 3a. Type locality: Borneo: Kalimantan Timur, market in Samarinda.
Holotype: MZB 10705.
Distribution: Mahakam River basin, Borneo (Ng, 2003e).
Clarias nigromarmoratus Poll, 1967
Clarias nigromarmoratus Poll, 1967: 229, fig. 106. Type locality: Riv. Luíta, Angola. Holotype: MD 6484.
Distribution: central Congo River system (Teugels, 1986b).
Clarias olivaceus Fowler, 1904
Chlarias olivaceus Fowler, 1904: 499, pl. 28 (lower figure). Type locality: Padang, Sumatra. Holotype: ANSP
27280.
Distribution: Highland rivers of western Sumatra (Tan & Ng, 2000).
Clarias pachynema Boulenger, 1903
Clarias pachynema Boulenger, 1903c: 438. Type locality: Ja River, southern Cameroon. Lectotype: BMNH
1903.7.28.191, illustrated in, and designated by, Teugels (1980: 686, fig. 61; 214.5 mm TL).
Clarias esamesae Boulenger, 1911e: 372. Type locality: Esamesa, south Cameroon. Holotype: BMNH 1911.5.30.28.
Clarias oxycephalus Boulenger, 1913: 67. Type locality: Nyong River, S. Cameroon. Holotype: BMNH

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1913.10.29.7.
Distribution: Coastal rivers of south Cameroon, Oowe system and central Congo River system (Teugels, 1986b).
Clarias planiceps Ng, 1999
Clarias planiceps Ng, 1999d: 22, figs. 4b, 5, 6. Type locality: Borneo: Sarawak, Belakin area, Ulu Sungai Anap.
Holotype: SMK uncataloged.
Distribution: Upper reaches of the Rajang and Kapuas Rivers, western Borneo, and Kayan River, eastern Borneo(Ng, 1999d).
Clarias platycephalus Boulenger, 1902
Clarias platycephalus Boulenger, 1902d: 35, pl. 10 (fig. 1). Type locality: Monsembé, Haut-Congo. Holotype:
BMNH 1901.12.21.43.
Clarias ekibondoi Fowler, 1936b: 300, figs. 63–66. Type locality: Ekibondo's Village, Belgain Congo. Holotype:
ANSP 65886.
Clarias varispinis Fowler, 1936b: 301, figs. 67–70. Type locality: Nola, Middle Congo. Holotype: ANSP 65887.
Distribution: lower and central Congo River basin (Teugels, 1986b).
Clarias pseudoleiacanthus Sudarto, Teugels & Pouyaud, 2003
Clarias pseudoleiacanthus Sudarto, Teugels & Pouyaud, 2003: 156, figs. 2a–b. Type locality: Satong, peat swamp at
ca. 30 km after Ketapang on the road to Sudadana, Borneo, West Kalimantan, Indonesia. Holotype: MZB
10964.
Distribution: Borneo.
Clarias pseudonieuhofii Sudarto, Teugels & Pouyaud, 2004
Clarias pseudonieuhofii Sudarto, Teugels & Pouyaud, 2004: 15, fig. 3. Type locality: Semitau, near the Danau Sentarum Reserve, upstream Kapuas R. on the road between Sintang and Putussibau, West Kalimantna, Borneo,
Indonesia. Holotype: MZB 10966.
Distribution: Upper Kapuas River, West Kalimantan, Borneo (Sudarto et al., 2004: 16).
Clarias salae Hubrecht, 1881
Clarias salae Hubrecht, 1881: 68. Type locality: St. Paul’s River, Liberia. Lectotype: RMNH 5373 (smallest specimen); lectotype illustrated in, and designated by, Teugels (1986a: 103, fig. 54).
Distribution: Konkoure system, Guinea, to the Cavally River, Côte d’Ivoire (Teugels, 1986b).
Clarias stappersii Boulenger, 1915
Clarias Stappersii Boulenger, 1915: 168. Type locality: Ruisseau affluent de la Lukinda, Tanganika-Moero. Holotype: MRAC 14139.
Distribution: Luapula-Moero system, Kafue River, upper Zambezi River and Cunene River (Teugels, 1986b).
Clarias submarginatus Peters, 1882
Clarias submarginatus Peters, 1882: 74. Type locality: Westafrica (Tooxlong River). Lectotype: ZMB 11895 (largest specimen), designated by Sydenham (1978: 358); lectotype illustrated by Teugels (1986a: 107, fig. 56).
Clarias hollyi Fowler, 1936b: 304, figs. 79–82. Type locality: Thirty kilometers east of Kribi, Cameroons. Holotype:
ANSP 66118. Preoccupied by Clarias hollyi Pellegrin, 1929; apparently not replaced.
Distribution: Kribi and Lobi Rivers, south Cameroon (Teugels, 1986b).
Clarias sulcatus Ng, 2004
Clarias sulcatus Ng, 2004d: 290, fig. 1. Type locality: Malaysia: Terengganu: Pulau Redang, stream on east slope ofwest ridge. Holotype: ZRC 22666.
Distribution: Rendang Island, off northeastern Malay Peninsula (Ng, 2004d).
Clarias teijsmanni Bleeker, 1857
Clarias Teijsmanni Bleeker, 1857a: 344. Type locality: Tjikoppo, provinciae Buitenzorg, 900 metr. p.m. supra mare[Java]. Holotype (110 mm TL): possibly AMS B.8031 (1), or BMNH 1863.12.4.62 (1).
Clarias pulcher Popta, 1904: 179. Type locality: le Howong, Bornéo central. Holotype RMNH 7542; described in
more detail, with illustration of holotype, in Popta (1906: pl. 1, fig. 2).
Distribution: Kapuas River, Borneo (Roberts, 1989a), and Java.
Clarias theodorae Weber, 1897
Clarias theodorae Weber, 1897: 150. Type locality: Natal: Umhloti-Fluss, Süd-Afrika. Holotype: ZMA 100647;

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holotype illustrated by Teugels (1986a: 111, fig. 58).
Clarias amplexicauda Boulenger, 1902d: 36, pl. 10 (fig. 2). Type locality: Banzyville, [Haut Ubangi]. Holotype:
MRAC 1158.
Clarias Fouloni Boulenger, 1905b: 644. Type locality: Lake Bangwelo. Holotype: BMNH 1905.11.10.8.
Clarias macrurus Boulenger, 1915: 168. Type locality: Ruisseau affluent de la Lukinda, Tanganika-Moero. Holotype: MRAC 14256.
Distribution: Zambezi, Kafue, Shire, upper Congo, Chobe, Okavango, Cunene, Pungwe, Sabi, Lundi, Limpopo,
Incomati, Pongola, Umgeni rivers; Lakes Tanganyika, Bangweulu Kobo, Niumbe, Mweru, Malawi, and Sibaya(Teugels, 1986b).
Clarias werneri Boulenger, 1906
Clarias Werneri Boulenger, 1906c: 569. Type locality: Gondokoro, and Lake Victoria (at Bunjako, Buganga, andMunyonyo), Uganda [restricted to Buganga, by lectotype designation]. Lectotype: BMNH 1906.5.30.165; designated by Teugels (1986a: 126).
Distribution: Lakes Victoria, Kioga, Edward, and Tanganyika; upper Nile River (Teugels, 1986b).
Species inquirendae, Clarias
Clarias abbreviatus Valenciennes, in Cuvier & Valenciennes, 1840b: 386 (286 in the Strasbourg deluxe edition). Type
locality: Macao. Syntypes: MNHN a-9592 (2).
Clarias Assamensis Day, 1877 (in Day, 1875–78): 485. Type locality: Upper and lower Assam. Syntypes: Possibly AMS
B.7485 (1), ZSI F1268 (1, lost).
Cossyphus ater M’Clelland, 1844a: 403, pl. 22 (fig. 3). Type locality: Chusan, China. Holotype: Whereabouts unknown.
Macropteronotus charmuth La Cepède, 1803: 84, 85. Type locality: Nile R. Types: based on numerous literature sources.
Possibly a synonym of Clarias anguillaris (Linnaeus, 1758) or Clarias gariepinus (Burchell, 1822) (Teugels, 1986a:
35; Teugels, 1986b: 71, 79; Seegers, 1996: 205), depending on the selection of a single primary type specimen.
Macropteronotus hexacicinnus La Cepède, 1803: 84, 88, pl. 2 (fig. 3). Type locality: China. No types known; based on an
unpublished painting.
Clarias marpus Valenciennes, in Cuvier & Valenciennes, 1840b: 378 (280 in the Strasbourg deluxe edition). Type locality: Calcutta,.... Pondichéry ...[Vizagapatam], .... l’Irouaddi.... . Syntypes: MNHN b-0685 (2).
Clarias nebulosus Deraniyagala, 1958: 133, pl. 2 (figs. 3–5, 7). Type locality: Kaluganga at Ratnapura, Ceylon. Holotype: NMSL FF721 (lost).
Clarias sauteri Regan, 1908b: 151. Type locality: Kagi, Formosa. Syntypes: BMNH 1908.5.27.16–22 (7).
Clarias tenuis Güntert, 1938: 302. Type locality: China. Syntypes (2): NMW 47074 (2).
Clarias thienemanni Ahl, 1934: 235. Type locality: Porsea, Toba-See, Sumatra. Syntypes (4): ZMB 20931, ZMB 20934,
ZMB 20944.

DINOTOPTERUS Boulenger, 1906
Dinotopterus Boulenger, 1906f: 550. Type species: Dinotopterus cunningtoni Boulenger, 1906. Type by monotypy.
Gender: Masculine.
Revision: Greenwood (1961); sensu lato, including species now in Bathyclarias.
Dinotopterus atribranchus Greenwood, 1961
Dinotopterus atribranchus Greenwood, 1961: 228. Type locality: Lake Nyasa. Holotype: BMNH 1960.2.29.17.
Distribution: Lake Malawi (Teugels, 1986b).
Dinotopterus cunningtoni Boulenger, 1906
Dinotopterus cunningtoni Boulenger, 1906f: 550, pl. 31. Type locality: Mpala, Lake Tanganyika. Syntypes: BMNH
1906.9.8.62 (1), BMNH 1906.9.8.63 (1).
Distribution: Lake Tanganyika (Teugels, 1986b).
Dinotopterus filicibarbis (Jackson, 1959)
Bathyclarias filicibarbis Jackson, 1959: 120, fig. 5. Type locality: Nkata Bay, Lake Nyasa. Holotype: BMNH

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1955.6.14.1.
Distribution: Lake Malawi (Teugels, 1986b).
Dinotopterus foveolatus (Jackson, 1955)
Clarias foveolatus Jackson, 1955: 681, unnumbered figure. Type locality: Nkata Bay, Lake Nyasa. Holotype:
BMNH 1955.6.15.1.
Distribution: Lake Malawi (Teugels, 1986b).
Dinotopterus gigas (Jackson, 1959)
Bathyclarias gigas Jackson, 1959: 125, fig. 7. Type locality: [Nkata Bay] Lake Nyasa. Holotype: BMNH
1960.2.29.35 (head and skin).
Distribution: Lake Malawi (Teugels, 1986b).
Dinotopterus jacksoni Greenwood, 1961
Dinotopterus jacksoni Greenwood, 1961: 222. Type locality: Nkata Bay, Lake Nyasa. Holotype: BMNH
1960.2.29.26.
Distribution: Lake Malawi (Teugels, 1986b).
Dinotopterus loweae (Jackson, 1959)
Bathyclarias loweae Jackson, 1959: 114, fig. 1. Type locality: Nkata Bay, Lake Nyasa. Holotype: BMNH
1960.2.29.3.
Distribution: Lake Malawi (Teugels, 1986b).
Dinotopterus nyasensis (Worthington, 1933)
Clarias nyasensis Worthington, 1933b: 308, fig. 9. Type locality: Lake Nyasa. Lectotype: BMNH 1932.11.15.584,
designated by Greenwood (1961: 221).
Distribution: Lake Malawi (Teugels, 1986b).
DOLICHALLABES Poll, 1942
Dolichallabes Poll, 1942c: 95. Type species: Dolichallabes microphthalmus Poll, 1942. Type by monotypy. Gender:
Masculine.
Dolichallabes microphthalmus Poll, 1942
Dolichallabes microphthalmus Poll, 1942c: 97, figs. a, d. Type locality: Kunungu, Congo belge. Holotype: MRAC
44655.
Distribution: Congo River system (Teugels, 1986b).
Remarks: Redescribed in Devaere et al. (2004).
ENCHELOCLARIAS Herre & Myers, 1937
Encheloclarias Herre & Myers, 1937: 66. Type species: Heterobranchus tapeinopterus Bleeker, 1852. Type by original designation. Gender: Masculine.
Revision: Ng & Lim (1993).
Encheloclarias baculum Ng & Lim, 1993
Encheloclarias baculum Ng & Lim, 1993: 27, fig. 5. Type locality: Probably Borneo: Sambas, Kalimantan Barat.
Holotype: BMNH 1863.12.4.162.
Distribution: Probably Sambas, Borneo; known only from holotype (Ng & Lim, 1993).
Remarks: Register number of holotype of Encheloclarias baculum incorrectly reported as BMNH 1863.12.11.162 in
Ng & Lim (1993) and Eschmeyer et al. (1998).
Encheloclarias curtisoma Ng & Lim, 1993
Encheloclarias curtisoma Ng & Lim, 1993: 32, figs. 9–10. Type locality: Peninsular Malaysia: Selangor, northSelangor peat swamp forest, blackwater stream at 39 km milestone on road to Tanjong Malim from Sungei
Basar. Holotype: ZRC 14886.
Distribution: streams north of Selangor peat forest, Selangor, Malaysia (Ng & Lim, 1993).
Encheloclarias kelioides Ng & Lim, 1993
Encheloclarias kelioides Ng & Lim, 1993: 30, figs. 7–8. Type locality: Peninsular Malaysia: Pahang, south ofPekan, blackwater stream across road, at 69 km stone on road from Mersing to Pekan. Holotype: ZRC 29401.

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Distribution: Pahang, and Mawai, Johor, southeastern peninsular Malaysia (Ng & Lim, 1993).
Encheloclarias prolatus Ng & Lim, 1993
Encheloclarias prolatus Ng & Lim, 1993: 29, fig. 6. Type locality: Borneo: Sarawak, Kuching. Holotype: SMK
5686.
Distribution: Western Sarawak, Borneo (Ng & Lim, 1993).
Encheloclarias tapeinopterus (Bleeker, 1852)
Heterobranchus tapeinopterus Bleeker, 1852e: 732. Type locality: Provincia Toboali, in fluviis [Sumatra, Indonesia]. Holotype (124 mm TL): RMNH 6806; holotype illustrated in Ng & Lim (1993: fig. 3).
Distribution: Banka, Sumatra (Ng & Lim, 1993).
Encheloclarias velatus Ng & Tan, 2000
Encheloclarias velatus Ng & Tan, 2000: 536, fig. 1. Type locality: Sumatra: Jambi, Angso Duo market. Holotype:
MZB 9335.
Distribution: Batang Hari drainage, central Sumatra, and Binton (Ng & Tan, 2000).
GYMNALLABES Günther, 1867
Gymnallabes Günther, 1867b: 111. Type species: Gymnallabes typus Günther, 1867. Type by original designation.
Gender: Feminine.
Gymnallabes alvarezi Roman, 1970
Gymnallabes alvarezi Roman, 1970: 5, fig. 1. Type locality: Rio Kie, circa de Ebebiyin, Rio Muni, Guinea Ecuatorial. Holotype: at Inst. Biol. Aplicada, Barcelona.
Distribution: Kie and Muni Rivers, Equatorial Guinea (Teugels, 1986b).
Gymnallabes nops Roberts & Stewart, 1976
Gymnallabes nops Roberts & Stewart, 1976: 277, pl. 7 (fig. e), pl. 8 (fig. a). Type locality: Near Tadi, about 50 km
downstream from Luozi. Lat. 5º 14'S, long. 13º56'E; lower Zaïre or Congo River. Holotype: MCZ 50298; skeleton of holotype illustrated in Devaere et al. (2005).
Distribution: Lower Congo River (Teugels, 1986b).
Remarks: Redescribed in Devaere et al. (2005).
Gymnallabes typus Günther, 1867
Gymnallabes typus Günther, 1867b: 111, pl. 2 (fig. a). Type locality: West Africa, probably Old Calabar. Syntypes:
BMNH 1866.12.4.1–2 (2).
Gymnallabes typus heterocercalis Lönnberg, 1895: 184. Type locality: Cameroons. Syntypes (2): ZMUU 566. Originally as Gymnallabes typus forma heterocercalis.
Clariallabes longicaudatus Pappenheim, 1911: 519, fig. 3. Type locality: in der Mabelle, Süd-Kamerun. Holotype:
ZMB 18401.
Distribution: Niger River to Cameroon (Teugels, 1986b).
HETEROBRANCHUS Geoffroy Saint-Hilaire, 1809
Heterobranchus Geoffroy Saint-Hilaire, 1809: pls. 16–17. Type species: Heterobranchus bidorsalis Geoffroy SaintHilaire, 1809. Type by monotypy. Gender: Masculine.
Revision: Teugels, Denayer & Legendre (1990).
Monophyly: Agnèse & Teugels (2001b).
Heterobranchus bidorsalis Geoffroy Saint-Hilaire, 1809
Heterobranchus bidorsalis Geoffroy Saint-Hilaire, 1809: pls. 16–17. Type locality: Nil. Holotype: at MNHN (skeleton).
Heterobranchus Geoffroyi Valenciennes, in Cuvier & Valenciennes, 1840b: 392 (291 in the Strasbourg deluxe edition). Type locality: Nil. Holotype: at MNHN (skeleton). New name for Heterobranchus bidorsalis Geoffroy
Saint-Hilaire, 1809.
Heterobranchus Senegalensis Valenciennes, in Cuvier & Valenciennes, 1840b: 397 (295 in the Strasbourg deluxe
edition). Type locality: Sénégal. Holotype: at MNHN (skull).
Heterobranchus intermedius Günther, 1864: 22. Type locality: Khartoum, Upper Nile. Syntypes (3, one a skeleton):

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Distribution: Nile, Niger, Upper Volta, Guinea and Senegal Rivers, Chad basin (Teugels, 1986b).
Heterobranchus boulengeri Pellegrin, 1922
Heterobranchus Boulengeri Pellegrin, 1922d: 274. Type locality: Rivière Lukonzolwa. Syntypes (2): MRAC 14763(1), MRAC 14764 (1).
Distribution: Lake Moero, Lukonzolwa River and upper Congo River, Zimbabwe (Teugels, 1986b).
Heterobranchus isopterus Bleeker, 1863
Heterobranchus isopterus Bleeker, 1863a: 108, pl. 22 (fig. 1). Type locality: Guinea [Rio Boutry]. Holotype: RMNH
2977, or ZMB 6590.
Heterobranchus macronema Bleeker, 1863a: 109, pls. 21 (fig. 1), 22 (fig. 2). Type locality: Guinea [Dabo-Crom].
Syntypes (4): RMNH 2978 (3).
Distribution: West Africa, from Senegal to Ghana (Teugels, 1986b).
Heterobranchus longifilis Valenciennes, 1840
Heterobranchus longifilis Valenciennes, in Cuvier & Valenciennes, 1840b: 394 (293 in the Strasbourg deluxe edition), pl. 447. Type locality: Nil (Egypte). Holotype: MNHN b-0273.
Heterobranchus laticeps Peters, 1852: 682. Type locality: Tette, Boror (Mossambique). Syntypes: ZMB 3071 or
3072 (2), possibly ZMB 4741 (1), ZMB 6590 (1). Illustrated in Peters (1868c: 37, pl. 7, fig. 1).
Heterobranchus platycephalus Nichols & La Monte, 1934: 2, fig. 2. Type locality: Luluabourg, Kasai District, Belgian Congo. Holotype: AMNH 12356.
Clarias loangwensis Worthington, 1933a: 46, unnumbered figure. Type locality: Loangwa River, Zambezi system.
Holotype: BMNH 1932.12.16.501.
Distribution: Nile, Niger, Senegal, Congo and upper and middle Zambezi Rivers (Teugels, 1986b).
† Heterobranchus palaeindicus Lydekker, 1886
† Heterobranchus palæindicus Lydekker, 1886: 248, pl. 36, fig. 4. Type locality: Siwalik Hills, India. Lower
Pliocene. Holotype: BMNH (paleo) 16402c (middle part of skull).
Distribution: Siwalik Hills, India; Lower Pliocene.
HETEROPNEUSTES Müller, 1840
Heteropneustes Müller, 1840: 243. Type species: Silurus fossilis Bloch, 1794. Type by monotypy. Gender: Masculine.
Saccobranchus Valenciennes, in Cuvier & Valenciennes, 1840b: 399 (296 in Strasbourg deluxe edition). Type species: Silurus singio Hamilton, 1822. Type by monotypy. Gender: Masculine.
Clarisilurus Fowler, 1937: 133. Type species: Clarisilurus kemratensis Fowler, 1937. Type by original designation.
Gender: Masculine.
Phylogenetic position: Diogo et al. (2003a).
Review: Hora (1936a).
Remarks: The relative dates of publication of the names Heteropneustes and Saccobranchus have not been clearly
established and, therefore, priority of one name over the other is unknown. Herein, Heteropneustes is treated asvalid because of its widespread usage in recent times.
Heteropneustes fossilis (Bloch, 1794)
Silurus fossilis Bloch, 1794: 46, pl. 370 (fig. 2). Type locality: Tranquebar. Lectotype: ZMB 3074, designated by,
and illustrated in, Paepke (1999: 87, pl. 25, fig. 3).
Silurus singio Hamilton, 1822: 147, 374, pl. 37 (fig. 46). Type locality: Bengal. No types known.
Silurus laticeps Swainson, 1838: 345, fig. 84. Type locality: India. No types known. See also, Swainson (1839: 393).
Silurus biserratus Swainson, 1839: 306. Type locality: Bengal. No types known. Based on “Ham. pl. 37. f 46 ” [=Hamilton (1822: pl. 37, fig. 46)]. Unneeded new name for Silurus singio Hamilton, 1822.
Saccobranchus microcephalus Günther, 1864: 31. Type locality: Ceylon. Holotype: BMNH 1864.3.18.1.
Saccobranchus microps Günther, 1864: 31. Type locality: Ceylon. Syntypes: BMNH 1859.5.31.9–11 (3).
Distribution: Widely distributed in South Asia.
Remarks: Synonymy of Saccobranchus microps based on Pethiyagoda & Bahir (1998).

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Heteropneustes kemratensis (Fowler, 1937)
Clarisilurus kemratensis Fowler, 1937: 133, figs. 5–7. Type locality: Kemrat, Siam. Holotype: ANSP 67880.
Distribution: Mekong, Chao Phraya and Tapi River basins (Kottelat, 2001b).
Species inquirenda, Heteropneustes
Heteropneustes longipectoralis Rema Devi & Raghunathan, 1999: 109, pl. 2 (figs. 1–5). Type locality: ThirumurthiDam, Anamalai Hills, Western Ghats, Tamil Nadu, India. Holotype: ZSI/SRS F.4935.
HORAGLANIS Menon, 1950
Horaglanis Menon, 1950: 60. Type species: Horaglanis krishnai Menon, 1950. Type by original designation. Gender: Masculine.
Horaglanis alikunhii Subhash Babu & Nayer, 2004
Horaglanis alikunhii Subhash Babu & Nayer, 2004: 296, fig. 1. Type locality: Parappukara (10º23'N, 76º15'E), Trichur district, Kerala [India]; from a well. Holotype: ZSI Calicut. Regn. V/F 13137.
Distribution: Trichur District, Kerala, India (Subhash Babu & Nayer, 2004).
Horaglanis krishnai Menon, 1950
Horaglanis krishnai Menon, 1950: 64, pl. 1 (figs. 1–3). Type locality: Well at Kottayam, Kerala [India]. Holotype:ZSI F313/2.
Distribution: Two separate locations in Kottayam, Kerala State, India (Subhash Babu & Nayer, 2004).
PLATYALLABES Poll, 1977
Platyallabes Poll, 1977: 129. Type species: Gymnallabes tihoni Poll, 1944. Type by original designation. Gender:
Feminine.
Platyallabes tihoni (Poll, 1944)
Gymnallabes tihoni Poll, 1944b: 79, fig. on p. 81. Type locality: Poste de peche de Kingabwa, Stanley-Pool. Neotype: MRAC 103307, designated by Derijst (1991: 43).
Distribution: Malebo Pool (Teugels, 1986b).
Remarks: Described initially on an illustration. Redescribed by Poll (1957: 240) with a new “holotype” which Derijst (1991: 43) regarded as, and effectively designated as, the neotype. Species redescribed in Devaere et al.
(2005).
PLATYCLARIAS Poll, 1977
Platyclarias Poll, 1977: 136. Type species: Platyclarias machadoi Poll, 1977. Type by original designation. Gender:
Masculine.
Platyclarias machadoi Poll, 1977
Platyclarias machadoi Poll, 1977: 141, figs. 7–8. Type locality: Haut Cuango, Cafunfo, rivière Borio, Angola. Holotype: MRAC 78-6-P-1345.
Distribution: Upper Cuango River (Teugels, 1986b).
TANGANIKALLABES Poll, 1943
Tanganikallabes Poll, 1943a: 127. Type species: Tanganikallabes mortiauxi Poll, 1943. Type by monotypy. Gender:
Feminine.
Tanganikallabes mortiauxi Poll, 1943
Tanganikallabes mortiauxi Poll, 1943a: 131, figs. 1–4. Type locality: lac Tanganyika, Albertville. Holotype: MRAC
63731; also illustrated in Poll (1946: 234, figs. 27, 28).
Distribution: Lake Tanganyika (Teugels, 1986b).
UEGITGLANIS Gianferrari, 1923
Uegitglanis Gianferrari, 1923: 1. Type species: Uegitglanis zammaranoi Gianferrari, 1923. Type by original desig-

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nation. Gender: Masculine.
Phylogeny: David (1936a).
Uegitglanis zammaranoi Gianferrari, 1923
Uegitglanis Zammaranoi Gianferrari, 1923: 1, pl. 1. Type locality: Pozzi di Uegit (El Uegit) Somalia italiana. Holotype: MSNM 37.
Distribution: Caves near the Uegit and the Uebi Scebeli rivers, Somalia (Teugels, 1986b).
Remarks: Systematic position discussed in David (1936a: 369).
XENOCLARIAS Greenwood, 1958
Xenoclarias Greenwood, 1958: 321. Type species: Clarias eupogon Norman, 1928. Type by original designation.
Gender: Masculine.
Xenoclarias eupogon (Norman, 1928)
Clarias eupogon Norman, 1928a: 105. Type locality: Lake Victoria (0º22'S, 34º14½'E). Holotype: BMNH
1928.5.24.57.
Distribution: Lake Victoria, in deep waters (Teugels, 1986b).
Xenoclarias holobranchus Greenwood, 1958
Xenoclarias holobranchus Greenwood, 1958: 322. Type locality: off the south-western tip of Ukerewe Island, Lake
Victoria. Holotype: BMNH 1958.7.8.1.
Distribution: Lake Victoria (Teugels, 1986b).

CLAROTEIDAE Bleeker, 1862
Claroteini Bleeker, 1862 (in Bleeker, 1862–63): 4. Type genus: Clarotes Kner, 1855.
Chrysichthyinae Regan, 1911: 561. Type genus: Chrysichthys Bleeker, 1858.
Auchenoglanidinae Jayaram, 1966b: 1119. Type genus: Auchenoglanis Günther, 1865.
Gephyroglanidini Jayaram, 1966b: 1071, 1099. Type genus: Gephyroglanis Boulenger, 1899.
Chrysichthyini Jayaram, 1966b: 1071.Type genus: Chrysichthys Bleeker, 1858; preoccupied by Regan (1911).
Taxonomic summaries: Risch (1986, as Bagridae, in part).
Reviews: Jayaram (1955a, 1966b, African genera); Risch (1992b, West Africa); Bailey & Stewart (1984, Lake Tanganyika).
Phylogeny: Jayaram (1976); Jayaram & Singh (1982); Mo (1991); Ng (2003j).
Key to genera: Poll & Gosse (1994).
17 genera, 83 species; including 1 fossil genus and 3 fossil species.
AMARGINOPS Nichols & Griscom, 1917
Amarginops Nichols & Griscom, 1917: 713. Type species: Amarginops platus Nichols & Griscom, 1917. Type bymonotypy. Gender: Masculine.
Amarginops hildae (Bell-Cross, 1973)
Chrysichthys hildae Bell-Cross, 1973: 7, fig. 1. Type locality: Buzi River, Mozambique, at approximately 20º22'S,
30º20'E. Holotype: QVMS 1827.
Distribution: Lower reaches of Buzi River, Mozambique (Skelton, 1993).
Remarks: Also treated in recent literature as Chrysichthys hildae.
Amarginops platus Nichols & Griscom, 1917
Amarginops platus Nichols & Griscom, 1917: 713, fig. 22–23. Type locality: Stanleyville, Congo. Holotype: AMNH
6528.
Distribution: Kisangani, Congo; known only from types (Risch, 1986).
ANASPIDOGLANIS Teugels, Risch, de Vos & Thys van den Audenaerde, 1991
Anaspidoglanis Teugels, Risch, de Vos & Thys van den Audenaerde, 1991: 513. Type species: Auchenoglanis mac-

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rostoma Pellegrin, 1909. Type by original designation. Gender: Masculine.
Revision: Mbega & Teugels (1998: 233).
Key: Geerinckx et al. (2003).
Anaspidoglanis akiri (Risch, 1987)
Auchenoglanis akiri Risch, 1987: 33, fig. 4. Type locality: Umayara Mba, Etche, Rivers State, Nigeria (5º12'N,
7º06'E). Holotype: MRAC 84-15-P-9; holotype illustrated in Geerinckx et al. (2003: 24, fig. 8c).
Distribution: Niger River delta, New Calabar and Bony Rivers, Nigeria (Geerinckx et al. 2003).
Remarks: Redescribed and assigned to Anaspidoglanis in Geerinckx et al. (2003).
Anaspidoglanis boutchangai (Thys van den Audenaerde, 1965)
Parauchenoglanis boutchangai Thys van den Audenaerde, 1965b: 347, fig. 1. Type locality: Rapides de la Ngounié
ou de la Louetsi à Lelamba, 2º12'S, 11º30'E [Gabon]. Holotype: MRAC 153160.
Distribution: Ngounié or Louetsi River, Gabon; known only from the type locality (Risch, 1986).
Anaspidoglanis macrostomus (Pellegrin, 1909)
Auchenoglanis macrostoma Pellegrin, 1909a: 67. Type locality: Ngomo (Ogôoué) [Gabon]. Holotype: MNHN
1909-0017. Also described as new in Pellegrin (1909d: 48, figs. 1a, 1b). Name spelled Auchenauglanis macrostom on p. 66.
Parauchenoglanis ansorgii Boulenger, 1912b: 20, pl. 22 (fig. 3). Type locality: N'Kutu sur les chutes du Loango.
Holotype: BMNH 1912.4.1.447.
Distribution: S. Cameroon, Gabon, and Congo River system (Risch, 1986).
AUCHENOGLANIS Günther, 1865
Auchenaspis Bleeker, 1858b: 198, 205, 208. Type species: Pimelodus biscutatus Geoffroy St. Hilaire, 1809. Type by
subsequent designation by Bleeker, 1862 (in Bleeker, 1862–63: 12). Gender: Feminine. Preoccupied by †Auchenaspis Egerton, 1857, in fossil fishes; replaced by Auchenoglanis Günther, 1865.
Auchenoglanis Günther, 1865b: 165. Type species: Pimelodus biscutatus Geoffroy St. Hilaire, 1809. Type by beinga replacement name. Gender: Masculine. Replacement for Auchenaspis Bleeker, 1858, preoccupied by † Auchenaspis Egerton, 1857, in fossil fishes.
Oxyglanis Vinciguerra, 1898: 249. Type species: Oxyglanis sacchii Vinciguerra, 1898. Type by monotypy. Gender:
Masculine.
Remarks: Included species and synonymy follows Teugels et al. (1991).
Auchenoglanis biscutatus (Geoffroy Saint-Hilaire, 1809)
Pimelodus biscutatus Geoffroy Saint-Hilaire, 1809: pl. 14 (fig. 1). Type locality: Nil. Holotype: MNHN a-8969.
Distribution: Nile, Niger, and Senegal rivers, and Lake Chad basin (Risch, 1986).
Auchenoglanis occidentalis (Valenciennes, 1840)
Pimelodus occidentalis Valenciennes, in Cuvier & Valenciennes, 1840b: 203 (150 in Strasbourg deluxe edition).
Type locality: Sénégal. Holotype: MNHN a-8971.
Oxyglanis Sacchii Vinciguerra, 1898: 250. Type locality: fiume Omo a Murzj. Holotype: MSNG 14506.
Auchenoglanis occidentalis tanganicanus Boulenger, 1906f: 553, pl. 33. Type locality: Kalambo,... the middle of thelake [Lake Tanganyika] and .... Mtomdwe Bay. Syntypes (4): BMNH 1906.9.6.37–38 (2), BMNH 1906.9.6.39(1), BMNH 1906.9.8.69 (1). Originally as Auchenoglanis occidentalis var. tanganicanus.
Auchenoglanis occidentalis tchadiensis Pellegrin, 1909b: 244. Type locality: Lac Chad à Bol. Syntypes: MNHN
1909-0425 (3), MNHN 1909-0426 (4). Originally as Auchenoglanis occidentalis var. tchadiensis.
Auchenoglanis acuticeps Pappenheim, in Pappenheim & Boulenger, 1914: 249, pl. 5 (fig. 1). Type locality: Albertsee. Syntypes (7): ZMB 19094 (1), ZMB 21317 (3).
Auchenoglanis Wittei Giltay, 1930a: 92, figs. 1–2. Type locality: Stanleyville. Holotype: MRAC 19802.
Distribution: East African lakes; Lake Chad; Nile, Congo, Omo and Giuba rivers (Risch, 1986).
BATHYBAGRUS Bailey & Stewart, 1984
Bathybagrus Bailey & Stewart, 1984: 8. Type species: Bathybagrus tetranema Bailey & Stewart, 1984. Type byoriginal designation. Gender: Masculine.

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Bathybagrus tetranema Bailey & Stewart, 1984
Bathybagrus tetranema Bailey & Stewart, 1984: 11, figs. 1–4. Type locality: Zambian waters of Lake Tanganyika.
Holotype: UMMZ 196110.
Distribution: Lake Tanganyika, Zambian portion (Risch, 1986).
CHRYSICHTHYS Bleeker, 1858
Chrysichthys Bleeker, 1858b: 60, 65. Type species: Pimelodus auratus Geoffroy St. Hilaire, 1809. Type by subsequent designation by Bleeker (1862–63: 9). Gender: Masculine.
Melanodactylus Bleeker, 1858b: 60, 65. Type species: Arius acutivelis Valenciennes, 1840. Type by subsequent designation by Bleeker (1862–63: 9). Gender: Masculine.
Chrysobagrus Boulenger, 1899a: 40. Type species: Chrysobagrus brevibarbis Boulenger, 1899. Type by subsequent
designation, apparently by Jordan (1920: 485); see Eschmeyer et al. (1998) for details. Gender: Masculine.
Key to species: Jayaram (1966b).
Chrysichthys aluuensis Risch, 1985
Chrysichthys aluuensis Risch, 1985b: 185, figs. 2–3. Type locality: Omuehuechi-Aluu, New Calabar River, Rivers
State, Nigeria. Holotype: MRAC 84-20-P-1.
Distribution: New Calabar River; known only from type locality (Risch, 1986).
Chrysichthys ansorgii Boulenger, 1910
Chrysichthys ansorgii Boulenger, 1910: 558. Type locality: Quanza River at Dondo and from the Bengo River. Syntypes (numerous): ANSP 37906 (1), BMNH 1911.6.1.116–124 (9), BMNH 1911.6.1.126 (1), USNM 86636 (1),
ZMB 18228 (1).
Distribution: Cuanza and Bengo Rivers, Angola (Risch, 1986).
Chrysichthys auratus (Geoffroy Saint-Hilaire, 1809)
Pimelodus auratus Geoffroy Saint-Hilaire, 1809: pl. 14 (figs. 3–4). Type locality: Nil. Holotype: at MNHN. Nameavailable from caption to plate, described as Porcus auratus in Geoffroy Saint-Hilaire (1827: 302).
Bagrus capito Valenciennes, in Cuvier & Valenciennes, 1840a: 430 (319 of Strasbourg deluxe edition). Type locality: Nil. Syntypes: MNHN b-0605 (1), MNHN b-0606 (1).
Chrysichthys macrops Günther, 1864: 71. Type locality: Nile, Chartoum. Syntypes (7): BMNH 1860.11.9.125 (1),
BMNH 1862.6.17.60–64 (5).
Gephyroglanis tilhoi Pellegrin, 1909b: 243. Type locality: Bol, Lac Chad. Holotype: MNHN 1909-0424.
Chrysichthys filamentosus Boulenger, 1912b: 19, pl. 18 (fig. 2). Type locality: Chiloango ville, Loango à N'Kutu,
Luali à Lundo et Buco Zau, Lebuzi à Kuka Muno. Syntypes: BMNH 1912.4.1.440 (1), BMNH 1912.4.1.441(1), BMNH 1912.4.1.442–443 (2), BMNH 1912.4.1.444 (1), BMNH 1912.4.1.445–446 (2), MRAC 1668–69(2), MRAC 1670 (1), ZMB 8820 (1).
Chrysichthys longifilis Pfaff, 1933: 289, figs. 5–6. Type locality: Kabara. Holotype (29.2 cm): ZMUC P 28668.
Distribution: Nile, Niger, Volta, Senegal rivers, Lake Chad basin, and coastal rivers of western Africa, from Liberia
to Angola (Risch, 1986).
Remarks: The name Bagrus auratus in Hyrtl (1859: 17) is sometimes treated as an available name but is more likely
intended as a reference to this species.
Chrysichthys bocagii Boulenger, 1910
Chrysichthys bocagii Boulenger, 1910: 557. Type locality: Bengo River at Dondo, Angola. Syntypes (2): BMNH
1911.6.1.114–115 (2).
Distribution: Bengo River, Angola (Risch, 1986).
Chrysichthys brachynema Boulenger, 1900
Chrysichthys brachynema Boulenger, 1900c: 480. Type locality: Kalambo [Lake Tanganyika]. Syntype or lectotype:
BMNH 1906.9.6.33 [see Remarks].
Chrysichthys myriodon Boulenger, 1900a: 139, pl. 51. Type locality: Albertville, Tembwi [Lake Tanganyika]. Syntypes: BMNH 1906.9.6.27 (1), BMNH 1906.9.6.28–31 (4), BMNH 1906.9.6.32 (1, skeleton), MRAC 325 (1).
Distribution: Lake Tanganyika (Risch, 1986).
Remarks: Risch (1986: 15) and Eschmeyer et al. (1998) stated that a lectotype was designated for Chrysichthys

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brachynema Boulenger, but neither indicated the source of that designation and I have not found an explicit lectotype designation; it may have been inadvertently designated by Risch (1986). The publication date for Boulenger (1900c) was reported as 1 Nov. 1900, while that of Boulenger (1900a) was December, 1900, which gives
Chrysichthys brachynema priority.
Chrysichthys brevibarbis (Boulenger, 1899)
Chrysobagrus brevibarbis Boulenger, 1899a: 40, pl. 21 (fig. 1). Type locality: Boma. Holotype: MRAC 16.
Distribution: Congo River basin, below Stanley Falls (Risch, 1986).
Chrysichthys cranchii (Leach, 1818)
Pimelodus Cranchii Leach, in Tuckey, 1818: 409. Type locality: Lower Zaire. Holotype: BMNH 2005.5.17.4.
Chrysichthys börressoni Lönnberg, 1924: 620. Type locality: Lubosi river, a tributary to Luala river, Lower Congo.
Holotype: NRM 9942.
Distribution: Lualaba River system, Congo basin (Risch, 1986).
Chrysichthys dageti Risch, 1992
Chrysichthys dageti Risch, 1992a: 152, fig. 1. Type locality: Riv. Nanga entre Mboukoumassi et le lac Nanga,
Congo (04º15'S, 11º48'E). Holotype: MRAC 90-57-P-1273.
Distribution: Nanga River and tributaries, Congo (Risch, 1992a).
Chrysichthys delhezi Boulenger, 1899
Chrysichthys delhezi Boulenger, 1899b: 108, pl. 42 (fig. 2). Type locality: Boma. Lectotype: MRAC 952 (1)or
MRAC 953 (1). Boulenger (1911: 225, fig. 262) illustrated a specimen and referred to it as the type, from the
MRAC collection. If identifiable, that specimen is the lectotype of the species. If the illustrated specimen cannot
be identified, the two speciemens listed above, plus BMNH 1899.6.27.49 (1), are syntypes.
? Chrysichthys delhezi thomasi Chabanaud, 1934: 209. Type locality: Congo. Syntypes: MNHN 1937-0074 (1),
MNHN 1937-0075 (1).
Distribution: Lower Congo River system (Risch, 1986).
Chrysichthys duttoni Boulenger, 1905
Chrysichthys Duttoni Boulenger, 1905a: 641. Type locality: Lusambo, on the Kasai River. Holotype: MRAC 1342.
Distribution: Lualaba River system, Congo basin (Risch, 1986).
Chrysichthys furcatus Günther, 1864
Chrysichthys furcatus Günther, 1864: 430. Type locality: West Africa. Holotype: BMNH 1864.7.22.1.
Distribution: Coastal rivers of west Africa, from Liberia to Cabinda, Angola (Risch, 1986).
Chrysichthys grandis Boulenger, 1917
Chrysichthys grandis Boulenger, 1917b: 367. Type locality: Kilewa Bay [Lake Tanganyika]. Holotype: MRAC
14347.
Distribution: Lake Tanganyika (Risch, 1986).
Chrysichthys graueri Steindachner, 1911
Chrysichthys graueri Steindachner, 1911c: 529. Type locality: Tanganyika, am Ende des nordwestlichen Ufers. Syntypes: NMW 47106 (1), NMW 47107 (1). Species illustrated and described in more detail in Steindachner(1911e: 1178, pl. 3).
Distribution: Lake Tanganyika (Risch, 1986).
Chrysichthys habereri Steindachner, 1912
Chrysichthys habereri Steindachner, 1912: 445. Type locality: Dscha, Nebenfluss des Kongo, im Bezirke Molundu.
Holotype: NMW 47108. Species also described as new and illustrated in Steindachner (1913: 32, fig. 8 and pl.
4).
Distribution: Congo River system, below Stanley Falls (Risch, 1986).
Chrysichthys helicophagus Roberts & Stewart, 1976
Chrysichthys helicophagus Roberts & Stewart, 1976: 276, pl. 6 (fig. e). Type locality: Near Tadi, about 50 km downstream from Luozi. Lat. 5º14'S, Long. 13º56'E; lower Zaïre or Congo River. Holotype: MCZ 50292.
Distribution: Lower Congo River, below Malebo Pool (Risch, 1986).
Chrysichthys johnelsi Daget, 1959
Chrysichthys johnelsi Daget, 1959: 691, fig. 2. Type locality: Gambie à Bafoulabé (parc national du Niokolo-Koba).

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Syntypes (3): MNHN 1959-0146 (2) plus specimen described in Johnels (1954: 386), now NRM 13899.
Distribution: Rivers of the upper Guinea region, including the Geba, Taja, Moa, Mano, Loffa, St. Pauls, St. Johns,
Nipoué and Tano rivers (Risch, 1986).
Remarks: Redescribed, with comments on relationships and zoogeography, by Risch & Thys van den Audenaerde(1985a).
Chrysichthys laticeps Pellegrin, 1932
Chrysichthys laticeps Pellegrin, 1932: 166. Type locality: Congo, près de Brazzaville. Holotype: MNHN 1931-0159(but listed as 0158 in the publication).
Distribution: Congo River system (Risch, 1986).
Chrysichthys levequei Risch, 1988
Chrysichthys levequei Risch, 1988: 3, fig. 1. Type locality: du Samou, (bassin du Konkouré) en Guinée. Holotype:
MNHN 1986-0563.
Distribution: Konkouré River basin, Guinea (Risch, 1988).
Chrysichthys longibarbis (Boulenger, 1899)
Chrysobagrus longibarbis Boulenger, 1899a: 41, pl. 21 (fig. 2). Type locality: Léopoldville. Syntypes: BMNH
1899.2.20.18 (1), MRAC 108 (1).
Distribution: Congo River basin, below Stanley Falls (Risch, 1986).
Chrysichthys longidorsalis Risch & Thys van den Audenaerde, 1981
Gephyroglanis velifer Thys van den Audenaerde, 1965a: 265, fig. 1. Type locality: Ekongolo, Cameroon. Holotype:
MRAC 153158. Preoccupied in Chrysichthys by Chrysichthys velifer Norman, 1923; replaced by Chrysichthys
longidorsalis Risch & Thys van den Audenaerde, 1981.
Chrysichthys longidorsalis Risch & Thys van den Audenaerde, 1981: 247. Type locality: Ekongolo. Holotype:
MRAC 153158. Replacement for Gephyroglanis velifer Thys van den Audenaerde, 1965, preoccupied in
Chrysichthys by Chrysichthys velifer Norman, 1923 (a junior synonym of Chrysichthys maurus).
Distribution: Nyong River (Risch & Thys van den Audenaerde, 1985b), Middle Sanaga River, Cameroon (Risch,1986).
Chrysichthys longipinnis (Boulenger, 1899)
Gephyroglanis longipinnis Boulenger, 1899b: 109, pl. 41 (fig. 4). Type locality: Dolo, Stanley Pool [Congo River].
Lectotype: MRAC 958, designated by Risch (1981: 510).
Gephyroglanis gigas Pellegrin, 1922d: 278. Type locality: Kwamouth. Holotype: MRAC 14771.
Chrysichthys magnus Pellegrin, 1922d: 276. Type locality: Kwamouth. Syntypes (4): MRAC 14766 (1), MRAC
14767 (1), MRAC 14768–69 (2).
Distribution: Congo-Lualaba River system (Risch, 1986).
Chrysichthys mabusi Boulenger, 1905
Chrysichthys mabusi Boulenger, 1905b: 645. Type locality: Lake Bangwelo. Holotype: BMNH 1905.11.10.9.
Distribution: Congo-Lualaba River system, above Stanley Falls (Risch, 1986).
Chrysichthys macropterus Boulenger, 1920
Chrysichthys macropterus Boulenger, 1920a: 27, pl. 24. Type locality: Stanley Falls, ... d’Avakubi sur Ituri, et ...
Bafwasende sur la Lindi; Congo Belge. Syntypes (6): BMNH 1919.9.10.252 (1), BMNH 1919.9.10.253 (1),
MRAC 7118 (1), MRAC 7577 (1), MRAC 7578 (1).
Distribution: Congo River basin (Risch, 1986).
† Chrysichthys macrotis Van Neer, 1994
† Chrysichthys macrotis Van Neer, 1994: 100, text figs. 3–5, pl. 3 (fig. 1). Type locality: Nkondo-Kaiso area,
Knondo Formation, Albertine Rift Valley. Holotype: [apparently at MRAC] NK 1365'89 (neurocranium).
Distribution: Nkondo-Kaiso area, Lake Albert Rift, Uganda; Pliocene, Nkondo Formation (Van Neer, 1994; Murray& Budney, 2003).
Remarks: Considered to be a member of the Claroteidae by Gayet & Meunier (2003).
† Chrysichthys mahengeensis Murray & Budney, 2003
† Chrysichthys mahengeensis Murray & Budney, 2003: 985, fig. 1. Type locality: Mahenge, Singida Plateau, Tanzania, 4º47'38"S, 34º15'28"E, about 53 km west of the town of Singida; Eocene. Holotype: National Museum of

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Tanzania WM 386/96.
Distribution: Known only from type locality.
Chrysichthys maurus (Valenciennes, 1840)
Bagrus maurus Valenciennes, in Cuvier & Valenciennes, 1840a: 431 (319 of Strasbourg deluxe edition). Type locality: Sénégal. Holotype: MNHN 0000-1198.
Gephyroglanis lowei Norman, 1923b: 582. Type locality: Bandama River. Holotype: BMNH 1923.3.2.40.
Chrysichthys velifer Norman, 1923b: 582. Type locality: Bandama River. Holotype: BMNH 1923.3.2.41.
Distribution: West Africa (Risch, 1986).
Chrysichthys nigrodigitatus (La Cepède, 1803)
Pimelodus nigrodigitatus La Cepède, 1803: 95, 103. Type locality: [No locality stated; Senegal, according to Risch,1986]. Holotype: MNHN 0000-0108.
Arius acutivelis Valenciennes, in Cuvier & Valenciennes, 1840b: 85 (64 of Strasbourg deluxe edition). Type locality:
Gorée, ... haut Sénégal. Syntypes: MNHN a-8968 (2) and possibly MNHN 0000-0108.
Chrysichthys acutirostris Günther, 1864: 431. Type locality: Golungo Alto, Angola. Holotype: BMNH
1864.7.13.41.
Chrysichthys Büttikoferi Steindachner, 1894: 60, pl. 3 (fig. 1). Type locality: Fischermann-See [Liberia]. Holotype:
RMNH 5341; figured by Boeseman (1963: pl. 1, fig. 2).
Chrysichthys camaronensis Günther, 1899: 729, pl. 44. Type locality: Camaroons. Holotype: BMNH 1871.11.20.21.
As Chrysichthys camaronensis in text (p. 729, 732) and Chrysichthys cameronensis on plate; first reviser may
be Eschmeyer et al. (1998).
Chrysichthys ogowensis Günther, 1899: 723, fig. 5. Type locality: Kondo-Kondo, on the Ogowe River [Gabon].
Holotype: BMNH 1896.5.5.66.
Distribution: West Africa from Senegal to Cabinda, Angola (Risch, 1986).
Remarks: See Adépo-Gourène et al. (1997) for population differentiation.
Chrysichthys nyongensis Risch & Thys van den Audenaerde, 1985
Chrysichthys longidorsalis nyongensis Risch & Thys van den Audenaerde, 1985: 91. Type locality: Ebongo, Akonolinga et en amont des chutes, 12 km SE d’Esaka. Holotype: MRAC 73-29-P-1404 [or 1407].
Distribution: Nyong River, Cameroon (Risch, 1986).
Remarks: Risch (1986) reported catalog number of the holotype as MRAC 73-29-P-1404, but Eschmeyer et al.
(1998) listed MRAC 73-29-P-1407 as holotype, with -1404 as a paratype.
Chrysichthys ogooensis (Pellegrin, 1900)
Gephyroglanis ogooensis Pellegrin, 1900: 180. Type locality: Adouma (Ogôoué). Holotype: MNHN 1886-0422.
Gephyroglanis ogoensis Boulenger, 1901a: 227. Type locality: Adouma (Ogôoué). Holotype: MNHN 1886-0422.
Unjustified emendation or misspelling of Gephyroglanis ogooensis Pellegrin, 1900.
Distribution: Ogowe-Nyanga River basin, Gabon (Risch, 1986).
Remarks: Redescribed as Gephyroglanis ogooensis in Risch (1985a).
Chrysichthys okae Fowler, 1949
Chrysichthys okae Fowler, 1949: 261, figs. 53–58. Type locality: Oka, Congo system. Holotype: ANSP 71909.
Distribution: Central Congo basin, in tributary streams (Risch, 1986).
Chrysichthys ornatus Boulenger, 1902
Chrysichthys ornatus Boulenger, 1902d: 39, pl. 11 (fig. 2). Type locality: l’Ubangi à Banzyville [et] Monsembé,
Haut Congo. Syntypes (4): BMNH 1901.12.21.48–50 (3), MRAC 1174 (1).
Distribution: Congo River, below Stanley Falls (Risch, 1986).
Chrysichthys persimilis Günther, 1899
Chrysichthys persimilis Günther, 1899: 727, pl. 43; fig. 7. Type locality: Gaboon. Holotype: BMNH 1867.5.22.1.
Distribution: Gabon, Congo (Risch, 1986).
Remarks: Treated in Paugy & Bénech (1989: 302) as a synonym of Chrysichthys auratus (Geoffroy Saint-Hilaire,1809).
Chrysichthys platycephalus Worthington & Ricardo, 1937
Chrysichthys platycephalus Worthington & Ricardo, 1937: 1092, fig. 6. Type locality: Mpulungu-Niamkolo and the

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estuary of the Ngere River, Lake Tanganyika basin. Syntypes (38): BMNH 1936.6.15.849–852 (4), BMNH
1936.6.15.853–856 (4), BMNH 1936.6.15.857–878 (22).
Distribution: Lake Tanganyika (Risch, 1986).
Chrysichthys polli Risch, 1987
Chrysichthys polli Risch, 1987: 25, fig. 2. Type locality: Yelelola (falls). Lower Zaire (5º43'S, 13º34'E). Holotype:
MRAC 47814.
Distribution: Lower Congo River basin (Risch, 1987).
Chrysichthys punctatus Boulenger, 1899
Chrysichthys punctatus Boulenger, 1899b: 107, pl. 42 (fig. 1). Type locality: Stanley Pool and Kutu. Syntypes (9):BMNH 1899.11.27.25–26 (2), MRAC 950 (1), MRAC 951 (1).
Distribution: Congo River system below Stanley Falls (Risch, 1986).
Chrysichthys rueppelli Boulenger, 1907
Chrysichthys rueppelli Boulenger, 1907a: 337, pl. 60 (fig. 2). Type locality: Lower Nile and freshwater pools near
Ghet-el-Nassara, Lake Menzaleh. Syntypes (4): BMNH 1850.7.29.17 (1), BMNH 1907.12.2.2131–2132 (2).
Distribution: Nile River (Risch, 1986).
Chrysichthys sharpii Boulenger, 1901
Chrysichthys Sharpii Boulenger, 1901b: 81. Type locality: Lake Mwero. Holotype: BMNH 1900.12.31.14 (dryskin).
Distribution: Lake Mweru, Congo Basin; known only from holotype (Risch, 1986).
Chrysichthys sianenna Boulenger, 1906
Chrysichthys sianenna Boulenger, 1906f: 551, pl. 32 (fig. 2). Type locality: Niamkolo and .... Mbete [Lake Tanganyika]. Syntypes (3): BMNH 1906.9.8.65–66 (2), BMNH 1906.9.8.67 (1).
Distribution: Lake Tanganyika (Risch, 1986).
Chrysichthys stappersii Boulenger, 1917
Chrysichthys stappersii Boulenger, 1917b: 366. Type locality: Kilewa Bay [Lake Tanganyika]. Holotype: MRAC
14236; illustrated in Poll (1946: 211, figs. 18, 19).
Distribution: Lake Tanganyika (Risch, 1986).
Chrysichthys teugelsi Risch, 1987
Chrysichthys teugelsi Risch, 1987: 22, fig. 1. Type locality: Cavally River at Tai (5º52'N, 7º27'W), Ivory Coast.
Holotype: MRAC 80-19-P-189.
Distribution: Cavally River, Côte d’Ivoire (Risch, 1987).
Chrysichthys thonneri Steindachner, 1912
Chrysichthys thonneri Steindachner, 1912: 446. Type locality: Dschaflusse und oberen Kongo. Syntypes (3): NMW
10643 (1), NMW 10645 (1). Illustrated and described in more detail in Steindachner (1913: 33, fig. 9 and pl. 5).
Distribution: Congo-Lualaba River system (Risch, 1986).
Chrysichthys thysi Risch, 1985
Chrysichthys thysi Risch, 1985b: 189, figs. 5–6. Type locality: Makokou, riv. Ivindo. Holotype: MRAC 73-2-P1791.
Distribution: Ogowe-Nyanga River basin, Gabon (Risch, 1986).
Chrysichthys uniformis Pellegrin, 1922
Chrysichthys uniformis Pellegrin, 1922d: 277. Type locality: Poko (Uelé). Holotype: MRAC 7477.
Distribution: Congo, known only from holotype (Risch, 1986).
Chrysichthys wagenaari Boulenger, 1899
Chrysichthys wagenaari Boulenger, 1899a: 39, pl. 20 (fig. 1). Type locality: Upoto [Congo basin]. Holotype:
MRAC 147.
Distribution: Ubangi River and Middle Congo River system below Stanley Falls (Risch, 1986).
Chrysichthys walkeri Günther, 1899
Chrysichthys walkeri Günther, 1899: 720, fig. 3. Type locality: River Prah, Gold Coast. Syntypes (3): BMNH
1899.12.22.20–22 (3).
Distribution: Western Africa, from Ghana to Cabinda, Angola (Risch, 1986).

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Species inquirendae, Chrysichthys
Chrysichthys coriscanus Günther, 1899: 724. Type locality: Corisco Isld. [Equatorial Guinea]. Syntypes (2): BMNH
1896.5.5.64–65 (2). Treated in recent literature as a synonym of either Chrysichthys auratus or Chrysichthys
nigrodigitatus.
Chrysichthys kingsleyae Günther, 1899: 728, pl. 45 (fig. a); fig. 8. Type locality: River Ogowe. Holotype: BMNH
1899.12.23.1. Treated in recent literature either as a synonym of Chrysichthys walkeri Günther, 1899 (Risch, 1986:27) or Chrysichthys auratus (Geoffroy Saint-Hilaire, 1809) (Paugy & Bénech 1989: 302).
Chrysichthys lagoensis Günther, 1899: 725, fig. 6. Type locality: Lagos. Holotype: BMNH 1866.3.8.16. Treated as synonym of Chrysichthys nigrodigitatus or Chrysichthys maurus.
CLAROTES Kner, 1855
Clarotes Kner, 1855b: 313. Type species: Clarotes heuglini Kner, 1855. Type by monotypy. Gender: Masculine.
Gonocephalus Kner, 1855b: 313. Type species: Clarotes heuglini Kner, 1855. Type by monotypy. Gender: Masculine. Alternate new name for Clarotes. Preoccupied by Gonocephalus Kaup, 1825, in Reptilia.
Octonematichthys Bleeker, 1858b: 60, 65. Type species: Bagrus nigrita Valenciennes, 1840. Type by monotypy.
Gender: Masculine.
Clarotes bidorsalis Pellegrin, 1938
Clarotes bidorsalis Pellegrin, 1938b: 220, fig. on p. 221. Type locality: Giuba [Somalia]. Holotype: MNHN 19380139.
Distribution: Juba River, Somalia (Risch, 1986).
Clarotes laticeps (Rüppell, 1829)
Pimelodus laticeps Rüppell, 1829: 7, pl. 1 (fig. 2). Type locality: Cairo. Holotype: SMF 226.
Bagrus nigrita Valenciennes, in Cuvier & Valenciennes, 1840a: 426 (316 of Strasbourg deluxe edition), pl. 416.
Type locality: Sénégal. Syntypes: MNHN 0000-1204 (1), MNHN a-1113 (2).
Clarotes heuglini Kner, 1855b: 313, pls. 1–2. Type locality: bei Girf in der Nahe von Chartoum [Sudan]. Holotype:at NMW.
Chrysichthys macropogon Norman, 1928b: 385. Type locality: Kiyawa River (near Katagum), Northern Provinces,
Nigeria. Holotype: BMNH 1928.7.3.70.
Chrysichthys pitmani Fowler, 1936b: 312, figs. 97–100. Type locality: Athi River Crossing 16 miles north-northeastof Kibwezi, Kenya Colony. Holotype: ANSP 65888.
Distribution: East Africa, Nile, Niger and Senegal Rivers; Lake Chad (Risch, 1986).
Clarotes macrocephalus Daget, 1954
Clarotes macrocephalus Daget, 1954: 257, fig. 95. Type locality: Diafarabé [Niger River]. Syntypes (10): MNHN
1954-0003 (1).
Distribution: Niger, Volta and Senegal Rivers (Risch, 1986).
Remarks: MNHN 1954-0003 is listed in MNHN catalog as a holotype.
† EAGLESOMIA White, 1934
† Eaglesomia White, 1934: 70. Type species: † Chrysichthys eaglesomei White, 1926, by original designation. Gender: Feminine.
Remarks: Included in the Claroteidae in Gayet & Meunier (2003).
† Eaglesomia eaglesomei (White, 1926)† Chrysichthys eaglesomei White, 1926: 59, pls. 14–15. Type locality: Ameki, Nigeria, Eocene. Holotype: BMNH
P.11944–11946 (posterior half of skull with anterior vertebral complex).
Remarks: Redescribed in White (1934: 70, pl. 8, 9, pl. 10, fig. 28).
GEPHYROGLANIS Boulenger, 1899
Gephyroglanis Boulenger, 1899a:42. Type species: Gephyroglanis congicus Boulenger, 1899. Type by monotypy.

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Gender: Masculine.
Gephyroglanis congicus Boulenger, 1899
Gephyroglanis congicus Boulenger, 1899a: 42, pl. 20 (fig. 2). Type locality: Upoto [Congo basin]. Holotype: MRAC
146.
Distribution: Congo-Lualaba River system (Risch, 1986).
Gephyroglanis gymnorhynchus Pappenheim, 1914
Gephyroglanis gymnorhynchus Pappenheim, in Pappenheim & Boulenger, 1914: 248, pl. 4 (fig. 6). Type locality:
Aruwimi bei Basoko. Holotype: ZMB 19092.
Distribution: Aruwimi River and Port Congo (Risch, 1986).
Gephyroglanis habereri Steindachner, 1912
Gephyroglanis habereri Steindachner, 1912: 449. Type locality: Dscha, Nebenflusse des Kongo, im Bezirke Molundu. Holotype: NMW 46570. Species illustrated and described in more detail in Steindachner (1913: 36, pl. 6).
Distribution: Congo River; known only from holotype (Risch, 1986).
GNATHOBAGRUS Nichols & Griscom, 1917
Gnathobagrus Nichols & Griscom, 1917: 711. Type species: Gnathobagrus depressus Nichols & Griscom, 1917.
Type by monotypy. Gender: Masculine.
Gnathobagrus depressus Nichols & Griscom, 1917
Gnathobagrus depressus Nichols & Griscom, 1917: 712, fig. 21. Type locality: Boma, Congo. Holotype: AMNH
6647.
Distribution: Congo River at Boma; known only from type locality (Risch, 1986).
LIAUCHENOGLANIS Boulenger, 1916
Liauchenoglanis Boulenger, 1916: 314. Type species: Liauchenoglanis maculatus Boulenger, 1916. Type by monotypy. Gender: Masculine.
Remarks: Treated in recent literature as valid or as a synonym of Notoglanidium.
Liauchenoglanis maculatus Boulenger, 1916
Liauchenoglanis maculatus Boulenger, 1916: 314, fig. 186. Type locality: N. Sherbo dist., Sierra Leone. Syntypes:
BMNH 1915.5.19.9–12 (4).
Distribution: Sierra Leone (Risch, 1986).
Remarks: Also treated in recent literature as Notoglandium maculatus [sic, N. maculatum].
LOPHIOBAGRUS Poll, 1942
Lophiobagrus Poll, 1942b: 318. Type species: Lophiobagrus lestradei Poll, 1942. Type by monotypy. Gender: Masculine.
Lophiobagrus aquilus Bailey & Stewart, 1984
Lophiobagrus aquilus Bailey & Stewart, 1984: 24, figs. 5b, 6b, 8. Type locality: Lake Tanganyika. Holotype:
UMMZ 199928.
Distribution: Lake Tanganyika; known only from type series (Risch, 1986).
Lophiobagrus asperispinis Bailey & Stewart, 1984
Lophiobagrus asperispinis Bailey & Stewart, 1984: 33, figs. 6d, 9. Type locality: Lake Tanganyika. Holotype:
MRAC 14359A.
Distribution: Lake Tanganyika (Risch, 1986).
Lophiobagrus brevispinis Bailey & Stewart, 1984
Lophiobagrus brevispinis Bailey & Stewart, 1984: 35, figs. 5c, 6c, 10. Type locality: Lake Tanganyika. Holotype:
UMMZ 199930.
Distribution: Lake Tanganyika (Risch, 1986).
Lophiobagrus cyclurus (Worthington & Ricardo, 1937)
Chrysichthys cyclurus Worthington & Ricardo, 1937: 1093, fig. 7. Type locality: Lake Tanganyika. Lectotype:
BMNH 1936.6.15.879 (49 mm SL), designated by Bailey & Stewart (1984: 20).

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Lophiobagrus lestradei Poll, 1942b: 320, fig. on p. 321. Type locality: Nyanza, Lake Tanganyika. Holotype: MRAC
54924.
Distribution: Lake Tanganyika (Risch, 1986).
NOTOGLANIDIUM Günther, 1903
Notoglanidium Günther, 1903: 336. Type species: Notoglanidium walkeri Günther, 1903. Type by monotypy. Gender: Neuter.
Notoglanidium pallidum Roberts & Stewart, 1976
Notoglanidium pallidum Roberts & Stewart, 1976: 276, pl. 7 (figs. a–c). Type locality: Near Bulu, about 15 km
downstream from Luozi. Lat. 5º01'S, Long. 14º01'E; lower Zaïre or Congo River. Holotype: MCZ 50236.
Distribution: Lower Zaire (Risch, 1986).
Notoglanidium thomasi Boulenger, 1916
Notoglanidium thomasi Boulenger, 1916: 313, fig. 185. Type locality: Victoria and Pujehun [Sierra Leone]. Syntypes: BMNH 1915.5.21.10 (1), BMNH 1915.5.27.3–4 (2).
Distribution: Sierra Leone (Risch, 1986).
Notoglanidium walkeri Günther, 1903
Notoglanidium walkeri Günther, 1903: 337, pl. 32. Type locality: River Ibbi [Ghana]. Syntypes (2): BMNH
1903.4.24.78–79 (2).
Distribution: Ghana and Côte d’Ivoire (Risch, 1986).
PARAUCHENOGLANIS Boulenger, 1911
Parauchenoglanis Boulenger, 1911a: 364. Type species: Pimelodus guttatus Lönnberg, 1895. Type by subsequent
designation by Jordan (1920: 538). Gender: Masculine.
Remarks: Included species follows Teugels et al. (1991); species synonymies follow Risch (1986).
Revision: Geerinckx et al. (2004), with key to species.
Parauchenoglanis ahli (Holly, 1930)
Auchenoglanis ahli Holly, 1930: 201, pl. 1 (fig. 9). Type locality: Gebirchsbache von Bakoko, Kamerun. Syntypes:
NMW 13895–99 (5); syntype illustrated in Geerinckx et al. (2004: 787, fig. 8i).
Distribution: Cross River basin, western Cameroon (Geerinckx et al., 2004; known only from types).
Parauchenoglanis altipinnis (Boulenger, 1911)
Auchenoglanis altipinnis Boulenger, 1911e: 372, fig. 288. Type locality: Ja River at Esamesa, Cameroon. Holotype:
BMNH 1911.5.30.30; illustrated in Geerinckx et al. (2004: 787, fig. 8d).
Distribution: Congo, Nyong, Ogowe and Chiloango River basins (Geerinckx et al., 2004: 798).
Parauchenoglanis balayi (Sauvage, 1879)
Pimelodus Balayi Sauvage, 1879: 98. Type locality: Lopé [Ogôoué River]. Holotype: MNHN a-0898. Described in
more detail in Sauvage (1880c: 44).
Pimelodus Guirali Thominot, 1886: 162. Type locality: [Not stated; as Rivière San Benito in MNHN catalog]. Holotype: MNHN 1885-0427; illustrated in Geerinckx et al. (2004: 787, fig, 8a).
Auchenoglanis ballayi Boulenger, 1901: 295. Type locality: Lopé [Ogôoué River]. Holotype: MNHN a-0898. Incorrect subsequent spelling or unjustified emendation of Pimelodus balayi Sauvage, 1879.
Auchenoglanis pulcher Boulenger, 1902g: 267, pl. 29 (figs. 2–2a). Type locality: Lindi River, which flows into theCongo at the Stanley Falls. Syntypes (several): BMNH 1902.4.14.19–20 (2), IRSNB 10 (2), MSNM 35 (1).
Auchenoglanis ubangensis Boulenger, 1902d: 41, pl. 11 (fig. 3). Type locality: l’Ubangi à Banzyville. Holotype:
MRAC 1215.
Auchenoglanis ballayi gravoti Pellegrin, 1907b: 321. Type locality: N'Tem, Cameroun. Syntypes: MNHN 19070158 (1), MNHN 1907-0159 (1), MNHN 1907-0160 (1). Originally proposed as Auchenoglanis ballayi var.
gravoti.
Auchenoglanis pietschmanni Holly, 1926: 158. Type locality: Bamfluss, Flußgebiet des Sanaga in Kamerun. Syntypes (5): NMW 7802 (1), NMW 7803 (1), NMW 7804 (1), NMW 78025 (1), NMW 7806 (1). Illustrated anddescribed in more detail in Holly (1927b: 208, fig. 3).

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Parauchenoglanis buettikoferi (Popta, 1913)
Auchenoglanis büttikoferi Popta, 1913: 237, pl. 10. Type locality: Warri, in Upper Nigeria. Holotype: RMNH 8860;
illustrated in Geerinckx et al. (2004: 787, fig. 8g).
Distribution: Upper Nigeria; known only from the holotype (Risch, 1986; Geerinckx et al., 2004).
Parauchenoglanis longiceps (Boulenger, 1913)
Auchenoglanis longiceps Boulenger, 1913: 68. Type locality: Nyong River, S. Cameroon. Holotype: BMNH
1913.10.29.12.
Auchenoglanis maculosus Holly, 1927b: 211, fig. 4. Type locality: Kamerun. Syntypes: NMW 7800 (1), NMW 7801
(1).
Distribution: Nyong River, central and southern Cameroon (Geerinckx et al., 2004: 800).
Parauchenoglanis monkei (Keilback, 1910)
Pimelodus guttatus Lönnberg, 1895: 184. Type locality: river N’Dian [Cameroon]. Syntypes: NRM (2, apparentlylost), ZMUU 567 (2). One syntype illustrated in Teugels et al. (1991: 502, fig. 1). Preoccupied by Pimelodus
guttatus La Cepède, 1803; replaced by Parauchenoglanis loennbergi Fowler, 1958.
Auchenoglanis monkei Keilhack, 1910: 120. Type locality: in einem Kleinen Bache im Urwald 2 km südlich vonLogobaba (4º2'N, 9º45'Ö), Kamerun. Syntypes (3): ZMB 17771 (1), ZMB 21318 (2).
Parauchenoglanis loennbergi Fowler, 1958: 14. Type locality: river N’Dian [Cameroon]. Syntypes: NRM (2, lost?),ZMUU 567 (2). Replacement name for Pimelodus guttatus Lönnberg, 1895, preoccupied by Pimelodus guttatus
La Cepède, 1803.
Auchenoglanis fasciatus Gras, 1961: 404, fig. 2. Type locality: Bokoutou, Bas-Dahomey. Syntypes (2): MNHN
1960-0392 (2); one syntype illustrated in Geerinckx et al. (2004: 787, fig. 8b).
Distribution: Ouémé River, Benin, to the Sanaga River, Cameroon (Geerinckx et al., 2004 [as Parauchenoglanis
guttatus]).
Parauchenoglanis ngamensis (Boulenger, 1911)
Auchenoglanis ngamensis Boulenger, 1911a: 371, fig. 287. Type locality: Okovango River. Holotype: BMNH
1910.5.31.35; illustrated in Geerinckx et al. (2004: 787, fig. 8e).
Distribution: Okavango and upper Zambezi basins, Kasai River, Congo River basin (Skelton, 1993; Geerinckx et al.,2004).
Remarks: First transferred into Parauchenoglanis by Skelton (1993).
Parauchenoglanis pantherinus (Pellegrin, 1929)
Auchenoglanis pantherinus Pellegrin, 1929b: 363. Type locality: Nyabessan (Ntem), Cameroun. Syntypes (4):MNHN 1929-0029 (1), MNHN 1929-0030 (1), MNHN 1929-0031 (2).
Distribution: Ntem River, Cameroon (Risch, 1986; Geerinckx et al., 2004: 801).
Parauchenoglanis punctatus (Boulenger, 1902)
Auchenoglanis punctatus Boulenger, 1902g: 267, pl. 29 (figs. 1–1a). Type locality: Lindi River, which flows into theCongo at the Stanley Falls. Holotype: IRSNB 9.
Auchenoglanis iturii Steindachner, 1911b: 530. Type locality: Aus dem Ituriflüss bei Mawambi. Syntypes: NMW
47474 (1), NMW 47475 (1), NMW 47476 (1); NMW 47475 illustrated in Geerinckx et al. (2004: 787, fig. 8c).
Species described in more detail in Steindachner (1911e: 1185).
Auchenoglanis grandis Fowler, 1936b: 312, fig. 96. Type locality: Nola, Middle Congo. Holotype: ANSP 66125.
Distribution: lower and central Congo River system; San Benito and Ogowe River basins (Geerinckx et al., 2004:
796).
PARDIGLANIS Poll, Lanza & Romoli Sassi, 1972
Pardiglanis Poll, Lanza & Romoli Sassi, 1972: 330. Type species: Pardiglanis tarabinii Poll, Lanza & Romoli
Sassi, 1972. Type by original designation. Gender: Masculine.
Pardiglanis tarabinii Poll, Lanza & Romoli Sassi, 1972
Pardiglanis tarabinii Poll, Lanza & Romoli Sassi, 1972: 338, figs. 1–7. Type locality: Gelib, fleuve Juba, Somalie
méridionale. Holotype: MZUF 1579.

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Distribution: Juba River, Somalia (Risch, 1986).
Remarks: See De Vos (2001b) for comments on this species.
PHYLLONEMUS Boulenger, 1906
Phyllonemus Boulenger, 1906f: 552. Type species: Phyllonemus typus Boulenger, 1906. Type by original designation. Gender: Masculine.
Phyllonemus brichardi Risch, 1987
Phyllonemus brichardi Risch, 1987: 29, fig. 3. Type locality: Ubwari, face est, lac Tanganyika (Zaire). Holotype:
MRAC 83-19-P-14.
Distribution: Lake Tanganyika (Risch, 1987).
Phyllonemus filinemus Worthington & Ricardo, 1937
Phyllonemus filinemus Worthington & Ricardo, 1937: 1096, fig. 9. Type locality: Kibwesi [and] Kigoma, Lake Tanganyika. Syntypes (8): BMNH 1936.6.15.925 (8).
Distribution: Lake Tanganyika (Risch, 1986).
Phyllonemus typus Boulenger, 1906
Phyllonemus typus Boulenger, 1906f: 552, pl. 32 (fig. 3). Type locality: Niamkolo [Lake Tanganyika]. Holotype:
BMNH 1906.9.8.68.
Distribution: Lake Tanganyika (Risch, 1986).
PLATYGLANIS Daget, 1979
Platyglanis Daget, 1979: 821. Type species: Platyglanis depierrei Daget, 1979. Type by original designation. Gender: Masculine.
Platyglanis depierrei Daget, 1979
Platyglanis depierrei Daget, 1979: 821, figs. 1–2. Type locality: Djim (affluent du Mbam lequel se jette dans la Sanaga en aval des chutes Nachtingal) aux environs du pont franchissant la rivière sur la piste qui mène de Ngila a
Ngorro [Cameroon]. Holotype: MNHN 1978-0759.
Distribution: Sanaga Basin, Cameroon; known only from the type locality (Risch, 1986).
RHEOGLANIS Poll, 1966
Rheoglanis Poll, 1966: 425. Type species: Rheoglanis dendrophorus Poll, 1966. Type by original designation. Gender: Masculine.
Rheoglanis dendrophorus Poll, 1966
Rheoglanis dendrophorus Poll, 1966: 425, unnumbered figure. Type locality: Kinsuka, rapides du fleuve. Holotype:
MRAC 153744.
Distribution: Lower Congo River, in rapids (Risch, 1986).
Species inquirendae, Claroteidae or Bagridae
Mystus fluviatilis Klein 1775 [Not seen].
Leptoglanis bouilloni Poll, 1959: 97, pl. 25 (fig. 1). Type locality: Stanley Pool. Holotype: MRAC 118507. Althoughoriginally considered an amphiliid, Roberts (2003) suggested that the holotype was a claroteid or bagrid.

CRANOGLANIDIDAE Myers, 1931
Cranoglanidae Myers, 1931: 261. Type genus: Cranoglanis Peters, 1881.
Taxonomic overview: Jayaram (1956b); Ng & Kottelat (2000d).
1 genus, 3 species; no named fossil taxa.
CRANOGLANIS Peters, 1881
Cranoglanis Peters, 1881b: 1030. Type species: Cranoglanis sinensis Peters, 1881. Type by monotypy. Gender:

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Masculine.
Anopleutropius Vaillant, 1893b: 198. Type species: Anopleutropius henrici Vaillant, 1893. Type by monotypy. Gender: Masculine.
Pseudeutropichthys Koller, 1926: 74. Type species: Pseudeutropichthys multiradiatus Koller, 1926. Type by original
designation. Gender: Masculine. Also appeared in Koller (1927: 28).
Remarks: Considered to be a monotypic genus by Liu et al. (2005), with only Cranoglanis bouderius as valid.
Cranoglanis bouderius (Richardson, 1846)
Bagrus bouderius Richardson, 1846: 283. Type locality: Lian Jiang, Lianzhou, Guangdong Prov., China. Neotype:
ZMB 11325, designated by Ng & Kottelat (2000d: 851). Originally as ? Bagrus (an Pimelodus ?) bouderius.
Based solely on an illustration by Reeves; which was reproduced in Jayaram (1956b: pl. 18) and Whitehead(1970: pl. 19b).
Macrones sinensis Bleeker, 1872: 153. Type locality: Lian Jiang, Guangdon Prov., Lianzhou, China. Neotype: ZMB
11325 designated by Ng & Kottelat (2000d: 851). Based originally on a painting, which was reproduced in
Jayaram & Boeseman (1976: pl. 1).
Cranoglanis sinensis Peters, 1881b: 1030, [pl. 1] (fig. 1). Type locality: Lian Jiang, Lianzhou, Guangdong Prov.,
China. Holotype: ZMB 11325. Preoccupied in Cranoglanis by Macrones sinensis Bleeker, 1872.
Distribution: Southern China.
Cranoglanis henrici (Vaillant, 1893)
Anopleutropius henrici Vaillant, 1893b: 199. Type locality: Rivière Noir, Tonkin [Vietnam]. Syntypes: MNHN
1892-0259 (1), MNHN 1892-0260 (1).
Distribution: Red River basin, northern Vietnam and southern China (Ng & Kottelat, 2000d).
Cranoglanis multiradiatus (Koller, 1926)
Pseudeutropichthys multiradiatus Koller, 1926: 74. Type locality: Kan-Kongfluß auf Hainan [China]. Syntypes:
NMW 5059–5061 (3). Also appeared as new in Koller (1927: 29, pl. 1, fig. 2).
Distribution: Hainan Island, China.

DIPLOMYSTIDAE Eigenmann, 1890
Diplomystidae Eigenmann, 1890: 14. Type genus: Diplomystes Bleeker, 1858.
Reviews: Azpelicueta (1994a, 1994b), Argentina; Arratia (1987) Chile.
Phylogeny: Arratia (1987).
Taxonomic summary: Ferraris (2003a).
2 genera, 6 species; no named fossil taxa.
DIPLOMYSTES Bleeker, 1858
Diplomyste Duméril, 1856: 487. Type species: Arius papillosus Valenciennes, 1840. Type by monotypy. Gender:
Masculine. Predates Diplomystes Bleeker, 1858, but the latter in wide use.
Diplomystes Bleeker, 1858b: 63. Type species: Arius papillosus Valenciennes, 1840. Type by monotypy. Gender:
Masculine.
Diplomystax Günther, 1864: 180. Type species: Arius papillosus Valenciennes, 1840. Type by original designation.
Gender: Masculine. Unneeded replacement for Diplomyste Duméril and Diplomystes Bleeker.
Diplomystes camposensis Arratia, 1987
Diplomystes camposensis Arratia, 1987: 44, fig. 19. Type locality: Lago Riñihue, Chile. Holotype: IZUA 3302.
Distribution: Valdivia region, southern Chile (Arratia, 1987).
Diplomystes chilensis (Molina, 1782)
Silurus chilensis Molina, 1782: 225. Type locality: Chile. Type(s): Whereabouts unknown.
Arius papillosus Valenciennes in Cuvier & Valenciennes, 1840b: 118 (88 in the Strasbourg deluxe edition), pl. 431.
Type locality: Rivières de Valparaíso et de San-Jago du Chili. Syntypes: MNHN 0000-1212 (1), MNHN b-0584(1), MNHN b-0585 (4), MNHN a-9028 (1), MNHN uncataloged (skeleton, missing).

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Arius cacharioides Leybold, 1859: 1083, pls. 1–2. Type locality: Ad thermos Colina, dictas in flumine, Río Seco
(Chile). Syntypes (100): Whereabouts unknown.
? Arius micropterus Philippi, 1866: 713. Type locality: (Chile). Type(s): Whereabouts unknown.
Arius squalus Philippi, 1866: 713. Type locality: Peine, Santiago, Chile. Type(s): Whereabouts unknown.
? Arius synodon Philippi, 1866: 714. Type locality: (Chile). Type(s): Whereabouts unknown.
? Arius villosus Philippi, 1866: 712. Type locality: (Chile). Type(s): Whereabouts unknown.
Distribution: Rivers near Valparaiso and Santiago, central Chile (Arratia 1987).
Remarks: Arratia (1987) reports the species as either extinct or endangered. The imprecise type localities of many of
the names listed in the synonymy make it impossible to know whether the names belong here or with anotherspecies of Diplomystes.
Diplomystes nahuelbutaensis Arratia, 1987
Diplomystes nahuelbutaensis Arratia, 1987: 33, fig. 11. Type locality: Río Cautín, Lautaro, Chile. Holotype: CAS
55423.
Distribution: Bío-Bío River basin and Loncomilla River, Chile (Arratia, 1987; Ruiz & Berra, 1994, with biologicalnotes).
OLIVAICHTHYS Arratia, 1987
Olivaichthys Arratia, 1987: 66. Type species: Diplomystes viedmensis MacDonagh, 1931. Type by original designation. Gender: Masculine.
Key: Azpelicueta (1994a, as Diplomystes).
Review: Azpelicueta (1994b, as Diplomystes).
Olivaichthys cuyanus (Ringuelet, 1965)
Diplomystes viedmensis cuyanus Ringuelet, 1965: 91. Type locality: Arroyo Yaucha (Vilucó, Mendoza) [Argentina].
Holotype: MLP 286.
Distribution: Colorado River and tributaries; Desaguadero-Salado River basin, including the rivers San Juan, Mendoza, Tunuyán and temporary environments connected to those rivers, Argentina (Azelpelicueta, 1994b).
Olivaichthys mesembrinus (Ringuelet, 1982)
Diplomystes viedmensis mesembrinus Ringuelet, 1982: 349, figured on p. 350. Type locality: Desembocadura delRío Senguer en el lado Musters (45º60'S, 49º10'W) [Chubut, Argentina]. Holotype: ILPLA 8452.
Distribution: Chubut and Senguer River basins, Argentina (Azpelicueta, 1994b).
Remarks: Redescribed in Azpelicueta & Gosztonyi (1998) as Diplomystes mesembrinus.
Olivaichthys viedmensis (MacDonagh, 1931)
Diplomystes viedmensis MacDonagh, 1931: 65. Type locality: río Negro, Argentina, frente a Viedma. Holotype:
MLP 214.
Distribution: South America: Negro River and tributaries in Argentina (Azpelicueta, 1994b).

DORADIDAE Bleeker, 1858
Doradini Bleeker, 1858b: 48, 52. Type genus: Doras La Cepède, 1803.
Remarks: The family-group names Centrochirinae, Hemidoradinae, and Lithodoradinae were proposed by Fowler(1951b: 3) in a manner that made the names available when published, but the names were rendered unavailable in1961 by Art. 13 of the Second Edition of the Code (ICZN, 1961). The names apparently remain unavailable, despite
Art. 13.2.1 of the current version of the Code (ICZN, 1999), because they were treated as unavailable in Ferraris &de Pinna (1999: 8).
Revision: Eigenmann (1925).
Review: Fernández-Yépez (1968, Venezuela).
Phylogeny: Higuchi (1992), Moyer et al. (2004).
Taxonomic summary: Sabaj (2002); Sabaj & Ferraris (2003).
30 genera, 77 species; no named fossil taxa.

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Incertae sedis
Doras fimbriatus Kner, 1855
Doras loricatus Kner, 1853c: 146, fig. 2 on unnumbered plate. Type locality: [Not stated]. Holotype: NMW 45407–
09 (1 of 4). Name made available from caption to illustration of a swimbladder; associated specimen is the holotype, if recognizable. Illustrated and described in more detail, as Doras fimbriatus, in Kner (1855: 134, pl. 3, fig.
5) with locality given as Rio Guaporé. Considered a nomen oblitum in Sabaj & Ferraris (2003).
Doras fimbriatus Kner, 1855a: 134, pl. 3 (fig. 5). Type locality: Rio Guaporé. Syntypes: NMW 45407–09 (4).
Distribution: Guaporé River basin, Bolivia and Brazil (Sabaj & Ferraris, 2003).
Remarks: Generic placement of this species unresolved (Sabaj & Ferraris, 2003).
Doras punctatus Kner, 1853
Doras (Corydoras) punctatus Kner, 1853c: 146, fig. 5 on unnumbered plate. Type locality: [Not stated]. Holotype:
NMW 45434–41 (1 of 14) or RMNH 2966 (1). Name made available from caption to illustration of a swimbladder; associated specimen is the holotype, if recognizable. Illustrated and described in more detail in Kner (1855:
136, pl. 6, fig. 10), with locality given as Mato-grosso und Rio Guaporé.
Distribution: Amazon River basin (Sabaj & Ferraris, 2003).
Remarks: Generic placement of this species unresolved (Sabaj & Ferraris, 2003).
Oxydoras eigenmanni Boulenger, 1895
Oxydoras eigenmanni Boulenger, 1895b: 524. Type locality: Descalvados, Matto Grosso. Syntypes: BMNH
1895.5.17.50–52 (3), BMNH 1895.5.17.53 (1); one syntype illustrated in Boulenger (1896c: pl. 4, fig. 3).
Distribution: Amazon and upper Paraguay River basins (Sabaj & Ferraris, 2003).
Remarks: Generic placement of this species unresolved (Sabaj & Ferraris, 2003).
ACANTHODORAS Bleeker, 1862
Cataphractus Edwards, in Catesby, 1771: appendix. Type species: Silurus cataphractus Linnaeus, 1758. Type by
absolute tautonymy. Gender: Masculine. Considered a nomen oblitum (Sabaj & Ferraris, 2003).
Acanthodoras Bleeker, 1862 (in Bleeker, 1862–63): 5. Type species: Silurus cataphractus Linnaeus, 1758. Type byoriginal designation. Gender: Masculine.
Acanthodoras cataphractus (Linnaeus, 1758)
Silurus cataphractus Linnaeus, 1758: 307. Type locality: America. Syntype: BMNH 1853.11.12.193 (1). BMNH
syntype illustrated in Gronovius (1754: pl. 3, figs. 3 & 4); another syntype illustrated in Catesby (1731–1743:
19, pl. 9).
Cataphractus Americanus Bloch & Schneider, 1801: 107, pl. 28. Type locality: America, pedalis. Syntype: BMNH
1853.11.12.193 (1). BMNH syntype figured in Gronovius (1754: pl. 3, figs. 3 & 4). Unneeded new name for
Silurus cataphractus Linnaeus.
Cataphractus americanus La Cepède, 1803: 124, 126. Type locality: dans la Caroline. Syntypes: On several literature accounts, including Catesby (1731–1743), Gronovius (1754), Bonnaterre (1788), and Gmelin (1789); only
extant syntype: BMNH 1853.11.12.193 (1) illustrated in Gronovius (1754: pl. 3, figs. 3 & 4); another syntype
illustrated in Catesby (1731–1743: 19, pl. 9). Name apparently proposed independently of Cataphractus americanus Bloch & Schneider. Type locality reported by La Cepède was based on account in Catesby.
Doras Blochii Valenciennes, in Cuvier & Valenciennes, 1840b: 277 (207 of Strasbourg deluxe edition). Type locality: [Not stated]. Syntype: BMNH 1853.11.12.193 (1) and literature sources. BMNH syntype figured in
Gronovius (1754: pl. 3, figs. 3 & 4).
Callichthys asper Gronow, in Gray, 1854: 157. Type locality: Americes Meridionalis rivulis. Syntype: BMNH
1853.11.12.193 (1). BMNH syntype figured in Gronovius (1754: pl. 3, figs. 3 & 4).
Distribution: Amazon River basin and coastal drainages of French Guiana, Guyana and Suriname (Sabaj & Ferraris,2003).
Acanthodoras depressus (Steindachner, 1881)
Doras (Rhinodoras) depressus Steindachner, 1881c: 103, pl. 1 (fig. 3). Type locality: Ausstände (Lago Alexo) am

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mittleren Laufe des Amazonen-Stromes. Holotype: NMW 46870.
Distribution: Amazon and Negro River basins, Brazil (Sabaj & Ferraris, 2003).
Acanthodoras spinosissimus (Eigenmann & Eigenmann, 1888)
Doras brunnescens Jardine, in Schomburgk, 1841: 163. Type locality: Upper Essequibo [Guyana]. No types known.
Doras spinosissimus Eigenmann & Eigenmann, 1888b: 161. Type locality: Coary [Brazil]. Holotype: MCZ 7222.
Distribution: Amazon and Essequibo River basins (Sabaj & Ferraris, 2003).
Remarks: Acanthodoras spinosissimus treated as valid, following Sabaj & Ferraris (2003), in which Doras brunnescens is treated as a nomen oblitum.
Species inquirendae, Acanthodoras
Doras castaneo-ventris Jardine, in Schomburgk, 1841: 161, pl. 3. Type locality: river Pasawiri. No types known.
Doras polygramma Kner, 1853c: 145, fig. 1 on unnumbered plate. Type locality: [Not stated]. Holotype: NMW 46846(99 mm SL, 1 of 2). Name made available from caption to illustration of a swimbladder; associated specimen is theholotype, if recognizable. Treated in Kner (1855: 126–127) as Doras cataphractus, based on specimens from “Rio
Guaporé and Barra do Rio negro.”Remarks: Catalog number for type of Doras polygramma incorrectly reported as NMW 46844 by Sabaj and Ferraris(2003).
AGAMYXIS Cope, 1878
Agamyxis Cope, 1878: 679. Type species: Doras pectinifrons Cope, 1870. Type by monotypy. Gender: Feminine.
Agamyxis albomaculatus (Peters, 1877)
Doras albomaculatus Peters, 1877: 470. Type locality: Calabozo [Venezuela]. Syntypes (2): ZMB 10043 (1).
Distribution: Orinoco River basin, Venezuela (Sabaj & Ferraris, 2003).
Agamyxis pectinifrons (Cope, 1870)
Doras pectinifrons Cope, 1870b: 568. Type locality: Pebas, Ecuador [now Peru]. Holotype: ANSP 8346.
Doras (Agamyxis) flavopictus Steindachner, 1908c: 84. Type locality: Iquitos [Amazonasgebiet]. Syntypes: NMW
46598–99 (2).
Distribution: Amazon River basin (Sabaj & Ferraris, 2003).
AMBLYDORAS Bleeker, 1862
Amblydoras Bleeker, 1862 (in Bleeker, 1862–63): 5. Type species: Doras affinis Kner, 1855. Type by original designation. Gender: Masculine.
Zathorax Cope, 1871b: 112. Type species: Zathorax monitor Cope, 1872. Type by subsequent monotypy. Gender:
Masculine. Appeared first with no included species, subsequently described with one species (Cope, 1872a:
271).
Amblydoras affinis (Kner, 1855)
Doras affinis Kner, 1855a: 121, pl. 2 (fig. 1). Type locality: Rio branco und Guaporé. Syntypes (11): NMW 46822–
23 (4), NMW 46826 (3), RMNH 2973 (1).
Amblydoras truncatus Bleeker, 1863b: 18. Type locality: flumine Guapore. Holotype: RMNH 2973. Holotype isalso a syntype of Doras affinis Kner.
Distribution: Guaporé, Branco and Essequibo River basins (Sabaj & Ferraris, 2003).
Amblydoras bolivarensis (Fernández-Yépez, 1968)
Hildadoras bolivarensis Fernández-Yépez, 1968: 43, fig. 18. Type locality: Qda. El Pilón, al este del Río Parguaza,
afluente (margen derecho) del Río Orinoco, frente a Pararuma [Estado Bolívar, Venezuela]. Holotype: FMNH
84067.
Distribution: Orinoco River basin, Venezuela (Sabaj & Ferraris, 2003).
Amblydoras gonzalezi (Fernández-Yépez, 1968)
Zathorax gonzalezi Fernández-Yépez, 1968: 70, fig. 36. Type locality: Laguna “Punta Vista,” a orillas del Río
Caroní (margen izquierda) en el Parque Cachamay [Venezuela]. Holotype: MBUCV V-3940.

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FERRARISDistribution: Orinoco River basin and Casiquiare Canal (Sabaj & Ferraris, 2003).
Amblydoras monitor (Cope, 1872)
Zathorax monitor Cope, 1872a: 272, pl. 4 (figs. 1a–1c). Type locality: small streams tributary to the Ambyiacu, aswell as ... the river itself ... which empties into the Amazon near Pebas, in Eastern Ecuador, some distance eastof the Napo. Lectotype: ANSP 8296, designated by and illustrated in Eigenmann (1925: 363, pl. 16, fig. 3).
Distribution: Upper Amazon River basin (Sabaj & Ferraris, 2003).
Amblydoras nauticus (Cope, 1874)
Zathorax nauticus Cope, 1874b: 133. Type locality: Nauta [Upper Amazon]. Syntypes: ANSP 21390–95 (6).
Distribution: Upper Amazon River basin, Peru (Sabaj & Ferraris, 2003).
Remarks: Syntype series contains two species; with four specimens of Amblydoras monitor Cope and two representing the species indicated here (Sabaj & Ferraris, 2003). A lectotype designation is necessary to formalize thevalidity of this name.
ANADORAS Eigenmann, 1925
Anadoras Eigenmann, 1925: 327. Type species: Doras grypus Cope, 1872. Type by original designation. Gender:
Masculine.
Anadoras grypus (Cope, 1872)
Doras grypus Cope, 1872a: 270, pl. 15 (figs. 1, 1a). Type locality: small streams tributary to the Ambyiacu, as well
as ... the river itself ... which empties into the Amazon near Pebas, in Eastern Ecuador, some distance east of the
Napo. Syntypes: ANSP 8345 (1), ANSP 16460 (1).
Distribution: Upper Amazon River basin (Sabaj & Ferraris, 2003).
Anadoras regani (Steindachner, 1908)
Doras regani Steindachner, 1908f: 163. Type locality: Fischmerke von Pará [Amazonasgebiete innerhalb Brasiliens]. Syntypes: NMW 46889 (4).
Distribution: Amazon and Oyapock River basins (Sabaj & Ferraris, 2003).
Anadoras weddellii (Castelnau, 1855)
Doras weddellii Castelnau, 1855: 48, pl. 17 (fig. 1). Type locality: petites flaques d’eau de la province des Chiquitos.
Holotype: MNHN 0000-4155.
Distribution: Upper Mamoré, Paraguay and Pilcomayo River basins (Sabaj & Ferraris, 2003).
ANDUZEDORAS Fernández-Yépez, 1968
Anduzedoras Fernández-Yépez, 1968: 28. Type species: Anduzedoras arleoi Fernández-Yépez, 1968. Type by original designation. Gender: Masculine.
Anduzedoras oxyrhynchus (Valenciennes, 1821)
Doras Oxyrhynchus Valenciennes, in Humboldt & Valenciennes, 1821: 184. Type locality: Probablement ... originaires d’Amérique. Holotype: MNHN 0000-4190.
Doras (Corydoras) ophthalmus Kner, 1853c: 146, fig. 3 on unnumbered plate. Type locality: [Not stated]. Holotype:
NMW 45418-20 (1 of 4). Name made available from caption to illustration of a swimbladder; associated specimen is the holotype, if recognizable. Illustrated and described in more detail, as Doras (Oxydoras) lipophthalmus, in Kner (1855: 147, pl. 5, fig. 8), with locality given as: Rio negro.
Doras (Oxydoras) lipophthalmus Kner, 1855a: 147, pl. 5 (fig. 8). Type locality: Rio negro [Brazil]. Syntypes: NMW
45418-20 (4).
Opsodoras steindachneri Eigenmann, 1925: 354. Type locality: Mouth of Rio Negro [Brazil]. Holotype: NMW
46379.
Anduzedoras arleoi Fernández-Yépez, 1968: 29, fig. 9. Type locality: Río Autana, al este de Puerto Ayacucho, Terr.
Fed. Amazonas [Venezuela]. Holotype: FMNH 84068.
Distribution: Negro and upper Orinoco River basins, Brazil and Venezuela (Sabaj & Ferraris, 2003).
ASTRODORAS Bleeker, 1862
Astrodoras Bleeker, 1862 (in Bleeker, 1862–63): 5. Type species: Doras asterifrons Kner, 1853. Type by original

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designation. Gender: Masculine.
Astrodoras asterifrons (Kner, 1853)
Doras asterifrons Kner, 1853c: 146, fig. 4 on unnumbered plate. Type locality: not stated. Holotype: NMW 46412 (1of 2), NMW 46611 (1 of 6), NMW 58211 (1 of 3) or RMNH 2972 (1). Name made available from caption toillustration of a swimbladder; associated specimen is the holotype, if recognizable. Species later illustrated
described in more detail in Kner (1855: 123, pl. 2, fig. 2), with the locality given as: Barra do Rio negro und R.
Guaporé.
Distribution: Amazon River basin (Sabaj & Ferraris, 2003).
CENTROCHIR Spix & Agassiz, 1829
Centrochir Spix & Agassiz, 1829: 14. Type species: Doras crocodili Humboldt, 1821. Type by monotypy. Gender:
Feminine.
Centrochir crocodili (Humboldt, 1821)
Doras Crocodili Humboldt, in Humboldt & Valenciennes, 1821: 181, pl. 48 (fig. 2). Type locality: Rio
Magdalena...Nouvelle-Grenade...entre Pinto et Mompox, par les 9º et 9º1/2 de latitude...[&] près du confluent
du Rio Cauca [Colombia]. No types known.
Doras longispinis Steindachner, 1878a: 89. Type locality: Magdalenen-Stromes [Colombia]. Syntypes: NMW 46624(4). Species illustrated and described in more detail in Steindachner (1879d: 39, pl. 4, fig. 2, and pl. 5, fig. 1).
Distribution: Magdalena River basin, Colombia (Sabaj & Ferraris, 2003).
Remarks: Redescribed in Schmidt (1928).
CENTRODORAS Eigenmann, 1925
Centrodoras Eigenmann, 1925: 309. Type species: Doras brachiatus Cope, 1872. Type by original designation.
Gender: Masculine.
Centrodoras brachiatus (Cope, 1872)
Doras brachiatus Cope, 1872a: 270. Type locality: Between the mouth of the Rio Negro and the Huallaga, in the
Marañon. Holotype: ANSP 8342. Appeared first as name only in Cope (1871a: 55).
Rhinodoras amazonum Steindachner, 1875: 141, pl. 2. Type locality: Amazonenstrom bei Teffé [brasilianische].
Holotype: at NMW, but not found in 2006 (M. Sabaj, pers. commun.).
Distribution: Amazon River basin, except Negro River (Sabaj & Ferraris, 2003).
Centrodoras hasemani (Steindachner, 1915)
Oxydoras (Rhinodoras) amazonum hasemani Steindachner, 1915e: 68. Type locality: Mündung des Rio Negro.
Holotype: at NMW, but not found in 2006 (M. Sabaj, pers. comm.). Originally as Oxydoras (Rhinodoras) amazonum var. hasemani.
Distribution: Negro River basin, Brazil (Sabaj & Ferraris, 2003).
DORAOPS Schultz, 1944
Doraops Schultz, 1944c: 270. Type species: Doraops zuloagai Schultz, 1944. Type by original designation. Gender:
Masculine.
Doraops zuloagai Schultz, 1944
Doraops zuloagai Schultz, 1944c: 271, pl. 7 (fig. a). Type locality: Río de Los Pajaros, 3 km above Lago Maracaibo.
Holotype: USNM 121015.
Distribution: Lake Maracaibo basin including lake itself and Apon, Santa Ana, Catatumbo, Escalante and de Los
Pajaros River basins, Colombia (Sabaj & Ferraris, 2003).
DORAS La Cepède, 1803
Doras La Cepède, 1803: 116. Type species: Silurus carinatus Linnaeus, 1766. Type by subsequent designation by
Bleeker (1862–63: 5). Gender: Masculine.
Mormyrostoma Miranda Ribeiro, 1911: 192. Type species: Silurus carinatus Linnaeus, 1766. Type by original designation. Gender: Neuter.

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FERRARISDoras carinatus (Linnaeus, 1766)
Silurus carinatus Linnaeus, 1766: 504. Type locality: Surinami. Type(s): Whereabouts unknown.
Distribution: Essequibo River basin and other coastal drainages east to mouth of Amazon River, possibly in lower
Amazon and lower Orinoco River basins (Sabaj & Ferraris, 2003).
Doras micropoeus (Eigenmann, 1912)
Hemidoras micropoeus Eigenmann, 1912b: 195. Type locality: Wismar, British Guiana. Holotype: CM 1636
(whereabouts unknown).
Distribution: Essequibo, Demerara and Corantijn River basins, possibly other coastal drainages of northern SouthAmerica above mouth of Amazon River (Sabaj & Ferraris, 2003).
FRANCISCODORAS Eigenmann, 1925
Franciscodoras Eigenmann, 1925: 317. Type species: Doras marmoratus Reinhardt, 1874. Type by original designation. Gender: Masculine.
Franciscodoras marmoratus (Reinhardt, 1874)
Doras marmoratus Reinhardt, in Lütken, 1874c: 30. Type locality: in flumine Rio das Velhas [in provincia Minasgeraës circa oppidulum Lagoa Santa, Brasiliae centralis]. Holotype: ZMUC P 29669. Illustrated and describedin Lütken (1875: 146 (and p. III of summary), pl. 1, fig. 1).
Distribution: São Francisco River basin, Brazil (Sabaj & Ferraris, 2003).
HASSAR Eigenmann & Eigenmann, 1888
Hassar Eigenmann & Eigenmann, 1888b: 158. Type species: Oxydoras orestes [= Oxydoras orestis] Steindachner,
1875. Type by subsequent designation by Eigenmann (1910: 394). Gender: Masculine. Originally proposed as asubgenus of Hemidoras.
Hassar affinis (Steindachner, 1881)
Oxydoras affinis Steindachner, 1881c: 107, pl. 1 (fig. 1). Type locality: Rio Puty [Brazil]. Holotype: NMW 45394.
Originally as: Oxydoras affinis ? (an Oxyd. Orestes var.?).
Hassar woodi Fowler, 1941a: 139, figs. 35–37. Type locality: Rio Parnahyba, Therezina, Piauhy [Brazil]. Holotype:
ANSP 69392.
Hassar iheringi Fowler, 1941a: 140, figs. 38–40. Type locality: Rio Parnahyba, Therezina, Piauhy [Brazil]. Holotype: ANSP 69393.
Distribution: Parnaíba River basin, Brazil (Sabaj & Ferraris, 2003).
Hassar orestis (Steindachner, 1875)
Oxydoras Orestis Steindachner, 1875: 138, pl. 1. Type locality: Rio Xingu (bei den Wasserfällen) und Rio Iça [brasilianische]. Syntypes: NMW 45427–30 (7), NMW 78651 (1). Steindachner consistently used Oxydoras orestis
in 1875, but later changed the spelling to Oxydoras orestes, which had been widely adopted.
Hemidoras notospilus Eigenmann, 1912b: 196, pl. 19 (fig. 2). Type locality: Crab Falls, British Guiana. Holotype:
FMNH 53184 (missing).
? Hassar ucayalensis Fowler, 1940a: 228, figs. 15–17. Type locality: Ucayali River basin, Contamana, Peru. Holotype: ANSP 68647.
Distribution: Amazon, Orinoco and Essequibo River basins (Sabaj & Ferraris, 2003).
Remarks: Inclusion of Hassar ucayalensis Fowler and, therefore, the distribution of the species into Peru must be
considered tentative (Sabaj & Ferraris, 2003).
Hassar wilderi Kindle, 1895
Hassar wilderi Kindle, 1895: 251. Type locality: Trocera, on Tocantins, Brazil. Lectotype: CAS 60711; designatedby, and illustrated in, Eigenmann (1925: 364, pl. 22, fig. 2); subsequent lectotype designation in Myers &Weitzman (1956: 4) not valid.
Distribution: Tocantins River basin, Brazil (Sabaj & Ferraris, 2003).
HEMIDORAS Bleeker, 1858
Hemidoras Bleeker, 1858b: 53, 54. Type species: Doras (Oxydoras) stenopeltis Kner, 1855. Type by monotypy.

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Gender: Masculine.
Hemidoras morrisi Eigenmann, 1925
Hemidoras morrisi Eigenmann, 1925: 347, pl. 27 (fig. 3). Type locality: Iquitos. Holotype: CAS 52130.
Distribution: Upper Amazon River basin (Sabaj & Ferraris, 2003).
Remarks: Status uncertain, possibly a junior synonym of Opsodoras boulengeri (Steindachner) (Sabaj & Ferraris,2003).
Hemidoras stenopeltis (Kner, 1855)
Doras (Oxydoras) stenopeltis Kner, 1855a: 142, pl. 4 (fig. 7). Type locality: Rio negro. Syntypes: NMW 45443 (2).
Distribution: Amazon River basin (Sabaj & Ferraris, 2003).
HYPODORAS Eigenmann, 1925
Hypodoras Eigenmann, 1925: 329. Type species: Hypodoras forficulatus Eigenmann, 1925. Type by original designation. Gender: Masculine.
Hypodoras forficulatus Eigenmann, 1925
Hypodoras forficulatus Eigenmann, 1925: 330, pl. 25 (fig. 3). Type locality: Iquitos. Holotype: CAS 37246.
Distribution: Upper Amazon River basin, Peru (Sabaj & Ferraris, 2003).
KALYPTODORAS Higuchi, Britski & Garavello, 1990
Kalyptodoras Higuchi, Britski & Garavello, 1990: 220. Type species: Kalyptodoras bahiensis Higuchi, Britski &Garavello, 1990. Type by original designation. Gender: Masculine.
Kalyptodoras bahiensis Higuchi, Britski & Garavello, 1990
Kalyptodoras bahiensis Higuchi, Britski & Garavello, 1990: 222, fig. 1. Type locality: Brazil, Bahia State, Rio Paraguaçu, lower course, downstream from Bananeiras dam, portion just before construction site of Pedra do Cavalodam (12º32'05"S, 38º29'30"W to 12º35'50"S, 39º05'15"W). Holotype: MZUSP 38565.
Distribution: Paraguaçu River in Bahia State, Brazil (Sabaj & Ferraris, 2003).
LEPTODORAS Boulenger, 1898
Leptodoras Boulenger, 1898b: 477. Type species: Oxydoras acipenserinus Günther, 1868. Type by subsequent designation by Eigenmann (1910: 395). Gender: Masculine.
Taxonomy: Böhlke (1970).
Revision, with key to species: Sabaj (2005).
Leptodoras acipenserinus (Günther, 1868)
Oxydoras acipenserinus Günther, 1868a: 475. Type locality: Xeberos [Upper Amazons]. Holotype: BMNH
1867.6.13.32. Species illustrated and described in more detail in Günther (1868b: 230, pl. 20).
Distribution: Upper Amazon River basin, Brazil, Ecuador, and Peru (Sabaj, 2005).
Leptodoras cataniai Sabaj, 2005
Leptodoras cataniai Sabaj, 2005: 667, fig. 22. Type locality: Venezuela, Amazonas, río Casiquiare from mouth of
río Pamoni to 4 km below mouth, west of Pamoni (2º48'N, 65º57'W). Holotype: ANSP 180918.
Distribution: Rio Negro and Casiquiare Canal in Brazil and Venezuela (Sabaj, 2005).
Leptodoras copei (Fernández-Yépez, 1968)
Anduzedoras copei Fernández-Yépez, 1968: 31, fig. 10. Type locality: una laguna al lado del Río Capanaparo [Venezuela]. Holotype: FMNH 84069.
Distribution: upper Orinoco, Negro, and Amazon River basins, Brazil, Peru and Venezuela (Sabaj, 2005).
Leptodoras hasemani (Steindachner, 1915)
Hemidoras hasemani Steindachner, 1915b: 218. Type locality: Amazonas, Südamerikas. Lectotype: NMW 46381(82 mm SL, largest of 4 specimens); lectotype designated by Sabaj (2005: 654). Species illustrated anddescribed in more detail in Steindachner (1915e: 75, pl. 10, figs. 4–7), with locality as: des Rio branco bei Boa
Vista und Serra grande und an der Mündung des Rio negro.
Distribution: Branco, Orinoco and Essequibo River basins (Sabaj, 2005).
Leptodoras juruensis Boulenger, 1898

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FERRARISLeptodoras juruensis Boulenger, 1898b: 478. Type locality: Jurua River, Brazil. Holotype: BMNH 1898.10.11.25.
Distribution: Main channel of Amazon River basin (Sabaj, 2005).
Leptodoras linnelli Eigenmann, 1912
Leptodoras linnelli Eigenmann, 1912b: 191, pl. 17, fig. 1; pl. 18, fig. 1. Type locality: Tumatumari [lower Potaro
River], British Guiana. Holotype: FMNH 53561.
Distribution: Upper Orinoco (Mavaca River), Essequibo and Demerara River basins and northern (left bank) tributaries to lower Amazon (Sabaj, 2005).
Leptodoras myersi Böhlke, 1970
Leptodoras myersi Böhlke, 1970: 54, fig. 1. Type locality: Peru, vicinity of Iquitos, Rio Amazonas (Marañon)
between Isla Iquitos and Isla Lapuna, near Isla Lapuna shore. Holotype: ANSP 112318.
Distribution: Upper Amazon River basin, near Iquitos, Peru (Sabaj, 2005).
Leptodoras nelsoni Sabaj, 2005
Leptodoras nelsoni Sabaj, 2005: 663, fig. 20. Type locality: Colombia, Meta, río Metica (Meta-Orinoco dr.), ca. 1.5
km east of Rajote, Plancha 267 (3º56'N, 73º3'W). Holotype: ANSP 180917.
Distribution: middle Orinoco River basin, Colombia and Venezuela (Sabaj, 2005).
Leptodoras praelongus (Myers & Weitzman, 1956)
Hassar praelongus Myers & Weitzman, 1956: 2, fig. 1. Type locality: São Gabriel Rapids of the Rio Negro, Amazonas, Brazil. Holotype: SU 48679.
Distribution: Negro and Orinoco River basins, Brazil and Venezuela (Sabaj, 2005).
Leptodoras rogersae Sabaj, 2005
Leptodoras rogersae Sabaj, 2005: 665, fig. 21. Type locality: Venezuela, Bolivar, río Orinoco (Atlantic Dr.), river
channel at Palua just below mouth of río Caroni, 181 nautical miles upstream from sea buoy (8º21'N, 62º43'W).
Holotype: USNM 226307.
Distribution: Lower to middle Orinoco River, primarily in main channel (Sabaj, 2005).
LITHODORAS Bleeker, 1862
Lithodoras Bleeker, 1862 (in Bleeker, 1862–63): 5. Type species: Doras lithogaster Kner, 1855. Type by original
designation. Gender: Masculine.
Lithodoras dorsalis (Valenciennes, 1840)
Doras dorsalis Valenciennes, in Cuvier & Valenciennes, 1840b: 284 (211 in Strasbourg deluxe edition). Type locality: [Not stated]. Holotype: MNHN 0000-4156. On Doras carinatus of Valenciennes (in Humboldt & Valenciennes, 1821: p. 184).
Doras papilionatus De Filippi, 1853: 167. Type locality: in flumine Amazonum. Holotype: MZUT 229.
Doras lithogaster Kner, 1855a: 132. Type locality: Forte do Rio branco. Syntypes (2): at NMW, but not found in2006 (M. Sabaj, pers. commun.).
Megalodoras paucisquamatus Van der Stigchel, 1946: 71, fig. 1. Type locality: Brazil. Holotype: RMNH 15480.
Distribution: Amazon River basin and estuary near Cayenne, French Guiana (Keith & Merona, 2000; Sabaj & Ferraris, 2003).
MEGALODORAS Eigenmann, 1925
Megalodoras Eigenmann, 1925: 306. Type species: Megalodoras irwini Eigenmann, 1925. Type by original designation. Gender: Masculine.
Hoplodoras Eigenmann, 1925: 310. Type species: Doras uranoscopus Eigenmann & Eigenmann, 1888. Type byoriginal designation. Gender: Masculine. Spelled Haplodoras on p. 311, Hoplodaros on fig. 12c, and Hoplodoras in key and main account. First reviser apparently Eschmeyer & Bailey, in Eschmeyer (1990).
Deltadoras Fernández-Yépez, 1968: 36. Type species: Deltadoras guayoensis Fernández-Yépez, 1968. Type byoriginal designation. Gender: Masculine.
Remarks: Sabaj & Ferraris (2003: 462) served as first reviser in selecting Megalodoras as valid over Hoplodoras.
Megalodoras guayoensis (Fernández-Yépez, 1968)
Deltadoras guayoensis Fernández-Yépez, 1968: 37, fig. 14. Type locality: Caño de Guayo, Territorio Delta Ama-

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Zootaxa 1418 © 2007 Magnolia Press · 173curo, Venezuela. Holotype: MBUCV V-15211.
Hoplodoras ramirezi Fernández-Yépez, 1968: 47, fig. 19. Type locality: bajo Orinoco [Venezuela]. Holotype: AFY
66117 (missing).
Distribution: Orinoco River basin, Venezuela (Sabaj & Ferraris, 2003).
Remarks: Sabaj & Ferraris (2003: 462) served as first reviser in selecting Deltadoras guayoensis as the valid namefor this species.
Megalodoras uranoscopus (Eigenmann & Eigenmann, 1888)
Doras uranoscopus Eigenmann & Eigenmann, 1888b: 159. Type locality: Lake Hyanuary [Brazil]. Holotype: MCZ
7235.
? Oxydoras (Rhinodoras) huberi Steindachner, 1911a: 324. Type locality: Rio Tocantins, bei Cametá [Amazonasgebiete]. Holotype: NMW 12615.
Doras libertatis Miranda Ribeiro, 1912: 20. Type locality: Manáos [Brazil]. Holotype (112 mm): MNRJ 718 (missing). Holotype illustrated in Eigenmann (1925: pl. 7).
Megalodoras irwini Eigenmann, 1925: 308, pl. 25 (fig. 2). Type locality: Iquitos. Holotype: CAS 80243. Species
account appears twice in paper; the first (p. 307) based on a single specimen (CAS 20735) that was apparently
intended as the holotype of another new species description, which was changed before publication, except for
word “holotype”in the species account, which was first treated as lapsus calami by Sabaj & Ferraris (2003).
Distribution: Amazon, Tocantins and Essequibo River basins (Sabaj & Ferraris, 2003).
NEMADORAS Eigenmann, 1925
Nemadoras Eigenmann, 1925: 359. Type species: Oxydoras elongatus Boulenger, 1898. Type by original designation. Gender: Masculine.
Nemadoras elongatus (Boulenger, 1898)
Oxydoras elongatus Boulenger, 1898e: 424, pl. 40 (fig. 4). Type locality: Rio Jurua, an affluent of the Amazons,Brazil. Holotype: BMNH 1897.12.1.47.
Opsodoras parallelus Eigenmann, 1925: 350, pl. 19 (fig. 3). Type locality: Iquitos. Holotype: CAS 60229.
Distribution: Amazon River basin (Sabaj & Ferraris, 2003).
Remarks: Holotype of Oxydoras elongatus Boulenger was apparently registered twice, with the second registration
number, BMNH 1971.4.13.1, appearing in several recent publications.
Nemadoras hemipeltis (Eigenmann, 1925)
Opsodoras hemipeltis Eigenmann, 1925: 349, pl. 19 (fig. 2). Type locality: Rio Ucayali at Contamana. Holotype:
CAS 60236.
Distribution: Amazon River basin (Sabaj & Ferraris, 2003).
Nemadoras humeralis (Kner, 1855)
Doras humeralis Kner, 1855a: 140, pl. 4 (fig. 6). Type locality: Barra do Rio negro. Syntypes: NMW 45410 (2).
Oxydoras bachi Boulenger, 1898e: 423, pl. 40 (fig. 3). Type locality: Rio Jurua, an affluent of the Amazons, Brazil.
Holotype: BMNH 1897.12.1.46.
Distribution: Amazon River basin (Sabaj & Ferraris, 2003).
Nemadoras leporhinus (Eigenmann, 1912)
Hemidoras leporhinus Eigenmann, 1912b: 195, pl. 19 (fig. 1). Type locality: Tumatumari [lower Potaro River], British Guiana. Holotype: FMNH 53185.
Distribution: Orinoco, Branco and Essequibo River basins (Sabaj & Ferraris, 2003).
Nemadoras trimaculatus (Boulenger, 1898)
Oxydoras trimaculatus Boulenger, 1898e: 422, pl. 40 (fig. 1). Type locality: Rio Jurua, an affluent of the Amazons,Brazil. Syntypes: BMNH 1897.12.1.41–43 (3).
Leptodoras trimaculatus Fowler, 1914: 264, fig. 14. Type locality: Rupununi River, British Guiana ... in the highlands of British Guiana, approximately secured in North Latitude 2º to 3º, and West Longitude 50º20'. Holotype:
ANSP 39342. Originally considered by Fowler (1958) to have been preoccupied in Opsodoras by Opsodoras
trimaculatus (Boulenger, 1898); and replaced by Opsodoras ogilviei Fowler, 1958. More recently, considered to
be preoccupied in Nemadoras (Sabaj & Ferraris, 2003).

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FERRARISOpsodoras ogilviei Fowler, 1958: 13. Type locality: Rupununi River, British Guiana. Holotype: ANSP 39342.
Replacement for Leptodoras trimaculatus Fowler, 1914; then preoccupied in Opsodoras by Opsodoras trimaculatus (Boulenger, 1898).
Distribution: Amazon, Orinoco and Essequibo River basins (Sabaj & Ferraris, 2003).
OPSODORAS Eigenmann, 1925
Opsodoras Eigenmann, 1925: 348. Type species: Opsodoras orthacanthus Eigenmann, 1925. Type by original designation. Gender: Masculine.
Opsodoras boulengeri (Steindachner, 1915)
Hemidoras boulengeri Steindachner, 1915b: 218. Type locality: Mündung des Rio negro. Holotype: at NMW, but
not found in 2006 (M. Sabaj, pers. comm.). Species illustrated and described in more detail, as Hemidoras (Leptodoras) boulengeri, in Steindachner (1915e: 63, pl. 8, figs. 1–3).
Distribution: Amazon River basin, Bolivia and Brazil (Sabaj & Ferraris, 2003).
Opsodoras morei (Steindachner, 1881)
Oxydoras Morei Steindachner, 1881c: 106, pl. 1 (fig. 2). Type locality: Rio Negro. Holotype: NMW 45433.
Distribution: Negro River basin, Brazil (Sabaj & Ferraris, 2003).
Opsodoras stuebelii (Steindachner, 1882)
Oxydoras Stübelii Steindachner, 1882b: 175. Type locality: Aus dem Huallaga. Syntypes (3): at NMW, but not foundin 2006 (M. Sabaj, pers. commum.). Species illustrated and described in more detail in Steindachner (1882c: 5,
pl. 3, figs. 1–1b).
Opsodoras orthacanthus Eigenmann, 1925: 351, pl. 22 (fig. 3). Type locality: Iquitos. Holotype: CAS 60231.
Distribution: Amazon River basin (Sabaj & Ferraris, 2003).
Opsodoras ternetzi Eigenmann, 1925
Opsodoras ternetzi Eigenmann, 1925: 353. Type locality: Tapajos at Santarem. Holotype: CAS 60230.
Distribution: Amazon River basin (Sabaj & Ferraris, 2003).
ORINOCODORAS Myers, 1927
Orinocodoras Myers, 1927: 124. Type species: Orinocodoras eigenmanni Myers, 1927. Type by original designation. Gender: Masculine.
Orinocodoras eigenmanni Myers, 1927
Orinocodoras eigenmanni Myers, 1927: 124. Type locality: Venezuela: Caño de Quiribana near Caicara. Holotype:SU 58747.
Distribution: Orinoco River basin, Venezuela (Sabaj & Ferraris, 2003).
OXYDORAS Kner, 1855
Oxydoras Kner, 1855a: 115. Type species: Doras niger Valenciennes, 1821. Type by subsequent designation by
Eigenmann & Eigenmann (1890: 246). Gender: Masculine. Designation of Oxydoras kneri Bleeker, 1862[=Doras (Oxydoras) niger of Kner, 1855a] as type species in Bleeker (1862–63: 5) is invalid because O. kneri
was not an available name in Kner (1855).
Pseudodoras Bleeker, 1858b: 53, 54. Type species: Doras niger Valenciennes, 1821. Type by subsequent designation by Jordan (1919a: 278). Gender: Masculine.
Hildadoras Fernández-Yépez, 1968: 41. Type species: Hildadoras orinocensis Fernández-Yépez, 1968. Type byoriginal designation. Gender: Masculine.
Oxydoras kneri Bleeker, 1862
Oxydoras kneri Bleeker, 1862 (in Bleeker, 1862–63): 5. Type locality: Cujaba. Holotype: at NMW. Based on Doras(Oxydoras) niger of Kner (1855), from Cujaba.
Distribution: Paraná River basin (Sabaj & Ferraris, 2003).
Oxydoras niger (Valenciennes, 1821)
Doras niger Valenciennes, in Humboldt & Valenciennes, 1821: 184. Type locality: Probablement ... Originairesd’Amérique. Holotype: MNHN 0000-4189.

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? Corydoras edentatus Spix, in Spix & Agassiz, 1829: pl. 5. Type locality: [Not stated]. Holoype: Whereabouts
unknown (Kottelat, 1988). Name available from plate, with illustrated specimen the holotype. Corresponds to
Doras humboldti Spix & Agassiz, 1829.
? Doras humboldti Spix & Agassiz, 1829: 14, pl. 5. Type locality: fluvio S. Francisci mediae Brasiliae. Syntypes:
Whereabouts unknown (Kottelat, 1988).
Rhinodoras prianomus Cope, 1874b: 134. Type locality: Nauta [upper Amazon]. Holotype: ANSP 21203.
Rhinodoras teffeanus Steindachner, 1875: 145, pl. 3. Type locality: Teffé am Amazonenstrom [brasilianische]. Syntypes: NMW 44568–69 (2).
? Oxydoras holdeni Fernández-Yépez, 1968: 54, fig. 24. Type locality: Río Apure, frente a Mango Verde [Venezuela]. Holotype: AFY 51265 (missing).
Distribution: Amazon, São Francisco and Essequibo River basins, possibly Orinoco River basin, (Sabaj & Ferraris,2003).
Remarks: Synonymy based on Sabaj & Ferraris (2003). Inclusion of Doras humboldti and Corydoras edentatus insynonymy tentative, in the absence of types. Agassiz (1831: conspectus) serves as first reviser in treating Doras
humboldti as valid over Corydoras edentatus.
Oxydoras sifontesi Fernández-Yépez, 1968
Oxydoras sifontesi Fernández-Yépez, 1968: 58, fig. 28. Type locality: Laguna rebalsera “El Medio,” Ciudad Bolívar[Venezuela]. Holotype: MBUCV V-1584.
Hildadoras orinocensis Fernández-Yépez, 1968: 41, fig. 17. Type locality: Boca del caño San Juan, Terr. Delta
Amacuro [Venezuela]. Holotype: MBUCV V-3939.
Distribution: Orinoco River basin (Sabaj & Ferraris, 2003).
Remarks: Sabaj & Ferraris (2003) acted as first reviser in giving precedence to O. sifontesi.
PHYSOPYXIS Cope, 1871
Physopyxis Cope, 1871b: 112. Type species: Physopyxis lyra Cope, 1871. Type by monotypy. Gender: Feminine.
More complete description in Cope (1872a: 273).
Revision, with key to species: de Sousa & Rapp Py-Daniel (2005).
Physopyxis ananas de Sousa & Rapp Py-Daniel, 2005
Physopyxis ananas de Sousa & Rapp Py-Daniel, 2005: 630, fig. 5. Type locality: Jutaí River, Solimões River basin,
Amazonas State, Brazil (02º45'40''S; 66º47'59''W). Holotype: INPA 24324.
Distribution: Amazon (including Negro River) and Essequibo River basins (de Sousa & Rapp Py-Daniel, 2005).
Physopyxis cristata de Sousa & Rapp Py-Daniel, 2005
Physopyxis cristata de Sousa & Rapp Py-Daniel, 2005: 632, fig. 7. Type locality: mouth of igarapé Zamula, Negro
River, Amazonas State, Brazil (00º51'48''S; 62º46'44''W). Holotype: INPA 24325.
Distribution: Negro River basin (Amazon River drainage), Amazonas State, Brazil (de Sousa & Rapp Py-Daniel,
2005).
Physopyxis lyra Cope, 1871
Physopyxis lyra Cope, 1871b: 112. Type locality: Amazon. Holotype: ANSP 8282. Illustrated and described in moredetail in Cope (1872a: 273, pl. 5, figs. 1a–1c), with locality as Ambyiacu River, Ecuador [now Peru].
Distribution: Amazon River basin, Peru and Brazil (de Sousa & Rapp Py-Daniel, 2005).
PLATYDORAS Bleeker, 1862
Platydoras Bleeker, 1862 (in Bleeker, 1862–63): 5. Type species: Silurus costatus Linnaeus, 1758. Type by original
designation. Gender: Masculine.
Platydoras armatulus (Valenciennes, 1840)
Doras armatulus Valenciennes, in Cuvier & Valenciennes, 1840b: 273 (204 in Strasbourg deluxe edition). Type
locality: Brésil, ... le Parana, mais non au-dessous du 27º30' de latitude sud. Holotype: MNHN 0000-4152.
Distribution: Paraná River basin (Sabaj & Ferraris, 2003).
Platydoras costatus (Linnaeus, 1758)
Silurus costatus Linnaeus, 1758: 306. Type locality: Indiis. Type(s): Whereabouts unknown. Based on account in

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Gronovius (1756: 24–25, pl. 5, figs. 1 & 2).
Mystus ascita Gronow, in Gray, 1854: 156. Type locality: Not stated. Type(s): Whereabouts unknown. Based on
account in Gronovius (1756: 24–25, pl. 5, figs. 1 & 2). Preoccupied by Mystus ascita Walbaum (1792); apparently not replaced.
? Doras dentatus Kner, 1855a: 118, pl. 3 (fig. 3). Type locality: Surinam. Holotype: NMW 46869. Originally proposed as Doras dentatus?
? Doras helicophilus Günther, 1868a: 475. Type locality: Surinam [Maroni River]. Syntypes: BMNH 1866.8.10.1–3(3).
Distribution: Amazon, Tocantins, Parnaíba, Orinoco, Essequibo River basins and coastal drainages in French Guiana
and Suriname (Sabaj & Ferraris, 2003).
Species inquirenda, Platydoras
Doras Hancockii Valenciennes, in Cuvier & Valenciennes, 1840b: 279 (207 in Strasbourg deluxe edition). Type locality:
not stated. Holotype: BMNH 1857.6.13.163. Based on Doras costata of Hancock (1828), from Demerara [Guyana].
PTERODORAS Bleeker, 1862
Pterodoras Bleeker, 1862 (in Bleeker, 1862–63): 5. Type species: Doras granulosus Valenciennes, 1821. Type byoriginal designation. Gender: Masculine.
Apuredoras Fernández-Yépez, 1950c: 195. Type species: Apuredoras rivasi Fernández-Yépez, 1950. Type by original designation. Gender: Masculine.
Parapterodoras Risso & Morra, 1964: 1. Type species: Parapterodoras paranensis Risso & Morra, 1964. Type byoriginal designation. Gender: Masculine.
Sachsdoras Fernández-Yépez, 1968: 66. Type species: Sachsdoras apurensis Fernández-Yépez, 1968. Type by original designation. Gender: Masculine.
Pterodoras granulosus (Valenciennes, 1821)
Doras granulosus Valenciennes, in Humboldt & Valenciennes, 1821: 184. Type locality: Probablement .... Originaires d’Amérique. Holotype: MNHN 0000-4187.
? Doras maculatus Valenciennes, 1836, (in Valenciennes, 1835–47): pl. 5 (fig. 3). Type locality: not stated. Holotype: MNHN b-0201 (1 of 2). Name available from plate that, according to Valenciennes (in Cuvier & Valenciennes) 1840b: 281 (210 in Strasbourg deluxe edition), was based on single specimen, but description in Cuvier& Valenciennes (1840b: 281 (209–211 in Strasbourg deluxe edition) based on holotype (MNHN 0000-4187) ofDoras granulosus Valenciennes, 1821, and two specimens (MNHN b-0201) with locality as Buénos-Ayres.
? Doras murica Kner, 1855a: 129. Type locality: Cujaba. Holotype: at NMW. Name credited to Natterer; however,availability of name appears attributable solely to Kner.
? Doras laevigatulus Berg, 1901: 298. Type locality: Dock Sud, que comunica con el Río de la Plata y el Riachuelode la Boca. Holotype: MACN 2118.
? Doras lentiginosus Eigenmann, 1917a: 401, pl. 40. Type locality: Santarem [Brazil]. Holotype: FMNH 58056.
? Silurus 12-radiatus Larrañaga, 1923: 386. Type locality: [Uruguay]. No types known.
? Silurus armatus Larrañaga, 1923: 377. Type locality: [Uruguay]. No types known. Originally as Silurus (armado)
armatus.
? Parapterodoras paranensis Risso & Morra, 1964: 2, pl. 1. Type locality: Río Paraná, frente a Corrientes. Holotype: Mus. Cien. Nat. del Chaco X-64-1.
Distribution: Amazon and Paraná River basins and coastal drainages in Guyana and Suriname (Sabaj & Ferraris,2003).
Pterodoras rivasi (Fernández-Yépez, 1950)
Apuredoras rivasi Fernández-Yépez, 1950c: 195, fig. on p. 196. Type locality: Río Apure, 2 km al este de San
Fernando. Holotype: AFY 51062 (whereabouts unknown).
Pterodoras angeli Fernández-Yépez, 1968: 65, fig. 32. Type locality: Río Arauca, Edo. Apure [Venezuela]. Holo-CHECKLIST OF CATFISHES, RECENT AND FOSSIL

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type: AFY 66803 (whereabouts unknown). Name also spelled Ptedoras angelis in original description. Pterodoras angeli selected as the valid name by Sabaj & Ferraris (2003: 465).
Sachsdoras apurensis Fernández-Yépez, 1968: 66, fig. 33. Type locality: Río Orinoco, cerca de la boca del Río
Apure [Venezuela]. Holotype: AFY 66802 (whereabouts unknown).
Distribution: Orinoco River basin, Colombia and Venezuela (Sabaj & Ferraris, 2003).
RHINODORAS Bleeker, 1862
Rhinodoras Bleeker, 1862 (in Bleeker, 1862–63): 5. Type species: Doras (Oxydoras) dorbignyi Kner, 1855. Type byoriginal designation. Gender: Masculine.
Rhinodoras boehlkei Glodek, Whitmire & Orcés, 1976
Rhinodoras boehlkei Glodek, Whitmire & Orcés, 1976: 3, fig. 1. Type locality: Eastern Ecuador, Río Bobonaza
between Montalvo and Chicherato. Holotype: FMNH 79203.
Distribution: Upper Amazon River basin (Sabaj & Ferraris, 2003).
Rhinodoras dorbignyi (Kner, 1855)
Doras (Oxydoras) d’Orbignyi Kner, 1855a: 149, pl. 5 (fig. 9). Type locality: Rio de la Plata. Holotype: NMW
44555. Name also spelled Doras (Oxydoras) d’Orbigny in original description. Rhinodoras dorbignyi selected
as the valid name over Rhinodoras dorbigny by Sabaj & Ferraris (2003: 465). Name credited to Krøyer inHeckel manuscript; however, availability of name appears attributable solely to Kner.
Doras nebulosus Eigenmann & Kennedy, 1903: 500. Type locality: either in Mato Grosso [at Descalvados, Brazil]
or [in the Paraguay at] Asuncion. Holotype: CAS 31186.
Distribution: Paraná River basin (Sabaj & Ferraris, 2003).
Remarks: Catalog number of holotype of Doras (Oxydoras) d’Orbignyi incorrectly reported as NMW 44551 in
Eschmeyer et al. (1998) and Sabaj & Ferraris (2003).
Rhinodoras thomersoni Taphorn & Lilyestrom, 1984
Rhinodoras thomersoni Taphorn & Lilyestrom, 1984a: 87, fig. 1. Type locality: Venezuela noroccidental. Cuencadel Lago Maracaibo, Estado Zulia, desembocadura del Río Catatumbo, cerca de El Congo. Holotype: MCNG
368.
Distribution: Santa Ana, Catatumbo and Escalante river basins, along southwest shore of Lake Maracaibo (Sabaj &Ferraris, 2003).
RHYNCHODORAS Klausewitz & Rössel, 1961
Rhynchodoras Klausewitz & Rössel, 1961: 45. Type species: Rhynchodoras xingui Klausewitz & Rössel, 1961.
Type by original designation. Gender: Masculine.
Rhynchodoras woodsi Glodek, 1976
Rhynchodoras woodsi Glodek, 1976: 44, fig. 1. Type locality: Eastern Ecuador, Moreta Bobonaza, between Sarayacu and Montalvo. Río Bobonaza, a tributary of Río Pastaza, itself a tributary of Río Marañon in Peru. Holotype: FMNH 77008.
Distribution: Pastaza River basin in Marañon River drainage, Ecuador (Sabaj & Ferraris, 2003).
Rhynchodoras xingui Klausewitz & Rössel, 1961
Rhynchodoras xingui Klausewitz & Rössel, 1961: 46, fig. 1. Type locality: Südamerika, Brasilien, Oberlauf des Rio
Xingu. Holotype: SMF 5281.
Distribution: Upper Xingu River basin, Brazil (Sabaj & Ferraris, 2003).
SCORPIODORAS Eigenmann, 1925
Scorpiodoras Eigenmann, 1925: 324. Type species: Doras heckelii Kner, 1855. Type by original designation. Gender: Masculine.
Autanadoras Fernández-Yépez, 1950a: 8. Type species: Autanadoras milesi Fernández-Yépez, 1950. Type by original designation. Gender: Masculine.
Scorpiodoras heckelii (Kner, 1855)
Doras Heckelii Kner, 1855a: 125, pl. 3 (fig. 4). Type locality: R. negro. Holotype: NMW 46613.

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FERRARISDoras Calderonensis Vaillant, 1880b: 154. Type locality: Caldéron (Haute-Amazone). Holotype: MNHN a-1980.
Autanadoras milesi Fernández-Yépez, 1950a: 8, pl. 1 (fig. 4). Type locality: Río Autana [Venezuela]. Holotype:
MBUCV V-15210.
Distribution: Negro, Orinoco, and upper Amazon River basins, Brazil and Venezuela (Sabaj & Ferraris, 2003).
TRACHYDORAS Eigenmann, 1925
Trachydoras Eigenmann, 1925: 337. Type species: Trachydoras atripes Eigenmann, 1925. Type by original designation. Gender: Masculine.
Trachydoras brevis (Kner, 1853)
Doras (Corydoras) brevis Kner, 1853c: 146, fig. 8 on unnumbered plate. Type locality: [Not stated]. Holotype:
NMW 45397–99 (1 of 5), NMW 46375 (1 of 2) or RMNH 2965 (1). Name made available from caption to illustration of a swimbladder; associated specimen is the holotype, if recognizable. Species later described and illustrated in Kner (1855a: 138–140, pl. 6, fig. 11), on 8 specimens with locality given as Barra do Rio negro; andswimbladder illustration in Kner (1853c) corrected from fig. 6 to fig. 8.
Distribution: Negro and Essequibo River basins (Sabaj & Ferraris, 2003).
Trachydoras microstomus (Eigenmann, 1912)
Hemidoras microstomus Eigenmann, 1912b: 193, pl. 18 (fig. 2). Type locality: Rockstone, British Guiana. Holotype:
FMNH 53206.
Distribution: Amazon, Orinoco and Essequibo River basins (Sabaj & Ferraris, 2003).
Trachydoras nattereri (Steindachner, 1881)
Oxydoras Nattereri Steindachner, 1881c: 104, pl. 2 (fig. 1). Type locality: Amazonen-Strome bei Teffe [Brazil].
Holotype: at NMW, but not found in 2006 (M. Sabaj, pers. commun.).
Distribution: Amazon River basin (Sabaj & Ferraris, 2003).
Trachydoras paraguayensis (Eigenmann & Ward, 1907)
Hemidoras paraguayensis Eigenmann & Ward in Eigenmann, McAtee & Ward, 1907: 116, pl. 34 (fig. 1). Type
locality: Paraguay River at ... Corumba [Brazil]. Holotype: CAS 37216.
Distribution: Paraná River basin (Sabaj & Ferraris, 2003).
Trachydoras steindachneri (Perugia, 1897)
Oxydoras Steindachneri Perugia, 1897: 20. Type locality: Rio Beni, Bolivia. Syntypes: MSNG 8834 (17).
Oxydoras trachyparia Boulenger, 1898e: 423, pl. 40 (fig. 2). Type locality: Rio Jurua, an affluent of the Amazons,Brazil. Syntypes: BMNH 1897.12.1.44–45 (2).
Trachydoras atripes Eigenmann, 1925: 339, pl. 26 (fig. 4). Type locality: Brook near River Itaya, above Iquitos[Peru]. Holotype: CAS 60234.
Distribution: Amazon River basin (Sabaj & Ferraris, 2003).
WERTHEIMERIA Steindachner, 1877
Wertheimeria Steindachner, 1877b: 659. Type species: Wertheimeria maculata Steindachner, 1877. Type by monotypy. Gender: Feminine.
Relationships: Higuchi et al. (1990).
Wertheimeria maculata Steindachner, 1877
Wertheimeria maculata Steindachner, 1877b: 660, pl. 10. Type locality: Rio Jequitinhonha [Brazil]. Syntypes:
NMW 43460 (1, lost), NMW 43461 (1).
Distribution: Jequitinhonha River basin, Brazil (Sabaj & Ferraris, 2003).
Species inquirenda, Doradidae
Doras insculptus Miranda Ribeiro, 1912: 22. Type locality: Manáos [Brazil]. Lectotype: MNRJ 656a, designated by
Miranda Ribeiro (1953: 398). Validity and generic placement considered uncertain in Sabaj & Ferraris (2003); likely
as Anadoras or Amblydoras.

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HEPTAPTERIDAE Gill, 1861
Pteronotidae Adams, in Adams et al., 1854: 107. Type genus: Pteronotus Swainson, 1839. Permanently invalid, because
Pteronotus Swainson is a junior homonym of Pteronotus Gray, 1837 (Mammalia).
Heptapterinae Gill, 1861c: 54. Type genus: Heptapterus Bleeker, 1858.
Rhamdiae Bleeker, 1862 (in Bleeker, 1862–63): 11, 60. Type genus: Rhamdia Bleeker, 1858.
Taxonomic summary: Gosline (1941), Bockmann & Guazzelli (2003).
Phylogeny: Lundberg & Mc Dade (1986), Ferraris (1988a, Nemuroglanis group); Bockmann (1994, 1998).
24 genera, 189 species; no named fossil taxa.
Incertae sedis
Imparales panamensis Bussing, 1970
Imparales panamensis Bussing, 1970: 7, fig. 2. Type locality: Panamá: Veraguas Prov., creek crossing road on s sideof Santa Fe (elev. 305 m). Holotype: USNM 204692.
Distribution: River basin(s) draining Veraguas Province, Tuira River basin, and Frijoles River, Panama (Bockmann& Guazzelli, 2003).
Remarks: Not related to Imparales, or its senior synonym Nemuroglanis (Bockmann & Ferraris, 2005), but notplaced elsewhere.
Pimelodus (Rhamdia) parvus Boulenger, 1898
Pimelodus (Rhamdia) parvus Boulenger, 1898c: 7. Type locality: du Rio Santiago, du Rio Zamora et du Rio Bomboiza. Syntypes: BMNH 1898.11.4.5–6, BMNH 1898.11.4.7, ZMUT 1543 (2), ZMUT 1544 (2), ZMUT 1691
(4).
Distribution: Marañon River basin, Ecuador.
Remarks: Silfvergrip (1996: 29) referred this species to Nannorhamdia, but Bockmann & Guazzelli (2003) disagreed with that placement and, instead, left the species unassigned.
ACENTRONICHTHYS Eigenmann & Eigenmann, 1889
Acentronichthys Eigenmann & Eigenmann, 1889b: 28. Type species: Acentronichthys leptos Eigenmann & Eigenmann, 1889. Type by original designation. Gender: Masculine.
Remarks: Treated as a synonym of Heptapterus by Miranda Ribeiro (1962b).
Acentronichthys leptos Eigenmann & Eigenmann, 1889
Acentronichthys leptos Eigenmann & Eigenmann, 1889b: 29. Type locality: Sao Matheos [=São Mateus, State of
Espírito Santo, Brazil]. Holotype: MCZ 7532.
Distribution: Coastal streams from Rio de Janeiro to Santa Catarina State, and in São Mateus, in Espírito Santo
State, Brazil (Bockmann & Guazzelli, 2003).
BRACHYGLANIS Eigenmann, 1912
Brachyglanis Eigenmann, 1912b: 156. Type species: Brachyglanis frenata Eigenmann, 1912. Type by original designation. Gender: Masculine.
Remarks: First published as Breviglanis in Eigenmann (1910: 384), but without diagnosis or available species; namewas apparently never made available by its use with an available species name.
Brachyglanis frenatus Eigenmann, 1912
Brachyglanis frenata Eigenmann, 1912b: 156. Type locality: Amatuk [Essequibo River basin], British Guiana.
Holotype: CM 1670, not found at FMNH.
Distribution: Essequibo, Negro, Orinoco, and Urubu River basins of Brazil, Guyana, and Venezuela (Bockmann &Guazzelli, 2003).
Brachyglanis magoi Fernández-Yépez, 1967
Brachyglanis magoi Fernández-Yépez, 1967: 162, pl. 1. Type locality: río Uruyén medio. Holotype: AFY 56611.

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FERRARISDistribution: Caroní River basin in Orinoco River drainage, Venezuela (Bockmann & Guazzelli, 2003).
Brachyglanis melas Eigenmann, 1912
Brachyglanis melas Eigenmann, 1912b: 157, pl. 11 (fig. 2). Type locality: Crab Falls, British Guiana. Holotype:
FMNH 53217.
Distribution: Essequibo River basin, Guyana (Bockmann & Guazzelli, 2003).
Brachyglanis microphthalmus Bizerril, 1991
Brachyglanis microphthalmus Bizerril, 1991: 2, fig. 1. Type locality: Brasil, Pará, Município de Oriximiná, Bacia do
rio Trombetas, drenagem do Lago Sapucuá, Igarapé Saracá. Holotype: MNRJ 11788.
Distribution: Trombetas River basin, Brazil (Bockmann & Guazzelli, 2003).
Brachyglanis nocturnus Myers, 1928
Brachyglanis nocturnus Myers, 1928: 84. Type locality: Rock-pools at São Gabriel Rapids, Rio Negro, Brazil. Holotype: CAS 63393.
Distribution: Negro River basin, Brazil (Bockmann & Guazzelli, 2003).
Brachyglanis phalacra Eigenmann, 1912
Brachyglanis phalacra Eigenmann, 1912b: 157, pl. 12 (fig. 1), pl. 13 (fig. 1). Type locality: Amatuk [Essequibo
River basin], British Guiana. Holotype: FMNH 53216.
Distribution: Essequibo River basin, Guyana (Bockmann & Guazzelli, 2003).
BRACHYRHAMDIA Myers, 1927
Brachyrhamdia Myers, 1927: 123. Type species: Brachyrhamdia imitator Myers, 1927. Type by original designation. Gender: Feminine.
Brachyrhamdia heteropleura (Eigenmann, 1912)
Pimelodus heteropleurus Eigenmann, 1912b: 176, pl. 16 (fig. 2). Type locality: Rupununi Pan [Essequibo Riverbasin], British Guiana. Holotype: FMNH 53336.
Distribution: Corantijn, Essequibo and Negro River basins (Bockmann & Guazzelli, 2003).
Brachyrhamdia imitator Myers, 1927
Brachyrhamdia imitator Myers, 1927: 123. Type locality: Caño de Quiribana, near Caicara [Venezuela]. Holotype:
CAS 54359.
Distribution: Caura River basin in Orinoco River drainage, Venezuela (Bockmann & Guazzelli, 2003).
Brachyrhamdia marthae Sands & Black, 1985
Brachyrhamdia marthae Sands & Black, 1985: 58 (4), illustrated on p. 58 (7). Type locality: Peru, aquarium import.
Holotype: RMNH 29424.
Distribution: Juruá and Mamoré/Madeira River basins (Bockmann & Guazzelli, 2003).
Remarks: Redescribed, as Rhamdia marthae, in Zarske (2003).
Brachyrhamdia meesi Sands & Black, 1985
Brachyrhamdia meesi Sands & Black, 1985: 58 (5), illustrated on p. 58 (7). Type locality: Near Belem, Brazil. Holotype: RMNH 29425.
Distribution: Amazon River basin, Brazil (Bockmann & Guazzelli, 2003).
Brachyrhamdia rambarrani (Axelrod & Burgess, 1987)
Pimelodella rambarrani Axelrod & Burgess, in Axelrod, 1987: 24, figures on pp. 24– 25. Type locality: reportedly
from the Rio Unini, a tributary of the Rio Negro, Amazonas, Brazil. Holotype: MZUSP 37693.
Distribution: Unini River basin in upper Negro River drainage, Brazil (Bockmann & Guazzelli, 2003).
CETOPSORHAMDIA Eigenmann & Fisher, 1916
Cetopsorhamdia Eigenmann & Fisher, in Eigenmann, 1916: 83. Type species: Cetopsorhamdia nasus Eigenmann &Fisher, 1916. Type by original designation. Gender: Feminine.
Cetopsorhamdia boquillae Eigenmann & Fisher, 1922
Cetopsorhamdia boquillae Eigenmann & Fisher, in Eigenmann, 1922b: 37, pl. 1 (fig. 3). Type locality: Boquilla[Colombia]. Holotype: FMNH 55212.
Distribution: Cauca River basin, Colombia (Maldonado-Ocampo et al., 2005).

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Cetopsorhamdia filamentosa Fowler, 1945
Cetopsorhamdia filamentosa Fowler, 1945c: 4, figs. 5–6. Type locality: San Ramón, Vallé de Chanchamayo, at 800
meters elevation, Peru. Holotype: ANSP 71637.
Distribution: Tulumayo River basin in upper Ucayali River drainage, Peru (Bockmann & Guazzelli, 2003).
Cetopsorhamdia iheringi Schubart & Gomes, 1959
Cetopsorhamdia iheringi Schubart & Gomes, 1959: 1, fig. 1. Type locality: Ribeirão Areia Branca, próximo à
estrada carroçável para Piraçununga, bacia do Rio Mogi Guaçu, Município de Descalvado [Brazil]. Holotype:
EEBP 368.
Distribution: Upper reaches of the Paraná and São Francisco River basins, Brazil (Bockmann & Guazzelli, 2003).
Cetopsorhamdia insidiosa (Steindachner, 1915)
Imparfinis insidiosus Steindachner, 1915e: 61, pl. 12 (fig. 7). Type locality: Rio branco bei Bem Querer und im Rio
Surumú, einem Nebenflusse des Rio Miang [Brazil]. Syntypes: NMW 46095.
Distribution: Branco River basin, Brazil (Bockmann & Guazzelli, 2003).
Cetopsorhamdia molinae Miles, 1943
Cetopsorhamdia molinae Miles, 1943a: 26. Type locality: Bugalagrande [Colombia]. Holotype: at ICNMHN probably lost (Cala, 1981: 1).
Cetopsorhamdia shermani Schultz, 1944c: 220, pl. 2 (fig. c). Type locality: Río Guárico and tributaries between San
Sebastián and San Casimiro (Orinoco system), Estado de Aragua, Venezuela. Holotype: USNM 121216.
Distribution: Magdalena, Orinoco and Tocantins River basins (Bockmann & Guazzelli, 2003); also Cauca River,Colombia (Maldonado-Ocampo et al., 2005).
Cetopsorhamdia nasus Eigenmann & Fisher, 1916
Cetopsorhamdia nasus Eigenmann & Fisher, in Eigenmann, 1916: 83. Type locality: Honda, Colombia. Holotype:
FMNH 58126; holotype illustrated in Eigenmann (1922b: 36, fig. 10; pl. 4, fig. 1).
Distribution: Magdalena and Cauca River basins, Colombia (Maldonado-Ocampo et al., 2005).
Cetopsorhamdia orinoco Schultz, 1944
Cetopsorhamdia orinoco Schultz, 1944c: 225, pl. 3 (fig. a). Type locality: Río Torbes, 1 km. above Táriba, Orinoco
system, Venezuela. Holotype: USNM 121214.
Distribution: Napo and Orinoco River basins (Bockmann & Guazzelli, 2003).
Cetopsorhamdia phantasia Stewart, 1985
Cetopsorhamdia phantasia Stewart, 1985a: 340, fig. 1. Type locality: Ecuador, Río Jivino, west of Limoncocha,
approx. 0º24.6'S, 76º39.0'W. Holotype: FMNH 94601.
Distribution: Napo River basin, Ecuador (Bockmann & Guazzelli, 2003).
Cetopsorhamdia picklei Schultz, 1944
Cetopsorhamdia picklei Schultz, 1944c: 222, pl. 2 (fig. d). Type locality: Río Motatán, 4 km. above Motatán, Maracaibo basin [Venezuela]. Holotype: USNM 121217.
Distribution: Maracaibo Lake basin, Venezuela (Bockmann & Guazzelli, 2003).
CHASMOCRANUS Eigenmann, 1912
Chasmocranus Eigenmann, 1912b: 160. Type species: Chasmocranus longior Eigenmann, 1912. Type by original
designation. Gender: Masculine.
Remarks: Named Chasmocephalus in Eigenmann (1910: 384) but without diagnosis or included available species;name was apparently never made available by its use with an available species name.
Chasmocranus brachynemus Gomes & Schubart, 1958
Chasmocranus brachynema Gomes & Schubart, 1958: 413, fig. 1. Type locality: Rio Mogi Guaçu, Cachoeira deEmas, Município de Piraçununga, Estado de São Paulo [Brazil]. Holotype: EEBP 617.
Distribution: Mogi Guaçu River basin in upper Paraná River drainage, Brazil (Bockmann & Guazzelli, 2003).
Chasmocranus brevior Eigenmann, 1912
Chasmocranus brevior Eigenmann, 1912b: 162, pl. 15 (fig. 1). Type locality: Waratuk, British Guiana. Holotype:
FMNH 53213.
Distribution: Mana and Maroni River basins, and Potaro River, Guianas (Bockmann & Guazzelli, 2003).

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Chasmocranus chimantanus Inger, 1956
Chasmocranus chimantanus Inger, 1956: 425, fig. 92. Type locality: Rio Abácapa on west side of Chimantá-tepui[Venezuela]. Holotype: FMNH 45704.
Distribution: Caroní and Carapo River basins, Orinoco River drainage, Venezuela (Bockmann & Guazzelli, 2003).
Chasmocranus longior Eigenmann, 1912
Chasmocranus longior Eigenmann, 1912b: 160, pl. 14 (fig. 2). Type locality: Amatuk, British Guiana. Holotype:
FMNH 53208.
Distribution: Caribbean coastal drainages from Guyana to Surinam; Negro, Orinoco and Xingu River basins (Bockmann & Guazzelli, 2003).
Chasmocranus lopezi Miranda Ribeiro, 1968
Chasmocranus Lopezi Miranda Ribeiro, 1968: 4, fig. 2. Type locality: Corrego Quente — Cubatão ? — Estado deSão Paulo — Brasil. Holotype: MNRJ 9705.
Distribution: São Paulo State, Brazil (Bockmann & Guazzelli, 2003).
Chasmocranus peruanus Eigenmann & Pearson, 1942
Chasmocranus peruanus Eigenmann & Pearson, in Eigenmann & Allen, 1942: 91, pl. 3 (figs. 3–4). Type locality:
Puerto Melendez below Pongo de Manseriche, Rio Marañon. Syntypes: CAS 63627 (2).
Distribution: Upper Amazon River basin, Peru (Bockmann & Guazzelli, 2003).
Chasmocranus quadrizonatus Pearson, 1937
Chasmocranus quadrizonatus Pearson, 1937: 93. Type locality: Tingo de Pauca, Rio Marañon, Peru. Holotype: CAS
63628.
Distribution: Marañon River basin, Peru (Bockmann & Guazzelli, 2003).
Chasmocranus rosae Eigenmann, 1919
Chasmocranes Rosae Eigenmann, 1919a: 127. Type locality: Río Negro, Villavicencio [Colombia]. Holotype:
FMNH 55140; holotype illustrated in Eigenmann (1922b: pl. 1, fig. 4).
Distribution: Meta River basin, Colombia (Bockmann & Guazzelli, 2003).
Chasmocranus surinamensis (Bleeker, 1862)
Heptapterus surinamensis Bleeker, 1862a: 387. Type locality: Surinama. Holotype: RMNH 2984; holotype illustrated in Bleeker (1864: 91, pl. 15, fig. 1).
Distribution: Suriname River basin (Bockmann & Guazzelli, 2003).
Remarks: Mees (1967: 226) interpreted the type locality as the Suriname River, presumably not far from Paramaribo(Bockmann & Guazzelli, 2003).
Chasmocranus truncatorostris Borodin, 1927
Chasmocranus truncatorostris Borodin, 1927b: 5, fig. 4. Type locality: Col. Hansa Joinville, Sta. Catharina Prov.,
Brazil. Holotype: AMNH 8640.
Distribution: Sta. Catharina Province, Brazil; known only from the type locality (Bockmann & Guazzelli, 2003).
GLADIOGLANIS Ferraris & Mago-Leccia, 1989
Gladioglanis Ferraris & Mago-Leccia, 1989: 166. Type species: Gladioglanis machadoi Ferraris & Mago-Leccia,
1989. Type by original designation. Gender: Masculine.
Gladioglanis conquistador Lundberg, Bornbusch & Mago-Leccia, 1991
Gladioglanis conquistador Lundberg, Bornbusch & Mago-Leccia, 1991: 192, fig. 7A. Type locality: Ecuador, Napo
Province, Río Napo basin, Río Aguarico sub-basin, north tributary of Río Cuyabeno, ca 3 km above Laguna
Grande de Cuyabeno, 0º1.5'S, 76º13.2'W. Holotype: FMNH 99288.
Distribution: Capim, Napo, Purus and Solimões Rivers, Amazon basin (Bockmann & Guazzelli, 2003).
Gladioglanis machadoi Ferraris & Mago-Leccia, 1989
Gladioglanis machadoi Ferraris & Mago-Leccia, 1989: 167, fig. 1. Type locality: Venezuela, Territorio Federal
Amazonas, Departamento Río Negro, Caño Manu of Casiquiare canal, approx. 250 m upstream of Solano,
2º00'N, 66º57'W. Holotype: MBUCV V-16000.
Distribution: Negro and Orinoco River basins, Brazil and Venezuela (Bockmann & Guazzelli, 2003).

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GOELDIELLA Eigenmann & Norris, 1900
Goeldiella Eigenmann & Norris, 1900: 353. Type species: Pimelodus eques Müller & Troschel, 1848. Type by original designation. Gender: Feminine.
Goeldiella eques (Müller & Troschel, 1848)
Pimelodus eques Müller & Troschel, in Schomburgk, 1848: 628. Type locality: in allen Flüssen Guiana’s. Holotype:
ZMB 3049. Also described as new in Müller & Troschel (1849: 5).
Distribution: Coastal Rivers of the Guianas and Amazon River basin (Bockmann & Guazzelli, 2003).
HEPTAPTERUS Bleeker, 1858
Heptapterus Bleeker, 1858b: 197, 208. Type species: Pimelodus mustelinus Valenciennes, 1835. Type by monotypy.
Gender: Masculine.
Review: Buckup (1988, southern Brazil and Uruguay); Mees (1967, Suriname).
Heptapterus bleekeri Boeseman, 1953
Heptapterus bleekeri Boeseman, 1953: 3, fig. 1a. Type locality: Marowijne basin, Nassau Mountains in brooklet[Suriname]. Holotype: RMNH 19421 (larger of 2 specimens).
Distribution: Maroni River basin, and State of Amapá, Brazil (Bockmann & Guazzelli, 2003).
Heptapterus fissipinnis Miranda Ribeiro, 1911
Heptapterus fissipinnis Miranda Ribeiro, 1911: 240. Type locality: corrego affluente do Rio Estrella [probably in theState of Rio de Janeiro, Brazil]. Holotype: at MNRJ (current whereabouts unknown). Misspelled Heptapterusñssipinnis in heading for account.
Distribution: Stream tributary to the Estrela River, Brazil (Bockmann & Guazzelli, 2003).
Heptapterus multiradiatus Ihering, 1907
Heptapterus multiradiatus Ihering, 1907: 20. Type locality: ribeirão do Alto da Serra, Estado de S. Paulo [Brazil].
Holotype: MZUSP 294.
Distribution: Upper Tietê River basin, Brazil (Bockmann & Guazzelli, 2003).
Heptapterus mustelinus (Valenciennes, 1835)
Pimelodus mustelinus Valenciennes, 1835, in Valenciennes, 1835–47: pl. 2 (fig. 1). Type locality: Rio de la Plata.
Holotype: at MNHN. Name available from caption to plate, with illustrated specimen the holotype, if recognizable; species later described in Cuvier & Valenciennes (1840b: 165; 122 of Strasbourg deluxe edition).
Heptapterus eigenmanni Steindachner, 1907e: 487. Type locality: Maldonado. Lectotype: MCZ 7597, designated by
Buckup (1988: 650), but listed in the publication as MCZ 7579.
Distribution: La Plata and Uruguay River basins, and coastal drainages of southern Brazil (Bockmann & Guazzelli,2003).
Heptapterus stewarti Haseman, 1911
Heptapterus stewarti Haseman, 1911b: 376, pl. 74. Type locality: Serrinha Paraná, in a creek about one-half milefrom the Rio Iguassú. Holotype: FMNH 54234.
Distribution: Iguaçu River basin, Brazil (Bockmann & Guazzelli, 2003).
Heptapterus sympterygium Buckup, 1988
Heptapterus sympterygium Buckup, 1988: 644, fig. 1. Type locality: Canal at east side of highway BR-471 nearpumping house of the Departamento Nacional de Obras de Saneamento (DNOS), 52º31'W, 32º32'S, Ecological
Station of Taim (EET), Rio Grande, State of Rio Grande do Sul (RS), Brazil. Holotype: MZUSP 19179.
Distribution: Coastal lowlands of eastern Rio Grande do Sul State, Brazil (Bockmann & Guazzelli, 2003).
Heptapterus tapanahoniensis Mees, 1967
Heptapterus tapanahoniensis Mees, 1967: 223, fig. 2. Type locality: Tapanahoni, about two kilometres downstreamof its confluence with the Paloemeu [Marowijne River basin, Suriname]. Holotype: RMNH 25546.
Distribution: Marowijne/Maroni and Sinnamary River basins (Bockmann & Guazzelli, 2003).
Heptapterus tenuis Mees, 1986
Heptapterus tenuis Mees, 1986: 322, fig. 3. Type locality: Crique Cascade Moyen, Maroni [French Guiana]. Holotype: RMNH 29422.
Distribution: Maroni River basin, French Guiana (Bockmann & Guazzelli, 2003).

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Species inquirenda, Heptapterus
Heptapterus ornaticeps Ahl, 1936: 110. Type locality: Rio de Janeiro [probably incorrect]. Holotype: ZMB 20933.
Known only from holotype, which Bockmann & Guazzelli (2003) indicated was likely to have been incorrectly
reported to have originated from the vicinity of Rio de Janeiro.
HORIOMYZON Stewart, 1986
Horiomyzon Stewart, 1986b: 47. Type species: Horiomyzon retropinnatus Stewart, 1986. Type by original designation. Gender: Masculine.
Horiomyzon retropinnatus Stewart, 1986
Horiomyzon retropinnatus Stewart, 1986b: 48, fig. 1. Type locality: Ecuador, Napo Province, Río Napo at Añangu,
just downstream from confluence with Río Añangu Cocha, lat. 0º31.6'S, long. 76º22.9'W. Holotype: FMNH
96553.
Distribution: Amazon and Napo River basins (Bockmann & Guazzelli, 2003).
IMPARFINIS Eigenmann & Norris, 1900
Imparfinis Eigenmann & Norris, 1900: 351. Type species: Imparfinis piperatus Eigenmann & Norris, 1900. Type bymonotypy. Gender: Masculine.
Nannorhamdia Regan, 1913d: 467. Type species: Nannorhamdia spurrellii Regan, 1913. Type by monotypy. Gender: Feminine.
Imparfinis borodini Mees & Cala, 1989
Imparfinis longicauda Borodin, 1927b: 4, fig. 3. Type locality: Franca, Rio Grande, Prov. of S. Paulo, Brazil. Holotype: AMNH 8639. Preoccupied in Imparfinis by Pimelodus (Rhamdia) longicauda Boulenger, 1887; replaced
by Imparfinis borodini Mees & Cala, 1989.
Imparfinis borodini Mees & Cala, 1989: 387. Type locality: Franca, Rio Grande, Prov. of S. Paulo, Brazil. Holotype:
AMNH 8639. Replacement for Imparfinis longicauda Borodin, 1927.
Distribution: Upper Paraná, São Francisco and Tocantins River basins, Brazil (Bockmann & Guazzelli, 2003).
Imparfinis cochabambae (Fowler, 1940)
Pimelodella cochabambae Fowler, 1940c: 80, figs. 35–37. Type locality: Boca Chapare, Cochabamba, Bolivia.
Holotype: ANSP 69066.
Distribution: Beni River basin, Bolivia (Bockmann & Guazzelli, 2003).
Imparfinis guttatus (Pearson, 1924)
Nannorhamdia guttatus Pearson, 1924: 11, pl. 2 (fig. 2). Type locality: Popoi River, Upper Beni [Bolivia]. Holotype:
CAS 63689.
Distribution: Mamoré and Pilcomayo River basins, Bolivia and Peru (Bockmann & Guazzelli, 2003).
Imparfinis hasemani Steindachner, 1915
Imparfinis hasemani Steindachner, 1915e: 45, figs. 1–3. Type locality: Rio Surumú bei Serra do Mello, dem Rio
branco bei Bem Querer und dem Rio Tapajos bei Santarem [Brazil]. Syntype: NMW 46093:1 (1).
Distribution: Branco and Tapajós River basins, Brazil (Bockmann & Guazzelli, 2003).
Imparfinis hollandi Haseman, 1911
Imparfinis hollandi Haseman, 1911b: 383, pl. 48 (fig. 2). Type locality: Porto União da Victoria, Rio Iguassú. Holotype: FMNH 54244.
Distribution: Iguaçu River basin (Bockmann & Guazzelli, 2003).
Imparfinis lineatus (Bussing, 1970)
Nannorhamdia lineata Bussing, 1970: 1, fig. 1. Type locality: Costa Rica: Puntarenas Prov., Quebrada 36 (elev. 80m) 12 km W of Pueblo Río Claro at Interamerican Highway. Holotype: LACM 30688-1.
Distribution: Streams in Puntarenas Province, Costa Rica (Bockmann & Guazzelli, 2003).
Imparfinis longicauda (Boulenger, 1887)
Pimelodus (Rhamdia) longicauda Boulenger, 1887c: 275, pl. 20 (fig. 2). Type locality: Canelos [Ecuador]. Syn-CHECKLIST OF CATFISHES, RECENT AND FOSSIL

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types: BMNH 1880.12.8.100–104 (5).
Distribution: Bobonaza River basin in upper Pastaza River drainage, Ecuador (Bockmann & Guazzelli, 2003).
Imparfinis microps Eigenmann & Fisher, 1916
Imparfinis microps Eigenmann & Fisher, in Eigenmann, 1916: 82. Type locality: Rio Negro at Villavicencio,
Colombia. Holotype: FMNH 57793, illustrated in Eigenmann (1922b: 285, pl. 4, figs. 2–3). Also described as
new in Eigenmann (1919a: 128).
Distribution: Meta River basin, Colombia (Bockmann & Guazzelli, 2003).
Imparfinis minutus (Lütken, 1874)
Rhamdia minuta Lütken, 1874c: 35. Type locality: in provincia Minas-geraës circa oppidulum Lagoa Santa, praecipue in flumine Rio das Velhas et affluentibus. Syntypes: ZMUC P 29661 (1), ZMUC P 29662 (1), ZMUC P
29663 (1). Illustrated and described in more detail in Lütken (1875: 179–180, pl. 3, fig. 6).
Distribution: Upper São Francisco River basin, Brazil (Bockmann & Guazzelli, 2003).
Imparfinis mirini Haseman, 1911
Imparfinis mirini Haseman, 1911a: 318, pl. 47. Type locality: Near Piracicaba above the falls of Rio Piracicabamirini [Brazil]. Holotype: FMNH 54335.
Distribution: Upper Araguaia and Paraná River basins, Brazil (Bockmann & Guazzelli, 2003).
Imparfinis nemacheir (Eigenmann & Fisher, 1916)
Nannorhamdia nemacheir Eigenmann & Fisher, in Eigenmann, 1916: 83. Type locality: Girardot, Colombia. Holotype: FMNH 58127, illustrated in Eigenmann (1922b: 41, fig. 13; pl. 4, fig. 4).
Distribution: Upper Amazon and Orinoco River basins (Bockmann & Guazzelli, 2003); Magdalena, Cauca and Patia
River basins, Colombia (Maldonado-Ocampo et al., 2005).
Imparfinis pijpersi (Hoedeman, 1961)
Cetopsorhamdia pijpersi Hoedeman, 1961: 132, fig. 2. Type locality: Surinam: ... Sipaliwini river, sta. 43, 20 km
from frontier with Brazil. Holotype: ZMA 102230.
Distribution: Corantijn River basin, Suriname (Bockmann & Guazzelli, 2003).
Imparfinis piperatus Eigenmann & Norris, 1900
Imparfinis piperatus Eigenmann & Norris, 1900: 352. Type locality: S. Paulo, Brazil. Holotype: CAS 63636.
Distribution: In tributaries of upper Paraná River and/or coastal streams of São Paulo State, Brazil (Bockmann &Guazzelli, 2003).
Imparfinis pristos Mees & Cala, 1989
Imparfinis pristos Mees & Cala, 1989: 391, fig. 3. Type locality: Colombia: … second tributary, left side, from themouth of the Rio Tomo, Vichada. Holotype: ICNMHN 1401.
Distribution: Negro and Orinoco River basins (Bockmann & Guazzelli, 2003).
Imparfinis pseudonemacheir Mees & Cala, 1989
Imparfinis pseudonemacheir Mees & Cala, 1989: 387, fig. 2. Type locality: Venezuela: … Rio Tucuragua, near
Caicara, Bolívar. Holotype: RMNH 30489 (37.5 mm SL specimen).
Distribution: Orinoco and Ucayali River basins (Bockmann & Guazzelli, 2003).
Imparfinis schubarti (Gomes, 1956)
Nannorhamdia schubarti Gomes, 1956: 404, fig. 1. Type locality: da Cachoeira de Emas, Rio Mogi Guaçu, e do
Córrego do Tijuco Prêto, um seu afluente, Município de Pirassununga, São Paulo [Brazil]. Holotype: EEBP
379.
Distribution: Upper Paraná River basin, Brazil (Bockmann & Guazzelli, 2003).
Imparfinis spurrellii (Regan, 1913)
Nannorhamdia spurrellii Regan, 1913d: 467. Type locality: R. Condoto [Colombia]. Holotype: BMNH
1913.10.1.41.
Distribution: San Juan River basin, Colombia (Bockmann & Guazzelli, 2003).
Imparfinis stictonotus (Fowler, 1940)
Nannorhamdia stictonotus Fowler, 1940c: 66, figs. 21–22. Type locality: Todos Santos, Rio Chapare, Bolivia. Holotype: ANSP 68892.
Distribution: Mamoré/Madeira, Paraguay, and Ucayali River basins (Bockmann & Guazzelli, 2003).

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FERRARISLEPTORHAMDIA Eigenmann, 1918
Leptoglanis Eigenmann, 1912b: 158. Type species: Leptoglanis essequibensis Eigenmann, 1912. Type by original
designation. Gender: Masculine. Preoccupied by Leptoglanis Boulenger, 1902, in fishes; replaced by Leptorhamdia Eigenmann, 1918.
Leptorhamdia Eigenmann, 1918a: 260. Type species: Leptoglanis essequibensis Eigenmann, 1912. Type by being a
replacement name. Gender: Feminine. Replacement for Leptoglanis Eigenmann, 1912; preoccupied by Leptoglanis Boulenger, 1902.
Leptorhamdia essequibensis (Eigenmann, 1912)
Leptoglanis essequibensis Eigenmann, 1912b: 158, pl. 13 (fig. 2). Type locality: Crab Falls, British Guiana. Holotype: FMNH 53331.
Distribution: Essequibo and lower Tocantins River basins (Bockmann & Guazzelli, 2003).
Leptorhamdia marmorata Myers, 1928
Leptorhamdia marmorata Myers, 1928: 83. Type locality: Rock-pools at São Gabriel Rapids, Rio Negro, Brazil.
Holotype: CAS 63671.
Distribution: Negro River basin, Brazil and Venezuela (Bockmann & Guazzelli, 2003).
Leptorhamdia schultzi (Miranda Ribeiro, 1964)
Rhamdella schultzi Miranda Ribeiro, 1964: 1, fig. 1. Type locality: Alto Rio Xingú [Brazil]. Holotype: MNRJ 9549.
Distribution: Upper Xingu River basin, Brazil (Bockmann & Guazzelli, 2003).
Remarks: Placement in Leptorhamdia follows Bizerril & Borba (1994).
MASTIGLANIS Bockmann, 1994
Mastiglanis Bockmann, 1994: 762. Type species: Mastiglanis asopos Bockmann, 1994. Type by original designation. Gender: Masculine.
Mastiglanis asopos Bockmann, 1994
Mastiglanis asopos Bockmann, 1994: 762, fig. 1. Type locality: Brazil, Pará, Igarapé Saracazinho, tributary of rio
Trombetas, near Porto Trombetas. Holotype: MNRJ 12227.
Distribution: Amazon, Capim and Orinoco River basins (Bockmann & Guazzelli, 2003).
MYOGLANIS Eigenmann, 1912
Myoglanis Eigenmann, 1912b: 159. Type species: Myoglanis potaroensis Eigenmann, 1912. Type by original designation. Gender: Masculine.
Myoglanis aspredinoides Do Nascimiento & Lundberg, 2005
Myoglanis aspredinoides Do Nascimiento & Lundberg, 2005: 39, fig. 1. Type locality: Venezuela, Estado Amazonas, Río Ventuari, Laguna Pavón, in pool behind beach of Río Ventuari, on south side of river, ca. 12 km fromits confluence with Río Orinoco; 4º4'N, 66º56'W. Holotype: MBUCV V-25388.
Distribution: Ventuari River, Orinoco River basin, Venezuela (Do Nascimiento & Lundberg, 2005).
Myoglanis koepckei Chang, 1999
Myoglanis koepckei Chang, 1999: 434, fig. 1. Type locality: Peru, Departamento Loreto, Provincia Maynas, Quebrada of the Mishana camp, small tributary of the Río Nanay, 03º52'S, 73º29'W, 107 m elevation. Holotype:
MUSM 10045.
Distribution: Upper Amazon River basin, Peru (Bockmann & Guazzelli, 2003).
Myoglanis potaroensis Eigenmann, 1912
Myoglanis potaroënsis Eigenmann, 1912b: 159, pl. 14 (fig. 1). Type locality: Creek at Tukeit, British Guiana. Holotype: FMNH 53333.
Distribution: Essequibo River basin, Guyana (Bockmann & Guazzelli, 2003).
NANNOGLANIS Boulenger, 1887
Nannoglanis Boulenger, 1887c: 278. Type species: Nannoglanis fasciatus Boulenger, 1887. Type by monotypy.
Gender: Masculine.

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Nannoglanis fasciatus Boulenger, 1887
Nannoglanis fasciatus Boulenger, 1887c: 278, pl. 21 (fig. 3). Type locality: eastern Ecuador. Syntypes: BMNH
1880.12.8.149–150 (2).
Distribution: Napo River basin, Ecuador (Bockmann & Guazzelli, 2003).
NEMUROGLANIS Eigenmann & Eigenmann, 1889
Nemuroglanis Eigenmann & Eigenmann, 1889b: 29. Type species: Nemuroglanis lanceolatus Eigenmann & Eigenmann, 1889. Type by original designation. Gender: Masculine.
Imparales Schultz, 1944b: 93. Type species: Imparales mariai Schultz, 1944. Type by original designation. Gender:
Masculine.
Medemichthys Dahl, 1961: 490. Type species: Medemichthys guayaberensis Dahl, 1961. Type by original designation. Gender: Masculine.
Revision: Bockmann & Ferraris (2005).
Nemuroglanis lanceolatus Eigenmann & Eigenmann, 1889
Nemuroglanis lanceolatus Eigenmann & Eigenmann, 1889b: 29. Type locality: Jutahy [Brazil]. Holotype: MCZ
8169.
Distribution: Napo and Solimões River basins, Brazil and Ecuador (Bockmann & Guazzelli, 2003).
Nemuroglanis mariai (Schultz, 1944)
Imparales mariai Schultz, 1944b: 94, fig. 1. Type locality: Río Meta system at Villavicencio, Colombia (Orinoco
drainage). Holotype: USNM 121251, holotype illustrated in Bockmann & Ferraris (2005: fig. 1c).
Medemichthys guayaberensis Dahl, 1961: 491, figure on p. 493. Type locality: small brook near the Expedition’scamp I, on the left side of the Guayabero River [Colombia]. Holotype: probably lost (Cala, 1981).
Distribution: Meta and Guaviare River basins, Colombia (Bockmann & Guazzelli, 2003, Bockmann & Ferraris,2005).
Nemuroglanis pauciradiatus Ferraris, 1988
Nemuroglanis pauciradiatus Ferraris, 1988a: 510, fig. 1. Type locality: Venezuela: Territorio Federal Amazonas;
morichal 26.9 km from Puerto Ayacucho along Puerto Ayacucho to Caicara highway. Holotype: MBUCV V16450.
Distribution: Negro, Orinoco and Sanabani River basins, Brazil and Venezuela (Bockmann & Guazzelli, 2003).
PARIOLIUS Cope, 1872
Pariolius Cope, 1872a: 289. Type species: Pariolius armillatus Cope, 1872. Type by monotypy. Gender: Masculine.
Pariolius armillatus Cope, 1872
Pariolius armillatus Cope, 1872a: 289. Type locality: the Ambyiacu [Peru]. Holotype: at ANSP; not ANSP 22004.
Distribution: Upper Amazon River basin, Brazil and Peru (Bockmann & Guazzelli, 2003).
Remarks: Redescribed in Gosline (1940a). ANSP 22004 is a trichomycterid, thus cannot be the holotype.
PHENACORHAMDIA Dahl, 1961
Phenacorhamdia Dahl, 1961: 504. Type species: Phenacorhamdia macarenensis Dahl, 1961. Type by original designation. Gender: Feminine.
Phenacorhamdia anisura (Mees, 1987)
Heptapterus anisurus Mees, 1987a: 451, fig. 1. Type locality: Río Guarapiche near Maturin, Monagas, Venezuela.
Holotype: RMNH 30499.
Distribution: Orinoco River basin, Venezuela (Bockmann & Guazzelli, 2003).
Phenacorhamdia boliviana (Pearson, 1924)
Imparfinis bolivianus Pearson, 1924: 12, pl. 2 (fig. 3). Type locality: Huachi [Bolivia]. Syntypes: CAS 63632 (2).
Distribution: Upper Mamoré/Madeira River basin, Bolivia (Bockmann & Guazzelli, 2003).
Phenacorhamdia hoehnei (Miranda Ribeiro, 1914)
Nannoglanis hoehnei Miranda Ribeiro, 1914: 3, pl. 1 (fig. 1). Type locality: Rio Taquary, Matto Grosso [Brazil].
Lectotype: MNRJ 787A, designated by Miranda Ribeiro (1953: 403).

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FERRARISDistribution: Taquari River basin, Paraguay River drainage, Brazil (Bockmann & Guazzelli, 2003).
Phenacorhamdia macarenensis Dahl, 1961
Phenacorhamdia macarenensis Dahl, 1961: 504, figure on p. 507. Type locality: Caño Lozada [Colombia]. Holotype: probably lost (Cala, 1981: 1).
Distribution: Guaviare River basin, Colombia (Bockmann & Guazzelli, 2003).
Phenacorhamdia nigrolineata Zarske, 1998
Phenacorhamdia nigrolineata Zarske, 1998: 27, fig. 1. Type locality: Tierra Roja na der Strabe Campoverde —Tournavista, nahe Pucallpa, Departamento Ucayali, Peru. Holotype: MTD F 20 728.
Distribution: Ucayali River basin, Peru (Bockmann & Guazzelli, 2003).
Phenacorhamdia somnians (Mees, 1974)
Heptapterus somnians Mees, 1974: 184, fig. 36. Type locality: Sangadina, a tributary of the Rio das Mortes, 2 km
from Xaventina, Mato Grosso, Brazil. Holotype: BMNH 1971.7.29.4.
Distribution: Das Mortes River basin, Brazil (Bockmann & Guazzelli, 2003).
Phenacorhamdia tenebrosa (Schubart, 1964)
Imparfinis tenebrosus Schubart, 1964a: 9, fig. 3. Type locality: Rio Mogi Guaçu, entre o pesqueiro da EE [=Estação
Experimental de Piscicultura de Pirassununga] e o pesqueiro do Sr. Firmino (Faz. Graciosa) ca. 1 km abaixo da
Cachoeira [Brazil]. Syntypes: EEBP 109 (1), EEBP 197 (5), EEBP 630 (4).
Distribution: Upper Paraná and São Francisco River basins, Brazil (Bockmann & Guazzelli, 2003).
Phenacorhamdia unifasciata Britski, 1993
Phenacorhamdia unifasciata Britski, 1993: 42, fig. 1. Type locality: Ribeirão Santana, afluente do Rio São Bartolomeu (15º50'N, 47º40'W), Distrito Federal [Brazil]. Holotype: MZUSP 36691.
Distribution: Upper Paraná River basin, Brazil (Bockmann & Guazzelli, 2003).
PIMELODELLA Eigenmann & Eigenmann, 1888
Pimelodella Eigenmann & Eigenmann, 1888b: 131. Type species: Pimelodus cristatus Müller & Troschel, 1848.
Type by original designation. Gender: Feminine.
Typhlobagrus Miranda Ribeiro, 1907b: [1]. Type species: Typhlobagrus kronei Miranda Ribeiro, 1907. Type bymonotypy. Gender: Masculine.
Caecorhamdella Borodin, 1927a: 1. Type species: Caecorhamdella brasiliensis Borodin, 1927. Type by original
designation. Gender: Feminine.
Revision: Eigenmann (1917b).
Review: Guazzelli (1997, southeastern Brazil).
Pimelodella altipinnis (Steindachner, 1864)
Pimelodus altipinnis Steindachner, 1864a: 37. Type locality: Britisch-Guiana. Holotype: NMW 45601. Illustrated
and described in more detail in Steindachner (1864b: 213, pl. 2, figs. 3–4).
Distribution: Essequibo River basin, Guyana (Bockmann & Guazzelli, 2003).
Pimelodella australis Eigenmann, 1917
Pimelodella laticeps australis Eigenmann, 1917b: 243. Type locality: Uruguayana [State of Rio Grande do Sul, Brazil]. Holotype: FMNH 57962.
Pimelodela garbei Miranda Ribeiro, 1918d: 639. Type locality: Itaqui —Rio Grande do Sul [Brazil]. Lectotype:
MNRJ 923A, designated by Miranda Ribeiro (1953: 403). Name clearly is a lapsus for Pimelodella garbei.
Distribution: Uruguay River basin, and coastal river drainages of southeastern Brazil (Bockmann & Guazzelli,2003).
Pimelodella avanhandavae Eigenmann, 1917
Pimelodella avanhandavae Eigenmann, 1917b: 240, pl. 29 (fig. 3). Type locality: Rio Tietê at Salto Avanhandava,
above the fall [Brazil]. Holotype: FMNH 57981.
Distribution: Tietê River basin, Brazil (Bockmann & Guazzelli, 2003).
Pimelodella boliviana Eigenmann, 1917
Pimelodella boliviana Eigenmann, 1917b: 245, pl. 31 (fig. 2). Type locality: Santa Cruz de la Sierra, Bolivia. Holotype: FMNH 57976.

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Distribution: Mamoré River basin, Bolivia (Bockmann & Guazzelli, 2003).
Pimelodella boschmai Van der Stigchel, 1964
Pimelodella boschmai Van der Stigchel, 1964: 327, fig. 1. Type locality: Mogi-Guassu (river), below the Emas falls,... Pirassununga, Brazil. Holotype: RMNH 23248.
Pimelodella insignis Schubart, 1964a: 6, fig. 1. Type locality: Rio Mogi Guaçu, abaixo do pesqueiro da EEBP;
Cochoeira [sic] de Emas, na topava; Rio Mogi Guaçu, abaixo do pesqueiro; Rio Mogi Guaçu, abaixo da cachoeira; Rio Mogi Guaçu; Rio Mogi Guaçu, pesqueiro da EEBP [Brazil]. Syntypes: EEBP 497, EEBP 497 [bis],
EEBP 498, EEBP 499, EEBP 620, EEBP 620a, EEBP 647.
Distribution: Mogi Guaçu River basin, Brazil (Bockmann & Guazzelli, 2003).
Pimelodella brasiliensis (Steindachner, 1877)
Pimelodus (Pseudorhamdia) brasiliensis Steindachner, 1877b: 608, pl. 7. Type locality: Rio Parahyba [Brazil].
Holotype: NMW 45612.
Distribution: Paraíba do Sul River basin, Brazil (Bockmann & Guazzelli, 2003).
Pimelodella breviceps (Kner, 1857)
Pimelodus breviceps Kner, 1857: 418. Type locality: Marabitanos [Brazil]. Holotype: NMW 45615.
Distribution: Upper Negro River basin, Brazil (Bockmann & Guazzelli, 2003).
Pimelodella buckleyi (Boulenger, 1887)
Pimelodus buckleyi Boulenger, 1887c: 275, pl. 20 (fig. 1). Type locality: Canelos [Ecuador]. Syntypes: BMNH
1880.12.8.98–99 (2).
Pimelodella copei Fowler, 1915a: 216, fig. 5. Type locality: Ambyiacu River, near Pebas, Ecuador. Holotype: ANSP
8362.
Distribution: Ambyiacu and Bobonaza River basins, upper Amazon basin (Bockmann & Guazzelli, 2003).
Pimelodella chagresi (Steindachner, 1876)
Pimelodus (Pseudorhamdia) Chagresi Steindachner, 1876: 584. Type locality: Rio Chagres und dessen Nebenflusse
bei Obispo [Panama]. Syntypes: at NMW.
Distribution: Chagres River basin, Panama (Bockmann & Guazzelli, 2003); Magdalena, Cauca, Patia, andCatatumbo River basins, Colombia (Maldonado-Ocampo et al., 2005).
Pimelodella chaparae Fowler, 1940
Pimelodella chaparae Fowler, 1940c: 75, figs. 29–30. Type locality: Boca Chapare, Cochabamba, Bolivia. Holotype: ANSP 69021.
Distribution: Beni River basin, Bolivia (Bockmann & Guazzelli, 2003).
Pimelodella conquetaensis Ahl, 1925
Pimelodella conquetaensis Ahl, 1925: 106. Type locality: Rio Coquetá [sic Caquetá], S.O. Columbien. Holotype:
ZMB 32030.
Distribution: Caquetá River basin in upper Japurá River drainage, Colombia (Bockmann & Guazzelli, 2003).
Pimelodella cristata (Müller & Troschel, 1848)
Pimelodus cristatus Müller & Troschel, in Schomburgk, 1848: 628. Type locality: Takutu und Mahu [Branco Riverbasin, Guyana]. Syntypes: ZMB 3052 (1), ZMB 3053 (1).
Distribution: Guyana and French Guiana (Bockmann & Guazzelli, 2003).
Remarks: Also cited as new in Müller & Troschel (1849: 4).
Pimelodella cruxenti Fernández-Yépez, 1950
Pimelodella cruxenti Fernández-Yépez, 1950a: 5, pl. 1 (fig. 2). Type locality: Río Autana, Venezuela. Holotype:
AFY 48161; whereabouts unknown.
Distribution: Autana River, Orinoco River drainage, Venezuela (Bockmann & Guazzelli, 2003).
Pimelodella cyanostigma (Cope, 1870)
Rhamdia cyanostigma Cope, 1870b: 569. Type locality: Pebas, Ecuador. Syntypes: ANSP 8381–8383 (3).
Distribution: Ampyiacu River, upper Amazon basin, Peru (Bockmann & Guazzelli, 2003).
Pimelodella dorseyi Fowler, 1941
Pimelodella dorseyi Fowler, 1941a: 127, figs. 4–7. Type locality: Rio Salgade, Icó, Ceará [Brazil]. Holotype: ANSP
69375.

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Pimelodella eigenmanni (Boulenger, 1891)
Pimelodus (Pimelodella) eigenmanni Boulenger, 1891: 232, footnote. Type locality: Macacos [Brazil]. Holotype:
BMNH 1889.11.14.6.
Distribution: Paraíba do Sul River basin, Brazil (Bockmann & Guazzelli, 2003).
Pimelodella eigenmanniorum (Miranda Ribeiro, 1911)
Rhamdia eigenmanniorum Miranda Ribeiro, 1911: 273. Type locality: from the Rio Parahyba and from Macacos[Brazil]. Syntypes (5): possibly MCZ 7438 (5), MCZ 7510 (1).
Distribution: Paraíba do Sul River basin and coastal rivers of Rio de Janeiro State, Brazil (Bockmann & Guazzelli,2003).
Pimelodella elongata (Günther, 1860)
Pimelodus elongatus Günther, 1860: 238, pl. 10 (fig. b). Type locality: Fresh waters of Esmeraldas, Ecuador. Syntypes: BMNH 1860.6.16.182–185, BMNH 1860.6.16.186–189, BMNH 1860.6.16.192, BMNH 1860.6.16.194.
Distribution: Esmeraldas River basin, Ecuador (Bockmann & Guazzelli, 2003).
Pimelodella enochi Fowler, 1941
Pimelodella enochi Fowler, 1941a: 130, figs. 12–15. Type locality: Açude Piloes, Parahyba, Brazil. Holotype:
ANSP 69378.
Distribution: Paraíba River basin in Paraíba State, Brazil (Bockmann & Guazzelli, 2003).
Pimelodella eutaenia Regan, 1913
Pimelodella eutaenia Regan, 1913d: 466. Type locality: Rio Condoto and the Rio Sipi [Colombia]. Syntypes (5, 75–
145 mm TL): BMNH 1913.10.1.37–40.
Distribution: San Juan River basin, Colombia (Bockmann & Guazzelli, 2003).
Pimelodella figueroai Dahl, 1961
Pimelodella figueroai Dahl, 1961: 498, figure on p. 500. Type locality: Caño Lozada, approx. 17 kilometers aboveits junction with Guayabero River [Colombia]. Holotype: probably lost (Cala, 1981).
Distribution: Guaviare River basin, Colombia (Bockmann & Guazzelli, 2003).
Pimelodella geryi Hoedeman, 1961
Pimelodella geryi Hoedeman, 1961: 134. Type locality: French Guiana: ... mainland, Litany River, village Aloiké,
sta. 29S. Holotype: ZMA 100168.
Distribution: Maroni River basin, Suriname (Bockmann & Guazzelli, 2003).
Pimelodella gracilis (Valenciennes, 1835)
Pimelodus gracilis Valenciennes, 1835 (in Valenciennes, 1835–47): pl. 2 (fig. 5). Type locality: [not stated]. Holotype: Possibly MNHN a-9284 or MNHN 1-9285. Name available from plate; with illustrated specimen theholotype, if identifiable. Described in Cuvier & Valenciennes (1840b: 181 (134 in Strasbourg deluxe edition)),with the locality as: Corrientes dans le Parana et les autres rivières au-dessus de 28º de latitud sud [Argentina].
Distribution: Orinoco, Amazon and La Plata River basins (Bockmann & Guazzelli, 2003).
Remarks: Bockmann & Guazzelli (2003) list MNHN a-9284 as the holotype of Pimelodus gracilis, but without justifying this statement. If the illustrated specimen cannot be identified, the statement of Bockmann & Guazzelli(2003) cannot be taken to be a de facto lectotype designation.
Pimelodella griffini Eigenmann, 1917
Pimelodella griffini Eigenmann, 1917b: 250, pl. 32 (fig. 3). Type locality: mountain rills near Sapucay, Paraguay.
Holotype: FMNH 57974.
Distribution: Lower Paraguay River basin, Paraguay (Bockmann & Guazzelli, 2003).
Pimelodella grisea (Regan, 1903)
Pimelodus (Pimelodella) griseus Regan, 1903b: 625. Type locality: Durango, Sapayo and Vaqueria Rivers, NW
Ecuador. Syntypes: BMNH 1902.5.27.36 (1), BMNH 1902.7.29.47 (1), BMNH 1902.7.29.58 (1).
Distribution: Durango, Sapayo and Vaqueria Rivers, NW Ecuador (Bockmann & Guazzelli, 2003).
Pimelodella hartii (Steindachner, 1877)
Pimelodus (Pseudorhamdia) Hartii Steindachner, 1877b: 611. Type locality: Rio Parahyba [Brazil]. Holotype:
NMW 45784.

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Distribution: Paraíba do Sul River basin, Brazil (Bockmann & Guazzelli, 2003).
Pimelodella hartwelli Fowler, 1940
Pimelodella hartwelli Fowler, 1940a: 222, figs.4–7. Type locality: Ucayali River basin, Contamana, Peru. Holotype:
ANSP 68644.
Distribution: Ucayali River basin, Peru (Bockmann & Guazzelli, 2003).
Pimelodella hasemani Eigenmann, 1917
Pimelodella hasemani Eigenmann, 1917b: 241, pl. 30 (fig. 1). Type locality: San Antonio de Rio Madeira. Holotype: FMNH 57980.
Distribution: Madeira River basin (Bockmann & Guazzelli, 2003).
Pimelodella howesi Fowler, 1940
Pimelodella howesi Fowler, 1940c: 77, figs. 32–34. Type locality: Boca Chapare, Chimore River, Bolivia. Holotype:
ANSP 69036.
Distribution: Beni River basin, Bolivia (Bockmann & Guazzelli, 2003).
Pimelodella itapicuruensis Eigenmann, 1917
Pimelodella itapicuruensis Eigenmann, 1917b: 247, pl. 31 (fig. 3). Type locality: Queimadas, Rio Itapicurú [Brazil].
Holotype: FMNH 57986.
Distribution: Itapicurú River basin, Bahia, Brazil (Bockmann & Guazzelli, 2003).
Pimelodella kronei (Miranda Ribeiro, 1907)
Typhlobagrus kronei Miranda Ribeiro, 1907b: [1], fig. on p. [1]. Type locality: Iguape, em aguas das cavernas do
Iporanga, também no estado de S. Paulo [Brazil]. Holotype: MNRJ 836.
Caecorhamdella brasiliensis Borodin, 1927a: 1, fig. 1c. Type locality: Estado de São Paulo, Brazil. Holotype:
AMNH 8604.
Distribution: Caves in the Ribeira do Iguape River basin, in São Paulo State, Brazil (Bockmann & Guazzelli, 2003).
Remarks: Synonymy based on Trajano & Britski (1992); If Caecorhamdella brasiliensis is later determined to be a
distinct species of Pimelodella, it would require a new name, inasmuch as it would be preoccupied in Pimelodella by Pimelodella brasiliensis (Steindachner, 1877).
Pimelodella lateristriga (Lichtenstein, 1823)
Pimelodes lateristrigus Lichtenstein, 1823: 112. Type locality: Brasilien. Holotype: ZMB 3038.
Pimelodus lateristrigus Müller & Troschel, 1849: 3. Type locality: Brasilien. Holotype: Probably ZMB 3038.
Distribution: Paraíba do Sul River basin, Brazil (Bockmann & Guazzelli, 2003).
Pimelodella laticeps Eigenmann, 1917
Pimelodella laticeps Eigenmann, 1917b: 243, pl. 30 (fig. 2). Type locality: Sapucay, Paraguay. Holotype: FMNH
57969.
Distribution: Lower Paraguay River basin, Paraguay (Bockmann & Guazzelli, 2003).
Pimelodella laurenti Fowler, 1941
Pimelodella laurenti Fowler, 1941a: 132, figs. 16–19. Type locality: Jatobá, Rio São Francisco, Pernambuco [Brazil]. Holotype: ANSP 69380 [incorrectly published as 19380].
Distribution: Lower São Francisco River basin, Brazil (Bockmann & Guazzelli, 2003).
Pimelodella linami Schultz, 1944
Pimelodella linami Schultz, 1944c: 210, pl. 1 (fig. d). Type locality: Río Torbes, 1 km. above Táriba, Orinoco system [Venezuela]. Holotype: USNM 121132.
Distribution: Orinoco River basin, Venezuela (Bockmann & Guazzelli, 2003).
Pimelodella macrocephala (Miles, 1943)
Nannorhamdia macrocephala Miles, 1943a: 25. Type locality: Valle del Cauca [Colombia]. Holotype: ICNMHN,
probably lost (Cala, 1981).
Distribution: Cauca River basin, Colombia (Bockmann & Guazzelli, 2003; Maldonado-Ocampo et al., 2005).
Pimelodella macturki Eigenmann, 1912
Pimelodella macturki Eigenmann, 1912b: 170, pl. 16 (fig. 1). Type locality: Creek in Mora Passage, British Guiana.
Holotype: FMNH 53234.
Distribution: Approuague, Corantijn, and Nickerie River basins, and eastern coastal drainages of Suriname (Bock-

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mann & Guazzelli, 2003).
Pimelodella martinezi Fernández-Yépez, 1970
Pimelodella martinezi Fernández-Yépez, 1970: pl. 35. Type locality: bajo Unare [Venezuela]. Holotype: AFY 70403; whereabouts unknown.
Distribution: Unare River basin, Venezuela (Bockmann & Guazzelli, 2003).
Pimelodella meeki Eigenmann, 1910
Pimelodella eigenmanni Meek, 1905: 241. Type locality: Sao Paulo, Brazil. Holotype: FMNH 3400; holotype illustrated in Eigenmann (1917b: pl. 32, fig. 1). Preoccupied in Pimelodella by Pimelodus (Pimelodella) eigenmanni
Boulenger, 1891; replaced by Pimelodella meeki Eigenmann, 1910.
Pimelodella meeki Eigenmann, 1910: 389. Type locality: São Paulo. Holotype: FMNH 3400; holotype illustrated in
Eigenmann (1917b: pl. 32, fig. 1). Replacement for Pimelodella eigenmanni Meek, 1905.
Distribution: Tietê River basin, Brazil (Bockmann & Guazzelli, 2003).
Pimelodella megalops Eigenmann, 1912
Pimelodella megalops Eigenmann, 1912b: 169, pl. 15 (fig. 2). Type locality: Tumatumari, British Guiana. Holotype:
FMNH 53231.
Distribution: Approuague River, French Guiana, and Essequibo River, Guyana (Bockmann & Guazzelli, 2003).
Pimelodella megalura Miranda Ribeiro, 1918
Pimelodella megalura Miranda Ribeiro, 1918d: 638. Type locality: S. Luiz de Caceres: Matto-Grosso [Brazil]. Lectotype: MNRJ 865A, designated by Miranda Ribeiro (1953: 403).
Distribution: Upper Paraguay River basin, Brazil (Bockmann & Guazzelli, 2003).
Pimelodella metae Eigenmann, 1917
Pimelodella metae Eigenmann, 1917b: 244, pl. 31 (fig. 1). Type locality: Rio Negro, Villavicencio [Colombia].
Holotype: FMNH 58441.
Distribution: Meta River basin, Colombia (Bockmann & Guazzelli, 2003).
Pimelodella modesta (Günther, 1860)
Pimelodus modestus Günther, 1860: 239, pl. 10 (fig. c). Type locality: Fresh waters of Esmeraldas, Ecuador. Syntypes: BMNH 1860.6.16.190–191 (2).
Distribution: Esmeraldas River basin, Ecuador (Bockmann & Guazzelli, 2003); Patia River, Colombia (MaldonadoOcampo et al., 2005).
Pimelodella montana Allen, 1942
Pimelodella montana Allen, in Eigenmann & Allen, 1942: 100, pl. 13 (fig. 2). Type locality: Rio Huallaga, Huánuco
[Peru]. Syntypes: CAS 63719 (5).
Distribution: Upper Marañon River basin, Peru (Bockmann & Guazzelli, 2003).
Pimelodella mucosa Eigenmann & Ward, 1907
Pimelodella mucosa Eigenmann & Ward, in Eigenmann, McAtee & Ward, 1907: 114, pl. 32 (fig. 1). Type locality:
Bahia Negra [Paraguay]. Holotype: CAS 63720.
Distribution: Paraguay River basin, Paraguay (Bockmann & Guazzelli, 2003).
Pimelodella nigrofasciata (Perugia, 1897)
Pimelodus nigrofasciatus Perugia, 1897: 18. Type locality: Rio Beni, Missioni Mosetenes, Bolivia. Holotype:
MSNG 8039 (reported to be missing in Eschmeyer et al., 1998).
Distribution: Beni River basin, Bolivia (Bockmann & Guazzelli, 2003).
Pimelodella notomelas Eigenmann, 1917
Pimelodella notomelas Eigenmann, 1917b: 244, pl. 30 (fig. 3). Type locality: San Luiz de Caceres [Brazil]. Holotype: FMNH 57967.
Distribution: Upper Paraguay River basin, Brazil (Bockmann & Guazzelli, 2003).
Pimelodella odynea Schultz, 1944
Pimelodella chagresi odynea Schultz, 1944c: 213, pl. 2 (fig. a). Type locality: Río San Juan at the bridge south ofMene Grande, Motatán system, Maracaibo basin [Venezuela]. Holotype: USNM 121133.
Distribution: Lake Maracaibo basin, Venezuela (Bockmann & Guazzelli, 2003).
Pimelodella ophthalmica (Cope, 1878)

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Pimelodus ophthalmicus Cope, 1878: 675. Type locality: Peruvian Amazon. Holotype: ANSP 21102.
Distribution: Upper Amazon River basin, Peru (Bockmann & Guazzelli, 2003).
Pimelodella pallida Dahl, 1961
Pimelodella pallida Dahl, 1961: 501, figure on p. 503. Type locality: Caño Lozada, about 500 meters above its junction with Guayabero River [Colombia]. Holotype: probably lost (Cala, 1981).
Distribution: Guaviare River basin, Colombia (Bockmann & Guazzelli, 2003).
Pimelodella pappenheimi Ahl, 1925
Pimelodella pappenheimi Ahl, 1925: 107. Type locality: Rio Pedro bei Humboldt, Sta Catharina, Brasilien. Syntypes: ZMB 31951 (2).
Distribution: Itajaí-Açú, Itapocú and Cubatão River basins in Santa Catarina State, and Paranaguá River basin in
Paraná State, Brazil (Bockmann & Guazzelli, 2003).
Pimelodella parnahybae Fowler, 1941
Pimelodella parnahybae Fowler, 1941a: 129, figs. 8–11. Type locality: Rio Parnahyba, Therezina, Piauhy [Brazil].
Holotype: ANSP 69377.
Distribution: Parnaíba River basin, Brazil (Bockmann & Guazzelli, 2003).
Pimelodella parva Güntert, 1942
Pimelodella parva Güntert, 1942: 34, fig. 4. Type locality: Río Paraguay, Paraguay. Holotype: NMBA 5302.
Distribution: Paraguay River basin, Paraguay (Bockmann & Guazzelli, 2003).
Pimelodella pectinifer Eigenmann & Eigenmann, 1888
Pimelodella pectinifer Eigenmann & Eigenmann, 1888b: 132. Type locality: Campos [Brazil]. Holotype: MCZ
7508.
Distribution: Muriaé River basin, Brazil (Bockmann & Guazzelli, 2003).
Pimelodella peruana Eigenmann & Myers, 1942
Pimelodella peruana Eigenmann & Myers, in Eigenmann & Allen, 1942: 101, pl. 3 (fig. 5). Type locality: Inahuaya,
Rio Ucayali [Peru]. Holotype: CAS 63721.
Distribution: Ucayali River basin, Peru (Bockmann & Guazzelli, 2003).
Pimelodella peruensis Fowler, 1915
Pimelodella peruense Fowler, 1915a: 214, fig. 4. Type locality: Peruvian Amazon. Holotype: ANSP 21932.
Distribution: Upper Amazon River basin, Peru (Bockmann & Guazzelli, 2003).
Pimelodella procera Mees, 1983
Pimelodella procera Mees, 1983: 49, fig. 3. Type locality: French Guiana: Crique Balaté. Holotype: RMNH 28588.
Distribution: Maroni River basin, French Guiana (Bockmann & Guazzelli, 2003).
Pimelodella reyesi Dahl, 1964
Pimelodella reyesi Dahl, in Dahl & Medem, 1964: 37, figure on p. 503. Type locality: Río Manso del Sinú, Colombia. Holotype: Whereabouts unknown.
Distribution: Sinú River basin, Colombia (Bockmann & Guazzelli, 2003).
Pimelodella roccae Eigenmann, 1917
Pimelodella roccae Eigenmann, 1917b: 240, pl. 35 (fig. 7). Type locality: Lower Urubamba Valley. Holotype: MCZ
30975. Illustrated and described more fully in Eigenmann & Allen (1942: 98, pl. 10, fig. 4).
Distribution: Urubamba and Beni River basins of Bolivia and Peru (Bockmann & Guazzelli, 2003).
Remarks: Eigenmann & Allen (1942: 98) reported that the holotype came from: Rio Comberciato, 1800 feet elevation.
Pimelodella rudolphi Miranda Ribeiro, 1918
Pimelodella rudolphi Miranda Ribeiro, 1918d: 637. Type locality: Mercado de São Paulo [Brazil]. Lectotype:
MNRJ 857A, designated by Miranda Ribeiro (1953: 404).
Distribution: São Paulo State, Brazil (Bockmann & Guazzelli, 2003).
Pimelodella serrata Eigenmann, 1917
Pimelodella serrata Eigenmann, 1917b: 235, pl. 29 (fig. 1). Type locality: San Joaquin, Bolivia. Holotype: FMNH
57979.
Distribution: Possibly from the Machupo River basin in upper Guaporé River drainage, Bolivia (Bockmann & Guaz-

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zelli, 2003).
Pimelodella spelaea Trajano, Reis & Bichuette, 2004
Pimelodella spelaea Trajano, Reis & Bichuette, 2004: 317, fig. 1. Type locality: Brazil, State of Goiás, São Domingos, upper Tocantins River basin, subterranean stream tributary to São Bernardo River inside São Bernardo
Cave, 13º49'S, 46º21'W. Holotype: MZUSP 81726.
Distribution: subterranean stream tributary to São Bernardo River inside São Bernardo Cave, Tocantins River basin,
Brazil (Trajano et al., 2004: 319).
Pimelodella steindachneri Eigenmann, 1917
Pimelodella steindachneri Eigenmann, 1917b: 237. Type locality: Maues, Rio Madeira... Para... Cudajas... Santarem... Rio Puty... Manacapurú [Brazil]. Syntypes: MCZ 7542a, MCZ 7566a–b, MCZ 7567a, MCZ 7547a,
MCZ 7587a, MCZ 7588a.
Distribution: Amazon River basin, Brazil (Bockmann & Guazzelli, 2003).
Pimelodella taeniophora (Regan, 1903)
Pimelodus (Pimelodella) taeniophorus Regan, 1903b: 625. Type locality: Descalvados, Matto Grosso [Brazil]. Syntypes: BMNH 1895.5.17.27–28 (2).
Distribution: Paraguay River basin, Brazil (Bockmann & Guazzelli, 2003).
Pimelodella taenioptera Miranda Ribeiro, 1914
Pimelodella taenioptera Miranda Ribeiro, 1914: 5, figure on p. 5. Type locality: Tapyrapoan, Rio Sepotuba [MatoGrosso, Brazil]. Lectotype: MNRJ 691A, designated by Miranda Ribeiro (1953: 404).
Distribution: Upper Paraguay River basin, Brazil (Bockmann & Guazzelli, 2003).
Pimelodella tapatapae Eigenmann, 1920
Pimelodella tapatapae Eigenmann, 1920a: 5. Type locality: Mouth of Rio Tapa Tapa, Venezuela. Holotype: CAS
57469.
Distribution: Tapa Tapa River basin in Lake Valencia drainage, Venezuela (Bockmann & Guazzelli, 2003).
Pimelodella transitoria Miranda Ribeiro, 1907
Pimelodella transitoria Miranda Ribeiro, 1907a: 186. Type locality: Ribeirão do Alambary —Iporanga [Brazil].
Holotype: at MNRJ.
Distribution: Ribeira de Iguape River basin, Brazil (Bockmann & Guazzelli, 2003).
Pimelodella vittata (Lütken, 1874)
Pseudorhamdia vittatus Lütken, 1874c: 34. Type locality: in flumine Rio das Velhas, in rivulis affluentibus, lacusculisque vicinis [Brazil]. Syntypes: ZMB 9175, ZMUC P 29652–ZMUC P 29657 (1 each).. Illustrated anddescribed in more detail in Lütken (1875: 173–174, figure on p. 173).
Distribution: Upper São Francisco River basin, Brazil (Bockmann & Guazzelli, 2003).
Pimelodella wesselii (Steindachner, 1877)
Pimelodus (Pseudorhamdia) Wesselii Steindachner, 1877b: 614, footnote. Type locality: Essequibo [Guyana]. Holotype: NMW 79188.
Distribution: Essequibo River basin, Guyana (Bockmann & Guazzelli, 2003).
Pimelodella witmeri Fowler, 1941
Pimelodella witmeri Fowler, 1941a: 133, figs. 20–22. Type locality: Rio Jaguaribe, Orós, Ceará [Brazil]. Holotype:
ANSP 69383.
Distribution: Jaguaribe River basin, Brazil (Bockmann & Guazzelli, 2003).
Pimelodella yuncensis Steindachner, 1902
Pimelodella yuncensis Steindachner, 1902: 135. Type locality: Pacasmayo, Nord Peru. Holotype: at NMW.
Distribution: Jequetepeque River basin, Peru (Bockmann & Guazzelli, 2003).
Species inquirenda, Pimelodella
Pimelodus bahianus Castelanu, 1855: 35, pl. 16, fig. 2. Type locality: eaux douces des environs de Bahia [Brazil]. Syntypes: MNHN b-0612 (2). Tentatively assigned to Pimelodella in Silfvergrip (1996: 16). Not treated in Bockmann &Guazzelli, 2003).

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Pimelodella rendahli Ahl, 1925: 106. Type locality: [No locality stated]. Holotype: ZMB 32031; known only from holotype.
RHAMDELLA Eigenmann & Eigenmann, 1888
Rhamdella Eigenmann & Eigenmann, 1888b: 129. Type species: Rhamdia eriarcha Eigenmann & Eigenmann,
1888. Type by original designation. Gender: Feminine. Originally proposed as a subgenus of Rhamdia.
Rhamdella aymarae Miquelarena & Menni, 1999
Rhamdella aymarae Miquelarena & Menni, 1999: 203, fig. 1. Type locality: Argentina, Salta Province, Río Itiyuro,
500 m below the Itiyuro Dam, route 34 between Estación Pocitos and Araguay, 22º07'S, 63º11'W. Holotype:
ILPLA 611.
Distribution: Itiyuro River basin, Argentina (Bockmann & Guazzelli, 2003).
Rhamdella eriarcha (Eigenmann & Eigenmann, 1888)
Rhamdia eriarcha Eigenmann & Eigenmann, 1888b: 129. Type locality: Rio Grande do Sul [Brazil]. Holotype:
MCZ 27272.
Rhamdella straminea Cope, 1894: 93, pl. 8 (fig. 10). Type locality: Rio Grande do Sul [Brazil]. Syntypes (5): Possibly ANSP 21604 (1), ANSP 39835–36 (2), ANSP 21581–84 (4), ANSP 23216 (1).
Rhamdella lemai Bertoletti, 1967: 78, fig. 2. Type locality: Pôrto Alegre, Guaíba River (Ponta Grossa) [Brazil].
Holotype: MRCN 1897.
Distribution: Jacuí River basin, Brazil (Bockmann & Guazzelli, 2003).
Rhamdella exsudans (Jenyns, 1842)
Pimelodus exsudans Jenyns, 1842 in Jenyns, 1840–42: 111. Type locality: ? Rio de Janeiro [Brazil]. Syntypes:
Department of Zoology, University of Cambridge V.132 (2).
Distribution: Rio de Janeiro State, Brazil (Bockmann & Guazzelli, 2003).
Rhamdella gilli (Starks, 1906)
Rhamdia gilli Starks, 1906: 769, pl. 65 (fig. 1). Type locality: Eten, Peru, in the Rio Eten. Holotype: USNM 53472.
Distribution: Eten River basin, Peru (Bockmann & Guazzelli, 2003).
Rhamdella ignobilis Steindachner, 1907
Rhamdella ignobilis Steindachner, 1907e: 484. Type locality: Flusse Cubataõ um Staate Santa Catharina bei Theresopolis (Brasilien). Syntypes: at NMW.
Distribution: Cubatão River basin in Santa Catarina State, Brazil (Bockmann & Guazzelli, 2003).
Rhamdella jenynsii (Günther, 1864)
Pimelodus jenynsii Günther, 1864: 128. Type locality: Rio de Janeiro [Brazil]. Type(s): at Department of Zoology,
University of Cambridge. Name based on one or more specimens identified in Jenyns (1840–42: 110) as
Pimelodus gracilis, which were deposited at the University of Cambridge.
Distribution: Rio de Janeiro State, Brazil (Bockmann & Guazzelli, 2003).
Rhamdella leptosoma Fowler, 1914
Rhamdella leptosoma Fowler, 1914: 260, fig. 12. Type locality: Rupununi River, British Guiana ... in the highlandsof British Guiana, approximately secured in North Latitude 2º to 3º, and West Longitude 50º20'. Holotype:
ANSP 39340.
Distribution: Rupununi River basin, Guyana (Bockmann & Guazzelli, 2003).
Rhamdella longipinnis Borodin, 1927
Rhamdella longipinnis Borodin, 1927b: 6. Type locality: Prov. St. Paulo, Brazil. Holotype: AMNH 8642.
Distribution: São Paulo State, Brazil (Bockmann & Guazzelli, 2003).
Rhamdella longiuscula Lucena & da Silva, 1991
Rhamdella longiuscula Lucena & da Silva, 1991: 32, figs. 2a, 3a. Type locality: Brasil: Rio Grande do Sul: arroio
Passo do Alto, próximo a localidade de Mineral, São Nicolau (aproximadamente 55º20'O — 28º08'S). Holotype: MCP 14441.
Distribution: Middle Uruguay River basin, Brazil (Bockmann & Guazzelli, 2003).
Rhamdella montana Eigenmann, 1913
Rhamdella montana Eigenmann, 1913: 421, pl. 32 (fig. 1). Type locality: Queta. Small brook in highlands southeast

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of Tarma [Peru]. Holotype: FMNH 56067.
Distribution: Ucayali River basin, Peru (Bockmann & Guazzelli, 2003).
Rhamdella papariae Fowler, 1941
Rhamdella papariae Fowler, 1941a: 135, figs. 27–30. Type locality: Lago Papary, Rio Grande do Norte, Brazil.
Holotype: ANSP 69387.
Distribution: Lake Parary basin (?), Brazil (Bockmann & Guazzelli, 2003).
Rhamdella robinsoni Fowler, 1941
Rhamdella robinsoni Fowler, 1941a: 135, figs. 23–26. Type locality: São José do Egito, Pernambuco [Brazil]. Holotype: ANSP 69386.
Distribution: Lower São Francisco River basin, Brazil (Bockmann & Guazzelli, 2003).
Rhamdella rusbyi Pearson, 1924
Rhamdella rusbyi Pearson, 1924: 12, pl. 2 (fig. 4). Type locality: Rio Colorado, Lower Bopi, Bolivia. Holotype:
CAS 63728.
Distribution: Beni River basin, Bolivia (Bockmann & Guazzelli, 2003).
Rhamdella wolfi Fowler, 1941
Rhamdella wolfi Fowler, 1941a: 136, figs. 31–34. Type locality: Rio Choró, Ceará [Brazil]. Holotype: ANSP 69388.
Distribution: Choró River basin, Brazil (Bockmann & Guazzelli, 2003).
RHAMDIA Bleeker, 1858
Pteronotus Swainson, 1839: 190, 309. Type species: Heterobranchus 5-tentaculatus Sp. Ag. [sic Heterobranchus 6tentaculatus Spix & Agassiz, 1829; = Heterobranchus sextentaculatus ]. Type by original designation. Gender:
Masculine. Preoccupied by Pteronotus Gray, 1837 in Mammalia, not replaced. See Miller (1905) for comments.
Rhamdia Bleeker, 1858b: 197, 207. Type species: Pimelodus sebae Cuvier, 1829. Type by subsequent designation
by Gill (1861c: 48). Gender: Feminine.
Pimelenotus Gill, 1858: 387. Type species: Pimelenotus vilsoni Gill, 1858. Type by subsequent designation by
Eigenmann & Eigenmann (1890: 116). Gender: Masculine. Listed a Pimelotus on p. 387, but corrected inprinted corrigenda to Pimelenotus.
Caecorhamdia Norman, 1926c: 325. Type species: Caecorhamdia urichi Norman, 1926. Type by monotypy. Gender: Feminine.
Revision: Silfvergrip (1996).
Phylogeny: Perdices et al. (2002, Central American species).
Remarks: The name Rhamdia micayi Eigenmann (in Pearson, 1924: 11) has been treated in some literature as available, and as a valid species. However, the name was originally published without distinguishing features (Bockmann & Guazzelli, 2003) and has apparently not yet been made available.
Rhamdia enfurnada Bichuette & Trajano, 2005
Rhamdia enfurnada Bichuette & Trajano, 2005: 589, fig. 1. Type locality: Brazil, southwestern State of Bahia, Serrado Ramalho, município de Coribe, Povoado de Descoberto, Gruna do Enfurnado (13º38'44"S 44º12'05.9"W,elevation 628 m), middle São Francisco River basin. Holotype: MZUSP 87776.
Distribution: Gruna do Enfurnado (13º38'44"S 44º12'05.9"W, elevation 628 m), Serra do Ramalho, Bahia State,
middle São Francisco River basin, northeastern Brazil (Bichuette & Trajano, 2005).
Rhamdia foina (Müller & Troschel, 1848)
Pimelodus foina Müller & Troschel, in Schomburgk, 1848: 628. Type locality: an den felsenreichen Stellen desTakutu [Guyana]. Holotype: ZMB 3039. Also cited as new in Müller & Troschel (1849: 5).
Rhamdia holomelas rupununi Fowler, 1914: 258, fig. 11. Type locality: Rupununi River, British Guiana ... in thehighlands of British Guiana, approximately secured in North Latitude 2º to 3º, and West Longitude 50º20'.
Holotype: ANSP 39339.
Distribution: Essequibo River, Guyana; Branco, Negro, Tocantins, and Trombetas River basins, Brazil (Bockmann& Guazzelli, 2003).
Rhamdia guasarensis Do Nascimiento, Provenzano & Lundberg, 2004
Rhamdia guasarensis Do Nascimiento, Provenzano & Lundberg, 2004: 565, figs. 1–4. Type locality: Surgencia del

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Tigre at 2.5 km W of Cerro Yolanda, Río Guasare basin, Sierra de Perijá, Estado Zulia, Venezuela (10º52'53"N,
72º30'03"W. Holotype: MBUCV V-29604.
Distribution: Guasare River basin, northwestern Venezuela (Do Nascimiento et al., 2004).
Rhamdia humilis (Günther, 1864)
Pimelodus humilis Günther, 1864: 129. Type locality: Venezuela. Lectotype: BMNH 1965.2.19.1, designated by Silfvergrip (1996: 25).
Rhamdia guairensis Eigenmann, 1920a: 6. Type locality: Rio Guaire near Caracas [Venezuela]. Lectotype: CAS
76655, illustrated in, and designated by, Silfvergrip (1996: 34, pl. 1, figs. 3, 4).
Nannorhamdia benedettii Fernández-Yépez & Martín Salazar, 1952: 35, figure on p. 37. Type locality: Quebrada
Ojo de Agua, Baruta [Venezuela]. Holotype: MHNLS 82.
Distribution: Rivers around Caracas, along the Caribbean coast of Venezuela (Bockmann & Guazzelli, 2003).
Rhamdia itacaiunas Silfvergrip, 1996
Rhamdia itacaiunas Silfvergrip, 1996: 83, pl. 2 (fig. 1). Type locality: Brazil, Est. Amazonas, Rio Tocantins basin,‘Igarapé Repartimento’. Holotype: INPA 7985.
Distribution: Middle Tocantins River basin, Brazil (Bockmann & Guazzelli, 2003).
Rhamdia jequitinhonha Silfvergrip, 1996
Rhamdia jequitinhonha Silfvergrip, 1996: 84, pl. 3 (fig. 2). Type locality: Brasil, Est. Minas Gerais, Rio Araçuaí,Santa Rita. Holotype: MZUSP 38630.
Distribution: Jequitinhonha River basin, Brazil (Bockmann & Guazzelli, 2003).
Rhamdia laluchensis Weber, Allegrucci & Sbordoni, 2003
Rhamdia laluchensis Weber, Allegrucci & Sbordoni, 2003: 275, fig. 1. Type locality: Mexico: Chiapas: Municipio
Ocozocoautla, subterranean watercourse in Sistema de La Lucha (17º03'40"N, 93º53'23"W), entrance approximately 5 km southwest of village of Aguablanca situated at extreme eastern shore of Malpaso Reservoir (Presa
Nezahualcoyotl). Holotype: ZMH 9373.
Distribution: Chiapas, Mexico; known only from type locality (Weber et al., 2003).
Rhamdia laticauda (Kner, 1857)
Pimelodus laticaudus Kner, 1857: 420. Type locality: Mexico. Syntypes: at NMW (not found; Silfvergrip, 1996:
16).
Pimelodus hypselurus Günther, 1864: 126. Type locality: Mexico. Holotype: BMNH 1858.11.22.32.
Pimelodus motaguensis Günther, 1864: 127. Type locality: Rio Motagua. Holotype: BMNH 1865.4.29.39.
Pimelodus petenensis Günther, 1864: 126. Type locality: Lake Peten. Holotype: BMNH 1864.1.26.371.
Pimelodus polycaulus Günther, 1864: 131. Type locality: Rio San Geronimo, Guatemala. Holotype: BMNH
1864.1.26.93.
Pimelodus salvini Günther, 1864: 130. Type locality: Rio de San Geronimo, Guatemala. Holotype: BMNH
1861.8.2.16.
Pimelodus (Rhamdia) brachypterus Cope, 1867: 404. Type locality: Orizava, Mexico. Holotype: ANSP 16471.
Rhamdia sacrificii Barbour & Cole, 1906: 156, pl. 2. Type locality: Sacrificial Cenote, near Chichen Itza, Yucatan[Mexico]. Lectotype: MCZ 29073, designated by Silfvergrip (1996: 30).
Rhamdia cabrerae Meek, 1906: 93. Type locality: Amatitlan, Guatemala. Holotype: FMNH 5501.
Pimelodus brachycephalus Regan, 1907b: 258. Type locality: Guatemala, Rio Nacasil. Lectotype: BMNH
1875.6.9.5, designated by Silfvergrip (1996: 31).
Pimelodus Rogersi Regan, 1907b: 259. Type locality: Costa Rica, Irazu. Lectotype: BMNH 1907.2.11.1, designatedby Silfvergrip (1996: 31).
Rhamdia regani Meek, 1907b: 144. Type locality: Turrialba, Costa Rica. Holotype: FMNH 6019.
Rhamdia underwoodi Regan, 1907, in Regan, 1906–08: 135, pl. 23 (fig. 4). Type locality: Costa Rica, Juan Viñas.
Lectotype: BMNH 1907.6.28.33, designated by Silfvergrip (1996: 32).
Rhamdia amatitlanensis Fowler, 1936a: 518, fig. 10. Type locality: Stream issuing from Lake Amatitlan [Guatemala]. Holotype: ANSP 64136.
Rhamdia laticauda typhla Greenfield, Greenfield & Woods, 1982: 564, fig. 1. Type locality: A cave in Belize, in anarea known as Las Cuevas, near Millionario in the Mountain Pine Ridge (16º45'N 89º00'W). Holotype: FMNH

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71605.
Rhamdia zongolicensis Wilkens, 1993: 375, fig. 1. Type locality: Cueva del Ostoc, Sierra de Zongolica (Veracruz,
Mexico), road from Comalapa to Tezonapa. Holotype: ZMH 9249.
Distribution: Atlantic slope river basins from central Mexico to northern Panama, in (Miller et al., 2005).
Rhamdia laukidi Bleeker, 1858
Rhamdia laukidi Bleeker, 1858b: 208. Type locality: Venezuela, Est. Amazonas, first Río Casiquiare caño, ca. 5 min.
from confluence of Río Casiquiare and Río Orinoco left side, Caño Caripo, ca. 3 km from mouth; 03 06'N, 65
50'W. Neotype: ANSP 174652, illustrated in, and designated by, Silfvergrip (1996: 17, pl. 3, figs. 3, 4).
Pimelodus holomelas Günther, 1863: 442. Type locality: Essequibo [Guyana]. Lectotype: BMNH 1864.1.21.8, designated by Silfvergrip (1996: 23).
Rhamdia tenella Eigenmann & Eigenmann, 1888b: 127. Type locality: Cudajas [Brazil]. Holotype: MCZ 7547.
Distribution: Amazon, Essequibo, and Orinoco River basins (Bockmann & Guazzelli, 2003).
Rhamdia macuspanensis Weber & Wilkens, 1998
Rhamdia macuspanensis Weber & Wilkens, 1998: 1000, fig. 3. Type locality: Watercourse in the Grutas de Agua
Blanca, Balneario de Agua Blanca at km 64.9 of the road 186 (carretera federal) from Villahermosa to Chetumal, ca. 20 km se. of Macuspana, Tabasco, Mexico (17º37'15.5"N, 92º28'22.6"W). Holotype: ZMH 8874.
Distribution: Grutas de Agua Blanca, at the northern edge of the Sierra del Norte de Chiapas, Tabasco, Mexico(Weber & Wilkens, 1998).
Rhamdia muelleri (Günther, 1864)
Pimelodus mülleri Günther, 1864: 119. Type locality: Para, Brazil; River Capin, Para, Brazil; Surinam [restricted toRiver Capin by lectotype designation]. Lectotype: BMNH 1849.11.8.101 (133.6 mm SL), designated by Silfvergrip (1996: 23).
Pimelodus (Rhamdia) Knerii Steindachner, 1877b: 631, footnote. Type locality: Marabitanos [Brazil]. Lectotype:
NMW 45790, designated by Silfvergrip (1996: 27).
Rhamdia obesa Eigenmann & Eigenmann, 1888b: 124. Type locality: Teffé [Brazil]. Holotype: MCZ 7518.
Distribution: Amazon, Ampyiacu, Aripuanã, Capim, Essequibo, Madeira, Negro, Orinoco, Solimões, Tocantins andTrombetas River basins (Bockmann & Guazzelli, 2003).
Remarks: Lectotype of Pimelodus mülleri Günther, 1864, incorrectly listed by Silfvergrip (1996) and elsewhere as:
BMNH 1849.11.8.
Rhamdia nicaraguensis (Günther, 1864)
Pimelodus nicaraguensis Günther, 1864: 125. Type locality: Lake of Nicaragua. Holotype: BMNH 1864.1.26.212.
Pimelodus managuensis Günther, 1867c: 603. Type locality: Lake Managua [Nicaragua]. Holotype: BMNH
1865.7.20.37.
Rhamdia alfaroi Fowler, 1932b: 382, figure on p. 381. Type locality: Escobal, 400 meters elevation, Costa Rica.
Holotype: ANSP 53934.
Rhamdia luigiana Villa, 1977: 133, fig. 1. Type locality: Nicaragua: Departamento de Granada; Lake Nicaragua,
between Ometepe and Zapatera Islands. Holotype: USNM 217535.
Distribution: Rivers of Costa Rica and Nicaragua (Bockmann & Guazzelli, 2003).
Rhamdia parryi Eigenmann & Eigenmann, 1888
Rhamdia Parryi Eigenmann & Eigenmann, 1888b: 130. Type locality: Rio Zanaleneo, near Tonala, Chiapas, Mexico. Lectotype: MCZ 27273, designated by Silfvergrip (1996: 29).
Distribution: Pacific draining rivers of Central America, from Tapanatepec River, Mexico, to southern Guatemala(Miller et al., 2005).
Remarks: Treated by Silfvergrip (1996) and Bockmann & Guazzelli (2003) as a synonym of Rhamdia laticauda(Kner), but valid in Miller et al. (2005).
Rhamdia poeyi Eigenmann & Eigenmann, 1888
Rhamdia poeyi Eigenmann & Eigenmann, 1888b: 127. Type locality: Goyaz [Brazil]. Holotype: MCZ 8196.
Distribution: Mamoré, and Tocantins River basins, Brazil, and upper Napo River, Ecuador (Bockmann & Guazzelli,2003).
Rhamdia quelen (Quoy & Gaimard, 1824)

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? Silurus quadrimaculatus Bloch, 1794: 37, pl. 368 (fig. 2). Type locality: Amerika. Holotype: ZMB 2944 (probably
lost; Silfvergrip, 1996: 9).
? Silurus erythropterus Bloch, 1794: 42, pl. 369, fig. 2. Type locality: Amerika. Syntypes: ZMB 3048 (1), ZMB
8786 (1).
Pimelodus quelen Quoy & Gaimard, 1824, in Freycinet, 1824–1825: 228, pl. 49 (fig. 3). Type locality: Peru, Depto
Loreto, right bank quebradita tributary to R. Samiria between Caño Pastos and Hamburgo, by neotype designation. Neotype: NRM 16091, illustrated in, and designated by, Silfvergrip (1996: 12, 97, 139: pl. 7, figs. 2, 3).
Pimelodus sebae Cuvier, 1829: 294. Type locality: Peru, Depto Loreto, right bank quebradita tributary to R. Samiria
between Caño Pastos and Hamburgo, by neotype designation. Neotype: NRM 16091, designated by Silfvergrip(1996: 13). Name made available by reference to Seba (1734–1765: pl. 29, fig. 5), for which the underlying
specimen is apparently lost.
Pimelodus namdia Cuvier, 1829: 294. Type locality: Peru, Depto Loreto, right bank quebradita tributary to R.
Samiria between Caño Pastos and Hamburgo, by neotype designation. Neotype: NRM 16091, designated by
Silfvergrip (1996: 12).
? Heterobranchus sextentaculatus Spix & Agassiz, 1829: 28, pl. 11. Type locality: Peru, Depto Loreto, right bankquebradita tributary to Río Samiria between Caño Pastos and Hamburgo, by neotype designation. Neotype:
NRM 16091, designated by Silfvergrip (1996: 13). As Heterobranchus 6-tentaculatus on caption to plate.
Pimelodus sapo Valenciennes, 1835 (in Valenciennes, 1835–47): pl. 2 (fig. 6). Type locality: [not stated]. Holotype:
MNHN 0000-1576. Name available from plate; described in Cuvier & Valenciennes (1840b: 179; 133 of Strasbourg deluxe edition), with type locality given as: Buénos-Ayres.
Pimelodus Hilarii Valenciennes, in Cuvier & Valenciennes, 1840b: 180 (134 in Strasbourg deluxe edition). Type
locality: les riviéres qui se jettent dans celle de Saint-François au Brésil … Monté-Vidéo [restricted to: SaintFrançois, by lectotype designation]. Lectotype: MNHN a-9415, designated by Silfvergrip (1996: 14, 147).
Pimelodus Pentlandii Valenciennes, in Cuvier & Valenciennes, 1840b: 183 (135 in the Strasbourg deluxe edition),pl. 435. Type locality: de l’Apurimac. Lectotype: BMNH 1862.11.15.11, designated by Silfvergrip (1996: 14,
139).
Pimelodus Stegelichii Müller & Troschel, in Schomburgk, 1848: 628. Type locality: Britisch-Guiana. Lectotype:
ZMB 3043 (182.2 mm SL), designated by Silfvergrip (1996: 15). Also reported as new, with types listed as
coming from Surinam in Müller & Troschel (1849: 3).
Pimelodus Sellonis Müller & Troschel, 1849: 2. Type locality: Brasilien. Holotype: ZMB 3041.
Pimelodus Deppei Müller & Troschel, 1849: 3. Type locality: Sandwich-Inseln [in error; (Silfvergrip, 1996: 15)
indicated that the specimens were probably from southern Mexico]. Lectotype: ZMB 3046, designated by Silfvergrip (1996: 15, 133).
Pimelodus musculus Müller & Troschel, 1849: 4. Type locality: America. Holotype: ZMB 3048. Holotype is also a
syntype of Silurus erythropterus Bloch, 1794.
Pimelenotus Vilsoni Gill, 1858: 391. Type locality: Trinidad. Holotype: possibly USNM 5927 (Silfvergrip, 1996:
22).
Pimelodus cinerascens Günther, 1860: 237, pl. 10 (fig. a). Type locality: Fresh waters of Guayaquil and Esmeraldas,
Ecuador. Lectotype: BMNH 1860.6.16.193, designated by Silfvergrip (1996: 22, 134).
Pimelodus guatemalensis Günther, 1864: 122. Type locality: Guatemala, Huamuchal. Lakes with brackish water.
Lectotype: BMNH 1864.1.26.210, designated by Silfvergrip (1996: 24).
Pimelodus wuchereri Günther, 1864: 123. Type locality: Bahia. Lectotype: BMNH 1864.1.19.18, designated by Silfvergrip (1996: 24).
Pimelodus godmanni Günther, 1864: 124. Type locality: Guatemala, Mexico … Lower Vera Paz … Rio Motagua …
Mexico. Syntypes: BMNH 1856.3.17.39–40 (2), BMNH 1856.3.17.38 (1), BMNH 1864.1.26.99–100 (2). Lectotype designation of BMNH 1864.1.26.94 by Silfvergrip (1996: 24, 133), is invalid, inasmuch as the statedlocality of that specimen: Rio San Geronimo is not one of the localities reported in Günther and, therefore, thespecimen was not one of the original syntypes.
? Pimelodus micropterus Günther, 1864: 124. Type locality: Guatemala … River of San Geronimo. Holotype:
BMNH 1864.1.26.92.

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lost (Silfvergrip, 1996: 25). Silfvergrip (1996: 25) interpreted the locality of the original types as western Mexico.
Pimelodus wagneri Günther, 1868c: 474. Type locality: Pacific and Atlantic rivers of Panama. Type(s): at NMW.
Type series may include NMW 45618, 45619 (Silfvergrip, 1996: 26). Species based on specimens from “RioChagres, Guajaquil und Neu-Granada” in Kner & Steindachner (1864: 52) identified as Pimelodella cinerascens.
Rhamdia dorsalis Gill, 1870: 94. Type locality: the Maranon, or Upper Amazon, and Napo Rivers. Holotype:
USNM 35334.
Pimelodus (Rhamdia) Parahybae Steindachner, 1877b: 615. Type locality: Rio Parahyba, Brasilien. Syntypes:
NMW 45852.
Pimelodus (Rhamdia) Queleni cuprea Steindachner, 1877b: 623. Type locality: aus dem Parahyba bei Juiz de Fora
(im mittleren lauf des Stromes). Lectotype: NMW 45877: 1, designated by Silfvergrip (1996: 146). Originallyas Pimelodus (Rhamdia) Queleni (var. cuprea).
Pimelodus (Rhamdia) Cuyabae Steindachner, 1877b: 633, footnote. Type locality: Cuyaba. Lectotype: NMW
45919: 1, designated by Silfvergrip (1996: 28).
Rhamdia bransfordii Gill, 1877: 337. Type locality: Panama … Camp Marie Caretta. Holotype: USNM 16674.
Pimelodus bathyurus Cope, 1878: 674. Type locality: Peruvian Amazon. Lectotype: ANSP 21437, designated by
Silfvergrip (1996: 28).
Rhamdia oaxacae Meek, 1902: 74, pl. 14. Type locality: Rio Quiotepec, Cuicatlan, Oaxaca, Mexico. Holotype:
FMNH 3716 (Originally published as 3717, see FMNH online catalog).
Rhamdia depressa Barbour & Cole, 1906: 155, pl. 1. Type locality: Ikil Cenote, near Chichen-Itza, Yucatan [Mexico]. Lectotype: MCZ 29072, designated by Silfvergrip (1996: 30, 133).
Rhamdia barbata Meek, 1907a: 106. Type locality: San Francisco, Guatemala. Holotype: FMNH 5906.
Pimelodus Boucardi Regan, 1907b: 258. Type locality: Yucatan. Holotype: BMNH 1880.7.13.33.
Rhamdia heteracantha Regan, 1907, in Regan, 1906–08: 134. Type locality: Costa Rica, Juan Viñas. Holotype:
BMNH 1907.6.28.32.
Rhamdia nasuta Meek, 1909: 207. Type locality: Buenos Aires de Terraba, Costa Rica. Holotype: FMNH 6480.
Rhamdia branneri Haseman, 1911b: 377, pl. 75. Type locality: Creek of the Rio Iguassú near Serrinha, Paraná, Brazil. Holotype: FMNH 54235.
Rhamdia branneri voulezi Haseman, 1911b: 378, pl. 76. Type locality: Porto União da Victoria, Rio Iguassú. Holotype: FMNH 54238.
Rhamdia mounseyi Regan, 1913c: 282. Type locality: River Ucayali, Peru. Lectotype: BMNH 1913.7.30.13, designated by Silfvergrip (1996: 33, 139).
Rhamdia riojae Fowler, 1915a: 209, fig. 2. Type locality: Rioja, near Moyabamba and Baka Puerto, on or near the
lower course of the Huallagua River, Peru. Holotype: ANSP 21101.
Rhamdia ortoni Fowler, 1915a: 211, fig. 3. Type locality: Peruvian Amazon. Holotype: ANSP 21928.
Rhamdia microps Eigenmann, in Eigenmann & Fisher, 1917: 394, pl. 38. Type locality: Uruguayana. Holotype:
FMNH 58285.
Rhamdia pubescens Miranda Ribeiro, 1920: 11, sixth unnumbered plate. Type locality: Urucum (proximo deCorumbá), Matto-Grosso [Brazil]. Lectotype: MNRJ 925A, designated by Miranda Ribeiro (1953: 404), but
perhaps specimen not isolated. Lectotype also selected by Silfvergrip (1996: 34), and listed as MNRJ 925.
Silurus ribularis Larrañaga, 1923: 376. Type locality: Peru, Depto Loreto, right bank quebradita tributary to R.
Samiria between Caño Pastos and Hamburgo, by neotype designation. Neotype: NRM 16091, designated by
Silfvergrip (1996: 35). Name spelled Silurus rivularis in Silfvergrip (1996) and Bockmann & Guazzelli (2003),
the latter of which includes a comment that ribularis is a misspelling, but no internal evidence supporting thatclaim was found.
Silurus 9-radiatus Larrañaga, 1923: 385. Type locality: Peru, Depto Loreto, right bank quebradita tributary to R.
Samiria between Caño Pastos and Hamburgo. Neotype: NRM 16091, designated by Silfvergrip (1996: 35).
Apparently an alternate new name for Silurus ribularis Larrañaga, 1923.

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Caecorhamdia urichi Norman, 1926c: 325, fig. 1. Type locality: Pool in interior of the Guacharo Cave, Trinidad.
Lectotype: BMNH 1926.7.28.1 (99.7 mm SL, smaller of 2 specimens), designated by Silfvergrip (1996: 35).
Rhamdia guatemalensis muriei Hubbs, 1935: 7, pl. 4 (fig. 1). Type locality: a stagnant aguada … at edge of swamp,in the high tropical “bush” region at Uaxactun, Petén, Guatemala. Holotype: UMMZ 97881.
Rhamdia guatemalensis decolor Hubbs, 1936: 201, pl. 1 (fig. 3). Type locality: Mexico … San Bulha Cave, Motul,
Yucatan. Holotype: UMMZ 102217.
Rhamdia guatemalensis stygaea Hubbs, 1936: 203, pl. 1 (fig. 2). Type locality: Mexico … San Isidro Cave in Salar
Colony, near Merida. Holotype: UMMZ 102218.
Rhamdia saijaensis Rendahl, 1941: 2. Type locality: Rio Saija, südl. von Buenaventura, zwischen B. und Guapi,
Kolumbiens. Holotype: NRM 10675.
Rhamdia duquei Eigenmann & Pearson, in Eigenmann & Allen, 1942: 93. Type locality: Rio Urubamba, Santa Ana.
Syntypes: SU 57895 (36). Although the original description indicates the type series consisted of a “type” andparatypes, there were no characteristics presented to indicate which of the specimens was considered the holotype. As such, the type series must be treated as syntypes.
Rhamdia sebae Martyi Güntert, 1942: 33. Type locality: Riactis Canâwé, Dept. Ita, Paraguay. Holotype: NMBA
5279.
Rhamdia lehmanni Dahl, 1961: 487, figure on p. 489. Type locality: a small pool in a brook tributary to the Guayabero River, approx. 1500 meters from Expedition’s camp No. 1, and about 1000 meters from the mouth of thebrook, on the left side of the river. Holotype: probably lost (Cala, 1981; Silfvergrip, 1996: 37).
Distribution: Rivers from Mexico to central Argentina (Bockmann & Guazzelli, 2003).
Rhamdia reddelli Miller, 1984
Rhamdia reddelli Miller, 1984: 136, fig. 1. Type locality: Cueva del Nacimiento del Río San Antonio, ca. 9 km SW
of Acatlán, Oaxaca, on Atlantic slope of eastern Mexico. Holotype: UMMZ 211164; illustrated in Miller et al.
(2005: 175, fig. 6-185).
Distribution: Caves of Papaloapan River basin, Atlantic slope, Mexico (Miller et al., 2005).
Remarks: Treated as a synonym of Rhamdia laticauda (Kner) by Silfvergrip (1996) and Bockmann & Guazzelli,2003).
Rhamdia xetequepeque Silfvergrip, 1996
Rhamdia xetequepeque Silfvergrip, 1996: 98, pl. 2 (fig. 3). Type locality: Peru, Jequetepeque River, just W of Chilete (06º19'90"S, 78º25'90"W). Holotype: ROM 70112.
Distribution: Jequetepeque River basin, Peru (Bockmann & Guazzelli, 2003).
RHAMDIOGLANIS Ihering, 1907
Rhamdioglanis Ihering, 1907: 16. Type species: Rhamdioglanis frenatus Ihering, 1907. Type by original designation. Gender: Masculine.
Rhamdioglanis frenatus Ihering, 1907
Rhamdioglanis frenatus Ihering, 1907: 16. Type locality: riverlets of the Island of S. Sebastião, State of S. Paulo.
Lectotype: MZUSP 154, designated by Britski (1969: 205).
Distribution: Streams in the Island of São Sebastião, São Paulo State, Brazil (Bockmann & Guazzelli, 2003).
Rhamdioglanis transfasciatus Miranda Ribeiro, 1908
Rhamdioglanis transfasciatus Miranda Ribeiro, 1908b: [3]. Type locality: Rio Bethary [Brazil]. Syntypes: MNRJ
965 (2).
Distribution: Coastal streams of Brazil from Rio de Janeiro to Santa Catarina State (Bockmann & Guazzelli, 2003).
RHAMDIOPSIS Haseman, 1911
Rhamdiopsis Haseman, 1911b: 375. Type species: Rhamdiopsis moreirai Haseman, 1911. Type by monotypy. Gender: Feminine.
Rhamdiopsis microcephala (Reinhardt, 1874)
Rhamdia microcephala Reinhardt, in Lütken, 1874c: 35. Type locality: in flumine Rio das Velhas. Syntypes: ZMUC
P 29658 (1), ZMUC P 29659 (1), ZMUC P 29660 (1). Illustrated and described in more detail in in Lütken

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FERRARIS

(1875: 177–179, pl. 3, fig. 7).
Distribution: Upper Paraná and São Francisco River basins, Brazil (Bockmann & Guazzelli, 2003).
Rhamdiopsis moreirai Haseman, 1911
Rhamdiopsis moreirai Haseman, 1911b: 375, pl. 73. Type locality: Serrinha Paraná, Rio Iguassú. Holotype: FMNH
54372.
Distribution: Iguaçu and Ribeira de Iguape River basins, Brazil (Bockmann & Guazzelli, 2003).
TAUNAYIA Miranda Ribeiro, 1918
Taunayia Miranda Ribeiro, 1918d: 642. Type species: Taunayia marginata Miranda Ribeiro, 1918. Type by monotypy. Gender: Feminine.
Taunayia bifasciata (Eigenmann & Norris, 1900)
Nannoglanis bifasciatus Eigenmann & Norris, 1900: 350. Type locality: S. Paulo, Brazil. Holotype: CAS 75771.
Taunayia marginata Miranda Ribeiro, 1918d: 642. Type locality: Piquete [Brazil]. Holotype: MZUSP 2273; holotype illustrated in Oliveira & Britski (2000: 129, fig. 1).
Distribution: Upper Paraíba do Sul and Tietê River basins, Brazil (Bockmann & Guazzelli, 2003).
Remarks: Synonymy based on Oliveira & Britski (2000), in which the species is redescribed.
Species inquirenda, Heptapteridae
Pimelodus velifer Humboldt, in Humboldt & Valenciennes, 1821: 171. Type locality: Río Magdalena, Colombia. No
types known.

† HYPSIDORIDAE Grande, 1987
† Hypsidoridae Grande, 1987: 28. Type genus: †Hypsidoris Lundberg & Case, 1970.
1 genus, 2 species, all taxa based only on fossils.
† HYPSIDORIS Lundberg & Case, 1970
† Hypsidoris Lundberg & Case, 1970: 452. Type species: †Hypsidoris farsonensis Lundberg & Case, 1970. Type byoriginal designation. Gender: Feminine.
† Hypsidoris farsonensis Lundberg & Case, 1970.
† Hypsidoris farsonensis Lundberg & Case, 1970: 452, pls. 1– 2; text figs. 1– 2. Type locality: Laney Shale member,
Green River Formation, Sweetwater County, near Farson, Wyoming. Holotype: PU 20570 (impressions of complete skeleton, in dorsal and ventral view); holotype illustrated in Grande (1987: figs. 1–2).
Distribution: Early Middle Eocene Lake Gosiute deposits of the Laney Shale Member (Grande, 1987).
† Hypsidoris oregonensis Grande & de Pinna, 1998
† Hypsidoris oregonensis Grande & de Pinna, 1998: 454, figs. 3–10. Type locality: Clarno Formation, Jefferson
County, north central Oregon. Holotype: FMNH PF10719 (impression of nearly complete skeleton).

ICTALURIDAE Gill, 1861
Ictaluri Gill, 1861c: 49. Type genus: Ictalurus Rafinesque, 1820.
Ameiurina Günther, 1864: 98. Type genus: Ameiurus Rafinesque, 1820.
Ichthaelurinae Meek, 1904: 10. Type genus: Ichthaelurus Cope, 1868.
† Astephinae Grande & Lundberg, 1988: 146. Type genus: † Astephus Cope, 1873.
Reviews: Lundberg (1975, fossil forms).
Identification guide: Page & Burr (1991, Canada and U.S.).
Checklist: Espinosa Pérez, et al. (1993, Mexico).
Phylogeny: Le Grande (1981, chromosomal evolution); Hardman (2002); Lundberg (1982, 1992).

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7 genera, 64 species; 1 fossil genus and 14 fossil species.
AMEIURUS Rafinesque, 1820
Ameiurus Rafinesque, 1820c: 359. Type species: Silurus lividus Rafinesque, 1820. Type designated by plenary powers (ICZN, Opinion 1584; see Bailey & Robins, 1988, for justification). Gender: Masculine. Also described as
new in Rafinesque (1820a: 65). Originally proposed as subgenus of Ictalurus.
Amiurus Agassiz, 1846: 17. Type species: Silurus lividus Rafinesque, 1820. Unjustified emendation of Ameiurus
Rafinesque, 1820. Gender: Masculine.
Gronias Cope, 1864: 231. Type species: Gronias nigrilabris Cope, 1864. Type by monotypy. Gender: Masculine.
Remarks: See Bailey & Robins (1988) for information on type species.
Relationships: Hardman & Page (2003).
Ameiurus brunneus Jordan, 1877
Amiurus brunneus Jordan, 1877a: 366. Type locality: South Fork of Ocmulgee R., at Flat Shoals, Georgia. Lectotype: USNM 20148, designated by Jordan & Evermann (1896: 142).
Distribution: Atlantic coast basins from southern Virginia to Georgia; Gulf Coast basins of Georgia, Alabama andFlorida (Page & Burr, 1991).
Ameiurus catus (Linnaeus, 1758)
Silurus catus Linnaeus, 1758: 305. Type locality: America; Asia [the latter in error]. No types known; based on illustrations in Catesby (1731–1743: pl. 23), and Marcgravius (1648: 173).
Pimelodus albidus Lesueur, 1819: 148. Type locality: la Delaware [River]. Type(s): Whereabouts unknown (Gilbert,
1998).
Pimelodus lynx Girard, 1859b: 160. Type locality: Potomac River, in a deep run, a tributary of Patapsco River, and at
Carlisle, Pa [U. S. A.]. Syntypes: at USNM, but not found.
Ichthaelurus Kevinskii Stauffer, 1869: 578. Type locality: Susquehanna River drainage, Lancaster Co., Pennsylvania. Holotype: Whereabouts unknown.
Ichthaelurus McCaskei Stauffer 1869: 578. Type locality: Shenk's Ferry, Susquehanna River (or tributary thereof),
Lancaster Co., Pennsylvania. Holotype: Whereabouts unknown.
Amiurus lophius Cope, 1870a: 487. Type locality: Washington, D. C. market, which came from the lower course of
the Potomac River. Syntypes (3): ANSP 8461–62 (2).
Amiurus niveiventris Cope, 1870a: 488. Type locality: Neuse River, N. Ca. [North Carolina, U.S.A.]. Syntypes:
ANSP 8466–67 (2).
Distribution: Atlantic and Gulf coast drainages from Hudson River to Pascagoula River, Mississippi; introduced
elsewhere (Page & Burr, 1991).
† Ameiurus hazenensis (Baumgartner, 1982)† Ictalurus (Amiurus) hazenensis Baumgartner, 1982: 38. Type locality: Laminated diatomaceous beds of Miocene
Middle Truckee Formation (ca. 39º 31'36"N, 119º 06'27"W), Two Tips Quadrangle, T19N, R26E, Sec. 7 (center), near Nazen, Churchill County. Nevada. Holotype: UMMP V74320 (anterior portion of fish).
Distribution: Churchill County, Nevada, Middle Truckee Formation; Miocene.
† Ameiurus lavetti (Lundberg, 1975)† Ictalurus lavetti Lundberg, 1975: 31, fig. 4 a–e; pl. 11, fig. a. Type locality: Sec. 7, T. 11 S, R. 37 W, Old Vincent
Ranch, 22 mi. NE of Wallace, Logan County, Kansas; middle Pliocene.
Distribution: Olallala Group, Logan County Kansas, and Basal Ash Hollow formation, South Dakota; lower to middle Pliocene (Lundberg, 1975).
† Ameiurus leidyi (Lundberg, 1975)† Ictalurus leidyi Lundberg, 1975: 35, fig. 4 f; pl. 5, fig. a, pl. 8, fig. k. Type locality: Observation Quarry, NW 1/4,
SE 1/4, Sec. 4, T. 31 N, R. 47 W. Dawes County, Nebraska; early Miocene. Holotype: AMNH 10509 (nearlycomplete skull).
Distribution: Observation Quarry, Nebraska; Valentine Formation, Nebraska; Ash Hollow Formation, Nebraska;
lower Pliocene (Lundberg, 1975).
† Ameiurus macgrewi (Lundberg, 1975)

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FERRARIS† Ictalurus macgrewi Lundberg, 1975: 29, fig. 3; pl. 10, fig. a. Type locality: Banks of Horse Creek, Sec. 3, T. 17 N,
R. 66 W, Laramie County, Wyoming; middle Miocene. Holotype: University of Wyoming 3298.
Distribution: Banks of Horse Creek, Laramie County, Wyoming; middle Miocene (Lundberg, 1975).
Ameiurus melas (Rafinesque, 1820)
Silurus melas Rafinesque, 1820b: 51. Type locality: Ohio River. No types known. Also described as new as Pimelodus melas in Rafinesque (1820a: 66 and 1820c: 360).
Pimelodus catulus Girard, 1858b: 208, pl. 41 (figs. 4–6). Type locality: Fort Smith, Arkansas. Syntypes: USNM 926(1), USNM 927 (5, missing).
Pimelodus confinis Girard, 1859b: 159. Type locality: Root River, near Racine, Wis. Syntypes: MCZ 35993 (1),
USNM 1514 (7), USNM 1515 (3).
Pimelodus cupreoides Girard, 1859b: 159. Type locality: Aux Plaines, Ill. Syntypes: USNM 1497 (4).
Amiurus obesus Gill 1861b: 45. Type locality: Nebraska. Syntypes: USNM 8968 (3).
Amiurus brachyacanthus Cope, 1880: 35. Type locality: Wallace Creek, headwaters of Medina River, Bandera Co.,
Texas. Syntypes: ANSP 20527–28 (2).
Amiurus cragini Gilbert, 1884: 10. Type locality: “The Lake” (a blind arm of the Arkansas River), at Garden City,
Finney Co., Kansas. Holotype: USNM 36814. Also appeared as new (and synonymized in a footnote) in Gilbert(1885: 512).
Distribution: North American Great Lakes, Hudson Bay, Mississippi River basin and Gulf Coast basins; introduced
elsewhere (Page & Burr, 1991).
Ameiurus natalis (Lesueur, 1819)
Pimelodus natalis Lesueur, 1819: 154. Type locality: Haut-Canada. No types known. Placed on Official List (ICZN,
Opinion 1584).
Silurus lividus Rafinesque, 1820b: 50. Type locality: Ohio River. No types known. As Pimelodus lividus in
Rafinesque (1820a: 65; and 1820c: 359).
Silurus lividus Fuscatus Rafinesque, 1820b: 51. Type locality: Ohio River. No types known. Originally as Silurus
lividus var. Fuscatus.
Silurus (Pimelodus) coenosus Richardson, 1836: 132. Type locality: Penetanguisheue, Lake Huron. Holotype:
Whereabouts unknown.
Pimelodus ailurus Girard, 1858b: 210. Type locality: Lake Amelia, near Fort Snelling, Minnesota. Syntypes: USNM
904 (2, originally 1), USNM 905 (1, missing).
Pimelodus antoniensis Girard, 1858b: 209. Type locality: Near San Antonio, Tex. Holotype: USNM 923.
Pimelodus felinus Girard, 1858b: 209. Type locality: Trib. of Gypsum creek, Canadian. Lectotype: USNM 924, designated by Jordan & Evermann (1896: 140).
Pimelodus puma Girard, 1859b: 160. Type locality: Charleston, S. C. [South Carolina]. Holotype: USNM 1536.
Amiurus natalis analis Jordan, 1877c: 87, pl. 18 (fig. 31). Type locality: Little Red River, Judsonia, White Co.,
Arkansas. Holotype: USNM 31088 (missing).
Amiurus erebennus Jordan, 1877c: 85, pl. 13 (figs. 19–20). Type locality: St. John's River, Florida. Holotype: USNM
19093.
Amiurus bolli Cope, 1880: 35. Type locality: Little Wichita River, in northern Texas. Syntypes: ANSP 20512–13 (2).
Amiurus prosthistius Cope, 1884a: 132. Type locality: Batsto River, Mullica System, New Jersey. Syntypes: ANSP
20546–49 (4).
Distribution: Atlantic and Gulf coast drainages, from New York to northern Mexico, Great Lakes and Mississippi
River basin (Page & Burr, 1991).
Remarks: Amiurus erebennus Jordan, 1877, was treated as a valid subspecies in Gilbert (1998).
Ameiurus nebulosus (Lesueur, 1819)
Pimelodus nebulosus Lesueur, 1819: 149. Type locality: Philadelphie .... la Delaware. No types known.
Pimelodus atrarius De Kay, 1842: 185, pl. 36 (fig. 116). Type locality: Wappinger's Creek, trib. to Hudson R.,
Dutchess Co., New York. Syntypes: New York State coll. (whereabouts unknown; Gilbert, 1998).
Pimelodus pullus DeKay, 1842: 184, pl. 37 (fig. 117). Type locality: Lake Pleasant, Lake Janet and other lakes in n.
districts, New York. Syntypes: New York State coll. (whereabouts unknown; Gilbert, 1998).

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Pimelodus vulgaris Thompson, 1842: 138. Type locality: Lake Champlain, Vermont. Holotype: Whereabouts
unknown.
? Pimelodus felis Agassiz, 1850: 281. Type locality: Lake Superior. Type(s): Whereabouts unknown.
Pimelodus marmoratus Holbrook, 1855: 54, pl. 6 (fig. 4). Type locality: Altamaha River, Georgia. Holotype: USNM
9031. As Pimelodus marginatus on table of plates (p. 58).
Pimelodus marginatus Holbrook, 1855: 58, pl. 6 (fig. 4). Type locality: Altamaha River, Georgia. Holotype: USNM
9031. Name made available from table of plates for illustration corresponding to species account of Pimelodus
marmoratus Holbrook.
Pimelodus dekayi Girard, 1859b: 160. Type locality: Oswego, Lake Ontario, and in four mile creek, near Oswego[New York]. Syntypes: USNM 1512 (missing), USNM 1513 (4).
Pimelodus hoyi Girard, 1859b: 159. Type locality: In the neighborhood of Racine, Wis. Syntypes: MCZ 35945 (1),
USNM 1539 (3).
Pimelodus vulpeculus Girard, 1859b: 160. Type locality: Charleston, S.C. [South Carolina]. Holotype: USNM 1530.
Gronias nigrilabris Cope, 1864: 231. Type locality: Conestoga Creek, trib. to Susquehanna River, Lancaster Co.,
Pennsylvania. Syntypes: ANSP 22082–83 (2).
Amiurus mispilliensis Cope, 1870a: 486. Type locality: Mispillion Creek, a sluggish stream in the southern part of
the State of Delaware. Holotype: ANSP 8536.
Ictalurus nebulosus pannonicus Harka & Pinter, 1990: 69, figs. 3–4. Type locality: Tisza R. at Poroszló, Hungary.
Holotype: MNSB 87.1.1. [Name based on an introduced population.]
Distribution: Atlantic and Gulf coast drainages, from Canada to Alabama; Great Lakes, Hudson Bay, and Mississippi River basin; introduced elsewhere in North America (Page & Burr, 1991); also in Europe and New
Zealand.
Remarks: Pimelodus marmoratus Holbrook, 1855, was treated as a valid subspecies, with no synonyms, in Gilbert(1998).
† Ameiurus pectinatus (Cope, 1874)† Ictalurus pectinatus Cope, 1874a: 49. Type locality: Florissant lake beds, Oligocene, Florissant, Colorado. Holotype: USNM 4086.
Distribution: Florissant lake beds, Colorado; Oligocene (Lundberg, 1975).
Ameiurus platycephalus (Girard, 1859)
Pimelodus platycephalus Girard, 1859b: 161. Type locality: In the neighborhood of Anderson, S.C. [South Carolina]. Lectotype: ANSP 8473, designated by Yerger & Relyea (1968: 364).
Distribution: Atlantic drainages of Virginia, the Carolinas, and Georgia (Page & Burr, 1991).
† Ameiurus reticulatus Smith, Morgan & Gustafson, 2000
† Ameiurus reticulatus Smith, Morgan and Gustafson, 2000: 16, figs. 9 b, c. Type locality: Blufftop local fauna, Ringold Formation, Franklin Co., Washington; middle Pliocene. Holotype: UMMP 104009, complete mandible.
Distribution: Ringold Formation, Washington; Miocene (Smith, Morgan and Gustafson, 2000).
† Ameiurus sawrockensis (Smith, 1962)† Ictalurus sawrockensis Smith, 1962: 527, fig. 8. Type locality: Beaver locality, Sawrock Canyon, west side of Sec.
36, T. 34 S, 312 W. Seward County, Kansas; upper Pliocene. Holotype: UMMP V37066 (pectoral-fin spine).
† Ictalurus benderensis Smith, 1962: 527, fig. 9. Type locality: Bender local fauna, Rexroad formation above
Rexroad local fauna, Cottrell pasture NW 1/4, SE 1/4, Sec. 16., T. 33 S, R. 29 W. Mead County, Kansas, upper
Pliocene. Holotype: UMMP V37664 (pectoral-fin spine).
Distribution: Sawrock Canyon local fauna, Rexroad Formation, Kansas; upper Pliocene (Lundberg, 1975).
Ameiurus serracanthus (Yerger & Relyea, 1968)
Ictalurus serracanthus Yerger & Relyea, 1968: 373, fig. 4. Type locality: N. shore of Lake Talquin, 6.8 mi. westsouthwest of Ochlockonee R., Gadsden Co., Florida. Holotype: USNM 231711.
Distribution: Gulf Coast drainages of northern Florida, western Georgia, and eastern Alabama (Page & Burr, 1991).
† Ameiurus vespertinus (Miller & Smith, 1967)† Ictalurus vespertinus Miller & Smith, 1967: 15. Type locality: roadcuts on state highway 45 about 1.5 mi. SE of
Fossil Butte, sec. 16, T. 4 S, R 1 W. Glenns Ferry Formation, lacustrine facies, late Pliocene. Holotype: UMMP

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FERRARISV55561 (right lower jaw).
† Ictalurus peregrinus Lundberg, 1975: 39, fig. 5 a–e; pl. 7, fig. c, d; pl. 8, fig. a. Type locality: Sec. 14, T. 21 S, R.
37 E, Malheur County, Oregon. Upper member of the Juntura Formation, Black Butte local fauna. Holotype:
UO 28035 (pectoral spine).
Distribution: Glenns Ferry Formation; Chalk Hills Formation; Chalk Butte Formation; Rome Beds, Oregon; Juntura
Formation, Black Butte local fauna, Pliocene-Pleistocene (Lundberg, 1975).
Remarks: Synonymy based on Lundberg (1992).
Species inquirenda, Ameiurus
Silurus Xanthocephalus Rafinesque, 1820b: 51. Type locality: Ohio, Kentucky and Licking rivers. No types known. See
also Rafinesque (1820a: 66; and 1820c: 360) as Pimelodus Xanthocephalus.
† ASTEPHUS Cope, 1873
† Astephus Cope, 1873: 638. Type species: Pimelodus antiquus Leidy, 1873. Type by subsequent designation by
Lundberg (1975: 5). Gender: Masculine. Originally proposed as a subgenus of Rhineastes Cope.
Remarks: Placement within the Ictaluridae follows Grande & Lundberg (1988).
Revision: Grande & Lundberg (1988).
Phylogenetic position: Grande & Lundberg (1988).
† Astephus antiquus (Leidy, 1873)† Pimelodus antiquus Leidy, 1873a: 99. Type locality: Bridger Formation, junction of Big Sandy and Green River,Wyoming. Holotype: USNM 2179 (pectoral spine, dentary). Illustrated and described in more detail in Leidy(1873b: 193, pl. 322, figs. 44–45). Holotype also illustrated in Lundberg (1975: fig. 1d; pl. 7, fig. f) and Grande& Lundberg (1988: fig. 1a).
† Rhineastes arcuatus Cope, 1873: 641. Type locality: Bridger Formation, Wyoming. Holotype: USNM 3985 (pectoral spine); holotype illustrated in Grande & Lundberg (1988: fig. 1b).
† Rhineastes calvus Cope, 1873: 640. Type locality: Bridger Formation, Mammoth Buttes, South Bitter Creek, Wyoming. Holotype: USNM 3980 (partial neurocranium); holotype illustrated in Cope (1884b: 65, pl. 5, figs. 3–4),
Lundberg (1975: pl. 4, fig. c), and Grande & Lundberg (1988: fig. 1c).
† Ameiurus primaevus Eastman, 1917: 293. Type locality: not data [almost certainly Green River Shales, Wyoming].
Holotype: USNM 8122 (nearly complete fish); holotype illustrated in Grande & Lundberg (1988: fig. 1e).
Distribution: Bridger Formation and Green River Formation (Lundberg, 1975).
Remarks: Synonymy follows Grande & Lundberg (1988), which treats the brief comments in Leidy (1873a: 99) as
sufficient to make the name Pimelodus antiquus available. However Woodward (1901: 329) treated Rhineastes
arcuatus as valid, with Pimelodus antiquus as a synonym, based on the observation that the account of P. antiquus in Leidy (1873a: 99) did not make the name available (nomen nudum) and the first description by Leidy(1873b: 193) that made the name available was published after that of R. arcuatus Cope.
† Astephus resimus Lundberg, 1975
† Astephus resimus Lundberg, 1975: 9, pl. 4, figs. a, b. Type locality: Bridger B Formation, middle Eocene; Pinnacle
Rock between Little America and Twin Buttes, Bridger Basin, Sweetwater County, Wyoming. Holotype:
USNM 18104 (nearly complete neurocranium); holotype illustrated in Grande & Lundberg (1988: fig. 2).
Distribution: Bridger B formation, Wyoming; middle Eocene; known only from type specimen (Lundberg, 1975).
ICTALURUS Rafinesque, 1820
Ictalurus Rafinesque, 1820c: 355. Type species: Silurus cerulescens Rafinesque, 1820. Type by subsequent designation by Gill (1861a: 49). Gender: Masculine. Originally proposed as a subgenus of Pimelodus.
Elliops Rafinesque, 1820c: 356. Type species: Silurus maculatus Rafinesque, 1820. Type by subsequent designation
by Jordan & Gilbert (1877: 87). Gender: Masculine. Proposed as a section of the subgenus Ictalurus. Also proposed as new in Rafinesque (1820a: 62).
Synechoglanis Gill, 1859a: 39. Type species: Synechoglanis beadlei Gill, 1859. Type by monotypy. Gender: Mascu-

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line.
Ichthaelurus Cope, 1868: 237. Type species: Silurus cerulescens Rafinesque, 1820. Type by being a replacement
name. Unjustified emendation of Ictalurus Rafinesque, 1820. Gender: Masculine.
Villarius Rutter, 1896: 256. Type species: Villarius pricei Rutter, 1896. Type by subsequent designation by Jordan &Evermann (1898: 2789). Gender: Masculine.
Haustor Jordan & Evermann, 1896: 135. Type species: “Gadus lacustris Walbaum, 1792”, now Pimelodus nigricans
Le Sueur, 1819 [see Remarks]. Type fixed by provision of Art. 70.3 (ICZN, 1999). Gender: Masculine. Originally proposed as a subgenus of Ameiurus.
Istlarius Jordan & Snyder, 1899: 118. Type species: Istlarius balsanus Jordan & Snyder, 1899. Type by original designation. Gender: Masculine.
Ichthyaelurus Meek, 1904: 10. Type species: Silurus cerulescens Rafinesque, 1820. Type by being a replacement
name. Unjustified emendation of Ictalurus Rafinesque, 1820. Gender: Masculine.
Systematics: Smith (1987, fossil forms).
Remarks: The generic name Haustor was originally proposed as a subgenus of Ameiurus, with the type species
Gadus lacustris Walbaum, a species that, at that time, was considered to be an ictalurid catfish [but see Speirs(1952) for the current placement of that name as a species of Lota]. Jordan (1920: 473) clarified his earlier
usage of Gadus lacustris as type species by equating that name with Pimelodus nigricans Le Sueur, 1819,
which is treated here as a junior synonym of Ictalurus punctatus. I interpret Jordan’s action as consistent withthe provisions of the Code that cover the discovery of a misidentified type species (Art. 70.3) and selection of a
new type species that reflects the “taxonomic species actually involved in the misidentification (Art. 70.3.2).”
As such, the name Haustor must be retained as a genus of ictalurid catfish with Pimelodus nigricans Le Sueur,
1819 as its type.
Ictalurus australis (Meek, 1904)
Amiurus australis Meek, 1904: 13. Type locality: Rio Forlón, Forlón, Tamaulipas, México. Holotype: FMNH 4474.
Distribution: Blanco to Panuco Rivers, Vera Cruz, Mexico (Espinosa Pérez, et al., 1993).
Ictalurus balsanus (Jordan & Snyder, 1899)
Istlarius balsanus Jordan & Snyder, 1899: 118, fig. 2. Type locality: Río Ixtla at Puente de Ixtla, Morelos, Mexico.
Holotype: SU 6149.
Istlarius balsanus occidentalis de Buen, 1946b: 119. Type locality: Río Huámito at La Huacana, Río Balsas drainage, Michoacan, Mexico. Syntypes: Whereabouts unknown.
Distribution: Balsas River basin, Mexico (Espinosa Pérez, et al., 1993).
Ictalurus dugesii (Bean, 1880)
Amiurus dugèsii Bean, 1880b: 304. Type locality: Río Turbio, Guanajuato, Mexico. Lectotype: USNM 23123, illustrated in, and designated by (in plate caption), Jordan & Evermann (1900: 3236, pl. 26).
Distribution: Lerma-Santiago, Ameca, Armeria basins and Chapala Lake, Mexico (Espinosa Pérez, et al., 1993).
† Ictalurus echinatus Lundberg, 1975
† Ictalurus echinatus Lundberg, 1975: 25, pl. 7, figs. a, i. Type locality: Verdigre Quarry on the E bank of an
unnamed tributary of Verdigre Creek, in the NW 1/4, SE 1/4, Sec. 16, T. 29 N, R. 7 W, approximately 7.5 mi S
and 5.5 mi W of the town of Verdigre, Knox County, Nebraska. Valentine Formation, late Miocene or early
Pliocene. Holotype: UW 3290.
Distribution: Valentine Formation, Nebraska; late Miocene or early Pliocene (Lundberg, 1975).
Ictalurus furcatus (Valenciennes, 1840)
Pimelodus furcatus Valenciennes, in Cuvier & Valenciennes, 1840b: 136 (102 in Strasbourg deluxe edition). Type
locality: l’Ohio ... le Wabash, ... le Mississipi. Syntypes: MNHN a-8834 (1), MNHN a-9458 (1), MNHN a-8402(1).
Pimelodus affinis Baird & Girard, 1854: 26. Type locality: Rio Grande del Norte. Lectotype: USNM 838, designatedby Gilbert (1998: 206). Specimen listed as a type illustrated in Evermann & Kendall (1894: 126, pl. 13, top),possibly qualifies as an earlier lectotype designation if illustrated specimen is identifiable.
Amiurus meridionalis Günther, 1864: 102. Type locality: Rio Usumacinta, Guatemala. Syntypes: BMNH
1865.4.29.38 (1), BMNH 1865.4.29.41–42 (2); one syntype illustrated in Günther (1868c: pl. 81, fig. 1).

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Amiurus pondersosus Bean, 1880a: 286. Type locality: Mississippi River near St. Louis. Holotype: USNM 287439(skeleton) and USNM 23388 (cast).
Distribution: Mississippi River basin; Gulf Coast basins from Alabama to Mexico (Page & Burr, 1991), and southward to the Usumacinta River basin, Guatemala (Miller et al., 2005).
Remarks: Although the name Pimelodus furcatus has long been treated as valid in Ictalurus, it is actually an
unneeded new name for Pimelodus cauda-furcatus Lesueur, which makes the name a junior synonym of Ictalurus punctatus. The name is retained here as valid, pending action to correct this problem.
† Ictalurus lambda Hubbs & Hibbard, 1951
† Ictalurus lambda Hubbs & Hibbard, 1951: 9. Type locality: Ogallala Formation, Trego County, Kansas. Holotype:
KU 6887 (partial left pectoral-fin spine).
Distribution: Ogallala Formation; Box Butte Co. and Verdigre Quarry, Knox Co., Nebraska; Lapara Creek Locality,
Bee Co. Texas (Lower Pliocene or upper Miocene); McGehee Fauna Locality, Alachua Co. Florida (middle
Pliocene)(Lundberg, 1975).
Ictalurus lupus (Girard, 1858)
Pimelodus lupus Girard, 1858b: 211. Type locality: Headwaters of Rio Pecos, Texas. Syntypes: USNM 915 (1),
USNM 916 (3).
Distribution: Rio Grande River basin (Page & Burr, 1991).
Ictalurus mexicanus (Meek, 1904)
Amiurus mexicanus Meek, 1904: 15. Type locality: Tributary to Rio Valles, Rascon, San Luis Potosi, Rio Panuco
basin, Mexico. Holotype: FMNH 4507.
Distribution: Panuco River basin, Mexico (Miller et al., 2005).
Ictalurus ochoterenai (de Buen, 1946)
Haustor ochoterenai de Buen, 1946a: 271, fig. 1. Type locality: Isla Patos, Mexico. Holotype: at Instituto de Biologia, Mexico.
Distribution: Isla Patos, Mexico (de Buen, 1946a).
Remarks: Treated as valid in Lundberg (1992), but as a synonym of Ictalurus dugesii in Alverez (1966) and Miller etal. (2005).
Ictalurus pricei (Rutter, 1896)
Villarius pricei Rutter, 1896: 257. Type locality: San Bernardino Creek, Rio Yaqui, Arizona, or n. border of Sonora,
Mexico. Syntypes (7): SU 4826 (5), BMNH 1900.9.29.179 (1).
Amiurus meeki Regan, 1907, in Regan, 1906–08: 140, pl. 23 (fig. 5). Type locality: Río Paphigochic, Miñaca, Chihuahua, Mexico. Holotype: BMNH 1905.12.6.348.
Distribution: Yaqui and Casas Grandes River basins, Mexico and southern U. S. A. (Page & Burr, 1991).
Ictalurus punctatus (Rafinesque, 1818)
Silurus punctatus Rafinesque, 1818a: 355. Type locality: Ohio River. No types known.
Pimelodus cauda-furcatus Lesueur, 1819: 152. Type locality: l’Ohio ... Pittsbourg [Gilbert (1998: 210) stated thatthe types were collected from Wabash and Mississippi Rivers]. Syntypes: MNHN a-8402 (1), MNHN a-8834(1).
Pimelodus nigricans Lesueur, 1819: 153, pl. 11 (lowest fig.). Type locality: Lac Erié, l’Ontario, etc. No types
known.
Silurus argentinus Rafinesque, 1820b: 50. Type locality: Lower parts of Ohio River. No types known.
Pimelodus argyrus Rafinesque, 1820c: 358. Type locality: [Ohio River]. No types known. Also in Rafinesque(1820a: 64).
Silurus cerulescens melanurus Rafinesque, 1820b: 49. Type locality: Ohio River. No types known. Originally as Silurus cerulescens var. melanurus.
Silurus maculatus Rafinesque, 1820b: 48. Type locality: Pittsburg and in Kentucky River, Ohio River. No types
known. Also in Rafinesque (1820a: 62 and 1820c: 356) as Pimelodus maculatus. Preoccupied by Silurus maculatus Thunberg 1792.
Silurus maculatus Erythroptera Rafinesque, 1820a: 49. Type locality: Ohio River. No types known. Originally as
Silurus maculatus var. Erythroptera.

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Silurus pallidus Rafinesque, 1820b: 49. Type locality: Ohio River. No types known.
Silurus pallidus Lateralis Rafinesque, 1820b: 49. Type locality: Ohio River. No types known. Originally as Silurus
pallidus var. Lateralis. Described as new as Pimelodus pallidus var. Lateralis in Rafinesque (1820a: 63; and1820c: 358).
Silurus pallidus Leucoptera Rafinesque, 1820b: 49. Type locality: Ohio River. No types known. Originally as Silurus pallidus var. Leucoptera. Described as new as Pimelodus pallidus var. Leucoptera in Rafinesque (1820a:
63; and 1820c: 358).
Silurus pallidus Marginatus Rafinesque, 1820b: 49. Type locality: Ohio River. No types known. Originally as Silurus pallidus var. Marginatus. Described as Pimelodus pallidus var. Marginata in Rafinesque (1820a: 63 and1820c: 358).
Silurus (Pimelodus) borealis Richardson, 1836: 135. Type locality: Pine Island Lake, lat. 54ºN. Holotype: Whereabouts unknown (Gilbert, 1998).
Silurus (Pimelodus) nigrescens Richardson, 1836: 134. Apparently an unintentional new name for Pimelodus nigricans Lesueur.
Pimelodus furcifer Valenciennes, in Cuvier & Valenciennes, 1840b: 139 (103 of Strasbourg deluxe edition). Type
locality: Surinam [in error, actually North America]. Syntype: MNHN 0000-1192 (1).
Pimelodus gracilis Hough, 1852: 26. Type locality: Oswegatchie River and Indian rivers, and other streams tributary
to St. Lawrence River [New York]. Syntype: USNM 1532 (1). Preoccupied by Pimelodus gracilis Valenciennes,
1835; replaced by Pimelodus houghi Girard, 1859.
Pimelodus olivaceus Girard, 1858b: 211, pl. 41 (figs. 1–3). Type locality: Ft. Pierre, and Yellowstone Nebraska;
Milk R., ?Montana; near mouth of Poteau R., and Arkansas R., Arkansas. Syntypes: MCZ 35911; USNM 906(3), USNM 8449 (1), USNM 908 (1), USNM 909 (5), USNM 910 (1), USNM 911 (1, missing). Illustration of
one of the types appeared in Suckley (1860: pl. 42).
Synechoglanis Beadlei Gill, 1859a: 40. Type locality: St. Catherines, between Hamilton and Niagara Falls, Ontario,
Canada. Holotype: USNM 5115.
Pimelodus graciosus Girard, 1859b: 161. Type locality: Prairie mer Rouge, La. [Louisiana]. Syntypes: USNM 1511
(2).
Pimelodus houghi Girard, 1859b: 159. Type locality: Oswegatchie River and Indian rivers, and other streams tributary to St. Lawrence River [New York]. Syntype: USNM 1532 (1). Replacement for Pimelodus gracilis Hough,1852; preoccupied by Pimelodus gracilis Valenciennes, 1835.
Pimelodus megalops Girard, 1859b: 161. Type locality: Black Warrior and Bigsby rivers, Ala. Syntype: USNM
1535 (1).
Pimelodus Hammondii Abbott, 1861: 568. Type locality: Fort Reily. Syntypes: ANSP 22065–66 (2).
Ictalurus simpsonii Gill, 1861b: 43. Type locality: Big Sandy R., Kansas. Syntypes (2): Whereabouts unknown (Gilbert, 1998).
Pimelodus notatus Abbott, 1861: 569. Type locality: Fort Reily, Kansas [probably in error, Gilbert (1998: 222)].
Holotype: ANSP 8449. Preoccupied by Pimelodus notatus Jardine, 1841.
Ichthaelurus robustus Jordan, 1877c: 76, pls. (fig. 3), 2 (fig. 4 & 4b). Type locality: Exact locality unknown [Ohio or
Mississippi rivers]. Holotype: USNM 20056.
Ictalurus okeechobeensis Heilprin, 1887: 129, pl. 18. Type locality: Kissimmee River, tributary to Lake
Okeechobee, Florida. Holotype: ANSP 8443.
Ictalurus anguilla Evermann & Kendall, 1898: 125, pl. 6 (fig. 1). Type locality: Atchafalaya River, above Morgan
City, Pointe Coupee Parish, Louisiana. Holotype: USNM 48788.
† Amiurus decorus Hay, 1924: 1, pl. 1, fig. 1. Type locality: Garvin Gulley locality, 2 1/4 miles north of Navasota,
Texas, Upper Miocene. Holotype: partial pectoral-fin spine; Whereabouts unknown.
Distribution: Great Lakes/Saint Lawrence basin, Hudson Bay, and Mississippi River basin; possibly native to Atlantic and Gulf Coast basins; widely introduced (Page & Burr, 1991).
Remarks: Inclusion of Amiurus decorus in the synonymy is based on Lundberg (1975: 25).
† Ictalurus rhaeas (Cope, 1891)† Rhineastes rhaeas Cope, 1891: 3. Type locality: Cypress Hills Formation, middle lower Oligocene, Cypress Hills,

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FERRARISSaskatchewan, Canada. Holotype: NMC 6209 (vertebral centrum).
† Ameiurus cancellatus Cope, 1891: 4. Type locality: Cypress Hills Formation, middle lower Oligocene, Cypress
Hills, Saskatchewan, Canada. Holotype: at Canadian Geol. Surv. Museum, Ottawa (vertebrae).
† Ameiurus maconnellii Cope, 1891: 5. Type locality: middle lower Oligocene, Cypress Hills, Saskatchewan, Canada. Holotype: at Canadian Geol. Surv. Museum, Ottawa (vertebrae).
Distribution: Cypress Hills Formation, middle lower Oligocene, Cypress Hills, Saskatchewan, Canada (Lundberg,1975).
Remarks: † Rhineastes rhaeas selected as valid name by first reviser action of Lundberg (1975: 17).
† Ictalurus spodius Smith, 1987
† Ictalurus spodius Smith, 1987: 381, fig. 1. Type locality: Gravel pit near Jocotepec, Jalisco, Mexico. Holotype:
UMMP V62528 (nearly complete neurocranium).
Distribution: Jocotepec, Jalisco, Mexico (Smith, 1987).
Species inquirendae, Ictalurus
Silurus Cerulescens Rafinesque, 1820b: 49. Type locality: Ohio River. No types known. Described as Pimelodus cerulescens in Rafinesque (1820a: 63 and 1820c: 357). Possibly a senior synonym of Ictalurus punctatus or Ictalurus furcatus.
Pimelodus vulpes Girard, 1858a: 170. Type locality: Live Oak Creek, Camanche Spring, Rio Leona Piedra Painte,
Devil's R., Rio Salado, Texas. Syntypes: MCZ 35937 (1), USNM 843 (2), USNM 844 (1, missing), USNM 845 (4),USNM 846 (4), USNM 847 (1, missing), USNM 848 (1, missing), USNM 849 (2, missing); syntypes illustrated in
Girard (1859a: 33, pl. 18). Thought by Gilbert (1998: 227) to have been based on specimens of two species (Ictalurus punctatus and Ictalurus lupus). In the absence of a lectotype, the status of this name is unresolved.
NOTURUS Rafinesque, 1818
Noturus Rafinesque, 1818b: 41. Type species: Noturus flavus Rafinesque, 1818. Type by monotypy. Gender: Masculine.
Schilbeodes Bleeker, 1858b: 258. Type species: Silurus gyrinus Mitchill, 1817. Type by monotypy. Gender: Masculine.
Pimelodon Lesueur, in Vaillant, 1896a: 28. Type species: Pimelodon insignarius Vaillant, 1896. Type by monotypy.
Gender: Masculine.
Rabida Jordan & Evermann, 1896: 144. Type species: Noturus furiosus Jordan & Meek, 1889. Type by original designation. Originally proposed as a subgenus of Schilbeodes. Gender: Feminine.
Revision: Taylor (1969).
Phylogeny: LeGrande (1981), Grady (1987), Grady & LeGrande (1992), Hardman (2004).
Remarks: The name Noturus latifrons appears in Jordan (1885) but was not made available therein and must betreated as a nomen nudum; see Gilbert (1998: 217) for discussion.
Noturus albater Taylor, 1969
Noturus albater Taylor, 1969: 144, frontispiece, pls. 4 (fig. 4), 11 (fig. 2). Type locality: White River, at Forsyth, on
Hwy. M80, Taney Co., Missouri. Holotype: UMMZ 151171.
Distribution: White and Saint Francis River systems, Arkansas and Missouri (Taylor, 1969).
Noturus baileyi Taylor, 1969
Noturus baileyi Taylor, 1969: 141, pl.4 (fig. 3), pl. 11 (fig. 1). Type locality: The lower portion of Abrams Creek,
Great Smoky Mountains National Park. Holotype: USNM 201602.
Distribution: Citico Creek, Little Tennessee River basin, Tennessee; Abrams Creek population extirpated (Page &Burr, 1991).
Noturus crypticus Burr, Eisenhour & Grady, 2005
Noturus crypticus Burr, Eisenhour & Grady, 2005: 794, fig. 2c. Type locality: Tennessee, Greene County, Little
Chucky Creek at mouth of Jackson Branch, 36º07'11"N, 83º02'04"W. Holotype: SIUC 52379.
Distribution: French Broad River basin, eastern Tennessee (Burr et al., 2005).

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Noturus elegans Taylor, 1969
Noturus elegans Taylor, 1969: 150, pl. 4 (figs. 5–6), pl. 12. Type locality: Fallen Timber Creek, at Hwy. 90, 8 mi. SE
of Glasgow, Barren Co., Kentucky. Holotype: UMMZ 167597.
Distribution: Green River system, Kentucky and Tennessee River systems, Tennessee (Page & Burr, 1991; Burr etal., 2005).
Noturus eleutherus Jordan, 1877
Noturus eleutherus Jordan, 1877a: 371. Type locality: French Broad River, Tennessee. Holotype: USNM 29678;
holotype illustrated in Jordan (1877c: 73, pl. 40, figs. 62–63a).
Schilbeodes gallowayi Fowler, 1945d: 122, figs. 155–157. Type locality: South Fork of Holston River above Bluff
City, Sullivan Co., Tennessee. Holotype: ANSP 54723.
Distribution: Ohio, Mississippi, Tennessee, and Ouachita and Red River systems (Page & Burr, 1991).
Noturus exilis Nelson, 1876
Noturus exilis Nelson, 1876: 51. Type locality: Mackinaw Cr., McLean Co., Illinois. Lectotype: INHS 37651, designated by Sabaj et al. (1997: 270).
Noturus elassochir Swain & Kalb, 1883: 639. Type locality: Illinois River, Napierville, Illinois. Holotype: USNM
29677.
Distribution: Mississippi River basin, in upland creeks (Taylor, 1969).
Phylgeography: Hardy et al. (2002).
Noturus fasciatus Burr, Eisenhour & Grady, 2005
Noturus fasciatus Burr, Eisenhour & Grady, 2005: 785, fig. 2a. Type locality: Tennessee, Hickman County, Brushy
Fork Creek, off Brushy Road, 1.6 km W of junction with Tennessee Highway 48, N of Aetna, 35º40'25"N,
87º31'03"W. Holotype: SIUC 52378.
Distribution: Duck River basin and nearby tributaries of the Tennessee River, Tennessee (Burr et al., 2005).
Noturus flavater Taylor, 1969
Noturus flavater Taylor, 1969: 204, pl. 4 (fig. 15), pl. 20 (fig. 2). Type locality: Flat Creek, at Hwy. M39, 12 mi. NE.
of Cassville, Barry Co., Missouri. Holotype: UMMZ 151322.
Distribution: Upper White River system, Arkansas and Missouri (Page & Burr, 1991).
Noturus flavipinnis Taylor, 1969
Noturus flavipinnis Taylor, 1969: 201, pl. 4 (fig. 14), pl. 20 (fig. 1). Type locality: Hines Creek, Clinton, Tennessee.
Holotype: USNM 163801.
Distribution: Upper Tennessee River basin (Taylor, 1969).
Noturus flavus Rafinesque, 1818
Noturus flavus Rafinesque, 1818b: 41. Type locality: Ohio River [now: Eagle Creek (a tributary to the lower Kentucky River which is a tributary to the Ohio River), from a pool below bridge at state highway 36, 3.5 miles eastof Jonesville in Grant Co., Kentucky, by neotype designation]. Neotype: USNM 202494, designated by Taylor(1969: 118).
Noturus luteus Rafinesque, 1819: 421. Type locality: [No locality stated]. No types known.
Noturus occidentalis Gill, 1861b: 45. Type locality: Platte River. Holotype: Whereabouts unknown (Gilbert, 1998).
Noturus platycephalus Günther, 1864: 104. Type locality: North America. Syntypes (2): BMNH 2005.9.21.1.
Distribution: Mississippi River system, Mohawk and Hudson Rivers, Great Lakes-Saint Lawrence River basin (Taylor, 1969).
Noturus funebris Gilbert & Swain, 1891
Noturus funebris Gilbert & Swain, in Gilbert, 1891: 153. Type locality: in a Spring Run tributary to North River near
Tuscaloosa, Alabama. Lectotype: USNM 36696, designated by Taylor (1969: 98).
Distribution: Gulf Coast basins east of Mississippi River (Taylor, 1969).
Noturus furiosus Jordan & Meek, 1889
Noturus furiosus Jordan & Meek, in Jordan, 1889: 351, pl. 43 (figs. 1–1b). Type locality: Neuse River at Millburnie,
near Raleigh, North Carolina. Lectotype: USNM 39932, illustrated in, as designated by (in plate caption), Jordan & Evermann (1900: 3237, pl. 29, figs. 69, 69a, 69b); same specimen designated as lectotype by Taylor(1969: 186).

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Noturus gilberti Jordan & Evermann, 1889
Noturus gilberti Jordan & Evermann, in Jordan, 1889: 352, pl. 43 (figs. 2–2b). Type locality: Roanoke River, Salem,
Virginia. Lectotype: USNM 3993, designated by (in caption to plate) Jordan & Evermann (1900: 3236, pl. 28);
same specimen designated as lectotype by Taylor (1969).
Distribution: Roanoke River basin, Virginia; possibly introduced elsewhere (Page & Burr, 1991).
Noturus gladiator Thomas & Burr, 2004
Noturus gladiator Thomas & Burr, 2004: 353, figs. 2–3. Type locality: Tennessee: Hardeman County: Mississippi
River basin, Pleasant Run (Hatchie River) at Summerville-Bolivar Rd., just upstream of Potters Creek, ca. 2.5
km W of Bolivar; 35º15'14"N, 089º02'05"W. Holotype: SIUC 47997.
Distribution: Mississippi River tributaries of western Tennessee and Mississippi (Thomas & Burr, 2004).
Noturus gyrinus (Mitchill, 1817)
Silurus gyrinus Mitchill, 1817: 289. Type locality: Wallkill [New York]. No types known.
Noturus sialis Jordan, 1877c: 102, pl. 42 (fig. 68), pl. 43 (fig. 69). Type locality: White River, Indianapolis, Indiana.
Holotype: Whereabouts unknown.
Distribution: Atlantic draining Rivers, below Fall line; Mississippi River basin; rivers draining Gulf of Mexico(north of Rio Grande); and Great Lakes; introduced into many additional rivers (Taylor, 1969).
Noturus hildebrandi (Bailey & Taylor, 1950)
Schilbeodes hildebrandi Bailey & Taylor, 1950: 32, pl. 1; pl. 2 (figs. e–f). Type locality: Brushy Creek, about 1 mile
above its mouth in Homochitto River, T.4N, R2E., Sect. 14, at the tiny hamlet of Homochitto, Amite County,
Mississippi. Holotype: UMMZ 157620.
Noturus hildebrandi lautus Taylor, 1969: 138, pl. 4 (fig. 2), pl. 10 (fig. 2). Type locality: North Fork of Obion River,at Tennessee Highway 69, Henry County, Tennessee. Holotype: USNM 201665.
Distribution: Left bank tributaries of Mississippi River, in Mississippi, Kentucky, and Tennessee (Taylor, 1969).
Noturus insignis (Richardson, 1836)
Pimelodus insigne Richardson, 1836: 132. Type locality: Philadelphia. Holotype: MNHN 0000-3053. Available
from reference to “Pimelodus ... insigne (livrée, Le Sueur).”
Pimelodus lemniscatus Valenciennes, in Cuvier & Valenciennes, 1840b: 144 (107 in Strasbourg deluxe edition).
Type locality: Philadelphia. Holotype: MNHN 0000-3053. On Pimélode livrée of Lesueur (1819).
Noturus marginatus Baird, in Cope, 1868: 237. Type locality: Sinking Creek of Kanawha River; head of JamesRiver, Virginia; and Susquehanna River at Carlisle, Pennsylvania. Syntypes (more than 20): ANSP 8431–32 (2),possibly ANSP 8435 (1), MCZ 8371 (1), MCZ 35955 (1), MNHN 0000-0340 (1), MNHN 0000-0341 (1),
UMMZ 138221 (1), USNM 1571 (9).
Pimelodon insignarius Vaillant, 1896a: 28, pl. 24. Type locality: Philadelphia. Holotype: MNHN 0000-3053.
Schilbeodes marginatus atrorus Hubbs & Raney, 1944: 7, 10, pl. 1. Type locality: Wolf Creek, between Bastian andNovis, Bland Co., Virginia. Holotype: UMMZ 139452.
Distribution: Atlantic coastal rivers between New York and Georgia, generally at or above Fall Line and New and
Kanawha River basins; introduced elsewhere (Talyor, 1969),
Remarks: The name Pimelode livrée in Lesueur (1819: 155) was considered by Taylor (1969: 83) to be an informalname and therefore not available. The name was accompanied by a brief description which, when assigned aname by Richardson (1836: 132), was sufficient to make Richardson’s name available, with the specimen examined by Lesueur as the holotype.
Noturus lachneri Taylor, 1969
Noturus lachneri Taylor, 1969: 54, pl. 3 (fig. 2), pl. 5 (fig. 2). Type locality: Middle Fork of the Saline River at
Arkansas Highway 7 crossing, 11.2 miles north of Mountain Valley, Garland Co., Arkansas. Holotype: USNM
201592.
Distribution: Saline and Ouachita River systems, Arkansas (Page & Burr, 1991).
Noturus leptacanthus Jordan, 1877
Noturus leptacanthus Jordan, 1877a: 352. Type locality: Silver Creek, 1 mi. above its junction with the Etowah R.,
near Rome, Georgia. Holotype: apparently destroyed (Taylor, 1969).

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Distribution: Southeastern United States, from Atlantic and Gulf of Mexico draining rivers (Taylor, 1969).
Noturus miurus Jordan, 1877
Noturus miurus Jordan, 1877a: 371. Type locality: Maumee River basin, lower Wabash River basin and White Rivernear Indianapolis, Indiana [restricted to White River near Indianapolis, by lectotype designation]. Lectotype:
MNHN a-1308, designated by Taylor (1969: 197).
Distribution: Great Lakes basin, Mississippi River basin, Pearl River system, and smaller rivers between the Pearland Mississippi (Taylor, 1969).
Noturus munitus Suttkus & Taylor, 1965
Noturus munitus Suttkus & Taylor, 1965: 171, figs. 1, 2. Type locality: Pearl River, 2.6 mi. east of Sandy Hook,
Marion Co., Mississippi. Holotype: TU 26250.
Distribution: Gulf Coast basins from Georgia to Louisiana, in several disjunct populations (Page & Burr, 1991).
Noturus nocturnus Jordan & Gilbert, 1886
Noturus nocturnus Jordan & Gilbert, 1886: 6. Type locality: Poteau River, ... Washita and Saline Rivers, [Arkansas
River tributaries]; [restricted to Saline River near Benton railroad bridge by lectotype designation]. Lectotype:
USNM 36461, designated by use of term “type” in figure caption in Evermann & Kendall (1894: pl. 11, top),
and approximate size of illustrated specimen which were used by Taylor (1969: 79) to identify the individual.
Distribution: Lower and central Mississippi River basin and other rivers draining into Gulf of Mexico (Taylor,1969).
Noturus phaeus Taylor, 1969
Noturus phaeus Taylor, 1969: 100, pl. 3 (fig. 8), pl. 7 (fig. 2). Type locality: North Fork of the Obion River at state
highway 69, Henry County, Tennessee. Holotype: USNM 202143.
Distribution: Lower Mississippi River valley and headwaters of Bayou Teche, Louisiana (Taylor, 1969).
Noturus placidus Taylor, 1969
Noturus placidus Taylor, 1969: 168, pls. 4 (fig. 9), 15 (fig. 2), 17 (fig. 2). Type locality: Neosho River, just S. of U.S.
Hwy. 50, near Emporia, sec. 7, T. 19 S., R. 12 E., Lyon Co., Kansas. Holotype: UMMZ 167653.
Distribution: Neosho River and tributaries, Cottonwood River and Illinois River, Mississippi River basin (Taylor,1969).
Noturus stanauli Etnier & Jenkins, 1980
Noturus stanauli Etnier & Jenkins, 1980: 19, figs. 1–2. Type locality: Clinch R. at Frost Ford, Hancock Co., Tennessee. Holotype: TU 110929.
Distribution: Clinch and Duck Rivers, Tennessee River basin, Tennessee (Page & Burr, 1991).
Noturus stigmosus Taylor, 1969
Noturus stigmosus Taylor, 1969: 173, pls. 2 (fig. 4), 4 (fig. 11), 16 (fig. 1), 18 (fig. 1). Type locality: Huron River,just south of pool, below North Territorial Road, north of Dexter, sec. 13, T. 1 S., R. 4 E., Washtenaw Co., Michigan. Holotype: UMMZ 165843.
Distribution: Lake Erie and Ohio River basin (Thomas & Burr, 2004).
Noturus taylori Douglas, 1972
Noturus taylori Douglas, 1972: 785, fig. 1. Type locality: South Fork of Caddo R., 1.6 km southeast of Hopper and0.8 km south of state hwy. 240, Montgomery Co., Arkansas. Holotype: USNM 206181.
Distribution: Caddo, Ouachita and Little Missouri River basins, Arkansas (Page & Burr, 1991).
Noturus trautmani Taylor, 1969
Noturus trautmani Taylor, 1969: 156, pls. 4 (fig. 7), 13 (fig. 1). Type locality: Big Darby Creek, 1 mile south of Fox,
southeastern Jackson Township, Pickaway County, Ohio. Holotype: UMMZ 187098.
Distribution: Big Darby Creek, Scioto River basin, Ohio (Taylor, 1969).
Species inquirenda, Noturus
Silurus mollis Hermann, 1804: 309. Type locality: America. No types known. Treated by Hubbs & Raney (1944) and,
subsequently, Gilbert (1998: 220) as a senior synonym of Silurus gyrinus Mitchell, but Taylor (1969: 52) rejectedthat conclusion and, instead, questioned whether the name could be unambiguously associated with any species. In

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the absence of a primary type specimen, the name is listed here as species inquirenda.
PRIETELLA Carranza, 1954
Prietella Carranza, 1954: 130. Type species: Prietella phreatophila Carranza, 1954. Type by original designation.
Gender: Feminine.
Prietella lundbergi Walsh & Gilbert, 1995
Prietella lundbergi Walsh & Gilbert, 1995: 851, figs. 1 (top), 2 c–d. Type locality: a subsurface thermal spring(Nacimiento de San Rafael de Los Castros) approximately 15 km WNW of Ciudad Mantae Tamaulipas state,
México (Río Tamesí drainage). Holotype: UNAM P-7705.
Distribution: Two springs of Tamesí River drainage, Tamaulipas state, Mexico (Miller et al., 2005).
Prietella phreatophila Carranza, 1954
Prietella phreatophila Carranza, 1954: 132, pl. 1. Type locality: Well at base of Sierra de Santa Rosa, ca. 27º55'N,
101º07'W, Múzquiz, Coahuila state, Mexico. Holotype: Author's personal coll.; illustrated in Miller et al. (2005:
165, fig. 6.171).
Distribution: Cases in Bravo River basin, Coahuila, from near Múzquiz northward to about 50 km SW of Ciudad
Acuña and to near Allende, Mexico (Miller et al., 2005).
PYLODICTIS Rafinesque, 1819
Pylodictis Rafinesque, 1819: 422. Type species: Pylodictis limosus Rafinesque, 1819. Type by monotypy. Gender:
Masculine. Placed on the Official List (ICZN, Opinion 1584). Proposed as new, but spelled Pilodictis in
Rafinesque (1820c: 361, 1820a: 67).
Ilictis Rafinesque, 1820c: 360. Type species: Silurus limosus Rafinesque, 1820 [=Pylodictis limosus Rafinesque,1819]. Type by monotypy. Gender: Masculine. Originally proposed as a section of the subgenus Ictalurus. Alsoin Rafinesque (1820: 66).
Leptops Rafinesque, 1820c: 358. Type species: Silurus viscosus Rafinesque, 1820. Type by subsequent designation
by Jordan & Gilbert (1877: 87). Gender: Masculine. Originally proposed as a section of the subgenus Ictalurus.
Also proposed as new in Rafinesque (1820a: 64).
Opladelus Rafinesque, 1820c: 359. Type species: Silurus nebulosus Rafinesque, 1820. Type by monotypy. Gender:
Masculine. Originally proposed as a subgenus of Ictalurus. Also in Rafinesque (1820a: 64–65).
Hopladelus Gill 1862: 51. Type species: Silurus nebulosus Rafinesque, 1820. Type by monotypy. Gender: Masculine. Unjustified emendation or misspelling of Opladelus Rafinesque, 1820.
Pelodichthys Jordan, 1877b: 10. Type species: Pylodictis limosus Rafinesque, 1819. Type by monotypy. Gender:
Masculine. Unjustified emendation of Pylodictis Rafinesque, 1819.
Remarks: The name Pylodictis is treated as valid, with the type species Pylodictis limosus (with Silurus olivaris as asenior subjective synonym) following Opinion 1584 of the Code. However, see Markle (1997: 446) in which it
shown that Pylodictis limosus was a mythical fish, based on a drawing made byAudubon.
Pylodictis olivaris (Rafinesque, 1818)
Silurus olivaris Rafinesque, 1818a: 355. Type locality: Ohio River. No types known. Placed on the Official List
(ICZN, Opinion 1584).
Pylodictis limosus Rafinesque, 1819: 422. Type locality: Ohio River mouth. No types known. Described as new as
Pimelodus limosus in Rafinesque (1820a: 67; and 1820c: 361); and as Silurus limosus in Rafinesque (1820b:
51).
Pimelodus aeneus Lesueur, 1819: 150. Type locality: l’Ohio. Type(s): Whereabouts unknown (Gilbert, 1998).
Silurus cupreus Rafinesque, 1820b: 51. Type locality: Ohio River. No types known. Described as Pimelodus cupreus
in Rafinesque (1820a: 65 and 1820c: 359).
Silurus nebulosus Rafinesque, 1820b: 50. Type locality: Ohio River. No types known. Described as Pimelodus nebulosus in Rafinesque (1820a: 64 and 1820c: 358).
Silurus viscosus Rafinesque, 1820b: 50. Type locality: Near Louisville, Kentucky, at falls, Ohio River. No types
known. Described as Pimelodus viscosus in Rafinesque (1820a: 64; and 1820c: 358).
Pimelodus lutescens Rafinesque, 1832: 20. Type locality: Tennessee River. No types known.

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Pimelodus punctulatus Valenciennes, in Cuvier & Valenciennes, 1840b: 134 (100 of Strasbourg deluxe edition).
Type locality: Newharmony. Lectotype: MNHN a-9459. Lectotype designated by Eschmeyer (in Eschmeyer etal. 1998: 1406). Although the description of the species was based entirely on one specimen, a second specimen, MNHN a-8965, from New Orleans, was listed, making both specimens syntypes. Bertin & Esteve (1950:24) listed MNHN a-9459 as the holotype and Gilbert (1998: 225) incorrectly treated that statement as a defacto
lectotype designation.
Distribution: Great Lakes and Mississippi River basins, Gulf Coast drainages from Georgia to Mexico; introduced
elsewhere (Page & Burr, 1991); reported from the Pleistocene of Florida by Young & Laerm (1993).
SATAN Hubbs & Bailey, 1947
Satan Hubbs & Bailey, 1947: 4. Type species: Satan eurystomus Hubbs & Bailey, 1947. Type by original designation. Gender: Masculine.
Satan eurystomus Hubbs & Bailey, 1947
Satan eurystomus Hubbs & Bailey, 1947: 8, pl. 1 (figs. 1–3). Type locality: An artesian well (depth 381 m), near San
Antonio, ca. 29º30'N, 98º30'W, Bexar Co., Texas. Holotype: UMMZ 190922.
Distribution: Artesian wells near San Antonio, Texas (Page & Burr, 1991).
TROGLOGLANIS Eigenmann, 1919
Trogloglanis Eigenmann, 1919b: 397. Type species: Trogloglanis pattersoni Eigenmann, 1919. Type by monotypy.
Gender: Masculine.
Trogloglanis pattersoni Eigenmann, 1919
Trogloglanis pattersoni Eigenmann, 1919b: 397, figs. 1–2. Type locality: Artesian well in San Antonio, Edwards
Aquifer, ca. 29º30'N, 98º30'W, Bexar Co., Texas. Holotype: CAS 58675.
Distribution: Artesian wells near San Antonio, Texas (Page & Burr, 1991).
Remarks: Anatomical description in Lundberg (1982).

LACANTUNIIDAE Rodiles-Hernández, Hendrickson & Lundberg, 2005
Lacantuniidae Rodiles-Hernández, Hendrickson & Lundberg, in Rodiles-Hernández et al., 2005: 6. Type genus: Lacantunia Rodiles-Hernández, Hendrickson & Lundberg, 2005.
1 Genus, 1 species; no fossil species.
LACANTUNIA Rodiles-Hernández, Hendrickson & Lundberg, 2005
Lacantunia Rodiles-Hernández, Hendrickson & Lundberg, in Rodiles-Hernández et al., 2005: 6. Type species:
Lacantunia enigmatica Rodiles-Hernández, Hendrickson & Lundberg. Type by original designation. Gender:
Feminine.
Remarks: Treated as the type of the family Lacantuniidae, in Rodiles-Hernandez et al. (2005: 5), or as incertae sedis
within the Siluriformes (2005: 6).
Lacantunia enigmatica Rodiles-Hernández, Hendrickson & Lundberg, 2005
Lacantunia enigmatica Rodiles-Hernández, Hendrickson & Lundberg, in Rodiles-Hernández et al., 2005: 10, fig. 1.
Type locality: México, Chiapas, Río Usumacinta basin, Río Lacantún, Selva Lacandona, Reserva de la Biosfera
Montes Azules, 16º08.083'N, 90º55.317'W. Holotype: ECO-SC 3859.
Distribution: Usumacinta River basin, Chiapas, Mexico (Rodiles-Hernández et al., 2005).

LORICARIIDAE Rafinesque, 1815
Loricaria Rafinesque, 1815: 89. Type genus: Loricaria Linnaeus, 1758.
Hypostomiden Kner, 1853b: 279. Type genus: Hypostomus La Cepède, 1803.
Ancistri Kner, 1854: 6. Type genus: Ancistrus Kner, 1854.

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Acanthicini Bleeker, 1862 (in Bleeker, 1862–63): 2. Type genus: Acanthicus Spix & Agassiz, 1829.
Acestrini Bleeker, 1862 (in Bleeker, 1862–63): 4. Type genus: Acestra Kner, 1853. Permanently invalid, following Art.
39 of the Code, because the underlying generic name is preoccupied.
Hypoptopominae [sic, Hypoptopomatinae] Eigenmann & Eigenmann, 1890: 8, 12, 353, 388. Type genus: Hypoptopoma
Günther, 1868.
Neoplecostominae Regan, 1904b: 306. Type genus: Neoplecostomus Eigenmann & Eigenmann, 1888.
Lithogeneinae [sic, for Lithogeninae] Gosline, 1947: 80. Type genus: Lithogenes Eigenmann, 1909.
Chaetostomidi Fowler, 1958: 14. Type genus: Chaetostomus Kner, 1846.
Farlowellidi Fowler, 1958: 14. Type genus: Farlowella Eigenmann & Eigenmann, 1889.
Harttiinae Boeseman, 1971: 4, 10. Type genus: Harttia Steindachner, 1877.
Acestridiinae Isbrücker & Nijssen, 1974a: 67, 68. Type genus: Acestridium Haseman, 1911.
Loricariichthyina Isbrücker, 1979a: 87, 89. Type genus: Loricariichthys Bleeker, 1862.
Metaloricariina Isbrücker, 1979a: 88, 90. Type genus: Metaloricaria, Isbrücker, 1975.
Hemiodontichthyina Isbrücker, 1979a: 88, 89. Type genus: Hemiodontichthys Bleeker, 1862.
Otocinclini Isbrücker, 1979b: 114. Type genus: Otocinclus Cope, 1871.
Planiloricariina Isbrücker, 1979a: 87, 89. Type genus: Planiloricaria Isbrücker, 1971.
Reganellina Isbrücker, 1979a: 87, 89. Type genus: Reganella Eigenmann, 1905.
Rineloricariina Isbrücker, 1979a: 87, 89. Type genus: Rineloricaria Bleeker, 1862.
Lithoxina Isbrücker, 1980: 77. Type genus: Lithoxus Eigenmann, 1910.
Pseudacanthicini Isbrücker, 1980: 76. Type genus: Pseudacanthicus Bleeker, 1862.
Ricolina Isbrücker, 1981: 54. Type genus: Ricola Isbrücker & Nijssen, 1978.
Hopliancistrini Isbrücker & Nijssen, 1989: 543. Type genus: Hopliancistrus Isbrücker & Nijssen, 1989.
Spectracanthicina Isbrücker & Nijssen, 1989: 544. Type genus: Spectracanthicus Nijssen & Isbrücker, 1987.
Otothyrini Schaefer, 1991: 31. Type genus: Otothyris Myers, 1927.
Lampiellini Isbrücker, in Isbrücker et al., 2001: 20. Type genus Lampiella Isbrücker, 2001.
Corymbophanini Armbruster, 2004a: 59. Type genus: Corymbophanes Eigenmann, 1909.
Pterygoplichthini [sic, for Pterygoplichthyini] Armbruster, 2004a: 61. Type genus: Pterygoplichthys Gill, 1858.
Rhinelepini Armbruster, 2004a: 61. Type genus: Rhinelepis Spix & Agassiz, 1829 [not Valenciennes, as stated in Armbruster].
Taxonomic summary: Gosline (1947); Isbrücker (1979b, 1980, 1992a, 2001); Reis et al. (2003); Ferraris (2003b, Neoplecostominae; 2003c, Loricariinae); Fisch-Muller (2003, Ancistrinae); Schaefer (2003c, Hypoptopomatinae); Weber(2003, Hypostominae).
Phylogeny: Boeseman (1971); Schaefer (1987, 1998); Montoya-Burgos et al. (1997, 1998, 2002, molecular phylogeny);
Armbruster (1998a, Rhinelepis group); Schaefer (1991, Hypoptopomatinae); Armbruster (2004a, Ancistrinae andHypostominae); Gauger & Buckup (2005, Otothyrini).
Review: Aquino (1997, Hypoptopomatinae of Argentina), Weber et al. (1992, Paraguay).
Remarks: The taxonomy of this some portions of this family is very unsettled at this time. Monophyly of the family
appears to be strongly supported, except, possibly, for the inclusion of Lithogenes (see Schaefer, 2003d, and Armbruster, 2004a for comments). Two subfamilies, Loricariinae and Hypoptopomatinae, as used in Reis et al. (2003),
appear to be generally regarded as monophyletic, but the monophyly and composition of the other subfamilies inReis et al. (2003) are currently being examined and likely to be altered substantially in the future (see, for example,
Armbruster, 2004a). Isbrücker & Nijssen (1992b) summarized the extent of sexual dimorphism in the family; Muller& Weber (1992) examined the possible taxonomic value of dental characteristics.
96 genera, 716 species; no named fossil taxa.
Incertae sedis
Hypostomus itacua Valenciennes, 1836
Hypostomus itacua Valenciennes, 1836, in Valenciennes, 1835–47: pl. 7 (fig. 1). Type locality: not stated [considered to be restricted to tributaries of La Plata]. Holotype not found in MNHN. Name available from caption to

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plate, with illustrated specimen the holotype.
Distribution: Middle Paraná River basin.
Remarks: Type locality from description of species in Cuvier & Valenciennes (1840: 506), but Weber (2003) indicated that redescription in Cuvier & Valenciennes (1840: 505) does not agree with the illustration in Valenciennes (1836). Redescription is apparently the basis of decisions by Günther (1864: 243) and Isbrücker (1980:
50) to place the species in the ancistrin genus Hemiancistrus.
Plecostomus obtusirostris (Steindachner, 1907)
Plecostomus obtusirostris Steindachner, 1907e: 490. Type locality: Rio Cubatao, Santa Catherina by Theresopolis,
Brazil. Holotype: at NMW (not found; Weber, 2003).
Distribution: Coastal drainages of southeastern Brazil.
Remarks: Weber (2003) disagreed with the placement of this species in Hypostomus by Isbrücker (1980: 29) inasmuch he indicated that the description included mention of hypertrophied odontodes on the sides of the head ofthe examined specimen.
ACANTHICUS Spix & Agassiz, 1829
Acanthicus Spix & Agassiz, 1829: 2. Type species: Acanthicus hystrix Spix & Agassiz, 1829. Type by monotypy.
Gender: Masculine.
Review: Isbrücker & Nijssen (1988).
Acanthicus adonis Isbrücker & Njissen, 1988
Acanthicus adonis Isbrücker & Nijssen, 1988: 166, fig. 5. Type locality: Brasilien, Est. Pará, Rio Tocantins bei Cametá (0º14'S, 49º30.5'W). Holotype: MZUSP 38580.
Distribution: Lower Tocantins River basin, Brazil (Fisch-Muller, 2003).
Acanthicus hystrix Spix & Agassiz, 1829
Acanthicus hystrix Spix & Agassiz, 1829: 3, pl. 1 (fig. 1). Type locality: flumine Amazonum. Holotype: Whereabouts unknown (Kottelat, 1988) and possibly destroyed (Terofal, 1983).
Rinelepis acanthicus Valenciennes, in Cuvier & Valenciennes, 1840b: 487 (360 in Strasbourg deluxe edition). Type
locality: flumine Amazonum. Holotype: destroyed (Terofal, 1983). Replacement name for Acanthicus hystrix
Spix & Agassiz, 1829, then considered congeneric with, and preoccupied by, Rinelepis histrix Valenciennes(1840), which was based on the older, but unpublished, Vandelli manuscript name Loricaria histrix.
Distribution: Amazon River basin (Fisch-Muller, 2003).
ACESTRIDIUM Haseman, 1911
Acestridium Haseman, 1911a: 319. Type species: Acestridium discus Haseman, 1911. Type by monotypy. Gender:
Neuter.
Revision: Retzer et al. (1999).
Acestridium colombiense Retzer, 2005
Acestridium colombiensis Retzer, 2005: 2, fig. 1. Type locality: Colombia: Guainia State: Orinoco Basin: Río Inirida, lagoon at 1 km up river from Puerto Inirida. Holotype: FMNH 115255.
Distribution: Orinoco River basin, Colombia (Retzer, 2005).
Acestridium dichromum Retzer, Nico & Provenzano, 1999
Acestridium dichromum Retzer, Nico & Provenzano, 1999: 316, fig. 3(a–c). Type locality: Pozo Lucas, tributary of
Río Orinoco, 7 km south of San Fernando de Atabapo, 4º02'N, 67º42'W, Amazonas State, Venezuela. Holotype:
MBUCV V-26780.
Distribution: Upper Orinoco and Negro River basins, Venezuela (Schaefer, 2003c).
Acestridium discus Haseman, 1911
Acestridium discus Haseman, 1911a: 319, pls. 50 (fig. 1), 51. Type locality: Igarapé de Cachoeira Grande, near
Manaos, Brazil. Holotype: FMNH 54339.
Distribution: Lower Negro River basin, Brazil (Schaefer, 2003c).
Acestridium martini Retzer, Nico & Provenzano, 1999
Acestridium martini Retzer, Nico & Provenzano, 1999: 318, fig. 3 (d–f). Type locality: Caño Pozo Azul, tributary of

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Río Orinoco, near Balneario Pozo Azul, 5º45'49"N, 67º29'21"W, Amazonas State, Venezuela, elev. 75 m. Holotype: MCNG 36484.
Distribution: Upper Orinoco and Negro River basins, Venezuela (Schaefer, 2003c).
ANCISTRUS Kner, 1854
Ancistrus Kner, 1854: 272. Type species: Hypostomus cirrhosus Valenciennes, 1836. Type by subsequent designation by Bleeker (1862–63: 2). Gender: Masculine.
Xenocara Regan, 1904b: 251. Type species: Chaetostomus latifrons Günther, 1869. Type by subsequent designation
by Eigenmann (1910: 405, 410). Gender: Neuter.
Thysanocara Regan, 1906a: 96. Type species: Hypostomus cirrhosus Valenciennes, 1836. Type by monotypy. Gender: Neuter. Proposed as a subgenus of Xenocara.
Pristiancistrus Fowler, 1945a: 121. Type species: Pristiancistrus eustictus Fowler, 1945. Type by original designation. Gender: Masculine.
Monophyly and systematics: Schaefer (1986); Fisch-Muller (1999).
Key: Fisch-Muller et al. (2005, species lacking adipose fin).
Ancistrus aguaboensis Fisch-Muller, Mazzoni & Weber, 2001
Ancistrus aguaboensis Fisch-Muller, Mazzoni & Weber, 2001: 293, fig. 4 (middle). Type locality: Brazil: Goiás:
Minaçú: córrego Água Boa: small tributary of rio Bonito, a left bank tributary of the upper rio Tocantins, 10 km
from Minaçú on road Minaçú-Campinaçu (13º35'S 48º14'W). Holotype: MNRJ 20850.
Distribution: Upper Tocantins River basin, Brazil (Fisch-Muller, 2003).
Ancistrus bodenhameri Schultz, 1944
Ancistrus brevifilis bodenhameri Schultz, 1944c: 305, pl. 10 (fig. A). Type locality: Río San Pedro at the bridge
south of Mene Grande, Motatán system; Venezuela. Holotype: USNM 121066.
Distribution: Eastern and western Maracaibo lake tributaries, Venezuela (Fisch-Muller, 2003).
Ancistrus bolivianus (Steindachner, 1915)
Xenocara boliviana Steindachner, 1915e: 95, pl. 9 (figs. 5–6). Type locality: Rio Songo [= Zongo], Nord Yungas,
Bolivia. Syntypes: NMW 43475 (2), NMW 43476 (27). Species described conditionally “Xenocara bufonia(C.V.) Regan, juv.? (an Xenocara boliviana n. sp.).”Distribution: Beni, Mamoré and Madre de Dios River basins (Fisch-Muller, 2003).
Ancistrus brevifilis Eigenmann, 1920
Ancistrus brevifilis Eigenmann, 1920a: 7, pl. 1. Type locality: El Concejo, Rio Tiquirito, Venezuela. Holotype: CAS
64609.
Distribution: Tuy River basin, Venezuela (Fisch-Muller, 2003).
Ancistrus brevipinnis (Regan, 1904)
Xenocara brevipinnis Regan, 1904b: 257. Type locality: Rio Grande do Sul [Sistema da Laguna dos Patos, Brazil].
Holotype: BMNH 1891.3.16.76, illustrated in Muller (1989: 894, fig. 3, tab. 3).
Distribution: Laguna dos Patos basin, Brazil (Fisch-Muller, 2003).
Ancistrus bufonius (Valenciennes, 1840)
Hypostomus bufonius Valenciennes, in Cuvier & Valenciennes, 1840b: 511 (377 in Strasbourg deluxe edition). Type
locality: Rio Apurimac, qui descend des montagnes du haut Pérou, à environ 2000 mètres de hauteur. Syntypes:
MNHN 0000-2228 (2).
Hypostomus calamita Valenciennes, in Cuvier & Valenciennes, 1840b: 515 (380 in Strasbourg deluxe edition). Type
locality: dans les montagnes du haut Pérou, dans le rio Apurimac. Syntypes: MNHN 0000-2227 (2).
Distribution: Apurimac River basin, Peru (Fisch-Muller, 2003).
Ancistrus caucanus Fowler, 1943
Ancistrus caucanus Fowler, 1943a: 253, figs. 41–43. Type locality: Sonsón, Cauca River basin, Colombia. Holotype: ANSP 70516.
Distribution: Cauca River basin, Colombia; known only from the holotype (Fisch-Muller, 2003).
Ancistrus centrolepis Regan, 1913
Ancistrus centrolepis Regan, 1913d: 470. Type locality: River Choco, San Juan, Colombia. Syntypes: BMNHCHECKLIST OF CATFISHES, RECENT AND FOSSIL

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1910.7.11.122 (1), BMNH 1913.10.1.58 (1).
Ancistrus melas Eigenmann, 1916: 83. Type locality: Condoto [San Juan basin, Colombia]. Holotype: FMNH
58339, illustrated in Eigenmann (1922b: 87, fig. 20; pl. 12, fig. 2).
Ancistrus baudensis Fowler, 1945a: 122, fig. 33–36. Type locality: Alto Rio Baudó, Pacific slope at 3000 feet elevation, Colombia. Holotype: ANSP 71709.
Distribution: Coastal drainages of northern Andes, rivers of northern Colombia (Fisch-Muller, 2003).
Ancistrus chagresi Eigenmann & Eigenmann, 1889
Ancistrus chagresi Eigenmann & Eigenmann, 1889b: 47. Type locality: Rio Chagres [Panama]. Syntypes (2, 14–19cm): MCZ 8026 (2). Name made available by diagnostic features in key on p. 47.
Distribution: Panama Canal Zone, Chagres, Gatun and Chorrera River basins, Panama (Fisch-Muller, 2003).
Ancistrus cirrhosus (Valenciennes, 1836)
Hypostomus cirrhosus Valenciennes, 1836, in Valenciennes, 1835–47: pl. 7 (fig. 3). Type locality: Not stated. Holotype: at MNHN; not found (Fisch-Muller, 2003). Name available from caption to plate, with the illustrated specimen the holotype.
Distribution: Paraná River basin, Argentina (Fisch-Muller, 2003).
Remarks: Species subsequently described in Cuvier & Valenciennes (1840b: 511, 378 in Strasbourg deluxe edition),with the statement that the species was found in Missiones and Buénos-Ayres. However, Fisch-Muller (2003)
presented evidence that the collection was likely to have come from Corrientes. Isbrücker’s (1980: 68) designation of MNHN b-0603 as lectotype is invalid, since the illustrated specimen must be considered the holotype.
Fisch-Muller (2003) suggested, instead, that the holotype was not preserved.
Ancistrus claro Knaack, 1999
Ancistrus claro Knaack, 1999a: 71, fig. 1. Type locality: Brazil, Mato Grosso, Rio Claro, above the bridge, in the
direction of Chappada dos Guimaraes. Holotype: MCP 28667.
Distribution: Cuiabá River basin in upper Paraguay River drainage, Brazil (Fisch-Muller, 2003).
Ancistrus clementinae Rendahl, 1937
Ancistrus clementinae Rendahl, 1937: 4, fig. 2 (upper). Type locality: Ecuador, Syst. Rio de Clementina [=Río Pozuelos], nw. von Babahoyo. Holotype: NRM 10369.
Distribution: Pozuelos River basin in the Guayas River drainage, Ecuador (Fisch-Muller, 2003).
Ancistrus cryptophthalmus Reis, 1987
Ancistrus cryptophthalmus Reis, 1987: 82, fig. 1b. Type locality: Caverna Passa Três (about 300 m into the cave),
Rio São Vicente system, São Domingos, Goias, Brazil. Holotype: MZUSP 28809.
Distribution: São Vicente and Angélica-Bezerra Cave system in upper Paranã River, Tocantins River drainage, Brazil (Fisch-Muller, 2003).
Remarks: See Reis (1992) for additional information.
Ancistrus cuiabae Knaack, 1999
Ancistrus cuiabae Knaack, 1999d: 151, figure on p. 150. Type locality: Brazil, Mato Grosso, Pantanal, 36 km SE
Pocone, in a permanent remnant water of Rio Bento Gome [=Gomes]. Holotype: MCP 28671.
Distribution: Cuiabá River basin in upper Paraguay River drainage, Brazil (Fisch-Muller, 2003).
Ancistrus damasceni (Steindachner, 1907)
Xenocara damasceni Steindachner, 1907d: 292. Type locality: dem Parnahyba bei Victoria und Sa. Filomena [Brazil]. Syntypes: NMW 43489 (4), NMW 43490 (7), ZSM 7579, and additional specimens.
Distribution: Upper Parnaíba River basin, Brazil (Fisch-Muller, 2003).
Ancistrus dolichopterus Kner, 1854
Hypostoma punctatum Jardine, in Schomburgk, 1841: 145, fig. on p. 145 (Schomb. Drawings, No 68). Type locality:
Rio Branco [Brazil]. Holotype: not preserved (Isbrücker, 1980: 72). Preoccupied by Hypostomus punctatus
Valenciennes, 1840.
Ancistrus dolichopterus Kner, 1854: 274, pl. 3 (fig. 1). Type locality: Barra do Rio negro [Brazil]. Syntypes: NMW
46276: 1–2 (2), NMW 47164: 1–2 (2). Originally as Anc. dolichopterus.
Distribution: Upper and middle Brazilian Amazon River basin; Negro, lower Trombetas, Tefé, Madeira and Tapajós
River basins, (Fisch-Muller, 2003).

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Ancistrus dubius Eigenmann & Eigenmann, 1889
Ancistrus cirrhosus dubius Eigenmann & Eigenmann, 1889b: 48. Type locality: Gurupa [Brazil]. Holotype: MCZ
7993 (but published as 7983). Name made available by diagnostic features in key on p. 47.
Distribution: Amazon River basin; middle Paraná and Paraguay River basins (Fisch-Muller, 2003).
Remarks: Eschmeyer et al. (1998: 498), Fisch-Muller (2003: 375), and the MCZ fish collection catalog, list MCZ
7993 as a syntype of Ancistrus cirrhosus dubius. However, the notation in Eigenmann & Eigenmann (1889b:48) includes the word “type” only in front of the number 7983 [sic for 7993] for a single specimen from Gurupa,
which appropriately makes that specimen the holotype.
Ancistrus erinaceus (Valenciennes, 1840)
Hypostomus erinaceus Valenciennes, in Cuvier & Valenciennes, 1840b: 510 (376 in Strasbourg deluxe edition).
Type locality: Chili [in error]. Holotype: MNHN a-9568.
Distribution: Unknown; known only from holotype (Fisch-Muller, 2003).
Remarks: Type locality, if intended to refer to Chile, must be in error inasmuch as no specimens of Ancistrus havesubsequently been found in any of the river basins of that country.
Ancistrus eustictus (Fowler, 1945)
Pristiancistrus eustictus Fowler, 1945a: 121, figs. 29–32. Type locality: Alto Rio Baudó, Pacific slope at 3000 feetelevation, Colombia. Holotype: ANSP 71710.
Distribution: Upper Baudó River basin, Colombia; known only from the holotype (Fisch-Muller, 2003).
Ancistrus formoso Sabino & Trajano, 1997
Ancistrus formoso Sabino & Trajano, 1997: 74, fig. 4. Type locality: Buraco do Ducho cave, Serra da Bodoquena,
Município de Bonito (21º08'S, 56º28'W), State of Mato Grosso do Sul, Brazil. Holotype: MZUSP 51386.
Distribution: Buraco do Ducho Cave in Formoso phreatic system, Brazil (Fisch-Muller, 2003).
Ancistrus fulvus (Holly, 1929)
Xenocara fulva Holly, 1929: 119. Type locality: Alto Rio Acara (Brasilien). Holotype: NMW 57203.
Distribution: Acará River basin, lower Amazon drainage, Brazil; known only from the holotype (Fisch-Muller,
2003).
Ancistrus galani Pérez & Viloria, 1994
Ancistrus galani Pérez & Viloria, 1994: 105, fig. 3. Type locality: Cueva de Los Laureles (Zu. 31), Río Socuy, Sierrade Perijá, Zulia State; Venezuela, 72º27'42"W, 10º45'04"N; elevation 650 m. Holotype: MBLUZ 04351.
Distribution: Los Laureles Cave in Socuy River basin, Lake Maracaibo drainage Venezuela (Fisch-Muller, 2003).
Ancistrus gymnorhynchus Kner, 1854
Ancistrus gymnorhynchus Kner, 1854: 275. Type locality: Puerto Cabello [Venezuela]. Holotype: NMW 43495.
Originally as Anc. gymnorhynchus; redescribed and illustrated by Steindachner (1917: 93, pl. 9, figs. 7–8).
Xenocara rothschildi Regan, 1905b: 242. Type locality: San Esteban, near Porto Cabello, Venezuela. Syntypes:
BMNH 1904.11.9.27–31 (5).
Distribution: Coastal rivers of Carabobo and Yaracuy States; upper Pao River basin in Orinoco River drainage, Venezuela (Fisch-Muller, 2003).
Ancistrus heterorhynchus (Regan, 1912)
Xenocara heterorhynchus Regan, 1912b: 668, pl. 76 (fig. 2). Type locality: Uruhuasi [Inambari basin] Peru (4000ft). Syntypes: BMNH 1911.12.20.35–36 (2).
Distribution: Inambari River basin in Madre de Dios River drainage, Peru; known only from type series (FischMuller, 2003).
Ancistrus hoplogenys (Günther, 1864)Chætostomus hoplogenys Günther, 1864: 247. Type locality: River Capin, Para [Brazil]. Syntypes: BMNH
1849.11.8.89–91 (3).
Chaetostomus alga Cope, 1872a: 287, pl. 15 (fig. 3). Type locality: The Ambyiacu river, which empties into the
Amazon near to Pebas, in Eastern Equador, some distance east of the Napo [Peru]. Syntypes: ANSP 16461 (1),ANSP 16462 (1).
Chaetostomus tectirostris Cope, 1872a: 288, pl. 15 (fig. 2). Type locality: The Ambyiacu river, which empties into
the Amazon near to Pebas, in Eastern Equador, some distance east of the Napo [Peru]. Syntypes: ANSP 8298

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(1), ANSP 8300 (1).
Chaetostomus (Ancistrus) cirrhosus punctata Steindachner, 1881c: 123. Type locality: Rio Branco und Rio Guapore
in Mato Grosso, Coary und Tabatinga [Amazonas, Brazil]. Syntypes: NMW 47163 (3), NMW 47279: 1–3 (3),
NMW 47280: 1–2 (2), NMW 47281: 1–2 (2), NMW 47288 (1). Originally as Chaetostomus (Ancistrus) cirrhosus var. punctata.
Distribution: Amazon, Essequibo, and Paraguay River basins (Fisch-Muller, 2003).
Ancistrus jataiensis Fisch-Muller, Cardoso, da Silva & Bertaco, 2005
Ancistrus jataiensis Fisch-Muller, Cardoso, da Silva & Bertaco, 2005b: 567, fig. 4. Type locality: Brazil: Goias:
Mambaí: córrego Jataí, tributary of the rio Vermelho (14º29'S, 46º6'W), rio Tocantins basin. Holotype: MCP
35244.
Distribution: upper Tocantins River basin, Brazil (Fisch-Muller et al., 2005b).
Ancistrus jelskii (Steindachner, 1876)
Chaetostomus Jelskii Steindachner, 1876: 603. Type locality: Amable Maria, Monterico, den Gebirgsbächen derhohen Anden in Peru [Tulumayo River basin, tributary of Péréné River]. Syntypes: at NMW.
Distribution: Tulumayo River basin in upper Ucayali River drainage, Peru (Fisch-Muller, 2003).
Remarks: Validity of species questioned by Fisch-Muller (2003).
Ancistrus latifrons (Günther, 1869)
Chaetostomus latifrons Günther, 1869: 426. Type locality: Upper Amazons, Peruvian Amazons. Holotype: BMNH
1869.5.21.4; holotype illustrated in Regan (1904b: 253, pl. 15, fig. 1).
Distribution: Upper Amazon and Solimões River basins, Brazil, Peru (Fisch-Muller, 2003).
Ancistrus leucostictus (Günther, 1864)
Chaetostomus leucostictus Günther, 1864: 248. Type locality: Essequibo [Guyana]. Possible holotype: BMNH
1864.1.21.85.
Distribution: Essequibo River basin, possibly all main rivers of Guianas (Fisch-Muller, 2003).
Remarks: Type status of specimen registered as holotype questioned by Muller (1989: 886; also Fisch-Muller,
2003).
Ancistrus lineolatus Fowler, 1943
Ancistrus lineolatus Fowler, 1943a: 255, figs. 44–47. Type locality: Florencia, Rio Orteguasa, Colombia. Holotype:
ANSP 70517.
Distribution: Orteguaza River basin in upper Japurá River drainage, Colombia; known only from holotype (FischMuller, 2003).
Ancistrus lithurgicus Eigenmann, 1912
Ancistrus lithurgicus Eigenmann, 1912b: 241, pl. 25 (fig. 3). Type locality: Crab Falls; Lower Essequibo, British
Guiana. Holotype: FMNH 53091
Distribution: Essequibo River basin, Guyana (Fisch-Muller, 2003).
Ancistrus macrophthalmus (Pellegrin, 1912)
Xenocara macrophthalma Pellegrin, 1912: 271, fig. 1. Type locality: l’Orénoque [Venezuela]. Holotype: MNHN
1887-0650.
Distribution: Orinoco River and lower parts of some of its tributaries, Venezuela (Fisch-Muller, 2003).
Ancistrus maculatus (Steindachner, 1881)
Chaetostomus (Ancistrus) cirrhosus maculatus Steindachner, 1881c: 123. Type locality: Cudajas, Obidos, dem Rio
Tajapouru [Brazil], und dem Rio Chagres [Panama]. Syntypes: NMW 47282 (1), NMW 47289: 1–2 (2), NMW
47290 (1), NMW 55035 (1). Originally as Chaetostomus (Ancistrus) cirrhosus var. maculatus.
Distribution: Amazon River basin, Brazil (Fisch-Muller, 2003).
Remarks: Type series mixed; specimens from Chagres River belong to Ancistrus chagresi (Fisch-Muller, 2003). A
lectotype designation of a specimen from Brazil is necessary to stabilize the validity of this name.
Ancistrus malacops (Cope, 1872)
Chaetostomus malacops Cope, 1872a: 287, pl. 5 (figs. 2a–b). Type locality: The Ambyiacu river, which empties into
the Amazon near to Pebas, in Eastern Equador, some distance east of the Napo [Peru]. Syntypes: ANSP 8299
(2).

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Remarks: Validity of species uncertain (Fisch-Muller, 2003).
Ancistrus maracasae Fowler, 1946
Ancistrus maracasae Fowler, 1946: 2, figs. 1–4. Type locality: Maracas River, Trinidad. Holotype: ANSP 71723.
Distribution: Maracas River basin, Trinidad and Tobago; known only from the holotype (Fisch-Muller, 2003).
Ancistrus martini Schultz, 1944
Ancistrus triradiatus martini Schultz, 1944c: 302, pl. 9 (fig. C). Type locality: Río Táchira, 7 km. north of San Antonio, Catatumbo system; Venezuela. Holotype: USNM 121064.
Distribution: Zulia River basin in the Catatumbo River drainage, Lake Maracaibo drainage, Venezuela (FischMuller, 2003).
Ancistrus megalostomus Pearson, 1924
Ancistrus megalostomus Pearson, 1924: 23, pl. 7 (fig. 1). Type locality: Huachi; at the junction of the Bopi andCochabamba rivers, altitude 2,235 feet, Bolivia. Syntypes: CAS 64614 (2).
Distribution: Beni River basin in Madeira River drainage, Bolivia; known only from the syntypes (Fisch-Muller,
2003).
Ancistrus minutus Fisch-Muller, Mazzoni & Weber, 2001
Ancistrus minutus Fisch-Muller, Mazzoni & Weber, 2001: 298, fig. 5 (middle). Type locality: Brazil: Goiás: Minaçú:
córrego Batéias, a left bank tributary of the upper rio Tocantins (13º49'S 48º20'W). Holotype: MNRJ 20851.
Distribution: Upper Tocantins tributaries in Goiás State upstream of the dam of Serra da Mesa, Brazil (Fisch-Muller,
2003).
Ancistrus montanus (Regan, 1904)
Xenocara montana Regan, 1904b: 258, pl. 14 (fig. 6). Type locality: Tumupara [=Tumupasa], Andes of Bolivia,
1500 ft. Holotype: BMNH 1902.12.18.4.
Distribution: Beni River basin in upper Madeira River drainage, Bolivia (Fisch-Muller, 2003).
Ancistrus multispinis (Regan, 1912)
Xenocara multispinis Regan, 1912b: 668, pl. 76 (fig. 1). Type locality: Humboldt and Novo Rivers, Sta. Catherina,
S. E. Brazil [restricted to Humboldt River, Rio Itapocú basin by lectotype designation]. Lectotype: BMNH
1910.7.26.32, illustrated in, and designated by, Muller (1989: 894, fig. 4, tab. 3).
Hemiancistrus albocinctus Ahl, 1936: 111. Type locality: Umbegung von Rio de Janeiro [Brazil]. Holotype: ZMB
20955.
Distribution: Atlantic coastal drainages of eastern Brazil, from Macacú River to Maquiné River (Fisch-Muller,
2003).
Ancistrus nationi (Fernández-Yépez, 1972)
Lasiancistrus nationi Fernández-Yépez, 1972a: 23, pl. 23. Type locality: Venezuela, la estación No 140 del Complejo Hidrográfico (4), Río Yaracuy. Holotype: personal collection of Fernández-Yépez, current whereabouts
unknown.
Distribution: Yaracuy River basin, Venezuela (Fisch-Muller, 2003).
Remarks: Placement in Ancistrus follows Armbruster (2005: 567).
Ancistrus nudiceps (Müller & Troschel, 1848)
Hypostomus nudiceps Müller & Troschel, 1848: 631. Type locality: dem Takutu, Britisch-Guiana. Holotype: ZMB
3180.
Distribution: Takutu River basin in upper Branco River drainage, Guyana, known only from the holotype (FischMuller, 2003).
Remarks: Redescribed in Eigenmann (1912b: 236, footnote 24).
Ancistrus occidentalis (Regan, 1904)
Xenocara occidentalis Regan, 1904b: 257, pl. 14 (fig. 5). Type locality: Canelos [=Chambria], E. Ecuador. Syntypes: BMNH 1880.12.8.69–74 (6).
Distribution: Bobonaza River basin, Ecuador (Fisch-Muller, 2003).
Ancistrus occloi Eigenmann, 1928
Ancistrus occloi Eigenmann, in Myers, 1928: 86. Type locality: Ollantaytambo, Rio Urubamba, Peru, 9000 feet.

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Holotype: CAS 66847.
Distribution: Upper Urubamba River basin, Peru (Fisch-Muller, 2003).
Remarks: Redescribed in Eigenmann & Allen (1942: 197, pl. 10 figs. 1–2).
Ancistrus parecis Fisch-Muller, Cardoso, da Silva & Bertaco, 2005
Ancistrus parecis Fisch-Muller, Cardoso, da Silva & Bertaco, 2005a: 529, fig. 3. Type locality: Brazil: Mato Grosso:
Campos de Júlio: rio Formiga (tributary of rio Juruena, upper rio Tapajós basin), on the road BR-364 (MT-235),
between Campos de Júlio and Sapezal, 13º41'01"S 59º12'11"W. Holotype: MCP 35570.
Distribution: Upper Tapajós River basin, Mato Grosso, Brazil (Fisch-Muller, Cardoso, da Silva & Bertaco, 2005a).
Ancistrus pirareta Muller, 1989
Ancistrus pirareta Muller, 1989: 891, fig. 2 (middle). Type locality: Salto Pirareta, sud de Piribebuy, affluent du Rio
Tebicuary-mi (Cordillera, Paraguay), 25º30'S/56º55'W. Holotype: MHNG 2450.10.
Distribution: Upper Tebicuary-mi River basin in lower Paraguay River drainage, Paraguay (Fisch-Muller, 2003).
Remarks: Redescribed in Muller (1990).
Ancistrus piriformis Muller, 1989
Ancistrus piriformis Muller, 1989: 887, fig. 1 (middle). Type locality: Rio Acaray, en dessous du lac de barrage (AltoParaná, Paraguay), 25º23'S/54º42'W. Holotype: MHNG 2450.11.
Distribution: Acaray River basin in middle Paraná River drainage, Paraguay (Fisch-Muller, 2003).
Remarks: Redescribed in Muller (1990).
Ancistrus ranunculus Muller, Rapp Py-Daniel & Zuanon, 1994
Ancistrus ranunculus Muller, Rapp Py-Daniel & Zuanon, 1994: 290, fig. 1 (middle). Type locality: Brazil, Pará, Rio
Xingú, furo do Tucum Seco, near Arroz Cru. Holotype: INPA 9509.
Distribution: Xingu and Tocantins River basins, Brazil (Fisch-Muller, 2003).
Ancistrus reisi Fisch-Muller, Cardoso, da Silva & Bertaco, 2005
Ancistrus reisi Fisch-Muller, Cardoso, da Silva & Bertaco, 2005b: 564, fig. 3. Type locality: Brazil: Goias: Mambaí:córrego das Dores, tributary of the rio Vermelho (14º29'S, 46º6'W), rio Tocantins basin. Holotype: MCP 34818.
Distribution: upper Tocantins River basin, Brazil (Fisch-Muller et al., 2005b).
Ancistrus spinosus Meek & Hildebrand, 1916
Ancistrus spinosus Meek & Hildebrand, 1916: 252. Type locality: Rio Calobre, tributary of the Rio Bayano, Panama. Holotype: FMNH 8942.
Distribution: Pacific and Caribbean river drainages of eastern Panama (Fisch-Muller, 2003).
Remarks: Fisch-Muller (2003) questioned the identity of the specimen registered as holotype.
Ancistrus stigmaticus Eigenmann & Eigenmann, 1889
Ancistrus stigmaticus Eigenmann & Eigenmann, 1889b: 48. Type locality: Sao Matheos; Goyaz [restricted toGoyaz, = Rio Araguaia drainage, upper course of Rio Vermelho at Goias, 15º47'S, 50º07'W, by lectotype designation]. Lectotype: MCZ 77659, illustrated in, and designated by, Muller (1989: 895, fig. 5). Name made available by diagnostic features in key on p. 47.
Distribution: Upper Araguaia River basin, Brazil (Fisch-Muller, 2003).
Ancistrus tamboensis Fowler, 1945
Ancistrus tamboensis Fowler, 1945c: 9, figs. 13–15. Type locality: Satipo, at 600 meters elevation in upper Tambobasin, Rio Ucayali drainage, Peru. Holotype: ANSP 71643.
Distribution: Tambo River basin in upper Ucayali drainage, Peru, known only from holotype (Fisch-Muller, 2003).
Ancistrus taunayi Miranda Ribeiro, 1918
Ancistrus taunayi Miranda Ribeiro, 1918d: 631. Type locality: Itaqui, Rio Lageado, Rio Grande do Sul [Brazil].
Lectotype: MZUSP 975, designated by Britski (1969: 208) and illustrated in Muller (1989: 896, fig. 6, tab. 3).
Distribution: Middle Uruguay River basin (Fisch-Muller, 2003).
Ancistrus temminckii (Valenciennes, 1840)
Hypostomus Temminckii Valenciennes, in Cuvier & Valenciennes, 1840b: 514 (380 in Strasbourg deluxe edition).
Type locality: Cayenne [in error, interpreted as Suriname, region near Paramaribo, by Boeseman (1972: 305)].
Holotype: RMNH 3123.
Plecostomus aculeatus Gronow, in Gray, 1854: 158. Type locality: America Meridionali. Based on descriptions and

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illustrations in Gronovius (1763: no. 393) and Seba (1734–65: pl. 29, fig. 12). Syntypes: Whereabouts
unknown.
Distribution: Saramacca, Suriname, and Maroni River basins, Suriname (Fisch-Muller, 2003).
Ancistrus tombador Fisch-Muller, Cardoso, da Silva & Bertaco, 2005
Ancistrus tombador Fisch-Muller, Cardoso, da Silva & Bertaco, 2005b: 560, fig. 1. Type locality: Brazil: Mato
Grosso: Porto dos Gaúchos, irarapé Ribeirão Preto, on the road MT-338, about 26 km SE from Porto dos
Gaúchos (11º39'27"S, 57º12'7"W), a tributary of rio Arinos, upper rio Tapajos basin. Holotype: MCP 33000.
Distribution: Arinos River tributaries, upper Tapajós River basin, Brazil (Fisch-Muller, et al., 2005b).
Ancistrus trinitatis (Günther, 1864)
Chaetostomus trinitatis Günther, 1864: 246. Type locality: Trinidad. Syntypes: based on specimens reported on by
Gill (1858: 409) as Ancistrus guacharote, possibly deposited at USNM, but whereabouts unknown.
Distribution: Trinidad Island, Trinidad and Tobago (Fisch-Muller, 2003).
Remarks: See comments on types and question about validity of species in Regan (1904b: footnote on p. 238; 1906:
95). Generic allocation based on Armbruster (2004a: 13), although Armbruster (2005: 567) suggests that eventhat is uncertain.
Ancistrus triradiatus Eigenmann, 1918
Ancistrus triradiatus Eigenmann, 1918b: 680. Type locality: Quebrada Cramalote, Villavicencio; Andes east of
Bogotá [upper Meta basin, Colombia]. Holotype: CAS 60164; holotype illustrated in Eigenmann (1922b: 224,
pl. 13, figs. 3–4).
Distribution: Middle and lower Orinoco River tributaries; Valencia Lake and Los Guayos River basin; southern tributaries of Lake Maracaibo basin (Fisch-Muller, 2003).
Ancistrus variolus (Cope, 1872)
Chaetostomus variolus Cope, 1872a: 288. Type locality: The Ambyiacu river, which empties into the Amazon near
to Pebas, in Eastern Equador, some distance east of the Napo [Peru]. Syntypes: ANSP 21284 (1), ANSP 21285(1).
Distribution: Ampiyacu River basin, Peru (Fisch-Muller, 2003).
Ancistrus verecundus Fisch-Muller, Cardoso, da Silva & Bertaco, 2005
Ancistrus verecundus Fisch-Muller, Cardoso, da Silva & Bertaco, 2005a: 527, fig. 2. Type locality: Brazil:
Rondônia: Vilhena: igarapé Piracolina, about 6 km W of Vilhena, on the road BR-364, upper rio Madeira basin,
12º43'33"S 60º11'34"W. Holotype: MCP 35572.
Distribution: Upper Madeira River basin, Rondônia, Brazil (Fisch-Muller et al., 2005a).
APHANOTORULUS Isbrücker & Nijssen, 1983
Aphanotorulus Isbrücker & Nijssen, 1983a: 105. Type species: Aphanotorulus frankei Isbrücker & Nijssen, 1983.
Type by original designation. Gender: Masculine.
Revision: Armbruster & Page (1996) and Armbruster (1998b).
Aphanotorulus ammophilus Armbruster & Page, 1996
Aphanotorulus ammophilus Armbruster & Page, 1996: 385, fig. 2 (middle). Type locality: Venezuela, Estado
Cojedes, Río San Carlos, R. Portuguesa drainage at Caño Hondo, 2 km west of Las Vegas on the road from Las
Vegas to Campo Allegre – 9º31'51"N, 68º39'39"W. Holotype: INHS 32035.
Distribution: Portuguesa River basin, Venezuela (Weber, 2003).
Aphanotorulus unicolor (Steindachner, 1908)
Plecostomus unicolor Steindachner, 1908f: 164. Type locality: Rio Purús. Lectotype: NMW 44271: 2, designated byArmbruster (1998b: 254).
Plecostomus madeiræ Fowler, 1913: 571, fig. 23. Type locality: Madeira River, about 200 miles east of W. long.
63º54'W, Brazil. Holotype: ANSP 39312.
Plecostomus popoi Pearson, 1924: 20, pl. 3 (fig. 2). Type locality: Popoi River, Upper Río Beni, Bolivia. Holotype:
CAS 77346.
Plecostomus micropunctatus La Monte, 1935: 1, fig. 1. Type locality: Rio Purus: [...] in the vicinity of the mouth of
Rio Macauhan, a tributary of Rio Iaco which, in turn, is a tributary of Rio Purus (70º15'W, 7º30'S). Holotype:

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AMNH 12598.
Plecostomus chaparae Fowler, 1940c: 81, figs. 38–40. Type locality: Boca Chapare, Río Chimore, Cochabamba,
Bolivia. Holotype: ANSP 69067.
Aphanotorulus frankei Isbrücker & Nijssen, 1983a: 108, figs. 1–2. Type locality: [...] du Pérou, Dept. Ucayali, Prov.
Coronel Portillo, système du Rio Aguaytia. Rio Neshua (environ 08º36'S, 74º50'W), 60 km S.O. de Pucallpa, lelong de la route de Tingo Maria. Holotype: ZMA 116.640.
Distribution: Upper Amazon River basin (Weber, 2003).
Remarks: Synonymy based on Armbruster (1998b: 254).
APISTOLORICARIA Isbrücker & Nijssen, 1986
Apistoloricaria Isbrücker & Nijssen, 1986b: 103. Type species: Apistoloricaria condei Isbrücker & Nijssen, 1986.
Type by original designation. Gender: Feminine.
Revision and key to species: Nijssen & Isbrücker (1988).
Apistoloricaria condei Isbrücker & Nijssen, 1986
Apistoloricaria condei Isbrücker & Nijssen, 1986b: 104, fig. 1. Type locality: L’Equateur, Prov. Napo, bassin duNapo (haut Amazone) embouchure du Rio Tiputini dans le Rio Napo au confluent de l’affluent principal, en eau
profunde (00º48.9'S, 75º32.5'O). Holotype: FMNH 94683.
Distribution: Upper Amazon drainage, Napo River basin, Ecuador (Nijssen & Isbrücker, 1988).
Apistoloricaria laani Nijssen & Isbrücker, 1988
Apistoloricaria laani Nijssen & Isbrücker, 1988: 34, figs. 2–4. Type locality: Colombie, Dept. Meta, Río Metica, àEl Aviso, environ 3º59'N, 72º59'O. Holotype: ANSP 131483.
Distribution: Orinoco River drainage, Meta River basin, Colombia (Nijssen & Isbrücker, 1988).
Apistoloricaria listrorhinos Nijssen & Isbrücker, 1988
Apistoloricaria listrorhinos Nijssen & Isbrücker, 1988: 36, figs. 5–7. Type locality: Colombie, Dept. Meta, Río Metica, à environ. 1.5 km E. de Rajote, 3º56'N, 73º03'O. Holotype: ANSP 131482.
Distribution: Orinoco River drainage, Metica River basin, Colombia (Nijssen & Isbrücker, 1988).
Apistoloricaria ommation Nijssen & Isbrücker, 1988
Apistoloricaria ommation Nijssen & Isbrücker, 1988: 37, figs. 8–10. Type locality: Pérou, Dept. Loreto, environs
d’Iquitos, Rio Amazonas (Maranon) entre l’île Iquitos et l’île Lapuna, près du rivage de l’île Lapuna. Holotype:
ANSP 138932.
Distribution: Lower Marañon River, Upper Amazon River drainage, Peru (Nijssen & Isbrücker, 1988).
APOSTURISOMA Isbrücker, Britski, Nijssen & Ortega, 1983
Aposturisoma Isbrücker, Britski, Nijssen & Ortega, 1983: 34. Type species: Aposturisoma myriodon Isbrücker, Britski, Nijssen & Ortega, 1983. Type by original designation. Gender: Neuter.
Aposturisoma myriodon Isbrücker, Britski, Nijssen & Ortega, 1983
Aposturisoma myriodon Isbrücker, Britski, Nijssen & Ortega, 1983: 35, figs. 3–6. Type locality: Pérou, Dept. Ucayali, Prov. Coronel Portillo, bassin du Aguaytia, Rio Huacamayó (environ 09º00'S, 75º29.0'O) près de la roule dePucallpa à Tingo Mario, environ 8 km N.E. du village d’Aguaytia. Holotype: MZUSP 15328.
Distribution: Upper Amazon River drainage, Aguaytia River basin, Peru (Isbrücker et al., 1983).
BARYANCISTRUS Rapp Py-Daniel, 1989
Baryancistrus Rapp Py-Daniel, 1989: 245. Type species: Hypostomus niveatus Castelnau, 1855. Type by original
designation. Gender: Masculine.
Baryancistrus demantoides Werneke, Sabaj, Lujan & Armbruster, 2005
Baryancistrus demantoides Werneke, Sabaj, Lujan & Armbruster, 2005: 535, fig. 2. Type locality: Venezuela, Amazonas, río Orinoco drainage, río Ventuari near ornamental fish market in river, 04.07565º, –066.89285º. MCNG
54029.
Distribution: Orinoco River at confluence with Ventuari River and lower Ventuari River, Venezuela (Werneke,
Sabaj, Lujan & Armbruster, 2005).

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Hemiancistrus longipinnis Kindle, 1895: 255. Type locality: Trocera on Tocantins, Brazil. Holotype: MCZ 29051.
Distribution: Tocantins River basin, Brazil (Fisch-Muller, 2003).
Baryancistrus niveatus (Castelnau, 1855)
Hypostomus niveatus Castelnau, 1855: 43, pl. 21 (fig. 3). Type locality: du Rio Araguay, de la province de Goyaz[Brazil]. Possible syntypes: MNHN a-9453 and MNHN a-9454 (dry specimens).
Distribution: Tocantins, Xingu, Tapajós, and Trombetas River basins, Brazil (Fisch-Muller, 2003).
Remarks: Literature mentions 2 syntypes (e.g., Regan, 1904b, Bertin & Estève, 1950), but the description appears to
be based on a single alcohol-preserved specimen (Fisch-Muller, 2003) which was not found at MNHN.
BROCHILORICARIA Isbrücker & Nijssen, 1979
Brochiloricaria Isbrücker & Nijssen, in Isbrücker, 1979a: 90. Type species: Brochiloricaria chauliodon Isbrücker,
1979. Type by original designation. Gender: Feminine.
Brochiloricaria chauliodon Isbrücker, 1979
Brochiloricaria chauliodon Isbrücker, 1979a: 102, fig. 15. Type locality: Argentine, Prov. Entre Rios, Isla El
Dorado, Paraná Guaza (localité non précisée). Holotype: ZSM 23342.
Distribution: La Plata River basin, Argentina (Isbrücker, 1979).
Brochiloricaria macrodon (Kner, 1853)
Loricaria macrodon Kner, 1853a: 12, 15, pl. 2. Type locality: Aus dem Cujabaflusse [Brazil]. Holotype: NMW
45087. Originally as Lor. macrodon.
Distribution: Paraguay River basin, Brazil.
CHAETOSTOMA Tschudi, 1846
Chaetostoma Tschudi, 1846: 25. Type species: Chaetostoma loborhynchos Tschudi, 1846. Type by monotypy. Gender: Neuter.
Chaetostomus Kner, 1854: 271, 286. Type species: Chaetostoma loborhynchos Tschudi, 1846. Type by being a
replacement name. Unjustified emendation of Chaetostoma Tschudi, 1846. Gender: Masculine.
Hypocolpterus Fowler, 1943a: 259. Type species: Hypocolpterus analis Fowler, 1943. Type by original designation.
Gender: Masculine.
Keys: Schultz (1944c: 287, Venezuela) and Fernández-Yépez (1945: 29, Venezuela).
Monophyly: Schaefer (1986).
Biogeography: Lasso & Provenzano (1998: 62).
Chaetostoma aburrense (Posada, 1909)
Hypostomus aburrensis Posada, 1909: 297. Type locality: Medellín (?). Holotype: Whereabouts unknown.
Distribution: Cauca River basin, Colombia (Fisch-Muller, 2003).
Chaetostoma aequinoctiale Pellegrin, 1909
Chaetostomus aequinoctialis Pellegrin, 1909c: 518. Type locality: Rio Pove, Santo Domingo de los Colorados (560mètres), Équateur. Holotype: MNHN 1904-0017.
Distribution: Guayas River basin, Ecuador (Fisch-Muller, 2003).
Chaetostoma alternifasciatum Fowler, 1945
Chaetostoma alternifasciatum Fowler, 1945a: 116, fig. 21–24. Type locality: Morelia, Rio Caquetá basin, Colombia.
Holotype: ANSP 71711.
Distribution: Caqueta River basin in upper Japurá River drainage, Colombia (Fisch-Muller, 2003).
Chaetostoma anale (Fowler, 1943)
Hypocolpterus analis Fowler, 1943a: 261, figs. 56–59. Type locality: Florencia, Rio Orteguasa, Colombia. Holotype: ANSP 70525.
Distribution: Orteguasa River basin, upper Japurá River drainage, Colombia (Fisch-Muller, 2003).
Chaetostoma anomalum Regan, 1903
Chaetostomus anomalus Regan, 1903a: 599. Type locality: Merida, Venezuela, altitude 1500 meters and from theAlbireggas and Milla rivers, above Merida, altitude 3500 meters. Syntypes: BMNH 1903.6.30.91–101, USNMCHECKLIST OF CATFISHES, RECENT AND FOSSIL

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133135.
Distribution: Lake Maracaibo basin, upper Chama, Capazon and Escalante River basins, Venezuela (Fisch-Muller,
2003).
Chaetostoma branickii Steindachner, 1881
Chaetostomus branickii Steindachner, 1881c: 118, pl. 6 (fig. 1). Type locality: Callacate [Peru]. Syntypes: NMW
47270 (2), NMW 47271 (3).
Distribution: Upper Chamaya River basin in upper Marañon River drainage, Peru (Fisch-Muller, 2003).
Remarks: Fisch-Muller (2003) clarified the type locality, as follows: Cajamarca: Rio Chota (= rio Chotano), alt. ca.
1465–1585 m., ca. 12 km from Cutervo N of Chota; 06º25'S / 78º56'W.
Chaetostoma breve Regan, 1904
Chaetostomus brevis Regan, 1904b: 247, pl. 13 (fig. 3). Type locality: Bomboiza and Zamora Rivers [Santiago Riverbasin], Eastern Ecuador. Syntypes (6): BMNH 1898.11.4.33–36 (4).
Distribution: Zamora River basin in upper Marañon River drainage, Ecuador (Fisch-Muller, 2003).
Chaetostoma brevilabiatum Dahl, 1942
Chaetostomus brevilabiatus Dahl, 1942: 80, fig. 1. Type locality: Rio Volcán near its junction to Rio San Bartolomé(tributary to Rio Magdalena on the left side, between Rios Nare and Ité) municipio of Remedios, department of
Antióquia, Republic of Colombia. Altitude approximately 600 m. Magdalena system. Holotype: at ZMUL.
Distribution: Magdalena River basin, Colombia; known only from the holotype (Fisch-Muller, 2003).
Chaetostoma dermorhynchum Boulenger, 1887
Chaetostomus dermorhynchus Boulenger, 1887c: 277, pl. 22 (upper fig.). Type locality: Canelos [Bobonaza River];eastern Ecuador. Syntypes: BMNH 1880.12.8.64–66 (3).
Distribution: Pastaza, upper Bobonaza, Napo, and upper Curaray River basins, Ecuador (Fisch-Muller, 2003).
Chaetostoma dorsale Eigenmann, 1922
Chaetostomus dorsalis Eigenmann, 1922b: 226. Type locality: Quebrada Cramalote, Villavicencio [Meta Riverbasin, Colombia]. Holotype: CAS 77093.
Distribution: Upper Meta and upper Apuré River basins, Colombia and Venezuela (Fisch-Muller, 2003).
Chaetostoma dupouii Fernández-Yépez, 1945
Chaetostoma dupouii Fernández-Yépez, 1945: 27. Type locality: el rio Encantado, en el sitio indicado en los planos
adjuntos y más o menos a un kilómetro más arriba de la confluencia do los ríos Encantado y Río Grande, tributario éste del Tuy; Venezuela. Holotype: collection of AFY; current whereabouts unknown.
Distribution: Tuy River basin, Venezuela (Fisch-Muller, 2003).
Chaetostoma fischeri Steindachner, 1879
Chaetostomus Fischeri Steindachner, 1879b: 150. Type locality: Mamoni-Fluss, bei Chepo im Isthmus von Panama.
Syntypes: NMW 47170 (1), NMW 47171 (1), NMW 47172 (2), NMW 47173 (1), NMW 54991 (2). Illustrated
and described in more detail in Steindachner (1879f: 162, pl. 4).
Distribution: Pacific versant rivers: Bayano, Tuira, San Juan and Guyas River basins; Caribbean versant Rivers:
Chagres, Atrato, Sinú and Magdalena River basins (Fisch-Muller, 2003).
Chaetostoma greeni Isbrücker, 2001
Chaetostomus maculatus Regan, 1904b: 246, pl. 14 (fig. 4). Type locality: Rozmaiu, Upper Peru. Syntypes: BMNH
1903.10.12.3–4 (2). Preoccupied by Chaetostomus cirrhosus var. maculata Steindachner, 1881; replaced by
Chaetostoma greeni Isbrücker (2001).
Chaetostoma greeni Isbrücker in Isbrücker et al., 2001: 24. Type locality: Rozmaiu, Upper Peru. Syntypes: BMNH
1903.10.12.3–4 (2). Replacement for Chaetostomus maculatus Regan, 1904, preoccupied by Chaetostomus(Ancistrus) cirrhosus var. maculata Steindachner, 1881.
Distribution: Roz River basin, Peru (Fisch-Muller, 2003).
Chaetostoma guairense Steindachner, 1881
Chaetostomus guairensis Steindachner, 1881c: 121, pl. 3 (fig. 1). Type locality: dem Guaire bei Carácas [Venezuela]. Syntypes: NMW 47183 (2), NMW 47184 (2), NMW 47185 (3).
Distribution: Tuy and Guaire River and Lake Valencia basins, Venezuela (Fisch-Muller, 2003).
Chaetostoma jegui Rapp Py-Daniel, 1991

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Chaetostoma jegui Rapp Py-Daniel, 1991: 240, fig. 2 (middle). Type locality: Brazil: Federal Territory of Roraima:
Rio Uraricoera, Maracá Island, rocks above Furo Maracá, 3º21'N 61º26'W. Holotype: INPA 2822.
Distribution: Uraricoera and Tacutu River basins, Branco River drainage, Brazil (Fisch-Muller, 2003).
Chaetostoma lepturum Regan, 1912
Chaetostomus lepturus Regan, 1912b: 667, pl. 75 (fig. 1). Type locality: Rio Tamana, Rio San Juan, Choco, S. W.
Colombia. Syntypes: BMNH 1910.7.11.116–118 (3).
Distribution: Upper San Juan River basin, Colombia (Fisch-Muller, 2003).
Chaetostoma leucomelas Eigenmann, 1918
Chaetostomus leucomelas Eigenmann, 1918b: 681. Type locality: Rio Patia, halfway between the Rios Magui andTelembi [Colombia]. Holotype: CAS 60167; holotype illustrated in Eigenmann (1922b: 84, pl. 12, figs. 5–6).
Distribution: Patia and upper Cauca River basins, Colombia (Fisch-Muller, 2003, Maldonado-Ocampo et al., 2005).
Chaetostoma lineopunctatum Eigenmann & Allen, 1942
Chaetostoma lineopunctata Eigenmann & Allen, 1942: 192, pl. 6 (fig. 1). Type locality: Rio Azupizú [Rio Pachitea
drainage, Peru]. Holotype: CAS 64650.
Distribution: Aguaytia, Pachitea and Pisqui River basins, Ucayali River drainage, Upper Amazon system, Peru
(Fisch-Muller, 2003).
Chaetostoma loborhynchos Tschudi, 1846
Chaetostoma loborhynchos Tschudi, 1846: 26, pl. 4. Type locality: dem Rio Tullumayo in der Montaña de Vitoc, am
Ostabhang der Anden; Peru. Holotype: NMW 47190.
Distribution: Tambo River basin, Ucayali River drainage, Peru (Fisch-Muller, 2003).
Chaetostoma machiquense Fernández-Yépez & Martín Salazar, 1953
Chaetostoma machiquensis Fernández-Yépez & Martín Salazar, 1953: 238, fig. on p. 239. Type locality: Río Negro,
a 16 kms. Oeste de Machiques; Perijá; Hoya del Lago de Maracaibo, Edo. Zulia, Venezuela. Holotype: MHNLS
1558.
Distribution: Negro River basin in the Lake Maracaibo drainage, Venezuela (Fisch-Muller, 2003).
Chaetostoma marcapatae Regan, 1904
Chaetostomus marcapatae Regan, 1904b: 246, pl. 14 (fig. 1). Type locality: Marcapata Valley, E. Peru. Holotype:
BMNH 1902.5.29.211.
Distribution: Inambari River basin in the Madeira River drainage, Peru; known only from the holotype (FischMuller, 2003).
Chaetostoma marginatum Regan, 1904
Chaetostomus marginatus Regan, 1904b: 249, pl. 13 (fig. 1). Type locality: Salidero, N.W. Ecuador. Syntypes (3):
BMNH 1901.8.3.27–28 (2), possibly BMNH 1902.5.27.44 (1), but recorded as coming from Rio Durango inregister.
Distribution: Bogota, Chimbo and San Juan River basins, Colombia, Ecuador (Fisch-Muller, 2003).
Remarks: Fisch-Muller (2003) clarified type locality as follows: Ecuador: Esmeraldas: rio Bogota, near Bulún (=Pulún), alt. 50–110 m, ca. 01º05'N 78º40'W.
Chaetostoma marmorescens Eigenmann & Allen, 1942
Chaetostoma marmorescens Eigenmann & Allen, 1942: 193. Type locality: Huanacachupa Creek, near Huánuco,
elevation 6000 feet. ... A small tumultuous stream from the eastern slope of the Central Cordillera to the Huallaga, a league above Huánuco [Peru]. Holotype: IU 15403 (135 mm specimen); whereabouts unknown, apparently not at CAS.
Distribution: Upper Huallaga River basin, Marañon River drainage, Peru (Fisch-Muller, 2003).
Chaetostoma microps Günther, 1864
Chaetostomus microps Günther, 1864: 250. Type locality: Andes of Western Ecuador. Syntypes: BMNH
1860.6.16.137–143 (7). Syntypes redescribed and illustrated in Regan (1904b: 251, pl. 14, fig. 3)
Distribution: Andes of western Ecuador (Fisch-Muller, 2003).
Chaetostoma milesi Fowler, 1941
Chaetostomus milesi Fowler, 1941c: 2, figs. 1–5. Type locality: Honda, Colombia. Holotype: ANSP 69330.
Distribution: Magdalena River, Colombia, and Apuré River basin, Venezuela (Fisch-Muller, 2003).

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Chaetostoma mollinasum Pearson, 1937
Chaetostomus mollinasus Pearson, 1937: 96, pl. 13 (fig. 1). Type locality: Balsas, 3500 feet, and Cajamarca, 9843
feet, Peru. Syntypes: CAS 64653 (8), CAS 64654 (5).
Distribution: Marañon and Cajamarca River basins, Peru (Fisch-Muller, 2003).
Chaetostoma niveum Fowler, 1944
Chaetostoma niveum Fowler, 1944b: 238, figs. 14–16. Type locality: clear waters of Rio Jurubidá, Nuquí [northwestern Colombia]. Holotype: ANSP 71432.
Distribution: Jurubidá River basin, Colombia (Fisch-Muller, 2003).
Chaetostoma nudirostre Lütken, 1874
Chaetostomus nudirostris Lütken, 1874a: 207. Type locality: Valencia [Venezuela]. Holotype: ZMUC P 30168,
illustrated by Steindachner (1881c: 120, pl. 5 figs. 2, 2a).
Distribution: Lake Valencia basin, Venezuela (Fisch-Muller, 2003).
Chaetostoma palmeri Regan, 1912
Chaetostomus palmeri Regan, 1912b: 667, pl. 75 (fig. 3). Type locality: Rio Tamana, Rio San Juan, Choco, S. W.
Colombia. Syntypes: BMNH 1910.7.11.120–121 (2).
Distribution: Tamana River basin in the San Juan River drainage, Colombia (Fisch-Muller, 2003).
Chaetostoma patiae Fowler, 1945
Chaetostoma patiae Fowler, 1945a: 118, figs. 25–28. Type locality: Rio Patia, at 3000 feet elevation, Pacific slope ofsouthwestern Colombia. Holotype: ANSP 71716.
Distribution: Patia River basin, Colombia (Fisch-Muller, 2003, Maldonado-Ocampo et al., 2005).
Chaetostoma paucispinis Regan, 1912
Chaetostomus paucispinis Regan, 1912b: 667, pl. 75 (fig. 2). Type locality: Tado, Rio San Juan, Choco, Colombia.
Holotype: BMNH 1910.7.11.119.
Distribution: San Juan River basin, Colombia (Fisch-Muller, 2003).
Chaetostoma pearsei Eigenmann, 1920
Chaetostomus pearsei Eigenmann, 1920a: 8, pl. 2. Type locality: Rio Castaño [Lake Valencia basin] at Maracay,
under rocks, Venezuela. Holotype: CAS 64655.
Distribution: Lake Valencia and Tuy River basins, Venezuela (Fisch-Muller, 2003).
Chaetostoma sericeum Cope, 1872
Chaetostomus sericeus Cope, 1872a: 288. Type locality: The Ambyiacu river, which empties into the Amazon near
to Pebas, in Eastern Equador, some distance east of the Napo [Peru]. Lectotype: ANSP 22005, designated by
Fowler (1915a: 234).
Distribution: Ampiyacu River basin, Peru (Fisch-Muller, 2003).
Chaetostoma sovichthys Schultz, 1944
Chaetostoma anomala sovichthys Schultz, 1944c: 292, pl. 8 (fig. b). Type locality: near the bridge over the Río San
Pedro, a tributary of the Río Motatán, southeast of Mene Grande, in the Maracaibo basin; Venezuela. Holotype:
USNM 121053.
Distribution: Lake Maracaibo, Motatán River and tributaries, Venezuela (Fisch-Muller, 2003); Catatumbo Riverbasin, Colombia (Maldonado-Ocampo et al., 2005).
Chaetostoma stannii, 1874
Chaetostomus Stannii Lütken, 1874a: 206. Type locality: Puerto Cabello [Venezuela]. Holotype: ZMUC P 30169,
illustrated in Steindachner (1881c: 120, pl. 5 fig. 4, 4a).
Distribution: Caribbean coastal drainages of Venezuela: Aroa, Tocuyo, Urama and Yaracuy River basins (FischMuller, 2003).
Chaetostoma tachiraense Schultz, 1944
Chaetostoma tachiraensis Schultz, 1944c: 288, pl. 7 (fig.c). Type locality: Río Táchira, 7 km. north of San Antonio,
Estado de Táchira, Venezuela; the Río Táchira is a tributary of the Río Zulia, Catatumbo system, Maracaibo
basin. Holotype: USNM 121052.
Distribution: Catatumbo River basin in the Lake Maracaibo drainage (up to altitudes between 1,000 and 2,000 m
above sea level), Colombia and Venezuela (Fisch-Muller, 2003).

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Chaetostoma taczanowskii Steindachner, 1882
Chaetostomus Taczanowskii Steindachner, 1882b: 177. Type locality: Rio de Tortora [Peru]. Species illustrated anddescribed in more detail in Steindachner (1882c: 23, pl. 5, figs. 2, 2a). Lectotype: NMW 47219 (9 cm); designated by Isbrücker (1980: 64).
Distribution: Huallaga and Urubamba River basins, Peru (Fisch-Muller, 2003).
Remarks: Isbrücker (1980: 64) interpreted original description to be based on a single specimen, which he concluded
must be the holotype. The later description (Steindachner, 1882c: 23, pl. 5, figs. 2, 2a) indicated that more than
two specimens were examined, which requires that the examined specimens be considered as syntypes. However, since there was no clear indication of multiple specimens in the original description, Isbrücker’s (1980:64) interpretation of the 9 cm specimen as the holotype became a de facto lectotype designation. Type locality
interpreted by Fisch-Muller (2003) as: Amazon: Upper rio Huambo basin, Río de la Tortora, 06º26' S 77º18' W.
Redescribed in Eigenmann & Allen (1942: 192).
Chaetostoma thomsoni Regan, 1904
Chaetostomus thomsoni Regan, 1904b: 250, pl. 14 (fig. 2). Type locality: Villeta [Magdalena basin], Colombia. Syntypes: BMNH 1902.5.15.28–30 (3).
Distribution: Magdalena River basin, Colombia (Fisch-Muller, 2003, Maldonado-Ocampo et al., 2005).
Chaetostoma vagum Fowler, 1943
Chaetostomus vagus Fowler, 1943a: 258, figs. 52–55. Type locality: Florencia, Rio Orteguasa, Colombia. Holotype:
ANSP 70521.
Distribution: Orteguaza River basin, upper Japurá River drainage, Colombia (Fisch-Muller, 2003).
Chaetostoma vasquezi Lasso & Provenzano, 1998
Chaetostoma vasquezi Lasso & Provenzano, 1998: 55, fig. 1b. Type locality: Raudales del Caño El Cambur, afluentedel Río Caura, 20 km al suroeste del puente del mismo río, Estado Bolívar, Venezuela. Holotype: MHNLS
8791.
Distribution: Orinoco, Caura and Caroni River basins, Venezuela (Fisch-Muller, 2003).
Chaetostoma venezuelae (Schultz, 1944)
Corymbophanes venezuelae Schultz, 1944d: 41, pl. 1 (fig. 1). Type locality: Río Caripe, Caripito, Venezuela. Holotype: USNM 120752.
Distribution: San Juan River basin, Venezuela (Fisch-Muller, 2003).
Chaetostoma yurubiense Ceas & Page, 1996
Chaetostoma yurubiense Ceas & Page, 1996: 672, fig. 2 (top). Type locality: Río Mayorica (Río Yaracuy system) at
Hwy 3, 9 km N Albarico near the town of Mayorica, State of Lara, Venezuela (10º25'43"N, 68º40'47"W). Holotype: INHS 34942.
Distribution: Yaracuy, Urama and Aroa River basins, Venezuela (Fisch-Muller, 2003).
CORDYLANCISTRUS Isbrücker, 1980
Cordylancistrus Isbrücker, 1980: 48. Type species: Pseudancistrus torbesensis Schultz, 1944. Type by original designation. Gender: Masculine.
Monophyly: Schaefer (1986: 103).
Cordylancistrus daguae (Eigenmann, 1912)
Hemiancistrus daguae Eigenmann, 1912a: 11. Type locality: Caldas, Colombia. Holotype: FMNH 56052, illustrated
in Eigenmann (1922b: 78, pl. 12, figs. 1–2).
Distribution: Dagua River basin, Colombia (Fisch-Muller, 2003).
Cordylancistrus perijae Pérez & Provenzano, 1996
Cordylancistrus perijae Pérez & Provenzano, 1996: 29, fig. 2 (top). Type locality: Venezuela, Zulia State, Maracaibo
basin, Sierra de Perijá, Caño Colorado, tributary to the Palmar River, 100 meters from dam site (under construction), approx. 10º37'12"N 72º25'34"W; elevation 150 m. Holotype: MBLUZ 4413.
Distribution: Palmar and Socuy River basins, Lake Maracaibo drainage, Venezuela (Fisch-Muller, 2003).
Cordylancistrus platycephalus (Boulenger, 1898)
Chaetostomus platycephalus Boulenger, 1898c: 12. Type locality: Rio Bomboiza [Zamora basin], Equateur oriental.

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Syntypes: BMNH 1898.11.4.42 (1), MZUT 1551 (1).
Distribution: Zamora River basin in upper Marañon River drainage, Ecuador (Fisch-Muller, 2003).
Cordylancistrus platyrhynchus (Fowler, 1943)
Hemiancistrus platyrhynchus Fowler, 1943a: 252, figs. 36–39. Type locality: Florencia, Colombia. Holotype: ANSP
70512.
Distribution: Upper Caquetá River basin, upper Japurá River drainage, Colombia (Fisch-Muller, 2003).
Cordylancistrus torbesensis (Schultz, 1944)
Pseudancistrus torbesensis Schultz, 1944c: 296, pl. 8 (fig. c). Type locality: 1 km. above Táriba in the Río Torbes[Apuré River basin], Orinoco system, Venezuela. Holotype: USNM 121001.
Distribution: Torbes River basin in Apuré River drainage, Venezuela (Fisch-Muller, 2003).
CORUMBATAIA Britski, 1997
Corumbataia Britski, 1997: 231. Type species: Corumbataia cuestae Britski, 1997. Type by original designation.
Gender: Feminine.
Corumbataia cuestae Britski, 1997
Corumbataia cuestae Britski, 1997: 234, fig. 3. Type locality: Cachoeira de Analândia, rio Corumbataí, Analândia,
SP [Brazil]. Holotype: MZUSP 51222.
Distribution: Tietê River basin, Brazil (Schaefer, 2003c).
Corumbataia tocantinensis Britski, 1997
Corumbataia tocantinensis Britski, 1997: 237, fig. 4. Type locality: rio Vermelho, Goiás, GO [Brazil]. Holotype:
MZUSP 51223.
Distribution: Araguaia and Tocantins River basins, Brazil (Schaefer, 2003c).
CORYMBOPHANES Eigenmann, 1909
Corymbophanes Eigenmann, 1909a: 5. Type species: Corymbophanes andersoni Eigenmann, 1909. Type by monotypy. Gender: Masculine.
Revision: Armbruster et al. (2000).
Corymbophanes andersoni Eigenmann, 1909
Corymbophanes andersoni Eigenmann, 1909a: 5. Type locality: Aruataima Falls, Upper Potaro River, British Guiana. Holotype: FMNH 52675, illustrated in Eigenmann (1912b: pl. 27, fig.1).
Distribution: Essequibo River drainage, upper Potaro River, upstream Kaieteur Falls, Guyana (Weber, 2003).
Corymbophanes kaiei Armbruster & Sabaj, 2000
Corymbophanes kaiei Armbruster & Sabaj, in Armbruster et al., 2000: 1002, fig. 3 (middle). Type locality: Guyana,
Potaro-Siparuni Region, Oung Creek (Chenapou River-Potaro River Drainage) about one hour hike southwestof coordinates 04º58'26"N, 59º34'41"W (mouth of Chenapou River). Holotype: UG/CSBD F644.
Distribution: Essequibo river drainage, upper Potaro River basin, Guyana (Weber, 2003).
CROSSOLORICARIA Isbrücker, 1979
Crossoloricaria Isbrücker, 1979a: 90. Type species: Loricaria variegata Steindachner, 1879. Type by original designation. Gender: Feminine.
Crossoloricaria bahuaja Chang & Castro, 1999
Crossoloricaria bahuaja Chang & Castro, 1999: 83, fig. 1. Type locality: Peru, Departamento Madre de Dios, Provincia Tambopata, Río Tambopata, 12º47'05"S, 69º18'55"W. Holotype: MUSM 9916.
Distribution: Madre de Dios River basin in Peru, Grande River in Bolivia and Manuripe River in Pando (Chang &Castro, 1999).
Crossoloricaria cephalaspis Isbrücker, 1979
Crossoloricaria cephalaspis Isbrücker, 1979a: 102, fig. 18. Type locality: Colombie, Río Magdalena. Holotype:
BMNH 1947.7.1.228.
Distribution: Magdalena River basin, Colombia (Isbrücker, 1979).
Crossoloricaria rhami Isbrücker & Nijssen, 1983

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Crossoloricaria rhami Isbrücker & Nijssen, 1983b: 9, fig. 1. Type locality: Pérou, Dept. Ucayali, Prov. Coronel Portillo, Bassin du Rio Aguaytia, Rio Huacamayo (ca. 9ºS, 75º29'O) près de la route de Pucallpa à Tingo Maria(Carretera Federico Badadre), à environ 8 km au N. E. du village Aguaytia. Holotype: ZMA 116391.
Distribution: Upper Amazon River basin (Isbrücker & Nijssen, 1983).
Crossoloricaria variegata (Steindachner, 1879)
Loricaria variegata Steindachner, 1879b: 151. Type locality: Mamoni (Mamoni-Flusses, bei Chepo in Isthmus vonPanama). Holotype: NMW 45138. Species illustrated and described in more detail in Steindachner (1879f: 165,
pl. 3).
Distribution: Mamoni, Tuira and Yape River basins in Panama; San Juan and Sinu River basins in Colombia (Ferraris, 2003c).
Crossoloricaria venezuelae (Schultz, 1944)
Loricaria variegata venezuelae Schultz, 1944c: 329, pl. 12 (fig. C). Type locality: Río Palmar at the bridge about 70
km southwest of Maracaibo [Venezuela]. Holotype: USNM 121108.
Distribution: Lake Maracaibo drainage (Schultz, 1944); also Magdalena River basin, Colombia (MaldonadoOcampo et al., 2005).
CTENILORICARIA Isbrücker & Nijssen, 1979
Cteniloricaria Isbrücker & Nijssen, in Isbrücker, 1979a: 91. Type species: Loricaria platystoma Günther, 1868.
Type by original designation. Gender: Feminine.
Cteniloricaria fowleri (Pellegrin, 1908)
Oxyloricaria Fowleri Pellegrin, 1908: 126. Type locality: Rivière Camopi (Guyane française). Holotype: MNHN
1901-0372.
Distribution: Oyapock River basin, Brazil and French Guiana (le Bail et al., 2000).
Remarks: Species redescribed in le Bail et al. (2000).
Cteniloricaria maculata (Boeseman, 1971)
Parasturisoma maculata Boeseman, 1971: 33, pl. 5. Type locality: Sipaliwini, near airstrip, upper Corantijn Riverbasin, Surinam. Holotype: RMNH 26381.
Distribution: Maroni and Corantijn River basins (le Bail et al., 2000).
Remarks: Species redescribed in le Bail et al. (2000).
Cteniloricaria platystoma (Günther, 1868)
Loricaria platystoma Günther, 1868a: 478. Type locality: Surinam. Lectotype: BMNH 1866.8.14.124, designated byIsbrücker (1979a: 113). Illustrated and described in more detail in Günther (1868b: 236, fig. 4).
Distribution: Suriname.
DASYLORICARIA Isbrücker & Nijssen, 1979
Dasyloricaria Isbrücker & Nijssen, in Isbrücker, 1979a: 90. Type species: Loricaria filamentosa Steindachner, 1878.
Type by original designation. Gender: Feminine.
Dasyloricaria capetensis (Meek & Hildebrand, 1913)
Loricaria capetensis Meek & Hildebrand, 1913: 80. Type locality: Río Capetí, Río Tuyra basin, Panama. Holotype:
FMNH 7582.
Distribution: Tuira River basin, Panama (Ferraris, 2003c).
Dasyloricaria filamentosa (Steindachner, 1878)
Loricaria filamentosa Steindachner, 1878a: 90. Type locality: Magdalenen-Stromes [Colombia]. Lectotype: NMW
44874, designated by Isbrücker (1979a: 113). Illustrated and described in more detail in Steindachner (1879d:
45, pl. 9).
Distribution: Magdalena River basin and Catatumbo River, Colombia (Maldonado-Ocampo et al., 2005).
Dasyloricaria latiura (Eigenmann & Vance, 1912)
Loricaria filamentosa latiura Eigenmann & Vance, in Eigenmann, 1912a: 13. Type locality: Boca de Certegai,
Colombia. Syntypes (12): originally CM 3806 and IU 12695, now FMNH 55115 (7), USNM 79219 (1), plus 4
additional specimens (whereabouts unknown).

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Distribution: Tuira River basin, Panama, and Atrato River basin, Colombia (Ferraris, 2003c).
Dasyloricaria seminuda (Eigenmann & Vance, 1912)
Loricaria filamentosa seminuda Eigenmann & Vance, in Eigenmann, 1912a: 13. Type locality: Colombia, Girardot.
Holotype: FMNH 55117; holotype illustrated in Eigenmann (1922b: 91, pl. 14, figs. 1, 2).
Distribution: Magdalena River basin, Colombia (Maldonado-Ocampo et al., 2005).
Dasyloricaria tuyrensis (Meek & Hildebrand, 1913)
Loricaria tuyrensis Meek & Hildebrand, 1913: 81. Type locality: Rio Tuyra, Boca de Cupe, Panama. Holotype:
FMNH 7583.
Distribution: Tuira River basin, Panama (Ferraris, 2003c).
DEKEYSERIA Rapp Py-Daniel, 1985
Dekeyseria Rapp Py-Daniel, 1985: 178. Type species: Dekeyseria amazonica Rapp Py-Daniel, 1985. Type by original designation. Gender: Feminine.
Zonancistrus Isbrücker in Isbrücker et al., 2001: 23. Type species: Ancistrus brachyurus Kner, 1854. Type by original designation. Gender: Masculine.
Remarks: The recently proposed genus Zonancistrus Isbrücker was considered by Fisch-Muller (2003) to be invalid.
Dekeyseria amazonica Rapp Py-Daniel, 1985
Dekeyseria amazonica Rapp Py-Daniel, 1985: 180, fig. 2. Type locality: Complexo do Janauacá, Rio Solimões,Amazonas, Brasil. Holotype: INPA 104.4.
Distribution: Amazon River basin, primarily above Manaus, Brazil (Fisch-Muller, 2003).
Dekeyseria brachyura (Kner, 1854)
Ancistrus brachyurus Kner, 1854: 279, pl. 4 (fig. 1). Type locality: Barra do Rio negro [=Manaus, Brazil]. Holotype:
NMW 47268. Originally as Anc. brachyurus.
Distribution: Lower Negro River basin, Brazil (Fisch-Muller, 2003).
Dekeyseria niveata (La Monte, 1929)
Plecostomus niveatus La Monte, 1929: 3, fig. 2. Type locality: Caño Pescado; about five miles north of Esmeralda,
at an elevation of 325 feet, Upper Orinoco drainage, Venezuela. Holotype: AMNH 9601.
Distribution: Upper Orinoco River basin, Venezuela (Fisch-Muller, 2003).
Dekeyseria picta (Kner, 1854)
Ancistrus pictus Kner, 1854: 277, pl. 4 (fig. 2). Type locality: Barra do Rio negro [=Manaus, Brazil]. Syntypes:
NMW 47200 (2), NMW 47201 (1), NMW 47202 (1). Originally as Anc. pictus.
Distribution: Lower Negro River basin, Brazil (Fisch-Muller, 2003).
Dekeyseria pulchra (Steindachner, 1915)
Aneistrus (Hemiancistrus) pulcher Steindachner, 1915b: 219. Type locality: Rio Negro bei Moura [Brazil]. Holotype: at NMW. Described in more detail and illustrated in Steindachner (1915e: 89, pl. 11, figs. 1, 2). Genericname clearly a misspelling of Ancistrus.
Distribution: Upper Negro and Orinoco rivers and Canal Casiquiare basin, Brazil, Colombia, Venezuela (FischMuller, 2003).
Dekeyseria scaphirhyncha (Kner, 1854)
Ancistrus scaphirhynchus Kner, 1854: 280, pl. 3 (fig. 2). Type locality: Barra do Rio negro [Manaus, Brazil]. Syntypes (4): NMW 47207–10. Originally as Anc. scaphirhynchus.
Distribution: Negro River basin, Brazil (Fisch-Muller, 2003).
Remarks: Redescribed by Rapp Py-Daniel (1985: 181, figs.).
DELTURUS Eigenmann & Eigenmann, 1889
Delturus Eigenmann & Eigenmann, 1889b: 45. Type species: Delturus parahybae Eigenmann & Eigenmann, 1889.
Type by original designation. Gender: Masculine.
Carinotus La Monte, 1933a: 1. Type species: Plecostomus (Carinotus) carinotus La Monte, 1933. Type by original
designation. Gender: Masculine.
Delturus angulicauda (Steindachner, 1877)

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FERRARISPlecostomus angulicauda Steindachner, 1877b: 672, pl. 12. Type locality: Rio Mucuri bei Santa Clara; Rio
Parahyba. Syntypes: NMW 44069–70 (only 2 found).
Distribution: Mucuri River basin, Brazil (Weber, 2003).
Delturus carinotus (La Monte, 1933)
Plecostomus (Carinotus) carinotus La Monte, 1933a: 2, fig. 1(a). Type locality: Rio Doce, Estado do Espirito Santo,
eastern Brazil. Holotype: AMNH 11911.
Distribution: Doce River basin, Brazil.
Delturus parahybae Eigenmann & Eigenmann, 1889
Delturus parahybæ Eigenmann & Eigenmann, 1889b: 45. Type locality: Parahyba [Brazil]. Syntypes (2, 28 cm):
MCZ 7726 (2).
Distribution: Paraíba do Sul River basin, Brazil (Weber, 2003).
DENTECTUS Martín Salazar, Isbrücker & Nijssen, 1982
Dentectus Martín Salazar, Isbrücker & Nijssen, 1982: 127. Type species: Dentectus barbarmatus Martín Salazar,
Isbrücker & Nijssen, 1982. Type by original designation. Gender: Masculine.
Dentectus barbarmatus Mártin Salazar, Isbrücker & Nijssen, 1982
Dentectus barbarmatus Martín Salazar, Isbrücker & Nijssen, 1982: 130, fig. 3. Type locality: Venezuela, Estado
Cojedes, Río Salinas, a tributary of the Río Pao Viejo, N.E. of El Baul (9º13'N, 68º07'W). Holotype: MBUCV
V-12780.
Distribution: Western Orinoco River basin, Venezuela (Mártin Salazar, Isbrücker & Nijssen, 1982).
DOLICHANCISTRUS Isbrücker, 1980
Dolichancistrus Isbrücker, 1980: 47. Type species: Pseudancistrus pediculatus Eigenmann, 1918. Type by original
designation. Gender: Masculine.
Monophyly: Schaefer (1986: 103).
Dolichancistrus atratoensis (Dahl, 1960)
Pseudancistrus atratoênsis Dahl, 1960a: 455, fig. on p. 456. Type locality: Quebrada La Noche, tributary to theupper Atrato, approximate height above sea level 550 m; western Colombia. Holotype: ICNMHN 51.
Distribution: Upper Atrato River basin, Colombia (Fisch-Muller, 2003).
Dolichancistrus carnegiei (Eigenmann, 1916)
Pseudancistrus carnegiei Eigenmann, 1916: 85. Type locality: Rio San Gil, Santander, Colombia. Holotype: FMNH
58350, illustrated in Eigenmann (1922b: 79, pl. 10, fig. 3).
Distribution: Magdalena River basin, Colombia (Fisch-Muller, 2003).
Dolichancistrus cobrensis (Schultz, 1944)
Pseudancistrus pediculatus cobrensis Schultz, 1944c: 299, pl. 9 (fig. B). Type locality: Río Cobre, tributary to Río
Quinta, latter tributary to Río La Grita, below La Grita, Catatumbo system, Maracaibo basin, Venezuela. Holotype: USNM 121036.
Distribution: Catatumbo River basin, Lake Maracaibo drainage, Venezuela (Fisch-Muller, 2003).
Dolichancistrus fuesslii (Steindachner, 1911)
Ancistrus füsslii Steindachner, 1911b: 373. Type locality: Sosomoco, Ostkolumbien, in 800 m Höhe. Holotype:
NMW 48026.
Distribution: Western Colombia; known only from the holotype (Fisch-Muller, 2003).
Dolichancistrus pediculatus (Eigenmann, 1918)
Pseudancistrus pediculatus Eigenmann, 1918b: 679. Type locality: Río Negro at Villavicencio, Colombia. Holotype: FMNH 58352; holotype illustrated in Eigenmann (1922b: 78, 224, fig. 17; pl. 10, fig. 4)
Distribution: Upper Meta River basin, Colombia (Fisch-Muller, 2003); Catatumbo River basin (Maldonado-Ocampo
et al., 2005).
Dolichancistrus setosus (Boulenger, 1887)
Chaetostomus setosus Boulenger, 1887b: 349. Type locality: Andes of Columbia. Syntypes: BMNH 1880.2.26.9–10(2). Redescription of syntypes in Regan (1904b: 239, pl. 12, fig. 2).

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Distribution: Andes of Colombia (Fisch-Muller, 2003).
EPACTIONOTUS Reis & Schaefer, 1998
Epactionotus Reis & Schaefer, 1998: 4. Type species: Epactionotus bilineatus Reis & Schaefer, 1998. Type by original designation. Gender: Masculine.
Epactionotus aky Azpelicueta, Casciotta, Almirón & Körber, 2004
Epactionotus aky Azpelicueta, Casciotta, Almirón & Körber, 2004: 82, figs. 1–2. Type locality: Argentina, Misiones, Río Uruguay basin, Arroyo Garibaldi (26º38'46.1"S, 53º59'55"W). Holotype: AI 124.
Distribution: Yabotí-Guazú basin, Uruguay River system, Misiones, Argentina (Azpelicueta et al., 2004).
Epactionotus bilineatus Reis & Schaefer, 1998
Epactionotus bilineatus Reis & Schaefer, 1998: 5, fig. 3. Type locality: Mouth of arroio do Ouro, which flows into
rio Maquiné, between Barra do Ouro and Maquiné, 29º35'S, 50º17'W, Rio Grande do Sul State, Brazil. Holotype: MCP 20279.
Distribution: Maquiné and Três Forquilhas River basins, Brazil (Schaefer, 2003c).
Epactionotus gracilis Reis & Schaefer, 1998
Epactionotus gracilis Reis & Schaefer, 1998: 12, fig. 6. Type locality: Rio Jordão at Jordão Alto, 28º36'S, 49º29'W,
Nova Veneza, Santa Catarina State, Brazil. Holotype: MCP 20282.
Distribution: Araranguá River basin, Brazil (Schaefer, 2003c).
Epactionotus itaimbezinho Reis & Schaefer, 1998
Epactionotus itaimbezinho Reis & Schaefer, 1998: 9, fig. 5. Type locality: Rio Canoas, tributary of rio Mampituba,
ca. 8 km from Praia Grande in the direction of Mãe dos Homens, 29º14'S, 50º01'W, Rio Grande do Sul, Brazil.
Holotype: MCP 20281.
Distribution: Mampituba River basin, Brazil (Schaefer, 2003c).
Epactionotus yasi Almirón, Azpelicueta & Casciotta, 2004
Epactionotus yasi Almirón, Azpelicueta & Casciotta, 2004: 138, figs. 1–3. Type locality: Argentina, province of
Misiones, río Iguazú basin, arroyo Lobo (25º42'57"S, 54º05'59"W). Holotype: MACN-ict 8649.
Distribution: Iguaçu River basin (Almirón, Azpelicueta & Casciotta, 2004).
EURYCHEILICHTHYS Reis & Schaefer, 1993
Eurycheilus Reis & Schaefer, 1992: 216. Type species: Eurycheilus pantherinus Reis & Schaefer, 1992. Type bymonotypy. Gender: Masculine. Preoccupied by Eurycheilus Septfontaine, 1970, in fossil cephalopods; replacedby Eurycheilichthys Reis & Schaefer, 1993.
Eurycheilichthys Reis & Schaefer, 1993: 894. Type species: Eurycheilus pantherinus Reis & Schaefer, 1992. Typeby being a replacement name. Gender: Masculine. Replacement for Eurycheilus Reis & Schaefer, 1992; preoccupied by Eurycheilus Septfontaine, 1970, in fossil cephalopods.
Eurycheilichthys limulus Reis & Schaefer, 1998
Eurycheilichthys limulus Reis & Schaefer, 1998: 15, fig. 7. Type locality: Rio Jacuí on road RS-010, between Marauand Passo Fundo, 28º18'S, 52º18'W, Rio Grande do Sul State, Brazil. Holotype: MCP 20283.
Distribution: Upper Jacuí River basin, Brazil (Schaefer, 2003c).
Eurycheilichthys pantherinus (Reis & Schaefer, 1992)
Eurycheilus pantherinus Reis & Schaefer, 1992: 217, figs. 1–5. Type locality: Brazil: Rio Grande do Sul, creek tributary of rio dos Touros, at road from Silveira to Rondinha, Bom Jesus, approx. 28º42'S, 50º10'W. Holotype:
MCP 13077.
Distribution: Upper and middle Uruguay River basin, Brazil (Schaefer, 2003c).
EXASTILITHOXUS Isbrücker & Nijssen, 1979
Exastilithoxus Isbrücker & Nijssen, in Isbrücker, 1979a: 91. Type species: Pseudacanthicus fimbriatus Steindachner,
1915. Type by original designation. Gender: Masculine.
Monophyly: Schaefer (1986: 103).
Review: Isbrücker & Nijssen (1985).

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FERRARISExastilithoxus fimbriatus (Steindachner, 1915)
Pseudocanthicus (Lithoxus) fimbriatus Steindachner, 1915a: 201. Type locality: Coquenanfluss [=Kukenan River,also Cuquenan, Caroni basin, Venezuela]. Holotype: NMW 44418. Species illustrated and described in moredetail in Steindachner (1915e: 92, pl. 10, figs. 1–3) with name as Pseudacanthicus (Lithoxus) fimbriatus. Holotype illustrated in Isbrücker & Nijssen (1985).
Distribution: Caroni and Matacuni River basins, Orinoco drainage, Venezuela (Fisch-Muller, 2003).
Exastilithoxus hoedemani Isbrücker & Nijssen, 1985
Exastilithoxus hoedemani Isbrücker & Nijssen, 1985: 227, fig. 3 (middle). Type locality: Brazil, Est. Amazonas, Rio
Marauiá, foot of the Sierra Neblina, Cachoeira “Pora Comeschie”, Rio Negro basin. Holotype: INPA 506.
Distribution: Maurauiá River in the upper Negro River drainage, Brazil (Fisch-Muller, 2003).
FARLOWELLA Eigenmann & Eigenmann, 1889
Acestra Kner, 1853a: 29. Type species: Acestra acus Kner, 1853. Type by subsequent designation by Bleeker (1862–
63: 4). Gender: Feminine. Preoccupied by Acestra Bonaparte, 1846 in fishes and by Dallas, 1852 in Hemiptera;replaced by Farlowella Eigenmann & Eigenmann, 1889.
Farlowella Eigenmann & Eigenmann, 1889b: 32. Type species: Acestra acus Kner, 1853. Type by being a replacement name. Gender: Feminine. Replacement for Acestra Kner, 1853.
Revision: Retzer & Page (1997).
Farlowella acus (Kner, 1853)
Acestra acus Kner, 1853a: 13, 29, pl. 8 (fig. 1). Type locality: Caracas [Venezuela]. Holotype: NMW 47795.
Loricaria scolopacina De Filippi, 1853: 166. Type locality: in rivis prov. Venezuelae, Caracas. Holotype: MZUT 22.
Distribution: Lake Valencia and Torito River basins, Venezuela (Retzer & Page, 1997).
Farlowella amazona (Günther, 1864)
Acestra amazonum Günther, 1864: 261. Type locality: Santarem, River Amazons [Brazil]. Holotype: BMNH
1856.3.25.22.
Acestra gladiolus Günther, 1864: 261. Type locality: Rio Cupai [Brazil]. Lectotype: BMNH 1853.3.19.66, designated by Isbrücker (1979a: 114).
Farlowella carinata Garman in Eigenmann & Eigenmann, 1889b: 32. Type locality: Santarem, Teffé, Gurupa, Obidos, Juthay, Tabatinga [restricted to Gurupá [Brazil] by lectotype designation]. Lectotype: MCZ 9830, designated by Isbrücker (1979a: 114).
Farlowella pseudogladiolus Steindachner, 1910c: 405, figured on p. 405. Type locality: Amazonenstromes ohne
nähere Angabe des Fundortes, wahrscheinlich aus dem Rio negro. Holotype: NMW 46498.
Farlowella paranaënse Meinken, 1937: 78. Type locality: mittleren Paranà -gebiete [Argentina]. Holotype: lost
(Eschmeyer et al., 1998).
Farlowella pleurotaenia Miranda Ribeiro, A., 1939: 360. Type locality: Pará [Brazil]. Holotype: MNRJ 758.
Farlowella oliveirae Miranda Ribeiro, A., 1939: 359. Type locality: Pará [Brazil]. Holotype: MNRJ 757.
Distribution: Amazon, Tocantins, and La Plata River basins (Retzer & Page, 1997).
Remarks: Van der Stigchel (1946: 185) tentatively placed Farlowella gladiolus as a synonym of Farlowella amazonum, thereby acting as first reviser and making F. amazona the valid name for this species.
Farlowella colombiensis Retzer & Page, 1997
Farlowella colombiensis Retzer & Page, 1997: 62, fig. 33. Type locality: Colombia, Meta Department, area of
Guaicaramo, Río Upia drainage. Holotype: SU 23733.
Distribution: Orinoco River drainage, Meta River basin, Colombia (Retzer & Page, 1997).
Farlowella curtirostra Myers, 1942
Farlowella curtirostra Myers, 1942: 102, fig. 11. Type locality: Quebrada Tabor, tributary of the Río Motatan system, 30 km north of Trujillo, Maracaibo basin, Venezuela. Holotype: SU 36508.
Distribution: Lake Maracaibo drainage, Venezuela (Retzer & Page, 1997).
Farlowella gracilis Regan, 1904
Farlowella gracilis Regan, 1904b: 303, pl. 20 (fig. 3). Type locality: Río Caqueta, Cauca Valley, s. Colombia. Holotype: BMNH 1902.5.29.180.

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Farlowella boliviana Steindachner, 1910c: 404. Type locality: Kolombien. Holotype: at NMW, but apparently lost,
Isbrücker (1979a: 114).
Distribution: Caqueta River basin, Upper Amazon River basin, Colombia (Retzer & Page, 1997).
Farlowella hahni Meinken, 1937
Farlowella hahni Meinken, 1937: 77, fig. 2. Type locality: Río Paranà, ca. 2 km east of Ayolas, 27º24'S, 56º46'W,
Misiones Dept., Paraguay. Neotype: UMMZ 228132, designated by Retzer & Page (1997: 76).
Distribution: Middle Paraná River basin, Argentina and Paraguay (Retzer & Page, 1997).
Farlowella hasemani Eigenmann & Vance, 1917
Farlowella hasemani Eigenmann & Vance, 1917: 301, pl. 30 (figs. 1–2). Type locality: Para [Brazil]. Holotype:
FMNH 55089.
Distribution: Pará State, Brazil (Retzer & Page, 1997).
Farlowella henriquei Miranda Ribeiro, 1918
Farlowella henriquei Miranda Ribeiro, 1918d: 636. Type locality: Rio Vermelho, aff. do Araguaya (Sta. Rita das
Antas) [Brazil]. Holotype: MZUSP 2159 [reported as MNRJ 2159 in Retzer & Page (1997: 71)].
Distribution: Araguaia River basin, Brazil (Retzer & Page, 1997).
Farlowella isbrueckeri Retzer & Page, 1997
Farlowella isbruckeri Retzer & Page, 1997: 55, fig. 24. Type locality: Brazil, Mato Grosso State, small river onhighway from Cuibá to Porto Velho, approximately 32 km from Lacerda. Holotype: MZUSP 37641.
Distribution: Upper Paraguay River basin, Brazil (Retzer & Page, 1997).
Farlowella jauruensis Eigenmann & Vance, 1917
Farlowella jauruënsis Eigenmann & Vance, 1917: 300, pl. 30 (fig. 3). Type locality: Jaura [Brazil]. Holotype:
FMNH 55088.
Distribution: Upper Paraguay River basin, Brazil (Retzer & Page, 1997).
Farlowella knerii (Steindachner, 1882)
Acestra Knerii Steindachner, 1882b: 177. Type locality: Canelos [Ecuador]. Lectotype: NMW 47796, designated byIsbrücker (1979a: 114). Illustrated and described in more detail in Steindachner (1882c: 26, pl. 7, figs. 1, 1a).
Distribution: Upper Amazon River drainage, Napo and Pastaza River basins, Ecuador (Retzer & Page, 1997).
Farlowella mariaelenae Martín Salazar, 1964
Farlowella mariaelenae Martín Salazar, 1964: 246, fig. 1. Type locality: río Salinas, brazo del río Pao Viejo, al noreste de El Baúl, brazo del río Portuguesa, Edo. Cojedes [Venezuela]. Holotype: MBUCV V-11938.
Distribution: Western Orinoco River basin and, possibly, Caribbean coastal rivers, Venezuela (Retzer & Page, 1997).
Farlowella martini Fernández-Yépez, 1972
Farlowella martini Fernández-Yépez, 1972a: 22, pl. 15. Type locality: la Estación 114 del Complejo Hidrográfico
(04) “Rio Yaracuy” [Coastal Caribbean drainage, Venezuela]. Syntypes (2; as male and female “holotypes”):
originally in personal collection of Fernández-Yépez, current whereabouts unknown.
Distribution: Caribbean coastal rivers in Venezuela (Retzer & Page, 1997).
Farlowella nattereri Steindachner, 1910
Farlowella nattereri Steindachner, 1910c: 403, figured on p. 404. Type locality: Aus dem mittleren Amazonas-Gebiete ohne nähere Angabe des Fundortes, wahrsheinlich aus dem Rio negro. Holotype: NMW 46497.
Farlowella hargreavesi Eigenmann, 1912b: 252, pl. 32 (fig. 3). Type locality: British Guiana. Holotype: Museum ofGeorgetown (apparently lost); holotype illustrated in Eigenmann (1912b: pl. 32, fig. 3).
Farlowella azygia Eigenmann & Vance, 1917: 299, pl. 29 (figs. 1–2). Type locality: Santarem. Holotype: FMNH
55091.
Farlowella acestrichthys Pearson, 1924: 25, pl. 8 (fig. 2). Type locality: Rurrenabaque, Bolivia. Syntypes: CAS
77322 (2), UMMZ 66480 (1).
Distribution: Upper Amazon and Essequibo River basins (Retzer & Page, 1997).
Farlowella odontotumulus Retzer & Page, 1997
Farlowella odontotumulus Retzer & Page, 1997: 57, fig. 28. Type locality: Ecuador, Sucumbios Province, Río
Aguarico, 4.7 km north of Lago Agrio at bridge on road to El Conejo, 0º07'42"S, 76º52'42"W. Holotype: FMNH
99135.

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FERRARISDistribution: Aguarico River basin in Ecuador and Mavaca River basin in Venezuela (Retzer & Page, 1997).
Farlowella oxyrryncha (Kner, 1853)
Acestra oxyrryncha Kner, 1853a: 13, 31, pl. 8 (fig. 2). Type locality: Rio Mamoré – Cachoeira da Bananeira [Brazil].
Holotype: NMW 47797. Name spelled Acestra oxyrrhyncha in figure captions, Acestra oxyrryncha in text;
Acestra oxyrryncha treated as valid following first reviser action of Isbrücker (1980: 100).
Acestra gladius Boulenger, 1898e: 425, pl. 41 (fig. 2). Type locality: Rio Jurua, an affluent of the Amazons, Brazil.
Lectotype: BMNH 1897.12.1.80, designated by Isbrücker (1979a: 114).
Distribution: Amazon and Orinoco River basins and coastal rivers of NE Brazil (Retzer & Page, 1997).
Farlowella paraguayensis Retzer & Page, 1997
Farlowella paraguayensis Retzer & Page, 1997: 78, fig. 54. Type locality: Paraguay, Canendiyu Department,
Arroyo Carimbatay, 15.6 km WSW Curuguaty, 24º31'06"S, 56º43'30"W. Holotype: UMMZ 228129.
Distribution: Paraguay River basin (Retzer & Page, 1997).
Farlowella platorynchus Retzer & Page, 1997
Farlowella platorynchus Retzer & Page, 1997: 71, fig. 44. Type locality: Peru, Ucayali Department, Pucallpa, Yarinachocha, Río Ucayali drainage, 8º23'S, 74º32'W. Holotype: MHNG 2389.57.
Distribution: Upper and lower Amazon River basin (Retzer & Page, 1997).
Farlowella reticulata Boeseman, 1971
Farlowella reticulata Boeseman, 1971: 37, pl. 7 (right). Type locality: Maka Creek, left tributary of the Lawa River
10 km S of Stoelmanseiland, Marowijne River basin, Surinam. Lectotype: ZMA 106174, designated byIsbrücker (1979a: 114).
Distribution: Essequibo, Marowijne, and Oyapock River basins (Retzer & Page, 1997).
Farlowella rugosa Boeseman, 1971
Farlowella rugosa Boeseman, 1971: 40, pl. 7 (middle). Type locality: Kamaloea (or Saloea) Creek, right tributary of
the Marowijne River, 9 km SE of the outlet of Gran Creek, French Guyane. Lectotype: ZMA 106208, designated by Isbrücker (1979a: 114).
Farlowella parvicarinata Boeseman, 1971: 42, pl. 7 (left). Type locality: Right tributary of the Nickerie River, 12km WSW of Stondansie Falls, Surinam. Lectotype: ZMA 105823, designated by Isbrücker (1979a: 114).
Distribution: Essequibo, Marowijne, and Corantijn River (Retzer & Page, 1997).
Remarks: Farlowella rugosa treated as the valid name following first reviser action of Retzer & Page (1997: 72).
Farlowella schreitmuelleri Ahl, 1937
Farlowella schreitmülleri Ahl, 1937b: 446. Type locality: Santarem [Brazil]: Lectotype ZMB 20956, designated byIsbrücker (1979a: 114).
Farlowella latisoma Miranda Ribeiro, A., 1939: 361. Type locality: Oriximiná, Amazonas [Brazil]. Holotype:
MNRJ 761.
Distribution: Lower Amazon River basin, Brazil (Retzer & Page, 1997).
Farlowella smithi Fowler, 1913
Farlowella smithi Fowler, 1913: 574, fig. 24. Type locality: Rio Madeira, about 200 miles east of W. Long. 62º20',
Brazil. Holotype: ANSP 39313.
Distribution: Middle Madeira River basin, Brazil (Retzer & Page, 1997).
Farlowella taphorni Retzer & Page, 1997
Farlowella taphorni Retzer & Page, 1997: 51, fig. 19. Type locality: Venezuela, Merida State, Río Muyapas (RíoTorondoy-Lago Maracaibo drainage), 4.5 km S of Nueva Bolivai on highway 1. Holotype: INHS 60352.
Distribution: Torondoy River basin, Maracaibo Lake drainage, Venezuela (Retzer & Page, 1997).
Farlowella venezuelensis Martín Salazar, 1964
Farlowella acus venezuelensis Martín Salazar, 1964: 250. Type locality: río Colorado, de la Hoya del río Guarapiche, en San Antonio de Maturín, Monagas [Venezuela]. Lectotype: MBUCV V-26150, designated by Retzer &Page (1997: 64).
Distribution: Guaripiche River basin, lower Orinoco River basin, Venezuela (Retzer & Page, 1997).
Farlowella vittata Myers, 1942
Farlowella vittata Myers, 1942: 103, fig. 12. Type locality: tributaries of the Rio Uribanto, from San Cristobal to

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Ilanos, Venezuela. Holotype: SU 36512.
Farlowella angosturae Martín Salazar, 1964: 251, fig. 3. Type locality: Caño Largo al oeste de Cuidad Bolívar, Edo.
Bolívar [Venezuela]. Holotype: MBUCV V-11939.
Farlowella agustini Martín Salazar, 1964: 254, fig. 5. Type locality: Una quebrada límite entre los Estados Cojedesy Carabobo, carretera Campo Carabobo Taguanes, afluente del río Chirigua que a su vez lo es del río Pao [Venezuela]. Holotype: MBUCV V-11940.
Farlowella guaricensis Martín Salazar, 1964: 253, fig. 4. Type locality: Río Guárico, en el puente de Uverito, carreteria San Juan de Los Morros a Ortiz, Guárico [Venezuela]. Holotype: MBUCV V-11941.
Farlowella roncallii Martín Salazar, 1964: 258, fig. 6. Type locality: la quebrada El Ahorcado Aguirre, afluente delRío Tirgua Carabobo [Venezuela]. Holotype: MBUCV V-11949.
Distribution: Orinoco River basin (Retzer & Page, 1997).
FONCHIIICHTHYS Isbrücker & Michels, 2001
Fonchiiichthys Isbrücker & Michels, in Isbrücker et al., 2001: 18. Type species: Loricaria uracantha Kner, 1863.
Type by original designation. Gender: Masculine.
Fonchiiichthys rupestris (Schultz, 1944)
Loricaria uracantha rupestre Schultz, 1944c: 325, pl. 12 (figs. a–b). Type locality: Río San Pedro at bridge south ofMene Grande, Río Motatán system, Maracaibo basin [Venezuela]. Holotype: USNM 121102.
Distribution: Rivers of the Lake Maracaibo basin, in lower and middle reaches of rivers, Venezuela (Ferraris,
2003c).
Fonchiiichthys uracanthus (Kner, 1863)
Loricaria uracantha Kner, 1863: 228, fig. 18. Type locality: Rio Chagres, Nordseite von Panama. Syntypes (apparently 2): at NMW, probably lost (Isbrücker, 1979a: 112). Originally as Lor. uracantha. Also appeared in Kner &Steindachner (1864: 56, pl. 6, fig. 3).
Loricaria bransfordi Gill, 1877: 338. Type locality: Isthmus of Panama, Empire Station. Holotype: USNM 16673.
Distribution: Atlantic and Pacific versant rivers of Costa Rica and Panama (Bussing, 1998).
Remarks: Diagnosis and ecological notes in Bussing (1998: 163), as Rineloricaria uracantha.
FURCODONTICHTHYS Rapp Py-Daniel, 1981
Furcodontichthys Rapp Py-Daniel, 1981: 2. Type species: Furcodontichthys novaesi Rapp Py-Daniel, 1981. Type byoriginal designation. Gender: Masculine.
Furcodontichthys novaesi Rapp Py-Daniel, 1981
Furcodontichthys novaesi Rapp Py-Daniel, 1981: 4, figs. 1–6. Type locality: rio Solimões, lago Tefé, Caititu, Amazonas, Brasil. Holotype: INPA T.79-014.
Distribution: Solimões River basin, Brazil (Rapp Py-Daniel, 1981).
HARTTIA Steindachner, 1877
Harttia Steindachner, 1877b: 668. Type species: Harttia loricariformis Steindachner, 1877. Type by monotypy. Gender: Feminine.
Quiritixys Isbrücker, in Isbrücker et al., 2001: 21. Type species: Harttia leiopleura Oyakawa, 1993. Type by original
designation. Gender: Feminine.
Harttia carvalhoi Miranda Ribeiro, 1939
Harttia carvalhoi Miranda Ribeiro, P., 1939a: 11, pl. 2 (upper). Type locality: Rio Paquequer, Estado do Rio de Janeiro [Brazil]. Syntypes: MNRJ 1038 (11).
Distribution: Paquequer River basin in Paraíba do Sul River drainage, Brazil.
Harttia depressa Rapp Py-Daniel & Oliveira, 2001
Harttia depressa Rapp Py-Daniel & Oliveira, 2001: 87, fig. 5. Type locality: Brazil: Amazonas, rio Pitinga, rio
Uatumã basin, below the Paranapanema Enterprise dam, Cachoeira 40 Ilhas (59º34'24"W, 0º53'20"S). Holotype: INPA 3901.
Distribution: Uatumã River basin, Brazil (Rapp Py-Daniel & Oliveira, 2001).

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Harttia dissidens Rapp Py-Daniel & Oliveira, 2001
Harttia dissidens Rapp Py-Daniel & Oliveira, 2001: 91, fig. 8. Type locality: Brazil: Pará: rio Tapajós, Pimental(56º15'16"W, 4º35'22"S). Holotype: INPA 7739.
Distribution: Tapajós River basin, Brazil (Rapp Py-Daniel & Oliveira, 2001).
Harttia duriventris Rapp Py-Daniel & Oliveira, 2001
Harttia duriventris Rapp Py-Daniel & Oliveira, 2001: 84, fig. 3. Type locality: Brazil: Pará: rio Tocantins, Igarapé
Bacuri (49º30'32"W, 4º29'44"S). Holotype: INPA 2833.
Distribution: Tocantins River basin, above Tucurui dam, Brazil (Rapp Py-Daniel & Oliveira, 2001).
Harttia garavelloi Oyakawa, 1993
Harttia garavelloi Oyakawa, 1993: 13, fig. 4. Type locality: Rio Fanado, ponte de cidade de Minas Novas, 17º15'S,
42º35'W, MG [Brazil]. Holotype: MZUSP 43266.
Distribution: Fanado and Araçuaí River basins, Brazil (Oyakawa, 1993).
Harttia gracilis Oyakawa, 1993
Harttia gracilis Oyakawa, 1993: 16, fig. 5. Type locality: Riacho afluente dos rio São João municipio de Fortalezade Minas, próximo à estrada de ferro Minas-Perobas, 20º58'S, 46º47'W, MG [Brazil]. Holotype: MZUSP 43267.
Distribution: São João and upper Paraná River basin, Brazil (Oyakawa, 1993).
Harttia guianensis Rapp Py-Daniel & Oliveira, 2001
Harttia guianensis Rapp Py-Daniel & Oliveira, 2001: 88, fig. 6. Type locality: French Guiana: Approuague River,
Saut Athanase (52º1'W, 4º11'N). Holotype: MNHN 1998-0395.
Distribution: Sinnamary and Approuague River basins (Rapp Py-Daniel & Oliveira, 2001).
Harttia kronei Miranda Ribeiro, 1908
Harttia kronei Miranda Ribeiro, 1908b: [2], figs. 3–3a. Type locality: Rio Bethary [Brazil]. Lectotype: MNRJ 713A,
designated by Miranda Ribeiro (1953: 400).
Distribution: Ribeira de Iguape River basin, Brazil.
Harttia leiopleura Oyakawa, 1993
Harttia leiopleura Oyakawa, 1993: 5, fig. 1. Type locality: Riacho afluente do Córrego Mutuca, à direita da estrada
Belo Horizonte Nova Lima, km 20, 20º60'S, 43º55'W, bacia do rio das Velhas MG [Brazil]. Holotype: MZUSP
43264.
Distribution: Das Velhas River basin, Brazil (Oyakawa, 1993).
Harttia longipinna Langeani, Oyakawa & Montoya-Burgos, 2001
Harttia longipinna Langeani, Oyakawa & Montoya-Burgos, 2001: 137, fig. 1. Type locality: Brazil: Minas Gerais
State: rio Pará, bridge on road BR 262 between Nova Serrana and Pará de Minas, approximately 19º53'S,
44º52'W. Holotype: MZUSP 54579.
Distribution: Upper São Francisco River basin, Brazil (Langeani, Oyakawa & Montoya-Burgos, 2001).
Harttia loricariformis Steindachner, 1877
Harttia loricariformis Steindachner, 1877b: 669, pl. 6 (figs. 2, 2a–b). Type locality: Oberer Lafu des Rio Parahyba
und dessen Nebenflüsse. Lectotype: NMW 46346, designated by Isbrücker (1979a: 113).
Distribution: Paraíba do Sul River basin, Brazil.
Harttia merevari Provenzano, Machado-Allison, Chernoff, Willink & Petry, 2005
Harttia merevari Provenzano, Machado-Allison, Chernoff, Willink & Petry, 2005: 520, figs. 1–2. Type locality:
Venezuela, Bolívar State, Caura River at the top of Salto Pará waterfalls, among rocks on the eastern side of theriver. Holotype: MBUCV V-26578.
Distribution: Upper Caura River, Orinoco River basin, Venezuela (Provenzano, et al., 2005).
Harttia novalimensis Oyakawa, 1993
Harttia novalimensis Oyakawa, 1993: 8, fig. 2. Type locality: Riacho afluente do Córrego Mutuca, à direita daestrada Belo Horizonte Nova Lima, km 20, 20º60'S, 43º55'W, bacia do rio das Velhas, MG [Brazil]. Holotype:
MZUSP 43262.
Distribution: Das Velhas River basin, Brazil (Oyakawa, 1993).
Harttia punctata Rapp Py-Daniel & Oliveira, 2001
Harttia punctata Rapp Py-Daniel & Oliveira, 2001: 82, fig. 2. Type locality: Brazil: Pará: rio Tocantins, on rocks

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just below the Tucurui dam (49º38'10"W, 3º49'43"S). Holotype: INPA 4714.
Distribution: Upper and middle Tocantins River basin, Brazil (Rapp Py-Daniel & Oliveira, 2001).
Harttia rhombocephala Miranda Ribeiro, 1939
Harttia rhombocephala Miranda Ribeiro, P., 1939a: 11, pl. 2 (lower). Type locality: Rio Farias [Brazil]. Holotype:
MNRJ 712.
Distribution: Farias River basin, Brazil (Miranda Ribeiro, P., 1939a).
Harttia surinamensis Boeseman, 1971
Harttia surinamensis Boeseman, 1971: 28, pl. 3. Type locality: Grandam, Gran Rio, upper Suriname River, Surinam. Holotype: RMNH 26388 (188 mm specimen, holotype may not have been separated from paratypes).
Distribution: Coastal rivers from Suriname River east to Approuague River, French Guiana (le Bail et al., 2000).
Remarks: Species redescribed by le Bail et al. (2000).
Harttia torrenticola Oyakawa, 1993
Harttia torrenticola Oyakawa, 1993: 10, fig. 3. Type locality: Riacho afluente do rio Paraopebá, Água Limpa,
município de Noeda, 3 km da BR 040, acima das cachoeiras, 44º02'W, 20º20'S, bacia do rio São Francisco, MG[Brazil]. Holotype: MZUSP 43283.
Distribution: Upper São Francisco River basin, Brazil (Oyakawa, 1993).
Harttia trombetensis Rapp Py-Daniel & Oliveira, 2001
Harttia trombetensis Rapp Py-Daniel & Oliveira, 2001: 89, fig. 7. Type locality: Brazil: Pará: rio Trombetas, Cachoeira Vira-Mundo (57º03'38"W, 1º02'04"S). Holotype: INPA 4715.
Distribution: Trombetas River basin, above Porteira Falls, Brazil (Rapp Py-Daniel & Oliveira, 2001).
Harttia uatumensis Rapp Py-Daniel & Oliveira, 2001
Harttia uatumensis Rapp Py-Daniel & Oliveira, 2001: 85, fig. 4. Type locality: Brazil: Amazonas: rio Uatumã, atSanta Luzia (59º28'36"W, 1º50'13"S). Holotype: INPA 2974.
Distribution: Uatumã River basin, Brazil (Rapp Py-Daniel & Oliveira, 2001).
HARTTIELLA Boeseman, 1971
Harttiella Boeseman, 1971: 25. Type species: Harttia crassicauda Boeseman, 1953. Type by original designation.
Gender: Feminine.
Harttiella crassicauda (Boeseman, 1953)
Harttia crassicauda Boeseman, 1953: 10, fig. 2. Type locality: Nassau Mountains, in creek [Suriname]. Holotype:
RMNH 19418 (largest of 15 specimens). Holotype may not have been separated from paratypes.
Distribution: Marowijne River basin, Suriname (le Bail et al., 2000).
Remarks: Species redescribed in le Bail et al. (2000).
HEMIANCISTRUS Bleeker, 1862
Hemiancistrus Bleeker, 1862 (in Bleeker, 1862–63): 2. Type species: Ancistrus medians Kner, 1854. Type by original designation. Gender: Masculine.
Remarks: Monophyly of genus is not supported by anatomical characters (Schaefer, 1986: 105) or mitochondrial
DNA (Montoya-Burgos et al., 2002: 379), but alternate taxonomy has not yet been proposed for the placementof all of the species listed below. However, Armbruster (2004) indicated that some species were clearly distinct
and he listed them in the “Hemiancistrus annectens group.”
Hemiancistrus annectens (Regan, 1904)
Ancistrus annectens Regan, 1904b: 225, pl. 11 (fig. 2). Type locality: St. Javier [San Javier de Cachaví] and the Rio
Durango [tributary of Rio Bogotá], N. W. Ecuador. Syntype (2): BMNH 2005.9.20.1 (1), BMNH 1902.5.27.49(1).
Distribution: Pacific coastal rivers of NW Ecuador (Cayapas basin) and of SW Colombia (Patia basin), Colombia,
Ecuador (Fisch-Muller, 2003).
Hemiancistrus aspidolepis (Günther, 1867)
Chaetostomus aspidolepis Günther, 1867c: 603. Type locality: Veragua [probably Veraguas province], Panama.
Holotype: BMNH 1866.11.5.1 (skin).

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Gatun, Panama; and New Granada. Syntypes: FMNH 26265 (14) and others [see Remarks].
Distribution: Gatun River basin, Caribbean coastal drainage, Panama (Fisch-Muller, 2003).
Remarks: Specimens from several literature sources cited by Eigenmann (1922b: 69) from Panama, including about
150 specimens of Plecostomus plecostomus mentioned by Meek & Hildebrand (1916: 247) should be considered as part of the syntype series of Plecostomus plecostomus panamensis, but only those listed above appear tohave been examined by Eigenmann.
Hemiancistrus chlorostictus Cardoso & Malabarba, 1999
Hemiancistrus chlorostictus Cardoso & Malabarba, 1999: 144, fig. 3. Type locality: Passo Fundo dam, Gerasul
UHPF, Ronda Alta, Rio Grande do Sul, Brazil (approximately 27º40'09"S, 52º45'25"W). Holotype: MCP
21153.
Distribution: Upper Uruguay River basin, Passo Fundo dam, Brazil (Fisch-Muller, 2003).
Hemiancistrus fugleri Ovchynnyk, 1971
Hemiancistrus fugleri Ovchynnyk, 1971: 108, figs. 12–15. Type locality: Ecuador, Province Esmeraldas, Rio
Bogota, Parroquia Cadondelet (78º45' west longitude. 1º6' north latitude). Holotype: MSUM 5133.
Distribution: Cayapas River basin, Ecuador, known only from holotype (Fisch-Muller, 2003).
Hemiancistrus fuliginosus Cardoso & Malabarba, 1999
Hemiancistrus fuliginosus Cardoso & Malabarba, 1999: 148, fig. 6. Type locality: rio Jacutinga, road Seara-Concórdia (BR283), Concórdia, Santa Catarina, Brazil (approx. 27º10'S, 52º09'W). Holotype: MCP 21155.
Distribution: Upper and middle Uruguay River basin, Brazil (Fisch-Muller, 2003).
Hemiancistrus guahiborum Werneke, Armbruster, Lujan & Taphorn, 2005
Hemiancistrus guahiborum Werneke, Armbruster, Lujan & Taphorn, 2005: 544, fig. 2. Venezuela, Amazonas, río
Ventuari at raudales Tencua 56 km ESE of San Juan de Manapiare, 05.04968º, -065.62722º Holotype: MCNG
51994.
Distribution: middle and upper Orinoco River basin (Werneke, Armbruster, Lujan & Taphorn, 2005).
Hemiancistrus hammarlundi Rendahl, 1937
Hemiancistrus hammarlundi Rendahl, 1937: 2, fig. 1 (up). Type locality: Ecuador, Syst. Rio de Clementina [= Rio
Pozuelos] nw. von Babahoyo. Holotype: NRM 10370.
Distribution: Guayas River basin, Ecuador (Fisch-Muller, 2003).
Hemiancistrus holostictus Regan, 1913
Hemiancistrus holostictus Regan, 1913d: 469. Type locality: Not stated [San Juan River basin, Colombia]. Holotype: BMNH 1913.10.1.57.
Distribution: San Juan River basin, Colombia (Fisch-Muller, 2003).
Remarks: Redescribed by Eigenmann (1922b: 73, pl. 9 fig. 4).
Hemiancistrus landoni Eigenmann, 1916
Hemiancistrus landoni Eigenmann, 1916: 84. Type locality: Naranjito [Chimbo tributary, Babahoyo basin], Ecuador. Holotype: CAS 59939; holotype illustrated in Eigenmann (1922b: 72, text fig. 16; pl. 9 fig. 3, pl. 10 fig. 2).
Distribution: Guayas River basin, Ecuador (Fisch-Muller, 2003).
Hemiancistrus macrops (Lütken, 1874)
Chaetostomus macrops Lütken, 1874a: 209. Type locality: Surinam. Holotype: ZMUC P 30142, illustrated in Steindachner (1881c: 124, pl. 5, figs. 3–3a).
Distribution: Suriname (Fisch-Muller, 2003).
Hemiancistrus maracaiboensis Schultz, 1944
Hemiancistrus maracaiboensis Schultz, 1944c: 317, pl. 10 (fig. c). Type locality: near the mouth of the Río Concha
in Lago Maracaibo; Venezuela. Holotype: USNM 121012.
Distribution: Lake Maracaibo basin, Venezuela (Fisch-Muller, 2003).
Hemiancistrus medians (Kner, 1854)
Ancistrus medians Kner, 1854: 256. Type locality: [No locality stated]. Holotype: at NMW. Originally as Anc. medians.
Distribution: Maroni River basin, French Guiana and Suriname (Fisch-Muller, 2003).

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Remarks: Type locality interpreted by Isbrücker (1992b) as Rivière Marouini, Maroni system, French Guiana.
Hemiancistrus megacephalus (Günther, 1868)
Chaetostomus megacephalus Günther, 1868a: 476. Type locality: Surinam. Holotype: BMNH 1978.9.12.3.
Described in more detail in Günther (1868b: 232).
Distribution: imprecise locality in Suriname, and Essequibo basin, Guyana (Fisch-Muller, 2003).
Hemiancistrus megalopteryx Cardoso, 2004
Hemiancistrus megalopteryx Cardoso, 2004: 174, fig. 1. Type locality: Brazil: Santa Catarina: Braço do Norte: rio
Braço do Norte, downstream dam (tributary of rio Tubarão), approx. 28º16'38"S 49º11'1"W. Holotype: MCP
32229.
Distribution: Tubarão River basin, Santa Catarina State, Brazil (Cardoso, 2004).
Hemiancistrus meizospilos Cardoso & Pezzi da Silva, 2004
Hemiancistrus meizospilos Cardoso & Pezzi da Silva, 2004: 4, fig. 6. Type locality: Brazil: Santa Catarina: Coronel
Freitas: rio Chapecó (tributary of the rio Uruguai basin). Holotype: MCP 34091.
Distribution: Chapecó River system, Uruguay River basin, Brazil (Cardoso & Pezzi da Silva, 2004).
Hemiancistrus micrommatos Cardoso & Lucinda, 2003
Hemiancistrus micrommatos Cardoso & Lucinda, 2003: 77, fig. 3 (lower). Type locality: Brazil: Goiás; rio
Tocantins, at site of future hydroelectric dam of Serra da Mesa, between Minaçu & Cavalcante, Minaçu,
approximatively 13º50'S 48º19'W. Holotype: MNRJ 20873.
Distribution: Lower Amazon Basin, Upper Rio Tocantins and tributaries, Brazil (Fisch-Muller, 2003).
Hemiancistrus punctulatus Cardoso & Malabarba, 1999
Hemiancistrus punctulatus Cardoso & Malabarba, 1999: 152, fig. 8. Type locality: rio dos Sinos, João Fernandes
beach, Caraá, Rio Grande do Sul, Brazil (approx. 29º45'87"S, 50º25'68"W). Holotype: MCP 21154.
Distribution: Laguna dos Patos basin, Brazil (Fisch-Muller, 2003).
Hemiancistrus spilomma Cardoso & Lucinda, 2003
Hemiancistrus spilomma Cardoso & Lucinda, 2003: 75, fig. 2 (lower). Type locality: Brazil: Tocantins; Porto
Franco, rio Tocantins near mouth of rio Lajeado. Holotype: MCP 28425.
Distribution: Lower Amazon Basin, lower and upper Tocantins drainage, Brazil (Cardoso & Lucinda, 2003).
Hemiancistrus spinosissimus Cardoso & Lucinda, 2003
Hemiancistrus spinosissimus Cardoso & Lucinda, 2003: 79, fig. 4 (lower). Type locality: Brazil: Goiás; rio Maranhão, near mouth of the rio Tocantinzinho, border of Niquelândia and Campinaçu, 13º56'25''S 48º19'08''W.
Holotype: MCP 26239.
Distribution: Lower Amazon Basin, Middle and Upper Rio Tocantins and tributaries, Brazil (Cardoso & Lucinda,
2003).
Hemiancistrus subviridis Werneke, Sabaj, Lujan & Armbruster, 2005
Hemiancistrus subviridis Werneke, Sabaj, Lujan & Armbruster, 2005: 538, fig. 4. Type locality: Venezuela, Amazonas, río Orinoco 117 km W of La Esmeralda, 03.28998º, –66.60004º. Holotype: MCNG 54032.
Distribution: Upper Orinoco River, Venezuela (Werneke, Sabaj, Lujan & Armbruster, 2005).
Hemiancistrus votouro Cardoso & Pezzi da Silva, 2004
Hemiancistrus votouro Cardoso & Pezzi da Silva, 2004: 2, fig. 1. Type locality: Brazil: Rio Grande do Sul: Benjamin Constant: arroio Lageado Grande (tributary of the rio Passo Fundo, Rio Uruguai basin), ca. 2.5 km NE
from Votouro Indian Reserve (27º26'50"S, 52º37'5"W). Holotype: MCP 33594.
Distribution: Passo Fundo system, Uruguay River basin, Brazil (Cardoso & Pezzi da Silva, 2004).
Hemiancistrus wilsoni Eigenmann, 1918
Hemiancistrus wilsoni Eigenmann, 1918b: 678. Type locality: Truando [tributary of Río Atrato, Colombia]. Holotype: FMNH 58550; holotype illustrated in Eigenmann (1922b: 74, pl. 10, fig. 1).
Distribution: Truando River basin, Atrato River drainage, Colombia (Fisch-Muller, 2003).
HEMILORICARIA Bleeker, 1862
Hemiloricaria Bleeker, 1862 (in Bleeker, 1862–63): 3. Type species: Hemiloricaria caracasensis Bleeker, 1862.
Type by original designation. Gender: Feminine.

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Remarks: Removed from the synonymy of Rineloricaria in Isbrücker (2001: 23).
Hemiloricaria altipinnis (Breder, 1925)
Loricaria altipinnis Breder, 1925: 1, fig. 1. Type locality: Río Chico, Darien, Panama. Holotype: AMNH 8404.
Distribution: Chico River basin, Panama (Ferraris, 2003c).
Hemiloricaria aurata Knaack, 2002
Hemiloricaria aurata Knaack, 2002b: 58, unnumbered figures. Type locality: Mündang eines lkeinen Zuflusses in
den Rio Aquidaban, Paraguay, Dpto. de Concepcion, Strecke Loreto-San Carlos. Holotype: ZMA 123591.
Distribution: Paraguay (Knaack, 2002b).
Hemiloricaria beni (Pearson, 1924)
Loricaria beni Pearson, 1924: 25, pl. 5 (fig. 4). Type locality: Lagoons, Lake Rogoagua [Beni River basin, Bolivia].
Holotype: CAS 28772.
Distribution: Beni River basin, Bolivia (Ferraris, 2003c).
Hemiloricaria cacerensis (Miranda Ribeiro, 1912)
Loricaria cacerensis Miranda Ribeiro, 1912: 11. Type locality: Caceres, M. Grosso (Aguas do Paraguay) [Brazil].
Lectotype: MNRJ 645A, designated by Miranda Ribeiro (1953: 400).
Distribution: Mato Grosso, Brazil (Ferraris, 2003c).
Hemiloricaria caracasensis Bleeker, 1862
Loricaria rostrata van der Hoeven, 1849: 90. Type locality: Caracas [Venezuela]. Holotype: RMNH 3120. Preoccupied by Loricaria rostrata Spix & Agassiz, 1829.
Hemiloricaria caracasensis Bleeker, 1862 (in Bleeker, 1862–63): 3. Type locality: Caracas [Venezuela]. Holotype:
RMNH 3120; holotype illustrated by Boeseman (1972: pl. 1), as Hemiloricaria caracassensis.
Distribution: Caribbean coastal drainages, Venezuela (Ferraris, 2003c).
Remarks: Hemiloricaria caracasensis was incorrectly reported to be a replacement name for Loricaria rostrata in
Ferraris (2003c). The two names were actually independently based on the same specimen.
Hemiloricaria castroi (Isbrücker & Nijssen, 1984)
Rineloricaria castroi Isbrücker & Nijssen, 1984a: 93, fig. 1. Type locality: Brazil, Est. Pará, Trapiche da Reserva
Biológica de Trombetas, Rio Trombetas. Holotype: MZUSP 15731.
Distribution: Trombetas River basin, Brazil (Ferraris, 2003c).
Hemiloricaria eigenmanni (Pellegrin, 1908)
Loricaria Eigenmanni Pellegrin, 1908: 125. Type locality: Sarare (Venezuela). Lectotype: MNHN 1899-0118, designated by Isbrücker (1979a: 113).
Distribution: Venezuela (?).
Hemiloricaria fallax (Steindachner, 1915)
Loricaria (Loricariichthys) fallax Steindachner, 1915e: 101, pl. 13 (figs. 1–4). Type locality: Rio Rupununi, dem
Rio Parimé dei Serra do Banco, aus einem Igarapé bei Capivari, dem Rio Branco bei Bem Querer und Conçeicão, einem Flusse bei Magauarý an der Braganzabahn bei Pará. Lectotype: NMW 44864, designated byIsbrücker & Nijssen (1979: 198).
Distribution: Upper Rupununi and Branco River basins (Ferraris, 2003c).
Hemiloricaria formosa (Isbrücker & Njissen, 1979)
Rineloricaria formosa Isbrücker & Nijssen, 1979: 192, fig. 1. Type locality: Colombia, Lagoon about 1 km upriverfrom Puerto Inírida, Río Orinoco basin. Holotype: FMNH 83713.
Distribution: Western Orinoco and upper Amazon River basins (Ferraris, 2003c).
Hemiloricaria hasemani (Isbrücker & Nijssen, 1979)
Rineloricaria hasemani Isbrücker & Nijssen, 1979: 200, fig. 4. Type locality: Brazil, Est. Pará, Maguarý near
Belém, along the Bragança railroad (Bragança, 1º02'S, 46º46'W), in forest streams. Holotype: NMW 44865.
Distribution: Lower Amazon or Tocantins River basin, Brazil (Ferraris, 2003c).
Hemiloricaria hoehnei (Miranda Ribeiro, 1912)
Loricaria hoehnei Miranda Ribeiro, 1912: 11. Type locality: Coxim, no Rio Paraguay [Brazil]. Holotype: MNRJ
650.
Distribution: Paraguay River basin, Brazil (Ferraris, 2003c).

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Hemiloricaria jubata (Boulenger, 1902)
Loricaria jubata Boulenger, 1902a: 70. Type locality: Northwestern Ecuador, St. Javier (60 feet), and the Rio
Durango, elev. (350 feet) [restricted to the Durango River by lectotype designation]. Lectotype: BMNH
1902.5.27.45, designated by Isbrücker (1979a: 112).
Distribution: Mira and Esmeraldas River basins, Pacific drainage, Ecuador (Ferraris, 2003c); and Patia River basin,
Colombia (Maldonado-Ocampo et al., 2005).
Hemiloricaria konopickyi (Steindachner, 1879)
Loricaria konopickyi Steindachner, 1879a: 34. Type locality: dem mittleren Laufe des Amazonenstromes. Holotype:
NMW 44934. Illustrated and described in more detail in Steindachner (1879e: 45 pl. 6, fig. 3).
Distribution: Amazon River basin, Brazil (Ferraris, 2003c).
Hemiloricaria lanceolata (Günther, 1868)
Loricaria lanceolata Günther, 1868a: 477. Type locality: Xeberos. Holotype: BMNH 1867.6.13.79. Described in
more detail in Günther (1868b: 235, fig. 3). Holotype redescribed and illustrated in Isbrücker (1973: 78, fig. 1)
as Rineloricaria lanceolata.
Distribution: Upper Amazon River basin, Peru (Ferraris, 2003c).
Hemiloricaria magdalenae (Steindachner, 1879)
Loricaria Magdalenae Steindachner, 1879d: 74. Type locality: Magdalenen-Stromes [Colombia]. Lectotype: NMW
45080, designated by Isbrücker (1979a: 112); syntype illustrated in Boeseman (1972: pl. 2).
Distribution: Sinú, Magdalena and, possibly, Catatumbo River basins, Colombia (Ferraris, 2003c).
Hemiloricaria melini (Schindler, 1959)
Loricariichthys melini Schindler, 1959: 387, fig. 1. Type locality: Rio Solimoes beim Ort Manacapuru, nahe derMündung des Rio Negro [Brazil]. Holotype: NRM 10830.
Distribution: Amazon River basin, Brazil (Ferraris, 2003c).
Hemiloricaria morrowi (Fowler, 1940)
Rhineloricaria morrowi Fowler, 1940a: 244, figs. 39– 41. Type locality: Ucayali River basin, Contamana, Peru.
Holotype: ANSP 68663.
Distribution: Ucayali River basin, Peru (Ferraris, 2003c).
Hemiloricaria nigricauda (Regan, 1904)
Loricaria nigricauda Regan, 1904b: 275, pl. 16 (fig. 2). Type locality: Porto Real, Province Rio Janeiro [Brazil].
Lectotype: BMNH 1891.6.16.32, designated by Isbrücker (1979a: 113).
Distribution: Coastal streams in Rio de Janeiro State, Brazil (Ferraris, 2003c).
Hemiloricaria parva (Boulenger, 1895)
Loricaria parva Boulenger, 1895b: 527. Type locality: Descalvados, Matto Grosso [Brazil]. Lectotype: BMNH
1895.5.17.91, designated by Isbrücker (1979a: 112); one syntype illustrated in Boulenger (1896: pl. 7, fig. 1).
Distribution: Paraguay River basin, Argentina and Brazil (Ferraris, 2003c).
Hemiloricaria phoxocephala (Eigenmann & Eigenmann, 1889)
Loricaria phoxocephala Eigenmann & Eigenmann, 1889b: 37. Type locality: Coary [Brazil]. Lectotype: MCZ 8030,
designated by Isbrücker (1979a: 112).
Distribution: Amazon River basin, Brazil (Ferraris, 2003c).
Hemiloricaria platyura (Müller & Troschel, 1848)
Loricaria platyura Müller & Troschel, 1848: 631. Type locality: dem Rupununi [Guyana]. Holotype: ZMB 3166.
Loricaria submarginatus Eigenmann, 1909a: 10. Type locality: Creek below Potaro Landing [Guyana]. Holotype:
FMNH 53080.
Distribution: Various coastal rivers from near Amazon mouth to Essequibo River, Guyana (le Bail et al., 2000).
Remarks: Species redescribed in le Bail et al. (2000).
Hemiloricaria sneiderni (Fowler, 1944)
Loricaria sneiderni Fowler, 1944b: 240, figs. 17–19. Type locality: Río Jurubidá, Nuquí, in brook [Colombia].
Holotype: ANSP 71433.
Distribution: Jurubidá River basin, Pacific Drainage, Colombia (Ferraris, 2003c).
Hemiloricaria stewarti (Eigenmann, 1909)

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Loricaria stewarti Eigenmann, 1909a: 9. Type locality: Chipoo Creek, a tributary of the Ireng [Guyana]. Holotype:
FMNH 53330; holotype illustrated in Eigenmann (1912b: pl. 30, fig. 5).
Distribution: Coastal rivers of the Guianas, and upper Branco River Basin (Ferraris, 2003c).
Remarks: Species redescribed by le Bail et al. (2000).
Hemiloricaria teffeana (Steindachner, 1879)
Loricaria teffeana Steindachner, 1879a: 34. Type locality: Teffé, Amazonenstrom [Brazil]. Lectotype: NMW 45134:
1, designated by Isbrücker (1979a: 112). Illustrated and described in more detail in Steindachner (1879e: 44, pl.
6, fig. 2).
Loricaria Valenciennesii Vaillant, 1880b: 157. Type locality: Calderon [Brazil]. Holotype: MNHN a-1985.
Distribution: Amazon River basin, Brazil (Ferraris, 2003c).
Hemiloricaria wolfei (Fowler, 1940)
Rhineloricaria wolfei Fowler, 1940a: 241, figs.33–35. Type locality: Ucayali River basin, Contamana, Peru. Holotype: ANSP 68660.
Distribution: Ucayali River basin, Peru (Ferraris, 2003c).
HEMIODONTICHTHYS Bleeker, 1862
Hemiodontichthys Bleeker, 1862 (in Bleeker, 1862–63): 4. Type species: Hemiodon acipenserinus Kner, 1853. Typeby original designation. Gender: Masculine.
Hemiodontichthys acipenserinus (Kner, 1853)
Hemiodon acipenserinus Kner, 1853a: 12, 28, pl. 7 (fig. 2a–c). Type locality: Rio Guaporé, Matogrosso (Juquia)[Brazil]. Lectotype: NMW 46139, designated by Isbrücker & Nijssen (1974b: 204). Originally as Hem. acipenserinus.
Distribution: Amazon, Essequibo, Oyapock, and Paraguay River basins (Ferraris, 2003c).
Remarks: Species redescribed by le Bail et al. (2000).
HEMIPSILICHTHYS Eigenmann & Eigenmann, 1889
Xenomystus Lütken, 1874a: 217. Type species: Xenomystus gobio Lütken, 1874. Type by original designation. Gender: Masculine. Preoccupied by Xenomystus Günther, 1868, in fishes; replaced by Hemipsilichthys Eigenmann& Eigenmann, 1889.
Hemipsilichthys Eigenmann & Eigenmann, 1889b: 46. Type species: Xenomystus gobio Lütken, 1874. Type by
being a replacement name. Gender: Masculine. Replacement name for Xenomystus Lütken, 1874, preoccupied
by Xenomystus Günther, 1868, in fishes.
Upsilodus Miranda Ribeiro, 1924: 365. Type species: Upsilodus victori Miranda Ribeiro, 1924. Type by monotypy.
Gender: Masculine.
Remarks: The name Psilichthys is found in Steindachner (1907a: 82) and treated in subsequent literature as the proposal of a new generic name, with Psilichthys cameroni Steindachner, 1907, as type. However the name isapparently a lapsus calami for Hemipsilichthys, which is used extensively in Steindachner (1907e), including
mention of its being a replacement for Xenomystus Lütken.
Revision and key to species: Pereira & Reis (2002), but with most species subsequently transferred into Pareiorhaphis by Pereira (2005).
Hemipsilichthys gobio (Lütken, 1874)
Xenomystus gobio Lütken, 1874a: 217, pl. 4 (fig. 1). Type locality: not stated. Holotype: ZMUC 30173.
Upsilodus victori Miranda Ribeiro, 1924: 366. Type locality: Paquequer ad Theresopolis, Rio de Janeiro [Brazil].
Lectotype: MNRJ 639, designated by Gosline (1947: 100).
Distribution: Paraíba do Sul River basin, Brazil (Weber, 2003).
Remarks: Type locality of Xenomystus gobio given by Pereira et al. (2000: 382) as Paraíba do Sul River basin, Brazil.
Hemipsilichthys nimius Pereira, Reis, Souza & Lazzarotto, 2003
Hemipsilichthys nimius Pereira, Reis, Souza & Lazzarotto, 2003: 5, fig. 1. Type locality: Brazil: Rio de Janeiro,
Parati, rio Carrasquinho below the Cachoeira do Tobogã, upper Perequê-Açu Basin, Penha, ca. 7.5 km west of

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highway BR 101, on road from Parati to Cunha (23º12'51"S, 44º47'28"W. Holotype: MCP 33049.
Distribution: Perequê-Açu and Taquari Rivers, Rio de Janeiro State, Brazil (Pereira et al., 2003).
Hemipsilichthys papillatus Pereira, Oliveira & Oyakawa, 2000
Hemipsilichthys papillatus Pereira, Oliveira & Oyakawa, 2000: 378, fig. 1. Type locality: Brazil: Minas Gerais: rio
Paraíba do Sul drainage, County of Rio Preto, ribeirão Santana, tributary of rio Preto (approx. 22º02'S
43º47'W). Holotype: MZUSP 53085.
Distribution: Preto River basin of Paraíba do Sul River drainage, Brazil (Weber, 2003).
HISONOTUS Eigenmann & Eigenmann, 1889
Hisonotus Eigenmann & Eigenmann, 1889b: 40. Type species: Hisonotus notatus Eigenmann & Eigenmann, 1889.
Type by original designation. Gender: Masculine.
Hisonotus depressicauda (Miranda Ribeiro, 1918)
Otocinclus depressicauda Miranda Ribeiro, 1918d: 635. Type locality: Sorocaba [Paraná River basin, Brazil]. Lectotype: MZUSP 1004, designated by Britski (1969: 208).
Distribution: Tietê River basin, Brazil (Schaefer, 2003c).
Hisonotus depressinotus (Miranda Ribeiro, 1918)
Microlepidogaster depressinotus Miranda Ribeiro, 1918d: 633. Type locality: Piracicaba [São Paulo State, Brazil].
Holotype: MZUSP 2156.
Distribution: Tietê River basin, Brazil (Schaefer, 2003c).
Hisonotus francirochai (Ihering, 1928)
Otocinclus francirochai Ihering, 1928a: 2, fig. 1. Type locality: Creeks by Pirangy, headwaters of the rio Turvo, Brazil. Lectotype: MZUSP 3258, designated by Britski (1969: 209).
Distribution: Grande River basin, Brazil (Schaefer, 2003c).
Hisonotus insperatus Britski & Garavello, 2003
Hisonotus insperatus Britski & Garavello, 2003: 588, fig. 1. Type locality: Brazil, São Paulo State, Botucatu, Rio
Capivara, Rio Tietê drainage. Holotype: MZUSP 78957.
Distribution: Upper Paraná River basin, São Paulo and Mato Grosso states, Brazil (Britski & Garavello, 2003).
Hisonotus laevior Cope, 1894
Hisonotus laevior Cope, 1894: 95, pl. 8 [sic, 7] (figs. 12, 12a–b). Type locality: Rio Jacuhy, Rio Grande do Sul, Brazil. Holotype: ANSP 21563.
Distribution: Jacuí River basin, Brazil (Schaefer, 2003c).
Hisonotus leptochilus Cope, 1894
Hisonotus leptochilus Cope, 1894: 96, pl. 8 [sic, 7] (figs. 11 a–b). Type locality: Rio Grande do Sul, Brazil. Holotype: ANSP 21564.
Distribution: Jacuí River basin, Brazil (Schaefer, 2003c).
Hisonotus leucofrenatus (Miranda Ribeiro, 1908)
Otocinclus leucofrenatus Miranda Ribeiro, 1908b: [2]. Type locality: Rio das Pedras [Brazil]. Type(s): at MNRJ,
current whereabouts unknown.
Distribution: Ribeira de Iguape River basin, Brazil (Schaefer, 2003c).
Hisonotus maculipinnis (Regan, 1912)
Otocinclus maculipinnis Regan, 1912b: 668, pl. 77 (fig. 3). Type locality: Rio de la Plata, South America. Syntypes:
BMNH 1909.4.2.19–22 (4).
Distribution: Paraná/Paraguay and La Plata River basins, Argentina (Schaefer, 2003c).
Hisonotus nigricauda (Boulenger, 1891)
Otocinclus nigricauda Boulenger, 1891: 234, pl. 25 (fig. 3). Type locality: Rio Grande do Sul, Brazil. Syntypes:
BMNH 1891.3.16.53–62.
Distribution: Paraná/Paraguay River basin, Brazil (Schaefer, 2003c).
Hisonotus notatus Eigenmann & Eigenmann, 1889
Hisonotus notatus Eigenmann & Eigenmann, 1889b: 41. Type locality: Santa Cruz and Juiz de Fora [Brazil]. Syntypes (96): ANSP 166924 (1), BMNH 1904.1.28.13–16 (4), MCZ 7764 (91), MCZ 8177 (1).

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Remarks: Eigenmann & Eigenmann (1890: 391) included the name minutus [interpreted in context as Hisonotus
minutus] in the discussion of H. notatus. This has been interpreted as a lapsus for H. notatus by Isbrücker (1980:84) and Schaefer (1996b: 1034) although not so noted in the printed corrigenda associated with the volume. The
name is not associated with an indication and not treated as valid elsewhere in the volume, so there is no reason
to consider this as an available name.
Hisonotus paulinus (Regan, 1908)
Otocinclus paulinus Regan, 1908f: 799, fig. 208. Type locality: Rio Piracicaba, San Paulo, Brazil. Holotype: BMNH
1907.7.6.9.
Distribution: Tietê River basin, Brazil (Schaefer, 2003c).
Hisonotus ringueleti Aquino, Schaefer & Miquelarena, 2001
Hisonotus ringueleti Aquino, Schaefer & Miquelarena, 2001: 3, fig. 1. Type locality: Uruguay, Rivera State, upper
Uruguay River drainage, Quaraí River drainage, creek at Km 18 of route joining Santa do Livramento, Brazil,
and Rivera, Uruguay; close to border (ca. 31º00'S, 55º30'W). Holotype: ILPLA 886.
Distribution: Upper Uruguay River basin, Uruguay (Schaefer, 2003c).
Hisonotus taimensis (Buckup, 1981)
Microlepidogaster taimensis Buckup, 1981: 22, figs. 1–7. Type locality: Novo canal do Arroio Taim, Estação
Ecológica do Taim, Rio Grande do Sul, Brazil. Holotype: MZUSP 14133.
Distribution: Mirim Lagoon drainage, Brazil (Schaefer, 2003c).
HOPLIANCISTRUS Isbrücker & Nijssen, 1989
Hopliancistrus Isbrücker & Nijssen, 1989: 543. Type species: Hopliancistrus tricornis Isbrücker & Nijssen, 1989.
Type by original designation. Gender: Masculine.
Hopliancistrus tricornis Isbrücker & Nijssen, 1989
Hopliancistrus tricornis Isbrücker & Nijssen, 1989: 543, fig. 6. Type locality: Brasilien, Est. Pará, Poça de Pedra no
Rio Tapajós, São Luis. Holotype: MZUSP 22007.
Distribution: Tapajós and Xingu River basins, Brazil (Fisch-Muller, 2003).
HYPANCISTRUS Isbrücker & Nijssen, 1991
Hypancistrus Isbrücker & Nijssen, 1991: 347. Type species: Hypancistrus zebra Isbrücker & Nijssen, 1991. Type byoriginal designation. Gender: Masculine.
Monophyly: Armbruster (2002).
Hypancistrus inspector Armbruster, 2002
Hypancistrus inspector Armbruster, 2002: 87, fig. 2 (middle). Type locality: Venezuela, Amazonas, Dpto. Casiquiare, río Casiquiare, río Negro-río Amazonas drainage, approximatively 10 river km above the río Negro (below
Solano), 1º58'N, 67º05'W. Holotype: MCNG 12133.
Distribution: Upper Negro, Casiquiare, and upper Orinoco River basin, Venezuela (Fisch-Muller, 2003).
Hypancistrus zebra Isbrücker & Nijssen, 1991
Hypancistrus zebra Isbrücker & Nijssen, 1991: 348, fig. 1 (middle). Type locality: Brazil, Est. Pará, anastomoses of
Rio Xingú, about one hour upstream of Altamira by speedboat (Altamira: 03º13'S 53º15'W). Holotype: MZUSP
41668.
Distribution: Xingu River basin, Brazil (Fisch-Muller, 2003).
HYPOPTOPOMA Günther, 1868
Hypoptopoma Günther, 1868a: 477. Type species: Hypoptopoma thoracatum Günther, 1868. Type by monotypy.
Gender: Neuter.
Aristommata Holmberg, 1893a: 96. Type species: Aristommata inexpectata Holmberg, 1893. Type by monotypy.
Gender: Feminine.
Diapeltoplites Fowler, 1915a: 237. Type species: Hypoptopoma gulare Cope, 1878. Type by original designation.
Gender: Masculine. Originally proposed as a subgenus of Hypoptopoma.

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Hypoptopoma bilobatum Cope, 1870
Hypoptopoma bilobatum Cope, 1870b: 566. Type locality: Pebas, Ecuador [now Peru]. Syntypes: ANSP 8280 (2).
Distribution: Amazon River basin, Peru (Schaefer, 2003c).
Hypoptopoma guianense Boeseman, 1974
Hypoptopoma guianense Boeseman, 1974: 259, fig. 1; pl. 1. Type locality: Left tributary of the Nickerie River, a few
km upstream from the Stondansi Falls, Surinam. Holotype: RMNH 26919.
Distribution: Nickerie River basin, Suriname (Schaefer, 2003c).
Hypoptopoma gulare Cope, 1878
Hypoptopoma gulare Cope, 1878: 679. Type locality: Peruvian Amazon. Holotype: ANSP 21477.
Distribution: Amazon River basin (Schaefer, 2003c).
Hypoptopoma inexspectatum (Holmberg, 1893)
Aristommata inexpectata Holmberg, 1893a: 96. Type locality: Río Paraguay, Formosa Prov., Argentina. Holotype:
MACN 5164. Also in Holmberg (1893b: 354).
Hypoptopoma guentheri Boulenger, 1895b: 526. Type locality: Descalvados, Mato Grosso [Brazil]. Syntypes:
BMNH 1895.5.17.77–82 (6).
Distribution: Paraná/Paraguay River basin (Schaefer, 2003c).
Remarks: Redescribed in Aquino & Miquelarena (2001), as Hypoptopoma inexpectata.
Hypoptopoma joberti (Vaillant, 1880)
Otocinclus joberti Vaillant, 1880a: 147. Type locality: Calderon, Brésil. Holotype: MNHN a-1966.
Distribution: Amazon River basin, Brazil (Schaefer, 2003c).
Hypoptopoma psilogaster Fowler, 1915
Hypoptopoma psilogaster Fowler, 1915a: 235, fig. 9. Type locality: Peruvian Amazon. Holotype: ANSP 21922.
Distribution: Amazon River basin, Colombia and Peru (Schaefer, 2003c).
Hypoptopoma steindachneri Boulenger, 1895
Hypoptopoma steindachneri Boulenger, 1895b: 526. Type locality: Amazonenstrome zunächst der mündung des Rio
Negro. Holotype: NMW 46272; holotype illustrated in Schaefer (1996b:1033, fig. 1). Based on specimen
described and illustrated in Steindachner (1879e: 47, pl. 7, fig. 1) as Hypoptopoma thoracatum.
Distribution: Amazon River basin (Schaefer, 2003c).
Hypoptopoma thoracatum Günther, 1868
Hypoptopoma thoracatum Günther, 1868a: 477. Type locality: Xeberos [upper Amazon River, Peru]. Holotype:
BMNH 1867.6.13.38. Illustrated and described in more detail in Günther (1868b: 234, fig. 2).
Distribution: Amazon River basin (Schaefer, 2003c).
HYPOSTOMUS La Cepède, 1803
Hypostomus La Cepède, 1803: 145. Type species: Hypostomus guacari La Cepède, 1803. Type by monotypy. Gender: Masculine.
Cochliodon Heckel, in Kner, 1854: 265. Type species: Hypostomus cochliodon Kner, 1854. Type by monotypy. Gender: Masculine.
Cheiridodus Eigenmann, 1922b: 70. Type species: Plecostomus hondae Regan, 1912. Type by original designation.
Gender: Masculine.
Watawata Isbrücker & Michels, in Isbrücker et al., 2001: 21. Type species: Hypostomus microstomus Weber, 1987.
Type by original designation. Gender: Feminine.
Remarks: Plecostomus Gronovius (1763: 127) was published in a work rejected by the ICZN (Opinion 261) for
nomenclatural purposes; the name appears on the Official Index of Rejected and Invalid Generic Names In
Zoology (Melville & Smith, 1987). Subsequent uses of Plecostomus were summarized in Isbrücker (1980). The
recently proposed genus Watawata Isbrücker & Michels was considered by Weber (2003) to be invalid.
Revision: Armbruster (2003a, Hypostomus cochliodon group).
Reviews: Boeseman (1968, Suriname), Weber (1986, Paraguay), Reis et al. (1990, Rio Grande do Sul), López &Miquelarena (1991a, b, Argentina); Mazzoli et al. (1994, Paraiba do Sul, Brazil), and Le Bail et al. (2000,
French Guiana).

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Molecular phylogeny: Montoya-Burgos et al. (2002, mitochondrial D-loop sequences).
Historical biogeography: Montoya-Burgos (2003).
Hypostomus affinis (Steindachner, 1877)
Plecostomus affinis Steindachner, 1877b: 685. Type locality: aus dem Rio Mucury bei Santa Clara [...] im Rio
Parahyba und dessen Nebenflusse Muriahé, im Rio Mucury und S. Antonio bei S. Antonio de Ferros [restricted
to Rio Parahyba (as Campos, in the rio Paraíba do Sul basin), Brazil, by lectotype designation ]. Lectotype:
NMW 44041:1; designated by Mazzoni et al. (1994: 5 and 7).
Distribution: Paraíba do Sul River basin, Brazil (Weber, 2003).
Hypostomus agna (Miranda Ribeiro, 1907)
Plecostomus ãgnã Miranda Ribeiro, 1907a: 188. Type locality: Rio da Ribeira: da Iporanga —S. Paulo [Brazil].
Holotype: at MNRJ.
Distribution: Ribeira de Iguapé River basin, Brazil (Weber, 2003; Oyakawa et al., 2005).
Hypostomus alatus Castelanau, 1855
Hypostomus alatus Castelnau, 1855: 41, pl. 20 (fig. 1). Type locality: rio Sabara, dans la province de Minas Geraës
(Brésil). Holotype: MNHN a-9441.
Distribution: Upper São Francisco River basin, Brazil (Weber, 2003).
Hypostomus albopunctatus (Regan, 1908)
Plecostomus albopunctatus Regan, 1908f: 797, pl. 49 (fig. 1). Type locality: Rio Piracicaba, Sao Paulo, Brazil. Lectotype: BMNH 1907.7.6.15; designated by Weber (1987: 279).
Distribution: Upper Paraná River basin, (Weber, 2003).
Hypostomus ancistroides (Ihering, 1911)
Plecostomus ancistroides Ihering, 1911: 396. Type locality: Rio Tatuhy, affluente do lado esquerdo do Rio Sorocaba,
Rio Piracicaba, Est. de S. Paulo [Brazil]. Syntypes: Whereabouts unknown, not found at MZUSP (Britski, 1969:
200).
Distribution: Tietê River basin, Brazil (Weber, 2003); Ribeira de Iguape River basin (Oyakawa et al., 2005).
Hypostomus angipinnatus (Leege, 1922)
Plecostomus angipinnatus Leege, 1922: 152, pl. 9 (fig. 1). Type locality: Mato Grosso (?) [Brazil]. Holotype: atPhyletisch Museum, Jena (not found, Weber, 2003).
Distribution: La Plata drainage: possibly Paraguay basin (Weber, 2003).
Hypostomus argus (Fowler, 1943)
Plecostomus argus Fowler, 1943a: 249, figs. 28–31. Type locality: Villavicencio, Río Meta basin, Colombia. Holotype: ANSP 70510.
Distribution: Upper Meta River basin, Colombia (Weber, 2003).
Hypostomus asperatus Castelnau, 1855
Hypostomus asperatus Castelnau, 1855: 41, pl. 20 (fig. 2). Type locality: de l’Araguay [Brazil]. Holotype: MNHN a9442.
Distribution: Tocantins River basin, Brazil (Weber, 2003).
Hypostomus aspilogaster (Cope, 1894)
Plecostomus aspilogaster Cope, 1894: 100, pl. 7 [pl. 7 reversed with pl. 8] (fig. 14). Type locality: from the Jacuhy,
Rio Grande do Sul. Lectotype: ANSP 21781 designated by Reis et al. (1990: 733).
Distribution: Uruguay River basin and Laguna dos Patos drainage, Brazil (Weber, 2003).
Remarks: Redescribed by Reis et al. (1990: 733).
Hypostomus atropinnis (Eigenmann & Eigenmann, 1890)
Plecostomus lima atropinnis Eigenmann & Eigenmann, 1890: 399, 410. Type locality: Goiás, Brazil. Holotype:
MCZ 27265.
Distribution: Tocantins River basin, Brazil (Weber, 2003).
Hypostomus auroguttatus Kner, 1854
Hypostomus auroguttatus Kner, 1854: 269, pl. 2 (fig. 3). Type locality: [Not explicitly given, apparently Rio
Parahyba, Brazil (Weber, 2003)]. Syntypes: at NMW. Originally proposed as Hyp. auroguttatus.
Plecostomus Lütkeni Steindachner, 1877c: 217. Type locality: Im Rio Parahyba, R. Mucuri, R. San Antonio, R.

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Quenda, R. de Pedral [restricted to Rio Parahyba (= rio Paraíba do Sul), by lectotype designation]. Lectotype:
NMW 44196: 1; designated by Mazzoni et al. (1994: 10).
Distribution: Paraíba do Sul River basin, Brazil (Weber, 2003).
Hypostomus bolivianus (Pearson, 1924)
Plecostomus bolivianus Pearson, 1924: 21, pl. 4 (fig. 1). Type locality: Popoi River, Upper Beni, Bolivia. Holotype:
CAS 77347.
Distribution: Beni River basin, Bolivia (Weber, 2003).
Hypostomus borellii (Boulenger, 1897)
Plecostomus borellii Boulenger, 1897c: 2. Type locality: Mission de San Francisco [Pilcomayo River, Bolivia]. Syntypes: BMNH 1897.1.27.19 (1), MZUT 1393 (1).
Distribution: Upper and middle Pilcomayo River basin, (Weber, 2003).
Hypostomus boulengeri (Eigenmann & Kennedy, 1903)
Plecostomus boulengeri Eigenmann & Kennedy, 1903: 502. Type locality: Matto Grosso or Asuncion. Holotype:
CAS 58554.
Distribution: Paraguay River basin (Weber, 2003).
Remarks: Redescription in Weber (1986: 982).
Hypostomus brevicauda (Günther, 1864)
Plecostomus brevicauda Günther, 1864: 235. Type locality: Bahia [Brazil]. Syntypes: BMNH 1864.1.19.16–17 (2).
Distribution: coastal drainages of Eastern Brazilian (Weber, 2003).
Hypostomus brevis (Nichols, 1919)
Plecostomus brevis Nichols, 1919a: 413, 423. Type locality: no Estado de S. Paulo [...] in the state of São Paulo[Brazil]. Holotype: AMNH 7150.
Distribution: Upper Paraná River basin or eastern Brazilian coastal drainage; known only from holotype (Weber,2003).
Hypostomus carinatus (Steindachner, 1881)
Plecostomus carinatus Steindachner, 1881c: 108, pl. 4 (fig. 2). Type locality: aus dem Amazonen-Strome, ohne
nähere Angabe des Fundortes; [...], von Jatuarana [=probably in the Parintins, near Barreirinha], Ueranduba[=lago Iranduba], und aus dem See Saraca [=lago de Sarac]. Syntypes: ?MCZ 27266 (1), NMW 44094 (1),
NMW 44096 (1).
Distribution: Middle Amazon River basin, Brazil (Weber, 2003).
Remarks: Some diagnostic characters in Rapp Py-Daniel (1988: 17 and tab. 3).
Hypostomus carvalhoi (Miranda Ribeiro, 1937)
Plecostomus carvalhoi Miranda Ribeiro, 1937: 54. Type locality: Rio Granjeiro— Crato— Ceará. Syntypes: MNRJ
924 (4, of which only 2 were reportedly found by Weber, 2003).
Distribution: Jaguaribe River basin, Brazil (Weber, 2003).
Remarks: Lectotype designation of specimen as MNRJ 924A by Miranda Ribeiro (1953: 401) considered not toclearly refer to a single specimen and therefore not valid by Weber (2003: 357).
Hypostomus cochliodon Kner, 1854
Hypostomus cochliodon Kner, 1854: 265, pl. 2 (fig. 1). Type locality: Rio Cuyaba. Syntypes: NMW 44101 (1),
NMW 46277 (1), NMW 59395 (1), RMNH D1897 (1).
Distribution: Paraguay and middle Paraná River basins (Weber, 2003).
Remarks: The names Loricaria melanoptera and Cochliodon hypostomus were published by Kner (1854: 265) assynonyms of H. cochliodon and, therefore, not available. Species redescribed in Armbruster (2003a), but see
Hollanda Carvalho & Weber (2004: 972) for comments on the identity of Armbruster’s material.
Hypostomus commersoni Valenciennes, 1836
Hypostomus Commersoni Valenciennes, 1836, in Valenciennes, 1835–47: pl. 7 (fig. 2). Type locality: not stated[considered to have come from La Plata River, Uruguay]. Holotype or lectotype: MNHN a-9444. Name madeavailable from caption to plate, with illustrated specimen the holotype (if recognizable; see Remarks).
Described in Cuvier & Valenciennes (1840b: 495, 366 of Strasbourg deluxe edition).
Plecostomus spiniger Hensel, 1870: 73. Type locality: Rio Cadea [=Cadeia River, Rio Grande do Sul, Brazil]. Holo-

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type: ZMB 7444.
Plecostomus limosus Eigenmann & Eigenmann, 1888b: 167. Type locality: Rio Grande do Sul [Brazil]. Lectotype:
MCZ 7869, designated by Reis et al. (1990: 737). Name made available by diagnostic features in key on p. 168.
Distribution: Middle and lower Paraná and Uruguay River basins; Laguna dos Patos basin (Weber, 2003).
Remarks: Weber (1986) selected MNHN a-9444 as a lectotype for Hypostomus commersoni. If this specimen was
the specimen illustrated in Valenciennes (1836), it is actually the holotype, and no lectotype would have beennecessary. However, if the illustrated specimen could not be identified, Weber’s lectotype designation would
stand. Type locality of Hypostomus commersoni discussed in Weber (1986: 994).
Hypostomus coppenamensis Boeseman, 1969
Hypostomus coppenamensis Boeseman, 1969: 120, fig. 3. Type locality: tributary of Left Coppename River, 3º54'N,
56º46'W, Surinam. Holotype: ZMA 105856.
Distribution: Upper Coppename River basin, Suriname (Weber, 2003).
Hypostomus corantijni Boeseman, 1968
Hypostomus corantijni Boeseman, 1968: 40, pl. 4 (fig. 1). Type locality: Sipaliwini River, Surinam. Holotype:
RMNH 25471 (largest specimen, if not separated).
Distribution: Corantijn River basin, upstream first cataracts, Suriname (Weber, 2003).
Hypostomus crassicauda Boeseman, 1968
Hypostomus crassicauda Boeseman, 1968: 42, pl. 17 (fig. 1). Type locality: Sipaliwini River, Surinam. Holotype:
RMNH 25489.
Distribution: Upper Sipaliwini River basin, Suriname (Weber, 2003).
Hypostomus derbyi (Haseman, 1911)
Plecostomus derbyi Haseman, 1911b: 384, pl. 79. Type locality: Porto União da Victoria [Iguaçu River basin, Brazil]. Holotype: FMNH 54246.
Distribution: Iguaçu River and Uruga-í Creek basins, Paraguayan tributaries of Paraná River, (Weber, 2003).
Remarks: Redescribed in Gomez et al. (1990) from Argentina.
Hypostomus dlouhyi Weber, 1985
Hypostomus dlouhyi Weber, 1985: 956, fig. 1 (middle). Type locality: système du Rio Paraná, sur un bras du lac deretenue du Rio Yguazú [non Iguaçu, Brésil] à la hauteur de la localité Juan E. O’Leary, 55º20' Ouest, 25º25'30"
Sud, Paraguay, dèp. Alto Parana. Holotype: MHNG 2229.43.
Distribution: Yguazú River basin in middle Paraná River drainage, Paraguay (Weber, 2003).
Hypostomus eptingi (Fowler, 1941)
Chaetostomus eptingi Fowler, 1941a: 158, figs. 66–68. Type locality: Forteleza, Ceará [Brazil]. Holotype: ANSP
69447.
Distribution: Coastal drainages of northern Brazil (Weber, 2003).
Hypostomus ericae Hollanda Carvalho & Weber, 2004
Hypostomus ericae Hollanda Carvalho & Weber, 2004: 955, fig. 5. Type locality: Brazil: Goiás: Tocantins Riverbasin: Rio Tocantins, Porto Alfredinho, 14º02' 22.1"S, 48º31'28.6"W. Holotype: MNRJ 27861.
Distribution: Upper Tocantins River basin, Brazil (Hollanda Carvalho & Weber, 2004).
Remarks: Included in Hypostomus cochliodon group by Hollanda Carvalho & Weber (2004).
Hypostomus ericius Armbruster, 2003
Hypostomus ericius Armbruster, 2003a: 25, fig. 9. Type locality: Peru. Amazonas: Río Marañon dr. 1 km upstream
from Caterpiza. Holotype: MUSM 17441.
Distribution: Maranon, Itaya, and Nanay Rivers, Upper Amazon River basin, Peru (Armbruster, 2003a).
Remarks: Included in the Hypostomus cochliodon species group by Armbruster (2003a).
Hypostomus fluviatilis (Schubart, 1964)
Plecostomus fluviatilis Schubart, 1964b: 7, figs. 1–4. Type locality: Corrego da Lazica, perto de Ouro Fino (MG);
CachoeXra do Espraiado, no alto Mogi Guaçu, acima do Soledade, Ouro Fino (MG); Alto Mogi Guaçu, 5 km
jusante de Inconfidentes (mun. Ouro Fino, MG). Syntypes: Whereabouts unknown, not found (Weber, 2003).
Distribution: Grande River basin, Brazil (Weber, 2003).
Hypostomus fonchii Weber & Montoya-Burgos, 2002

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Hypostomus fonchii Weber & Montoya-Burgos, 2002: 357, fig. 3 (b). Type locality: Peru: Ucayali: Quebrada John,
lower part, near mouth in the Rio Pauya, in shallow water with emerging rocks, 06º 36' 7.0"S/75º56'26.3"W(GPS), alt. 360 m., Rio Cushabatay drainage, Rio Ucayali basin (Station 1). Holotype: MHNG 2613.66.
Distribution: Cushabatay River basin, Peru (Weber, 2003).
Hypostomus francisci (Lütken, 1874)
Plecostomus francisci Lütken, 1874c: 30. Type locality: in flumine Sti. Francisci [=São Francisco River, Brazil].
Holotype: ZMUC P 30171. Originally proposed as Pl. francisci.
Distribution: Upper São Francisco River basin, Brazil (Weber, 2003).
Hypostomus garmani (Regan, 1904)
Plecostomus garmani Regan, 1904b: 214, pl. 10 (fig. 1). Type locality: Rio das Velhas, Eastern Brazil. Holotype:
BMNH 1904.1.28.3.
Distribution: São Francisco River basin, Brazil (Weber, 2003).
Hypostomus goyazensis (Regan, 1908)
Plecostomus goyazensis Regan, 1908f: 798, fig. 207(a). Type locality: Goyaz [Brazil]. Holotype: BMNH
1889.11.14.49.
Distribution: Upper Araguaia River basin, Brazil (Weber, 2003).
Hypostomus gymnorhynchus (Norman, 1926)
Plecostomus gymnorhynchus Norman, 1926b: 95. Type locality: Iponcin Creek, into Approuague River, French Guiana. Holotype: BMNH 1926.3.2.745.
Distribution: Guianan coastal drainages, from Oyapock westward to Maroni basin, French Guiana (Weber, 2003).
Remarks: Redescribed in Le Bail et al. (2000: 254).
Hypostomus hemicochliodon Armbruster, 2003
Hypostomus hemicochliodon Armbruster, 2003a: 29, fig. 12. Type locality: Peru. Amazonas: Río Nanay, Río Amazonas dr. at Pampa Chica, 4.54 km from Iquitos center, 269º bearing, 03º45'07"S, 73º16'59"W. Holotype:
MUSM 17442.
Distribution: Marañon, Itaya, and Nanay, Orosa and Napo Rivers, Upper Amazon River basin, Negro, Tapajos and
Xingu Rivers of Brazil, and Orinoco River basin, (Armbruster, 2003a).
Remarks: Included in the Hypostomus cochliodon species group by Armbruster (2003a).
Hypostomus hemiurus (Eigenmann, 1912)
Plecostomus hemiurus Eigenmann, 1912b: 224, fig. 34, pl. 25 (fig. 1). Type locality: Amatuk [Potaro River], British
Guiana. Holotype: FMNH 53110.
Distribution: Coastal drainages of Guyana (Weber, 2003).
Remarks: Appeared first as name only in Eigenmann (1910: 407).
Hypostomus hermanni (Ihering, 1905)
Plecostomus Hermanni Ihering, 1905: 560. Type locality: Rio Piracicaba, São Paulo, Brazil. Holotype: BMNH
1905.6.9.5.
Distribution: Tietê River basin, Brazil (Weber, 2003).
Hypostomus hondae (Regan, 1912)
Plecostomus hondae Regan, 1912b: 666, pl. 76 (fig. 3). Type locality: Honda, Colombia (300–400 ft.). Syntypes:
BMNH 1909.7.23.43–44 (2).
Cochliodon pospisili Schultz, 1944c: 312, pl. 11 (figs. c–d). Type locality: Rio Palmar near Totuma, about 100 km.
southwest of Maracaibo, Venezuela. Holotype: USNM 121003.
Distribution: Magdalena River and Lake Maracaibo basins (Weber, 2003); also Sinú and Atrato basins (Armbruster,
2003a).
Remarks: Redescribed in Lilyestrom (1984) and Armbruster (2003a); included by Armbruster in the Hypostomus
cochliodon species group.
Hypostomus hoplonites Rapp Py-Daniel, 1988
Hypostomus hoplonites Rapp Py-Daniel, 1988: 14, fig. 2(6). Type locality: Brasil: Amazon (Rio Solimões, complexo de lagos do Janauacá). Holotype: INPA 109.2.
Distribution: Middle Amazon basin, Brazil (Weber, 2003).

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Hypostomus iheringii (Regan, 1908)
Plecostomus iheringii Regan, 1908f: 795, pl. 47 (fig. 1). Type locality: Rio Piracicaba, San Paulo, Brazil. Holotype:
BMNH 1907.7.6.13.
Distribution: Tietê River basin, Brazil (Weber, 2003).
Hypostomus interruptus (Miranda Ribeiro, 1918)
Plecostomus interruptus Miranda Ribeiro, 1918d: 632. Type locality: Rio Juquiá [Brazil]. Holotype: MZUSP 2110.
Distribution: Ribeira de Iguape River basin, Brazil (Weber, 2003; Oyakawa et al., 2005).
Remarks: Type locality interpreted by Britski (1969: 209) as rio Juquiá, Poço Grande, Estado de São Paulo, Brazil.
Hypostomus isbrueckeri Reis, Weber & Malabarba, 1990
Hypostomus isbrueckeri Reis, Weber & Malabarba, 1990: 752, fig. 22. Type locality: Rio Conceição, Ijui, Rio
Grande do Sul, Brazil. Holotype: MCP 10488.
Distribution: Middle and upper Uruguay River basin, Brazil (Weber, 2003).
Hypostomus jaguribensis (Fowler, 1915)
Plecostomus jaguribensis Fowler, 1915b: 264, fig. 3. Type locality: Rio Jaguaribé at Barro Alto, Brazil. Holotype:
ANSP 39930.
Distribution: Jaguaribe River basin, Brazil (Weber, 2003).
Hypostomus johnii (Steindachner, 1877)
Plecostomus Johnii Steindachner, 1877b: 691. Type locality: Rio Puty und Rio Preto [Brazil]. Syntypes: MCZ 7831
(1), MCZ 7863 (4), MCZ 7864 (2), NMW 44191 (2), NMW 44192 (2), NMW 44193 (2).
Distribution: Parnaíba and São Francisco River basins (Weber, 2003).
Hypostomus laplatae (Eigenmann, 1907)
Plecostomus laplatae Eigenmann, 1907: 450, pl. 21 (fig. 1). Type locality: Buenos Aires [Argentina]. Holotype: at
Princeton Univ.
Plecostomus taeniatus Regan, 1908d: 358. Type locality: Río La Plata, Argentina. Syntypes: BMNH 1908.8.29.17(1), ZMB 16817 (1).
Plecostomus rachovii Regan, 1913e: 555. Type locality: Near Rio Janeiro [Probably incorrect (Weber, 2003)]. Holotype: BMNH 1913.10.30.15.
Plecostomus commersonoides Marini, Nichols & La Monte, 1933: 3. Type locality: Dársena Norte, Buenos Aires,
Argentina. Holotype: AMNH 12243.
Distribution: La Plata River basin (Weber, 2003).
Remarks: Redescribed in López & Miquelarena (1991a: 28).
Hypostomus latifrons Weber, 1986
Hypostomus latifrons Weber, 1986: 991, fig. 6 (middle). Type locality: Río Aguaray-guazú, bassin du río Paraguay
système du rio Paraná, Transchaco 117 km 58º00'00" Ouest, 24º22'50" Sud, dép. Presidente Hayes, Paraguay.
Holotype: MHNG 2256.67.
Distribution: Paraguay River basin, Paraguay (Weber, 2003).
Hypostomus latirostris (Regan, 1904)
Plecostomus latirostris Regan, 1904b: 213, pl. 11 (fig. 1). Type locality: River Jungada, Matto Grosso ... and from
Goyaz [Brazil]. Syntypes (3): BMNH 1892.4.20.26–27 (2).
Distribution: Paraguay River basin (Weber, 2003).
Hypostomus levis (Pearson, 1924)
Rhinelepis levis Pearson, 1924: 22, pl. 5 (fig. 1). Type locality: Huachi, altitude 2,235 feet. Huachi overlooks thejunction of the Bopi and Cochabamba rivers which here form the Beni, Bolivia. Holotype: CAS 77349.
Distribution: Cochabamba and Popoi Rivers, upper Beni River basin, Madeira River drainage of Bolivia (Armbruster, 2003a).
Remarks: Redescribed and placed in the Hypostomus cochliodon species group by Armbruster (2003a).
Hypostomus lexi (Ihering, 1911)
Plecostomus lexi Ihering, 1911: 387. Type locality: Rio Pardo, perto de Barretos (Est. S. Paulo) quasi na foz do rioque é affluente do Rio Grande, por sua vez tributario do lado esquerdo do Rio Paraná [Brazil]. Holotype:
MZUSP 2126.

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Distribution: Grande River basin, Brazil (Weber, 2003).
Hypostomus lima (Reinhardt, 1874)
Plecostomus lima Reinhardt, in Lütken, 1874c: 29. Type locality: in rivulis flumini Rio da Velhas affluentibus [as“Ribeirão do Mato” og andre smaa Bifloder til Rio d. Velhas, in Lhtken, 1875]. Syntypes: BMNH 1876.1.10.1–
2 (2), MNHN 0000-9573 (1), NMW 44194 (1), NMW 44195 (1), SMNS 2050 (1), ZMUC P 30143–ZMUC P
30148 (1 each).
Distribution: São Francisco River basin, Brazil (Weber, 2003).
Hypostomus longiradiatus (Holly, 1929)
Plecostomus longiradiatus Holly, 1929: 118. Type locality: aus dem Rio Guaná [sic, Guamá (Weber, 2003)], vermutlich einem Nebenflusse des Madeira, der selbst ein Nebenfluß des Amazonenstromes ist [...] (Brasilien).
Holotype: Whereabouts unknown.
Distribution: Guamá River basin, Brazil (Weber, 2003).
Hypostomus luteomaculatus (Devincenzi 1942)
Plecostomus luteomaculatus Devincenzi, in Devincenzi & Teague, 1942: 20, pl. 3 (fig. 3). Type locality: Río Uruguay [at Paysandú, Uruguay]. Holotype: MHNM CI 359.
Distribution: Uruguay River basin, (Weber, 2003).
Hypostomus luteus (Godoy, 1980)
Plecostomus luteus Godoy, 1980: 29, fig. 15. Type locality: Rio Pelotas, Volta Grande 2, Marcelino Ramos, RS[Brazil]. Holotype: personal collection of M. Godoy.
Distribution: Upper Uruguay River basin, Brazil (Weber, 2003).
Remarks: Redescribed by Reis et al. (1990: 741).
Hypostomus macrophthalmus Boeseman, 1968
Hypostomus pseudohemiurus macrophthalmus Boeseman, 1968: 56, pl. 10 (fig. 1). Type locality: Sipaliwini River,near air strip, Surinam. Holotype: RMNH 25514.
Distribution: Sipaliwini River basin, Suriname (Weber, 2003).
Hypostomus macrops (Eigenmann & Eigenmann, 1888)
Plecostomus macrops Eigenmann & Eigenmann, 1888b: 170. Type locality: Rio das Velhas [Brazil]. Holotype:
MCZ 7888. Name made available by diagnostic features in key on p. 170.
Distribution: Das Velhas River basin, Brazil (Weber, 2003).
Hypostomus macushi Armbruster & de Souza, 2005
Hypostomus macushi Armbruster & de Souza, 2005: 3, fig. 1. Type locality: Guyana, Rupunini (Region 9). Ireng
River, 6.9 km WW Karasabai, Takutu River – Negro River drainage, 04.01957º, – 059.60170º. Holotype: UG/
CSBD 11047.
Distribution: Lower and middle Rupununi River, Essequibo River, Takutu River and Ireng River, Guyana (Armbruster & de Souza, 2005).
Remarks: Placed into the Hypostomus cochliodon species group by Armbruster & de Souza (2005) .
Hypostomus margaritifer (Regan, 1908)
Plecostomus margaritifer Regan, 1908f: 796, pl. 47 (fig. 2). Type locality: Rio Piracicaba, San Paulo, Brazil. Holotype: BMNH 1907.7.6.14.
Plecostomus margaritifer butantanis Ihering, 1911: 394. Type locality: no Rio Pinheiros, perto de São Paulo (Capital) [Brazil]. Holotype: possibly at MZUSP, but not found.
Distribution: Upper and middle Paraná River basin, (Weber, 2003).
Hypostomus meleagris (Marini, Nichols & La Monte, 1933)
Plecostomus meleagris Marini, Nichols & La Monte, 1933: 4. Type locality: Southeastern Brazil [apparently inupper Paraná drainage]. Holotype: AMNH 12246.
Distribution: Upper and Middle Paraná River basin (Weber, 2003).
Hypostomus micromaculatus Boeseman, 1968
Hypostomus micromaculatus Boeseman, 1968: 49, 77 [addendum], pl. 7 (figs. 1–4). Type locality: Mamadam (falls),
Surinam River, Surinam. Holotype: RMNH 25487 (largest specimen).
Distribution: Upper and middle Suriname River basin, Suriname (Weber, 2003).

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Hypostomus microstomus Weber, 1987
Hypostomus microstomus Weber, 1987: 275, fig. 2 (middle). Type locality: système du río Paraná, río Alto-Paraná,Paraguay, dpt Itapua, 15 km SE Encarnacion, Campichuelo, 55º45' Ouest, 27º26' Sud. Holotype: MHNG
2367.90.
Distribution: Middle Paraná River basin (Weber, 2003).
Hypostomus mutucae Knaack, 1999
Hypostomus mutucae Knaack, 1999b: 102, fig. p. 102, Type locality: Brazil, Mato Grosso, rio Mutuca. Holotype:
MCP 28669.
Distribution: Cuiabá River basin, Brazil (Weber, 2003).
Hypostomus myersi (Gosline, 1947)
Plecostomus myersi Gosline, 1947: 116, pl. 5 (fig. 9). Type locality: Rio Iguaçu at Porto União, State of Paraná [Brazil]. Holotype: MNRJ 4251.
Distribution: Iguaçu River and Uruga-í Creek basins, Brazil (Weber, 2003); Argentina (Gomez et al., 1990).
Remarks: Redescribed in Gomez et al. (1990) from Argentina.
Hypostomus nematopterus Isbrücker & Nijssen, 1984
Hypostomus nematopterus Isbrücker & Nijssen, 1984c: 9, fig. 1 (middle). Type locality: French Guyana, Alikene
Creek, left bank tributary to Camopi River, Oyapock river system. Holotype: IRSNB 689.
Distribution: Oyapock River basin, French Guiana; known only from the types (Weber, 2003).
Hypostomus niceforoi (Fowler, 1943)
Hemiancistrus niceforoi Fowler, 1943a: 250, figs. 32–35. Type locality: Florencia, Río Orteguasa, Colombia. Holotype: ANSP 70511.
Distribution: Upper Japurá River basin, Colombia (Weber, 2003).
Hypostomus nickeriensis Boeseman, 1969
Hypostomus nickeriensis Boeseman, 1969: 125, fig. 4a. Type locality: Stondansie Falls, Nickerie River, Surinam.
Holotype: ZMA 105.765.
Distribution: Upper Nickerie River basin, Suriname (Weber, 2003).
Hypostomus niger (Marini, Nichols & La Monte, 1933)
Plecostomus niger Marini, Nichols & La Monte, 1933: 4. Type locality: Southeastern Brazil. Holotype: AMNH
12245.
Distribution: Rivers of southeastern Brazil; known only from holotype (Weber, 2003).
Hypostomus nigromaculatus (Schubart, 1964)
Plecostomus nigromaculatus Schubart, 1964b: 2, fig. 2. Type locality: Rio Mogi Guaçu, Cachoeira de Emas, naregião da corredeira (topava) (Mun. Pirassununga, Est. São Paulo) [Brazil]. Holotype: EEBP 304 (whereabouts
unknown).
Distribution: Upper and middle Paraná River basin, Brazil (Weber, 2003).
Hypostomus nudiventris (Fowler, 1941)
Plecostomus nudiventris Fowler, 1941a: 147, figs. 44–46. Type locality: Rio Choró, Ceará, near Forteleza [Brazil].
Holotype: ANSP 69402.
Distribution: Coastal drainages of northern Brazil (Weber, 2003).
Hypostomus occidentalis Boeseman, 1968
Hypostomus gymnorhynchus occidentalis Boeseman, 1968: 47, pl. 6 (figs. 1–4). Type locality: Surinam River near
Brokopondo, Surinam. Holotype: RMNH 25520.
Distribution: Suriname River basin, Suriname (Weber, 2003).
Hypostomus oculeus (Fowler, 1943)
Panaque oculeus Fowler, 1943a: 256, figs.48–51. Type locality: Florencia, Rio Orteguasa, Colombia. Holotype:
ANSP 70518.
Distribution: Marañon, Napo, and Orteguasa River basins, upper Amazon drainage (Armbruster, 2003a); upper
Japurá River basin, Colombia (Weber, 2003).
Remarks: Redescribed in Armbruster (2003a) and placed in the Hypostomus cochliodon species group.
Hypostomus pagei Armbruster, 2003

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Hypostomus pagei Armbruster, 2003a: 41, figs. 14 b, 18. Type locality: Venezuela, Yaracuy, Rio Tupe, Rio Aroa dr.
ca. 12 km N Aroa on Highway 3; 10º30'31"N, 68º52'55"W. Holotype: MCNG 24696.
Distribution: Aroa, Tocuyo and Yaracuy River basins, Caribbean drainage, Venezuela (Armbruster, 2003a).
Remarks: Included in the Hypostomus cochliodon species group by Armbruster (2003a) .
Hypostomus pantherinus Kner, 1854
Hypostomus pantherinus Kner, 1854: 267. Type locality: Rio Guaporé, ohne nähere Angaben [Brazil]. Holotype:
NMW 44206. Originally proposed as Hyp. pantherinus.
Distribution: Madeira River basin, Brazil; known only from holotype (Weber, 2003).
Hypostomus papariae (Fowler, 1941)
Plecostomus plecostomus papariae Fowler, 1941a: 145, figs. 41–43. Type locality: Lago Papary, Rio Grande doNorte [Brazil]. Holotype: ANSP 69398.
Distribution: Grande do Norte River basin, Brazil (Weber, 2003).
Hypostomus paranensis Weyenbergh, 1877
Hypostomus paranensis Weyenbergh, 1877: 9. Type locality: las lagunas al rededor de Sante Fe [Argentina]. Syntypes: Whereabouts unknown.
Plecostomus cordovae Günther, 1880a: 11. Type locality: Cordova [Argentina]. Holotype: BMNH 1878.4.4.1.
Distribution: Bermejo, Dulce, Uno, Secundo and middle Paraná River basins (Weber, 2003).
Remarks: Hypostomus paranensis Weyenbergh treated as nomen oblitum by Weber (2003), without explanation.
Hypostomus paucimaculatus Boeseman, 1968
Hypostomus paucimaculatus Boeseman, 1968: 52, pl. 8 (fig. 1). Type locality: Surinam River, near Brokopondo,
Surinam. Holotype: RMNH 25468.
Distribution: Upper and middle Suriname River basin, Suriname (Weber, 2003).
Hypostomus paucipunctatus Hollanda Carvalho & Weber, 2004
Hypostomus paucipunctatus Hollanda Carvalho & Weber, 2004: 962, fig. 7. Type locality: Brazil: Pará: municipalityof Caldeirão, Itacaiúnas River, Igarapé Pojuca, Serra dos Carajás (5º52'S, 50º32'W, coordinates of Itacaiúnas
River, near Castanhal). Holotype: MZUSP 82271.
Distribution: Itacaiúnas River basin, Pará, Brazil (Hollanda Carvalho & Weber, 2004).
Remarks: Included in Hypostomus cochliodon species group by Hollanda Carvalho & Weber (2004).
Hypostomus paulinus (Ihering, 1905)
Plecostomus paulinus Ihering, 1905: 560. Type locality: Rio Piracicaba, São Paulo, Brazil. Holotype: BMNH
1905.6.9.4.
Distribution: Tietê River basin, Brazil, Paraná River basin, Paraguay (Weber, 2003).
Hypostomus piratatu Weber, 1986
Hypostomus piratatu Weber, 1986: 987, fig. 3 (middle). Type locality: Bras intermittant du Río Tebicuary, bassin du
rio Paraguay, systeme du Río Paraná; Sapucái, dép. Paraguari, Paraguay. Holotype: MHNG 2265.03.
Distribution: Eastern Paraguay River tributaries, Paraguay (Weber, 2003).
Hypostomus plecostomoides (Eigenmann, 1919)
Cochliodon plecostomoides Eigenmann, 1919a: 133. Type locality: Quebrada Cramalote, Villavicencio [Colombia].
Holotype: CAS 82501 [lost in mail (Eschmeyer et al., 1998)]. Also described as new in Eigenmann (1922b:
225, pl. 11, figs. 1–3).
Distribution: Orinoco River basin and Tuy River and Valencia Lake basins of Venezuela (Armbruster, 2003a).
Remarks: Redescribed in Lilyestrom (1984) and Armbruster (2003a); placed by Armbruster in Hypostomus cochliodon species group.
Hypostomus plecostomus (Linnaeus, 1758)
Acipenser plecostomus Linnaeus, 1758: 238. Type locality: India [probably Suriname]. Syntypes: NRM 32 (3, of
which one is illustrated in Linnaeus (1754: pl. 28, fig. 4)), and specimen also illustrated in Gronovius (1754: 24,
pl. 3, figs. 1–2).
Hypostomus guacari La Cepède, 1803: 144, 145, pl. 4 (fig. 2). Type locality: Les rivieres de l’Amerique. Syntypes:
Based on numerous literature sources, including account of Acipenser plecostomus Linnaeus, 1758; therefore
syntypes of that species are also syntypes of this one.

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Loricaria flava Shaw, 1804: 38, pl. 101. Type locality: Indian Seas. Based on numerous literature sources, including
account of Acipenser plecostomus Linnaeus, 1758; therefore syntypes of that species are also syntypes of thisone.
Plecostomus bicirrosus Gronow, in Gray, 1854: 158. Type locality: Americes Meridionali fluminibus. Holotype or
syntype: Specimen illustrated in Gronovius (1754: 24, pl. 3, figs. 1–2).
Plecostomus brasiliensis Bleeker, 1862 (in Bleeker, 1862–63): 2. Type locality: [not stated]. Based on description of
Hypostomus plecostomus in Cuvier & Valenciennes (1840). Described in more detail in Bleeker (1864: 7), withlocalities listed as follows: Surinama, Mejico, Cujaba, Chili. Boeseman (1968: 38) designated a lectotype,
RMNH 3102, but its validity needs to be clarified, as it was apparently based on the description in Bleeker(1864) and not the earlier paper.
Distribution: Coastal drainages of the Guianas (Weber, 2003).
Remarks: Proposed neotype designations for Acipenser plecostomus and Hypostomus guacari (RMNH 18240 for
both names) in Boeseman (1968: 11) are invalid inasmuch as the NRM syntypes are still extant.
Hypostomus pseudohemiurus Boeseman, 1968
Hypostomus pseudohemiurus Boeseman, 1968: 54, pl. 9 (fig. 1). Type locality: Kabalebo River, Corantijn Riverbasin, Surinam. Holotype: RMNH 25516 (largest specimen).
Distribution: Corantijn River basin, Suriname (Weber, 2003).
Hypostomus punctatus Valenciennes, 1840
Hypostomus punctatus Valenciennes, in Cuvier & Valenciennes, 1840b: 493 (364 in Strasbourg deluxe edition).
Type locality: Rio-Janéiro. Holotype: at MNHN, but not found by Weber (2003).
Distribution: coastal drainages of southeastern Brazil (Weber, 2003).
Hypostomus pusarum (Starks, 1913)
Plecostomus pusarum Starks, 1913: 36, pl. 6. Type locality: at Ceará Mirim [Brazil]. Holotype: SU 22225.
Distribution: coastal drainages of northern Brazil (Weber, 2003).
Hypostomus pyrineusi (Miranda Ribeiro, 1920)
Cochliodon pyrineusi Miranda Ribeiro, 1920: 9, pls. 3& 4. Type locality: provavelmente Jamary [Brazil]. Holotype:
MNRJ 863.
Distribution: Madeira, Napo, Ucayali, Marañon and upper Amazon River basins (Armbruster, 2003a).
Remarks: Redescribed and included in Hypostomus cochliodon species group in Armbruster (2003a).
Hypostomus regani (Ihering, 1905)
Plecostomus regani Ihering, 1905: 558. Type locality: Rio Piracicaba, São Paulo, Brazil. Lectotype: BMNH
1905.6.7.2, designated by Reis et al. (1990: 745).
Distribution: Paraná, Paraguay and Uruguay River basins (Weber, 2003).
Remarks: Redescribed by Reis et al. (1990: 745).
Hypostomus robinii Valenciennes, 1840
Hypostomus Robinii Valenciennes, in Cuvier & Valenciennes, 1840b: 501 (370 in Strasbourg deluxe edition). Type
locality: de la Trinité [...] des affluens da la Plata [restricted to La Trinité by lectotype designation]. Lectotype:
MNHN a-9569, designated by Boeseman (1968: 36).
Distribution: Trinidad Island, Trinidad and Tobago (Weber, 2003).
Hypostomus rondoni (Miranda Ribeiro, 1912)
Plecostomus rondoni Miranda Ribeiro, 1912: 6. Type locality: S. Manoel— Rio Tapajós [Brazil]. Holotype: MNRJ
741. Specific name spelled Plecostomus rondini in heading of account, but corrected in attached printed corrigendum.
Distribution: Tapajós River basin, Brazil (Weber, 2003).
Hypostomus roseopunctatus Reis, Weber & Malabarba, 1990
Hypostomus roseopunctatus Reis, Weber & Malabarba, 1990: 756, figs. 2, 26. Type locality: Rio Pelotas at roadfrom Esmeralda to Anita Garibaldi, Rio Grande do Sul, Brazil. Holotype: MCP 12239.
Distribution: Uruguay River basin, (Weber, 2003).
Hypostomus saramaccensis Boeseman, 1968
Hypostomus saramaccensis Boeseman, 1968: 58, pl. 2 (fig. 1). Type locality: Feddiprati (rapids), middle Saramacca

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River, Surinam. Holotype: RMNH 25488 (largest).
Distribution: Coastal drainages of Suriname (Weber, 2003).
Hypostomus scabriceps (Eigenmann & Eigenmann, 1888)
Plecostomus commersonii scabriceps Eigenmann & Eigenmann, 1888b: 168. Type locality: Sao Matheos [Brazil].
Syntypes: MCZ 7894 (2), BMNH 1904.1.28.2 (1). Name made available from characters in key on p. 168.
Distribution: São Mateus River basin, Brazil (Weber, 2003).
Hypostomus scaphyceps (Nichols, 1919)
Plecostomus scaphyceps Nichols, 1919a: 425. Type locality: Cerqueira Cezar, Estado de São Paulo [Brazil]. Holotype: AMNH 7152. Name spelled Plecostomus scaplyceps on p. 416 [in Portuguese portion of original description].
Distribution: Paranapanema River basin, Brazil (Weber, 2003).
Remarks: Fowler (1954: 194) acted as first reviser and selected Plecostomus scaphyceps as the valid name for thisspecies.
Hypostomus sculpodon Armbruster, 2003
Hypostomus sculpodon Armbruster, 2003a: 49, fig. 21. Type locality: Venezuela, Amazonas, Río Casiquiare, Río
Negro dr., Departmento Río Negro, at Piedra Carguana (at second camp, left descending bank of river in portionof river between mouths of Río Siapa and Río Pasimoni), 01º58.611'N, 66º32.359'W. Holotype: MCNG 37042.
Distribution: Upper Negro and upper Orinoco River basins, Venezuela (Armbruster, 2003a).
Remarks: Included in the Hypostomus cochliodon species group by Armbruster (2003a).
Hypostomus seminudus (Eigenmann & Eigenmann, 1888)
Plecostomus seminudus Eigenmann & Eigenmann, 1888b: 169. Type locality: Brazil (?). Holotype: MCZ 28981.
Name available from description in key on p. 169.
Distribution: Probably Brazil; known only from holotype (Weber, 2003).
Hypostomus simios Hollanda Carvalho & Weber, 2004
Hypostomus simios Hollanda Carvalho & Weber, 2004: 967, fig. 9. Type locality: Brazil: Amapá: bridge on the roadto Serra do Navio, Cupixi river. Holotype: MZUSP 82268.
Distribution: Cupixi River basin, Amapá, Brazil (Hollanda Carvalho & Weber, 2004).
Remarks: Included in Hypostomus cochliodon species group by Hollanda Carvalho & Weber (2004).
Hypostomus sipaliwinii Boeseman, 1968
Hypostomus sipaliwinii Boeseman, 1968: 60, pl. 12 (fig. 1). Type locality: Sipaliwini River, upper Corantijn Riverbasin, Surinam. Holotype: RMNH 25481 (larger specimen).
Distribution: Upper Corantijn River basin, Suriname (Weber, 2003).
Hypostomus soniae Hollanda Carvalho & Weber, 2004
Hypostomus soniae Hollanda Carvalho & Weber, 2004: 965, fig. 8. Type locality: Brazil: Pará: temporary pools on
the river sides of the Rio Tapajós, between Vila Nova and Urua. Holotype: MZUSP 82272.
Distribution: Tapajós River basin, Pará, Brazil (Hollanda Carvalho & Weber, 2004).
Remarks: Included in Hypostomus cochliodon species group by Hollanda Carvalho & Weber (2004).
Hypostomus strigaticeps (Regan, 1908)
Plecostomus strigaticeps Regan, 1908f: 796, pl. 48 (fig. 1). Type locality: Rio Piracicaba, San Paulo, Brazil. Syntypes: BMNH 1907.7.6.10–12 (2 or 3).
Distribution: Tietê River basin, Brazil (Weber, 2003).
Hypostomus subcarinatus Castelnau, 1855
Hypostomus subcarinatus Castelnau, 1855: 42, p. 21 (fig. 1). Type locality: des rivières de la province des Mines[Brazil]. Holotype: MNHN a-9575.
Distribution: Coastal drainages of eastern Brazil, possibly including São Francisco River basin (Weber, 2003).
Hypostomus surinamensis Boeseman, 1968
Hypostomus surinamensis Boeseman, 1968: 61, pl. 13 (fig. 1). Type locality: Surinam River near Brokopondo, Surinam. Holotype: RMNH 25497 (larger specimen).
Distribution: Suriname River basin, Suriname (Weber, 2003).
Hypostomus tapanahoniensis Boeseman, 1969

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Hypostomus gymnorhynchus tapanahoniensis Boeseman, 1969: 129, figs. 5 (b, d). Type locality: Upper Tapanahoni
River, Surinam. Holotype: RMNH 25476. Originally proposed as H. gymnorhynchus tapanahoniensis.
Distribution: Maroni River basin (Weber, 2003).
Hypostomus taphorni (Lilyestrom, 1984)
Cochliodon taphorni Lilyestrom, 1984: 43, fig. 8. Type locality: Río Botanamo, cerca del puente en la vía a
Bochinche, Edo, Bolívar, Venezuela. Holotype: MCNG 8084.
Distribution: Cuyuni River basin, Essequibo drainage, Venezuela (Armbruster, 2003a), Rupununi River and Takutu
River basins, Guyana (Armbruster & de Souza, 2005).
Remarks: Redescribed and included in Hypostomus cochliodon species group in Armbruster (2003a).
Hypostomus tapijara Oyakawa, Akama & Zanata, 2005
Hypostomus tapijara Oyakawa, Akama & Zanata, 2005: 19, fig. 7. Type locality: Brazil: São Paulo, rio Catas Altas,
Ribeira. Holotype: MZUSP 55204.
Distribution: Ribeira de Iguape River basin, Brazil (Oyakawa et al., 2005).
Hypostomus ternetzi (Boulenger, 1895)
Plecostomus ternetzi Boulenger, 1895b: 525. Type locality: Paraguay. Holotype: BMNH 1895.5.17.64; holotype
illustrated in Boulenger (1896c: pl. 5).
Distribution: Middle Paraná, Paraguay, and Uruguay River basins (Weber, 2003).
Hypostomus tietensis (Ihering, 1905)
Plecostomus tietensis Ihering, 1905: 559. Type locality: Rio Tietè, São Paulo, Brazil. Holotype: BMNH 1905.6.9.1.
Distribution: Tietê River basin, Brazil (Weber, 2003).
Hypostomus topavae (Godoy, 1969)
Plecostomus topavae Godoy, 1969: 176, fig. 1. Type locality: na região da topava [...] Cachoeira de Emas, Rio Mogi
Guassu, Estado de São Paulo, Brasil. Holotype: EEBP 315a.
Distribution: Grande River basin, Brazil; known only from holotype (Weber, 2003).
Hypostomus unae (Steindachner, 1878)
Plecostomus Unae Steindachner, 1878b: 383. Type locality: Rio Una (südlich von Bahia) [Brazil]. Syntypes: BMNH
1862.11.23.12 (1), NMW 44259 (4).
Distribution: Una River basin in Bahia State, Brazil (Weber, 2003).
Remarks: This name was based on descriptions of Plecostomus Robinii (not Valenciennes) in Günther (1864: 236)
and Steindachner (1877b: 676).
Hypostomus uruguayensis Reis, Weber & Malabarba, 1990
Hypostomus uruguayensis Reis, Weber & Malabarba, 1990: 760, fig. 30. Type locality: Rio Uruguay at “Rancho da
Amizade”, São Borja, Rio Grande do Sul, Brazil. Holotype: MCP 11874.
Distribution: Uruguay River basin (Weber, 2003).
Hypostomus vaillanti (Steindachner, 1877)
Plecostomus Vaillanti Steindachner, 1877c: 225. Type locality: Rio Preto [at Villa de Santa Rita, Bahia, Brazil]. Syntypes: MCZ 7842 (2), NMW 44273 (1), NMW 44276 (3), NMW 44277 (1).
Distribution: Preto River basin of São Francisco River drainage, Brazil (Weber, 2003).
Hypostomus variipictus (Ihering, 1911)
Plecostomus variipictus Ihering, 1911: 390. Type locality: Rio Pardo, perto de Barretos, Estado de São Paulo, quasiem sua foz no Rio Grande, tributario do lado esquerdo do Rio Paraná, Brazil. Holotype: MZUSP 2114.
Distribution: Grande River basin, Brazil (Weber, 2003).
Hypostomus varimaculosus (Fowler, 1945)
Plecostomus varimaculosus Fowler, 1945a: 113, fig. 14–16. Type locality: Morelia, Río Caquetá drainage, Colombia. Holotype: ANSP 71707.
Distribution: Upper Japurá River basin, Upper Amazon drainage of Brazil and Colombia (Weber, 2003).
Hypostomus variostictus (Miranda Ribeiro, 1912)
Plecostomus variostictus Miranda Ribeiro, 1912: 6. Type locality: Coxim, M[ato] Grosso [Brazil]. Holotype: MNRJ
1072.
Distribution: Upper Paraguay River basin, Brazil; only known from holotype (Weber, 2003).

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Hypostomus ventromaculatus Boeseman, 1968
Hypostomus ventromaculatus Boeseman, 1968: 65, pl. 15 (fig. 1). Type locality: Surinam River between Afobaka
and Brokopondo, Surinam. Holotype: RMNH 25507.
Distribution: Coastal drainages of northern South America from Oyapock River to Suriname River (Weber, 2003).
Remarks: Redescribed in Le Bail et al. (2000: 256).
Hypostomus vermicularis (Eigenmann & Eigenmann, 1888)
Plecostomus vermicularis Eigenmann & Eigenmann, 1888b: 171. Type locality: Rio Parahyba; Rio Janeiro; Mendez; Macacos, Goiás [Brazil]. Syntypes (37, 8–30 cm): BMNH 1904.1.28.4–5 (2), MCZ 7814 (1), MCZ 7848
(12), MCZ 7849 (1), MCZ 7850 (4), MCZ 7851 (1), MCZ 7857 (1), MCZ 91352 (12), NMW 44279 (1), USNM
123010 (3).
Distribution: Coastal drainages of eastern Brazil (Weber, 2003).
Remarks: Type series apparently mixed (Weber, 2003), and in need of a lectotype designation to stabilize current
usage.
Hypostomus waiampi Hollanda Carvalho & Weber, 2004
Hypostomus waiampi Hollanda Carvalho & Weber, 2004: 955, fig. 2. Type locality: Brazil: Amapá, bridge on theroad to Serra do Navioo, Cupixi river. Holotype: MZUSP 82269.
Distribution: Cupixi River, Amapá, Brazil (Hollanda Carvalho & Weber, 2004).
Remarks: Included in Hypostomus cochliodon species group by Hollanda Carvalho & Weber (2004).
Hypostomus watwata Hancock, 1828
Hypostomus watwata Hancock, 1828: 246. Type locality: Demerara [...] on the sea-shores [now: Berbice River,
Guyana, by neotype designation]. Neotype: BMNH 1932.11.10.31; designated by, and illustrated in, Boeseman(1868: 71, pl. 16, fig. 1).
Hypostomus verres Valenciennes in Cuvier & Valenciennes, 1840b: 494 (365 in Strasbourg deluxe edition). Type
locality: envoyés de Cayenne [French Guiana]. Lectotype: MNHN a-9450, designated by Boeseman (1968: 70).
Distribution: coastal drainages of northern South America from Oyapock River to Demerara River, and Venezuela(Weber, 2003).
Remarks: Redescribed in Le Bail et al. (2000: 260).
Hypostomus winzi (Fowler, 1945)
Plecostomus winzi Fowler, 1945b: 9, figs. 4–7. Type locality: Honda, Colombia. Holotype: ANSP 71623.
Distribution: Magdalena River basin, Colombia, known only from holotype (Weber, 2003).
Hypostomus wuchereri (Günther, 1864)
Plecostomus wuchereri Günther, 1864: 235. Type locality: Bahia, Brazil. Syntypes (4): BMNH 1863.3.27.15 (1).
Distribution: Paraguaçu River basin (?), Brazil (Weber, 2003).
ISBRUECKERICHTHYS Derijst, 1996
Isbrueckerichthys Derijst, 1996: 64. Type species: Hemipsilichthys duseni Miranda Ribeiro, 1907. Type by original
designation. Gender: Masculine.
Revision and key to species: Pereira & Reis (2002).
Isbrueckerichthys alipionis (Gosline, 1947)
Pareiorhaphis alipionis Gosline, 1947: 104. Type locality: Rio Bethary, São Paulo [Brazil]. Holotype: MNRJ 4241.
Distribution: Betari River basin, Ribeira de Iguape River drainage, Brazil (Weber, 2003).
Isbrueckerichthys duseni (Miranda Ribeiro, 1907)
Hemipsilichthys duseni Miranda Ribeiro, 1907a: 187, figure on page 187. Type locality: do Paraná ..[&]... do
Ribeirão do Monjolinho, á 300 metros sobre o mar [Iporanga River, São Paulo, Brazil]. Lectotype: MNRJ 772,
designated by Gosline (1947: 102).
Distribution: Upper Ribeira de Iguape River in Paraná State, Brazil (Weber, 2003).
Isbrueckerichthys epakmos Pereira & Oyakawa, 2003
Isbrueckerichthys epakmos Pereira & Oyakawa, 2003: 6, fig. 2. Type localilty: Brazil. São Paulo: Tapiraí: rio
Ribeira de Laguna drainage, rio Verde at Piúva, on road to Rio Verde, 23º58'22"S, 47º34'23"W. Holotype:
MZUSP 79804.

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ISORINELORICARIA Isbrücker, 1980
Isorineloricaria Isbrücker, 1980: 15. Type species: Plecostomus spinosissimus Steindachner, 1880. Type by original
designation. Gender: Feminine.
Isorineloricaria spinosissima (Steindachner, 1880)
Plecostomus spinosissimus Steindachner, 1880b: 98, pl. 5 (figs. 1, 1a). Type locality: Guayaquil [Ecuador]. Holotype: NMW 55027.
Plecostomus festae Boulenger, 1898c: 11. Type locality: Rio Vinces and Rio Peripa, Equateur. Syntypes: BMNH
1898.11.4.32 (1), ZMUT 1518 (1).
Distribution: Pacific versant rivers of Ecuador (Weber, 2003).
IXINANDRIA Isbrücker & Nijssen, 1979
Ixinandria Isbrücker & Nijssen, in Isbrücker, 1979a: 91. Type species: Loricaria steinbachi Regan, 1906. Type byoriginal designation. Gender: Feminine.
Ixinandria montebelloi (Fowler, 1940)
Canthopomus montebelloi Fowler, 1940c: 55, figs. 9–11. Type locality: Monte Bello, Tarija, Bolivia. Holotype:
ANSP 68832.
Distribution: Southern Bolivia.
Ixinandria steinbachi (Regan, 1906)
Loricaria Steinbachi Regan, 1906a: 97. Type locality: Salta, Argentina. Lectotype: BMNH 1906.5.31.37, designatedby Isbrücker (1979a: 113), and illustrated in Isbrücker (1979a: 91, figs. 1 a–c, 2, 3).
Distribution: Salado River basin, Argentina.
Remarks: Species reviewed by Ringuelet et al. (1967).
KRONICHTHYS Miranda Ribeiro, 1908
Kronichthys Miranda Ribeiro, 1908b: [1]. Type species: Kronichthys subteres Miranda Ribeiro, 1908. Type bymonotypy. Gender: Masculine.
Remarks: Also described as new in Miranda Ribeiro (1908a).
Kronichthys heylandi (Boulenger, 1900)
Plecostomus heylandi Boulenger, 1900b: 165. Type locality: Mountain stream in the Province São Paulo [...] 400
feet above sea-level near Santos [Brazil]. Holotype: BMNH 1899.12.18.1.
Distribution: coastal streams of eastern Brazil, between Santos and Rio de Janeiro (Weber, 2003).
Remarks: Redescribed in Gosline (1947: 106).
Kronichthys lacerta (Nichols, 1919)
Plecostomus lacerta Nichols, 1919a: 414. Type locality: Poço Grande (Estado de São Paulo) Rio Juquia, Brazil.
Holotype: AMNH 7151.
Distribution: Baia de Paranaguá basin and Ribeira de Iguape River basin, Brazil (Weber, 2003).
Remarks: Redescribed in Armbruster & Page (1997: 228).
Kronichthys subteres Miranda Ribeiro, 1908
Kronichthys subteres Miranda Ribeiro, 1908b: [2], fig. 1. Type locality: Rios Bethary, Pardo e Iporanga [Brazil].
Syntypes: MNRJ 655 (4).
Distribution: Ribeira de Iguape River basin, Brazil (Weber, 2003).
Remarks: Also described as new in Miranda Ribeiro (1908a).
LAMONTICHTHYS Miranda Ribeiro, 1939
Lamontichthys Miranda Ribeiro, P., 1939a: 12. Type species: Harttia filamentosa La Monte, 1935. Type by original
designation. Gender: Masculine.
Revision: Isbrücker & Nijssen (1978b).
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Harttia filamentosa La Monte, 1935: 5, fig. 4. Type locality: Brazilian Amazonas, Rio Jurua, in the vicinity of themouth of the Rio Embira, a tributary of Rio Tarauaca, which, in turn, is a tributary of Rio Jurua, 70º15'W,
7º30'S. Holotype: AMNH 12616.
Harttia filamentissima Eigenmann & Allen, 1942: 211, pl. 8 (figs. 1–2). Type locality: Rio Huallaga [Peru]. Holotype: CAS 28541.
Distribution: Western Amazon River basin (Ferraris, 2003c).
Lamontichthys llanero Taphorn & Lilyestrom, 1984
Lamontichthys llanero Taphorn & Lilyestrom, 1984: 96, fig. 2. Type locality: Venezuela, Estado Portuguesa, Río
Guanare Viejo cerca de Hoyada. Holotype: MCNG 10000.
Distribution: Guanare Viejo River basin, Orinoco River drainage, Venezuela (Taphorn & Lilyestrom, 1984).
Lamontichthys maracaibero Taphorn & Lilyestrom, 1984
Lamontichthys maracaibero Taphorn & Lilyestrom, 1984b: 94, fig. 1. Type locality: Venezuela, Estado Zulia, Distrito Miranda, Río Motatán, en el puente en Agua Viva. Holotype: MCNG 3592.
Distribution: Lake Maracaibo basin, Venezuela (Taphorn & Lilyestrom, 1984).
Lamontichthys stibaros Isbrücker & Nijssen, 1978
Lamontichthys stibaros Isbrücker & Nijssen, 1978b: 66, fig. 8. Type locality: Ecuador, Prov. Pastaza, Río Amazonas
system, Río Bobonaza at Chicherota, 2º25'S, 76º38'W, altitude approximately 260–280 m., upper Río Pastaza.
Holotype: USNM 167914.
Distribution: Bobonaza River basin, Ecuador (Isbrücker & Nijssen, 1978b).
LAMPIELLA Isbrücker, 2001
Lampiella Isbrücker, in Isbrücker et al., 2001: 19. Type species: Otocinclus gibbosus Miranda Ribeiro, 1908. Typeby original designation. Gender: Feminine.
Remarks: Genus created to accommodate Otocinclus gibbosus Miranda Ribeiro, which Schaefer (1997) removed
from Otocinclus, but did not place elsewhere.
Lampiella gibbosa (Miranda Ribeiro, 1908)
Otocinclus gibbosus Miranda Ribeiro, 1908b: [2], figs. 2–2a. Type locality: Rio Bethary [Brazil]. Holotype: MNRJ
1048.
Distribution: Ribeira de Iguape River basin, Brazil (Schaefer, 2003c).
LASIANCISTRUS Regan, 1904
Lasiancistrus Regan, 1904b: 224. Type species: Chaetostomus heteracanthus Günther, 1869. Type by subsequent
designation by Eigenmann (1910: 409). Gender: Masculine. Proposed originally as a subgenus of Ancistrus
Kner.
Phylogeny: Armbruster & Provenzano (2000).
Revision: Heitmans, et al. (1983), Armbruster (2005).
Key to species: Armbruster (2005: 552).
Lasiancistrus caucanus Eigenmann, 1912
Lasiancistrus caucanus Eigenmann, 1912a: 11. Type locality: Cartago; the Cauca, Colombia. Holotype: FMNH
56034; holotype illustrated in Eigenmann (1922: 76, pl. 11, fig. 4).
Hemiancistrus mayoloi Eigenmann, 1912a: 10. Type locality: Istmina, San Juan River, Colombia. Holotype: FMNH
56036.
Ancistrus planiceps Meek & Hildebrand, 1913: 79. Type locality: Rio Tuyra, Boca de Cupe, Panama. Holotype:
FMNH 7580.
Lasiancistrus volcanensis Dahl, 1942: 83, fig. 3. Type locality: Rio Volcán near its junction to Rio San Bartolomé...tributary of Rio Magdalena, on the left side, between Rios Narés and Ité, municipio of Remedios, departmentof Antióquia, Republic of Colombia). Altitude approx. 600 m. Holotype: at ZMUL.
Distribution: Magdalena, Atrato, San Juan, Cauca, Tuyra, and Bayano River basin, Colombia and Panama (Armbruster, 2005).
Remarks: Armbruster (2005: 555) acted as first reviser in selecting Lasiancistrus caucanus as the valid name for the

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species; Maldonado-Ocampo et al. (2005), however, continue to treat Lasiancistrus mayoloi and L. volcanensis
as distinct species.
Lasiancistrus guacharote (Valenciennes, 1840)
Hypostomus guacharote Valenciennes, in Cuvier & Valenciennes, 1840b: 508 (375 in Strasbourg deluxe edition).
Type locality: Les eaux douces de Porto-Ricco [=Puerto Rico, in error]. Syntype: MNHN a-9567 (1).
Ancistrus mystacinus Kner, 1854: 276. Type locality: Carracas [Venezuela]. Holotype: NMW 44200. Originally as
Anc. mystacinus.
Lasiancistrus maracaiboensis Schultz, 1944c: 314, pl. 11 (fig. a). Type locality: Río Socuy, 3 km. above its mouth.
Venezuela. Holotype: USNM 121038.
Distribution: Lake Maracaibo basin, Venezuela (Armbruster, 2005).
Lasiancistrus heteracanthus (Günther, 1869)
Chaetostomus heteracanthus Günther, 1869: 425, fig. 3. Type locality: Upper Amazons, Peruvian Amazons. Holotype: BMNH 1869.5.21.3.
Distribution: Napo River, Ecuador and Peru (Armbruster, 2005).
Remarks: Type locality interpreted by Eigenmann & Eigenmann (1891: 41; see also Eigenmann & Allen 1942: 18)
as Marañon River, but see different interpertation in Armbruster (2005).
Lasiancistrus saetiger Armbruster, 2005
Lasiancistrus saetiger Armbruster, 2005: 559, fig. 7. Type locality: Brazil, Pará, rio Guamá near Ourém, 1º34'07"S,
47º10'08"W. Holotype: MCP 37942.
Distribution: Guama River, Capim River basin, Brazil (Armbruster, 2005).
Lasiancistrus schomburgkii (Günther, 1864)
Hypostomus pictus Castelnau, 1855: 44, pl. 22 (fig. 2). Type locality: l’Ucayale [Peru]. Holotype: MNHN a-9573.
Preoccupied in Hemiancistrus by Ancistrus pictus Kner, 1854; replaced by Hemiancistrus castelnaui Miranda
Ribeiro, 1911.
Chaetostomus schomburgkii Günther, 1864: 245. Type locality: British Guiana. Syntypes: BMNH 1845.3.5.26–27(2).
Hemiancistrus castelnaui Miranda Ribeiro, 1911: 58. Type locality: l’Ucayale [Peru]. Holotype: MNHN a-9573.
Replacement for Hypostomus pictus Castelnau, 1855, which was then preoccupied in Hemiancistrus by Ancistrus pictus Kner, 1854.
Ancistrus multispinis Holly, 1929: 119. Type locality: Mercado Blèin (Brasilien). Holotype: at NMW.
Hemiancistrus caquetae Fowler, 1945a: 115, figs. 17–20. Type locality: Morelia, Río Caquetá drainage, Colombia.
Holotype: ANSP 71708.
Lasiancistrus scolymus Nijssen & Isbrücker, 1985: 242, fig. 1. Type locality: Brazil, Est. Mato Grosso do Sul, Rio
Aripuanã, Humboldt (10º10'S 59º27'W), Rio Madeira system. Holotype: MZUSP 26809.
Lasiancistrus guapore Knaack, 2000a: 57, figure on p. 58. Type locality: Brasilien, Mato Grosso, im Hauptstrom desRio Guaporé im Stadtgebiet von Pontes E Lacerda; S 15º07.627', W 58º57.786'. Holotype: MCP 28678.
Distribution: Amazon, Upper Orinoco and Upper Essequibo River basins (Armbruster, 2005).
Lasiancistrus tentaculatus Armbruster, 2005
Lasiancistrus tentaculatus Armbruster, 2005: 565, fig. 9. Type locality: río Las Marias, near El Portrero, below themetal bridge. 09º05'N, 69º39'W [Venezuela]. Holotype: MCNG 19744.
Distribution: Orinoco River basin and Valencia Lake basin, Venezuela and Colombia (Armbruster, 2005).
LEPORACANTHICUS Isbrücker & Nijssen, 1989
Leporacanthicus Isbrücker & Nijssen, 1989: 544. Type species: Leporacanthicus galaxias Isbrücker & Nijssen,
1989. Type by original designation. Gender: Masculine.
Leporacanthicus galaxias Isbrücker & Nijssen, 1989
Leporacanthicus galaxias Isbrücker & Nijssen, 1989: 546, fig. 10. Type locality: Brasilien, Est. Pará, Rio Tocantins,
Lagoa em frenta a Jacobal. Holotype: MZUSP 24136.
Distribution: Southern tributaries of middle and lower Amazon, including Madeira, Tocantins and Guamá rivers;
Ventuari River basin, upper Orinoco drainage (Provenzano, 1995; Fisch-Muller, 2003).

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Leporacanthicus heterodon Isbrücker & Nijssen, 1989
Leporacanthicus heterodon Isbrücker & Nijssen, 1989: 547, fig. 12. Type locality: Brasilien, Est. Mato Grosso, Rio
Xingú, Cachoeira von Martius, oberer Xingú. Holotype: IRSNB 693.
Distribution: Xingu River basin, Brazil (Fisch-Muller, 2003).
Leporacanthicus joselimai Isbrücker & Nijssen, 1989
Leporacanthicus joselimai Isbrücker & Nijssen, 1989: 546, fig. 11. Type locality: Brasilien, Est. Pará, Maloquinha,
perta de Itaituba, Rio Tapajós. Holotype: MZUSP 21921.
Distribution: Tapajós River basin, Brazil (Fisch-Muller, 2003).
Leporacanthicus triactis Isbrücker, Nijssen & Nico, 1992
Leporacanthicus triactis Isbrücker, Nijssen & Nico, 1992: 3, fig. 3. Type locality: Venezuela, Territorio Federal
Amazonas, oberer Orinoco, Caño Mavaquita etwa einen Kilometer flussaufwärts von der Mündung in den Río
Mavaca, ungefähr 2º12'30"N, 65º05'30"W. Holotype: MCNG 25357.
Distribution: Upper Orinoco River basin (Fisch-Muller, 2003).
LEPTOANCISTRUS Meek & Hildebrand, 1916
Leptoancistrus Meek & Hildebrand, 1916: 254. Type species: Acanthicus canensis Meek & Hildebrand, 1913. Typeby original designation. Gender: Masculine.
Monophyly: Schaefer (1986: 108).
Leptoancistrus canensis (Meek & Hildebrand, 1913)
Acanthicus canensis Meek & Hildebrand, 1913: 80. Type locality: Rio Cana, Cana, Darien, Panama. Holotype:
FMNH 7581. Species illustrated and described in more detail by Meek & Hildebrand (1916: 254, pl. 11).
Distribution: Pacific drainages of eastern Panama; Armila River basin on Caribbean slope, Panama (Fisch-Muller,
2003).
Leptoancistrus cordobensis Dahl, 1964
Leptoancistrus cordobensis Dahl, in Dahl et al., 1964: 32, unnum. plate. Type locality: río Batatal, Alto río Uré[upper San Jorge tributary, Magdalena basin, Colombia]. Holotype: Whereabouts unknown (possibly at ICNMHN).
Distribution: Upper Sinú River basin, and upper San Jorge River basin, Magdalena River drainage, Colombia(Fisch-Muller, 2003).
LIMATULICHTHYS Isbrücker & Nijssen, 1979
Limatulichthys Isbrücker & Nijssen, in Isbrücker, 1979a: 91. Type species: Loricaria punctata Regan, 1904. Type byoriginal designation. Gender: Masculine.
Limatulichthys griseus (Eigenmann, 1909)
Loricaria griseus Eigenmann, 1909a: 8. Type locality: Conawaruk [=Konawaruk, Guyana]. Holotype: FMNH
53077; holotype illustrated in Eigenmann (1912b: pl. 30, fig. 2; pl. 32, fig. 2).
Distribution: Essequibo and western Orinoco River basins (Ferraris, 2003c).
Limatulichthys petleyi (Fowler, 1940)
Loricaria punctata Regan, 1904b: 285, pl. 17 (fig. 1). Type locality: Manaos, Rio Negro, middle Amazon, and Portodo Moz, lower Amazon. Syntypes: BMNH 1893.4.24.18, and at MCZ. Preoccupied by Loricaria punctata
Kner, 1854 [now Pterygoplichthys punctatus], apparently not replaced.
Rhineloricaria petleyi Fowler, 1940a: 243, figs. 36–38. Type locality: Ucayali River basin, Contamana, Peru. Holotype: ANSP 68661.
Loricariichthys parnahybae Fowler, 1941a: 161, figs. 69–71. Type locality: Rio Parnahyba, Therizina, Piauhy [Brazil]. Holotype: ANSP 69451.
Distribution: Amazon, Tocantins and Parnaíba River basins (Ferraris, 2003c).
Remarks: Recently removed from the synonymy of Limatulichthys griseus (Eigenmann, 1909) by Isbrücker (2001).
LIPOPTERICHTHYS Norman, 1935
Lipopterichthys Norman, 1935b: 627. Type species: Lipopterichthys carrioni Norman, 1935. Type by original desig-

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nation. Gender: Masculine.
Monophyly: Schaefer (1986: 109).
Lipopterichthys carrioni Norman, 1935
Lipopterichthys carrioni Norman, 1935b: 628, fig. on p. 628 (top). Type locality: the Zamora River, near Loja, Ecuador. Holotype: BMNH 1933.5.29.1.
Distribution: Marañon River basin, Ecuador (Fisch-Muller, 2003).
LITHOGENES Eigenmann, 1909
Lithogenes Eigenmann, 1909a: 6. Type species: Lithogenes villosus Eigenmann, 1909. Type by original designation.
Gender: Masculine.
Phylogenetic position: Schaefer (2003d).
Lithogenes valencia Provenzano, Schaefer, Baskin & Royero, 2003
Lithogenes valencia Provenzano, Schaefer, Baskin & Royero, 2003: 564, fig. 1. Type locality: Venezuela, Est. Carabobo, Lago de Valencia basin. Holotype: MBUCV V-30740.
Distribution: Lake Valencia basin, northern Venezuela (Provenzano et al., 2003).
Lithogenes villosus Eigenmann, 1909
Lithogenes villosus Eigenmann, 1909a: 6. Type locality: Aruataima Falls, Upper Potaro. [Guyana]. Holotype:
FMNH 52960; holotype illustrated in Eigenmann (1912b: pl. 26, figs. 2–4) and Schaefer (2003d: fig. 2).
Distribution: Essequibo River basin, Potaro River, Guyana.
Remarks: Redescribed in Schaefer (2003d: 8).
LITHOXUS Eigenmann, 1910
Lithoxus Eigenmann, 1910: 405. Type species: Lithoxus lithoides Eigenmann, 1910. Type by original designation (p.
412). Gender: Masculine. Diagnostic characters given in key in footnote.
Paralithoxus Boeseman, 1982: 46. Type species: Ancistrus bovallii Regan, 1906. Type by original designation. Gender: Masculine. Proposed as a subgenus of Lithoxus. Paralithoides appears in the abstract of Boeseman (1982:41) and is treated here as an error.
Revision: Boeseman (1982), Nijssen & Isbrücker (1990)
Monophyly: Schaefer (1986: 110).
Lithoxus boujardi Muller & Isbrücker, 1993
Lithoxus boujardi Muller & Isbrücker, 1993: 72, fig. 1. Type locality: Guyane française, bassin de l’Approuague,
Arataye, entre le Saut Japigny et le Saut Pararé. Holotype: MNHN 1992-1321.
Distribution: Approuague and Oyapock River basins, French Guiana (Fisch-Muller, 2003).
Remarks: Redescribed in Le Bail et al. (2000: 242, fig.).
Lithoxus bovallii (Regan, 1906)
Ancistrus Bovallii Regan, 1906a: 96. Type locality: the Kaat River, tributary to the Treng River, Upper Potaro, British Guiana. Lectotype: BMNH 1905.11.1.43, designated by Nijssen & Isbrücker (1990: 332).
Distribution: Ireng River basin in upper Negro River drainage, Guyana (Fisch-Muller, 2003).
Remarks: Redescribed in Schindler (1996).
Lithoxus lithoides Eigenmann, 1910
Lithoxus lithoides Eigenmann, 1910: 412. Type locality: Cataracts of British Guiana. Holotype: FMNH 53557; holotype illustrated in Eigenmann (1912b: 242, pl. 29, figs. 1, 2, 4).
Distribution: Essequibo River, Guyana and upper Corantijn River basin, Suriname (Fisch-Muller, 2003).
Lithoxus pallidimaculatus Boeseman, 1982
Lithoxus (Paralithoxus) pallidimaculatus Boeseman, 1982: 50, pl. 3 (top). Type locality: Kwambaolo Creek, righttributary of Sara Creek above Dam, Suriname River system, Surinam. Holotype: RMNH 28368.
Distribution: Suriname River basin, Suriname (Fisch-Muller, 2003).
Lithoxus planquettei Boeseman, 1982
Lithoxus (Paralithoxus) planquettei Boeseman, 1982: 53, pl. 4 (top). Type locality: Crique Boulenger, Comté system, French Guyana. Holotype: RMNH 28304 (1 of 3).

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Distribution: Atlantic coastal drainages from Maroni to Kaw River basins, French Guiana (Fisch-Muller, 2003).
Remarks: Redescribed in Le Bail et al. (2000: 244).
Lithoxus stocki Nijssen & Isbrücker, 1990
Lithoxus stocki Nijssen & Isbrücker, 1990: 329, fig. 2. Type locality: Marouini River, downstream of village Epoia,
French Guiana. Holotype: IRSNB 639.
Distribution: Maroni and Mana River basins, French Guiana (Fisch-Muller, 2003).
Remarks: Redescribed in Le Bail et al. (2000: 246).
Lithoxus surinamensis Boeseman, 1982
Lithoxus (Paralithoxus) surinamensis Boeseman, 1982: 54, pl. 5 (top). Type locality: near Awaradam, Gran Rio,
upper Suriname River system, Surinam. Holotype: RMNH 28361 (largest specimen).
Distribution: Gran Rio River basin in upper Suriname River drainage, Suriname (Fisch-Muller, 2003).
LORICARIA Linnaeus, 1758
Loricaria Linnaeus, 1758: 307. Type species: Loricaria cataphracta Linnaeus, 1758. Type by monotypy. Gender:
Feminine.
Fusiloricaria Fowler, 1940a: 247. Type species: Loricaria clavipinna Fowler, 1940. Type by original designation.
Gender: Feminine. Originally proposed as a subgenus of Loricaria.
Revision: Isbrücker (1981).
Loricaria apeltogaster Boulenger, 1895
Loricaria apeltogaster Boulenger, 1895b: 528. Type locality: Paraguay. Lectotype: BMNH 1895.5.17.105, designated by Isbrücker (1979a: 110) and illustrated in Isbrücker (1979a: 107, figs. 20a, 21a); one syntype illustrated
in Boulenger (1896c: pl. 7, fig. 2).
Distribution: Uruguay, Paraná and Paraguay River basins (Isbrücker, 1981).
Loricaria cataphracta Linnaeus, 1758
Loricaria cataphracta Linnaeus, 1758: 307. Type locality: In America meridionali. Lectotype: NRM 33, designatedby Isbrücker (1979a: 110). Earlier neotype designation by Isbrücker (1972: 173) invalidated by rediscovery of
syntype series.
Loricaria cirrhosa Bloch & Schneider, 1801: 125, pl. 34. Type locality: America meridionali. Lectotype: ZMB
3160, designated by, and illustrated in, Isbrücker (1972: 170, figs. 4–5).
Loricaria setifera La Cepède, 1803: 140, 141. Type locality: In America meridionali. Syntype: NRM 33. On
Gronovius (1763: n. 69), Bloch (1794: pl. 375, fig. 3), Loricaria cataphracta Linnaeus, Seba (1734–65: pl. 29,
fig. 14), and other literature sources.
Loricaria Dentata Shaw, 1804: 37. Type locality: Indian Seas, in error [South America]. Types: On Gronovius(1763: 127, pl. 2, figs. 1–2), Bloch (1794: pl. 375, figs. 3–4), and Loricaria cataphracta Linnaeus.
Plecostomus flagellaris Gronow in Gray, 1854: 158. Type locality: Surinam. Holotype: BMNH 1863.11.12.195(dorsal surface of body) and 196 (ventral surface); holotype redescribed and illustrated in Isbrücker (1972: fig.
7).
Loricaria carinata Castelnau, 1855: 46, pl. 23 (fig. 3). Type locality: De la rivières Amazones. Holotype: MNHN a9562; holotype redescribed and illustrated in Isbrücker (1972: 178, fig. 8).
Loricaria dura Bleeker, 1862 (in Bleeker, 1862–63): 3. Type locality: Surinama. Lectotype: NRM 33. Unneeded
new name for Loricaria cataphracta Linnaeus.
Distribution: Amazon River basin, and coastal rivers of the Guianas, Brazil, French Guiana, Guyana, Suriname(Isbrücker, 1981).
Remarks: Species redescribed by le Bail et al. (2000); see Isbrücker (1972) for comments on the identity of the species.
Loricaria clavipinna Fowler, 1940
Loricaria clavipinna Fowler, 1940a: 247, figs. 44–47. Type locality: Ucayali River basin, Contamana, Peru. Holotype: ANSP 68665; holotype illustrated in Isbrücker (1981: figs. 9–10).
Distribution: Amazon River basin, Brazil, Peru (Isbrücker, 1981).
Loricaria lata Eigenmann & Eigenmann, 1889

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Loricaria lata Eigenmann & Eigenmann, 1889b: 36. Type locality: Goyaz [Brazil]. Lectotype: MCZ 46721, designated by, and illustrated in, Isbrücker (1972: 179, figs. 9–10).
Distribution: Araguaia River basin, Brazil (Isbrücker, 1981).
Loricaria nickeriensis Isbrücker, 1979
Loricaria nickeriensis Isbrücker, 1979a: 97, figs. 11–13. Type locality: Surinam, district Nickerie, rapide dans la rivière Fallawatra, 5 km SSW des chutes Stondansie. Holotype: ZMA 107561.
Distribution: Nickerie and Marowijne River basins (Isbrücker, 1981).
Loricaria parnahybae Steindachner, 1907
Loricaria parnahybae Steindachner, 1907b: 153. Type locality: dem Rio Parnahyba an der Mündung eines Baches
bei Victoria [Brazil]. Lectotype: NMW 44854, designated by Isbrücker (1979a: 110).
Loricaria piauhiae Fowler, 1941a: 163, figs. 72–74. Type locality: Rio Parnahyba, Therezina, Piauhy [Brazil]. Holotype: ANSP 69452; holotype illustrated in Isbrücker (1981: fig. 7).
Distribution: Coastal rivers of NE Brazil and French Guiana (Isbrücker, 1981).
Loricaria piracicabae Ihering, 1907
Loricaria piracicabae Ihering, 1907: 28. Type locality: Piracicaba, Piracicaba River, State of S. Paulo [Brazil].
Holotype: MZUSP 2182.
Distribution: Piracicaba River basin, Brazil (Isbrücker, 1981).
Loricaria simillima Regan, 1904
Loricaria simillima Regan, 1904b: 292, pl. 17 (fig. 2). Type locality: Canelos, e. Ecuador. Lectotype: BMNH
1880.12.8.77, designated by Isbrücker (1979a: 110).
Distribution: Amazon, Orinoco and La Plata River basins (Isbrücker, 1981).
Loricaria tucumanensis Isbrücker, 1979
Loricaria tucumanensis Isbrücker, 1979a: 98, fig. 14. Type locality: Argentine, Prov. Tucuman, San Miguel deTucuman, 26º47'S, 65º15'W (Rio Sali ?). Holotype: USNM 88669.
Distribution: Dulce River basin, Argentina (Isbrücker, 1981).
LORICARIICHTHYS Bleeker, 1862
Plecostomus Swainson, 1839: 304. Type species: Loricaria maculata Bloch, 1794. Type by monotypy. Gender:
Masculine. Originally as a subgenus of Loricaria. Predates Loricariichthys Bleeker, but not treated as valid
since before 1899 and, therefore, permanently suppressed.
Loricariichthys Bleeker, 1862 (in Bleeker, 1862–63): 3. Type species: Loricaria maculata Bloch, 1794. Type byoriginal designation. Gender: Masculine.
Parahemiodon Bleeker, 1862: 373. Type species: Parahemiodon typus Bleeker, 1862. Type by original designation.
Gender: Masculine.
Loricariichthys acutus (Valenciennes, 1840)
Loricaria acuta Valenciennes, in Cuvier & Valenciennes, 1840b: 472 (349 of Strasbourg deluxe edition), pl. 452.
Type locality: On peut croire qu’it venait du Brésil. Holotype: at MNHN (but apparently lost).
Distribution: Brazil (?).
Loricariichthys anus (Valenciennes, 1836)
Loricaria anus Valenciennes, 1836, in Valenciennes, 1835–47: pl. 6 (fig. 1). Type locality: [Not stated]. Holotype:
MNHN a-9456 or MNHN a-9455. Name made available from illustration of single specimen, which must be
considered the holotype if it can be recognized. Described in Cuvier & Valenciennes (1840b: 470 [347 of Strasbourg deluxe edition]).
Distribution: Coastal rivers of southern Brazil, Uruguay and lower Paraná River basins (Ferraris, 2003c).
Remarks: Redescribed in Cuvier & Valenciennes (1840: 470 (347 in Strasbourg deluxe edition) from two specimens,with locality stated as: sur les bordes de la rivière de La Plata près de Buénos-Ayres. Lectotype designation by
Isbrücker (1979a: 113) of MNHN a-9456 (396 mm) would be valid, if illustrated specimen can not be identified.
Loricariichthys brunneus (Hancock, 1828)
Loricaria brunnea Hancock, 1828: 247. Type locality: Venezuela, the branches and lakes of the Orinoko [sic].

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Type(s): Whereabouts unknown (Isbrücker, 1979a: 113).
Distribution: Orinoco River basin (Ferraris, 2003c).
Loricariichthys cashibo (Eigenmann & Allen, 1942)
Loricaria cashibo Eigenmann & Allen, 1942: 205. Type locality: Lago Cashiboya [Peru]. Holotype: IU 15416 (131mm FL; whereabouts unknown, not found by Isbrücker & Nijssen, 1976b: 110).
Distribution: Upper Amazon River basin, Peru (Ferraris, 2003c).
Loricariichthys castaneus (Castelnau, 1855)
Loricaria castanea Castelnau, 1855: 46, pl. 23 (fig. 4). Type locality: donné a Rio de Janeiro, comme venant des rivières du voisinage [Brazil]. Holotype: MNHN a-9561.
Loricaria Spixii Steindachner, 1881b: 97. Type locality: Rio Parahyba, Santa Cruz, Rio Quenda, Rio Muriahe, Sao
Matheos, Sambaia, Rio Grande do Sul [restricted to Rio Quenda by lectotype designation] . Lectotype: NMW
45113, designated by Isbrücker (1979a: 113). Illustrated and described in more detail in Steindachner (1881d: 4,
pl. 2).
Distribution: Coastal streams of SE Brazil between southern São Paulo and northern Espírito Santo, including the
Paraíba do Sul River basin, Brazil (Ferraris, 2003c).
Loricariichthys chanjoo (Fowler, 1940)
Parahemiodon chanjoo Fowler, 1940a: 240, figs. 30–32. Type locality: Ucayali River basin, Contamana, Peru.
Holotype: ANSP 68656.
Distribution: Ucayali River basin, Peru (Ferraris, 2003c).
Loricariichthys derbyi Fowler, 1915
Loricariichthys derbyi Fowler, 1915b: 267, fig. 4. Type locality: Rio Jaguaribe at Barro Alto, Brazil. Holotype:
ANSP 39932.
Distribution: Jaguaribe River basin, Brazil (Ferraris, 2003c).
Loricariichthys edentatus Reis & Pereira, 2000
Loricariichthys edentatus Reis & Pereira, 2000: 1041, fig. 8. Type locality: rio Uruguay at brazo Itape, south of Isla
Canario, Concepción del Uruguay, Argentina (32º29'S, 58º14'W). Holotype: MACN 4041.
Distribution: Lower Uruguay River basin, Argentina (Reis & Pereira, 2000).
Loricariichthys hauxwelli Fowler, 1915
Loricariichthys hauxwelli Fowler, 1915a: 238, fig. 10. Type locality: Ambyiacu River, Ecuador. Holotype: ANSP
8301.
Distribution: Ampyiacu River basin, Peru (Ferraris, 2003c).
Loricariichthys labialis (Boulenger, 1895)
Loricaria labialis Boulenger, 1895b: 527. Type locality: Paraguay. Lectotype: BMNH 1895.5.17.98, designated byIsbrücker (1979a: 113); one syntype illustrated in Boulenger (1896c: pl. 7, fig. 1).
Distribution: Paraguay and middle Paraná River basin (Ferraris, 2003c).
Remarks: Reviewed in Ringuelet et al. (1967), diagnosed in Britski et al. (1999).
Loricariichthys maculatus (Bloch, 1794)
Loricaria maculata Bloch, 1794: 73, pl. 375 (figs. 1–2). Type locality: [No locality stated]. Lectotype: ZMB 3163,
designated by, and illustrated in, Isbrücker (1971a: 12, figs. 3–4), with the locality stated as Surinam .... District
Suriname, ditches at Cultuurtuin at Paramaribo West.
Loricaria accipenser Shaw, 1804: 36. Type locality: Indian seas, in error [South America]. Types: On Bloch (1794:
pl. 375, fig. 1), Gronovius (1763: 127, no. 391), and Loricaria cataphracta Linnaeus.
Plecostomus cataphractus Gronow in Gray, 1854: 158. Type locality: [No locality stated]. Neotype: ZMB 3163, designated by, and illustrated in, Isbrücker (1971a: 15, figs. 3–4).
Loricaria amazonica Castelnau, 1855: 46, pl. 23 (fig. 2). Type locality: [Not stated]. Holotype: MNHN a-9560.
? Parahemiodon typus Bleeker, 1862a: 373. Type locality: Surinama. Holotype: RMNH 3121; holotype illustrated in
Bleeker (1864a: 20, pl. 6, fig. 1; pl. 13, fig. 1).
? Loricaria parahemiodon Günther, 1864: 258. Type locality: Surinama. Holotype: RMNH 3121. Unneeded newname for Parahemiodon typus Bleeker.
Distribution: Rivers of Suriname (Ferraris, 2003c).

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Remarks: Redescribed in Isbrücker (1971a: 12).
Loricariichthys melanocheilus Reis & Pereira, 2000
Loricariichthys melanocheilus Reis & Pereira, 2000: 1042, fig. 9. Type locality: mouth of rio Ijuí-Mirim, tributary of
rio Ijuí, Pirapó, Rio Grande do Sul, Brazil (rio Uruguay drainage, 28º03'S, 55º11'W). Holotype: MCP 16961.
Distribution: Lower Paraná and Uruguay River basins, Brazil (Reis & Pereira, 2000).
Loricariichthys microdon (Eigenmann, 1909)
Loricaria microdon Eigenmann, 1909a: 7. Type locality: Rupununi, British Guiana. Holotype: FMNH 53555; holotype illustrated in Eigenmann (1912b: pl. 30, fig. 1; pl. 32, fig. 1).
Distribution: Rupununi River basin, Guyana (Ferraris, 2003c).
Loricariichthys nudirostris (Kner, 1853)
Loricaria nudirostris Kner, 1853a: 12, 22, pl. 4. Type locality: Barra do Rio negro [Manaus, Brazil]. Lectotype:
NMW 45092: 1, designated by Isbrücker (1979a: 113). Originally as Lor. nudirostris.
Distribution: Amazon River basin, Brazil (Ferraris, 2003c).
Loricariichthys platymetopon Isbrücker & Nijssen, 1979
Loricariichthys platymetopon Isbrücker & Nijssen, 1979: 203, fig. 7. Type locality: Paraguay, Lago Ypacarai, San
Bernardino, 25º16'S, 57º16'W, Río Paraguay system. Holotype: USNM 181580.
Distribution: La Plata River basin (Ferraris, 2003c).
Loricariichthys rostratus Reis & Pereira, 2000
Loricariichthys rostratus Reis & Pereira, 2000: 1044, fig. 10. Type locality: rio São Francisco Falso, limit between
Diamante do Oeste and Santa Helena, Paraná, Brazil (upper rio Paraná drainage, 24º54'S, 54º12'W). Holotype:
MCP 16962.
Distribution: Paraná River basin, above Paraguay River mouth (Reis & Pereira, 2000).
Loricariichthys stuebelii (Steindachner, 1882)
Loricaria Stübelii Steindachner, 1882b: 175. Type locality: Aus dem Huallaga [Peru]. Lectotype: NMW 45110, designated by Isbrücker (1979a: 113). Illustrated and described in more detail in Steindachner (1882c: 7, pl. 3, figs.
2, 2a, 2b).
Distribution: Huallaga River basin, Upper Amazon drainage, Peru (Ferraris, 2003c).
Loricariichthys ucayalensis Regan, 1913
Loricariichthys ucayalensis Regan, 1913c: 282. Type locality: River Ucayali, Peru. Lectotype: BMNH
1913.7.30.36, designated by Isbrücker & Nijssen (1979: 208).
Distribution: Ucayali River basin, Upper Amazon drainage, Peru (Ferraris, 2003c).
MACROTOCINCLUS Isbrücker & Seidel, 2001
Macrotocinclus Isbrücker & Seidel, in Isbrücker et al., 2001: 20. Type species: Otocinclus affinis Steindachner,
1877. Type by original designation. Gender: Masculine.
Remarks: This genus corresponds to the “affinis” clade of Otocinclus as diagnosed in Schaefer (1997) and is the sister group to Otocinclus sensu stricto.
Macrotocinclus affinis (Steindachner, 1877)
Otocinclus affinis Steindachner, 1877c: 221, pl. 1 (figs. 1, 1a, 1b). Type locality: Flüssen bei S. Crux in der umgebung von Rio de Janeiro, Brazil. Lectotype: NMW 45368:1, designated by, and illustrated in [but labeled as
NMW 45368:2], Schaefer (1997: 49).
Distribution: Vicinity of Rio de Janeiro, Brazil (Schaefer, 2003c).
Macrotocinclus flexilis (Cope, 1894)
Otocinclus fimbriatus Cope, 1894: 98, pl. 9 (figs. 16, 16a–b). Type locality: Rio Jacuhy [Jacuí], Rio Grande do Sul,
Brazil. Syntypes: ANSP 21752–55 (4), ANSP 21585–97 (14); syntype ANSP 21754 illustrated in Schaefer(1997: fig. 34a).
Otocinclus flexilis Cope, 1894: 97, pl. 8 (figs. 13, 13a–b). Type locality: Rio Jacuhy [Jacuí], Rio Grande do Sul, Brazil. Lectotype: ANSP 21756, designated by, and illustrated in, Schaefer (1997: 53, fig. 33).
Otocinclus arnoldi Regan, 1909: 234. Type locality: Rio de la Plata, South America. Holotype: BMNH
1908.12.5.13; holotype ilustrated in Schaefer (1997: fig. 34b).

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Distribution: Middle and lower Paraná/Paraguay, Uruguay and La Plata basins and Atlantic coastal streams ofSoutheastern Brazil (Schaefer, 2003c).
Remarks: Otocinclus flexilis treated as valid by first reviser action of Eigenmann (1910: 412). Redescribed in
Aquino (1996).
MEGALANCISTRUS Isbrücker, 1980
Megalancistrus Isbrücker, 1980: 52. Type species: Chaetostomus gigas Boulenger, 1895. Type by original designation. Gender: Masculine.
Megalancistrus barrae (Steindachner, 1910)
Ancistrus barrae Steindachner, 1910a: 58. Type locality: Rio San Francisco bei Barra [Brazil]. Syntypes: NMW
48019 (1), NMW 48020 (1).
Distribution: São Francisco River basin, Brazil (Fisch-Muller, 2003).
Megalancistrus parananus (Peters, 1881)
Pterygoplichthys (Ancistrus) parananus Peters, 1881a: 17. Type locality: Parana fluss bei La Paz in Entre-Rios[Argentina]. Holotype: ZMB 11328.
Chaetostomus aculeatus Perugia, 1891: 637. Type locality: Asuncion (Rio Paraguay) [Paraguay]. Holotype: MSNG
8937.
Chaetostomus gigas Boulenger, 1895b: 526. Type locality: Paraguay. Holotype: BMNH 1895.5.17.72; holotype
illustrated in Boulenger (1896c: pl. 6).
Distribution: Paraguay, Paraná, and Uruguay River basins (Fisch-Muller, 2003).
Remarks: Redescribed as Pterygoplichthys aculeatus in Pignalberi et al. (1970: 379).
METALORICARIA Isbrücker, 1975
Metaloricaria Isbrücker, 1975b: 2. Type species: Metaloricaria paucidens Isbrücker, 1975. Type by original designation. Gender: Feminine.
Review: Isbrücker & Nijssen (1982).
Metaloricaria nijsseni (Boeseman, 1976)
Harttia nijsseni Boeseman, 1976: 170, pl. 8. Type locality: Sipaliwini River, southwestern Surinam. Holotype:
RMNH 27498.
Distribution: Suriname, Saramacca, Nickerie, and Corantijn River basins, Suriname (Isbrücker & Nijssen, 1982).
Metaloricaria paucidens Isbrücker, 1975
Metaloricaria paucidens Isbrücker, 1975b: 2, pls 1–3. Type locality: French Guiana, creek at right bank of Ouaqui
River, upstream of Saut Bali, Maroni River system. Holotype: IRSNB 549.
Distribution: Oyapock, Sinnamary and Marowijne River basins (le Bail et al., 2000).
Remarks: Species redescribed by le Bail et al. (2000).
MICROLEPIDOGASTER Eigenmann & Eigenmann, 1889
Microlepidogaster Eigenmann & Eigenmann, 1889a: 8. Type species: Microlepidogaster perforatus Eigenmann &Eigenmann, 1889. Type by original designation. Gender: Feminine.
Remarks: Type species written as Microlepidogaster perforated in generic account, but as M. perforatus in species
account.
Microlepidogaster perforatus Eigenmann & Eigenmann, 1889
Microlepidogaster perforatus Eigenmann & Eigenmann, 1889a: 9. Type locality: Rio Carandahy, Brazil. Holotype:
MCZ 8181.
Distribution: Carandaí River in upper Paraná River basin, Brazil (Schaefer, 2003c).
NANNOPTOPOMA Schaefer, 1996
Nannoptopoma Schaefer, 1996a: 915. Type species: Otocinclus spectabilis Eigenmann, 1914. Type by original designation. Gender: Neuter.
Nannoptopoma spectabile (Eigenmann, 1914)

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FERRARIS

Otocinclus spectabilis Eigenmann, 1914b: 229. Type locality: Quebrada Cramalote, 4º07'N, 73º36'W, Dpto. Meta,
Villavicencio, Colombia. Holotype: CAS 33806.
Distribution: Amazon and Orinoco River basins, Colombia (Schaefer, 2003c).
Nannoptopoma sternoptychum Schaefer, 1996
Nannoptopoma sternoptychum Schaefer, 1996a: 920, fig. 4. Type locality: Quebrada Mariposa at Cusco-Amazonico
Lodge, 12º36'S, 69º11'W, Puerto Maldonado, Tambopata, Madre de Dios, Peru. Holotype: MUSM 4097.
Distribution: Lower Amazon River basin and tributaries below 200 m elevation (Schaefer, 2003c).
NEBLINICHTHYS Ferraris, Isbrücker & Nijssen, 1986
Neblinichthys Ferraris, Isbrücker & Nijssen, 1986: 70. Type species: Neblinichthys pilosus Ferraris, Isbrücker &Nijssen, 1986. Type by original designation. Gender: Masculine.
Neblinichthys pilosus Ferraris, Isbrücker & Nijssen, 1986
Neblinichthys pilosus Ferraris, Isbrücker & Nijssen, 1986: 70, fig. 1 (top). Type locality: Venezuela, Territorio Federal Amazonas; Dept. Rio Negro, Rio Baria basin. Rio Mawarinuma tributary at Neblina base camp, on rightbank in riffle, 0º55'N, 66º10'W, elevation 120 m. Holotype: AMNH 56137.
Distribution: Upper Baria River basin, Venezuela (Ferraris et al., 1986).
Neblinichthys yaravi (Steindachner, 1915)
Ancistrus (Hemiancistrus) yaravi Steindachner, 1915e: 87. Type locality: Rio Coquenan [=Kukenan, Caroni basin],
Venezuela. Holotype: at NMW.
Neblinichthys roraima Provenzano, Lasso & Ponte, 1995: 245, fig. 1. Type locality: Venezuela: Estado Bolivar: río
Caroni system, first creek (quebrada) NW of base camp Roraima tepui, tributary of río Kukenan (5º15'N,
60º40'W), at an elevation of 1200–1400 m. Holotype: MHNLS 8753.
Distribution: Upper Caroni drainage, Venezuela (Fisch-Muller, 2003).
Remarks: Synonymy follows Armbruster (pers. commun., 2005).
NEOPLECOSTOMUS Eigenmann & Eigenmann, 1888
Neoplecostomus Eigenmann & Eigenmann, 1888b: 171. Type species: Plecostomus microps Steindachner, 1877.
Type by original designation. Gender: Masculine. Proposed originally as a subgenus of Plecostomus.
Neoplecostomus espiritosantensis Langeani, 1990
Neoplecostomus espiritosantensis Langeani, 1990: 16, fig. 3. Type locality: rio Jacu (braço sul), Vitor Hugo, Mun.
de Domingos Martins, ES [Brazil]. Holotype: MZUSP 38573.
Distribution: South America: Jacu and São Lourenço River basins, eastern slope of Serra do Mar, Brazil (Langeani,1990).
Neoplecostomus franciscoensis Langeani, 1990
Neoplecostomus franciscoensis Langeani, 1990: 22, fig. 5. Type locality: riacho afluente do córrego da Mutuca, àdireita da estrada Belo Horizonte-Nova Lima, Km 20, MG, 20º06'S,43º55'W [Brazil]. Holotype: MZUSP
38577.
Distribution: Headwaters of Das Velhas and Paraopeba River basins, São Francisco River drainage, Brazil(Langeani, 1990).
Neoplecostomus granosus (Valenciennes, 1840)
Hypostomus granosus Valenciennes in Cuvier & Valenciennes, 1840b: 502 (371 in Strasbourg deluxe edition). Type
locality: Cayenne, ... Rio-Janéiro [both localities apparently in error (Langeani, 1990)]. Syntypes: MNHN a9566 (3), MNHN b-0597 (6), MNHN b-0598 (6).
Distribution: Southeastern Brazil (?).
Neoplecostomus microps (Steindachner, 1877)
Plecostomus microps Steindachner, 1877b: 688, pl. 13. Type locality: der Umgebung von Rio Janeiro [Brazil]. Lectotype: NMW 45337.1, designated by Langeani (1990: 8).
Distribution: Paraíba do Sul River basin, Brazil (Langeani, 1990).
Neoplecostomus paranensis Langeani, 1990
Neoplecostomus paranensis Langeani, 1990: 12, figs. 2, 6. Type locality: Rio Cubatão, Faz. Santa Carlota, Cajurú,

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SP [Brazil]. Holotype: MZUSP 38572.
Distribution: Upper Paraná River basin, Brazil (Langeani, 1990).
Neoplecostomus ribeirensis Langeani, 1990
Neoplecostomus ribeirensis Langeani, 1990: 19, fig. 4. Type locality: riacho afluente do rio Bananal, km 7 darodovia Biguá-Iguape (SP-222), SP [Brazil]. Holotype: MZUSP 38576.
Distribution: Ribeira de Iguape River basin, Brazil (Langeani, 1990).
Neoplecostomus variipictus Bizerril, 1995
Neoplecostomus variipictus Bizerril, 1995b: 695, figs. 1–2. Type locality: rio Santo Antônio, tributário do rio Bengala, bacia do rio Paraíba do Sul, Firburgo, RJ [Brazil]. Holotype: MNRJ 12806.
Distribution: Paraíba do Sul River basin, Brazil (Bizerril, 1995b).
NIOBICHTHYS Schaefer & Provenzano, 1998
Niobichthys Schaefer & Provenzano, 1998: 222. Type species: Niobichthys ferrarisi Schaefer & Provenzano, 1998.
Type by original designation. Gender: Masculine.
Niobichthys ferrarisi Schaefer & Provenzano, 1998
Niobichthys ferrarisi Schaefer & Provenzano, 1998: 226, fig. 4. Type locality: Municipio Autónomo Río Negro, Río
Baria, approximately 200 m above Neblina base camp, 0º55'N, 66º10'W, Amazonas State, Venezuela. Holotype:
MBUCV V-20225.
Distribution: Upper Baria River, Negro River basin, Venezuela (Schaefer, 2003c).
OLIGANCISTRUS Rapp Py-Daniel, 1989
Oligancistrus Rapp Py-Daniel, 1989: 246. Type species: Chaetostomus punctatissimus Steindachner, 1881. Type byoriginal designation. Gender: Masculine.
Oligancistrus punctatissimus (Steindachner, 1881)
Chaetostomus punctatissimus Steindachner, 1881c: 119, pl. 3 (fig. 3). Type locality: aus dem Amazonen-Stromeohne nähere Angabe des Fundortes [Brazil]. Holotype: NMW 47206.
Distribution: Xingu River basin, Brazil (Fisch-Muller, 2003).
OTOCINCLUS Cope, 1871
Otocinclus Cope, 1871b: 112. Type species: Otocinclus vestitus Cope, 1872. Type by subsequent monotypy. Gender:
Masculine. Genus originally proposed without species; species name first added in Cope (1872a).
Revision, phylogeny and biogeography: Schaefer (1997).
Otocinclus bororo Schaefer, 1997
Otocinclus bororo Schaefer, 1997: 69, fig. 43. Type locality: Barra do Bugres, stream crossing road between Barrado Bugres and Cáceres, 51 km S of Barra Do Bugres, ca. 15º20'S, 57º12'W, Mato Grosso, Brazil. Holotype:
MCP 19285.
Distribution: Upper Paraguay River basin, Brazil (Schaefer, 2003c).
Otocinclus caxarari Schaefer, 1997
Otocinclus caxarari Schaefer, 1997: 71, fig. 44. Type locality: Region between Guajara-Mirim and Mato Grosso, ca.
10º48'S, 65º23'W, Rio Solimoes/Madeira/Guaporé drainage, Brazil/Bolivia border. Holotype: MZUSP 51102.
Distribution: Upper Madeira River basin, Brazil (Schaefer, 2003c).
Otocinclus cocama Reis, 2004
Otocinclus cocama Reis, 2004: 110, figs. 1–4. Type locality: quebrada Yanayacu (approx. 04º55'S, 073º43'W), tributary to the caño of the cocha Supay in Jenaro Herrera, Provincia Requena, Departamento Loreto, Peru. Holotype: MUSM 20686.
Distribution: Upper Amazon River basin, Peru (Reis, 2004).
Otocinclus hasemani Steindachner, 1915
Otocinclus hasemani Steindachner, 1915c: 348. Type locality: Engenhão de Agua, im flußgebiete des Paranahyba[Maranhão State, Brazil]. Lectotype: NMW 45369: 1, designated by, and illustrated in, Schaefer (1997: 62, fig.
38a).

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FERRARISDistribution: Tocantins and Parnaíba River basins, Brazil (Schaefer, 2003c).
Otocinclus hoppei Miranda Ribeiro, 1939
Otocinclus hoppei Miranda Ribeiro, A., 1939: 359. Type locality: Belém, Pará, Brasil-Norte. Lectotype: MNRJ
1030, designated by Miranda Ribeiro (1953: 401); lectotype illustrated in Schaefer (1997: fig. 39a).
Distribution: Amazon River basin, Brazil (Schaefer, 2003c).
Otocinclus huaorani Schaefer, 1997
Otocinclus huaorani Schaefer, 1997: 74, fig. 46. Type locality: Trib. to Río San Miguel (km 50, 4.5 km S of Tipischca), ca. 0º10'N, 76º13'W, Río Amazonas/Napo drainage, Napo, Ecuador. Holotype: FMNH 101555.
Distribution: Western upper Amazon and Orinoco River basins (Schaefer, 2003c).
Otocinclus macrospilus Eigenmann & Allen, 1942
Otocinclus macrospilus Eigenmann & Allen, 1942: 201, pl. 13 (fig. 1). Type locality: Rio Morona [Upper Amazon
basin, Peru]. Holotype: CAS 77335; holotype illustrated in Schaefer (1997: fig. 42a).
Distribution: Amazon River basin, Peru (Schaefer, 2003c).
Otocinclus mariae Fowler, 1940
Otocinclus mariae Fowler, 1940c: 83, figs. 41–43. Type locality: Boca Chapare, Río Chimore [Río Mamoré system]
Bolivia. Holotype: ANSP 69068; holotype illustrated in Schaefer (1997: fig. 41a).
Distribution: Upper Madeira and lower Amazon River basins, Bolivia (Schaefer, 2003c).
Otocinclus mimulus Axenrot & Kullander, 2003
Otocinclus mimulus Axenrot & Kullander, 2003: 251, fig. 1. Type locality: Paraguay: Departamento Caaguazú: río
Paraná drainage, small stream (arroyo) at Estancia María Belén, 8 km from Colone Patricio Colman,
25º40'13"S, 55º5'52"W. Holotype: MNHNP uncat (currently as NRM 43480).
Distribution: Monday River, Paraná River basin, Paraguay (Axenrot & Kullander, 2003).
Otocinclus mura Schaefer, 1997
Otocinclus mura Schaefer, 1997: 77, fig. 47. Type locality: Rio Sanabani, munícXpio de Silves (Lago Canaçari drainage into Rio Solimões mainstem), ca. 2º51'40"S, 58º15'40"W, rio Solimões/Sanabani/Lago Canaçari drainage,
Brazil. Holotype: MZUSP 51101.
Distribution: Upper Amazon River basin, Brazil (Schaefer, 2003c).
Otocinclus tapirape Britto & Moreira, 2002
Otocinclus tapirape Britto & Moreira, 2002: 1064, fig. 1. Type locality: Brazil: Goiás, córrego Água Parada, tributary of Rio Pintado, 11 km west from Novo Planalto, rio Araguaia system, 13º7'S, 49º62'W. Holotype: MZUSP
73975.
Distribution: Upper and middle Araguaia River basin, Brazil (Schaefer, 2003c).
Otocinclus vestitus Cope, 1872
Otocinclus vestitus Cope, 1872a: 283, pl. 4 (figs. 2a–c). Type locality: Tributaries of the Ambyiacu [which empties
into the Amazon near to Pebas, in Eastern Equador, some distance east of the Napo]. Lectotype: ANSP 8283,
designated by, and illustrated in, Schaefer (1997: 46, fig. 29).
Distribution: Amazon and lower [sic, upper] Paraná River basins (Schaefer, 2003c).
Otocinclus vittatus Regan, 1904
Otocinclus vittatus Regan, 1904b: 267, pl. 15 (fig. 3). Type locality: Descalvados, Mato Grosso, Brazil. Lectotype:
BMNH 1895.5.17.83, designated by, and illustrated in, Schaefer (1997: 57, fig. 36a).
Rhinolepis paraguensis Borodin, 1927b: 7. Type locality: Paraguay. Holotype: AMNH 8641.
Distribution: Amazon, Orinoco, Paraná/Paraguay, Xingu and Tocantins River basins (Schaefer, 2003c).
Otocinclus xakriaba Schaefer, 1997
Otocinclus xakriaba Schaefer, 1997: 79, fig. 48. Type locality: Rio Pau de Colher, on road between Guanambi andMutas, 14º12'48"S, 42º49'29"W, rio São Francisco drainage, Bahia, Brazil. Holotype: MZUSP 51103.
Distribution: São Francisco River basin, Brazil (Schaefer, 2003c).
OTOTHYRIS Myers, 1927
Otothyris Myers, 1927: 128. Type species: Otothyris canaliferus Myers, 1927. Type by original designation. Gender: Feminine.

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Revision: Garavello et al. (1998).
Otothyris juquiae Garavello, Britski & Schaefer, 1998
Otothyris juquiae Garavello, Britski & Schaefer, 1998: 8, fig. 4. Type locality: Juquiá creek at Cantagalo farm, SãoPaulo State, Brazil. Holotype: MZUSP 10309.
Distribution: Ribeira de Iguape River basin, Brazil (Schaefer, 2003c).
Otothyris lophophanes (Eigenmann & Eigenmann, 1889)
Rhinelepis lophophanes Eigenmann & Eigenmann, 1889a: 9. Type locality: Santa Cruz, Brazil. Holotype: MCZ
8164.
Otocinclus cephalacanthus Miranda Ribeiro, 1911: 93. Type locality: Brazil. Holotype: MNRJ 1050.
Otothyris canaliferus Myers, 1927: 128. Type locality: Hills in the vicinity of Rio de Janeiro, Brazil. Lectotype:
MCZ 31577, designated in Garavello et al. (1998: 7).
Distribution: Atlantic coastal streams of Rio de Janeiro State, Brazil (Schaefer, 2003c).
Otothyris rostrata Garavello, Britski & Schaefer, 1998
Otothyris rostrata Garavello, Britski & Schaefer, 1998: 10, fig. 5. Type locality: Osório, Canal between lagoons
Emboaba and Emboabinha, on the highway between Osório and Tramandai, Rio Grande do Sul State, Brazil.
Holotype: MCP 14812.
Distribution: Atlantic coast of Santa Catarina and Rio Grande do Sul States, Brazil (Schaefer, 2003c).
Otothyris travassosi Garavello, Britski & Schaefer, 1998
Otothyris travassosi Garavello, Britski & Schaefer, 1998: 12, fig. 6. Type locality: Barra Sêca River at Linhares toSão Mateus Road, Espírito Santo State, Brazil. Holotype: MNRJ 10177.
Distribution: Atlantic coast of southeastern Brazil (Schaefer, 2003c).
OTOTHYROPSIS Ribeiro, Carvalho & Melo, 2005
Otothyropsis Ribeiro, Carvalho & Melo, 2005: 491. Type species: Otothyropsis marapoama Ribeiro, Carvalho &Melo, 2005, by original designation. Gender: Feminine.
Otothyropsis marapoama Ribeiro, Carvalho & Melo, 2005
Otothyropsis marapoama Ribeiro, Carvalho & Melo, 2005: 491, fig. 1. Type locality: Brazil, São Paulo, rio Tietê
basin, Município de Marapoama, ribeirão Cubatão, road between Marapoama and Elisiário (21º11'35"S,
49º07'22"W). Holotype: LIRP 5640.
Distribution: Tietê River, Paraná River basin, Brazil (Ribeiro et al., 2005).
OXYROPSIS Eigenmann & Eigenmann, 1889
Oxyropsis Eigenmann & Eigenmann, 1889b: 39. Type species: Oxyropsis wrightiana Eigenmann & Eigenmann,
1889. Type by original designation. Gender: Feminine.
Revision: Aquino & Schaefer (2002).
Oxyropsis acutirostris Miranda Ribeiro, 1951
Oxyropsis acutirostris Miranda Ribeiro, 1951a: 2, pl. 1 (fig. 2), pl. 2 (fig. 2). pl. 3 (fig. 2). Type locality: Paricachoeira, rio Taquié, afl. do Vaupes que é afl. do Negro, Amazonas [Brazil]. Holotype: MNRJ 5745.
Distribution: Orinoco and Negro River basins (Schaefer, 2003c).
Oxyropsis carinata (Steindachner, 1879)
Hypoptopoma carinatum Steindachner, 1879e: 48, pl. 4 (fig. 3). Type locality: Einem nebenflusse des Amazonenstromes an der Peruanische Grenze. Holotype: NMW 46267; holotype illustrated in Aquino & Schaefer (2002:
fig. 6).
Distribution: Middle and upper Amazon River basin (Schaefer, 2003c).
Oxyropsis wrightiana Eigenmann & Eigenmann, 1889
Oxyropsis wrightiana Eigenmann & Eigenmann, 1889b: 39. Type locality: Lago Hyanuary [Brazil]. Holotype: MCZ
8055; holotype illustrated in Aquino & Schaefer (2002: fig. 4).
Oxyropsis wrightii Eigenmann & Eigenmann, 1891: 40. Type locality: Lago Hyanuary. Holotype: MCZ 8055.
Unjustified emendation of Oxyropsis wrightiana.
Distribution: Middle and upper Amazon River basin (Schaefer, 2003c).

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FERRARISPANAQOLUS Isbrücker & Schraml, 2001
Panaqolus Isbrücker & Schraml, in Isbrücker et al., 2001: 20. Type species: Panaque gnomus Schaefer & Stewart,
1993. Type by original designation. Gender: Masculine.
Revision: Schaefer & Stewart (1993) as Panaque dentex species group.
Species key: Chockley & Armbruster (2002), as Panaque dentex species group.
Remarks: This genus corresponds to the monophyletic assembledge called the Panaque dentex species group by
Schaefer & Stewart (1993). The recognition of this group as a separate genus was rejected by Chockley & Armbruster (2002) and Weber (2003).
Panaqolus albomaculatus (Kanazawa, 1958)
Panaque albomaculatus Kanazawa, 1958: 327, fig. 1 (top). Type locality: Ecuador, tributary of the Rio Suno, upper
Napo River, 0º47' south latitude, 77º16' west longitude, Rio Pucuno. Holotype: USNM 167909.
Distribution: Headwaters of Napo, Marañon and Ucayali rivers, Ecuador, Peru (Fisch-Muller, 2003).
Remarks: Redescribed by Schaefer & Stewart (1993: 327, fig. 13).
Panaqolus changae (Chockley & Armbruster, 2002)
Panaque changae Chockley & Armbruster, 2002: 83, fig. 1 (middle). Type locality: Peru: Loreto: río Itaya, río Amazonas drainage, 11 km SSW center of Iquitos at bearing 39º, 3º49'47.6"S 73º18'2.9"W. Holotype: MUSM
17107.
Distribution: Itaya and Momon River basins, Peru (Fisch-Muller, 2003).
Panaqolus dentex (Günther, 1868)
Chaetostomus dentex Günther, 1868a: 477. Type locality: Xeberos. Holotype: BMNH 1867.6.13.37. Described in
more detail in Günther (1868b: 233).
Distribution: Napo, Pastaza and Marañon River basins (Fisch-Muller, 2003).
Remarks: Redescribed by Schaefer & Stewart (1993: 321, fig. 9), with type locality given as: Peru, Departamento
Loreto, Xeberos (or Jeberos), upper Río Aipena system, tributary to Río Huallaga near its confluence with Río
Marañon, approx. 5º18'S 76º17'W.
Panaqolus gnomus (Schaefer & Stewart, 1993)
Panaque gnomus Schaefer & Stewart, 1993: 333, fig. 17 (top). Type locality: Ecuador, Morona-Santiago Province,
Río Cushuimi at Cushuimi, approx. 2º39'S 77º43'W, just over 300 m alt., upper Río Morona/ Marañon basin.
Holotype: FMNH 70860.
Distribution: Pastaza and upper Marañon basins (Fisch-Muller, 2003).
Panaqolus maccus (Schaefer & Stewart, 1993)
Panaque maccus Schaefer & Stewart, 1993: 335, figs. 18, 19. Type locality: Venezuela, Estado Portuguesa, Río Las
Marinas, upstream from bridge on Route 5 east of Guanare, tributary of Río Portuguesa, Río Orinoco basin,
approx. 9º05'N 69º40'W. Holotype: MBUCV V-24010.
Distribution: Apuré and Caroni River basins, Venezuela (Fisch-Muller, 2003).
Panaqolus nocturnus (Schaefer & Stewart, 1993)
Panaque nocturnus Schaefer & Stewart, 1993: 330, fig. 15. Type locality: Peru, Departamento Amazonas, Río Santiago, near la Poza, Río Marañon basin, 4º01'24"S 77º45'06"W. Holotype: LACM 41729-51.
Distribution: Santiago and Pastaza River basins, upper Napo drainage (Fisch-Muller, 2003).
Panaqolus purusiensis (La Monte, 1935)
Panaque purusiensis La Monte, 1935: 4, fig. 3. Type locality: Rio Purus, Brazil; in the vicinity of the mouth of Rio
Macauhan [=Macauá], a tributary of Rio Yaco [=Iaco], which, in turn, is a tributary of Rio Purus (69ºW, 9º20'S).
Holotype: AMNH 12600; holotype illustrated in Schaefer & Stewart (1991: fig. 12).
Distribution: Macauá River basin, upper Purus drainage, Brazil (Fisch-Muller, 2003).
PANAQUE Eigenmann & Eigenmann, 1889
Panaque Eigenmann & Eigenmann, 1889b: 44. Type species: Chaetostomus nigrolineatus Peters, 1877. Type byoriginal designation. Gender: Masculine.
Panaque cochliodon (Steindachner, 1879)

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Chaetostomus cochliodon Steindachner, 1879c: 194. Type locality: Aus dem Cauca, dem grössten Nebenflusse desMagdalenen-Stromes [Colombia]. Syntypes: NMW 47297 (1), NMW 47298 (1). Also described as new in
Steindachner (1879g: 187); described in more detail and illustrated in Steindachner (1880b: 63, pl. 4).
Chaetostomus gibbosus Steindachner, 1880b: 63, pl. 4. Type locality: Aus dem Cauca, dem grössten Nebenflusse
des Magdalenen-Stromes [Colombia]. Syntypes: NMW 47297 (1), NMW 47298 (1). Alternate name for Chaetostomus cochliodon proposed as Chaetostomus cochliodon (sive gibbosus).
Distribution: Cauca and Magdalena River basins, Colombia (Fisch-Muller, 2003).
Panaque nigrolineatus (Peters, 1877)
Chaetostomus nigrolineatus Peters, 1877: 471, fig. on an unnumb. pl. ff. p. 556. Type locality: Calabozo [Guaricó
River], Venezuela. Syntypes: ZMB 10046 (1), ZMB 10047 (1).
Distribution: Orinoco River and middle and lower Amazon River tributaries (Fisch-Muller, 2003).
Panaque suttonorum Schultz, 1944
Panaque suttoni Schultz, 1944c: 308, pl. 10 (fig. b). Type locality: Río Negro below the mouth of the Río Yasa,
Maracaibo basin, Venezuela. Holotype: USNM 121033.
Distribution: Western and eastern tributaries of Maracaibo Lake, Venezuela (Fisch-Muller, 2003).
Remarks: Schultz (1944c: 311) indicated that the species was to be named in honor of “Dr. and Mrs. Fredrick A. Sutton,” which prompted Isbrucker (2001: 30) to correct the name to suttonorum.
PARALORICARIA Isbrücker, 1979
Paraloricaria Isbrücker, 1979a: 87, 90. Type species: Loricaria vetula Valenciennes, 1836. Type by original designation. Gender: Feminine.
Paraloricaria agastor Isbrücker, 1979
Paraloricaria agastor Isbrücker, 1979a: 103, figs. 20 (right) –21 (right). Type locality: Paraguay (rio ?) Paraguay.
Holotype: BMNH 1895.5.17.108.
Distribution: Paraguay River basin, Paraguay (Ferraris, 2003c).
Paraloricaria commersonoides (Devincenzi, 1943)
Loricaria commersonoides Devincenzi, 1943: 3, fig. 2. Type locality: río Uruguay, frente a la ciudad de Paysandú[Uruguay]. Holotype: at MHNM (apparently lost, Olazarri et al., 1970).
Distribution: Uruguay River basin (Ferraris, 2003c).
Remarks: Redescribed in Ringuelet et al. (1967).
Paraloricaria vetula (Valenciennes, 1836)
Loricaria vetula Valenciennes, 1836, in Valenciennes, 1835–47: pl. 6 (fig. 2). Type locality: [Not stated]. Holotype:
MNHN a-8996. Described in Cuvier & Valenciennes (1840b: 466, 344 in Strasbourg deluxe edition) with locality stated as: des environs de Buénos-Ayres.
Distribution: La Plata River basin (Ferraris, 2003c).
Remarks: Reviewed in Ringuelet et al. (1967).
PARANCISTRUS Bleeker, 1862
Parancistrus Bleeker, 1862 (in Bleeker, 1862–63): 2. Type species: Hypostomus aurantiacus Castelnau, 1855. Typeby original designation. Gender: Masculine.
Acanthodemus Marschall, 1873: 63. Type species: Hypostomus aurantiacus Castelnau, 1855. Type by subsequent
designation by Jordan (1919a: 264). Gender: Masculine.
Monophyly: Rapp Py-Daniel (1989).
Parancistrus aurantiacus (Castelnau, 1855)
Hypostomus aurantiacus Castelnau, 1855: 43, pl. 21 (fig. 2). Type locality: l’Ucayale. Holotype: MNHN a-9452(dry). Name spelled Hypostomus aurentiacus on figure caption. Eschmeyer, et al. (1998: 165) acted as firstreviser in selecting the spelling H. aurantiacus over H. aurentiacus.
Hypostomus nigricans Castelnau, 1855: 44, pl. 22 (fig. 1). Type locality: l’Amazone. Holotype: MNHN a-9576.
Hypostomus vicinus Castelnau, 1855: 45, pl. 23 (fig. 1). Type locality: l’Ucayale. Holotype: MNHN a-9572.
Distribution: Ucayali, Tocantins and Xingu Rivers (Fisch-Muller, 2003).

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Remarks: Redescribed by Rapp Py-Daniel (1989: 239), in which the three names are placed together in synonymy
for the first time. Therefore, Rapp Py-Daniel acted as first reviser in selecting H. aurantiacus as valid.
Parancistrus nudiventris Rapp Py-Daniel & Zuanon, 2005
Parancistrus nudiventris Rapp Py-Daniel & Zuanon, 2005: 573, fig. 1. Type locality: Brazil; Pará State; rio Xingu,
bedrock at ilha do Bacabal, 3º23'19" S, 51º43'24" W. Holotype: INPA 15037.
Distribution: Xingu River, Brazil (Rapp Py-Daniel & Zuanon, 2005).
PAREIORHAPHIS Miranda Ribeiro, 1918
Pareiorhaphis Miranda Ribeiro, 1918a: 106. Type species: Hemipsilichthys calmoni [= Psilichthys cameroni Steindachner, 1907]. Type by subsequent designation by Regan (1920b: 14). Gender: Feminine. Type designation
made with the unjustified emended name of Hemipsilichthys calmoni, and, erroneously, with Ribeiro as author.
Subsequent type designation by Gosline (1947: 102) of Hemipsilichthys duseni is invalid.
Revision and key to species: Pereira & Reis (2002) as Hemipsilichthys, in part.
Pareiorhaphis azygolechis (Pereira & Reis, 2002)
Hemipsilichthys azygolechis Pereira & Reis, 2002: 129, fig. 20 (middle). Type locality: Brazil: Paraná: Guaratuba;
rio Araraquara ca. 8 km N of Guaruva (25º57'58"S 48º49'43"W). Holotype: MCP 19670.
Distribution: São João River basin, Brazil (Weber, 2003).
Pareiorhaphis bahianus (Gosline, 1947)
Corymbophanes bahianus Gosline, 1947: 107, pl. 3 (fig. 6). Type locality: Fazenda Almada, Ilheus, Bahia [Brazil].
Holotype: MNRJ 4243.
Distribution: Small drainages near Ilhéus, between Pado and Contas River basins, Brazil (Weber, 2003).
Pareiorhaphis cameroni (Steindachner, 1907)
Psilichthys cameroni Steindachner, 1907a: 82. Type locality: aus dem Fluss Cubataõ im Staate Santa Catharina, Brazilien. Lectotype: ZSM 4837, designated by Pereira & Reis (2002: 104, 106). Illustrated and described in moredetail in Steindachner (1907e: 475, pls. 1, 2) as Hemipsilichthys cameroni.
Hemipsilichthys calmoni Steindachner, 1908a: 31. Emendation of Psilichthys cameroni [see Remarks].
Distribution: Cubatão (southern), Tubarão and Araranguá River basins, Brazil (Weber, 2003).
Remarks: Steindachner (1908a: 31) proposed to correct the name cameroni to calmoni in order to reflect the correctspelling of the patronym which was stated in Steindachner (1907c: 418) to honor “Dr. Miguel Calmon.” However, the spelling cameroni appears in the minutes of the meeting, which constitutes the original proposal of thename (Steindachner, 1907a) as well as the invariant spelling in the text and plate captions of the subsequentdescription (Steindachner, 1907e). Despite this, Pereira & Reis (2002: 104, 106), and Weber (2003: 353) treat
cameroni as an incorrect spelling. Species redescribed, as Hemipsilichthys calmoni, in Steindachner (1910d:
430).
Pareiorhaphis cerosus (Miranda Ribeiro, 1951)
Hemipsilichthys cerosus A. Miranda Ribeiro, in P. Miranda Ribeiro, 1951c: xxxix. Type locality: Not explicitly
given. Lectotype: MNRJ 648. Lectotype illustrated in, and designated, by Pereira & Reis (2002: 115, fig. 11).
Earlier lectotype proposal in Miranda Ribeiro (1953: 400) considered to be invalid by Pereira & Reis (2002:
117) and Weber (2003).
Distribution: Eastern Brazilian coastal basin; known only from two type specimens (Weber, 2003).
Pareiorhaphis eurycephalus (Pereira & Reis, 2002)
Hemipsilichthys eurycephalus Pereira & Reis, 2002: 131, fig. 21 (middle). Type locality: Brazil: Santa Catarina:
Urubici: creek tributary of Rio Canoas East of vila São José, on road to serra do Corvo Branco (28º03'33"S49º25'42"W). Holotype: MCP 19778.
Distribution: Upper Canoas River basin, Brazil (Weber, 2003).
Pareiorhaphis garbei (Ihering, 1911)
Hemipsilichthys garbei Ihering, 1911: 399. Type locality: no curso superior, montanha acima, do Rio Macahé, noEst. do Rio de Janeiro [...] im Staat Rio de Janiero, im Gebirgeflusse Macahé (ca. 10–15m. Breite, flach und
Steinig) im Oberlaufe, d.h. am Gebirgsrande [Brazil]. Lectotype: MZUSP 1163, designated by, and illustratedin, Oliveira (1997: 117, fig. 1).

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Distribution: Macaé River basin, Brazil (Weber, 2003).
Remarks: Redescribed in Oliveira (1997).
Pareiorhaphis hypselurus (Pereira & Reis, 2002)
Hemipsilichthys hypselurus Pereira & Reis, 2002: 136, fig. 23 (middle). Type locality: Brazil: Rio Grande do Sul:
Maquiné; rio Forqueta tributary to rio Maquiné, 29º32'42"S 50º14'21"W. Holotype MCP 19665.
Distribution: Maquiné, Três Forquillas, Mampituba and Araranguá River basins, Brazil (Weber, 2003).
Pareiorhaphis hystrix (Pereira & Reis, 2002)
Hemipsilichthys hystrix Pereira & Reis, 2002: 134, fig. 22 (middle). Type locality: Brazil: Rio Grande do Sul: Bom
Jesus; creek tributary of rio dos Touros, on road from Silveira to Rondinha, 28º39'18"S 50º18'25"W. Holotype
MCP 19779.
Distribution: Headwaters of Pelotas, Ijui and Piratinim River basins; upper Taquari River basin, Brazil (Weber,2003).
Pareiorhaphis mutuca (Oliveira & Oyakawa, 1999)
Hemipsilichthys mutuca Oliveira & Oyakawa, 1999: 75, fig. 1 (top). Type locality: Brazil: Minas Gerais: Rio das
Velhas drainage, County of Nova Lima, Córrego Mutuca, at right side of the road from Belo Horizonte to Nova
Lima, km 20 (approx. 20º60'S, 43º55'W). Holotype: MZUSP 37148.
Distribution: Headwaters of Das Velhas River basin, Brazil (Weber, 2003).
Pareiorhaphis nudulus (Reis & Pereira, 1999)
Hemipsilichthys nudulus Reis & Pereira, 1999: 46, fig. 2 (top). Type locality: Rio Jordão, at Jordão Alto (approx.
28º35'S, 49º27'W), Nova Veneza, Santa Catarina, Brazil. Holotype: MCP 20278.
Distribution: Araranguá, Mapituba, Três Forquilhas, and Maquiné River basins, Brazil (Weber, 2003).
Pareiorhaphis parmula Pereira, 2005
Pareiorhaphis parmula Pereira, 2005: 272, fig. 1. Type locality: Brazil: Paraná: Lapa: rio Iguaçu basin, rio dos
Patos, tributary to rio da Várzea, on road PR-427 from Lapa to Campo do Tenente, 25º50'36.8"S049º43'39.2"W. Holotype: MCP 35826.
Distribution: Dos Patos River, tributary of Iguaçu River, Parana River basin, Brazil (Pereira, 2005).
Pareiorhaphis regani (Giltay, 1936)
Hemipsilichthys regani Giltay, 1936: 6, fig. 2. Type locality: Rio Curi Curiay, au S. du Rio Negro (Amazone, Brésil).
Holotype: IRSNB 47.
Distribution: Curicuriari and possibly Jaú River basins in the Negro River drainage, Brazil (Weber, 2003).
Pareiorhaphis splendens (Bizerril, 1995)
Hemipsilichthys splendens Bizerril, 1995a: 118, fig. 2. Type locality: Brazil: State of Santa Catarina, river Cubatão:near to the city of Joinville. Holotype: MNRJ 13325.
Distribution: Cubatão, Itajaí-Açu, Cubatão, Nhundiaquara and São João River basins, Brazil (Weber, 2003).
Pareiorhaphis steindachneri (Miranda Ribeiro, 1918)
Hemipsilichthys steindachneri Miranda Ribeiro, 1918a: 107. Type locality: Flum. Rio Itapucú (Santa Catharina)[Brazil]. Lectotype: NMW 92790, designated by Pereira & Reis (2002: 109). Originally proposed as H. steindachneri, but included in a new genus Pareiorhaphis.
Pseudancistrus luderwaldti Miranda Ribeiro 1918d: 632. Type locality: Hansa, Santa Catharina, Brasil. Holotype:
MZUSP 2109.
Distribution: Cubatão, Itapocu, and Itajaí-Açu River basins, Brazil (Weber, 2003).
Remarks: Priority of Hemipsilichthys steindachneri over Pseudancistrus luderwaldti has not been clearly established and may not be resolved by dates of publication; precedence in the absence of a clear indiction of priority
may be by first reviser action of Pereira & Reis (2002), which is tentatively followed here.
Pareiorhaphis stephanus (Oliveira & Oyakawa, 1999)
Hemipsilichthys stephanus Oliveira & Oyakawa, 1999: 77, fig. 4 (top). Type locality: Brazil: Minas Gerais: Rio
Jequitinhonha drainage, County of Diamantina, ribeirão das Pedras, ca. 1300 masl, 3 km north of Diamantina(approx. 18º10'S, 43º37'W). Holotype: MZUSP 36971.
Distribution: Jequitinhonha River basin, Brazil (Weber, 2003).
Pareiorhaphis stomias (Pereira & Reis, 2002)

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rio Mãe Luzia at Forquilha, 28º27'66"S 49º30'06"W. Holotype MCP 19666.
Distribution: Ararangua and Tubarão River basins, Brazil (Weber, 2003).
Pareiorhaphis vestigipinnis (Pereira & Reis, 1992)
Hemipsilichthys vestigipinnis Pereira & Reis, 1992: 113, fig. 2(b). Type locality: Creek tributary of the Rio Caveirasat Painel, on the road from São Joaquim to Lares (approximately 27º55'S, 50º05'W) Lajes, Santa Catarina, Brazil. Holotype: MCP 14344.
Distribution: Headwaters of Canoas River basin, Brazil (Weber, 2003).
PAREIORHINA Gosline, 1947
Pareiorhina Gosline, 1947: 104. Type species: Rhinelepis rudolphi Miranda Ribeiro, 1911. Type by original designation. Gender: Feminine.
Key to species: Chamon et al. (2005: 555).
Pareiorhina brachyrhyncha Chamon, Aranda & Buckup, 2005
Pareiorhina brachyrhyncha Chamon, Aranda & Buckup, 2005: 551, fig. 1. Type locality: Brazil, State of São Paulo,
Município Pindamonhangaba, Bairro Ribeirão Grande (or Teteqüera) and adjacent tributary ditch (Rio Paraíba
do Sul drainage), Vera Cruz farm, 22º46'14.1"S, 45º26'42.5"W. Holotype: MNRJ 25033.
Distribution: Paraíba do Sul River basin, São Paulo State, Brazil (Chamon et al., 2005).
Pareiorhina carrancas Bockmann & Ribeiro, 2003
Pareiorhina carrancas Bockmann & Ribeiro, 2003: 232, fig. 1. Type locality: Brazil, Minas Gerais: minicípio of
Carrancas, córrego Debaixo da Serra (an upper course tributary of córrego do Beijinho, a left margin affluent of
rio Grande at Reservoir of Camargos), in piedmont of Serra de Carrancas (44º36'08.7"W, 21º26'39.7"S). Holotype: LIRP 2280.
Distribution: Upper Rio Grande basin, of Upper Parana River system, Brazil (Bockmann & Ribeiro, 2003).
Pareiorhina rudolphi (Miranda Ribeiro, 1911)
Plecostomus (Rhinelepis) microps Ihering, 1907: 24. Type locality: Piquete (Lorena), Estado de São Paulo [Brazil].
Holotype: MZUSP 2154. Previously unpublished illustration of holotype reproduced in Bockmann & Ribeiro(2003: fig. 6). Preoccupied by Plecostomus microps Steindachner, 1876; replaced by Rhinelepis rudolphi
Miranda Ribeiro, 1911.
Rhinelepis rudolphi Miranda Ribeiro, 1911: 84. Type locality: Piquete (Lorena), Estado de São Paulo [Brazil]. Holotype: MZUSP 2154. Replacement name for Plecostomus microps Ihering, 1907.
Distribution: Paraíba do Sul River near Lorena, São Paulo State, Brazil (Weber, 2003).
PAROTOCINCLUS Eigenmann & Eigenmann, 1889
Parotocinclus Eigenmann & Eigenmann, 1889b: 41. Type species: Otocinclus maculicauda Steindachner, 1877.
Type by monotypy. Gender: Masculine. Proposed originally as a subgenus of Hisonotus.
Revision: Garavello (1976).
Reviews: Schaefer & Provenzano (1993, Guiana Shield); Lacerda & Evers (1996, Brazil).
Key: Garavello & Britski (2003, eastern Brazilian coastal rivers).
Parotocinclus amazonensis Garavello, 1976
Parotocinclus amazonensis Garavello, 1976: 7, figs. 21–22. Type locality: Ilha Sorubim, Rio Solimões, Amazonas,
Brazil. Holotype: MZUSP 10145.
Distribution: Amazon River basin, Brazil (Schaefer, 2003c).
Parotocinclus aripuanensis Garavello, 1988
Parotocinclus aripuanensis Garavello, 1988: 122, fig. 3. Type locality: Ingazeiro, 20 km upstream of Boca do Rio
Canumã, Aripuanã, MT, Brazil. Holotype: MZUSP 36899.
Distribution: Aripuanã River basin, Brazil (Schaefer, 2003c).
Parotocinclus bahiensis (Miranda Ribeiro, 1918)
Microlepidogaster bahiensis Miranda Ribeiro, 1918d: 635. Type locality: Villa Nova, E. da Bahia [Brazil]. Syntypes: MZUSP 1071, lost (Britski, 1969).

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Distribution: Bahia State, Brazil (Schaefer, 2003c).
Parotocinclus bidentatus Gauger & Buckup, 2005
Parotocinclus bidentatus Gauger & Buckup, 2005: 512, fig. 5. Type locality: rio Pirapetinga (left bank tributary of
rio Paraíba do Sul), Resende (22º28'08"S 44º26'48"W), Rio de Janeiro State, Brazil. Holotype: MNRJ 27962.
Distribution: Pirapetinga and Calçado Rivers, Paraíba do Sul River basin, Rio de Janeiro, Brazil (Gauger & Buckup,2005).
Parotocinclus britskii Boeseman, 1974
Parotocinclus britskii Boeseman, 1974: 267, fig. 2; pl. 2. Type locality: Left tributary of Coppename River, Surinam, 3º51'N, 56º55'W. Holotype: ZMA 106593.
Distribution: Orinoco River basin and Atlantic coastal drainages of the Guianas (Schaefer, 2003c).
Parotocinclus cearensis Garavello, 1976
Parotocinclus cearensis Garavello, 1976: 14, figs. 19–20. Type locality: Cachoeira do Gusmão, Ipu, Ceará, Brazil.
Holotype: MNRJ 10176.
Distribution: Ceará State, Brazil (Schaefer, 2003c).
Parotocinclus cesarpintoi Miranda Ribeiro, 1939
Paraotocinclus cesarpintoi Miranda Ribeiro, P., 1939b: 364, fig. Type locality: Vila de Quebrângulo, no Estado deAlagôas [Brazil]. Syntypes (2, 45–55 mm TL): possibly MNRJ 1022 (2 of 7), or MNRJ 1024 (2).
Distribution: Paraíba do Sul River basin, Brazil (Schaefer, 2003c).
Parotocinclus collinsae Schmidt & Ferraris, 1985
Parotocinclus collinsae Schmidt & Ferraris, 1985: 341, figs. 1–3. Type locality: Tributary to Takutu River, ca. 2 mi.
from Mazarhally Takutu Lumber Camp in Takutu mountains, 6º15'N, 59º5'W, Essequibo, Guyana. Holotype:
AMNH 55433.
Distribution: Essequibo River basin, Guyana (Schaefer, 2003c).
Parotocinclus cristatus Garavello, 1976
Parotocinclus cristatus Garavello, 1976: 6, figs. 9–10. Type locality: Fazenda Almada, Ilhéus, Bahia, Brazil. Holotype: MNRJ 10132.
Distribution: Coastal rivers near Ilhéus, Bahia State, Brazil (Schaefer, 2003c).
Parotocinclus doceanus (Miranda Ribeiro, 1918)
Microlepidogaster doceanus Miranda Ribeiro, 1918d: 634. Type locality: Rio Doce, E. Santo [Brazil]. Holotype:
MZUSP 1016.
Distribution: Doce River basin, Brazil (Schaefer, 2003c).
Parotocinclus eppleyi Schaefer & Provenzano, 1993
Parotocinclus eppleyi Schaefer & Provenzano, 1993: 46, figs. 5–6. Type locality: Caño Curicurito, ca. 1 km aboveits mouth into the Río Autana, Amazonas, Venezuela, 4º47'N, 67º25'W. Holotype: MBUCV V-22530.
Distribution: Upper and middle Orinoco River basin, Venezuela (Schaefer, 2003c).
Parotocinclus haroldoi Garavello, 1988
Parotocinclus haroldoi Garavello, 1988: 118, fig. 1. Type locality: Córrego do Otaviano, Poço do Sanharó, Riacho
Sanharó, Piauí, Brazil. Holotype: MNRJ 10531.
Distribution: Piauí State, Brazil (Schaefer, 2003c).
Parotocinclus jimi Garavello, 1976
Parotocinclus jimi Garavello, 1976: 9, figs. 25–26. Type locality: Rio do Peixe, small tributary of Rio de Contas,
Fazenda Pedra Branca, Itagibá, Bahia, Brazil. Holotype: MZUSP 12133.
Distribution: Bahia State, Brazil (Schaefer, 2003c).
Parotocinclus jumbo Britski & Garavello, 2002
Parotocinclus jumbo Britski & Garavello, 2002: 280, fig. 1. Type locality: Brazil: Paraíba: rio Paraíba do Norte, atbridge on road PB 408, NW of Umbuzeiro (7 38'27"S 35 42'30"W). Holotype: MZUSP 69513.
Distribution: Coastal rivers in Ceará, Paraíba, Pernambuco and Alagoas States, Brazil (Schaefer, 2003c).
Parotocinclus longirostris Garavello, 1988
Parotocinclus longirostris Garavello, 1988: 120, fig. 2. Type locality: Rio Preto da Eva, Manaus-Itacoatiara highway, km 80, Manaus, Amazonas, Brazil. Holotype: MZUSP 36891.

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Parotocinclus maculicauda (Steindachner, 1877)
Otocinclus maculicauda Steindachner, 1877c: 222, pl. 1 (figs. 2–2b). Type locality: Santa Cruz, Rio Quenda in
vicinity of Rio de Janeiro, Brazil. Lectotype: NMW 45381: 1, designated by, and illustrated in, Schaefer (1996b:
1034, fig. 2).
Parotocinclus steindachneri Di Caporiacco, 1948: 201, fig. 2. Type locality: Rio Novo, Santa Catarina, Brazil. Syntypes: MZUF 5565 (1).
Distribution: Coastal rivers between Santa Catarina and Rio de Janeiro States, Brazil (Schaefer, 2003c).
Parotocinclus minutus Garavello, 1976
Parotocinclus minutus Garavello, 1976: 15, figs. 13–14. Type locality: Rio Vasa-Barris, Canudos, Bahia, Brazil.
Holotype: MNRJ 10135.
Distribution: Vasa-Barris River basin, Brazil (Schaefer, 2003c).
Parotocinclus muriaensis Gauger & Buckup, 2005
Parotocinclus muriaensis Gauger & Buckup, 2005: 514, fig. 7. Type locality: rio Muriaé river (left bank tributary of
rio Paraíba do Sul), approx. 21º15'S 41º45'W, Município de Itaperuna, Rio de Janeiro State, Brazil. Holotype:
MNRJ 28528.
Distribution: Itaperuna River, Paraiba do Sul River basin, Rio de Janeiro, Brazil (Gauger & Buckup, 2005).
Parotocinclus planicauda Garavello & Britski, 2003
Parotocinclus planicauda Garavello & Britski, 2003: 254, fig. 1. Type locality: Brazil. Minas Gerais, Coroaci,
Suaçui Pequeno River, barro do Rochedo, at Antonio Pereira de Oliveira farm, 20 km from Coroaci. Holotype:
MZUSP 75071.
Distribution: Doce river basin, Minas Gerais, Brazil (Garavello & Britski, 2003).
Parotocinclus polyochrus Schaefer, 1988
Parotocinclus polyochrus Schaefer, 1988: 184, figs. 1–2. Type locality: Río Mawarinuma tributary at Neblina basecamp, Terr. Federal Amazonas, Venezuela, 0º55'N, 66º10'W, elev. 120 m. Holotype: AMNH 74482.
Distribution: Baria River, Upper Negro River basin, Venezuela (Schaefer, 2003c).
Parotocinclus prata Ribeiro, Melo & Pereira, 2002
Parotocinclus prata Ribeiro, Melo & Pereira, 2002: 218, fig. 1. Type locality: Brazil: Minas Gerais: rio São Francisco basin, município de Presidente Olegário, headwaters of rio da Prata, stream tributary of ribeirão Quiricó,
fazenda São Zeferino, near Galena; approximately 18º22'S 46º14.3'W. Holotype: MZUSP 68359.
Distribution: Upper São Francisco River basin, Brazil (Schaefer, 2003c).
Parotocinclus spilosoma (Fowler, 1941)
Plecostomus spilosoma Fowler, 1941a: 152, figs. 53–55. Type locality: Campina Grande, Parahyba, Brazil. Holotype: ANSP 69410.
Distribution: Paraíba State, Brazil (Schaefer, 2003c).
Parotocinclus spilurus (Fowler, 1941)
Plecostomus spilurus Fowler, 1941a: 148, figs. 47–49. Type locality: Rio Salgade, Icó, Ceará, Brazil. Holotype:
ANSP 69403.
Distribution: Ceará State, Brazil (Schaefer, 2003c).
PECKOLTIA Miranda Ribeiro, 1912
Peckoltia Miranda Ribeiro, 1912: 7. Type species: Chaetostomus vittatus Steindachner, 1881. Type by monotypy.
Gender: Feminine.
Peckoltichthys Miranda Ribeiro, 1917: 49. Type species: Peckoltichthys filicaudatus Miranda Ribeiro, 1917. Typeby monotypy. Gender: Masculine.
Ancistomus Isbrücker & Seidel, in Isbrücker et al., 2001: 17. Type species: Ancistrus snethlageae Steindachner,
1911. Type by original designation. Gender: Masculine.
Sophiancistrus Isbrücker & Seidel, in Isbrücker et al., 2001: 21. Type species: Hemiancistrus ucayalensis Fowler,
1940. Type by original designation. Gender: Masculine.
Remarks: Miranda Ribeiro submitted a manuscript for publication in which Peckoltichthys was proposed as a

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replacement for Peckoltia, which he considered to be preoccupied by an identical generic name in Botany. The
publication of that manuscript was delayed until1920 (Miranda Ribeiro, 1920) and was preceded by Miranda
Ribeiro (1917), in which Peckoltichthys was published with only one included species, P. filicaudatus, and no
indication that the name was intended as a replacement. Therefore, Peckoltichthys must be considered to havebeen proposed with P. filicaudatus as type, by monotypy and not, as indicated in Miranda Ribeiro (1920) as areplacement name with Chaetostomus vittatus as type. The newly proposed generic names Sophiancistrus
Isbrücker & Seidel and Ancistomus Isbrücker & Seidel were considered by Fisch-Muller (2003) to be unwarranted.
Peckoltia arenaria (Eigenmann & Allen, 1942)
Hemiancistrus arenarius Eigenmann & Allen, 1942: 185, pl. 6 (fig. 2). Type locality: Yurimaguas. Holotype: CAS
77323.
Distribution: Upper Amazon River basin, Peru (Fisch-Muller, 2003).
Peckoltia bachi (Boulenger, 1898)
Chaetostomus bachi Boulenger, 1898e: 425, pl. 41 (fig. 1). Type locality: Rio Jurua, an affluent of the Amazons,Brazil. Holotype: BMNH 1897.12.1.61.
Distribution: Juruá River basin, Brazil (Fisch-Muller, 2003).
Peckoltia braueri (Eigenmann, 1912)
Hemiancistrus braueri Eigenmann, 1912b: 232, pl. 28 (fig. 1). Type locality: Takutu, British Guiana. Holotype:
ZMB 3174 (larger specimen).
Distribution: Upper Negro and Branco River basins; Brazil and Guyana (Armbruster & Werneke, 2005).
Remarks: Redescribed in Armbruster & Werneke (2005:2).
Peckoltia brevis (La Monte, 1935)
Hemiancistrus brevis La Monte, 1935: 3, fig. 2. Type locality: Rio Purus, Brazil; in the vicinity of the mouth of Rio
Macauhan, a tributary of Rio Yaco, which, in turn, is a tributary of Rio Purus (69ºW., 9º20'S.). Holotype:
AMNH 12602.
Distribution: Purus River basin, Brazil (Fisch-Muller, 2003).
Peckoltia cavatica Armbruster & Werneke, 2005
Peckoltia cavatica Armbruster & Werneke, 2005: 8, figs. 2 (bottom), 4. Type locality: Guyana, Rupununi (Region9), 3.7 km SSE Massara, 03.86228º, –059.28439º. Holotype: UG/CSBD 11043.
Distribution: Upper Rupununi River, Guyana (Armbruster & Werneke, 2005).
Peckoltia filicaudata (Miranda Ribeiro, 1917)
Peckoltichthys filicaudatus Miranda Ribeiro, 1917: 49. Type locality: Fluvio Solimões [Brazil]. Holotype: MNRJ
969.
Distribution: Upper Amazon River basin, Brazil; known only from holotype (Fisch-Muller, 2003).
Peckoltia furcata (Fowler, 1940)
Chaetostomus furcatus Fowler, 1940a: 238, figs. 28–29. Type locality: Ucayali River basin, Contamana, Peru. Holotype: ANSP 68655.
Distribution: Ucayali River basin, Peru; known only from holotype (Fisch-Muller, 2003).
Peckoltia kuhlmanni (Miranda Ribeiro, 1920)
Peckoltichthys kuhlmanni Miranda Ribeiro, 1920: 10, pl. 5 (middle). Type locality: Tapajóz [Brazil]. Lectotype:
MNRJ 2044A, designated by Miranda Ribeiro (1953: 401), but specimen not isolated.
Distribution: Tapajós River basin, Brazil (Fisch-Muller, 2003).
Peckoltia oligospila (Günther, 1864)
Chaetostomus oligospilus Günther, 1864: 244. Type locality: River Capin [Brazil]. Holotype: BMNH 1849.11.8,
illustrated in Regan (1904b: 232, pl. 12 fig. 1).
Distribution: Lower Amazon River basin, Brazil (Fisch-Muller, 2003).
Peckoltia sabaji Armbruster, 2003
Peckoltia sabaji Armbruster, 2003b: 6, figs. 2–4. Type locality: Guyana: Rupununi (Region 9), Essequibo River Dr.,
Rupununi R. 5.9 km WSW village of Sand Creek, 02.96656º, –059.56943º. Holotype: UG/CSBD 11041.
Distribution: Rupununi, Essequibo and Takutu River basins, Guyana, and Casiquiare and Orinoco basins, Venezuela

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(Armbruster, 2003b).
Peckoltia snethlageae (Steindachner, 1911)
Ancistrus snethlageae Steindachner, 1911a: 328. Type locality: Rio Tapajoz bei Villa Braga und Goyana [Brazil].
Syntypes: NMW 48049 (1), NMW 48050 (1).
Distribution: Tapajós River basin, Brazil (Fisch-Muller, 2003).
Remarks: Recently placed by Isbrücker & Seidel (in Isbrücker, 2001a: 17) in the monotypic genus Ancistomus.
Peckoltia ucayalensis (Fowler, 1940)
Hemiancistrus ucayalensis Fowler, 1940a: 235, figs. 24–25. Type locality: Ucayali River, Contamana, Peru. Holotype: ANSP 68651.
Distribution: Ucayali River basin, Peru (Fisch-Muller, 2003).
Peckoltia vermiculata (Steindachner, 1908)
Ancistrus vittatus vermiculata Steindachner, 1908f: 166. Type locality: dem mittleren Laufe des Amazonestrommes,
den Gewässern um Pará, Brasiliens. Syntypes: NMW 48056 (1), NMW 48059 (1), NMW 48064 (1). Originallyas Ancistrus vittatus var. vermiculata.
Distribution: Middle and lower Amazon River basin, Brazil (Fisch-Muller, 2003).
Peckoltia vittata (Steindachner, 1881)
Chaetostomus vittatus Steindachner, 1881c: 115, pl. 2 (fig. 5). Type locality: Amazonen-Strom, Tajapouru, Xingu
bei Porto de Moz, Rio Madeira [Brazil]. Syntypes: MCZ 7999 (1), MCZ 8017 (1), NMW 47225 (1), NMW
47226 (1), NMW 47227 (1), NMW 47228 (2).
Distribution: Middle and lower Amazon River basin, Brazil (Fisch-Muller, 2003).
PLANILORICARIA Isbrücker, 1971
Planiloricaria Isbrücker, 1971b: 276. Type species: Pseudohemiodon (Planiloricaria) cryptodon Isbrücker, 1971.
Type by original designation. Gender: Feminine. Originally proposed as a subgenus of Pseudohemiodon.
Remarks: Diagnosed and distinguished in key from related genera in Isbrücker & Nijssen (1986a).
Planiloricaria cryptodon (Isbrücker, 1971)
Pseudohemiodon (Planiloricaria) cryptodon Isbrücker, 1971b: 278, fig. 3. Type locality: Peru: Río Ucayali near
Pucallpa. Holotype: ZFMK 1865. Holotype illustrated in Isbrücker (1979b: fig. 4).
Distribution: Ucayali, Purus, and Mamoré River basins, Upper Amazon drainage (Isbrücker & Nijssen, 1986a).
Remarks: Redescribed in Isbrücker & Nijssen (1986a).
POGONOPOMA Regan, 1904
Pogonopoma Regan, 1904b: 205. Type species: Plecostomus wertheimeri Steindachner, 1867. Type by subsequent
designation by Eigenmann (1910: 407). Gender: Neuter. Originally proposed as subgenus of Plecostomus.
Pogonopomoides Gosline, 1947: 109. Type species: Rhinelepis parahybae Steindachner, 1877. Type by original designation. Gender: Neuter.
Revision: Quevedo & Reis (2002).
Pogonopoma obscurum Quevedo & Reis, 2002
Pogonopoma obscurum Quevedo & Reis, 2002: 405, fig. 3 (middle). Type locality: Rio Canoas at road from Anita
Garibaldi to Abdon Batista (approximatively 27º39'S, 51º05'W) Santa Catarina, Brazil. Holotype MCP 25036.
Distribution: Upper Uruguay River basin, Brazil (Weber, 2003).
Pogonopoma parahybae (Steindachner, 1877)
Rhinelepis Parahybae Steindachner, 1877c: 218, pl. 2. Type locality: aus dem Parahyba [Brazil]. Syntypes: MCZ
7756 (4), NMW 44556 (1), NMW 44557 (1), NMW 44558 (1).
Distribution: Paraíba do Sul River basin, Brazil (Weber, 2003).
Remarks: Redescribed (in Pogonopomoides) in Armbruster (1998a: 629).
Pogonopoma wertheimeri (Steindachner, 1867)
Plecostomus Wertheimeri Steindachner, 1867c: 119. Type locality: dem Fluss Mucuri im gleichnamige Districte bei
Santa Clara, in Brasilien. Holotype: NMW 44288; illustrated in Steindachner (1867f: 701, pl. 1).
Distribution: Mucuri and São Mateus River basins, Brazil (Weber, 2003).
Remarks: Redescription in Armbruster (1998a: 629).

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PROLORICARIA Isbrücker, 2001
Proloricaria Isbrücker, in Isbrücker et al., 2001: 21. Type species: Loricaria prolixa Isbrücker & Nijssen, 1978.
Type by original designation. Gender: Feminine.
Proloricaria lentiginosa (Isbrücker, 1979)
Loricaria prolixa lentiginosa Isbrücker, 1979a: 97, fig. 9. Type locality: Brésil, Est. Sao Paulo, haut bassin du Rio
Parana, Represa de Volta Grande, Rio Grande. Holotype: MZUSP 13188.
Distribution: Upper Paraná River basin, Brazil (Isbrücker, 1979).
Proloricaria prolixa (Isbrücker & Nijssen, 1978)
Loricaria prolixa Isbrücker & Nijssen, 1978a: 188, fig. 5. Type locality: Brazil, Est. São Paulo, Rio Piracicaba
through River Tietê, Rio Paraná system. Holotype: NMW 45091.
Distribution: Upper Paraná River drainage, La Plata basin, Brazil (Isbrücker & Nijssen, 1978).
PSEUDACANTHICUS Bleeker, 1862
Pseudacanthicus Bleeker, 1862 (in Bleeker, 1862–63): 2. Type species: Hypostomus serratus Valenciennes, 1840.
Type by original designation. Gender: Masculine.
Stoneiella Fowler, 1914: 271. Type species: Stoneiella leopardus Fowler, 1914. Type by original designation. Gender: Feminine.
Monophyly: Schaefer (1986: 113).
Pseudacanthicus fordii (Günther, 1868)
Chaetostomus Fordii Günther, 1868a: 476. Type locality: Surinam [Probably Suriname (Günther, 1868b)]. Syntypes(4): BMNH 1866.8.14.148 and BMNH 1866.8.14.152 (4), BMNH 1866.8.14.150 (1), BMNH 1866.8.14.151(1). Species illustrated and described in more detail, with additional comments on type locality, in Günther(1868b: 231, pl. 21).
Distribution: Coastal drainages of Suriname (Fisch-Muller, 2003).
Pseudacanthicus histrix (Valenciennes, 1840)
Rinelepis histrix Valenciennes, in Cuvier & Valenciennes, 1840b: 486 (359 in Strasbourg deluxe edition). Type
locality: [No locality stated]. No types known; description based on unpublished manuscript and illustrations byVandelli. Illustration of type reproduced by Capello (1870: pl. 7), and França (1922: fig. 9).
Loricaria spinosae Ferreira, in França, 1922: 84, fig. 9. Type locality: in Fluminis Amazonici. No types known.
Distribution: Negro and lower Amazon River basins, Brazil (Fisch-Muller, 2003).
Pseudacanthicus leopardus (Fowler, 1914)
Stoneiella leopardus Fowler, 1914: 271, fig. 17. Type locality: Rupununi River, British Guiana ... in the highlands ofBritish Guiana, approximately secured in North Latitude 2º to 3º, and West Longitude 50º20'. Holotype: ANSP
39345.
Distribution: Rupununi River basin, Guyana (Fisch-Muller, 2003).
Pseudacanthicus serratus (Valenciennes, 1840)
Hypostomus serratus Valenciennes, in Cuvier & Valenciennes, 1840b: 503 (372 in Strasbourg deluxe edition). Type
locality: rapportée de Surinam. Holotype: RMNH 3125.
Distribution: Coastal drainages of the Guianas (Fisch-Muller, 2003).
Pseudacanthicus spinosus (Castelnau, 1855)
Hypostomus spinosus Castelnau, 1855: 45, pl. 22 (fig. 3). Type locality: la rivière des Amazones. Holotype: MNHN
a-9577.
Distribution: Amazon River basin, Brazil (Fisch-Muller, 2003).
PSEUDANCISTRUS Bleeker, 1862
Pseudancistrus Bleeker, 1862 (in Bleeker, 1862–63): 2. Type species: Hypostomus barbatus Valenciennes, 1840.
Type by original designation. Gender: Masculine.
Lithoxancistrus Isbrücker, Nijssen & Cala, 1988: 14. Type species: Lithoxancistrus orinoco Isbrücker, Nijssen &Cala, 1988. Type by original designation. Gender: Masculine.

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Monophyly: Synonymy and species composition follows Armbruster (2004a, b).
Pseudancistrus barbatus (Valenciennes, 1840)
Hypostomus barbatus Valenciennes, in Cuvier & Valenciennes, 1840b: 506 (374 in Strasbourg deluxe edition). Type
locality: La Mana [French Guiana]. Holotype: MNHN a-9564.
Hypostomus guttatus Valenciennes, in Cuvier & Valenciennes, 1840b: 508 (375 in Strasbourg deluxe edition). Type
locality: Surinam. Holotype: RMNH 3126, illustrated in Bleeker (1864: 10, pl. 2 fig. 2, pl. 3 fig. 3).
Distribution: Major river basins of Guianas: Oyapock, Mana, Maroni, Suriname, Corantijn, and Essequibo (FischMuller, 2003).
Pseudancistrus brevispinis (Heitmans, Nijssen & Isbrücker, 1983)
Lasiancistrus brevispinis Heitmans, Nijssen & Isbrücker, 1983: 38, fig. 4 (middle). Type locality: Surinam, Nickerie
River system, district Nickerie, Fallawatra River, rapid 5 km S. W. of Stondansie Fall. Holotype: ZMA 107.740.
Distribution: Atlantic coastal drainages of the Guianas, from Nickerie to Oyapock River basins, (Fisch-Muller,
2003).
Remarks: Redescribed in Le Bail et al. (2000: 236, fig.).
Pseudancistrus coquenani (Steindachner, 1915)
Ancistrus (Pseudancistrus) coquenani Steindachner, 1915e: 90, pl. 11 (fig. 3). Type locality: dem Coquenanflusse,
einem Nebenflusse des Caroni in Venezuela. Syntypes (3): NMW 48023 (2). Name proposed conditionally as“coquenani n. sp.? an A. güntheri Regan female.”Distribution: Kukenan River basin in upper Caroni River drainage, Venezuela (Fisch-Muller, 2003).
Pseudancistrus depressus (Günther, 1868)
Chaetostomus depressus Günther, 1868a: 476. Type locality: Surinam. Holotype: BMNH 1866.8.14.139. Describedin more detail in Günther (1868b: 232).
Distribution: Unknown locality in Suriname (Fisch-Muller, 2003).
Pseudancistrus genisetiger Fowler, 1941
Pseudancistrus genisetiger Fowler, 1941a: 155, figs. 59–62. Type locality: Rio Jaguaribe, Orós, Ceará [Brazil].
Holotype: ANSP 69441.
Distribution: Jaguaribe River basin, Brazil (Fisch-Muller, 2003).
Pseudancistrus guentheri (Regan, 1904)
Ancistrus guentheri Regan, 1904b: 241, pl. 12 (fig. 3). Type locality: British Guiana. Holotype: BMNH 1978.3.2.1.
Distribution: Unknown locality in Guyana (Fisch-Muller, 2003).
Pseudancistrus longispinis (Heitmans, Nijssen & Isbrücker, 1983)
Lasiancistrus longispinis Heitmans, Nijssen & Isbrücker, 1983: 45, fig. 8 (middle). Type locality: French Guiana,
Oyapock River system: Camopi River at Pauwé Jean-Jean, upstream of Saut Mauvais (03º11'N 52º22'W). Holotype: IRSNB 612.
Distribution: Oyapock River basin, French Guiana (Fisch-Muller, 2003).
Pseudancistrus niger (Norman, 1926)
Hemiancistrus niger Norman, 1926b: 96. Type locality: Oyapock River at Sant [sic] Cafesoca, French Guiana. Lectotype: BMNH 1926.3.2.756, designated by Heitmans et al. (1983: 35).
Distribution: Oyapock River basin, French Guiana (Fisch-Muller, 2003).
Pseudancistrus nigrescens Eigenmann, 1912
Pseudancistrus nigrescens Eigenmann, 1912b: 234, pl. 25 (fig. 2). Type locality: Amatuk, British Guiana. Holotype:
FMNH 53105.
Distribution: Upper Potaro River basin, Guyana (Fisch-Muller, 2003).
Pseudancistrus orinoco (Isbrücker, Nijssen & Cala, 1988)
Lithoxancistrus orinoco Isbrücker, Nijssen & Cala, 1988: 14, fig. 1 (top). Type locality: Colombie, Province deVichada, Rio Orinoco, à la cascade (=raudal) près de l’embouchure du Rio Tuparro. Holotype: ICNMHN 1200.
Distribution: Orinoco River basin (Fisch-Muller, 2003).
Pseudancistrus papariae Fowler, 1941

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Pseudancistrus papariae Fowler, 1941a: 157, figs. 63–65. Type locality: Lago Papary, Rio Grande do Norte [Brazil]. Holotype: ANSP 69442.
Distribution: Lake Papari and Jaguaribe River basin, Brazil (Fisch-Muller, 2003).
Pseudancistrus sidereus Armbruster, 2004
Pseudancistrus sidereus Armbruster, 2004b: 8, fig. 3. Type locality: Venezuela, Amazonas, Rio Orinoco dr.: 175.6,
Río Siapa from 10 to 15 km downstream, Río Casiquiare – Río Negro dr., 01.50000º, –065.71667º. Holotype:
MCNG 26125.
Distribution: Upper Orinoco River and Casiquaire, Amazonas, Venezuela (Armbruster, 2004b).
PSEUDOHEMIODON Bleeker, 1862
Pseudohemiodon Bleeker, 1862 (in Bleeker, 1862–63): 3. Type species: Hemiodon platycephalus Kner, 1853. Typeby original designation. Gender: Masculine.
Pseudohemiodon amazonus (Delsman, 1941)
Loricaria apeltogaster amazonum Delsman, 1941: 80. Type locality: Trombetas-river, near Obidos [Brazil]. Holotype: IRSNB 546. Originally as Loricaria apeltogaster var. amazonum.
Distribution: Lower Amazon River basin, Brazil.
Pseudohemiodon apithanos Isbrücker & Nijssen, 1978
Pseudohemiodon apithanos Isbrücker & Nijssen, 1978a: 195, fig. 6. Type locality: Ecuador, Prov. Napo, Río
Conejo, a tributary of Río San Miguel, vicinity of Santa Cecilia, 0º06'N, 76º51'W, upper Río Amazonas system.
Holotype: ANSP 134370.
Distribution: San Miquel River basin, Western Amazon River drainage, Ecuador.
Pseudohemiodon devincenzii (Soriano-Señorans, 1950)
Loricaria devincenzii Soriano-Señorans, 1950: 265. Type locality: sobre fondo de arena fangosa con red de playa, a
200 metros aqua abajo d la desembocadura del arroyo Espinillar, en el Río Uruguay, Rep. Oriental del Uruguay,
departamento de Salto. Holotype: MHNM (apparently lost, Olazarri et al., 1970).
Distribution: Uruguay River basin, Uruguay.
Pseudohemiodon laminus (Günther, 1868)
Loricaria lamina Günther, 1868a: 478. Type locality: Xeberos (Upper Amazon). Lectotype: BMNH 1867.6.13.33,
designated by Isbrücker (1973: 186). Illustrated and described in more detail in Günther (1868b: 239, figs. 6, 7).
Distribution: Upper Amazon River basin, Peru.
Pseudohemiodon laticeps (Regan, 1904)
Loricaria laticeps Regan, 1904b: 295, pl. 20 (fig. 1). Type locality: Paraguay. Lectotype: BMNH 1895.5.17.113,
designated by Isbrücker (1973: 187).
Distribution: Uruguay, Paraná and Paraguay River basins.
Pseudohemiodon platycephalus (Kner, 1853)
Hemiodon platycephalus Kner, 1853a: 12, 25, pl. 1 (fig. 6), pl. 6 (fig. 2a–b). Type locality: Rio Cuiaba [Brazil].
Holotype: lost (Isbrücker, 1980: 122). Originally as Hem. ? platycephalus.
Distribution: Paraguay River basin, Brazil.
Pseudohemiodon thorectes Isbrücker, 1975
Pseudohemiodon thorectes Isbrücker, 1975a: 87, fig. 1. Type locality: Bolivia, Est. Santa Cruz, Buena Vista,
17º28'S, 63º37'W, west of Río Palacios, tributary of Río Mamoré, which is a tributary of Río Madeira, Rio Amazonas system. Holotype: BMNH 1927.10.4.43.
Distribution: Mamoré River basin, Bolivia.
PSEUDOLITHOXUS Isbrücker & Werner, 2001
Pseudolithoxus Isbrücker & Werner, in Isbrücker et al., 2001: 21. Type species: Lasiancistrus tigris Armbruster &Provenzano, 2000. Type by original designation. Gender: Masculine.
Remarks: This genus corresponds to the Lasiancistrus anthrax species group of Armbruster & Provenzano (2000).
Pseudolithoxus anthrax (Armbruster & Provenzano, 2000)
Lasiancistrus anthrax Armbruster & Provenzano, 2000: 243, fig. 1B. Type locality: Venezuela, Estado Amazonas,

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Río Orinoco backwater behind sand beach ca. 0.5 hr. upstream from Isla Temblador, 03º04'N 66º28'W. Holotype: MBUCV V-18384.
Distribution: Upper and lower Orinoco tributaries, Caura and Aro rivers, Venezuela (Fisch-Muller, 2003).
Pseudolithoxus dumus (Armbruster & Provenzano, 2000)
Lasiancistrus dumus Armbruster & Provenzano, 2000: 246, fig. 3B. Type locality: Venezuela, Estado Amazonas,
Río Orinoco, backwater behind sand playa circa 0.5 hr. upstream from Isla Temblador, 03º04'N 66º28'W. Holotype: MBUCV V-17544.
Distribution: Upper Orinoco, upper Negro and Casiquiare River basins, Venezuela (Fisch-Muller, 2003).
Pseudolithoxus nicoi (Armbruster & Provenzano, 2000)
Lasiancistrus nicoi Armbruster & Provenzano, 2000: 249, fig. 4B. Type locality: Venezuela, Estado Amazonas,
Departamento Río Negro, Río Manipitare (Río Siapa drainage), from about 5 to 8 km upstream from confluencewith Río Siapa. Holotype: MCNG 37033.
Distribution: Upper Negro River basin, including Canal Casiquiare basin, Venezuela (Fisch-Muller, 2003).
Pseudolithoxus tigris (Armbruster & Provenzano, 2000)
Lasiancistrus tigris Armbruster & Provenzano, 2000: 251, fig. 5B. Type locality: Venezuela, Estado Amazonas,
backwater of Río Orinoco behind sand playa circa 0.5 hr. upstream from Isla Temblador, 03º04'N 66º28'W.
Holotype: MBUCV V-17546.
Distribution: Upper Orinoco and Ventuari River basins, Venezuela (Fisch-Muller, 2003).
PSEUDOLORICARIA Bleeker, 1862
Pseudoloricaria Bleeker, 1862 (in Bleeker, 1862–63): 3. Type species: Loricaria laeviuscula Valenciennes, 1840.
Type by original designation. Gender: Feminine.
Revision: Isbrücker & Nijssen (1976a).
Pseudoloricaria laeviuscula (Valenciennes, 1840)
Loricaria læviuscula Valenciennes, in Cuvier & Valenciennes, 1840b: 476 (p. 352 of Strasbourg deluxe edition).
Type locality: [Locality unknown]. Holotype: MNHN b-0365.
Distribution: Lower and middle Amazon basin, including Negro and Branco rivers, Brazil (Isbrücker & Nijssen,
1976a).
PSEUDORINELEPIS Bleeker, 1862
Pseudorinelepis Bleeker, 1862 (in Bleeker, 1862–63): 3. Type species: Rinelepis genibarbis Valenciennes, 1840.
Type by original designation. Gender: Feminine.
Canthopomus Eigenmann, 1910: 407. Type species: Rhinelepis genibarbis Valenciennes, 1840. Type by original
designation. Gender: Feminine. Designation by Eigenmann & Allen (1942: 183) of Rhinelepis agassizii Steindachner as type is invalid.
Monistiancistrus Fowler, 1940a: 236. Type species: Monistiancistrus carachama Fowler, 1940. Type by original
designation. Gender: Masculine.
Relationships: Armbruster (1998a: 631).
Pseudorinelepis genibarbis (Valenciennes, 1840)
Rinelepis genibarbis Valenciennes, in Cuvier & Valenciennes, 1840b: 484 (357 in Strasbourg deluxe edition), pl.
453. Type locality: probablement du Brésil. Holotype (15 pouces): at MNHN.
Rhinelepis Agassizii Steindachner, 1877c: 228. Type locality: am See Manacapouru [Brazil]. Syntypes: MCZ 8007
(1), NMW 44559 (3), NMW 445560 (1), NMW 44561 (1).
Plecostomus pellegrini Regan, 1904b: 218. Type locality: from Upper Amazon. Holotype: MNHN a-3956.
Monistiancistrus carachama Fowler, 1940a: 236, figs. 26–27. Type locality: Ucayali River basin, Contamana, Peru.
Holotype: ANSP 68654.
Distribution: Upper Amazon River basin (Weber, 2003).
Remarks: Redescribed in Armbruster & Hardman (1999).
PSEUDOTOCINCLUS Nichols, 1919

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Pseudotocinclus Nichols, 1919b: 534. Type species: Pseudotocinclus intermedius Nichols, 1919. Type by monotypy.
Gender: Masculine.
Revision, with key to species (Takako et al., 2005: 501).
Pseudotocinclus juquiae Takako, Oliveira & Oyakawa, 2005
Pseudotocinclus juquiae Takako, Oliveira & Oyakawa, 2005: 504, fig. 4. Type locality: Brazil, São Paulo, first tributary of the rio Juquiá, near ribeirão das Antas on the road from São Lourenço da Serra to Juquitiba,
23º59'49.1"S, 46º56'01.0"W, Juquitiba. Holotype: MZUSP 49333.
Distribution: Juquiá River basin, Brazil (Takako et al., 2005).
Pseudotocinclus parahybae Takako, Oliveira & Oyakawa, 2005
Pseudotocinclus parahybae Takako, Oliveira & Oyakawa, 2005: 503, fig. 3. Type locality: Brazil, São Paulo, a tributary of the ribeirão Grande, Fazenda São Sebastião do Ribeirão Grande, rio Paraíba do Sul basin, approximately 22º46'S, 45º27'W, Pindamonhangaba. Holotype: MZUSP 49122.
Distribution: Ribeirão Grande tributary, Paraíba do Sul River basin, Brazil (Takako et al., 2005).
Pseudotocinclus tietensis (Ihering, 1907)
Otocinclus (Microlepidogaster) tietensis Ihering, 1907: 26. Type locality: Rio Tieté, São Paulo, Brazil. Holotype: at
DZSASP [now MZUSP] (not found).
Pseudotocinclus intermedius Nichols, 1919b: 534. Type locality: Campo Grande, Mato Grosso, Brazil. Holotype:
AMNH 7177.
Distribution: Upper Tietê River at Paranapiacaba, Brazil (Schaefer, 2003c).
PSEUDOTOTHYRIS Britski & Garavello, 1984
Pseudotothyris Britski & Garavello, 1984: 232. Type species: Otocinclus obtusus Miranda Ribeiro, 1911. Type byoriginal designation. Gender: Feminine.
Pseudotothyris janeirensis Britski & Garavello, 1984
Pseudotothyris janeirensis Britski & Garavello, 1984: 234, figs. 9–10. Type locality: Rio dos Macacos, Represa
Engenho da Serra, Paulo de Frontin, Rio de Janeiro, Brazil. Holotype: MNRJ 10278.
Distribution: Rio de Janeiro State, Brazil (Schaefer, 2003c).
Pseudotothyris obtusa (Miranda Ribeiro, 1911)
Otocinclus obtusus Miranda Ribeiro, 1911: 95, pl. 29 (fig. 3). Type locality: Brazil. Lectotype: MNRJ 1026A, designated by Miranda Ribeiro (1953: 401). As Otocinclus obtusos at heading of account and index (p. 498), andOtocinclus obtusus in etymology, index (p. 497), figure caption, and elsewhere in the text. Britski & Garavello(1984: 226) treated O. obtusa as the valid name, with obutsos as a misspelling, thereby acting as first reviser.
Distribution: Southeastern Brazil (Schaefer, 2003c).
PTEROSTURISOMA Isbrücker & Nijssen, 1978
Pterosturisoma Isbrücker & Nijssen, 1978b: 69. Type species: Harttia microps Eigenmann & Allen, 1942. Type byoriginal designation. Gender: Neuter.
Pterosturisoma microps (Eigenmann & Allen, 1942)
Harttia microps Eigenmann & Allen, 1942: 211, pl. 8 (figs. 3–4); pl. 9 (fig. 1). Type locality: Iquitos [Peru]. Lectotype: CAS 28543, designated by Isbrücker & Nijssen (1978b: 71).
Distribution: Upper Amazon River basin, Peru (Eigenmann & Allen, 1942).
PTERYGOPLICHTHYS Gill, 1858
Pterygoplichthys Gill, 1858: 408. Type species: Hypostomus duodecimalis Valenciennes, 1840. Type by subsequent
designation in Bleeker (1862–63: 2) with genus spelled Pterygophlichtys. Gender: Masculine.
Liposarcus Günther, 1864: 238. Type species: Hypostomus multiradiatus Hancock, 1828. Type by subsequent designation by Jordan (1919b: 332). Gender: Masculine
Glyptoperichthys Weber, 1991: 639. Type species: Ancistrus lituratus Kner, 1854. Type by original designation.
Gender: Masculine.
Remarks: Revised by Weber (1992), with Liposarcus and Glyptoperichthys also treated as valid. Generic synonymy

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herein based on Armbruster (2004a).
Pterygoplichthys ambrosettii (Holmberg, 1893)
Liposarcus ambrosettii Holmberg, 1893b: 354. Type locality: Río Paraguay, in front of Formosa. No types known.
Pterygoplichthys anisitsi Eigenmann & Kennedy, 1903: 503. Type locality: Laguna of the Rio Paraguay at Asuncion. Holotype: IU 9873; whereabouts unknown.
Pterygoplichthys juvens Eigenmann & Kennedy, 1903: 504. Type locality: Asuncion, Río Paraguay, Paraguay. Holotype: CAS 59784.
Ancistrus multiradiatus alternans Regan, 1904b: 229. Type locality: Paraguay and Southern Bolivia [restricted byWeber (1992: 28) to: Paraguyan Chaco [...] région Vila Concepción (=Concepción) — Caraya Vuelta -2/3 Piste
Pozo Colorado, probablement Waikthlatingmayalwa (=? Mission Inglesia)]. Lectotype: BMNH 1898.7.4.5, designated by Weber (1992: 28). Originally proposed as A. multiradiatus var. alternans.
Distribution: Paraguay, middle Paraná, Bermejo, and Uruguay River basins (Weber, 2003).
Remarks: Weber (1992, 2003) treated Liposarcus ambrosettii Holmberg, 1893, as a nomen oblitum, but its use in
several publications negate that status. Redescribed as Liposarcus anisitsi in Weber (1992: 13).
Pterygoplichthys disjunctivus (Weber, 1991)
Liposarcus disjunctivus Weber, 1991: 638. Type locality: Rio Madeira, système de l’Amazonas, Restauracão, Amazonas, Brésil. Holotype: MZUSP 28360; illustrated in Weber (1992: pl. 9).
Distribution: Madeira River basin, Bolivia and Brazil (Weber, 2003).
Pterygoplichthys etentaculatus (Spix & Agassiz, 1829)
Hypostoma etentaculatum Spix & Agassiz, 1829: 7, pl. 4 (fig. 1). Type locality: in Brasilae septentrioralis fluviis[now: São Francisco River at Januaria, Minas Gerais State, Brazil, by neotype designation]. Neotype: MZUSP
35821, designated by Weber (1992: 25).
Hypostomus duodecimalis Valenciennes in Cuvier & Valenciennes, 1840b: 498 (367 in Strasbourg deluxe edition),pl. 454. Type locality: dans la rivière Saint-François au Brésil. Holotype: MNHN a-9446.
Hypostomus brevitentaculatus Ranzani, 1841: 63. Type locality: nelle aqua dolci della Provincia di S. Paulo nel
Brasile. Holotype: at MZUB. Described in more detail in Ranzani (1842: 329).
Ancistrus longimanus Kner, 1854: 283, pl. 5 (fig. 1). Type locality: [Not stated]. Syntypes (2): at NMW. Originallyproposed as Anc. longimanus.
Distribution: São Francisco River basin, Brazil (Weber, 2003).
Pterygoplichthys gibbiceps (Kner, 1854)
Ancistrus gibbiceps Kner, 1854: 284, pl. 5 (fig. 2). Type locality: aus dem Rio Negro bei Marabitanos [Brazil]. Holotype: at NMW. Originally proposed as Anc. gibbiceps.
Liposarcus altipinnis Günther, 1864: 239. Type locality: Rio Cupai [Brazil]. Holotype: BMNH 1853.3.19.34.
Distribution: Middle and upper Amazon and Orinoco basins (Weber, 2003).
Remarks: Redescribed in Weber (1992: 17), with type locality of Liposarcus altipinnis as: Cupari River, Tapajós
River basin, Amazon drainage, Amazonas State, Brazil.
Pterygoplichthys joselimaianus (Weber, 1991)
Glyptoperichthys joselimaianus Weber, 1991: 640, illustrated in Weber (1992: 20, pl. 15b). Type locality: Rio Araguaya, affl. du Tocantins, système de l’Amazone, Aruanã, Goiàs, Brésil. Holotype: MZUSP 4873.
Distribution: Tocantins River basin, Brazil (Weber, 2003).
Pterygoplichthys lituratus (Kner, 1854)
Ancistrus lituratus Kner, 1854: 285, pl. 5 (fig. 3). Type locality: aus dem Rio Guaporé bei cidade do Matogrosso[Amazon system, Brazil]. Lectotype: NMW 16416; designated by Weber (1992: 28). Originally as Anc. lituratus.
Distribution: Madeira River basin, Bolivia and Brazil (Weber, 2003).
Remarks: Redescribed in Weber (1992: 28).
Pterygoplichthys multiradiatus (Hancock, 1828)
Hypostomus multiradiatus Hancock, 1828: 246. Type locality: Demerara [in title] [...] Lakes [...] on the slime. Holotype: BMNH 1857.6.13.162.
Distribution: Orinoco River basin, possibly also in Guyana (Weber, 2003).

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Remarks: Redescription and type locality interpretation in Weber (1992: 9, 27).
Pterygoplichthys pardalis (Castelnau, 1855)
Hypostomus pardalis Castelnau, 1855: 42, pl. 20 (fig. 3). Type locality: l’Amazone. Holotype: MNHN a-9574.
Liposarcus varius Cope, 1872a: 284. Type locality: The Ambyiacu river, which empties into the Amazon near toPebas, in Eastern Equador, some distance east of the Napo [Peru]. Lectotype: ANSP 21931, designated by
Fowler (1915: 233).
Liposarcus jeanesianus Cope, 1874b: 135. Type locality: Nauta, Peru [= Marañon River, Amazon basin]. Lectotype:
ANSP 8241, designated by Weber (1992: 27).
Distribution: Lower, middle and upper Amazon River basin (Weber, 2003).
Remarks: Redescribed in Weber (1992: 10).
Pterygoplichthys parnaibae (Weber, 1991)
Glyptoperichthys parnaibae Weber, 1991: 641. Type locality: Lac de Parnaguá, Rio Paraim, bassin de Parnaíba, sys.
côtier brésilien; Maranhão, Brésil. Holotype: NMW 48034, illustrated in Weber (1992: 21, pl. 16 b).
Distribution: Parnaíba River basin, Brazil (Weber, 2003).
Pterygoplichthys punctatus (Kner, 1854)
Loricaria punctata Kner, 1854: 281. Type locality: S. Vincente [...] aus einer Lache [Brazil]. Holotype: NMW
76587.
Distribution: Madeira, Purus, Jurua and Marañon River basins (Weber, 2003).
Remarks: Redescribed Weber (1992: 22).
Pterygoplichthys scrophus (Cope, 1874)
Liposarcus scrophus Cope, 1874b: 136. Type locality: Nauta [Peru]. Syntypes: USNM 132587 (2).
Distribution: Marañon and Ucayali River basins, Peru (Weber, 2003).
Remarks: Redescribed in Page et al. (1996: 186).
Pterygoplichthys undecimalis (Steindachner, 1878)
Chaetostomus undecimalis Steindachner, 1878a: 90. Type locality: Cienaga Grande de Santa Marta, Magdalena
basin, 40 km east of Barranquilla, Colombia. Lectotype: NMW 47224, designated in Weber (1992: 26); illustrated in Steindachner (1879d: 43, pl. 8).
Distribution: Magdalena River basin, Colombia (Weber, 2003); also Cauca and Catatumbo River basins (Maldonado-Ocampo et al., 2005).
Pterygoplichthys xinguensis (Weber, 1991)
Glyptoperichthys xinguensis Weber, 1991: 640. Type locality: Rio Fresco, affl. du Rio Xingu, système de l’Amazone; Aldeio Gorotire, municipe de São Felix do Xingu, Pará, Brésil. Holotype: MZUSP 35961, illustrated inWeber (1992: pl. 14, fig. b).
Distribution: Xingu River basin, Brazil (Weber, 2003).
Pterygoplichthys zuliaensis Weber, 1991
Pterygoplichthys zuliaensis Weber, 1991: 638. Type locality: Río Santa Ana, bassin du Maracaibo, Hacienda Río
Grande, 9º36'20"N, 72º07'00"W, état de Zulia, Venezuela. Holotype: MBUCV V-14653; holotype illustrated inWeber (1992: pl. 4b).
Distribution: Lake Maracaibo basin, Venezuela (Weber, 2003).
PYXILORICARIA Isbrücker & Nijssen, 1984
Pyxiloricaria Isbrücker & Nijssen, 1984b: 163. Type species: Pyxiloricaria menezesi Isbrücker & Nijssen, 1984.
Type by original designation. Gender: Feminine.
Pyxiloricaria menezesi Isbrücker & Nijssen, 1984
Pyxiloricaria menezesi Isbrücker & Nijssen, 1984b: 164, fig. 1. Type locality: Brazil, Mato Grosso do Sul, Lagoas
marginais da rodovia Transpantaneira, município de Miranda (20º10'S, 56º19'W).
Distribution: Paraguay River basin, Brazil (Isbrücker & Nijssen, 1984).
REGANELLA Eigenmann, 1905
Hemiodon Kner, 1853a: 24. Type species: Hemiodon depressus Kner, 1853. Type by subsequent designation by

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FERRARISBleeker (1862–63: 4). Gender: Masculine. Preoccupied by Hemiodon Swainson, 1840, in Mollusca; replaced byReganella Eigenmann, 1905.
Reganella Eigenmann, 1905: 793. Type species: Hemiodon depressus Kner, 1853. Type by being a replacement
name. Gender: Feminine. Replacement for Hemiodon Kner, 1853.
Reganella depressa (Kner, 1853)
Hemiodon depressus Kner, 1853a: 12, 27, pl. 7 (fig. 1a–c). Type locality: Rio negro und Marabitanos [restricted to
Marabitanos by lectotype designation]. Lectotype: NMW 9438, designated by Isbrücker & Nijssen (1974b:
219). Originally as Hem. depressus.
Distribution: Negro, Branco, and Tapajós River basins, Brazil (Ferraris, 2003c).
RHADINOLORICARIA Isbrücker & Nijssen, 1974
Rhadinoloricaria Isbrücker & Nijssen, 1974a: 73. Type species: Loricaria macromystax Günther, 1869. Type byoriginal designation. Gender: Feminine.
Rhadinoloricaria macromystax (Günther, 1869)
Loricaria macromystax Günther, 1869: 426, figs. 5–6. Type locality: Peruvian Amazonas. Holotype: BMNH
1869.5.21.8.
Distribution: Upper Amazon River basin (Ferraris, 2003c).
RHINELEPIS Spix & Agassiz, 1829
Rhinelepis Spix & Agassiz, 1829: 2. Type species: Rhinelepis aspera Spix & Agassiz, 1829. Type by monotypy.
Gender: Feminine.
Phylogenetic position: Armbruster (1998b).
Rhinelepis aspera Spix & Agassiz, 1829
Rhinelepis aspera Spix & Agassiz, 1829: 4, pl. 2 (figs. 1–2). Type locality: in flumine S. Francisci [Brazil]. Holotype: Whereabouts unknown (Kottelat, 1988).
Distribution: São Francisco and upper Paraná River basins, Brazil (Weber, 2003).
Rhinelepis strigosa Valenciennes, 1840
Rhinelepis strigosa Valenciennes, in Cuvier & Valenciennes, 1840: 480 [354 in Strasbourg deluxe edition]. Type
locality: dans le Parana et d’autres rXvières de la province de Corrientes [Argentina]. Holotype: MNHN a-9571.
Distribution: Paraná and Uruguay River basins (Weber, 2003).
RICOLA Isbrücker & Nijssen, 1978
Ricola Isbrücker & Nijssen, 1978a: 182. Type species: Loricaria macrops Regan, 1904. Type by original designation. Gender: Feminine.
Ricola macrops (Regan, 1904)
Loricaria macrops Regan, 1904b: 290, pl. 17 (fig. 3). Type locality: the R. de la Plata. Lectotype: BMNH
1868.9.16.1, designated by Isbrücker & Nijssen (1978a: 184).
Distribution: Lower Paraná River basin (Isbrücker & Nijssen, 1978).
Remarks: Reviewed in Ringuelet et al. (1967).
RINELORICARIA Bleeker, 1862
Rineloricaria Bleeker, 1862 (in Bleeker, 1862–63): 3. Type species: Loricaria lima Kner, 1853. Type by original
designation. Gender: Feminine.
Leliella Isbrücker, in Isbrücker et al., 2001: 20. Type species: Rineloricaria heteroptera Isbrucker & Nijssen, 1975.
Type by original designation. Gender: Feminine.
Rineloricaria aequalicuspis Reis & Cardoso, 2001
Rineloricaria aequalicuspis Reis & Cardoso, 2001: 324, fig. 3. Type locality: Brazil: Rio Grande do Sul: rio TrêsForquilhas, opposie to church of Vila Itati (29º30'43"S 50º05'31"W). Holotype: MCP 25852.
Distribution: Três Forquilhas and Maquiné River basins, Brazil (Reis & Cardoso, 2001).
Rineloricaria cadeae (Hensel, 1868)

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Loricaria cadeae Hensel, 1868: 369. Type locality: Rio Cadéa [Rio Grande do Sul, Brazil]. Lectotype: ZMB 7430,
designated by Isbrücker (1979a: 112).
Distribution: Laguna dos Patos basin, Brazil (Ferraris, 2003c).
Rineloricaria catamarcensis (Berg, 1895)
Loricaria catamarcensis Berg, 1895: 137. Type locality: Provincia Catamarca, Argentina. Holotype: at MACN (notfound, Isbrücker, 1979a: 113).
Distribution: Tala and Salí River basins in Catamarca Province, Argentina (Ringuelet et al., 1967).
Remarks: Reviewed in Ringuelet et al. (1967).
Rineloricaria cubataonis (Steindachner, 1907)
Loricaria cubataonis Steindachner, 1907e: 489. Type locality: aus dem Flusse Cubatão, im Staate Santa Catharina
bei Theoresopolis (Brasilien). Holotype: NMW (not found, Isbrücker, 1979a: 113).
Distribution: Coastal Atlantic rivers of southeastern Brazil (Ferraris, 2003c).
Rineloricaria felipponei (Fowler, 1943)
Loricaria felipponei Fowler, 1943b: 319, figs. 7–9. Type locality: Uruguay. Holotype: ANSP 70324.
Distribution: Lower La Plata River basin, Uruguay (Ferraris, 2003c).
Rineloricaria henselii (Steindachner, 1907)
Loricaria henselii Steindachner, 1907e: 488. Type locality: Aus dem Flusse Cubatão im Staate Santa Catherina bei
Theresopolis (Brasilien). Holotype: at NMW (not found, Isbrücker, 1979a: 113).
Distribution: Cubatão River in southern Santa Catarina State, Brazil (Ferraris, 2003c).
Rineloricaria heteroptera Isbrücker & Nijssen, 1976
Rineloricaria heteroptera Isbrücker & Nijssen, 1976b: 112, fig. 1. Type locality: Brazil, Est. Amazonas. Creek inreserve Ducke, near Manaus (03º06'S, 60º00'W). Holotype: IRSNB 555.
Distribution: Amazon River basin, near Manaus, Brazil (Ferraris, 2003c).
Remarks: Recently designated as type of a monotypic genus, Leliella, by Isbrücker (2001: 20).
Rineloricaria jaraguensis (Steindachner, 1909)
Loricaria jaraguensis Steindachner, 1909a: 196. Type locality: dem Jaraguá und dessen Nebenflussen in brasilianischen Staate Sa. Catharina. Lectotype: NMW 44886, designated by Isbrücker (1979a: 113).
Distribution: Jaraguá River basin in Santa Catarina State, Brazil (Ferraris, 2003c).
Rineloricaria kronei (Miranda Ribeiro, 1911)
Loricaria kronei Miranda Ribeiro, 1911: 132, pl. 34 (fig. 2). Type locality: Rio Iporanga, S. Paulo [Brazil]. Holotype: MNRJ (not found).
Distribution: Ribeira de Iguape River basin, Brazil (Ferraris, 2003c).
Rineloricaria latirostris (Boulenger, 1900)
Loricaria latirostris Boulenger, 1900b: 165. Type locality: Southern Brazil, Province São Paulo, Mogy-guassu River
about 250 miles inland of Santos. Lectotype: BMNH 1899.12.18.6, designated by Isbrücker (1979a: 112).
Loricaria paulina Boulenger, 1900b: 166. Type locality: Southern Brazil, Province São Paulo, Mogy-guassu River.
Lectotype: BMNH 1899.12.18.8, designated by Isbrücker (1979a: 112).
Distribution: Upper Paraná River basin, Brazil (Ferraris, 2003c).
Remarks: Loricaria latirostris treated as valid following first reviser action of Regan (1904b: 283).
Rineloricaria lima (Kner, 1853)
Loricaria lima Kner, 1853a: 12, 25, pl. 6 (fig. 1a–b). Type locality: aus Brasilien durch Natterer, ober ohne nähere
Angabe des Fundortes. Holotype: NMW (not found, Isbrücker, 1980: 107). Originally as Lor.? lima.
Distribution: Brazil (Ferraris, 2003c).
Rineloricaria longicauda Reis, 1983
Rineloricaria longicauda Reis, 1983: 64, figs. 5–7. Type locality: Banhado do Taim, município de Rio Grande, RS[Brazil]. Holotype: MZUSP 16078.
Distribution: Coastal drainages of Rio Grande do Sul State, between Chuí and Tramandaí, Brazil (Reis, 1983).
Rineloricaria maquinensis Reis & Cardoso, 2001
Rineloricaria maquinensis Reis & Cardoso, 2001: 321, fig. 1. Type locality: Brazil: Rio Grande do Sul: arroio ÁguaParada, tributary to rio Maquiné (approximately 29º40'S 50º12'W), Maquiné. Holotype: MCP 25817.

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FERRARISDistribution: Maquiné and Araranguá River basins in southern Brazil (Reis & Cardoso, 2001).
Rineloricaria microlepidogaster (Regan, 1904)
Loricaria microlepidogaster Regan, 1904b: 283, pl. 15 (fig. 4). Type locality: Rio Grande do Sul [Brazil]. Holotype:
BMNH 1884.2.5.41.
Distribution: Laguna dos Patos basin, Brazil (Ferraris, 2003c).
Rineloricaria microlepidota (Steindachner, 1907)
Loricaria lima microlepidota Steindachner, 1907b: 154. Type locality: dem Juruá [Brazil]. Syntypes: at NMW (notfound: Isbrücker, 1979a: 113). Originally as Loricaria lima var. microlepidota.
Distribution: Juruá River basin, Brazil (Ferraris, 2003c).
Rineloricaria misionera Rodríguez & Miquelarena, 2005
Rineloricaria misionera Rodríguez & Miquelarena, 2005: 5, fig. 1. Type locality: arroyo Cuña-Pirú (27º08'S,
54º54'W) Canguás Department, Misiones Province, Argentina. Holotype: ILPLA 1698.
Distribution: Paraná and Uruguay River basins, Misiones, Argentina (Rodríguez & Miquelarena, 2005).
Rineloricaria pareiacantha (Fowler, 1943)
Loricaria pareiacantha Fowler, 1943b: 321, figs. 10–12. Type locality: Rio Santa Lucia, Canelones [Uruguay].
Holotype: ANSP 67815.
Distribution: Santa Lucia River basin, Uruguay (Ferraris, 2003c).
Rineloricaria pentamaculata Langeani & Araujo, 1994
Rineloricaria pentamaculata Langeani & Araujo, 1994: 153, fig. 1. Type locality: rio Turvo, Ourinhos, São Paulo,
Brasil. Holotype: MZUSP 43051.
Distribution: Turvo River basin in São Paulo State, Brazil (Langeani & Araujo, 1994).
Rineloricaria quadrensis Reis, 1983
Rineloricaria quadrensis Reis, 1983: 66, figs. 15–17. Type locality: Lagoa dos Quadros, município de Osório, RS[Brazil]. Holotype: MZUSP 14363.
Distribution: Coastal streams and lagoons in Rio Grande do Sul, between Tramandaí and Torres, Brazil (Reis, 1983).
Rineloricaria steindachneri (Regan, 1904)
Loricaria steindachneri Regan, 1904b: 281. Type locality: Eastern Brazil, from the R. Preto to the R. Parahyba. Lectotype: NMW 45016: 1, designated by Isbrücker (1979a: 113).
Distribution: Coastal rivers of northeastern Brazil (Ferraris, 2003c).
Rineloricaria strigilata (Hensel, 1868)
Loricaria strigilata Hensel, 1868: 368. Type locality: Südbrasilien, in eimen steinigen Bache bei Santa Cruz. Holotype: ZMB 7429.
Distribution: Laguna dos Patos basin, Brazil (Ferraris, 2003c).
Rineloricaria thrissoceps (Fowler, 1943)
Loricaria thrissoceps Fowler, 1943b: 322, figs. 13–15. Type locality: Río Santa Lucia, Canelones [Uruguay]. Holotype: ANSP 67796.
Distribution: Santa Lucia River basin, Uruguay (Ferraris, 2003c).
SCHIZOLECIS Britski & Garavello, 1984
Schizolecis Britski & Garavello, 1984: 228. Type species: Microlepidogaster guntheri Miranda Ribeiro, 1918. Typeby original designation. Gender: Feminine.
Schizolecis guntheri (Miranda Ribeiro, 1918)
Microlepidogaster guntheri Miranda Ribeiro, 1918d: 634. Type locality: Praia do Piraíque, Ilha de São Sebastião,Estado de São Paulo [Brazil]. Lectotype: MZUSP 1015, designated by Britski (1969: 208).
Pseudotocinclus ribeiroi Gomes, 1955: 222, pl. 1. Type locality: Rio Branco, Bacia do rio Itanhaem, São Paulo, Brazil. Holotype: EEBP 110.
Distribution: Southeastern Brazil (Schaefer, 2003c).
Remarks:There is some controversy regarding the correct spelling of the valid name for this species. The name waslisted as Microlepidogaster guntheri Miranda Ribeiro (1918d: 634) with a statement in a footnote on that pagethat the species name was a patronym for a “Sr. F. Gunther.” However, Isbrücker (1980: 85–86) reported that the

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species was listed as Microlepidogaster güntheri in a later article (Miranda Ribeiro, 1918e: 717) in the same
volume of the journal. If the two papers were published simultaneously, the two spellings would need to betreated as alternate spellings that require further action. Isbrücker (1980: 85–86) noted both spellings andtreated M. guentheri as valid, and would thereby act as first reviser, if necessary.
SCOBINANCISTRUS Isbrücker & Nijssen, 1989
Scobinancistrus Isbrücker & Nijssen, 1989: 542. Type species: Scobinancistrus pariolispos Isbrücker & Nijssen,
1989. Type by original designation. Gender: Masculine.
Scobinancistrus aureatus Burgess, 1994
Scobinancistrus aureatus Burgess, 1994: 237, figure on p. 236. Type locality: Ilha da Fazenda, Rio Xingu, EstadoPara, Brazil. Holotype: MZUSP 47690.
Distribution: Xingu River basin, Brazil (Fisch-Muller, 2003).
Scobinancistrus pariolispos Isbrücker & Nijssen, 1989
Scobinancistrus pariolispos Isbrücker & Nijssen, 1989: 542, fig. 1. Type locality: Brasilien, Est. Pará, Rio
Tocantins, Jatobal. Holotype: INPA 1076.
Distribution: Tocantins and Tapajós River basins, Brazil (Fisch-Muller, 2003).
SPATULORICARIA Schultz, 1944
Spatuloricaria Schultz, 1944c: 287, 334. Type species: Spatuloricaria phelpsi Schultz, 1944. Type by original designation. Gender: Feminine.
Euacanthagenys Fowler, 1945a: 123. Type species: Euacanthagenys caquetae Fowler, 1945. Type by original designation. Gender: Feminine.
Spatuloricaria atratoensis Schultz, 1944
Spatuloricaria atratoensis Schultz, 1944a: 155. Type locality: Río Truando, tributary of Río Atrato, Colombia.
Holotype: USNM 93810.
Distribution: Atrato River basin, Colombia (Ferraris, 2003c).
Spatuloricaria caquetae (Fowler, 1943)
Loricaria caquetae Fowler, 1943a: 261, figs. 60–62. Type locality: Florencia, Rio Orteguasa, Colombia. Holotype:
ANSP 70526.
Distribution: Orteguasa River basin, Colombia (Ferraris, 2003c).
Spatuloricaria curvispina (Dahl, 1942)
Loricaria curvispina Dahl, 1942: 85, figs. 5–7. Type locality: Río Batatal, tributary to Río San Jorge, which is next
to Rio Cauca the largest tributary to Río Magdalena. Department of Bolivar, Republic of Colombia. Holotype:possibly at ZMUL.
Distribution: Magdalena River basin, Colombia (Ferraris, 2003c).
Spatuloricaria euacanthagenys Isbrücker, 1979
Euacanthagenys caquetae Fowler, 1945a: 125, figs. 37–40. Type locality: Morelia, Rio Caquetá drainage, Colombia. Holotype: ANSP 71718; holotype illustrated in Isbrücker (1979a: 111, figs. 24, 25, 26, 27b). Preoccupied in
Spatuloricaria by Loricaria caquetae Fowler, 1943; replaced by Spatuloricaria euacanthagenys Isbrücker,
1979.
Spatuloricaria euacanthagenys Isbrücker, 1979a: 113, figs. 24–26. Type locality: Morelia, Río Caquetá drainage,
Colombia. Holotype: ANSP 71718; holotype illustrated in Isbrücker (1979a: 113, figs 24, 25, 26, 27b). Replacement for Euacanthagenys caquetae Fowler, 1945.
Distribution: Caquetá River basin, Colombia (Ferraris, 2003c).
Spatuloricaria evansii (Boulenger, 1892)
Loricaria Evansii Boulenger, 1892b: 10, pl. 1. Type locality: Jangada [Matto Grosso, Brazil]. Holotype: BMNH
1892.4.20.29; holotype illustrated in Isbrücker (1973: 174, fig. 1).
? Loricaria cirrhosa Perugia, 1897: 22. Type locality: Rio Beni, Missioni Mosetenes, Bolivia. Lectotype: MSNG
8850 (1); lectotype designated by, and illustrated in, Isbrücker (1973: 174, fig. 2). Preoccupied by Loricaria cirrhosa Bloch & Schneider, 1801; apparently not replaced.

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FERRARISDistribution: Paraguay and Beni River basins (Ferraris, 2003c).
Spatuloricaria fimbriata (Eigenmann & Vance, 1912)
Loricaria fimbriata Eigenmann & Vance, in Eigenmann, 1912a: 12. Type locality: Boca de Certegai [Colombia].
Holotype: FMNH 55117; holotype illustrated in Eigenmann (1922b: 93, pl. 15, figs.1, 2).
Distribution: Tuira River, Panama and Magdalena River and Boca de Certegai, Colombia (Ferraris, 2003c).
Spatuloricaria gymnogaster (Eigenmann & Vance, 1912)
Loricaria gymnogaster Eigenmann & Vance, in Eigenmann, 1912a: 12. Type locality: Apulo [Colombia]. Holotype:
CAS 28773; holotype illustrated in Eigenmann (1922b: 92, pl. 14, figs. 3–4).
Distribution: Upper Magdalena River basin, Colombia (Maldonado-Ocampo et al., 2005).
Spatuloricaria lagoichthys (Schultz, 1944)
Loricaria gymnogaster lagoichthys Schultz, 1944c: 331, pl. 13. Type locality: Río Palmar near Totuma, about 100
km southwest of Maracaibo [Venezuela]. Holotype: USNM 121092. Teeth of holotype illustrated in Isbrücker
(1979a: 111, fig. 27a).
Distribution: Lake Maracaibo basin, Venezuela (Ferraris, 2003c).
Spatuloricaria nudiventris (Valenciennes, 1840)
Loricaria nudiventris Valenciennes, in Cuvier & Valenciennes, 1840b: 469 (346 of Strasbourg deluxe edition). Type
locality: la rivière de Saint-François au Brésil. Holotype: MNHN a-9558.
Distribution: São Francisco River basin, Brazil (Ferraris, 2003c).
Spatuloricaria phelpsi Schultz, 1944
Spatuloricaria phelpsi Schultz, 1944c: 335, pl. 14. Type locality: Río Socuy, 3 km above its mouth, Maracaibo
basin, Venezuela. Holotype: USNM 121121. Teeth of holotype illustrated in Isbrücker (1979a: 112, fig. 27c).
Distribution: Lake Maracaibo basin, Venezuela (Ferraris, 2003c).
Spatuloricaria puganensis (Pearson, 1937)
Loricaria puganensis Pearson, 1937: 96. Type locality: Pusoc, Rio Marañon, Peru. Lectotype: CAS 28776, designated by Isbrücker (1979a: 113).
Distribution: Marañon River basin, Peru (Ferraris, 2003c).
SPECTRACANTHICUS Nijssen & Isbrücker, 1987
Spectracanthicus Nijssen & Isbrücker, 1987: 93. Type species: Spectracanthicus murinus Nijssen & Isbrücker,
1987. Type by original designation. Gender: Masculine.
Spectracanthicus murinus Nijssen & Isbrücker, 1987
Spectracanthicus murinus Nijssen & Isbrücker, 1987: 94, fig. 1 (middle). Type locality: Brésil, Est. Pará, système du
Rio Tapajós: Poça de Pedra no Rio Tapajós, São Luis. Holotype: MZUSP 22011.
Distribution: Tapajós River basin, Brazil (Fisch-Muller, 2003).
SQUALIFORMA Isbrücker & Michels, 2001
Squaliforma Isbrücker & Michels, in Isbrücker et al., 2001: 22. Type species: Hypostomus horridus Kner, 1854.
Type by original designation. Gender: Feminine.
Squaliforma annae (Steindachner, 1881)
Plecostomus annae Steindachner, 1881c: 112, pl. 3 (figs. 2–2a). Type locality: Pará [= Belem, Brazil]. Holotype:
NMW 44073. Listed as Chaetostomus annae in figure caption (p. 146).
Distribution: Guamá River basin, Brazil (Weber, 2003).
Squaliforma biseriata (Cope, 1872)
Plecostomus biseriatus Cope, 1872a: 285. Type locality: the Amazon, between the mouth of the Rio Negro and the
Peruvian Amazon or Ucayale River. Holotype: ANSP 8279.
Distribution: Upper Amazon River basin, Brazil (Weber, 2003).
Squaliforma emarginata (Valenciennes, 1840)
Hypostomus emarginatus Valenciennes, in Cuvier & Valenciennes, 1840b: 500 (369 in Strasbourg deluxe edition).
Type locality: Probablement orXginaire du Brésil. Holotype: MNHN a-9447 (dry).
Distribution: Lower, middle and upper Amazon River basin (Weber, 2003).

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Squaliforma gomesi (Fowler, 1941)
Plecostomus iheringi Fowler, 1941a: 150, figs. 50–52. Type locality: Ceará [Brazil]. Holotype: ANSP 69409. Preoccupied by Plecostomus iheringii Regan, 1908; replaced by Plecostomus gomesi Fowler, 1942.
Plecostomus gomesi Fowler, 1942: [1]. Type locality: Ceará [Brazil]. Holotype: ANSP 69409. Replacement namefor Plecostomus iheringi Fowler, 1941; preoccupied by Plecostomus iheringii Regan, 1908.
Distribution: Jaguaribe River basin, Brazil (Weber, 2003).
Squaliforma horrida (Kner, 1854)
Hypostomus horridus Kner, 1854: 259, pl. 1 (fig. 1). Type locality: Forte do Principe am Rio Guaporé, Brazil. Syntypes (3): NMW 16325 (1), NMW 86604 (1). Originally proposed as Hyp. horridus.
Distribution: Madeira River basin, Brazil (Weber, 2003).
Squaliforma phrixosoma (Fowler, 1940)
Plecostomus phrixosoma Fowler, 1940a: 233, figs. 21–23. Type locality: Ucayali River basin, Contamana, Peru.
Holotype: ANSP 68650.
Distribution: Upper Amazon River basin, Peru (Weber, 2003).
Squaliforma scopularia (Cope, 1871)
Plecostomus scopularius Cope, 1871a: 55. Type locality: the Amazon above the mouth of the Rio Negro. Lectotype:
ANSP 8081, designated by Fowler (1915: 233); illustrated in Cope (1872a: pl. 16, no. 1).
Distribution: Middle Amazon River basin, Brazil (Weber, 2003).
Squaliforma squalina (Jardine, 1841)
Hypostoma squalinum Jardine, in Schomburgk, 1841: 142, pl. 2. Type locality: Rios Branco, Negro, and Essequibo.
No types known.
Distribution: Negro and Branco River basins; Essequibo River basin (Weber, 2003).
Squaliforma tenuicauda (Steindachner, 1878)
Plecostomus tenuicauda Steindachner, 1878a: 90. Type locality: Magdalenen-Stromes [Colombia]. Syntypes:
MSNG 8856 (1), NMW 42596 (1), NMW 44263 (1), NMW 44264 (1), NMW 44265 (1), NMW 44266 (3),
NMW 44268 (1), NMW 44294 (1), ZMUC P 30172 (1). One syntype illustrated in Steindachner (1879d: pl. 6).
Distribution: Magdalena River basin, Colombia (Weber, 2003).
Squaliforma tenuis (Boeseman, 1968)
Hypostomus tenuis Boeseman, 1968: 64, pl. 14 (fig. 3). Type locality: Near Paramaribo, Surinam. Holotype: RMNH
16198.
Distribution: Possibly Suriname River basin; known only from holotype (Weber, 2003).
Remarks: Type locality considered to be in error by Weber (2003).
Squaliforma villarsi (Lütken, 1874)
Plecostomus villarsi Lütken, 1874a: 211. Type locality: Caracas. Syntypes: ZMUC P 30149 (1), ZMUC P 30150 (1).
Distribution: Venezuela (Weber, 2003).
Squaliforma virescens (Cope, 1874)
Plecostomus virescens Cope, 1874b: 137. Type locality: not explicitly given [Upper Amazon]. Syntypes: ANSP
21280 (4).
Distribution: Upper Amazon River basin, Peru (?)(Weber, 2003).
STURISOMA Swainson, 1838
Sturisoma Swainson, 1838: 333. Type species: Loricaria rostrata Spix & Agassiz, 1829. Type by monotypy. Gender: Neuter. See also Swainson (1839: 304).
Oxyloricaria Bleeker, 1862 (in Bleeker, 1862–63): 3. Type species: Loricaria barbata Kner, 1853. Type by original
designation. Gender: Feminine.
Parasturisoma Miranda Ribeiro, 1911: 109. Type species: Loricaria (Rineloricaria) brevirostris Eigenmann &Eigenmann, 1889. Type by monotypy. Gender: Neuter.
Sturisoma aureum (Steindachner, 1900)
Loricaria aurea Steindachner, 1900: 206. Type locality: Bodega central, Rio Magdalena [Colombia]. Holotype: at
NMW.

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Sturisoma barbatum (Kner, 1853)
Loricaria barbata Kner, 1853a: 12, 23, pl. 5. Type locality: Rio Cuiabá, Cujaba-Fluss [Brazil]. Holotype: NMW
46155. Originally Lor. barbata.
Distribution: Paraguay River basin, Brazil (Ferraris, 2003c).
Sturisoma brevirostre (Eigenmann & Eigenmann, 1889)
Loricaria brevirostris Eigenmann & Eigenmann, 1889b: 35. Type locality: Iça [Brazil]. Holotype: MCZ 8095.
Distribution: Içá River basin, Brazil (Ferraris, 2003c).
Sturisoma dariense (Meek & Hildebrand, 1913)
Oxyloricaria dariensis Meek & Hildebrand, 1913: 81. Type locality: Rio Tuyra, Boca de Cupe, Panama. Holotype:
FMNH 7584.
Distribution: Tuira River basin, Panama (Ferraris, 2003c).
Sturisoma festivum Myers, 1942
Sturisoma festivum Myers, 1942: 100, fig. 8. Type locality: Río Monay, 35 km north of Trujillo, Motatan system,
Maracaibo basin, Venezuela. Holotype: SU 36505.
Distribution: Lake Maracaibo basin (Ferraris, 2003c).
Sturisoma frenatum (Boulenger, 1902)
Loricaria frenata Boulenger, 1902a: 69. Type locality: Northwestern Ecuador, St. Javier (60 feet), Saildero (350feet), and Rio Durango (350 feet) [restricted to Salidero, Ecuador by lectotype designation]. Lectotype: BMNH
1901.8.3.29, designated by Isbrücker (1979a: 113).
Distribution: Pacific versant rivers, Ecuador (Ferraris, 2003c).
Sturisoma guentheri (Regan, 1904)
Oxyloricaria guentheri Regan, 1904b: 299, pl. 18 (fig. 1). Type locality: Xeberos, Upper Amazon [Peru]. Holotype:
BMNH 1867.6.13.36.
Distribution: Upper Amazon River basin, Peru (Ferraris, 2003c).
Sturisoma kneri Ghazzi, 2005
Sturisoma kneri Ghazzi, 2005: 559, fig. 1. Type locality: Venezuela, Lago Maracaibo. Holotype: MZUT 764.
Distribution: Maracaibo Lake basin, Venezuela (Ghazzi, 2005).
Remarks: Previous citations of this name (as well as Loricaria kneri) were shown to be unavailable for purposes ofzoological nomenclature (Ghazzi, 2005).
Sturisoma lyra (Regan, 1904)
Oxyloricaria lyra Regan, 1904b: 300, pl. 19 (fig. 2). Type locality: Rio Jurua. Lectotype: BMNH 1897.12.1.64, designated by Isbrücker (1979a: 113).
Distribution: Juruá River basin, middle Amazon River drainage (Ferraris, 2003c).
Sturisoma monopelte Fowler, 1914
Sturisoma monopelte Fowler, 1914: 274, fig. 18. Type locality: Rupununi River, British Guiana ... in the highlandsof British Guiana, approximately secured in North Latitude 2º to 3º, and West Longitude 50º20'. Holotype:
ANSP 39346.
Distribution: Rupununi River basin, Guyana (Ferraris, 2003c).
Sturisoma nigrirostrum Fowler, 1940
Sturiosoma nigrirostrum Fowler, 1940a: 249, figs. 48–50. Type locality: Ucayali River basin, Contamana, Peru.
Holotype: ANSP 68666.
Distribution: Ucayali River basin, Peru (Ferraris, 2003c).
Sturisoma panamense (Eigenmann & Eigenmann, 1889)
Loricaria panamensis Eigenmann & Eigenmann, 1889b: 34. Type locality: Panama. Holotype: MCZ 8126.
Distribution: Pacific slope rivers of Panama, Colombia, and Ecuador (Ferraris, 2003c); Cauca River basin, Colombia (Maldonado-Ocampo et al., 2005).
Sturisoma robustum (Regan, 1904)
Oxyloricaria robusta Regan, 1904b: 300, pl. 19 (fig. 1). Type locality: Rio Paraguay. Lectotype: BMNH
1895.5.17.89, designated by Isbrücker (1979a: 113).

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Distribution: La Plata River basin (Ringuelet et al., 1967).
Remarks: Reviewed in Ringuelet et al. (1967).
Sturisoma rostratum (Spix & Agassiz, 1829)
Loricaria rostrata Spix & Agassiz, 1829: 5, pl. 3. Type locality: Brasiliae fluviis. Holotype: lost (Isbrücker, 1979a:
113).
Distribution: Brazil (Ferraris, 2003c).
Sturisoma tenuirostre (Steindachner, 1910)
Oxyloricaria tenuirostris Steindachner, 1910b: 410. Type locality: dem Rio Meta in Venezuela. Holotype: NMW
45453.
Distribution: Meta River basin, Venezuela (Ferraris, 2003c).
STURISOMATICHTHYS Isbrücker & Nijssen, 1979
Sturisomatichthys Isbrücker & Nijssen, in Isbrücker, 1979a: 91. Type species: Oxyloricaria leightoni Regan, 1912.
Type by original designation. Gender: Masculine.
Sturisomatichthys caquetae (Fowler, 1945)
Harttia caquetae Fowler, 1945a: 126, figs. 41–43. Type locality: Morelia, Rio Caquetá drainage, Colombia. Holotype: ANSP 71719.
Distribution: Caquetá River basin, Colombia (Ferraris, 2003c).
Sturisomatichthys citurensis (Meek & Hildebrand, 1913)
Oxyloricaria citurensis Meek & Hildebrand, 1913: 82. Type locality: Río Cupe at Cituro, Panama, Tuyra Riverbasin. Holotype: FMNH 7585.
Distribution: Tuira and Bayano River basins, Panama (Ferraris, 2003c).
Sturisomatichthys leightoni (Regan, 1912)
Oxyloricaria leightoni Regan, 1912b: 669, pl. 77 (fig. 2). Type locality: Honda, Colombia, 300–400 ft. Lectotype:
BMNH 1909.7.23.45, designated by Isbrücker (1979a: 113).
Distribution: Upper Magdalena and Cauca River basins, Colombia (Maldonado-Ocampo et al., 2005).
Sturisomatichthys tamanae (Regan, 1912)
Oxyloricaria tamanae Regan, 1912b: 669, pl. 77 (figs.1, 1a & b). Type locality: the Rio Tamana, Río San Juan,
Choco, S.W. Colombia. Lectotype: BMNH 1910.7.11.133, designated by Isbrücker (1979a: 113), and illustrated
in Isbrücker (1979a: 96, fig. 7).
Distribution: San Juan River basin, Colombia (Ferraris, 2003c).
Species inquirendae, Loricariidae
Microlepidogaster bourguyi Miranda Ribeiro, 1911: 90. Type locality: Brasil. Syntypes (12): possibly NMRJ 1036.
Ancistrus salgadae Fowler, 1941: 154. Fig. 57. Type locality: Rio Salgado, Icá, Ceará; eastern Brazil. Holotype: ANSP
69440. A juvenile, identification uncertain (S. Fisch-Muller, 2003).
Ancistrus mattogrossenssis Miranda Ribeiro, 1912: 5. Type locality: Not stated. Syntypes (2): Whereabouts unknown,
not found at MNRJ or MZUSP (Fisch-Muller, 2003). Original spelling of name considered to be incorrect by FischMuller (2003) and corrected the spelling to Ancistrus mattogrossensis, following Miranda Ribeiro (1918c: 114).

MALAPTERURIDAE Bleeker, 1858
Malapterurini Bleeker, 1858b: ix. Type genus: Malapterurus La Cepède, 1803.
Revision: Poll & Gosse (1969); Norris (2002).
Taxonomic summary: Gosse (1986a).
Phylogeny: Howes (1985).
Reviews: Teugels (1992b), west Africa.
2 genera, 19 species; no named fossil taxa.

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Malapterurus La Cepède, 1803: 90. Type species: Silurus electricus Gmelin, 1789. Type by designation under plenary powers; placed on Official List (ICZN Opinion 93, Direction 56). Gender: Masculine.
Anacanthus Minding 1832: 117. Type species: Silurus electricus Gmelin, 1789. Type by monotypy. Preoccupied byAnacanthus Gray, 1830, in fishes. Gender: Masculine.
Malopterurus Agassiz, 1846: 223. Type species: Silurus electricus Gmelin, 1789. Type by being a replacement
name. Unjustified emendation of Malapterurus La Cepède, 1803. Gender: Masculine.
Remarks: Malapterus Jarocki, 1822, Malapturus Swainson, 1838, and Malacopterurus Gill, 1890, are considered to
be incorrect subsequent spellings, and therefore unavailable names, in ICZN Direction 56. Forsskål (1775: 16)
applied the generic name of the electric ray, Torpedo, to his account of the electric catfish of the Nile.
Malapterurus barbatus Norris, 2002
Malapterurus barbatus Norris, 2002: 88, fig. 49. Type locality: Sierra Leone, Sewa River system, Bagbe River atYfin, 8º42'N, 11º05'W. Holotype: MNHN 1990-100.
Distribution: Western Guinean rivers from the Kolente River, Sierra Leone, to the Borlor River, Liberia (Norris,2002).
Malapterurus beninensis Murray, 1855
Malapterurus Beninensis Murray, 1855: 20, pl. Type locality: Old Calabar. Syntypes: BMNH 1856.1.16.1 (2), possibly NMSZ 1855.27 (2).
? Malapterurus affinis Günther, 1864: 220. Type locality: Old Calabar. Syntypes: BMNH 1859.11.20.1 (1), BMNH
1864.7.9.1 (1).
Distribution: Coastal plains of west central Africa, from the Volta River, Ghana, to the Shiloango River, Angola(Norris, 2002).
Remarks: Synonymy of Malapterurus affinis with M. beninensis considered tentative by Norris (2002).
Malapterurus cavalliensis Roberts, 2000
Malapterurus cavalliensis Roberts, 2000: 8. Type locality: Ivory Coast: rivière Cavally directly W of Tiéouléoula
about 18 km S of Tai. Holotype: BMNH 2000.3.3.7.
? Malapterurus thysi Norris, 2002: 81, fig. 45. Type locality: Ivory Coast, Cess (Nipoué) River at Toyebli, 6º38'N,
8º29'W. Holotype: MRAC 73-10-P-4144.
Distribution: Cess and Cavally Rivers, western Côte d’Ivoire (Norris, 2002).
Malapterurus electricus (Gmelin, 1789)
Silurus electricus Bonnaterre, 1788: 156, pl. 62 (fig. 245). Type locality: Nile and rivers of Africa. No types known.
Silurus electricus Gmelin, 1789: 1354. Type locality: Rosetta, branch of the Nile River, Egypt. Neotype: BMNH
1907.12.2.2547, designated in Poll & Gosse (1969: 5). Placed on Official list as type of Malapterurus (ICZN
Direction 57).
Distribution: Nile River basin; Western African rivers south to the Niger River basin (Norris, 2002).
Malapterurus leonensis Roberts, 2000
Malapterurus leonensis Roberts, 2000: 9. Type locality: Sierra Leone, Moyamba. Holotype: BMNH
1976.11.12.122.
? Malapterurus stiassnyae Norris, 2002: 48, fig. 23. Type locality: Sierra Leone, River Moa drainage, Tiwai Island,
7º33'N, 11º21'W. Holotype: AMNH 59488.
Distribution: Upper Guinean basins of western Africa, between the Bofon River, Guinea, and St. Paul River, Liberia(Norris, 2002).
Malapterurus melanochir Norris, 2002
Malapterurus melanochir Norris, 2002: 109, fig. 65. Type locality: Congo, Congo R. basin, Ruki River at Eala,00º04'N, 18º20'E. Holotype: MRAC 46166.
Distribution: Central Congo River basin and upper Lualaba River system (Norris, 2002).
Malapterurus microstomus Poll & Gosse, 1969
Malapterurus microstoma Poll & Gosse, 1969: 8, pls. 1, 2 (fig. b), 3 (fig. b). Type locality: Yangambi, Congo R.,
Zaire. Holotype: MRAC 164701.
Distribution: Congo River basin (Norris, 2002).

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Malapterurus minjiriya Sagua, 1987
Malapterurus minjiriya Sagua, 1987: 78, fig. 1. Type locality: Lake Kainji, lower Niger basin, Nigeria. Holotype:
BMNH 1979.3.5.405.
Distribution: Niger and Volta River basins; possibly also White Nile and Omo River (Norris, 2002).
Malapterurus monsembeensis Roberts, 2000
Malapterurus monsembeensis Roberts, 2000: 9. Type locality: Upper Congo at Monsembe. Holotype: BMNH
1899.8.22.6.
? Malapterurus gossei Norris, 2002: 102, fig. 60. Type locality: Congo, Congo R. basin, Léopoldville (Ouest) (=Kinshasa), 4º20'N, 15º15'E. Holotype: MRAC 88017.
Distribution: Congo River system (Norris, 2002).
Malapterurus occidentalis Norris, 2002
Malapterurus occidentalis Norris, 2002: 73, fig. 40. Type locality: Gambia, Mc Carthy Id. 13º32' [N], 14º46' E [sic,W]. Holotype: BMNH 1901.12.28.77.
Distribution: Middle Gambia River, Gambia, and Geba River, Guinea-Bissau (Norris, 2002).
Malapterurus oguensis Sauvage, 1879
Malapterurus electricus oguensis Sauvage, 1879: 99. Type locality: Lopé, pays des Okanda, fleuve Ogôoué. Holotype: MNHN a-0889. Originally proposed as Malapterurus electricus var. oguensis. Described in more detailand illustrated in Sauvage (1880c: 45, pl. 1, fig. 3) as Malapterurus electricus, var. ogooensis.
Distribution: Ivindo River, Ogowe River basin; and Nyanga River and Kouilou-Niari Systems of western Africa.
Malapterurus punctatus Norris, 2002
Malapterurus punctatus Norris, 2002: 91, fig. 51. Type locality: Liberia, Cavally River drainage, unspecified tributary waters of Duobe River at Duo Town, ca. 5º41'N, 8º06'W. Holotype: MRAC 80-36-P-2154.
Distribution: Eastern Guinean rivers, from the St. Paul River, Sierra Leone, to the Cavally River, Liberia (Norris,2002).
Malapterurus shirensis Roberts, 2000
Malapterurus shirensis Roberts, 2000: 10. Type locality: Chiromo, lower Shire. Holotype: BMNH 1921.9.6.53.
? Malapterurus zambeziensis Norris, 2002: 44, fig. 20. Type locality: Zambia/Zimbabwe, Zambezi River drainage,
Lake Kariba at Andora Harbor. Holotype: MRAC 187094.
Distribution: Zambezi River basin (Norris, 2002).
Malapterurus tanganyikaensis Roberts, 2000
Malapterurus tanganyikaensis Roberts, 2000: 11. Type locality: Lake Tanganyika. Holotype: BMNH
1936.6.15.1234.
Malapterurus polli Norris, 2002: 111, fig. 67. Type locality: Tanzania: au large de Karema, à 660–700 m de la côte,± 6º49'N, 30º26'E. Holotype: MRAC 90328.
Distribution: Lake Tanganyika (Norris, 2002).
Malapterurus tanoensis Roberts, 2000
Malapterurus tanoensis Roberts, 2000: 11. Type locality: Unvaiye lagoon, Tano basin, sw. Ghana. Holotype: USNM
316021.
? Malapterurus murrayi Norris, 2002: 94, fig. 54. Type locality: Ghana, Tano River near Wiawso, slow pool, 6º12'N
2º29'W. Holotype: BMNH 1969.4.28.29.
Distribution: Ofin and Tano Rivers, Ghana (Norris, 2002).
Malapterurus teugelsi Norris, 2002
Malapterurus teugelsi Norris, 2002: 52, fig. 26. Type locality: Guinea, Ndyarendi, Kogon River, ca. 11º21'N,
14º30'W. Holotype: MRAC 92-59-P-1680.
Distribution: Kogon River basin, Guinea (Norris, 2002).
PARADOXOGLANIS Norris, 2002
Paradoxoglanis Norris, 2002: 117. Type species: Paradoxoglanis caudivittatus Norris, 2002. Type by original designation. Gender: Masculine.
Paradoxoglanis caudivittatus Norris, 2002

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Paradoxoglanis caudivittatus Norris, 2002: 120, fig. 72. Type locality: Congo, Lomela, N. Sankuru, petits ruisseauxaffl. riv. Lomela, 2º00'–2º19'S, 22º58'–23º15'E. Holotype: MRAC 98452.
Distribution: Central portions of the Congo Basin (Norris, 2002).
Paradoxoglanis cryptus Norris, 2002
Paradoxoglanis cryptus Norris, 2002: 128, fig. 78. Type locality: Congo, Aketi, riv. Kagala, affl. riv. Itimbiri, rive
droite, 2º44'N, 23º46'E. Holotype: MRAC 88-25-P-2231.
Distribution: Kagal River, Itimbiri basin, Congo System (Norris, 2002).
Paradoxoglanis parvus Norris, 2002
Paradoxoglanis parvus Norris, 2002: 126, fig. 76. Type locality: Congo, Congo R. drainage, Mawuya, riv. Mawuya,
terr. de Libenge, 3º14'N 18º41'E. Holotype: MRAC 167535.
Distribution: middle Congo River basin (Norris, 2002).

MOCHOKIDAE Jordan, 1923
Synodontini Bleeker, 1862 (in Bleeker, 1862–63): 6. Type genus: Synodontis Cuvier, 1816.
Rhinoglanina Günther, 1864: 4, 10, 216. Type genus: Rhinoglanis Günther, 1864.
Mochokidae Jordan, 1923: 150. Type genus: Mochokus Joannis, 1835.
Chiloglanidinae Riehl & Baensch, 1990: 396. Type genus: Chiloglanis Peters, 1868.
Remarks: The name Simuldentinae, published in Taverne & Aloulou-Triki (1974) is not available as a family groupname because it is not based on an available generic name.
Taxonomic summary: Gosse (1986b).
Reviews: Paugy & Roberts (1992, West Africa); Skelton (1993, 2001, Southern Africa); Seegers (1996, Lake Rukwa
basin); Skelton, et al. (1985, Okavango basin).
Key to genera: Poll & Gosse (1994).
10 genera, 188 species; no named fossil taxa.
ACANTHOCLEITHRON Nichols & Griscom, 1917
Acanthocleithron Nichols & Griscom, 1917: 720. Type species: Acanthocleithron chapini Nichols & Griscom, 1917.
Type by monotypy. Gender: Neuter.
Acanthocleithron chapini Nichols & Griscom, 1917
Acanthocleithron chapini Nichols & Griscom, 1917: 721, fig. 26. Type locality: Avakubi [Ituri River], Congo. Holotype: AMNH 6548.
Distribution: Ituri River and Congo River from Kisangani to Stanley Pool (Gosse, 1986b).
ATOPOCHILUS Sauvage, 1879
Atopochilus Sauvage, 1879: 97. Type species: Atopochilus savorgnani Sauvage, 1879. Type by monotypy. Gender:
Masculine.
Atopochilus chabanaudi Pellegrin, 1938
Atopochilus Chabanaudi Pellegrin, 1938a: 375, fig. 1. Type locality: Brazzaville (Congo). Holotype: MNHN 19370111.
Distribution: Congo River at Stanley Pool (Gosse, 1986b).
Remarks: Catalog number for holotype of Atopochilus Chabanaudi mistakenly reported as MNHN 37-311 by Bertin& Estève (1950).
Atopochilus christyi Boulenger, 1920
Atopochilus Christyi Boulenger, 1920a: 33, fig. 17. Type locality: Stanley-Falls, ... d’Avakubi sur l’Ituri. Syntypes(2): Possibly BMNH 1919.9.10.282 (1), MRAC 7239 (1), MRAC 7350 (1).
Distribution: Congo River basin (Gosse, 1986b).
Atopochilus macrocephalus Boulenger, 1906
Atopochilus macrocephalus Boulenger, 1906a: 112. Type locality: Fort Don Carlos, in the province of Loanda, at the

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junction of the Cambo and Kwango (or Cuango) Rivers, Angola. Holotype: BMNH 1904.5.2.182.
Distribution: Congo River basin (Gosse, 1986b).
Atopochilus mandevillei Poll, 1959
Atopochilus mandevillei Poll, 1959: 104, pl. 18 (figs. 2a–c). Type locality: Stanley Pool. Holotype: MRAC 100503.
Distribution: Congo River at Stanley Pool (Gosse, 1986b).
Atopochilus pachychilus Pellegrin, 1924
Atopochilus pachychilus Pellegrin, 1924c: 487. Type locality: Kanda kanda, dans la Lubilonji, tributaire du Kasaï,affluent du Congo. Syntypes (2): MNHN 1924-0213 (1), MRAC 14920 (1).
Distribution: Kasai River, Congo basin; known only from type locality (Gosse, 1986b).
Atopochilus savorgnani Sauvage, 1879
Atopochilus Savorgnani Sauvage, 1879: 98. Type locality: Doumé [Fl. Ogôoué, aux chutes de Doumé (Congofrançais)]. Holotype: MNHN a-0899. Described in more detail and illustrated in Sauvage (1880c: 43. pl. 3, fig.
3).
Distribution: Upper Ogowe River, Muni River Basin, Equatorial Guinea, and Ntem River, southern Cameroon(Gosse, 1986b).
Atopochilus vogti Pellegrin, 1922
Atopochilus Vogti Pellegrin, 1922c: 350. Type locality: Riv. Wamé, Afrique orientale. Holotype: MNHN 1922-0021.
Distribution: Wami River, Tanzania (Gosse, 1986b; Seegers, 2003).
BRACHYSYNODONTIS Bleeker, 1862
Brachysynodontis Bleeker, 1862 (in Bleeker, 1862–63): 6. Type species: Synodontis batensoda Rüppell, 1832. Typeby original designation. Gender: Feminine.
Remarks: Treated in Willoughby (1994) as a synonym of Synodontis, without comment.
Brachysynodontis batensoda (Rüppell, 1832)
Synodontis batensoda Rüppell, 1832: 6, pl. 3 (fig. 2). Type locality: Nil, bei Kaïro. Holotype: SMF 2636 (dry).
Synodontes Ruppelli Swainson, 1838: 339, fig. 82. Type locality: [No locality stated]. No types known.
Distribution: Nile, Chad, Niger, Senegal and Gambia river basins (Gosse, 1986b).
Remarks: Synonymy follows Grant (2004).
CHILOGLANIS Peters, 1868
Chiloglanis Peters, 1868b: 599. Type species: Chiloglanis deckenii Peters, 1868. Type by monotypy. Gender: Masculine.
Reviews: Jubb & Le Roux (1969, southern Africa); Roberts (1989b, Cameroon), Seegers (1996, Lake Rukwa).
Remarks: The name Chiloglanis athiensis appears in Copley (1941: 15) but as name only and therefore not an available name.
Chiloglanis angolensis Poll, 1967
Chiloglanis angolensis Poll, 1967: 255, fig. 120. ruisseau à forêt-galerie, traversant la route Chibia-Jau, à la bifurcation Jau-Onguéria, quelques km au sud de la Chibia, environs de Sá de Bandeira, alt. 1.500 m., Angola. Holotype: MD 1875.
Distribution: known only from type locality (Gosse, 1986b).
Chiloglanis anoterus Crass, 1960
Chiloglanis anoterus Crass, 1960: 446, fig. 2 (a–d). Type locality: Upper Pivaan River (Pongola River system), altitude ± 4600 ft. (30º28'E, 27º30'S), Natal. Holotype: AMG P1157.
Distribution: Phongolo and Incomati River system tributaries (Skelton, 1993).
Chiloglanis asymetricaudalis De Vos, 1993
Chiloglanis asymetricaudalis De Vos, 1993: 154, figs. 2–4. Type locality: Nyakabuye, rivière Rubyiro, (bassin de la
Ruzizi), près de la confluence avec la rivière Ntandwe, Rwanda (2º34'S, 29º02'E). Holotype: MRAC 87-05-P263.
Distribution: Ruzizi River basin, Lake Tanganyika drainage (De Vos, 1993).
Chiloglanis batesii Boulenger, 1904

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Chiloglanis Batesii Boulenger, 1904a: 19. Type locality: South Cameroon; Efulen and stream tributaries of the Lobi
River, 15 or 20 miles S. W. of Efulen. Syntypes (5): BMNH 1904.7.1.97–100 (4), BMNH 1904.7.1.127 (1).
Chiloglanis micropogon Poll, 1952: 228, figs. 3–4. Type locality: riv. Nzokwe, affluent del la riv. Ulindi (Territ.
Kabare). Holotype: MRAC 91479.
Distribution: Lualaba and Kasai River, Congo Basin; Upper Niger; Meme, Mungo, Lobi Rivers and coastal rivers of
Cameroon; Kaduna River, Nigeria (Gosse, 1986b).
Remarks: Chiloglanis micropogon has been generally treated as valid, but recently treated by Roberts (1989b: 169)
as a synonym of C. batesii.
Chiloglanis benuensis Daget & Stauch, 1963
Chiloglanis benuensis Daget & Stauch, 1963: 98, fig. 3. Type locality: Bénoué à Lakdo. Holotype: MNHN 19621273.
Distribution: Niger River basin; known only from type locality (Gosse, 1986b).
Chiloglanis bifurcus Jubb & Le Roux, 1969
Chiloglanis bifurcus Jubb & Le Roux, 1969: 17, figs. 9–9a. Type locality: Crocodile River, Incomati River system,
Montrose Farm, Nelspruit District, Southern Africa. Holotype: AMG P996.
Distribution: Crocodile-Incomati basin (Skelton, 1993).
Remarks: See Kelynhans & James (1995) for comments on species.
Chiloglanis brevibarbis Boulenger, 1902
Chiloglanis brevibarbis Boulenger, 1902f: 224, pl. 17 (figs. 2, 2a, 2b). Type locality: Mathoiya River, in the Kenya
district, East Africa. Holotype: BMNH 1902.5.26.19.
Distribution: Tana System and Athi River (Gosse, 1986b).
Chiloglanis cameronensis Boulenger, 1904
Chiloglanis cameronensis Boulenger, 1904a: 18. Type locality: Efulen, South Cameroon. Syntypes (7): BMNH
1904.7.1.90–96 (7).
Distribution: coastal rivers of Cameroon, Equatorial Guinea, and Gabon, Congo River basin (Gosse, 1986b).
Chiloglanis carnosus Roberts & Stewart, 1976
Chiloglanis carnosus Roberts & Stewart, 1976: 282, pl. 8 (figs. b–d). Type locality: Near village of Inga, a few kilometers upstream and on the opposite side of the river from Inga hydroelectric dam. Lat. 5º27.5'S, Long.
13º36.0'E; lower Zaïre or Congo River. Holotype: MCZ 50464.
Distribution: Lower Congo River basin; known only from type locality (Gosse, 1986b).
Chiloglanis congicus Boulenger, 1920
Chiloglanis congicus Boulenger, 1920a: 32, fig. 16. Type locality: Stanley-Falls, Congo Belge. Holotype: MRAC
7159.
Distribution: Stanley Falls, Stanley Pool, Congo River basin (Gosse, 1986b).
Chiloglanis deckenii Peters, 1868
Chiloglanis Deckenii Peters, 1868b: 599, pl. 2. Type locality: Africa orientalis. Syntypes: ZMB 6844 (3), ZMB
16387.
Synodontis eurystomus Pfeffer, 1889: 14. Type locality: Rufu bei Korogwe. Lectotype: ZMH H387, designated by
Ladiges et al. (1958: 159).
Distribution: Kenya and Tanzania (Gosse, 1986b).
Chiloglanis disneyi Trewavas, 1974
Chiloglanis disneyi Trewavas, 1974: 365, figs. 6 (left), 8. Type locality: R. Wowe, Mungo system, Cameroon. Holotype: BMNH 1973.5.14.342.
Distribution: Wowe River, Mungo basin; Known only from type locality (Gosse, 1986b).
Chiloglanis elisabethianus Boulenger, 1915
Chiloglanis elisabethianus Boulenger, 1915: 171. Type locality: Rivière Lubumbashi, à Elisabethville. Holotype:
MRAC 12126.
Distribution: Luapula-Moero River, Congo River basin (Gosse, 1986b).
Chiloglanis emarginatus Jubb & Le Roux, 1969
Chiloglanis emarginatus Jubb & Le Roux, 1969: 21, figs. 12–12a. Type locality: Lekkerloop River, tributary of the

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Komati River of the Incomati River system, on the farm Vergelegen, Carolina District, Southern Africa. Holotype: AMG P953.
Distribution: Phongolo and Incomati Rivers, South Africa and Swaziland, and Pungwe and Zambezi Rivers, Zimbabwe (Skelton, 1993).
Chiloglanis fasciatus Pellegrin, 1936
Chiloglanis fasciatus Pellegrin, 1936a: 58. Type locality: Cubango, Angola. Holotype: MNHN 1936-0158.
Distribution: Zambezi River system, Angola and Botswana (Gosse, 1986b).
Chiloglanis harbinger Roberts, 1989
Chiloglanis harbinger Roberts, 1989b: 175, fig. 13. Type locality: Cameroun, R. Lokunje near Bipindi. Holotype:
CAS 60811.
Distribution: Lokunje River basin, southeastern Cameroon (Roberts, 1989b).
Chiloglanis kalambo Seegers, 1996
Chiloglanis kalambo Seegers, 1996: 377, figs. 279–281. Type locality: Kanyele River, a southeastern affluent of theupper Kalambo River, on road from Sumbawanga to Mbala, 34 km N of the Tanzania–Zambia border, Lake
Tanganyika drainage, Rukwa region, western Tanzania (8º25'S, 31º30'E). Holotype: MRAC 94-34-P-1079.
Distribution: Upper Kalambo River basin, Tanzania (Seegers, 1996).
Chiloglanis lamottei Daget, 1948
Chiloglanis lamottei Daget, 1948: 38, fig. 12. Type locality: Guineé Fse: Mt. Nimba, Zié, alt. 475–500 m. Holotype:
MNHN 1949-0056.
Distribution: Known only from the type locality (Gosse, 1986b).
Chiloglanis lufirae Poll, 1976
Chiloglanis lufirae Poll, 1976: 97, fig. 52. Type locality: Muye (affluent droit de la Lufira), alt. 800–900 m.
[Upemba National Park]. Holotype: MRAC 79-01-P-4728.
Distribution: Lufira River tributary; known only from type locality (Gosse, 1986b).
Chiloglanis lukugae Poll, 1944
Chiloglanis lukugae Poll, 1944a: 7, figs. 3–5. Type locality: Sange, région d’Albertville, Congo belge. Holotype:IRSNB 72; illustrated in Poll (1946: 224, fig. 23).
Distribution: Luapula River basin; tributaries of Lake Tanganyika, Angola, affl. of Cuango River (Gosse, 1986b).
Remarks: Gosse (1986b) stated that the type locality is a small tributary of the Luvua River on road between
Albertville and Kiambi but not near Albertville as has been stated elsewhere.
Chiloglanis macropterus Poll & Stewart, 1975
Chiloglanis macropterus Poll & Stewart, 1975: 152, fig. 1. Type locality: Luongo River, at ferry crossing, 53 km. S.
of Kawambwa, Zambia. Holotype: ROM 28035.
Distribution: Luongo River, Zambia (Gosse, 1986b).
Chiloglanis marlieri Poll, 1952
Chiloglanis marlieri Poll, 1952: 226, figs. 1–2. Type locality: riv. Ndakirwa à Meshe, affluent de la Luhoho. Holotype: MRAC 91478.
Distribution: Lualaba basin (Gosse, 1986b).
Chiloglanis mbozi Seegers, 1996
Chiloglanis mbozi Seegers, 1996: 224, figs. 161–162. Type locality: Hanseketwa River, a brook which is an affluentof the Momba River, draining the Mbozi block via the Msangano trough, 25 km east of Tunduma on the road to
Mbeya, Mbeya Region, western Tanzania (9º08'S, 32º52'E). Holotype: MRAC 94-34-P-927.
Distribution: Hanseketwa River, Momba River basin, Tanzania (Seegers, 1996).
Chiloglanis microps Matthes, 1965
Chiloglanis microps Matthes, 1965: 188, figs. 3–4. Type locality: riv. Lukima à Kiamakoto (Lufira). Holotype:
MRAC 140908.
Distribution: Lufira River basin (Gosse, 1986b).
Chiloglanis modjensis Boulenger, 1904
Chiloglanis modjensis Boulenger, 1904b: 332, pl. 31 (figs. 3, 3a, 3b). Type locality: Modjo River [Webi Shebelibasin]. Syntypes (2): BMNH 1905.7.25.103 (1).

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Chiloglanis neumanni Boulenger, 1911
Chiloglanis neumanni Boulenger, 1911a: 481, fig. 359. Type locality: Upper Bubu River, Masailand. Lectotype:
BMNH 1905.7.25.49; designated by, and illustrated in, Seegers (1996: 214, fig. 152).
Distribution: Limpopo River system, Cunene, Kafue, Zambezi, Kiavango, and upper Congo Rivers, Lakes Malawi
and Kariba (Gosse, 1986b; Skelton, 1993).
Chiloglanis niger Roberts, 1989
Chiloglanis niger Roberts, 1989b: 173, figs. 1f, 2f, 4e–f, 6f, 12. Type locality: Cameroun, Niger basin, R. Menchum
below high waterfall near Befang, Bamenda highlands. Holotype: CAS 60809.
Distribution: Menchum River, Niger River basin, Cameroon (Roberts, 1989b).
Chiloglanis niloticus Boulenger, 1900
Chiloglanis niloticus Boulenger, 1900d: 522. Type locality: island of Arko, Soudan. Syntypes (several): BMNH
1907.12.2.2466–73 (8), MNHN 1907-0230 (1), MSNG 14420 (1).
Distribution: Nile and Niger River basins (Gosse, 1986b).
Chiloglanis normani Pellegrin, 1933
Chiloglanis normani Pellegrin, 1933a: 113, fig. on p. 114. Type locality: Danané (Côte d’Ivoire). Syntypes (2):MNHN 1932-0301 (1), NMBA 4253 (1).
Distribution: Cavally River system; known only from type locality (Gosse, 1986b).
Chiloglanis occidentalis Pellegrin, 1933
Chiloglanis occidentalis Pellegrin, 1933a: 112, fig. on p. 112. Type locality: Douékoué, ... Man (Côte d’Ivoire). Syntypes (2): MNHN 1932-0300 (1), NMBA 4252 (1).
Chiloglanis niloticus waterloti Daget, 1954: 304, fig. 115. Type locality: Banamanan, cercle de Kissidougou[Guinea]. Syntypes: MNHN 1935-0224 (1), MNHN 1960-0472 (5), MNHN 1960-0473 (29 or 32), MNHN
1960-0474 (18 or 15).
Distribution: Bafing and Baoule rivers, upper Senegal, upper Niger, Sassandra, upper St. Paul, Kolente andKonkoure Rivers (Gosse, 1986b).
Chiloglanis paratus Crass, 1960
Chiloglanis paratus Crass, 1960: 452, fig. 4 (a–d). Type locality: Concrete wall of Pongola River barrage, altitude ±1000 ft. (31º30'E, 27º23'S), Natal. Holotype: AMG P1154.
Distribution: Phongolo, Incomati and Limpopo river systems, southern Africa (Skelton, 1993).
Chiloglanis pojeri Poll, 1944
Chiloglanis pojeri Poll, 1944a: 8, figs. 6–8. Type locality: Mambwe, région d’Albertville. Holotype: IRSNB 74;
illustrated in Poll (1946: 225, fig. 24).
Distribution: Luvua and Lualaba River, upper Congo Basin; Koki River of Lake Tanganyika (Gosse, 1986b).
Chiloglanis polyodon Norman, 1932
Chiloglanis polyodon Norman, 1932: 184, fig. 3. Type locality: Headwaters of Bagbwe River, Sierra Leone. Holotype: BMNH 1932.5.18.64.
Distribution: Bagbwe River basin, Sierra Leone; known only from type locality (Gosse, 1986b).
Chiloglanis polypogon Roberts, 1989
Chiloglanis polypogon Roberts, 1989b: 168, figs. 1b, 2b, 3c–d, 6a–b, 8. Type locality: Cameroun, Cross basin, highgradient stream along road from Mamfe to Bamenda, 88–94 km SW of Bamenda. Holotype: CAS 60790.
Distribution: Cross River basin, Cameroon (Roberts, 1989b).
Chiloglanis pretoriae van der Horst, 1931
Chiloglanis pretoriae van der Horst, 1931: 248, fig. 2. Type locality: Crocodile River, Pretoria District, Transvaal.
Holotype: at TM, current whereabouts unknown.
Chiloglanis pumilus van der Horst, 1931: 250, fig. 4. Type locality: Aapies River and Crocodile River, Pretoria Dist.,
Transvaal. Syntypes: SAIAB 30011 (1), SAIAB 30012 (1).
Distribution: Incomati, Limpopo, middle and lower Zambezi, Pungwe and Busi basins, southern Africa (Skelton,1993).
Chiloglanis reticulatus Roberts, 1989

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Chiloglanis reticulatus Roberts, 1989b: 165, figs. 1a, 2a, 3a–b, 5, 7. Type locality: Cameroun, Congo basin, R.
Mwamedjwel, a very small stream 2–3 km W of Yokadouma. Holotype: CAS 60786.
Distribution: Northwestern portion of the Congo River basin, Cameroon and Congo (Roberts, 1989b).
Chiloglanis rukwaensis Seegers, 1996
Chiloglanis rukwaensis Seegers, 1996: 227, figs. 163–164. Type locality: Chiwanda River (also spelled Chuwanda
or Chwanda River), a small river, Momba drainage, 32 km northwest of Tunduma near Chiwanda (or Chuwanda), on the road to Sumbawanga, western Lake Rukwa drainage, Mbeya Region, western Tanzania,(09º10'S, 32º33'E). Holotype: MRAC 94-34-P-932.
Distribution: Chiwanda River, Momba River basin, Tanzania (Seegers, 1996).
Chiloglanis ruziziensis De Vos, 1993
Chiloglanis ruziziensis De Vos, 1993: 162, figs. 6–7. Type locality: Nyakabuye, rivière Rubyiro (bassin de la Ruzizi), pres de la confluence avec la rivière Ntandwe, Rwanda (2º34'S, 29º02'E). Holotype: MRAC 87-05-P-264.
Distribution: Ruzizi River basin, Lake Tanganyika drainage (De Vos, 1993).
Chiloglanis sanagaensis Roberts, 1989
Chiloglanis sanagaensis Roberts, 1989b: 169, figs. 1c, 2c, 3 (e–f), 6 (c–d), 9. Type locality: Cameroun, Sanaga
basin, gravel bars and riffles in lower 1–2 km of R. Nchit where it flows into R. Mbam. Holotype: CAS 60794.
Distribution: Sanaga River basin (Roberts, 1989b).
Chiloglanis sardinhai Ladiges & Voelker, 1961
Chiloglanis sardinhai Ladiges & Voelker, 1961: 139, pl. 7 (figs. 12 and 13). Type locality: Mujije (der Mujije ist ein
Zufluss des Longa, Cuanza-Sul, Angolas). Holotype: ZMH H1317.
Distribution: Longa River basin; known only from type locality (Gosse, 1986b).
Chiloglanis somereni Whitehead, 1958
Chiloglanis somereni Whitehead, 1958: 199, figs. 2–3. Type locality: Waroya River, Nyanza Province, Kenya; at
34º30'E, 0º10'N. Holotype: BMNH 1958.7.18.1.
Distribution: Westward flowing rivers of Kenya into Lake Victoria (Gosse, 1986b).
Chiloglanis swierstrai van der Horst, 1931
Chiloglanis swierstrai van der Horst, 1931: 249, fig. 3. Type locality: Crocodile River, Pretoria District, Transvaal.
Holotype: SAIAB 30013.
Chiloglanis engiops Crass, 1960: 451, fig. 3 (a–d). Type locality: Lower Pivaan Rier (Pongola River system), altitude ± 2000 ft. (31º11'E, 27º25'S), Natal. Holotype: AMG P1156.
Distribution: Lowveld and warmer portions of Phongolo, Incomati and Limpopo river basins, southern Africa (Skelton, 1993).
Chiloglanis trilobatus Seegers, 1996
Chiloglanis trilobatus Seegers, 1996: 230, figs. 165–167. Type locality: Piti River, an affluent of the Rungwa River,
63 km S of Rungwa Village on the road from Itigi to Makambako or Mbeya, eastern Lake Rukwa drainage,
western Tanzania (07º27'S, 33º25'E). Holotype: MRAC 94-34-P-938.
Distribution: Rungwizi River basin, Lake Rukwa basin, Tanzania (Seegers, 1996).
Chiloglanis voltae Daget & Stauch, 1963
Chiloglanis voltae Daget & Stauch, 1963: 99, fig. 4. Type locality: Bougouri Ba au pont de Nabéré. Holotype:
MNHN 1962-1280.
Distribution: Volta and upper Bénoué River systems (Gosse, 1986b).
EUCHILICHTHYS Boulenger, 1900
Euchilichthys Boulenger, 1900d: 522. Type species: Atopochilus guentheri Schilthuis, 1891. Type by subsequent
designation by Jordan (1920: 488). Gender: Masculine.
Euchilichthys astatodon (Pellegrin, 1928)
Atopochilus astatodon Pellegrin, 1928b: 107, fig. 1. Type locality: Luluaburg Saint-Joseph, Congo belge. Syntypes(6): MNHN 1928-0017 (1), MNHN 1928-0018 (1), MRAC 19942 (1), NMBA 3721 (1), NMBA 3723–24 (2),
NMBA 3726 (1).
Distribution: Kasai River (Gosse, 1986b).

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Euchilichthys boulengeri Nichols & La Monte, 1934: 3, fig. 3. Type locality: Luluabourg, Kasai District, Belgian
Congo. Holotype: AMNH 12357.
Distribution: Kasai River (Gosse, 1986b).
Euchilichthys dybowskii (Vaillant, 1892)
Chiloglanis Dybowskii Vaillant, 1892b: 2. Type locality: l’Oubanghi. Syntypes: MNHN 1892-0081 (2).
Euchilichthys habereri Steindachner, 1912: 447. Type locality: Dscha, Süd. Kamerun. Holotype: NMW 10700.
Illustrated and described in more detail in Steindachner (1913: 45, fig. 9 and pl. 1 (figs. 2–2a).
Distribution: Ja River, Congo basin, Cameroon (Gosse, 1986b).
Euchilichthys guentheri (Schilthuis, 1891)
Atopochilus güntheri Schilthuis, 1891: 86, pl. 6 (fig. 2). Type locality: Stanley Pool. Holotype: BMNH 1899.9.6.6.
Distribution: Congo River basin, up to Bangweulu River system (Gosse, 1986b).
Euchilichthys royauxi Boulenger, 1902
Euchilichthys royauxi Boulenger, 1902d: 46, pl. 13. Type locality: l’Ubangi à Banzyville. Syntypes (3): BMNH
1901.12.26.46–47 (2), MRAC 1186 (1).
Distribution: Congo River basin, up to Moero Lake system (Gosse, 1986b).
HEMISYNODONTIS Bleeker, 1862
Hemisynodontis Bleeker, 1862 (in Bleeker, 1862–63): 6. Type species: Pimelodus membranaceus Geoffroy St.
Hilaire, 1809. Type by original designation. Gender: Feminine.
Remarks: Treated in Willoughby (1994) as a synonym of Synodontis, without comment.
Hemisynodontis membranacea (Geoffroy Saint-Hilaire, 1809)
Pimelodus membranaceus Geoffroy Saint-Hilaire, 1809: pl. 13 (figs. 1–2). Type locality: Fl. Nil (Egypte). Holotype:
MNHN 0000-4197. Name available from caption on plate, described as Synodontis membranaceus in Geoffroy
Saint-Hilaire (1827: 297).
Synodontis Guentheri Vaillant, 1893a: 16. Type locality: Karthoum. Syntypes: BMNH 1862.6.17.160 (1), BMNH
1862.6.17.161–162 (2), BMNH 1865.11.15.13 (1).
Distribution: Nile, Niger, Senegal, Gambia, and Volta River basins, Chad system (Gosse, 1986b).
MICROSYNODONTIS Boulenger, 1903
Microsynodontis Boulenger, 1903e: 26. Type species: Microsynodontis batesii Boulenger, 1903. Type by monotypy.
Gender: Feminine.
Microsynodontis armatus Ng, 2004
Microsynodontis armatus Ng, 2004h: 10, fig. 6. Type locality: Gabon: Ogooué-Ivindo province, Ivindo River drainage, Balé Creek, 0º31'19"N, 12º47'58"E. Holotype: CU 89392.
Distribution: Ivindo River basin, Gabon (Ng, 2004h: 15).
Microsynodontis batesii Boulenger, 1903
Microsynodontis batesii Boulenger, 1903e: 26, pl. 4. Type locality: Mvile River, southern Cameroon. Syntypes (several; apparently more than 6, from notes found in BMNH register): BMNH 1903.7.28.105–110 (6).
Microsynodontis Christyi Boulenger, 1920a: 32, fig. 15. Type locality: Poko, Congo Belge. Syntypes (4): BMNH
1919.9.10.281 (1), MRAC 7168–70 (3).
Distribution: Ntem River basin, southern Cameroon and northern Gabon, and Campo. Ivindo, Lobé, Nyong and Sanaga River basins, southern and central Cameroon (Ng, 2004h: 9).
Remarks: Ng (2004h: 2) stated that Microsynodontis christyi should be treated as valid, but provided no further comments. The BMNH syntype of M. christyi has had an incorrect jar label, with the registration number listed as
BMNH 1919.9.10.381.
Microsynodontis emarginata Ng, 2004
Microsynodontis emarginatus Ng, 2004h: 15, fig. 8. Type locality: Gabon: Haut-Ogooué Province, Motobo I village,
Kiéne creek, 1º32'14.1"S, 13º32'43.5"E. Holotype: CU 89393.
Distribution: Ogooué River basin, Gabon (Ng, 2004h: 19).

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Microsynodontis hirsuta Ng, 2004
Microsynodontis hirsutus Ng, 2004h: 20, fig. 10. Type locality: Gabon: Woleu-Ntem province, Ngomo creek, where
it crosses Oyem-Minvoul road, 1º41'30.0"N, 11º39'18.9"E. Holotype: CU 87040.
Distribution: Nye River of Ntem River basin, Gabon (Ng, 2004h: 23).
Microsynodontis laevigata Ng, 2004
Microsynodontis laevigatus Ng, 2004h: 24, fig. 11. Type locality: Gabon: Ogooué-Ivindo province, Ivindo Riverdrainage, small creek flowing into Ivindo River, Makokou, 0º35'8"N, 12º51'22"E. Holotype: CU 88265.
Distribution: Ivindo River basin, Gabon (Ng, 2004h: 27).
Microsynodontis lamberti Poll & Gosse, 1963
Microsynodontis lamberti Poll & Gosse, 1963: 61, pl. 4 (fig. 1). Type locality: Rivière Lilanda, Yangole, Congo centrale. Holotype: MRAC 137837.
Distribution: Lilanda River, Congo River basin; known only from type locality (Gosse, 1986b).
Microsynodontis nannoculus Ng, 2004
Microsynodontis nannoculus Ng, 2004h: 27, fig. 12. Type locality: Equatorial Guinea: Mami River, a tributary of
Kyé River. Holotype: MRAC 173145.
Distribution: Kyé River basin, Ntem River drainage, Equatorial Guinea (Ng, 2004h: 31).
Microsynodontis nasutus Ng, 2004
Microsynodontis nasutus Ng, 2004h: 32, fig. 14. Type locality: Gabon: Woleu-Ntem province, Okano River on rapids 0.5 km S of village of Na, 0º48'35"N, 11º38'47"E. Holotype: CU 89394.
Distribution: Okano River, Ogooué River basin, Gabon (Ng, 2004h: 35).
Microsynodontis notata Ng, 2004
Microsynodontis notatus Ng, 2004h: 35, fig. 15. Type locality: Gabon: Ezanga River, about midway between Lake
Ezanga and Ogooué mainstream. Holotype: MRAC 80-51-P-839.
Distribution: Lower Ogooué River, Gabon (Ng, 2004h: 39).
Microsynodontis polli Lambert, 1958
Microsynodontis polli Lambert, 1958: 42, fig. 1. Type locality: Rivière Gbin, Guinée Forestière, Guinée Française.
Holotype: MRAC 119457.
Distribution: Gbin River, Upper Niger; known only from type locality (Gosse, 1986b).
Microsynodontis vigilis Ng, 2004
Microsynodontis vigilis Ng, 2004h: 39, fig. 16. Type locality: Gabon: Moyen-Ogooué province, 12 km N of Lambaréné, branch of Nzorbang creek near village of same name, 0º34'6.2"N, 10º12'46.4"E. Holotype: CU 87039.
Distribution: Lower Ogooué River basin, Gabon (Ng, 2004h: 43).
MOCHOKIELLA Howes, 1980
Mochokiella Howes, 1980: 165. Type species: Mochokiella paynei Howes, 1980. Type by original designation. Gender: Feminine.
Mochokiella paynei Howes, 1980
Mochokiella paynei Howes, 1980: 165, figs. 1, 2b. Type locality: Kassawe Forest Reserve, Sierra Leone. Holotype:
BMNH 1979.8.22.1.
Distribution: Kassawe Forest Reserve, Sierra Leone; known only from type locality (Gosse, 1986b).
MOCHOKUS Joannis, 1835
Mochokus Joannis, 1835: [Cl. IV, pl. 8]. Type species: Mochokus niloticus Joannis, 1835. Type by monotypy. Gender: Masculine.
Mochocus Günther, 1864: 217. Type species: Mochokus niloticus Joannis, 1835. Gender: Masculine. Unjustified
emendation of Mochokus Joannis, 1835.
Rhinoglanis Günther, 1864: 216. Type species: Rhinoglanis typus Günther, 1864. Type by original designation. Gender: Masculine.
Mochokus brevis Boulenger, 1906
Mochocus brevis Boulenger, 1906d: 148. Type locality: Fashoda and Lake No, White Nile. Syntypes (43): BMNH

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1907.12.2.2529–2542 (14), BMNH 1907.12.2.2544 (1), MNHN 1907-0233 (2).
Distribution: Nile and Chad basins (Gosse, 1986b).
Mochokus niloticus Joannis, 1835
Mochokus Niloticus Joannis, 1835: [Cl. IV, pl. 8]. Type locality: Nil. Holotype: Whereabouts unknown; not found in
MNHN by Boulenger (1900d: 526) or subsequently.
Rhinoglanis typus Günther, 1864: 216, unnumbered figure. Type locality: Gondokoro. Holotype: BMNH
1981.4.13.1.
Rhinoglanis Vannutellii Vinciguerra, 1898: 254. Type locality: lago Rodolfo. Syntypes: MSNG 14419 (4).
Distribution: Nile and Niger basins (Gosse, 1986b).
SYNODONTIS Cuvier, 1816
Synodontis Cuvier, 1816: 203. Type species: Silurus clarias Linnaeus, 1758. Type by subsequent designation by
Bleeker, 1862 (in Bleeker, 1862–63): 6. Gender: Feminine.
Leiosynodontis Bleeker, 1862 (in Bleeker, 1862–63): 6. Type species: Synodontis maculosus Rüppell, 1829. Type byoriginal designation. Gender: Feminine.
Pseudosynodontis Bleeker, 1862 (in Bleeker, 1862–63): 6. Type species: Synodontis serratus Rüppell, 1829. Typeby original designation. Gender: Feminine.
Revision: Vaillant (1895b, 1896b); Poll (1971, with keys to species).
Reviews: Willoughby (1994, Lake Kainji, Nigeria); Bruwer & van der Bank (2003, southern Africa).
Keys: Skelton & White (1990, southern Africa); Bruwer & van der Bank (2003).
Remarks: The name Synodontes published in Minding (1832: 116) is considered an incorrect subsequent spellingand not an available name.
Synodontis acanthomias Boulenger, 1899
Synodontis acanthomias Boulenger, 1899a: 46, pl. 23. Type locality: Boma et Léopoldville [restricted to Boma bylectotype designation]. Lectotype: MRAC 18, designated by Poll (1971: 282).
Synodontis pfefferi Steindachner, 1912: 447. Type locality: Dscha, Nebenfluss des Congo im Bezirke Molundu dessüdlichen Kamerun. Holotype: NMW 9023. Species illustrated and described in more detail in Steindachner(1913: 43, pl. 8).
Distribution: Congo River basin, except central forest (Gosse, 1986b).
Synodontis afrofischeri Hilgendorf, 1888
Synodontis Afro-Fischeri Hilgendorf, 1888: 77. Type locality: Victoria-Nyanza (Ukerewe-See). Holotype: ZMB
12745. Holotype illustrated in Poll (1971: fig. 28) and Seegers (1996: fig. 171).
Distribution: Lakes Victoria, Nabugabo, Kioga, Ihema; Nile, Kagera, Kingani Malagarasi Rivers (Gosse, 1986b);
Piti River, Lake Rukwa basin (Seegers, 1996).
Synodontis alberti Schilthuis, 1891
Synodontis alberti Schilthuis, 1891: 88. Type locality: Kinshasa, Stanley Pool. Lectotype: BMNH 1899.9.6.9, designated by, and illustrated in, Poll (1971: 252, fig. 114).
Distribution: Congo River basin (Gosse, 1986b).
Synodontis albolineata Pellegrin, 1924
Synodontis albolineatus Pellegrin, 1924b: 320, fig. 1. Type locality: Madjingo, rivière Djoua (Gabon). Holotype:
MNHN 1924-0147, illustrated in Poll (1971: fig. 68).
Distribution: Ogowe River basin; known only from type locality (Gosse, 1986b).
Synodontis angelica Schilthuis, 1891
Synodontis angelica Schilthuis, 1891: 87. Type locality: Léopoldville, Stanley Pool. Holotype: BMNH 1899.9.6.8;
holotype illustrated in Poll (1971: fig. 110, top).
Synodontis tholloni Boulenger, 1901a: 304. Type locality: bassin de l’Ogooué (?). Holotype: MNHN 1890-0030.
Synodontis angelicus zonatus Poll, 1933: 134, fig. 11. Type locality: Lukulu (riv. Lukulu). Holotype: MRAC 37904;
holotype illustrated in Poll (1971: fig. 110, bottom). Originally as Synodontis angelicus var. zonatus.
Synodontis werneri Boeseman, 1957: 146, pl. 5. Type locality: Stanley Pool, Belgian Congo. Holotype: RMNH
20913.

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Distribution: Congo River basin, except in central forest (Gosse, 1986b).
Synodontis annectens Boulenger, 1911
Synodontis annectens Boulenger, 1911d: 56. Type locality: Nianimaru, Gambia; Culufi and Geba Rivers, Portuguese
Guinea [restricted to Nianimaru by lectotype designation]. Lectotype: BMNH 1901.12.28.60, designated byPoll (1971: 131).
Distribution: Gambia; Geba basin, Guinea Bissau; and Taja basin, Sierra Leone (Gosse, 1986b).
Synodontis ansorgii Boulenger, 1911
Synodontis ansorgii Boulenger, 1911b: 375. Type locality: Geba and Culufi Rivers, Portuguese Guinea [restricted toGeba River, by lectotype designation]. Lectotype: BMNH 1912.4.1.128, designated by,and illustrated in, Poll
(1971: 135, fig. 56).
Distribution: Geba River basin, Guinnea-Bissau; and Taja River basin, Sierra Leone (Gosse, 1986b).
Synodontis arnoulti Roman, 1966
Synodontis arnoulti Roman, 1966: 141, pl. 7 (fig. 14). Type locality: le Kou, affluent de la Volta Noire, à 30 km au
Nord de Bobo-Dioulasso. Holotype: MRAC 141742.
Distribution: Volta River basin (Gosse, 1986b).
Synodontis aterrima Poll & Roberts, 1968
Synodontis aterrimus Poll & Roberts, 1968: 297, figure on p. 298. Type locality: Bokuma, Bassin du Congo. Holotype: MRAC 101466.
Distribution: Central Congo River basin (Gosse, 1986b).
Synodontis bastiani Daget, 1948
Synodontis bastiani Daget, 1948: 35, fig. 10. Type locality: Côte d’Ivoire: Bouaflé. Holotype: MNHN 1949-0055.
Synodontis dageti Poll & Roman, 1967: 185, fig. 3. Type locality: [68 km south of Bolo-Dioulasso], Haut Comoé.
Holotype: MRAC 141982.
Synodontis eburneensis Daget, 1965: 473. Type locality: le Bandama au lieu dit Lamto, à mi-chemin entre Toumodi
et Tiassalé (Côte d’Ivoire). Holotype: MNHN 1964-0266; holotype illustrated in Poll (1971: fig. 92).
Distribution: Sassandra, Bandama, Bia, and Comoré basins, Côte d’Ivoire (Gosse, 1986b).
Remarks: Synonymy of these three names first proposed by Paugy & Roberts (1992).
Synodontis batesii Boulenger, 1907
Synodontis Batesii Boulenger, 1907b: 50. Type locality: Ja River, South Cameroon. Lectotype: BMNH
1907.5.22.179; designated by, and illustrated in, Poll (1971: 331, fig. 150).
Distribution: Central Congo River basin; Ja River and Nyong River, Cameroon; Muni River, Equatorial Guinea(Gosse, 1986b).
Synodontis brichardi Poll, 1959
Synodontis brichardi Poll, 1959: 100, pl. 17 (figs. 2 a–c). Type locality: Léopoldville, début des rapides à Kinsuka.
Holotype: MRAC 100494.
Distribution: Lower Congo River rapids (Gosse, 1986b).
Synodontis budgetti Boulenger, 1911
Synodontis budgetti Boulenger, 1911a: 403, fig. 305. Type locality: Lokoja, Upper Niger [Nigeria]. Holotype:
BMNH 1904.1.20.42.
Distribution: Niger and Oueme Rivers, Nokuoue Lake (Gosse, 1986b).
Synodontis camelopardalis Poll, 1971
Synodontis camelopardalis Poll, 1971: 430, fig. 202; pls. 7 (fig. 4), 12 (fig. 14). Type locality: Eala, riv. Ruki. Holotype: MRAC 47654.
Distribution: Central Congo River basin; known only from type locality (Gosse, 1986b).
Synodontis caudalis Boulenger, 1899
Synodontis caudalis Boulenger, 1899a: 44, pl. 22 (fig. 1). Type locality: Boma, Matadi, Léopoldville [restricted to
Matadi by lectotype designation]. Lectotype: MRAC 57, designated by, and illustrated in, Poll (1971: 274, fig.
124).
Distribution: (Gosse, 1986b).
Synodontis caudovittata Boulenger, 1901

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Synodontis caudovittatus Boulenger, 1901c: 10. Type locality: Mouth of Lake No, White Nile. Lectotype: BMNH
1907.12.2.2360, designated by, and illustrated in, Poll (1971: 104).
Distribution: Nile River basin (Gosse, 1986b).
Synodontis centralis Poll, 1971
Synodontis centralis Poll, 1971: 435, fig. 204; pls. 7(fig. 7), 12 (fig. 12). Type locality: Ndwa village, Kunungu,
Zaire. Holotype: MRAC 57198.
Distribution: Central Congo River basin (Gosse, 1986b).
Synodontis clarias (Linnaeus, 1758)
Silurus clarias Linnaeus, 1758: 306. Type locality: habitat in Americae, Africae fluviis. Types: See Remarks, below.
Silurus callarias Bloch & Schneider, 1801: 379. Type locality: in fluviis Americae australis et Nilo. Syntypes: on
Silurus clarias Linnaeus, and Bloch (1782: pl. 35, figs. 1–2). An unneeded new name for Silurus clarias Linnaeus, 1758.
Pimelodus synodontis Geoffroy Saint-Hilaire, 1809: pl. 12 (figs. 5–6). Type locality: Nil. Holotype: MNHN 00004194. Name available on caption to plate. Species described as Synodontis macrodon by Geoffroy Saint-Hilaire(1827: 295).
Synodontis macrodon Geoffroy Saint-Hilaire, 1827: 295. Type locality: Fl. Nil (Egypte). Holotype: MNHN 00004194. New name for Pimelodus synodontis Geoffroy Saint-Hilaire, 1809.
Distribution: Nile, Niger, Senegal, Gambia, and Volta River basins, Chad system (Gosse, 1986b).
Remarks: Silfvergrip (1996: 8–9) discussed the specimens that formed the type series for Silurus clarias. Two of thefour type specimens are extant, neither of which is a species of Synodontis. Silfvergrip (1996: 9) selected “Hasselquist’s specimen” (NRM 7044), which was subsequently identified as a specimen of Bagrus docmak (Silfvergrip, pers. commun., 2003), as the lectotype of Silurus clarias. The nomenclatural impact of this selection
would result in Bagrus docmak (Forsskål, 1775) becoming a junior synonym of Silurus clarias, and the genera
Synodontis Cuvier, 1816, and Bagrus Bosc, 1816, becoming synonyms. Because of the nomenclatural impact ofSilfvergrip’s lectotype designation on two relatively well known African catfishes, his designation is not followed here but it left for a more thorough review in the future.
Synodontis comoensis Daget & Lévêque, 1981
Synodontis comoensis Daget & Lévêque, 1981: 45, figs. 1–5. Type locality: rivière Comoé au niveau du bac deGansé, Parc National de la Comoé (Côte d’Ivoire), 8º39'N, 3º56'W. Holotype: MNHN 1980-1636.
Distribution: Comoé River, Côte d’Ivoire; known only from type locality (Gosse, 1986b).
Synodontis congica Poll, 1971
Synodontis congicus Poll, 1971: 440, fig. 206; pls. 7 (fig. 6), 12 (fig. 16). Type locality: Gangala na Bodio, riv.
Dungu. Holotype: MRAC 165695.
Distribution: Congo River basin, except lower Congo, central basin, Luapula-Moero, and high Katanga (Gosse,
1986b).
Synodontis contracta Vinciguerra, 1928
Synodontis contractus Vinciguerra, 1928: 24, pl. 1. Type locality: fiume Rubi a Buta [upper Congo basin]. Lectotype: MSNG 26493, designated by Poll (1971: 383).
Synodontis davidi Axelrod, 1970: 33, figs. Type locality: Stanley Pool near Kinshasha, Congo. Type(s): Whereabouts unknown.
Distribution: Congo River basin (Gosse, 1986b).
Synodontis courteti Pellegrin, 1906
Synodontis Courteti Pellegrin, 1906b: 473. Type locality: Fort-Archambault (Chari) [Soudan]. Holotype: MNHN
1904-0083; illustrated in Poll (1971: 129, fig. 54).
Synodontis thomasi Chabanaud, 1934: 219. Type locality: Afrique équatorial française. Type(s) (510 mm): Whereabouts unknown.
Distribution: Chad and Niger basins (Gosse, 1986b).
Synodontis cuangoana Poll, 1971
Synodontis cuangoanus Poll, 1971: 446, fig. 208; pl. 7 (fig. 10), pl. 12 (fig. 17). Type locality: Cuango, Cafunfo(8º47'S, 18º01'E). Holotype: MRAC 172253.

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Distribution: Congo River basin, south of Kasai, Zaire and Angola (Gosse, 1986b).
Synodontis decora Boulenger, 1899
Synodontis decorus Boulenger, 1899a: 49, pl. 25. Type locality: Léopoldville et Nouvelle-Anvers; [restricted to
Nouvelle-Anvers by lectotype designation]. Lectotype: MRAC 154, designated by Poll (1971: 291).
Synodontis vittatus Boulenger, 1920a: 31, pl. 25. Type locality: Stanleyville. Lectotype: MRAC 7119, designated byPoll (1971: 295).
Distribution: Congo River basin, except Luapula River system (Gosse, 1986b).
Synodontis dekimpei Paugy, 1987
Synodontis dekimpei Paugy, 1987: 357, fig. 1. Type locality: rivière Konkouré à Soukya (aval du pont routier Kindia-Telimélé en amont de Tondo) (Guinée). Holotype: MRAC 81-20-P-11.
Distribution: Konkouré River, Guinea (Paugy, 1987).
Synodontis depauwi Boulenger, 1899
Synodontis depauwi Boulenger, 1899a: 45, pl. 22 (fig. 2). Type locality: Léopoldville. Lectotype: MRAC 111, designated by Poll (1971: 278).
Distribution: Congo River at Stanley Pool (Gosse, 1986b).
Synodontis dhonti Boulenger, 1917
Synodontis dhonti Boulenger, 1917b: 367. Type locality: Kilewa Bay [Lake Tanganyika]. Holotype: MRAC 14344;
holotype illustrated in Poll (1946: 211, figs. 21–22, and 1971: fig. 168).
Synodontis irsacae Matthes, 1959b: 78. Type locality: Kalundu, nord du Lac Tanganyika. Holotype: MRAC 130315.
Distribution: Lake Tanganyika (Gosse, 1986b).
Synodontis dorsomaculata Poll, 1971
Synodontis dorsomaculatus Poll, 1971: 449, fig. 210; pl. 7 (fig. 8), pl. 12 (fig. 18). Type locality: Kadia, Kisale.
Holotype: MRAC 71395.
Distribution: Upper Lualaba (Katanga), Congo River basin (Gosse, 1986b).
Synodontis euptera Boulenger, 1901
Synodontis eupterus Boulenger, 1901c: 11. Type locality: Mouth of Lake No, White Nile. Holotype (147 mm SL):BMNH 1907.12.2.2376; holotype redescribed and illustrated in Boulenger (1907a: 370, pl. 68), with locality as
Goz-abu-Gumah, White Nile; holotype also illustrated in Poll (1971: fig. 44).
Synodontis macrepipterus Pellegrin, 1922b: 222. Type locality: Fort Crampel (Gribingui). Holotype: MNHN 19220031.
Distribution: White Nile, Chad Basin (Gribinqui), Niger and Volta Rivers (Gosse, 1986b).
Synodontis fascipinna Nichols & La Monte, 1953
Synodontis fascipinna Nichols & La Monte, 1953: 2, fig. 2. Type locality: Birao, eastern French Equatorial Africa.
Holotype: AMNH 19841.
Distribution: Headwaters of the Chari River basin; known only from type locality (Gosse, 1986b).
Synodontis filamentosa Boulenger, 1901
Synodontis filamentosus Boulenger, 1901c: 10. Type locality: Mouth of Lake No, White Nile. Holotype: BMNH
1907.12.2.2377; holotype illustrated in Boulenger (1907: pl. 69) and Poll (1971: fig. 42).
Synodontis augierasi Pellegrin, 1929a: 136, fig.1. Type locality: [Niger R. at Koulikoro, Mali, w. Africa.] Syntypes:
MNHN 1928-0243 (1), MNHN 1928-0244 (1).
Distribution: Nile, Chad, Niger, and Volta basins (Gosse, 1986b).
Synodontis flavitaeniata Boulenger, 1919
Synodontis flavitaeniatus Boulenger, 1919: 161. Type locality: Eala [Congo River basin]. Holotype: MRAC 1369;
holotype illustrated in Poll (1971: fig. 170).
Distribution: Central Congo River basin (Gosse, 1986b).
Synodontis frontosa Vaillant, 1895
Synodontis frontosus Vaillant, 1895a: 48. Type locality: Nil [White Nile, Sudan]. Holotype: MNHN 0000-4208.
Illustrated in Vaillant (1895b: pl. 10, fig. 8) and described in more detail in Vaillant (1896b: 146).
Synodontis Citernii Vinciguerra, 1898: 247. Type locality: Lago Rodolfo. Holotype: MSNG 14423 (dry).
Distribution: Nile basin; Lakes Rudolf, Albert, Eduard and Kioga; Chad basin; Uebi Giupa (Uebi Shebeli) (Gosse,

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1986b).
Synodontis fuelleborni Hilgendorf & Pappenheim, 1903
Synodontis fuelleborni Hilgendorf & Pappenheim, 1903: 265. Type locality: Rukwa-See. Lectotype: ZMB 16309,
designated by, and illustrated in, Poll (1971: 320, fig. 146); lectotype also illustrated in Seegers (1996: fig. 177).
Distribution: Rukwa Lake and Rufigi basin (Gosse, 1986b).
Synodontis gambiensis Günther, 1864
Synodontis gambiensis Günther, 1864: 214. Type locality: Gambia. Holotype: BMNH 2005.9.9.1; holotype illustrated in Poll (1971: fig. 20).
Synodontis gambiensis latifrons Blache, 1964: 201, fig. 112. Type locality: Lake Tchad basin. Syntypes: Possibly
MNHN 1959-0495 (2), MNHN 1959-0500 (5), MNHN 1959-0524 (8).
Distribution: Niger, Chad, Gambia, Geba, Taja, Waange, Volta, and Oueme basins (Gosse, 1986b).
Remarks: Considered to be a synonym of Synodontis schall by Lévêque et al. (1989: 120) and Paugy (in Lévêque etal., 1992: 561). Register number for holotype of Synodontis gambiensis Günther, given in Poll (1971) andEschmeyer et al. (1998) as BMNH 1863.16.4.40, is incorrect; it not a possible register number in the BMNH
system.
Synodontis geledensis Günther, 1896
Synodontis geledensis Günther, 1896b: 220, fig. 2. Type locality: Geledi on the Shebeli. Holotype: BMNH
1895.12.31.69.
Distribution: Uebi Shebeli, Somalia; Lorian Swamps and Uasso Nyiro (Uebi Giuba basin), Kenya (Gosse, 1986b).
Synodontis gobroni Daget, 1954
Synodontis gobroni Daget, 1954: 301, fig. 113. Type locality: Mopti, Markala. Syntypes (4): MNHN 1954-0005 (1),
MNHN1960-0471 (1).
Distribution: Niger River basin (Gosse, 1986b).
Remarks: Treated in Willoughby (1994: 28) as a synonym of Synodontis guttatus Günther, 1865.
Synodontis granulosa Boulenger, 1900
Synodontis granulosus Boulenger, 1900c: 480. Type locality: North end of Lake Tanganyika. Lectotype: BMNH
1906.9.6.40, designated by, and illustrated in, Poll (1971: 303, fig. 138).
Distribution: Lake Tanganyika (Gosse, 1986b).
Synodontis greshoffi Schilthuis, 1891
Synodontis greshoffi Schilthuis, 1891: 87. Type locality: Kinshasa, Stanley Pool. Holotype: BMNH 1899.9.6.7;
holotype illustrated in Poll (1971: fig. 112).
Synodontis holopercnus Boulenger, 1920a: 28. Type locality: Bosabangi, Avakubi,& Poko, Congo Belge. Syntypes(5): BMNH 1919.9.10.264-265 (2), MRAC 7616 (1), MRAC 7775 (1), MRAC 7829 (1).
Synodontis pantherinus Boulenger, 1920b: 149. Type locality: Ituri à Avakubi. Holotype: MRAC 14513.
Distribution: Congo River basin, except Luapula River system and lower Congo (Gosse, 1986b).
Synodontis guttata Günther, 1865
Synodontis guttatus Günther, 1865a: 452. Type locality: Niger. Holotype: BMNH 1865.4.6.4 (29 inches, dry); holotype illustrated in Boulenger (1911a: fig. 335) and Poll (1971: fig. 24)
Distribution: Lower Niger basin (Gosse, 1986b).
Synodontis haugi Pellegrin, 1906
Synodontis Haugi Pellegrin, 1906a: 470. Type locality: Ngomo (Ogôoué). Holotype: MNHN 1906-0209; holotype
illustrated in Poll (1971: fig. 52).
Distribution: Ogowe River basin (Gosse, 1986b).
Synodontis iturii Steindachner, 1911
Synodontis melanostictus iturii Steindachner, 1911c: 534. Type locality: Ituriflüsse. Holotype: NMW 46049; holotype illustrated in Poll (1971: fig. 160). Originally as Synodontis melanostictus var. iturii.
Synodontis ituriensis Boulenger, 1920a: 28. Type locality: Mawambi sur l’Ituri, Congo Belge. Holotype: MRAC
7576.
Distribution: Ituri River (Gosse, 1986b).
Synodontis katangae Poll, 1971

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Synodontis katangae Poll, 1971: 454, fig. 212; pls. 7 (fig. 11), 12 (fig. 19). Type locality: Lukonzolwa, lac Moëro.
Holotype: MRAC 14231.
Distribution: Congo River basin, high Katanga and Luapula-Moero (Gosse, 1986b).
Synodontis khartoumensis Abu Gideiri, 1967
Synodontis khartoumensis Abu Gideiri, 1967: 133, fig. 1. Type locality: Blue Nile, two miles from place where theBlue Nile joins the White Nile to form the main Nile at Khartoum, Sudan. Holotype: at Sudan Natural History
Museum.
Distribution: Nile basin and Lake Albert (Gosse, 1986b).
Synodontis koensis Pellegrin, 1933
Synodontis koensis Pellegrin, 1933a: 110, fig. on p. 111. Type locality: Man (rivière Ko). Holotype: NMBA 4500;
holotype illustrated in Poll (1971: fig. 72).
Distribution: Nzo River of Sassandra River basin, Côte d’Ivoire (Gosse, 1986b).
Synodontis laessoei Norman, 1923
Synodontis laessoei Norman, 1923c: 696. Type locality: Kokema River, a tributary of the Quanza River, Angola;
altitude 4000 ft. Holotype: BMNH 1923.8.15.3; holotype illustrated in Poll (1971: fig. 176).
Distribution: Kolema River of Quanza River basin, Angola; known only from type locality (Gosse, 1986b).
Synodontis leopardina Pellegrin, 1914
Synodontis leopardinus Pellegrin, 1914: 26. Type locality: Pays des Barotsés (Ht.-Zambèze). Holotype: MNHN
1913-0321; holotype illustrated in Poll (1971: fig. 162).
Synodontis jallae Gilchrist & Thompson, 1917: 561. Type locality: Shesheke, S. Rhodesia. Holotype: SAM 14290[now at AMG].
Distribution: Cunene, Upper Zambezi and Okovango river basins (Skelton, 1993).
Synodontis leoparda Pfeffer, 1896
Synodontis leopardus Pfeffer, 1896: 35. Type locality: Rufu bei Korogwe. Holotype: at ZMH (Gosse, 1986: 132).
Distribution: Coastal rivers of Tanzania and Somalia (Gosse, 1986b).
Remarks: Considered by De Vos (2001a: 49) as a possible synonym of Synodontis zanzibaricus Peters, 1868.
Synodontis levequei Paugy, 1987
Synodontis levequei Paugy, 1987: 361, fig. 3. Type locality: rivière Kakrima (basin du Konkouré), près de Kasséri(Guinée). Holotype: MNHN 1987-0931.
Distribution: Kakrima River (Paugy, 1987).
Synodontis longirostris Boulenger, 1902
Synodontis longirostris Boulenger, 1902d: 44, pl. 12 (fig. 2). Type locality: Rivière Yembe à Banzyville. Holotype:
MRAC 1243.
Synodontis ovidius Lönnberg & Rendahl, 1920: 173, figs 4–5. Type locality: Mukimbungu, Lower Congo. Holotype: NRM 9913.
Distribution: Congo River basin (Gosse, 1986b).
Synodontis longispinis Pellegrin, 1930
Synodontis Batesi longispinis Pellegrin, 1930: 207. Type locality: la Sangha. Holotype: MNHN 1929-0245; holotype illustrated in Poll (1971: fig. 182). Originally as Synodontis Batesi var. longispinis nov. var.
Distribution: Sangha River, Congo River basin (Gosse, 1986b).
Synodontis lufirae Poll, 1971
Synodontis lufirae Poll, 1971: 459, fig. 214; pl. 7 (fig. 12). Type locality: Lac de Koni, Lufira. Holotype: MRAC
168400.
Distribution: Lufira River, Congo River basin (Gosse, 1986b).
Synodontis macrophthalma Poll, 1971
Synodontis macrophthalmus Poll, 1971: 207, fig. 98; pls. 3 (fig. 15), 10 (fig. 5). Type locality: Ampem, Volta Lake
(Ghana). Holotype: MRAC 168500.
Distribution: Volta Lake, Ghana (Gosse, 1986b).
Synodontis macrops Greenwood, 1963
Synodontis macrops Greenwood, 1963: 66. Type locality: tributary stream of the Aswa River, Teso district, Uganda.

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Holotype: BMNH 1961.12.1.319; holotype illustrated in Poll (1971: fig. 88).
Distribution: Aswa River basin; Nile River system, Uganda (Gosse, 1986b).
Synodontis macrostigma Boulenger, 1911
Synodontis macrostigma Boulenger, 1911a: 432, fig. 325. Type locality: Okovango River. Lectotype: BMNH
1910.5.31.37, designated by Poll (1971: 345). A previous lectotype designation may have been made by Gilchrist & Thompson (1913: 462) in the caption to fig. 109, in which the illustrated specimen is labeled as thetype, if the illustrated specimen can be identified.
Distribution: Kafue and Mashi Rivers, Zambezi River basin; Okovango and Cunene Rivers (Gosse, 1986b; Skelton,1993).
Synodontis macrostoma Skelton & White, 1990
Synodontis macrostoma Skelton & White, 1990: 281, fig. 3. Type locality: Upper Zambezi River at Katima Mulilo,
Caprivi, Namibia. Holotype: AMG P11687.
Distribution: Cunene, Okavango, upper Zambezi and Kafur river basins (Skelton, 1993).
Synodontis manni De Vos, 2001
Synodontis manni De Vos, 2001a: 42, figs. 2–3. Type locality: Kenya: Tana River at Baomo area near Baomo village, 1º55'S, 40º08'E. Holotype: NMK FW-600/1.
Distribution: Tana River, Kenya (De Vos, 2001a).
Synodontis marmorata Lönnberg, 1895
Synodontis marmoratus Lönnberg, 1895: 186. Type locality: Muddy rivulet at Bonge [Cameroon]. Lectotype:
ZMUU, 38.5 mm SL specimen, designated by, and illustrated in, Poll (1971: 93, fig. 34).
Synodontis marmoratus truncatus Holly, 1927a: 8. Type locality: Njongfluß, Kamerun. Treated by Gosse (1986) as
possible synonym of S. marmoratus.
Distribution: Sanaga and Nyong Rivers (Gosse, 1986b).
Synodontis matthesi Poll, 1971
Synodontis matthesi Poll, 1971: 462, fig. 216; pls. 7 (fig. 14), 12 (fig. 20). Type locality: Mtera, riv. Ruaha (Tanzania). Holotype: ZMA 109743.
Distribution: Rufigi River basin, Tanzania (Gosse, 1986b).
Synodontis melanoptera Boulenger, 1903
Synodontis melanopterus Boulenger, 1903d: 327, pl. 29 (fig. 1). Type locality: Oguta, Niger Delta. Lectotype:
BMNH 1902.10.25.3, designated and illustrated in Poll (1971: 114, fig. 46).
Distribution: Porto Novo, Benin; Niger, Ouémé and Ogun River basins (Gosse, 1986b).
Synodontis multimaculata Boulenger, 1902
Synodontis multimaculatus Boulenger, 1902d: 43, pl. 12 (fig. 1). Type locality: l’Ubangi à Banzyville. Holotype:
MRAC 1314; illustrated in Poll (1971: 306, fig. 140).
Distribution: Ubangi River, Congo River basin (Gosse, 1986b).
Synodontis multipunctata Boulenger, 1898
Synodontis multipunctatus Boulenger, 1898d: 497. Type locality: Lake Tanganyika. Holotype: BMNH 1898.9.9.76;
illustrated and described in more detail in Boulenger (1898f: 24, pl. 8), with locality as Sumba.
Distribution: Lake Tanganyika (Gosse, 1986b).
Synodontis nebulosa Peters, 1852
Synodontis nebulosus Peters, 1852: 682. Type locality: Tette [Zambezi River, Mozambique]. Holotype: ZMB 3120;
illustrated in Peters (1868c: 28, pl. 5, fig. 1).
Distribution: Lower and middle Zambezi River basin (Skelton, 1993).
Synodontis nigrita Valenciennes, 1840
Synodontis nigrita Valenciennes, in Cuvier & Valenciennes, 1840b: 265 (197 in Strasbourg deluxe edition), pl. 441.
Type locality: Sénégal. Holotype: MNHN 0000-3051; illustrated in Poll (1971: 59, fig. 14).
Synodontis ornatus Pappenheim, in Pappenheim & Boulenger, 1914: 251, pl. 5 (fig. 2). Type locality: Albert See.
Syntypes: ZMB 19097 (8).
Distribution: Nile, Chad, Niger, Senegal, Gambia, Casamance, Geba, Kolente and Volta River basins; coastal rivers
from Ghana to Nigeria (Gosse, 1986b).

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Synodontis nigriventris David, 1936
Synodontis nigriventris David, 1936b: 417. Type locality: Mongende; Ikengo; Basongo; Koteli; Buta; Budjala;
Léopoldville; Flandria [restricted to Buta by lectotype designation]. Lectotype: MRAC 30312, designated by,
and illustrated in, Poll (1971: 397, fig. 186).
Distribution: Central Congo River basin (Gosse, 1986b).
Synodontis nigromaculata Boulenger, 1905
Synodontis nigromaculatus Boulenger, 1905b: 645. Type locality: Lake Bangwelo. Holotype: BMNH
1905.11.10.10; holotype illustrated in Poll (1971: fig. 148).
Synodontis melanostictus Boulenger, 1906f: 553, pl. 34. Type locality: Lofu, Lake Tanganyika. Holotype: BMNH
1906.9.8.72.
Synodontis colyeri Boulenger, 1923: 438. Type locality: Mansa River, N. Rhodesia. Holotype: SAM 14529 [now at
AMG].
Distribution: Okavango and upper Zambezi River basins; Upper Congo River basin, Zambia; Lake Tanganyika andKasai River (Skelton, 1993).
Synodontis njassae Keilhack, 1908
Synodontis njassae Keilhack, 1908: 168.Type locality: Njassa-See. Lectotype: ZMB 18191, designated by, and illustrated in, Poll (1971: 337, fig. 154).
Distribution: Lake Malawi (Gosse, 1986b).
Synodontis notata Vaillant, 1893
Synodontis Maculatus Vaillant, 1892b: 2. Type locality: Riv. Oubangui à Bangui. Holotype: apparently MNHN
1892-0080. Nomen oblitum, see Remarks.
Synodontis notatus Vaillant, 1893a: 17. Type locality: Riv. Oubangui à Bangui. Holotype: apparently MNHN 18920080. Proposed as a replacement for Synodontis maculatus, Vaillant, 1892, considered by Vaillant to be effectively preoccupied by Synodontis maculosus Rüppell, 1829. Poll (1971: fig. 116) illustrated MNHN 1886-0436
and referred to it as the lectotype. Nomen protectum.
Synodontis notatus binotata Pellegrin, 1926: 205. Type locality: Bolobo, Congo belge. Syntypes (2): MRAC 49686,
MNHN 1926-0195. Originally as Synodontis notatus var. binotata.
Synodontis notatus ocellatus Poll, 1938: 404, fig. 9. Type locality: environs de Maka, riv. Londo [Katanga, bassin duCongo]. Holotype: MRAC 49686. Originally as Synodontis notatus var. ocellatus.
Distribution: Congo River basin (Gosse, 1986b).
Remarks: Synodontis notata Vaillant, 1893, has been in wide use since its proposal (see Gosse, 1986b, for list of
publications) and considered here to be a nomen protectum. The holotype for Synodontis maculatus Vaillant,
1892, and S. notatus Vaillant, 1893, is listed by Bertin & Estève (1950) and other sources as MNHN 1886-0436,
but Gosse (1986: 135) indicated that the holotype was actually MNHN 1892-0080, which was collected by
Dybowski from the Oubangui, as stated in Vaillant (1892).
Synodontis nummifer Boulenger, 1899
Synodontis nummifer Boulenger, 1899a: 47, pl. 24. Type locality: Léopoldville. Holotype: MRAC 107.
Distribution: Congo River basin (Gosse, 1986b).
Synodontis obesus Boulenger, 1898
Synodontis obesus Boulenger, 1898a: 415. Type locality: Gaboon and Opobo River, Old Calabar [restricted to
Opobo River by lectotype designation]. Lectotype: BMNH 1896.5.5.67, designated by and illustrated in Poll
(1971: 96, fig. 36).
Synodontis loppei Pellegrin, 1927: 365. Type locality: Edea [Cameroon]. Syntypes: MHNLR P.197 (1), MNHN
1927-0278 (1). Illustrated in Pellegrin (1928a: 5, unnumbered figure).
Distribution: Coastal rivers from Ghana to Gabon; basins of the Comoë, Pra, Volta, Momo, Cross, Wouri, Sanaga
Nyong, Kribi and Lobé Rivers (Gosse, 1986b).
Synodontis ocellifer Boulenger, 1900
Synodontis ocellifer Boulenger, 1900e: 514. Type locality: Kunchow Creek [River Gambia]. Lectotype: BMNH
1901.7.17.7; designated by, and illustrated in, Poll (1971: 101, fig. 38).
Distribution: Senegal, Gambia, Volta, Chad, and Niger basins (Gosse, 1986b).

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Synodontis omias Günther, 1864
Synodontis omias Günther, 1864: 213. Type locality: West Africa [Upper Niger River]. Holotype: BMNH
1863.12.9.3 (skeleton).
Distribution: Niger River basin (Gosse, 1986b).
Synodontis ornatipinnis Boulenger, 1899
Synodontis ornatipinnis Boulenger, 1899b: 111, pl. 43 (fig. 2). Type locality: Coquilhatville et Bikoro (Lac Tumba)[restricted to Coquilhatville by lectotype designation]. Lectotype: MRAC 968, designated by Poll (1971: 299).
Distribution: Congo River system (Gosse, 1986b).
Synodontis ornatissima Gosse, 1982
Synodontis ornatus Boulenger, 1920a: 29, fig. 14. Type locality: Poko, Congo Belge. Holotype: MRAC 7144. Preoccupied by Synodontis ornatus Pappenheim, 1914; replaced by Synodontis ornatissimus Gosse, 1982.
Synodontis ornatissimus Gosse, 1982: 48. Type locality: Poko, Congo Belge. Holotype: MRAC 7144. Replacement
for Synodontis ornatus Boulenger, 1920; preoccupied by Synodontis ornatus Pappenheim, 1914.
Distribution: Ubangi River and its tributaries, Congo River basin (Gosse, 1986b).
Synodontis pardalis Boulenger, 1908
Synodontis pardalis Boulenger, 1908a: 30. Type locality: a waterfall of the Libi River, near the Ja River (Congo System), South Cameroon. Lectotype: BMNH 1909.4.29.91, designated by, and illustrated in, Poll (1971: 334, fig.
152).
Distribution: Ja River, Congo River basin, Cameroon (Gosse, 1986b).
Synodontis petricola Matthes, 1959
Synodontis petricola Matthes, 1959b: 78. Type locality: lac Tanganyika. Holotype: MRAC 130357; holotype illustrated in Poll (1971: fig. 192).
Distribution: Lake Tanganyika (Gosse, 1986b).
Synodontis pleurops Boulenger, 1897
Synodontis pleurops Boulenger, 1897b: 423. Type locality: Stanley Falls, upper Congo. Holotype: BMNH
1897.9.30.13; holotype illustrated in Poll (1971: fig. 120).
Distribution: Congo River basin, except the Luapula system and the lower Congo River (Gosse, 1986b).
Synodontis polli Gosse, 1982
Synodontis eurystomus Matthes, 1959b: 77. Type locality: lacTanganyika, confiné au littoral rocheux du lac. Holotype: MRAC 130440. Holotype illustrated in Matthes (1962: pl. 1, fig. c) and Poll (1971: fig. 190). Preoccupied
by Synodontis eurystomus Pfeffer, 1889; replaced by Synodontis polli Gosse, 1982.
Synodontis polli Gosse, 1982: 48. Type locality: Luhanga, Lake Tanganyika. Holotype: MRAC 130440. Replacement for Synodontis eurystomus Matthes, 1959.
Distribution: Lake Tanganyika (Gosse, 1986b).
Synodontis polyodon Vaillant, 1895
Synodontis polyodon Vaillant, 1895a: 48. Type locality: Ogôoué. Holotype: MNHN 1886-0433; holotype illustrated
in Vaillant (1895b: pl. 9, fig. 1), and described in more detail in Vaillant (1896b: 127).
Distribution: Ogowe River basin (Gosse, 1986b).
Synodontis polystigma Boulenger, 1915
Synodontis polystigma Boulenger, 1915: 170. Type locality: Rivière Luapula, à Kasenga; lac Moero, à Lukonzolwa[restricted to lac Moëro by lectotype designation]. Lectotype: MRAC 14174, designated by, and illustrated in,Poll (1971: 356, fig. 164).
Distribution: Luapula-Moëro Rivers, Congo River basin (Gosse, 1986b).
Synodontis pulcher Poll, 1971
Synodontis pulcher Poll, 1971: 465, fig. 218; pls. 7 (fig. 13), 12 (fig. 21). Type locality: Stanley Pool. Holotype:
MRAC 168399.
Distribution: Stanley Pool, Congo River basin (Gosse, 1986b).
Synodontis punctifer Daget, 1965
Synodontis punctifer Daget, 1965: 472. Type locality: le Nzo à Guiglo, Côte d’Ivoire. Holotype: MNHN 1964-0265;
holotype illustrated in Poll (1971: fig. 90).

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Distribution: Nzo River, Sassandra River basin (Gosse, 1986b).
Synodontis punctulata Günther, 1889
Synodontis punctulatus Günther, 1889: 71, pl. 8 (fig. a). Type locality: River Ruva, in the Arusha country. Lectotype:
BMNH 1887.11.3.47; designated by, and illustrated in, Poll (1971: 233, fig. 108).
Distribution: Ruwa River, Tanzania; Uebi Shebeli, Somalia (Gosse, 1986b).
Remarks: Considered to be a possible synonym of Synodontis zanzibaricus Peters, 1868 (De Vos 2001a: 49).
Synodontis rebeli Holly, 1926
Synodontis rebeli Holly, 1926: 157. Illustrated in Holly (1927b: 215, figs. 5–6). Type locality: Bamfluß, Flußgebiet
des Sanaga in Kamerun. Lectotype: NMW 7796: 1; designated by, and illustrated in, Poll (1971: 155, fig. 70).
Distribution: Sanaga River basin, Cameroon (Gosse, 1986b).
Synodontis resupinata Boulenger, 1904
Synodontis resupinatus Boulenger, 1904c: 199, pl. 8. Type locality: Lokoja, Northern Nigeria. Holotype: BMNH
1904.1.20.52.
Distribution: Niger River basin (Gosse, 1986b).
Synodontis ricardoae Seegers, 1996
Synodontis ricardoae Seegers, 1996: 245, figs. 178–180. Type locality: Kazizi, some kilometers northeast of Muze,at the northwestern shore of Lake Rukwa, Rukwa region, western Tanzania (07º37'N, 31º36'E). Holotype:
MRAC 94-34-P-1093.
Distribution: Lake Rukwa basin (Seegers, 1996).
Synodontis robbianus Smith, 1875
Synodontis robbianus Smith, 1875: 92, fig. 1. Type locality: Ikorofiong, Old Calabar River. Syntype: BMNH
1874.5.23.3 (1); illustrated in Poll (1971: fig. 26) as holotype.
Distribution: Lower Niger River and Cross River (Gosse, 1986b).
Synodontis robertsi Poll, 1974
Synodontis robertsi Poll, 1974: 442, fig. 1. Type locality: Riv. Lukenie à Elombe, ferry landing, lat. 2º49'S, long.
18º14'E. Holotype: MRAC 189535.
Distribution: Lukenie River, Congo River basin; known only from type locality (Gosse, 1986b).
Synodontis ruandae Matthes, 1959
Synodontis ruandae Matthes, 1959a: 62. Type locality: Riv. Kagera à Rusumu, Ruanda. Holotype: MRAC 130345;
holotype illustrated in Poll (1971: 178, fig. 82).
Distribution: Kagera River basin, Nile-Victoria system (Gosse, 1986b).
Synodontis rufigiensis Bailey, 1968
Synodontis rufigiensis Bailey, 1968: 346, fig. 1. Type locality: Lake Lugongwe, at Utete on the lower Rufiji, Tanzania. Holotype: BMNH 1968.6.12.1; holotype illustrated in Poll (1971: 428, fig. 200).
Distribution: Rufigi River basin, Tanzania (Gosse, 1986b).
Synodontis rukwaensis Hilgendorf & Pappenheim, 1903
Synodontis zambezensis rukwaensis Hilgendorf & Pappenheim, 1903: 267. Type locality: Rukwa Sees. Lectotype:
ZMB 16311; designated by, and illustrated in, Seegers (1996: 249, fig. 182).
Synodontis maculipinna Norman, 1922: 687. Type locality: Mpanganye, Rufigi River, Tanganyika Territory. Lectotype: BMNH 1922.4.18.27, designated by, and illustrated in, Poll (1971: 376, fig. 174); lectotype also illustrated
in Seegers (1996: fig. 184).
Synodontis wamiensis Lohberger, 1930: 91, fig. 1. Type locality: aus dem Wamiflusse und Nebenflüssen [30 km.
north of Bahamojo, Tanzania]. Holotype: NMW 18866; illustrated in Seegers (1996: fig. 185).
Distribution: Lake Rukwa basin, Ugalla basin Malagarasi system, and coastal drainages of Tangazni south of thePangani River (Seegers, 1996).
Remarks: Synonymy based on Seegers (1996).
Synodontis schall (Bloch & Schneider, 1801)
Silurus schall Bloch & Schneider, 1801: 385. Type locality: Habitat Nilum. Holotype: ZMB 3110 (lost; Paepke,
1999: 99).
Pimelodus clarias Geoffroy Saint-Hilaire, 1809: pl. 13 (figs. 3–4). Type locality: Nil, Egypt. Holotype: Possibly

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FERRARISMNHN 0000-4203 (1) or MNHN 0000-4205 (1 of 2). Name available from plate, with illustrated specimen theholotype, if identifiable. Described as Synodontis clarias in Geoffroy Saint-Hilaire (1827: 299). Independenceof this name and Silurus clarias Linnaeus not clearly established, but treated as independent in literature (e. g.,
Poll, 1971).
Synodontis maculosus Rüppell, 1829: 10, pl. 3 (fig. 1). Type locality: die Märkte von Cairo. Lectotype: SMF 653,
designated by Poll (1971: 50).
Synodontis arabi Valenciennes, in Cuvier & Valenciennes, 1840b: 261 (194 in Strasbourg deluxe edition). Type
locality: Nil, Egypte. Syntypes: MNHN 0000-4203 (1), MNHN 0000-4205 (2). Unneeded new name for Silurus
schall Bloch & Schneider, 1801.
Synodontis smithii Günther, 1896b: 222, pl. 9. Type locality: [Not stated; introduction indicates specimen obtained“en route to Lake Rudolph”; BMNH register lists Lake Stephanie [sic, Lac Stefanie, Kenya] and Poll (1971)
listed Lake Rudolph, but these are in contradiction to statements in introduction]. Holotype: BMNH
1895.12.31.68.
Distribution: Nile basin, Abaia, Stephanie and Rudolf Lakes, Chad, Niger, Senegal, Uebi Guiba (Uebi Shebeli)basins and possibly Tana and Volta rivers (Gosse, 1986b).
Synodontis schoutedeni David, 1936
Synodontis schoutedeni David, 1936b: 416. Type locality: Kungungu; Koteli; Basongo; Mongende; Budjala;Bumba; Flandria [restricted to Kungungu by lectotype designation]. Lectotype: MRAC 38093; designated by,
and illustrated in, Poll (1971: 393, fig. 184).
Distribution: Central Congo River basin (Gosse, 1986b).
Synodontis serpentis Whitehead, 1962
Synodontis serpentis Whitehead, 1962b: 100, figs. 1a, 1b. Type locality: Athi River at Jilore, thirty miles from rivermouth, Kenya. Holotype: BMNH 1962.1.19.1.
Distribution: Athi and Tana River basins, Kenya (Gosse, 1986b).
Synodontis serrata Rüppell, 1829
Synodontis serratus Rüppell, 1829: 8, pl. 2 (fig. 1). Type locality: Cairo. Holotype: SMF 2649 (dry).
Synodontis humeratus Valenciennes, in Cuvier & Valenciennes, 1840b: 264 (196 in Strasbourg deluxe edition). Type
locality: Nil. No types known; based on a drawing.
Distribution: Nile basin (Gosse, 1986b).
Synodontis smiti Boulenger, 1902
Synodontis smiti Boulenger, 1902d: 45, pl. 12 (fig. 3). Type locality: l’Ubangi à Banzyville. Lectotype: MRAC
1266, designated in Poll (1971: 316).
Synodontis tenuis Nichols & Griscom, 1917: 717, fig. 25. Type locality: Stanleyville, Congo. Holotype: AMNH
6535.
Distribution: Ubangi and Lualaba Rivers, Congo River basin (Gosse, 1986b).
Synodontis soloni Boulenger, 1899
Synodontis soloni Boulenger, 1899b: 110, pl. 43 (fig. 1). Type locality: Congo. Lectotype: MRAC 155385, designated by Poll (1971: 296).
Distribution: Stanley Pool rapids and Libenge, Congo River basin (Gosse, 1986b).
Synodontis sorex Günther, 1864
Synodontis sorex Günther, 1864: 211. Type locality: Chartoum, upper Nile. Lectotype: BMNH 1862.6.17.69, designated by Poll (1971: 65).
Distribution: Nile, Chad, Niger and Volta basins (Gosse, 1986b).
Remarks: Lectotype register number incorrectly listed as BMNH 1862.1.17.69 in Poll (1971) and Eschmeyer et al.
(1998).
Synodontis steindachneri Boulenger, 1913
Synodontis steindachneri Boulenger, 1913: 69. Type locality: Nyong River, S. Cameroon. Holotype: BMNH
1913.10.29.16; holotype illustrated in Boulenger (1916: fig. 188) and Poll (1971: fig. 162).
Distribution: Nyong River, Cameroon (Gosse, 1986b).
Synodontis tanganaicae Borodin, 1936

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Synodontis serratus tanganaicae Borodin, 1936: 9. Type locality: Kasanga, Lake Tanganyika. Lectotype: MCZ
32538; designated by De Vos & Thys van den Audenaerde (1998: 148).
Synodontis lacustricolus Poll, 1953: 157, fig. 18a; pl. 6 (fig. 3). Type locality: Stat. 123, au large de Karéma [Lac
Tanganyika]. Holotype: IRSNB 197.
Distribution: Lake Tanganyika (De Vos & Thys van den Audenaerde, 1998: 153).
Remarks: Synonymy first proposed in De Vos & Thys van den Audenaerde (1998: 153), with valid name as Synodontis tanganyicae.
Synodontis tessmanni Pappenheim, 1911
Synodontis tessmanni Pappenheim, 1911: 523, fig. 5. Type locality: Akonangi, im Kje [Ntem basin]. Holotype (12.8cm SL): ZMB 18413; illustrated in Poll (1971: fig. 62), as lectotype, but a holotype was clearly stated (as “Typeder Art”) in description.
Synodontis hollyi ntemensis Pellegrin, 1929b: 367. Type locality: Nyabessan (Ntem), Cameroun. Holotype: MNHN
1929-0051. Originally as Synodontis Hollyi var. ntemensis nov. var.
Distribution: Ntem River, Cameroon (Gosse, 1986b).
Synodontis thamalakanensis Fowler, 1935
Synodontis thamalakanensis Fowler, 1935a: 274, fig. 12. Type locality: Thamalakane River, Maun, BechuanalandProtectorate. Holotype: ANSP 53245.
Distribution: Okavango and Upper Zambezi River basins (Skelton, 1993).
Synodontis thysi Poll, 1971
Synodontis thysi Poll, 1971: 210, fig. 100; pls. 3 (fig. 16), 10 (fig. 6). Type locality: Mange-Ferry, riv. Little Scarcies
ou Kaba (Sierra Leone). Holotype: MRAC 168314.
Distribution: Little Scarcies River, Sierra Leone; known only from type locality (Gosse, 1986b).
Synodontis tourei Daget, 1962
Synodontis tourei Daget, 1962: 113, fig. 38; pl. 11 (figs. 24–25). Type locality: Bafing à Ballay, Guinée. Lectotype:
MNHN 1959-0109; designated by Poll (1971: 185).
Distribution: Bafing River, Senegal River basin, Guinea (Gosse, 1986b).
Synodontis unicolor Boulenger, 1915
Synodontis unicolor Boulenger, 1915: 170. Type locality: Lac Moero et rivière Luapula, à Kasenga [restricted to
Kasenga by lectotype designation]. Lectotype: MRAC 14223; designated by, and illustrated in, Poll (1971: 359,
fig. 166).
Distribution: Luapula-Moero system, Congo River basin (Gosse, 1986b).
Synodontis vanderwaali Skelton & White, 1990
Synodontis vanderwaali Skelton & White, 1990: 284, fig. 5. Type locality: Okavango River, Namibia. Holotype:
AMG P5796.6.
Distribution: Cunene, Okavango and upper Zambezi basins (Skelton, 1993).
Synodontis velifer Norman, 1935
Synodontis velifer Norman, 1935a: 219, fig. 3. Type locality: Ejura, Ashanti Forest, Gold Coast. Holotype: BMNH
1934.8.31.146; holotype illustrated in Poll (1971: 164, fig. 74).
Distribution: Sassandra, Bandama and Volta River basins (Gosse, 1986b).
Synodontis vermiculata Daget, 1954
Synodontis vermiculatus Daget, 1954: 292, fig. 109. Type locality: Mopti, riv. Niger (Mali). Neotype: MNHN 19611129; designated by Poll (1971: 170).
Distribution: Niger River basin (Gosse, 1986b).
Synodontis victoriae Boulenger, 1906
Synodontis victoriae Boulenger, 1906b: 438. Type locality: Entebbe and Buganga, Lake Victoria [restricted to
Buganga by lectotype designation]. Lectotype: BMNH 1906.5.30.190; designated by, and illustrated in, Poll
(1971: 120, fig. 50).
Distribution: Victoria Lake basin, Kioga Lake and Victoria Nile, Malagarasi River (Gosse, 1986b).
Synodontis violacea Pellegrin, 1919
Synodontis violaceus Pellegrin, 1919b: 212. Type locality: Gribingui. Lectotype: MNHN 1919-0246, designated by,

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and illustrated in, Poll (1971: 149, fig. 66).
Distribution: Chad, Niger and Volta basins (Gosse, 1986b).
Synodontis voltae Roman, 1975
Synodontis voltae Roman, 1975: 45, unnumbered figure. Type locality: dans la Bougouriba, affluent de la Volta
Noire. Holotype: MHV V5641.
Distribution: Upper Volta River basin; known only from type locality (Gosse, 1986b).
Synodontis waterloti Daget, 1962
Synodontis waterloti Daget, 1962: 111, fig. 37; pl. 10 (fig. 23). Type locality: Friguiagbé, près Kindia, Guinée. Holotype: MNHN 1935-0223.
Distribution: Waanje and Taja Rivers, Sierra Leone; St. John River basin, Liberia; Cess River basin, Côte d’Ivoire;
Pra River basin, Ghana (Gosse, 1986b).
Synodontis woosnami Boulenger, 1911
Synodontis woosnami Boulenger, 1911a: 424, fig. 319. Type locality: Okovango [Botswana]. Holotype: BMNH
1910.5.31.36.
Distribution: Cunene, Okavango and upper Zambezi River basins (Skelton, 1993).
Synodontis xiphias Günther, 1864
Synodontis xiphias Günther, 1864: 215. Type locality: West Africa. Holotype: BMNH 1863.11.9.1 (dry); holotype
illustrated in Boulenger (1911a: fig. 339) and Poll (1971: fig. 22).
Synodontis labeo Günther, 1865a: 453. Type locality: West Africa, probably Niger R. Holotype: BMNH 1865.5.9.8(stuffed); illustrated in Boulenger (1911a: fig. 337).
Distribution: Niger River basin (Gosse, 1986b).
Remarks: Synodontis labeo Günther, 1865, is treated by Willoughby (1994: 28) as probably valid.
Synodontis zambezensis Peters, 1852
Synodontis zambezensis Peters, 1852: 682. Type locality: riv. Zambèze, Mozambique. Lectotype: ZMB 3119; designated by, and illustrated in, Poll (1971: 225, fig. 104). Illustrated in Peters (1868c: 31, pl. 5, figs. 2–3).
Synodontis zambesensis Günther, 1864: 214. Type locality: Zambezi. Lectotype: ZMB 3119, designated by, andillustrated in, Poll (1971: 225, fig. 104). Unjustified emendation of Synodontis zambezensis Peters, 1852.
Distribution: Pongolo, Lompopo, Pungwe and Zambezi River basins, Rukwa Lake, Luangwe River of Moero basin.
(Gosse, 1986b).
Synodontis zanzibarica Peters, 1868
Synodontis zanzibaricus Peters, 1868b: 600. Type locality: Wahrscheinlich von Mombas [Kenya]. Lectotype: ZMB
6846; designated by, and illustrated in, Poll (1971: 229, fig. 106).
Distribution: Mombassa, Kenya (Gosse, 1986b).
Species inquirendae, Synodontis
Synodontis vaillanti Boulenger, 1897b: 424. Type locality: Congo français (Bangui). Holotype: MNHN 1892-0275.
Treated by Gosse (1986:148) as doubtfully valid.
Synodontis Hollyi Pellegrin, 1929b: 365. Type locality: Ndjibula ..., Nanga Eboko (Sanaga), Cameroun. Syntypes:
MNHN 1929-0047 (1), MNHN 1929-0048 (1), MNHN 1929-0049 (1), MNHN 1929-0050 (1). Type series apparently mixed; with specimens representing both Synodontis obesus Boulenger, 1898, and Synodontis rebeli Holly,
1926.
Synodontis galinae Kochetov, 1998: 4, 2 figs. Type locality: White Nile. Type(s): Whereabouts unknown.

NEMATOGENYIIDAE Bleeker, 1862
Nematogenyini Bleeker, 1862 (in Bleeker, 1862–63): 16. Type genus: Nematogenys Girard, 1855.
Taxonomic summary: de Pinna (2003).
1 genus, 2 species; including 1 named fossil species.

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NEMATOGENYS Girard, 1855
Nematogenys Girard, 1855: 198. Type species: Trichomycterus inermis Guichenot, 1848. Type by monotypy. Gender: Feminine.
Phylogeny: Arratia & Chang (1975).
† Nematogenys cuivi Azpelicueta & Rubilar, 1998
† Nematogenys cuivi Azpelicueta & Rubilar, 1998: 476, figs. 2, 3. Type locality: Cura-Mallín Formation, Miocene,Northern slope of the Cerro Rucañanco (38º41'S, 71º11'W), about 1,250 m a.s.l., Lonquimay, Chile. Holotype:
CPUC Lonq (R)/7 (neurocranium with Weberian complex, branchial and hyoid arches, pectoral girdle andspine, some vertebrae, and ribs).
Distribution: Cura-Mallin Formation, south-central Chile (Azpelicueta & Rubilar, 1998).
Nematogenys inermis (Guichenot, 1848)
Trichomycterus inermis Guichenot, 1848: 312, pl. 9 (fig. 2). Type locality: Chile. No types known.
Nematogenys nigricans Philippi, 1866: 716. Type locality: Chile. No types known.
Nematogenys pallidus Philippi, 1866: 716. Type locality: Chile. No types known.
Distribution: Isolated localities in Concepción, Rancagua and Angol, Chile. Formerly widespread throughout muchof Central Chile (de Pinna, 2003).

PANGASIIDAE Bleeker, 1858
Pangasini Bleeker, 1858b: 49, 63. Type genus: Pangasius Valenciennes, 1840.
Pangasianodonidi Fowler, 1958: 14. Type genus: Pangasianodon Chevey, 1931.
Revision: Roberts & Vidthayanon (1991).
Review: Vidthayanon & Roongthongbaisuree (1993, Thailand).
Identification guide: Jayaram (1977d, South Asia).
Phylogeny: Pouyaud et al. (2000); Pouyaud et al. (2004).
Remarks: Several phylogenetic studies indicate that the species included in this family may constitute a natural group,
but the group is likely nested within the Schilbidae. As such, the Pangasiidae may not deserve continued recognition
as a family but is so recognized herein due to current usage. See Fumihito (1989) for morphological comparisonsamong some species.
5 genera, 30 species; including 1 named fossil genus and 2 fossil species.
† CETOPANGASIUS Roberts & Jumnongthai, 1999
† Cetopangasius Roberts & Jumnongthai, 1999: 177. Type species: †Cetopangasius chaetobranchus Roberts &Jumnongthai, 1999. Type by original designation. Gender: Masculine. Also spelled Cetopagansius in text. Cetopangasius treated here as the valid spelling.
† Cetopangasius chaetobranchus Roberts & Jumnongthai, 1999
† Cetopangasius chaetobranchus Roberts & Jumnongthai, 1999: 177, figs.16–21. Type locality: Thailand, Northcentral Phetchabun Province, Ban Nong Pla; Miocene. Holotype: Department of Natural Resources, Thailand(DNR) TF 5013 (entire skeleton).
Distribution: Thailand; Miocene (Roberts & Jumnongthai, 1999).
HELICOPHAGUS Bleeker, 1858
Helicophagus Bleeker, 1858b: 28, 45. Type species: Helicophagus typus Bleeker, 1858. Type by original designation. Gender: Masculine.
Helicophagus leptorhynchus Ng & Kottelat, 2000
Helicophagus leptorhynchus Ng & Kottelat, 2000c: 55, figs. 1, 2a. Type locality: Thailand Ubon Ratchathani Provinve, Mun River at Bung Wai, about 7 km W of Ubon Ratchathani (15º12'30"N, 104º47'30"E). Holotype:
USNM 288676.
Distribution: Mekong and Chao Phraya River basins (Kottelat, 2001b).

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Helicophagus typus Bleeker, 1858
Helicophagus typus Bleeker, 1858a: 46. Type locality: Palembang, in fluvimine Mussi. Holotype: BMNH
1863.12.4.118; illustrated in Bleeker (1862–63: pl. 79 [= Silur. pl. 31], fig. 2).
Distribution: Sumatra and southeastern Borneo (Roberts & Vidthayanon, 1991; Musikasinthorn et al., 1998).
Helicophagus waandersii Bleeker, 1858
Helicophagus Waandersii Bleeker, 1858b: 175. Type locality: Palembang, in fluviis [Sumatra]. Holotype (340 mm
TL): BMNH 1863.12.4.89; illustrated in Bleeker (1862–63: pl. 80 [= Silur. pl. 32]).
Distribution: Mekong and Chao Phraya Rivers, and Sumatra (Roberts & Vidthayanon, 1991).
PANGASIANODON Chevey, 1931
Pangasianodon Chevey, 1931: 538. Type species: Pangasianodon gigas Chevey, 1931. Type by monotypy. Gender:
Masculine.
Remarks: Treatment of this genus as valid follows Rainboth (1996).
Pangasianodon gigas Chevey, 1931
Pangasianodon gigas Chevey, 1931: 538, figs. 1–2, pl. 1. Type locality: Cambodge. No types known (cast of type in
Musée Economique de Phnom-Pehn).
Pangasius paucidens Fang & Chaux, in Chaux & Fang, 1949b: 344, fig. 6. Type locality: Cambodge. Holotype:
MNHN 1966-0730.
Distribution: Mekong River basin; stocked in various reservoirs in Thailand (Kottelat, 2001b).
Pangasianodon hypophthalmus (Sauvage, 1878)
Helicophagus hypophthalmus Sauvage, 1878: 235. Type locality: Laos. Lectotype: MNHN a-0745, designated byKottelat (1984b: 812).
Pangasius sutchi Fowler, 1937: 141, figs. 27–29. Type locality: Bangkok, Siam. Holotype: ANSP 67902.
Distribution: Mekong and Chao Phraya River basin (Kottelat, 2001b); in cultivation elsewhere in Asia, including
Viet Nam and Myanmar.
PANGASIUS Valenciennes, 1840
Pangasius Valenciennes, in Cuvier & Valenciennes, 1840b: 45 (34 of Strasbourg deluxe edition). Type species: Pangasius buchanani Valenciennes, 1840 (= Pimelodus pangasius Hamilton, 1822). Type by monotypy. Gender:
Masculine.
Pseudopangasius Bleeker, 1862c: 399. Type species: Pangasius polyuranodon Bleeker, 1852. Type by original designation. Gender: Masculine. Also in Bleeker (1862–63: 14, 75).
Neopangasius Popta, 1904: 180. Type species: Neopangasius nieuwenhuisii Popta, 1904. Type by monotypy. Gender: Masculine.
Sinopangasius Chang & Wu, 1965: 11, 13. Type species: Sinopangasius semicultratus Chang & Wu, 1965. Type byoriginal designation. Gender: Masculine.
Pangasius bocourti Sauvage, 1880
Pangasius (Pseudopangasius) Bocourti Sauvage, 1880b: 229. Type locality: Phnom-Penh [Cambodia]. Holotype:
MNHN 0000-9528.
Pangasius altifrons Durand, 1940: 23, pl. 5. Type locality: Tonlé-Sap [Cambodia]. Holotype: at Institut Océanographique Nhatrang.
Distribution: Mekong and Chao Phraya River basins (Roberts & Vidthayanon, 1991; Kottelat, 2001b).
Pangasius conchophilus Roberts & Vidthayanon, 1991
Pangasius conchophilus Roberts & Vidthayanon, 1991: 114, figs. 1b, 2j, 5. Type locality: Thabo, Nongkhai prov.
[Thailand]. Holotype: NIFI 2227.
Distribution: Mekong, Bangpakong, and Chao Phraya River basins (Roberts & Vidthayanon, 1991; Kottelat, 2001b).
Pangasius djambal Bleeker, 1846
Pangasius djambal Bleeker, 1846b: 290. Type locality: Batavia, in fluviis. Possible syntypes: BMNH 1863.12.4.81(1, 400+ mm TL), RMNH 6854 (1), RMNH 31192 (6), RMNH 8069 (1).
Pangasius bedado Roberts, 1999: 110, figs. 1–3. Type locality: Palembang, Sumatra. Holotype: MZB 2598.

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Distribution: Java and Borneo (Roberts & Vidthayanon, 1991), Sumatra (Roberts, 1999).
Remarks: Roberts & Vidthayanon (1991:116) designated a neotype (RMNH 6854) for Pangasius djambal althoughseveral specimens from Bleeker’s collections, some of which may be syntypes, were reported on therein. The“neotype” was incorrectly listed as a lectotype by Eschmeyer et al. (1998). Synonymy follows Gustiano et al.
(2004). Register number of syntype of Pangasius djambal incorrectly reported as BMNH 1863.12.11.81 inEschmeyer et al. (1998).
Pangasius elongatus Pouyaud, Gustiano & Teugels, 2002
Pangasius elongatus Pouyaud, Gustiano & Teugels, 2002: 248, fig. 5. Type locality: Vietnam: Lower Mekong River
Delta. Holotype: MZB 10890.
Distribution: Lower reaches of the Mekong, Chao Phraya, and Bangpakong Rivers (Pouyaud et al., 2002).
Pangasius humeralis Roberts, 1989
Pangasius humeralis Roberts, 1989a: 131, figs. 100, 101c. Type locality: Kapuas basin, fish market at Sintang,
Western Borneo (Kalimantan Barat, Indonesia). Holotype: MZB 3680.
Distribution: Kapuas River basin, western Borneo (Roberts, 1989a; Roberts & Vidthayanon, 1991).
† Pangasius indicus (Marck, 1876)† Brachyspondylus indicus Marck, 1876a: 412, pl. 24 (fig. 2). Type locality: Padang, Sumatra; Tertiary. Holotype: at
Royal Geol. Mus. Dresden (imperfect fish). Also in Marck (1876b), priority of publication not established.
† Pseudeutropius verbeekii Günther, 1876: 435, pl. 15 (fig. 2). Type locality: Highlands of Padang, Sumatra; Tertiary. Syntypes (2): BMNH 47527 (postcranial fish).
Distribution: Padang, Sumatra; Tertiary.
Remarks: Gunther (1876: 436) noted that his specimens were likely conspecific with that of Marck, but considered
Marck’s name inaccurately reflected the geographic distribution and was, therefore, inappropriate. Woodward(1901: 326) lists † Brachyspondylus indicus, as a junior synonym of † Pseudeutropius verbeekii, which was followed in the literature. Also see Sanders (1934) for summary of the taxonomic history of these names. Roberts& Jumnongthai (1999) treated this as a species of Pangasius, apparently following Sanders (1934: 22). Patterson (1993) expressed skepticism of the reported Eocene age of these fossils.
Pangasius kinabatanganensis Roberts & Vidthayanon, 1991
Pangasius kinabatanganensis Roberts & Vidthayanon, 1991: 123, figs. 2P, 10. Type locality: Kinabatangan River at
Deramakot [Borneo, Malaysia]. Holotype: FMNH 68042.
Distribution: Kinabatangan River basin, northeastern Borneo (Roberts & Vidthayanon, 1991).
Pangasius krempfi Fang & Chaux, 1949
Pangasius krempfi Fang & Chaux, in Chaux & Fang, 1949b: 343, fig. 5. Type locality: en mer .... à Bong-Lao [Vietnam]. Holotype: MNHN 1966-0729.
Sinopangasius semicultratus Chang & Wu, 1965: 11, 13, figs. 1–4. Type locality: Off Paihai, Kwangtung, China.
Holotype: ASIZB 56-1174.
Distribution: Mekong and Hue River basins, and along coast of South China Sea of Vietnam and Guandong, China(Kottelat, 2001b).
Pangasius kunyit Pouyaud, Teugels & Legendre, 1999
Pangasius kunyit Pouyaud, Teugels & Legendre, 1999: 251, fig. 2. Type locality: Sangasanga Village, 30 km southeast of the town of Samarinda, delta of Mahakam River, East Kalimantan, Indonesia. Holotype: MZB 10009.
Distribution: Mahakam River delta, Borneo, Indonesia (Pouyaud et al., 1999).
Pangasius larnaudii Bocourt, 1866
Pangasius Larnaudii Bocourt, 1866: 15, pl. 1 (figs. 2, 2a). Type locality: Ajuthia. Syntypes: MNHN 0000-1549 (1),
MNHN a-9423 (1).
Pangasius taeniura Fowler, 1935b: 98, fig. 19. Type locality: Bangkok, Siam. Holotype: ANSP 61753.
Pangasius burgini Fowler, 1937: 141, figs. 24–26. Type locality: Bangkok, Siam. Holotype: ANSP 67901.
Distribution: Mekong, Meklong and Chao Phraya River basins (Roberts & Vidthayanon, 1991; Kottelat, 2001b).
Pangasius lithostoma Roberts, 1989
Pangasius lithostoma Roberts, 1989a: 132, fig. 102. Type locality: Market at Sintang, Western Borneo (Kalimantan
Barat, Indonesia). Holotype: MZB 3678.

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Pangasius macronema Bleeker, 1851
Pangasius macronema Bleeker, 1851a: 11. Type locality: Banjermassing, in fluviis. Holotype (116 mm): possibly
NMV 45892 (1), RMNH 6855 (10), BMNH 1863.12.4.66 (1).
Pangasius siamensis Steindachner, 1878c: 393. Type locality: Menam-Fluss bei Bangkok. Syntypes: NMW 45469(1), NMW 76998 (1).
? Pangasius aequilabialis Fowler, 1937: 140, figs. 20–23. Type locality: Bangkok, Siam. Holotype: ANSP 67897.
Distribution: Mekong and Chao Phraya River basins, Java, and Borneo (Kottelat, 2001b).
Remarks: BMNH lists BMNH 1863.12.4.66 (1, 144 mm SL) as a type of Pangasius macronema but is too large tobe the holotype.
Pangasius mahakamensis Pouyaud, Gustiano & Teugels, 2002
Pangasius mahakamensis Pouyaud, Gustiano & Teugels, 2002: 246, fig. 4. Type locality: Indonesia, East Kalimantan Province, Mahakam River at Samarinda. Holotype: MZB 10886.
Distribution: Asia: Mahakam River of eastern Kalimantan, Indonesia (Pouyaud et al., 2002).
Pangasius mekongensis Gustiano, Teugels & Pouyaud, 2003
Pangasius mekongensis Gustiano, Teugels & Pouyaud, 2003: 370, fig. 7. Type locality: [Lower Mekong River, Vietnam]. Holotype: MZB 10847.
Distribution: Mekong River, Vietnam (Gustiano, Teugels & Pouyaud, 2003).
Pangasius myanmar Roberts & Vidthayanon, 1991
Pangasius myanmar Roberts & Vidthayanon, 1991: 131, fig. 17. Type locality: Rangoon, [Myanmar]. Holotype: SU
33787.
Distribution: Myanmar, known only from types (Roberts & Vidthayanon, 1991).
Pangasius nasutus (Bleeker, 1863)
Pseudopangasius nasutus Bleeker, 1863f: 72. Type locality: Bandjermassin, in fluviis. Holotype (270 mm TL): possibly BMNH 1863.12.4.113 (207 mm SL, ~ 255 mm TL). Also described as new in Bleeker (1863e).
Pangasius ponderosus Myers, in Herre & Myers, 1937: 67, pl. 6. Type locality: Chandra Dam, Perak, Malay Peninsula. Holotype: SU 14162.
Distribution: Sumatra, Borneo and Malay Peninsula (Roberts & Vidthayanon, 1991).
Pangasius nieuwenhuisii (Popta, 1904)
Neopangasius Nieuwenhuisii Popta, 1904: 180. Type locality: le Bo, Bornéo central. Holotype: RMNH 7546;
described in more detail, with illustration of holotype, in Popta (1906: 30, pl. 1, figs. 3a, 3b).
Distribution: Mahakam River basin, eastern Borneo (Roberts & Vidthayanon, 1991).
Pangasius pangasius (Hamilton, 1822)
Pimelodus pangasius Hamilton, 1822: 163, 376, pl. 33 (fig. 52). Type locality: Estuaries of Bengal. No types known.
Pachypterus luridus Swainson 1839: 306. No types known. Made available by reference to “Ham. p. 163, f. 62 [sic,52]” [= Hamilton (1822, 163, pl. 33, fig. 52)]. Unneeded new name for Pimelodus pangasius Hamilton, 1822.
Pangasius Buchanani Valenciennes, in Cuvier & Valenciennes, 1840b: 45 (34 of Strasbourg deluxe edition), pl. 425.
Type locality: Estuaries of Bengal. No types known. Unneeded new name for Pimelodus pangasius Hamilton,
1822, apparently to avoid tautonymy.
Pangasius pangasius godavarii David, 1962: 151, fig. 3g. Type locality: Godavary River, at Rajahmundry [India].
Syntypes: at Central Inland Fisheries Research Institute’s Tank Fisheries Unit, Bangalore, India.
Pangasius pangasius upiensis Srivastava, 1968: 97, fig. 60. Type locality: Bale-ka-Maidan, R. Rohini, Gorakhpur,
Uttar Pradesh, India. Holotype: at Zool. Mus. Gorakhpur Univ., India.
Distribution: Rivers and estuaries of Indian subcontinent (Roberts & Vidthayanon, 1991).
Remarks: Redescribed in Hora (1938b).
Pangasius polyuranodon Bleeker, 1852
Pangasius polyuranodon Bleeker, 1852b: 425. Type locality: Bandjermassing, in fluviis. Holotype (160 mm TL):
possibly RMNH 6855.
Pangasius juaro Bleeker, 1852d: 589. Type locality: Palembang, in fluviis. Holotype (336 mm TL): possibly BMNH
1863.12.4.79 (1) or RMNH 6855 (1 of larger specimens).

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Distribution: Rivers of Sumatra and northern, western and southern Borneo (Pouyaud et al., 2002).
Pangasius rheophilus Pouyaud & Teugels, 2000
Pangasius rheophilus Pouyaud & Teugels, 2000: 194, figs. 1–2. Type locality: Bahau River, tributary of upper
Kayan at Longpujungan, Bulungan Regency, Kalimantan Timur, Indonesia. Holotype: MZB 10010.
Distribution: Bahau River, tributary of upper Kayan at Longpujungan, Kalimantan Timur, Indonesia (Pouyaud &Teugels, 2000).
Pangasius sabahensis Gustiano,Teugels & Pouyaud, 2003
Pangasius sabahensis Gustiano,Teugels & Pouyaud, 2003: 372, fig. 8. Type locality: [Kinabatangan River basin,
North Borneo, Sabah State, Malaysia]. Holotype: MZB 10847.
Distribution: Kinabatangan River basin, North Borneo, Sabah State, Malaysia (Gustiano et al., 2003).
Pangasius sanitwongsei Smith, 1931
Pangasius sanitwongsei Smith, 1931: 29, figs. 13–14. Type locality: Menam Chao Phya at Koh Yai, Central Siam.
Holotype: Siam Department of Fisheries (apparently lost).
Pangasius beani Smith, 1931: 26. Type locality: Klong Ban Poh, off Lopburi, near Ayuthia, Central Siam. Holotype:
USNM 90308.
Distribution: Mekong and Chao Phraya River basins (Roberts & Vidthayanon, 1991; Kottelat, 2001b).
Remarks: Roberts & Vidthayanon (1991: 138) acting as first reviser, selected Pangasius sanitwongsei as valid.
Species inquirendae, Pangasius
Pangasius delicatissimus Bleeker, 1863 (in Bleeker, 1862–63): 73, footnote. Type locality: Krawang, Java. No types
known, based on a drawing.
Pangasius hoeksi Hardenberg, 1948: 412. Type locality: Kapuas River (W. Borneo). Holotype (275 mm): Whereabouts
unknown.
PSEUDOLAIS Vaillant, 1902
Pseudolais Vaillant, 1902: 51. Type species: Pseudolais tetranema Vaillant, 1902. Type by monotypy. Gender: Masculine.
Pteropangasius Fowler, 1937:142. Type species: Pangasius cultratus Smith, 1931. Type by original designation.
Gender: Masculine.
Pseudolais micronemus (Bleeker, 1847)
Pangasius micronemus Bleeker, 1847b: 8. Type locality: Java. Syntypes: Possibly BMNH 1863.12.4.82 (1, 250 mm
SL), RMNH 6856 (3). Also in Bleeker (1847: 166).
Pangasius rios Bleeker, 1851d: 205. Type locality: Bandjermassing, in fluviis. Holotype (115 mm TL): possibly
BMNH 1863.12.4.92 (~ 104 mm SL).
Pseudolais tetranema Vaillant, 1902: 52, fig. 3. Type locality: Tepoe, bords du Mahakam, Bornéo central. Holotype:
Possibly RMNH 7821.
Pangasius de Zwaani Weber & de Beaufort, 1912: 535, pl. 12 (fig. 3). Type locality: Taluk, Sumatra. Holotype:
ZMA 113011; illustration of holotype reproduced in Weber & de Beaufort (1913: fig. 103).
Pangasius tubbi Inger & Chin, 1959: 287, fig. 47. Type locality: Confluence of the Deramakot River with the Kinabatangan River, Kinabatangan District, North Borneo. Holotype: FMNH 68047.
Distribution: Mekong and Hue Rivers, Malay Peninsula, Sumatra, Java, and Borneo (Roberts & Vidthayanon,1991).
Pseudolais pleurotaenia (Sauvage, 1878)
Pangasius pleurotaenia Sauvage, 1878: 235. Type locality: Laos. Lectotype: MNHN 0000-9529 designated by, andillustrated in, Kottelat (1984b: 813, fig. 8a). One of two syntypes illustrated in Sauvage (1881:169, pl. 8, fig. 6).
Pangasius cultratus Smith, 1931: 25. Type locality: Tapi River near Bandon, peninsular Siam. Holotype: USNM
90306.
Pangasius fowleri Smith, 1931: 28. Type locality: Lopburi River at Lopburi, Central Siam. Holotype: USNM 90309.
Distribution: Mekong, Meklong, Tapi and Chao Phraya River basins (Roberts & Vidthayanon, 1991; Kottelat,

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2001b).

PIMELODIDAE
Pimelodini Bonaparte, 1838: 131. Type genus: Pimelodus La Cepède, 1803.
Pimelodinae Swainson, 1838: 331, 338. Type genus: Pimelodus La Cepède, 1803.
Sorubinae Swainson, 1838: 356. Type genus: Sorubium Swainson, 1838.
Hypophthalmini Bleeker, 1862 (in Bleeker, 1862–63): 15. Type genus: Hypophthalmus Spix & Agassiz, 1829.
Calophysinae Eigenmann, 1890: 12. Type genus: Calophysus Müller & Troschel, 1842.
Luciopimelodinae Driver, 1919: 451. Type genus: Luciopimelodus Eigenmann & Eigenmann, 1888.
Pinirampidae Fernández-Yépez, 1965: 12. Type genus: Pinirampus Bleeker, 1858.
Brachyplatystomatini Lundberg & Akama, 2005: 496. Type genus: Brachyplatystoma Bleeker, 1862.
Remarks: Priority of the publications by Bonaparte and Swainson has not yet been established. Thus, the authorship ofthe valid family name is not fixed. Callophysidae, based on the unjustified emendation Callophysus, has been usedon occasion.
Taxonomic summary: Lundberg & Littmann (2003).
Phylogeny: Nass (1991).
Review: Mago-Leccia et al. (1986, Venezuela); Castro (1986a, Colombian Amazon and Orinoco basins).
29 genera, 93 species; 4 named fossil species.
AGUARUNICHTHYS Stewart, 1986
Aguarunichthys Stewart, 1986a: 662. Type species: Aguarunichthys torosus Stewart, 1986. Type by original designation. Gender: Masculine.
Aguarunichthys inpai Zuanon, Rapp Py-Daniel & Jégu, 1993
Aguarunichthys inpai Zuanon, Rapp Py-Daniel & Jégu, 1993: 258, figs. 2b, 6. Type locality: Brazil: Amazonas
State: Solimões River drainage, north of the Marchantaria Island, at about 15 km above confluence with Rio
Negro drainage. Holotype: INPA 5398.
Distribution: Middle Amazon River basin, Brazil (Lundberg & Littmann, 2003).
Aguarunichthys tocantinsensis Zuanon, Rapp Py-Daniel & Jégu, 1993
Aguarunichthys tocantinsensis Zuanon, Rapp Py-Daniel & Jégu, 1993: 252, figs. 2a, 4. Type locality: Brazil: ParáState: Tocantins River drainage, rapids above Marabá. Holotype: INPA 5400.
Distribution: Tocantins River basin, Brazil (Lundberg & Littmann, 2003).
Aguarunichthys torosus Stewart, 1986
Aguarunichthys torosus Stewart, 1986a: 663, figs. 7–8. Type locality: Peru, Departamento Amazonas, Río Cenepa,
1.6 km west of Huampami and near Peruvian military camp Chavez Valdivia, about 210 m elev., Approx.
4º28'S, 78º10'W. Holotype: LACM 39651-1.
Distribution: Cenepa River basin, Amazon River drainage, Peru (Lundberg & Littmann, 2003).
BAGROPSIS Lütken, 1874
Bagropsis Lütken, 1874c: 32. Type species: Bagropsis reinhardti Lütken, 1874. Type by monotypy. Gender: Feminine.
Bagropsis reinhardti Lütken, 1874
Bagropsis Reinhardti Lütken, 1874a: 32. Type locality: in flumine Rio das Velhas. Syntypes: BMNH 1876.1.10.9(1), NMW 45905 (1), ZMUC P 29630 (1), ZMUC P 29631 (1), ZMUC P 29632 (1). Species illustrated anddescribed in Lütken (1875: 160 (and p. V of summary), pl. 1, fig. 2).
Distribution: Das Velhas River basin, São Francisco River drainage, Brazil (Lundberg & Littmann, 2003).
BERGIARIA Eigenmann & Norris, 1901
Bergiella Eigenmann & Norris, 1900: 355. Type species: Pimelodus westermanni Reinhardt, 1874. Type by original

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designation. Gender: Feminine. Preoccupied by Bergiella Baker, 1897 (Hymenoptera), replaced by Bergiaria
Eigenmann & Norris, 1901.
Bergiaria Eigenmann & Norris, 1901: 272. Type species: Pimelodus westermanni Reinhardt, 1874. Type by being a
replacement name. Gender: Feminine. Replacement for Bergiella Eigenmann & Norris, 1900.
Bergiaria platana (Steindachner, 1908)
Bergiella platana Steindachner, 1908d: 111. Type locality: La Plata. Holotype: at NMW.
Distribution: Paraná River basin, Argentina (Lundberg & Littmann, 2003).
Bergiaria westermanni (Reinhardt, 1874)
Pimelodus Westermanni Reinhardt, in Lütken, 1874a: 33. Type locality: in flumine Rio das Velhas. Syntypes:
ZMUC P 29638 (1), ZMUC P 29639 (1). Species illustrated and described in more detail in Lütken (1875: 167(and p. VI of summary), pls. 2 (fig. 4), pl. 3 (fig. 4a–b)).
Distribution: Das Velhas River basin in São Francisco River drainage, Brazil (Lundberg & Littmann, 2003).
BRACHYPLATYSTOMA Bleeker, 1862
Piramutana Bleeker, 1858b: 356. Type species: Bagrus piramuta Kner, 1857. Type by monotypy. Gender: Feminine.
Piratinga Bleeker, 1858b: 355. Type species: Bagrus reticulatus Kner, 1857. Type by subsequent designation by
Bleeker (1862–63): 11. Gender: Feminine.
Brachyplatystoma Bleeker, 1862 (in Bleeker, 1862–63): 10. Type species: Platystoma vaillanti Valenciennes, 1840.
Type by original designation. Gender: Neuter.
Malacobagrus Bleeker, 1862 (in Bleeker, 1862–63): 11. Type species: Pimelodes filamentosus Lichtenstein, 1819.
Type by original designation. Gender: Masculine.
Taenionema Eigenmann & Bean, 1907: 662. Type species: Taenionema steerei Eigenmann & Bean, 1907. Type byoriginal designation. Gender: Neuter. Preoccupied by Taenionema Banks, 1905 (Plecoptera), and by Taenionema Bolivar, 1906 (Orthoptera); replaced by Goslinia Myers, 1941.
Goslinia Myers, 1941: 88. Type species: Taenionema steerei Eigenmann & Bean, 1907. Type by being a replacement name. Gender: Feminine. Replacement for Taenionema Eigenmann & Bean, 1907, preoccupied by Taenionema Banks, 1905.
Ginesia Fernández-Yépez, 1951: [1]. Type species: Ginesia cunaguaro Fernández-Yépez, 1951. Type by original
designation. Gender: Feminine.
Merodontotus Britski, 1981: 109. Type species: Merodontotus tigrinus Britski, 1981. Type by original designation.
Gender: Masculine.
Review: Miranda Ribeiro (1918f, Brazil).
Remarks: Treatment of Brachyplatystoma as valid over older names follows Lundberg & Littmann (2003) and Lundberg & Akama (2005). Synonymy of Merodontodus and Goslinia follows Lundberg & Akama (2005).
Brachyplatystoma capapretum Lundberg & Akama, 2005
Brachyplatystoma capapretum Lundberg & Akama, 2005: 501, fig. 8. Type locality: Brazil, Amazonas State, Rio
Tefé sand beach, Lago Mucura, Supiã-Pucu, Tefé, 3º22'S, 64º43'W. Holotype: MZUSP 78481.
Distribution: Amazon River basin of Brazil and Peru (Lundberg & Akama, 2005).
Brachyplatystoma filamentosum (Lichtenstein, 1819)
Pimelodes filamentosus Lichtenstein, 1819: 60. Type locality: Brasilien. Holotype: ZMB 2973.
Platystoma affine Valenciennes, in Cuvier & Valenciennes, 1840b: 24 (18 in the Strasbourg deluxe edition). Type
locality: [not stated; apparently from Rio de Janeiro, Brazil]. Holotype: MNHN a-9360 (mounted).
Platystoma gigas Günther, 1872: 450. Type locality: River Huallaga, Upper Amazon [Peru]. Holotype: BMNH
1872.12.3.1 (stuffed).
Piratinga pirá-aïba Goeldi, 1898: 464, 477, pl. (fig. 4). Type locality: [Pará, Brazil]. Holotype: Whereabouts
unknown. Name available from caption on plate.
Brachyplatystoma goeldii Eigenmann & Bean, 1907: 661, fig. 1. Type locality: Brazil: Amazon River, between Paraand Manaos. Holotype: USNM 52561.
Distribution: Amazon and Orinoco River basins and major rivers of the Guianas and NE Brazil (Lundberg & Litt-

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mann, 2003).
Brachyplatystoma juruense (Boulenger, 1898)
Platystoma juruense Boulenger, 1898e: 421, pl. 39. Type locality: Rio Jurua, an affluent of the Amazons, Brazil.
Holotype: BMNH 1897.11.26.11.
Ginesia cunaguaro Fernández-Yépez, 1951: [2], unnumbered figure. Type locality: Río Apure, 3 km east of San
Fernando de Apure, Venezuela. Holotype: AFY 51189; current whereabouts unknown.
Distribution: Amazon and Orinoco River basins (Lundberg & Littmann, 2003).
Brachyplatystoma platynemum Boulenger, 1898
Brachyplatystoma platynema Boulenger, 1898b: 477. Type locality: Brazil, Pará. Holotype: BMNH 1898.10.11.20.
Taenionema steerei Eigenmann & Bean, 1907: 662, fig. 2. Type locality: Brazil, Amazon River, between Para andManaos. Holotype: USNM 52571.
Distribution: Amazon and Orinoco River basins (Lundberg & Littmann, 2003).
† Brachyplatystoma promagdalena Lundberg, 2005
† Brachyplatystoma promagdalena Lundberg, 2005: 599, figs. 1a, 2a, 3, 4a, 5a, 6a. Type locality: Miocene Villavieja Formation of the Honda Group in the southern end of the río Magdalena basin near the town of Villavieja,
Huila Department, Colombia at approx. 3º5'N, 75º13'W. Holotype: IGM 183062 (partial vertebral column,
Weberian complex).
Distribution: La Venta formation, central Colombia; middle Miocene (Lundberg, 2005).
Brachyplatystoma rousseauxii (Castelnau, 1855)
Bagrus rousseauxii Castelnau, 1855: 32, pl. 14 (fig. 1). Type locality: rivière des Amazones. Holotype: MNHN a9457 (dry).
Bagrus Goliath Kner, 1857: 379. Type locality: Salto Theotonio [Brazil]. Holotype: at NMW.
Brachyplatystoma paraense Steindachner, 1909a: 195. Type locality: Fischmarkt von Pará [Brazil]. Holotype:
NMW 47590. Illustrated and described in more detail in Steindachner (1915e: 65, fig. 4, pl. 13, fig. 5).
Distribution: Amazon and Orinoco River basins and other major rivers of South America east of the Andes mountains (Lundberg & Littmann, 2003).
Brachyplatystoma tigrinum (Britski, 1981)
Merodontotus tigrinus Britski, 1981: 110, figs. 1–2. Type locality: Brazil, Cachoeira do Teotônio, rio Madeira, Território de Rondônia. Holotype: MZUSP 14004.
Distribution: Amazon River basin (Lundberg & Littmann, 2003).
Brachyplatystoma vaillantii (Valenciennes, 1840)? Silurus Vaillantii Cuvier, 1816: 204. Type locality: Brazil. Based on “bagre primus” Marcgravius (1648: 173).
Originally as Sil. Nob. Vaillantii.
Platystoma Vaillantii Valenciennes, in Cuvier & Valenciennes, 1840b: 21 (16 of Strasbourg deluxe edition), pl. 423.
Type locality: de Cayenne et de Suriname. Syntypes: MNHN b-0158 (1), MNHN b-0159 (1). Based in part on“bagre primus” Marcgravius (1648: 173).
Bagrus reticulatus Kner, 1857: 376, pl. 1 (fig. 1). Type locality: Salto Theotonio, am Flusse Araguay, Forte do Rio
branco und Rio Madeira [Brazil]. Holotype: at NMW.
Bagrus piramuta Kner, 1857: 382. Type locality: Barra do Rio negro und Borba Rio Madeira [Brazil]. Syntypes: at
NMW.
Brachyplatystoma parnahybae Steindachner, 1908e: 126. Type locality: Rio Parnahyba. Holotype: at NMW.
Distribution: Amazon and Orinoco River basins and major rivers of the Guianas and NE Brazil (Lundberg & Littmann, 2003).
Remarks: Inclusion of Silurus Vaillanti in the synonymy is based on the citation of the same species from Marcgravius (1648) in the text of that species as well as Platystoma Vaillantii Valenciennes.
Species inquirenda, Brachyplatystoma
Bagrus punctulatus Castelnau, 1855: 33, pl. 14 (fig. 2). Type locality: de la rivière des Amazones [Brazil]. Holotype: at
MNHN, but current whereabouts unknown.

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CALOPHYSUS Müller & Troschel, 1842
Calophysus Müller & Troschel, in Müller, 1842b: 310. Type species: Pimelodes macropterus Lichtenstein, 1819.
Type by subsequent designation by Bleeker (1862–63: 12). Gender: Masculine. Also as new in Müller (1843:
318).
Callophysus Müller & Troschel, 1848: 629. Type species: Pimelodes macropterus Lichtenstein, 1819. Type by subsequent designation by Bleeker (1862–63: 12). Gender: Masculine. Unjustified emendation of Calophysus, anda spelling that was in wide use.
Pimeletropis Gill, 1859b: 196. Type species: Pimeletropis lateralis Gill, 1859. Type by monotypy. Gender: Feminine.
Pseudocallophysus Bleeker, 1862 (in Bleeker, 1862–63): 12. Type species: Pimelodus ctenodus Spix & Agassiz,
1829. Type by original designation. Gender: Masculine.
Calophysus macropterus (Lichtenstein, 1819)
Pimelodes macropterus Lichtenstein, 1819: 59. Type locality: Brasilien. Holotype: ZMB 3055.
Pimelodus ctenodus Spix & Agassiz, 1829: 21, pl. 8a. Type locality: aequitorialis fluviis [Brazil]. Holotype: Whereabouts unknown (Kottelat, 1988).
Pimeletropis lateralis Gill, 1859b: 196. Type locality: Amazon River. Holotype: at Lyceum Natural History, NY(whereabouts unknown).
Distribution: Amazon and Orinoco River basins (Lundberg & Littmann, 2003).
CHEIROCERUS Eigenmann, 1917
Cheirocerus Eigenmann, 1917a: 398. Type species: Cheirocerus eques Eigenmann, 1917. Type by original designation. Gender: Masculine.
Sovichthys Schultz, 1944c: 190. Type species: Sovichthys abuelo Schultz, 1944. Type by original designation. Gender: Masculine.
Revision: Stewart & Pavlik (1985).
Cheirocerus abuelo (Schultz, 1944)
Sovichthys abuelo Schultz, 1944c: 191, pl. 1 (fig. a). Type locality: Venezuela, Río de Los Pájaros, 3 km above Lago
Maracaibo, at depth of 15 ft. Holotype: USNM 121183.
Distribution: Lake Maracaibo basin (Lundberg & Littmann, 2003).
Cheirocerus eques Eigenmann, 1917
Cheirocerus eques Eigenmann, 1917a: 398, pl. 39. Type locality: Villa Bella [Bolivia]. Holotype: FMNH 58255.
Distribution: Amazon River basin (Lundberg & Littmann, 2003).
Cheirocerus goeldii (Steindachner, 1908)
Pimelodina goeldii Steindachner, 1908c: 83. Type locality: Rio Purus [Brazil]. Holotype: NMW 45503.
Pimelodus leptus Eigenmann & Pearson, in Eigenmann & Allen, 1942: 104, pl. 4 (fig. 3). Type locality: Río
Pachitea [Peru]. Holotype: CAS 47288.
Distribution: Purus River basin (Lundberg & Littmann, 2003).
DUOPALATINUS Eigenmann & Eigenmann, 1888
Duopalatinus Eigenmann & Eigenmann, 1888b: 136. Type species: Platystoma emarginatum Valenciennes, 1840.
Type by monotypy. Gender: Masculine.
Duopalatinus emarginatus (Valenciennes, 1840)
Platystoma emarginatum Valenciennes, in Cuvier & Valenciennes, 1840b: 25 (19 of Strasbourg deluxe edition).
Type locality: rivière de Saint-François. Holotype: MNHN a-9353 (mounted).
Distribution: São Francisco River basin, Brazil (Lundberg & Littmann, 2003).
Duopalatinus peruanus Eigenmann & Allen, 1942
Duopalatinus peruanus Eigenmann & Allen, 1942: 107, pl. 4 (fig. 4). Type locality: Rio Puinagua, mouth of Rio
Pacaya [Río Ucayali system, Peru]. Holotype: CAS 63630.
Distribution: Amazon and Orinoco River (Lundberg & Littmann, 2003).

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FERRARISEXALLODONTUS Lundberg, Mago-Leccia & Nass, 1991
Exallodontus Lundberg, Mago-Leccia & Nass, 1991: 843. Type species: Exallodontus aguanai Lundberg, MagoLeccia & Nass, 1991. Type by original designation. Gender: Masculine.
Exallodontus aguanai Lundberg, Mago-Leccia & Nass, 1991
Exallodontus aguanai Lundberg, Mago-Leccia & Nass, 1991: 847, figs. 2–10, 13. Type locality: Venezuela, Territorio Federal Delta Amacuro, Río Orinoco near Los Castillos, 159 n mi from sea buoy at the terminus of the shipnavigation channel in Boca Grande, 8º32'N, 62º23'W, 20–30 m. Holotype: MBUCV V-18930.
Distribution: Amazon and Orinoco River basins (Lundberg & Littmann, 2003).
HEMISORUBIM Bleeker, 1862
Hemisorubim Bleeker, 1862 (in Bleeker, 1862–63): 10. Type species: Platystoma platyrhynchos Valenciennes, 1840.
Type by original designation. Gender: Masculine.
Hemisorubim platyrhynchos (Valenciennes, 1840)
Platystoma platyrhynchos Valenciennes in Cuvier & Valenciennes, 1840b: 27 (20 of Strasbourg deluxe edition).
Type locality: [Not stated]. Holotype: MNHN 0000-1203.
Distribution: Amazon, Maroni, Orinoco, and Paraná River basins (Lundberg & Littmann, 2003).
HYPOPHTHALMUS Cuvier, 1829
Hypophthalmus Cuvier, 1829: 293. Type species: Hypophthalmus edentatus Spix & Agassiz, 1829. Type by subsequent designation by Bleeker (1862–63: 15). Gender: Masculine.
Hypophthalmus Spix & Agassiz, 1829: 16. Type species: Hypophthalmus edentatus Spix & Agassiz, 1829. Type by
subsequent designation Bleeker (1862–63: 15). Gender: Masculine.
Notophthalmus Hyrtl, 1859: 17. Type species: Hypophthalmus marginatus Valenciennes, 1840. Type by monotypy.
Gender: Masculine. Preoccupied by Notophthalmus Rafinesque, 1820 (Amphibia); apparently not replaced.
Pseudohypophthalmus Bleeker, 1862 (in Bleeker, 1862–63): 15. Type species: Hypophthalmus fimbriatus Kner,
1857. Type by original designation. Gender: Masculine.
Review: Lopez-Fernandez & Winemiller (2000, Venezuela).
Phylogeny: Howes (1983a), Lundberg et al. (1991).
Hypophthalmus edentatus Spix & Agassiz, 1829
Hypophthalmus edentatus Spix & Agassiz, 1829: 16, pl. 9. Type locality: influviis Brasiliae equitorialis. Syntypes(2): Possibly MHNN 706 (2); see Kottelat (1988: 24) for discussion.
Hypophthalmus Spixii Valenciennes in Cuvier & Valenciennes, 1840b: 231 (172 of Strasbourg deluxe edition). Type
locality: la partie la plus septentrionale du Brésil. Possible syntypes: MHNN 706 (2). Unneeded new name for
Hypophthalmus edentatus Spix & Agassiz, 1829.
Distribution: Amazon and Orinoco River basins and Atlantic coastal rivers of Guyana and Suriname (Lundberg &Littmann, 2003).
Hypophthalmus fimbriatus Kner, 1857
Hypophthalmus fimbriatus Kner, 1857: 444, pl. 9 (fig. 30). Type locality: Rio negro [Brazil]. Syntype: NMW 50519(1).
Distribution: Amazon River at Santarém and Negro River basin in Brazil and Venezuela (Lundberg & Littmann,
2003).
Hypophthalmus marginatus Valenciennes, 1840
Hypophthalmus marginatus Valenciennes, in Cuvier & Valenciennes, 1840b: 225 (168 of Strasbourg deluxe edition),pl. 439. Type locality: Cayenne, ... Surinam. Syntypes: MNHN a-8961 (1), MNHN a-8963 (1), RMNH 1932(1), RMNH 1933 (1).
Distribution: Amazon and Orinoco River basins and major rivers of French Guiana and Suriname (Lundberg & Littmann, 2003).
Hypophthalmus oremaculatus Nani & Fuster, 1947
Hypophthalmus oremaculatus Nani & Fuster, 1947: 3, figs. 2–3. Type locality: Puerto Gaboto, Rio Paraná, curso

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inferior [Argentina]. Holotype: MACN 3496.
Distribution: Paraná River basin, Brazil and Argentina (Lundberg & Littmann, 2003).
Species inquirendae, Hypophthalmus
Hypophthalmus longifilis Valenciennes in Cuvier & Valenciennes, 1840b: 230 (171 of Strasbourg deluxe edition). Type
locality: Surinam. Syntypes: RMNH 1931 (1), RMNH 2974 (1), RMNH 2988 (1).
Hypophthalmus perporosus Cope, 1878: 673. Type locality: Peru, probably Nauta. Holotype: USNM 132589.
Hypophthalmus devall Röhl, 1942: 383. Type locality: Venezuela, rios Aupre y Orinoco. Type(s): Whereabouts
unknown.
IHERINGICHTHYS Eigenmann & Norris, 1900
Iheringichthys Eigenmann & Norris, 1900: 354. Type species: Pimelodus labrosus Lütken, 1874. Type by original
designation. Gender: Masculine.
Iheringichthys labrosus (Lütken, 1874)
Pimelodus labrosus Lütken, 1874b: 200. Type locality: La Plata. Syntypes: ZMUC P 29633 (1), ZMUC P 29634 (1),
ZMUC P 29635 (1), ZMUC P 29636 (1)..
Distribution: Paraná River basin (Lundberg & Littmann, 2003).
Iheringichthys megalops Eigenmann & Ward, 1907
Iheringichthys megalops Eigenmann & Ward, in Eigenmann, McAtee & Ward, 1907: 115, pl. 32 (figs. 3–4). Type
locality: Paraguay, Bahia Negra, Río Paraguay. Holotype: CAS 63631.
Distribution: Paraná River basin, Paraguay (Lundberg & Littmann, 2003).
LEIARIUS Bleeker, 1862
Leiarius Bleeker, 1862 (in Bleeker, 1862–63): 10. Type species: Arius longibarbis Castelnau, 1855. Type by original
designation. Gender: Masculine.
Sciadeoides Eigenmann & Eigenmann, 1888b: 136. Type species: Sciades marmoratus Gill, 1870. Type by monotypy. Gender: Masculine. Originally proposed as a subgenus of Sciades.
Leiarius marmoratus (Gill, 1870)
Sciades marmoratus Gill, 1870: 95. Type locality: Peru or Ecuador, Amazon system, Río Marañon or Río Napo.
Holotype: USNM 8447.
Distribution: Amazon, Essequibo, and Orinoco River basins (Lundberg & Littmann, 2003).
Leiarius pictus (Müller & Troschel, 1849)
Bagrus (Sciades) pictus Müller & Troschel, 1849: 8, pl. 1 (fig. 1). Type locality: unbekannt. Holotype: ZMB 2991.
Distribution: Amazon and Orinoco River basins (Lundberg & Littmann, 2003).
Species inquirendae, Leiarius
Pimelodus arekaima Jardine, in Schomburgk, 1841: 178, pl. 5. Type locality: Upper Essequibo, Rio Branco. No types
known.
Arius longibarbis Castelnau, 1855: 36, pl. 15 (fig. 2). Type locality: rivière des Amazones. Holotype: MNHN 0000-1189.
Originally as Arius ? longibarbis.
Pimelodus multiradiatus Kner, 1857: 414. Type locality: Borba am Rio Madeira und Forte do Rio branco am Rio Facutu[Brazil]. Syntypes: NMW 16515 (1), NMW 16518 (1).
LUCIOPIMELODUS Eigenmann & Eigenmann, 1888
Luciopimelodus Eigenmann & Eigenmann, 1888b: 122. Type species: Pimelodus pati Valenciennes, 1835. Type byoriginal designation. Gender: Masculine.
Luciopimelodus pati (Valenciennes, 1835)
Pimelodus Pati Valenciennes, 1835, in Valenciennes, 1835–47: pl. 1 (figs. 7–9). Type locality: Not stated [Corri-

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FERRARIS

entes, Argentina]. Probable holotype: MNHN a-9419 (1, dry). Name available from plate; mentioned in Valenciennes (1847: 7) from Corrientes. Described in Cuvier & Valenciennes (1840b: 176 (131 of Strasbourg deluxe
edition)).
Silurus 11-radiatus Larrañaga, 1923: 386. Type locality: Uruguay. No types known. Corresponds to Silurus pati on
p. 376.
Distribution: La Plata and Blanco River basins (Lundberg & Littmann, 2003).
MEGALONEMA Eigenmann, 1912
Megalonema Eigenmann, 1912b: 150. Type species: Megalonema platycephalum Eigenmann, 1912. Type by original designation. Gender: Neuter.
Megalonema argentina (MacDonagh, 1938)
Perugia argentina MacDonagh, 1938: 157. Type locality: Argentina, rio Paraná, Posadas, Territorio de Misiones.
Holotype: MLP 5.V.I.35.
Distribution: Paraná River basin, Argentina (Lundberg & Littmann, 2003).
Megalonema pauciradiatum Eigenmann, 1919
Megalonema pauciradiatum Eigenmann, in Driver, 1919: 455. Type locality: Paraguay, Villa Rica. Holotype: CAS
63672.
Distribution: Paraná River basin, Paraguay (Lundberg & Littmann, 2003).
Megalonema platanum (Günther, 1880)
Pimelodus platanus Günther, 1880a: 10. Type locality: Parana. Holotype: BMNH 1872.6.8.18.
Distribution: Paraná River basin (Lundberg & Littmann, 2003).
Megalonema platycephalum Eigenmann, 1912
Megalonema platycephalum Eigenmann, 1912b: 150, fig. 31, pl. 10 (fig. 2). Type locality: Tumatumari, British Guiana. Holotype: FMNH 53224.
Distribution: Amazon, Essequibo, and Orinoco River basins (Lundberg & Littmann, 2003).
Megalonema psammium Schultz, 1944
Megalonema platycephalum psammium Schultz, 1944c: 216, pl. 2 (fig. b). Type locality: Venezuela, Rio Palmar at
the bridge 70 km southwest of Maracaibo. Holotype: USNM 121175.
Distribution: Lake Maracaibo basin, Colombia, Venezuela (Lundberg & Littmann, 2003).
Megalonema xanthum Eigenmann, 1912
Megalonema xanthum Eigenmann, 1912a: 16. Type locality: Girardot [Colombia]. Holotype: FMNH 56032; holotype illustrated in Eigenmann (1922b: 35, pl. 3, fig. 3) as Perugia xanthus.
Distribution: Magdalena River basin, Colombia (Lundberg & Littmann, 2003; Maldonado-Ocampo et al., 2005).
PARAPIMELODUS La Monte, 1933
Parapimelodus La Monte, 1933b: 226. Type species: Pimelodus valenciennis Lütken, 1874. Type by original designation. Gender: Masculine.
Revision: Lucena et al. (1992).
Parapimelodus nigribarbis (Boulenger, 1889)
Pimelodus (Pseudorhamdia) nigribarbis Boulenger, 1889: 266. Type locality: Brazil, Rio Grande do Sul,
Camaquam River. Lectotype: BMNH 1889.8.24.6, designated by Lucena et al. (1992: 145).
Distribution: Laguna dos Patos basin, Brazil (Lundberg & Littmann, 2003).
Parapimelodus valenciennis (Lütken, 1874)
Pimelodus valenciennis Lütken, 1874b: 200. Type locality: La Plata. Holotype: ZMUC P 29637.
Pimelodus Spegazzinii Perugia, 1891: 632. Type locality: Rio Durazno. Syntypes: MSNG 8026 (2).
Distribution: Paraná River basin (Lundberg & Littmann, 2003).
PERRUNICHTHYS Schultz, 1944
Perrunichthys Schultz, 1944c: 229. Type species: Perrunichthys perruno Schultz, 1944. Type by original designation. Gender: Masculine.

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Perrunichthys perruno Schultz, 1944
Perrunichthys perruno Schultz, 1944c: 230, pl. 3 (fig. b); fig. 3. Type locality: Venezuela, Río Negro, below themouth of the Río Yasa, about 75 km. south of Rosario, west side of Lago Maracaibo. Holotype: USNM 121189.
Distribution: Lake Maracaibo basin (Lundberg & Littmann, 2003).
PHRACTOCEPHALUS Spix & Agassiz, 1829
Phractocephalus Spix & Agassiz, 1829: 10. Type species: Phractocephalus bicolor Spix & Agassiz, 1829. Type bymonotypy. Gender: Masculine.
Pirarara Spix & Agassiz, 1829: 23. Type species: Silurus hemioliopterus Bloch & Schneider, 1801. Type by monotypy. Gender: Feminine. Appeared first as name in synonymy under Phractocephalus and in legend for pl. 6;
made available by Bleeker (1862–63: 11) but with authorship to Spix & Agassiz.
Remarks: Phractocephalus selected as valid by first reviser action of Agassiz (in Spix & Agassiz, 1831: Conspectus); see Kottelat (1989b: 321) for details.
Phractocephalus hemioliopterus (Bloch & Schneider, 1801)
Silurus hemioliopterus Bloch & Schneider, 1801: 385. Type locality: [in flumine Maranham Brasiliae].
Pimelodus grunniens Humboldt, in Humboldt & Valenciennes, 1821: 172. Type locality: le Bas-Orénoque. No types
known.
Phractocephalus bicolor Spix & Agassiz, 1829: 23, pl. 6. Type locality: in fluvio Amazonum [Brazil]. Whereabouts
unknown (Kottelat, 1988). As Pirarara bicolor on plate.
Distribution: Amazon and Orinoco River basins (Lundberg & Littmann, 2003).
Remarks: See Boeseman (1983b) for comments on taxonomy and nomenclature.
† Phractocephalus nassi Lundberg & Aguilera, 2003
† Phractocephalus nassi Lundberg & Aguilera, 2003: 101, figs. 3, 5a, 5b, 5e, 5h, 5k, 5l. Type locality: El Mamón oil
field, 350 m north of oil well number 1, locality USB 56-FU, 11º15'N, 70º13'W, Urumaco Formation, middle
member, Falcón State, northwestern Venezuela. Holotype: MCN.USB OL-2142.
Distribution: Urumaco Formation, Falcón State, Venezuela, upper Miocene (Lundberg & Aguilera, 2003).
Remarks: Described earlier by Lundberg et al. (1988) as Phractocephalus hemioliopterus.
PIMELODINA Steindachner, 1877
Pimelodina Steindachner, 1877a: 149. Type species: Pimelodina flavipinnis Steindachner, 1877. Type by monotypy.
Gender: Feminine.
Revision: Stewart (1986a).
Pimelodina flavipinnis Steindachner, 1877
Pimelodina flavipinnis Steindachner, 1877a: 150, pl. 13 (fig. 2) [as Pimelodus (Pimelodina) flavipinnis]. Type locality: Amazonenstrom bei Para [Brazil]. Holotype: NMW 45498.
Pimelodina nasus Eigenmann & Eigenmann, 1888b: 120. Type locality: Para [Brazil]. Holotype: MCZ 7490.
Distribution: Amazon and Orinoco River basins (Lundberg & Littmann, 2003).
PIMELODUS La Cepède, 1803
Pimelodus La Cepède, 1803: 93. Type species: Pimelodus maculatus La Cepède, 1803. Type by subsequent designation, by Gill (1861b). Gender: Masculine.
Pseudariodes Bleeker, 1862 (in Bleeker, 1862–63): 11. Type species: “Pseudariodes clarias = Silurus clarias Bl.”[= Pimelodus Blochii Valenciennes, 1840, nec. Silurus clarias Linnaeus, 1758]. Type by original designation.
Gender: Masculine.
Pseudorhamdia Bleeker, 1862 (in Bleeker, 1862–63): 11. Type species: Pimelodus maculatus La Cepède, 1803.
Type by original designation. Gender: Feminine.
Pimelodus absconditus Azpelicueta, 1995
Pimelodus absconditus Azpelicueta, 1995: 72, figs. 1–2. Type locality: Argentina, Misiones, San Javier, UruguayRiver. Holotype: MLP 8781.
Distribution: Paraná River basin (Lundberg & Littmann, 2003).

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FERRARISPimelodus albicans (Valenciennes, 1840)
Arius albidus Valenciennes, 1835, in Valenciennes, 1835–47: pl. 3, fig. 2. Type locality: Not stated. Holotype:
MNHN a-9400. Name made available by caption on plate.
Arius albicans Valenciennes, in Cuvier & Valenciennes, 1840b: 80 (60 of Strasbourg deluxe edition). Type locality:
Buénos-Ayres. Syntype: MNHN a-9400 (dry). Unneeded new name or lapsus calami for Arius albidus Valenciennes.
Silurus muticus Larrañaga, 1923: 386. Type locality: Uruguay. No types known.
Distribution: Paraná River basin, Argentina (Lundberg & Littmann, 2003).
Remarks: Treatment of Pimelodus albicans as valid follows Lundberg & Littmann (2003), in which Arius albidus
Valenciennes, 1835, was treated as a nomen oblitum.
Pimelodus albofasciatus Mees, 1974
Pimelodus albofasciatus Mees, 1974: 137, pl. 4. Type locality: Suriname, Sipaliwini. Holotype: RMNH 26156.
Distribution: Amazon, Orinoco, upper Corantijn and Sipaliwini River basins (Lundberg & Littmann, 2003).
Pimelodus altissimus Eigenmann & Pearson, 1942
Pimelodus altissimus Eigenmann & Pearson, in Eigenmann & Allen, 1942: 106, pl. 5 (fig. 5). Type locality: Rio
Ucayali, near Orellana [Peru]. Holotype: CAS 55369.
Distribution: Amazon River basin (Lundberg & Littmann, 2003).
Pimelodus atrobrunneus Vidal & Lucena, 1999
Pimelodus atrobrunneus Vidal & Lucena, 1999: 123, figs. 1–3. Type locality: Brasil: rio Ligeiro na estrada entre
Marcelino Ramos e Maximiliano de Almeida, Marcelino Ramos, 27º38'S–51º52'O, Rio Grande do Sul. Holotype: MCP 19678.
Distribution: Upper Uruguay River basin, Brazil (Lundberg & Littmann, 2003).
Pimelodus blochii Valenciennes, 1840
Pimelodus Blochii Valenciennes, in Cuvier & Valenciennes, 1840b: 188 (139 of Strasbourg deluxe edition). Type
locality: Surinam. Syntypes: MNHN b-0072 (1), ZMB 2995 (1). Based in part on Silurus clarias of Bloch(1782: pl. 35, figs. 1–2), not Silurus clarias Linnaeus, 1758.
Pseudorhamdia macronema Bleeker, 1864a: 79, pl. 13 (fig. 7), pl. 14. Type locality: Surinama. Lectotype: RMNH
3069, designated by Boeseman (1972: 317).
Pseudorhamdia piscatrix Cope, 1870b: 569. Type locality: Pebas [Peru]. Syntypes: ANSP 8386–87 (2).
Distribution: Gulf of Paria, Amazon, Corantijn, Essequibo and Orinoco River basins (Lundberg & Littmann, 2003);
Magdalena River basin (Maldonado-Ocampo et al., 2005).
Remarks: Eschmeyer et al. (1998) and Lundberg & Littmann (2003) list MNHN b-0072 as lectotype of Pimelodus
blochii, without citing a published source of the lectotype designation and, instead, apparently relying on a notation in the MNHN catalog to that effect. No published designation has been found, so the two specimens aretentatively retained as syntypes here.
Pimelodus brevis Marini, Nichols & La Monte, 1933
Pimelodus brevis Marini, Nichols & La Monte, 1933: 1, fig. 1. Type locality: Argentina, Rio de la Plata, San
Fernando. Holotype: AMNH 12240 (missing).
Distribution: Paraná River basin; Durazno River, Argentina (Lundberg & Littmann, 2003).
Pimelodus coprophagus Schultz, 1944
Pimelodus clarias coprophagus Schultz, 1944c: 203, fig. 2. Type locality: Venezuela, Río Agua Caliente, 2 to 3 km
above the southwestern corner of Lago Maracaibo. Holotype: USNM 121150.
Distribution: Lake Maracaibo basin (Lundberg & Littmann, 2003).
Pimelodus fur (Reinhardt, 1874)
Pseudorhamdia fur Reinhardt, in Lütken, 1874c: 33. Type locality: in flumine Rio das Velhas. Syntypes: NMW
44443 (3), NMW 44763 (2), SMNS 2026 (1), ZMB 9181 (2), ZMUC P 29643–ZMUC P 29651 (1 each). Illustrated and described in more detail in Lütken (1875: 169 (and p. VI of summary), pl. 2, fig. 3; pl. 3, fig. 3a).
Distribution: Das Velhas River basin, São Francisco River drainage, Brazil (Lundberg & Littmann, 2003).
Pimelodus garciabarrigai Dahl, 1961
Pimelodus garcia-barrigai Dahl, 1961: 494. Type locality: Colombia, Deep pool in Caño Lozada, about 11 km

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above its junction with the Guayabero River. Holotype: ICNMHN 744 (Cala, 1981).
Distribution: Guayabero River basin, Orinoco River drainage, Colombia (Lundberg & Littmann, 2003).
Pimelodus grosskopfii Steindachner, 1879
Pimelodus (Pimelodus) Grosskopfii Steindachner, 1879c: 194. Type locality: Río Cauca [Colombia]. Syntypes:
NMW 45781 (4), NMW 45782 (1). Also described as new in Steindachner (1879g: 186); described in moredetail and illustrated in Steindachner (1880b: 57, pl. 1 figs. 1, 1a).
Pimelodus longifilis Posada, 1909: 294. Type locality: aus dem Cauca [Colombia]. No types known.
Distribution: Magdalena River basin and Lake Maracaibo basins (Lundberg & Littmann, 2003).
Pimelodus heraldoi Azpelicueta, 2001
Pimelodus heraldoi Azpelicueta 2001: 194, figs. 1–2 Type locality: Brazil, Estado de São Paulo: Município de Pirassununga, rio Mogi Guaçu in Emas. Holotype: MZUSP 22713.
Distribution: Upper Paraná River basin, Brazil (Lundberg & Littmann, 2003).
Pimelodus jivaro Eigenmann & Pearson, 1942
Pimelodus jivaro Eigenmann & Pearson, in Eigenmann & Allen, 1942: 105, pl. 4 (fig. 2). Type locality: Rio Morona[Peru]. Syntypes (5; 85–105 mm): CAS 55891 (3).
Distribution: Upper Amazon River basin, Ecuador, Peru (Lundberg & Littmann, 2003).
Pimelodus maculatus La Cepède, 1803
Pimelodus maculatus La Cepède, 1803: 94. Type locality: Le grand fleuve de la Plata, ... Buénos-Ayres, ainsi qu’à laEncenada. No types known.
Distribution: Paraná and São Francisco River basins (Lundberg & Littmann, 2003).
Pimelodus microstoma Steindachner, 1877
Pimelodus microstoma Steindachner, 1877b: 604, footnote. Type locality: Brazil, von Irisanga, Rio branco und Barrado Rio negro. Syntypes: NMW 45823 (1), NMW 45824 (2).
Distribution: Amazon River basin, Brazil (Lundberg & Littmann, 2003).
Pimelodus mysteriosus Azpelicueta, 1998
Pimelodus mysteriosus Azpelicueta, 1988: 88, figs. 1–3, 5–6, 8, 10, 12, 14. Type locality: Argentina, Misiones, Candelaria, arroyo Anselmo. Holotype: MLP 9191.
Distribution: Paraná River basin (Lundberg & Littmann, 2003).
Pimelodus navarroi Schultz, 1944
Pimelodus grosskopfii navarroi Schultz, 1944c: 207, pl. 1 (fig. c). Type locality: Venezuela, Río Palmar at the bridge
70 km southwest of Maracaibo. Holotype: USNM 121174.
Distribution: Lake Maracaibo basin (Lundberg & Littmann, 2003).
Pimelodus ornatus Kner, 1857
Pimelodus ornatus Kner, 1857: 411, pl. 6 (fig. 18). Type locality: Surinam, dem Rio negro und Cujaba [Brazil]. Syntypes: NMW 45832 (2), NMW 45843 (1).
Megalonema rhabdostigma Fowler, 1914: 256, fig. 10. Type locality: Rupununi River, British Guiana ... in the highlands of British Guiana, approximately secured in North Latitude 2º to 3º, and West Longitude 50º20'. Holotype:
ANSP 39338.
Distribution: Amazon, Corantijn, Essequibo, Orinoco and Paraná River basins and major rivers of the Guianas (Lundberg & Littmann, 2003).
Pimelodus ortmanni Haseman, 1911
Pimelodus ortmanni Haseman, 1911b: 379, pl. 50 (fig. 2). Type locality: Brazil, Porto União da Victoria, Rio
Iguassú. Holotype: FMNH 54240.
Distribution: Paraná River basin, Brazil (Lundberg & Littmann, 2003).
Pimelodus paranaensis Britski & Langeani, 1988
Pimelodus paranaensis Britski & Langeani, 1988: 410, figs. 1–2. Type locality: Ilha Solteira, Rio Paraná, SP [Brazil]. Holotype: MZUSP 23089.
Distribution: Upper Paraná River basin, Brazil (Lundberg & Littmann, 2003).
Pimelodus pictus Steindachner, 1877
Pimelodus pictus Steindachner, 1877a: 144. Type locality: aus dem Amazonenstrome auf peruanishen Gebiete, und

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aus dem Hyavary. Syntypes: NMW 45859 (3).
Distribution: Amazon and Orinoco River basins (Lundberg & Littmann, 2003).
Pimelodus platicirris Borodin, 1927
Pimelodus platicirris Borodin, 1927c: 2. Type locality: Salto de Pirassunungo, Rio Mogy Guassu, São Paulo Prov.,
Brazil. Holotype: AMNH 8628.
Distribution: Paraná River basin, Brazil; known only from the holotype (Lundberg & Littmann, 2003).
Pimelodus punctatus (Meek & Hildebrand, 1913)
Megalonema punctatum Meek & Hildebrand, 1913: 77. Type locality: Río Tuyra, Boca de Cupe, Panama. Holotype:
FMNH 7577.
Megalonema robustum Meek & Hildebrand, 1913: 78. Type locality: Río Tuyra, Marriganti, Panama. Holotype:
FMNH 7578.
Distribution: Tuira River basin, Panama (Lundberg & Littmann, 2003).
Species inquirendae, Pimelodus
Pimelodus rigidus Spix & Agassiz, 1829: 19, pl. 7 (fig. 2). Type locality: Brazil aequatoriali. Holotype: Whereabouts
unknown (Kottelat, 1988).
Pseudariodes pantherinus Lütken, 1874b: 192. Type locality: Caracas. Syntypes: BMNH 1876.1.10.10 (1), NMW 45851(1), USNM 44970 (1), ZMUC P 29640 (1), ZMUC P 29641 (1), ZMUC P 29642 (1).
Piramutana macrospila Günther, 1880a: 10, pl. 2. Type locality: Rio de la Plata. Holotype: BMNH 1878.9.10.14.
PINIRAMPUS Bleeker, 1858
Pinirampus Bleeker, 1858b: 198. Type species: Pimelodus pinirampus Agassiz [= Pimelodus pirinampu Spix &Agassiz, 1829]. Type by monotypy. Gender: Masculine.
Pirinampus Günther, 1864: 135. Type species: Pimelodus pirinampu Spix & Agassiz, 1829. Type by being a
replacement name. Gender: Masculine. Unjustified emendation of Pinirampus Bleeker, 1858.
Perugia Eigenmann & Norris, 1900: 355. Type species: Pirinampus [sic] agassizii Steindachner, 1876. Type byoriginal designation. Gender: Feminine.
Pinirampus pirinampu (Spix & Agassiz, 1829)
Pimelodus barbancho Humboldt, in Humboldt & Valenciennes, 1821: 172. Type locality: Guarico, l’Apure etd’autres rivières des steppes de Venezuela. Holotype: at MNHN.
Pimelodus pirinampu Spix & Agassiz, 1829: 20, pl. 8. Type locality: Brasiliae fluviis. Type(s): Whereabouts
unknown (Kottelat, 1988).
Pimelodus insignis Jardine, in Schomburgk, 1841: 180, pl. 6. Type locality: Rio Branco [Brazil]. No types known.
Galeichthys araguayensis Castelnau, 1855: 37, pl. 17 (fig. 3). Type locality: Rio Araguay [Brazil]. Holotype:
MNHN a-9366 (mounted).
Pinirampus typus Bleeker, 1862 (in Bleeker, 1862–63): 11. Type locality: Brasiliae fluviis. Type(s): Whereabouts
unknown. Unneeded replacement for Pimelodus pinirampu Spix & Agassiz, 1829, apparently to avoid Stricklandian tautonymy.
Pirinampus Agassizii Steindachner, 1876: 607, pl. 12. Type locality: Pará [Brazil]. Holotype: NMW 45955.
Distribution: Amazon, Essequibo, Orinoco, and Paraná River basins (Lundberg & Littmann, 2003).
Remarks: Pimelodus barbancho treated here as a nomen oblitum, following Lundberg & Littmann (2003).
PLATYNEMATICHTHYS Bleeker, 1858
Platynematichthys Bleeker, 1858b: 356. Type species: Bagrus punctulatus Kner, 1857. Type by monotypy. Gender:
Masculine.
Platypogon Starks, 1913: 28. Type species: Platypogon caerulorostris Starks, 1913. Type by monotypy. Gender:
Masculine.
Platynematichthys notatus (Jardine, 1841)
Pimelodus notatus Jardine, in Schomburgk, 1841: 181, pl. 7. Type locality: Fort St. Joaquim on the Rio Branco [Bra-CHECKLIST OF CATFISHES, RECENT AND FOSSIL

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zil]. No types known.
Bagrus punctulatus Kner, 1857: 380. Type locality: Forte do Principe am Rio Guaporé und Rio branco [Brazil].
Holotype: at NMW. Preoccupied by Bagrus punctulatus Castelnau, 1855; replaced by Bagrus nigropunctatus
Kner, 1858.
Bagrus nigropunctatus Kner, 1858: 345. Type locality: Forte do Principe am Rio Guapore und Rio branco [Brazil].
Holotype: at NMW. Replacement for Bagrus punctulatus Kner, 1857; preoccupied by Bagrus punctulatus
Castelnau, 1855.
Platypogon caerulorostris Starks, 1913: 29, pl. 5. Type locality: Brazil, Pará. Holotype: SU 22228.
Distribution: Amazon and Orinoco River basins (Lundberg & Littmann, 2003).
PLATYSILURUS Haseman, 1911
Platysilurus Haseman, 1911a: 320. Type species: Platysilurus barbatus Haseman, 1911. Type by monotypy. Gender:
Masculine.
Platysilurus malarmo Schultz, 1944
Platysilurus malarmo Schultz, 1944c: 234, fig. 3g, pl. 3 (fig. c). Type locality: Venezuela, Lago Maracaibo near themouth of the Río Concho. Holotype: USNM 121179.
Distribution: Lake Maracaibo basin (Lundberg & Littmann, 2003).
Platysilurus mucosus (Vaillant, 1880)
Platystoma mucosa Vaillant, 1880b: 151. Type locality: à Caldéron (Haute-Amazone). Holotype: MNHN a-1955.
Duoplatinus goeldii Steindachner, 1908b: 65. Type locality: Rio Purus [Brazil]. Holotype: at NMW.
Platysilurus barbatus Haseman, 1911: 320, pl. 52. Type locality: São Antonio de R. Madeira [Brazil]. Holotype:
FMNH 60306.
Distribution: Amazon and Orinoco River basins (Lundberg & Littmann, 2003).
Platysilurus olallae (Orcés, 1977)
Duopalatinus olallae Orcés, 1977: 78, fig. 4. Type locality: el bajo Bobonaza [Ecuador]. Holotype: at MEPN (132mm, catalog number not stated).
Distribution: Ecuador.
Remarks: Not treated in Lundberg & Littmann (2003).
PLATYSTOMATICHTHYS Bleeker, 1862
Platystomatichthys Bleeker, 1862 (in Bleeker, 1862–63): 10. Type species: Platystoma sturio Kner, 1857. Type byoriginal designation. Gender: Masculine.
Platystomatichthys sturio (Kner, 1857)
Platystoma sturio Kner, 1857: 395, pl. 3 (figs. 9–9a). Type locality: Rio branco [Brazil]. Holotype: at NMW.
Distribution: Amazon River basin (Lundberg & Littmann, 2003).
PROPIMELODUS Lundberg & Parisi, 2002
Propimelodus Lundberg & Parisi, 2002: 77. Type species: Pimelodus eigenmanni van der Stigchel, 1946. Type byoriginal designation.
Propimelodus eigenmanni (van der Stigchel, 1946)
Pimelodus eigenmanni van der Stigchel, 1946: 64. Type locality: Brazil. Holotype: RMNH 15465. Preoccupied by
Pimelodus eigenmanni Boulenger, 1891 [now Pimelodella eigenmanni, in the Heptapteridae]; apparently not
replaced.
Distribution: Amazon, Approuague, Kourou, and Oyapock River basins (Lundberg & Littmann, 2003).
Remarks: Pimelodus eigenmanni is tentatively treated as valid, despite being a junior primary homonym, following
Art. 23.9.5 of the Code of Zoological Nomenclature. Redescribed in Lundberg & Parisi (2002).
PSEUDOPLATYSTOMA Bleeker, 1862
Pseudoplatystoma Bleeker, 1862 (in Bleeker, 1862–63): 10. Type species: Silurus fasciatus Linnaeus, 1766. Type byoriginal designation. Gender: Neuter.

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FERRARISHemiplatystoma Bleeker, 1862 (in Bleeker, 1862–63): 10. Type species: Platystoma tigrinum Valenciennes, 1840.
Type by original designation. Gender: Neuter.
Remarks: Eigenmann & Eigenmann (1888b) serve as first revisers in giving precedence to Pseudoplatystoma overHemiplatystoma.
Pseudoplatystoma corruscans (Spix & Agassiz, 1829)
Platystoma corruscans Spix & Agassiz, 1829: 26 [pl. 13]. Type locality: in flumine S. Francisci [Brazil]. Holotype:
Whereabouts unknown (Kottelat, 1988). As Sorubim caparary on plate.
Sorubim caparary Spix in Spix & Agassiz, 1829: pl. 13. Type locality: in flumine S. Francisci [Brazil]. Holotype:
Whereabouts unknown (Kottelat, 1988). Name available from caption on plate.
Platystoma coruscans Valenciennes, in Cuvier & Valenciennes, 1840b: 17 (13 in the Strasbourg deluxe edition).
Misspelling or emendation of Platystoma corruscans Spix & Agassiz.
Silurus macrocephalus Larrañaga, 1923: 386. Type locality: Uruguay. No types known. Appears to correspond to
Silurus mangrullo on p. 377, which is a nomen nudum.
Distribution: São Francisco and Paraná River basins (Lundberg & Littmann, 2003)
Remarks: Agassiz (in Spix & Agassiz, 1831: conspectus) serves as first reviser in giving precedence to Platystoma
corruscans over Sorubim caparary.
Pseudoplatystoma fasciatum (Linnaeus, 1766)
Silurus fasciatus Linnaeus, 1766: 505. Type locality: Brasilia, Surinami. Holotype: Whereabouts unknown.
Platystoma artedii Günther, 1864: 106. Type locality: South America. Syntypes: On Mystus no. 6 in Seba (1734–65:
84, pl. 29, fig. 6) and Mystus no. 386 of Gronovius (1784: pl. 125).
Pseudoplatystoma fasciatum intermedium Eigenmann & Eigenmann, 1888b: 138. Type locality: Obidos. Holotype:
MCZ 7321.
Pseudoplatystoma fasciatum brevefile Eigenmann & Eigenmann, 1889b: 31. Type locality: Goyaz [Brazil]. Holotype: MCZ 7317.
Pseudoplatystoma fasciatum reticulatum Eigenmann & Eigenmann, 1889b: 30. Type locality: Rio Negro [Brazil].
Holotype: MCZ 23813.
Pseudoplatystoma fasciatum nigricans Eigenmann & Eigenmann, 1889b: 31. Type locality: Xingu. Syntypes (2):MCZ 7301 (1), MCZ 7311 (1).
Distribution: Amazon, Corantijn, Essequibo, Orinoco, and Paraná River basins (Lundberg & Littmann, 2003);
Magdalena River basin (Maldonado-Ocampo et al., 2005).
Pseudoplatystoma tigrinum (Valenciennes, 1840)
Platystoma truncatum Spix & Agassiz, 1829: 27, pl. 13a. Type locality: Brazil, Japurá and Solimoen. Type (s):Whereabouts unknown (Kottelat, 1988).
Platystoma tigrinum Valenciennes, in Cuvier & Valenciennes, 1840b: 10 (8 of Strasbourg deluxe edition), pl. 422.
Type locality: probablement originaire du Brésil. Holotype: MNHN a-9354 (mounted).
Distribution: Amazon and Orinoco River basins (Lundberg & Littmann, 2003).
Remarks: Synonymy follows Lundberg & Littmann (2003) which differs from most previous accounts, in which
Platystoma truncatum was treated as a synonym of Pseudoplatystoma fasciatum. Because of this, Platystoma
truncatum became the senior synonym for a widely used Pseudoplatystoma tigrinum. P. truncatum was treatedas a nomen oblitum in Lundberg & Littmann (2003), but without supporting documentation. Use of P. truncatum as a valid name has not been found in surveyed compilations of late 19th or 20th century and, therefore, theuse of P. tigrinum as valid is continued here.
Species inquirendae, Pseudoplatystoma
Platystoma pardalis Valenciennes, 1836, in Valenciennes, 1835–47: pl. 4 (fig. 2). Type locality: Buenos Aires, Argentina. Holotype: MNHN a-8833. Name available from plate, mentioned in Valenciennes (1847: 6), as Platystoma
panthale. Described in Cuvier & Valenciennes (1840b: 15 (12 of Strasbourg deluxe edition)), as Platystoma pardale.
Platystoma orbignianus Valenciennes, 1836, in Valenciennes, 1835–47: pl. 4 (fig. 3). Type locality: Brazil. Holotype:
MNHN b-0160. Name available from plate, description in Valenciennes (1847: 6), as Platystoma orbignyanum.

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Description in Cuvier & Valenciennes (1840b: 12, [9 of Strasbourg deluxe edition]).
Platystoma panthale Valenciennes, 1847: 6. Type locality: [Not stated]. Holotype: MNHN a-8833. New name proposed
for Platystoma pardalis Valenciennes, 1836, without explanation.
Platystoma punctifer Castelnau, 1855: 40, pl. 19 (fig. 2). Type locality: de l’ Amazone. Holotype: MNHN 0000-1582.
SORUBIM Cuvier, 1829
Sorubim Cuvier, 1829: 293. Type species: Silurus lima Bloch & Schneider, 1801. Type by subsequent designation,
apparently by Kottelat (1988). Gender: Masculine.
Platystoma Spix & Agassiz, 1829: 23. Type species: Silurus lima Bloch & Schneider, 1801. Type by subsequent designation by Jordan (1917: 131). Gender: Neuter. Preoccupied by Platystoma Meigen, 1803 (Diptera).
Sorubim Spix & Agassiz, 1829: 24. Type species: Silurus lima Bloch & Schneider, 1801. Type by subsequent designation by Bleeker (1862–63: 10). Gender: Masculine. First appeared in synonymy of Platystoma Spix & Agassiz and therefore not available, but made available because of use as a valid name by Bleeker (1862–63: 10)[ICZN art. 11.6.1]. Preoccupied by Sorubim Cuvier, 1829.
Sorubium Swainson, 1838: 356. Type species: Silurus lima Bloch & Schneider, 1801. Type by being a replacement
name. Unneeded replacement for Sorubim Spix & Agassiz, 1829.
Abron Gistel, 1848: X. Type species: Silurus lima Bloch & Schneider, 1801. Type by being a replacement name.
Gender: Neuter. Replacement for Platystoma Valenciennes [= Spix & Agassiz, 1829].
Remarks: Sorubim was apparently adopted by Cuvier (1829) from the Spix & Agassiz (1829) manuscript, but thename was published first in the former book.
Revision: Littmann (1998).
Sorubim cuspicaudus Littmann, Burr & Nass, 2000
Sorubim cuspicaudus Littmann, Burr & Nass, 2000: 903, fig. 2b, 3, 4c. Type locality: Colombia, Departamento de
Tolima, Río Magdalena drainage, at Puerto Soplaviento. Holotype: FMNH 56223.
Distribution: Magdalena and Sinu River basins, Colombia, and Lake Maracaibo (Lundberg & Littmann, 2003).
Sorubim elongatus Littmann, Burr, Schmidt & Isern, 2001
Sorubim elongatus Littmann, Burr, Schmidt & Isern, 2001: 4, fig. 2. Type locality: Peru: Departamento Loreto,
Maynas Province, Ullpa Caño, 50 m upstream of confluence with Moena Caño, Río Itaya drainage, Río Amazonas basin (approx. 3º46'20"S 73º14'17"W). Holotype: SIUC 30303.
Distribution: Amazon, Essequibo, and Orinoco River basins (Lundberg & Littmann, 2003).
Sorubim lima (Bloch & Schneider, 1801)
Silurus lima Bloch & Schneider, 1801: 384. Type locality: in flumine Maranham Brasiliae. Holotype: ZMB 3185
(stuffed).
Sorubim infraoculare Spix, in Spix & Agassiz, 1829: pl. 15. Type locality: Brasiliae aequatorialis fluviis. Holotype:
Whereabouts unknown (Kottelat, 1988). Name available from caption on plate.
Platystoma Luceri Weyenbergh, 1877: 10, pl. 3 (figs. 1–3). Type locality: Santa-Fe [Argentina]. Holotype: Whereabouts unknown.
Sorubim latirostris Miranda Ribeiro, 1920: 14, pl. 12. Type locality: Amazonas [Brazil]. Holotype: MNRJ 631.
Distribution: Amazon, Orinoco, Paraná and Parnaíba River basin (Lundberg & Littmann, 2003).
Sorubim maniradii Littmann, Burr & Buitrago-Suárez, 2001
Sorubim maniradii Littmann, Burr & Buitrago-Suárez, 2001: 88, fig. 1. Type locality: Ecuador, Napo State, Río
Yasuni, Río Napo drainage, 0º59'06''S 75º25'36''W. Holotype: FMNH 108814.
Distribution: Amazon River basin (Lundberg & Littmann, 2003).
Sorubim trigonocephalus Miranda Ribeiro, 1920
Sorubim trigonocephalus Miranda Ribeiro, 1920: 12, pls. 10, 11. Type locality: Porto Velho [Brazil]. Holotype:
MNRJ 882.
Distribution: Madeira and Tapajós River basins, Brazil (Lundberg & Littmann, 2003).
SORUBIMICHTHYS Bleeker, 1862
Sorubimichthys Bleeker, 1862, in Bleeker, 1862–63: 10. Type species: Sorubim jandia Spix & Agassiz, 1829. Type342 · Zootaxa 1418 © 2007 Magnolia Press

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by original designation. Gender: Masculine.
Pteroglanis Eigenmann & Pearson, in Pearson, 1924: 9. Type species: Pteroglanis manni Eigenmann & Pearson,
1924. Type by monotypy. Gender: Masculine.
Sorubimichthys planiceps (Spix & Agassiz, 1829)
Platystoma planiceps Spix & Agassiz, 1829: 25, pl. 12. Type locality: Brazil: in fluminae Amazonum, Solimoëns,Rio Negro. Possible syntype or lectotype: MHNN 811(Kottelat, 1988).
Sorubim Pirauáca Spix, in Spix & Agassiz, 1829: pl. 12. Type locality: Brazil: in fluminae Amazonum, Solimoëns,Rio Negro. Possible holotype: MHNN 811(Kottelat, 1988). Name available from caption on plate; as Platystoma planiceps in text.
Sorubim jandia Spix & Agassiz, 1829: pl. 14. Type locality: equatorialis fluviis [Brazil]. Holotype: Whereabouts
unknown (Kottelat, 1988). Name available from caption on plate; as Platystoma spatula in text.
Platystoma spatula Spix & Agassiz, 1829: 26, pl. 14. Type locality: equatorialis fluviis [Brazil]. Types: Whereabouts unknown (Kottelat, 1988).
Sorubimichthys ortoni Gill, 1870: 94. Type locality: Maranon, or Upper Amazon, and Napo Rivers. Holotype:
USNM 8238.
Pteroglanis manni Eigenmann & Pearson, in Pearson, 1924: 9, pl. 1 (fig. 1). Type locality: Bolivia, Little Rio Negro,
tributary to Rio Beni. Holotype: CAS 59623.
Distribution: Amazon and Orinoco River basins (Lundberg & Littmann, 2003).
Remarks: Lundberg et al. (1989) reviewed the nomenclature of this species. Agassiz (in Spix & Agassiz, 1831: conspectus) serves as first reviser in treating Platystoma planiceps as valid over Sorubim Pirauáca, and Platystoma
spatula over Sorubim jandia. Precedence of Platystoma planiceps over Platystoma spatula may be by action of
Lundberg et al. (1989).
STEINDACHNERIDION Eigenmann & Eigenmann, 1919
Steindachneria Eigenmann & Eigenmann, 1888b: 137. Type species: Steindachneria amblyurus Eigenmann &Eigenmann, 1888. Type by original designation. Gender: Feminine. Preoccupied by Steindachneria Goode &Bean, 1888 (in fishes), replaced by Steindachneridion Eigenmann & Eigenmann, 1919.
Steindachneridion Eigenmann & Eigenmann, 1919: 525. Type species: Steindachneria amblyurus Eigenmann &Eigenmann, 1888. Type by being a replacement name. Gender: Neuter. Replacement for Steindachneria Eigenmann & Eigenmann, 1888.
Revision, with key to species: Garavello (2005).
Steindachneridion amblyurum (Eigenmann & Eigenmann, 1888)
Steindachneria amblyurus Eigenmann & Eigenmann, 1888b: 137. Type locality: Jequitinhonha [Brazil]. Syntypes(2, 35–38 cm): MCZ 7324 (2).
Distribution: Jequitinhonha River basin, Brazil (Garavello, 2005).
Steindachneridion doceana (Eigenmann & Eigenmann, 1889)
Steindachneria doceana Eigenmann & Eigenmann, 1889b: 30. Type locality: Rio Doce [Brazil]. Syntypes (9): MCZ
23792 (1), MCZ 23793 (1), MCZ 23794 (7); one syntype illustrated in Garavello (2005: 614, fig. 6).
Distribution: Doce River basin, Brazil (Garavello, 2005).
† Steindachneridion iheringi (Woodward, 1899)† Arius iheringi Woodward, 1899: 64, pl. 2 (fig. 1), pl. 3 (fig. 2). Type locality: Taubaté, Estado do Sao Paulo, Brazil; Tertiary Lignite. Syntypes: BMNH P.9220 (skull), Museu Paulista (nearly complete specimen), Senckenberg Museum (partial specimen).
Distribution: Paraíba River basin, Estado do São Paulo, Brazil; Tertiary (Santos, 1973).
Remarks: Species assigned to the Pimelodidae and the genus Steindachneridion by Santos (1973).
Steindachneridion melanodermatum Garavello, 2005
Steindachneridion melanodermatum Garavello, 2005: 618, fig. 9. Type locality: Brazil, Paraná State, municipality of
Quedas do Iguaçu, rio Iguaçu, down river of the Salto Osório dam (approx. 25º35'S 53º05'W). Holotype:
MZUSP 87983.
Distribution: Iguaçu River, upper Paraná River Basin, Brazil (Garavello, 2005).

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Steindachneridion parahybae (Steindachner, 1877)
Platystoma Parahybae Steindachner, 1877b: 640, pl. 9. Type locality: Rio Parahyba bei Juiz de Fora; Rio Jequitinhonha [Brazil]. Syntypes: MCZ 25521 (1, skeleton), MCZ 7295 (1), MCZ 7323 (2), NMW (?).
Distribution: Paraíba do Sul River basin, Brazil (Garavello, 2005).
Remarks: Garavello (2005: 610) incorrectly reported the catalog number of one lot of the types as MCZ 73231.
Redescribed in Oliveira & de Moraes (1997).
Steindachneridion punctatum (Miranda Ribeiro, 1918)
Steindachneria scripta punctata Miranda Ribeiro, 1918d: 642. Type locality: Itaqui, Rio Grande do Sul [Brazil].
Holotype: MNRJ 1167. Originally Steindachneria scripta var. punctata.
Distribution: Upper Uruguay and Paraná River basin, Brazil (Garavello, 2005).
Steindachneridion scriptum (Miranda Ribeiro, 1918)
Steindachneria scripta Miranda Ribeiro, 1918d: 640. Type locality: Itaqui [Rio Grande do Sul, Brazil]. Lectotype:
MZUSP 2286, designated by Britski (1969).
Distribution: Upper Paraná and Uruguay River basins, Brazil (Garavello, 2005).
† Steindachneridion silvasantosi Figueiredo & Costa-Carvlho, 1999
† Steindachneridion silvasantosi Figueiredo & Costa-Carvlho, 1999: 685, figs. 1–4. Type locality: Tremembé Formation, Lacustrine sedimentary deposit that outcrops in the mine Nossa Mina Senhora da Guia, Tremembé
county, city of Taubaté, São Paulo State [Brazil]; Tertiary (? Oligocene). Holotype: DGM 1291-P; partial skull,including neurocranium and suspensorium.
Distribution: Paraíba River basin, São Paulo State, Brazil; Tertiary (Figueiredo & Costa-Carvlho, 1999).
ZUNGARO Bleeker, 1858
Zungaro Bleeker, 1858b: 196. Type species: Zungaro humboldtii Bleeker [=Pimelodus zungaro Humboldt, 1821].
Type by absolute tautonymy. Gender: Masculine.
Paulicea Ihering, 1898: 108. Type species: Paulicea jahu Ihering, 1898. Type by subsequent designation by Eigenmann (1910). Gender: Feminine.
Remarks: Generic synonymy based on Silfvergrip (1992: 306).
Zungaro jahu (Ihering, 1898)
Paulicea jahu Ihering, 1898: 108. Type locality: Sao Paulo [Brazil]. Types: Whereabouts unknown.
Distribution: Paraná-Paraguay River basin (Lundberg & Littmann, 2003).
Zungaro zungaro (Humboldt, 1821)
Pimelodus zungaro Humboldt, in Humboldt & Valenciennes, 1821: 170, pl. 46 (fig. 1). Type locality: Tomependa,
Río Marañon [Amazon River basin, Peru]. Holotype: Whereabouts unknown.
Bagrus flavicans Castelnau, 1855: 31, pl. 13 (fig. 2). Type locality: Brazil, l’Amazone. Holotype: MNHN a-8823(mounted).
Zungaro humboldtii Bleeker, 1858b: 207. Type locality: Tomedenda, Río Marañon [Amazon River basin, Peru].
Holotype: Whereabouts unknown. Unneeded replacement name for Pimelodus zungaro, presumably to avoid
tautomymy.
Platystoma Lütkeni Steindachner, 1876: 609, pl. 13. Type locality: Laufe des Amazonenstromes gefangen. Syntypes:
at NMW.
Distribution: Amazon and Orinoco River basins (Lundberg & Littmann, 2003).
ZUNGAROPSIS Steindachner, 1908
Zungaropsis Steindachner, 1908b: 67. Type species: Zungaropsis multimaculatus Steindachner, 1908. Type bymonotypy. Gender: Feminine.
Zungaropsis multimaculatus Steindachner, 1908
Zungaropsis multimaculatus Steindachner, 1908b: 67. Type locality: Rio Xingu. Holotype: NMW 43537.
Remarks: Considered to be a valid species, in a genus near to Zungaro by A. Silfvergrip (pers. commun., 2003).

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Species inquirenda, Pimelodidae
Pimelodus (Bagrus) maculatus Jardine, in Schomburgk, 1841: 175, fig. Type locality: Most rivers of Guyana, also Rio
Negro and Amazon River. No types known. Preoccupied by Pimelodus maculatus La Cepède, 1803.

PLOTOSIDAE Bleeker, 1858
Plotosichthyoidei Bleeker, 1858b: 49, 310. Type genus: Plotosus La Cepède, 1803.
Taxonomic summaries: Taylor & Gomon (1986, Africa); Tilak (1970a, South Asia); Hoese & Hanley, in Paxton et al.
(1989, Australia).
Reviews: Allen (1989, Australia); Allen (1991, New Guinea); Ferraris (1999a, western central Pacific).
Identification guide: Jayaram (1982, South Asia).
Remarks: Whitley (1940b: 407) noted that the names Plotosus laticeps, Copidoglanis labiosus, Copidoglanis labrosus,
Copidoglanis levis and Copidoglanis curtus, all found in Saville-Kent (1889), are nomina nuda and therefore
unavailable. The name Plotosus flavolineatus found in Whitley (1941a: 311) is also a nomen nudum and not available.
10 genera, 35 species; no named fossil taxa.
Incertae sedis
Plotosus argenteus Zietz, 1896
Plotosus argenteus Zietz, 1896: 410, pl. 16 (fig. 7). Type locality: Barcoo River, Finke River at Hermannsburg andIdracowra, and in Plam Creek and Ilpilla Creek, [Central Australia]; [ restricted to Cooper Creek, near
Innamincka, (which is apparently the locality listed by Zeitz (1896:411) as Barcoo River) by lectotype designation]. Lectotype: SAMA F1090, designated by Unmack (2001: 57).
Distribution: Cooper Creek, Bulloo and Finke Rivers, internal drainage system, Australia (Allen, 1989).
Remarks: Treated in recent literature as valid either as Neosilurus argenteus or Porochilus argenteus.
Copidoglanis rendahli Whitley, 1928
Copidoglanis obscurus Rendahl, 1922: 173, fig. 5. Type locality: Glencoe, Hermit Hill, nw. Australia. Lectotype:ZMUO J6341, designated by Pethon (1969: 5). Preoccupied by Copidoglanis obscurus Günther, 1864, replaced
by Copidoglanis rendahli Whitley, 1928.
Copidoglanis rendahli Whitley, 1928: 214. Type locality: Glencoe, Hermit Hill, N.W. Australia. Lectotype: ZMUO
J6341 designated by Pethon (1969: 5) for Copidoglanis obscurus Rendahl. Replacement for Copidoglanis
obscurus Rendahl 1922; preoccupied by Copidoglanis obscurus Günther, 1864.
Distribution: Widely distributed in northern Australia in isolated localities, including Fitzroy, Ord, Jardin Rivers,
and streams of Arnhem Land (Allen, 1989).
Remarks: Treated in recent literature as valid either as Neosilurus rendahli or Porochilus rendahli.
ANODONTIGLANIS Rendahl, 1922
Anodontiglanis Rendahl, 1922: 168. Type species: Anodontiglanis dahli Rendahl, 1922. Type by original designation. Gender: Masculine.
Anodontiglanis dahli Rendahl, 1922
Anodontiglanis dahli Rendahl, 1922: 169, figs. 2–3. Type locality: Glencoe, Northern Territory, Australia. Holotype:ZMUO J361.
Distribution: Northern Australia, including Fitzroy, Daly, East Alligator, Roper, Mitchell and Archer Rivers (Allen,1989), in lentic and lotic freshwater habitats (Paxton et al., 1989).
CNIDOGLANIS Günther, 1864
Cnidoglanis Günther, 1864: 27. Type species: Plotosus megastomus Richardson, 1845. Type by subsequent designation by Jordan (1919b: 332). Gender: Masculine.
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Choeroplotosus Kner, 1866: 545. Choeroplotosus decemfilis Kner, 1867. Type by subsequent monotypy. Nameappeared first without an available species name. Also described as new in Kner (1867: 300). Gender: Masculine.
Neoplotosus Castelnau, 1875: 45. Type species: Neoplotosus waterhousii Castelnau, 1875. Type by monotypy. Gender: Masculine.
Ostophycephalus Ogilby, 1899: 155. Type species: Ostophycephalus duriceps Ogilby, 1899. Type by original designation. Gender: Masculine.
Remarks: Eschmeyer & Bailey (1990: 92) interpreted the availability of Choeroplotosus as follows: “In 1866: 545,
Kner listed one species as, "Choeroplotosus limbatus (Plotos. limbatus ? C. V.)", but in 1867 (p. 300) he
described the species as Choeroplotosus decemfilis n. sp., with "Syn.? an Plotosus limbatus C. V. Apparently
the genus can date to Kner (1866: 545) with no definitely-included named species, with species added in 1867.”
Cnidoglanis macrocephalus (Valenciennes, 1840)
Plotosus macrocephalus Valenciennes, in Cuvier & Valenciennes, 1840b: 428 (317 in Strasbourg deluxe edition), pl.
449. Type locality: prise à Timor [in error, possibly for Tasmania, Australia]. Holotype: MNHN a-8921.
Plotosus megastomus Richardson, 1845, in Richardson, 1844–48: 31, pl. 21 (figs. 1–3). Type locality: Sydney Cove;
Australia. Holotype: BMNH 1974.5.22.1.
Choeroplotosus decemfilis Kner, 1867: 300, pl. 12 (fig. 1 [as Choeroplotosus limbatus]). Type locality: Von Sidney.
Syntypes (2): NMW 47098 (1), NMW 47100 (1).
Cnidoglanis bostockii Castelnau, 1873: 140. Type locality: Fremantle, Western Australia. Syntypes (2): Whereabouts unknown.
Neoplotosus waterhousii Castelnau, 1875: 45. Type locality: Adelaide, South Australia. Holotype: MNHN a-9545.
Ostophycephalus duriceps Ogilby, 1899: 156. Type locality: St. Vincent’s Gulf, South Australia. Holotype: SAMA
F1093.
Distribution: Kirra, Queensland to Jervis Bay, New South Wales, and Kingston, South Australia to Houtman Abrolhos Islands, Western Australia and Duck River, Tasmania, in near-shore and reef habitats (Paxton et al., 1989).
Remarks: Kowarsky (1976) discussed the valid name for, and distribution of, this species.
EURISTHMUS Ogilby, 1899
Euristhmus Ogilby, 1899: 154. Type species: Plotosus elongatus Castelnau, 1878. Type by original designation.
Gender: Masculine.
Exilichthys Whitley, 1933: 65. Type species: Cnidoglanis nudiceps Günther, 1880. Type by original designation.
Gender: Masculine.
Euristhmus lepturus (Günther, 1864)
Cnidoglanis lepturus Günther, 1864: 28. Type locality: Sydney, New South Wales [Australia]. Syntypes (2, one
stuffed): BMNH 1864.1.17.33 (1).
Plotosus elongatus Castelnau, 1878a: 237. Type locality: Brisbane River [Queensland, Australia]. Holotype: Whereabouts unknown [possibly MNHN a-2783].
Distribution: Exmouth Gulf, Western Australia to Sydney, New South Wales, and New Guinea, in freshwater, estuarine and near-shore marine habitats (Paxton et al., 1989; Allen, 1997: 60).
Euristhmus microceps (Richardson, 1845)
Plotosus microceps Richardson, 1845, in Richardson, 1844–48: 31, pl. 21 (figs. 4–7). Type locality: North-west
coast of Australia. Holotype: BMNH 1846.3.3.2.
Cnidoglanis microcephalus Günther, 1864: 28. Type locality: North-west coast of Australia. Holotype: BMNH
1846.3.3.2. Unjustified emendation of Plotosus microceps Richardson, 1845.
Distribution: Shark Bay to Broome, Western Australia, in near-shore habitats over soft bottoms (Paxton et al., 1989).
Euristhmus nudiceps (Günther, 1880)
Cnidoglanis nudiceps Günther, 1880b: 49. Type locality: Arafura Sea. Holotype or lectotype: BMNH
1879.5.14.590; illustrated in Weber & de Beaufort (1913: 232).
Distribution: Shark Bay, Western Australia to Brisbane Queensland, Australia, in near-shore habitats over soft bottoms (Paxton et al., 1989); also New Guinea (Allen, 1997: 60).

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Remarks: Paxton et al. (1989: 223) list a holotype for Cnidoglanis nudiceps, but the Natural History Museum in
London lists a second specimen (BMNH 1890.2.26.165) from the Challenger Expedition taken in the Arafura
Sea that may also have been examined by Günther. There is no evidence in the original description whether
more than one specimen was examined. Therefore, even if the second specimen was originally a syntype, itwould have been rendered a paralectotype by the de facto lectotype designation in Paxton et al. (1989).
NEOSILUROIDES Allen & Feinberg, 1998
Neosiluroides Allen & Feinberg, 1998: 11. Neosiluroides cooperensis Allen & Feinberg, 1998. Type by original designation. Gender: Masculine.
Neosiluroides cooperensis Allen & Feinberg, 1998
Neosiluroides cooperensis Allen & Feinberg, 1998: 12, fig. 2. Type locality: Callamurra Waterhole, Coopers Creek,
11.3 km east of Innamincka, South Australia, ca. 27º41'S, 140º51'E. Holotype: AMS I.18699-001.
Distribution: Coopers Creek system, Lake Eyre drainage (Allen & Feinberg, 1998).
NEOSILURUS Steindachner, 1867
Neosilurus Steindachner, 1867a: 7. Type species: Neosilurus hyrtlii Steindachner, 1867. Type by monotypy. Gender:
Masculine. Also appeared as new in Steindachner (1867d: 11).
Neosilurus Castelnau, 1878a: 238. Type species: Neosilurus australis Castelnau, 1878. Type by monotypy. Gender:
Masculine. Preoccupied by Neosilurus Steindachner, 1867, in fishes; replaced by Cainosilurus Macleay, 1881.
Eumeda Castelnau, 1878b: 143. Type species: Eumeda elongata Castelnau, 1878. Type by monotypy. Gender: Feminine.
Cainosilurus Macleay, 1881: 211. Type species: Neosilurus australis Castelnau, 1878. Type by being a replacement
name. Gender: Masculine. Replacement for Neosilurus Castelnau, 1878; preoccupied by Neosilurus Steindachner, 1867.
Lambertia Perugia, 1894: 550. Type species: Lambertia atra Perugia, 1894. Type by monotypy. Gender: Feminine.
Preoccupied by Lambertia Robineau-Desvoidy, 1863, in Diptera; replaced by Lambertichthys Whitley, 1938.
Anyperistius Ogilby, 1908: 3, 11. Type species: Anyperistius perugiae Ogilby, 1908. Type by original designation.
Gender: Masculine. Spelled two ways originally: Anyperistius and Anyperisteus. First reviser apparently
Eschmeyer & Bailey (in Eschmeyer, 1990) who selected Anyperistius.
Lambertichthys Whitley, 1938: 223. Type species: Lambertia atra Perugia, 1894. Type by being a replacement
name. Gender: Masculine. Replacement for Lambertia Perugia, 1894; preoccupied by Lambertia RobineauDesvoidy, 1863, in Diptera.
Neosilurus ater (Perugia, 1894)
Lambertia atra Perugia, 1894: 551. Type locality: Inawi, Papuasia orientale. Syntypes: MSNG 1663 (1), MSNG
8140 (1, missing), ZMA 113360 (1).
Neosilurus mediobarbis Ogilby, 1908: 12. Type locality: ? Queensland, Australia. Holotype: QM (lost: Paxton et al.,1989).
Lambertichthys ater sepikensis Whitley, 1956b: 68. Type locality: Upper Sepik River, New Guinea. Holotype: AMS
IA.7278; illustrated in Whitley (1956a: 44, fig. 4).
Distribution: Northern Australia and central-southern New Guinea; in freshwater (Allen, 1989).
Neosilurus brevidorsalis (Günther, 1867)
Copidoglanis brevidorsalis Günther, 1867a: 66. Type locality: Nicol Bay, Cape York, Queensland, Australia. Holotype: BMNH 1867.5.13.11.
Anyperistius perugiae Ogilby, 1908: 11. Type locality: Inawe, St. Joseph River, New Guinea. Type(s): at MSNG
and/or BMNH 1965.1.26.1 (1). Based on specimen(s) identified by Perugia (1894: 552) as Eumeda elongata(not of Castelnau).
Neosilurus bartoni Regan, 1908c: 153. Type locality: Sogeri, New Guinea. Syntypes: BMNH 1905.8.15.9–10 (2).
Distribution: Cape York Peninsula, northern Australia, and central-southern New Guinea; in streams, turbid backwaters, and lagoons (Allen, 1989).
Remarks: BMNH 1965.1.26.1 was sent to BMNH on exchange from MSNG and is apparently a type of Anyperistius

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perugiae Ogilby, 1908.
Neosilurus coatesi (Allen, 1985)
Tandanus coatesi Allen, 1985: 252, fig. 3. Type locality: Ninar River, ca. 6 km west of Maprik, Papua New Guinea,
ca. 3º37'S, 143º00'E. Holotype: WAM P.27839-007.
Distribution: Sepik River basin, in streams of Torracelli Mountains (Allen et al., 1992).
Neosilurus equinus (Weber, 1913)
Copidoglanis equinus Weber, 1913: 527, fig. 11. Type locality: Lorentz-Fluss [New Guinea]. Syntypes (13): ZMA
111105 (7), ZMA 111106 (1), ZMA 111107 (4), ZMA 111108 (2).
Distribution: Southern Papua (formerly Irian Jaya), New Guinea (Allen, 1996).
Neosilurus gjellerupi (Weber, 1913)
Copidoglanis gjellerupi Weber, 1913: 528. Type locality: Kaiserin Augusta-fluss [New Guinea]. Holotype: ZMA
111091.
Distribution: Ramu River basin, Papua New Guinea (Allen et al., 1992) and northern Papua (formerly Irian Jaya),
New Guinea (Allen, 1996).
Remarks: The name Copidoglanis papuensis that is found in the caption of figures 11 and 12 in Hase (1914: 540)
appears to have been published accidentally inasmuch as the species account indicates that the specimens wereregarded as having been identified as Copidoglanis gjellerupi (Weber, 1913). This was treated by Eschmeyer etal. (1998: 1283) as unavailable, because it was not treated as the valid name of a species when proposed.
Neosilurus gloveri Allen & Feinberg, 1998
Neosilurus gloveri Allen & Feinberg, 1998: 13, fig. 3. Type locality: Main Spring, Dalhousie Springs, South Australia, ca. 26º25'S, 135º30'E. Holotype: SAMA F.4159.
Distribution: Dalhousie artesian springs, Finke River basin, southern Australia (Allen & Feinberg, 1998).
Neosilurus hyrtlii Steindachner, 1867
Neosilurus Hyrtlii Steindachner, 1867d: 14, pl. 1 (figs. 3–3a). Type locality: Fitzroy-Flusse bei Rockhampton, OstAustralien. Syntypes: NMW 45340 (5), NMW 45341 (2).
Silurichthys australis Castelnau, 1875: 45. Type locality: Cape York [Queensland, Australia]. Holotype: Whereabouts unknown.
Neosilurus australis Castelnau, 1878a: 239. Type locality: Freshwater lagoons of Rockhampton, Queensland, Australia. Holotype: Whereabouts unknown.
Eumeda elongata Castelnau, 1878b: 144. Type locality: Brisbane River, Rockhampton [Queensland, Australia].
Holotype: MNHN a-2173.
Neosilurus robustus Ogilby, 1908: 13. Type locality: Keppel Bay, Queensland, Australia. Holotype: Whereabouts
unknown, not at QM.
Copidoglanis glencoensis Rendahl, 1922: 170, fig. 4. Type locality: Glencoe R., nw. Australia. Lectotype: ZMUO
J5254; Lectotype designated by Pethon (1969: 4).
Neosilurus mortoni Whitley, 1941b: 7, fig. 7. Type locality: Yam Creek (nine miles from Brock's Creek, railway line
south from Darwin) Northern Territory of Australia. Holotype: AMS IA.4824.
Distribution: Widely distributed in northern Australia; in freshwater (Allen, 1989).
Neosilurus idenburgi (Nichols, 1940)
Copidoglanis idenburgi Nichols, 1940: 1. Type locality: Bernhard Camp (altitude 75 meters), Idenburg River, Netherland New Guinea. Holotype: AMNH 15034.
Distribution: Ramu River basin, Papua New Guinea (Allen, 1992) and northern Papua (formerly Irian Jaya), New
Guinea (Allen, 1996).
Neosilurus mollespiculum Allen & Feinberg, 1998
Neosilurus mollespiculum Allen & Feinberg, 1998: 16, fig. 6. Type locality: Running River, Burdekin River system,
Queensland, ca. 19º07'S, 145º50'E, Australia. Holotype: QM I.30685.
Distribution: Isdell, Carson, Drysdale, Katherine, Fergusson, and Burdekin Rivers, northern Australia (Allen &Feinberg, 1998).
Neosilurus novaeguineae (Weber, 1907)
Copidoglanis novae-guineae Weber, 1907: 226. Type locality: Sentani-See, Nord-Neu-Guinea. Syntypes (13):348 · Zootaxa 1418 © 2007 Magnolia Press

FERRARISFMNH 52388 (1), NMW 46696 (1), RMNH 7980 (1), ZMA 112670 (8).
Copidoglanis novae-guineae niger Nichols, 1940: 1. Type locality: Bernhard Camp (altitude 75 meters), Idenburg
River, Netherland New Guinea. Holotype: AMNH 15035.
Distribution: Ramu River basin, Papua New Guinea (Allen et al., 1992) and northern Papua (formerly Irian Jaya),
New Guinea (Allen, 1996).
Neosilurus pseudospinosus Allen & Feinberg, 1998
Neosilurus pseudospinosus Allen & Feinberg, 1998: 15, figs. 4–5. Type locality: Rocky pool of Ord R. on Old Lissadel Station, Kimberley Dist., Western Australia, ca. 16º40'S, 128º83'E. Holotype: WAM P.28505-007.
Distribution: Ord River, Western Australia (Allen & Feinberg, 1998).
OLOPLOTOSUS Weber, 1913
Oloplotosus Weber, 1913: 521 Type species: Oloplotosus mariae Weber, 1913. Type by monotypy. Gender: Masculine.
Key: Allen (1985; 1991).
Oloplotosus luteus Gomon & Roberts, 1978
Oloplotosus luteus Gomon & Roberts, in Roberts, 1978: 47, fig. 25. Type locality: Shallow, turbid backwater of Ok
Tedi and lowermost half km of small tributary about 16 km NE of Nigerum, 915 km upriver from Toro Pass,
5º34.1'S, 141º15.0'E; Fly River, Papua New Guinea. Holotype: AMS I.27093-001.
Distribution: Upper Fly River, New Guinea (Roberts, 1978).
Oloplotosus mariae Weber, 1913
Oloplotosus Mariae Weber, 1913: 522, fig. 3. Type locality: Lorentz-Fluss, Sabang; Lorentz-Fluss, Alkmaar [NewGuinea]. Syntypes (5): AMNH 9500 (1), ZMA 111103 (1), ZMA 111104 (2).
Distribution: Southern Papua (formerly Irian Jaya), New Guinea (Allen, 1996).
Oloplotosus torobo Allen, 1985
Oloplotosus torobo Allen, 1985: 248, fig. 1. Type locality: Soro River at n. end of Lake Kutubu, ca. 4 km east ofMoro landing strip, Papua New Guinea, ca. 6º23'S, 143º15'E, 0–3 m. Holotype: WAM P.28158-003.
Distribution: Soro River, Papua New Guinea (Allen, 1985).
PARAPLOTOSUS Bleeker, 1863
Paraplotosus Bleeker, 1863 (in Bleeker, 1862–63): 100. Type species: Plotosus albilabris Valenciennes, 1840. Typeby monotypy. Gender: Masculine.
Endorrhis Ogilby, 1898b: 283. Type species: Copidoglanis longifilis Macleay, 1881. Type by original designation.
Gender: Feminine.
Revision: Allen (1998).
Paraplotosus albilabris (Valenciennes, 1840)
Plotosus albilabris Valenciennes, in Cuvier & Valenciennes, 1840b: 427 (316 in Strasbourg deluxe edition). Type
locality: Batavia. Holotype: MNHN a-9544.
Plotosus macrophthalmus Bleeker, 1846a: 179. Type locality: Batavia. Syntypes (size and number not stated):
Whereabouts unknown.
Copidoglanis longifilis Macleay, 1881: 207. Type locality: Long Island, Torres Strait, Queensland, Australia. Syntypes: AMS I.16269-001 (2).
Distribution: Indo-Australian Archipelago, New Guinea and Australia; often in coral reef habitats (Allen, 1998).
Paraplotosus butleri Allen, 1998
Paraplotosus butleri Allen, 1998: 129, fig. 3. Type locality: Near Broome, ca. 17º58'S, 122º14'E, Western Australia.
Holotype: WAM P.27368-021.
Distribution: Northern Australia, in coastal reefs (Allen, 1998).
Paraplotosus muelleri (Klunzinger, 1880)
Cnidoglanis mülleri Klunzinger, 1880: 411. Type locality: Port Darwin [Australia]. Holotype: SMNS 2519.
Distribution: Dampier Archipelago to the eastern Gulf of Carpentaria, northern Australia; in turbid coastal reefs(Allen, 1998).

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PLOTOSUS La Cepède, 1803
Plotosus La Cepède, 1803: 129. Type species: Platystacus anguillaris Bloch, 1794. Type by monotypy. Gender:
Masculine.
Deportator Gistel, 1848: X. Type species: Platystacus anguillaris Bloch, 1794. Type by being a replacement name.
Gender: Masculine. Unneeded replacement for Plotosus La Cepède, 1803, which was considered by Gistel to be
preoccupied by Plotus Linnaeus, 1766, in Aves.
Remarks: Plotosis (in Duméril, 1856: 340), Plotoseus (in Lesson, 1826, in Lesson 1826–31: pl. 31, 1831: 435) andPlotosius (in Siebold, 1846: 228) are incorrect subsequent spellings of Plotosus and not available names.
Key: Gomon & Taylor (1982).
Plotosus canius Hamilton, 1822
Plotosus canius Hamilton, 1822: 142, 374, pl. 15 (fig. 44). Type locality: rivers of the southern parts of Bengal. No
types known.
Plotosus unicolor Valenciennes, in Cuvier & Valenciennes, 1840b: 426 (316 in Strasbourg deluxe edition). Type
locality: Java. Holotype: MNHN a-8924.
Plotosus viviparus Bleeker, 1846a: 182. Type locality: Batavia. Syntypes (size and number not stated): possibly
RMNH 8066 (1), RMNH 15875 (6), SMNS 10624 (1).
Plotosus horridus Bleeker, 1846a: 183. Type locality: Batavia. Type(s) (size and number not stated): Whereabouts
unknown.
Plotosus multiradiatus Bleeker, 1846a: 183. Type locality: Batavia. Type (s) (size and number not stated): Whereabouts unknown.
Distribution: Coastal regions of Thailand, Sundaland, Sulawesi, Moluccas, and India, and into lower Mekong River(Kottelat, 2001b).
Remarks: Synonymy based on Bleeker (1862–63: 99). Plotosus caesius in Cuvier (1829, 1836, etc.) and Hyrtl(1859: 5, 6, 17) are considered to be incorrect spellings for P. canius and not the proposal of an available name.
Silurus unitius of Hora (1933: 133) is not available, as it was not treated as a valid species.
Plotosus fisadoha Ng & Sparks, 2002
Plotosus fisadoha Ng & Sparks, 2002: 565, fig. 1. Type locality: Madagascar: southeastern coast of Fanarantsoa
Province; Farafangana market. Holotype: UMMZ 235269.
Distribution: Southeastern Madagascar (Ng & Sparks, 2002).
Plotosus limbatus Valenciennes, 1840
Plotosus limbatus Valenciennes in Cuvier & Valenciennes, 1840b: 422 (313 in Strasbourg deluxe edition). Type
locality: Côte de Malabar ... Pondichéry; [restricted to Malabar by lectotype designation]. Lectotype: MNHN a9546, designated by Gomon & Taylor (1982: 9).
Copidoglanis obscurus Günther, 1864: 26. Type locality: Probably from Australia. Holotype: BMNH
1852.9.13.164.
Distribution: Western Indian Ocean and Arabian Sea, in marine and brackish waters (Taylor & Gomon, 1986).
Plotosus lineatus (Thunberg, 1787)
Silurus lineatus Thunberg, 1787: 31, footnote 13. Type locality: [East Indian Seas]. Type(s): ZMUC P 28555.
Described in more detail and illustated in Thunberg (1791: 191, pl. 6).
Platystacus anguillaris Bloch, 1794: 61, pl. 373 (figs. 1, 2). Type locality: Not stated [given as Tranquebar in Bloch& Schneider, 1801]. Lectotype: ZMB 3078, designated by Taylor & Gomon (1986: 160).
Plotosus thunbergianus La Cepède, 1803: 693, 694. Type locality: La mer des grandes Indes. Holotype: ZMUC P
28555. Unneeded new name for Silurus lineatus Thunberg, 1787.
Plotoseus ikapor Lesson, 1826, in Lesson 1826–31: pl. 31, fig. 3. Type locality: baie d’Offeach de l’ile de Waigiou.
Holotype: Whereabouts unknown. Name made available by figure caption, with illustrated specimen the holotype; described in Lesson (1831: 132).
Plotosus marginatus Bennett, 1830: 691. Type locality: Sumatra. Syntype: BMNH 1855.12.26.452 (1).
Plotosus vittatus Swainson, 1839: 307. Type locality: Not stated [given as Tranquebar in Bloch & Schneider, 1801].
Unneeded new name for Platystacus anguillaris Bloch. Based on “Bl. 373, fig. 1 ” [= Bloch (1794: pl. 373 (fig.
1)].

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FERRARISPlotosus castaneus Valenciennes, in Cuvier & Valenciennes, 1840b: 421 (312 in Strasbourg deluxe edition). Type
locality: Mahé sur la côte malabare. Holotype: MNHN a-8929.
Plotosus lineatus Valenciennes in Cuvier & Valenciennes, 1840b: 412 (306 in the Strasbourg deluxe edition). Type
locality: la mer des Indes ... la mer Rouge ... la Séchelles ... la côte de Malabar, Trincomalé, Pondichéry,
Amboine ... Célèbes, îles des Amis, îles de la Société, Macao ... Philippines. Syntypes:MNHN a-8930 (1),
MNHN a-8931 (2), MNHN a-8932 (10 or 11), MNHN a-8936 (1). Preoccupied in Plotosus by Silurus lineatus
Thunberg, 1787.
Plotosus castaneoïdes Bleeker, 1851f: 490. Type locality: Rio [=Riouw], in mari. Syntypes (2, 160–170 mm TL): at
RMNH.
Plotosus arab Bleeker, 1862a: 111. Type locality: Not stated [given as Tranquebar in Bloch & Schneider, 1801].
Syntypes: at MNHN, ZMB 3078 (1), ZMB 3079 (1). Name made available by reference to Plotosus anguillarisof La Cepède (1803: 130, pl. 3 fig. 2), which is based on specimens and Platystacus anguillaris Bloch, 1794.
Plotosus brevibarbus Bessednov, 1967: 446, fig. 3. Type locality: 21º10' с. ш, 108º30'в. д, 29 m [Gulf of Tonkin,South China Sea]. Holotype: Mus. TINRO 3667.
Distribution: Indian Ocean and Western Pacific, in marine waters, but sometimes entering freshwaters in East Africa
and Madagascar (Taylor & Gomon, 1986); eastern Mediterranean (Golani, 2002).
Remarks: The name Silurus Arab in Forsskål (1775: xvi) is not available (M. Kottelat, pers. commun.). The namePlotosus malignus appears in the account of Plotosus lineatus Valenciennes, but it not treated as valid and istherefore not available from that publication. The name does not appear to have been treated as valid subsequently. See Smith (1941) for nomenclatural comments regarding this species.
Plotosus nkunga Gomon & Taylor, 1982
Plotosus nkunga Gomon & Taylor, 1982: 3, pl. 1. Type locality: South Africa, Transkei, Bashee River (32º15'S,
28º55'E). Holotype: SAM 18457.
Distribution: Southern Africa from Boknes to Boteler Point, and possibly to Zanzibar, in marine waters but also
entering freshwater (Taylor & Gomon, 1986).
Plotosus papuensis Weber, 1910
Plotosus papuensis Weber, 1910: 228. Type locality: Lorentz-Fluss [New Guinea]. Syntypes: FMNH 52494 (1),
ZMA 111093–97 (5), ZMA 111098–100 (5), ZMA 111513 (1).
Distribution: Southern Papua (formerly Irian Jaya), New Guinea (Allen, 1996).
Species inquirenda, Plotosus
Plotosus abbreviatus Boulenger, 1895a: 247. Type locality: Mouth of Baram River, Sarawak. Holotype: BMNH
1894.8.3.35.
POROCHILUS Weber, 1913
Porochilus Weber, 1913: 523. Type species: Porochilus obbesi Weber, 1913. Type by monotypy. Gender: Masculine.
Porochilus meraukensis (Weber, 1913)
Copidoglanis meraukensis Weber, 1913: 529. Type locality: Merauke, Süd-Neu Guinea. Syntypes: RMNH 10865(1), ZMA 111092 (1).
Distribution: Southern Papua (formerly Irian Jaya), New Guinea (Allen, 1996).
Porochilus obbesi Weber, 1913
Porochilus Obbesi Weber, 1913: 523, figs. 4–5. Type locality: Lorentz-Fluss, ... bei Sabang, New Guinea. Syntypes(6): ZMA 111101 (3), ZMA 111102 (1), RMNH 10864 (1).
Distribution: Southern Papua (formerly Irian Jaya), New Guinea (Allen, 1996).
TANDANUS Mitchell, 1838
Tandanus Mitchell, 1838: 95. Type species: Plotosus (Tandanus) tandanus Mitchell, 1838. Type by monotypy. Gender: Masculine. Originally proposed as a subgenus of Plotosus.
Copidoglanis Günther, 1864: 25. Type species: Plotosus (Tandanus) tandanus Mitchell, 1838. Type by subsequent

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designation by Jordan (1919b: 332) with name was misprinted Copiodoglanis. Gender: Masculine.
Tandanus bostocki Whitley, 1944
Plotosus unicolor Castelnau, 1873: 141. Type locality: Interior of Western Australia. Syntype: NMV 51851 (1). Preoccupied by Plotosus unicolor Valenciennes, 1840, replaced by Tandanus bostocki Whitley, 1944.
Tandanus bostocki Whitley, 1944: 260. Type locality: Interior of Western Australia. Syntype: NMV 51851 (1).
Replacement for Plotosus unicolor Castelnau, 1873, preoccupied by Plotosus unicolor Valenciennes, 1840.
Remarks: The statement in Whitley (1944: 260) that designates a holotype for the species is in error, since he clearlystates elsewhere that Tandanus bostocki is intended to replace the preoccupied name of Castelnau and, thereby,
takes the same specimen as holotype.
Distribution: Coastal drainages of southwestern Australia, from the Frankland River to Moore River (Allen, 1989).
Tandanus tandanus (Mitchell, 1838)
Plotosus (Tandanus) tandanus Mitchell, 1838: 44, 95, pl. 5 (fig. 2). Type locality: Lagoon near Tangulda, Namoi
River, New South Wales; river between Gwydir River and MacIntyre River, New South Wales. No types
known.
Distribution: Murray-Darling River system and coastal drainages of eastern Australia (Allen, 1989).
Remarks: See Musyl & Keenan (1996) for comments on populational variation that may represent unrecognized
additional species.
Species inquirenda, Plotosidae
Plotosus nigricans Valenciennes, in Cuvier & Valenciennes, 1840b: 412, 417 (310 in Strasbourg deluxe edition). Type
locality: Not stated. Holotype: at MNHN (skeleton). Species name mentioned in generic account as well as accountof Plotosus lineatus. Reference made to published comments about the skeleton by Cuvier in Regne Animal, which
are adequate to act as an indication, even though Valenciennes never provides a more complete treatment of thespecies. Gomon & Taylor (1982: 10), disagree and treated the name as a nomen nudum, but later argue that a
specimen previously treated as the holotype of the species, MNHN a-9433, must not be considered as one of the“original type-specimens.”

PSEUDOPIMELODIDAE Lundberg, Bornbusch & Mago-Leccia, 1991
Pseudopimelodinae Lundberg, Bornbusch & Mago-Leccia, 1991: 204. Type genus: Pseudopimelodus Bleeker, 1858.
Remarks: The name Pseudopimelodidae was proposed first by Fernández-Yépez & Martín Salazar (1953) in a mannerthat made the name available when published, but was rendered unavailable in Art. 13 of the Second Edition of theCode (ICZN, 1961). The name apparently remains unavailable, despite Art. 13.2.1 of the current version of the Code(ICZN, 1999), because the name was treated as unavailable in Ferraris & de Pinna (1999: 8).
Taxonomic summary: Shibatta (2003a).
Phylogeny: Shibatta (1998, 2003b).
5 genera, 29 species; no named fossil taxa.
BATROCHOGLANIS Gill, 1858
Batrochoglanis Gill, 1858: 389. Type species: Pimelodus raninus Valenciennes, 1840. Type by original designation.
Gender: Masculine.
Remarks: Batrachoglanis, first used in Gill (1861c), is either a misspelling or unjustified emendation of Batrochoglanis Gill, 1858.
Batrochoglanis acanthochiroides (Güntert, 1942)
Pseudopimelodus acanthochiroides Güntert, 1942: 29. Type locality: Santander, Colombia. Syntypes: NMBA 5277–
78 (2).
Pseudopimelodus villosus butcheri Schultz, 1944c: 199, pl. 1 (fig. b). Type locality: Río San Juan near bridge southof Mene Grande, tributary to Rio Motatán, Maracaibo basin, Venezuela. Holotype: USNM 121270.
Distribution: Catatumbo River basin in Lake Maracaibo drainage (Shibatta, 2003a).

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Batrochoglanis melanurus Shibatta & Pavanelli, 2005
Batrochoglanis melanurus Shibatta & Pavanelli, 2005: 23, figs. 1–2. Type locality: Córrego Cancela, affluent of rio
Cuiabá, rio Paraguai basin; 14º42.501S/ 56º15.850W; Nobres; State of Mato Grosso; Brazil. Holotype: MZUSP
87240.
Distribution: Cuiabá River, Paraguay River basin, Mato Grosso. Brazil (Shibatta & Pavanelli, 2005).
Batrochoglanis raninus (Valenciennes, 1840)
Pimelodus raninus Valenciennes, in Cuvier & Valenciennes, 1840b: 157 (117 in the Strasbourg deluxe edition), pl.
434. Type locality: La Mana ... Rio-Janéiro. Syntypes: MNHN a-9942 (3).
Pseudopimelodus acanthochirus Eigenmann & Eigenmann, 1888b: 122. Type locality: Brazil. Syntypes (5): MCZ
8133 (1), MCZ 8157 (1), MCZ 7732 (2), MCZ 7332 (1).
Distribution: Amazon River basin (Shibatta, 2003a).
Batrochoglanis transmontanus (Regan, 1913)
Pseudopimelodus transmontanus Regan, 1913d: 467. Type locality: Condoto ..., San Juan and the Tamana ...., andthe Durango, W. Ecuador. Syntypes (9): BMNH 1910.7.11.104, BMNH 1910.7.11.105, BMNH 1913.10.1.36,
BMNH 1902.5.27.37–40.
Distribution: Baudó, San Juan, Patia and Durango River basins, Pacific slope drainages (Shibatta, 2003a).
Batrochoglanis villosus (Eigenmann, 1912)
Pseudopimelodus villosus Eigenmann, 1912b: 152, fig. 32, pl. 10 (fig. 1). Type locality: Potaro Landing, British
Guiana. Holotype: FMNH 53219.
Distribution: Demerara, Essequibo, Orinoco and Amazon River basins (Shibatta, 2003a).
CEPHALOSILURUS Haseman, 1911
Cephalosilurus Haseman, 1911a: 317. Type species: Cephalosilurus fowleri Haseman, 1911. Type by monotypy.
Gender: Masculine.
Cephalosilurus albomarginatus (Eigenmann, 1912)
Pseudopimelodus albomarginatus Eigenmann, 1912b: 153, pl. 11 (fig. 1). Type locality: Tukeit, British Guiana.
Holotype: FMNH 53221.
Distribution: Tukeit, Guyana (Shibatta, 2003a).
Cephalosilurus apurensis (Mees, 1978)
Pseudopimelodus apurensis Mees, 1978b: 253, pl. 1. Type locality: Río Arichuna, near San Pedro, Rincón hondo,
Apure, Venezuela. Holotype: RMNH 27644.
Distribution: Arichuna River basin, Apure State, Venezuela (Shibatta, 2003a).
Cephalosilurus fowleri Haseman, 1911
Cephalosilurus fowleri Haseman, 1911a: 317, pl. 46. Type locality: Rio São Francisco, Cidade de Barra, Bahia, Brazil. Holotype: FMNH 54254.
Distribution: São Francisco River basin, Brazil (Shibatta, 2003a).
Cephalosilurus nigricaudus (Mees, 1974)
Pseudopimelodus nigricauda Mees, 1974: 218, figs. 37d, 38d; pls. 11–12. Type locality: Sipaliwini, Suriname.
Holotype: RMNH 26739.
Distribution: Sipaliwini River basin, Suriname (Shibatta, 2003a).
LOPHIOSILURUS Steindachner, 1877
Lophiosilurus Steindachner, 1877a: 154. Type species: Lophiosilurus alexandri Steindachner, 1877. Type by monotypy. Gender: Masculine.
Lophiosilurus alexandri Steindachner, 1877
Lophiosilurus Alexandri Steindachner, 1877a: 154, pl. 15. Type locality: Aus Brasilien, wahrscheinlich aus dem
Amazonenstrome. Holotype: NMW 46123.
Pseudopimelodus Agassizi Steindachner, 1880b: 61. Type locality: Aus Brasilien, wahrscheinlich aus dem Amazonenstrome. Holotype: NMW 46123. Unneeded new name, or lapsus calami for Lophiosilurus alexandri Steindachner, 1877, originally written as Pseudop. Agassizi.

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Distribution: São Francisco River basin, Brazil (Shibatta, 2003a).
Remarks: See Pinto & Marzulo (1975) for comments on morphology.
MICROGLANIS Eigenmann, 1912
Microglanis Eigenmann, 1912b: 130, 155. Type species: Microglanis poecilus Eigenmann, 1912. Type by original
designation. Gender: Masculine.
Key: Bertaco & Cardoso (2005), and Shibatta & Benine (2005), southern Brazil.
Microglanis ater Ahl, 1936
Microglanis ater Ahl, 1936: 109. Type locality: Brazil. Holotype: ZMB 20932.
Distribution: Brazil (Shibatta, 2003a).
Microglanis cibelae Malabarba & Mahler, 1998
Microglanis cibelae Malabarba & Mahler, 1998: 249, figs. 1, 4–6. Type locality: arroio do Ouro, Rio Maquiné,
between Maquiné and Barra do Ouro, Rio Grande do Sul, Brasil. Holotype: MZUSP 48653.
Distribution: Coastal drainages in northern Rio Grande do Sul and Santa Catarina States, Brazil (Shibatta, 2003a;
Bertaco & Cardoso, 2005).
Microglanis cottoides (Boulenger, 1891)
Pimelodus (Pseudopimelodus) cottoides Boulenger, 1891: 233, pl. 25 (fig. 2). Type locality: Rio Camaquã, Rio
Grande do Sul, Brazil. Syntypes: BMNH 1891.3.16.36–45 and BMNH 1891.3.16.106 (lots combined, 12 specimens) and BMNH 1889.8.24.5 (1).
Distribution: Laguna dos Patos and Uruguay River basins, Brazil (Shibatta, 2003a; Bertaco & Cardoso, 2005).
Microglanis eurystoma Malabarba & Mahler, 1998
Microglanis eurystoma Malabarba & Mahler, 1998: 248, fig. 3. Type locality: Rio Uruguay, Itá, Santa Catarina, Brazil. Holotype: MCP 13405.
Distribution: Upper Uruguay River basin, Brazil (Shibatta, 2003a; Bertaco & Cardoso, 2005).
Microglanis garavelloi Shibatta & Benine, 2005
Microglanis garavelloi Shibatta & Benine, 2005: 580, figs. 1–2. Type locality: Brazil, Paraná: Taquari stream,
23º12'24"S 50º56'50"W, Município de Jataizinho. Holotype: MZUSP 88006.
Distribution: Paranapanema and Tietê River basins, Brazil (Shibatta & Benine, 2005).
Microglanis iheringi Gomes, 1946
Microglanis iheringi Gomes, 1946: 9, pl. 1. Type locality: Río Turmero, Aragua, Venezuela. Holotype: FMNH
35350.
Distribution: Turmero River basin, Venezuela (Shibatta, 2003a).
Microglanis malabarbai Bertaco & Cardoso, 2005
Microglanis malabarbai Bertaco & Cardoso, 2005: 62, fig. 2. Type locality: arroio Albino, tributary of rio Ijuí (rio
Uruguay drainage), 28º08'10"S, 54º55'28"W, São Pedro do Butiá, Rio Grande do Sul, Brazil. Holotype: MCP
35941.
Distribution: Middle Uruguay River basin, Brazil (Bertaco & Cardoso, 2005).
Microglanis nigripinnis Bizerril & Perez-Neto, 1992
Microglanis nigripinnis Bizerril & Perez-Neto, 1992: 97, figs. 1–2. Type locality: Brazil, Estado do Rio de Janeiro,
rio Macacu, Município de Cachoeira do Macacu, Cidade de Papucaia. Holotype: MZUSP 42308.
Distribution: Macacu River basin in Rio de Janeiro State, Brazil (Shibatta, 2003a).
Microglanis parahybae (Steindachner, 1880)
Pseudopimelodus Parahybae Steindachner, 1880b: 60 footnote, pl. 1 (figs. 2–2b). Type locality: Rio Parahyba undvon Santa Cruz [Brazil]. Syntypes: MCZ 8161 (25), MCZ 8162 (6), NMW 44433 (9), NMW44436 (1).
Distribution: Paraíba do Sul River basin, Brazil (Shibatta, 2003a).
Microglanis pellopterygius Mees, 1978
Microglanis pellopterygius Mees, 1978b: 256, pl. 2. Type locality: Tributary stream of Río Aguarico at Santa
Cecilia, 0º06'N, 76º51'W, Napo, Ecuador. Holotype: ANSP 130437.
Distribution: Aguarico River basin, Ecuador (Shibatta, 2003a).
Microglanis poecilus Eigenmann, 1912

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Microglanis poecilus Eigenmann, 1912b: 155, pl. 12 (fig. 2). Type locality: Below Packeoo Falls, British Guiana.
Holotype: FMNH 46365.
Distribution: Essequibo River basin and rivers of French Guiana (Shibatta, 2003a).
Microglanis secundus Mees, 1974
Microglanis secundus Mees, 1974: 235, fig. 40 e; pl. 14. Type locality: Sipaliwini, Suriname. Holotype: RMNH
26525.
Distribution: Catatumbo River basin; Venezuela and Colombia; Sipaliwini River basin, Suriname (Shibatta, 2003a).
Microglanis variegatus Eigenmann & Henn, 1914
Microglanis variegatus Eigenmann & Henn, in Eigenmann, Henn & Wilson, 1914: 14. Type locality: Near Vinces,
Ecuador. Holotype: CAS 17971; holotype illustrated in Eigenmann (1922b: 33, pl. 2, figs. 3–4).
Distribution: Forest pool near Vinces, Ecuador (Shibatta, 2003a).
Microglanis zonatus Eigenmann & Allen, 1942
Microglanis zonatus Eigenmann & Allen, 1942: 89, pl. 3 (figs. 1–2). Type locality: Rio Morona (?). Holotype: CAS
17970.
Distribution: Upper Amazon River basin, Peru (Shibatta, 2003a).
PSEUDOPIMELODUS Bleeker, 1858
Pseudopimelodus Bleeker, 1858b: 196, 204, 207. Type species: Pimelodus bufonius Valenciennes, 1840. Type by
subsequent designation by Gill (1861c). Gender: Masculine.
Pseudopimelodus bufonius (Valenciennes, 1840)
Pimelodus Bufonius Valenciennes, in Cuvier & Valenciennes, 1840b: 154 (115 in the Strasbourg deluxe edition).
Type locality: Cayenne. Holotype: RMNH (lost).
Distribution: Rivers of northeastern South America from Lake Maracaibo basin to eastern Brazil (Shibatta, 2003a)
and Magdalena and Cauca River basins, Colombia (Maldonado-Ocampo et al., 2005).
Pseudopimelodus charus (Valenciennes, 1840)
Pimelodus charus Valenciennes, in Cuvier & Valenciennes, 1840b: 159 (118 in the Strasbourg deluxe edition). Type
locality: Rio Sabara [Brazil]. No types known; description based on an unpublished illustration, reproduced in
Mees (1974, frontispiece).
Distribution: São Francisco River basin, Brazil (Shibatta, 2003a).
Pseudopimelodus mangurus (Valenciennes, 1835)
Pimelodus mangurus Valenciennes, 1835, in Valenciennes, 1835–47: pl. 1 (figs. 4–6). Type locality: Río de la Plata.
Possible holotype: MNHN a-8401(1), MNHN a-9417 (1). Name available from caption on plate, so illustrated
specimen is the holotype, if identifiable. Described in Cuvier & Valenciennes (1840b: 156 (116 of Strasbourg
deluxe edition)), with locality given as Rio-Janéiro.
Pseudopimelodus roosevelti Borodin, 1927b: 1, fig. 1. Type locality: Parassununga, Estado São Paulo, Brazil. Holotype: AMNH 8638.
Distribution: Uruguay, Paraná, Paraguay and La Plata River basins (Shibatta, 2003a).
Pseudopimelodus pulcher (Boulenger, 1887)
Pimelodus (Pseudopimelodus) pulcher Boulenger, 1887c: 276, pl. 21 (fig. 1). Type locality: Canelos, e. Ecuador.
Syntypes: BMNH 1880.12.8.105–107 (3).
Pseudopimelodus variolosus Miranda Ribeiro, 1914: 4, pl. 1 (fig. 2), pl. 2 (figs. 1–2). Type locality: Coxim, Rio
Taquary, M. Grosso [Brazil]. Lectotype: MNRJ 818A, designated by Miranda Ribeiro (1953: 404).
Distribution: Upper Amazon River basin (Shibatta, 2003a).
Pseudopimelodus schultzi (Dahl, 1955)
Zungaro zungaro schultzi Dahl, 1955: 13. Type locality: Cereté, Colombia. Holotype: Whereabouts unknown (possibly at ICNMHN).
Distribution: Magdalena River basin, Colombia (Shibatta, 2003a).
Species inquirenda, Pseudopimelodidae

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Zungaro mathisoni Fernández-Yépez, 1972a: 22, pl. 41. Type locality: Río Yaracuy drainage, Venezuela. Holotype:author’s personal collection; whereabouts unknown.

SCHILBIDAE Bleeker, 1858
Ailichthyoidei Bleeker, 1858b: ix, 49, 248. Type genus: Ailia Gray, 1830.
Schilbeini Bleeker, 1858b: 49, 250, 256. Type genus: Schilbe Oken, 1817.
Siluranodontinae Regan, 1911: 568. Type genus: Siluranodon Bleeker, 1858.
Taxonomic summaries: De Vos (1984b, 1986, Africa).
Reviews: Reizer et al. (1980, Senegal), De Vos (1995, Africa); De Vos (1992, West Africa).
Identification guide: Jayaram (1977, South Asia).
Key to genera: Trewavas (1943, Africa); Poll & Gosse (1994, Africa).
Remarks: Although the family group name is often spelled Schilbeidae, the spelling used here, Schilbidae, was considered to be correct by Steyskal (1980: 174) and is followed herein. The monophyly of this family, as currently populated, is unlikely. Species of the Pangasiidae and possibly Horabagrus may be nested within the family, and thespecies of Neotropius and Pseudeutropius may be more closely related to some species of the Bagridae. See Mo(1991), Pouyaud et al. (2000, 2004), Diogo et al. (2004) for details. Hardmann (2005) further suggested that African
species of the Schilbidae may be more closely related to other African catfishes than to any Asian member of the
Schilbidae, in which case the name Ailiidae would be the valid name for the Asian group.
14 genera, 62 species; no named fossil taxa.
AILIA Gray, 1830
Acanthonotus Gray, 1830: pl. 85 (fig. 1). Type species: Silurus (Acanthonotus) cuvieri Gray, 1830. Type by monotypy. Gender: Masculine. Name made available by figure caption. Originally proposed as a subgenus of Silurus.
Subsequently published in Gray (1831: 8), but with Acanthonotus Hardwickii as the only included species. Preoccupied by Acanthonotus Goldfuss, 1809 (and possibly earlier), in Mammalia.
Ailia Gray, 1830: pl. 85 (fig. 2). Type species: Malapterus (Ailia) bengalensis Gray, 1830. Type by monotypy. Gender: Feminine. Originally proposed as a subgenus of Malapterus. Name made available by caption of plate; text
published later in Gray (1831: 8).
Ailiichthys Day, 1872: 712. Type species: Ailiichthys punctata Day, 1872. Type by monotypy. Gender: Masculine.
Review: Hora (1941); Jayaram (1963).
Ailia coila (Hamilton, 1822)
Malapterurus coila Hamilton, 1822: 158, 375. Type locality: Fresh water rivers of Bengal. No types known. Unpublished Hamilton illustration reproduced in Gray (1830: pl. 85, fig. 2), as Malapterus (Ailia) bengalensis.
Malapterus (Ailia) bengalensis Gray, 1830: pl. 85 (fig. 2). Type locality: India. No types known; name based on previously-unpublished illustration by Hamilton. Name made available by figure caption; brief diagnosis publishedin Gray (1831: 8), with locality stated as “Rivers Bengal.”
Silurus (Acanthonotus) Cuvieri Gray, 1830: pl. 85 (fig. 1). Type locality: India. No types known. Name made available by plate legend; text published later in Gray (1831: 8), as Acanthonotus Hardwickii.
Acanthonotus Hardwickii Gray, 1831: 8 [pl. 85 (fig. 1), of Gray, 1830]. Type locality: India. No types known.
Unneeded new name for Silurus (Acanthonotus) Cuvieri Gray, 1830.
Ailia affinis Günther, 1864: 56. Type locality: Kulu and Assam, India. Syntypes (3, in two lots): BMNH
1860.3.19.867–868 (2), BMNH 2005.5.17.3 (1).
Distribution: Rivers of northern India and Deccan, including Indus, Ganges, Yamuna, Brahmaputra, Mahanadi andKrishna Rivers (Jayaram, 1963).
Ailia punctata (Day, 1872)
Ailiichthys punctata Day, 1872: 713. Type locality: Jumna at and below Delhi, also in the lower Punjaub rivers. Syntypes: AMS B.7570 (1), BMNH 1889.2.1.2585–87 (3), BMNH 1889.2.1.2588–90 (3), BMNH 1889.2.1.2591(1, skeleton), BMNH 1889.2.1.2592–93 (2), NMW 47849 (1), NMW 47863 (1), RMNH 2763 (1), ZMB 11213(1), ZSI 420 (1, lost).

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FERRARISDistribution: Indus, Yamuna and Ganges Rivers, northern India (Jayaram, 1963).
CLUPISOMA Swainson, 1838
Clupisoma Swainson, 1838: 347, 351, 354. Type species: Silurus (Clupisoma) argentata Swainson, 1839. Type by
subsequent monotypy. Gender: Neuter. Originally proposed as a subgenus of Silurus, without any included species; first species added in Swainson (1839: 306).
Schilbeichthys Bleeker, 1858b: 255, 256. Type species: Silurus garua Hamilton, 1822. Type by monotypy. Gender:
Masculine.
Key to species: Datta & Karmakar (1980).
Clupisoma bastari Datta & Karmakar, 1980
Clupisoma bastari Datta & Karmakar, 1980: 193, figs. 1–2. Type locality: Indravati River (a tributary of GodavariRiver) at Lohandigura, 33 kms west of Jagdalpur, Bastar District, Madhya Pradesh, India. Holotype: ZSI
FF1342.
Distribution: Godavari River basin, Madhya Pradesh, India (Jayaram, 1999).
Clupisoma garua (Hamilton, 1822)
Silurus garua Hamilton, 1822: 156, 375, pl. 21 (fig. 50). Type locality: Fresh water rivers of the Gangetic provinces.
No types known.
Silurus (Clupisoma) argentata Swainson, 1839: 306. Type locality: Fresh water rivers of the Gangetic provinces. No
types known. Made available by reference to “Ham. pl. 21, f. 50” [= Hamilton, 1822: pl. 21 (fig. 50)]. Unneeded
new name for Silurus garua Hamilton, 1822.
Distribution: Throughout northern India, but not south of the Mahanadi River, Pakistan, Bangladesh, and Nepal(Jayaram, 1999).
Clupisoma longianalis (Huang, 1981)
Platytropius longianalis Huang, 1981: 438, fig. 5. Type locality: Puer county, Xiaoganlanba, Yunnan, China. Holotype: KIZ 735118.
Distribution: Lancangjiang [= Mekong River] (Chu et al., 1999).
Clupisoma montana Hora, 1937
Clupisoma montana Hora, 1937b: 673, figs. 7–8. Type locality: Teesta River, below Darjeeling, India. Type(s): ZSIF 12472/1.
? Pseudeutropius murius batarensis Shrestha, 1980: 212, fig. 97. Type locality: Batar, Trisuli river, Nuwakot Dist.,
Bagmati Zone, Nepal. Holotype (220 mm): at Museum of Zoology Department, Institute of Science, Tribuhaven University, Nepal.
Distribution: Teesta, Jamuna and Sone Rivers, India, Nepal (Jayaram, 1999).
Clupisoma naziri Mirza & Awan, 1973
Clupisoma naziri Mirza & Awan, 1973: 152, fig. 2. Type locality: Indus River at Jinnah Barrage, Pakistan. Holotype: GCM 10.
Distribution: NWFP, Punjab, Azad Kashmir, Indus River basin, Pakistan (Mirza, 2003).
Clupisoma nujiangense Chen, Ferraris & Yang, 2005
Clupisoma nujiangense Chen, Ferraris & Yang, 2005: 566, fig. 1. Type locality: China, Yunnan Province, Baoshan
City, Longling County, Mungnuo Township, Sanjiangkou, Nu Jiang (=Salween River), 24º25'18.6"N,
98º58'21'2"E. Holotype: KIZ 200310118.
Distribution: Upper Salween River, Yunnan, China (Chen et al., 2005).
Clupisoma prateri Hora, 1937
Clupisoma prateri Hora, 1937b: 671, fig. 6. Type locality: Burma. Holotype: Asiatic Soc. Bengal 213 or 219 [now at
ZSI].
Distribution: Lower and middle reaches of Irrawaddy River, Sittang, Bago and Salween river basins, Myanmar (Ferraris, 2004).
Remarks: Redescribed in Ferraris (2004: 6, fig. 2).
Clupisoma roosae Ferraris, 2004
Clupisoma roosae Ferraris, 2004: 2, fig. 1. Type locality: Myanmar, Kachin State, Myitkyina market. Holotype:

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NRM 40030.
Distribution: Upper Irrawaddy River basin, Myanmar (Ferraris, 2004).
Clupisoma sinense (Huang, 1981)
Platytropius sinensis Huang, 1981: 437, figs. 1–4. Type locality: Puer county, Xiaoganlanba, Yunnan, China. Holotype: KIZ 735124.
Distribution: Mekong River basin (Rainboth, 1996), including Lancangjiang (= upper Mekong River of Yunnan)(Chu et al., 1999), and Malay Peninsula (Zakaria-Ismail, 1992; Kottelat, 2001b).
Remarks: Placement of this species in Clupisoma follows Ng (1999b) and Chen et al. (2005).
EUTROPIICHTHYS Bleeker, 1862
Eutropiichthys Bleeker, 1862b: 398. Type species: Pimelodus vacha Hamilton, 1822. Type by original designation.
Gender: Masculine. Also proposed as new in Bleeker (1862–63: 14).
Revision: Hora (1937e).
Eutropiichthys burmannicus Day, 1877
Eutropiichthys burmannicus Day, 1877 (in Day, 1875–78): 490. Type locality: Burma. Possible syntypes: ZSI B.38–
39 (2), ZSI B.219 (1, lost), ZSI B.306 (1). Originally as: “Variety Eutropiichthys burmannicus,” in the accountof Eutropiichthys vacha.
Distribution: Irrawaddy, lower Salween and smaller intervening river basins.
Eutropiichthys murius (Hamilton, 1822)
Pimelodus murius Hamilton, 1822: 195, 378. Type locality: Mahananda river [Bengal]. No types known. Unpublished Hamilton illustration reproduced in Hora (1929: pl. 22, fig. 2).
Pachypterus melanurus Swainson, 1839: 306. Type locality: Mahananda river [Bengal]. No types known. Unneeded
new name for Pimelodus murius Hamilton, 1822.
Distribution: Northern India, Bangladesh and Pakistan (Jayaram, 1999).
Eutropiichthys vacha (Hamilton, 1822)
Pimelodus vacha Hamilton, 1822: 196, 378, pl. 19 (fig. 64). Type locality: Larger fresh water rivers of the Gangetic
provinces. No types known.
Pachypterus punctatus Swainson, 1839: 306. Type locality: Larger fresh water rivers of the Gangetic provinces. No
types known. Made available by reference to “Ham. p. 196, f. 64 ” [= Hamilton, 1822: 196, p. 19, fig. 64)].
Unneeded new name for Pimelodus vacha Hamilton, 1822.
Distribution: Northern India, south to the Mahanadi River, Bangladesh, Nepal and Pakistan (Jayaram, 1999).
IRVINEIA Trewavas, 1943
Irvineia Trewavas, 1943: 165. Type species: Irvineia voltae Trewavas, 1943. Type by monotypy. Gender: Feminine.
Irvineia orientalis Trewavas, 1964
Irvineia orientalis Trewavas, 1964: 390, figs. 1, 2, 3 b–c. Type locality: Villaggio Duca delgi Abruzzi, Uebi Scebeli(Shebeli) system. Holotype: MSNG 39551; holotype illustrated in De Vos (1995: 358, fig. 226).
Distribution: Juba-Uebi Shebeli system (De Vos, 1995).
Irvineia voltae Trewavas, 1943
Irvineia voltae Trewavas, 1943: 165, fig. 1. Type locality: Above the Senchi Rapids, River Volta, Gold Coast. Syntypes (3): BMNH 1944.2.9.19–21 (3); one syntype illustrated in De Vos (1995: 362, fig. 229).
Distribution: lower Volta River basin (De Vos, 1995).
LAIDES Jordan, 1919Laïs Bleeker, 1857b: 473. Type species: Pangasius hexanema Bleeker, 1852. Type by monotypy. Gender: Masculine. Preoccupied by Lais Gistel, 1848, in Tunicata, replaced by Laides Jordan, 1919.
Laides Jordan, 1919b: 293. Type species: Pangasius hexanema Bleeker, 1852. Type by being a replacement name.
Gender: Masculine. Replacement for Lais Bleeker, 1857; preoccupied by Lais Gistel, 1848, in Tunicata.
Laides hexanema (Bleeker, 1852)
Pangasius hexanema Bleeker, 1852d: 588. Type locality: Palembang, Batavia. Syntypes (2, 102–152 mm TL): pos-

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FERRARIS

sibly BMNH 1863.12.4.1007 (1, 157 mm TL), NMV A928 (1).
Distribution: Thailand to Indonesia, in large rivers, including the Mekong (Rainboth, 1996).
Laides longibarbis (Fowler, 1934)
Pangasius longibarbis Fowler, 1934a: 87, fig. 27. Type locality: Me Kong at Chieng Sen, North Siam. Holotype:
ANSP 59441.
Distribution: Mekong, Mehklong, and Chao Phraya River basins (Ng, 1999b).
Remarks: Redescribed by Ng (1999b).
NEOTROPIUS Kulkarni, 1952
Pachypterus Swainson, 1838: 346 et seq. Type species: Silurus atherinoides Bloch, 1794. Type by subsequent designation by Swain (1882: 281). Gender: Masculine. Originally without any included species. Species first added
in Swainson (1839: 306).
Neotropius Kulkarni, 1952: 231. Type species: Neotropius khavalchor Kulkarni, 1952. Type by monotypy. Gender:
Masculine.
Review: Hora (1941).
Remarks: Swain (1882: 281) and (following Swain) Jordan (1919a: 205) treated Pachypterus Swainson as preoccupied in Coleoptera, presumably basing their decision on the following notation in Agassiz (1846: 271): “Pachypterus Sol. Col., 1833.” However, it appears that Agassiz was referring to usage of the name in Dejean, 1834,
in which a manuscript by Solier was cited as the source of Pachypterus. According to Neave (1940: 515)
Dejean’s use of the name was as a nomen nudum and Pachypterus was not used in a way that would have madeit available in Coleoptera until Lucas, 1846, which postdates the first proposal of Pachypterus. However, thewritings of Swain and Jordan may have stifled use of Pachypterus Swainson as valid, and the younger name
Neotropius, which has been in widespread usage for the past one-half century, is used here. The species listed
below are similar overall to those of Pseudeutropius and the two genera may not both represent natural groups.
Neotropius acutirostris (Day, 1870)
Pseudeutropius acutirostris Day, 1870c: 618. Type locality: Throughout Burma. Syntypes: AMS B.7967 (1), BMNH
1889.2.1.2413 (1), BMNH 1889.2.1.2462 (1), NMW 48327 (1), ZSI F484 (1), ZSI A.505 (1 lost).
Distribution: Irrawaddy, Sittang, and Bago Rivers, Myanmar.
Neotropius atherinoides (Bloch, 1794)
Silurus Atherinoides Bloch, 1794: 48, pl. 371 (fig. 1). Type locality: Tranquebar. Holotype: ZMB 3013.
Pimelodus angius Hamilton, 1822: 180, 377, pl. 29 (fig. 59). Type locality: Rivers of Bengal. No types known.
Pimelodus urua Hamilton, 1822: 177, 377. Type locality: Rivers and ponds of the northern parts of Bengal. No types
known. Unpublished Hamilton illustration reproduced in Hora (1929: pl. 21, fig. 4).
Pachypterus trifasciatus Swainson, 1839: 306. Type locality: Rivers of Bengal. No types known. Name made available by reference to “Ham. P. 180, f. 59 ” [= Hamilton (1822: 180, pl. 29, fig. 59)]. Unneeded new name for
Pimelodus angius Hamilton, 1822.
Pseudeutropius atherinoides walkeri Chaudhuri, 1912: 444, pl. 41 (fig. 3). Type locality: Siripur, Saran, Bihar[North India]. Syntypes (2): at ZSI. Originally as Pseudeutropius atherinoides var. walkeri.
Distribution: Widely distributed in India, except for Assam and Kerala; also in Pakistan, Nepal, and Bangladesh(Jayaram, 1999).
Neotropius khavalchor Kulkarni, 1952
Neotropius khavalchor Kulkarni, 1952: 232, fig. 1. Type locality: Panchaganga River, near Kolhapur, Bombay State,
India. Holotype: ZSI F647/2.
Distribution: Krishna River basin, Maharashtra and Andhra Pradesh, India (Jayaram, 1999).
Species inquirenda, Neotropius
Bagrus Exodon Cuvier & Valenciennes, in Valenciennes, 1832b: 385, pl. 4 (fig. 1). Type locality: Bengal. Syntypes:
MNHN a-8960 (2), MNHN b-0680 (1). Also in Cuvier & Valenciennes (1840a: 395).

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PARAILIA Boulenger, 1899
Parailia Boulenger, 1899b: 105. Type species: Parailia congica Boulenger, 1899. Type by monotypy. Gender: Feminine.
Physailia Boulenger, 1901e: 445. Type species: Physailia pellucida Boulenger, 1901. Type by monotypy. Gender:
Feminine.
Parailia congica Boulenger, 1899
Parailia congica Boulenger, 1899b: 106, pl. 41 (fig. 2). Type locality: Ebinga (rivière du lac Léopold II). Lectotype:
MRAC 942, designated by De Vos (1995: 297).
Parailia longifilis Boulenger, 1902d: 37, pl. 10 (fig. 3). Type locality: l’Ubangi à Banzyville. Lectotype: MRAC
1309 (1), designated by De Vos (1995: 295).
Distribution: Congo River basin (De Vos, 1995).
Parailia occidentalis (Pellegrin, 1901)
Ailia occidentalis Pellegrin, 1901: 331. Type locality: Cap Lopez, Congo français. Holotype: MNHN 1885-0404.
Physailia ansorgii Boulenger, 1910: 557. Type locality: Quanza River at Cunga, Angola. Syntypes (2): BMNH
1911.6.1.107 (1), NMW 45490 (1); NMW syntype illustrated in De Vos (1995: 305, fig. 208).
Physailia villiersi Boulenger, 1912b: 17, pl. 17 (fig. 6). Type locality: de la Lucala près de Cabinda, du Chiloango à
Mayili, de la Luali à Lundo, et de la Luculla. Syntypes: ANSP 38756–58 (3), BMNH 1912.4.1.419–420 (2),
BMNH 1912.4.1.421 (1), BMNH 1912.4.1.422–424 (3), BMNH 1912.4.1.425–429 (5), FMNH 56151 (2),
MRAC 1656–1659 (4), MRAC 1660–1661 (2), MRAC 1662 (1), NMW 45489 (16), ZMB 18818 (9).
Distribution: Ogowe, Congo, Quanza, Luculla, Chiloango Rivers (De Vos, 1995).
Parailia pellucida (Boulenger, 1901)
Physailia pellucida Boulenger, 1901e: 445. Type locality: Omdurman, Nile River. Syntypes (numerous): BMNH
1907.12.2.1942–1956 (14 or 15), BMNH 1907.12.2.1940–1941 (2), ? MCZ 32074 (1), MNHN 1907-0215 (3),
MNHN 1907-0216 (3).
Distribution: Nile River, Chad Basin, several west African river basins (De Vos, 1995).
Parailia somalensis (Vinciguerra, 1897)
Ailia somalensis Vinciguerra, 1897: 346. Type locality: Lugh, Fl. Ganana, Somaliland. Syntypes: BMNH
1961.5.3.12 (1), MSNG 14612 (4); one MSNG syntype illustrated in de Vos (1995: 321, fig. 212).
Physailia somalensis tanensis Whitehead, 1962a: 101, fig. 2. Type locality: Tana River at Hola, Kenya. Holotype:
BMNH 1961.5.3.7.
Distribution: Ganana and lower Tana Rivers (De Vos, 1995).
Parailia spiniserrata Svensson, 1933
Parailia spiniserrata Svensson, 1933: 73, pl. 4 (fig. 2); fig. 24. Type locality: MacCarthy Island Area, Gambia
River, British West Africa. Syntypes (about 25): NRM 11144 (4), NRM 14453 (1), NRM 14454 (8), NRM
21144 (2), NRM 31144 (1), NRM 31145 (1), NRM 31146 (4), NRM 31147 (1); one syntype illustrated in De
Vos (1995: 326, fig. 215).
Distribution: Gambia, Geba, and Jong River basins (De Vos, 1995).
PAREUTROPIUS Regan, 1920
Ansorgia Boulenger, 1912b: 17. Type species: Ansorgia vittata Boulenger, 1912. Type by monotypy. Gender: Feminine. Preoccupied by Ansorgia Warren, 1899, in Lepidoptera; replaced by Ansorgiichthys Whitley, 1935.
Pareutropius Regan, 1920a: 105. Type species: Pareutropius micristius Regan, 1920. Type by monotypy. Gender:
Masculine.
Eutropiellus Nichols & La Monte, 1933b: 5. Type species: Eutropiellus kasai Nichols & La Monte, 1933. Type byoriginal designation. Gender: Masculine.
Ansorgiichthys Whitley, 1935: 249. Type species: Ansorgia vittata Boulenger, 1912. Type by being a replacement
name. Gender: Masculine. Replacement for Ansorgia Boulenger, 1912; preoccupied by Ansorgia Warren, 1899,
in Lepidoptera.
Revision: Thys van den Audenaerde (1964, as Eutropiellus).
Pareutropius buffei (Gras, 1961)

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FERRARISEutropiellus buffei Gras, 1961: 406, fig. 3. Type locality: Bas-Ouémé, Bas-Dahomey. Syntypes (5): Lab. d'Hydrobiol. Cotonou, Benin (1), MNHN 1958-0001 (4).
Eutropiellus vandeweyeri Thys van den Audenaerde, 1964: 225, fig. 3. Type locality: Environment de Aba, Lower
Nigeria Delta, Nigeria. Holotype: MRAC 140281.
Distribution: Lower Ouémé River, Cross and Niger Rivers (De Vos, 1995).
Pareutropius debauwi (Boulenger, 1900)
Eutropius debauwi Boulenger, 1900a: 138, pl. 50 (fig. 2). Type locality: Uerre. Syntypes: BMNH 1900.12.13.8 (1),
MRAC 347 (1).
Ansorgia vittata Boulenger, 1912b: 17, pl. 19 (fig. 2). Type locality: Lower Congo. Syntypes: ANSP 38734–35 (2),
BMNH 1912.4.1.414 (1), BMNH 1912.4.1.415–417 (3), BMNH 1912.4.1.418 (1, skeleton), MRAC 1653 (1),
MRAC 1654 (1), MRAC 1655 (1), ZMB 18817 (1). Name as Ansorgia vitata on p. 17, A. vittata on plate legend(p. 28) and on plate; De Vos (1986: 37) cited both spellings and used Ansorgia vittata, thereby acting as firstreviser.
Ansorgia vittata bistriata Giltay, 1930b: 393, fig. 1. Type locality: Buta. Holotype: MRAC 20584.
Eutropiellus kasai Nichols & La Monte, 1933b: 5, fig. 4. Type locality: Luluabourg, Kasai, Belgian Congo. Holotype: AMNH 12338.
Distribution: Congo River system, Chiloango River, the Kouilou and Ogowe Rivers (De Vos, 1995).
Pareutropius longifilis (Steindachner, 1914)
Eutropius longifilis Steindachner, 1914: 537. Type locality: Deutsch-Ostafrika [Kiperege]. Syntypes: NMW 46507(2). Species illustrated and described in more detail in Steindachner (1915d: 75, pl. 3, fig. 2), with locality as:
Wasserlauf bei dem Dorfe Keperege; one syntype illustrated in De Vos (1995: 347, fig. 222).
Pareutropius micristius Regan, 1920a: 105. Type locality: Tanganyika Territory, Morogoro. Syntypes (7): BMNH
1920.3.8.10–12 (3).
Distribution: eastward flowing rivers north of the Ruvuma, Lake Chiuta, Ruvuma system, Lake Chilwa (De Vos,
1995).
Pareutropius mandevillei Poll, 1959
Pareutropius mandevillei Poll, 1959: 92, pl. 16 (fig. 4a). Type locality: Stanley Pool. Holotype: MRAC 98387.
Distribution: Congo River basin (De Vos, 1995).
PLATYTROPIUS Hora, 1937
Platytropius Hora, 1937a: 352. Type species: Pseudeutropius siamensis Sauvage, 1883. Type by original designation. Gender: Masculine.
Nemasiluroides Fowler, 1937: 137. Type species: Nemasiluroides furcatus Fowler, 1937. Type by original designation. Gender: Masculine.
Remarks: Platytropius Hora was published in January, 1937, and therefore has priority over Nemasiluroides Fowler,
1937, which was published on 19 May, 1937. A second paper by Hora (1937c: 39) which has been cited as theoriginal proposal of Platytropius (e. g., Eschmeyer, 1990) was not published until November, 1937.
Platytropius siamensis (Sauvage, 1883)
Pseudeutropius siamensis Sauvage, 1883b: 154. Type locality: Mé-Nam, Siam. Holotype: MNHN a-5002.
Nemasiluroides furcatus Fowler, 1937: 138, figs. 16–19. Type locality: Bangkok, Siam. Holotype: ANSP 67893.
Distribution: Chao Phraya River basin, Thailand.
PROEUTROPIICHTHYS Hora, 1937
Proeutropiichthys Hora, 1937a: 353. Type species: Eutropius macropthalmos Blyth, 1860. Type by original designation. Gender: Masculine.
Review: Hora (1941).
? Proeutropiichthys buchanani (Valenciennes, 1840)
Bagrus Buchanani Valenciennes, 1840, in Jacquemont, 1835–44: pl. 16 (fig. 3). Type locality: Hindustan, India.
Holotype: MNHN a-5721(1 of 3) or MNHN a-5722 (1 of 3). Name made available by figure caption.
Distribution: Hindustan, India.

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Remarks: Illustrated specimen should be treated as the holotype, if identifiable from among the specimens listed
herein. Inclusion into Proeutropiichthys based on personal observation of types.
? Proeutropiichthys goongwaree (Sykes, 1839)
Hypophthalmus Goongwaree Sykes, 1839: 163. Type locality: the Mota Mola near Poona, India. No types known.
Also described as new in Sykes (1840: 60); illustrated and described in more detail in Sykes (1841: 369, pl. 64,
fig. 3).
Distribution: Southern India (Jayaram, 1999).
Remarks: Placed in Pseudeutropius by Günther (1864) and Day (1875–78), then into Eutropiichthys by Hora
(1937e), where it has remained until now. However, the described species and illustrations are clearly not of a
species of either of these genera and more likely that of a species of Proeutropiichthys.
Proeutropiichthys macropthalmos (Blyth, 1860)
Eutropius macropthalmos Blyth, 1860: 156. Type locality: Tenasserim [Myanmar]. Type(s): Whereabouts unknown.
Proeutropiichthys taakree burmanicus Tilak, 1982: 35, figs. 1, 3, 5. Type locality: Burma. Holotype: ZSI FF771.
Distribution: Irrawaddy, Sittang, and Bago River basins, Myanmar.
Proeutropiichthys taakree (Sykes, 1839)
Hypophthalmus taakree Sykes, 1839: 163. Type locality: Deccan, India. No types known. Also described as new inSykes (1840: 60); illustrated and described more fully in Sykes (1841: 369, pl. 64, fig. 4).
Schillee Sykesii Jerdon, 1849: 335. Type locality: Cavery River, southern India. No types known.
Pseudeutropius longimanus Günther, 1864: 60. Type locality: India. Holotype: BMNH 1857.6.13.88 (skin).
Pseudeutropius megalops Günther, 1864: 60. Type locality: Godaveri at Mahadespur, Orissa, Central India. Holotype: BMNH 1860.3.19.784.
Distribution: Western Ghats, Kerala, Maharashtra, India (Jayaram, 1999).
PSEUDEUTROPIUS Bleeker, 1862
Pseudeutropius Bleeker, 1862b: 398. Type species: Eutropius brachypopterus Bleeker, 1858. Type by original designation. Gender: Masculine. Also in Bleeker (1862–63: 14, 74).
Pseudeutropius brachypopterus (Bleeker, 1858)
Eutropius brachypopterus Bleeker, 1858b: 169. Type locality: Palembang [Sumatra]. Holotype (115 mm TL):
Whereabouts unknown; holotype illustrated in Bleeker (1862–63: 71, pl. 75 [= Silur. pl. 27], fig. 1).
Distribution: Kapuas River (Roberts, 1989a); Sumatra.
Pseudeutropius mitchelli Günther, 1864
Pseudeutropius mitchelli Günther, 1864: 59. Type locality: Madras Presidency [India]. Syntypes: BMNH
1863.12.18.134–135 (2).
Distribution: Kerala, India (Jayaram, 1999).
Pseudeutropius moolenburghae Weber & de Beaufort, 1913
Pseudeutropius moolenburghae Weber & de Beaufort, 1913: 249, fig. 100. Type locality: Sumatra (Batang Hari
river) [Indonesia]. Syntypes: AMNH 9283 (1), ZMA 112681 (6).
Distribution: Batang Hari River, Sumatra (Weber & de Beaufort, 1913); Kapuas River (Roberts, 1989a).
SCHILBE Oken, 1817
Schilbe Oken, 1817: 1183. Type species: Silurus mystus Linnaeus, 1758. Type by monotypy. Gender: Masculine.
Pusichthys Swainson, 1838: 348, et seq. Type species: Schilbe uranoscopus Rüppell, 1832. Type by subsequentmonotypy. Gender: Masculine. Originally proposed as a subgenus of Silurus, without any included species.
Species first added in Swainson (1839: 307).
Eutropius Müller & Troschel, 1849: 6. Type species: “Bagrus schilboides Val. (Hypophthalmus niloticus Rüppell)”[= Bagrus schilbeides Valenciennes, 1840]. Type by monotypy. Gender: Masculine. Originally proposed as asubgenus of Bagrus.
Proeutropius Fowler, 1936b: 307. Type species: Silurus congensis Leach, 1818. Type by original designation. Gender: Masculine. Originally proposed as a subgenus of Eutropius.
Review: De Vos & Lévêque (1983), west Africa (as Eutropius).

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FERRARISKey: De Vos (1984a) Quanza and Bengo rivers, Angola (as Eutropius).
Schilbe angolensis (De Vos, 1984)
Eutropius angolensis De Vos, 1984a: 13, figs. 4, 5d. Type locality: Quanza river at Quimbango, Angola, 10º51'S,
17º30'E. Holotype: MRAC 78-6-P-899.
Distribution: Upper Quanza River, Angola (De Vos, 1995).
Schilbe banguelensis (Boulenger, 1911)
Eutropius banguelensis Boulenger, 1911a: 282, fig. 231. Lake Bangwelu. Holotype: BMNH 1907.9.30.8.
Eutropius nasalis Boulenger, 1915: 169. Type locality: Lac Moero. Lectotype: MRAC 14163; lectotype designatedby, and illustrated in, De Vos (1995: 124, fig. 158, bottom) as lectoholotype.
Distribution: Chambezi River, Lake Bangweulu, Luapula River and Lake Mweru (De Vos, 1995).
Schilbe bocagii (Guimarães, 1884)
Eutropius Bocagii Guimarães, 1884a: 85, unnumbered pl. Type locality: Dondo (Quanza fl.), Angola. Holotype: MB(destroyed, De Vos, 1995: 128). Illustration of holotype reproduced in De Vos (1984a: 5, fig. 1a).
Eutropius ansorgii Boulenger, 1910: 555. Type locality: Quanza River at Cunga, Angola. Syntypes (3): BMNH
1911.6.1.100–101 (2), NMW 46502 (1).
Eutropius seraoi Boulenger, 1910: 556. Type locality: Bengo River and from the Lucalla River at Lucalla, Angola.
Syntypes (10): ANSP 37956 (1), BMNH 1911.6.1.102–105 (4), BMNH 1911.6.1.106 (1), NMW 46524 (2),
ZMB 18226 (1); one NMW syntype illustrated in De Vos (1984a: fig. 3, and 1995: 129, fig. 160f).
Eutropius eclipsis Fowler, 1919b: 270, fig. 12. Type locality: Quanza River at Cunga, Angola. Holotype: USNM
42342. Illustration of holotype reproduced in De Vos (1984a: 5, fig. 1b).
Distribution: Lower Quanza River and Bengo Rivers, Angola (De Vos, 1995).
Schilbe brevianalis (Pellegrin, 1929)
Eutropius brevianalis Pellegrin, 1929b: 362. Type locality: Dehane (Nyong), Cameroun. Syntypes: MNHN 19280354 (1), MNHN 1928-0355 (1).
Distribution: coastal rivers in Nigeria and Cameroon (De Vos, 1995).
Schilbe congensis (Leach, 1818)
Silurus Congensis Leach, in Tuckey, 1818: 409. Type locality: Lower Congo. Holotype: BMNH 2005.5.17.6.
Eutropius congoensis Boulenger, 1899b: 105. Type locality: Lower Congo. Holotype: BMNH 2005.5.17.6. Unjustified emendation of Silurus congensis (De Vos, 1995).
Eutropius congolensis Boulenger, 1901a: 268. Type locality: Lower Congo. Holotype: BMNH 2005.5.17.6. Unjustified emendation of Silurus congensis Leach, 1818.
Distribution: Congo River system (De Vos, 1995).
Schilbe djemeri (Thys van den Audenaerde & De Vos, 1982)
Eutropius djemeri Thys van den Audenaerde & De Vos, 1982: 179, figs. 1–2. Type locality: River Djerem, below
falls at the rapids, 6–7 km south of Mbakaou, Cameroon. Holotype: MRAC 73-18-P-2310.
Distribution: Upper Sanaga River basin, Cameroon (De Vos, 1995).
Schilbe durinii (Gianferrari, 1932)
Eutropius Durinii Gianferrari, 1932: 141. Type locality: Lago Tanganyika. Holotype: MSNM 36; illustrated in De
Vos (1995: 146, fig. 169).
Distribution: Unknown (De Vos, 1995).
Remarks: Validity of species questioned by De Vos (1986, 1995), who indicated that the type locality is almost certainly wrong.
Schilbe grenfelli (Boulenger, 1900)
Eutropius grenfelli Boulenger, 1900a: 137, pl. 50 (fig. 1). Type locality: Bolobo. Holotype: BMNH 1900.6.23.6.
Eutropius gastratus Nichols & Griscom, 1917: 708, fig. 20. Type locality: Rungu, Congo. Holotype: AMNH 6404.
Eutropius bomae Lönnberg & Rendahl, 1922: 126. Type locality: Boma, lower Congo. Holotype: NRM 6024.
Distribution: Congo River basin, Ogowe, Nyanga, and Ntem-Campo River basins, Nyong and Lokondje basins (DeVos, 1995).
Schilbe intermedius Rüppell, 1832
Schilbe intermedius Rüppell, 1832: 6. Type locality: Nil. Syntypes: BMNH 1850.7.29.14 (1), SMF 202 (1), SMF

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2625 (1, dry), SMF 6403 (1, skeleton). BMNH specimen illustrated by Seegers (1996: 184, fig. 129) as holotype.
Schilbe auratus Joannis, 1835: [Cl. IV, pl. 5]. Type locality: Nil. Possible holotype: MNHN b-0346.
Schilbe Senegallus Valenciennes, in Cuvier & Valenciennes, 1840a: 378 (281 of Strasbourg deluxe edition). Type
locality: Sénégal. Syntypes (huit pouces): at MNHN.
Bagrus depressirostris Peters, 1852: 682. Type locality: Mozambique. Syntypes: BMNH 1863.4.30.15 (1), ZMB
3015 (2), ZMB 3016 (1). Illustrated and described in more detail in Peters (1868c: 25, pl. 4, fig. 5) as Eutropius
depressirostris.
Schilbe dispila Günther, 1864: 51. Type locality: River Niger, West Africa; Upper Nile, 500 miles south of Chartoum. Syntypes: BMNH 1852.2.22.17 (1), BMNH unregistered (2).
Schilbe senegalensis Günther, 1864: 51. Type locality: Senegal. Syntypes:at MNHN (not found). Unjustified emendation of Schilbe senegallus Valenciennes, 1840.
Schilbe senegalensis fasciata Steindachner, 1870: 983, pl. 6 (figs. 1–2). Type locality: Senegal bei St. Louis,
Dagana, Podor, und Bakel. Syntypes: NMW 44579–81 (61). Originally as Schilbe senegalensis var. fasciata.
Schilbe steindachneri Guimaraes, 1884b: 1, pl. 1 (figs. 1–2). Type locality: Cunene. Holotype: MB (destroyed; De
Vos, 1995: 162).
Schilbe bouvieri Rochebrune, 1885: 95. Type locality: [Casamance R., Senegal, w. Africa]. Type(s): at Museo Bouvieri.
Schilbe Emini Pfeffer, 1896: 32. Type locality: Muhale-Bach, Uniamwesi. Holotype: ZMB 12851.
Eutropius lemairii Boulenger, 1900a: 138, pl. 50 (fig. 3). Type locality: Lofoi, Katanga. Syntypes: BMNH
1900.12.13.24 (1), MRAC 293 (1).
Schilbe palmeri Svensson, 1933: 71, pl. 3 (fig. 4). Type locality: Gambia River, Mc Carthy Island area. Syntypes:
NRM 11145 (3), NRM 14452 (1), NRM 21145 (1).
Distribution: Widely distributed in Sub-Saharan Africa (De Vos, 1995).
Remarks: See De Vos & Skelton (1990) for discussion of validity and correct name of the species.
Schilbe laticeps (Boulenger, 1899)
Eutropius laticeps Boulenger, 1899b: 105, pl. 41 (fig. 3) [not fig. 2 as stated in text]. Type locality: Kutu, Lac
Léopold II. Holotype: MRAC 941.
Distribution: Congo River basin (De Vos, 1995).
Schilbe mandibularis (Günther, 1867)
Eutropius mandibularis Günther, 1867b: 112. Type locality: Bossumprah River, Gold Coast. Holotype: deposited at
Liverpool Museum. Holotype considered to be lost by Boulenger (1911: 290) and Trewavas (1943: 168).
Eutropius liberiensis Hubrecht, 1881: 69. Type locality: St. Paul’s River, Liberia. Holotype: RMNH 5328.
Eutropius mentalis Boulenger, 1901a: 269. Type locality: River Prah, Gold Coast. Syntypes: BMNH 1899.12.22.8–
10 (3).
Distribution: West Africa, from the St. Paul to the Prah River (De Vos, 1995).
Remarks: See De Vos (1983) for comments on taxonomy.
Schilbe marmoratus Boulenger, 1911
Schilbe marmoratus Boulenger, 1911f: 222. Type locality: la rivière Sankuru (Kasaï). Holotype (70 mm): at MHNL.
Schilbe congolensis Steindachner, 1912: 445. Type locality: Moloundou, Dscha, Kamerun. Syntypes: NMW 10698–
99 (2). Species described in more detail in Steindachner (1913: 30). Preoccupied in Schilbe by Eutropius congolensis Boulenger, 1901.
Distribution: Congo River basin and Shiloango system (De Vos, 1995).
Schilbe micropogon (Trewavas, 1943)
Eutropius micropogon Trewavas, 1943: 168. Type locality: River Volta, Gold Coast. Syntypes (3): BMNH
1944.2.9.16–17 (2), BMNH 1944.2.9.18 (1).
Distribution: lower course of west African rivers, from Senegal to Cameroon (De Vos, 1995).
Schilbe moebiusii (Pfeffer, 1896)
Eutropius möbiusii Pfeffer, 1896: 30. Type locality: Kingani-Fluss. Syntypes: ZMB 13680 (8, ? now 3); one syntype
illustrated in De Vos (1995: 235, fig. 185).

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FERRARISEutropius moebii Boulenger, 1911a: 293. Type locality: Kingani -Fluss. Syntypes: ZMB 13680 (8, ? now 3). Unjustified emendation of Eutropius möbiusii Pfeffer, 1896.
Distribution: Coastal rivers of Tanzania, including the Kingani, Rufiji, and Wani basins (De Vos, 1995).
Schilbe multitaeniatus (Pellegrin, 1913)
Eutropius multitaeniatus Pellegrin, 1913b: 273. Type locality: l’Ogôoué (Ngomo). Syntypes: MNHN 1908-0230(1), MNHN 1908-0231 (1), MNHN 1913-0266 (1).
Eutropius multilineatus Boulenger, 1916: 293. Unjustified emendation of Eutropius multitaeniatus Pellegrin, 1913
(de Vos, 1995).
Eutropius cameronensis Pellegrin, 1929b: 361. Type locality: Akonolinga (Nyong), ... Mfida (Nyong), .... Nyabessan (Ntem); Cameroun. Syntypes: MNHN 1928-0351 (1), MNHN 1928-0352 (1), MNHN 1928-0353 (1).
Distribution: Dja, Nyong, Igowe, Nyang, and N’dogo Rivers (De Vos, 1995).
Remarks: See De Vos (1983) for comments on taxonomy.
Schilbe mystus (Linnaeus, 1758)
Silurus mystus Linnaeus, 1758: 305. Type locality: In Nilo. Holotype: NRM 70; illustrated in De Vos (1995: 247, fig.
191).
Hypophthalmus niloticus Rüppell, 1829: 6, pl. 1 (fig. 1). Type locality: Nil. Syntypes: BMNH 1850.7.29.16 (1), possibly RMNH 2985 (1), SMF 752 [or 725] (1), SMF 2637 (1, dry), SMF 6050 (1, skeleton), SMF 6051 (1, skeleton), SMF 6053 (1).
Bagrus Adansonii Valenciennes, in Cuvier & Valenciennes, 1840a: 391 (290 of Strasbourg deluxe edition), pl. 414.
Type locality: Sénégal. Holotype: MNHN a-8670 (dry).
Schilbe Hasselquistii Valenciennes, in Cuvier & Valenciennes, 1840a: 377 (280 of Strasbourg deluxe edition). Type
locality: Nil. Holotype: MNHN b-0599.
Bagrus schilbeides Valenciennes, in Cuvier & Valenciennes, 1840a: 389 (289 of Strasbourg deluxe edition). Type
locality: Nil. Syntypes: MNHN b-0600 (1), MNHN b-0601 (1). Unneeded new name for Hypophthalmus niloticus Rüppell, 1829.
Eutropius obtusirostris Günther, 1864: 53. Type locality: India [in error]. Syntypes: BMNH 1842.3.26.28–29 (2).
Eutropius altipinnis Steindachner, 1894: 57, pl. 1 (fig. 1). Type locality: St. Paul -Flusse, bei Soforch Place. Syntypes: NMW 46501 (1), RMNH 5397 (2).
Distribution: Nile River basin and West Africa (De Vos, 1995).
Remarks: De Vos & Skelton (1990: 324, fig. 1) illustrated the holotype of Silurus mystus Linnaeus, and discussed
nomenclatural history of the specimen but, unfortunately, mislabeled the photographed specimen as NRM 63(rather than the correct number: NRM 70). De Vos (1995) concluded that Eutropius obtusirostris types probably
originated in West Africa.
Schilbe nyongensis (De Vos, 1981)
Eutropius nyongensis De Vos, 1981: 968, fig. 4. Type locality: Nyong river, above falls at 12 km. S.-E. of Eseka,
Cameroon. Holotype: MRAC 75-4-P-43.
Distribution: Nyong River, Cameroon, known only from type locality (De Vos, 1995).
Schilbe tumbanus (Pellegrin, 1926)
Eutropius tumbanus Pellegrin, 1926: 203. Type locality: Tondu (Lac Tumba); Congo belge. Syntypes (7): MNHN
1926-0161(1), MNHN 1926-0162 (1), MNHN 1926-0163 (1), MRAC 19685-86 (1), MRAC 19686 (1), MRAC
19687 (1), MRAC 19688 (1).
Distribution: Middle Congo River basin, abundant in Lake Tumba (De Vos, 1995).
Schilbe uranoscopus Rüppell, 1832
Schilbe uranoscopus Rüppell, 1832: 4, pl. 1 (fig. 1). Type locality: Ägypten, Nil bei Cairo. Lectotype: BMNH
1850.7.29.22, designated by De Vos (1986: 45).
Schilbe Isidori Valenciennes, in Cuvier & Valenciennes, 1840a: 375 (278 of Strasbourg deluxe edition), pl. 412.
Type locality: Egypte. Syntypes (no larger than 10 pouces): at MNHN.
Distribution: Nile River basin, Turkana system, Chad basin, Niger system, and Cross River (De Vos, 1995).
Schilbe yangambianus (Poll, 1954)
Eutropius yangambianus Poll, 1954: 61, fig. 4. Type locality: riv. Lubulu Iles du fleuve. Holotype: MRAC 123625.

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Distribution: Congo River basin and one location in upper Zambezi River (Skelton, 1993; De Vos, 1995).
Schilbe zairensis De Vos, 1995
Schilbe zairensis De Vos, 1995: 285, fig. 201. Type locality: Lemfu, Inkisi River, Lower Zaire Basin, Zaire, ca.
5º18'S, 15º13'E. Holotype: MRAC 86-21-P-98.
Distribution: Lower Congo River basin (De Vos, 1995).
SILONIA Swainson, 1838
Silonia Swainson, 1838: 345 et seq. Type species: Ageniosus (Silonia) lurida Swainson, 1838. Type by monotypy.
Gender: Feminine. Originally proposed as a subgenus of Ageniosus.
Silundia Valenciennes, in Cuvier & Valenciennes, 1840b: 48 (36 of Strasbourg deluxe edition). Type species: Silundia gangetica Valenciennes, 1840. Type by subsequent designation by Bleeker (1862b: 399). Gender: Feminine.
Silondia Günther, 1864: 65. Type species: Silundia gangetica Valenciennes, 1840. Unjustified emendation of Silundia Valenciennes, 1840. Gender Feminine.
Silonopangasius Hora, 1937a: 352. Type species: Ageneiosus childreni Sykes, 1839. Type by original designation.
Gender: Masculine.
Silonia childreni (Sykes, 1839)
Ageneiosus childreni Sykes, 1839: 165. Type locality: Deccan, India. No types known. Also proposed as new inSykes (1840: 62). Illustrated and described in more detail in Sykes (1841: 375, pl. 66, fig. 3).
Silundia Sykesii Day, 1876: 569. Type locality: Deccan and Kurnool, India. Syntypes: AMS B.8084 (1), NMW
44624 (1), ZSI F1230 (1). Originally as “Silundia Sykesii, n.s.? = ? Ageneiosus Childreni, Sykes, Bleeker, &Jerdon.”Distribution: Cauvery, Godavari and Krishna River basins, India (Jayaram, 1999).
Silonia silondia (Hamilton, 1822)
Pimelodus silondia Hamilton, 1822: 160, 375, pl. 7 (fig. 50). Type locality: Gangetic estuaries. No types known.
Ageniosus (Silonia) lurida Swainson, 1838: 345, fig. 85. Type locality: Gangetic estuaries. No types known. Name
made available to reference to Ham. pl. 7 fig. 50 [= Hamilton, 1822, pl. 7, fig. 50]. Unneeded new name for
Pimelodus silondia Hamilton.
Silundia Gangetica Valenciennes, in Cuvier & Valenciennes, 1840b: 49 (36 of Strasbourg deluxe edition), pl. 426.
Unneeded new name for Pimelodus silondia Hamilton, 1822, apparently to avoid tautonymy.
Distribution: Northern India, Bangladesh, Myanmar and Nepal (Jayaram, 1999).
Remarks: Redescribed in Hora (1938a).
SILURANODON Bleeker, 1858
Siluranodon Bleeker, 1858b: 253, 255, 256. Type species: Silurus auritus Geoffroy St. Hilaire, 1809. Type by monotypy. Gender: Masculine.
Siluranodon auritus (Geoffroy Saint-Hilaire, 1809)
Silurus auritus Geoffroy Saint-Hilaire, 1809: pl. 11 (figs. 1–2). Type locality: fl. Nil, Egypt. Possible holotype or
syntypes: MNHN a-8954 (4), MNHN a-8956 (1). Name available from caption on plate, with illustrated specimen the holotype, if identifiable. Description of species in Geoffroy St. Hilaire (1827: 291–299).
Distribution: Nile, Chad, Niger, Volta, Comoe Rivers (De Vos, 1995).
Genus inquirendum, Schilbidae
Schillee Jerdon, 1849: 335. Type species: apparently not designated; name based on two species. Name credited toCuvier, which may refer to Schilbe, but included species belong to two other genera.
SCOLOPLACIDAE Bailey & Baskin, 1976
Scoloplacinae Bailey & Baskin, 1976: 5. Type genus: Scoloplax Bailey & Baskin, 1976.
Revision: Schaefer et al. (1989).

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FERRARISPhylogeny: Schaefer (1990).
Taxonomic summary: Schaefer (2003a).
1 genus, 4 species; no named fossil taxa.
SCOLOPLAX Bailey & Baskin, 1976
Scoloplax Bailey & Baskin, 1976: 5. Type species: Scoloplax dicra Bailey & Baskin, 1976. Type by original designation. Gender: Feminine.
Scoloplax dicra Bailey & Baskin, 1976
Scoloplax dicra Bailey & Baskin, 1976: 7, figs. 1–3. Type locality: Isolated Ox-Bow lagoon off the Río Iténez, ca.
400 m southwest of the river at a point opposite Costa Marques (Brazil), Dept. of Beni, Bolivia, 12º28.38'S,
64º16.59'W. Holotype: AMNH 32482.
Distribution: Amazon and Paraguay River basins (Schaefer, 2003a).
Scoloplax distolothrix Schaefer, Weitzman & Britski, 1989
Scoloplax distolothrix Schaefer, Weitzman & Britski, 1989: 191, figs. 5, 6. Type locality: Small tributary of Rio
Batovi, upper Rio Xingu, Mato Grosso, Brazil, approx. 12º58'S, 53º37'W. Holotype: MZUSP 39065.
Distribution: Tocantins/Araguaia, Xingu and Paraguay River basins (Schaefer, 2003a).
Scoloplax dolicholophia Schaefer, Weitzman & Britski, 1989
Scoloplax dolicholophia Schaefer, Weitzman & Britski, 1989: 196, figs. 9, 10. Type locality: In igarapé, trib. of Rio
Tarumãzinho, approximately 45 km north of Manaus, Amazonas, Brazil, 2º42'S, 60º03'W. Holotype: MZUSP
6788.
Distribution: Negro River basin, Brazil (Schaefer, 2003a).
Scoloplax empousa Schaefer, Weitzman & Britski, 1989
Scoloplax empousa Schaefer, Weitzman & Britski, 1989: 194, figs. 7, 8. Type locality: Rio Ivinheima 70 km
upstream from its confluence with Rio Paraná and Rio dos Bandeirantes, Mato Grosso, Brazil, approx. 22º35'S,
53º30'W. Holotype: MZUSP 39075.
Distribution: Amazon and Paraguay/Paraná River basins (Schaefer, 2003a).

SILURIDAE Cuvier, 1816
Siluroïdes Cuvier, 1816: 199. Type genus: Silurus Linnaeus, 1758.
Kryptopterini Bleeker, 1862 (in Bleeker, 1862–63): 18, 85. Type genus: Kryptopterus Bleeker, 1857.
Phalacronotini Bleeker, 1862 (in Bleeker, 1862–63):18, 90. Type genus: Phalacronotus Bleeker, 1857.
Monophyly: Bornbusch (1991b).
Phylogeny: Bornbusch (1991b; 1995).
Revision: Kobayakawa (1989, Silurus sensu lato, including Pterocryptis).
Review: Chen (1977, China).
Checklist: Haig (1952, with key to genera).
Identification guide: Jayaram (1977d, South Asia).
Remarks: Although the monophyly of the family appears to be well supported (see Bornbusch, 1991b), the limits ofmany of the included genera are not clear. As such, the composition of the genera as listed herein are likely tochange with additional study.
12 genera, 94 species; 3 named fossil species of uncertain status.
Incertae sedis
Kryptopterus indicus Datta, Barman & Jayaram, 1987
Kryptopterus indicus Datta, Barman & Jayaram, 1987: 29, fig. 1. Type locality: Hornbill Point, Namdapha River,Namdapha Wildlife Sanctuary, Arunachal Pradesh, India. Holotype: ZSI FF1699.
Distribution: Known only from type locality (Jayaram, 1999).

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Remarks: Generic placement undetermined. Illustration of type clearly not that of a species of Kryptopterus.
BELODONTICHTHYS Bleeker, 1857
Belodontichthys Bleeker, 1857b: 472. Type species: Belodontichthys macrochir Bleeker, 1857. Type by monotypy.
Gender: Masculine.
Phylogeny: Howes & Fumihito (1991).
Belodontichthys dinema (Bleeker, 1851)
Wallago dinema Bleeker, 1851d: 202. Type locality: Banjermasin. Syntypes (2, 190, 195 mm TL): possibly RMNH
6834 (1 or 2 of 5. Syntype illustrated in Bleeker (1862–63: pl.86 [= Silur. 38], fig. 2) as Belodontichthys macrochir.
Belodontichthys macrochir Bleeker, 1857b: 472. Type locality: Banjermasin. Syntypes (2, 190, 195 mm TL): possibly RMNH 6834 (1 or 2 of 5). Unneeded new name for Wallago dinema Bleeker, 1851.
Distribution: Laos, Vietnam, Thailand, Malay Peninsula, Sumatra and Borneo (Roberts, 1989a).
Remarks: BMNH 1863.12.4.54 (1, 272 mm SL) is incorrectly registered as a type of Belodontichthys macrochir [=Wallago dinema], but is too long to be either of the syntypes.
Belodontichthys truncatus Kottelat & Ng, 1999
Belodontichthys truncatus Kottelat & Ng, 1999: 388, fig. 1. Type locality: Chao Phraya River, flood-waters 17.5 km
north of Ayuthya, Maharaj Prov., Thailand. Holotype: UMMZ 235105.
Distribution: Mekong and Chao Phraya River basins (Kottelat, 2001b).
CERATOGLANIS Myers, 1938
Ceratoglanis Myers, 1938: 98. Type species: Hemisilurus scleronema Bleeker, 1863. Type by original designation.
Gender: Masculine.
Ceratoglanis pachynemus Ng, 1999
Ceratoglanis pachynema Ng, 1999c: 390, figs. 3, 5, 6. Type locality: Prachinburi Market, Thailand. Holotype: CAS
96577.
Distribution: Chao Phraya and Mekong River basins (Ng, 1999c).
Ceratoglanis scleronemus (Bleeker, 1863)
Hemisilurus scleronema Bleeker, 1863 (in Bleeker, 1862–63): 93, pl. 101 [= Silur. pl. 53]. Type locality: Java
(Krawang). Holotype (399 mm TL): possibly RMNH 2918 (1). Also described as new in Bleeker (1863e and1863f).
Distribution: Pahang River basin, peninsular Malaysia, Baram, Barito, Kapuas and Rejang river basins, Borneo,
Citarum River basin, Java, and the Batang Hari and Siak River basins, Sumatra (Ng, 1999c).
HEMISILURUS Bleeker, 1857
Hemisilurus Bleeker, 1857b: 472. Type species: Wallago heterorhynchus Bleeker, 1853. Type by monotypy. Gender:
Masculine.
Diastatomycter Vaillant, 1891: 182. Type species: Diastatomycter chaperi Vaillant, 1891. Type by monotypy. Gender: Masculine. Also appeared in Vaillant (1893c: 61).
Phylogeny and biogeography: Bornbusch & Lundberg (1989).
Hemisilurus heterorhynchus (Bleeker, 1853)
Wallago heterorhynchus Bleeker, 1853b: 514. Type locality: Moara kompeh, Sumatrae orientalis, in fluviis. Holotype (326 mm TL): RMNH 6849.
Diastatomycter Chaperi Vaillant, 1891: 182. Type locality: Knapei River, Borneo. Holotype: MNHN 1891-0458.
Also described as new in Vaillant (1893c: 61); redescribed and illustrated in Vaillant (1894b: 70, pl. 2, fig. 2).
Distribution: Sumatra, and Borneo (Roberts, 1989a).
Remarks: BMNH 1863.12.4.58 (~205 mm SL) is incorrectly listed as a type of Wallago heterorhynchus as the species was named on a single individual and this specimen is too small.
Hemisilurus mekongensis Bornbusch & Lundberg, 1989
Hemisilurus mekongensis Bornbusch & Lundberg, 1989: 435, figs. 1–4. Type locality: Thailand, Ubon Ratchathani

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Province, Khong Chiam District, Mun River at Ban Dan, upstream of confluence with Mekong River, 15º18'N,
105º31'E. Holotype: UMMZ 214565.
Distribution: Mekong River basin (Kottelat, 2001b).
Hemisilurus moolenburghi Weber & de Beaufort, 1913
Hemisilurus moolenburghi Weber & de Beaufort, 1913: 212, figs. 84–85. Type locality: Batang Hari, Sumatra. Syntypes: ZMA 113564 (2).
Distribution: Batang Hari, Sumatra, and western Borneo (Roberts, 1989a).
KRYPTOPTERUS Bleeker, 1857
Kryptopterus Bleeker, 1857b: 472. Type species: Kryptopterus micropus Bleeker, 1857. Type by subsequent designation by Bleeker (1862: 395, or 1862–63: 18). Gender: Masculine.
Kryptopterichthys Bleeker, 1857b: 472. Type species: Silurus palembangensis Bleeker, 1852. Type by subsequent
designation by Bleeker (1862: 395, or 1862–63: 18). Gender: Masculine.
Cryptopterus Günther, 1864: 6, 38. Type species: Kryptopterus micropus Bleeker, 1857. Type by being a replacement name. Gender: Masculine. Unjustified emendation of Kryptopterus. Preoccupied by Cryptopterus Kaup,
1860, in fishes.
Cryptopterella Fowler, 1944a: 1. Type species: Cryptopterella beldti Fowler, 1944. Type by original designation.
Gender: Feminine.
Remarks: First reviser action in making Kryptopterus the valid name for this taxon appears to have been accomplished in Haig (1952: 106), in which Kryptopterichthys is treated as a junior synonym of Kryptopterus,
although the names were attributed to Bleeker (1858) which was, until recently, thought to be the source of bothnames.
Kryptopterus baramensis Ng, 2002
Kryptopterus baramensis Ng, 2002c: 68, figs. 1–2. Type locality: Borneo: Sarawak, Sungai Akah, above Long
Tebangan, 300 m below confluence with Sungai Pahang, below falls, 3º22'12"N, 114º56'06"E. Holotype: ROM
72477.
Distribution: Baram River basin, northern Borneo (Ng, 2002c).
Kryptopterus bicirrhis (Valenciennes, 1840)
Silurus bicirrhis Valenciennes, in Cuvier & Valenciennes, 1840a: 367 (272 of Strasbourg deluxe edition), pl. 411.
Type locality: Java. Holotype: MNHN a-9932; radiograph of holotype in Roberts (1989a: fig. 113, lower).
Cryptopterus amboinensis Günther, 1864: 40, 429. Type locality: Amboyna [apparently in error]. Syntypes (2):
BMNH 1855.3.24.14 (1).
Distribution: Mekong and Chao Phraya River basins, and Sundaland (Kottelat, 2001b).
Kryptopterus cryptopterus (Bleeker, 1851)
Silurus cryptopterus Bleeker, 1851b: 270. Type locality: Bandjarmassing. Holotype (110 mm TL): RMNH 6840 (1of 16).
Kryptopterus micropus Bleeker, 1857b: 472. Type locality: Bandjarmassing. Holotype (110 mm TL): RMNH 6840
(1 of 16). Unneeded replacement name for Silurus cryptopterus Bleeker, 1851, apparently to avoid tautonymy.
Distribution: Borneo and Sumatra, and possibly Tapi River (Ng, 2003i: 8).
Kryptopterus dissitus Ng, 2001
Kryptopterus dissitus Ng, 2001a: 198, fig. 1. Type locality: Laos: Champasak Province, Mekong River at Ban Hang
Khone, just downstream from Khone falls. Holotype: UMMZ 238017.
Distribution: Chao Phraya and Mekong River basins (Ng, 2001a).
Kryptopterus eugeneiatus (Vaillant, 1893)
Callichrous eugeneiatus Vaillant, 1893c: 61. Type locality: riv. Knapei et Sebroeang, près Smitow, Bornéo. Syntypes: MNHN 1891-0459 (1), MNHN 1891-0460 (1).
Distribution: Kapuas River basin (Roberts, 1989a).
Kryptopterus geminus Ng, 2003
Kryptopterus geminus Ng, 2003i: 2, figs 1, 2a. Type locality: Cambodia: Stung Treng, Mekong River 2 km downstream from mouth of Tonle San on sandbars, Mekong River drainage, 13º31'N, 105º56'E. Holotype: UMMZCHECKLIST OF CATFISHES, RECENT AND FOSSIL

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234664.
Distribution: Mekong, Mae Khlong, Bang Pakong and Chao Phraya River basins (Ng, 2003i).
Kryptopterus hesperius Ng, 2002
Kryptopterus hesperius Ng, 2002c: 70, fig. 3. Type locality: Thailand: Kanchanaburi Province, Kwae Noi River at
Sai Yok. Holotype: ZMUC P28551.
Distribution: Mae Khlong River basin, western Thailand (Ng, 2002c).
Kryptopterus lais (Bleeker, 1851)
Silurus laïs Bleeker, 1851e: 428. Type locality: Sambas, in fluviis. Holotype (125 mm TL): RMNH 6839 (larger of2).
Distribution: Sambas, Kapuas, Kahajan and Barito rivers, Borneo (Roberts, 1989a).
Kryptopterus limpok (Bleeker, 1852)
Silurus limpok Bleeker, 1852d: 583. Type locality: Palembang, Sumatra. Holotype (175 mm TL): possibly BMNH
1863.12.4.100 (150 mm SL).
Distribution: Mekong and Chao Phraya River basins, Malay Peninsula, Sumatra and Borneo (Kottelat, 2001b).
Kryptopterus lumholtzi Rendahl, 1922
Cryptopterus lumholtzi Rendahl, 1922: 200. Type locality: Bulungan, northeastern Borneo. Holotype: ZMUO
J5310.
Distribution: Bulungan, northeastern Borneo (Rendahl, 1922).
Kryptopterus macrocephalus (Bleeker, 1858)
Kryptopterichthys macrocephalus Bleeker, 1858b: 293. Type locality: Sumatra ?; Padang ?, in fluviis. Holotype (113mm TL): possibly BMNH 1863.12.4.99 (96 mm SL) or BMNH 1863.12.4.101. Illustrated in Bleeker (1862–63:
pl. 89 [= Silur. pl. 41], fig. 1).
Distribution: Peninsular Thailand, Malay Peninsula, Sumatra and Borneo (Roberts, 1989a).
Kryptopterus minor Roberts, 1989
Kryptopterus minor Roberts, 1989a: 149, fig. 115. Type locality: Mainstream of Sungai Pinoh at Nanga Saian, 45
km S of Nangapinoh, Western Borneo (Kalimantan Barat, Indonesia). Holotype: MZB 3638. Name spelled
Kryptopterus minimus on p. 146. As there has not been a first reviser action to fix the name, it is done here, with
Kryptopterus minor as the valid name.
Distribution: Kapuas River, western Borneo (Roberts, 1989a).
Kryptopterus mononema (Bleeker, 1847)
Silurus mononema Bleeker, 1847b: 8. Type locality: Surakarta, Java. Syntypes: possibly BMNH 1863.12.4.93 (1,145 mm SL), RMNH 6835 (2). One syntype illustrated in Bleeker (1862–63: 87, pl. 91 [= Silur pl. 43], fig. 1).
Also described as new in Bleeker (1847: 166).
Distribution: Java, Indonesia.
Kryptopterus palembangensis (Bleeker, 1852)
Silurus palembangensis Bleeker, 1852d: 584. Type locality: Palembang, in fluviis. Holotype (169 mm TL): Whereabouts unknown.
Distribution: Sumatra.
Remarks: BMNH 1863.12.4.101 (120 mm SL) is incorrectly registered as a type of Silurus palembangensis but
appears to be too small to be the holotype.
Kryptopterus paraschilbeides Ng, 2003
Kryptopterus paraschilbeides Ng, 2003d: 3, fig. 1. Type locality: Cambodia: Kompong Chhnang, Tonle Sap River,
17 km upstream from Kompong Chhnang. Holotype: UMMZ 238788.
Distribution: Mekong River basin (Ng, 2003d).
Kryptopterus piperatus Ng, Wirjoatmodjo & Hadiaty, 2004
Kryptopterus piperatus Ng, Wirjoatmodjo & Hadiaty, 2004: 92, fig. 1. Type locality: Indonesia, Sumatra: Aceh
Selatan: Sungai Lembang in front of camp, Suag Balimbing Research Station, Gunung Leuser National Park.
Holotype: MZB 8717.
Distribution: Lembang River, Alas River basin, Sumatra, Indonesia (Ng et al., 2004).
Kryptopterus sabanus (Inger & Chin, 1959)

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FERRARISOmpok sabanus Inger & Chin, 1959: 282. Type locality: Segama River at the Segama Estate near Lahad Datu,
Lahad Datu District, East Coast Residency, North Borneo. Holotype: FMNH 44828.
Distribution: Kapuas and Kinabatangan Rivers, northern Borneo (Roberts, 1989a).
Kryptopterus schilbeides (Bleeker, 1858)
Hemisilurus schilbeides Bleeker, 1858b: 297. Type locality: Palembang, Sumatra; Bandjermassin, Borneo. Syntypes(2: 96–99 mm TL): possibly BMNH 1863.12.4.157 (1, 79 mm SL, 91 mm TL), BMNH 1864.5.15.6 (1). Illustrated in Bleeker (1862–63: pl. 90 [=Silur. pl. 42], fig. 4).
Distribution: Lower Mekong River to Indonesia, in rivers, canals, ditches and swamps (Rainboth, 1996).
Remarks: Register number of syntype of Hemisilurus schilbeides incorrectly reported as BMNH 1863.12.11.157 in
Eschmeyer et al. (1998).
Species inquirendae, Kryptopterus
Cryptopterella beldti Fowler, 1944a: 2, unnumbered figure. Type locality: Borneo (aquarium fish). Holotype: ANSP
71571.
MICRONEMA Bleeker, 1857
Micronema Bleeker, 1857b: 472. Type species: Silurus hexapterus Bleeker, 1851. Type by monotypy. Gender: Neuter.
Micronema cheveyi (Durand, 1940)
Cryptopterus Cheveyi Durand, 1940: 19, pl. 4. Type locality: Rivière de Kaskos [ Cambodia]. Holotype: possibly at
Institut Océanographique Nhatrang.
Distribution: Mekong River basin (Chu et al., 1999), in rivers and canals (Rainboth, 1996).
Micronema hexapterus (Bleeker, 1851)
Silurus hexapterus Bleeker, 1851d: 203. Type locality: Bandjermassing. Holotype (145 mm TL): Whereabouts
unknown.
Distribution: Thailand to Indonesia, in rivers, streams, and canals (Rainboth, 1996).
Remarks: BMNH 1863.12.4.73 is incorrectly registered as the holotype of Silurus hexapterus Bleeker, but it is muchlarger (159 mm SL) than the listed length of the holotype.
Micronema moorei (Smith, 1945)
Kryptopterus moorei Smith, 1945: 342, fig. 78. Type locality: Menam Chao Phya, near Paknampo, central Thailand.
Holotype: USNM 109787.
Distribution: Chao Phraya and lower Mekong River basins, in streams and canals of the floodplain (Rainboth, 1996).
Remarks: Treated as a probable synonym of Kryptopterus cheveyi in Kottelat (2001b).
Micronema platypogon (Ng, 2004)
Kryptopterus platypogon Ng, 2004f: 2, figs. 1, 2a. Type locality: Borneo: Sarawak, Rajang River drainage, market at
Sibu, 2º17'18.6"N 111º49'49.2"E. Holotype: ZRC 45838.
Distribution: Rajang River basin, northern Borneo (Ng, 2004f).
OMPOK La Cepède, 1803
Ompok La Cepède, 1803: 49. Type species: Ompok siluroides La Cepède, 1803. Type by monotypy. Gender: Masculine.
Callichrous Hamilton, 1822: 149. Type species: Silurus pabda Hamilton, 1822. Type by subsequent designation by
Bleeker (1862: 395 or 1862–63: 17). Gender: Masculine. Originally proposed as a subgenus of Silurus.
Pseudosilurus Bleeker, 1857b: 472. Type species: Wallago leiacanthus Bleeker, 1853. Type by monotypy. Gender:
Masculine.
Silurodes Bleeker, 1857b: 472. Type species: Silurus macronema Bleeker, 1851. Type by monotypy. Gender: Masculine. Also in Bleeker (1858b: 255, 256, 271).
Revision: Ng (2003b, Ompok hypophthalmus group); Tan & Ng (1996, Ompok leiacanthus species group).
Review: Parameswaran et al. (1967, Indian species).

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Ompok bimaculatus (Bloch, 1794)
Silurus bimaculatus Bloch, 1794: 24, pl. 364. Type locality: Tranquebar. Lectotype: ZMB 2916, designated by, andillustrated in, Paepke (1999: 139, pl. 25, fig. 2).
Callichrous ceylonensis Günther, 1864: 46. Type locality: Ceylon. Syntypes: BMNH 1852.2.19.107–108 (2),
BMNH 1853.3.30.60 (1).
Distribution: Widely distributed in south and Southeast Asia, including Mekong River (Kottelat, 2001b), but probably more restricted to just south Asia and Sri Lanka.
Remarks: See Jayaram (1977a) for comments on type locality of Silurus bimaculatus, and Rema Devi &Emilyamma (1997) for comments on its identity.
Ompok binotatus Ng, 2002
Ompok binotatus Ng, 2002b: 26, fig. 1. Type locality: Indonesia: Borneo, Kalimantan Barat, Sungai Mandai Kechilnear its confluence with Kapuas mainstream, 18 km west southwest of Putussibau, 0º48'N, 112º47'E. Holotype:
FMNH 94243.
Distribution: Kapuas River basin, Borneo (Ng, 2002b).
Ompok borneensis (Steindachner, 1901)
Callichrous (Silurodes) borneensis Steindachner, 1901: 445, pl. 18 (fig. 3). Type locality: Baram-Flusse, Borneo.
Holotype (8.4 cm): at NMW or SMF.
Distribution: Baram River basin, Borneo (Steindachner, 1901).
Ompok fumidus Tan & Ng 1996
Ompok fumidus Tan & Ng 1996: 537, figs. 3, 4c. Type locality: North Selengor peat swamp forest, 43 km on roadfrom Tanjong Malim to Sungai Besar, Selangor, Malaysia. Holotype: ZRC 15049.
Distribution: peninsular Malaysia (Tan & Ng, 1996).
Ompok hypophthalmus (Bleeker, 1846)
Silurus hypophthalmus Bleeker, 1846a: 149. Type locality: Batavia. Type(s) (size and number not stated): Whereabouts unknown.
Silurus macronema Bleeker, 1851: 203. Type locality: Bandjermassing. Holotype (140 mm TL): possibly BMNH
1863.12.4.155 (121 mm SL); illustrated in Bleeker (1862–63: 83, pl. 88 [= Silur. pl. 40], fig. 2), as Silurodes
macronema.
Distribution: Ciliwung and Brantas River basins, Java, and Barito River basin, southern Borneo (Ng, 2003b).
Ompok jaynei Fowler, 1905
Ompok jaynei Fowler, 1905: 466, fig. 3. Type locality: Baram basin, Baram region of Sarawak, Borneo. Holotype:
ANSP 114890.
Distribution: Baram River basin, Borneo (Fowler, 1905).
Ompok leiacanthus (Bleeker, 1853)
Wallago leiacanthus Bleeker, 1853a: 189. Type locality: Marwang, in fluviis [now: East end of Danau Arang Arang,
Jambi Prov., Sumatra, Indonesia, by neotype designation]. Neotype: ZRC 38538, designated by Tan & Ng
(1996: 532).
Distribution: Sumatra.
Ompok malabaricus (Valenciennes, 1840)
Silurus Malabaricus Valenciennes, in Cuvier & Valenciennes, 1840a: 353 (262 of Strasbourg deluxe edition). Type
locality: Malabar. Syntypes: MNHN b-0607 (2).
Distribution: Goa, Kerala, India (Jayaram, 1999).
Ompok miostomus Vaillant, 1902
Wallago miostoma Vaillant, 1902: 44. Type locality: Tepoe, bords du Mahakam, Bornéo central. Syntypes: RMNH
7811 (2); syntype illustrated in Roberts (1982b: fig. 3b).
Distribution: Mahamam River basin, central Borneo (Vaillant, 1902).
Ompok pabda (Hamilton, 1822)
Silurus pabda Hamilton, 1822: 150, 374, pl. 25 (fig. 47). Type locality: ponds and rivers of Bengal. No types known.
Silurus (Callichrus) vittatus Swainson 1839: 306. Type locality: ponds and rivers of Bengal. No types known. Name
made available by reference to “Ham. pl. 25, f. 47 ” [= Hamilton (1822: p. 25, fig. 47)]. Unneeded new name for372 · Zootaxa 1418 © 2007 Magnolia Press

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Silurus pabda Hamilton, 1822.
Distribution: Brahmaputra and Ganges River basins, India and Bangladesh; Pakistan; Myanmar (Jayaram, 1999).
Ompok pabo (Hamilton, 1822)
Silurus pabo Hamilton, 1822: 153, 375, pl. 22 (fig. 48). Type locality: Brahmaputra River, towards Assam. No types
known.
Silurus (Callichrus) erythrogaster Swainson, 1839: 306. Type locality: Brahmaputra River, towards Assam. No
types known. Name made available by reference to “Ham. pl. 17 f. 48 ” [sic pl. 22, fig. 48] (= Hamilton, 1822:
pl. 22, fig. 48). Unneeded new name for Silurus pabo Hamilton, 1822.
Distribution: Brahmaputra, Ganges, and Yamuna River basins, India and Bangladesh; Pakistan; Myanmar(Jayaram, 1999).
Ompok pinnatus Ng, 2003
Ompok pinnatus Ng, 2003f: 48, fig. 1. Type locality: Cambodia, Tonle Sap at Koompong Chhnang, fishing lot 9 insecond channel E of town. Holotype: UMMZ 232679.
Distribution: Chao Phraya River, Thailand and Mekong River basin, Cambodia (Ng, 2003f).
Ompok platyrhynchus Ng & Tan, 2004
Ompok platyrhynchus Ng & Tan, 2004: 2, figs. 1, 2a. Type locality: Borneo: Brunei Darussalam, Temburong district: Belalong sub-basin Sungai Esu, about 15 minutes upstream of Kuala Belalong Field Studies Centre(04º32'17.9"N 115º09'35.2"E). Holotype: ZRC 48678.
Distribution: Temburong River basin, northern Borneo (Ng & Tan, 2004).
Ompok pluriradiatus Ng, 2002
Ompok pluriradiatus Ng, 2002b: 28, fig. 3. Type locality: Indonesia: Borneo: Kalimantan Timur: Mahakam Riverdrainage, a swift blackwater stream entering Mahakam River from the left side downriver of Nuarapahu(0º14'S, 116º07'E). Holotype: MZB 5951.
Distribution: Mahakam River basin, eastern Borneo (Ng, 2002b).
Ompok rhadinurus Ng, 2003
Ompok rhadinurus Ng, 2003b: 1299, figs. 3, 5b. Type locality: Peninsular Malaysia, Selangor, North Selangor Peat
Swamp Forest, irrigation canal on western boundary. Holotype: ZRC 14897.
Distribution: Bernam, Endau, Pahang, Pattani and Perak River basins, peninsular Malaysia, Kapuas River basin,
Borneo, and Batang Hari, Deli, Indragiri and Musi River basins, Sumatra (Ng, 2003b).
Ompok sindensis (Day, 1877)
Callichrous Sindensis Day, 1877 (in Day, 1875–78): 476, pl. 110 (fig. 1). Type locality: Sind, from the Indus [Pakistan]. Holotype: ZSI 505.
Distribution: Sindh, Pakistan (Mirza, 2003).
Ompok urbaini (Fang & Chaux, 1949)
Cryptopterus urbaini Fang & Chaux, in Chaux & Fang, 1949a: 197, fig. 2. Type locality: Cambodge. Holotype:
MNHN 1966-0706.
Distribution: Mekong, Chao Phraya and Pasak River basins, Southeast Asia (Ng, 2003b).
Remarks: Ng (2003b) lists three syntypes, but a holotype was clearly indicated in the description.
Ompok weberi (Hardenberg, 1936)
Callichrous weberi Hardenberg, 1936a: 232. Type locality: Padang Tikarbay, Kapuas River basin, western Borneo.
Holotype (79 mm): Whereabouts unknown (see Ng & Siebert, 2002, for comments).
Distribution: Kapuas River, Borneo (Ng & Siebert, 2002).
Remarks: Redescribed in Ng & Siebert (2002).
Species inquirendae, Ompok
Ompok siluroïdes La Cepède, 1803: 49, 50, pl. 1 (fig. 2). Type locality: not stated [Asia]. Holotype: MNHN a-8669.
Silurus lamghur Heckel, 1838: 82, pl. 12 (figs. 5–6). Type locality: Kashmir. Holotype: NMW 44689.
Silurus canio Hamilton, 1822: 151, 374. Type locality: Ponds in the north-east parts of Bengal. No types known. Unpublished Hamilton illustration reproduced in Hora (1929: pl. 20, fig. 5).

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Silurus chechra Hamilton, 1822: 152, 375. Type locality: Kusi river [India]. No types known.
Silurus duda Hamilton, 1822: 152, 375. Type locality: river Kusi [India]. No types known.
Silurus (Callichrus) affinis Swainson, 1839: 306. Type locality: Kusi River [India]. No types known. Name made available by reference to “Ham. 152 (duda)” [= Hamilton, 1822: 152]. Unneeded new name for Silurus duda Hamilton,
1822.
Silurus (Callichrus) immaculatus Swainson, 1839: 306. Type locality: Bengal. No types known. Name made available byreference to “Ham. 151 (canio)” [= Hamilton, 1822: 151]. Unneeded new name for Silurus canio Hamilton, 1822.
Silurus (Callichrus) nebulosus Swainson, 1839: 306. Type locality: Kusi River [India]. No types known. Name madeavailable by reference to “Ham. 152 (nebulosus)” [=Hamilton, 1822: 152)]. Unneeded new name for Silurus
chechra Hamilton, 1822.
Silurus anostomus Valenciennes, in Cuvier & Valenciennes, 1840a: 363 (269 of Strasbourg deluxe edition), pl. 410. Type
locality: Bengale. Syntypes: MNHN a-8949 (2), MNHN a-8950 (2).
Silurus microcephalus Valenciennes, in Cuvier & Valenciennes, 1840a: 365 (271 of Strasbourg deluxe edition). Type
locality: Bengale. Holotype (6 pouces): at MNHN.
Silurus Mysoricus Valenciennes, in Cuvier & Valenciennes, 1840a: 364 (270 of Strasbourg deluxe edition). Type locality:
Mysore. Syntypes: MNHN 0000-3107 (2).
Silurus indicus M’Clelland & Griffith, in M’Clelland, 1842: 583. Type locality: Loodianah, the Punjab, and Cabool
River at Jullalabad. Holotype: Whereabouts unknown, possibly at BMNH.
Cryptopterus latovittatus Playfair, 1867: 16. Type locality: Cachar, eastern provinces of British India, within the watershed of the Burhampooter. Syntypes: BMNH 1867.2.14.72–73.
Pseudosilurus macropthalmos Blyth, 1860: 156. Type locality: Tenasserim. Holotype: Whereabouts unknown.
Callichrous nigrescens Day, 1870c: 616. Type locality: Throughout the branches of the Irrawaddi, in the Pegu and Sittoung rivers, Burma. Syntypes: AMS B.7636 (1), ZSI A.500 (2, lost).
Callichrous notatus Day, 1870c: 616. Type locality: Rivers of Burma. Possible syntypes: AMS B.7982 (2), ZSI 1275 (1),ZSI A.499 (2, lost).
Callichrous egertonii Day, 1872: 710. Type locality: Subhimalayan range in the Punjaub. Syntype: AMS B.8065 (1).
Wallago krattensis Fowler, 1934b: 335, fig. 1. Type locality: Kratt, southeast Siam. Holotype: ANSP 60177.
Silurus goae Haig, 1952: 77, fig. on p. 78. Type locality: Goa, India. Holotype: SU 41889. Referred to Ompok rather than
Silurus by Kobayakawa (1989: 176).
PHALACRONOTUS Bleeker, 1857
Phalacronotus Bleeker, 1857b: 472. Type species: Silurus phalacronotus Bleeker, 1851. Type by absolute tautonymy. Gender: Masculine. Type designations of Silurus leptonema in Bleeker (1862: 395 and 1862–63: 18) areinvalid.
Phalacronotus apogon (Bleeker, 1851)
Silurus apogon Bleeker, 1851c: 67. Type locality: Banjermasin. Syntypes (4, 118–175 mm TL): RMNH 6843 (inpart).
Silurus leptonema Bleeker, 1852: 584. Type locality: Palembang, Sumatra. Holotype (268 mm TL): possibly at
RMNH.
Silurus micropogon Bleeker, 1855: 418, 419. Type locality: Pontianak; Bandjermasin, Borneo. Apparently an
unneeded new name for Silurus apogon Bleeker, 1851.
Distribution: Mekong and Chao Phraya river basins, Malay Peninsula, Sumatra and Borneo (Kottelat, 2001b).
Phalacronotus bleekeri (Günther, 1864)
Cryptopterus bleekeri Günther, 1864: 44. Type locality: Siam. Syntypes (2): BMNH 1863.12.4.105, and possibly
BMNH 1862.11.1.208.
Micronema bleekeri Bocourt, 1866: 17, pl. 1 (figs. 3, 3a–c). Type locality: [Menam R. at Bangkok]. Syntypes:
MNHN 0000-1546 (3).
Distribution: Mekong and Chao Phraya River basins, and Sundaland (Kottelat, 2001b).
Remarks: Günther (1864) cited Micronema bleekeri Bocourt in his account of Cryptopterus bleekeri. However, he
did not cite a Bocourt publication in his list of cited literature (pp. vii–x) and available evidence indicates that374 · Zootaxa 1418 © 2007 Magnolia Press

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Bocourt’s publication did not appear until 1866. Therefore, Günther’s name has priority and Bocourt’s must betreated as independently proposed and available.
Phalacronotus micronemus (Bleeker, 1846)
Silurus micronemus Bleeker, 1846b: 289. Type locality: Batavia. Holotype: RMNH 6841.
Silurus phalacronotus Bleeker, 1851e: 429. Type locality: Sambas. Holotype (220 mm TL): Whereabouts unknown.
Phalacronotus micruropterus Bleeker, 1857b: 473. Type locality: Sambas, in fluviis. Holotype (220 mm TL):
Whereabouts unknown. Unneeded replacement name for Silurus phalacronotus Bleeker, 1851, to avoid Stricklandian tautonymy.
Micronema typus Bleeker, 1858b: 300. Type locality: Batavia. Holotype: RMNH 6841. Unneeded new name for Silurus micronemus Bleeker, 1846.
Kryptopterus deignani Fowler, 1937: 136, figs. 10–12. Type locality: Me Poon, Siam. Holotype: ANSP 67884.
Distribution: Mekong and Chao Phraya River basins, and Sundaland (Kottelat, 2001b).
Remarks: BMNH 1863.12.4.75 (209 mm SL), BMNH 1863.12.4.95 (222 mm SL, 249 mm TL), and BMNH
1863.12.4.96 (122 mm SL) are incorrectly registered as the types of Silurus micronemus.
Phalacronotus parvanalis (Inger & Chin, 1959)
Kryptopterus parvanalis Inger & Chin, 1959: 284, fig. 46. Type locality: Kinabatangan River at Deramakot, Kinabatangan District, North Borneo. Holotype: FMNH 68014.
Distribution: Kinabatangan River at Deramakot, North Borneo, Malaysia (Inger & Chin, 1959).
PINNIWALLAGO Gupta, Jayaram & Hajela, 1981
Pinniwallago Gupta, Jayaram & Hajela, 1981: 291. Type species: Pinniwallago kanpurensis Gupta, Jayaram &Hajela, 1981. Type by original designation. Gender: Masculine.
Pinniwallago kanpurensis Gupta, Jayaram & Hajela, 1981
Pinniwallago kanpurensis Gupta, Jayaram & Hajela, 1981: 290, fig. 1. Type locality: ‘Bara Tal’ near village of Bhitargaon, Tehsil, Ghatampur, Kanpur, Uttar Pradesh, India. Holotype: ZSI FF1443.
Distribution: Known only from the type locality in Uttar Pradesh, India (Jayaram, 1999).
PTEROCRYPTIS Peters, 1861
Pterocryptis Peters, 1861: 712. Type species: Pterocryptis gangelica Peters, 1861. Type by monotypy. Gender: Feminine.
Apodoglanis Fowler, 1905: 463. Type species: Apodoglanis furnessi Fowler, 1905. Type by original designation.
Gender: Masculine.
Hito Herre, 1924a: 702. Type species: Hito taytayensis Herre, 1924. Type by original designation. Gender: Masculine.
Hitoichthys Herre 1924b: 1570. Type species: Hitoichthys taytayensis Herre, 1924. Type by original designation.
Gender: Masculine.
Penesilurus Herre, 1924a: 703. Type species: Penesilurus palavanensis Herre, 1924. Type by original designation.
Gender: Masculine. Also described as new in Herre (1924b: 1570); earliest publication not established.
Herklotsella Herre, 1933: 179. Type species: Herklotsella anomala, Herre, 1933. Type by original designation. Gender: Feminine. Publication date stated to be Dec. 1933, but perhaps did not appear until later (Eschmeyer, 1998);
priority over Herklotsella Fowler, 1934, which was published in January, 1934, is not clearly established.
Review: Ng & Freyhof (2001a), Vietnam.
Remarks: The revision by Kobayakawa (1989) included many of these species in an expanded Silurus sensu lato.
Haig (1952: 82) gave precedence to Hito over Hitoichthys, after concluding that priority of publication could
not be determined.
Pterocryptis anomala (Herre, 1933)
Herklotsella anomala Herre, 1933: 179. Type locality: Hong Kong market. Holotype: SU 26769.
Silurus sinensis Hora, 1937d: 343, figs. 8 (c–d). Type locality: Lunchow, China. Holotype: Originally ZMFMIB
13692, apparently now at ASIZB. Name proposed for specimen identified as Silurus wynaadensis in Tchang(1936: 35). Preoccupied by Silurus sinensis La Cepède, 1803; replaced by Silurus gilberti Hora, 1938.

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Silurus gilberti Hora, 1938c: 243. Type locality: Lunchow, China. Holotype: Originally ZMFMIB 13692, apparently
now at ASIZB. Replacement name for Silurus sinensis Hora, 1937.
Distribution: Rivers draining southeastern China, from Minjiang and Pearl Rivers to vicinity of Hong Kong (Ng &Chan, 2005).
Remarks: Synonymy follows Ng & Chan (2005).
Pterocryptis berdmorei (Blyth, 1860)
Silurichthys Berdmorei Blyth, 1860: 156. Type locality: Tenasserim provinces. Holotype: ZSI 481 (whereabouts
unknown).
Silurus morehensis Arunkumar & Tombi Singh, 1997: 73, fig. 1. Type locality: Moreh Bazar, Moreh, Manipur,
India. Holotype: MUMF 2211/1A.
Distribution: Southeastern Myanmar and possibly western Thailand (Ng & Freyhof, 2001a), Manipur, India (Arunkumar & Tombi Singh, 1997).
Pterocryptis bokorensis (Pellegrin & Chevey, 1937)
Penesilurus bokorensis Pellegrin & Chevey, 1937: 315. Type locality: [Bokor, Cambodia, elev. 800–1000 m]. Holotype: MNHN 1936-0167.
Distribution: Cambodia.
Remarks: Ng & Kottelat (1998b) indicate that the species is known only from holotype, but Rainboth (1996) implies
that it is a food fish, known from upland streams of the Mekong basin in Cambodia.
Pterocryptis buccata Ng & Kottelat, 1998
Pterocryptis buccata Ng & Kottelat, 1998b: 394, figs. 1, 2a, 3a–b. Type locality: Thailand: Kanchanaburi Province,
Amphoe Sai Yok, Mae Khlong basin. Holotype: ZRC 41496.
Distribution: Meklong River basin, Kanchanaburi Province, Thailand; in caves and epigean habitats (Ng & Kottelat,
1998b).
Pterocryptis burmanensis (Khin-Thant, 1966)
Silurus burmanensis Khin-Thant, 1966: 219, pls. 1–3. Type locality: Inlé Lake, s. Shan State, 23º35'N, 96º57'E,
Burma, elev. 2915 ft. Holotype: Zoological Museum, Art and Sciences University of Rangoon 332.
Distribution: Inle Lake, Myanmar.
Pterocryptis cochinchinensis (Valenciennes, 1840)
Silurus Cochinchinensis Valenciennes, in Cuvier & Valenciennes, 1840a: 352 (262 of Strasbourg deluxe edition).
Type locality: [Cochinchine]. Syntypes: MNHN 0000-0573 (1), MNHN b-0602 (1).
Distribution: Coastal Rivers of central Vietnam, between Vinh and An Lao (Ng & Freyhof, 2001a); Xijiang [= WestRiver], Jiulongjiang, Hainan Island (Chu et al., 1999); Nam Xam basin, Laos (Kottelat, 2001b).
Pterocryptis crenula Ng & Freyhof, 2001
Pterocryptis crenula Ng & Freyhof, 2001a: 630, fig. 6. Type locality: Vietnam: Quang Ninh province, Hai Ninh district, torrent at km 5 on road from Bac Phong Sinh to Mong Cai, 21º35'31"N, 107º43'52"E. Holotype: ZRC
46317.
Distribution: coastal rivers of northeastern Vietnam (Ng & Freyhof, 2001a).
Pterocryptis cucphuongensis (Mai, 1978)
Silurus cucphuongensis Mai, 1978: 245, fig. 112. Type locality: Cuc Phurong, Vietnam. Holotype: DVZUT 345.
Distribution: Song Iuong drainage, northern Vietnam (Ng & Freyhof, 2001a).
Pterocryptis furnessi (Fowler, 1905)
Apodoglanis furnessi Fowler, 1905: 463, fig. 2. Type locality: Baram River, Borneo. Holotype: ANSP 114894; holotype illustrated in Bornbusch (1991a: 1071, fig. 1).
Distribution: Niah and Baram River basins, Sarawak (Bornbusch, 1991a).
Remarks: Redescribed in Bornbusch (1991a).
Pterocryptis gangelica Peters, 1861
Pterocryptis gangelica Peters, 1861: 712. Type locality: Ganges River. Holotype: ZMB 4796; holotype illustrated in
Bornbusch (1991a: 1077, fig. 4).
Silurus afghana Günther, 1864: 34. Type locality: Afghanistan [in error: Assam, India]. Holotype: BMNH
1860.3.19.755.

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Silurus dukai Day, 1873: 239. Type locality: Darjeeling, India. Syntypes: AMS B.7571 (1), ZSI 1118 (1, lost), ZSI
1232 (1, lost).
Distribution: Northern India (Ng & Chan, 2005).
Remarks: Coad (1981b: 16) discussed the mistake in the reported type locality of Silurus afghana. Synonymy of P.
gangelica and S. afghana follows Ng & Chan (2005).
Pterocryptis inusitata Ng, 1999
Pterocryptis inusitata Ng, 1999a: 372, figs. 1–2. Type locality: Laos: Mekong Basin, Nam Theun watershed, Nam
Ong at Ban Don. Holotype: ZRC 41455.
Distribution: Nam Kading basin, Mekong River system, Laos (Ng, 1999a).
Pterocryptis taytayensis (Herre, 1924)
Hito taytayensis Herre, 1924a: 703. Type locality: Small fresh-water creek, near Taytay, Palawan, Philippines. Holotype: BSMP 9357 (presumed destroyed).
Hitoichthys taytayensis Herre 1924b: 1570. Type locality: Small fresh-water creek, near Taytay, Palawan, Philippines. Holotype: BSMP 9357 (presumed destroyed).
Penesilurus palavanensis Herre, 1924a: 704. Type locality: Lake Manguao, Palawan, Philippines. Holotype: BSMP
(considered destroyed). Also described as new in Herre (1924b: 1570), earliest not established.
Distribution: Palawan, Philippines.
Pterocryptis torrentis (Kobayakawa, 1989)
Silurus torrentis Kobayakawa, 1989: 171, fig. 33. Type locality: Lampae stream, Khaoluk village, Trang, Thailand.
Holotype: NSMT-P 50234.
Distribution: Trang and Chataburi, Thailand (Kobayakawa, 1989); Myanmar localities reported by Kobayakawa
may refer to specimens of Pterocryptis berdmorei (Ng & Freyhof, 2001a).
Pterocryptis verecunda Ng & Freyhof, 2001
Pterocryptis verecunda Ng & Freyhof, 2001a: 636, fig. 9. Type locality: Vietnam: Hai Phong province, Cat Ba
Island, Stream near entrance of Trung Trang cave, (20º47'17"N, 107º00'04"E). Holotype: ZRC 46316.
Distribution: Cat Ba Island, northeastern Vietnam (Ng & Freyhof, 2001a).
Pterocryptis wynaadensis (Day, 1873)
Silurus punctatus Day, 1868: 155. Type locality: Stream in Wynaad, India, elev. 3000 ft. Possible syntypes: AMS
B.7990 (1), BMNH 1868.5.14.6–7 (2), BMNH 1889.2.1.2521–2522 (2), NMW 77251 (1), ZMB 11221 (1), ZSI
461 (1), ZSI 1233 (1), ZSI A.480 (1). Preoccupied by Silurus punctatus Rafinesque, 1818, and Silurus punctatus Cantor, 1842.
Silurus wynaadensis Day, 1873c: 237. Type locality: in the Wynaad, in a stream about 3000 ft. above the level of thesea [India]. Possible syntypes: AMS B.7990 (1), BMNH 1868.5.14.6–7 (2), BMNH 1889.2.1.2521–2522 (2),NMW 77251 (1), ZMB 11221 (1), ZSI 461 (1), ZSI 1233 (1), ZSI A.480 (1). Replacement for Silurus punctatus
Day, 1868.
Distribution: Kerala, India (Ng & Kottelat, 1998b; Jayaram, 1999).
SILURICHTHYS Bleeker, 1856
Silurichthys Bleeker, 1856: 417, 418. Type species: Silurus phaiosoma Bleeker, 1851. Type by monotypy. Gender:
Masculine.
Revision: Ng & Ng (1998).
Silurichthys citatus Ng & Kottelat, 1997
Silurichthys citatus Ng & Kottelat, 1997: 204, fig. 1. Type locality: Kalimantan Barat, Sungai Sekumpai, a small forest stream flowing into Sungai Pinoh, 0º32'S, 111º39.5'E, Borneo. Holotype: MZB 3670.
Distribution: Kapuas River basin, western Borneo (Ng & Ng, 1998).
Silurichthys gibbiceps Ng & Ng, 1998
Silurichthys gibbiceps Ng & Ng, 1998: 301, figs. 1b, 1j, 2b, 3b, 6. Type locality: Kalimantan Tengah, Sungai Barito
basin, Sungai Paku-merah (0º35.171'S, 115º11.398'E) [Indonesia]. Holotype: MZB 6101.
Distribution: Barito River basin, southern Borneo (Ng & Ng, 1998).
Silurichthys hasseltii Bleeker, 1858

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Silurichthys Hasseltii Bleeker, 1858: 270. Type locality: Tjisekat, in fluviis. Neotype: RMNH 2992. Illustrated in
Bleeker (1862–63: pl. 87 [= Silur. pl. 39], fig. 1).
Distribution: Western Java, Bangka, Bintan, Batam, Singapore, and southern peninsular Malaysia (Ng & Ng, 1998).
Remarks: Species named on illustration that may have been based on RMNH 2992 or on another specimen that was
not preserved. In the absence of conclusive evidence that the RMNH specimen was the basis of the illustration,
Ng & Ng (1998: 305) selected RMNH 2992 as the neotype of the species.
Silurichthys indragiriensis Volz, 1904
Silurichthys indragiriensis Volz, 1904: 464. Type locality: Kwantang River, near Djapura (Indragari), Sumatra.
Holotype: NMW 44622.
Distribution: Central Sumatra, Bintan, Bangka, Billiton and peninsular Malaysia (Ng & Ng, 1998).
Silurichthys marmoratus Ng & Ng, 1998
Silurichthys marmoratus Ng & Ng, 1998: 310, figs. 1e, 1f, 2e, 3e, 11. Type locality: Sarawak: Sungai Sebiris, 13.8
km after Kampung Puteh turnoff, towards Lundu on Sematan-Lundu road (1º41'32"N, 109º47'0.8"E). Holotype:ZRC 40293.
Distribution: Northern and western Borneo (Sarawak, Brunei, and Kalimantan Barat) (Ng & Ng, 1998).
Silurichthys phaiosoma (Bleeker, 1851)
Silurus phaiosoma Bleeker, 1851: 428. Type locality: Sambas, in fluviis. Holotype (82 mm TL): RMNH 6831.
Distribution: Northwestern Borneo (Ng & Ng, 1998).
Remarks: BMNH 1863.12.4.109 (102 mm SL) and BMNH 1864.5.15.5 (79 mm SL, 100 mm TL) are registered as
types, but are too large to be the holotype.
Silurichthys sanguineus Roberts, 1989
Silurichthys sanguineus Roberts, 1989a: 151, fig. 119. Type locality: Sungai Tekam, a small forest stream, where it
enters Kapuas mainstream about 5–6 km upstream from Sanggau, Western Borneo (Kalimantan Barat, Indonesia). Holotype: MZB 3673; holotype illustrated in Ng & Ng (1998: fig. 13).
Distribution: Kapuas River basin, western Borneo; known only from holotype (Ng & Ng, 1998).
Silurichthys schneideri Volz, 1904
Silurichthys schneideri Volz, 1904: 463. Type locality: Upper Langkat, Danau near Sukaranda [Sumatra]. Holotype:
NMW 44623.
Silurichthys leucopodus Fowler, 1939: 56, figs. 4–6. Type locality: Waterfall at Trang, Siam. Holotype: ANSP
68463.
Distribution: Northern Sumatra, northern peninsular Malaysia, southern and southeastern Thailand, and southern
Cambodia (Ng & Ng, 1998).
SILURUS Linnaeus, 1758
Silurus Linnaeus, 1758: 304. Type species: Silurus glanis Linnaeus, 1758. Type by Linnaean tautonymy. On Official
List (ICZN Opinion 77 and Direction 56). Gender: Masculine.
Glanis Agassiz, 1857: 333. Type species: Glanis aristotelis Agassiz, 1857. Type by monotypy. Gender: Masculine.
Preoccupied by Glanis Spix & Agassiz, 1829, in fishes.
Parasilurus Bleeker, 1862b: 392, 394. Type species: Silurus japonicus Temminck & Schlegel, 1846. Type by original designation. Gender: Masculine. Also in Bleeker (1862–63: 17).
Revision: Kobayakawa (1989), Silurus sensu lato, including Pterocryptis.
Remarks: The name Duanensis appears in the title of Hu et al. (2004) as a generic name for a species referred to in
the text as a species of Silurus. Duanensis is considered a lapsus calami and, therefore, a nomen nudum.
Silurus aristotelis Agassiz, 1857
Glanis Aristotelis Agassiz, 1857: 333. Type locality: Greece. Syntypes: MCZ 7938 (6), USNM 55895 (1).
Silurus (Parasilurus) aristotelis Garman, 1890: 56. Type locality: Acheloüs River, Acarnania, Greece. Syntypes:
MCZ 7938 (6), USNM 55895 (1).
Distribution: Balkan Peninsula, Greece (Kobayakawa, 1989).
Silurus asotus Linnaeus, 1758
Silurus Asotus Linnaeus, 1758: 304. Type locality: Asia. Type(s): Whereabouts unknown. Neotype designation by

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Chen (1977: 206, 216) apparently valid inasmuch as ten specimens were all designated as a collective neotype(M. Kottelat, pers. commun.).
Silurus inermis Houttuyn, 1782: 338. Type locality: Japanse. No types known. Preoccupied by Silurus inermis Linnaeus, 1766; effectively replaced by Silurus imberbis Gmelin, 1789.
Silurus dauuricus Pallas 1787: 359, pl. 11 (fig. 11). Type locality: Ingoda, Onone et Arguno Dauuriae fluuiis.
Type(s) (20" 3'" TL): possibly at ZIN or ZMB.
Silurus imberbis Gmelin, 1789:1361. Type locality: Japonia. On Silurus inermis Houttuyn and effectively replacingthat name. Apparently intended as a new name to avoid the homonymy with Silurus inermis Linnaeus, 1766.
Centranodon japonicus La Cepède, 1803: 138, 139. Type locality: Japan. On Silurus inermis Houttuyn (1782: 338,
no. 27) and Silurus imberbis Gmelin.
Silurus punctatus Cantor, 1842: 485. Type locality: Chusan [China]. Syntypes: BMNH 1968.3.11.29 (3); possibly
BMNH 1843.7.21.6, BMNH 1843.7.21.24, and BMNH 1843.7.21.25 (not found), also BMNH 1860.3.19.736–
737 (2), BMNH 1860.3.19.785–786 (2). Preoccupied by Silurus punctatus Rafinesque, 1818; apparently not
replaced.
Silurus xanthosteus Richardson, 1845: 133, pl. 56 (figs. 12–14). Type locality: Chusan, Canton, China. Syntypes (3):
BMNH 1968.3.11.29 (1) and specimen used for drawing by Reeves.
Silurus japonicus Temminck & Schlegel, 1846, in Temminck & Schlegel, 1843–50: 226, pl. 104 (fig. 1). Type locality: Nagasaki, Japan. Lectotype: RMNH D675, designated by Boeseman (1947: 169).
Silurus cinereus Dabry de Thiersant, 1872: 189, pl. 47 (fig. 1). Type locality: Yang-tsee-kiang [China]. Type(s): possibly at MNHN.
Silurus bedfordi Regan, 1908g: 61, pl. 2 (fig. 3). Type locality: Kimhoa [and] Chong-ju, Corea. Syntypes: BMNH
1907.12.10.66 (1), BMNH 1907.12.10.67 (1).
? Parasilurus asotus longus Wu, 1930a: 255, fig. 1. Type locality: Creek at Tian-Tai Mountain, Chekiang [Zhejiang], China. Syntypes (2): possibly at MNHN. Originally as Parasilurus asotus var. longus.
Distribution: Zhujiang [= Pearl River], Changjiang [=Yangtze River], Huanghe [= Yellow River], Heilongjiang [=Amur River], China (Chu et al., 1999); Southern Hokkaido to Kyushu, Japan, as well as Taiwan, and northern
Vietnam (Matsuura et al., 2000); Common throughout Korea (Mori, 1952).
Remarks: Treated as valid as Parasilurus asotus in some recent publications.
Silurus biwaensis (Tomoda, 1961)
Parasilurus biwaensis Tomoda, 1961: 348, fig. 1. Type locality: Offshore of Onoé, Lake Biwa-ko, Japan. Holotype:
MIKU 34407 [now at FAKU].
Distribution: Lake Biwa, Japan (Matsuura et al. 2000); reported from the Ueno Formation, Pliocene of ancient Lake
Biwa, by Kobayakawa & Okuyama (1994a, b).
Silurus duanensis Hu, Lan & Zhang, 2004
Silurus duanensis Hu, Lan & Zhang, 2004: 586, 589, figs. 1–2. Type locality: underground rivers of Disu Town
(23º34'N, 108º01'E), Du’an County, Guangxi [China]. Holotype: ASIZB 73176.
Distribution: Hongushui River basin, Pearl River drainage, China (Hu et al., 2004).
Silurus glanis Linnaeus, 1758
Silurus glanis Linnaeus, 1758: 304. Type locality: Oriente, minus frequens in Europae lacubus. Syntypes: BMNH
1853.11.12.168 (1, skin), NRM 59 (1). Placed on Official List as type of Silurus (ICZN Direction 57).
Silurus silurus Wulff, 1765: 33. Type locality: Germany. No types known.
Silurus glanis aralensis Kessler, 1872: 48. Type locality: Amu-Darya, Syr-Darya, and Zeravshan rivers, cent. Asia.
Syntype (3): ZISP 2071 (1). Originally as Silurus glanis var. aralensis.
? Silurus chantrei Sauvage, 1882: 163. Type locality: Fleuve Koura à Tiflis. Syntypes: MNHN a-3932 (2); describedin more detail and illustrated in Sauvage (1884b: 19, pl. 1, fig. 1).
Distribution: Danube basin, all Baltic basin drainages, Elbe River, Lake Constance, Murtensee, and Rivers draining
the Caspian, Azov and Aral Seas (Lelek, 1987); Lake Urmia, Tedhzhen River and possibly Tigris River basin(Coad, 1995), widely introduced elsewhere in Europe, including Great Britain, France, Italy, the Iberian Peninsula, and rivers flowing into the Arctic Ocean.
Silurus grahami Regan, 1907

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Silurus grahami Regan, 1907a: 64. Type locality: Chien Kiung Lake, 30 mi. southeast of Yunnan Fu, China. Holotype: BMNH 1907.5.4.46.
Distribution: Fuxian Lake, Yunnan, China (Chu et al., 1999)
Silurus lanzhouensis Chen, 1977
Silurus lanzhouensis Chen, 1977: 210, pl. 2 (fig. 8). Type locality: [Yellow River system, Lanchow and Inner Mongolia, China]. Syntypes: Mus. Inst. Hydrobiol. Acad. Sinica, Hupei [LAN] 001-003 (3), [NEI] 004-005 (2),
[NEI] 769739-42 (4).
Distribution: Changjiang [=Yangtze River], Zhujiang [= Pearl River], Minjiang (Chu et al., 1999).
Silurus lithophilus (Tomoda, 1961)
Parasilurus lithophilus Tomoda, 1961: 350, fig. 2. Type locality: Near Onoé, Lake Biwa-ko, Japan. Holotype:
MIKU 34411 [now at FAKU].
Distribution: Lake Biwa and Lake Yogo, Japan (Masuda et al., 1984).
Silurus mento Regan, 1904
Silurus mento Regan, 1904a: 192. Type locality: Sea of Tien [Tien Chih], Yunnan Fu, China, elev. 6000 ft. Syntypes:
BMNH 1904.1.26.40–41 (2).
Distribution: Lakes of Dianchi and Yilong, Yunnan, China (Chu et al., 1999)
Silurus meridionalis Chen, 1977
Silurus soldatovi meridionalis Chen, 1977: 209, pl. 2 (fig. 7). Type locality: [Yangtze R. system, China]. Syntypes: ?
Mus. Inst. Hydrobiol. Acad. Sinica, Hupei 101–102 (2), 0719 (1), 66.5.194–195 (2), 731419 (1), 731494 (1),746003 (1), 746008–09 (2).
Distribution: Yangtze River basin, China.
Silurus microdorsalis (Mori, 1936)
Parasilurus microdorsalis Mori, 1936: 671, pl. 24 (fig. 1). Type locality: River Rakuto at Ei-yo, South Chosen[Korea]. Holotype: At Preparatory Department of Keijo Imperial University.
Distribution: Yalujiang [= Amnok River], China (Chu et al., 1999); Upper reaches of the Yalu, Han, Kim, Naktong
and Anpyeng Rivers, Korea (Mori, 1952).
Silurus soldatovi Nikolsky & Soin, 1948
Silurus soldatovi Nikolsky & Soin, 1948: 1359, fig. 1. Type locality: [Amur River, Lake Kabar at Elabuga, Khabarovskyi Krai, Russia]. Holotype: ZMMU P-6505.
Distribution: Heilongjiang [=Amur River] China and Russia; Liaohe, China (Chu et al., 1999)
Silurus triostegus Heckel, 1843
Silurus triostegus Heckel, 1843: 1090, pl. 13 (fig. 1). Type locality: dem Tigris bei Mossul. Syntypes (4): at NMW,
SMF 2623 (1).
Distribution: Tigris River basin (Coad, 1995); Euphrates River, Turkey (Ünlü & Bozkurt, 1996).
WALLAGO Bleeker, 1851
Wallago Bleeker, 1851b: 265. Type species: Silurus Mülleri Bleeker, 1846. Type by subsequent designation by
Bleeker (1862: 17, 79). Gender: Masculine.
Silurodon Kner, 1866: 546. Type species: Silurodon hexanema Kner, 1866. Type by monotypy. Gender: Masculine.
Also described as new in Kner (1867: 305).
Wallagonia Myers, 1938: 98. Type species: Wallago leerii Bleeker, 1851. Type by original designation. Gender:
Feminine.
Remarks: See Myers (1938: 98, 1948: 19) and Kottelat (2000b: 87) concerning the history of problems surrounding
designation of type species of Wallago.
Revision, with key to species: Roberts (1982b).
Wallago attu (Bloch & Schneider, 1801)
Silurus attu Bloch & Schneider, 1801: 378, pl. 75. Type locality: in lacubus Malabariae. Holotype: ZMB 8783 (dry,lost). As Silurus athu in text and index, Silurus attu on plate; first reviser apparently Bleeker (1862–63: 79) inwhich Wallago attu is treated as valid over Wallago atthu [sic.]).
Silurus boalis Hamilton, 1822: 154, 375, pl. 29 (fig. 49). Type locality: Rivers not only of the Gangetic provinces,

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but all over India, and is occasionally found in ponds. No types known.
Silurus (Callichrus) macrostomus Swainson, 1839: 306. Type locality: Rivers not only of the Gangetic provinces,
but all over India, and is occasionally found in ponds. No types known. Name made available by reference to“Ham. 154, pl. 29, fig. 49 ” [= Hamilton, 1822: 154, 375, pl. 29 (fig. 49)]. Unneeded new name for Silurus boalis Hamilton, 1822.
Silurus Wallagoo Valenciennes, in Cuvier & Valenciennes, 1840a: 354. Type locality: Bengale, .... la côte de Coromandel, .... pays des Birmans. Syntypes: based on specimens at MNHN and literature accounts in Bloch &Schneider (1801), Russell (1803), and Hamilton (1822).
Silurus Mülleri Bleeker, 1846b: 289. Type locality: Batavia. Type(s): Whereabouts unknown.
Wallago Russellii Bleeker, 1854c: 108. Type locality: Calcutta, in fluminae Hooghly ... Batavia, in fluviis. Syntypes(4, 285–485 mm TL): Whereabouts unknown, and numerous literature citations.
Wallago attu valeya Deraniyagala, 1953: 45. Type locality: Yakvala, Ceylon. Holotype: NMSL FF187; illustrated in
Deraniyagala (1952: pl. 13, as Wallagonia attu).
Distribution: India, Malay Peninsula, Sumatra, Java, and Mekong and Chao Phraya River basins (Kottelat, 2001b).
Remarks: Redescribed in Hora (1939b) as Wallagonia attu.
Wallago leerii Bleeker, 1851
Wallago Leerii Bleeker, 1851e: 427. Type locality: Sambas et Palembang, in fluviis. Syntypes (2, 225, 230 mm TL):RMNH 6833 (2 of 4).
Wallago nebulosus Vaillant, 1902: 46. Type locality: Tepoe, bords du Mahakam, Bornéo central. Holotype: RMNH
7812; holotype illustrated by Roberts (1982b: fig. 3a).
Wallagonia tweediei Hora & Misra, in Hora & Gupta, 1941: 18, figs. 2–3. Type locality: Kuala Tahan, Pahang,
Malaysia. Holotype: ZRC 350 (plaster cast with head and fins built in); ZSI F13365/1 (right anterior gill arch ofholotype).
Distribution: Malay Peninsula, Sumatra and Borneo (Kottelat, 2001b; Ng, 2004a).
Wallago maculatus Inger & Chin, 1959
Wallago maculatus Inger & Chin, 1959: 279. Type locality: Kinabatangan River at Deramakot, Kinabatangan District, North Borneo. Holotype: FMNH 68038.
Distribution: Kinabatangan River, Borneo; known only from type series (Roberts (1982b).
Wallago micropogon Ng, 2004
Wallago micropogon Ng, 2004a: 93, fig. 1. Type locality: Cambodia: Stung Treng morning market (13º30.0'N,
105º58.0'E). Holotype: UMMZ 232320.
Distribution: Mekong River basin of Cambodia, Laos and Vietnam, and middle Chao Phraya River basin (Ng,2004a).
Species inquirenda, Wallago
Silurodon hexanema Kner, 1866: 546. Type locality: Von Schanghai. Holotype (6 1/3"): at NMW. Described in moredetail and illustrated in Kner (1867: 305, pl. 12, fig. 2 [as Wallago attu ?]).
Species inquirendae, Siluridae
Silurus sinensis La Cepède, 1803: 58, 82, pl. 2 (fig. 1). Type locality: China. No types known: based on an unpublisheddrawing.
Silurus sinensis M’Clelland, 1844a: 402. Type locality: China. No types known. Preoccupied by Silurus sinensis La
Cepède, 1803.
Silurichthys basilewskii Bleeker, 1858b: 256. Type locality: China borealis. Based on specimens identified as Silurusasotus in Basilewky (1855). Originally Silurichthys ? basilewskii.
Belodontichthys javanensis Hardenberg, 1938: 311. Type locality: Fish market of Batavia. Holotype (215 mm): Whereabouts unknown.
† Silurus altus Sytchevskaya, 1989. [No other information; in Gayet & Meunier, 2003].

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Distribution: middle Miocene to lower Pliocene, Russia; Lower/Middle Miocene, China; Miocene or Pliocene of
Tueirusia Formation, Russia (Gayet & Meunier, 2003).
† Heterobranchus austriacus Thenius, 1952. [No other information; in Gayet & Meunier, 2003].
Distribution: Bruun-Vösendorf, Austria, Pannonian (Gayet & Meunier, 2003).
Remarks: Included in Siluridae and probably Silurus by Gayet & Meunier (2003), following Gaudant (1994).
† Silurus glanis atavus Bogačew, 1924. [No additional information; from Weiler, 1956: 187.]

SISORIDAE Bleeker, 1858
Sisorichthyoidei Bleeker, 1858b: 48, 50. Type genus: Sisor Hamilton, 1822.
Glyptosterni Gill, 1861c: 53. Type genus: Glyptosternon M’Clelland, 1842.
Erethistides Bleeker, 1862 (in Bleeker, 1862–63): 13. Type genus: Erethistes Müller & Troschel, 1849.
Bagarina Günther, 1864: 3, 9, 183. Type genus: Bagarius Bleeker, 1854.
Exostomatina Günther, 1864: 264. Type genus Exostoma Blyth, 1860.
Continae de Pinna, 1996: 64. Type genus: Conta Hora, 1950.
Glyptothoracini de Pinna, 1996: 64. Type genus: Glyptothorax Blyth, 1860.
Laguviini de Pinna, 1996: 65. Type genus: Laguvia Hora, 1921.
Pseudecheneidina de Pinna, 1996: 64. Type genus: Pseudecheneis Blyth, 1860.
Revision: Hora & Silas (1952a, glyptosternoids).
Review: Chu (1986, western China); Chu & Kuang (1990, Yunnan, China); Chu & Mo (1999, China); Menon (1999,
India); Mirza & Hameed (1974, Pakistan).
Phylogeny: de Pinna (1996); Diogo et al. (2002, 2003b); Hora & Silas (1952b, glyptosternoids), Chu (1979, glyptosternoids), He (1996b, glyptosternoids), Peng et al. (2004, glyptosternoids), Guo et al. (2005, glyptosternoids).
Identification guide: Jayaram (1979, South Asia).
Historical biogeography: Hora & Silas (1952b), He (1995, glyptosternoids).
Remarks: In 1996, de Pinna removed several taxa from the Sisoridae and placed them into the family Erethistidae. Thosetaxa, placed herein within the genera Conta, Erethistes, Erethistoides, Hara and Pseudolaguvia, together with thesubsequently named genera Ayarnangra and Caelatoglanis, appear instead to be part of a natural subgroup within
the Sisoridae that includes the genus Glyptothorax and possibly others, but not the glyptosternoid genera (Ferraris &Britz, 2005; pers. obs.; and pers. commun., H. H. Ng, 2005). Therefore, the taxa placed in the Erethistidae by dePinna are returned to the Sisoridae.
22 genera, 167 species; 1 named fossil species.

Incertae sedis
Laguvia manipurensis Arunkumar, 2000
Laguvia manipurensis Arunkumar, 2000: 194, fig. 1. Type locality: Lairaok Maru stream near Moreh, 110 km from
Imphal City, Manipur, India. Holotype: MUMF 3001/1A.
Distribution: Yu River basin, Chindwin River basin, Manipur, India (Arunkumar, 2000).
Remarks: Although described as a species of Laguvia, neither the description nor the illustration are sufficient toassign this species to genus or even family.
AYARNANGRA Roberts, 2001
Ayarnangra Roberts, 2001: 83. Type species: Ayarnangra estuarius Roberts, 2001. Type by original designation.
Gender: Masculine.
Ayarnangra estuarius Roberts, 2001
Ayarnangra estuarius Roberts, 2001: 84, figs. 1–3. Type locality: Pathein Chaung (=Ngawan Chaung) near Pathein,
lower Ayeyarwaddy basin, Myanmar. Holotype: KUMF 3190.

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BAGARIUS Bleeker, 1854
Bagarius Bleeker, 1854c: 121. Type species: Pimelodus bagarius Hamilton, 1822. Type by monotypy. Gender: Masculine.
Revision: Roberts (1983).
Bagarius bagarius (Hamilton, 1822)
Pimelodus bagarius Hamilton, 1822: 186, 378, pl. 7 (fig. 62). Type locality: Ganges River. No types known.
Pimelodus platespogon Valenciennes, 1840, in Jacquemont, 1835–44: pl. 18 (fig. 3). Type locality: l’Inde. Holotype:
MNHN 0000-2904 (1 of 2). Name made available by plate caption, therefore the illustrated specimen is theholotype if distinguishable.
Bagarius Buchanani Bleeker, 1854c: 121. Type locality: Calcutta, in flumine Hooghly; Surakata, Javae centrals, influmine Pepeh. Syntype (3): possibly NMV 46015 (1). Considered to be an unneeded new name for Pimelodus
bagarius Hamilton by Roberts (1983), to avoid Stricklandian tautonymy.
Distribution: Mekong, Chao Phraya and Ganges River basins (Kottelat, 2001b).
Remarks: Redescribed in Hora (1939a).
† Bagarius gigas Gunther, 1876
† Bagarius gigas Gunther, 1876: 436, pl. 16, fig. 1. Type locality: Highlands of Padang, Sumatra; Tertiary. Holotype
BMNH 47513 (Imperfect pectoral arch, etc.).
Distribution: Highlands of Padang, Sumatra; reportedly from Eocene, but age questioned by Gayet & Meunier(2003).
Remarks: See Sanders (1934) for additional information.
Bagarius rutilus Ng & Kottelat, 2000
Bagarius rutilus Ng & Kottelat, 2000b: 10, fig. 3. Type locality: Market in Hanoi, Viet Nam. Holotype: ZRC 40440.
Distribution: Nam Xan and Nam Ma basins, Laos, and Red River basin, Viet Nam and China (Kottelat, 2001b).
Bagarius suchus Roberts, 1983
Bagarius suchus Roberts, 1983: 442, figs. 2c, 4. Type locality: Thailand, Kemrat. Holotype: ANSP 89521.
Distribution: Mekong and Chao Phraya River basins (Kottelat, 2001b).
Bagarius yarrelli (Sykes, 1839)
Bagrus Yarrelli Sykes, 1839: 163. Type locality: Mota Mola at Poona, Deccan, India. Type(s): Whereabouts
unknown. Also described as new in Sykes (1840: 60); illustrated and described in more detail in Sykes (1841:
370, pl. 65, fig. 1).
Pimelodus Carnaticus Jerdon, 1849: 341. Type locality: Bowany River, southern India. No types known.
Bagarius lica Volz, 1903a: 557. Type locality: Moresi R. at Palembang, Sumatra. Holotype: at NMBE. Described in
more detail in Volz (1903b: 391).
Bagarius Nieuwenhuisii Popta, 1904: 190. Type locality: le Bo, Bornéo central. Holotype: RMNH 7561; describedin more detail, with illustration of holotype, in Popta (1906: 66, pl. 4, fig. 14).
Distribution: Widely distributed in southern and southeastern Asia (Kottelat, 2001b).
CAELATOGLANIS Ng & Kottelat, 2005
Caelatoglanis Ng & Kottelat, 2005: 14. Type species: Caelatoglanis zonatus Ng & Kottelat, 2005. Type by original
designation. Gender: Masculine.
Caelatoglanis zonatus Ng & Kottelat, 2005
Caelatoglanis zonatus Ng & Kottelat, 2005: 14, fig. 2. Type locality: Myanmar, Kayin State, stream “Chon Son”
between Kyondaw and Phadaw, about 20 km NW of Payathouzu, (at border with Thailand), 15º25'N, 98º15'E.
Holotype: ZRC 49885.
Distribution: Ataran River basin, Myanmar (Ng & Kottelat, 2005).
CONTA Hora, 1950
Conta Hora, 1950: 194. Type species: Pimelodus conta Hamilton, 1822. Type by original designation. Gender: Fem-CHECKLIST OF CATFISHES, RECENT AND FOSSIL

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inine.
Conta conta (Hamilton, 1822)
Pimelodus conta Hamilton, 1822: 191, 378. Type locality: River Mahananda [Bengal]. No types known. Previously
unpublished Hamilton illustration reproduced in Hora (1929: pl. 21, fig. 8), and subsequently reproduced in
Hora (1950: 297, fig. 4a) and Ng (2005a: 28, fig. 6).
Hara elongata Day, 1872: 704. Type locality: Stream near Garrow hills. Holotype: ZSI 436. Illustration of holotype
first published in Day (1877: pl. 92, fig. 5) and reproduced in Ng (2005a: 28, fig. 7).
Distribution: Bhareli and Mahananda Rivers, northeast Bengal, Garo Hills, Meghalaya; and Bangladesh (Jayaram,
1979); also Sarda River, Uttar Pradesh (Tilak, 1987).
Conta pectinata Ng, 2005
Conta pectinata Ng, 2005a: 24, fig. 1. Type locality: India: Assam: Dibrugarh. Holotype: ZRC 49672.
Distribution: Middle Brahmaputra River basin, Assam, India (Ng, 2005a).
ERETHISTES Müller & Troschel, 1849
Erethistes Müller & Troschel, 1849: 12. Type species: Erethistes pusillus Müller & Troschel, 1849. Type by monotypy. Gender: Masculine.
Erethistes maesotensis Kottelat, 1983
Erethistes maesotensis Kottelat, 1983: 71, figs. 1–2. Type locality: Thailand: Tak province: Mae Nam Moei, 5 km W
of Mae Sot (16º41'N, 98º31'E). Holotype: MHNG 2096.63.
Distribution: Mae Nam Moei, Salween River basin, Thailand (Kottelat, 1983).
Erethistes pusillus Müller & Troschel, 1849
Erethistes pusillus Müller & Troschel, 1849: 12, pl. 1 (fig. 3). Type locality: Assam. Holotype: ZMB 3102; holotype
illustration reproduced by Hora (1950: 187, pl. 1, fig. 5, as Erethistes pussilus).
Distribution: Ganges River basin, Assam, India.
ERETHISTOIDES Hora, 1950
Erethistoides Hora, 1950: 190. Type species: Erethistoides montana Hora, 1950. Type by original designation. Gender: Masculine.
Erethistoides ascita Ng & Edds, 2005
Erethistoides ascita Ng & Edds, 2005a: 240, fig. 1. Type locality: Nepal: Jhapa, Bhadrapur, Mechi River at Bhadrapur, 26º32'17.9"N, 88º6'6.1"E. Holotype: KU 35016.
Distribution: Rivers of lowland plains of southeastern Nepal (Ng & Edds, 2005a).
Erethistoides cavatura Ng & Edds, 2005
Erethistoides cavatura Ng & Edds, 2005a: 243, fig. 5. Type locality: Nepal: Chitawan, Dhungre River at Sauraha.
Holotype: OSUS 15572.
Distribution: Narayani River basin, Nepal (Ng & Edds, 2005a).
Erethistoides montana Hora, 1950
Erethistoides montana Hora, 1950: 191, pl. 1 (figs. 10–12). Type locality: Streamlets round about Tangla, Darrang
district, Assam [India]. Holotype: ZSI F314/2.
Distribution: Brahmaputra River basin, India (Ng, 2005f).
Erethistoides pipri Hora, 1950
Erethistoides montana pipri Hora, 1950: 193, pl. 1 (figs. 7–9). Type locality: Pipri, Rihand River, Mirzapur district,
U. P. [India]. Holotype: ZSI F315/2.
Distribution: Rihand River, Uttar Pradesh, India (Hora, 1950); known only from holotype (Jayaram, 1979).
Erethistoides sicula Ng, 2005
Erethistoides sicula Ng, 2005f: 2, fig. 1. Type locality: India: West Bengal, Schutunga River (tributary of the Mansai
River) as Ansole, 26º22'24"N 89º11'17"E. Holotype: UMMZ 243718.
Distribution: Mansai River drainage, Brahmaputra River basin, India (Ng, 2005f: 6).
EUCHILOGLANIS Regan, 1907

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Chimarrichthys Sauvage, 1874: 332. Type species: Chimarrichthys davidi Sauvage, 1874. Type by monotypy. Gender: Masculine.
Euchiloglanis Regan, 1907c: 158. Type species: Chimarrichthys davidi Sauvage, 1874. Type by being a replacement
name. Gender: Masculine. Proposed as a replacement for Chimarrichthys Sauvage, 1874, then considered to be
preoccupied by Cheimarrichthys Haast, 1874, in fishes.
Coraglanis Hora & Silas, 1952a: 12. Type species: Euchiloglanis kishinouyei Kimura, 1934. Type by original designation. Gender: Masculine.
Remarks: Chimarrichthys Sauvage, 1874, is not actually preoccupied by Cheimarrichthys Haast, 1874, but the namehas apparently not been used as valid since Regan (1907c) proposed Euchiloglanis as a replacement. The prevailing usage requirements of the Code require that Euchiloglanis be treated as valid here. Coraglanis Hora &Silas is generally treated as valid as a monotypic genus for Coraglanis kishinouyei in Indian literature. He(1996b) indicated that Euchiloglanis kishinouyei shared derived characters uniquely with Euchiloglanis davidi
and placed the two into Euchiloglanis.
Euchiloglanis davidi (Sauvage, 1874)
Chimarrichthys Davidi Sauvage, 1874: 333. Type locality: Yao-Tchy, Thibet oriental. Syntypes: BMNH 1923.3.13.1(1), MNHN 0000-6273 (3), MNHN 0000-6274 (3).
Distribution: Chinijiang, Sichuan (Chu et al., 1999); eastern Tibet.
Euchiloglanis kishinouyei Kimura, 1934
Euchiloglanis Kishinouyei Kimura, 1934: 178, pl. 6. Type locality: Kwan-hsien, Szechwan Province, China. Holotype: at Laboratory of Biological Department of the Shanghai Science Institute, Shanghai.
Distribution: Jinshajiang [upper Yangtze River basin] (Chu et al., 1999).
EXOSTOMA Blyth, 1860
Exostoma Blyth, 1860: 155. Type species: Exostoma berdmorei Blyth, 1860. Type by subsequent designation, by
Bleeker (1862–1863: 13). Gender: Neuter.
Remarks: See Regan (1923) for comments on this genus.
Exostoma berdmorei Blyth, 1860
Exostoma Berdmorei Blyth, 1860: 155. Type locality: Tenasserim [Burma]. Syntypes: ASB 597 (2), (now at ZSI).
Possible syntype: ASB 600.
Distribution: Eastern Myanmar.
Exostoma labiatum (M’Clelland, 1842)
Glyptosternon labiatus M’Clelland, 1842: 588. Type locality: Mishmee Hills Upper Assam [India]. Holotype:
BMNH 1860.3.19.97.
Distribution: Yaluzangbujiang [= upper Brahmaputra River], and Irrawaddy River (Chu et al., 1999).
Species inquirendae, Exostoma
Glyptosternum chaudhurii Hora, 1923b: 41, fig. 7. Type locality: Putao Plains, northern frontier of Burma. Holotype:ZSI F9741/1. Treated as a synonym of either E. labiatum or E. vinciguerrae.
Glyptosternum stuarti Hora, 1923b: 39, pl. 2 (figs. 4–6). Type locality: Nam-Yak River at Tanja, on the northern frontierof Burma. Holotype: ZSI F9742/1. Treated either as valid or as a synonym of E. labiatum.
Exostoma Vinciguerrae Regan, 1905a: 184. Type locality: Khakhyen Hills, upper Burma. Holotype: BMNH
1893.2.16.17. Treated either as valid or as a synonym of E. labiatum. See Steinitz (1961) for comments on taxonomy.
GAGATA Bleeker, 1858
Gagata Bleeker, 1858b: 204, 206. Type species: Pimelodus gagata Hamilton, 1822. Type by absolute tautonymy.
Gender: Feminine.
Callomystax Günther, 1864: 218. Type species: Pimelodus gagata Hamilton, 1822. Type by being a replacement
name. Gender: Masculine. Unneeded replacement for Gagata Bleeker, 1858.

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Revision: Hora & Law (1941); Roberts & Ferraris (1998).
Gagata cenia (Hamilton, 1822)
Pimelodus cenia Hamilton, 1822: 174, 376, pl. 31 (fig. 57). Type locality: Northern parts of Bengal, where it frequents rivers. No types known.
Distribution: Ganges, Indus and Mahanadi river basins (Roberts & Ferraris, 1998).
Gagata dolichonema He, 1996
Gagata dolichonema He, 1996a: 380, fig. 1. Type locality: Daojieba of Baoshan County, (24º41'N, 99º10'E), Yunnan
Province, China. Holotype: IHASW 791.
Gagata gasawyuh Roberts & Ferraris, 1998: 325, figs. 6–7. Type locality: Myanmar, Tenasserim River mainstream
upstream from Kita (or Htee-tah). Holotype: CAS 95544.
Distribution: Upper Salween River basin, China and Irrawaddy, Salween and Tenasserim River basins, Myanmar(Roberts & Ferraris, 1998).
Remarks: Synonymy based on recent examination of specimens from the vicinity of the type locality of Gagata dolichonema, and photograph of the holotype.
Gagata gagata (Hamilton, 1822)
Pimelodus gagata Hamilton, 1822: 197, 379, pl. 39 (fig. 65). Type locality: Fresh water rivers and estuaries of Bengal. No types known.
Gagata typus Bleeker, 1863c: 90. Type locality: Fresh water rivers and estuaries of Bengal. No types known.
Unneeded new name for Pimelodus gagata Hamilton, 1822.
Distribution: Ganges River basin, India and Bangladesh (Roberts & Ferraris, 1998).
Gagata itchkeea (Sykes, 1839)
Phractocephalus itchkeea Sykes, 1839: 164. Type locality: Deccan, India. Type(s): at BMNH. Also described as
new in Sykes (1840: 61) Illustrated and described in more detail in Sykes (1841: 373, pl. 67, fig. 1).
Arius pumilus Valenciennes, 1840, in Jacquemont, 1835–44: pl. 18 (fig. 1). Type locality: Inde. Syntypes: MNHN
0000-1208 (8). Name made available from caption on plate.
Distribution: Narmada, Krishna and Cauvery River basins, India (Roberts & Ferraris, 1998).
Remarks: Synonymy of Arius pumilus based on examination of types; illustrated specimen of A. pumilis is arguably
a holotype, but it could not be readily distinguished from remaining specimens.
Gagata melanoptera Roberts & Ferraris, 1998
Gagata melanopterus Roberts & Ferraris, 1998: 330, fig. 10. Type locality: Myanmar: Yangon Division, Hlaing
River, 16º53'41"N, 96º05'28"E. Holotype: USNM 348852.
Distribution: Irrawaddy, Rangoon, Sittang and lower Salween River basins, Myanmar (Roberts & Ferraris, 1998).
Gagata pakistanica Mirza, Parveen & Javed, 1999
Gagata pakistanica Mirza, Parveen & Javed, 1999: 1, fig. 1. Type locality: River Indus near Ghazi, Pakistan. Holotype: Disposition not stated in publication, but presumably deposited at GCM.
Distribution: Indus River, Pakistan (Mirza, 2003).
Gagata sexualis Tilak, 1970
Gagata sexualis Tilak, 1970b: 207, fig. 1. Type locality: North Koel River at Daltonganj (Chotanagpur) [South
Bihar, India]. Holotype: ZSI F5592/2.
Gagata youssoufi Ataur Rahman, 1976: 5, fig. 1. Type locality: river Meghna near Chandpur, Bangladesh. Holotype:at Museum of Freshwater Fisheries Research Station, Chandpur, Bangladesh.
Distribution: Ganges and Brahmaputra River basins (Roberts & Ferraris, 1998).
Remarks: Synonymy based on Heok Hee Ng (pers. comm., 2003).
GLARIDOGLANIS Norman, 1925
Glaridoglanis Norman, 1925b: 574. Type species: Exostoma andersonii Day, 1870. Type by monotypy. Gender:
Masculine.
Glaridoglanis andersonii (Day, 1870)
Exostoma andersonii Day, 1870b: 524. Type locality: Hotham; and Ponsee, China. Possible syntypes: ZSI F917374/1 [? = A.595] (2), ZSI A.596 (2, lost), AMS B.8081 (1).

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GLYPTOSTERNON M’Clelland, 1842
Glyptosternon M’Clelland, 1842: 584. Type species: Glyptosternon reticulatus M’Clelland, 1842. Type by subsequent designation by Bleeker (1862–63: 12). Gender: Neuter
Glyptosternum Agassiz, 1846: 164. Type species: Glyptosternon reticulatus M’Clelland, 1842. Unjustified emendation of Glyptosternon M’Clelland, 1842. Gender: Neuter.
Parexostoma Regan, 1905a: 182. Type species: Exostoma stoliczkae Day, 1877. Type by subsequent designation,
apparently by Jordan (1920: 515). Gender: Neuter.
Remarks: Blyth (1860: 154) restricted Glyptosternon to a single species, but did not, by that action, designate thatspecies, G. reticulatus, as type. See Regan (1923) for comments on this genus.
Glyptosternon akhtari Silas, 1952
Glyptosternum akhtari Silas, in Hora & Silas, 1952a: 11, pl. 1 (figs. 4–6). Type locality: Bamian River, Oxus watershed, Afghanistan. Holotype: ZSI F643/2.
Distribution: Bamian River, Oxus watershed, Afghanistan (Hora & Silas, 1952a).
Glyptosternon maculatum (Regan, 1905)
Parexostoma maculatum Regan, 1905a: 183. Type locality: Lhasa, Tibet. Syntypes: BMNH 1904.12.28.87–88 (2).
Distribution: Yaluzangbujiang [= upper Brahmaputra River] (Chu et al., 1999).
Glyptosternon malaisei Rendahl & Vestergren, 1941
Glyptosternon malaisei Rendahl & Vestergren, 1941: 213. Type locality: Kambaiti, Kachin State, Irrawaddy Riverdrainage, Burma, 7000 ft. elev. Holotype: NRM 10721.
Distribution: Irrawaddy River drainage, northern Myanmar (Rendahl & Vestergren, 1941).
Glyptosternon reticulatum M’Clelland, 1842
Glyptosternon reticulatus M’Clelland, 1842: 584. Type locality: Sir-i-Chusma, at the source of the Cabul River[Afghanistan]. No types preserved. Name spelled Glyptosternon retuculatus and Glyptosternon reticulatus in
original description; Kullander et al. (1999: 146) apparently serve as first revisers in selecting reticulatus.
Exostoma stoliczkae Day, 1877b: 782. Type locality: Upper waters of the Indus River. Possible syntypes: AMS I.122(1), ZSI F497 (1), ZSI 1196–98 (3, lost).
Exostoma gracile Gratzianov, 1907: 58. Type locality: Naryn R. drainage, upper Syr-Darya R. basin, Namanganskaya Obl., Uzbekistan. Holotype: ZMMU P-785.
Exostoma labrax Gratzianov, 1907: 59. Type locality: Garm dist., mountain spring Kartveng at kishlak Shulmak,
Garmskaya Obl., Tadzhikistan. Syntypes: ZMMU P-2034 (2).
Distribution: Upper Jhelum basin and Gilgit River, Indus River drainage, Pakistan and Kashmir (Kullander, 1999).
Remarks: See Steinitz (1961) for comments on taxonomy. Redescribed in Hora (1932).
GLYPTOTHORAX Blyth, 1860
Glyptothorax Blyth, 1860: 154. Type species: Glyptosternon striatus M’Clelland, 1842. Type by subsequent designation by Bleeker (1862– 1863: 13). Gender: Masculine.
Aglyptosternon Bleeker, 1862 (in Bleeker, 1862–63): 12. Type species: Silurus cous Linnaeus, 1766. Type by original designation. Gender: Neuter.
Euclyptosternum Günther, 1864: 183. Type species: Silurus cous Linnaeus, 1766. Unjustified emendation of Aclyptosternon [sic, Aglyptosternon Bleeker, 1862]. Gender: Neuter.
Pteroglanis Fowler, 1934a: 92. Type species: Pteroglanis horai Fowler, 1934. Type by original designation. Preoccupied by Pteroglanis Eigenmann & Pearson, 1924, in fishes, replaced by Pteropsoglanis Fowler, 1934. Gender: Masculine.
Pteropsoglanis Fowler, 1934b: 351. Type species: Pteroglanis horai Fowler, 1934. Type by being a replacement
name. Replacement for Pteroglanis Fowler, 1934; preoccupied by Pteroglanis Eigenmann & Pearson, 1924.
Gender: Masculine.
Sundagagata Boeseman, 1966: 243. Type species: Sundagagata robusta Boeseman, 1966. Type by original designation. Gender: Feminine.

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Paraglyptothorax Li, 1986: 524. Type species: Glyptosternum pallozonum Lin, 1934. Type by original designation.
Gender: Masculine. Originally proposed as a subgenus of Glyptothorax.
Superglyptothorax Li, 1986: 524. Type species: Glyptothorax coheni Ganguly, Datta & Sen, 1972. Type by original
designation. Gender: Masculine. Originally proposed as a subgenus of Glyptothorax.
Key: Mo & Chu (1986, Chinese species).
Review: Menon, M. A. S. (1955, India and Burma, with key), Sufi (1963, Pakistan).
Remarks: Extension of the eastern limit of the distribution of this genus into the Black Sea of Turkey noted by Coad
& Delmastro (1985).
Glyptothorax alaknandi Tilak, 1969
Glyptothorax brevipinnis alaknandi Tilak, 1969: 42, figs. 8–11. Type locality: Alaknanda River, near Srinagar, dist.
Pauri Garhwal, Uttar Pradesh, India. Holotype: ZSI F6154/2.
Distribution: Alaknanda River, Uttar Pradesh, India (Jayaram, 1999).
Glyptothorax anamalaiensis Silas, 1951
Glyptothorax anamalaiensis Silas, 1951b: 370. Type locality: Streams at base of Anamalai Hills, South India. Holotype: ZSI F629/2; illustrated in Silas (1951a: pl. 1, figs 1–3), as Glyptothorax prox. madraspatanus Day.
Distribution: Anamalai Hills, Kerala, India (Jayaram, 1999).
Glyptothorax annandalei Hora, 1923
Glyptothorax annandalei Hora, 1923b: 14, pl. 1 (fig. 3). Type locality: Nierolay stream, Bhavani River at the base of
Nilgiri Hills [India]. Holotype: ZSI F10135/1.
Distribution: Yaluzanbujiang [= Brahmaputra River], China (Chu & Mo, 1999).
Glyptothorax armeniacus (Berg, 1918)
Glyptosternum armeniacum Berg, 1918: 146. Type locality: River Mukhlassi-darasi, headwaters of Euphrates. Syntypes (5): possibly ZSI F11319/1 (1), ZIN 20806 (4); ZIN syntype illustrated in Berg (1931: pl. 1, figs. 3–5; fig.
6).
Distribution: Tigris River basin, Iran (Coad, 1995); Euphrates River basin, Turkey (Berg, 1918).
Glyptothorax botius (Hamilton, 1822)
Pimelodus botius Hamilton, 1822: 192, 378. Type locality: Northern rivers of Bengal [now India: west Bengal:
Hooghly River at Kalna, 23º13'30.0"N, 88º22'39.0"E by neotype designation]. Neotype: ZRC 50223; designated by, and illustrated in, Ng (2005e: 3, fig. 1).
Distribution: Hooghly River, India (Ng, 2005e: 8).
Remarks: Redescribed in Ng (2005e).
Glyptothorax brevipinnis Hora, 1923
Glyptothorax brevipinnis Hora, 1923b: 16, pl. 1 (fig. 4). Type locality: Unknown locality [probably India]. Syntypes: ZSI F10134/1 (4).
Distribution: Not known (Jayaram, 1999).
Glyptothorax buchanani Smith, 1945
Glyptothorax buchanani Smith, 1945: 402, fig. 89. Type locality: The Metum, a small swift affluent of the Mechem,
tributary of the Meping, in northern Thailand. Holotype: USNM 117754.
Distribution: Me Ping, Chao Phraya Basin, Thailand (Smith, 1945).
Glyptothorax burmanicus Prashad & Mukerji, 1929
Glyptothorax burmanicus Prashad & Mukerji, 1929: 184, pl. 7 (fig. 3); fig. 5. Type locality: Sankha, a large hillstream, midway between Kamaing and Mogaung, Myitkyina dist. [Burma]. Holotype: ZSI F10877/1.
Distribution: Irrawaddy and Salween Rivers (Chu & Mo, 1999).
Glyptothorax callopterus Smith, 1945
Glyptothorax callopterus Smith, 1945: 400, fig. 87. Type locality: Waterfall stream on Kao Chong, near Trang, in
Peninsular Thailand. Holotype: USNM 109820.
Distribution: Peninsular Thailand.
Glyptothorax cavia (Hamilton, 1822)
Pimelodus cavia Hamilton, 1822: 188, 378. Type locality: Northern rivers of Bengal. No types known.
Euglyptosternum lineatum Day, 1877 (in Day, 1875–78): 500, pl. 116 (fig. 7). Type locality: Jumna, near Suddya in

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upper Assam. Possible syntypes: AMS B.7509 (1), ZSI F1312 (1).
Distribution: Ganges River basin.
Glyptothorax coheni Ganguly, Datta & Sen, 1972
Glyptothorax coheni Ganguly, Datta & Sen, 1972: 342, figs. 3–4. Type locality: Subarnarekha River, Chotanagpur
Plateau, Bihar, India. Holotype: USNM 205612.
Distribution: Subarnarekha River, Bihar, India (Jayaram, 1999).
Glyptothorax conirostris (Seindachner, 1867)
Glyptosternum conirostre Steindachner, 1867e: 532, pl. 6 (fig. 2). Type locality: Simla. Holotype: at NMW.
Glyptosternum modestum Day, 1872: 714. Type locality: upper portion of the Jumna. Possible syntypes: AMS
B.7562 (1), AMS B.7564 (1), ZMB 2765 (1).
Distribution: Jamuna River, Mahananda River basin and Indus River basin (Jayaram, 1999).
Glyptothorax cous (Linnaeus, 1766)
Silurus cous Linnaeus, 1766: 504. Type locality: Syria. Possible syntype: BMNH 1955.6.25.2 (1).
Distribution: Tigris River basin, Iran (Coad, 1995).
Glyptothorax dakpathari Tilak & Husain, 1976
Glyptothorax dakpathari Tilak & Husain, 1976: 229, figs. 1–8. Type locality: Yamuna R., below Barrage, Dakpathar, District Dehra Dun, Uttar Pradesh, India. Holotype: ZSI NRS V-988.
Distribution: Yamuna River, Uttar Pradesh, India (Jayaram, 1999).
Remarks: Treated as a synonym of Glyptothorax gracilis by Menon (1999).
Glyptothorax davissinghi Manimekalan & Das, 1998
Glyptothorax davissinghi Manimekalan & Das, 1998: 87, unnumbered figure. Type locality: Karim Puzha,
Maancheri, Nilambur Reserve Forest, Kerala, India. Holotype: ZSIC 6008.
Distribution: Kerala, India (Manimelakan & Das, 1998).
Glyptothorax deqinensis Mo & Chu, 1986
Glyptothorax deqinensis Mo & Chu, 1986: 345, 350, fig. 6. Type locality: Deqin county (28º30'N, 99º00'E) Yunnan[China]. Holotype: KIZ 748621.
Distribution: upper Lancangjiang [= Mekong River], China (Chu et al., 1999).
Glyptothorax dorsalis Vinciguerra, 1890
Glyptothorax dorsalis Vinciguerra, 1890: 246, pl. 7 (fig. 4). Type locality: Meetan, Birmania [Myanmar]. Holotype:
MSNG 14417.
Distribution: Upper Irrawaddy River basin (pers. obs.); Nujiang [= Salween River], China (Chu et al., 1999).
Glyptothorax exodon Ng & Rachmatika, 2005
Glyptothorax exodon Ng & Rachmatika, 2005: 251, fig. 1. Type locality: Borneo: Kalimantan Barat, Sungai
Tekelan. Holotype: MZB 9940.
Distribution: Kapuas River basin, western Borneo (Ng & Rachmatika, 2005).
Glyptothorax fokiensis (Rendahl, 1925)
Glyptosternum fokiensis Rendahl, 1925: 307. Type locality: Lan-Hao, Lien-Cheng-Hsien, Fokien, s. China. Syntypes: NRM 10018 (2).
Glyptosternon punctatum Nichols, 1941: 1, figs. 1–2. Type locality: Kiating (Loshan), Szechwan Prov., China, elev.
1100 ft. Holotype: AMNH 15218.
Distribution: Changjiang [= Yangtze River], rivers east of Yuanjiang (Chu et al., 1999; as Glyptothorax fukiensis
fukiensis).
Remarks: Redescribed as Glyptothorax fukiensis in Xie et al. (2001).
Glyptothorax fuscus Fowler, 1934
Glyptothorax fuscus Fowler, 1934a: 89, figs. 31–33. Type locality: Chantaboon, Southeast Siam. Holotype (59 mm):ANSP 59358.
Distribution: Mekong and Chao Phraya River basins (Kottelat, 2001b).
Glyptothorax garhwali Tilak, 1969
Glyptothorax garhwali Tilak, 1969: 37, figs. 1–4. Type locality: Alaknanda River, near Srinagar, dist. Pauri Garhwal, Uttar Pradesh, India. Holotype: ZSI F6152/2.

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Distribution: Alaknanda River, Uttar Pradesh, India (Jayaram, 1999).
Glyptothorax gracilis (Günther, 1864)
Glyptosternum gracile Günther, 1864: 186. Type locality: Nepal. Holotype: BMNH 1845.1.9.846.
Distribution: Sikkim, India and Nepal (Jayaram, 1999).
Glyptothorax hainanensis (Nichols & Pope, 1927)
Glyptosternon hainanensis Nichols & Pope, 1927: 333, fig. 7. Type locality: Nodoa, Hainan [China]. Holotype:
AMNH 8362.
Distribution: Hainan Island, China (Chu et al., 1999, as Glyptothorax fukiensis hainanensis).
Glyptothorax honghensis Li, 1984
Glyptothorax fukiensis honghensis Li, 1984a: 66, fig. 1. Type locality: Hekou, Weishan, Nanjian, Hong River basin,
Yunnan Province, China. Syntypes: KIZ 6440430 (1), KIZ 6440474 (1), KIZ 6507022 (1), KIZ 6507133 (1),KIZ 6507134 (1), KIZ 6507137 (1). Name spelled Glyptothorax fukiensis honghensis on p. 66 and Glyptothorax fukiensis honghenensis on p. 69; G. honghensis treated as the valid spelling here, based on presumed etymology of name.
Distribution: Northeastern Laos, and Red River basin of Vietnam and China (Kottelat, 2001b).
Glyptothorax horai (Fowler, 1934)
Pteroglanis horai Fowler, 1934a: 92, figs. 37–40. Type locality: Sop Lao, in Maun Luang, Southeast Shan States[Burma]. Holotype: ANSP 59462.
Distribution: Mekong River basin, Myanmar.
Glyptothorax housei Herre, 1942
Glyptothorax housei Herre, 1942: 117, fig. 1. Type locality: Rapids in a mountain stream in the Anamallai Hills,
Puthototam Estate, four miles east of Valparai, Pollachi district, South India. Holotype: SU 36531.
Distribution: Kerala, India (Jayaram, 1999).
Glyptothorax indicus Talwar, 1991
Glyptothorax horai Shaw & Shebbeare, 1936: 188, unnumbered pl. Type locality: Streams of Terai, northern Bengal.
Holotype: ZSI F11376/1. Preoccupied in Glyptothorax by Pteroglanis horai Fowler, 1934; replaced by Glyptothorax indicus Talwar, 1991.
Glyptothorax indicus Talwar, in Talwar & Jhingran, 1991: 654, fig. 210. Type locality: Streams of Terai, n. Bengal.
Holotype: ZSI F11376/1. Replacement name for Glyptothorax horai Shaw & Shebbeare, 1936, preoccupied in
Glyptothorax by Pteroglanis horai Fowler, 1934.
Distribution: Kosi River, Rihand River and rivers of the Vindhya Mountain range, India, as well as the Punjab and
Nepal (Jayaram, 1999).
Glyptothorax interspinalus (Mai, 1978)
Glyptosternon interspinalum Mai, 1978: 271, fig. Type locality: Creeks in n. Vietnam. Syntypes: probably at
DVZUT.
? Glyptothorax merus Li, 1984b: 79, fig. 2. Type locality: Jingdong Co., Yunnan, China. Holotype: KIZ 737159.
Distribution: Nam Ma Basin, Laos, and Red River basin of China and Vietnam (Kottelat, 2001b).
Glyptothorax jalalensis Balon & Hensel, 1970
Glyptothorax jalalensis Balon & Hensel, 1970: 160, fig. 1. Type locality: Kabul River tributary, near Jalal-Abad,
Afghanistan. Holotype: SNMB RY 2176.
Distribution: Kabul River basin, Afghanistan (Balon & Hensel, 1970).
Glyptothorax kashmirensis Hora, 1923
Glyptothorax kashmirensis Hora, 1923b: 22, fig. 2. Type locality: Kashmir Valley. Syntypes: ZSI F10270/1 (2).
Distribution: Jhelum River, Kashmir (Kullander, 1999); NWFP, Azad Kashmir (Mirza, 2003).
Remarks: See Rashida et al. (1996) for comments on the taxonomic position of this species.
Glyptothorax kurdistanicus (Berg, 1931)
Glyptosternum kurdistanicum Berg, 1931: 1267, fig. 1; pl. 1 (fig. 2). Type locality: Kurdistan, at the village of Germau (or Germav), at the height of 1500 m, ... Germau is situated in latitude 36º southeast of Serdesht on theWestern slope of the Sur-kei Range, in the basin of the river Bané, tributary to the Little Zab, which is a tributary to the Tigris R. Holotype: ZIN 20780.

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FERRARISDistribution: Tigris River basin, Iran (Coad, 1995).
Glyptothorax lampris Fowler, 1934
Glyptothorax lampris Fowler, 1934a: 91, figs. 34–36. Type locality: Chieng Mai, North Siam. Holotype: ANSP
59357.
Distribution: Mekong and Chao Phraya River basins, and rivers of southeastern Thailand (Kottelat, 2001b); in
medium-sized upland rivers (Rainboth, 1996).
Glyptothorax laosensis Fowler, 1934
Glyptothorax laosensis Fowler, 1934a: 88, figs. 28–30. Type locality: Bua Yai, East Siam. Holotype: ANSP 59412.
Distribution: Mekong and Chao Phraya River basins (Kottelat, 2001b); in small to medium-sized streams (Rainboth,
1996).
Glyptothorax lonah (Sykes, 1839)
Bagrus lonah Sykes, 1839: 164. Type locality: Deccan, India. Holotype: BMNH 1860.3.19.756 or BMNH
1860.3.19.757. Also described as new in Sykes (1840: 61); illustrated and described in more detail in Sykes(1841: 371).
Glyptosternum dekkanense Günther, 1864: 187. Type locality: Dekkan. Holotype: BMNH 1860.3.19.757.
Distribution: Deccan, Godavari and Krishna River basins, India (Jayaram, 1999).
Remarks: Holotype of Bagrus lonah Sykes has not been clearly established; see Günther (1864: 187), and Day(1873d: 748) for comments.
Glyptothorax longicauda Li, 1984
Glyptothorax longicauda Li, 1984b: 82, fig. 3. Type locality: Tengchong, Yunnan, China. Holotype: KIZ 764126.
Distribution: Irrawaddy River basin, China (Chu et al., 1999).
Glyptothorax longjiangensis Mo & Chu, 1986
Glyptothorax longjiangensis Mo & Chu, 1986: 344, 349, fig. 5. Type locality: Longjiang river (upper tributary of
Irrawaddy river), Tengchong County (25º00'N, 98º30'E), Yunnan, China. Holotype: KIZ 764246.
Distribution: Irrawaddy River basin, Yunnan, China (Chu et al., 1999).
Glyptothorax macromaculatus Li, 1984
Glyptothorax macromaculatus Li, 1984b: 82, fig. 7. Type locality: Yangbi Xian, Yunnan, China. Holotype: KIZ
748804.
Distribution: Mekong River basin (Chu et al., 1999; Kottelat, 2001b).
Glyptothorax madraspatanus (Day, 1873)
Glyptosternum madraspatanum Day, 1873b: 526. Type locality: Bowany River, Neilgherries, Madras State, India.
Possible syntypes: AMS B.7759 (1), AMS B.8004 (1), NMW 46582–83 (2), RMNH 2739 (1), ZMB 10798 (1),
ZMB 10829 (1), ZMB 11208 (1), ZSI F1235 (1), ZSI F1313 (1).
Distribution: Western Ghats, Anaimalai Hills, Nilgiri Hills, and Cauvery River (Jayaram, 1999).
Glyptothorax major (Boulenger, 1894)
Akysis major Boulenger, 1894a: 246. Type locality: Senah, Tagora and Baram rivers, Sarawak. Syntypes: BMNH
1892.9.2.59 (1), BMNH 1892.10.7.26 (1), BMNH 1893.3.6.173–177 (5), BMNH 1893.3.6.178 (1, skeleton).
Glyptosternum kükenthali Steindachner, 1901: 448, pl. 18 (figs. 5, 5a). Type locality: Baram-Flusse, Borneo. Holotype: SMF 752.
Distribution: Malay Peninsula and western Borneo (Roberts, 1989a).
Glyptothorax manipurensis Menon, 1955
Glyptothorax manipurensis Menon (A. G. K.), 1955: 23, fig. 1. Type locality: Barak River at Karong, Naga Hills,
Manipur State, India. Holotype: ZSI F738/2.
Distribution: Barak River, Manipur, India (Jayaram, 1999).
Remarks: Redescribed in Kosygin & Vishwanath (2005).
Glyptothorax minimaculatus Li, 1984
Glyptothorax minimaculatus Li, 1984b: 81, fig. 5. Type locality: Tengchong Xian, Yunnan, China. Holotype: KIZ
764336.
Distribution: Irrawaddy River basin, Yunnan, China (Chu et al., 1999).
Glyptothorax minutus Hora, 1921

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Glyptothorax minutus Hora, 1921a: 180, fig. 1. Type locality: Imphal stream near Karong [Kameng], Manipur Valley, India. Syntypes (4): lost (Hora, 1921a: 182).
Distribution: Manipur, India (Hora, 1921a).
Glyptothorax naziri Mirza & Naik, 1969
Glyptothorax naziri Mirza & Naik, 1969: 123, figs. 1–2. Type locality: Zhob River, Baluchistan, Pakistan. Holotype:
GCM 6.
Distribution: NWFP, Punjab, Balochistan, Azad Kashmir, Pakistan (Mirza, 2003).
Glyptothorax nelsoni Ganguly, Datta & Sen, 1972
Glyptothorax nelsoni Ganguly, Datta & Sen, 1972: 341, fig. 1. Type locality: Subarnarekha River, Chotanagpur Plateau, Bihar, India. Holotype: USNM 205611.
Distribution: Subarnarekha River, Bihar, India (Jayaram, 1999).
Glyptothorax nieuwenhuisi (Vaillant, 1902)
Glyptosternon Nieuwenhuisi Vaillant, 1902: 72, 162, figs. 14–15. Type locality: Le Bloeoe, Bornéo central. Syntypes: MNHN 1891-0484 (1), MNHN 1891-0485 (1), MNHN 1891-0486 (1), MNHN 1891-0487 (1), MNHN
1903-0189 (1).
Distribution: Bloeoe River, Borneo.
Glyptothorax obscurus Li, 1984
Glyptothorax obscura Li, 1984b: 86, fig. 1. Type locality: Jingdon Xian, Yunnan, China. Holotype: KIZ 737197.
Distribution: Upper Red River basin, China (Li, 1984b).
Glyptothorax pallozonus (Lin, 1934)
Glyptosternum pallozonum Lin, 1934: 228, figs. 7–8. Type locality: Loh Fau Shan, Poh-lo District, Kwangtung,China. Holotype: Fisheries Experiment Station, Canton G10.
Distribution: Dongjiang, Guandong, China (Chu et al., 1999).
Glyptothorax panda Ferraris & Britz, 2005
Glyptothorax panda Ferraris & Britz, 2005: 376, fig. 1. Type locality: Myanmar, Kachin State: Upper Irrawaddy
River drainage, hill stream 8 miles from Kamaing on road to Tanai, vicinity of Myitkyina. Holotype: USNM
384824.
Distribution: Upper Irrawaddy River basin, Myanmar (Ferraris & Britz, 2005).
Glyptothorax pectinopterus (M’Clelland, 1842)
Glyptosternon pectinopterus M’Clelland, 1842: 587. Type locality: The mountains of Simla. No types known.
Distribution: Ganges, Sutlej and upper Indus River basins, south Asia (Kullander, 1999).
Glyptothorax platypogon (Valenciennes, 1840)
Pimelodus platypogon Valenciennes, in Cuvier & Valenciennes, 1840b: 152 (113 of Strasbourg deluxe edition).
Type locality: Java. Possible syntypes: MNHN b-0196 (2), MNHN 0000-2903 (2), MNHN 0000-2904 (2);RMNH (not found); SMF 649 (2).
Pimelodus cyanochloros Bleeker, 1847b: 11. Type locality: Java. Syntypes: SMNS 10569 (6). Also described as new
in Bleeker (1847a: 168).
Sundagagata robusta Boeseman, 1966: 243, fig. 1. Type locality: River near Buitenzorg, Java. Holotype: RMNH
25264.
Distribution: Java; western Borneo (Roberts, 1989a).
Remarks: Sundagagata robusta Boeseman, 1966, was considered by Kottelat et al. (1993: 76) to be an abnormal
specimen of Glyptothorax platypogon (Valenciennes, 1840).
Glyptothorax platypogonides (Bleeker, 1855)
Pimelodus platypogonides Bleeker, 1855a: 272. Type locality: Lahat, in fluviis. Syntypes (4, 70–85 mm TL): possibly BMNH 1863.12.4.154 (1), RMNH 6912 (4), RMNH 15289 (3).
Callomystax Schmidti Volz, 1904: 470. Type locality: Simbolon Gebirge, cent. Sumatra, Indonesia, elev. 1400 m.
Syntypes (5): MHNG 683.22 (1), NMBA 2827 (1).
Distribution: Sumatra (Ng & Rachmatika, 2005).
Remarks: Register number of syntype of Pimelodus platypogonides incorrectly reported as BMNH 1863.12.11.154
in Eschmeyer et al. (1998). Name of species frequently cited as Glyptothorax platypogonoides.

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FERRARISGlyptothorax prashadi Mukerji, 1932
Glyptothorax prashadi Mukerji, 1932: 281, fig. 1. Type locality: Kyenchaung, Mergui District, Lower Burma. Holotype: ZSI F11334/1.
Distribution: Southeastern Myanmar and Peninsular Thailand (Jayaram, 1999).
Glyptothorax punjabensis Mirza & Kashmiri, 1971
Glyptothorax conirostris punjabensis Mirza & Kashmiri, 1971: 88, fig. 1. Type locality: Rawal Dam, Rawalpindi,
Pakistan. Holotype: GCM 7.
Distribution: Soan River, Jhelum canal, Kabul River, and Siran River, Pakistan, and Kotli, Azad Kashmir (Jayaram,
1999).
Remarks: Treated as a possible synonym of Glyptothorax kashmirensis Hora, 1923, by Kullander (1999).
Glyptothorax quadriocellatus (Mai, 1978)
Glyptosternon quadriocellatum Mai, 1978: 272, fig. 121. Type locality: Fast-running creeks, n. Vietnam. possibly at
DVZUT.
Glyptosternon minutum Mai, 1978: 274, fig. 122. Type locality: Fast-running creeks, n. Vietnam. Type (s): at
DVZUT. Preoccupied in Glyptothorax by Glyptothorax minutus Hora, 1921; replaced by Glyptothorax spectrum Kottelat, 2001.
Glyptothorax spectrum Kottelat, 2001a: 55. Type locality: Fast-running creeks, n. Vietnam. Type (s): at DVZUT.
Replacement name for Glyptosternon minutum Mai, 1978, preoccupied in Glyptothorax by Glyptothorax minutus Hora, 1921.
Distribution: Yuanjiang, China (Chu et al., 1999); northern Viet Nam (Kottelat, 2001a).
Remarks: Kottelat (2001a: 54) synonymized G. quadriocellatum and G. minutum and selected G. quadriocellatum as
the valid name.
Glyptothorax saisii (Jenkins, 1910)
Glyptosternum saisii Jenkins, 1910: 128, fig.; pl. 6 (fig. 6). Type locality: Sita Nullah stream, Paresnath Hills, Bihar,
w. Bengal, elev. 2150 ft. Holotype: ZSI F2583/1.
Distribution: Bihar, Uttar Pradesh, and Maharashtra, India (Jayaram, 1999).
Remarks: Redescribed in Tilak & Husain (1978).
Glyptothorax siamensis Hora, 1923
Glyptothorax siamensis Hora, 1923a: 168, pl. 12 (figs. 1–3). Type locality: Nakon Sritamarat Hills, Siam. Holotype:ZSI F10548/1.
Distribution: Thailand and Peninsular Malaysia (Ng & Rachimatika, 2005).
Glyptothorax silviae Coad, 1981
Glyptothorax silviae Coad, 1981a: 291, figs. 1–3. Type locality: Iran, Khuzestan, stream 3 km south of Bagh-e
Malek, tributary to Rud-e Zard or Ab-e Ala in the drainage of the Jarrahi River, 31º29'N, 49º54'30"E, elev. 660
m. Holotype: NMC 79-0390A.
Distribution: Tigris River and possibly Gulf basins (Coad, 1995).
Glyptothorax sinensis (Regan, 1908)
Glyptosternum sinense Regan, 1908a: 110, pl. 4 (fig. 3). Type locality: Tunting [=Tungting], China. Holotype:
BMNH 1907.11.26.4.
Distribution: lower and middle Changjiang [= Yangtze River], China (Chu et al., 1999); also reported from Manipur,
India (Kosygin & Vishwanath, 2005) .
Remarks: Redescribed as Glyptothorax sinense in Xie et al. (2001).
Glyptothorax steindachneri (Pietschmann, 1913)
Glyptosternum steindachneri Pietschmann, 1913a: 93. Type locality: dem Tigris, Mosul. Syntypes (2): at NMW.
Distribution: Tigris River basin (Coad, 1995).
Glyptothorax stocki Mirza & Nijssen, 1978
Glyptothorax stocki Mirza & Nijssen, 1978: 79, fig. 1. Type locality: Bhed Nullah, small stream on G.T. road to
Rawalpindi, 7 mi. from Lahore, Pakistan. Holotype: ZMA 114763.
Distribution: NWFP, Punjab, Azad Kashmir, Pakistan (Mirza, 2003).
Glyptothorax stolickae (Steindachner, 1867)

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Glyptosternum Stoličkae Steindachner, 1867e: 533, pl. 6 (fig. 1). Type locality: Simla. Syntypes: NMW 76606 (3).
Distribution: Western Himalaya, Shimla, Ganga, India (Jayaram, 1999).
Glyptothorax striatus (M’Clelland, 1842)
Glyptosternon Striatus M’Clelland, 1842: 587, pl. 6 (in part). Type locality: Kasyah mountains. Syntypes (3):
BMNH 1860.3.19.95 (1, 78 mm SL).
Distribution: Kashi and Garro Hills, India (Jayaram, 1999).
Glyptothorax sufii Bashir & Mirza, 1975
Glyptothorax telchitta sufii Bashir & Mirza, 1975: 96, fig. 1. Type locality: River Sutlej, Lahore District, Pakistan.
Holotype: GCM F-13.
Distribution: Indus River basin, Pakistan (Bashir & Mirza, 1975).
Glyptothorax sykesi (Day, 1873)
Glyptosternum sykesi Day, 1873d: 748. Type locality: Deccan, India. Holotype: BMNH 1860.3.19.756. Name proposed conditionally for specimen in Günther (1864: 187) identified as Glyptosternum lonah, which Day contended was not the holotype and, in fact, not even conspecific with G. lonah.
Distribution: Peninsular India.
Glyptothorax telchitta (Hamilton, 1822)
Pimelodus telchitta Hamilton, 1822: 185, 378. Type locality: Fresh water rivers of Bengal and Behar [now: India:
west Bengal: Hooghly River at Kalna, 23º13'30.0"N, 88º22'39.0"E, by neotype designation]. Neotype: UMMZ
244946; designated by, and illustrated in, Ng (2005e:10, fig. 5). Unpublished Hamilton illustration reproducedin Hora (1929: pl. 21, fig. 3). Name spelled Pimelodus telchita on p. 378; first reviser may be Eschmeyer et al.
(1998), but not yet clearly established to be earliest.
Distribution: Widely distributed in India (Jayaram, 1999).
Glyptothorax trewavasae Hora, 1938
Glyptothorax trewavasae Hora, 1938d: 373, pl. 7 (figs. 3–4). Type locality: Yenna Valley, Satara dist., Maharashtra,
India. Holotype: ZSI F9723/1.
Distribution: Krishna River basin, Yenna, Kayana, and Tunga Rivers, India (Jayaram, 1999).
Glyptothorax trilineatus Blyth, 1860
Glyptothorax trilineatus Blyth, 1860: 154. Type locality: Tenasserim [Burma]. Syntypes: ZSI F10380/1 (2).
Glyptothorax trilineatoides Li, 1984b: 87, fig. 5. Type locality: Tengchong Xian, Yunnan, China. Holotype: KIZ
764336.
Distribution: Nujiang [=Salween River], Irrawaddy River (Chu et al., 1999).
Glyptothorax ventrolineatus Vishwanath & Linthoingambi, 2005
Glyptothorax ventrolineatus Vishwanath & Linthoingambi, 2005: 201, fig. 1. Type locality: Iril river, Ukhrul district, Manipur, India. Holotype: MUMFL022I.
Distribution: Iril River, Chindwin River basin, Manipur, India (Vishwanath & Linthoingambi, 2005).
Glyptothorax zanaensis Wu, He & Chu, 1981
Glyptothorax zanaensis Wu, He & Chu, 1981: 74, fig. 1 (1-b). Type locality: Qinghai-Xizang plateau region, China.
Syntypes: IHASW 606164–66 (3), IHASW 606168 (1), IHASW 606170–74 (5), IHASW 606178 (1), IHASW
6006651–53 (3).
Glyptothorax longinema Li, 1984b: 81, fig. 6. Type locality: Lushui Xian, Yunnan, China. Holotype: KIZ 741097.
Glyptothorax rubermentus Li, 1984b: 83, 88, fig. 8. Type locality: Baoshan Xian, Yunnan, China. Holotype: KIZ
749356.
Distribution: Mekong and Salween River basins, China (Chu et al., 1999, as Glyptothorax zainaensis; Kottelat,
2001b).
Glyptothorax zhujiangensis Lin, 2003
Glyptothorax zhujiangensis Lin, 2003: 159, 161. Type locality: Baishuidai Streamin Yaxi Town (22º15'N,
112º59'25"E), Xinhui City, middle-southern Guandong, China. Holotype: ZX 970901 (housed in IHASW).
Distribution: Zhujiang [= Pearl River] China (Lin, 2003).
Species inquirendae, Glyptothorax

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FERRARISGlyptosternum laak Popta, 1904: 190. Type locality: le Howong, Bornéo central. Syntypes: RMNH 7562 (3); describedin more detail, with an illustration of one of the syntypes, in Popta (1906: 71, pl. 4, figs. 15a, 15b).
Glyptosternum tiong Popta, 1904: 191. Type locality: le Kajan, Bornéo central. Syntypes: RMNH 7564 (2); described in
more detail, with an illustration of one of the syntypes in Popta (1906: 75, pl. 4, fig. 16).
Glyptothorax conirostre poonaensis Hora, 1938d: 368, pl. 7 (figs. 5–6). Type locality: Mula Mutha R. at Poona, Maharashtra, Bombay State, India. Holotype: ZSI F12126/1. Originally as Glyptothorax conirostre var. poonaensis.
Treated as valid or as a synonym of Glyptothorax conirostris (Steindachner, 1867).
GOGANGRA Roberts, 2001
Gangra Roberts & Ferraris, 1998: 333. Type species: Pimelodus viridescens Hamilton, 1822. Type by original designation. Gender: Feminine. Preoccupied by Gangra Walker, 1862; in Lepidoptera, replaced by Gogangra Roberts, 2001.
Gogangra Roberts, 2001: 83. Type species: Pimelodus viridescens Hamilton, 1822. Type by being a replacement
name. Gender: Feminine. Replacement for Gangra Roberts & Ferraris, 1998; preoccupied by Gangra Walker,
1862, in Lepidoptera.
Gogangra laevis Ng, 2005
Gogangra laevis Ng, 2005c: 280, fig. 1. Type locality: Bangladesh: Gowain River and Khal at Gowainghat. Holotype: UMMZ 244603.
Distribution: Yamuna and Meghna Rivers, lower Brahmaputra River basin, Bangladesh (Ng, 2005c).
Gogangra viridescens (Hamilton, 1822)
Pimelodus viridescens Hamilton, 1822: 173, 377, pl. 11 (fig. 56). Type locality: Rivers in the northern parts of Bengal. No types known.
Nangra punctata Day, 1877 (in Day, 1875–78): 494, pl. 115 (fig. 8). Type locality: Sone River at Bheer Bhoom in
Bengal, India. Syntype: AMS B.7566 (1).
Distribution: Ganges and Brahmaputra River basins (Roberts & Ferraris, 1998; Ng, 2005c).
HARA Blyth, 1860
Hara Blyth, 1860: 152. Type species: “Hara buchanani nobis; Pimelodus hara, B.H.” [= Pimelodus hara Hamilton,
1822]. Type by original designation. Gender: Feminine.
Laguvia Hora, 1921b: 739. Type species: Pimelodus asperus M’Clelland, 1844. Type by subsequent designation, by
Jordan (1923: 148). Gender: Feminine.
Remarks: Jayaram (1973c, 1999) lists Laguvia shawi as the type species of Laguvia, by subsequent designation, but
it is not stated where this designation was published. If published before 1923, it would have priority over thedesignation by Jordan (1923). Therefore, the type species as stated above must be considered tentative until thedetails of the designation of L. shawi are clarified. Synonymy of Laguvia in Hara follows Ng & Kottelat (2005:
20).
Hara filamentosa Blyth, 1860
Hara filamentosa Blyth, 1860: 152. Type locality: Tenasserim [Burma]. Syntypes: ZSI 585 (6).
Distribution: Widely distributed in Myanmar.
Hara hara (Hamilton, 1822)
Pimelodus hara Hamilton, 1822: 190, 378. Type locality: River Kosi [now Hooghly River south of Ranaghat, by
neotype designation]. Neotype: UMMZ 244697, designated by, and illustrated in, Ng & Kottelat (2005: 20, fig.
5). Previously unpublished Hamilton illustration reproduced in Hora (1929: pl. 21, fig. 5) and reproduced in Ng& Kottelat (2005: 20, fig. 6).
Pimelodus asperus M’Clelland, 1844a: 404, pl. 24 (fig. 2). Type locality: Chusan, China [now Hooghly River southof Ranaghat, by neotype designation]. Neotype: UMMZ 244697, designated by, and illustrated in, Ng & Kottelat (2005: 19, fig. 5).
Hara buchanani Blyth, 1860: 152. Type locality: River Kosi [now Hooghly River south of Ranaghat, by neotype
designation]. Neotype: UMMZ 244697, designated by, and illustrated in, Ng & Kottelat (2005: 20, fig. 5) as

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neotype of Pimelodus hara. Unneeded new name for Pimelodus hara Hamilton, 1822.
Distribution: Bihar, Uttar Pradesh, Orissa, northern Bengal and Assam, India; Bangladesh and Myanmar [questionable] (Jayaram, 1979).
Remarks: Redescribed in Tilak (1978).
Hara horai Misra, 1976
Hara horai Misra, 1976: 245, pl. 9 (figs. 1–3). Type locality: Terai and Duars, n. Bengal. Syntype: ZSI FF955 (formerly ZSI F 11390/1), illustrated in Hora (1950: pl. 2, figs. 1–3) as Hara hara.
Distribution: Terai Duars, northern Bengal, India (Misra, 1976).
Remarks: Neotype designation by Tilak & Talwar (1976: 246) [ZSI FF955] invalid inasmuch as that specimen isapparently one of the original syntypes (M. Kottelat, pers. commum.). Redescribed in Tilak (1978).
Hara jerdoni Day, 1870
Hara jerdoni Day, 1870a: 39, pl. 4 (figs. 2a–c). Type locality: Sylhet district. Syntypes (2): ZSI 431 (1).
Distribution: Sylhet district, Bangladesh (Jayaram, 1979).
Remarks: Redescribed in Husain & Tilak (1978).
Hara saharsai Datta Munshi & Srivastava, 1988
Hara saharsai Datta Munshi & Srivastava, 1988: 265, fig. 5. Type locality: Kosi Belt, n. Bihar, India. Syntypes:
Bhagalpur Univ., Dept. Zool. Mus. 67 (13). Also appeared as new in Srivastava (1989: 121, fig. 1).
Distribution: northern Bihar, India (Datta Munshi & Srivastava, 1988).
Hara serrata Vishwanath & Kosygin, 2000
Hara serratus Vishwanath & Kosygin, 2000: 143, figs. 2–5. Type locality: Jiri river at Jiribam, Manipur, India.
Holotype: MUMF 2508.
Distribution: Barak and Jiri Rivers, Manipur, India (Vishwanath & Kosygin, 2000).
MYERSGLANIS Hora & Silas, 1952
Myersglanis Hora & Silas, 1952a: 19. Type species: Exostoma blythii Day, 1870. Type by original designation. Gender: Masculine.
Key: Vishwanath & Kosygin (1999: 296).
Myersglanis blythii (Day, 1870)
Exostoma Blythii Day, 1870b: 525. Type locality: not stated. Syntypes (2): Asiatic Society of Bengal 599 (2) [now at
ZSI] (2), possibly ZSI 2361 (1).
Distribution: Possibly Pharping, Nepal (Regan, 1907c; Hora & Silas, 1952a).
Myersglanis jayarami Vishwanath & Kosygin, 1999
Myersglanis jayarami Vishwanath & Kosygin, 1999: 291, pl. 1. Type locality: Laniye river at Jessami, Manipur,
India (94º32'E, 25º38'N). Holotype: MUMF 2138.
Distribution: Lainye River, Chindwin River basin, Manipur, India (Vishwanath & Kosygin, 1999).
NANGRA Day, 1877
Nangra Day, 1877 (in Day, 1875–78): 493. Type species: Pimelodus nangra Hamilton, 1822. Type by absolute tautonymy. Gender: Feminine.
Revision: Hora & Law (1941); Roberts & Ferraris (1998).
Remarks: See Jayaram (1972a) for comments on the validity of the genus.
Nangra assamensis Sen & Biswas, 1994
Nangra assamensis Sen & Biswas, 1994: 441, fig. 1; pl. 1. Type locality: Brahmaputra River at Neematighat, 14
kms. from Jorhat, Assam [India]. Holotype: ZSI V/F/ERS/53.
Nangra carcharhinoides Roberts & Ferraris, 1998: 338, fig. 16. Type locality: Ganges River at Patna, India. Holotype: CAS 95566.
Distribution: Ganges and Brahmaputra Rivers, India (Roberts & Ferraris, 1998).
Nangra bucculenta Roberts & Ferraris, 1998
Nangra bucculenta Roberts & Ferraris, 1998: 336, fig. 14. Type locality: Ganges River delta, Tangail District, North
Central Region, Bangladesh. Holotype: CAS 95564.

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Nangra nangra (Hamilton, 1822)
Pimelodus nangra Hamilton, 1822: 193, 378, pl. 11 (fig. 63). Type locality: Kosi River [now: Ganges River at Patna,
India, by neotype designation]. Neotype: CAS 96626, designated by Roberts & Ferraris (1998: 341).
Nangra Buchanani Day, 1877 (in Day, 1875–78): 494, pl. 113 (fig. 3). Kosi River [now: Ganges River at Patna,
India, by neotype designation]. Neotype: CAS 96626. Replacement for Pimelodus nangra Hamilton, 1822, toavoid tautomymy.
Distribution: Ganges and Indus River basins (Roberts & Ferraris, 1998).
Nangra ornata Roberts & Ferraris, 1998
Nangra ornata Roberts & Ferraris, 1998: 341, fig. 19. Type locality: Bangladesh: Gowain River and Khal at
Gowainghat, northern Sylhet Province (Surma or Meghna watershed). Holotype: UMMZ 233236.
Distribution: Surma or Meghna watershed, Bangladesh (Roberts & Ferraris, 1998).
Nangra robusta Mirza & Awan, 1973
Nangra robusta Mirza & Awan, 1973: 145, fig. 1. Type locality: Indus River at Jinnah Barrage near Kalabagh, Pakistan. Holotype: GCM 9.
Distribution: Indus River, Pakistan (Mirza & Awan, 1973).
OREOGLANIS Smith, 1933
Oreoglanis Smith, 1933: 70. Type species: Oreoglanis siamensis Smith, 1933. Type by original designation. Gender:
Masculine.
Paroreoglanis Pellegrin, 1936: 244. Type species: Paroreoglanis delacouri Pellegrin, 1936. Type by monotypy.
Gender: Masculine.
Oreoglanis delacouri (Pellegrin, 1936)
Paroreoglanis delacouri Pellegrin, 1936b: 244. Type locality: Xieng Khouang in upper Laos. Lectotype: MNHN
1936-0031 (103.4 mm SL), designated by Ng & Kottelat (1999: 379).
Distribution: Nam Ngiap basin, Mekong drainage, Laos (Ng & Rainboth, 2001).
Oreoglanis frenatus Ng & Rainboth, 2001
Oreoglanis frenatus Ng & Rainboth, 2001: 7, fig. 6. Type locality: Laos: Xieng Khouang Province, Nam Ka basin,
Houay Kheua at Hwy 7 bridge, 19º38'N, 103º28'E. Holotype: UMMZ 236811.
Distribution: Nam Ka Basin, Mekong drainage, Laos (Ng & Rainboth, 2001).
Oreoglanis hypsiurus Ng & Kottelat, 1999
Oreoglanis hypsiurus Ng & Kottelat, 1999: 376, fig. 1. Type locality: Upper Nam Theun, ca. 1 km upstream of confluence with Houay Nuok Lan, 18º04'09"N, 105º29'44"E, Kham-mouan Province, Laos. Holotype: ZRC 40440.
Distribution: Nam Kading basin, Mekong drainage, Laos (Kottelat, 2001b).
Oreoglanis infulatus Ng & Freyhof, 2001
Oreoglanis infulatus Ng & Freyhof, 2001b: 1165, figs. 1–3. Type locality: Vietnam: Ha Tinh Province, stream at Son
Kim, a tributary of Song Lam (18º24'25"N, 105º11'10"E). Holotype: ZFMK 35719.
Distribution: Lam River headwaters, central Vietnam (Ng & Freyhof, 2001b).
Oreoglanis insignis Ng & Rainboth, 2001
Oreoglanis insignis Ng & Rainboth, 2001: 15, fig. 8. Type locality: China: Yunnan Province, Baoshan Prefecture,
Longchuanjiang and Dajiang, near Qushi. Holotype: KIZ 9810191.
Distribution: Upper Irrawaddy River basin, China and Myanmar, possibly upper Salween basin (Ng & Rainboth,2001).
Oreoglanis lepturus Ng & Rainboth, 2001
Oreoglanis lepturus Ng & Rainboth, 2001: 17, fig. 10. Type locality: Laos: Bolikamsai Province, Nam Phao about 2
km from Vietnam border, 18º23'N, 105º19'E. Holotype: UMMZ 236814.
Distribution: Nam Phao basin, Mekong drainage, northeastern Laos (Ng & Rainboth, 2001).
Oreoglanis macronemus Ng, 2004
Oreoglanis macronemus Ng, 2004e: 209, fig. 1. Type locality: Laos: Xieng Khouang. Holotype: BMNH
1933.8.19.51.

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Distribution: Nam Ngiap or Nam Sen River basins, Xieng Khouang region, Laos (Ng, 2004e).
Oreoglanis macropterus (Vinciguerra, 1890)
Exostoma macropterum Vinciguerra, 1890: 253, pl. 8 (fig. 5). Type locality: Paese dei Catcin, Birmania [Myanmar].
Syntypes: BMNH 1893.2.16.18 (1), MSNG 14410 (3), NMW 46488 (1), RMNH 10236 (1), USNM 44805 (1)Distribution: Irrawaddy River basin; Salween River, China (Ng & Rainboth, 2001).
Remarks: See Steinitz (1961) for comments on taxonomy.
Oreoglanis setiger Ng & Rainboth, 2001
Oreoglanis setiger Ng & Rainboth, 2001: 23, fig. 12. Type locality: Laos: Louang Namtha Province, Nam Ma Oun,
21º05'N, 101º04'E. Holotype: UMMZ 236813.
Distribution: Nam Oun basin, Mekong River drainage, Laos, and upper Mekong River southwestern China (Ng &Rainboth, 2001).
Oreoglanis siamensis Smith, 1933
Oreoglanis siamensis Smith, 1933: 72, pl. 3 (figs. 1–2); fig. 4. Type locality: Kang River near the base of Doi
Angka, northern Siam. Holotype: KUMF 0172.
Distribution: Mae Nam Ping basin, Chao Phraya drainage, northern Thailand (Ng & Rainboth, 2001).
Remarks: Statements in Menon & Yazdani (1968) and Eschmeyer et al. (1998) which indicated that ZSI F12233/1
might be the holotype of Oreoglanis siamensis are in error. Smith (1933: 74) clearly stated that two specimens
were examined, which he identified as the “type” and “topotype,” and that the specimen deposited in the IndianMuseum (now ZSI) was the topotype.
PARACHILOGLANIS Wu, He & Chu, 1981
Parachiloglanis Wu, He & Chu, 1981: 76, 79. Glyptosternum hodgarti Hora, 1923. Type by original designation.
Gender: Masculine.
Parachiloglanis hodgarti (Hora, 1923)
Glyptosternum hodgarti Hora, 1923b: 38, pl. 2 (figs. 1–3). Type locality: Pharping, Nepal. Holotype: ZSI F1553/1.
Distribution: Yaluzangbujiang [= upper Brahmaputra River] (Chu et al., 1999); Abor Hills, Meghalaya, Darjeeling,
Kali River, Teesta valley and Nepal (as Euchiloglanis hodgarti) (Jayaram, 1999).
PAREUCHILOGLANIS Pellegrin, 1936
Pareuchiloglanis Pellegrin, 1936b: 245. Type species: Pareuchiloglanis poilanei Pellegrin, 1936. Type by monotypy. Gender: Masculine.
Remarks: Included species follows He (1996b).
Review: Ding (2003, western China).
Pareuchiloglanis anteanalis Fang, Xu & Cui, 1984
Pareuchiloglanis anteanalis Fang, Xu & Cui, 1984: 209, fig.1. Type locality: Wudu County, Gansu Province, China.
Holotype: Shaanxi Inst. Zool. 82VI9565.
Distribution: Wudu County, Gansu Province, China (Fang, et al., 1984).
Pareuchiloglanis feae (Vinciguerra, 1890)
Exostoma Feae Vinciguerra, 1890: 256, pl. 8 (fig. 6). Type locality: Taó, e Iadó, Birmania [Myanmar]. Syntypes:
BMNH 1893.2.16.19–20 (2), BMNH 1893.2.16.21–22 (2), BMNH 1893.2.16.23 (1, skeleton), MSNG 14411(4), MSNG 14412 (3), MSNM 38 (1), NMW 9064–65 (2), RMNH 10237 (3), USNM 44743 (3).
Distribution: Irrawaddy and Salween Rivers, China (Chu et al., 1999).
Pareuchiloglanis gongshanensis Chu, 1981
Pareuchiloglanis gongshanensis Chu, 1981: 28, 31, fig. 2. Type locality: Gongshan Xian, Yunnan, China. Holotype:
KIZ 731202.
Distribution: Upper Nujiang [= Salween River], China (Chu et al., 1999).
Pareuchiloglanis gracilicaudatus (Wu & Chen, 1979)
Euchiloglanis gracilicaudata Wu & Chen, 1979: 294, fig. 4. Type locality: Za-Qu, upper Lancang Jiang, Nangqen,
Qinghai, China. Syntypes: Qinghai Inst. Biol. (12).
Distribution: Lancangjiang [= Mekong River], China (Chu et al., 1999).

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Pareuchiloglanis kamengensis (Jayaram, 1966)
Euchiloglanis kamengensis Jayaram, 1966d: 85, fig. 1. Type locality: Norgum River at Kalaktang, 1370 m alt.,
Kameng Frontier Division, N.E.F.A. Holotype: ZSI F2105/2.
Distribution: Yaluzangbujiang [= upper Brahmaputra River], Irrawaddy, Nujiang [= Salween River], Lancangjiang[= Mekong River], China (Chu et al., 1999).
Pareuchiloglanis longicauda (Yue, 1981)
Euchiloglanis longicauda Yue, in Zheng, 1981: 183, fig. 151. Type locality: Panyang R. (Hong-Shui R. system),
Bama Co., Guangxi Prov., China. Holotype: ASIZB and Fish. Res. Inst., Guangxi Zhuangzu Autonomous
Region 750443.
Distribution: Nanpanjiang, Beipaniang, Hongshuihe, China (Chu et al., 1999).
Pareuchiloglanis macropterus Ng, 2004
Pareuchiloglanis macropterus Ng, 2004g: 3, fig. 1. Type locality: China, Yunnan province, Salween River (Nujiang)
drainage; Laowo River, a tributary of Salween River (Nujiang), ca. 5 km to Liuku on Yongping–Liuku Road,
25º50'18.6"N 98º53'46.8"E; 900 m asl. Holotype: ZRC 49124.
Distribution: Salween and Irrawaddy River basins, southern China (Ng, 2004g).
Pareuchiloglanis macrotremus (Norman, 1925)
Euchiloglanis macrotrema Norman, 1925b: 570. Type locality: Ngoi-Tio, Col des Nuages, Tonkin, Vietnam, elev.
4500–6500 ft. Syntypes: BMNH 1925.2.19.5–6 (2).
Distribution: Yuanjiang, China (Chu et al., 1999); Tonkin, Vietnam.
Pareuchiloglanis myzostomus (Norman, 1923)
Euchiloglanis myzostoma Norman, 1923a: 562. Type locality: Yunnan, China. Syntypes: BMNH 1923.2.21.40–49(10).
Distribution: Lancangjiang [= Mekong River], China (Chu et al., 1999).
Pareuchiloglanis nebulifer Ng & Kottelat, 2000
Pareuchiloglanis nebulifer Ng & Kottelat, 2000b: 11, fig. 4. Type locality: Small creek, trib. of Houai Siam,
upstream of Ban Kangpabong, Houaphan Prov., Laos, 20º19'36"N, 104º25'01"E. Holotype: ZRC 45706.
Distribution: Nam Xam and Nam Ma basins, Ma River Drainage, Laos (Kottelat, 2001b).
Pareuchiloglanis poilanei Pellegrin, 1936
Pareuchiloglanis poilanei Pellegrin, 1936b: 246. Type locality: Annam [Vietnam]. Syntypes: MNHN 1936-0005(1), MNHN 1936-0006 (1), MNHN 1936-0007 (1), MNHN 1936-0008 (1), MNHN 1936-0009 (1), MNHN
1936-0010 (1), MNHN 1936-0011 (1), MNHN 1936-0012 (1), MNHN 1936-0013 (1), MNHN 1936-0014 (1),
MNHN 1936-0015 (1), MNHN 1935-0016 (1), MNHN 1936-0017 (1), MNHN 1936-0018 (1), MNHN 19360019 (1), MNHN 1936-0020 (4).
Distribution: Central Vietnam.
Pareuchiloglanis rhabdurus Ng, 2004
Pareuchiloglanis rhabdurus Ng, 2004g: 7, fig. 3. Type locality: Vietnam: Ha Giang province, Red River (SongHong) drainage, Vi Xuyen district, Cao Bo stream (Bac Trao river) near camp 1, 22º45'18.0"N, 104º52'11'4"E.
Holotype: AMNH 211153.
Distribution: Song Lo River, Red River basin, northern Vietnam (Ng, 2004g).
Pareuchiloglanis robustus Ding, Fu & Ye, 1991
Pareuchiloglanis robusta Ding, Fu & Ye, 1991: 369, fig. 1. Type locality: a stream (Qing Yijing), Sichuan, China.
Holotype: SPNRI 8801.
Distribution: Yangtze River basin, China (Ding et al., 1991).
Pareuchiloglanis sichuanensis Ding, Fu & Ye, 1991
Pareuchiloglanis sichuanensis Ding, Fu & Ye, 1991: 371, fig. 2. Type locality: Baoxing, Sichuan Province, China.
Holotype: Sichuan Agric. Univ. 6-1481.
Distribution: Yangtze River basin, China (Ding et al., 1991).
Pareuchiloglanis sinensis (Hora & Silas, 1952)
Euchiloglanis sinensis Hora & Silas, 1952a: 17, fig. 2. Type locality: presumably .... from some part of China, possibly Yunnan. Holotype: ZSI F12208/1.

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Distribution: Jinshajiang, China (Chu et al., 1999).
Pareuchiloglanis songdaensis Nguyen & Nguyen, 2001
Pareuchiloglanis songdaensis Nguyen & Nguyen, 2001: 67, fig. 1. Type locality:[Da River, Muong Te, Lai Chau
Town and Phong Tho, Vietnam]. Holotype (115 mm SL): LM 2001.01 H.
Distribution: Da River, Vietnam (Nguyen & Nguyen, 2001).
Pareuchiloglanis songmaensis Nguyen & Nguyen, 2001
Pareuchiloglanis songmaensis Nguyen & Nguyen, 2001: 68, fig. 2. Type locality: [Ma River, Song Ma district, Son
La Province, Vietnam]. Holotype (82 mm SL): LM 2001.02H.
Distribution: Ma River, Vietnam (Nguyen & Nguyen, 2001).
Pareuchiloglanis tianquanensis Ding & Fang, 1997
Pareuchiloglanis tianquanensis Ding & Fang, 1997: 17, fig. 1. Type locality: a stream (Upper Qing Yi River), altitude 2010 m., Tianquan County, Sichuan Province [China]. Holotype: SPNRI 920185.
Distribution: Upper Qing Yi River basin, Sichuan, China (Ding & Fang, 1997).
PSEUDECHENEIS Blyth, 1860
Pseudecheneis Blyth, 1860: 154. Type species: Glyptosternon sulcatus M’Clelland, 1842. Type by monotypy. Gender: Feminine.
Parapseudecheneis Hora, in Hora & Chabanaud, 1930: 216. Type species: Pseudecheneis paviei Vaillant, 1892.
Type by monotypy. Gender: Feminine.
Propseudecheneis Hora, 1937d: 348.Type species: Propseudecheneis tchangi Hora, 1937. Type by original designation. Gender: Feminine.
Review: Hora (1952); Hora & Chabanaud (1930); Chu (1982), China.
Phylogeny: Chu (1982), Zhou & Zhou (2005).
Pseudecheneis crassicauda Ng & Edds, 2005
Pseudecheneis crassicauda Ng & Edds, 2005b: 2, fig. 1. Type locality: Nepal: Mewa Khola (River), Dhakuta District, 27º0'N, 87º20'E. Holotype: BMNH 1958.9.1.8.
Distribution: Tamur River basin, Kosi drainage, Nepal (Ng & Edds, 2005b).
Pseudecheneis immaculata Chu, 1982
Pseudecheneis immaculatus Chu, 1982: 428, 447, fig. 1. Type locality: Baijixun (upper Lancang River), Weixi
County, Yunnan, China. Holotype: KIZ 748742.
Distribution: upper Langcanjiang [= Mekong River], China (Chu et al., 1999).
Pseudecheneis paviei Vaillant, 1892
Pseudecheneis paviei Vaillant, 1892a: 126. Type locality: Near Lai Chau, Tonkin, northern Vietnam. Holotype:
MNHN 1892-0049; redescribed and illustrated in Vaillant (1904: 464, pl. 22, fig. 3). Holotype illustrated in
Hora & Chabanaud (1930: 217, fig. 2).
Pseudecheneis intermedius Chu, 1982: 430, 447, fig. 2. Type locality: Dongbao (a stream at upper reaches of BabianRiver, belonging to the Red River drainage), Jingdong County [Yunnan, China]. Holotype: KIZ 737173.
Distribution: Northern Vietnam and Yunnan, China (Kottelat, 2001a).
Remarks: Synonymy follows Ng & Edds (2005b: 12).
Pseudecheneis serracula Ng & Edds, 2005
Pseudecheneis serracula Ng & Edds, 2005b: 6, fig. 3. Type locality: Nepal: Mugu/Bajura, Jhugala, Karnali River,purchased at Jhugala, 29º31'18.0"N, 81º46'48.0"E. Holotype: KU 29554.
Distribution: Karnali and Gandaki River basins, Nepal (Ng & Edds, 2005).
Pseudecheneis sulcata (M’Clelland, 1842)
Glyptosternon sulcatus M’Clelland, 1842: 587, pl. 6 (in part). Type locality: Kasyah Hills. Holotype: BMNH
2005.5.17.5 (76 mm SL).
Distribution: Lancangjiang [ = Mekong River], Nujiang [= Salween River], Yaluzangbujiang [= upper Brahmaputra
River], Irrawaddy River, China (Chu et al., 1999).
Pseudecheneis sulcatoides Zhou & Chu, 1992
Pseudecheneis sulcatoides Zhou & Chu, 1992: 111, 115, figs. 1–3. Type locality: Yangbi (25º40'N, 99º57'E), [Lan-400 · Zootaxa 1418 © 2007 Magnolia Press

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cangjiang River (=upper Mekong)] Yunnan Province, China. Holotype: KIZ 839059.
Distribution: Upper Mekong River basin, Yunnan, China (Zhou & Chu, 1992).
Pseudecheneis sympelvica Roberts, 1998
Pseudecheneis sympelvicus Roberts, 1998: 290, fig. 1. Type locality: Nam Veo, tributary of Nam Phao 25 km east of
Lak Sao, Nam Theun watershed, Mekong Basin, central Laos. Holotype: ZRC 40359.
Distribution: Nam Theun watershed, Mekong River basin, Laos (Roberts, 1998).
Pseudecheneis tchangi Hora, 1937
Propseudecheneis tchangi Hora, 1937d: 348, fig. 11b. Type locality: Yunnan, China. Holotype: Zoological Museumof Fan Memorial Institute, Beijing, 12016.
Distribution: Red River, Yunnan, China (Hora, 1937d).
PSEUDEXOSTOMA Chu, 1979
Pseudexostoma Chu, 1979: 78, 81. Type species: Glyptosternum yunnanensis Tchang, 1935. Type by original designation. Gender: Neuter.
Pseudexostoma brachysoma Chu, 1979
Pseudexostoma yunnanensis brachysoma Chu, 1979: 78, 81, fig. 3. Type locality: Hill stream running into the Nu
River, near Laowo, Yunlong Xian [Yunnan, China]. Holotype: KIZ 742093.
Distribution: Nujiang [= Salween River], China (Chu et al., 1999).
Pseudexostoma yunnanense (Tchang, 1935)
Glyptosternum yunnanensis Tchang, 1935b: 174, fig. 1. Type locality: Yunnan, China. Holotype: ASIZB [=ZMFMIB] 12027.
Distribution: Irrawaddy River, China (Chu et al., 1999).
PSEUDOLAGUVIA Misra, 1976
Pseudolaguvia Misra, 1976: 253. Type species: Glyptothorax tuberculatus Prashad & Mukerji, 1929. Type by original designation. Gender: Feminine.
Remarks: See Jayaram (1973c) for comments on this genus, but as Laguvia.
Pseudolaguvia foveolata Ng, 2005
Pseudolaguvia foveolata Ng, 2005b: 174, fig. 1. Type locality: India: West Bengal: Tista River at Tista barrage,
26º45'10"N, 88º34'11"E. Holotype: UMMZ 244867.
Distribution: Tista River, Brahamputra River basin, India (Ng, 2005b).
Pseudolaguvia inornata Ng, 2005
Pseudolaguvia inornata Ng, 2005g: 36, fig. 1. Type locality: Bangladesh: Chittagong District, Koilla Khal (creek),
10 km E of Feni-Chittagong highway on road to Ramgarh, 22º55'N, 91º36'E. Holotype: UMMZ 245580.
Distribution: Feni River basin, Bangladesh (Ng, 2005g).
Pseudolaguvia kapuri (Tilak & Husain, 1975)
Laguvia ribeiroi kapuri Tilak & Husain, 1975: 1, figs. 1–3. Type locality: Padhoi river, near Kalsia Ghat, Saharanpur, (U. P.) [India]. Holotype: ZSI/NRS 836.
Distribution: Padhoi River and Dhamola River, Saharanpur, Uttar Pradesh (Jayaram, 1999).
Remarks: Redescribed in Gupta (1982).
Pseudolaguvia muricata Ng, 2005
Pseudolaguvia muricata Ng, 2005g: 41, fig. 3. Type locality: Bangladesh: Sylhet District, Rangapani Khal (creek), 6
km WNW of Sylhet-Shillong highway, 25º10'N, 92º6'E. Holotype: UMMZ 245581.
Distribution: Brahmaputra River basin, Bangladesh (Ng, 2005g).
Pseudolaguvia ribeiroi (Hora, 1921)
Laguvia ribeiroi Hora, 1921b: 741, pl. 29 (fig. 3). Type locality: Khoila River, tributary of Teesta at Jalpaiguri[India]. Holotype: ZSI F 10086/1.
Distribution: Teesta River system, north Bengal; Kosi River system Behar; Rihand River system, Uttar Pradesh, andrivers of Nepal (Jayaram, 1979, 1999).
Pseudolaguvia shawi (Hora, 1921)

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Laguvia shawi Hora, 1921b: 740, pl. 29 (fig. 2). Type locality: Mahanadi River below Darjeeling, India. Holotype:ZSI F10085/1.
Distribution: North Bengal, India, and Bangladesh (Jayaram, 1999); Brahmaputra River basin, Assam, India (Arunkumar, 2000).
Pseudolaguvia tenebricosa Britz & Ferraris, 2003
Pseudolaguvia tenebricosa Britz & Ferraris, 2003: 2, fig. 1. Type locality: Myanmar, Kayin Division, Pathe Chaung,hill stream, 13 miles east of Taungoo, 19º01'11"N, 96º35'33"E. Holotype: USNM 373293.
Distribution: Sittang and upper Irrawaddy River basins, Myanmar (Britz & Ferraris, 2003).
Pseudolaguvia tuberculata (Prashad & Mukerji, 1929)
Glyptothorax tuberculatus Prashad & Mukerji, 1929: 182, pl. 7 (fig. 2); fig. 4. Type locality: Sankha, a large hillstream, midway between Kamaing and Mogaung, Myitkyina dist., Upper Burma. Holotype: ZSI F10876/1.
Distribution: Myitkyina District, Kachin State, Myanmar (Prashad & Mukerji, 1929).
SISOR Hamilton, 1822
Sisor Hamilton, 1822: 208, 379. Type species: Sisor rabdophorus Hamilton, 1822. Type by monotypy. Gender: Masculine.
Revision: Ng (2003c).
Sisor barakensis Vishwanath & Darshan, 2005
Sisor barakensis Vishwanath & Darshan, 2005: 1952, fig. 1a. Type locality: River Barak, Jiri, Manipur, India. Holotype: MUMF 3131. Holotype illustrated as Image 1 on web version of paper.
Distribution: Barak River, Brahmaputra River basin, India (Vishwanath & Darshan, 2005).
Sisor chennuah Ng & Lahkar, 2003
Sisor chennuah Ng & Lahkar, in Ng, 2003c: 2876, fig. 5. Type locality: India, Assam State, Brahmaputra Riverdrainage, Dibrugarh. Holotype: NRM 40420.
Distribution: Brahmaputra River basin, India and Bangladesh (Ng, 2003c).
Sisor rabdophorus Hamilton, 1822
Sisor rabdophorus Hamilton, 1822: 208, 379. Type locality: Northern rivers of Bengal and Behar [now India: West
Bengal State, Bhagirathi River at crossing point between Kalna (Barddhaman District) and Nisinghapur (Nadia
District) (23º13'33" N, 88º32'41.4"E), by neotype designation]. Neotype: ZRC 45829, designated by Ng (2003c:
2873, 2881).
Distribution: Ganges River basin, West Bengal, India (Ng, 2003c).
Remarks: Name often misspelled Sisor rhabdophorus, following the incorrect subsequent spelling in Cuvier &Valenciennes (1840).
Sisor rheophilus Ng, 2003
Sisor rheophilus Ng, 2003c: 2877, fig. 6. Type locality: India: Uttar Pradesh State, Kali Nadi River, near Muzaffarnagar. Holotype: UMMZ 189651.
Distribution: Upper and middle Ganges River basin, Bihar State and Uttar Pradesh, India (Ng, 2003c).
Sisor tortosus Ng, 2003
Sisor tortosus Ng, 2003c: 2878, fig. 7. Type locality: India: Bihar State, Ganges River at Patna. Holotype: CAS
96629.
Distribution: Middle Ganges River basin, Bihar and Deli States, India (Ng, 2003c).
Species inquirenda, Sisoridae
Exostoma Oschanini Herzenstein, 1889a: 70. Upper Amu-Darya R., upper Syr-Derya R. basin (Ugam R, Tashkent),
Uzbekistan. Syntypes: BMNH 1888.11.24.1 (1), ZIN 8055 (1), ZIN 8056 (1), ZIN 8057 (2). Also in Herzenstein(1889b:120).

TRICHOMYCTERIDAE Bleeker, 1858

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Eremophilini Bonaparte, 1846: 5. Type genus: Eremophilus Humboldt, 1805.
Trichomycterini Bleeker, 1858b: 49, 250, 257. Type genus: Trichomycterus Valenciennes, 1832.
Vandelliini Bleeker, 1862 (in Bleeker, 1862–63): 17. Type genus: Vandellia Valenciennes, 1846.
Stegophilina Günther, 1864: 5, 12, 276. Type genus: Stegophilus Reinhardt, 1859.
Pareiodontinae Eigenmann, 1918c: 261. Type genus: Pareiodon Kner, 1855.
Tridentinae Eigenmann, 1918c: 275. Type genus: Tridens Eigenmann & Eigenmann, 1889.
Glanapteryginae Myers, 1944: 592. Type genus: Glanapteryx Myers, 1927.
Sarcoglanidinae Myers & Weitzman, 1966: 278. Type genus: Sarcoglanis Myers & Weitzman, 1966.
Trichogeninae Isbrücker, 1986: 276. Type genus: Trichogenes Britski & Ortega, 1983.
Copionodontinae de Pinna, 1992b: 179. Type genus: Copionodon de Pinna, 1992.
Taxonomic summary: de Pinna & Wosiacki (2003).
Biogeography: Arratia (1990).
Revision: Eigenmann (1918a); Tchernavin (1944, Trichomcyterinae).
Phylogeny: Baskin (1973), de Pinna (1989b, Glanapteryginae); de Pinna (1992b); Costa (1994); Bockmann et al. (2004).
Review: Miranda Ribeiro (1946, 1951b, Brazilian Stegophilinae); Miranda Ribeiro (1947, Brazilian Vandelliinae);
Miranda Ribeiro (1949b, 1956, 1957, Brazilian Trichomycterinae).
41 genera, 207 species; no named fossil taxa (but see Propygidium in section below entitled “Names wrongly treated as
Siluriformes”).
ACANTHOPOMA Lütken, 1892
Acanthopoma Lütken, 1892: 57. Type species: Acanthopoma annectens Lütken, 1892. Type by monotypy. Gender:
Neuter.
Acanthopoma annectens Lütken, 1892
Acanthopoma annectens Lütken, 1892: 53, figures on p. 56. Type locality: Huallaga. Holotype: MTD F5245.
Distribution: Upper and middle Amazon River basin, Brazil and Peru (de Pinna & Wosiacki, 2003).
AMMOGLANIS Costa, 1994
Ammoglanis Costa, 1994: 208. Type species: Ammoglanis diaphanus Costa, 1994. Type by original designation.
Gender: Masculine.
Ammoglanis diaphanus Costa, 1994
Ammoglanis diaphanus Costa, 1994: 208, fig. 1. Type locality: Stream trib. to Rio Javaés, Rio Araguaia basin, ca. 40
km north of Araguaçú, 12º24'S, 49º58'W, Tocantin state, Brazil. Holotype: MNRJ 12442.
Distribution: Stream tributary to Javaés River, Araguaia River basin, Brazil (de Pinna & Wosiacki, 2003).
Ammoglanis pulex de Pinna & Winemiller, 2000
Ammoglanis pulex de Pinna & Winemiller, 2000: 257, fig. 1. Type locality: Venezuela: Estado Amazonas: Río Paria
Grande at bridge on road between Pto. Ayacucho and Samariapo (5º23'N 67º37'W). Holotype: MBUCV V29040.
Distribution: Paria Grande River, Pamoni River and Caño Garrapata, Venezuela (de Pinna & Wosiacki, 2003).
APOMATOCEROS Eigenmann, 1922
Apomatoceros Eigenmann, 1922a: 113. Type species: Apomatoceros alleni Eigenmann, 1922. Type by monotypy.
Gender: Masculine.
Apomatoceros alleni Eigenmann, 1922
Apomatoceros alleni Eigenmann, 1922a: 113, pls. 3 (figs. 1–4), 4 (fig. 9). Type locality: Río Morona, upper Amazon
system, Peru. Holotype: CAS 56174.
Distribution: Amazon River basin, Peru (de Pinna & Wosiacki, 2003).
BULLOCKIA Arratia, Chang, Menu-Marque & Rojas, 1978
Bullockia Arratia, Chang, Menu-Marque & Rojas, 1978: 162, 187. Type species: Hatcheria maldonadoi Eigenmann,
1920. Type by monotypy. Gender: Feminine.

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Bullockia maldonadoi (Eigenmann, 1920)
Hatcheria Maldonadoi Eigenmann, 1920c: 53. Type locality: lower course of the Rio Nonguen where it passes
through the ground of the Agricultural School, at the outskirts of Concepción, Chile. Lectotype: CAS 63842.
Illustrated and described in more detail in Eigenmann (1927: 39, pl. 8, figs. 2–2b); with the lectotype established in caption to pl. 8.
Hatcheria bullocki Fowler, 1940b: 180, figs. 14–15. Type locality: Angol, Chile. Holotype: ANSP 69145. Böhlke
(1984: 163) noted that stated locality in the original description is Angol, but catalog and label say “El Vergal.”
Distribution: Chile (de Pinna & Wosiacki, 2003).
Remarks: The description of Hatcheria maldonadoi stated that the Type (= Holotype) was illustrated. However,there was no illustration associated with that publication, so all of the examined specimens need to be treated as
syntypes. The illustrated specimen in Eigenmann (1927) was labeled as the type, which is sufficient to betreated as a lectotype designation.
COPIONODON de Pinna, 1992
Copionodon de Pinna, 1992b: 181. Type species: Copionodon pecten de Pinna, 1992. Type by original designation.
Gender: Masculine.
Copionodon lianae Campanario & de Pinna, 2000
Copionodon lianae Campanario & de Pinna, 2000: 370, figs. 1–2. Type locality: Brazil: Bahia: Rio Grisante (Rio
Mucujê basin), approx. 13º08'S 41º17'W. Holotype: MZUSP 42469.
Distribution: Grisante River, tributary to Mucujê River, Paraguaçu River basin, Brazil (de Pinna & Wosiacki, 2003).
Copionodon orthiocarinatus de Pinna, 1992
Copionodon orthiocarinatus de Pinna, 1992b: 188, figs. 3–4. Type locality: Rio Mucujê, trib. of Rio Paraguaçu,
Bahia, Brazil, ca. 13º00'S, 41º23'W, elev. 1200 m. Holotype: MZUSP 42463.
Distribution: Mucujê River, tributary of Paraguaçu River, Brazil (de Pinna & Wosiacki, 2003).
Copionodon pecten de Pinna, 1992
Copionodon pecten de Pinna, 1992b: 182, figs. 1–2. Type locality: Rio Mucujê, trib. of R. Paraguaçu, Bahia, Brazil,
ca. 13º00'S, 41º23'W, elev. 1200 m. Holotype: MZUSP 42461.
Distribution: Mucujê River, tributary of Paraguaçu River, Brazil (de Pinna & Wosiacki, 2003).
EREMOPHILUS Humboldt, 1805
Eremophilus Humboldt, 1805a: 17. Type species: Eremophilus mutisii Humboldt, 1805. Type by monotypy. Gender:
Masculine.
Thrichomycterus Humboldt, 1805a: 18. Type species: Eremophilus mutisii Humboldt, 1805. Type by monotypy.
Gender: Masculine. Given as an alternative name, but never used subsequently, for Eremophilus Humboldt.
Trachypoma Giebel, 1871: 97. Type species: Trachypoma marmoratum Giebel, 1871. Type by original designation.
Gender: Neuter. Preoccupied by Trachypoma Günther, 1859, in fishes; apparently not replaced.
Eremophilus mutisii Humboldt, 1805
Eremophilus mutisii Humboldt, 1805a: 18, pl. 6. Type locality: petite rivière de Bogota, qui forme la fameuse catarate de Tequendama, Colombia. No types known.
? Trachypoma marmoratum Giebel, 1871: 97. Type locality: obern Amazonenstrom. Type(s): Whereabouts
unknown.
Distribution: Bogotá River basin; probably introduced to Ubaté, Chinquinquirá and Tundama valleys, Colombia (dePinna & Wosiacki, 2003).
GLANAPTERYX Myers, 1927
Glanapteryx Myers, 1927: 128. Type species: Glanapteryx anguilla Myers, 1927. Type by original designation.
Gender: Feminine.
Glanapteryx anguilla Myers, 1927
Glanapteryx anguilla Myers, 1927: 129. Type locality: Rapids of São Gabriel, Rio Negro system, Brazil. Holotype:
CAS 56048.

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FERRARISDistribution: Negro and Orinoco River basins, Brazil and Venezuela (Nico & de Pinna, 1996; de Pinna & Wosiacki,2003).
Remarks: Redescribed in de Pinna (1989b), with notes on distribution in Nico & de Pinna (1996).
Glanapteryx niobium de Pinna, 1998
Glanapteryx niobium de Pinna, 1998b: 36, fig. 1. Type locality: Brazil, State of Amazonas, Pico da Neblina National
Park, Morro dos Seis Lagos (approx. 0º17'N 66º41'W), Lago Esperança. Holotype: INPA 12421.
Distribution: Morro dos Seis Lagos, Negro River basin, Brazil (de Pinna & Wosiacki, 2003).
GLAPHYROPOMA de Pinna, 1992
Glaphyropoma de Pinna, 1992b: 194. Type species: Glaphyropoma rodriguesi de Pinna, 1992. Type by original designation. Gender: Feminine.
Glaphyropoma rodriguesi de Pinna, 1992
Glaphyropoma rodriguesi, 1992b: 196, figs. 7–8. Type locality: Rio Mucujê, trib. of Rio Paraguaçu, Bahia, Brazil,
ca. 13º00'S, 41º23'W, elev. 1200 m. Holotype: MZUSP 42465.
Distribution: Mucujê River, tributary of Paraguaçu River, Brazil (de Pinna & Wosiacki, 2003).
HAEMOMASTER Myers, 1927
Haemomaster Myers, 1927: 131. Type species: Haemomaster venezuelae Myers, 1927. Type by original designation. Gender: Masculine.
Haemomaster venezuelae Myers, 1927
Haemomaster venezuelae Myers, 1927: 131. Type locality: Venezuela, Playa Matepalma, Orinoco. Holotype: CAS
55882.
Distribution: Amazon and Orinoco River basins, Brazil and Venezuela (de Pinna & Wosiacki, 2003).
Remarks: Redescribed in Schmidt (1985).
HATCHERIA Eigenmann, 1909
Hatcheria Eigenmann, 1909b: 250. Type species: Hatcheria patagoniensis Eigenmann, 1909. Type by original designation. Gender: Feminine.
Revision: Arratia & Menu-Marque (1981).
Hatcheria macraei (Girard, 1855)
Thrichomycterus macraei Girard, 1855: 245. Type locality: Near Uspullata, east side of the Cordilleras, Argentina,
elev. 7000 ft. Syntypes: MCZ 8298 (1), USNM 1546 (orig. 3, now 1).
Pygidium Burmeisteri Berg, 1895: 128, pl. 2 (fig. 1). Type locality: Provincia Mendoza (Río Mendoza), Argentina.
Holotype: at MACN.
Hatcheria patagoniensis Eigenmann, 1909b: 250, pl. 33 (fig. 2), pl. 34 (fig. 1). Type locality: Río Blanco at base ofAndes, Patagonia, Argentina, 47º30'S, 72ºW. Holotype: Whereabouts unknown.
Hatcheria titcombi Eigenmann, 1918c: 692. Type locality: Rio Comajo; tributary of Lake Traful, tributary to Rio
Limay, Argentina. Holotype: CAS 28557; holotype illustrated in Eigenmann (1918a: 284, pl. 44, fig. 2).
Hatcheria pique MacDonagh, 1938: 171, figs. 23–24. Type locality: Río Colorado cerca de su desembocadura,
vecino a la estación Pedro Luro del F. C. S., Argentina. Holotype: MLP 15.X.36.
Distribution: Cis-Andean rivers of Argentina, between 29º and 45º30'S (de Pinna & Wosiacki, 2003).
HENONEMUS Eigenmann & Ward, 1907
Henonemus Eigenmann & Ward, in Eigenmann, McAtee & Ward, 1907: 118. Type species: Stegophilus intermedius
Eigenmann & Eigenmann, 1889. Type by original designation. Gender: Masculine.
Cobitiglanis Fowler, 1914: 268. Type species: Ochmacanthus taxistigma Fowler, 1914. Type by original designation. Gender: Masculine. Proposed originally as a subgenus of Ochmacanthus.
Henonemus intermedius (Eigenmann & Eigenmann, 1889)
Stegophilus intermedius Eigenmann & Eigenmann, 1889b: 54. Type locality: Goyaz [Brazil]. Holotype: MCZ 9842.
Distribution: Araguaia River basin, Brazil (de Pinna & Wosiacki, 2003).

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Henonemus macrops (Steindachner, 1882)
Stegophilus macrops Steindachner, 1882b: 178. Type locality: Amazonenstrom. Holotype: at NMW. Species illustrated and described in more detail in Steindachner (1882c: 28, pl. 6, figs. 2, 2a).
Distribution: Amazon River basin, Brazil (de Pinna & Wosiacki, 2003).
Henonemus punctatus (Boulenger, 1887)
Stegophilus punctatus Boulenger, 1887c: 279, pl. 21 (fig. 4). Type locality: Canelos. Holotype: BMNH
1880.12.8.89.
Distribution: Amazon River basin of Brazil, Ecuador and Peru (de Pinna & Wosiacki, 2003).
Henonemus taxistigma (Fowler, 1914)
Ochmacanthus taxistigma Fowler, 1914: 268, fig. 16. Type locality: Rupununi River, British Guiana ... in the highlands of British Guiana, approximately secured in North Latitude 2º to 3º, and West Longitude 50º20'. Holotype:
ANSP 39344.
Distribution: Rupununi River basin, Guyana (de Pinna & Wosiacki, 2003).
HOMODIAETUS Eigenmann & Ward, 1907
Homodiaetus Eigenmann & Ward, in Eigenmann, McAtee & Ward, 1907: 117. Type species: Homodiaetus anisitsi
Eigenmann & Ward, 1907. Type by original designation. Gender: Masculine.
Revision: Koch (2002).
Homodiaetus anisitsi Eigenmann & Ward, 1907
Homodiaetus anisitsi Eigenmann & Ward, in Eigenmann, McAtee & Ward, 1907: 119, pl. 34 (figs. 2–3). Type locality: small creek at Villa Rica, Paraguay. Holotype: CAS 37276; holotype illustrated in Eigenmann (1918a: 351;
pl. 56, fig. 3, 5).
Homodiaetus vazferreirai Devincenzi, in Devincenzi & Vaz-Ferreira, 1939: 168, figured. Type locality: río Uruguay,
en las inmediaciones de la ciudad de Paysandú, Uruguay. Holotype: MHNM CI 345.
Distribution: Paraná-Paraguay River basin (Koch, 2002).
Homodiaetus banguela Koch, 2002
Homodiaetus banguela Koch, 2002: 42, fig. 4. Type locality: Brasil, Rio de Janeiro: Silva Jardim (rio São João, no
cruzamento com a BR-101). Holotype: MCP 19619.
Distribution: São João River, State of Rio de Janeiro, southeastern Brazil (Koch, 2002).
Homodiaetus graciosa Koch, 2002
Homodiaetus graciosa Koch, 2002: 44, fig. 5. Type locality: Brasil, Paraná, Morretes (rio Sagrado, posto florestal).
Holotype: MCP 19618.
Distribution: Coastal basins of southeastern Brazil in states of Paraná and São Paulo (Koch, 2002).
Homodiaetus passarellii (Miranda Ribeiro, 1944)
Stegophilus passarellii Miranda Ribeiro, 1944c: 1, fig. on p. 2. Type locality: Córrego do Barro Branco (conhecido
atualmente como Roncador) com nascentes na Serra do Barro Branco e desaguando no canal Saracuruna— um
dos formadores do rio Estrêla (hoje também conhecido como Canal), e que se lança no fundo de baia de Guanabara no local denominado “Boqueirão” estrada de Magé, Estado do Rio de Janeiro [Brazil]. Holotype: MNRJ
3783.
Distribution: Coastal basins of Rio de Janeiro State, southeastern Brazil (Koch, 2002).
ITUGLANIS Costa & Bockmann, 1993
Ituglanis Costa & Bockmann, 1993: 44. Type species: Pygidium proops parahybae Eigenmann, 1918. Type by original designation. Gender: Masculine.
Ituglanis amazonicus (Steindachner, 1882)
Trichomycterus amazonicus Steindachner, 1882b: 178. Type locality: Cudajas [Brazil]. Holotype: NMW 43306.
Species illustrated and described in more detail in Steindachner (1882c: 29, pl. 6, figs. 4, 4a).
Distribution: Amazon River basin, Brazil, and rivers of French Guiana (de Pinna & Wosiacki, 2003).
Ituglanis bambui Bichuette & Trajano, 2004
Ituglanis bambui Bichuette & Trajano, 2004: 245, fig. 1. Type locality: Brazil: Goiás: São Domingos: Terra Ronca

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State Park: Angélica Cave: upper tributary of main subterranean stream, formed by infiltration of epikarstic
water, 13º31'W, 46º23'W. Holotype: MZUSP 79860.
Distribution: São Domingos karst area, Tocantins River basin, Brazil (Bichuette & Trajano, 2004).
Ituglanis eichorniarum (Miranda Ribeiro, 1912)
Trichomycterus eichorniarum Miranda Ribeiro, 1912: 27. Type locality: Cáceres, nas margens do Paraguay (M.
Grosso) [Brazil]. Lectotype: MNRJ 780A, designated by Miranda Ribeiro (1953: 405).
Distribution: Upper Paraguay River basin, Brazil (de Pinna & Wosiacki, 2003).
Ituglanis epikarsticus Bichuette & Trajano, 2004
Ituglanis epikarsticus Bichuette & Trajano, 2004: 248, fig. 4. Type locality: Brazil: Goiás: São Domingos: Terra
Ronca State Park: São Mateus Cave: rimstone dams (travertine basins) fed by infiltration of epikarstic water(13º40'S, 46º22'W). Holotype: MZUSP 79869.
Distribution: São Domingos karst area, Tocantins River basin, Brazil (Bichuette & Trajano, 2004).
Ituglanis gracilior (Eigenmann, 1912)
Pygidium gracilior Eigenmann, 1912b: 213. Type locality: Erukin, British Guiana. Holotype: FMNH 53264; holotype illustrated in Eigenmann (1918a: 326; pl. 50, fig. 3).
Distribution: Guyana (de Pinna & Wosiacki, 2003).
Ituglanis guayaberensis (Dahl, 1960)
Pygidium metae guayaberensis Dahl, 1960b: 307, fig. 2. Type locality: Small pond in a brook trib. to Río Guayabero, ca. 1 km from mouth of the brook In a brook approx. 1500 m. from Camp 1, same locality as the holotype
of Leyvaichthys castaneus, Orinoco System, Colombia. Holotype: Whereabouts unknown (possibly at ICNMHN).
Distribution: Guayabero River basin, Orinoco drainage, Colombia (de Pinna & Wosiacki, 2003).
Ituglanis herberti (Miranda Ribeiro, 1940)
Trichomycterus Herberti Miranda Ribeiro, 1940: 60, fig. Type locality: Rio Bodoquena, Pantanal, sur de Mato
Grosso [Brazil]. Syntypes (7): MNRJ 1428 (2), MNRJ 1429 (5).
Distribution: Bodoquena River, Paraguay River basin, Brazil (de Pinna & Wosiacki, 2003).
Ituglanis laticeps (Kner, 1863)
Trichomycterus laticeps Kner, 1863: 228, fig. 17. Type locality: Fundort, wie die vorige Art [in reference to Trichomycterus taenia, from “Westabhange der Andes im Staate Ecuador”]. Also described in Kner & Steindachner(1864: 54, pl. 6, fig. 2).
Distribution: Ecuador (de Pinna & Wosiacki, 2003).
Ituglanis macunaima Datovo & Landim, 2005
Ituglanis macunaima Datovo & Landim, 2005: 456, fig. 1. Type locality: Brazil, Mato Grosso, Cocalinho; rio Araguaia basin, Corixo da Saudade (Corixinho); 14º17'20.6"S 51º9'12.1"W. Holotype: MZUSP 88452.
Distribution: Araguaia River basin, Brazil (Datovo & Landim, 2005).
Ituglanis metae (Eigenmann, 1918)
Pygidium metae Eigenmann, 1918c: 694. Type locality: Barrigona, Colombia. Holotype: CAS 58138; holotype illustrated in Eigenmann (1918a: 312, pl. 47, fig. 5).
Distribution: Colombia (de Pinna & Wosiacki, 2003).
Ituglanis nebulosus de Pinna & Keith, 2003
Ituglanis nebulosus de Pinna & Keith, 2003: 874, figs. 1, 2. Type locality: French Guiana, Arataye river (Tributaryof Approuague river), near the natural preserve “Reserve naturelle des Nouragues” (04º00'22"N, 52º36'34"W).
Holotype: MNHN 2001-1128.
Distribution: Approuague River basin, French Guiana (de Pinna & Keith, 2003).
Ituglanis parahybae (Eigenmann, 1918)
Pygidium proops parahybae Eigenmann, 1918a: 332. Type locality: Rio Parahyba, Brazil. Holotype: FMNH 58576.
Distribution: Paraíba do Sul and São João River basins, Brazil (de Pinna & Wosiacki, 2003).
Ituglanis parkoi (Miranda Ribeiro, 1944)
Pygidium parkoi Miranda Ribeiro, 1944b: 1, fig. Type locality: rio Iticoaí (entre os rios das Pedras e Ituí) que
deságua no Javari, afluente do Amazonas, Benjamin Constant, Estados do Amazonas [Brazil]. Holotype: MNRJCHECKLIST OF CATFISHES, RECENT AND FOSSIL

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3849.
Distribution: Amazon River basin, Brazil (de Pinna & Wosiacki, 2003).
Ituglanis passensis Fernández & Bichuette, 2002
Ituglanis passensis Fernández & Bichuette, 2002: 274, fig. 1. Type locality: Brazil: Goiás: São Domingos: Passa
Três cave: subterranean stream in the Tocantins drainage, 13º36'S 46º23'W. Holotype: MCP 27382.
Distribution: Passa Três cave system in São Domingos, Goiás, Brazil (de Pinna & Wosiacki, 2003).
Ituglanis proops (Miranda Ribeiro, 1908)
Trichomycterus proöps Miranda Ribeiro, 1908b: [3], fig. 4. Type locality: Rio Ribeira, Iguape, [Brazil]. Lectotype:
MNRJ 781A, designated by Miranda Ribeiro (1953: 405).
Distribution: Ribeira de Iguape River basin, Brazil (de Pinna & Wosiacki, 2003).
Ituglanis ramiroi Bichuette & Trajano, 2004
Ituglanis ramiroi Bichuette & Trajano, 2004: 250, fig. 7. Type locality: Brazil: Goiás: São Domingos: Terra Ronca
State Park: São Bernardo Cave: side pool fed by small water inlet (13º49'S, 46º21'W), for about 700 m fromcave entrance. Holotype: MZUSP 79865.
Distribution: São Domingos karst area, Tocantins River basin, Brazil (Bichuette & Trajano, 2004).
LISTRURA de Pinna, 1988
Listrura de Pinna, 1988: 114. Type species: Listrura nematopteryx de Pinna, 1988. Type by original designation.
Gender: Feminine.
Listrura boticario de Pinna & Wosiacki, 2002
Listrura boticario de Pinna & Wosiacki, 2002: 721, fig. 1. Type locality: Brazil, State of Paraná, Município de
Guaraqueçaba, pool adjacent to Rio da Figueira (tributary of Rio Morato, itself tributary to Rio Guaraqueçaba,an isolated coastal basin that drains directly into the Baía das Laranjeiras), inside the nature preserve “Reserva
Particular do Patrimônio Natural Salto Morato” (25º16'S, 48º12'W, UTM: 7.212.500–7.215.400). Holotype:
MZUSP 69573.
Distribution: Da Figueira and Guaraqueçaba River basins, Paraná State, Brazil (de Pinna & Wosiacki, 2003).
Listrura camposi (Miranda Ribeiro, 1957)
Eremophilus camposi Miranda Ribeiro, 1957: 72, fig. Type locality: Rio Poço Grande, [trib. of the Rio Juquiá,
Fazenda Poço Grande, Juquiá, ca. 24º15'S, 47º37'W, Sao Paulo State, Brazil]. Holotype: MZUSP 3426.
Distribution: Poço Grande River, tributary of the Juquiá River, São Paulo State, and Ribeirão da Ilha, Florianópolis,
Santa Catarina State, Brazil (de Pinna & Wosiacki, 2003).
Listrura nematopteryx de Pinna, 1988
Listrura nematopteryx de Pinna, 1988: 115, figs. 1–2. Type locality: Small marsh which is source of creek later joining Ribeirão Imbariê, trib. of Rio Estrela, near 58 km mark of old road leading to Petrópolis (Antiga Rio-Petrópolis, also called “Estrada Automóvel Club”), Município de Magé, Localidade de Piabetá, Rio de Janeiro, Brazil(22º36'36"S 43º11'26"W). Holotype: MZUSP 36974.
Distribution: Imbariê Creek, basin, Estrela River system, Piabetá, Rio de Janeiro State, and Picinguaba, São Paulo
State, Brazil (de Pinna & Wosiacki, 2003).
Listrura tetraradiata Landim & Costa, 2002
Listrura tetraradiata Landim & Costa, 2002: 152, fig. 1. Type locality: Brazil: Estado do Rio de Janeiro: Municipio
de Saquarema, rio da Represa, a tributary of rio Bom Sucesso, rio Ibicuíba basin of the Lagoa de Araruama system, Palmital, approximately 5 km north from Bacaxá, about 22º50'S, 42º28'W. Holotype: MZUSP 52572.
Distribution: Ibicuíba River, Araruama Lagoon system, Brazil (de Pinna & Wosiacki, 2003).
MALACOGLANIS Myers & Weitzman, 1966
Malacoglanis Myers & Weitzman, 1966: 281. Type species: Malacoglanis gelatinosus Myers & Weitzman, 1966.
Type by original designation. Gender: Masculine.
Malacoglanis gelatinosus Myers & Weitzman, 1966
Malacoglanis gelatinosus Myers & Weitzman, 1966: 282, figs. 3–4. Type locality: Small caño of the Río Orteguaza,a short distance above its junction with the Rio Caquetá, Caquetá Province, Colombia. Holotype: SU 50754.

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FERRARISDistribution: Caquetá River basin, Colombia (de Pinna & Wosiacki, 2003).
MEGALOCENTOR de Pinna & Britski, 1991
Megalocentor de Pinna & Britski, 1991: 115. Type species: Megalocentor echthrus de Pinna & Britski, 1991. Typeby original designation. Gender: Masculine.
Megalocentor echthrus de Pinna & Britski, 1991
Megalocentor echthrus de Pinna & Britski, 1991: 116, figs. 3, 4 (a–b). Type locality: Rio Madeira, Calama, Praia do
Caraparu, Estado do Amazonas, Brazil. Holotype: MZUSP 41879.
Distribution: Amazon and Orinoco River basins (de Pinna & Wosiacki, 2003).
MICROCAMBEVA Costa & Bockmann, 1994
Microcambeva Costa & Bockmann, 1994a: 718. Type species: Microcambeva barbata Costa & Bockmann, 1994.
Type by original designation. Gender: Feminine.
Microcambeva barbata Costa & Bockman, 1994
Microcambeva barbata Costa & Bockmann, 1994a: 718, figs. 2–3. Type locality: Brazil: Estado do Rio de Janeiro:rio São João, near Gaviões, Município de Silva Jardim. Holotype: MZUSP 43678.
Distribution: Atlantic coastal drainages of Rio de Janeiro and Espírito Santo states, Brazil (de Pinna & Wosiacki,2003).
Microcambeva ribeirae Costa, Lima & Bizerril, 2004
Microcambeva ribeirae Costa, Lima & Bizerril, 2004: 3, figs. 1, 2. Type locality: Brazil: Estado de São Paulo:
Município de Pedro de Toledo, Rio São Lourencinho, no Bairro São Lourenço. Holotype: MZUSP 84301.
Distribution: Ribeira do Iguape River basin, SE Brazil (Costa et al., 2004).
MIUROGLANIS Eigenmann & Eigenmann, 1889
Miuroglanis Eigenmann & Eigenmann, 1889b: 55. Type species: Miuroglanis platycephalus Eigenmann & Eigenmann, 1889. Type by original designation. Gender: Masculine.
Miuroglanis platycephalus Eigenmann & Eigenmann, 1889
Miuroglanis platycephalus Eigenmann & Eigenmann, 1889b: 56. Type locality: Jutahy [Brazil]. Holotype: MCZ
8172.
Distribution: upper Amazon River basin, Brazil (de Pinna & Wosiacki, 2003).
OCHMACANTHUS Eigenmann, 1912
Ochmacanthus Eigenmann, 1912: 213. Type species: Ochmacanthus flabelliferus Eigenmann, 1912. Type by original designation. Gender: Masculine.
Gyrinurus Miranda Ribeiro, 1912: 27. Type species: Gyrinurus batrachostoma Miranda Ribeiro, 1912. Type bymonotypy. Gender: Masculine.
Ochmacanthus alternus Myers, 1927
Ochmacanthus alternus Myers, 1927: 129. Type locality: Venezuela: Caño de Quiribana, near Caicara. Lectotype:
CAS 13522.
Distribution: Negro and Orinoco River basins, Brazil and Venezuela (de Pinna & Wosiacki, 2003).
Ochmacanthus batrachostoma (Miranda Ribeiro, 1912)
Gyrinurus batrachostoma Miranda Ribeiro, 1912: 28, pl. Type locality: rio Paraguay em S. Luiz de Cáceres [ Mato
Grosso, Brazil]. Holotype: MNRJ 786.
Distribution: Paraguay River basin, Brazil (de Pinna & Wosiacki, 2003); la Plata basin, Argentina (Casciotta & Almirón, 1996).
Ochmacanthus flabelliferus Eigenmann, 1912
Ochmacanthus flabelliferus Eigenmann, 1912b: 213. Type locality: Konawaruk, British Guiana. Holotype: FMNH
53263; holotype illustrated in Eigenmann (1918a: 357; pl. 55, fig. 5).
Distribution: River drainages in Guyana and Venezuela (de Pinna & Wosiacki, 2003).
Ochmacanthus orinoco Myers, 1927

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Ochmacanthus orinoco Myers, 1927: 130. Type locality: Venezuela: Playa Matepalma, Orinoco. Holotype: CAS
76355.
Distribution: Negro and Orinoco River basins, Brazil and Venezuela (de Pinna & Wosiacki, 2003).
Ochmacanthus reinhardtii (Steindachner, 1882)
Stegophilus Reinhardtii Steindachner, 1882b: 178. Type locality: Amazonenstrom und Rio Iça. Syntypes: NMW
44735 (2), NMW 50603 (3). Species illustrated and described in more detail in Steindachner (1882c: 28, pl. 6,
fig. 1).
Distribution: Amazon River basin, Brazil, and drainages in French Guyana (de Pinna & Wosiacki, 2003).
PARACANTHOPOMA Giltay, 1935
Paracanthopoma Giltay, 1935: 1. Type species: Paracanthopoma parva Giltay, 1935. Type by original designation.
Gender: Neuter.
Phylogeny: Schmidt (1993).
Paracanthopoma parva Giltay, 1935
Paracanthopoma parva Giltay, 1935: 1, figs. 1–3. Type locality: Rio Catrymany supérieur [Brazil]. Holotype:IRSNB 43.
Distribution: Amazon basin, Brazil, and Essequibo River basin, Guyana (de Pinna & Wosiacki, 2003).
Remarks: Redescribed, with notes on biology, in Schmidt (1993).
PARASTEGOPHILUS Miranda Ribeiro, 1946
Parastegophilus Miranda Ribeiro, 1946: 12. Type species: Stegophilus maculatus Steindachner, 1879. Type by original designation. Gender: Masculine.
Parastegophilus maculatus (Steindachner, 1879)
Stegophilus maculatus Steindachner, 1879a: 32. Type locality: La Plata, Prov. of Buenos Aires, Argentina. Holotype:
NMW 57222. Species illustrated and described in more detail in Steindachner (1879e: 25, pl. 4, fig. 2).
Distribution: Lower Paraná and Uruguay River basins, Argentina (de Pinna & Wosiacki, 2003).
Parastegophilus paulensis (Miranda Ribeiro, 1918)
Pseudostegophilus paulensis Miranda Ribeiro, 1918e: 727. Type locality: Avanhandava, Rio Tieté, Estado de São
Paulo [Brazil]. Holotype: MZUSP 2272.
Pseudostegophilus scarificator Ihering, 1930: 100, pl. 13 (fig. 2). Type locality: São Paulo [Brazil]. Syntypes (2):Whereabouts unknown.
Distribution: Upper Paraná River basin, Brazil (de Pinna & Wosiacki, 2003).
PARAVANDELLIA Miranda Ribeiro, 1912
Paravandellia Miranda Ribeiro, 1912: 28. Type species: Paravandellia oxyptera Miranda Ribeiro, 1912. Type bymonotypy. Gender: Feminine.
Branchioica Eigenmann, 1918c: 702. Type species: Branchioica bertoni Eigenmann, 1918. Type by original designation. Gender: Feminine.
Pleurophysus Miranda Ribeiro, 1918d: 636. Type species: Pleurophysus hydrostaticus Miranda Ribeiro, 1918. Typeby monotypy. Gender: Masculine.
Parabranchioica Devincenzi & Vaz-Ferreira, 1939: 5. Type species: Parabranchioica teaguei Devincenzi & VazFerreira, 1939. Type by monotypy. Gender: Feminine.
Paravandellia oxyptera Miranda Ribeiro, 1912
Paravandellia oxyptera Miranda Ribeiro, 1912: 29. Type locality: Caceres, margens do Rio Paraguay [Brazil].
Holotype: MNRJ 790.
Branchioica bertoni Eigenmann, 1918c: 703. Type locality: Asuncion, Paraguay. Holotype: CAS 63840; holotype
illustrated in Eigenmann (1918a: 368, pl. 53, figs. 3–5) as Branchioica bertonii.
Pleurophysus hydrostaticus Miranda Ribeiro, 1918d: 636. Type locality: Rio Claro [Brazil]. Lectotype: MZUSP
2214, designated by Britski (1969: 206).
Vandellia hematophaga Guimarães, 1935: 300, figs. 1–4. Type locality: Río Tietê, Salto, São Paulo [Brazil]. Lecto-

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type: SU 16766, designated by Böhlke (1953: 45).
Parabranchioica teaguei Devincenzi & Vaz-Ferreira, 1939: 5, unnumbered figure. Type locality: río Uruguay, en las
inmediaciones de la ciudad de Paysandú [Uruguay]. Type(s): at MHNM.
Distribution: Paraná, Paraguay and Uruguay River basins, Brazil, Paraguay, and Uruguay (de Pinna & Wosiacki,2003).
Paravandellia phaneronema (Miles, 1943)
Branchioica phaneronema Miles, 1943a: 32, fig. 8. Type locality: various points in the Upper Cauca valley. Holotype: lost (Román-Valencia, 1998). Described again in Miles (1943b: 367, figs. 1–3).
Branchioica magdalenae Miles, 1943a: 33. Type locality: Río Magdalena, Colombia. Holotype: lost (Román-Valencia, 1998). Described again in Miles (1943b: 368).
Distribution: Magdalena and Cauca River basins, Colombia (de Pinna & Wosiacki, 2003).
Remarks: Although both of these nominal species were described twice in 1943, the text of Miles (1943b) indicatesthat the names were first made available in Miles (1943a). Román-Valencia (1998) treated Branchioica
magdalenae as a junior synonym of Paravandellia phaneronema, thereby acting as first reviser.
PAREIODON Kner, 1855
Pareiodon Kner, 1855a: 160. Type species: Pareiodon microps Kner, 1855. Type by monotypy. Gender: Masculine.
Centrophorus Kner, 1859: 167 (footnote). Type species: Pareiodon microps Kner, 1855. Type by being a replacement name. Gender: Masculine. Unneeded replacement for Pareiodon Kner, 1855, then considered preoccupied
by Parodon. Preoccupied by Centrophorus Müller & Henle, 1837, in Elasmobranchs.
Astemomycterus Guichenot, 1860: 525. Type species: Trichomycterus pusillus Castelnau, 1855. Type by original
designation. Gender: Masculine.
Pareiodon microps Kner, 1855
Pareiodon microps Kner, 1855a: 160, fig. 2. Type locality: Borba (?) [Río Madeira]. Syntypes: NMW 45486 (2).
Trichomycterus pusillus Castelnau, 1855: 50, pl. 24 (fig. 4). Type locality: l’Araguay et [...] l’Amazone. Syntypes:
MNHN 0000-1210 (4).
Distribution: Amazon River basin, Brazil (de Pinna & Wosiacki, 2003).
Remarks: Priority of publication of the two included synonyms may not have yet been established, inasmuch as theprecise dates of publication of the two works were not listed in Eschmeyer et al. (1998) or other relevant publications. If priority cannot be used to resolve this issue, precedence must be given to Pareiodon microps by firstreviser action of Eigenmann & Eigenmann (1889: 346).
PLECTROCHILUS Miranda Ribeiro, 1917
Plectrochilus Miranda Ribeiro, 1917: 50. Type species: Plectrochilus machadoi Miranda Ribeiro, 1917. Type bymonotypy. Gender: Masculine.
Plectrochilus diabolicus (Myers, 1927)
Urinophilus diabolicus Myers, 1927: 132. Type locality: Iquitos, Peru. Holotype: CAS 59940.
Distribution: Amazon River basin, Brazil and Peru (de Pinna & Wosiacki, 2003).
Plectrochilus machadoi Miranda Ribeiro, 1917
Plectrochilus machadoi Miranda Ribeiro, 1917: 50, three unnumbered figures. Type locality: Fluvio Solimões, [Brazil]. Holotype: MNRJ 978.
Vandellia hasemani Eigenmann, 1918a: 363, figs. 33–35, pl. 53 (fig. 3). Type locality: Río Mamoré, Alto Amazonas,
Bolivia. Holotype: FMNH 58523.
Distribution: Amazon River basin, Brazil and Peru (de Pinna & Wosiacki, 2003).
Plectrochilus wieneri (Pellegrin, 1909)
Vandellia wieneri Pellegrin, 1909e: 199, unnumbered figure. Type locality: [Río Napo, near mouth of Río Misahually, Ecuador]. Holotype: MNHN a-9934.
Distribution: Napo River basin, Ecuador (de Pinna & Wosiacki, 2003).
PSEUDOSTEGOPHILUS Eigenmann & Eigenmann, 1889

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Pseudostegophilus Eigenmann & Eigenmann, 1889b: 54. Type species: Stegophilus nemurus Günther, 1869. Typeby original designation. Gender: Masculine.
Pseudostegophilus haemomyzon (Myers, 1942)
Homodiaetus haemomyzon Myers, 1942: 98, fig. 4. Type locality: Río Guarico at Calabozo, Venezuela. Holotype:SU 36500.
Distribution: Orinoco River basin, Venezuela (de Pinna & Wosiacki, 2003).
Pseudostegophilus nemurus (Günther, 1869)
Stegophilus nemurus Günther, 1869: 429. Type locality: Upper Amazon River, Peru. Holotype: BMNH 1869.5.21.9.
Distribution: Amazon River basin, Brazil and Peru (de Pinna & Wosiacki, 2003).
PYGIDIANOPS Myers, 1944
Pygidianops Myers, 1944: 592. Type species: Pygidianops eigenmanni Myers, 1944. Type by original designation.
Gender: Masculine.
Key: Schaefer et al. (2005: 5).
Pygidianops cuao Schaefer, Provenzano, de Pinna & Baskin, 2005
Pygidianops cuao Schaefer, Provenzano, de Pinna & Baskin, 2005: 5, figs. 1, 2. Type locality: Venezuela, Estado
Amazonas, Río Cuao at Raudal Guacamaya, 8.1 miles upstream from Raudal El Danto, 05º07.71'N,
67º31.53'W. Holotype: MBUCV V-30917.
Distribution: Cuao River, Orinoco basin, Venezuela (Schaefer et al., 2005).
Pygidianops eigenmanni Myers, 1944
Pygidianops eigenmanni Myers, 1944: 592, pl. 52 (fig. 1), pl. 53 (figs. 3–5). Type locality: Rock pools below São
Gabriel Rapids, Rio Negro, Brazil. Holotype: CAS 11120.
Distribution: Negro River basin, Brazil (de Pinna & Wosiacki, 2003).
Pygidianops magoi Schaefer, Provenzano, de Pinna & Baskin, 2005
Pygidianops magoi Schaefer, Provenzano, de Pinna & Baskin, 2005: 10, figs. 5, 6. Type locality: Venezuela, Estado
Delta Amacuro, Río Orinoco at Puerto Cabrian, 8º34.8'N, 62º15.9'W. Holotype: MBUCV V-31035.
Distribution: Lower Orinoco River, between Ciudad Bolivar and Barrancas, Venezuela (Schaefer et al., 2005).
RHIZOSOMICHTHYS Miles, 1943
Bathophilus Miles, 1942: 57. Type species: Pygidium totae Miles, 1942. Type by monotypy. Gender: Masculine.
Preoccupied by Bathophilus Giglioli, 1882, in fishes; replaced by Rhizosomichthys Miles, 1943, and Bathypygidium Whitley, 1947.
Rhizosomichthys Miles, 1943b: 369. Type species: Pygidium totae Miles, 1942. Type by being a replacement name.
Gender: Masculine. Replacement for Bathophilus Miles, 1942; preoccupied by Bathophilus Giglioli, 1882.
Bathypygidium Whitley, 1947: 150. Type species: Pygidium totae Miles, 1942. Type by being a replacement name.
Gender: Neuter. Replacement for Bathophilus Miles, 1942; preoccupied by Bathophilus Giglioli, 1882.
Rhizosomichthys totae (Miles, 1942)
Pygidium totae Miles, 1942: 55, unnumbered figure. Type locality: Lago de Tota, Boyacá, Cordillera Oriental,
Colombia, 3060 metros. Holotype: ICNMNH 20 [? now ICNMHN 353].
Distribution: Lake Tota basin, Colombia (de Pinna & Wosiacki, 2003).
SARCOGLANIS Myers & Weitzman, 1966
Sarcoglanis Myers & Weitzman, 1966: 279. Type species: Sarcoglanis simplex Myers & Weitzman, 1966. Type byoriginal designation. Gender: Masculine.
Sarcoglanis simplex Myers & Weitzman, 1966
Sarcoglanis simplex Myers & Weitzman, 1966: 279, figs. 1–2. Type locality: Rock pools below São Gabriel Rapids,of the Rio Negro (below the town of Uaupés, formerly São Gabriel), Estado de Amazonas, Brazil. Holotype: SU
50189.
Distribution: Upper Negro River basin, Brazil (de Pinna & Wosiacki, 2003).

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Schultzichthys Dahl, 1960b: 312. Type species: Schultzichthys gracilis Dahl, 1960. Type by monotypy. Gender:
Masculine.
Schultzichthys bondi (Myers, 1942)
Acanthopoma bondi Myers, 1942: 97, fig. 5. Type locality: Río Apuré at San Fernando de Apuré, Venezuela. Holotype: SU 36498.
Distribution: Amazon and Orinoco River basins (de Pinna & Wosiacki, 2003).
Schultzichthys gracilis Dahl, 1960
Schultzichthys gracilis Dahl, 1960b: 312, figs. 4–5. Type locality: foot of a rapid in Caño Lozada, about 15 km
above its junction with the Guayabero River [Río Orinoco system, Colombia]. Holotype: Whereabouts
unknown (possibly at ICNMHN).
Distribution: Guayabero River, Orinoco River basin, Colombia (de Pinna & Wosiacki, 2003).
SCLERONEMA Eigenmann, 1918
Scleronema Eigenmann, 1918c: 691. Type species: Scleronema operculatum Eigenmann, 1918. Type by original
designation. Gender: Neuter.
Scleronema angustirostre (Devincenzi, 1942)
Pygidium angustirostris Devincenzi, in Devincenzi & Teague, 1942: 30, pl. 4 (fig. 3). Type locality: La Cañeda delas Piedras, Uruguay. Holotype: at MHNM (not found).
Distribution: Uruguay (de Pinna & Wosiacki, 2003).
Scleronema minutum (Boulenger, 1891)
Trichomycterus minutus Boulenger, 1891: 235, pl. 26 (fig. 3). Type locality: San Lorenzo district [= São Lourenço
das Missões], Brazil. Syntypes: BMNH 1891.3.16.84–86 (3).
Distribution: Rio Grande do Sul State, Brazil (de Pinna & Wosiacki, 2003); la Plata basin, Argentina (Casciotta &Almirón, 1996).
Scleronema operculatum Eigenmann, 1918
Scleronema operculatum Eigenmann, 1918c: 691. Type locality: Cacequy, Brazil. Holotype: FMNH 58080; holotype illustrated in Eigenmann (1918a: 281, pl. 44, fig. 1).
Distribution: Rio Grande do Sul State, Brazil (de Pinna & Wosiacki, 2003).
SILVINICHTHYS Arratia, 1998
Silvinichthys Arratia, 1998: 356. Type species: Trichomycterus mendozensis Arratia, Chang, Menu-Marque & Rojas,
1978. Type by monotypy. Gender: Masculine.
Silvinichthys bortayro Fernández & de Pinna, 2005
Silvinichthys bortayro Fernández & de Pinna, 2005: 101, figs. 1–2. Type locality: Argentina, Provincia de Salta,
Departamento Capital, artificial well near Río Arenales at San Luis, 1273 m elevation, 12 m depth, 24º50'S,
65º30'W. Holotype: FML 2590.
Distribution: Artificial wells in Salta Province, Argentina (Fernández & de Pinna, 2005).
Silvinichthys mendozensis (Arratia, Chang, Menu-Marque & Rojas, 1978)
Trichomycterus mendozensis Arratia, Chang, Menu-Marque & Rojas, 1978: 170, figs. 8–10. Type locality: freshwater of the preandean range of Mendoza, Argentina ..., where it occupies a restricted environment between 1,500
to 1,700 m above sea level. Holotype: FFSUC IC 118-27.
Distribution: Mendoza River basin, Argentina (de Pinna & Wosiacki, 2003).
STAUROGLANIS de Pinna, 1989
Stauroglanis de Pinna, 1989a: 6. Type species: Stauroglanis gouldingi de Pinna, 1989. Type by original designation.
Gender: Masculine.
Stauroglanis gouldingi de Pinna, 1989
Stauroglanis gouldingi de Pinna, 1989a: 7, figs. 1–3. Type locality: Cachoeira do Aracu, R. Daraá (R. Negro drainage system), Estado do Amazonas, Brazil. Holotype: MZUSP 31088.

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Distribution: Daraá River, Negro River basin, Brazil (de Pinna & Wosiacki, 2003).
STEGOPHILUS Reinhardt, 1859
Stegophilus Reinhardt, 1859: 5 Type species: Stegophilus insidiosus Reinhardt, 1859. Type by monotypy. Gender:
Masculine.
Stegophilus insidiosus Reinhardt, 1859
Stegophilus insidiosus Reinhardt, 1859: 5, pl. 2. Type locality: Rio das Velhas, Rio São Francisco system, Brazil.
Syntypes: BMNH 1875.5.22.1 (1), ZMUC P 30152 (1), ZMUC P 30153 (1).
Distribution: São Francisco River basin, Brazil (de Pinna & Wosiacki, 2003).
Stegophilus panzeri (Ahl, 1931)
Henonemus panzeri Ahl, 1931: 210, fig. 3. Type locality: Rio Capim [Brazil]. Holotype: at ZMB (orig. no. 656).
Distribution: Lower Amazon River basin, Brazil (de Pinna & Wosiacki, 2003).
Stegophilus septentrionalis Myers, 1927
Stegophilus septentrionalis Myers, 1927: 130. Type locality: Venezuela: Santa Barbara, Orinoco. Holotype: CAS
64597.
Distribution: Orinoco River basin, Venezuela (de Pinna & Wosiacki, 2003).
STENOLICMUS de Pinna & Starnes, 1990
Stenolicmus de Pinna & Starnes, 1990: 77. Type species: Stenolicmus sarmientoi de Pinna & Starnes, 1990. Type byoriginal designation. Gender: Masculine.
Stenolicmus sarmientoi de Pinna & Starnes, 1990
Stenolicmus sarmientoi de Pinna & Starnes, 1990: 77, fig. 1–2. Type locality: Río Matos, trib. of Río Apere, 48 km
east of San Borja, Ballivia Prov., Depto. Beni, Bolivia, 14º55'S, 66º17'W. Holotype: USNM 301664.
Distribution: Upper Apere River basin, Bolivia (de Pinna & Wosiacki, 2003).
TRICHOGENES Britski & Ortega, 1983
Trichogenes Britski & Ortega, 1983: 211. Type species: Trichogenes longipinnis Britski & Ortega, 1983. Type byoriginal designation. Gender: Masculine.
Trichogenes longipinnis Britski & Ortega, 1983
Trichogenes longipinnis Britski & Ortega, 1983: 212, figs. 1–2. Type locality: Cachoeira do Amor, km 3 da estrada
Parati-Ubatuba, SP [Brazil] Holotype: MZUSP 16099.
Distribution: Atlantic coastal drainages, northern São Paulo State, Brazil (de Pinna & Wosiacki, 2003).
TRICHOMYCTERUS Valenciennes, 1832
Trichomycterus Valenciennes, 1832a: 348. Type species: Trichomycterus nigricans Valenciennes, 1832. Type bymonotypy. Gender: Masculine.
Review: Fernández (1999, Argentina).
Trichomycterus albinotatus Costa, 1992
Trichomycterus albinotatus Costa, 1992: 102, figs. 1–2. Type locality: Estado do Rio de Janeiro: Visconde de Maua,
1 km O de la Ville de Maromba, rio Preto, bassin du rio Paraíba [22º30'S 44º15'], Brazil. Holotype: MZUSP
42312.
Distribution: Preto River, Paraíba do Sul River basin, Brazil (de Pinna & Wosiacki, 2003).
Trichomycterus alternatus (Eigenmann, 1918)
Pygidium alternatum Eigenmann, 1918c: 700. Type locality: Rio Doce, Brazil. Holotype: FMNH 58082; holotype
illustrated in Eigenmann (1918a: 336, pl. 52, fig. 3).
Pygidium florense Miranda Ribeiro, 1943: 1, fig. Type locality: rio das Flores, próximo a Ipiabas (estação de Pandiá
Calógeras), Estado do Rio de Janeiro [Brazil]. Holotype: MNRJ 3751.
Pygidium travassosi Miranda Ribeiro, 1949a: 145, fig. 2. Type locality: Rio das Pedras, Fazenda Penedo, Agulhas
Negras, Estado do Rio, Rio de Janeiro, Brazil. Holotype: MNRJ 5424.
Distribution: Atlantic coastal river basins, Rio de Janeiro and Espírito Santo States, Brazil (de Pinna & Wosiacki,

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2003), Jequitinhonha River basin, Minas Gerais (Triques & Vono, 2004).
Trichomycterus alterus (Marini, Nichols & La Monte, 1933)
Pygidium alterum Marini, Nichols & La Monte, 1933: 2, fig. 2. Type locality: Rio de los Sauces, La Rioja, Argentina. Holotype: AMNH 12241.
Distribution: Humahuaca, Los Sauces River and Valle Guanchin (La Rioja), Argentina (de Pinna & Wosiacki, 2003).
Trichomycterus areolatus Valenciennes, 1846
Trichomycterus areolatus Valenciennes, in Cuvier & Valenciennes, 1846: 492. Type locality: la rivière de San Jago,
Santiago. Syntypes: MNHN 0000-3167 (3), MNHN a-6310 (2).
Trichomycterus maculatus Valenciennes, in Cuvier & Valenciennes, 1846: 493. Type locality: Santiago du Chili.
Syntypes: MNHN 0000-3166 (1), MNHN 0000-4236 (10).
Trichomycterus marmoratus Philippi, 1866: 714. Type locality: Chile. No types known.
Trichomycterus palleus Philippi, 1866: 715. Type locality: Chile. No types known.
Trichomycterus tigrinus Philippi, 1866: 714. Type locality: Chile. No types known.
Distribution: Pacific draining rivers of Central Chile (de Pinna & Wosiacki, 2003).
Trichomycterus arleoi (Fernández-Yépez, 1972)
Pygidium arleoi Fernández-Yépez, 1972a: 21, pl. 34. Type locality: Estación -140 — de la Cuenca del Rio Yaracuy,
Venezuela. Holotype: collection of author, whereabouts unknown.
Distribution: Yaracuy River basin, Venezuela (de Pinna & Wosiacki, 2003).
Trichomycterus auroguttatus Costa, 1992
Trichomycterus auroguttatus Costa, 1992: 105, fig. 6. Type locality: Rio de Janeiro Estado do Rio de Janeiro: 1 km à
l’ouest de la ville de Visconde de Mauá, rio Marimbondo, affluent du rio Preto, bassin du rio Paraíba, Brazil.
Holotype: MZUSP 43341.
Distribution: Marimbondo River, Preto River system, Paraíba do Sul River basin, Brazil (de Pinna & Wosiacki,2003).
Trichomycterus bahianus Costa, 1992
Trichomycterus bahianus Costa, 1992: 105, fig. 5. Type locality: Estado da Bahia: Una, 6,5 km au sud-est de la villede Sao José, dans um ruisseau affluent du ribeirão Caveira, bassin du rio Una, Bahia, Brazil. Holotype: MZUSP
43340.
Distribution: Ribeirão Caveira tributary, Una River basin, State of Bahia, Brazil (de Pinna & Wosiacki, 2003).
Trichomycterus banneaui (Eigenmann, 1912)
Pygidium banneaui Eigenmann, 1912a: 19. Type locality: Bernal Creek near Honda, Colombia. Holotype: FMNH
56025; holotype illustrated in Eigenmann (1918a: 318, pl. 48, fig. 1).
Distribution: River drainages of Tolima and Cundinamarca, Colombia (Maldonado-Ocampo et al., 2005).
Remarks: Treated as a junior synonym of Trichomycterus bogotensis (Eigenmann, 1912), in de Pinna & Wosiacki(2003), who also apparently acted as first reviser, but treated as a valid species in Maldonado-Ocampo et al.
(2005).
Trichomycterus barbouri (Eigenmann, 1911)
Pygidium barbouri Eigenmann, 1911: 214, pl. 32. Type locality: Río Beni, e. Bolivia. Holotype: MCZ 29313.
Distribution: Amazon River basin, Bolivia, and la Plata River basin, Argentina (Fernández, 2000b); Beni Riverbasin, Bolivia (de Pinna & Wosiacki, 2003).
Remarks: Redescribed in Fernández (2000).
Trichomycterus belensis Fernández & Vari, 2002
Trichomycterus belensis Fernández & Vari, 2002: 739, fig. 1. Type locality: Argentina, Provincia de Catamarca,
Departamento Belén, stream tributary to Laguna Blanca, 11 km northeast from Belén on provincial road 43
along route from Belén to Antogagasta de la Sierra, near Los Nacimientos de San Antonio (approximately
26º30'S, 67º03'W). Holotype: FML 2530.
Distribution: Laguna Blanca basin, Argentina (de Pinna & Wosiacki, 2003).
Trichomycterus bogotensis (Eigenmann, 1912)
Pygidium bogotense Eigenmann, 1912a: 18. Type locality: On the plains of Bogota, at an elevation of nearly nine
thousand feet, ..., Chapinero, Colombia. Holotype: FMNH 56030; holotype illustrated in Eigenmann (1918a:

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315, pl. 49, fig. 3).
Distribution: Upper Cauca, Calima and San Juan rivers, Colombia (Maldonado-Ocampo et al., 2005).
Trichomycterus bomboizanus (Tortonese, 1942)
Pygidium bomboizanum Tortonese, 1942: 113, pl. 1. Type locality: Rio Bomboiza (Ecuador merid.) Ecuador. Holotype: MZUT 3553.
Distribution: Bomboiza River basin, Ecuador (de Pinna & Wosiacki, 2003).
Trichomycterus borellii Boulenger, 1897
Trichomycterus borellii Boulenger, 1897c: 3. Type locality: Mission d’Aguairenda, Tala, and Lesser, Argentina.
Syntypes: BMNH 1897.1.27.26 (1), ZMUT 1396 (6), ZMUT 1397 (2), ZMUT 13998 (1).
Pygidium Schmidti Berg, 1897: 266. Type locality: Río de Belén (Provincia de Catamarca), Argentina. Syntypes:
BMNH 1898.9.23.3 (1), MACN 2361 (1), MACN 4595 (1), MACN 5164 (2), MACN 5176 (1).
Distribution: Salta, Catamarca and Mendoza, Argentina; Aguairenda, Bolivia (de Pinna & Wosiacki, 2003).
Trichomycterus boylei Nichols, 1956
Pygidium boylei Nichols, 1956: 1, fig. 1. Type locality: dry country at Tilcara, Argentina, elevation 8000 feet,
Argentina. Holotype: AMNH 20299.
Distribution: Grande River basin, Argentina (de Pinna & Wosiacki, 2003).
Trichomycterus brasiliensis Reinhardt, 1874
Trichomycterus brasiliensis Reinhardt, in Lütken, 1874c: 29. Type locality: in Rio das Velhas. Syntypes: NMW
85270 (3), SMNS 2021 (1), ZMB 9171 (2), ZMUC P 30154–ZMUC P 30166 (1 each).
Trichomycterus brasiliensis tristis Lütken, 1875: 137. Type locality: Rio das Velhas, Minas Gerais, Brazil. Holotype:
ZMUC P 30167.
Distribution: Upper São Francisco River, State of Minas Gerais and in smaller adjoining basins in Southeastern Brazil (de Pinna & Wosiacki, 2003).
Trichomycterus caliensis (Eigenmann, 1912)
Pygidium caliense Eigenmann, 1912a: 18. Type locality: Cali, Colombia. Holotype: FMNH 56029.
Distribution: Calima River basin, Colombia (de Pinna & Wosiacki, 2003).
Trichomycterus candidus (Miranda Ribeiro, 1949)
Eremophilus candidus Miranda Ribeiro, 1949b: 2, unnumbered plate. Type locality: pequeno córrego que cai noribeirão Espírito Santo, afluente do Rio Claro e este do Sapucaí, que deságua no Rio Grande — Município deConceição Aparecida, Estado de Minas Gerais, Brazil. Holotype: MNRJ 5209.
Distribution: Grande River basin, Minas Gerais State, Brazil (de Pinna & Wosiacki, 2003).
Remarks: Redescribed as Eremophilus candidus in Barbosa & Costa (2003a).
Trichomycterus castroi de Pinna, 1992
Trichomycterus castroi de Pinna, 1992a: 90, figs. 1–3. Type locality: Branch of the Rio Iguaçu, near the pointcrossed by Curitiba-Paranaguá Road, State of Paraná, Brazil, ca. 25º26’S, 49º06’W. Holotype: MZUSP 36964.
Distribution: Iguaçu River basin, Brazil (de Pinna & Wosiacki, 2003).
Trichomycterus catamarcensis Fernández & Vari, 2000
Trichomycterus catamarcensis Fernández & Vari, 2000: 990. Type locality: Provincia de Catamarca, Departamento
Belén, stream tributary to Laguna Blanca, 11 km northeast from Belén on provincial road 43 along route from
Belén to Antofagasta de la Sierra, near Los Nacimientos de San Antonio (approximately 26º 30'S, 67º03'W),
elevation 3500m, Argentina. Holotype: FML 2507.
Distribution: Laguna Blanca basin in Catamarca Province, Argentina (de Pinna & Wosiacki, 2003).
Trichomycterus caudofasciatus Alencar & Costa, 2004
Trichomycterus caudofasciatus Alencar & Costa, 2004: 5, fig. 2. Type locality: Brazil: Estado de Minas Gerais:
Município de Alto Caparaó, Rio Caparaó, Rio Itabapoana basin, Alto Caparaó, 20º25'53.9"S, 41º51'56.8"W.
Holotype. UFRJ 6002.
Distribution: Upper Itabapoana River basin, southeastern Brazil (Alencar & Costa, 2004).
Trichomycterus celsae Lasso & Provenzano, 2003
Trichomycterus celsae Lasso & Provenzano, 2003: 1140, fig. 1. Type locality: Venezuela, Estado Bolívar, río
Kukenán, cabeceras, valle entre los tepuyes Roraima y Kukenán, Gran Sabana (05º06'30"N, 60º49'48"O). Holo-

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type: MHNLS 6453.
Distribution: Kukenán River, Caroní River basin, Venezuela; known only from type locality (Lasso & Provenzano,2003).
Trichomycterus chaberti Durand, 1968
Trichomycterus chaberti Durand, 1968: 344, fig. 3. Type locality: Umayalanta [Cave], Bolivia. Holotype: MNHN
1968-0217.
Distribution: Umayalanta Cave system, Bolivia (de Pinna & Wosiacki, 2003).
Trichomycterus chapmani (Eigenmann, 1912)
Pygidium chapmani Eigenmann, 1912a: 18. Type locality: Boquia, Colombia. Holotype: FMNH 56027; holotype
illustrated in Eigenmann (1918a: 309, fig. 13; pl. 47, fig. 2).
Distribution: Upper Cauca River basin, and Dagua, Calima and San Juan Rivers, Colombia (Maldonado-Ocampo etal., 2005).
Trichomycterus chiltoni (Eigenmann, 1920)
Pygidium chiltoni Eigenmann, 1920c: 54. Type locality: lower course of the Rio Nonguen where it passes throughthe ground of the Agricultural School, at the outskirts of Concepción, Chile. Lectotype: CAS 57596. Also
described as new in Eigenmann (1927: 40, pl. 8, figs. 1, 1a, and pl. 13, figs. 5–6), with lectotype indicated in
caption of pl. 8: “Type: 15059 I. U. M., 168 mm.”
Distribution: Western drainages in central Chile (de Pinna & Wosiacki, 2003).
Trichomycterus chungaraensis Arratia, 1983
Trichomycterus chungaraensis Arratia, 1983: 67, fig. 2. Type locality: Streams of Vertiente de Mal Paso, Chungará
Lake, 4500 m above sea level, North Chile, South America, Chile. Holotype: FFSUC IC 290878A.
Distribution: Streams of Chungará Lake, Chile (de Pinna & Wosiacki, 2003).
Trichomycterus concolor Costa, 1992
Trichomycterus concolor Costa, 1992: 107, fig. 9. Type locality: ruisseau 20 km au sud de la ville de Garapuava,bassin du rio São Francisco, Minas Gerais, Brazil. Holotype: MZUSP 43347.
Distribution: Upper São Francisco River basin, Brazil (de Pinna & Wosiacki, 2003).
Trichomycterus conradi (Eigenmann, 1912)
Pygidium conradi Eigenmann, 1912b: 212. Type locality: Amatuk, British Guiana. Holotype: FMNH 53721; holotype illustrated in Eigenmann (1918a: 325; pl. 50, fig. 2).
Distribution: River drainages in Guyana and Venezuela (de Pinna & Wosiacki, 2003).
Trichomycterus corduvensis Weyenbergh, 1877
Trichomycterus corduvensis Weyenbergh, 1877: 11, pl. 3 (figs. 1–2). Type locality: Rio Primero, y en las acequias deCórdoba, Argentina. Syntype: MSNG 9020 (1).
Distribution: Primeiro River basin, Argentina (de Pinna & Wosiacki, 2003).
Trichomycterus davisi (Haseman, 1911)
Pygidium davisi Haseman, 1911b: 380, pl. 77; fig. 1. Type locality: Serrinha Paraná, Rio Iguaçu system, Brazil.
Holotype: FMNH 60309; holotype illustrated in Eigenmann (1918a: 334; pl. 51, fig. 5).
Distribution: Iguaçu and Ribeira de Iguape River basins, Brazil (de Pinna & Wosiacki, 2003).
Trichomycterus diabolus Bockmann, Casatti & de Pinna, 2004
Trichomycterus diabolus Bockmann, Casatti & de Pinna, 2004: 227, figs. 1–2. Type locality: Brazil: São Paulo: Rio
Paranapanema basin, município de Teodoro Sampaio, Morro do Diabo State Park, Cóão Carlos (22º35'28.0"S,
52º14'38.1"W). Holotype: MZUSP 78860.
Distribution: Paranapanema River basin, southeastern Brazil (Bockmann et al., 2004).
Trichomycterus dispar (Tschudi, 1846)
Pygidium dispar Tschudi, 1846: 22, pl. 3. Type locality: ... meisten Flüssen der Cordillera; ich habe ein Exemplar
ahf einer Höhe von mehr als 14000 Fussü. M. gefangen; aber immer nur auf dem Hochlande zwischen den
beiden Gebirgsketten und am Ostabhange der Anden wie an dem der Cordillera, Peru. Syntype: MHNN 767 (1).
Distribution: Peruvian Andes (de Pinna & Wosiacki, 2003).
Trichomycterus dorsotriatus (Eigenmann, 1918)
Pygidium dorsotriatum Eigenmann, 1918c: 695. Type locality: Villavicencio, Colombia. Holotype: FMNH 58096;

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holotype illustrated in Eigenmann (1918a: 320, pl. 48, fig. 3). Appeared only as Pygidium dorsotriatum in
Eigenmann (1918c) but as Pygidium dorsostriatum in later Eigenmann publications (e. g., Eigenmann, 1918a:320); dorsotriatum regarded as a typographical error in Eschmeyer et al. (1998) and de Pinna & Wosiacki(2003), but no corrigendum by Eigenmann has been reported.
Distribution: River drainages in Villavicencio, Colombia (de Pinna & Wosiacki, 2003).
Trichomycterus duellmani Arratia & Menu-Marque, 1984
Trichomycterus duellmani Arratia & Menu-Marque, 1984: 510, figs. 14–15. Type locality: Río Tupiza, 12.5 km SE
Tupiza, Bolivia, ... 22º33'S, 65º45'W. Holotype: KU 20191.
Distribution: Tupiza River basin, Bolivia (de Pinna & Wosiacki, 2003).
Trichomycterus emanueli (Schultz, 1944)
Pygidium emanueli emanueli Schultz, 1944c: 259, pl. 5 (fig. c). Type locality: Río Chama at Estanques, Estado deMérida, Venezuela. Holotype: USNM 121223.
Distribution: Chama River basin, Venezuela (de Pinna & Wosiacki, 2003).
Trichomycterus fassli (Steindachner, 1915)
Pygidium fassli Steindachner, 1915a: 200. Type locality: Rio Songo im Distrikt Nord-Yungas, Bolivia. Syntypes (4):
NMW 44470 (1). Illustrated and described in more detail as Pygidium fasslii in Steindachner (1915e: 97, pl. 13,
figs. 1–2).
Distribution: Songo River basin, Bolivia (de Pinna & Wosiacki, 2003).
Trichomycterus gabrieli (Myers, 1926)
Pygidium gabrieli Myers, 1926: 151. Type locality: São Gabriel rapids, Rio Negro, in rock-pools, Amazon system,
Brazil. Syntypes: CAS 64583 (4), SU 36556 (1).
Distribution: Upper Negro River, Brazil (de Pinna & Wosiacki, 2003).
Trichomycterus giganteus Lima & Costa, 2004
Trichomycterus giganteus Lima & Costa, 2004: 1, fig. 1. Type locality: Brazil: Estado do Rio de Janeiro: Município
do Rio de Janeiro, Campo Grande, Rio Guandu-Mirim, Rio Guandu basin. Holotype: UFRJ 5999.
Distribution: Upper Guandu River basin, southeastern Brazil (Lima & Costa, 2004).
Trichomycterus goeldii Boulenger, 1896
Trichomycterus goeldii Boulenger, 1896b: 154. Type locality: Colonia Alpina, Organ Mts in the Province Rio Janeiro, at an altitude of nearly 2600 feet, Brazil. Syntypes: BMNH 1896.4.4.7–8 (2).
Distribution: Mountain ranges of coastal basins of Rio de Janeiro State, Brazil (de Pinna & Wosiacki, 2003).
Trichomycterus gorgona Fernández & Schaefer, 2005
Trichomycterus gorgona Fernández & Schaefer, 2005: 69, figs. 1–3. Type locality: Colombia, Departamento Cauca,
Isla Gorgona, freshwater stream near northeast end of island, 02º59'N, 78º11'30"W. Holotype: ANSP 149946.
Distribution: Gorgona Island, Pacific coast of Colombia (Fernández & Schaefer, 2005).
Trichomycterus guaraquessaba Wosiacki, 2005
Trichomycterus guaraquessaba Wosiacki, 2005: 51, fig. 2. Type locality: rio Bracinho, Fazenda Salto Dourado,
Município de Guaraqueçaba, Paraná, Brazil. Holotype: MPEG 7916.
Distribution: Bracinho River, Atlantic coastal basin, southern Brazil (Wosiacki, 2005).
Trichomycterus guianensis (Eigenmann, 1909)
Pygidium guianensis Eigenmann, 1909a: 11. Type locality: Aruataima Falls, Upper Potaro, British Guiana. Holotype: FMNH 52676; holotype illustrated in Eigenmann (1918a: 325, pl. 50, fig. 1) as Pygidium guianense.
Distribution: Rivers of the Guianas and Venezuela (de Pinna & Wosiacki, 2003).
Trichomycterus hasemani (Eigenmann, 1914)
Pygidium hasemani Eigenmann, 1914a: 48. Type locality: Santarem, Pará, Brazil. Holotype: FMNH 56424; holotype illustrated in Eigenmann (1918a: 326, pl. 50, fig. 4).
Distribution: Amazon River basin (de Pinna & Wosiacki, 2003).
Trichomycterus heterodontus (Eigenmann, 1918)
Pygidium heterodontum Eigenmann, 1918c: 692. Type locality: Rio Mendoza, Palmira, Argentina, 900 m, Argentina. Holotype: CAS 58139; holotype illustrated in Eigenmann (1918a: 296, pl. 44, fig. 4).
Distribution: Mendoza River basin, Argentina (de Pinna & Wosiacki, 2003).

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Trichomycterus iheringi (Eigenmann, 1918)
Pygidium iheringi Eigenmann, 1918c: 697. Type locality: São Paulo in coastal streams and Parana basin, Brazil.
Holotype: CAS 64585; holotype illustrated in Eigenmann (1918a: 330, pl. 50, fig. 5) and Wosiacki (2005: 58,
fig. 3).
Distribution: Ribeira do Iguape River basin, Brazil (de Pinna & Wosiacki, 2003).
Remarks: Redescribed in Wosiacki (2005: 56).
Trichomycterus immaculatus (Eigenmann & Eigenmann, 1889)
Pygidium immaculatum Eigenmann & Eigenmann, 1889b: 52. Type locality: Juiz de Fora, Rio Parahybuna, Sao
Matheos, Goyaz [Brazil]. Syntypes (14): MCZ 8266 (1), MCZ 8300 (10), MCZ 8302 (1), MCZ 8305 (1), MCZ
8307 (1). Species illustrated in Eigenmann (1918a: 334, pl. 52, fig.1).
Distribution: Paraíbuna River, Paraíba do Sul River basin, Brazil (de Pinna & Wosiacki, 2003).
Trichomycterus itacambirussu Triques & Vono, 2004
Trichomycterus itacambirussu Triques & Vono, 2004: 165, fig. 2. Type locality: Brazil: Minas Gerais: County ofGrão Mogol, Rio Jequitinhonha basin: drainage of Rio Itacambiruçu: Córrego do Cambral, 42º45'W 16º33'S.
Holotype: MZUSP 58493.
Distribution: Jequitinhonha River basin, Minas Gerais, Brazil (Triques & Vono, 2004).
Trichomycterus itacarambiensis Trajano & de Pinna, 1996
Trichomycterus itacarambiensis Trajano & de Pinna, 1996: 86, figs. 1–2. Type locality: Creek inside Olhos d’Aguacave, Município de Itacarambi, Minas Gerais, Brazil, 15º06'06"S, 44º09'30"W. Holotype: MZUSP 42469.
Distribution: Olhos d’Água Cave in State of Minas Gerais, Brazil (de Pinna & Wosiacki, 2003).
Trichomycterus itatiayae Miranda Ribeiro, 1906
Trichomycterus brasiliensis itatiayae Miranda Ribeiro, 1906: 178, pl. 1 (figs. a–c). Type locality: ... do Itatiaya, ...
em afluente do Parahyba, Rio de Janeiro State, Brazil. Lectotype: MNRJ 792, designated by Caramaschi & Caramaschi (1991: 223).
Distribution: Rivers in Itatiaia Mountains, upper reaches of Paraíba do Sul River basin, Brazil (de Pinna & Wosiacki,2003).
Remarks: Type locality was interpreted by Caramaschi & Caramaschi (1991: 223) as: Ribeirão da Tapera (tributaryof the Ribeirão Bonito, Rio Paraíba do Sul drainage), Mont-Serrat, Itatiaia (ca. 22º27'S, 44º50'W, ca. 800 m
a.s.l.), Rio de Janeiro State, Brazil.
Trichomycterus jacupiranga Wosiacki & Oyakawa, 2005
Trichomycterus jacupiranga Wosiacki & Oyakawa, 2005: 469, figs. 5–6. Type locality: rio do Queimado, ParqueEstadual de Jacupiranga, Cajati [São Paulo, Brazil]. Holotype: MZUSP 67818.
Distribution: Queimado River, São Paulo, Brazil (Wosiacki & Oyakawa, 2005).
Trichomycterus jequitinhonhae Triques & Vono, 2004
Trichomycterus jequitinhonhae Triques & Vono, 2004: 167, fig. 3. Type locality: Brazil: Minas Gerais: County ofCoronel Murta: Rio Jequitinhonha basin: Córrego Laranjeiras, 42º18'W 16º45'S. Holotype: MZUSP 58497.
Distribution: Jequitinhonha River basin, Minas Gerais, Brazil (Triques & Vono, 2004).
Trichomycterus johnsoni (Fowler, 1932)
Pygidium johnsoni Fowler, 1932a: 367, figure on p. 364. Type locality: Descalvados, Matto Grosso, Brazil. Holotype: ANSP 53873.
Distribution: Paraná River basin, Mato Grosso, Brazil, and Corrientes, Argentina (de Pinna & Wosiacki, 2003).
Trichomycterus knerii Steindachner, 1882
Trichomycterus Knerii Steindachner, 1882a: 142. Type locality: Canelos [Ecuador]. Syntype: NMW 43328 (1). Species illustrated and described in more detail in Steindachner (1882d: 81, pl. 5, figs. 1, 1a).
Distribution: Canelos, Ecuador (de Pinna & Wosiacki, 2003).
Trichomycterus landinga Triques & Vono, 2004
Trichomycterus landinga Triques & Vono, 2004: 162, fig. 1. Type locality: Brazil: Minas Gerais: County of Coronel
Murta: Rio Jequitinhonha basin: Córrego Moquém, 42º35'W 16º45'S. Holotype: MZUSP 58496.
Distribution: Jequitinhonha River basin, Minas Gerais, Brazil (Triques & Vono, 2004).
Trichomycterus latidens (Eigenmann, 1918)

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Pygidium latidens Eigenmann, 1918c: 693. Type locality: Small creek near mouth of Rio Calima, Colombia. Holotype: IU 13801 (whereabouts unknown); holotype illustrated in Eigenmann (1918a: 312, pl. 47, fig. 4).
Distribution: Calima River basin, Colombia (de Pinna & Wosiacki, 2003).
Trichomycterus latistriatus (Eigenmann, 1918)
Pygidium latistriatum Eigenmann, 1918c: 696. Type locality: Quebrada de Pinchote, Santander, Colombia. Holotype: FMNH 58449; holotype illustrated in Eigenmann (1918a: 321, pl. 48, fig. 4).
Distribution: Gaira, Cesar and Ariguani rivers, Magdalena River basin, Colombia (Maldonado-Ocampo et al.,2005).
Trichomycterus laucaensis Arratia, 1983
Trichomycterus laucaensis Arratia, 1983: 74, fig. 8. Type locality: System of Lauca River, Parinacota, 4,390 m
above sea level, northern Chile. Holotype: FFSUC IC 160878A.
Distribution: Lauca River basin, Chile (de Pinna & Wosiacki, 2003).
Trichomycterus lewi Lasso & Provenzano, 2003
Trichomycterus lewi Lasso & Provenzano, 2003: 1144, fig. 7. Type locality: Venezuela, Estado Bolívar, río
Kukenán, cabeceras, valle entre los tepuyes Roraima y Kukenán, Gran Sabana (05º06'30"N, 60º49'48"O). Holotype: MHNLS 6454.
Distribution: Kukenán River, Caroni River basin, Venezuela; known only from type locality (Lasso & Provenzano,2003).
Trichomycterus longibarbatus Costa, 1992
Trichomycterus longibarbatus Costa, 1992: 104, fig. 4. Type locality: près de la ville de Santa Tereza, Espirito
Santo, Brazil. Holotype: MZUSP 43339.
Distribution: Near Santa Tereza, Espirito Santo State, Brazil (de Pinna & Wosiacki, 2003).
Trichomycterus maracaiboensis (Schultz, 1944)
Pygidium banneaui maracaiboensis Schultz, 1944c: 262, pl. 6 (fig. b). Type locality: Río San Juan near bridge,
south of Mene Grande, tributary to Río Motatán, Maracaibo basin, Venezuela. Holotype: USNM 121227.
Distribution: San Juan River, tributary to Motatán River, Lake Maracaibo basin, Venezuela (de Pinna & Wosiacki,2003).
Trichomycterus mboycy Wosiacki & Garavello, 2004
Trichomycterus mboycy Wosiacki & Garavello, 2004: 8, fig. 5. Type locality: Brazil: State of Paraná: município deFoz do Jordão, Jordão reservoir, rio Jordão near at its mouth, tributary of rio Iguaçu (rio Paraná Basin), 25º45'S,
52º10'W. Holotype: MPEG 6695.
Distribution: Jordão River, Iguaçu River basin, Brazil (Wosiacki & Garavello, 2004).
Trichomycterus meridae Regan, 1903
Trichomycterus meridae Regan, 1903b: 624. Type locality: Merida, Venezuela, and from the Rio Albireggas above
Merida, altitude 3500 metres, Venezuela. Syntypes: BMNH 1903.4.28.35-38 (4), BMNH 1906.6.30.77–79 (3),
USNM 133136 (1).
Distribution: Albirregas River basin, Venezuela (de Pinna & Wosiacki, 2003).
Trichomycterus migrans (Dahl, 1960)
Pygidium migrans Dahl, 1960b: 309, fig. 3. Type locality: in front of Camp 1, in the Guayabero River [Orinoco System, Colombia]. Holotype: ICNMHN 399.
Distribution: Guayabero River, Orinoco River basin, Colombia (de Pinna & Wosiacki, 2003).
Trichomycterus mimonha Costa, 1992
Trichomycterus mimonha Costa, 1992: 106, fig. 7. Type locality: 2 km au nord de la ville de Piquete, rio Benfica,bassin du rio Paraíba, Sao Paulo, Brazil. Holotype: MZUSP 43343.
Distribution: Benfica River, Paraíba do Sul River basin, Brazil (de Pinna & Wosiacki, 2003).
Trichomycterus mirissumba Costa, 1992
Trichomycterus mirissumba Costa, 1992: 107, fig. 8. Type locality: Visconde de Mauá, 1 km à l’est de la ville deMaromba, rio Preto, près du confluent avec le ruisseau Santa Clara, bassin du rio Paraíba, Rio de Janeiro, Brazil. Holotype: MZUSP 43345.
Distribution: Preto River, Paraíba do Sul River basin, Brazil (de Pinna & Wosiacki, 2003).

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Trichomycterus mondolfi (Schultz, 1945)
Pygidium mondolfi Schultz, 1945: 29, fig. 1. Type locality: Quebrado Chacaito near Caracas but in Estado deMirando, Río Tuy system, Venezuela. Holotype: USNM 120377.
Distribution: Tuy River basin, Caribbean Drainage, Venezuela (de Pinna & Wosiacki, 2003).
Trichomycterus motatanensis (Schultz, 1944)
Pygidium emanueli motatanensis Schultz, 1944c: 260, pl. 6 (fig. a). Type locality: Río San Juan at the bridge southof Mene Grande, Motatán system, Maracaibo basin Venezuela. Holotype: USNM 121232.
Distribution: San Juan River, Motatán drainage, Lake Maracaibo basin, Venezuela (de Pinna & Wosiacki, 2003).
Trichomycterus naipi Wosiacki & Garavello, 2004
Trichomycterus naipi Wosiacki & Garavello, 2004: 2, fig. 2. Type locality: Brazil: State of Paraná: município de
Tijucas do Sul, rio São João, tributary of rio da Várzea (Iguaçu Basin), near Lagoinha, 25º55'S, 49º11'W. Holotype: MPEG 6699.
Distribution: da Várzea River, Iguaçu River basin, Brazil (Wosiacki & Garavello, 2004).
Trichomycterus nigricans Valenciennes, 1832
Trichomycterus nigricans Valenciennes, 1832a: 348. Type locality: ruisseaux de Sainte-Catherine du Brèsil, Santa
Catarina, Brazil. Holotype: apparently MNHN b-0251.
Distribution: Coastal drainages in Santa Catarina State, Brazil (de Pinna & Wosiacki, 2003).
Remarks: Species redescribed by Arratia (1998) with MNHN b-0251 listed as the holotype.
Trichomycterus nigromaculatus Boulenger, 1887
Trichomycterus nigromaculatus Boulenger, 1887b: 349. Type locality: Andes of Columbia. Syntypes: BMNH
1880.2.26.16–17 (2).
Distribution: Colombian Andes (de Pinna & Wosiacki, 2003).
Trichomycterus pantherinus Alencar & Costa, 2004
Trichomycterus pantherinus Alencar & Costa, 2004: 2, fig. 1. Type locality: Brazil: Estado do Espírito Santo:
Município de Sant Leopoldina, below waterfall near Fazenda Sete Quedas, Rio da Prata, Rio Santa Maria da
Vitória basin, 20º03'15.9"S, 40º32'20.5"W. Holotype: UFRJ 6001.
Distribution: Santa Maria da Vitória River basin, Southeastern Brazil (Alencar & Costa, 2004).
Trichomycterus paolencis (Eigenmann, 1918)
Pygidium paolence Eigenmann, 1918c: 698. Type locality: Sâo Paulo in the Parana basin and (?) in coastal streams.
... Alto da Serra, Rio Tieté, Sâo Paulo. ... Mogy das Cruces, Rio Tieté, São Paulo, Brazil. Holotype: FMNH
58085; holotype illustrated in Eigenmann (1918a: 332, pl. 51, fig. 3).
Distribution: Paraná River basin, São Paulo State, Brazil (de Pinna & Wosiacki, 2003).
Trichomycterus papilliferus Wosiacki & Garavello, 2004
Trichomycterus papilliferus Wosiacki & Garavello, 2004: 5, figs. 3, 4. Type locality: Brazil: State of Paraná: município de Foz do Jordão, Jordão reservoir, rio Jordão near at its mouth, tributary of rio Iguaçu (rio Paraná Basin),
25º45'S, 52º10'W. Holotype: MPEG 6692.
Distribution: Jordão River, Iguaçu River basin, Brazil (Wosiacki & Garavello, 2004).
Trichomycterus paquequerensis (Miranda Ribeiro, 1943)
Pygidium paquequerense Miranda Ribeiro, 1943: 2, fig. Type locality: rio Paquequer Grande, Estado do Rio de Janeiro [Brazil]. Holotype: MNRJ 1159.
Distribution: Paquequer River, Paraíba do Sul River basin, Brazil (de Pinna & Wosiacki, 2003).
Trichomycterus piurae (Eigenmann, 1922)
Pygidium punctulatum piurae Eigenmann, 1922b: 63. Type locality: Piura, Peru. Holotype: CAS 58119.
Distribution: Piura River basin, Peru (de Pinna & Wosiacki, 2003).
Trichomycterus plumbeus Wosiacki & Garavello, 2004
Trichomycterus plumbeus Wosiacki & Garavello, 2004: 13, fig. 8. Type locality: Brazil: State of Paraná: municípiode Foz do Jordão, Jordão reservoir, rio Jordão near at its mouth, tributary of rio Iguaçu (rio Paraná Basin),
25º45'S, 52º10'W. Holotype: MPEG 6689.
Distribution: Jordão River, Iguaçu River basin, Brazil (Wosiacki & Garavello, 2004).
Trichomycterus potschi Barbosa & Costa, 2003

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Trichomycterus potschi Barbosa & Costa, 2003b: 282, fig. 1. Type locality: Brazil: Estado do Rio de Janeiro: rio das
Cachoeiras, serra do Matutu, Município de Mangaratiba. Holotype: MCP 29061.
Distribution: Coastal river basins between Mangaratiba and Itaguaí, Rio de Janeiro State, Brazil (Barbosa & Costa,
2003b).
Trichomycterus pradensis Sarmento-Soares, Martins-Pinheiro, Aranda & Chamon, 2005
Trichomycterus pradensis Sarmento-Soares, Martins-Pinheiro, Aranda & Chamon, 2005: 291, fig. 1. Type locality:
Brazil: Bahia: Jucuruçu, rio Jucuruçu, 2 km before the city of Jucuruçuon road Itamaraju-Jucuruçu, middle ofrio Jucuruçu basin, 16º50'10"S 40º08'40"W. Holotype: MNRJ 28483.
Distribution: Jucuruçu River, a coastal river basin of SE Brazil (Sarmento-Soares et al., 2005).
Trichomycterus pseudosilvinichthys Fernández & Vari, 2004
Trichomycterus pseudosilvinichthys Fernández & Vari, 2004: 876, figs. 1–2. Type locality: Argentina, Provincia de
La Rioja, Departamento Chilecito, Río Amarillo at Famatina, a small assemblage of houses near Fundición de
Oro Santa Florentina (28º55'S, 67º31'W), on east slope of Sierra de Famatina. Holotype: FML 2588.
Distribution: La Rioja province, Argentina (Fernández & Vari, 2004).
Trichomycterus punctatissimus Castelnau, 1855
Trichomycterus punctatissimus Castelnau, 1855: 49, pl. 24 (fig. 3). Type locality: De l’Araguay. Holotype: MNHN
b-0610.
Distribution: Araguaia River basin, Brazil (de Pinna & Wosiacki, 2003).
Trichomycterus punctulatus Valenciennes, 1846
Trichomycterus punctulatus Valenciennes, in Cuvier & Valenciennes, 1846: 488, pl. 552. Type locality: la rivière deLima, Peru. Syntypes: MNHN 0000-3168 (5).
Distribution: Western Peru (de Pinna & Wosiacki, 2003).
Trichomycterus ramosus Fernández, 2000
Trichomycterus ramosus Fernández, 2000a: 350, fig. 1. Type locality: Catamarca: Departamento Belém: Laguna
Blanca, 3680 m elevation, approximately 26º30'S, 67º03'W, Argentina. Holotype: FML 2070.
Distribution: Laguna Blanca basin, Catamarca Province, Argentina (de Pinna & Wosiacki, 2003).
Trichomycterus regani (Eigenmann, 1918)
Pygidium regani Eigenmann, 1918c: 696. Type locality: Tado, Rio San Juan, Colombia. Holotype: CAS 64591;
holotype illustrated in Eigenmann (1918a: 323, pl. 48, fig. 5).
Distribution: San Juan River basin, Colombia (de Pinna & Wosiacki, 2003).
Trichomycterus reinhardti (Eigenmann, 1918)
Pygidium reinhardti Eigenmann, 1918c: 699. Type locality: Burmier on the Rio Itabira, a tributary of the Rio das
Velhas, Brazil. Holotype: FMNH 58081; holotype illustrated in Eigenmann (1918a: 333, pl. 51, fig. 4).
Distribution: Upper São Francisco River basin, Brazil (de Pinna & Wosiacki, 2003).
Trichomycterus retropinnis Regan, 1903
Trichomycterus retropinnis Regan, 1903b: 624. Type locality: St. Augustin, Andes of Colombia, elev. 5000 feet.
Syntypes (2): BMNH 1899.8.21.12–13 (2).
Distribution: Upper Magdalena River basin, Colombia (Maldonado-Ocampo et al., 2005).
Trichomycterus riojanus (Berg, 1897)
Pygidium riojanum Berg, 1897: 269. Type locality: un arroyo de la Cordillera de La Rioja, Argentina. Holotype:
MACN 5175 (destroyed, Fernández & Schaefer, 2003: 357).
Distribution: La Rioja Range, Argentina (de Pinna & Wosiacki, 2003).
Trichomycterus rivulatus Valenciennes, 1846
Trichomycterus rivulatus Valenciennes, in Cuvier & Valenciennes, 1846: 495. Type locality: ruisseaux qui se jettent
dans le lac de Titicaca, vaste mer alpine, peuplée par les Orestias sans ventrales, ou dans les afluens de l’Apurimac, l’une des sources de l’Amazone ... du Guasacona, Peru. Syntype: MNHN b-0586 (1).
Trichomycterus barbatula Valenciennes, in Cuvier & Valenciennes, 1846: 498. Type locality: Guasacona et du Riode Pontezualo près coroico, par une hauteur de terize à quatorze milles pieds et à una latitude de seize à dix-septdegrés nord. Syntypes: MNHN 0000-4077 (1), MNHN 0000-4078 (5) and MNHN b-0587 (3).
Trichomycterus gracilis Valenciennes, in Cuvier & Valenciennes, 1846: 497. Type locality: Rio de Azangaro près de422 · Zootaxa 1418 © 2007 Magnolia Press

FERRARISGuasacona, dans le Rio de Pontezualo près de Coroico, et enfin dans le lac de la Compucila dans les Andes, àl’ouest de Cuzco, par la hautes de quatorze mille pieds, Peru. Syntypes: MNHN 0000-3129 (2), MNHN 00004063 (2), MNHN a-9766 (6), MNHN b-0588 (2).
Trichomycterus incae Valenciennes, in Cuvier & Valenciennes, 1846: 496. Type locality: Rio Guatanai à Cuzco,
Peru. Syntype: MNHN a-8986 (1).
Trichomycterus pentlandi Castelnau, 1855: 49, pl. 24 (fig. 1). Type locality: lac situé prés de la mission de Sarayacu,
qui communiqué avec la riviér d’Ucayale, Peru. Holotype: MNHN b-0608.
Trichomycterus pictus Castelnau, 1855: 50, pl. 24 (fig. 2). Type locality: grand lac de Titicaca. Syntypes: MNHN b0609 (3).
Trichomycterus pardus Cope, 1874b: 132. Type locality: Upper Amazon. Possible holotype: ANSP 22004.
Trichomycterus poeyanus Cope, 1877: 47. Type locality: Arequipa, Peru. Syntypes: ANSP 21382-83 (2).
Pygidium oroyae Eigenmann & Eigenmann, 1889b: 51. Type locality: Pochachara, Oroya River [Brazil]. Syntypes(8): MCZ 3955 (8).
Trichomycterus eigenmanni Boulenger, 1898c: 8. Type locality: Cumbaca [Brazil]. Holotype: MCZ 8301.
Pygidium quechuorum Steindachner, 1900: 207. Type locality: Arequipa, Río Chile, Südperu. Syntypes (5): at
NMW.
Pygidium tiraquae Fowler, 1940c: 92, fig. 52. Type locality: Tiraque, Cochabamba Department, Bolivia. Holotype:
ANSP 69126.
Pygidium atochae Allen, in Eigenmann & Allen, 1942: 156, pl. 13 (figs. 3–5). Type locality: Rio de Atocha, Atocha,
Bolivia. Holotype: CAS 64576.
Distribution: High-altitude lakes and streams in the central Andean range (including Lakes Titicaca and Poopó),from Lake Junin in the north to Chilean region of Tarapacá in the south, spanning Western Bolivia, Peru andNorthern Chile (de Pinna & Wosiacki, 2003).
Trichomycterus roigi Arratia & Menu-Marque, 1984
Trichomycterus roigi Arratia & Menu-Marque, 1984: 494, figs. 1–10. Type locality: Río Pastos Chicos, Jujuy, northof Argentina; 23º24'S —66º35'W. Holotype: MLP 29-8-83-1.
Distribution: Pastos Chicos River basin, Argentina (de Pinna & Wosiacki, 2003).
Trichomycterus romeroi (Fowler, 1941)
Pygidium romeroi Fowler, 1941c: 4, figs. 6–8. Type locality: Honda, Colombia. Holotype: ANSP 69331.
Distribution: Honda River basin, Colombia (de Pinna & Wosiacki, 2003).
Trichomycterus santaeritae (Eigenmann, 1918)
Pygidium santae-ritae Eigenmann, 1918a: 341, pl. 52 (fig. 5). Type locality: Santa Rita, Rio Preto, Brazil. Holotype:
FMNH 58577.
Distribution: Preto River, Paraíba do Sul River basin, Brazil (de Pinna & Wosiacki, 2003).
Trichomycterus spegazzinii (Berg, 1897)
Pygidium Spegazzinii Berg, 1897: 267. Type locality: Río de Cachi (Provincia de Salta), ..., á una altura de 2500 á
2800 metros sobre el nivel del mar, Argentina. Syntypes: BMNH 1898.9.23.1–2 (2), MACN 4925 (19), MACN
5173 (2), SMF 831 (1).
Distribution: Provinces of Salta and Catamarca in Argentina (de Pinna & Wosiacki, 2003).
Trichomycterus spelaeus Do Nascimiento, Villarreal & Provenzano, 2001
Trichomycterus spelaeus Do Nascimiento, Villarreal & Provenzano, 2001: 21, fig. 1. Type locality: Cueva Punto
Fijo, en el caserío Punto Fijo a 7,5 km al N. del Cerro Yolanda. 590 m.s.n.m., cuenca del Río Guasare, Edo.
Zulia, Venezuela (10º57'10"N 72º28'06"O). Holotype: MBUCV V-29602.
Distribution: Punto Fijo Cave, upper Guasare River basin, Venezuela (de Pinna & Wosiacki, 2003).
Trichomycterus spilosoma (Regan, 1913)
Pygidium spilosoma Regan, 1913d: 468. Type locality: Rio Sipi and Rio Tamana, Colombia. Syntypes (3, 130–250
mm TL): BMNH 1910.7.11.106–107 (2), BMNH 1910.7.11.108 (1, skeleton), BMNH 1910.7.11.15 (1).
Distribution: Sipi and Tamana River basins, Colombia (de Pinna & Wosiacki, 2003).
Trichomycterus stawiarski (Miranda Ribeiro, 1968)
Pygidium stawiarski Miranda Ribeiro, 1968: 1. Type locality: Pequeno córrego pertencente á bacia do Rio Paraná —CHECKLIST OF CATFISHES, RECENT AND FOSSIL

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localidade de Bituruna —Est. do Paraná, Paraná, Brazil. Holotype: MNRJ 9739.
Distribution: Iguaçu River basin, Brazil (de Pinna & Wosiacki, 2003).
Trichomycterus stellatus (Eigenmann, 1918)
Pygidium stellatum Eigenmann, 1918a: 308, pl. 47 (fig. 1). Type locality: Quebrada Sarjento, Colombia. Holotype:
FMNH 58101.
Distribution: Rivers of Andean eastern cordillera, Colombia (de Pinna & Wosiacki, 2003).
Trichomycterus straminius (Eigenmann, 1918)
Pygidium straminium Eigenmann, 1918c: 694. Type locality: Quebrada del Mango, Santander, Colombia. Holotype:
FMNH 58105; holotype illustrated in Eigenmann (1918a: 313, pl. 49, fig. 1) as Pygidium stramineum.
Distribution: Andean streams of Cundinamarca and Santander, Colombia (Maldonado-Ocampo et al., 2005).
Trichomycterus striatus (Meek & Hildebrand, 1913)
Pygidium striatum Meek & Hildebrand, 1913: 78. Type locality: Río Cana at Cana, Panama. Holotype: FMNH
7579.
Pygidium septentrionale Behre, 1928: 309, pl. 18. Type locality: small streams tributary to Rio Chiriqui del Tire
above Caldera, Pacific slope of Panama, altitude about 4,000 feet ... Quebrada Salão, Panama, ca. elev. 4000 ft.
Holotype: FMNH 59522.
Distribution: Rivers of Panama and Costa Rica (de Pinna & Wosiacki, 2003); Catatumbo, Magdalena, Cauca andDagua rivers, Colombia (Maldonado-Ocampo et al., 2005).
Trichomycterus taczanowskii Steindachner, 1882
Trichomycterus Taczanowskii Steindachner, 1882b: 177. Type locality: Rio Huambo, Rio de Tortora [Peru]. Syntype: NMW 43387 (1). Species illustrated and described in more detail in Steindachner (1882c: 22, pl. 4, figs. 1,
1a, 1b).
Distribution: Peru (de Pinna & Wosiacki, 2003).
Trichomycterus taenia Kner, 1863
Trichomycterus taenia Kner, 1863: 228, fig. 16. Type locality: Vom Westabhange der Andes im Staate Ecuador,
Ecuador. Holotype: at NMW. Also appeared in Kner & Steindachner (1864: 52, pl. 6, fig. 1).
Distribution: Ecuadorian Andes (de Pinna & Wosiacki, 2003); Magdalena and Patia River basins, Colombia (Maldonado-Ocampo et al., 2005).
Trichomycterus taeniops Fowler, 1954
Pygidium tenue Fowler, 1945c: 6, figs. 7–9. Type locality: Acobamba, near Tarma at 2000 meters elevation, Rio
Ucayali basin, Peru. Holotype: ANSP 71638. Preoccupied in Trichomycterus by Trichomycterus tenuis Weyenbergh, 1877; replaced by Trichomycterus taeniops Fowler, 1954.
Trichomycterus taeniops Fowler, 1954: 36, fig. 635. Type locality: Acobamba, near Tarma, at 2000 meters elevation,
Rio Ucayali basin, Peru, elev. 2000 m, Peru. Replacement for Pygidium tenue Fowler, 1945; preoccupied in Trichomycterus by Trichomycterus tenuis Weyenburgh, 1877.
Distribution: Ucayali River basin (elevation 2000 m), Peru (de Pinna & Wosiacki, 2003).
Trichomycterus taroba Wosiacki & Garavello, 2004
Trichomycterus taroba Wosiacki & Garavello, 2004: 10, fig. 6. Type locality: Brazil: State of Paraná: município deFoz do Jordão, Jordão reservoir, rio Jordão near at its mouth, tributary of rio Iguaçu (rio Paraná Basin), 25º45'S,
52º10'W. Holotype: MPEG 6689.
Distribution: Jordão River, Iguaçu River basin, Brazil (Wosiacki & Garavello, 2004).
Trichomycterus tenuis Weyenbergh, 1877
Trichomycterus tenuis Weyenbergh, 1877: 12, pl. 3 (figs. a–c). Type locality: una pequeña laguna, en la Sierra de
Córdoba, cerca de la Villa Cruz-del-eje, Argentina. Syntypes: MSNG 8852 (2).
Distribution: Córdoba Sierra, Argentina (de Pinna & Wosiacki, 2003).
Trichomycterus transandianus (Steindachner, 1915)
Pygidium taenia transandianum Steindachner, 1915e: 100, pl. 12 (fig. 6). Type locality: Gebirgsbach im Cañon delGallo, einem rechten Seitentale des Rio Combeima in der Zentral-Cordillere, Columbien, in einer Höhe von1800 m. Syntypes (2): NMW 44475. Originally as Pygidium taenia Var. nova transandianum.
Distribution: Combeima River basin, central Andean cordillera, Colombia (de Pinna & Wosiacki, 2003).

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Trichomycterus trefauti Wosiacki, 2004
Trichomycterus trefauti Wosiacki, 2004: 3, fig. 2. Type locality: riacho Andrequicé, tributary of rio Paraúna, itself atributary of rio das Velhas (rio São Francisco basin), approximately 18º30'S, 43º30'W, Município de Trinta Réis,
Minas Gerais, Brazil. Holotype: MZUSP 79911.
Distribution: Upper São Francisco River basin, Brazil (Wosiacki, 2004).
Trichomycterus triguttatus (Eigenmann, 1918)
Pygidium triguttatum Eigenmann, 1918a: 339, pl. 52 (fig. 4). Type locality: Jacarehy, São Paulo, Brazil. Holotype:
FMNH 58670.
Distribution: São Paulo State, Brazil (de Pinna & Wosiacki, 2003).
Trichomycterus tupinamba Wosiacki & Oyakawa, 2005
Trichomycterus tupinamba Wosiacki & Oyakawa, 2005: 466, figs. 2–4. Type locality: Brazil, São Paulo: rio Betari,tributary of rio Ribeira de Iguape, in front of the camping area of Núcleo Santana, Parque Estadual Turístico do
Alto Ribeira. Holotype: MZUSP 62382.
Distribution: Betari and Iporanga rivers, Ribeira de Iguape River basin, Brazil (Wosiacki & Oyakawa, 2005).
Trichomycterus unicolor (Regan, 1913)
Pygidium unicolor Regan, 1913d: 468. Type locality: Condoto, Colombia. Syntypes: BMNH 1913.10.1.42–43 (2).
Distribution: San Juan River basin, Colombia (de Pinna & Wosiacki, 2003).
Trichomycterus variegatus Costa, 1992
Trichomycterus variegatus Costa, 1992: 103, fig. 3. Type locality: Estado de Minas Gerais: ville de São Roque deMinas, rio do Peixe, affluent du haut rio Sao Francisco [Brazil]. Holotype: MZUSP 42316.
Distribution: Upper São Francisco River basin, Brazil (de Pinna & Wosiacki, 2003).
Trichomycterus venulosus (Steindachner, 1915)
Pygidium venulosum Steindachner, 1915a: 199. Type locality: Paramo de Cruz verde, östliche Cordillere, Columbien, in 3000 m. Syntypes: NMW 44476 (2). Species illustrated and described in more detail in Steindachner(1915e: 85, pl. 121, figs. 3–4).
Distribution: Streams of eastern Andean cordillera, Colombia (Maldonado-Ocampo et al., 2005).
Trichomycterus vermiculatus (Eigenmann, 1918)
Pygidium vermiculatum Eigenmann, 1918c: 699. Type locality: Juiz de Fora, Rio Parahyba, Brazil. Holotype:
FMNH 58077; holotype illustrated in Eigenmann (1918a: 335, pl. 52, fig. 2).
Distribution: Paraíba do Sul River, State of Minas Gerais, Brazil (de Pinna & Wosiacki, 2003).
Trichomycterus vittatus Regan, 1903
Trichomycterus vittatus Regan, 1903b: 623. Type locality: Marcapata Valley, E. Peru. Holotype: BMNH
1902.5.29.210.
Distribution: Eastern Peru (de Pinna & Wosiacki, 2003).
Trichomycterus weyrauchi (Fowler, 1945)
Pygidium weyrauchi Fowler, 1945c: 7, figs. 10–12. Type locality: Acobamba, near Tarma, at 2200 meters elevation,
Rio Ucayali drainage, Peru. Holotype: ANSP 71639.
Distribution: Ucayali River basin (elev. 2900 m), Peru (de Pinna & Wosiacki, 2003).
Trichomycterus yuska Fernández & Schaefer, 2003
Trichomycterus yuska Fernández & Schaefer, 2003: 355, figs. 1–2. Type locality: Argentina: Provincia de Catamarca, Departamento Tinogasta, Arroyo Aguas Calientes, 4050 m elevation. Holotype: FML 2535.
Distribution: Known only from the type locality, a high elevation stream in western Argentina (Fernández &Schaefer, 2003).
Trichomycterus zonatus (Eigenmann, 1918)
Pygidium zonatum Eigenmann, 1918a: 330, pl. 51 (fig. 1). Type locality: Agua Quente, São Paulo, Brazil. Holotype:
FMNH 58573.
Trichomycterus cubataonis Bizerril, 1994: 618, figs. 1–3. Type locality: Rio Cubatão, Joinville, Estado de Santa
Catarina [Brazil]. Holotype: MNRJ 12490.
Distribution: Coastal rivers from Santa Catarina to São Paulo States, Brazil (de Pinna & Wosiacki, 2003).

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TRIDENS Eigenmann & Eigenmann, 1889
Tridens Eigenmann & Eigenmann, 1889b: 53. Type species: Tridens melanops Eigenmann & Eigenmann, 1889.
Type by original designation. Gender: Masculine.
Tridens melanops Eigenmann & Eigenmann, 1889
Tridens melanops Eigenmann & Eigenmann, 1889b: 53. Type locality: Iça [Brazil]. Syntypes (27): CAS 64598 (1),
BMNH 1889.11.14.73 (1), MCZ 8137 (1), MCZ 1566385 (6), MCZ 156639 (2, c&s), MZUSP uncat [ex. MCZ
156638] (1), USNM 41522 (1), USNM 120296 (3).
Distribution: Amazon River basin, Brazil (de Pinna & Wosiacki, 2003).
TRIDENSIMILIS Schultz, 1944
Tridensimilis Schultz, 1944c: 266. Type species: Tridensimilis venezuelae Schultz, 1944. Type by original designation. Gender: Masculine.
Tridensimilis brevis (Eigenmann & Eigenmann, 1889)
Tridens brevis Eigenmann & Eigenmann, 1889b: 54. Type locality: Tabatinga [Brazil]. Holotype: MCZ 8160.
Distribution: Amazon River basin, Brazil (de Pinna & Wosiacki, 2003).
Tridensimilis venezuelae Schultz, 1944
Tridensimilis venezuelae Schultz, 1944c: 267, pl. 6 (fig. c). Type locality: Río Negro, below the mouth of the Río
Yasa, Maracaibo basin, Venezuela. Holotype: USNM 121290.
Distribution: Orinoco River basin, Venezuela (de Pinna & Wosiacki, 2003).
TRIDENTOPSIS Myers, 1925
Tridentopsis Myers, 1925b: 84. Type species: Tridentopsis pearsoni Myers, 1925. Type by original designation.
Gender: Feminine.
Tridentopsis cahuali Azpelicueta, 1990
Tridentopsis cahuali Azpelicueta, 1990: 982, figs. 1–4. Type locality: Estancia El Bagual, Formosa Province, Argentina, 26º10'53"S, 58º56'39"W. Holotype: MLP 5-IX-89-1.
Distribution: Paraguay River basin, Argentina (de Pinna & Wosiacki, 2003).
Tridentopsis pearsoni Myers, 1925
Tridentopsis pearsoni Myers, 1925b: 84. Type locality: Lagoons at Lake Rogoagua, Bolivia. Holotype: CAS 28258.
Distribution: Upper Amazon River basin, Bolivia (de Pinna & Wosiacki, 2003).
Tridentopsis tocantinsi La Monte, 1939
Tridentopsis tocantinsi La Monte, 1939: 1. Type locality: Rio Tocantins, northeastern Brazil. Holotype: AMNH
13967.
Distribution: Tocantins River basin, Brazil (de Pinna & Wosiacki, 2003).
TYPHLOBELUS Myers, 1944
Typhlobelus Myers, 1944: 593. Type species: Typhlobelus ternetzi Myers, 1944. Type by original designation. Gender: Masculine.
Key: Schaefer et al. (2005).
Typhlobelus guacamaya Schaefer, Provenzano, de Pinna & Baskin, 2005
Typhlobelus guacamaya Schaefer, Provenzano, de Pinna & Baskin, 2005: 14, figs. 7, 8. Type locality: Venezuela,
Estado Amazonas, Río Cuao at Raudal Guacamaya, 8.1 miles upstream from Raudal El Danto, 05º07.71'N,
67º31.53'W. Holotype: MBUCV V-30936.
Distribution: Cuao River, Orinoco River basin, Venezuela (Schaefer et al., 2005).
Typhlobelus lundbergi Schaefer, Provenzano, de Pinna & Baskin, 2005
Typhlobelus lundbergi Schaefer, Provenzano, de Pinna & Baskin, 2005: 10, figs. 5, 6. Type locality: Venezuela,
Estado Delta Amacuro, Río Orinoco at Los Castillos, upstream of east end of Caño Limon, 8º31.2'N, 62º35.1'W.
Holotype: MBUCV V-31040.
Distribution: Lower Orinoco River, between Ciudad Bolivar and Los Castillos, Venezuela (Schaefer et al., 2005).
Typhlobelus macromycterus Costa & Bockmann, 1994

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Typhlobelus macromycterus Costa & Bockmann, 1994b: 68, figs. 1–3. Type locality: Brazil: Estado do Pará, Rio
Tocantins near Tucuruí. Holotype: MNRJ 12129.
Distribution: Tocantins River near Tucuruí, Pará State, Brazil (de Pinna & Wosiacki, 2003).
Typhlobelus ternetzi Myers, 1944
Typhlobelus ternetzi Myers, 1944: 593, pl. 52 (fig. 2), pl. 53 (figs. 6–8). Type locality: Rock pools below São Gabriel
Rapids, Rio Negro, Brazil. Holotype: CAS 11118.
Distribution: Upper Negro River basin, Brazil (de Pinna & Wosiacki, 2003).
VANDELLIA Valenciennes, 1846
Vandellia Valenciennes, in Cuvier & Valenciennes, 1846: 386. Type species: Vandellia cirrhosa Valenciennes, 1846.
Type by monotypy. Gender: Feminine.
Urinophilus Eigenmann, 1918a: 358. Type species: Vandellia sanguinea Eigenmann, 1918. Type by subsequent designation by Eigenmann (1920d: 441). Gender: Masculine. Originally proposed without any included species;
species first assigned, and a type species designated, in Eigenmann (1920d).
Vandellia beccarii Di Caporiacco, 1935
Vandellia beccarii Di Caporiacco, 1935: 59. Type locality: Rockstone, in flumine Essequibo dicto, in Guiana Britannica. Holotype: MZUF 5506.
Distribution: Orinoco River basin and rivers of Guyana (de Pinna & Wosiacki, 2003).
Remarks: Redescribed in Schmidt (1987).
Vandellia cirrhosa Valenciennes, 1846
Vandellia cirrhosa Valenciennes, in Cuvier & Valenciennes, 1846: 386, pl. 547. Type locality: [Probably from
America]. Syntypes (2): MNHN a-6308.
Vandellia gigantea Cornalia, 1849: 15, figs. 4–5. Type locality: [Fl. Amazonum et Napo]. Type(s): lost (Cagnolaro& Violani, 1988).
Vandellia plazaii Castelnau, 1855: 51, pl. 28 (fig. 1). Type locality: rio Ucayale (Pérou). Holotype: MNHN a-6309.
Vandellia Balzanii Perugia, 1897: 23. Type locality: Rio Beni, Missioni Mosetenes, Bolivia. Holotype: MSNG 8848.
Urinophilus erythrurus Eigenmann, 1922b: 114, pls. 3 (figs. 5–7), 4 (10–16). Type locality: Río Morona, Peru.
Holotype: CAS 64599.
Distribution: Amazon River basin (de Pinna & Wosiacki, 2003).
Remarks: Vandellia gigantea Cornalia, 1849, was treated as a nomen oblitum by Cagnolaro & Violani (1988), but
considered to be available, albeit a junior synonym, in de Pinna & Wosiacki (2003).
Vandellia sanguinea Eigenmann, 1918
Vandellia sanguinea Eigenmann, 1918c: 701. Type locality: San Antonio de Rio Madeira. Holotype: FMNH 58086;
holotype illustrated in Eigenmann (1918a: 365, pl. 53, fig. 2).
Distribution: Amazon, Orinoco and Essequibo River basins (de Pinna & Wosiacki, 2003).
Genus inquirendum, Trichomycteridae
Pygidium Meyen, 1835, in Meyen, 1834–35: 475. Type species: Pygidium fuscum Meyen, 1835. Type by monotypy.
Gender: Neuter. Familial assignment of this genus is uncertain (cf. Tchernavin, 1944).
Remarks: The family group name Pygidiidae was proposed in Eigenmann & Eigenmann (1888a: 649) on Pygidium
Meyen, 1835.
Species inquirenda, Trichomycteridae
Pygidium fuscum Meyen, 1835,in Meyen, 1834–35: 369. Type locality: Peru. Holotype: Whereabous unknown. Familial
assignment of this species is uncertain (see Tchernavin, 1944).

UNPLACED NAMESSILURIFORMES Incertae sedis
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† BACHMANNIA Dolgopol de Saez, 1945
† Bachmannia Dolgopol de Saez, 1945: 453. Type species: † Bachmannia chabutensis Dolgopol de Saez, 1945.
Type by Monotypy. Gender: Feminine.
† Bachmannia chabutensis Dolgopol de Saez, 1945
† Bachmannia chabutensis Dolgopol de Saez, 1945: 453, figure on p. 453. Type locality: de la laguna del Hunco ydel Mirador de Chubut [Argentina]; Tertiary. Holotype: MLP 40-V-17-1a (impression).
† Arius argentinus Dolgopol de Saez, 1945: 455, figure on p. 455. Type locality: La Laguna del Huncho y del Niradorde, Chabut, Argentina; Tertiary. Holotype: MLP 40-V-17-3 (impression of cranium and partial vertebral column).
Distribution: Miocene of Argentina (Gayet & Meunier, 2003).
Remarks: Synonymy from Gayet & Meunier (2003), which appears to be based on Arratia & Cione (1996).
CONORHYNCHOS Bleeker, 1858
Conostome Duméril, 1856: 484. Type species: Pimelodus conirostris Cuvier, 1836. Type by original designation.
Gender: Feminine. Possibly preoccupied by Conostoma Hodgson, 1842, in birds and not treated as valid inrecent times. Considered a nomen oblitum.
Conorhynchos Bleeker, 1858: 191, 205, 209. Type species: Pimelodus conirostris Cuvier, 1836. Type by monotypy.
Gender: Neuter. New name for Conostome Duméril, which is apparently preoccupied by Conostoma Hodgson,
1842, in birds. Name often misspelled Conorhynchus following the spelling in Bleeker (1862-63).
Conorhynchus Bleeker, 1863 (in Bleeker, 1862-63): 12. Type species: Pimelodus conirostris Cuvier, 1836. Type byoriginal designation. New spelling for Conorhynchos. Preoccupied by Conorhynchus Motschousky, 1860, in
Coleoptera.
Conorhynchichthys Regan, 1908, in Regan, 1906-08: 192. Type species: Pimelodus conirostris Cuvier, 1836. Typeby being a replacement name. Gender: Masculine. Replacement for Conorhynchus Bleeker, 1863 [=Conorhynchos Bleeker, 1858], preoccupied by Conorhynchus Motschousky, 1860.
Remarks: Initially included in the Pimelodidae, but that placement questioned in Reis et al. (2003).
Conorhynchos conirostris (Cuvier, 1836)
Pimelodus conirostris Cuvier, 1836: 541, footnote 12. Type locality: rivière de Saint-François. Holotype: MNHN a9413. Described and illustrated in Cuvier & Valenciennes (1840b: 204 (156 in Strasbourg deluxe edition), pl.
436).
Conorhynchus glaber Steindachner, 1877b: 637, pl. 8. Type locality: einem flusse bei Porto Seguro [apparently inerror]. Holotype: at NMW.
Distribution: São Francisco River basin, Brazil.
† EOPEYERIA Whitley, 1947
† Ariopsis Peyer, 1928: 43. Type species: † Ariopsis aegyptiacus Peyer, 1928. Type by monotypy. Gender: Feminine. Preoccupied by Ariopsis Gill, 1861, in Recent fishes. Replaced by † Peyeria Whitley, 1940.
† Peyeria Whitley, 1940a: 242. Type species: † Ariopsis aegyptiacus Peyer, 1928. Type by being a replacement
name. Gender: Feminine. Replacement for † Ariopsis Peyer, 1928, preoccupied by Ariopsis Gill, 1861, in
Recent fishes. But, † Peyeria Whitley preoccupied by † Peyeria Weiler, 1935, in fossil Chondrichthyes.
† Eopeyeria Whitley, 1947: 150. Type species: †Ariopsis aegyptiacus Peyer, 1928. Type by being a replacement
name. Gender: Feminine. Replacement for † Peyeria Whitley, 1940, preoccupied by † Peyeria Weiler, 1935, infossil Chondrichthyes.
† Eopeyeria aegyptiaca (Peyer, 1928)† Ariopsis aegyptiacus Peyer, 1928: 43, pl. 5, pl. 6 (fig. 1). Type locality: Qasr-es-Sagha-Stufe, Norden des Fajum.
Ägyptens; Eocene. Holotype: at Bayerische Staatssammlung, München.
Distribution: Fajum, Egypt; Eocene.
† FAJUMIA Stromer, 1904
† Fajumia Stromer, 1904: 3. Type species: † Fajumia schweinfurthi Stromer, 1904. Type by monotypy. Gender:

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Feminine.
Remarks: Placed by Jordan (1923) into the Bagridae, but placed in the Ariidae by Peyer (1928). Gayet & Meunier(2003) place this genus as Siluriformes incertae sedis, possibly belonging within the Arioida.
† Fajumia menoni Sahni & Mishra, 1975
† Fajumia menoni Sahni & Mishra, 1975: 7, pl. 1 (figs. 3 a–c). Type locality: Bluish-grey stage (Middle Eocene) at
Gabhatad, Western India; Middle-Eocene. Holotype: LUVP 11140 (cranium).
Distribution: Western India, middle Eocene (Sahni & Mishra, 1975).
Remarks: Placed in the Bagridae by Sahni & Mishra (1975), following placement of genus in that family by Jordan(1923).
† Fajumia misrai Sahni & Mishra, 1975
† Fajumia misrai Sahni & Mishra, 1975: 8, pl. 1 (figs. 4–5). Type locality: Bluish-grey shales of Babia Stage (Middle Eocene) at Nareda, Western India. Holotype: LUVP 11142 (cranium).
Distribution: Western India; middle Eocene (Sahni & Mishra, 1975).
Remarks: Placed in the Bagridae by Sahni & Mishra (1975), following placement of genus in that family by Jordan(1923).
† Fajumia schweinfurthi Stromer, 1904
† Fajumia Schweinfurthi Stromer, 1904: 3, pl. 1 (figs. 1–2). Type locality: Mitteleocän am Nordrande des Fajûm,Aegypten. Holotype: in Munich (? at Bayerische Staatssammlung, München) (nearly complete cranium).
Distribution: Fajum, Egypt; middle Eocene.
† Fajumia stromeri Peyer, 1928
† Fajumia Stromeri Peyer, 1928: 33, fig. 3, pl. 4 (fig. 1). Type locality: Qasr-es-Sagha-Stufe, Norden des Fajum.
Ägyptens; Eocene. Holotype: at Bayerische Staatssammlung, München.
Distribution: Fajum, Egypt; middle Eocene.
HORABAGRUS Jayaram, 1955
Horabagrus Jayaram, 1955b: 261. Type species: Pseudobagrus brachysoma Günther, 1864. Type by original designation. Gender: Masculine.
Remarks: Horabagrus was initially placed in the Bagridae and has subsequently been suggested to be more closely
related to species of the Schilbidae (K. C. Jayaram, pers. comm.) or as a separate family that also includes somespecies currently placed in the Schilbidae (de Pinna, 1998a, Hardmann, 2005).
Horabagrus brachysoma (Günther, 1864)
Pseudobagrus brachysoma Günther, 1864: 86. Type locality: Cochinchina. Holotype: BMNH 1862.9.18.5 (skin).
Pseudobagrus chryseus Day, 1865b: 290. Type locality: Kurriavanoor River, Kurriapudnam or Cochin, India. Holotype: Possibly AMS B.7922 (1), BMNH 1865.7.17.5-6 (2), MCZ 4280 (1), NMW 44177 (1), RMNH 6870 (1),ZMB 9575 (1), ZSI 475 (1, lost). Also described and illustrated in Day (1865a: 185, pl. 13, fig. 2).
Distribution: Vernbanad Lake and estuaries of Kerala and southwestern Karnataka, India (Pethiyagoda & Kottelat,1994).
Remarks: See Jayaram (1952) for taxonomic comments.
Horabagrus nigricollaris Pethiyagoda & Kottelat, 1994
Horabagrus nigricollaris Pethiyagoda & Kottelat, 1994: 110, fig. 13. Type locality: India: Kerala: Chalakudy River,
26 km upstream of Chalakudy town, near Vettilappara. Holotype: AMS I.34198-001.
Distribution: Chalakudy River, Kerala, India (Pethiyagoda & Kottelat, 1994).
PHREATOBIUS Goeldi, 1905
Phreatobius Goeldi, 1905: 549. Type species: Phreatobius cisternarum Goeldi, 1905. Type by monotypy. Gender:
Masculine.
Remarks: Originally treated as a genus of the Trichomycteridae, in recent years this genus has been placed within
the Heptapteridae (e. g., Bockmann & Guazzelli, 2003), following Buckup (1988) and the unpublished study by
Bockmann (1998). However, Muriel-Cunha & de Pinna (2005) suggests that the placement of this genus and its
single named species is unsettled. The family-group name Phreatobinae was proposed for this genus by Reichel

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(1927: 383).
Phreatobius cisternarum Goeldi, 1905
Phreatobius cisternarum Goeldi, 1905: 549. Type locality: Wasser einer Binnenlandzisterne tief im Innern der Mündung des Amazonenstromes vorgelagertern Rieseninsel Marajó [Brazil]. Syntypes (2): MHNG 2623.30 (1),plus one of the following: FMNH 58580 (1), MHNG 1213.97 (1), or MHNG 1505.91 (2 entire specimens plus
one sectioned and mounted on microscopic preparations).
Distribution: Marajó island, mouth of the Amazon River, Brazil, in cisterns (Bockmann & Guazzelli, 2003).
Remarks: See Reichel (1927), Carvalho (1967) and Muriel-Cunha & de Pinna (2005) for further information on thisenigmatic species.
† PLIOSILURUS Weiler, 1956.
† Pliosilurus Weiler, 1956: 180. Type species: † Pliosilurus primus Weiler, 1956. Type by monotypy. Gender: Masculine.
Remarks: Listed as † Pliosilurus primus gen. & sp. nov., which, if published before 1931, would have made the typedesignation by original designation, but not after 1930.
† Pliosilurus primus Weiler, 1956
† Pliosilurus primus Weiler, 1956: 180, figs. 1–2; pl. 11. Type locality: Willershausen bek Göttingen [Pliocene, Germany]. Holotype: Geologischen Instituts der Universität Göttingen 4836 (head, pectoral girdle, portion of vertebral column and anal fin).
Distribution: Germany; Pliocene.
† RHINEASTES Cope, 1872
† Rhineastes Cope, 1872b: 486. Type species: † Rhineastes peltatus Cope, 1872. Type by original designation. Gender: Masculine.
Remarks: † Rhineastes has generally been placed in the Ariidae, but Lundberg (1992: 396) considered that placement in error and, instead, treated the genus as Incertae sedis in the Siluriformes, which is followed here.
† Rhineastes peltatus Cope, 1872
† Rhineastes peltatus Cope, 1872b: 486. Type locality: Bridger Formation, South Bitter Creek, Wyoming. Holotype
USNM 3984 (supraoccipital bone and dorsal spine); holotype illustrated in Cope (1884b: 63, pl. 5, figs. 1–2).
Distribution: Bridger Formation, Wyoming; known only from type locality (Lundberg, 1975).
† Rhineastes smithii Cope, 1872
† Rhineastes smithii Cope, 1872b: 486. Type locality: Bridger Formation, South Bitter Creek, Wyoming. Holotype:
USMN 3978 (pectoral spine); holotype illustrated in Cope (1884b: 64, pl. 5, figs. 5–11).
Distribution: Bridger Formation, Wyoming (Lundberg, 1975).
Remarks: Inclusion in † Rhineastes considered tentative by Lundberg (1975).
Species inquirenda, † Rhineastes† Rhineastes radulus Cope, 1873: 639. Type locality: Bridger Formation at Cottonwood Creek, Wyoming; Eocene. Holotype: USNM 4099 (fragments); holotype illustrated in Cope (1884b: 67, pl. 5, figs. 14–17). Identity uncertain (Lundberg, 1975).
† SOCNOPAEA Stromer, 1904
† Socnopaea Stromer, 1904: 6. Type species: † Socnopaea grandis Stromer, 1904. Type by monotypy. Gender: Feminine.
Remarks: Jordan (1923) placed this genus and † Fajumia into the Bagridae, but Peyer (1928) placed both genera in
the Ariidae.
† Socnopaea grandis Stromer, 1904
† Socnopaea grandis Stromer, 1904: 6, pl. 1 (fig. 3). Type locality: Mitteleocän am Nordrande des Fajûm, Aegypten.
Holotype: in Munich (partial cranium).
Distribution: Fajum, Egypt; middle Eocene.

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† Socnopaea horai Sahni & Mishra, 1975: 10, pl. 2 (fig. 1). Type locality: Ossiferus gypseous shales of Babia stage(Middle Eocene) at Harudi, Western India. Holotype: LUVP 11145 (cranium).
Genera inquirendae, Siluriformes
Tachysurus La Cepède, 1803: 150. Type species: Tachysurus sinensis La Cepède, 1803. Type by monotypy. Gender:
Masculine. The type species is considered to be an unrecognizable siluriform by Wheeler & Baddokwaya (1981)
and Taylor (1986a).
Laimumena Sauvage, 1884a: 147. Type species: Laimumena barbonica Sauvage, 1884. Type by monotypy. Gender:
Feminine. Description of type species is not recognizable to genus or even family.
Trachymochlus Hoedeman, 1961: 137. Type species: Trachymochlus cupido Hoedeman, 1961. Type by monotypy. Gender: Masculine.
Species inquirendae, Siluriformes
Silurus undecimalis Linnaeus, 1758: 305. Type locality: Not stated. Type(s): originally part of collection at Museum
Adolphi Friderici but whereabouts unknown, apparently not at NRM. Type locality stated as “Surinami” in Linnaeus(1764: 97).
Silurus luvur Molina, 1782: 346. Type locality: Chile. No types known.
Silurus schilby Sonnini, 1799: 256, pl. 23 (fig. 1). Type locality: Nil. On p. 295 there is note about Silurus schilbe niloticus of Hasselquist that suggests that this may not be intended to be the proposal of a new name.
Silurus gurgu Bloch & Schneider, 1801: 388. Type locality: in Nilo. No types known, based on literature account. Proposed conditionally.
Silurus minutus Schneider, in Bloch & Schneider, 1801: 389. Type locality: India. No types known, based on unpublished literature source. Proposed conditionally.
Tachysurus sinensis La Cepède, 1803: 150, 151, pl. 5 (fig. 2). Type locality: China. No types known; based on a Chinese
painting; illustration in La Cepède reproduced in Wheeler & Baddokwaya (1981). Treated in many Chinese works
as valid in Arius, but Wheeler & Baddokwaya (1981) and Taylor (1986a) indicate that it is an unrecognizable siluriform from the description and/or illustration.
Silurus ichneumon Hermann, 1804: 309. Type locality: Cairo. No types known.
Pimelodes fossor Lichtenstein, 1823: 112. Type locality: Cape of Good Hope, South Africa. Type(s): at ZMB.
Pimelodus octocirrhus Cuvier, 1829: 294. Type locality: not stated. Available from footnote as, “Pim. octocirrhus, N.,
Seb., III, xxix, 1” [= Seba (1734–65: pl. 29, fig. 1)].
Pimelodus javus Valenciennes, in Cuvier & Valenciennes, 1840b: 187 (139 in Strasbourg deluxe edition). Type locality:
Java. Holotype (4 ½ pouces): at MNHN. Status uncertain (Roberts, 1993: 47).
Pimelodus tachisurus Valenciennes, in Cuvier & Valenciennes, 1840b: 163 (121 of Strasbourg deluxe edition). Type
locality: China. No types known; based on a Chinese painting. Unneeded new name for Tachisurus chinensis [sic,for sinensis] La Cepède.
Pimelodus Cantonensis Valenciennes, in Cuvier & Valenciennes, 1840b: 142 (106 of Strasbourg deluxe edition). Type
locality: Canton [China]. No types known. Based solely on an illustration.
Bagrus chinta Valenciennes, in Cuvier & Valenciennes, 1840a: 445 (330 of Strasbourg deluxe edition). Type locality:[Vizagapatam]. No types known, based on account and illustration in Russell (1803: pl. 167).
Pimelodus pusillus Ranzani, 1842: 332, pl. 27. Type locality: Unknown [apparently Brazil, based on other species inpublication]. Holotype: MZUB 933. Name appeared first in Ranzani (1841: 64) as a nomen nudum.
† Pimelodus Sadleri Heckel, 1849: 19. Type locality: Bihar Comitat, Hungary; Miocene. Holotype ? (fin ray fragments).
Illustrated and described in more detail in Heckel (1850: 213, pl. 16, fig. 3).
Silurus ferox Gronow, in Gray, 1854: 135. Type locality: not stated. No types known.
Rhamdia javanica Bleeker, 1858b: 139. Type locality: Java. Holotype: at MNHN. Unjustified emendation of Pimelodus
javus Valenciennes, 1840.

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Laimumena barbonica Sauvage, 1884a: 147. Type locality: Réunion. No types known. Not found in catalogs of MNHN
(pers. obs.) and not seen on shelves.
† Silurus Serdicensis Toula, 1889: 108, pl. 9. Type locality: im Westen von Sofia. Upper Tertiary. Syntypes: (fragmentary
remains).
† Silurus gaudryi Leriche, 1900: 181, pl. 1, figs. 7–12. Type locality: Cuis, Monthelon [Marne, Lower Eocene]. Types: atlaboratoire de Géologie de l’Université de Lille (dorsal fin spines, pectoral fin spine). Originally Silurus (?)gaudryi.
Remarks: As Pimelodus gaudryi in Leriche (1901:165).
† Silurus pliocaenicus Leidenfrost, 1925, 121, text fig. 2 and pl. 6. Type locality: pannonisch-pontischen Tonen von
Rákos bei Budapest [Hungary, Pliocene]. Holotype: at Samml. der Kön. ung. Geologischen Anstalt, Budapest, partial neurocranium.
Remarks: Possibly a bagrid (S. L.), or a silurid (Gayet & Meunier, 2003: 503).
† Silurus stenocephalus Leidenfrost, 1925, 122, pl. 7. Type locality: Pannonisch-pontischen Tonen von Rákos bei Budapest [Hungary, Pliocene]. Holotype: at Samml. der Kön. ung. Geologischen Anstalt, Budapest, partial neurocranium.
† Rhineastes grangeri Hussakof, 1932: 17, fig. 26. Type locality: Tung Gur beds, 50 miles southeast of Iren Dabasu,
Inner Mongolia; Pliocene. Holotype: at AMNH (right pectoral fin spine).
Distribution: Known only from the type locality, in central Asia.
Trachymochlus cupido Hoedeman, 1961: 137, fig. 4. Type locality: Nickerie, entrance of Cupido creek into Maratakka
River, Suriname. Holotype: ZMA 102236.
† Schmidelia graciliformis Berry. [no other information; from Dolgopol de Saez (1945)].
† Clarias pliocaenicus Sauvage (in Depéret, 1885). [no other information; from Gayet & Meunier (2003)]† Ariopsis peyeria Eocene, North Africa [no additional information, From Romer. Probably a mistake for Ariopsis Peyer]

Names wrongly treated as Siluriformes† Arius lemoinei of Eocene of France [No additional information]
Remarks: Considered an acipenserid by Gayet & Meunier (2003).
† BRACHYSPONDYLUS Marck, in Marck & Schlüter, 1868: 283. Type species: † Brachspondylus cretaceus Marck,
1868. Type by monotypy. Gender: Masculine.
Remarks: Listed by Jordan (1923: 643) as a cyprinid with †Brachyspondylus saropterix Marck, 1876, as orthotype.
Brachyspondylus saropterix has been considered to be a siluriform taxon, most recently by Gayet & Meunier(2003), but † Brachyspondylus is actually available from Marck & Schlüter (1868: 283), with † Brachyspondylus cretaceus (283, pl. 43, fig. 2) as type by monotypy. This species is a scaly fish, probably a cyprinid, so that †Brachyspondylus is not a catfish name. See Sanders (1934: 30) for discussion of the taxonomic history of thisname.
† BUCKLANDIUM König, 1825
† Bucklandium König, 1825: 4. Type species: † Bucklandium diluvii König, 1825. Type by monotypy. Gender: Neuter.
† Glyptocephalus Agassiz, 1843 (in Agassiz, 1833–43): 264. Type species: † Glyptocephalus radiatus Agassiz,
1843. Type by monotypy. Gender: Masculine. Preoccupied by Glyptocephalus Gottsche, 1834, in Recent fishes(Pleuronectidae).
† Glyptocara Gill, 1888: 926. Replacement for † Glyptocephalus Agassiz, 1844, preoccupied by Glyptocephalus
Gottsche, 1834.
Remarks: The name Ephippus Owenii appears in Agassiz (1843 (in Agassiz, 1833–43): 264) in the account of the
newly proposed name Glyptocephalus radiatus. It is not treated as a valid name and is therefore not available
from this usage. Gayet & Meunier (2003) place † Bucklandium as Siluriformes incertae sedis, possibly belonging within the Arioida, which appears to follow Woodward (1889b: 208), in which the name is treated as a siluriform fish near to Auchenoglanis. However, the illustration provided by Woodward is apparently not that of a432 · Zootaxa 1418 © 2007 Magnolia Press

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siluriform fish and after close examination of the type specimen, Ralf Britz (pers. commun.) concluded that thebones are definitely not those of a catfish.
† Bucklandium diluvii König, 1825
† Bucklandium diluvii König, 1825: 4, pl. 8, no. 91. Type locality: Isle of Sheppey, Lower Eocene, London Clay.
Holotype: BMNH P.9230 (cranium and pectoral arch). Redescribed and illustrated in Woodward (1889b: 208,
pl. 22).
† Glyptocephalus radiatus Agassiz, 1843 (in Agassiz, 1833–43): 264. Type locality: [Argiles de Londres] Sheppy.
Holotype: BMNH P.9230 (cranium and pectoral arch).
Distribution: London Clay, Lower Eocene.
Remarks: † Glyptocephalus radiatus Agassiz, 1843, is an unneeded new name for † Bucklandium diluvii König,1825, which Agassiz apparently did not associate with the examined specimen.
† PROPYGIDIUM Bocchino, 1964: 186. Type species: † Propygidium primaevus Bocchino, 1964. Type by original designation. Gender: Neuter.
Remarks: Although originally described as a member of the siluriform family Trichomycteridae, the material was
reexamined and interpreted as a member of the Perciformes by Cione & Torno (1988).
† Propygidium primaevus Bocchino, 1964: 186, fig. on p. 189. Type locality: Serie Andesítica Lask Bayas, to the west of
Rio Negro Province, Argentina; Eocene/Oligocene. [Corrected to: Cerro David, 0.5 km from the road from San Carlos de Bariloche to Chenqueniyeu, to the west of the Province of Rio Negro, from the middle levels of the Yirihuau
Formation (Upper Oligocene-Lower Miocene), by Cione & Torno (1988)]. Holotype: División de Paleontología
Vertebrados del Museo de La Plata 21-871, partial skeletal impression.
† TREWAVASIA White & Moy Thomas, 1941
† Xenopholis Davis, 1887: 548. Type species: † Xenopholis carinatus Davis, 1887, by monotypy. Gender: Feminine.
Preoccupied by Xenopholis Peters, 1869, in Reptiles; replaced by † Trewavasia White & Moy Thomas, 1941,and †Xenopholoides Fowler, 1958.
†Trewavasia White & Moy Thomas, 1941: 400. Type species: † Xenopholis carinatus Davis, 1887. Gender: Feminine. Type by being a replacement name. Replacement for † Xenopholis Davis, 1887; preoccupied by Xenopholis Peters, 1869, in Reptiles. Gender: Feminine.
† Xenopholoides Fowler, 1958: 13. Type species: † Xenopholis carinatus Davis, 1887. Gender: Feminine. Type by
being a replacement name. Replacement for † Xenopholis Davis, 1887; preoccupied by Xenopholis Peters,
1869, in Reptiles.
Remarks: Described by Davis (1887) as probably showing a “closer connection with siluroids than with any othergroup” but clearly not a siluriform fish. Most likely an Acanthopterygian.
† Trewavasia carinata (Davis, 1887)† Xenopholis carinatus Davis, 1887: 549, pl. 29, fig. 4. Type locality: Mt. Lebanon, Syria, Hard Chalk.

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Appendix 1. Siluriform genera and species named from otoliths. Unless otherwise noted, all names were found in
Weiler’s (1968) catalogue of fish otoliths as belonging to catfishes
Generic names† Claibornichthys Frizzel & Dante, 1965: 697. Type species: † Claibornichthys troelli Frizzel & Dante, 1965. Type byoriginal designation. Gender: Masculine.
Remarks: † Otolithus (Sciaenidarum) decipiens Koken, 1888, tentatively included as a second species of this genus
by Frizzel & Dante (1965).
† Vorhisia Frizzell, 1965: 179. Type species: † Vorhisia vulpes Frizzell, 1965. Type by original designation. Gender:
Feminine.
Remarks: The family group name † Vorhisiidae was proposed in the same publication.
Species names(Names listed by date of publication)† Otolithus (incertae sedis) crassus Koken, 1884: 559, pl. 12, fig. 13. Type locality: Heaton Hill, Isle of Wight. Holotype: at Roy. Mus. Nat. Hist. Berlin (otolith), but apparently lost (Stinton, 1977: 71).
Remarks: Placed in Arius by Weiler (1968). Treated as valid as † Tachysurus crassus by Stinton (1977: 71).
† Otolithus (Sciaenidarum) decipiens Koken, 1888: 285, pl. 19 (figs. 5%6). Type locality: Calyborne-Schicten. Syntypes(2): Whereabouts unknown..
Remarks: Not listed among siluriform otolith based names in Weiler (1968), but treated a possible second species of† Claibornichthys (as † Otolithus (Arius) decipiens) in Frizzel & Dante (1965).
† Arius baroni Newton, 1889: 207, pl. 21, fig. 7. Type locality: Ankoala, Madagascar; Eocene. Holotype: at BMNH, inthe Baron collection (1 of 8 otoliths). Name available from figure caption, thus illustrated specimen is the holotype.
† Raja similis Woodward, 1889a: 86, pl. 4, figs. 4–5. Type locality: Hampshire, England; Eocene. (Otoliths).
Remarks: Name corrected to † Arius similis on plate caption. Treated as a synonym of † Tachysurus crassus (Koken)
by Stinton (1977: 71).
† Otolithus (Arius) danicus Koken, 1891: 81, fig. 1. Type locality: Copenhagen; lower Eocene. Based on otolith identified as † Otolithus cf. crassus in Koken (1885: 116, pl. 5, fig. 29).
† Otolithus (Arius) germanicus Koken, 1891: 81, pl. 1, fig. 3, and pl. 6, fig. 8. Type locality: Lattorf, Westeregeln, andOsterwedingen, Lower Oligocene; Sollingern, Weinheim, Waldboeckelheim, Middle Oligocene. (otoliths).
† Otolithus (Arius) vanigonis Koken, 1891: 81, pl. 6, fig. 4. Type locality: Mitteloligöcan, Waldböeckelheim.
† Otolithus (Arius) Lerichei Priem, 1906: 277, figs. 46–47. Type locality: problement du Thanétien ou Yprésien ? desenvirons de Reims, éocènes du bassin Parisien. Holotype (partial Lapillus): Collection Bourdot.
† Otolithus (Siluridarum ?) incertus Priem, 1906: 277, figs. 48–49. Type locality: problement d’Hérouval (Yprésien
supérieur), éocènes du bassin Parisien. Holotype: ? MNHN.
† Otolithus (Arius ?) moravicus Schubert, 1908: 106, abb. 3. Type locality: Marnes de Pausram (Lower Ologocene).
Holotype: GBW 1908/01/4.
Remarks: Nolf (1981: 136) listed this name as a rejected species.
† Otolithus (Arius) angelicus Bassoli, 1909: 41, fig. 1. Type locality: Headon Member, Solent formation, Brockenhurst,
Hampshire, Britain. Holotype: (Lapillus).
Remarks: Treated as valid as † Galeichthys angelicus by Stinton (1977: 73).
† Otolithus (Arius) crassus bartonensis Shepherd, 1916: 183, fig. 155 (5). Type locality: Ober- Eozän, England. Originally † Otolithus (Arius) crassus var. bartonensis.
Remarks: Treated as a synonym of † Tachysurus crassus (Koken) by Stinton (1977: 71).
† Otolithus (Arius) danicus bartonensis Shepherd, 1916: 180, fig. 157 (11). Type locality: Upper Eocene, England. Originally † Otolithus (Arius) danicus var. bartonensis.
Remarks: Treated as a synonym of † Tachysurus planus (Frost), as well † Tachysurus crassus (Koken) as by Stinton

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(1977: 70).
† Otolithus (Arius ?) parvus Schubert, 1916: 287, pl. 7 (fig. 24). Type locality: Barton Formation (Upper Eocene) at Barton Cliff, England. Syntypes: GBW 1916/01/25, 25a.
Remarks: Treated as a synonym of † Tachysurus crassus (Koken) by Stinton (1977: 71). Nolf (1981: 136) listed thisname as a rejected species.
† Otolithus (Arius) newtoni Schubert, 1916: 286, pl. 7 (fig. 3). Type locality: Barton Formation (Upper Eocene) at Barton
Cliff, England. Syntypes: GBW 1916/01/3 (or MGB 0427, see Stinton, 1977: 72).
Remarks: Treated as a synonym of † Tachysurus crassus (Koken) by Stinton (1977: 71). Nolf (1981: 136) treatedthis name as a synonym of † Arius crassus (Koken).
† Otolithus (Arius) tenuis Frost, 1925: 28, pl. 2 (fig. 37). Type locality: Hoofdsossiel-zone, Al. Geureugah (Nisam),Sumatra; upper Tertiary (Neogene). Holotype: (? Lapillus).
Remarks: Name spelled † Otolithus (Arius) tenius on plate, which is treated here as an invalid name.
† Otolithus (Arius) africanus Frost, 1926: 84, pl. 18, figs. 7–8. Type locality: Ameki, Nigeria; Eocene. Syntypes (5):(Lapillus).
† Otolithus (Arius) amekiensis Frost, 1926: 84, pl. 18, fig. 10. Type locality: Ameki, Nigeria; Eocene. Holotype: (Lapillus).
† Otolithus (Arius) angulatus Frost, 1926: 84, pl. 18, fig. 9. Type locality: Ameki, Nigeria; Eocene. Syntypes (3): (Lapillus).
† Otolithus (Arius ?) glaber Voigt, 1926: 177, pl. 2, figs. 23–24. Type locality: Bavaria, Cretaceous.
† Otolithus (Arius) jaekeli Richter 1928: 138, pl. 1, fig. 3. Type locality: Pomerania, Cretaceous.
† Otolithus (Arius) aequus Frost, 1934: 504, pl. 14 (fig. 15). Type locality: Barton Formation, Barton, Hampshire; Upper
Eocene. Holotype: BMNH P 22732 (Lapillus).
Remarks: Treated as valid as † Galeichthys aequus by Stinton (1977: 71).
† Otolithus (Arius) planus Frost, 1934: 504, pl. 14 (fig. 14). Type locality: Barton Formation, Barton, Hampshire; Upper
Eocene. Holotype: BMNH P 22730 (otolith).
Remarks: Treated as valid as † Tachysurus planus by Stinton (1977: 70).
† Arius rutschi Casier, 1958: 22, pl. 2 (figs. 12, 15). Type locality: l’Ile de la Barbade; formation de Scotland superieure.
Holotype: TLLS 80. (Lapillus).
† Arius cavatus Stinton, 1962: 82, pl. 19 (fig. 6). Type locality: Borneo, Lower Pliocene. Holotype: (Lapillus).
† Tachysurus oblongus Stinton, 1962: 81, pl. 19 (fig. 3). Type locality: Borneo, Miocene–Pliocene. Type(s): Whereabouts unknown.
† Netuma radiata Stinton, 1962: 81, pl. 1 (fig. 17). Type locality: Borneo, Pliocene.
† Netuma regularis Stinton, 1962: 82, pl. 19 (fig. 5). Type locality: Borneo, Pliocene.
† Otolithus (Arius) rotundatus Roedel, 1930: 52, pl. 1, fig. 17. Type locality: Köthen und Frqankfurt [Germany], Eocene.
Holotype: (Lapillus).
† Vorhisia vulpes Frizzell, 1965: 180, figs, 2, 3. Type locality: Upper Cretaceous (Maestrichtian): along left bank ofGrand River; Sec. 19, T20N, R20W; Corson County, South Dakota. Holotype: Frizzell personal collection, no number (otolith: left lapillus).
Distribution: Fox Hill Formation: Irish Creek lithofacies, Bullhead lithofacies, and Colgate lithofacies, South
Dakota, USA; Upper Cretaceous (Frizzell & Koenig, 1973).
† Claibornichthys troelli Frizzel & Dante, 1965: 697, pl. 86 (figs. 1, 2, 8, 17, 18, 19). Type locality: Stone City Beds:
Brazos River, west of Bryan Texas. Holotype: USNM 23371 (otolith).
† Diplomystes rudis Stinton, 1966: 423, pl. 66, fig. 12. Type locality: London Clay, England.
Distribution: London Clay, England, Lower Eocene (Weiler, 1968).

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Appendix 2. Institutional codes
AFY
AI
AMG
AMNH
AMS
ANSP
ASIZB
BCUE
BKNU
BMNH
BSMP

Personal collection of Agustín Fernández-Yépez; portions now at SCN and MBUCV, Venezuela.
Asociación Ictiológica, La Plata, Argentina.
Albany Museum, Grahamstown, South Africa.
American Museum of Natural History, New York, U.S.A.
Australian Museum, Sydney, N. S. W., Australia.
Academy of Natural Sciences, Philadelphia, Pennsylvania, U.S.A.
Academia Sinica Institute of Zoology, Beijing, China.
Department of Biology, Ch’ongju University of Education, Korea.
Kunsan National University, Department of Biology, Korea.
Natural History Museum, London. Formerly British Museum (Natural History), U.K.
Department of Agriculture, Bureau of Science, Manila, Philippines. Evidently entire collection destroyed
during World War II.
CAS
California Academy of Sciences, San Francisco, California, U.S.A.
CM
Carnegie Museum, Pittsburgh, Pennsylvania, U.S.A. Now at FMNH.
CPUC
Colecciones Paleontológicas, Departamento Ciencias de la Tierra, Universidad de Concepción, Concepción, Chile.
CSIRO
Commonwealth Science & Industrial Research Organization, Division of Marine Research, Hobart, Tasmania, Australia.
CU
Cornell University Museum of Vertebrates, Ithaca, New York, U.S.A.
DGM
Paleontological collection of the Setor de Paleontologia, Departamento Nacional de Produço Mineral, Universisade do Estado do Rio de Janeiro, Brazil.
DVZUT
Department of Vertebrate Zoology, University of Tông-Hop, Hanoi, Vietnam.
DZSASP
Departamento de Zoologia, Secretaria da Agricultura, São Paulo, Brazil. Formerly Museu Paulista.
ECO-SCEl Colegio de la Frontera Sur, San Cristóbal, Chiapas, Mexico.
EEBP
Estaçao Experimental de Biologia e Piscicultura de Pirassununga, Brazil.
FAKU
Kyoto University, Department of Fisheries, Faculty of Agriculture, Japan.
FFSUC
Faculty of Forestry Sciences, Universidad de Chile, Ichthyological Collection, Santiago, Chile.
F/GUZ
Department of Zoology, Gauhati University, Guwahati, Assam, India.
FML
Instituto Fundacion Miguel Lillo, Tucuman, Argentina.
FMNH
Field Museum of Natural History, Chicago, Illinois, U.S.A.
FRLM
Faculty of Fisheries, Mie University, Fisheries Research Laboratory, Mie-ken, Japan.
GBW
Collection du Gelogische Bundesanstald, Vienna, Austria.
GCM
Government College, Department of Zoology, Lahore, Pakistan.
GSJ
Geological Society of Japan.
ICNMHN Instituto de Ciencias Naturales, Museo de Historia Natural, Universidad Nacional de Colombia, Bogotá,
Colombia. .
IHASWInstitute Hydrobiology Academy Sinica, Wuhan, China.
ILPLA
Museo de La Plata, Instituto de Limnologia, La Plata, Argentina.
IMCN
Natural Sciences Museum Federico Carlos Lehmann V.- INCIVA Cali, Colombia
INHS
Illinois Natural History Survey, Champaigne, Illinois, U.S.A.
INPA
Instituto Nacional de Pesquisas da Amazonia, Manaus, Amazonas, Brazil.
INVEMAR Instituto de Investigaciones Marinas de Punta de Betin, Santa Marta, Colombia.
IRSNB
Institut Royal des Sciences Natureles de Belgique, Brussels, Belgium.
IU
Indiana University, Bloomington, Indiana, U.S.A. Now distributed among several institutions, primarily
CAS, UMMZ and USNM; many IU types remain unaccounted for.
IUQ
Laboratorio de Ictiología, Departamento de Biología, Universidad del Quindío, Armenia, Colombia.
IZUA
Universidad Austral de Chile, Instituto de Zoologia, Valdivia, Chile.
JFBM
Bell Museum Fish Collection at the University of Minnesota.

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JNU
KIZ
KU
KUMF
LACM
LIVCM
LUVP
MACN
MB
MBLUZ
MBUCV
MCN.USB
MCNG
MCP
MCZ
MD
MEPN
MHNC
MHNG
MHNL
MHNLR
MHNLS
MHNM
MHNN
MHV
MIKU
MLP
MNHN
MNHNP
MNKP
MNRJ
MNSB
MPEG
MRAC
MRCN
MSINR
MSNG
MSNM
MSUM
MTD
MUMF
MUSM
MZB
MZUF
MZS
MZUSP
MZUT
NIFI
NMBA

Department of Biology, Ji Nan University, China.
Kunming Institute of Zoology, Chinese Academy of Sciences, Kunming, Yunnan, China.
University of Kansas, Museum of Natural History, Lawrence, Kansas, U.S.A.
Kasetsart University Museum of Fisheries, Bangkok, Thailand.
Los Angeles County Museum of Natural History, Los Angeles, California, U.S.A.
World Museum Liverpool, U.K.
Vertebrate Palaeontology Laboratory, Geology Department, Luchnow Uiversity, Luchnow, India.
Museo Argentino de Ciencias Naturales, Bernardino Rivadavia, Capital Federal, Argentina.
Universidade de Lisboa, Museu Bocage, Lisboa, Portugal.
Museo de Biología de la Universidad del Zulia, Venezuela.
Universidad Central de Venezuela, Museo de Biologia, Caracas, Venezuela.
Museo de Ciencias Naturales, Universidad Simón Bolívar, Caracas, Venezuela.
Museo de Ciencias Naturales, Guanare, Venezuela.
Pontifícia Universidade Católica do Rio Grande do Sul, Museu de Ciências, Rio Grande do Sul, Porto
Alegre, Brazil.
Museum of Comparative Zoology, Harvard University, Cambridge, Massachusetts, U.S.A.
Museo do Dundo, Dundo, Angola.
Escuela Polytecnica Nacional, Quito, Ecuador.
Museo de Historia Naturale de Cochabamba, Bolivia.
Museum d’Histoire Naturelle, Geneve, Switzerland.
Musée d’Hisoire Naturelle du Luxembourg.
Museum d’Histoire Naturelle, La Rochelle, France.
Museo de Historia Natural de La Salle, Caracas, Venezuela.
Museo de Historia Natural de Montevideo, Montevideo, Uruguay.
Museum d’Histoire Naturelle, Neuchatel, Switzerland.
Musée de Haute Volta, Ouagadougou, Upper Volta.
Marine Biological Institute, Kyoto University, Japan. Specimens now at FAKU.
Museo de La Plata, La Plata, Argentina.
Muséum National d’Histoire Naturelle, Paris, France.
Museo Nacional de Historia Natural de Paraguay.
Museo de Historia Natural Noel Kempff Mercado, Santa Cruz, Bolivia.
Universidade Federal do Rio de Janeiro, Museu Nacional, Rio de Janeiro, Brazil.
Museum of Natural Sciences, Budapest, Hungary.
Museu Paraenese ‘Emilio Goeldi’, Para, Brazil.
Musée Royal de l’Afrique Centrale, Tervuren, Belgium.
Museu Rio-Grandense de Ciências Naturais, Brazil.
Museum Sichuan Institute of Natural Resources, China.
Museo Civico di Storia Naturale di Genova ‘Giacomo Doria’, Genova, Italy.
Museo Civico di Storia Naturale, Milano, Italy.
Michigan State University, University Museum, East Lansing, Michigan, U.S.A.
Staatliches Museum für Tierkunde, Dresden, Germany.
Department of Life Sciences, Manipur University Museum of Fishes, Canchipur, Manipur, India.
Museo de Historia Natural de la Universidad Nacional Mayor de San Marcos, Lima, Peru.
Museum Zoologicum Bogoriense, Bogor, Indonesia.
Universita di Firenze, Museo Zoologico de la Specola, Firenze, Italy.
Universite de Strasbourg, Musée de Zoologie, Strasbourg, France.
Universidade de São Paulo, Museu de Zoologia, São Paulo, Brazil.
Universita di Torino, Museo Zoologico, Torino, Italy.
National Inland Fisheries Institute, Fish Taxonomy Division, Bangkok, Thailand.
Naturhistorisches Museum Basel, Basel, Switzerland.

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NMBE
NMC
NMK
NMSL
NMSZ
NMV
NMW
NRM
NSMT
NTM
OSUS
PEM
QM
QVMS
RMNH
ROM
SAIAB

Naturhistorisches Museum Bern, Bern, Switzerland.
National Museums of Canada, Ottawa, Canada.
National Museum, Nairobi, Kenya.
National Museum, Sir Marcus Fernando Mawatha, Colombo, Sri Lanka.
National Museums of Scotland, Edinburgh, Scotland.
National Museum of Victoria, Melbourne, Victoria, Australia.
Naturhistorisches Museum, Vienna, Austria.
Swedish Museum of Natural History (Naturhistoriska Riksmuseet), Stockholm, Sweden.
National Science Museum, Department of Zoology, Tokyo, Japan.
Northern Territory Museum of Arts & Sciences, Darwin, Northern Territory, Australia.
Oklahoma State University, Department of Zoology, Stillwater, Oklahoma, U.S.A.
Port Elizabeth Museum, Port Elizabeth, South Africa.
Queensland Museum, Brisbane, Queensland, Australia.
Queen Victoria Museum, Harare, Zimbabwe.
Rijksmuseum van Natuurlijke Historie, Leiden, Netherlands.
Royal Ontario Museum, Toronto, Ontario, Canada.
South African Institute for Aquatic Biodiversity; formerly J. L. B. Smith Institute of Ichthyology, Grahamstown, South Africa (RUSI).
SAM
South African Museum, Cape Town, South Africa.
SAMASouth Australian Museum, Adelaide, South Australia, Australia.
SBM
Sabah Museum, Sabah, East Malaysia.
SCNU
South China Normal University, Guangzhou, China.
SIUC
Southern Illinois University (Carbondale), University Museum, Carbondale, Illinois, U.S.A.
SMF
Natur-Museum und Forschungs-Institut Senckenberg, Frankfurt-am-Main, Germany.
SMK
Sarawak Museum, Kuching, Sarawak, Malaysia.
SMNS
Staatliches Museum fur Naturkunde in Stuttgart, Ludwigsburg, Germany.
SNMB
Slovak National Museum, Bratislava, Czechoslovakia.
SPNRI
Sichuan Province, Natural Resources Institute, China.
SRS/ZSI
Southern Regional Station, Zoological Survey of India, Madras, India.
STRI
Smithsonian Tropical Research Institute, Panama.
SU
Stanford University, Palo Alto, California, U.S.A. Specimens on long term loan to CAS.
TM
Transvaal Museum, Pretoria, South Africa.
TU
Tulane University, Department of Zoology, New Orleans, Louisiana, U.S.A.
UAB
Universidad Autonoma de Barcelona, Departamento de Biología, Barcelona, Spain.
UBJTL
Universidad Bogotá Jorge Tadeo Lozano, Colombia.
UFRJ
Universidade Federal do Rio de Janeiro, Brazil.
UG/CSBD University of Guyana, Center for the study of Biological Diversity, Georgetown, Guyana.
UMMP
University of Michigan Museum of Paleontology, Ann Arbor, Michigan, U.S.A.
UMMZ
University of Michigan Museum of Zoology, Ann Arbor, Michigan, U.S.A.
UNAM
Universidad Nacional Autonoma de Mexico, Departmento de Zoologia, Instituto de Biologia, Mexico, DF,
Mexico.
UO
University of Oregon, Corvallis, Oregon, U.S.A.
USNM
National Museum of Natural History, Washington, D.C., U.S.A. Formerly United States National Museum.
UW
University of Washington, College of Fisheries, Seattle, Washington, U.S.A.
WAM
Western Australian Museum, Perth, Western Australia, Australia.
ZFMK
Zoologisches Forschunginstitut und Museum Alexander Koenig, Bonn, Germany.
ZIN
Zoological Institute, Academy of Sciences, St. Petersburg, Russia.
ZISP
Zoological Institute, Academy of Sciences, Leningrad, Russia.
ZMA
Universiteit van Amsterdam, Zoologisch Museum, Amsterdam, The Netherlands.
ZMB
Universitat Humboldt, Museum fur Naturkunde, Berlin, Germany.

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ZMH
ZMMU
ZMUB
ZMUC
ZMUL
ZMUO
ZMUT
ZMUU
ZRC
ZSI
ZSIC
ZSI/NRSZSI/SRS
ZVC-P
ZSM
ZX

Zoological Museum Fan Memorial Institute of Biology, Tsing Hua University, Peiping, China.
Universitat Hamburg, Zoologisches Institut und Museum, Hamburg, Germany.
Zoological Museum, Moscow University, Russia.
Universitetet i Bergen, Zoologisk Museum, Bergen, Norway.
Kobenhavns Universitet Zoologisk Museum (Zoological Museum, University of Copenhagen), Copenhagen, Denmark.
Universitetets Lund, Zoologiska Museet, Lund, Sweden.
Universitetets I Oslo, Zoologisk Museum, Oslo, Norway.
University of Tokyo, Department of Zoology, University Museum, Tokyo, Japan.
Zoologiska Museet, Uppsala Universitet, Uppsala, Sweden.
Zoological Reference Collection, National University of Singapore, Singapore.
Zoological Survey of India, Kolkata [Calcutta], India.
Zoological Survey of India, Chennai, India.
Northern Regional Station, Zoological Survey of India, Dehradun, India.
Southern Regional Station, Zoological Survey of India, Madras, India.
Facultad de Ciencias, Universidad de la República, Montevideo, República Oriental del Uruguay
Zoologische Museum Staatssammlung, München, Germany.
Zhuhai City Fisheries Science Research Institute, Guandong Province, China.

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Appendix 3. Taxonomic publications on Siluriformes issued in 2006
The following publications were issued in 2006, after the cutoff date for this Checklist. The publications are listed inalphabetical order of authorship, with an abbreviated summary of the taxonomic and nomenclatural actions taken, tomake the checklist as up to date as possible. New names and other names that were subject to nomenclatural action havebeen noted and are included in the index. However, new names and any other proposed taxonomic changes were notincorporated into the statistics provided in Table 1.
Alencar, A.R. & Costa, W.J.E.M. (2006) Trichomycterus pauciradiatus, a new catfish species from the upper rio Paranábasin, southeastern Brazil (Siluriformes: Trichomycteridae). Zootaxa, 1269, 43–49.
Trichomycterus pauciradiatus Alencar & Wilson, 2006: 44, fig. 1. Type locality: Brazil: Estado de Minas Gerais: Municipio de Carrancas, córrego Debaixo da Serra, stream tributary of córego Água Limba, rio Paraná basin, 21°26'40"S,
44°36'09"W. Holotype: UFRJ 5831.
Distribution: Upper Paraná River basin, southeastern Brazil.
Almirón, A., Azpelicueta, M. de las M., Casciotta, J. & Litz, T. (2006) A new species of Hisonotus (Siluriformes, Loricariidae, Otothyrini) from the Republica Oriental del Uruguay. Revue suisse de Zoologie, Annales de la Société
zoologique suisse et du Muséum d'Histoire naturelle de Genève, 113, 87–94.
Hisonotus charrua Almirón Azpelicueta, Casciotta & Litz, 2006: 88, figs. 1–3. Type locality: República Oriental del
Uruguay, Departamento Tacuarembó, río Uruguay basin, Cañada de Los Peña (31°39.09'S, 56°12.32'W). Holotype:
ZVC-P 5639.
Distribution: Uruguay River and La Plata River mouth, Uruguay.
Betancur-R., R. & Acero P., A. (2006) A new species of Notarius (Siluriformes: Ariidae) from the Colombian Pacific.
Zootaxa, 1249, 47–59.
Notarius armbrusteri Betancur-R. & Acero P., 2006: 50, figs. 2–3. Type locality: fish market of Buenaventura, Valle delCauca, Colombia. Holotype: INVEMAR-PEC 6677.
Distribution: Eastern Pacific coast, at Valle del Cauca, Colombia.
Key: Species of Notarius of Eastern Pacific on p. 58.
Casciotta, J., Azpelicueta, M. de las M., Almirón, A. & Litz, T. (2006) Hisonotus candombe, a new species from the río
Uruguay basin in the República Oriental del Uruguay (Siluriformes, Loricariidae, Otothyrini). Spixiana, 26 (2),
147–152.
Hisonotus candombe Casciotta, Azpelicueta, Almirón & Litz, 2006: 147, figs. 1–3. Type locality: República Oriental del
Uruguay, Departamento Salto, río Uruguay basin, arroyo Palomas (31°04'43"S – 57°37'26"W. Holotype: ZVC-P
5595.
Distribution: lower Uruguay River basin, Uruguay.
Covain, R., Le Bail, P.Y., Sagnes, P. & Fisch-Muller, S. (2006) Species of the genus Harttia (Siluriformes: Loricariidae)
in French Guiana: Morphology, taxonomy and distribution. Cybium, 30, 3–18.
Day, J. J. & Wilkinson, M. (2006) On the origin of the Synodontis catfish species flock from Lake Tanganyika. Biology
Letters, doi:10.1098/rsbl.2006.0532, 1–5.
Diogo, R. & Bills, R. (2006) Osteology and myology of the cephalic region and pectoral girdle of the South African catfish Austroglanis gilli, with comments on the autapomorphies and phylogenetic relationships of the Austroglanididae (Teleostei: Siluriformes). Animal Biology, 56, 39–62.
Diogo, R., Chardon, M. & Vandewalle, P. (2006) Osteology and myology of the cephalic region and pectoral girdle of the
Cetopsis coecutiens Spix & Agassiz, 1829, comparison with other cetopsids, and comments on the synapomorphiesand phylogenetic relationships of the Cetopsidae (Teleostei: Siluriformes). Belgian Journal of Zoology, 136, 3–13.
Diogo, R., Chardon, M. & Vandewalle, P. (2006) Osteology and myology of the cephalic region and pectoral girdle of the
Nematogenys inermis Guichenot, 1848, with comments on the autapomorphies and phylogenetic relationships of the
Nematogenyidae (Teleostei: Siluriformes). Belgian Journal of Zoology, 136, 15–24.
Do Nascimiento, C. & Provenzano, F. (2006) The genus Henonemus (Siluriformes: Trichomycteridae) with a descriptionof a new species from Venezuela. Copeia, 2006, 198–205.

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Henonemus triacanthopomus Do Nascimiento & Provenzano, 2006: 199, fig. 1. Type locality: Venezuela, Estado Delta
Amacuro, beach at Caño Macareo, 8°49'16"N, 62°00'00"W. Holotype: MBUCV-V 29526
Distribution: Apure and Arauca Rivers and in Caño Macareo in the Orinoco Delta, Venezuela.
Egge, J.J.D. & Simons, A.M. (2006) The challenge of truly cryptic diversity: diagnosis and description of a new madtom
catfish (Ictaluridae: Noturus). Zoologica Scripta, 35, 581–595.
Noturus maydeni Egge, 2006: 588, fig. 7. Type locality: Strawberry River at Simstown Public Access, 4 mi N of Evening
Shade, 1.7 mi off U.S. Hwy 167 on Simstown Road (036°05'52"N, 091°36'27"W), Sharp Co., AR [USA]. Holotype:
JFBM 39465.
Distribution: Black and St. Francis river basins, Mississippi River drainage, Arkansas and Missouri, USA (Egge &Simons, 2006).
Remarks: Not distinguishable morphologically from Noturus albater; diagnosis based on karyotypes, allozyme variation, and DNA sequences.
Fernández, L. & Osinaga, K. (2006) A new Trichomycterus (Siluriformes: Trichomycteridae) from Aguarague National
Park of the Bolivian preandean region, with comments on the relationships within the genus. Environmental Biologyof Fishes, 75, 385–393.
Trichomycterus aguarague Fernández & Osinaga, 2006: 386, fig. 1. Type locality: Bolivia, Tarija, Province Gran Chaco,
National Park Aguarague, Quebrada Timboycito, a Río Caiguami tributary, elevation 700–800 m, 21°30'S, 63°60'W.
Holotype: MNKP 4012.
Distribution: Caiguami River tributaries, Pilcomayo River basin, Paraná River system, Bolivia.
Friel, J.P. & Vigliotta, T.R., (2006) Synodontis acanthoperca, a new species from the Ogôoué River system, Gabon withcomments on spiny ornamentation and sexual dimorphism in mochokid catfishes (Siluriformes: Mochokidae). Zootaxa, 1125, 45–56.
Synodontis acanthoperca Friel & Vigliotta, 2006: 47, figs. 1–3. Type locality: Gabon, Haut-Ogôoué Province, OgôouéRiver at and below the Rapids of Massoukou (Masuku), 1°39'30"S 13°32'14"E. Holotype: CU 89005.
Distribution: Louétsi and Ogôoué Rivers, Ogôoué River basin, Gabon.
Jayaram, K.C. (2006) Catfishes of India, Narendra Publishing House, Delhi, xxii +323 p., 11 pl.
Pterocryptis barakensis Vishwanath & Sharma, in Jayaram, 2006: 99, pl. 5, fig. 2. Type locality: India, Manipur, Tamenglong District, Barak River at Vanchengphai, village. Holotype: MUMF 4018.
Distribution: Barak River, Brahmaputra River basin, India.
Horabagrinae new subfamily [Schilbidae], Jayaram, 2006: 141. Type genus: Horabagrus Jayaram, 1955.
Horaglanidinae new subfamily [Clariidae], Jayaram, 2006: 309. Type genus: Horaglanis Menon, 1950.
Janssen, G., Devaere, S. Weekers, P. & Adriaens, D. (2006) Phylogenetic and biogeographical analysis of African air–breathing catfish (Siluriformes: Clariidae): inferred from ribosomal genes and spacers sequences with emphasize on
anguilliformity. Molecular Phylogeny and Evolution 38: 65–78.
Jerep, F.C., Shibatta, O.A., Pereira, E.H.L. & Oyakawa, O.T. (2006) Two new species of Isbrueckerichthys Derijst, 1996
(Siluriformes: Loricariidae) from the rio Paranapanema basin, Brazil. Zootaxa, 1372, 53-68.
Isbrueckerichthys calvus Jerep, Shibatta, Pereira & Oyakawa, 2006: 60, fig. 2. Type locality: Brazil; Paraná State; Apucarana; rio Tibagi basin; córrego Juruba; 23°34’44.6”S/51°22’12.6”W. Holotype: MZUEL 3714. Distribution: Tributary streams of the Taquara River, Tibagi River basin, Paraná State, Brazil.
Isbrueckerichthys saxicola Jerep, Shibatta, Pereira & Oyakawa, 2006: 55, fig. 1. Type locality: Brazil; Paraná State;
Londrina; rio Tibagi basin; ribeirão Jacutinga; 23°14´30S/51°13´05”W. Holotype. MZUEL 3716. Distribution:
Ribeirão Jacutinga headwaters, Tibagi River basin, Paraná State, Brazil.
Koblmüller, S., Sturmbauer C., Verheyen, E., Meyer, A. & Salzburger, W. (2006) Mitochondrial phylogeny and phylogeography of East African squeaker catfishes (Siluriformes: Synodontis). BMC Evolutionary Biology, 2006, 6 (49)(http://www.biomedcentral.com/1471-2148/6/49).
Mori, H. & Shibatta, O.A. (2006) A new species of Microglanis Eigenmann, 1912 (Siluriformes, Pseudopimelodidae)
from rio São Francisco basin, Brazil. Zootaxa, 1302, 31–42.
Microglanis leptostriatus Mori & Shibatta, 2006: 33, fig. 1. Type locality: rio Verde Grande, 16º 39'S/46º11'57.8"W,Montes Claros, State of Minas Gerais, Brazil. Holotype: MZUSP 85985.
Distribution: São Francisco River basin, Brazil.

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Murdy, E.O. & Ferraris, C.J., Jr. (2006) A revision of the marine catfish genus Euristhmus (Teleostei: Siluriformes: Plotosidae). The Beagle, Records of the Museums and Art Galleries of the Northern Territory, 22, 77-90.
Euristhmus microphthalmus Murdy & Ferraris, 2006: 84, figs. 3, 6. Type locality: AUSTRALIA. Northern Territory.
Woods Inlet, Darwin Harbour, 12°30'S, 130°45'E. Holotype: NTM S. 11242-001.
Distribution: Northern Territory, Australia and Irian Jaya, Indonesia, in nearshore habitats over soft bottoms.
Euristhmus sandrae Murdy & Ferraris, 2006: 88, figs 3, 9. Type locality: AUSTRALIA. Western Australia. ExmouthGulf, 21°42'S, 114°48'E, 9 m. Holotype: WAM P.32730.001.
Distribution: Western Australia from south of Rowley Shoals and Exmouth Gulf, in waters as deep as 80m over soft
bottoms.
Cnidoglanis lepturus Günther, 1864, Lectotype: BMNH 1864.1.17.33, designated on p. 82.
Key: species of Euristhmus on p. 80.
Near, T. J. & Hardman, M. (2006) Phylogenetic relationships of Noturus stanauli and N. crypticus (Siluriformes: Ictaluridae), two imperiled freshwater fish species from the southeastern United States. Copeia, 2006, 378–383.
Ng, H.H. (2006) A Phylogenetic Analysis of the Asian Catfish Family Sisoridae (Teleostei: Siluriformes), and the Evolution of Epidermal Characters in the Group. Unpublished Ph.D. dissertation, University of Michigan, Ann Arbor.
Ng, H.H. (2006) Erethistoides infuscatus, a new species of catfish (Teleostei: Erethistidae) from South Asia. Ichthyological Explorations of Freshwaters, 17, 283–287.
Erethistoides infuscatus Ng, 2006: 284, fig. 1. Type locality: India: Meghalaya, East Khasi Hills, Umsing River. Holotype: UMMZ 245695.
Distribution: Brahmaputra and Meghna River drainages, northeast India and Bangladesh.
Ng, H.H. (2006) The identity of Batasio tengana (Hamilton, 1822), with the description of two new species of Batasio
from north-eastern India (Teleostei: Bagridae). Journal of Fish Biology, 68 (suppl. A), 101–118.
Batasio tengana Hamilton, 1822: Neotype designated by Ng, 2006: 103, fig. 1. Type locality: India: West Bengal: Tista
River at Tista barrage; 26°45'10"N; 88°34'11"E. Neotype: UMMZ 244796.
Distribution: Ganges and Brahmaputra River basins.
Batasio fasciolatus Ng, 2006: 107, fig. 4. Type locality: India: West Bengal, market at Malbazar: 26°32'30"N;
88°44'17"E. Holotype: UMMZ 244798.
Distribution: Tista River basin, Brahmaputra River system.
Batasio spilurus Ng, 2006: 110, fig. 5. Type locality: India: Assam, Dibrugarh district, 27°29'N: 94°54'E. Holotype: ZRC
49133.
Distribution: Brahmaputra River basin (Ng, 2006).
Ng, H.H. (2006) Pseudecheneis suppaetula, a new species of glyptosternine catfish (Teleostei: Sisoridae) from India.
Zootaxa, 1267, 59–68.
Pseudecheneis suppaetula Ng, 2006: 60, fig. 1. Type locality: India: Himachal Pradesh, Ganges River drainage, upper
reaches of Giri River, in Chhaila area (in the vicinity of Kotkhai), 31°6'15"N 77°25'56"E. Holotype: NRM 36977.
Distribution: Ganges River basin, Himachal Pradesh, India.
Ng, H.H. (2006) Akysis longifilis, a new species of catfish (Teleostei: Akysidae) from Myanmar. Zootaxa, 1150, 19–30.
Akysis longifilis Ng, 2006: 20, fig. 1. Type locality: Myanmar: Bago division, Pyu township, Pyu stream (tributary of Sittang River) ca. 229 km from Yangon, 18°29'N, 96°26'E. Holotype: UMMZ 246172.
Distribution: Sittang River basin, Myanmar.
Ng, H.H. (2006) The identity of Pseudecheneis sulcata (M’Clelland, 1842), with descriptions of two new species ofrheophilic catfish (Teleostei: Sisoridae) from Nepal and China. Zootaxa, 1254, 45–68.
Pseudecheneis eddsi Ng, 2006: 51, fig. 4. Type locality: Nepal: Tanahun, Khairenitar, Seti River (Ganges River drainage), 28°2'0.0"N 84°4'0.0"E. Holotype: KU 36872.
Distribution: Gandaki River basin, Nepal.
Pseudecheneis stenura Ng, 2006: 57, fig. 5. Type locality: China: Yunnan, Baoshan Prefecture, Longchuanjiang at Lianmengjie bridge (Irrawaddy River drainage). Holotype: KIZ 199811999.
Distribution: Irrawaddy River basin, Yunnan, China.
Ng, H.H. (2006) Pseudolaguvia ferula, a new species of sisoroid catfish (Teleostei: Erethistidae) from India. Zootaxa,
1229, 59–68.
Pseudolaguvia ferula Ng, 2006: 60, fig. 1. Type locality: India: West Bengal, Tista River at Tista Barrage, 26°45'10"N

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88°34'11"E. Holotype: UMMZ 245985.
Distribution: Tista River, West Bengal, India.
Ng, H.H. & Bailey, R.M. (2006) Chiloglanis productus, a new species of suckermouth catfish (Siluriformes: Mochokidae) from Zambia. Occasional Papers of the University of Michigan Museum of Zoology, 738, 1–13.
Chiloglanis productus Ng, & Bailey, 2006: 2, figs. 1–3. Type locality: Zambia: Lunzua stream, 11.3 km SSE of Mpulungu, about 1 km N of bridge on Mbala–Mpulungu road, 8°49'S 31°10'E. Holotype: UMMZ 199816.
Distribution: Lunzua River basin, Lake Tanganyika drainage, Zambia.
Ng, H.H. & Lim, K.K.P. (2006) Two new species of Leiocassis (Teleostei: Bagridae), riverine catfishes from northeast
Borneo. Ichthyological Exploration of Freshwaters, 17, 165–172.
Leiocassis collinus Ng & Lim, 2006: 166, fig. 1. Type locality: Borneo: Sabah: Danum Valley, Segama River drainage,
Sungai Palum Tambun, tributary of Sungai Segama, upstream of Danum Valley Field Center. Holotype: ZRC 46154.
Distribution: Kalabakan and Segama River basins, Sabah, Borneo.
Leiocassis tenebricus Ng & Lim, 2006: 166, fig. 1. Type locality: Borneo: Kalimantan Timur: Kayan River drainage,
Sungai Nah, tributary to Kayan River ca. 20 minutes upstream of confluence with Iwan River, 1°57'43.2"N
115°6'35.4"E at 550 m asl;. Holotype: MZB 10718.
Distribution: Kayan River basin, northeastern Borneo.
Ortega-Lara, A. & Lehmann A., P. (2006) Cruciglanis, a new genus of pseudopimelodid catfish (Ostariophysi: Siluriformes) with description of a new species from the Colombian Pacific coast. Neotropical Ichthyology, 4, 147–156.
Cruciglanis Ortega-Lara & Lehmann, 2006: 149. Type species: Cruciglanis pacifici Ortega-Lara & Lehmann, 2006, byoriginal designation. Gender: Masculine.
Cruciglanis pacifici Ortega-Lara & Lehmann, 2006: 150, fig. 1. Type locality: Colombia, Valle del Cauca Department,
near Buenaventura city, San Cipriano River where it crosses San Cipriano village, confluence of the La Sardina
stream, approximately 500 m before the confluence with Dagua River, Dagua River basin, 03º50.543'N
76º54.068'W, 84 m of altitude. Holotype: IMCN 2359.
Distribution: Dagua River basin and Anchicayá River basin, Pacific versant, Colombia.
Parisi, B.M., Lundberg, J.G. & DoNascimiento, C. (2006) Propimelodus caesius a new species of long-finned pimelodid
catfish (Teleostei: Siluriformes) from the Amazon Basin, South America. Proceedings of the Academy Of Natural
Sciences, Philadelphia, 155, 67–78.
Propimelodus caesius Parisi, Lundberg & DoNascimiento, 2006: 68, figs. 1–2. Type locality: Brazil, Pará State, Rio
Amazonas above Rio Trombetas, 20 km above Obidos, collected with 3 m bottom trawl in channel 4–7 m deep,
1°56'07.3"S, 55°41'18.5"W. Holotype: MZUSP 88582.
Distribution: Amazon River main stem and lower reaches of many tributary rivers, Brazil and Peru.
Peng, Z.-G., Ho, S.-W., Zhang, Y.-G. & He, S.-P. (2006) Uplift of the Tibetan plateau: Evidence from divergence times ofglyptosternoid catfishes. Molecular Phylogenetics and Evolution, 39, 568–572.
Provenzano, F. & Milani, N. (2006) Cordylancistrus nephelion (Siluriformes, Loricariidae), a new and endangered species of suckermouth armored catfish from the Tuy River, north-central Venezuela. Zootaxa, 1116, 29–41.
Cordylancistrus nephelion Provenzano & Milani, 2006: 31, figs. 1–2. Type locality: Venezuela, Caribbean Sea basin,
Tuy River system, Mesia River, tributary of the Guare River, near Village Corocito, approximately 10°12'N,
67°05'W. Holotype: MBUCV-V-21800.
Distribution: Tuy River basin, Venezuela (Provenzano & Milani, 2006).
Key: species of Cordylancistrus on p. 38.
Reis, R.E. & Borges, T.A.K. (2006) The South American catfish genus Entomocorus (Ostariophysi: Siluriformes:
Auchenipteridae), with the description of a new species from the Paraguay River basin. Copeia, 2006, 412–422.
Entomocorus radiosus Reis & Borges, 2006: 416, figs. 6–7. Type locality: Brazil, Mato Grosso, Rio Paraguay at neighborhood of Cáceres, approx. 16°03'S, 057°42'W. Holotype: MCP 35902.
Distribution: Paraguay River basin (Reis & Borges, 2006).
Key: species of Entomocorus p. 419.
Reis, R.E., Pereira, E.H.L. & Armbruster, J.A.W. (2006) Delturinae, a new loricariid catfish subfamily (Teleostei, Siluriformes), with revisions of Delturus and Hemipsilichthys. Zoological Journal of the Linnean Society, 147, 277–299.
Delturus brevis Reis & Pereira, 2006: 295, fig. 12. Type locality: Rio Araçuaí upstream of its mouth on Rio Jequitinhonha, Araçuaí, Minas Gerais, Brazil. Holotype: MZUSP 69858.
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Distribution: Jequitinhonha River basin, Minas Gerais, Brazil.
Delturinae new subfamily Armbruster, Reis & Pereira, 2006: 279. Type genus: Delturus Eigenmann & Eigenmann,
1889.
Key: subfamilies of Loricariidae, p. 279.
Key: Hemipsilichthys on p. 281.
Delturus parahybae Lectotype: MCZ 7726, designated on p. 293.
Delturus angulicauda Lectotype: NMW 44069, designated on p. 288.
Hemipsilichthys papillatus holotype illustrated on p. 285, fig. 6.
Hemipsilichthys nimius holotype illustrated on p. 287, fig. 8.
Retzer, M. E. (2006) A new species of Farlowella Eigenmann and Eigenmann (Siluriformes: Loricariidae), a stick catfish from Bolivia. Zootaxa, 1282, 59–68.
Farlowella altocorpus Retzer, 2006: 60, figs. 1, 2. Type locality: Bolivia: La Paz State: Río Beni Basin: Río Coroico,
Caranavi. Holotype: INHS 99773.
Distribution: Coroico River, Beni River basin, Bolivia.
Salcedo, N.J. (2006) New species of Chaetostoma (Siluriformes: Loricariidae) from central Peru. Copeia, 2006, 60–67.
Chaetostoma changae, Salcedo, 2006: 61, fig. 1. Type locality: Peru, Departamento de Huánuco, vicinity of Tingo
María, back-water near Puerto Nuevo, flowing into Río Tullamayo, 09°18'S, 75°59'W, 649 m. Holotype: ANSP
179125.
Distribution: Huallaga River basin at Tingo María, Peru.
Salcedo, N.J. (2006) Two new species of Chaetostoma (Siluriformes: Loricariidae) from the Huallaga River in central
Peru. Ichthyological Exploration of Freshwaters, 17, 207–220 [not seen].
Chaetostoma daidalmatos Salcedo, 2006
Chaetostoma stroumpoulos Salcedo, 2006
Sarmento-Soares, L.M., Martins-Pinheiro, R.F., Aranda, A.T. & Chamon, C.C. (2006) Microglanis pataxo, a new catfish
from southern Bahia coastal rivers, northeastern Brazil (Siluriformes: Pseudopimelodidae). Neotropical Ichthyology,
4, 157–166.
Microglanis pataxo Sarmento-Soares, Martins-Pinheiro, Aranda & Chamon, 2006: 158, Fig. 1. Type locality: Brazil,
Bahia: Itamarajú, Jundiar creek on road BR-101 after joint with road to Jucuruçu, in the neighbourhoods of the cityof Itamaraju (17º01'35"S 39º35'57"W). Holotype: MNRJ 28397.
Distribution: Peruipe, Jucurucu and Cahy Rivers, northeastern Brazil.
Sarmento-Soares, L.M., Martins-Pinheiro, R.F., Aranda A.T. & Chamon, C.C. (2006) Ituglanis cahyensis, a new catfish
from Bahia, Brazil (Siluriformes: Trichomycteridae). Neotropical Ichthyology, 4, 309–318.
Ituglanis cahyensis Sarmento-Soares, Martins-Pinheiro, Aranda & Chamon, 2006: 310, fig. 1. Type locality: Brazil,
Bahia: Prado, rio Palmares on road Guarany- Corumbau, in direction to Corumbau after the joint with the road to themouth of rio Cahy, (16º57'48"S, 39º16'33"W). Holotype: MNRJ 28404.
Distribution: Palmares River, Cahy River basin, Southeastern Bahia State, Brazil.
Sullivan J.P., Lundberg, J.G. & Hardman, M. (2006) A phylogenetic analysis of the major groups of catfishes (Teleostei:
Siluriformes) using rag1 and rag2 nuclear gene sequences. Molecular Phylogenetics and Evolution, 41, 636–662.
Tamang, L., Chaudhury, S. & Chauhury, D. (2006) On a new record of freshwater fish, Pseudolaguvia shawi (Hora) fromArunachal Pradesh, India (Teleostomi, Erethistidae). Zoos' Print Journal, 21 (11), 2443–2446.
Thomson, A.W. & Page, L.M. (2006) Genera of the Asian catfish families Sisoridae and Erethistidae (Teleostei: Siluriformes). Zootaxa, 1345, 1–90.
Diagnoses and review of named taxa above species level.
Synonymy of Hara and Erethistes.
Vari, R. P. & Ferraris, C.J., Jr. (2006) The Catfish genus Tetranematichthys (Auchenipteridae). Copeia, 2006, 168–180.
Tetranematichthys wallacei Vari & Ferraris, 2006: 169, figs. 1a, 2, 4. Type locality: Brazil, Amazonas, upper Rio Negro,
São Pedro, mouth of Igarapé do Ibará, approximately 0°15'S, 66°46'W. Holotype: MZUSP 31096,
Distribution: Orinoco, Negro Amazon and Tocantins River basins.
Villa-Verde, L. & Costa, W.J.E.M. (2006) A new glanapterygine catfish of the genus Listrura (Siluriformes: Trichomycteridae) from the southeastern Brazilian coastal plains. Zootaxa, 1142, 43–50.

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Listrura picinguabae Villa-Verde & Costa, 2006: 44, figs. 1–3. Type locality: Brazil: Estado de São Paulo: Município deUbatuba, serra do Mar, Picinguaba, small stream tributary to rio da Fazenda, on small road near Km 11 of the roadBR-101, Parque Estadual da Serra do Mar, about 23°20'S 44°45'W. Holotype: UFRJ 6111, 48.6 mm SL.
Distribution: da Fazenda River tributaries, São Paulo State, southeastern Brazil.
Vishwanath, W. & Darshan, A. (2006) A new species of the genus Batasio Blyth (Teleostei: Bagridae) from Manipur,
India. Zoos' Print Journal, 21, 2160–2163.
Batasio niger Vishwanath & Darshan, 2006: 2160, fig. 1. Type locality: Khujailok stream, Chandel district, Manipur,
India. Holotype: MUMF 9028.
Distribution: Chindwin River basin, Manipur, India.
Remarks: Publication includes illustration of tooth plates only; illustration of holotype available only on web versionof paper.
Wright, J.J. & Page, L.M. (2006) Taxonomic revision of Lake Tanganyikan Synodontis (Siluriformes: Mochokidae).
Florida Museum of Natural History Bulletin, 46, 99–154.
Synodontis grandiops Wright & Page, 2006: 109, figs. 3c, 8, 9, 10. Type locality: Mwakizega coastline, L. Tanganyika.
Holotype: BMNH 1982.4.13.4785.
Distribution: Lake Tanganyika.
Synodontis ilebrevis Wright & Page, 2006: 117, figs. 5d, 13, 14, 15. Type locality: Lake Tanganyika at Chaitika, Zambia.
Holotype: UF 160942.
Distribution. Lake Tanganyika, only from the Cape Chaitika area, Zambia.
Synodontis lucipinnis Wright & Page, 2006: 126, figs. 4a, 17, 18. Type locality: Zambia, Mpulungu, Musende Rocks,
08°46'00"S, 031°51'00"E. Holotype: SAIAB 39577.
Distribution: Lake Tanganyika, only from the Musende Rocks area, Zambia.
Key: Synodontis species of Lake Tanganyika.

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Ng, H.H. (2003a) Mystus impluviatus: a new species of bagrid catfish (Teleostei: Bagridae) from eastern Borneo. Copeia,

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2003, 373–378.
Ng, H.H. (2003b) A review of the Ompok hypophthalmus group of silurid catfishes with the description of a new species
from South-east Asia. Journal of Fish Biology, 62, 1296–1311.
Ng, H.H. (2003c) A revision of the south Asian sisorid catfish genus Sisor (Teleostei: Siluriformes). Journal of Natural
History, 37, 2871–2883.
Ng, H.H. (2003d) Kryptopterus paraschilbeides, a new species of catfish (Teleostei: Siluridae) from mainland Southeast
Asia. The Natural History Journal of the Chulalongkorn University, 3, 1–8.
Ng, H.H. (2003e) Arius verrucosus, a new species of freshwater ariid catfish (Teleostei: Ariidae) from the Mekong River.
Occasional Papers of the Museum of Zoology, University of Michigan, 734, 1–14.
Ng, H.H. (2003f) Ompok pinnatus, a new species of silurid catfish (Teleostei: Siluriformes: Siluridae) from mainland
Southeast Asia. Proceedings of the Biological Society of Washington, 116, 47–51.
Ng, H.H. (2003g) Clarias nigricans, a new species of clariid catfish (Teleostei: Siluriformes) from eastern Borneo. The
Raffles Bulletin of Zoology, 51, 393–398.
Ng, H.H. (2003h) Clarias insolitus, a new species of clariid catfish (Teleostei: Siluriformes) from southern Borneo. Zootaxa, 284, 1–8.
Ng, H.H. (2003i) Kryptopterus geminus, a new species of silurid catfish (Teleostei: Siluridae) from mainland Southeast
Asia. Zootaxa, 305, 1–11.
Ng, H.H. (2003j) Phylogeny and systematics of Bagridae. In: Arratia, G., Kapoor, B.G., Chardon, M. & Diogo, R. (Eds.),
Catfishes, Vol. 1, Science Publishers, Enfield, NH, USA, pp. 439–464.
Ng, H.H. (2004a) Wallago micropogon: a new species of silurid catfish (Teleostei: Siluridae) from mainland Southeast
Asia. Copeia, 2004, 92–97.
Ng, H.H. (2004b) New estuarine species of Mystus (Teleostei: Bagridae) from the Malay Peninsula. Copeia, 2004, 883–
887.
Ng, H.H. (2004c) Batasio elongatus, a new species of bagrid catfish from southwest Myanmar (Siluriformes: Bagridae).
Ichthyological Exploration of Freshwaters, 15, 67–70.
Ng, H.H. (2004d) Clarias sulcatus, a new walking catfish (Teleostei: Clariidae) from Pulau Redang. Ichthyological
Exploration of Freshwaters, 15, 289–294.
Ng, H.H. (2004e) Oreoglanis macronemus, a new species of glyptosternine catfish (Teleostei: Siluriformes: Sisoridae)
from northern Laos. The Raffles Bulletin of Zoology, 52, 209–213.
Ng, H.H. (2004f) Kryptopterus platypogon, a new silurid catfish (Teleostei: Siluridae) from Borneo. Zootaxa, 398, 1–8.
Ng, H.H. (2004g) Two new glyptosternine catfishes (Teleostei: Sisoridae) from Vietnam and China. Zootaxa, 428, 1–12.
Ng, H.H. (2004h) The Microsynodontis (Teleostei: Siluriformes: Mochokidae) of the lower Guinea region, west central
Africa, with the description of eight new species. Zootaxa, 531, 1–52.
Ng, H.H. (2004i) Rita macracanthus, a new riverine catfish (Teleostei: Bagridae) from South Asia. Zootaxa, 568, 1–12.
Ng, H.H. (2005a) Conta pectinata, a new erethistid catfish (Teleostei: Erethistidae) from northeast India. Ichthyological
Exploration of Freshwaters, 16, 23–28.
Ng, H.H. (2005b) Pseudolaguvia foveolata, a new catfish (Teleostei: Erethistidae) from northeast India. Ichthyological
Exploration of Freshwaters, 16, 173–178.
Ng, H.H. (2005c) Gogangra laevis, a new species of riverine catfish from Bangladesh (Teleostei: Sisoridae). Ichthyological Exploration of Freshwaters, 16, 279–286.
Ng, H.H. (2005d) Amblyceps carinatum, a new species of hillstream catfish from Myanmar (Teleostei: Amblycipitidae).
The Raffles Bulletin of Zoology, 53, 243–249.
Ng, H.H. (2005e) Glyptothorax botius (Hamilton, 1822), a valid species of catfish (Teleostei: Sisoridae) from northeast
India. Zootaxa, 930, 1–19.
Ng, H.H. (2005f) Erethistoides sicula, a new catfish (Teleostei: Erethistidae) from India. Zootaxa, 1021, 1–12.
Ng, H.H. (2005g) Two new species of Pseudolaguvia (Teleostei: Erethistidae) from Bangladesh. Zootaxa, 1044, 35–47.
Ng, H.H. & Chan, B.P.-L. (2005) Revalidation and redescription of Pterocryptis anomala (Herre, 1933), a catfish (Teleostei: Siluridae) from southern China. Zootaxa, 1060, 51–64.
Ng, H.H. & Dodson, J.J. (1999) Morphological and genetic descriptions of a new species of catfish, Hemibagrus
chrysops, from Sarawak, east Malaysia, with an assessment of phylogenetic relationships (Teleostei: Bagridae). The
Raffles Bulletin of Zoology, 47, 45–57.
Ng, H.H. & Edds, D.R. (2004) Batasio macronotus, a new species of bagrid catfish from Nepal (Teleostei: Bagridae).
Ichthyological Exploration of Freshwaters, 15, 295–300.
Ng, H.H. & Edds, D.R. (2005a) Two new species of Erethistoides (Teleostei: Erethistidae) from Nepal. Ichthyological

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Exploration of Freshwaters, 16, 239–248.
Ng, H.H. & Edds, D.R. (2005b) Two new species of Pseudecheneis, rheophilic catfishes (Teleostei: Sisoridae) from
Nepal. Zootaxa, 1047, 1–19.
Ng, H.H. & Ferraris, C.J., Jr. (2000) A review of the genus Hemibagrus in southern Asia, with descriptions of two new
species. Proceedings of the California Academy of Sciences, Series 4, 52, 125–142.
Ng, H.H. & Freyhof, J. (2001a) A review of the catfish genus Pterocryptis (Siluridae) in Vietnam, with the description of
two new species. Journal of Fish Biology, 59, 624–644.
Ng, H.H. & Freyhof, J. (2001b) Oreoglanis infulatus, a new species of glyptosternine catfish (Siluriformes: Sisoridae)
form central Vietnam. Journal of Fish Biology, 59, 1164–1169.
Ng, H.H. & Freyhof, J. (2003) Akysis clavulus, a new species of catfish (Teleostei: Akysidae) from central Vietnam. Ichthyological Exploration of Freshwaters, 14, 311–316.
Ng, H.H. & Freyhof, J. (2005) A new species of Pseudomystus (Teleostei: Bagridae) from Central Vietnam. Copeia,
2005, 745–750.
Ng, H.H. & Hadiaty, R.K. (2005) Two new bagrid catfishes (Teleostei: Bagridae) from the Alas River drainage, northern
Sumatra. Ichthyological Exploration of Freshwaters, 16, 83–92.
Ng, H.H. & Kottelat, M. (1996) Akysis fuscus, a new species of catfish (Teleostei: Akysidae) from the Kapuas basin,
Borneo. Ichthyological Exploration of Freshwaters, 7, 19–26.
Ng, H.H. & Kottelat, M. (1997) Silurichthys citatus, a new species of silurid catfish from western Borneo (Teleostei: Siluriformes). Ichthyological Exploration of Freshwaters, 7, 203–208.
Ng, H.H. & Kottelat, M. (1998a) Hyalobagrus, a new genus of miniature bagrid catfish from southeast Asia (Teleostei:
Siluriformes). Ichthyological Exploration of Freshwaters, 9, 335–346.
Ng, H.H. & Kottelat, M. (1998b) Pterocryptis buccata, a new species of catfish from western Thailand (Teleostei: Siluridae) with epigean and hypogean populations. Ichthyological Research, 45, 393–399.
Ng, H.H. & Kottelat, M. (1998c) The catfish genus Akysis Bleeker (Teleostei: Akysidae) in Indochina, with descriptionsof six new species. Journal of Natural History, 32, 1057–1097.
Ng, H.H. & Kottelat, M. (1999) Oreoglanis hypsiurus, a new species of glyptosternine catfish (Teleostei: Sisoridae). Ichthyological Exploration of Freshwaters, 10, 375–380.
Ng, H.H. & Kottelat, M. (2000a) A review of the genus Amblyceps (Osteichthyes: Amblycipitidae) in Indochina, withdescriptions of five new species. Ichthyological Exploration of Freshwaters, 11, 335–348.
Ng, H.H. & Kottelat, M. (2000b) Description of three new species of catfishes (Teleostei: Akysidae and Sisoridae) from
Laos and Vietnam. Journal of South Asian Natural History, 5, 7–15.
Ng, H.H. & Kottelat, M. (2000c) Helicophagus leptorhynchus, a new species of molluscivorous catfish from Indochina(Teleostei: Pangasiidae). The Raffles Bulletin of Zoology, 48, 55–58.
Ng, H.H. & Kottelat, M. (2000d) Cranoglanis henrici (Vaillant, 1893) a valid species of cranoglanidid catfish from
Indochina (Teleostei: Cranoglanididae). Zoosystema, 22, 847–852.
Ng, H.H. & Kottelat, M. (2001) A review of the genus Batasio (Teleostei: Bagridae) in Indochina, with the description of
B. tigrinus sp. n. from Thailand. Revue suisse de Zoologie, Annales de la Société zoologique suisse et du Muséumd'Histoire naturelle de Genève, 108, 495–511.
Ng, H.H. & Kottelat, M. (2003) Parakysis notialis, a new species of akysid catfish from Borneo (Siluriformes:
Akysidae). Ichthyological Research, 50, 48–51.
Ng, H.H. & Kottelat, M. (2004) Akysis vespa, a new species of catfish (Siluriformes: Akysidae) from the Ataran Riverdrainage (Myanmar). Ichthyological Exploration of Freshwaters, 25, 193–200.
Ng, H.H. & Kottelat, M. (2005) Caelatoglanis zontatus, a new genus and species of the Erethistidae (Teleostei: Siluriformes) from Myanmar, with comments on the nomenclature of Laguvia and Hara species. Ichthyological Exploration of Freshwaters, 16, 13–22.
Ng, H.H. & Lim, K.K.P. (1995) A revision of the southeast Asian catfish genus Parakysis (Teleostei: Akysidae), withdescriptions of two new species. Ichthyological Exploration of Freshwaters, 6, 255–266.
Ng, H.H. & Lim, K.K.P. (2000) Hemileiocassis panjang, a new genus and new species of bagrid catfish from Java
(Teleostei, Siluriformes). Beaufortia, 50, 191–195.
Ng, H.H. & Lim, K.K.P. (2005) The identity of Pseudomystus moeschii (Boulenger, 1890), with the description of two
new species of bagrid catfishes from Southeast Asia (Teleostei: Bagridae). Zootaxa, 851, 1–18.
Ng, H.H., Martin-Smith, K.M. & Ng, P.K.L. (2000) Hemibagrus furcatus, a new species of bagrid catfish (Teleostei: Siluriformes) from Sabah, east Malaysia. The Raffles Bulletin of Zoology, 48, 65–69.
Ng, H.H. & Ng, P.K.L. (1998) A revision of the South-east Asian catfish genus Silurichthys. Journal of Fish Biology, 52,

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291–333.
Ng, H.H. & Ng, P.K.L. (2001) A revision of the akysid catfish genus Acrochordonichthys Bleeker. Journal of Fish Biology, 58, 386–418.
Ng, H.H. & Rachmatika, I. (1999) The catfishes (Teleostei: Siluriformes) of Bentuang Karimun National Park, West Kalimantan, Indonesia. The Raffles Bulletin of Zoology, 47, 167–183.
Ng, H.H. & Rachmatika, I. (2005) Glyptothorax exodon, a new species of rheophilic catfish from Borneo (Teleostei:
Sisoridae). The Raffles Bulletin of Zoology, 53, 251–255.
Ng, H.H. & Rainboth, W.J. (1999) The bagrid catfish genus Hemibagrus (Teleostei: Siluriformes) in central Indochina
with a new species from the Mekong River. The Raffles Bulletin of Zoology, 47, 555–576.
Ng, H.H. & Rainboth, W.J. (2001) A review of the sisorid catfish genus Oreoglanis (Siluriformes: Sisoridae) withdescriptions of four new species. Occasional Papers of the Museum of Zoology, University of Michigan, 732, 1–34.
Ng, H.H. & Rainboth, W.J. (2005) Four new species of Akysis (Teleostei: Siluriformes: Akysidae) from mainland Southeast Asia, with comments on A. similis. The Raffles Bulletin of Zoology, Supplement 13, 33–42.
Ng, H.H. & Sabaj, M.H. (2005) Akysis hardmani (Siluriformes: Akysidae), a new species of catfish from Thailand. Ichthyological Exploration of Freshwaters, 16, 215–222.
Ng, H.H. & Siebert, D.J. (1998) A revision of the akysid catfish genus Breitensteinia Steindachner with descriptions of
two new species. Journal of Fish Biology, 53, 645–657.
Ng, H.H. & Siebert, D.J. (2002) Rediscovery and redescription of Ompok weberi (Hardenberg, 1936), a poorly known
species of silurid catfish (Teleostei: Siluriformes) from Borneo. The Raffles Bulletin of Zoology, 50, 169–173.
Ng, H.H. & Siebert, D.J. (2004) A new species of the catfish genus Akysis (Siluriformes: Akysidae) from southern
Borneo. Zootaxa, 733, 1–8.
Ng, H.H. & Siebert, D.J. (2005) Pseudomystus stenogrammus, a new species of bagrid catfish from Borneo (Teleostei,
Bagridae). Zootaxa, 813, 1–7.
Ng, H.H. & Sparks, J.S. (2002) Plotosus fisadoha, a new species of marine catfish (Teleostei: Siluriformes: Plotosidae)
from Madagascar. Proceedings of the Biological Society of Washington, 115, 80–85.
Ng, H.H. & Sparks, J.S. (2003) The ariid catfishes (Teleostei: Siluriformes: Ariidae) of Madagascar, with the descriptionof two new species. Occasional Papers of the Museum of Zoology, University of Michigan, 735, 1–21.
Ng, H.H. & Sparks, J.S. (2005) Revision of the endemic Malagasy catfish family Anchariidae (Teleostei: Siluriformes),
with descriptions of a new genus and three new species. Ichthyological Exploration of Freshwaters, 16, 303–323.
Ng, H.H. & Tan, H.H. (1999) The fishes of the Endau drainage, Peninsular Malaysia with descriptions of two new species of catfishes (Teleostei: Akysidae, Bagridae). Zoological Studies, 38, 350–366.
Ng, H.H. & Tan, H.H. (2000) A new species of Encheloclarias (Siluriformes: Clariidae) from Sumatra. Journal of Fish
Biology, 57, 536–539.
Ng, H.H. & Tan, H.H. (2002) Redescription of Acrochordonichthys ischnosoma Bleeker, 1858, a poorly known speciesof akysid catfish (Teleostei: Siluriformes) from Sumatra and Java. The Raffles Bulletin of Zoology, 50, 449–452.
Ng, H.H. & Tan, H.H. (2004) Ompok platyrhynchus, a new silurid catfish (Teleostei: Siluridae) from Borneo. Zootaxa,
580, 1–11.
Ng, H.H., Wirjoatmodjo, S. & Hadiaty, R.K. (2001a) Mystus punctifer, a new species of bagrid catfish (Teleostei: Siluriformes) from northern Sumatra. The Raffles Bulletin of Zoology, 49, 355–358.
Ng, H.H., Wirjoatmodjo, S. & Hadiaty, R.K. (2001b) Hemibagrus caveatus, a new species of bagrid catfish (Teleostei:
Siluriformes) from northern Sumatra. The Raffles Bulletin of Zoology, 49, 359–361.
Ng, H.H., Wirjoatmodjo, S. & Hadiaty, R.K. (2004) Kryptopterus piperatus, a new species of silurid catfish (Teleostei:
Siluridae) from northern Sumatra. Ichthyological Exploration of Freshwaters, 15, 91–95.
Ng, P.K.L. & Lim, K.K.P. (1993) The Southeast Asian catfish genus Encheloclarias (Teleostei: Clariidae), with descriptions of four new species. Ichthyological Exploration of Freshwaters, 4, 21–37.
Ng, P.K.L. & Ng, H.H. (1995) Hemibagrus gracilis, a new species of large riverine catfish (Teleostei: Bagridae) from
Peninsular Malaysia. The Raffles Bulletin of Zoology, 43, 133–142.
Nguyen, H.D. & Nguyen, V.H. (2001) [Two newly found fish species of Pareuchiloglanis genus (Sisoridae, Siluriformes) in Vietnam]. Tap chí Sinh Hoc [Journal of Biology], 23, 66–71. [In Vietnamese, with English title and summary; English title only roughly agrees with Vietnamese title]
Nichols, J.T. (1919a) "Cascudos" brazileiros do genero Plecostomus do Museu Paulista (Brasilian catfishes of the genus
Plecostomus from the Museu Paulista). Revista do Museu Paulista, 11, 409–426.
Nichols, J.T. (1919b) Um novo genero de cascudos da familia Loricariidae. Revista do Museu Paulista, 11, 533–535,
539–540.

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Nichols, J.T. (1925) Some Chinese fresh-water fishes. X.— Subgenera of bagrin catfishes. XI.— Certain apparently
undescribed carps from Fukien. XII.— A small goby from the central Yangtze. XIII.— A new minnow referred to
Leucogobio. XIV.— Two apparently undescribed fishes from Yunnan. American Museum Novitates, 185, 1–7.
Nichols, J.T. (1926a) Some Chinese fresh-water fishes. XV.— Two apparently undescribed catfishes from Fukien.
XVI.— Concerning gudgeons related to Pseudogobio, and two new species of it. XVII.— Two new rhodeins. American Museum Novitates, 214, 1–7.
Nichols, J.T. (1926b) Some Chinese fresh-water fishes. XVIII.— New species in recent and earlier Fukien collections.
American Museum Novitates, 224, 1–7.
Nichols, J.T. (1930) Some Chinese fresh-water fishes. XXVI.— Two new species of Pseudogobio. XXVII.—A new catfish from northeastern Kiangsi. American Museum Novitates, 440, 1–5.
Nichols, J.T. (1931) Some Chinese fresh-water fishes. XXVIII.— A collection from Chungan Hsien, northwestern Fukien. American Museum Novitates, 449, 1–3.
Nichols, J.T. (1940) Results of the Archbold expeditions. No. 30.— New catfishes from northern New Guinea. American
Museum Novitates, 1093, 1–3.
Nichols, J.T. (1941) Four new fishes from western China. American Museum Novitates, 1107, 1–3.
Nichols, J.T. (1943) The Fresh-water Fishes of China, Natural History of Central Asia, Vol. 9, American Museum of
Natural History, New York, 322 p., 10 pl.
Nichols, J.T. (1956) A new pygidiin catfish from Argentina. American Museum Novitates, 1760, 1–3.
Nichols, J.T. & Griscom, L. (1917) Fresh-water fishes of the Congo basin obtained by the American Museum Congo
expedition, 1909–1915. Bulletin of the American Museum of Natural History, 37, 653–756, pls. 64–83.
Nichols, J.T. & La Monte, F.R. (1933a) A new catfish, Amphilius pictus, and a discussion of a small Liberian collectionof fishes. American Museum Novitates, 626, 1–3.
Nichols, J.T. & La Monte, F.R. (1933b) New fishes from the Kasai district of the Belgian Congo. American Museum
Novitates, 656, 1–6.
Nichols, J.T. & La Monte, F.R. (1934) More new fishes from the Kasai district of the Belgian Congo. American Museum
Novitates, 723, 1–6.
Nichols, J.T. & La Monte, F.R. (1953) Two new African catfishes from eastern French Equatorial Africa. American
Museum Novitates, 1648, 1–4.
Nichols, J.T. & Pope, C.H. (1927) The fishes of Hainan. Bulletin of the American Museum of Natural History, 54, 321–
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Nico, L.G. & Pinna, M.C.C., de. (1996) Confirmation of Glanapteryx anguilla (Siluriformes, Trichomycteridae) in the
Orinoco River basin, with notes on the distribution and habits of the Glanapteryginae. Ichthyological Exploration ofFreshwaters, 7, 27–32.
Nielsen, J.G. (1974) Fish Types in the Zoological Museum of Copenhagen, Copenhagen, 115 p.
Nijssen, H. (1970) Revision of the Surinam catfishes of the genus Corydoras La Cepède, 1803 (Pisces, Siluriformes,
Callichthyidae). Beaufortia, 18, 1–75.
Nijssen, H. (1971) Two new species and one new subspecies of the South American catfish genus Corydoras (Pisces,
Siluriformes, Callichthyidae). Beaufortia, 19, 89–98.
Nijssen, H. (1972) Records of the catfish genus Corydoras from Brazil and French Guiana with descriptions of eight new
species (Pisces, Siluriformes, Callichthyidae). Netherlands Journal of Zoology, 21, 412–433, pls. 1–3.
Nijssen, H. & Isbrücker, I.J.H. (1967) Notes on the Guiana species of Corydoras La Cepède, 1803, with descriptions ofseven new species and designation of a neotype for Corydoras punctatus (Bloch, 1794) –(Pisces, Cypriniformes,
Callichthyidae). Zoologische Mededelingen (Leiden), 42, 21–50, pls. 1–5.
Nijssen, H. & Isbrücker, I.J.H. (1970) The South American catfish genus Brochis Cope, 1872 (Pisces, Siluriformes, Callichthyidae). Beaufortia, 18, 151–168.
Nijssen, H. & Isbrücker, I.J.H. (1971) Two new species of the catfish genus Corydoras from Brazil and Peru (Pisces, Siluriformes, Callichthyidae). Beaufortia, 18, 183–189.
Nijssen, H. & Isbrücker, I.J.H. (1975) Cataphractus punctatus Bloch, 1794 (Pisces, Siluriformes, Callichthyidae):request for invalidation of neotype and validation of a rediscovered syntype as lectotype. Bulletin of Zoological
Nomenclature, 32, 63–64, pl. 1.
Nijssen, H. & Isbrücker, I.J.H. (1976a) The South American plated catfish genus Aspidoras R. von Ihering, 1907, withdescriptions of nine new species from Brazil (Pisces, Siluriformes, Callichthyidae). Bijdragen tot de Dierkunde, 46,
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Nijssen, H. & Isbrücker, I.J.H. (1976b) Corydoras ornatus, a new species of callichthyid catfish from the Rio Tapajós

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drainage, Brazil (Pisces, Siluriformes, Callichthyidae). Bulletin Zoölogisch Museum, Universiteit van Amsterdam, 5,
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Nijssen, H. & Isbrücker, I.J.H. (1976c) A new callichthyid catfish, Corydoras gracilis, from Brazil. Tropical Fish Hobbyist, 25, 90–98.
Nijssen, H. & Isbrücker, I.J.H. (1979) Chronological enumeration of nominal species and subspecies of Corydoras(Pisces, Siluriformes, Callichthyidae). Bulletin Zoölogisch Museum, Universiteit van Amsterdam, 6, 129–135.
Nijssen, H. & Isbrücker, I.J.H. (1980a) On the identity of Corydoras nattereri Steindachner 1877 with the description ofa new species Corydoras prionotos (Pisces, Siluriformes, Callichthyidae). Beaufortia, 30, 1–9.
Nijssen, H. & Isbrücker, I.J.H. (1980b) A review of the genus Corydoras La Cepède, 1803 (Pisces, Siluriformes, Callichthyidae). Bijdragen tot de Dierkunde, 50, 190–220.
Nijssen, H. & Isbrücker, I.J.H. (1980c) Aspidoras virgulatus n. sp., a plated catfish from Espírto Santo, Brazil (Pisces,
Siluriformes, Callichthyidae). Bulletin Zoölogisch Museum, Universiteit van Amsterdam, 7, 133–139.
Nijssen, H. & Isbrücker, I.J.H. (1980d) Three new Corydoras species from French Guiana and Brazil (Pisces, Siluriformes, Callichthyidae). Netherlands Journal of Zoology, 30, 494–503.
Nijssen, H. & Isbrücker, I.J.H. (1982) Corydoras boehlkei, a new catfish from the Rio Caura system in Venezuela(Pisces, Siluriformes, Callichthyidae). Proceedings of the Academy of Natural Sciences, Philadelphia, 134, 139–
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Nijssen, H. & Isbrücker, I.J.H. (1983a) Brochis britskii, a new species of plated catfish from the upper Rio Paraguai system, Brazil (Pisces, Siluriformes, Callichthyidae). Bulletin Zoölogisch Museum, Universiteit van Amsterdam, 9,
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Nijssen, H. & Isbrücker, I.J.H. (1983b) Review of the genus Corydoras from Colombia, with descriptions of two new
species (Pisces, Siluriformes, Callichthyidae). Beaufortia, 33, 53–71.
Nijssen, H. & Isbrücker, I.J.H. (1983c) Sept espèces nouvelles de poissons-chats cuirassés du genre Corydoras La
Cepède, 1803, de Guyane française, de Bolivie, d'Argentine, du Surinam et du Brésil (Pisces, Siluriformes, Callichthyidae). Revue Française d'Aquariologie et Herpetologie, 10, 73–84.
Nijssen, H. & Isbrücker, I.J.H. (1985) Lasiancistrus scolymus, a new species of mailed catfish from Rio Aripuanã, Est.
Mato Grosso do Sul, Brazil (Pisces, Siluriformes, Loricariidae). Bijdragen tot de Dierkunde, 55, 242–248.
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FERRARISINDEX
Names in bold indicate valid taxa; page numbers in italics indicate primary entry for the name.
5-tentaculatus, Heterobranchus, 197
6-tentaculatus, Heterobranchus, 197,
200
7-radiatus, Silurus, 122
9-radiatus, Silurus, 201
11-radiatus, Silurus, 335
12-radiatus, Silurus, 177
16-radiatus, Silurus, 44
38-radiatus, Silurus, 70Aaaldereni, Hemipimelodus, 32
abbreviatus, Bagrus, 95
abbreviatus, Clarias, 148
abbreviatus, Plotosus, 351
aboinensis, Clarias (Clarioides), 144
Abron, 342
absconditus, Pimelodus, 336
abuelo, Cheirocerus, 332
abuelo, Sovichthys, 332
aburrense, Chaetostoma, 227
aburrensis, Hypostomus, 227
Acanthicini, 217
Acanthicus, 217, 218
acanthicus, Rinelepis, 218
Acanthicus adonis, 218
Acanthicus canensis, 266
Acanthicus hystrix, 218
Acanthobunocephalus, 57
Acanthobunocephalus nicoi, 57
acanthochiroides, Batrochoglanis,352
acanthochiroides, Pseudopimelodus,
352
acanthochirus, Pseudopimelodus, 353
Acanthocleithron, 303
Acanthocleithron chapini, 303
Acanthodemus, 278
Acanthodoras, 167
Acanthodoras cataphractus, 167
Acanthodoras depressus, 167
Acanthodoras spinosissimus, 168
acanthomias, Synodontis, 311
Acanthonotus, 356
Acanthonotus Hardwickii, 356
acanthoperca, Synodontis, 441
Acanthopoma, 403
Acanthopoma annectens, 403
Acanthopoma bondi, 413
accipenser, Loricaria, 270
Acentronichthys, 180
Acentronichthys leptos, 180
Acestra, 217, 237
Acestra acus, 237
Acestra amazonum, 237
Acestra gladiolus, 237

Acestra gladius, 239
Acestra Knerii, 238
Acestra oxyrrhyncha, 239
Acestra oxyrryncha, 239
acestrichthys, Farlowella, 238
Acestridiinae, 217
Acestridium, 217, 218
Acestridium colombiense, 218
Acestridium colombiensis, 218
Acestridium dichromum, 218
Acestridium discus, 218
Acestridium martini, 218
Acestrini, 217
acicularis, Sperata,106
Acipenser plecostomus, 258, 259
acipenserinus, Hemiodon, 247
acipenserinus, Hemiodontichthys, 247
acipenserinus, Leptodoras, 172
acipenserinus, Oxydoras, 172
acrensis, Corydoras, 112
acrocephalus, Arius, 47
Acrochordonichthyini, 12
Acrochordonichthys, 12
Acrochordonichthys Büttikoferi, 13
Acrochordonichthys chamaeleon, 12
Acrochordonichthys falcifer, 12
Acrochordonichthys guttatus, 12
Acrochordonichthys gyrinus, 12
Acrochordonichthys ischnosoma, 12
Acrochordonichthys mahakamensis,
12
Acrochordonichthys melanogaster, 13
Acrochordonichthys obscurus, 13
Acrochordonichthys pachyderma, 13
Acrochordonichthys platycephalus, 13
Acrochordonichthys rugosus, 13
Acrochordonichthys septentrionalis,
13
Acrochordonichthys strigosus, 13
Acrochordonichthys varius, 13
Acrochordonichthys zonatus, 13
aculeatus, Chaetostomus, 272
aculeatus, Leiocassis, 92
aculeatus, Plecostomus, 224
aculeatus, Pterygoplichthys, 272
acus, Acestra, 237
acus, Farlowella, 237
acus venezuelensis, Farlowella, 239
acuta, Loricaria, 269
acuticeps, Auchenoglanis, 154acutirostris, Oxyropsis, 276
acutirostris, Arius, 31
acutirostris, Chrysichthys, 158
acutirostris, Neotropius, 359
acutirostris, Oxyropsis, 276

CHECKLIST OF CATFISHES, RECENT AND FOSSIL

acutirostris, Pseudeutropius, 359
acutivelis, Arius, 155, 158
acutus, Arius, 51
acutus, Corydoras, 112
acutus, Loricariichthys, 269
Adansonii, Bagrus, 365
Adelopeltis, 107
Adelopeltis angusticeps, 107
Adelopeltis laticeps, 107
adiposalis, Pseudobagrus, 101
adolfoi, Corydoras, 112
adonis, Acanthicus, 218
adspersus, Callichthys, 127
aegyptiaca, Eopeyeria, 428
aegyptiacus, Ariopsis, 428
Aelurichthys, 37
Aelurichthys isthmensis, 38Ælurichthys longispinis, 38Ælurichthys nuchalis, 38
Aelurichthys panamensis, 37, 38
Aelurichthys pinnimaculatus, 38
Aelurichthys scutatus, 38
aeneum, Hoplosoma, 112
aeneus, Ariodes, 49
aeneus, Corydoras, 112
aeneus, Pimelodus, 215
aequalicuspis, Rineloricaria, 293
aequibarbis, Arius, 55
aequilabialis, Pangasius, 327
aequinoctiale, Chaetostoma, 227
aequinoctialis, Chaetostomus, 227
aequus, Otolithus (Arius), 435
affine, Platystoma, 330
affinis, Ailia, 356
affinis, Amblydoras, 168
affinis, Auchenipterus(Pseudauchenipterus), 76
affinis, Bagrus, 107affinis, Batasio, 85
affinis, Callichthys, 111
affinis, Doras, 168
affinis, Hassar, 171affinis, Hypostomus, 251affinis, Macrotocinclus, 271
affinis, Malapterurus, 301
affinis, Otocinclus, 271
affinis, Oxydoras, 171affinis, Pimelodus, 208
affinis, Plecostomus, 251affinis, Pseudauchenipterus, 76affinis, Silurus (Callichrus), 374
afghana, Silurus, 376
africana, Arius, 31
africana, Arius falcarius, 31
africanus, Otolithus (Arius), 435

Zootaxa 1418 © 2007 Magnolia Press · 549

afrofischeri, Synodontis, 311
Afro-fischeri, Synodontis, 311
Agamyxis, 168
Agamyxis albomaculatus, 168
Agamyxis pectinifrons, 168
Agassizi, Pseudopimelodus, 353
agassizii, Cathorops, 39
agassizii, Corydoras, 112
Agassizii, Pinirampus, 339
Agassizii, Rhinelepis, 289
agassizii, Tachisurus, 39
agastor, Paraloricaria, 278
agboyiensis, Clarias, 139
agboyiensis, Clarias (Clarioides), 139
Agenciosus polystictus, 70
Ageneiosi, 68
Ageneiosus, 68
Ageneiosus (Pseudageneiosus)
therezinae, 69
Ageneiosus armatus, 68, 69
Ageneiosus atronasus, 68
Ageneiosus barranquerensis, 69
Ageneiosus brevifilis, 68, 69
Ageneiosus brevis, 69
Ageneiosus caucanus, 70
Ageneiosus childreni, 366
Ageneiosus dentatus, 70
Ageneiosus freiei, 70
Ageneiosus gabardinii, 69
Ageneiosus guianensis, 70
Ageneiosus inermis, 69
Ageneiosus madeirensis, 69
Ageneiosus magoi, 69
Ageneiosus marmoratus, 69
Ageneiosus marquesi, 70
Ageneiosus melanopogon, 68
Ageneiosus militaris, 68, 69
Ageneiosus mino, 38
Ageneiosus ogilviei, 69
Ageneiosus pardalis, 70
Ageneiosus parnaguensis, 70
Ageneiosus piperatus, 70
Ageneiosus polystictus, 70
Ageneiosus quadrifilis, 78
Ageneiosus rondoni, 69
Ageneiosus sebae, 69
Ageneiosus ucayalensis, 70
Ageneiosus uruguayensis, 70
Ageneiosus valenciennesi, 69
Ageneiosus virgo, 70
Ageneiosus vittatus, 70
Ageneisus polystictus, 70
Ageniosus (Silonia) diaphina, 86
Ageniosus (Silonia) lurida, 366
Ageniosus axillaris, 69
Ageniosus militaris, 69
Ageniosus porphyreus, 70
ageneiosus, Batrachocephalus, 38
Agenius, 68

Aglyptosternon, 387
Agmus, 58
Agmus lyriformis, 60agna, Hypostomus, 251ãgnã, Plecostomus, 251
agricolus, Bagrus, 107
aguaboensis, Ancistrus, 219
aguadulce, Cathorops, 39
aguadulce, Galeichthys, 39
aguanai, Exallodontus, 333
aguarague, Trichomycterus, 441
Aguarunichthys, 329
Aguarunichthys inpai, 329
Aguarunichthys tocantinsensis, 329
Aguarunichthys torosus, 329
aguilerae, Silvaichthys, 76
agustini, Farlowella, 240ahli, Auchenoglanis, 162ahli, Parauchenoglanis, 162
Ailia, 356
Ailia affinis, 356
Ailia coila, 356
Ailia occidentalis, 360
Ailia punctata, 356
Ailia somalensis, 360
Ailichthyoidei, 356
Ailiichthys, 356
Ailiichthys punctata, 356
Ailurichthys, 37
ailurus, Pimelodus, 205
akamai, Micromyzon, 61
akhtari, Glyptosternon, 387
akhtari, Glyptosternum, 387
akiri, Anaspidoglanis, 154
akiri, Auchenoglanis, 154aky, Epactionotus, 236
Akyses, 12
Akysidae, 12
Akysis, 12
Akysis alfredi, 16
Akysis armatus, 98
Akysis baramensis, 16
Akysis brachybarbatus, 13
Akysis clavulus, 14
Akysis clinatus, 14
Akysis ephippifer, 14
Akysis filifer, 16
Akysis fuliginatus, 14
Akysis fuscus, 16
Akysis hendricksoni, 14
Akysis heterurus, 14
Akysis inermis, 16
Akysis kurzii, 18
Akysis leucorhynchus, 16
Akysis longifilis, 442
Akysis macronema, 16, 17
Akysis maculipinnis, 14
Akysis major, 391
Akysis meridionalis, 17

550 · Zootaxa 1418 © 2007 Magnolia Press

Akysis microps, 14
Akysis nitidus, 17
Akysis pictus, 14
Akysis prashadi, 14
Akysis pseudobagarius, 16, 17
Akysis recavus, 14
Akysis similis, 17
Akysis sinensis, 17
Akysis sinesis, 17
Akysis variegatus, 15
Akysis variegatus variegatus, 14
Akysis varius, 15
Akysis vespa, 15
alaknandi, Glyptothorax, 388
alaknandi, Glyptothorax brevipinnis,388
alasensis, Mystus, 94
alatus, Arius, 53
alatus, Hypostomus, 251
albater, Aspidoras, 108
albater, Noturus, 211, 441
alberti, Synodontis, 311
albescens, Glanidium, 75
albicans, Arius, 337
albicans, Bagrus, 54
albicans, Pimelodus, 337
albicollaris, Leiocassis, 104
albicrux, Trachelyopterus, 79
albicrux, Trachycorystes, 79
albidus, Arius, 337
albidus, Callichthys, 128
albidus, Pimelodus, 40, 204
albifasciatus, Bunocephalus, 62
albilabris, Bagrus, 95
albilabris, Paraplotosus, 349
albilabris, Plotosus, 349
albinotatus, Trichomycterus, 414
albocinctus, Hemiancistrus, 223
albofasciatus, Pimelodus, 337
albolineata, Synodontis, 311
albolineatus, Corydoras, 112
albolineatus, Mystus, 94
albolineatus, Synodontis, 311
albomaculatus, Agamyxis, 168
albomaculatus, Doras, 168
albomaculatus, Panaqolus, 277
albomaculatus, Panaque, 277
albomarginatus, Cephalosilurus, 353
albomarginatus, Leiocassis, 101
albomarginatus, Pseudobagrus, 101
albomarginatus, Pseudopimelodus,
353
albopunctatus, Clarias, 139
albopunctatus, Hypostomus, 251
albopunctatus, Plecostomus, 251
aleppensis, Macrones, 97
aleuropsis, Bunocephalus, 58
alexandri, Lophiosilurus, 353
alfaroi, Rhamdia, 199

FERRARIS

alfredi, Akysis, 16
alfredi, Pseudobagarius, 16alga, Chaetostomus, 221
alikunhii, Horaglanis, 152
alipionis, Hemipsilichthys, 262
alipionis, Isbrueckerichthys, 262
Allabenchelys, 137
Allabenchelys attemsi, 137
Allabenchelys brevior, 137
Allabenchelys dhonti, 142
Allabenchelys engelseni, 142
Allabenchelys laticeps, 137
Allabenchelys longicauda, 137
Allabenchelys manyangae, 137
Allabenchelys pietschmanni, 138
alleni, Apomatoceros, 403
alluaudi, Clarias, 139
aloikae, Bunocephalus amaurus, 58
alta, Tympanopleura, 69
altae, Centromochlus, 73
altamazonicum, Lepthoplosternum,
128
alternatum, Pygidium, 414
alternatus, Trichomycterus, 414
alternifasciatum, Chaetostoma, 227
alternus, Ochmacanthus, 409
alterum, Pygidium, 415
alterus, Trichomycterus, 415
altifrons, Pangasius, 325
altipinnis, Auchenoglanis, 162
altipinnis, Eutropius, 365
altipinnis, Hemiloricaria, 245
altipinnis, Liposarcus, 291
altipinnis, Loricaria, 245
altipinnis, Parauchenoglanis, 162
altipinnis, Pimelodella, 189
altipinnis, Pimelodus, 189
altissimus, Pimelodus, 337
altocorpus, Farlowella, 444
altus, Silurus, 381
alula, Doumea, 24
aluuensis, Chrysichthys, 155alvarezi, Gymnallabes, 150
amandajanea, Corydoras, 112
amapaensis, Corydoras, 113
Amaralia, 57
Amaralia hypsiura, 57
Amarginops, 153
Amarginops hildae, 153
Amarginops platus, 153
amatitlanensis, Rhamdia, 198
amaurus, Bunocephalus 58
amazona, Farlowella, 237
amazonae, Helogenes, 134amazonensis, Parotocinclus, 281
amazonica, Dekeyseria, 118
amazonica, Loricaria, 270
amazonicus, Bunocephalus, 59
amazonicus, Dysichthys, 59

amazonicus, Ituglanis, 406
amazonicus, Trichomycterus, 406
amazonum, Acesta, 237
amazonum, Loricaria apeltogaster,288
amazonum, Rhinodoras, 170
amazonus, Pseudohemiodon, 288
ambiacus, Corydoras, 113
amblops, Felichthys, 79
amblops, Trachelyopterus, 79
Amblycepinae, 17
Amblyceps, 17, 98
Amblyceps apangi, 17
Amblyceps arnunachalensis, 17
Amblyceps caecutiens, 17, 18
Amblyceps cæcutiens, 18
Amblyceps carinatum, 18
Amblyceps deyi, 17
Amblyceps foratum, 18
Amblyceps horae, 99
Amblyceps inermis, 17
Amblyceps laticeps, 18
Amblyceps macropterus, 18
Amblyceps mangois, 18
Amblyceps marginatoides, 19
Amblyceps marginatus, 19
Amblyceps mucronatum, 18
Amblyceps murraystuarti, 18
Amblyceps murray-stuarti, 18
Amblyceps platycephalus, 18
Amblyceps serratum, 18
Amblyceps tenuispinis, 18
Amblyceps variegatum, 19
Amblycipitidae, 17
Amblydoras, 168, 179
Amblydoras affinis, 168
Amblydoras bolivarensis, 168
Amblydoras gonzalezi, 168
Amblydoras monitor, 169, 169
Amblydoras nauticus, 169
Amblydoras truncatus, 168
amblyurum, Steindachneridion, 343
amblyurus, Steindachneria, 343
amboinensis, Cryptopterus, 369
ambrosettii, Liposarcus, 291
ambrosettii, Pterygoplichthys, 291
ambyiacus, Auchenipterus, 71
Ameiurina, 203
Ameiurus, 203, 204
Ameiurus brunneus, 204
Ameiurus cancellatus, 211
Ameiurus catus, 204
Ameiurus hazenensis, 204
Ameiurus lavetti, 204
Ameiurus leidyi, 204
Ameiurus macgrewi, 204
Ameiurus maconnellii, 211
Ameiurus melas, 205
Ameiurus natalis, 205

CHECKLIST OF CATFISHES, RECENT AND FOSSIL

Ameiurus nebulosus, 205
Ameiurus pectinatus, 206
Ameiurus platycephalus, 206
Ameiurus primaevus, 207
Ameiurus reticulatus, 206
Ameiurus sawrockensis, 206
Ameiurus serracanthus, 206
Ameiurus vespertinus, 206
amekiensis, Otolithus (Arius), 435
amemiyae, Aoria, 91
amemiyai, Aoria, 91
americanus, Cataphractus, 167
Amissidens, 32
Amissidens hainesi, 32
Amiurus, 204
Amiurus australis, 208
Amiurus bolli, 205
Amiurus brachyacanthus, 205
Amiurus brunneus, 204
Amiurus cragini, 205
Amiurus decorus, 210
Amiurus dugèsii, 208
Amiurus erebennus, 205
Amiurus lophius, 204
Amiurus meeki, 209
Amiurus meridionalis, 208
Amiurus mexicanus, 209
Amiurus mispilliensis, 206
Amiurus natalis analis, 205
Amiurus niveiventris, 204
Amiurus obesus, 205
Amiurus prosthistius, 205
Ammoglanis, 403
Ammoglanis diaphanus, 403
Ammoglanis pulex, 403
ammophilus, Aphanotorulus, 225
amphibelus, Corydoras, 113
Amphiliidae, 20
Amphilius, 20, 21
Amphilius atesuensis, 21
Amphilius baudoni, 21
Amphilius baudoni uniformis, 21
Amphilius baudoni var. uniformis., 21
Amphilius brevidorsalis, 23
Amphilius brevis, 21
Amphilius cryptobullatus, 21
Amphilius grammatophorus, 22, 23
Amphilius grammatophorus
brevipinna, 22
Amphilius grammatophorus
inaequalis, 23
Amphilius grammatophorus
marmoratus, 23
Amphilius grammatophorus var.
brevipinna, 23
Amphilius grammatophorus var.
inaequalis, 23
Amphilius grandis, 23
Amphilius hargeri, 23

Zootaxa 1418 © 2007 Magnolia Press · 551

Amphilius jacksonii, 21
Amphilius kakrimensis, 21
Amphilius kivuensis, 21
Amphilius krefftii, 23
Amphilius lamani, 22
Amphilius lampei, 22
Amphilius laticaudatus, 22
Amphilius lentiginosus, 22
Amphilius longirostris, 22
Amphilius maesii, 22
Amphilius natalensis, 22
Amphilius nigricaudatus, 21
Amphilius nigricaudatus
multipunctata, 21
Amphilius nigricaudatus var.
multipunctata, 21
Amphilius notatus, 27
Amphilius opisthophthalmus, 22
Amphilius oxyrhinus, 23
Amphilius pictus, 21
Amphilius platychir, 22
Amphilius platychir cubangoensis, 23
Amphilius platychir var. cubangoensis,
23
Amphilius pulcher, 23
Amphilius pulcher ephippiata, 23
Amphilius rheophilus, 23
Amphilius transvaaliensis, 23
Amphilius uranoscopus, 23
Amphilius zairensis, 23
amphiloxa, Cetopsis, 131
amphiloxus, Hemicetopsis, 131
amplexicauda, Clarias, 148
Anacanthus, 301
Anadoras, 169, 179
Anadoras grypus, 169
Anadoras regani, 169
Anadoras weddellii, 169
anale, Chaetostoma, 227
analis, Amiurus natalis, 205
analis, Hypocolpterus, 227
analis, Leiocassis (Dermocassis), 101
analis, Pseudobagrus, 101
analis, Trachycorystes, 80
anamalaiensis, Glyptothorax, 388
ananas, Physopyxis, 176
Anaspidoglanis, 153
Anaspidoglanis akiri, 154
Anaspidoglanis boutchangai, 154
Anaspidoglanis macrostomus, 154
Anchariidae, 29
Ancharius, 29
Ancharius brevibarbis, 29
Ancharius fuscus, 29
Ancharius griseus, 29
Ancistomus, 283
Ancistri, 216
ancistroides, Hypostomus, 251
ancistroides, Plecostomus, 251

Ancistrus, 216, 219
Ancistrus (Hemiancistrus) yaravi, 273
Ancistrus (Pseudancistrus) coquenani,
287
Ancistrus aguaboensis, 219
Ancistrus annectens, 242
Ancistrus barrae, 272
Ancistrus baudensis, 220
Ancistrus bodenhameri, 219
Ancistrus bolivianus, 219
Ancistrus Bovallii, 267
Ancistrus brachyurus, 234
Ancistrus brevifilis, 219
Ancistrus brevifilis bodenhameri, 219
Ancistrus brevipinnis, 219
Ancistrus bufonius, 219
Ancistrus caucanus, 219
Ancistrus centrolepis, 219
Ancistrus chagresi, 220, 222
Ancistrus cirrhosus, 220
Ancistrus cirrhosus dubius, 221
Ancistrus claro, 220
Ancistrus clementinae, 220
Ancistrus cryptophthalmus, 220
Ancistrus cuiabae, 220
Ancistrus damasceni, 220
Ancistrus dolichopterus, 220
Ancistrus dubius, 221
Ancistrus erinaceus, 221
Ancistrus eustictus, 221
Ancistrus formoso, 221
Ancistrus fulvus, 221
Ancistrus füsslii, 235
Ancistrus galani, 221
Ancistrus gibbiceps, 291
Ancistrus guacharote, 225
Ancistrus guentheri, 287
Ancistrus gymnorhynchus, 221
Ancistrus heterorhynchus, 221
Ancistrus hoplogenys, 221
Ancistrus jataiensis, 222
Ancistrus jelskii, 222
Ancistrus latifrons, 222
Ancistrus leucostictus, 222
Ancistrus lineolatus, 222
Ancistrus lithurgicus, 222
Ancistrus lituratus, 290, 291
Ancistrus longimanus, 291
Ancistrus macrophthalmus, 222
Ancistrus maculatus, 222
Ancistrus malacops, 222
Ancistrus maracasae, 223
Ancistrus martini, 223
Ancistrus mattogrossensis, 300
Ancistrus mattogrossenssis, 300
Ancistrus medians, 242, 243
Ancistrus megalostomus, 223
Ancistrus melas, 220
Ancistrus minutus, 223

552 · Zootaxa 1418 © 2007 Magnolia Press

Ancistrus montanus, 223
Ancistrus multiradiatus alternans, 291
Ancistrus multispinis, 223
Ancistrus multispinis, 265
Ancistrus mystacinus, 265
Ancistrus nationi, 223
Ancistrus nudiceps, 223
Ancistrus occidentalis, 223
Ancistrus occloi, 223
Ancistrus parecis, 224
Ancistrus pictus, 234, 265
Ancistrus pirareta, 224
Ancistrus piriformis, 224
Ancistrus planiceps, 264
Ancistrus ranunculus, 224
Ancistrus reisi, 224
Ancistrus salgadae, 300
Ancistrus scaphirhynchus, 234
Ancistrus snethlageae, 283, 285
Ancistrus spinosus, 224
Ancistrus stigmaticus, 224
Ancistrus tamboensis, 224
Ancistrus taunayi, 224
Ancistrus temminckii, 224
Ancistrus tombador, 225
Ancistrus trinitatis, 225
Ancistrus triradiatus, 225
Ancistrus triradiatus martini, 223
Ancistrus variolus, 225
Ancistrus verecundus, 225
Ancistrus vittatus var. vermiculata, 285
Ancistrus vittatus vermiculata, 285
andamanensis, Arius, 56
andersoni, Liobagrus, 19
andersoni, Corymbophanes, 232
Andersonia, 23Andersonia brevior, 24Andersonia leptura, 24Andersonia pellegrini, 24
andersonii, Exostoma, 386
andersonii, Glaridoglanis, 386
Andinichthyidae, 30
Andinichthys, 30
Andinichthys bolivianensis, 30
Anduzedoras, 169
Anduzedoras arleoi, 169
Anduzedoras copei, 172
Anduzedoras oxyrhynchus, 169
anduzei, Ernstichthys, 60
anduzei, Trachelyopterichthys, 78
Aneistrus (Hemiancistrus) pulcher,234
Anemanotus, 37
anfractus, Clarias, 139
angeli, Pterodoras, 177angelica, Synodontis, 311
angelicus, Otolithus (Arius), 434
angelicus zonatus, Synodontis, 311
angelis, Ptedoras, 178

FERRARIS

angipinnatus, Hypostomus, 251
angipinnatus, Plecostomus, 251
angius, Pimelodus, 359
angolensis, Chiloglanis, 304
angolensis, Clarias, 139angolensis, Doumea, 25
angolensis, Eutropius, 363
angolensis, Schilbe, 363
angolensis macronema, Chlarias, 141angosturae, Farlowella, 240
anguilla, Ictalurus, 210
anguilla, Glanapteryx, 404
anguillaris, Clarias, 140, 144
anguillaris, Platystacus, 350, 351
anguillaris, Silurus, 138, 140, 143
anguillaris nigeriensis, Clarias, 140
anguillicauda, Liobagrus, 19
Anguilloclarias, 139
angulatus, Arius, 55
angulatus, Otolithus (Arius), 435
angulicauda, Plecostomus, 235
anguilicauda, Delturus, 234, 444
angusticeps, Adelopeltis, 107angustifrons, Anoplopterus, 27angustifrons, Tetracamphilius, 27angustirostre, Scleronema, 413
angustirostris, Pygidium, 413
anisitsi, Liposarcus, 291
anisitsi, Pterygoplichthys, 291
anisitsi, Homodiaetus, 406
anisura, Phenacorhamdia, 188
anisurus, Bagrus, 90
anisurus, Heptapterus, 188
anisurus, Pimelodus, 98
annae, Chaetostomus, 297
annae, Plecostomus, 297
annae, Squaliforma, 297
annandalei, Glyptothorax, 388
annectens, Acanthopoma, 403
annectens, Ancistrus, 242
annectens, Hemiancistrus, 242
annectens, Synodontis, 312
Anodontiglanis, 345
Anodontiglanis dahli, 345
anomala, Herklotsella, 375
anomala, Pterocryptis, 375
anomala sovichthys, Chaetostoma, 230
anomalopteryx, Parakysis, 15
anomalum, Chaetostoma, 227
anomalus, Chastostomus, 227
Anopleutropius, 165
Anopleutropius henrici, 165
Anoplopterus, 21
Anoplopterus angustifrons, 27
Anoplopterus longirostris, 22
Anoplopterus uranoscopus, 21, 23
anostomus, Silurus, 374
anoterus, Chiloglanis, 304
Ansorgia, 360

Ansorgia, vitata, 360
Ansorgia vittata, 360, 361
Ansorgia vittata bistriata, 361
ansorgii, Chrysichthys, 155ansorgii, Eutropius, 363
ansorgii, Parauchenoglanis, 154ansorgii, Phractura, 26ansorgii, Physalia, 360
ansorgii, Synodontis, 312
Ansorgiichthys, 360
anteanalis, Pareuchiloglanis, 398
anthrax, Lasiancistrus, 288
anthrax, Pseudolithoxus, 288
antiquus, Astephus, 207
antiquus, Pimelodus, 207
antoniensis, Pimelodus, 205
anus, Loricaria, 269
anus, Loricariichthys, 269
Anyperistius, 347
Anyperistius perugiae, 347
aor, Pimelodus, 106aor, Sperata, 106
aor sarwari, Aorichthys, 107
aorella, Sperata, 107
aorellus, Bagrus, 107
Aoria, 106
Aoria amemiyae, 91
Aoria amemiyai, 91
Aoria henryi, 100
Aoria hoi, 107
Aoria lacus, 108
Aoria rendahli, 108
Aoria virgatus, 101
Aorichthys, 106
Aorichthys aor sarwari, 107
aorides, Bagrus, 106
aorinus, Bagrus, 107
apangi, Amblyceps, 17
apeltogaster, Loricaria, 268
apeltogaster amazonum, Loricaria,
288
apeltogaster var. amazonum,
Loricaria, 288
Aphanotorulus, 225
Aphanotorulus ammophilus, 225
Aphanotorulus frankei, 225, 226
Aphanotorulus unicolor, 225
Apistoloricaria, 226
Apistoloricaria condei, 226
Apistoloricaria laani, 226
Apistoloricaria listrorhinos, 226
Apistoloricaria ommation, 226
apithanos, Pseudohemiodon, 288
Apodoglanis, 375
Apodoglanis furnessi, 375, 376
apogon, Phalacronotus, 374
apogon, Silurus, 374
Apomatoceros, 403
Apomatoceros alleni, 403

CHECKLIST OF CATFISHES, RECENT AND FOSSIL

Aposturisoma, 226
Aposturisoma myriodon, 226
approuaguensis, Corydoras, 113
Apuredoras, 177
Apuredoras rivasi, 177
apurensis, Cephalosilurus, 353
apurensis, Pseudopimelodus, 353
apurensis, Sachsdoras, 177, 178
apus, Channallabes, 137
apus, Gymnallabes, 137
aquilus, Lophiobagrus, 161arab, Plotosus, 351
arabi, Synodontis, 321
arafurensis, Arius, 49
araguaiaensis, Corydoras, 113
araguayensis, Galeichthys, 339
aralensis, Silurus glanis, 379
arcana, Cetopsis, 131
arcifer, Callichthys, 111
arcuatus, Corydoras, 113
arcuatus, Gogo, 29
arcuatus, Rhineastes, 207
areio, Corydoras, 113
arekaima, Pimelodus, 334
arenaria, Peckoltia, 284
arenarius, Arius, 34
arenarius, Bagrus (Ariodes), 34
arenarius, Hemiancistrus, 284
arenatus, Arius, 39
arenatus, Cathorops, 39
areolatus, Trichomycterus, 415
Argeini, 63
argentata, Silurus (Clupisoma), 357
argenteus, Arius, 56
argenteus, Neosilurus, 345
argenteus, Pimelodus, 53
argenteus, Plotosus, 345
argenteus, Porochilus, 345
argentina, Corydoras punctatus, 122argentina, Megalonema, 335
argentina, Perugia, 335
argentinus, Arius, 428
argentinus, Silurus, 209
argentivittatus, Macrones, 100
argentivittatus, Pelteobagrus, 100
Arges, 63
Arges boulengeri, 63
Arges brachycephalus, 63
Arges chotae, 63
Arges cirratus, 64
Arges eigenmanni, 64
Arges festae, 64
Arges fissidens, 64
Arges heterodon, 64
Arges homodon, 64
Arges longifilis, 65
Arges marmoratus, 65
Arges orientalis, 65
Arges peruanus, 66

Zootaxa 1418 © 2007 Magnolia Press · 553

Arges regani, 66
Arges retropinna, 66
Arges sabalo, 63, 67
Arges simonsii, 67
Arges stübeli, 67
Arges taczanowskii, 67
Arges theresiae, 67
Arges vaillanti, 67
Arges whymperi, 68
argus, Hypostomus, 251
argus, Plecostomus, 251
argyropleuron, Arius, 51
argyropleuron, Plicofollis, 51
argyrus, Pimelodus, 209
Arii, 30
Ariidae, 30
Ariodes, 34
Ariodes aeneus, 49
Ariodes macrocephalus, 51
Arius tonggol, 52
Arioida, 429, 432
arioides, Bagrus, 40, 54
Ariopsis, 32
Ariopsis, 428
Ariopsis aegyptiacus, 428
Ariopsis assimilis, 33
Ariopsis bonillai, 33
Ariopsis felis, 33
Ariopsis guatemalensis, 33
Ariopsis lentiginosa, 33
Ariopsis robertsi, 48
Ariopsis seemanni, 33
Ariopsis peyeria, 432
aripuanensis, Parotocinclus, 281
Aristommata, 249
Aristommata inexpectata, 249, 250
Aristotelis, Glanis, 378
aristotelis, Silurus, 378
aristotelis, Silurus (Parasilurus), 378
Arius, 30, 34
Arius (?) bartonensis, 56
Arius (Hemiarius) carinatus, 42
Arius (Hemiarius) danielsi, 42
Arius (Hemiarius) nudidens, 56
Arius acrocephalus, 47
Arius acutirostris, 31
Arius acutivelis, 155, 158
Arius acutus, 51
Arius aequibarbis, 55
Arius africana, 31
Arius alatus, 53
Arius albicans, 337
Arius albidus, 337
Arius andamanensis, 56
Arius angulatus, 55
Arius arafurensis, 49
Arius arenarius, 34
Arius arenatus, 39
Arius argenteus, 56

Arius argentinus, 428
Arius argyropleuron, 51
Arius arius, 34
Arius armiger, 46
Arius assimilis, 33, 434
Arius augustus, 46
Arius australis, 47
Arius baroni, 434
Arius Belangerii, 54
Arius bicolor, 34
Arius bleekeri, 46
Arius boakeii, 34
Arius bonneti, 36, 56
Arius borneënsis, 56
Arius Brandtii, 54
Arius brevirostris, 31
Arius brunellii, 31
Arius Buchanani, 34
Arius burmanicus, 42
Arius cacharioides, 166
Arius caelatoides, 55
Arius cælatus, 46
Arius Capellonis, 32
Arius cavatus, 435
Arius cærulescens, 33
Arius chondropterygioides, 55
Arius chondropterygius, 55
Arius clavispinosus, 36
Arius cleptolepis, 47
Arius clijpeaster, 55
Arius clypeastroides, 55
Arius coatesi, 47
Arius cochinchinensis, 34
Arius cookei, 50
Arius cous, 31
Arius crassus, 435
Arius crossocheilos, 52
Arius cruciger, 106
Arius curtisii, 47
Arius dasycephalus, 39
Arius dayi, 49
Arius despaxi, 36
Arius Dieperinki, 32
Arius digulensis, 47
Arius dioctes, 44
Arius dispar, 34
Arius doriae, 41
Arius dussumieri, 52
Arius Dutemplei, 56
Arius egertoni, 56
Arius egertoni belgicus, 56
Arius elatturus, 50
Arius equestris, 33
Arius falcarius, 34
Arius falcarius africana, 31
Arius falcarius var. africana, 31
Arius fangi, 34
Arius festae, 56
Arius festinus, 34

554 · Zootaxa 1418 © 2007 Magnolia Press

Arius fissus, 39
Arius Fraasi, 56
Arius froggatti, 42
Arius Fürthii, 39
Arius gagora, 34
Arius gagoroides, 35
Arius gigas, 31, 56
Arius goniaspis, 52
Arius graeffei, 47
Arius grandicassis, 49, 50
Arius grandoculis, 53
Arius granducolis, 53
Arius granosus, 54
Arius granulatus, 32
Arius guatemalensis, 33
Arius hainesi, 32
Arius hamiltonis, 51
Arius hardenbergi, 36
Arius hastatus, 105
Arius Heckelii, 55
Arius heudelotii, 31
Arius heward-belli, 56
Arius hypophthalmus, 39, 40
Arius iheringi, 343
Arius insculptus, 50
Arius insidiator, 45
Arius jella, 36
Arius kanganamanensis, 39
Arius Kessleri, 50
Arius kirkii, 52
Arius kitsoni, 56
Arius kutchensis, 56
Arius laeviceps, 55
Arius lagoensis, 31
Arius laticeps, 41
Arius latirostris, 47
Arius latiscutatus, 31
Arius layardi, 51
Arius leiotetocephalus, 52
Arius lemoinei, 432
Arius leptonotacanthus, 34
Arius longibarbis, 334
Arius luniscutis, 36
Arius macracanthus, 36
Arius macrocephalus, 51
Arius macronotacanthus, 47
Arius macruropterygius, 55
Arius maculatus, 34, 35
Arius madagascariensis, 35
Arius magatensis, 52
Arius malabaricus, 35
Arius manillensis, 35
Arius manjong, 55
Arius mastersi, 45
Arius melanochir, 41
Arius melanopterygius, 56
Arius melanopus, 40
Arius mercatoris, 31
Arius microcephalus, 35

FERRARIS

Arius microgastropterygius, 55
Arius micronotacanthus, 55
Arius micropterus, 166
Arius microstomus, 39
Arius micruropterygius, 36
Arius midgleyi, 47
Arius militaris, 51
Arius molliceps, 55
Arius multiradiatus, 40
Arius nasutus, 49
Arius neogranatensis, 50
Arius nigricans, 54
Arius nox, 38, 39
Arius nuchalis, 41
Arius nucleus, 57
Arius obesus, 81
Arius oetik, 35
Arius oncina, 75
Arius oncinus, 75
Arius osculus, 50
Arius papillosus, 165
Arius parkii, 32
Arius parmocassis, 50
Arius parvipinnis, 47
Arius paucus, 48
Arius pavimentatus, 105
Arius pectoralis, 48
Arius phrygiatus, 32
Arius physacanthus, 36
Arius pidada, 55
Arius planiceps, 50
Arius platypogon, 54
Arius platystomus, 52
Arius pleurops, 39
Arius polystaphylodon, 52
Arius proximus, 48, 49
Arius pumilus, 386
Arius puncticulatus, 41
Arius quadriscutis, 37
Arius ritoides, 106
Arius robertsi, 48
Arius rostratus, 55
Arius rugispinis, 32
Arius russi, 57
Arius rutschi, 435
Arius sagoroides, 45
Arius satparanus, 56
Arius schlegeli, 56
Arius sciurus, 35
Arius seemanni, 33
Arius serratus, 49
Arius similis, 434
Arius sinensis, 55
Arius solidus, 39
Arius spatula, 42, 43
Arius squalus, 166
Arius stauroforus, 32
Arius stirlingi, 46
Arius stricticassis, 50

Arius subrostratus, 35
Arius sumatranus, 36
Arius synodon, 166
Arius taylori, 40, 48
Arius tenuispinis, 52, 56
Arius tonggol, 52
Arius truncatus, 43
Arius tuyra, 40
Arius uncinatus, 36
Arius utarus, 48
Arius utik, 35
Arius vandeli, 50
Arius variolosus, 41
Arius venosus, 36
Arius verrucosus, 45
Arius villosus, 166
Arius viviparus, 55
arleoi, Anduzedoras, 169
arleoi, Pygidium, 415
arleoi, Trichomycterus, 415
armatulus, Doras, 176
armatulus, Platydoras, 176
armatus, Ageneiosus, 68, 69
armatus, Akysis, 98
armatus, Callichthys, 113
armatus, Corydoras, 113
armatus, Macrones, 94
armatus, Microsynodontis, 309
armatus, Mystus, 94, 98
armatus, Nanobagrus, 98
armatus, Silurus, 177
armbrusteri, Notarius, 440
armeniacum, Glyptosternum, 388
armeniacus, Glyptothorax, 388
armiger, Arius, 46
armiger, Mystus, 94
armiger, Nemapteryx, 46
armillatus, Pariolius, 188
arnoldi, Otocinclus, 271
arnoulti, Synodontis, 312
arnunachalensis, Amblyceps, 17
artedii, Platystoma, 341
ascita, Erethistoides, 384
ascita, Mystus, 177
ascita, Silurus, 54
asopos, Mastiglanis, 187
asotus, Parasilurus, 379
asotus, Silurus, 378, 381
asotus longus, Parasilurus, 379
asper, Callichthys, 110, 167
aspera, Rhinelepis, 293
asperatus, Hypostomus, 251
asperispinis, Lophiobagrus, 161
asperus, Pimelodus, 395
Aspidobagrus, 93
aspidolepis, Chaetostomus, 242
aspidolepis, Hemiancistrus, 242
Aspidoradidi, 108
Aspidoras, 108

CHECKLIST OF CATFISHES, RECENT AND FOSSIL

Aspidoras albater, 108
Aspidoras belenos, 108
Aspidoras brunneus, 109
Aspidoras carvalhoi, 109
Aspidoras depinnai, 109
Aspidoras eurycephalus, 109
Aspidoras fuscoguttatus, 109
Aspidoras lakoi, 109
Aspidoras maculosus, 109
Aspidoras menezesi, 109
Aspidoras microgalaeus, 109
Aspidoras pauciradiatus, 109
Aspidoras poecilus, 109
Aspidoras psammatides, 110
Aspidoras raimundi, 110
Aspidoras rochai, 110
Aspidoras spilotus, 110
Aspidoras taurus, 110
Aspidoras velites, 110
Aspidoras virgulatus, 110
aspilogaster, Hypostomus, 251
aspilogaster, Plecostomus, 251
Aspistor, 36
Aspistor hardenbergi, 36
Aspistor luniscutis, 36
Aspistor parkeri, 36
Aspistor quadriscutis, 37
Aspredinichthys, 57
Aspredinichthys filamentosus, 57
Aspredinichthys tibicen, 58
Aspredinidae, 57
aspredinoides, Myoglanis, 187
Aspredo, 57, 58
Aspredo aspredo, 58
Aspredo batrachus, 58
Aspredo filamentosus, 57
Aspredo gronovii, 58, 60
Aspredo sexcirrhis, 61
Aspredo sicuephorus, 58
Aspredo sicyephorus, 58
Aspredo spectrum, 61
Aspredo tibicen, 57, 58
Aspredo verrucosa, 60
aspredo, Aspredo, 58
aspredo, Silurus, 58
assamensis, Clarias, 148assamensis, Nangra, 396
assimilis, Ariopsis, 33
assimilis, Arius,33
astatodon, Atopochilus, 308
astatodon, Euchilichthys, 308
Astemomycterus, 411
Astephinae, 203
Astephus, 203, 207
Astephus antiquus, 207
Astephus resimus, 207
asterifrons, Astrodoras, 170
asterifrons, Doras, 169, 170
Asterophysi, 68

Zootaxa 1418 © 2007 Magnolia Press · 555

Asterophysus, 68, 70
Asterophysus batrachus, 70
Astroblepidae, 63
Astroblepiformes, 63
Astroblepus, 63
Astroblepus boulengeri, 63
Astroblepus brachycephalus, 63
Astroblepus caquetae, 63
Astroblepus chapmani, 63
Astroblepus chimborazoi, 63
Astroblepus chotae, 63
Astroblepus cirratus, 64
Astroblepus cyclopus, 64
Astroblepus cyclopus santanderensis,
67
Astroblepus eigenmanni, 64
Astroblepus festae, 64
Astroblepus fissidens, 64
Astroblepus formosus, 64
Astroblepus frenatus, 64
Astroblepus grixalvii, 64
Astroblepus grixalvii micrescens, 65
Astroblepus guentheri, 64
Astroblepus heterodon, 64
Astroblepus homodon, 64
Astroblepus jurubidae, 65
Astroblepus labialis, 65
Astroblepus latidens, 65
Astroblepus longiceps, 65
Astroblepus longifilis, 65
Astroblepus mancoi, 65
Astroblepus mariae, 65
Astroblepus marmoratus, 65
Astroblepus micrescens, 65
Astroblepus mindoensis, 65
Astroblepus nicefori, 65
Astroblepus nicéfori, 65
Astroblepus orientalis, 65
Astroblepus peruanus, 66
Astroblepus phelpsi, 66
Astroblepus pholeter, 66
Astroblepus pirrensis, 66
Astroblepus praeliorum, 66
Astroblepus prenadillus, 66
Astroblepus regani, 66
Astroblepus rengifoi, 66
Astroblepus retropinnus, 66
Astroblepus riberae, 66
Astroblepus rosei, 66
Astroblepus sabalo, 67
Astroblepus santanderensis, 67
Astroblepus simonsii, 67
Astroblepus stuebeli, 67
Astroblepus supramollis, 67
Astroblepus taczanowskii, 67
Astroblepus theresiae, 67
Astroblepus trifasciatus, 67
Astroblepus ubidiai, 67
Astroblepus unifasciatus, 67

Astroblepus vaillanti, 67
Astroblepus vanceae, 67
Astroblepus ventralis, 68
Astroblepus whymperi, 68
Astrodoras, 169
Astrodoras asterifrons, 170
Astrophysi, 68
Astrophysus, 70
asymetricaudalis, Chiloglanis, 304
atahualpa, Paracetopsis, 135
atavus, Silurus glanis, 382
ater, Cossyphus, 139, 148ater, Galeichthys, 43ater, Microglanis, 354
ater, Neosilurus, 347
ater sepikensis, Lambertichthys, 347
aterrima, Synodontis, 312
aterrimus, Synodontis, 312
atesuensis, Amphilius, 21
atherinoides, Neotropius, 359
atherinoides, Silurus, 359
atherinoides walkeri, Pseudeutropius,
359
athiensis, Chiloglanis, 304
athu, Silurus, 380
atochae, Pygidium, 423
Atopochilus, 303
Atopochilus astatodon, 308
Atopochilus chabanaudi, 303
Atopochilus christyi, 303
Atopochilus güntheri, 309
Atopochilus macrocephalus, 303
Atopochilus mandevillei, 304
Atopochilus pachychilus, 304
Atopochilus savorgnani, 304
Atopochilus vogti, 304
atra, Lambertia, 347
atrarius, Pimelodus, 205
atratoensis, Dolichancistrus, 235
atratoensis, Spatuloricaria, 296
atratoênsis, Pseudancistrus, 235
atribranchus, Dinotopterus, 148
atrifasciatus, Mystus, 94
atripes, Trachydoras, 179
atripinnis, Hemipimelodus, 34
atrizona, Hoplomyzon, 61
atrobrunneus, Pimelodus, 337
atronasus, Ageneiosus, 68
atropersonatus, Corydoras, 113
atropinnis, Hypostomus, 251
atroplumbeus, Tachysurus, 32
atrorus, Schilbeodes marginatus, 213attemsi, Allabenchelys, 137
attemsi, Clariallabes, 137attu, Silurus, 380
attu, Wallago, 380
attu, Wallagonia, 381
attu valeya, Wallago, 381
aubentoni, Mystus, 91

556 · Zootaxa 1418 © 2007 Magnolia Press

Auchenauglanis macrostom, 154
Auchenaspis, 154
Auchenipterichthys, 71
Auchenipterichthys coracoideus, 71
Auchenipterichthys dantei, 71
Auchenipterichthys longimanus, 71
Auchenipterichthys punctatus, 71
Auchenipterichthys thoracatus, 71
Auchenipteridae, 68
Auchenipterini, 68
Auchenipterus, 68, 71
Auchenipterus (Pseudauchenipterus)
affinis, 76
Auchenipterus (Pseudauchenipterus)
Jequitinhonhae, 76
Auchenipterus (Pseudauchenipterus)
striatulus, 80
Auchenipterus (Pseudepapterus)
hasemani, 76
Auchenipterus ambyiacus, 71
Auchenipterus brachyurus, 72
Auchenipterus brevibarbis, 80
Auchenipterus brevior, 72
Auchenipterus britskii, 72
Auchenipterus ceratophysus, 79
Auchenipterus demerarae, 72
Auchenipterus dentatus, 72
Auchenipterus fordicei, 72
Auchenipterus furcatus, 76
Auchenipterus glaber, 80
Auchenipterus Heckelii, 73
Auchenipterus immaculatus, 80
Auchenipterus insignis, 79
Auchenipterus isacanthus, 80
Auchenipterus lacustris, 79
Auchenipterus longimanus, 71
Auchenipterus maculosus, 79
Auchenipterus Magdalenae, 79
Auchenipterus menezesi, 72
Auchenipterus nigripinnis, 72
Auchenipterus nuchalis, 72
Auchenipterus obscurus, 81
Auchenipterus osteomystax, 72
Auchenipterus paysanduanus, 72
Auchenipterus punctatus, 71
Auchenipterus robustus, 80
Auchenipterus thoracatus, 71
Auchenipterus thoracicus, 71
Auchenipterus trachycorystes, 80, 81
Auchenoglanidinae, 153
Auchenoglanis, 153, 154
Auchenoglanis acuticeps, 154
Auchenoglanis ahli, 162
Auchenoglanis akiri, 154
Auchenoglanis altipinnis, 162
Auchenoglanis ballayi, 162
Auchenoglanis ballayi gravoti, 162
Auchenoglanis ballayi var. gravoti,
162

FERRARISAuchenoglanis biscutatus, 154
Auchenoglanis büttikoferi, 163
Auchenoglanis fasciatus, 163
Auchenoglanis grandis, 163
Auchenoglanis iturii, 163
Auchenoglanis longiceps, 163
Auchenoglanis macrostoma, 154
Auchenoglanis maculosus, 163
Auchenoglanis monkei, 163
Auchenoglanis ngamensis, 163
Auchenoglanis occidentalis, 154
Auchenoglanis occidentalis
tanganicanus, 154
Auchenoglanis occidentalis
tchadiensis, 154
Auchenoglanis occidentalis var.
tanganicanus, 154
Auchenoglanis occidentalis var.
tchadiensis, 154
Auchenoglanis pantherinus, 163
Auchenoglanis pietschmanni, 162
Auchenoglanis pulcher, 162
Auchenoglanis punctatus, 163
Auchenoglanis ubangensis, 162
Auchenoglanis Wittei, 154
Auchenopterus, 71
augierasi, Synodontis, 314
augusta, Nemapteryx, 46
augustus, Arius, 46
aulometopon, Netuma, 56
aulopygia, Tatia, 77
aulopygius, Centromochlus, 77
aurantiacus, Bagrus, 101
aurantiacus, Hypostomus, 278
aurantiacus, Parancistrus, 278
aurantiacus, Pseudobagrus, 101, 108
aurata, Hemiloricaria, 245
auratus, Bagrus, 155
auratus, Chrysichthys, 155, 158, 160
auratus, Pimelodus, 155
auratus, Porcus, 155
auratus, Schilbe, 364
aurea, Loricaria, 298
aureatus, Scobinancistrus, 296
aureum, Sturisoma, 298
auritus, Siluranodon, 366
auritus, Silurus, 366
aurofrenatus, Corydoras, 113
auroguttatus, Hypostomus, 251
auroguttatus, Trichomycterus, 415
australe, Corydoras, 60
australe, Dysichthys, 118
australis, Amiurus, 208
australis, Arius, 47
australis, Ictalurus, 208
australis, Pimelodella, 189
australis, Neosilurus, 347, 348
austriacus, Heterobranchus, 382
Austroglanididae, 81

Austroglanis, 81
Austroglanis barnardi, 81
Austroglanis gilli, 81
Austroglanis sclateri, 81
Autanadoras, 178
Autanadoras milesi, 178, 179
avanhandavae, Pimelodella, 189
axelrodi, Corydoras, 113
axillaris, Ageniosus, 69
Ayarnangra, 382
Ayarnangra estuarius, 382
aymarae, Rhamdella, 196
azureus, Galeichthys, 33
azygia, Farlowella, 238
azygolechis, Hemipsilichthys, 279
azygolechis, Pareiorhaphis, 279
Bbachi, Chaetostomus, 284
bachi, Oxydoras, 174bachi, Peckoltia, 284
Bachmannia, 428
Bachmannia chabutensis, 428
baculum, Encheloclarias, 149
badeli, Trachycorystes insignis, 80
baderi, Corydoras, 113
Bagarina, 382
Bagarius, 382, 383
Bagarius bagarius, 383
Bagarius Buchanani, 383
Bagarius gigas, 383
Bagarius lica, 383
Bagarius Nieuwenhuisii, 383
Bagarius rutilus, 383
Bagarius suchus, 383
Bagarius yarrelli, 383
bagarius, Bagarius, 383
bagarius, Pimelodus, 383
Bagre, 30, 37
Bagre bagre, 37, 97
Bagre marinus, 37
Bagre panamensis, 38
Bagre pinnimaculatus, 38
bagre, Bagre, 37, 97
bagre, Silurus, 37, 38
bagre, Stearopterus, 37
Bagreidae, 30
Bagrichthyes, 81
Bagrichthyoidei, 81
Bagrichthys, 81, 82
Bagrichthys hypselopterus, 82
Bagrichthys macracanthus, 82
Bagrichthys macropterus, 82
Bagrichthys majusculus, 83
Bagrichthys micranodus, 83
Bagrichthys obscurus, 83
Bagrichthys vaillantii, 83
Bagridae, 81
Bagroides, 82, 83
Bagroïdes, 83

CHECKLIST OF CATFISHES, RECENT AND FOSSIL

Bagroïdes macracanthus, 82
Bagroides macropterus, 82, 83
Bagroides melanopterus, 83
Bagroides melapterus, 83
Bagroïdes melapterus, 83
Bagroides Vaillantii, 83
Bagroidinae, 82
Bagropsis, 329
Bagropsis reinhardti, 329
Bagrus, 81, 83
Bagrus (Ariodes) arenarius, 34
Bagrus (Ariodes) Meyenii, 52
Bagrus (Bagrus) ramentosus, 108
Bagrus (Sciades) emphysetus, 53, 54
Bagrus (Sciades) pictus, 334
Bagrus abbreviatus, 95
Bagrus Adansonii, 365
Bagrus affinis, 85, 107
Bagrus agricolus, 107
Bagrus albicans, 54
Bagrus albilabris, 95
Bagrus anisurus, 90
Bagrus aorellus, 107
Bagrus aorides, 106
Bagrus aorinus, 107
Bagrus arioides, 40, 54
Bagrus aurantiacus, 101
Bagrus auratus, 155
Bagrus bajad, 83
Bagrus barbatus, 44
Bagrus bayad macropterus, 83
Bagrus bayad var. macropterus, 84
Bagrus bilineatus, 49
Bagrus Birmannus, 95
Bagrus bouderius, 165
Bagrus buchanani, 361
Bagrus caeruleus, 84
Bagrus capensis, 43
Bagrus capito, 155
Bagrus carchariorhijnchos, 49
Bagrus chinta, 431
Bagrus corsula, 89
Bagrus couma, 53
Bagrus crinalis, 55
Bagrus degeni, 84
Bagrus depressirostris, 364
Bagrus docmac niger, 84
Bagrus docmak, 84, 313
Bagrus doroides, 45
Bagrus exodon, 359
Bagrus filamentosus, 84
Bagrus flavicans, 344
Bagrus flavus, 90
Bagrus fuscus, 95
Bagrus gagorides, 35
Bagrus genidens, 43, 44
Bagrus goliath, 331
Bagrus gulioides, 95
Bagrus Halepensis, 93, 94, 96, 97

Zootaxa 1418 © 2007 Magnolia Press · 557

Bagrus heterurus, 97
Bagrus hoevenii, 88
Bagrus hypselopterus, 82
Bagrus Javensis, 45
Bagrus Keletius, 98
Bagrus koenigi, 84
Bagrus laevigatus, 49
Bagrus Lamarrii, 106, 107
Bagrus leucophasis, 96
Bagrus limbatus, 107
Bagrus lonah, 391
Bagrus lubosicus, 84
Bagrus macronemus, 37, 97
Bagrus macronemus, 93, 97
Bagrus Malabaricus, 96
Bagrus maurus, 158
Bagrus melas, 95
Bagrus meridionalis, 84
Bagrus mesops, 53
Bagrus micracanthus, 96
Bagrus micropogon, 92, 93
Bagrus montanus, 96
Bagrus nemurus, 87, 90
Bagrus netuma, 48, 49
Bagrus nigriceps, 96
Bagrus nigrita, 160
Bagrus nigropunctatus, 340
Bagrus oculatus, 96
Bagrus orientalis, 84
Bagrus passany, 54
Bagrus pemecus, 53
Bagrus planiceps, 90
Bagrus poecilopterus, 93
Bagrus proops, 54
Bagrus punctatus, 91
Bagrus punctulatus, 331, 339, 340
Bagrus reticulatus, 330, 331
Bagrus rhodonotus, 49
Bagrus rhodopterygius, 95
Bagrus rousseauxii, 331
Bagrus schilbeides, 362, 365
Bagrus Schlegelii, 36, 95
Bagrus shuwaiensis, 84
Bagrus Sieboldii, 90
Bagrus sinensis, 108
Bagrus singaringan, 97
Bagrus sondaicus, 45
Bagrus stenomus, 103, 104
Bagrus sumatranus, 36
Bagrus temminckianus, 55
Bagrus Temminckii, 55
Bagrus thalassinus, 49
Bagrus trachacanthus, 89
Bagrus trachipomus, 55
Bagrus ubangensis, 84
Bagrus urostigma, 84
Bagrus ussuriensis, 92, 100
Bagrus vachellii, 101
Bagrus venaticus, 55

Bagrus vertagus, 55
Bagrus wolffii, 98
Bagrus Wyckii, 91
Bagrus Yarrelli, 383
bahianus, Corymbophanes, 279
bahianus, Pareiorhaphis, 279
bahianus, Pimelodus, 195
bahianus, Trichomycterus, 415
bahiensis, Galeichthys, 38bahiensis, Kalyptodoras, 172
bahiensis, Microlepidogaster, 281
bahiensis, Parotocinclus, 281
bahuaja, Crossoloricaria, 232
baileyi, Noturus, 211
bajad, Bagrus, 83
bajad, Silurus, 83
balayi, Parauchenoglanis, 162
balayi, Pimelodus, 162
ballayi, Auchenoglanis, 162
ballayi gravoti, Auchenoglanis, 162
ballayi var. gravoti, Auchenoglanis,
162
balsanus, Ictalurus, 208
balsanus, Istlarius, 208
balsanus occidentalis, Istlarius, 208
Balzanii, Vandellia, 427
bambui, Ituglanis, 406
banguela, Homodiaetus, 406
banguelensis, Eutropius, 363
banguelensis, Schilbe, 363
bankae, Chaca, 135
bankanensis, Chaca, 135banneaui, Pygidium, 415
banneaui, Trichomycterus, 415
banneaui maracaiboensis, Pygidium,
420
barakensis, Pterocryptis, 441
barakensis, Sisor, 402
baramensis, Akysis, 16
baramensis, Hemibagrus, 87
baramensis, Kryptopterus, 369
baramensis, Liocassis, 93
baramensis, Macrones, 87
baramensis, Pseudobagarius, 16barbancho, Pimelodus, 339
barbarmatus, Dentectus, 235
barbata, Loricaria, 298, 299
barbata, Microcambeva, 409
barbata, Rhamdia, 201
barbatula, Trichomycterus, 422
barbatum, Sturisoma, 299
barbatus, Bagrus, 44barbatus, Callichthys, 129
barbatus, Hypostomus, 286, 287
barbatus, Malapterurus, 301
barbatus, Platysilurus, 340
barbatus, Pseudancistrus, 287
barbatus, Scleromystax, 129
barbatus, Xyliphius, 62558 · Zootaxa 1418 © 2007 Magnolia Press

barbonica, Laimumena, 431, 432
barbouri, Pygidium, 415
barbouri, Trichomycterus, 415
barbus, Genidens, 44
barbus, Pimelodus, 44
barnardi, Austroglanis, 81
barnardi, Gephyroglanis, 81baroni, Arius, 434
Baronis Mülleri, Pimelodus(Rhamdia), 201
barrae, Ancistrus, 272
barrae, Megalancistrus, 272
barranquerensis, Ageneiosus, 69
bartonensis, Arius bartonensis, 56
bartonensis, Otolithus (Arius) crassus,
434
bartonensis, Otolithus (Arius) danicus,
434
bartoni, Neosilurus, 347
Baryancistrus, 226
Baryancistrus demantoides, 226
Baryancistrus longipinnis, 227
Baryancistrus niveatus, 227
basilewskii, Silurichthys, 381
bastari, Clupisoma, 357
bastiani, Synodontis, 312
batarensis, Pseudeutropius murius,
357
Batasinae, 81
Batasio, 81, 84
Batasio affinis, 85
Batasio batasio, 85, 86
Batasio buchanani, 84, 85
Batasio dayi, 85
Batasio elongatus, 85
Batasio fasciolatus, 442
Batasio havmolleri, 85
Batasio macronotus, 85
Batasio merianiensis, 85
Batasio niger, 445
Batasio pakistanicus, 85
Batasio sharavatiensis, 86
Batasio spilurus, 442
Batasio tengana, 86, 442
Batasio tigrinus, 86
Batasio travancoria, 86
batasio, Batasio, 85, 86
batasio, Pimelodus, 85
batasius, Pimelodus, 85, 98
batensoda, Brachysynodontis, 304
batensoda, Synodontis, 304
Batesi longispinis, Synodontis, 316
Batesi var. longispinis, Synodontis, 316
batesii, Chiloglanis, 304
batesii, Microsynodontis, 309
batesii, Synodontis, 312
Bathophilus, 412
Bathybagrus, 154
Bathybagrus tetranema, 154, 155

FERRARIS

Bathycetopsis, 131
Bathycetopsis oliveirai, 131, 132
Bathyclarias, 136, 148
Bathyclarias euryodon, 136
Bathyclarias filicibarbis, 148
Bathyclarias gigas, 149
Bathyclarias ilesi, 136
Bathyclarias longibarbis, 136
Bathyclarias loweae, 149
Bathyclarias rotundifrons, 136
Bathyclarias worthingtoni, 136
Bathypygidium, 412
bathyurus, Pimelodus, 201
Batrachocephalinae, 30
Batrachocephalus, 30, 38
Batrachocephalus ageneiosus, 38
Batrachocephalus micropogon, 38
Batrachocephalus mino, 38
Batrachoglanis, 352
batrachostoma, Gyrinurus, 409
batrachostoma, Ochmacanthus, 409
batrachus, Aspredo, 58
batrachus, Asterophysus, 70
batrachus, Clarias, 140
batrachus, Silurus, 140
Batrochoglanis, 352
Batrochoglanis acanthochiroides, 352
Batrochoglanis melanurus, 353
Batrochoglanis raninus, 353
Batrochoglanis transmontanus, 353
Batrochoglanis villosus, 353
batu, Clarias, 140
baudensis, Ancistrus, 220baudoensis, Cetopsis, 131
baudoênsis, Pseudocetopsis, 131
baudoni, Amphilius, 21
baudoni uniformis, Amphilius, 21
baudoni var. uniformis, Amphilius, 21
bayad, Porcus, 83
bayad macropterus, Porcus, 83
bayad var. macropterus, Porcus, 84
Beadlei, Synechoglanis, 207, 210
beani, Pangasius, 328
beccarii, Vandellia, 427
bedado, Pangasius, 325
bedfordi, Silurus, 379
Belangerii, Arius, 54
beldti, Cryptopterella, 369, 371
belenos, Aspidoras, 108
belensis, Trichomycterus, 415
belgicus, Arius egertoni, 56
Belodontichthys, 368
Belodontichthys dinema, 368
Belodontichthys javanensis, 381
Belodontichthys macrochir, 368
Belodontichthys truncatus, 368
Belonoglanis, 24
Belonoglanis brieni, 24
Belonoglanis curvirostris, 24

Belonoglanis nudipectus, 24
Belonoglanis tenuis, 24
benderensis, Ictalurus, 206
benedettii, Nannorhamdia, 198
beninensis, Malapterurus, 301
bengalensis, Malapterus (Ailia), 356
beni, Hemiloricaria, 245
beni, Lepthoplosternum, 128beni, Loricaria, 245
benjamini, Entomocorus, 74
benuensis, Chiloglanis, 305
berdmorei, Exostoma, 385
berdmorei, Pterocryptis, 376, 377
Berdmorei, Silurichthys, 376
Bergiaria, 329
Bergiaria platana, 330
Bergiaria westermanni, 330
Bergiella, 329
Bergiella platana, 330
berneyi, Neoarius, 47
berneyi, Tachysurus (Pararius), 47
bernhardi, Hemipimelodus, 39
bertoni, Branchioica, 410
bertoni, Corydoras, 130
bertonii, Branchioica, 410
bicarinatus, Chaenothorax, 111, 125bicirrhis, Kryptopterus, 369
bicirrhis, Silurus, 369
bicirrosus, Plecostomus, 259
bicolor, Arius, 34
bicolor, Bunocephalus, 59
bicolor, Corydoras, 114
bicolor, Hemipimelodus, 34
bicolor, Leiocassis, 104
bicolor, Phractocephalus, 336
bicolor, Pirarara, 336
bidentatus, Parotocinclus, 282
bidorsalis, Clarotes, 160
bidorsalis, Heterobranchus, 150
bifasciata, Taunayia, 203
bifasciatus, Corydoras, 114
bifasciatus, Nannoglanis, 203
biffi, Notarius, 49bifidus, Bunocephalus, 59bifurcus, Chiloglanis, 305
bilineata, Netuma, 49
bilineatus, Bagrus, 49
bilineatus, Corydoras, 114
bilineatus, Epactionotus, 236
bilineatus, Pimelodus, 107bilobatum, Hypoptopoma, 250
bimaculatus, Macrones, 94
bimaculatus, Mystus, 94
bimaculatus, Ompok, 372
bimaculatus, Silurus, 372
binotata, Synodontis notatus, 318
binotatus, Ompok, 372
Birmannus, Bagrus, 95
biscutatus, Auchenoglanis, 154

CHECKLIST OF CATFISHES, RECENT AND FOSSIL

biscutatus, Pimelodus, 154biseriata, Squaliforma, 297
biseriatus, Plecostomus, 297
bistriata, Ansorgia vittata, 361
biwaensis, Parasilurus, 379
biwaensis, Silurus, 379
bleekeri, Arius, 46
bleekeri, Cryptopterus, 374
bleekeri, Heptapterus, 184
bleekeri, Macrones, 89, 94
bleekeri, Micronema, 374
bleekeri, Mystus, 94
bleekeri, Nemapteryx, 46
bleekeri, Paracetopsis, 134
bleekeri, Phalacronotus, 374
blochi, Corydoras, 114
blochi vittatus, Corydoras, 126
Blochii, Cotylephorus, 61
blochii, Doras, 167
Blochii, Galeichthys, 38
Blochii, Osteogeneiosus, 51
blochii, Pimelodus, 336, 337
blohmi, Epapterus, 74
Blythii, Exostoma, 396
Blythii, Macrones, 85
blythii, Myersglanis, 396bo, Macrones, 88
boakeii, Arius, 34
boalis, Silurus, 380
bocagii, Chrysichthys, 155
Bocagii, Eutropius, 363
bocagii, Schilbe, 363
bockmanni, Glanidium, 75
bocourti, Heterobagrus, 94
bocourti, Mystus, 94
bocourti, Pangasius, 325
Bocourti, Pangasius(Pseudopangasius), 325
bodenhameri, Ancistrus, 219
boehlkei, Corydoras, 114
boehlkei, Rhinodoras, 178boemia, Tatia, 77
boesemani, Corydoras, 114
bogotense, Pygidium, 415
bogotensis, Trichomycterus, 415
bokorensis, Penesilurus, 376
bokorensis, Pterocryptis, 376
bolivarensis, Amblydoras, 168
bolivarensis, Hildadoras, 168
boliviana, Farlowella, 238
boliviana, Phenacorhamdia, 188boliviana, Pimelodella, 189boliviana, Xenocara, 219
bolivianensis, Andinichthys, 30
bolivianus, Ancistrus, 219
bolivianus, Corydoras, 117bolivianus, Hypostomus, 252bolivianus, Imparfinis, 188bolivianus, Plecostomus, 252

Zootaxa 1418 © 2007 Magnolia Press · 559

bolli, Amiurus, 205bolteni, Callichthys callichthys, 111
bomae, Eutropius, 363
bomboides, Pseudomystus, 103
bomboizanum, Pygidium, 416
bomboizanus, Trichomycterus, 416
bondi, Acanthopoma, 413
bondi, Corydoras, 114
bondi, Schultzichthys, 413
bondi coppenamensis, Corydoras,115
bongan, Hemibagrus, 87
bongan, Macrones, 87bonillai, Ariopsis, 33
bonillai, Galeichthys, 33
bonneti, Arius, 36, 56
boquillae, Cetopsorhamdia, 181
borealis, Silurus (Pimelodus), 210
borellii, Hypostomus, 252borellii, Plecostomus, 252borellii, Trichomycterus, 416
borneensis, Callichrous (Silurodes),372
borneensis, Cephalocassis, 41, 42
borneensis, Ompok, 372
borneënsis, Arius, 56
borneënsis, Pimelodus, 41borodini, Imparfinis, 185bororo, Otocinclus, 274
börressoni, Chrysichthys, 156bortayro, Silvinichthys, 413
boschmai, Pimelodella, 190
bostocki, Tandanus, 352
bostockii, Cnidoglanis, 346
boticario, Listrura, 408
botius, Glyptothorax, 388
botius, Pimelodus, 388
Boucardi, Pimelodus, 201
bouderius, Bagrus, 165
bouderius, Cranoglanis, 165
bouilloni, Leptoglanis, 164
boujardi, Lithoxus, 267
boulengeri, Arges, 63,boulengeri, Astroblepus, 63
boulengeri, Euchilichthys, 309
boulengeri, Hemidoras, 175
boulengeri, Hemidoras (Leptodoras),175
boulengeri, Heterobranchus, 151boulengeri, Hypostomus, 252boulengeri, Opsodoras, 172, 175
boulengeri, Plecostomus, 252bourguyi, Microlepidogaster, 300
boutchangai, Anaspidoglanis, 154
boutchangai, Parauchenoglanis, 154
bouvieri, Schilbe, 364
bovallii, Ancistrus, 267
bovallii, Lithoxus, 267
bovei, Peltura, 26
bovei, Phractura, 26

boylei, Pygidium, 416
boylei, Trichomycterus, 416
brachiatus, Centrodoras, 170
brachiatus, Doras, 170
Brachspondylus cretaceus, 432
brachyacanthus, Amiurus, 205
brachybarbatus, Akysis, 13brachycephalus, Arges, 63
brachycephalus, Astroblepus, 63
brachycephalus, Pimelodus, 198
Brachyglanis, 180
Brachyglanis frenata, 180
Brachyglanis frenatus, 180
Brachyglanis magoi, 180
Brachyglanis melas, 181
Brachyglanis microphthalmus, 181
Brachyglanis nocturnus, 181
Brachyglanis phalacra, 181
Brachymystus, 87brachynema, Chrysichthys, 155
brachynema, Chasmocranus, 182
brachynemus, Chasmocranus, 182
Brachyplatystoma, 329, 330
Brachyplatystoma capapretum, 330
Brachyplatystoma filamentosum, 330
Brachyplatystoma goeldii, 330
Brachyplatystoma juruense, 331
Brachyplatystoma paraense, 331
Brachyplatystoma parnahybae, 331
Brachyplatystoma platynema, 331
Brachyplatystoma platynemum, 331
Brachyplatystoma promagdalena, 331
Brachyplatystoma rousseauxii, 331
Brachyplatystoma tigrinum, 331
Brachyplatystoma vaillantii, 331
Brachyplatystomatini, 329
brachypopterus, Eutropius, 362
brachypopterus, Pseudeutropius, 362
brachypterus, Pimelodus (Rhamdia),
198
Brachyrhamdia, 181
Brachyrhamdia heteropleura, 181
Brachyrhamdia imitator, 181
Brachyrhamdia marthae, 181
Brachyrhamdia meesi, 181
Brachyrhamdia rambarrani, 181
brachyrhyncha, Pareiorhina, 281
brachysoma, Clarias, 140
brachysoma, Horabagrus, 429
brachysoma, Pseudobagrus, 429
brachysoma, Pseudexostoma, 401
Brachyspondylus, 432
Brachyspondylus indicus, 326
Brachyspondylus saropterix, 82
Brachystacus, 135
Brachysynodontis, 304
Brachysynodontis batensoda, 304
brachyura, Dekeyseria, 234
brachyurus, Ancistrus, 234

560 · Zootaxa 1418 © 2007 Magnolia Pressbrachyurus, Auchenipterus, 72
brachyurus, Euanemus, 72
Branchioica, 410
Branchioica bertoni, 410
Branchioica bertonii, 410
Branchioica magdalenae, 411
Branchioica phaneronema, 411
Branchiosteus, 17
Brandtii, Arius, 54
branickii, Chaetostoma, 228
branickii, Chaetostomus, 228
branneri, Rhamdia, 201
branneri voulezi, Rhamdia, 201
bransfordi, Loricaria, 240
bransfordii, Rhamdia, 201
brashnikowi, Leiocassis, 100
brashnikowi, Macrones (Leiocassis),100
brashnikowi, Pelteobagrus, 100
brasiliensis, Caecorhamdella, 189, 190
brasiliensis, Pimelodella, 190, 192
brasiliensis, Pimelodus(Pseudorhamdia), 192
brasiliensis, Plecostomus, 259
brasiliensis, Trichomycterus, 416
brasiliensis itatiayae, Trichomycterus,419
brasiliensis tristis, Trichomycterus,
416
braueri, Hemiancistrus, 284
braueri, Peckoltia, 284
breei, Corydoras, 114
Breitensteinia, 15
Breitensteinia cessator, 15
Breitensteinia hypselurus, 15
Breitensteinia insignis, 15
breve, Chaetostoma, 228
brevianalis, Eutropius, 363
brevianalis, Pseudobagrus, 101
brevianalis, Schilbe, 363
brevibarbis, Ancharius, 29
brevibarbis, Auchenipterus, 80
brevibarbis, Chiloglanis, 305
brevibarbis, Chrysichthys, 156
brevibarbis, Chrysobagrus, 155, 156
brevibarbis, Clariallabes, 137
brevibarbis, Gogo, 29
brevibarbus, Plotosus, 351
brevicauda, Hypostomus, 252
brevicauda, Phractura, 26
brevicauda, Plecostomus, 252
brevicaudatus, Leiocassis, 102
brevicaudatus, Pseudobagrus, 102
Brevicephaloides, 139
Breviceps, 37
breviceps, Clarias, 141
breviceps, Liocassis, 103
breviceps, Pimelodella, 190
breviceps, Pimelodus, 190

FERRARIS

breviceps, Pseudomystus, 103
brevicorpus, Coreobagrus, 86
brevidorsalis, Amphilius, 23
brevidorsalis, Copidoglanis, 347
brevidorsalis, Neosilurus, 347
brevifilis, Ageneiosus, 68, 69
brevifilis, Ancistrus, 219
brevifilis bodenhameri, Ancistrus, 219
Breviglanis, 180
brevilabiatum, Chaetostoma, 228
brevilabiatus, Chaetostomus, 228
brevinuchalis, Clarias, 140
brevior, Allabenchelys, 137
brevior, Andersonia, 24
brevior, Auchenipterus, 72
brevior, Chasmocranus, 182
brevipinna, Amphilius
grammatophorus, 22
brevipinna, Amphilius
grammatophorus var., 23
brevipinnis, Ancistrus, 219
brevipinnis, Glyptothorax, 388
brevipinnis, Xenocara, 219
brevipinnis alaknandi, Glyptothorax,
388
brevirostre, Sturisoma, 299
brevirostris, Arius, 31
brevirostris, Corydoras, 114
brevirostris, Loricaria, 299
brevirostris, Loricaria (Rineloricaria),
298
brevis, Ageneiosus, 69
brevis, Amphilius, 21
brevis, Chaetostomus, 228
brevis, Delturus, 443
brevis, Doras (Corydoras), 179
brevis, Hemiancistrus, 284
brevis, Hypostomus, 252
brevis, Leptoglanis, 28
brevis, Mochocus, 310
brevis, Mochokus, 310
brevis, Peckoltia, 284
brevis, Pimelodus, 337
brevis, Plecostomus, 252
brevis, Trachydoras, 179
brevis, Tridens, 426
brevis, Tridensimilis, 426
brevis, Zaireichthys, 28
brevispinis, Lasiancistrus, 287
brevispinis, Lophiobagrus, 161
brevispinis, Pseudancistrus, 287
brevitentaculatus, Hypostomus, 291
brichardi, Phyllonemus, 164
brichardi, Synodontis, 312
brieni, Belonoglanis, 24britskii, Auchenipterus, 72
britskii, Brochis, 114
britskii, Corydoras, 114
britskii, Parotocinclus, 282

broadbenti, Tachysurus, 51
Brochiloricaria, 227
Brochiloricaria chauliodon, 227
Brochiloricaria macrodon, 227
Brochis, 111
Brochis britskii, 114
Brochis coeruleus, 111, 125
Brochis dipterus, 111, 125
Brochis multiradiatus, 120
Brochis splendens, 125
Brontes, 63
Brontes prenadilla, 63, 66
brunellii, Arius, 31
brunnea, Loricaria, 269
brunnea, Tatia, 77
brunnescens, Doras, 168
brunneus, Ameiurus, 204
brunneus, Amiurus, 204
brunneus, Aspidoras, 109
brunneus, Loricariichthys, 269
Brustiarius, 38
Brustiarius nox, 38
Brustiarius solidus, 39
buccata, Pterocryptis, 376
bucculenta, Nangra, 396
Buchanani, Arius, 34
Buchanani, Bagarius, 383
buchanani, Bagrus, 361
Buchanani, Batasio, 84, 85
buchanani, Chaca, 136buchanani, Glyptothorax, 388
buchanani, Hara, 395
Buchanani, Nangra, 397
Buchanani, Pangasius, 325, 327
buchanani, Proeutropiichthys, 361
buchanani, Rama, 105
Buchanani, Rita, 105, 106
Bucklandium, 432
Bucklandium diluvii, 432, 433
buckleyi, Pimelodella, 190
buckleyi, Pimelodus, 190
budgetti, Clarias, 140
budgetti, Synodontis, 312
buettikoferi, Clarias, 140
buettikoferi, Parauchenoglanis, 163
buffei, Eutropiellus, 361
buffei, Pareutropius 360
bufonia, Xenocara, 219
bufonius, Ancistrus, 219
bufonius, Pimelodus, 355
bufonius, Pseudopimelodus, 355
bufonius, Xenocara, 219
bullocki, Hatcheria, 404
Bullockia, 403
Bullockia maldonadoi, 404
bulumae, Clarias, 140
Bunocephalichthys, 58
Bunocephalichthys gronovii, 60
Bunocephalini, 57

CHECKLIST OF CATFISHES, RECENT AND FOSSIL

Bunocephalus, 57, 58
Bunocephalus albifasciatus, 62
Bunocephalus aleuropsis, 58
Bunocephalus amaurus, 58
Bunocephalus amaurus aloikae, 58
Bunocephalus amaurus sipaliwini, 58
Bunocephalus amazonicus, 59
Bunocephalus bicolor, 59
Bunocephalus bifidus, 59
Bunocephalus carvalhoi, 59
Bunocephalus chamaizelus, 59
Bunocephalus colombianus, 59
Bunocephalus coracoideus, 59
Bunocephalus depressus, 62
Bunocephalus dolichurus, 62
Bunocephalus doriae, 59
Bunocephalus haggini, 59
Bunocephalus hypsiurus, 57
Bunocephalus iheringii, 59
Bunocephalus knerii, 59
Bunocephalus larai, 59
Bunocephalus melas, 58
Bunocephalus minutus, 59
Bunocephalus quadriradiatus, 60
Bunocephalus retropinnis, 59
Bunocephalus rugosus, 60
Bunocephalus salathei, 59
Bunocephalus scabriceps, 58, 60
Bunocephalus verrucosus, 60burgessi, Corydoras, 114
burgini, Pangasius, 326
burmanensis, Pterocryptis, 376
burmanensis, Silurus, 376
burmanica, Cochlefelis, 42burmanica, Olyra, 99
burmanicus, Arius, 42burmanicus, Glyptothorax, 388
burmanicus, Proeutropiichthys
taakree, 362
burmannicus, Eutropiichthys, 358
Burmeisteri, Pygidium, 405
burmensis, Chaca, 135butantanis, Plecostomus margaritifer,
256
butcheri, Pseudopimelodus villosus,
352
buthupogon, Clarias, 141butleri, Paraplotosus, 349
Büttikoferi, Acrochordonichthys, 13
Büttikoferi, Auchenoglanis, 163
Büttikoferi, Clarias, 141
Büttikoferi, Chrysichthys, 158
Ccabrerae, Rhamdia, 198
cacerensis, Hemiloricaria, 245
cacerensis, Loricaria, 245
cacharioides, Arius, 166
cadeae, Loricaria, 294
cadeae, Rineloricaria, 293

Zootaxa 1418 © 2007 Magnolia Press · 561

Caecorhamdella, 189
Caecorhamdella brasiliensis, 189, 192
Caecorhamdia, 197
Caecorhamdia urichi, 197, 202
caecutiens, Amblyceps, 17, 18
Caelatoglanis, 382, 383
Caelatoglanis zonatus, 383
caelata, Nemapteryx, 46cælatus, Arius, 46
caelatoides, Arius, 55
caerulescens, Pimelodus, 81
caeruleus, Bagrus, 84
caerulorostris, Platypogon, 339, 340
caesius, Plotosus, 350
caesius, Propimelodus, 443
cahuali, Tridentopsis, 426
cahyensis, Ituglanis, 444
caiapo, Cetopsis, 131
Cainosilurus, 347
calamita, Hypostomus, 219
Calichthys, 110caliense, Pygidium, 416
caliensis, Trichomycterus, 416
callarias, Silurus, 313
Callichrous, 371
Callichrous (Silurodes) borneensis,372
Callichrous ceylonensis, 372
Callichrous egertonii, 374
Callichrous eugeneiatus, 369
Callichrous nigrescens, 374
Callichrous notatus, 374
Callichrous Sindensis, 373
Callichrous weberi, 373
Callichthini, 108
Callichthyidae, 108
Callichthys, 108, 110
Callichthys adspersus, 127
Callichthys affinis, 111
Callichthys albidus, 128
Callichthys arcifer, 111
Callichthys armatus, 113
Callichthys asper, 110, 167
Callichthys barbatus, 129
Callichthys callichthys, 110
Callichthys callichthys bolteni, 111
Callichthys callichthys demararae, 111
Callichthys chiquitos, 128
Callichthys exaratus, 129
Callichthys fabricioi, 111
Callichthys hemiphractus, 111
Callichthys kneri, 111
Callichthys laevigatus, 127
Callichthys littoralis, 127
Callichthys longifilis, 129
Callichthys loricatus, 111
Callichthys melampterus, 127, 128
Callichthys oibaensis, 111
Callichthys paleatus, 122

Callichthys pectoralis, 128
Callichthys pictus, 129
Callichthys serralabium, 111
Callichthys splendens, 125
Callichthys sulcatus, 129
Callichthys taiosh, 125
Callichthys tamoata, 111
Callichthys thoracatus, 129
callichthys, Callichthys, 110callichthys bolteni, Callichthys, 111callichthys demararae, Callichthys,111
Callomystax, 385
Callomystax Schmidti, 392
callopterus, Glyptothorax, 388
calmoni, Hemipsilichthys, 279
Calophysinae, 329
Calophysus, 329, 332
Calophysus macropterus, 332
calvarius, Silurus, 99, 100
calvus, Isbrueckerichthys, 441
calvus, Rhineastes, 207
camaronensis, Chrysichthys, 158
camelopardalis, Synodontis, 312
cameronensis, Chiloglanis, 305
cameronensis, Chrysichthys, 158
cameronensis, Clarias, 144
cameronensis, Eutropius, 365
cameroni, Hemipsilichthys, 279
cameroni, Pareiorhaphis, 279
cameroni, Psilichthys, 247, 279
camerunensis, Clarias, 139, 141, 144
camerunensis, Leptoglanis, 28
camerunensis, Zaireichthys, 28
camposensis, Diplomystes, 165
camposi, Eremophilus, 408
camposi, Listrura, 408
canaliferus, Otothyris, 275, 276
canarensis, Mystus, 96
cancellatus, Ameiurus, 211
candidus, Eremophilus, 416
candidus, Trichomycterus, 416
candiru, Cetopsis, 131
candombe, Hisonotus, 440
canensis, Acanthicus, 266
canensis, Leptoancistrus, 266
canio, Silurus, 373, 374
canius, Plotosus, 350
Canthopomus, 289
Canthopomus montebelloi, 263
Cantonensis, Pimelodus, 431
cantoris, Osteogeneiosus, 51
capapretum, Brachyplatystoma, 330
caparary, Sorubim, 341
Capellonis, Arius, 32
capensis, Bagrus, 43
capensis, Clarias, 143
capetensis, Dasyloricaria, 233
capetensis, Loricaria, 233

562 · Zootaxa 1418 © 2007 Magnolia Press

capito, Bagrus, 155
capitulum, Macrones fortis, 88
caquetae, Astroblepus, 63
caquetae, Corydoras, 119
caquetae, Euacanthagenys, 296
caquetae, Harttia, 300
caquetae, Hemiancistrus, 265
caquetae, Loricaria, 296
caquetae, Spatuloricaria, 296
caquetae, Sturisomatichthys, 300
caracasensis, Hemiloricaria, 244, 245
caracassensis, Hemiloricaria, 245
carachama, Monistiancistrus, 289
carcharhinoides, Nangra, 396
carchariorhijnchos, Bagrus, 49
carcio, Pimelodus, 85, 98
carinata, Farlowella, 237
carinata, Loricaria, 268
carinata, Oxyropsis, 276
carinata, Trewavasia, 433
carinatum, Amblyceps, 18
carinatum, Hypoptopoma, 276
carinatus, Arius (Hemiarius), 42
carinatus, Cinetodus, 42
carinatus, Doras, 171, 173
carinatus, Hypostomus, 252
carinatus, Plecostomus, 252
carinatus, Silurus, 170, 171
carinatus, Xenopholis, 433
Carinotus, 234
carinotus, Delturus, 235
carinotus, Plecostomus (Carinotus),
234, 235
carlae, Corydoras, 115
Carnaticus, Pimelodus, 383
carnegiei, Dolichancistrus, 235
carnegiei, Pseudancistrus, 235
carnosus, Chiloglanis, 305
carnosus, Pseudomystus, 103
carolinensis, Mystus, 37
carrancas, Pareiorhina, 281
carrioni, Lipopterichthys, 266, 267
carsonii, Clarias, 145
carvalhoi, Aspidoras, 109
carvalhoi, Bunocephalus, 59
carvalhoi, Harttia, 240
carvalhoi, Hypostomus, 252
carvalhoi, Plecostomus, 252
Cascadura, 108, 127
Cascadura maculocephala, 127, 128
Cascaduridi, 108
cashibo, Loricaria, 270
cashibo, Loricariichthys, 270
castanea, Loricaria, 270
castaneoïdes, Plotosus, 351
castaneo-ventris, Doras, 168
castaneus, Helogenes, 134
castaneus, Leyvaichthys, 134, 407
castaneus, Loricariichthys, 270

FERRARIS

castaneus, Mystus, 94
castaneus, Plotosus, 351
castelnaui, Hemiancistrus, 265
castroi, Hemiloricaria, 245
castroi, Rineloricaria, 245
castroi, Trichomycterus, 416
catamarcensis, Loricaria, 294
catamarcensis, Rineloricaria, 294
catamarcensis, Trichomycterus, 416
cataniai, Leptodoras, 172
cataphracta, Loricaria, 268, 270
Cataphractops, 127
Cataphractus, 110, 167
Cataphractus americanus, 167
Cataphractus callichthys, 111
Cataphractus depressus, 111
Cataphractus punctatus, 111, 123
cataphractus, Plecostomus, 270
cataphractus, Silurus, 167, 168
cataractus, Clarias, 141
cataractus, Phagorus, 141
Catastoma, 48
catharinensis, Glanidium, 75
Cathorops, 39
Cathorops agassizii, 39
Cathorops aguadulce, 39
Cathorops arenatus, 39
Cathorops dasycephalus, 39
Cathorops fuerthii, 39
Cathorops hypophthalmus, 40
Cathorops mapale, 40
Cathorops melanopus, 40
Cathorops multiradiatus, 40
Cathorops spixii, 40
Cathorops steindachneri, 40
Cathorops tuyra, 40
catulus, Pimelodus, 205
catus, Ameiurus, 204
catus, Silurus, 204
caucanus, Ageneiosus, 70
caucanus, Ancistrus, 219
caucanus, Lasiancistrus, 264
caudalis, Synodontis, 312
cauda-furcatus, Pimelodus, 209
caudimaculatus, Corydoras, 115
caudofasciatus, Trichomycterus, 416
caudivittatus, Paradoxoglanis, 302
caudovittata, Synodontis, 312
caudovittatus, Synodontis, 313
cavalliensis, Malapterurus, 301
cavasius, Pimelodus, 95
cavasius, Mystus, 95
cavatica, Peckoltia, 284
cavatura, Erethistoides, 384
cavatus, Arius, 435
caveatus, Hemibagrus, 87
cavernicola, Clarias, 141
cavia, Glyptothorax, 388
cavia, Pimelodus, 388

caxarari, Otocinclus, 274
cayennae, Hoplosternum thoracatum,128cæcutiens, Amblyceps, 18cæcutiens, Silurus, 131cælatus, Arius, 46cælatus, Callichthys, 111cærulescens, Arius, 33cærulescens, Pimelodus, 81
cearensis, Parotocinclus, 282
celsae, Trichomycterus, 416
cenia, Gagata, 386
cenia, Pimelodus, 386
centralis, Clariallabes, 137
centralis, Clarias, 137
centralis, Synodontis, 313
centralus, Hemibagrus, 87
Centranodon japonicus, 379
Centrochir, 170
Centrochir crocodili, 170
Centrochirinae, 166
Centrodoras, 170
Centrodoras brachiatus, 170
Centrodoras hasemani, 170
centrolepis, Ancistrus, 219
Centromochli, 68
Centromochlus, 68, 73
Centromochlus (Gephyromochlus)
leopardus, 75
Centromochlus altae, 73
Centromochlus aulopygius, 77
Centromochlus concolor, 73
Centromochlus creutzbergi, 77
Centromochlus dunni, 77
Centromochlus existimatus, 73
Centromochlus gyrinus, 77
Centromochlus heckelii, 73
Centromochlus intermedius, 77
Centromochlus macracanthus, 73
Centromochlus megalops, 73
Centromochlus perugiae, 73
Centromochlus punctatus, 73
Centromochlus reticulatus, 73
Centromochlus romani, 73
Centromochlus schultzi, 73
Centromochlus Steindachneri, 73
Centrophorus, 411
cephalacanthus, Otocinclus, 276
cephalaspis, Crossoloricaria, 232
Cephalocassis, 41
Cephalocassis borneensis, 41, 42
Cephalocassis jatia, 41
Cephalocassis manillensis, 41
Cephalocassis melanochir, 41
Cephalocassis Stormii, 44, 45
Cephalosilurus, 353
Cephalosilurus albomarginatus, 353
Cephalosilurus apurensis, 353
Cephalosilurus fowleri, 353

CHECKLIST OF CATFISHES, RECENT AND FOSSIL

Cephalosilurus nigricaudus, 353
Ceratocheilus, 71
Ceratocheilus osteomystax, 71, 72
Ceratoglanis, 368
Ceratoglanis pachynema, 368
Ceratoglanis pachynemus, 368
Ceratoglanis scleronemus, 368
ceratophysus, Auchenipterus, 79
ceratophysus, Trachelyopterus, 79
Ceratorhynchus, 68
cerosus, Hemipsilichthys, 279
cerosus, Pareiorhaphis, 279
Cerulescens, Pimelodus, 211
Cerulescens, Silurus, 207, 208, 211
cerulescens melanurus, Silurus, 209
cerulescens var. melanurus, Silurus,
209
cervinus, Corydoras, 115
cesarpintoi, Glanidium, 75
cesarpintoi, Paraotocinclus, 282
cesarpintoi, Parotocinclus, 282
cessator, Breitensteinia, 15
Cetopagansius, 324
Cetopangasius, 324
Cetopangasius chaetobranchus, 324
Cetopsidae, 130
Cetopsidium, 130
Cetopsidium ferreirai, 130
Cetopsidium minutum, 130
Cetopsidium morenoi, 130
Cetopsidium orientale, 130
Cetopsidium pemon, 131
Cetopsidium roae, 131
Cetopsini, 130
Cetopsis, 130, 131
Cetopsis amphiloxa, 131
Cetopsis arcana, 131
Cetopsis baudoensis, 131
Cetopsis caiapo, 131
Cetopsis candiru, 131
Cetopsis chalmersi, 132
Cetopsis coecutiens, 131
Cetopsis fimbriata, 132
Cetopsis gobioides, 131, 132
Cetopsis jurubidae, 132
Cetopsis macroteronema, 133
Cetopsis montana, 132
Cetopsis motatanensis, 132
Cetopsis occidentalis, 134, 135
Cetopsis oliveirai, 132
Cetopsis orinoco, 132
Cetopsis othonops, 132
Cetopsis parma, 132
Cetopsis pearsoni, 132
Cetopsis plumbea, 133
Cetopsis plumbeus, 133
Cetopsis sandrae, 133
Cetopsis sarcodes, 133
Cetopsis Spixii, 131

Zootaxa 1418 © 2007 Magnolia Press · 563

Cetopsis starnesi, 133
Cetopsis umbrosa, 133
Cetopsis ventralis, 135
Cetopsogiton, 135
Cetopsorhamdia, 181
Cetopsorhamdia boquillae, 181
Cetopsorhamdia filamentosa, 182
Cetopsorhamdia iheringi, 182
Cetopsorhamdia insidiosa, 182
Cetopsorhamdia molinae, 182
Cetopsorhamdia nasus, 181, 182
Cetopsorhamdia orinoco, 182
Cetopsorhamdia phantasia, 182
Cetopsorhamdia picklei, 182
Cetopsorhamdia pijpersi, 186
Cetopsorhamdia shermani, 182
ceylonensis, Callichrous, 372
chabanaudi, Atopochilus, 303
chaberti, Trichomycterus, 417
chabutensis, Bachmannia, 428
Chaca, 135
Chaca bankae, 135
Chaca bankanensis, 135
Chaca burmensis, 135
Chaca chaca, 136
Chaca hamiltonii, 135, 136
Chaca lophioides, 135, 136
chaca, Chaca, 136
chaca, Platystacus, 135, 136
Chacidae, 135
Chacini, 135
Chaenothorax, 111
Chaenothorax bicarinatus, 111, 125
Chaenothorax eigenmanni, 125
Chaenothorax multiradiatus, 120
chaetobranchus, Cetopangasius, 324
Chaetostoma, 227
Chaetostoma aburrense, 227
Chaetostoma aequinoctiale, 227
Chaetostoma alternifasciatum, 227
Chaetostoma anale, 227
Chaetostoma anomala sovichthys, 230
Chaetostoma anomalum, 227
Chaetostoma branickii, 228
Chaetostoma breve, 228
Chaetostoma brevilabiatum, 228
Chaetostoma changae, 444
Chaetostoma daidalmatos, 444
Chaetostoma dermorhynchum, 228
Chaetostoma dorsale, 228
Chaetostoma dupouii, 228
Chaetostoma fischeri, 228
Chaetostoma greeni, 228
Chaetostoma guairense, 228
Chaetostoma jegui, 228
Chaetostoma lepturum, 229
Chaetostoma leucomelas, 229
Chaetostoma lineopunctata, 229
Chaetostoma lineopunctatum, 229

Chaetostoma loborhynchos, 227, 229
Chaetostoma machiquense, 229
Chaetostoma machiquensis, 229
Chaetostoma marcapatae, 229
Chaetostoma marginatum, 229
Chaetostoma marmorescens, 229
Chaetostoma microps, 229
Chaetostoma milesi, 229
Chaetostoma mollinasum, 230
Chaetostoma niveum, 230
Chaetostoma nudirostre, 230
Chaetostoma palmeri, 230
Chaetostoma patiae, 230
Chaetostoma paucispinis, 230
Chaetostoma pearsei, 230
Chaetostoma sericeum, 230
Chaetostoma sovichthys, 230
Chaetostoma stannii, 230
Chaetostoma stroumpoulos, 444
Chaetostoma tachiraense, 230
Chaetostoma tachiraensis, 230
Chaetostoma taczanowskii, 231
Chaetostoma thomsoni, 231
Chaetostoma vagum, 231
Chaetostoma vasquezi, 231
Chaetostoma venezuelae, 231
Chaetostoma yurubiense, 231
Chaetostomidi, 217
Chaetostomus, 217
Chaetostomus (Ancistrus) cirrhosus
maculatus, 222
Chaetostomus (Ancistrus) cirrhosus
punctata, 222
Chaetostomus (Ancistrus) cirrhosus
var. maculatus, 222
Chaetostomus (Ancistrus) cirrhosus
var. punctata, 222
Chaetostomus aculeatus, 272
Chaetostomus aequinoctialis, 227
Chaetostomus alga, 221
Chaetostomus annae, 297
Chaetostomus anomalus, 227
Chaetostomus aspidolepis, 242
Chaetostomus bachi, 284
Chaetostomus branickii, 228
Chaetostomus brevilabiatus, 228
Chaetostomus brevis, 228
Chaetostomus cirrhosus var. maculata,
228
Chaetostomus cochliodon, 278
Chaetostomus dentex, 277
Chaetostomus depressus, 287
Chaetostomus dermorhynchus, 228
Chaetostomus dorsalis, 228
Chaetostomus eptingi, 253
Chaetostomus Fischeri, 228
Chaetostomus Fordii, 286
Chaetostomus furcatus, 284
Chaetostomus gibbosus, 278

564 · Zootaxa 1418 © 2007 Magnolia Press

Chaetostomus gigas, 272
Chaetostomus guairensis, 228
Chaetostomus heteracanthus, 264, 265Chætostomus hoplogenys, 221
Chaetostomus Jelskii, 222
Chaetostomus latifrons, 219, 222
Chaetostomus lepturus, 229
Chaetostomus leucomelas, 229
Chaetostomus leucostictus, 222
Chaetostomus macrops, 243
Chaetostomus maculatus, 228
Chaetostomus malacops, 222
Chaetostomus marcapatae, 229
Chaetostomus marginatus, 229
Chaetostomus megacephalus, 244
Chaetostomus microps, 229
Chaetostomus milesi, 229
Chaetostomus mollinasus, 230
Chaetostomus nigrolineatus, 277, 278
Chaetostomus nudirostris, 230
Chaetostomus oligospilus, 284
Chaetostomus palmeri, 230
Chaetostomus paucispinis, 230
Chaetostomus pearsei, 230
Chaetostomus platycephalus, 231
Chaetostomus punctatissimus, 274
Chaetostomus schomburgkii, 265
Chaetostomus sericeus, 230
Chaetostomus setosus, 235
Chaetostomus Stannii, 230
Chaetostomus Taczanowskii, 231
Chaetostomus tectirostris, 221
Chaetostomus thomsoni, 231
Chaetostomus trinitatis, 225
Chaetostomus undecimalis, 292
Chaetostomus vagus, 231
Chaetostomus variolus, 225
Chaetostomus vittatus, 283, 285
chagresi, Ancistrus, 220, 222
chagresi, Pimelodella, 190
Chagresi, Pimelodus(Pseudorhamdia), 190
chagresi odynea, Pimelodella, 193
chalmersi, Cetopsis, 132
chamaeleon, Acrochordonichthys, 12
chamaeleon, Sosia, 12
Chamaigenes, 57
chamaizelus, Bunocephalus, 59
Chandramara, 86
Chandramara chandramara, 86, 105chandramara, Chandramara, 86, 105chandramara, Pimelodus, 86
chandramara, Rama, 86changae, Chaetostoma, 444
changae, Panaqolus, 277
changae, Panaque, 277
changi, Pseudobagrus, 103
chanjoo, Loricariichthys, 270
chanjoo, Parahemiodon, 270

FERRARIS

Channallabes, 137
Channallabes apus, 137
chantrei, Silurus, 379
chaparae, Pimelodella, 190
chaparae, Plecostomus, 226
Chaperi, Diastatomycter, 368
chapini, Acanthocleithron, 303
chapmani, Astroblepus, 63
chapmani, Cyclopium, 63
chapmani, Pygidium, 417
chapmani, Trichomycterus, 417
chappuisi, Doumea, 25
chaquensis, Epapterus, 74
charmuth, Macropteronotus, 138, 148
charrua, Hisonotus, 440
charus, Pimelodus, 355
charus, Pseudopimelodus, 355
chaseni, Leiocassis, 93
Chasmocephalus, 182
Chasmocranes rosae, 183
Chasmocranus, 182
Chasmocranus brachynema, 182
Chasmocranus brachynemus, 182
Chasmocranus brevior, 182
Chasmocranus chimantanus, 183
Chasmocranus longior, 182, 183
Chasmocranus lopezi, 183
Chasmocranus peruanus, 183
Chasmocranus quadrizonatus, 183
Chasmocranus rosae, 183
Chasmocranus surinamensis, 183
Chasmocranus truncatorostris, 183
chaudhurii, Glyptosternum, 385
chauliodon, Brochiloricaria, 227
chechra, Silurus, 374
Cheiridodus, 250
Cheirocerus, 332
Cheirocerus abuelo, 332
Cheirocerus eques, 332
Cheirocerus goeldii, 332
chennuah, Sisor, 402
Cheveyi, Cryptopterus, 371
cheveyi, Micronema, 371
childreni, Ageneiosus, 366
childreni, Silonia, 366
chilensis, Diplomystes, 165
chilensis, Silurus, 165
Chiloglanidinae, 303
Chiloglanis, 303, 304
Chiloglanis angolensis, 304
Chiloglanis anoterus, 304
Chiloglanis asymetricaudalis, 304
Chiloglanis athiensis, 304
Chiloglanis batesii, 304
Chiloglanis benuensis, 305
Chiloglanis bifurcus, 305
Chiloglanis brevibarbis, 305
Chiloglanis cameronensis, 305
Chiloglanis carnosus, 305

Chiloglanis congicus, 305
Chiloglanis deckenii, 304, 305
Chiloglanis disneyi, 305
Chiloglanis dybowskii, 309
Chiloglanis elisabethianus, 305
Chiloglanis emarginatus, 305
Chiloglanis engiops, 308
Chiloglanis fasciatus, 306
Chiloglanis harbinger, 306
Chiloglanis kalambo, 306
Chiloglanis lamottei, 306
Chiloglanis lufirae, 306
Chiloglanis lukugae, 306
Chiloglanis macropterus, 306
Chiloglanis marlieri, 306
Chiloglanis mbozi, 306
Chiloglanis micropogon, 305
Chiloglanis microps, 306
Chiloglanis modjensis, 306
Chiloglanis neumanni, 307
Chiloglanis niger, 307
Chiloglanis niloticus, 307
Chiloglanis niloticus waterloti, 307
Chiloglanis normani, 307
Chiloglanis occidentalis, 307
Chiloglanis paratus, 307
Chiloglanis pojeri, 307
Chiloglanis polyodon, 307
Chiloglanis polypogon, 307
Chiloglanis pretoriae, 307
Chiloglanis productus, 443
Chiloglanis pumilus, 307
Chiloglanis reticulatus, 307
Chiloglanis rukwaensis, 308
Chiloglanis ruziziensis, 308
Chiloglanis sanagaensis, 308
Chiloglanis sardinhai, 308
Chiloglanis somereni, 308
Chiloglanis swierstrai, 308
Chiloglanis trilobatus, 308
Chiloglanis voltae, 308
chiltoni, Pygidium, 417
chiltoni, Trichomycterus, 417
chimantanus, Chasmocranus, 183
Chimarrhoglanis, 21
Chimarrhoglanis Leroyi, 21, 23
Chimarrichthys, 385
Chimarrichthys Davidi, 385
chimborazoi, Astroblepus, 63
chimborazoi, Cyclopium, 63
chinensis, Macrones, 88
chinensis, Pseudobagrus, 101chinensis, Tachisurus, 431
chinta, Bagrus, 431
chiquitos, Callichthys, 128
Chlarias, 138
Chlarias angolensis macronema, 141
Chlarias olivaceus, 146
chlorostictus, Hemiancistrus, 243

CHECKLIST OF CATFISHES, RECENT AND FOSSIL

Choeroplotosus, 346
Choeroplotosus decemfilis, 346
Choeroplotosus limbatus, 346
chondropterygioides, Arius, 55
chondropterygius, Arius, 55
chotae, Arges, 63
chotae, Astroblepus, 63
christyi, Atopochilus, 303
Christyi, Microsynodontis, 309
chrysea, Rita, 105
chryseus, Pseudobagrus, 429
Chrysichthyinae, 153
Chrysichthys, 153, 155
Chrysichthys acutirostris, 158
Chrysichthys aluuensis, 155
Chrysichthys ansorgii, 155
Chrysichthys auratus, 155, 158, 160
Chrysichthys bocagii, 155
Chrysichthys börressoni, 156
Chrysichthys brachynema, 155
Chrysichthys brevibarbis, 156
Chrysichthys büttikoferi, 158
Chrysichthys camaronensis, 158
Chrysichthys cameronensis, 158
Chrysichthys coriscanus, 160
Chrysichthys cranchii, 156
Chrysichthys cyclurus, 161
Chrysichthys dageti, 156
Chrysichthys delhezi, 156
Chrysichthys delhezi thomasi, 156
Chrysichthys duttoni, 156
Chrysichthys eaglesomei, 160
Chrysichthys filamentosus, 155
Chrysichthys furcatus, 156
Chrysichthys grandis, 156
Chrysichthys graueri, 156
Chrysichthys habereri, 156
Chrysichthys helicophagus, 156
Chrysichthys hildae, 153
Chrysichthys johnelsi, 156
Chrysichthys kingsleyae, 160
Chrysichthys lagoensis, 160
Chrysichthys laticeps, 157
Chrysichthys levequei, 157
Chrysichthys longibarbis, 157
Chrysichthys longidorsalis, 157
Chrysichthys longidorsalis nyongensis,
158
Chrysichthys longifilis, 155
Chrysichthys longipinnis, 157
Chrysichthys mabusi, 157
Chrysichthys macropogon, 160
Chrysichthys macrops, 155
Chrysichthys macropterus, 157
Chrysichthys macrotis, 157
Chrysichthys magnus, 157
Chrysichthys mahengeensis, 157
Chrysichthys maurus, 157, 158, 160
Chrysichthys myriodon, 155

Zootaxa 1418 © 2007 Magnolia Press · 565

Chrysichthys nigrodigitatus, 158, 160
Chrysichthys nyongensis, 158
Chrysichthys ogooensis, 158
Chrysichthys ogowensis, 158
Chrysichthys okae, 158
Chrysichthys ornatus, 158
Chrysichthys persimilis, 158
Chrysichthys pitmani, 160
Chrysichthys platycephalus, 158
Chrysichthys polli, 159
Chrysichthys punctatus, 159
Chrysichthys rueppelli, 159
Chrysichthys sharpii, 159
Chrysichthys sianenna, 159
Chrysichthys stappersii, 159
Chrysichthys teugelsi, 159
Chrysichthys theobaldi, 105
Chrysichthys thonneri, 159
Chrysichthys thysi, 159
Chrysichthys uniformis, 159
Chrysichthys velifer, 157, 158
Chrysichthys wagenaari, 159
Chrysichthys walkeri, 159, 160
Chrysobagrus, 155
Chrysobagrus brevibarbis, 155, 156
Chrysobagrus longibarbis, 157
chrysops, Hemibagrus, 87chungaraensis, Trichomycterus, 417
cibelae, Microglanis, 354
cinerascens, Pimelodella, 201
cinerascens, Pimelodus, 200
cinereus, Silurus, 379
Cinetodus, 42
Cinetodus carinatus, 42
Cinetodus conorhynchus, 42
Cinetodus crassilabris, 42
Cinetodus froggatti, 42
cirratus, Arges, 64
cirratus, Astroblepus, 64
cirrhosa, Loricaria, 268, 296
cirrhosa, Vandellia, 427
cirrhosus, Ancistrus, 220
cirrhosus, Hypostomus, 219, 220
cirrhosus dubius, Ancistrus, 221
cirrhosus maculatus, Chaetostomus(Ancistrus), 222
cirrhosus punctata, Chaetostomus(Ancistrus), 222
cirrhosus var. maculata,
Chaetostomus, 228
cirrhosus var. maculatus,
Chaetostomus (Ancistrus), 222
cirrhosus var. punctata, Chaetostomus(Ancistrus), 222
cisternarum, Phreatobius, 429, 430
citatus, Silurichthys, 377
Citernii, Synodontis, 314
citurensis, Oxyloricaria, 300
citurensis, Sturisomatichthys, 300

Claibornichthys, 434
Claibornichthys troelli, 434, 435
clandestinus, Tetracamphilius, 27
Clariallabes, 137
Clariallabes attemsi, 137
Clariallabes brevibarbis, 137
Clariallabes centralis, 137
Clariallabes dumerili, 138
Clariallabes heterocephalus, 137
Clariallabes laticeps, 137
Clariallabes longicauda, 137
Clariallabes longicaudatus, 150
Clariallabes manyangae, 137
Clariallabes melas, 137
Clariallabes melas lembesseensis, 137
Clariallabes mutsindoziensis, 137
Clariallabes petricola, 138
Clariallabes pietschmanni, 138
Clariallabes platyprosopos, 138
Clariallabes simeonsi, 138
Clariallabes teugelsi, 138
Clariallabes uelensis, 138
Clariallabes variabilis, 138
Clarias, 136, 138
Clarias (Allabenchelys) dumerililongibarbis, 138
Clarias (Allabenchelys) submarginatus
thysvillensis, 141
Clarias (Allabenchelys) uelensis, 138
Clarias (Clarioides) aboinensis, 144
Clarias (Clarioides) agboyiensis, 139
Clarias (Clarioides) curtus, 140
Clarias (Clarioides) isheriensis, 139
Clarias abbreviatus, 148
Clarias agboyiensis, 139
Clarias albopunctatus, 139
Clarias alluaudi, 139
Clarias amplexicauda, 148
Clarias anfractus, 139
Clarias angolensis, 139
Clarias anguillaris, 140, 144, 148
Clarias anguillaris nigeriensis, 140
Clarias Assamensis, 148
Clarias batrachus, 140
Clarias batu, 140
Clarias brachysoma, 140
Clarias breviceps, 141
Clarias brevinuchalis, 140
Clarias budgetti, 140
Clarias buettikoferi, 140
Clarias bulumae, 140
Clarias buthupogon, 141
Clarias büttikoferi, 141
Clarias cameronensis, 144
Clarias camerunensis, 139, 141, 144
Clarias capensis, 143
Clarias carsonii, 145
Clarias cataractus, 141
Clarias cavernicola, 141

566 · Zootaxa 1418 © 2007 Magnolia Press

Clarias centralis, 137
Clarias confluentus, 140
Clarias congicus, 142
Clarias dahomeyensis, 142
Clarias dayi, 141
Clarias depressus, 143
Clarias dhonti, 142
Clarias dialonensis, 142
Clarias dolloi, 142
Clarias dorsimarmoratus, 141
Clarias duchaillui, 141
Clarias dumerilii, 142
Clarias dussumieri, 142
Clarias ebriensis, 142
Clarias ekibondoi, 147
Clarias engelseni, 142
Clarias esamesae, 146
Clarias eupogon, 153
Clarias falconeri, 142
Clarias fouloni, 148
Clarias foveolatus, 149
Clarias fuscus, 142
Clarias gabonensis, 142
Clarias gariepinus, 140, 143, 148
Clarias gilli, 146
Clarias gracilis, 144
Clarias guentheri, 143
Clarias guineensis, 141
Clarias hasselquistii, 140
Clarias hilgendorfi, 139
Clarias hilli, 144
Clarias hollyi, 144, 147
Clarias insolitus, 144
Clarias intermedius, 144
Clarias jaensis, 144
Clarias kapuasensis, 144
Clarias kingsleyae, 144
Clarias laeviceps, 142, 144
Clarias laeviceps dialonensis, 142
Clarias lamottei, 144
Clarias lazera, 143
Clarias leiacanthus, 145
Clarias liacanthus, 145
Clarias liberiensis, 141
Clarias lindicus, 141
Clarias liocephalus, 145
Clarias loangwensis, 151
Clarias longibarbis, 136, 138
Clarias longiceps, 143
Clarias longior, 145
Clarias lualae, 142
Clarias maclareni, 145
Clarias macracanthus, 143
Clarias macrocephalus, 145
Clarias macromystax, 145
Clarias macrurus, 148
Clarias malaris, 143
Clarias marpus, 148
Clarias megapogon, 141

FERRARIS

Clarias meladerma, 145
Clarias melanoderma, 146
Clarias melanopogon, 145
Clarias melas, 137
Clarias melasoma, 146
Clarias Mellandi, 146
Clarias microphthalmus, 143
Clarias microstomus, 146
Clarias monkei, 141
Clarias moorii, 143
Clarias mossambicus, 143
Clarias mülleri, 143
Clarias N'gamensis, 146
Clarias nebulosus, 148
Clarias neumanni, 145
Clarias ngamensis, 146
Clarias ngola, 142
Clarias nieuhofii, 139, 146
Clarias nigeriae, 141
Clarias nigricans, 146
Clarias nigromarmoratus, 146
Clarias notozygurus, 143
Clarias nyasensis, 149
Clarias obscurus, 141
Clarias olivaceus, 146
Clarias ornatus, 145
Clarias orontis, 143
Clarias oxycephalus, 146
Clarias pachynema, 146
Clarias parvimanus, 140
Clarias pentapterus, 146
Clarias phillipsi, 145
Clarias planiceps, 147
Clarias platycephalus, 139, 147
Clarias pliocaenicus, 432
Clarias poensis, 141
Clarias pseudoleiacanthus, 147
Clarias pseudonieuhofii, 147
Clarias pulcher, 147
Clarias pulicaris, 142
Clarias punctatus, 140
Clarias robecchii, 143
Clarias salae, 147
Clarias sauteri, 148
Clarias Senegalensis, 140
Clarias smithii, 143
Clarias stappersii, 147
Clarias submarginatus, 147
Clarias sulcatus, 147
Clarias Syriacus, 143
Clarias teijsmanni, 147
Clarias tenuis, 148
Clarias theodorae, 139, 147
Clarias thienemanni, 148
Clarias tsanensis, 143
Clarias vandenhoutei, 144
Clarias varispinis, 147
Clarias vinciguerrae, 143
Clarias walkeri, 141

Clarias werneri, 148
Clarias xenodon, 143
Clarias youngicus, 145
Clarias zygouron, 141
clarias, Silurus, 311, 313, 321, 336,
337
clarias, Pimelodus, 320
clarias, Pseudariodes, 336
clarias, Synodontis, 313
clarias coprophagus, Pimelodus, 337
Clariidae, 136
Clariini, 136
Clarioides, 139
Clarisilurus, 151
Clarisilurus kemratensis, 151, 152
claro, Ancistrus, 220
Claroteidae, 153
Claroteini, 153
Clarotes, 153, 160
Clarotes bidorsalis, 160
Clarotes heuglini, 160
Clarotes laticeps, 160
Clarotes macrocephalus, 160
clauseni, Phractura, 26
clavipinna, Loricaria, 268
clavispinosus, Arius, 36
clavulus, Akysis, 14
clementinae, Ancistrus, 220
cleptolepis, Arius, 47
clijpeaster, Arius, 55
clinatus, Akysis, 14, 357
Clupisoma bastari, 357
Clupisoma garua, 357
Clupisoma longianalis, 357
Clupisoma montana, 357
Clupisoma naziri, 357
Clupisoma nujiangense, 357
Clupisoma prateri, 357
Clupisoma roosae, 357
Clupisoma sinense, 358
clypeastroides, Arius, 55
Cnidoglanis, 345
Cnidoglanis bostockii, 346
Cnidoglanis lepturus, 346, 442
Cnidoglanis macrocephalus, 346
Cnidoglanis microcephalus, 346
Cnidoglanis mülleri, 349
Cnidoglanis nudiceps, 346
coatesi, Arius, 47
coatesi, Neoarius, 47
coatesi, Neosilurus, 348
coatesi, Tandanus, 348
Cobitiglanis, 405
cobrensis, Dolichancistrus, 235
cobrensis, Pseudancistrus pediculatus,
235
cocama, Otocinclus, 274
cochabambae, Imparfinis, 185
cochabambae, Pimelodella, 185

CHECKLIST OF CATFISHES, RECENT AND FOSSIL

cochinchinensis, Arius, 34
cochinchinensis, Pterocryptis, 376
cochinchinensis, Silurus, 376
cochlearis, Hemipimelodus, 43
Cochlefelis, 42
Cochlefelis burmanica, 42
Cochlefelis danielsi, 42
Cochlefelis spatula, 43
Cochliodon, 250
Cochliodon hypostomus, 252
Cochliodon plecostomoides, 258
Cochliodon pospisili, 254
Cochliodon pyrineusi, 259
Cochliodon taphorni, 261
cochliodon, Chaetostomus, 278
cochliodon, Hypostomus, 250, 252,
253-262
cochliodon, Panaque, 277
cochui, Corydoras, 115
coecutiens, Cetopsis, 131
coecutiens, Silurus, 131
coelestinus, Bagrus (Bagrus), 53
coerulescens, Pimelodus, 81
coeruleus, Brochus, 111, 125
coheni, Glyptothorax, 388, 389
coila, Ailia, 356
coila, Malapterurus, 356
colcloughi, Hemipimelodus, 51
Collettii, Heptapterus, 99
collinsae, Parotocinclus, 282
collinus, Leiocassis, 443
colombianus, Bunocephalus, 59
colombiense, Acestridium, 218
colombiensis, Acestridium, 218
colombiensis, Farlowella, 237
colvillii, Macrones, 97
colyeri, Synodontis, 318
colymbetes, Euanemus, 71, 72
commersoni, Hypostomus, 252
Commersonii, Pimelodus, 43, 44
commersonii scabriceps, Plecostomus,
260
commersonoides, Loricaria, 278
commersonoides, Paraloricaria, 278
commersonoides, Plecostomus, 255
comoensis, Synodontis, 313
conchophilus, Pangasius, 325
concolor, Centromochlus, 73
concolor, Corydoras, 115concolor, Tatia, 73
concolor, Trichomycterus, 417
condei, Apistoloricaria, 226
condiscipulus, Corydoras, 115
confinis, Pimelodus, 205
confluentus, Clarias, 140congensis, Schilbe, 363
congensis, Silurus, 362, 363
congica, Parailia, 360
congica, Synodontis, 313

Zootaxa 1418 © 2007 Magnolia Press · 567

congicus, Chiloglanis, 305
congicus, Clarias, 142congicus, Gephyroglanis, 160, 161
congicus, Synodontis, 313
congolensis, Eutropius, 363
congolensis, Schilbe, 364
conirostre, Glyptosternum, 389
conirostre poonaensis, Glyptothorax,
395
conirostris, Conorhynchos, 428
conirostris, Glyptothorax, 389, 395
conirostris, Pimelodus, 428
conirostris punjabensis, Glyptothorax,
393
Conorhynchichthys, 428
Conorhynchos, 53, 428
Conorhynchos conirostris, 428
Conorhynchos nelsoni, 53
Conorhynchus, 428
conorhynchus, Cinetodus, 42conorhynchus, Tetranesodon, 42
Conorhynchus glaber, 428
Conostome, 428
conquetaensis, Pimelodella, 190
conquistador, Gladioglanis, 183conradi, Pygidium, 417
conradi, Trichomycterus, 417
Conta, 382, 383
Conta conta, 384
Conta pectinata, 384
conta, Conta, 384
conta, Pimelodus, 383, 384
Continae, 382
contracta, Synodontis, 313
contractus, Synodontis, 313
cookei, Arius, 50
cookei, Notarius, 50
cooperensis, Neosiluroides, 347
copei, Anduzedoras, 172
copei, Corydoras, 115
copei, Leptodoras, 172
copei, Pimelodella, 190
Copidoglanis, 351
Copidoglanis brevidorsalis, 347
Copidoglanis curtus, 345
Copidoglanis equinus, 348
Copidoglanis gjellerupi, 348
Copidoglanis glencoensis, 348
Copidoglanis idenburgi, 348
Copidoglanis labiosus, 345
Copidoglanis labrosus, 345
Copidoglanis levis, 345
Copidoglanis longifilis, 349
Copidoglanis meraukensis, 351
Copidoglanis novae-guineae niger,
349
Copidoglanis obscurus, 345, 350
Copidoglanis papuensis, 348
Copidoglanis rendahli, 345

Copionodon, 403, 404
Copionodon lianae, 404
Copionodon orthiocarinatus, 404
Copionodon pecten, 404
Copionodontinae, 403
coppenamensis, Corydoras, 115
coppenamensis, Corydoras bondi, 115
coppenamensis, Hypostomus, 253
coprophagus, Pimelodus, 337
coprophagus, Pimelodus clarias, 337
coquenani, Ancistrus (Pseudancistrus),
287
coquenani, Pseudancistrus, 287
coracoideus, Auchenipterichthys, 71
coracoideus, Bunocephalus, 59
coracoideus, Dysichthys, 58, 59
coracoideus, Trachycorystes, 71
Coraglanis, 385
Coraglanis kishinouyei, 385
corantijni, Hypostomus, 253cordobensis, Leptoancistrus, 266
Cordorinus, 111
cordovae, Plecostomus, 258
corduvensis, Trichomycterus, 417
Cordylancistrus, 231, 443
Cordylancistrus daguae, 231
Cordylancistrus nephelion, 443
Cordylancistrus perijae, 231
Cordylancistrus platycephalus, 231
Cordylancistrus platyrhynchus, 232
Cordylancistrus torbesensis, 232
Coreobagrus, 86, 100
Coreobagrus brevicorpus, 86Coreobagrus ichikawai, 86
Coreobagrus okadai, 86
coriaceus, Trachelyopterus, 79
coriaceus maculosus, Trachelyopterus,
79
coriatae, Corydoras, 115
coriscanus, Chrysichthys, 160
corruscans, Platystoma, 341
corruscans, Pseudoplatystoma, 341
corsula, Bagrus, 89
corsula, Macrones, 90
corsula, Mugil, 89
corsula, Pimelodus, 89
cortesi, Corydoras, 124
Corumbataia, 232
Corumbataia cuestae, 232
Corumbataia tocantinensis, 232
coruscans, Pimelodes, 37
coruscans, Platystoma, 341
Corydoradinae, 108
Corydoras, 108, 111
Corydoras acrensis, 112
Corydoras acutus, 112
Corydoras adolfoi, 112
Corydoras aeneus, 112
Corydoras agassizii, 112

568 · Zootaxa 1418 © 2007 Magnolia Press

Corydoras albolineatus, 112
Corydoras amandajanea, 112
Corydoras amapaensis, 113
Corydoras ambiacus, 113
Corydoras amphibelus, 113
Corydoras approuaguensis, 113
Corydoras araguaiaensis, 113
Corydoras arcuatus, 113
Corydoras areio, 113
Corydoras armatus, 113
Corydoras atropersonatus, 113
Corydoras aurofrenatus, 113
Corydoras australe, 118
Corydoras axelrodi, 113
Corydoras baderi, 113
Corydoras bertoni, 130
Corydoras bicolor, 114
Corydoras bifasciatus, 114
Corydoras bilineatus, 114
Corydoras blochi, 114
Corydoras blochi vittatus, 126
Corydoras boehlkei, 114
Corydoras boesemani, 114
Corydoras bolivianus, 117
Corydoras bondi, 114
Corydoras bondi coppenamensis, 115
Corydoras breei, 114
Corydoras brevirostris, 114
Corydoras britskii, 114
Corydoras burgessi, 114
Corydoras caquetae, 119
Corydoras carlae, 115
Corydoras caudimaculatus, 115
Corydoras cervinus, 115
Corydoras cochui, 115
Corydoras concolor, 115
Corydoras condiscipulus, 115
Corydoras copei, 115
Corydoras coppenamensis, 115
Corydoras coriatae, 115
Corydoras cortesi, 124
Corydoras crimmeni, 115
Corydoras cruziensis, 115
Corydoras crypticus, 115
Corydoras davidsandsi, 116
Corydoras delphax, 116
Corydoras difluviatilis, 116
Corydoras diphyes, 116
Corydoras dubius, 126
Corydoras duplicareus, 116
Corydoras edentatus, 176
Corydoras ehrhardti, 116
Corydoras eigenmanni, 129
Corydoras elegans, 111, 116
Corydoras elegans nijsseni, 121
Corydoras ellisae, 116
Corydoras ephippifer, 116
Corydoras episcopi, 126
Corydoras eques, 112, 116

FERRARIS

Corydoras esperanzae, 117
Corydoras evelynae, 117
Corydoras filamentosus, 117
Corydoras flaveolus, 117
Corydoras fowleri, 117
Corydoras funnelli, 119
Corydoras garbei, 117
Corydoras geoffroy, 111, 117
Corydoras geryi, 117
Corydoras gomezi, 117
Corydoras gossei, 118
Corydoras gracilis, 118
Corydoras grafi, 113
Corydoras griseus, 118
Corydoras guapore, 118
Corydoras guianensis, 118
Corydoras habrosus, 118
Corydoras haraldschultzi, 118
Corydoras hastatus, 112, 118
Corydoras heteromorphus, 118
Corydoras imitator, 118
Corydoras incolicana, 118
Corydoras isbrueckeri, 119
Corydoras julii, 119
Corydoras juquiaae, 121
Corydoras juquiae, 121
Corydoras kanei, 119
Corydoras kronei, 129
Corydoras lacerdai, 119
Corydoras lamberti, 119
Corydoras latus, 119
Corydoras leopardus, 119
Corydoras leucomelas, 119
Corydoras loretoensis, 119
Corydoras loxozonus, 119
Corydoras macropterus, 130
Corydoras macrosteus, 112
Corydoras maculatus, 122
Corydoras maculifer, 119
Corydoras mamore, 120
Corydoras mamoré, 120
Corydoras marmoratus, 122
Corydoras melanistius, 120
Corydoras melanistius brevirostris,
114
Corydoras melanistius longirostris,
113
Corydoras melanotaenia, 120
Corydoras melini, 120
Corydoras meridionalis, 116
Corydoras metae, 120
Corydoras micracanthus, 120
Corydoras microcephalus, 122
Corydoras microps, 112
Corydoras multimaculatus, 120
Corydoras multiradiatus, 120
Corydoras myersi, 123
Corydoras nanus, 120
Corydoras napoensis, 120

Corydoras narcissus, 120
Corydoras nattereri, 121
Corydoras nattereri triseriatus, 121
Corydoras negro, 121
Corydoras nijsseni, 121
Corydoras noelkempffi, 121
Corydoras octocirrus, 117
Corydoras oelemariensis, 114
Corydoras oiapoquensis, 121
Corydoras ornatus, 121
Corydoras orphnopterus, 121
Corydoras osteocarus, 121
Corydoras ourastigma, 121
Corydoras oxyrhynchus, 121
Corydoras paleatus, 122
Corydoras panda, 122
Corydoras pantanalensis, 122
Corydoras paragua, 122
Corydoras parallelus, 122
Corydoras pastazensis, 122
Corydoras pastazensis orcesi, 122
Corydoras paucerna, 122
Corydoras pauciradiatus, 109
Corydoras pestai, 116
Corydoras pinheiroi, 122
Corydoras polystictus, 123
Corydoras potaroensis, 123
Corydoras prionotos, 130
Corydoras pulcher, 123
Corydoras punctatus, 123
Corydoras punctatus argentina, 122
Corydoras punctatus sipaliwini, 125
Corydoras pygmaeus, 123
Corydoras rabauti, 123
Corydoras raimundi, 110
Corydoras reticulatus, 123
Corydoras revelatus, 123
Corydoras reynoldsi, 123
Corydoras robineae, 124
Corydoras robustus, 124
Corydoras sanchesi, 124
Corydoras saramaccensis, 124
Corydoras sarareensis, 124
Corydoras schultzei, 112
Corydoras schwartzi, 124
Corydoras schwartzi surinamensis,
126
Corydoras semiaquilus, 124
Corydoras semiscutatus, 125
Corydoras septentrionalis, 124
Corydoras serratus, 124
Corydoras seussi, 124
Corydoras similis, 124
Corydoras simulatus, 124
Corydoras sipaliwini, 125
Corydoras sodalis, 125
Corydoras solox, 125
Corydoras spectabilis, 125
Corydoras spilurus, 125

CHECKLIST OF CATFISHES, RECENT AND FOSSIL

Corydoras splendens, 125
Corydoras steindachneri, 125
Corydoras stenocephalus, 125
Corydoras sterbai, 125
Corydoras surinamensis, 126
Corydoras sychri, 126
Corydoras treitlii, 126
Corydoras trilineatus, 126
Corydoras tukano, 126
Corydoras undulatus, 126
Corydoras venezuelanus, 112
Corydoras virescens, 123
Corydoras virginiae, 126
Corydoras vittatus, 126
Corydoras weitzmani, 126
Corydoras wotroi, 120
Corydoras xinguensis, 126
Corydoras zygatus, 126
Corymbophanes, 217, 232
Corymbophanes andersoni, 232
Corymbophanes bahianus, 279
Corymbophanes kaiei, 232
Corymbophanes venezuelae, 231
Cossyphus, 139
Cossyphus ater, 139, 148costatus, Platydoras, 176
costatus, Silurus, 176
cottoides, Microglanis, 354
cottoides, Pimelodus(Pseudopimelodus), 354
Cotylephorus, 61
Cotylephorus blochii, 61
cotylephorus, Platystacus, 61
couma, Bagrus, 53
couma, Sciades, 53
courteti, Synodontis, 313
cous, Arius, 31
cous, Glyptothorax, 31, 389
cous, Silurus, 31, 387, 389
cragini, Amiurus, 205
cranchii, Chrysichthys, 156
cranchii, Pimelodus, 156
Cranoglanididae, 164
Cranoglanidae, 164
Cranoglanis, 164
Cranoglanis bouderius, 165
Cranoglanis henrici, 165
Cranoglanis multiradiatus, 165
Cranoglanis sinensis, 164, 165
crassicauda, Harttia, 242
crassicauda, Harttiella, 242
crassicauda, Hypostomus, 253
crassicauda, Pseudecheneis, 400
crassilabris, Cinetodus, 42
crassilabris, Hemipimelodus, 42
crassilabris, Leiocassis, 92
crassilabris, Liocassis, 92
crassilabris macrops, Leiocassis, 92
crassirostris, Leiocassis. 92

Zootaxa 1418 © 2007 Magnolia Press · 569

crassirostris, Liocassis, 92
crassus, Arius, 435
crassus, Otolithus cf., 434
crassus, Otolithus (incertae sedis), 434
crassus, Tachysurus, 434, 435
cratensis, Trachycorystes, 80
crenula, Pterocryptis, 376
cretaceus, Brachyspondylus, 432
creutzbergi, Centromochlus, 77
creutzbergi, Tatia, 77
crimmeni, Corydoras, 115
crinalis, Bagrus, 55
cristata, Pimelodella, 190
cristata, Physopyxis, 176
cristatus, Parotocinclus, 282
cristatus, Pimelodus, 189, 190
crocodili, Centrochir, 170
Crocodili, Doras, 170
crossocheilos, Arius, 52
Crossoloricaria, 232
Crossoloricaria bahuaja, 232
Crossoloricaria cephalaspis, 232
Crossoloricaria rhami, 232
Crossoloricaria variegata, 233
Crossoloricaria venezuelae, 233
cruciger, Arius, 106
Cruciglanis, 443
Cruciglanis pacifici, 443
cruxenti, Pimelodella, 190
cruziensis, Corydoras, 115
Cryptarius, 43
Cryptarius daugueti, 43
Cryptarius truncatus, 43
crypticus, Corydoras, 115
crypticus, Noturus, 211, 442
cryptobullatus, Amphilius, 21
cryptodon, Planiloricaria, 285
cryptodon, Pseudohemiodon(Planiloricaria), 285
cryptophthalmus, Ancistrus, 220
Cryptopterella, 369
Cryptopterella beldti, 369, 371
Cryptopterus, 369
Cryptopterus amboinensis, 369
Cryptopterus bleekeri, 374
Cryptopterus cheveyi, 371
Cryptopterus latovittatus, 374
Cryptopterus lumholtzi, 370
Cryptopterus urbaini, 373
cryptopterus, Kryptopterus, 369
cryptopterus, Silurus, 369
cryptus, Paradoxoglanis, 303
Cteniloricaria, 233
Cteniloricaria fowleri, 233
Cteniloricaria maculata, 233
Cteniloricaria platystoma, 233
ctenodus, Pimelodus, 332
cuangoana, Synodontis, 313
cuangoanus, Synodontis, 313

cuao, Pygidianops, 412
cubangoensis, Amphilius platychir, 23
cubataonis, Loricaria, 294
cubataonis, Rineloricaria, 294
cubataonis, Trichomycterus, 425
cucphuongensis, Pterocryptis, 376
cucphuongensis, Silurus, 376
cucuhyensis, Pseudepapterus, 76
cuestae, Corumbataia, 232
cuiabae, Ancistrus, 220
cuivi, Nematogenys, 324
cultratus, Pangasius, 328
cunaguaro, Ginesia, 330, 331
cunningtoni, Dinotopterus, 148
cupido, Trachymochlus, 431, 432
cuprea, Pimelodus (Rhamdia) Queleni,201
cupreoides, Pimelodus, 205
cupreus, Silurus, 215
curtirostra, Farlowella, 237
curtisii, Arius, 47
curtisoma, Encheloclarias, 149
curtus, Clarias (Clarioides), 140
curtus, Copidoglanis, 345
curvirostris, Belonoglanis, 24
curvispina, Loricaria, 296
curvispina, Spatuloricaria, 296
cuspicaudus, Sorubim, 342
cuvieri, Genidens, 44
Cuvieri, Silurus (Acanthonotus), 356
Cuyabae, Pimelodus (Rhamdia), 201
cuyanus, Diplomystes viedmensis, 166
cuyanus, Olivaichthys, 166
cyanochloros, Pimelodus, 392
cyanostigma, Pimelodella, 190
cyanostigma, Rhamdia, 190
Cyclopidae, 63
Cyclopium, 63
Cyclopium chapmani, 63
Cyclopium chimborazoi, 63
Cyclopium humboldtii, 63, 64
Cyclopium mariae, 65
Cyclopium mindoense, 65
Cyclopium pirrense, 66
Cyclopium trifasciatum, 67
Cyclopium ubidiai, 67
Cyclopium unifasciatum, 67
Cyclopium vanceae, 67
Cyclopium ventrale, 68
cyclopium, Pimelodus, 64
cyclopum, Pimelodus, 64
cyclopus, Astroblepus, 64
cyclopus santanderensis, Astroblepus,
67
cyclurus, Chrysichthys, 161
cyclurus, Lophiobagrus, 161Ddageti, Chrysichthys, 156
dageti, Synodontis, 312

570 · Zootaxa 1418 © 2007 Magnolia Press

daguae, Cordylancistrus, 231
daguae, Hemiancistrus, 231
dahli, Anodontiglanis, 345
dahomeyensis, Clarias, 142
daidalmatos, Chaetostoma, 444
daillyi, Hoplosternum littorale, 128
dakpathari, Glyptothorax, 389
dalungshanensis, Mystus, 95
damasceni, Ancistrus, 220
damasceni, Xenocara, 220
danicus, Otolithus (Arius), 434
danielsi, Arius (Hemiarius), 42
danielsi, Cochlefelis, 42
dantei, Auchenipterichthys, 71
dariense, Sturisoma, 299
dariensis, Oxyloricaria, 299
dasycephalus, Arius, 39
dasycephalus, Cathorops, 39
Dasyloricaria, 233
Dasyloricaria capetensis, 233
Dasyloricaria filaentosa, 233
Dasyloricaria latiura, 233
Dasyloricaria seminuda, 234
Dasyloricaria tuyrensis, 234
daugueti, Cryptarius, 43
Daugueti, Hemipimelodus, 43
dauuricus, Silurus, 379
Davalla, 68
Davalla Schomburgkii, 68, 69
davalla, Hypothalmus, 69
Davidi, Chimarrichthys, 385
davidi, Euchiloglanis, 385
davidi, Synodontis, 313
davidsandsi, Corydoras, 116
davisi, Pygidium, 417
davisi, Trichomycterus, 417
davissinghi, Glyptothorax, 389
dawalla, Hypothalmus, 69
dayi, Arius, 49
dayi, Batasio, 85
dayi, Clarias, 141
dayi, Hemipimelodus, 46Dayi, Macrones, 85
dayi, Nedystoma, 46
de Zwaani, Pangasius, 328
debauwi, Eutropius, 361
debauwi, Pareutropius, 361
Decapogon, 127
Decapogon urostriatum, 127
Decapogon urotriatum, 127
Decapogon verissimi, 127
decaradiatus, Trachelyichthys, 78
decemfilis, Trachelyichthys, 346
decipiens, Otolithus (Sciaenidarum),434
deckenii, Chiloglanis, 304, 305
decolor, Rhamdia guatemalensis, 202
decora, Synodontis, 314
decorus, Amiurus, 210

FERRARIS

decorus, Synodontis, 314
degeni, Bagrus, 84
deignani, Kryptopterus, 375
dekayi, Pimelodus, 206
Dekeyseria, 234
Dekeyseria amazonica, 234
Dekeyseria brachyura, 234
Dekeyseria niveata, 234
Dekeyseria picta, 234
Dekeyseria pulchra, 234
Dekeyseria scaphirhyncha, 234
dekimpei, Synodontis, 314
dekkanense, Glyptosternum, 391
delacouri, Oreoglanis, 397
delacouri, Paroreoglanis, 397
delhezi, Chrysichthys, 156delhezi thomasi, Chrysichthys, 156delicata, Psammphiletria, 27
delicatissimus, Pangasius, 328
delphax, Corydoras, 116
Deltadoras, 173
Deltadoras guayoensis, 173, 174
Delturinae, 444
Delturus, 234, 444
Delturus angulicauda, 234, 444
Delturus brevis, 443
Delturus carinotus, 235
Delturus parahybae, 234, 235, 444
demantoides, Baryancistrus, 226
demararae, Callichthys callichthys,111
demerarae, Auchenipterus, 72
dendrophorus, Rheoglanis, 164
Dentata, Loricaria, 268
dentatus, Ageneiosus, 70
dentatus, Auchenipterus, 72
dentatus, Doras, 177
Dentectus, 235
Dentectus barbarmatus, 235
dentex, Chaetostomus, 277
dentex, Panaqolus, 277
dentex, Panaque, 277
Denticetopsis, 133
Denticetopsis epa, 133
Denticetopsis iwokrama, 133
Denticetopsis macilenta, 133
Denticetopsis praecox, 133
Denticetopsis royeroi, 133
Denticetopsis sauli, 133, 134
Denticetopsis seducta, 134
depauwi, Synodontis, 314
depierrei, Platyglanis, 164
depinnai, Aspidoras, 109
Deportator, 350
Deppei, Pimelodus, 200
depressa, Harttia, 240
depressa, Hemiodon, 293
depressa, Reganella, 293
depressa, Rhamdia, 201

depressa, Tocantinsia, 78
depressicauda, Hisonotus, 248
depressicauda, Otocinclus, 248
depressinotus, Hisonotus, 248
depressinotus, Microlepidogaster, 248
depressirostris, Bagrus, 364
depressirostris, Eutropius, 364
depressus, Acanthodoras, 167
depressus, Bunocephalus, 62
depressus, Cataphractus, 111
depressus, Chaetostomus, 287
depressus, Clarias, 143
depressus, Doras (Rhinodoras), 167
depressus, Gnathobagrus, 161
depressus, Hemiodon, 292, 293
depressus, Pseudancistrus, 287
depressus, Pterobunocephalus, 62
deqinensis, Glyptothorax, 389
derbyi, Hypostomus, 253
derbyi, Loricariichthys, 270
derbyi, Plecostomus, 253
Dermocassis, 92
dermorhynchum, Chaetostoma, 228
dermorhynchus, Chaetostomus, 228
despaxi, Arius, 36
devall, Hypophthalmus, 334
devincenzii, Loricaria, 288
devincenzii, Pseudohemiodon, 288
deyi, Amblyceps, 17dhonti, Allabenchelys, 142
dhonti, Clarias, 142
dhonti, Synodontis, 314
diabolicus, Plectrochilus, 411
diabolicus, Urinophilus, 411
diabolus, Trichomycterus, 417
dialonensis, Clarias, 142dialonensis, Clarias laeviceps, 142
Dianema, 108, 127
Dianema longibarbis, 127
Dianema urostriatum, 127
Dianemidi, 108
Diapeltoplites, 249
diaphanus, Ammoglanis, 403
diaphina, Ageniosus (Silonia), 86
Diastatomycter, 368
Diastatomycter Chaperi, 368
Diasternum, 127
dibrugarensis, Macrones montanus, 96
dichromum, Acestridium, 218
dicra, Scoloplax, 367
Dieperinki, Arius, 32
difluviatilis, Corydoras, 116
digulensis, Arius, 47
diluvii, Bucklandium, 432, 433
dinema, Belodontichthys, 368
dinema, Wallago, 368
Dinotopteroides, 139
Dinotopteroides prentissgrayi, 139,
146

CHECKLIST OF CATFISHES, RECENT AND FOSSIL

Dinotopterus, 148
Dinotopterus atribranchus, 148
Dinotopterus cunningtoni, 148
Dinotopterus euryodon, 136
Dinotopterus filicibarbis, 148
Dinotopterus foveolatus, 149
Dinotopterus gigas, 149
Dinotopterus ilesi, 136
Dinotopterus jacksoni, 149
Dinotopterus jallae, 146
Dinotopterus longibarbis, 136
Dinotopterus loweae, 149
Dinotopterus nyasensis, 149
Dinotopterus rotundifrons, 136
Dinotopterus worthingtoni, 136
dioctes, Arius, 44
dioctes, Hemiarius, 44
diphyes, Corydoras, 116
Diplomystax, 165
Diplomyste, 165
Diplomystes, 165
Diplomystes camposensis, 165
Diplomystes chilensis, 165
Diplomystes mesembrinus, 166
Diplomystes nahuelbutaensis, 166
Diplomystes rudis, 435
Diplomystes viedmensis, 166
Diplomystes viedmensis cuyanus, 166
Diplomystes viedmensis mesembrinus,
166
Diplomystidae, 165
dipterus, Brochus, 111, 125
discus, Acestridium, 218
disjunctivus, Liposarcus, 291
disjunctivus, Pterygoplichthys, 291
disneyi, Chiloglanis, 305
dispar, Arius, 34
dispar, Pygidium, 417
dispar, Trichomycterus, 417
dispilurus, Epapterus, 74
dissidens, Harttia, 241dissitus, Kryptopterus, 369
distolothrix, Scoloplax, 367
djambal, Pangasius 325
djemeri, Eutropius, 363
djemeri, Schilbe, 363
dlouhyi, Hypostomus, 253
doceana, Steindachneria, 343
doceana, Steindachneridion, 343
doceanus, Microlepidogaster, 282
doceanus, Parotocinclus, 282
docmac niger, Bagrus, 84
docmak, Bagrus, 84, 313
docmak, Silurus, 84
Doiichthyidae, 30
Doiichthys, 30, 46
Doiichthys novaeguineae, 46
Doiichthys novae-guineae, 46
Dolichallabes, 149

Zootaxa 1418 © 2007 Magnolia Press · 571

Dolichallabes microphthalmus, 149
Dolichamphilius, 24
Dolichamphilius brieni, 24
Dolichamphilius longiceps, 24
Dolichancistrus, 235
Dolichancistrus atratoensis, 235
Dolichancistrus carnegiei, 235
Dolichancistrus cobrensis, 235
Dolichancistrus fuesslii, 235
Dolichancistrus pediculatus, 235
Dolichancistrus setosus, 235
dolicholophia, Scoloplax, 367
dolichopterus, Ancistrus, 220
dolichurus, Bunocephalus, 62
dolichurus, Pterobunocephalus, 62
dolloi, Clarias, 142
Doradidae, 166
Doradini, 166dorae, Leptoglanis, 28
dorae, Zaireichthys, 28
Doraops, 170
Doraops zuloagai, 170
Doras, 166, 170
Doras (Agamyxis) flavopictus, 168
Doras (Corydoras) brevis, 179
Doras (Corydoras) ophthalmus, 169
Doras (Corydoras) punctatus, 167
Doras (Oxydoras) d’Orbignyi, 178
Doras (Oxydoras) lipophthalmus, 169
Doras (Oxydoras) niger, 175
Doras (Oxydoras) stenopeltis, 171, 172
Doras (Rhinodoras) depressus, 167
Doras affinis, 168
Doras albomaculatus, 168
Doras armatulus, 176
Doras asterifrons, 169, 170
Doras Blochii, 167
Doras brachiatus, 170
Doras brunnescens, 168
Doras Calderonensis, 179
Doras carinatus, 171, 173
Doras castaneo-ventris, 168
Doras cataphractus, 168
Doras Crocodili, 170
Doras dentatus, 177
Doras dorsalis, 173
Doras fimbriatus, 167
Doras granulosus, 177
Doras grypus, 169
Doras Hancockii, 177
Doras Heckelii, 178
Doras helicophilus, 177
Doras humboldti, 176
Doras humeralis, 174
Doras insculptus, 179
Doras laevigatulus, 177
Doras lentiginosus, 177
Doras libertatis, 174
Doras lithogaster, 173

Doras longispinis, 170
Doras loricatus, 167
Doras maculatus, 177
Doras marmoratus, 171
Doras micropoeus, 171
Doras murica, 177
Doras nebulosus, 178
Doras niger, 175
Doras Oxyrhynchus, 169
Doras papilionatus, 173
Doras pectinifrons, 168
Doras polygramma, 168
Doras punctatus, 167
Doras regani, 169
Doras spinosissimus, 168
Doras uranoscopus, 173, 174
Doras weddellii, 169
d’Orbigny, Doras (Oxydoras), 178
d’Orbignyi, Doras (Oxydoras), 178dorbignyi, Doras (Oxydoras), 178dorbigny, Rhinodoras, 178dorbignyi, Rhinodoras, 178
doriae, Arius, 41
doriae, Bunocephalus, 59
doriae, Liocassis, 93
doroides, Bagrus, 45
dorsale, Chaetostoma, 228
dorsalis, Chaetostomus, 228
dorsalis, Doras, 173
dorsalis, Glyptothorax, 389
dorsalis, Lithodoras, 173
dorsalis, Rhamdia, 201
dorseyi, Pimelodella, 190
dorsimarmoratus, Clarias, 141dorsomaculata, Synodontis, 314
dorsomaculatus, Synodontis, 314
dorsostriatum, Pygidium, 417
dorsotriatum, Pygidium, 417
dorsotriatus, Trichomycterus, 417
Doumea, 20, 24
Doumea alula, 24
Doumea angolensis, 25
Doumea chappuisi, 25
Doumea scaphyrhynchura, 27
Doumea thysi, 25
Doumea typica, 24, 25
Doumeinae, 20
dowii, Leptarius, 53
dowii, Sciades, 53
Duanensis, 378
duanensis, Silurus, 379
dubia, Netuma, 54
dubius, Ancistrus, 221
dubius, Corydoras, 126
duchaillui, Clarias, 141
duda, Silurus, 374
duellmani, Trichomycterus, 418
dugesii, Ictalurus, 208, 209
dugèsii, Amiurus, 208

572 · Zootaxa 1418 © 2007 Magnolia Press

dukai, Silurus, 377
dumerili, Clariallabes, 138
dumerili, Leiocassis, 93
dumerili, Rhinobagrus, 92, 93
dumerili longibarbis, Clarias(Allabenchelys), 138
dumerilii, Clarias, 142
dumus, Lasiancistrus, 289
dumus, Pseudolithoxus, 289
dunni, Centromochlus, 77
dunni, Tatia, 77
duodecimalis, Hypostomus, 290, 291
Duopalatinus, 332
Duopalatinus emarginatus, 332
Duopalatinus olallae, 340
Duopalatinus peruanus, 332
Duoplatinus goeldii, 340
duplicareus, Corydoras, 116
dupouii, Chaetostoma, 228
Dupouyichthys, 60
Dupouyichthys sapito, 60
duquei, Rhamdia, 202dura, Loricaria, 268
duriceps, Ostophycephalus, 346
durinii, Eutropius, 363
durinii, Schilbe, 363
duriventris, Harttia, 241
duseni, Hemipsilichthys, 262, 279
duseni, Isbrueckerichthys, 262
Dussumieri, Arius, 52
dussumieri, Clarias, 142
dussumieri, Plicofollis, 52
Dutemplei, Arius, 56
duttoni, Chrysichthys, 156
dybowskii, Chiloglanis, 309
dybowskii, Euchilichthys, 309
Dysichthys, 58
Dysichthys amazonicus, 59
Dysichthys australe, 60
Dysichthys coracoideus, 58, 59
Dysichthys quadriradiatus, 60Eeaglesomei, Eaglesomia, 160
Eaglesomia, 160
Eaglesomia eaglesomei, 160
ebriensis, Clarias, 142
eburneensis, Synodontis, 312
echinatus, Ictalurus, 208
echthrus, Megalocentor, 409
eclipsis, Eutropius, 363
eddsi, Pseudecheneis, 442
edentatus, Corydoras, 176
edentatus, Hypophthalmus, 333
edentatus, Loricariichthys, 270
egertoni, Arius, 56
Egertoni, Silurus, 56
egertoni belgicus, Arius, 56
egertonii, Callichrous, 374
ehrhardti, Corydoras, 116

FERRARIS

eichorniarum, Ituglanis, 407
eichorniarum, Trichomycterus, 407
Eidouxii, Galeichthys, 38
eigenmanni, Arges, 64
eigenmanni, Astroblepus, 64
eigenmanni, Chaenothorax, 125
eigenmanni, Corydoras, 129
eigenmanni, Galeichthys, 33
eigenmanni, Heptapterus, 184
eigenmanni, Hemiloricaria, 245
Eigenmanni, Loricaria, 245
eigenmanni, Orinocodoras, 175
eigenmanni, Oxydoras, 167
eigenmanni, Pimelodella, 191, 340
eigenmanni, Pimelodella, 193
eigenmanni, Pimelodus, 340
eigenmanni, Pimelodus (Pimelodella),191
eigenmanni, Propimelodus, 340
eigenmanni, Pygidianops, 412
eigenmanni, Trichomycterus, 423
eigenmanniorum, Pimelodella, 191
eigenmanniorum, Rhamdia, 191
ekibondoi, Clarias, 147
elassochir, Noturus, 212
elatturus, Arius, 50
electricus, Malapterurus, 301
electricus, Silurus, 301
electricus ogooensis, Malapterurus,
302
electricus oguensis, Malapterurus, 302
elegans, Corydoras, 111, 116
elegans, Noturus, 212
elegans nijsseni, Corydoras, 121eleutherus, Noturus, 212
elisabethianus, Chiloglanis, 305
Ellenriederii, Leiocassis, 104
Elliops, 207
ellisae, Corydoras, 116
Ellisichthys, 127
elongata, Eumeda, 347, 348
elongata, Hara, 384
elongata, Olyra, 99
elongata, Pimelodella, 191
elongatus, Batasio, 73
elongatus, Macrones, 89
elongatus, Nemadoras, 174
elongatus, Oxydoras, 174
elongatus, Pangasius, 326
elongatus, Pimelodus, 191
elongatus, Plotosus, 346
elongatus, Sorubim, 342
elongatus hongus, Hemibagrus, 89
emanueli, Trichomycterus, 418
emanueli emanueli, Pygidium, 418
emanueli motatanensis, Pygidium, 421
emarginata, Microsynodontis, 309
emarginata, Squaliforma, 297
emarginatum, Platystoma, 332

emarginatus, Chiloglanis, 305
emarginatus, Duopalatinus, 332
emarginatus, Hypostomus, 297
emarginatus, Liocassis, 102
emarginatus, Microsynodontis, 309
emarginatus, Pseudobagrus, 100
Emini, Schilbe, 364
emmelane, Tachysurus, 40
emphysetus, Bagrus (Sciades), 53, 54
empousa, Scoloplax, 367
Encheloclarias, 149
Encheloclarias baculum, 149
Encheloclarias curtisoma, 149
Encheloclarias kelioides, 149
Encheloclarias prolatus, 150
Encheloclarias tapeinopterus, 150
Encheloclarias velatus, 150
Endorrhis, 349
enfurnada, Rhamdia, 197
engelseni, Allabenchelys, 142engelseni, Clarias, 142
engiops, Chiloglanis, 308
enigmatica, Lacantunia, 216
enochi, Pimelodella, 191
Entomocorus, 74
Entomocorus benjamini, 74
Entomocorus gameroi, 74
Entomocorus melaphareus, 74
Entomocorus radiosus, 443
Eobagrus, 87
Eobagrus hoerdzanicus, 87
Eomacrones, 87
Eomacrones wilsoni, 87
Eopeyeria, 428
Eopeyeria aegyptiaca, 428
epa, Denticetopsis, 133
Epactionotus, 236
Epactionotus aky, 236
Epactionotus bilineatus, 236
Epactionotus gracilis, 236
Epactionotus itaimbezinho, 236
Epactionotus yasi, 236
epakmos, Isbrueckerichthys, 262
Epapterus, 74
Epapterus blohmi, 74
Epapterus chaquensis, 74
Epapterus dispilurus, 74
ephippiata, Amphilius pulcher, 23
ephippifer, Akysis, 14
ephippifer, Corydoras, 116
epikarsticus, Ituglanis, 407
episcopi, Corydoras, 126eppleyi, Parotocinclus, 282
eptingi, Chaetostomus, 253
eptingi, Hypostomus, 253
equatorialis, Tachysurus, 40
eques, Cheirocerus, 332
eques, Corydoras, 112, 116
eques, Goeldiella, 184

CHECKLIST OF CATFISHES, RECENT AND FOSSIL

eques, Pimelodus, 184
equestris, Arius, 33
equinus, Copidoglanis, 348
equinus, Neosilurus, 348
erebennus, Amiurus, 205
Eremophilini, 403
Eremophilus, 403, 404
Eremophilus camposi, 408
Eremophilus candidus, 416
Eremophilus mutisii, 404
Erethistes, 382, 384, 444
Erethistes maesotensis, 384
Erethistes pusillus, 384
Erethistes pussilus, 384
Erethistidae, 382
Erethistides, 382
Erethistoides, 382
Erethistoides ascita, 384
Erethistoides cavatura, 384
Erethistoides infuscatus, 442
Erethistoides montana, 384
Erethistoides montana pipri, 384
Erethistoides pipri, 384
Erethistoides sicula, 384
eriarcha, Rhamdella, 196
eriarcha, Rhamdia, 196
ericae, Hypostomus, 253
ericius, Hypostomus, 253
erinaceus, Ancistrus, 221
erinaceus, Hypostomus, 221
Ernstichthys, 60
Ernstichthys anduzei, 60
Ernstichthys intonsus, 60
Ernstichthys megistus, 60
erythrogaster, Silurus (Callichrus),
373
Erythroptera, Silurus maculatus, 209
erythrurus, Urinophilus, 427
esamesae, Clarias, 146
esmeraldas, Paracetopsis, 135
esperanzae, Corydoras, 117
espiritosantensis, Neoplecostomus,
273
essequibensis, Leptoglanis, 187
essequibensis, Leptorhamdia, 187
estuarius, Ayarnangra, 382
etentaculatum, Hypostoma, 291
etentaculatus, Pterygoplichthys, 291
Euacanthagenys, 296
Euacanthagenys caquetae, 296
euacanthagenys, Spatuloricaria, 296
Euanemini, 68
Euanemus, 68, 71
Euanemus brachyurus, 72
Euanemus colymbetes, 71, 72
Euanemus longipinnis, 74
Euanemus nigripinnis, 72
Euchilichthys, 308
Euchilichthys astatodon, 308

Zootaxa 1418 © 2007 Magnolia Press · 573

Euchilichthys boulengeri, 309
Euchilichthys dybowskii, 309
Euchilichthys guentheri, 309
Euchilichthys habereri, 309
Euchilichthys royauxi, 309
Euchiloglanis, 384
Euchiloglanis davidi, 385
Euchiloglanis gracilicaudata, 398
Euchiloglanis hodgarti, 398
Euchiloglanis kamengensis, 399
Euchiloglanis kishinouyei, 385
Euchiloglanis longicauda, 399
Euchiloglanis macrotrema, 399
Euchiloglanis myzostoma, 399
Euchiloglanis sinensis, 399
Euclyptosternum, 387
eugeneiatus, Callichrous, 369
eugeneiatus, Kryptopterus, 369
Euglyptosternum lineatum, 388
Eumeda, 347
Eumeda elongata, 347, 348
eupogoides, Pseudobagrus, 107
eupogon, Clarias, 153
eupogon, Pelteobagrus, 100
eupogon, Pseudobagrus, 100
eupogon, Xenoclarias, 153
euptera, Synodontis, 314
eupterus, Synodontis, 314
Euristhmus, 346
Euristhmus lepturus, 346
Euristhmus microceps, 346
Euristhmus microphthalmus, 442
Euristhmus nudiceps, 346
Euristhmus sandrae, 442
eurycephalus, Aspidoras, 109
eurycephalus, Hemipsilichthys, 279
eurycephalus, Pareiorhaphis, 279
Eurycheilichthys, 236
Eurycheilichthys limulus, 236
Eurycheilichthys pantherinus, 236
Eurycheilus, 236
Eurycheilus pantherinus, 236
euryodon, Bathyclarias, 136
euryodon, Dinotopterus, 136
eurystoma, Microglanis, 354
eurystomus, Satan, 216
eurystomus, Synodontis, 305, 319
eustictus, Ancistrus, 221
eustictus, Pristiancistrus, 219, 221
eutaenia, Pimelodella, 191
Eutropiellus, 360
Eutropiellus buffei, 361
Eutropiellus kasai, 360, 361
Eutropiellus vandeweyeri, 361
Eutropiichthys, 358
Eutropiichthys burmannicus, 358
Eutropiichthys murius, 358
Eutropiichthys vacha, 358
Eutropius, 362

Eutropius altipinnis, 365
Eutropius angolensis, 363
Eutropius ansorgii, 363
Eutropius banguelensis, 363
Eutropius bocagii, 363
Eutropius bomae, 363
Eutropius brachypopterus, 362
Eutropius brevianalis, 363
Eutropius cameronensis, 365
Eutropius congolensis, 363, 364
Eutropius debauwi, 361
Eutropius depressirostris, 364
Eutropius djemeri, 363
Eutropius durinii, 363
Eutropius eclipsis, 363
Eutropius gastratus, 363
Eutropius grenfelli, 363
Eutropius laticeps, 364
Eutropius lemairii, 364
Eutropius liberiensis, 364
Eutropius longifilis, 361
Eutropius macropthalmos, 361, 362
Eutropius mandibularis, 364
Eutropius mentalis, 364
Eutropius micropogon, 364
Eutropius möbiusii, 364, 365
Eutropius moebii, 365
Eutropius multilineatus, 365
Eutropius multitaeniatus, 365
Eutropius nasalis, 363
Eutropius nyongensis, 365
Eutropius obtusirostris, 365
Eutropius seraoi, 363
Eutropius tumbanus, 365
Eutropius yangambianus, 365
Evansii, Loricaria, 296
evansii, Spatuloricaria, 296
evelynae, Corydoras, 117evermanni, Tachysurus, 40
Exallodontus, 333
Exallodontus aguanai, 333
exaratus, Callichthys, 129
Exastilithoxus, 236
Exastilithoxus fimbriatus, 237
Exastilithoxus hoedemani, 237
Exilichthys, 346
exilis, Noturus, 212
exilis, Trachelyichthys, 78
existimatus, Centromochlus, 73
exodon, Bagrus, 359
exodon, Glyptothorax, 389
Exostoma, 382, 385
Exostoma andersonii, 386
Exostoma berdmorei, 385
Exostoma Blythii, 396
Exostoma Feae, 398
Exostoma gracile, 387
Exostoma labiatum, 385
Exostoma labrax, 387

574 · Zootaxa 1418 © 2007 Magnolia Press

Exostoma macropterum, 398
Exostoma Oschanini, 402
Exostoma stoliczkae, 387
Exostoma Vinciguerrae, 385
Exostomatina, 382
exsudans, Pimelodus, 196
exsudans, Rhamdella, 196
Eydouxii, Galeichthys, 38
Ffabricioi, Callichthys, 111
Fajumia, 428
Fajumia menoni, 429
Fajumia misrai, 429
Fajumia schweinfurthi, 428, 429
Fajumia stromeri, 429
falcarius, Arius, 34
falcarius, Mystus, 95
falcarius africana, Arius, 31
falcarius var. africana, Arius, 31
falcifer, Acrochordonichthys, 12
falconeri, Clarias, 143
fallax, Hemiloricaria, 245
fallax, Loricaria (Loricariichthys), 245
fangi, Arius, 34
fangi, Pseudobagrus, 101
Farlowella, 217, 237
Farlowella acestrichthys, 238
Farlowella acus, 237
Farlowella acus venezuelensis, 239
Farlowella agustini, 240
Farlowella altocorpus, 444
Farlowella amazona, 237
Farlowella amazonum, 237
Farlowella angosturae, 240
Farlowella azygia, 238
Farlowella boliviana, 238
Farlowella carinata, 237
Farlowella colombiensis, 237
Farlowella curtirostra, 237
Farlowella gladiolus, 237
Farlowella gracilis, 237
Farlowella guaricensis, 240
Farlowella hahni, 238
Farlowella hargreavesi, 238
Farlowella hasemani, 238
Farlowella henriquei, 238
Farlowella isbruckeri, 238
Farlowella jauruensis, 238
Farlowella jauruënsis, 238
Farlowella knerii, 238
Farlowella latisoma, 239
Farlowella mariaelenae, 238
Farlowella martini, 238
Farlowella nattereri, 238
Farlowella odontotumulus, 238
Farlowella oliveirae, 237
Farlowella oxyrryncha, 239
Farlowella paraguayensis, 239
Farlowella paranaënse, 237

FERRARIS

Farlowella parvicarinata, 239
Farlowella platorynchus, 239
Farlowella pleurotaenia, 237
Farlowella pseudogladiolus, 237
Farlowella reticulata, 239
Farlowella roncallii, 240
Farlowella rugosa, 239
Farlowella schreitmuelleri, 239
Farlowella schreitmülleri, 239
Farlowella smithi, 239
Farlowella taphorni, 239
Farlowella venezuelensis, 239
Farlowella vittata, 239
Farlowellidi, 217
farsonensis, Hypsidoris, 203fasciata, Phractura, 26
fasciata, Schilbe senegalensis, 364
fasciatum, Pseudoplatystoma, 341
fasciatum brevefile, Pseudoplatystoma,341
fasciatum intermedium,
Pseudoplatystoma, 341
fasciatum nigricans,Pseudoplatystoma, 341
fasciatum reticulatum,
Pseudoplatystoma, 341
fasciatus, Auchenoglanis, 163
fasciatus, Chiloglanis, 306
fasciatus, Nannoglanis, 187, 188
fasciatus, Noturus, 212
fasciatus, Silurus, 340, 341
fasciolatus, Batasio, 442
fascipinna, Synodontis, 314
fassli, Pygidium, 418
fassli, Trichomycterus, 418
fasslii, Pygidium, 418
feae, Exostoma, 398
feae, Pareuchiloglanis, 398
feliceps, Galeichthys, 43
Felichthys, 37
Felichthys amblops, 79
Felichthys filamentosus, 37
Felichthys flavescens, 76
Felichthys stauroforus, 32
felinus, Pimelodus, 205
felipponei, Loricaria, 294
felipponei, Rineloricaria, 294
felis, Ariopsis, 33
felis, Pimelodus, 206
felis, Silurus, 33
ferox, Silurus, 431
ferrarisi, Niobichthys, 274
ferreirai, Cetopsidium, 130
ferula, Pseudolaguvia, 442
festae, Arges, 64
festae, Arius, 56
festae, Astroblepus, 64
festae, Plecostomus, 263
festinus, Arius, 34

festivum, Sturisoma, 299
figueroai, Pimelodella, 191
filamentissima, Harttia, 264
filamentosa, Cetopsorhamdia, 182
filamentosa, Dasyloricaria, 233
filamentosa, Hara, 395
filamentosa, Harttia, 263, 264
filamentosa, Loricaria, 233
filamentosa, Synodontis, 314
filamentosa latiura, Loricaria, 233
filamentosa seminuda, Loricaria, 234
filamentosum, Brachyplatystoma, 330
filamentosus, Aspredinichthys, 57
filamentosus, Aspredo, 57
filamentosus, Bagrus, 84
filamentosus, Chrysichthys, 155filamentosus, Corydoras, 117
filamentosus, Felichthys, 37
filamentosus, Lamontichthys, 263
filamentosus, Pimelodes, 330
filamentosus, Synodontis, 314
filamentus, Hemibagrus, 88
filamentus, Macrones (Hemibagrus),
88
filicaudata, Peckoltia, 284
filicaudatus, Peckoltichthys, 283, 284
filicibarbis, Bathyclarias, 148
filicibarbis, Dinotopterus, 148
filifer, Akysis, 16
filifer, Pseudobagarius, 16
filinemus, Phyllonemus, 164
fimbriata, Cetopsis, 132
fimbriata, Loricaria, 297
fimbriata, Spatuloricaria, 297
fimbriatus, Doras, 167
fimbriatus, Exastilithoxus, 237
fimbriatus, Hypophthalmus, 333
fimbriatus,Otocinclus, 271
fimbriatus, Pseudacanthicus(Lithoxus), 236, 237
fimbriatus, Pseudocanthicus(Lithoxus), 237
firestonei, Paramphilius, 25
fisadoha, Plotosus, 350
fischeri, Chaetostoma, 228
Fischeri, Chaetostomus, 228
fisheri, Trachelyopterus, 79
fisheri, Trachycorystes, 79
fissidens, Arges, 64
fissidens, Astroblepus, 64
fissipinnis, Heptapterus, 184
fissus, Arius, 39
flabelliferus, Ochmacanthus, 409
flagellaris, Plecostomus, 268
flava, Loricaria, 259
flavater, Noturus, 212
flaveolus, Corydoras, 117
flavescens, Bagrus, 54
flavescens, Felichthys, 76

CHECKLIST OF CATFISHES, RECENT AND FOSSIL

flavescens, Pseudauchenipterus, 76
flavicans, Bagrus, 344
flavipinnis, Noturus, 212
flavipinnis, Pimelodina, 336
flavipinnis, Pimelodus (Pimelodina),
336
flavipinnis, Pseudomystus, 103
flavitaeniata, Synodontis, 314
flavitaeniatus, Synodontis, 314
flavolineatus, Plotosus, 345
flavomaculatus, Leptoglanis, 28
flavomaculatus, Zaireichthys, 28
flavopictus, Doras (Agamyxis), 168
flavus, Bagrus, 90
flavus, Hyalobagrus, 92
flavus, Noturus, 211, 212
flexilis, Macrotocinclus, 271
flexilis, Otocinclus, 271, 272
florense, Pygidium, 414
fluviatilis, Hypostomus, 253
fluviatilis, Leiocassis, 85
fluviatilis, Mystus, 164
fluviatilis, Plecostomus, 253
Fluvidraco, 101
foina, Pimelodus, 197
foina, Rhamdia, 197
fokiensis, Glyptosternum, 389
fokiensis, Glyptothorax, 389
fonchii, Hypostomus, 253
Fonchiiichthys, 240
Fonchiiichthys rupestris, 240
Fonchiiichthys uracanthus, 240
foratum, Amblyceps, 18
fordicei, Auchenipterus, 72
Fordii, Chaetostomus, 286
fordii, Pseudacanthicus, 286
forficulatus, Hypodoras, 172
formosa, Hemiloricaria, 245
formosa, Rineloricaria, 245
formosanus, Liobagrus, 19
formoso, Ancistrus, 221
formosus, Astroblepus, 64
fortis, Hemibagrus, 88
fortis, Macrones, 88
fortis capitulum, Macrones, 88
fortis var. capitulum, Macrones, 88
fossilis, Heteropneustes, 151
fossilis, Silurus, 151
fossor, Pimelodes, 431
Fouloni, Clarias, 148
foveolata, Pseudolaguvia, 401
foveolatus, Clarias, 149
foveolatus, Dinotopterus, 149
fowleri, Cephalosilurus, 353
fowleri, Corydoras, 117
fowleri, Cteniloricaria, 233
Fowleri, Oxyloricaria, 233
fowleri, Pangasius, 328
Fraasi, Arius, 56

Zootaxa 1418 © 2007 Magnolia Press · 575

francirochai, Hisonotus, 248
francirochai, Otocinclus, 248
francisci, Hypostomus, 254
francisci, Plecostomus, 254
Franciscodoras, 171
Franciscodoras marmoratus, 171
franciscoensis, Neoplecostomus, 273
frankei, Aphanotorulus, 225, 226
freiei, Ageneiosus, 70
frenata, Brachyglanis, 180
frenata, Loricaria, 299
frenatum, Sturisoma, 299
frenatus, Astroblepus, 64
frenatus, Brachyglanis, 180
frenatus, Oreoglanis, 397
frenatus, Rhamdioglanis, 202
froggatti, Arius, 42froggatti, Cinetodus, 42frontosa, Synodontis, 314
frontosus, Synodontis, 314
fuelleborni, Synodontis, 315
fuerthii, Cathorops, 39
fuesslii, Dolichancistrus, 235
fugleri, Hemiancistrus, 243fui, Pseudobagrus, 107
fukiensis, Glyptothorax, 389
fukiensis fukiensis, Glyptothorax, 389
fukiensis hainanensis, Glyptothorax,390
fukiensis honghensis, Glyptothorax,390
fukiensis honghenensis, Glyptothorax,
390
fuliginatus, Akysis, 14
fuliginosus, Hemiancistrus, 243
fulva, Xenocara, 221
fulvidraco, Pelteobagrus, 100
fulvi-draco, Pimelodus, 100
fulvus, Ancistrus, 221
fumidus, Ompok, 372
fumosus, Pseudomystus, 103
funebris, Noturus, 212
funnelli, Corydoras, 119
fur, Pimelodus, 337
fur, Pseudorhamdia, 337
furcata, Peckoltia, 284
furcatus, Auchenipterus, 76
furcatus, Chaetostomus, 284
furcatus, Chrysichthys, 156
furcatus, Hemibagrus, 88
furcatus, Ictalurus, 208, 211
furcatus, Nemasiluroides, 361
furcatus, Pimelodus, 208, 209
furcifer, Pimelodus, 210
Furcodontichthys, 240
Furcodontichthys novaesi, 240
furiosus, Noturus, 211, 212
furnessi, Apodoglanis, 375, 376
furnessi, Pterocryptis, 376

Fürthii, Arius, 39
Fuscatus, Silurus lividus, 205
fuscoguttatus, Aspidoras, 109
fuscum, Pygidium, 427
fuscus, Akysis, 16
fuscus, Ancharius, 29
fuscus, Bagrus, 95
fuscus, Clarias, 142
fuscus, Glyptothorax, 389
fuscus, Leiocassis, 103
fuscus, Macropteronotus, 142
fuscus, Neobagrus, 19, 20
fuscus, Pseudobagarius, 16
fuscus, Pseudomystus, 103
Fusiloricaria, 268
füsslii, Ancistrus, 235
Ggabardinii, Ageneiosus, 69
gabonensis, Clarias, 142
gabrieli, Pygidium, 418
gabrieli, Trichomycterus, 418
gadense, Nigerium, 98
Gagata, 385
Gagata cenia, 386
Gagata dolichonema, 386
Gagata gagata, 386
Gagata gasawyuh, 386
Gagata itchkeea, 386
Gagata melanoptera, 386
Gagata melanopterus, 386
Gagata pakistanica, 386
Gagata sexualis, 386
Gagata typus, 386
Gagata youssoufi, 386
gagata, Gagata, 385, 386
gagora, Arius, 34
gagora, Pimelodus, 34
gagorides, Bagrus, 35
gagoroides, Arius, 35
galani, Ancistrus, 221
galaxias, Leporacanthicus, 265
galaxias, Tatia, 77
galeatus, Silurus, 79
galeatus, Trachelyopterus, 79
Galeichthys, 43
Galeichthys aequus, 435
Galeichthys aguadulce, 39
Galeichthys angelicus, 434
Galeichthys araguayensis, 339
Galeichthys ater, 43
Galeichthys azureus, 33
Galeichthys bahiensis, 38
Galeichthys Blochii, 38
Galeichthys bonillai, 33
Galeichthys eigenmanni, 33
Galeichthys, Eidouxii, 38
Galeichthys Eydouxii, 38
Galeichthys feliceps, 43
Galeichthys gilberti, 33

576 · Zootaxa 1418 © 2007 Magnolia Press

Galeichthys Gronovii, 37
Galeichthys guentheri, 33
Galeichthys ocellatus, 43
Galeichthys Parrae, 38
Galeichthys peruvianus, 43
Galeichthys simonsi, 33
Galeichthys stanneus, 49
Galeichthys xenauchen, 33
galinae, Synodontis, 323
gallowayi, Schilbeodes, 212
gambensis, Pimelodus, 54
gambiensis, Synodontis, 315
gambiensis latifrons, Synodontis, 315
gameroi, Entomocorus, 74
gangelica, Pterocryptis, 375, 376, 377
gangetica, Silundia, 366
Gangra, 395
garavelloi, Harttia, 241
garavelloi, Microglanis, 354
garbei, Corydoras, 117garbei, Hemipsilichthys, 279
garbei, Pareiorhaphis, 279
garbei, Pimelodela, 189
garciabarrigai, Pimelodus, 337
garcia-barrigai, Pimelodus, 337
garhwali, Glyptothorax, 389
gariepinus, Clarias, 140, 143, 148gariepinus, Silurus (Heterobranchus),
143
garmani, Hypostomus, 254
garmani, Plecostomus, 254
garua, Clupisoma, 357
garua, Silurus, 357
gasawyuh, Gagata, 386
gastratus, Eutropius, 363
Gastrodermus, 111
gaudryi, Silurus, 432
Gelanoglanis, 73, 74
Gelanoglanis nanonocticolus, 74
Gelanoglanis stroudi, 74
gelatinosus, Malacoglanis, 408
geledensis, Synodontis, 315
geminus, Kryptopterus, 369
genibarbis, Pseudorinelepis, 289
genibarbis, Rinelepis, 289
Genidens, 32, 43
Genidens barbus, 44
Genidens cuvieri, 44
Genidens genidens, 44
Genidens granulosus, 44
Genidens machadoi, 44
Genidens planifrons, 44
Genidens valenciennesii, 44
genidens, Genidens, 44
genidens, Pimelodus, 43, 44
genisetiger, Pseudancistrus, 287
geoffroy, Corydoras, 111, 117
Geoffroyi, Heterobranchus, 150
Gephyroglanidini, 153

FERRARIS

Gephyroglanis, 153, 160
Gephyroglanis barnardi, 81
Gephyroglanis congicus, 160, 161
Gephyroglanis gigas, 157
Gephyroglanis gilli, 81
Gephyroglanis gymnorhynchus, 161
Gephyroglanis habereri, 161
Gephyroglanis longipinnis, 157
Gephyroglanis lowei, 158
Gephyroglanis ogoensis, 158
Gephyroglanis ogooensis, 158
Gephyroglanis rotundiceps, 28
Gephyroglanis sclateri, 81
Gephyroglanis tilhoi, 155
Gephyroglanis velifer, 157
Gephyromochlus, 75
germanicus, Otolithus (Arius), 434
geryi, Corydoras, 117
geryi, Pimelodella, 191
gibbiceps, Ancistrus, 291
gibbiceps, Pterygoplichthys, 291
gibbiceps, Silurichthys, 377
gibbosa, Lampiella, 264
gibbosus, Chaetostomus, 278
gibbosus, Otocinclus, 264
gigantea, Vandellia, 427
giganteus, Trichomycterus, 418
gigas, Arius, 31, 56
gigas, Bagarius, 383
gigas, Bathyclarias, 149
gigas, Chaetostomus, 272
gigas, Dinotopterus, 149
gigas, Gephyroglanis, 157
gigas, Pangasianodon, 325
gigas, Platystoma, 330
gigas, Xiurenbagrus, 20
gilberti, Galeichthys, 33gilberti, Noturus, 213
gilberti, Silurus, 376
gilli, Austroglanis, 81
gilli, Clarias, 146
gilli, Gephyroglanis, 81
gilli, Rhamdella, 196
gilli, Rhamdia, 196
Ginesia, 330
Ginesia cunaguaro, 330, 331
gjellerupi, Copidoglanis, 348
gjellerupi, Neosilurus, 348
glaber, Auchenipterus, 80
glaber, Conorhynchus, 428
glaber, Otolithus (Arius ?), 435
gladiator, Noturus, 213
Gladioglanis, 183
Gladioglanis conquistador, 183
Gladioglanis machadoi, 183
gladiolus, Acestra, 237
gladius, Acestra, 239
gladysae, Phractura, 26
Glanapteryginae, 403

Glanapteryx, 403, 404
Glanapteryx anguilla, 404
Glanapteryx niobium, 405
Glanidium, 75
Glanidium albescens, 75
Glanidium bockmanni, 75
Glanidium catharinensis, 75
Glanidium cesarpintoi, 75
Glanidium leopardum, 75
Glanidium melanopterum, 75
Glanidium neivai, 77
Glanidium piresi, 78
Glanidium ribeiroi, 75
Glanis, 37, 378
Glanis Aristotelis, 378
glanis aralensis, Silurus, 379
glanis, Silurus, 378, 379
Glaphyropoma, 405
Glaphyropoma rodriguesi, 405
Glaridoglanis, 386
Glaridoglanis andersonii, 386
glencoensis, Copidoglanis, 348
gloveri, Neosilurus, 348
Glyptocephalus, 432
Glyptocephalus radiatus, 432, 433
Glyptoperichthys, 290
Glyptoperichthys joselimaianus, 291
Glyptoperichthys parnaibae, 292
Glyptoperichthys xinguensis, 292
Glyptosterni, 382
Glyptosternon, 382, 387
Glyptosternon akhtari, 387
Glyptosternon hainanensis, 390
Glyptosternon interspinalum, 390
Glyptosternon labiatus, 385
Glyptosternon maculatum, 387
Glyptosternon malaisei, 387
Glyptosternon minutum, 393
Glyptosternon Nieuwenhuisi, 392
Glyptosternon pectinopterus, 392
Glyptosternon punctatum, 389
Glyptosternon quadriocellatum, 393
Glyptosternon reticulatum, 387
Glyptosternon reticulatus, 387
Glyptosternon retuculatus, 387
Glyptosternon striatus, 387, 394
Glyptosternon sulcatus, 400
Glyptosternum, 387
Glyptosternum akhtari, 387
Glyptosternum armeniacum, 388
Glyptosternum chaudhurii, 385
Glyptosternum conirostre, 389
Glyptosternum dekkanense, 391
Glyptosternum fokiensis, 389
Glyptosternum gracile, 390
Glyptosternum hodgarti, 398
Glyptosternum kükenthali, 391
Glyptosternum kurdistanicum, 390
Glyptosternum laak, 395

CHECKLIST OF CATFISHES, RECENT AND FOSSIL

Glyptosternum madraspatanum, 391
Glyptosternum modestum, 389
Glyptosternum pallozonum, 388, 392
Glyptosternum saisii, 393
Glyptosternum sinense, 393
Glyptosternum Stoličkae, 393
Glyptosternum steindachneri, 393
Glyptosternum stuarti, 385
Glyptosternum sykesi, 394
Glyptosternum tiong, 395
Glyptosternum yunnanensis, 401
Glyptothoracini, 382
Glyptothorax, 382, 387
Glyptothorax alaknandi, 388
Glyptothorax anamalaiensis, 388
Glyptothorax annandalei, 388
Glyptothorax armeniacus, 388
Glyptothorax botius, 388
Glyptothorax brevipinnis, 388
Glyptothorax brevipinnis alaknandi,
388
Glyptothorax buchanani, 388
Glyptothorax burmanicus, 388
Glyptothorax callopterus, 388
Glyptothorax cavia, 388
Glyptothorax coheni, 388, 389
Glyptothorax conirostre poonaensis,
395
Glyptothorax conirostris, 389, 395
Glyptothorax conirostris punjabensis,393
Glyptothorax cous, 31, 389
Glyptothorax dakpathari, 389
Glyptothorax davissinghi, 389
Glyptothorax deqinensis, 389
Glyptothorax dorsalis, 389
Glyptothorax exodon, 389
Glyptothorax fokiensis, 389
Glyptothorax fukiensis fukiensis, 389
Glyptothorax fukiensis hainanensis,
390
Glyptothorax fukiensis honghensis,
390
Glyptothorax fukiensis honghenensis,
390
Glyptothorax fuscus, 389
Glyptothorax garhwali, 389
Glyptothorax gracilis, 389, 390
Glyptothorax hainanensis, 390
Glyptothorax honghensis, 390
Glyptothorax horai, 390
Glyptothorax housei, 390
Glyptothorax indicus, 390
Glyptothorax interspinalus, 390
Glyptothorax jalalensis, 390
Glyptothorax kashmirensis, 390, 393
Glyptothorax kurdistanicus, 390
Glyptothorax lampris, 391
Glyptothorax laosensis, 391

Zootaxa 1418 © 2007 Magnolia Press · 577

Glyptothorax lonah, 391, 394
Glyptothorax longicauda, 391
Glyptothorax longinema, 394
Glyptothorax longjiangensis, 391
Glyptothorax macromaculatus, 391
Glyptothorax madraspatanus, 391
Glyptothorax major, 391
Glyptothorax manipurensis, 391
Glyptothorax merus, 390
Glyptothorax minimaculatus, 391
Glyptothorax minutus, 391, 392, 393
Glyptothorax naziri, 392
Glyptothorax nelsoni, 392
Glyptothorax nieuwenhuisi, 392
Glyptothorax obscura, 392
Glyptothorax obscurus, 392
Glyptothorax pallozonus, 392
Glyptothorax panda, 392
Glyptothorax pectinopterus, 392
Glyptothorax platypogon, 392
Glyptothorax platypogonides, 392
Glyptothorax platypogonoides, 392
Glyptothorax prashadi, 393
Glyptothorax punjabensis, 393
Glyptothorax quadriocellatus, 393
Glyptothorax rubermentus, 394
Glyptothorax saisii, 393
Glyptothorax siamensis, 393
Glyptothorax silviae, 393
Glyptothorax sinensis, 393
Glyptothorax spectrum, 393
Glyptothorax steindachneri, 393
Glyptothorax stocki, 393
Glyptothorax stolickae, 393
Glyptothorax striatus, 394
Glyptothorax sufii, 394
Glyptothorax sykesi, 394
Glyptothorax telchitta, 394
Glyptothorax telchitta sufii, 394
Glyptothorax trewavasae, 394
Glyptothorax trilineatoides, 394
Glyptothorax trilineatus, 394
Glyptothorax tuberculatus, 401, 402
Glyptothorax ventrolineatus, 394
Glyptothorax zainaensis, 394
Glyptothorax zanaensis, 394
Glyptothorax zhujiangensis, 394
Gnathobagrus, 161
Gnathobagrus depressus, 161
gnomus, Panaqolus, 277
gnomus, Panaque, 277
goae, Silurus, 374
Gobibagrus, 87
Gobibagrus hoerdzanicus, 87
gobio, Hemipsilichthys, 247
gobio, Xenomystus, 247
gobioides, Cetopsis, 131, 132gobroni, Synodontis, 315
godavarii, Pangasius pangasius, 327

godfreyi, Tachysurus (Pararius), 45
godmanni, Pimelodus, 200
Goeldiella, 184
Goeldiella eques, 184
goeldii, Brachyplatystoma, 330
goeldii, Cheirocerus, 332
goeldii, Duoplatinus, 340
goeldii, Pimelodina, 332
goeldii, Trichomycterus, 418
Gogangra, 395
Gogangra laevis, 395
Gogangra viridescens, 395
Gogo, 29
Gogo arcuatus, 29
Gogo brevibarbis, 29
Gogo ornatus, 29
gogra, Phractocephalus, 105
gogra, Rita, 105Gogrius, 105Gogrius sykesii, 105
Goliath, Bagrus, 331
gomesi, Plecostomus, 298
gomesi, Squaliforma, 298
gomezi, Corydoras, 117
gongshanensis, Pareuchiloglanis, 398
goniaspis, Arius, 52
Gonocephalus, 160
gonzalezi, Amblydoras, 168
gonzalezi, Zathorax, 168
goodi, Paramphilius, 25
goongwaree, Hypophthalmus, 362
goongwaree, Proeutropiichthys, 362
gorgona, Trichomycterus, 418
Goslinia, 330
gossei, Corydoras, 118
gossei, Malapterurus, 302
gouldingi, Helogenes, 134
gouldingi, Stauroglanis, 413
goyazensis, Hypostomus, 254
goyazensis, Plecostomus, 254
gracile, Exostoma, 387
gracile, Glyptosternum, 390
gracilicaudata, Euchiloglanis, 398
gracilicaudatus, Pareuchiloglanis,398
graciliformis, Schmidelia, 432
gracilior, Ituglanis, 407
gracilior, Pygidium, 407
gracilis, Clarias, 144
gracilis, Corydoras, 118
gracilis, Epactionotus, 236
gracilis, Farlowella, 237
gracilis, Glyptothorax, 389, 390
gracilis, Harttia, 241
gracilis, Hemibagrus, 88
gracilis, Pimelodella, 191
gracilis, Pimelodus, 191, 196, 210
gracilis, Osteogeneiosus, 51
gracilis, Pseudepapterus, 76

578 · Zootaxa 1418 © 2007 Magnolia Press

gracilis, Pseudobagrus, 102
gracilis, Schultzichthys, 413
gracilis, Trichomycterus, 422
graciosa, Homodiaetus, 406
graciosus, Pimelodus, 210
Graeffei, Arius, 47
graeffei, Neoarius, 47
grafi, Corydoras, 113
grahami, Silurus, 379
grammatophorus, Amphilius, 22, 23
grammatophorus brevipinna,
Amphilius, 22
grammatophorus inaequalis,
Amphilius, 23
grammatophorus marmoratus,
Amphilius, 23
grandicassis, Arius, 49, 50
grandicassis, Notarius, 50
grandiops, Synodontis, 445
grandis, Amphilius, 23
grandis, Auchenoglanis, 163
grandis, Chrysichthys, 156
grandis, Parakysis 15
grandis, Socnopaea, 430
grandiscutata, Rita, 105
grandoculis, Arius, 53
grandoculis, Potamarius, 53
granducolis, Arius, 53
grangeri, Rhineastes, 432
granosus, Arius, 54
granosus, Hypostomus, 273
granosus, Neoplecostomus, 273
granulosa, Synodontis, 315
granulatus, Arius, 32
granulosus, Doras, 177
granulosus, Genidens 44
granulosus, Pterodoras, 177
granulosus, Synodontis, 315
graueri, Chrysichthys, 156
gravoti, Auchenoglanis ballayi, 162
greeni, Chaetostoma, 228
grenfelli, Eutropius, 363
grenfelli, Schilbe, 363
greshoffi, Synodontis, 315
griffini, Pimelodella, 191
grisea, Pimelodella, 191
griseus, Ancharius, 29
griseus, Corydoras, 118
griseus, Limatulichthys, 266
griseus, Loricaria, 266
griseus, Pimelodus (Pimelodella), 191
grixalvii, Astroblepus, 63, 64
grixalvii micrescens, Astroblepus, 65
Gronias, 204
Gronias nigrilabris, 204, 206
Gronovii, Galeichthys, 37
gronovii, Aspredo, 58, 60
grosskopfii, Pimelodus, 338

FERRARIS

grosskopfii, Pimelodus (Pimelodus),
338
grosskopfii navarroi, Pimelodus, 338
grunniens, Pimelodus, 336
grypus, Anadoras, 169
grypus, Doras, 169
guacamaya, Typhlobelus, 426
guacari, Hypostomus, 250, 258
guacharote, Ancistrus, 225
guacharote, Hypostomus, 265
guacharote, Lasiancistrus, 265
guahiborum, Hemiancistrus, 243
guairense, Chaetostoma, 228
guairensis, Chaetostomus, 228
guairensis, Rhamdia, 198
guapore, Corydoras, 118
guapore, Lasiancistrus, 265
guaraquessaba, Trichomycterus, 418
guaricensis, Farlowella, 240
guasarensis, Rhamdia, 197
guatemalensis, Ariopsis, 33
guatemalensis, Arius, 33
guatemalensis, Pimelodus, 200
guatemalensis decolor, Rhamdia, 202
guatemalensis muriei, Rhamdia, 202
guatemalensis stygaea, Rhamdia, 202
guayaberensis, Medemichthys, 188
guayaberensis, Ituglanis, 407
guayaberensis, Pygidium metae, 407
guayoensis, Deltadoras, 173
guayoensis, Megalodoras, 173
guentheri, Ancistrus, 287
guentheri, Astroblepus, 64
guentheri, Atopochilus, 308
guentheri, Clarias, 143
guentheri, Euchilichthys, 309
guentheri, Galeichthys, 33
guentheri, Hypoptopoma, 250
guentheri, Microlepidogaster, 296
guentheri, Pseudancistrus, 287
guentheri, Oxyloricaria, 299
guentheri, Sturisoma, 299
guentheri, Stygogenes, 64
Guentheri, Synodontis, 309
guianense, Hypoptopoma, 250
guianense, Pygidium, 418
guianensis, Ageneiosus 70
guianensis, Corydoras, 118
guianensis, Harttia, 241
guianensis, Pygidium, 418
guianensis, Trichomycterus, 418
guineensis, Clarias, 141
Guirali, Pimelodus, 162
Guiritinga, 43
gulare, Hypoptopoma, 249, 250
gulio, Bagrus, 93
gulio, Mystus, 95
gulio, Pimelodus, 93, 95
gulioides, Bagrus, 95

gulosus, Tachisurus, 40
guntheri, Microlepidogaster, 295
guntheri, Schizolecis, 295
güntheri, Atopochilus, 309
güntheri, Ancistrus, 287
güntheri, Microlepidogaster, 296
guppyi, Pseudauchenipterus, 76
gurgu, Silurus, 431
guttata, Synodontis, 315
guttatus, Acrochordonichthys, 12
guttatus, Hemibagrus, 88
guttatus, Hypostomus, 287
guttatus, Imparfinis, 185
guttatus, Nannorhamdia, 185
guttatus, Parauchenoglanis, 163
guttatus, Pimelodus, 88, 162, 163
guttatus, Synodontis, 315
Guyanancistrus, 287
Gymnallabes, 137, 150
Gymnallabes alvarezi, 150
Gymnallabes apus, 137
Gymnallabes nops, 150
Gymnallabes tihoni, 152
Gymnallabes typus, 150
Gymnallabes typus forma
heterocercalis, 150
Gymnallabes typus heterocercalis, 150
gymnogaster, Loricaria, 297
gymnogaster, Spatuloricaria, 297
gymnogaster lagoichthys, Loricaria,
297
gymnorhynchus, Ancistrus, 221
gymnorhynchus, Gephyroglanis, 161
gymnorhynchus, Hypostomus, 254
gymnorhynchus, Plecostomus, 254
gymnorhynchus occidentalis,
Hypostomus, 257
gymnorhynchus tapanahoniensis,
Hypostomus, 261
gyrina, Tatia, 77
Gyrinurus, 409
Gyrinurus batrachostoma, 409
gyrinus, Acrochordonichthys 12
gyrinus, Centromochlus, 77
gyrinus, Noturus, 213
gyrinus, Silurus, 211, 213, 214Hhabereri, Chrysichthys, 156
habereri, Euchilichthys, 309
habereri, Gephyroglanis, 161
habrosus, Corydoras, 118
Haemomaster, 405
Haemomaster venezuelae, 405
haemomyzon, Homodiaetus, 412
haemomyzon, Pseudostegophilus, 412
haggini, Bunocephalus, 59
hahni, Farlowella, 238
hainanensis, Glyptosternon, 390
hainanensis, Glyptothorax, 390

CHECKLIST OF CATFISHES, RECENT AND FOSSIL

hainanensis, Glyptothorax fukiensis,
390
hainanensis, Hemibagrus, 88
hainanensis, Leiocassis, 88
hainesi, Amissidens, 32
hainesi, Arius, 32
Halepensis, Bagrus, 93, 94, 96, 97
hamiltonii, Chaca, 135, 136
hamiltonii, Plotosus (Clarias), 140
hamiltonis, Arius, 51
hammarlundi, Hemiancistrus, 243
Hammondii, Pimelodus, 210
Hancockii, Doras, 177
Haplodoras, 173
Hara, 382, 395, 444
Hara buchanani, 395
Hara elongata, 384
Hara filamentosa, 395
Hara hara, 395, 396
Hara horai, 396
Hara jerdoni, 396
Hara Malabarica, 96
Hara saharsai, 396
Hara serrata, 396
Hara serratus, 396
hara, Hara, 396
hara, Pimelodus, 395, 396
haraldschultzi, Corydoras, 118
harbinger, Chiloglanis, 306
hardenbergi, Arius, 36
hardenbergi, Aspistor, 36
Hardwickii, Acanthonotus, 356
Hargeri, Amphilius, 23
hargreavesi, Farlowella, 238
Harmandi, Hemiarius, 31
haroldoi, Parotocinclus, 282
hartii, Pimelodella, 191
Hartii, Pimelodus (Pseudorhamdia),
191
Harttia, 217, 240, 440
Harttia caquetae, 300
Harttia carvalhoi, 240
Harttia crassicauda, 242
Harttia depressa, 240
Harttia dissidens, 241
Harttia duriventris, 241
Harttia filamentissima, 264
Harttia filamentosa, 263, 264
Harttia garavelloi, 241
Harttia gracilis, 241
Harttia guianensis, 241
Harttia kronei, 241
Harttia leiopleura, 240, 241
Harttia longipinna, 241
Harttia loricariformis, 240, 241
Harttia merevari, 241
Harttia microps, 290
Harttia nijsseni, 272
Harttia novalimensis, 241

Zootaxa 1418 © 2007 Magnolia Press · 579

Harttia punctata, 241
Harttia rhombocephala, 242
Harttia surinamensis, 242
Harttia torrenticola, 242
Harttia trombetensis, 242
Harttia uatumensis, 242
Harttiella, 242
Harttiella crassicauda, 242
Harttiinae, 217
hartwelli, Pimelodella, 192
hasemani, Auchenipterus(Pseudepapterus), 76
hasemani, Centrodoras, 170
hasemani, Farlowella, 238
hasemani, Hemidoras, 172
hasemani, Hemiloricaria, 245
hasemani, Imparfinis, 185
hasemani, Leptodoras, 172
hasemani, Otocinclus, 274
hasemani, Oxydoras (Rhinodoras)amazonum, 170
hasemani, Pimelodella, 192
hasemani, Pseudepapterus 76
hasemani, Pygidium, 418
hasemani, Rineloricaria, 245
hasemani, Trichomycterus, 418
hasemani, Vandellia, 411
Hassar, 171
Hassar affinis, 171
Hassar iheringi, 171
Hassar orestis, 171
Hassar praelongus, 173
Hassar ucayalensis, 171
Hassar wilderi, 171
Hassar woodi, 171
Hasselquistii, Clarias, 140
Hasselquistii, Schilbe, 365
hasseltii, Silurichthys, 377
hassleriana, Netuma, 56
hastatus, Arius, 105
hastatus, Corydoras, 112, 118
Hatcheria, 405
Hatcheria bullocki, 404
Hatcheria macraei, 405
Hatcheria Maldonadoi, 403, 404
Hatcheria patagoniensis, 405
Hatcheria pique, 405
Hatcheria titcombi, 405
haugi, Synodontis, 315
Haustor, 208
Haustor ochoterenai, 209
hauxwelli, Loricariichthys, 270
havmolleri, Batasio, 85
havmolleri, Mystus, 85
hazenensis, Ameiurus, 204
hazenensis, Ictalurus (Amiurus), 204
Heckelii, Arius, 55
heckelii, Auchenipterus, 73
heckelii, Centromochlus, 73

Heckelii, Doras, 178
heckelii, Scorpiodoras, 178
Helicophagus, 324
Helicophagus hypophthalmus, 325
Helicophagus leptorhynchus, 324
Helicophagus typus, 324, 325
Helicophagus waandersii, 325
helicophagus, Chrysichthys 156
helicophilus, Doras, 177
Helogenes, 130, 134
Helogenes amazonae, 134
Helogenes castaneus, 134
Helogenes gouldingi, 134
Helogenes marmoratus, 134
Helogenes marmoratus uruyensis, 134
Helogenes unidorsalis, 134
Helogenes uruyensis, 134
Helogenidae, 130
hematophaga, Vandellia, 410
Hemiancistrus, 218, 242
Hemiancistrus albocinctus, 223
Hemiancistrus annectens, 242
Hemiancistrus arenarius, 284
Hemiancistrus aspidolepis, 242
Hemiancistrus braueri, 284
Hemiancistrus brevis, 284
Hemiancistrus caquetae, 265
Hemiancistrus castelnaui, 265
Hemiancistrus chlorostictus, 243
Hemiancistrus daguae, 231
Hemiancistrus fugleri, 243
Hemiancistrus fuliginosus, 243
Hemiancistrus guahiborum, 243
Hemiancistrus hammarlundi, 243
Hemiancistrus holostictus, 243
Hemiancistrus landoni, 243
Hemiancistrus longipinnis, 227
Hemiancistrus macrops, 243
Hemiancistrus maracaiboensis, 243
Hemiancistrus mayoloi, 264
Hemiancistrus medians, 243
Hemiancistrus megacephalus, 244
Hemiancistrus megalopteryx, 244
Hemiancistrus meizospilos, 244
Hemiancistrus micrommatos, 244
Hemiancistrus niceforoi, 257
Hemiancistrus niger, 287
Hemiancistrus platyrhynchus, 232
Hemiancistrus punctulatus, 244
Hemiancistrus spilomma, 244
Hemiancistrus spinosissimus, 244
Hemiancistrus subviridis, 244
Hemiancistrus ucayalensis, 283, 285
Hemiancistrus votouro, 244
Hemiancistrus wilsoni, 244
Hemiarius, 44
Hemiarius dioctes, 44
Hemiarius harmandi, 31
Hemiarius insidiator, 44

580 · Zootaxa 1418 © 2007 Magnolia Press

Hemiarius sona, 44
Hemiarius stormi, 45
Hemiarius stormii, 45
Hemiarius verrucosus, 45
Hemibagrus, 87
Hemibagrus baramensis, 87
Hemibagrus bongan, 87
Hemibagrus caveatus, 87
Hemibagrus centralus, 87
Hemibagrus chrysops, 87
Hemibagrus elongatus hongus, 89
Hemibagrus filamentus, 88
Hemibagrus fortis, 88
Hemibagrus furcatus, 88
Hemibagrus gracilis, 88
Hemibagrus guttatus, 88
Hemibagrus hainanensis, 88
Hemibagrus hoevenii, 88
Hemibagrus hongus, 89
Hemibagrus imbrifer, 89
Hemibagrus johorensis, 89
Hemibagrus macropterus, 87, 89, 91
Hemibagrus major, 89
Hemibagrus maydelli, 89
Hemibagrus menoda, 89
Hemibagrus microphthalmus, 90
Hemibagrus nemurus, 90
Hemibagrus olyroides, 90
Hemibagrus peguensis, 90
Hemibagrus planiceps, 90
Hemibagrus pluriradiatus, 90
Hemibagrus punctatus, 91
Hemibagrus sabanus, 91
Hemibagrus spilopterus, 91
Hemibagrus taphrophilus, 108
Hemibagrus variegatus, 91
Hemibagrus velox, 91
Hemibagrus vietnamicus, 91
Hemibagrus vietnammicus, 91
Hemibagrus wyckii, 91
Hemibagrus wyckioides, 91
Hemicetopsis, 131
Hemicetopsis amphiloxus, 131
Hemicetopsis macilentus, 133
Hemicetopsis minutus, 130
Hemicetopsis morenoi, 130
Hemicetopsis othonops, 132
hemicochliodon, Hypostomus, 254
Hemidoradinae, 166
Hemidoras, 171
Hemidoras (Leptodoras) boulengeri,
175
Hemidoras boulengeri, 175
Hemidoras hasemani, 172
Hemidoras leporhinus, 174
Hemidoras micropoeus, 171
Hemidoras microstomus, 179
Hemidoras morrisi, 172
Hemidoras notospilus, 171

FERRARISHemidoras paraguayensis, 179
Hemidoras stenopeltis, 172
Hemileiocassis, 91
Hemileiocassis panjang, 91
Hemiloricaria, 244
Hemiloricaria altipinnis, 245
Hemiloricaria aurata, 245
Hemiloricaria beni, 245
Hemiloricaria cacerensis, 245
Hemiloricaria caracasensis, 244, 245
Hemiloricaria caracassensis, 245
Hemiloricaria castroi, 245
Hemiloricaria eigenmanni, 245
Hemiloricaria fallax, 245
Hemiloricaria formosa, 245
Hemiloricaria hasemani, 245
Hemiloricaria hoehnei, 245
Hemiloricaria jubata, 246
Hemiloricaria konopickyi, 246
Hemiloricaria lanceolata, 246
Hemiloricaria magdalenae, 246
Hemiloricaria melini, 246
Hemiloricaria morrowi, 246
Hemiloricaria nigricauda, 246
Hemiloricaria parva, 246
Hemiloricaria phoxocephala, 246
Hemiloricaria platyura, 246
Hemiloricaria sneiderni, 246
Hemiloricaria stewarti, 246
Hemiloricaria teffeana, 247
Hemiloricaria wolfei, 247
Hemiodon, 292
Hemiodon acipenserinus, 247
Hemiodon depressus, 292, 293
Hemiodon platycephalus, 288
Hemiodontichthyina, 217
Hemiodontichthys, 217, 247
Hemiodontichthys acipenserinus, 247
hemioliopterus, Phractocephalus, 336
hemioliopterus, Silurus, 336
hemipeltis, Nemadoras, 174
hemipeltis, Opsodoras, 174
hemiphractus, Callichthys, 111
Hemipimelodinae, 30
Hemipimelodus, 30, 41
Hemipimelodus aaldereni, 32
Hemipimelodus atripinnis, 34
Hemipimelodus bernhardi, 39
Hemipimelodus bicolor, 34
Hemipimelodus cochlearis, 43
Hemipimelodus colcloughi, 51
Hemipimelodus crassilabris, 42
Hemipimelodus daugueti, 43
Hemipimelodus dayi, 46
Hemipimelodus intermedius, 42
Hemipimelodus macrocephalus, 41
Hemipimelodus macrorhynchus, 32
Hemipimelodus papillifer, 48
Hemipimelodus siamensis, 41

Hemipimelodus sundanensis, 56
Hemipimelodus taylori, 48
Hemipimelodus velutinus, 48
Hemiplatystoma, 341
Hemipsilichthys, 247, 444
Hemipsilichthys azygolechis, 279
Hemipsilichthys calmoni, 279
Hemipsilichthys cameroni, 279
Hemipsilichthys cerosus, 279
Hemipsilichthys duseni, 262, 279
Hemipsilichthys eurycephalus, 279
Hemipsilichthys garbei, 279
Hemipsilichthys gobio, 247
Hemipsilichthys hypselurus, 280
Hemipsilichthys hystrix, 280
Hemipsilichthys mutuca, 280
Hemipsilichthys nimius, 247, 444
Hemipsilichthys nudulus, 280
Hemipsilichthys papillatus, 248, 444
Hemipsilichthys regani, 280
Hemipsilichthys splendens, 280
Hemipsilichthys steindachneri, 280
Hemipsilichthys stephanus, 280
Hemipsilichthys stomias, 281
Hemipsilichthys vestigipinnis, 281
Hemisilurus, 368
Hemisilurus heterorhynchus, 368
Hemisilurus mekongensis, 368
Hemisilurus moolenburghi, 369
Hemisilurus schilbeides, 371
Hemisilurus scleronema, 368
Hemisorubim, 333
Hemisorubim platyrhynchos, 333
Hemisynodontis, 309
Hemisynodontis membranacea, 309
hemiurus, Hypostomus, 254
hemiurus, Plecostomus, 254
hendricksoni, Akysis, 14
henni, Hexanematichthys, 56
Henonemus, 405
Henonemus intermedius, 405
Henonemus macrops, 406
Henonemus panzeri, 414
Henonemus punctatus, 406
Henonemus taxistigma, 406
Henonemus triacanthopomus, 440
henrici, Anopleutropius, 165
henrici, Cranoglanis, 165
henriquei, Farlowella, 238
henryi, Aoria, 100
henselii, Loricaria, 294
henselii, Rineloricaria, 294
Heptapteridae, 180
Heptapterinae, 180
Heptapterus, 180, 184
Heptapterus anisurus, 188
Heptapterus bleekeri, 184
Heptapterus collettii, 99
Heptapterus eigenmanni, 184

CHECKLIST OF CATFISHES, RECENT AND FOSSIL

Heptapterus fissipinnis, 184
Heptapterus multiradiatus, 184
Heptapterus mustelinus, 184
Heptapterus ñssipinnis, 184
Heptapterus ornaticeps, 185
Heptapterus somnians, 189
Heptapterus stewarti, 184
Heptapterus surinamensis, 183
Heptapterus sympterygium, 184
Heptapterus tapanahoniensis, 184
Heptapterus tenuis, 184
heraldoi, Pimelodus, 338
herberti, Ituglanis, 407
Herberti, Trichomycterus, 407
Herklotsella, 375
Herklotsella anomala, 375
hermanni, Hypostomus, 254
Hermanni, Plecostomus, 254
herzbergii, Sciades, 53
Herzbergii, Silurus, 53
herzensteini, Leiocassis, 92
herzensteini, Macrones, 92
hesperius, Kryptopterus, 370
heteracantha, Rhamdia, 201
heteracanthus, Chaetostomus, 264,265
heteracanthus, Lasiancistrus, 265
Heterobagrus, 94
Heterobagrus bocourti, 94
Heterobranchoides, 139
Heterobranchus, 136, 150
Heterobranchus 5-tentaculatus, 197
Heterobranchus 6-tentaculatus, 197,
200
Heterobranchus austriacus, 382
Heterobranchus bidorsalis, 150
Heterobranchus boulengeri, 151
Heterobranchus Geoffroyi, 150
Heterobranchus intermedius, 150
Heterobranchus isopterus, 151
Heterobranchus laticeps, 151
Heterobranchus longifilis, 151
Heterobranchus macronema, 151
Heterobranchus palæindicus, 151
Heterobranchus platycephalus, 151
Heterobranchus Senegalensis, 150
Heterobranchus sextentaculatus, 197,
200
Heterobranchus tapeinopterus, 149,
150
heterocephalus, Clariallabes, 137
heterocercalis, Gymnallabes typus,
150
heterodon, Arges, 64
heterodon, Astroblepus, 64
heterodon, Leporacanthicus, 266
heterodontum, Pygidium, 418
heterodontus, Trichomycterus, 418
heteromorphus, Corydoras, 118

Zootaxa 1418 © 2007 Magnolia Press · 581

heteropleura, Brachyrhamdia, 181
heteropleurus, Pimelodus, 181
Heteropneustes, 136, 151
Heteropneustes fossilis, 151
Heteropneustes kemratensis, 152
Heteropneustes longipectoralis, 152
Heteropneustidae, 136
heteroptera, Rineloricaria, 293, 294
heterorhynchus, Ancistrus, 221
heterorhynchus, Hemisilurus, 368
heterorhynchus, Wallago, 368
heterorhynchus, Xenocara, 221
heterurus, Akysis, 14
heterurus, Bagrus, 97
heterurus, Zaireichthys, 28
Heudelotii, Arius, 31
heuglini, Clarotes, 160
heward-belli, Arius, 56
hexacicinnus, Macropteronotus, 148
hexadactylus, Silurus, 61
hexanema, Laides, 358
hexanema, Pangasius, 358
hexanema, Silurodon, 380, 381
Hexanematichthys, 45
Hexanematichthys henni, 56
Hexanematichthys hymenorrhinos, 54
Hexanematichthys leptaspis, 47
Hexanematichthys leptocassis, 45
Hexanematichthys mastersi, 45
Hexanematichthys sagor, 45
Hexanematichthys sundaicus, 45
Hexanematichthys surinamensis, 33
hexapterus, Micronema, 371
hexapterus, Silurus 371
heylandi, Kronichthys, 263
heylandi, Plecostomus, 263
Hilarii, Pimelodus, 200
Hildadoras, 175
Hildadoras bolivarensis, 168
Hildadoras orinocensis, 175, 176
hildae, Amarginops, 153
hildae, Chrysichthys, 153
hildebrandi, Noturus, 213
hildebrandi, Schilbeodes, 213
hildebrandi lautus, Noturus, 213
hilgendorfi, Clarias, 139
hilli, Clarias, 144
hirsuta, Microsynodontis, 310
hirsutus, Liocassus, 107
hirsutus, Microsynodontis, 310
Hisonotus, 248
Hisonotus candombe, 440
Hisonotus charrua, 440
Hisonotus depressicauda, 248
Hisonotus depressinotus, 248
Hisonotus francirochai, 248
Hisonotus insperatus, 248
Hisonotus laevior, 248
Hisonotus leptochilus, 248

Hisonotus leucofrenatus, 248
Hisonotus maculipinnis, 248
Hisonotus nigricauda, 248
Hisonotus notatus, 248
Hisonotus paulinus, 249
Hisonotus ringueleti, 249
Hisonotus taimensis, 249
histrix, Pseudacanthicus, 286
histrix, Rinelepis, 218, 286
Hito, 375
Hito taytayensis, 375, 377
Hitoichthys, 375
Hitoichthys taytayensis, 375, 377
hodgarti, Euchiloglanis, 398
hodgarti, Glyptosternum, 398
hodgarti, Parachiloglanis, 398
hoedemani, Exastilithoxus, 237
hoehnei, Hemiloricaria, 245
hoehnei, Loricaria, 245
hoehnei, Nannoglanis, 188
hoehnei, Phenacorhamdia, 188
hoeksi, Pangasius, 328
hoerdzanicus, Eobagrus. 87
hoerdzanicus, Gobibagrus, 87
hoevenii, Bagrus, 88
hoevenii, Hemibagrus, 88
Hoffstetterichthys pucai, 30
hoi, Aoria, 107
holdeni, Oxydoras, 176
hollandi, Imparfinis, 185
Hollyi, Clarias, 144
hollyi, Clarias, 147
Hollyi, Synodontis, 323
Hollyi ntemensis, Synodontis, 322
holobranchus, Xenoclarias, 153
holomelas, Pimelodus, 199
holomelas rupununi, Rhamdia, 197
holopercnus, Synodontis, 315
holostictus, Hemiancistrus, 243
Homodiaetus, 406
Homodiaetus anisitsi, 406
Homodiaetus banguela, 406
Homodiaetus graciosa, 406
Homodiaetus haemomyzon, 412
Homodiaetus passarellii, 406
Homodiaetus vazferreirai, 406
homodon, Arges, 64
homodon, Astroblepus, 64
hondae, Hypostomus, 254
hondae, Plecostomus, 250, 254
honghenensis, Glyptothorax fukiensis,
390
honghensis, Glyptothorax, 390
honghensis, Glyptothorax fukiensis,
390
hongus, Hemibagrus, 89
hongus, Hemibagrus elongatus, 89
Hopladelus, 215
Hopliancistrini, 217

582 · Zootaxa 1418 © 2007 Magnolia Press

Hopliancistrus, 217, 249
Hopliancistrus tricornis, 249
Hoplisoma, 111
Hoplodaros, 173
Hoplodoras, 173
Hoplodoras ramirezi, 174
hoplogenys, Ancistrus, 221
hoplogenys, Chætostomus, 221
Hoplomizoninae, 57
Hoplomyzon, 57, 61
Hoplomyzon atrizona, 61
Hoplomyzon atrizona petroleus, 61
Hoplomyzon megistus, 60
Hoplomyzon papillatus, 61
Hoplomyzon sexpapilostoma, 61
Hoplomyzontinae, 57
hoplonites, Hypostomus, 254
Hoplosoma aeneum, 112
Hoplosterninae, 108
Hoplosternum, 108, 127
Hoplosternum littorale, 127
Hoplosternum littorale daillyi, 128
Hoplosternum magdalenae, 128
Hoplosternum oronocoi, 129
Hoplosternum punctatum, 127, 128
Hoplosternum schreineri, 128
Hoplosternum shirui, 128
Hoplosternum stevardii, 128
Hoplosternum thoracatum cayennae,
128
Hoplosternum thoracatum
surinamensis, 129
hoppei, Otocinclus, 275
Horabagrinae, 441
Horabagrus, 356, 429, 441
Horabagrus brachysoma, 429
Horabagrus nigricollaris, 429
horae, Amblyceps, 99
horae, Olyra, 99
Horaglanidinae, 441
Horaglanis, 152, 441
Horaglanis alikunhii, 152
Horaglanis krishnai, 152
horai, Glyptothorax, 390
horai, Glyptothorax, 390
horai, Hara, 396
horai, Mystus, 95
horai, Mystus (Mystus) vittatus, 95
horai, Olyra, 99
horai, Pteroglanis, 387, 390
horai, Socnopaea, 431
Horiomyzon, 185
Horiomyzon retropinnatus, 185
horrida, Squaliforma, 298
horridus, Hypostomus, 297, 298
horridus, Plotosus, 350
Hosii, Liocassis, 93
houghi, Pimelodus, 210
housei, Glyptothorax, 390

FERRARIS

howesi, Pimelodella, 192
howong, Macrones, 88
hoyi, Pimelodus, 206
huaorani, Otocinclus, 275
huberi, Oxydoras (Rhinodoras), 174
humboldti, Doras, 176
Humboldtii, Cyclopium, 63, 64
humboldtii, Zungaro, 344
humeralis, Doras, 174
humeralis, Nemadoras, 174
humeralis, Pangasius, 326
humeratus, Synodontis, 321
humilis, Pimelodus, 198
humilis, Rhamdia, 198
hwanghoensis, Leiocassis, 107
Hyalobagrus, 92
Hyalobagrus flavus, 92
Hyalobagrus leiacanthus, 92
Hyalobagrus ornatus, 92hydrostaticus, Pleurophysus, 410
hymenorrhinos, Hexanematichthys, 54
Hypancistrus, 249
Hypancistrus inspector, 249
Hypancistrus zebra, 249
Hypocolpterus, 227
Hypocolpterus analis, 227
Hypodoras, 172
Hypodoras forficulatus, 172
Hypophthalmini, 329
Hypophthalmus, 329, 333
Hypophthalmus devall, 334
Hypophthalmus edentatus, 333
Hypophthalmus fimbriatus, 333
Hypophthalmus goongwaree, 362
Hypophthalmus longifilis, 334
Hypophthalmus marginatus, 333
Hypophthalmus niloticus, 362, 365
Hypophthalmus nuchalis, 71, 72
Hypophthalmus oremaculatus, 333
Hypophthalmus perporosus, 334
Hypophthalmus Spixii, 333
Hypophthalmus taakree, 362
hypophthalmus, Helicophagus, 325
hypophthalmus, Arius, 39, 40
hypophthalmus, Cathorops, 40
hypophthalmus, Pangasianodon, 325
hypophthalmus, Ompok, 371, 372
hypophthalmus, Silurus, 372
Hypoptopoma, 217, 249
Hypoptopoma bilobatum, 250
Hypoptopoma carinatum, 276
Hypoptopoma guentheri, 250
Hypoptopoma guianense, 250
Hypoptopoma gulare, 249, 250
Hypoptopoma inexpectata, 250
Hypoptopoma inexspectatum, 250
Hypoptopoma joberti, 250
Hypoptopoma psilogaster, 250
Hypoptopoma steindachneri, 250

Hypoptopoma thoracatum, 249, 250
Hypoptopomatinae, 217
Hypoptopominae, 217
Hypostoma etentaculatum, 291
Hypostoma punctatum, 220
Hypostoma squalinum, 298
Hypostomiden, 216
Hypostomus, 216, 217, 250
Hypostomus aburrensis, 227
Hypostomus affinis, 251
Hypostomus agna, 251
Hypostomus alatus, 251
Hypostomus albopunctatus, 251
Hypostomus ancistroides, 251
Hypostomus angipinnatus, 251
Hypostomus argus, 251
Hypostomus asperatus, 251
Hypostomus aspilogaster, 251
Hypostomus atropinnis, 251
Hypostomus aurantiacus, 278
Hypostomus auroguttatus, 251
Hypostomus barbatus, 286, 287
Hypostomus bolivianus, 252
Hypostomus borellii, 252
Hypostomus boulengeri, 252
Hypostomus brevicauda, 252
Hypostomus brevis, 252
Hypostomus brevitentaculatus, 291
Hypostomus bufonius, 219
Hypostomus calamita, 219
Hypostomus carinatus, 252
Hypostomus carvalhoi, 252
Hypostomus cirrhosus, 219, 220
Hypostomus cochliodon, 250, 252,
253-262
Hypostomus commersoni, 252
Hypostomus coppenamensis, 253
Hypostomus corantijni, 253
Hypostomus crassicauda, 253
Hypostomus derbyi, 253
Hypostomus dlouhyi, 253
Hypostomus duodecimalis, 290, 291
Hypostomus emarginatus, 297
Hypostomus eptingi, 253
Hypostomus ericae, 253
Hypostomus ericius, 253
Hypostomus erinaceus, 221
Hypostomus fluviatilis, 253
Hypostomus fonchii, 253
Hypostomus francisci, 254
Hypostomus garmani, 254
Hypostomus goyazensis, 254
Hypostomus granosus, 273
Hypostomus guacari, 250, 258, 259
Hypostomus guacharote, 265
Hypostomus guttatus, 287
Hypostomus gymnorhynchus, 254
Hypostomus gymnorhynchus
occidentalis, 257

CHECKLIST OF CATFISHES, RECENT AND FOSSIL

Hypostomus gymnorhynchus
tapanahoniensis, 261
Hypostomus hemicochliodon, 254
Hypostomus hemiurus, 254
Hypostomus hermanni, 254
Hypostomus hondae, 254
Hypostomus hoplonites, 254
Hypostomus horridus, 297, 298
Hypostomus iheringii, 255
Hypostomus interruptus, 255
Hypostomus isbrueckeri, 255
Hypostomus itacua, 217
Hypostomus jaguribensis, 255
Hypostomus johnii, 255
Hypostomus laplatae, 255
Hypostomus latifrons, 255
Hypostomus latirostris, 255
Hypostomus levis, 255
Hypostomus lexi, 255
Hypostomus lima, 256
Hypostomus longiradiatus, 256
Hypostomus luteomaculatus, 256
Hypostomus luteus, 256
Hypostomus macrophthalmus, 256
Hypostomus macrops, 256
Hypostomus macushi, 256
Hypostomus margaritifer, 256
Hypostomus meleagris, 256
Hypostomus micromaculatus, 256
Hypostomus microstomus, 250, 257
Hypostomus multiradiatus, 290, 291
Hypostomus mutucae, 257
Hypostomus myersi, 257
Hypostomus nematopterus, 257
Hypostomus niceforoi, 257
Hypostomus nickeriensis, 257
Hypostomus niger, 257
Hypostomus nigricans, 278
Hypostomus nigromaculatus, 257
Hypostomus niveatus, 226, 227
Hypostomus nudiceps, 223
Hypostomus nudiventris, 257
Hypostomus occidentalis, 257
Hypostomus oculeus, 257
Hypostomus pagei, 257
Hypostomus pantherinus, 258
Hypostomus papariae, 258
Hypostomus paranensis, 258
Hypostomus pardalis, 292
Hypostomus paucimaculatus, 258
Hypostomus paucipunctatus, 258
Hypostomus paulinus, 258
Hypostomus pictus, 265
Hypostomus piratatu, 258
Hypostomus plecostomoides, 258
Hypostomus plecostomus, 258
Hypostomus pseudohemiurus, 259
Hypostomus pseudohemiurus
macrophthalmus, 256

Zootaxa 1418 © 2007 Magnolia Press · 583

Hypostomus punctatus, 220, 259
Hypostomus pusarum, 259
Hypostomus pyrineusi, 259
Hypostomus regani, 259
Hypostomus robinii, 259
Hypostomus rondoni, 259
Hypostomus roseopunctatus, 259
Hypostomus saramaccensis, 259
Hypostomus scabriceps, 260
Hypostomus scaphyceps, 260
Hypostomus sculpodon, 260
Hypostomus seminudus, 260
Hypostomus serratus, 286
Hypostomus simios, 260
Hypostomus sipaliwinii, 260
Hypostomus soniae, 260
Hypostomus spinosus, 286
Hypostomus strigaticeps, 260
Hypostomus subcarinatus, 260
Hypostomus surinamensis, 260
Hypostomus tapanahoniensis, 260
Hypostomus taphorni, 261
Hypostomus tapijara, 261
Hypostomus Temminckii, 224
Hypostomus tenuis, 298
Hypostomus ternetzi, 261
Hypostomus tietensis, 261
Hypostomus topavae, 261
Hypostomus unae, 261
Hypostomus uruguayensis, 261
Hypostomus vaillanti, 261
Hypostomus variipictus, 261
Hypostomus varimaculosus, 261
Hypostomus variostictus, 261
Hypostomus ventromaculatus, 262
Hypostomus vermicularis, 262
Hypostomus verres, 262
Hypostomus vicinus, 278
Hypostomus waiampi, 262
Hypostomus watwata, 262
Hypostomus winzi, 262
Hypostomus wuchereri, 262
Hypothalmus dawalla, 69
Hypselobagrus, 93
Hypselobagrus macronema, 97
hypselopterus, Bagrichthys, 82
hypselopterus, Bagrus, 82
hypselurus, Breitensteinia, 15
hypselurus, Hemipsilichthys, 280
hypselurus, Pareiorhaphis, 280
hypselurus, Pimelodus, 198
Hypsidoridae, 203
Hypsidoris, 203
Hypsidoris farsonensis, 203
Hypsidoris oregonensis, 203
hypsiura, Amaralia, 57
hypsiurus, Bunocephalus, 57
hypsiurus, Oreoglanis, 397
hyrtlii, Neosilurus, 347, 348

hystrix, Acanthicus, 218
hystrix, Hemipsilichthys, 280
hystrix, Pareiorhaphis, 280
Iichikawai, Coreobagrus, 86ichikawai, Pseudobagrus, 86
ichneumon, Coreobagrus, 431
Ichthaelurinae, 203
Ichthaelurus, 203, 208
Ichthaelurus Kevinskii, 204
Ichthaelurus McCaskei, 204
Ichthaelurus robustus, 210
Ichthyaelurus, 208
Ictaluri, 203
Ictaluridae, 203
Ictalurus, 203, 207
Ictalurus (Amiurus) hazenensis, 204
Ictalurus anguilla, 210
Ictalurus australis, 208
Ictalurus balsanus, 208
Ictalurus benderensis, 206
Ictalurus dugesii, 208, 209
Ictalurus echinatus, 208
Ictalurus furcatus, 208, 211
Ictalurus lambda, 209
Ictalurus lavetti, 204
Ictalurus leidyi, 204
Ictalurus lupus, 209, 211
Ictalurus macgrewi, 205
Ictalurus mexicanus, 209
Ictalurus nebulosus pannonicus, 206
Ictalurus ochoterenai, 209
Ictalurus okeechobeensis, 210
Ictalurus pectinatus, 206
Ictalurus peregrinus, 207
Ictalurus pricei, 209
Ictalurus punctatus, 208, 209, 211
Ictalurus rhaeas, 210
Ictalurus sawrockensis, 206
Ictalurus serracanthus, 206
Ictalurus simpsonii, 210
Ictalurus spodius, 211
Ictalurus vespertinus, 206
idenburgi, Copidoglanis, 348
idenburgi, Neosilurus, 348
ignobilis, Rhamdella, 196
iheringi, Arius, 343
iheringi, Cetopsorhamdia, 182
iheringi, Hassar, 171
iheringi, Microglanis, 354
iheringi, Plecostomus, 298
iheringi, Pygidium, 419
iheringi, Steindachneridion, 343
iheringi, Trichomycterus, 419
Iheringichthys, 334
Iheringichthys labrosus, 334
Iheringichthys megalops, 334
iheringii, Bunocephalus, 59
iheringii, Hypostomus, 255

584 · Zootaxa 1418 © 2007 Magnolia Press

iheringii, Plecostomus, 255, 298
ikapor, Plotoseus, 350
ikiensis, Pseudobagrus, 102
ilebrevis, Synodontis, 445
ilesi, Bathyclarias, 136
ilesi, Dinotopterus, 136
Ilictis, 215
imberbis, Silurus, 70, 379
imbrifer, Hemibagrus, 89
imitator, Brachyrhamdia, 181
imitator, Corydoras, 118
immaculata, Pseudecheneis, 400
immaculatum, Pygidium, 419
immaculatus, Auchenipterus, 80
immaculatus, Pseudecheneis, 400
immaculatus, Silurus (Callichrus), 374
immaculatus, Trichomycterus, 419
Imparales, 180, 188
Imparales mariai, 188
Imparales panamensis, 180
Imparfinis, 185
Imparfinis bolivianus, 188
Imparfinis borodini, 185
Imparfinis cochabambae, 185
Imparfinis guttatus, 185
Imparfinis hasemani, 185
Imparfinis hollandi, 185
Imparfinis insidiosus, 182
Imparfinis lineatus, 185
Imparfinis longicauda, 185
Imparfinis longicauda, 185
Imparfinis microps, 186
Imparfinis minutus, 186
Imparfinis mirini, 186
Imparfinis nemacheir, 186
Imparfinis pijpersi, 186
Imparfinis piperatus, 185, 186
Imparfinis pristos, 186
Imparfinis pseudonemacheir, 186
Imparfinis schubarti, 186
Imparfinis spurrellii, 186
Imparfinis stictonotus, 186
Imparfinis tenebrosus, 189
impluviatus, Mystus, 95
inaequalis, Amphilius
grammatophorus, 23
incae, Trichomycterus, 423
Incaichthys suarezi, 30
incolicana, Corydoras, 118
indicus, Brachyspondylus, 326
indicus, Glyptothorax, 390
indicus, Kryptopterus, 367
indicus, Pangasius, 326
indicus, Pimelodus, 19
indicus, Silurus, 374
indragiriensis, Silurichthys, 378
ineac, Phractura, 28
ineac, Trachyglanis, 28
inermis, Ageneiosus, 69

FERRARIS

inermis, Akysis, 16
inermis, Amblyceps, 17
inermis, Nematogenys, 324
inermis, Pseudobagarius, 16
inermis, Silurus, 69, 379
inermis, Trichomycterus, 324
inexpectata, Aristommata, 249, 250
inexpectata, Hypoptopoma, 250
inexspectatum, Hypoptopoma, 250
infraoculare, Sorubim, 342
infulatus, Oreoglanis, 397
infuscatus, Erethistoides, 442
ingluvies, Osteogeneiosus, 51
inornata, Pseudolaguvia, 401
inornatus, Liocassis, 103
inornatus, Pseudomystus, 103
inpai, Aguarunichthys, 329
insculptus, Arius, 50
insculptus, Doras, 179
insculptus, Notarius, 50
insidiator, Arius, 45
insidiator, Hemiarius, 44
insidiosa, Cetopsorhamdia, 182
insidiosus, Imparfinis, 182
insidiosus, Stegophilus, 414
insignarius, Pimelodon, 211, 213
insigne, Pimelodus, 213
insignis, Auchenipterus, 79
insignis, Breitensteinia, 15
insignis, Noturus, 213
insignis, Oreoglanis, 397
insignis, Pimelodella, 190
insignis, Pimelodus, 339
insignis, Trachelyopterus, 79
insignis badeli, Trachycorystes, 80
insignis peloichthys, Trachycorystes,80
insolitus, Clarias, 144
inspector, Hypancistrus, 249
insperatus, Hisonotus, 248
insularum, Netuma, 50
intermedia, Phractura, 26
intermedia, Tatia, 77
intermedium, Pseudoplatystoma
fasciatum, 341
intermedius, Centromochlus, 77intermedius, Clarias, 144intermedius, Hemipimelodus, 42intermedius, Henonemus, 405
intermedius, Heterobranchus, 150
intermedius, Pelteobagrus, 100intermedius, Pseudecheneis, 400
intermedius, Pseudobagrus, 100intermedius, Pseudotocinclus, 290
intermedius, Schilbe, 363
intermedius, Stegophilus, 405
intermedius, Trachyglanis, 28interruptus, Hypostomus, 255
interruptus, Plecostomus, 255

interspinalum, Glyptosternon, 390
interspinalus, Glyptothorax, 390
intonsus, Ernstichthys, 60
inusitata, Pterocryptis, 377
irsacae, Synodontis, 314
Irvineia, 358
Irvineia orientalis, 358
Irvineia voltae, 358
irwini, Megalodoras, 173, 174
isacanthus, Auchenipterus, 80
isbrueckeri, Farlowella, 238
isbrueckeri, Corydoras, 119
isbrueckeri, Farlowella, 238
isbrueckeri, Hypostomus, 255
Isbrueckerichthys, 262, 441
Isbrueckerichthys alipionis, 262
Isbrueckerichthys calvus, 441
Isbrueckerichthys duseni, 262
Isbrueckerichthys epakmos, 262
Isbrueckerichthys saxicola, 441
ischnosoma, Acrochordonichthys, 12
isheriensis, Clarias (Clarioides), 139
Isidori, Schilbe, 365
isopterus, Heterobranchus, 151
Isorineloricaria, 263
Isorineloricaria spinosissima, 263
isthmensis, Aelurichthys, 38
Istlarius, 208
Istlarius balsanus, 208
Istlarius balsanus occidentalis, 208
itacaiunas, Rhamdia, 198
itacambirussu, Trichomycterus, 419
itacarambiensis, Trichomycterus, 419
itacua, Hypostomus, 217
itaimbezinho, Epactionotus, 236
itapicuruensis, Pimelodella, 192
itatiayae, Trichomycterus, 419
itchkeea, Gagata, 386
itchkeea, Phractocephalus, 386
Ituglanis, 406
Ituglanis amazonicus, 406
Ituglanis bambui, 406
Ituglanis cahyensis, 444
Ituglanis eichorniarum, 407
Ituglanis epikarsticus, 407
Ituglanis gracilior, 407
Ituglanis guayaberensis, 407
Ituglanis herberti, 407
Ituglanis laticeps, 407
Ituglanis macunaima, 407
Ituglanis metae, 407
Ituglanis nebulosus, 407
Ituglanis parahybae, 407
Ituglanis parkoi, 407
Ituglanis passensis, 408
Ituglanis proops, 408
Ituglanis ramiroi, 408
ituriensis, Synodontis, 315
iturii, Auchenoglanis, 163

CHECKLIST OF CATFISHES, RECENT AND FOSSIL

iturii, Synodontis, 315
iwokrama, Denticetopsis, 133
Ixinandria, 263
Ixinandria montebelloi, 263
Ixinandria steinbachi, 263
izabalensis, Potamarius, 53
Jjacksonensis, Netuma thalassina, 49
jacksoni, Dinotopterus, 149
jacksonii, Amphilius, 21
jacupiranga, Trichomycterus, 419
jaekeli, Otolithus (Arius), 435
jaensis, Clarias, 144
jagur, Macropteronotus, 140
jaguribensis, Hypostomus, 255
jaguribensis, Plecostomus, 255
jahu, Paulicea, 344
jahu, Zungaro, 344
jalalensis, Glyptothorax, 390
jallae, Dinotopterus, 146
jallae, Synodontis, 316
jandia, Sorubim, 342, 343
janeirensis, Pseudotothyris, 290japonicus, Centranodon, 379
japonicus, Silurus, 378, 379
jaraguensis, Loricaria, 294
jaraguensis, Rineloricaria, 294
jataiensis, Ancistrus, 222
jatia, Cephalocassis, 41
jatius, Pimelodus, 41
jauruensis, Farlowella, 238
jauruënsis, Farlowella, 238
javanensis, Belodontichthys, 381
javanica, Rhamdia, 431
Javensis, Bagrus, 45
javus, Pimelodus, 431
jayarami, Myersglanis, 396
jaynei, Ompok, 372
jeanesianus, Liposarcus, 292
jegui, Chaetostoma, 228
jella, Arius, 36
jelskii, Ancistrus, 222
Jelskii, Chaetostomus, 222
jenynsii, Pimelodus, 196
jenynsii, Rhamdella, 196
jequitinhonha, Rhamdia, 198
Jequitinhonhae, Auchenipterus(Pseudauchenipterus), 76
jequitinhonhae, Pseudauchenipterus,
76
jequitinhonhae, Trichomycterus, 419
jerdoni, Hara, 396
jimi, Parotocinclus, 282
jivaro, Pimelodus, 338
joberti, Hypoptopoma, 250
joberti, Otocinclus, 250
johannae, Septobranchus, 42johnelsi, Chrysichthys, 156
johnii, Hypostomus, 255

Zootaxa 1418 © 2007 Magnolia Press · 585

Johnii, Plecostomus, 255
johnsoni, Pygidium, 419
johnsoni, Trichomycterus, 419
johorensis, Hemibagrus, 89
johorensis, Mystus, 89
jokeannae, Trachycorystes, 80
jordani, Tachisurus, 33
joselimai, Leporacanthicus, 266
joselimaianus, Glyptoperichthys, 291
joselimaianus, Pterygoplichthys, 291
juaro, Pangasius, 327
jubata, Hemiloricaria, 246
jubata, Loricaria, 246
julii, Corydoras, 119
jumbo, Parotocinclus, 282
juquiaae, Corydoras, 121
juquiae, Corydoras, 121
juquiae, Otothyris, 276
juquiae, Pseudotocinclus, 290
jurubidae, Astroblepus, 65
jurubidae, Cetopsis, 132
jurubidae, Pseudocetopsis, 132
juruense, Brachyplatystoma, 331
juruense, Platystoma, 331
juruensis, Leptodoras, 172
juvens, Pterygoplichthys, 291
Kkaiei, Corymbophanes, 232
kaifenensis, Leiocassis, 107
kajan, Macrones, 88
kakrimensis, Amphilius, 21
kalambo, Chiloglanis, 306
Kalyptodoras, 172
Kalyptodoras bahiensis, 172
kamengensis, Euchiloglanis, 399
kamengensis, Pareuchiloglanis, 399
kanei, Corydoras, 119
kanganamanensis, Arius, 39
kanpurensis, Pinniwallago, 375
kapuasensis, Clarias, 144
kapuri, Laguvia ribeiroi, 401
kapuri, Pseudolaguvia, 401
kasai, Eutropiellus, 360, 361
kashmirensis, Glyptothorax, 390, 393
katangae, Synodontis, 315
Keletius, Bagrus, 98
kelioides, Encheloclarias, 149
kempi, Olyra, 99
kemratensis, Clarisilurus, 151, 152
kemratensis, Heteropneustes, 152
kessleri, Arius, 50
kessleri, Notarius, 50
Ketengus, 45
Ketengus typus, 45
Kevinskii, Ichthaelurus, 204
khartoumensis, Synodontis, 316
khavalchor, Neotropius, 359
kinabatanganensis, Pangasius, 326
kingi, Liobagrus, 19

kingsleyae, Clarias, 144
kingsleyae, Chrysichthys, 160
kirkii, Arius, 52
kishinouyei, Coraglanis, 385
kishinouyei, Euchiloglanis, 385
kitsoni, Arius 56
kivuensis, Amphilius, 21
kneri, Callichthys, 111
kneri, Loricaria, 299
kneri, Oxydoras, 175
kneri, Sturisoma, 299
Knerii, Acestra, 238
knerii, Bunocephalus, 59
knerii, Farlowella, 238
knerii, Pimelodus (Rhamdia), 199
knerii, Trichomycterus, 419
koenigi, Bagrus, 84
koensis, Synodontis, 316
koepckei, Myoglanis, 187
konopickyi, Hemiloricaria, 246
konopickyi, Loricaria, 246
koreanus, Pseudobagrus, 102
krattensis, Wallago, 374
krefftii, Amphilius, 23
krempfi, Pangasius, 326
krishnai, Horaglanis, 152
krishnensis, Mystus, 89
kronei, Corydoras, 129
kronei, Harttia, 241
kronei, Loricaria, 294
kronei, Pimelodella, 192
kronei, Rineloricaria, 294
kronei, Typhlobagrus, 189, 192
Kronichthys, 263
Kronichthys heylandi, 263
Kronichthys lacerta, 263
Kronichthys subteres, 263
Kryptopterichthys, 369
Kryptopterichthys macrocephalus, 370
Kryptopterini, 367
Kryptopterus, 367, 369
Kryptopterus baramensis, 369
Kryptopterus bicirrhis, 369
Kryptopterus cheveyi, 371
Kryptopterus cryptopterus, 369
Kryptopterus deignani, 375
Kryptopterus dissitus, 369
Kryptopterus eugeneiatus, 369
Kryptopterus geminus, 369
Kryptopterus hesperius, 370
Kryptopterus indicus, 367
Kryptopterus lais, 370
Kryptopterus limpok, 370
Kryptopterus lumholtzi, 370
Kryptopterus macrocephalus, 370
Kryptopterus micropus, 369
Kryptopterus minimus, 370
Kryptopterus minor, 370
Kryptopterus mononema, 370

586 · Zootaxa 1418 © 2007 Magnolia Press

Kryptopterus moorei, 371
Kryptopterus palembangensis, 370
Kryptopterus paraschilbeides, 370
Kryptopterus parvanalis, 375
Kryptopterus piperatus, 370
Kryptopterus platypogon, 371
Kryptopterus sabanus, 370
Kryptopterus schilbeides, 371
kryptos, Xiliphius, 62
kuhlmanni, Peckoltia, 284
kuhlmanni, Peckoltichthys, 284
kükenthali, Glyptosternum, 391
kunyit, Pangasius, 326
kurdistanicum, Glyptosternum, 390
kurdistanicus, Glyptothorax, 390
kurzii, Akysis, 18
kutchensis, Arius, 56
kuturnee, Phractocephalus, 105
kuturnee, Rita, 105
kyphus, Pelteobagrus, 82
kyphus, Pseudobagrus, 82
Llaak, Glyptosternum, 395
laani, Apistoloricaria, 226
labeo, Synodontis, 323
labialis, Astroblepus, 65
labialis, Loricaria, 270
labialis, Loricariichthys, 270
labiatum, Exostoma, 385
labiatus, Glyptosternon, 385
labiosus, Copidoglanis, 345
labrax, Exostoma, 387
labrosus, Copidoglanis, 345
labrosus, Iheringichthys, 334
labrosus, Pimelodus, 334
labrosus, Xyliphius, 62
Lacantunia, 216
Lacantunia enigmatica, 216
Lacantuniidae, 216
lacerdai, Corydoras, 119
lacerta, Kronichthys, 263
lacerta, Plecostomus, 263
lachneri, Noturus, 213
lacus, Aoria, 108
lacustricolus, Synodontis, 322
lacustris, Auchenipterus, 79
lacustris, Trachelyopterus, 79
lacustris, Gadus, 208
laessoei, Synodontis, 316
laeviceps, Arius, 55
laeviceps, Clarias, 144
laeviceps dialonensis, Clarias, 142
laevigata, Microsynodontis, 310
laevigatulus, Doras, 177
laevigatus, Bagrus, 49
laevigatus, Callichthys, 127
laevigatus, Microsynodontis, 310
laevior, Hisonotus, 248
laevis, Gogangra, 395

FERRARIS

laevis, Platystacus, 58
laeviuscula, Loricaria, 289
laeviuscula, Pseudoloricaria, 289
lagoensis, Arius, 31
lagoensis, Chrysichthys, 160
lagoichthys, Loricaria gymnogaster,297
lagoichthys, Spatuloricaria, 297
Laguvia, 382, 395
Laguvia manipurensis, 382
Laguvia ribeiroi, 401
Laguvia ribeiroi kapuri, 401
Laguvia shawi, 395, 402
Laguviini, 382
Laides, 358
Laides hexanema, 358
Laides longibarbis, 359
Laimumena, 431
Laimumena barbonica, 431, 432
Lais, 358
lais, Kryptopterus, 370
Laïs, 358
laïs, Silurus, 370
lakoi, Aspidoras, 109
laluchensis, Rhamdia, 198
lamani, Amphilius, 22
Lamarrii, Bagrus, 106, 107
lambda, Ictalurus, 209
lamberti, Corydoras, 119
lamberti, Microsynodontis, 310
Lambertia, 347
Lambertia atra, 347
Lambertichthys, 347
Lambertichthys ater sepikensis, 347
lamghur, Silurus, 373
lamina, Loricaria, 288
laminus, Pseudohemiodon, 288
Lamontichthys, 263
Lamontichthys filamentosus, 263
Lamontichthys llanero, 264
Lamontichthys maracaibero, 264
Lamontichthys stibaros, 264
lamottei, Chiloglanis, 306
lamottei, Clarias, 144
lampei, Amphilius, 22
Lampiella, 217, 264
Lampiella gibbosa, 264
Lampiellini, 217
lampris, Glyptothorax, 391
lanceolata, Hemiloricaria, 246
lanceolata, Loricaria, 246
lanceolata, Rineloricaria, 246
lanceolatus, Nemuroglanis, 188
landanensis, Tachysurus, 57
landinga, Trichomycterus, 419
landoni, Hemiancistrus, 243
lanzhouensis, Silurus, 380
laosensis, Glyptothorax, 391
laplatae, Hypostomus, 255

laplatae, Plecostomus, 255
larai, Bunocephalus, 59
larnaudii, Pangasius, 326
Lasiancistrus, 264
Lasiancistrus anthrax, 288
Lasiancistrus brevispinis, 287
Lasiancistrus caucanus, 264
Lasiancistrus dumus, 289
Lasiancistrus guacharote, 265
Lasiancistrus guapore, 265
Lasiancistrus heteracanthus, 265
Lasiancistrus longispinis, 287
Lasiancistrus maracaiboensis, 265
Lasiancistrus mayoloi, 265
Lasiancistrus nationi, 223
Lasiancistrus nicoi, 289
Lasiancistrus saetiger, 265
Lasiancistrus schomburgkii, 265
Lasiancistrus scolymus, 265
Lasiancistrus tentaculatus, 265
Lasiancistrus tigris, 288, 289
Lasiancistrus volcanensis, 264
lata, Loricaria, 268
lateralis, Pimeletropis, 332
Lateralis, Pimelodus pallidus var., 210
Lateralis, Silurus pallidus, 210
lateristriga, Pimelodella, 192
lateristrigus, Pimelodes, 192
lateristrigus, Pimelodus, 192
laticauda, Rhamdia, 198, 199, 202
laticauda typhla, Rhamdia, 198
laticaudatus, Amphilius, 22
laticaudus, Pimelodus, 198
laticeps, Adelopeltis, 107
laticeps, Allabenchelys, 137
laticeps, Amblyceps, 17
laticeps, Arius, 41
laticeps, Chrysichthys, 157
laticeps, Clariallabes, 137
laticeps, Clarotes, 160
laticeps, Eutropius, 364
laticeps, Heterobranchus, 151
laticeps, Ituglanis, 407
laticeps, Loricaria, 288
laticeps, Olyra, 17, 18
laticeps, Pimelodella, 192
laticeps, Pimelodus, 160
laticeps, Plotosus, 345
laticeps, Pseudohemiodon, 288
laticeps, Schilbe, 364
laticeps, Silurus, 151
laticeps, Trichomycterus, 407
laticeps australis, Pimelodella, 189
latidens, Astroblepus, 65
latidens, Pygidium, 420
latidens, Trichomycterus, 419
latifrons, Ancistrus, 222
latifrons, Chaetostomus, 219, 222
latifrons, Hypostomus, 255

CHECKLIST OF CATFISHES, RECENT AND FOSSIL

latifrons, Noturus, 211
latifrons, Synodontis gambiensis, 315
latirostris, Arius, 47
latirostris, Hypostomus, 255
latirostris, Loricaria, 294
latirostris, Neoarius, 47
latirostris, Plecostomus, 255
latirostris, Rineloricaria, 294
latirostris, Sorubim, 342
latiscutatus, Arius, 31
latisoma, Farlowella, 239
latistriatum, Pygidium, 420
latistriatus, Trichomycterus, 420
latiura, Dasyloricaria, 233
latiura, Loricaria filamentosa, 233
latovittatus, Cryptopterus, 374
latus, Corydoras, 119
laucaensis, Trichomycterus, 420
laukidi, Rhamdia, 199
laurenti, Pimelodella, 192
lautus, Noturus hildebrandi, 213
lavetti, Ameiurus, 204
lavetti, Ictalurus, 204
layardi, Arius, 51
lazera, Clarias, 143læviuscula, Loricaria, 289
leerii, Wallago, 380, 381
lehmanni, Rhamdia, 202
leiacanthus, Clarias, 145
leiacanthus, Hyalobagrus, 92
leiacanthus, Leiocassis, 104
leiacanthus, Ompok, 371, 372
leiacanthus, Pseudomystus, 104
leiacanthus, Wallago, 371, 372
Leiarius, 334
Leiarius marmoratus, 334
Leiarius pictus, 334
leidyi, Ameiurus, 204
leidyi, Ictalurus, 204leightoni, Oxyloricaria, 300
leightoni, Sturisomatichthys, 300
Leiocassis, 92
Leiocassis (Dermocassis) analis, 101
Leiocassis aculeatus, 92
Leiocassis albicollaris, 104
Leiocassis albomarginatus, 101
Leiocassis bicolor, 104
Leiocassis brashnikowi, 100
Leiocassis brevicaudatus, 102
Leiocassis chaseni, 93
Leiocassis collinus, 443
Leiocassis crassilabris, 92
Leiocassis crassilabris macrops, 92
Leiocassis crassirostris, 92
Leiocassis ellenriederii, 104
Leiocassis fluviatilis, 85
Leiocassis fuscus, 103
Leiocassis hainanensis, 88
Leiocassis herzensteini, 92

Zootaxa 1418 © 2007 Magnolia Press · 587

Leiocassis hwanghoensis, 107
Leiocassis kaifenensis, 107
Leiocassis leiacanthus, 104
Leiocassis longibarbus, 92
Leiocassis longirostris, 93
Leiocassis lui, 102
Leiocassis macropterus, 83
Leiocassis mahakamensis, 104
Leiocassis micropogon, 93
Leiocassis microps, 100
Leiocassis moeschii, 105
Leiocassis myersi, 104
Leiocassis nitidus, 82
Leiocassis omeihensis, 102
Leiocassis poecilopterus, 93
Leiocassis regani, 93
Leiocassis robustus, 104
Leiocassis saravacensis, 93
Leiocassis similis, 108
Leiocassis sinyanensis, 108
Leiocassis tenebricus, 443
Leiocassis tenuifurcatus, 93
Leiocassis trilineatus, 103
Leiocassis ussuriensis, 101
Leiocassis virgatus, 101
leiopleura, Harttia, 240, 241
Leiosynodontis, 311
leiotetocephalus, Arius, 52
Leliella, 293, 294
lemai, Rhamdella, 196lemairii, Eutropius, 364
lembesseensis, Clariallabes melas, 137
lemniscatus, Pimelodus, 213
lemoinei, Arius, 432
lentiginosa, Ariopsis, 33
lentiginosa, Loricaria prolixa, 286
lentiginosa, Proloricaria, 286
lentiginosus, Amphilius, 22
lentiginosus, Doras, 177
lentiginosus, Tachisurus, 33
leonensis, Malapterurus, 301
leoparda, Synodontis, 316
leopardina, Synodontis, 316
leopardinus, Synodontis, 316
leopardinus, Trachelyopterus, 80
leopardinus, Trachycorystes, 80
leopardum, Glanidium, 75
leopardus, Centromochlus(Gephyromochlus), 75
leopardus, Corydoras, 119
leopardus, Pseudacanthicus, 286
leopardus, Stoneiella, 286
leopardus, Synodontis, 316
Leporacanthicus, 265
Leporacanthicus galaxias, 265
Leporacanthicus heterodon, 266
Leporacanthicus joselimai, 266
Leporacanthicus triactis, 266
leporhinus, Hemidoras, 174

leporhinus, Nemadoras, 174
leptacanthus, Noturus, 213
Leptarius, 53
Leptarius dowii, 53
leptaspis, Hexanematichthys, 47
leptaspis, Neoarius, 47
Lepthoplosternum, 128
Lepthoplosternum altamazonicum,
128
Lepthoplosternum beni, 128
Lepthoplosternum pectorale, 128
Lepthoplosternum stellatum, 128
Lepthoplosternum tordilho, 129
Lepthoplosternum ucamara, 129
Leptoancistrus, 266
Leptoancistrus canensis, 266
Leptoancistrus cordobensis, 266
leptocassis, Hexanematichthys, 45
leptochilus, Hisonotus, 248
Leptodoras, 172
Leptodoras acipenserinus, 172
Leptodoras cataniai, 172
Leptodoras copei, 172
Leptodoras hasemani, 172
Leptodoras juruensis, 172
Leptodoras linnelli, 173
Leptodoras myersi, 173
Leptodoras nelsoni, 173
Leptodoras praelongus, 173
Leptodoras rogersae, 173
Leptodoras trimaculatus, 174
Leptoglanidinae, 20
Leptoglaninae, 20
Leptoglanis, 20, 25
Leptoglanis, 187
Leptoglanis bouilloni, 164
Leptoglanis brevis, 28
Leptoglanis brieni, 24
Leptoglanis camerunensis, 28
Leptoglanis dorae, 28
Leptoglanis essequibensis, 187
Leptoglanis flavomaculatus, 28
Leptoglanis mandevillei, 28
Leptoglanis wamiensis, 29
Leptoglanis xenognathus, 25
leptonema, Silurus, 374
leptonotacanthus, Arius, 34
Leptops, 215
Leptorhamdia, 187
Leptorhamdia essequibensis, 187
Leptorhamdia marmorata, 187
Leptorhamdia schultzi, 187
leptorhynchus, Helicophagus, 324
leptos, Acentronichthys, 180
leptosoma, Rhamdella, 196
leptostriatus, Microglanis, 441
leptura, Andersonia, 24
lepturum, Chaetostoma, 229
lepturus, Chaetostomus, 229

588 · Zootaxa 1418 © 2007 Magnolia Press

lepturus, Cnidoglanis, 346, 442
lepturus, Euristhmus, 346
lepturus, Oreoglanis, 397
lepturus, Xyliphius, 62
leptus, Pimelodus, 332
Lerichei, Otolithus (Arius), 434
leroyi, Chimarrhoglanis, 21, 23
lestradei, Lophiobagrus, 161, 162
leucofrenatus, Hisonotus, 248
leucofrenatus, Otocinclus, 248
leucomelas, Chaetostoma, 229
leucomelas, Chaetostomus, 229
leucomelas, Corydoras, 119
leucophasis, Bagrus, 96
leucophasis, Mystus, 96
leucopodus, Silurichthys, 378
Leucoptera, Pimelodus pallidus, 210
Leucoptera, Silurus pallidus, 210
leucorhynchus, Akysis, 16
leucorhynchus, Pseudobagarius, 16
leucostictus, Ancistrus, 222
leucostictus, Chaetostomus, 222
levequei, Chrysichthys, 157
levequei, Synodontis, 316
levis, Copidoglanis, 345
levis, Hypostomus, 255
levis, Rhinelepis, 255
lewi, Trichomycterus, 420
lexi, Hypostomus, 255
lexi, Plecostomus, 255
Leyvaichths, 134
Leyvaichthys, 134
Leyvaichthys castaneus, 134, 407
liacanthus, Clarias, 145
lianae, Copionodon, 404
Liauchenoglanis, 161
Liauchenoglanis maculatus, 161liberiensis, Clarias, 141liberiensis, Eutropius, 364
libertatis, Doras, 174
lica, Bagarius, 383
lima, Hypostomus, 256
lima, Loricaria, 294
lima, Plecostomus, 256
lima, Rineloricaria, 294
lima, Silurus, 342
lima, Sorubim, 342
lima atropinnis, Plecostomus, 251
lima microlepidota, Loricaria, 295
lima var. microlepidota, Loricaria, 295
Limatulichthys, 266
Limatulichthys griseus, 266
Limatulichthys petleyi, 266
limbatus, Bagrus, 107
limbatus, Choeroplotosus, 346
limbatus, Plotosus, 108, 346, 350
limosus, Pimelodus, 215
limosus, Plecostomus, 253
limosus, Pylodictis, 215

FERRARIS

limosus, Silurus, 215
limpok, Kryptopterus, 370
limpok, Silurus, 370
limulus, Eurycheilichthys, 236
linami, Pimelodella, 192
lindica, Phractura, 26
lindicus, Clarias, 141
lineata, Nannorhamdia, 185
lineatum, Euglyptosternum, 388
lineatus, Imparfinis, 185
lineatus, Plotosus, 350
lineatus, Plotosus, 351, 352
lineatus, Silurus, 350
lineolatus, Ancistrus, 222
lineopunctata, Chaetostoma, 229
lineopunctatum, Chaetostoma, 229
linnelli, Leptodoras 173
Liobagrus, 19
Liobagrus andersoni, 19
Liobagrus anguillicauda, 19
Liobagrus formosanus, 19
Liobagrus kingi, 19
Liobagrus marginatoides, 19
Liobagrus marginatus, 19
Liobagrus mediadiposalis, 19
Liobagrus nantoensis, 19
Liobagrus nantoënsis, 19
Liobagrus nigricauda, 20
Liobagrus obesus, 20
Liobagrus reinii, 19, 20
Liobagrus styani, 20
Liobagrus sugubii, 20
Liobagrus xiurenensis, 20
Liocassis, 92
Liocassis baramensis, 93
Liocassis breviceps, 103
Liocassis crassilabris, 92
Liocassis crassirostris, 92
Liocassis doriae, 93
Liocassis emarginatus, 102
Liocassis hosii, 93
Liocassis inornatus, 103
Liocassis longirostris, 92, 93
Liocassis merabensis, 93
Liocassis moeschii, 104
Liocassis naso, 108
Liocassis rugosus, 104
Liocassis saravacensis, 93
Liocassis siamensis, 104
Liocassis torosilabris, 108
Liocassis truncatus, 103
Liocassis vaillanti, 105
Liocassus hirsutus, 107
liocephalus, Clarias, 145
Liosomadoras, 75
Liosomadoras morrowi, 75
Liosomadoras oncinus, 75
lipophthalmus, Doras (Oxydoras), 169
Lipopterichthys, 266

Lipopterichthys carrioni, 266, 267
Liposarcus, 290
Liposarcus altipinnis, 291
Liposarcus ambrosettii, 291
Liposarcus anisitsi, 291
Liposarcus disjunctivus, 291
Liposarcus jeanesianus, 292
Liposarcus scrophus, 292
Liposarcus varius, 292
liropus, Tachysurus, 41
listrorhinos, Apistoloricaria, 226
Listrura, 408
Listrura boticario, 408
Listrura camposi, 408
Listrura nematopteryx, 408
Listrura picinguabae, 444
Listrura tetraradiata, 408
Lithodoradinae, 166
Lithodoras, 173
Lithodoras dorsalis, 173
lithogaster, Doras, 173
Lithogeneinae, 217
Lithogenes, 217, 267
Lithogenes valencia, 267
Lithogenes villosus, 267
lithoides, Lithoxus, 267
lithophilus, Parasilurus, 380
lithophilus, Silurus, 380
lithostoma, Pangasius, 326
Lithoxancistrus, 286
Lithoxancistrus orinoco, 286, 287
Lithoxina, 217
Lithoxus, 217, 267
Lithoxus (Paralithoxus)
pallidimaculatus, 267
Lithoxus (Paralithoxus) planquettei,
267
Lithoxus (Paralithoxus) surinamensis,
268
Lithoxus boujardi, 267
Lithoxus bovallii, 267
Lithoxus lithoides, 267
Lithoxus pallidimaculatus, 267
Lithoxus planquettei, 267
Lithoxus stocki, 268
Lithoxus surinamensis, 268
lithurgicus, Ancistrus, 222
littorale, Hoplosternum, 127
littorale daillyi, Hoplosternum, 128
littoralis, Callichthys, 127
lituratus, Ancistrus, 290, 291
lituratus, Pterygoplichthys, 291
lividus, Silurus, 204, 205
lividus Fuscatus, Silurus, 205
lividus var. Fuscatus, Silurus, 205
livrée, Pimélode, 213
llanero, Lamontichthys, 264
loangwensis, Clarias, 151
loborhynchos, Chaetostoma, 227, 229

CHECKLIST OF CATFISHES, RECENT AND FOSSIL

lombarderoi, Xyliphius, 62
lonah, Bagrus, 391
lonah, Glyptothorax, 391, 394
longianalis, Clupisoma, 357
longianalis, Platytropius, 357
longibarbatus, Trichomycterus, 420
longibarbis, Arius, 334
longibarbis, Bathyclarias, 136
longibarbis, Chrysichthys, 157
longibarbis, Chrysobagrus, 157
longibarbis, Clarias, 136, 138
longibarbis, Clarias (Allabenchelys)
dumerili, 138
longibarbis, Dianema, 127
longibarbis, Dinotopterus, 136
longibarbis, Laides, 359
longibarbis, Pangasius, 359
longibarbus, Leiocassis, 92
longicauda, Allabenchelys, 137
longicauda, Clariallabes, 137
longicauda, Euchiloglanis, 399
longicauda, Glyptothorax, 391
longicauda, Imparfinis, 185
longicauda, Pareuchiloglanis, 399
longicauda, Phractura, 26
longicauda, Pimelodus (Rhamdia), 185
longicauda, Rineloricaria, 294
longicaudata, Olyra, 99
longicaudatus, Olyra, 99
longicaudatus, Clariallabes, 150
longicehpalus, Tachisurus, 39
longicephalus, Tachisurus, 39
longiceps, Astroblepus, 65
longiceps, Auchenoglanis, 163
longiceps, Clarias, 143
longiceps, Dolichamphilius, 24
longiceps, Osteogeneiosus, 51
longiceps, Parauchenoglanis, 163
longidorsalis, Chrysichthys, 157
longidorsalis nyongensis,
Chrysichthys, 158
longifilis, Akysis, 442
longifilis, Arges, 65
longifilis, Astroblepus, 65
longifilis, Callichthys, 129
longifilis, Chrysichthys, 155
longifilis, Copidoglanis, 349
longifilis, Eutropius, 361
longifilis, Heterobranchus, 151
longifilis, Hypophthalmus, 334
longifilis, Parailia, 360
longifilis, Pareutropius, 361
longifilis, Pimelodus, 338
longimanus, Ancistrus, 291
longimanus, Auchenipterichthys, 71
longimanus, Auchenipterus, 71
longimanus, Pseudeutropius, 362
longinema, Glyptothorax, 394
longior, Chasmocranus, 182, 183

Zootaxa 1418 © 2007 Magnolia Press · 589

longior, Clarias, 145
longipectoralis, Heteropneustes, 152
longipinna, Harttia, 241
longipinnis, Bariancistrus, 227
longipinnis, Chrysichthys, 157
longipinnis, Euanemus, 74
longipinnis, Gephyroglanis, 157
longipinnis, Hemiancistrus, 227
longipinnis, Rhamdella, 196
longipinnis, Trichogenes, 414
longiradiatus, Hypostomus, 256
longiradiatus, Plecostomus, 256
longirostris, Amphilius, 22
longirostris, Anoplopterus, 22
longirostris, Corydoras melanistius,
113
longirostris, Leiocassis, 93
longirostris, Liocassis, 92, 93
longirostris, Parakysis, 16
longirostris, Parotocinclus, 282
longirostris, Synodontis, 316
longispinis, Ælurichthys, 38
longispinis, Doras, 170
longispinis, Lasiancistrus, 287
longispinis, Pseudancistrus, 287
longispinis, Synodontis, 316
longispinis, Synodontis Batesi, 316
longisrostris, Nkondobagrus, 99
longiuscula, Rhamdella, 196
longjiangensis, Glyptothorax, 391
longus, Parasilurus asotus, 379
lopezi, Chasmocranus, 183
Lophiobagrus, 161
Lophiobagrus aquilus, 161
Lophiobagrus asperispinis, 161
Lophiobagrus brevispinis, 161
Lophiobagrus cyclurus, 161
Lophiobagrus lestradei, 161, 162
lophioides, Chaca, 135, 136
Lophiosilurus, 353
Lophiosilurus alexandri, 353
lophius, Amiurus, 204
lophophanes, Otothyris, 276
lophophanes, Rhinelepis, 276
loppei, Synodontis, 318
loretoensis, Corydoras, 119
Loricaria, 216, 268
Loricaria (Loricariichthys) fallax, 245
Loricaria accipenser, 270
Loricaria acuta, 269
Loricaria altipinnis, 245
Loricaria amazonica, 270
Loricaria anus, 269
Loricaria apeltogaster, 268
Loricaria apeltogaster amazonum, 288
Loricaria apeltogaster var. amazonum,
288
Loricaria aurea, 298
Loricaria barbata, 298, 299

Loricaria beni, 245
Loricaria bransfordi, 240
Loricaria brevirostris, 299
Loricaria brunnea, 269
Loricaria cacerensis, 245
Loricaria cadeae, 294
Loricaria capetensis, 233
Loricaria caquetae, 296
Loricaria carinata, 268
Loricaria cashibo, 270
Loricaria castanea, 270
Loricaria catamarcensis, 294
Loricaria cataphracta, 268, 270
Loricaria cirrhosa, 268, 296
Loricaria clavipinna, 268
Loricaria commersonoides, 278
Loricaria cubataonis, 294
Loricaria curvispina, 296
Loricaria Dentata, 268
Loricaria devincenzii, 288
Loricaria dura, 268
Loricaria Eigenmanni, 245
Loricaria Evansii, 296
Loricaria felipponei, 294
Loricaria filamentosa, 233, 234
Loricaria filamentosa latiura, 233
Loricaria filamentosa seminuda, 234
Loricaria fimbriata, 297
Loricaria flava, 259
Loricaria frenata, 299
Loricaria griseus, 266
Loricaria gymnogaster, 297
Loricaria gymnogaster lagoichthys,
297
Loricaria henselii, 294
Loricaria hoehnei, 245
Loricaria jaraguensis, 294
Loricaria jubata, 246
Loricaria kneri, 299
Loricaria konopickyi, 246
Loricaria kronei, 294
Loricaria labialis, 270
Loricaria laeviuscula, 289
Loricaria læviuscula, 289
Loricaria lamina, 288
Loricaria lanceolata, 246
Loricaria lata, 268
Loricaria laticeps, 288
Loricaria latirostris, 294
Loricaria lima, 293, 294
Loricaria lima microlepidota, 295
Loricaria lima var. microlepidota, 295
Loricaria macrodon, 227
Loricaria macromystax, 293
Loricaria macrops, 293
Loricaria maculata, 269, 270
Loricaria magdalenae, 246
Loricaria melanoptera, 252
Loricaria microdon, 271

590 · Zootaxa 1418 © 2007 Magnolia Press

Loricaria microlepidogaster, 295
Loricaria nickeriensis, 269
Loricaria nigricauda, 246
Loricaria nudirostris, 271
Loricaria nudiventris, 297
Loricaria panamensis, 299
Loricaria parahemiodon, 270
Loricaria pareiacantha, 295
Loricaria parnahybae, 269
Loricaria parva, 246
Loricaria paulina, 294
Loricaria phoxocephala, 246
Loricaria piauhiae, 269
Loricaria piracicabae, 269
Loricaria platystoma, 233
Loricaria platyura, 246
Loricaria prolixa, 286
Loricaria prolixa lentiginosa, 286
Loricaria puganensis, 297
Loricaria punctata, 266, 292
Loricaria rostrata, 245, 298, 300
Loricaria scolopacina, 237
Loricaria setifera, 268
Loricaria simillima, 269
Loricaria sneiderni, 246
Loricaria spinosae, 286
Loricaria Spixii, 270
Loricaria Steinbachi, 263
Loricaria steindachneri, 295
Loricaria stewarti, 247
Loricaria strigilata, 295
Loricaria Stübelii, 271
Loricaria submarginatus, 246
Loricaria teffeana, 247
Loricaria thrissoceps, 295
Loricaria tucumanensis, 269
Loricaria tuyrensis, 234
Loricaria uracantha, 240
Loricaria uracantha rupestre, 240
Loricaria Valenciennesii, 247
Loricaria variegata, 232, 233
Loricaria variegata venezuelae, 233
Loricaria vetula, 278loricariformis, Harttia, 240, 241
Loricariichthyina, 217
Loricariichthys, 217, 269
Loricariichthys acutus, 269
Loricariichthys anus, 269
Loricariichthys brunneus, 269
Loricariichthys cashibo, 270
Loricariichthys castaneus, 270
Loricariichthys chanjoo, 270
Loricariichthys derbyi, 270
Loricariichthys edentatus, 270
Loricariichthys hauxwelli, 270
Loricariichthys labialis, 270
Loricariichthys maculatus, 270
Loricariichthys melanocheilus, 271
Loricariichthys melini, 246

FERRARIS

Loricariichthys microdon, 271
Loricariichthys nudirostris, 271
Loricariichthys parnahybae, 266
Loricariichthys platymetopon, 271
Loricariichthys rostratus, 271
Loricariichthys stuebelii, 271
Loricariichthys ucayalensis, 271
Loricariidae, 216
loricatus, Callichthys, 111
loricatus, Doras, 167
loweae, Bathyclarias, 149
loweae, Dinotopterus, 149lowei, Gephyroglanis, 158
loxozonus, Corydoras, 119
lualae, Clarias, 142
lubosicus, Bagrus, 84
lucenai, Trachelyopterus, 80
Luceri, Platystoma, 342
Luciopimelodinae, 329
Luciopimelodus, 329, 334
Luciopimelodus pati, 334
lucipinnis, Synodontis, 445
luderwaldti, Pseudancistrus, 280
lufirae, Chiloglanis, 306
lufirae, Synodontis, 316
lui, Leiocassis, 102
luigiana, Rhamdia, 199
lukugae, Chiloglanis, 306
lukugae, Phractura, 26
lumholtzi, Cryptopterus, 370
lumholtzi, Kryptopterus, 370
lundbergi, Prietella, 215
lundbergi, Typhlobelus, 426
luniscutis, Arius, 36
luniscutis, Aspistor, 36
lupus, Ictalurus, 209, 211
lupus, Pimelodus, 209
lurida, Ageniosus (Silonia), 366
luridus, Macrones, 90
luridus, Pachypterus, 327
luteomaculatus, Hypostomus, 256
luteomaculatus, Plecostomus, 256
lutescens, Pimelodus, 215
luteus, Hypostomus, 256
luteus, Noturus, 212
luteus, Oloplotosus, 349
luteus, Plecostomus, 256
Lütkeni, Platystoma, 344
Lütkeni, Plecostomus, 251
luvur, Silurus, 431
lynx, Pimelodus, 204lyra, Oxyloricaria, 299
lyra, Physopyxis, 176lyra, Sturisoma, 299
lyriformis, Agmus, 60Mmabusi, Chrysichthys, 157
macarenensis, Phenacorhamdia, 188,
189

maccus, Panaqolus, 277
maccus, Panaque, 277
macgrewi, Ameiurus, 204
macgrewi, Ictalurus, 205
machadoi, Genidens, 44
machadoi, Gladioglanis, 183
machadoi, Platyclarias, 152
machadoi, Plectrochilus, 411
machadoi, Tachysurus, 44
machiquense, Chaetostoma, 229
machiquensis, Chaetostoma, 229
macilenta, Denticetopsis, 133
macilentus, Hemicetopsis, 133
maclareni, Clarias, 145
maconnellii, Ameiurus, 211
macracanthus, Arius, 36
macracanthus, Bagrichthys, 82
macracanthus, Bagroïdes, 82
macracanthus, Centromochlus, 73
macracanthus, Clarias, 143
macracanthus, Pseudobagrichthys, 82
macracanthus, Rita, 106
macraei, Hatcheria, 405
macraei, Thrichomycterus, 405
macrepipterus, Synodontis, 314
macrocephala, Nannorhamdia, 192
macrocephala, Pimelodella, 192
macrocephalus, Ariodes, 51
macrocephalus, Arius, 51
macrocephalus, Atopochilus, 303
macrocephalus, Clarias, 145
macrocephalus, Clarotes, 160
macrocephalus, Cnidoglanis, 346
macrocephalus, Hemipimelodus, 41
macrocephalus, Kryptopterichthys,370
macrocephalus, Kryptopterus, 370
macrocephalus, Osteogeneiosus, 51
macrocephalus, Plotosus, 346
macrocephalus, Silurus, 341
macrochir, Belodontichthys, 368
macrodon, Brochiloricaria, 227
macrodon, Loricaria, 227
macrodon, Synodontis, 313
macromaculatus, Glyptothorax, 391
macromycterus, Typhlobelus, 426
macromystax, Clarias, 145
macromystax, Loricaria, 293
macromystax, Rhadinoloricaria, 293
macronema, Akysis, 16
macronema, Chlarias angolensis, 141
macronema, Heterobranchus, 151
macronema, Hypselobagrus, 97
macronema, Pangasius, 327
macronema, Pseudorhamdia, 337
macronema, Silurodes, 372
macronema, Silurus, 371, 372
macronemus, Bagrus, 37, 97
macronemus, Bagrus, 93, 97

CHECKLIST OF CATFISHES, RECENT AND FOSSIL

macronemus, Oreoglanis, 397
macronemus, Pseudobagarius, 16Macrones, 85, 106
Macrones (Hemibagrus) filamentus, 88
Macrones (Hemibagrus) wyckioides,
91
Macrones (Leiocassis) brashnikowi,
100
Macrones (Liocassis) taeniatus, 102
Macrones (Pseudobagrus) tenuis, 103
Macrones aleppensis, 97
Macrones argentivittatus, 100
Macrones armatus, 94
Macrones baramensis, 87
Macrones bimaculatus, 94
Macrones bleekeri, 89, 94
Macrones Blythii, 85
Macrones bo, 88
Macrones bongan, 87
Macrones chinensis, 88
Macrones colvillii, 97
Macrones corsula, 90
Macrones Dayi, 85
Macrones elongatus, 88
Macrones fortis, 88
Macrones fortis capitulum, 88
Macrones fortis var. capitulum, 88
Macrones herzensteini, 92
Macrones howong, 88
Macrones kajan, 88
Macrones luridus, 90
Macrones medianalis, 102
Macrones merianiensis, 85
Macrones microphthalmus, 90
Macrones montanus dibrugarensis, 96
Macrones montanus var.
dibrugarensis, 96
Macrones peguensis, 90
Macrones pluriradiatus, 90
Macrones pratti, 102
Macrones pulcher, 97
Macrones rufescens, 94, 97
Macrones sinensis, 165
Macrones vittata, 106
Macronichthys, 106
Macronoides, 85, 87
Macronoides wilsoni, 87
macronotacanthus, Arius, 47
macronotus, Batasio, 85
macrophthalma, Synodontis, 316
macrophthalma, Xenocara, 222
macrophthalmus, Ancistrus, 222
macrophthalmus, Hypostomus, 256
macrophthalmus, Hypostomus
pseudohemiurus, 256
macrophthalmus, Plotosus, 349
macrophthalmus, Synodontis, 316
macropogon, Chrysichthys, 160
macrops, Chaetostomus, 243

Zootaxa 1418 © 2007 Magnolia Press · 591

macrops, Chrysichthys, 155
macrops, Hemiancistrus, 243
macrops, Henonemus, 406
macrops, Hypostomus, 256
macrops, Leiocassis crassilabris, 92
macrops, Loricaria, 293
macrops, Plecostomus, 256
macrops, Ricola, 293
macrops, Stegophilus, 406
macrops, Synodontis, 316
Macropterobagrus, 87
Macropteronotus, 138
Macropteronotus charmuth, 138, 148
Macropteronotus fuscus, 142
Macropteronotus hexacicinnus, 148
Macropteronotus jagur, 140
Macropteronotus magur, 140
macropterum, Exostoma, 398
macropterus, Amblyceps, 18
macropterus, Bagrichthys, 82
macropterus, Bagroides, 82, 83
macropterus, Bagroïdes, 82
macropterus, Bagrus bayad, 83
macropterus, Calophysus, 332
macropterus, Chiloglanis, 306
macropterus, Chrysichthys, 157
macropterus, Corydoras, 130
macropterus, Hemibagrus, 87, 89, 91
macropterus, Leiocassis, 83
macropterus, Pareuchiloglanis, 399
macropterus, Pimelodes, 332
macropterus, Porcus bayad, 83
macropterus, Oreoglanis, 398
macropterus, Scleromystax, 130
macropthalmos, Eutropius, 361, 362
macropthalmos, Proeutropiichthys,362
macropthalmos, Pseudosilurus, 374
macrorhynchus, Hemipimelodus, 32
macrospila, Piramutana, 339
macrospilus, Otocinclus, 275
macrosteus, Corydoras, 112
macrostigma, Synodontis, 317
macrostoma, Auchenoglanis, 154
macrostoma, Synodontis, 317
macrostomus, Anaspidoglanis, 154
macrostomus, Silurus (Callichrus), 381
macroteronema, Cetopsis, 133
macrotis, Chrysichthys, 157
Macrotocinclus, 271
Macrotocinclus affinis, 271
Macrotocinclus flexilis, 271
macrotrema, Euchiloglanis, 399
macrotremus, Pareuchiloglanis, 399
macrura, Phractura, 27
macruropterygius, Arius, 55
macrurus, Clarias, 148
macturki, Pimelodella, 192
maculata, Cteniloricaria, 233

maculata, Loricaria, 269, 270
maculata, Parasturisoma, 233
maculata, Wertheimeria, 179
maculatum, Glyptosternon, 387
maculatum, Parexostoma, 387
maculatus, Ancistrus, 222
maculatus, Arius, 34, 35
maculatus, Chaetostomus, 228
maculatus, Chaetostomus (Ancistrus)
cirrhosus, 222, 228
maculatus, Chaetostomus (Ancistrus)
cirrhosus var., 222, 228
maculatus, Corydoras, 122
maculatus, Doras, 177
maculatus, Liauchenoglanis, 161
maculatus, Loricariichthys, 270
maculatus, Nothoglanidium, 161
maculatus, Parastegophilus, 410
maculatus, Pimelodus, 336, 338, 345
maculatus, Pimelodus, 209
maculatus, Pimelodus (Bagrus), 345
maculatus, Silurus, 35, 207, 209
maculatus, Stegophilus, 410
maculatus, Synodontis, 318
maculatus, Trichomycterus, 415
maculatus, Wallago, 381
maculatus Erythroptera, Silurus, 209
maculatus var. Erythroptera, Silurus,
209
maculicauda, Otocinclus, 281, 283
maculicauda, Parotocinclus, 283
maculifer, Corydoras, 119
maculipinna, Synodontis, 320
maculipinnis, Akysis, 14
maculipinnis, Hisonotus, 248
maculipinnis, Otocinclus, 248
maculocephala, Cascadura, 127, 128
maculosus, Aspidoras, 109
maculosus, Auchenipterus, 79
maculosus, Auchenoglanis, 163
maculosus, Synodontis, 311, 318, 321
maculosus, Trachelyopterus coriaceus,
79
macunaima, Ituglanis, 407
macushi, Hypostomus, 256
macuspanensis, Rhamdia, 199
madagascariensis, Arius, 35
madeiræ, Plecostomus, 225
madeirensis, Ageneiosus, 69
madraspatanum, Glyptosternum, 391
madraspatanus, Glyptothorax, 388,
391
maesii, Amphilius, 22
maesotensis, Erethistes, 384
magatensis, Arius, 52
Magdalenae, Auchenipterus, 79
magdalenae, Branchioica, 411
magdalenae, Hemiloricaria, 246
magdalenae, Hoplosternum, 128

592 · Zootaxa 1418 © 2007 Magnolia Press

Magdalenae, Loricaria, 246
magdalenae, Trachycorystes, 79
magdalenae, Xyliphius, 62
magnus, Chrysichthys, 157
magoi, Ageneiosus, 69
magoi, Brachyglanis, 180
magoi, Pygidianops, 412
magur, Macropteronotus, 140
mahakamensis, Acrochordonichthys,
12
mahakamensis, Leiocassis, 104
mahakamensis, Pangasius, 327
mahakamensis, Pseudomystus, 104
mahengeensis, Chrysichthys, 157
major, Akysis, 391
major, Glyptothorax, 391
major, Hemibagrus, 89
majusculus, Bagrichthys, 83
malabarbai, Microglanis, 354
Malabarica, Hara, 96
malabaricus, Arius, 35
Malabaricus, Bagrus, 96
malabaricus, Mystus, 96
malabaricus, Ompok, 372
malabaricus, Silurus, 372
Malacobagrus, 330
Malacoglanis, 408
Malacoglanis gelatinosus, 408
malacops, Ancistrus, 222
malacops, Chaetostomus, 222
Malacopterurus, 301
malaisei, Glyptosternon, 387
Malapteruridae, 300
Malapterurini, 300
Malapterurus, 300, 301
Malapterurus affinis, 301
Malapterurus barbatus, 301
Malapterurus beninensis, 301
Malapterurus cavalliensis, 301
Malapterurus coila, 356
Malapterurus electricus, 301
Malapterurus electricus oguensis, 302
Malapterurus electricus, var.
ogooensis, 302
Malapterurus electricus var. oguensis,
302
Malapterurus gossei, 302
Malapterurus leonensis, 301
Malapterurus melanochir, 301
Malapterurus microstoma, 301
Malapterurus microstomus, 301
Malapterurus minjiriya, 302
Malapterurus monsembeensis, 302
Malapterurus murrayi, 302
Malapterurus occidentalis, 302
Malapterurus oguensis, 302
Malapterurus polli, 302
Malapterurus punctatus, 302
Malapterurus shirensis, 302

FERRARISMalapterurus stiassnyae, 301
Malapterurus tanganyikaensis, 302
Malapterurus tanoensis, 302
Malapterurus teugelsi, 302
Malapterurus thysi, 301
Malapterurus zambeziensis, 302
Malapterus, 301
Malapterus, 301, 356
Malapterus (Ailia) bengalensis, 356
Malapturus, 301
malaris, Clarias, 143
malarmo, Platysilurus, 340
maldonadoi, Bullockia, 404
Maldonadoi, Hatcheria, 403, 404
malignus, Plotosus, 351
Malopterurus, 301
mamore, Corydoras, 120
mamoré, Corydoras, 120
managuensis, Pimelodus, 199
mancoi, Astroblepus, 65
mandevillei, Atopochilus, 304
mandevillei, Leptoglanis, 28
mandevillei, Pareutropius, 361
mandevillei, Zaireichthys, 28
mandibularis, Eutropius, 364
mandibularis, Schilbe, 364
mangois, Amblyceps, 18
mangois, Pimelodus, 18
mangrullo, Silurus, 341
mangurus, Pimelodus, 355
mangurus, Pseudopimelodus, 355
manillensis, Arius, 35
manillensis, Cephalocassis, 41
Manillensis, Pimelodus, 41
manipurensis, Glyptothorax, 391
manipurensis, Laguvia, 382
maniradii, Sorubim, 342
manjong, Arius, 55
manni, Pteroglanis, 343
manni, Synodontis, 317
manyangae, Allabenchelys, 137
manyangae, Clariallabes, 137
mapale, Cathorops, 40
maquinensis, Rineloricaria, 294
maracaibero, Lamontichthys, 264
maracaiboensis, Hemiancistrus, 243
maracaiboensis, Lasiancistrus, 265
maracaiboensis, Pygidium banneaui,
420
maracaiboensis, Trichomycterus, 420
maracasae, Ancistrus, 223
marapoama, Otothyropsis, 276
marcapatae, Chaetostoma, 229
marcapatae, Chaetostomus, 229
margaritifer, Hypostomus, 256
margaritifer, Plecostomus, 256
margaritifer butantanis, Plecostomus,
256

marginata, Pimelodus pallidus var.,210
marginata, Taunayia, 203
marginatoides, Amblyceps, 19
marginatoides, Liobagrus, 19
marginatum, Chaetostoma, 229
marginatus, Amblyceps, 19
marginatus, Chaetostomus, 229
marginatus, Hypophthalmus, 333
marginatus, Liobagrus, 19
marginatus, Noturus, 213
marginatus, Pimelodus, 206
marginatus, Plotosus, 350
marginatus, Silurus pallidus, 210
marginatus, Silurus pallidus var., 210
marginatus atrorus, Schilbeodes, 213
mariae, Astroblepus, 65
mariae, Cyclopium, 65
mariae, Oloplotosus, 349
mariae, Otocinclus, 275
mariaelenae, Farlowella, 238
mariai, Imparales, 188
mariai, Nemuroglanis, 188
marinus, Bagre, 37
marinus, Silurus, 37, 38, 44
marlieri, Chiloglanis, 306
marmorata, Leptorhamdia, 187
marmorata, Synodontis, 317
marmoratum, Trachypoma, 404
marmoratus, Ageneiosus, 69
marmoratus, Amphilius
grammatophorus, 23
marmoratus, Arges, 65
marmoratus, Astroblepus, 65
marmoratus, Corydoras, 122
marmoratus, Helogenes, 134
marmoratus, Doras, 171
marmoratus, Franciscodoras, 171
marmoratus, Leiarius, 334
marmoratus, Pimelodus, 206
marmoratus, Schilbe, 364
marmoratus, Sciades, 334
marmoratus, Silurichthys, 378
marmoratus, Synodontis, 317
marmoratus, Trichomycterus, 415
marmoratus truncatus, Synodontis, 317
marmorescens, Chaetostoma, 229
marpus, Clarias, 148
marquesi, Ageneiosus, 70
marthae, Brachyrhamdia, 181
marthae, Rhamdia, 181
martinezi, Pimelodella, 193
martini, Acestridium, 218
martini, Ancistrus, 223
martini, Ancistrus triradiatus, 223
martini, Farlowella, 238
Martyi, Rhamdia sebae, 202
mastersi, Arius, 45
mastersi, Hexanematichthys, 45

CHECKLIST OF CATFISHES, RECENT AND FOSSIL

Mastiglanis, 187
Mastiglanis asopos, 187
mathisoni, Zungaro, 356
matthesi, Synodontis, 317
mattogrossensis, Ancistrus, 300
mattogrossenssis, Ancistrus, 300
maurus, Bagrus, 158
maurus, Chrysichthys, 157, 158, 160
maydelli, Hemibagrus, 89
maydelli, Mystus (Mystus), 89
maydeni, Noturus, 441
mayoloi, Hemiancistrus, 264
mayoloi, Lasiancistrus, 265
mazatlana, Netuma, 54
mboycy, Trichomycterus, 420
mbozi, Chiloglanis, 306
McCaskei, Ichthaelurus, 204
Medemichthys, 188
Medemichthys guayaberensis, 188
mediadiposalis, Liobagrus, 19
medianalis, Macrones, 102
medianalis, Pseudobagrus, 102
medians, Ancistrus, 242, 243
medians, Hemincistrus, 243
mediobarbis, Neosilurus, 347
meeki, Amiurus, 209
meeki, Pimelodella, 193
meesi, Brachyrhamdia, 181
megacephalus, Chaetostomus, 244
megacephalus, Hemiancistrus, 244
Megalancistrus, 272
Megalancistrus barrae, 272
Megalancistrus parananus, 272
Megalechis, 129
Megalechis picta, 129
Megalechis thoracata, 129
Megalocentor, 409
Megalocentor echthrus, 409
Megalodoras, 173
Megalodoras guayoensis, 173
Megalodoras irwini, 173, 174
Megalodoras paucisquamatus, 173
Megalodoras uranoscopus, 174
Megalonema, 335
Megalonema argentina, 335
Megalonema pauciradiatum, 335
Megalonema platanum, 335
Megalonema platycephalum, 335
Megalonema platycephalum
psammium, 335
Megalonema psammium, 335
Megalonema punctatum, 339
Megalonema rhabdostigma, 338
Megalonema robustum, 339
Megalonema xanthum, 335
megalops, Centromochlus, 73
megalops, Iheringichthys, 334
megalops, Pimelodella, 193
megalops, Pimelodus, 210

Zootaxa 1418 © 2007 Magnolia Press · 593

megalops, Pseudeutropius, 362
megalopteryx, Hemiancistrus, 244
megalostomus, Ancistrus, 223
megalura, Pimelodella, 193
megapogon, Clarias, 141
megastomus, Plotosus, 345, 346
megistus, Ernstichthys, 60
megistus, Hoplomyzon, 60
meizospilos, Hemiancistrus, 244
mekongensis, Hemisilurus, 368
mekongensis, Pangasius, 327
meladerma, Clarias, 145
melampterus, Callichthys, 127, 128
melanistius, Corydoras, 120
melanistius brevirostris, Corydoras,
114
melanistius longirostris,
Corydoras,113
melanocheilus, Loricariichthys, 271
melanochir, Arius, 41
melanochir, Cephalocassis, 41
melanochir, Malapterurus, 301
Melanodactylus, 155melanoderma, Clarias, 146
melanodermatum, Steindachneridion,
343
melanogaster, Acrochordonichthys, 13
melanogaster, Pimelodus, 12, 13
melanopogon, Ageneiosus, 68
melanopogon, Clarias, 145
melanops, Tridens, 426
melanoptera, Gagata, 386
melanoptera, Loricaria, 252
melanoptera, Synodontis, 317
melanopterum, Glanidium, 75
melanopterus, Bagroides, 83
melanopterus, Gagata, 386
melanopterus, Synodontis, 317
melanopterus, Xyliphius, 62melanopterygius, Arius, 56
melanopus, Arius, 40
melanopus, Cathorops, 40
melanostictus, Synodontis, 318
melanostictus iturii, Synodontis, 315
melanostictus var. iturii, Synodontis,
315
melanotaenia, Corydoras, 120
melanurus, Batrochoglanis, 353
melanurus, Pachypterus, 358
melanurus, Silurus cerulescens, 209
melanurus, Silurus cerulescens, var.,
209
melaphareus, Entomocorus, 74
melapterus, Bagroides, 83
melapterus, Bagroïdes, 83
melas, Ameiurus, 205
melas, Ancistrus, 220
melas, Bagrus, 95
melas, Brachyglanis, 181

melas, Bunocephalus, 58
melas, Clariallabes, 137
melas, Clarias, 137
melas, Pimelodus, 205
melas, Silurus, 205
melas lembesseensis, Clarias, 137
melasoma, Clarias, 146
meleagris, Hypostomus, 256
meleagris, Plecostomus, 256
melini, Corydoras, 120
melini, Hemiloricaria, 246
melini, Loricariichthys, 246
Mellandi, Clarias, 146
membranacea, Hemisynodontis, 309
membranaceus, Pimelodus, 309
mendozensis, Silvinichthys, 413
mendozensis, Trichomycterus, 413
menezesi, Aspidoras, 109
menezesi, Auchenipterus, 72
menezesi, Pyxiloricaria, 292
menoda, Hemibagrus, 89
menoda, Pimelodus, 89
menoni, Fajumia, 429
mentalis, Eutropius, 364
mento, Silurus, 380
merabensis, Liocassis, 93
meraukensis, Copidoglanis, 351
meraukensis, Porochilus, 351
mercatoris, Arius, 31
merevari, Harttia, 241
merianiensis, Batasio, 85
merianiensis, Macrones, 85
meridae, Trichomycterus, 420
meridionalis, Akysis, 17
meridionalis, Amiurus, 208
meridionalis, Bagrus, 84
meridionalis, Corydoras, 116
meridionalis, Pseudobagarius, 17
meridionalis, Silurus, 380
meridionalis, Silurus soldatovi, 380
Merodontotus, 330
Merodontotus tigrinus, 330, 331
merus, Glyptothorax, 390
mesembrinus, Diplomystes, 166
mesembrinus, Diplomystes viedmensis,
166
mesembrinus, Olivaichthys, 166
mesops, Bagrus, 53
metae, Corydoras, 120
metae, Ituglanis, 407
metae, Pimelodella, 193
metae, Pygidium, 407
metae guayaberensis, Pygidium, 407
Metaloricaria, 217, 272
Metaloricaria nijsseni, 272
Metaloricaria paucidens, 272
Metaloricariina, 217
mexicanus, Amiurus, 209
mexicanus, Ictalurus, 209

594 · Zootaxa 1418 © 2007 Magnolia Press

meyenii, Bagrus (Ariodes), 52
mica, Mystus, 100
mica, Pelteobagrus, 100
micayi, Rhamdia, 197
micracanthus, Bagrus, 96
micracanthus, Corydoras, 120
micranodus, Bagrichthys, 83
micrescens, Astroblepus, 65
micrescens, Astroblepus grixalvii, 65
micristius, Pareutropius, 360, 361
Microcambeva, 409
Microcambeva barbata, 409
Microcambeva ribeirae, 409
microcephala, Rhamdia, 202
microcephala, Rhamdiopsis, 202
microcephalus, Arius, 35
microcephalus, Cnidoglanis, 346
microcephalus, Corydoras, 122
microcephalus, Pseudarius, 35
microcephalus, Saccobranchus, 151
microcephalus, Silurus, 374
microceps, Euristhmus, 346
microceps, Plotosus, 346
Microcorydoras, 112
microdon, Loricaria, 271
microdon, Loricariichthys, 271
microdorsalis, Parasilurus, 380
microdorsalis, Silurus, 380
microgalaeus, Aspidoras, 109
microgastropterygius, Arius, 55
Microglanis, 354
Microglanis ater, 354
Microglanis cibelae, 354
Microglanis cottoides, 354
Microglanis eurystoma, 354
Microglanis garavelloi, 354
Microglanis iheringi, 354
Microglanis leptostriatus, 441
Microglanis malabarbai, 354
Microglanis nigripinnis, 354
Microglanis parahybae, 354
Microglanis pataxo, 444
Microglanis pellopterygius, 354
Microglanis poecilus, 354
Microglanis secundus, 355
Microglanis variegatus, 355
Microglanis zonatus, 355
Microlepidogaster, 272
Microlepidogaster bahiensis, 281
Microlepidogaster bourguyi, 300
Microlepidogaster depressinotus, 248
Microlepidogaster doceanus, 282
Microlepidogaster guentheri, 296
Microlepidogaster guntheri, 295
Microlepidogaster güntheri, 296
Microlepidogaster perforated, 272
Microlepidogaster perforatus, 272
Microlepidogaster taimensis, 249
microlepidogaster, Loricaria, 295

FERRARIS

microlepidogaster, Rineloricaria, 295
microlepidota, Loricaria lima, 295
microlepidota, Rineloricaria, 295
micromaculatus, Hypostomus, 256
micrommatos, Hemiancistrus, 244
Micromyzon, 61
Micromyzon akamai, 61
Micronema, 371
Micronema bleekeri, 374
Micronema cheveyi, 371
Micronema hexapterus, 371
Micronema moorei, 371
Micronema platypogon, 371
Micronema typus, 375
micronemus, Pangasius, 328
micronemus, Phalacronotus, 375
micronemus, Pseudolais, 328
micronemus, Silurus, 375
micronotacanthus, Arius, 55
microphthalmus, Brachyglanis, 181
microphthalmus, Clarias, 143
microphthalmus, Dolichallabes, 149
microphthalmus, Euristhmus, 442
microphthalmus, Hemibagrus, 90
microphthalmus, Macrones, 90
micropoeus, Doras, 171
micropoeus, Hemidoras, 171
micropogon, Bagrus, 92, 93
micropogon, Batrachocephalus, 38
micropogon, Chiloglanis, 305
micropogon, Eutropius, 364
micropogon, Leiocassis, 93
micropogon, Schilbe, 364
micropogon, Silurus, 374
micropogon, Wallago, 381
microps, Akysis, 14
microps, Chaetostoma, 229
microps, Chaetostomus, 229
microps, Chiloglanis, 306
microps, Corydoras, 112
microps, Harttia, 290
microps, Imparfinis, 186
microps, Leiocassis, 100
microps, Neoplecostomus, 273
microps, Pareiodon, 411
microps, Pelteobagrus, 100
microps, Plecostomus, 273, 281
microps, Plecostomus (Rhinelepis),281
microps, Pterosturisoma, 290
microps, Rhamdia, 201
microps, Saccobranchus, 151
micropterus, Arius, 166
micropterus, Pimelodus, 200
micropunctatus, Plecostomus, 225
micropus, Kryptopterus, 369
microstoma, Malapterurus, 301
microstoma, Pimelodus, 338
microstomus, Arius, 39

microstomus, Clarias, 146
microstomus, Hemidoras, 179
microstomus, Hypostomus, 250, 257
microstomus, Malapterurus, 301
microstomus, Trachydoras, 179
Microsynodontis, 309
Microsynodontis armatus, 309
Microsynodontis batesii, 309
Microsynodontis Christyi, 309
Microsynodontis emarginata, 309
Microsynodontis emarginatus, 309
Microsynodontis hirsuta, 310
Microsynodontis hirsutus, 310
Microsynodontis laevigata, 310
Microsynodontis laevigatus, 310
Microsynodontis lamberti, 310
Microsynodontis nannoculus, 310
Microsynodontis nasutus, 310
Microsynodontis notata, 310
Microsynodontis notatus, 310
Microsynodontis polli, 310
Microsynodontis vigilis, 310
micruropterus, Phalacronotus, 375
micruropterygius, Arius, 36
midgleyi, Arius, 47
midgleyi, Neoarius, 47
migrans, Pygidium, 420
migrans, Trichomycterus, 420
milesi, Autanadoras, 178, 179
milesi, Chaetostoma, 229
milesi, Chaetostomus, 229
militaris, Ageneiosus, 68, 69
militaris, Ageniosus, 68
militaris, Arius, 51
militaris, Osteogeneiosus, 51
militaris, Silurus, 51, 68, 69
mimonha, Trichomycterus, 420
mimulus, Otocinclus, 275
mindoense, Cyclopium, 65
mindoensis, Astroblepus, 65
minimaculatus, Glyptothorax, 391
minimus, Kryptopterus, 370
minjiriya, Malapterurus, 302
mino, Ageneiosus, 38
mino, Batrachocephalus, 38
minor, Kryptopterus, 370
minuta, Rhamdia, 186
minutum, Cetopsidium, 130
minutum, Glyptosternon, 393
minutum, Scleronema, 413
minutus, Ancistrus, 223
minutus, Bunocephalus, 59
minutus, Hemicetopsis, 130
minutus, [Hisonotus], 249
minutus, Imparfinis, 186
minutus, Glyptothorax, 391, 392, 393
minutus, Parotocinclus, 283
minutus, Trachyglanis, 28
minutus, Silurus, 431

CHECKLIST OF CATFISHES, RECENT AND FOSSIL

minutus, Trichomycterus, 413
miostoma, Wallago, 372
miostomus, Ompok, 372
mirini, Imparfinis, 186
mirissumba, Trichomycterus, 420
misionera, Rineloricaria, 295
mispilliensis, Amiurus, 206
misrai, Fajumia, 429
misrai, Mystus, 97
mitchelli, Pseudeutropius, 362
Miuroglanis, 409
Miuroglanis platycephalus, 409
miurus, Noturus, 214
möbiusii, Eutropius, 364
Mochocus, 310
Mochocus brevis, 310
Mochokidae, 303
Mochokiella, 310
Mochokiella paynei, 310
Mochokus, 303, 310
Mochokus brevis, 310
Mochokus niloticus, 310, 311
modesta, Pimelodella, 193
modestum, Glyptosternum, 389
modestus, Pimelodus, 193
modjensis, Chiloglanis, 306
moebii, Eutropius, 365
moebiusii, Schilbe, 364
moeschii, Leiocassis, 105
moeschii, Liocassis, 104
moeschii, Pseudomystus 104
molinae, Cetopsorhamdia, 182
mollespiculum, Neosilurus, 348
molliceps, Arius, 55
mollinasum, Chaetostoma, 230
mollinasus, Chaetostomus, 230
mollis, Silurus, 214mondolfi, Pygidium, 421
mondolfi, Trichomycterus, 421
mong, Pimelodus, 55
mongallensis, Slatinia, 24
Monistiancistrus, 289
Monistiancistrus carachama, 289
monitor, Amblydoras, 169monitor, Zathorax, 168, 169
monkei, Auchenoglanis, 163
monkei, Clarias, 141
monkei, Parauchenoglanis, 163mononema, Kryptopterus, 370
mononema, Silurus, 370
monopelte, Sturisoma, 299monsembeensis, Malapterurus, 302
montana, Cetopsis, 132montana, Clupisoma, 357
montana, Erethistoides, 384
montana, Pimelodella, 193montana, Rhamdella, 196montana, Xenocara, 223montana pipri, Erethistoides, 384

Zootaxa 1418 © 2007 Magnolia Press · 595

montanus, Ancistrus, 223montanus, Bagrus, 96
montanus, Mystus, 96
montanus dibrugarensis, Macrones, 96montebelloi, Canthopomus, 263
montebelloi, Ixinandria, 263
moolenburghae, Pseudeutropius, 362
moolenburghi, Hemisilurus, 369
moorei, Kryptopterus, 371
moorei, Micronema, 371
moorii, Clarias, 143
moravicus, Otolithus (Arius ?), 434
morehensis, Silurus, 376
morei, Opsodoras, 175
Morei, Oxydoras, 175
moreirai, Rhamdiopsis, 202, 203
morenoi, Cetopsidium, 130
morenoi, Hemicetopsis, 130
Mormyrostoma, 170
morrisi, Hemidoras, 172
morrowi, Hemiloricaria, 246morrowi, Liosomadoras, 75
morrowi, Rhineloricaria, 246
mortiauxi, Tanganikallabes, 152
mortoni, Neosilurus, 348
mossambicus, Clarias, 143
motaguensis, Pimelodus, 198
motatanensis, Cetopsis, 132
motatanensis, Pseudocetopsis
plumbeus, 132
motatanensis, Pygidium emanueli, 421
motatanensis, Trichomycterus, 421
mounseyi, Rhamdia, 201
mucosa, Pimelodella, 193
mucosa, Platystoma, 340
mucosus, Platysilurus, 340
mucronatum, Amblyceps, 18
muelleri, Paraplotosus, 349
muelleri, Rhamdia, 199
Mugil corsula, 89
mukherjii, Mystus, 95
mülleri, Clarias, 143mülleri, Cnidoglanis, 349
mülleri, Pimelodus, 199
Mülleri, Silurus, 380, 381
multilineatus, Eutropius, 365
multimaculata, Synodontis, 317
multimaculatus, Corydoras, 120
multimaculatus, Synodontis, 317
multimaculatus, Zungaropsis, 344
multipunctata, Amphilius
nigricaudatus, 21
multipunctata, Synodontis, 317
multipunctatus, Synodontis, 317
multiradiatus, Arius, 40
multiradiatus, Brochis, 120
multiradiatus, Cathorops, 40
multiradiatus, Chaenothorax, 120
multiradiatus, Corydoras, 120

multiradiatus, Cranoglanis, 165
multiradiatus, Heptapterus, 184
multiradiatus, Hypostomus, 290, 291
multiradiatus, Mystus, 96
multiradiatus, Pimelodus, 334
multiradiatus, Plotosus, 350
multiradiatus, Pseudeutropichthys, 165
multiradiatus, Pterygoplichthys, 291
multiradiatus alternans, Ancistrus, 291
multispinis, Ancistrus, 223
multispinis, Ancistrus, 265
multispinis, Xenocara, 223
multitaeniatus, Eutropius, 365
multitaeniatus, Schilbe, 365
munitus, Noturus, 214
mura, Otocinclus, 275
muriaensis, Parotocinclus, 283
murica, Doras, 177
muricata, Pseudolaguvia, 401
muriei, Rhamdia guatemalensis, 202
murinus, Spectracanthicus, 297
murius, Eutropiichthys, 357
murius, Pimelodus, 357
murius batarensis, Pseudeutropius,
357
murrayi, Malapterurus, 302
murraystuarti, Amblyceps, 18
murray-stuarti, Amblyceps, 18
musaica, Tatia, 77
musculus, Pimelodus, 200
mustelinus, Heptapterus, 184
mustelinus, Pimelodus, 184
muticus, Silurus, 337
mutisii, Eremophilus, 404
mutsindoziensis, Clariallabes, 137
mutuca, Hemipsilichthys, 280
mutuca, Pareiorhaphis, 280
mutucae, Hypostomus, 257
myanmar, Pangasius, 327
Myersglanis, 396
Myersglanis blythii, 396
Myersglanis jayarami, 396
myersi, Corydoras, 123
myersi, Hypostomus, 257
myersi, Leiocassis, 104
myersi, Leptodoras, 173
myersi, Plecostomus, 257
myersi, Pseudomystus, 104
Myoglanis, 187
Myoglanis aspredinoides, 187
Myoglanis koepckei, 187
Myoglanis potaroensis, 187
Myoglanis potaroënsis, 187
myriodon, Aposturisoma, 226
myriodon, Chrysichthys, 155
Mysoricus, Silurus, 374
mystacinus, Ancistrus, 265
mysteriosus, Pimelodus, 338
mysticetus, Mystus, 96

596 · Zootaxa 1418 © 2007 Magnolia Press

Mystus, 93
Mystus (Mystus) maydelli, 89
Mystus (Mystus) vittatus horai, 95
Mystus alasensis, 94
Mystus albolineatus, 94
Mystus armatus, 94, 98
Mystus armiger, 94
Mystus ascita, 177
Mystus atrifasciatus, 94
Mystus aubentoni, 91
Mystus bimaculatus, 94
Mystus bleekeri, 94
Mystus bocourti, 94
Mystus canarensis, 96
Mystus carolinensis, 37
Mystus castaneus, 94
Mystus cavasius, 95
Mystus dalungshanensis, 95
Mystus falcarius, 95
Mystus fluviatilis, 164
Mystus gulio, 95
Mystus havmolleri, 85
Mystus horai, 95
Mystus impluviatus, 95
Mystus johorensis, 89
Mystus krishnensis, 89
Mystus leucophasis, 96
Mystus malabaricus, 96
Mystus mica, 100
Mystus misrai, 97
Mystus montanus, 96
Mystus mukherjii, 95
Mystus multiradiatus, 96
Mystus mysticetus, 96
Mystus nigriceps, 96
Mystus oculatus, 96
Mystus olyroides, 90
Mystus pahangensis, 90
Mystus pelusius, 96
Mystus pulcher, 97
Mystus punctifer, 97
Mystus rhegma, 97
Mystus rufescens, 94, 97
Mystus sabanus, 91
Mystus seengtee, 97
Mystus singaringan, 97
Mystus spinipectoralis, 97
Mystus stigmaturus, 85
Mystus tengara, 97, 98
Mystus vittatus, 98
Mystus wolffii, 98
mystus, Schilbe, 365
mystus, Silurus, 362, 365
myzostoma, Euchiloglanis, 399
myzostomus, Pareuchiloglanis, 399
NN'gamensis, Ciarias, 146
N'gamensis, Clarias, 146
nahuelbutaensis, Diplomystes, 166

FERRARIS

naipi, Trichomycterus, 421
namdia, Pimelodus, 200
Nangra, 396
Nangra assamensis, 396
Nangra bucculenta, 396
Nangra Buchanani, 397
Nangra carcharhinoides, 396
Nangra nangra, 397
Nangra ornata, 397
Nangra punctata, 395
Nangra robusta, 397
nangra, Nangra, 397
nangra, Pimelodus, 396, 397
nannoculus, Microsynodontis, 310
Nannoglanis, 187
Nannoglanis bifasciatus, 203
Nannoglanis fasciatus, 187, 188
Nannoglanis hoehnei, 188
Nannoptopoma, 272
Nannoptopoma spectabile, 272
Nannoptopoma sternoptychum, 273
Nannorhamdia, 180, 185
Nannorhamdia benedettii, 198
Nannorhamdia guttatus, 185
Nannorhamdia lineata, 185
Nannorhamdia macrocephala, 192
Nannorhamdia nemacheir, 186
Nannorhamdia schubarti, 186
Nannorhamdia spurrellii, 185, 186
Nannorhamdia stictonotus, 186
Nanobagrus, 98
Nanobagrus armatus, 98
Nanobagrus nebulosus, 98
Nanobagrus stellatus, 98
nanonocticolus, Gelanoglanis, 74
nantoensis, Liobagrus, 19
nantoënsis, Liobagrus 19
nanus, Corydoras, 120
napoensis, Corydoras, 120
narcissus, Corydoras, 120
nasalis, Eutropius, 363
naso, Liocassis, 108
Nasocassis, 92
nassi, Phractocephalus, 336
nasus, Cetopsorhamdia, 181, 182
nasus, Pimelodina, 336
nasuta, Psammphiletria, 27
nasuta, Rhamdia, 201
nasutus, Arius, 49
nasutus, Microsynodontis, 310
nasutus, Pangasius, 327
nasutus, Pseudopangasius, 327
natalensis, Amphilius, 22
natalis, Ameiurus, 205
natalis, Pimelodus, 205
natalis analis, Amiurus, 205
nationi, Ancistrus, 223
nationi, Lasiancistrus, 223
nattereri, Corydoras, 121

nattereri, Farlowella, 238
Nattereri, Oxydoras, 179
nattereri, Trachydoras, 179
nattereri triseriatus, Corydoras, 121
nauticus, Amblydoras, 169
nauticus, Zathorax, 169
navarroi, Pimelodus, 338
navarroi, Pimelodus grosskopfii, 338
naziri, Clupisoma, 357
naziri, Glyptothorax, 392
Neblinichthys, 273
Neblinichthys pilosus, 273
Neblinichthys roraima, 273
Neblinichthys yaravi, 273
nebulifer, Pareuchiloglanis, 399
nebulosa, Synodontis, 317
nebulosus, Ameiurus, 205
nebulosus, Clarias, 148
nebulosus, Doras, 178
nebulosus, Ituglanis, 407
nebulosus, Nanobagrus, 98
nebulosus, Pimelodus, 205, 215
nebulosus, Silurus, 215
nebulosus, Silurus (Callichrus), 374
nebulosus, Synodontis, 317
nebulosus, Wallago, 381
nebulosus pannonicus, Ictalurus, 206
Nedystoma, 46
Nedystoma dayi, 46
Nedystoma novaeguineae, 46
negro, Corydoras, 121
neivai, Glanidium, 77
neivai, Tatia, 77
nella, Pimelodus, 52
nella, Plicofollis, 52
nelsoni, Conorhynchos, 53
nelsoni, Glyptothorax, 392
nelsoni, Leptodoras, 173
nelsoni, Potamarius, 53
nemacheir, Imparfinis, 186
nemacheir, Nannorhamdia, 186
Nemadoras, 174
Nemadoras elongatus, 174
Nemadoras hemipeltis, 174
Nemadoras humeralis, 174
Nemadoras leporhinus, 174
Nemadoras trimaculatus, 174
Nemapteryx, 46
Nemapteryx armiger, 46
Nemapteryx augusta, 46
Nemapteryx bleekeri, 46
Nemapteryx caelata, 46
Nemapteryx macronotacantha, 46
Nemapteryx nenga, 46, 47
Nemasiluroides, 361
Nemasiluroides furcatus, 361
Nematogenyidae, 323
Nematogenyini, 323
Nematogenys, 323, 324

CHECKLIST OF CATFISHES, RECENT AND FOSSIL

Nematogenys cuivi, 324
Nematogenys inermis, 324
Nematogenys nigricans, 324
Nematogenys pallidus, 324
nematophorus, Platystacus, 61
nematopterus, Hypostomus, 257
nematopteryx, Listrura, 408
Nemuroglanis, 180, 188
Nemuroglanis lanceolatus, 188
Nemuroglanis mariai, 188
Nemuroglanis pauciradiatus, 188
nemurus, Bagrus, 87, 90
nemurus, Hemibagrus, 90
nemurus, Pseudostegophilus, 412
nemurus, Stegophilus, 412
nenga, Nemapteryx, 46, 47
nenga, Pimelodus, 47
Neoarius, 33, 47
Neoarius berneyi, 47
Neoarius coatesi, 47
Neoarius graeffei, 47
Neoarius latirostris, 47
Neoarius leptaspis, 47
Neoarius midgleyi, 47
Neoarius paucus, 48
Neoarius pectoralis, 48
Neoarius taylori, 48
Neoarius utarus, 48
Neoarius velutinus, 48
Neobagrus, 19
Neobagrus fuscus, 19, 20
neogranatensis, Arius, 50
neogranatensis, Notarius, 50
Neopangasius, 325
Neopangasius Nieuwenhuisii, 325, 327
Neoplecostominae, 217
Neoplecostomus, 217, 273
Neoplecostomus espiritosantensis,
273
Neoplecostomus franciscoensis, 273
Neoplecostomus granosus, 273
Neoplecostomus microps, 273
Neoplecostomus paranensis, 273
Neoplecostomus ribeirensis, 274
Neoplecostomus variipictus, 274
Neoplotosus, 346
Neoplotosus waterhousii, 346
Neosiluroides, 347
Neosiluroides cooperensis, 347
Neosilurus, 347
Neosilurus ater, 347
Neosilurus argenteus, 345
Neosilurus australis, 347, 348
Neosilurus bartoni, 347
Neosilurus brevidorsalis, 347
Neosilurus coatesi, 348
Neosilurus equinus, 348
Neosilurus gjellerupi, 348
Neosilurus gloveri, 348

Zootaxa 1418 © 2007 Magnolia Press · 597

Neosilurus hyrtlii, 347, 348
Neosilurus idenburgi, 348
Neosilurus mediobarbis, 347
Neosilurus mollespiculum, 348
Neosilurus mortoni, 348
Neosilurus novaeguineae, 348
Neosilurus pseudospinosus, 349
Neosilurus rendahli, 345
Neosilurus robustus, 348
Neotropius, 356, 359
Neotropius acutirostris, 359
Neotropius atherinoides, 359
Neotropius khavalchor, 359
nephelion, Cordylancistrus, 443
Netuma, 48
Netuma aulometopon, 56
Netuma bilineata, 49
Netuma dubia, 54
Netuma hassleriana, 56
Netuma insularum, 50
Netuma mazatlana, 54
Netuma osakae, 49
Netuma planifrons, 44
Netuma proxima, 48
Netuma radiata, 435
Netuma thalassina, 48
Netuma thalassina jacksonensis, 49
netuma, Bagrus, 49
neumanni, Chiloglanis, 307
neumanni, Clarias, 145
newtoni, Otolithus (Arius), 435
ngamensis, Clarias, 146ngamensis, Auchenoglanis, 163
ngamensis, Parauchenoglanis, 163
ngola, Clarias, 142
nicaraguensis, Pimelodus, 199
nicaraguensis, Rhamdia, 199
nicefori, Astroblepus, 65
nicéfori, Astroblepus, 65
niceforoi, Hemiancistrus, 257
niceforoi, Hypostomus, 257
nickeriensis, Hypostomus, 257
nickeriensis, Loricaria, 269
nicoi, Acanthobunocephalus, 57
nicoi, Lasiancistrus, 289
nicoi, Pseudolithoxus, 289
nieuhofii, Clarias, 139, 146
Nieuwenhuisi, Glyptosternon, 392
nieuwenhuisi, Glyptothorax, 392
Nieuwenhuisii, Bagarius, 383
Nieuwenhuisii, Neopangasius, 325,
327
nieuwenhuisii, Pangasius, 327
niger, Bagrus docmac, 84
niger, Batasio, 445
niger, Chiloglanis, 307
niger, Copidoglanis novae-guineae,
349
niger, Doras, 175

niger, Doras (Oxydoras), 175
niger, Hemiancistrus, 287
niger, Hypostomus, 257
niger, Oxydoras, 175
niger, Plecostomus, 257
niger, Pseudancistrus, 287
nigeriae, Clarias, 141
nigeriensis, Clarias anguillaris, 140
Nigerium, 98
Nigerium gadense, 98
Nigerium wurnoense, 98, 99
nigrescens, Callichrous, 374
nigrescens, Pseudancistrus, 287
nigrescens, Silurus (Pimelodus), 210
nigribarbis, Parapimelodus, 335
nigribarbis, Pimelodus(Pseudorhamdia), 335
nigricans, Arius, 54
nigricans, Clarias, 146
nigricans, Hypostomus, 278
nigricans, Nematogenys, 324
nigricans, Pimelodus, 208, 209, 210
nigricans, Plotosus, 352
nigricans, Pseudoplatystoma
fasciatum, 341
nigricans, Trichomycterus, 414, 421
nigricauda, Hemiloricaria, 246
nigricauda, Hisonotus, 248
nigricauda, Liobagrus, 20
nigricauda, Loricaria, 246
nigricauda, Otocinclus, 248
nigricauda, Pseudopimelodus, 353
nigricaudatus, Amphilius, 21
nigricaudatus multipunctata,Amphilius, 21
nigricaudus, Cephalosilurus, 353
nigriceps, Bagrus, 96
nigriceps, Mystus, 96
nigricollaris, Horabagrus, 429
nigricollis, Tympanopleura, 68
nigrilabris, Gronias, 204, 206
nigripinnis, Auchenipterus, 72
nigripinnis, Euanemus, 72
nigripinnis, Microglanis, 354
nigrirostrum, Sturiosoma, 299
nigrita, Bagrus, 160
nigrita, Synodontis, 317
nigriventris, Synodontis, 318
nigrodigitatus, Chrysichthys, 158, 160
nigrodigitatus, Pimelodus, 158
nigrofasciata, Pimelodella, 193
nigrofasciatus, Pimelodus, 193
nigrolineata, Phenacorhamdia, 189
nigrolineatus, Chaetostomus, 277, 278
nigrolineatus, Panaque, 278
nigrolineatus, Pseudauchenipterus, 76
nigromaculata, Synodontis, 318
nigromaculatus, Hypostomus, 257
nigromaculatus, Plecostomus, 257

598 · Zootaxa 1418 © 2007 Magnolia Press

nigromaculatus, Synodontis, 318
nigromaculatus, Trichomycterus, 421
nigromarmoratus, Clarias, 146
nigropunctatus, Bagrus, 340
nijsseni, Corydoras, 121
nijsseni, Corydoras elegans, 121
nijsseni, Harttia, 272
nijsseni, Metaloricaria, 272
niloticus, Chiloglanis, 307
niloticus, Hypophthalmus, 362, 365
niloticus, Mochokus, 310, 311
niloticus, Silurus schilbe, 431
niloticus waterloti, Chiloglanis, 307
nimius, Hemipsilichthys, 247, 444
Niobichthys, 274
Niobichthys ferrarisi, 274
niobium, Glanapteryx, 405
nitidus, Akysis, 17, 82
nitidus, Leiocassis, 82
nitidus, Pelteobagrus, 82
nitidus, Pseudobagarius, 17
nitidus, Pseudobagrus, 82
niveata, Dekeyseria, 234
niveatus, Baryancistrus, 227
niveatus, Hypostomus, 226, 227
niveatus, Plecostomus, 234
niveiventris, Amiurus, 204
niveum, Chaetostoma, 230
njassae, Synodontis, 318
Nkondobabrus, 99
Nkondobagrus longisrostris, 99
nkunga, Plotosus, 351
nocturnus, Brachyglanis, 181
nocturnus, Noturus, 214
nocturnus, Panaqolus, 277
nocturnus, Panaque, 277
nodosus, Pseudauchenipterus, 76
nodosus, Silurus, 75, 76
noelkempffi, Corydoras, 121
nops, Gymnallabes, 150
normani, Chiloglanis, 307
Notarius, 49, 440
Notarius armbrusteri, 440
Notarius biffi, 49
Notarius cookei, 50
Notarius grandicassis, 50
Notarius insculptus, 50
Notarius kessleri, 50
Notarius neogranatensis, 50
Notarius osculus, 50
Notarius planiceps, 50
notata, Microsynodontis, 310
notata, Synodontis, 318
notatus, Amphilius, 27
notatus, Callichrous, 374
notatus, Hisonotus, 248
notatus, Microsynodontis, 310
notatus, Pimelodus, 210, 339
notatus, Platynematichthys, 339

FERRARISnotatus, Synodontis, 318
notatus, Tetracamphilius, 27
notatus binotata, Synodontis, 318
notatus ocellatus, Synodontis, 318
notatus var. binotata, Synodontis, 318
notatus var. ocellatus, Synodontis, 318
notialis, Parakysis, 16
Notoglanidium, 161, 162
Notoglandium maculatus, 161
Notoglanidium pallidum, 162
Notoglanidium thomasi, 162
Notoglanidium walkeri, 162
notomelas, Pimelodella, 193
Notophthalmus, 333
notospilus, Hemidoras, 171
notozygurus, Clarias, 143
Noturus, 211
Noturus albater, 211
Noturus baileyi, 211
Noturus crypticus, 211, 442
Noturus elassochir, 212
Noturus elegans, 212
Noturus eleutherus, 212
Noturus exilis, 212
Noturus fasciatus, 212
Noturus flavater, 212
Noturus flavipinnis, 212
Noturus flavus, 211, 212
Noturus funebris, 212
Noturus furiosus, 211, 212
Noturus gilberti, 213
Noturus gladiator, 213
Noturus gyrinus, 213
Noturus hildebrandi, 213
Noturus hildebrandi lautus, 213
Noturus insignis, 213
Noturus lachneri, 213
Noturus latifrons, 211
Noturus leptacanthus, 213
Noturus luteus, 212
Noturus marginatus, 213
Noturus maydeni, 441
Noturus miurus, 214
Noturus munitus, 214
Noturus nocturnus, 214
Noturus occidentalis, 212
Noturus phaeus, 214
Noturus placidus, 214
Noturus platycephalus, 212
Noturus sialis, 213
Noturus stanauli, 214, 442
Noturus stigmosus, 214
Noturus taylori, 214
Noturus trautmani, 214
novaeguineae, Nedystoma, 46
novaeguineae, Neosilurus, 348
novaesi, Furcodontichthys, 240
novae-guineae, Copidoglanis, 348
novae-guineae, Doiichthys, 46

novae-guineae niger, Copidoglanis,
349
novalimensis, Harttia, 241nox, Arius, 38, 39
nox, Brustiarius, 38
ntemensis, Synodontis Hollyi, 322
nuchalis, Ælurichthys, 38
nuchalis, Arius, 41
nuchalis, Auchenipterus, 72
nuchalis, Hypophthalmus, 71, 72
nucleus, Arius, 57
nudiceps, Ancistrus, 223
nudiceps, Cnidoglanis, 346
nudiceps, Euristhmus, 346
nudiceps, Hypostomus, 223
nudiceps, Pelteobagrus, 100
nudiceps, Pseudobagrus, 100
nudidens, Arius (Hemiarius), 56
nudipectus, Belonoglanis, 24
nudirostre, Chaetostoma, 230
nudirostris, Chaetostomus, 230
nudiventris, Hypostomus, 257
nudirostris, Loricaria, 271
nudirostris, Loricariichthys, 271
nudiventris, Plecostomus, 257
nudiventris, Loricaria, 297
nudiventris, Parancistrus, 279
nudiventris, Spatuloricaria, 297
nudulus, Hemipsilichthys, 280
nudulus, Pareiorhaphis, 280
nujiangense, Clupisoma, 357
nummifer, Synodontis, 318
nyasensis, Clarias, 149nyasensis, Dinotopterus, 149
nyongensis, Chrysichthys, 158
nyongensis, Chrysichthys
longidorsalis, 158
nyongensis, Eutropius, 365
nyongensis, Schilbe, 365ñssipinnis, Heptapterus, 184O
oaxacae, Rhamdia, 201
obbesi, Porochilus, 351
obesa, Rhamdia, 199
obesus, Amiurus, 205
obesus, Arius, 81
obesus, Liobagrus, 20
obesus, Synodontis, 318, 323
oblongus, Tachysurus, 435
obscura, Glyptothorax, 392
obscurum, Pogonopoma, 285
obscurus, Acrochordonichthys 13
obscurus, Auchenipterus, 81
obscurus, Bagrichthys, 83
obscurus, Clarias, 141
obscurus, Copidoglanis, 345, 350
obscurus, Glyptothorax, 392
obtusa, Pseudotothyris, 290
obtusirostris, Eutropius, 365

CHECKLIST OF CATFISHES, RECENT AND FOSSIL

obtusirostris, Plecostomus, 218
obtusos, Otocinclus, 290
obtusus, Otocinclus, 290
occidentalis, Ailia, 360
occidentalis, Ancistrus, 223
occidentalis, Auchenoglanis, 154
occidentalis, Cetopsis, 134, 135
occidentalis, Chiloglanis, 307
occidentalis, Hypostomus, 257
occidentalis, Hypostomus
gymnorhynchus, 257
occidentalis, Istlarius balsanus, 208
occidentalis, Malapterurus, 302
occidentalis, Noturus, 212
occidentalis, Parailia, 360
occidentalis, Pimelodus, 154
occidentalis, Xenocara, 223
occidentalis tanganicanus,
Auchenoglanis, 154
occidentalis tchadiensis,
Auchenoglanis, 154
occidentalis var. tanganicanus,
Auchenoglanis, 154
occidentalis var. tchadiensis,
Auchenoglanis, 154
occloi, Ancistrus, 223
ocellatus, Galeichthys, 43
ocellatus, Silurus, 35
ocellatus, Synodontis notatus, 318
ocellifer, Synodontis, 318
Ochmacanthus, 409
Ochmacanthus alternus, 409
Ochmacanthus batrachostoma, 409
Ochmacanthus flabelliferus, 409
Ochmacanthus orinoco, 409
Ochmacanthus reinhardtii, 410
Ochmacanthus taxistigma, 405, 406
ochoterenai, Haustor, 209ochoterenai, Ictalurus, 209
octocirrhus, Pimelodus, 431
octocirrus, Corydoras, 117
Octonematichthys, 160
oculatus, Bagrus, 96
oculatus, Mystus, 96
oculeus, Hypostomus, 257
oculeus, Panaque, 257
odontotumulus, Farlowella, 238
odynea, Pimelodella, 193
odynea, Pimelodella chagresi, 193
oelemariensis, Corydoras, 114
oetik, Arius, 35
ogilviei, Ageneiosus, 69
ogilviei, Opsodoras, 174, 175
ogoensis, Gephyroglanis, 158
ogooensis, Chrysichthys, 158
ogooensis, Gephyroglanis, 158
ogooensis, Malapterurus electricus,
var., 302
ogowensis, Chrysichthys, 158

Zootaxa 1418 © 2007 Magnolia Press · 599

oguensis, Malapterurus, 302
oguensis, Malapterurus electricus, 302
oguensis, Malapterurus electricus var.,302
oiapoquensis, Corydoras, 121
oibaensis, Callichthys, 111
okadai, Coreobagrus, 86
okae, Chrysichthys, 158
okeechobeensis, Ictalurus, 210
olallae, Duopalatinus, 340
olallae, Platysilurus, 340
Oligancistrus, 274
Oligancistrus punctatissimus, 274
oligospila, Peckoltia, 284
oligospilus, Chaetostomus, 284
olivaceus, Chlarias, 146
olivaceus, Clarias, 146
olivaceus, Pimelodus, 210
Olivaichthys, 166
Olivaichthys cuyanus, 166
Olivaichthys mesembrinus, 166
Olivaichthys viedmensis, 166
olivaris, Pylodictis, 215
olivaris, Silurus, 215
oliveirae, Farlowella, 237
oliveirai, Bathycetopsis, 131, 132
oliveirai, Cetopsis, 132
Oloplotosus, 349
Oloplotosus luteus, 349
Oloplotosus mariae, 349
Oloplotosus torobo, 349
Olyra, 81, 99
Olyra burmanica, 99
Olyra elongata, 99
Olyra horae, 99
Olyra horai, 99
Olyra kempi, 99
Olyra laticeps, 17, 18
Olyra longicaudata, 99
Olyra longicaudatus, 99
Olyrinae, 81
olyroides, Hemibagrus, 90
olyroides, Mystus, 90
omeihensis, Leiocassis, 102
omeihensis, Pseudobagrus, 102
omias, Synodontis, 319
ommation, Apistoloricaria, 226
Ompok, 371
Ompok bimaculatus, 372
Ompok binotatus, 372
Ompok borneensis, 372
Ompok fumidus, 372
Ompok hypophthalmus, 371, 372
Ompok jaynei, 372
Ompok leiacanthus, 371, 372
Ompok malabaricus, 372
Ompok miostomus, 372
Ompok pabda, 372
Ompok pabo, 373

Ompok pinnatus, 373
Ompok platyrhynchus, 373
Ompok pluriradiatus, 373
Ompok rhadinurus, 373
Ompok sabanus, 371
Ompok siluroides, 371
Ompok siluroïdes, 373
Ompok sindensis, 373
Ompok urbaini, 373
Ompok weberi, 373
oncina, Arius, 75
oncinus, Arius, 75
oncinus, Liosomadoras, 75
ondon, Pseudobagrus, 102
operculatum, Scleronema, 413
ophthalmica, Pimelodella, 193
ophthalmicus, Pimelodus, 194
ophthalmus, Doras (Corydoras), 169
opisthophthalmus, Amphilius, 22
Opladelus, 215
Opsodoras, 175
Opsodoras boulengeri, 172, 175
Opsodoras hemipeltis, 174
Opsodoras morei, 175
Opsodoras ogilviei, 174, 175
Opsodoras orthacanthus, 175
Opsodoras parallelus, 174
Opsodoras steindachneri, 169
Opsodoras stuebelii, 175
Opsodoras ternetzi, 175
Opsodoras trimaculatus, 174
orbignianus, Platystoma, 341
orbignyanum, Platystoma, 341
orcesi, Corydoras pastazensis, 122oregonensis, Hypsidoris, 203oremaculatus, Hypophthalmus, 333
Oreoglanis, 397
Oreoglanis delacouri, 397
Oreoglanis frenatus, 397
Oreoglanis hypsiurus, 397
Oreoglanis infulatus, 397
Oreoglanis insignis, 397
Oreoglanis lepturus, 397
Oreoglanis macronemus, 397
Oreoglanis macropterus, 398
Oreoglanis setiger, 398
Oreoglanis siamensis, 397, 398
orestes, Oxydoras, 171
Orestis, Oxydoras, 171orientale, Cetopsidium, 130orientale, Pseudocetopsis, 130orientalis, Arges, 65
orientalis, Astroblepus, 65
orientalis, Bagrus, 84orientalis, Irvineia, 358
orinocensis, Hildadoras, 175, 176orinoco, Cetopsis, 132orinoco, Cetopsorhamdia, 182
orinoco, Lithoxancistrus, 286, 287

600 · Zootaxa 1418 © 2007 Magnolia Pressorinoco, Ochmacanthus, 409
orinoco, Pseudancistrus, 287
orinoco, Pseudocetopsis plumbeus,
132
Orinocodoras, 175
Orinocodoras eigenmanni, 175
ornata, Nangra, 397
ornaticeps, Heptapterus, 185
ornatipinnis, Synodontis, 319
ornatissima, Synodontis, 319
ornatissimus, Synodontis, 319
ornatus, Chrysichthys, 158
ornatus, Clarias, 145
ornatus, Corydoras, 121ornatus, Gogo, 29
ornatus, Hyalobagrus, 92
ornatus, Pimelodus, 338
ornatus, Pseudobagrus, 92
ornatus, Pseudomystus, 104ornatus, Synodontis, 317, 319
oronocoi, Hoplosternum, 129
orontis, Clarias, 143
oroyae, Pygidium, 423
orphnopterus, Corydoras, 121
orthacanthus, Opsodoras, 175
orthiocarinatus, Copionodon, 404
ortmanni, Pimelodus, 338
ortoni, Rhamdia, 201
ortoni, Sorubimichthys, 343
osakae, Netuma, 49
Oschanini, Exostoma, 402
osculus, Arius, 50
osculus, Notarius, 50
Osteobagrus, 106
osteocarus, Corydoras, 121
Osteogaster, 112
Osteogeneiosus, 51
Osteogeneiosus blochii, 51
Osteogeneiosus cantoris, 51
Osteogeneiosus gracilis, 51
Osteogeneiosus ingluvies, 51
Osteogeneiosus longiceps, 51
Osteogeneiosus macrocephalus, 51
Osteogeneiosus militaris, 51
Osteogeneiosus valenciennesi, 51
Osteogeniosus sthenocephalus, 51
Osteogeniosus stenocephalus, 51
Osteomystax, 71
osteomystax, Auchenipterus, 72
osteomystax, Ceratocheilus, 71, 72
Ostophycephalus, 346
Ostophycephalus duriceps, 346
othonops, Cetopsis, 132
othonops, Hemicetopsis, 132
Otocinclini, 217
Otocinclus, 217, 264, 271, 274
Otocinclus (Microlepidogaster)tietensis, 290
Otocinclus affinis, 271

FERRARIS

Otocinclus arnoldi, 271
Otocinclus bororo, 274
Otocinclus caxarari, 274
Otocinclus cephalacanthus, 276
Otocinclus cocama, 274
Otocinclus depressicauda, 248
Otocinclus fimbriatus, 271
Otocinclus flexilis, 271, 272
Otocinclus francirochai, 248
Otocinclus gibbosus, 264
Otocinclus hasemani, 274
Otocinclus hoppei, 275
Otocinclus huaorani, 275
Otocinclus joberti, 250
Otocinclus leucofrenatus, 248
Otocinclus macrospilus, 275
Otocinclus maculicauda, 281, 283
Otocinclus maculipinnis, 248
Otocinclus mariae, 275
Otocinclus mimulus, 275
Otocinclus mura, 275
Otocinclus nigricauda, 248
Otocinclus obtusos, 290
Otocinclus obtusus, 290
Otocinclus paulinus, 249
Otocinclus spectabilis, 273
Otocinclus tapirape, 275
Otocinclus vestitus, 274, 275
Otocinclus vittatus, 275
Otocinclus xakriaba, 275
Otolithus (Arius) aequus, 435
Otolithus (Arius) africanus, 435
Otolithus (Arius) amekiensis, 435
Otolithus (Arius) angelicus, 434
Otolithus (Arius) angulatus, 435
Otolithus (Arius) crassus bartonensis,
434
Otolithus (Arius) danicus, 434
Otolithus (Arius) danicus bartonensis,
434
Otolithus (Arius) decipiens, 434
Otolithus (Arius) germanicus, 434
Otolithus (Arius ?) glaber, 435
Otolithus (Arius) jaekeli, 435
Otolithus (Arius) Lerichei, 434
Otolithus (Arius ?) moravicus, 434
Otolithus (Arius) newtoni, 435
Otolithus (Arius ?) parvus, 435
Otolithus (Arius) planus, 435
Otolithus (Arius) rotundatus, 435
Otolithus (Arius) tenius, 435
Otolithus (Arius) tenuis, 435
Otolithus (Arius) vanigonis, 434
Otolithus (incertae sedis) crassus, 434
Otolithus (Sciaenidarum) decipiens,
434
Otolithus (Siluridarum ?) incertus, 434
Otothyrini, 217, 440
Otothyris, 217, 275

Otothyris canaliferus, 275, 276
Otothyris juquiae, 276
Otothyris lophophanes, 276
Otothyris rostrata, 276
Otothyris travassosi, 276
Otothyropsis, 276
Otothyropsis marapoama, 276
ourastigma, Corydoras, 121
ovidius, Synodontis, 316
oxycephalus, Clarias, 146
Oxydoras, 175
Oxydoras (Rhinodoras) amazonum
hasemani, 170
Oxydoras (Rhinodoras) huberi, 174
Oxydoras acipenserinus, 172
Oxydoras affinis, 171
Oxydoras bachi, 174
Oxydoras eigenmanni, 167
Oxydoras elongatus, 174
Oxydoras holdeni, 176
Oxydoras kneri, 175
Oxydoras Morei, 175
Oxydoras Nattereri, 179
Oxydoras niger, 175
Oxydoras orestes, 171
Oxydoras Orestis, 171
Oxydoras sifontesi, 176
Oxydoras steindachneri, 179
Oxydoras Stübelii, 175
Oxydoras trachyparia, 179
Oxydoras trimaculatus, 174
Oxyglanis, 154
Oxyglanis sacchii, 154
Oxyloricaria, 298
Oxyloricaria citurensis, 300
Oxyloricaria dariensis, 299
Oxyloricaria Fowleri, 233
Oxyloricaria guentheri, 299
Oxyloricaria leightoni, 300
Oxyloricaria lyra, 299
Oxyloricaria robusta, 299
Oxyloricaria tamanae, 300
Oxyloricaria tenuirostris, 300
oxyptera, Paravandellia, 410
oxyrhinus, Amphilius, 23
oxyrhynchus, Anduzedoras, 169
oxyrhynchus, Corydoras, 121
Oxyrhynchus, Doras, 169
Oxyropsis, 276
Oxyropsis acutirostris, 276
Oxyropsis carinata, 276
Oxyropsis wrightiana, 276
Oxyropsis wrightii, 276
oxyrrhyncha, Acestra, 239
oxyrryncha, Acestra, 239
oxyrryncha, Farlowella, 239
P
pabda, Ompok, 372
pabda, Silurus, 371, 372

CHECKLIST OF CATFISHES, RECENT AND FOSSIL

pabo, Ompok, 373
pabo, Silurus, 373
pachychilus, Atopochilus, 304
pachyderma, Acrochordonichthys, 13
pachynema, Ceratoglanis, 368
pachynemus, Ceratoglanis, 368
pachynema, Clarias, 146
Pachypterus, 359
Pachypterus luridus, 327
Pachypterus melanurus, 358
Pachypterus punctatus, 358
Pachypterus trifasciatus, 359
Pachyula, 42
pacifici, Cruciglanis, 443
pagei, Hypostomus, 257
pahangensis, Mystus, 90
pakistanica, Gagata, 386
pakistanicus, Batasio, 85
palavanensis, Penesilurus, 375, 377
palæindicus, Heterobranchus, 151
paleatus, Callichthys, 122
paleatus, Corydoras, 122
palembangensis, Kryptopterus, 370
palembangensis, Silurus, 369, 370
palleus, Trichomycterus, 415
pallida, Pimelodella, 194
pallida, Sosia chamaeleon, 13
pallidimaculatus, Lithoxus, 267
pallidimaculatus, Lithoxus(Paralithoxus), 267
pallidum, Notoglanidium, 162
pallidus, Nematogenys, 324
pallidus, Silurus, 210
pallidus Lateralis, Silurus, 210
pallidus Leucoptera, 210
pallidus Marginatus, Silurus, 210
pallidus var. Lateralis, Pimelodus, 210
pallidus var. Lateralis, Silurus, 210
pallidus var. Leucoptera, Pimelodus,
210
pallidus var. Leucoptera, Silurus, 210
pallidus var. Marginata, Silurus, 210
pallidus var. Marginatus, Pimelodus,
210
pallidus var. Marginatus, Silurus, 210
pallozonum, Glyptosternum, 388, 392
pallozonus, Glyptothorax, 392
palmeri, Chaetostoma, 230
palmeri, Chaetostomus 230
palmeri, Schilbe, 364
panamense, Sturisoma, 299
panamensis, Aelurichthys, 37, 38
panamensis, Bagre, 38
panamensis, Imparales, 180
panamensis, Loricaria, 299
panamensis, Plecostomus plecostomus,
243
Panaqolus, 277
Panaqolus albomaculatus, 277

Zootaxa 1418 © 2007 Magnolia Press · 601

Panaqolus changae, 277
Panaqolus dentex, 277
Panaqolus gnomus, 277
Panaqolus maccus, 277
Panaqolus nocturnus, 277
Panaqolus purusiensis, 277
Panaque, 277
Panaque albomaculatus, 277
Panaque changae, 277
Panaque cochliodon, 277
Panaque dentex, 277
Panaque gnomus, 277
Panaque maccus, 277
Panaque nigrolineatus, 278
Panaque nocturnus, 277
Panaque oculeus, 257
Panaque purusiensis, 277
Panaque suttoni, 278
Panaque suttonorum, 278
panda, Corydoras, 122
panda, Glyptothorax, 392
Pangasianodon, 324
Pangasianodon gigas, 325
Pangasianodon hypophthalmus, 325
Pangasianodonidi, 324
Pangasiidae, 324
Pangasini, 324
Pangasius, 324, 325
Pangasius (Pseudopangasius)
bocourti, 325
Pangasius aequilabialis, 327
Pangasius altifrons, 325
Pangasius beani, 328
Pangasius bedado, 325
Pangasius bocourti, 325
Pangasius Buchanani, 325, 327
Pangasius burgini, 326
Pangasius conchophilus, 325
Pangasius cultratus, 328
Pangasius de Zwaani, 328
Pangasius delicatissimus, 328
Pangasius djambal, 325, 326
Pangasius elongatus, 326
Pangasius fowleri, 328
Pangasius hexanema, 358
Pangasius hoeksi, 328
Pangasius humeralis, 326
Pangasius indicus, 326
Pangasius juaro, 327
Pangasius kinabatanganensis, 326
Pangasius krempfi, 326
Pangasius kunyit, 326
Pangasius larnaudii, 326
Pangasius lithostoma, 326
Pangasius longibarbis, 359
Pangasius macronema, 327
Pangasius mahakamensis, 327
Pangasius mekongensis, 327
Pangasius micronemus, 328

Pangasius myanmar, 327
Pangasius nasutus, 327
Pangasius nieuwenhuisii, 327
Pangasius pangasius, 327
Pangasius pangasius godavarii, 327
Pangasius pangasius upiensis, 327
Pangasius paucidens, 325
Pangasius pleurotaenia, 328
Pangasius polyuranodon, 325, 327
Pangasius ponderosus, 327
Pangasius rheophilus, 328
Pangasius rios, 328
Pangasius sabahensis, 328
Pangasius sanitwongsei, 328
Pangasius siamensis, 327
Pangasius sutchi, 325
Pangasius taeniura, 326
Pangasius tubbi, 328
panjang, Hemileiocassis, 91
pannonicus, Ictalurus nebulosus, 206
pantanalensis, Corydoras, 122
panthale, Platystoma, 341, 342
pantherinus, Auchenoglanis, 163
pantherinus, Eurycheilichthys, 236
pantherinus, Eurycheilus, 236
pantherinus, Hypostomus, 258
pantherinus, Parauchenoglanis, 163
pantherinus, Pseudariodes, 339
pantherinus, Synodontis, 315
pantherinus, Trichomycterus, 421
panzeri, Henonemus, 414
panzeri, Stegophilus, 414
paolence, Pygidium, 421
paolencis, Trichomycterus, 421
papariae, Hypostomus, 258
papariae, Plecostomus plecostomus,
258
papariae, Pseudancistrus, 287
papariae, Rhamdella, 197
papilionatus, Doras, 173
papillatus, Hemipsilichthys, 248, 444
papillatus, Hoplomyzon, 61
papillifer, Hemipimelodus, 48
papilliferus, Trichomycterus, 421
papillosus, Arius, 165
pappenheimi, Pimelodella, 194
papuensis, Copidoglanis, 348
papuensis, Plotosus, 351
paquequerense, Pygidium, 421
paquequerensis, Trichomycterus, 421
Parabranchioica, 410
Parabranchioica teaguei, 411
Paracanthopoma, 410
Paracanthopoma parva, 410
Paracetopsis, 134
Paracetopsis atahualpa, 135
Paracetopsis bleekeri, 134, 135
Paracetopsis esmeraldas, 135
Parachiloglanis, 398

602 · Zootaxa 1418 © 2007 Magnolia Press

Parachiloglanis hodgarti, 398
Paradiplomystes, 37
Paradoxoglanis, 302
Paradoxoglanis caudivittatus, 302
Paradoxoglanis cryptus, 303
Paradoxoglanis parvus, 303
paraense, Brachyplatystoma, 331
Paraglyptothorax, 388
paragua, Corydoras, 122paraguayensis, Farlowella, 239
paraguayensis, Hemidoras, 179
paraguayensis, Trachydoras, 179
paraguensis, Rhinolepis, 275
Parahemiodon, 269
Parahemiodon chanjoo, 270
Parahemiodon typus, 269, 270
parahemiodon, Loricaria, 270
parahybae, Delturus, 234, 235, 444
parahybae, Ituglanis, 407
parahybae, Microglanis, 354
parahybae, Pimelodus (Rhamdia), 201
Parahybae, Platystoma, 344
parahybae, Pogonopoma, 285
parahybae, Pseudopimelodus, 354
parahybae, Pseudotocinclus, 290
parahybae, Pygidium proops, 406, 407
Parahybae, Rhinelepis, 285
parahybae, Steindachneridion, 344
Parailia, 360
Parailia congica, 360
Parailia longifilis, 360
Parailia occidentalis, 360
Parailia pellucida, 360
Parailia somalensis, 360
Parailia spiniserrata, 360
Parakysidae, 12
Parakysis, 12, 15
Parakysis anomalopteryx, 15
Parakysis grandis, 15
Parakysis longirostris, 16
Parakysis notialis, 16
Parakysis verrucosa, 15, 16
Parakysis verrucosus, 16
Paralithoides, 267
Paralithoxus, 267
parallelus, Corydoras, 122parallelus, Opsodoras, 174
Paraloricaria, 278
Paraloricaria agastor, 278
Paraloricaria commersonoides, 278
Paraloricaria vetula, 278
Paramphilius, 25
Paramphilius firestonei, 25
Paramphilius goodi, 25
Paramphilius teugelsi, 25
Paramphilius trichomycteroides, 25paranaënse, Farlowella, 237
paranaensis, Pimelodus, 338
parananus, Megalancistrus, 272

FERRARIS

parananus, Pterygoplichthys(Ancistrus), 272
Parancistrus, 278
Parancistrus aurantiacus, 278
Parancistrus nudiventris, 279
paranensis, Hypostomus, 258
paranensis, Neoplecostomus, 273
paranensis, Parapterodoras, 177
Paraotocinclus cesarpintoi, 282
Paraphractura, 26
Paraphractura tenuicauda, 26, 27
Parapimelodus, 335
Parapimelodus nigribarbis, 335
Parapimelodus valenciennis, 335
Paraplotosus, 349
Paraplotosus albilabris, 349
Paraplotosus butleri, 349
Paraplotosus muelleri, 349
Parapseudecheneis, 400
Parapterodoras, 177
Parapterodoras paranensis, 177
Pararius, 48
paraschilbeides, Kryptopterus, 370
Parasilurus, 378
Parasilurus asotus longus, 379
Parasilurus biwaensis, 379
Parasilurus lithophilus, 380
Parasilurus microdorsalis, 380
Parastegophilus, 410
Parastegophilus maculatus, 410
Parastegophilus paulensis, 410
Parasturisoma, 298
Parasturisoma maculata, 233
paratus, Chiloglanis, 307
Parauchenipterus, 79
Parauchenipterus paseae, 79
Parauchenoglanis, 162
Parauchenoglanis ahli, 162
Parauchenoglanis altipinnis, 162
Parauchenoglanis ansorgii, 154
Parauchenoglanis balayi, 162
Parauchenoglanis boutchangai, 154
Parauchenoglanis buettikoferi, 163
Parauchenoglanis longiceps, 163
Parauchenoglanis monkei, 163
Parauchenoglanis ngamensis, 163
Parauchenoglanis pantherinus, 163
Parauchenoglanis punctatus, 163
Paravandellia, 410
Paravandellia oxyptera, 410
Paravandellia phaneronema, 411
pardale, Platystoma, 341
pardalis, Ageneiosus, 70
pardalis, Hypostomus, 292
pardalis, Platystoma, 341, 342
pardalis, Pterygoplichthys, 292
pardalis, Synodontis, 319
Pardiglanis, 163
Pardiglanis tarabinii, 163

pardus, Trichomycterus, 423
parecis, Ancistrus, 224pareiacantha, Loricaria, 295
pareiacantha, Rineloricaria, 295
Pareiodon, 403, 411
Pareiodon microps, 411
Pareiodontinae, 403
Pareiorhaphis, 247, 279
Pareiorhaphis alipionis, 262
Pareiorhaphis azygolechis, 279
Pareiorhaphis bahianus, 279
Pareiorhaphis cameroni, 279
Pareiorhaphis cerosus, 279
Pareiorhaphis eurycephalus, 279
Pareiorhaphis garbei, 279
Pareiorhaphis hypselurus, 280
Pareiorhaphis hystrix, 280
Pareiorhaphis mutuca, 280
Pareiorhaphis nudulus, 280
Pareiorhaphis parmula, 280
Pareiorhaphis regani, 280
Pareiorhaphis splendens, 280
Pareiorhaphis steindachneri, 280
Pareiorhaphis stephanus, 280
Pareiorhaphis stomias, 280
Pareiorhaphis vestigipinnis, 281
Pareiorhina, 281
Pareiorhina brachyrhyncha, 281
Pareiorhina carrancas, 281
Pareiorhina rudolphi, 281
Pareuchiloglanis, 398
Pareuchiloglanis anteanalis, 398
Pareuchiloglanis feae, 398
Pareuchiloglanis gongshanensis, 398
Pareuchiloglanis gracilicaudatus, 398
Pareuchiloglanis kamengensis, 399
Pareuchiloglanis longicauda, 399
Pareuchiloglanis macropterus, 399
Pareuchiloglanis macrotremus, 399
Pareuchiloglanis myzostomus, 399
Pareuchiloglanis nebulifer, 399
Pareuchiloglanis poilanei, 398, 399
Pareuchiloglanis rhabdurus, 399
Pareuchiloglanis robusta, 399
Pareuchiloglanis robustus, 399
Pareuchiloglanis sichuanensis, 399
Pareuchiloglanis sinensis, 399
Pareuchiloglanis songdaensis, 400
Pareuchiloglanis songmaensis, 400
Pareuchiloglanis tianquanensis, 400
Pareutropius, 360
Pareutropius buffei, 360
Pareutropius debauwi, 361
Pareutropius longifilis, 361
Pareutropius mandevillei, 361
Pareutropius micristius, 360, 361
Parexostoma, 387
Parexostoma maculatum, 387
pariolispos, Scobinancistrus, 296

CHECKLIST OF CATFISHES, RECENT AND FOSSIL

Pariolius, 188
Pariolius armillatus, 188
parkeri, Aspistor, 36
parkeri, Silurus, 36
parkii, Arius, 32
parkoi, Ituglanis, 407
parkoi, Pygidium, 407
parma, Cetopsis, 132
parmocassis, Arius, 50
parnaguensis, Ageneiosus, 70
parnahybae, Brachyplatystoma, 331
parnahybae, Loricaria, 269
parnahybae, Loricariichthys, 266
parnahybae, Pimelodella, 194
parnaibae, Glyptoperichthys, 292
parnaibae, Pterygoplichthys, 292
Paroreoglanis, 397
Paroreoglanis delacouri, 397
Parotocinclus, 281
Parotocinclus amazonensis, 281
Parotocinclus aripuanensis, 281
Parotocinclus bahiensis, 281
Parotocinclus bidentatus, 282
Parotocinclus britskii, 282
Parotocinclus cearensis, 282
Parotocinclus cesarpintoi, 282
Parotocinclus collinsae, 282
Parotocinclus cristatus, 282
Parotocinclus doceanus, 282
Parotocinclus eppleyi, 282
Parotocinclus haroldoi, 282
Parotocinclus jimi, 282
Parotocinclus jumbo, 282
Parotocinclus longirostris, 282
Parotocinclus maculicauda, 283
Parotocinclus minutus, 283
Parotocinclus muriaensis, 283
Parotocinclus planicauda, 283
Parotocinclus polyochrus, 283
Parotocinclus prata, 283
Parotocinclus spilosoma, 283
Parotocinclus spilurus, 283
Parotocinclus steindachneri, 283
Parrae, Galeichthys, 38
parryi, Rhamdia, 199
parva, Hemiloricaria, 246
parva, Loricaria, 246
parva, Paracanthopoma, 410
parva, Pimelodella, 194
parva, Pseudotatia, 77
parvanalis, Kryptopterus, 375
parvanalis, Phalacronotus, 375
parvicarinata, Farlowella, 239
parvimanus, Clarias, 140
parvipinnis, Arius, 47
parvus, Otolithus (Arius ?), 435
parvus, Paradoxoglanis, 303
parvus, Pimelodus (Rhamdia), 180
paseae, Parauchenipterus, 79

Zootaxa 1418 © 2007 Magnolia Press · 603

passany, Bagrus, 54
passany, Sciades, 54
passarellii, Homodiaetus, 406
passarellii, Stegophilus, 406
passensis, Ituglanis, 408
pastazensis, Corydoras, 122
pastazensis orcesi, Corydoras, 122
patagoniensis, Hatcheria, 405
pataxo, Microglanis, 444
pati, Luciopimelodus, 334
pati, Pimelodus, 334
pati, Silurus, 335
patiae, Chaetostoma, 230
pattersoni, Trogloglanis, 216
paucerna, Corydoras, 122
paucidens, Metaloricaria, 272
paucidens, Pangasius, 325
paucimaculatus, Hypostomus, 258
paucipunctatus, Hypostomus, 258
pauciradiatum, Megalonema, 335
pauciradiatus, Aspidoras, 109
pauciradiatus, Corydoras, 109
pauciradiatus, Nemuroglanis, 188
pauciradiatus, Trichomycterus, 440
paucispinis, Chaetostoma, 230
paucispinis, Chaetostomus, 230
paucisquamatus, Megalodoras, 173
paucus, Arius, 48
paucus, Neoarius, 48
paulensis, Parastegophilus, 410
paulensis, Pseudostegophilus, 410
Paulicea, 344
Paulicea jahu, 344
paulina, Loricaria, 294
paulinus, Hisonotus, 249
paulinus, Hypostomus, 258
paulinus, Otocinclus, 249
paulinus, Plecostomus, 258
paviei, Pseudecheneis, 400
pavimentatus, Arius, 105
paynei, Mochokiella, 310
paysanduanus, Auchenipterus, 72
pearsei, Chaetostoma, 230
pearsei, Chaetostomus, 230
pearsoni, Cetopsis, 132
pearsoni, Tridentopsis, 426
Peckoltia, 283
Peckoltia arenaria, 284
Peckoltia bachi, 284
Peckoltia braueri, 284
Peckoltia brevis, 284
Peckoltia cavatica, 284
Peckoltia filicaudata, 284
Peckoltia furcata, 284
Peckoltia kuhlmanni, 284
Peckoltia oligospila, 284
Peckoltia sabaji, 284
Peckoltia snethlageae, 285
Peckoltia ucayalensis, 285

Peckoltia vermiculata, 285
Peckoltia vittata, 285
Peckoltichthys, 283
Peckoltichthys filicaudatus, 283, 284
Peckoltichthys kuhlmanni, 284
pecten, Copionodon, 404
pectinata, Conta, 384
pectinatus, Ameiurus, 206
pectinatus, Ictalurus, 206
pectinatus, Tetracamphilius, 27
pectinidens, Pimelodus, 46
pectinifer, Pimelodella, 194
pectinifrons, Agamyxis, 168
pectinifrons, Doras, 168
pectinopterus, Glyptosternon, 392
pectinopterus, Glyptothorax, 392
pectorale, Lepthoplosternum, 128
pectoralis, Arius, 48
pectoralis, Callichthys, 128
pectoralis, Neoarius, 48
pediculatus, Dolichancistrus, 235
pediculatus, Pseudancistrus, 235
pediculatus cobrensis, Pseudancistrus,
235
peguensis, Hemibagrus, 90
peguensis, Macrones, 90
pellegrini, Andersonia, 24
pellegrini, Plecostomus, 289
pellopterygius, Microglanis, 354
pellucida, Parailia, 360
pellucida, Physailia, 360
Pelodichthys, 215
peloichthys, Trachelyopterus, 80
peloichthys, Trachycorystes insignis,
80
peltatus, Rhineastes, 430
Pelteobagrini, 81
Pelteobagrus, 81, 99
Pelteobagrus argentivittatus, 100
Pelteobagrus brashnikowi, 100
Pelteobagrus eupogon, 100
Pelteobagrus fulvidraco, 100
Pelteobagrus intermedius, 100
Pelteobagrus kyphus, 82
Pelteobagrus mica, 100
Pelteobagrus microps, 100
Pelteobagrus nitidus, 82
Pelteobagrus nudiceps, 100
Pelteobagrus ramentosus, 108
Pelteobagrus ussuriensis, 100
Pelteobagrus vachellii, 101
Pelteobagrus virgatus, 101
Pelteobagrus virgatus vinhensis, 101
Peltura, 26
Peltura Bovei, 26
pelusius, Mystus, 96
pelusius, Silurus, 96
pemecus, Bagrus, 53
pemon, Cetopsidium, 131

604 · Zootaxa 1418 © 2007 Magnolia Press

Penesilurus, 375
Penesilurus bokorensis, 376
Penesilurus palavanensis, 375, 377
pentamaculata, Rineloricaria, 295
pentapterus, Clarias, 146
pentlandi, Trichomycterus, 423
Pentlandii, Pimelodus, 200
peregrinus, Ictalurus, 207
perforated, Microlepidogaster, 272
perforatus, Microlepidogaster, 272
perijae, Cordylancistrus, 231
perporosus, Hypophthalmus, 334
Perrunichthys, 335
Perrunichthys perruno, 335, 336
perruno, Perrunichthys, 335, 336
persimilis, Chrysichthys, 158
personatus, Callychthys, 129
peruana, Pimelodella, 194
peruanus, Arges, 66
peruanus, Astroblepus, 66
peruanus, Chasmocranus, 183
peruanus, Duopalatinus, 332
peruense, Pimelodella, 194
peruensis, Pimelodella, 194
Perugia, 339
Perugia argentina, 335
Perugia xanthus, 335
perugiae, Anyperistius, 347
perugiae, Centromochlus, 73
peruvianus, Galeichthys, 43
pestai, Corydoras, 116
Petacara, 62
petenensis, Pimelodus, 198
petleyi, Limatulichthys, 266
petleyi, Rhineloricaria, 266
petricola, Clariallabes, 138
petricola, Synodontis, 319
petroleus, Hoplomyzon atrizona, 61
Peyeria, 428
peyeria, Ariopsis, 432
pfefferi, Synodontis, 311
phaeus, Noturus, 214
Phagorus, 139
Phagorus cataractus, 141
phaiosoma, Silurichthys, 378
phaiosoma, Silurus, 377, 378
phalacra, Brachyglanis, 181
Phalacronotini, 367
Phalacronotus, 367, 374
Phalacronotus apogon, 374
Phalacronotus bleekeri, 374
Phalacronotus micronemus, 375
Phalacronotus micruropterus, 375
Phalacronotus parvanalis, 375
phalacronotus, Silurus, 374, 375
phaneronema, Branchioica, 411
phaneronema, Paravandellia, 411
phantasia, Cetopsorhamdia, 182
phelpsi, Astroblepus, 66

FERRARIS

phelpsi, Spatuloricaria, 296, 297
Phenacorhamdia, 188
Phenacorhamdia anisura, 188
Phenacorhamdia boliviana, 188
Phenacorhamdia hoehnei, 188
Phenacorhamdia macarenensis, 188,
189
Phenacorhamdia nigrolineata, 189
Phenacorhamdia somnians, 189
Phenacorhamdia tenebrosa, 189
Phenacorhamdia unifasciata, 189
philippinus, Pseudarius, 35
phillipsi, Clarias, 145
pholeter, Astroblepus, 66
phoxocephala, Hemiloricaria, 246
phoxocephala, Loricaria, 246
Phractocephalus, 336
Phractocephalus bicolor, 336
Phractocephalus gogra, 105
Phractocephalus hemioliopterus, 336
Phractocephalus itchkeea, 386
Phractocephalus kuturnee, 105
Phractocephalus nassi, 336
Phractura, 26
Phractura ansorgii, 26
Phractura bovei, 26
Phractura brevicauda, 26
Phractura clauseni, 26
Phractura fasciata, 26
Phractura gladysae, 26
Phractura ineac, 28
Phractura intermedia, 26
Phractura lindica, 26
Phractura longicauda, 26
Phractura lukugae, 26
Phractura macrura, 27
Phractura scaphyrhynchura, 27
Phractura tenuicauda, 27
Phreatobinae, 429
Phreatobius, 429
Phreatobius cisternarum, 429, 430
phreatophila, Prietella, 215
phrixosoma, Plecostomus, 298
phrixosoma, Squaliforma, 298
phrygiatus, Arius, 32
Phyllonemus, 164
Phyllonemus brichardi, 164
Phyllonemus filinemus, 164
Phyllonemus typus, 164
physacanthus, Arius, 36
Physailia, 360
Physailia ansorgii, 360
Physailia pellucida, 360
Physailia somalensis tanensis, 360
Physailia villiersi, 360
Physopyxis, 176
Physopyxis ananas, 176
Physopyxis cristata, 176
Physopyxis lyra, 176

piauhiae, Loricaria, 269
picinguabae, Listrura, 444
picklei, Cetopsorhamdia, 182
picta, Dekeyseria, 234
picta, Megalechis, 129
pictus, Akysis, 14
pictus, Amphilius, 21
pictus, Ancistrus, 234, 265
pictus, Bagrus (Sciades), 334
pictus, Callichthys, 129
pictus, Hypostomus, 265
pictus, Leiarius, 334
pictus, Pimelodus, 338
pictus, Trichomycterus, 423
pidada, Arius, 55
pietschmanni, Allabenchelys, 138
pietschmanni, Auchenoglanis, 162
pietschmanni, Clariallabes, 138
pijpersi, Cetopsorhamdia, 186
pijpersi, Imparfinis, 186
pilosus, Neblinichthys, 273
Pimelenotus, 197
Pimelenotus Vilsoni, 197, 200
Pimeletropis, 332
Pimeletropis lateralis, 332
Pimelode livrée, 213
Pimelodela garbei, 189
Pimelodella, 189
Pimelodella altipinnis, 189
Pimelodella australis, 189
Pimelodella avanhandavae, 189
Pimelodella boliviana, 189
Pimelodella boschmai, 190
Pimelodella brasiliensis, 190, 192
Pimelodella breviceps, 190
Pimelodella buckleyi, 190
Pimelodella chagresi, 190
Pimelodella chagresi odynea, 193
Pimelodella chaparae, 190
Pimelodella cinerascens, 201
Pimelodella cochabambae, 185
Pimelodella conquetaensis, 190
Pimelodella copei, 190
Pimelodella cristata, 190
Pimelodella cruxenti, 190
Pimelodella cyanostigma, 190
Pimelodella dorseyi, 190
Pimelodella eigenmanni, 191, 193,
340
Pimelodella eigenmanniorum, 191
Pimelodella elongata, 191
Pimelodella enochi, 191
Pimelodella eutaenia, 191
Pimelodella figueroai, 191
Pimelodella garbei, 189
Pimelodella geryi, 191
Pimelodella gracilis, 191
Pimelodella griffini, 191
Pimelodella grisea, 191

CHECKLIST OF CATFISHES, RECENT AND FOSSIL

Pimelodella hartii, 191
Pimelodella hartwelli, 192
Pimelodella hasemani, 192
Pimelodella howesi, 192
Pimelodella insignis, 190
Pimelodella itapicuruensis, 192
Pimelodella kronei, 192
Pimelodella lateristriga, 192
Pimelodella laticeps, 192
Pimelodella laticeps australis, 189
Pimelodella laurenti, 192
Pimelodella linami, 192
Pimelodella macrocephala, 192
Pimelodella macturki, 192
Pimelodella martinezi, 193
Pimelodella meeki, 193
Pimelodella megalops, 193
Pimelodella megalura, 193
Pimelodella metae, 193
Pimelodella modesta, 193
Pimelodella montana, 193
Pimelodella mucosa, 193
Pimelodella nigrofasciata, 193
Pimelodella notomelas, 193
Pimelodella odynea, 193
Pimelodella ophthalmica, 193
Pimelodella pallida, 194
Pimelodella pappenheimi, 194
Pimelodella parnahybae, 194
Pimelodella parva, 194
Pimelodella pectinifer, 194
Pimelodella peruana, 194
Pimelodella peruense, 194
Pimelodella peruensis, 194
Pimelodella procera, 194
Pimelodella rambarrani, 181
Pimelodella rendahli, 196
Pimelodella reyesi, 194
Pimelodella roccae, 194
Pimelodella rudolphi, 194
Pimelodella serrata, 194
Pimelodella spelaea, 195
Pimelodella steindachneri, 195
Pimelodella taeniophora, 195
Pimelodella taenioptera, 195
Pimelodella tapatapae, 195
Pimelodella transitoria, 195
Pimelodella vittata, 195
Pimelodella wesselii, 195
Pimelodella witmeri, 195
Pimelodella yuncensis, 195
Pimelodes filamentosus, 330
Pimelodes fossor, 431
Pimelodes lateristrigus, 192
Pimelodes macropterus, 332
Pimelodidae, 329
Pimelodina, 336
Pimelodina flavipinnis, 336
Pimelodina goeldii, 332

Zootaxa 1418 © 2007 Magnolia Press · 605

Pimelodina nasus, 336
Pimelodinae, 329
Pimelodini, 329
Pimelodon, 211
Pimelodon insignarius, 211, 213
Pimelodus, 336
Pimelodus (Bagrus) maculatus, 345
Pimelodus (Pimelodella) eigenmanni,
191, 193
Pimelodus (Pimelodella) griseus, 191
Pimelodus (Pimelodella) taeniophorus,
195
Pimelodus (Pimelodina) flavipinnis,
336
Pimelodus (Pimelodus) grosskopfii,
338
Pimelodus (Pseudopimelodus)
cottoides, 354
Pimelodus (Pseudopimelodus) pulcher,
355
Pimelodus (Pseudorhamdia)
brasiliensis, 190
Pimelodus (Pseudorhamdia) Chagresi,
190
Pimelodus (Pseudorhamdia) Hartii,
191
Pimelodus (Pseudorhamdia)
nigribarbis, 335
Pimelodus (Pseudorhamdia) Wesselii,
195
Pimelodus (Rhamdia) Baronis Mülleri,
201
Pimelodus (Rhamdia) brachypterus,
198
Pimelodus (Rhamdia) Cuyabae, 201
Pimelodus (Rhamdia) Knerii, 199
Pimelodus (Rhamdia) longicauda, 185
Pimelodus (Rhamdia) Parahybae, 201
Pimelodus (Rhamdia) parvus, 180
Pimelodus (Rhamdia) Queleni cuprea,
201
Pimelodus absconditus, 336
Pimelodus aeneus, 215
Pimelodus affinis, 208
Pimelodus ailurus, 205
Pimelodus albicans, 337
Pimelodus albidus, 40, 204
Pimelodus albofasciatus, 337
Pimelodus altipinnis, 189
Pimelodus altissimus, 337
Pimelodus angius, 359
Pimelodus anisurus, 98
Pimelodus antiquus, 207
Pimelodus antoniensis, 205
Pimelodus aor, 106
Pimelodus arekaima, 334
Pimelodus argenteus, 53
Pimelodus argyrus, 209
Pimelodus arius, 34

Pimelodus asperus, 395
Pimelodus atrarius, 205
Pimelodus atrobrunneus, 337
Pimelodus auratus, 155
Pimelodus bagarius, 383
Pimelodus bahianus, 195
Pimelodus balayi, 162
Pimelodus barbancho, 339
Pimelodus barbus, 44
Pimelodus batasio, 84, 85
Pimelodus Batasius, 85, 98
Pimelodus bathyurus, 201
Pimelodus bilineatus, 107
Pimelodus biscutatus, 154
Pimelodus blochii, 336, 337
Pimelodus borneënsis, 41
Pimelodus botius, 388
Pimelodus Boucardi, 201
Pimelodus brachycephalus, 198
Pimelodus breviceps, 190
Pimelodus brevis, 337
Pimelodus buckleyi, 190
Pimelodus Bufonius, 355
Pimelodus caerulescens, 81
Pimelodus cærulescens, 81
Pimelodus Cantonensis, 431
Pimelodus carcio, 85, 98
Pimelodus Carnaticus, 383
Pimelodus catulus, 205
Pimelodus cauda-furcatus, 209
Pimelodus cavasius, 95
Pimelodus cavia, 388
Pimelodus cenia, 386
Pimelodus chandramara, 86
Pimelodus charus, 355
Pimelodus cinerascens, 200
Pimelodus clarias, 320
Pimelodus clarias coprophagus, 337
Pimelodus coerulescens, 81
Pimelodus Commersonii, 43, 44
Pimelodus confinis, 205
Pimelodus conirostris, 428
Pimelodus conta, 383, 384
Pimelodus coprophagus, 337
Pimelodus cranchii, 156
Pimelodus cristatus, 189, 190
Pimelodus ctenodus, 332
Pimelodus cupreoides, 205
Pimelodus cupreus, 215
Pimelodus cyanochloros, 392
Pimelodus cyclopum, 64
Pimelodus dekayi, 206
Pimelodus Deppei, 200
Pimelodus eigenmanni, 340
Pimelodus elongatus, 191
Pimelodus eques, 184
Pimelodus exsudans, 196
Pimelodus felinus, 205
Pimelodus felis, 206

606 · Zootaxa 1418 © 2007 Magnolia Press

Pimelodus foina, 197
Pimelodus fur, 337
Pimelodus furcatus, 208, 209
Pimelodus furcifer, 210
Pimelodus gagata, 385, 386
Pimelodus gagora, 34
Pimelodus gambensis, 54
Pimelodus garciabarrigai, 337
Pimelodus garcia-barrigai, 337
Pimelodus gaudryi, 432
Pimelodus genidens, 43, 44
Pimelodus godmanni, 200
Pimelodus gracilis, 191, 196, 210
Pimelodus graciosus, 210
Pimelodus grosskopfii, 338
Pimelodus grosskopfii navarroi, 338
Pimelodus grunniens, 336
Pimelodus guatemalensis, 200
Pimelodus guirali, 162
Pimelodus gulio, 93, 95
Pimelodus guttatus, 88, 162, 163
Pimelodus Hammondii, 210
Pimelodus hara, 395, 396
Pimelodus heraldoi, 338
Pimelodus heteropleurus, 181
Pimelodus Hilarii, 200
Pimelodus holomelas, 199
Pimelodus houghi, 210
Pimelodus hoyi, 206
Pimelodus humilis, 198
Pimelodus hypselurus, 198
Pimelodus indicus, 19
Pimelodus insigne, 213
Pimelodus insignis, 339
Pimelodus jatius, 41
Pimelodus javus, 431
Pimelodus jenynsii, 196
Pimelodus jivaro, 338
Pimelodus labrosus, 334
Pimelodus lateristrigus, 192
Pimelodus laticaudus, 198
Pimelodus laticeps, 160
Pimelodus lemniscatus, 213
Pimelodus leptus, 332
Pimelodus limosus, 215
Pimelodus lividus, 205
Pimelodus longifilis, 338
Pimelodus lupus, 209
Pimelodus lutescens, 215
Pimelodus lynx, 204
Pimelodus maculatus, 209, 336, 338,
345
Pimelodus managuensis, 199
Pimelodus mangois, 18
Pimelodus mangurus, 355
Pimelodus Manillensis, 41
Pimelodus marginatus, 206
Pimelodus marmoratus, 206
Pimelodus megalops, 210

FERRARISPimelodus melanogaster, 12, 13
Pimelodus melas, 205
Pimelodus membranaceus, 309
Pimelodus menoda, 89
Pimelodus micropterus, 200
Pimelodus microstoma, 338
Pimelodus modestus, 193
Pimelodus mong, 55
Pimelodus motaguensis, 198
Pimelodus mülleri, 199
Pimelodus multiradiatus, 334
Pimelodus murius, 358
Pimelodus musculus, 200
Pimelodus mustelinus, 184
Pimelodus mysteriosus, 338
Pimelodus namdia, 200
Pimelodus nangra, 396, 397
Pimelodus natalis, 205
Pimelodus navarroi, 338
Pimelodus nebulosus, 205, 215
Pimelodus nella, 52
Pimelodus nenga, 47
Pimelodus nicaraguensis, 199
Pimelodus nigricans, 208, 209, 210
Pimelodus nigrodigitatus, 158
Pimelodus nigrofasciatus, 193
Pimelodus notatus, 210, 339
Pimelodus occidentalis, 154
Pimelodus octocirrhus, 431
Pimelodus olivaceus, 210
Pimelodus ophthalmicus, 194
Pimelodus ornatus, 338
Pimelodus ortmanni, 338
Pimelodus pallidus var. Lateralis, 210
Pimelodus pallidus var. Leucoptera,
210
Pimelodus pallidus var. Marginata,
210
Pimelodus pangasius, 325, 327
Pimelodus paranaensis, 338
Pimelodus pati, 334
Pimelodus pectinidens, 46
Pimelodus Pentlandii, 200
Pimelodus petenensis, 198
Pimelodus pictus, 338
Pimelodus pinirampus, 339
Pimelodus pirinampu, 339
Pimelodus platanus, 335
Pimelodus platespogon, 383
Pimelodus platicirris, 339
Pimelodus platycephalus, 206
Pimelodus platychir, 21, 22
Pimelodus platypogon, 392
Pimelodus platypogonides, 392
Pimelodus pleurostigma, 13
Pimelodus polycaulus, 198
Pimelodus pullus, 205
Pimelodus puma, 205
Pimelodus punctatus, 339

Pimelodus punctulatus, 216
Pimelodus pusillus, 431
Pimelodus quelen, 200
Pimelodus rama, 105
Pimelodus raninus, 352, 353
Pimelodus rigidus, 339
Pimelodus rita, 106
Pimelodus Rogersi, 198
Pimelodus rugosus, 13
Pimelodus Sadleri, 431
Pimelodus sagor, 45
Pimelodus salvini, 198
Pimelodus sapo, 200
Pimelodus sebae, 197, 200
Pimelodus seengtee, 97
Pimelodus Sellonis, 200
Pimelodus silondia, 366
Pimelodus sona, 45
Pimelodus Spegazzinii, 335
Pimelodus spixii, 40
Pimelodus Stegelichii, 200
Pimelodus synodontis, 313
Pimelodus tachisurus, 431
Pimelodus telchita, 394
Pimelodus telchitta, 394
Pimelodus tengana, 86
Pimelodus tengara, 97
Pimelodus thunberg, 35
Pimelodus urua, 359
Pimelodus vacha, 358
Pimelodus valenciennis, 335
Pimelodus variegatus, 13, 14, 15
Pimelodus velifer, 203
Pimelodus versicolor, 44
Pimelodus viridescens, 395
Pimelodus viscosus, 215
Pimelodus vulgaris, 206
Pimelodus vulpeculus, 206
Pimelodus vulpes, 211
Pimelodus wagneri, 201
Pimelodus Westermanni, 329, 330
Pimelodus wuchereri, 200
Pimelodus Xanthocephalus, 207
Pimelodus zonatus, 13
Pimelodus zungaro, 344
Pimelotus, 197
pinheiroi, Corydoras, 122
Pinirampidae, 329
Pinirampus, 329, 339
Pinirampus pirinampu, 339
Pinirampus typus, 339
pinirampus, Pimelodus, 339
pinnatus, Ompok, 373
pinnimaculatus, Aelurichthys, 38
pinnimaculatus, Bagre, 38
Pinniwallago, 375
Pinniwallago kanpurensis, 375
piperata, Tympanopleura, 68, 70
piperatus, Ageneiosus, 70

CHECKLIST OF CATFISHES, RECENT AND FOSSIL

piperatus, Imparfinis, 185, 186
piperatus Kryptopterus, 370
pipri, Erethistoides, 384
pipri, Erethistoides montana, 384
pique, Hatcheria, 405
piracicabae, Loricaria, 269
Piramutana, 330
Piramutana macrospila, 339
Pirarara, 336
Pirarara bicolor, 336
pirareta, Ancistrus, 224
piratatu, Hypostomus, 258
Piratinga, 330
Piratinga pirá-aïba, 330
Pirauáca, Sorubim, 343
pirá-aïba, Piratinga, 330
piresi, Glanidium, 78
piresi, Tocantinsia, 78
piriformis, Ancistrus, 224
pirinampu, Pimelodus, 339
pirinampu, Pinirampus, 339
Pirinampus, 339
Pirinampus agassizii, 339
pirrense, Cyclopium, 66
pirrensis, Astroblepus, 66
piscatrix, Pseudorhamdia, 337
pitmani, Chrysichthys, 160
piurae, Pygidium punctulatum, 421
piurae, Trichomycterus, 421
placidus, Noturus, 214
planicauda, Parotocinclus, 283
planiceps, Ancistrus, 264
planiceps, Arius, 50
planiceps, Bagrus, 90
planiceps, Clarias, 147
planiceps, Hemibagrus, 90
planiceps, Notarius, 50
planiceps, Platystoma, 343
planiceps, Sorubimichthys, 343
planifrons, Genidens, 44
planifrons, Netuma, 44
Planiloricaria, 217, 285
Planiloricaria cryptodon, 285
Planiloricariina, 217
planquettei, Lithoxus, 267
planquettei, Lithoxus (Paralithoxus),
267
planus, Otolithus (Arius), 435
planus, Tachysurus, 434, 435
platana, Bergiaria, 330
platana, Bergiella, 330
platanum, Megalonema, 335
platanus, Pimelodus, 335
platespogon, Pimelodus, 383
platicirris, Pimelodus, 339
platorynchus, Farlowella, 239
platus, Amarginops, 153
Platyallabes, 152
Platyallabes tihoni, 152

Zootaxa 1418 © 2007 Magnolia Press · 607

Platycephaloides, 139
platycephalum, Megalonema, 335
platycephalum psammium,
Megalonema, 335
platycephalus, Acrochordonichthys,12, 13platycephalus, Amblyceps, 18
platycephalus, Ameiurus, 206
platycephalus, Chaetostomus, 231
platycephalus, Chrysichthys, 158
platycephalus, Clarias, 139, 147
platycephalus, Cordylancistrus, 23
platycephalus, Hemiodon, 288
platycephalus, Heterobranchus, 151
platycephalus, Miuroglanis, 409
platycephalus, Noturus, 212
platycephalus, Pimelodus, 206
platycephalus, Pseudohemiodon, 288
platychir, Amphilius, 22
platychir, Pimelodus, 21, 22
platychir cubangoensis, Amphilius, 23
Platyclarias, 152
Platyclarias machadoi, 152
Platydoras, 176
Platydoras armatulus, 176
Platydoras costatus, 176
Platyglanis, 164
Platyglanis depierrei, 164
platymetopon, Loricariichthys, 271
platynema, Brachyplatystoma, 331
Platynematichthys, 339
Platynematichthys notatus, 339
platynemum, Brachyplatystoma, 331
Platypogon, 339
Platypogon caerulorostris, 339, 340
platypogon, Arius, 54
platypogon, Glyptothorax, 392
platypogon, Kryptopterus, 371
platypogon, Micropogon, 371
platypogon, Pimelodus, 392
platypogon, Sciades, 54
platypogonides, Glyptothorax, 392
platypogonides, Pimelodus, 392
platypogonoides, Glyptothorax, 392
platyprosopos, Clariallabes, 138
platyrhynchos, Hemisorubim, 333
platyrhynchos, Platystoma, 333
platyrhynchus, Cordylancistrus, 232
platyrhynchus, Hemiancistrus, 232
platyrhynchus, Ompok, 373
Platysilurus, 340
Platysilurus barbatus, 340
Platysilurus malarmo, 340
Platysilurus mucosus, 340
Platysilurus olallae, 340
Platysomatos, 61
Platystacinae, 57
Platystacus, 57, 61
Platystacus anguillaris, 350, 351

Platystacus chaca, 135, 136
Platystacus cotylephorus, 61
Platystacus laevis, 58
Platystacus nematophorus, 61
Platystacus verrucosus, 58, 60
Platystoma, 342
Platystoma affine, 330
Platystoma artedii, 341
Platystoma corruscans, 341
Platystoma coruscans, 341
Platystoma emarginatum, 332
Platystoma juruense, 331
Platystoma Luceri, 342
Platystoma lütkeni, 344
Platystoma mucosa, 340
Platystoma orbignianus, 341
Platystoma orbignyanum, 341
Platystoma panthale, 341, 342
Platystoma parahybae, 344
Platystoma pardale, 341
Platystoma pardalis, 341, 342
Platystoma planiceps, 343
Platystoma platyrhynchos, 333
Platystoma punctifer, 342
Platystoma seenghala, 107
Platystoma spatula, 343
Platystoma sturio, 340
Platystoma tigrinum, 341
Platystoma truncatum, 341
Platystoma Vaillantii, 331
platystoma, Cteniloricaria, 233
platystoma, Loricaria, 233
Platystomatichthys, 340
Platystomatichthys sturio, 340
platystomus, Arius, 52
platystomus, Plicofollis, 52
Platytropius, 361
Platytropius longianalis, 357
Platytropius siamensis, 361
Platytropius sinensis, 358
platyura, Hemiloricaria, 246
platyura, Loricaria, 246
plazaii, Vandellia, 427
plecostomoides, Cochliodon, 258
plecostomoides, Hypostomus, 258
Plecostomus, 250
Plecostomus (Carinotus) carinotus,
234, 235
Plecostomus (Rhinelepis) microps, 281
Plecostomus aculeatus, 224
Plecostomus affinis, 251
Plecostomus ãgnã, 251
Plecostomus albopunctatus, 251
Plecostomus ancistroides, 251
Plecostomus angipinnatus, 251
Plecostomus angulicauda, 235
Plecostomus annae, 297
Plecostomus argus, 251
Plecostomus aspilogaster, 251

608 · Zootaxa 1418 © 2007 Magnolia Press

Plecostomus bicirrosus, 259
Plecostomus biseriatus, 297
Plecostomus bolivianus, 252
Plecostomus borellii, 252
Plecostomus boulengeri, 252
Plecostomus brasiliensis, 259
Plecostomus brevicauda, 252
Plecostomus brevis, 252
Plecostomus carinatus, 252
Plecostomus carvalhoi, 252
Plecostomus chaparae, 226
Plecostomus commersonii scabriceps,
260
Plecostomus commersonoides, 255
Plecostomus cordovae, 258
Plecostomus derbyi, 253
Plecostomus festae, 263
Plecostomus flagellaris, 268
Plecostomus fluviatilis, 253
Plecostomus francisci, 254
Plecostomus garmani, 254
Plecostomus gomesi, 298
Plecostomus goyazensis, 254
Plecostomus gymnorhynchus, 254
Plecostomus hemiurus, 254
Plecostomus Hermanni, 254
Plecostomus heylandi, 263
Plecostomus hondae, 250, 254
Plecostomus iheringi, 298
Plecostomus iheringii, 255, 298
Plecostomus interruptus, 255
Plecostomus jaguribensis, 255
Plecostomus johnii, 255
Plecostomus lacerta, 263
Plecostomus laplatae, 255
Plecostomus latirostris, 255
Plecostomus lexi, 255
Plecostomus lima, 256
Plecostomus lima atropinnis, 251
Plecostomus limosus, 253
Plecostomus longiradiatus, 256
Plecostomus luteomaculatus, 256
Plecostomus luteus, 256
Plecostomus Lütkeni, 251
Plecostomus macrops, 256
Plecostomus madeiræ, 225
Plecostomus margaritifer, 256
Plecostomus margaritifer butantanis,
256
Plecostomus meleagris, 256
Plecostomus microps, 273, 281
Plecostomus micropunctatus, 225
Plecostomus myersi, 257
Plecostomus niger, 257
Plecostomus nigromaculatus, 257
Plecostomus niveatus, 234
Plecostomus nudiventris, 257
Plecostomus obtusirostris, 218
Plecostomus paulinus, 258

FERRARISPlecostomus pellegrini, 289
Plecostomus phrixosoma, 298
Plecostomus plecostomus panamensis,243
Plecostomus plecostomus papariae,
258
Plecostomus popoi, 225
Plecostomus pusarum, 259
Plecostomus rachovii, 255
Plecostomus regani, 259
Plecostomus Robinii, 261
Plecostomus rondini, 259
Plecostomus rondoni, 259
Plecostomus scaphyceps, 260
Plecostomus scaplyceps, 260
Plecostomus scopularius, 298
Plecostomus seminudus, 260
Plecostomus spilosoma, 283
Plecostomus spilurus, 283
Plecostomus spiniger, 252
Plecostomus spinosissimus, 263
Plecostomus strigaticeps, 260
Plecostomus tenuicauda, 298
Plecostomus ternetzi, 261
Plecostomus tietensis, 261
Plecostomus topavae, 261
Plecostomus Unae, 261
Plecostomus unicolor, 225
Plecostomus vaillanti, 261
Plecostomus variipictus, 261
Plecostomus varimaculosus, 261
Plecostomus variostictus, 261
Plecostomus vermicularis, 262
Plecostomus villarsi, 298
Plecostomus virescens, 298
Plecostomus Wertheimeri, 285
Plecostomus winzi, 262
Plecostomus wuchereri, 262
plecostomus, Acipencer, 258
plecostomus, Hypostomus, 258
plecostomus, Plecostomus, 243
plecostomus panamensis, Plecostomus,
243
plecostomus papariae, Plecostomus,
258
Plectrochilus, 411
Plectrochilus diabolicus, 411
Plectrochilus machadoi, 411
Plectrochilus wieneri, 411
Pleurophysus, 410
Pleurophysus hydrostaticus, 410
pleurops, Arius, 39
pleurops, Synodontis, 319
pleurostigma, Pimelodus, 13
pleurotaenia, Farlowella, 237
pleurotaenia, Pangasius, 328
pleurotaenia, Pseudolais, 328
Plicofollis, 32, 51
Plicofollis argyropleuron, 51

Plicofollis crossocheilos, 52Plicofollis dussumieri, 52
Plicofollis nella, 52
Plicofollis platystomus, 52Plicofollis polystaphylodon, 52
Plicofollis tenuispinis, 52Plicofollis tonggol, 52
pliocaenicus, Silurus, 432
Pliosilurus, 430
Pliosilurus primus, 430
Plotoseus, 350
Plotoseus ikapor, 350
Plotosichthyoidei, 345
Plotosidae, 345
Plotosis, 350
Plotosius, 350
Plotosus, 345, 350
Plotosus (Clarias) hamiltonii, 140
Plotosus (Tandanus) tandanus, 351,
352
Plotosus abbreviatus, 351
Plotosus albilabris, 349
Plotosus argenteus, 345
Plotosus brevibarbus, 351
Plotosus caesius, 350
Plotosus canius, 350
Plotosus castaneoïdes, 351
Plotosus castaneus, 351
Plotosus elongatus, 346
Plotosus fisadoha, 350
Plotosus flavolineatus, 345
Plotosus horridus, 350
Plotosus laticeps, 345
Plotosus limbatus, 108, 346, 350
Plotosus lineatus, 350
Plotosus lineatus, 351, 352
Plotosus macrocephalus, 346
Plotosus macrophthalmus, 349
Plotosus malignus, 351
Plotosus marginatus, 350
Plotosus megastomus, 345, 346
Plotosus microceps, 346
Plotosus multiradiatus, 350
Plotosus nigricans, 352
Plotosus nkunga, 351
Plotosus papuensis, 351
Plotosus thunbergianus, 350
Plotosus unicolor, 350, 352
Plotosus vittatus, 350
Plotosus viviparus, 350
plumbea, Cetopsis, 133
plumbeus, Cetopsis, 132, 133
plumbeus, Trichomycterus, 421
plumbeus motatanensis,
Pseudocetopsis, 132
plumbeus orinoco, Pseudocetopsis,
132
pluriradiatus, Hemibagrus, 90
pluriradiatus, Macrones, 90

CHECKLIST OF CATFISHES, RECENT AND FOSSIL

pluriradiatus, Ompok, 373
poecilopterus, Bagrus, 93
poecilopterus, Leiocassis, 93
poecilus, Aspidoras, 109
poecilus, Microglanis, 354
poensis, Clarias, 141
poeyanus, Trichomycterus, 423
poeyi, Rhamdia, 199
Pogonopoma, 285
Pogonopoma obscurum, 285
Pogonopoma parahybae, 285
Pogonopoma wertheimeri, 285
Pogonopomoides, 285
poilanei, Pareuchiloglanis, 398, 399
pojeri, Chiloglanis, 307
polli, Chrysichthys, 159
polli, Malapterurus, 302
polli, Microsynodontis, 310
polli, Synodontis, 319
polycaulus, Pimelodus, 198
polygramma, Doras, 168
polyochrus, Parotocinclus, 283
polyodon, Chiloglanis, 307
polyodon, Synodontis, 319
polypogon, Chiloglanis, 307
polystaphylodon, Arius, 52
polystaphylodon, Plicofollis, 52
polystictus, Agenciosus, 70
polystictus, Ageneisus, 70
polystictus, Ageneiosus, 70
polystictus, Corydoras, 123
polystigma, Synodontis, 319
polyuranodon, Pangasius, 325, 327
ponderosus, Pangasius, 327
poonaensis, Glyptothorax conirostre,395
popoi, Plecostomus, 185
Porcinae, 81
Porcus, 81, 83
Porcus auratus, 155
Porcus bayad, 83
Porcus bayad macropterus, 83
Porochilus, 351
Porochilus meraukensis, 351
Porochilus obbesi, 351
Porochilus rendahli, 345
porosus, Silurus, 95
porosus, Trachycorystes, 81
porphyreus, Ageniosus, 70
pospisili, Cochliodon, 254
Potamarius, 53
Potamarius grandoculis, 53
Potamarius izabalensis, 53
Potamarius nelsoni, 53
potaroensis, Corydoras, 123
potaroensis, Myoglanis, 187
potaroënsis, Myoglanis, 187
potschi, Trichomycterus, 421
pradensis, Trichomycterus, 422

Zootaxa 1418 © 2007 Magnolia Press · 609

praecox, Pseudocetopsis, 133
praecox, Denticetopsis, 133
praeliorum, Astroblepus, 66
praelongus, Hassar, 173
praelongus, Leptodoras, 173
Prajadhipokia, 94
Prajadhipokia rex, 94
prashadi, Akysis, 14
prashadi, Glyptothorax, 393
prata, Parotocinclus, 283
prateri, Clupisoma, 357
pratti, Macrones, 102
pratti, Pseudobagrus, 102prenadilla, Brontes, 63, 66
prenadillus, Astroblepus, 66
prentissgrayi, Dinotopteroides, 139,
146
pretoriae, Chiloglanis, 307
prianomus, Rhinodoras, 176
pricei, Ictalurus, 209
pricei, Villarius, 208, 209
Prietella, 215
Prietella lundbergi, 215
Prietella phreatophila, 215
primaevus, Ameiurus, 207
primaevus, Propygidium, 433
primus, Pliosilurus, 430
prionotos, Corydoras, 130
prionotos, Scleromystax, 130
Pristiancistrus, 219
Pristiancistrus eustictus, 219, 221
pristos, Imparfinis, 186
procera, Pimelodella, 194
productus, Chiloglanis, 443
Proeutropiichthys, 361
Proeutropiichthys buchanani, 361
Proeutropiichthys goongwaree, 362
Proeutropiichthys macropthalmos,
362
Proeutropiichthys taakree, 362
Proeutropiichthys taakree burmanicus,
362
Proeutropius, 362
prolatus, Encheloclarias, 150
prolixa, Loricaria, 286
prolixa, Proloricaria, 286
prolixa lentiginosa, Loricaria, 286
Proloricaria, 286
Proloricaria lentiginosa, 286
Proloricaria prolixa, 286
promagdalena, Brachyplatystoma,331
proops, Bagrus, 54
proops, Ituglanis, 408
proops, Sciades, 54
proöps, Trichomycterus, 408
proops parahybae, Pygidium, 406, 407
Prophagorus, 139
Propimelodus, 340

Propimelodus caesius, 443
Propimelodus eigenmanni, 340
Propseudecheneis, 400
Propseudecheneis tchangi, 400, 401
Propygidium, 403, 433
Propygidium primaevus, 433
prosthistius, Amiurus, 205
proxima, Netuma, 49
proximus, Arius, 48, 49
psammatides, Aspidoras, 110
psammium, Megalonema, 335
psammium, Megalonema
platycephalum, 335
Psammphiletria, 27
Psammphiletria delicata, 27
Psammphiletria nasuta, 27
Pseudacanthicini, 217
Pseudacanthicus, 217, 286
Pseudacanthicus (Lithoxus) fimbriatus,
237
Pseudacanthicus fimbriatus, 236
Pseudacanthicus fordii, 286
Pseudacanthicus histrix, 286
Pseudacanthicus leopardus, 286
Pseudacanthicus serratus, 286
Pseudacanthicus spinosus, 286
Pseudageneiosus, 68
Pseudancistrus, 286
Pseudancistrus atratoênsis, 235
Pseudancistrus barbatus, 287
Pseudancistrus brevispinis, 287
Pseudancistrus carnegiei, 235
Pseudancistrus coquenani, 287
Pseudancistrus depressus, 287
Pseudancistrus genisetiger, 287
Pseudancistrus guentheri, 287
Pseudancistrus longispinis, 287
Pseudancistrus luderwaldti, 280
Pseudancistrus niger, 287
Pseudancistrus nigrescens, 287
Pseudancistrus orinoco, 287
Pseudancistrus papariae, 287
Pseudancistrus pediculatus, 235
Pseudancistrus pediculatus cobrensis,
235
Pseudancistrus sidereus, 288
Pseudancistrus torbesensis, 231, 232
Pseudariodes, 336
Pseudariodes pantherinus, 339
Pseudarius, 34
Pseudarius microcephalus, 35
Pseudarius philippinus, 35
Pseudauchenipterini, 68
Pseudauchenipterus, 68, 75
Pseudauchenipterus affinis, 76
Pseudauchenipterus flavescens, 76
Pseudauchenipterus guppyi, 76
Pseudauchenipterus jequitinhonhae,
76

610 · Zootaxa 1418 © 2007 Magnolia Press

Pseudauchenipterus nigrolineatus, 76
Pseudauchenipterus nodosus, 76
Pseudecheneidina, 382
Pseudecheneis, 382, 400
Pseudecheneis crassicauda, 400
Pseudecheneis eddsi, 442
Pseudecheneis immaculata, 400
Pseudecheneis immaculatus, 400
Pseudecheneis intermedius, 400
Pseudecheneis paviei, 400
Pseudecheneis serracula, 400
Pseudecheneis stenura, 442
Pseudecheneis sulcata, 400, 442
Pseudecheneis sulcatoides, 400
Pseudecheneis suppaetula, 442
Pseudecheneis sympelvica, 401
Pseudecheneis sympelvicus, 401
Pseudecheneis tchangi, 401
Pseudepapterus, 76
Pseudepapterus cucuhyensis, 76
Pseudepapterus gracilis, 76
Pseudepapterus hasemani, 76
Pseudeutropichthys, 165
Pseudeutropichthys multiradiatus, 165
Pseudeutropius, 356, 357, 362
Pseudeutropius acutirostris, 359
Pseudeutropius atherinoides, 359
Pseudeutropius atherinoides walkeri,
359
Pseudeutropius brachypopterus, 362
Pseudeutropius longimanus, 362
Pseudeutropius megalops, 362
Pseudeutropius mitchelli, 362
Pseudeutropius moolenburghae, 362
Pseudeutropius murius batarensis, 357
Pseudeutropius siamensis, 361
Pseudeutropius verbeekii, 326
Pseudexostoma, 401
Pseudexostoma brachysoma, 401
Pseudexostoma yunnanense, 401
Pseudexostoma yunnanensis
brachysoma, 401
Pseudocanthicus (Lithoxus) fimbriatus,
237
Pseudobagarius, 16
Pseudobagarius alfredi, 16
Pseudobagarius baramensis, 16
Pseudobagarius filifer, 16
Pseudobagarius fuscus, 16
Pseudobagarius hardmani, 16
Pseudobagarius inermis, 16
Pseudobagarius leucorhynchus, 16
Pseudobagarius macronemus, 16
Pseudobagarius meridionalis, 17
Pseudobagarius nitidus, 17
Pseudobagarius pseudobagarius, 17
Pseudobagarius similis, 17
Pseudobagarius sinensis, 17
pseudobagarius, Akysis, 16, 17

FERRARIS

pseudobagarius, Pseudobagarius, 17
Pseudobagrichthys, 82
Pseudobagrichthys macracanthus, 82
Pseudobagrus, 86, 100, 101
Pseudobagrus adiposalis, 101
Pseudobagrus albomarginatus, 101
Pseudobagrus analis, 101
Pseudobagrus aurantiacus, 101, 108
Pseudobagrus brachysoma, 429
Pseudobagrus brevianalis, 101
Pseudobagrus brevicaudatus, 102
Pseudobagrus changi, 103
Pseudobagrus chinensis, 101
Pseudobagrus chryseus, 429
Pseudobagrus emarginatus, 100
Pseudobagrus eupogoides, 107
Pseudobagrus eupogon, 100
Pseudobagrus fangi, 101
Pseudobagrus fui, 107
Pseudobagrus gracilis, 102
Pseudobagrus henryi, 100
Pseudobagrus ichikawai, 86
Pseudobagrus ikiensis, 102
Pseudobagrus intermedius, 100
Pseudobagrus koreanus, 102
Pseudobagrus kyphus, 82
Pseudobagrus medianalis, 102
Pseudobagrus nitidus, 82
Pseudobagrus nudiceps, 100
Pseudobagrus omeihensis, 102
Pseudobagrus ondon, 102
Pseudobagrus ornatus, 92
Pseudobagrus pratti, 102
Pseudobagrus taeniatus, 102
Pseudobagrus taiwanensis, 102
Pseudobagrus tenuis, 102
Pseudobagrus tokiensis, 103
Pseudobagrus trilineatus, 103
Pseudobagrus truncatus, 103
Pseudobagrus ussuriensis, 101
Pseudobagrus virgatus, 101
Pseudobagrus wangi, 108
Pseudobagrus wittenburgii, 108
Pseudobagrus wui, 101
Pseudocallophysus, 332
Pseudocanthicus (Lithoxus) fimbriatus,
237
Pseudocetopsis, 131
Pseudocetopsis baudoênsis, 131
Pseudocetopsis jurubidae, 132
Pseudocetopsis orientale, 130
Pseudocetopsis plumbeus
motatanensis, 132
Pseudocetopsis plumbeus orinoco, 132
Pseudocetopsis praecox, 133
Pseudodoras, 175
pseudogladiolus, Farlowella, 237
Pseudohemiodon, 288

Pseudohemiodon (Planiloricaria)
cryptodon, 285
Pseudohemiodon amazonus, 288
Pseudohemiodon apithanos, 288
Pseudohemiodon devincenzii, 288
Pseudohemiodon laminus, 288
Pseudohemiodon laticeps, 288
Pseudohemiodon platycephalus, 288
Pseudohemiodon thorectes, 288
pseudohemiurus, Hypostomus, 259
pseudohemiurus macrophthalmus,
Hypostomus, 256
Pseudohypophthalmus, 333
Pseudolaguvia, 382, 401
Pseudolaguvia ferula, 442
Pseudolaguvia foveolata, 401
Pseudolaguvia inornata, 401
Pseudolaguvia kapuri, 401
Pseudolaguvia muricata, 401
Pseudolaguvia ribeiroi, 401
Pseudolaguvia shawi, 401, 444
Pseudolaguvia tenebricosa, 402
Pseudolaguvia tuberculata, 402
Pseudolais, 328
Pseudolais micronemus, 328
Pseudolais pleurotaenia, 328
Pseudolais tetranema, 328
pseudoleiacanthus, Clarias, 147
Pseudolithoxus, 288
Pseudolithoxus anthrax, 288
Pseudolithoxus dumus, 289
Pseudolithoxus nicoi, 289
Pseudolithoxus tigris, 289
Pseudoloricaria, 289
Pseudoloricaria laeviuscula, 289
Pseudomystus, 103
Pseudomystus bomboides, 103
Pseudomystus breviceps, 103
Pseudomystus carnosus, 103
Pseudomystus flavipinnis, 103
Pseudomystus fumosus, 103
Pseudomystus fuscus, 103
Pseudomystus inornatus, 103
Pseudomystus leiacanthus, 104
Pseudomystus mahakamensis, 104
Pseudomystus moeschii, 104
Pseudomystus myersi, 104
Pseudomystus ornatus, 104
Pseudomystus robustus, 104
Pseudomystus rugosus, 104
Pseudomystus siamensis, 104
Pseudomystus sobrinus, 104
Pseudomystus stenogrammus, 104
Pseudomystus stenomus, 104
Pseudomystus vaillanti, 105
pseudonemacheir, Imparfinis, 186
pseudonieuhofii, Clarias, 147
Pseudopangasius, 325
Pseudopangasius nasutus, 327

CHECKLIST OF CATFISHES, RECENT AND FOSSIL

Pseudopimelodidae, 352
Pseudopimelodinae, 352
Pseudopimelodus, 352, 355
Pseudopimelodus acanthochirus, 353
Pseudopimelodus Agassizi, 353
Pseudopimelodus albomarginatus, 353
Pseudopimelodus apurensis, 353
Pseudopimelodus bufonius, 355
Pseudopimelodus charus, 355
Pseudopimelodus mangurus, 355
Pseudopimelodus nigricauda, 353
Pseudopimelodus parahybae, 354
Pseudopimelodus pulcher, 355
Pseudopimelodus roosevelti, 355
Pseudopimelodus schultzi, 355
Pseudopimelodus transmontanus, 353
Pseudopimelodus variolosus, 355
Pseudopimelodus villosus, 353
Pseudopimelodus villosus butcheri,352
Pseudoplatystoma, 340
Pseudoplatystoma corruscans, 341
Pseudoplatystoma fasciatum, 341
Pseudoplatystoma fasciatum
intermedium, 341
Pseudoplatystoma fasciatum nigricans,
341
Pseudoplatystoma fasciatum
reticulatum, 341
Pseudoplatystoma tigrinum, 341
Pseudorhamdia, 336
Pseudorhamdia fur, 337
Pseudorhamdia macronema, 337
Pseudorhamdia piscatrix, 337
Pseudorhamdia vittatus, 195
Pseudorinelepis, 289
Pseudorinelepis genibarbis, 289
Pseudosilurus, 371
Pseudosilurus macropthalmos, 374
pseudosilvinichthys, Trichomycterus,
422
pseudospinosus, Neosilurus, 349
Pseudostegophilus, 411
Pseudostegophilus haemomyzon, 412
Pseudostegophilus nemurus, 412
Pseudostegophilus paulensis, 410
Pseudostegophilus scarificator, 410
Pseudosynodontis, 311
Pseudotatia, 77
Pseudotatia parva, 77
Pseudotocinclus, 289
Pseudotocinclus intermedius, 290
Pseudotocinclus juquiae, 290
Pseudotocinclus parahybae, 290
Pseudotocinclus ribeiroi, 295
Pseudotocinclus tietensis, 290
Pseudotothyris, 290
Pseudotothyris janeirensis, 290
Pseudotothyris obtusa, 290

Zootaxa 1418 © 2007 Magnolia Press · 611

Psilichthys, 247
Psilichthys cameroni, 247, 279
psilogaster, Hypoptopoma, 250
Ptedoras angelis, 178
Pterobunocephalus, 62
Pterobunocephalus depressus, 62
Pterobunocephalus dolichurus, 62
Pterocryptis, 367, 375
Pterocryptis anomala, 375
Pterocryptis barakensis, 441
Pterocryptis berdmorei, 376, 377
Pterocryptis bokorensis, 376
Pterocryptis buccata, 376
Pterocryptis burmanensis, 376
Pterocryptis cochinchinensis, 376
Pterocryptis crenula, 376
Pterocryptis cucphuongensis, 376
Pterocryptis furnessi, 376
Pterocryptis gangelica, 375, 376
Pterocryptis inusitata, 377
Pterocryptis taytayensis, 377
Pterocryptis torrentis, 377
Pterocryptis verecunda, 377
Pterocryptis wynaadensis, 377
Pterodoras, 177
Pterodoras angeli, 177, 178
Pterodoras granulosus, 177
Pterodoras rivasi, 177
Pteroglanis, 343, 387
Pteroglanis horai, 387, 390
Pteroglanis manni, 343
Pteronotidae, 180
Pteronotus, 180, 197
Pteropangasius, 328
Pteropsoglanis, 387
Pterosturisoma, 290
Pterosturisoma microps, 290
Pterygophlichtys, 290
Pterygoplichthys, 217, 290
Pterygoplichthys (Ancistrus)
parananus, 272
Pterygoplichthys aculeatus, 272
Pterygoplichthys ambrosettii, 291
Pterygoplichthys disjunctivus, 291
Pterygoplichthys etentaculatus, 291
Pterygoplichthys gibbiceps, 291
Pterygoplichthys joselimaianus, 291
Pterygoplichthys juvens, 291
Pterygoplichthys lituratus, 291
Pterygoplichthys multiradiatus, 291
Pterygoplichthys pardalis, 292
Pterygoplichthys parnaibae, 292
Pterygoplichthys punctatus, 292
Pterygoplichthys scrophus, 292
Pterygoplichthys undecimalis, 292
Pterygoplichthys xinguensis, 292
Pterygoplichthys zuliaensis, 292
pubescens, Rhamdia, 201
pucai, Hoffstetterichthys, 30

puganensis, Loricaria, 297
puganensis, Spatuloricaria, 297
pulcher, Amphilius, 23
pulcher, Aneistrus (Hemiancistrus),
234
pulcher, Auchenoglanis, 162
pulcher, Corydoras, 123
pulcher, Clarias, 147
pulchra, Dekeyseria, 234
pulcher, Macrones, 97
pulcher, Mystus, 97
pulcher, Pimelodus(Pseudopimelodus), 355
pulcher, Pseudopimelodus, 355
pulcher, Synodontis, 319
pulcher ephippiata, Amphilius, 23
pulex, Ammoglanis, 403
pulicaris, Clarias, 142
pullus, Pimelodus, 205
puma, Pimelodus, 205
pumilus, Arius, 386
pumilus, Chiloglanis, 30
punctata, Ailia, 356
punctata, Ailiichthys, 356
punctata, Chaetostomus (Ancistrus)
cirrhosus, 222
punctata, Harttia, 241
punctata, Loricaria, 266, 292
punctata, Steindachneria scripta, 344
punctata, Nangra, 395
punctata, Tatia, 73
punctatissimus, Chaetostomus, 274
punctatissimus, Oligancistrus, 274
punctatissimus, Trichomycterus, 422
punctatum, Glyptosternon, 389
punctatum, Hoplosternum, 127, 128
punctatum, Hypostoma, 220
punctatum, Megalonema, 339
punctatum, Steindachneridion, 344
punctatus, Doras (Corydoras), 167
punctatus, Auchenipterichthys, 71
punctatus, Auchenipterus, 71
punctatus, Auchenoglanis, 163
punctatus, Bagrus, 91
punctatus, Cataphractus, 111, 123
punctatus, Centromochlus, 73
punctatus, Chrysichthys, 159
punctatus, Clarias, 140
punctatus, Corydoras, 123
punctatus, Doras, 167
punctatus, Hemibagrus, 91
punctatus, Henonemus, 406
punctatus, Hypostomus, 220, 259
punctatus, Ictalurus, 208, 209, 211
punctatus, Malapterurus, 302
punctatus, Pachypterus, 358
punctatus, Parauchenoglanis, 163
punctatus, Pimelodus, 339
punctatus, Pterygoplichthys, 266, 292

612 · Zootaxa 1418 © 2007 Magnolia Press

punctatus, Silurus, 209, 377, 379
punctatus, Stegophilus, 406
punctatus argentina, Corydoras, 122
punctatus sipaliwini, Corydoras, 125
puncticulatus, Arius, 41
punctifer, Mystus, 97
punctifer, Platystoma, 342
punctifer, Synodontis, 319
punctulata, Synodontis, 320
punctulatum piurae, Pygidium, 421
punctulatus, Bagrus, 331, 339, 340
punctulatus, Hemiancistrus, 244
punctulatus, Pimelodus, 216
punctulatus, Synodontis, 320
punctulatus, Trichomycterus, 422
punjabensis, Glyptothorax, 393
punjabensis, Glyptothorax conirostris,
393
purusiensis, Panaqolus, 277
purusiensis, Panaque, 277
pusarum, Hypostomus, 259
pusarum, Plecostomus, 259
Pusichthys, 362
pusillus, Erethistes, 384
pusillus, Pimelodus, 431
pusillus, Trichomycterus, 411
pussilus, Erethistes, 384
Pygidianops, 412
Pygidianops cuao, 412
Pygidianops eigenmanni, 412
Pygidianops magoi, 412
Pygidiidae, 427
Pygidium, 427
Pygidium alternatum, 414
Pygidium alterum, 415
Pygidium angustirostris, 413
Pygidium arleoi, 415
Pygidium atochae, 423
Pygidium banneaui, 415, 420
Pygidium banneaui maracaiboensis,
420
Pygidium barbouri, 415
Pygidium bogotense, 415
Pygidium bomboizanum, 416
Pygidium boylei, 416
Pygidium Burmeisteri, 405
Pygidium caliense, 416
Pygidium chapmani, 417
Pygidium chiltoni, 417
Pygidium conradi, 417
Pygidium davisi, 417
Pygidium dispar, 417
Pygidium dorsostriatum, 417
Pygidium dorsotriatum, 417
Pygidium emanueli emanueli, 418
Pygidium emanueli motatanensis, 421
Pygidium fassli, 418
Pygidium fasslii, 418
Pygidium florense, 414

FERRARIS

Pygidium fuscum, 427
Pygidium gabrieli, 418
Pygidium gracilior, 407
Pygidium guianense, 418
Pygidium guianensis, 418
Pygidium hasemani, 418
Pygidium heterodontum, 418
Pygidium iheringi, 419
Pygidium immaculatum, 419
Pygidium johnsoni, 419
Pygidium latidens, 420
Pygidium latistriatum, 420
Pygidium metae, 407
Pygidium metae guayaberensis, 407
Pygidium migrans, 420
Pygidium mondolfi, 421
Pygidium oroyae, 423
Pygidium paolence, 421
Pygidium paquequerense, 421
Pygidium parkoi, 407
Pygidium proops parahybae, 406, 407
Pygidium punctulatum piurae, 421
Pygidium quechuorum, 423
Pygidium regani, 422
Pygidium reinhardti, 422
Pygidium riojanum, 422
Pygidium romeroi, 423
Pygidium santae-ritae, 423
Pygidium Schmidti, 416
Pygidium septentrionale, 424
Pygidium Spegazzinii, 423
Pygidium spilosoma, 423
Pygidium stawiarski, 423
Pygidium stellatum, 424
Pygidium stramineum, 424
Pygidium straminium, 424
Pygidium striatum, 424
Pygidium taenia transandianum, 424
Pygidium tenue, 424
Pygidium tiraquae, 423
Pygidium totae, 412
Pygidium travassosi, 414
Pygidium triguttatum, 425
Pygidium unicolor, 425
Pygidium venulosum, 425
Pygidium vermiculatum, 425
Pygidium weyrauchi, 425
Pygidium zonatum, 425
pygmaeus, Corydoras, 123
pygmaeus, Silurus, 132
Pylodictis, 215
Pylodictis limosus, 215
Pylodictis olivaris, 215
pyrineusi, Cochliodon, 259
pyrineusi, Hypostomus, 259
Pyxiloricaria, 292
Pyxiloricaria menezesi, 292
Q
quadrensis, Rineloricaria, 295

quadricostatus, Silurus, 122
quadrifilis, Ageneiosus, 78
quadrifilis, Tetranematichthys, 78
quadrimaculatus, Silurus, 200
quadriocellatum, Glyptosternon, 393
quadriocellatus, Glyptothorax, 393
quadriradiatus, Bunocephalus, 60
quadriradiatus, Dysichthys, 60
quadriscutis, Arius, 37
quadriscutis, Aspistor, 37
quadrizonatus, Chasmocranus, 183
quechuorum, Pygidium, 423
quelen, Pimelodus, 200
quelen, Rhamdia, 199
Queleni cuprea, Pimelodus (Rhamdia),
201
Quiritixys, 240
Rrabauti, Corydoras, 123
rabdophorus, Sisor, 402
Rabida, 211
rachovii, Plecostomus, 255
radiata, Netuma, 435
radiatus, Glyptocephalus, 432, 433
radiosus, Entomocorus, 443
radulus, Rhineastes, 430
raimundi, Aspidoras, 110
raimundi, Corydoras, 110
Raja similis, 434
Rama, 105
Rama buchanani, 105
Rama chandramara, 86
Rama rama, 105rama, Pimelodus, 105rama, Rama, 105
rambarrani, Brachyrhamdia, 181
rambarrani, Pimelodella, 181
ramentosus, Bagrus (Bagrus), 108
ramentosus, Pelteobagrus, 108
ramirezi, Hoplodoras, 174
ramiroi, Ituglanis, 408
ramosus, Trichomycterus, 422
raninus, Batrochoglanis, 353
raninus, Pimelodus, 352, 353
ranunculus, Ancistrus, 224rebeli, Synodontis, 320, 323
recavus, Akysis, 14
reddelli, Rhamdia, 202
Reganella, 217, 292
Reganella depressa, 293
Reganellina, 217
regani, Anadoras, 169regani, Arges, 66
regani, Astroblepus, 66
regani, Doras, 169regani, Hypostomus, 259
regani, Hemipsilichthys, 280
regani, Leiocassis, 93regani, Pareiorhaphis, 280

CHECKLIST OF CATFISHES, RECENT AND FOSSIL

regani, Plecostomus, 259
regani, Pygidium, 422
regani, Rhamdia, 198
regani, Trichomycterus, 422
reinhardti, Bagropsis, 329
reinhardti, Pygidium, 422
reinhardti, Trichomycterus, 422
reinhardtii, Ochmacanthus, 410
Reinhardtii, Stegophilus, 410
reinii, Liobagrus, 19, 20
reisi, Ancistrus, 224
rendahli, Aoria, 108
rendahli, Copidoglanis, 345
rendahli, Neosilurus, 345
rendahli, Pimelodella, 196
rendahli, Porochilus, 345
rengifoi, Astroblepus, 66
resimus, Astephus, 207
resupinata, Synodontis, 320
resupinatus, Synodontis, 320
reticulata, Farlowella, 239
reticulata, Tatia, 73
reticulatum, Glyptosternon, 387
reticulatum, Pseudoplatystoma
fasciatum, 341
reticulatus, Ameiurus, 206
reticulatus, Bagrus, 330, 331
reticulatus, Centromochlus, 73
reticulatus, Chiloglanis, 307
reticulatus, Corydoras, 123
reticulatus, Glyptosternon, 387
retropinna, Arges, 66
retropinnatus, Horiomyzon, 185retropinnis, Bunocephalus, 59
retropinnis, Trichomycterus, 422
retropinnus, Astroblepus, 66
retuculatus, Glyptosternon, 387
revelatus, Corydoras, 123rex, Prajadhipokia, 94reyesi, Pimelodella, 194
reynoldsi, Corydoras, 123
rhabdophorus, Sisor, 402
rhabdostigma, Megalonema, 338
rhabdurus, Pareuchiloglanis, 399
Rhadinoloricaria, 293
Rhadinoloricaria macromystax, 293
rhadinurus, Ompok, 373
rhaeas, Ictalurus, 210
rhaeas, Rhineastes, 210
Rhamdella, 196
Rhamdella aymarae, 196
Rhamdella eriarcha, 196
Rhamdella exsudans, 196
Rhamdella gilli, 196
Rhamdella ignobilis, 196
Rhamdella jenynsii, 196
Rhamdella lemai, 196
Rhamdella leptosoma, 196
Rhamdella longipinnis, 196

Zootaxa 1418 © 2007 Magnolia Press · 613

Rhamdella longiuscula, 196
Rhamdella montana, 196
Rhamdella papariae, 197
Rhamdella robinsoni, 197
Rhamdella rusbyi, 197
Rhamdella schultzi, 187
Rhamdella straminea, 196
Rhamdella wolfi, 197
Rhamdia, 197
Rhamdia alfaroi, 199
Rhamdia amatitlanensis, 198
Rhamdia barbata, 201
Rhamdia branneri, 201
Rhamdia branneri voulezi, 201
Rhamdia bransfordii, 201
Rhamdia cabrerae, 198
Rhamdia cyanostigma, 190
Rhamdia depressa, 201
Rhamdia dorsalis, 201
Rhamdia duquei, 202
Rhamdia eigenmanniorum, 191
Rhamdia enfurnada, 197
Rhamdia eriarcha, 196
Rhamdia foina, 197
Rhamdia gilli, 196
Rhamdia guairensis, 198
Rhamdia guasarensis, 197
Rhamdia guatemalensis decolor, 202
Rhamdia guatemalensis muriei, 202
Rhamdia guatemalensis stygaea, 202
Rhamdia heteracantha, 201
Rhamdia holomelas rupununi, 197
Rhamdia humilis, 198
Rhamdia itacaiunas, 198
Rhamdia javanica, 431
Rhamdia jequitinhonha, 198
Rhamdia laluchensis, 198
Rhamdia laticauda, 198, 199, 202
Rhamdia laticauda typhla, 198
Rhamdia laukidi, 199
Rhamdia lehmanni, 202
Rhamdia luigiana, 199
Rhamdia macuspanensis, 199
Rhamdia marthae, 181
Rhamdia micayi, 197
Rhamdia microcephala, 202
Rhamdia microps, 201
Rhamdia minuta, 186
Rhamdia mounseyi, 201
Rhamdia muelleri, 199
Rhamdia nasuta, 201
Rhamdia nicaraguensis, 199
Rhamdia oaxacae, 201
Rhamdia obesa, 199
Rhamdia ortoni, 201
Rhamdia parryi, 199
Rhamdia poeyi, 199
Rhamdia pubescens, 201
Rhamdia quelen, 199

Rhamdia reddelli, 202
Rhamdia regani, 198
Rhamdia riojae, 201
Rhamdia sacrificii, 198
Rhamdia saijaensis, 202
Rhamdia sebae Martyi, 202
Rhamdia tenella, 199
Rhamdia underwoodi, 198
Rhamdia xetequepeque, 202
Rhamdia zongolicensis, 199
Rhamdiae, 180
Rhamdioglanis, 202
Rhamdioglanis frenatus, 202
Rhamdioglanis transfasciatus, 202
Rhamdiopsis, 202
Rhamdiopsis microcephala, 202
Rhamdiopsis moreirai, 202, 203
rhami, Crossoloricaria, 232
rhegma, Mystus, 97
Rheoglanis, 164
Rheoglanis dendrophorus, 164
rheophilus, Amphilius, 23
rheophilus, Pangasius, 328
rheophilus, Sisor, 402
Rhineastes, 207, 430
Rhineastes arcuatus, 207
Rhineastes calvus, 207
Rhineastes grangeri, 432
Rhineastes peltatus, 430
Rhineastes radulus, 430
Rhineastes rhaeas, 210
Rhineastes smithii, 430
Rhinelepis, 217, 293
Rhinelepis Agassizii, 289
Rhinelepis aspera, 293
Rhinelepis levis, 255
Rhinelepis lophophanes, 276
Rhinelepis parahybae, 285
Rhinelepis rudolphi, 281
Rhinelepis strigosa, 293
Rhineloricaria morrowi, 246
Rhineloricaria petleyi, 266
Rhineloricaria wolfei, 247
Rhinobagrus, 92
Rhinobagrus dumerili, 92, 93
Rhinodoras, 178
Rhinodoras amazonum, 170
Rhinodoras boehlkei, 178
Rhinodoras dorbignyi, 178
Rhinodoras prianomus, 176
Rhinodoras teffeanus, 176
Rhinodoras thomersoni, 178
Rhinoglanina, 303
Rhinoglanis, 303, 310
Rhinoglanis typus, 310, 311
Rhinoglanis Vannutellii, 311
Rhinolepis paraguensis, 275
Rhizosomichthys, 412
Rhizosomichthys totae, 412

614 · Zootaxa 1418 © 2007 Magnolia Press

rhodonotus, Bagrus, 49
rhodopterygius, Bagrus, 95
rhombocephala, Harttia, 242
Rhynchodoras, 178
Rhynchodoras woodsi, 178
Rhynchodoras xingui, 178ribeirae, Microcambeva, 409
ribeirensis, Neoplecostomus, 274
ribeiroi, Glanidium, 75
ribeiroi, Laguvia, 401ribeiroi, Pseudolaguvia, 401ribeiroi, Pseudotocinclus, 295
ribeiroi kapuri, Laguvia, 401
riberae, Astroblepus, 66
ribularis, Silurus, 201
ricardoae, Synodontis, 320
Ricola, 217, 293
Ricola macrops, 293
Ricolina, 217
rigidus, Pimelodus, 339
Rinelepis acanthicus, 218
Rinelepis genibarbis, 289
Rinelepis histrix, 218, 286
Rineloricaria, 217, 244, 293
Rineloricaria aequalicuspis, 293
Rineloricaria cadeae, 293
Rineloricaria castroi, 245
Rineloricaria catamarcensis, 294
Rineloricaria cubataonis, 294
Rineloricaria felipponei, 294
Rineloricaria formosa, 245
Rineloricaria hasemani, 245
Rineloricaria henselii, 294
Rineloricaria heteroptera, 293, 294
Rineloricaria jaraguensis, 294
Rineloricaria kronei, 294
Rineloricaria lanceolata, 246
Rineloricaria latirostris, 294
Rineloricaria lima, 294
Rineloricaria longicauda, 294
Rineloricaria maquinensis, 294
Rineloricaria microlepidogaster, 295
Rineloricaria microlepidota, 295
Rineloricaria misionera, 295
Rineloricaria pareiacantha, 295
Rineloricaria pentamaculata, 295
Rineloricaria quadrensis, 295
Rineloricaria steindachneri, 295
Rineloricaria strigilata, 295
Rineloricaria thrissoceps, 295
Rineloricaria uracantha, 240
Rineloricariina, 217
ringueleti, Hisonotus, 249
riojae, Rhamdia, 201
riojanum, Pygidium, 422
riojanus, Trichomycterus, 422
rios, Pangasius, 328
Rita, 81, 105
Rita Buchanani, 105, 106

FERRARISRita chrysea, 105
Rita gogra, 105
Rita grandiscutata, 105
Rita kuturnee, 105
Rita macracanthus, 106
Rita rita, 106
Rita sacerdotum, 106
rita, Pimelodus, 106
rita, Rita, 106
Ritae, 81
ritoides, Arius, 106
rivasi, Apuredoras, 177rivasi, Pterodoras, 177
rivularis, Silurus, 201
rivulatus, Trichomycterus, 422
roae, Cetopsidium, 131
robbianus, Synodontis, 320
robecchii, Clarias, 143robertsi, Ariopsis, 48
robertsi, Arius, 48
robertsi, Synodontis, 320
robineae, Corydoras, 124robinii, Hypostomus, 259
Robinii, Plecostomus, 261
robinsoni, Rhamdella, 197
robusta, Nangra, 397
robusta, Oxyloricaria, 299
robusta, Pareuchiloglanis, 399
robusta, Sundagagata, 387, 392
robustum, Megalonema, 339
robustum, Sturisoma, 299
robustus, Auchenipterus, 80
robustus, Corydoras, 124robustus, Ichthaelurus, 210
robustus, Leiocassis, 104
robustus, Neosilurus, 348
robustus, Pareuchiloglanis, 399
robustus, Pseudomystus, 104
roccae, Pimelodella, 194
rochai, Aspidoras, 108, 110
rodriguesi, Glaphyropoma, 405
rogersae, Leptodoras, 173
Rogersi, Pimelodus, 198
roigi, Trichomycterus, 423
romani, Centromochlus, 73
romani, Tatia, 73
romeroi, Pygidium, 423
romeroi, Trichomycterus, 423
roncallii, Farlowella, 240
rondoni, Ageneiosus, 69
rondini, Hypostomus, 259
rondoni, Plecostomus, 259
roosae, Clupisoma, 357
roosevelti, Pseudopimelodus, 355
roraima, Neblinichthys, 273
rosae, Chasmocranus, 183rosei, Astroblepus, 66
roseopunctatus, Hypostomus, 259
rostrata, Loricaria, 245, 298, 300

rostrata, Otothyris, 276
rostratum, Sturisoma, 300
rostratus, Arius, 55
rostratus, Loricariichthys, 271
rothschildi, Xenocara, 221
rotundatus, Otolithus (Arius), 435
rotundiceps, Gephyroglanis, 28
rotundiceps, Zaireichthys, 28
rotundifrons, Bathyclarias, 136
rotundifrons, Dinotopterus, 136
rousseauxii, Bagrus, 331rousseauxii, Brachyplatystoma, 331
royauxi, Euchilichthys, 309
royeroi, Denticetopsis, 133
ruandae, Synodontis, 320
rubermentus, Glyptothorax, 394
rudis, Diplomystes, 435
rudolphi, Pareiorhina, 281
rudolphi, Pimelodella, 194
rudolphi, Rhinelepis, 281
rueppelli, Chrysichthys, 159
rufescens, Macrones, 97
rufescens, Mystus, 94, 97
rufigiensis, Synodontis, 320
rugispinis, Arius, 32
rugosa, Farlowella, 239
rugosus, Acrochordonichthys, 13
rugosus, Bunocephalus, 60
rugosus, Liocassis, 104rugosus, Pimelodus, 13
rugosus, Pseudomystus, 104
rukwaensis, Chiloglanis, 308
rukwaensis, Synodontis, 320
rukwaensis, Synodontis zambezensis,
320
rupestre, Loricaria uracantha, 240
rupestris, Fonchiiichthys, 240
rupununi, Rhamdia holomelas, 197
rusbyi, Rhamdella, 197
Russellii, Wallago, 381
russi, Arius, 57
rutilus, Bagarius, 383
rutschi, Arius, 435
ruziziensis, Chiloglanis, 308
Ssabahensis, Pangasius, 328
sabaji, Peckoltia, 284
sabalo, Arges, 63, 67
sabalo, Astroblepus, 67
sabanus, Hemibagrus, 91
sabanus, Kryptopterus, 370
sabanus, Mystus, 91
sabanus, Ompok, 371
Sacchii, Oxyglanis, 154
Saccobranchus, 151
Saccobranchus microcephalus, 151
Saccobranchus microps, 151
sacerdotum, Rita, 106
Sachsdoras, 177

CHECKLIST OF CATFISHES, RECENT AND FOSSIL

Sachsdoras apurensis, 177, 178
sacrificii, Rhamdia, 198
Sadleri, Pimelodus, 431
saetiger, Lasiancistrus, 265sagor, Hexanematichthys, 45
sagor, Pimelodus, 45
sagoroides, Arius, 45
saharsai, Hara, 396
saijaensis, Rhamdia, 202
saisii, Glyptosternum, 393
saisii, Glyptothorax, 393
salae, Clarias, 147
salathei, Bunocephalus, 59
salgadae, Ancistrus, 300
salmacis, Scleromystax, 130
salvini, Pimelodus, 198
sanagaensis, Chiloglanis, 308
sanchesi, Corydoras, 124
sandrae, Cetopsis, 133
sandrae, Euristhmus, 442
sanghensis, Trachyglanis, 28
sanguinea, Vandellia, 427
sanguineus, Silurichthys, 378
sanitwongsei, Pangasius, 328
santaeritae, Trichomycterus, 423
santae-ritae, Pygidium, 423
santanderensis, Astroblepus, 67
santanderensis, Astroblepus cyclopus,
67
sapito, Dupouyichthys, 60
sapo, Pimelodus, 200
saramaccensis, Corydoras, 124
saramaccensis, Hypostomus 259
sarareensis, Corydoras, 124
saravacensis, Leiocassis, 93
saravacensis, Liocassis, 93
sarcodes, Cetopsis, 133
Sarcogenys, 48
Sarcoglanidinae, 403
Sarcoglanis, 403, 412
Sarcoglanis simplex, 412
sardinhai, Chiloglanis, 308
sarmientoi, Stenolicmus, 414
saropterix, Brachyspondylus, 82, 432
Satan, 216
Satan eurystomus, 216
satparanus, Arius, 56
sauli, Denticetopsis, 133, 134
sauteri, Clarias, 148savorgnani, Atopochilus, 303, 304
sawrockensis, Ameiurus, 206
sawrockensis, Ictalurus, 206
saxicola, Isbrueckerichthys, 441
scabriceps, Bunocephalus, 58, 60
scabriceps, Hypostomus, 260
scabriceps, Plecostomus commersonii,
260
scaphirhyncha, Dekeyseria, 234
scaphirhynchus, Ancistrus, 234

Zootaxa 1418 © 2007 Magnolia Press · 615

scaphyceps, Hypostomus, 260
scaphyceps, Plecostomus, 260
scaphyrhynchura, Doumea, 27
scaphyrhynchura, Phractura, 27
scaplyceps, Plecostomus, 260
scarificator, Pseudostegophilus, 410
schall, Silurus, 320
schall, Synodontis, 315, 320
Schilbe, 356, 362
Schilbe angolensis, 363
Schilbe auratus, 364
Schilbe banguelensis, 363
Schilbe bocagii, 363
Schilbe bouvieri, 364
Schilbe brevianalis, 363
Schilbe congensis, 363
Schilbe congolensis, 364
Schilbe djemeri, 363
Schilbe durinii, 363
Schilbe emini, 364
Schilbe grenfelli, 363
Schilbe Hasselquistii, 365
Schilbe intermedius, 363
Schilbe Isidori, 365
Schilbe laticeps, 364
Schilbe mandibularis, 364
Schilbe marmoratus, 364
Schilbe micropogon, 364
Schilbe moebiusii, 364
Schilbe multitaeniatus, 365
Schilbe mystus, 365
Schilbe nyongensis, 365
Schilbe palmeri, 364
Schilbe senegalensis, 364
Schilbe senegalensis fasciata, 364
Schilbe Senegallus, 364
Schilbe steindachneri, 364
Schilbe tumbanus, 365
Schilbe uranoscopus, 362, 365
Schilbe yangambianus, 365
Schilbe zairensis, 366
Schilbeichthys, 357
Schilbeidae, 356
schilbeides, Bagrus, 362, 365
schilbeides, Hemisilurus, 371
schilbeides, Kryptopterus, 371
Schilbeini, 356
Schilbeodes, 211
Schilbeodes gallowayi, 212
Schilbeodes hildebrandi, 213
Schilbeodes marginatus atrorus, 213
Schilbidae, 324, 356, 429
schilboides, Bagrus, 362
schilby, Silurus, 431
Schillee Sykesii, 362
Schizolecis, 295
Schizolecis guntheri, 295
schlegeli, Arius, 56
Schlegelii, Bagrus, 36, 95

Schmidelia graciliformis, 432
Schmidti, Callomystax, 392
Schmidti, Pygidium, 416
schneideri, Silurichthys, 378
Schomburgkii, Davalla, 68, 69
schomburgkii, Chaetostomus, 265
schomburgkii, Lasiancistrus, 265
schoutedeni, Synodontis, 321
schreineri, Hoplosternum, 128
schreitmuelleri, Farlowella, 239
schreitmülleri, Farlowella, 239
schubarti, Imparfinis, 186
schubarti, Nannorhamdia, 186
schultzei, Corydoras, 112
schultzi, Centromochlus, 73
schultzi, Leptorhamdia, 187
schultzi, Pseudopimelodus, 355
schultzi, Rhamdella, 187
schultzi, Zungaro zungaro, 355
Schultzichthys, 413
Schultzichthys bondi, 413
Schultzichthys gracilis, 413
schwartzi, Corydoras, 124
schwartzi surinamensis, Corydoras,
126
schweinfurthi, Fajumia, 428, 429
Sciadeichthys, 53
Sciadeichthys (Selenaspis) walrechti,
53
Sciadeoides, 334
Sciadeops, 53
Sciades, 53
Sciades couma, 53
Sciades dowii, 53
Sciades herzbergii, 53
Sciades marmoratus, 334
Sciades passany, 54
Sciades platypogon, 54
Sciades proops, 54
Sciades troschelii, 53, 54
sciurus, Arius, 35
sclateri, Austroglanis, 81
sclateri, Gephyroglanis, 81
Scleromystax, 112, 129
Scleromystax barbatus, 129
Scleromystax macropterus, 130
Scleromystax prionotos, 130
Scleromystax salmacis, 130
Scleronema, 413
Scleronema angustirostre, 413
Scleronema minutum, 413
Scleronema operculatum, 413
scleronema, Hemisilurus, 368
scleronemus, Ceratoglanis, 368
Scobinancistrus, 296
Scobinancistrus aureatus, 296
Scobinancistrus pariolispos, 296
scolopacina, Loricaria, 237
Scoloplacidae, 366

616 · Zootaxa 1418 © 2007 Magnolia Press

Scoloplacinae, 366
Scoloplax, 366, 367
Scoloplax dicra, 367
Scoloplax distolothrix, 367
Scoloplax dolicholophia, 367
Scoloplax empousa, 367
scolymus, Lasiancistrus, 265
scopularia, Squaliforma, 298
scopularius, Plecostomus, 298
Scorpiodoras, 178
Scorpiodoras heckelii, 178
scripta, Steindachneria, 344
scripta punctata, Steindachneria, 344
scriptum, Steindachneridion, 344
scrophus, Liposarcus, 292
scrophus, Pterygoplichthys, 292
sculpodon, Hypostomus, 260
scutatus, Aelurichthys, 38
sebae, Ageneiosus, 69
sebae, Pimelodus, 197, 200
sebae Martyi, Rhamdia, 202
secundus, Microglanis, 355
seducta, Denticetopsis, 134
seemanni, Ariopsis, 33
seemanni, Arius, 33
seenghala, Platystoma, 107
seenghala, Sperata, 107
seengtee, Mystus, 97
Seengtee, Pimelodus, 97
Selenaspis, 53
Sellonis, Pimelodus, 200
semiaquilus, Corydoras, 124
semicultratus, Sinopangasius, 325, 326
seminuda, Dasyloricaria, 234
seminuda, Loricaria filamentosa, 234
seminudus, Hypostomus, 260
seminudus, Plecostomus, 260
semiscutatus, Corydoras, 125
Senegalensis, Clarias, 140
Senegalensis, Heterobranchus, 150
senegalensis, Schilbe, 364
senegalensis fasciata, Schilbe, 364
Senegallus, Schilbe, 364
sepikensis, Lambertichthys ater, 347
septentrionale, Pygidium, 424
septentrionalis, Acrochordonichthys,13
septentrionalis, Corydoras, 124septentrionalis, Stegophilus, 414
Septobranchus, 42
Septobranchus johannae, 42
seraoi, Eutropius, 363
Serdicensis, Silurus, 432
sericeum, Chaetostoma, 230
sericeus, Chaetostomus, 230
serpentis, Synodontis, 321
serracanthus, Ameiurus, 206
serracanthus, Ictalurus, 206
serracula, Pseudecheneis, 400

FERRARIS

serralabium, Callichthys, 111
serrata, Hara, 396
serrata, Pimelodella, 194
serrata, Synodontis, 321
serratum, Amblyceps, 18
serratus, Arius, 49
serratus, Corydoras, 124
serratus, Hara, 396
serratus, Hypostomus, 286
serratus, Pseudacanthicus, 286
serratus, Synodontis, 311, 321
serratus tanganaicae, Synodontis, 322
setifera, Loricaria, 268
setiger, Oreoglanis, 398
setosus, Chaetostomus, 235
setosus, Dolichancistrus, 235
seussi, Corydoras, 124
sexcirrhis, Aspredo, 61
sexpapilostoma, Hoplomyzon, 61
sextentaculatus, Heterobranchus, 197,
200
sexualis, Gagata, 386
sharavatiensis, Batasio, 86
sharpii, Chrysichthys, 159
shawi, Laguvia, 395, 402
shawi, Pseudolaguvia, 401, 444
shermani, Cetopsorhamdia, 182
shirensis, Malapterurus, 302
shirui, Hoplosternum, 128
shuwaiensis, Bagrus, 84
sialis, Noturus, 213
siamensis, Glyptothorax, 393
siamensis, Hemipimelodus, 41
siamensis, Liocassis, 104
siamensis, Oreoglanis, 397, 398
siamensis, Pangasius, 327
siamensis, Platytropius, 361
siamensis, Pseudeutropius, 361
siamensis, Pseudomystus, 104sianenna, Chrysichthys, 159
sichuanensis, Pareuchiloglanis, 399
sicuephorus, Aspredo, 58
sicula, Erethistoides, 384
sicyephorus, Aspredo, 58
Sieboldii, Bagrus, 90
sifontesi, Oxydoras, 176
Silondia, 366
silondia, Silonia, 366
Silonia, 366
Silonia childreni, 366
Silonia silondia, 366
Silonopangasius, 366
Silundia, 366
Silundia gangetica, 366
Silundia Sykesii, 366
Siluranodon, 356, 366
Siluranodon auritus, 366
Siluranodontinae, 356
Silurichthys, 377

Silurichthys australis, 348
Silurichthys basilewskii, 381
Silurichthys berdmorei, 376
Silurichthys citatus, 377
Silurichthys gibbiceps, 377
Silurichthys hasseltii, 377
Silurichthys indragiriensis, 378
Silurichthys leucopodus, 378
Silurichthys marmoratus, 378
Silurichthys phaiosoma, 378
Silurichthys sanguineus, 378
Silurichthys schneideri, 378
Siluridae, 367
Silurodes, 371
Silurodes macronema, 372
Silurodon, 380
Silurodon hexanema, 380, 381
siluroides, Ompok, 371
Siluroïdes, 367
siluroïdes, Ompok, 373
Silurus, 375, 378
Silurus (?)gaudryi, 432
Silurus (Acanthonotus) cuvieri, 356
Silurus (Callichrus) affinis, 374
Silurus (Callichrus) erythrogaster, 373
Silurus (Callichrus) immaculatus, 374
Silurus (Callichrus) macrostomus, 381
Silurus (Callichrus) nebulosus, 374
Silurus (Callichrus) vittatus, 372
Silurus (Clupisoma) argentata, 357
Silurus (Heterobranchus) gariepinus,
143
Silurus (Parasilurus) aristotelis, 378
Silurus (Pimelodus) nigrescens, 210
Silurus 11-radiatus, 335
Silurus 12-radiatus, 177
Silurus 16-radiatus, 44
Silurus 38-radiatus, 70
Silurus 7-radiatus, 122
Silurus 9-radiatus, 201
Silurus afghana, 376
Silurus altus, 381
Silurus anguillaris, 138, 140, 143
Silurus anostomus, 374
Silurus apogon, 374
Silurus arab, 351
Silurus argentinus, 209
Silurus aristotelis, 378
Silurus armatus, 177
Silurus ascita, 54
Silurus asotus, 378, 381
Silurus aspredo, 58
Silurus Atherinoides, 359
Silurus athu, 380
Silurus attu, 380
Silurus auritus, 366
Silurus bagre, 37
Silurus bajad, 83
Silurus batrachus, 140

CHECKLIST OF CATFISHES, RECENT AND FOSSIL

Silurus bedfordi, 379
Silurus bicirrhis, 369
Silurus bimaculatus, 372
Silurus biserratus, 151
Silurus biwaensis, 379
Silurus boalis, 380
Silurus burmanensis, 376
Silurus callarias, 313
Silurus callichthys, 110
Silurus calvarius, 99, 100
Silurus canio, 373, 374
Silurus carinatus, 170, 171
Silurus cataphractus, 167
Silurus catus, 204
Silurus cæcutiens, 131
Silurus Cerulescens, 207, 208, 211
Silurus cerulescens melanurus, 209
Silurus cerulescens var. melanurus,
209
Silurus chantrei, 379
Silurus chechra, 374
Silurus chilensis, 165
Silurus cinereus, 379
Silurus clarias, 311, 313, 321, 336, 337
Silurus Cochinchinensis, 376
Silurus coecutiens, 131, 132
Silurus congensis, 362, 363
Silurus costatus, 176
Silurus cous, 31, 387, 389
Silurus cryptopterus, 369
Silurus cucphuongensis, 376
Silurus cupreus, 215
Silurus dauuricus, 379
Silurus docmak, 84
Silurus duanensis, 379
Silurus duda, 374
Silurus dukai, 377
Silurus Egertoni, 56
Silurus electricus, 301
Silurus fasciatus, 340, 341
Silurus felis, 33
Silurus ferox, 431
Silurus fossilis, 151
Silurus galeatus, 79
Silurus garua, 357
Silurus gilberti, 375, 376
Silurus glanis, 378, 379
Silurus glanis aralensis, 379
Silurus glanis atavus, 382
Silurus goae, 374
Silurus grahami, 379
Silurus gurgu, 431
Silurus gyrinus, 211, 213, 214
Silurus hemioliopterus, 336
Silurus Herzbergii, 53
Silurus hexadactylus, 61
Silurus hexapterus, 371
Silurus hypophthalmus, 372
Silurus ichneumon, 431

Zootaxa 1418 © 2007 Magnolia Press · 617

Silurus imberbis, 70, 379
Silurus indicus, 374
Silurus inermis, 69, 379
Silurus japonicus, 378, 379
Silurus laïs, 370
Silurus lamghur, 373
Silurus lanzhouensis, 380
Silurus laticeps, 151
Silurus leptonema, 374
Silurus lima, 342
Silurus limosus, 215
Silurus limpok, 370
Silurus lineatus, 350, 351
Silurus lithophilus, 380
Silurus lividus, 204, 205
Silurus lividus Fuscatus, 205
Silurus lividus var. Fuscatus, 205
Silurus luvur, 431
Silurus macrocephalus, 341
Silurus macronema, 371, 372
Silurus maculatus, 35, 207, 209
Silurus maculatus Erythroptera, 209
Silurus maculatus var. Erythroptera,
209
Silurus Malabaricus, 372
Silurus mangrullo, 341
Silurus marinus, 37, 38, 44
Silurus melas, 205
Silurus mento, 380
Silurus meridionalis, 380
Silurus microcephalus, 374
Silurus microdorsalis, 380
Silurus micronemus, 375
Silurus micropogon, 374
Silurus militaris, 51, 68, 69
Silurus minutus, 431
Silurus mollis, 214
Silurus mononema, 370
Silurus morehensis, 376
Silurus Mülleri, 380, 381
Silurus muticus, 337
Silurus Mysoricus, 374
Silurus mystus, 362, 365
Silurus nebulosus, 215
Silurus nodosus, 75, 76
Silurus ocellatus, 35
Silurus olivaris, 215
Silurus pabda, 371, 372
Silurus pabo, 373
Silurus palembangensis, 369, 370
Silurus pallidus, 210
Silurus pallidus Lateralis, 210
Silurus pallidus Leucoptera, 210
Silurus pallidus Marginatus, 210
Silurus pallidus var. Lateralis, 210
Silurus pallidus var. Leucoptera, 210
Silurus pallidus var. Marginatus, 210
Silurus parkeri, 36
Silurus pati, 335

Silurus pelusius, 96
Silurus phaiosoma, 377, 378
Silurus phalacronotus, 374, 375
Silurus pliocaenicus, 432
Silurus porosus, 95
Silurus punctatus, 209, 377, 379
Silurus pygmaeus, 132
Silurus quadricostatus, 122
Silurus quadrimaculatus, 200
Silurus ribularis, 201
Silurus rivularis, 201
Silurus schall, 320
Silurus schilbe niloticus, 431
Silurus schilby, 431
Silurus Serdicensis, 432
Silurus silurus, 379
Silurus sinensis, 375, 376, 381
Silurus singio, 151
Silurus soldatovi, 380
Silurus soldatovi meridionalis, 380
Silurus stenocephalus, 432
Silurus torrentis, 377
Silurus triostegus, 380
Silurus undecimalis, 431
Silurus unitius, 350
Silurus Vaillantii, 331
Silurus verrucosus, 60
Silurus viscosus, 215
Silurus vittatus, 98
Silurus wallagoo, 381
Silurus wynaadensis, 375, 377
Silurus Xanthocephalus, 207
Silurus xanthosteus, 379
silurus, Silurus, 379
Silvaichthys, 76
Silvaichthys aguilerae, 76
silvasantosi, Steindachneridion, 344
silviae, Glyptothorax, 393
Silvinichthys, 413
Silvinichthys bortayro, 413
Silvinichthys mendozensis, 413
simeonsi, Clariallabes, 138
similis, Akysis, 17
similis, Corydoras, 124
similis, Leiocassis, 108
similis, Pseudobagarius, 17
similis, Raja, 434
simillima, Loricaria, 269
simios, Hypostomus, 260
simonsi, Galeichthys, 33
simonsii, Arges, 67
simonsii, Astroblepus, 67
simplex, Sarcoglanis, 412
simplex, Tatia, 78
simpsonii, Ictalurus, 210
simulatus, Corydoras, 124
Simuldentinae, 303
Sindensis, Callichrous, 373
sindensis, Ompok, 373

618 · Zootaxa 1418 © 2007 Magnolia Presssinesis, Akysis, 17sinense, Clupisoma, 358
sinense, Glyptosternum, 393
sinensis, Akysis, 17sinensis, Arius, 55
sinensis, Bagrus, 108
sinensis, Cranoglanis, 164, 165
sinensis, Euchiloglanis, 399
sinensis, Glyptothorax, 393
sinensis, Macrones, 165
sinensis, Pareuchiloglanis, 399
sinensis, Platytropius, 358
sinensis, Pseudobagarius, 17sinensis, Silurus, 375, 376, 381
sinensis, Tachysurus, 431
singaringan, Bagrus, 97
singaringan, Mystus, 97
Sinopangasius, 325
Sinopangasius semicultratus, 325, 326
sinyanensis, Leiocassis, 108
sipaliwini, Bunocephalus amaurus, 58
sipaliwini, Corydoras, 125
sipaliwini, Corydoras punctatus, 125
sipaliwini, Hypostomus, 260
Sisor, 382, 402
Sisor barakensis, 402
Sisor chennuah, 402
Sisor rabdophorus, 402
Sisor rhabdophorus, 402
Sisor rheophilus, 402
Sisor tortosus, 402
Sisorichthyoidei, 382
Sisoridae, 382
Slatinia, 24
Slatinia mongallensis, 24smithi, Farlowella, 239
smithii, Clarias, 143smithii, Rhineastes, 430
smithii, Synodontis, 321
smiti, Synodontis, 321
sneiderni, Hemiloricaria, 246
sneiderni, Loricaria, 246
snethlageae, Ancistrus, 283, 285
snethlageae, Peckoltia, 285
sobrinus, Pseudomystus, 104
Socnopaea, 430
Socnopaea grandis, 430
Socnopaea horai, 431
sodalis, Corydoras, 125
soldatovi, Silurus, 380
soldatovi meridionalis, Silurus, 380
solidus, Arius, 39
solidus, Brustiarius, 39soloni, Synodontis, 321
solox, Corydoras, 125
somalensis, Ailia, 360
somalensis, Parailia, 360
somalensis tanensis, Physailia, 360
somereni, Chiloglanis, 308

FERRARIS

somnians, Heptapterus, 189
somnians, Phenacorhamdia, 189
sona, Hemiarius, 45
sona, Pimelodus, 45
sondaicus, Bagrus, 45
songdaensis, Pareuchiloglanis, 400
songmaensis, Pareuchiloglanis, 400
soniae, Hypostomus, 260
Sophiancistrus, 283
sorex, Synodontis, 321
Sorubim, 281, 342
Sorubim caparary, 341
Sorubim cuspicaudus, 342
Sorubim elongatus, 342
Sorubim infraoculare, 342
Sorubim jandia, 342, 343
Sorubim latirostris, 342
Sorubim lima, 342
Sorubim maniradii, 342
Sorubim Pirauáca, 343
Sorubim trigonocephalus, 342
Sorubimichthys, 342
Sorubimichthys ortoni, 343
Sorubimichthys planiceps, 343
Sorubinae, 329
Sorubium, 329, 342
Sosia, 12
Sosia chamaeleon, 12, 13
Sosia chamaeleon pallida, 13
Sosia chamaeleon var. pallida, 13
Sovichthys, 332
Sovichthys abuelo, 332
sovichthys, Chaetostoma, 230
sovichthys, Chaetostoma anomala, 230
spatula, Arius, 42, 43
spatula, Cochlefelis, 43
spatula, Platystoma, 343
Spatuloricaria, 296
Spatuloricaria atratoensis, 296
Spatuloricaria caquetae, 296
Spatuloricaria curvispina, 296
Spatuloricaria euacanthagenys, 296
Spatuloricaria evansii, 296
Spatuloricaria fimbriata, 297
Spatuloricaria gymnogaster, 297
Spatuloricaria lagoichthys, 297
Spatuloricaria nudiventris, 297
Spatuloricaria phelpsi, 296, 297
Spatuloricaria puganensis, 297
spectabile, Nannoptopoma, 272
spectabilis, Corydoras. 125
spectabilis, Otocinclus, 272, 273
Spectracanthicina, 217
Spectracanthicus, 217, 297
Spectracanthicus murinus, 297
spectrum, Aspredo, 61
spectrum, Glyptothorax, 393
Spegazzinii, Pimelodus, 335
Spegazzinii, Pygidium, 423

spegazzinii, Trichomycterus, 423
spelaea, Pimelodella, 195
spelaeus, Trichomycterus, 423
Sperata, 106
Sperata acicularis, 106
Sperata aor, 106
Sperata aorella, 107
Sperata sarwari, 107
Sperata seenghala, 107
spilomma, Hemiancistrus, 244
spilopterus, Hemibagrus, 91
spilosoma, Parotocinclus, 283
spilosoma, Plecostomus, 283
spilosoma, Pygidium, 423
spilosoma, Trichomycterus, 423
spilotus, Aspidoras, 110
spilurus, Batasio, 442
spilurus, Corydoras, 125
spilurus, Parotocinclus, 283
spilurus, Plecostomus, 283
spiniger, Plecostomus, 252spinipectoralis, Mystus, 97
spiniserrata, Parailia, 360
spinosae, Loricaria, 286
spinosissimus, Acanthodoras, 168
spinosissimus, Doras, 168
spinosissimus, Hemiancistrus, 244
spinosissima, Isorineloricaria, 263
spinosissimus, Plecostomus, 263
spinosus, Ancistrus, 224
spinosus, Hypostomus, 286
spinosus, Pseudacanthicus, 286
spixii, Cathorops, 40
Spixii, Cetopsis, 131
Spixii, Hypophthalmus, 333
Spixii, Loricaria, 270
spixii, Pimelodus, 40
splendens, Brochis, 125
splendens, Callichthys, 125
splendens, Corydoras, 125
splendens, Hemipsilichthys, 280
splendens, Pareiorhaphis, 280
spodius, Ictalurus, 211
spurrellii, Imparfinis, 186
spurrellii, Nannorhamdia, 185, 186
Squaliforma, 297
Squaliforma annae, 297
Squaliforma biseriata, 297
Squaliforma emarginata, 297
Squaliforma gomesi, 298
Squaliforma horrida, 298
Squaliforma phrixosoma, 298
Squaliforma scopularia, 298
Squaliforma squalina, 298
Squaliforma tenuicauda, 298
Squaliforma tenuis, 298
Squaliforma villarsi, 298
Squaliforma virescens, 298
squalina, Squaliforma, 298

CHECKLIST OF CATFISHES, RECENT AND FOSSIL

squalinum, Hypostoma, 298
squalus, Arius, 166
stanauli, Noturus, 214, 442
stanneus, Galeichthys, 49
stannii, Chaetostoma, 230
Stannii, Chaetostomus, 230
stappersii, Chrysichthys, 159
stappersii, Clarias, 147
starnesi, Cetopsis, 133
stauroforus, Arius, 32
stauroforus, Felichthys, 32
Stauroglanis, 413
Stauroglanis gouldingi, 413
stawiarski, Pygidium, 423
stawiarski, Trichomycterus, 423
Stearopterus, 37
Stearopterus bagre, 37
steerei, Taenionema, 330, 331
Stegelichii, Pimelodus, 200
Stegophilina, 403
Stegophilus, 403, 414
Stegophilus insidiosus, 414
Stegophilus intermedius, 405
Stegophilus macrops, 406
Stegophilus maculatus, 410
Stegophilus nemurus, 412
Stegophilus panzeri, 414
Stegophilus passarellii, 406
Stegophilus punctatus, 406
Stegophilus Reinhardtii, 410
Stegophilus septentrionalis, 414
steinbachi, Ixinandria, 263
Steinbachi, Loricaria, 263
steindachneri, Cathorops, 40, 48
Steindachneri, Centromochlus, 73
steindachneri, Corydoras, 125
steindachneri, Glyptosternum, 393
steindachneri, Glyptothorax, 393
steindachneri, Hemipsilichthys, 280
steindachneri, Hypoptopoma, 250
steindachneri, Loricaria, 295
Steindachneri, Oxydoras, 179
steindachneri, Opsodoras, 169
steindachneri, Pareiorhaphis, 280
steindachneri, Parotocinclus, 283
steindachneri, Pimelodella, 195
steindachneri, Rineloricaria, 295
steindachneri, Schilbe, 364
steindachneri, Synodontis, 321
steindachneri, Tachysurus, 40
steindachneri, Trachydoras, 169
Steindachneria, 343
Steindachneria amblyurus, 343
Steindachneria doceana, 343
Steindachneria scripta, 344
Steindachneria scripta punctata, 344
Steindachneridion, 343
Steindachneridion amblyurum, 343
Steindachneridion doceana, 343

Zootaxa 1418 © 2007 Magnolia Press · 619

Steindachneridion iheringi, 343
Steindachneridion melanodermatum,
343
Steindachneridion parahybae, 344
Steindachneridion punctatum, 344
Steindachneridion scriptum, 344
Steindachneridion silvasantosi, 344
stellatum, Lepthoplosternum, 128
stellatum, Pygidium, 424
stellatus, Nanobagrus, 98
stellatus, Trichomycterus, 424
stenocephalus, Corydoras, 125
stenocephalus, Osteogeniosus, 51
stenocephalus, Silurus, 432
stenogrammus, Pseudomystus, 104
Stenolicmus, 414
Stenolicmus sarmientoi, 414
stenomus, Bagrus, 103, 104
stenomus, Pseudomystus, 104
stenopeltis, Doras (Oxydoras), 171,172
stenopeltis, Hemidoras, 172
stenura, Pseudecheneis, 442
stephanus, Pareiorhaphis, 280
sterbai, Corydoras, 125
sternoptychum, Nannoptopoma, 273
stevardii, Hoplosternum, 128
stewarti, Hemiloricaria, 246
stewarti, Heptapterus, 184
stewarti, Loricaria, 247
sthenocephalus, Osteogeniosus, 51
stiassnyae, Malapterurus, 301
stibaros, Lamontichthys, 264
stictonotus, Imparfinis, 186
stictonotus, Nannorhamdia, 186
stigmaticus, Ancistrus, 224
stigmaturus, Mystus, 85
stigmosus, Noturus, 214
stirlingi, Arius, 46
stocki, Glyptothorax, 393
stocki, Lithoxus, 268
Stoličkae, Glyptosternum, 393
stolickae, Glyptothorax, 393
stoliczkae, Exostoma, 387
stomias, Hemipsilichthys, 281
stomias, Pareiorhaphis, 280
Stoneiella, 286
Stoneiella leopardus, 286
Stormi, Hemiarius, 45
Stormii, Cephalocassis, 44, 45
stormii, Hemiarius, 45
straminea, Rhamdella, 196
stramineum, Pygidium, 424
straminium, Pygidium, 424
straminius, Trichomycterus, 424
Strephon, 63
striatulus, Auchenipterus(Pseudauchenipterus), 80
striatulus, Trachelyopterus, 80

striatum, Pygidium, 424
Striatus, Glyptosternon, 387, 394
striatus, Glyptothorax, 394
striatus, Trichomycterus, 424
stricticassis, Arius, 50
strigata, Tatia, 78
strigaticeps, Hypostomus, 260
strigaticeps, Plecostomus, 260
strigilata, Loricaria, 295
strigilata, Rineloricaria, 295
strigosa, Rhinelepis, 293
strigosus, Acrochordonichthys, 13
stromeri, Fajumia, 429
stroudi, Gelanoglanis, 74
stroumpoulos, Chaetostoma, 444
stuarti, Glyptosternum, 385
stübeli, Arges, 67
Stübelii, Loricaria, 271
Stübelii, Oxydoras, 175
stuebeli, Astroblepus, 67
stuebelii, Loricariichthys, 271
stuebelii, Opsodoras, 175
sturio, Platystoma, 340
sturio, Platystomatichthys, 340
Sturiosoma nigrirostrum, 299
Sturisoma, 298
Sturisoma aureum, 298
Sturisoma barbatum, 299
Sturisoma brevirostre, 299
Sturisoma dariense, 299
Sturisoma festivum, 299
Sturisoma frenatum, 299
Sturisoma guentheri, 299
Sturisoma kneri, 299
Sturisoma lyra, 299
Sturisoma monopelte, 299
Sturisoma nigrirostrum, 299
Sturisoma panamense, 299
Sturisoma robustum, 299
Sturisoma rostratum, 300
Sturisoma tenuirostre, 300
Sturisomatichthys, 300
Sturisomatichthys caquetae, 300
Sturisomatichthys citurensis, 300
Sturisomatichthys leightoni, 300
Sturisomatichthys tamanae, 300
styani, Liobagrus, 20
stygaea, Rhamdia guatemalensis, 202
Stygogenes, 63
Stygogenes guentheri, 64
Stygogenes humboldtii, 64
Suarezi, Incaichthys, 30
subcarinatus, Hypostomus, 260
submarginatus, Clarias, 147
submarginatus, Loricaria, 246
submarginatus thysvillensis, Clarias(Allabenchelys), 141
subrostratus, Arius, 35
subteres, Kronichthys, 263

620 · Zootaxa 1418 © 2007 Magnolia Press

subviridis, Hemiancistrus, 244
suchus, Bagarius, 383
sufii, Glyptothorax, 394
sufii, Glyptothorax telchitta, 394
sugubii, Liobagrus, 20
sulcata, Pseudecheneis, 400
sulcatoides, Pseudecheneis, 400
sulcatus, Callichthys, 129
sulcatus, Clarias, 147
sulcatus, Glyptosternon, 400
sumatranus, Arius, 36
sumatranus, Bagrus, 36
Sundagagata, 387
Sundagagata robusta, 387, 392
sundaicus, Hexanematichthys, 45
sundanensis, Hemipimelodus, 56
Superglyptothorax, 388
suppaetula, Pseudecheneis, 442
supramollis, Astroblepus, 67
surinamensis, Chasmocranus, 183
surinamensis, Corydoras, 126
surinamensis, Corydoras schwartzi,
126
surinamensis, Harttia, 242
surinamensis, Heptapterus, 183
surinamensis, Hexanematichthys, 33
surinamensis, Hoplosternum
thoracatum, 129
surinamensis, Hypostomus, 260
surinamensis, Lithoxus, 268
surinamensis, Lithoxus (Paralithoxus),
268
sutchi, Pangasius, 325
suttoni, Panaque, 278
suttonorum, Panaque, 278
swierstrai, Chiloglanis, 308
sychri, Corydoras, 126
sykesi, Glyptosternum, 394
sykesi, Glyptothorax, 394
sykesii, Gogrius, 105
Sykesii, Schillee, 362
Sykesii, Silundia, 366
sympelvica, Pseudecheneis, 401
sympelvicus, Pseudecheneis, 401
sympterygium, Heptapterus, 184
Synechoglanis, 207
Synechoglanis Beadlei, 207, 210
synodon, Arius, 166
Synodontes, 311
Synodontini, 303
Synodontis, 303, 311
Synodontis acanthomias, 311
Synodontis acanthoperca, 441
Synodontis afrofischeri, 311
Synodontis Afro-fischeri, 311
Synodontis alberti, 311
Synodontis albolineata, 311
Synodontis albolineatus, 311
Synodontis angelica, 311

FERRARIS

Synodontis angelicus, 311
Synodontis angelicus var. zonatus, 311
Synodontis angelicus zonatus, 311
Synodontis annectens, 312
Synodontis ansorgii, 312
Synodontis arabi, 321
Synodontis arnoulti, 312
Synodontis aterrima, 312
Synodontis aterrimus, 312
Synodontis augierasi, 314
Synodontis bastiani, 312
Synodontis batensoda, 304
Synodontis batesi longispinis, 316
Synodontis batesi var. longispinis, 316
Synodontis batesii, 312
Synodontis brichardi, 312
Synodontis budgetti, 312
Synodontis camelopardalis, 312
Synodontis caudalis, 312
Synodontis caudovittata, 312
Synodontis caudovittatus, 313
Synodontis centralis, 313
Synodontis citernii, 314
Synodontis clarias, 313, 320
Synodontis colyeri, 318
Synodontis comoensis, 313
Synodontis congica, 313
Synodontis congicus, 313
Synodontis contracta, 313
Synodontis contractus, 313
Synodontis courteti, 313
Synodontis cuangoana, 313
Synodontis cuangoanus, 313
Synodontis dageti, 312
Synodontis davidi, 313
Synodontis decora, 314
Synodontis decorus, 314
Synodontis dekimpei, 314
Synodontis depauwi, 314
Synodontis dhonti, 314
Synodontis dorsomaculata, 314
Synodontis dorsomaculatus, 314
Synodontis eburneensis, 312
Synodontis euptera, 314
Synodontis eupterus, 314
Synodontis eurystomus, 305, 319
Synodontis fascipinna, 314
Synodontis filamentosa, 314
Synodontis filamentosus, 314
Synodontis flavitaeniata, 314
Synodontis flavitaeniatus, 314
Synodontis frontosa, 314
Synodontis frontosus, 314
Synodontis fuelleborni, 315
Synodontis galinae, 323
Synodontis gambiensis, 315
Synodontis gambiensis latifrons, 315
Synodontis geledensis, 315
Synodontis gobroni, 315

Synodontis grandiops, 445
Synodontis granulosa, 315
Synodontis granulosus, 315
Synodontis greshoffi, 315
Synodontis guentheri, 309
Synodontis guttata, 315
Synodontis guttatus, 315
Synodontis haugi, 315
Synodontis Hollyi, 323
Synodontis Hollyi ntemensis, 322
Synodontis holopercnus, 315
Synodontis humeratus, 321
Synodontis ilebrevis, 445
Synodontis irsacae, 314
Synodontis ituriensis, 315
Synodontis iturii, 315
Synodontis jallae, 316
Synodontis katangae, 315
Synodontis khartoumensis, 316
Synodontis koensis, 316
Synodontis labeo, 323
Synodontis lacustricolus, 322
Synodontis laessoei, 316
Synodontis leopardina, 316
Synodontis leopardinus, 316
Synodontis leoparda, 316
Synodontis leopardus, 316
Synodontis levequei, 316
Synodontis longirostris, 316
Synodontis longispinis, 316
Synodontis loppei, 318
Synodontis lucipinnis, 445
Synodontis lufirae, 316
Synodontis macrepipterus, 314
Synodontis macrodon, 313
Synodontis macrophthalma, 316
Synodontis macrophthalmus, 316
Synodontis macrops, 316
Synodontis macrostigma, 317
Synodontis macrostoma, 317
Synodontis maculatus, 318
Synodontis maculipinna, 320
Synodontis maculosus, 311, 318, 321
Synodontis manni, 317
Synodontis marmorata, 317
Synodontis marmoratus, 317
Synodontis marmoratus truncatus, 317
Synodontis matthesi, 317
Synodontis melanoptera, 317
Synodontis melanopterus, 317
Synodontis melanostictus, 318
Synodontis melanostictus iturii, 315
Synodontis melanostictus var. iturii,
315
Synodontis multimaculata, 317
Synodontis multimaculatus, 317
Synodontis multipunctata, 317
Synodontis multipunctatus, 317
Synodontis nebulosa, 317

CHECKLIST OF CATFISHES, RECENT AND FOSSIL

Synodontis nebulosus, 317
Synodontis nigrita, 317
Synodontis nigriventris, 318
Synodontis nigromaculata, 318
Synodontis nigromaculatus, 318
Synodontis njassae, 318
Synodontis notata, 318
Synodontis notatus, 318
Synodontis notatus binotata, 318
Synodontis notatus ocellatus, 318
Synodontis notatus var. binotata, 318
Synodontis notatus var. ocellatus, 318
Synodontis nummifer, 318
Synodontis obesus, 318, 323
Synodontis ocellifer, 318
Synodontis omias, 319
Synodontis ornatipinnis, 319
Synodontis ornatissima, 319
Synodontis ornatissimus, 319
Synodontis ornatus, 317, 319
Synodontis ovidius, 316
Synodontis pantherinus, 315
Synodontis pardalis, 319
Synodontis petricola, 319
Synodontis pfefferi, 311
Synodontis pleurops, 319
Synodontis polli, 319
Synodontis polyodon, 319
Synodontis polystigma, 319
Synodontis pulcher, 319
Synodontis punctifer, 319
Synodontis punctulata, 320
Synodontis punctulatus, 320
Synodontis rebeli, 320, 323
Synodontis resupinata, 320
Synodontis resupinatus, 320
Synodontis ricardoae, 320
Synodontis robbianus, 320
Synodontis robertsi, 320
Synodontis ruandae, 320
Synodontis rufigiensis, 320
Synodontis rukwaensis, 320
Synodontis schall, 315, 320
Synodontis schoutedeni, 321
Synodontis serpentis, 321
Synodontis serrata, 321
Synodontis serratus, 311, 321
Synodontis serratus tanganaicae, 322
Synodontis smithii, 321
Synodontis smiti, 321
Synodontis soloni, 321
Synodontis sorex, 321
Synodontis steindachneri, 321
Synodontis tanganaicae, 321
Synodontis tanganyicae, 322
Synodontis tenuis, 321
Synodontis tessmanni, 322
Synodontis thamalakanensis, 322
Synodontis tholloni, 311

Zootaxa 1418 © 2007 Magnolia Press · 621

Synodontis thomasi, 313
Synodontis thysi, 322
Synodontis tourei, 322
Synodontis unicolor, 322
Synodontis vaillanti, 323
Synodontis vanderwaali, 322
Synodontis velifer, 322
Synodontis vermiculata, 322
Synodontis vermiculatus, 322
Synodontis victoriae, 322
Synodontis violacea, 322
Synodontis violaceus, 322
Synodontis vittatus, 314
Synodontis voltae, 323
Synodontis wamiensis, 320
Synodontis waterloti, 323
Synodontis werneri, 311
Synodontis woosnami, 323
Synodontis xiphias, 323
Synodontis zambesensis, 323
Synodontis zambezensis, 323
Synodontis zambezensis rukwaensis,
320
Synodontis zanzibarica, 323
Synodontis zanzibaricus, 316, 320, 323
synodontis, Pimelodus, 313
Syriacus, Clarias, 143Ttaakree, Hypophthalmus, 362
taakree, Proeutropiichthys, 362
taakree burmanicus,
Proeutropiichthys, 362
tachiraense, Chaetostoma, 230
tachiraensis, Chaetostoma, 230
Tachisurus agassizii, 39
Tachisurus gulosus, 40
Tachisurus jordani, 33
Tachisurus lentiginosus, 33
Tachisurus longicehpalus, 39
Tachisurus longicephalus, 39
Tachisurus upsulonophorus, 44
tachisurus, Pimelodus, 431
Tachysurus, 48, 431
Tachysurus (Pararius) berneyi, 47
Tachysurus (Pararius) godfreyi, 45
Tachysurus atroplumbeus, 32
Tachysurus broadbenti, 51
Tachysurus crassus, 434, 435
Tachysurus emmelane, 40
Tachysurus equatorialis, 40
Tachysurus evermanni, 40
Tachysurus landanensis, 57
Tachysurus liropus, 41
Tachysurus machadoi, 44
Tachysurus oblongus, 435
Tachysurus planus, 434, 435
Tachysurus sinensis, 431
Tachysurus steindachneri, 40
taczanowskii, Arges, 67

taczanowskii, Astroblepus, 67
taczanowskii, Chaetostoma, 231
Taczanowskii, Chaetostomus, 231
taczanowskii Trichomycterus, 424
taenia, Trichomycterus, 407, 424
taenia transandianum, Pygidium, 424
taeniatus, Macrones (Liocassis), 102
taeniatus, Plecostomus, 255
taeniatus, Pseudobagrus, 102
taeniatus, Trachelyopterichthys, 78
taeniatus, Trachelyopterus, 78
Taenionema, 330
Taenionema steerei, 330, 331
taeniophora, Pimelodella, 195
taeniophorus, Pimelodus(Pimelodella), 195
taeniops, Trichomycterus, 424
taenioptera, Pimelodella, 195
taeniura, Pangasius, 326
taimensis, Hisonotus, 249
taimensis, Microlepidogaster, 249
taiosh, Callichthys, 125
taiwanensis, Pseudobagrus, 102
tamanae, Oxyloricaria, 300
tamanae, Sturisomatichthys, 300
tamboensis, Ancistrus, 224
tamoata, Callichthys, 111
Tandanus, 351
Tandanus bostocki, 352
Tandanus coatesi, 348
Tandanus tandanus, 352
tandanus, Tandanus, 352
tanensis, Physailia somalensis, 360
tanganaicae, Synodontis, 321
tanganaicae, Synodontis serratus, 322
tanganicanus, Auchenoglanis
occidentalis, 154
Tanganikallabes, 152
Tanganikallabes mortiauxi, 152
tanganyicae, Synodontis, 322
tanganyikaensis, Malapterurus, 302
tanoensis, Malapterurus, 302
tapanahoniensis, Heptapterus, 184
tapanahoniensis, Hypostomus, 260
tapanahoniensis, Hypostomus
gymnorhynchus, 261
tapatapae, Pimelodella, 195
tapeinopterus, Encheloclarias, 150
tapeinopterus, Heterobranchus, 149,
150
taphorni, Cochliodon, 261
taphorni, Farlowella, 239
taphorni, Hypostomus, 261
taphrophilus, Hemibagrus, 108
tapijara, Hypostomus, 261
tapirape, Otocinclus, 275
tarabinii, Pardiglanis, 163
taroba, Trichomycterus, 424
Tatia, 77622 · Zootaxa 1418 © 2007 Magnolia Press

Tatia aulopygia, 77
Tatia boemia, 77
Tatia brunnea, 77
Tatia concolor, 73
Tatia creutzbergi, 77
Tatia dunni, 77
Tatia galaxias, 77
Tatia gyrina, 77
Tatia intermedia, 77
Tatia musaica, 77
Tatia neivai, 77
Tatia perugiae, 73
Tatia punctata, 73
Tatia reticulata, 73
Tatia romani, 73
Tatia simplex, 78
Tatia strigata, 78
taunayi, Ancistrus, 224
Taunayia, 203
Taunayia bifasciata, 203
Taunayia marginata, 203
taurus, Aspidoras, 110
taxistigma, Henonemus, 406
taxistigma, Ochmacanthus, 405, 406
taylori, Arius, 40, 48
taylori, Hemipimelodus, 48
taylori, Neoarius, 48
taylori, Noturus, 214
taytayensis, Hito, 375, 377
taytayensis, Hitoichthys, 375, 377
taytayensis, Pterocryptis, 377
tchadiensis, Auchenoglanis
occidentalis, 154tchangi, Pseudecheneis, 401
teaguei, Parabranchioica, 410, 411
teaguei, Trachelyopterus, 80
teaguei, Trachycorystes, 80
tectirostris, Chaetostomus, 221
teffeana, Hemiloricaria, 247
teffeana, Loricaria, 247
teffeanus, Rhinodoras, 176
teijsmanni, Clarias, 147
telchita, Pimelodus, 394
telchitta, Glyptothorax, 394
telchitta, Pimelodus, 394
telchitta sufii, Glyptothorax, 394
temminckianus, Bagrus, 55
temminckii, Ancistrus, 224
Temminckii, Bagrus, 55
Temminckii, Hypostomus, 224
tenebricosa, Pseudolaguvia, 402
tenebricus, Leiocassis, 443
tenebrosa, Phenacorhamdia, 189
tenebrosus, Imparfinis, 189
tenella, Rhamdia, 199
tengana, Batasio, 86, 442
tengana, Pimelodus, 86
tengara, Mystus, 97, 98
tengara, Pimelodus, 97

FERRARIS

tenius, Otolithus (Arius), 435
tentaculatus, Lasiancistrus, 265
tenue, Pygidium, 424
tenuicauda, Paraphractura, 26, 27
tenuicauda, Phractura, 27
tenuicauda, Plecostomus, 298
tenuicauda, Squaliforma, 298
tenuifurcatus, Leiocassis, 93
tenuirostre, Sturisoma, 300
tenuirostris, Oxyloricaria, 300
tenuis, Belonoglanis, 24
tenuis, Clarias, 148
tenuis, Heptapterus, 184
tenuis, Hypostomus, 298
tenuis, Macrones (Pseudobagrus), 103
tenuis, Otolithus (Arius), 435
tenuis, Pseudobagrus, 102
tenuis, Squaliforma, 298
tenuis, Synodontis, 321
tenuis, Trichomycterus, 424
tenuispinis, Amblyceps, 18
tenuispinis, Arius, 52, 56
tenuispinis, Plicofollis, 52
ternetzi, Hypostomus, 261
ternetzi, Opsodoras, 175
ternetzi, Plecostomus, 261
ternetzi, Typhlobelus, 426, 427
tessmanni, Synodontis, 322
Tetracamphilius, 27
Tetracamphilius angustifrons, 27
Tetracamphilius clandestinus, 27
Tetracamphilius notatus, 27
Tetracamphilius pectinatus, 27
tetranema, Bathybagrus, 154, 155
tetranema, Pseudolais, 328
Tetranematichthys, 78
Tetranematichthys quadrifilis, 78
Tetranematichthys wallacei, 444
Tetranesodon, 42
Tetranesodon conorhynchus, 42tetraradiata, Listrura, 408
teugelsi, Chrysichthys, 159
teugelsi, Clariallabes, 138teugelsi, Malapterurus, 302
teugelsi, Paramphilius, 25
thalassina, Netuma, 49
thalassina jacksonensis, Netuma, 49
thalassinus, Bagrus, 49
thamalakanensis, Synodontis, 322
theobaldi, Chrysichthys, 105
theodorae, Clarias, 139, 147
theresiae, Arges, 67
theresiae, Astroblepus, 67
therezinae, Ageneiosus(Pseudageneiosus), 69
thienemanni, Clarias, 148
tholloni, Synodontis, 311
thomasi, Chrysichthys delhezi, 156
thomasi, Notoglanidium, 162

thomasi, Synodontis, 313
thomersoni, Rhinodoras, 178
thomsoni, Chaetostoma, 231
thomsoni, Chaetostomus, 231
thonneri, Chrysichthys, 159
thoracata, Megalechis, 129
thoracatum, Hypoptopoma, 249, 250
thoracatum cayennae, Hoplosternum,
128
thoracatum surinamensis,
Hoplosternum, 129
thoracatus, Auchenipterichthys, 71
thoracatus, Auchenipterus, 71
thoracatus, Callichthys, 129
thoracicus, Auchenipterus, 71
thorectes, Pseudohemiodon, 288
Thrichomycterus, 404
Thrichomycterus macraei, 405
thrissoceps, Loricaria, 295
thrissoceps, Rineloricaria, 295
thunberg, Pimelodus, 35
thunbergianus, Plotosus, 350
Thysanocara, 219
thysi, Chrysichthys, 159
thysi, Doumea, 25
thysi, Malapterurus, 301
thysi, Synodontis, 322
thysvillensis, Clarias (Allabenchelys)
submarginatus, 141
tianquanensis, Pareuchiloglanis, 400
tibicen, Aspredinichthys, 58
tibicen, Aspredo, 57, 58
tietensis, Hypostomus, 261
tietensis, Otocinclus(Microlepidogaster), 290
tietensis, Plecostomus, 261
tietensis, Pseudotocinclus, 290
tigrinum, Brachyplatystoma, 331
tigrinum, Pseudoplatystoma, 341
tigrinus, Batasio, 86
tigrinus, Merodontotus, 330, 331
tigrinus, Trichomycterus, 415
tigris, Lasiancistrus, 288, 289
tigris, Pseudolithoxus, 289
tihoni, Gymnallabes, 152
tihoni, Platyallabes, 152
tilhoi, Gephyroglanis, 155
tiong, Glyptosternum, 395
tiraquae, Pygidium, 423
titcombi, Hatcheria, 405
tocantinensis, Corumbataia, 232
tocantinsensis, Aguarunichthys, 329
tocantinsi, Tridentopsis, 426
Tocantinsia, 78
Tocantinsia depressa, 78
Tocantinsia piresi, 78
tokiensis, Pseudobagrus, 103
tombador, Ancistrus, 225
tonggol, Ariodes, 52

CHECKLIST OF CATFISHES, RECENT AND FOSSIL

tonggol, Arius, 52
tonggol, Plicofollis, 52
topavae, Hypostomus, 261
topavae, Plecostomus, 261
torbesensis, Cordylancistrus, 232
torbesensis, Pseudancistrus, 231, 232
tordilho, Lepthoplosternum, 129
torobo, Oloplotosus, 349
torosilabris, Liocassis, 108
torosus, Aguarunichthys, 329
Torpedo, 301
torrenticola, Harttia, 242
torrentis, Pterocryptis, 377
torrentis, Silurus, 377
tortosus, Sisor, 402
totae, Pygidium, 412
totae, Rhizosomichthys, 412
tourei, Synodontis, 322
trachacanthus, Bagrus, 89
Trachelyichthys, 78
Trachelyichthys decaradiatus, 78
Trachelyichthys exilis, 78
Trachelyopterichthys, 78
Trachelyopterichthys anduzei, 78
Trachelyopterichthys taeniatus, 78
Trachelyopterini, 68
Trachelyopterus, 68, 79
Trachelyopterus albicrux, 79Trachelyopterus amblops, 79
Trachelyopterus ceratophysus, 79
Trachelyopterus coriaceus, 79
Trachelyopterus coriaceus maculosus,
79
Trachelyopterus fisheri, 79
Trachelyopterus galeatus, 79
Trachelyopterus insignis, 79
Trachelyopterus lacustris, 79
Trachelyopterus leopardinus, 80
Trachelyopterus lucenai, 80
Trachelyopterus peloichthys, 80
Trachelyopterus striatulus, 80
Trachelyopterus taeniatus, 78
Trachelyopterus teaguei, 80
trachipomus, Bagrus, 55
Trachycorystes, 80
Trachycorystes albicrux, 79Trachycorystes analis, 80
Trachycorystes coracoideus, 71
Trachycorystes cratensis, 80
Trachycorystes fisheri, 79
Trachycorystes insignis badeli, 80
Trachycorystes insignis peloichthys, 80
Trachycorystes jokeannae, 80
Trachycorystes leopardinus, 80
Trachycorystes magdalenae, 79
Trachycorystes porosus, 81
Trachycorystes teaguei, 80
Trachycorystes trachycorystes, 80
Trachycorystes typus, 81

Zootaxa 1418 © 2007 Magnolia Press · 623

trachycorystes, Auchenipterus, 80
trachycorystes, Trachycorystes, 80
Trachydoras, 179
Trachydoras atripes, 179
Trachydoras brevis, 179
Trachydoras microstomus, 179
Trachydoras nattereri, 179
Trachydoras paraguayensis, 179
Trachydoras steindachneri, 179
Trachyglanis, 28
Trachyglanis ineac, 28
Trachyglanis intermedius, 28
Trachyglanis minutus, 28
Trachyglanis sanghensis, 28
Trachymochlus, 431
Trachymochlus cupido, 431, 432
trachyparia, Oxydoras, 179
Trachypoma, 404
Trachypoma marmoratum, 404
transandianum, Pygidium taenia, 424
transandianus, Trichomycterus, 424
transfasciatus, Rhamdioglanis, 202
transitoria, Pimelodella, 195transmontanus, Batrochoglanis, 353
transmontanus, Pseudopimelodus, 353
transvaaliensis, Amphilius, 23
trautmani, Noturus, 214
travancoria, Batasio, 86
travassosi, Otothyris, 276
travassosi, Pygidium, 414
trefauti, Trichomycterus, 425
treitlii, Corydoras, 126
trewavasae, Glyptothorax, 394
Trewavasia, 433
Trewavasia carinata, 433
triacanthopomus, Henonemus, 440
triactis, Leporacanthicus, 266
Trichogenes, 403, 414
Trichogenes longipinnis, 414
Trichogeninae, 403
Trichomycteridae, 403
Trichomycterini, 403
trichomycteroides, Paramphilius, 25
Trichomycterus, 403, 413
Trichomycterus aguarague, 441
Trichomycterus albinotatus, 414
Trichomycterus alternatus, 414
Trichomycterus alterus, 415
Trichomycterus amazonicus, 406
Trichomycterus areolatus, 415
Trichomycterus arleoi, 415
Trichomycterus auroguttatus, 415
Trichomycterus bahianus, 415
Trichomycterus banneaui, 415
Trichomycterus barbatula, 422
Trichomycterus barbouri, 415
Trichomycterus belensis, 415
Trichomycterus bogotensis, 415
Trichomycterus bomboizanus, 416

Trichomycterus borellii, 416
Trichomycterus boylei, 416
Trichomycterus brasiliensis, 416
Trichomycterus brasiliensis itatiayae,
419
Trichomycterus brasiliensis tristis, 416
Trichomycterus caliensis, 416
Trichomycterus candidus, 416
Trichomycterus castroi, 416
Trichomycterus catamarcensis, 416
Trichomycterus caudofasciatus, 416
Trichomycterus celsae, 416
Trichomycterus chaberti, 417
Trichomycterus chapmani, 417
Trichomycterus chiltoni, 417
Trichomycterus chungaraensis, 417
Trichomycterus concolor, 417
Trichomycterus conradi, 417
Trichomycterus corduvensis, 417
Trichomycterus cubataonis, 425
Trichomycterus davisi, 417
Trichomycterus diabolus, 417
Trichomycterus dispar, 417
Trichomycterus dorsotriatus, 417
Trichomycterus duellmani, 418
Trichomycterus eichorniarum, 407
Trichomycterus eigenmanni, 423
Trichomycterus emanueli, 418
Trichomycterus fassli, 418
Trichomycterus gabrieli, 418
Trichomycterus giganteus, 418
Trichomycterus goeldii, 418
Trichomycterus gorgona, 418
Trichomycterus gracilis, 422
Trichomycterus guaraquessaba, 418
Trichomycterus guianensis, 418
Trichomycterus hasemani, 418
Trichomycterus herberti, 407
Trichomycterus heterodontus, 418
Trichomycterus iheringi, 419
Trichomycterus immaculatus, 419
Trichomycterus incae, 423
Trichomycterus inermis, 324
Trichomycterus itacambirussu, 419
Trichomycterus itacarambiensis, 419
Trichomycterus itatiayae, 419
Trichomycterus jacupiranga, 419
Trichomycterus jequitinhonhae, 419
Trichomycterus johnsoni, 419
Trichomycterus knerii, 419
Trichomycterus landinga, 419
Trichomycterus laticeps, 407
Trichomycterus latidens, 419
Trichomycterus latistriatus, 420
Trichomycterus laucaensis, 420
Trichomycterus lewi, 420
Trichomycterus longibarbatus, 420
Trichomycterus maculatus, 415
Trichomycterus maracaiboensis, 420

624 · Zootaxa 1418 © 2007 Magnolia Press

Trichomycterus marmoratus, 415
Trichomycterus mboycy, 420
Trichomycterus mendozensis, 413
Trichomycterus meridae, 420
Trichomycterus migrans, 420
Trichomycterus mimonha, 420
Trichomycterus minutus, 413
Trichomycterus mirissumba, 420
Trichomycterus mondolfi, 421
Trichomycterus motatanensis, 421
Trichomycterus naipi, 421
Trichomycterus nigricans, 414, 421
Trichomycterus nigromaculatus, 421
Trichomycterus palleus, 415
Trichomycterus pantherinus, 421
Trichomycterus paolencis, 421
Trichomycterus papilliferus, 421
Trichomycterus paquequerensis, 421
Trichomycterus pardus, 423
Trichomycterus pauciradiatus, 440
Trichomycterus pentlandi, 423
Trichomycterus pictus, 423
Trichomycterus piurae, 421
Trichomycterus plumbeus, 421
Trichomycterus poeyanus, 423
Trichomycterus potschi, 421
Trichomycterus pradensis, 422
Trichomycterus proöps, 408
Trichomycterus pseudosilvinichthys,
422
Trichomycterus punctatissimus, 422
Trichomycterus punctulatus, 422
Trichomycterus pusillus, 411
Trichomycterus ramosus, 422
Trichomycterus regani, 422
Trichomycterus reinhardti, 422
Trichomycterus retropinnis, 422
Trichomycterus riojanus, 422
Trichomycterus rivulatus, 422
Trichomycterus roigi, 423
Trichomycterus romeroi, 423
Trichomycterus santaeritae, 423
Trichomycterus spegazzinii, 423
Trichomycterus spelaeus, 423
Trichomycterus spilosoma, 423
Trichomycterus stawiarski, 423
Trichomycterus stellatus, 424
Trichomycterus straminius, 424
Trichomycterus striatus, 424
Trichomycterus taczanowskii, 424
Trichomycterus taenia, 407, 424
Trichomycterus taeniops, 424
Trichomycterus taroba, 424
Trichomycterus tenuis, 424
Trichomycterus tigrinus, 415
Trichomycterus transandianus, 424
Trichomycterus trefauti, 425
Trichomycterus triguttatus, 425
Trichomycterus tupinamba, 425

FERRARIS

Trichomycterus unicolor, 425
Trichomycterus variegatus, 425
Trichomycterus venulosus, 425
Trichomycterus vermiculatus, 425
Trichomycterus vittatus, 425
Trichomycterus weyrauchi, 425
Trichomycterus yuska, 425
Trichomycterus zonatus, 425
tricornis, Hopliancistrus, 249
Tridens, 403, 426
Tridens brevis, 426
Tridens melanops, 426
Tridensimilis, 426
Tridensimilis brevis, 426
Tridensimilis venezuelae, 426
Tridentinae, 403
Tridentopsis, 426
Tridentopsis cahuali, 426
Tridentopsis pearsoni, 426
Tridentopsis tocantinsi, 426
trifasciatum, Cyclopium, 67
trifasciatus, Astroblepus, 67
trifasciatus, Pachypterus, 359
trigonocephalus, Sorubim, 342
triguttatum, Pygidium, 425
triguttatus, Trichomycterus, 425
trilineatoides, Glyptothorax, 394
trilineatus, Corydoras, 126
trilineatus, Glyptothorax, 394
trilineatus, Leiocassis, 103
trilineatus, Pseudobagrus, 103
trilobatus, Chiloglanis, 308
trimaculatus, Leptodoras, 174
trimaculatus, Nemadoras, 174
trimaculatus, Opsodoras, 174
trimaculatus, Oxydoras, 174trinitatis, Ancistrus, 225trinitatis, Chaetostomus, 225
triostegus, Silurus, 380
triradiatus, Ancistrus, 225
triradiatus martini, Ancistrus, 223
triseriatus, Corydoras nattereri, 121
tristis, Trichomycterus brasiliensis,
416
troelli, Claibornichthys, 434, 435
Trogloglanis, 216
Trogloglanis pattersoni, 216
trombetensis, Harttia, 242
troschelii, Sciades, 53, 54
truncatorostris, Chasmocranus, 183
truncatum, Platystoma, 341
truncatus, Amblydoras, 168
truncatus, Arius, 43
truncatus, Belodontichthys, 368
truncatus, Cryptarius, 43
truncatus, Liocassis, 103
truncatus, Pseudobagrus, 103
truncatus, Synodontis marmoratus, 317
tsanensis, Clarias, 143

tubbi, Pangasius, 328
tucumanensis, Loricaria, 269
tukano, Corydoras, 126
tumbanus, Eutropius, 365
tumbanus, Schilbe, 365
tupinamba, Trichomycterus, 425
tuyra, Arius, 40
tuyra, Cathorops, 40
tuyrensis, Dasyloricaria, 234
tuyrensis, Loricaria, 234
tweediei, Wallagonia, 381
Tympanopleura, 68
Tympanopleura alta, 69
Tympanopleura nigricollis, 68
Tympanopleura piperata, 68, 70
typhla, Rhamdia laticauda, 198
Typhlobagrus, 189
Typhlobagrus kronei, 189, 192
Typhlobelus, 426
Typhlobelus guacamaya, 426
Typhlobelus lundbergi, 426
Typhlobelus macromycterus, 426
Typhlobelus ternetzi, 426
typica, Doumea, 24, 25
typus, Gagata, 386
typus, Gymnallabes, 159
typus, Helicophagus, 324, 325
typus, Ketengus, 45
typus, Micronema, 375
typus, Parahemiodon, 269, 270
typus, Phyllonemus, 164
typus, Pinirampus, 339
typus, Rhinoglanis, 310, 311
typus, Trachycorystes, 81
typus heterocercalis, Gymnallabes,
150
U
uatumensis, Harttia, 242
ubangensis Auchenoglanis, 162
ubangensis, Bagrus, 84
ubidiai, Astroblepus, 67
ubidiai, Cyclopium, 67
ucamara, Lepthoplosternum, 129
ucayalensis, Ageneiosus, 70
ucayalensis, Hemiancistrus, 283, 285
ucayalensis, Hassar, 171
ucayalensis, Loricariichthys, 271
ucayalensis, Peckoltia, 285
Uegitglanis, 152
Uegitglanis zammaranoi, 152, 153
uelensis, Clariallabes, 138
uelensis, Clarias (Allabenchelys), 138
umbrosa, Cetopsis, 133
unae, Hypostomus, 261
Unae, Plecostomus, 261
uncinatus, Arius, 36
undecimalis, Chaetostomus, 292
undecimalis, Pterygoplichthys, 292
undecimalis, Silurus, 431

CHECKLIST OF CATFISHES, RECENT AND FOSSIL

underwoodi, Rhamdia, 198
undulatus, Corydoras, 126
unicolor, Aphanotorulus, 225
unicolor, Plecostomus, 225
unicolor, Plotosus, 350, 352
unicolor, Pygidium, 425
unicolor, Synodontis, 322
unicolor, Trichomycterus, 425
unidorsalis, Helogenes, 134
unifasciata, Phenacorhamdia, 189
unifasciatum, Cyclopium, 67
unifasciatus, Astroblepus, 67
uniformis, Amphilius baudoni, 21
uniformis, Chrysichthys, 159
unitius, Silurus, 350
upiensis, Pangasius pangasius, 327
Upsilodus, 247
Upsilodus victori, 247
upsulonophorus, Tachisurus, 44
uracantha, Loricaria, 240
uracantha, Rineloricaria, 240
uracantha rupestre, Loricaria, 240
uracanthus, Fonchiiichthys, 240
uranoscopus, Amphilius, 23
uranoscopus, Anoplopterus, 21, 23
uranoscopus, Doras, 173, 174
uranoscopus, Megalodoras, 174
uranoscopus, Schilbe, 362, 365
urbaini, Cryptopterus, 373
urbaini, Ompok, 373
urichi, Caecorhamdia, 197, 202
Urinophilus, 427
Urinophilus diabolicus, 411
Urinophilus erythrurus, 427
urostigma, Bagrus, 84
urostriatum, Decapogon, 127
urostriatum, Dianema, 127
urotriatum, Decapogon, 127
urua, Pimelodus, 359
uruguayensis, Ageneiosus, 70
uruguayensis, Hypostomus, 261
uruyensis, Helogenes, 134
uruyensis, Helogenes marmoratus, 134
ussuriensis, Bagrus, 92, 100
ussuriensis, Leiocassis, 101
ussuriensis, Pelteobagrus, 100
ussuriensis, Pseudobagrus, 101
utarus, Arius, 48
utarus, Neoarius, 48
utik, Arius, 35
Vvacha, Eutropiichthys, 358
vacha, Pimelodus, 358
vachellii, Bagrus, 101
vachellii, Pelteobagrus, 101
vagum, Chaetostoma, 231
vagus, Chaetostomus, 231
vaillanti, Arges, 67
vaillanti, Astroblepus, 67

Zootaxa 1418 © 2007 Magnolia Press · 625

vaillanti, Hypostomus, 261
vaillanti, Liocassis, 105
Vaillanti, Plecostomus, 261
vaillanti, Pseudomystus, 105
vaillanti, Synodontis, 323
vaillantii, Bagrichthys, 83
Vaillantii, Bagroides, 83
vaillanti, Brachyplatystoma, 331
Vaillanti, Platystoma, 330, 331
vaillantii, Silurus, 331
valencia, Lithogenes, 267
valenciennesi, Ageneiosus, 69
valenciennesi, Osteogeneiosus, 51
valenciennesii, Genidens, 44
Valenciennesii, Loricaria, 247
valenciennis, Parapimelodus, 335
valenciennis, Pimelodus, 335
valeya, Wallago attu, 381
vanceae, Astroblepus, 67
vanceae, Cyclopium, 67
vandeli, Arius, 50
Vandellia, 403, 427
Vandellia Balzanii, 427
Vandellia beccarii, 427
Vandellia cirrhosa, 427
Vandellia gigantea, 427
Vandellia hasemani, 411
Vandellia hematophaga, 410
Vandellia plazaii, 427
Vandellia sanguinea, 427
Vandellia wieneri, 411
Vandelliini, 403
vandenhoutei, Clarias, 144
vanderwaali, Synodontis, 322
vandeweyeri, Eutropiellus, 361
vanigonis, Otolithus (Arius), 434
Vannutellii, Rhinoglanis, 311
variabilis, Clariallabes, 138variegata, Crossoloricaria, 233
variegata, Loricaria, 232, 233
variegata venezuelae, Loricaria, 233
variegatum, Amblyceps, 19
variegatus, Pimelodus, 13, 14, 15
variegatus, Akysis, 15variegatus, Akysis variegatus, 14variegatus, Hemibagrus, 91
variegatus, Microglanis, 355
variegatus, Trichomycterus, 425
variipictus, Hypostomus, 261
variipictus, Neoplecostomus, 274
variipictus, Plecostomus, 261
varimaculosus, Hypostomus, 261
varimaculosus, Plecostomus, 261
variolosus, Arius, 41
variolosus, Pseudopimelodus, 355
variolus, Ancistrus, 225
variolus, Chaetostomus, 225
variostictus, Hypostomus, 261
variostictus, Plecostomus, 261

varispinis, Clarias, 147varius, Acrochordonichthys, 13
varius, Akysis, 15
varius, Liposarcus, 292
vasquezi, Chaetostoma, 231
vazferreirai, Homodiaetus, 406
velatus, Encheloclarias, 150
velifer, Chrysichthys, 157, 158
velifer, Gephyroglanis, 157
velifer, Pimelodus, 203
velifer, Synodontis, 322
velites, Aspidoras, 110
velox, Hemibagrus, 91
velutinus, Hemipimelodus, 48
velutinus, Neoarius, 48
venaticus, Bagrus, 55
venezuelae, Chaetostoma, 231
venezuelae, Corymbophanes, 231
venezuelae, Crossoloricaria, 233
venezuelae, Haemomaster, 405
venezuelae, Loricaria variegata, 233
venezuelae, Tridensimilis, 426
venezuelanus, Corydoras, 112
venezuelensis, Farlowella, 239
venezuelensis, Farlowella acus, 239
venosus, Arius, 36ventrale, Cyclopium, 68
ventralis, Astroblepus, 68
ventralis, Cetopsis, 135
ventromaculatus, Hypostomus, 262
venulosum, Pygidium, 425
venulosus, Trichomycterus, 425
verbeekii, Pseudeutropius, 326
verecunda, Pterocryptis, 377
verecundus, Ancistrus, 225verissimi, Decapogon, 127vermicularis, Hypostomus, 262
vermicularis, Plecostomus, 262
vermiculata, Ancistrus vittatus, 285
vermiculata, Peckoltia, 285
vermiculata, Synodontis, 322
vermiculatum, Pygidium, 425
vermiculatus, Synodontis, 322
vermiculatus, Trichomycterus, 425
verres, Hypostomus, 262
verrucosa, Aspredo, 60
verrucosa, Parakysis, 15, 16
verrucosus, Arius, 45
verrucosus, Bunocephalus, 60
verrucosus, Hemiarius, 45
verrucosus, Parakysis, 16
verrucosus, Platystacus, 58, 60
verrucosus, Silurus, 60
versicolor, Pimelodus, 44
vertagus, Bagrus, 55
vespa, Akysis, 15vespertinus, Ameiurus, 206
vespertinus, Ictalurus, 206
vestigipinnis, Hemipsilichthys, 281

626 · Zootaxa 1418 © 2007 Magnolia Press

vestigipinnis, Pareiorhaphis, 281
vestitus, Otocinclus, 274, 275
vetula, Loricaria, 278
vetula, Paraloricaria, 278
vicinus, Hypostomus, 278
victori, Upsilodus, 247
victoriae, Synodontis, 322
viedmensis, Diplomystes, 166
viedmensis, Olivaichthys, 166
viedmensis cuyanus, Diplomystes, 166
viedmensis mesembrinus, Diplomystes,
166
vietnamicus, Hemibagrus, 91
vietnammicus, Hemibagrus, 91
vigilis, Microsynodontis, 310
Villarius, 208
Villarius pricei, 208, 209
villarsi, Plecostomus, 298
villarsi, Squaliforma, 298
villiersi, Physailia, 360
villosus, Arius, 166
villosus, Batrochoglanis, 353
villosus, Lithogenes, 267
villosus, Pseudopimelodus, 353
villosus butcheri, Pseudopimelodus,
352
Vilsoni, Pimelenotus, 197, 200
vinciguerrae, Clarias, 143
Vinciguerrae, Exostoma, 385
vinhensis, Pelteobagrus virgatus, 101
violacea, Synodontis, 322
violaceus, Synodontis, 322
virescens, Corydoras, 123
virescens, Plecostomus, 298
virescens, Squaliforma, 298
virgatus, Aoria, 101
virgatus, Leiocassis, 101
virgatus, Pelteobagrus, 101
virgatus, Pseudobagrus, 101
virgatus vinhensis, Pelteobagrus, 101
virginiae, Corydoras, 126
virgo, Ageneiosus, 70
virgulatus, Aspidoras, 110
viridescens, Gogangra, 395
viridescens, Pimelodus, 395
viscosus, Pimelodus, 215
viscosus, Silurus, 215
vitata, Ansorgia, 361
vittata, Ansorgia, 360, 361
vittata, Farlowella, 239
vittata, Macrones, 106
vittata, Peckoltia, 285
vittata, Pimelodella, 195
vittata bistriata, Ansorgia, 361
vittatus, Ageneiosus, 70
vittatus, Chaetostomus, 283, 284, 285
vittatus, Corydoras, 126
vittatus, Corydoras blochi, 126
vittatus, Mystus, 98

FERRARIS

vittatus, Otocinclus, 275
vittatus, Plotosus, 350
vittatus, Pseudorhamdia, 195
vittatus, Silurus, 98
vittatus, Silurus (Callichrus), 372
vittatus, Synodontis, 314
vittatus, Trichomycterus, 425
vittatus horai, Mystus (Mystus), 95
vittatus vermiculata, Ancistrus, 285
viviparus, Arius, 55
viviparus, Plotosus, 350
vogti, Atopochilus, 304
volcanensis, Lasiancistrus, 264
voltae, Chiloglanis, 308
voltae, Irvineia, 358
voltae, Synodontis, 323
Vorhisia, 434
Vorhisia vulpes, 434, 435
Vorhisiidae, 434
votouro, Hemiancistrus, 244
voulezi, Rhamdia branneri, 201
vulgaris, Pimelodus, 206
vulpeculus, Pimelodus, 206
vulpes, Pimelodus, 211
vulpes, Vorhisia, 434, 435
Wwaandersii, Helicophagus, 325
wagenaari, Chrysichthys, 159
wagneri, Pimelodus, 201
waiampi, Hypostomus, 262
walkeri, Chrysichthys, 159, 160
walkeri, Clarias, 141
walkeri, Notoglanidium, 162
walkeri, Pseudeutropius atherinoides,359
wallacei, Tetranematichthys, 444
Wallago, 380
Wallago attu, 380
Wallago attu valeya, 381
Wallago dinema, 368
Wallago heterorhynchus, 368
Wallago krattensis, 374
Wallago leerii, 380, 381
Wallago leiacanthus, 371, 372
Wallago maculatus, 381
Wallago micropogon, 381
Wallago miostoma, 372
Wallago nebulosus, 381
Wallago Russellii, 381
Wallagonia, 380
Wallagonia tweediei, 381
wallagoo, Silurus, 381
walrechti, Sciadeichthys (Selenaspis),
53
wamiensis, Leptoglanis, 29
wamiensis, Synodontis, 320
wamiensis, Zaireichthys, 29
wangi, Pseudobagrus, 108
Watawata, 250

waterhousii, Neoplotosus, 346
waterloti, Chiloglanis niloticus, 307
waterloti, Synodontis, 323
watwata, Hypostomus, 262
weberi, Callichrous, 373
weberi, Ompok, 373
weddellii, Anadoras, 169
weddellii, Doras, 169
weitzmani, Corydoras, 126
werneri, Clarias, 148
werneri, Synodontis, 311
Wertheimeri, Plecostomus, 285
wertheimeri, Pogonopoma, 285
Wertheimeria, 179
Wertheimeria maculata, 179
wesselii, Pimelodella, 195
Wesselii, Pimelodus (Pseudorhamdia),195
westermanni, Bergiaria, 330
Westermanni, Pimelodus, 329, 330
weyrauchi, Pygidium, 425
weyrauchi, Trichomycterus, 425
whymperi, Arges, 68
whymperi, Astroblepus, 68
wieneri, Plectrochilus, 411
wieneri, Vandellia, 411
wilderi, Hassar, 171
wilsoni, Eomacrones, 87
wilsoni, Hemiancistrus, 244
wilsoni, Macronoides, 87
winzi, Hypostomus, 262
winzi, Plecostomus, 262
witmeri, Pimelodella, 195
Wittei, Auchenoglanis, 154wittenburgii, Pseudobagrus, 108
wolfei, Hemiloricaria, 247
wolfei, Rhineloricaria, 247
Wolffii, Bagrus, 98
wolffii, Mystus, 98
wolfi, Rhamdella, 197
woodi, Hassar, 171
woodsi, Rhynchodoras, 178
woosnami, Synodontis, 323
worthingtoni, Bathyclarias, 136
worthingtoni, Dinotopterus, 136
wotroi, Corydoras, 120
wrightiana, Oxyropsis, 276
wrightii, Oxyropsis, 276
wuchereri, Hypostomus, 262
wuchereri, Pimelodus, 200
wuchereri, Plecostomus, 262
wui, Pseudobagrus, 101
wurnoense, Nigerium, 98, 99
Wyckii, Bagrus, 91
wyckii, Hemibagrus, 91
wyckioides, Hemibagrus, 91
wyckioides, Macrones (Hemibagrus),
91
wynaadensis, Pterocryptis, 377

CHECKLIST OF CATFISHES, RECENT AND FOSSIL

wynaadensis, Silurus, 375, 377
X
xakriaba, Otocinclus, 275
Xanthocephalus, Pimelodus, 207
Xanthocephalus, Silurus, 207
xanthosteus, Silurus, 379
xanthum, Megalonema, 335
xanthus, Perugia, 335
xenauchen, Galeichthys, 33
Xenocara, 219
Xenocara boliviana, 219
Xenocara brevipinnis, 219
Xenocara bufonia, 219
Xenocara damasceni, 220
Xenocara fulva, 221
Xenocara heterorhynchus, 221
Xenocara macrophthalma, 222
Xenocara montana, 223
Xenocara multispinis, 223
Xenocara occidentalis, 223
Xenocara rothschildi, 221
Xenoclarias, 153
Xenoclarias eupogon, 153
Xenoclarias holobranchus, 153
xenodon, Clarias, 143
xenognathus, Leptoglanis, 25
Xenomystus, 247
Xenomystus gobio, 247
Xenopholis, 433
Xenopholis carinatus, 433
Xenopholoides, 433
xetequepeque, Rhamdia, 202
Xiliphius, 62
Xiliphius kryptos, 62
xinguensis, Corydoras, 126
xinguensis, Glyptoperichthys, 292
xinguensis, Pterygoplichthys, 292
xingui, Rhynchodoras, 178
xiphias, Synodontis, 323
Xiurenbagrus, 20
Xiurenbagrus gigas, 20
Xiurenbagrus xiurenensis, 20
xiurenensis, Xiurenbagrus, 20
Xyliphius, 62
Xyliphius barbatus, 62
Xyliphius kryptos, 62
Xyliphius labrosus, 62
Xyliphius lepturus, 62
Xyliphius lombarderoi, 62
Xyliphius magdalenae, 62
Xyliphius melanopterus, 62Yyangambianus, Eutropius, 365
yangambianus, Schilbe, 365
yaravi, Ancistrus (Hemiancistrus), 273
yaravi, Neblinichthys, 273
yarrelli, Bagarius, 383
Yarrelli, Bagrus, 383
yasi, Epactionotus, 236

Zootaxa 1418 © 2007 Magnolia Press · 627

youngicus, Clarias, 145
youssoufi, Gagata, 386
yuncensis, Pimelodella, 195
yunnanense, Pseudexostoma, 401
yunnanensis, Glyptosternum, 401
yunnanensis brachysoma,
Pseudexostoma, 401
yurubiense, Chaetostoma, 231
yuska, Trichomycterus, 425Zzainaensis, Glyptothorax, 394
Zaireichthys, 25, 28
Zaireichthys brevis, 28
Zaireichthys camerunensis, 28
Zaireichthys dorae, 28
Zaireichthys flavomaculatus, 28
Zaireichthys heterurus, 28
Zaireichthys mandevillei, 28
Zaireichthys rotundiceps, 28
Zaireichthys wamiensis, 29
Zaireichthys zonatus, 28, 29
zairensis, Amphilius, 23
zairensis, Schilbe, 366
zambesensis, Synodontis, 323
zambezensis, Synodontis, 323
zambezensis rukwaensis, Synodontis,
320
zambeziensis, Malapterurus, 302
zammaranoi, Uegitglanis, 152, 153
zanaensis, Glyptothorax, 394
zanzibarica, Synodontis, 323
zanzibaricus, Synodontis, 316, 320,
323
Zathorax, 168
Zathorax gonzalezi, 168
Zathorax monitor, 168, 169
Zathorax nauticus, 169zebra, Hypancistrus, 249
zhujiangensis, Glyptothorax, 394
Zonancistrus, 234
zonatum, Pygidium, 425
zonatus, Acrochordonichthys, 13
zonatus, Caelatoglanis, 383
zonatus, Microglanis, 355
zonatus, Pimelodus, 13
zonatus, Synodontis angelicus, 311
zonatus, Trichomycterus, 425
zonatus, Zaireichthys, 28, 29
zongolicensis, Rhamdia, 199
zuliaensis, Pterygoplichthys, 292
zuloagai, Doraops, 170
Zungaro, 344
Zungaro humboldtii, 344
Zungaro jahu, 344
Zungaro mathisoni, 356
Zungaro zungaro, 344
Zungaro zungaro schultzi, 355
zungaro, Zungaro, 344
zungaro schultzi, Zungaro 355

Zungaropsis, 344
Zungaropsis multimaculatus, 344
zygatus, Corydoras, 126
zygouron, Clarias, 141

628 · Zootaxa 1418 © 2007 Magnolia Press

FERRARIS