〔植探報Vol. 22 : 135 ～ 147，2006〕

東ティモールにおけるマメ類および共生微生物
遺伝資源多様性の保全，2005 年
友岡 憲彦 1)・阿部 健一 2)・Duncan A. VAUGHAN1) ・加賀 秋人 1)・
伊勢村 武久 1)・黒田 洋輔 1)
1) 農業生物資源研究所・遺伝資源研究グループ・集団動態研究チーム
2) 国立民族学博物館・地域研究交流企画センター

Conservation of Legume - Symbiotic Rhizobia Genetic
Diversity in East Timor, 2005
Norihiko TOMOOKA1), Ken-ichi ABE2), Duncan A. VAUGHAN1),
Akito KAGA1), Takehisa ISEMURA1) and Yosuke KURODA1)
1) National Institute of Agrobiological Sciences , Kannondai 2-1-2, Tsukuba, Ibaraki
305-8602, Japan2) The Japan Center for Area Studies , National Museum of Ethnology , 10-1 Senri
Expo Park, Suita, Osaka 565-8511, Japan

Summary
A field survey was conducted in East Timor from May 16 to 24, 2005. As a result, 2accessions of cultivated mungbean (Vigna radiata var. radiata ), 7 of wild mungbean (V. radiatavar. sublobata ), 19 of cultivated rice bean (V. umbellata ), 6 of wild rice bean (V. umbellata ), 5 ofcowpea (V. unguiculata ), 3 of V. trinervia , 1 of V. reflexo-pilosa , 4 of soybean (Glycine max ), 1 ofGlycine sp., 1 of hyacinth bean (Lablab purpureus ) and 3 of common bean (Phaseolus vulgaris )were collected and conserved.
Introduction
Timor is located in the eastern end of Lesser Sunda Islands, surrounded by Indonesia, Papua
New Guinea and Australia(Fig.1). The island is about 800 km length (East to west) and 100 km
width (North to South). It has the highest peak of 2,963 m and has diverse ecological zones.
Timor is divided into approximately two equal area West Timor of Indonesia and East Timor.
Biologically it is located in the transitional zone between Southeastern and Australian floraand fauna, so that unique genetic resources are expected. To understand and to conserve
legume genetic resources of East Timor, a survey was planned. In this survey, conservation of
traditional legume crops, wild relatives and their symbiotic root nodules (rhizobia) are the main
- 135 -

objectives.
Methods
The field survey was conducted in East Timor from May 16 to 24, 2005 (Table 1). The routeof the survey is shown in Fig. 2. Seeds, herbarium specimens and if available root nodules werecollected. Information on collection sites including village name, altitude, latitude, longitude,
habitat and other ecological data together with detailed sketch maps of the collection sites wererecorded as passport data.
Results and Discussion
A summary list of collected materials is shown in Table 2 and detailed passport data ispresented in Table 3. The map locality of the collection sites of wild Vigna spp. is shown in Fig. 3.
Identification of Vigna spp. was done based on the key prepared by Tomooka et al. (2002).
Leguminous crops
Among leguminous crops, rice bean (Vigna umbellata ) and cowpea (V. unguiculata ) are themost popular and are cultivated throughout East Timor. Nineteen accessions of rice bean and 5
accessions of cowpea are collected. However in the market of Maubisse, located at the highest
altitude of ca. 1,500 m, we could not find cowpea or rice bean and only common bean (Phaseolus

vulgaris ) is sold there. Around Maubisse, we observed several common bean fields on road sideslopes together with sweet potato, taro and corn. Three accessions were collected. Mungbean
is sold in some markets such as Baucau and Viqueque. Mungbean seems not so popular in East
Timor. Soybean (Glycine max : three yellow and one black seeded type) is found in markets ofhigher elevation such as Aileu and Ermera provinces. Soybean seeds are cooked by boiling
together with corn. This type of food seems very popular in East Timor and rice bean, cowpea,
common bean are used in this way. These legume seeds are sold mixed with corn seeds in manymarkets. Around Baucau area, we often found velvet bean (Mucuna pruriens var. utilis ) growing
in home gardens. In the area around Bobonaro, hyacinth bean (Lablab purpureus ) fields are
frequently seen. However, we could not collect any seed samples for these two species. Theseleguminous crops seem to have been cultivated in East Timor for long time.
Wild legumes
Eighteen wild legume accessions consist of 4 Vigna and 1 Glycine species were collected.
Among wild legumes, wild mungbean (V. radiata var. sublobata ) and wild rice bean (V .

umbellata ) are the two commonly found species.
Wild mungbean prefers rather dry lowland to mid altitude grassland habitat where eucalypt
trees are sporadically growing (Fig. 3). The altitude of collection sites in this type of habitat
ranged from 2 m to 712 m. However, in one place near Maubisse, we found a wild mungbean
population growing on bare gravel soil at an altitude of 1,298 m (ET21, Table 3). Plants in thispopulation showed unique growth characters which are completely different from the otherpopulations. ET21 plants have spreading long branches crawling on the ground. Leaflets are
extremely small. They have shorter pods (3-4 cm) containing fewer seeds (8-9). They have
- 136 -

adapted to the specific habitat which is characterized by very dry, cold and windy environment
located near the top of mountain road. A population of wild Glycine sp. (ET22) was also foundgrowing in this place. In other typical grassland habitats, wild mungbean showed twining
growth habit and pod length and number of seeds per pod ranged from 4.5 to 6 cm and from12 to 15 seeds, respectively.
Wild rice bean prefers a wetter and cooler habitat compared with wild mungbean. The
altitude of collection sites ranged from 550 to 1,645 m with an average of 992 m (Fig. 3). Outof 7 populations of wild rice bean, plants which have atypical seed coat color were found in 5
populations. These plants with atypical seed coat color (black and pale brown) are considered
to be derived from natural hybridization between wild and cultivated rice bean. Plants in a
population (ET23) showed brighter yellow standard petal color with purple pigment outside.
Their leaves are densely hairy. This wild rice bean population was collected in the highest
altitude (1,645 m) among those collected in East Timor.
Three accessions of V. trinervia were collected from high altitude places (Fig. 3; 1,297- 1,400 m). They are growing on road side slopes with dry soil conditions. Plants arecharacterized by their densely hairy stems and leaves. Local people said goats like to eat thisplant. An accession of V. reflexo-pilosa was collected near Gleno at an altitude of 800 m. The
habitat is a wet road side slope.
Acknowledgements
This research is supported by the Grant-in-Aid for Scientific Research No.16310160, Japan
Society for the Promotion of Science.
References
Tomooka N., D. A. Vaughan, H. Moss and M. Maxted (2002) The Asian Vigna : genus Vigna
subgenus Ceratotropis genetic resources. Kluwer Academic Publishers. 270 pages.

和文摘要
東ティモールは，インドネシア（東南アジア）とニューギニア，オーストラリア大陸との間に位置する長さ 800km 幅 100km 程度の島である．アジアの生物相からオーストラリアの生物相へと移行するウォーレス線の東に位置し，島ごとに独自の生物相をもつウォーレシアと呼ばれる興味深い地域にあたる．アジア的要素として水田，水牛による蹄耕がみられ，オーストラリア的要素としてはユーカリの林が印象的である．太古から繰り返しアジアからの民族移動の波が押し寄せ，多様な民族が島の各地で多様な文化・言語を維持しつつ生活している．我々はこれまでに東南アジア各地に続いてパプアニューギニアでの調査を行ってきた．本調査では，東ティモールのほぼ全域にわたって，道路沿いに調査を行い，伝統的マメ科作物，その近縁野生種および共生している根粒菌の収集保全を行った．

- 137 -

Fig. 1. Locality of East Timor

Fig. 2. Exploration routes and main cities in East Timor

Fig. 3. Collection sites of wild Vigna species in East Timor- 138 -

Table 1. Itinerary of the field survey in East Timor, 2005
Date

Day

Itinerary

Stay

16-MayMon

Narita 11:00 -- GA881 -- 16:50 Bali, Indonesia

Bali

17-MayTueBali 9:10 -- MZ8480 -- 12:00 Dili, East Timor

Dili

18-MayWed

Dili -- Manatuto -- Baucau -- Com

Com

19-MayThu

Com -- Baucau -- Viqueque -- Manatuto -- Dili

Dili

20-MayFri

Dili -- Aileu -- Ainaro

Ainaro21-MaySat

Ainaro -- Bobonaro -- Maliana -- Batugade -- Dili

Dili

22-MaySun

Dili -- Ermera -- Letefoho -- Atsutabe -- Dili

Dili

23-MayMon

Dili 12:45 -- MZ8490 -- 13:40 Bali, Indonesia 22:00 -- GA880

Bali

24-MayTue-- 8:50 Narita

Table 2. Summary list of collected accessions in East Timor
Species

StatusNo. ofaccessions

Vigna radiata

cultivated2

Baucau, Viqueque

300 m

wild

7

Dili, Manatuto, Ainaro

2 - 1298 m

Vigna reflexo-pilosa wild

1

Ermera800 m

Vigna trinervia

wild

3

Aileu, Ermera

1298 - 1400 m

Vigna unguiculata

cultivated5

Baucau, Lautem, Viqueque, Bobonaro

300 - 755 m

Vigna umbellata

cultivated19

Baucau, Viqueque, Cova Lima, Bobonaro, Ermera 300 - 1225 m

wild

8

Baucau, Viqueque, Ainaro, Liquica, Ermera

550 - 1645 m

Glycine max

cultivated

4

Aileu, Ermera

933 - 1225 m

Glycine sp.

wild

1

Ainaro1298 mPhaseolus vulgaris cultivated3

Baucau, Ermera

350 - 1400 m

Total

53

Province- 139 -

Altitude range

Table 3. Passport data of the collected materials in East Timor (2005)
収集品のパスポートデータ
No.

Coll.
Date

Coll.
No.

Species

StatusCollection Site

Site
No.

Altitude(m)

Latitude/Longitude1

May 18 2005ET-1

Vigna radiata var. wild
sublobata

5km E of Dili, Dili Province1

S08-33-40.4 190
E125-38-11.3

2

May 18 2005ET-2

Vigna radiata var. wild
sublobata

5km W of Manatuto, Manatuto
Province2

S08-28-46.1 2
E125-55-08.1

3

May 18 2005ET-3Vigna unguiculata cultivated Baucau new market, Baucau
Province3

S08-27
E126-27350

4May 18 2005ET-4

Phaseolus vulgaris cultivated Baucau new market, Baucau
Province3

S08-27
E126-27350

5May 18 2005ET-5Vigna radiata

cultivated Baucau new market, Baucau
Province3

S08-27
E126-27350

6May 18 2005ET-6-1

Vigna umbellata

cultivated Baucau old market, Baucau
Province

4

S08-27
E126-27300

7May 18 2005ET-6-2

Vigna umbellata

cultivated Baucau old market, Baucau
Province

4

S08-27
E126-27300

8May 18 2005ET-7Vigna umbellata

cultivated E of Baucau, Baucau Province

5

S08-28-12.2 311
E126-29-09.9

9

May 18 2005ET-8Vigna unguiculata cultivated A little S of Lautem, Lautem
Province

6

S08-25-12.5 340
E126-55-28.1

Vigna umbellata

20km SW of Baucau, Baucau
Province

7

S08-34-30.0 550
E126-23-15.9

11 May 19 2005ET-10-1 Vigna umbellata

cultivated Kota Berkoli, Baucau Province

8

S08-36-26.2 680
E126-22-57.5

12 May 19 2005ET-10-2 Vigna umbellata

cultivated Kota Berkoli, Baucau Province

8

S08-36-26.2 680
E126-22-57.5

13 May 19 2005ET-10-3 Vigna umbellata

cultivated Kota Berkoli, Baucau Province

8

S08-36-26.2 680
E126-22-57.5

wild

9

S08-43-24.9 960
E126-22-01.3

15 May 19 2005ET-12-1 Vigna unguiculata cultivated Viqueque market, ViquequeProvince

10

S08-51
E126-21300

16 May 19 2005ET-12-2 Vigna umbellata

cultivated Viqueque market, ViquequeProvince

10

S08-51
E126-21300

17 May 19 2005ET-12-3 Vigna umbellata

cultivated Viqueque market, ViquequeProvince

10

S08-51
E126-21300

cultivated Viqueque market, ViquequeProvince

10

S08-51
E126-21300

10 May 19 2005ET-9

14 May 19 2005ET-11

Vigna umbellata

wildnear Ossu, Viqueque Province

18 May 19 2005ET-13

Vigna radiata

19 May 19 2005ET-14

Vigna radiata var. wild
sublobata

50km S of Manatuto, Manatuto
Province

11

S08-57-16.9 264
E125-59-30.7

20 May 19 2005ET-15

Vigna radiata var. wild
sublobata

45km S of Manatuto, Manatuto
Province

12

S08-53-06.5 712
E125-59-12.7- 140 -

Shading

DisturbancePopulation sizeroad side
grassland

light

high

sporadicallynear thebeachmedium medium a few
plantsHabitat

Growthstage

Seed Sample

Herba Rhizo-rium -bium

flowering → silt
mature

yes

bulk

yes

no

pod length 4.5~5 cm, black podwith short brown hair, 12~13
seeds per pod, seed blackishwith reticulation, hilum slightly
protruding

floweringyes

indiyesvidualno

Soilmarketharvested

yes

bulk

no

no

pod not fully mature, seedblackish with reticulation,
hilum slightly protruding
shiny black seedmarketharvested

yes

bulk

no

no

purple with mottled seedmarketharvested

yes

bulk

no

no

dull green seedmarketharvested

yes

bulk

no

no

dark red seedmarketharvested

yes

bulk

no

no

tan seed

yes

bulk

no

no

yes

bulk

no

no

pod length 10cm, pod pale
brown, 10 seeds per pod, dark
red seedpale brown seedfarmer's
field

medium low

5x5m

farmer's
house

maturesandy

Remarkssilt

harvestedroad side
grasslandlightmedium 2 x 10 m flowering → black
mature
silt

yes

bulk

yes

yes

pod length 6~7 cm, 10~11
seeds per pod, pale brown withblack mottle seed, leaflet shapevariable

farmer's
field

light

low

10 x 10 m mature

clay

yes

bulk

no

no

pod length 8~9 cm, 10 seeds
per pod, smoky red seedfarmer's
field

light

low

10 x 10 m mature

clay

yes

bulk

no

no

pod length 8~9 cm, 9~10 seeds
per pod, dark red seedfarmer's
field

light

low

10 x 10 m mature

clay

yes

bulk

no

no

pale yellowish brown seedroad side
grasslandlightmedium a few
plants

silt

yes

bulk

yes

yes

floweringmarketharvested

yes

bulk

no

no

pod length 6~7 cm, 10 seeds
per pod, pale brown with black
mottle seedshiny black seedmarketharvested

yes

bulk

no

no

pale to dark shiny red seedmarketharvested

yes

bulk

no

no

pale yellowish brown seedmarketharvested

yes

bulk

no

no

dull green seed
pod length 5~5.5 cm, 12~13
seeds per pod, seed blackishwith reticulation, hilum slightly
protruding, large leaflet with
red vein
pod length 5~5.5 cm, 14~15
seeds per pod, seed blackishwith reticulation, hilum slightly
protruding

grassland
witheucaypt

lightmedium sporadicallyflowering → silt
mature

yes

bulk

yes

yes

grassland
witheucalyptmedium medium sporadicallyflowering → silt
mature

yes

bulk

no

no

- 141 -

Table 3. (continued).
No.

Coll.
Date

Coll.
No.

Species

StatusCollection Site

Site
No.

Latitude /Longitude

Altitude(m)

21 May 20 2005ET-16

Vigna umbellata

wild

10km S of Dili, Dili Province

13

S08-36-12.8 630
E125-34-56.0

22 May 20 2005ET-17

Vigna trinervia

wild

20km S of Dili, Aileu Province

14

S08-38-17.9 1297
E125-36-01.5

23 May 20 2005ET-18

Glycine max

cultivated Aileu market, Aileu Province

15

24 May 20 2005ET-19

Glycine max

cultivated Aileu market, Aileu Province

15

25 May 20 2005ET-20-1 Vigna umbellata

cultivated Aileu market, Aileu Province

15

26 May 20 2005ET-20-2 Vigna umbellata

cultivated Aileu market, Aileu Province

15

S08-43-44.0 933
E125-34-00.1
S08-43-44.0 933
E125-34-00.1
S08-43-44.0 933
E125-34-00.1
S08-43-44.0 933
E125-34-00.1
S08-48-06.4 1298
E125-34-55.1

27 May 20 2005ET-21

Vigna radiata var. wild
sublobata

10km NW of Maubisse, Ainaro
Province

16

28 May 20 2005ET-22

Glycine sp.

wild

10km NW of Maubisse, Ainaro
Province

16

S08-48-06.4 1298
E125-34-55.1

29 May 20 2005ET-23

Vigna umbellata

wild

5km S of Maubisse, Ainaro Prov- 17
ince

S08-53-31.9 1645
E125-35-33.2

30 May 21 2005ET-24

Vigna radiata var. wild
sublobata

5km S of Ainaro Province18

S09-02-14.6 516
E125-30-31.6

Vigna radiata var. wild
10km S of Ainaro Province
sublobata32 May 21 2005ET-26 Vigna umbellata cultivated 30km SE of Bobonaro, Cova
Lima Province
33 May 21 2005ET-27-1 Vigna umbellata cultivated 10km SE of Bobonaro, Cova
Lima Province

19

S09-03-42.8 330
E125-31-41.1
S09-07400
E125-25
S09-04-01.6 600
E125-22-59.7

34 May 21 2005ET-27-2 Vigna umbellata

21

S09-04-01.6 600
E125-22-59.7

35 May 21 2005ET-28

22

36

22

S09-01-40.5 755
E125-22-21.4
S09-01-40.5 755
E125-22-21.4
S09-01-40.5 755
E125-22-21.4
S09-01-40.5 755
E125-22-21.4
S08-37-41.2 630
E125-27-53.8

31 May 21 2005ET-25

37
38
39

cultivated 10km SE of Bobonaro, Cova
Lima Province

Vigna unguiculata cultivated 5km E of Bobonaro, Bobonaro
Province
May 21 2005ET-29-1 Vigna unguiculata cultivated 5km E of Bobonaro, Bobonaro
Province
May 21 2005ET-29-2 Vigna umbellata cultivated 5km E of Bobonaro, Bobonaro
Province
May 21 2005ET-29-3 Vigna umbellata cultivated 5km E of Bobonaro, Bobonaro
Province
15km SW of Dili, Dili Province
May 22 2005ET-30 Vigna umbellata wild

40 May 22 2005ET-31

Vignareflexo-pilosa

wild

20
21

22
22
23

5km S of Gleno, Ermera Province 24- 142 -

S08-44-28.2 800
E125-26-05.1

Habitat

Shading

DisturbancePopulation size

Growthstage

Soil

Seed Sample

Herba Rhizo-rium -bium

Remarks

road side
grasslandlightmedium sporadicallyflowering → silt
mature

yes

bulk

yes

yes

pod length 7 cm (by herbarium), 9 seeds per pod, true
black seed, a bit larger thantypical wild seed

gully
besidemountain
road

open

high

flowering → gravel
mature

yes

bulk

yes

no

pod length 5.5~6 cm, 14~15
seeds per pod, long brownhairs on mature pod, pods easily dettached from peduncle,
flower clean yellow, stem, leaf,
pod hairy, goat like to eat

marketharvested

yes

bulk

no

no

yellow with brown hilum seedmarketharvested

yes

bulk

no

no

marketharvested

yes

bulk

no

no

whitish yellow with pale brown
hilum seeddark red seedmarketharvested

yes

bulk

no

no

pale yellowish brown seedsporadicallygully near open
the top ofmountain
road

high

sporadicallyflowering → gravel
mature

yes

bulk

yes

yes

pod length 3~4 cm, 8~9 seeds
per pod, blackish with reticulation seed, very small seed,hilum slightly protruding,spreading growth habit, small
leafletgully near open
the top ofmountain
road

high

sporadicallyflowering → gravel
mature

yes

bulk

yes

no

pod length 2 cm, 5 seeds perpod, pale brown short hairs onmature podroad side
grassland

light

high

sporadicallyflowering → silt
mature

yes

bulk

yes

yes

road side
grasslandmedium medium several
plantsmaturegravel
(dry)

yes

bulk

yes

no

pod length 7~8 cm, 9~10 seeds
per pod, true black seed, flower
bright yellow, leaf hairy
pod length 5.5~6 cm, 13~14
seeds per pod, blackish withreticulation seed, hilum slightly
protruding

river side

medium medium one plant flowering

gravelyes

yes

no seednofromfarmerfromfarmerharvestedyesindiyesvidualbulk
noharvested

yes

bulk

no

no

from
farmerharvested

yes

bulk

no

no

from
farmerfromfarmerfromfarmerfromfarmerroad side
grassland

harvested

yes

bulk

no

no

3 seeds only, red seed, rather
round seed shapeplant mixed with corn, Nov./
Dec. → May, pale to dark red
seedplant mixed with corn, Nov./
Dec. → May, pale yellowish
brown seedshiny black seed

harvested

yes

bulk

no

no

shiny black seed

harvested

yes

bulk

no

no

pale to dark red seed

harvested

yes

bulk

no

no

pale yellowish brown seedmedium medium several
plantsfloweringyes

indiyesvidualyesroad side
wet slope

medium medium several
plantsmature

yes

bulk

yes

no

pod not yet matured, seeds
from herbarium specimen,
white mark on leaflet
pod length 7~8 cm, pod black,11~13 seeds per pod, balck
seed, hairless stem

clay
(wet)

- 143 -

Table 3. (continued).
No.

Coll.
Date

Coll.
No.

Species

StatusCollection Site

Site
No.

Latitude /Longitude

Altitude(m)

41 May 22 2005ET-32-1 Vigna umbellata

wild

10km S of Letefoho, Ermera
Province

25

S08-52-14.8 1147
E125-24-47.2

42 May 22 2005ET-32-2 Vigna umbellata

wild

10km S of Letefoho, Ermera
Province

25

S08-52-14.8 1147
E125-24-47.2

43 May 22 2005ET-33

Vigna trinervia

wild

15km S of Letefoho, Ermera
Province

26

S08-53-07.1 1400
E125-25-04.4

44 May 22 2005ET-34

Vigna umbellata

wild

15km S of Letefoho, Ermera
Province
May 22 2005ET-35-1 Phaseolus vulgaris cultivated 15km S of Letefoho, Ermera
Province
May 22 2005ET-35-2 Phaseolus vulgaris cultivated 15km S of Letefoho, Ermera
Province
May 22 2005ET-36-1 Vigna umbellata cultivated Atsabe coffee shop, Ermera
Province
May 22 2005ET-36-2 Vigna umbellata cultivated Atsabe coffee shop, Ermera
Province
cultivated Atsabe coffee shop, Ermera
May 22 2005ET-37-1 Glycine max
Province
cultivated Atsabe coffee shop, Ermera
May 22 2005ET-37-2 Glycine max
Province
May 22 2005ET-38 Vigna trinervia
wild
3km N of Letefoho, Ermera
Province

26

S08-53-07.1 1400
E125-25-04.4
S08-53-07.1 1400
E125-25-04.4
S08-53-07.1 1400
E125-25-04.4
S08-55-26.3 1225
E125-23-52.3
S08-55-26.3 1225
E125-23-52.3
S08-55-26.3 1225
E125-23-52.3
S08-55-26.3 1225
E125-23-52.3

45
46
47
48
49
50
51

- 144 -

26
26
27
27
27
27
28

S08-49-23.1 1310
E125-26-01.2

Habitat

Shading

DisturbancePopulation size

Growthstage

Soil

Seed Sample

Herba Rhizo-rium -bium

Remarks

road side
grasslandlightmedium several
plantsmaturegravel

yes

bulk

yes

no

pod length 7~7.5cm, pod
brown, 9~11 seeds per pod,pale brown with slightly black
mottled seedroad side
grasslandlightmedium several
plantsmaturegravel

yes

bulk

yes

no

road side
grasslandlightmedium several
plantsmature

dry

yes

bulk

yes

no

pod length 7~7.5cm, pod
brown, 9~11 seeds per pod,pale brown seed
pod length 5.5~6 cm, pod
brown with long hair,
12~13seeds per pod, blackishwith reticulation seedroad side
grasslandroad sideslope fieldroad sideslope fieldfromfarmerfromfarmerfromfarmerfromfarmerroad sideslope fieldlightgravel

yes

bulk

no

no

light

medium several
matureplantsnone
5 x 10 m mature

gravel

yes

bulk

no

no

pod length 7~7.5 cm, 9~10
seeds per pod, true black seedpurple seed

light

none

gravel

yes

bulk

no

no

white seed

harvested

yes

bulk

no

no

red seed

harvested

yes

bulk

no

no

pale yellowish brown seed

harvested

yes

bulk

no

no

harvested

yes

bulk

no

no

whitish yellow with brown
hilum seedblack with black hilum seed

yes

bulk

no

no

light

5 x 10 m mature

medium several
plantsmaturegravel- 145 -

pod length 5.5~6 cm, pod
brown with long hair, 11~12
seeds per pod

2005ET-3
Vigna unguiculata

2005ET-8
Vigna unguiculata

2005ET-6-1
Vigna umbellata

2005ET-10-3Vigna umbellata

2005ET-9
Vigna umbellata

2005ET-5
Vigna radiata

2005ET-1
Vigna radiata

2005ET-21
Vigna radiata

2005ET-33
Vigna trinervia

2005ET-31
Vigna reflexo-pilosa

2005ET-4
Phaseolus vulgaris

2005ET-6-2
Vigna umbellata

2005ET-18
Glycine max

2005ET-37
Glycine max- 147 -

2005ET-22
Glycine sp.