柴田 泰利・丸山 宗利・保科 英人・岸本 年郎・直海 俊一郎・
野村 周平・Volker Puthz・島田 孝・渡辺 泰明・山本 周平

日本産ハネカクシ科総目録（昆虫綱：甲虫目）

Reprinted from Bulletin of the Kyushu University Museum No. 11
pp. 69-218, March 2013

九州大学総合研究博物館研究報告

Bull. Kyushu Univ. Museum
No. 11, 69-218, 2013

日本産ハネカクシ科総目録（昆虫綱：甲虫目）
柴田泰利 1, 11・丸山宗利 2, 11・保科英人 3・岸本年郎 4・直海俊一郎 5・野村周平 6・
Volker Puthz7・島田 孝 8・渡辺泰明 9・山本周平 101

東京都町田市・2九州大学総合研究博物館・3福井大学教育学部・
4
自然環境研究センター・5千葉県千葉市・6国立科学博物館・
7
Burgmuseum Schlitz・8 ㈱静環検査センター・9 東京農業大学・
10
九州大学大学院生物資源環境科学府・11
編集担当（編集より後の著者はラテンアルファベット順）

Catalogue of Japanese Staphylinidae（Insecta: Coleoptera）
Yasutoshi Shibata1, 11, Munetoshi Maruyama2, 11, Hideto Hoshina3, Toshio Kishimoto4,
Shun-Ichiro Naomi5, Shûhei Nomura6, Volker Puthz7, Takashi Shimada8,
Yasuaki Watanabe9, Shûhei Yamamoto10

1

Machida-shi, Tokyo, 2The Kyushu University Museum,3
Faculty of Education, Fukui University, 4 Japan Wildlife Research Center,
5
Chiba-shi, Chiba, 6National Museum of Nature and Science, Tokyo,
7
Burgmuseum Schlitz, 8Seikan Kensa Center Inc., 9Tokyo University of Agriculture,
10
Graduate School of Bioresource and Bioenvironmental Sciences, Kyushu University, 11Editors

要旨．甲虫最大の一群，ハネカクシ科（398属2,262種44亜種）
を対象とし，2012年上半期までに記載された日本産種を目録化した.
Abstract. A catalogue of the Japanese species of Staphylinidae (398 genera, 2,262 species, 44 subspecies), which is oneof the largest families of Coleoptera, is presented.

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日本産ハネカクシ科総目録（昆虫綱：甲虫目）
Catalogue of Japanese Staphylinidae (Insecta: Coleoptera)

はじめに Introduction
ハネカクシ科甲虫は世界に ５ 万 ８ 千種あまりが知られていて， 37万種といわれている甲虫目の中でも大きな科の一つである．その生活範囲は広く，地球上の多様な環境に適応して生息している．
こ の 科 の 世 界 の 種 類 を ま と め た 目 録 に はJunk & Schenkling（ 1910–1940）の「 Coleopterorum
Catalogus 」が あ り， さ ら に 日 本 を 含 む 旧 北 区 の も の と し て はWinkler（ 1924–1932）の「 Catalogus
Coleopterorum Regionis Palaearcticae 」がある．その後は長いあいだ， 目録が出ることはなかったが，2001年にHerman による「 Catalog of the Staphylinidae I-VII 」が出版された．ただこのシリーズにはデオキノコムシ亜科， アリヅカムシ亜科， アリガタハネカクシ亜科とヒゲブトハネカクシ亜科は入っていない．2003年になり，Löbl & Smetana による「 Catalogue of Palaearctic Coleoptera 」の １ 巻が発刊され，
現在 ７ 巻までが出版されている．２ 巻（2004）がハネカクシ科を含み，デオキノコムシとアリヅカムシがそれぞれ亜科としてハネカクシ科の一員として取り扱われている．この二大冊は情報量も多く， ハネカ
クシの分類には非常に大切な文献である．
いっぽう， 日本のハネカクシ科に関連のある目録としては， 安立綱光（1957）の「 The Staphylinid
Fauna of Japan 」があり720種が収載されていて， 当時は日本のハネカクシ相の基礎資料として広く利用された．つぎに， 柴田泰利による日本産ハネカクシ科目録（I）（V）– （ 1976–1985）があり， 当時のハネカク
シ科からヒゲブトハネカクシ亜科を除き679種を取り上げている．その後は九州大学農学部昆虫学教室と日本野生生物研究センター（1989）による「日本産昆虫総目録 I–II+ 索引」，同（1990）によるその追加・訂正が出たが， ハネカクシ科はⅠ巻と追加・訂正にあり1,012種が記録されている．デオキノコムシとアリ
ヅカムシは独立の科としている．
ここ4， 50年間の日本や外国のハネカクシ専門家による研究の進展に伴い日本産ハネカクシ科の種数は急速に増加し， 2009年に新たにこの科に加わったコケムシを合わせて現在2,260種にもなっている．1957
年の安立の日本産ハネカクシの目録に収載された種数720種に比べて約 ３ 倍の増加である．そこで今回，国内のハネカクシの専門家と共同して新たな知見を含む日本産ハネカクシ科の目録を編纂することにした．
この目録が多くの方々に広く利用されることを期待し， 補遺の出版に向け， 内容のさらなる充実に心がけるつもりである．
About 58,000 species of rove beetles have been described globally. The family is currently ranked as the largest
within Coleoptera, an order consisting of 370,000 species. As a group, staphylinids display a wide range of lifestyles
and have colonised diverse terrestrial environments.
Junk & Schenkling (1910–1940) published a catalogue of the family for the world species in their “Coleopterorum
Catalogus”; subsequently, Winkler (1924–1932) made a catalogue for the Palearctic species: “Catalogus Coleopterorum
Regionis Palaearcticae.” However, since then, no catalogue was published for several decades. In 2001, Herman
published a series of volumes in his “Catalog of the Staphylinidae I–VII.” However, this series did not include thesubfamilies Scaphidiinae, Pselaphinae, Paederinae and Aleocharinae. From 2003, Löbl & Smetana began publishingthe “Catalogue of Palaearctic Coleoptera”, now at Volume 7. Volume 2 (2004) contains Staphylinidae (including
Scaphidiinae and Pselaphinae). Together, the Herman and Löbl & Smetana catalogues are currently amongst the mostimportant pieces of literature for staphylinid taxonomy.
For Japanese staphylinids, Adachi (1957) published “The Staphylinid Fauna of Japan” which includes 720 species.

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Yasutoshi Shibata, Munetoshi Maruyama, Hideto Hoshina, Toshio Kishimoto, Shun-Ichiro Naomi, Shûhei Nomura, Volker Puthz, Takashi Shimada, Yasuaki Watanabe, Shûhei Yamamoto

It was widely used as a basic information source for the Japanese species. Later, Shibata (1976–1985) published hisseries “Catalogues of Japanese Staphylinidae (I)–(V)” which treated the 679 species known at that time excluding
Aleocharinae. After that, the Entomological Laboratory (Kyushu University) and Japan Wildlife Research Center(1989, 1990) published “A Check List of Japanese Insects”, which listed 1,012 species, but treated Scaphidiinae andPselapininae as distinct families.
For the past 40–50 years, the number of staphylinid species has been rapidly increasing thanks to studies by Japaneseand foreign researchers. Presently, about 2,260 species, including a recently added subfamily, Scydmaeninae, are
known from Japan. Hence, the known Japanese staphylinid diversity has approximately tripled since Adachi (1957). In
this article, we present a comprehensive catalogue of Japanese Staphylinidae, which incorporates many new species andupdated nomenclature.
We hope this catalogue will be utilized by many people, and intend to augment the contents in future supplements.
We thank Dr. Joseph Parker for reviewing the English text.

凡例 Explanatory note⑴

目録はLöbl & Smetana（ 2004 ）の目録を参考にして作成したものであるが， 一部追加や変更， また新たな体系の提案などを含む．

⑵

2012年前半までに発表された日本産（千島列島の歯舞諸島，色丹島，国後島，択捉島を含む）ハネカクシ科を対象としている．

⑶

各亜科の著者名を表1に示し，地名の略号を表 ２ に一覧した．

⑷

すべての属， 種に和名をつけたが， 新たにつけた和名には新称， 変更したときは改称を括弧内に表した．
「 HN 」は異物同名，「 RN 」は置換名を示す．

⑴

This catalogue is based on that of Löbl & Smetana (2004), but includes many new additions and changes.

⑵This catalogue includes those species published up to the first half of 2012.

⑶

The author(s) of each subfamily is shown in Table 1, and abbreviations of local names are listed in Table 2.

⑷

We provide Japanese names for all genera and species and indicate new names and new name changes using 新
称 [new name] and 改称 [rename], respectively. “HN” denotes a homonym, and “RN” a replacement name.

表 １．日本産ハネカクシ科の亜科と担当著者
Table 1. Subfamilies of Japanese Staphylinidae and authors.
Omaliinae ヨツメハネカクシ亜科

渡辺泰明（ Y. Watanabe ）Proteininae ハバビロハネカクシ亜科

柴田泰利（ Y. Shibata ）Micropeplinae チビハネカクシ亜科

柴田泰利（ Y. Shibata ）Dacycerinae ニセマキムシ亜科

柴田泰利（ Y. Shibata ）Pselaphinae アリヅカムシ亜科

野村周平（ S. Nomura ）Tachyporinae シリホソハネカクシ亜科

山本周平（ S. Yamamoto ）Trichophyinae ホソヒゲハネカクシ亜科

柴田泰利（ Y. Shibata ）Aleocharinae ヒゲブトハネカクシ亜科

丸山宗利（ M. Maruyama ）・岸本年郎（ T. Kishimoto ）2013 The Kyushu

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日本産ハネカクシ科総目録（昆虫綱：甲虫目）
Catalogue of Japanese Staphylinidae (Insecta: Coleoptera)

Apateticinae オサシデムシモドキ亜科

柴田泰利（ Y. Shibata ）Scaphidiinae デオキノコムシ亜科

保科英人（ H. Hoshina ）Piestinae ヒラタハネカクシ亜科

直海俊一郎（ S. Naomi ）Osoriinae ツツハネカクシ亜科

直海俊一郎（ S. Naomi ）Oxytelinae セスジハネカクシ亜科

島田

Oxyporinae オオキバハネカクシ亜科

柴田泰利（ Y. Shibata ）Megalopsidiinae メダカオオキバハネカクシ亜科

直海俊一郎（ S. Naomi ）Steninae メダカハネカクシ亜科

直海俊一郎（ S. Naomi ）・Volker Puthz

Euaesthetinae チビフトハネカクシ亜科

柴田泰利（ Y. Shibata ）Scydmaeninae コケムシ亜科

保科英人（ H. Hoshina ）Pseudopsinae スジヒラタハネカクシ亜科

柴田泰利（ Y. Shibata ）Paederinae アリガタハネカクシ亜科

渡辺泰明（ Y. Watanabe ）Staphylininae ハネカクシ亜科

柴田泰利（ Y. Shibata ）孝（ T. Shimada ）表 ２．地名略号一覧（アルファベット順は別表1を参照）．
Table 2. Abbreviations of local names (See, Appendix 1 for alphabetical list).
略号 Abbreviations
日本本土 Mainland Japan

略号 Abbreviations

MJ

伊豆諸島 Izu-shotô（Islands）

IzIss

北海道 Hokkaidô

HOK

大島（伊豆）Ôshima（Izu）

Iz-O

本州 Honshû

HON

新島 Nii-jima

Iz-N

四国 Shikoku

SHI

神津島 Kôzu-shima

Iz-Kz

九州 Kyûshû

KYU

三宅島 Miyake-jimaIz-My

御蔵島 Mikura-jima

Iz-Mk

八丈島 Hachijô-jima

Is-Hch

青ヶ島 Ao-ga-shima

Iz-A

北海道沖島嶼 Islands off Hokkaidô（千島 Chishima（=Kuril）

Chs ）国後島 Kunashiri-tô（=Kunashir）

Kun

択捉島 Etorofu-tô（=Iturup）

Etr

父島 Chichi-jima

Og-cc

利尻島 Rishiri-tô

Rsr

母島 Haha-jima

Og-hh

礼文島 Rebun-tôReb

弟島 Otôto-jima

Og-ot

大島（渡島）Ôshima（Oshima）

OOs

北硫黄島 Kita-iô-jimaKIw

奥尻島 Okushiri-tôOks

硫黄島 Iô-jima

Iw

南硫黄島 Minami-iô-jima

MIw

小笠原諸島 Ogasawara-shotôOg

本州沖島嶼 Islands off Honshû
飛島 Tobi-shimaTob

粟島 Awa-shima

Aws

小豆島 Shôdo-shima

Shd

佐渡島 Sado-ga-shimaSad

沖ノ島（四国）Oki-no-shima（SHI）

Okns

淡路島 Awaji-shima

Awj

大島（紀伊）Ôshima（Kii）

KOs

隠岐 OkiOki2013 The Kyushu

四国沖島嶼 Islands off Shikoku

九州沖島嶼 Islands off Kyûshû
沖ノ島（筑前）Oki-no-shima（Chikuzen） Kokn

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Yasutoshi Shibata, Munetoshi Maruyama, Hideto Hoshina, Toshio Kishimoto, Shun-Ichiro Naomi, Shûhei Nomura, Volker Puthz, Takashi Shimada, Yasuaki Watanabe, Shûhei Yamamoto

略号 Abbreviations

略号 Abbreviations
戸島 He-jima

Hej

横当島（吐噶喇）Yokoate-jima（Tokara）T-Yok

樺島 Kaba-shima

Kbs

宝島（吐噶喇）Takara-jima（Tokara）

T-Tak

平戸 Hirado

Hrd

与論島 Yoron-tô

Yor

対馬 TsushimaTsm

喜界島 Kikai-jimaKik

壱岐 IkiIki

奄美大島 Amami-ôshima

Amm

男女群島 Danjo-guntôDnj

徳之島 Toku-no-shima

Tok

五島列島 Gotô-rettô

Got

沖永良部島 Okinoerabu-jimaOke

中通島 Nakadôri-jima

Nkd

沖縄島（本島）Okinawa-jima

Okn

福江島 Fukue-jima

Fuk伊是名島 Izena-jimaIzn

天草 Amakusa

Amk

伊平屋島 Iheya-jima

Ihy

上甑島 Kami-koshiki-jima

KKs

久米島 Kume-jimaKum

下甑島 Shimo-koshiki-jima

SKs

北大東島 Kita-daitô-jima

KDt

南大東島 Minami-daitô-jima

MDt

RK

宮古島 Miyako-jima

Myk

屋久島 Yaku-shimaYak伊良部島 Irabu-jima

Irb

種子島 Tane-ga-shimaTan（八重山諸島 Yaeyama-shotôYae）

薩摩黒島 Kuro-shima

Krs

多良間島 Tarama-jimaTrm

口永良部島 Kuchinoerabu-jima

Kue

石垣島 Ishigaki-jimaIsh（トカラ列島 Tokara-rettôT-R）

西表島 Iriomote-jima

Iri

中之島（吐噶喇）Nakano-shima（Tokara）T-Nak

波照間島 Hateruma-jimaHat

悪石島（吐噶喇）Akuseki-jima（Tokara）T-Aks

与那国島 Yonaguni-jima

Yon

南西諸島 Nansei-shotô（Ryukyu）

諏訪瀬島（吐噶喇）Suwanose-jima（Tokara）T-Sw

総目録 Catalogue

Family Staphylinidae Latreille, 1802 ハネカクシ科
Subfamily Omaliinae MacLeay, 1825 ヨツメハネカクシ亜科（28属176種）
Tribe Anthophagini Thomson, 1859
Genus Acidota Stephens, 1829 ナガヨツメハネカクシ属
crenata japonica Y. Watanabe, 1990c: 145 ナガヨツメハネカクシ HOK, HON
daisetsuzana Y. Watanabe, 1990c: 146 ダイセツナガヨツメハネカクシ（和名新称）HOK
quadrata Zetterstedt, 1838: 53 (Omalium) カクナガヨツメハネカクシ（和名新称）HOK; Russia, Mongolia, Europe
Genus Anthobium Leach, 1819 ハバビロヨツメハネカクシ属
subgenus Anthobium Leach, 1819

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Catalogue of Japanese Staphylinidae (Insecta: Coleoptera)hammondi Y. Watanabe, 1989: 53 (Lathrimaeum) ハモンドハバビロヨツメハネカクシ（和名新称）Japan
Genus Brathinus LeConte, 1852 ヒョウタンヨツメハネカクシ属
oculatus Lewis, 1886: 85 ヒョウタンヨツメハネカクシ（和名改称）HOK, HON= japonicus Nakane, 1961: 3
shikokuensis Y. Watanabe & Satô, 1981: 615 シコクヒョウタンヨツメハネカクシ（和名改称）SHI
Genus Camioleum Lewis, 1893 ヨツメツヤシデムシモドキ属
loripes Lewis, 1893b: 395 ヨツメツヤシデムシモドキ（和名改称）HON
Genus Geodromicus Redtenbacher, 1857 ミズギワヨツメハネカクシ属
amamianus Y. Watanabe, 1990c: 284 (Psephidonus) アマミミズギワヨツメハネカクシ（和名新称）Amm
aokii Y. Watanabe, 1972: 111 (Psephidonus) アオキミズギワヨツメハネカクシ HOK
caliginosus Sharp, 1889: 470 (Anthophagus) クロミズギワヨツメハネカクシ HOK, HON, SHI; South Korea, North
Korea= lewisi Cameron, 1930: 183
hammondi Y. Watanabe, 1990c: 269 (Psephidonus) ハモンドミズギワヨツメハネカクシ（和名新称）HON, KYU
hermani Y. Watanabe, 1991b: 43 [RN] (Psephidonus) ハーマンミズギワヨツメハネカクシ（和名新称）HOK, HON,
SHI, KYU, Iz-Mk; South Korea= pusillus Y. Watanabe, 1990c: 279 [HN] (Psephidonus)
japonicus Y. Watanabe, 1990c: 280 (Psephidonus) ヤマトミズギワヨツメハネカクシ（和名新称）HOK, HON
kirschenblati Tikhomirova, 1973: 155 キタミズギワヨツメハネカクシ（和名新称）HOK; Russia (Far East)= iburinus Y. Watanabe, 1990c: 278 (Psephidonus)
lestevoides Sharp, 1889: 470 (Anthophagus) ミズギワヨツメハネカクシ HOK, HON, SHI; South Korea, Russia (Far East)
miyamorii Y. Watanabe, 1990c: 287 (Psephidonus) ミヤモリミズギワヨツメハネカクシ（和名新称）HOK
sibiricus Bernhauer, 1915a: 69 ミズギワヨツメハネカクシ（和名新称）HON; Korea, China, Russia (Sakhalin,Siberia)= niponensis Y. Watanabe, 1990c: 275 (Psephidonus)
sinuatus Sharp, 1889: 470 (Anthophagus) クロツヤミズギワヨツメハネカクシ HOK, HON; China
sprius Y. Watanabe, 1990c: 272 (Psephidonus) ホソミズギワヨツメハネカクシ（和名新称）HON
suensoni Bernhauer, 1936: 304 ミヤマミズギワヨツメハネカクシ HON
Genus Lesteva Latreille, 1797 ヒメミズギワヨツメハネカクシ属（和名新称）
crassipes Sharp, 1889: 471 ムモンヒメミズギワヨツメハネカクシ（和名改称）HON, SHI, KYU
distincta Y. Watanabe, 1990c: 178 カスガヒメミズギワヨツメハネカクシ（和名新称）HON, SHI, KYU; Korea
fenestrata Sharp, 1874a: 97 フタモンヒメミズギワヨツメハネカクシ（和名改称）HON, SHI, KYU
gracilis Y. Watanabe, 1990c: 157 ホソヒメミズギワヨツメハネカクシ（和名新称）HON
hammondi Y. Watanabe, 1990c: 166 ハモンドヒメミズギワヨツメハネカクシ（和名新称）HON
japonica Y. Watanabe, 1990c: 172 ウスグロヒメミズギワヨツメハネカクシ（和名新称）HON, SHI
kojohamana Y. Watanabe, 1990c: 157 エゾヒメミズギワヨツメハネカクシ（和名新称）HOK
lewisi Cameron, 1930: 182 ルイスヒメミズギワヨツメハネカクシ（和名新称）HON, SHI, KYU, Shd

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Yasutoshi Shibata, Munetoshi Maruyama, Hideto Hoshina, Toshio Kishimoto, Shun-Ichiro Naomi, Shûhei Nomura, Volker Puthz, Takashi Shimada, Yasuaki Watanabe, Shûhei Yamamoto

minima Y. Watanabe, 1990c: 153 ヒメミズギワヨツメハネカクシ（和名新称）HON
miyabi Y. Watanabe, 1990c: 175 キタヒメミズギワヨツメハネカクシ（和名新称）HON; South Korea, North Korea
miyamorii Y. Watanabe, 1990c: 174 ミヤモリヒメミズギワヨツメハネカクシ（和名新称）HOK, HON
nipponica Y. Watanabe, 1990c: 164 ニホンヒメミズギワヨツメハネカクシ（和名新称）HON
okiana Y. Watanabe, 2004a: 74 オキフタモンヒメミズギワヨツメハネカクシ（和名改称）Okipersimilis Cameron, 1930: 182 ニセフタモンヒメミズギワヨツメハネカクシ（和名新称）HON, SHI
plagiata Sharp, 1889: 472 ネアカヒメミズギワヨツメハネカクシ
HON, SHI, KYU（和名改称）
shibatai Y. Watanabe, 1990c: 158 シバタヒメミズギワヨツメハネカクシ（和名新称）HON
shimadai Y. Watanabe, 2004a: 71 シマダヒメミズギワヨツメハネカクシ（和名改称）Oki
shikokuana Y. Watanabe, 1990c: 160 シコクヒメミズギワヨツメハネカクシ（和名新称）SHI
tsushimae Y. Watanabe, 1990c: 170 ツシマヒメミズギワヨツメハネカクシ（和名新称）Tsm
Genus Liophilydrodes Nakane & Sawada, 1983 ヒラタヨツメハネカクシ属（和名新称）
flavipennis Y. Watanabe, 1990c: 303 キバネヒラタヨツメハネカクシ（和名新称）HON
pullus Nakane & Sawada, 1956b: 184 (Philydrodes) クロヒラタヨツメハネカクシ（和名改称）HOK, HON
puncticollis Nakane & Sawada, 1956b: 183 (Philydrodes) エゾヒラタヨツメハネカクシ（和名改称）HOK; Russia(Sakhalin)
subtilis Sharp, 1889: 471 (Anthophagus) アカヒラタヨツメハネカクシ（和名改称）HON
suzukii Y. Watanabe, 1990c: 305 ユミオリヒラタヨツメハネカクシ（和名新称）HON
troglophilus Nakane & Sawada, 1956b: 184 (Philydrodes) アナヒラタヨツメハネカクシ（和名新称）HON
yamanakai Y. Watanabe, 1990c: 301 タテヤマヒラタヨツメハネカクシ（和名新称）HON
Genus Mannerheimia Mäklin, 1880 セマルヨツメハネカクシ属（和名新称）
curtella Sharp, 1889: 473 (Omalium) セマルヨツメハネカクシ（和名新称）HON, SHI
evias Y. Watanabe, 1990c: 84 ミヤマセマルヨツメハネカクシ（和名新称）HON
japonica Watanabe, 1990c: 87 ヤマトセマルヨツメハネカクシ（和名新称）HON
tsurumii Y. Watanabe, 1990c: 83 ニッコウセマルヨツメハネカクシ（和名新称）HON
yasudai Y. Watanabe, 1990c: 86 ダイセツセマルヨツメハネカクシ（和名新称）HOK
Genus Olophrum Erichson, 1839 ハネナガヨツメハネカクシ属（和名新称）
arrowi Scheerpeltz, 1929a: 105 アロウハネナガヨツメハネカクシ（和名改称）HOK, HON, Oki, SHI, KYU
japonicum Scheerpeltz, 1929a: 110 ヤマトハネナガヨツメハネカクシ（和名新称）HON
mutatum Scheerpeltz, 1929a: 100 ニセカクムネハネナガヨツメハネカクシ（和名改称）Japan
simplex Sharp, 1874a: 97 ナガサキハネナガヨツメハネカクシ（和名改称）HON, KYU; China
subsolanum Y. Watanabe, 1990c: 132 カントウハネナガヨツメハネカクシ（和名新称）HON
tadashii Y. Watanabe, 1990c: 135トゲアシハネナガヨツメハネカクシ（和名新称）HON
vicinum Sharp, 1889: 472 カクムネハネナガヨツメハネカクシ（和名改称）HON
yasudai Y. Watanabe, 1990c: 139 ダイセツカクムネハネナガヨツメハネカクシ（和名新称）HOK
Genus Orochares Kraatz, 1857 ツヤヨツメハネカクシ属（和名新称）2013 The Kyushu

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Catalogue of Japanese Staphylinidae (Insecta: Coleoptera)

debilis Sharp, 1889: 472 (Amphichroum) キバネツヤヨツメハネカクシ HOK, HON, SHI, KYU= japonica Cameron, 1933: 168 (Orochares)
Genus Philydrodes Bernhauer, 1929 カタホソヨツメハネカクシ属（和名新称）
Subgenus Minyphilydrodes Watanabe, 1990
asamanis Y. Watanabe, 1990c: 337 アサマカタホソヨツメハネカクシ（和名新称）HON
biguttatus Y. Watanabe, 1990c: 343 フタモンカタホソヨツメハネカクシ（和名新称）HON
chichibunis Y. Watanabe, 1990c: 347 チチブカタホソヨツメハネカクシ（和名新称）HON
flavidus Y. Watanabe, 1990c: 344 キイロカタホソヨツメハネカクシ（和名新称）HON
ishidai Nakane & Sawada, 1956b: 182 モンカタホソヨツメハネカクシ（和名改称）HON
kumosoensis Y. Watanabe, 2012a: 20 クモソウカタホソヨツメハネカクシ（和名改称）SHI
maculatus Y. Watanabe, 1990c: 340 ハクバカタホソヨツメハネカクシ（和名新称）HON
niger Y. Watanabe, 1990c: 349 クロカタホソヨツメハネカクシ（和名新称）HON
parallelus Y. Watanabe, 1990c: 338 シナノミヤマカタホソヨツメハネカクシ（和名新称）HON
shimomurai Y. Watanabe, 1990c: 346 シモムラカタホソヨツメハネカクシ（和名新称）HON
watanabei R. Dvořák, 1957: 9 ワタナベカタホソヨツメハネカクシ（和名改称）HON
yujii Y. Watanabe, 2012a: 17 アワカタホソヨツメハネカクシ（和名改称）SHI
Subgenus Philydrodes Bernhauer, 1929
aquatilis Sharp, 1889: 469 (Anthophagus) クロカタホソヨツメハネカクシ（和名改称）HON= japonicum Yoshida & Nomura, 1952: 6 (Coryphium)dewanus Y. Watanabe, 1990c: 325 デワカタホソヨツメハネカクシ（和名新称）HON
gigas Y. Watanabe, 1990c: 317 オオクロカタホソヨツメハネカクシ（和名新称）HON
grandis Nakane & Sawada, 1956b: 181 オオカタホソヨツメハネカクシ（和名新称）HON
hikosanensis Nakane & Sawada, 1956b: 180 オオズカタホソヨツメハネカクシ（和名改称）KYU, SHI
horioi Nakane & Sawada, 1956b: 180 ミヤマカタホソヨツメハネカクシ（和名改称）HON
kawaseorum Hayashi, 1994a: 133 カワセカタホソヨツメハネカクシ（和名新称）HON
michikoae Y. Watanabe, 1990c: 323 ミチノクカタホソヨツメハネカクシ（和名新称）HON
minipunctatus Y. Watanabe, 1990c: 321 アルプスカタホソヨツメハネカクシ（和名新称）HON
rubidus Nakane & Sawada, 1956b: 180 アカバカタホソヨツメハネカクシ（和名新称）HON
rufescens Y. Watanabe, 1990c: 327 シコクカタホソヨツメハネカクシ（和名新称）SHI
sanukiensis Y. Watanabe, 2009c: 343 サヌキカタホソヨツメハネカクシ（和名改称）SHI
similis Nakane & Sawada, 1956b: 181 ニセミヤマカタホソヨツメハネカクシ（和名新称）HON
takaoensis Nakane & Sawada, 1956b: 182 タカオカタホソヨツメハネカクシ（和名新称）HON
tsurugisanus Y. Watanabe, 1990c: 326 ツルギカタホソヨツメハネカクシ（和名新称）SHI
yoshidai Y. Watanabe, 1990c: 328 ヨシダカタホソヨツメハネカクシ（和名新称）SHI
zipanguensis Y. Watanabe, 1990c: 333 ニホンカタホソヨツメハネカクシ（和名新称）HON
Genus Trigonodemus LeConte, 1863 ヨツメシデムシモドキ属（和名新称）
lebioides Kraatz, 1877: 105 (Arimimelus) クロモンヨツメシデムシモドキ（和名改称）HON, Oki, SHI, KYU; China

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Yasutoshi Shibata, Munetoshi Maruyama, Hideto Hoshina, Toshio Kishimoto, Shun-Ichiro Naomi, Shûhei Nomura, Volker Puthz, Takashi Shimada, Yasuaki Watanabe, Shûhei Yamamoto

Tribe Aphaenostemmini Peyerimhoff, 1914
Genus Giulianium Moore, 1976 ハマベヨツメハネカクシ属
alaskanum Ahn & Ashe, 1999: 162 ハマベヨツメハネカクシ（和名改称）HOK; USA (Alaska)
Tribe Coryphiini Jacobson, 1908
Subtribe Boreaphilina Zerche, 1990
Genus Archaeoboreaphilus Zerche, 1990 カクムネヨツメハネカクシ属（和名新称）
fuscus Y. Watanabe, 1988: 53 (Coryphium) スホウカクムネヨツメハネカクシ（和名新称）HON
iwakisanensis Y. Watanabe, 1988: 54 (Coryphium) イワキカクムネヨツメハネカクシ（和名新称）HON
miyamorii Y. Watanabe, 1988: 49 (Coryphium) ミヤモリカクムネヨツメハネカクシ（和名新称）HOK, HON
nikkoensis Sawada, 1964a: 36 (Boreaphilus) ニッコウカクムネヨツメハネカクシ（和名新称）HON
tateoi Y. Watanabe, 1988: 52 (Coryphium) ヤマトカクムネヨツメハネカクシ（和名新称）HON
yasutoshii Y. Watanabe, 1988: 58 (Coryphium) クロカクムネヨツメハネカクシ（和名新称）HON
Genus Boreaphilus Sahlberg, 1832 ムネボソヨツメハネカクシ属
graciliformis Zerche, 1990: 183 ホソムネボソヨツメハネカクシ（和名新称）HON; South Korea
hokkaidensis Zerche, 1993: 347 エゾムネボソヨツメハネカクシ（和名新称）HOK; Russia (Sakhalin)japonicus Sharp, 1874a: 96 ムネボソヨツメハネカクシ HON, Oki, SHI, KYU; South Korea, China, Russia (Far East)lewisianus Sharp, 1874a: 96 ルイスムネボソヨツメハネカクシ（和名新称）HON, SHI
temporalis Zerche, 1990: 189 ヨシノムネボソヨツメハネカクシ（和名新称）HON
Subtribe Coryphiina Jacobson, 1908
Genus Coryphium Stephens, 1834 ズビロチビヨツメハネカクシ属
yasudai Y. Watanabe, 2007a: 299 ズビロチビヨツメハネカクシ HOKGenus Eudectus Redtenbacher, 1857 シリブトヨツメハネカクシ属
japonicus Zerche, 1990: 157 エゾシリブトヨツメハネカクシ（和名新称）HOK
rufulus Weise, 1877a: 96 シリブトヨツメハネカクシ HOK, HON, Iz-Hch, SHI, KYU
Genus Planeboreaphilus Shibata, 1970 イブシヨツメハネカクシ属（和名新称）
coriaceoides Shibata, 1970: 59 イブシヨツメハネカクシ（和名新称）HOK, HON
Tribe Eusphalerini Hatch, 1957
Genus Eusphalerum Kraatz, 1857 ハナムグリヨツメハネカクシ属（和名新称）
Subgenus Eusphalerum Kraatz, 1857
Group of E. pollens ハナムグリヨツメハネカクシ種群

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Catalogue of Japanese Staphylinidae (Insecta: Coleoptera)amamiense Y. Watanabe, 1993: 803 アマミハナムグリヨツメハネカクシ（和名改称）Amm
aoba Y. Watanabe, 1990c: 219 オオツキハナムグリヨツメハネカクシ（和名新称）HON
bonjuense Y. Watanabe, 2003b: 398 ボンジュサンハナムグリヨツメハネカクシ（和名改称）HON
bosatsu Y. Watanabe, 1990c: 198 ボサツハナムグリヨツメハネカクシ（和名新称）HON
daisetsuense Y. Watanabe, 1990c: 215 ダイセツハナムグリヨツメハネカクシ（和名新称）HOK
dewanum Y. Watanabe, 1990c: 212 デワハナムグリヨツメハネカクシ（和名新称）HON
dogoenze Y. Watanabe & Shimada, 2006: 197ドウゴハナムグリヨツメハネカクシ（和名改称）Okiezoense Y. Watanabe, 1990c: 217 エゾハナムグリヨツメハネカクシ（和名新称）HOK
hidakanum Y. Watanabe, 1990c: 216 ヒダカハナムグリヨツメハネカクシ（和名新称）HOK
ideyu Y. Watanabe, 1990c: 189 イデユハナムグリヨツメハネカクシ（和名新称）HON
insulare Y. Watanabe & Shimada, 2006: 200 オキハナムグリヨツメハネカクシ（和名改称）Okikaiji Y. Watanabe, 1990c: 197 マストミハナムグリヨツメハネカクシ（和名新称）HON
komatsudakense Y. Watanabe, 2012b: 85 コマツダケハナムグリヨツメハネカクシ（和名改称）HON
kamui Y. Watanabe, 1990c: 213 カムイハナムグリヨツメハネカクシ（和名新称）HOK
kana Y. Watanabe, 1993: 807 カナハナムグリヨツメハネカクシ（和名改称）Amm
kasumi Y. Watanabe, 1990c: 210 カスミハナムグリヨツメハネカクシ（和名新称）HON
kumoma Y. Watanabe, 1990c: 196 クモマハナムグリヨツメハネカクシ（和名新称）HON
lewisi Cameron, 1930: 182 (Anthobium) ルイスハナムグリヨツメハネカクシ（和名改称）KYU
okutamaense Y. Watanabe, 1990c: 195 オクタマヨツメハナムグリハネカクシ（和名新称）HON
pollens Sharp, 1889: 476 (Anthobium) クロハナムグリヨツメハネカクシ（和名改称）HON, SHI, KYU, Yakporoshiriense Y. Watanabe, 1990c: 206 ポロシリハナムグリヨツメハネカクシ（和名新称）HOK
sagiri Y. Watanabe, 1990c: 203 サギリハナムグリヨツメハネカクシ（和名新称）HON
saigokuense Y. Watanabe, 1990c: 191 サイゴクハナムグリヨツメハネカクシ（和名新称）HON, SHI
sanae Y. Watanabe, 1990c: 211 ホソハナムグリヨツメハネカクシ（和名新称）HON
satsuki Y. Watanabe, 1990c: 201 サツキハナムグリヨツメハネカクシ（和名新称）HON
shibatai Y. Watanabe, 1990c: 207 シバタハナムグリヨツメハネカクシ（和名新称）HON
shikokuense Y. Watanabe, 1990c: 185 シコクハナムグリヨツメハネカクシ（和名新称）SHI
solitare Sharp, 1874a: 98 (Anthobium) ハラグロハナムグリヨツメハネカクシ（和名新称）HON, SHI, KYU; Taiwan= rufiventre Cameron, 1930: 182 (Anthobium)
takane Y. Watanabe, 1990c: 193 タカネハナムグリヨツメハネカクシ（和名新称）HON
tsugaruense Y. Watanabe, 2003b: 400 ツガルハナムグリヨツメハネカクシ（和名改称）HON
yakushimanum Y. Watanabe, 1990c: 190 ヤクシマハナムグリヨツメハネカクシ（和名新称）Yakyamanami Y. Watanabe, 1990c: 209 ジョウシュウハナムグリヨツメハネカクシ（和名新称）HON
yamauchii Y. Watanabe, 2003b: 396 ヤマウチハナムグリヨツメハネカクシ（和名改称）HON
yukiguni Y. Watanabe, 1990c: 205 エチゴハナムグリヨツメハネカクシ（和名新称）HON
Group of E. japonicum ケバネハナムグリヨツメハネカクシ種群
eifleri Zerche, 2007: 430 アキヨツメハナムグリハネカクシ（和名新称）HON
floricola Y. Watanabe, 2012b: 88 ミヤマハナムグリヨツメハネカクシ（和名改称）HON
hirtellum Fauvel, 1901a: 2 (Anthobium) ケハダハナムグリヨツメハネカクシ（和名新称）HON
japonicum Bernhauer, 1909a: 52 (Anthobium) ニホンハナムグリヨツメハネカクシ（和名新称）HON, Oki2013 The Kyushu

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Yasutoshi Shibata, Munetoshi Maruyama, Hideto Hoshina, Toshio Kishimoto, Shun-Ichiro Naomi, Shûhei Nomura, Volker Puthz, Takashi Shimada, Yasuaki Watanabe, Shûhei Yamamoto

kirizumiense Y. Watanabe, 1990c: 230 キリズミハナムグリヨツメハネカクシ（和名新称）HON
kyushuense Y. Watanabe, 1990c: 221 キュウシュウハナムグリヨツメハネカクシ（和名新称）KYU
parallelum Sharp, 1889: 476 (Anthobium) キイロハナムグリヨツメハネカクシ（和名改称）HOK, HON, SHI, KYU
senjoense Y. Watanabe, 1990c: 236 センジョウハナムグリヨツメハネカクシ（和名新称）HON
shizuokanum Y. Watanabe, 1990c: 227 シズオカハナムグリヨツメハネカクシ（和名新称）HON
tachiyazawanum Y. Watanabe, 1990c: 231 ウゼンハナムグリヨツメハネカクシ（和名新称）HON
towadaense Y. Watanabe, 1990c: 233トワダハナムグリヨツメハネカクシ（和名改称）HON
tsukushiense Y. Watanabe, 1990c: 223 ツクシハナムグリヨツメハネカクシ（和名新称）KYU
yasukawai Y. Watanabe, 1990c: 228 ヤスカワハナムグリヨツメハネカクシ（和名新称）HON
Group of E. michinoku ハダバネハナムグリヨツメハネカクシ種群
kaoru Y. Watanabe, 1990c: 242 シンシュウハナムグリヨツメハネカクシ（和名新称）HON
michinoku Y. Watanabe, 1990c: 239 ミチノクハナムグリヨツメハネカクシ（和名新称）HON
sasajii Hayashi, 2007b: 103 ササジハナムグリヨツメハネカクシ（和名改称）HON
Tribe Omaliini McLeay, 1825
Genus Acrolocha Thomson, 1858 モモブトヨツメハネカクシ属（和名新称）
horiguchii Y. Watanabe, 2007b: 433 モモブトヨツメハネカクシ HON
kanagawana Y. Watanabe, 2007c: 437 カナガワモモブトヨツメハネカクシ HON
miyamorii Y. Watanabe, 1990c: 68 ミヤモリモモブトヨツメハネカクシ（和名新称）HOK, Chs
Genus Acruliopsis Zerche, 2003 ヘリトゲヨツメハネカクシ属（和名新称）
denticollis Sharp, 1889: 475 (Omalium) ヘリトゲヨツメハネカクシ HOK, HON, Oki, SHI, KYU
nipponica Y. Watanabe, 1980b: 274 (Pycnoglypta) ヒメヘリトゲヨツメハネカクシ HOK, HON
watanabei Zerche, 2003: 307 ワタナベヘリトゲヨツメハネカクシ（和名新称）HON
Genus Carcinocephalus Bernhauer, 1903 ニセヨツメハネカクシ属
satoi Hayashi, 2007a: 49 サトウニセヨツメハネカクシ（和名新称）HON
Genus Nipponophloeostiba Watanabe, 1962 チビヒラタヨツメハネカクシ属（和名新称）
verrucifera Y. Watanabe, 1962b: 78 チビヒラタヨツメハネカクシ Iz-My, Iz-Mk, Iz-Hch, Yak, Iri
Genus Omalium Gravenhorst, 1802 ヨツメハネカクシ属
diffine Sharp, 1889: 474 ニセセミゾヨツメハネカクシ（和名新称）SHI, KYU, Yak
hibernum Hayashi, 2006: 75 フユセミゾヨツメハネカクシ（和名新称）HON
japonicum Sharp, 1874a: 98 セミゾヨツメハネカクシ HON, SHI, KYU
niponense Sharp, 1889: 474 ムナクボヨツメハネカクシ HOK, HON, SHI, KYU
shibatai Y. Watanabe, 1990c: 94 シバタセミゾヨツメハネカクシ（和名新称）HON
tenue Weise, 1877a: 95 アカチャセミゾヨツメハネカクシ（和名新称）HON

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Catalogue of Japanese Staphylinidae (Insecta: Coleoptera)

Genus Phloeonomus Heer, 1839 チビキカワヨツメハネカクシ属（和名新称）
Subgenus Phloeonomodes Smetana, 1964
japonicus Y. Watanabe, 1963b: 164 ナミチビキカワヨツメハネカクシ（和名新称）HON
Subgenus Phloeonomus Heer, 1839
kyushuensis Y. Watanabe, 1962a: 17 (Paraphloeostiba) チビキカワヨツメハネカクシ（和名新称）HOK, HON, KYU,
Ish, Iri
Genus Phloeostiba Thomson, 1858 ヒラタヨツメハネカクシ属
kamijoi Y. Watanabe, 2009a: 71 カミジョウヒラタヨツメハネカクシ HOK
plana Paykull, 1792: 145 (Staphylinus) ヒラタヨツメハネカクシ HOK, HON, SHI; Russia, Europe
Genus Xylodromus Heer, 1839 キカワヨツメハネカクシ属（和名新称）
daimio Sharp, 1889: 475 (Omalium) ダイミョウキカワヨツメハネカクシ（和名新称）HON, KYU
Genus Xylostiba Ganglbauer, 1895 ホソヨツメハネカクシ属（和名新称）
tahirai Y. Watanabe, 1997b: 760 ホソヨツメハネカクシ HON, Oki

Subfamily Proteininae Erichson, 1839 ハバビロハネカクシ亜科（２属26種）Tribe Proteinini Erichson, 1839
Genus Megarthrus Stephens, 1829 ハバビロハネカクシ属
aino Cuccodoro, 1996: 478 アイノハバビロハネカクシ（和名新称）HON
conformis Sawada, 1962b: 11 カドホホハバビロハネカクシ（和名新称）HON, SHI
conspirator Cuccodoro, 1996: 481 ニッコウハバビロハネカクシ（和名新称）HOK, HON, SHI; Russia (Sakhalin)constrictus Cuccodoro, 1996: 482 キタザワハバビロハネカクシ（和名新称）HON
convexus Sharp, 1874a: 100 セマルハバビロハネカクシ HOK, HON, SHI, KYU, Yakcorticalis Sharp, 1889: 468 エダモンハバビロハネカクシ HON, SHI, KYU
denticollis Beck, 1817: 26 (Omalium) キンキハバビロハネカクシ HON; Russia, Syria, Turkey, Europe
hemipterus Illiger, 1794: 597 (Silpha) カバイロハバビロハネカクシ HOK, HON, SHI; China, Russia, Europe= heterops Sawada, 1962b: 14
impressicollis Eppelsheim, 1893a: 67 キチャバネハバビロハネカクシ（和名新称）HON; Russia
incubifer Cuccodoro, 1996: 486 クロチャバネハバビロハネカクシ（和名新称）HOK, SHI
japonicus Sharp, 1874a: 99 ハバビロハネカクシ HOK, HON, Iz-My, Iz-Hch, SHI, KYU, Tsm, SKs, Yak; China,
Russiamontanus Sawada, 1962b: 14 イナゴユハバビロハネカクシ（和名改称）HOK, HON, SHI, Yak; China, Russia= subangulatus Sawada, 1962b: 14
parallelus Sharp, 1874a: 99 ムツバハバビロハネカクシ（和名新称）HOK, HON, Oki, SHI, KYU, Tsm, Yak

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Yasutoshi Shibata, Munetoshi Maruyama, Hideto Hoshina, Toshio Kishimoto, Shun-Ichiro Naomi, Shûhei Nomura, Volker Puthz, Takashi Shimada, Yasuaki Watanabe, Shûhei Yamamoto= subparallelus Sawada, 1962b: 12
sawadai Cuccodoro, 1996: 492 サワダハバビロハネカクシ（和名新称）HON
scriptus Sharp, 1889: 469 クロモンハバビロハネカクシ HOK, HON, Oki, SHI, KYU
shibatai Sawada, 1962b: 13 シバタハバビロハネカクシ（和名新称）HON
Genus Proteinus Latreille, 1797 チビハバビロハネカクシ属
awanus Hayashi, 2008: 172 アワチビハバビロハネカクシ SHI
crassicornis Sharp, 1874a: 100 チビハバビロハネカクシ HOK, HON, Iz-Hch, SHI, KYU; South Korea, China,
Russiaezoensis Hayashi, 1988: 17 キタチビハバビロハネカクシ HOK
gotoi Hayashi, 1986: 111 ゴトウチビハバビロハネカクシ HON
nitiduloides Hayashi, 2008: 169 コゲチャチビハバビロハネカクシ SHI
sawadai Hayashi, 1988a: 18 サワダチビハバビロハネカクシ HON
shibatai Hayashi, 1986: 109 シバタチビハバビロハネカクシ HON
taichii Hayashi, 2008: 166 ツルギチビハバビロハネカクシ SHItateoitoi Hayashi, 1988: 20 イトウチビハバビロハネカクシ HON
yoshidai Hayashi, 2000: 92 ヨシダチビハバビロハネカクシ SHI

Subfamily Micropeplinae Leach, 1815 チビハネカクシ亜科（１属８種）
Genus Micropeplus Latreille, 1809 セスジチビハネカクシ属
fulvus japonicus Sharp, 1874a: 101 セスジチビハネカクシ HON, SHI, KYU, Tsm; China, India
hiromasai Y. Watanabe & Shibata, 1964: 64 ナガセスジチビハネカクシ（和名改称）HOK, HON, Oki, KYU
horii Y. Watanabe, 2004b: 80 ホリセスジチビハネカクシ HOK
kikuchii Y. Watanabe, 1975: 323 キクチセスジチビハネカクシ HON
okiensis Y. Watanabe, 1990b: 37 オキセスジチビハネカクシ HON, Okisatoi Y. Watanabe, 1975: 319 サトウセスジチビハネカクシ HOK
sharpi Sawada, 1964a: 35 シャープセスジチビハネカクシ HON, SHI
yasutoshii Y. Watanabe, 1975: 309 ヒメセスジチビハネカクシ HON

Subfamily Dasycerinae Reitter, 1887 ニセマキムシ亜科（１属１種）
Genus Dasycerus Brongniart, 1800 イトヒゲニセマキムシ属
japonicus Nakane, 1963: 22 イトヒゲニセマキムシ HOK, HON, SHI

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日本産ハネカクシ科総目録（昆虫綱：甲虫目）
Catalogue of Japanese Staphylinidae (Insecta: Coleoptera)

Subfamily Pselaphinae Latreille, 1802 アリヅカムシ亜科（83属326種23亜種）
Supertribe Batrisitae Reitter, 1882 ムネトゲアリヅカムシ上族
Tribe Batrisini Reitter, 1882
Subtribe Batrisina Reitter, 1882
Genus Arthromelodes Jeannel, 1954 ヒサゴアリヅカムシ属（和名新称）
aizuanus Nomura, 1991: 355 アイヅヒサゴアリヅカムシ（和名新称）HON
corniventris Nomura, 1991: 347トゲヒサゴアリヅカムシ（和名新称）HON
crucifer Nomura, 1991: 358 ヤクヒサゴアリヅカムシ（和名新称）Yak
dilatatus Raffray, 1909: 25 (Batrisocenus) ハラビロヒサゴアリヅカムシ（和名新称）HON; China?
daibosatsuanus daibosatsuanus Nomura, 1991: 352 ダイボサツヒサゴアリヅカムシ（和名新称）HON
d. fujimontanus Nomura, 1991: 354 HON
d. shiranemontanus Nomura, 1991: 353 HON
giganteus Nomura, 1991: 341 オオヒサゴアリヅカムシ Amm, Tokgyoja Nomura, 1991: 345 ギョウジャヒサゴアリヅカムシ（和名新称）HON
hikosanus Nomura, 1991: 349 ヒコサンヒサゴアリヅカムシ（和名新称）KYU
kiiensis Nomura, 1991: 343 キイヒサゴアリヅカムシ（和名新称）HON
loebli Nomura, 1991: 351 シコクヒサゴアリヅカムシ SHI
mercurius Nomura, 1991: 348 キュウシュウヒサゴアリヅカムシ（和名新称）KYU
optatus Sharp, 1874b: 112 (Batrisus) ナガサキヒサゴアリヅカムシ（和名新称）KYU
pilicollis Nomura, 1991: 342 エンシュウヒサゴアリヅカムシ（和名新称）HON
punctifrons Nomura, 1991: 340 アマミヒサゴアリヅカムシ Amm
saikaiensis Nomura, 1991: 356 サイカイヒサゴアリヅカムシ KYU
sinuatipes Nomura, 1991: 354 ソリアシヒサゴアリヅカムシ（和名新称）HON
thysanoventris Nomura, 1991: 346 フサヒサゴアリヅカムシ SHIwatanabei Arai, 2002: 275 ワタナベヒサゴアリヅカムシ HON
Genus Babascenellus Nomura, 1995 ババツノヒゲアリヅカムシ属（和名新称）
macroscapus Nomura, 1995a: 125 ババツノヒゲアリヅカムシ Myk, Ish, Iri, Hat, Yon
Genus Basitrodes Jeannel, 1958 ツノアリヅカムシ属（和名新称）
godzilla Nomura, 2003: 175 ゴジラツノアリヅカムシ HON
hakusanus Nomura, 2002: 327 ハクサンツノアリヅカムシ HON
kasaharai Nomura, 2002: 322 カサハラツノアリヅカムシ HON
oscillator Sharp, 1883: 309 (Batrisus) ジョウエツツノアリヅカムシ HOK, HON
vestitus Sharp, 1883: 307 (Batrisus) チュウゼンジツノアリヅカムシ HOK, HON; Russia
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Yasutoshi Shibata, Munetoshi Maruyama, Hideto Hoshina, Toshio Kishimoto, Shun-Ichiro Naomi, Shûhei Nomura, Volker Puthz, Takashi Shimada, Yasuaki Watanabe, Shûhei Yamamoto

cornutus Kurbatov, 1984: 145 クナシリツノアリヅカムシ（和名新称）Kun
Genus Batriscenaulax Jeannel, 1958 ハケスネアリヅカムシ属（和名新称）
kleinzach Nomura, 1991: 324 ムナビロハケスネアリヅカムシ Ish, Iri
kunigamensis Nomura, 1991: 326 クニガミハケスネアリヅカムシ Okn
longipes longipes Jeannel, 1958: 57 ナガスネアリヅカムシ HOK, HON, Aws, SHI, KYU
l. hayato Nomura, 1991: 331 KYU
l. kumaso Nomura, 1991: 331 KYU
modestus Sharp, 1874b: 116 (Batrisus) ハケスネアリヅカムシ HON, Sad, SHI, KYU= furuhatai Kubota, 1944: 6 (Batrisocenus)
nagensis Nomura, 1991: 327 ナゴハケスネアリヅカムシ Okn
Genus Batriscenellus Jeannel, 1958 ハラクボアリヅカムシ属（和名新称）
Subgenus Batriscenellinus Nomura, 1991 ウエノツヤアリヅカムシ亜属（和名新称）
uenoi Nomura, 1991: 321 ウエノツヤアリヅカムシ HOK, HON, SHI, KYU
Subgenus Batriscenellus Jeannel, 1958 ハラクボアリヅカムシ亜属（和名新称）
fallax Sharp, 1883: 318 (Batrisus) ヒメハラクボアリヅカムシ HOK, Kun, HON= insularis Kurbatov, 1990a: 71
fragilis Sharp, 1883: 317 (Batrisus) ホソハラクボアリヅカムシ HOK, HON, SHI, KYU
japonicus Sharp, 1883: 318 (Batrisus) ハラクボアリヅカムシ（和名新称）HOK, HON, Sad, KYU
kujumontanus Nomura, 1991: 306 キュウシュウハラクボアリヅカムシ HON, KYU
nodulifer Nomura, 1991: 309 ブンゴハラクボアリヅカムシ（和名新称）KYU
omogensis Nomura, 1991: 307 オモゴハラクボアリヅカムシ SHI
puncticollis Sharp, 1883: 316 (Batrisus) アラメハラクボアリヅカムシ HON= punctatus Nomura, 1991: 303
vicarius Löbl, 1973: 322 チョウセンハラクボアリヅカムシ（和名新称）Tsm; North Korea, South Korea, Cheju-do,
RussiaSubgenus Nipponoscenellus Nomura, 1991 ニッポンアナバラアリヅカムシ亜属（和名新称）
transformis Nomura, 1991: 311 ニッポンアナバラアリヅカムシ HON
Subgenus Scaioscenellus Jeannel, 1958 アナバラアリヅカムシ亜属（和名新称）
insulicola Nomura, 1991: 316 シマツヤアリヅカムシ Oke
ohishii Nomura, 1991: 315 オオイシツヤアリヅカムシ HON, KYU
orientalis Löbl, 1973: 322 (Batrisiella) HOK?（註１）; South Korea, China= brachygaster Nomura & Lee, 1993: 13
pilosellus Nomura, 1991: 318 ケモンツヤアリヅカムシ HON, KYU
sakaii Nomura, 1991: 320 サカイツヤアリヅカムシ Okn
shiranui Nomura, 1991: 314 シラヌイツヤアリヅカムシ（和名新称）KYU
similis Sharp, 1883: 319 (Batrisus) アナバラアリヅカムシ HON, Sad
Genus Batrisceniola Jeannel, 1958 アナズアリヅカムシ属（和名新称）2013 The Kyushu

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Catalogue of Japanese Staphylinidae (Insecta: Coleoptera)dissimilis Sharp, 1874b: 116 (Batrisus) アナズアリヅカムシ HON, Sad, Oki, SHI
hiranoi Nomura, 1991: 362 ヒラノアナズアリヅカムシ HON
semipunctulata Raffray, 1909: 25 (Batrisocenus) メンガタアリヅカムシ HOK, HON, KYU
Genus Batrisodellus Jeannel, 1958 ニッポンムネトゲアリヅカムシ属（和名新称）
acuminatus Sharp, 1883: 307 (Batrisus)トガリムネトゲアリヅカムシ KYU
cerberus Tanabe & Nakane, 1989: 739 ゴンゲンホラトゲアリヅカムシ（和名新称）KYU
coprea Tanabe & Nakane, 1989: 738 クゴウホラトゲアリヅカムシ（和名新称）HON
cristatus Jeannel, 1958: 32 (Basitrodes)トサカトゲアリヅカムシ HON
laticollis Jeannel, 1958: 29 (Basitrodes) ヒロムネトゲアリヅカムシ（和名新称）KYU
longulus Jeannel, 1958: 30 (Basitrodes) ナガムネトゲアリヅカムシ HON
nipponensis Raffray, 1909: 23 (Batrisodes) ニッポンムネトゲアリヅカムシ HON, Yak, Kue= risor Tanabe & Nakane, 1989: 735= vulgaris Raffray, 1909: 24 (Batrisodes)
palpalis Sharp, 1883: 306 (Batrisus) ヨコヅナトゲアリヅカムシ HON
punctipennis Sharp, 1883: 305 (Batrisus) アナバネムネトゲアリヅカムシ（和名新称）HON
Genus Batrisodes Reitter, 1882 オニアリヅカムシ属（和名新称）
Subgenus Batrisodes Reitter, 1882 オニアリヅカムシ亜属（和名新称）
galloisi Jeannel, 1958: 24 ガロアオニアリヅカムシ HON
vargus Kurbatov, 1992a: 30 クナシリオニアリヅカムシ（和名新称）Kun
Subgenus Excavodes Park, 1951 コオニアリヅカムシ亜属（和名新称）
acutifrons Jeannel, 1958: 20 コオニアリヅカムシ HON
angustus Sharp, 1874b: 113 (Batrisus) オオオニアリヅカムシ（和名新称）HON
caviceps Sharp, 1883: 308 (Batrisus) オサムネトゲアリヅカムシ（和名改称）SHI, KYU
dorsalis Jeannel, 1958: 18 タカオトゲアリヅカムシ HON
ornatifrons Sharp, 1883: 313 (Batrisus) ニッコウコオニアリヅカムシ HON
rugicollis Sharp, 1883: 313 (Batrisus) ハコネトゲアリヅカムシ HON
sennin Nomura, 2007a: 55 センニントゲアリヅカムシ Yon
stipes Sharp, 1874b: 115 (Batrisus) アラメトゲアリヅカムシ HOK, HON, SHI, KYU= coiffaiti Jeannel, 1958: 42 (Speobatrisodes)
Subgenus Pubimodes Park, 1951 タマヒゲオニアリヅカムシ亜属（和名新称）
globulifer Jeannel, 1958: 25 タマヒゲオニアリヅカムシ（和名新称）HON, KYU
harmandi Raffray, 1904: 34 アルマンオニアリヅカムシ（和名新称）HON; Russia= epistomalis Raffray, 1904: 35
Genus Batrisoschema Reitter, 1884 クチキムネトゲアリヅカムシ属（和名新称）
euplectiforme Sharp, 1883: 303 (Batrisus) クチキムネトゲアリヅカムシ HON, SHI, KYU
Genus Batrisus Aubé, 1833 チイロアリヅカムシ属（和名新称）2013 The Kyushu

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Yasutoshi Shibata, Munetoshi Maruyama, Hideto Hoshina, Toshio Kishimoto, Shun-Ichiro Naomi, Shûhei Nomura, Volker Puthz, Takashi Shimada, Yasuaki Watanabe, Shûhei Yamamoto= Genus Batristilbus Raffray, 1909
politus Sharp, 1883: 310 エグリチイロアリヅカムシ HOK, HON, SHI, KYU, Iki
trichothorax Tanokuchi, 1988a: 72 (Batristilbus) ケムネチイロアリヅカムシ HON, KYU
Genus Coryphomodes Jeannel, 1960 フタアナムネトゲアリヅカムシ属（和名新称）
dionysius Schaufuss, 1888: 12 [RN] (Batrisus) フタアナムネトゲアリヅカムシ HON, Oki, KYU= brevispinus Jeannel, 1958: 48 (Coryphomus)= spinicollis Sharp, 1883: 304 [HN] (Batrisus)
Genus Dendrolasiophilus Nomura, 2010 クサアリアリヅカムシ属
concolor Sharp, 1883: 310 (Batrisus) ツヤクサアリアリヅカムシ HOK, HON, SHI, KYU
nishikawai Nomura, 2008a: 140 ニシカワクサアリアリヅカムシ HOK, HON
Genus Kigatrodes Jeannel, 1958 キガムネトゲアリヅカムシ属（和名新称）
gracilis Sharp, 1883: 315 (Batrisus) ホソキガムネトゲアリヅカムシ（和名新称）HON
solitarius Sharp, 1883: 314 (Batrisus) キガムネトゲアリヅカムシ（和名新称）HON
Genus Maajappia Nomura, 2010 モノノケアリヅカムシ属
omotonis Nomura, 2010a: 57 オモトモノノケアリヅカムシ IshGenus Petaloscapus Jeannel, 1958 ヒゲブトムネトゲアリヅカムシ属（和名新称）
basicornis Sharp, 1883: 312 (Batrisus) ヒゲブトムネトゲアリヅカムシ HON, Sad
fissifrons fissifrons Sharp, 1883: 311 (Batrisus) ヒゴヒゲブトムネトゲアリヅカムシ（和名新称）KYU
f. stricticollis Jeannel, 1958: 37 HON
hirsutus Tanokuchi, 1986: 271 ハママツムネトゲアリヅカムシ HON
nasutus Jeannel, 1958: 36 ナスヒゲブトムネトゲアリヅカムシ HON, KYU
ornatus Sharp, 1874b: 114 (Batrisus) カザリヒゲブトムネトゲアリヅカムシ HON, Sad, KYU, Gotsasajii Nomura, 2001c: 309 ササジムネトゲアリヅカムシ HON
temporalis temporalis Jeannel, 1958: 37 エラハリムネトゲアリヅカムシ HON
t. yoshidai Nomura, 2001c: 315 SHIGenus Physomerinus Jeannel, 1952 モモブトアリヅカムシ属（和名新称）
hasegawai Nomura, 1991: 336 ハセガワモモブトアリヅカムシ Ishpedator Sharp, 1883: 319 (Batrisus) モモコブアリヅカムシ HON, SHI, KYU, Got, Yak; China= sauciipes Raffray, 1904: 50 (Batrisocenus)
schenklingi Raffray, 1912a: 105 (Batrisocenus) シェンクリングモモブトアリヅカムシ HOK?（註２）, Ish, Iri; Taiwan= akanensis Löbl, 1973: 324
Genus Speobatrisodes Jeannel, 1958 ホラアナトゲアリヅカムシ属（和名新称）
punctaticeps Jeannel, 1958: 41 ホラアナトゲアリヅカムシ SHI2013 The Kyushu

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Catalogue of Japanese Staphylinidae (Insecta: Coleoptera)

Genus Tribasodes Jeannel, 1958 ホソムネトゲアリヅカムシ属（和名新称）
dentatus Jeannel, 1958: 47 ホソムネトゲアリヅカムシ HON
kamedai Nomura, 2012: 13 クラホソムネトゲアリヅカムシ Okn
longicornis Sharp, 1883: 304 (Batrisus) ヒゲボソムネトゲアリヅカムシ HON , KYU
longipes Jeannel, 1958: 45ドウボソムネトゲアリヅカムシ HON
Genus Tribasodites Jeannel, 1960 カザリムネトゲアリヅカムシ属（和名新称）
coiffaiti Jeannel, 1958: 17 (Batrisodes) ヒメムネトゲアリヅカムシ HON, SHI, KYU
picticornis Nomura, 1986: 499 カザリムネトゲアリヅカムシ Okn, Iri
Genus Trisinus Raffray, 1894 フサヒゲアリヅカムシ属（和名新称）= Genus Batrisoplisus Raffray, 1908
amamianus Nomura, 1991: 287 (Batrisoplisus) アマミフサヒゲアリヅカムシ Amm
constrictus Nomura, 1991: 293 (Batrisoplisus) クビレメダカアリヅカムシ Ammesakii Nomura, 1991: 282 (Batrisoplisus) エサキフサヒゲアリヅカムシ KYU
galloisi Jeannel, 1958: 54 (Batrisoplisus) ガロアフサヒゲアリヅカムシ HON
inermis Nomura, 1991: 296 (Batrisoplisus) シリコブメダカアリヅカムシ（和名新称）SHI
monostatos Nomura, 1991: 289 (Batrisoplisus) オキナワメダカアリヅカムシ Okn, Okeokamotoi Nomura, 1991: 284 (Batrisoplisus) オカモトフサヒゲアリヅカムシ（和名新称）HON
parallelus Nomura, 1991: 297 (Batrisoplisus) ツツメダカアリヅカムシ（和名新称）Yak
raffrayi Jeannel, 1958 56 (Batrisoplisus) ラフレイフサヒゲアリヅカムシ HOK, HON, SHI, KYU, Yak= coiffaiti Jeannel, 1958: 56 (Batrisoplisus)
sagamianus Nomura, 1991: 275 (Batrisoplisus) サガミフサヒゲアリヅカムシ HON
satsumanus Nomura, 1991: 278 (Batrisoplisus) サツマフサヒゲアリヅカムシ（和名新称）KYU
sawadai Nomura, 1991: 277 (Batrisoplisus) サワダフサヒゲアリヅカムシ（和名新称）HON
subtilis Nomura, 1991: 278 (Batrisoplisus) ブンゴフサヒゲアリヅカムシ（和名新称）KYU
tamino Nomura, 1991: 292 (Batrisoplisus) タミノメダカアリヅカムシ Okn
tanabei Nomura, 1991: 284 (Batrisoplisus) タナベフサヒゲアリヅカムシ（和名新称）KYU
tenuiformis Nomura, 1991: 281 (Batrisoplisus) アズマフサヒゲアリヅカムシ HON
torticornis Nomura, 1991: 290 (Batrisoplisus) ネジレヒゲメダカアリヅカムシ Ish, Iritosanus Nomura, 1991: 283 (Batrisoplisus)トサフサヒゲアリヅカムシ HON, SHI, KYU= antennatus Weise, 1877a: 97 [HN] (Batrisus)= nomurai Löbl, 1998a: 463 [RN] (Batrisoplisus)
uroceratus uroceratus Nomura, 1991: 295 (Batrisoplisus) シリトゲメダカアリヅカムシ（和名改称）KYU
u. brevispinis Nomura, 1991: 296 (Batrisoplisus) KYU, Amk
venustus Jeannel, 1958: 55 (Batrisoplisus) ヒメフサヒゲアリヅカムシ HON, SHI
volcaninsulanus Nomura, 1991: 294 (Batrisoplisus) クチノエラブメダカアリヅカムシ（和名新称）Kue
Supertribe Clavigeritae Leach, 1815 ヒゲブトアリヅカムシ上族
Tribe Clavigerini Leach, 1815

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Yasutoshi Shibata, Munetoshi Maruyama, Hideto Hoshina, Toshio Kishimoto, Shun-Ichiro Naomi, Shûhei Nomura, Volker Puthz, Takashi Shimada, Yasuaki Watanabe, Shûhei Yamamoto

Subtribe Clavigerina Leach, 1815
Genus Anaclasiger Raffray, 1890 ネジレヒゲブトアリヅカムシ属（和名新称）
sinuaticollis Raffray, 1890: 216 ネジレヒゲブトアリヅカムシ Kue; Taiwan, Thailand, Malaysia, Singapore
Genus Articerodes Raffray, 1890 シマヒゲブトアリヅカムシ属（和名新称）
kishimotoi Nomura, 2001d: 348 オガサワラヒゲブトアリヅカムシ Og-hh
kurosawai Nomura, 2001d: 344 クロサワヒゲブトアリヅカムシ Og-hh, Og-ot
Genus Diartiger Sharp, 1883 ヤマトヒゲブトアリヅカムシ属（和名新称）
fossulatus fossulatus Sharp, 1883: 330 コヤマトヒゲブトアリヅカムシ HOK, HON
f. dentipes Nomura & Lee, 1992: 77 SHI (W-S); Korea, Cheju-do
f. hirashimai Nomura, 1997a: 86 KYU (C)f. imasakai Nomura, 1997a: 85 KYU (Shimabara)
f. ispartae Karaman, 1969: 50 (Coiffaitius) HON
f. izuinsulicola Nomura, 1997a: 82 HON (Is-Kz)
f. morimotoi Nomura, 1997a: 84 HON, Oki, SHI, KYU (N)f. tadauchii Nomura & Komatsu, 2012: 10 Tsm
f. toshioi Nomura & Komatsu, 2012: 12 HON (Iz-Mk)
kubotai Nomura, 1997a: 87 クボタヤマトヒゲブトアリヅカムシ HON, SHI, KYU
spinipes Sharp, 1883: 331 ヤマトヒゲブトアリヅカムシ KYU (C-S)
Genus Microdiartiger Sawada, 1963 ヒメヒゲブトアリヅカムシ属（和名新称）
japonicus Sawada, 1964b: 13 ヒメヒゲブトアリヅカムシ HON, SHI, KYU
Genus Micrelytriger Nomura, 1997 コバネヒゲブトアリヅカムシ属（和名改称）（註3）
mirabilis Nomura, 1997b: 116 コバネヒゲブトアリヅカムシ Yak, Amm
nakatai Nomura, 1997b: 120 オキナワコバネヒゲブトアリヅカムシ Okn
Genus Triartiger Kubota, 1944 ミフシヒゲブトアリヅカムシ属
reductus Nomura, 1997c: 441 ツシマミフシヒゲブトアリヅカムシ Tsm
Supertribe Euplectitae Streubel, 1839 ナガアリヅカムシ上族
Tribe Bythinoplectini Schaufuss, 1890
Subtribe Bythinoplectina Schaufuss, 1890
Genus Nipponozethus Coulon, 1989 ヒメシュモクアリヅカムシ属（和名新称）
delicatulus Jeannel, 1958: 5 (Zethopsus)トサヒメシュモクアリヅカムシ HON, SHI, KYU
lativentris Jeannel, 1958: 6 (Zethopsus) ヒメシュモクアリヅカムシ SHI, KYU

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日本産ハネカクシ科総目録（昆虫綱：甲虫目）
Catalogue of Japanese Staphylinidae (Insecta: Coleoptera)

Subtribe Pyxidicerina Raffray, 1903
Genus Parapyxidicerus Sawada, 1964 シュモクアリヅカムシ属（和名新称）
carinatus Sawada, 1964b: 11 シュモクアリヅカムシ HON, SHI, KYU
Tribe Dimerini Raffray, 1908
Genus Octomicrus Schaufuss, 1877 ヤツメアリヅカムシ属
enkaizanus Nomura, 2010b: 225 ヤツメアリヅカムシ HON
Tribe Euplectini Streubel, 1839
Subtribe Euplectina Streubel, 1839
Genus Euplectus Leach, 1817 ナガアリヅカムシ属（和名新称）
Subgenus Euplectus Leach, 1817 ナガアリヅカムシ亜属（和名新称）
sawadaianus Nomura, 2001a: 149 [RN] サワダナガアリヅカムシ HON= nanus Sawada, 1956b: 102 (Phthartomerus) [HN]
Subgenus Diplectellus Jeannel, 1950 オウシュウナガアリヅカムシ亜属（和名新称）
punctatus Mulsant & Rey, 1861: 76 オウシュウナガアリヅカムシ（和名新称）Kun; Europe, Caucasus, Irkutsk
Incertae sedis (subgenus?) 亜属不明
doryphorus Kurbatov, 1991b: 74 (Phthartomerus) シコクナガアリヅカムシ Kun, SHI
rubicundus Kurbatov, 1988: 1086 キタナガアリヅカムシ Kun, HON
Genus Leptoplectus Casey, 1908 コナガアリヅカムシ属（和名新称）
nipponensis Jeannel, 1958: 10 (Euplectus)トサコナガアリヅカムシ（和名新称）SHI
remyi Jeannel, 1961: 428 (Euplectus) コナガアリヅカムシ HON; India, Nepal, Swizerland, Senegal
solivagus Kurbatov, 1991b: 75 ソリヴァグスコナガアリヅカムシ（和名新称）Kun
occultus Kurbatov, 1991b: 76 オクルトゥスコナガアリヅカムシ（和名新称）Kun
illex Kurbatov, 1992b: 43 イレックスコナガアリヅカムシ（和名新称）Kun
Tribe Mayetini Winkler, 1925
Genus Mayetia Musant & Rey, 1875 ツチアリヅカムシ属（和名新称）
Subgenus Promayetia Coiffait, 1962 イシイツチアリヅカムシ亜属（和名新称）
ishiiana Nomura & Naomi, 1997: 557 イシイツチアリヅカムシ HON
Tribe Trichonychini Reitter, 1882
Subtribe Bibloporina Park, 1951
Genus Aphilia Reitter, 1882

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Yasutoshi Shibata, Munetoshi Maruyama, Hideto Hoshina, Toshio Kishimoto, Shun-Ichiro Naomi, Shûhei Nomura, Volker Puthz, Takashi Shimada, Yasuaki Watanabe, Shûhei Yamamoto

longicollis Jeannel, 1958: 7 (Pseudozibus) ホソヒメアリヅカムシ HON, SHI, KYU
Genus Bibloporus Thomson, 1859 ヤマトキカワアリヅカムシ属（和名新称）
Subgenus Bibloporus Thomson, 1859 ヤマトキカワアリヅカムシ亜属（和名新称）= Subgenus Synbibloporus Sawada, 1956
japonicus Sawada, 1956b: 101 ヤマトキカワアリヅカムシ HON, SHI, KYU
neglectus Kurbatov, 1993a: 28 クナシリキカワアリヅカムシ（和名新称）Kun
pallidus Raffray, 1912b: 426 パッリドゥスキカワアリヅカムシ（和名新称）HON (Kioto)
ponderosus Kurbatov, 1991a: 56 チシマキカワアリヅカムシ HOK, Kun, HON
pubens Kurbatov, 1991b: 72 プベンスキカワアリヅカムシ（和名新称）Kun
Genus Kuriporus Kurbatov, 1991 ケシツブキカワアリヅカムシ属（和名新称）
creator Kurbatov, 1991b: 71 ケシツブキカワアリヅカムシ Kun, SHI
Genus Piptoncus Kurbatov, 1991 ヨコヅナキカワアリヅカムシ属（和名新称）
duplex sobrinus Kurbatov, 1991b: 73 ヨコヅナキカワアリヅカムシ Kun, SHI
Subtribe Panaphantina Jeannel, 1950
Genus Acetalius Sharp, 1883 ハラフトアリヅカムシ属
dubius Sharp, 1883: 322 ハラフトアリヅカムシ HON, SHI, KYU, Tsm
pilosus Besuchet, 1985: 763 ケブカハラフトアリヅカムシ SHI, KYU
Genus Forinus Kurbatov, 1991 フォリヌスチビナガアリヅカムシ属（和名新称）
secundus Kurbatov, 1992b: 41 フォリヌスチビナガアリヅカムシ（和名新称）Kun
Genus Labroplectus Kurbatov, 1992 ラブロチビナガアリヅカムシ属（和名新称）
depositor Kurbatov, 1993a: 26 ラブロチビナガアリヅカムシ（和名新称）Kun
Genus Philiopsis Raffray, 1893 チビナガアリヅカムシ属（和名新称）
sulcifrons Jeannel, 1958: 8 チビナガアリヅカムシ（和名新称）（註4）SHI
Genus Philoscotus Sawada, 1957 オチバアリヅカムシ属
brevis Sawada, 1957: 24 マルガタオチバアリヅカムシ HON, SHI, KYU, Sad= coiffaiti Jeannel, 1959: 17 (Typhlacetalius)
longulus Sawada, 1957: 23 ナガオチバアリヅカムシ HON, Sad, SHI, KYU= pallidus Sawada, 1957: 24 (Philoscotus)
persimilis Nomura, 1988b: 743 オチバアリヅカムシ HON
rostralis Nomura, 1988b: 746 クチボソオチバアリヅカムシ HON, SHI, KYU, Yak
septentrionalis Nomura, 1988a: 520 キタオチバアリヅカムシ HON

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Catalogue of Japanese Staphylinidae (Insecta: Coleoptera)

sylvestris Nomura, 1988b: 739 モリオチバアリヅカムシ HON
tenuis Nomura, 1988a: 518 ホソオチバアリヅカムシ HON, KYU
venustus Nomura, 1988a: 516 ヒメオチバアリヅカムシ KYU
Subtribe Trimiina Bowman, 1934
Genus Saulcyella Reitter, 1882 コブヒゲアリヅカムシ属（和名新称）
schmidtii Märkel, 1845: 259 (Euplectus) コブヒゲアリヅカムシ HOK, HON; Russia, Europe
Supertribe Goniaceritae Reitter, 1882 オノヒゲアリヅカムシ上族
Tribe Brachyglutini Raffray, 1904
Subtribe Brachyglutina Raffray, 1904
Genus Batraxis Reitter, 1882 アトキリアリヅカムシ属（和名新称）
kawaharai Maruyama & Sugaya, 2004: 322 キタアトキリアリヅカムシ HOK
splendida Nomura, 1986: 501 ツヤアトキリアリヅカムシ Amm, Tok, Okn, Iri; South Korea
Genus Eupines King, 1866 チビマルアリヅカムシ属（和名新称）
sphaerica Motschulsky, 1851: 492 (Bryaxis) チビマルアリヅカムシ Ish, Iri; India, Indochina, Singapore, Malaysia(Borneo), Indonesia (Java, Sumatra, Sulawesi), New Guinea
Genus Noduliceps Jeannel, 1958 エビスアリヅカムシ属（和名新称）
latifrons Sharp, 1883: 325 (Bryaxis) エビスアリヅカムシ HON, SHI
Genus Pedisinops Newton & Chandler, 1989 サンゴアリヅカムシ属（和名新称）= Genus Halohermatus Sawada, 1991
regulus Sawada, 1991: 150 (Halohermatus) サンゴアリヅカムシ Ish, Iri
Genus Physoplectus Reitter, 1882 イソアリヅカムシ属（和名新称）= Genus Thalassomerus Sawada, 1992
miyakei Sawada, 1992: 58 (Thalassomerus) アマミイソアリヅカムシ Amm, Okn, Ish, Yon
reikoae Sawada, 1992: 56 (Thalassomerus) ヒラズイソアリヅカムシ HON
Genus Prosthecarthron Raffray, 1914 アシベアリヅカムシ属（和名新称）= Genus Barbiera Jeannel, 1952
sauteri Raffray, 1914: 3 (Prosthecarthron) アシベアリヅカムシ HON, SHI, KYU, Okn, Ish; Iri; Taiwan, North
Korea= palpalis Löbl, 1974: 97 (Briara)

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Yasutoshi Shibata, Munetoshi Maruyama, Hideto Hoshina, Toshio Kishimoto, Shun-Ichiro Naomi, Shûhei Nomura, Volker Puthz, Takashi Shimada, Yasuaki Watanabe, Shûhei Yamamoto

Genus Reichenbachia Leach, 1826 ホテイアリヅカムシ属（和名新称）
pulla Sharp, 1874b: 123 (Bryaxis) ホテイアリヅカムシ（和名新称）KYU
Genus Rybaxis Saulcy, 1876 ダイコクアリヅカムシ属
infuscata Raffray, 1909: 33 ヒメダイコクアリヅカムシ HON
kolorevi Kurbatov, 1984: 146 クナシリダイコクアリヅカムシ HOK, Kun
lamellifera Löbl, , 1973b: 326 チョウセンダイコクアリヅカムシ HON, KYU; North Korea
nigrescens Jeannel, 1958: 98 クロダイコクアリヅカムシ HON
pinguis Kurbatov, 1990b: 143 ペンギンダイコクアリヅカムシ KYU; Russia (Ussuri)
princeps Sharp, 1874b: 118 (Bryaxis) ダイコクアリヅカムシ HON, SHI, KYU
Genus Triomicrus Sharp, 1883 マルムネアリヅカムシ属
hamifer Löbl, Kurbatov & Nomura, 1998a: 85 コザマルムネアリヅカムシ Okn
melini Löbl, Kurbatov & Nomura, 1998a: 85 オキナワマルムネアリヅカムシ Okn
protervus Sharp, 1874b: 121 (Bryaxis) マルムネアリヅカムシ HON, Sad, SHI, KYU, Hej= galloisi Jeannel, 1958: 72
simplex Sharp, 1883: 326 ツヤマルムネアリヅカムシ HON, SHI, KYU, Got
sternalis Löbl, Kurbatov & Nomura, 1998a: 100トガリマルムネアリヅカムシ HON
sublaevis Raffray, 1909: 31 スベマルムネアリヅカムシ HON, KYU
Genus Trissemus Jeannel, 1949 エンマアリヅカムシ属
Subgenus Trissemus Jeannel, 1949 エンマアリヅカムシ亜属（和名新称）
alienus Sharp, 1874b: 120 (Bryaxis) ナミエンマアリヅカムシ HON, SHI, KYU, Hrd; North Korea, Russia?
clavatus Motschulsky, 1851: 491 (Bryaxis) リュウキュウヒゲブトエンマアリヅカムシ Okn, Yon; India
crassipes Sharp, 1874b: 125 (Bryaxis) アシブトエンマアリヅカムシ HON, KYU; North Korea
cubitus Sharp, 1874b: 122 (Bryaxis) ヒロエンマアリヅカムシ HON, SHI, KYU
curtus Sharp, 1874b: 124 (Bryaxis) ヒゲブトエンマアリヅカムシ HON, KYU= antilope Raffray, 1909: 30 (Reichenbachia)diffinis Sharp, 1883: 324 (Bryaxis) フトエンマアリヅカムシ HON, SHI
galloisi Jeannel, 1958: 101 ガロアエンマアリヅカムシ（和名新称）HON
implicitus Raffray, 1912a: 108 (Reichenbachia) タイワンヒゲブトエンマアリヅカムシ HON, SHI, KYU, T-Tak;
Taiwan, Vietnam
mundus Sharp, 1874b: 122 (Bryaxis) フタフシエンマアリヅカムシ HON, KYU
pseudalienus Kurbatov, 1990a: 76 エゾエンマアリヅカムシ（和名新称）HOK, Kun, OOs, HON; Russia (Sakhalin)
Tribe Bythinini Raffray, 1890
Genus Bryaxis Kugelann, 1794 オノヒゲアリヅカムシ属
affinis Sharp, 1883: 326 (Bythinus) ネブトオノヒゲアリヅカムシ HON, SHI, KYU
bushido Löbl, Kurbatov & Nomura, 1998b: 257 ブシドウオノヒゲアリヅカムシ SHI2013 The Kyushu

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Catalogue of Japanese Staphylinidae (Insecta: Coleoptera)

coiffaiti Jeannel, 1958: 88 コアフェーオノヒゲアリヅカムシ SHI
extremalis Kurbatov, 1990a: 75 チシマオノヒゲアリヅカムシ HOK, Kun, HON, SHI
gracilipalpis Jeannel, 1958: 90トサオノヒゲアリヅカムシ HON, SHI, KYU
harmandi Raffray, 1909: 38 アルマンオノヒゲアリヅカムシ HON
heian Löbl, Kurbatov & Nomura, 1998b: 231 ヘイアンオノヒゲアリヅカムシ HON
hisamatsui Löbl, Kurbatov & Nomura, 1998b: 248 ヒサマツオノヒゲアリヅカムシ SHI
hoko Löbl, Kurbatov & Nomura, 1998b: 243 ホコオノヒゲアリヅカムシ HON
humilis Raffray, 1909: 37 エクボオノヒゲアリヅカムシ HON; Russia (Far East)
iriomotensis Löbl, Kurbatov & Nomura, 1998b: 265 イリオモテオノヒゲアリヅカムシ Irijaponicus Sharp, 1874b: 125 (Bythinus) ヤマトオノヒゲアリヅカムシ HON, Sad, SHI, KYU, Hrd, Got; Russia (Far East)= brevipalpis Jeannel, 1958: 89
jomon Löbl, Kurbatov & Nomura, 1998b: 233 ジョウモンオノヒゲアリヅカムシ HON, SHI
kamakura Löbl, Kurbatov & Nomura, 1998b: 235 カマクラオノヒゲアリヅカムシ HON
karate Löbl, Kurbatov & Nomura, 1998b: 245 カラテオノヒゲアリヅカムシ HON
katana Löbl, Kurbatov & Nomura, 1998b: 241 カタナオノヒゲアリヅカムシ HON, SHI
kofun Löbl, Kurbatov & Nomura, 1998b: 232 コフンオノヒゲアリヅカムシ HON
koltzei Reitter, 1887: 269 (Bythinus) ウスリーオノヒゲアリヅカムシ HON; Cheju-do, North Korea, Russia= kintaro kintaro Nomura, 1995b: 130= kintaro odaiensis Nomura, 1995b: 134
mayumi Löbl, Kurbatov & Nomura, 1998b: 255 マユミオノヒゲアリヅカムシ HON, KYU
naginata Löbl, Kurbatov & Nomura, 1998b: 244 ナギナタオノヒゲアリヅカムシ HON
peckorum Löbl, Kurbatov & Nomura, 1998b: 240 ペックオノヒゲアリヅカムシ SHI
platalea Löbl, Kurbatov & Nomura, 1998b: 236 ヘラオノヒゲアリヅカムシ SHIreversus Sharp, 1883: 327 (Bythinus) ミフシオノヒゲアリヅカムシ HON, SHI, KYU
samurai Löbl, Kurbatov & Nomura, 1998b: 240 サムライオノヒゲアリヅカムシ HON
sauteri Raffray, 1909: 39 ザウターオノヒゲアリヅカムシ HON, SHI
sawadai Löbl, Kurbatov & Nomura, 1998b: 237 サワダオノヒゲアリヅカムシ HON
sparcepunctatus Jeannel, 1958: 93 コブオノヒゲアリヅカムシ HON, SHI
subseriatus Weise, 1877a: 98 (Bythinus) ナガオノヒゲアリヅカムシ HON, SHI, KYU= frontalis Jeannel, 1958: 87
sumo Löbl, Kurbatov & Nomura, 1998b: 263 スモウオノヒゲアリヅカムシ HON
tanto Löbl, Kurbatov & Nomura, 1998b: 245 タントウオノヒゲアリヅカムシ HON
taradakensis Löbl, Kurbatov & Nomura, 1998b: 247 タラダケオノヒゲアリヅカムシ KYU
tetralobus Löbl, Kurbatov & Nomura, 1998b: 264 カクオノヒゲアリヅカムシ HON
trichosus Jeannel, 1958: 94 ケブカオノヒゲアリヅカムシ HON
yari Löbl, Kurbatov & Nomura, 1998b: 246 ヤリオノヒゲアリヅカムシ HON
Incertae sedis (genus Bryaxis?) 属不確定
grandicollis Jeannel, 1958: 95 リュウガドウエンマアリヅカムシ SHI2013 The Kyushu

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Yasutoshi Shibata, Munetoshi Maruyama, Hideto Hoshina, Toshio Kishimoto, Shun-Ichiro Naomi, Shûhei Nomura, Volker Puthz, Takashi Shimada, Yasuaki Watanabe, Shûhei Yamamoto

Genus Bythoxenites Jeannel, 1958 カマヒゲアリヅカムシ属（和名新称）
brevicornis Löbl & Kurbatov, 1996: 927 シコクカマヒゲアリヅカムシ（和名新称）SHI
brevipilis Löbl & Kurbatov, 1996: 930 キョウトカマヒゲアリヅカムシ HON
frontalis Löbl & Kurbatov, 1996: 935 ジョウシュウカマヒゲアリヅカムシ（和名新称）HON
japonicus Yoshida & Nomura, 1952: 7 (Machaerites) ホラアナカマヒゲアリヅカムシ（和名改称）HON
longiceps Löbl & Kurbatov, 1996: 934 ナガカマヒゲアリヅカムシ（和名新称）HON
longicornis Löbl & Kurbatov, 1996: 925 ヒゲナガカマヒゲアリヅカムシ（和名新称）SHI
major Tanokuchi, 1990: 37 オオカマヒゲアリヅカムシ HON
pubiceps Löbl & Kurbatov, 1996: 932 ケブカカマヒゲアリヅカムシ（和名新称）HON
rimicola Tanokuchi, 1990: 40 ミネイシハラカマヒゲアリヅカムシ（和名新称）SHI
sulcithorax Tanokuchi, 1990: 34 ネジレカマヒゲアリヅカムシ HON= diversicornis Löbl & Kurbatov, 1996: 929= torticornis Löbl & Kurbatov, 1996: 928
Genus Tychobythinus Ganglbauer, 1896 オニオノヒゲアリヅカムシ属（和名新称）
aino Kurbatov, 1993b: 57 アイノオニオノヒゲアリヅカムシ Kun, HON
japonicus Löbl & Kurbatov, 1996: 921 ヤマトオニオノヒゲアリヅカムシ（和名新称）HON
Tribe Cyathigerini Schaufuss, 1872
Genus Plagiophorus Motschulsky, 1851 ダルマアリヅカムシ属（和名新称）= Genus Cyathiger King, 1865
= Genus Paracyathiger Jeannel, 1951
fujiyamai Kubota, 1944: 7 (Cyathiger) フジヤマダルマアリヅカムシ HON, Iz-Hch, SHI; Amm, Ish; South Korea,
Chinahispidus Sugaya, Nomura & Burckhardt, 2004: 150 ヤエヤマダルマアリヅカムシ Ish, Iri
Tribe Iniocyphini Park, 1951
Subtribe Natypleurina Newton & Thayer, 1992
Genus Morana Sharp, 1874 マメアリヅカムシ属
angustata Arai, 2003: 216 サツナンマメアリヅカムシ Amm, Tokbara Tanabe & Nakane, 1990: 28 ヤクシママメアリヅカムシ KYU, Yak, Tan, Krs, Kue
crassicostata Arai, 2003: 197 アマミマメアリヅカムシ Amm, T-Tak
deigo Arai, 2003: 199 オキナワマメアリヅカムシ Okn
donan Arai, 2003: 219 ヨナグニマメアリヅカムシ Yon
discedens Sharp, 1874b: 118 マメアリヅカムシ HON, Sad, SHI, KYU, Hrd, Got, Tanelegans Tanokuchi, 1988b: 529トウカイマメアリヅカムシ HON, SHI
kazuyoae Arai, 2003: 203 カズヨマメアリヅカムシ Oknlata Arai, 2003: 196 ヒロマメアリヅカムシ Yak

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日本産ハネカクシ科総目録（昆虫綱：甲虫目）
Catalogue of Japanese Staphylinidae (Insecta: Coleoptera)

maruyamai Arai, 2003: 212 タバゲマメアリヅカムシ T-Nak
oni Tanabe & Nakane, 1990: 27 オニマメアリヅカムシ KYU
puella Tanabe & Nakane, 1990: 26 カワラマメアリヅカムシ KYU
rhynchocephala Arai, 2003: 217 ヤエヤママメアリヅカムシ Yon
Genus Nipponobythus Jeannel, 1958 オオズアリヅカムシ属（和名新称）
expectatus Löbl, 1965d: 492 アナオオズアリヅカムシ（和名新称）HON
latifrons Jeannel, 1958: 81 オオズアリヅカムシ HON, Oki, SHI
omissus Löbl, 1965d: 494 ウンゼンオオズアリヅカムシ KYU; Korea, China
plagiatus Jeannel, 1958: 79 サルタオオズアリヅカムシ（和名改称）SHI
syrbatoides Jeannel, 1958: 79 モリオオズアリヅカムシ HON
Genus Takaorites Jeannel, 1958 ネジレヒゲアリヅカムシ属（和名新称）= Genus Bythonesiotes Jeannel, 1958 [HN]= Genus Bythiotes Newton & Chandler, 1989 [RN]
coiffaiti Jeannel, 1958: 76 (Bythonesiotes) アキヨシヒロズアリヅカムシ（和名改称）HON
longicornis Jeannel, 1958: 77 (Bythonesiotes) ヒゲナガヒロズアリヅカムシ SHI
torticornis Jeannel, 1958: 74 ネジレヒゲアリヅカムシ HON
Genus Tenguobythus Arai & Nomura, 2007 テングアリヅカムシ属
decoratus Arai & Nomura, 2007: 416 ヤエヤマテングアリヅカムシ Ish, Iri, Yon
nasalis Arai & Nomura, 2007: 410 テングアリヅカムシ HON, Iz-My, SHI, KYU
pentagonus Arai & Nomura, 2007: 413 オキナワテングアリヅカムシ Okn
rhinoceros Arai & Nomura, 2007: 412 アマミテングアリヅカムシ Amm
yama Arai & Nomura, 2007: 414 ヤマテングアリヅカムシ Okn
Tribe Tychini Raffray, 1904
Genus Hyugatychus Nomura, 1996 ヒュウガモリアリヅカムシ属（和名新称）
teizonagatomoi Nomura, 1996: 269 ヒュウガモリアリヅカムシ HON, SHI, KYU
tokunoshimensis Nomura, 1996: 272トクノシマモリアリヅカムシ Tok
Genus Tainochus Kurbatov, 1992 ムクゲアリヅカムシ属（和名新称）
imperator Kurbatov, 1992c: 143 チシマムクゲアリヅカムシ HOK, Kun, HON
insulicola Nomura & Lee, 1992: 74 (Atychodea) シマムクゲアリヅカムシ HON; Cheju-do
iwaoi Nomura, 1996: 250 オカモトムクゲアリヅカムシ HON
minutus Nomura, 1996: 258 シコクムクゲアリヅカムシ SHInitidus Nomura, 1996: 253 ツヤムクゲアリヅカムシ HON
puncticeps Nomura, 1996: 256 キュウシュウムクゲアリヅカムシ KYU

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Yasutoshi Shibata, Munetoshi Maruyama, Hideto Hoshina, Toshio Kishimoto, Shun-Ichiro Naomi, Shûhei Nomura, Volker Puthz, Takashi Shimada, Yasuaki Watanabe, Shûhei Yamamoto

Genus Tychus Leach, 1817 モリアリヅカムシ属（和名新称）
crassicornis Raffray, 1909:40 モリアリヅカムシ HON; China (Yunnan)?
dichotomus Nomura & Lee, 1992: 72 タイリクモリアリヅカムシ Kun, HON, KYU, Tsm; South Korea= kurilensis Kurbatov, 1992c: 142
yezoensis Nomura, 1996: 265 エゾモリアリヅカムシ HOK
yukihikoi Nomura, 1996: 262 ヒラノモリアリヅカムシ HON
Supertribe Pselaphitae Latreille, 1802 ヒゲナガアリヅカムシ上族
Tribe Ctenistini Blanchard, 1845
Genus Centrotoma Heyden, 1849 クロジュズヒゲアリヅカムシ属（和名新称）
prodiga Sharp, 1874b: 107 クロジュズヒゲアリヅカムシ HON, KYU
Genus Ctenisodes Raffray, 1897 ホソクシヒゲアリヅカムシ属（和名新称）= Genus Pilopius Casey, 1897
discedens Sharp, 1883: 296 (Ctenistes) ホソクシヒゲアリヅカムシ HON, Aws, Sad, KYU, Tsm
Genus Ctenistes Reichenbach, 1816 クシヒゲアリヅカムシ属（和名新称）
Subgenus Ctenistes Reichenbach, 1816 クシヒゲアリヅカムシ亜属（和名新称）
oculatus Sharp, 1874b: 110 クシヒゲアリヅカムシ HON, KYU
Genus Poroderus Sharp, 1883 カギクシヒゲアリヅカムシ属（和名新称）= Genus Poroderopsis Jeannel, 1960
armatus Sharp, 1874b: 111 (Ctenistes) カギクシヒゲアリヅカムシ HON, KYU
medius Sharp, 1874b: 111 (Ctenistes) ツヤクシヒゲアリヅカムシ HON, KYU, Hrd
mimeticus Sharp, 1883: 295 (Ctenistes)トゲクシヒゲアリヅカムシ（和名新称）HON, KYU
similis Sharp, 1874b: 112 (Ctenistes) ススキクシヒゲアリヅカムシ HON, KYU; North Korea
Genus Sognorus Reitter, 1882 ヒメクシヒゲアリヅカムシ属（和名新称）
breviceps Sharp, 1883: 296 (Ctenistes) ヒメクシヒゲアリヅカムシ HON, KYU, Hrd, Tsm; South Korea, North Korea
Tribe Hybocephalini Raffray, 1890
Genus Apharinodes Raffray, 1890 オオウロコアリヅカムシ属（和名新称）
papageno Nomura, 1989: 279 オオウロコアリヅカムシ Okn
Genus Stipesa Sharp, 1874 ハナダカアリヅカムシ属（和名新称）
rudis Sharp, 1874b: 109 ハナダカアリヅカムシ HON, SHI, KYU
Tribe Odontalgini Jeannel, 1949

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Catalogue of Japanese Staphylinidae (Insecta: Coleoptera)

Genus Odontalgus Raffray, 1877 スジバネアリヅカムシ属（和名新称）
coreanus Löbl, 1973: 329 チョウセンスジバネアリヅカムシ KYU; North Korea
masaoi Arai & Nomura, 2003: 194 スジバネアリヅカムシ HON
Tribe Pselaphini Latreille, 1802
Genus Hirashimanymus Nomura, 1990 カギアシアリヅカムシ属（和名新称）
schistodactyroides Nomura, 1990: 52 カギアシアリヅカムシ Ish, Iri, Yon; Taiwan
Genus Pselaphaulax Reitter, 1909 ヤマトヒゲナガアリヅカムシ属（和名新称）
japonicus Raffray, 1909: 42 (Pselaphus) ヤマトヒゲナガアリヅカムシ（和名改称）HOK, HON
Genus Pselaphogenius Reitter, 1910 アラメヒゲナガアリヅカムシ属
debilis Sharp, 1883: 328 (Pselaphus) アラメヒゲナガアリヅカムシ KYU (Nagasaki)
dentipenis Sawada, 1969: 13 ダイセンヒゲナガアリヅカムシ HON (Chûgoku)
gotoinsularis Nomura, 1998: 33 (Dicentrius) ゴトウヒゲナガアリヅカムシ Nkd
katsuyukii Nomura, 2001b: 284 タカオヒゲナガアリヅカムシ HON (Takao-san, Tôkyô)lanceolatus Sawada, 1969: 16 アマミヒゲナガアリヅカムシ Ammorientalis Besuchet, 1961b: 39 カギヒゲナガアリヅカムシ HON (North to Central)= uncifer Tanokuchi, 1981: 351
paradoxus paradoxus Sawada, 1969: 7 サイゴクヒゲナガアリヅカムシ HON (Kinki)
p. chiyokoae Nomura, 2000: 147 KYU (Saga, Fukuoka)
p. iyonis Nomura, 2000: 146 SHI (Ehime)
p. magnioculatus Sawada, 1971a: 89 HON (Chûgoku), KYU
patrius Nomura, 1999a: 261 マツウラヒゲナガアリヅカムシ KYU (Saga, N. Nagasaki)
seihiensis Nomura, 1999: 264 セイヒヒゲナガアリヅカムシ KYU (Seihi, Nagasaki)
shibatai Sawada, 1969: 12 シバタヒゲナガアリヅカムシ SHI
shimabaranus Nomura, 1999: 266 シマバラヒゲナガアリヅカムシ KYU (Shimabara, Tara-dake)shintaro Nomura, 1999: 266 タラダケヒゲナガアリヅカムシ KYU (Tara-dake)spinifer spinifer Sawada, 1969: 1トゲヒゲナガアリヅカムシ HON (Kinki)s. tuberculipenis Sawada, 1969: 4 HON (Ômine-san)s. uncinatus Sawada, 1969: 3 HON (Izumi-katsuragi-san)tridentatus tridentatus Sawada, 1969: 4 タカネヒゲナガアリヅカムシ HON (High mountain of C, Ibuki-yama)t. vicinus Sawada, 1969: 6 HON (Hokuriku)
tubipenis Sawada, 1969: 14 ツシマヒゲナガアリヅカムシ Tsmyoshidai Sawada, 1968: 3 ヨシダヒゲナガアリヅカムシ SHI
yoshii yoshii Sawada, 1969: 8 ヨシイヒゲナガアリヅカムシ HON (Ômine-san, Ôdaigahara)
y. hamatus Sawada, 1969: 11 HON (Yoshinoyama)
y. securiger Sawada, 1969: 11 HON (Wakayama)

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Yasutoshi Shibata, Munetoshi Maruyama, Hideto Hoshina, Toshio Kishimoto, Shun-Ichiro Naomi, Shûhei Nomura, Volker Puthz, Takashi Shimada, Yasuaki Watanabe, Shûhei Yamamoto

Genus Pselaphus Herbst, 1792 ヒゲナガアリヅカムシ属（和名新称）
lewisi Sharp, 1883: 329 ルイスヒゲナガアリヅカムシ（和名改称）SHI, KYU（註5）Genus Tyraphus Sharp, 1874 シバチアリヅカムシ属（和名新称）
nitidus Raffray, 1909: 43 シバチアリヅカムシ（和名新称）HON; North Korea, Russia (Far East)
Tribe Tmesiphorini Jeannel, 1949
Genus Raphitreus Sharp, 1883 ヒゲブトエクボアリヅカムシ属（和名新称）
speratus Sharp, 1874b: 109 (Tmesiphorus) ヒゲブトエクボアリヅカムシ HON, SHI, KYU
Genus Saltisedes Kubota, 1944 セダカアリヅカムシ属（和名新称）
brunneus Kubota, 1944: 9 セダカアリヅカムシ HOK, HON, SHI, KYU
Genus Tmesiphorus LeConte, 1849 ヒゲカタアリヅカムシ属（和名新称）
costatus Weise, 1877a: 99 ミスジヒゲカタアリヅカムシ HON
crassicornis Sharp, 1883: 299 ヒゲカタアリヅカムシ HOK, HON, Sad, KYU
princeps Sharp, 1883: 299 オオヒゲカタアリヅカムシ（和名新称）OOs, HON
Tribe Tyrini Reitter, 1882
Subtribe Somatipionina Jeannel, 1949
Genus Himepion Nomura & Hlaváč, 2003 エヒメアリヅカムシ属（和名新称）
cyathicornis Nomura & Hlaváč, 2003: 162 エヒメアリヅカムシ HON, SHI
Subtribe Tyrina Reitter, 1882
Genus Labomimus Sharp, 1883 アシナガアリヅカムシ属（和名新称）
reitteri Sharp, 1883: 300 アシナガアリヅカムシ HON
shibatai Sawada, 1961a: 43 オオアシナガアリヅカムシ HON, SHI
Genus Lasinus Sharp, 1874 オオトゲアリヅカムシ属（和名新称）
monticola Sawada, 1961a: 41 ヤマオオトゲアリヅカムシ HON, Sad, SHI, KYU, Hrd, Gotspinosus Sharp, 1874b: 106 オオトゲアリヅカムシ HOK, KYU; Russia
Genus Tyrus Aubé, 1833 コケアリヅカムシ属（和名新称）
japonicus Sharp, 1883: 302 コケアリヅカムシ（和名新称）HON, KYU
註 １：北海道からの記録は疑問．すなわち日本産ではない可能性が高い．台湾「唖口 (Akau) 」産標本のラベルのつづりが「阿寒 (Akan) 」と誤読されたのではないか？ Nomura (2004) 参照．

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Catalogue of Japanese Staphylinidae (Insecta: Coleoptera)註 ２：北海道からの記録は疑問．台湾「唖口 (Akau) 」産標本のラベルのつづりが「阿寒 (Akan) 」と誤読された可能性が高い．Nomura (2004) 参照．
註 ３：属記載時に与えられた属和名は「コバネアリヅカムシ属」であったが，これは「～ヒゲブト～」を落としたものであり．適切でない．
註 ４：Löbl & Besuchet, 2004 では， 本 種 はPhiliopsis とPseudoplectus の 双 方 に 記 述 が ある．確 かに 本 種 はPseudoplectus である疑いはあるが，今回本種をPseudoplectus 属と再同定した確実な記録を見出すことができなかったため，古い方を採ってPhiliopsisとした．
註 ５： Löbl & Besuchet, 2004では本属の日本産種として以下のような記述があるが，今回，日本からの確実な記録を見出すことはできなかった：(Pselaphus) striatus Löbl, 1974: 99 日本，北朝鮮，イラン，アフガニスタン

Subfamily Tachyporinae MacLeay, 1825 シリホソハネカクシ亜科（16属134種２亜種）
Tribe Deropini Smetana, 1983 ホソミズギワハネカクシ族（和名改称）
Genus Derops Sharp, 1889 ホソミズギワハネカクシ属（和名改称）
coreanus Y. Watanabe, 1969: 627 (Paraleaster) カンコクホソミズギワハネカクシ Tsm; South Korea, China, Russia(Ussuri)japonicus Sawada, 1956a: 42 (Paraleaster) ホソミズギワハネカクシ KYUlongicornis Sharp, 1889: 418 ヒゲナガホソミズギワハネカクシ HOK, HON, Oki, SHI, KYU; China, Russia (Far East)= kasugaensis Sawada, 1956a: 43 (Paraleaster)okinawanus Y. Watanabe, 1985: 447 オキナワホソミズギワハネカクシ Okn
yaeyamanus Kishimoto, 2001: 176 ヤエヤマホソミズギワハネカクシ（和名改称）Iri
Tribe Megarthropsini Cameron, 1919 ハバビロシリホソハネカクシ族
Genus Lacvietina Herman, 2004 アバタシリホソハネカクシ属
takashii Hayashi, 2003a: 114 (Peitawopsis) アバタシリホソハネカクシ Ish, Iri
Tribe Mycetoporini Thomson, 1859 キノコハネカクシ族
Genus Bolitobius Leach, 1819 ニセキノコハネカクシ属（和名新称）
longicornis L.-Z. Li & Sakai, 1998: 85 ヒゲナガニセキノコハネカクシ（和名新称）SHI
okamotoi L.-Z. Li & Sakai, 1998: 82 クロモンニセキノコハネカクシ（和名新称）SHI
parasetiger Schülke, 1993: 77 ルイスニセキノコハネカクシ（和名新称）HOK, HON, SHI, Tsm; North Korea,
China, Russia (Far East)
princeps Sharp, 1874a: 19 (Megacronus) チャバネニセキノコハネカクシ（和名改称）HON, SHI, KYU, Yak; China
setiger Sharp, 1874a: 18 (Megacronus) カタモンニセキノコハネカクシ（和名改称）HOK, HON, SHI, KYU; China,
Russia (Far East)

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Yasutoshi Shibata, Munetoshi Maruyama, Hideto Hoshina, Toshio Kishimoto, Shun-Ichiro Naomi, Shûhei Nomura, Volker Puthz, Takashi Shimada, Yasuaki Watanabe, Shûhei Yamamoto

Genus Bryophacis Reitter, 1909 キタキノコハネカクシ属（和名新称）
rufus punctipennis Thomson, 1861: 172 (Lordithon) キタキノコハネカクシ（和名新称）HOK; Russia, Mongolia, N
Europe= yasudai L.-Z. Li & Sakai, 1997: 91 (Ischnosoma)
Genus Bryoporus Kraatz, 1857 イクビキノコハネカクシ属（和名新称）
gracilis Sharp, 1888: 462 (Megacronus) アカイクビキノコハネカクシ（和名改称）HOK, HON, SHI, KYU, Tsm,
Tan, Yak; China
naomii L.-Z. Li, 1998: 17 ナオミイクビキノコハネカクシ（和名改称）HOK, HON, SHI, KYU
okamotoi L.-Z. Li, 1998: 20 オカモトイクビキノコハネカクシ（和名新称）SHI
ryukyuensis L.-Z. Li, Du & Lai, 1998: 52 リュウキュウイクビキノコハネカクシ（和名新称）Amm, Iriwadai Scheerpeltz, 1959: 97 ワダイクビキノコハネカクシ（和名新称）HON
Genus Carphacis Gozis, 1886 ハネスジキノコハネカクシ属
gigas Schülke, 1995a: 68 オオハネスジキノコハネカクシ（和名新称）HON, SHI, KYU; Russia (Far East)
kawanabei L.-Z. Li & Ohbayashi, 1998: 89 カワナベハネスジキノコハネカクシ（和名新称）SHI
nikkoensis Schülke, 1995a: 69 ニッコウハネスジキノコハネカクシ HOK, HON, KYU
paramerus Schülke, 1995a: 63 ニセハネスジキノコハネカクシ（和名新称）HOK, HON, KYU
striatus Oliver, 1795: [42]: 28 (Staphylinus) [HN] ハネスジキノコハネカクシ HOK, HON, SHI, KYU; South Korea,
Russia (Siberia), Europe, Algeria, Morocco
zerchei Schülke, 1995a: 72 ツェルヘハネスジキノコハネカクシ（和名新称）HON, SHI
Genus Ischnosoma Stephens, 1829 イクビハネカクシ属
bolitobioides Bernhauer, 1923a: 126 (Mycetoporus) トウヨウイクビハネカクシ（和名新称）HON, SHI; Taiwan,
China, Thailand, Oriental Region
convexum Sharp, 1888: 463 (Mycetoporus) ハネナガイクビハネカクシ（和名新称）HON, SHI, KYU, Yak, Okn,
Amm, Ish; Taiwan, Hong Kong, China, Thailand, Myanmar, India, Malaysia (Borneo), Indonesia (Sumatra),Solomon, Australian Region
discoidale Sharp, 1888: 463 (Mycetoporus) ヤマトイクビハネカクシ HON, SHI, KYU; Taiwan, China, Thailand,India
duplicatum Sharp, 1888: 464 (Mycetoporus) フタスジイクビハネカクシ HON, SHI, KYU; Taiwan, China, Thailand,
India, Nepal
fusciventre Tikhomirova, 1973: 160 (Bryoporus) キタイクビハネカクシ（和名新称）HOK; China, Russia (Far East)= paradiscoidale L.-Z. Li & Sakai, 1996a: 77
lewisium Sharp, 1874a: 19 (Mycetoporus) ルイスイクビハネカクシ（和名新称）SHI, KYU, Tsm= irregulare L.-Z. Li & Sakai, 1996a: 75
longicorne Mäklin, 1847: 12 (Mycetoporus) エゾイクビハネカクシ（和名新称）HOK; Russia (Siberia), Turkey,
Europe, N Africa, Nearctic Region= yasudai L.-Z. Li & Sakai, 1996a: 79
quadriguttatum japonicum Kocian, 2003: 39 ヒゲブトイクビハネカクシ（和名新称）HON

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Catalogue of Japanese Staphylinidae (Insecta: Coleoptera)rosti Bernhauer, 1923a: 127 (Mycetoporus) オオイクビハネカクシ（和名新称）HOK, HON; China, Russia
splendidum Gravenhorst, 1806: 24 (Tachinus) イクビハネカクシ（和名新称）HOK; China, Russia (Siberia, Far East),
India, Caucasus, Europe, N Africa, Nearctic Region
Genus Lordithon Thomson, 1859 キノコハネカクシ属
Subgenus Bobitobus Tottenham, 1939 ズナガキノコハネカクシ亜属（和名新称）
daimio Sharp, 1888: 456 (Bolitobius) ダイミョウキノコハネカクシ HOK, HON, SHI, KYU
elegantulus L.-Z. Li & Sakai, 1996b: 251 ツマグロアカキノコハネカクシ HON, SHI, KYU
hokkaidensis L.-Z. Li & Sakai, 1996b: 254 エゾキノコハネカクシ（和名新称）HOK
hosodai Katayama & T. Ito, 2010: 296 ホソダキノコハネカクシ（和名新称）HON
irregularis Weise, 1877a: 93 (Bolitobius) ハスオビキノコハネカクシ HON, SHI, Shd, KYU, Amm; China
niponensis Cameron, 1933: 170 (Bolitobius) チビクロモンキノコハネカクシ HOK, HON, SHI, KYU; China
ohbayashii L.-Z. Li & Zhao, 1999: 250 オオバヤシキノコハネカクシ（和名新称）HOK; Russia (Siberia, Far East)
principalis Sharp, 1888: 456 (Bolitobius) ハラオビキノコハネカクシ HOK, HON, SHI, Yaksemirufus Sharp, 1888: 457 (Bolitobius) クロモンキノコハネカクシ HOK, HON, SHI, KYU; China
takashii Katayama & T. Ito, 2010: 294 ムモンチャバネキノコハネカクシ（和名新称）HON, SHI, KYU
Subgenus Lordithon Thomson, 1859 キノコハネカクシ亜属（和名新称）
aitai Li & Ohbayashi, 1997: 95 ズグロキノコハネカクシ Tsmbicolor Gravenhorst, 1806: 28 (Tachinus) ハラグロキノコハネカクシ HOK, HON, SHI, KYU; South Korea, China,
Russia (Far East, Siberia), Europe= felix Sharp, 1888: 459 (Bolitobius)
cinctiventris Sharp, 1888: 458 (Bolitobius) ハネモンキノコハネカクシ（和名新称）HON, SHI, KYU
humeronotatus L.-Z. Li & Ohbayashi, 1999: 232 アカチャモンキノコハネカクシ（和名新称）HON, SHI
japonicus Sharp, 1874a: 20 (Bolitobius) ニホンキノコハネカクシ（和名新称）HON, SHI, KYU
lewisi Cameron, 1933: 171 (Bolitobius) ルイスキノコハネカクシ（和名新称）HON, Oki, SHI, KYU
limbifer Fauvel, 1901a: 3 (Bolitobius) ショウモンキノコハネカクシ（和名新称）HON, SHI, KYU= nigriventris Cameron, 1944b: 158 (Bolitobius)
optatus Sharp, 1888: 462 (Megacronus) モンナシキノコハネカクシ（和名新称）HON
pallidiceps Sharp, 1888: 459 (Bolitobius) ニセハネモンハネカクシ（和名新称）HON, SHI, KYU; Taiwan
sakaii L.-Z. Li & Ohbayashi, 1999: 239 サカイキノコハネカクシ（和名新称）HOK, HON, SHI, KYU
sharpianus Scheerpeltz, 1933: 1490 (Bolitobius) [RN] シャープキノコハネカクシ（和名新称）HON, SHI, KYU= sharpi Cameron, 1930: 208 (Bolitobius) [HN]simplex Sharp, 1888: 459 (Bolitobius) ムネアカキノコハネカクシ HON, SHI, KYU, Tsm; South Korea= melanurus Fauvel, 1901a: 4 (Bolitobius)
trimaculatus Fabricius, 1793: 533 (Oxyporus) ウスモンキノコハネカクシ HOK, HON, SHI, KYU; China, Russia(Siberia, Far East), Iran, Turkey, Europe= breviceps Sharp, 1888: 458 (Bolitobius)
Genus Mycetoporus Mannerheim, 1830 ニセイクビハネカクシ属（和名新称）
japonicus L.-Z. Li & Ohbayashi, 1996: 239 ニセイクビハネカクシ（和名新称）HOK, HON

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Yasutoshi Shibata, Munetoshi Maruyama, Hideto Hoshina, Toshio Kishimoto, Shun-Ichiro Naomi, Shûhei Nomura, Volker Puthz, Takashi Shimada, Yasuaki Watanabe, Shûhei Yamamoto

Genus Parabolitobius L.-Z. Li, M.-J. Zhao & Sakai, 2000 チャキノコハネカクシ属（和名新称）
prolongatus Sharp, 1888: 460 (Megacronus) アカチャキノコハネカクシ HOK, HON, SHI, KYU; China
yezoensis L.-Z. Li, Zhao & Sakai, 2000: 14 コゲチャキノコハネカクシ（和名新称）Reb; China, Russia
Tribe Tachyporini MacLeay, 1825 シリホソハネカクシ族
Genus Cilea Jacquelin du Val, 1856 ツヤヒメマルクビハネカクシ属（和名新称）
limbifera Motschulsky, 1858c: 222 (Erchomus) フタテンツヤヒメマルクビハネカクシ（和名改称）（註６）HON,
Iz-My, Iz-Hch, KYU, Tsm, Yak, T-Nak, T-Tak, Amm, Okn, Myk, Ish, Iri, Hat, Yon; Taiwan, India, Nepal,
Pakistan, Oriental Region
Genus Coproporus Kraatz, 1857 チビマルクビハネカクシ属（和名新称）
evanescens Boheman, 1858: 26 (Tachyporus) アカセマルチビマルクビハネカクシ（和名改称）Iz-Hch, Og-Cc, OgHh, Yak, Amm, Okn, Iri; China, Malaysia, Thailand= insulicola Y. Watanabe & Shibata, 1972: 66 (Paracilea)
micropennis Nakane & Sawada, 1960: 125 (Erchomus) ミナミチビマルクビハネカクシ KYU, Yak, T-Nak, RK;
Taiwanminimus Motschulsky, 1858c: 220 (Erchomus) チビマルクビハネカクシ（和名新称）Og; Taiwan, Hong Kong,
China, Philippines, Myanmar, Vietnam, Malaysia, Singapore, Indonesia (Sumatra), India, Sri Lanka, Seychelles
scitulus Weise, 1877a: 91 (Erchomus) クロチビマルクビハネカクシ HON, Iz-My, Iz-Mk, Iz-Hch, SHI, KYU; China,
Russia (Far East)
Genus Nitidotachinus Campbell, 1993 ヒゲナガマルクビハネカクシ属（和名新称）
adachii Y. Watanabe & Shibata, 1961b: 37 (Tachinus) クロヒゲナガマルクビハネカクシ HON, SHI, KYU
excellens excellens Bernhauer, 1938b: 23 (Tachinus) ヒゲナガマルクビハネカクシ HON, KYU; South Korea, China,
Russia (Far East)= sawadai Y. Watanabe & Shibata, 1961b: 36 (Tachinus)
impunctatus Sharp, 1888: 384 (Tachinus) ムネアカヒゲナガマルクビハネカクシ（和名改称）HOK, Reb, HON, SHI;
China
japonicus Schülke, 2000: 906 ヤマトヒゲナガマルクビハネカクシ（和名新称）HON
Genus Sepedophilus Gistel, 1856 ヒメキノコハネカクシ属
apicalis Naomi & Maruyama, 1997a: 251 ヘリグロヒメキノコハネカクシ（和名新称）HON, Okn
apterus Naomi & Maruyama, 1998: 69 フトムクゲヒメキノコハネカクシ（和名新称）HON, SHI
armatus Sharp, 1888: 455 (Conosoma) クロゲヒメキノコハネカクシ HOK, HON, Iz-My, Iz-Mk, Iz-Hch, SHI, KYU,
Yak, Tan, Okn; China
consimilis Naomi & Maruyama, 1998: 71 チャマルムクゲヒメキノコハネカクシ（和名新称）KYU
elegantissimus Naomi & Maruyama, 1997b: 68 フタマルモンヒメキノコハネカクシ（和名新称）Okn
exiguus Naomi & Maruyama, 1997b: 66 チビムナビロヒメキノコハネカクシ（和名新称）Ish; Taiwan
facilis facilis Naomi & Maruyama, 1998: 55 ムクゲヒメキノコハネカクシ（和名新称）HON, KYU

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Catalogue of Japanese Staphylinidae (Insecta: Coleoptera)f. tsushimanis Naomi & Maruyama, 1998: 60 Tsm
fimbriatus Sharp, 1888: 454 (Conosoma) オオヒメキノコハネカクシ HOK, HON, SHI, KYU, Yak; South Korea,
Taiwangermanus Sharp, 1874a: 17 (Conurus) ムクゲヒメキノコハネカクシ HOK, HON, Iz-O, Iz-Mk, Awj, Oki, SHI, KYU,
Tsm, SKs, Yak; China= rufipennis Adachi, 1937: 58 (Conosoma)
glabratus glabratus Naomi & Maruyama, 1997b: 62 ムナビロフタモンヒメキノコハネカクシ（和名新称）HON, KYU
g. okinawanis Naomi & Maruyama, 1997b: 65 Amm, Okn
hirsutus Naomi & Maruyama, 1998: 60 チャイロムクゲヒメキノコハネカクシ（和名新称）HON, KYU
humeralis Eppelsheim, 1886: 36 (Conurus) ネアカヒメキノコハネカクシ（和名新称）HOK; Taiwan, China, Russia(Far East)
insulicola Naomi & Maruyama, 1998: 61 ツヤチャイロムクゲヒメキムクゲノコハネカクシ（和名新称）Okn
iriomotensis Naomi & Maruyama, 1997a: 253 ヨツグロモンヒメキノコハネカクシ（和名新称）Ish, Iriishiiellus Naomi & Maruyama, 1998: 64 イシイムクゲヒメキノコハネカクシ（和名新称）HOK, HON
kobensis Cameron, 1933: 171 (Conosoma) カタモンヒメキノコハネカクシ（和名新称）HON, Iz-Mk, Iz-My
lewisi Cameron, 1930: 208 (Conosoma) ルイスヒメキノコハネカクシ（和名新称）SHI, Amm
littoreus Linnaeus, 1758: 422 (Staphylinus) ハネナガヒラタヒメキノコハネカクシ（和名新称）HOK, HON; China,
Iran, Russia (Siberia), Caucasus, Europe, USAlongipennis Cameron, 1930: 208 (Conosoma) クロヒゲヒメキノコハネカクシ（和名新称）HON
nomurai Naomi & Maruyama, 1998: 62 ノムラムクゲヒメキノコハネカクシ（和名新称）HON, SHI
planipennis Nakane, 1964b: 245 ヒラタヒメキノコハネカクシ HOK
pumilus Sharp, 1874a: 18 (Conurus) ハスモンヒメキノコハネカクシ HOK, HON, Iz-Mk, Iz-Hch, SHI, KYU, SKs,
Tan, Yak, Amm
quadrifurcatus Naomi & Maruyama, 1997a: 249 ムナグロヒメキノコハネカクシ（和名新称）Amm, Okn
satoshii Naomi & Maruyama, 1998: 68 イトウムクゲヒメキノコハネカクシ（和名新称）KYU
simulans Nakane & Sawada, 1960: 124 ヤクシマヒメキノコハネカクシ KYU, Tsm, Yak; South Korea
subarmatus Naomi & Maruyama, 1997a: 246 ニセクロゲヒメキノコハネカクシ（和名新称）HON, Kue; South Korea
tibialis Sharp, 1888: 454 (Conosoma) ヒメキノコハネカクシ HOK, HON, SHI, KYU, Yak; South Korea
tristiculus Weise, 1877a: 92 (Conosoma) ツヤヒメキノコハネカクシ（和名新称）HON, SHI, Iz-My, Iz-Mk; Russia(Siberia, Far East)
varicornis Sharp, 1888: 455 (Conosoma) クロヒメキノコハネカクシ HOK, HON, Iz-Mk, Iz-My, Oki, SHI, KYU,
SKs, Yak, Amm; South Korea, China
wankowiczi Pandellé, 1869: 294 (Conurus) フタモンヒメキノコハネカクシ（和名交換）（註７）HOK, HON, SHI,KYU; China, Russia (Siberia, Far East), Belarus, Latvia
Genus Tachinus Gravenhorst, 1802 マルクビハネカクシ属
Subgenus Japanotachinus Ullrich, 1975 ニホンマルクビハネカクシ亜属（和名新称）
nakanei Ullrich, 1975: 288 ヒゲブトマルクビハネカクシ HOK, HON, Oki, SHI, KYU
Subgenus Latotachinus Ullrich, 1975 ヒゲフトナガマルクビハネカクシ亜属（和名新称）
punctiventris Sharp, 1888: 385 クロツヤマルクビハネカクシ HOK, HON, SHI, KYU

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Yasutoshi Shibata, Munetoshi Maruyama, Hideto Hoshina, Toshio Kishimoto, Shun-Ichiro Naomi, Shûhei Nomura, Volker Puthz, Takashi Shimada, Yasuaki Watanabe, Shûhei Yamamoto

strigiventris Bernhauer, 1907: 387 ウスイロマルクビハネカクシ（和名新称）HON
Subgenus Tachinoderus Motschulsky, 1858 コマルクビハネカクシ亜属（和名新称）
diminutus Sharp, 1888: 385 コクロマルクビハネカクシ HON, Iz-Mk, SHI, KYU
iriomotensis L.-Z. Li, 1994: 664 イリオモテマルクビハネカクシ（和名新称）Ish, Iri; Taiwan
kobensis Cameron, 1933: 172 ムナグロマルクビハネカクシ（和名新称）HON, Oki, SHI, KYU, Yaknaomii L.-Z. Li, 1994: 661 ナオミマルクビハネカクシ（和名新称）Amm; Taiwan, China
nigriceps nigriceps Sharp, 1888: 384 クロズマルクビハネカクシ HOK, HON, SHI, KYU, Tsm, Yak; South Korea,
China, Russia
yamato Hayashi, 2003b: 134 (Paratachinus) ヒメマルクビハネカクシ（和名新称）HON
Subgenus Tachinus Gravenhorst, 1802 マルクビハネカクシ亜属（和名新称）
bipustulatus Fabricius, 1793: 533 (Oxyporus) チビヒゲマルクビハネカクシ（和名新称）HOK, HON, SHI; China,
Russia (Siberia, Far East), Kazakhstan, Europe= nigrinus Sharp, 1888: 382
elongatus Gyllenhal, 1810: 251 ナガマルクビハネカクシ HOK, HON; China, N Kuril, Russia (Siberia, Far East),
Mongolia, Caucasus, Europe, USA (Alaska)= kobai Kano, 1933: 99 (Mycetoporus)
gelidus Eppelsheim, 1893: 41 アカバマルクビハネカクシ HOK, HON, SHI, KYU; South Korea, China, Russia(Siberia, Far East), Mongolia= luridus Sharp, 1888: 381 [HN]
= sharpi Bernhauer & Schubert, 1916: 486 [RN]
japonicus Sharp, 1888: 379 ヤマトマルクビハネカクシ HON, Sad, Oki, SHI, KYU; Taiwan, China
javanus Cameron, 1937: 35 シバタマルクビハネカクシ（和名新称）HON, SHI; China, Russia (Siberia, Far East),
India, SE Asia
= shibatai Hayashi, 1987: 13
kinoshitai L.-Z. Li, 1995: 60 キノシタマルクビハネカクシ（和名新称）HON, SHI
mimulus Sharp, 1874a: 16 キベリマルクビハネカクシ HOK, HON, Sad, Iz-Mk, SHI, KYU
obesus Weise, 1877a: 92 ニセヤマトマルクビハネカクシ（和名新称）HON, SHI, KYU= kasugensis T. Ito, 1993a: 63
pallipes rishirianus Y. Watanabe & Shibata, 1965b: 322 キタマルクビハネカクシ HOK, Rsr
sibiricus Sharp, 1888: 383 カタアカマルクビハネカクシ HOK, HON, SHI, KYU; South Korea, North Korea, China,
Mongolia, Russia (Siberia, Far East), C Europe= bidens Sharp, 1888: 380
trifidus Sharp, 1888: 380 ネアカマルクビハネカクシ HON, SHI, KYU
yezoensis L.-Z. Li, 1995: 60 エゾマルクビハネカクシ（和名新称）HOK; South Korea
Incertae sedis (subgenus?) 亜属不明
sanguinithorax Bernhauer, 1939b: 157 キュウシュウマルクビハネカクシ（和名新称）KYU
Genus Tachyporus Gravenhorst, 1802 シリホソハネカクシ属
abdominalis Fabricius, 1781: 501 (Oxyporus) チャイロシリホソハネカクシ（和名新称）HON; China, Russia(Siberia, Sakhalin), Mongolia, Caucasus, Europe, USA

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Catalogue of Japanese Staphylinidae (Insecta: Coleoptera)celatoides Schülke, 1998: 382 ニセクロズシリホソハネカクシ（和名新称）HON
celatus Sharp, 1874a: 17 クロズシリホソハネカクシ HOK, HON, Iz-My, SHI, KYU, Yak; South Korea, China,
Russia (Siberia, Far East), W Asia
cephalotes Schülke, 1998: 377 ザウターシリホソハネカクシ（和名新称）HON
chrysomelinus Linnaeus, 1758: 423 (Staphylinus) シリホソハネカクシ（和名新称）HOK; China, Russia (Siberia,
Far East), Mongolia, C Asia, Europe, USA
dispar Paykull, 1789: 77 (Staphylinus) クロズムネチャシリホソハネカクシ（和名新称）HON; Europe, USA= immaculatus Luze, 1901: 169
frontalis Cameron, 1933: 171 セッツシリホソハネカクシ（和名新称）HON, SHI
japonicus Schülke, 1998: 374 ヤマトシリホソハネカクシ（和名新称）HON, SHI
oculatus Sharp, 1888: 452 ズビロシリホソハネカクシ（和名新称）HON, KYU
orthogrammus Sharp, 1888: 453 キベリシリホソハネカクシ HON, SHI, KYU; China
picturatus Reitter, 1913: 121 (Leucoparyphus) ハネモンシリホソハネカクシ（和名新称）HOK, HON; Taiwan,
China, Russia (Far East)
pusillus Gravenhorst, 1806: 9 ハバビロシリホソハネカクシ（和名新称）SHI; China, Russia (Siberia, Far East), India,
Syria, Turkey, Europe, N Africa
suavis Sharp, 1888: 452 ツヤグロシリホソハネカクシ HON, SHI, KYU; South Korea
terminalis Sharp, 1888: 451 キイロシリホソハネカクシ（和名新称）HOK, HON, KYU; China, Russia (Siberia,
Sakhalin)註 ６：これまでフタテンヒメマルクビハネカクシ Cilea silphoides (Linnaeus, 1767)として日本から記録されてきたが，実際は S. limbifera (Motschulsky, 1858)とするのが妥当のようである(Hayashi, 2002)．標本の再調査といった課題が残るものの，ここでは後者として扱っておく．
註 ７：これまで日本から報告されたフタモンヒメキノコハネカクシ Sepedophilus bipustulatus (Gravenhorst, 1802) とされてきた種は，S. wankowiczi (Pandellé, 1869)と置き換えるのが妥当のようである(Yamamoto, submitted)．和名は混乱を与えないよう，従来通り，フタモンヒメキノコハネカクシを踏襲する．また，他に本属種として S.
testaceus (Fabricius, 1793)も記録されているが(Horion, 1967)，疑わしいので本目録からは除外する．

Subfamily Trichophyinae Thomson, 1859 ホソヒゲハネカクシ亜科（１属３種）
Genus Trichophya Mannerheim, 1830 ホソヒゲハネカクシ属
japonica Y. Watanabe & Shibata, 1962: 95 ニホンホソヒゲハネカクシ HOK, HON
punctata Naomi, 1996b: 71 ナガホソヒゲハネカクシ HOK
uenoi Naomi, 1995d: 347 ウエノホソヒゲハネカクシ HOK

Subfamily Aleocharinae Fleming, 1821 ヒゲブトハネカクシ亜科（102属357種１亜種）

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Yasutoshi Shibata, Munetoshi Maruyama, Hideto Hoshina, Toshio Kishimoto, Shun-Ichiro Naomi, Shûhei Nomura, Volker Puthz, Takashi Shimada, Yasuaki Watanabe, Shûhei Yamamoto

Tribe Aleocharini Fleming, 1821 ヒゲブトハネカクシ族
Subtribe Aleocharina Fleming, 1821 ヒゲブトハネカクシ亜族
Genus Aleochara Gravenhorst, 1802 ヒゲブトハネカクシ属
Subgenus Aleochara Gravenhorst, 1802
clavigera Sharp, 1874a: 7 オオヒゲブトハネカクシ（和名新称）Japan; South Korea, China
coreana Bernhauer, 1926: 271 マルアカヒゲブトハネカクシ（和名新称）Japan; North Korea, South Korea= lewisia Bernhauer, 1936: 324
curtula Goeze, 1777: 730 (Staphylinus) ナカアカヒゲブトハネカクシ HOK, HON, Sad, Oki, SHI, KYU, Tsm;
Holoarctic Region, S America= discoidea Sharp, 1874a: 7
kochi Bernhauer, 1941: 211 コッホヒゲブトハネカクシ（和名新称）HON; South Korea
lata Gravenhorst, 1802: 186 ヒゲブトハネカクシ HOK, HON, KYU; South Korea, North Korea, China, Russia,
Holoarctic Region
parens Sharp, 1874a: 6 コクロヒゲブトハネカクシ HOK, HON, Iz-Iss, KIw, SHI, KYU, Yak; South Korea, North
Korea, China
Subgenus Baryodma Thomson, 1858
niponensis Sharp, 1888: 281 ニッポンヒゲブトハネカクシ（和名新称）HON, KYU 種復活予定
Subgenus Coprochara Mulsant & Rey, 1874
bipustulata Linnaeus, 1760: 232 (Staphylinus) 日本に分布しない (Yamamoto & Maruyama, submitted)
squalithorax Sharp, 1888: 282 フトツヤケシヒゲブトハネカクシ HOK, HON; South Korea
verna Say, 1833 フタモンヒゲブトハネカクシ( 和名交換予定) HOK, HON, SHI, KYU; Holoarctic Region
Subgenus Emplenota Casey, 1884
fucicola Sharp, 1874a: 9 ツヤケシヒゲブトハネカクシ HON, Iz-Hch, Awj, SHI, KYU, Tsm, Amk, KKs; South
Korea, North Korea, Hong Kong, China, France= variolosa Weise, 1877a: 89 (Homalota)
hayamai Yamamoto & Maruyama, 2012: 23 ハヤマツヤケシヒゲブトハネカクシ HON, KYU, Oki, KKs
puetzi Assing, 1995: 220 (Emplenota) キタツヤケシヒゲブトハネカクシ HOK, Oks; North Korea, Russia (Chs,
Sakhalin, Primorsky, Kamchatka)
segregata Yamamoto & Maruyama, 2012: 20 ニセツヤケシヒゲブトハネカクシ HOK, Oks, HON, Sad, Awj, SHI,KYU, Tan
yamato Yamamoto & Maruyama, 2012: 26 オオツヤケシヒゲブトハネカクシ HON
Subgenus Maseochara Sharp 1883
hiranoi Yamamoto & Maruyama, 2009: 65 キノカワヒゲブトハネカクシ HON
Subgenus Triochara Bernhauer, 1901
nubis Assing, 1995: 230 (Triochara) エゾセスジヒゲブトハネカクシ HOK; Russia (Chs, Sakhalin, Kamchatka)
trisulcata Weise, 1877a: 88 ホソセスジヒゲブトハネカクシ HON, Sad, SHI, KYU, Tsm, SKs, Tan, Amm, Tok, Oke;South Korea, Hong Kong, China
zerchei Assing, 1995: 231 (Triochara) ニセセスジヒゲブトハネカクシ HOK, HON, Awj, SHI, KYU; South Korea,
North Korea, Russia (Chs, Sakhalin, Primorsky, Kamchatka)2013 The Kyushu

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日本産ハネカクシ科総目録（昆虫綱：甲虫目）
Catalogue of Japanese Staphylinidae (Insecta: Coleoptera)

Subgenus Xenochara Mulsant & Rey, 1874
asiatica Kraatz, 1859:15 アジアヒゲブトハネカクシ（和名新称）HON, KYU; China, India, Nepal, Oriental Region= japonica Sharp, 1874a: 8
fumata Gravenhorst, 1802: 96 イブシヒゲブトハネカクシ（和名新称）HON; China, Russia, Holoarctic Region
lygaea Kraatz, 1862: 317 クロホソヒゲブトハネカクシ（和名新称）Japan; China, Russia
peninsulae Bernhauer, 1936: 325 シマバラヒゲブトハネカクシ（和名新称）KYU; South Korea
peregrina Sharp, 1874a: 9 イコクヒゲブトハネカクシ（和名新称）Japanpraesul Sharp, 1874a: 9 セナガヒゲブトハネカクシ HON, Awj; China, Russia
puberula Klug, 1832: 139 ウスアカヒゲブトハネカクシ HOK, HON, Tsm, Yak, Yor, Okn, KDt, MDt; Cosmopolitan
Genus Creochara Cameron, 1939 ショウリョウヒゲブトハネカクシ属
brevipennis Bernhauer, 1903: 21 (Myrmedonia) ショウリョウヒゲブトハネカクシ HON, SHI, KYU, Tsm, Yak, Amm,
Okn; Taiwan, Vietnam, Indonesia, India
Genus Plesiochara Sawada, 1989 クサビラハネカクシ属（和名新称）
fusca Sawada, 1989b: 304 クサビラハネカクシ（和名新称）HON
nitida Sawada, 1990: 551 ツヤクサビラハネカクシ（和名新称）HON
Genus Tinotus Sharp, 1883 チビヒゲブトハネカクシ属（和名新称）
japonicus Cameron, 1933: 217 チビヒゲブトハネカクシ（和名改称）HON
Genus Tetrasticta Kraatz, 1857 メダカヒゲブトハネカクシ属
laeta Maruyama & Sugaya, 2002: 17 メダカヒゲブトハネカクシ Iri; Taiwan
Tribe Athetini Casey, 1910 ヒメハネカクシ族
Subtribe Athetina Fleming, 1821 ヒメハネカクシ亜族
Genus Acrotona Thomson, 1859 ホソヒメハネカクシ属（和名新称）
grata Cameron, 1933: 216 (Atheta) ウレシヒメハネカクシ（和名新称）HOK, HON, SHI, KYU; Taiwan, North
Korea, Russia
inornata Kraatz, 1859: 39 (Homalota) オボコヒメハネカクシ（ 和 名 新 称 ）HON, KYU; India, China, Oriental
Region= taedia Cameron, 1933: 215
lutulenta Sharp, 1888: 293 (Homalota) オワイヒメハネカクシ（和名新称）HON, SHI, KYU: North Korea
neglecta Cameron, 1933: 215 (Atheta) キヅカヌヒメハネカクシ（和名新称）KYU; China
pseudotenera Cameron, 1933: 216 (Atheta) ムコヒメハネカクシ（和名新称）HOK, HON, SHI, KYU; Europe
vivida Sharp, 1874a: 15 (Homalota) ゲンキヒメハネカクシ（和名新称）HON, KYU; North Korea
yosii Sawada, 1970a: 53 (Ischnopoda) ヨシイヒメハネカクシ（和名新称）HON
Genus Adota Casey, 1910 ウシオヒメハネカクシ属（和名新称）2013 The Kyushu

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Yasutoshi Shibata, Munetoshi Maruyama, Hideto Hoshina, Toshio Kishimoto, Shun-Ichiro Naomi, Shûhei Nomura, Volker Puthz, Takashi Shimada, Yasuaki Watanabe, Shûhei Yamamoto

madida Bernhauer, 1907: 400 (Atheta) カマクラウシオヒメハネカクシ（和名新称）HON, KYU
magnipennis Bernhauer, 1943: 184 (Atheta) ウシオヒメハネカクシ 学名変更予定 HON, KYU
maritima Mannerheim, 1843: 224 (Homalota) オオウシオヒメハネカクシ（和名新称）Japanushio Sawada, 1971: 304 (Ischnopoda) HON, KYU 下位同物異名予定
vacillator Cameron, 1933: 211 (Atheta) ヨロメキウシオヒメハネカクシ（和名新称）HON
Genus Alevonota Thomson, 1858 ハラナガヒメハネカクシ属（和名新称）
japonica Cameron, 1933: 218 ハラナガヒメハネカクシ（和名新称）HON
Genus Aloconota Thomson, 1858ドウナガヒメハネカクシ属（和名新称）
Subgenus Aloconota Thomson, 1858
bulbosa Sawada, 1989b: 299 カイドウナガヒメハネカクシ（和名新称）HON
cuspidata Sawada, 1971b: 294 (Tomoglossa) セトドウナガヒメハネカクシ（和名新称）HON
punctifoveata Sawada, 1970a: 44 (Tomoglossa) シナノドウナガヒメハネカクシ（和名新称）HON
Genus Amischa Thomson, 1858 ズンドウヒメハネカクシ属（和名新称）
Subgenus Amischa Thomson, 1858
analis Gravenhorst, 1802: 76 (Aleochara) ズンドウヒメハネカクシ（和名新称）HOK; Russia, Iran, Kazakhstan,
Uzbekistan, Europe, USA
niponensis Sharp, 1888: 292 (Homalota) ニッポンズンドウヒメハネカクシ（和名新称）KYU
Genus Atheta Thomson, 1858 ヒメハネカクシ属
Subgenus Anopleta Mulsant & Rey, 1874
okamotoi Sawada, 1989b: 294 オカモトヒメハネカクシ（和名新称）HON
Subgenus Atheta Thomson, 1858
allocera Eppelsheim, 1893: 34 エッペルシェイムヒメハネカクシ（和名新称）HON; Russia, Europe= allocera ontakeana Sawada, 1989b: 288
brunneipennis Thomson, 1852: 135 (Homalota) チャバネヒメハネカクシ（和名新称）HOK; Russia, Europe
coriaria Kraatz, 1856: 282 (Homalota) ナメシヒメハネカクシ（和名新称）HON; North Korea, China, Europe= academica Sawada, 1976: 125 (Taxicera)crassa Sawada, 1990: 545 オオヒメハネカクシ（和名新称）HON
euryptera Stephens, 1832: 135 (Aleochara) フトハネヒメハネカクシ（和名新称）HON, Yak; China, Russia, Europe= euryptera japonica Bernhauer, 1907: 407
sauteri Bernhauer, 1907: 407 ザウターヒメハネカクシ（和名新称）HON, SHI, KYU; Taiwan, China, North Korea,
Nepal, India= reitteriana Bernhauer, 1939a: 109
separata Sawada, 1970a: 47 (Ischnopoda) サワダヒメハネカクシ（和名新称）HON
settsuensis Cameron, 1933: 213 セッツヒメハネカクシ（和名新称）HON
transfuga Sharp, 1874a: 13 (Homalota) クボタヒメハネカクシ（和名新称）HON, SHI, KYU= kubotai Bernhauer, 1943: 185

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日本産ハネカクシ科総目録（昆虫綱：甲虫目）
Catalogue of Japanese Staphylinidae (Insecta: Coleoptera)

Subgenus Badura Mulsant & Rey, 1873
nana Kraatz, 1859: 36 (Homalota) コビトヒメハネカクシ（和名新称）HON; Taiwan, China, North Korea, India,
Oriental Region= kanagawana Bernhauer, 1907: 399
ririkoae Sawada, 1989b: 285 リリコヒメハネカクシ（和名新称）HON
tokiokai Sawada, 1971b: 306 (Ischnopoda)トキオカヒメハネカクシ HOK, HON, KYU
Subgenus Bessobia Thomson, 1858
occulta Erichson, 1837: 393 (Homalota) エリクソンヒメハネカクシ（和名新称）HON; China, Russia, Europe= erichsoni Bernhauer, 1907: 393
Subgenus Ceritaxa Mulsant & Rey, 1873
kanagawaensis Scheerpeltz, 1960: 614 カナガワヒメハネカクシ（和名新称）HON
Subgenus Chaetida Mulsant & Rey, 1874
subantennata Cameron, 1933: 214 ナガサキヒメハネカクシ（和名新称）KYU
subasperata Bernhauer, 1907: 399 ヤマナカヒメハネカクシ（和名新称）HON
Subgenus Coprothassa Thomson, 1859
melanaria Mannerheim, 1830: 70 (Bolitochara) クロヒメハネカクシ（和名新称）HOK; China, Russia, Europe
Subgenus Datomicra Mulsant & Rey, 1874
dadopora Thomson, 1867: 379 エゾヒメハネカクシ（和名新称）HOK; China, India, Europe= poroshirica Sawada, 1978: 243
gallina Sawada, 1989a: 1 メンドリヒメハネカクシ（和名新称）HOK
Subgenus Datostiba Sawada, 1976
denticauda Bernhauer, 1907: 401 シリバヒメハネカクシ（和名新称）HON, KYU; China
dentiventris Bernhauer, 1907: 412 HON 下位同物異名予定
lewisiana Cameron, 1933: 214 ルイスヒメハネカクシ（和名新称）HOK, HON, SHI, KYU; North Korea, China,
Oriental Region
nikkoana Sawada, 1978: 241 ニッコウヒメハネカクシ（和名新称）HON
Subgenus Dimetrota Mulsant & Rey, 1873
aeneipennis Thomson, 1856: 103 (Homalota) ツヤバネヒメハネカクシ（和名新称）HOK, HON, SHI; North Korea,
Russia= tenuidacta Sawada, 1970: 52
atramentaria Gyllenhall, 1810: 408 (Aleochara) スミイロヒメハネカクシ（和名新称）HOK, HON, KYU; Palearctic
Region, Africa
nikkoensis Sawada, 1989b: 290トチギヒメハネカクシ（和名新称）HON
nobukii Sawada, 1989b: 4 ダイセツヒメハネカクシ（和名新称）HOK
shigae Sawada, 1976: 57 (Cadaverota) シガヒメハネカクシ（和名新称）HON
weisei Bernhauer, 1907: 401 ズグロアカヒメハネカクシ（和名改称）HOK, HON, Tsm; North Korea, China= multispina Sawada, 1970a: 51
yamamotoi Sawada, 1989b: 292 ヤマモトヒメハネカクシ（和名新称）SHI
Subgenus Eubadura Sawada, 1990
akiensis Sawada, 1990: 543 アキヒメハネカクシ（和名新称）HON

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Yasutoshi Shibata, Munetoshi Maruyama, Hideto Hoshina, Toshio Kishimoto, Shun-Ichiro Naomi, Shûhei Nomura, Volker Puthz, Takashi Shimada, Yasuaki Watanabe, Shûhei Yamamoto

Subgenus Microdota Mulsant & Rey, 1873
formicetorum Bernhauer, 1907: 400 アリスヒメハネカクシ（和名新称）HON; North Korea
inari Sawada, 1978: 246 イナリヒメハネカクシ（和名新称）HON
kawachiensis Sawada, 1974: 158 カワチヒメハネカクシ（和名新称）HON
kobensis Cameron, 1933: 212 コウベヒメハネカクシ（和名新称）HOK, HON; North Korea
koreana Bernhauer, 1923a: 128 コウライヒメハネカクシ（和名新称）Japan; North Korea, China= bulbosa Sawada, 1974: 179
muris Sawada, 1974: 176 コネズミヒメハネカクシ（和名新称）HON, SHI, KYU
nakanei Sawada, 1970b: 46 (Ischnopoda) ナカネヒメハネカクシ（和名新称）HON
nopporensis Sawada, 1974: 160 ノッポロヒメハネカクシ（和名新称）HOK
ontakensis Sawada, 1974: 155 オンタケヒメハネカクシ（和名新称）HON
oviformis Sawada, 1970a: 61 (Ischnopoda) タマゴヒメハネカクシ（和名新称）HON
puthzi Sawada, 1974: 170 プーツヒメハネカクシ（和名新称）HON
silvatica Bernhauer, 1907: 405 モリヒメハネカクシ（和名新称）HON; North Korea
spiniventris Bernhauer, 1907: 402 ハラトゲヒメハネカクシ（和名新称）HON
spinicauda Bernhauer, 1907: 404 シリトゲヒメハネカクシ（和名新称）HON 種復活予定
spinula Sawada, 1970a: 60 コトゲヒメハネカクシ（和名新称）HON
subcrenulata Bernhauer, 1907: 403 ナミバラヒメハネカクシ（和名新称）HOK, HON; North Korea, China
sublaevigata Bernhauer, 1907: 405 ミジンヒメハネカクシ（和名新称）HON
uncinata Sawada, 1971a: 80 (Ischnopoda) アキヨシヒメハネカクシ（和名新称）HON
unidentata Sawada, 1971a: 82 (Ischnopoda) ヒトバヒメハネカクシ（和名新称）HON; Oriental Region
vagans Bernhauer, 1907: 404 ホウロウヒメハネカクシ（和名新称）HON, KYU; Taiwan, China= bichuensis Cameron, 1933: 212
vulpina Sawada, 1974: 170 キツネヒメハネカクシ（和名新称）HON
yoshidai Sawada, 1974: 173 ヨシダヒメハネカクシ（和名新称）SHI
Subgenus Orientatheta Sawada, 1985
chinkiangensis Bernhauer, 1939b: 145トウアヒメハネカクシ（和名新称）KYU; North Korea, China
Subgenus Paradota Sawada, 1989
liberta Sawada, 1989a: 274 ジユウヒメハネカクシ（和名新称）HON
Subgenus Philhygra Mulsant & Rey, 1873
palustris Kiesenwetter, 1844: 318 ヌマチヒメハネカクシ（ 和 名 新 称 ）HON; Korea, China, Russia, Mongolia,
Europe, USA
pseudoelongatula Bernhauer, 1907: 411 オカヤマヒメハネカクシ（和名新称）HON; South Korea, China
sparsa Bernhauer, 1907: 410 ウスヒメハネカクシ（和名新称）HOK, HON, SHI; Korea= lucidula Cameron, 1933: 211
yokkaichiana Bernhauer, 1907: 410 ヨッカイチヒメハネカクシ（和名新称）HON, KYU; Korea, China= unzensis Cameron, 1933: 211
Subgenus Suensonia Bernhauer, 1936
obsoletepunctata Bernhauer, 1936: 314 セーンソンヒメハネカクシ（和名新称）KYU
Subgenus Tetropa Mulsant & Rey, 1874

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Catalogue of Japanese Staphylinidae (Insecta: Coleoptera)neolata Sawada, 1976: 31 [RN] サワダヒメハネカクシ（和名新称）HON= prolata Sawada, 1970a: 63 (Ischnopoda) [HN]
Genus Caenogluta Sawada, 1977ドロヒメハネカクシ属（和名新称）
ocyusina Bernhauer, 1907: 406 (Atheta)ドロヒメハネカクシ（和名新称）HON
Genus Geostiba Thomson, 1858 コバネヒメハネカクシ属（和名新称）
circellaris Gravenhorst, 1806: 155 (Aleochara) コバネヒメハネカクシ（和名改称）HOK; North Korea, Russia,
Turkey, Europe, USAdaisetsuana Sawada, 1989a: 7 ダイセツコバネヒメハネカクシ（和名新称）HOK
Genus Hydrosmecta Thomson, 1858 ミズギワヒメハネカクシ属（和名新称）
longula Heer, 1839: 334 (Homalota) ミズギワヒメハネカクシ属（和名新称）HON; Europe= thinobioides Kraatz, 1854: 125 (Homalota)
Genus Liogluta Thomson, 1858 ツヤヒメハネカクシ属（和名新称）
constricta Sawada, 1970a: 48 (Ischnopoda) クビレツヤヒメハネカクシ（和名新称）HOK, HON
distans Sharp, 1874a: 14 (Homalota) ハルカツヤヒメハネカクシ（和名新称）HOK, HON
iturupensis Bernhauer, 1907: 408 (Atheta) エトロフツヤヒメハネカクシ（和名新称）HOK, Chs
libraria Sawada, 1989b: 297 ホンツヤヒメハネカクシ（和名新称）HON
ursi Sawada, 1972: 53 (Atheta) ウルシツヤヒメハネカクシ（和名新称）HOK
yasudai Sawada, 1989a: 9 ヤスダツヤヒメハネカクシ（和名新称）HOK
yezoensis Sawada, 1989a: 12 エゾツヤヒメハネカクシ（和名新称）HOK
Genus Mimatheta Cameron, 1920 マガイヒメハネカクシ（和名新称）
nipponensis Cameron, 1933: 217 マガイヒメハネカクシ（和名新称）HON
Genus Nehemitropia Lohse, 1971 ツヤケシヒメハネカクシ属（和名新称）
lividipennis Mannerheim, 1830: 79 (Oxypoda) ツヤケシヒメハネカクシ（ 和 名 改 称 ）HOK, HON, SHI, KYU;
Taiwan, North Korea, China, Europe= sordida Marsham, 1802: 513 (Staphylinus)
milu Likovský, 1977: 209 ミルツヤケシヒメハネカクシ（和名改称）Japan; China
Genus Neodinaraea Pace, 1983 コブヒメハネカクシ属（和名新称）
tuberculata Bernhauer, 1907: 391 (Leptusa) コブヒメハネカクシ（和名新称）HON
Genus Nepalota Pace, 1987 オニヒメハネカクシ属（和名新称）
angusticavata Assing, 2003a: 155 オニヒメハネカクシ（和名新称）HON, SHI
kyushuica Assing, 2003a: 157 キュウシュウオニヒメハネカクシ（和名新称）KYU
laticavata Assing, 2003a: 159 ナラオニヒメハネカクシ（和名新称）HON

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Yasutoshi Shibata, Munetoshi Maruyama, Hideto Hoshina, Toshio Kishimoto, Shun-Ichiro Naomi, Shûhei Nomura, Volker Puthz, Takashi Shimada, Yasuaki Watanabe, Shûhei Yamamoto

naomii Assing, 2003a: 161 ナオミオニヒメハネカクシ（和名新称）SHI
rectisulcata Assing, 2003a: 159 ヤマナシオニヒメハネカクシ（和名新称）HON
Genus Nikkostiba Pace, 1984 ニッコウヒメハネカクシ属（和名新称）
loebli Pace, 1984: 899 ニッコウヒメハネカクシ（和名新称）HON
Genus Notothecta Thomson, 1858 ヒロムネヒメハネカクシ属（和名新称）
effecta Sawada, 1989b: 276 (Atheta) ヒロムネヒメハネカクシ（和名新称）HON
incola Sawada, 1989b: 276 (Atheta) キヨタキヒロムネヒメハネカクシ（和名新称）HON
kashimaensis Sawada, 1989b: 281 (Atheta) カシマヒロムネヒメハネカクシ（和名新称）HON
longisetosa Sawada, 1970a: 50 (Atheta) ケナガヒロムネヒメハネカクシ（和名新称）HON
watanabei Sawada, 1989b: 283 (Atheta) ワタナベヒロムネヒメハネカクシ（和名新称）HON
Genus Pelioptera Kraatz, 1857 ハバムネヒメハネカクシ属（和名新称）
Subgenus Pelioptera Kraatz, 1857
exasperata Kraatz, 1859: 32 (Homalota) アラハダハバムネヒメハネカクシ（和名新称）Japan; India, SE Asia
micans Kraatz, 1857b: 56 ツヤハバムネヒメハネカクシ（和名新称）Japan; China, India, Oriental Region
testaceipennis Motschulsky, 1858c: 251 (Homalota) キバネハバムネハネカクシ RK; China, SE Asia
= luchuensis Cameron, 1933: 213
Incertae sedis (subgenus?) 亜属不明
babai Sawada, 1989b: 301 ババハバムネヒメハネカクシ（和名新称）HON
flavonitescens Bernhauer, 1939a: 108 (Atheta) キュウシュウハバムネヒメハネカクシ（和名新称）KYU
ocyamensis Bernhauer, 1914b: 39 (Atheta) オカヤマハバムネヒメハネカクシ（和名新称）HON
Genus Plataraea Thomson, 1858 コガシラヒメハネカクシ属（和名新称）
Subgenus Plataraea Thomson, 1858
brunnea Fabricius, 1798: 180 (Staphylinus) ウスアカコガシラヒメハネカクシ（和名新称）Japan; Europe
punctifrons Sawada, 1970a: 56 (Ischnopoda) コガシラヒメハネカクシ（和名新称）HON
Genus Psammostiba Sawada, 1976 ヒゲナガヒメハネカクシ属
hilleri Weise, 1877a: 90 (Homalota) ヒゲナガヒメハネカクシ HON, KYU= multipunctata Sawada, 1971b: 301
jessoensis Brundin, 1943: 22 (Atheta) エゾヒゲナガヒメハネカクシ HOK, HON; North Korea, Russia
kamtschatica Brundin, 1943: 21 (Atheta) ホッポウヒゲナガヒメハネカクシ HOK, HON; Russia
Genus Salinamexus Moore & Legner, 1977 ハマハネカクシ属
browni Moore & Legner, 1977: 464 ハマハネカクシ HON; Mexico
hayamai Maruyama, 2011b: 108 ヒメハマハネカクシ HON
koreanus Jeon & Ahn, 2007: 193 チョウセンハマハネカクシ HON; Korea

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日本産ハネカクシ科総目録（昆虫綱：甲虫目）
Catalogue of Japanese Staphylinidae (Insecta: Coleoptera)

Genus Saphocallus Sharp, 1888 カクバラハネカクシ属（和名新称）
parviceps Sharp, 1888: 288 カクバラハネカクシ（和名新称）HON, KYU
Genus Schistoglossa Kraatz, 1856 マルムネヒメハネカクシ属（和名新称）
yosiiana Sawada, 1970a: 44 マルムネヒメハネカクシ（和名新称）HON
Genus Shigatheta Sawada, 1976 シガヒメハネカクシ属（和名新称）
tortuosa Sawada, 1970a: 45 (Ischnopoda) シガヒメハネカクシHON
Genus Stethusa Casey, 1910 ムネヒメハネカクシ属（和名新称）
miyamotoi Sawada, 1955: 85 (Ischnopoda) ムネヒメハネカクシ（和名新称）T-Nak
Genus Tomoglossa Kraatz, 1856 ツツヒメハネカクシ属（和名新称）
luteicornis Erichson, 1837: 332 (Homalota) ツツヒメハネカクシ（和名新称）Japan; China, Russia, Europe, Africa
Genus Tropimenelytron, Pace, 1983 スジバネヒメハネカクシ属（和名新称）
unicum Bernhauer, 1907: 409 (Atheta) スジバネヒメハネカクシ（和名新称）HON; North Korea
Subtribe Schistogeniina Fenyes, 1918 アバタヒメハネカクシ亜族（和名新称）
Genus Schistogenia Kraatz, 1857 アバタヒメハネカクシ属（和名新称）
crenicollis Kraatz, 1857a: 40 アバタヒメハネカクシ（和名新称）HON; Hong Kong, China, India, SE Asia
Subtribe Taxicerina Lohse, 1989 ヤサヒメハネカクシ亜族（和名新称）
Genus Taxicera Mulsant & Rey, 1873 ヤサヒメハネカクシ属（和名新称）
Subgenus Taxicera Mulsant & Rey, 1873
pseudorphana Cameron, 1933: 216 (Atheta) ヤサヒメハネカクシ( 和名新称 )HON
Subtribe Thamiaraeina Fenyes, 1921 ショウジョウハネカクシ亜族（和名新称）
Genus Thamiaraea Thomson, 1858 ショウジョウハネカクシ属（和名新称）
Subgenus Thamiaraea Thomson, 1858
diffinis Sharp, 1888: 292 ショウジョウハネカクシ（和名改称）HON
japonica Cameron, 1933: 216 ヒメショウジョウハネカクシ（和名新称）HON
Tribe Autaliini Thomson, 1859 スジハラハネカクシ族（和名新称）
Genus Autalia Leach, 1819 スジハラハネカクシ属（和名新称）
puncticollis Sharp, 1864: 45 クロスジハラハネカクシ（和名新称）HON; Europe, USA

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Yasutoshi Shibata, Munetoshi Maruyama, Hideto Hoshina, Toshio Kishimoto, Shun-Ichiro Naomi, Shûhei Nomura, Volker Puthz, Takashi Shimada, Yasuaki Watanabe, Shûhei Yamamoto

rivularis Gravenhorst, 1802: 73 (Aleochara) スジハラハネカクシ（ 和 名 新 称 ）HOK; China, Russia, Caucasus,
Europe, USArufula Sharp, 1888: 371 アカスジハラハネカクシ（和名改称）HOK, HON, KYU; China= pulchella Sawada, 1964a: 38
Tribe Deinopsini Sharp, 1883 ヌレチハネカクシ族（和名新称）
Genus Deinopsis Mattews, 1838 ヌレチハネカクシ属（和名新称）
minor Klimaszewski, 1979: 45 ヒメヌレチハネカクシ（和名新称）HON
modesta Sharp, 1874a: 16 ヌレチハネカクシ（和名新称）Japan; Russia
Tribe Falagriini Mulsant & Rey, 1873 セミゾハネカクシ族（和名新称）
Genus Falagria Leach, 1819 セミゾハネカクシ属
caesa Erichson, 1837: 295 クロセミゾハネカクシ HOK, HON, Sad, KYU; South Korea, North Korea, China, Russia,
Europe, Africa, USA= sulcata Paykull, 1789: 32 (Staphylinus)
Genus Falagrioma Casey, 1906 ヒメセミゾハネカクシ属
myrmecophila Sharp, 1888: 294 (Falagria) クサアリセミゾハネカクシ（和名新称）HOK, HON, SHI
simplex Sharp, 1874a: 2 (Falagria) ヒメセミゾハネカクシ（和名新称）HON; Taiwan
Genus Myrmecocephalus MacLeay, 1873 キバネセミゾハネカクシ属（和名新称）
concinnus Erichson, 1839a: 51 (Falagria) アメイロセミゾハネカクシ HON, SHI, Yak; Russia (Siberia), China,
Philippines, Indonesia (Java), India, Hawaii, Europe, S America= foveus Sharp, 1874a: 3 (Falagria)
hisamatsui Sawada & Nakane, 1954: 7 (Falagria) ヒサマツセミゾハネカクシ（和名新称）SHI
japonicus Bernhauer, 1939a: 105 (Falagria) ウスチャセミゾハネカクシ HOK, HON, SHI, KYU
sapidus Sharp, 1874a: 2 (Falagria) キバネセミゾハネカクシ HON, Sad, KYU, SKs, Tan, Yak, Amm; South Korea,
North Korea, Oriental Region
Genus Myrmecopora Saulcy, 1865 ウミセミゾハネカクシ属（和名新称）
Subgenus Lamproxenusa Assing, 1997
algarum Sharp, 1874a: 12 (Tachyusa) ウミセミゾハネカクシ HOK, HON, KYU
reticulata Assing, 1997: 344 ホソウミセミゾハネカクシ（和名新称）HON; North Korea, Russia
rufescens Sharp, 1874a: 11 (Tachyusa) ウスアカウミセミゾハネカクシ（和名新称）Japan; South Korea
Tribe Gymnusini Heer, 1839トマムハネカクシ族（和名新称）
Genus Gymnusa Gravenhorst, 1806トマムハネカクシ属（和名新称）2013 The Kyushu

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Catalogue of Japanese Staphylinidae (Insecta: Coleoptera)

miyashitai Naomi, 1994a: 47トマムハネカクシ（和名新称）HOK
Tribe Homalotini Heer, 1839 カレキハネカクシ族（和名新称）
Genus Bolitochara Mannerheim, 1830 ムネクボハネカクシ属（和名新称）
lobata Sawada, 1970b: 41 (Ditropalia) ヤマムネクボハネカクシ（和名新称）HON
obliqua Erichson, 1837: 298 ハスオビムネクボハネカクシ（和名新称）HON; Caucasus, Europe
varipes Sharp, 1888: 371 ハラモンムネクボハネカクシ HON
Genus Euryusa Erichson, 1837 アシボソシリアゲハネカクシ属
Subgenus Ectolabrus Sharp, 1888
hagai Maruyama & Miyake, 2003: 72 ケシアシボソシリアゲハネカクシ HOK
minor Maruyama & Hlaváč, 2002: 178 ヒメアシボソシリアゲハネカクシ HOK, HON, SHI
pipitzi Eppelsheim, 1887a: 229 (Silusa) アシボソシリアゲハネカクシHOK, Kun, HON, SHI, KYU, Tsm, Amm;
Russia (Far East), Korea, Austria= laticollis Sharp, 1888: 370 (Ectolabrus)
Genus Heterota Mulsant & Rey, 1874 ウシオハネカクシ属
arenaria Cameron, 1920: 251 サンゴウシオハネカクシ Ish; Indonesia, Singapore
ororum Maruyama, 2011a: 98 ウシオハネカクシ HON
sunjaei Park & Ahn, 2008: 111 キアシウシオハネカクシ HON; South Korea
Genus Leptusa Kraatz, 1856 コケホソハネカクシ属（和名新称）
Subgenus Aphaireleptusa Pace, 1996
puthzi Assing, 2002b: 981 プーツツヤケシコケホソハネカクシ（和名新称）HON
Subgenus Drepanoleptusa Pace, 1982
deplanata Sawada, 1970a: 40 (Sipalia) シガコケホソハネカクシ（和名新称）HON
ruficornis Cameron, 1933: 210 アカヒゲコケホソハネカクシ（和名新称）HON
yamato Sawada, 1990: 541 (Sipalia) ヤマトコケホソハネカクシ（和名新称）HON
Subgenus Dysleptusa Pace, 1982
honshuica Assing, 2002b: 975 ホンシュウコケホソハネカクシ（和名新称）HON
Subgenus Eospisalia Pace, 1982
ishizuchiensis Pace, 1982b: 584 イシズチコケホソハネカクシ（和名新称）SHI
kitazawai kitazawai Sawada, 1970a: 41 (Sipalia) キタザワコケホソハネカクシ（和名新称）HON
k. fujiensis Pace, 1982a:140 HON
lackneri Assing, 2011: 268 ツシマコケホソハネカクシ（和名新称）Tsm
taichii Kishimoto, 2008: 152 オオダイコケホソハネカクシ（和名新称）HON
Subgenus Heteroleptusa Pace, 1989
japonica Cameron, 1933: 209 ニホンコケホソハネカクシ（和名新称）KYU
Incertae cedis (subgenus?) 亜属不明

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Yasutoshi Shibata, Munetoshi Maruyama, Hideto Hoshina, Toshio Kishimoto, Shun-Ichiro Naomi, Shûhei Nomura, Volker Puthz, Takashi Shimada, Yasuaki Watanabe, Shûhei Yamamoto

cornigera Assing, 2002b: 997 (Eospisalia?) カイコケホソハネカクシ（和名新称）HON
Genus Phymatura Sahlberg, 1876 キノコツヤケシハネカクシ属（和名新称）
japonica Cameron, 1930: 210 ニホンキノコツヤケシハネカクシ（和名新称）HON 種復活予定
crenulicauda Bernhauer, 1907: 398 (Atheta) キノコツヤケシハネカクシ（和名新称）HON; North Korea
Genus Tachyusida Mulsant & Rey, 1872 コゲチャクチキハネカクシ属
velox Sharp, 1888: 372 コゲチャクチキハネカクシHON, SHI, KYU
Subtribe Gyrophaenina Kraatz, 1856 キノコツヤハネカクシ亜族（和名新称）
Genus Brachida Mulsant & Rey, 1872 オチバハネカクシ属（和名新称）
brachyptera Kishimoto, 2000: 75 アマミコバネオチバハネカクシ（和名改称）Amm, Tok
brevipennis Bernhauer, 1939a: 103 コバネオチバハネカクシ（和名改称）HON, SHI, KYU
clara Weise, 1877a: 90 (Homalota) オチバハネカクシ（和名改称）HON, SHI, KYU, Tsm, Got, Yak, Tan, T-R,
Okn, Ish
Genus Gyrophaena Mannerheim, 1830 キノコツヤハネカクシ属（和名改称）
Subgenus Acanthophaena Cameron, 1834
appendiculata Motschulsky, 1858c: 228 オオメキノコツヤハネカクシ（和名改称）HON, KYU, Tan, Yak, Amm;
Philippines, Malaysia, India, Oriental Region
Subgenus Gyrophaena Mannerheim, 1830
bernhaueri Jacobson, 1909: 533 [RN] ニホンキノコツヤハネカクシ（和名新称）HON= japonica Bernhauer, 1914a: 9 [RN]
= puncticollis Bernhauer, 1907: 390 [HN]biseriata Cameron, 1933: 175 フタバキノコツヤハネカクシ（和名新称）HON
cariniventris Cameron, 1933: 173 セスジキノコツヤハネカクシ（和名新称）HON
cicatricosa Motschulsky, 1858c: 231 ミナミキノコツヤハネカクシ（和名新称）Og; Hong Kong, India, Oriental Region
defecta Cameron, 1933: 209 HON 下位同物異名予定
futamata Cameron, 1933: 209 フタマタキノコツヤハネカクシ（和名新称）HON
harmandi Fauvel, 1901a: 4 アルマンキノコツヤハネカクシ（和名新称）Japanlaevior Cameron, 1933: 174 スベキノコツヤハネカクシ（和名改称）HON
lewisiana Cameron, 1933: 175 ルイスキノコツヤハネカクシ（和名新称）HON
mitsumata Cameron, 1933: 209 ミツマタキノコツヤハネカクシ（和名新称）HON
niponensis Cameron, 1933: 174 ニッポンキノコツヤハネカクシ（和名改称）HOK, HON; South Korea, China
puncticeps Bernhauer, 1907: 389 ナミキノコツヤハネカクシ（和名新称）HON
rugilipennis Cameron, 1933: 174 シワバネキノコツヤハネカクシ（和名新称）HON
sapporensis Sharp, 1888: 376 キタキノコツヤハネカクシ（和名改称）HOK
settsuensis Cameron, 1933: 208 セッツキノコツヤハネカクシ（和名新称）HON
triquetra Weise, 1877a: 91 ミツバキノコツヤハネカクシ（和名新称）HON

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Catalogue of Japanese Staphylinidae (Insecta: Coleoptera)tsushimana Bernhauer, 1936: 310 ツシマキノコツヤハネカクシ（和名新称）Tsm
yotsudeba Cameron, 1933: 208 ヨツデバキノコツヤハネカクシ（和名新称）HON
yotsumata Cameron, 1933: 208 ヨツマタキノコツヤハネカクシ（和名新称）HON
Subgenus Leptarthrophaena Scheerpeltz, 1948
hanedai Sawada, 1970b: 39 ハネダキノコツヤハネカクシ（和名新称）HON
Subtribe Homalotina Heer, 1839 カレキハネカクシ亜族（和名新称）
Genus Anomognathus Solier, 1849 ハラトゲカレキハネカクシ属（和名新称）
armatus Sharp, 1888: 294 (Thectura) ハラトゲカレキハネカクシ（和名改称）HON, KYU, SKs
Genus Homalota Mannerheim, 1830 カレキハネカクシ属（和名新称）
fraterna Sharp, 1888: 376 (Epipeda) カレキハネカクシ（和名新称）HON; Taiwan, China
mikado Likovský, 1984: 6 [RN] KYU; Taiwan= granigera Sharp, 1888: 375 [HN] (Epipeda) ツブツブカレハハネカクシ（和名新称）( 属所属変更予定)
sauteri Bernhauer, 1907: 391 カレキハネカクシ（和名新称）Japan; Taiwan
Subtribe Silusina Fenyes, 1918 クチヒゲハネカクシ亜族（和名新称）
Genus Coenonica Kraatz, 1857 カレハハネカクシ属（和名新称）
anteatomica Pace, 1993b: 133 ヘイワカレハハネカクシ（和名新称）Japan
lewisa Sharp, 1874a: 14 (Homalota) ルイスカレハハネカクシ（和名改称）HON; North Korea
puncticollis Kraatz, 1857b: 46 カレハハネカクシ（和名新称）Yak; India, Pan Tropics
sharpi Fauvel, 1901b: 66 [RN] (Leptusa) シャープカレハハネカクシ（和名改称）HON, KYU= impresicollis Sharp, 1888: 371 [HN] (Leptusa)
Genus Neosilusa Cameron, 1920 ヒゲナガカレハハネカクシ属（和名新称）
ceylonica Kraatz, 1857a: 8 (Stenusa) HON; Taiwan, China= crassicornis Sharp, 1888: 374 (Silusa) ヒゲナガカレハハネカクシ（和名新称）種復活予定
Genus Silusa Erichson, 1837 クチヒゲハネカクシ属（和名新称）
Subgenus Silusa Erichson, 1837
biimpressa Bernhauer, 1907: 390 フタテンクチヒゲハネカクシ（和名新称）HON
conferta Sharp, 1888: 374 ハコネコケホソハネカクシ（和名新称）HON 属所属変更予定
lanuginosa Sharp, 1888: 375 クチヒゲハネカクシ
KYU, SKs; China（和名改称）
punctipennis Sharp, 1888: 373 アラメコケホソハネカクシ（和名新称）HON 属所属変更予定
rorida Sharp, 1888: 373 ヒメツヤケシコケホソハネカクシ（和名新称）HON 属所属変更予定
Subgenus Stenusa Kraatz, 1856
rugosa Sharp, 1888: 372 ツヤケシコケホソハネカクシ（和名改称）HON, SKs 属所属変更予定

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Yasutoshi Shibata, Munetoshi Maruyama, Hideto Hoshina, Toshio Kishimoto, Shun-Ichiro Naomi, Shûhei Nomura, Volker Puthz, Takashi Shimada, Yasuaki Watanabe, Shûhei Yamamoto

Tribe Hoplandriini Casey, 1910 ニセヒゲブトハネカクシ族（和名新称）
Genus Hoplandria Kraatz, 1857 オオニセヒゲブトハネカクシ属（和名新称）
convexa Weise, 1877: 88 オオニセヒゲブトハネカクシ（和名新称）HON
spiniventris Bernhauer, 1907: 392 ハサミニセヒゲブトハネカクシ（和名改称）HON
Genus Omoplandria Cameron, 1949 ヒメハナハネカクシ属（和名新称）
gyrophaenula Sharp, 1888: 294 (Homalota) ミチコヒメハナハネカクシ HON
Genus Pseudoplandria Fenyes, 1921 ニセヒゲブトハネカクシ属（和名新称）
sakuradanii Sawada, 1990: 548 サクラダニニセヒゲブトハネカクシ（和名新称）HON; North Korea
Tribe Hygronomini Thomson, 1859 日本に分布しない
Subtribe Hygronomina Thomson, 1859
Genus Protinodes Sharp, 1888 ズングリハネカクシ属（和名新称）族所属変更予定
puncticollis Sharp, 1888: 378 ズングリハネカクシ（和名新称）HON
Tribe Hypocyphtini Laporte, 1835 ケシハネカクシ族（和名新称）
Genus Cypha Leach, 1819 タマキノコハネカクシ属（和名新称）
nomurai Naomi, 1987c: 274 (Hypocyptus) ニホンタマキノコハネカクシ Okn
Genus Holobus Solier, 1849 ハダニカブリケシハネカクシ属（和名改称）
antennatatus Bernhauer, 1907: 388 (Oligota) マルハダニカブリケシハネカクシ属（和名改称）Japan
kashmirica benefica Naomi, 1984: 517 (Oligota) ヒメハダニカブリケシハネカクシ HON, Iz-Hch, SHI, KYU
kurama Nakane, 1963: 21 (Oligota) コマルハダニカブリケシハネカクシ属（和名改称）HON
yasumatsui Kistner, 1969: 142 (Oligota) ハダニカブリケシハネカクシ HON, Iz-Hch, Awj, SHI, KYU; South Korea
Genus Oligota Mannerheim, 1830 ケシハネカクシ属（和名新称）
japonica Bernhauer, 1923b: 146 ケシハネカクシ（和名新称）HON
parvula Cameron, 1933: 173 ミジンケシハネカクシ（和名新称）HON
Tribe Leucocraspedini Fenyes, 1921 マルケシハネカクシ族（和名新称）
Genus Leucocraspedum Kraatz, 1859 マルケシハネカクシ属（和名新称）
pallidum Cameron, 1933: 172 HON, KYU; Taiwan 下位同物異名予定
parvum Cameron, 1949: 470 Yak, Amm 下位同物異名予定
rufotestaceum Bernhauer, 1927: 97 マルケシハネカクシ（和名新称）Tsm

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日本産ハネカクシ科総目録（昆虫綱：甲虫目）
Catalogue of Japanese Staphylinidae (Insecta: Coleoptera)

Tribe Liparocephalini Fenyes, 1918 ウミハネカクシ族
Genus Amblopusa Casey, 1894 ノコバウミハネカクシ属
pacifica Sawada, 1991: 147 (Boreorhadinus) ノコバウミハネカクシ HOK; USA (Alaska), Canada
hokkaidona Ahn & Ashe, 1996: 142 ヒメノコバウミハネカクシ HOK
magna Zerche, 1998: 106 オオノコバウミハネカクシ HOK; Russia (Primorsky, N Kuril)
Genus Diaulota Casey, 1894 ツツムネウミハネカクシ属
aokii Sawada, 1971c: 104 アオキツツムネウミハネカクシ HOK, HON, Iz-O, Shd; South Korea, USA (Alaska)
pacifica Sawada, 1971: 101 ホソツツムネウミハネカクシ HON, SHI, Shd; South Korea
uenoi Sawada, 1955: 82 (Genoplectes) シリブトウミハネカクシ HON, SHI, T-R; South Korea
Genus Halorhadinus Sawada, 1971 イソハネカクシ属
aequalis Sawada, 1971c: 92 クロイソハネカクシ HON; South Korea
inaequalis Sawada, 1971c: 95 ワカサイソハネカクシ HON; South Korea
sawadai Maruyama & M. Hayashi, 2009: 72 ウマヅライソハネカクシ HON
Genus Liparocephalus Mäklin, 1853 オオズウミハネカクシ属
litoralis Kirschenblat, 1938: 532 ホテイウミハネカクシHOK, Chs; Russia (Komandor)
tokunagai Sakaguti, 1944: 20 オオズウミハネカクシ HON, SHI, KYU
Genus Paramblopusa Ahn & Ashe, 1996 ニセノコバウミハネカクシ属
borealis Casey, 1906: 355 (Amblopusa) ニセノコバウミハネカクシHOK; Canada, USA
Tribe Lomechusini Fleming, 1821 アリノスハネカクシ族
Subtribe Lomechusina Fleming, 1821 ハケゲアリノスハネカクシ亜族
Genus Lomechusa Gravenhorst, 1806 ハケゲアリノスハネカクシ属
sinuata Sharp, 1888: 288 (Atemeles) ハケゲアリノスハネカクシ HOK, HON, SHI, KYU; Russia (Sakhalin)
Genus Lomechusoides Tottenham, 1939 タカネアリノスハネカクシ属
amurensis Wasmann, 1897: 247 (Lomechusa) Japan; Russia, Mongolia= suensoni Bernhauer, 1936: 323 (Lomechusa) タカネアリノスハネカクシ HON 種復活予定= hosodai Sawada, 1994: 363 (Lomechusa)
Subtribe Myrmedoniina Thomson, 1867 アリノスハネカクシ亜族
Genus Drusilla Leach, 1819 ニセセミゾハネカクシ属（和名新称）
Subgenus Drusilla Leach, 1819
aino Nakane, 1963: 21 ニセセミゾハネカクシHOK, Reb, HON; China

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Yasutoshi Shibata, Munetoshi Maruyama, Hideto Hoshina, Toshio Kishimoto, Shun-Ichiro Naomi, Shûhei Nomura, Volker Puthz, Takashi Shimada, Yasuaki Watanabe, Shûhei Yamamoto

nomurai Maruyama, 2000b: 362 ツシマニセセミゾハネカクシ Tsmsparsa Sharp, 1874a: 4 (Santhota) アカニセセミゾハネカクシHON, SHI, KYU, IzIss, Got, SKs; South Korea
Genus Pella Stephens, 1835 クサアリハネカクシ属
beijingorum Pace, 1998: 976 (Zyras (Pella)) ヒメクサアリハネカクシHOK, HON; Russia (Far East), China
comes Sharp, 1874a: 10 (Myrmedonia) クロツヤクサアリハネカクシ HON, SHI, KYU
japonica Sharp, 1888: 290 (Myrmedonia) ネアカクサアリハネカクシ HOK, HON, SHI, KYU; South Korea, Russia
indiscreta Sharp, 1888: 291 (Myrmedonia) チゴクサアリハネカクシ HOK, HON, SHI
kidaorum Maruyama, 2006: 73 エゾクサアリハネカクシ HOK
kinomurai Maruyama, 2006: 96 アカアシクサアリハネカクシ HOK, HON
horii Maruyama, 2006: 37 クマハネカクシ HOK, HON
masakoae Maruyama, 2006: 156 オオクサアリハネカクシ HOK, HON, SHI; South Korea
socia Sharp, 1874a: 10 (Myrmedonia) コクロツヤクサアリハネカクシ HOK, HON, SHI, KYU
spreta Sharp, 1888: 291 (Myrmedonia) メダカクサアリハネカクシ HOK, HON
Genus Tetrabothrus Bernhauer, 1915 コンボウヒゲブトハネカクシ属
japonicus Nakane, 1991: 111 コンボウヒゲブトハネカクシ HON, SHI, KYU, Yak, Amm, Ish, Iri, Yon; South Korea
septentrionalis Kishimoto, 1997: 446 キタコンボウヒゲブトハネカクシ HOK, HON, SHI, Tsm; South Korea
validus Maruyama & Kishimoto, 1999: 513 オニコンボウヒゲブトハネカクシ HON
Genus Zyras Stephens, 1835 アリノスハネカクシ属
Subgenus Zyras Stephens, 1835
cylindricornis M. Dvořák, 1981: 56 HON; South Korea, China 下位同物異名予定
fugax Sharp, 1888: 289 (Myrmedonia) クロヒゲアリノスハネカクシ HON; South Korea
haworthi Stephens, 1832: 126 (Aleochara) モンクロアリノスハネカクシ HON, KYU; Europe
iridescens Sawada, 1970a: 49 (Bolitochara) ヒゲブトアリノスハネカクシ HON
optatus Sharp, 1888: 289 (Myrmedonia) コモンクロアリノスハネカクシ HON, Iz-My, Yak, Kum; Russia
particornis Sharp, 1888: 290 (Myrmedonia) シロヒゲアリノスハネカクシ HON; South Korea, North Korea, China,
Russiapictus Sharp, 1874a: 11 (Ilyobates) クビアカアリノスハネカクシ HON, KYU; South Korea, North Korea
Subtribe Termitozyrina Seevers, 1957 オオシロアリハネカクシ亜族
Genus Hodotermophilus Naomi & Terayama, 1986 オオシロアリハネカクシ属
gloriosus Naomi & Terayama, 1986a: 507 オオシロアリハネカクシ Yak
Tribe Mesoporini Cameron, 1959 クチキケシハネカクシ族
Genus Dictyon Fauvel, 1900 ツチケシハネカクシ属（和名新称）
insulicola Kishimoto, 1999: 207 シマツチケシハネカクシ Og

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日本産ハネカクシ科総目録（昆虫綱：甲虫目）
Catalogue of Japanese Staphylinidae (Insecta: Coleoptera)

Genus Kistnerium Naomi & Iwata, 1996 クチキケシハネカクシ属（和名新称）
japonicum Naomi & Iwata, 1996: 78 クチキケシハネカクシ（和名改称）HON, KYU
Tribe Myllaenini Ganglbauer, 1895 ミギワハネカクシ族
Genus Bryothinusa Casey, 1904 ナギサハネカクシ属
algarum Sawada, 1971c: 90 ウスアカナギサハネカクシ HON, KYU, SKs
minuta Sawada, 1955: 83 (Halesthenus) ホソナギサハネカクシ T-R; South Korea
nakanei Sawada, 1955: 85 (Halesthenus) ナカネナギサハネカクシ HON, KYU, T-R; South Korea
sakishimana Sawada, 1991: 144 サキシマナギサハネカクシ Iritsutsuii Sawada, 1955: 84 (Halesthenus) キイロナギサハネカクシ HON, T-R; South Korea
Genus Myllaena Erichson, 1837 ミギワハネカクシ属
graeca Kraatz, 1858: 54 オウシュウミギワハネカクシ（和名新称）HON; Turkistan, Caucasus, Europe
japonica Sharp, 1888: 377 ミギワハネカクシ（和名改称）HON, SHI, KYU, Tan, Yak; North Korea, China
lewisi Cameron, 1933: 173 ルイスミギワハネカクシ（和名新称）HON
rufotestacea Cameron, 1933: 173 ヒメミギワハネカクシ（和名新称）HON
torrentum Cameron, 1933: 172 オオミギワハネカクシ（和名新称）HON
Tribe Oxypodini Thomson, 1859 ゴミハネカクシ族
Subtribe Oxypodina Thomson, 1859 ゴミハネカクシ亜族（和名新称）
Genus Aspidobactrus Sharp, 1888 マルムネアリヤドリ属
claviger Sharp, 1888: 284 マルムネアリヤドリ HON
formicae Maruyama, 2000a: 168 ヤママルムネアリヤドリ HON
semiorbiculatus Maruyama, 2000a: 157 ニシマルムネアリヤドリ HON, SHI, KYU
toyodai Maruyama, 2000a: 162 ヒメマルムネアリヤドリ HON, KYU
Genus Diploxenus Kistner & Akatsuka, 1981 ケシアリヤドリ属
kubotai Kistner & Akatsuka, 1981: 179 ケシアリヤドリ HON
Genus Homoeusa Kraatz, 1856 ヒラタアリヤドリ属
acuminata Märkel, 1842: 143 (Euryusa) HON; Europe 日本に分布しない(Maruyama, in prep.)japonica Sharp, 1874a: 5 ヤマトヒラタアリヤドリ HON, KYU
laevigata Sharp, 1888: 283 ツヤヒラタアリヤドリ HON
longicornia Sharp, 1888: 283 ヒゲナガヒラタアリヤドリHOK, HON
prolongata Sawada, 1970a: 57 ヒラタアリヤドリ HON
Genus Amarochara Thomson, 1858 フトヒゲゴミハネカクシ属（和名新称）
flavicornis Bernhauer, 1907: 412 フトヒゲゴミハネカクシ（和名新称）HON

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Yasutoshi Shibata, Munetoshi Maruyama, Hideto Hoshina, Toshio Kishimoto, Shun-Ichiro Naomi, Shûhei Nomura, Volker Puthz, Takashi Shimada, Yasuaki Watanabe, Shûhei Yamamoto

Genus Calodera Mannerheim, 1830 ハラクボゴミハネカクシ属（和名新称）
desdemona Sharp, 1888: 286 ハラクボゴミハネカクシ（和名新称）HON
protensa Mannerheim, 1830: 86 オウシュウハラクボゴミハネカクシ（和名新称）Japan; Europe
Genus Devia Blackwelder, 1952 ニセゴミハネカクシ属（和名新称）
prospera Erichson, 1839a: 143 (Oxypoda) ニセゴミハネカクシ属（和名新称）HOK; Europe, Russia (Siberia), USA
Genus Losiusa Seevers, 1978 ナガアリヤドリ属（和名新称）
oxypodina Sharp, 1888: 284 (Thiasophila) ナガアリヤドリ（和名新称）HON
Genus Ocalea Erichson, 1837 日本に分布しない
japonica Sharp, 1874a: 4 キュウシュウクサビラハネカクシ KYU 所属変更予定
Genus Oreuryalea Assing & Maruyama, 2002 ウミゴミハネカクシ属（和名新称）
watanabei Assing & Maruyama, 2002: 217 ウミゴミハネカクシ HOK; Russia (Primorsky, Sakhalin)
Genus Oxypoda Mannerheim, 1830 ゴミハネカクシ属
Subgenus Baeoglena Thomson, 1858
producta Sawada, 1970b: 51 シナノゴミハネカクシ（和名新称）HON
Subgenus Bessopora Thomson, 1859
imadatei Sawada, 1970b: 56 イマダテゴミハネカクシ（和名新称）HON
kobensis Cameron, 1933: 218 コウベゴミハネカクシ（和名新称）HON
Subgenus Mycetodrepa Thomson, 1859
renominata Bernhauer & Scheerpeltz, 1926: 757 [RN] フォーベルゴミハネカクシ（和名新称）HON= japonica Fauvel, 1901a: 5 [HN]
Subgenus Oxypoda Mannerheim, 1830
japonica Sharp, 1874a: 12 オオゴミハネカクシ（和名新称）Japanluridipennis Sharp, 1888: 285 ツヤゴミハネカクシ（和名改称）HON; North Korea= lewisi Cameron, 1933: 218
obtusa Sawada, 1970b: 53 ホソゴミハネカクシ（和名新称）HON
Subgenus Podoxya Mulsant & Rey, 1875
kuramotoi Sawada, 1971a: 87 クラモトゴミハネカクシ（和名新称）HON
proba Sharp, 1874a: 13 フトゴミハネカクシ（和名新称）Japansauteri Bernhauer, 1907: 414 ザウターゴミハネカクシ（和名新称）HON
shimabarana Bernhauer, 1939b: 150 シマバラゴミハネカクシ（和名新称）KYU
subrufa Sharp, 1888: 285 アカゴミハネカクシ HON, KYU; North Korea
Insertae cedis (subgenus?) 亜属所属不明
laeta Weise, 1877a: 97 ヒメゴミハネカクシ（和名新称）HON
hilaris Sharp, 1888: 285 ヒメキノコツヤケシハネカクシ（和名新称）HON 所属変更予定

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Catalogue of Japanese Staphylinidae (Insecta: Coleoptera)

Genus Parocyusa Bernhauer, 1902 ナナフシハネカクシ属（和名新称）
Subgenus Tetralaucopora Bernhauer, 1902
japonica Cameron, 1933: 217 (Chilopora) ナナフシハネカクシ（和名新称）KYU
Genus Pentanota Bernhauer, 1902 イツホシハネカクシ属（和名新称）
alpicola Sawada, 1988: 37 イツホシハネカクシ（和名新称）HON
Genus Porocallus Sharp, 1888 ツマキリチビハネカクシ属（和名新称）
insignis Sharp, 1888: 287 ツマキリチビハネカクシ HOK, HON, SHI, KYU: Russia (Maritime), North Korea, China
Genus Rhomphocallus Assing, 2003 ツヤケシゴミハネカクシ属（和名新称）
maruyamai Assing, 2003b: 170 マルヤマツヤケシゴミハネカクシ（和名新称）HOK, Yak
princeps Sharp, 1874a: 6 (Microglotta) ツヤケシゴミハネカクシ（和名新称）HON
Genus Thiasophila Kraatz, 1856 アリヤドリ属
rufescens Sharp, 1874a : 5 ホソヒラタアリヤドリ（和名新称）Japan 属所属変更予定
Subtribe Tachyusina Thomson, 1859 ホソクロハネカクシ亜族（和名新称）
Genus Gnypeta Thomson, 1858 クビレハネカクシ属（和名新称）
Subgenus Gnypeta Thomson, 1858
ainu Sawada, 1976: 131 アイヌクビレハネカクシ（和名新称）HOK
aokii Sawada, 1970a: 42 クビレハネカクシ（和名新称）HON
Genus Ischnopoda Stephens, 1835 ヒメクビレハネカクシ属（和名新称）
semiopaca Bernhauer, 1939a: 106 (Tachyusa) ヒメクビレハネカクシ（和名新称）KYU
wei Pace, 1993a: 88 (Tachyusa) シナヒメクビレハネカクシ（和名新称）HON; China
Genus Tachyusa Erichson, 1837 ホソクロハネカクシ属（和名新称）
Subgenus Tachyusa Erichson, 1837
coarctata Erichson, 1837: 308 ホソクロハネカクシ（和名改称）HOK, HON; North Korea, China, Russia (Siberia),
Europereitteri Bernhauer, 1939a: 106 ライターホソクロハネカクシ（和名新称）KYU; China
Tribe Placusiini Mulsant & Rey, 1871トゲジリハネカクシ族（和名新称）
Genus Placusa Erichson, 1837トゲジリハネカクシ属（和名新称）
tachypoloides Waltl, 1838: 268 (Aleochara) トゲジリハネカクシ（和名改称）KYU; South Korea, North Korea,
Russia (Siberia), Europe

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Yasutoshi Shibata, Munetoshi Maruyama, Hideto Hoshina, Toshio Kishimoto, Shun-Ichiro Naomi, Shûhei Nomura, Volker Puthz, Takashi Shimada, Yasuaki Watanabe, Shûhei Yamamoto

Tribe Pronomaeini Mulsant & Rey, 1873 クチナガハネカクシ族（和名新称）
Genus Brachypronomaea Sawada, 1956 サンゴハネカクシ属（和名新称）
esakii Sawada, 1956c: 197 エサキサンゴハネカクシ Okn, Ish
Tribe Termitohospitini Seevers, 1941 シロアリヤドリ族
Subtribe Hetairotermitina Seevers, 1957 シロアリヤドリ亜族（和名新称）
Genus Japanophilus Maruyama & Iwata, 2002 ダルマシロアリヤドリ属
hojoi Maruyama & Iwata, 2002: 427 ダルマシロアリヤドリ Yak, T-Nak, T-Sw
Genus Sinophilus Kistner, 1985 シロアリヤドリ属
yukoae Maruyama & Iwata, 2002: 422 シロアリヤドリ Ihy, Ish, Iri
Tribe Trichopseniini LeConte & Horn, 1883 シロアリハネカクシ族
Genus Trichopsenius Horn, 1877 シロアリハネカクシ属
brunneus Naomi & Terayama, 1986b: 702 ウスチャシロアリハネカクシ Tok
crassicornis Naomi & Terayama, 1996: 84 ヤエヤマシロアリハネカクシ（和名新称）Irijaponicus Seevers, 1957: 271 シロアリハネカクシ HON
matsumotoi Naomi & Terayama, 1986b: 700 マツモトシロアリハネカクシYak
serratus Naomi & Terayama, 1986b: 701 タテミゾシロアリハネカクシAmm
Genus Platyolopsis Scheerpeltz, 1959 ニセトビムシハネカクシ属（和名新称）族所属変更予定
paradoxa Bernhauer, 1907: 413 ニセトビムシハネカクシ（和名新称）HON

Subfamily Apateticinae Fauvel, 1859 オサシデムシモドキ亜科（２属２種）
Genus Apatetica Westwood, 1848 オサシデムシモドキ属
princeps Sharp, 1874c: 421 (Trygaeus) オサシデムシモドキ HON, SHI, KYU
Genus Nodynus Waterhouse, 1876 シデムシモドキ属
leucofasciatus Lewis, 1879: 460 シラオビシデムシモドキ HOK, HON, Sad, SHI, KYU

Subfamily Scaphidiinae Latreille, 1807 デオキノコムシ亜科（12属89種）

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Catalogue of Japanese Staphylinidae (Insecta: Coleoptera)

Tribe Cypariini Achard, 1924
Genus Cyparium Erichson, 1845 カメノコデオキノコムシ属
jiroi Ogawa & Sakai, 2011: 130 マメカメノコデオキノコムシ KYU
laevisternale Nakane, 1963: 31 コカメノコデオキノコムシ HON, SHI, KYU
mikado Achard, 1923: 109 カメノコデオキノコムシ HOK, HON, SHI, KYU, Tsm; South Korea, China
Tribe Scaphiini Achard, 1924
Genus Ascaphium Lewis, 1893 スジデオキノコムシ属
apicale Lewis, 1893a: 290 ヒメセスジデオキノコムシ HON, SHI, KYU
sulcipenne Lewis, 1893a: 289 アカアシスジデオキノコムシ HON
tibiale Lewis, 1893a: 289 ホソスジデオキノコムシ HON, SHI, KYU
Genus Episcaphium Lewis, 1893 アカバデオキノコムシ属
semirufum Lewis, 1893a: 291 アカバデオキノコムシ HON, SHI, KYU= ruficolle Lewis, 1893a: 291
Tribe Scaphidiini Latreille, 1807
Genus Scaphidium Olivier, 1790 デオキノコムシ属
amamiense Hoshina & Morimoto, 1999: 90 アマミデオキノコムシ Amm, Tok
amurense Solsky, 1871: 350 ツシマデオキノコムシ Japan, Tsm; Korea, Russia= tsushimense Shirozu & Morimoto, 1963: 74
brunneum Hoshina & Morimoto, 1999: 87 チャイロデオキノコムシ Okn
chujoi Löbl, 1967: 129 チュウジョウデオキノコムシ SHI, KYU
emarginatum Lewis, 1893a: 291 エグリデオキノコムシ HOK, HON, SHI, KYU
femorale Lewis, 1893a: 292 ヒメデオキノコムシ HON, SHI, KYU
inagoense Kimura, 2008: 160 ミヤマデオキノコムシ HON, SHI
incisum Lewis, 1893a: 294 ヒメクロデオキノコムシ HOK, HON, KYU
japonum Reitter, 1877: 369 ヤマトデオキノコムシ HOK, HON, SHI, KYU, Tsm= longipes Lewis, 1893a: 292
kumejimaense Hoshina & Maruyama, 1999: 480 クメジマデオキノコムシ Kum, Ishlewisi Achard, 1923: 108 (Scaphidiolum) セモンコガタデオキノコムシ KYU
montivagum Shirozu & Morimoto, 1963: 70 コヒメデオキノコムシ HON; China
morimotoi Löbl, 1982b: 101 [RN] クロツヤデオキノコムシ Yor= takahashii Shirozu & Morimoto, 1963: 64 [HN]okinawaense Hoshina & Morimoto, 1999: 91 オキナワデオキノコムシ Okn
optabile Lewis, 1893a: 290 (Scaphium) コクロデオキノコムシ KYU
reitteri Lewis, 1879: 460 ヘリアカデオキノコムシ HON, SHI, KYU, Tsm, Amm

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Yasutoshi Shibata, Munetoshi Maruyama, Hideto Hoshina, Toshio Kishimoto, Shun-Ichiro Naomi, Shûhei Nomura, Volker Puthz, Takashi Shimada, Yasuaki Watanabe, Shûhei Yamamoto= insulare Achard, 1922: 38
rufopygum Lewis, 1893a: 293 シリアカデオキノコムシ HON, SHI, KYU
sakura Hoshina 2001: 101 サクラデオキノコムシ Okn, Kum
taichii Kimura, 2008: 158 ヤクシマデオキノコムシ Yak
takemurai Nakane, 1956: 30 タケムラデオキノコムシ KYUyasumatsui Shirozu & Morimoto, 1963: 67 カギアシデオキノコムシ SHI, KYU
Tribe Scaphisomatini Casey, 1894
Genus Baeocera Erichson, 1845 マメデオキノコムシ属
abnormalis Nakane, 1963: 22 マルマメデオキノコムシ HON, KYU
caliginosa Löbl, 1984: 186 オキナワマメデオキノコムシ KYU, Okn
curtula Achard, 1923: 117 マメデオキノコムシ HON, SHI, KYU
frater Löbl, 1969: 334 (Eubaeocera) オオマメデオキノコムシ HON, SHI
micros Achard, 1923: 116 (Scaphosoma) ナガサキマメデオキノコムシ KYU
myrmidon Achard, 1923: 116 (Scaphosoma) ヒゼンマメデオキノコムシ KYU; Taiwan
nakanei Löbl, 1968: 1 (Eubaeocera) ナカネマメデオキノコムシ HON
satana Nakane, 1963: 22 アカマメデオキノコムシ KYU; China
sordida Löbl, 1984: 183 イシヅチマメデオキノコムシ SHI
takizawai Löbl, 1984: 190 ヤエヤママメデオキノコムシ IshGenus Caryoscapha Ganglbauer, 1899 アゴヒゲケシデオキノコムシ属
seorsum Löbl, 1965a: 1 アゴヒゲケシデオキノコムシ HOK, HON
Genus Pseudobironium Pic, 1920 ツブデオキノコムシ属
lewisi Achard, 1923: 118 ツブデオキノコムシ HON, SHI, KYU
Genus Scaphisoma Leach, 1815 ケシデオキノコムシ属
adnexum Löbl, 1972: 117 クロミケシデオキノコムシ HON
austerum Löbl, 1965b: 1 ニセツマキケシデオキノコムシ HON
brunneonotatum Pic, 1923: 17 チャモンケシデオキノコムシ Okn, Ish, Iri; Taiwan, China, Vietnam
castaneipenne Reitter, 1877: 370 クリイロケシデオキノコムシ HON, SHI, KYU, Tsm
colasi Löbl, 1965c: 55 コーラケシデオキノコムシ HOK, HON
conforme okinawense Löbl, 1982b: 104 オキナワケシデオキノコムシ Iz-Hch, Okn
conspicuum Löbl, 1982a: 19 コゲチャケシデオキノコムシ HON, SHI, Fukcrassipes Achard, 1923: 115 アシブトケシデオキノコムシ HON
danielae Löbl, 1982c: 332 クロホシケシデオキノコムシ SHI
galloisi Achard, 1923: 114 ガロアケシデオキノコムシ HON, Tsm, SHI= harmandi Achard, 1923: 114
gentile Löbl, 1982c: 329 アカグロケシデオキノコムシ HON, SHI

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Catalogue of Japanese Staphylinidae (Insecta: Coleoptera)

hadrops Löbl, 1965c: 57 エゾケシデオキノコムシ HOK
haemorrhoidale Reitter, 1877: 369 ツマキケシデオキノコムシ HON, SHI, KYU, Tsm; China, North Korea, Russia(Siberia)= lewisi Achard, 1923: 112
hisamatsui Löbl, 1982c: 328 ヒサマツケシデオキノコムシ SHIindubium Löbl, 1965c: 50 チャバネケシデオキノコムシ HON; Russia (Siberia)
insulanum Löbl, 1982b: 102 アマミケシデオキノコムシ Amm
iriomotense Löbl, 1977: 163 イリオモテケシデオキノコムシ Irijanczyki Löbl, 1965c: 52 ヤンチクケシデオキノコムシ HON= ignotum Löbl, 1965c: 53
japonicum Löbl, 1965c: 47 ヤマトケシデオキノコムシ HON, SHI; China, Russia (Siberia)
jelineki Löbl, 1965c: 56 イエリネクケシデオキノコムシ HON
laevigatum Löbl, 1970: 755 ツヤケシデオキノコムシ HON; Taiwan, China, Russia
pecki Löbl, 1982a: 21 ペックケシデオキノコムシ SHI
perkinsi Scott, 1908 パーキンスケシデオキノコムシ Og-cc; Hawaii
rubrum Reitter, 1877: 370 アカミケシデオキノコムシ HON, SHI, KYU, Tsm, T-Nak, Okn, Iri= ustulatum Achard, 1923: 112
rufum Achard, 1923: 115 アカケシデオキノコムシ HOK, HON, SHI, KYU, Tsm, Amm, Okn, Ish, Iri; Korea, China
sakaii Löbl, 1988b: 1133 リュウキュウケシデオキノコムシ RK
sasagoense Löbl, 1965c: 49 ササゴケシデオキノコムシ HON
taiwanum Löbl, 1980: 106 タイワンデオキノコムシ HON; Taiwan, Russia
tamaninii Löbl, 1965b: 3 タマニーニケシデオキノコムシ HOK, HON; Russia (Siberia)tricolor Heller, 1917: 46 ミイロケシデオキノコムシ Og-hh; Philippines, Oriental Region
unicolor Achard, 1923: 113 キョウトケシデオキノコムシ HON, SHI, KYU; Taiwan, China, Russia
Genus Scaphobaeocera Csiki, 1909 ホテイケシデオキノコムシ属
abnormalis Löbl, 1981: 240ドウキョウホテイケシデオキノコムシ（和名改称）SHI
dorsalis Löbl, 1980: 118 アジアホテイケシデオキノコムシ Yon; SE Asia, China, Taiwan, South Korea
gracilis Löbl, 1981: 235 アカホソホテイケシデオキノコムシ（和名改称）HON
hisamatsui Hoshina, 2008b: 141 ヤエヤマホテイケシデオキノコムシ Ish, Iriinexpecta Löbl, 1981: 237 コホテイケシデオキノコムシ（和名改称）HON
japonica Reitter, 1880: 49 (Toxidium)トビイロホテイケシデオキノコムシ（和名改称）HON, KYU
pecki Lobl, 1981: 238 ペックホテイケシデオキノコムシ（和名改称）HON, SHI, South Korea
smetanai Löbl, 1981: 233 スメタナホテイケシデオキノコムシ（和名改称）HON, SHI
sunadai Hoshina & Sugaya, 2003: 36 オキナワホテイデオキノコムシ Okn
variabilis Löbl, 1981: 234 カワリホテイケシデオキノコムシ（和名改称）HON
Genus Scaphicoma Motschulsky, 1863 オオナガケシデオキノコムシ属
hiranoi Hoshina, 2008a: 57 ヤエヤマナガケシデオキノコムシ Ish2013 The Kyushu

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Yasutoshi Shibata, Munetoshi Maruyama, Hideto Hoshina, Toshio Kishimoto, Shun-Ichiro Naomi, Shûhei Nomura, Volker Puthz, Takashi Shimada, Yasuaki Watanabe, Shûhei Yamamoto

Genus Scaphoxium Löbl, 1979 ナガケシデオキノコムシ属
japonicum Löbl, 1981: 242 ナガケシデオキノコムシ SHIhiranoi Hoshina, 2008a: 57 ヤエヤマナガケシデオキノコムシ Ishkunigamiense Hoshina & Sugaya, 2003: 39 クニガミナガケシデオキノコムシ Okn
saigoi Hoshina, 2008c: 2 サイゴウナガケシデオキノコムシ Amm
taiwanum Löbl, 1980: 121 タイワンナガケシデオキノコムシ Ish, Iri; Taiwan, China, Thailand, Nepal, India
Genus Toxidium LeConte, 1860 ホソケシデオキノコムシ属
aberrans Achard, 1923: 119 ヤマトホソケシデオキノコムシ HON, SHI, Amm

Subfamily Piestinae Erichson, 1839 ヒラタハネカクシ亜科（３属19種）Genus Eupiestus Kraatz, 1859 ミナミヒラタハネカクシ属（和名新称）
iriomotensis Naomi, 2011b: 60 ワタナベミナミヒラタハネカクシ Iri
Genus Piestoneus Sharp, 1889 オオヒラタハネカクシ属（和名新称）
lewisii Sharp, 1889: 465 ルイスオオヒラタハネカクシ（和名改称）HOK, HON, Oki, SHI, KYU; South Korea= deboiseae Naomi, 1995e: 772
monticola Naomi, 1995e: 773 ミヤマオオヒラタハネカクシ（和名新称）HON
oharai Naomi, 1995e: 775 オオハラオオヒラタハネカクシ（和名新称）HOK, HON, SHI, KYU, Tsm
Genus Siagonium Kirby & Spence, 1815 ヤマヒラタハネカクシ属（和名新称）
Species group of S. debile Sharp ヒメヒラタハネカクシ種群
debile Sharp, 1889: 464 ヒメヒラタハネカクシ HON, SHI
deboiseae Naomi, 1994b: 152 デボアヒメヒラタハネカクシ（和名新称）HON
hammondi Naomi, 1994b: 150 ハモンドヒメヒラタハネカクシ（和名新称）SHI, KYU
puthzi Naomi, 1994b: 149 プーツヒメヒラタハネカクシ（和名新称）HOK, KYU
Species group of S. gracile Sharp ホソヒラタハネカクシ種群
crowsoni Naomi & Nakane, 1995: 7 クローソンホソヒラタハネカクシ（和名新称）HON
densepunctatum Naomi & Nakane, 1995: 8 リュウキュウホソヒラタハネカクシ（和名新称）Okn
emotoi Naomi, 2006c: 41 エモトホソヒラタハネカクシ（和名改称）HON
gracile Sharp, 1889: 464 ホソヒラタハネカクシ HOK, HON, Iz-Hch, SHI, KYU, Yak= kojimai Takai & Nakane, 1985: 98
kurinoense Takai & Nakane, 1985: 103 クリノホソヒラタハネカクシ（和名新称）KYU
yamashitai Takai & Nakane, 1985: 97 ヤマシタホソヒラタハネカクシ（和名改称）HON, SHI= nikkoense Takai & Nakane, 1985: 99
Species group of S. nobile Sharp セミゾヒラタハネカクシ種群
incertum Takai & Nakane, 1985: 101 キノカワセミゾヒラタハネカクシ（和名新称）HON

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Catalogue of Japanese Staphylinidae (Insecta: Coleoptera)

nakanei Naomi, 1995b: 149 ナカネセミゾヒラタハネカクシ HOK
nobile Sharp, 1889: 463 セミゾヒラタハネカクシ HON, SHI, KYU
Species group of S. vittatum Sharp モンヒラタハネカクシ種群
haroldi Weise, 1879: 148 アカモンヒラタハネカクシ（和名改称）HON, SHI, KYU, Iz-My, Iz-Mk, Yak; Taiwan
vittatum Fauvel, 1875: i モンヒラタハネカクシ（和名改称）HOK; Taiwan, South Korea, Russia (Siberia)

Subfamily Osoriinae Erichson, 1839 ツツハネカクシ亜科（14属28種）
Tribe Eleusinini Sharp, 1887 ウスバツツハネカクシ族（和名新称）
Genus Eleusis Laporte, 1835 ウスバツツハネカクシ属（和名新称）
coarctata Sharp, 1889: 466 オオウスバツツハネカクシ（和名改称）HOK, HON, Oki, SHI, KYU, Tsm, Amk; South
Korea
humilis Erichson, 1840: 839 (Isomalus) ホソウスバツツハネカクシ（和名改称）Iz-My, KYU, Yak, Ish, Iri; Taiwan,
China, Bhutan , SE Asia, North America, South America= subtilis Sharp, 1889: 467
quadridens Fauvel, 1904: 84トゲムネウスバツツハネカクシ（和名新称）Amm; SE Asia
terminata Fauvel, 1869: 494 キウスバツツハネカクシ（和名改称）Og, Okn, Myk, Ish, Iri; Taiwan, SE Asia, Sri
Lanka= kraatzi Fauvel, 1878: 207
Genus Zeoleusis Steel, 1950 ネッタイツツハネカクシ属（和名新称）
fusciceps Kraatz, 1859: 184 (Isomalus) ヒメネッタイツツハネカクシ（和名新称）Okn, Iri; India, Singapore, Indonesia,
Philippines, SE Asia
Tribe Leptochirini Sharp, 1887 オニツツハネカクシ族（和名新称）
Genus Priochirus Sharp, 1887 ツノツツハネカクシ属（和名新称）
Subgenus Euleptarthrus Jacobson, 1908
japonicus Sharp, 1889: 468 クロツヤツノツツハネカクシ（和名改称）HON, Sad, SHI, KYU, SKs, Okn; Taiwan,
China= freyi Bernhauer, 1941: 209
masahiroi Naomi, 1996a: 6 サイトウツノツツハネカクシ（和名新称）HON, KYU
Subgenus Leiochirus Greenslade, 1971
yasutoshii Naomi, 1996a: 9 シバタツノツツハネカクシ（和名新称）Amm
Subgenus Paraborolinus Nakane & Sawada, 1960
yakushimensis Herman, 2001: 41 ヤクツノツツハネカクシ（和名改称）KYU, Yak, Kue, Tan= bicorrnis Nakane & Sawada, 1960: 121

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Yasutoshi Shibata, Munetoshi Maruyama, Hideto Hoshina, Toshio Kishimoto, Shun-Ichiro Naomi, Shûhei Nomura, Volker Puthz, Takashi Shimada, Yasuaki Watanabe, Shûhei Yamamoto

Subgenus Plastus Bernhauer, 1903
kimurai Naomi, 1996a; 14 キムラツノツツハネカクシ（和名新称）Okn
kishimotoi Naomi, 2004b: 99 キシモトツノツツハネカクシ（和名改称）Ish
Tribe Osoriini Erichson, 1839 ツツハネカクシ族
Genus Atopocnemius Bernhauer, 1914 ツヤツツハネカクシ属（和名新称）
nomurai Naomi, 1986c：40 ノムラツヤツツハネカクシ（和名改称）Amm
Genus Mimogonellus Fagel, 1955 クチキツツハネカクシ属（和名新称）
japonicus Naomi, 1987b: 15 (Saegerius) ナガクチキツツハネカクシ（和名改称）HON, KYU, Tsm, T-Yok, Amm,
Tok, Oke, Ishyasutoshii Naomi, 1987b: 17 (Saegerius) シバタクチキツツハネカクシ（和名改称）KYU
Genus Mimogonus Fauvel, 1903 オチバツツハネカクシ属（和名新称）
microps Sharp, 1889: 412 (Osorius) ヒメオチバツツハネカクシ（和名改称）HON, KYU; Taiwan; Korea, China
Genus Osorius Guerin-Meneville, 1829 ツツハネカクシ属
angustulus Sharp, 1874a: 89 フトツツハネカクシ HOK, HON, Iz-Mk, Oki, SHI, KYU, Kokn, Tan, Tsm, Kue; South
Korea
taurus Sharp, 1889: 411 ツノフトツツハネカクシ HON, Iz-Mk, Iz-My, SHI, KYU, Tsm, Yak, Kue; South Korea,
China, Oriental Region
Tribe Thoracophorini Reitter, 1909 ホソツツハネカクシ族（和名新称）
Subtribe Clavilispinina Newton & Thayer, 1992 ウスイロツツハネカクシ亜族（和名新称）
Genus Clavilispinus Bernhauer, 1926 ウスイロツツハネカクシ属（和名新称）
exiguus Erichson, 1840: 830 (Lispinus) ミナミウスイロツツハネカクシ（和名改称）Og, KYU, MIw, Yak, Oke, Amm;
Taiwan, China, Pan Tropics
Subtribe Lispinina Bernhauer & Schubert, 1910 ヒメツツハネカクシ亜族（和名新称）
Genus Lispinus Erichson, 1839 ヒメツツハネカクシ属（和名新称）
jambar Naomi, 1996c: 763 ヤンバルヒメツツハネカクシ（和名新称）T-Nak, Amm, Okn, Ish, Iritakahashii Naomi, 2012a: 150; タカハシヒメツツハネカクシ（和名新称）Ish
Genus Nacaeus Blackwelder, 1942 モリツツハネカクシ属（和名新称）
iriomotensis Naomi, 1997d: 137 イリオモテモリツツハネカクシ（和名新称）Okn, Irijaponicus Cameron, 1930: 181 (Lispinus) ニホンモリツツハネカクシ（和名新称）HON, Oki, KYU, Amm, Irilongulus Sharp, 1874a: 101 (Lispinus) ナガモリツツハネカクシ（和名改称）HON, Iz-My, Iz-Mk, Iz-Hch, Oki, SHI,

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Catalogue of Japanese Staphylinidae (Insecta: Coleoptera)

KYU, Tsm, SKs, Yak; South Korea, Taiwan
newtoniellus Naomi, 1997d: 135 ニュートンモリツツハネカクシ（和名新称）Oke, Okn
Genus Neolosus Blackwelder, 1942 カクムネツツハネカクシ属（和名新称）
aper Sharp, 1889: 466 (Lispinus) ニホンカクムネツツハネカクシ（和名改称）HON, SHI, KYU; South Korea
Subtribe Thoracophorina Reitter, 1909 ホソツツハネカクシ亜族（和名新称）
Genus Tetrapleurus Bernhauer, 1914 ハネスジツツハネカクシ属（和名新称）
paralellus Bernhauer, 1917: 41 ミナミハネスジツツハネカクシ（和名新称）Ish; Taiwan
Genus Thoracophorus Motschulsky, 1837 ホソツツハネカクシ属（和名新称）
certatus Sharp, 1889: 465 ヤマトホソツツハネカクシ（和名改称）HON, SHI, KYU, Iz-Mk, Iz-Hch, Og-Hh, Oki,
Yak, T-Nak, Amm, Iri; Taiwan
Incertae cedis (subtribe?) 亜族所属不明
Genus Allotrochus Fagel, 1955 ミナミツツハネカクシ属（和名新称）
takahashii Naomi & Irmler, 2012: 107 タカハシミナミツツハネカクシ（和名新称）Ish

Subfamily Oxytelinae Fleming, 1821 セスジハネカクシ亜科（14属86種）
Tribe Blediini Ádám, 2001
Genus Bledius Leach, 1819 カワベハネカクシ属
Subgenus Astycops Thomson, 1859
sellatus Sharp, 1889: 413 キベリイブシカワベハネカクシ（和名新称）HOK, Rsr, HON; North Korea, China
Subgenus Barygus Schiodte, 1866
japonicus Bernhauer, 1936: 305 ヤマトカワベハネカクシ（和名新称）Japan; China
obihiroensis Nakane, 1963: 21 アカバカワベハネカクシ HOK
pallipes Gravenhorst, 1806: 197 (Oxytelus) キアシカワベハネカクシ HOK; China, Russia, Europe
vitaensis Bernhauer, 1938a: 25 ビタカワベハネカクシ（和名新称）Japan
Subgenus Bledius Leach, 1819
gigantulus Bernhauer, 1922: 224 キョクトウオオツノハネカクシ（和名新称）HON, SHI; Taiwan, China, Mongolia
gyotokui Nakane, 1963: 21 チクゴオオツノハネカクシ（和名改称）KYU
salsus Miyatake, 1963: 106 オオツノハネカクシ HON, Oki, SHI, KYU; South Korea, China,
setonis Miyatake, 1967: 94 [RN] コツノハネカクシ SHI= rotundicollis Miyatake, 1963: 109 [HN]2013 The Kyushu

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Yasutoshi Shibata, Munetoshi Maruyama, Hideto Hoshina, Toshio Kishimoto, Shun-Ichiro Naomi, Shûhei Nomura, Volker Puthz, Takashi Shimada, Yasuaki Watanabe, Shûhei Yamamoto

tricornis Herbst, 1784: 149 (Staphylinus) ミツノハネカクシ（和名新称）Japan; China, Vietnam, Mongolia, Iran,
Turkey, Syria, Afganistan, C Asia, Algeria, Europe, Russia, Caucasus
yezoensis Nakane, 1963: 21 エゾオオツノハネカクシ（和名改称）HOK, Rsr; South Korea
Subgenus Hesperophilus Stephens, 1829
cribratus Sharp, 1889: 413 ツマキカワベハネカクシ HON
fragilis Sharp, 1874a: 90 キバネカワベハネカクシ（和名新称）KYU; Taiwan
obtusus Sharp, 1889: 413 ツヤムネカワベハネカクシ HON; China
Subgenus Pucerus Mulsant & Rey, 1878
curvicornis Sharp, 1889: 412 キベリカワベハネカクシ HOK, HON, Oki, Tan, Tsm; North Korea, Chinaorphanus Sharp, 1874a: 91 ヒゲナガカワベハネカクシ HON, KYU; Taiwan, China
Incertae sedis (subgenus?) 亜属不明
lucidus Sharp, 1874a: 90 ツマグロカワベハネカクシ HON, KYU, T-Aks; Taiwan, China, Oriental Region, New
Guinea, Australia, Madagascar, Congo, Kenya
Tribe Coprophilini Heer, 1839
Genus Coprophilus Latreille, 1829 キノカワハネカクシ属（和名新称）
adachii Y. Watanabe & Shibata, 1961c: 43 (Elonium) アカバキノカワハネカクシ HOK, HON
impressus Sharp, 1889: 415 セミゾキノカワハネカクシ HOK, HON, SHI= fulvomaculatus Adachi, 1935: 125
simplex Sharp, 1889: 415 キノカワハネカクシ HOK, HON, SHI
Genus Coprotrichus Hayashi, 2005 ニセキノカワハネカクシ属（和名新称）
kameii Hayashi, 2005: 65 ニセキノカワハネカクシ HON
Tribe Euphanini Reitter, 1909
Genus Deleaster Erichson, 1839 ハラビロハネカクシ属（和名新称）
yokoyamai Adachi, 1935: 126 ハラビロハネカクシ HOK, HON
Genus Syntomium Curtis, 1828ドウガネアナバケハネカクシ属（和名新称）
japonicum Y.Watanabe & Shibata, 1960: 103ドウガネアナバケハネカクシ HON, SHI
Tribe Oxytelini Fleming, 1821
Genus Anotylus Thomson, 1859 ヒメセスジハネカクシ属（和名新称）
amicus Bernhauer, 1923a: 122 (Oxytelus) コバネアシベセスジハネカクシ HON, SHI, KYU, Ish, Iri; Taiwan, Korea,
Chinaantennarius Bernhauer, 1907: 377 (Oxytelus) アバタセスジハネカクシ HON, SHI
cognatus Sharp, 1874a: 94 (Oxytelus) セスジハネカクシ HOK, Reb, HON, Aws, Iz-Hch, SHI, KYU; Chs, South2013 The Kyushu

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Catalogue of Japanese Staphylinidae (Insecta: Coleoptera)

Korea, China
crassicornis Sharp, 1874a: 92 (Oxytelus) ヒゲブトセスジハネカクシ HON, Iz-Hch, SHI, KYU; Taiwan, South Korea,
North Korea, China
funebris Bernhauer, 1907: 376 (Oxytelus) イブシセスジハネカクシ HON, Iz-Hch, SHI; China
glaerosus Wollaston, 1854: 610 (Oxytelus) ナンヨウヒメヒラタセスジハネカクシ（和名新称）Og, KIw; Taiwan,
China, Oriental Region, USA, C America, West Indies, Africa
gregarius Sharp, 1889: 414: 87 (Oxytelus) イブシヒメセスジハネカクシ（和名新称）HON, Taiwan
hosodai T. Ito, 1991b; 87 ホソダセスジハネカクシ HON
japonicus Cameron, 1930: 183 (Oxytelus) ヒラタセスジハネカクシ HON, Iz-Mk, Oki, SHI, SKs, Tanlaticornis Sharp, 1874a: 95 (Oxytelus) ヒメクロセスジハネカクシ HON, Iz-Mk, Iz-Hch, SHI, KYU, Tsm; China
latiusculus Kraatz, 1859: 176 (Oxytelus) チビクロセスジハネカクシ HOK, Reb, Rsr, HON, SHI; Taiwan, China,
Oriental Region, Africa, Madagascar, New Guinea, Australia, Middle East, Cyprus, Sicily= ganglbaueri Bernhauer, 1907: 375
lewisius Sharp, 1874a: 95 ルイスツヤセスジハネカクシ（和名改称）HON, Iz-My, Iz-Mk, Iz-Hch, SHI, KYU, Tsm,SKs, Tan, Yak, Amm, Okn, Ihy; Taiwan, China
marginatus Weise, 1877a: 96 (Oxytelus) オオツヤセスジハネカクシ（和名新称）HON, SHI, KYU= sharpianus Cameron, 1930: 184 (Oxytelus)
mimulus Sharp, 1874a: 94 シワバネセスジハネカクシ（和名改称）HOK, HON, Iz-Hch, Oki, SHI, KYU; China,
Russia, USA, Chile
nitidifrons Wollaston, 1871: 411 (Oxytelus) ツヤセスジハネカクシ（ 和 名 新 称 ）Japan; China, India, Oriental
Region, St. Helena, Palau, Hawaii, Madagascar, Açores, Madeira
pusillimus, Kraatz, 1859: 177 (Oxytelus) ツヤヒメセスジハネカクシ（ 和 名 新 称 ）HOK; Taiwan, China, Nepal,
Pakistan, Oriental Region, Africa, Madagascar, Australia, Fiji
= pygmaeus Kraatz, 1859: 176 (Oxytelus) [HN]
= sauteri Bernhauer, 1907: 375 (Oxytelus)
vicinus Sharp, 1874a: 94 (Oxytelus)トビイロセスジハネカクシ HOK, HON, SHI, KYU; China
Genus Aploderus Stephens, 1833 ムネミゾハネカクシ属（和名新称）
watanabei Shimada, 2002: 233 ワタナベムネミゾハネカクシ HON
Genus Carpelimus Leach, 1819 ヒメユミセミゾハネカクシ属（和名新称）
Subgenus Boopinus Klima, 1904
siamensis Fauvel, 1886: 144 (Trogophloeus) キバネヒメユミセミゾハネカクシ（和名改称）HON, Oki, SHI, Yak,
Okn; China, Oriental Region, Madagascar, Australia
Subgenus Carpelimus Leach, 1819
angusticollis Bernhauer, 1907: 373 (Trogophloeus)トビイロヒメユミセミゾハネカクシ（和名新称）HON, KYU, SKs,
MDt
= sharpianus Cameron, 1933: 169 (Trogophloeus)
biimpressus Cameron, 1933: 168 (Trogophloeus) ムナビロヒメユミセミゾハネカクシ（和名新称）HON, Oki, SHI;
Tsm; China

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Yasutoshi Shibata, Munetoshi Maruyama, Hideto Hoshina, Toshio Kishimoto, Shun-Ichiro Naomi, Shûhei Nomura, Volker Puthz, Takashi Shimada, Yasuaki Watanabe, Shûhei Yamamoto

pseudovagus Gildenkov, 2002: 1467 マガイヒメユミセミゾハネカクシ（和名新称）HON
sedatus Sharp, 1889: 417 (Trogophloeus) ホソヒメユミセミゾハネカクシ（和名新称）HON, KYU; China
vagus Sharp, 1889: 417 (Trogophloeus) ニセヒメユミセミゾハネカクシ（和名改称）HON, Oki, SHI, KYU, Tsm, Iri,Okn, KDt, MDt; Taiwan, South Korea, China
Subgenus Paratrogophloeus Hatch, 1957
longicollis Bernhauer, 1907: 373 (Trogophloeus) イブシニセユミセミゾハネカクシ（和名新称）HON, SHI
porphyreus Gildenkov, 2002: 1467 コイブシニセユミセミゾハネカクシ（和名新称）HON
Subgenus Troginus Mulsant & Rey, 1878
despectus fujiensis Gildenkov, 2002: 1474 フジツツニセユミセミゾハネカクシ（和名新称）HON; China
exiguus Erichson, 1839b: 604 (Trogophloeus) チビツツニセユミセミゾハネカクシ（和名改称）HON, SHI, KYU,
Tsm, Tan, Yak; China, Mongolia, Oriental Region, Australia, Europe, Russia, West India, Africa
niponensis Gildenkov, 2002: 1469 ヤマトツツニセユミセミゾハネカクシ（和名新称）HON
Genus Ochthephilus Mulsant & Rey, 1856 ヨコセミゾハネカクシ属（和名新称）
Subgenus Ochthephilus Mulsant & Rey, 1856
antennatus Y. Watanabe & Shibata, 1961a: 7 (Ancyrophorus) ヒゲブトヨコセミゾハネカクシ HOK, HON; South
Korea
japonicus Y. Watanabe & Shibata, 1961a: 6 (Ancyrophorus) ヒゲナガヨコセミゾハネカクシ HOK, HON, SHI
laevis Y. Watanabe & Shibata, 1961a: 8 (Ancyrophorus) ツヤヨコセミゾハネカクシ HOK, HON
vulgaris Y. Watanabe & Shibata, 1961a: 7 (Ancyrophorus) ナミヨコセミゾハネカクシ HOK, HON, SHI; South Korea
Genus Oxytelopsis Fauvel, 1895 ムネトゲセスジハネカクシ属（和名新称）
shibatai T. Ito, 1987: 75 ムネトゲセスジハネカクシ HON, SHI, KYU
Genus Oxytelus Gravenhorst, 1802 セスジハネカクシ属
Subgenus Oxytelus Gravenhorst, 1802
bengalensis Erichson, 1840: 789 クロズマグソセスジハネカクシ HON, SHI, KYU, Okn; Taiwan, South Korea(Cheju-do), China, Oriental Region=opacifrons Sharp, 1874a: 93
incisus Motschulsky, 1858a: 504 アカセスジハネカクシ HON, Iz-Hch, Og, SHI, KYU, Yak, Amm, Okn, MDt;
Taiwan, China, South Korea, Pan Tropics, Iran, Nepal, Pakistan, Yemen, Canada= laevior Sharp, 1874a: 92
iners Weise, 1877a: 94 ヒメキバネセスジハネカクシ（和名新称）HON, Yakmigrator Fauvel, 1904: 100 ヒメアカセスジハネカクシ HON, Oki, KYU, Tsm, T-Aks, Okn; Taiwan, China, South
Korea, Pan Tropics, Europe=akazawensis Bernhauer, 1907: 379
nigriceps Kraatz, 1859: 171 クロズセスジハネカクシ HON, SHI, KYU, Yak, Amm, Okn; Taiwan, South Korea,
China, Oriental Region, New Guinea
piceus Linnaeus, 1767: 686 (Staphylinus) キバネセスジハネカクシ HOK, HON, Sad, Iz-My, Iz-Hch, Og, SHI,KYU, T-Nak; North Korea, South Korea, China, Oriental Region, Europe, Africa

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Catalogue of Japanese Staphylinidae (Insecta: Coleoptera)= japonicus Motschulsky, 1861: 10
Subgenus Tanycraerus Thomson, 1859
houomontis T. Ito, 1994e: 333 [RN] ホウオウセスジハネカクシ HOK, HON= montivagus T. Ito, 1994a: 42 [HN]
jessoensis Bernhauer, 1907: 378 エゾセスジハネカクシ（和名新称）HOK
puncticeps Kraatz, 1859: 176 カタツノセスジハネカクシ Okn, Ish, Iri; Taiwan, China, SE Asia, India, Sri Lanka,
Africa, Madagascar
takahashii T. Ito, 1994a: 44 タカハシセスジハネカクシ HON; China, South Korea
varipennis Kraatz, 1859: 172 ニセキバネセスジハネカクシ Japan, Oki, Tsm; China, South Korea, China, India,
Sri Lanka
Genus Platystethus Mannerheim, 1830 ヒメカワベハネカクシ属（和名新称）
Subgenus Craetopycrus Tottenham, 1939
alutaceus Thomson, 1861: 123 ヒガワヒメカワベハネカクシ（和名新称）Japan; Russia, Mongolia, Afganistan,
Europe, USA
cornutus Gravenhorst, 1802: 109 (Oxytelus) ヘリグロヒメカワベハネカクシ（和名新称）HOK; China, India, Nepal,
Oriental Region, Palearctic Region
operosus Sharp, 1874a: 91 クロヒメカワベハネカクシ HOK, HON, Iz-My, Iz-Hch, SHI, Kue; Taiwan, South Korea,
North Korea, China
Incertae sedis (subgenus?) 亜属不明
americanus Erichson, 1840: 784 クロツヤヒメカワベハネカクシ（和名改称）HON; Hawaii, Canada, North America,
Mexicojaponicus Bernhauer, 1907: 374 ヤマトツヤヒメカワベハネカクシ（和名新称）HON
laesicollis Lokay, 1918b: 21 ニセクロツヤヒメカワベハネカクシ（和名新称）HON
quedenfeldti Weise, 1879: 147 キバネツヤヒメカワベハネカクシ（和名新称）HON
Genus Thinobius Kiesenwetter, 1844 スソハダケハネカクシ属（和名新称）
Subgenus Platyderothinophilus Scheerpeltz, 1959
ootsukai Naomi, 1995a: 22 オオツカスソハダケハネカクシ HON
yabakeinis Naomi, 1995a: 23 ニシスソハダケハネカクシ（和名新称）SHI, KYU
Subgenus Thinobius Kiesenwetter, 1844
japonicus Naomi, 1995a: 20 ヤマトスソハダケハネカクシ HON
kuroshio Sawada, 1971d: 327 (Yoshiityphlus) クロシオスソハダケハネカクシ（和名新称）HON
nomurai Naomi, 1995a: 18 エゾスソハダケハネカクシ（和名新称）HOK
takashii Naomi, 2004d: 156 ツシマスソハダケハネカクシ（和名新称）Tsm
Genus Thinodromus Kraatz, 1858 ユミセミゾハネカクシ属（和名新称）
Subgenus Amisammus Gozis, 1886
japonicus Cameron, 1933: 183 (Trogophloeus) ヤマトニセユミセミゾハネカクシ HON, Oki, SHI, Tsm, SKs; South
Korea (Cheju-do)

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Yasutoshi Shibata, Munetoshi Maruyama, Hideto Hoshina, Toshio Kishimoto, Shun-Ichiro Naomi, Shûhei Nomura, Volker Puthz, Takashi Shimada, Yasuaki Watanabe, Shûhei Yamamoto

Subgenus Thinodromus Kraatz, 1858
deceptor Sharp, 1889: 416 (Trogophloeus) アカアシユミセミゾハネカクシ HON, SHI, KYU, Tsm; China, South
Korea
proprius Gildenkov, 2003: 584 コバネユミセミゾハネカクシ（和名新称）HON
pseudoeminens Gildenkov, 2003: 586 シンニセユミセミゾハネカクシ（和名新称）HON
sericatus Sharp, 1889: 416 (Trogophloeus) ユミセミゾハネカクシ HOK, HON, SHI, KYU; South Korea= eminens Sharp, 1889: 416 (Trogophloeus)

Subfamily Oxyporinae Fleming, 1821 オオキバハネカクシ亜科（２属23種）Genus Oxyporus Fabricius, 1775 オオキバハネカクシ属
aokii R. Dvořák, 1956b: 60 アオキオオキバハネカクシ（和名新称）HON
basicornis Cameron, 1930: 184 ヒメオオキバハネカクシ HON, SHI, KYU; South Korea
basiventris Jarrige, 1948: 40 ネアカオオキバハネカクシ HOK, HON, SHI, KYU; South Korea, China
crocatus Fauvel, 1901a: 2 アカチャオオキバハネカクシ（和名新称）HON; South Korea
germanus Sharp, 1889: 409 イクチオオキバハネカクシ HON, Oki, SHI; South Korea, North Korea, China
imitator Nakane & Sawada, 1956a: 119 マガイオオキバハネカクシ（和名新称）HON
japonicus Sharp, 1889: 407 オオキバハネカクシ HOK, HON, SHI, KYU; South Korea, North Korea= nagayamai Nakane & Sawada, 1956a: 118
lewisi Cameron, 1930: 184 ルイスオオキバハネカクシ（和名新称）HON
maculiventris Sharp, 1889: 408 アカバオオキバハネカクシ HON, SHI; China= flavescens Nakane & Sawada, 1956a: 117
niger Sharp, 1889: 407 クロオオキバハネカクシ HOK, HON, SHI, KYU; South Korea
parcus Sharp, 1889: 408 オオズオオキバハネカクシ HOK, HON, Oki, SHI, KYU; South Korea= jozanus Nakane & Sawada, 1956a: 117= nishimurai Nakane & Sawada, 1956a: 117
rufus osawai Nakane & Sawada, 1956a: 120 ムネアカオオキバハネカクシ HON, SHI, KYU; South Korea
tosensis Takeuchi, 1941: 151トウセムネアカオオキバハネカクシ（和名新称）HON
triangulum Sharp, 1889: 407 ハスオビオオキバハネカクシ HOK, HON, Oki, SHI, KYU; South Korea= pullus Nakane & Sawada, 1956a: 117
Genus Pseudoxyporus Nakane & Sawada, 1956 ニセオオキバハネカクシ属（和名新称）
angusticeps Bernhauer 1938a: 33 (Oxyporus) ホソニセオオキバハネカクシ（和名改称）HON, SHI; China
biguttatus Sharp, 1889: 411 (Oxyporus) カドムネオニセオキバハネカクシ（和名改称）HON, SHI= aizuanus Nakane & Sawada, 1956a: 122= varicolor Nakane & Sawada, 1956a: 122= yanoi Nakane & Sawada, 1956a: 122
cyanipennis Kirshenblat, 1938: 528 (Oxyporus) ルリバネニセオオキバハネカクシ（和名改称）HOK; Russia

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Catalogue of Japanese Staphylinidae (Insecta: Coleoptera)(South Ussuri)
gnatho Sharp, 1889: 410 (Oxyporus) マルモンニセオオキバハネカクシ（和名改称）HON, SHI, KYU
hoplites Sharp, 1889: 410 (Oxyporus) ヨコモンニセオオキバハネカクシ（和名改称）HON
humeralis Sharp, 1889: 410 (Oxyporus) カタモンニセオオキバハネカクシ（和名改称）HON
longipes Sharp, 1889: 409 (Oxyporus) アシナガニセオオキバハネカクシ（和名改称）HON, SHI; South Korea,
China= kurosawai Nakane & Sawada, 1956a: 121= narumii Nakane & Sawada, 1956a: 121
masatoi T. Ito, 1999a: 255 ツルギニセオオキバハネカクシ（和名改称）SHI
sakagutii Nakane & Sawada, 1956a: 122 ハネビロニセオオキバハネカクシ（和名改称）HON, SHI, KYU= nigrofemoralis Nakane & Sawada, 1956a: 122

Subfamily Megalopsidiinae Leng, 1920 メダカオオキバハネカクシ亜科（１属４種）
Genus Megalopinus Eichelbaum, 1915 メダカオオキバハネカクシ属
Subgenus Megalopinus Eichelbaum, 1915
flavomaculatus Naomi, 1986b: 351 キモンメダカオオキバハネカクシ Tsmhirashimai Naomi, 1986b: 348 ヒラシマメダカオオキバハネカクシ Amm, Tok, Okn, Kum; Taiwan
japonicus Nakane, 1957: 53 (Megalopsidia) ナカネメダカオオキバハネカクシ（和名改称）HON, SHI, Tsm, Ishtomishimai Naomi, 1996c: 69 キボシメダカオオキバハネカクシ KYU

Subfamily Steninae MacLeay, 1825 メダカハネカクシ亜科(1, 2)（２属258種８亜種）⑴

325th contribution to the knowledge of Steninae (V. Puthz)⑵

46th contribution to the knowledge of Steninae (S.-I. Naomi)
メダカハネカクシ亜科に関しては，種と種群に関する系統的に配列したリストを準備することができたので，甲虫学者

はそれらに関する仮説を利用して，この亜科の地域ファウナを，明瞭に研究できるようになる．私達は，より一層の研究・調査により，日本列島から知られたメダカハネカクシ属およびヒョウタンメダカハネカクシ属の種に関する系統的体系の仮説が検証されることを希望する．
As far as the clade Steninae is concerned, a phylogenetically arranged list of the species and species groups can beprepared such that coleopterists can study specifically the regional fauna of the subfamily, using the hypothesis about
them. We hope that our hypothesis on the phylogenetic system of the *Stenus *and *Dianous *species from theJapanese Archipelago will be tested by further investigations.
Genus Dianous Leach, 1819 ヒョウタンメダカハネカクシ属

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Yasutoshi Shibata, Munetoshi Maruyama, Hideto Hoshina, Toshio Kishimoto, Shun-Ichiro Naomi, Shûhei Nomura, Volker Puthz, Takashi Shimada, Yasuaki Watanabe, Shûhei Yamamoto

moritai Naomi, 1997a: 1 モリタヒョウタンメダカハネカクシ（和名新称）Amm, Tokuedai Naomi, 2010b: 41 ウエダヒョウタンメダカハネカクシ Amm
shibatai Sawada, 1960: 10 オオヒョウタンメダカハネカクシ HON
iwakisanus Watanabe, 1984: 134 ツガルヒョウタンメダカハネカクシ HON
gongen Watanabe, 1984: 131 ホソヒョウタンメダカハネカクシ HON
viridicatus Naomi, 1997a: 3 ミドリヒョウタンメダカハネカクシ（和名新称）HON
japonicus Sawada, 1960: 9 ニホンヒョウタンメダカハネカクシ（和名改称）HON, SHI, KYU= septentrionalis Naomi, 1988d: 50= morimotoi Naomi, 1988d: 48
amamiensis Sawada, 1960: 11 アマミヒョウタンメダカハネカクシ Amm
yoshidai Naomi, 1988d: 51 ヨシダヒョウタンメダカハネカクシ Iri
Genus Stenus Lateille, 1797 メダカハネカクシ属
Species group of S. comma LeConte オオフタホシメダカハネカクシ種群
biguttatus (Linnaeus), 1758: 422 ヒメフタホシメダカハネカクシ HOK; China, Palearctic Region
alienus Sharp, 1874a: 81 ホソフタホシメダカハネカクシ HOK, HON, Oki, SHI, KYU; Taiwan, South Korea,
North Korea, China, Russia
anthracinus Sharp, 1889: 326 スジナガメダカハネカクシ（和名改称）HOK, HON, SHI, KYU; China= yanoi Sawada & Nakane, 1954: 8
kobensis Cameron, 1930: 205 ナガメダカハネカクシ HOK, HON, Oki, SHI, KYU
comma LeConte, 1863: 50 オオフタホシメダカハネカクシ HOK; North Korea, China, Russia, Holoarctic Region
tenuipes Sharp, 1874a: 80 フタホシメダカハネカクシ HOK, HON, SHI, KYU; South Korea, China
macies Sharp, 1874a: 83 ドウボソナガメダカハネカクシ（和名改称）HOK, HON, Iz-My, Iz-Mk, Iz-Hch, Oki, SHI,KYU, Tsm, Yak; China
Species group of S. clavicornis (Scopoli) アラハダナガメダカハネカクシ種群
juno Paykull, 1789: 33 ミスジオオメダカハネカクシ（和名新称）HOK, HON; China, Russia, Europe, N America
indagator Eppelsheim, 1887b: 428 オオメダカハネカクシ HOK, HON, SHI; China, Russia (W Siberia)= lentus Sharp, 1889: 326
yamato Naomi, 1988e: 57 ヤマトナガメダカハネカクシ（和名改称）HON, SHI
lewisius Sharp, 1874a: 82 ルイスナガメダカハネカクシ（和名改称）HOK, HON, SHI, KYU= gatenpio Naomi, 1990c: 99
sauteri Bernhauer, 1907: 379 ザウテルナガメダカハネカクシ（和名改称）HOK, HON, SHI
nishikawai Naomi, 1988e: 58 ニシカワナガメダカハネカクシ（和名改称）HOK, HON, KYU
amurensis Eppelsheim, 1886: 43 アムールナガメダカハネカクシ（和名新称）HOK; China, Russia, E Asia
calliceps Bernhauer, 1916c: 28 ヤマヒザグロメダカハネカクシ（和名改称）HON, SHI, KYU; China
disterminus Puthz, 1993b: 179 ヒメヒザグロメダカハネカクシ（和名新称）HON
distans Sharp, 1889: 327 ヒザグロメダカハネカクシ HOK, HON, SHI, KYU; South Korea, North Korea, Taiwan,
China, Russia
formosanus Benick, 1914: 285 タイワンナガメダカハネカクシ（和名新称）Yon; Taiwan, China
clavicornis Scopoli, 1763: 100 (Staphylinus) アラハダナガメダカハネカクシ HOK; South Korea, China, Russia,

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Catalogue of Japanese Staphylinidae (Insecta: Coleoptera)

Europe, N America
laborator Sharp, 1889: 327 ニセナガメダカハネカクシ（和名改称）HON
takedai Naomi, 1990c: 103 タケダナガメダカハネカクシ（和名新称）HON
Species group of S. humilis Erichson ホソヒメメダカハネカクシ種群
japonicus Sharp, 1874a: 84 ヒメメダカハネカクシ HOK, HON, Iz-Hch, SHI, KYU, Yak; China, Russia= nipponensis Cameron, 1930b: 205
mammops Casey, 1884: 97 キタヒメメダカハネカクシ（和名改称）HOK, Rsr, Tsm; China, Russia, E Asia, N America
hammondi Puthz, 1970: 31 ハモンドヒメメダカハネカクシ（和名新称）HOK; China, Russia (Siberia)
expugnator Ryvkin, 1987a: 123 クナシリヒメメダカハネカクシ（和名新称）HOK, Kun, HON
takashii Naomi, 2004a: 15 ワタナベヒメメダカハネカクシ（和名新称）HON
nipponomontanus Naomi, 1988c: 48 オオダイヒメメダカハネカクシ（和名改称）HON
zaishin zaishin Naomi, 1990c: 102 ザイシンヒメメダカハネカクシ（和名新称）SHI
z. ohbayashii Naomi, 1997e: 141 SHI
lanceolatus Naomi, 1997e: 135トガリヒメメダカハネカクシ（和名新称）HON
yasutoshii Naomi, 1997e: 137 シバタヒメメダカハネカクシ（和名新称）HON
giushin Naomi, 1990c: 101 ギュウシンヒメメダカハネカクシ（和名新称）KYU
satushin Naomi & Puthz, 1994a: 212 サツシンヒメメダカハネカクシ（和名新称）SHI
wasamatanus Puthz, 2001b: 44 ワサマタヒメメダカハネカクシ（和名新称）HON
ochiba Naomi & Puthz, 1994a: 213 オチバヒメメダカハネカクシ（和名新称）HON
kokie Naomi, 2004a: 13 コキエヒメメダカハネカクシ（和名新称）HON
toukin Naomi & Puthz, 1994a: 216トウキンヒメメダカハネカクシ（和名新称）HON
hagai Naomi, 1997e: 138 ハガヒメメダカハネカクシ（和名新称）HON
corrugatus Naomi & Shimada, 2008: 54 シワバネヒメメダカハネカクシ（和名新称）Oki
zimmermanni zimmermanni Puthz, 1968: 43 チンメルマンヒメメダカハネカクシ（和名改称）HON
z. curvaticellus Naomi, 1997e: 142 HON
serratimarginatus Naomi, 1997e: 139 ボウソウヒメメダカハネカクシ（和名新称）HON
sakura Hromádka, 1979d: 115 サクラヒメメダカハネカクシ（和名改称）HON
Species group of S. circularis (Gravenhorst) マルメダカハネカクシ種群
sexualis Sharp, 1874a: 84 コツヤメダカハネカクシ（和名改称）HON; China
Species group of S. javanus Cameron ジャワメダカハネカクシ種群
riukiuensis Puthz, 1973a: 6 リュウキュウメダカハネカクシ Amm, Okn; Taiwan
Species group of S. palposus Zetterstedt ヒゲメダカハネカクシ種群
ruralis Erichson, 1840: 697 アレチメダカハネカクシ（和名改称）HOK; South Korea, North Korea, China, Russia,
EuropeSpecies group of S. melanarius Stephens コクロメダカハネカクシ種群
boops Ljungh, 1810: 158 カワラコクロメダカハネカクシ（和名新称）HOK; China, Russia, Europe
melanarius Stephens, 1833: 299 コクロメダカハネカクシ HOK, HON, SHI, KYU, Tan; Holoarctic Region, Oriental
Region= verecundus Sharp, 1874a: 81
hiroyukii Puthz, 2001c: 103 ヒラノコクロメダカハネカクシ（和名新称）HON

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Yasutoshi Shibata, Munetoshi Maruyama, Hideto Hoshina, Toshio Kishimoto, Shun-Ichiro Naomi, Shûhei Nomura, Volker Puthz, Takashi Shimada, Yasuaki Watanabe, Shûhei Yamamoto

Species group of S. canaliculatus Gyllenhal スジメダカハネカクシ種群
raddei Ryvkin, 1987b: 156 ラッデスジメダカハネカクシ（和名新称）HON; Russia (Amur)
shogun Puthz, 1987: 45 ショウグンスジメダカハネカクシ（和名改称）HON
Species group of S. incanus Erichson ギンゲメダカハネカクシ種群
puberulus Sharp, 1874a: 83 ヒメギンゲメダカハネカクシ（和名改称）HOK, HON, Oki, SHI, KYU= sasajii Naomi, 1988c: 43
Species group of S. gibbicollis J. Sahlberg クビブトメダカハネカクシ種群
mikado Hromádka, 1979d: 117 ミカドクビブトメダカハネカクシ（和名改称）HOK, HON= nomurai Naomi, 1988c: 45
pudefactus Ryvkin, 1987a: 124 タイリククビブトメダカハネカクシ（和名新称）HOK; Kun
miyama Naomi, 1988c: 47 ミヤマクビブトメダカハネカクシ（和名新称）HON
Species group of S. crassus Stephens フトケブカメダカハネカクシ種群
hirtiventris Sharp, 1889: 328 ケブカメダカハネカクシ HON, SHI; China
tomitaorum Naomi, 1989b: 38トミタケブカメダカハネカクシ（和名新称）HOK
pilosiventris Bernhauer, 1915a: 70 ヒメケブカメダカハネカクシ（和名新称）HON; South Korea, North Korea,
China, Russia
immarginatus Mäklin, 1853: 192 キタケブカメダカハネカクシ（和名新称）HOK; Holoarctic Region
Species group of S. opticus Gravenhorst ホソケブカメダカハネカクシ種群
nubilus Ryvkin, 1987c: 270 ヒメホソケブカメダカハネカクシ（和名新称）HOK; Russia (Siberia)
Species group of S. pallitarsis Stephens アシベメダカハネカクシ種群
aestivus Naomi, 1988a: 28 オオアシベメダカハネカクシ（和名改称）HOK
latitarsis Sharp, 1889: 329 ヒラアシアシベメダカハネカクシ（和名改称）HOK, HON
depressus Puthz, 1973b: 88 ヒラタアシベメダカハネカクシ（和名改称）HOK, HON; Taiwan, China, Russia (Ussuri)
benefactor Ryvkin, 1985: 853 リブキンアシベメダカハネカクシ（和名新称）HOK, HON, SHI, KYU; Russia= kanmiyai Naomi, 1988a: 31
Species group of S. piliferus Motschulsky カタビロメダカハネカクシ種群
concinnus Sharp, 1889: 332 マメカタビロメダカハネカクシ HON
hirtellus Sharp, 1874a: 86 タチゲカタビロメダカハネカクシ（和名改称）KYU; Taiwan, China
oblitus Sharp, 1874a: 87 ツヤカタビロメダカハネカクシ（和名改称）HON; South Korea, China, Vietnam, Russia
piliferus Motschulsky, 1858a: 514 カタビロメダカハネカクシ（和名新称）Iri; Taiwan, China, Nepal, India, SE Asia,
Australia, Bismark Archipelago
Species group of S. rorellus Fauvel ネッタイマメメダカハネカクシ種群
confertus Sharp, 1889: 331 ウスグロマメメダカハネカクシ HON
dissimilis Sharp, 1874a: 87 キアシマメメダカハネカクシ（和名改称）HON, KYU; Taiwan, China
Species group of S. similis (Herbst) カワベメダカハネカクシ種群
bohemicus Machulka, 1947: 88 ボヘミアカワベメダカハネカクシ（和名新称）HOK; South Korea, China, Mongolia,
Europeaquilonalis Naomi & Puthz, 1997 (Naomi, 1997g) : 748 クロカワベメダカハネカクシ（和名新称）HOK, HON;
Russia (Sakhalin)mercator Sharp, 1889: 333 カラカネカワベメダカハネカクシ（和名改称）HOK, HON, SHI, KYU; South Korea,

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日本産ハネカクシ科総目録（昆虫綱：甲虫目）
Catalogue of Japanese Staphylinidae (Insecta: Coleoptera)

North Korea, China
sedatus Sharp, 1889: 333 タチゲカワベメダカハネカクシ（和名改称）HOK, HON; North Korea, China, Russia= weisei Bernhauer, 1912: 259
cicindeloides Schaller, 1783: 324 (Staphylinus) アシマダラカワベメダカハネカクシ（和名改称）HOK, HON, Oki,
SHI, KYU; Taiwan, South Korea, Vietnam, Palearctic Region= cicindela Sharp, 1874a: 85
kiesenwetteri Rosenhauer, 1856: 76 アカボシカワベメダカハネカクシ（和名新称）HOK; Russia, Europe
Species group of S. cylindricollis Fauvel マルクビメダカハネカクシ種群
velox Sharp, 1889: 333 ツヤマルクビメダカハネカクシ（和名改称）HON, Oki, SHI
hanami Hromádka, 1979a: 65 ハナミマルクビメダカハネカクシ（和名改称）HON; Taiwan
plagiocephalus Benick, 1940: 567 ヒメマルクビメダカハネカクシ（和名新称）Okn
Species group of S. coelogaster Champion ハラボソメダカハネカクシ種群
oshimaensis Naomi, 1990d: 198 オオシマハラボソメダカハネカクシ（和名新称）Amm, Ish
Species group of S. wasmanni Fauvel ワスマンメダカハネカクシ種群
spurius Benick, 1929b: 260 リュウキュウツヤメダカハネカクシ（和名新称）Iri; Taiwan, China, Philippines= takaoensis Puthz, 1971: 20
Species group of S. flavidulus Sharp ズグロメダカハネカクシ種群
flavidulus flavidulus Sharp, 1889: 334 ズグロメダカハネカクシ HON, Iz-Hch, KYU, Yak; China
f. paederinus Champion, 1924: 160 Amm, Yor, Tok, Oke, Okn, Ish, Iri; Taiwan, India, Oriental Region
Species group of S. bispinus Motschulsky フタトゲメダカハネカクシ種群
currax Sharp, 1874a: 88 ツヤハネブトメダカハネカクシ（和名改称）HON, SHI, KYU; Taiwan, China, Oriental Region
Species group of S. alpicola Fauvel コウザンメダカハネカクシ種群
noctivagus Casey, 1884: 121 ケージーメダカハネカクシ HOK; Russia, Kyrgyztan, Kazakhstan, Nearctic Region
Species group of S. rugipennis Sharp キアシホソメダカハネカクシ種群
rugipennis Sharp, 1874a: 85 キアシホソメダカハネカクシ HOK, HON, Iz-Mk, Iz-Hch, SHI, KYU, Og, Yak, T-Nak,
Amm, Tok, Okn; Taiwan, China, Russia (Siberia)= namazu Hromádka, 1979b: 101= sharpianus Cameron, 1930: 205
Species group of S. abdominalis Fauvel ハラブトメダカハネカクシ種群
bicolon Sharp, 1889: 328 フタテンハラブトメダカハネカクシ（和名改称）HON, SHI, KYU
coronatus zipanguensis Puthz, 1968: 47 オオハラブトメダカハネカクシ（和名改称）HON= watanabei Puthz, 1968: 49
mysticus Sharp, 1888: 330 ハラスジメダカハネカクシ HON, SHI, KYU
sharpi Bernhauer & Schubert, 1911: 185 オオツヤメダカハネカクシ HON, SHI, KYU= palpalis Sharp, 1889: 329
Species group of S. gestroi Fauvel オオメメダカハネカクシ種群
gestroi Fauvel, 1895: 212 オオメメダカハネカクシ Iz-Hch, Yak, T-Nak, Amm, Okn= takara Nakane, 1963: 21
Species group of S. guttalis Fauvel ウスフタテンメダカハネカクシ種群
guttalis Fauvel, 1895b: 212 ウスフタテンメダカハネカクシ Ish; Taiwan, China

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Yasutoshi Shibata, Munetoshi Maruyama, Hideto Hoshina, Toshio Kishimoto, Shun-Ichiro Naomi, Shûhei Nomura, Volker Puthz, Takashi Shimada, Yasuaki Watanabe, Shûhei Yamamoto= guttalis ishigakiensis Naomi, 1988b: 39
Species group of S. cirrus Benick クロツヤタチゲメダカハネカクシ種群
sawadaianus Hromádka, 1979b: 100 サワダタチゲメダカハネカクシ（和名改称）HON, SHI
sugiei Naomi, 1997b: 3 スギエタチゲメダカハネカクシ（和名新称）HON
longisetosus Puthz, 1993a: 160 シコクタチゲメダカハネカクシ（和名新称）SHI
sakaii Naomi & Puthz, 1996: 154 サカイタチゲメダカハメカクシ（和名新称）SHI
yashiro Naomi & Puthz, 1994b: 257 ヤシロタチゲメダカハネカクシ（和名新称）KYU
nyoirin Naomi, 2004f: 21 ニョイリンタチゲメダカハネカクシ（和名新称）HON
kanzeon Naomi, 2004f: 19 カンゼオンタチゲメダカハネカクシ（和名新称）HON
nachiensis Puthz, 2001b: 53 ナチタチゲメダカハネカクシ（和名新称）HON
domburi Hromádka, 1979b: 101ドンブリタチゲメダカハネカクシ（和名改称）SHI
cirriformis Naomi, 1988b: 34 ナガタチゲメダカハネカクシ HON, SHI
amamiensis Naomi, 1988b: 36 アマミタチゲメダカハネカクシ（和名改称）Amm
punctifer Naomi, 1988b: 35 ミナミタチゲメダカハネカクシ（和名改称）Okn= jambar Naomi, 1990b: 52
tokunis Naomi, 1998b: 389トクノシマタチゲメダカハネカクシ（和名新称）Tok
Species group of S. indubius Sharp コバネメダカハネカクシ種群
indubius Sharp, 1889: 330 コバネメダカハネカクシ HON= tanuki Hromádka, 1979c: 107= vernalis Naomi, 1997g: 750
kiyosumiensis Naomi & Takeda, 1991: 10 キヨスミコバネメダカハネカクシ（和名改称）HON
ohishii Naomi, 1987a: 5 オオイシコバネメダカハネカクシ（和名改称）SHI, KYU
fusciceps Naomi, 2006a: 72 シコクコバネメダカハネカクシ（和名新称）SHI
kyushuensis Naomi & Puthz, 2006b (Naomi, 2006b): 74 キュウシュウコバネメダカハネカクシ（和名新称）KYU
hypovelox Naomi & Puthz, 2006b (Naomi, 2006b): 76ドウゴコバネメダカハネカクシ（和名新称）SHI
wakayamanus Puthz, 2001c: 43 ワカヤマコバネメダカハネカクシ（和名新称）
acerrimus Naomi, 2006b: 81 ヒラコバネメダカハネカクシ（和名新称）HON
cucumeroides Naomi & Puthz, 2006b (Naomi, 2006b): 83 コイチャコバネメダカハネカクシ（和名新称）HON
unagi Hromádka, 1979c: 105 イワワキコバネメダカハネカクシ（和名改称）HON= ookami Hromádka, 1979c: 106
fugu Hromadka, 1979c: 104 キイコバネメダカハネカクシ（和名改称）HON= sakana Hromádka, 1979c: 107
ukon Naomi & Puthz, 2006b (Naomi, 2006b): 91 ウコンコバネメダカハネカクシ（和名新称）HON
sawadai Hromádka, 1979c: 107 サワダコバネメダカハネカクシ（和名改称）HON
kishimotoi Naomi & Puthz, 2006b (Naomi, 2006b): 97 キシモトコバネメダカハネカクシ（和名新称）HON
yokozekii Naomi, 2006b: 99 ヨコゼキコバネメダカハネカクシ（和名新称）HON
tateoi Naomi, 2006b: 101 イトウコバネメダカハネカクシ（和名新称）HON
serratus Naomi, 2006b: 103 ユザワコバネメダカハネカクシ（和名新称）HON
sadoensis Naomi, 2006b: 105 サドコバネメダカハネカクシ（和名新称）HON
subserratus Naomi, 2006b: 107 シビツコバネメダカハネカクシ（和名新称）HON

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Catalogue of Japanese Staphylinidae (Insecta: Coleoptera)naomii Puthz, 2001a: 38 ナオミコバネメダカハネカクシ（和名新称）HON
takane Naomi, 1987a: 8 タカネコバネメダカハネカクシ（和名改称）HON
unicoloratus Naomi & Puthz, 2006 (Naomi, 2006b): 114 ヒトイロコバネメダカハネカクシ（和名新称）HON
tohokuensis Naomi & Puthz, 2006 (Naomi, 2006b): 116トウホクコバネメダカハネカクシ（和名新称）HON
autumnalis Naomi, 1997c: 15 アカアシコバネメダカハネカクシ（和名新称）SHI
unzenmontis Naomi & Puthz, 1996: 156 ウンゼンコバネメダカハネカクシ（和名新称）KYU
inimitabilis Puthz, 1993a: 162 ニセコバネメダカハネカクシ（和名新称）SHI
Species group of S. asyura Naomi ハヤシメダカハネカクシ種群⑴ dubitativus-subgroup
dubitativus Puthz, 1993b: 181 アマミハヤシメダカハネカクシ（和名新称）Amm
keman Naomi, 1990b: 45 ケマンハヤシメダカハネカクシ（和名新称）HON= haniwa Hromádka, 1990a: 55
utan Naomi, 1998b: 388 ウータンハヤシメダカハネカクシ（和名新称）HON
incommodus Puthz, 1993a: 146 キョウトハヤシメダカハネカクシ（和名新称）HON
hayashii Puthz, 2003: 17 ナラハヤシメダカハネカクシ（和名新称）HON
ambiguellus Naomi, 1998a: 101 ムネスジハヤシメダカハネカクシ（和名新称）HON
nyorai Naomi, 1990a: 8 ニョライハヤシメダカハネカクシ（和名新称）HON
geisha Puthz, 2001b: 45 シラネハヤシメダカハネカクシ（和名新称）HON
inamatus Puthz, 1993a: 149 チャバネハヤシメダカハネカクシ（和名新称）HON
etsukoae Naomi, 1987a: 2 キタハヤシメダカハネカクシ（和名新称）HOK
puthziellus Naomi, 1997c: 18 プーツハヤシメダカハネカクシ（和名新称）HON
kazami Naomi, 1988f: 80 カザミハヤシメダカハネカクシ（和名改称）HON
silvaticulus Naomi, 1997c: 20 ウスチャハヤシメダカハネカクシ（和名新称）HON
imadatei Naomi & Puthz, 1996: 158 イマダテハヤシメダカハネカクシ（和名新称）HON⑵ kasumi-subgroupbishamon Naomi, 1998b: 390 ビシャモンハヤシメダカハネカクシ（和名新称）SHI
carura Naomi, 1989c: 48 カルラハヤシメダカハネカクシ（和名新称）SHI
kasumi Naomi, 1987a: 6 カスミハヤシメダカハネカクシ（和名改称）HON
houou Naomi, 2010a: 35 ホウオウハヤシメダカハネカクシ（和名改称）HON
yasuhikoiellus Naomi, 2010a: 36 ヤスヒコハヤシメダカハネカクシ（和名改称）HON
inaequatus Puthz, 1993a: 150 クロフハヤシメダカハネカクシ（和名新称）HON
fukuimontium Puthz, 1993b: 183 フクイハヤシメダカハネカクシ（和名新称）HON
sawadaiellus Naomi & Puthz, 1994c: 299 サワダハヤシメダカハネカクシ（和名新称）HON
tengu Hromádka, 1990a: 57 テングハヤシメダカハネカクシ（和名新称）HON
izanagi Naomi & Puthz, 1994c: 303 イザナギハヤシメダカハネカクシ（和名新称）HON
kumoma Naomi, 1987a: 7 クモマハヤシメダカハネカクシ（和名改称）HON
fujimontis Puthz, 2001c: 46 フジハヤシメダカハネカクシ（和名新称）HON
fujiensis Puthz, 2001b: 48 スバルハヤシメダカハネカクシ（和名新称）HON
taoi Naomi, 1990a: 9 タオハヤシメダカハネカクシ（和名新称）HON⑶ hannia-subgroup2013 The Kyushu

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Yasutoshi Shibata, Munetoshi Maruyama, Hideto Hoshina, Toshio Kishimoto, Shun-Ichiro Naomi, Shûhei Nomura, Volker Puthz, Takashi Shimada, Yasuaki Watanabe, Shûhei Yamamoto

zdenae Hromádka, 1990b: 127 ズデナハヤシメダカハネカクシ（和名新称）HON= bunraku Hromádka, 1990b: 128
hannia Naomi, 1990c: 104 ハンニャハヤシメダカハネカクシ（和名新称）HON
oni Naomi, 1988f: 81 オニハヤシメダカハネカクシ（和名改称）HON
haginoi Naomi, 1997c: 16 ハギノハヤシメダカハネカクシ（和名新称）HON⑷ asyura-subgroupkoinobori Hromádka, 1980: 115 アメイロハヤシメダカハネカクシ（和名改称）HOK
maruyamai Naomi, 2004c: 106 マルヤマハヤシメダカハネカクシ（和名改称）HON
jurojin Naomi, 2004c: 108 ジュロウジンハヤシメダカハネカクシ（和名改称）HON
biwa Hromádka, 1980b: 113トワダハヤシメダカハネカクシ（和名改称）HON
asyura asyura Naomi, 1988f: 75 ハヤシメダカハネカクシ（和名改称）HON
a. hakonensis Naomi, 2004g: 25 HON
daikoku Naomi, 2004g: 29 ダイコクハヤシメダカハネカクシ（和名新称）HON
basara Naomi, 1988f: 76 バサラハヤシメダカハネカクシ（和名改称）HON
santira Naomi, 1988f: 78 サンチラハヤシメダカハネカクシ（和名改称）HON
hijiri Naomi, 1989c: 45 ヒジリハヤシメダカハネカクシ（和名新称）HON
bicara Naomi, 1988f: 79 ビカラハヤシメダカハネカクシ（和名改称）HON
yukawai Naomi, 2004a: 186 ユカワハヤシメダカハネカクシ（和名新称）HON
oisami Naomi & Puthz, 1994c: 304 オイサミハヤシメダカハネカクシ（和名新称）HON
okamotoi Naomi, 1989a: 2 オカモトハヤシメダカハネカクシ（和名新称）HON, SHI
Species group of S. cephalotes Sharp オチバメダカハネカクシ種群⑴ cephalotes-subgroup
cephalotes Sharp, 1889: 331 オチバメダカハネカクシ（和名改称）KYU, Tsm= nagasakianus Bernhauer, 1936: 239
alter Puthz, 2003: 15 シマバラオチバメダカハネカクシ（和名新称）KYU
matobai Naomi, 2012: 279 マトバオチバメダカハネカクシ（和名新称）KYU
kagura Naomi & Puthz, 1993: 308 カグラオチバメダカハネカクシ（和名新称）HON
toshiharui Naomi, 1990b: 46 ヨシダオチバメダカハネカクシ（和名新称）KYU
oblongulus Naomi, 1998a: 99 キリシマオチバメダカハネカクシ（和名新称）KYU
inexoratus Puthz, 1993a: 155 イシズチオチバメダカハネカクシ（和名新称）SHI
inaestimatus Puthz, 1993a: 154 アリミネオチバメダカハネカクシ（和名新称）HON
coffeatus Naomi, 2012: 291 コイチャオチバメダカハネカクシ（和名新称）SHI
akome Naomi, 1989d: 166 アコメオチバメダカハネカクシ（和名新称）KYU
kaguyahime Naomi, 1990b: 50 カグヤヒメオチバメダカハネカクシ（和名新称）KYU
otohime Naomi, 1990b: 48 オトヒメオチバメダカハネカクシ（和名新称）KYU
ryugu Naomi, 1989c: 51 リュウグウオチバメダカハネカクシ（和名新称）Tsm
ruricolaris Naomi, 1998a: 102 タチバナオチバメダカハネカクシ（和名新称）KYU
shuheii Naomi, 1990a: 2 ノムラオチバメダカハネカクシ（和名新称）HON
brendelli Naomi, 1997g: 755 ブレンデルオチバメダカハネカクシ（和名新称）KYU
hirashimai Naomi, 1989c: 49 ヒラシマオチバメダカハネカクシ（和名新称）KYU

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Catalogue of Japanese Staphylinidae (Insecta: Coleoptera)

yamajii Naomi & Puthz, 1994c: 305 ヤマジオチバメダカハネカクシ（和名新称）HON⑵ coiffaitiellus-subgroup
coiffaitiellus Naomi & Puthz, 1993: 303 コアッフェオチバメダカハネカクシ（和名新称）HON
inclarescens Puthz, 1993a: 151 ミイロオチバメダカハネカクシ（和名新称）HON
productus Naomi, 2007: 71 ヒメミイロオチバメダカハネカクシ（和名新称）HON
hoshinai Naomi, 2012: 299 ホシナオチバメダカハネカクシ（和名新称）HON
aoi Naomi & Puthz, 1993: 305 アオイオチバメダカハネカクシ（和名新称）HON⑶ syugen-subgroupsyugen Naomi, 1989a: 6 シュゲンオチバメダカハネカクシ（和名新称）HON
himiko Naomi, 1989a: 4 ヒミコオチバメダカハネカクシ（和名新称）HON
dainichi Puthz, 2001b: 50 ダイニチオチバメダカハネカクシ（和名新称）HON
syaca Naomi, 1989c: 46 シャカオチバメダカハネカクシ（和名新称）HON
yoshidai Naomi, 1997g: 757トクシマオチバメダカハネカクシ（和名新称）SHI
maiko Naomi & Puthz, 1993: 306 マイコオチバメダカハネカクシ（和名新称）HON
hagoromo Naomi, 1988b: 38 ハゴロモオチバメダカハネカクシ（和名改称）Ish, Iri, Yon
intermixtus Puthz, 1993a: 164 ハラスジオチバメダカハネカクシ（和名新称）HON
uneme Naomi, 1989a: 8 ウネメオチバメダカハネカクシ（和名新称）SHI
hime Naomi & Puthz, 1994b: 259 ヒメオチバメダカハネカクシ（和名新称）HON
kajika Naomi & Puthz, 1994b: 261 カジカオチバメダカハネカクシ（和名新称）HON
satsuki Naomi, 1987a: 1 サツキオチバメダカハネカクシ（和名改称）SHI
hotei Naomi, 1998b: 391 ホテイオチバメダカハネカクシ（和名新称）SHI⑷ daimio-subgroupdaimio Naomi, 1989d: 162 ダイミョウオチバメダカハネカクシ（和名新称）KYU
imasakai Naomi, 1989d: 163 イマサカオチバメダカハネカクシ（和名新称）KYU
inassuetus Puthz, 1993a: 157 フタカドオチバメダカハネカクシ（和名新称）SHI
inaspectus Puthz, 1993a: 159 マルカドオチバメダカハネカクシ（和名新称）SHI
bifurcatus Naomi, 2011a: 47 フタマタオチバメダカハネカクシ（和名新称）HON
warabi Naomi & Nomura, 1990: 48 キンキオチバメダカハネカクシ（和名新称）HON⑸ okiensis-subgroup
okiensis Naomi & Shimada, 2008: 55 オキオチバメダカハネカクシ（和名新称）Okitsurusakii Naomi, 1998a: 100 ツルサキオチバメダカハネカクシ（和名新称）HON
akojagai Hromádka, 1982: 132 ダイセンオチバメダカハネカクシ（和名新称）HON
enma Naomi, 1990a: 6 エンマオチバメダカハネカクシ（和名新称）HON
amma Naomi & Nomura, 1990: 45 オオサカオチバメダカハネカクシ（和名新称）HON
gagyumontis Naomi, 1990a: 4 ガギュウオチバメダカハネカクシ（和名新称）HON
masatakai Naomi & Nomura, 1990: 47 サトウオチバメダカハネカクシ（和名新称）HON
amagasui Naomi, 2007: 73 アマガスオチバメダカハネカクシ（和名新称）HON
miroku Naomi, 2006a: 30 ミロクオチバメダカハネカクシ（和名新称）HON
uenoi Naomi & Nomura, 1990: 46 ウエノオチバメダカハネカクシ（和名新称）HON
bosatsu Naomi, 1989d: 159 ボサツオチバメダカハネカクシ（和名新称）HON

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Yasutoshi Shibata, Munetoshi Maruyama, Hideto Hoshina, Toshio Kishimoto, Shun-Ichiro Naomi, Shûhei Nomura, Volker Puthz, Takashi Shimada, Yasuaki Watanabe, Shûhei Yamamotovolkeri Naomi, 1998b: 385 プーツオチバメダカハネカクシ（和名新称）HON
ferus Naomi, 2012: 305 オワセオチバメダカハネカクシ（和名新称）HON
testaceopiceus Bernhauer, 1938a: 31 ベルンハウアオチバメダカハネカクシ（和名改称）HON
amida Naomi, 1989d: 164 アミダオチバメダカハネカクシ（和名新称）SHI
pubicornis Naomi, 1998b: 387 ワカヤマオチバメダカハネカクシ（和名新称）HON⑹ rufescens-subgroup (monophyletic subgroup)
ohtoensis Naomi, 2006a: 38 ナンキオチバメダカハネカクシ（和名新称）HON
cygnipenis Puthz, 2001b: 52 ツヤズオチバメダカハネカクシ（和名新称）HON
ebisu Naomi, 2006a: 47 エビスオチバメダカハネカクシ（和名新称）HON
olliformis olliformis Naomi, 2006a: 44 オオミネオチバメダカハネカクシ（和名新称）HON
o. owasenus Naomi, 2006a: 46 HON
agrestis Naomi, 2006a: 42 キイオチバメダカハネカクシ（和名新称）HON
jukata Hromádka, 1982: 135 コウヤオチバメダカハネカクシ（和名改称）HON
jamesashei Naomi, 2012: 312 アッシェオチバメダカハネカクシ（和名新称）HON
ichikawai Naomi, 2006a: 49 イチカワオチバメダカハネカクシ（和名新称）HON
acutatus Naomi, 2012: 310トガリオチバメダカハネカクシ（和名新称）HON
rufescens Sharp, 1874a: 88トビイロオチバメダカハネカクシ（和名改称）HON
daigonis Naomi & Puthz, 1993: 310 ダイゴウオチバメダカハネカクシ（和名新称）HON
wasabi Hromádka, 1982: 133 ワサビオチバメダカハネカクシ（和名改称）HON
ingens ingens Naomi, 2006a: 62 コモノオチバメダカハネカクシ（和名新称）HON
i. ryugatakensis Naomi, 2006a: 64 HON
lubomiri Naomi, 2006a: 64 ハロマドカオチバメダカハネカクシ（和名新称）HON
ichihashii ichihashii Naomi, 2006a: 67 イチハシオチバメダカハネカクシ（和名新称）HON
i. miunensis Naomi, 2006a: 70 HON
mikawanis Naomi, 1997f: 6006 ミカワオチバメダカハネカクシ（和名新称）HON
gyrosus Naomi, 2006a: 72 ホズミオチバメダカハネカクシ（和名新称）HON
nakanei nakanei Hromádka, 1982: 132 ナカネオチバメダカハネカクシ（和名改称）HON= ignorabilis Naomi, 1997f: 607
n. awajinis Naomi, 1997f: 609 HON

Subfamily Euaesthetinae Thomson, 1859 チビフトハネカクシ亜科（４属39種）Tribe Euaesthetini Thomson, 1859
Genus Edaphus Motschulsky, 1856 スジツヤチビハネカクシ属
bannadakemontis Puthz, 2010d: 82 バンナダケスジツヤチビハネカクシ（和名新称）Ishbeszedesi Reitter, 1914: 189 ヒメスジツヤチビハネカクシ HON; China, Turkey, Holoarctic Region= piceus Cameron, 1933: 1692013 The Kyushu

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Catalogue of Japanese Staphylinidae (Insecta: Coleoptera)bifoveolifrons Puthz, 2010d: 83 フタアナスジツヤチビハネカクシ（和名新称）Tok
callosifrons Puthz, 2010d: 89 コブスジツヤチビハネカクシ（和名新称）Ishcarinicollis Bernhauer, 1907: 380 スジツヤチビハネカクシ HON, SHI, Yak; China
cloanthus Puthz, 2010d: 77 ヤエヤマスジツヤチビハネカクシ（和名新称）Ish, Iridiversipunctatus Puthz, 2010d: 88 ヤンバルスジツヤチビハネカクシ（和名新称）Okn
fuscipennis Puthz, 2010d: 80 チャイロスジツヤチビハネカクシ（和名新称）Iri
impressipennis Puthz, 2010d: 82 ナガスジツヤチビハネカクシ（和名新称）Amm
iriomoteanus Puthz, 2010d: 79 イリオモテスジツヤチビハネカクシ（和名新称）Iri
ishigakiensis Puthz, 2010d: 81 イシガキスジツヤチビハネカクシ（和名新称）Ishjaponicus Sharp, 1889: 325 ミナミスジツヤチビハネカクシ（和名新称）KYU, T-Aks; Thailand
kojimai Puthz, 2010d: 91 コジマスジツヤチビハネカクシ（和名新称）Okn
kumejimanus Puthz, 2010d: 85 クメジマスジツヤチビハネカクシ（和名新称）Kum
mezentius Puthz, 2010d: 77 アマミスジツヤチビハネカクシ（和名新称）Amm
nakayamensis Puthz, 1980: 243 キタスジツヤチビハネカクシ HOK
napporoensis Puthz, 1980: 244 ウスアカスジツヤチビハネカクシ HOK
nipponensis Puthz, 1975: 181 ニホンスジツヤチビハネカクシ HON, KYU
nomurai Puthz, 2010c: 288: ノムラスジツヤチビハネカクシ（和名新称）Amm, Ish, Iri; Taiwan
ogatai Puthz, 2010d 90 オガタスジツヤチビハネカクシ（和名新称）Yon
okinawaensis Puthz, 1980: 241 オキナワスジツヤチビハネカクシ Okn, Amm, Tokomotomontis Puthz, 2010d: 88 オモトスジツヤチビハネカクシ（和名新称）Ish
perdifficilis Puthz, 2010d: 82 ノソコダケスジツヤチビハネカクシ（和名新称）Ishperexillis Puthz, 2010d: 85 クニガミスジツヤチビハネカクシ（和名新称）Okn
perversus Puthz, 1994: 390 [RN] ヤマトスジツヤチビハネカクシ（和名改称）HON= japonicus Bernhauer, 1907: 380 (Euaesthetus) [HN]
riukiuensis Puthz, 1980: 242 リュウキュウスジツヤチビハネカクシ Oknsergestus Puthz, 2010d: 76トクノシマスジツヤチビハネカクシ（和名新称）Tok
siluvius Puthz, 2010d: 77 ユワンダケスジツヤチビハネカクシ（和名新称）Okn, Amm
takaii Puthz, 2010d: 83 タカイスジツヤチビハネカクシ（和名新称）Amm
tokaraensis Puthz, 2010d: 87トカラスジツヤチビハネカクシ（和名新称）T-Nak, T-Aks
yonaguniensis Puthz, 2010d: 87 ヨナグニスジツヤチビハネカクシ（和名新称）Yon
Genus Euaesthetus Gravenhorst, 1806 チビフトハネカクシ属
hammondi Puthz, 1976: 46 ハモンドチビフトハネカクシ HON; China
nitidulus Sharp, 1874a: 78 ムナクボチビフトハネカクシ HON, KYU
nomurai Puthz, 2010a: 11 キタチビフトハネカクシ（和名改称）HON
Genus Octavius Fauvel, 1873 ヒメチビフトハネカクシ属（和名新称）
flavescens Kistner, 1961: 20 (Doletica) アメイロチビフトハネカクシ（和名新称）SHI, Ish; Taiwan, Philippines,
Thailand, Indonesia (Sumatra), India

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Yasutoshi Shibata, Munetoshi Maruyama, Hideto Hoshina, Toshio Kishimoto, Shun-Ichiro Naomi, Shûhei Nomura, Volker Puthz, Takashi Shimada, Yasuaki Watanabe, Shûhei Yamamoto

Tribe Stenaesthetini Bernhauer & Schubert, 1911
Genus Stenaesthetus Sharp, 1874 ニセメダカハネカクシ属
insulanus Puthz, 2010b: 55 サキシマニセメダカハネカクシ（和名新称）Ish, Irilackneri Puthz, 2011: 17 アマミニセメダカハネカクシ（和名新称）Amm
okinawaensis Puthz, 1986: 299 オキナワニセメダカハネカクシ Amm, Okn, Kum
sunioides Sharp, 1874a: 80 ニセメダカハネカクシ HON, Iz-Hch, Og, Okn, KYU, Yak, Okn; Taiwan, China, Nepal,
Pakistan, India, Oriental Region

Subfamily Scydmaeninae Leach, 1815 コケムシ亜科（13属67種１亜種）
Tribe Cephenniini Reitter, 1882
Genus Cephennodes Reitter, 1884 ムナビロコケムシ属
Subgenus Cephennodes Reitter, 1884
araiorum Jałoszyński, 2003: 226 (Chelonoidum) イソニセムナビロコケムシ HON
japonicus Sharp, 1886: 50 (Cephennium) ヤマトムナビロコケムシ KYU, South Korea
mizunoi Hoshina & Kishimoto, 2004: 2 チウンムナビロコケムシ Og
moderatus Kurbatov, 1995: 951(Chelonoidum) クナシリムナビロコケムシ Kun, HON, SHI
pullatus Kurbatov, 1995: 948 (Chelonoidum) オオムナビロコケムシ SHI
torosus Kurbatov, 1995: 957(Chelonoidum) バンドウムナビロコケムシ HON
vafer Kurbatov, 1995: 946 チシマムナビロコケムシ Kun, HON, SHI, KYU
yonaguniensis Hoshina, 2012b: 426 ヨナグニムナビロコケムシ Yon
Subgenus Fusionodes Jałoszyński 2007
loebli Kurbatov, 1995: 954 (Chelonoidum) イシヅチムナビロコケムシ SHI, HON
Insertae sedis (subgenus?) 亜属不明
besucheti Kurbatov, 1995: 954 (Chelonoidum) ミヤマムナビロコケムシ SHIGenus Cephennomicrus Reitter, 1907 ハラビロコケムシ属
cactiformis Jałoszyński & Hoshina, 2003: 58 (Neseuthia) ツシマハラビロコケムシ Tsm
disjunctus Jałoszyński, S. Arai & K. Arai, 2004: 62 (Neseuthia) リュウキュウハラビロコケムシ（和名新称）Okn
hobbiti Jałoszyński & Hoshina, 2003: 53 (Neseuthia) ミナミハラビロコケムシ Ishiriomotensis Jałoszyński, 2009: 9 イリオモテハラビロコケムシ（和名新称）Irijaponigenus Jałoszyński & Hoshina, 2003: 55 (Neseuthia)トクノシマハラビロコケムシ Tok
nagoanus Jałoszyński, 2009: 12 ナゴハラビロコケムシ（和名新称）Okn
nomurai Jałoszyński & Hoshina, 2003: 49 (Neseuthia) ヤエヤマハラビロコケムシ Ish, Irin. disjuncta Jałoszyński, S. Arai & K. Arai, 2004: 62 Okn
okinawanus Jałoszyński, S. Arai & K. Arai, 2004: 59 (Neseuthia) オキナワハラビロコケムシ（和名新称）Okn

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Catalogue of Japanese Staphylinidae (Insecta: Coleoptera)

pseudojaponigenus Jałoszyński, 2009: 17 オモトハラビロコケムシ（和名新称）Ishtanakai Hoshina, 2012a: 191 カワラハラビロコケムシ HON
Tribe Cyrtoscydmini L. W. Schaufuss, 1889
Genus Euconnus Thomson, 1859 ヒメコケムシ属
Subgenus Cladoconnus Reitter, 1909
matsuae Hoshina, 2004a: 119 マツクシヒゲヒメコケムシ HON
Subgenus Euconnus Thomson, 1859
japonicus Sharp, 1874b: 127 (Scydmaenus) ヤマトヒメコケムシ HON, KYU
ohnoensis Hoshina, 2006: 44 ロクロシヒメコケムシ HON
Subgenus Euconophron Reitter, 1909
fustiger Sharp, 1874b: 128 (Scydmaenus) シリブトヒメコケムシ HON, SHI, KYU
kojiroi Hoshina, 2004b: 17 コジロウヒメコケムシ HON
matsunoyamensis Hoshino & Nagano, 2005: 2 マツノヤマヒメコケムシ HON
unpini Hoshina, 2007: 18 ウンピンヒメコケムシ HON
Subgenus Napochus Thomson, 1859
lewisii Sharp, 1886: 47 ルイスヒメコケムシ HON, KYU= miyawakianus Franz, 1976: 57
Subgenus Pycnophus Casey, 1897
impar Sharp, 1886: 46 バンドウツヤヒメコケムシ HON, KYU= otorii Hoshina & Arai, 2003: 30
tsugaruensis Hoshina & Arai, 2003: 28 ツガルツヤヒメコケムシ HON
Incertae sedis (subgenus?) 亜属不明
dulcis Sharp, 1886: 47 カワラヒメコケムシ HON, KYU
oscillans Sharp, 1886: 48 ハラボソヒメコケムシ KYU
raucus Sharp, 1886: 48 アカチャヒメコケムシ KYU
schoenfeldti Reitter, 1891: 19 チャイロヒメコケムシ Japan
Genus Horaeomorphus Schaufuss, 1889トゲアシオオコケムシ属
sakishimanus Jałoszyński, 2003: 224 サキシマトゲアシオオコケムシ Ish, Iri
Genus Microscydmus Saulcy & Croissandeau, 1893 コヒメコケムシ属
Subgenus Microscydmus Saulcy & Croissandeau, 1893
debilis Sharp, 1874b: 127 (Scydmaenus) コヒメコケムシ HON, SHI, KYU
Genus Neuraphes Thomson, 1859 ムナスジコケムシ属
Subgenus Pararaphes Reitter, 1891
niponensis Franz, 1976: 53 エゾムナスジコケムシ HOK2013 The Kyushu

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Yasutoshi Shibata, Munetoshi Maruyama, Hideto Hoshina, Toshio Kishimoto, Shun-Ichiro Naomi, Shûhei Nomura, Volker Puthz, Takashi Shimada, Yasuaki Watanabe, Shûhei Yamamoto

Genus Scydmoraphes Reitter, 1891 カクムネコケムシ属
japonicus Franz, 1976: 54 ヤマトカクムネコケムシ HOK, HON
Genus Stenichnus Thomson, 1859 マルムネコケムシ属
bellulus Jałoszyński, 2004d: 158 ヤエヤママルムネコケムシ Ish, Iriminipollens Jałoszyński, 2004d: 170 ヒメマルムネコケムシ HOK
mitai Jałoszyński, 2006: 54トカラマルムネコケムシ T-R
pollens Sharp, 1886: 49 (Scydmaenus) チンゼイマルムネコケムシ KYU
sakurayamanus Jałoszyński & K. Arai, 2009: 66 サクラヤママルムネコケムシ HON
totoro Jałoszyński, 2004d: 170トトロマルムネコケムシ Ish, Iri
Genus Syndicus Motschulsky, 1851 モモスジオオコケムシ属
Subgenus Syndicus Motschulsky, 1851
yaeyamensis Jałoszyński, 2004a: 68 ヤエヤマモモスジオオコケムシ Ish, Iri
Tribe Eutheiini Casey, 1897
Genus Eutheia Steohens, 1830 ニセナガコケムシ属
horii Jałoszyński, 2004b: 133 エゾニセナガコケムシ HOK
rufa Jałoszyński, 2004b: 130 サビニセナガコケムシ HOK
yoshidai Hoshina, 2007: 7 アワニセナガコケムシ SHIGenus Paraneseuthia Franz, 1986 ナガコケムシ属
holzneri Franz, 1976: 51 (Eutheia) コゲチャナガコケムシ HON
inexpectata Jałoszyński, 2006: 133 サガミナガコケムシ HOK
paradoxa K. Sawada, 1962a: 19 (Euthiconus) チビナガコケムシ HON, KYU
Genus Veraphis Casey, 1897 ヒラタコケムシ属
engelmarki niponensis Jałoszyński & Hoshina, 2005: 128 タイリクヒラタコケムシ HOK
hisamatsui Jałoszyński & Hoshina, 2005: 108 エゾヒラタコケムシ HOK
horianus Jałoszyński & Hoshina, 2005: 123 アイヌヒラタコケムシ HOK
ishikawai Hisamatsu, 1985: 13 ヒゲブトヒラタコケムシ HOK
japonicus Sawada, 1962a: 20 ホソヒラタコケムシ HOK, HON, SHI
kurbatovi Jałoszyński & Hoshina, 2005: 118 ミチノクヒラタコケムシ HON
oodaigaharensis Jałoszyński & Hoshina, 2005: 110 オオダイヒラタコケムシ HON
sawadai Jałoszyński & Hoshina, 2005: 116 ムツヒラタコケムシ HON
tottoriensis Jałoszyński & Hoshina, 2005: 112 サンインヒラタコケムシ HON
Tribe Scydmaenini Leach, 1815

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Catalogue of Japanese Staphylinidae (Insecta: Coleoptera)

Genus Scydmaenus Latreille, 1802 コケムシ属
Subgenus Scydmaenus
takaranus Nakane, 1963: 22 ヒメムクゲコケムシ T-Tak
Subgenus Mascarensia Franz, 1973
kasugaensis Franz, 1976: 58 カスガコケムシ HON= honshuensis Franz, 1976: 57
Incertae sedis (subgenus?) 亜属不明
angustulus Csiki, 1919: 78 [RN] ムネボソコケムシ HON= angustus Sharp, 1886: 49 [HN] (Eumicrus)
cribratus Sharp, 1886: 50 (Eumicrus) アイヌコケムシ HOK, HON
reversus Sharp, 1874b: 128 (Scydmaenus) ヒゼンコケムシ KYU
vestitus Sharp, 1874b: 126 (Eumicrus) ムクゲコケムシ HON, KYU, Yae; Taiwan
curtipennis Sharp, 1886: 49 (Eumicrus) フトコケムシ KYU

Subfamily Pseudopsinae Ganglbauer, 1895 スジヒラタハネカクシ亜科（１属１種）
Genus Pseudopsis Newman, 1834 スジヒラタハネカクシ属
watanabei Herman, 1975: 274 スジヒラタハネカクシ HOK, Rsr, HON

Subfamily Paederinae Fleming, 1821 アリガタハネカクシ亜科（37属281種９亜種）
Tribe Paederini Fleming, 1821
Subtribe Acanthoglossina Coiffait, 1982
Genus Acanthoglossa Kraatz, 1859 ケブカハネカクシ属（和名新称）
hirtella Sharp, 1889: 320 (Eomedon) ケブカハネカクシ（和名新称）Awj, SHI, KYU; South Korea
Subtribe Astenina Hatch, 1957
Genus Astenus Dejean, 1833 シリグロハネカクシ属（和名新称）
Subgenus Astenognathus Reitter, 1909
indicus Kraatz, 1859: 148 (Sunius) キバネシリグロハネカクシ（和名新称）Japan; Taiwan, China, Oriental Region,
Australian region, Canarias, Africa= oculatus Sharp, 1874a: 72 (Sunius)maculipennis Kraatz, 1859: 148 (Sunius) フタホシシリグロハネカクシ HON, Iz-My, Iz-Hch, SHI, KYU, Tan, Yak,
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Yasutoshi Shibata, Munetoshi Maruyama, Hideto Hoshina, Toshio Kishimoto, Shun-Ichiro Naomi, Shûhei Nomura, Volker Puthz, Takashi Shimada, Yasuaki Watanabe, Shûhei Yamamoto= bicolon Sharp, 1874a: 72 (Sunius)porosus Sharp, 1889: 322 (Sunius) ツヤケシシリグロハネカクシ（和名新称）HON, SHI; South Korea
Subgenus Astenopleuritus Coiffait, 1982
leptocerus Eppelsheim, 1895: 64 (Sunius) ホソシリグロハネカクシ（和名新称）Og; Oriental Region, Ethiopian
RegionSubgenus Astenus Dejean, 1833
angulatus Sharp, 1874a: 70 (Neognatus) シリグロハネカクシ（和名改称）KYU, Amm, Tok; Russia (Siberia), China
suffusus Sharp, 1874a: 73 (Sunius) オオシリグロハネカクシ HOK, HON, Iz-My, Iz-Hch, Oki, SHI, KYU, Yak= flavipennis Bernhauer, 1927: 93
Subgenus Eurysunius Reitter, 1909
setifer Cameron, 1930: 206トゲトゲハネカクシ HON, KYU
Incertae sedis (subgenus?) 亜属不明
brevipes Sharp, 1874a: 71 (Sunius) ヒメシリグロハネカクシ HON, Iz-My, Iz-Mk, Iz-A, Og, Oki, SHI, KYU, Amm,
Okn; Taiwan, South Korea
chloroticus Sharp, 1874a: 74 (Sunius) ヤマトシリグロハネカクシ HON, Iz-Hch, Iz-A, Og, SHI, KYU, Tsm, Tan,
Yak, Amm; Taiwan, China
japonicus Zhao & Sakai, 1999: 167 カントウシリグロハネカクシ（和名新称）HON
latifrons Sharp, 1874a: 70 (Sunius) キアシハラグロハネカクシ（和名改称）HON, Iz-A, Oki, KOs, KYU, Kbs, Nkd,
Tan, Amm, Okn, Ish; Taiwan= histrio Sharp, 1874a: 71
naomii Zhao, 1996: 259 ヤエヤマシリグロハネカクシ（和名新称）Ish, Iriohbayashii Zhao, Li & Sakai, 1997: 85 アマミシリグロハネカクシ（和名新称）Amm
sakaii Zhao & Li, 1998: 56トカラシリグロハネカクシ（和名新称）T-R
shibatai T. Ito, 1995a: 30 ヒゲノシリグロハネカクシ Amm, Tok, Ish, Irispinosus Zhao & Sakai, 1997: 87トゲシリグロハネカクシ（和名新称）Ishtaiwanus T. Ito, 1996c: 87 タイワンシリグロハネカクシ（和名新称）Ish; Taiwan
yonezoi T. Ito, 1995a: 32 リュウキュウシリグロハネカクシ Amm, Tok, Oke, Okn, Yon
Genus Nazeris Fauvel, 1873 アバタコバネハネカクシ属
angustus T. Ito, 1992b: 128 ホソアバタコバネハネカクシ（和名新称）HON
densiventris Koch, 1939: 158 ツクシアバタコバネハネカクシ（和名新称）KYU
gotoi T. Ito, 1986: 89 アマミアバタコバネハネカクシ（和名改称）Amm
hasegawai hasegawai Nakane & Sawada, 1954: 737 オゼアバタコバネハネカクシ（和名新称）HON
h. dewanus T. Ito, 1992b: 127 HON
hayashii T. Ito, 1999b: 131 ミエアバタコバネハネカクシ（和名新称）HON
hikosanus T. Ito, 1991a: 7 ヒコサンアバタコバネハネカクシ（和名新称）SHI, KYU
hisamatsui T. Ito, 1991a: 8 イヨアバタコバネハネカクシ（和名新称）SHI, KYU
izuensis T. Ito, 2009c: 149 イズアバタコバネハネカクシ HON
masaohayashii T. Ito, 1991c: 129 カイアバタコバネハネカクシ（和名新称）HON
naomii T. Ito, 1992b: 134 クジュウアバタコバネハネカクシ（和名新称）KYU

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Catalogue of Japanese Staphylinidae (Insecta: Coleoptera)nipponicus T. Ito, 1991c: 133 カナガワアバタコバネハネカクシ（和名新称）HON
ohkurai T. Ito, 1992b: 130 タカオアバタコバネハネカクシ（和名新称）HON
okinawanus okinawanus T. Ito, 1986: 91 オキナワアバタコバネハネカクシ Okn
o. amamianus T. Ito, 1994c: 102 Tok
omogonis T. Ito, 1991a: 10 オモゴアバタコバネハネカクシ（和名新称）SHI
optatus Sharp, 1889: 322 (Mesunius) ヒメアバタコバネハネカクシ HON, SHI
pacificus T. Ito, 1990b: 99 ムロトアバタコバネハネカクシ（和名新称）SHI
shibatai T. Ito, 1990b: 97 キイアバタコバネハネカクシ（和名新称）HON
suensoni suensoni Bernhauer, 1936: 306 ウンゼンアバタコバネハネカクシ（和名新称）KYU
s. higonis T. Ito, 1992b: 133 KYUtaichii T. Ito, 2008a: 180 ソバツブアバタコバネハネカクシ（和名新称）HON
validus T. Ito, 1991c: 131 ニセアバタコバネハネカクシ（和名新称）HON
watanabei T. Ito, 1990a: 70 アマギアバタコバネハネカクシ（和名新称）HON
wollastoni wollastoni Sharp, 1874a: 68 (Mesunius) アバタコバネハネカクシ HON, Oki, SHI
w. harimanus T. Ito, 1991c: 128 HON
w. kyotensis T. Ito, 1991c: 128 HON
w. peninsularis T. Ito, 1991c: 129 HON
yaeyamanus T. Ito, 2008a: 177 ヤエヤマアバタコバネハネカクシ（和名新称）Ishyasuakii T. Ito, 2002: 247 ヤマトアバタコバネハネカクシ（和名新称）HON
yoshidai T. Ito, 1996e: 213 アワアバタコバネハネカクシ（和名新称）SHI
Subtribe Cryptobiina Casey, 1905
Genus Cephalochetus Kraatz, 1859 ナガズハネカクシ属（和名新称）
rufus Cameron, 1918: 78 (Calliderma) ナガズハネカクシ T-Nak, Ish; Singapore, Indonesia (Sumatra), India
Genus Homaeotarsus Hochhuth, 1851 ヤマトナガエハネカクシ属（和名新称）
Subgenus Hesperobium Casey, 1886
japonicus Sharp, 1874a: 60 (Cryptobium) ヤマトナガエハネカクシ HOK, HON, Iz-Mk, KYU, Okn; Taiwan, South
Korea, North Korea, China
Genus Monocrypta Casey, 1905 ナガエハネカクシ属（和名新称）
apicata Sharp, 1874a: 59 (Cryptobium) ナガエハネカクシ（和名新称）KYU
pectoralis Sharp, 1874a: 60 (Cryptobium) アカバナガエハネカクシ HON, Iz-Hch, KYU, Tan; Taiwan, South Korea,
China, Thailand
Genus Ochthephilum Stephens, 1829 カワベナガエハネカクシ属（和名新称）
biwakense T. Ito, 2008b: 29 ビワコカワベナガエハネカクシ（和名新称）HON
chibaense Naomi, 1995c: 165 チバカワベナガエハネカクシ（和名新称）HON
cuneatum Sharp, 1889: 253 (Cryptobium) カワベナガエハネカクシ HON, Sad, Tsm; Taiwan

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Yasutoshi Shibata, Munetoshi Maruyama, Hideto Hoshina, Toshio Kishimoto, Shun-Ichiro Naomi, Shûhei Nomura, Volker Puthz, Takashi Shimada, Yasuaki Watanabe, Shûhei Yamamoto

densipenne Sharp, 1889: 254 (Cryptobium) クロカワベナガエハネカクシ HOK, HON, KYU; South Korea, China
harusawai T. Ito, 1996b: 78 アマミカワベナガエハネカクシ（和名新称）Amm
hokkaidense T. Ito, 2008b: 30 エゾカワベナガエハネカクシ（和名新称）HOK
okinawaense Y. Watanabe, 2008a: 199 オキナワカワベナガエハネカクシ Okn
shibatai T. Ito, 1996b: 81 サキシマカワベナガエハネカクシ（和名新称）Ish, Irishinanense T. Ito, 2008b: 30 シナノカワベナガエハネカクシ（和名改称）HON
Incertae sedis (genus ?) 属不確定
kurosai T. Ito, 1996b: 77 ツマアカカワベナガエハネカクシ HON, SHI, KYU, Yak
Subtribe Echiasterina Casey, 1905
Genus Echiaster Erichson, 1839 ホソクビサビハネカクシ属（和名新称）
unicolor japonicus Bernhauer, 1923a: 123 ホソクビサビハネカクシ（和名新称）Iz-My, SHI, Shd, Tsm
Subtribe Lathrobiina Laporte, 1835
Genus Domene Fauvel, 1873 マルズハネカクシ属
Subgenus Macromene Coiffait, 1982
crassicornis Sharp, 1874a: 59 (Lathrobium) マルズハネカクシ HOK, HON, Awj, Oki, Shd, SHI, KYU; South
Korea, China
curtipennis Sharp, 1889: 261 コマルズハネカクシ HOK, HON, Iz-O, Iz-My, SHI, KYU, Tsm; South Korea, China
daimio Sharp, 1889: 260 ダイミョウマルズハネカクシ HON
orbiculata Sharp, 1889: 260 マルムネマルズハネカクシ HON, Sad, SHI
stilicoides Sharp, 1874a: 58 (Lathrobium) カラカネマルズハネカクシ（和名改称）KYU
Genus Lathrobium Gravenhorst, 1802 ナガハネカクシ属
Subgenus Lathrobium Gravenhorst, 1802
Group of L. dignum ナガハネカクシ種群
cylindricum Bernhauer, 1938a: 37 ウンゼンナガハネカクシ（和名改称）KYU; China
dignum Sharp, 1874a: 55 ナガハネカクシ（和名改称）HOK, Rsr, HON, SHI, KYU; South Korea, North Korea,
China, Russia
ishiharai Hayashi, 1994b: 145 イシハラナガハネカクシ HON, SHI, KYU
kobense Sharp, 1874a: 57 アカバチビナガハネカクシ HON, SHI
monilicorne Sharp, 1889: 259 ヒゲブトナガハネカクシ HOK, HON; China
scabripenne Sharp, 1874a: 58 ナガサキナガハネカクシ KYUGroup of L. nomurai/ harimanum オオコバネナガハネカクシ種群
aioiense Y. Watanabe, 2002b：442 アイオイオオコバネナガハネカクシ HON
awajishimanum Y. Watanabe, 2001a: 38 アワジオオコバネナガハネカクシ Awj
daisenense Y. Watanabe, 1987a: 8 ダイセンオオコバネナガハネカクシ（和名改称）HON
daisensanum Y. Watanabe, 1998: 92 ダイセンサンオオコバネナガハネカクシ（和名改称）SHI2013 The Kyushu

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Catalogue of Japanese Staphylinidae (Insecta: Coleoptera)

fujimotoi Y. Watanabe, 2001a: 41 フジモトオオコバネナガハネカクシ KYU
harimanum Y. Watanabe, 1986c: 688 ハリマオオコバネナガハネカクシ（和名改称）HON
hikosanense Y. Watanabe, 1998: 91 ヒコサンオオコバネナガハネカクシ（和名改称）KYU
iwamiense Y. Watanabe, 1991c: 151 イワミオオコバネナガハネカクシ（和名改称）HON
kamezawai Y. Watanabe, 2005b: 29 カメザワオオコバネナガハネカクシ SHIkanmuriense Y. Watanabe, 2002b: 439 カンムリオオコバネナガハネカクシ HON
kasagatanum Y. Watanabe, 2002b: 445 カサガタオオコバネナガハネカクシ HON
kasumiense Y. Watanabe, 2002b: 444 カスミオオコバネナガハネカクシ HON
katsumiae Y. Watanabe & Yoshida, 2007: 59 カツミオオコバネナガハネカクシ SHIkishuense Y. Watanabe, 1991c: 145 キイオオコバネナガハネカクシ（和名改称）HON
konpira Y. Watanabe, 1991c: 149 コンピラオオコバネナガハネカクシ（和名改称）SHI
morii Y. Watanabe, 2002b: 444 モリオオコバネナガハネカクシ HON
moritai Y. Watanabe, 1998: 94 モリタオオコバネナガハネカクシ（和名改称）HON
narutoensis Y. Watanabe, 2010b: 67 ナルトオオコバネナガハネカクシ SHI
nomurai Nakane, 1955: 29 オオコバネナガハネカクシ KYUohdaiense Y. Watanabe, 1998: 90 オオダイオオコバネナガハネカクシ（和名改称）HON
ohtakisanum Y. Watanabe, 2010b: 63 オオタキサンオオコバネナガハネカクシ SHItanakai Y. Watanabe, 1998: 86 タナカオオコバネナガハネカクシ（和名改称）HON
tosanum Y. Watanabe, 1987a: 11トサオオコバネナガハネカクシ（和名改称）SHI
tsurugisanum Y. Watanabe, 1991c: 153 ツルギオオコバネナガハネカクシ（和名改称）SHI
uenoi Y. Watanabe, 1980a: 21 ウエノオオコバネナガハネカクシ（和名改称）HON
Group of L. pollens/shingon コバネナガハネカクシ種群
aonoi Y. Watanabe, 2011c: 177 アオノコバネナガハネカクシ HON
arakawai Y. Watanabe, 1992a: 39 タニガワコバネナガハネカクシ HON
awanum Y. Watanabe, 1991a: 35 アワコバネナガハネカクシ SHIdensum Bernhauer, 1936: 307 オカヤマコバネナガハネカクシ
HON（和名改称）
dozenense Y. Watanabe & Shimada, 2005: 604ドウゼンコバネナガハネカクシ Oki
gomadanzanum Y. Watanabe, 2005c: 595 ゴマダンコバネナガハネカクシ HON
hakusanum Y. Watanabe & Hoshina, 2003: 8 ハクサンコバネナガハネカクシ HON
hayashii Y. Hayashi, 1999: 147 コンゴウコバネナガハネカクシ（和名新称）HON
hirakuranum Y. Watanabe, 2005a: 323 ヒラクラコバネナガハネカクシ HON
hisamatsui Y. Watanabe & Yoshida, 2009: 89 イシヅチコバネナガハネカクシ SHI
ishidai Hayashi, 1996: 19 タンバコバネナガハネカクシ（和名新称）HON
itsukushimanum Y. Watanabe, 2011b: 47 イツクシマコバネナガハネカクシ HON
izumoense Y. Watanabe, 2010a: 79 イズモコバネナガハネカクシ HON
japonicum japonicum Bernhauer, 1907: 384 リシリコバネナガハネカクシ Rsr, Etr= rishiriense Y. Watanabe & Y. Shibata, 1965b: 318
j. kunashiriense Y. Watanabe, 2004c: 39 Kun
kusamai Y. Watanabe, 1999b: 76 スルガコバネナガハネカクシ（和名改称）HON
masatoi Y. Watanabe, 2010a: 76 マサトコバネナガハネカクシ HON

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Yasutoshi Shibata, Munetoshi Maruyama, Hideto Hoshina, Toshio Kishimoto, Shun-Ichiro Naomi, Shûhei Nomura, Volker Puthz, Takashi Shimada, Yasuaki Watanabe, Shûhei Yamamoto

matobai Y. Watanabe, 2005a: 314 マトバコバネナガハネカクシ HON
mayasanense Y. Watanabe, 1992b: 189 マヤサンコバネナガハネカクシ HON
minakawai Y. Watanabe, 2004c: 41 エトロフコバネナガハネカクシ（和名改称）Etr
nagashimanum Y. Watanabe, 2005c: 597 ナガシマコバネナガハネカクシ HON
nanseiense Y. Watanabe, 2005c: 598 ナンセイコバネナガハネカクシ HON
nasuense Y. Watanabe, 1992a: 43 ナスコバネナガハネカクシ HON
nishikawai Y. Watanabe, 1986c: 692 ニシカワコバネナガハネカクシ HON
ohkurai Hayashi, 1996: 16 アマイシコバネナガハネカクシ（和名新称）HON
okamotoi Y. Watanabe, 2011b: 50 クレコバネナガハネカクシ HON
omogoense Y. Watanabe, 1991a: 33 オモゴコバネナガハネカクシ SHI
onodai Y. Watanabe, 1996: 219 コシキコバネナガハネカクシ SKsowaseanum Y. Watanabe, 2005c: 590 オワセコバネナガハネカクシ HON
pollens Sharp, 1889: 254 コバネナガハネカクシ HON
sanukiense Y. Watanabe, 1991a 29 サヌキコバネナガハネカクシ SHI
shingon Y. Watanabe, 1992b: 193 コウヤコバネナガハネカクシ HON
shizuokaense Y. Watanabe, 1986c: 694 シズオカコバネナガハネカクシ HON
shotaroi Y. Watanabe, 2005c: 593 ショウタロウコバネナガハネカクシ HON
susamiense Y. Watanabe, 2005a: 317 スサミコバネナガハネカクシ HON
susumui Y. Watanabe, 1984: 135 オバココバネナガハネカクシ HON
suzukii Y. Watanabe, 2011c: 181 ヒゼンコバネナガハネカクシ HON
tadaorum Y. Watanabe, 2008b: 187 タダコバネナガハネカクシ HON
taichii Y. Watanabe, 2008b: 184 タイチコバネナガハネカクシ HON
taishakuense Y. Watanabe, 2011b: 53 タイシャクキョウコバネナガハネカクシ HON
tokushimanum Y. Watanabe & Yoshida, 2009: 93トクシマコバネナガハネカクシ SHI
toyodai Y. Watanabe, 2005a: 321トヨダコバネナガハネカクシ HON
uozumii Y. Watanabe, 2002c: 13 ウオズミコバネナガハネカクシ SHI
yokozekii Y. Watanabe, 2005a: 319 ヨコゼキコバネナガハネカクシ HON
yosiianum Y. Watanabe, 1999a: 13 ヨシイコバネナガハネカクシ SHIGroup of L. brachypterum/monticola ヒメコバネナガハネカクシ種群
adachii Y. Watanabe, 2010c: 261 アダチヒメコバネナガハネカクシ HON
brachypterum Sharp, 1889: 255 ヒメコバネナガハネカクシ HON
isense Y. Watanabe, 2006a: 53 イセヒメコバネナガハネカクシ HON
kagaense Y. Watanabe & Hoshina, 2003: 8 カガヒメコバネナガハネカクシ HON
kasaharai Y. Watanabe, 2002a: 315 カサハラヒメコバネナガハネカクシ HON
kinokuniense Y. Watanabe, 2006a: 57 キノクニヒメコバネナガハネカクシ HON
koyasanum Y. Watanabe, 2006a: 48 コウヤヒメコバネナガハネカクシ HON
kurosawai Y. Watanabe, 2001c: 359 クロサワヒメコバネナガハネカクシ HON
masaoi Y. Watanabe, 1999c: 109 キイヒメコバネナガハネカクシ HONmasumotoi Y. Watanabe, 2011a: 103 マスモトヒメコバネナガハネカクシ HON
monticola Sharp, 1889: 255 ナガサキヒメコバネナガハネカクシ（和名改称）KYU

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Catalogue of Japanese Staphylinidae (Insecta: Coleoptera)nabetaniense Y. Watanabe, 1997a: 144 ナベタニヒメコバネナガハネカクシ HON
nankiense Y. Watanabe, 2006a: 55 ナンキヒメコバネナガハネカクシ HON
notoense Y. Watanabe, 1997a: 140 ノトヒメコバネナガハネカクシ HON
ohtohense Y. Watanabe, 2006a: 51 オウトウヒメコバネナガハネカクシ HON
okiense Y. Watanabe & Shimada, 2004: 294 オキヒメコバネナガハネカクシ（和名改称）Okisasajii Y. Watanabe, 2001b: 223 ササジヒメコバネナガハネカクシ HON
satoi Y. Watanabe, 2003a: 151 サトウヒメコバネナガハネカクシ（和名改称）HON
shinanense Y. Watanabe, 2011a: 106 シナノヒメコバネナガハネカクシ HON
shiritakanum Y. Watanabe, 1997a: 142 シリタカヒメコバネナガハネカクシ HON
sugiei Y. Watanabe, 1997a: 136 スギエヒメコバネナガハネカクシ HON
tamotsui Y. Watanabe, 1994: 349 イシハラヒメコバネナガハネカクシ（和名改称）SHI
wasamatanum Y. Watanabe, 2006a: 59 ワサマタヒメコバネナガハネカクシ HON
Group of L. tahirai チビコバネナガハネカクシ種群
kanayamaense Y. Watanabe, 2001d: 468 カナヤマチビコバネナガハネカクシ HON
nidoagense Y. Watanabe, 2001d: 471 ニドアゲチビコバネナガハネカクシ HON
nikkoense Y. Watanabe, 2001d: 472 ニッコウチビコバネナガハネカクシ HON
tahirai Y. Watanabe, 2001d: 465 チビコバネナガハネカクシ HON
Genus Nipponolathrobium Y. Watanabe, 2012 チカコバネナガハネカクシ属（和名新称）
daibosatsu T. Ito, 2009a: 28 (Lathrobium) ダイボサツチカナガハネカクシ（和名新称）HON
kitayamai T. Ito, 2009a 26 (Lathrobium)トビラチカナガハネカクシ（和名新称）HON
moritai Y. Watanabe, 2012c: 340 サンプクチカコバネナガハネカクシ（和名新称）HON
iyaense Y. Watanabe, 2012c: 343 イヤチカコバネナガハネカクシ（和名新称）SHI
yozawanum Y. Watanabe, 1980a: 24 (Lathrobium) チカコバネナガハネカクシ（和名改称）HON
Genus Lobrathium Mulsant & Rey, 1878 ツツナガハネカクシ属（和名新称）
Subgenus Lobrathium Mulsant & Rey, 1878
amamiense T. Ito, 1996d: 116 アマミツツナガハネカクシ（和名新称）Amm
cribricolle Sharp, 1889: 256 (Lathrobium) キモンツツナガハネカクシ（和名改称）HON, Okn, Kum
hokkaidense T. Ito, 1996a: 6 エゾツツナガハネカクシ（和名新称）HOK
isamutanakai T. Ito, 2009b: 33 タナカキモンナガハネカクシ HON
ishidai T. Ito, 1996a: 4 ナスツツナガハネカクシ（和名新称）HON
ishizuchiense T. Ito, 1996d: 110 イシヅチツツナガハネカクシ（和名新称）SHI
mizunoi T. Ito, 1996a: 3 ゴザイシツツナガハネカクシ（和名新称）HON
nipponense T. Ito, 1995b: 37 ホソクロツツナガハネカクシ（和名改称）HON, KYU
nudum Sharp, 1874a: 55 (Lathrobium) ツツナガハネカクシ（和名改称）HON, SHI, KYU, Yak; Russia
ohkurai T. Ito, 1996a: 1 ヨシノツツナガハネカクシ（和名改称）HON
okamotoi T. Ito, 1995b: 40 アワツツナガハネカクシ（和名改称）SHI
partitum Sharp, 1874a: 56 (Lathrobium) サキアカバツツナガハネカクシ（和名改称）HOK, HON, SHI, Iz-Hch,
KYU, Tsm; China

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Yasutoshi Shibata, Munetoshi Maruyama, Hideto Hoshina, Toshio Kishimoto, Shun-Ichiro Naomi, Shûhei Nomura, Volker Puthz, Takashi Shimada, Yasuaki Watanabe, Shûhei Yamamoto

riozoi Y. Watanabe, 1972: 117 (Lathrobium) チビアカツツナガハネカクシ（和名改称）HOK
ryukyuense T. Ito, 1996d: 114 リュウキュウツツナガハネカクシ（和名新称）Tan, T-Nak, Amm, Tok, Oke, Okn, Yon
sasajii T. Ito, 2007: 87 キイツツナガハネカクシ（和名改称）HON
shibatai shibatai T. Ito, 1995b: 41 ヤマトツツナガハネカクシ（和名新称）HON
s. varium T. Ito, 1995b: 43 HON
yoshidai yoshidai Adachi, 1955: 30 (Lathrobium) ヨシダツツナガハネカクシ（和名改称）HOK, Rsr
y. sadoensis Y. Watanabe & Baba, 1973: 260 (Lathrobium) Sad
Genus Platydomene Ganglbauer, 1895ドウガネナガハネカクシ属（和名新称）
anguina Sharp, 1874a: 57 (Lathrobium) アカチャドウガネナガハネカクシ（和名改称）HON; W Asia
carinicollis Sharp, 1889: 255 (Lathrobium) ヒラタドウガネナガハネカクシ（和名改称）HON, KYU
daibosatsuensis Y. Watanabe, 2009b: 245 ダイボサツドウガネナガハネカクシ HON
flavipes Y. Watanabe, 2009b: 248 キアシドウガネナガハネカクシ HON
funebris Sharp, 1889: 256 (Lathrobium) クロツヤドウガネナガハネカクシ（和名改称）HON, SHI
hakusana Y. Watanabe, 2008c: 332 ハクサンドウガネナガハネカクシ HON
hirogawarana Y. Watanabe, 2008c: 339 ヒロガワラドウガネナガハネカクシ HON
iidesana Y. Watanabe, 2009b: 251 イイデドウガネナガハネカクシ HON
kojimai Y. Watanabe, 2008c: 335 コジマドウガネナガハネカクシ HON
nikkoensis Y. Watanabe, 2008c: 336 ニッコウドウガネナガハネカクシ HON
nobilis Sawada, 1965: 11 (Lobrathium)ドウガネナガハネカクシ HON
Genus Pseudobium Mulsant & Rey, 1878 ニセナガハネカクシ属（和名新称）
yutakakimurai Hayashi, 1994a: 137 ニセナガハネカクシ（和名新称）HON
Genus Pseudolathra Casey, 1905 スジナガハネカクシ属（和名新称）
Subgenus Pseudolathra Casey, 1905
lineata Herman, 2003: 7 [RN] スジナガハネカクシ（和名新称）HON; Taiwan, China, Oriental Region= seriatum Sharp, 1889: 259 (Lathrobium)
regulare Sharp, 1889: 258 (Lathrobium) カクムネスジナガハネカクシ（和名改称）HON, SHI
unicolor Kraatz, 1859: 117 (Lathrobium) ツマグロスジナガハネカクシ（和名改称）HON, Og-ch, Oki, KYU, Tsm,
Yak, Amm, Okn, Kum; Taiwan, China, Bhutan, Nepal, Pakistan, India, Oriental Region
Genus Tetartopeus Czwalina, 1888 コガシラナガハネカクシ属（和名新称）
niger LeConte, 1863: 43 (Lathrobium) コガシラナガハネカクシ（和名改称）HON; Russia, USA (Alaska)= cognatus Sharp, 1889: 257 (Lathrobium)
fragilis Sharp, 1889: 258 (Lathrobium) ホソキアシコガシラナガハネカクシ（和名改称）HON; South Korea
fulvipes Adachi, 1955: 30 (Lathrobium) クロコガシラナガハネカクシ（和名改称）SHI
pallipes Sharp, 1889: 257 (Lathrobium) キアシコガシラナガハネカクシ（和名改称）HOK, HON Oki, Tsm, Okn;
South Korea, China, Russia

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Catalogue of Japanese Staphylinidae (Insecta: Coleoptera)

Subtribe Medonina Casey, 1905
Genus Charichirus Sharp, 1889 アジアトガリハネカクシ属（和名新称）
chinensis Boheman, 1858: 32 (Lathrobium) アジアトガリハネカクシ（和名新称）Amm, Okn; Taiwan, China, India,
Oriental Region
Genus Hypomedon Mulsant & Rey, 1878 ヒメトガリハネカクシ属（和名新称）
debilicornis Wollaston, 1857: 194 (Lithocharis) ヒメトガリハネカクシ（和名改称）HON, Iz-My, Iz-Hch, SHI, KYU
Tsm, Yak, Amm; Taiwan, South Korea, China, India, Cosmopolitan
Genus Isocheilus Sharp, 1889 ニセトガリハネカクシ属
staphylinoides Kraatz, 1859: 134 (Lithocharis) ニセトガリハネカクシ HON, Iz-My, Iz-Hch, Oki, SHI, Okns, KYU,
Tsm, Yak; Taiwan, South Korea, Russia, Singapore, India, Sri Lanka, Africa
Genus Lithocharis Dejean, 1833 クロズトガリハネカクシ属（和名新称）
dissimilis Sharp, 1874a: 66 ニセクロズトガリハネカクシ（和名新称）KYU, HON
nigriceps Kraatz, 1859: 139 クロズトガリハネカクシ HON, Sad, Iz-Hch, Iz-My, Iz-Mk, Og, Oki, SHI, KYU, Tan,Yak Amm, Okn, Iri, MDt, KDt; Taiwan, South Korea, China, India, Oriental Region
ochracea Gravehorst, 1802: 59 (Paederus) キバネクロズトガリハネカクシ（ 和 名 新 称 ）HON, KYU; China,
Kazakhstan, Turkey, Cosmopolitan
sororcula Kraatz, 1859: 140 セトクロズトガリハネカクシ（和名新称）SHI (Hakata-jima, Ô-shima); India, Oriental
Regionsubopaca Bernhauer, 1907: 382 (Medon) ツヤケシクロズトガリハネカクシ（和名新称）HON
uvida Kraatz, 1859: 138 アジアクロズトガリハネカクシ（和名新称）HON; South Korea, Taiwan, China, Singapore,
Indonesia (Sumatra), India, Sri Lanka
Genus Medon Stephens, 1833トガリハネカクシ属
basipennis Bernhauer, 1927: 93 ムネミゾトガリハネカクシ（和名新称）Japan, Oki
confertus Sharp, 1889: 265トガリハネカクシ（和名新称）KYU; Taiwan
discedens Sharp, 1889: 266 アカアシクロトガリハネカクシ HON
lewisius Sharp, 1874a: 65 (Lithocharis) ネアカトガリハネカクシ HON, Oki, Tsm, SHI; Taiwan
prolixus Sharp, 1874a: 65 (Lithocharis) ウミベトガリハネカクシ HON, Iz-My, Iz-Hch, SHI, KYU
rubeculus Sharp, 1889: 264 ムネアカトガリハネカクシ（和名新称）HOK, HON, SHI, Awj; South Korea
spadiceus Sharp, 1889: 265 チャバネトガリハネカクシ HON
submaculatus Sharp, 1889: 266 ヨコモントガリハネカクシ HON, Oki, SHI, KYU, Tsm, Yak; Taiwan
sulcifrons Bernhauer, 1907: 383 ミゾトガリハネカクシ HOK, HON
Genus Panscopaeus Sharp, 1889 マメトガリハネカクシ属（和名新称）
lithocharoides Sharp, 1874a: 63 (Scopaeus) マメトガリハネカクシ（和名改称）HON, Oki, SHI, KYU, Tsm; China,
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Yasutoshi Shibata, Munetoshi Maruyama, Hideto Hoshina, Toshio Kishimoto, Shun-Ichiro Naomi, Shûhei Nomura, Volker Puthz, Takashi Shimada, Yasuaki Watanabe, Shûhei Yamamoto

yakushimanus T. Ito, 1992a: 61 (Achenomorphus) ヤクマメトガリハネカクシ（和名新称）Yak; Taiwan, China
Genus Thinocharis Kraatz, 1859 ヒゲボソトガリハネカクシ属（和名新称）
basicornis Sharp, 1874a: 63 (Scopaeus) アカバヒゲボソトガリハネカクシ（和名新称）SHI, KYU
japonica Bernhauer, 1909c: 199 [RN] ヒゲボソトガリハネカクシ（和名新称）HON, SHI= brevicornis Bernhauer, 1907: 381 [HN]
Subtribe Paederina Fleming, 1821
Genus Megalopaederus Scheerpeltz, 1957 オオアリガタハネカクシ属（和名新称）
kurosawai Y. Watanabe, 1986b: 167 クロサワオオアリガタハネカクシ（和名改称）HON
lewisi Cameron, 1930: 206 (Paederus) ルイスオオアリガタハネカクシ（和名改称）HOK, HON
poweri Sharp, 1874a: 74 (Paederus) オオアリガタハネカクシ（和名改称）HON
wadai Scheerpeltz, 1957: 472 ワダオオアリガタハネカクシ（和名改称）HON, Shd, KYU
Genus Paederus Fabricius, 1775 アリガタハネカクシ属
Subgenus Eopaederus Scheerpeltz, 1957
tamulus Erichson, 1840: 661 アシグロアリガタハネカクシ KYU; Taiwan, China, Bhutan, Singapore, Indonesia (Java,
Sumatra), India, Sri Lanka
Subgenus Heteropaederus Scheerpeltz, 1957
fuscipes Curtis, 1826: pl.108 アオバアリガタハネカクシ HOK, Rsr, HON, Sad, Iz-My, Iz-Hch, Oki, SHI, KYU, Tsm,
Tan, Yak, T-R, Amm, Okn, Kum, Ish, Iri, Yon, KDt, MDt; Taiwan, China, South Korea, North Korea, Palearctic
Region, Australia, Africa= idae Sharp, 1874a: 75
Subgenus Paederus Fabricius, 1775
angustipennis Bernhauer, 1936: 240 ホソバアリガタハネカクシ（和名新称）HON
japonicus Bernhauer, 1915b: 265 ニホンアリガタハネカクシ（和名新称）HON
parallelus Weise, 1877b: 368 エゾアリガタハネカクシ HOK, Chs, Rsr, HON, Sad; South Korea, North Korea, China
Incertae sedis (subgenus?) 亜属不明
jessoensis Bernhauer, 1935: 44 キタアリガタハネカクシ（和名新称）HOK
kozuensis Cameron, 1930: 206 コウズアリガタハネカクシ（和名新称）HON
Subtribe Scopaeina Mulsant & Rey, 1878
Genus Scopaeus Erichson, 1839 ヒメクビボソハネカクシ属
complex Sharp, 1874a: 61 ヤマトヒメクビボソハネカクシ（和名新称）Japan
currax Sharp, 1889: 261 ヒメクビボソハネカクシ HON, Oki, SHI, Awj; China
curraxoides Adachi, 1955: 26 ニセヒメクビボソハネカクシ（和名新称）HON
filiformis Wollaston, 1867: 243 ホソヒメクビボソハネカクシ（和名新称）Japan; Taiwan, China, Nepal, Oriental
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Catalogue of Japanese Staphylinidae (Insecta: Coleoptera)japonicus Cameron, 1933: 169 マヤサンヒメクビボソハネカクシ（和名新称）HON
limbatus Kraatz, 1859 キベリヒメクビボソハネカクシ HON, Okn, Ish; China, Afghanistan, Nepal, North Pakistan,
Oriental Region
naomii Frisch, 2003: 688 キタヒメクビボソハネカクシ（和名新称）HON
nipponensis Frisch, 2003: 664 ニホンヒメクビボソハネカクシ（和名新称）HON
paliferus Frisch, 2011 ヘラヒメクビボソハネカクシ HOK, HON, KYU; Russia (East Siberia)schuelkei Frisch, 2003: 686 シュルケヒメクビボソハネカクシ（和名新称）HON, KYU, Yak; Taiwan, Hong Kong,
China, SE Asia
testaceus Motschulsky, 1858b: 642 アカヒメクビボソハネカクシ（和名新称）HOK, HON, SHI, KYU, Okn; Taiwan,
China, India, SE Asia
= quadraticeps Bernhauer, 1938a: 36= rufescens Adachi, 1955: 26
trapeziceps Frisch, 2003: 665 アジアヒメクビボソハネカクシ（和名新称）HON; South Korea, China
viriliformis Frisch, 2003: 670 コバネヒメクビボソハネカクシ（和名新称）HON, SHI, MIw
virilis Sharp, 1874a: 62 チビヒメクビボソハネカクシ（和名改称）HON, Iz-Mk, Iz-Hch, Oki, SHI, Okns, Tan, Yak,
Okn; China, Oriental Region
yoshidai Adachi, 1955: 27 カナガワヒメクビボソハネカクシ（和名新称）HON
Subtribe Stilicina Casey, 1905
Genus Rugilus Leach, 1819 クビボソハネカクシ属
Subgenus Eurystilicus Fagel, 1953
ceylanensis Kraatz, 1859: 126 (Stilicus) キバネクビボソハネカクシ HON, Iz-My, Iz-Hch, SHI, KYU, Yak, Amm,
Okn, Ish, Yon; Taiwan, South Korea, North Korea, China, India, Oriental Region
rufescens Sharp, 1874a: 61 (Stilicus) クビボソハネカクシ HOK, HON, Iz-My, Iz-Mk, Iz-Hch, Oki, SHI, KYU, Tsm,
Yak; Taiwan, South Korea, North Korea, China, Russia, Oriental Region
Subgenus Rugilus Leach, 1819
japonicus Y. Watanabe, 1961: 351 ツマキクビボソハネカクシ Iz-My, Iz-Mk, Iz-Hch, Iz-Mk, Tan, Yak, Amm, Okn
longipennis Sharp, 1889: 321 (Stilicus) ハネナガクビボソハネカクシ HOK, Rsr, HON, SHI; South Korea, China
Incertae sedis (subgenus?) 亜属不明
subtilis Erichson, 1840: 631 クロクビボソハネカクシ HON; Europe
Genus Stilicoderus Sharp, 1889 オオクビボソハネカクシ属
japonicus Shibata, 1968: 8 ヤマトオオクビボソハネカクシ（和名改称）HOK, HON, SHI; China
signatus Sharp, 1889: 321 オオクビボソハネカクシ HOK, HON, SHI, Tsm, Yak; China
Genus Stiliderus Motschulsky, 1858 フトクビボソハネカクシ属
duplicatus T. Ito, 1984: 59 (Stilicoderus) フトクビボソハネカクシ Amm
Subtribe Stilicopsina Casey, 1905

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Yasutoshi Shibata, Munetoshi Maruyama, Hideto Hoshina, Toshio Kishimoto, Shun-Ichiro Naomi, Shûhei Nomura, Volker Puthz, Takashi Shimada, Yasuaki Watanabe, Shûhei Yamamoto

Genus Stilicopsis Sachse, 1852 アミメクビボソハネカクシ属（和名新称）
insulicola T. Ito, 1992a: 59 アミメクビボソハネカクシ（和名新称）Yak, Tok
Genus Enallagium Bernhauer, 1915 ヤマトコマルズハネカクシ属（和名新称）
japonicum Nakane & Sawada, 1960: 122 ヤマトコマルズハネカクシ（和名新称）HON, SHI, Yak, Oke
Genus Ophryomedon Wasmann, 1916 フチドリツヤケシハネカクシ属（和名新称）
crenatus Wasmann, 1916: 204 フチドリツヤケシハネカクシ（和名新称）HON, KYU; China, Malaysia, India= marginatus Naomi, 1995c: 162
Genus Sunesta Blackwelder, 1939 タチゲクビボソハネカクシ属
setigera Sharp, 1874a: 67 (Acanthoglossa) タチゲクビボソハネカクシ HON, Iz-My, Iz-Mk, Iz-Hch, Og, SHI, KYU,
Tan, Tsm, T-Yok, Amm; Taiwan, South Korea, Thailand
Tribe Pinophilini Nordmann, 1837
Subtribe Pinophilina Nordmann, 1837
Genus Pinophilus Gravenhorst, 1802 クビブトハネカクシ属
formosae Bernhauer, 1935: 42 クビブトハネカクシ（和名新称）Irb, Trm, Ish, Yon; Taiwan, Oriental Region
javanus Erichson, 1840: 672 クロクビブトハ ネカクシ HON, KYU, Tsm, Ish; China, Sunda, Indonesia (Java,
Sumatra), India, Sri Lanka= insignis Sharp, 1874a: 77
lewisius Sharp, 1874a: 77 ルイスクビブトハネカクシ HON, Iz-My, Iz-Mk, Iz-Hch, SHI, KYU, Tsm, Tan, Yak;
Taiwan, South Korea
punctatissimus Sharp, 1889: 323 オオクビブトハネカクシ HON, KYU, Tan, Yak, Ish, Iri; China
rufipennis Sharp, 1874a: 78 アカバクビブトハネカクシ HON, Iz-Hch, Sad, KYU, Yak, T-R, Amm, Tok, Okn, Myk,Iri, Yon; South Korea, China, Iransauteri Bernhauer, 1935: 42 ゾウタークビブトハネカクシ（和名新称）Okn, Kum; Taiwan
Subtribe Procirrina Bernhauer & Schubert, 1912
Genus Oedichirus Erichson, 1839 クロバネアリガタハネカクシ属（和名新称）
Subgenus Oedichirus Erichson, 1839
chapmani Cameron, 1940: 250 チャップマンクロバネアリガタハネカクシ（和名新称）Ish; Vietnam
idae Sharp, 1874a: 76 ナガクロバネアリガタハネカクシ HON, KYU, Tok
kiuchii Sawada, 1964a: 36 シコククロバネアリガタハネカクシ（和名新称）SHI
kuroshio Hayashi, 1989: 161 クロシオクロバネアリガタハネカクシ（和名新称）Iri; Taiwan (Lanyu)
lewisius Sharp, 1874a: 76 クロバネアリガタハネカクシ HON, Aws, Iz-Mk, Iz-Hch, Oki, SHI, KYU, Tsm, Tan, Yak,
Okn; South Korea, North Korea

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日本産ハネカクシ科総目録（昆虫綱：甲虫目）
Catalogue of Japanese Staphylinidae (Insecta: Coleoptera)

Genus Palaminus Erichson, 1839 アラハダドウナガハネカクシ属（和名新称）
formosae Cameron, 1949: 465 タイワンアラハダドウナガハネカクシ Yak, Amm, Okn; Taiwan
japonicus Cameron, 1930: 206 アラハダドウナガハネカクシ HON, Iz-My, Iz-Hch, SHI, Okns; South Korea, Taiwan
Genus Procirrus Latreille, 1829 クビナガハネカクシ属（和名新称）
Subgenus Procirrus Latreille, 1829
lewisii Sharp, 1889: 324 クビナガハネカクシ HON, Og, SHI, KYU, Kue, Tok, Ish; Thailand, Oriental Region

Subfamily Staphylininae Latreille, 1802 ハネカクシ亜科（59属288種３亜種）
Tribe Diochini Casey, 1906
Genus Diochus Erichson, 1839 コガシラホソハネカクシ属
japonicus Cameron, 1930: 206 コガシラホソハネカクシ HON, Oki, SHI, KYU, Tsm, Amm; South Korea, China
Tribe Othinini Thomson, 1859
Genus Atrecus Jacquelin du Val, 1856 カタアカホソハネカクシ属
pilicornis Paykull, 1790: 135 (Staphylinus) カタアカホソハネカクシ（和名改称）HOK, HON; China, Mongolia,
Russia (Siberia), Europe
Genus Othius Stephens, 1829 ホソハネカクシ属
amamianus T. Ito, 1993b: 144 アマミホソハネカクシ（和名新称）Amm
latus Sharp, 1874a: 51トビイロホソハネカクシ HOK, HON, KYU; Russia (Sakhalin), China= ozakii T. Ito, 1993b: 143
medius Sharp, 1874a: 50 ウスアカバホソハネカクシ HOK, Oks, HON, Sad, SHI, KYU, Tsm, Yak; South Korea,
China= kusuii T. Ito, 1993b: 147= yakushimanus T. Ito, 1993b: 148
rosti Bernhauer, 1907: 385 ロストホソハネカクシ（和名新称）HOK, HON; North Korea, Chs, Russia (Sakhalin,
Maritime, Ussuri)
rufipennis Sharp, 1874a: 49 アカバホソハネカクシ HOK, Sad, KYU; South Korea, North Korea, China
Tribe Staphylinini Latreille, 1802
Subtribe Amblyopinina Seevers, 1914
Genus Heterothops Stephens, 1829 チビツヤムネハネカクシ属
cognatus Sharp, 1874a: 20 ホソチビツヤムネハネカクシ（和名改称）HOK, HON, Aws, Sad, Iz-My, Iz-Mk, Iz-Hch,2013 The Kyushu

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Yasutoshi Shibata, Munetoshi Maruyama, Hideto Hoshina, Toshio Kishimoto, Shun-Ichiro Naomi, Shûhei Nomura, Volker Puthz, Takashi Shimada, Yasuaki Watanabe, Shûhei Yamamoto

SHI, KYU, SKs, Yak; South Korea, China
ishiharai T. Ito, 1994d: 177 アバタチビツヤムネハネカクシ（和名新称）HON; China
rotundiceps Sharp, 1889: 35 オオメチビツヤムネハネカクシ HON, Yak; South Korea
Subtribe Anisolina Hayashi, 1993
Genus Amichrotus Sharp, 1889 ズマルハネカクシ属
apicipennis Sharp, 1889: 115 ズマルハネカクシ HOK, HON, Sad, Oki, SHI, KYU; Taiwan, China
Genus Anisolinus Sharp, 1889 ブチヒゲハネカクシ属
elegans Sharp, 1889: 114 ツヤケシブチヒゲハネカクシ HOK, HON, Oki, SHI, KYU
hayashii Sawada, 1961b: 4 クロブチヒゲハネカクシ HON
picticornis Sharp, 1889: 113 カタモンブチヒゲハネカクシ HONtaoi Naomi, 1981: 113 ミヤマブチヒゲハネカクシ HON
tsurugiensis Sawada, 1961: 5 ハネアカブチヒゲハネカクシ SHIGenus Hesperosoma Scheerpeltz, 1965 キバナガマルズハネカクシ属（和名新称）
excellens Bernhauer, 1939a: 100 (Amichrotus) キバナガマルズハネカクシ（和名新称）HON(?); China, Vietnam
Genus Tympanophorus Nordmann, 1837 ツヤケシアバタハネカクシ属
hayashidai Shibata, 1975: 21 マルズツヤケシアバタハネカクシ（和名改称）HON, KYU; South Korea, China
sauteri Bernhauer, 1907: 385 カクツヤケシアバタハネカクシ HON, Sad, SHI, KYU, Tan, Yak; South Korea, China
Subtribe Philonthina Kirby, 1837
Genus Belonuchus Nordmann 1837 ヒラズハネカクシ属
rufoniger Fauvel, 1895: 266 アカ バ ヒラ ズ ハ ネカクシ KYU; Taiwan, Hong Kong, China, Indonesia (Java),
Myanmar, India (Assam), West Indies
Genus Bisnius Stephens, 1829 ニセコガシラハネカクシ属（和名新称）
germanus Sharp, 1874a: 41 (Philonthus) チャイロニセコガシラハネカクシ（和名改称）HON, Iz-My, Iz-Mk, Iz-Hch,Oki, SHI, KYU, SKs, Yak, Ammmacies Sharp, 1874a: 41 (Philonthus) マルズニセコガシラハネカクシ（和名改称）SHI, Shd, KYU, Tsmparcus Sharp, 1874a: 40 (Philonthus) オオズニセコガシラハネカクシ（和名改称）HOK, HON, SHI, Shd; China,
Mongolia, Australia, Europe
sordidus Gravenhorst, 1802: 176 (Philonthus) クロニセコガシラハネカクシ（ 和 名 新 称 ）Japan; China, Russia,
Mongolia, India, Nepal, Europe, Cosmopolitan
Genus Cafius Stephens, 1832 アバタウミベハネカクシ属（和名新称）
Subgenus Pseudoremus Blackwelder, 1943

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日本産ハネカクシ科総目録（昆虫綱：甲虫目）
Catalogue of Japanese Staphylinidae (Insecta: Coleoptera)algarum Sharp, 1874a: 38 (Philonthus) ヒメアバタウミベハネカクシ（和名改称）HON, Sad, Iz-My, Iz-Mk, Iz-Hch,Oki, SHI, KYU, Amm; South Korea, Hong Kong, China, Russia
histrio Sharp, 1874a: 31 (Philonthus) ホソアバタウミベハネカクシ HOK, HON, Sad, Iz-My, Iz-Mk, Iz-Hch, SHI,
Tan, Yak; South Korea, North Korea, Hong Kong, Russia
mimulus Sharp, 1874a: 38 (Philonthus) ヒラタアバタウミベハネカクシ（和名新称）HON, KYU; South Korea
nauticus Fairmaire, 1849: 288 (Philonthus) ミナミアバタウミベハネカクシ Amm, Okn, Trm, Ish, Iri; Taiwan, Hong
Kong, Vietnam, Pan Tropics
rufescens Sharp, 1889: 44 アカバアバタウミベハネカクシ（和名改称）HOK, HON, Oki, SHI, Shd, KYU, Tsm, Tan,Amm, Iri; South Korea, Hong Kong, China, Vietnam
vestitus Sharp, 1874a: 37 (Philonthus) オオアバタウミベハネカクシ（和名改称）Reb, HON, Aws, Sad, Awj, Oki,
Shd, SHI, KYU, Tsm, Tan, Yak; South Korea, North Korea, Russia
Genus Erichsonius Fauvel, 1874 ムネスジミギワハネカクシ属（和名新称）
Subgenus Erichsonius Fauvel, 1874
watanabei Uhlig, 1992: 72 オオヌマムネスジミギワハネカクシ（和名新称）HOK, HON
Subgenus Sectophilonthus Tottenheim, 1949
hokkaidoensis Uhlig & Y. Watanabe, 1992: 24 キタムネスジミギワハネカクシ（和名新称）HOK
itoi Uhlig & Y. Watanabe, 1992: 39 オオダイムネスジミギワハネカクシ（和名新称）HON
japonicus Cameron, 1933: 170 (Actobius) ツヤムネスジミギワハネカクシ（和名改称）HOK, HON, SHI, KYU
kobensis Cameron, 1933: 169 (Actobius) クロムネスジミギワハネカクシ（和名改称）HON
naomii Uhlig & Y. Watanabe, 1992: 66 リュウキュウムネスジミギワハネカクシ（和名新称）Ish; Taiwan
sawadai Uhlig & Y. Watanabe, 1992: 45 サワダムネスジミギワハネカクシ（和名新称）HOK, HON, SHI
shibatai Y. Watanabe & Uhlig, 1992: 28 タンザワムネスジミギワハネカクシ（和名新称）HON
sobosanus Y. Watanabe, 1963a: 8 ソボムネスジミギワハネカクシ（和名改称）HON, SHI, KYU
vulgaris Uhlig & Y. Watanabe, 1992: 58 ユノコヤムネスジミギワハネカクシ（和名新称）HON
Genus Gabrius Stephens, 1829 ホソコガシラハネカクシ属
abas Smetana, 1984a 135 アバスホソコガシラハネカクシ（和名新称）HON, SHI, Tsm
damon Smetana, 1984a: 135 ダモンホソコガシラハネカクシ（和名新称）HON
demades Smetana, 1984a: 138 デマーデスホソコガシラハネカクシ（和名新称）HON, SHI
egens Sharp, 1874a: 44 (Philonthus) チャバネホソコガシラハネカクシ（和名新称）HON, SHI; South Korea (Cheju-do)
gelo Smetana, 1984b: 649 ニッコウホソコガシラハネカクシ（和名新称）HON
io Smetana, 1984a: 133 イオホソコガシラハネカクシ（和名新称）HON
japonicus Shibata, 1991: 90 ヤマトホソコガシラハネカクシ HOK, HON, SHI
kobayashii Smetana, 1984a: 128 コバヤシホソコガシラハネカクシ HON
kuanshanensis Bernhauer, 1914c: 66 (Philonthus) カンザンホソコガシラハネカクシ（和名新称）HOK; HON, China
lackneri Schillhammer, 2011: 140トウホクホソコガシラハネカクシ（和名新称）HON
nepos Smetana, 1984a: 146 ネポスホソコガシラハネカクシ（和名新称）HON, SHI
nigritulus Gravenhorst, 1802: 41 (Staphylinus) ナガサキホソコガシラハネカクシ（和名新称）HON, SHI, KYU;
China, India, Nepal, Mongolia, Russia, Europe, Cosmopolitan

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Yasutoshi Shibata, Munetoshi Maruyama, Hideto Hoshina, Toshio Kishimoto, Shun-Ichiro Naomi, Shûhei Nomura, Volker Puthz, Takashi Shimada, Yasuaki Watanabe, Shûhei Yamamoto

okamotoi Shibata, 1991: 85 オカモトホソコガシラハネカクシ HON, SHI
ophion Smetana, 1984a: 146 オフィオンホソコガシラハネカクシ（和名新称）HOK, HON, SHI, Tsm; Chs, South
Korea, North Korea= gladipenis Cho & Lee, 1997: 275
philo Smetana, 1984a: 140 ファイロホソコガシラハネカクシ（和名新称）HON, SHI
praesignis Schillhammer, 2001: 394 タイワンホソコガシラハネカクシ（和名新称）HON, SHI; Taiwan, South Korea
schillhammeri Shibata, 1996: 146 [RN] オオホソコガシラハネカクシ HON= multipunctatus Shibata, 1991: 88 [HN]
septempunctatus Cameron, 1930: 207 (Philonthus) コウベホソコガシラハネカクシ（和名新称）HON, SHI, KYU
sharpianus Cameron, 1930: 207 (Philonthus) シャープホソコガシラハネカクシ HON, Oki, SHI, KYU, Tsm; China= conicus Bernhauer, 1939a: 99 (Philonthus)= demarcates Tottenham, 1940: 68
subdepressus Bernhauer, 1939a: 97 (Philonthus) ヒラタホソコガシラハネカクシ HOK, HON, SHI, KYU; China
unzenensis Bernhauer, 1939b: 153 (Philonthus) コゲチャホソコガシラハネカクシ HON, KYU; South Korea, Russia
yamanei Smetana, 1984a: 130 ヤマネホソコガシラハネカクシ HON
Genus Gabronthus Tottenham, 1955 チビコガシラハネカクシ属（和名新称）
maritimus Motschulsky, 1858b: 661 (Gabrius) ミゾナシチビコガシラハネカクシ（和名新称）HON, SHI, Yak, Ish;
Taiwan, Israel, Lebanone, Saudi Arabia, Europe
pygmaeus Kraatz, 1859: 93: (Philonthus) ツヤチビコガシラハネカクシ（和名新称）Japan; Oriental Region
sulcifrons Sharp, 1889: 39 (Philonthus) タテミゾチビコガシラハネカクシ（和名改称）HON, Sad, Iz-My, Iz-Hch, Oki
SHI, KYU, Tsm, Yak, Amm, Iri; Taiwan, South Korea, Hong Kong, China, New Zealand, Finland
thermarum Aubé, 1850: 316 (Philonthus) ズグロチビコガシラハネカクシ（和名新称）HON, SHI, Okn; Taiwan,
China, Russia, Europe, Cosmopolitan
Genus Hesperus Fauvel, 1874 ムネスジハネカクシ属（和名新称）
ignoratus T. Ito, 1994b: 81 クロツヤムネスジハネカクシ（和名新称）HON, SHI, Tsm; Taiwan
ornatus Sharp, 1889: 43 タテミゾムネスジハネカクシ（和名改称）HON, SHI, KYU
tiro Sharp, 1874a: 49 (Philonthus) ツマグロムネスジハネカクシ（和名改称）HOK, HON, Oki, SHI, Awj, Shd,
KYU; China
Genus Mentophilonthus Levasseur, 1966 オオメコガシラハネカクシ属（和名新称）
notabilis Kraatz, 1859: 79 (Philonthus) スソアカオオメコガシラハネカクシ（和名改称）T-Tak, Kik, Okn, Izn, Trm,Iri, Hat; Taiwan, Hong Kong, Nepal, Philippines, IriIndia, Sri Lanka
Genus Neobisnius Ganglbauer, 1895 ヒメホソハネカクシ属
inornatus Sharp, 1889: 43 (Actobius) スソアカヒメホソハネカクシ HON, Iz-Hch, Oki, Tsm, Yak, Amm, Okn; South
Korea, China
praelongus Gemminger & Harold, 1868: 591 (Philonthus) ミナミヒメホソハネカクシ（和名新称）Ish, Iri; Taiwan,
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Catalogue of Japanese Staphylinidae (Insecta: Coleoptera)

pumilus Sharp, 1874a: 49 (Philonthus) アカバヒメホソハネカクシ HOK, HON, Sad, Iz-Hch, Oki, SHI, KYU, Tsm;
Taiwan, South Korea, Hong Kong, China, Russia
Genus Philonthus Stephens, 1829 コガシラハネカクシ属
Subgenus Onychophilonthus Neresheimer & Wagner, 1924
bicaudus Sharp, 1889: 36 オオトゲツメコガシラハネカクシ（和名新称）HON
lederi Eppelsheim, 1893: 49 シレトコトゲツメコガシラハネカクシ（和名新称）HOK, HON; China, Russia, Russia(Siberia), Mongolia, Europe
nakanei Sawada, 1965: 14トゲツメコガシラハネカクシ HOK, HON, SHI; China
tarsalis Smetana, 1963: 412 キタトゲツメコガシラハネカクシ（和名新称）HON; Chs, Russia
Subgenus Philonthus Stephens, 1829
addendus Sharp, 1867: 440 ズマルオオコガシラハネカクシ（和名改称）HOK, HON; South Korea, North Korea,
China, Mongolia, Russia (Siberia), Europe
aeneipennis Boheman, 1858: 30 ドウガネコガシラハネカク HON, Sad, Iz-My, Og, Oki, SHI, Shd, KYU, Tsm, Got,
Tan, Yak, Yor, KDt, MDt; South Korea, China, Taiwan, Malaysia, Indonesia (Sumatra), India, Sunda, New Guinea,
Mauritiusalbilabris Nordmann, 1837: 86 ツヤキアシコガシラハネカクシ（和名新称）HOK, HON, SHI, Amm; Hong Kong,
Russia= frater Bernhauer, 1907: 387
amamiensis Hayashi, 1993: 78 アマミヒラタコガシラハネカクシ（和名新称）Amm, Tok
amicus Sharp, 1874a: 45 ズホソヒゲナガコガシラハネカクシ（和名新称）HON, Iz-N, Iz-Hch, Tob, Sad, Oki, KYU,
Yak, Oke, Yor, Kum; Hong Kong, China, Russia
aokii R. Dvořák, 1958: 135 アオキコガシラハネカクシ（和名新称）HON, SHI
azabuensis R. Dvořák, 1958: 137 チャイロキヌコガシラハネカクシ HOK, HON, SHI
caeruleipennis Mannerheim, 1830: 27 (Staphylinus) ルリコガシラハネカクシ HOK, HON, Sad, Oki, SHI, KYU,
Tsm; Chs, South Korea, Canada, USA
chujoi R. Dvořák, 1958: 138 チュウジョウコガシラハネカクシ（和名新称）HON; South Korea
cunctator Sharp, 1889: 41ドウバネコガシラハネカクシ HOK, HON, SHI; South Korea
daimio Sharp, 1889: 42 エゾスジコガシラハネカクシ HOK, HON; South Korea
debilis Gravenhorst, 1802: 35 (Philonthus) ヒゲブトコガシラハネカクシ（和名新称）HOK, HON; China, Russia,
Europe, USA
decoloratus Kirshenblat, 1933: 101 ツヤクロオオコガシラハネカクシ（和名改称）HOK, HON, SHI; South Korea,
China, Mongolia, Russia
depressipennis Sharp, 1889: 37 ヒラタカクコガシラハネカクシ HOK, HON, SHI, KYU; South Korea
discoideus Gravenhorst, 1802: 38 チビカクコガシラハネカクシ HON, Iz-Hch, Og, Iw, KIw, SHI, Tan, Oke, Yor,
Okn, Ish, Iri; Hong Kong, China, Russia, Cosmopolitan
discrepens Sharp, 1889: 38 ツヤアカバコガシラハネカクシ HON
eidmannianus Scheerpeltz, 1929b: 121 アイドマンコガシラハネカクシ（和名新称）KYU, Tan; China, Russia= diallus Tottenham, 1953: 145
flavipes Kraatz, 1859: 88 キアシキヌツヤコガシラハネカクシ（和名新称）Amm; Hong Kong, China, SE Asia

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Yasutoshi Shibata, Munetoshi Maruyama, Hideto Hoshina, Toshio Kishimoto, Shun-Ichiro Naomi, Shûhei Nomura, Volker Puthz, Takashi Shimada, Yasuaki Watanabe, Shûhei Yamamoto

flavocinctus Motschulsky, 1858i: 663 クロズマルコガシラハネカクシ（和名新称）Og, Amm, Ish; Taiwan, Hong
Kong, China, India, Sri Lanka, Mauritius
freyi Bernhauer, 1939c: 591 ヒメチャバネコガシラハネカクシ（和名新称）HON; China
gastralis Sharp, 1874a: 48 チャバネコガシラハネカクシ HOK, Rsr, HON, SHI, Okns, KYU, Tsmgemellus Kraatz, 1859: 91 タテミゾコガシラハネカクシ（和名新称）Ish; Hong Kong, China, India, Sunda
geminus Kraatz, 1859: 87 フチホソアカバコガシラハネカクシ（和名新称）Amm; Hong Kong, China, Oriental Region
havelkai R. Dvořák, 1958: 140 ハベルカコガシラハネカクシ（和名新称）HON, SHI; South Korea
japonicus Sharp, 1874a: 40 クロオオコガシラハネカクシ（和名改称）HOK, HON, Awj, SHI, KYU, Yak, SKs; South
Korea, China, Chs, Russia (Siberia, Sakhalin)= bernhaueri Roubal, 1909c: 374 [HN]
= binderi Roubal, 1910: 263 [RN]
jonenensis Sawada, 1965: 12 タカネコガシラハネカクシ（和名新称）HOK, HON
kamimurai Sawada, 1965: 15 (Paragabrius) カミムラコガシラハネカクシ HOK, HON
kobensis Sharp, 1874a: 46 フタイロコガシラハネカクシ HON, Oki, SHI, KYU
lewisius Sharp, 1874a: 42 オオドウガネコガシラハネカクシ HON, Sad, Iz-N, Iz-Hch, Oki, SHI, KYU, Tsm, Yak,
T-Nak, Amm, Okn, Kum, Iri, Hat; Taiwan, South Korea, Hong Kong, China, Vietnam
liopterus Sharp, 1889: 36 コゲ チャコガシラハネカクシ HOK, HON, SHI, KYU, Tsm; South Korea, Russia,
Tajikistan
longicornis Stephens, 1832: 237 ヒゲナガコガシラハネカクシ HOK, HON, SHI, Shd, KYU; Taiwan, South Korea,
Hong Kong, China, Russia, Cosmopolitan= scybalarius Nordmann, 1837: 94
macrocephalus Sharp, 1889: 42 アカヒラタコガシラハネカクシ HON, Iz-Mk, KYU
micanticollis Sharp, 1889: 36 ニジムネコガシラハネカクシ HOK, HON, SHI, KYU; South Korea, China
minimus Cameron, 1933: 170 ツヤヒメコガシラハネカクシ（和名新称）HON
minutus Boheman, 1848: 283 ゴミコガシラハネカクシ HON, Sad, Og, SHI, Ish, Iri; Taiwan, South Korea, Hong Kong,
China, Oriental Region, Europe, Africa= mutans Sharp, 1874a: 43
nudus Sharp, 1874a: 36 ツヤウミベコガシラハネカクシ（和名改称）HOK, Reb, HON, Sad, Iz-Hch, Oki, SHI, Shd,
KYU, SKs; South Korea, North Korea, Chs, Russia, Canada, USA= grossepunctatus Bernhauer, 1939b: 154
numata R. Dvořák, 1958: 139 キアシチビコガシラハネカクシ HOK, HON, Oki, SHI, KYU; South Korea, Chinaoberti Eppelsheim, 1889: 174 ニセクロオオコガシラハネカクシ（和名改称）HOK, HON, SHI, KYU; South Korea,
North Korea, China, Chs, Russia, Mongolia= pseudojaponicus Bernhauer, 1936: 307
ohizumi R. Dvořák, 1958: 136 オオイズミコガシラハネカクシ（和名新称）HON, SHI, Yak; South Korea
parvicornis Gravenhorst, 1802: 23 (Staphylinus) ヒメヒゲナガコガシラハネカクシ（和名新称）HON; Taiwan, China,
Russia, India, Europe= agilis Gravenhorst, 1806: 77 (Staphylinus)
productus Kraatz, 1859: 89 チビカドマルコガシラハネカクシ（和名新称）Okn, Ish, Yon; Taiwan, Hong Kongprolatus prolatus Sharp, 1874a: 46 チャアシヒラタコガシラハネカクシ（和名新称）HON, KYU, SHI; Australian Region2013 The Kyushu

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Catalogue of Japanese Staphylinidae (Insecta: Coleoptera)p. ezoensis Hayashi, 1993: 75 Oks
quediodes Sharp, 1874a: 38 デメコガシラハネカクシ（和名新称）KYU
quisquiliarius Gyllenhal, 1810: 335 (Staphylinus) ヒメクロコガシラハネカクシ（和名新称）HON, Og, KYU, Okn,
Iri; China, Russia, Europe, Cosmopolitan
rectangulus Sharp, 1874a: 42 カクコガシラハネカクシ HOK, HON, Sad, Iz-O, Iz-Hch, Og, Oki, SHI, KYU, Dnj,
Tsm, Yak, Kum, KDt, MDt; Taiwan, South Korea, Hong Kong, China, Tahiti, Europe, USA
rotundicollis Ménétriés, 1832: 145 (Staphylinus) アオドウバネコガシラハネカクシ（和名新称）HOK, HON; South
Korea, China, Europe
spadiceus Sharp, 1889: 41 ヒラタコガシラハネカクシ HOK, HON, Sad, SHI, KYU, Tsm; South Korea, China
spinipes Sharp, 1874a: 39 オオアカバコガシラハネカクシ HOK, HON, Iz-O, SHI, KYU, Tsm, Yak, Trm; South
Korea, Taiwan, China, Russia, E Asia, Europe
strandi Smetana, 1959: 161 [RN] ストランドコガシラハネカクシ（和名新称）HOK, HON; Russia (Siberia), Mongolia= jenseni A. Strand, 1941: 36 [HN]
sublucanus Herman, 2001: 48 [RN] キヌコガシラハネカクシ HON, Oki, Iz-My, SHI, KYU, Yak, Amm; South
Korea, China= sericans Sharp, 1874a: 45 [HN]subvarians Sawada, 1965: 13 アシナガコガシラハネカクシ HOK, HON, SHI; China
tardus Kraatz, 1859: 84 ヘリアカバコガシラハネカクシ HON, Sad, Iz-My, Iz-Mk, Iz-Hch, Oki, SHI, Shd, KYU,
Tsm, SKs, Yak, Okn, Trm; Taiwan, China, West India, Sri Lanka= solidus Sharp, 1874a: 43
variipennis Kraatz, 1859: 85 ホソアカバコガシラハネカクシ（和名新称）SHI, Amm, Kik, KDt; Taiwan, Hong Kong,
China
virgatus Sharp, 1889: 38 クロスジコガシラハネカクシ HON, SHI; South Korea, China, Russia
wuesthoffi Bernhauer, 1939a: 98 ニセヒゲナガコガシラハネカクシ（和名改称）HOK, HON, Iz-Hch, Sad, Oki, SHI,KYU, SKs, Yak, Yon; South Korea, North Korea, China, Russia (Chs, Sakhalin)yokohamus R. Dvořák, 1958: 138 ヨコハマコガシラハネカクシ HON, SHI
Genus Pseudohesperus Hayashi, 2008 ムネスジコガシラハネカクシ属
rutiliventris Sharp, 1874a: 47 (Philonthus) ムネスジコガシラハネカクシ HOK, HON, Sad, Oki, SHI, KYU, Tsm,
Yak; South Korea, China, Chs, Russia (Sakhalin, Siberia)= tokyanus R. Dvořák, 1957a: 9 (Philonthus)
Genus Phucobius Sharp, 1874 アカバウミベハネカクシ属（和名新称）
densipennis Bernhauer, 1931: 131 リュウキュウアカバウミベハネカクシ（和名改称）Amm, Oke, Okn, Kum, Trm,
Ish, Iri, Yon
simulator Sharp, 1874a: 35 ツヤケシアカバウミベハネカクシ（和名改称）HOK, HON, Tob, Aws, Sad, Iz-Hch, Awj,
Oki, SHI, Shd, KYU, Got, Tan, Yak; South Korea, China, Chs, Russia
Genus Rabigus Mulsant & Rey, 1876 チャバネコガシラハネカクシ属
inconstans Sharp, 1889: 40 (Philonthus) ホソチャバネコガシラハネカクシ HOK, HON, SHI; South Korea, North

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Yasutoshi Shibata, Munetoshi Maruyama, Hideto Hoshina, Toshio Kishimoto, Shun-Ichiro Naomi, Shûhei Nomura, Volker Puthz, Takashi Shimada, Yasuaki Watanabe, Shûhei Yamamoto

Korea, China, Mongolia, Russia
Genus Remus Holme, 1837 ムネスジウミベハネカクシ属
corallicola Fairmaire, 1849: 289 ムネスジウミベハネカクシ Iri; Taiwan, Hong Kong, SE Asia, Tahiti, New Caledonia,
Australia, Ceychelles, Mauritius, Madagascar
Subtribe Quediina Kraatz, 1857
Genus Acylophorus Nordmann, 1837 ヒゲオレハネカクシ属
honshuensis R. Dvořák, 1957: 10 クロヒゲオレハネカクシ HOK, HON
ruficollis Motschulsky, 1858b: 657 ムネアカヒゲオレハネカクシ Amm, Okn, Ish, Iri; Nepal, Oriental Region
Genus Indoquedius Blackwelder, 1952 オオメツヤムネハネカクシ属（和名新称）
juno Sharp, 1874a: 24 (Quedius) ヤマトオオメツヤムネハネカクシ（和名改称）HOK, HON, Awj, Oki, SHI, KYU,
Tsm
praeditus Sharp, 1889: 29 (Quedius) アカアシオオメツヤムネハネカクシ（和名改称）HOK, HON, SHI; South Korea,
China
Genus Pseudorientis Y.Watanabe, 1970 ヒメツヤムネハネカクシ属（和名新称）
shinobuae Y. Watanabe, 1970: 72 ヒメツヤムネハネカクシ（和名新称）HON
Genus Quedius Stephens, 1829 ツヤムネハネカクシ属
Subgenus Distichalius Casey, 1915
annectens Sharp, 1889: 32 クロスジツヤムネハネカクシ HON, SHI
hideyoi Hayashi, 2012: 79 アマミクロスジツヤムネハネカクシ Amm, Tokjaponicus Sharp, 1874a: 26 アカバツヤムネハネカクシ HOK, Rsr, HON, Aws, Sad, SHI, Shd, KYU, Tsm, SKs;South Korea, Russia
kamchaticus Smetana, 1976: 26 カムチャツカツヤムネハネカクシ（和名新称）HOK; Kamchatka
lewisius Sharp, 1874a: 27 ルイスツヤムネハネカクシ HON, Sad, SHI, KYU
pretiosus Sharp, 1874a: 26 ニジツヤムネハネカクシ KYU; China
Subgenus Microsaurus Dejean, 1833
abnormalis Sharp, 1889: 30 ミヤマツヤムネハネカクシ HON
adustus Sharp, 1889: 32 チャイロツヤムネハネカクシ HOK, HON, Oki, SHI, KYU, SKs; China
amamiensis Y. Watanabe, 1990d: 69 (Velleius) アマミクシヒゲツヤムネハネカクシ（和名改称）Amm
aurorus Herman, 2001: 49 [RN] エゾツヤムネハネカクシ HOK, HON, KYU= laticollis Sharp, 1889: 31 [HN]babai Uéno & Y. Watanabe, 1966: 331 ミョウコウツヤムネハネカクシ HON
cephalotes Uéno & Y. Watanabe, 1966: 326 ラカンツヤムネハネカクシ SHIdaibosatsunis Y. Watanabe, 1986a: 19 カントウツヤムネハネカクシ HON
dilatatus Fabricius, 1787: 220 (Staphylinus) クシヒゲツヤムネハネカクシ（和名改称）HOK, HON, Sad, Aws, Oki,2013 The Kyushu

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Catalogue of Japanese Staphylinidae (Insecta: Coleoptera)

SHI; South Korea, China, Turkey, Europe, Russia
elongatus Naomi, 1986a: 244 (Velleius) ナガクシヒゲツヤムネハネカクシ（和名改称）SHI, HON; South Korea
fodinarum Uéno & Y. Watanabe, 1966: 323 キウラツヤムネハネカクシ KYU
hakusanus Y. Watanabe, 1987c: 326 ハクサンツヤムネハネカクシ HON
higonis Uéno & Y. Watanabe, 1970: 14 ヒゴツヤムネハネカクシ KYUhirticornis Sharp, 1889: 31 ツノヒゲツヤムネハネカクシ HOK, HON
horii Y. Watanabe, 2003c: 26 ラッコツヤムネハネカクシ HOK
iriei Uéno & Y. Watanabe, 1970: 10 クマツヤムネハネカクシ KYUkaminakaensis Y. Watanabe & Yoshida, 2008: 135 カミナカツヤムネハネカクシ SHI
kawasawai Uéno & Y. Watanabe, 1970: 15トサヤマツヤムネハネカクシ SHIkiuchii Y. Watanabe & Yoshida, 1970: 2 リュウノイワヤツヤムネハネカクシ SHIkojii Y. Watanabe & Yoshida, 2008: 138 コウジツヤムネハネカクシ SHIkuniakii Y. Watanabe, 1990a: 26 ヤツガタケヒラタツヤムネハネカクシ HON
moritai Y. Watanabe, 1990a: 30 ヤクシヒラタツヤムネハネカクシ HON
ohkawai Y. Watanabe, 2008d: 28 ナエバツヤムネハネカクシ HON
onodai Y. Watanabe, 1986a: 22 ヒロガワラツヤムネハネカクシ HON
ontakesanus Y. Watanabe, 2000: 46 オンタケツヤムネハネカクシ HON
parviceps Sharp, 1874a: 25 コガシラツヤムネハネカクシ HON, Oki, SHI, KYU
pectinatus Sharp, 1874a: 24 (Velleius) クシヒゲツヤムネハネカクシ（和名改称）HON, SHI, KYU, Tsm; Taiwan,
South Korea, China
senjonis Y. Watanabe, 1987c: 330 センジョウツヤムネハネカクシ HON
setosus Sharp, 1889: 29 (Velleius) コクシヒゲツヤムネハネカクシ（和名改称）HOK, HON, SHI, KYU, Yak; South
Korea
shimomurai Y. Watanabe, 1990a: 31 クモトリヒラタツヤムネハネカクシ HON
simulans Sharp, 1874a: 25 ナミツヤムネハネカクシ HOK, Reb, Rsr, HON, Sad, Iz-My, Iz-Mk, Shd, KYU: South Korea,
Chinasugai Uéno & Y. Watanabe, 1966: 333 フジツヤムネハネカクシ HON
suzukii Y. Watanabe, 1987c: 328 スズキツヤムネハネカクシ HON
toyoshimai Y. Watanabe, 2009d: 16 オオキソコマツヤムネハネカクシ HON
troglodytes Uéno & Y. Watanabe, 1966: 327 カンナツヤムネハネカクシ HON
tsurugiensis Y. Watanabe & Yoshida, 1970: 4 ツルギツヤムネハネカクシ SHIuchikawai Y. Watanabe, 1981: 19 カザアナツヤムネハネカクシ HON
uenoi Y. Watanabe, 1979: 12 ウエノツヤムネハネカクシ HON
umbratus Uéno & Y. Watanabe, 1966: 335 クラサワツヤムネハネカクシ HON
watanabei Solodovnikov, 2012: 25 [RN] ヤマトクシヒゲツヤムネハネカクシ（和名改称）HON= japonicus Y. Watanabe, 1990d: 67 (Velleius) [HN]
yasuhikoi Sawada, 1964a: 37 キソコマツヤムネハネカクシ HON
Subgenus Quedionuchus Sharp, 1884
armipes Sharp, 1889: 34 (Quedionuchus)トゲアシツヤムネハネカクシ HON, KYU; Chs, China
planatus Sharp, 1889: 34 (Quedionuchus) チャイブシツヤムネハネカクシ HOK, HON

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Yasutoshi Shibata, Munetoshi Maruyama, Hideto Hoshina, Toshio Kishimoto, Shun-Ichiro Naomi, Shûhei Nomura, Volker Puthz, Takashi Shimada, Yasuaki Watanabe, Shûhei Yamamoto

samuraicus Bernhauer & Schubert, 1916: 433 [RN] イブシツヤムネハネカクシ HON, SHI; South Korea, Russia= concolor Sharp, 1889: 34 (Quedionuchus) [HN]
Subgenus Quedius Stephens, 1829
viduus Sawada, 1965: 17 ジョウネンツヤムネハネカクシ HON
Subgenus Raphirus Stephens, 1829
boops Gravenhorst, 1802: 78 ヒメオオメツヤムネハネカクシ（和名改称）HON (?); Russia (Siberia), Turkistan,
Europelateralis Gravenhorst, 1802: 35 (Staphylinus) キベリツヤムネハネカクシ Japan (?); Palearctic Region
marginipennis Bernhauer, 1939a: 103 カミコウチツヤムネハネカクシ HON
maruyamai Smetana, 2002: 8 リシリツヤムネハネカクシ（和名新称）HOK, Rsr
multipunctatus Sharp, 1889: 33 ミドリオオメツヤムネハネカクシ（和名改称）HON, SHI
sugayai Smetana, 2002: 10 ダイセツツヤムネハネカクシ（和名新称）HOK
Incertae sedis (subgenus?) 亜属不明
imbecillus Sharp, 1889: 31 ハコネツヤムネハネカクシ HON, SHI
tokiensis Bernhauer, 1913a: 132トウキョウツヤムネハネカクシ（和名新称）HON
Genus Quwatanabius Smetana, 2002 マルズツヤムネハネカクシ属（和名新称）
flavicornis Sharp, 1889: 30 (Quedius) ツマグロマルズツヤムネハネカクシ（和名改称）HON, SHI, KYU; Taiwan
Subtribe Staphylinina Latreille, 1802
Genus Agelosus Sharp, 1889 ハバビロオオハネカクシ属（和名新称）
carinatus carinatus Sharp, 1874a: 32 (Goerius) アカバハバビロオオハネカクシ（和名改称）HOK, HON, Sad, Aws;
South Korea, Chinac. brevipennis Naomi, 1983: 585 SHI, KYU
unicolor unicolor Naomi, 1983: 585 クロバネハバビロオオハネカクシ（和名改称）SHI, KYU
u. masaoi Hayashi, 1991: 183 SHIGenus Aulacocypus Müller, 1925 ツツガタハネカクシ属（和名新称）
gloriosus Sharp, 1874a: 34 (Ocypus) キンバネツツガタハネカクシ（和名改称）HON; South Korea, China
parvulus Sharp, 1874a: 33 (Ocypus) チビドウガネツツガタハネカクシ（和名改称）HOK, HON, SHI; South Korea,
China, Russia (Far East)= jureceki Rambousek, 1923: 54 (Staphylinus)
Genus Creophilus Leach, 1819 オオハネカクシ属
maxillosus Linnaeus, 1758: 421 (Staphylinus) オオハネカクシ HOK, HON, Iz-Hch, SHI, KYU, Yak, Amm, Okn,
Ihy, Ish, Iri; Russia (Siberia), Taiwan, South Korea, North Korea, China, India, Hawaii, Europe, N Africa, USA= imbecillus Sharp, 1874a: 28= subfasciatus Sharp, 1874a: 28

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Catalogue of Japanese Staphylinidae (Insecta: Coleoptera)

Genus Dinothenarus Thomson, 1858 ビロウドハネカクシ属（和名新称）
Subgenus Parabemus Reitter, 1909
ganglbauerianus Bernhauer, 1939a: 101 (Staphylinus) ビロウドハネカクシ（和名新称）HON
Genus Eucibdelus Kraatz, 1859 ハイイロハネカクシ属
Subgenus Eucibdelus Kraatz, 1859ishigakiensis Hayashi, 1997: 30 イシガキハイイロハネカクシ（和名新称）Ish
Subgenus Pareucibdelus Hayashi, 1997
japonicus Sharp, 1874a: 29 ハイイロハネカクシ HON, Aws, SHI, KYU, SKs; South Korea= aokii R. Dvořák, 1956a: 30= chujoi R. Dvořák, 1956a: 32= verae R. Dvořák, 1956a: 33
Genus Hadropinus Sharp, 1889 ハマベオオハネカクシ属
fossor Sharp, 1889: 116 ハマベオオハネカクシ HOK, Rsr, HON; South Korea, China, Chs, Russia (Sakhalin)
Genus Liusus Sharp, 1889 カタモンハネカクシ属
hilleri Weise, 1877a: 93 (Hadrotes) カタモンハネカクシ HOK, Rsr, HON, Aws, Oki, SHI, KYU, Tsm; South Korea,
Russiahumeralis Matsumura, 1911: 113 (Staphylinus) キタカタモンハネカクシ HOK; Chs, Russia (Sakhalin), South Korea,
China
Genus Miobdelus Sharp, 1889 ホソサビイロモンキハネカクシ属
brevipennis Sharp, 1889: 112 ホソサビイロモンキハネカクシ HON
insignis Müller, 1926: 41 (Staphylinus) クロバネホソサビイロモンキハネカクシ（和名新称）Japan; China= fokiensis Bernhauer, 1933: 32 (Staphylinus)
Genus Ocypus Leach, 1819 マルズオオハネカクシ属（和名新称）
Subgenus Ocypus Leach, 1819
weisei Harold, 1877: 344 キンボシマルズオオハネカクシ（和名改称）HOK, HON, SHI, KYU, Tsm; South Korea,
North Korea, China
Subgenus Pseudocypus Mulsant & Rey, 1876
lewisius Sharp, 1874a: 33 クロサビイロマルズオオハネカクシ（和名改称）HON, Oki, SHI; South Korea, China= kobensis Cameron, 1930b: 207
nigroaeneus Sharp, 1889: 109 クロマルズオオハネカクシ（和名改称）HOK, HON, SHI; South Korea, North Korea,
China, Mongolia, Russia= rambouseki Müller, 1925: 46 (Staphylinus)
shiretokensis Hayashi, 2008a: 199 シレトコクロマルズオオハネカクシ（和名改称）HOK
Incertae sedis (subgenus?) 亜属不明
japonicus Sawada, 1965: 16 ジョウネンマルズオオハネカクシ（和名新称）HON

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Yasutoshi Shibata, Munetoshi Maruyama, Hideto Hoshina, Toshio Kishimoto, Shun-Ichiro Naomi, Shûhei Nomura, Volker Puthz, Takashi Shimada, Yasuaki Watanabe, Shûhei Yamamoto

Genus Ontholestes Ganglbauer, 1895 サビハネカクシ属
gracilis Sharp, 1874a: 28 (Leistrophus) サビハネカクシ HOK, HON, Oki, SHI, KYU, Tsm; South Korea, North Korea,
Chinaoculatus Sharp, 1874a: 29 (Leistrophus) ヒゲブトサビハネカクシ（和名新称）KYU; China
Genus Philetaerius Sharp, 1889 オオズハイイロハネカクシ属
elegans Sharp, 1889: 119 オオズハイイロハネカクシ HOK, HON, SHI, KYU
Genus Phytolinus Sharp, 1889 アシビロハイイロハネカクシ属（和名新称）
lewisii Sharp, 1889: 121 キオビアシビロハイイロハネカクシ（和名改称）HON, SHI= tsuguriensis R. Dvořák, 1956a: 30 (Eucibdelus)
monticola Naomi, 1984a: 386 ヒトオビアシビロハイイロハネカクシ（和名改称）HON
variegatus Sawada & Nakane, 1954: 8 シラフアシビロハイイロハネカクシ（和名改称）HON, SHI, KYU
Genus Platydracus Thomson, 1858トガリオオズハネカクシ属（和名新称）
Subgenus Platydracus Thomson, 1858
amamiensis T. Ito, 1982: 61 アマミ
トガリオオズハネカクシ（和名新称）Amm
brachycerus Smetana & Davies, 2000: 39 [RN] ヒメクロトガリオオズハネカクシ（和名改称）HOK, HON, SHI,KYU, SKs; South Korea= brevicornis Weise, 1877b: 367 [HN] (Ocypus)
brevicornis Motschulsky, 1861: 11 (Trichoderma) アカバトガリオオズハネカクシ（和名改称）HOK, HON, Aws,
Sad, Oki, SHI, KYU, Got, Tan; South Korea, North Korea, China, Russia= paganus Sharp, 1874a: 30 (Staphylinus)
chalcescens Sharp, 1889: 117 (Staphylinus) アオグロトガリオオズハネカクシ（和名新称）HON
circumcinctus Bernhauer, 1914d: 102 (Staphylinus)ドウガネトガリオオズハネカクシ（和名改称）HON, SHI; Taiwan,
South Korea, North Korea, India
consularis Bernhauer, 1915a: 72 (Staphylinus) ハリマトガリオオズハネカクシ（和名新称）HON
dauricus Mannerheim, 1830: 22 (Staphylinus) アオドウガネトガリオオズハネカクシ（ 和 名 新 称 ）Japan; China,
Mongolia, Russia (Siberia)
inornatus Sharp, 1874a: 31 (Staphylinus) クロガネトガリオオズハネカクシ（和名改称）HOK, HON, Aws, SHI,KYU, Tsm; South Korea, China
kiushiuensis Bernhauer, 1939b: 155 (Staphylinus) キュウシュウトガリオオズハネカクシ（和名新称）KYU
plebejus Bernhauer, 1915a: 73 (Staphylinus) ツシマトガリオオズハネカクシ（和名新称）Tsm; South Korea
sharpi Fauvel, 1901a: 5 [RN] カラカネトガリオオズハネカクシ（和名改称）HOK, HON, SHI, KYU; China, Russia(Sakhalin)= subaeneus Sharp, 1874a: 31 [HN] (Staphylinus)
vicarius Sharp, 1889: 117 (Staphylinus) ヒメアカバトガリオオズハネカクシ（和名改称）HON; South Korea
Genus Protocypus Müller, 1923 サビイロモンキハネカクシ属
amagiensis Naomi & Tahira, 2009: 93 アマギサビイロモンキハネカクシ HON

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Catalogue of Japanese Staphylinidae (Insecta: Coleoptera)dorsalis Sharp, 1889: 109 (Ocypus) サビイロモンキハネカクシ HON
fujiensis Naomi, 1992b: 6 (Ocypus) フジサビイロモンキハネカクシ（和名新称）HON
hagai Naomi, 1992c: 139 (Ocypus) ダイセンサビイロモンキハネカクシ（和名新称）HON
hakusanus Naomi, 1992a: 57 (Ocypus) ハクサンサビイロモンキハネカクシ（和名新称）HON
hidanus Y. Watanabe, 1987b: 43 (Ocypus) ヒダサビイロモンキハネカクシ HON
iyonus Naomi, 1992c: 137 (Ocypus) イヨサビイロモンキハネカクシ（和名新称）SHI
izayoi Naomi, 1992b: 5 (Ocypus) ヤツサビイロモンキハネカクシ（和名新称）HON
kendappa Naomi, 1992c: 144 (Ocypus) イシヅチサビイロモンキハネカクシ（和名新称）HON, SHI
kiimontanus Naomi, 1992b: 11 (Ocypus) キイサビイロモンキハネカクシ（和名新称）HON
kisonus Naomi, 1992c: 142 (Ocypus) キソサビイロモンキハネカクシ（和名新称）HON
nemotoi Naomi, 1992a: 60 (Ocypus) シラネサビイロモンキハネカクシ（和名新称）HON
ohtsukaorum Naomi, 1992a: 63 (Ocypus) ソボサビイロモンキハネカクシ（和名新称）KYU
sabi Naomi, 1992c: 141 (Ocypus) ヒロガワラサビイロモンキハネカクシ（和名新称）HON
scutiger Sharp, 1889: 110 (Ocypus) オオサビイロモンキハネカクシ HON; South Korea= septentrionalis Y. Watanabe, 1984: 139 (Ocypus)
smetanai Naomi, 1992a: 55 (Ocypus) ハヤチネサビイロモンキハネカクシ（和名新称）HON
watarasensis Naomi, 2004d: 154 (Ocypus) ワタラセサビイロモンキハネカクシ HON
yamato Naomi, 1992b: 9 (Ocypus) ヤマトサビイロモンキハネカクシ（和名新称）HON
yoroi Naomi, 1992a: 59 (Ocypus) ミョウコウサビイロモンキハネカクシ（和名新称）HON
yuinus Naomi, 1992a: 60 (Ocypus) ユイサビイロモンキハネカクシ（和名新称）HON
Genus Staphylinus Linne, 1758 ダイミョウハネカクシ属
daimio Sharp, 1889: 118 ダイミョウハネカクシ HOK, HON; China
Subtribe Tanygnathinina Jacobson, 1909
Genus Atanygnathus Jacobson, 1909 ハラホソハネカクシ属
terminalis Erichson, 1839b: 418 (Tanygnathus) チャムネハラホソハネカクシ HOK, HON, SHI, KYU, Amm, MDt;
China, Oriental Region, Europe
juang Smetana, 1995: 136 クロハラホソハネカクシ（和名新称）HON, Amm, Okn, Ish, Iri; Taiwan
Subtribe Xanthopygina Sharp, 1884
Genus Algon Sharp, 1874 ムネビロハネカクシ属
grandicollis Sharp, 1874a: 23 ムネビロハネカクシ HOK, HON, Aws, Sad, Awj, Oki, SHI, Shd, KYU, Tsm, Yak;South Korea, North Korea, China
Genus Rientis Sharp, 1874 アバタツヤムネハネカクシ属
parviceps Sharp, 1874a: 21 アバタツヤムネハネカクシ HON, KYU

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Yasutoshi Shibata, Munetoshi Maruyama, Hideto Hoshina, Toshio Kishimoto, Shun-Ichiro Naomi, Shûhei Nomura, Volker Puthz, Takashi Shimada, Yasuaki Watanabe, Shûhei Yamamoto

Tribe Xantholinini Erichson, 1839
Genus Gauropterus C. G. Thomson, 1860 ムネミゾナガハネカクシ属（和名新称）
fulgidus Fabricius, 1787: 220 (Staphylimus) キバネムネミゾナガハネカクシ（和名改称）HON, KYU; South Korea,
China, Myanmar, Indonesia (Sumatra), Sri Lanka, Europe
Genus Hypnogyra Casey, 1906 ツツガタナガハネカクシ属（和名新称）
tenebrosa Bordoni, 1997: 174 クロバネツツガタナガハネカクシ（和名改称）HOK, HON, Sad, SHI, KYU; Russia
tubula Sharp, 1889: 250 (Xantholinus) アカバネツツガタナガハネカクシ（和名改称）HOK, HON, Oki, SHI, KYU;
South Korea, China
Genus Megalinus Mulsant & Rey, 1877 キバネナガハネカクシ属
japonicus Sharp, 1874a: 52 (Xantholinus) オオキバネナガハネカクシ HOK, HON, SHI, KYU; South Korea, North
Korea, China
mixtus Sharp, 1874a: 53 (Xantholinus) ホソキバネナガハネカクシ（和名新称）HOK, HON, Sad, SHI, KYU, Tsm,
Yak; South Korea, China
suffusus Sharp, 1874a: 52 (Xantholinus) キバネナガハネカクシ HOK, HON, Iz-O, Iz-My, Iz-Mk, SHI, Shd, KYU,
Tsm, SKs, Yak; Taiwan, China
Genus Liotesba Scheerpeltz, 1965 ツヤクビナガハネカクシ属（和名新称）
punctiventris Sharp, 1889: 252 (Xantholinus) アカバツヤクビナガハネカクシ（和名新称）HON, Iz-Mk, SHI, KYU;
Tsm
Genus Medhiama Bordoni, 2002 ホソナガハネカクシ属
paupera Sharp, 1889: 250 (Xantholinus) ウスイロホソナガハネカクシ HOK, HON, SHI; Chs, China, India, Nepal
Genus Gyrohypnus Leach, 1819 アバタナガハネカクシ属（和名新称）
keijiroi Y. Watanabe & Shibata, 1965b: 319 (Xantholinus) コゲチャアバタナガハネカクシ（和名新称）Reb, Rsr
Genus Metolinus Cameron, 1920 アシビロナガハネカクシ属（和名新称）
leucocnemis Kraatz, 1859: 107 (Metoponcus) ヤクアシビロナガハネカクシ（和名新称）Yak; Sri Lanka, India,
Nepal, Myanmar, Vietnam, Indonesia
planulatus Sharp, 1889: 252 (Leptacinus) ヒゴアシビロナガハネカクシ（和名新称）HON, KYU; China
Genus Nudobius C. G. Thomson, 1860 ツヤナガハネカクシ属
lentus Gravenhorst, 1806: 101 (Xantholinus) アカバネツヤナガハネカクシ（和名新称）HOK; Chs, China, Russia(Siberia), Turkey, Europe
pleuralis Sharp, 1874a: 53 (Xantholinus) ツマキツヤナガハネカクシ HOK, HON, Sad, Oki, SHI, KYU, Tsm; South
Korea, China= apicipennis Sharp, 1889: 253

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Catalogue of Japanese Staphylinidae (Insecta: Coleoptera)

sejunctus Y. Watanabe & Shibata, 1965a: 91 ヒゲブトツヤナガハネカクシ HON; Taiwan= formosanus Shibata, 1973: 123
Genus Pachycorynus Motschulsky, 1858 ヒラズホソクビハネカクシ属（和名新称）
dimidiatus Motschulsky, 1858c: 205 ミナミヒラズホソクビハネカクシ（和名新称）Iri, Hat; Taiwan, India, Oriental
Region, New Guinea
Genus Phacophallus Coiffait, 1956 ヨツミゾナガハネカクシ属（和名新称）
flavipennis Kraatz, 1859: 111 (Leptacinus) キバネヨツミゾナガハネカクシ（和名改称）HON, SHI, KYU, Tan, Yak,
Kue, Okn, Iri; Taiwan, China, India, Sri Lanka, Nepal, Myanmar, Malaysia, Cambodia, Laos, Vietnam, Malaysia(Borneo), Philippines, Australia
japonicus Cameron, 1933: 169 (Leptacinus) クロズヨツミゾナガハネカクシ（和名改称）HON, Tsm, Yak; South
Korea, North Korea, China, Thailand, Malaysia, Indonesia (Sumatra, Java)= chinensis Cameron, 1940: 251 (Leptacinus)
pallidipennis Motschulsky, 1858c: 206 (Leptacinus) オオヨツミゾナガハネカクシ（ 和 名 新 称 ）HON; China,
Thailand, Bhutan, Indonesia, Singapore, Sri Lanka, India, Philippines, Europe, USA= tricolor Kraatz, 1859: 110 (Leptacinus)
parumpunctatus Gyllenhal, 1827: 481 (Staphylinus) フトヨツミゾナガハネカクシ（和名新称）SHI (?), Yak (?);
Cyprus, Turkey, Europe
Genus Stenistoderus Jacquelin du Val, 1856 ホソムネハネカクシ属
nothus Erichson, 1839a: 338 (Leptacinus) チャイロホソムネハネカクシ HON, Sad, SHI; Iraq, Europe, Canary, USA
Genus Xantholinus Dejean, 1821 ムネスジナガハネカクシ属
angusticeps Sharp, 1889: 251 ナガズムネスジナガハネカクシ（和名新称）KYU
cunctator Sharp, 1889: 251 ムネスジナガハネカクシ HON, SHI
Genus Xanthophius Motschulsky, 1860 ヒメナガハネカクシ属
angustus Sharp, 1874a: 54 ズグロヒメナガハネカクシ HON, Iz-Hch, Og, Tan; Taiwan, South Korea, China
Genus Zeteotomus Jacquelin du Val, 1856 ズナガホソクビハネカクシ属
maximus Bernhauer, 1907: 384 (Metoponcus) ズナガホソクビハネカクシ HOK, HON, SHI, KYU, Yak; China, Chs,
Russia (Siberia)
dilatipennis Kirshenblat, 1948: 48 (Metpponcus) クナシリズナガホソクビハネカクシ（和名新称）Kun; Russia (Far East)

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Yasutoshi Shibata, Munetoshi Maruyama, Hideto Hoshina, Toshio Kishimoto, Shun-Ichiro Naomi, Shûhei Nomura, Volker Puthz, Takashi Shimada, Yasuaki Watanabe, Shûhei Yamamoto

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Yasutoshi Shibata, Munetoshi Maruyama, Hideto Hoshina, Toshio Kishimoto, Shun-Ichiro Naomi, Shûhei Nomura, Volker Puthz, Takashi Shimada, Yasuaki Watanabe, Shûhei Yamamoto

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Yasutoshi Shibata, Munetoshi Maruyama, Hideto Hoshina, Toshio Kishimoto, Shun-Ichiro Naomi, Shûhei Nomura, Volker Puthz, Takashi Shimada, Yasuaki Watanabe, Shûhei Yamamoto

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Yasutoshi Shibata, Munetoshi Maruyama, Hideto Hoshina, Toshio Kishimoto, Shun-Ichiro Naomi, Shûhei Nomura, Volker Puthz, Takashi Shimada, Yasuaki Watanabe, Shûhei Yamamoto

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Naomi S.-I. 1981: On the genus Anisolinus Sharp (Coleoptera, Staphylinidae) with description of a new species.
Kontyû 49: 109-115.
Naomi S.-I. 1982: Description of a new subgenus Paramichrotus of the genus Amichrotus from Taiwan (Coleoptera,
Staphylinidae). Transactions of the Shikoku Entomological Society 16: 37-39.
Naomi S.-I. 1983: Revision of the subtribe Xanthopygina (Coleoptera, Staphylinidae) of Japan. III. Kontyû 51: 582592.
Naomi S.-I. 1984a: Studies on the genus Phytolinus Sharp (Coleoptera, Staphylinidae) from Japan and Taiwan, withdescriptions of two new species. Kontyû 52: 382-389.
Naomi S.-I. 1984b: Studies on the genus Oligota Mannerheim (Coleoptera, Staphylinidae) from Japan. 1. On thespecies-group of O. yasumatsui Kistner, with description of a new subspecies. Kontyû 52: 516-521.
Naomi S.-I. 1986a: Taxonomic study on the genus Velleius Mannerheim of Japan, with description of a new species(Coleoptera, Staphylinidae). Transactions of the Shikoku Entomological Society 17: 239-246.

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Naomi S.-I. 1986b: A taxonomic study on the subfamily Megalopininae (Coleoptera, Oxyporidae) of Japan, withdescriptions of two new species. Kontyû 54: 344-352.
Naomi S.-I. 1986c: Taxonomic study on the subfamily Osoriinae (Coleoptera, Oxytelidae) from Japan, I. Elytra, Tokyo
14: 33-42.
Naomi S.-I. 1987a: Studies on the subfamily Steninae (Coleoptera, Oxyporidae) from Japan, VII. Subgenus Parastenusof the genus Stenus Latreille (Part 2). Akitu 87: 1-9.
Naomi S.-I. 1987b: Taxonomic study on the subfamily Osoriinae (Coleoptera, Oxytelidae) from Japan, II. Elytra
Tokyo 15: 12-18.
Naomi S.-I. 1987c: Discovery of the subfamily Hypocyptinae (Coleoptera, Staphylinidae) from Japan, with descriptionof a new species. Transactions of the Shikoku Entomological Society 18: 273-275.
Naomi S.-I. 1988a: Studies on the subfamily Steninae (Coleoptera, Oxyporidae) from Japan. II. Subgenus Hemistenus
on the genus Stenus Latreille. Proceedings of the Japanese Society of Systematic Zoology 37: 27-33.
Naomi S.-I. 1988b: Studies on the subfamily Steninae (Coleoptera, Oxyporidae) from Japan. VI. Subgenus Parastenusof the genus Stenus, Part1. Elytra, Tokyo 16: 33-40.
Naomi S.-I. 1988c: Studies on the subfamily Steninae (Coleoptera, Oxyporidae) from Japan. III. New or little known
species of the subgenus Nestus of the genus Stenus Latreile. Proceedings of the Japanese Society of Systematic
Zoology 38: 43-50.
Naomi S.-I. 1988d: Studies on the subfamily Steninae (Coleoptera, Oxyporidae) from Japan. I. New or little known
species of the genus Dianous Leach (Coleoptera, Oxyporidae). Transactions of the Shikoku Entomological Society
19: 47-54.
Naomi S.-I. 1988e: Studies on the subfamily Steninae (Coleoptera, Oxyporidae) from Japan. IV. New or little known
species of the subgenus Stenus s. str. of the genus Stenus Latreille (Coleoptera, Oxyporidae). Transactions of the
Shikoku Entomological Society 19: 55-62.
Naomi S.-I. 1988f: Studies on the subfamily Steninae (Coleoptera, Oxyporidae) from Japan. IX. Subgenus Hypostenusof the genus Stenus Latreille, Part 1. Elytra, Tokyo 16: 73-84.
Naomi S.-I. 1989a: Studies on the subfamily Steninae (Coleoptera, Oxyporidae) from Japan. X. Subgenus Hypostenusof the genus Stenus Latreille (Part 2). Akitu (N. S.) 105: 1-11.
Naomi S.-I. 1989b: Studies on the subfamily Steninae (Coleoptera, Oxyporidae) from Japan. V. Subgenus Tesnusof the genus Stenus Latreille, with a key to the species of the subgenera Tesnus and Nestus. Proceedings of theJapanese Society of Systematic Zoology 39: 37-42.
Naomi S.-I. 1989c: Studies on the subfamily Steninae (Coleoptera, Oxyporidae) from Japan. XII. Subgenus
Hypostenus of the genus Stenus Latreille, Part 4. Elytra , Tokyo 17: 45-52.
Naomi S.-I. 1989d: Studies on the subfamily Steninae (Coleoptera, Oxyporidae) from Japan. XIII. Subgenus
Hypostenus of the genus Stenus Latreille, Part 5. Elytra, Tokyo 17: 159-167.
Naomi S.-I. 1990a: Studies on the subfamily Steninae (Coleoptera, Oxyporidae) from Japan. XI. Subgenus Hypostenusof the genus Stenus Latreille (Part 3). Akitu (N. S.) 113: 1-10.
Naomi S.-I. 1990b: Studies on the subfamily Steninae (Coleoptera, Oxyporidae) from Japan. XIV. Subgenus
Hypostenus of the genus Stenus Latreille, Part 6. Elytra, Tokyo 18: 45-53.
Naomi S.-I. 1990c: Studies on the subfamily Steninae (Coleoptera, Oxyporidae) from Japan. XVII. Descriptions offive new species of the genus Stenus Latreille. Natural History Research 1: 99-107.

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Yasutoshi Shibata, Munetoshi Maruyama, Hideto Hoshina, Toshio Kishimoto, Shun-Ichiro Naomi, Shûhei Nomura, Volker Puthz, Takashi Shimada, Yasuaki Watanabe, Shûhei Yamamoto

Naomi S.-I. 1990d: Studies on the subfamily Steninae (Coleoptera, Oxyporidae) from Japan. XV. Subgenus
Hypostenus of the genus Stenus Latreille, Part 7. Elytra, Tokyo 18: 197-207.
Naomi S.-I. 1992a: Taxonomic studies on the subgenus Protocypus Müller of the genus Ocypus Leach (Coleoptera:
Staphylininae) of Japan, I. Natural History Research 2: 53-64.
Naomi S.-I. 1992b: Taxonomic studies on the subgenus Protocypus Müller of the genus Ocypus Leach (Coleoptera:
Staphylininae) of Japan. II. Akitu 127: 1-12.
Naomi S.-I. 1992c: Taxonomic studies on the subgenus Protocypus Müller of the genus Ocypus Leach (Coleoptera:
Staphylininae) of Japan. III. Japanese Journal of Entomology 60: 137-147.
Naomi S.-I. 1994a: Description of a new species of genus Gymnusa Gravenhorst (Coleoptera: Staphylinidae) from
Hokkaido, Japan. Natural History Research 3: 47-50.
Naomi S.-I. 1994b: Revision of the debile group of the genus Siagonium Kirby et Spence from Japan (Coleoptera,
Staphylinidae, Piestinae). Entomological Review of Japan 49: 145-154.
Naomi S.-I. 1995a: Taxonomic study on the genus Thinobius Kiesenwetter from Japan (Coleoptera, Staphylinidae).
Entomological Review of Japan 50: 17-25.
Naomi S.-I. 1995b: Revision of the subfamily Piestinae (Coleoptera: Staphylinidae) from Japan, I. Natural History
Research 3: 141-151.
Naomi S.-I. 1995c: Descriptions of two new species of the subfamily Paederinae (Coleoptera, Staphylinidae) fromJapan. Japanese Journal of Systematic Entomology 1: 161-168.
Naomi S.-I. 1995d: Description of a new species of the genus Trichophya (Coleoptera, Staphylinidae) from Japan.
In: Watanabe Y., Satô M. & Owada M. (eds.): Beetles and Nature. Special Bulletin of the Japanese Society ofColeopterology (Tokyo) 4: 347-350.
Naomi S.-I. 1995e: Revision of the genus Piestoneus Sharp (Coleoptera: Staphylinidae) from Japan. Japanese Journalof Entomology 63: 763-780.
Naomi S.-I. 1996a: Revision of the tribe Leptochirini Sharp (Coleoptera, Staphylinidae) from Japan. Japanese Journalof Systematic Entomology 2: 1-17.
Naomi S.-I. 1996b: Two new species of the family Staphylinidae (Coleoptera) from Japan. New Entomologist 45: 6973.
Naomi S.-I. 1996c: Revision of the genera Lispinus Erichson and Neolosus Blackwelder (Coleoptera: Staphylinidae)
from Japan. Japanese Journal of Entomology 64: 762-771.
Naomi S.-I. 1997a: Two new Dianous Leach (Coleoptera, Staphylinidae) from Japan, with taxonomic notes on someJapanese Stenus Latreille. New Entomologist 46: 1-7.
Naomi S.-I. 1997b: A new species of Stenus cirrus-group (Coleoptera, Staphylinidae, Steninae) from Japan, withdiscussion on the morphology of spermatheca. Entomological Review of Japan 52: 1-7.
Naomi S.-I. 1997c: Four new species of the subgenus Parastenus of the genus Stenus Latreille (Coleoptera,
Staphylinidae) from Japan. Japanese Journal of Systematic Entomology 3: 15-23.
Naomi S.-I. 1997d: A revision of the genus Nacaeus Blackwelder (Coleoptera, Staphylinidae) from Japan. Japanese
Journal of Entomology 65: 127-142.
Naomi S.-I. 1997e: Four new species and two new subspecies of the subgenus Nestus of the genus Stenus Latreille(Coleoptera, Staphylinidae) from Japan. Natural History Research 4: 135-143.
Naomi S.-I. 1997f: Taxonomic study on Stenus wasabi Hromádka and its allied species (Coleoptera; Staphylinidae;

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Steninae), with discussion of the internal structure of the aedeagus. Japanese Journal of Entomology 65: 600-611.
Naomi S.-I. 1997g: Four new species of the genus Stenus (Coleoptera, Staphylinidae), with redescriptions of two
interesting species from Japan. Japanese Journal of Entomology 65: 745-759.
Naomi S.-I. 1998a: Four new brachypterous species of the subgenus Hypostenus of the genus Stenus (Coleoptera:
Staphylinidae) from Japan. Entomological Science 1: 99-104.
Naomi S.-I. 1998b: Six new brachypterous species of the genus Stenus (Coleoptera: Staphylinidae) from Japan.
Entomological Science 1: 385-392.
Naomi S.-I. 2004a: Taxonomic study of Stenus bicara Naomi and its allied species (Coleoptera: Staphylinidae) fromJapan. Esakia 44: 183-189.
Naomi S.-I. 2004b: Description of a new species of the genus Priochirus Sharp (Coleoptera: Staphylinidae) fromJapan. Entomological Review of Japan 59: 99-104.
Naomi S.-I. 2004c: Taxonomic study on Stenus biwa Hromádka and its allied species (Coleoptera: Staphylinidae)
from Japan. Entomological Review of Japan 59: 105-112.
Naomi S.-I. 2004d: Descriptions of two new species of the family Staphylinidae (Coleoptera) from Japan.
Entomological Review of Japan 59: 153-159.
Naomi S.-I. 2004e: Descriptions of two new species of the genus Stenus Latreille (Coleoptera: Staphylinidae) fromJapan. Natural History Research 8: 13-18.
Naomi S.-I. 2004f: Two new species of the S. cirrus-Group of the genus Stenus Latreille (Coleoptera, Staphylinidae)
from Japan. Natural History Research 8: 19-25.
Naomi S.-I. 2004g: Taxonomic study on Stenus asyura Naomi and its allied species (Coleoptera: Staphylinidae) fromJapan. Natural History Research 8: 27-32.
Naomi S.-I. 2006a: Taxonomic revision of the genus Stenus Latreille, 1797 (Coleoptera, Staphylinidae, Steninae) ofJapan: Species group of S. (Hypostenus) rufescens Sharp. Natural History Research Special Issue 9: 1-81.
Naomi S.-I. 2006b: Taxonomic revision of the genus Stenus Latreille, 1797 (Coleoptera, Staphylinidae, Steninae) ofJapan: Species group of S. indubius Sharp. Japanese Journal of Systematic Entomology 12: 39-120.
Naomi S.-I. 2006c: Description of a new species of the genus Siagonium Kirby et Spence from Japan (Coleoptera,
Staphylinidae, Piestinae). Natural History Research 9: 41-44.
Naomi S.-I. 2007: Descriptions of two new species of the subgenus Hypostenus of the genus Stenus Latreille(Coleoptera, Staphylinidae) from Japan. Natural History Research 9: 71-75.
Naomi S.-I. 2010a: Descriptions of two new species of the genus Stenus Latreille (Coleoptera: Staphylinidae: Steninae)
from Chubu District, Central Japan. Natural History Research 11: 35-40.
Naomi S.-I. 2010b: A new species of the genus Dianous Leach (Coleoptera: Staphylinidae: Steninae) from the Nansei
Islands, Japan. Natural History Research 11: 41-45.
Naomi S.-I. 2011a: A new species of the genus Stenus Latreille, 1797 (Coleoptera, Staphylinidae, Steninae) from Mie
Pref., central Japan. Japanese Journal of Systematic Entomology 17: 47-50.
Naomi S.-I. 2011b: Discovery of the genus Eupiestus Kraatz, 1859 (Coleoptera, Staphylinidae, Piestinae) from Japan,
with description of a new species from Iriomote Is., Nansei Islands. Japanese Journal of Systematic Entomology
17: 59-64.
Naomi S.-I. 2011c: Two new synonyms of Staphylinidae (Coleoptera) from Japan. Japanese Journal of Systematic
Entomology 17: 65-66.

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Yasutoshi Shibata, Munetoshi Maruyama, Hideto Hoshina, Toshio Kishimoto, Shun-Ichiro Naomi, Shûhei Nomura, Volker Puthz, Takashi Shimada, Yasuaki Watanabe, Shûhei Yamamoto

Naomi S.-I. 2012a: A new species of the genus Lispinus Erichson (Staphylinidae, Osoriinae, Thoracophorini) from
Nansei Islands, Japan. Japanese Journal of Systematic Entomology 18: 149-153.
Naomi S.-I. 2012b: Taxonomic revision of the genus Stenus Latreille, 1797 (Coleoptera, Staphylinidae, Steninae) ofJapan: Species group of S. cephalotes Sharp (Part 1). Japanese Journal of Systematic Entomology 18: 269-318.
Naomi S.-I. & Irmler U. 2012: First record of the genus Allotrochus Fagel, 1955 (Coleoptera: Staphylinidae:
Osoriinae) from the Oriental region, with description of the new species A. takahashii from Ishigaki Is., Nansei
Islands, Japan. Biogeography 14: 105-110.
Naomi S.-I. & Iwata R. 1996: Kistnerium japonicum Naomi et Iwata n. gen. et n. sp., the first representative of theTribe Mesoporini (Coleoptera, Staphylinidae, Aleocharinae) from Japan, associated with termite (Isoptera:
Rhinotermitidae). Sociobiology 28: 73-81.
Naomi S.-I. & Maruyama M. 1997a: A Revision of the genus Sepedophilus Gistel (Coleoptera; Staphylinidae;
Tachyporinae) from Japan: Species group of S. armatus Sharp. Japanese Journal of Systematic Entomology 3:
239-257.
Naomi S.-I. & Maruyama M. 1997b: Revision of the genus Sepedophilus Gistel (Coleoptera; Staphylinidae;
Tachyporinae) from Japan: Species group of S. glabratus and S. exiguus. Entomological Review of Japan 52: 6171.
Naomi S.-I. & Maruyama M. 1997c: Redescriptions of two interesting species of the genus Sepedophilus Gistel(Coleoptera: Staphylinidae) from Japan. Entomological Review of Japan 52: 73-78.
Naomi S.-I. & Maruyama M. 1998: A Revision of the genus Sepedophilus Gistel (Coleoptera, Staphylinidae,
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Yasutoshi Shibata, Munetoshi Maruyama, Hideto Hoshina, Toshio Kishimoto, Shun-Ichiro Naomi, Shûhei Nomura, Volker Puthz, Takashi Shimada, Yasuaki Watanabe, Shûhei Yamamoto

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Yasutoshi Shibata, Munetoshi Maruyama, Hideto Hoshina, Toshio Kishimoto, Shun-Ichiro Naomi, Shûhei Nomura, Volker Puthz, Takashi Shimada, Yasuaki Watanabe, Shûhei Yamamoto

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Yasutoshi Shibata, Munetoshi Maruyama, Hideto Hoshina, Toshio Kishimoto, Shun-Ichiro Naomi, Shûhei Nomura, Volker Puthz, Takashi Shimada, Yasuaki Watanabe, Shûhei Yamamoto

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Watanabe Y. 2001c: A new species of the group of Lathrobium brachypterum (Coleoptera, Staphylinidae) from2013 The Kyushu

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Yasutoshi Shibata, Munetoshi Maruyama, Hideto Hoshina, Toshio Kishimoto, Shun-Ichiro Naomi, Shûhei Nomura, Volker Puthz, Takashi Shimada, Yasuaki Watanabe, Shûhei Yamamoto

southern Aizu in northeastern Honshu, Japan. Elytra, Tokyo 29: 358-363.
Watanabe Y. 2001d: Four new species of apterous Lathrobium (Coleoptera, Staphylinidae) from central Honshu,
Japan. Elytra, Tokyo 29: 465-475.
Watanabe Y. 2002a: A new species of the group of Lathrobium brachypterum (Coleoptera, Staphylinidae) from the
Bôsô Peninsula in central Honshu, Japan. Elytra, Tokyo 30: 314−319.
Watanabe Y. 2002b: Five new staphylinids of the group of Lathrobium nomurai (Coleoptera, Staphylinidae) fromwestern Honshu, Japan. Elytra, Tokyo 30: 439−449.
Watanabe Y. 2002c: A new species of the group of Lathrobium pollens (Coleoptera, Staphylinidae) from southern
Shikoku, Japan. Journal of the Speleological Society of Japan 27: 12−16.
Watanabe Y. 2003a: Lathrobium satoi (Coleoptera, Staphylinidae), a new staphylinid beetle from Tôkai District incentral Honshu, Japan. Special Bulletin of the Japanese Society of Coleopterolology 6: 151−155.
Watanabe Y. 2003b: Notes on the genus Eusphalerum (Coleoptera, Staphylinidae) from Aomori Prefecture, Japan.
Elytra, Tokyo 31: 395−402.
Watanabe Y. 2003c: A new species of the group of Quedius yasuhikoi (Coleoptera, Staphylinidae) from Hokkaido,
Japan. Journal of the Speleological Society of Japan 28: 26−30.
Watanabe Y. 2004a: Two new species of the genus Lesteva (Coleoptera, Staphylinidae) from the Island of Dôgo of the
Oki Islands, West Japan. Elytra 32: 71−77.
Watanabe Y. 2004b: A remarkable new species of the genus Micropeplus (Coleoptera, Staphylinidae) from Hokkaido,
northeast Japan. Elytra, Tokyo 32: 79−84.
Watanabe Y. 2004c: Lathrobium japonicum and its new relatives (Coleoptera, Staphylinidae) from the Kuril Islands.
Biodiversity and Biogeography of the Kuril Islands and Sakhalin (2004), 1: 37−44.
Watanabe Y. 2005a: Apterous Lathrobium (Coleoptera, Staphylinidae) from the Kii Peninsula in Japan. 1. Group of
Lathrobium singon. Elytra, Tokyo 33: 313−325.
Watanabe Y. 2005b: A new subterranean Lathrobium (Coleoptera, Staphylinidae) from northwestern Shikoku, Japan.
Journal of the Speleological Society of Japan 39: 29−33.
Watanabe Y. 2005c: Apterous Lathrobium (Coleoptera, Staphylinidae) from the Kii Peninsula in Japan. 2. Group of
Lathrobium pollens. Elytra, Tokyo 33: 589−601.
Watanabe Y. 2006a: Apterous Lathrobium (Coleoptera, Staphylinidae) from the Kii Peninsula in Japan. 3. Group of
Lathrobium brachypterum. Elytra, Tokyo 34: 47−62.
Watanabe Y. 2006b: On the systematic status of Lathrobium (s. str.) japonicum kunashiriense Y. Watanabe(Coleoptera, Staphylinidae). Biodiversity and Biogeography of the Kuril Islands and Sakhalin (2006), 2: 160.
Watanabe Y. 2007a: A new species of the genus Coryphium (Coleoptera, Staphylinidae) from Hokkaido, Japan.
Elytra, Tokyo 35: 299−304.
Watanabe Y. 2007b: Two new species of the genus Acrolocha (Coleoptera, Staphylinidae) from Japan. Elytra, Tokyo
35: 433−440.
Watanabe Y. 2008a: A new apterous Ochthephilum (Coleoptera, Staphylinidae) from the Island of Okinawa–hontô ofthe Ryukyus, Japan. Elytra, Tokyo 36: 199−203.
Watanabe Y. 2008b: Two new species of Lathrobium (Coleoptera, Staphylinidae) from Mt. Mayasan of Hyôgo
Prefecture in western Honshu, Japan. Special Publication of the Japan Coleopterological Society 2: 183−190.
Watanabe Y. 2008c: New brachypterous Platydomene (Coleoptera, Staphylinidae) from mountain areas of central

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Honshu, Japan. Elytra, Tokyo 36: 331−341.
Watanabe Y. 2008d: A new subterranean species of the genus Quedius (Coleoptera, Staphylinidae) from central
Honshu, Japan. Journal of the Speleological Society of Japan 33: 28−32.
Watanabe Y. 2009a: A new species of the genus Phloeostiba (Coleoptera, Staphylinidae) from Hokkaido, Japan.
Elytra, Tokyo 37: 71−75.
Watanabe Y. 2009b: More new brachypterous species of the group of Platydomene nobilis (Coleoptera, Staphylinidae)
from northeastern Honshu, Japan. Elytra, Tokyo 37: 245−253.
Watanabe Y. 2009c: A new species of Philydrodes (Coleoptera, Staphylinidae) from eastern Shikoku, Japan. Japanese
Journal of Systematic Entomology 15: 343−346.
Watanabe Y. 2009d: A new species of the group of Quedius abnormalis (Coleoptera, Staphylinidae) from Mt. Kiso−
komagatake in central Honshu, Japan. Journal of the Speleological Society of Japan 34: 15−21.
Watanabe Y. 2010a: Lathrobium densum and its two new relatives (Coleoptera, Staphylinidae) from western Honshu,
Japan. Elytra, Tokyo 38: 73−82.
Watanabe Y. 2010b: More new species of the group of Lathrobium nomurai (Coleoptera, Staphylinidae) from eastern
Shikoku, Japan. Japanese Journal of Systematic Entomology 16: 63−69.
Watanabe Y. 2010c: Lathrobium brachypterum and its new relative (Coleoptera, Staphylinidae) from central Honshu,
Japan. Elytra, Tokyo 38: 257−265.
Watanabe Y. 2011a: Two new species of the brachypterous Lathrobium (Coleoptera, Staphylinidae) from the Northern
Japanese Alps in central Honshu, Japan. Special Publication of the Japanese Society of Scarabaeoidology 1: 103108.
Watanabe Y. 2011b: Three new apterous Lathrobium (Coleoptera, Staphylinidae) from Hiroshima Prefecture inwestern Honshu, Japan. Elytra, Tokyo New Series 1: 47-55.
Watanabe Y. 2011c: Two new species of apterous Lathrobium (Coleoptera, Staphylinidae) from Okayama Prefecture
in western Honshu, Japan. Japanese Journal of Systematic Entomology 17: 177-183.
Watanabe Y. 2012a: Two new species of the Subgenus Minyphilydrodes (Coleoptera, Staphylinidae) from eastern
Shikoku, Japan. Japanese Journal of Systematic Entomology 18: 17-22.
Watanabe Y. 2012b: Two new species of Eusphalerum (Coleoptera, Staphylinidae) from the Southern Japanese Alpsin central Honshu, Japan. Elytra, Tokyo New Series 2: 85-90.
Watanabe Y. 2012c: Description of a new paederine genus Nipponolathrobium (Coleoptera, Staphylinidae), with two
new species from Japan. Japanese Journal of Systematic Entomology 18: 335-345.
Watanabe Y. & Baba K. 1973: Staphylinid beetles found in old gold mines of the island of Sado, central Japan.
Annotationes Zoologicae Japonenses 46: 259-265.
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in central Honshu, Japan. Memoirs of the Research and Education Center for Regional Environment Fukui
University, “Nature and Environment of The Sea of Japan Districts” 10: 7−14.
Watanabe Y. & Satô M. 1981: A new Brathinus (Coleoptera, Staphylinidae) from Shikoku, Japan. Kontyû 49: 615619.
Watanabe Y. & Shibata Y. 1960: Description of a new species of genus Syntomium in Japan (Col. Staphylinidae).
Journal of Agricultural Science (Tokyo) 6: 103-105.
Watanabe Y. & Shibata Y. 1961a: On the genus Ancyrophorus in Japan with descriptions of four new species (Col.

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Yasutoshi Shibata, Munetoshi Maruyama, Hideto Hoshina, Toshio Kishimoto, Shun-Ichiro Naomi, Shûhei Nomura, Volker Puthz, Takashi Shimada, Yasuaki Watanabe, Shûhei Yamamoto

Staphylinidae). Journal of Agricultural Science (Tokyo) 7: 6-9.
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Staphylinidae). Journal of Agricultural Science (Tokyo, Commemoration Issue of 70th Anniversary) 7: 35-40.
Watanabe Y. & Shibata Y. 1961c: A revision of the genus Elonium Leach in Japan (Col. Staph.). Journal of
Agricultural Science (Tokyo) 7: 43-45.
Watanabe Y. & Shibata Y. 1962: Description of a new species of the genus Trichophya Mannerheim from Japan (Col.
Staphylinidae). Journal of Agricultural Science (Tokyo) 7: 95-96.
Watanabe Y. & Shibata Y. 1964: On the genus Micropeplus Latr. of Japan with descriptions of a new and an
unrecorded species (Col. Staphylinidae). Journal of Agricultural Science (Tokyo) 10: 67-70.
Watanabe Y. & Shibata Y. 1965a: A revision of the genus Nudobius Thomson of Japan (Col. : Staphylinidae). Journalof Agricultural Science (Tokyo) 10: 91-94.
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Watanabe Y. & Shibata Y. 1972: The Staphylinid-fauna of Yaku-shima Island, Japan, with descriptions of a new
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from eastern Shikoku, Japan. Japanese Journal of Systematic Entomology 13: 59-62.
Watanabe Y. & Yoshida M. 2008: Two new species of the group of Quedius abnormalis (Coleoptera, Staphylinidae)
from eastern Shikoku, Japan. Japanese Journal of Systematic Entomology 14: 135−140.
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Catalogue of Japanese Staphylinidae (Insecta: Coleoptera)

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167-169.

別表1．アルファベット順地名略号一覧．
Appendix 1. Alphabetical list of abbreviations of local names.
Amk

天草 Amakusa — Off Kyûshû

Amm

奄美大島 Amami-ôshima — 南西諸島 Nansei-shotô (Ryukyu)

Awj

淡路島 Awaji-shima — 本州沖 Off HonshûAws

粟島 Awa-shima — 本州沖 Off Honshû

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Yasutoshi Shibata, Munetoshi Maruyama, Hideto Hoshina, Toshio Kishimoto, Shun-Ichiro Naomi, Shûhei Nomura, Volker Puthz, Takashi Shimada, Yasuaki Watanabe, Shûhei Yamamoto

Chs

千島 Chishima (=Kuril) — 北海道沖 Off Hokkaidô

Dnj

男女群島 Danjo-guntô — 九州沖 Off Kyûshû

Etr

択捉島 Etorofu-tô (=Iturup) — 北海道沖 Off HokkaidôFuk

福江島 Fukue-jima — 九州沖 Off Kyûshû

Got

五島列島 Gotô-rettô — 九州沖 Off Kyûshû

Hat

波照間島 Hateruma-jima — 南西諸島 Nansei-shotô (Ryukyu)

Hej

戸島 He-jima — 九州沖 Off Kyûshû

HOK

北海道 Hokkaidô — 日本本土 Mainland JapanHON

本州 Honshû — 日本本土 Mainland Japan

Hrd

平戸 Hirado — 九州沖 Off Kyûshû

Ihy

伊平屋島 Iheya-jima — 南西諸島 Nansei-shotô (Ryukyu)Iki

壱岐 Iki — 九州沖 Off Kyûshû

Irb

伊良部島 Irabu-jima — 南西諸島 Nansei-shotô (Ryukyu)

Iri

西表島 Iriomote-jima — 南西諸島 Nansei-shotô (Ryukyu)

Ish

石垣島 Ishigaki-jima — 南西諸島 Nansei-shotô (Ryukyu)

Is-Hch八丈島 Hachijô-jima — 伊豆諸島 Izu-shotôIw

硫黄島 Iô-jima — 小笠原諸島 Ogasawara-shotôIz-A

青ヶ島 Ao-ga-shima — 伊豆諸島 Izu-shotô

IzIss

伊豆諸島 Izu-shotôIz-Kz

神津島 Kôzu-shima — 伊豆諸島 Izu-shotôIz-Mk

御蔵島 Mikura-jima — 伊豆諸島 Izu-shotôIz-My

三宅島 Miyake-jima — 伊豆諸島 Izu-shotôIzn

伊是名島 Izena-jima — 南西諸島 Nansei-shotô (Ryukyu)

Iz-N

新島 Nii-jima — 伊豆諸島 Izu-shotôIz-O

大島（伊豆）Ôshima (Izu) — 伊豆諸島 Izu-shotôKbs

樺島 Kaba-shima — 九州沖 Off Kyûshû

KDt

北大東島 Kita-daitô-jima — 南西諸島 Nansei-shotô (Ryukyu)

Kik

喜界島 Kikai-jima — 南西諸島 Nansei-shotô (Ryukyu)

KIw

北硫黄島 Kita-iô-jima — 小笠原諸島 Ogasawara-shotô

KKs

上甑島 Kami-koshiki-jima — 九州沖 Off KyûshûKokn

沖ノ島（筑前）Oki-no-shima — 九州沖 Off KyûshûKOs

大島（紀伊）Ôshima (Kii) — 本州沖 Off Honshû

Krs

薩摩黒島 Kuro-shima — 南西諸島 Nansei-shotô (Ryukyu)

Kue

口永良部島 Kuchinoerabu-jima — 南西諸島 Nansei-shotô (Ryukyu)

Kum

久米島 Kume-jima — 南西諸島 Nansei-shotô (Ryukyu)

Kun

国後島 Kunashiri-tô (=Kunashir) — 北海道沖 Off Hokkaidô

KYU

九州 Kyûshû — 日本本土 Mainland Japan

MDt

南大東島 Minami-daitô-jima — 南西諸島 Nansei-shotô (Ryukyu)

MIw

南硫黄島 Minami-iô-jima — 小笠原諸島 Ogasawara-shotô

Myk

宮古島 Miyako-jima — 南西諸島 Nansei-shotô (Ryukyu)

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Nkd

中通島 Nakadôri-jima — 九州沖 Off Kyûshû

Og

小笠原諸島 Ogasawara-shotôOg-cc

父島 Chichi-jima — 小笠原諸島 Ogasawara-shotôOg-hh

母島 Haha-jima — 小笠原諸島 Ogasawara-shotôOg-ot

弟島 Otôto-jima — 小笠原諸島 Ogasawara-shotôOke

沖永良部島 Okinoerabu-jima — 南西諸島 Nansei-shotô (Ryukyu)Oki

隠岐 Oki — 本州沖 Off Honshû

Okn

沖縄島（本島）Okinawa-jima — 南西諸島 Nansei-shotô (Ryukyu)

Okns

沖ノ島（四国）Oki-no-shima (SHI) — 四国沖 Off ShikokuOks

奥尻島 Okushiri-tô — 北海道沖 Off Hokkaidô

OOs

大島（渡島）Ôshima (Oshima) — 北海道沖 Off HokkaidôReb

礼文島 Rebun-tô — 北海道沖 Off HokkaidôRK

南西諸島 Nansei-shotô (Ryukyu)

Rsr

利尻島 Rishiri-tô — 北海道沖 Off Hokkaidô

Sad

佐渡島 Sado-ga-shima — 本州沖 Off Honshû

Shd

小豆島 Shôdo-shima — 四国沖 Off Shikoku

SHI

四国 Shikoku — 日本本土 Mainland JapanSKs

下甑島 Shimo-koshiki-jima — 九州沖 Off KyûshûT-Aks

悪石島（吐噶喇）Akuseki-jima (Tokara) — 南西諸島 Nansei-shotô (Ryukyu)Tan

種子島 Tane-ga-shima — 南西諸島 Nansei-shotô (Ryukyu)

T-Nak

中之島（吐噶喇）Naka-no-shima (Tokara) — 南西諸島 Nansei-shotô (Ryukyu)

Tob

飛島 Tobi-shima — 本州沖 Off Honshû

Tok

徳之島 Toku-no-shima — 南西諸島 Nansei-shotô (Ryukyu)

T-R

吐噶喇列島 Tokara-rettô — 南西諸島 Nansei-shotô (Ryukyu)

Trm

多良間島 Tarama-jima — 南西諸島 Nansei-shotô (Ryukyu)

Tsm

対馬 Tsushima — 九州沖 Off Kyûshû

T-Sw

諏訪瀬島（吐噶喇）Suwanose-jima (Tokara) — 南西諸島 Nansei-shotô (Ryukyu)

T-Tak

宝島（吐噶喇）Takara-jima (Tokara) — 南西諸島 Nansei-shotô (Ryukyu)

T-Yok

横当島（吐噶喇）Yokoate-jima (Tokara) — 南西諸島 Nansei-shotô (Ryukyu)

Yae

八重山諸島 Yaeyama-shotô — 南西諸島 Nansei-shotô (Ryukyu)

Yak

屋久島 Yaku-shima — 南西諸島 Nansei-shotô (Ryukyu)

Yon

与那国島 Yonaguni-jima — 南西諸島 Nansei-shotô (Ryukyu)

Yor

与論島 Yoron-tô — 南西諸島 Nansei-shotô (Ryukyu)

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218 University Museum柴田 泰利・丸山 宗利・保科 英人・岸本 年郎・直海 俊一郎・
野村 周平・Volker Puthz・島田 孝・渡辺 泰明・山本 周平

日本産ハネカクシ科総目録（昆虫綱：甲虫目）

Reprinted from Bulletin of the Kyushu University Museum No. 11
pp. 69-218, March 2013