..

fJ -lt~~±~~~·.:k· ;c..t

Transactions and Proceedingsof thePalaeontological Society of Japan:-Jew Series

No. 53

Palaeontological Society of japan
Apr. lOth, 1964

CONTENTS
TRANSACTIONS
Page

•163.

Upper Miocene foraminifera from the Kiyosumi Formation, Boso Peninsula.............................................................. Naoaki AoKI 163

4&l.

Diphyphyllum from ltoshiro, Fukui Prefecture, Central japan ............. .
. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Hisayoshi !Go 170

465.

Notes on a Cretaceous Nautiloid from Kyushu ........................... .
. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Tatsuro MA TSUIIIOTO and Masahisa AMANO 173

466.

Carboniferous Conodonts from the Omi Limestone, Niigata Prefecture,
Central japan (Studies of Asian Condont:s. Pan I) ....................... .
. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Hisayoshi lc;o and Toshio KoiKE 179

PROCEEDINGS . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 194

President : Teiichi KOBA YASH!
Councillors:

Kiyoshi ASANO (Editor of " Fossils"). Riuji Et\DO. Haruyoshi FUJIIIIOTO,

Shoshiro HANZAWA, Tetsuro IlANA! (Secretary), Wataru HASHI~IOTO (Treasurer).
Kotora HATAI, lchiro HAYASAKA, Hiroshi OZAKI (Planning), Teiichi KOBAYASHI,
Tatsuro l\IATSUMOTO (Editor of Special Publications), .Masao MINATO, TokioSIIIKAl\IA (Membership), Fuyuji TAKAI (Editor), Ryuzo TORIYAMA
Assistant Secretary:

Takeo ICHIKAWA

All Communications relating to this journal should be addressed to the
PALAEONTOLOGICAL SOCIETY OF JAPAN
Geological Institute, Faculty of Science, University of Tokyo, Japan

Trans. Proc. Palaeont. Soc. Japan. N. S .. No. 53. pp. 163-169. pl. 25. Apr. 10. 196,1

46:~.

UPPER MIOCENE FORAMINIFERA FRO:\I THE
KIYOSUMI FORMATION. BOSO PENINSULA*NAOAKI AOKIInstitute of Geology and i'Vlineralogy. Tokyo University of Educationm~,'1~f.:;.lf'lifl®il· ~">_l·.fffiq1lfiil.t'fr1L.!R: tNNJIU:V!El'!!!ll!ltil · ~li'ilif7 *- :t'">.±~tM:IIt
fill. t.1~fdl~b·lt"C 11 fil!~iiettL-t-.:. tfifi s tl!'3:'..l<> <tr,'fifwr a{i*for permission to publish this paper andfor kind information on stratigraphy;
Dr. Takayasu Ucmo of the Universityof Tokyo for helpful suggestions; Dr.
Yokichi TAKAYANAGI of the Tohoku'Cniversity for kind permission to observe his collection from Nobori; Prof.
Wataru HASIIIli!OTO of the Tokyo University of Education for supervision andencouragements.

Introduction
Little has been studied on the foraminifera from the upper rvliocene rocksof the Boso Peninsula. because they arevery sparsely fossiliferous.
In 1962.
SArTO reported five planktonic species
from the Kiyosumi formation. including
G!obigerina nepenthes TODD and Globorotalia menardii miocenica PALMER, andhe considered that the middle horizonof the Kiyosumi formation is correlativewith the Venezuelan Globorotalia menardii menardii/Gfobigerina nepenthes Zone.
Previously. KOMATSU (1958 \IS! worked out the \liocene stratigraphy of thepeninsula and he examined the foraminiferal assemblages for zoning and correlation. Recently. the writer observed
the Komatsu's collection now stored in
our institute. besides some materials
newly obtained from the type sectionof the Kiyosumi. The purpose of thispaper is to record the several important
constituents of the Kiyosumi fauna andto describe five new species.
Acknowledgements are due to thefollowing persons: !vir. Naomiki Ko:\IATSU of the Teikoku Oil Company

Fauna
Collections were made from the three
sections taken along the small tributaries of the Obitsu River. in Kazusamachi. Kimitsu-gun. central part of the
Boso Peninsula. Despite of its meagre
population. about 110 benthonic species
were found in 35 samples. and the fauna
is dominantly composed of the follO\Ving species (in the order of abundance).
Urigerina proboscidea (Sc:Hw ACER)
Stilostomella ketienziensis (bliiZAKI)
Bulimina striata D'ORBIGKY
Nm1ion cf. parkerae UciiiO.. :Vodosaria" longiscata D'ORBIG:-Y
Rolmlus pseudorotulatus AsA:-<oBolit•inila quad1·itatera (SCH\\' AGEl~)"Eponides" umbonatus (HEL·ss). yar.
Cassidufina cf. subglobosa BRADY
Gyroidina komatsui AoKI. n. sp.

* J{eceived June 15, 1963: read at 8'1th
meeting of the Society at Sendai June 1. 1963

1 ~i3

Naoal•i AOKI

164

·Ur,igeriua cf. allilaeusis As A :-~oNonion pompi!ioides (FtCIITEL and ~Iot.t.)
Pullenia bulloides (o'ORBIGNY)
Sphaeroiditza bulloides (o'OR IIIGNY)
Cassidulhza sp.
l'lanulina wuellerstorfi (ScHWAGER)
Cibicides asa11oi MATSUl'AGA
Ortlzomorplzina obitsuensis AoKJ. n. sp.
Gyrnidina profunda AoKt. n. sp.
Fissurina miuima AoKt, n. sp.
Para}issurina kiyosumiensis AoKI, n. sp.
Lagena lzispit!a REuss
Lagena cf. asjJera REuss
Oolina me!o D'OR BIGNY
Pyrgo murrhina (SCIIW.\GEI~)
Cibicitles sp.
Gyroiclilla altiformis STEWAllT and STEWART".\'odosaria" tosta ScuwAGER
Bulimina rostra/a BRADY
8igmoili11a St'h/zmzberKeri S11. \'ESTR 1
Ortl:omorphina adt•ena (Cusu~tA:>: and\liocene to Pleistocene of Japan and it
is one of the dominantly occurred species in the Kiyosum1 formation.

Fissurina minima AOKT, n. sp.
Pl. 25, figs. la, b.

l'arojissurina sp. cf. P. dorbig11ymw(WIES:'<ER)

Test very small in size. unilocular,
compressed, almost circular in side view.
slightly longer than broad, lenticular inapertural view. thickness about two
thirds of the height of the test: marginsubacute, not keeled: wall smooth. finely
perforate: aperture a short slit on the
periphery. with a short entosolenian
tube. Length 0.15 mm .. width 0.14 rnm ..
thickness 0.09 mm. (holotype).
Holotype. Reg. no. 68001, from a riverside exposure (Sasa River). ca. 800 m.
south of Katakura, Kazusa-machi, Kimitsu-gun, Chiba Prefecture: sample no.
385. lower part of the Kiyosumi formation. upper Ivliocene. Collected by N.

Epistominella ·rotunda (HUSEZI"-IA andKG:\IATSU.

LAI;\.111'\G)

Epislomina elegans (o'ORBIGNY)

Fissuri11a amzecte11s BuRROWs and HoLLA:"DMARUHASI)

P!eurostome!la altermms SCHWAGER

Description of Species

Robulus pseudorotulatus AsA:>:O

This new species is characterized by
its very minute and circular test anda very short entosolenian tube.
Fissurina minima is common but restricted to the mudstone member of thelower part of the Kiyosumi formation.

Pl. 25. fig. 20.

Rolmlus pse~~tlorotulatus .-\.SAi'O. 1938, Tohok11
Uniz• .. Sci. Rep., ser. 2. 1•. 19. no. 2. p. 201.
pl. 25, figs. l. 3. -1: pl. 26. fig. 28: pl. 31.
tigs. 3, 6. - - AsA:'iO, 1951. Illust. Cat.
Japan. Tert. Forum .. pt. 15. p. 6. tfs. 28.
29.

The Kiyosumi specimens are smaller·(less than 2 mm. in diameter) than the
typical Pliocene form of this species andhave 7-9 chambers in the tina! whorl.
The large, slightly protruded umbilicus
and narrow peripheral keel are distinctive characters of this species.
It is widely known from the middle

Parafissurina kiyosumiensis
AOKI, n. sp.
Pl. 25, figs. 2a. b.

Test medium or large for the genus.
unilocular, compressed. nearly circularin outline, but apertural end somewhat
protruding roundly from the generaloutline of the test. thickness nearly two
thirds of the length, lenticular both incross section and in peripheral view:
periphery acute. with a narrow keel.
which broader at the lower half of the4rn.

Miocene Kiyosmni Foraminifera

test: wall smooth. translucent. finely
perforate; aperture eccentric, an arched
slit, parallel to the periphery, with along entosolenian tube extending to thebase of the interior. Length 0.26 mm ..
width 0.23 mm., thickness 0.16 mm. (holotype).
Holotype, Reg. no. 68002, from a riverside exposure (Kurotaki River). ca. 800 m.
north of Godai-batake. Kazusa-machi.
Kimitsu-gun. Chiba Prefecture: sample
no. 279, middle part of the Kiyosumi
formation. upper ~liocene. Collected by

N. 1\Q\IATSU.
This species is similar to "Fissurino
margi11ata (l\lo:>:TAGU)'' rarely found in
the Pliocene rocks of Japan. so it hasoften been confused with the latter
species.
Parajlssurina /~iyoszmtilmsis is frequently found in the lower half of the Kiyosumi formation.

Hi5

Test small in size. un iserial. straight.
subcylindrical, sides nearly parallel orslightly tapering: initial end roundedwith a button of clear shell material;chambers distinct. about six in numer.
appressed in the earlier. inflated andnearly as broad as high in the later
half: sutures horizontal. somewhat indistinct in the earlier. distinct and depressed later; wall finely perforate.
ornamented by numerous weak. longitudinal striations, becoming smooth atthe last few chambers; aperture terminal. central. a round pore in a thick andshort neck, without any tooth. Length
0.46 mm .. width ca. 0.16 mm. 'holotype).
Holotype. Reg. no. 68003. from a riYerside exposure 1Sasa River), ca. 300m
east of Katakura. Kazusa-machi. Kimitsu-gun. Chiba Prefecture: sample no.
361. upper part of the Kiyosumi formation, upper \liocene. Collected by :\.
KOi\lATSU.

·· .Vodosaria" longiscala o'ORBIGNY
Pl. 25. figs. 10. 11.

cVodosaria lrmgiscata o'ORTJIG:-IY, 18•16, Foram.
Foss. Rass. Tert. Vie1111e. p. 32. pl. 1. figs.
J0-12 (fide ELI.IS and MESST!\A, 1940 et
seq .. Catalogue of Foraminifera).-- As,\:"0. 1953, Toltoku U11i1'., lust (;eo/. Pal..
Short Pap., 110. 5. p. 8. pl. 1, figs. 20-22.

The specimens from the Vienna Basin
were re-studied by l\IARKS (1952. Cushman Found Foram. Res .. Contr., t•. 2. pt.
2, p. 45) and Nodosaria arzmdinea ScHWAGER. 1866, from Kar Nicobar was assumed to be a synonym of this species.
Fragments of this species are abundantly found in the Kiyosumi formation.

Ortlzomorphina. obitsuensis
AoKI. n. sp.

Pl. 25, fig. 3.

This species is characterized by smaller-sized test with weak longitudinal
striations on the surface and is easily
distinguishable from the species belonging to the genus Ortlwmorphina STAI:--1FORTH. 1952.
0. obitsuensis is frequently found in
the Kiyosumi formation and sporadicallyobserved in the Pliocene of the Boso
Peninsula.

Stilostomdla ketienziensis (ISHIZi\Kl)
Pl. 25, fig. '1.

E/lips0110dosaria kelienziensis lsHIZAKI. 1943.
Nat. Hist. Soc. Taiu•an, Trans .. 1'. 33, nos.
2-12-24.1. p. 684, tfs. 1. 6. ll. -As.'\NO.
1951. lllust. Cat. ]apall. Tert. Foram .. pt.
15. p. 28, tfs. 121, 122.

Several related forms to this Japanese
species had been described, but the Boso

Naoaki AOKI

lGG

specimens were compared only with the
Kechienji (Ketienzi) specimens and foundto be identical. The surface of thisspecies is smooth or ornamented with asingle or double rows of numerous. \Veak,longitudinal spines on the lower half ofeach chamber.
Fragments of this species are abundant throughout the Kiyosumi formation.
It is also common in the Pliocene formations of the Boso Peninsula.

25. --ASANO, 1958, Tohoku Uniz•.. Sci.
Rep .. ser. 2. 11. 24. p. 8, pl. 1. figs. 4. 7, 8.
--MARKS, 1.951. Cusltmau Found. Foram.
Res., Co11Jr., t•. 2. pt. 2, p. 58.

This species is abundant and one of
the most commonly occurred species in
the Kiyosumi formation.
Ut,igerina cf. akilaensis ASA:'\0
Pl. 25, Jig. 9.
Cf. Ut,igerina allifaensis AsAl\0, 1950, Il/ust.

Bolil•inita quadrilatera (SCIIW AGER)
Pl. 25, fig. 6.

Textularia quadrilatera SCHWAGER. 1866,
NoMra F:xped., Ceo/. Theil., 1•. 2. p. 253.
pl. 7, fig. 10.

This species is common in the middle
horizon of the Kiyosumi formation.
Tosaia /l(wzawai TAKA Y AC\AGI
Pl. 25. figs. 12a. b.

Tosaia lumzawai TAKAYANAGI. 1953. Tolwku
Unit-., /liSt. Ceo/. Pal., Short Pap .. no. 5,
p. 30. pl. 4, figs. 7. - - GALLITF.LLI, 1957.

U.S. Nat. Mus .. Hull.. no. 215. p. 151,
pl. 34. fig. 18.

This species is first described fromthe upper Miocene Nobori formation.,
Kochi Prefecture. It is yery rare andseveral specimens were found from six
samples of the Kiyosumi formation. All
are smaller and triserial throughout.
Hulimina striata D'ORillGNY
Pl. 25. figs. 14. 15.
Bulinina striata

o'ORAIC:'\Y.

CnsH:'-1.-\X and

1938, Cushman Lab. Foram. Res.,
Contr .. t'. J.J, 110 4. p. 90. (>1. 16. figs. 1-3--As.-\1'0. 1953. Tolloku Uniz·., !nsf. Geo/.
PARKER.

Pal.. S/rort Pap .. no. 5, p. 6, pl. 2. figs. 2·1.

Cat. Japan. Tert. Foram., pt. 2, p. 14. tfs.
60-62.

This form has highly raised or lamellated. longitudinal costae. which are discontinuous at the sutures. It is similar
to UtJigerina a!?itaensis ASA:-\0 widely
known from the Pliocene to Recent deposits of Japan, but it differs in having
a shorter test and deeply incised sutures.
lt may represent a variety of thisspecies.
It is frequently found in the Kiyosumi
formation and common in a sample
from the middle part of the Sasa Rivet·
section.
lh•igerina proboscidea SCHWAGER
Pl. 25. fig. 7. 8.

l.}rigerina proboscidea ScHWAGER.
1866.
Nm•ara Exped .. Ceo!. Thai!. z·. 2. p. 250.
pl. 7. fig. 96. --CUsll!\tA:--:. 1939, Ceo!.
Soc. Japan, four .. 1-'· 46, uo. 546, p. 151.
pl. 10, fig. 13. --Ct:sH:\JA:o-: and Tooo.
!941, Cushman Lab. Foram. Res., Contr., l'.
17. 110. 3. p. 73, pl. 17. fig. 9; pl. 19. figs.
3-9.

-

ASAJ'I:o. 1953, Tohoku U11iv ..

!nsf.

Ceo/. Pal., Short Pap .. 110. 5. p. 10. pl. 2.
figs. 29-31. --CTIIJJ. 1959. Osaka Jlus.
Sal. /list .. Bull., uo. 11. p. 26, pl. 7, lig.
5.

It is commonly and characteristicallyfound in the upper l\liocene rocks of468.

1lfiocene Kiyosum·i Fol'a.·minifera

japan and is the most common species
throughout the Kiyosumi formation.
Pleurostomella altemans Scuw ACER
PL 2S, fig. '-'·

Pleurostomella altemcms ScHWAGER, 1866.
.Vo!'ara E:rped .. Geol. Thai!. 1•. 2. p. 238,
pL 6, ligs. 79, 80. --CtJSHJ\IAX'. 1939,(,'1'01. Soc. Japan, jour .. /J. 46. no. fi·16. p.
152. pL 10. lig. 11.

This Kar Nicobar species is sporadically found in the Kiyosumi formation.
It has been recorded fro:n the Nobori
formation. Shikoku.
"J:'ponides" umbonatus REUSS. var.
Pl. 2:J. figs. 19a-c.

The periphery of Kiyosumi specimens
is narrowly rounded, differing from the
Pliocene ones of 1his species ( AsA1\0,
1951. Illust. Cat. Japan. Tert. Foram .. pt.
J.J, p. 12, tfs. 91, 921. however. the intermediate forms between the two areoften found in the present collection.
JV!nximum diameter of the largest specimen is 0.1 mm.
This variety is frequently found in
the Kiyosumi formation and is also observed very rarely in the lower Plioceneof the Boso Peninsula.

IG7

creasing moderately in size as added ;sutures distinct, slightly depressed, nearly radial. on the both sides; aperture
ventral. an elongate narrow slit, archedalong the base of the apertural faceextending toward the umbilical area;
wall thin, smooth, finely perforate.
Length 0.28 mm., width 0.22 mm., thickness 0.20 mm. (holotype).
Holotype, Reg. no. 68016, from a riverside exposure (Kurotaki River), ca. 700 m.
NW of Godai-batake. Kazusa-machi.
Kimitsu-gun, Chiba Prefecture: sample
no. 287. lower part of the Kiyosumi
formation, upper i\liocene. Collected by
N. Koro.IATSU.
The present species is similar to
Gyroidina nipponica lSIIIZAKI (19-14. Nat.
Ilist. Soc. Taiwan, Trwts., t•. 34, no. 244,
p. 102. pl. 3, fig. 3) from ~liyazaki, butdiffers in its larger number of chambersin the final whorl.
This species is named in honor of J'vlr.
Naomiki 1\.Cll'llATSU, oil-geologist. Teil<oku
Oil Company. in recognition of his workon the ::\liocene stratigraphy of the Boso
Peninsula.
Gyroidina lwmatsui is commonly foundin the lower half of the Sasa and Kurotaki J{iver sections of the Kiyosumi
formation.
Gyroidina profunda AOKI. n. sp.
Pl. 2:J. figs. 17a-c.

Gyroidina komalsui AoKr. n. sp.
Pl. 25, figs. 16a-c.

Test small. trochoid. ovate and slightly
lobulate in side view. thickness about
two thirds of the length, uneqally biconvex, dorsal surface almost flat or moreor less convex. ventral side roundly
convex; periphery broadly rounded;chambers 7 or 8 in the final whorl, in-

Test medium, trochoid, O\'ate in sideview. nearly equally biconvex, or ventral side more convex than the dorsal
side. consisting of 2Y:i or 3 whorls; periphery narrowly rounded and lobulate
yery slightly at the later part of the
test: chambers not inflated, about 7 in
the final whorl ; sutures somewhat indis·
tinct, oblique or nearly radial. slightly
curved. !lush on the dorsal side. radial,

168

Naoalci AOKIvery slightly depressed on the ventral
side; aperture ventral, an arched slit.
at the middle part of the basal marginof the apertural face: wall smooth. very
finely perforate. Length 0.44 nun .. width
0.38 mm .. thickness 0.28 mm. (holotype).
Holotype, Reg. no. 68017, from a river·side exposure (Sasa River), ca. 400 m.
south of I<atakura. Kazusa-machi. Kimi·
tsu-gun. Chiba Prefecture ; sample no.
368. middle part of the Kiyosumi for·mation. upper ~Iiocene. Collected by N.
KOMATSU.

This species is very similar to Gyroi·
dina gemma BANDY (1953. ]. Pal., v. 27.
p. 179, pl. 23, fig. 4) from off California,
but differs in having a more rounded andsmooth periphery and fewer chambers
in a whorl.
Gyroidina profunda is frequently foundin the Kiyosumi formation and also
common in the .. lowest Pliocene" rocksof the Boso Peninsula.

Nonion cf. parkerac Vcmo
Pl. 25, figs. 18a, b.
Cf. Nonion

parllerae Uc111o. 1960. Cushman
Fowul. Foram. Res .. Spec. l'u/Jl .. 110. 5. p.
60, pl. 4. Jigs. 9, 10.
Cf. No11io11 bnrleemms C"VILLIA~ISON). BANDY.
1961. Micropaleontology, 11. 7. no. 1. p. 16.
pl. 5, figs. 6.
Noni011 nicobarense CusH:o.!Al':. Isn!WADA. Ht·
GUCIII and KIKUCHI, ]apau. tlssoc. Petr.
Tech.. four .. v. 27, 110. 3. pl. opposite
p. 79, fig. 5.

Nonion zaaudamae bosoensis Ktl\\'ANO. 1962.
Res. lnst. Natur. Resources. 1\fisc. Rep.,
11os. 58-59, pl. 19, fig. 9. (uomen nudum).

The Boso specimens are not variablein morphology and are very closely simi·
lar to this Californian species. The
typical Recent form is more compressedwith a slightly more angular edge. but
the di ffcrence is not signi licant. This

Explanation of Plate 25(a, side or dorsal view: b, edge view: c. ventral view. unless otherwise noted.
Figs. 1, 3-6. 8-10. 12. 13, 15, 17. 18, 20, from Katakura: figs. 2, 7,
H. 16, 19, from Godai-batake: fig. 11. from Fudago.)
Fig.
Fig.

Fissurina minima AoKJ. n. sp., xl50. Holotype. Reg. no. 68001.
Pamfissurina kiyosumiensis AoKI. n. sp., x93. Holotype. Reg. no. ()8002.
Fig.
Ortlwmorphina obitsuensis Aoro, n. sp .. x69. Holotype. Reg. no. 68003.
Fig. 4.
Stilostomel/a ketienziensis (IsH IZAKI). X 69. Reg. no. 6800~.
Fig. 5.
Pleurostomella altertWIIS ScHWAGER, x59. Reg. no. 68005.
Fig. 6.
Bolit•inita quadrilatera (Sell wAGER). x 48. Reg. no. 68006.
Figs. 7. 8. Uvigerina proboscidea ScHWAGER. x53. Reg. nos. 68007, 68008.
Fig. 9.
U1•igeri11a cf. akitaeusis AsANO, x85. Reg. no. 68009.
Figs. 10, 11. "Nodosaria" longiscala o'ORllJG:-:v, x26. Reg. nos. 68010, 68011.
Fig. 12.
Tosaia llanzawai TAKAYt\NAGI, xlOO. Reg. no. 68012. b. basal view.
Fig. 13.
Cassidulina cf. subglofwsa BRADY, X 100. I{eg. no. 68013.
Figs. H. 15. Bulimi1w striata o'OJWIG!':Y. X 48. Reg. nos. 68011, 68015.
Fig. 16.
Gyroidina komatsui AOKI. n. sp., x93. Holotypc. Reg. no. 68016.
Gyroidina profunda AoKJ. n. sp .. x59. Holotype. Reg. no. 68017.
Fig. 17.
Fig. 18.
Nonion cf. parkerae Ucruo. x 69. Reg. no. 68018.
Fig. 19.
"E'pm1ides" umbonatus (R~:uss). var., x93. Reg. no. 68019.
Rolmlus pseudorolulatus i\sANO. X 29. Reg. no. 68020.
Fig. 20.
1.
2.
3.

AOK!:

Miocene Kiyosumi Foraminifera

Plate 25

4a3.

Miocene Kiyoswni Fomminifent

species is characterized by a narrowly
rounded periphery, limbate sutures and\·ery coarsely perforate wall. and it
resembles olnoma!in a umbilicatula HE·
IW:\·ALLEN and EARI.AC\D (1932. Dis·cot•e7)' Rep., 1•. 4, p. 426. pl. 14. figs. :1042) from south of the Falkland Islands.
and Nonio11 padanum PERCO:\IG (1952.
VII Cmwegno Naz. JHet. Petr.. Sez. 1.
Prestampia, p. 17. pl. 2. figs. I. 2) fromthe Italian 1\lio·Pliocene. The latter twopriorly established species have not been
examined for comparison, but the pre·sent species seems to be nearly identical.
It has already been recorded as ··No·nion 11icobarense CusH~IA:\" from Japan.
but the 1\.ar Nicobar species has a"rounded" periphery and fewer chambersin the final whorl (CUSI-1:\IAN. 1939. li. S.
Geo/. Sun•., l'ro/ Pap. 191. p. 17. pl. 4.
fig. 16).
The present species is congeneric with
Nautilus pompilioides FICHTEL and l\Ior.r..
1798. and may belong to the genus
Melonis I'vlo!\TFORT, 1808. emended by
VoLOSHINOVA(1958. Microfauna SSSR.

/.\', p. 148) or (;£welinonion HOFKER. 1951
( 1956. Skrifter Unit•. Zoo/. 11Jus. 1\.oben-

169

haven. Xf'·. p. 116).
Nonion cf. parlwrae is common in the

Kiyosumi formation and also in the
Pliocene of the Boso Peninsula.
Cassidulina cf. subglobosa 13RAilY
Pl. 25. fig. 13.
Cf. Cassidulina subglobosa BRADY, 1884. Rep.
Voy. Challenger. Zoo/., l'. 9, p. 430. pl. 54.
fig. 11.

Kiyosumi specimens referable to thiscosmopolitan species. have very small·
sized tests. The figured hypotype isthe largest specimen obtained.
It is frequent in the Kiyosumi forma·
tion.
References
KoMATSU. N. (1958) : Geology of the central
part of the Bnso Peninsula. Unpublishedmaster·s thesis. Tokyo Uni vcrsity of
Education.
SAITO, T. (1962) : Notes on Globigerina lte·
pentlws Tor>o. 1957. Palaeon/. Soc. Japan.
Trans. Proc., n. s .. 110. 18. p. :l31-342, pl.
51-52----·----

Boso
Godai·batake
FudagoKatakura
KazusaKechienji

JjJ~

m:a:tlll
:fl.

m:

K it
J: tt
f.li~~

Kimitsu
Kiyosumi
Kurotaki

t~I

~

m iG
.'1.\ ~

~obori~Obitsu
Sa sa

tj\

w:

•

Trans. Proc. Palaeont. Soc. Japan. X S., No. 53, pp. 170-172. Apr. 10. 1964

-164.

Dlf'HYPHYLLUM FROM ITOSHIRO. FUKUIPREFECTURE, CENTRAL JAPAN*
HISAYOSHI IGO
Institute of Geology and Mineralogy, Tokyo University of Education

fnl~Hlil.Tiff&E'lilff. Diphyphyllum : ffii:J.rlliH~IfNEi f-J·jJi;-=.)C~-t- r., ~i1oW,O)i~f't~li 1lU1liUIJ'!!!

mm~~~l:m:~-c. IHJ.§(tlj:--.i.:. .J: ">"(lilf1f ~~tt..:,0-IQI Diphyphyllum deliculmn MI:'\ATO
et KATO ::.jJi~C:.'J.~:bh-9.111llf!Jla-~H\ L.f-:O)"'CifC~L- '*f'ta-~~~ f..:cfii ~ 7-.. ~---------

foraminiferal faunule to be probably
Moscovian and the corals are about thesame in age or a little younger, but as
young as Artinskian".
Although his fusulinids are in very
poor state of preservation, his conclusion
as to the Moscovian age of them wasupheld by the subsequent discoveries of
the Fusulina Zone in the Hida massif(leo, 1956). A Caninia faunule was also
found in the upper part of the Fusulina
Zone below the Triticites Zone in the
Ichinotani Formation distributed in the
Fukuji district about 60 km northeast ofltoshiro (FUJIMOTO and lGO. 1958: IGO.
1961 ). Therefore, the geologica! age of
the Shimozaisho Limestone seems to be
1\loscovian and there is no possibility of
it extending upwards into the Permian.
As will be described later Diphyphyllwn
cfr. delicutum which was newly foundfrom the Lop of the Shimozaisho Lime·stone resembles Diphyphyllum delicutum
MINATO and KATO and is also related
to D. equiseptatum YARE and H.\YASAKA.
IJoth species were discriminated fromthe Nagaiwa Series of the Kitakami
massif. As already pointed out by 1\II·
NATO (1955) and MINATO and KATO
(1957) this type of Diphyphyllum is a longranging type and it persisted from late

KO!\ISHI i1951) worked out the strati·
graphy and paleontology of the fossil·
iferous limestone distributed narrowly
in the Itoshiro area. Ono County, Fukui
Prefecture. central japan. According tohim. the Paleozoic rocks distributed inthis area are surrounded by the so-called" Ilida i'vletamorphic Complex" and areslightly metamorphosed both dynamicallyand thermally. He subdivided the Paleo·zoic rocks as follows in descending order;
Shimozaisho alternation of clayslate andlimestone.......... 210m thick.
Shimozaisho Limestone ...... 250m thick.

KO'KISlll reported several poorly pre·1Jerved fusulinids and corals from theupper part of the Shimozaisho Limestone.
such as Eosclwbertella or Sdwbertella,
Fusulina ? sp. A. F. ? sp. B, cfr. F. /an·ceolata (LEE and CHEN), Caninia sp. A.
sp. nov. ?. C. cfr. mapingense. C. cfr.
simplisepiala and C. ? spp. indet.
Concerning the geological age of thisfauna he mentioned (KONISHI, 1954, p. 12)
as follows " Through their comparison
with the species known from Asia andits adjaccnce. the writer contends the* Received June 12. 1963: read at the
Meeting of the Palaeontological Society ofjapan held in Sendai City. June I, 1963.

170

464.

!Jiphyphyllum from Itosh iro

Early Carboniferous to Early Permian.
Therefore, the present discovery of Diphyphyllum in the Shimozaisho Limestone is
not a strong positive suggestion of the
Moscovian age. however, the close similarity between this specimen and D.
deliwlum should not be neglected in a
discussion on the geologic age of thementioned limestone.

171

disposed. Diameter of corallite 7 to
10 mm in full growth. Septa of twoorders. rather short and slightly flexuous.
1\lajor septa 17 to 19 in number andattain to about one half to one third ofcorallite radius. J\linor septa alternatewith major septa. short and less than
one half length of major septa. Dissepiments in one row and concentricallyarranged.
In longitudinal section tabulae numerous, about 10 in distance of one millimeter. Inner tabulae essentially horizontal. rarely undulating and uniting
with not so clearly differentiated inclined
outer tabulae. Dissepimentarium narrow,usually in one row and rarely in two
rows. No axial structure observed.
Remarlls :-The present specimen closely resembles DiphyjJhyllum delicutuml\li:\ATO and KATO in corallite size, short
septa. septal number and arrangementof tabulae. but D. tlelicutum possesses

Description of Fossil

Genus Diphyphyllum Lo:-~soALE, 18-15

Dipll)'Phyllum sp .. cfr. D. delicutum]'VI!NATO and KATO
Text·tigs. A-E.
cfr. Diphyphyflum delicutum }.IJ:":\TO and K.>.·

1957. Trans. Proc. Pal. Soc. japan.
N. S., No. 28. p. 137. texl·figs. A-C.

TO.

Corallum fasiculate and phaceloid.
Carallites cylindrical and rather closely

.,
A

B

E
D

0

10 MM

c
Text-figs. A-E. !>ip/iyplzyllmn sp., cfr. /J. delicu.fum MJNATo and
A. Longitudinal section, slight!}' modified: B. Longitudinal
section, partly tangential: C. Longitudinal section. slightly repro·
duced: D. Longitudinal section: E. Transverse section.
KATO:

Hi.sa.·uosh i JG<J

17:!

slightly more numerous tabulae in lhecorresponding distance. It is also related
to IJ. equiseptalum in many respects. D.
equiseptalum has more numerous septa
and slightly larger corallite. Owing to
the poor state of preservation. specificdetermination is reserved.
Occurrence :-!\ONISHI's locality (a) Valley, Itoshiro, Ono County, Fukui Prefecture. Reg. no. 21714.
References Cited
Ft.·JI~toTo.

H. and lGo. H. (1958): Strati·
graphic position of the corals in thelchinot ani Formation (Carhonif~~rous).
Fukuji district. Hid a massif. central
Japan. japan Acad. !'roc .. l'O/. 31. p. IS9163.
leo. H. (1956): On the Carboniferous andPermian of the Fukuji district. Hida
massif. with special reference to thefusulinid zones of the lchinotani Group.

Ceo/. Soc. Japan, ]our.. l'O/. 62. p. 217-240.
- - , (1961): 1\Iiddle Carboniferous corals
from the lchinotani Formation (Upper
Paleozoic corals from Fukuji, southeastern part of the llida massif. Part 3).
Pal. Soc. japan. Traus. Proc .. N. S .. no. 43,
p. 127-137. pis. 18. 19.
Kol'JSllt. K (1954) : Notes on the Moscovian(?) deposits in Itoshiromura. Fukui, japan. Ceo/. Soc. japan. ]our .. vo/. 60. p.
7-17, pl. I.
Mi:>:AT<>. M. (1955): japanese Carboniferous
and Permian corals. Hokkaido Univ .. Fac.
Sci., jour.. ser. IV. vol. 9, p. 1-202. p\s.
1-45.
- - and KATo, M. (1957): Two Carboni·ferous corals from the Kitakami Mountains. northeast Honshu. Japan. Pal. Soc.,
Japan. Trans. Proc.. N. S., 110. 28. p. 1371~2. text-figs. A-C.
YABE. H. and HAY:\SAKA, I. (1915):
Paleozoic corals from Japan. Korea and China.
Ill. Ceo!. Soc. Tol1yo. ]our .. vo/. 22. p.
127-1·12.

-------------------------------Fukujilchinotani
ltoshiro«J .11!!
-· / f't:riiltts

Ono County
Shimozaisho:;k~lfll!;-fi't:liJT

Trans. Proc. Palaeont. Soc. Japan. K S., ~o. 53, pp. 173-178. pl. 26. Apr. 10. 1964

465.

NOTES ON A CRETACEOUS NAUTILOID FROlvl KYUSHU*
T A TSURO ivlATSUl\IOTO
Department of Geology, Kyushu University
and

MASAHISA Ai'.. tANO
Department of Geology, Kumamoto University;JL!·}[ii(.:O)!~i!P.~~ ·j trHJ'iHt:t-i:

lfu!Je);:;!Jrl:~·=H'1f0) ~lllli.* i;lllfflilli~'Hf't- r{l'lffl~

niles ~iJP. L. t.:$~0)i'"Cf(,£0)~ftJJ'!,O) ~Hi:P~-c. ~~I?< "'ff$-'..; I-.-.::. 7(Gray so.

:.t) 11·~::> 1 jWO)

:to 5 tst.J:I. ·1tHt.:tJ:nQ.I?itt.:, ..:nLt Pa1·acymatoceras Fi\O)~f:i'fi~t~~T ~, ,.;f>:M\I!!IiJ~O)PA;;!.:;til\ P. as per CW~#I~H- ~ =- 7 :,~Jl'(:) .!:: JTIJJ;R1.:.. ~O)n}!iJiil'jl*'.:.m• · ,i';z.,t;·.~ z,,
ill!*CQ) Paracymatoceras 0) ~~fil.(i.:.Lt.: 0) lt;z..tJ:ts:l. •O)i.:::.><~L. i£;~~ T""-~"C~ 1.>, £1 G1~l:tl/Jtff ') -c:t-o ~. Cymatoceras semilobatus ( 1 './ Fffifti;Q)J:f!r,7 '" !::' 7 :.t) "?C. celtotiWII·

<

<

s

.t IJ"4.i.!l(;·j:iJJ1il.·, ;;J>:f!!in: P.asper O)[i'(,f::fffifl·· "?-'!"if!itf"Jt.:
Cymatoceras Q)~~fltJJtil•l?. iltWU: Vt:~1t.Lt.:tO)fl•l1 . .!:: <1.:.r
illlF-liffi7f!J~O) :b O)~.LJ-iHJii.>P. L 't: J: <lt~~Lt.cl ·.!::. I~W~-c~t.cl. ·,t~
i~ n!i · "}( YIJ !', ~ensis(!<k#IC7)-tl~=-7:.--)
fJlft~~~"->$~0)------

Introductory Notes

Nautilo:ds are rather rare in the Cre·taceous of Kyushu. Yi\JIE and Sllll\TIZU(1924: described Cymaloceras pseudoat/asfrom the middle part of the Senonian
llimenoura Croup. Amakusa. western
Kyushu. This was represented by asingle. well preserved specimen. In thesubsequent collection some fragmentary
specimens have been added to it fromthe Himenoura Group. but they are in·sutllcient for accurate identification. In
a recent collection from the Albian
Yatsushiro Formation, Kumamoto Prefecture. one of us (T. l\1.) has recognizeda small but interesting example of either
Jleminautilus or Paracenoceras, which* Received 7 November 1963: read JO
November 1963 at the 86th Meeting of the
Palaeontological Society of Japan.

*

awaits a full palaeontological description.
Whether Cymaloceras /sukushiense (Ko·BA YASIII. 1954) (em. KU\L\IEL, 1956). fromthe Tertiary sandstone exposed in thecity of Fukuoka. is a relict or a derived
fossil is a problem to be settled.
In this paper we describe an interest·
ing specimen of lhe cymatoceratinc nau·tiloid recently obtained from the middle
part (Lower Cenomanian) of the Goshonoura Group of Shishi-jima. Kagoshima
Prefecture.
Before entering into the description.
short remarks are to be gi vcn on thestratigraphic occurrence of the specimen(see Fig. 1).
The stratigraphic sequence of the Goshonoura Group in Shishi-jima has recently been described by one of us(A~IA~O. 1962. p. 2-9). As in the typearea of Goshonoura island (see l'viA TSU·

'l'atslli'O MATSUiVJOTO and Masahisa AMA.NO

17-l:VIOTO. 1960, p. 53), the group is divided
into three formations. lower, middle andupper. The middle division in Shishi·
jima consists of the following four mem·bers in ascending order:(a)Silty sandstone. about 180m. with
Nipponitrigonia.
(b) Conglomerate and coarse-sandstone.
about 110m.
(c) Sandstone, about 150 rn. rich in tri·
gonians.
(d) Siltstone. about 220m. with Grayso·
niles.

0

The nautiloid specimen was obtained
from the third member, which consists
primarily of fine· to medium-grainedsandstones. with intercalated dark grey
siltstone in the upper part. The sandstone may be occasionally conglomeratic
but the conglomerate does not form acontinuous bed. The sandstone is massive in some parts and well bedded inothers. This member is rich in mollusca,
especially in trigonians. The fossils arewell accumulated in some parts to form
fossiliferous beds and lenticular bodies

2KM

l===lo==...tFig. 1. Geological sketch map of Shishi-jima (adapted from A\tA:"iO. 1962).
I. II. III: Lower. i\.fiddle and Upper Formations of the Goshonouru Group: a-d:
Members a. d of the l'vliddle Goshonoura: IV: Himcnoura Group (Senonian):
T: Lower Tertiary: X : Type locality of !'amcymafocuas tmzgllaicum. Inset isa map of Kyushu. indicating the location of Shishi·jima at the center of a small
circle and also those of Fukuoka and Kumamoto with F and K.

46.'>.

Cretaceous l\la1tliloid from J(yush H

but ma ...- be scattered in other parts.
The mode of occurrence of the describednautiloid belongs to the latter category.
The fossil was preserved in a massive,
grey. fine to medium-grained sandstonewhich contains Atanthotri![onia pustulosa(NAGAO). A. ogawai (YEHARAJ and A.
dilapsa (YEIIARA\. It is fragmentaryand in its broken body-chamber small
gastropod shells (probably referable toTurritella) are embedded.
On the evidence of ammonites. Graysonites sp. cf. G. foun/aini YoUNG, Graysonites spp. indet. a and {3 (see MATSUMOTO. 1960), the fourth member is undoubtedly assigned to the Lower Cenomanian. We (Al\IANO et al. in MATst·MOTO, 1960) have, furthermore. concluded
that the l\1iddle Formation of the Goshonoura Group is wholl)r referable to theLower Cenomanian on the grounds of thestratigraphic relation and the contained
fossils. Therefore the Sandstone l'v1ember (c), in which the described nautiloid
was found. is regarded as a lower partof the Lower Cenomanian.
The described specimen is now preserved in the Type-specimen Room of
the Department of Geology, Kyushu University. Ful<uoka, with register number,GK. H640L

17!5

Paracymatoceras /zmghaicum sp. nov.
Pl. 26. figs. ln-b: text-figs. 2. 3

I!olotvpe :-GK. H6401, an incompletely
preserved body-chamber and the lastportion of n phragmocone, in which thetest is partly preserved.
Description :-The conch is involute andthe umbilicus is closed. The body chamber is subquadrate in section. slightly
broader than high. The measurements
show 61 mm in height and 68 mm inbreadth of the whorl, but the specimen
is slightly deformed. The venter isbroad with a shallow median groove.
The flank is gently inflated and abruptly
bent at the umbilical shoulder to forma nearly vertical umbilical walL The
maximum breadth is somewhat below
the mid-height.

Systematic Description
Family Nautilidae n'ORBIG:-IY, 1840
Subfamily Cymatoccratinae SI'ATH. 1927
Genus Pamcymatoceras SPATH, 1927

1)'pe-species :-Nautilus asJJcr (01'1'F.L)
ZITTEL. 1868.
Remarks:- This genus. established by
SPATH (1927, p. 21), has received goodcomments by i\'l!LI.ER and HARRIS (1945,
p. 9J and also by Ku:>.DIEL (1956. p. 427:.

Fig. 2.

Paracymatoceras tung!taicrm1

sp. nov.
llolotype, GK. HG-101. natural
cross section of the body-chamber. xl.

17G

Tatsuro MATSUMOTO ancl J[asahisa A.MANO

30 mm

Fig. 3.

Paracymatoceras t:mghaicum sp. nov. External
suture of the holotype. GK. H6401.

The ribbing is of cymatoceroid type.
consisting of numerous low ribs which
run almost radially, with slight flexuo·sity. on the flank. curved backward on
the Yentrolateral part and cross theventer with a strong backward sinus.
forming chevrons. On the internal mould
the ribs are impressed on the venterbut the impression is very weak on theflank. The ribs are close-set, being
separated by somewhat narrower inter·spaces.
The suture is sinuous. consisting of abroad and low ventral lobe, considerably
high, asymmetric, ventrolateral saddle. abroad and moderately deep lateral lobe.
a small saddle close to the umbilicus
and a shallow and small umbilical lobe.
The internal suture is not clearly exposed. The siphuncle crosses the septum
at a point somewhat dorsad from thecenter.
Remarks :-Although a single. imperfectly preserved specimen is at our
disposal, the observed characters are so
distinctive that the establishment of a
new species is justifiable as discussed
below.
Comparis011 :-In the sinuous suture.
subquadrate whorl section with a ventral
groove and cymatoceroid ribbing thisspecies is generally allied to Paracymalo·
ceras asper (OPPEL) (ZITTEL. 1868, p. 48,
pl. 3, fig. 1 ; l\liLLER and HARRIS. 19-15,
p. 9. pl. figs. 1-3; Kui\Ii\IEL, 1956, p. 427,
text-fig. 23G, pl. 19. figs. 1, 2), the type-species of Paracymatoceras SPATH. 1927.
Accordingly it is to be referred to thegenus Paracymatoceras.
Besides the type-species. which occurs
in the Tithonian of Europe, five distinct
species and a few indeterminable spec~esof Paracymatoceras have been known
from the Cretaceous of various areas in
the world (see KuMMEL, 1953, 1956). No
ventral groove is observed on all thesehitherto known Cretaceous species.
from which the present species is distinguished. 1\IILLER and HARRIS (1945)suggested that those without ventral
groove might be separated from thegroup of P. asper, but KU:\IMEL ( 1956)
holds that the genus Paracymatoceras isvariable in this respect. Anyhow it isinteresting to note that the present
species represents a Cretaceous exampleof the group of P. a,~per with a ventral
groove.
The suture of the present species is
not quite identical with that of P. asper.
In the latter species there is no \'entral
lobe and the dorsolateral saddle is larger
and more distant from the umbilicus
than in the present species.
Among the Cretaceous species, Paracy·
matoceras texanum (SHUMARD) 11860. p.
50). from the ·washita Group of Texas,
as described by i'viiLI.EH and HAr~RIS(1945, p. 10. pl. 3, figs. 1, 2: pl. 5. figs.
1-3: text-fig. 3) has deeper lateral lobe
than the present species and no ventral
lobe. It has a much more compressed

465.

C1·etaceous ~Vautiloid from Kyush·zt

conch than the present species.
As Ku~E\IEL (1956) has pointed out.
certoin species of Cymatoceras HYATT.
1884. have fairly sinuous sutures. being
trasi t·ional to Paracymatoceras. Thus.

Cymatoceras

semilobatus(SPE:-\(;LJ-:1~)(1910. p. 133 [9]. pl. 26 [11], fig .. (a, b),from the Lower Ootatoor Group of India.
has a lateral lobe which is similar inoutline to that of the present species.
but h:1s no ventral lobe and probably no
dorsolateral saddle. G)matoceras cenomanensis (SCHLUTER) (1876, p. 168. pl. '15.
figs. 1. 2), from the Cenomanian of Europe.
has a lateral lobe and a small dorso·
bteral lobe, which are similar to thoseof the present species. but has no ventral
lobe. Its conch is broader and has liner
ribs than ours. Cyma/oceras virgattwz
fSPE:-;GLER) (1910. p. 131, [7], pl. 26 [11],fig. 3a, b; pl. 27 [12]. fig. 7c). from theLower Ootatoor Group (Upper Albian)of India, has a ventral lobe as shallow
as that of the prcsen t species bu L its
ventrolateral saddle seems to be lower
than that of the latter. It has muchbroader whorl than ours and peculiar.
diverging ribs.
Occurrence :-Hirokino·tani. 1\.ashiwa·
guri, in the southern coastal area of
Shishi-jima, Kagoshima Prefecture: ap·
proximately 130.H'l8" East Long., 32'
15'52" North Lat.; in .4-canthotri)!nllia
bearing Sandstone 1\lember (the th:rdmember in ascending order) of thei\liddle Formation of the Goshonoura
Group. The specimen was collected by.M. A:\IA:-;o.

Concluding Remarks
A nautiloid recently obtained from the
middle part of the Goshonoura Group.
probably Lower Cenomanian, of Shishi·
jima, off the southwest coast of Kyushu,

177

represents a new species of Paracymato·
ceras. named P. tunghaicum. It has ashallow grooye along the mid-ventral line
like P. asper (OPPEL). the type-species,
from the Tithonian of Europe, being
dissimilar in this respect to other known
Cretaceous species of Paracymatoreras.
In suture it is similar to and somewhat
more sinuous than such species as C)·matoceras semilobatus (SPE:\GLER). fromthe Upper Albian of southern India.
\Vhether Paracyma/oceras tzmghaicum n.
sp. is a direct descendant of P. asper oris an offshoot from Cymatoceras by wayof a species with sinuous sutures may
be a question to be solved by further
discovery of various examples. especiallyfrom the Lower Cretaceous sequence.*

References Cited
lVl. (1962): The geologic history of
the Paleo·Shiranuhi bay in the Neo·Cre·
taceous Period. (Part :2 Regional Gco·logy).
E.umamo/o jour. Sci.. [H-1].
Ceo/.. Vol. 5, No. 1. pp. 1-36. 2 maps.
KoBAY,\SI!t. T. (1954): A new cymatoceratid
from the Palaeogene of northern Kyushu
in japan. japan. four. Ceo!. Ceogr .. Vol.
2·1, pp. 15-21. pl. 22.
Kc\·I:'.IEI.. B. (1953): Lower Cretaceous nau·tiloids from Texas. Breciora, :Vo. 19.
pp. 1-11. pis. 1. 2.
- - (1956): Post-Triassic nauriloid genera.
Bull. Jlus. Comp. Zoo/ .. lfan,ard Col/ ..
Vol. 114. No. 7. pp. 321-49,1. pis. 1-28.
J-1.\TSLT:\IOTO, T. (Hl60):
Graysonites (Cre·taccou,; ammonites) from Kyushu; with~otes on stratigraphy by M. A~IANO. T .
:\L\TSll\IOTO. H. OJ\,\I)A and H. OGUIH .
.1Iem. Fac. Sci .. Eyus/111 l.hziv .. [D]. Ceo/.,Fo/. 10. No. 1. pp. 1-11. pis. 6-8.
A:-.IA:"O,• Acknowledgements- We are indebted toDr. ltaru IL\Y.\:'.!1 and Dr. Tetsuro HANAI
who kindly helped this study. Miss !llisako
KJDo and JI.Iiss Tomoko l\·lt YAZAKJ assisted
us in preparing the manuscript.

178

Tatsuro M.-1TSUMOTO and Masahisa 1\MA.VO

MtLum, A. A. and HARRIS R. A. (1945):
North American Cymatoceratidae {Meso·
zoic Nautiloidea). jour. Paleont .. Vol. 19.
pp. 1-13. pis. l-16.
Sc!ILi'TEt~. C. (1876): Cephalopoden der ob.
eren deutschen Kreide. Pa/aeonfographica. Vol. 21. pt. 2. pp. 123-263 [1-144]
pis. 36-55.
SHU~IARD. B. F. (1860) : Descriptions of new
Cretaceous fossils from Texas. St. Louis
Acad. Sci. Trans .. Vol. 1. pp. 590-610(inaccessible here).
SPATH, L. F. (1927): Re\'ision of the Jurassic
cephalopod fauna of the Kachh (Cutch).
Alem. Gco{. Surv. India (Palaeont. Indica).
[N. S.]. Vol. 9. No. 2. pp. 1-84, pis. 1-7.
SPENGLER. E. (1910): Untersuchungen tiber
die stidindische Kreideformation. Die
Nautiliden und Belemniten des Trichino·

Goshonoura
1.ilJ l9r illi
Hirokino·tani !1\*(l).ft.

polydistrikts. Beitr. Palaeont. Osfeneicfl.
Ungarns u. Orients. Vol. 23. pp. 125-157[1-33]. pis. 26-29 [11-14].
YAtlE. H. and SHI:'-I!Zli S. {1924): A new
species of Nautilus. N. (Cymatoceras)
pseudo-atlas YABE and Sut~IIZU. from theupper Cretaceous of Amakusa. japan.
jour. Geo/. Geogr .. Vol. 3, No. 2. pp. 41!13, pl. 5.
ZITTEL, K. A. von (1868): PalaeontologischeStudien tiber die Grenzschichlen der Juraund Kreide-Formation im Gebiete der
Karpathen. Alpen und Apenninen. I. Abt·
hcil ung. Die Cephalopoden der Stramber·ger Schichlcn. Mus. K. Bayer. Staates
Palaeont. Mitt. (begonncn von OPPEl ..
fortgeselzt von ZITTEL). llol. 2, pp. 33ll8. pis. 1-24 in an atlas.

Kashiwaguri
Shishi·jima

Explanation of Plate 26
Fig. la-d. Paracymatoceras twzghaiwm sp. nov.
Ilololype. GK. H6,101. from a sandstone at Hirokino·tani. Kashiwaguri. southern coastof Shishi·jima. 1\agoshima Prefecture. Sandstone :Member. with Acantlzotrigonia. of
Middle Formation of the Goshonoura Group. Two lateral (a. b). ventral (c) and frontal(d) views. In Fig. ld the whorl section is not in a right position (cf. Text-fig. 2),
while lhe siphuncle is shown at the bottom. Figures are all of natural size.

MATSUMOTO and AMANO:

Cretaceous Nautiloid from Kyushu

Plate 26

Trans. Proc. Palaeont. Soc. Japan, 1\. S .. l\o. 53. pp. 179-193. pls. '27. 28. Apr. 10. 1964

4f5G.

CARBONIFEROUS CONODONTS FROlVI THE OMI LIMESTONE.
NIIGATA PREFECTURE. CENTRAL JAPAN(STUDIES OF ASIAN CONOD01\i"TS. PAHT I)*HISA YOSIII IGO and TOSHIO KOIKE
Institute of Geology and 1\linernlogy. Tokyo University of Education

Introduction and Acknowlegments
The Omi Limestone is distributed in
Nishikubiki County. Niigata Prefecture.
Central japan. It has been studied bymany workers. such as 1-IAYASAKA 11924).
KAWADi\ (1954) and FU.JITA (19581, but
no comprehensive paleontological study
has been published except for HAYA·
SAKA's classical work. We have been
studying various fossils from this huge
limestone mass and are very fortunate
to find an excellently preserved richconodont fauna in the lower part of thislimestone. There has been no report ofa rich conodont fauna in Japan except
for recent brief note of H.\ YASHI (1963).
V•i e studied this nevdy found conodont
fauna with keen interest. As the resultof the study we discriminated eleven
genera and sixteen species and became
to believe that this conodont fauna ap·
parently indicates Early Pennsylvanianor Late Namurian in age. This conclusionmust be noted about the geological ageof the \ower part of the Omi Limestonewhich was rather vague before our study ..
This paper is the first report of ourcontinuous work on conodonts from J apan and other Asian districts.
Acknowledgments are due to Drs.
Haruvoshi FUJIMOTO, professor emeritusof th~ Tokyo Uni vcrsity of Education,
Wataru HASHL\fOTO, professor of thesame university. Teiichi KOBAYASHI,
professor emeritus of the University ofTokyo and Teturo HA!\AI, assistant pro·fessor of the same uni\'ersity, for theirsuggestions and encouragements. During
the senior author's stay in Illinois he
was indebted to Dr. Charles COLI.INSON.
geologist of the Illinois State Geological
Survey, for his guidance for the conodont study. We also thank !vir. llidetoshi ToGASHI of the Omi Denka Co.
Ltd .. for his help to our fossil collectionin the field. A part of expenses of thiswork was defrayed from the Scientific
Expenditure Fund. Ministry of Education,
japanese Government.
Stratigraphic Summary

The Omi Limestone is a thick limestone mass ranging from the Early Car-• Received Jan.(}, 1964; read at the Annual
Meeting of the Society at Tokyo, Sep. 7. 1963.
l'i~lHisayosh·i /GO and 1imhio KoiKE

180

boniferous (Tournaisi:.m) to lJtc Middle
Permian in age. According to KAWADA(1954) and FUJ!TA (1958) this limestone-can be subclivide:-1 into five lithogenetic
units as follows in descending order.
Pe- .... . /'arajusulina and .Veoschu·agerinaZones
P 1 . . • . . . J>seudofusulina

ancl Pseudoscluca·
Zones
C, ...... Triticites Zone
C1 . . . . . . Fusulinella-Fusulina Zone
C1 . . . . . • Mi/lerella-Coral· Brachiopod Zone~:erina

The representing conodonts were col·lected from C1 of KAWADA and FUJITA.
The geological age of C1 was very obscure, but IIA 1 ASAKA once discriminatedthe following fossils, such as Amygdalop!zyllum giganteum (IIAYASAKA), Corzoenia ? omiensis (HAYASAK.-\). Axophyllum gracile HAYASAKA, Litlws/rolion
somaense YABE and 1-IAYASAKA, Sly!ido·
phy!lum sp., Chaetetes sp., Gigantoproductus edelburgensis (PHILLIPS), .l1•onia
cfr. aculeatus (:\L-\RTIN), Eclzinoconc!ws
punctatus var., Productus semireticulaius
1\IARTI:'\. Camarophoria globulina PHIL·
LIPS, 1\lartinia glabra detora PHILLIPS,
Reticula ria lin eat a ?\I A R TIN, Syringothyris
cuspidatus l\1ARTI:-<, Megistocrinus sp.,.-lctinocrinus aff. globrus PHILLIPS and-others.
Unfortunately precise stratigraphic
positions of these fossils are still obscure.
It seems to be necessary further inves·tigation of these fossils from Ct based
upon the present progressed paleontology. The occurrence of Syringothyris
cuspida/us from Ct indicates Tournaisian age, and Gigantoprodudus and·others are thought to be the Visean.
Recently SAKAGANII 1962) proved the·occurrence of Profw:.ulinella from theupper part of C1 and described several·Carbon:ferous bryozoa. KATO and NAK:\·
1\IVRA , 1£62) reported the occurrence of1

Eoasianites from the certain horizon ofc, and they briefly discussed about thegeological age of C1 . Judging fromthese evidence, KAWADA's Coral-Brachiopod Zone or FUJITA's Mi/lerellaCoral-Brachiopod Zone is thought to beequivalent to the Tournaisian to Bashikirian.
\-\'c collected many specimens of brachiopods. ammonites, corals and othersfrom about 100m. below of the SAKA·
GAl\lr's Profusulinella-!imestone.
They
are not throughly worked out, but wediscriminated the following ammonites
in addition to Eoasianiles sp., such as
Gastrioceras sp .. aff. G. reticulatus Yin
and Paralegoceras sp.. These ammonites
may indicate the Reticuloceras Zone ofwestern Europe. The Reticuloceras Zone
is the Upper Namurian and thought tobe equivalent to the Lower Pennsylvanian(l\looRE. 1948; ELIAS, 1956). Our conodont fauna was obtained from this ammonite-bearing limestone. and as will be
discussed later the geological age ofthis conodont fauna also indicates exactly same age with the preliminary
conclusion from these ammonites andbrachiopods.

Faunal Summary of Conodonts
The collections were made in the Nishiyama quarry of the Omi Denka Co. Ltd.
along the Omigawa River (Text-fig. 1).
Stratigraphic position of conodont-bearing bed is about 100m. below of the
SA KAGAl\II's l'rofusulinel/a·limestone.
\Ve have succeeded to separate numerous specimens of conodonts from grayto white sparry limestone by acetic acidtreatment and cliscriminatC'd the following species. namely :-Hindeoddla asialira leo and 1\oiKE. n. sp., H paradelicatula IGO and KOIKE, n. sp. H. sakagamii lGO and KOIKE. n. sp .. Synprionio-

-~!iii.

Carbonifemus Conodonts from Central Japan

1~1

N

1
QUARRY OF THE
OM! DENKA CO.

A

GONGfNYAMAITAGA~INE li;
~~~

~\

§
~
~
o
Text·fig. I.

0

IOOOm

Map ;;howhg the fossil lol:ality.

dina microdenla ELLISO~. S. collinsoni
lGo and KOII\E. n. sp., Fuprioniodina
dentata IGO and KoiKE. n. sp .. Ligonodina
hanaii IGO and KOIKE. n. sp .. Roundyn
subacordus IGU~"ELL). Lonchodina ? niP·ponica leo and KorKE. n. sp .. Ozarkodina
orientale lGO and KoiKE, n. sp .. Spatho·· gnathodus echigoensis IGO and KOIKE. n.
sp .. Idio~tnathodus togashii (Go and KOIKE.
n. sp., Streptognathodus jap01zicus (Go and
KOIKE. n. sp., S. extwnsus !GO and Koii<E.
n. sp .. Gnatlwdus opimus [Go and KOII\E.
n. sp. and Gnathodus sp.
llindeodella asiatica and fl. sakagamii
resemble H. armata STAt:FFER and PLUM·:'IIER which was described from the East~lountain Shale member. \Iineral Wells
formation. Texas. !/. paradelicatula isalso closely related with fl. delicatula
STAUFFER and Pr.tnl\IER from the Gra·
ford and i'VIineral \Veils formation in
Texas.
These three new species offfindeodella are very abundant through·out the collections. and they are related
to the mentioned American species whichrange from the Desmoinesian to 1vlissou·
rian. Owing to the fragmentary stateof these described foreign specimens.
detailed comparison seems to be clirTicult.
Thus we proposed new species hereinbased upon many complete specimens.
The genus Synprioniodina is known
from the Lower Devoni:w to Upper Car·
boniferous. but it is rather common in
the De\·onian and :\•lississippian rocks.
S. microdenta is very abundant in thetreated material and it is similar to theAmerican specimens which were de·scribed from the Cherokee Shale throughthe Deer Creek Limestone 1Eu.rso:-<.
1941) and also from the l'vliddle Dimple
formation 11\lorrowan) in Texas iELUSON
and GRAVES. 19H ~. ROIIDES '19;}2) de·
scribed Synprioniodina-assemblage from

182

llisayoshi !GO and Toshio Kol KE

the Pennsylvanian of Illinois and his
illustrated specimens quite agree with
our specimens. S. collinsoni is rather
rare, but having unique denticles and isvery distinct from other preYiousl~· de·scribed species.
£uprioniodina den/afa is not so com·
mon in our collections.
The genus
Euprionindina is rather long-ranged conodont and has been known from theLower Carboniferous to Triassic. Our
new species resembles E. Pulcella YOUNG·QUIST and DmvNs described from theLower Pennsylvanian of Iowa.
Ligonodina is also a long-ranged genus
of Coleodontidae and has been reported
from the Ordovician up to the Triassic
rocks. L. hanaii is a rare species inour collection and it may be relatedwith L. typa (GUNl\ELL) and also with
L. lexingtnnensis (GUNNELL) reported
from the Desmoinesian and I\ I issourian
of North America.
Roundya subacodus (Gtl!'\7-\ELL) is alsoa rare species in this fauna. Detailed
comparison between our forms and 1heAmerican ones seems to be difficult. butthey stand very close each other. According to Eu.JSO:'\ (1941), R. subacodus
ra11ges from the Desmoinesian throughVirgil ian.
Londwdina ? nipponica is also a rare
representative of this fauna. It resem·
bles L. ? ponderosa ELLISO:'\ in many
respects, if not entirely conspecific. EL·
J.JSON's species ranges from the Desmoi·
nesian to the Lower Permian in 1\lissouri and Kansas.
Ozarkodina orientale is a common spe·
cies in our fauna and is related to 0.
delicatula described by many ·workers
from the Kinkaid to various levels of
the Pennsylvanian in U.S. A. The l'vlississippian species of 0. compressa REXROAD described from Illinois is similar
to our 0. orientale.

Spathognathodus echignensis is a rather
common species in this fauna and it
slightly resembles S. minutus (ELLISO~)
discriminated from the Desmoinesian to
Big Rlue Series.
Idio~tnathodus lo;?ashii is a rather rarethroughout our collection.
This newspecies seems to be a primitiYe type
among this genus and resembles I. deli·t·atus GuN.!'\ELL persisting in the Desma·
inesian to Lower Virgilian in Kansasand Mis~ouri.
Two forms of Streptognathodus are
discriminated from this fauna. S. japonicus is very abundant and is a charac·
teristic representative in this fauna. but
it differs from many American species.
S. expansus is rare. but it resembles S.
u•abaunsensis Gt1NNELL and other Pen·nsyfvanian species of U.S. A ..
Gnathodus opimus is fairly common in
our collection, but it differs from mostof the American and European species.
As mentioned above the Omi conodont
fauna is dominated by Streptognatlwdus
japonicus, Gnathodus opimus. three new
species of Hindeodella, Synprioniodina
microden fa. SJwfhognatlwdus eclzigoen sis
and Ozarkodina orientale.
Up to date a few papers treated theLower Pennsylvanian or Upper NalilUrian conodonts were published, therefore.
it renders difficult to compare our faunawith those from foreign countries. The
Omi fauna is characterized by entire
lack of the typical l\'lississippian genera,such as Dollymae. Doliognathus, Scalio·
gnallllts, Staurognathus, Taphrognathus.
and others. It is also never overlookedthat the characteristic [vliddle Pennsyl·
vanian to Upper Triassic genus of Condo/ella is avoided in the fauna. From
the above mentioned evidences the pre·sent Omi conodont fauna apparently indicates Early Pennsylvanian or Late
Namurian in age, and it is thought to·46o.

Carboniferous Cu1wdonts fi'Om Central .Japanbe one of the richest conodont faunas ofthe mentioned age in the world.
Description of Species

Family Coleodonticlae BRA:\SONand l'viEHL. 194-1
Subfamily llincleodellinae BAss. 1959
Genus !Jindeodella BASSLER, 1925
Hindeodella asiatica leo

and KOIKE. n. sp.
Pl. '!.7. figs. 7 10Bar-like unit with anterior bar flexed
inward as well as downward approxi·
mately 90 degrees. anterior end further
curved toward posterior. and hook-likein shape. Posterior bar almost straight.
elliptical in transverse section. but flat·
tened except for blade-li l<e near posterior
end. Aboral side of bar sharply edged
except adjacent to small pulp cavity anddistinctly grooved along midline.
Denticles of posterior bar two sizes.
very closely set and directed posteriorly
and angled about 70 degrees with bar.
Large denticles sharply pointed andslightly curved inward. Smaller clcnti·
ctes needle·like. variable in size andshape. six to ten between adjacent larger
denticles. Denticles near posterior eilddiffer from those of most part of poste·rior bar. strongly angled posteriorly
and make an angle of ,10 degrees withbar. One or two slighlly smaller denti·
cles developed between adjacent larger
ones. ~lain cusp bicom·ex. larger thanany other denticles. arched posteriorly,
and curved inward.
Anterior bar angled downward justanterior to main cusp with abrupt inward
flexure at about one-third length ofanterior b:H from main cusp. Dcnticles

183

of anterior bar erect. discrete. larger
than small sized denticles of posteriorbar, and anteriorly increase in size.
Pulp cavity small with faint elevationof lips.
Remarks .·-Hindeodella asiatica closely
resembles H. armata STAUFFER and
PLU~II\IER described from the Pennsylvanian l'vlineral Wells formation in Tex.ls,
but our species is having more numerous
small denticles of the posterior bar andwell developed clcnticles of the anteriorbar. H. jragili~ HASS described fromthe Early lV!ississippian Chappel Limestone in Texas is also similar species
with our II. asiatica. but the present
species is easily distinguishable in more
numerous small sized denticles of theposterior bar and more distinctly discrete
larger denticles of the anterior bar. H.
bel/a STAUFFER and PLU:O.IMER was based
upon the fragmentary posterior end. The
posterior end of H. asiatica is almost
difficult to distinguish from //. bella.
Reg. no. 23002 (Holotype)
Hindeodella paradelicatula

leo and KoiKE, n. sp.
Pl. 27, figs. 3-5

B:1r slender. flexuous, very thin, andflattened in transverse section. Aboral
side sharply edged. but very fine shallow
groove developed except adjacent to oval
and small pulp cavity. Posterior end ofbar very thi.... hlade·like. and not gro·oved. Antenur tJar bends inward slight·
ly and its angle about 120 degrees \'v"ithbar.
Denticles of posterior bar with sharply
pointed end. two sizes of larger andsmaller which bend posteriorly and makean angle of about 50 to 70 degrees withbar. Large denticles deeply penetrated
into bar and markedly bend toward an·18-!

1hHayoshi ]GO and l'oshio KoiKE

terior. Smaller denticles about a half
length of larger ones and counted five
to seven between adjacent larger ones.
Denticles of posterior end make a small
angle with b:u· and alternate with larger
and smaller. Denticles of anterior bar
coalesced or discrete. seven or eight intotal number, and subequal in size andshape with larger denticles of posteriorbar. but two or three minute ones de·\'eloped near main cusp. l\lain cusp
largest in size. sharply pointed, and bi·
convex. Pulp cavity oval. small. anrlsurrounded by slightly elevated I ips.
Remarl~s :-Hindeodel/a paradelicalula
proposed herein is based upon complete
specimens and it resembles closely H.
de/icatu[a STAUFFER and PLU~IJ\IER de·scribed from the Graford formation in
Texas. General feature of the bar andthe arrangement of the denticles arealmost same each other except for more
flattened bar of our species. Lhving to
the fragmentary Texan species, detailed
comparison between both species seems
to be difficult.
Hindeodella salwgamii lc;oposteriorly. making an angle of 80 degrees in anterior part of bar. while about
30 to 40 degrees in posterior half. They
deeply penetrated into bar and stronglr
bend anteriorly. Denticles of anterior
bar two types in size. and almost samewith those of posterior bar except fewer
numbers of smaller denticles. !'.lain cusp
slightly larger than denticles of posterior
bar and directed posteriorly with slight·
ly larger angle than those of other dentides of posterior bar. Pulp cavity located just beneath main cusp, small.
narrow. and with faint elevation of lips.
Rl'marl~s :-Ilindeodella salwgamii re·sembles closely II. asiatica in mode ofinward curvature. but the present species
has a slight downward curvature of theanterior bar. ftat and thin both posterior
and anterior bars with almost similar
denticles.
Reg. no. 23007 (llolotypc).

Subfamily Ligonodininae IIASS, 1959
Genus Synprioniodina ULRICIT
and BASSLER, 1926

and KOIKE. n. sp.

Synprioniodina microdenta Eti.ISO'\

Pl. '27, figs. l. 2

Pl. 27. figs. 11-17

Bar consisting of slender and almost
straight posterior bar, and curved ante·rior bar. Posterior bar thin. flattened.
and finely grooved along sharply edgedaboral side. Anterior bar thinner thanposterior bar. blade-like, and angled
inward as well as downward. Inward
curvature with larger angle than down·
ward and makes an angle of about rightangle.
Position of curvature locatedabout middle of anterior bar.
Denticles of posterior bar consistingof larger and smaller ones. Larger den·tides biconvex. sharply pointed, directedS)•ntn·irmiodina sp. GtiNNELL, 1933. .four. Pal..
r·ot. 1. p. 269. pl. :n. fig. 6.
Synprioniodina miaodenta Eu.Jso:-;, 19·11.
Ibid .. rot. 15. p. 119. pl. 20, figs. 43-46:
ELLISON and GR.\\'ES, 1911,
11/issouri[inh•.. Sell. Min. Met .. Bulf .. Tech. Ser ..
to/. J.J. p. 3. 4. pl. 1. fig. lO.

Two h:us meet at forming of an angle
of nboul 40 degrees in common plane.
Posterior bar slender, very long, gently
arched. having sharply edged aboral side.
Denticles subequal. sharply pointed,
coalesced, and angled anteriorly to baseof bar at an angle of about 45 degrees.

l,6U.

Cm·bowiferous Conodonts from Centntl Japan

Large denticles about twice larger thansmaller ones and alternate in most partof posterior bar. Anti cusp short. withtriangular small denticles coalesced atbased and inclined anteriorly about anangle of 30 degrees with base of bar.
Main cusp at an apex of bifurcationof bars, sharply pointed. curved inward.
about twice as long as other denticles
and three times or more ncar base as
wide as other ones. and expanded atbase on inward side into wide flaring
lip of pulp cavity. while outer side lip
depressed in ward.
Aboral side of both bars trayersed byfine groove terminated in a deep conicalpulp cavity just below main cusp.
Remarks :-Our material is similar to
ELLiso:-.:'s specimens described from ~vlis­
souri and Texas in many respects. The
inclination of the denticles of both bars
and \\·eek expansion of lip of pulp cavityof the Japanese specimens slightly differ
from those of the American specimens.
Reg. no. 23015

Sy11prioniodina u1/linsoni lGo
and KOIKE. n. sp.
1'1. 27. figs. 18. 19

The blade·like bar meet at about rightangle. Posterior blade thin with sharpedged aboral side.
Denticles of posterior bar small. biconvex. coalesced near base and laterallycompressed, erect in anterior part andslightly curved posterior end.
Anticusp blade-like. with various si~ed
biconvex laterally compressed coalesced
denticles. and curved anteriorly. \laincusp large, biconvex. sharply pointed
with broad base. sharply curved poste·
riorly. and slightly eleYaled ridge along
midline of posterior side. but shallow
groove developed in anterior side.

185.

Aboral side of both blades traversedby distinct groove terminated in deep
conical pulp cavity beneath main cusp.
Pulp cavity with moderately elevatedtriangular lips.
Remarks :-The present new species
differs from the previously described
species in blade-like bars. shape of main
cusp and denticlcs of the posterior bar·and anticusp.
Reg. no. 23028 (llolotype·,.
Genus h,·uprioniodina ULRICH
and 13ASSLEI~. 1926

Euprioniodina dentata leo
and KOIKE. n. sp.
Pl. 28. ftg. 23

Bar-like units meet at forming of rightangle. Posterior bar thin with sharply
edged aboral side and pointed denticles.
Dent icles of posterior bar slightly convex inward. strongly arched toward
anterior, forming an angle of about 50
to 60 degrees. which subequal in shapeand si~e. laterally compressed. and biconvex with pointed end and broad base.
Anticusp short. having small triangulardenticles coalesced at base. and archedanteriorly about at right angle with baseof anticusp. ~-lain cusp at apex of bi·
furcalion of two blades. large. slurply
pointed. cun·ed outward with elevatedkeel along midline on inner side. andexpanded at base on inw:1rd side intoflaring lip of pulp cavity, but on outerside slightly concave.
Aboral side of posterior bar and anticusp grooved. and its opening terminatedin a deep pulp cavity of inward side ofmain cusp. extending to anticusp.
Remarks:- This species resembles Euprion iodina put cella YOU:\GQUIST and
Dmr:-:s described from the Early Pennsyl-

1H6

Hisnyoski [GO and Toshio KoiKE

vanian shale of Knoxville. Iowa. but thedcnticles of posterior bar of the former
are more anteriorly arched and h::tving
broad base.
Reg. no. 23026 (Holotype).
Genus Ligonodina BASSLER, \925

Ligonodina lzanaii !GOand KOIKE, n. sp.
Pl. 28. figS. 21. 22

Bar-like units meet at large angle.
Anticusp curved inward as well as dO\vnward. with two separated denticles so
far as observable. Posterior bar thin,with sharply edged aboral and oral sides.
and with one denticle so far as preserved.
Aboral side of bar finely grooved.
Denticles of anticusp sharply pointed,
curved inward. and circular in crosssection. Denticles of posterior bar laterally compreseed. and biconvex with sharp
pointed tips.
l\lain cusp very long,rather llexible. arched posteriorly, having
sharply pointed tip. Basal part of main
cusp subtriangular in cross section. but
circular in most part. Pulp cavity large
and subrhombic at <:bora! opening, deep,
and merges into main cusp.
Remarks :-Ligonodina lumaii resem·
bles closely L. typa tGu:-il\EI.L) than any
previously known species, but our form
has widely spaced denlicles of anticusp.and may have larger pulp cavity.
Reg. no. 23038 {Holotype).
Subfamily llibbardellinae 1\ltiLLER. 1956

Prioniodtts missouriensis Gt.;NNELL. l93l. Ibid ..
p. 246. pl. 2\l. fig. 9.

ldioprioniodus slriatus Gu:-;:-.EI.L, 1933. Ibid ..
z·ol. 7. p. 265, pl. 32, figs. 36. 37.
Hibbardella subacoda ELLISO:'\', 1941. Ibid.,z,of 15. p. 118. pl. 20, ligs. 22, 26.

Main cusp large, long, sharply pointed,
and slightly curved posteriorly andlaterally with sharpest curvature near
two-thirds length from pointed end.
Sharp-edged lateral ridges developed inmain cusp and continus downward alongeach limb of anterior arch. Posteriorside of main cusp grooved slightly ad·jacent to each of lateral ridges, butanterior side convex.
Dcnticles of anterior nrch well deve·
loped, discrete. sharply pointed andedged, biconvex. and arched anteriorly
and also inward slightly. Length ofposterior bar unknown.
Denticles ofposterior bar subequal with those ofanterior arch but smaller and erect.
Aboral side of main cusp excavat€dand subangular. Posterior bar distinctly
grooved. Pulp cavity beneath main cusp,
deep and conical.
Remarks :-The American specimens
described by GUNNELL (1933) and ELLISON(1941! are having slightly different curvature of the main cusp compared withthose of our specimens. but they arefragmental and further detailed comparison renders difficult. The Mississippian species of Roundya bamettana HASS
is having more distinctly discrete dentides.
Reg. no. 23027.

Genus Rouudya HASS, 1953
Subfamily Lonchodininae !·lAss. 1959

Rowzdya subacodus (GUN:>;ELL)

Genus Louchodina BASSLER, 1925

Pl. 28. fig. 20

Primzioclus subacodrts GUNNELl., 1931,
Pal., z•o/. 5, p. 246. pl. 29. fig. 9.

jour.

Lonchodina ? nipponica IGO
and KOIKE, n. sp.

MW.

Cwrbon·iferous Cunodonts from Central Japan

Pl. '!.7, fig. 20

Anterior bar longer than posterior bar,stought. subtriangular in cross section.
Denticles of anterior bar three. stought,
discrete, rather widely spaced with ob·
tuse tip, and almost straight to slightly
curved posteriorly. Main cusp largest
and rounded in cross seclion. Posterior
bar short with two small discrete denti·
des erected or slightly curved posteri·orly. Aboral side of bar broadly gro·oved, its opening continued from pulp
cavity. Pulp cavity large and shallow
beneath main cusp.
Remarks:- This new species resembles
Londzodina ? ponderosa ELLISON, but it
differs in short anterior and posteriorbars, and development of the denticles.
Short bar. fewer numbers of dcnticles,
large pulp cavity, and broadly grooved
aboral side of the bar differ from thetypical Lonclwdina. Generic position ofthis new species is tentative.
Reg. no. 23037 (llolotype).
Family Prioniodontidae BASSLE!{, 1925
Subfamily Prioniodontinae
BASSLER, 1925

187

base of blade.
Denticles of posterior blade seven,laterallr compressed some\vhat smaller
than those of anterior blade. closely set,
partly coalesced. making almost rightangle with base of blade. Apical denti·
cles laterally compressed. sharp-edged,
pointed, almost twice in width and lengthof other dcnticles, and arched making
an equal angle with those of denticles
of anterior blade.
Aboral side of blade sharply edged.
Pulp cavity long, extended into bothblades. narrow. deep, and with a moder·ately elevated flaring lip tapered intobase of apical denticle.
Remarks :-Ozarkodina orientale resem·
bles more closely 0. delicatula describedby STAUFFEH and PLU:Vll\IER (1932), GUN·:\ELL (1933). ELLISON (1941) and others
than any other described Pennsylvanianspec;es. It differs from 0. de!icatula inmore discrete denticles of the anterior
blade and fewer numbers of the denti·
cles in both blades. It also resembles
0. compressa REXROAD described fromthe Kinkaid of Illinois. but REXROAD's
species has more distinct and larger main
cusp than that of 0. orientale.
Reg. no. 23012 (1-lolotypc).

Genus Ozarkodina BRANSO:\
and rviEHL, 1933

Ozarkodina orientale leo
and KOIKE, n. sp.
Pl. 27. figs. 21-23

Bar blade-like. thin. orally arched. andlaterally slightly curved. Blade thickest
near base of denticles. Anterior blade
longer than posterior one.
Anterior
blade with six to seven laterally compressed subequal closely set denticles,
directed posteriorly in most part, inclined
at an angle of 20 to 30 degrees with

Subfamily Spathognathodontinae
HASS, 1959
Genus Spathognathodus BRA:\SONand l\lEHL, 1941

Spathognathodus echigoensis
IGO and KOIKE, n. sp.
Pl. 28. figs. 24. 25

Blade thin. laterally straight, slightly
arched with nearly straight aboral sideof anterior half.
Denticles subequal in size and shape,

188

Hisa.yosht' ]GO and Toshio 1\0IKE

but become smaller toward both anteriorly and posteriorly. coalesced deeply
with sharply pointed tip and grooved
between adjacent denticles. and angled
posteriorly. The first two anterior den·tides erect and very thin.
Aboral side of blade sharply edged,
grooved along midline except pulp
cavity. Pulp cavity large, deep, withexpanded lip which continues to almost
posterior end of blade. and traversed bylongitudinal grooye extending fromblade.
Remarl~s :-Spathognathodus echigoensis
resembles closely S. minufus (ELLISON),but it differs from the American species
in almost straight aboral side of blade
and subequal size of the denticles. andlarger pulp cavity.
Reg. no. 23036 (Holotype :.

Trough shallow, and restricted poste·
r:orly. two-thirds or less length of plate.
Blade shorter than plate so far as ex·
amined specimens. Denticles of blade
rather large compared with size of blade.
Aboral side of blade grooved along mid·line and its opening continued into expanded pulp cavity. Aboral side of pulp
cavity entirely smooth.
Remarks:~ This new species resembles
Idiogllallwdus delicatus Gu~:-;ELL, but it
can be distinguished easily from the latterby the longitudinal ridge ornamentationof the posterior part of platform.
Reg. no. 23034 (1-Iolotype).
c_;enus Streptognatlwdus STAUFFER
and PI.m.IMER. 1932
S/reptognathodus japo11icus !Goand KoiKE. n. sp.
Family ldiognathodontidae HARRIS.
and HOLLINGSWORTH. 1m3
Subfamily ldiognathodontinae HARRIS
and lloLLI:\GSWORTH. 1933
Genus Idiognathodus GuN;-.;EJ.J.. 1931
Irlio!.[nathodus togashii lc;oand 1\0JKE. n. sp.
Pl. 28. figs. l-tl

Plate lanceolate. pointed anteriorly aswell and posteriorly. widest near middle.
and steeply sided. Cup small and asymmetrical, outer side slightly 13rger thaninner side. Axis almost straight. but
feebly curved inward. Oral surface of
platform ornamented with parallel trans·verse ridges consisting of coalesced nodes
at about two-thirds anterior part of plate.
Posterior part of platform split by blade
and also ornamented with longitudinalridges consisting of coalesced nodes.

Pl. 28. figs. 5-13

Cup asymmetrical. pointed anteriorly,
widest near middle, outer side always
larger than inner side. Platform lan·ceolate and transverse section of oralsu~·face deeply concave. Oral surface of
platform ornamented with 1wo or morenode·lil<e parallel longitudinal ridges,
narrow and obsolescent toward anterior
end. l'vleclian trough rather shallow. de·epest ncar widest part of cup, slightly
curyed inward posteriorly.
Blade subequl in length to plate. verythin in posterior end. thickest just be·neath denticulated oral side. Denticlesof blade ten or more in mature specimen.
almost a half height of blade, but coa·lesced each other at about two-thirds
from base.
Denticles chevron-like inlateral view, largest near posterior end.
gradually decreasing in size toward plateand continued anteriorly as carina.
Carina short and one·fifth or less of

4(ifi.

Ca.rbonife;·olts Concdonts from Centml .JaJJII.n

length of plate, and merged into longi·
tudinal ridge of outer side.
Aboral side of blade finely grooved
along midline. Aboral side of expanded
pulp C:lYity entirely smooth. but rather
deeply grooved along midLne.
Remarlzs :-The present new species
has rather broad ,·ariation in size. shapeof plate, and oral side ornamentation of
platform. It resembles closely S. sui·cop!icalus YOF\GQUIST, HAWLEY andMILLER described from the Permian of
Idaho. but this japanese form has more
irregular parallel r:dges on platform. It
also slightly resembles S. eleganlula
STAUFFER and PI.lii\1:\IER. S. gracilis
STAUFFER and PUIM:\1ER. but it is easily
distinguishable therefrom by the oral
ornamentation of pia tform and shortcarina.
Reg. no. 23032 1Holotypel.
S/reptognathodus cxpansus leo

and KoiKE, n. sp.
Pl. 28. ftg. 14

Cup large. highly asymmetrical. point·
cd anteriorly, greatest width near middleof cup. outer side larger than inner side.
Ax:s curved inw:.~rd at junction of blade
and platform. Platform lanceolilte. andoral surface ornamented with fourteenparallel transverse ridges ending abrupt·
ly at median trough and slightly ra·
dialing from trough in anterior half ofcup. Two rows of various sized six
nodes developed in posterior part of
inner platform along axis and a row offewer numbers of node developed inouter platform. :\ledian trough shallow
and transverse section of oral surface of
platform slightly concave.
Blade subequal in length to cup. thintroughout. with nine dentic\es. Dentic\esof blade largest ncar posterior end,

18D

chevron-like in lateral view. and coa·lesced at base. Carina short and re·stricted poster;or end of cup. Aboral
surface of expanded pulp cavity smooth,grooved along midline. its opening con·rinuecl from blade.
Remarks:- It resembles ELLJSo~·s(19411 specimens of Streptognathodus u:abaunsensis Gunnell. but it has more
simple ornamentation of nodes ncar po·
sterior part of the platform and fewer
numbers of denticles of blade.
S. excelsus STAUFFEI~ and Pr.m.niER
and S. oppletus ELLISO:\ are also allied
species of S. expansus. Howe\·er. S. ex·
celsus has more complicated ornamenta·lion on the posterior part of cup, and S.
opletus has longer carina than those of
S. expansus.
Reg. no. 23033 iHololype'.

Genus Gnathodus PA:\DER. 18;)6
Gllatlzodus opimus IGO

and KOIKE. n. sp.
Pl. 28. figs. 15-18

Axis straight to slightly angled inward
at junction of blade and carina. Cup
highly asymmetrical. pointed anteriorly,
widest at two-thirds from anterior end,and outer side larger than inner side.
Oral surface of cup plain and smooth.
Platform grooved by shallow troughalong axis, deepest at posterior end. andornamented with node-like parallel ridges
longitudinal!~· along carina. Denticles ofcarina rounded in tip. fused, node-like.
and largest at widest part of platform.
11lade almost equal or slightly larger
than carina. very thin. and subequal in
thickness throughout. Denticles of blade
coalesced with sharp edged tip. andchevron-like. which largest at posterior
end and decreased in size anteriorly.

190

Hisa.yo:;hi lGO and Toshio I<OJ KE

Aboral side of blade sharply edged, finely
grooved along midline. merging into expanded pulp cavity. Aboral surface ofpulp cavity smooth and grooved along
midline and its opening conLinuecl fromblade.
Remarks :-Gnalhodus opimus resembles G. dilatus STAUFFER and PLU:'v!JVJER
and G. roundyi GUl\NELL in two parallelridges. but our specimens have rounded
node-like low denticles of carina andmore expanded pulp cavity. It is easily
distinguishable from other described
Gnathodus by smooth oral surface of cup.
Reg. no. 23030 (Holotype).

ReferencesW. (1935): Micropaleontology andstraligraphy of the Lower Pennsyh·anianof central Missouri. }our. Paleont.. 110/.
9, /lO. 6. p. 483-502, pl. 55.
BAKER. R. C. (1942) : The age and fossilsof the Olentangy Shale of central Ohio,
Amer. jour. Sci .. I'O! 240. 110. 2. p. 137143. pis. 1-3.
BRA:-;soN, E. B. and Mt::HL. M.G. (19-ll) :New and Iitle known Carboniferous conodont genera. Jour. Paleo11f., 110{. 15. no.
2. p. 97-106. pl. 19.
- - and (19H): Conodonts. ill Index
fossils of North America, by H. W.
Slll~IPER and R. R. SHROCK. \VILEY andBAILEY.

Expl::tnation of Plate 27(All ligs. X 75)

Figs. I. 2. Hindeodella sakagamii leo and KoiKE. n. sp.
I. Lateral view of the holotype. Reg. no. 23008.
2. Lateral l'iew of a complete specimen.
Figs. 3-6. Hindeode/la paradelicalula Teo and Kott\E, n. sp.
3. Lateral view of the holotype. Reg. no. 23006.
4. Lateral view of a complete specimen.
5. 6. Lateral view of two large imcomplete specimens.
Figs. 7-10. Himleodella asiatica leo and KotKt::. n. sp.
7. Lateral view of the holotype. Reg. no. 23002.
8, 9. Lateral views of two incomplete specimens.
10. Posterior end of the posterior bar of a large specimen.
Figs. 11-17. Synprioniodina microdenta ELLISON
II. 12. Two lateral views of the same specimen.
13. Lateral view of posterior bar.
1-!. Lateral view of a complete specimen.
15. 16. Two lateral views of the same specimen.
17. Lateral view.
Figs. 18. 19. Synj!rioniodina col/insoni Ic:o and Ko1 KE. n. sp.
18. Lateral view of the holotype. Reg. no. 23028.
19. Lateral view.
Fig. 20. Lonchodina ? nipponica leo and KoiKE. n. sp.
20. Lateral view of the holotype. Reg. no 23037.
Figs. 21-23. Ozarkodina orientale leo and KorKE. n. sp.
21. Lateral view of the holotype. Reg. no. 23012.
22, 23. Lateral views of two incomplete specimens.

!GO and K OIKE :Carboniferous Conodonts from Central Japan

Plate 27

466.

Cm·boni;(erons Concdonts jmm Cent1·al Japa.n

So1-:s. New York. p. 235-2·16. pis. ~)3-9-1.
CooPER, C. L. and SLoss. L. L. (1943): Cono·
dont fauna and distribution of a Lower:\iississippian black shale in Montana
and Alberta. jour. Paleont .. vo/. 11. no.
2. p. 168-176, pis. 28-29- - (1947): Upper Kinkaid (Mississippian)microfauna from Johnson Country. Illinois. !hid., vo/. 21. uo. 2. p. 81-94. pis.
19-23.
EDWARDS. W. and S-ruBnLF.FII::LD. C. J.
(1948): Marine bands and other faunalmarker-horizons in relation to the sedimentary cycles of the Middle Coal 1\•leasures of Not tinghamshire and Derbyshire.
with an appendix by K. C. DtiNIL-\111.
(Juart. jour. Geot. Soc. London. t•ot. 103,
pt. 4. 110. 412. p. 209-260. pis. 13-15.
Elltso:-.:. S. (1941): Revision of the Pennsylvanian conodonts : ]our. Pa/eont.. t•ol.
15. no. 2. p. 107-143. pis. 20-23.
- - (1946) : Conodonts as Paleozoic guidefossils: A mer. Assoc. Petr. Geo/. Bull.,t•ol. 30. lUI. 1. p. 93-110, 3 figs.
- - and GH,WEs. R. W. (19,11):Lower
Pennsylvanian (Dil]lple Limestone) conn·
donts of the Marathon region, Texas.
[]nil'. 1\lissouri. Sch. Min. and 1\le/. Bull ..
Tech. Ser .. t·ol. 1-J, no. 3. p. 1-13. pis. 1-3.
FAY. R. 0. (1952) : Catalogue of conodonts.
Kcmsas Ultit·. Pat. Contr., Vertebrata, art.
3, p. 1-206, figs. 1-109.
FPJITA. H (1958): Geology of the Omi Limestone (Permo-Carboniferous), Niigata
Prefecture.
Toholm Unil•., Ins/. Geo/.
Pal .. Con/r.. no. 48. p. 1-60.
Gu:o-:NELL, F. II. (1931): Conodonts from theFort Scott Limestone of 'Missouri. jour.
Paleont., t·ot . .'>. 110. 3. p. 244-253, pl. 29.
- - (1933) : Conodonts and fish remains
from the Cherokee. Kansas City, andWabaunsee Groups of Missouri andKansas. Ibid .. rot. 7. no. 3. p. 261-297,
pis. 31-33.
HARRIS. R. W. and Hou.t:-.:cswoRTII. R. V.
(1933): New Pennsylvanian conodontsfrom Oklahoma. Amer. jour. Sci. Ser.
5. t•ol. 25, 110. 147. p. 193-204, pl. 1.
liAR LT01'. B. II. (1933) : l\Iicropaleontologyof the Pennsylvanian Johns Valley Shale

191

of the Ouachita Mountains, Oklahoma
and its relationship to the :VIississippian
Caney Shale. ]our. Pa/eont .. t•o/. 7. 110. 1.
p. 3-29, pis. 1-7.
H... ss. W.ll. (1941): Morphology of conodonts. jour. Paleont .. z·ol. 15. 110. 1, p.
71-81, pis. 12-16.
- - (1953): Conodonts of the Barnett Formation of Texas. r..·. S. Geof. Sun·. Prof.
Paper 2-13-F, p. 69-94. pis. J.l-16.
- - (1959): Conodonts from the Chappel
Limestone of Texas: U.S. Geo/. Surz,_
Prof. Paper 29,1-]. p. 365-399. pis. 46-50.
- - , RHoDEs. F. H. T. and Mt!LLI,R, K. J.
(1962): Conodonts. In R. C. MoORE's
Treatise m1 Im•ertebrate Paleonlo/of!y. pt.
IV, p. W1-W97: \V246-W259.
HAYASAKA. !. (192-!): On the fauna of the
Anthracolithic LimestOne. Omi-mura in
the western part of Echigo. Tolwku Imp.
[.'nir .. Sci. Rep .. ser. 2 (Geo/.), 1'0!. 8. p.
1-82, pis. 1-7.
HA YASIII. S. (1963) : On the conodonts newly
discovered from the Ash io Mountains.
central Japan. Chikyulwglzku (!·:arth Science), 110. 68, p. 9-12. pl. 1 (in japanese).
HJGGI1'S, A. C. (1961): Some Namurian conodonts from Korth Staffordshire. Geo!.
Mag .. t•o/. 98, p. 210-224.
KATO, M. and ::\AKA:o.n:R.'\, K. (1962): A
Goniatite from the Omi Limestone. Chi·kyukagaku (Earth Science). 110. 63. p. 3334. pl. 6 (in Japanese).
KAWADA, S. (1954):
Stratigraphical andpaleontological studies of Omi Limestone
in the ltagamine district. Kiigata Prefecture. Tokyo Univ. Education, !nsf. Geo/.
Mi11 .. Studies, no 3. p. 15-27.
MILLER. A. K. and Yoc:--iGQUST. W. (1947):
Conodonts from the type section of the
Sweetland Creek Shale in Iowa. ]our.
Paleont .. 1•0/. 21. 110. 6. p. 501-517, pis.
72-75.
MtLLER. K. J. (1956):
Taxonomy. nomenclature. orientation, and stratigraphic
evaluation of conodonts. Ibid., t•o/. 30.
110- 6. p. 1324-1340. pl. J.l5.
REXROAD. C. B. (1957) : Conodonts from the
Chester Series in the type area of southwestern Illinois. Illinois Stale Geol. Sun• ..

192

Hisa.yoshi ]GO and Toshio l\OIKE

Rept. illl'est.. 199. p. 1-·13. pis. 1-'l.
- - (1958) : Conodonts from the Glen Dean
Formation (Chester) of rhe Illinois Basin.
Ibid .. 209, p. 1-27, pis. 1-6.
- - (1958) : The connodont homcomorphs
Taphrognatlws
and Streptagnathodus.
]our. l'aleont .. vol . .12. no. 6. p. 1158-1159.
- - and BllHTON, R. C. (1961): Conodontsfrom the Kinkaid Formation (Chester)in Illinois. Ibid.. t•ol. 35, 110. 6, p. 11431158. pis. 138-141.
- - and CoLLt:-=so:--.-. C. (1961): Preliminarr
range chart of conodonts from the
Chester Series (Mississippian) in the
Illinois Basin: llli11ois State Geol. Sun• ..
Cirwla r 319. p. 1-11.
- - and - - (19G3) : Conodonts from the
Sr. Louis Formation (Valmeyeran Series)
of Illinois. Indiana. and Missouri. Ibid ..
Circular 355. p. 1-28. pis. 1-2.

RHODES. F. H. T.

(1952): A classification ofPennsylvanian conodont assemblages.
four. Paleont., vo/. 26, no. 6. p. 886-901.
pis. 126-129.
SAKAGA:\11, S. (1962) : Discovery of Profusulinel/a from the Omi Limestone. Niigata
Prefecture. four. Geol. Soc. ]apa11. t•o/.
68, /1(). 7.97. p. 175.
STA:-<I.EY, E. A. (1958) : Some Mississippian
conodonts from the 1-1 igh Resistivity
Shale of the Nancy \Vatson No.1 Well innortheastern Mississippi. ]our. Paleont ..
t•o/ ..12. tzo. 3, p. 459-476, pis. 63-68.
STAUFFER, C. R. and PLt::\1.\IER, H.]. (1932):
Texas Pennsyh·anian conodonts and theirstratigraphic relations.
Unit•. Texas,
Bull. 3201. p. 13-50, pis. 1-4.
STt:RcEo". :M. T. and Yot::o-:cQUIST. W. (1949):
Allegheny conodonts from eastern Ohio.
.foul'. Paleont., 1'0!. 23, 110. 4. p. 380-386,

Explanation of Plate 28(All Jigs. x75)

ldiognathodus togashii IGo and Kon;E, n. sp.
Oral view of holotype. Reg. no. 23034.
2~1. Two lateral views and aboral dew of the same specimen.
Figs. 5-13. Streptognatlzodus japonicus !Go and KoiKE, n. sp.
5-7. Oral. lateral and aboral views of a small specimen.
8-10. Lateral. oral and aboral views of lhe holotype. Reg. no. 23032.
11-13. Oral. aboral and lateral views of a large specimen.
Fig. H. Streptognatlwdus expansus IGo and KorKE. n. sp.
1-1. Oral view of the holotype. Heg. no. 23033.
Figs. 15-18. Gnathodus opimus leo and KotKE. n. sp.
15-17. Oral. lateral and aboral views of the same specimen.
18. Oral ,·iew of the holotypc. Reg. no. 23030.
Fig. 19. Gnathodus sp.
19. Oral view of an incomplete specimen.
Fig. 20. Romrdya subacodus (GUNNr·:t.L)
20. Lateral view.
Fig. 21, 22. Ligonodina hanaii lc;o and KotKE. n. sp.
21. Lateral view of the holot.ype. Reg. no. 23038.
22. Lateral view of another specimen.
Fig. 23. J.:upl'ioniodilla dentata leo and KorKE. n. sp.
23. Lateral view of the holotype. Reg. no. 23026.
Figs. 2-1, 25. Spathognathodus echiJ.:oensis !Go and KoiKE. n. sp.
2-1. Lateral view.
25. Lateral view of the holotype. Reg. no. 23036.
Figs. 1-·1.
I.

Leo and K OIEE :Carbon1jerous Conodonts from Central Japan

Plate 28

5
7

9

25

4fi6.

Carbonij(,row.; Conodo11ts fmm Central Japanpis. 74-75.
T. L. and GoEBEL. E. D. (1963):
Preliminary report on conodonts of t.he
1\leramecian Stage (Upper :\lississippian)from the subsurface of western Kansas.
Kansas State c;eol. Sun•. !Jull .. 165. pt.
], p. 1-16.
You:-;GQUIST. W. and Do\1·1\s. I~. H. (1949):
Additional conodonts from the Pennsyl·
vanial of Iowa: ]our. Pateout. ro/. 23.
uo. 2. p. 161-171. pis. 30-31.
--and HEEZE!\, B. C. (19~8): Some Pennsylvanian conodonts from Iowa: Ibid., z•o!.
22. 110. li, p. 767-773. pl. 118.
- - . IIAWLEY, R. \V. and 1\·11LLER. :\. K.

Tllo~IPSO:'\,(1951) :Phosphoria conodonts fromsoutheastern Idaho: Ibid .. Pol. 25. no. 3.
p. 356-36 L pl. 54.
- - and MILLER. A. K. (19-!9): Conodontsfrom lhe Late i\Iississippian Pella bedsof south-central Iowa. Ibid .. vo/. 23. 110.
li. p. 617-622. pl. 101.
- - . --and Dow"\s, H. R (1950): Burling·
ton conodonts from Iowa. Ibid .. z,of. 2-J,
110. 5. p. 525-530, pl. 67.
- - and PETERSO:'\, R. F. (1947) : Conodontsfrom the Sheffield Formation of north·central Iowa. Ibid .. 1'0/. 21. no. 3. p.
2-12-253. pis. 36-38.

All specimens described herein were kept at the Institute of Geology and Mineralogy.
Tokyo University of Education.

194

PROCEEDINGS OF THE PALAEONTOLOGICAL SOCIETY
OF JAPAN

B;;$:1';~~~;.~19641flll:~~.:tD.to:1r~i:!: 1964
1f 1 Yi 18 · 19 B7L~Ii"l(~Y:~!f'bJ7Y{~Ji'Ff.¥:'<'~ 101 -It~l::..:to~·"LmlRU~ht.: (/Hiffi:tf 45 ~).

.ill\:

lj.i{........ ·'H'l;J'(::.-;..~:f-~?.z.. r.ilHU:Mf*~~ (Ontogeny and

Evolution) J
M~l~:mm~oo~~~t-~.. ~mili~~"Miogypsina ·· nepionic acceleration r.:."?~·"L... . ........................ £\:%~ '1i.
Nummulitidae. Orbiroidae. Lepidorbitoidi·
dae. Lepidocyclinidac. Amphistcginidae
lf1~1:t.~co*ile~"E .............. 'f<.fRIEJIY£!}..... ~n; Jh

Scutell idae coif!.[(l:l'l.im~7 :/~71 1·1.:..'1..~ (Vjf.M~'Ec.if!lfl:....
. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . f'A;;$:iJ£1ll;

li!K~ Dasycladaceae ~-fti:. "?l· ·"L .. JiJWI~i.l(On the evolution of Cycadeoidea .... Ji;Ji;~jilOntogeny. phylogenr and eYolution in Pale·
ontology. . ..
. .............. !j!Jj;{ --Jlil

Microplankton from the Asagai formationin the Joban coal·tield ................ .
. .. .. .. .. ..
Kiyoshi TAI\AII/\SHl
Cretaceous planktonic foraminiferal sue·cession in the Putah Creek. Yolo Napa
and Solane County. California .
. . . . . . . Yokichi TAKAYA:-.AGI
Two r.ew Permian fusulinid genera fromThailand ......................... Ryuzo
TORJYA:<I.-\ and Kametoshi KA:-;!\II':RA
Triassic coral fnunas from the Konosegroup in Kyushu ... Kametoshi KA:-;!-.JF:RA
Notes on a coral-brachiopod species as·

1'semblage in the lowest part of the
Akiyoshi limestone group. . .......... .
. . l'vlasamichi 0TA and Juichi YA:-.AGWA
A new species of Hamlingelfa (Brachiopoda)
in Japan. (f~~) ........ Takasyi 1-IA:o.L>.DA
Some Silurian brachiopods from Yokokura·
yam a, Kochi Prefecture. Japan ........ .
. . . . . . . . . . . . . . . . . . . . . . . . . . . . Mitsuo KooA
Some Permian Aviculopectinidae from the
Kitakami Massif. Northeastern Japan.
(f\'::im:) ................ Masafumi MuRATA
On some Cretaceous Corbulids from japan.
..................... Yoshihisa 0TA
On a new bake,·elloid. from the Lower
Cretaceous of Japan ....... ltaru IIAYA:\11
Middle Triassic Ammonoids from Japan.
... Yuji BA:"Do
Note on a new genus of Collignoniceratidae(Cretaceous Ammonoidea) ........... .
.................... Tatsuro MATSli:V!OTo
Ontogeny of Reesidites mirzimus ......... .
.............................. Jkuo 0BATA
On th.e protoconch of some gastropods ..
............................ Tsugio SHUTo
1\'aticid gastropods from the Miyazakigroup ..................... Tsugio SHUTO
On some Paleogene gastropods from north
Kyush u. Japan ........................ .
........ Tsugio SJWTO and Yoshiro 'UEOA
I~;IJ;i$&1JilHtfii:~~Illi'fHt.T:il:."?~·-c (-t~O· ...

.................................. 7.;:~mz.

lfl
1:'r
Jl!fi'J7;; 7Jli~~nl!'tff.I'Jl~Hlf ............... .
.............. .tN;I\Ji- · .!1M.l.JJ'.f~. ;f.:fti'd$

M 16 ITIIJT<I~lllb~"'f-~~7l0-:IJil~iiJ%t1ir:fi~.Hll.~

m11r .............................. l'Jilll 1tl~

7Jlt~:fi&<~h'ff,[.q:.l1!!f!~id~~Hs·· ....... ~IE

leiE

~

~

.

~

o

1963 ~l=f:lt1:.~Jl\ffifl I!Jj#iJlf'tn>~ec~nt..:.

01963 1F!lrc.a.~Jt(l)*fi:W:!lUI8 H''F~fliJ~~fllitc.il!i$tUI (I) l*fllr=P1£1~=ttft1fiQ)Iilf~J fij 8~~~o*tiJJ~fM!Iilc. i!!illl1'i3.ti (I) r 8 &O:l!iftJI!!~ooll!l.ni!IJ~Uf(l)lilf~J flh\: ~io!i~WJ~I~Htlil~ W!llf3eB~ (I)r *JllfiiW!@fmtttMt/'ti!EO: IC.~1¥ii!IJ#J{t.:0ll1\(l)lilf~J ~ El1!t~11RI.:. ~~ J~lEP.YliB'a (I) r~~;{ftL!!tm(l)i'f~J ~..:cn.-ttttffid!t Lt..:. .::(I) ; t;.i?!illJ~=:.tt(l)m~'1:.'UHt~Piii~JrlJJ~I.I:~lJ. ~nt.:.
1964 ~ 1 JJ 18 8 Jtffl*:"f!-c!18f!H ~n.t..:*~~:fr::.M ,-c~nY&O:Iilll&!fJJi!nq:k(l) .1: 5t~~lE ~.11-t.:.
a. 1'di1J~ 12*:m(l)~·.mt-trf.ii:t~it? -c~A?o. ~Y!ltti'im~ft.¥ 1. ooo fl:lt~}JI]~ft¥ 1. 500 R*~JltJ~i;\:1¥

1o.ooo l'iL:J.J:t-J-7.>. 15tt=~~Lt.~~jlji\(l):liJ1Sn:ts:~.·. f'E?'i-(l)ii:~l:t.=¥4 ?llc-t7.>.

b. lfii~J;l!.~: I :l. lli(~ (tefi~ · JIJ!.fg! · f.J'}(:5.-~l:r) a 2·1 lll.lllU~ ( !11 7 71 !1- ffl~*t.l 60 fjc) ~1\a!lc -to.
4. lf!;lll& (14.2x20 em) 1"1 2 Mct-IR!l: c -t 7.>.
5. :filill<!lt.IO fr.lfrlit60 W (1 .lf.lt 1 :ifl1.i"'D ~JlR!lcL. A~l::t.JHt..:lttl~ffl1~n:.-riJJ-liKU~
m~

.....

7.

J:JJ:Q)~lUU:·.:::t~~-ftlH:u;ft.ffi:a:~\"~tdHt!-t7.>. ~t..:lli{fln: 4 Filii0Jii""C:I1)~fl't~lt~ll!l2

jl.g.ift 12 J>j!~.:: :t ¢~lgj$.; .l: c::,-=~Jl&(l)~nJt-;F,~I.I;ft_ff!-t" 0(II 7. lil'llli!t. L II 8 t- II 7 t:.f:\U:If 7.> ,)

~(~

f}lj

Ill!

0

J;J.

J:

I .rM'W\$.i<HiW81964 ~ 6 Jl 7 [jm87 rm #JJ ii<
II&*~
I 1964 =¥ 5 fJ 5 0
196-t 1]! 9 .fJ 20 8
l:ll88(ill~~~*~
I 1964 ~ 8 J1 20 8m 87 J!li#JJ~ o?:tffl:k~r:): r¥r~ftl~ct~.(l)fi~1t.EJ 1:.'?~·-r(/)'/~;f{,;; .... -.A Cl!I:JliA er .itO
8

1:ll 88 ill!~~ (IAXli*'.'f.) : lli:M~~J(i!t.lliA ~Ft?'l-l'h)News;K I?< Musee Royal d'Histoire Naturelles de Belgique (l)~lH~t- L -c~ •t..: Prof. Dr. Victor
Emile Van Straelen it. Jiil:ili: Charles Darwin Foundation for the Galapagos lsland3 (I) President

c L. -c 7i7J{:::~· ;;t..iJ·I?M 1J. *~ 2 .fJ 29 f:HIH~·(I)~Ifl~!lt L -c 75 fl(l)~iffit-M t: t..:.

19641p4Jl 5 B
19641f,4Jjl0A

ffi

IS

~

***~~~00it!!'i!£~fi~P3fj

a *ir1:~1f!~~~

m s3 ~

40 0 f9

ffi

APIJ

~

~

#

il1li

B ::$:15'1:~1£:~ffli5' ·fell'mHI~•
em " r:~ ~ ~ Jj'( 8 4 1 8 o •>* F: w~ ~K z x- r, Otr z / ts

•w~•m~~~~~

a

m

~

(1964, 1, 18);$:~11 El ;;fi:ti ~¥;)~~c. ~' ? 0:<$:~1iti~~~:t3J: r.f 1:. tw:OOI*<b ?~~flO)ifEW::t3J: 0:~&-i-iif?O)~ fol!t:J c T ?.
**~~2*0)§!t:J~~T?~~~O)·~~fi?o1. ~:~5-c O)titJ,O)I:l:llltl~O)~fi.

2. ·~m;tM!ili~O)fllJitll.

3. tlf&O)t.:~O)~m~ • 3tiiii~'E'O)lH>O)tmlilL·1. !Jf~UJ~I~)J · ~MJJ:to J: lJIVf~*Mt.c t:>lJI::.~tn~-:.~-t-t Q;/J~QJ;&~H·UJI!!!tll 2 !#:UJ ~ (J(J~ij!t
1::.~-t;;,.:::.!: 0

~H*

m5*
mG*
m1*
ms*
~9*
~10*

< c

**O)§!t:J~i!T? t.:bb~~0)1it-i>H"t':<$:~1~:tT.!ii!O)!iJf~~fil.~~ft 1:. iJ>-r: ~ ?.
;;f;:~l±ti~~~:t3J: i:ll:. :h.I~OO~.:b ?Mi~.ffl<::~lij;-iot,'f-:>~fil.i:t!Ui-t ?.
~P\l-i-5ttt"t'~illi~i'! • %;JIJ~t11. • JtUIJ~fUJJ:U:~W:*.l'ilc T ?o~iM~fi!liiJi'Jl::O).A~$.\MIHII!Ui t.., t::#fl~-:> ~ ~ili£:l.~O)il51~J: ~ l'Ji::bb 1.> otUU*~Ii;;$:~11:. 10 !Fttl:~tl "t'.:b IJ ti~¥JJ~I<::-:>t 'l'i4UlO).:b? b O)"'f:, ~BU~/il5 ;80)/ft!l¥0).:b ~ t.: t 0)1~-:> ~ Jlll!iiN~~O);Jjii~J: ~ "'CJ:i::bb 1.>

0'!!rti!IJ~till:tm 2 ~0)§ !t:J~:ftUIJT 7, !*.A. i:ifi!~/il~O)j'ffi~I~J: 7,.
:81l~.l'illiti~~~~<::-:>~ 'l'ifitft..:~tlfO),:b 7.l#l~-:> ~iif~tHiiJ>ff£JIJL,, ~~O)~SI~J: ~"t'J£~1.>.
:ft~lif!l112 *'~J£bb ;~1-t.:idH,•ti'Jbbt..:tH'Ll!t..:; t.ct '• 4H.U:t~$0)~1!i~~~tl:F. 3 ~~~t;!l!

m12*

J£L-~$~~#~TQI:.c~"t'~1.>o~~O)ifltflili~~~l:)!f~ "[}£~ 0 0 ~ftlHUi~fl$ I, 000 F'J, ~}JI]~f.\$1. 500 fq, JfJ!IJ~~!f. to. ooo fl]JJ.J:c -t 7.1. :8'liH~Ji'll:t~'l!1ft'JA.O)ft~7J>t.c~ 'o :i£;>1-0)~t'J.Ii!F 4 iJB c -t 7.1.

~13*
~14*

;;f;:~O)gft~~ft·WM*·~~·~~~J:o.
~~-i>l~$Ul:W~t..,~-:t3J:r.f*~O)S-~m-tfiAO).:b?~:#f~.~--~O)S~Hl'~
S-tol:.ciJ:"t'~oo

~15*

1

15

2

*ftO)~AI:l~* S. if;Jl.
:g ct..,. ? t;;!'i=f:g ~m~~.H'!C Too ifWlli~l' $C. 1.f!illi~MHft~t 'o~RO)~!Jilill:.t IJ *~ ~~~~Ji:to J: r.f\lf:iet'i=F~ ~if!< 1:. 'IJ~-r: ~ 0 0'iitf6:t!J;;lli~~t:l,~rz:t-.H ''lli:.Ql! ~ H 1.>. ~.!&J;;l.IH~;~I]~!'aO)Ij1fJ> ;~t~ O)iJJi/1t.i!* I~J:? "['~lil ~:11.1.>.
:ftfiWNilliJ4.H-~O)Jfli>· ~;jljl,i£~~K:t3\ ''li:Mill ~ ~t. :.t:~~ ti:2<L-~f6~tfl!l!T oo~GfUiii'&ib o.mitti~ JU::i.iR!il<::{-tl!l!~~~~T 1.> oc

mt6:i(:
tf, 17*mts~<.:. cn:-c ~ Q, ~1'~*11il¥.llHH~t;tfU! L-~~ QJ ~a1.:. J: -:>"C~clb:<1>:~1:li~$-·(1ll!.irf§IJ~~Hfl <o -t' O)~*ll:.l.i~~iJ: ~f) *~.i!J!'SO)£Ji>:1Jil·-i>iJ!i:E:-t o.

*~1.:.1MHH~:R~/I!l¢ o ~ff:f'<:Rr1;w;l!HH~I.:.#;.bll1" ~ .::..!:: n:-c ~;;,.

m~O);ii~ liiWJIU't~iJ :~;.t-t o.

~au0•iJ~ot~~oq~~q~~.a-B•-r;;,.~~~~ftO)+~O)-JJ.~O)I:l:l~.a-&-:>

l'~.TI:Too~~~i~aO).:::.O)~-uJ:O)#tJ:~~IDBrf:Jt.: ollll:l'l:t3J: rYB.O):f!llffi ~liiHil L- t.:ami:!r t-:>~19*

l'i&~B-O)lR;1<.-i>~ltt.:~alilli~f@~-i>t3.-t7.lo~~I<::UH'f.VJ:t '~!'!lifi!!O)l:l:\~~~~~-t'O)~~fffiO)fj~-io~ 1-f.-t o 1:. ciJ:"t' ~ 7.1. {13. L,, ~m2o*f.&~O)ji~li~~~I<:J: I). ilJefili~O)~Ii.ii1JHI> 1:. :11.~~~ ?.

J,'(,~fc.J.O)~~mO)t-tfir;t 1 A.l ~~c:~?o~21~~*:t3J: r.fjl'f;fi_fc.l.liiif.liU~~-i>;!'nM\l,. ~~ O)i:JtilRJ: o~*J'jtJ-I~flEP~"S'~!JJ{ -i> ··~~

"too
~23:i(:

mlli~ftl:t?itBf~Jt~~m•t..,lfliiJt~O)~llll~t&?t 'l'~Br.a-t~~n-t .::,.
*~O)~i!t!FJi:r±m$1 ~ 1~~~~ IJ 12 f.! 31
1c.*!o.

~24**~~U&~W"toR~~~KM.t..,.

M1) ll'f:fiijjl~O):II~Ii~l'fm:iBS~~H:.J: 7.1.

~22*

s

s

-t'O)I:l:l8~ft0)~5J'O)=ttl:O)fillg-i>N~~:h.~~G

f~P.

UIJ