PhycologicalResearch..........................................................................................

Volume 64 • Issue 4 • October 2016..........................................................................................
ISSN 1322-0829..........................................................................................

The JapaneseSociety ofPhycology

Phycological Research
EDITOR-IN-CHIEF
Mitsunobu Kamiya
Faculty of Marine Bioscience, Fukui Prefectural University, Fukui, JapanASSOCIATE EDITORS
Jaanika Blomster
Dept. of Environmental
Sciences, University ofHelsinki, Finland

Niels Daugbjerg
Department of Biology,
University ofCopenhagen, Denmark

Daiske Honda
Department of Biology,
Faculty of Science andEngineering, Konan
University, Japan

Mitsunori Iwataki
Asian Natural
Environmental Science
Center, University ofTokyo, Japan

Regine Jahn
Botanic Garden and
Botanical Museum
Berlin-Dahlem, Freie
Universität Berlin,
Germany

Ulf Karsten
Institute of BiologicalSciences, University ofRostock, Germany

Christos Katsaros
Department of Botany,
Faculty of Biology,
University of Athens,
GreeceMasanobu Kawachi
Environmental Biology
Division, National
Institute forEnvironmental Studies,Japan

Jeong Ha Kim
Department ofBiological Sciences,
Sungkyunkwan
University, Korea

Tatyana A. Klochkova
Department of Biology,
Kongju National
University, Korea

Kazuhiro Kogame
Department of NaturalHistory Sciences,Hokkaido University

Showe-Mei Lin
Institute of Marine
Biology, National
Taiwan Ocean
University, Taiwan

Ichiro Mine
Graduate School ofKuroshio Science,
Kochi University, Japan

Navid Moheimani
School of Veterinaryand Life Sciences,
Murdoch University

Rei Narikawa
Department of BiologicalScience, Shizuoka
University, Japan

Wendy Nelson
National Institute of
Water & AtmosphericResearch & SBS,
University of Auckland,
New Zealand

Gregory N. Nishihara
Institute of East China
Sea Research,
Nagasaki University,
Japan

Tatsufumi Okino
Faculty of Environmental
Earth Science,
Hokkaido University,Japan

Hiroyuki Sekimoto
Department of Chemicaland Biological Sciences,
Faculty of Science,
Japan Women’sUniversity, Japan

John A. West
School of Botany,
University of Melbourne, Australia

Giuseppe C. Zuccarello
School of BiologicalSciences, Universityof Wellington,
New Zealand

EDITORIAL SECRETARY
Shinya Sato
Faculty of Marine Bioscience, Fukui Prefectural University, Fukui, JapanAIMS AND SCOPE Phycological Research is published by the JapaneseSociety of Phycology and complements the Japanese Journal of Phycology.
The Journal publishes international, peerreviewed research dealing withall aspects of phycology to facilitate the international exchange of results.
All articles are peer-reviewed by at least two researchers expert in thefield of the submitted paper. Phycological Research has been credited by
the International Association for Plant Taxonomy for the purpose ofregistration of new non-vascular plant names (including fossils). For
further information see http://bgbm3.bgbm.fu-berlin.de/iapt/registration/
journals/htm.
DISCLAIMER The Publisher, Japanese Society of Phycology and Editors
cannot be held responsible for errors or any consequences arising fromthe use of information contained in this journal; the views and opinions
expressed do not necessarily reflect those of the Publisher, JapaneseSociety of Phycology and Editors, neither does the publication ofadvertisements constitute any endorsement by the Publisher, JapaneseSociety of Phycology and Editors of the products advertised.

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Cover Primary segregative cell division in Struvea okamurae (Photos byA. Hasebe). The same apical portion of a cylindrical cell was photographedin order from left to right 72 min, 109 min, 126 min, 136 min, 148 min,294 min, and 492 min after the light period had begun. See Okuda et al.
in this issue for details.