Graduate School of Science and Technology
Degree Programs in Life and Earth Sciences
List of research fields（Doctoral Programs）
Doctoral Program in Biology
Field of Research

Faculty

Systematics andEvolutionary Biology

ISHIDA Ken-ichiro

Detailed Description of Research Field①②

Classification of micro-and macro-algae based on ultrastructure andmolecular phylogenetic analyses
The endosymbiotic acquisitions and evolution of plastids
Searching for new useful algae for algal biomass research
Taxonomy of reptiles based on morphological data
Molecular phylogeny and biogeography of reptiles and birds
Conservation genetics of amphibians and reptiles
Evo-Devo research of chordates
Comparative embryology of marine invertebrates, including bivalvesand echinoderms
Natural history and biodiversity of the Kingdom Fungi
Taxonomy and phylogenetic studies of the basal lineage of Fungi(Zygomycota and Chytridiomycota)
Fungal ecology focused on their interactions with other organisms andtheir life cycles
Natural history of placozoans, xenacoelomorphs, and echinoderms
Origins and evolution of deuterostomes and metazoans
Diversity and evolution of marine invertebrates
Classification of protists including microalgae based on ultrastructuralcharacters and molecular phylogenetic analyses
Searching for new useful algae for algal biomass research

HIROTA Mitsuru①②

Plant response to environmental changes, perspective from ecology
Ecosystem ecology focused on carbon cycling in terrestrial ecosystem

TANAKA Kenta①

Evolutionary ecology focusing on ecological and genetic adaptive
mechanisms in wild Arabidopsis
Conservation ecology in mountains, grasslands and forests

HONDA Masanao

WADA Hiroshi

DEGAWA Yousuke

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NAKANO Hiroaki

NAKAYAMA Takeshi

Ecology

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Population genetics/genomics and population demographic inferenceof several species (e.g. trees, fishes, insects, mammals)
Ecosystem management and conservation using molecular ecology
approaches
Impact of human activities on forest ecosystems and their history

TOQUENAGA Yukihiko①②

Experimental ecology with field and laboratory populations
Theoretical biology with mathematical models

OHASHI Kazuharu①

Foraging behavior of pollinators with special reference to their
cognitive abilities
The evolution of floral phenotypes via ecological interactions withflower visitors
Behavioral ecology and evolutionary ecology in terrestrial arthropods
Geographic variation in behavior and ecology, and speciation

TSUDA Yoshiaki②②
SATO Yukie①②

HARVEY Benjamin①②

AGOSTINI Sylvain①②

Marine community ecology, utilizing field surveys, aquarium
experiments and modelling.
Understanding the effects of environmental change on coastal
systems, such as ocean acidification, global warming, and marine
heatwaves
Marine ecophysiology, especially of scleractinian corals
Responses of marine organisms to climate change and anthropogenic
stressors

1Plant Physiology andDevelopmental
Biology

SUZUKI Iwane①②③

IWAI Hiroaki①②

Animal Physiology
and Developmental
Biology

SUZAKI Takuya① Molecular genetic studies on root nodule development during legumeRhizobium symbiosis② Studies on molecular mechanism of nitrogen nutrient response in plants

MAEDA Yoshiaki①②③

MINODA Ayumi① Studies on regulation of primary metabolism in algae as unicellularplant model systems② Studies on metal metabolism in photosynthetic organisms

IRVING Louis John①②

Effect of nutrient status on the host – parasite interaction
Exploring the influence of abiotic factors on grass plant growth andcompetition※ KOBAYASHI Satoru①②③

Common mechanisms regulating germline formation in animals
Genetic pathway regulating sex determination of germline in
Drosophila
Mechanism regulating germline-stem-cell maintenance in Drosophila

SASAKURA Yasunori

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Developmental mechanisms of animals
Metamorphosis of ascidians
Molecular biology of mimicry
Development and Evolution

CHIBA Chikafumi①②

Molecular mechanism of adult newt body-part regeneration
Molecular mechanisms of injury responses and injury-caused disorders
in mammalian tissues
Induction and regulatory mechanisms of transdifferentiation③
NIWA Ryusuke①

Functional analyses of genomes and chromosomes in algae
Production of useful metabolites by metabolic engineering in algae
Digital transformation of algal research④

Molecular, cellular, and systemic mechanisms of the interaction
between insects and parasitoid wasps
Mechanisms of interorgan communication in the regulation ofdevelopment, stem cell proliferation, post-mating responses, and aging
Studies on molecular mechanisms of cancer cachexia using Drosophila
as a model
Structural biology and chemical biology of insect growth control agents

YAGUCHI Shunsuke①②③

Axis specification/formation of the sea urchin embryo
Development of the serotonergic neurons in the sea urchin embryo
Evolution of the anterior neuroectoderm

OKAMOTO Naoki①②

Insect hormones and its regulation during development
Neuro-endocrine control of physiology and behavior in insects

SAKURAI Keisuke① Electrophysiological studies on molecular mechanisms of signaltransduction in retinal neurons② Studies on non-visual photoreceptor cells in CNSINABA Kazuo①②③

Structure, motility, and regulation of cilia and flagella
Diversity of cilia and eukaryotic evolution
Mechanism of fertilization and reproduction of marine organisms(protists, marine invertebrates and fishes)

CHIBA Tomoki①②③

Genetic analysis of selective protein degradation
Cell biology of the ubiquitin family
Knockout mice analysis of the ubiquitin system②③

Molecular andCellular Biology

Photosynthetic acclimation and signal perception to environmental
stress
Production of useful metabolites by metabolic engineering in algae
Application of quantum beams and nuclear resonance reaction forbiological breeding
Cell wall functions on the plant development and environmental
responses
Mechanisms of plant cell wall cross-linking

2

Molecular andCellular Biology

NAKANO Kentaro①②③

MIURA Kenji①②

ISHIKAWA Kaori

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SHIBA Kogiku①②

TSURUTA Fuminori①②③

Genomics andBioinformatics

Advanced Cellular
Biology

Investigation of signal transduction controlling cytoskeleton andmembrane dynamics
Studies on the molecular diversity and evolution of cytoskeleton and its
regulatory systems
Molecular biology of the mechanisms of cell division using yeast andprotistSignaling mechanisms for abiotic stress response and sugar
accumulation in plants
Production of useful proteins (pharmaceutical proteins, etc.) with plantbiotechnology
Production and evaluation of genome editing crops
Analyses of influences by mutations of mitochondrial DNA on cellular andphysiological functions
Studies on the interactions between nuclear-coded genes andmitochondrial functions
Investigation of disease mechanisms of mitochondria-related diseases
using model animals
Studies on regulatory mechanism of sperm motility in marineorganisms
Studies on regulatory mechanism of flagellar and ciliary motility inmarine organisms
Molecular basis of the developing brain regulated by microglia
Neuron-glia communication coordinating the brain environment in the
neonatal period
Mechanisms of the architecture of neural circuits influenced byenvironmental stresses

HIRAKAWA Yoshihisa①②③

Plastid evolution via secondary endosymbioses
CO2 fixation in microalgae
Genome evolution in microalgae

INAGAKI Yuji①②③

Molecular phylogeny of eukaryotes
Evaluation of the impact of lateral gene transfer to genome evolution
Estimation of protein functions combining evolutionary parameters
and tertiary structures

KUWAYAMA Hidekazu

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Molecular analysis of biological soliton in multicellular movement
Functional analysis of a genetic disease in intracellular signaling pathway
Memory of cell and spatio-temporal pattern recognition
Analyses of a novel anti-tumor factor and the mechanism of caffeinedependent enhancement of anticancer drugs

NAKADA Kazuto①②③

Functional morphology of mammalian mito-chondria
Generation of mouse models for mitochondrial DNA-based diseases
Therapeutics for mitochondrial DNA-based diseases

SAWAMURA Kyoichi

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Evolutionary Genetics
Genetic analysis of hybrid inviability and sterility in Drosophila
Genetic analysis of sexual isolation in DrosophilaInterspecific introgression in natural populations of Drosophila

HARADA Ryuhei①②③

Computational Biophysics and Theoretical Biology
Molecular dynamics simulations for analyzing biological functions
In silico drug design based on molecular simulations

NAKAYAMA Takuro①②

Symbiogenesis in unicellular organisms
Genomic research on evolution and diversity of protists*NAGAMUNE Kisaburo(NIH, Tokyo)①②①

Understanding the infectious mechanism of parasitic protozoa
Study about the unusual organelle of parasitic protozoa
Basic research for the development of anti-parasitic drug

*SHITARA Hiroshi(IGAKUKEN, Tokyo)①②③

Molecular genetics of mitochondrial DNA in mammals
Generation of new mouse strains using transgenic technology
Imaging techniques for visualizing mitochondria in mammals

3

Advanced Cellular
Biology*MATSUI Hisanori(Takeda Pharmaceutical
Company, Ltd. Fujisawa)*YABUKI Akinori(JAMSTEC, Yokosuka)

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Advanced Molecular
Biology*NORIO Tanaka(National Museum of
Nature and Science)

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Phylogenetic and taxonomic studies of aquatic vascular plants
Adaptive evolution of aquatic vascular plants
Distribution and geographic genetic structure of aquatic vascular plants
Conservation studies of rare and threatened aquatic species
Research on ex situ conservation in botanical gardens*HOSAKA Kentaro(National Museum of
Nature and Science)①*MASAKI Takashi(FFPRI, Tsukuba)①②③

Taxonomy, phylogenetics and biogeography of fungi, especiallymushrooms
Fungal diversity in the environment (soil, water and air)Natural history of fungi based on museum specimens, DNA and othermetadata
Population ecology of woody plans
Structure and dynamics of forest ecosystem
Growth management of forests*TAJIMA Yuko
(National Museum of
Nature and Science)①②③

Life history on marine mammals
Comparative morphology on marine mammals
Health assessments on marine mammals*CHIBA Youko(RIKEN, Wako)①②③

Search for novel metabolisms in microorganisms. (Prokaryote)
Diversity of CO2 fixation and amino acid synthetic pathways
Analysis of metabolic evolutionary by physical chemistry*FUJIWARA Sumire(AIST, Tsukuba)①②③

Basic studies of transcriptional regulation mechanisms in higher plants
Research and development of useful plants by modifications oftranscription factors or genes
Functional analyses of transcription factors in higher plants①②③

Research and development of biomass utilization process
Research and development of symbiosis based biotechnology
meta- and single-cell transcriptome analysis*MORIYA Shigeharu(RIKEN, Yokohama)※

Drug discovery research in the field of neuroscience,endocrinology(particularly neuroendocrinology and reproductive endocrinology,
and drug repurposing
Translational research for drug discovery
Diversity and classification of microbial eukaryotes
Ecological function and role of microbial eukaryotes in ocean
Diversity and functional evolution of RNA-editing and its related
phenomena in microbial eukaryotes
Monitoring of the diversity of microbial eukaryotes on ocean
environmental changes②③

The faculty member marked with ※ will be retired by March 31,2026.

Note: *Adjunct Professor of the Cooperative Graduate School4Doctoral Program in Agricultural Sciences＜Biosphere Resource Science and Technology＞
Field of Research
Plant Breeding

Animal Science
Genetic Science Field

Plant Genome
Sciences

Faculty
YOSHIOKA Yosuke

ASANO Atsushi

（

＊

）

Detailed Description of Research Field①

Study on conversation and efficient utilization of genetic
resources②③④

Genetic analysis of important traits in crops
Pollination biology for seed multiplication of crops
Development of digital phenotyping method①

Integrated physiology of homeostatic functions useful foranimal production②

Study on molecular and cellular basis for fertilization anddevelopment in model animal③

Development of reproductive and genomic
biotechnologies for livestock production①

Exploring molecular mechanism for fruit development intomatoIdentification of genes related to important breeding traits incrops and horticultural plants by genome analysis.
Rapid and efficient development of new cultivars by genome
editing technology.
Identification and characterization of genes controlling levels offunctional materials by metabolic and genome analysis of largescale tomato mutant population.
Innovation of gene modification technology by modified
CRISPR/Cas9 system.

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Olericulture andFloriculture

FUKUDA Naoya①

KANG Seung Won
NONAKA Satoko②③④

Horticulture and Crop Production Field⑤

Pomology andPostharvest
Physiology of
Fruit

SUGAYA Sumiko

Crop Science

MATSUKURA Chiaki①

Physiology of fruit during pre- and postharvest②

Environmental and chemical growth regulation on fruit
trees
Propagation of woody plants③

WANG Ning

Molecular and physiological dissections of useful traits
involved in agricultural production in vegetables andornamentals
Development of new genetic engineering technologies andnovel high quality varieties in vegetables and ornamentals
Development of information technologies for vegetables
and ornamentals production
Study of Sophisitication for transformation and genome
editing technology
Study for introducion of new traits into Cucubitaceae,
Solananceae, and Asteraceae via new breeding
technology, CRISPR/Cas9.

①

Comparative studies on the efficient crop production
systems and its management②

Establishment of sustainable crop production systems andits assessment③

Physiological and ecological researches for yield andquality of crops
Physiological research on the mechanisms and control ofstress tolerance in crop④⑤5Analysis of gene function for critical agronomic traits incrops

Plant Parasitic
Mycology

OKANE Izumi①

Plant Protection Field②③

AppliedEntomology and

FURUKAWA Seiichi

Zoology

ForestEcotopology①②③

KAMIJO Takashi
KAWADA Kiyokazu

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Forest Environmental Science Field

Conservation ofRegional
Resources

SEINO Tatsuyuki①

Systematics of plant parasitic fungi including symbiotic fungi,
particularly rust fungi, blue stain fungi, endophytes andmycorrhizal fungi.
Studies on ecology and physiology of these fungi and bacteria.
Functional analysis of genes associated with disease resistance inplant.
Insect immune mechanisms against pathogens and parasitoids
Elucidation of strategies of parasitoids to survive in the host
insect species
Improvement of biological control of insect pests
Dynamics and function of forest ecosystem
Vegetation science and management
Conservation and restoration of arid and semi-arid
ecosystem
Conservation of endangered species②③

Conservation genetics of tropical tree species, andphylogeography of forest tree species and genetic study of local
adaptation
Study on conservation of regional resources
Wildlife management and biodiversity conservation

TSUDA Yoshiaki

Environmental
Soil Chemistry

ASANO Maki①②③

Environmental chemistry of soils
Ecological studies on soil organic matter
Soil conservation under semi-arid grasslands

Environmental
Plant
Biochemistry

YAMAJI Keiko①

Effect of endophytic microbes on heavy-metal stress tolerancein plants
Effect of endophytic microbes on environmental stress
tolerance in plants
Effect of endophytic microbes on radio Cs accumulation inplants②③

Plant Molecular
Biology

SHIBA Hiroshi

Advanced Interdisciplinary Agricultural Science Field①

Molecular mechanisms of epigenetic regulation inheterosis②

Molecular mechanisms of epigenetic regulation in sexual
plant reproduction
Epigenetic engineering of plant development③
Metabolic
Network Biology

KUSANO Miyako①②③

Epigenetics

BUZAS Diana Mihaela①

Molecular genetic analysis of the perennial life history inArabidopsis halleri gemmifera②

Molecular ecology analysis of seasonal response in Wasabi
japonica
Dissection of memory DNA function in overwintering in
crucifers③

Plant Cell and
Synthetic
Biology

KINOSHITA Natsuko

Development of analytical platforms to capture
quantitative and qualitative changes of metabolite levels
Metabolic network biology using “omics” datasets
Flavor analysis of important crops and vegetables

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6

Plant and insect interaction
Production of high added value products in plants
Visualization of plant environmental response mechanisms

Agro-biological Sciences Field

Plant Stress
Biology*FUJITA Yasunari①(Japan International Res. Center
for Agricultural Sci. (JIRCAS))②

Molecular elucidation of stress tolerance mechanisms inplantsDevelopment of environmental stress-tolerant crops

Climate Change
ImpactAssessment on
Vegetation*MATSUI Tetsuya(Forestry and Forest Products
Research Institute(FFPRI))①

Relations between distributions of forest vegetation andclimatic conditions
Impact assessment and adaptation planning of climatechange on forest ecosystem functions and ecosystemservices

Tropical Forestry*TANI Naoki①(Japan International Res. Center
for Agricultural Sci. (JIRCAS))②

AnimalFunctional
Biology*SAKUMOTO Ryosuke(Institute of Livestock andGrassland Science, NARO)

Insect Functional
Regulation*TABATA Jun(Institute for Plant Protection,
NARO)

Environmental
Agronomy①*MINAMIKAWA Kazunori(Japan International Res. Center②
for Agricultural Sci. (JIRCAS))

②

①
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③
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Improvement of tropical forestry using indigenous genetic
resources in Southeast Asian tropical forest
Reproductive biology in Southeast Asian tropical forest andits application to sustainable forest management
Factors involved in the animal productive functions
Study on animal reproductive biology, especially on theestablishment of pregnancy and its maintenance in ruminants.
Development of effective technique to improve reproductive
performance of domestic animals
Chemical ecology of insects and associated plants
Development of insect functional regulation techniques
based on chemical ecological studies

Development and assessment of climate change mitigation andadaptation technologies in rice production
Observation and modeling of greenhouse gas emission andcarbon and nutrient cycling in rice production(＊) Please contact the Chair of the Doctoral Program in Agricultural Sciences (e-mail: sugaya.sumiko.fw#@#u.tsukuba.ac.jp) in
regard to this research field. (*Replace “#@#” with “@”.)*Adjunct professor of the Cooperative Graduate School (not assigned an academic advisor’s position for research students[kenkyusei]).

7Doctoral Program in Agricultural Sciences＜Appropriate Technology and Sciences for Sustainable Development＞
Field of Research
Environmental Colloid andInterfaceEngineering

Faculty
KOBAYASHI Motoyoshi

Detailed Description of Research Field①

SUGIMOTO Takuya②③

Eco-region Development Engineering Field

Bio-resource
Process andSystemEngineering

（

＊

）

①

②

Water and solute transportation in soil. Salinity anderosion of soil
Water resource engineering in arid land, water quality
control, water treatment
Physics and chemistry of soil, soil pollution soil, colloid andinterface
Resource and energy utilization using agricultural waste,biomass and organic wastewater based on bio-resource
recycling system
LCA, LCC, and simulator development for optimization
design of bio-resource conversion process and grasping of
biomass potential and its utilization

Watershed
Conservation

NASAHARA(NISHIDA) Kenlo①②③

Mechanism of sediment production and transport
Sabo planning in harmony with natural environment
Environmental analysis through remote sensing

Water
Resources
Management
Engineering

ISHII Atsushi①②③

Development and management of irrigation systems
Water resources evaluation for development
Participatory irrigation management①

Intelligent machinery and robotics for agricultural
production
System analysis for bioenergy production and utilization
Real-time crop monitoring systems for site-specificmanagement

ASADA Yohei

Bioproduction
andMachinery

Tofael AHAMED

Farmland
SystemEngineering

KOBAYASHI Motoyoshi

Food
Resources
Engineering②③

Food and Biomass Science Field①②

Farmland engineering, soil conservation engineering
Soil Physics, Environmental materials①②③

Micro/nano-engineering for advanced bioresource processing
Microchannel technology for advanced food processing
Formulation of food micro/nano-dispersions andevaluation of their gastrointestinal digestion
Effective utilization of food processing waste for value
addition

YAMASHITA Yuji
Marcos Antonio das
NEVES

④

Agri-Food
Process
Engineering※KITAMURA Yutaka①②

Milling and spray drying for health food production
Development of novel food by applying rice slurry

Chemistry of
Biomaterials

NAKAGAWA-IZUMI Akiko①②③

Chemistry for wood pulping and pulp bleaching
Chemical utilization of biomaterials and bio-refinery
Micro-analysis of wood components (lignin,tannin,carbohydrate and others) and the related
compounds

8

Food and Biomass Science Field

EngineeringofBiomaterials①②

OBATAYA Eiichi

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AgriculturalandBioresource①

SHUTO Hisato②

Economics①

Rural Development Economics Field
Eco-region Development Engineering Field

Resource
Economics
andDevelopment
Studies

SHUTO Hisato

Farm
Business andAgribusiness
Management

UJIIE Kiyokazu

ForestResource
Economics（

ForestResources
Sociology

KOHROKI Katsuhisa

Rural
Environment*MIYAMOTO Teruhito

Improvement

Nano andMicro-scale
Food
Analysis＊）Analysis of food industries with specific attention to issuesof productivity, R&D, scale economies, and economics oforganization
Economic analysis of agricultural and food security policies②

International trade analysis of agricultural commodities
and resources
Community development and resource management①②③

Farm production and supply economics under the risk
Farm and agribusiness firm management and marketing
Food consumption and consumer policy①②

Study on forest economics and policy
International comparative study on management andutilization of forest resources
International comparative study on production andMarketing of forest products

③

①
②
③

*YOSHIMOTO Shuhei(Institute for RuralEngineering, NARO)

Creation of paper-based electronics and sensors
Conservation of aging library collection and flooddamaged paper cultural heritage using salt water
immersion method
Synthesis of fluorine containing condensation polymers forcomposite materials
Chemical modification of poly (amino acid)s and polysaccharides
Property enhancement of biomaterials for highperformance musical instruments
Investigation on the mechanical properties of wood withrespect to its fiber-reinforced cellular structure, anddevelopment of technology for their effective utilization
Physical and chemical characterization of natural
adhesives such as Japanese lacquer and chitosan, anddevelopment of technology for their utilization①②③*MANO Junichi*GENKAWA Takuma①(Institute of Food
Research,NARO)②

9

Historical study of forest management in Japan
Socioeconomic study on regional forest management inJapan
Comparative study on forestry organizations
Irrigation and drainage management in farmland
Modeling, measurement and interpretation ofmass and energy flow in soil
Hydrological investigations and conservation ofgroundwater resources in rural areas

Development of analytical methods for evaluating foodquality
Development of food processing methods using
biotechnology

Food and Biomass Science Field

Sustainability ofBiomass
Resources

Regional Forest
Resource
Development

Development of biomass utilization technology using
microbialfunction*KOSUGI Akihiko(Japan International Res.
Center for Agricultural
Sci.(JIRCAS))*YAMADA Tatsuhiko(Forestry and Forest Products
Research Institute(FFPRI))①②③

Development of lignin based functional bio-materials
Chemical conversion of cellulosic biomass for preparing usefulchemicals, liquid fuels and fuel additives
Rapid analysis of lignocellulosics to evaluate potential of forest
biomass

Agricultural Economics and Sociology Field

International
Agriculture andForestry
Development*IIYAMA Miyuki(Japan International Res.
Center for Agricultural
Sci.(JIRCAS))①

Regional Forest
Resource
Development*ISHIZAKI Ryoko(Forestry and Forest Products
Research Institute(FFPRI))①②

Identification of social conflicts over forest resources
Study on how to lead rural development by utilizing
forest resources①②

Agricultural Workforce and Human Resource Development
Local Agricultural Support Systems to Revitalize Rural
Communities

Farming System* S AWA DA M a m o r u②

Trends and prospects of international agriculture research
agendas on global food systems.
Sustainable agricultural intensification of smallholder systems.

※Faculty members due to retire in March 2027(＊) Please contact the Vice Chair of the Doctoral Program in Agricultural Sciences (e-mail: ishii.atsushi.fu#@#u.tsukuba.ac.jp) in
regard to this research field. (*Replace “#@#” with “@”.)*Adjunct professor of the Cooperative Graduate School (not assigned an academic advisor’s position for research students[kenkyusei]).
(* E-mail address: add following domain name:@u.tsukuba.ac.jp . Or replace “#@#” with “@” . )10

Sub-Program in Advanced Agricultural Technology and Science cooperated with NARO*
Field of ResearchInnovative Crop
Production andQuality control
SystemFaculty(e-mail address)

Detailed Description of Research Field

TANAKA Tsuyoshi①(tstanaka#@#affrc.go.jp)②

MITSUNAGA Takayuki(aeiou#@#affrc.go.jp)③

KIMURA Toshiyuki

Comparative genomics for breeding and molecular
evolution study
Study on construction of pest-occurrence-prediction
system using statistical modeling
Study on application for agricultural research by using
LC-MS/MS(kmr#@#affrc.go.jp)
FUKATSU Tokihiro①(fukatsu#@#affrc.go.jp)②

SUGIURA Ryo(rsugiura#@#affrc.go.jp)③

TANAKA Daisuke

Farming systems to reduce labor, production costs andenvironmental loads
Research on remote sensing technology and image
processing technique for agricultural applications
Studies on cryopreservation of genetic resources
as a long-term conservation technique(dtanaka#@#affrc.go.jp)
Innovative Animal
Production System

SASAKI Keisuke①(ksuk#@#affrc.go.jp)②

TOHNO Masanori(tohno#@#affrc.go.jp)

Innovative Crop
Breeding andCultivation System

TANAKA Junichi(tanajun#@#affrc.go.jp)
MATSUI Katsuhiro(matsuik#@#affrc.go.jp)①②③

TAGUCHI Kazunori

Studies on measurement and improvement of quality, sensory
traits, and consumer satisfaction of animal products
Exploring the individual roles of plants, microbes, andanimals and their interactions in animal production
systems.

Developments of new crop breeding methods using
genome information
Study on efficient trait improvement of resource crops
and underutilized plants
Breeding and genetics of high performance varieties
by heterosis in root and tuber crops(ktaguchi#@#affrc.go.jp)
TATSUKI Miho①

(tatsuki#@#affrc.go.jp)②

KUNIHISA Miyuki

Study on fruit maturing, senescence and postharvest
technology of fruit tree
Research on the application of mass data for genomes inapple breeding(miyuky#@#affrc.go.jp)

ONOZAKI Takashi(onozaki#@#affrc.go.jp)①②

SHIBUYA Kenichi(shibuken#@#affrc.go.jp)③

TAKAHASHI Megumu(mtakahashi#@#affrc.go.jp)*NARO=National Agriculture and Food Research Organization

11

Breeding for disease resistance and improvement offlower vase life in ornamental crops
Study on regulatory mechanism of flower senescenceand postharvest technology in ornamental plants
Developments of stable production system for open-field
vegetables

Doctoral Program in Life and Agricultural Sciences(*Replace “#@#” with “@”.)Field of Research
Biochemistry of
Bioactive
Molecules

Faculty
USUI Takeo
SHIGEMORI Hideyuki
SUNOHARA Yukari
FURUKAWA Jun
YAMADA Kosumi
MATSUYAMA Shigeru

Detailed Description of Research Field
①
②
③

④

⑤
⑥
Chemical Life Science⑦⑧⑨Structural
Biochemistry

TANAKA Toshiyuki①②③

Functional Foods
and Food
Chemistry

YOSHIDA Shigeki①②③

Genomic Biology

TANIMOTO Keiji
ISHIDA Junji
KAKO Koichiro
DAITOKU Hiroaki①②③

Animal Life Science

Molecular andDevelopmental
Biology

KASHIWABARA Shin-ichi①②③

Biology for Gene
Regulation

KIMURA Keiji①②③

Identification of molecular targets of the bioactive compounds inmammalian and plant cells and their action mechanisms
Antioxidative responses to oxidative stresses
Elucidation of the molecular mechanisms of bioactive substances
involved in biological phenomena of plant (germination,
phototropism, gravitropism, senescence, etc.)
Search for bioactive compounds related to prevention of diseases(Alzheimer's disease, diabetes, osteoporosis, etc.) from edible
and medicinal plants)
Mechanisms how to accumulate various metals in plants
Signaling mechanisms about nutrient status in the organ to organinteractions in plants
Functional and structural analysis of plant growth regulators in
response to phototropic and gravitropic stimulation
Functional and structural analysis of extracellular plant
metabolites associated with allelopathy and their application inplant production
Semiochemicals mediating interactions among insects, plants andanimals
Analysis of the structure-function relationships of proteins
involved in signal transduction and transcription regulation
Analysis of the chromophore-protein interactions ofchromoprotein antitumor antibiotics
Protein engineering based on detailed structural information on
functional proteins
Structure and function of bioactive compounds in food
Production of bioactive compounds by using bioconversion
process
Development of industrial enzymes for food production
Molecular mechanisms of aging regulated by transcription andmetabolism
Protein methylation and its modifying enzymes involved in
hypertension
Mammalian epigenetics in genomic imprinting and gene
regulatory mechanisms in blood pressure homeostasis
Transcriptional and translational regulation of genes during
gametogenesis
Functional roles of proteins involved in fertilization, egg
activation, and early embryonic development
Development of reproductive and developmental technologies
for future life
Analysis for dynamics of mitotic chromosomes.
Analysis for function of condensin complex.
Analysis for novel function of the nucleolus.

12

Molecular
Microbial
Bioengineering※KOBAYASHI Michihiko
HASHIMOTO Yoshiteru

①
②
③
④
⑤

Screening of new metabolism, and functional analysis ofphysiological functions.
Metabolic engineering and screening/ analysis /design/
remodeling of useful enzymes and genes.
Functional analysis of enzymes involved in cleavage and synthesisof a C-N bond and their molecular evolution.
Development of super biological catalysts with novel functions of
microorganisms and their enzymes.
Functional analysis of nucleic acid-related enzymes and its
application to DNA/RNA engineering.

Applied Microbiology

AppliedMicrobiology

NOMURA Nobuhiko
UTADA S. Andrew
TOYOFUKU Masanori

①
②
③
④
⑤

Bacterial cell- cell communication and biofilm formation
Microfluidics for analysis of bacterial communities
Biophysical analysis of biofilm formation
Bacterial interactions through membrane vesicles
Molecular microbiology of environmental bacteria and theirapplications

Ecological
Molecular
Microbiology

TAKAYA Naoki
NAKAJIMA-KAMBE
Toshiaki
NAKAMURA Akira
YING Bei-Wen
TAKESHITA NorioYAWATA Yutaka①

Environmental response and morphogenesis of filamentous
fungi
Enzymology and molecular biology of microbial enzymes
Bacterial metabolisms and communication
Screening of novel microorganisms/genes with useful functions
and their engineering
Fermentative production of useful compounds from waste
biomass by metabolic engineering
Study on microbial catabolic pathway of L-form sugars
Development and application of host-vector system in Thermus
thermophilus
Experimental evolution for investigating the microbial survival
strategies
Multilevel omics analyses of the genome reduced Escherichia coli
Physiological functions of sulfur-containing amino acids and its
applications

②
③
④
⑤
⑥
⑦
⑧
⑨
⑩

Cell Cultivation
Engineering①

AOYAGI Hideki②③④
Biochemical Engineering

Bioreaction
Engineering

Biomimetic
Chemistry

ICHIKAWA Sosaku
HIRAKAWA Hidehiko

（

＊

）

Development of cultivation system for cell and protoplast withnovel functional activities and their biotechnological application
Analysis of naturally-occurring microbial symbiotic association,
construction of artificial symbiotic system and their applicationfor various bioprocesses
Cell cultivation engineering and development of novel bioreactors
Development of cultivation system for uncultured microbes(microbial dark matter), animal cells, and plant cells and theirbiotechnological application

①
②
③
④

Application of polymolecular aggregates for bioprocesses
Production of useful materials by enzymes and microorganisms
Development of tools for selective protein conjugation
Interdisciplinary studies for practical use of cytochrome P450s①②③

Enzyme isomerism leading chiral homogeneity
Characterization of polyelectrolyte complex
Polymer chemistry for exploration and simulation of biological
functions13Animal Life Science

AnimalBioresource
Engineering*INOUE Kimiko( RIKEN )①②③

Molecular
Neurobiology*DOI Motomichi( AIST )①②③

Applied Microbiology
Biochemical
Engineering
工学領域

Applied
BioengineeringofMicrobial
Ecosystems*TAMAKI Hideyuki( AIST )Evolutionary
Biology ofSymbiosis*FUKATSU Takema( AIST )Food MolecularEngineering*KOBORI Toshiro

①
②
③

①
②

（Institute of Food
Research, NARO)①②

Characterization of the germ cell genome using a nuclear transfer
techniqueAnalysis of the mechanisms for zygotic gene activation using a
nuclear transfer technique
Development of techniques for preservation of male germ cells
using microinsemination
Molecular analysis of nervous-system formation and maintenance
Development of screening systems for neuronal dysfunctions anddiseases
Development of live-cell imaging methods using fluorescent andluminescent techniques
Culturing the uncultured fastidious microorganisms in theenvironment and exploring their novel biological functions
Omics-driven discovery of novel microbial and genetic resources
Ecophysiology and diversity of uncultured microorganisms in theenvironments (gut, plants, deep subsurface, etc.)

Biological function, evolution and origin of endosymbiotic
associations between insects and microorganisms
Molecular, physiological and regulating mechanisms underlying
sophisticated inter-organismal interactions in symbiosis,
parasitism, manipulation and sociality
Screening and utilization of biomolecules for sensing food quality
Analyses on structure-function relationship of advanced glycationand products※Faculty members due to retire in March 2027(＊) Please contact the Chair of Doctoral Program in Life and Agricultural Sciences (e-mail: usui.takeo.kb#@#u.tsukuba.ac.jp) in
regard to this research field. (*Replace “#@#” with “@”.)*Adjunct professor of the Cooperative Graduate School (not assigned an academic advisor’s position for research students[kenkyusei]).
(NARO) = National Agriculture and Food Research Organization(RIKEN) = RIKEN(AIST) = National Institute of Advanced Industrial Science and Technology

14Doctoral Program in Bioindustrial Sciences
Research
Field
GenomeBiology

Faculty

Specialized Field※2 NAKAMURA Kouji

Molecular mechanism of protein secretion, Functional
analysis for functional RNA gene

KIKUCHI Akira

Stress physiology, Molecular breeding, Somatic embryogenesis※1 ONO Michiyuki

Molecular biology of photoperiodism and flower induction,
Plant biotechnology of developing new flower (color, shapeand more) and edible vaccine

Bioprocess
Engineering

NOMURA Nakao

Development of sustainable agriculture, Forestry andfisheries industry using bioengineering technique

Bioactive
NaturalProducts
Chemistry

SHIGEMORI
Hideyuki

Naturally occurring bioactive substances, Phototropism,
Gravitropism, Flowering, Apical dominance, Allelopathy,
Plant growth regulators, New drugs from unexplorednatural resources, Preventive medicines of lifestyle-relateddisease, Environmental preservation-type functional agentsMIYAMAE Yusaku

Molecular tools for control of cellular protein stability,
Establishment of drug screening system, Identification andmechanism elucidation of nuclear receptor ligand,Chemical biology on natural products

YAMADA Kosumi

Plant growth regulators, Phytohormones, Environmental
response in plants, Chemical communication in plants

NAKAJIMA-KAMBE
Toshiaki

Isolation and screening of microorganisms with potential
for bioproduction/biotransformation. (plastic degradation,
biotransformation of oil/fat-related biomass, and methane
conversion)

AOYAGI Hideki

Analysis of naturally-occurring microbial symbiotic
association, construction of artificial symbiotic system andtheir application for various bioprocesses※1 WATANABE N.
Kazuo

Biodiplomacy, Assessment of biodiversity, Sustainable use
for genetic resource, Biosafety, Access for bioresources andits appropriation

OGUCHI Taichi

Plant biotechnology, plant physiology on environmental
response, Environmental and health risk assessment ofbiotech plants, Detection method on biotech foods

Animal Cell
Biotechnology

ITO Yuzuru

Eco-system
Technology
Area

BioEnvironmental
ControlEngineering

UTSUMI Motoo

Basic technology of the regenerative medicine using humanstem cells (Quality control, mass cultivation,differentiation)
Drug discovery support technology (organ celldifferentiation for drug discovery using knowledge fromregenerative medicine, microphysiological system)
Functional analysis of marine microorganisms and its rolein cycling of matter, Bio eco-engineering

Bioresource
Development
Technology
Area

Food System※2 KITAMURA
Yutaka
KOKAWA Mito

Post-harvest technologies, Processing of functional foods,
Conversion and utilization of biomass and food waste, Nondestructive analysis of food quality using light

Biological andMaterial Cycles

YANG Yingnan

Photocatalytic technology, Solar light utilization system,
Bioreactor, High efficiency conversion and effective
utilization of bioresources, Renewable energy

Genetic
Resource
Science andTechnology
Area

Bioindustry
andBioscience
Area

Plant
Biotechnolo
gy on
Abiotic
StressesPlant Physiology
andBiotechnology

Plant
Physiology
andChemistry
Industrial
Microbiology
andBioresource
Science

Bioindustrial
Resources

15

Genetic
Resource
Science andTechnology
Area

Industrial
Sciences for
Genetic
Resources

Comparative genomes (plants), Environmental stressresponses, Transcriptional regulatory networks, Plant
genetic resources*MARUYAMA
Kyonoshin(JIRCAS)

Development of biotechnology using electrogenic bacteria,
Data-driven chemical biology research using a highthroughput electrochemical system, Development of
resource recovery technology using the interaction
between materials and bacteria

Electrochemical *OKAMOTO
Biotechnology
Akihiro(NIMS)※1 Faculty menbers due to retire in March 2026
※2 Faculty menbers due to retire in March 2027
Note: *Adjunct Professor of the Cooperative Graduate School(JIRCAS) = Japan International Research Center for Agricultural Sciences(NIMS)= National Institute for Materials Science16Doctoral Program in Geosciences
Field of
Research
Human
Geography

MATSUI Keisuke
jiji@geoenv.tsukuba.ac.jp

Cultural geography, Geography of tourism and religion, Theory ofcultural tourism

Regional
Geography

KUREHA Masaaki
mkureha@geoenv.tsukuba.ac.jp

Regional geography of Europe and Japan, Geography of tourism

Faculty

Detailed Description of Research Field

Regional geography of Australia, Urban geography, GIS
TSUTSUMI Jun
jtsu@geoenv.tsukuba.ac.jp
Geomorphology

Hydrological
Science

Atmospheric
Science

Geographical
Information
Science

IKEDA Atsushi
aikeda@geoenv.tsukuba.ac.jp

Cold region geomorphology, Permafrost monitoring, Mountainenvironments

HATTANJI Tsuyoshi
hattan@geoenv.tsukuba.ac.jp

Hydrogeomorphology, Landslides, Rock weathering, Karst
geomorphology

SEKIGUCHI Tomohiro
sekiguchi@ied.tsukuba.ac.jp

Sedimentary processes,Bedform,Experiment

ASANUMA Jun
asanuma@ied.tsukuba.ac.jp

Hydrometeorology, Land-vegetation-atmosphere system,
Atmospheric Turbulence

YAMANAKA Tsutomu
tyam@geoenv.tsukuba.ac.jp

Water and material cycle, Isotopic tracer, Eco-hydrometeorology

UEDA Hiroaki
hueda.hiroaki.gm@u.tsukuba.ac.jp

Atmosphere-ocean-land interaction involved in the climate
systemUENO Kenichiueno.kenichi.fw@un.tsukuba.ac.jp

Land-atmosphere interaction and precipitation system, Mountainweather and snow cover variations＊KUSAKA Hiroyuki
kusaka@ccs.tsukuba.ac.jp

Urban climatology, Mountain meteorology, Applied meteorology(wind energy prediction, biometeorology)

Remote Sensing, GIS, Global Environment, Water Quality of Lakes
Matsushita Bunkei
matsushita.bunkei.gn@u.tsukuba.ac.jp

Analysis ofEnvironmental
Dynamics

MORIMOTO Takehiro
tmrmt@geoenv.tsukuba.ac.jp

Agricultural and Rural geography, Sustainability of agriculture
and rural area, GIS

ONDA Yuichi
onda@geoenv.tsukuba.ac.jp

Transfer of radionuclides in Environment, Hydro-geomorphology,
Forest hydrology

TSUMUNE Daisuke
tsumune.daisuke.gw@u.tsukuba.ac.jp

Oceanic material cycle, simulation of oceanic radioactivity
dynamics.

KATO Hiroaki
kato.hiroaki.ka@u.tsukuba.ac.jp

Forest hydrology, Soil erosion, Environmental radioactivity

17

Paleobiological
Science

AGEMATSU Sachikoagematsu@geol.tsukuba.ac.jp

Conodont, Graptolite, Tentaculite, Paleozoic historical geology ofSoutheast Asia

TANAKA Kohei
koheitanaka@geol.tsukuba.ac.jp

Vertebrate paleontology and paleoecology

KAMATA Yoshihito
yoshi_kamata@geol.tsukuba.ac.jp

Geological evolution of Southeast Asia

FUJINO Shigehiro
shige-fujino@geol.tsukuba.ac.jp

Sedimentology and stratigraphy, Geological records of tsunamis
in Japan and Asian countries

YAGI Yuji
yagi-y@geol.tsukuba.ac.jp

Earthquake rupture process and seismicity

UJIIE Kohtaro
kujiie@geol.tsukuba.ac.jp

Structural geology and tectonics

OKUWAKI Ryo
rokuwaki@geol.tsukuba.ac.jp

Seismic source processes of earthquakes and non-earthquakes

MARUOKA Teruyuki
maruoka.teruyuki.fu@u.tsukuba.ac.jp

Isotope geology, Geochemistry

FUJISAKI Wataru
wataru-fujisaki@geol.tsukuba.ac.jp

History of life on earth, Tectonics

TSUNOGAE Toshiakitsunogae@geol.tsukuba.ac.jp

Petrology of metamorphic rocks, Collisional orogeny, Gondwana

IKEHATA Kei
ikkei@geol.tsukuba.ac.jp

Volcanology, Geochemistry

KUROSAWA Masanori
kurosawa@geol.tsukuba.ac.jp

Mineralogy, Fluid inclusion analysis

KYONO Atsushi
kyono@geol.tsukuba.ac.jp
IIZUKA Satoshi
iizuka@bosai.go.jp

Mineralogy, Crystallography, Mineral physics

SHIMOKAWA Shinyasimokawa@bosai.go.jp

Physical oceanography, Coastal disasters, Marine ecosystem

SHUSSE Yukari
shusse@bosai.go.jp

Clouds and precipitation meteorology, Rader meteorology

OceanAtmosphere
Interaction
SystemISHII Masayoshi
maish@mri-jma.go.jp

Oceanography, Atmosphere-Ocean Interactions, Climate Variations

KAJINO Mizuo
kajino@mri-jma.go.jp

Atmospheric Chemistry, Aerosol-Cloud-Radiation Interactions

Earth
Historical
Analysis

KOHNO Naoki
kohno@kahaku.go.jp

Paleobiology of Cenozoic animals (especially for aquatic animals)

SHIGETA Yasunari
shigeta@kahaku.go.jp

Paleobiology of cephalopoda

TSUTSUMI Yukiyasu
ytsutsu@kahaku.go.jp

Geochronology

Paleogeosphere
ScienceGeodynamics

Planetary
Resource
Geology

Petrology

Mineralogy

Water-related
Disaster Science

Atmosphere-ocean interaction, Meteorological disaster,
Extreme event＊The faculty member marked with (*) can be a supervisor for students in the atmospheric science field, as well as the spatial information
science field.

18Doctoral Program in Environmental Studies
Field of Research
Sustainability
Hydrology

https://www.envr.tsukuba.ac.jp/eng/

Faculty

Detailed Description of Research Field

TSUJIMURA Maki
ASANUMA Jun
ONDA Yuichi⚫ Groundwater contamination and human activities,
monitoring of water resources and water environment⚫ Dynamics of radio nuclides in water environment Hydrogeomorphology

Environmental
Sustainable Soil
Science

YAMAJI Keiko

Environmental
Microbiology

NOMURA Nobuhiko
TOYOFUKU Masanori
NAGAKUBO Toshiki*⚫ Mode of action of agrochemicals, Stress responses of plants⚫ Chemical response of plants and microorganisms in therhizospheric soil
⚫ Applied microbiology for bioremediation⚫ Microbial control for creating functional materials⚫ Applying physics and engineering techniques to understandbacterial behavior and biofilm formation

Sustainable Recycling
of Bio-resources

LEI Zhongfang
UTSUMI Motoo
YUAN Tian⚫ Techniques for improving water quality with lower load and friendly
symbiosis to environment⚫ Recycling and reusing of agricultural wastes and development offunctional food materials from them
⚫ Development of wastewater treatment technologies based on
ecosystem engineering⚫ Aquatic biogeochemistry and engineering⚫ Aquatic environmental remediation for sustainable water use⚫ Inhibition by microbial metabolite on water purification process
and development of its measures⚫ Toxicity assessment and remediation of environmental pollutants

Environmental
radiochemistry★SAKAGUCHI Aya⚫ Development of analytical techniques for environmental radionuclides⚫ Behavior of natural/artificial radionuclides in the environment⚫ Applications of natural/artificial radionuclides as tracers forenvironmental dynamics

Global Climate
Variability

KAMAE Yoichi⚫ Cloud Feedback and Climate Sensitivity⚫ Monsoon and Global Climate Variability⚫ Paleoclimate Modeling (Pliocene)

Water Resources
Management

ISHII Atsushi⚫ Development and management of irrigation systems⚫ Water resources evaluation for development⚫ Participatory irrigation management

YOKOI Tomoyuki

⚫
⚫
⚫
⚫

HIROTA Mitsuru⚫ Distribution of terrestrial plants and their adaptive strategy toenvironments

Engineering
Biodiversity andConservation Ecology

Ecosystem Ecology

OMORI Yuko*
MASUMOTO Shota*

Life history and biodiversity of insects
Pollination service and conservation of pollinators
Behavior and life history of wild bees
Interaction between alien plants and native insects⚫ Response to environmental change in alpine ecosystem: species,
community and ecosystem components⚫ Dynamics of bioelements in ecosystems⚫ Evaluation of the effects of global change to biosphere⚫ Aquatic Biogeochemistry and Engineering

Urban LandscapePlanning★MURAKAMI Akinobu★YAMAMOTO Sachiko⚫

History of dwelling environment Conservation of traditional built
environment,⚫ Urban planning, Regional planning

19

Environmental andSocio- economic
Policies

YABAR Helmut
MIZUNOYA Takeshi★KAIDA Naoko⚫ Evaluation and analysis of technological externalities, market
failure, revealed preference in demand and common property inthe ecosystem,⚫ Methods for comprehensive evaluation of the environment
integrated waste management systems: policy and planning⚫ Remote sensing and GIS application to environmental problems
Monitoring natural environment, Environmental change predictionand environmental impact assessment, Policies for environmental
preservation, Environmental policy, Environmental economics⚫

Environmental psychology, environmental economics, proenvironmental behavior, environmental decision-making

NASAHARA Kenlo⚫ Watershed management, Environmental monitoring and disasterprevention using satellite remote sensing

Soil Environment
Conservation

KOBAYASHI Motoyoshi
YAMASHITA Yuji
SUGIMOTO Takuya
ASADA Yohei⚫ Fundamental of colloid science and its application to soil andwater⚫ Water and solute transportation in soil. Salinity and erosion of soil
⚫ Water resource engineering in arid land, water quality control,
water treatment⚫ Physics and chemistry of soil, soil pollution, and soil colloid andinterface

Sustainability
Policies andDiplomacy

MATSUI Kenichi⚫

Environmental dispute resolution and diplomacy⚫

Rural resources management and forest conservation indeveloping nations,⚫

Environmental/water ethics and law⚫

Environmental and agricultural policies for sustainability⚫

Environmental disaster policies⚫

Traditional knowledge for sustainability

Integrated
Watershed
Management

Functional foodresources

ISODA Hiroko
MIYAMAE Yusaku
TAKAHASHI Shinya
Farhana FERDOUSI⚫ Mechanisms behind functional food resources for potential
applications in food and cosmetics.
⚫ Studies on small molecules that modulate the cellular metabolism⚫ Pigment Cell Research (Melanogenesis and Melanoma Research)⚫ Search for functional components from biomass⚫ Bioinformatics, Omics Research, Clinical trial, Epidemiology

Plant physiology

SUZUKI Iwane
MAEDA Yoshiaki⚫ Photosynthetic acclimation and signal perception to environmental
stress⚫ Production of useful metabolites by metabolic engineering in algae⚫ Application of quantum beams and nuclear resonance reaction forbiological breeding

Environmental
Disaster

UCHIDA TaroYAMAKAWA Yosuke*

Prevention
Regional Air
Pollution[Cooperative
graduate school
system: National
Institute forEnvironmental
Studies]

TAKAMI Akinori
SUGATA Seiji
NAGASHIMA Tatsuya⚫ Policy of natural disaster prevention, Strategy for natural disasterprevention, Risk management against natural disaster, Risk assessmentand mitigation for natural disasters, Crisis management for natural
disasters⚫ Observation and analysis of air pollution including PM2.5 in East
Asia and study of its health and climate impact⚫ Numerical study of regional air pollutants, Material transport in
the atmosphere⚫ Studies on Asian air pollution and its effects using chemicaltransport model

Faculty members marked by * cannot be assigned as thesis director, but can advise the student under the direction of a qualified
thesis director within the same research field.
★The faculty member marked with ★ is a dual-role faculty member and cannot be a primary supervisor.
E-mail addresses of the faculty members are available on the following web site:

20

https://www.envr.tsukuba.ac.jp/eng/