Bull. Natl. Mus. Nat. Sci., Ser. A, 47(1), pp. 19–29, February 22, 2021
DOI: 10.50826/bnmnszool.47.1_19

Inventory Studies on the Subfamily Pselaphinae (Coleoptera,
Staphylinidae) of Myanmar Part 3:
A List of Collected Species in
Tanintharyi Region in November 2018Shûhei Nomura1 and Mu Mu Aung2
1

Department of Zoology, National Museum of Nature and Science,
4–1–1 Amakubo, Tsukuba, Ibaraki 305–0005, Japan
E-mail: nomura@kahaku.go.jp
2
Forest Research Institute, Forest Department, Ministry of Natural Resources andEnvironmental Conservation, Yezin, Nay Pyi Taw, Myanmar
E-mail: mumuaung85@gmail.com(Received 11 November 2020; accepted 23 December 2020)

Abstract In the course of biological inventory by FRI, Myanmar and NMNS, Japan conductedin Tanintharyi Region in Nov. 2018, 50 pselaphine species of 35 genera were recognized. The following four species are recorded from Myanmar for the ﬁrst time: Trichonomorphus ursinus Raffray, 1890, Cratna abdominalis Löbl, 1975, Pareuplectops coomani Jeannel, 1957, Megatyrus
masumotoi Nomura et al., 2010.
Key words : Pselaphinae, Staphylinidae, Pselaphinae, fauna, Myanmar.

Introduction
During the biological inventory by the Forest
Research Institute (FRI), Yezin, Myanmar andthe National Museum of Nature and Science(NMNS), Tsukuba, Japan from 2016 to 2020, a
second survey of Pselaphine beetles was conducted in November 2018. In this survey, about
490 samples of pselaphine beetles were collected
in Tanintharyi Region by the ﬁrst author. These
pselaphine specimens were identiﬁed into 50
species belonging to 35 genera (some of which
could not be identiﬁed). The following 4 species
are recorded from Myanmar for the ﬁrst time:
Trichonomorphus ursinus Raﬀray, 1890, Cratna
abdominalis Löbl, 1975, Pareuplectops coomani
Jeannel, 1957, and Megatyrus masumotoi
Nomura et al., 2010. Thirty-four species indicated by *-mark were newly added to the list of
pselaphine species recognized from Tanintharyi© 2021 National Museum of Nature and Science

Region in this research project.
Materials and Methods
In this time, all of the survey were conductedin the Tanintharyi Nature Reserve (Figs. 1, 2A,
2E). Most of the pselaphine specimens in thisstudy were collected by the following methods:
portable light trap in Nakase system (NLT: Figs.
2B–D), extraction by simpliﬁed Winkler apparatuses (SWA) from leaf litter and hand sifting ofleaf litter (SLL: Fig. 2F). The NLTs each with aﬂuorescent tube 4W in the system of Dr. Yuta
Nakase were used for collecting pselaphines by
Nomura (see Nomura, 2010, 2013). They were
ﬁxed or hooked on a tree and lighted in evening
and they were collected in the next morning.
After that, many pselaphine specimens were
picked up in the sorting of collected materials.
These NLTs were separately settled and collectedon the high position (ca. 4 m above the ground:HP) and the low position (ca. 1 m above the20

S. Nomura and M. M. Aung

Fig. 1. Map of collecting sites of Pselaphines in Tanintharyi Region.
Myanmar; B. Enlarged map of Tanintharyi Region.

ground: LP). Otherwise, some specimens wereextracted from sifted leaf litter by the simpliﬁed
Winkler apparatus (SWA). And the other somewere collected by hand sifting leaf litter (SLL).
Collected specimens are tentatively preserved in
the collection of the National Museum of Natureand Science, Tokyo, Japan (NMNS).
Results
1) A List of Pselaphine Specimens Collectedfrom Tanintharyi Region in Nov. 2018
All collecting data of the records shown below
are abbreviated as follows. See the foregoing partfor the collecting methods abbreviated as NLTHP/LP, SWA, and SLL.

A. Position of Tanintharyi Region in

NLT-HP/LP: Nakase system light trap athigh position /low position
12-13HP: Pt. D3, at high position (5 m above
the ground), by Nakase system light trap, Tanyntharyi Nature Reserve, Tanintharyi Region,
N14°44′15″, E98° 11′17″, ca. 30 m alt., 12–13. xi.
2018, S. Nomura leg.
12-13LP: same as above, but, at low position(1 m above the ground).
14-15HP: same as above, but, at high position(5 m above the ground), 14–15. xi. 2018.
14-15LP: same as above, but, at low position(1 m above the ground).
16-17HP: same as above, but, Pt. D4, at highposition (5 m above the ground), N14°44′22″,
E98°11′42″, alt. ca. 100 m, 16–17. xi. 2018.
16-17LP: same as above, but, at low positionPselaphine Species from Myanmar 3

21

Fig. 2. Habitats and collecting methods of Pselaphines in Tanintharyi Region. A. A landscape of Tanintharyi
Nature Reserve (TNR); B. A collecting site of TNR where a Nakase system light trap (NLT) was set at highposition (HP); C. A NLT set at low position (LP) above the ground; D. Sorting of the material collected by
NLT; E. The other collecting point in TNR; F. A snapshot of sifting leaf litter (SLL) by Nomura in TNR.

(1 m above the ground).
SWA: simpliﬁed Winkler apparatus
12-SWA: Tanintharyi Nature Reserve, by simpliﬁed Winkler apparatus, Tanintharyi Region,
N14°40′49″, E98°20′49″, ca. 500 m alt., 12. xi.
2018, S. Nomura leg.

14-SWA: same as above, but ca. 550 m alt.,
14. xi. 2018
16-SWA: Pt. D4, Tanintharyi Nature Reserve,by simpliﬁed Winkler apparatus, Tanintharyi
Region, N14°44′24″, E 98°11′43″, ca. 100 m alt.,
16. xi. 2018, S. Nomura leg.

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S. Nomura and M. M. Aung

Fig. 3. Pselaphine species recognized in Tanintharyi Region 1/2. A. Bythinoplectina gen. sp. 2; B. B. gen. sp. 3;
C. B. gen. sp. 4; D. B. gen. sp. 5; E. Bibloplectus? sp. 1; F. Pseudoplectus sp. 1; G. Philotrimium abdominale(Motschulsky, 1851); H. Anama sp. 2; I. Batricrator sp. 1; J. Smetanabatrus sp. 1; K. Tribasodites sp. 1; L. T.
sp. 2; M. Coryphomodes sp. 1; N. C. sp. 2; O. Batrisina, gen. sp. 1; P. Trichonomorphus ursinus Raﬀray, 1890;
Q. Intestinalius sp. 1; R. Oxyomela? sp. 1; S. Trisinus sp. 2; T. Cratna abdominalis Löbl, 1975.

SLL: sifting leaf litter
12SLL: Tanintharyi Nature Reserve, by sifting
leaf litter, Tanintharyi Region, N14°40′49″,
E98°20′49″, ca. 500 m alt., 12. xi. 2018, S.
Nomura leg.
13SLL: same as above, but ca. 550 m alt., 13. xi.
2018.
14SLL: same as above, but 14. xi. 2018.
15SLL: Pt. D4, Tanintharyi Nature Reserve, bysimpliﬁed Winkler apparatus, Tanintharyi
Region, N14°44′24″, E98°11′43″, ca. 100 m alt.,
15. xi. 2018, S. Nomura leg.
16SLL: same as above, but 16. xi. 2018.
In the following list, newly recognized species inthis cooperative project is indicated by *-mark.
Supertribe Euplectitae
Tribe Bythinoplectini
Subtribe Bythinoplectina
1. Bythinoplectina gen. sp. 1
Specimens examined. 1 ex., 15SLL.
2. Bythinoplectina gen. sp. 2* (Fig. 3A)Specimens examined. 1 ex., 12-13LP; 1 ex.,
15SLL.

3. Bythinoplectina gen. sp. 3* (Fig. 3B)
Specimens examined. 1 ex., 12-13HP.
4. Bythinoplectina gen. sp. 4* (Fig. 3C)Specimens examined. 1 ex., 12-13LP.
5. Bythinoplectina gen. sp. 5* (Fig. 3D)Specimens examined. 1 ex., 12-13HP.
Tribe Euplectini
6. Euplectus sp. 1
Specimens examined. 1 ex., 14-15HP; 1 ex.,
14-15LP.
Tribe Trichonychini
Subtribe Panaphatina
7. Bibloplectus? sp. 1* (Fig. 3E)
Specimens examined. 1 ex., 12-13LP.
Remarks. The genus Bibloplectus is diversiﬁed
in Europe and West Asia (35 spp. by Löbl andLöbl, 2015). A few species have been known
from the Oriental Region. It is characterized by
the small and elongate body, the fore tibiae andhind trochanters of the male without denticule,Pselaphine Species from Myanmar 3

and the abdominal tergites IV and V subequal inlength.
8. Pseudoplectus? sp. 1* (Fig. 3F)
Specimens examined. 1 ex., 14-15HP.
Remarks. The genus Pseudoplectus was originally deﬁned from Europe. It includes one named
species from Europe and one from Japan (Löbland Löbl, 2015). It is characterized by the verysmall (less than 1 mm) and elongate body and thealmost equal size of the abdominal tergites IV
and V.
Subtribe Trimiina
9. Philotrimium abdominale (Motschulsky, 1851)*(Fig. 3G)
Specimens examined. 1 ex., 12SLL; 1 ex.,
14SLL; 1 ex., 17.SLL.
Remarks. This species was originallydescribed from Tenasserim, S Myanmar by
Motschulsky (1851). It was also recorded from
Kaeng Krachan National Park, Thailand as“Aphilia sp. 3”, by Nomura et al. (2010).
Supertribe Batrisitae
Tribe Batrisini
Subtribe Batrisina
10. Anama sp. 1
Specimens examined. 1 ex., 12-13HP; 1 ex.,
14-15HP.
Remarks. This species was already recorded
from Tanintharyi Nature Reserve in the lastreport of this survey in Jan. 2017 (Nomura andAung, 2020b).
11. Anama sp. 2* (Fig. 3H)Specimens examined. 2 exs., 16-17LP.
Remarks. This species is very similar to thelast species, however it is separated by the large
body and the small eyes.
12. Mnia sp. 1
Specimens examined. 1 ex., 15 SLL; 1 ex.,
16SLL.

23

13. Batricrator sp. 1* (Fig. 3I)
Specimens examined. 1 ex., 12-13HP; 1 ex.,12-13LP; 4 exs., 14-15HP; 1 ex., 14-15LP.
Remarks. The genus Batricrator was deﬁned
by Jeannel (1957) with the type species, B. trabisoides described from Tonkin (＝ Ha Noi), N
Vietnam. The undescribed species of this genus
was collected from Tanintharyi Nature Reserve
for the ﬁrst time in the present study. The same
species was already reported from Kaeng Krachan National Park, Thailand by Nomura et al.
(2010) as “Tribasodites sp. 4”.
14. Smetanabatrus sp. 1* (Fig. 3J)Specimens examined. 1 ex., 15SLL; 2 exs.,
16SWA.
Remarks. The genus Smetanabatrus wasdeﬁned by Yin and Li (2013) with the type species, S. kinabalu from Borneo. The second species was described from Myanmar (Shan State)(Yin and Li, 2015), and the third was discoveredfrom the Malay Peninsula (Yin and Cuccodoro,2018). A species of this genus collected from
Tanintharyi Nature Reserve was concerned as anundescribed species.
15. Tribasodites sp. 1* (Fig. 3K)
Specimens examined. 1 ex., 13SLL; 2 exs.,
14SWA; 1 ex., 15SLL; 2 exs., 17SLL.
Remarks. Four species of the genera Tribasodites and Coryphomodes shown here andbelow were collected from the leaf litter accumurated on the ground in the forest of Tanintharyi
Nature Reserve, by sifting leaf litter (SLL) or by
extraction using simpliﬁed Winkler apparatus(SWA).
16. Tribasodites sp. 2* (Fig. 3 L)Specimens examined. 1 ex., 12SWA; 1 ex.,
13SLL.
17. Coryphomodes sp. 1* (Fig. 3M)Specimens examined. 5 exs., 12SLL; 16 exs.,
13SLL; 3 exs., 14SWA; 14 exs., 15SLL; 5 exs.,
16SLL.

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S. Nomura and M. M. Aung

18. Coryphomodes sp. 2* (Fig. 3N)Specimens examined. 9 exs., 12SLL; 7 exs.,
13SLL; 1 ex., 14SLL; 2 exs., 14SWA; 2 exs.,
15SLL.
19. Batrisina, gen. sp. 1* (Fig. 3O)Specimens examined. 1 ex., 16-17LP.
Remarks. This species couldnʼt be identiﬁed ingenus level because it lacks the male sexual character. It is similar to the Oxyomera? sp. shown
below, but it was clearly diﬀerent by the large
size of the body.
20. Trichonomorphus ursinus Raﬀray, 1890*(Fig. 3P)
Specimens examined. 1 ex., 15SLL; 1 ex.,
17SLL; 2 exs., 16-17LP.
Remarks. This species was originallydescribed from Penang Is., Malaysia. It is characterized by the middle-sized, broad and ﬂattenedbody, and the very coarsely punctate dorsal surface. It was recorded from Myanmar for the ﬁrst
time in the present study. A total of 4 specimensof this species were collected by sifting leaf litter(SLL) and portable light traps (NLT).
21. Intestinalius sp. 1* (Fig. 3Q)
Specimens examined. 3 exs., 16SLL.
Remarks. The genus Intestinalius was deﬁned
by Kurbatov (2007) for the type species, I. quinquesulcatus (Raﬀray, 1894) known from Singapore. It was ﬁrstly recorded from Myanmar in
the present study. Eight species of this genus has
been known from SE Asia. This species is very
similar to the type species, I. quinquesulcatus,
however it is separated by the unmodiﬁed andfusiform palpomere IV.
22. Oxyomera? sp. 1* (Fig. 3R)Specimens examined. 2 exs., 16-17LP
Remarks. The genus Oxyomera was deﬁned by
Raﬀray (1890) from Singapore with the type species, O. denticollis Raﬀray, 1890. This species issimilar to the type species in the laterally spinulate pronotum and the rounded abdomen. But it
was separated by the abdominal tergite IV with-out short median longitudinal carina.
23. Trisinus sp. 2* (Fig. 3S)Specimens examined. 1 ex., 12-13HP; 1 ex.,12-13LP.
Remarks. This species is similar to the Japanese species, T. torticornis known from the
Ryukyus in the modiﬁed antennomere 6–11 in
the male. However, it is separable by the subglobose male antennomere 7 clearly larger than 8(not subglobose antennomere 7 smaller than 8 in
T. torticornis).
24. Cratna abdominalis Löbl, 1975* (Fig. 3T)Specimens examined. 1 ex., 16-17HP.
Remarks. This species was originallydescribed from Taiwan. It was recorded also
from Vietnam by Nomura and Pham (2019). It ischaracterized by the long and slender legs, maxillary palpi, and the dorsally ﬂattened abdominal
tergite IV without sexual patch in the male.
25. Cratna venusta Blattný, 1925
Specimens examined. 1 ex., 12SLL.
Remarks. This species was originallydescribed from Tenasserim, S Myanmar, by
Blattný (1925). It was also recognized in the last
survey in Jan. 2017. Its male character is
unknown because it was described on the basisof one female.
26. Batrisiella tenasserimi (Blattný, 1925)
Specimens examined. 4 exs., 12-13HP; 1 ex.,12-13LP; 6 exs., 14-15HP.
Remarks. This species was described as a new
species of the genus Bartisocenus Raﬀray, 1903.
It was reclassiﬁed to the genus Batrisiella Raffray, 1904 in Nomura and Aung (2020b).
27. Trisiniotus nitidulus (Motschulsky, 1851)
Specimens examined. 9 exs., 12-13HP; 5 exs.,
12-13LP; 1 ex., 14-15HP.
Remarks. This is one of the most common species collected by light traps in Tanintharyi Nature
Reserve. It was originally described as a memberof the genus Batrisus Aubé, 1833, however itPselaphine Species from Myanmar 3

25

was moved to the genus Trisiniotus Jeannel,
1960 by Nomura and Aung (2020b).

Supertribe Goniaceritae
Tribe Arnyllini

28. Arthromelodes reichenbachi (Motschulsky,
1851)
Specimens examined. 1 ex., 14-15HP.
Remarks. This species was originallydescribed as a new species of the genus Batrisus.
It was reclassiﬁed to the genus Arthromelodes
Jeannel, 1954 by Nomura et Aung (2020b). It ischaracterized by the weakly bent antennae andthe abdominal tergite IV with a large sexual
patch on dorsal side in the male.

30. Harmophorus gibbioides Motschulsky, 1951
Specimens examined. 1 ex., 14-15HP; 1 ex.,
16SWA; 1 ex., 16-17LP.

29. Arthromelodes sp. 1* (Fig. 4A)Specimens examined. 1 ex., 12-13HP.
Remarks. This species is similar to the former
species in having the large sexual patch on theabdominal segment IV in the male. However, it
is separable by the short and straight antenna.

32. Atenisodus sp. 1
Specimens examined. 1 ex., 15SLL; 4 exs.,
16SLL; 9 exs., 16SWA.

Tribe Brachyglutini
Subtribe Brachyglutina
31. Reichenbachia sp. 3
Specimens examined. 6 exs., 12-13HP; 1 ex.,
14-15HP; 1 ex.,16-17HP; 1 ex., 16-17LP.

33. Batraxis sp. 1* (Fig. 4B)
Specimens examined. 37 exs., 12SLL; 26 exs.,
12SWA; 16exs., 13SLL; 9 exs., 14SLL; 17 exs.,
14SWA; 88 exs., 15 SLL; 10 exs., 16SLL; 3 exs.,

Fig. 4. Pselaphine species recognized in Tanintharyi Region 2/2. A. Arthromelodes sp. 1; B. Batraxis sp. 1; C.
Sunorfa exsculpta (Schaufuss, 1887); D. Morana sp. 1; E. Maya dilatata (Motschulsky, 1851); F. Pareuplectops coomani Jeannel, 1957; G. P. tenasserimi (Blattný, 1925); H. Atychodea quadrifoveolata (Motschulsky,1851); I. Plagiophorus paradoxus (Motschulsky, 1851); J. P. sp. 4; K. P. sp. 5; L. Stipesa sp. 1; M. Ancystrocerus sp. 1; N. Megatyrus masumotoi Nomura et al., 2011.

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S. Nomura and M. M. Aung

16SWA.
Remarks. In the genus Batraxis, B. raﬀrayana
has been known from Tenasserim and Yangon(Nomura and Aung, 2020a). This species isclearly diﬀerent from B. raﬀrayana by having
the long and slender antennae and the peanutshaped antennomere X.
Tribe Iniocyphini
Subtribe Natypleurina
34. Sunorfa exsculpta (Schaufuss, 1887)* (Fig.
4C)
Specimens examined. 1 ex., 16-17LP.
Remarks. This species was originallydescribed by Schaufuss (1887) from Sumatra,
however it was recorded also from Tenasserim
by Blattný (1925).
35. Morana sp. 1* (Fig. 4D)Specimens examined. 1 ex., 16SWA.
Remarks. The genus Morana is widely distributed in East to Southeast Asia. Two species havebeen already known from Myanmar according to
Nomura and Aung (2020a).
36. Maya dilatata (Motschulsky, 1851)* (Fig.
4E)
Specimens examined. 3 exs., 16SWA.
Remarks. The genus Maya was established by
Blattný (1925) based on the type species M. dilatata discovered from Tenasserim, S Myanmar
and M. uzeli distributed in Ceylon (Sri Lanka).
This species is characterized by the small-sized(ca. 1.2 mm) and blackish body.
Tribe Proterini
37. Pareuplectops coomani Jeannel, 1957* (Fig.
4F)
Specimens examined. 10 exs., 12-13HP; 8
exs., 12-13LP; 4 exs., 14-15HP; 4 exs., 14-15LP;
1 ex., 16-17 L.
Remarks. This species is the type species ofthis genus, which was described by Jeannel(1957) from Tonkin (Ha Noi), N Vietnam. It is

recorded from Myanmar for the ﬁrst time in thepresent study.
38. Pareuplectops tenasserimi (Blattný, 1925)*(Fig. 4G)
Specimens examined. 18 exs., 12-13HP; 20
exs., 12-13LP; 15 exs., 14-15HP; 15 exs.,
14-15LP.
Remarks. This is the ﬁrst species of this genus
from Myanmar. It was originally described from
Tenasserim as a new species of the genus Epiplectus. Kurbatov and Cuccodoro (2009) moved
this species to Pareuplectops. In the same paper,
they synomymized E. novissimus Blattný, 1925
with this species.
Tribe Tychini
39. Atychodea quadrifoveolata (Motschulsky,1851)* (Fig. 4H)Specimens examined. 1 ex., 12-13HP; 1 ex.,
14-15HP.
Remarks. According to Nomura and Aung(2020a), this species was originally described asa species of Tychus, however it was moved to
Atychodea by Kurbatov and Sabella (2008).
Tribe Cyathigerini
40. Plagiophorus paradoxus Motschulsky, 1851*(Fig. 4I)
Specimens examined. 1 ex., 12-13HP; 2 exs.,
14-15HP; 2 exs., 16SWA; 1 ex., 16-17LP.
Remarks. Burckhardt and Löbl (2002) revalidated the genus Plagiophorus deﬁned by
Motschulsky (1851), and synonymized the following four genera with Plagiophorus:
Cyathiger King, 1865, Cyathigerodes Jeannel,
1951, Denicyathiger Jeannel, 1951, Paracyathiger Jeannel, 1951, Manuleiger Jeannel, 1961.
They also added a checklist of the species of thisgenus including 84 valid species. This species isvery important as it is the type species of thelarge genus Plagiophorus. It was redescribed indetail by Burckhardt and Löbl (2002) on thebasis of the holotype preserved in the ZoologicalPselaphine Species from Myanmar 3

Museum of the University, Moskow, Russia.
41. Plagiophorus sp. 4* (Fig. 4J)Specimens examined. 2 exs., 16SWA.
42. Plagiophorus sp. 5* (Fig. 4K)
Specimens examined. 1 ex., 16SWA.
Supertribe Pselaphitae
Tribe Ctenistini
43. Ctenisoschema coomani (Jeannel, 1957)
Specimens examined. 1 ex., 16-17HP.
Remarks. This species originally describedfrom Tonkin (Ha Noi), N Vietnam was ﬁrstly
recorded from Myanmar by Nomura and Aung(2020b).
44. Ctenistes sp. 1
Specimens examined. 1 ex., 16-17HP.
Tribe Hybocephalini
45. Stipesa sp. 1* (Fig. 4 L)
Specimens examined. 1 ex., 12-13HP; 2 exs.,
14-15.LP.
Remarks. Blattný (1925) described three species of the genus Filiger Schaufuss, 1877, which
was synonymized with Stipesa Sharp, 1874.
Then, three Filiger species were moved to
Stipesa. These three species are uniformly covered with coarse punctures and short hairs. However, the other species partly covered with long
hairs and scales were discovered from Tanintharyi Nature Reserve and recorded herein.
Tribe Tmesiphorini
46. Pseudophanias sp. 1
Specimens examined. 2 exs., 16SLL; 1 ex.,
17SLL.
Remarks. This species and the next species
were recorded from Tanintharyi Nature Reserveby Nomura and Aung, 2020b for the ﬁrst time.
Both species were collected also in 2018.

27

47. Pseudophanias sp. 2
Specimens examined. 6 exs., 15SLL; 9 exs.,
16SLL; 6 exs., 17SLL.
48. Ancystrocerus sp. 1* (Fig. 4M)
Specimens examined. 1 ex., 13SLL.
Remarks. The genus Ancystrocerus Raﬀray,1893 known from tropical Asia was classiﬁed
into the tribe Tyrini in Newton and Chandler(1989). However, it should be included in thetribe Tmesiphorini, because it has c-shaped
setose sulci just beneath antennal bases.
Tribe Tyrini
Subtribe Tyrina
49. Megatyrus masumotoi Nomura et al., 2011*(Fig. 4N)
Specimens examined. 1 ex., 13SLL; 1 ex.,
14SLL; 1 ex., 16SLL.
Remarks. The genus Megatyrus was established by Hlaváč and Nomura (2003) on the basisof three species collected from China and Vietnam. After that, Nomura et al. (2011) added three
new species from Thailand. At that time, M.
masumotoi was described from Kaeng Krachan
National Park. In the present study, this species
was discovered from Tanintharyi Nature
Reserve. It is the ﬁrst record of this genus andthis species from Myanmar.
Subtribe Centrophthalmina
50. Centrophthalmus helferi Blattný, 1925
Specimens examined. 1 ex., 14-15HP.
2) The table of pselaphine specimens collectedby NLTs in the survey in Nov. 2018
The weather conditions, habitats, trapped position, and the collected pselaphine specimens byportable light traps (NLT) are tabulated in Table
1. As shown in Table 1, 173 pselaphine specimens of 29 species were collected by NLTs in
total.

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S. Nomura and M. M. Aung
Table 1.
Date
Locality

Collected traps (set traps)
Habitat
Position
Bythinoplectina gen. sp. 2*
B. gen. sp. 3*
B. gen. sp. 4*
B. gen. sp. 5*
Euplectus sp. 1
Bibloplectus? sp. 1*
Pseudoplectus? sp. 1*
Anama sp. 1*
A. sp. 2*
Batriclator sp. 1*
Batrisina, gen. sp. 1
Trichonomorphus ursinus*
Oxyomera? sp. 1*
Trisinus sp. 2
Cratna abdominalis
Arthromelodes reichenbachi
A. sp. 1
Batrisiella tenasserimi
Trisiniotus nitidulus
Pareuplectops tenasserimi
P. coomani
Harmophorus gibbioides
Reichenbachia sp. 3
Sunorfa exsculpta*
Plagiophorus paradoxus*
Atychodea quadrifoveolata
Stipesa? sp. 1*
Ctenisoschema coomani
Centrophthalmus helferi
No. of species
No. of specimens

Condition and result of light trap survey in Myanmar in Nov. 2018.
12-13.xi.2018
Tanintharyi Nature
Reserve
D3, alt. 60 m

14-15.xi.2018
Tanintharyi Nature
Reserve
D3, alt. 60 m

16-17.xi.2018
Tanintharyi Nature
Reserve
D4, alt. 100 m

4 (4)forest5m

4 (4)forest5m

4 (4)forest
5m

1
1

4 (4)
forest
1m

1

1
1

1

1

1

1

1

1
1
4

6

6
1
15
4
1
1

1
1
1

2
1
2

15
4
1

1
14
56

9
39

Acknowledgements
We wish to express our hearty thanks to Dr.
Zi-Wei Yin of the Shanghai Normal University,
China for his critical reading of the manuscript.
This study was carried out by the integrated
research project “Biological Inventory with special attention to Myanmar” initiated by the
National Museum of Nature and Science, Japan
based on MoU between NMNS and Forest
Department, Ministry of Natural Resources andEnvironmental Conservation, Myanmar. We are
grateful to the Myanmar Forest Department,
Ministry of Natural Resources and Environmen-

14
41

1
2
1
1

1
1
5
20
8

2

1

5
22

1
1
1
1
1

2
3
4

Total

4 (4)
forest
1m

1

1

1
4
9
18
10

4 (4)
forest
1m

9
11

1
1
1
1
2
1
1
2
2
7
1
2
1
2
1
1
1
11
15
68
27
2
9
1
4
2
3
2
1
29
173

tal Conservation for their permission to carry out
ﬁeldwork in protected forests, and for their support and collaboration.
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29

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