ˏ˻ˏ͹ˏ̶͠ˡ‫ݖܩ‬቉ድǵፀ 3 ‫ش‬ǵ2003 ॷ 11 ೐අਬȆକ̥͙̹ͣͺΐͺȆͺές΃‫׫‬ᨆ͞ˠ˽ɹॶᩃᨆɴȲȿʳ᳋ᰖɹᅊਖ਼‫ݭ‬ଘգᅎɭз຦ɹྫ֝༎֪
*ǵ߂ǵីǵѵǵে⏋* ೠǵ‫ؤ‬ǵᅓǵ᱗**࣬ǵᅒǵ౩ǵχ⏋
Habitat Use of Fishes and Fishing Activity
in Plain Area of Southern Laos
IWATA Akihisa,* OHNISHI Nobuhiro* and KIGUCHI Yuka**
Fish fauna, habitats of aquatic bodies, habitat use of fishes, and local people’s fishingactivities were researched in the Ban hiang River system, one branch of the Mekong River,southern Laos.
Fish fauna of this area comprised 10 orders, 31 families and about 158 species. Aquaticbodies were divided into two large water areas, : permanent water areas and : temporary
water areas, and habitats were classified into 14 types. About 56 species of fishes were
found that clarified relation between their ecology and habitat, and 12 habitat use patterns
were recognized. Fishes inhabiting paddy fields were noteworthy in that they were notspontaneous arrivals from permanent water areas, but particular species specific to thishabitat.
A total of 26 types of fishing gear were found in this area. Local fishing practices are
based on a sound indigenous knowledge of the ecology of fishes. There were some differences
in sex and age among fishing gear users related with safety, distance from village and so on.
The implications of expansion of irrigation and aquaculture toward aquatic resourcesmanagement system were discussed.

1ȅ͉ȁ̲ȁ͛ȁͅǵആ‫ˏ˻ˏ׫‬ɻ͐Ͳ˽ͻͲຩҌɴఌᨗɇʶ⏋ϗ࡚ɭᮒ࡚ȵȲʱɲɐៀհ๭ɌȦᕶးɹ̀͌͠ɹ
μɫз຦ȵᅊ༎Ɍɩȷɛ⏏ᮒ࡚ȵʢɛʰɐ߂ᩄɹອɻɄɹ‫ݖܩ‬ɴȲȿʳΥ๖ᅋ෌ɹ‫ॻݜ‬ᅊ෌ɫϴ ᨊ ߂ ࡝ ߂ ࡝ ᭏ ˏ ˻ ˏ͹ ˏ ̶ ͠ ˡ ‫ ݖ ܩ‬቉ ድ ቉ ድ ዀ⏋Graduate School of Asian and African Area Studies, Kyoto
University** ͍˱Ͳ͹˗ˠ̒̍⏋Mekong Watch Japan*51

ˏ˻ˏ͹ˏ̶͠ˡ‫ݖܩ‬቉ድǵፀ 3 ‫ش‬ȡʳዝсˁఌȭ⏋ɗʶɭ‫ؾ‬ಁɴօʱՏɇʶɛ޽ዛ޽෵ɲອ‫ݖ‬ᄩ‫ޅ‬ɻ⏋߃းອᏙɭȡȦʚɤɩտᰩɭɌɩɹອᅋ႒ᅊᅋɴჼ‫ܧ‬ɫʢ޽߂ɲᢨ჉ˁɌɩȦʳ⏏
ǵആ‫ˏ˻ˏ׫‬ɹʛɲʰɑ⏋ɗɹៀฐɴȲȦɩγᅠ೑ధɹ߂ໜाɫȡʳ͍˱Ͳໜɻ̜̍̾̒ɴ
྇༑ˁеᒞɌɩᮖ‫׫‬ˁάʱ⏋ɗɹ༑‫ݖ‬ɴμ‫͞⏋ͻ͇Ͳ͑͊⏋ܕ‬ˠ˽⏋̊˓⏋ˡͲ̓˻ˏ⏋̜̠̾͌ɹ៟ 6 ˡ‫ܕ‬ˁ‫ي‬ʛɲȵʰ೎৿ɴ̜̠̾͌ɹ͍˱Ͳ̊̚͡ɴɗɹໜ‫ؤ‬ˁᬖȼ⏏যᬏᤂᮐ4,620 Ρ⏋༑‫ݖ‬ᯆዩᏝ 795,500 km2 ɹμɴᏝ 6,500 ΨϺɹϺǾȵಬʰɌɩȦʳ␩஗ 1996; ̠̊Ͳ
2002␭⏏
ǵ͍˱Ͳໜɹ༑‫ݖ‬ɴ‫ي‬ʚʶʳ 6 ˡ‫ܕ‬ɹȩɠ⏋͞ˠ˽ɻɗɹ‫ܤܕ‬ɹᏝ 90⎾ȵ͍˱Ͳໜɹᮃອ‫ݖ‬ɴ‫ي‬ʚʶɩȦʳ⏏Ʉɹ‫ܕ‬ɻ˓Ͳ̝˹̠‫ץ‬ऄԒ᭜ᨆɹ‫׌‬ആɴеᒞɌ⏋ɗɹ‫ޅܕ‬ˁμ‫̠̜̾⏋ܕ‬͌⏋͊͑Ͳ͇ͻ⏋̊˓⏋ˡͲ̓˻ˏɴஞɌ⏋ᯆዩ 236,800 km2 [Claridge et al. 1997]⏋1995 ॷɴȲȿʳϺ‫ؤ‬ɻ 450 ΨϺɫȡʳ⏏Ʉɹ‫ܕ‬ɹ 1998 ॷɴȲȿʳອᅋ෌ᅊᅋᩄɻ 42,800 t ɫȡʱ⏋ᕶᐌრɻෑʠɩᲞȦ⏏2000 ॷɫɹ͞ˠ˽‫ܕ‬຦ 1 Ϻȡɛʱɹॷᬚ᳋ᰖ༥ᢹᩄɻᏝ 8-10 kg ɫȡʳ[Modadugu et al. 2000]⏏ອᅋ႒ɹգᅎ঎ȵᲞȦɄɭɫ೑‫ؿ‬ɲ౗ೣɫ 1998 ॷ⏋‫ܕ‬຦ 1 Ϻȡɛʱɹॷᬚອᅋ႒њᐌᩄɻᏝ 6.4 kg ɫȡʱ⏃␩ອᅋং᳋కᨆНᅙᠩ֪‫ك‬Օഓవ 2001␭ʭʱ፤Տ⏆⏋͞ˠ˽ɴȲȦɩອᅋᣂ྇ȵෑʠɩᩂឹɲ৲ւˁഗɛɌɩȦʳɄɭȵࢅ౫ɴჿៗɫȷʳ⏏͞ˠ˽‫ܕ‬຦ɹອᅋ႒ɴࢪɐʳћࡔ঎ɻɄɹʭȩɴᲞȦȵ⏋᳋ˁೳషɭɌɛᇍᨥᰩ‫˪̚ͻ̮ڃ‬ɻ႕ɴᩂឹ
ɫ⏋Shoemarker et al. [2002] ɻȋ͞ˠ˽ϺɹᎦɴɦȽፀϠɹਖᮛᰩష‫ڃ‬ȍɭᝠჼɌɩȦʳʓɰɫȡʳ⏏৳ʰɻ͞ˠ˽‫׫‬ᨆɹ̫Ͳ̶ˎ˓ा༑‫ݖ‬ɴȡʳ೴ˁёɴ᏷ЂɌɩȲʱ⏋Ʉɹᠶయȴʰ‫׬‬Ꮴ៟፤ɐʳɭ⏋Ʉɹ‫ݖܩ‬ɫɻ 1 ॷ 1 Ϻȡɛʱ 6.2-9.4 kg ɹ̮ͻ̚˪ˁᰩʏɩȦʳɄɭɴɲʳ⏏̮ͻ̚˪ɹೳషɻз຦ɹᅊ༎‫ܞ‬ɫ஛ʶɛ˱˓ᰖ⏋̝˻͚˗ᰖˁ‫ي‬ʞμ͹ࢸ০ɹ᳋ɫȡʳȵ⏋ɗʶʰɻʓɭ˃ɰɹ‫غݭ‬ອᅒʦɗɹٞᦇɫ஛ᮃɇʶʳʢɹɫȡʳ⏏
ǵΥ౅⏋ອᅋᣂ྇ᅊᅋᩄɴ޽߂ɲ৯ᯱˁ‫ؖ‬ʖɐ᰸ຂϟ෌ɻ͞ˠ˽ɴȲȦɩʢɇʚɊʚɲ̺ͦ˻˙˪̜ȵ⏋ςɴ‫޺ܕ‬ลᬠςࢷɹ০ɫࣙᬖɌɩȦʳ [Modadugu et al. 2000]⏏ЁɹɭɄʸ⏋˱˓ʦ̠˓̘͡ˑ̴͞ˏፈȵ᰸ຂɇʶɩȦʳɴɐȹɲȦȵ⏋Ё৿⏋ঃ፴ܑɴ᰸ຂ຾ˁᦶଁɐʳɄɭɴɲʶɽ⏋ჼ‫ܧ‬ആ‫ˏ˻ˏ׫‬໤ࣳɹ‫ݖܩع‬ɫሾʰʶʳ˛̲᰸ຂɹʭȩɴගകᣂ྇ɴ޽߂ɲ৯ᯱˁ
‫ؖ‬ʖɐɽȴʱɫɲȼ⏋᰸ຂ᳋ɹዛᗈȵᩃ޺ɴఔ༑ɇʶʳɄɭɭȡȦʚɤɩ⏋͍˱ͲອᏙɹອ‫ݖ‬ᅊ઴Ꮩɴᩂ߂ɲ৯ᯱˁήȭ⏋‫ܧ‬ഀອᅋᣂ྇ȵᣒᇕ͹ᩄᇕɴ߂ȷȼ޴ࢅɐʳɄɭȵ‫׶‬ઉɇʶʳ⏏
ǵອᅋᣂ྇ɹ୫ᐚᇕգᅎˁᓒȭʳȩȭɫ⏋ʚɑࢪᢒᅊ႒ȵɰɄɫᅋ‫׺‬ɌɩᅊᔒɌ⏋ɰɹʭȩɲ‫ݭ‬ଘɫᅊ༎‫ر‬ˁԃȩɐʳɹȴɭȦɤɛ‫ݜ‬ኄ੻‫ݬ‬ȵέ‫خ‬๕ɫȡʳ⏏ᦏॷɴȲȿʳ͞ˠ˽ɹ᳋ᰖ
ɹՕᰖ࡝ᇕ቉ድɭɌɩɻ⏋Taki [1974; 1978], Roberts [1993], Roberts and Warren [1994] Ȳʭʃ Baird[1999] ȵȡɁʰʶʳ⏏ɌȴɌ⏋մ 2 ᓔɻ͡ˏͲ̺̫͞Ͳȴʰʹˑ˛Ͳ̍͑Ͳᦏ᧿ʚɫɹ‫׌‬ᨆɭ̮˪́ȴʰ˱ͻͲྡٞᦇɹ೎‫׫‬ᨆ‫ ৿⏋ݖܩ‬3 ᓔɻ˱ͻͲྡٞᦇɹ೎‫׫‬ᨆ‫ݖܩ‬ɹʛɭȦɤɛʭȩɴ⏋ᠭീȵ႕ࡴɹ‫ݖܩ‬ɴᭉࡴɇʶɩȦɛȵ⏋Kottelat [2001] ɴȲȦɩ՟ʠɩɄɹ‫ܕ‬ԃкɹ᳋52࣬ᅒ͹߂ី͹ೠ‫׫⏧ؤ‬ᨆ͞ˠ˽ɹॶᩃᨆɴȲȿʳ᳋ᰖɹᅊਖ਼‫ݭ‬ଘգᅎɭз຦ɹྫ֝༎֪

ᰖሂȵᐨᒧɇʶʳɴᕸɤɛ⏏‫ع‬ዛᰖɹᅊਖ਼‫ݭ‬ଘ᧧ିʦው֪ɴᬠɐʳᠭീɻ Taki [1978], Roberts[1993] Ȳʭʃ Roberts and Warren [1994] ɴʭʱɲɇʶɩȦʳʢɹɹ⏋ᅋ‫ݭ׺‬ଘʦІ͹዗᳋ɹᔒଁ
‫ݭ‬ଘɭᅊਖ਼‫ݭ‬ଘգᅎɴᬠɐʳ੻‫ݬ‬ɻӱՕɫȡʳɭɻៜȦᮑȦ⏏޽Ꮫ [1997] ʦቄພ͹̝͞͡ [1990]ɻആ‫ˏ˻ˏ׫‬ɹཁອ᳋ɹᅊ઴ᇕៗᠦˁ᝔ɤɩȦʳȵȦɑʶʢෙᠦᇕɲԒࢅɴᅥʚɤɩȦʳ⏏
Mekong River Commision [2001] ɻ͞ˠ˽ˁ‫ي‬ʠɩ⏋͍˱Ͳໜ༑‫ݖ‬ɴᅊਖ਼ɐʳςឹອᅋ᳋ዛ 48 ዛɹው֪ɭᅋ‫׺‬ɴɦȦɩ‫ݬ‬ٚɌɩȦʳȵ⏋ᅊਖ਼‫ݭ‬ଘɹ᠎Ᏻɲ៨ᥔɭɗʶʰɹգᅎɴɻៜ‫ؖ‬ɌɩȲʰɑ⏋᧧Տɇʶɛ᳋ዛʢз຦ɹᅊ෌Ϋᩂឹɫȡʳࢸ͹μ݀ɹዛᰖɻʓɭ˃ɰ‫ي‬ʚʶɩȦɲȦɹɫΫᦚɌɛʭȩɲ‫ݜ‬ኄᇕ੻‫ݬ‬ȵʓɭ˃ɰɲȦɭȦȩɹȵჼႥɫȡʳ⏏
ǵ͞ˠ˽ɫ‫ݖܩ‬቉ድˁ᝔ȩ᭬ɴອᅋ႒գᅎɭȦȩ᯵ᇿȵෑʠɩᩂឹɲ቉ድࢪᢒɭɲʳɄɭɻΫ៨Ɍɛဣȴʰʢᠶˁ৺ɛɲȦ⏏ɗɹμɫʢྫ֝༎֪ɻϺᬚɹ‫ۊ‬ʛɭອᅊᅊ႒ɹᅊ઴ɭˁᐄʉஞ
ဣɫȡʱ⏋ອᅋᣂ྇ȵᣒᇕ͹ᩄᇕɴ޴‫׋‬Ɍɛʱ⏋ʚɛ⏋ᅊ෌০઴ʦອᅋᣂ྇գᅎ˹˽̘͌ɴ޴
‫׋‬ȵᅊɍʶɽ⏋ɗʶʰɹ޴‫׋‬ȵྫ֝༎֪ɴሁஞᇕȴɦቁಁᬚɴ৯ᯱˁήȭʳ⏏ɇʰɴ᰸ຂϟ෌ɻອᅋᣂ྇ɴࢪɐʳዩෑᇕɲϺဤɹɻɛʰȷȴȿɭɌɩ⏋ɗɹ‫ۊ‬ဤȵਃഀɹᅊ෌০઴Ȳʭʃອᅋᣂ྇գᅎ˹˽̘͌ɴɰɹʭȩɲ৯ᯱˁʢɛʰɌɩʪȼɹȴ໶ᇿɇʶʳ⏏
ǵೣዣɻᘿᓔʰȵɄɹ 6 ॷᬚᠭീˁ᝔ɤɩȷɛ‫׫‬ᨆ͞ˠ˽ɹ̫Ͳ̯˛Ͳा༑‫ݖ‬ɹॶᩃᨆɴɰɹ
ʭȩɲ᳋ᰖȵᅊਖ਼ɌɩȦʳɹȴˁኍɐɭɭʢɴ⏋Ʉɹ‫ݖܩ‬ɴȲȿʳອ‫ݖ‬ᄩ‫ޅ‬ɹ‫ך‬Օˁ᝔Ȧ⏋Υಁᇕອ‫ݖ‬ˁμਕɭɌɩ⏋ᅋ‫ݭ׺‬ଘɭІ͹዗᳋ɹᔒଁ‫ݭ‬ଘɴ໶ᇿɌɲȵʰ⏋ɗʶəʶɹ᳋ዛȵȦȴɴ҅Ǿɹᅊਖ਼ᄩ‫ޅ‬ˁգᅎɌɩȦʳȴ౩ʰȴɴɐʳɄɭˁፀ 1 ɹᇿᇕɭɐʳ⏏ፀ 2 ɴɄɹ‫ݖܩ‬з຦ɹྫ֝༎֪ˁΫ៨ɹ੻‫ݬ‬ɴ‫ݜ‬ɧȦɩ៨ᥔɐʳ⏏ɇʰɴ⏋೎ᦏɹອ‫ݖ‬ᄩ‫ޅ‬ɹ޴‫׋‬ɭ᰸ຂϟ෌ɹಔ‫ؖ‬ȵЁ৿ɰɹʭȩɲ৯ᯱˁʢɛʰɌɩʪȼɹȴɴɦȦɩៜ‫ؖ‬ɐʳ⏏

2ȅ಺औ༹༷̤͍͢಺औ඾শ
ǵᠭീ‫ݖܩ‬ɴȲȿʳ᳋ᰖɹᅊਖ਼‫ݭ‬ଘգᅎɭྫ֝༎֪ɹᬠѥˁ౩ʰȴɴɐʳɛʠ⏋߂աɌɩГάɹʭȩɲᠭീˁ᝔ɤɛ⏏
(1) ᳋ᰖɹ஛ᮃᠭീ
ǵᘿᓔʰᕶʰȵହᐨ⏋ଝᐨ⏋լɌᐨፈɹྫԍɴʭɤɩອᅋ႒ˁ஛ᮃɌ⏋ฌೣˁсଁɐʳɭɭʢɴ‫ࡴܔ‬ሁ৿ɹฌೣˁԠሎ௣৯Ɍɛ⏏႕ɴଝᐨɻᐨᇿɹࢸɇȦʢɹʢьᅎɌɩІ͹዗᳋ɹ஛ᮃɴ
֗ʠɛ⏏‫ؾ‬ಁɴ‫ݖܩ‬з຦ȴʰ‫ؿ‬ዌ⏋գᅎ໬⏋ѝൔ⏋য়᠍ᅊ႒ɹᅊ઴⏋႕ɴᅋ‫ݭ׺‬ଘʦଁᬏɭው
֪ɴᬠʼʳᅊਖ਼‫ݭ‬ଘፈɹ੻‫ݬ‬ˁ਄ʳɛʠɴᓬ‫؜‬ʱᠭീˁ᝔ɤɛ⏏
(2) ྫ֝༎֪ɹჼ‫ݭ‬ᠭീǵᠭീ‫ݖܩ‬ɹз຦ȵອᅋᣂ྇ˁ஛஁ɌɩȦʳჼ‫ݭ‬ɴ᝔ȷ⏋஛஁႒⏋஛஁ᓔ⏋஛஁౅໬⏋஛஁‫ݭ‬ଘˁԠሎɫ៨᪼ɐʳɭɭʢɴ⏋஛஁ᓔȴʰ‫ؿ‬ዌ⏋գᅎ໬⏋ѝൔ⏋য়᠍ᅊ႒ɹᅊ઴⏋႕ɴᅋ‫ݭ׺‬ଘʦଁᬏɭው֪ɴᬠʼʳᅊਖ਼‫ݭ‬ଘፈɹ੻‫ݬ‬ˁ਄ʳɛʠɴᓬ‫؜‬ʱᠭീˁ᝔ɤɛ⏏
53ˏ˻ˏ͹ˏ̶͠ˡ‫ݖܩ‬቉ድǵፀ 3 ‫ش‬(3) ॑‫ݭ‬ᠭീǵᠭീ‫ݖܩ‬ɹ೴ɫ॑‫ݭ‬ɴ᝔ȷ⏋ɗɹ‫ݭ‬ଘɴι˃ɫȦʳອᅋᣂ྇ˁԠሎɫ៨᪼⏋‫ؾ‬ಁɴɗʶʰˁ଩ȩϺȴʰ‫ؿ‬ዌ⏋஛஁‫ݭ‬ଘ⏋஛஁౅໬⏋஛஁ᓔ⏋գᅎ໬⏋ѝൔ⏋য়᠍ᅊ႒ɹᅊ઴⏋႕ɴᅋ‫ݭ׺‬ଘʦଁᬏɭው֪ɴᬠʼʳᅊਖ਼‫ݭ‬ଘፈɹ੻‫ݬ‬ˁ਄ʳɛʠɴᓬ‫؜‬ʱᠭീˁ᝔ɤɛ⏏
(4) ᅊ෌ɴᬠɐʳ೴ɫɹᓬ‫؜‬ʱᠭീǵᠭീ‫ܩ‬ɹ೴ɴ᝔ȷ⏋ɗɹ೴ɹз຦ɴ‫ع‬ዛɹᅊ෌⏋႕ɴྫ֝ɭᦅ෌͹ക෌ɹᬠѥ⏋ອᅋᣂ྇፬
ჿɹ౅໬⏋᰸ຂϟ෌͹ອᅒ᰸ຂɹ೑ံ⏋ɗʶʰɴࢪɐʳз຦ɹᓒȭ౅ɲɰɹ੻‫ݬ‬ˁ਄ʳɛʠɴᓬ‫؜‬ʱᠭീˁ᝔ɤɛ⏏

ǵዛᰖɹ‫ࡴؾ‬ɻ௣৯ɇʶɛԠሎ⏋ʚɛɻฌೣɹᔀሗɫ៎࢘ɫȷʳ০ᣒˁʢɭɴ⏋Kottelat [2001]ˁؒᓒɴɌɩ᝔ʼʶɛ⏏ɛɞɌࡸкᰒ਎᫦ʦᲓൔɹ႕ਐˁᠭʏʳɛʠɹ᥉˛̒˪˽ᐻខᒞɫȡʳ̶̘̅̒˪˽ˁьᅎɌɲȿʶɽɲʰɲȦʭȩɲ᠎Ᏻɲ៎࢘ɻే៯ᇕ͹ಁᬚᇕɴέ‫خ‬ᔨɫȡɤɛɛʠ⏋Ё‫܌‬ɹ᳋ዛɴᬠɐʳ‫ࡴؾ‬ᐄഗɻȡȼʚɫಫࡴᇕɲʢɹɫȡʳ⏏ฌೣɻɐʏɩЁ৿ɹԊ‫ؾ‬቉ድɹɛʠອᅊᅊ႒ᣂ྇቉ድ́Ͳ̊ͻ (Living Aquatic Resources Research Center, LARReC) ɴѳ፬ɇʶɩȦʳ⏏ɲȲ⏋Ё‫܌‬ɹᠭീɻ͞ˠ˽‫ܕ‬ጓ߂࡝ (National University of Laos)⏋͞ˠ˽‫ܕ‬ጓᦅക቉ድଘ (National Agriculture and Forestry Research Institute, NAFRI) ɴȡʳອᅊᅊ႒ᣂ྇቉ድ́
Ͳ̊ͻɹ៰‫خ‬ɹʢɭɴ᝔ʼʶɛʢɹɫȡʳ⏏
ǵᠭീ‫ݖܩ‬ɹອ‫ݖ‬ᄩ‫ޅ‬ɹᅊਖ਼‫ݭ‬ଘȲʭʃໜा০઴ɹ‫ך‬Օɻ‫ܤ‬ೠ࡝Цᬠីఌᨆᑂ [1998] ɴȲȿʳ៨ᥔϟ᯵ˁ‫ݜ‬ኄ੻‫ݬ‬ɭɌɛ⏏ᇿȲʭʃዀɹ౗ೣ᠝ᝠ៨ɻΫᩃ͹‫ܮ‬ೣ␩1999␭ɴਃɤɛ⏏
ǵᠭീ‫⏋ܩ‬ᠭീ‫ܩ‬ɹᄩ‫ޅ‬ɹෙᅭ⏋ᠭീ౗ಁȲʭʃᠭീ౅໬ɻГάɹɭȲʱɫȡʳ⏃‫ ܒ‬1⏆⏏
(1) ̩̠̜͌͞Ͳ೴
ǵ͍˱ͲໜɹΥ߂ఌ༑ɫȡʳ̫Ͳ̯˛Ͳाά༑‫ݖ‬ɹ߂࣐ು‫ܩ‬ဣɴеᒞɐʳ೴ɫ⏋̍͑ͲͅͲ
ाɹ‫غ‬༑ဣȴʰᏝ 2 km ά༑ɴȡʳ⏏ధॷɴ 1 ঎ɻ༇ອɴʛʚʼʶ⏋ዝсʦɗɹЈɹс႒ɻ߂ଠ௖ˁ‫؝‬ȿʳ⏏ೣ༑ɹໜा০઴ɻဈɹᨆՕɹອཉȵཉȼ⏋ອᯆȵ໳ጓɛɲȦ Bc ݀ɫ⏋ɗʶɴɦʰɲʳໜᤄྲྀʦ໡ȵȡʳ⏏̫Ͳ̯˛ͲाȴʰͅͲ̺ɫளອɌϗ࡚ဌྰˁ᝔ȩ⏏ᮒ࡚ɫʢᅒɴ
ອɹɲȦ౗ȵᐚȼɭʦɻʱளອဌྰˁ᝔ȩ⏏1999 ॷ 7 ೐ 25 ౗⏃ᮒ࡚⏆ɴჼ‫ݭ‬ᠭീ⏋1999 ॷ 12
೐ 17-18 ౗⏃ϗ࡚⏆ɴ॑‫ݭ‬͹஛ᮃᠭീˁ᝔ɤɛ⏏2001 ॷ 3 ೐ 10-12 ౗⏃ϗ࡚⏆⏋6 ೐ 30 ౗Ȳʭʃ7 ೐ 1 ౗⏃ᮒ࡚⏆⏋2002 ॷ 8 ೐ 10-11 ౗⏃ᮒ࡚⏆ɴ೴຦ʍɹᓬ‫؜‬ʱᠭീ⏋ჼ‫ݭ‬ᠭീȲʭʃ஛ᮃᠭ
ീˁ᝔ɤɛ⏏
(2) ˰Ͳ˱˪೴Ȳʭʃɗɹᦏ᧿ǵ̫Ͳ̯˛Ͳा༑‫ݖ‬ɴȡʱ⏋Υᅱά༑ɫ‫׌‬үȴʰ‫غ‬༑ɐʳɄɹाɹϠ๖ఌ༑̍͑ͲͅͲाɹά༑ᨆɴеᒞɐʳ⏏ੈχᇕອ‫ݖ‬ɹ̍͑ͲͅͲाೣ༑ᨆɹʓȴ⏋ᮒ࡚ɴɻɄɹाɴɦɲȵʳ߂ࢸɹ໡ʦໜᤄྲྀ⏋ੈχᇕ͹ΥಁᇕᏳ༑⏋ະ࿭؅⏋༇ອകȵȡʳ⏏႕ɴ˽˓ɭ٧ɽʶʳ߂ៀฐɲ໡ɻ
54࣬ᅒ͹߂ី͹ೠ‫׫⏧ؤ‬ᨆ͞ˠ˽ɹॶᩃᨆɴȲȿʳ᳋ᰖɹᅊਖ਼‫ݭ‬ଘգᅎɭз຦ɹྫ֝༎֪

଎ 1 ȁ಺औ౷ത

ອᅋΫᩂឹɫȡʳ⏏ອᅒɴɻᮒອ߃ອᅒɹʓȴɴϗ࡚ဌྰᅒʢࡔ‫ܧ‬ɐʳ⏏ೣ༑ᨆɹໜा০઴ɻ
Bc ݀ɫȡʳ⏏1999 ॷ 7 ೐ 21 ౗⏋7 ೐ 26-27 ౗⏃ᮒ࡚⏆ɴ॑‫ݭ‬͹ჼ‫ݭ‬͹஛ᮃᠭീ⏋1999 ॷ 12 ೐
11-12 ౗⏃ϗ࡚⏆ɴ॑‫ݭ‬ᠭീˁ᝔ɤɛ⏏ʚɛ⏋˰Ͳ˱˪೴ᦏ᧿ɹࢸाɫ 7 ೐ 26 ౗⏃ᮒ࡚⏆⏋ჼ‫ݭ‬ᠭീˁ᝔ɤɛ⏏ɇʰɴ⏋2001 ॷ 3 ೐ 13-14 ౗⏃ϗ࡚⏆⏋6 ೐ 26-29 ౗Ȳʭʃ 7 ೐ 1-2 ౗⏃ᮒ࡚⏆ɴ͞˪˹ͻ೴⏋̠˱˗೴⏋˹ˏͲ̫Ͳ೴⏋͡Ͳ̜͌ा⏋̫̒˪ाɫჼ‫ݭ‬͹஛ᮃᠭീɌɛ⏏2002
ॷ 8 ೐ 12 ౗ɴɻ̠˱˗೴ɹ຦ᬚɹ᰸ຂ෌ᓔɴᓬ‫؜‬ʱᠭീˁ⏋8 ೐ 12-13 ౗⏃ᮒ࡚⏆ɴɻ̝Ͳ̝
˪͇˓೴ɭˤ̝ͻाɫ೴຦ʍɹᓬ‫؜‬ʱᠭീ⏋ჼ‫ݭ‬ᠭീȲʭʃ஛ᮃᠭീˁ᝔ɤɛ⏏8 ೐ 13 ౗ɭ
14 ౗ɴɻ˰Ͳ˱˪೴ɫ॑‫ݭ‬ᠭീˁ᝔ɤɛ⏏2001 ॷȲʭʃ 2002 ॷɴᠭീɌɛ‫ܩ‬ဣɻ˰Ͳ˱˪೴
ɹᦏ᧿ɫȡʳɛʠ⏋ᝠ 1 ɴɻ˰Ͳ˱˪೴ɭɌɩɗʶʰˁɭʱʚɭʠɛ⏏
(3) ˵̠͌˵˓೴
ǵ̨́ʭʱ‫ ᧑ܕ‬13 ‫ش‬ᐻɫᏝ 5 km ‫׌‬ആɴ᝔ɤɛ‫ܩ‬ဣɴȡʳ೴ɫ⏋ᮒອ߃ອᅒɫϗ࡚ဌྰɻ᝔ʼ
ʶɩȦɲȦ⏏ʚɛ⏋ᦏ᭰ɴੈ।ᇕອ‫ݖ‬ɻɲȦ⏏̊ͻ̩ˠाɴ༑ԂɐʳΥఌ༑ɹ೎Ϋ༑ᨆɴеᒞɐʳ⏏2000 ॷ 7 ೐ 18-19⏋26⏋28 ౗⏃ᮒ࡚⏆⏋2001 ॷ 6 ೐ 25 ౗⏃ᮒ࡚⏆⏋ε᭙‫ܩ‬ɹອᅒ⏋Ȳʭʃࢸाɫჼ‫ݭ‬͹஛ᮃᠭീˁ᝔ɤɛ⏏2002 ॷ 8 ೐ 15-16 ౗⏃ᮒ࡚⏆ɴ೴຦ʍɹᓬ‫؜‬ʱᠭീ⏋ჼ‫ݭ‬ᠭീˁ᝔ɤɛ⏏

55ˏ˻ˏ͹ˏ̶͠ˡ‫ݖܩ‬቉ድǵፀ 3 ‫ش‬(4) ̨Ͳˡ̼͌ͻ̜೴ǵ˵̠͌˵˓೴ˁ༑ʶʳ̊ͻ̩ˠाɹΥఌ༑ɹά༑ᨆɴȡɛʱ⏋˵̠͌˵˓೴ȴʰᏝ 1 km ɹ‫ܩ‬ဣɴеᒞɐʳ⏏ໜ᧑ᨆɴ๿ɇʶɛΥಁᇕອ‫ݖ‬ɫȡʳອɛʚʱɫ 2001 ॷ 3 ೐ 16 ౗⏃ϗ࡚⏆ɴ஛ᮃᠭീˁ᝔ɤɛ⏏
(5) ̊ͻ̩ˠाǵ‫ ᧑ܕ‬13 ‫ش‬ᐻɭɄɹाȵϯेɐʳ‫ܩ‬ဣɻϗ࡚ɫʢອɹ༲ʶɲȦੈχᇕອ‫ݖ‬ɫȡʳ⏏2001 ॷ 3
೐ 17 ౗⏃ϗ࡚⏆ɴ஛ᮃᠭീˁ᝔ɤɛ⏏ᠭീಁɻቆ໵ঈɹཌɭቆኊঈɹဈȵࡔ‫ܧ‬Ɍ⏋ອཉȵ༓ȼ⏋ອᯆȵ໳ጓɦ Bb ݀ɹໜा০઴ˁɌɩȦɛ⏏
(6) ̊͌͠ͻ೴ǵ̍͑ͲͅͲाɹ‫غ‬༑ဣȴʰᏝ 50 km Ϋ༑ɹ̫Ͳ̯˛Ͳाೣ༑ɹृࣳɴеᒞɐʳ⏏̫Ͳ̯˛Ͳाೣ༑ɹໜा০઴ɻဈɹᨆՕɹອཉȵཉȼ⏋ອᯆȵ໳ጓɛɲȦ Bc ݀ɫȡɤɛ⏏ྫ֝ɌɩȦʳ‫ݖܩ‬з຦ɴ 1999 ॷ 7 ೐ 22 ౗⏃ᮒ࡚⏆ɴჼ‫ݭ‬ᠭീˁ᝔ɤɛ⏏
(7) ̜̩̋̒˓́೴ǵ̫Ͳ̯˛Ͳाೣ༑ɭ‫ ᧑ܕ‬23 ‫ش‬ᐻȵϯेɐʳ‫ܩ‬ဣɴȡʳ೴⏏ໜा০઴ɻ Bc ݀ɞȵ࣬ဈ⏃˰ͻͲ⏆ȵȡʳ⏏1999 ॷ 7 ೐ 23-24 ౗⏃ᮒ࡚⏆ɴჼ‫ݭ‬͹஛ᮃᠭീˁ᝔ɤɛ⏏
(8) ̅Ͳ̸͠ᦏ᧿
ǵ᳋Ꭲˁ୫ɤɩᤉΫˁ๰ȦɩȦʳ‫ݖܩ‬з຦ɴ⏋1999 ॷ 7 ೐ 27 ౗⏃ᮒ࡚⏆ɴჼ‫ݭ‬ᠭീˁ᝔ɤɛ⏏
(9) ˵ʹˎ̠˭̜̒
ǵ˵ʹˎ̠˭̜̒ላɹμਕ‫Ͳ˛̯Ͳ̫⏏ܩ‬ाອᏙȴʰ஛‫؜‬ɇʶʳອᅋᣂ྇ɹʓȴɴ͍˱Ͳໜೣ
༑ʦ̊˓ȴʰɹᥥԂອᅋ႒ʢ༑ᦲɌɩȦʳ⏏1999 ॷ 7 ೐ 28-29 ౗⏃ᮒ࡚⏆⏋1999 ॷ 12 ೐ 19 ౗⏃ϗ࡚⏆⏋2000 ॷ 7 ೐ 17 ౗⏃ᮒ࡚⏆⏋2001 ॷ 3 ೐ 15 ౗⏃ϗ࡚⏆ɭ 7 ೐ 3 ౗⏃ᮒ࡚⏆⏋2002 ॷ 8 ೐
17 ౗⏃ᮒ࡚⏆ɴ॑‫ݭ‬ᠭീˁ᝔ɤɛ⏏
(10) ͞ˠ̫ˠ
ǵ̜̠̾͌ɭɹ‫ޅܕ‬ɴеᒞɐʳᅘ⏏̫Ͳ̯˛ͲाອᏙȴʰ஛‫؜‬ɇʶʳອᅋᣂ྇ɹʓȴɴ̜̠̾͌ȴʰɹᥥԂອᅋ႒ʢ༑ᦲɌɩȦʳ⏏1999 ॷ 7 ೐ 30 ౗⏃ᮒ࡚⏆ɴ॑‫ݭ‬ᠭീˁ᝔ɤɛ⏏
(11) ̫Ͳ̝Ͳ
ǵ͞ˠ̫ˠȴʰᏝ 20 km ά༑ɹ̫Ͳ̯˛Ͳाೣ༑ɹृࣳɴеᒞɐʳ⏏1999 ॷ 7 ೐ 30 ౗⏃ᮒ࡚⏆⏋॑‫ݭ‬ᠭീ⏋Ȳʭʃྫკ႒ˁ୫ɤɩᤉΫˁ๰ȦɩȦʳ‫ݖܩ‬з຦ɴჼ‫ݭ‬ᠭീˁ᝔ɤɛ⏏
(12) ̶ˎ͞Ͳ೴⏃̩͞Ͳ˵˓೴⏆ǵ‫ ᧑ܕ‬9 ‫ش‬ᐻɭ̫Ͳ̯˛Ͳाఌ༑ɹȩɠɹΥໜा⏋˵Ͳ̅˓ाɹϯे‫ܩ‬ဣɴȡʳ೴⏏
1999 ॷ 7 ೐ 31 ౗⏃ᮒ࡚⏆ɴ࣬ဈɹ࿠༑ᨆɫହᐨɴʭɤɩ᳋ᰖˁ஛‫؜‬ɌɩȦʳ‫ݖܩ‬з຦ɴჼ‫ݭ‬ᠭീˁ᝔ɤɛ⏏

56࣬ᅒ͹߂ី͹ೠ‫׫⏧ؤ‬ᨆ͞ˠ˽ɹॶᩃᨆɴȲȿʳ᳋ᰖɹᅊਖ਼‫ݭ‬ଘգᅎɭз຦ɹྫ֝༎֪

(13) ̨́ǵ̫Ͳ̯˛ͲाອᏙȴʰ஛‫؜‬ɇʶʳອᅋᣂ྇ɹʓȴɴ̊˓ȴʰɹᥥԂອᅋ႒ʢ༑ᦲɌɩȦʳ⏏
1999 ॷ 7 ೐ 31 ౗⏃ᮒ࡚⏆⏋Ȳʭʃ 2000 ॷ 7 ೐ 26 ౗⏃ᮒ࡚⏆ɴ॑‫ݭ‬ᠭീˁ᝔ɤɛ⏏
(14) ̶˝Ͳ̠͌೴ǵ̍͑ͲͅͲाΫ༑‫ݖ‬ɹε᭙ᨆɴеᒞɐʳ⏏Ʉɹ೴ɴԂʳଝմɹε᭙‫ܩ‬Օອ‫ݖ‬ɹᤉΫɴȡʳອɛʚʱ⏋Ȳʭʃɗɹดɴȡʳࢸྂ‫ܩ‬ɫ 2000 ॷ 7 ೐ 21 ౗⏃ᮒ࡚⏆ɴ஛ᮃᠭീˁ᝔ɤɛ⏏
(15) ͅͻ˵˓೴ǵ̶˝Ͳ̠͌೴ɭ‫ؾ‬෵ɴ̍͑ͲͅͲाΫ༑‫ݖ‬ɹε᭙ᨆɴеᒞɐʳε᭙‫ܩ‬ɹᢆ༅ɴȡʳອᅒɫ
2000 ॷ 7 ೐ 22-23 ౗⏃ᮒ࡚⏆ɴ஛ᮃᠭീˁ᝔ɤɛ⏏
(16) ̯Ͳ˹ˠ˗೴
ǵͅͻ˵˓೴ȴʰ‫ី׫‬ɴᏝ 2 km ɹ‫ܩ‬ဣɴȡʳ೴ɫɄɹ‫ݖܩ‬ˁБᝠɐʳ॑ȵᬖȴʶʳ⏏1999 ॷ
11 ೐ 15 ౗⏃ϗ࡚⏆ɴ॑‫ݭ‬ᠭീˁ᝔ɤɛ⏏
(17) ̨Ͳ˱͌೴
ǵ˵ʹˎ̠˭̜̒⏋̨́ᬚɴȡʳ೴ɫੈ।ᇕອ‫ݖ‬ɭɌɩɹ߂ȷȦ຾ȵȡʳ⏏Ʉɹ຾ɫ 2000 ॷ
7 ೐ 29-30 ౗⏃ᮒ࡚⏆ɴ஛ᮃᠭീˁ᝔ɤɛ⏏

3ȅࠫȁ‫ض‬3.1ȁ಺औ౷֖͈‫ݿ‬႒௖ǵᠭീೝᬚμ⏋‫غ‬៟ɫ 10 ᇿ⏋31 ዀ⏋158 ዛɹ᳋ᰖȵ஛ᮃʢɌȼɻᇿែተ᠗ɇʶɛ⏃ᝠ 1⏆⏏
ǵᲓᴏΫᇿȵ 74.1⎾⏃117 ዛ⏆ɭԃкɹᏝ 3/4 ˁ‫װ‬ʠʳΥ౅⏋˽̀˥ᇿɻ 12.7⎾⏃20 ዛ⏆ɭࢹɲȦ⏏Ʉʶɻ⏋ᲓᴏΫᇿɴ‫ي‬ʚʶʳ˱˓ዀȵ 72 ዛɫ೎ʢዛᰖధȵ޽ȼ⏋ԃкɹ 45.6⎾ˁ‫װ‬ʠ⏋ᐚȦɩ̝˻͚˗ዀ 16 ዛ⏃10.1⎾⏆⏋˧˧ዀ 9 ዛ⏃5.7⎾⏆ȵΫе 3 еɴеᒞɐʳɭȦȩჿᅓɴʭʳ⏏ʚɛ⏋Ʉɹȩɠ˱˓ዀɹ Cirrhinus cirrhosus⏋˱˓ Cyprinus carpio⏋̩˪ͤͲ
Hypophthalmichthys molitrix⏋ˡͩ˽͍̀ዀɹ̠˓̘͡ˑ̴͞ˏ Oreochromis niloticus ɹ 4 ዛᰖɻ᰸ຂɹɛʠɴውԂɇʶɛ‫޺ܕ‬ውԂዛɫȡʳ⏏ɇʰɴ⏋Pangasiidae ɹ Pangasius bocourti⏋̯̠͇ͤ̀ዀɹ Clarias batrachus ɻ͞ˠ˽‫׫‬ᨆɹ‫ܧ‬ഀዛɞȵ̊˓ɫ᰸ຂɇʶɛʢɹȵ‫ڃڧ‬ɭɌɩ॑‫ݭ‬ɫ᢫ެɇʶɩȦʳ⏏
3.2ȁ಺औ౷֖̤̫ͥͅ‫ݿ‬႒͈୆௳ાਫ਼͈ߊ໦
ǵᠭീ‫ ܩ‬17 ೴Ȳʭʃɗɹٞᦇɴࡔ‫ܧ‬ɐʳອ‫ݖ‬ˁ⏋ϗ࡚ɹອɹ೑ံ⏋ອᅠɹៀฐ⏋ঈᣒ⏋ອཉ⏋ᮒ࡚ɴȲȿʳЈɹອᅠɭɹᦹᐚਸፈˁᓒૅɌ⏋Гάɹʭȩɴ᳋ᰖɹᅊਖ਼‫ݭ‬ଘɭɌɩ‫ך‬ՕɌɛ⏏

A⏏Υಁᇕອ‫ݖ‬
A-1⏏Υಁᇕ๬ອ‫ݖ‬57

ˏ˻ˏ͹ˏ̶͠ˡ‫ݖܩ‬቉ድǵፀ 3 ‫ش‬ນ 1 ȁΨϋΪͺϋ୼କࠏ͈಺औ౷֖́‫ݿ̹̯ͦ෇ږ‬႒͈ςΑΠ1ǧOsteoglossiformesǵˏ̠ͦͩᇿ
ǵǣ Notopteridae ̠˧̠̠͇̊̀ዀ
Chitala ornata
1
Notopterus notopterus2ǧClupeiformesǵ̢˹Ͳᇿǵǣ Clupeidaeǵ̢˹Ͳዀ
Clupeichthys aesarnensis
3

2

3

4

R

5

6

D
D

RD
RD

R

D

7

ᠭǵീǵ‫ܩ‬ǵဣ
8
9 10 11

12

13

14

R

R

15

16

17

ǵǣ Sundasalangidae ˽Ͳ̋˵͞Ͳ˪˽ዀ

Sundasalanx mekongensis
4ǧCypriniformesǵ˱˓ᇿǵǣ Cyprinidaeǵ˱˓ዀ
Aaptosyax grypus
5
Albulichthys albuloides
6
Barbichthys laevis
7
Barbonymus altus
8
Barbonymus gonionotus
9
10 Boraras micros
11 Chela sp.
12 Cirrhinus cirrhosus
13 Cirrhinus ornatipinnis
14 Cirrhinus molitorella
15 Cirrhinus sp.
16 Crossocheilus reticulatus
17 Cyclocheilichthys apogon
18 Cyclocheilichthys armatus
19 Cyclocheilichthys armatus?
20 Cyclocheilichthys enoplos
21 Cyclocheilichthys furcatus?
22 Cyclocheilichthys repasson
23 Cyprinus carpio
24 Esomus metallicus
25 Esomus sp.
26 Garra fasciacauda
27 Hampala dispar
28 Henicorhynchus siamensis
29 Hypophthalmichthys molitrix
30 Hypsibarbus lagleri
31 Hypsibarbus malcomi
32 Hypsibarbus sp.
33 Labeo pierrei
34 Labeo barbatula
35 Labeo chrysophekadion
36 Labiobarbus leptocheila
37 Lobocheilos melanotaenia
38 Lobocheilos sp.
39 Macrocirichthys macrochirus

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̜̩̋̒˓́೴⏩3⏏̩̠̜͌͞Ͳ೴⏩4⏏̅Ͳ̸͠ᦏ᧿⏩5⏏˰Ͳ˱˪⏩6⏏˵ʹˎ̠˭̜̒⏩7⏏͞ˠ̫ˠ⏩
8⏏̫Ͳ̝Ͳ⏩9⏏̶ˎ͞Ͳ೴⏩10⏏̨́⏩11⏏˵̠͌˵˓೴⏩12⏏̯Ͳ˹ˠ˗೴⏩13⏏̶˝Ͳ̠͌೴⏩14⏏
ͅͻ˵˓೴⏩15⏏̨Ͳ˱͌೴⏩16⏏̨Ͳˡ̼͌ͻ̜೴⏩17⏏̊ͻ̩ˠा⏏

58࣬ᅒ͹߂ី͹ೠ‫׫⏧ؤ‬ᨆ͞ˠ˽ɹॶᩃᨆɴȲȿʳ᳋ᰖɹᅊਖ਼‫ݭ‬ଘգᅎɭз຦ɹྫ֝༎֪

1
40 Mystacoleucus atridorsalis
41 Mystacoleucus marginatus
42 Mystacoleucus sp.
43 Opsarius koratensis
44 Osteochilus hasseltii
45 Osteochilus lini
46 Osteochilus melanopleura
47 Osteochilus microcephalus
48 Osteochilus sp.
49 Oxygaster pointoni
50 Parachela maculicaua
51 Parachela siamensis
52 Parachela sp.
53 Paralaubuca barroni
54 Paralaubuca sp.
55 Paralaubuca typus
56 Poropuntius laoensis
57 Poropuntius sp.
58 Probarbus jullieni
59 Puntioplites falcifer
60 Puntioplites sp.
61 Puntius aurotaeniatus
62 Puntius aurotaeniatus?
63 Puntius brevis
64 Puntius jacobusboehlkei
65 Puntius orphoides
66 Puntius sp.
67 Raiamas guttatus
68 Rasbora aurotaenia
69 Rasbora borapetensis
70 Rasbora paviei
71 Rasbora spilocera
72 Rasbora rubrodorsalis
73 Rasbora trilineata
74 Rasbora sp.
75 Scaphognathops stejnegeri
76 Thynnichthys thynnoides
ǵǣ Balitoridaeǵ̢̨̊̓͠ዀ
77 Homaloptera sp.
78 Nemacheilus sp.
79 Nemacheilus sp.
ǵǣ Cobitidaeǵ̝˻͚˗ዀ
80 Acanthopsis sp.
81 Acanthopsis sp.
82 Acanthopsoides sp.
83 Botia eos
84 Botia helodes
85 Botia lecontei
86 Botia modesta
87 Botia morleti
88 Lepidocephalichthys hasseltii?
89 Lepidocephalichthys sp.
90 Lepidocephalichthys sp.
91 Lepidocephalichthys sp.

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59

ˏ˻ˏ͹ˏ̶͠ˡ‫ݖܩ‬቉ድǵፀ 3 ‫ش‬

1
92 Lepidocephalichthys sp.
93 Lepidocephalichthys sp.
94 Lepidocephalichthys sp.
95 Pangio anguilarisǧSiluriformesǵ̠͇̀ᇿǵǣ Bagridaeǵ˧˧ዀ
96 Bagrichthys obscurus
97 Hemibagrus aff. nemurus
98 Hemibagrus wyckioides
99 Mystus albolineatus
100 Mystus atrifasciatus
101 Mystus bocourti
102 Mystus mysticetus
103 Mystus sp.
104 Mystus sp.
ǵǣ Siluridaeǵ̠͇̀ዀ
105 Belodontichthys truncatus
106 Hemisilurus mekongensis
107 Kryptopterus apogon
108 Kryptopterus sp.
109 Ompok bimaculatus
110 Wallago attu
ǵǣ Pangasiidaeǵ̮Ͳˤ˹˗˽ዀ
111 Helicophagus waandersi
112 Pangasius bocourti
113 Pangasius concophilus
114 Pangasius pleurotaenia
115 Pangasius pleurotaenia?
116 Pangasius polyuranodon?
ǵǣ Schilubeidaeǵ˽˥̾͡ዀ
117 Laides longibarbis
ǵǣ Clariidaeǵ̯̠͇ͤ̀ዀ
118 Clarias batrachusǵǣ Sisoridaeǵ˹̅͡ዀ
119 Bagarius sp.
120 Glyptothorax sp.
ǵǣ Akysidaeǵˏ˥͖˹˽ዀ
121 Akysis sp.
ǧBeloniformesǵ̋̓ᇿǵǣ Adrianichthyidaeǵ͍̋ˡዀ
122 Oryzias pectoralis
123 Oryzias sp.
124 Oryzias sp.
ǵǣ Belonidaeǵ̋̓ዀ125 Xenentodon canciloides
ǵǣ Hemiramphidaeǵ˵͜͠ዀ
126 Dermogenys siamensisǧGasterosteiformesǵ̜˰˗ˠᇿ
ǵǣ Syngnathidaeǵ͜˗˻˗ˠዀ
127 Microphis sp.
ǵǣ Indostomidaeǵ˓Ͳ̝˽̜͌˽ዀ
128 Indostomus sp.
ǧSynbranchiformesǵ̊˗̠˧ᇿ
ǵǣ Chaudhuriidae ˡ˗̝˕͠ˏዀ

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60

࣬ᅒ͹߂ី͹ೠ‫׫⏧ؤ‬ᨆ͞ˠ˽ɹॶᩃᨆɴȲȿʳ᳋ᰖɹᅊਖ਼‫ݭ‬ଘգᅎɭз຦ɹྫ֝༎֪

1
129 Chaudhuria caudataǵǣ Mastacembelidae ̜˰˗̠˧ዀ
130 Macrognathus siamensis
131 Macrognathus semiocellatus
132 Mastacembelus armatus
133 Mastacembelus sp.
ǵǣ Synbranchidaeǵ̊˗̠˧ዀ
134 Monopterus albusǧPerciformesǵ˽̀˥ᇿǵǣ Chandidae ̊ˡ˵˳˓˹͐̍ዀ
135 Parambasis siamensis
ǵǣ Datnioididae
136 Datnioides undecimradiatus
ǵǣ Cichlidaeǵˡͩ˽͍̀ዀ
137 Oreochromis niloticus
ǵǣ Nandidaeǵ̠Ͳ̝˕˽ዀ
138 Nandus oxyrhynchusǵǣ Pristolepididae
139 Pristolepis fasciata
ǵǣ Odontobutidaeǵ̝Ͳ˱ዀ
140 Neodontobutis aurarmus
ǵǣ Gobiidaeǵ̩̄ዀ
141 Brachygobius mekongensis
142 Gobiopterus sp.
143 Oxyeleotris marmorataǵǣ Anabantidae ˥̨̓͠˗ˠዀ
144 Anabas testudineus
ǵǣ Osphronemidae ˠ˽̺̦ͦ͌˽ዀ
145 Betta smaragdina
146 Osphronemus exodon
147 Trichogaster trichopterus
148 Trichopsis schalleri
149 Trichopsis vittata
ǵǣ Channidae ̊˓ͩͲ̝˻͚˗ዀ
150 Channa gachua
151 Channa lusius
152 Channa maluria?
153 Channa micropertes
154 Channa striataǧPleuronectiformesǵˡͤ˓ᇿǵǣ Soleidae ˵˵˗˹̨˹̊ዀ
156 Brachirus harmandiǧTetraodontiformesǵ̶˫ᇿǵǣ Tetraodontidaeǵ̶˫ዀ
157 Monotrete suvattii
158 Monotrete sp.

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ˏ˻ˏ͹ˏ̶͠ˡ‫ݖܩ‬቉ድǵፀ 3 ‫ش‬

A-1-1⏏ອɛʚʱǵᮒೝɴᤉΫʦఔ႑‫ܩ‬ɴɫȷʳෑʠɩࢸៀฐɫ೎ʢέ࡫ࡴɲΥಁᇕອ‫ݖ‬ɫȡʳ⏏
A-1-2⏏Პеອᅒǵϗ࡚ɴອɻɲȦȵ⏋ᮒ࡚ɹᓚсೝɴɻΥಁᇕອ‫ݖ‬ɭɲʳ⏏Њᦏɹੈχᇕອ‫ݖ‬ɭɻ᧙ᤂᮐɴ
ȡʳ⏏
A-1-3⏏жеອᅒǵϗ࡚ɴອɻɲȦȵ⏋ᮒ࡚ɹᓚсೝɴɻΥಁᇕອ‫ݖ‬ɭɲʳဣɴȲȦɩᲞеᅒɭ‫ؾ‬෵ɞȵ⏋ᮒ࡚ɻٞᦇɹໜᤄྲྀɭᏳ༑ˁЂɌɩᦹᐚɐʳ⏏
A-1-4⏏ဌྰອᅒǵϗ࡚ɴɻೣഀॵΫȵɤɩȦɛжеᅒȵளອͅͲ̺ˁьᅎɌɩԗʃླྀອɐʳ⏏ளອɹᬠѥΫ⏋ੈχᇕອ‫ݖ‬ɫȡʳ͍˱Ͳໜఌ༑ʦ໡ɹٞᦇɴеᒞɐʳ⏏
A-1-5⏏ະ࿭؅͹ອ໗കǵ͍˱Ͳໜೣ༑⏋ఌ༑⏋ໜᤄྲྀ⏋ঃȦ໡⏋຾ɲɰɹࣳʦɗʶɴᦏȦ‫ݭ‬ଘɴȡʱ⏋ᮒ࡚ɴɻອ
໗ɐʳΥಁᇕອ‫ݖ‬ɫȡʳ⏏
A-2⏏Υಁᇕ༑ອ‫ݖ‬
A-2-1⏏ΥಁᇕᏳ༑ǵϗ࡚ɴອɻɲȦΥಁᇕອ‫ݖ‬ɫ⏋ᮒ࡚ɴɲʳɭᅒʦ຾⏋ໜᤄྲྀʦఌ༑ɲɰˁᦹᐚɇɓʳลᔨˁ೑ɐʳ⏏‫ݭ‬ଘɴʭʱᮒೝɴྂᅊඨ႒ʦୈອඨ႒ȵᑢᗛɐʳ⏏ঈᣒɻቆɫ⏋ៀฐɻࢸɇȦȵໜा০઴ɻဈɹᨆՕɹອཉȵཉȼ⏋ອᯆȵ໳ጓɛɲȦ Bc ݀ɫȡʳ⏏

B⏏ੈχᇕອ‫ݖ‬
B-1⏏ੈχᇕ๬ອ‫ݖ‬
B-1-1⏏ໜᤄྲྀǵ͍˱Ͳໜೣ༑ʦఌ༑ɴᦏஞɌɛ‫ݭ‬ଘɴȡʳ๬ອɹੈχᇕອ‫ݖ‬ɫ⏋ࣳᦇٞᦇɴɻະ࿭؅ʦ
༇ອകˁ೑ɐʳ⏏ᮒ࡚ɴɻ͍˱Ͳໜೣ༑ᨆʦఌ༑ᨆɭອᤉˁЂɌɩᦹᐚɐʳ⏏ঈᣒɻ໵ɫໍ
ອ͹ୈອ͹༝ᘽ͹༝ᧉɲʰʃɴྂᅊඨ႒ȵᑢᗛɐʳ⏏̩̠̜͌͞Ͳ೴ɹ̨Ͳ̊˓̶ˏ⏋˰Ͳ˱˪೴ɹ̨ͲˡͲ⏋̨Ͳ͍̩ͻͲȵɄʶɴȡɛʳ⏏
B-1-2⏏໡ǵໜᤄྲྀɴᰖаɐʳȵ⏋ອᅠɹៀฐȵ߂ȷȼ⏋ϗ࡚ɴॵΫȵɤɩȦɛອᤉȵᮒ࡚ɹອеɹΫ౥ɴʭʱ͍˱Ͳໜఌ༑ɭᦹᐚɐʳʢɹɹᤂᮐᇕɴɻɗʶɭ᧙Ȧ‫ݭ‬ଘɴȡʳ๬ອ‫ݖ‬ˁɄɄɫɻ
໡ɭ٧ʉɄɭɴɐʳ⏏˰Ͳ˱˪ᦏ᧿ɹ˽˓ȵɄʶɴȡɛʳ⏏
B-1-3⏏຾ǵໜᤄྲྀɴᰖаɐʳȵ⏋ᮒ࡚ɫʢٞᦇɹੈχᇕອ‫ݖ‬ɭᏳ༑ɲɰɴʭʱᦹᐚɌɲȦ๬ອ‫ݖ‬ˁɄ62࣬ᅒ͹߂ី͹ೠ‫׫⏧ؤ‬ᨆ͞ˠ˽ɹॶᩃᨆɴȲȿʳ᳋ᰖɹᅊਖ਼‫ݭ‬ଘգᅎɭз຦ɹྫ֝༎֪

Ʉɫɻ຾ɭ٧ʉɄɭɴɐʳ⏏
B-2⏏ੈχᇕ༑ອ‫ݖ‬
B-2-1⏏ੈχᇕᏳ༑ǵϗ࡚ʢອɹ༲ʶɲȦᏳ༑ɫ⏋ໜᤄྲྀʦ໡ɭᦹᐚɐʳ⏏ໜाᄩ‫ޅ‬ɻΫ៨ɹΥಁᇕᏳ༑ɭᰖаɐʳ⏏
B-2-2⏏͍˱Ͳໜఌ༑
ǵϗ࡚ɫʢອȵɲȼɲʰɲȦੈχᇕອ‫ݖ‬ɫȡʳȵ⏋‫ݭ‬ଘɴʭɤɩɻɄɹಁೝ⏋ອཉȵᘿɌȼ༓ȼɲʱϺȵਂ๰ɫʢाˁดుɫȷʳʭȩɴɲʳ⏏ʚɛ⏋͍˱Ͳໜೣ༑ɴປʏɩा३ȵႷȦ⏏ঈᣒɻቆ໵ɫ⏋ໜा০઴ɻဈɹᨆՕɹອཉȵཉȼ⏋ອᯆȵ໳ጓɛɲȦ Bc ݀ɫȡʳ⏏
B-2-3⏏͍˱Ͳໜೣ༑
ǵϗ࡚⏋ᮒ࡚ˁ‫ک‬ʼɑອȵࡔ‫ܧ‬ɌɩȦʳɭȦȩઝ٢ɫ⏋Ʉɹ‫ݖܩ‬ɴȲȿʳʢɤɭʢ࡫ࡴɌɛ
ੈχᇕອ‫ݖ‬ɫȡʳ⏏ɛɞɌ⏋ϗ࡚ɭᮒ࡚ɹອеेɻధ m ɴ‫ؖ‬ʉ⏏ঈᣒɻቆ໵ɫ⏋ໜा০઴ɻဈɹᨆՕɹອཉȵཉȼ⏋ອᯆȵ໳ጓɛɲȦ Bc ݀ɫȡʳ⏏
B-2-4⏏࣬ဈǵ͍˱Ͳໜೣ༑ʦఌ༑ɻቆ໵ɹঈᣒȵযǾɭᐚȼ⏏ɌȴɌ⏋ჼ‫ؿܩ‬ɫ˰ͻͲɭ٧ɽʶʳ‫ݭ‬ଘɻອঈȴʰ࣬ኂȵ᭠ᣱɌ⏋ाɻອɌʉȷˁȡɁɩ࣬আˁ༑ʶάʳਸ਼༑ᨆɭɲʳ⏏Ʉɹά༑ᨆɴɻʼɑȴɴኊঈȵࡔ‫ܧ‬ɐʳɄɭʢȡʳ⏏
B-2-5⏏ཌǵ͍˱Ͳໜೣ༑ʦఌ༑ɫ⏋႕ɴອཉɹȡʳཌˁჼ‫ؿܩ‬ɫͩͲɭ٧ʉ⏏

3.3ȁ‫ݿ‬႒͈୆‫ڰ‬ঃ͂୆௳ાਫ਼၌ဥ
ǵᠭീ‫ݖܩ‬ɫተ᠗ɇʶɛ᳋ᰖɹȩɠ⏋ɗɹᅊ༎‫ر‬ɭᅊਖ਼‫ݭ‬ଘɹգᅎɹᬠѥɴᬠɐʳ੻‫ݬ‬ȵ޽ȴʶࢹɲȴʶ਄ʰʶɛዛᰖɻᏝ 56 ዛɫȡʳ⏏๭ተɲዛధˁ៨ɇɲȦɹɻ዗᳋ɹຍ᭥ɫ⏋ࣚɻ‫ؾ‬ࡴɫȷʳȵዛᰖʚɫɻჼ‫ܧ‬ɴȲȦɩɻέ‫خ‬ᔨɲʢɹȵ‫ي‬ʚʶʳȴʰɫȡʳ⏏Гάɴɗʶʰˁෙᠦɐʳ⏏

A⏏Υಁᇕອ‫ݖ‬ɹʛɴᅊਖ਼ɐʳ᳋ᰖǵ˱˓ዀɹ Esomus metallicus⏃Ԡሎ 1⏆ɻ ᮒ࡚ɴжеອᅒɻʢɠʸ˃ɹɄɭ⏋Პеອᅒɫʢ҅кధɻෑʠɩ޽ȼ⏋᧑ᤉΫɹອɛʚʱɴʢՏჼɐʳ⏏ʚɛ⏋ᮒ࡚ɴ޽ᩄɹᮒȵᭇʱ⏋ȦɛʳɭɄʸȵອ༡ɌɴɲʳʭȩɲႥ઴ɹಁ⏋ೣዛȵ᧑ᤉΫɫʢ⏋ອཉɹෑʠɩ༓ȦᇤᕖႥɹອ
ɹ༑ʶɹμˁృɌȦΥಁᇕອ‫ݖ‬ˁິʠɩᇵ˃ɴ᧚ΫɌɩȦȼࠒȵ᠗ʠʰʶʳ⏏ອᅒԒʦɗʶ
ɴ᭰ஞɐʳఔ႑‫ܩ‬ɹອɛʚʱፈȴʰ⏋ʈ‫׋‬ሁ৿ɹմೝІ᳋ȴʰଁ᳋ʚɫ‫ଁع‬ᬏຍ᭥ȵ‫ؾ‬ಁɴ஛ᮃɇʶʳ⏏ɄɹɄɭȴʰ⏋ೣዛȵɄɹ‫ݭ‬ଘɫᑢຂɌ⏋ԃᅊ༎‫ر‬ˁȲȼɤɩȦʳɄɭɻተࡸ
63ˏ˻ˏ͹ˏ̶͠ˡ‫ݖܩ‬቉ድǵፀ 3 ‫ش‬ৢ૯ 1ȁEsomus metallicus ͈౽‫ݿ‬

ৢ૯ 2ȁLepidocephalichthys sp. ଼͈‫ݿ‬ɫȡʳ⏏ϗ࡚ɻອᅒʦᏳ༑ԒɹॵΫȵʱȴȿɛອɛʚʱɴ᠗ʠʰʶʳʢɹɹ⏋ੈχᇕອ‫ݖ‬ɫೣዛɹᅊਖ਼ȵተ᠗ɇʶɲȦ⏏‫ݖܩ‬з຦ɹ᠌ɫɻ⏋ᮒ࡚ɹɻɍʠɴੈχᇕອ‫ݖ‬ɭɻʚɞᦹᐚਸ
ȵɲȦອᅒɴອȵྊʚʱɻɍʠʳɭɐȽɴࢸ݀҅кȵʛʰʶʳɭȦȩ⏏ɄɹɄɭɻᘿᓔʰʢተ᠗Ɍɛ⏏Јɹອ‫ݖ‬ȴʰው֪ɌɩɄɲȦɹɴɄɹʭȩɲჼᢒȵȲȷʳɹɻ⏋ೣዛɹ‫׺‬ȵϗၥᓘχਸˁ୫ɠ⏋Ʉʶȵϗ࡚ɹᬚɻອᅒɹ‫ܤ‬μɴȡʱ⏋ອȵྊʚʳɭ‫ؾ‬ಁɴɗʶȵʈ‫׋‬ɐʳɭȦȩ‫خ‬ᔨਸȵᲞȦ⏏
ǵ˱˓ዀɹ Rasbora rubrodorsalis ʢᮒ࡚⏋ϗ࡚ˁɭȲɌɩੈχᇕອ‫ݖ‬ɴՏჼɓɑ⏋ϗ࡚ʢອɛʚʱɹʭȩɲέ࡫ࡴɲ‫ݭ‬ଘɫᣴϗɐʳဣɫմᓔɭ޴ʼʰɲȦȵ⏋ᲞеອᅒԒɫɻ҅кధȵࢹɲȦ⏏
ǵ̝˻͚˗ዀɹ Lepidocephalichthys spp.
⏃Ԡሎ 2⏆ʢ Esomus metallicus ɭ‫ؾ‬෵ɲᅊ༎‫ر‬ˁȲȼʳʢɹɭਲ਼ʼʶʳȵ⏋ೣዛɹ‫⏋غݭ‬Esomus metallicus ɭᅵɲʱ⏋ଁ᳋ɻྂ঎ȵȡʶɽᬏಁᬚ
ᅊࡔ‫خ‬ᔨɲɹɫ⏋ϗ࡚ɴɻອᅒɹঈཉȼ࿃ԂɌɩᣴϗɐʳ‫خ‬ᔨਸʢ‫ࡴو‬ɫȷɲȦ⏏
ǵΫ៨ 3 ዛɻϗ࡚ɹဌྰᅒɫʢተ᠗ɇʶ⏋႕ɴմ 2 ዛɻ዗᳋ʦᗐ᳋ʢ஛ᮃɇʶɛ⏏

B⏏ϗ࡚ɴɻੈχᇕ๬ອ‫ݖ‬ɴᅊਖ਼Ɍ⏋ᮒ࡚ɴɻΥಁᇕອ‫ݖ‬ɴʢଁ᳋ȵᅊਖ਼ɐʳ᳋ᰖǵɄɹʭȩɲᅊ༎‫ر‬ˁȲȼʳ᳋ᰖɻ⏋գᅎɐʳᅊ༎‫ݭ‬ଘɹेɴʭʱГάɹ 4 ̊˓̺ɴ‫ך‬Օɇʶʳ⏏
B-1⏏ΥಁᇕᏳ༑⏋жеອᅒ⏋Პеອᅒɫᑢຂɐʳ᳋ᰖǵ̯̠͇ͤ̀ዀɹ Clarias batrachus⏃Ԡሎ 3⏆⏋̜˰˗̠˧ዀɹ Macrognathus siamensis⏃Ԡሎ4⏆⏋˥̨̓͠˗ˠዀɹAnabas testudineus⏃Ԡሎ 5⏆⏋̊˓ͩͲ̝˻͚˗ዀɹ Channa striata⏃Ԡሎ 6⏆ȵɄʶɴȡɛʳ⏏Channa striata ɻᅤɹᗳȵອᯆɴុȦ᝵ɇʳʭȩɲ‫ݭ‬ଘ⏋ʚɛྂਸ⏋ୈອඨ႒ȵᑢᗛɐʳ‫ݭ‬ଘɴ༝ਸ‫׺‬ˁᅋ‫៊⏋׺‬᳋ɻʈ‫৿׋‬ʢɄʶʰˁࡪʳ⏏‫ݖܩ‬з຦ɹ੻‫ݬ‬ɴʭʶɽ⏋Clarias batrachus ɻʦʦອཉɹȡʳࣳᦇɹອঈɴ៊᳋ȵ‫ܤ‬ˁɲʰɌɩ͇˗Ͳ̝Ⴅɹ
ुˁᦶʱ⏋‫׺‬ˁѳᡸɐʳɭȦȩ⏏
64࣬ᅒ͹߂ី͹ೠ‫׫⏧ؤ‬ᨆ͞ˠ˽ɹॶᩃᨆɴȲȿʳ᳋ᰖɹᅊਖ਼‫ݭ‬ଘգᅎɭз຦ɹྫ֝༎֪

ৢ૯ 3ȁClarias batrachus ͈౽‫ݿ‬

ৢ૯ 4ȁMacrognathus siamensis ͈౽‫ݿ‬

ৢ૯ 5ȁAnabas testudineus ͈ဘ‫ݿ‬

ৢ૯ 6ȁChanna striata ͈౽‫ݿ‬ǵMacrognathus siamensis ˁ᭑ȼʓȴɹ 3 ዛɻជ֖ዷຩ٧ُ۬ࡲˁ೑Ɍ⏋ϗ࡚ɫʢྂ঎ȵӱՕɲ‫ݭ‬ଘɫȡʶɽ‫ܤ‬μɫᣴϗɌɩȦʳ‫خ‬ᔨਸȵȡʳ⏏ʚɛ⏋Macrognathus siamensis ʢкȵྂɤɩȦʶɽᬏಁᬚᅊࡔ‫خ‬ᔨɲɹɫ⏋մ 3 ዛɭ‫ؾ‬෵ɹ‫خ‬ᔨਸȵᓒȭʰʶʳ⏏
ǵɲȲ⏋Channa striata ɻϗ࡚ɹဌྰᅒɴȲȦɩ዗᳋ɭॾ᳋ȵ஛ᮃɇʶɛ⏏
B-2⏏ΥಁᇕᏳ༑⏋жеອᅒɫᑢຂɐʳ᳋ᰖǵ˱˓ዀɹ Puntius aurotaeniatus⏃Ԡሎ 7⏆⏋̊˗̠˧ዀɹ̊˗̠˧ Monopterus albus⏃Ԡሎ 8⏆ȵɄʶɴȡɛʳ⏏
ǵ̊˗̠˧ɻዷຩμȴʰᨧᏬˁ٧ُɐʳɄɭȵɫȷ⏋кȵྂɤɩȦʶɽᬏಁᬚᅊࡔ‫خ‬ᔨɲɹɫ⏋ྂ঎ɹȡʳঈᣒȵȡʶɽϗ࡚ɫʢжеອᅒʦΥಁᇕᏳ༑ɫᣴϗɌɩȦʳ‫خ‬ᔨਸȵᲞȦ⏏
B-3⏏ΥಁᇕᏳ༑⏋ໜᤄྲྀᑉᦇᨆɹະ࿭؅ɫᑢຂɐʳ᳋ᰖǵ˱˓ዀɹ Puntius brevis⏃Ԡሎ 9⏆⏋˽̀˥ዀɹ Parambasis siamensis⏃Ԡሎ 10⏆ȵɄʶɴȡɛʳ⏏ᮒ࡚ɫʢອᅒɴѢԂɐʳɄɭɻॕɫȡʳ⏏
B-4⏏жеອᅒ⏋ໜᤄྲྀᑉᦇᨆɹະ࿭؅ɫᑢຂɐʳ᳋ᰖǵ༑ອ‫ݖ‬ˁ߫ʚɲȦዛᰖɫ⏋ໍອඨ႒ፈɹອᗳȵᑢᗛɌɛᄩ‫ޅ‬ɴ޽ȼՏჼɐʳȵ⏋ᮒ࡚ɴɻɗʶʰɹɲȦжеອᅒʦໜᤄྲྀᑉᦇᨆɹະ࿭؅ɴʢᅊਖ਼ɐʳ⏏ʚɛ⏋ʓȴɭ᧓Ȧ⏋ϗ࡚ɫʢ
ᑢຂɐʳ⏏
65

ˏ˻ˏ͹ˏ̶͠ˡ‫ݖܩ‬቉ድǵፀ 3 ‫ش‬ৢ૯ 7ȁPuntius aurotaeniatus ͈౽‫ݿ‬

ৢ૯ 8ȁΗ;΢Ά Monopterus albus ͈౽‫ݿ‬

ৢ૯ 9ȁPuntius brevis ͈౽‫ݿ‬

ৢ૯ 10ȁParambasis siamensis ଼͈‫ݿ‬

ৢ૯ 11ȁOryzias pectoralis ଼͈‫ݿ‬

ৢ૯ 12ȁTrichopsis schalleri ଼͈‫ݿ‬ǵ͍̋ˡዀɹ͍̋ˡࣚ⏃Oryzias pectoralis ɲɰ⏆ɴ‫ي‬ʚʶʳዛᰖ⏃Ԡሎ 11⏆Ȳʭʃˠ˽̺ͦ
̦˽͌ዀɹ Trichogaster trichopterus, Trichopsis schalleri⏃Ԡሎ 12⏆, Trichopsis vittata ȵɄʶɴȡɛʳ⏏
ǵɲȲ⏋Trichogaster trichopterus ɭ Trichopsis schalleri ɻϗ࡚ɹဌྰᅒɫʢ஛ᮃɇʶ⏋႕ɴմᓔɻᅋ‫׺‬আʦ዗᳋ʢተ᠗ɇʶɛ⏏

C⏏ଁ᳋ɻੈχᇕອ‫ݖ‬ɴᅊਖ਼Ɍ⏋ᮒ࡚ɴΥಁᇕອ‫ݖ‬ˁᅋ‫ݭ׺‬ɭɐʳ᳋ᰖǵɄɹʭȩɲᅊ༎‫ر‬ˁȲȼʳ᳋ᰖɻ⏋գᅎɐʳᅊ༎‫ݭ‬ଘɹेɴʭʱГάɹ 2 ̊˓̺ɴ‫ך‬Օɇ
66

࣬ᅒ͹߂ី͹ೠ‫׫⏧ؤ‬ᨆ͞ˠ˽ɹॶᩃᨆɴȲȿʳ᳋ᰖɹᅊਖ਼‫ݭ‬ଘգᅎɭз຦ɹྫ֝༎֪

ʶʳ⏏
C-1⏏ଁ᳋ɻੈχᇕ༑ອ‫ݖ‬ɴᅊਖ਼ɐʳ᳋ᰖǵ̠͇̀ዀɹ Wallago attu⏃Ԡሎ 13⏆ȵɄʶɴȡɛʳ⏏ೣዛɹଁ᳋ɻ͍˱Ͳໜೣ༑ʦఌ༑ɴ
ᅊਖ਼ɌɩȦʳ⏏ᮒ࡚ɹɻɍʠ⏋߂ᩄɹᭇᮒɴʭʱອеɹਸ਼Ϋ౥ɫ⏋ϗ࡚ɴɻॵΫȵɤɩȦɛᏳ༑ȵླྀອɌɩໜᤄྲྀɭ͍˱Ͳໜɹ༑ອᨆȵᦹᐚɐʳ⏏ೣዛɻɄɹಁˁʛɻȴʰȦ߂ᒴˁɲɌɩໜᤄྲྀɴѢԂɐʳ⏏‫ݖܩ‬з຦ɹ੻‫ݬ‬ɴʭʶɽ⏋ະ࿭؅ʦອ໗കɫᅋ‫׺‬ɐʳɭȦȩ⏏ɗɹ৿⏋ໜᤄྲྀˁᔒଁ‫ݭ‬ଘɭɌɩգᅎɌ⏋ଁᬏɭɭʢɴ͍˱Ͳໜೣ༑ɴʢɰʳ⏏
C-2⏏ଁ᳋ɻੈχᇕ๬ອ‫ݖ‬ɴᅊਖ਼ɐʳ᳋ᰖǵ̠͇̀ዀɹ Ompok bimaculatus⏃Ԡሎ 14⏆⏋̊˓ͩͲ̝˻͚˗ዀɹ Channa lusius⏃Ԡሎ 15⏆⏋
Ch. micropertes ȵɄʶɴȡɛʳ⏏
ǵOmpok bimaculatus ɹଁ᳋ɻ໡ʦໜᤄྲྀɴᅊਖ਼ɌɩȦʳ⏏ᮒ࡚ɴໜᤄྲྀٞᦇɹະ࿭؅ɫ዗
᳋ȵ஛ᮃɇʶʳɄɭȴʰ⏋Ʉɹ‫ݭ‬ଘˁᅋ‫ݭ׺‬ଘɭɌɩգᅎɌɩȦʳɴ᧓ȦɲȦ⏏ɇʰɴᗐ᳋ɻжеອᅒʦੈχᇕ͹ΥಁᇕᏳ༑ɴʢѢԂɌɩଁᬏɐʳ⏏
ǵChanna lusius ɭ Ch. micropertes ɻঃȦᬖఔᇕɲອᯆˁ୫ɦ໡ʦໜᤄྲྀፈɹ๬ອ‫ݖ‬ɴ޽Ȧ⏏
ະ࿭؅ʦ༇ອകɫ዗᳋ȵ஛ᮃɇʶʳɄɭȴʰɄʶʰɹ‫ݭ‬ଘˁᅋ‫ݭ׺‬ଘɭɌɩգᅎɌɩȦʳɴ᧓ȦɲȦ⏏

ৢ૯ 13ȁWallago attu ͈ဘ‫ݿ‬

ৢ૯ 14ȁOmpok bimaculatus ͈౽‫ݿ‬

ৢ૯ 15ȁChanna lusius ͈౽‫ݿ‬67

ˏ˻ˏ͹ˏ̶͠ˡ‫ݖܩ‬቉ድǵፀ 3 ‫ش‬D⏏ଁ᳋ɻੈχᇕອ‫ݖ‬ɴᅊਖ਼Ɍ⏋ᮒ࡚ɴΥಁᇕອ‫ݖ‬ˁᔒଁ‫ݭ‬ଘɭɐʳ᳋ᰖǵɄɹʭȩɲᅊ༎‫ر‬ˁȲȼʳ᳋ᰖɻ⏋գᅎɐʳᅊ༎‫ݭ‬ଘɹेɴʭʱГάɹ 2 ̊˓̺ɴ‫ך‬Օɇʶʳ⏏
D-1⏏ଁ᳋ɻੈχᇕ༑ອ‫ݖ‬ɴᅊਖ਼ɐʳ᳋ᰖǵ˱˓ዀɹ Hampala dispar⏃Ԡሎ 16⏆⏋Henicorhynchus siamensis⏋˧˧ዀɹ Mystus mysticetus ȵɄʶɴȡɛʳ⏏
ǵHampala dispar ɹଁ᳋ɻ͍˱Ͳໜೣ༑͹ఌ༑ɴ޽ȼᅊਖ਼ɌɩȦʳȵʼɑȴɲȵʰៀฐɹ߂ȷɲ໡ɴʢՏჼɐʳ⏏዗᳋ɻ͍˱Ͳໜఌ༑ʦ໡ɴɦʰɲʳੈχᇕఌ༑ɫ஛ᮃɇʶʳɹɫɄʶʰɹ‫ݭ‬ଘȵᅋ‫ݭ׺‬ɫȡʳ‫خ‬ᔨਸȵᲞȦ⏏ɗʶʭʱ߂݀ɹॾ᳋ɴɲʳɭໜᤄྲྀʦɗʶɴɦʰɲʳжеອᅒɴѢԂɌɩᅊᔒɌ⏋ଁᬏɴЫɤɩଁ᳋ɹᅊਖ਼‫ݭ‬ଘɴʢɰʳ⏏
ǵHenicorhynchus siamensis ɭ Mystus mysticetus ɹଁ᳋ʢΫዛɭ‫ؾ‬෵ɲ‫ݭ‬ଘɴᅊਖ਼ɌɩȦʳ⏏
ᅋ‫ݭ׺‬ଘɻέ౩ɫȡʳȵ⏋ॾ᳋ɻໜᤄྲྀʦɗʶɴɦʰɲʳжеອᅒɴѢԂɌɩɗʶʰɹ‫ݭ‬ଘˁᔒଁ‫ݭ‬ଘɭɌɩգᅎɌɩȦʳ⏏
D-2⏏ଁ᳋ɻੈχᇕ๬ອ‫ݖ‬ɴᅊਖ਼ɐʳ᳋ᰖǵ̢˹Ͳዀɹ Clupeichthys aesarnensis⏋˱˓ዀɹ Rasbora aurotaenia⏋̋̓ዀɹ Xenentodon
canciloides ȵɄʶɴȡɛʳ⏏
ǵClupeichthys aesarnensis ɻঃȦᬖఔᇕɲອᯆˁ୫ɦ໡ʦໜᤄྲྀፈɹ๬ອ‫ݖ‬ɴ޽Ȧ⏏ೣዛɹ˹͞˽݀І᳋ɻᮒ࡚ɴະ࿭؅ʦອ໗കˁᔒଁ‫ݭ‬ଘɭɌɩգᅎɌɩȦʳ⏏
ǵRasbora aurotaenia ɭ Xenentodon canciloides ɻໜᤄྲྀ⏋໡ɲɰɴᅊਖ਼Ɍ⏋Ʉʶʰɹ‫ݭ‬ଘɫᑢຂɐʳ⏏ᗐ᳋ɻᮒೝɴΥಁᇕᏳ༑ɲɰɴѢԂɌɩଁᬏɐʳ⏏

E⏏ଁ᳋ɻੈχᇕ༑ອ‫ݖ‬ɴᅊਖ਼Ɍ⏋ᮒ࡚ɴੈχᇕ๬ອ‫ݖ‬ˁІ͹዗᳋ɹᔒଁ‫ݭ‬ଘɭɌɩգᅎɐ
ʳ᳋ᰖǵ˱˓ዀɹ Barbonymus altus⏃Ԡሎ 17⏆⏋Crossocheilus reticulatus⏃Ԡሎ 18⏆⏋Cyclocheilichthysৢ૯ 16ȁHampala dispar ͈ဘ‫ݿ‬68࣬ᅒ͹߂ី͹ೠ‫׫⏧ؤ‬ᨆ͞ˠ˽ɹॶᩃᨆɴȲȿʳ᳋ᰖɹᅊਖ਼‫ݭ‬ଘգᅎɭз຦ɹྫ֝༎֪

ৢ૯ 17ȁBarbonymus altus ͈౽‫ݿ‬

ৢ૯ 18ȁCrossocheilus reticulatus ͈౽‫ݿ‬ɹЗᬚ⏋Labiobarbus leptocheila, Lobocheilos melanotaenia, Mystacoleucus atridorsalis, Paralaubuca
typus⏋̝˻͚˗ዀɹ Acanthopsoides sp., ˧˧ዀɹ Hemibagrus aff. nemurus, ̮Ͳˤ˹˗˽ዀɹ
Pangasius pleurotaenia ȵɄʶɴȡɛʳ⏏
ǵΫ៨Ɍɛ˱˓ዀɹዛᰖɻІ͹዗᳋ȵໜᤄྲྀɲɰɹ๬ອ‫ݖ‬ɫ஛ᮃɇʶʳɄɭȴʰ⏋ɄɄˁᔒଁ‫ݭ‬ଘɭɌɩգᅎɌɩȦʳɄɭɻተࡸɫȡʳ⏏ɌȴɌ⏋ᅋ‫׺‬ʢɄɹ‫ݭ‬ଘɫ᝔ʼʶɩȦʳȴɰȩȴɻჼ‫ܧ‬ɹɭɄʸέ౩ɫȡʳ⏏Ʉʶʰɹዛᰖɹ‫׺‬ȵɰɹʭȩɲਸᣒȴʢៗ౩ɫȷɩȦɲȦȵ⏋ʢɌʢ༝ਸ‫׺‬ɫȡʶɽ⏋‫ވ‬ອಁɴອɭɭʢɴ‫׺‬ȵໜᤄྲྀɴԂʱ⏋ʈ‫׋‬ɌɛІ᳋ȵɄɄɫᇍᔒɌɩȦʳ‫خ‬ᔨਸʢȡʳȴʰɫȡʳ⏏
ǵHemibagrus aff. nemurus ɹଁ᳋ɻςɴ͍˱Ͳໜೣ༑⏋ఌ༑ɴᅊਖ਼ɐʳ⏏‫ݖܩ‬з຦ɹ᠌ɫɻ⏋ఌ༑ɹཌȵᅋ‫ݭ׺‬ଘɫȡʳɭȦȩ⏏ॾ᳋ɻఌ༑ɹʓȴ⏋ໜᤄྲྀɲɰɴѢԂɌɩଁᬏɐʳ⏏
ǵAcanthopsoides sp. ɭ Pangasius pleurotaenia ɹᅋ‫ݭ׺‬ଘɴᬠɐʳ੻‫ݬ‬ɻɲȦȵ⏋዗᳋⏋ॾ᳋ȵໜᤄྲྀɫ஛ᮃɇʶʳɄɭȴʰ⏋Ʉɹ‫ݭ‬ଘˁᔒଁ‫ݭ‬ଘɭɌɩգᅎɌɩȦʳʢɹɭਲ਼ʼʶʳ⏏
ǵɄʶʰɹ᳋ᰖɴɻ቉ድɹᦽࣙɴʭʱᅋ‫ݭ׺‬ଘɹ੻‫ݬ‬ȵ਄ʰʶʶɽ⏋C-1 ɴ‫ך‬Օɇʶʳዛᰖʢ‫ي‬ʚʶɩȦʳʢɹɭਲ਼ʼʶʳ⏏

F⏏ੈχᇕອ‫ݖ‬ɹʛɴᅊਖ਼ɐʳ᳋ᰖǵɄɹʭȩɲ‫ݭ‬ଘɫᅊ༎‫ر‬ˁȲȼʳ᳋ᰖɻ⏋գᅎɐʳᅊਖ਼‫ݭ‬ଘɹेɴʭʱГάɹ 2 ̊˓̺ɴ‫ך‬Օɇʶʳ⏏
F-1⏏ੈχᇕ๬ອ‫ݖ‬ɴᅊਖ਼ɐʳ᳋ᰖǵ˱˓ዀɹ Boraras micros⏃Ԡሎ 19⏆⏋Rasbora boprapetensis, R. spilocera, R. trilineata⏋̝˻͚˗ዀɹ Pangio anguilaris⏋˵͜͠ዀɹ Dermogenys siamensis, カウドゥリア科ɹ Chaudhuria
caudata, ̠Ͳ̝˕˽ዀɹNandus oxyrhynchus⏃Ԡሎ 20⏆⏋̝Ͳ˱ዀɹ Neodontobutis aurarmus⏋
̩̄ዀɹ Brachygobius mekongensis ȵɄʶɴȡɛʳ⏏
ǵɄʶʰɹ᳋ᰖɹȩɠ⏋Rasbora boprapetensis ɻϗ࡚ɴॵΫȵʳɄɭɻɲȦʢɹɹ༓ȼɩࢸ
69ˏ˻ˏ͹ˏ̶͠ˡ‫ݖܩ‬቉ድǵፀ 3 ‫ش‬ৢ૯ 19ȁBoraras micros ଼͈‫ݿ‬

ৢ૯ 20ȁNandus oxyrhynchus ͈౽‫ݿ‬

ৢ૯ 21ȁAaptosyax grypus ͈ဘ‫ݿ‬

ৢ૯ 22ȁProbarbus jullieni ͈ဘ‫ݿ‬ៀฐɲᄩ‫ޅ‬ɴ޽Ȧ⏏
F-2⏏ੈχᇕ༑ອ‫ݖ‬ɴᅊਖ਼ɐʳ᳋ᰖǵ ˱ ˓ ዀ ɹ Aaptosyax grypus ⏃Ԡ ሎ 21⏆⏋Probarbus jullieni ⏃Ԡ ሎ 22⏆⏋Raiamas guttatus,
Scaphognathops stejnegeri, Hypsibarbus spp., ̠͇̀ዀɹ Kryptopterus spp., ˧˧ዀɹ Mystus
albolineatus, M. atrifasciatus ȵɄʶɴȡɛʳ⏏
ǵProbarbus jullieni ɹଁ᳋ɻ͍˱Ͳໜೣ༑ʦఌ༑ɴᅊਖ਼Ɍ⏋ϗ࡚ɴ࣬ဈɴᮃʚʱᅋ‫׺‬ɐʳ[Roberts and Warren 1994]⏏ࢸ݀ɹᗐ᳋ɻ͍˱Ͳໜఌ༑ɫ஛ᮃɇʶʳ⏏Hypsibarbus ɴࣚɐʳዛᰖɻ‫ݖܩ‬з຦ɹ᠌ɫɻ⏋ᮒ࡚ɴ߂ᒴɫ࣬ဈɴᮃʚʱ⏋࣬ɹΫɴᅋ‫׺‬ɐʳɭȦȩ⏏ɗɹʓȴ⏋ɄɄɴȡɁɛዛᰖɹᅋ‫ݭ׺‬ଘɴᬠɐʳ੻‫ݬ‬ɻɲȦȵ⏋ೣᠭീ‫ݖ‬ɴȲȦɩΥಁᇕອ‫ݖ‬ȴʰɹ៨᪼ɻɲȼ⏋዗᳋ʢɌȼɻॾ᳋ȵ͍˱Ͳఌ༑ᨆɫ஛ᮃɇʶʳɄɭȴʰ⏋Ʉɹ‫ݭ‬ଘˁᔒଁ‫ݭ‬ଘɭɌɩգᅎɌɩȦʳɄɭɻተȴɫȡʳ⏏
3.4ȁਯྦྷ͈‫ݽ‬Ⴛ‫ڰ‬൲͂‫ݿ‬႒͈୆௳ાਫ਼၌ဥ
ǵ࣬ᅒ [2002] ɻೣᠭീ‫ݖܩ‬ȴʰ 25 ዛɹྫ໬ˁ‫ݬ‬ٚɌɩȦʳȵ⏋ɗʶʰɭ᳋ᰖɹᅊਖ਼‫ݭ‬ଘգᅎɭɹᬠᦹɴɦȦɩɻᦚʏʰʶɩȦɲȦ⏏ೣ፲ɫɻɗɹ੻‫ݬ‬ˁ᠎Ᏻɴ៨ᥔɐʳ⏏

70࣬ᅒ͹߂ី͹ೠ‫׫⏧ؤ‬ᨆ͞ˠ˽ɹॶᩃᨆɴȲȿʳ᳋ᰖɹᅊਖ਼‫ݭ‬ଘգᅎɭз຦ɹྫ֝༎֪

ৢ૯ 23ȁ̞݉࿌Ȫ΍όͻϋȫৢ૯ 24ȁए਀࿌Ȫ΃Πϋȫ(1) தȦᐨ⏃˵ʹˑͲ⏆⏃Ԡሎ 23⏆ǵᦲ।ɻΥಁᇕອ‫ݖ‬ɹжеອᅒ⏋Პеອᅒፈɫьᅎɇʶʳ⏏ᮒ࡚ɴȲȿʳਸ਼࿠ɲ‫ވ‬ອಁɴɻΫ៨Ɍɛ‫ݭ‬ଘɹʓȴɴ⏋͍˱Ͳໜఌ༑ɹ‫׶‬᭝঎ɹࢹɲȦࣳᦇ⏋Ȳʭʃϗ࡚ɫʢ຾ʦໜᤄྲྀɫьᅎɇʶʳɄɭʢȡʳ⏏ςɭɌɩଁϺߨਸɭӻጣȵᅎȦ⏋Rasbora, Esomus metallicus, Trichopsis,
Trichogaster trichopterus, Anabas testudineus, Channa striata, Clalias batrachus ፈȵ஛஁ɇʶʳ⏏
(2) ؕଝᐨ⏃ˡ̜Ͳ⏆⏃Ԡሎ 24⏆ǵ͍˱Ͳໜఌ༑⏋຾ፈɹੈχᇕອ‫ݖ‬ɫᅎȦʰʶʳ⏏ଁϺᅖਸȵьᅎɌ⏋ᖐɹʍɇȷɴ‫ࡴܔ‬Ɍɩᖐˁᦽʠ⏋ᝠࣝˁᧉ໻ɌɩȦʳ᳋ᰖˁ஛஁ɐʳ౅໬ɭ⏋ೠɫᦶɤɛ᣸‫ݭ‬ɴɄɹᐨˁ‫ࡴܔ‬Ɍɩ༑άɌɩȼʳ᳋ᰖˁ஁ʰȭʳ 2 ዛᰖɹ౅໬ȵȡʳ⏏ьᅎಁೝʦ஛஁ࢪᢒ᳋ዛɴᬠɐʳ੻‫ݬ‬ɻ਄ʰʶɲȴɤɛ⏏
(3) ‫܋‬ଝᐨ⏃˵̝˕Ͳ⏆ǵଁϺߨਸɴʭʱॷᬚˁɭȲɌɩьᅎɇʶʳȵ⏋႕ɴᮒ࡚⏋ᭇᮒ৿ɹ‫ވ‬ອಁɴ᳋ᰖȵ༎ᇍɴ᝔֪ɐʳᅊ઴ˁգᅎɌɩΥಁᇕᏳ༑⏋ະ࿭؅⏋ອ໗ക⏋‫ݩ‬ອɌɛжеອᅒɹʓȴ⏋ੈχᇕອ‫ݖ‬ɫȡʳ͍˱Ͳໜೣ༑⏋ఌ༑⏋ໜᤄྲྀ⏋໡⏋຾ፈ⏋᣸‫ݭ‬ȵʭȼɄɹᐨɹఌ഼ᨆȵ࡭ԃɴɻອ໗Ɍɲ
Ȧʭȩɲ༓ଘɫᅎȦʰʶʳ⏏ᐨɴ᳋ˁ᠛ৄɐʳɛʠɹ᰹ɻԂʶɲȦ⏏ອঈɴᐨˁᘺɭɌɩɌɽ
ʰȼ৺ɠ⏋᳋ȵԂʳ᯴‫غ‬ˁឿ៟ʰɤɩளɁʳ௭сˁᑵʱᦐɐ⏏ᐨɹΫˁᦲ᧌ɐʳɐʏɩɹ᳋ዛ
ȵ஛஁ࢪᢒɭɲʳȵ˱˓ዀ⏋̝˻͚˗ዀ⏋̠͇̀ዀ⏋̜˰˗̠˧ዀ⏋̊˓ͩͲ̝˻͚˗ዀɲɰ
ɹࢸ͹μ݀ዛȵ޽ȼ⏋߂݀᳋ȵ஛஁ɇʶʳɄɭɻॕɫȡʳ⏏
(4) ହᐨ⏃̼ͻ)ǵଁϺᅖਸȵ߫˃ɫьᅎɐʳྫԍɫ⏋ੈχᇕອ‫ݖ‬ɫȡʳ͍˱Ͳໜೣ༑⏋ఌ༑⏋᭭ࢀ႒ɹɲȦ໡ʦໜᤄྲྀፈ⏋͍˱Ͳໜೣ༑⏋ఌ༑ɫɻঈᣒɻ‫ک‬ʼɲȦȵອཉɹȡʳᨆՕʭʱɻ༓Ȧ࣬ဈٞᦇȵ߫ʚʶʳ⏏ጓɠᦉʛɹʓȴɴᖐΫȴʰହԂɐʳ౅໬ʢȡʳ⏏ςɴ᳋ɹᩄȵ޽Ȧᮒ࡚ɴ᝔ʼ
71ˏ˻ˏ͹ˏ̶͠ˡ‫ݖܩ‬቉ድǵፀ 3 ‫ش‬ʶʳ⏏౗ೣɫɻହᐨɹμ߈ᨆɴɄʶˁଝᑵʱࢊɓʳᏣȵȡʳȵ⏋ೣᠭീ‫ܩ‬ɫɻɄʶȵࡔ‫ܧ‬ɌɲȦ⏏ହԂ৿⏋ହɁɛϺȵອɹμɴԂɤɩᐨˁ‫ؚ܌‬ɐʳɛʠᏣˁਖឹɭɌɲȦȴʰɫȡʳ⏏ɛɞɌ⏋ອཉɹȡʳ‫ݭ‬ଘɫьᅎɐʳ‫غݭ‬ɻᏣˁɦȿʳɭȦȩ⏏ᐨɴȴȴʳ᳋ɻɐʏɩգᅎࢪᢒɭɇ
ʶʳȵ⏋ᕶсɹɛʠᐨᇿˁࢸɇȼɐʳɹȵࢅ౫ɫɻɲȼ⏋ςɴμ͹߂݀ɹ᳋ዛˁႰȩ⏏
(5) լᐨ⏃͐ͻͲ⏆ǵହᐨɭ‫ؾ‬෵ɴଁϺᅖਸȵੈχᇕອ‫ݖ‬ɫьᅎɐʳྫԍɫ⏋ьᅎ໬ȵ 2 ɦɴ߂աɇʶʳ⏏ʚɑ⏋͍˱Ͳໜೣ༑⏋ఌ༑ɹ༑ອ‫ݖ‬ɹᝠࣝȴʰμࣝɫ᝔ȩ༑Ɍլᐨɫȡʳ⏏ᖐȴʰᐨˁໜाԒɴହԂɌ⏋ອ༑ɴʚȴɓɩΥࡴ‫ך‬ᬚ༑άɐʳɹˁ৺ɤɩȴʰᐨˁளɁ⏋ʚɛΫ༑ᨆɴଖɤɩΥᦹɹ௭сˁᑵʱᦐɐ⏏ᧉ໻ਸɹ৔Ȧμ͹߂݀ɹ˱˓ዀ⏋̠͇̀ዀɲɰˁႰȩ⏏ʢȩ 1 ɦɻᐨˁЇ஘ȿʳеᒞȵΥࡴɴѳɛʶɩȦʳ౅໬ɫȡʳ⏏͍˱Ͳໜఌ༑ɹ༑ອ‫ݖ‬ɭ໡ʦ຾ɲɰɹ๬ອ‫ݖ‬Ȧɑʶɫʢ᝔ʼʶ⏋ହᐨɭᅵɲʱອᗳʦໍೠɲɰɹ᭭ࢀ႒ȵȡʳ‫ݭ‬ଘɫʢ᳋ȵ஛஁ɫȷʳɭȦȩգဣȵȡʳ⏏մᦚɹ౅໬ɭ‫ؾ‬෵ɴ஛஁ࢪᢒዛɻᧉ໻ਸɹ৔Ȧ˱˓ዀ⏋̠͇̀ዀɲɰɫȡʳȵ⏋៯ᒞ‫ݭ‬ଘȵອঈɴᦏȿʶɽঈᅊਸɹ৔Ȧ̝˻͚˗ዀ⏋˧˧ዀȵ⏋ʚɛ⏋޹౅ɴ៯ᒞɌɩᓂ౗ȡɁʶɽ޿ᬚʦᚚ౩ᚚಬɴ༎֪ɐʳ̠͇̀ዀ⏋̊˓ͩͲ̝˻͚˗ዀȵ஛஁ɇʶʳ⏏
(6) ‫ؘ‬ଝᐨ⏃̢˙Ͳ⏆⏃Ԡሎ 25⏆ǵଁϺߨਸɹʓȴᅖࡎ͹ߨࡎӻጣȵ 2 Ϻ 1 ᏼɫьȦ⏋ଁϺᅖਸȵьȩɄɭɻॕɫȡʳ⏏௭сɴɻ 2 ɦɹ౅໬ȵȡʳ⏏1 ɦɻɄʶˁ୫ɤɩອμˁմᦽɐʳ౅໬ɫ⏋ʓȴɻጓɠ๬ʚɤɛʚʚɫ
ອ༑ɭɭʢɴ༑άɐʳ᳋ˁ஛஁ɐʳ౅໬ɫȡʳ⏏Ȧɑʶɹ‫غݭ‬ʢ᣸‫ݭ‬ɹተȴɲ༓ଘȵ᧧ɽʶʳ⏏մᓔɻςɴΥಁᇕᏳ༑⏋຾ʦ໡ٞᦇɹະ࿭؅⏋৿ᓔɻᭇᮒ৿ɹ‫ވ‬ອɫ༎ᇍɴ༎֪ɐʳ᳋ᰖˁࢪᢒɭɌɩੈχᇕᏳ༑⏋͍˱Ͳໜఌ༑ʦ໡ɹࣳᦇ⏋Ȳʭʃɗʶʰɹ‫ݭ‬ଘɴɦʰɲʳΥಁᇕᏳ༑ɫ᝔ʼʶʳ⏏մᓔɹ౅໬ɫɻ Rasbora rubrodorsalis, Esomus metallicus, Lepidocephalichthys,
Anabas testudineus ፈɹࢸ͹μ݀᳋⏋৿ᓔɫɻ Rasbora aurotaenia, Xenentodon canciloides ፈɹʓȴɴ‫ވ‬ອಁɴϟఖᇕɴ༑άɌɩȷɛ Esomus metallicus ʦ Trichogaster trichopterus ፈɹΥಁᇕອ‫ݖ‬ɴᅊਖ਼ɐʳ᳋ᰖʢ஛ជɇʶʳ⏏

ৢ૯ 25ȁ஼਀࿌ȪΣͿϋȫ72࣬ᅒ͹߂ី͹ೠ‫׫⏧ؤ‬ᨆ͞ˠ˽ɹॶᩃᨆɴȲȿʳ᳋ᰖɹᅊਖ਼‫ݭ‬ଘգᅎɭз຦ɹྫ֝༎֪

ৢ૯ 26ȁన࿌ȪΐΛίȫ

ৢ૯ 27ȁన࿌ȪΠȜϋȫ(7) ᝬᐨ⏃˻̺̒⏋̜ͻͲ⏆⏃Ԡሎ 26⏋27⏆
ǵ̩̠̜͌͞Ͳ೴ɫɻ⏋Ʉɹྫԍɻ˻̺̒ɭ٧ɽʶ⏋͍˱Ͳໜఌ༑ɭໜᤄྲྀˁᐄʉΥಁᇕᏳ༑ɴ៯ᒞɇʶʳ⏏ᮒ࡚ɹɻɍʠ⏋߂ᩄɹᭇᮒɴʭʳອеɹਸ਼Ϋ౥ɫ͍˱Ͳໜఌ༑ɹອȵɄɄˁɭȲɌɩໜᤄྲྀɴ༑ʶᦉʞ⏏Ʉɹᐨɻɗɹ‫ݭ‬ଘɴጓɩʰʶɩȦʳೠɹఌ഼ɴ‫ࡴܔ‬ɇʶʳ⏏ᐨɹ៯ᒞȴʰ᳋ɹ஛஁ፈ⏋Υᦹɹс෌ɴɻភధɹଁϺᅖਸȵਃϟɐʳ⏏ࢪᢒ᳋ዛɻ Wallago attu ɹᅋ‫᧚׺‬Ϋᒴɞȵ⏋ʓȴɴཌྷკɇʶʳ˱˓ዀɲɰɹࢸ͹μ݀᳋ዛʢ̮ͻ̚˪ɹೳషɭɌɩգᅎɇʶ
ʳ⏏
ǵ̯Ͳ˹ˠ˗೴ʦ˵̠͌˵˓೴ɫɻ̜ͻͲɭ٧ɽʶʳ⏏ᮒ࡚ɹ‫ވ‬ອಁɴ៯ᒞɇʶʳȵ⏋Ʉʶʰɹ೴ɻ͍˱Ͳໜఌ༑ȴʰ᧙ᤂᮐɴȡʳɛʠ Wallago attu ɻ஛஁ɇʶɲȦ⏏ࢪᢒ᳋ዛɻ˱˓ዀ⏋̊˓ͩͲ̝˻͚˗ዀ⏋̯̠͇ͤ̀ዀ ʦ Anabas testudineus ɲɰɹࢸ͹μ݀᳋ዛɫȡʳ⏏
(8) ፊ⏃ดᒞȷ⏆Ʉɹྫԍɻ 3 ዛᰖ᠗ʠʰʶɛ⏏
8-1⏏ͦͻ̺⏃Ԡሎ 28⏆ǵᮒ࡚⏋‫ވ‬ອɌɩອ໗Ɍɛ͍˱Ͳໜೣ༑⏋ఌ༑⏋໡ፈɹੈχᇕອ‫ݖ‬ɹࣳᦇʦɗʶɴᦹᐚɐʳΥಁᇕອ‫ݖ‬ɹະ࿭؅ʦອ໗കɹ᳋ɹᦲʱ᧑ɴ៯ᒞɇʶʳ⏏ଁϺᅖਸɹьᅎɐʳྫԍɫ⏋μɴ᳋ˁ᠛ৄɐʳ᰹ɻԂʶɲȦ⏏μ͹߂݀ɹ Hemibagrus wyckioides, H. aff. nemurus⏋ಁɴɻ˱˓ዀɹ
Labeo ፈȵ஛஁ɇʶʳ⏏Channa striata ˁ஛஁ࢪᢒɭɐʳ᭬ɻ⏋ͦͻ̺ɹ೎ߡᨆˁዷຩμɴՏɌɩȲȷ⏋ɄɹμɴԂɤɛ᳋ȵዷຩˁُȭɲȦɛʠɴྔʶ๷ɵɹˁᬿȽ⏏˛̲ᅎɹͦͻ̺ɻੈχᇕອ‫ݖ‬ɹ͍˱Ͳໜఌ༑ɹाঈɴ‫ࡴܔ‬Ɍɩ៯ᒞɇʶ⏋ങГ޺ɹᨆՕɻ˵͞Ͳន⏃͠ͅ‫̢̲͠׋ݾ‬̚ͲᏙɹ‫ଁغ‬ᑥᐣɫ౗ೣɫɻᐨକɹᐨɭɌɩьᅎɇʶɩȦʳʢɹ⏆ɫȡʳ⏏
8-2⏏˵˓⏃Ԡሎ 29⏆ǵᮒ࡚⏋Υಁᇕອ‫ݖ‬ɫьᅎɇʶʳ⏏៯ᒞ‫ݭ‬ଘɻອᅒˁᑦȽࢸອᤉʦᏳ༑ɲɰɹ༑ʶɹȡʳ‫ݭ‬ଘɫȡʳ⏏ອ༑ɴЫɤɩΫάው֪ɐʳ᳋ᰖɹᅊ઴ˁգᅎɌɩȦʳ⏏ᦲ।ɻʢɰɌɹɦȦɛԂ‫ؤ‬ȵ 1 ɦɫ⏋ອ༑ɴࢪɌԂ‫ؤ‬ȵά༑ɴ‫ك‬ȼʭȩɴ៯ᒞɐʳȵ⏋μɴɻԂ‫ؤ‬ȵ 2 ɦȡʱ⏋ɗʶʰ73ˏ˻ˏ͹ˏ̶͠ˡ‫ݖܩ‬቉ድǵፀ 3 ‫ش‬ৢ૯ 28ȁᡫȪ‫؍‬౾̧ȫȪυȜίȫৢ૯ 29ȁᡫȪ‫؍‬౾̧ȫȪ΍ͼȫ

ৢ૯ 30ȁᡫȪ‫؍‬౾̧ȫȪ΍ͼΌΛίȫৢ૯ 31ȁᡫȪਸ౾̧ȫȪΠͽθͼͺϋȫɹ‫ك‬ȷȵɗʶəʶΫ༑͹ά༑‫ك‬ȷɫ⏋᧚Ϋ͹༑ά‫౅ؘ‬ɹ᳋ˁ஛஁ɫȷʳʢɹʢȡʳ⏏Rasbora
rubrodorsalis, Esomus metallicus, Trichopsis ɹЗᬚ⏋Trichogaster trichopterus, Anabas testudineus,
Channa striata, Clarias batrachus, Macrognathus siamensis ፈȵ஛஁ɇʶʳ⏏
8-3⏏˵˓˳̺̒⏃Ԡሎ 30⏆ǵɄʶɻ᳋ɫɻɲȼ⏋̤͇ˤ˛͡ᰖˁ஛஁ࢪᢒɭɐʳፊɫ⏋ᮒ࡚ɴᅤɲɰɹྂɤɛ‫ܩ‬ᯆɴԂ‫ؤ‬ɞȿՏɌɩ৿‫ץ‬ᨆˁ‫ݏ‬ʠ⏋μɴᔼɤɛˡ̢ʦ᳋⏋̮ͻ̚˪ˁԂʶɩ᠛ৄɐʳ⏏
(9) ፊ⏃ᑖᒞȷ⏆ǵ஛஁ࢪᢒɭɐʳ᳋ዛɴʭʱ০Ⴅȵᅵɲʱ⏋ʚɛ⏋៯ᒞ‫ݭ‬ଘʢ᧓ɤɩȦʳ⏏ɇʰɴ⏋ࢪᢒ᳋ዛɹ‫ؿ‬ዌˁЊɌɩɄʶʰˁ‫ך‬աɌɩȦʳ‫غݭ‬ȵ޽Ȧ⏏ȦɑʶʢଁϺᅖਸȵьᅎɐʳ⏏
9-1⏏̜˕͌˓ˏͲ⏃Ԡሎ 31⏆ǵɄʶɻ̊˗̠˧⏃ჼ‫ؿܩ‬ɻ˓ˏͲ⏆ɹʛˁ஛஁ɐʳፊɫȡʳ⏏ᮒ࡚⏋ϗ࡚ˁɭȲɌɩଁϺᅖਸȵжеອᅒʦະ࿭؅⏋ໜᤄྲྀፈ๬ອ‫ݖ‬ɹອཉɹ༓Ȧ‫ݭ‬ଘɴ៯ᒞɐʳ⏏Ʉɹዛᰖɻ̼̲ɹʭȩɲк݀ɫմᦽ৿ᦡˁᕶ‫ܧ‬ɴ᝔ȩɹɫ⏋ፊɴԂɤɛკ႒ˁᦣȵɇɲȦɛʠɴ‫؜‬ʱՏɌ‫ؤ‬ˁᯃ।ɴ74࣬ᅒ͹߂ី͹ೠ‫׫⏧ؤ‬ᨆ͞ˠ˽ɹॶᩃᨆɴȲȿʳ᳋ᰖɹᅊਖ਼‫ݭ‬ଘգᅎɭз຦ɹྫ֝༎֪

ᬏȼɌ⏋ɇʰɴɗɹᨆՕɴঈɹᬔɍɛԓፏႥɹጭɫᑂ˃ɞፏˁ୻Ԃɐʳ⏏ɄɹፏɻᙦɭɌɩล
ᔨɐʳɭ‫ؾ‬ಁɴɄɹ᳋ˁ᠛ৄɐʳɛʠɹ͊͊̀ᰖ⏋ᔼɤɛˡ̢ʦ᳋⏋̮ͻ̚˪ɭȦɤɛ᰹ˁԂʶʳᨆՕɴʢɲʳ⏏៯ᒞɐʳ᭬ɻ⏋ྫԍȵ࡭ԃɴອ໗ɌɲȦʭȩɴɐʳ⏏ፊɴԂɤɛკ႒ȵዷຩȴʰᨧᏬˁ٧ُɫȷɑɴྔʶ๷ɲɲȦʭȩɴɐʳɛʠɫȡʳ⏏
9-2⏏̜˕̮͌ͻ˱̜̒ǵɄʶɻ Hemibagrus aff. nemurus⏃ჼ‫ؿܩ‬ɻ̮ͻ˱̜̒⏆ˁ஛஁ࢪᢒɭɐʳፊɫȡʳ⏏ᮒ࡚ɴଁϺᅖਸȵ๬ອɹੈχᇕອ‫ݖ‬ɫȡʳ໡ɴ៯ᒞɐʳ⏏ፊɹ޺үɴᘽɹɦȦɛഘˁɇɌɩˡ̶͐͞ͻ˻͖ɌɩອঈɴໍʠʳɄɭʢȡʳ⏏᠛ৄɐʳɛʠɹ᰹ɭɌɩ̓͌˧ˏ͠ɹॾᛚˁԂʶʳ⏏
9-3⏏̜˕͌͞Ͳ⏃Ԡሎ 32⏆ǵᮒ࡚ɴଁϺᅖਸȵ༑ອ‫ݖ‬ɹੈχᇕອ‫ݖ‬ɹ͍˱Ͳໜఌ༑ɹाঈɴ៯ᒞɐʳ⏏ອ༑ɹ৑ȼɩອཉ
ɹȡʳཌȵ៯ᒞ‫ݭ‬ଘɭɌɩ᧧ɽʶʳɄɭȵ޽Ȧ⏏᳋ˁ᠛ৄɐʳɛʠɴɵȴʦ˹ͦˏ͠ᰖ⏋̓͌˧ˏ͠ɹॾᛚˁ᰹ɭɌɩμɴԂʶʳ⏏˱˓ዀɹࢸ݀ዛɫȡʳ Puntius, Opsarius, Mystacoleucus ɹЗᬚʦ Barbonymus, Hypsibarbus ɹЗᬚɹᗐ᳋ȵ஛஁ࢪᢒɭɲʳ⏏͞ͲɻɄʶʰɹᐵዌɫȡʳ⏏
(10) ፊ⏃ᘺɭɌ⏆⏃̅Ͳ⏆⏃Ԡሎ 33⏆ǵЈɹፊɭ᧓ɤɩԂ‫ؤ‬ɴʢɰɌȵɲȦ⏏ᮒ࡚ɴଁϺᅖਸȵΥಁᇕອ‫ݖ‬ɴᅊɍɛࢸອᤉɫ⏋μɴԂɤɛ᳋ȵອ༑ɹɛʠ᧚ʶɲȦʭȩɲਸ਼‫־‬ᨗɲ‫ݭ‬ଘɴ៯ᒞɐʳ⏏˵˓⏃8-2⏆ɭᰖаɌɛ᳋ዛȵ஛஁ɇʶʳ⏏
(11) ᳋ᕳᎢ⏃˽͌⏆⏃Ԡሎ 34⏆ǵϗ࡚ɴɲɤɩອȵৄȷɻɍʠ⏋ອཉȵ༓ȼɲɤɛະ࿭؅⏋຾⏋໡ɲɰɫьᅎɇʶʳ⏏ଁϺᅖਸɹʓȴ⏋ଁϺߨਸ⏋ᅖߨӻጣʢьᅎɐʳ⏏Ԃɤɛ᳋ˁΫᨆɴȡʳ‫ؤ‬ȴʰଝ摑ʛɫ‫؜‬ʱՏɐɭȦȩɄɹྫԍɹ႕ਸȴʰ⏋஛஁ࢪᢒɭɲʳ᳋ᰖɻȡʳዐ঎ɹ߂ȷɇɹ Channa striata, Clarias

ৢ૯ 32ȁᡫȪਸ౾̧ȫȪΠͽθρϋȫৢ૯ 33ȁᡫȪ၂̱͂ȫȪΕϋȫ
75

ˏ˻ˏ͹ˏ̶͠ˡ‫ݖܩ‬቉ድǵፀ 3 ‫ش‬batrachus, Anabas testudineus, Macrognathus siamensis ʦμ݀ɹ˱˓ዀ᳋ᰖɫȡʳɄɭȵ޽Ȧ⏏
(12) ാ༡ȿᎢ⏃ˡͻ⏆⏃Ԡሎ 35⏆ǵᮒ࡚⏋ςɴ໡ʦໜᤄྲྀȲʭʃ༑ʶɹᑃʦȴɲ͍˱Ͳໜఌ༑ɫଁϺᅖਸɴʭɤɩ᝔ʼʶʳ⏏߂ȷɲጭនɹɊʳႥɹྫԍɫ⏋Ʉʶˁອμɴໍʠɛ৿⏋ɗɹΫɴೠɹഘˁ޽ధዩʞ⏏ೠɹഘɴ᜜
ᮃɌɩȼʳ˛̲ᰖȲʭʃࢸ͹μ݀᳋ᰖ⏋ɗʶˁ஁ᰩɌɴᮃʚʳ̠͇̀ዀ⏋˧˧ዀ⏋̊˓ͩͲ̝
˻͚˗ዀɲɰɹ᳋ᰖȵ஛஁ࢪᢒɭɲʳ⏏
(13) ᒘ⏃Ԡሎ 36⏋37⏆ǵԂ‫ؤ‬ɴᙦȵȡʱ⏋μɴ᳋ȵԂʳɭɗʶȵᬔʚʳʭȩɲ෯ᦶɴɲɤɩȦʳ⏏Гάɹ 2 ዛᰖȵȡʳ⏏
13-1⏏̍͑Ͳ
ǵ̍͑Ͳɻጭនɫ⏋ᮒ࡚⏋༑ອɹੈχᇕອ‫ݖ‬ɫȡʳ͍˱Ͳໜೣ༑⏋ఌ༑ɹࣳ໤Ȧɴ⏋Ԃ‫ؤ‬ˁά༑үɴ‫ك‬ȿɩ៯ᒞɇʶʳ⏏ଁϺᅖਸȵьᅎɌ⏋߂݀ɹ Hemibagrus wyckioides, H. aff. nemurus, ˱˓ዀɹ Labeo ɹЗᬚፈȵ஛஁ࢪᢒ᳋ዛɭɲʳ⏏
13-2⏏̍͑Ͳ̮ͻ˱ͻǵ̍͑Ͳ̮ͻ˱ͻɻೠɹ഍ɫɫȷɛ፯Ⴅɹྫԍɫ⏋ϗ࡚ɴອཧȵᲞȼɲɤɛ᭬⏋Channa striata⏃ჼ‫⏆ͻ˱ͻ̮ؿܩ‬ȵອཧɹжȦອঈɹ˹˙̊͡ͻɴ᭯ʶʳɭȦȩᅊ઴ˁգᅎɌɩȦʳ⏏ϗ࡚ৢ૯ 34ȁ‫ٓݿ‬ᢌȪΑθȫ

ৢ૯ 35ȁড়૫̫ᢌȪ΃Ȝȫ

ৢ૯ 36ȁ᤭ȪΙλϋȫ

ৢ૯ 37ȁ᤭ȪΙλϋΩȜ΋Ȝȫ
76࣬ᅒ͹߂ី͹ೠ‫׫⏧ؤ‬ᨆ͞ˠ˽ɹॶᩃᨆɴȲȿʳ᳋ᰖɹᅊਖ਼‫ݭ‬ଘգᅎɭз຦ɹྫ֝༎֪

ɴଁϺᅖਸȵໜᤄྲྀɫьᅎɐʳ⏏
(14) ᩑʱ⏃Ԡሎ 38⏋39⏆
ǵᩑʱᩍˁьᅎɐʳྫ໬ɴɻГάɹ 2 ዛᰖȵ᠗ʠʰʶɛ⏏
14-1⏏̜̮̾̒˪
ǵ̜̮̾̒˪ɻᅖࡎӻጣȵࢃȴʰᦏȦᤂᮐɴȡʳΥಁᇕອ‫ݖ‬ɹອᅒʦະ࿭؅ɫᮒ࡚ɴ᝔ȩ⏏᰹ɭɌɩςɴ͊͊̀ˁьᅎɌ⏋Channa striata, Clarias batrachus, Anabas testudineus ʦ⏋ಁɭɌɩˡ˛͡ᰖˁ஛஁ɐʳ⏏
14-2⏏̴̜̾̒ˏ˪
ǵ̴̜̾̒ˏ˪ɻଁϺᅖਸȵ͍˱Ͳໜఌ༑ɹ༑ອɹੈχᇕອ‫ݖ‬ɫᮒ࡚ɴ᝔ȩ⏏᰹ɭɌɩ
10 cm еɹ̯̠͇ͤ̀ዀ᳋ᰖɹ༎᳋ˁ߂ȷɲᩑʱᩍɴᏣɫᐄʃɦȿ⏋߂݀ɹ̠͇̀ᰖɫȡʳ
Hemibagrus wyckioides ʦ H. aff. nemurus ˁ஛஁ɐʳ⏏
(15) ൯⏃͠ͻ⏆⏃Ԡሎ 40⏆ǵ͠ͻɻࢸ݀ɹ൯ɫ⏋Υಁᇕອ‫ݖ‬ɫȡʳᲞеອᅒɴ៯ᒞɇʶʳ⏏៯ᒞ‫ݭ‬ଘɻᅒᣴɌɹອȵ೎ᏺᇕɴᮃʚʳ೎άеອᅒɹອɹ༑ʶՏɌ‫ؤ‬ȵಔᦲɫȡʳȵ⏋μ᦭ɹອᅒɹᅮʦອᅒʭʱάଝɹΥಁᇕᏳ༑ɴ៯ᒞɇʶɩȦʳ‫غݭ‬ʢȡʳ⏏ᮒ࡚ȵɻɍʚʱ⏋ᅒඨȭȵᏺʼɤɛ৿ɴ៯ᒞɐʳ⏏៯ᒞс෌ɻᅒɹଘ೑ᓔɫȡʳଁϺᅖਸȵ‫׬‬Ⴖɫ᝔ȩȵ⏋ៀฐɹ߂ȷɲ‫غݭ‬ɻភధɹଁϺᅖਸȵԊ‫ؾ‬ɫɐʳ⏏ᮒ࡚ɹᬖࠃɭɭʢɴອᅒԒɫଁᬏ͹ᑢຂɌɛ᳋ȵᭇᮒɫ‫ވ‬ອɌɛ᭬⏋ອ༑ɴɹɤɩ
ው֪ɐʳᓃਸˁգᅎɌ⏋ອᅒȴʰˠͻ̫ͻ̶ͦͻɐʳອɭɭʢɴ༑άɐʳ᳋ˁ೎৿࣊ɴȡʳጭនɹޭɹᨆՕɴɛʠʳ⏏೎ᦏɻɄɹᨆՕȵ˵͞ͲនɹᝬɴɲɤɩȦʳ‫غݭ‬ʢ޽Ȧ⏏Ʉɹྫԍȵȡʳᅒɹᅮɻᦲ।ɹɗʶɴປʏᲞȼɲɤɩȲʱ⏋༑άɐʳʢɹɻɐʏɩྫკɇʶʳʭȩɴɲɤɩȦʳ⏏˱˓ዀɹ Esomus metallicus, Rasbora rubrodorsalis⏋̝˻͚˗ዀɹ Lepidocephalichthys ɹЗᬚ⏋̯̠͇ͤ̀ዀɹ Clarias batrachus⏋̜˰˗̠˧ዀɹ Macrognathus siamensis, ˥̨̓͠˗ˠዀ
ɹ Anabas testudineus⏋̊˓ͩͲ̝˻͚˗ዀɹ Channa striata ፈȵςɴ஛஁ɇʶʳ⏏

ৢ૯ 38ȁ೦ͤȪαΛΠΩ·ȫৢ૯ 39ȁ೦ͤȪαΛΠάͺ·ȫ77ˏ˻ˏ͹ˏ̶͠ˡ‫ݖܩ‬቉ድǵፀ 3 ‫ش‬ৢ૯ 41ȁକ̺̱Α΋ΛίȪ΃Εȫ

ৢ૯ 40ȁၴȪςȜȫ(16) ᎅɹࡎ⏃̶˙ˏ̒˪⏆ǵᎅɹࡎˁ‫׬‬ႶɫьᅎɌɩ᳋ˁ஛஁ɐʳɄɭɻɲȦȵ⏋ͦͻ̺ (8-1) ʦ̍͑Ͳ (13-1)⏋̅Ͳ (10)ɴ᳋ȵԂʳʭȩɴ᠛ࢷɐʳɛʠɴьᅎɇʶ⏋͠ͻ (15) ɹ̶˙Ͳ˽ɭɌɩʢਖɑᅎȦʰʶʳɹɫɄɄɫɻྫԍɹʁɭɦɭɌɩȡɁɛ⏏
(17) ᳋ࢊɓ‫⏆͌͡⏃ݭ‬ǵΥಁᇕອ‫⏋ݖ‬ςɴອᅒɴᦶʰʶʳԓ০ɹཉʛɫಁɭɌɩɗɹܑٞˁ໵ɹ‫ܤ‬ଝɫᲞȼɌɩȡʳ⏏ᦅс෌ɴਃϟɐʳଁϺᅖਸȵ᝔ȩ⏏ϗ࡚ɴອеȵάȵʱ⏋᳋ȵອɹȡʳɭɄʸɴᮃ‫غ‬ɐʳᅊ઴ˁգᅎɐʳ⏏͠ͻ (15) ɫ஛஁ɇʶʳ᳋ዛɭᰖаɌɛዛᰖȵࢪᢒɭɲʳ⏏
(18) ᬏനπᐨ⏃̋Ͳ̜Ͳ⏆ǵᬏɇ 3 m ʓɰɹጭጯɹӴɴሁ৹Ꮭ 15 cm ɹπȦങɹᐨȵɦȦɩȦʳ⏏ᮒ࡚ɹ޿ɴ⏋ଁϺᅖਸȵ̼̝̒͞˓̜ˁဣတɌɩ⏋Υಁᇕອ‫ݖ‬ɴȲȦɩˡ˛͡ᰖˁ஛஁ɐʳಁɴᅎȦʳ⏏
(19) ອɞɌ˽˱̺̒⏃ˡ̅⏆⏃Ԡሎ 41⏆ǵςɴϗ࡚ɹɻɍʠ⏋ଁϺᅖਸȵອᩄɹ޽ȼɲȦΥಁᇕອ‫ݖ‬ȴʰອˁȴȦՏɌɩ᳋ˁ஛஁ɐʳಁɴьᅎɐʳ⏏ȴȦՏɌɛອɻ˵ʹˑͲ (1) ɴȩȿɩ᳋ˁ‫ؚ܌‬ɐʳ⏏͠ͻ (15) ɫ஛஁ɇʶʳ᳋ዛɭᰖаɌɛዛᰖȵࢪᢒɭɲʳ⏏

4ȅაȁ݈
4.1ȁ‫ݿ‬႒͈୆௳ાਫ਼၌ဥ͂କ֖۪‫ޏ‬ǵആ‫ˏ˻ˏ׫‬ᡉ‫ܕ‬ɻཁອ᳋ȵዛధ͹ჼࡔᩄɭʢɴ޽ȦɄɭɫγᅠᇕɴሾʰʶɩȦʳ [Kottelat
1989]⏏ೣᠭീ‫ݖ‬ɫʢ޽ዛɹ᳋ᰖȵ᠗ʠʰʶ⏋ɗʶʰɹʓɭ˃ɰɻɇʚɊʚɲᅊਖ਼‫ݭ‬ଘˁው֪ɌɲȵʰգᅎɌɩȦɛ⏏ɗʶʰɹ᳋ᰖɻழᩆ‫ڃڧ‬ɭɌɩᩂឹɲ‫ڃ‬ᇿɫȡʳɹʛɲʰɑ⏋౗।ᅊ༎ɴȲȿʳᕶࢃ༥ᢹɴȴȴɐɄɭɹɫȷɲȦອᅋᣂ྇ɫȡʳ⏏Ʉɹʭȩɲ᳋ᰖɹᅊਖ਼‫ݭ‬ଘգᅎ
78࣬ᅒ͹߂ី͹ೠ‫׫⏧ؤ‬ᨆ͞ˠ˽ɹॶᩃᨆɴȲȿʳ᳋ᰖɹᅊਖ਼‫ݭ‬ଘգᅎɭз຦ɹྫ֝༎֪

ɴࢪਝɌɩ‫ݖܩ‬з຦ȵɇʚɊʚɲྫԍˁьᅎɌɩྫ֝༎֪ˁ᝔ɤɩȦʳɄɭɻ⏋‫ݖܩ‬з຦ȵɄɹ‫ݖܩ‬ɹ᳋ᰖɹᅊ઴ɴᬠɐʳሾᡫˁ‫מ‬Օɴ୫ɠ‫غ‬ʼɓɩȦʳ៷୞ɫʢȡʳ⏏Ʉɹʭȩɲϟёɻആ‫ˏ˻ˏ׫‬ɹЈɹ‫ݖܩ‬ɭʢԊᦲɐʳʢɹɫȡʳ␩ቄພ͹̝͞͡ 1990; Hubbel 1999; Claridge et al.
1997␭⏏ອᅋᣂ྇ɹ޽෵ਸȵɄʶʰɹ‫ܕ‬ɹయ‫׋‬ˁఌȭɩȦʳɹɻᬚ᧓Ȧɲȼ⏋͞ˠ˽ʢɗɹё޺ɫɻɲȦ⏏
ǵTaki [1978] ɻ͞ˠ˽ɹ͡ˏͲ̺̫͞Ͳȴʰʹˑ˛Ͳ̍͑Ͳᦏ᧿ʚɫɹ‫׌‬ᨆɭ̮˪́ȴʰ˱ͻͲྡٞᦇɹ೎‫׫‬ᨆɹ 2 ‫ݖܩ‬ɭആ‫ܩعˏ˻ˏ׫‬ɴȲȦɩ᳋ዛᏼଁˁៗ౩Ɍ⏋ɗʶʰˁປᥒɌɩᅊ
႒‫ܩ‬ჿ࡝ᇕᓒ࢘ˁɌɛ⏏ɇʰɴ⏋͞ˠ˽ȴʰ਄ʰʶɛ੻‫ݬ‬ˁμਕɭɌɩ͍˱Ͳໜμ༑ᨆɴᅊਖ਼ɐʳ᳋ᰖɹᅊਖ਼‫࡚⏋ି᧧ܩ‬፲ᇕው֪⏋ᰩਸ⏋ᅋ‫׺‬ೝɹ஠ࡴɴᬠɐʳ቉ድˁ᝔ɤɛ⏏ᅊਖ਼‫ܩ‬᧧ିɴᬠɌɩɻ⏋ອ‫ݖ‬ᄩ‫ޅ‬ˁ࣢‫ܩ‬ཙ༑⏋ж‫ܩ‬Ᏻ༑⏋ᑃʦȴɴ༑ʶʳອᤉˁ‫ي‬ʞ๬ອ‫⏋ݖ‬μ͹߂ៀฐɲж‫ܩ‬ໜाɹ 4 ɦɴ‫ך‬ՕɌ⏋ɗʶɴ༡ອ؅ˁ֓ȭɩ⏋ɗʶʰɹᅊਖ਼‫ܩ‬ɴՏჼɐʳ᳋ᰖˁ͠˽̜ˏ̺̒ɌɩȦʳ⏏࡚፲ᇕው֪ɹ቉ድɻж‫ܩ‬ໜाਸ᳋ᰖˁࢪᢒɭɌɩ 1 ॷᬚɹᐙᐚᠭീˁ᝔
Ȧ⏋ᠭീೝᬚμɐʏɩɹ೐ʢɌȼɻ 6 ‫܌‬ɹ᭧೐ᠭീɹȩɠ 5 ‫܌‬ГΫ஛ᮃɇʶɛዛᰖɭ⏋ᠭീೝᬚμ 3 Ͷ೐ГάʢɌȼɻ 6 ‫܌‬ɹ᭧೐ᠭീɹȩɠ 2 ‫ᦹ܌‬ᐚɌɩ஛ᮃɇʶɲȴɤɛዛᰖȵȡʳɄɭˁ᠗ʠ⏋մᓔˁࡴመਸ⏋৿ᓔˁው֪ਸ᳋ᰖɭࡴᒶɌɛ⏏ɇʰɴ⏋༡ອ؅ɴж‫ܩ‬ໜा‫ᧉ܌‬ਸ᳋ᰖɹʛɲʰɑ⏋Ᏻ༑ਸ⏋๬ອਸɽȴʱȴж‫ܩ‬ໜाࡴመਸ᳋ᰖʚɫʢȵѢԂɐʳɄɭˁተ᠗Ɍ
ɛ⏏Υ౅⏋ɄʶʰɹᠭീೝᬚˁɭȲɌɩкᬏ 3 cm ГάɹІ͹዗᳋ȵ஛ᮃɇʶɩȲʰɑ⏋ᅋ‫׺‬᝔֪ʦᅋ‫ݭ׺‬ଘ⏋І͹዗᳋ɹᔒଁ‫ݭ‬ଘȵ៨ᦚɇʶɩȦɲȦ⏏ೣ቉ድɴȲȦɩው֪ਸɹ᠗ʠʰʶɛ Paralaubuca typus, Wallago attu, Ompok bimaculatus, Hemibagrus aff. nemurus⏃Taki [1978] ɫɻMystus nemurus⏆ɻ Taki [1978] ɫɻж‫ܩ‬ໜाࡴመਸ᳋ᰖɴ‫ي‬ʚʶɩȦʳ⏏Ʉɹჿᅓɹʁɭɦɭ
Ɍɩ⏋ೣ቉ድȵΥಁᇕອ‫ݖ‬ɫɹᅋ‫ݭ׺‬ଘʦІ͹዗᳋ɹᔒଁ‫ݭ‬ଘɹᠭീɴᩂဣˁȲȦɛɄɭʢᓒ
ȭʰʶʳȵ⏋Ё৿ᠭീ‫ݖܩ‬ɹ᧓ȦˁᓒૅɌɲȵʰɇʰɲʳ቉ድȵਖឹɭਲ਼ʼʶʳ⏏
ǵRoberts [1993] Ȳʭʃ Roberts and Warren [1994] ɻ˱ͻͲྡٞᦇɹ೎‫׫‬ᨆ‫ݖܩ‬ɴᅊਖ਼ɐʳ᳋ᰖɹᅊ઴ˁ‫ݬ‬ٚɌɩȦʳ⏏І͹዗᳋ɴᬠɐʳ੻‫ݬ‬ɴʓɭ˃ɰʈʶʰʶɩȲʰɑ⏋ອ‫ݖ‬ᄩ‫ޅ‬ɻໜाɭອ໗കˁ߂աɐʳɴɭɰʚʱ⏋ᅊਖ਼‫ݭ‬ଘɹ᠎Ᏻɲ‫ך‬Օɻ᝔ʼʶɩȦɲȦ⏏ɌȴɌ⏋᳋ᰖɹው֪ˁ‫ع‬೐Ʌɭɴ៨ᥔɌɩȲʱ⏋ɗɹμɫᯃ‫ᧉ܌‬ਸ᳋ᰖɭ‫ᧉ܌‬ਸ᳋ᰖˁ‫ך‬ՕɌ⏋ɇʰɴ‫ᧉ܌‬ˁᑢຂɭᯃᑢຂɴ‫ך‬աɌɩȦʳ⏏ೣᠭീ‫ܩ‬ɴȲȦɩ⏋ੈχᇕອ‫ݖ‬ɫȡʳ͍˱Ͳໜೣ༑ʦ͍˱Ͳໜఌ
༑ɫ⏋᳋ᰖɹው֪ɭᅊ༎‫ر‬ɹᬠѥɴᬠʼʳ੻‫ݬ‬ˁɇʰɴ‫ؚ‬ᮃɐʳɄɭɻ⏋Ё৿ɹᠩᰍɫȡʳ⏏
ǵೣᠭീɹᐄഗ⏋ᠭീ‫ݖܩ‬ɹອᄩ‫ޅ‬ɻੈχᇕ͹Υಁᇕອ‫ݖ‬ˁ‫ي‬ʠɩ޽෵ਸɴ࢏ʛ⏋޽ȼɹ᳋ᰖ
ȵᅊ༎‫ر‬ɹ႕ࡴɹଁᬏຍ᭥ɫɗʶʰɹ‫ݭ‬ଘˁգᅎɌɩȦʳɄɭȵ౩ʰȴɴɲɤɛ⏏႕ɴᲞеອᅒɹ᳋ᰖᒴᮃɻዛ෯ଁɴȲȦɩႶᕶɹʢɹɫȡʱ⏋‫׬‬ɴੈχᇕອ‫ݖ‬ȴʰɹᦜԂɴʭʳʢɹɫɻɲȦȩȭɴ␩߂ីʓȴ 2001␭⏋Ʉɹ‫ݖܩ‬ɹ᳋ᰖᒴᮃɹᅊ႒޽෵ਸɹᲞɇɻ⏋Υಁᇕອ‫ݖ‬ɭੈχᇕອ‫ݖ‬ɭɹᬚ⏋ੈχᇕອ‫ݖ‬ɹໜᤄྲྀɭ͍˱Ͳໜɭɹᬚ⏋͍˱Ͳໜɹఌ༑ɭೣ༑ɹᬚˁΫάው֪
79ˏ˻ˏ͹ˏ̶͠ˡ‫ݖܩ‬቉ድǵፀ 3 ‫ش‬ɐʳɲɰ⏋ɗʶʰɹᅊ༎‫ر‬ɹμɫᅵɲɤɛອ‫ݖ‬ᄩ‫ޅ‬ɹᬚˁው֪ɐʳɄɭɴʭɤɩѳɛʶɩȦʳɭȦȩϟࡸɻ႕ɴ໶ᇿɇʶʳ⏏͍˱Ͳໜɴᅊਖ਼ɐʳ᳋ᰖɹው֪ɭΥಁᇕອ‫ݖ‬ɹᩂឹਸɻˡͲ̓˻ˏʦ͍˱Ͳ̊̚͡ɫೣᠭീɭᰖаɌɛᠭീˁࡸేɌɛᇐቄ [1967]⏋޽Ꮫ [1975] ʦ Taki [1978]ɹ቉ድଁഗɴȲȦɩʢ୭௎ɇʶɩȦʳ⏏᳋ᰖȵᅵɲɤɛອ‫ݖ‬ᄩ‫ޅ‬ɹᬚˁ৵਋ው֪ɐʳɄɭˁ‫خ‬ᔨɴɌɩȦʳɹɻ⏋‫׬‬ɴອɹ༑ʶȵᦹᐚɌɩȦʳɹɫɻɲȼ⏋ɗʶʰɹᬚɴ᳋ᰖɹው֪ˁᭀࢀ
ɐʳʭȩɲ᭭ࢀȵɲȦɭȦȩɄɭɫȡʳ⏏Ʉʶˁ‫[ ݻل‬1991] ɻᅊ႒࡝ᇕອ਌ᄩɭዌɌɩȦʳ⏏
ǵჼ‫ܧ‬ɹ౗ೣɫɻ⏋ੈχᇕອ‫ݖ‬ɻʢɠʸ˃ɹɄɭ⏋࡚፲ᇕɴੈχᇕອ‫ݖ‬ɭᦹᐚɐʳອᤉʦອᅒ
ɭȦɤɛΥಁᇕອ‫ݖ‬ȵ⏋ɇʚɊʚɲ᳋ᰖɴᅋ‫ݭ׺‬ଘʦІ͹዗᳋ɹᔒଁ‫ݭ‬ଘɭɌɩգᅎɇʶɩȦʳɭȦȩɄɭȴʰ⏋ɗʶʰɹ‫ݭ‬ଘȵཁອ᳋ᰖᒴᮃɹᅊ႒޽෵ਸɹᐣ୫ɴɻɛɐᩂឹɲ৲ւȵ໶ᇿɇʶɩȦʳ␩ຽऊ͹ᅒμ 1998; ᚹ‫ټ‬͹ອᢆ 1999; ႅᩃʓȴ 1988; ఱᚹ 1997; ఱᚹʓȴ 1988; ᦅ೴ᄩ‫ޅ‬఩Һ́Ͳ̊ͻ 1997␭⏏
ǵɌȴɌ⏋ჼ‫ܧ‬ɹ౗ೣɫɻ‫ݜ‬ᇻ఩Һϟ෌ɴʭʳ‫ݭܘ‬఩Һɹ᭬⏋ᅎອᤉɭகອᤉˁՕᮐɌ⏋ᅎອᤉɻ̮˓̺͞˓Ͳ‫׋‬ɇʶʳɭɭʢɴອᅒȴʰɹອɻᘺेɹᯃ।ɴ߂ȷȦகອᤉɴᘺɠʳɛʠ᳋ᰖȵອᅒˁգᅎɐʳɄɭȵέ‫خ‬ᔨɭɲɤɛ⏏ʚɛ⏋ອᅒʦᅎອᤉɴᦹᐚɐʳໜाɫɻᦅ෌ᅎ
‫؜‬ອ‫ݫ‬ʦቆᬿ‫ݨݫ‬ɹ៯ᒞɴʭʱ᳋ᰖɹው֪ȵᘿɌȼߺɁʰʶʳɄɭɭɲɤɛ⏏Ʉɹʭȩɴᅊ႒࡝ᇕອ਌ᄩȵᘿɌȼᭀࢀɇʶɛᐄഗ⏋఩Һ৿ɹອᅒʦɗʶɴЊ᭦ɐʳᅎອᤉɫɻ౩ʰȴɴອᅒᅊ઴Ꮩɹ᳋ᰖɹᅊ႒޽෵ਸɻཝࢹɌɩȦʳ␩ຽऊ͹ᅒμ 1998; ጩ 1985; 2000; ႅᩃʓȴ 2001; ‫ݻل‬
1991; ᦅ೴ᄩ‫ޅ‬఩Һ́Ͳ̊ͻ 1997; ఱᚹ 1984; ᅒμ 1999␭⏏
ǵɇʰɴ⏋ᅎອᤉɹሁᐻ‫׋‬ʦ⏋˱Ͳ˪͠ͻ̜Ϊᯆᡸࣳ‫׋‬ȵɐɐʛ⏋ɗɹȩȭໜाఒѹɹ᭬⏋ໜআ఩๭ɴʭʳाঈɹॶܹ‫׋‬ɴЊ᭦Ɍɩဈɭཌȵ༥߉Ɍɛʱ⏋жອᤉȵሁᐻ‫׋‬ɌɛɄɭɲɰɴ
ʭʱ⏋ອᄩ‫ޅ‬ɹ޽෵ਸȵ߂ȷȼ༥߉Ɍɛ⏏ɄʶɴЫɤɩ᳋ᰖɹዛధ⏋҅кధȵɭʢɴཝࢹɌɛ␩‫ ݻل‬1991␭⏏
ǵೣᠭീ‫ݖܩ‬ɴȲȦɩɻ⏋᳋ᰖɴᬠɐʳᅊ႒࡝ᇕອ਌ᄩɴࢪɌɩੲ৯ᯱˁ‫ؖ‬ʖɐʭȩɲϺဤᇕ‫ۊ‬ဤȵʓɭ˃ɰ᝔ʼʶɩȦɲȦɄɭʢȡʱᖪ߫ɲႥ઴ɫȡʳɭȦȭʳ⏏ɌȴɌ⏋˰Ͳ˱˪೴ᦏ
᧿ɴȡʳੈχᇕᏳ༑ɫɻ⏋ဌྰᅎອɹɛʠɹᦅ෌ᅎ‫؜‬ອ‫ݫ‬ȵল៯ɇʶɛɄɭɴʭʱ᳋ɹው֪ȵ1⏆ߺɁʰʶɛɹɴ֓ȭɩ⏋ɗɹά༑үɫɻອȵྡྷᅥɌɩᄩ‫ޅ‬ȵੲ‫׋‬Ɍɩ᳋ȵɭʶɲȼɲɤɛ⏏

ǵЁ౗ɹ͞ˠ˽ɴȲȦɩ᳋ᰖᒴᮃɴ৯ᯱˁήȭʳϺဤᇕ‫ۊ‬ဤɻ⏋ྫ֝༎֪ˁաɴɐʶɽဌྰϟ෌ɭ̋͌ল៯ɫȡʳ⏏ɄɄ 10 ॷഀ͞ˠ˽ɫɻᎦɹ࡭ԃᕶᐌ᧒ଁɹɛʠ⏋ϗ࡚ɴʢဌྰˁɌɩ
Ꭶˁᅊᅋɐʳకፑˁᦽʠɩȷɛ⏏ೣᠭീ‫ݖܩ‬ɫʢ̩̠̜͌͞Ͳ೴ʦ˰Ͳ˱˪೴ɹٞᦇɫဌྰᅒ
ȵᦶଁɇʶɩȦɛ⏏ೣᠭീ‫ݖܩ‬ɫɹဌྰϟ෌ɫɻ⏋Ϋ៨Ɍɛʭȩɴ‫؜‬ອᇿᇕɹᦅ෌ᅎ‫ݨݫ‬ˁল៯ɐʳɄɭʢȡʳȵ⏋Ʉʶʭʱʢ͍˱Ͳໜఌ༑ʦៀฐɹ߂ȷɲ໡ȴʰளອͅͲ̺ˁьᅎɌɩဌ1⏆ᛚ౩੯ᅊລȴʰɹኽѵɴʭʳ⏏

80࣬ᅒ͹߂ី͹ೠ‫׫⏧ؤ‬ᨆ͞ˠ˽ɹॶᩃᨆɴȲȿʳ᳋ᰖɹᅊਖ਼‫ݭ‬ଘգᅎɭз຦ɹྫ֝༎֪

ྰᅎອˁተѳɌɩȦʳ‫غݭ‬ȵ޽Ȧ⏏Ʉɹဣɫອ‫ݖ‬ᄩ‫ޅ‬ʍɹ৯ᯱɻࢹɲȦɭȦȭʳ⏏ᅎອᤉɻሁ
ᐻᇕɫΪᯆ˱Ͳ˪͠ͻ̜ᡸࣳɞȵ⏋U ࡓྋɫɻɲȼ⏋θࣳɻᑃӃహɫȡʳ⏏ʚɛ⏋ອঈɴɻȦɑʶɹ‫ݭ‬ଘɫʢ໵ȵྊʚɤɩୈອඨ႒ɹᅊᔒɌɩȦʳᨆՕʢȡʱ⏋᳋ᰖɹᅊਖ਼ᄩ‫ޅ‬ɭȦȩဣȴʰ೎ੲɭɻਲ਼ʼʶɲȴɤɛ⏏ᅎອᤉȴʰᅒʍɹᘺेɻࢹɲȼ⏋஘ȿ༑Ɍɇʶɛອɻ೎৿ɴɻ‫ܤ‬ᒽ⏃Ꮼ஗ʱ⏆ɹອᤉˁɭȲɌɩΥಁᇕᏳ༑ɴகອɇʶɩȦʳ⏏ჼ‫⏋ܧ‬ೣᠭീ‫ݖܩ‬ɫ᝔ʼʶɩȦʳဌྰ০઴ɻᅊ႒࡝ᇕອ਌ᄩɹែဣɴɛɩɽ߂ȷɲ‫ک‬ᰍɻɲȦɭՠుɇʶɛ⏏
ǵဌྰᅒɻೣഀॵΫȵɤɩȦɛ‫ݭ‬ଘȵϗ࡚ɴȲȦɩԗʃླྀອɐʳ⏋ϺဤᇕɴᦶʰʶɛృɌȦΥಁᇕອ‫ݖ‬ɫȡʳ⏏2001 ॷ 3 ೐ 11 ౗⏋̩̠̜͌͞Ͳ೴ɹဌྰᅒɫ஛ᮃᠭീˁ᝔ɤɛɭɄʸȋ3⏏ᐄഗȍɹ፲ɫ៨Ɍɛʭȩɴ⏋˱˓ዀɹ Esomus metallicus ɹ዗᳋͹ॾ᳋͹ଁ᳋ɭ Rasbora
rubrodorsalis ɹॾ᳋͹ଁ᳋⏋̝˻͚˗ዀɹ Lepidocephalichthys sp. ɹଁ᳋⏋̊˓ͩͲ̝˻͚˗ዀ
ɹ Channa striata ɹ዗᳋͹ॾ᳋⏋ˠ˽̺̦ͦ͌˽ዀɹ Trichogaster trichopterus ɹᅋ‫׺‬আɭ዗᳋͹ॾ᳋⏋Trichopsis schalleri ɹॾ᳋͹ଁ᳋ȵ஛ᮃɇʶɛ⏏Lepidocephalichthys Г޺ɻॾ᳋⏋ȡʳȦɻᅋ‫׺‬আʦ዗᳋ȵ᠗ʠʰʶɛɄɭȴʰ⏋Ʉɹ‫ݭ‬ଘɫϗ࡚ɫʢɄʶʰɹዛᰖȵᑢຂɌɩȦʳɄɭɻ
ተࡸɫȡʳ⏏Ʉɹȩɠ⏋Trichogaster trichopterus ɭ Trichopsis schalleri ɻϗ࡚ɫʢ๬ອਸɹੈχᇕອ‫ݖ‬ɫᑢຂɌɩȦʳȵ⏋Esomus metallicus, Rasbora rubrodorsalis, Channa striata ɹ 3 ዛᰖɻᘿᓔʰɹᠭീɫɻ⏋ਃഀɹᅊਖ਼‫ݭ‬ଘɫɻϗ࡚ɴȲȿʳᑢຂˁተ᠗ɌɩȦɲȦ⏏ృɌȼօՏɇʶɛ
Υಁᇕອ‫ݖ‬ɴѢԂɌɛɄʶʰɹዛᰖɴᅊ༎‫ر‬ɹ޴‫׋‬ȵᅊɍɩȦʳɄɭɻ⏋Ϻဤᇕ‫ۊ‬ဤɭᅊ႒ɹ
ࢪਝɭȦȩဣȴʰᖂ٢ཉȦჼᢒɭȦȭʭȩ⏏
ǵʚɛ⏋Ʉɹϟࡸɻອᅋᣂ྇ɹ‫֓ވ‬ɭȦȩᯆȴʰʢ໶ᇿɇʶʳȵ⏋ᅥઝɌɲȼɩɻȦȿɲȦɄɭɻဌྰᅒɹᄩ‫ޅ‬ɫȡʳ⏏з຦ɹ᠌ɴʭʳɭဌྰᅒɴɌɩȴʰዝɹࢀᛚ⏋႕ɴ˪̼͐͠ˡ͍͌˹ Leptocorisa chinensis ȵ߂ᩄɴᇍᅊɐʳɄɭɴɲɤɛɛʠ⏋Ʉʶˁᱤ᭑ɐʳɛʠɴຏᛚջˁьᅎɐʳਖឹȵᅊɍɛɭȦȩ⏏ʚɛ⏋ᔋషˁత॒Ɍɛ৿ɴ᳋ȵ๷˃ɫ༝ȴʃΫȵʳɭȦȩ⏏ᰩషɭɌɩգᅎɇʶʳ᳋ᰖɻ‫࡝׋‬႒ᣒɴູയɇʶɩȦɲȦɄɭȵմர೼Иɫȡʳȵ⏋ອᅒɴᅊਖ਼ɐ
ʳ᳋ᰖˁອᅋᣂ྇ɭɌɩգᅎɐʳ᭬ɹᠩᰍɭɌɩЁ৿ᩂ߂ɲ໶ઝˁɻʰɤɩʪȼਖឹȵȡʳ⏏
ǵ͞ˠ˽ɴɭɤɩᮚ֐ɻ‫ܕ‬ɹᢧ྇ˁ਄ʳɛʠɹᩂឹɲ᯵ᇿɹʁɭɦɫȡʳ⏏ɗɹɛʠ⏋ჼ‫ܧ‬ȡʳʢɹɴ֓ȭɩЁ৿޽ȼɹ̋͌ˁল៯ɐʳ៟ᅙȵȡʳ␩஗ 1996␭⏏̋͌ল៯ɴЫȩᅊ઴ᣂ྇ʦ
‫ݖܩ‬з຦ɹᅊ༎ʍɹ৯ᯱɻ޽߂ɲʢɹȵȡʱ⏋ອᅋᣂ྇ɭྫ֝༎֪ɴ߂ȷɲ޴‫׋‬ȵᅊɍɩȦʳϟёʢ‫ݬ‬ٚɇʶɩȦʳ␩ೠ‫ ؤ‬1999␭⏏ೣᠭീ‫ݖܩ‬ɫɻჼ‫ܧ‬ɹɭɄʸ̋͌ȵࡔ‫ܧ‬ɌɩȦɲȦ⏏
ǵೣᠭീ‫ݖܩ‬ɹ⏋ʓȴɹ‫ݖܩ‬ɴʛʰʶɲȦႥ໥ɭɌɩɻ 2002 ॷʭʱ̫Ͳ̯˛ͲाΫ༑ɹ́ͅͲᦏ᧿ɫᩆɹ஛஗ȵᬖࠃɇʶ⏋ፀ 2 ຍ᭥ɭɌɩ᪅᩿࣢ɹᬖᇍȵϜࡴɇʶɩȦʳɄɭɫȡʳ⏏ᩆɹᏄ᪹໬ɻ˹ˏͲ‫غ׋‬႒ˁьᅎɐʳ౅໬ȵɭʰʶʳ [Oxiana 2001]⏏Ʉɹ౅໬ɴࢪɌ⏋ᅣ [2000]
ɻ̺͞Ͳ̜கອμɹ˹ˏͲ‫غ׋‬႒ȲʭʃঘɇȦμɹ˹ˏͲ⏋ອ᪃⏋᪅⏋᩹⏋ϩ᩹⏋ˡ̝͊˗͌ȵᄩ‫ޅ‬ʍ༑Տɐʳ‫ک‬ᰍˁ୭௎ɌɩȦʳ⏏̶ˑ̴͠Ͳɫɻࡸ᭬ɴɄʶʰɹ႒ᣒȵܑٞɴ༑Տɐ
81ˏ˻ˏ͹ˏ̶͠ˡ‫ݖܩ‬቉ድǵፀ 3 ‫ش‬ʳϟఖȵᅊɍɩȦʳ⏏ɇʰɴ⏋‫ݖܩ‬з຦ȵອ᪃ˁьᅎɌɩᩆˁᏄᑇɐʳ౅໬ˁ᝔ȩʭȩɴɲʶɽ⏋ံลອ᪃ɹᙜຩˁُɤɩံลອ᪃μນɴɲʳɽȴʱɫɻɲȼ⏋ᕶးᅠɫ೑ล‫׋‬Ɍɛອ᪃ȵ᳋ᰖˁ‫ي‬ʞᅊ႒ɴᙢዩɐʳϟ઴ȵᅊɍʳ‫خ‬ᔨਸɻᲞȦ⏏᪅ɹ஛‫؜‬໬ɭɌɩɻᮚៗ஛‫؜‬໬ȵ஛ᅎɇʶʳ⏏Ʉɹ౅໬ɻᩃ޺ɴ᩿ቄˁ‫ݡ‬ዩɌɩॕቘᨧˁΫȴʰత॒ɐʳɹɫ⏋‫ܩ‬άອʦໜाɴཉխ
ɲອᣒູയˁ୛ȼ‫׶‬᭝ਸȵȡʳ␩ᅣ 2000␭⏏Ʉʶʰɹᄩ‫کޅ‬ᰍɴᬠѥɐʳ႒ᣒɹΥᨆɻອѶᆖʦ˓̊˓˓̊˓ᆖɹ؅‫܍‬ɭԊᦲɌɩȦʳ⏏ɄʶʰɹᆖຩɻᲞ࿣঎ɴᅊ႒࿣ᑘɇʶɛ؅‫܍‬႒ᣒˁ೑ɐʳອᅋ႒ˁᰩɌɩᇍᆗɐʳϟёȵ޽Ȧ⏏Еɴ⏋̫Ͳ̯˛ͲाΫ༑ɫᲞ࿣঎ɴູയɇʶʳ᳋ᰖȵঃ‫ݖ‬ው֪᳋ዛɫȡʶɽ⏋ອᅋᣂ྇ˁ޽ᅎɐʳɄɹ‫ܕ‬ɫ⏋ঃȦ፴ܑɫᆖຩȵᇍᅊɐʳ‫خ‬ᔨਸʢ‫ࡴو‬ɫȷʳʢɹɫɻɲȦ⏏Ʉʶʰɹ႒ᣒɹ߃းອ‫ݖ‬ʍɹ༑Տɭອᅋᣂ྇ᩄɹ޴֪ˁ̢͐̊͠
Ͳ˫ɐʳɭɭʢɴ⏋Ʉɹອ‫ݖ‬ɴȲȿʳ᳋ᰖɹው֪᝔֪ˁ‫ي‬ʠɛᅊ઴ˁ౛ਸ਼ɴៗ౩ɐʳਖឹȵȡ
ʳ⏏
4.2ȁူ૓মު͈ຽ‫ऻ̷͈ࢗ͂ݞ‬ǵອᅋᣂ྇ˁᅋɐʳ‫ݭ‬ଘɻԊ೑‫ܩ‬ɫȡʱ⏋೴‫׬‬еɫᅎᇰณȵ៯ࡴɇʶɩȦʳ‫غݭ‬ȵʓɭ˃ɰɫ
ȡʳ⏏͞ˠ˽ɫɻฌᲞɹΥᅱжȦाᑉɴеᒞɐʳȋा೴ȍ⏋ɗʶɴᐚȼॶ‫ܩ‬ɹȋॶ‫ܩ‬೴ȍ⏋ɗɌɩॶ‫ܩ‬ɹΫଝɫεɴȡʳȋε೴ȍȵȡʱ⏋Ʉʶʰ 3 ɦɴ‫ך‬Օɇʶʳ೴Ǿȵ႒Ǿϯழ⏋ϯ౫⏋॑‫ݭ‬ɲɰɹᡉ༎֪ˁɭȲɌɩȡʳ႕ࡴɹ‫ݖܩ‬ɹμɫʁɭɦɹ̢̜̒͘ˁɦȼɤɩȦʳɹȵಔᦲɫ
2⏆Ʉɹ̢̜̒͘Ԓɫɻʓȴɹ೴Ϻʢ͡ͻ͡ɇȭࡪʶɽয়᠍‫ܩ‬ɹգᅎˁኧ๬ɇʶɩȦʳʼ
ȡʳ⏏

ȿɫɻɲȦ⏏ɲȴɴɻʚɤɛȼ೴ɭᑉɹɲȦϺᬚɴʢ஛‫؜‬ˁ᠗ʠɩȦʳ‫غݭ‬ʢȡʳ⏏ʚɛ⏋ອᅋᣂ྇ɹѳᡸˁᇿᇕɭɌɛྫ֝ៀըɹʓȴɴ⏋႕ࡴɹ֪႒ɴࢪɐʳȲɗʶ⏋ᏄᮠѵЖፈɹɇʚɊʚɲឹ‫܍‬ɴʭɤɩ⏋೴ᘺ‫׬‬еʢɌȼɻ‫׬ݖܩ‬еɫࡪʰʶɩȦʳៀըȵȡʱ⏋ᐄഗᇕɴອᅋᣂ
྇ˁѳԃɌɩȦʳɭȦȩ⏋‫ܧ‬ഀɹᣂ྇፬ჿ౅໬ȵࡔ‫ܧ‬ɌɩȦʳ [Iwata in press]⏏Ʉɹʭȩɲը঎ɹμɫ‫ݖܩ‬з຦ɻΫ៨ɹʭȩɲྫ֝༎֪ˁ᝔ɤɩȷɛ⏏
ǵ͞ˠ˽ɫɻЁ౗ᇵ˃ɴ᰸ຂϟ෌ȵࣙᬖɇʶɩȦʳ⏏᰸ຂϟ෌ɴᬠɌɩ⏋ჼಁဣɫɻዛᗈˁՏ
ᗼ˵˓̀ɴʚɫᔒଁɌ᢫ެɐʳ‫ܕ‬ጓɹ᰸᳋‫ݭ‬ɻࡔ‫ܧ‬ɌɲȦ⏏ɌȴɌ⏋᰸ຂ᳋ɹ៊᳋ˁᰲᔒɌ⏋ɗʶʰɹዛᗈᅊᅋˁ᝔ɤɩȦʳ́Ͳ̊ͻɻ˵ʹˎ̠˭̜̒ላɹ‫ͻ˝ʹ˪̮⏋غݭ‬೴ɹላጓዛᗈ
ᅊᅋ́Ͳ̊ͻɫȡʳ⏏຦ᬚɹ᰸ຂ́Ͳ̊ͻʢ޽ȼࡔ‫ܧ‬Ɍ⏋ɛɭȭɽ˰Ͳ˱˪ᦏ᧿ɹ຦ᬚ෌ᓔɻ
˱˓⏋̠˓̘͡ˑ̴͞ˏ⏋Barbonymus gonionotus⏋̩˪ͤͲ⏋Cirrhinus cirrhosus ˁ᰸ຂɌɩȲʱ⏋ᕶՕɹ᰸ຂ຾ɫዛᗈȵᅊᅋɫȷʳ˱˓⏋̠˓̘͡ˑ̴͞ˏ⏋Barbonymus gonionotus ɹ዗᳋ʦ៊᳋ˁٞᦇɹ೴຦ɴ᢫ެɌɩȦʳ⏏೎ᦏɫɻ̊˓ȴʰዛᗈȵᇵ˃ɴᥥԂɇʶɩȦʳ⏏‫ܩ‬ӯɹϺɻ‫ܕ‬ᅋʭʱʢ‫ڃ‬ᣒȵᖪȦɭȦȩ⏏
ǵ᰸ຂ຾ɭɌɩɻਃഀȴʰࡔ‫ܧ‬ɌɩȦʳ຾ɴ֓ȭɩ⏋೎ᦏɫɻ᧑ᤉʦဌྰᅎອᤉɹ఩Һɹɛʠ

2⏆ᛚ౩੯ᅊລȴʰɹኽѵɴʭʳ⏏

82

࣬ᅒ͹߂ី͹ೠ‫׫⏧ؤ‬ᨆ͞ˠ˽ɹॶᩃᨆɴȲȿʳ᳋ᰖɹᅊਖ਼‫ݭ‬ଘգᅎɭз຦ɹྫ֝༎֪

ɴ஗ʱάɁɩອȵྊʚɤɛ‫ݭ‬ଘˁգᅎɐʳϟёȵ‫ވ‬ȭɩȦʳ⏏ʚɛ⏋ဌྰɹɛʠɴɓȷ๬ʠɛ
ᨆՕˁ᰸ຂ຾ɭɐʳёʢ᠗ʠʰʶʳ⏏
ǵ೴ᘺɫɹ᰸ຂɻ҅Ϻͤ̾͡ɭ೴ɹԊ೑຾ˁьɤɛϟ෌ɭɴ߂աɇʶʳ⏏҅Ϻɫ᰸ຂˁ᝔ȩϺɻዛᗈȵᣘԂɫȷʳɴ᣸ʳᣂᅋˁ೑ɌɩȦʳ⏏೴ᘺɫ᰸ຂˁ᝔ȩ‫⏋غݭ‬ዛᗈˁ೴ɹᣂᩆɫᣘ
Ԃɐʳ‫غݭ‬ɭ⏋ɲ˃ʰȴɹອᅋ෌ஹ֖̺ͦ˻˙˪̜ɴʭʱዛᗈȵံషɫఌᐌɇʶʳ‫غݭ‬ɭȵȡʳ⏏ɗɌɩ⏋Ȧɑʶɹ˭ͻ˽ʢዛᗈȵఔ༑ɇʶʳອ‫ݖ‬ɫɻྫ֝༎֪ȵ؏ᩂɴៀըɇʶʳɄɭɴɲʳ⏏Ʉɹʭȩɲёɻೣᠭീ‫ݖܩ‬ɫɻ˵̠͌˵˓೴ɹ೴ᘺԊ೑຾ɫ᠗ʠʰʶɛ⏏͞ˠ̫ˠᦏҶɹ́ͅͲ೴ɫɻਃഀ೴຦ɹອᅋ႒஛஁‫ݭ‬ଘɞɤɛ‫ݖך‬ȵ᰸ຂ຾ɴɲɤɛɛʠ⏋з຦ɹ஛஁‫ݭ‬ଘȵ೼ИɹੲȦଘɴ޴ೃˁоәɲȼɇʶɛ⏏Ʉɹ˭ͻ˽ɻ‫ܕ‬᭬ᇕ NGO ɹˠ̒˪˽̶ˎ͌ɹᢩ‫ܐ‬ࢪፑ̺ͦ˻˙˪̜ɴʭʳʢɹɫȡʳ⏏
ǵჼ‫͞ܧ‬ˠ˽ɫ᰸ຂϟ෌ɹɇʚɊʚɲ᠅ʛȵ᝔ʼʶɩȦʳȵ⏋Ȧɑʶʢ๘ᎦʢɌȼɻ౗ೣȵςࢷɌɩȦʳ [Modadugu et al. 2000]⏏ɗɌɩ⏋᰸ຂϟ෌ȵᬖࠃɇʶʳɭዛᗈʦɗʶˁ᰸ຂɐʳ຾ɴ౩ተɲ҅Ϻᇕଘ೑ณȵࢷԂɇʶʳɄɭɴʭʱ⏋ɄʶʚɫɴɲȴɤɛகЈᇕը঎ȵᅊɍ⏋ਃഀʭʱࡔ‫ܧ‬ɌɩȦɛԊ೑຾ʦԊ‫ؾ‬ɹᅎᇰณȵ༥߉ɐʳ⏏৿ᓔɻ͞ˠ˽ɹᕶးᄩ‫ޅ‬ɹμɫᅵɲɤɛ
ጓ‫ܩ‬೼Иɴ࣎зɌɩȦʳϺǾȵԊࡔɐʳɛʠɴᬏಁᬚȴȿɩօʱȡɁʰʶɩȷɛ˹˽̘͌ɫȡʳ⏏ΫᦚɌɛʭȩɴ⏋॑‫ݭ‬ςᒶᇕែဣȴʰʛʶɽѝғɹɲȦࢸ͹μ݀ɹ᳋ᰖɫʢ⏋з຦ɹᕶࢃ༥ᢹɹɛʠɴɻᩂឹɲອᅋᣂ྇ɫȡʳ⏏Ё৿⏋᰸ຂϟ෌ɭਃഀɹը঎ˁθጓɇɓʳʭȩɲը঎Ϋɹू߅ȵਖឹɫȡʳ⏏
ǵ᰸ຂϟ෌ɴʭʳອ‫ݖ‬ᅊ઴Ꮩʍɹ৯ᯱɴɻ⏋ɇʰɴ๖ɹʭȩɲʢɹȵȡʳ⏏Гάɴɗɹёˁ୲Ɂʳ⏏
ǵ̊˓ɫᣱȷɩȦʳϟёɻᅊ႒࡝ᇕ͹ᅊ઴࡝ᇕɲែဣȴʰʛɩཉխɫȡʳ⏏̯̠͇ͤ̀ᰖɻ
̊˓ɫʢ‫إ‬ȼʭʱᰩషɭɌɩঃȼգᅎɇʶɩȦɛ⏏͞ˠ˽ᯊ٨৿⏋ౙ̅ᦹɹࢭᬐࢃȵ೎߂кᬏ
1.5 m ɴʢɲʳȵ٢ȵੲȦˏ̶͠ˡ؅ᅋɹ Clarias gariepinus ˁ͞ˠ˽‫ܕ‬ԒɴࢷԂɌ⏋᰸ຂˁᬖࠃɌɛ⏏᭰‫ܕ‬ɹ̊˓ɫɻ⏋٢ɻчȦȵଁᬏȵᧆȼɩᆖຩɴ৑Ȧ Clarias macrocephalus ɭɄɹዛˁϯᮊɇɓ⏋ʭʱ߂ȷȼɩ٢ɹчȦ᳋ዛˁօʳɛʠɹ᠅ʛɭɌɩ Clarias macrocephalus ɹᮇɴ
Clarias gariepinus ɹᮁˁϯᮊɇɓɩ᰸ຂዛᗈɭɌɛ⏏Ʉʶʰȵᩃ޺ɴᦣɁՏɌɩ‫ܧ‬ഀዛɹ̢̒
̍ˁߥȩɽȴʱɫɲȼ⏋‫ܧ‬ഀዛɭϯᨗɌɩዛȵग‫ޙ‬ɌɩɌʚɤɛ‫ݖܩ‬ȵȡʳ␩ໜᩃ 2001␭⏏
ǵ͞ˠ˽ɫʢᇵ˃ɴᩃ޺ఔ༑ɇʶɩȦʳ˱˓ɻ⏋ɐɫɴ‫ؾ‬෵ɲɄɭȵ᝔ʼʶɛȦȼɦȴɹ‫ܕ‬ɫɻ⏋ອᗳˁᰩࢀɌອɹ࿡঎ˁ‫֓ވ‬ɇɓʳɭȦɤɛອ‫ݖ‬ᄩ‫ޅ‬ɹੲ‫׋‬ˁৄȷᣱɄɐɽȴʱɫɲȼ⏋Ј᳋ዛɹជᰩɴȲȦɩʢཉխɲ৯ᯱˁ‫ؖ‬ʖɐʭȩɴɲʱ⏋ˠͻ˽̜͞͠ˏɹʭȩɴ૴ᣌᩆˁȴȿɩɄɹዛᰖɹᱤ᭑ˁᦽʠɩȦʳ‫ܕ‬ɐʰȡʳ [Baird 1999]⏏ɇʰɴෑጩɲёɭɌɩ⏋1970ॷБɴˏ̶͠ˡɹᰩష‫׶‬ลˁఛȩᇿᇕɫ FAO ɴʭʱ⏋Lates niloticus ȵೣഀɻɄɹ᳋ȵȦɲȴɤɛ̲˪̜͠ˏྲྀɴఔ༑ɇʶɛ⏏ɭɄʸȵ⏋Ʉɹ᳋ɻ 200 ዛˁᣳȭʳ‫ܧ‬ഀዛˁ‫ྙޙ‬ɇɓɩ
83ˏ˻ˏ͹ˏ̶͠ˡ‫ݖܩ‬቉ድǵፀ 3 ‫ش‬Ɍʚɤɛ␩ೠά 1998␭⏏˱˓ɭ‫ؾ‬෵ɴ͞ˠ˽ɫᩃ޺ఔ༑ɇʶɩȦʳ̠˓̘͡ˑ̴͞ˏʢγᅠ‫ع‬‫ܩ‬ɫᅊ઴Ꮩʍᢦɹ৯ᯱˁ‫ؖ‬ʖɐɄɭȵ૴ਥɇʶɩȦʳ⏏2001 ॷɴ IUCN (International Union for
the Conservation of Nature and Natural Resources) ɻ⏋ೣഀɹᕶးᄩ‫ޅ‬ȴʰաɹ‫ݭ‬ଘɴውʳɄɭɫᄩ‫ޅ‬ɴੲ৯ᯱˁήȭʳ 100 ዛɹ֪ඨ႒ˁ͠˽̜ˏ̺̒Ɍɛȵ˱˓⏋Lates niloticus Ȳʭʃ̠˓̘͡ˑ̴͞ˏɭ‫ࣚؾ‬ɹˡͩ˽͍̀ Oreochromis mossambicus ȵɄʶɴ‫ي‬ʚʶɩȦʳ␩೛౗ృᓬ 2001; Global invasive species database 2003␭⏏ɗʶəʶɹ‫ݖܩ‬ɫອ‫ݖ‬ᅊ઴Ꮩɹ෯ᦶȵ᧓ȩɹɫ⏋Ʉʶʰɹёˁሁஞ͍˱Ͳໜ༑‫ݖ‬ɴȡɩɻʠʳ៱ɴɻȦȴɲȦ⏏ɌȴɌ⏋Ȧȼʰ͍˱Ͳໜອ‫ݖ‬ȵঃ߂ɞȴʰɭȦɤɩ⏋ɗɹ৯ᯱȵᇖံɫɻɲȦɞʸȩ⏏Ё৿ᩂ߂ɲᬠਕˁʢɤɩ‫ܧ‬ഀອ‫ݖ‬ᅊ઴Ꮩʍɹ৯ᯱˁ̢͐̊͠Ͳ˫ɐʳਖឹȵȡʳ⏏
ǵ᳋ᰖˁఔ༑ɐʳɄɭɻᄩ‫ޅ‬ʦዛᰖ෯ଁɴ৯ᯱˁήȭʳɽȴʱɫɻɲȦ⏏ѳԃᅊ႒࡝ɹែဣ
ɫɻᅊ႒޽෵ਸɭȦȩɭȷ⏋ዛ෯ଁɹ޽෵ਸɻʢɠʸ˃‫ؾ‬ዛԒɹ᧨Шᇕ޽෵ਸɹѳԃˁʢɗɹែএɴȦʶɩȦʳ␩ᵠᢆ͹ሼ؅ 1996␭⏏ᅊ႒ɻ‫ؾ‬ዛɫʢ᧨Шᇕɴᅵɲɤɛ႕ਐˁ୫ɤɛȦȼɦȴɹᮃ܎ȴʰଁጓɌɩȦʳ⏏႕ࡴɹ᧨Шᇕ႕ਐˁ୫ɦᮃ܎ˁɗʶɭɻᅵɲɤɛ႕ਐˁ୫ɦ
ᮃ܎ɹμɴఔ༑ɐʳɭ⏋ɗʶʰȵϯᨗɌɩ߃းɴɻࡔ‫ܧ‬Ɍɲȴɤɛ᧨Шᇕ႕ਐˁ୫ɦᮃ܎ȵՏჼɐʳ⏏ɄʶɻϺဤɴʭɤɩᣱɄɇʶɛ᧨Шᇕɲఉϕɫȡʳ␩ᨙ໨ 1997␭⏏ჼ‫͞ܧ‬ˠ˽ɫɻ
Barbonymus gonionotus ɹఔ༑ȵ‫ܩع‬ɫᇵ˃ɴ᝔ʼʶɩȦʳȵ⏋Ʉɹ᳋ɻ͍˱Ͳɴೣഀȴʰᅊਖ਼ɌɩȦʳ᳋ɲɹɫ޺ഀ᳋ɫȡʳ˱˓ʦ̠˓̘͡ˑ̴͞ˏɹёɴປʏʶɽ‫ܧ‬ഀᅊ઴Ꮩʍɹ৯ᯱɻ
ࢹɲȦɭȦȭʳȵ⏋᧨Шᇕ޽෵ਸɹែဣȴʰʛʳɭ‫ک‬ᰍɹȡʳ᝔ဤɫȡʳ⏏ఔ༑ɐʳ‫غݭ‬ɻ‫ؾ‬ΥອᏙԒɫ᧨Шᇕ႕ਐȵᅵɲʰɲȦᮃ܎ˁ᧧ʉɄɭȵਖឹɫȡʳ⏏
ǵ߃းອᅋᣂ྇ɹཝࢹɴ֓ȭ⏋᰸ຂϟ෌ɹୟ߂ɴЫȩ᰸ຂ᳋ɹಔ‫ؖ‬ɴʭɤɩ⏋షჿˁᇫɴᇵʳ᭬ɴ⏋Υ‫י‬ɫɠʬȩɰȦȦ߂ȷɇɹ᳋⏋Ȧʼʪʳ̘ͻ̸̶͡ˑ̒˹͖ɹʛˁᩂែɐʳʭȩɲѝғ៎ȵ৔ʚʱ⏋ࢸ͹μ݀᳋ᰖɹᮛឹȵཝࢹɐʳɭɐʶɽ⏋ჼ‫ܧ‬᝔ʼʶɩȦʳྫ֝༎֪ʦᅊ෌Ȳʭʃᰩయ‫׋‬ɴʢ߂ȷɲ৯ᯱˁήȭʳɞʸȩ⏏ʚɛ⏋᰸ຂ෌ɴਃϟɐʳϺɭɗȩɫɲȦ‫ݖܩ‬з຦ɹᬚɴ҅Ϻଘ਄ፈɹဣɴȲȦɩᐁཕᇕൔेȵঃȵʳ‫خ‬ᔨਸʢȡʳ⏏Ё৿⏋߃းອᅋᣂ྇ɹ޴‫׋‬ɴ֓ȭɩ᰸ຂϟ෌ɹ֪‫ك‬ɴʢ໶ઝˁ‫ك‬ȿɩʪȼਖឹȵȡʳɞʸȩ⏏ɇʰɴ⏋͍˱ͲໜອᏙɴᅊ
ਖ਼ɌɩȦʳ᳋ᰖɹᅊ༎‫⏋ر‬ᅊ઴ɹៗ౩ˁ౛ਸ਼ɴ᝔Ȧ⏋ࢰഀɹອᅋᣂ྇ɹᣒᇕ͹ᩄᇕ޴‫׋‬ɴ‫׸‬এɴࢪਝɫȷʳ‫ݜ‬ኄᇕ੻‫ݬ‬ˁҺȭʳɄɭȵ೚ʚʶʳ⏏
৫ȁৃǵЁ‫܌‬ɹ቉ድɴѤࡶˁɻȴɤɩȦɛɞȦɛ͞ˠ˽‫ܕ‬ጓ߂࡝ക࡝ᨆ˽˪˱Ͳ́Ͳ ˵˓̠ͤͻ̜࡝ᨆᬏ⏋̜Ͳ͠ ˵͓̍͑̒˪ᦅ࡝ᨆտ࡝ᨆᬏ⏋LARReC ɹ˵˓̺̚͞ͻ̜ǵ̢͖͇̍͞ͻଘᬏ⏋ˡ̲ͅͲ ̶˝˗̊̓Ͳລ⏋˹Ͳ̊˗˝Ͳ ʹˑ͞ͅͲລ⏋ʚɛᠭീɴ‫ؾ‬᝔ɇʶɛ͞ˠ˽‫ܕ‬ጓ߂࡝ɹ˽̒˪̅ͲͅͲ ̺͠˪˵ͻລ⏋˵ʹˎ̜̒ລ⏋͞ˠ͌ˏລ⏋̊ͻ̠Ͳລ⏋LARReC ɹ̅Ͳ̸Ͳລ⏋̊ͻʹ˝Ͳລɴཉᡚɐʳ⏏ʚɛ⏋ᠭീลೳʦฌೣɹѳ፬ፈɴਉ‫֐ת‬ȦɛɞȦɛ LARReC ɹ˵˓̺̚͞ͻ̜ǵ̢͖͇̍͞ͻଘᬏ⏋ˡ̲ͅͲ ̶˝˗84࣬ᅒ͹߂ី͹ೠ‫׫⏧ؤ‬ᨆ͞ˠ˽ɹॶᩃᨆɴȲȿʳ᳋ᰖɹᅊਖ਼‫ݭ‬ଘգᅎɭз຦ɹྫ֝༎֪

̊̓Ͳລɴઢᡚɹઝˁᝠɐʳ⏏ʚɛ⏋ჼ‫ܩ‬ᠭീɹ᭬ɇʚɊʚɲ੻‫ݬ‬ˁரњɇʶɛ೴Ϻʁɭʱʁɭʱɴਉ኎
ᅕɌΫɁʳ⏏
ǵೣ቉ድɹลЦˁήȭɩȦɛɞȷ⏋ԃᯆᇕɴᠭീˁఌ୫Ɍ⏋ɇʚɊʚɲᯆɫ‫֐ת‬Ȧɛɞȼɭɭʢɴᢵᩂɲ
੻‫ݬ‬ˁήȭɩȼɞɇɤɛϴᨊ߂࡝߂࡝᭏ˏ˻ˏ͹ˏ̶͠ˡ‫ݖܩ‬቉ድ቉ድዀɹ‫إ‬ाχᮁ‫ؿ‬᠕ఠஐ⏋ቄᅒᏛᨀմఠஐ⏋ጭᅒಅϓ֖ఠஐ⏋ࢸ‫ܮ‬ঔψລ⏋ϴᨊ߂࡝ആ‫ˏ˻ˏ׫‬቉ድ́Ͳ̊ͻɹ࡫ᚹ‫ٮ‬ᮁ֖ఠஐ⏋ᛚ౩੯ᅊລ⏋ኜକ߂࡝ࢸകЭ‫֖ڈ‬ଝ⏋մϴᨊ߂࡝ᦅ࡝ᨆɹകᩁᗔລɴ؄ȼਉ኎ᅕɌΫɁʳ⏏
ǵೣ቉ድɻ̜̊͜ᢧ܎ 1999 ॷ঎቉ድ֖ଁ B ֖ଁᅱ‫ ش‬99B3-103ȋ͞ˠ˽ Bang hiang ा༑‫ݖ‬з຦ɹᅊ෌ɴȲȿʳᅊ઴ᣂ྇գᅎɴᬠɐʳ቉ድŅШᐏᇕయ‫׋‬ɹѳࡔ͹ᐙମɭᄩ‫ޅ‬ѳԃɹែဣȴʰŅȍ⏋̜̊͜ᢧ܎ 2001 ॷ঎቉ድ֖ଁ B ֖ଁᅱ‫ ش‬D01B3-130ȋ͞ˠ˽ Bang hiang ा༑‫ݖ‬з຦ɹᅊ෌ɴȲȿʳᅊ઴ᣂ྇գᅎɴᬠɐʳ቉ድŅ‫ܩܧ‬ɹሾਜ਼ɹ៨᪼ɭಔ‫ؖ‬ŅȍȲʭʃ⏋ዀ࡝቉ድᢹ‫ݜ‬ᇻ቉ድ (A)(2) ᠩᰍᅱ‫ ش‬10041071ȋᅵᅊ઴Ꮩஞ៘ɴ
ᬠʼʳϺ‫ؤ‬ው֪ɭᣂ྇գᅎ˹˽̘͌ɹ޴ᢟȍ⏃቉ድБᝠᓔ⏧‫إ‬ाχᮁ⏆⏋ዀ࡝቉ድᢹជ֖ᩆμ൒ᇕ቉ድ୞ဣ
০ଁ̺ͦ˫͌͞ᠩᰍᅱ‫ ش‬10CE2001ȋˏ˻ˏ͹ˏ̶͠ˡɴȲȿʳ‫ݖܩ‬ᑂଁ⏧؅݀͹޴ࢅ͹᥊ଁȍ⏋ॶଁ 14 ॷ঎ϴᨊ߂࡝ఠᔒ቉ድ୷ᖂᢧ܎ፀΥ‫ش‬ϟ෌༠޺༐᧛֖ଁ⏃ቁೝ⏆ɹ֖ଁᩆɹΥᨆˁ‫؝‬ȿɩ᝔ʼʶɛʢɹɫȡʳ⏏
֨ȁဥȁ໲ȁࡃȏ೛౗ృᓬȑ2001 ॷ 5 ೐ 14 ౗⏃ϴᨊႆ޹՘⏆ȋ˓Ͳ̾ͻ̋ͻ֪ඨ႒⏧100 ˁ୭ࡴ⏋ᡰٚ⏧‫ܕ‬᭬ѳᡸลᬠȍ
Baird, I. G. 1999. Fishing for Sustainability in the Mekong Basin, Watershed 4(3): 54-56.
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ຽऊѳᅖ͹ᅒμ‫߅ڒ‬ᑂ⏏1998⏏ȏອᦇᄩ‫ޅ‬ɹѳԃŅᅊ႒ᒴᮃɹែဣȴʰŅȑѵ࣢኏⏏
஗ǵ‫⏏׭‬1996⏏ȏ͍˱ͲໜŅᬖᇍɭᄩ‫ޅ‬Ņȑ‫إ‬Ёೆ᭏⏏
ᚹ‫ټ‬ᮂ౩͹ອᢆ๭Υ⏏1999⏏ȋ᳋ᰖɹᅊਖ਼‫ݭ‬ଘɭɌɩɹອᅒᄩ‫ޅ‬ȍȏཁອᅊ႒ɹѳԃᅊ઴࡝Ņ਋ӯᅊ઴࡝ɴ‫ك‬ȿɩŅȑѵ࣢኏⏋76-85⏏
Global invasive species database. 2003. 100 of the worst. http://www.invasives.org/database/welcom/⏃2003 ॷ 8 ೐ 31౗⏆⏏
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203-211⏏
ႅᩃǵѹ͹Ᏻᢆ‫ٮ‬༠͹ϥ‫ؤ‬ਜ਼Υ೙͹ᮾ໡ч౅⏏2001⏏ȋ‫ן‬ುा༑‫ݖ‬ɹ 3 ̊˓̺ɹອᅒᬚɫɹ᳋ᰖሂɹປᥒȍȏ᳋ᰖ࡝ᮊ᠖ȑ48: 19-25⏏
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24-27⏏
ೠάǵ໨⏏1998⏏ȋˏˡ͍ዀȍȏ౗ೣ֪႒߂ᇑዀ 6 ᳋ᰖȑॶՂ኏⏋100-101⏏
ໜᩃǵ‫׭‬ᑂ⏏2001⏏ȏആ‫ܒݭ॑ˏ˻ˏ׫‬᫾␩᳋ᢤᑂ␭ȑেయ‫⏏ݟ‬
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ᅣǵ౩ᨀ⏏2000⏏ȋᦽʞ᩿࣢ᬖᇍɭᑵʱᦐɇʶʳ᩿ࢀȍȏˏ˻ˏᄩ‫ޅ‬ᇐೆ 2000/01ȑആ໽ᐁཕృ‫ݬ‬኏⏋37-62⏏
ጩǵૠϠ⏏1985⏏ȋᦅ෌ອᤉɹ᳋ᰖѳᡸɴɦȦɩȍȏཁອ᳋ȑ11: 64-72⏏
␲␲␲␲⏏2000⏏ȋᅒ‫ܘ‬ɴɦȿʳࢸɇɲ᳋᧑ȍȏਝᅎᅊ઴ू࡝ȑ3: 231-234⏏
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Hubbel, D. 1999. Food for People: Natural Fisheries of the Mekong River, Watershed 4(3): 22-36.
ቄພሁ᧑͹˭̦˽⏋̝͞͡⏏1990⏏ȏ᳋ᨲɭ̠ͤ̀˹ɹ቉ድŅ͐Ͳ˽ͻͲ͹ˏ˻ˏɹᰩϟయ‫׋‬Ņȑ࣬໳ೆঊ⏏
࣬ᅒ౩χ⏏2002⏏ȋ͞ˠ˽ᦅ຦ɹ‫ܧ‬ഀሾᡫŅ˵̫̠˭̜̒ላɹ̠͇̀ྫŅȍȏၖय़ᦅ෌ȑ46⏋ա‫ش‬1: 83-84⏏
Iwata, A. in press. Aquatic Eco-resources Management and Its Changes in Laos. Kyoto: Kyoto Univ. Press.
Mekong River Commission. 2001. Fish Migration and Spawning ver. 1: Mekong Mainstream (CD-ROM). Phnom Penh:
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Modadugu, V. Gupta, Bounthong Saphakdy, Lieng Khamsivilay. 2000. Review of Aquaculture Support to Lao PDR
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ᦅ೴ᄩ‫ޅ‬఩Һ́Ͳ̊ͻ⏏1997⏏ȏᅊ႒ሂѳԃɴᨗૅɌɛᦅ෌ອգే៯ɹ఩Һଝ໬ȑᦅ೴ᄩ‫ޅ‬఩Һ́Ͳ̊ͻ⏏
߂ីѵে͹࣬ᅒ౩χ͹ೠ‫ؤ‬ᅓ᱗͹˽̒˪˱Ͳ́Ͳǵ˵˓̠ͤ˗˽͹˵˓̺̚͞ͻ˽ǵ̢͖͇̍͞ͻ⏏
2001⏏ȋ͞ˠ˽ɹ߃ອᅒɹྫ֝ɭ᳋ᰖɹᅊ༎‫ر‬ȍȏၖय़ᦅ෌ȑ45⏋ա‫ ش‬2: 33-34⏏
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␲␲␲␲⏏1997⏏ȋཁອ᳋ɹᑢຂ‫ݭ‬ଘɭɌɩɹΥಁᇕອ‫ݖ‬ȍᬏᅒᖾ‫ٮ‬͹Ᏻᢆ‫ٮ‬༠ᑂ⏋౗ೣ᳋ᰖ࡝Цᇺѹȏ౗ೣɹॕࢹཁອ᳋ɹჼႥɭᏙᐏѳࡔŅʭʛȵȭʶ౗ೣᅋཁອ᳋Ņȑᐶೆଗ⏋194-204⏏
ఱᚹૠ໠͹ႅᩃǵѹ͹ࢸ໨ᰒᮁ⏏1988⏏ȋཁອ᳋ɹອᅒٞᦇɴȲȿʳΥಁᇕອ‫ݖ‬ʍɹѢԂɭᅋ‫׺‬ȍȏ౗ᅊ઴Ц᠖ȑ38: 35-47⏏
ᨙ໨ǵྃ⏏1997⏏ȋཁອ᳋‫҅౅ܩ‬кᒴɹ᧨Шᇕ႕ਸɭᏙᐏѳࡔȍᬏᅒᖾ‫ٮ‬͹Ᏻᢆ‫ٮ‬༠ᑂ⏋౗ೣ᳋ᰖ࡝Цᇺѹȏ౗ೣɹॕࢹཁອ᳋ɹჼႥɭᏙᐏѳࡔŅʭʛȵȭʶ౗ೣᅋཁອ᳋Ņȑᐶೆଗ⏋218-227⏏
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޽Ꮫѳ১⏏1975⏏ȋ͍˱Ͳ̊̚͡ɹ᳋ሂɭԒອᯆྫ෌ȍȏആ‫ˏ˻ˏ׫‬቉ድȑ13: 146-160⏏
␲␲␲␲⏏1997⏏ȋ᳋ᢤȍȏϟԎആ‫ˏ˻ˏ׫‬Ņᰟ‫ܤ‬͹ᅊ઴͹ᄩ‫ޅ‬ȑেయ኏⏋164-165⏏
ᅒμ᧑౩⏏1999⏏ȋອᅒٞᦇɹອᄩ‫ޅ‬ɹ᧓Ȧȵ̝˻͚˗ɹՕ॒ɭᅊਖ਼ࢌ঎ɴ‫ؖ‬ʖɐ৯ᯱȍȏ᳋ᰖ࡝ᮊ᠖ȑ46:
75-81⏏
ᇐቄᖾΥ⏏1967⏏ȏˡͲ̓̚ˑˏɹອᅋϟ੻ǵ༠޺ອᅋ‫أ‬ೆ 11ȑ౗ೣອᅋᣂ྇ѳᡸ‫ת‬Ц⏏
Ϋᩃᥞড়͹‫ܮ‬ೣΥᅖ⏏1999⏏ȏ᳋ɹՕᰖɹ‫ܒ‬᫾ȑആ༠߂࡝ՏႆЦ⏏
ᵠᢆȦɑʛ͹ሼ؅਒Υ⏏1996⏏ȏѳԃᅊ઴࡝ԂᬐŅ᧨Шࡎȴʰಕ៎ʚɫŅȑయΥᐵ‫غ‬Տႆ⏏

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