Degree Programs in Life and Earth Sciences

Graduate School of Science and Technology
Degree Programs in Life and Earth Sciences
List of research fields （Master's Programs）
Master's Program in Biology
Field of Research
Systematics andEvolutionary Biology

Faculty

Detailed Description of Research Field

ISHIDA Ken-ichiro① Classification of micro-and macro-algae based on ultrastructure andmolecular phylogenetic analyses② The endosymbiotic acquisitions and evolution of plastids③ Searching for new useful algae for algal biomass research

HONDA Masanao① Taxonomy of reptiles based on morphological data
② Molecular phylogeny and biogeography of reptiles and birds③ Conservation genetics of amphibians and reptiles

WADA Hiroshi① Evo-Devo research of chordates② Comparative embryology of marine invertebrates, including bivalves andechinoderms

DEGAWA Yousuke① Natural history and biodiversity of the Kingdom Fungi② Taxonomy and phylogenetic studies of the basal lineage of Fungi(Zygomycota and Chytridiomycota)③ Fungal ecology focused on their interactions with other organisms andtheir life cycles

NAKANO Hiroaki① Natural history of placozoans, xenacoelomorphs, and echinoderms② Origins and evolution of deuterostomes and metazoans③ Diversity and evolution of marine invertebrates

NAKAYAMA Takeshi①

YAHATA Kensuke①

Classification of protists including microalgae based on
ultrastructural characters and molecular phylogenetic analyses② Searching for new useful algae for algal biomass research②③

EcologyComparative morphological studies on ovarian structure and mode ofoogenesis in arthropods
Comparative studies on structures for appendage autotomy inarthropods
Phylogenetic studies of myriapods based on comparative
morphological methods

HIROTA Mitsuru① Plant response to environmental changes, perspective from ecology② Ecosystem ecology focused on carbon cycling in terrestrial ecosystem

TANAKA Kenta① Evolutionary ecology focusing on ecological and genetic adaptive
mechanisms in wild Arabidopsis② Conservation ecology in mountains, grasslands and forests

TSUDA Yoshiaki① Population genetics/genomics and population demographic inference ofseveral species (e.g. trees, fishes, insects, mammals)② Ecosystem management and conservation using molecular ecology
approaches③ Impact of human activities on forest ecosystems and their history

TOQUENAGA Yukihiko① Experimental ecology with field and laboratory populations② Theoretical biology with mathematical models

OHASHI Kazuharu① Foraging behavior of pollinators with special reference to their cognitive
abilities② The evolution of floral phenotypes via ecological interactions with flower
visitors

1

Degree Programs in Life and Earth Sciences
Ecology

SATO Yukie
HARVEY Benjamin

AGOSTINI Sylvain

Plant Physiology andDevelopmental Biology

KIKUCHI Akira
SUZUKI Iwane

IWAI Hiroaki※ONO Michiyuki
SUZAKI Takuya① Behavioral ecology and evolutionary ecology in terrestrial arthropods② Geographic variation in behavior and ecology, and speciation① Marine community ecology, utilizing field surveys, aquarium experiments
and modelling.
② Understanding the effects of environmental change on coastal systems,
such as ocean acidification, global warming, and marine heatwaves① Marine ecophysiology, especially of scleractinian corals② Responses of marine organisms to climate change and anthropogenic
stressors① Study on diversity of environmental stress responses in higher plants② Study on expression of totipotency in higher plants① Photosynthetic acclimation and signal perception to environmental
stress② Production of useful metabolites by metabolic engineering in algae③ Application of quantum beams and nuclear resonance reaction forbiological breeding① Cell wall functions on the plant development and environmental
responses② Mechanisms of plant cell wall cross-linking① Molecular mechanism of photoperiodic induction of flowering② Studies on genetically modified and genome edited plants①②

MAEDA Yoshiaki①②③

MINODA Ayumi①

IRVING Louis John

① Effect of nutrient status on the host – parasite interaction

Studies on regulation of primary metabolism in algae as unicellularplant model systems② Studies on metal metabolism in photosynthetic organisms②

Animal Physiology andDevelopmental Biology

Molecular genetic studies on root nodule development during
legume-Rhizobium symbiosis
Studies on molecular mechanism of nitrogen nutrient response inplantsFunctional analyses of genomes and chromosomes in algae
Production of useful metabolites by metabolic engineering in algae
Digital transformation of algal research※KOBAYASHI Satoru

SASAKURA Yasunori

CHIBA Chikafumi★NIWA Ryusuke

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YAGUCHI Shunsuke

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Exploring the influence of abiotic factors on grass plant growthand competition
Common mechanisms regulating germline formation in animals
Genetic pathway regulating sex determination of germline in Drosophila
Mechanism regulating germline-stem-cell maintenance in Drosophila
Developmental mechanisms of animals
Metamorphosis of ascidians
Molecular biology of mimicry
Development and Evolution
Molecular mechanism of adult newt body-part regeneration
Molecular mechanisms of injury responses and injury-caused disorders inmammalian tissues
Induction and regulatory mechanisms of transdifferentiation
Molecular, cellular, and systemic mechanisms of the interaction between
insects and parasitoid wasps
Mechanisms of interorgan communication in the regulation ofdevelopment, stem cell proliferation, post-mating responses, and aging
Studies on molecular mechanisms of cancer cachexia using Drosophila asa model
Structural biology and chemical biology of insect growth control agents
Axis specification/formation of the sea urchin embryo
Development of the serotonergic neurons in the sea urchin embryo
Evolution of the anterior neuroectoderm

2Degree Programs in Life and Earth Sciences
Animal Physiology andDevelopmental Biology

Molecular and Cellular
Biology

OKAMOTO Naoki① Insect hormones and its regulation during development② Neuro-endocrine control of physiology and behavior in insects

SAKURAI Keisuke①

INABA Kazuo

CHIBA Tomoki

NAKANO Kentaro

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MIURA Kenji①②

ISHIKAWA Kaori

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SHIBA Kogiku① Studies on regulatory mechanism of sperm motility in marine organisms② Studies on regulatory mechanism of flagellar and ciliary motility inmarine organisms

TSURUTA Fuminori①②③

Genomics andBioinformatics

Electrophysiological studies on molecular mechanisms of signaltransduction in retinal neurons
Studies on non-visual photoreceptor cells in CNS
Structure, motility, and regulation of cilia and flagella
Diversity of cilia and eukaryotic evolution
Mechanism of fertilization and reproduction of marine organisms(protists, marine invertebrates and fishes)
Genetic analysis of selective protein degradation
Cell biology of the ubiquitin family
Knockout mice analysis of the ubiquitin system
Investigation of signal transduction controlling cytoskeleton andmembrane dynamics
Studies on the molecular diversity and evolution of cytoskeleton and its
regulatory systems
Molecular biology of the mechanisms of cell division using yeast andprotistSignaling mechanisms for abiotic stress response and sugar accumulationin plants
Production of useful proteins (pharmaceutical proteins, etc.) with plantbiotechnology
Production and evaluation of genome editing crops
Analyses of influences by mutations of mitochondrial DNA on cellularand physiological functions
Studies on the interactions between nuclear-coded genes andmitochondrial functions
Investigation of disease mechanisms of mitochondria-relateddiseases using model animals

Molecular basis of the developing brain regulated by microglia
Neuron-glia communication coordinating the brain environment in the
neonatal period
Mechanisms of the architecture of neural circuits influenced byenvironmental stresses
Plastid evolution via secondary endosymbioses
CO2 fixation in microalgae
Genome evolution in microalgae

HIRAKAWA Yoshihisa①②③

INAGAKI Yuji

① Molecular phylogeny of eukaryotes② Evaluation of the impact of lateral gene transfer to genome evolution③ Estimation of protein functions combining evolutionary parameters andtertiary structures

KUWAYAMA Hidekazu① Molecular analysis of biological soliton in multicellular movement② Functional analysis of a genetic disease in intracellular signaling pathway③ Memory of cell and spatio-temporal pattern recognition④ Analyses of a novel anti-tumor factor and the mechanism of caffeinedependent enhancement of anticancer drugs① Functional morphology of mammalian mito-chondria② Generation of mouse models for mitochondrial DNA-based diseases③ Therapeutics for mitochondrial DNA-based diseases

NAKADA Kazuto

3

Degree Programs in Life and Earth Sciences
Genomics andBioinformatics

Advanced Cellular
Biology

NAKAMURA Kouji① Biochemical and molecular biological analysis of many roles ofbacteriophages, ranging from fundamental biological research to theiruse medical and industrial biotechnologies② Novel physiological functions of non-coding small RNAs and theirmechanisms of regulation of gene expressions③ Identification of novel RNA-binding proteins and analysis of their
physiological functions

SAWAMURA Kyoichi

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HARADA Ryuhei① Computational Biophysics and Theoretical Biology② Molecular dynamics simulations for analyzing biological functions③ In silico drug design based on molecular simulations

NAKAYAMA Takuro① Symbiogenesis in unicellular organisms② Genomic research on evolution and diversity of protists

ITO Yuzuru①②

Advanced Molecular
Biology

Evolutionary Genetics
Genetic analysis of hybrid inviability and sterility in Drosophila
Genetic analysis of sexual isolation in DrosophilaInterspecific introgression in natural populations of Drosophila

Basic technology of the regenerative medicine using human iPS/somatic
stem cells
Drug discovery support technology based on regenerative medicine
technology
Understanding the infectious mechanism of parasitic protozoa
Study about the unusual organelle of parasitic protozoa
Basic research for the development of anti-parasitic drug

*NAGAMUNE Kisaburo(NIH, Tokyo)①②③*MARUYAMA
Kyonoshin(JIRCAS, Tsukuba)① Comparative genomic research in crop plants② Transcriptional and metabolic network research in crop plants③ Development of improved crop varieties*SHITARA Hiroshi(IGAKUKEN, Tokyo)① Molecular genetics of mitochondrial DNA in mammals② Generation of new mouse strains using transgenic technology③ Imaging techniques for visualizing mitochondria in mammals*MATSUI Hisanori(Takeda Pharmaceutical
Company, Ltd.
Fujisawa)①*YABUKI Akinori(JAMSTEC, Yokosuka)① Diversity and classification of microbial eukaryotes② Ecological function and role of microbial eukaryotes in ocean③ Diversity and functional evolution of RNA-editing and its related
phenomena in microbial eukaryotes④ Monitoring of the diversity of microbial eukaryotes on ocean
environmental changes① Extracellular electron transfer mechanism in electrogenic bacteria② Data-driven chemical biology research using a high-throughput
electrochemical system③ Development of resource recovery technology using the interaction
between materials and bacteria① Taxonomy, phylogenetics and biogeography of fungi, especiallymushrooms② Fungal diversity in the environment (soil, water and air)
③ Natural history of fungi based on museum specimens, DNA and othermetadata① Population ecology of woody plans② Structure and dynamics of forest ecosystem③ Growth management of forests*OKAMOTO Akihiro(National Institute forMaterials Science)*HOSAKA Kentaro(National Museum of
Nature and Science)*MASAKI Takashi( FFPRI, Tsukuba )②

Drug discovery research in the field of neuroscience,
endocrinology (particularly neuroendocrinology andreproductive endocrinology, and drug repurposing
Translational research for drug discovery

4Degree Programs in Life and Earth Sciences
Advanced Molecular
Biology*TAJIMA Yuko
(National Museum of
Nature and Science)① Life history on marine mammals② Comparative morphology on marine mammals③ Health assessments on marine mammals*CHIBA Youko(RIKEN, Wako)① Search for novel metabolisms in microorganisms.(Prokaryote)② Diversity of CO2 fixation and amino acid synthetic pathways③ Analysis of metabolic evolutionary by physical chemistry*FUJIWARA Sumire(AIST, Tsukuba)①②*MORIYA shigeharu(RIKEN, Yokohama)
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Basic studies of transcriptional regulation mechanisms inhigher plants
Research and development of useful plants by modifications oftranscription factors or genes
Functional analyses of transcription factors in higher plants
Research and development of biomass utilization process
Research and development of symbiosis based biotechnology
meta- and single-cell transcriptome analysis

The faculty member marked with ※ will be retired by March 31,2026.

Note: *Adjunct Professor of the Cooperative Graduate School★The faculty member marked with ★ is a dual-role faculty member and cannot be a primary supervisor.

5

Degree Programs in Life and Earth Sciences

Master's Program in Agro-Bioresources Science and Technology
Field of Research
Plant Breeding

Faculty
YOSHIOKA Yosuke

Detailed Description of Research Field①

Study on conversation and efficient utilization of genetic
resources
Genetic analysis of important traits in crops
Pollination biology for seed multiplication of crops
Development of digital phenotyping method②③④
Animal Science

Plant Genome
Sciences

ASANO Atsushi

SUGIMOTO Koichi
TAKAYAMA Mariko★ LOMBARDO Fabien
Claude Renaud①

Integrated physiology of homeostatic functions useful foranimal production②

Study on molecular and cellular basis for fertilization anddevelopment in model animal③

Development of reproductive and genomic biotechnologies forlivestock production①

Exploring molecular mechanism for fruit development in tomato
Identification of genes related to important breeding traits in crops
and horticultural plants by genome analysis.
Rapid and efficient development of new cultivars by genome editing
technology.
Identification and characterization of genes controlling levels offunctional materials by metabolic and genome analysis of largescale tomato mutant population.
Innovation of gene modification technology by modified
CRISPR/Cas9 system.

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Agro-biological Sciences Field

Olericulture andFloriculture

FUKUDA Naoya
KANG Seung Won
NONAKA Satoko①

Molecular and physiological dissections of useful traits
involved in agricultural production in vegetables andornamentals
Development of new genetic engineering technologies andnovel high quality varieties in vegetables and ornamentals
Development of information technologies for vegetables andornamentals production
Study of Sophisitication for transformation and genome
editing technology
Study for introducion of new traits into Cucubitaceae,
Solananceae, and Asteraceae via new breeding technology,
CRISPR/Cas9.
Studies on fruit tree physiologies under cultivation andeffects of environmental factors on the physiology
Study on mechanisms of fruit tree flower development andpostharvest physiology of the fruit

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Pomology andPostharvest
Physiology ofFruitsSUGAYA Sumiko
SEKOZAWA Yoshihiko

Crop Science

MATSUKURA Chiaki
WANG Ning①②①

Comparative studies on the efficient crop production systems
and its management
Establishment of sustainable crop production systems and its
assessment
Physiological and ecological researches for yield and qualityof crops
Physiological research on the mechanisms and control ofstress tolerance in crops
Analysis of gene function for critical agronomic traits in crops②③④⑤
Plant Parasitic
Mycology

OKANE Izumi① Systematics of plant parasitic fungi including symbiotic fungi,
particularly rust fungi, blue stain fungi, endophytes and mycorrhizal
fungi.
② Studies on ecology and physiology of these fungi and bacteria.
③ Functional analysis of genes associated with disease resistance inplant.

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Degree Programs in Life and Earth Sciences

Agro-biological Sciences Field

AppliedEntomology
andZoology

FURUKAWA Seiichi
KURAMITSU Kazumu

ForestEcotopology

KAMIJO Takashi
KAWADA Kiyokazu

Conservationof Regional
Resources

SEINO Tatsuyuki
TSUDA Yoshiaki

Environmental
Soil Chemistry

ASANO Maki

Biological
Systems
Regulation
Science

KUSANO Miyako
SHIBA Hiroshi

Epigenetics

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BUZAS Diana Mihaela①②③

Insect immune mechanisms against pathogens and parasitoids
Elucidation of strategies of parasitoids to survive in the host insect
species
Improvement of biological control of insect pests
Ecology and ethology of parasitoids④

Dynamics and function of forest ecosystem
Vegetation science and management
Conservation and restoration of arid and semi-arid
Ecosystem
Conservation of endangered species①②

Genetic diversity of forest tree species
Evolution and local adaptation of forest tree species

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Study on conservation of regional resources
Environmental chemistry of forest soils
Soil ecological studies on soil organic matter
Soil conservation under grassland in Eurasian steppe①②Development of analytical platforms to capture quantitative andqualitative changes of metabolite levels
Metabolic network biology using “omics” datasets③

Flavor analysis of important crops and vegetables④

Molecular mechanisms of epigenetic regulation in heterosis⑤②③

Molecular mechanisms of epigenetic regulation in sexual plant
reproduction
Molecular genetic analysis of the perennial life history in Arabidopsis
halleri gemmifera
Molecular ecology analysis of seasonal response in Wasabi japonica
Dissection of memory DNA function in overwintering in crucifers①

Agricultural Economics and Sociology Field

Plant Cell and
Synthetic
Biology

KINOSHITA Natsuko①②③

Plant and insect interaction
Production of high added value products in plants
Visualization of plant environmental response mechanisms

AgriculturalandBioresource
Economics

SHUTO Hisato①

Analysis of food industries with specific attention to issues of
productivity, R&D, scale economies, and economics oforganization
Economic analysis of agricultural and food security policies

Resource
Economics andDevelopment
Studies

SHUTO Hisato

Farm BusinessandAgribusiness
Management

UJIIE Kiyokazu

ForestEconomics

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Farm production and supply economics under the risk
Farm and agribusiness firm management and marketing
Food consumption and consumer policy①

Study on forest policy and economics②

International comparative study on forest management and forest
products market
International comparative study on production and marketing offorest products③
ForestSociology

KOHROKI Katsuhisa

International trade analysis of agricultural commodities andresources
Community development and resource management①②③

Historical study of forest management in Japan
Socioeconomic study on regional forest management in Japan
Comparative study on forestry organizations

7

Degree Programs in Life and Earth Sciences
Food
Resources
Engineering

Marcos Antonio das
NEVES

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Environmental
Colloid andInterfaceEngineering

KOBAYASHI Motoyoshi①

SUGIMOTO Takuya②③

Bio-resource
Process andSystemEngineering
Watershed
Conservation

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NASAHARA(NISHIDA) Kenlo
YAMAKAWA Yosuke①②③

Micro / nano-engineering for advanced bioresource processing
Microchannel technology for advanced food processing
Formulation of food micro /nano-dispersions and evaluation of
their gastrointestinal digestion
Effective utilization of food processing waste for value addition
Water and solute transportation in soil.
Salinity and erosion of soil
Water resource engineering in arid land, water quality control,
water treatment
Physics and chemistry of soil, soil pollution,colloid and interface
Resource and energy utilization using agricultural waste, biomass
and organic wastewater based on bio-resource recycling system
LCA, LCC, and simulator development for optimization design ofbio-resource conversion process and grasping of biomass potential
and its utilization
Mechanism of sediment production and transport
Sabo planning in harmony with natural environment
Environmental analysis through remote sensing

Bioresource Environment Engineering Field

Water
Resources
Management
Engineering

ISHII Atsushi
ASADA Yohei①②③

Development and management of irrigation systems
Water resources evaluation for development
Participatory irrigation management

Farmland
SystemEngineering

KOBAYASHI Motoyoshi①②

Farmland engineering, soil conservation engineering
Soil Physics, Environmental materials

Bioproduction
andMachinery

Tofael AHAMED①

Intelligent machinery and robotics for agricultural production
System analysis for bioenergy production and utilization
Real-time crop monitoring systems for site-specific management

Agri-Food
Process
Engineering

KITAMURA Yutaka

Chemistry of
Biomaterials

NAKAGAWA-IZUMI Akiko

Engineering of
Biomaterials※ENOMAE Toshiharu

YAMASHITA Yuji

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OBATAYA Eiichi※KAJIYAMA Mikio

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Removal of food hazard by wet milling
Milling of components related to health function by spray dry
Development of novel food by applying rice slurry
Chemistry for wood pulping and pulp bleaching
Chemical utilization of biomaterials and bio-refinery
Micro-analysis of wood components (lignin, tannin, carbohydrate
and others) and the related compounds
Creation of paper-based electronics and sensors
Conservation of aging library collection and paper cultural heritage
Forest and marine biomass composites for eco-friendly packaging
Synthesis of fluorine containing condensation polymers forcomposite materials
Chemical modification of poly (amino acid)s and poly saccharides
Property enhancement of biomaterials for high-performancemusical instruments
Investigation on the mechanical properties of wood with respect
to its fiber-reinforced cellular structure, and development oftechnology for their effective utilization
Physical and chemical characterization of natural adhesives such as
Japanese lacquer and chitosan, and development of technology for
their utilization

8

Degree Programs in Life and Earth Sciences
Biochemistryof Bioactive
Molecules

USUI Takeo①

SUNOHARA Yukari②③

FURUKAWA Jun

Genomic
Biology

MATSUYAMA Shigeru
TANIMOTO Keiji
ISHIDA Junji
KAKO Koichiro④

Identification of molecular targets of the bioactive compounds inmammalian and plant cells and their action mechanisms
Antioxidative responses to oxidative stresses
Semiochemicals mediating interactions among insects, plants andanimals
Mechanisms how to accumulate various metals in plants①

Modification and function of methyltransferases②

Aging regulated by methylation and metabolism (C. elegans & mouse)③

Genomic imprinting④

Gene expression mechanism for homeostasis①

Analysis of the structure-function relationships of proteins involved insignal transduction and transcription regulation
Analysis of the chromophore-protein interactions of chromoprotein
antitumor antibiotics
Protein engineering based on detailed structural information on
functional proteins

DAITOKU Hiroaki
Structural
Biochemistry

TANAKA Toshiyuki②③
Molecular
Microbial
Bioengineering

KOBAYASHI Michihiko
HASHIMOTO Yoshiteru①②③

Applied Biochemistry Field④⑤
Bioreaction
Engineering

ICHIKAWA Sosaku①

Application of polymolecular aggregates for bioprocesses②

Production of useful materials by enzymes and microorganisms③

Development of tools for selective protein conjugation

TOYOFUKU Masanori

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YAWATA Yutaka
AOYAGI Hideki

Interdisciplinary studies for practical use of cytochrome P450s
Bacterial cell- cell communication and biofilm formation
Microfluidics for analysis of bacterial communities
Biophysical analysis of biofilm formation
Bacterial interactions through membrane vesicles
Molecular microbiology of environmental bacteria and theirapplications①

Development of cultivation system for cell and protoplast withnovel functional activities and their biotechnological application
Analysis of naturally-occurring microbial symbiotic association,
construction of artificial symbiotic system and their application forvarious bioprocesses
Cell cultivation engineering and development of novel bioreactors

HIRAKAWA HidehikoAppliedMicrobiology

NOMURA Nobuhiko
UTADA, Andrew S.

Cell Cultivation
Engineering②③

Biomimetic
Chemistry

Screening of new metabolism of natural and unnatural compounds,
and functional analysis of their physiological functions
Metabolic engineering and screening/analysis/design/remodeling of
useful enzymes and genes
Functional analysis of enzymes involved in cleavage and synthesis of a
C-N bond and their molecular evolution
Development of super biological catalysts with novel functions of
microorganisms and their enzymes
Functional analysis of gene promoters and their application to the
production of useful compounds.

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Development of cultivation system for uncultured microbes(microbial dark matter), animal cells, and plant cells and theirbiotechnological application①②

Studies on complex of protein and polymer
Basic and applied technical studies on polyelectrolyte gel

9Degree Programs in Life and Earth Sciences
Applied Biochemistry Field

Molecular andDevelopmental Biology

KASHIWABARA Shin-ichi①②③

Biology for
Gene
Regulation

KIMURA Keiji

Ecological
Molecular
Microbiology

TAKAYA Naoki①②③

NAKAMURA Akira
YING Bei-Wen
TAKESHITA Norio

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Applied Biochemistry Field⑦⑧

Functional
Foods andFood
Chemistry

YOSHIDA Shigeki

Environmental
Plant
Biochemistry

YAMAJI Keiko

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Bioprocess
Engineering

NOMURA Nakao

Bioactive
NaturalProducts
Chemistry

SHIGEMORI Hideyuki①②

Biosystem Sciences Field

MIYAMAE Yusaku

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Plant
Physiology andChemistry

Industrial
Microbiology
andBioresource
Science

YAMADA Kosumi

NAKAJIMA-KAMBE
Toshiaki

Analysis for dynamics of mitotic chromosomes.
Analysis for function of condensin complex.
Analysis for novel function of the nucleolus.
Environmental response and morphogenesis of filamentous fungi
Enzymology and molecular biology of microbial enzymes
Bacterial metabolisms and communication
Development and application of host-vector system in Thermus
thermophilus
Study on microbial catabolic pathway of L-form sugars
Multilevel analyses and computational prediction of microbial growth
dynamics
Experimental evolution for investigating the microbial survival
strategies
Physiological functions of sulfur-containing amino acids and its
applications
Structure and function of bioactive compounds in food
Production of bioactive compounds by using bioconversion process
Development of industrial enzymes for food production
Effect of endophytic microbes on heavy-metal stress tolerance inplants
Effect of endophytic microbes on environmental stress tolerance inplants
Effect of endophytic microbes on radio Cs accumulation in plants
Development of sustainable agriculture, forestry and fisheries
industry using bioengineering technique③
Chemical
BiologyTranscriptional and translational regulation of genes during
gametogenesis
Functional roles of proteins involved in fertilization, egg activation,and early embryonic development
Development of reproductive and developmental technologies for
future life①

Elucidation of the molecular mechanisms of bioactive substances
involved in biological phenomena of plant (germination,
phototropism, gravitropism, senescence, etc.).
Search for bioactive compounds related to prevention of diseases(Alzheimer's disease, diabetes, osteoporosis, etc.) from edible andmedicinal plants.
Isolation and structure elucidation of new bioactive compounds(antimicrobial, antitumor, etc.) from unexplored microorganisms.
Small molecule control for cellular protein stability and function
Development of drug screening systems by focusing on the uniquecharacter of target receptor
Chemical biology on natural products②

Isolation and identification of plant-derived bioactive compounds
against abiotic and biotic stimuli
Evaluation of their biosynthetic pathway and mode of action③

Application of these compounds to precision crop farming
Isolation and screening of microorganisms with potential forbioproduction/biotransformation. (plastic degradation,
biotransformation of oil/fat-related biomass, and methane
conversion)

10Degree Programs in Life and Earth Sciences
Bioindustrial
Resources

Biosystem Sciences Field

Animal Cell
Biotechnology※WATANABE N. Kazuo
OGUCHI Taichi

ITO Yuzuru①

Bodiplomacy, conservation and sustainable use for genetic
resources, Biosafety on transgenic plants, Access for bioresources
and its appropriation②

Plant biotechnology, plant physiology on environmental response,
Environmental and health risk assessment of biotech plants,
Detection method biotech foods
Basic technology of the regenerative medicine using human stem
cells (Quality control, mass cultivation, differentiation)
Drug discovery support technology (organ cell differentiation for
drug discovery using knowledge from regenerative medicine,
microphysiological system)①②

BioEnvironmental
ControlEngineering

UTSUMI Motoo

Diversity and function analysis of marine and freshwater
microorganisms and its role in cycling of matter, Bio ecoengineering

Food System

KITAMURA Yutaka
KOKAWA Mito

Post-harvest technologies, Processing of functional foods,
Conversion and utilization of biomass and food waste, Nondestructive analysis of food quality using light

Biological andMaterial
CyclesYANG Yingnan

Plant Stress
Biology

AnimalFunctional
BiologyAgro-biological Sciences Field①

Photocatalytic technology, Solar light utilization system,②

Bioreactor, High efficiency conversion and effective utilization ofbioresources, Renewable energy*FUJITA Yasunari(Japan International
Res. Center for
Agricultural Sci.
(JIRCAS) )①

Molecular elucidation of stress tolerance mechanisms in plants
Development of environmental stress-tolerant crops*SAKUMOTO Ryosuke(Institute of Livestockand Grassland Science,
NARO)

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InsectFunctional
Regulation* TABATA Jun(Institute for Plant
Protection,NARO)①②

Climate Change
ImpactAssessment on
Vegetation*MATUI Tetsuya(Forestry and Forest
Products ResearchInstitute(FFPRI))①

Tropical
Forestry*MATSUI*TANI
Naoki
Tetsuya①(Forestry(Japan International
and Forest
Products
Research
Res. Centerfor
Institute(FFPRI))
Agricultural Sci. (JIRCAS))②*MINAMIKAWA Kazunori(Japan International
Res. Center for
Agricultural Sci.
(JIRCAS))①

International
Agriculture andForestry
Development*IIYAMA Miyuki(Japan International Res.
Center for Agricultural
Sci. (JIRCAS) )①

Regional Forest
Resource
Development* ISHIZAKI Ryoko(Forestry and Forest
Products ResearchInstitute(FFPRI))①②

Identification of social conflicts over forest resources
Study on how to lead rural development by utilizing forest
resources

Farming System* S AWA DA M a m o r u①②

Agricultural Workforce and Human Resource Development
Local Agricultural Support Systems to Revitalize Rural Communities

Environmental
Agronomy

②

③

②

②

Agricultural Economics andSociology Field②

Factors involved in the animal productive functions.
Study on animal reproductive biology, especially on the establishment ofpregnancy and its maintenance in ruminants.
Development of effective technique to improve reproductive
performance of domestic animals
Chemical ecology of insects and associated plants
Development of insect functional regulation techniques based on
chemical ecological studies
Relations between distributions of forest vegetation and climaticconditions
Impact assessment and adaptation planning of climate change on
forest ecosystem functions and ecosystem services
Improvement of tropical forestry using indigenous genetic resources inSoutheast Asian tropical forest
Reproductive biology in Southeast Asian tropical forest and its
application to sustainable forest management
Development and assessment of climate change mitigation andadaptation technologies in rice production
Observation and modeling of greenhouse gas emission and carbon andnutrient cycling in rice production
Trends and prospects of international agriculture research agendas on
global food systems.
Sustainable agricultural intensification of smallholder systems.

11

Degree Programs in Life and Earth Sciences
Rural
Environment
Improvement* MIYAMOTO Teruhito

Source Environment Engineering Field* YOSHIMOTO Shuhei(Institute for RuralEngineering,NARO)

①
②
③
①
②

Nano andMicro-scale
Food
Analysis* MANO Junichi

Sustainability ofBiomass
Resources*KOSUGI Akihiko(Japan International Res.
Center for Agricultural
Sci. (JIRCAS) )Regional
ForestResource
Development*YAMADA Tatsuhiko(Forestry and Forest
Products ResearchInstitute (FFPRI))①

Development of lignin based functional bio-materials②

Chemical conversion of cellulosic biomass for preparing usefulchemicals, liquid fuels and fuel additives Rapid analysis of
lignocellulosics to evaluate potential of forest biomass

AnimalBioresource
Engineering*INOUE Kimiko( RIKEN )①

Characterization of the germ cell genome using a nuclear transfer
techniqueAnalysis of the mechanisms for zygotic gene activation using a nuclear
transfer technique
Development of techniques for preservation of male germ cells
using microinsemination

Development of analytical methods for evaluating food quality
Development of food processing methods using biotechnology* GENKAWA Takuma(Institute of Food
Research,NARO)
Development of biomass utilization technology using microbial
function②③

Applied Biochemistry Field※

Irrigation and drainage management in farmland
Modeling, measurement and interpretation of mass and energy flow in
soil
Hydrological investigations and conservation of groundwater resourcesin rural areas

Evolutionary
Biology ofSymbiosis*FUKATSU Takema(AIST)

Molecular
Neurobiology*DOI Motomichi(AIST)①②①②③

Applied
BioengineeringofMicrobial
Ecosystems*TAMAKI Hideyuki(AIST)

Food
Molecular
Engineering*KOBORI Toshiro（Institute of Food
Research,NARO)

①
②
③

①
②

Biological function, evolution and origin of endosymbiotic
associations between insects and microorganisms
Molecular, physiological and regulating mechanisms underlying
sophisticated inter-organismal interactions in symbiosis,
parasitism, manipulation and sociality
Molecular analysis of nervous-system formation and maintenance
Development of screening systems for neuronal dysfunctions anddiseases
Development of live-cell imaging methods using fluorescent andluminescent techniques
Culturing the uncultured fastidious microorganisms in the environment
and exploring their novel biological functions
Omics-driven discovery of novel microbial and genetic resources
Ecophysiology and diversity of uncultured microorganisms in theenvironments (gut, plants, deep subsurface, etc.)
Screening and utilization of biomolecules for sensing food quality.
Analyses on structure-function relationship of advanced glycation end
products.

The faculty member marked with ※ will be retired by March 31,2026.

*Adjunct professor of the Cooperative Graduate School (not assigned an academic advisor’s position for research students[kenkyusei]).
★The faculty member marked with ★ is a dual-role faculty member and cannot be a primary supervisor.
(＊) Please contact the Chair of Doctoral Program in Life and Agricultural Sciences (e-mail: kamijo.takashi.fw#@#u.tsukuba.ac.jp）) in
regard to this research field. (*Replace “#@#” with “@”.)12Degree Programs in Life and Earth Sciences

Master's Program in Geosciences / Geoenvironmental Science Field
The Master’s Program in Geoscience provides fundamental knowledge and practical skills as a prerequisite both for further study in
doctoral programs and for professional life. This program comprises two major fields: Geoenvironmental Sciences and Earth
Evolution Sciences. The former is comprised of eight research fields (human geography, regional geography, geomorphology,hydrological science, atmospheric science, geographical information science, terrestrial water cycle system, and atmosphere-oceaninteraction system). The latter is comprised of seven research fields (paleobiological science, paleogeosphere science, geodynamics,
planetary resource geology, petrology, mineralogy, and earth historical analysis). The research fields of the faculty members arelisted in the table below.
Field of Research

Faculty

Detailed Description of Research Field

Human Geography

MATSUI Keisuke
jiji@geoenv.tsukuba.ac.jp

Cultural geography, Geography of tourism and religion, Theory ofcultural tourism

Regional
Geography

KUREHA Masaaki
mkureha@geoenv.tsukuba.ac.jp

Regional geography of Europe and Japan, Geography of tourism
Regional geography of Australia, Urban geography, GIS

TSUTSUMI Jun
jtsu@geoenv.tsukuba.ac.jp
Geomorphology

Hydrological
Science

Atmospheric
Science

Geographical
Information
Science

IKEDA Atsushi
aikeda@geoenv.tsukuba.ac.jp

Cold region geomorphology, Permafrost monitoring,
Mountain environments

HATTANJI Tsuyoshi
hattan@geoenv.tsukuba.ac.jp

Hydrogeomorphology, Landslides, Rock weathering, Karst
geomorphology

SEKIGUCHI Tomohiro
sekiguchi@ied.tsukuba.ac.jp

Sedimentary processes, Bedform, Experiment

ASANUMA Jun
asanuma@ied.tsukuba.ac.jp

Hydrometeorology, Land-vegetation-atmosphere System,
Atmospheric Turbulence

TSUJIMURA Makimktsuji@geoenv.tsukuba.ac.jp

Groundwater hydrology, Groundwater and surface waterinteraction, Water governance in watershed

YAMANAKA Tsutomu
tyam@geoenv.tsukuba.ac.jp

Water and material cycle, Isotopic tracer, Eco-hydro-meteorology

UEDA Hiroaki
hueda.hiroaki.gm@u.tsukuba.ac.jp

Atmosphere-ocean-land interaction involved in the climate
systemUENO Kenichiueno.kenichi.fw@un.tsukuba.ac.jp

Land-atmosphere interaction and precipitation system, Mountainweather and snow cover variations＊KUSAKA Hiroyuki
kusaka@ccs.tsukuba.ac.jp

Urban climatology, Mountain meteorology, Applied meteorology(wind energy prediction, biometeorology)

Matsushita Bunkei
matsushita.bunkei.gn@u.tsukuba.ac.
jp

Remote Sensing, GIS, Global Environment, Water Quality of Lakes

MORIMOTO Takehiro
tmrmt@geoenv.tsukuba.ac.jp
Analysis ofEnvironmental
Dynamics

Agricultural and Rural geography, Sustainability of agriculture
and rural area, GIS

ONDA Yuichi
onda@geoenv.tsukuba.ac.jp

Transfer of radionuclides in Environment, Hydro-geomorphology,
Forest hydrology

TSUMUNE Daisuke
tsumune.daisuke.gw@u.tsukuba.
ac.jp

Oceanic material cycle, simulation of oceanic radioactivity
dynamics.

KATO Hiroaki
kato.hiroaki.ka@u.tsukuba.ac.jp

Forest hydrology, Soil erosion, Environmental radioactivity

13

Degree Programs in Life and Earth Sciences
Water-related
Disaster Science

OceanAtmosphere
Interaction System

IIZUKA Satoshi
iizuka@bosai.go.jp

Atmosphere-ocean interaction ,Meteorological disaster,
Extreme event

SHIMOKAWA Shinyasimokawa@bosai.go.jp

Physical oceanography, Coastal disasters, Marine ecosystem

SHUSSE Yukari
shusse@bosai.go.jp

Clouds and precipitation meteorology, Rader meteorology

ISHII Masayoshi
maish@mri-jma.go.jp

Oceanography, Atmosphere-Ocean Interactions, Climate Variations
Atmospheric Chemistry, Aerosol-Cloud-Radiation Interactions

KAJINO Mizuo
kajino@mri-jma.go.jp＊ The faculty member marked with (*) can be a supervisor for students in the atmospheric science field, as well as the spatial information
science field.

14

Degree Programs in Life and Earth Sciences

Master's Program in Geosciences / Earth Evolution Science Field
Paleobiological
Science

AGEMATSU Sachikoagematsu@geol.tsukuba.ac.jp

Conodont, Graptolite, Tentaculite, Paleozoic historicalgeology of Southeast Asia
Vertebrate paleontology and paleoecology

Paleogeosphere
Science

TANAKA Kohei
koheitanaka@geol.tsukuba.ac.jp
KAMATA Yoshihito
yoshi_kamata@geol.tsukuba.ac.jp
FUJINO Shigehiro
shige-fujino@geol.tsukuba.ac.jp

Sedimentology and stratigraphy, Geological records of tsunamis
in Japan and Asian countries

YAGI Yuji
yagi-y@geol.tsukuba.ac.jp

Earthquake rupture process and seismicity

UJIIE Kohtaro
kujiie@geol.tsukuba.ac.jp

Structural geology and tectonics

OKUWAKI Ryo
rokuwaki@geol.tsukuba.ac.jp

Seismic source processes of earthquakes and nonearthquakes
Petrology of metamorphic rocks, Collisional orogeny,
Gondwana

Geodynamics

Petrology

Planetary
Resource
Geology

TSUNOGAE Toshiakitsunogae@geol.tsukuba.ac.jp

Geological evolution of Southeast Asia

IKEHATA Kei
ikkei@geol.tsukuba.ac.jp

Volcanology, Geochemistry

MARUOKA Teruyuki
maruoka.teruyuki.fu@u.tsukuba.ac.jp

Isotope geology, Geochemistry

FUJISAKI Wataru

History of life on earth, Tectonics

wataru-fujisaki@geol.tsukuba.ac.jp

Mineralogy

Earth Historical
Analysis

KUROSAWA Masanori
kurosawa@geol.tsukuba.ac.jp

Mineralogy, Fluid inclusion analysis

KYONO Atsushi
kyono@geol.tsukuba.ac.jp

Mineralogy, Crystallography, Mineral physics

KOHNO Naoki
kohno@kahaku.go.jp

Paleobiology of Cenozoic animals (especially for aquatic
animals)

SHIGETA Yasunari
shigeta@kahaku.go.jp

Paleobiology of cephalopoda

TSUTSUMI Yukiyasu
ytsutsu@kahaku.go.jp

Geochronology

15Degree Programs in Life and Earth Sciences

Master’s Program in Environmental Sciences
Facultyhttps://www.envr.tsukuba.ac.jp/eng/
Detailed Description of Research Field

TSUJIMURA Maki

Age dating of groundwater/ spring water using CFCs/ tritium, Hydrogeological processes
by using the isotopes, Rainfall-runoff processes in mountainous catchment

ASANUMA Jun

Surface hydrology, Evapotranspiration and precipitation, Precipitation sources,
Hydrological cycle, flood and flood mitigation

ONDA Yuichi

Transfer of radionuclides in Environment, Hydro-geomorphology, Forest hydrology

KATO Hiroaki

Forest hydrology, Soil erosion, Environmental radioactivity★SAKAGUCHI Aya

Applications of natural/artificial radionuclides as tracers for environmental dynamics

KAMAE Yoichi

Energy balance among atmosphere-ocean-land system

YAMAJI Keiko

Chemical Interaction between Plants and Microorganisms in the Rhizosphere understress environments

SUNOHARA Yukari

Action mechanisms of bioactive substances that regulate plant growth or biological functionsSUZUKI Iwane

Photosynthetic mechanism of microalgae, Microalgal biomass and carbon-nitrogenmetabolism

MAEDA Yoshiaki

Molecular biology, Genome science, Biotechnology, Production of useful compounds andenvironmental cleanup with microalgae

NOMURA Nobuhiko

Bacterial cell-cell communication and bacterial biofilm

TOYOFUKU Masanori

Microbiology

LEI Zhongfang

Biological waste and wastewater treatment, Biogranulation, Resource and energy
recovery

YUAN Tian

Anaerobic digestion for waste and wastewater treatment, Toxicity assessment andremediation of environmental pollutants

UTSUMI Motoo

Aquatic Biogeochemistry and Engineering※ENOMAE Toshiharu

Environmental Materials Science

ISHII Astushi

Development and management of irrigation systems, Water resources evaluation fordevelopment, Participatory irrigation management

KAMIJO Takashi

Vegetation dynamics on volcano and revegetation of volcanically devastated sites

KAWADA Kiyokazu

Conservation and restoration of ecosystems

YOKOI Tomoyuki

Insect ecology, Behavior and life history of bees,
Pollination service and conservation of flower-visiting insects

16

Degree Programs in Life and Earth Sciences
HIROTA Mitsuru

OMORI Yuko

Ecosystem Ecology, Plant Physiological Ecology, Carbon cycle and greenhouse gases(GHGs) dynamics in terrestrial ecosystem, Response to environmental change in alpine
ecosystem: species, community and ecosystem components
Marine Biogeochemistry, Oceanic carbon cycle and sea-air interaction

MURAKAMI Akinobu

Landscape planning, Urban and rural planning

YAMAMOTO SachikoArchitectural planning, Regional planning

YABAR Helmut

Integrated waste management systems: policy and planning; environmental impact
assessment; GIS for environmental management: applications in flood analysis, air
pollution analysis, waste and wastewater management, renewable energy potential
Environmental economics, Environmental policy, Comprehensive evaluation ofenvironmental policy and technology, Socio-environmental system simulation

MIZUNOYA Takeshi

KAIDA Naoko

Environmental psychology, environmental economics, pro-environmental behavior,environmental decision-making

NASAHARA Kenlo

Environmental monitoring and disaster prevention using satellite remote sensing

KUSAKA Hiroyuki

Urban climatology, Mountain meteorology, Applied meteorology (wind energy
prediction, biometeorology)

MATSUSHITA Bunkei

Remote Sensing, Geo-ecology, Modeling

KOBAYASHI Motoyoshi

Environmental and Colloidal Engineering、Aggregation and Dispersion of Colloids,
Electrokinetics

SUGIMOTO Takuya

Colloid Transport Phenomena

YAMASHITA Yuji

Colloid facilitated Transportation.
Colloidal Aspects of Humic Substances.

ASADA Yohei

Rural environmental engineering/Planning, Agricultural hydraulics/hydraulics※KAJIYAMA Mikio

Synthetic study on material sciences, Synthesis and properties of hybrid polymers

MATSUI Kenichi

Environmental dispute resolution and diplomacy; rural development and sustainability;
environmental/water ethics and law; environmental and agricultural policies forsustainability; environmental disaster policies
Mechanisms behind functional food resources for potential applications in food andcosmetics

ISODA Hiroko

MIYAMAE Yusaku

TAKAHASHI Shinya

Chemical control of cellular protein stability and its biological function
Screening and mechanism analysis of bioactive substances that modulate intracellular
metabolism
Risk sciences of radiation and chemicals, Plant molecular biology/Plant physiology,
Environmental impact assessment

Farhana FERDOUSI

Bioinformatics, Omics Research, Clinical trial, Epidemiology

ASANO Maki

Soil Science

UCHIDA Taro

Policy and planning of natural disaster prevention, Sediment disaster mitigation,
Watershed management

17

Degree Programs in Life and Earth Sciences
YAMAKAWA Yosuke

Forest science, Risk assessment and mitigation for natural disasters

TAKAMI Akinori[National Institute forEnvironmental Studies]
SUGATA Seiji[National Institute forEnvironmental Studies]
NAGASHIMA Tatsuya[National Institute forEnvironmental Studies]

Observation and analysis of air pollution including PM2.5 in East Asia and study of its
health and climate impact
Simulation and analysis of regional air pollutants and related analyses including
observation and meteorology
Studies on Asian air pollution and its effects using chemical transport model※ The faculty member marked with ※ will retire in March 31,2026.

★The faculty member marked with ★ is a dual-role faculty member and cannot be a primary supervisor.

18

Degree Programs in Life and Earth Sciences

Master's Program in Mountain Studies
Faculty

http://mountain-studies.tsukuba.ac.jp/en/toppage/

Detailed Description of Research Field

KUREHA Masaaki

Geography of Tourism

MATSUI Keisuke

Human Geography

IKEDA Atsushi

Geomorphology

UENO Kenichi

Atmospheric Science

YAMANAKA Tsutomu

Hydrologic Science

HATTANJI Tsuyoshi

Geomorphology

YAGI Yuji

Seismology

OKUWAKI Ryo

Seismology, Earthquake and non-earthquake source processes, Seismic arrayprocessing, Environmental seismology

KAMATA Yoshihito

Paleogeosphere Science, Accretionary Geology, Micro-biostratigraphy

NAKAYAMA Takeshi

Plant Systematic Taxonomy

ISHIDA Kenichiro

Plant and Protist Phylogeny and Systematics

DEGAWA Yousuke

Mycology, Plant Systematic Taxonomy

TANAKA Kenta

Population Biology, Plant Reproductive Ecology

TOQUENAGA Yukihiko

Theoretical Ecology

OHASHI Kazuharu

Plant Evolutionary Ecology

SATO Yukie

Behavioral Ecology, Evolutionary Ecology※ENOMAE Toshiharu

Environmental Materials Science

KAMIJO Takashi

Plant Ecology

SEINO Tatsuyuki

Forest Ecology

KOHOROKI Katsuhisa

Forest Resource Sociology

OBATAYA Eiichi

Wood Materials Engineering

NAKAGAWA-IZUMI Akiko

Wood Science

TSUDA Yoshiaki

Molecular Ecology, Population Genetics

KAWADA Kiyokazu

Plant Ecology

YAMAKAWA Yosuke

Erosion Control Engineering, Forest Hydrology

TSUJIMURA Maki

Aquatic Environmental Science

HIROTA Mitsuru

Ecosystem Ecology

MATSUI Kenichi

Environmental Policy

YOKOI Tomoyuki

Insect Ecology, Behavioral Ecology, Conservation Ecology

YAHATA Kensuke

Arthropod Systematics and Comparative Morphology

MORIYA Shigeharu（RIKEN）

Biomass Utilization ,Biological Symbiosis, Microbial Ecology, Molecular Evolution

19

Degree Programs in Life and Earth Sciences
TANI Naoki（JIRCAS: Japan
International Research Center
for Agricultural Sciences）
MASAKI Takashi（Forest Research andTropical Forest Management, Molecular Ecology

Forest Ecology

Management Organization）
MATSUI Tetsuya（Forest Research andManagement Organization）
TANAKA Norio（ National Museum of Nature

Vegetation Science, Impact of Climate Change

Plant Phylogeny and Systematics, Aquatic Plants

Science）※ The faculty member marked with ※ will be retired by March 31,2026.

20