NEW OR LITTLE-KNOWN TIPULIDA FROM EASTERNASIA (DIPTERA), VII'
By Cnnrr,os P. AT,EXANDER
Of Amherst, Massachusetts
TWO P',ATES

The crane flies described herewith are almost all based on
extensive collections made in southern Japan and Formosa by
my friend Prof. Syuti Issiki, and on Mount Omei, Szechwan,
China, by a Canadian collector and secured through Mr. Herbert
s. Parish. A few additional Japanese species were collectedby Messers. Esaki, Sakaguchi, Ueno, and Uye. The types of
the new species are preserved in my collection through thekind g:enerosity of the entomologists above named, to whom
my deepest thanks are extended.
As a slight contribution to our knowledge of the fipulide ofJapan and Formosa, I am recording in full the species taken
by Professor Issiki on Mount Kirishima, in Kiushiu; on Yakushima Island, south of Kiushiu; and on Arisan, a mountainstation in Formosa.
TIPULIDA FROMMOUNTKIBISHIMA, KIUSHIU
Mount Kirishima, Kiushiu, altitude, 2,500 to 3,500 feet, May
3 to 7, L929; mostly at 2,500 feet, May 3, Lg29 (SUuti Issiki) .
ma'chidai' (Alex.).
Li.monia (Limonia)(Dic,ranorryia)
basitusca (Alex.).
Limonia
I'imonia (Dipranomaia) immodestoides (Alex.).
Li,monia (Dioranom'Aia) purwtulata (de Meij.).
Antocha (Antochn) bifila Llex.
T ricgphona in"sulana Alex.
Rhaphidolabis coneors Alex'
Dactglolabi.s longicauda .Llex,
P seu.ilolimnophila horii .Llex.
Limnophita (Tri.chol;imnophi.Io) caesietla .lllex.
saitama'e Alex.
I'imnoph;ila (Tri'cholimnophila\
Li,mnophila (Ephelia') dietziana Llex.
Limnophila (Prianolabis) liponeura sp. nov.
Limnophila (Prionolabis) submn'niln Alex,from" Contribution
Agricultural
College.

theDepartmentofEntomology,

Massachusetts
b0?

508

The Phi,l;i,ppineJotnnal of Science

N ippol:bnn ophila kiusiuens?s Alex.
Eleplw,ntorngia dietziana sp. nov.
Eri,optera (Empeda) microtrichia,ta Alex.
Erioptera (Ili8if,, asymmztrica Alex,
Ormoaia, iliaersipes .lJex.
Ortnosi.q, tekeuchii Llex.
Moloph.ilus albohalterata Llex.
Molophilus pegosus Alex.
TrpuLID.iE

taoM

KOSUGIDANI, YaKUSIIIMA

Yakushima, an island south of Kiushiu; collections made at
Kosugidani, altitude 2,500 feet, in rich native forest, April 29,
1929 (Syuti Issi&r). Esaki2 outlines the differences existing
between the mountainous, richly wooded island of Yakushima
and the indicated presencethereon of Palearctic forms, and the
lowlying nearby Tanegashima Island. The species of theaccompanyinglist are predominantly Palearetic in affinities andso confirm Esaki's statement.
Limonia (Di,scobola) margarita (Alex.).
Lim.onia (I)monia\ anth.racina Alex.
Limonia (Limonia \ nnci,idai (Aiex.).
(Lim,onial ntortacartha (Alex,),
LimniaLimonia (Limonia) f risirnrr sp. no\'.
Limonia (Limonia) yaku-"i in,ii,s:s sD. nov.
(
Limonia (Dicronornyia\ i',a,nr',.,;.1;.;,-{lex. ) .
Iimoruia (Geranomyia) rndiaiis -{lex.
Ant ocha ( Antocha) subco n.r1.,,,7,- rr srr. nui'.
Helius (Helhrc\ oblifercla-" -{lex.
Orimarga gakushimana sp. no\'.
T ricyphona yakushimo,na AIex.
I)icranota (Antalopina) gibberu ( -{lex. ) .
Ductylolttbis longicauda megustglota subsp. r'rov,
Pse u.d.olimnoph,ila h.orii AIex.
Umnophila (Prionolabis) Iipophleps sp, nov.
Limn.o1th,ila (Prionolabi,s\ subrnurida Al,ex.
Limnophila subnemnralis Alex., var.
Ulomorplta pol'gtricha Llex.
N'ippolimno1thila Eaku.sh.inr.erzsis
Alex.
Atarba (Atarbodes) mhtu.ticorrtissp. nov,
Lipsotlrix Eakushimae Alex,
Erioptera (Empe&t) mict'otrichtata Alex.
TIPULID.E FROM ARISAN, FOR,MOSA

Arisan, Formosa, altitude 6,000 to 8,000 feet, July 7 and 8,
1929; mostly at 7,300 feet, July 7,1929 (Sauti Issi,ki).
ro trichopygo Alex,"\'csoT,reC gtt a r omgia taiwania, Llex.
'Allattani

Kdzlem6nyek 23 (1926, I27.

I

42,4

Alerand,er: Tipul,id,p from Eastem

Asi,a, VII

bOg

Limonia. (Discobota) argus (Say).
Itimania ( Dicranomgin) shirakii- (.!irex,.t.
(Helius) liliputatats Atex.
!_"li*
Helitrs (Helius) rufithoras
Alii.
Tricgphorw arisana Alex.
Tricgphona formosarw Alex.
Ad,elphomgia, aria,na Alex.
UIa flaaidtibasis Alex.
Pseud,olimnophita
iUustris .Llex.
Pseudolimnophila
ma,rcid,a Alex.
Limnophila (Trichol:imnopnlby
ptatgstyla TnralleLo subsp.
nov.
I:i.,mtophila (Dicranophrigrr*id"rlrii,"es,tq, sp. nov.
Umn1pltiln(Dicranophragrr*i/lt"*.
7or*o'ro"uu.rorrnis
Elephantomyia(Eteihanio*ioi"tl
Arex.
(E lephantomyaO i"ii\*allephantomgia
Atex.
Atarba(Atarboiles)f"r"Loil.ir il;""

Atarba, (Atarbod,es) isei.kiana,rorr.
Atarba (Atarbodee) teptouantni' "p.
/ltix.
Cerato cheilw tinctipends .Llex,
Dasyntallom,yia signata Brun.
Eri.oFtera (Erioptera)
alboguttata Fodw,--'
Erioptera (Iti^sia) tenuisentL
Ai;;.
Erioptera (Empeda.) a"gusttstiim;i|re*,
D asy malophilu s na k o ensis AIex.
Molophilus aricolo sp. nov.
Molophilus nigritarsis
Alex,
Molophilus nigdtus
Alex.
TIPULIN.€
TIPULINI
TIPULA LATILIGULA Tp. noY.

General coloration yellowish gray,the prescutum with four
scarcely indicated brown stripes;
uot*n*
yellow; wing:s grayish, the stigma brown; obliterative
areas conspicuous; m_cu
near the base of cell 1st M,; malehypopygium with the median
0f tergite produced into a compressed:"ci-olbrade that is subtended by a smalr acute spine o'eitier side; eighth sternite
bearing a very broad, triangular tonguelike
lobe.
Male.-Length, ab_outf t mittimete"rs;wing, tZ.S.
Frontal prolongation of head relatively
short and stou!
brownish gray; nasus short and stoul.
Antenne of moderate
Iength, if bent backward extending
abount to the harteres orslightly beyond; the segments yetow,
their bases -,rot o"-*"u"."rydarkened, the outer segments rtigttty
infuscated. rr*o"r}ro*ish gray, the vertical tubercle uu"y io*.
Mesonotum yetowish gray, the -p"aslutumwith four scarcery
indicated brown stripes. pleura yellowish
gray. lfuil"r"Jp"f",
the knobs dark brown. Legs withthe coxe yellowish gray,

510

The Phikppine Jotnnal of Sciencetsso

the trochanters yellow; femora brownish yellow, the tips infuscated; tibiee and tarsi passing into dark brown. Wings somewhat teneral, grayish, the brown stigma darker; large andconspicuous o,bliterative areas before the stigma and especiallyacross the base of cell lst M' involving large portions of cells
R and Mu; veins dark. Venation: m-cu close to the inner endof cell 1st M, M3*, being less than one-half r-m; m-cu very
long and oblique.
Abdomen chiefly brownish gray, shrunken and distorted in thetype. Male hypopygium (Plate 2, fig. 19) with the tergite extensively fused with the sternite. Ninth tergite (Plate 2, ftg.
20) tridentate, the median region produced into a long compressed blade that is slightly upcurved at apex, subtended oneither side by an acute spine of less than one-half the length.
Outer dististyle, od,,a large, pale, boomerang-shapedlobe, bearing a tuft of dusky sete at tip. Ninth sternite, 9s, gently emarginate, with a small pencil of about three spinous setre on eitherside near the mid-line. Eighth sternite (Plate 2r fig.21) bearing on caudal margin a very large, broadly triangular, troughlike
lobe, the greatest width exceedingthe length.
H abi,tat.-Japan (Honshiu ) .
Holotype, male, Norikura-ga-take, Shinano, July 22, 1929 (M.
Ueno).
Ti,pula latiligula will probably be found to be strictly an
alpine species. The structure of the male h1'popygium readily
separatesit from all other regional forms.
LIM ONiIN,€
LIM ONIINI
LIMONI^{(DICRANOMT"IA)KANSUENSIS

sp. nov.

General coloration opaque brown I rostrum, palpi, and antennaeblack; wings strongly tinged with brown; cell lst M'?closed;
male hypopygium with the rostral prolongation of the ventral
dististyle short and stout, bearing two very unequal spines, theinner one very tiny.
Male.-Length, about 6.5 millimeters ; wing, 7.3.
Rostrum and palpi black. Antennre black throughout; flagellar segments oval, gradually decreasing in size outwardly.
Head yellowish gray.
Mesonotum broln, opaque, without clearly defined stripes;
secutellum and postnotum more pruinose. Pleura brown,
sparsely pruinose. Halteres pale, the knobs infuscated. Legs
with the coxe pale yellow, the fore coxre slightly infuscated on

42,1

Alesanl,er: Tipuliiln

from Eastem Asia, Vil

511

outer face at base; trochanters yellow; remainder of legs yellow,
the outer tarsal segments more infuscated. Wings (Plate 1,
fig. 1) strongly tinged with brown, the oval stigma still darker
brown; scarcely evident darkenings along cord and outer end ofcell lst Mr; veins brown, more yellowish brown basad of thecord. Venation Sc, ending just beyond the origin of Rs, Sc, a
short distance before this origin ; cell 1st M, closed; m-cu atfork of M; vein 2d A rather strongly sinuous.
Abdomen brown, the ventral dististyle of the hypopygium
paler. Male hypopygium (Plate 2, fig. 22) with the laterallobes of the tergite, 9f, conspicuous, each with a group of sete;median region of tergite further produced into a transverse
flaplike structure. Ventral dististyle, od, smaller than the basistyle, the rostral prolongation unusually short and stout, bearing at near midlength two very unequal spines, the outermost
longest, approximately one-half the length of the prolongation,
the inner spine very small. Dorsal dististyle a glently curved
blackened rod, the tip acute. Gonapophyses, gf, with the mesalapical lobe very long and slender.
(Kansu) .
Habitat.-China
Holotypg male, Hweihsien (ex Staudinger-Bang Haas).
The structure of the male hypopygium separates this species
from all similar forms in this general region.
LIMONIA(LIUONIA)YAKUSEIMENSIS rp. nov.

Mesonotal prescutum and scutal lobes black; remainder ofthorax brown; antennae brownish blaek throughout, basal segments subglobular; wings strongly suffused with brown; malehypopygium with the dorsal dististyle reduced to a miscroscopic
structure; rostral prolongation of ventral dististyle with two
distinctly separated spines of approximately equal size.
Male.-I'ength, about 5.6 millimeters; wing, 6.8.
Rostrum and palpi very much reduced, the former reddish
yellow. Antenna brownish black throughout; basal flagellar
segments subg:lobular,with very short apical necks; outer segments passing into oval, each with a pair of long, unilaterally
arranged verticils that are much longer than the segments;
outermost segments fusiform, the terminal segment longest,
about one-third Ionger than the penultimate. Head black, subnitidous.
Pronotum blackish. Mesonotal prascutum and scutal lobes
black, subnitidous; median region of soutum and the scutellum
pale, the surface sparsely pruinose; postnotum pale brown.

5L2

The Phili,ppine Journnl of Science:sso

Pleura brown, sparsely variegated with darker, the surface
more or less pruinose. Halteres of moderate length, pale, theknobs infuscated. Legs with the fore eoxa blackened,pruinosel
middle and hind coxe pale; trochanters obscure yellow; remainder of legs brown, the outer tarsal segments even darker.
Wings (Plate 1, fig. 2) with a strong brown suffusion, the oval
stigma only slightly darker; linear pale streaks in cell R, crossing
cell lst M, and in cell M, crossing m-cu into cell M"; veinsbrown. Venation: Sc, ending about opposite two-thirds thelength of Rs, Sc, at its tip; Rs sinuous, nearly straight on basal
fourth, the remainder more arcuated; cell lst M, closed; m-cu
strongly areuated, longer than the distal section of Cu, placed
shortly beyond the fork of M.
Abdominal tergites dark reddish brown, the extreme caudal
margins of the segments narrowly blackened; basal sternites
more yellowish; male hypopygium pale. Male hypopygium(Plate 2, frg. 23) with the tergite, 9f, transverse, the caudal
margin gently emarginate medially, with a group of long eoarse
sete on either side. Basistyle, b, with the ventromesal lobe
nearly apical in position. Dorsal dististyle reduced to a microscopic structure, as in the group. Ventral dististyle, od, oval,
the rostral prolongation elongate, bearing two slender rostralspines that arise from a common tubercle shortly beyond thebase of the prolongation.
Habi.tat.-Japan (Kiushiu).
Holotype, male, Kosugidani, Yakushima, altitude 2,500 feet,
April 29, 1929 (S. Issiki).
Umonia, galatshimensis is related to L. frauilulenta Alexander(Formosa) and,L. ubensis Alexander (Luzon), differing in thecoloration of the body and the structure of the male hypopygium,
especially the position and shape of the rostral spines of theventral dististyle.
LIMONIA(LIUONIA)TRISTINA

sD. nov.

Female.-Length, about 9 millimeters; wing, 9.
Closelyallied to L. (L.) anthracinn Alexander (Japan), differing especiallyin the uniformly darkenedwings and structure of the ovipositor.
Rostrum, palpi, and antenna black. Head black.
Mesonotumblack, the posterior portions of the scutal lobes
and the postnotum with piceous reflections. Pleura with thedorsopleural region and the propleura black, the remainder
chiefly dark castaneous,the dorsal portion of the sternopleurite

42,4Aleuand,er: Ti,puhid,afrom Eastern Asin, Vil513

blackened. Halteres pale, the knobs blackened. LeeF with thefore coxa blackened, the middle and posterior coxe paler;trochanters reddish brown; remainder of legs black, only thefemoral bases restrictedly pale. Wings with a strong blackish
suffusion, the costal and stigmal regions more suffused I vague,
scarcely apparent seams along cord and outer end of cell lst
M"i a conspicuous elouding along vein Cu. in cell M; veinsdark brown. Venation: Sc, ending before midlength of Rs, Sc,
at its tip; R, about one-half Rr*ri m-cu just before the forkof M.
Abdominal tergites black, the sternites yellow, the basal andintermediate sternites with the caudal margins broadly brownish
black. Ovipositor with the tergal valves much smaller, slenderer, and more curved than in antluracfum.
Habi,tat.-Japan (Kiushiu) .
Holotype, female, Kosugidani, Yakushima, altitude 2;500 feet,
April 29, 1929 (S. /ssiki).
LIXONIA(LIIIONIA)UNICINCTIFERA

sp. nov.

Generalcoloration brown; pleura yellow, striped longitudinallywith brown; femora yellow, the tips blackened;remainder oflegs snowy white, the tibia with a narrow black ring before midlength; wings tinged with yellowish gxay, the costal region
brighter; cell lst M, closed; abdominal segmentsannulated yellow and brownish black; male hypopygium with a single rostralspine that arises from a stout tubercle.
Male.-Length, about 7 millimeters; wing, 6.7.
Femnl,e.-Length, about 5.5 millimeters; wing, 5.5.
Rostrum gray, the mouth parts and palpi brownish yellow,
much redueed in size. Antenne with the scape dark brown,
the flagellum brownish black; flagellar segxnentsoval, passing
through elongate-oval;verticils longer than the segments; terminal segment about one.third longer than the penultimate.
Head dark brown.
Pronotum yellow. Mesonotal prescutum brown, the lateral
margins narrowly obscureyellow; scutal lobesdarkened; medianregion of scutum and the scutellum more pruinose; postnotum
slightly pruinose. Pleura yellow with a namow brown dorsomedian longitudinal stripe; ventral sternopleurite more or lessdarkened. Ilalteres dusky, the knobs dark brown. Legs withthe coxe obscure yellow; trochanters brown; femora yellow,
the tips conspieuouslyblack; tibie pure white with a blaek ring
before midlength, this about one-half as wide as the white basal
262890-{

5L4

The Phili,ppi,ne Jourrwl of Sci,ence

rcBo

portion; tarsi snowy white, including the last segment. Wings(Plate 1, fu. 3) tinged with yellowish gray or brown, the costal
region and outer portions of the radial cells clearer yellow;
stigma oval, dark brown; narrow and barely evident dark seams
along the cord; caudal and posterior regions of wing more infumed, especially in the female; veins brown, Sc more yellow.
Venation: Sc, ending beyond midlength of the short Rs, Sc, at
its tip; Rs angulated and weakly spurred close to origin; cell
Lst M, closed; m-cu close to fork of M; vein Zd A at, origin
running close to margin, thence approaching vein lst A.
Abdominal tergites brownish black, narrowly ringed caudallywith yellow; sternites yellow, narrowly ringed basally withbrown; hypopygium chiefly darkened. Male hypopygium (plate2, fig.24) with the caudal marg:in of tergite, gt, deeply emarginate. Dorsal dististyle a strongly chitinized rod, the apex
abruptly narrowed into a gently decurved point. Ventral dististyle, ad,, fleshy, the rostral prolongation short and stout, atbase bearing a large tumid lobe that is larger than the prolongation itself, extended into a single long straight spine. Gonapophyses, g, with the mesal apical lobe flattened, pale, terminating
in an acute point. .lEdeagus, o, broad, the apex dilated into
two divergent lobes, each lobe with four punctures.
H abi.tat.-China ( Szechwan) .
Holotype, male, Mount Omei, altitude 4,b00 feet, July 25, Lg?g
(ex Parish). Allotopotype, female.
Limoni,a unicinctif ero is very distinct from all regional species.
fn some respects it suggests members of the subgenus pseud,oglochina, but it is certainly a true Limonin.
EELIUS (EELIUS) COSToFIMBRIATUS gp. nov.

General coloration brown; legs dark brown, the tips of thetarsi paler; wings with a stronger brown tinge; costal fringeof male long and conspicuous; cell lst M, irregularly pentagonal, m being oblique in position.
Male.-Length, about 7.5 to 8 millimeters; wing, Z.
Female.-Length, about 9.b to l0.millimetersl wing, g.
Rostrum a little longer than the head, dark brown; palpi
black. Antenne black throughout, relatively small, a litile
longer than the rostrum. Head brownish black.
Mesonotum chiefly brown, the prascutum rich cinnamon brownwith darker markings; postnotum extensively pale. pleura
brownish yellow, indistinctly variegated with darker. Halteres
dirty white, the knobs infuscated. Legs with the coxe and42,4

Ale*and,er: T'ipulid,p from Eastern Asin, VII

515

trochanters testaceous; femora brown, their bases slightly paler;tibie and tarsi dark brown, the tips of the latter paling
to brownish yellow. Wings (Plate 1, fig. 4) strongly tinged
with brown, the stigmal region darker brown; veins brown.
Costal fringe of male long and conspicuous. Venation: Cell lst
M, irregularly pentangonal, m being oblique in position andusually longer than the second section of Mr*ri m-cu at orclose to fork of M.
Abdominal tergites dark brown, the sternites brownish yellow;
hypopygium brownish yellow. Male hypopygium much as intenui,stylus, the basistyles with a conspicuous spinulose lobe on
mesal face at base.
(Loochoo Islands).
Habitat.-Japan
Holotype, male, Kunjan, Okinawa, altitude 500 to 1,000 feet,
May, 1923 (5. Sakaguchi,), Allotopotype, female. Paratopotypes, 3 broken males.
Heli;us costofr,mbrinfzrs is closely allied to the Formosan Il.
tenuistulus Alexander, differing most evidently in the conspicuous costal fringe of the male.
HELIUS(EDLIUS)PALL-IDISSIMUS

sp. nov.

General coloration pale yellow, including the halteres and mostof the legs; head light gray; wings yellow, the veins deeper
yellow; cell lst M, rectangular; m-cu close to fork of M.
Female.-Length, about 7.8 to 8 millimetersl wing,6.6 to 7.
Rostrum of moderate length, a little longer than the combinedhead and neck, brownish yellow; basal segments of palpi pale,the outer segments brownish black. Antenna with the scapal
segments obscure yellow, the flagellum brownish black; basal
flagellar segments short and crowded, the outer segments longand slender; verticils elongate, especially on the outer segments.
Head light gray.
Thorax uniformly pale yellow. Halteres pale yellow. Legs
with the coxe and trochanters pale yellow; femora yellow, theremaining segrnents passing into yellowish brown. Wings(Plate 1, fig.5) tingedwith yellow; veins pale yellow to brightyellow. Venation: Sc, ending shortly before fork of Rs, Sc, at
its tip; anterior branch of Rs relatively short and nearly straightbeyond origin, the cell at margin correspondingly widened; cell
1st M, large, rectangular; m-cu close to fork of M.
Abdomen reddish yellow, the caudal margins of the segments
narrowly paler. Ovipositor with the valves very long andslender, dark horn-yellow.

516

The Pluilippine Journnl of Sci,ence:r,ao

H abi,tat.-China ( Szechwan).
Holotype, female, Mount Omei, altitude 4,b00 feet, July 2g,
L929 (ex Parish). Paratopotype, female, altitude g,000 feet,
July 25, L929.
In its unusually pale coloration, Helius pallid,issi,musresembles
H. uni,color (Brunetti), of the Himalayas, differing in the large
size and very different venation of the medial field.
ANTOCEA (ANTOCEA) SUBCONFLUENTA sp. nov,

General coloration of mesonotum dark gray, the humeral andlateral portions of the prescutum buffy; wings suffused with
gray; cell lst M, tending to be open by the atrophy of m; malehypopygium with the outer dististyle flattened, the outer marginnear tip with two or three appressed serrations; adeagus small.
Male.-Length, about 4.8 millimeters; wing, b.b.
Rostrum brownish yellow; palpi black. Antennre black
throughout. Head dark gray, the orbits clearer gray.
Mesonotal prrescutum almost covered by three confluent darkgray stripes, restricting the ground color to buffy humeral andlateral portions; posterior sclerites of mesonotum dark gray,the scutellum slightly paler. Pleura dark brownish gray, thepteropleurite more reddish brorvn. Halteres pale, the knobs
dusky. Legs with the coxe reddish brown, the fore coxa
darker; trochanters obscf re brownish yellow; remainder of legs
brown, the middle legs and all tarsi somewhat darker. Wings(Plate 1, fig. 6) with a strong gray suffusion, the stigma scarcely
indicated; veins pale brown. Anal angle of wing moderately
developed. Venation: Cell lst M, open by the partial atrophyof m.
Abdominal tergites brownish black, the basal sternites somewhat brighter; hypopygium obscure yellow. Male hypopyglum(Plate 2, fig. 25) with the caudal margin of the tergite, 9f,
transverse to gently concave,with a series of coarse, chiefly marginal sete. Outer dististyle, od, simple, the margin flattened
into a blade, with two or three appressedserrations before apex.
Gonapophyses, gt, appearing as slender sinuous yellow rods, thetips acute. .4Edeagus,o, small.
Habi.tat.-Japan (Kiushiu) .
Holotype, male, Kosugidani, Yakushima, altitude 2,500 feet,
April 29, 1929 (S. Issi,ki.).
Antochn subconfl,uenfo is apparently most closely related toA. confl,uenfaAlexander (eastern China), differing in the vena-

42,4Alenand,er: Tipulid,z from, Eastem Asia, VII

5L7tion and outline of the wing and in the structure of the malehypopygium.
ORIMARGACRUCIFORUIS

sp. nov.

General coloration plumbeous gray; halteres pale, the knobs
infuscated; wings long and narro\y, tinted with brown, the apex
infuscated, cell R, remaining pale; R, in approximate alignment
with the basal section of Rr_- to produce a f-shaped figure;medial forks short; cell 2d A long and narrow.
Female.-Length, about 6.5 millimeters; wing, 5.5 by 1.
Rostrum and palpi black. Antenna with the basal segment
black, the remaining segments dark brown; flagellar segments
globular, the outer segments passing into short-oval. Headpruinose.
Thorax blue-gray pruinose, the pleura somewhat clearer gray.
Halteres pale, the knobs infuscated. Legs with the coxa andtrochanters dark reddish brown; remainder of legs broken.
Wings (Plate l, fi9. 7) unusually long and narrow, as shownby the measurements; anal region greatly reduced; membrane
pale basally, gradually darkening outwardly, the apex narrowly
infuscated, interrupted by the entirely pale cell Ru; veins black.
Costal fringe relatively long and consp,icuous. Venation: Sc,
ending just beyond midlength of the long Rs, Sc, at its extreme
tip; a supernumerary crossvein in cell Sc, opposite origin ofRs; R, in approximate alignment with the origin of Ro*u.
Rs being in alignment with R, and Rro, lackingi R, and Rr*,nearly equal; basal section of Rr*u angulated and weakly
spurred before midlength; r-m far beyond level of R,; cell M,
shallow; m-cu opposite the basal fourth of Rs; cell 2d A longand narrow.
Abdominal tergites brownish black, the sternites somewhat
lighter brown. Ovipositor with the tergal valves slender but
relatively short.
(Szechwan).
Habitat.-China
Holotype, female, Mount Omei. altitude 6,000 feet, July 25,
1929 (ex Parish).
Ori,marga cruci,formis is very distinct from all described regional species.
OBIMARGAYAKUSEIUANAsP. nov.

General coloration gray; wings with a faint dusky tinge;
veins pale brown; medial field with the outer forks relatively
deep.

The Phi,lippi,ne Jourrwl of Science

518

rsso

Male.-Lengbh, about 6 to 6.5 millimeters; wing, 5.6 to 6.2.
Female.-Length, about 6.5 to 7 millimeters; wing, 6 to 6.5.
Rostrum gray; palpi black. Antenna black throughout ; flagellar segments short-oval, the outer segments more elongate.
Head gray.
Mesonotum light gray, the median region of the prascutum
and the scutal lobes somewhat darker gray. Pleura gray.
Halteres pale. Legs with the coxa dark; trochanters reddish
brown; remainder of legs brown. Wings (Plate 1, fig. 8) witha faint dusky tinge; veins pale brown, the outer costal reg:ion
more yellowish. Macrotrichia almost as in prwinosa. Venation: Sc long, Sc, ending just before the fork of Rs; free tip ofSc, somewhat variable in relation to the position of R,, the distance from one to two times the total length of the latter; Rr*,
considerably longer than R, alone; forks of medial field much
deeper than in pruinoso.
Abdomen, including the hypopygium, dark brown.
Habitat.-Japan (Kiushiu).
Holotype, male, Kosugidani, Yakushima, altitude 2,500 feet,
Allotopotype, female. ParatopoApril 29, 1929 (S. lssiki).
types, 6 males and females.
Orimarga yakushinzaizc is most nearly allied to O. prui,nosa
Alexander (Japan: Honshiu), differing in the larger size, darker
wings and veins, and the deep forks of the medial field.
ORIIIARGAOMEINA

sp. nov.

General coloration dark gray; wings milky white, the veins
pale yellowish brown to reddish brown; costal fringe of moderate
length only; R, lying beyond level of r-mi Rr*, variable inlength, from two to three times Rr; anal cells large.
Male.-Length, about 6 to 7 millimeters I wing, 5.5 to 6.8.
Female.-Length, about 7 to 7.5 millimeters; wing, 6.5 to 7.5.
Rostrum gray; palpi black. Antenne with the basal segmentof the scape varying from yellow to black, the remaining
segments black. Head gray.
Thorax dark gray, without distinct markings. Halteres palethroughout. Legs with the coxa reddish brorn'n; trochantersbrownish yellow; femora yellowish brown; tibiae and tarsi
brownish black to black. Wings (Plate 1, fig. 9) milky white,
the veins pale yellowish brown to reddish brorvn. Costal fringeof moderate length only; no macrotrichia on Rs. Venation: Sc
long, Sc, ending just before the fork Rs; R, far beyond r-m;

4?^4

Aleranfi,er: Tipuliila from Eastem Astn, VII

519

Rr*, in most cases elongate, approximately three times &alone,in casesshorter, approximately twice Rr; anal cells large.
Abdomen dark brown, more or less pruinose.
Habitat.-China (Szechwan).
Holotype, male, Mount Omei, altitude 4,500 feet, Aug:ust 2,
L929 (ex Parish). Allotopotype, female, August L4, L929.
Paratopotypes, several males and females, July and August,1929.
Or[,margaomefumis very closeto O. galrushim&ru&
sp,nov. andmay more properly be consideredas being only a variety of thelatter species,differing especiallyin the milky-white wings withpale veins, and in slight details of venation. The present species
appearsto vary to an unusual degree. The two types have thelargest measurementsgiven above, in conjunction with an unusually long Rr*r.
ORIUARGA SETICOSTA rD. nov.

General coloration brownish gray; thoracic pleura with anarrow black longitudinal stripe; halteres pale throughout;wings yellow, the veins darker yellow; costalfringe long and conspicuousin both sexes; R, and r-m in nearly transverse alignment.
Male.-Inngth, about 5 to 5.5 millimeters; wing, 4.5 to 4.8.
Fema,Ie.-Length, about 6.5 millimeters; wing, 5.5 to 6.
Rostrum and palpi black. Antenna n'ith the scapalsegments
black, the flagellum brown; flagellar segments short-oval, theouter segmentsbecomingslightly more elongate. Anterior partof head buffy, the remainder gray.
Thoracic notum brownish gray to gray, the prescutum witha slightly darker median stripe, the latter in casesweakly bifid;
humeral region brighter; posterior sclerites of notum darker,
the median area of scutum pale. Pleura brownish gray with anaruow but conspicuousblack longitudinal stripe extending fromabove the fore ccixa to beneath the wing root; ventral sternopleurite narrowly darkened. Halteres pale throughout. Legs
with the coxe and trochanters brownish yellow; femora brownish
yellow; remainder of legs gradually passing into brown, theterminal tarsal segmentsblackened. Wings (Plate 1, fig. 10)
relatively broad, especiallythe anal region, the widest point ofwing being at near midlength; ground color pale yellow, theveins darker yellow. Costal fringe of both sexes long andconspieuous. Venation: Sc, ending about opposite two-thirds

520

The Plai,lippi,neJorn'rwl of Scienca$sothe length of Rs, Sc, at its tip; free tip of Sc, faint, a little
more than its own length before R,; Rs relatively long, arcuated;
Rr*, lon8er than Rr*r; basal section of R.*u angulated before
midlength i Mr*, subequal to Mn; m-eu about opposite midlenethof Rs; anal cells unusually wide, especially cell 2d A.
Abdomen brown, the margins of the segmentsmore blackened;hypopygium brown.
Habitat.-China ( Szeehwan).
Holotype, male, Mount Omei, altitude 4,500 feet, August ,.
tg29 (ex Parish). Allotopotype, female. Paratopotypes, 5males and females, July 24 to August 8, 1929.
Orimnrga seticosta is readily distinguished by the diagnostic
features listed above, especially the long costal fringe in both
sexes'
PEDrcrrNr
DICRANOTA(AIiIALOPINA)

GIBBEBAKARAFUTONIS

gubsp. nov.

Close to the typical form, differing especially in the narrowwings of the male (and undoubtedly both sexes). The wing
(Plate 1, fig. 11) is of approximately equal width along the
middle third of the width, whereas in the male of typical gibbera(Plate l, fig. L2) the wing is widest opposite the termination of
the 2d anal vein, cell 2d A is wider and the auxillary margin
is gently incised.
Habitat.-Japan (Karafuto) .
Holotype, male, Maoka, July 28, L922 (7. Esaki). Paratopotypes,6 males; paratype, l male, Manui, August 3, 1922 (T.
Esaki).
Dicranota (Amnlopi,no) gibbera is closely allied to D. (4.\elegantu,la (Brunetti), of southeastern Asia. Amn'lopi'na, together with Rhaphidolobis and. Plectromgia, would seem to be
more coruectly referable to Dicranota as subgeneric groups,
although the distinctions existing between several of the socalled genera of pediciine crane flies are very slight. The relation of the narrow-rvinged males in this and other groups of Tipulide to closely allied forms rvith broad wings in the male isstill poorly understood and in the present case, at least, is best
expressed by a trinomial until more data become available.
Edwards * has discussed additional material of D. (4.) elegantuln from Assam and Pahang.
rJourn. Fed. Malay States Mus. 14 (1928) 127.

qz,1Alerander:

Tipulid,a from Eastetn Asia, VII

527

HEXATOMINIDACTXLOLABISLONGICAUDAITEGASTYLATA

aubsp. nov.

Male.-Length, about 5 to 6 millimeters; wing, 6 to 7.
General coloration of the head and thorax light blue-gray.
Mesonotal prescutum with four narrow dark brown stripes, theintermediate pair not reaching the suture. Legs brownish
black. Wings strongly tinged with brown, the transverse elements and certain of the longitudinal veins evidently seamedwith brown, to produce a streaked appearance. Venation: Sc,
ending a short distance before the end of Rs. Abdomen darkgray, the hypopygium dark, including the styli. Male hypopygium with the outer dististyle unusually large, even exceedingin size that of typical Longicaul,a, in length about two-thirds
that of the basistyle.
(Kiushiu) .
Habitat.-Japan
Holotype, male, Kosugidani, Yakushima, altitude 2,500 feet,
April 29, 1929 (S. Issi.ki).
LIMNOPIIILA(DICRANOPHRAGMA)DORSOLINEATA

sp. nov.

General coloration of mesonotum obscure yellow, the regionof the lateral prescutal stripes bordered anteriorly by narrowdark brown lines; basal segments of antenne pale; pleuragrayish brown, lined longitudinally with dark brown; femora
yellow, more darkened outwardly, the tips abruptly paler; wings
strongly suffused with dusky, with a brown pattern that is confined to the vicinity of the veins; wings (male) of approximately
equal width throughout their length.
Mole.-Length, about 4.6 millimeters; wing, 5.2 to 5.5.
Rostrum and palpi dark brown. Antenna with the scapeand basal tlVo or three flagellar segments yellow, the remainder passing into black. Head gray, the anterior vertex moreocherous.
Mesonotal prescutum with the restricted ground color darkbrown, represented by hook-shapedareas surrounding the usual
lateral stripes, together with an isolated oval brown spot onlateral portion, sometimes attached to the remaining dark area;
scutum and scutellum ocherous brown; postnotum dark brown,
pruinose. Pleura grayish brown, with three dark brown longitudinal stripes, the most dorsal lying just below the pale dorsopleural region, extending from the cervical sclerites to thepostnotum; second stripe very short, lying between the anepisternum and sternopleurite; third dark area occupying the522

The Philippine Jotnnnt of ScienceLsso

ventral sternopleurite. Halteres yellow, the knobs weakly infuscated. Legs with the coxe brown; trochanters browaishyellow; femora yellow, slightly darkened outwardly, the tips
abruptly paler; tibia and tarsi obscureyellow, the terminal tarsal
segments darkened; legs long, conspicuously setiferous. Wings(Plate 1, fig. 13) strongly suffused with dusky, the dark wash
including all of the cells excepting the prearcular and costa!
regions; dark brown pattern almost as in formosa. Wing ofapproximately equal width throughout the c6ntral third of the
Iength. Venation: Vein 2d A ending just before to opposite
the level of the origin of Rs.
Abdomen dark brolrn, the basistyles of the male hypopygium
more reddish brotvn. Ilale hlpopygrum (plate Z, fig.26) with
the inner dististyle, i.d, very stout and broad-based, the apex
blunt and fleshy; no conspicuous group of seta in the axilla ofdististyles.
Habi,tat.-Formosa.
Holotype, male, Arisan, altitude 6,b00 to 8,000 feet, July Z,
1929 (S. Issi.ki). Paratopotype, male.
Li,mnophi,ln dorsol;i,neatois readily distinguished from L. (D)formosa Alexander (Formosa) by the diagnostic features listed
above.
LIUNOPIIILA(PRIONOLABIS)LIPOPELEPS

BD. noy.

General coloration gray; antenne black throughout; halteresyellow; wings tinged with gray, the prearcular region lightyellow; cell M, lacking; male hypopygium with the inner dististyle bifid at apex; gonapophysesappearing as slender sinuous
blackened rods that bear two groups of spines along the outermargin.
Male.-Length, about 5.5 millimeters; wing, 6.b.
Female.-Lengf,h, about 5 millimeters; wing, 6.
Rostrum pruinose; palpi black. Antennre black, the basal segment pruinose; flagellar segments oval, gradually decreasing insize outwardly. Head dark gray, the broad anterior vertex
lighter gray.
Mesonotal prescutum black, pruinose with gray; no tuberculate pits; pseudosultural fovee black. Pleura black, graypruinose, more heavily so on the sternopleurite. Halteres ofmoderate length, yellow, the knobs brighter. Legs with thecoxe and trochanters dark, the former pruinose; femora
brownish black, their bases obscure yellow; remainder of legs
black. Wings (Plate 1, fig. 14) tinged with gray; prearcular

42,1

Alerand,er: T'ipulid,a from Eastem, Asia, VII523

region light yellow; stigma and vague seams along cord andouter end of cell lst M, slightly darker, than the ground color;
Marcrotrichia of veins sparse, caudad of theveins pale brown.
main branch of R being nearly confined to those veins distad of
the cord. Venation: Sc, extending to opposite the fork of Rz+g+r
Sc, not far from its tip; R, faint, less than R1..2; ceil Mf
lacking; m-cu at near midleneth of cell 1st Mr.
Abdomen, including the hypopygium, blaek, pruinose. Male
hypopygium (Plate 2, fig. 27) with the caudal margin of thetergite, 9f, gently notched. Outer dististyle, od, gradually narrowed to a slender blackened point, just beyond midlength bearing a small lateral spine. Inner dististyle, id, wit};, the apex
bifid. Gonapophyses, gr, entirely blackened, each appearing asa slender sinuous rod that terminates in a long apical spine; onouter margin with two groups of smaller spines, one group
being subapical, the other nearer midlength of the apophysis;
the number of spines in both these groups variable. .?Edeagus,o, highly compressed, as in the subgenus.
Habitat.-Japan (Kiushiu) .
Holotype, male, Kosugidani, Yakushima, altitude 21500 feet'
April 29, 1929 (,S. /ssiki). Allotopotype, female. Paratopotypes, 5 males and females.
The present species and the closely allied Z. (P.) Iiponeura
sp. nov. are placed in the subgenus Prtonolabi's with some slightquestion. The general structure and basic plan of the malehypopygium are the same, but the gonapophyses are rather remarkable in form and depart notably from the otherwise conservative type of the subgenus.
LIITNOPEILA(PRIONOLABIS)LIPONEURA

ap. noY.

Male.-T'ength, about 5.3 millimeters; wing, 6.7.
Female.-I'ength, about 5.8 millimeters; wing, 6.4.
Closely allied to L. (P.) li'pophlepssp. nov., differing especiallyin the structure of the male hypopygium, notably of the gonapophyses.
Coloration of the body more intensely black. Legs darkercolored, brownish black to black, only the femoral bases restrictedly obscure yellow. Wings darker, slightly broader thanin lipophleps, the cells being correspondingly widened. Male
hypopygium (Plate 2, fig. 28) with the outer dististyle, ocf,
broader than in li,pophleps. fnner dististyle, id, with the basemore expanded, the terminal taillike extension simple at apex.
Gonapophyses, gr, of somewhat remarkable form, as figured.

524

1e3o

The Philippine Jourtwl of Science(Kiushiu) .
Habitat.-Japan
Holotype, male, Mount Kirishima, altitude 3,000 to 3,500 feet,
May 4, 1929 (S. Issi,ki). Allotopotype, female, altitude 2'500
feet, May 3, I9Z:9 (S. /ssiki).
LIMNOPHILA(TBICEOLIMNOPHILA)SATSUIIICOLAsI). noY.

Size small (wing not over 6.5 millimeters) ; general coloration
gray; cephalic portion of prascutum not polished; wings
brownish yellow, the base brighter, the stigma darker; malehypopygium with the lobes of the tergite relatively long andnarrow; not widened distally; outer dististyle and outer arm of
the inner dististyle narrow.
Mule.-Length, about 4.5 to 5 millimeters; wing, 5.7 to 5.8.
Female.-Length, about 7 millimeters; wing, 6.
Antenne dark brown, the basal segments,a little paler; flagellar segments long-oval. Head gray.
Mesonotum gray, the prescutum with poorly defined brown
stripes, the anterior portion not polished black, as in pi'l;ifer.
Pleura gray. Halteres yellow. Legs with the coxa and trochanters yellow; femora and tibia yellow, the tips narrowly
infuscated; tarsi brownish black. Wings tinged with brownish
yellow, the base brighter yellow, the oval stigrna darker; very
indistinct seams along cord and outer end of cell I-st Mr; veinsdark brown. Macrotrichia of cells confined to those beyond
cord, in the allotype greatly reduced in number. Venation: Cell
M, present, abnormally lacking in one wing of the type; innerends of cells Rn, Ru, and lst M,, in nearly transverse alignment.
Abdomen brownish gray, the hypopygium somewhat brighter.
Male hypopygrum with the sublateral lobes of the tergite relatively long and narrow, nearly parallel-sided, the tips obtusely
rounded, not expanded outwardly as in caesiella,;emargination
separating the lobes subcircular, about twice as wide as the diameter of one lobe. Outer dististyle and outer arm of the innerdististyle both narrow.
Habitat.-J apan (Kiushiu) .
Holotype, male, Shiroyama Hill, city of Kagoshima, Aprtl 27,7929 (S. lssfke). Allotopotype, female. Paratopotype, male.
Li,mnophi.la satsumicola is most closely allied to L. caesi'ella
Alexander, differing especially in the size, and in the structureof the male hypopygium, such as the relatively long and slenderlobes of the tergite.
LIDINOPEILA(TRICEOLIIINOPHILA)PLATYSTYLAPARALLELA

Eubsp. noY.

Quite as in typical platgstgln Alexander except for details of
structure of the male hypopygium. Median lobe of the tergite

42,4Alenand,er: Tipulid,a from Eastern Asin, VII

SZElong, parallel-sided, the apex entire, not expanded as in the
typical form. Outer arm of the inner dististyle slender, only
weakly dilated at tip.
Habi,tat.-Formosa.
Holotype, male, Arisan, altitude 6,500 to 8,000 feet, July Z,
1929 (S. /sszki). Allotopotype, female. Paratopotype, male;paratypes, 2 males, Rantaizan, altitude 4,000 to 6,000 feet, May
20, L928 (S. /sse&i).
ATABBA (ATARBODES) ISSIKIANA sp. noy.

General coloration yellow; antenne pale throughout; mesonotal prescutum yellow, the median stripe bordered anteriorly
by 4 n-shaped black mark; each scutal lobe with two black
markings; scutellum bordered posteriorly by black; femora
yellow, the tips not darkened.
Male.-Length, about 5 to 5.3 millimeters; wing, G to 7.
Female.-Length, about G.b to Z millimeters; wing, 7 to 7.5.
Rostrum pale; palpi pale basally, the outer two segments
darkened. Antenne pale throughout, those of the female longer
and stouter than those of the male; flagellar segments of maleelongate-cylindrical, with long conspicuous verticils; of female
with shorter verticils. Head chiefly ocherous yellow.
Mesonotum yellow, the median region of the prescutum
bordered anteriorly by a.-, -shaped black mark, each scutal lobe
with two brownish black or black marks, the more lateral one
largest; scutellum yellow, bordered posteriorly by black, in somecases so extensively as to include almost the whole sclerite;
postnotum pale, more infuscated and weakly pruinose behind.
Pleura pale yellow. Halteres yellow. Legs yellow, the femoral
tips not darkened; terminal tarsal segments infuscated. Wings(Plate 1, fig. 15) yellow, the veins darker yellow. Macrotrichia
of veins relatively numerous. Venation: Sc of moderate length,
extending about to midlength of Rs, Sc, some distance from its
tip and shortly beyond origin of Rs; veins issuing from cell
1st M, divergent; m-cu beyond the form of M.
Abdomen, including the hypopygium, yellow, the extreme eaudal margins of the tergites infuscated; in the female the abdomen is more uniformly darkened.
Habitat.-Formosa.
Holofupe, male, Arisan, altitude ?,800 feet, July ?, 1g2g (S.
Issiki).
Allotopotype, female. Paratopotypes, 4 males andfemales, altitucle 6,500 to 8,000 feet, July ?, lg2g (S. Issi,ki\ ;paratypes, 1 female, Rantaizan, altitude G,000 to ?,000 feet,

526

The Plui,kppine Journal of Sci,ence:.r,soMay 21, 1928; 1 female, 7,000 feet, June 2, lg27 ; 1 female,
Sankakuho, altitude 7,000 to 8,000 feet, May 25,1923 (S. Issi,ki,).
A paratype from Rantaizan has the extreme tips of the femora
darkened but from the peculiar thoracic pattern would appear
to belong here.
The known species of Atarba in Japan and Formosa form agroup of closely allied forms that are separable chiefly on coloration. It is worthy of note that the antenna of the female ofthese species are distinctly longer and stouter than those of themale sex. The described species from this region may be separated as follows:
1. Thoracic dorsum yellow, variegated with black....
.-._-.._--...
Z.
Thoracic dorsum uniformly yellow.-.......-_4.
-_--_.."._-..--.-.-..-".-_-.
2. Femoral tips narrowly blackenedA. paltid.i.comis Edwards.
Femoral tips not darkened._-.._......-..--.._....--.-.
B.
3. Antenne dark brown or black throughout."..-..-__--A. fuseicornis Edwards.
Antenne pale throughout ------.....-....
-.--.-...A. issikia,no sp. nov.
4. General color of the mesonotum polished yellow; wings deep yellow.
A. IeptonantDo Alexander.
General color of mesonotum opaque testaceous yellow; wings pale yellow.
A' mirwticornis sp' nov'
ATARBA (ATARB'DE') lrrNuTrco*Nrs gp. noy.

Male.-Length, about 5 millimeters; \iling, 5.8.
Closely allied to A. (A.) leptonantha Alexander (Formosa),
differing especially in the shorter antenna and very pale yellow
coloration of the body and wings.
Antenne (male) small, if bent backward ending some distance
before the wing root; flag:ellar segments elongate-cylindrical,
the verticils slightly exceeding the segments. Head and thorax
dull testaceous yellow, not nitidous yellow as in leptoxantha;vague indications of a darker coloration surrounding the regionof the lateral prrescutal stripes. Legs broken beyond the trochanters. Wings paler yellow than in leptorontha.
Habi.tat.-Japan (Kiushiu) .
Holotype, male, Kosugidani, Yakushima, altitude 2,500 feet,
April 29, 1929 (S. Issi,ki).
ELEPHANTOMYIA(ELEPEANTOUYIA)DIETZIANA

sp. noy.

General coloration obscure yellow, the prescutum with a
median darkening in front; head gray; halteres pale yellow;
pleura more or less darkened; wings tinged with yellow, theoval stigma pale brown; abdominal segments bicolorous, theirbases obscure yellow, the apices more broadly blackened; malehypopygium with the gonapophysesbeyond midlength spinose.

42,+

Alesand,er: Ti,puliil,afrom Eastem Asia, VIIEZ|

Male.-Length, excluding rostrum, about 9 to 10 millimeters;
wing, 8,2 to 8.5; rostrum, about 7 to 7.5.
Femnle.-Length, excluding rostrum, about 7,5 to 8 millimeters; wing, 7.5 to 8; rostrum, about 7.
Rostrum black, elongate. Antennre brown, the scapal segments more pruinose; flagellar segmentsgradually increasing
in length and decreasingin diameter outwardly, with long conspicuous verticils that exceedthe segments; terminal segment
very small. Head gray, that postgene paler.
Pronotum dark brown, obscureyellow laterally. Mesonotum
obscurebrownish yellow, the prascutum conspicuouslydarkened
medially, this area usually becomingobsoletebefore the suture;
one specimen,a paratype male, has the entire thorax blackened,sparsely pruinose. Pleura more or less dark brownish gray,especiallyon the sternopleurite, in some specimensmore extensive to include almost the entire pleura. Halteres pale yellow.
Legs with the coxe and trochanters yellow; femora yellow basally, passing into brown at tips; tibie and tarsi yellowish
brown, the latter paler yellow on outer segments. Wings tinged
with yellow,the long oval stigma pale brown; veins pale brown,
the basalveins more luteous. Venation: Sc, ending oppositeor
just before the fork of Rs, Sc" closeto its tip; branchesof Rs
running approximately parallel to one another, cell R, at margin being much wider than cell Rr; anterior branch of Rs moreor less sinuousbeneaththe stigma; m-cu at or before midlengthof cell lst M,.
Abdomen bicolorous, the segments obscure yellow basally,brownish black apically, in the males the dark color becoming
more extensiveon the outer segments,which are entirely blackened; hypopygium with the basistyles pale. Male hypopygrum
with the gonapophysesprovided with a group of from ten to
twelve spines beyond midlength, beyond which point each apophysis is producedinto a long pale point.
H abi,tat.-Japan (Kiushiu) .
Holotype, male, Kirishima, altitude 2,500 feet, May 3, 192g
(S. /ssri/ci). Allotopotype, female. Paratopotypes,5 males andfernales.
Someyears ago, Dr. William G. Dietz sent me a brief diagnosis of an ElephantomAiapreservedin his collection, received
from Honshiu, Japan. I believe this representsthe present
species,and it is fitting to dedicate this distinct speciesof fly
to this distinguished student of the Tipulide. The spinous

528

Tlrc Pluil;tppi,neJornnal, of Sci,ence

L$ogonapopyses and marked prescutum separate the fly from thelarger and more-northern E. (8.) hokkaidenscs Alexander. The
somewhat similar E. (8.) serotina Alexander (Formosa) is distinguished by the coloration of the wings and body.
ERIOPTERINIGONOUYIA(PROGONOIIYIA)PEBTURBATA

sp. nov.

General coloration black, more or less pruinose; pleura black,variegated with obscure yellow; knobs of halteres darkened;legs black; wings tinged with dusky; male hypopygium withthree dististyles.
Male.-Length, about 4.5 millimeters; wing, 5.3.
Rostrum dark brown; palpi black. Antenna black throughout; flagellar segments decreasing in length and diameter outwardly, the verticils exceeding the segments in length. Headblack, sparsely dusted with gray.
Mesonotal prescutum black, very sparsely dusted with gray,the anterior lateral pretergites and humeral region restrictedly
obscure yellow; scutellum dusky at base, the posterior half lightyellow; postnotum dark gray. Pleura black, variegated with
obscure yellow, including the suture between the anepisternum
and sternopleurite, the meron and adjoining sclerites, and thedorsal portion of the pleurotergite. Halteres dirty white, theknobs infuscated. Legs with the coxe darkened on outer faces;
trochanters brownish yellow; remainder of legs black, the femoral bases scarcely brightened. Wings (Plate 1, fig. 16) witha weak dusky tinge, the stigmal region only restrictedly andvaguely darkened; veins bron'nish black. Venation: Sc, ending
just before or opposite the fork of Rs, Sc" some distance fromits tip, the distance between origin of Rs and Sc, about equal
to m-cu; cell R, deep; m-cu at fork of M.
Abdomen brownish black, including the hypopygium. Male
hypopygium (Pate 2, fi5. 29) with the outer lateral angle ofthe basistyle, b, produced into a stout setiferous lobe. Three
dististyles, iI; the outermost a glabrous rod that is nearly straight,
a little expanded at apex, at base with a consp'icuous rounded
lbbe; second dististyle a broad triangular blade that narrows toan acute apical beak; margin of style with a comb of nine orten straight yellow pegs; surface of this style with numerous
sete; inner, or third, dististyle bifid at apex into a long spine
and a more-flattened blade, the stem, with abundant long sete.
-Edeagus, p, flattened, terminating in a short curved spine,
surface of disk of adeagus with several long slender sete.

42,4

Alenand,er:Tipukda from Eastern Asia, VII

5Zg

Habitat.-China (Szechwan).
Holotype, male, Mount Omei, altitude 4,500feet, July 15, 1g2g
(ex Parish).
I cannot identify this fly with the Indian G. (P.) ruigri,pes(Brunetti), which has as synonymsG. ni,gra Brunetti, G. i,ncompletaBrunetti, and possibly G. gracili,sBrunetti. It superficiallyresemblesG, (P.) alboscutellnfoAlexander (Formosa), but the
male hypopygiumis so different that someof the parts are notreadily homologized. It is probablethat this specieswill befound not to be a Progonomgia in the strict interpretation of
the subgenus.
GONOMYIA(GONOMYIA)OMEIENSIS

rp. nov.

General coloration gray; antenna black throughout; scutellum
obscure yellow; pleura yellow, striped longitudinally with bluishgray, most evident on the anterior mesopleura; wings gray, thestigma more brownish; male hypopygium with the inner dististyle a bispinous flattened blade; phallosome asymmetrical,
bearing a blackened, spearlike, lateral arm.
Male.-Length, about 4.4 millimeters; wing, b.
Female.-Length, about 5 millimeters; wing, 5.4.
Rostrum orange; palpi black; in female, rostrum entirely
black. Antenna black throughout, the basal segments of flagellum large, the remainder becoming linear. Head dark gray.
Pronotum fulvous, darker laterally. Anterior lateral pretergites obscure yellow. Mesonotal prescutum brownish gray, thehumeral and lateral regions obscure yellow; scutum brownish
gray; scutellum obscure yellow; postnotum gray. Pleura paleyellow with two narrow blue-gray longitudinal stripes that are
most evident on the anepisternum and ventral sternopleurite,
the pteropleurite entirely pale. Halteres relatively elongate,
pale, the knobs infuscated. Legs with the coxe pale, the outerfaces of the fore and hind coxaeslightly infuscated; trochanterspale brown; remainder of legs brownish black, passing into
black. Wing:s (Plate 1, fig. 17) gxay, the stigma more brownish;veins brownish black. Venation: Sc, extending to shortly beyond the origin of Rs, Sc, a little removed from its tip; Rssinuous on basal half ; cell lst M" closed; m-cu elose to forkof M.
Abdominal tergites dark brown, the sternites paler, theextreme caudal margins of the outer segments narrowly pale.
Male hypopygium (Plate 2, fiq.30) with the apex of basistyle,b, produced into a small oval setiferous lobe. Outer dististyle,262390-6

530

The Philippine lotnna,I of Science$Bood, a pale flattened blade that is setiferous on outer margin, theinner margin glabrous. fnner dististyle, fd, a bispinous flattened blade, the outer spine longest, bearing a single powerfulseta near base; smaller spine more curved; apex of style bearing
two powerful fasciculate bristles, in addition to about four or fivesmaller sete. Phallosome, p, asymmetrical, a pale flattened
blade, at near midlength bearing a black spear-shaped arm on
one side only.
Habitat.-China( Szechwan).
Holotype, male, Mount Omei, altitude 4,b00 feet, August 14,
1929 (ex Parish). Allotopotype, female, August 7, J:g?;g.
Gonomyia ometensis is allied to G. affinis Brunetti (Hima_layas), differing in the striped pleura and details of venation.
I do not have a male of affi,nisfor comparison.
GONOMYIA(LIPOPELEPS)QUADRIFILA

ap. noy.

General coloration brownish gray; head yellow, the centerof the vertex with a small dark spot; pleura dark, striped withyellow; femora pale, with a subterminal dark ring; wings
dusky, the costal region yellowish white, the disk variegated withpale spots and flecks; sc short; anterior branch of Rs short andoblique; male hypopygium with three dististyles, two being very
Iong and filiform.
Male.-Length, about B millimeters; wing, 3.b.
Rostrum and palpi black. Antenna with the basal segments
obscure yellow, the slender outer segments of the flagellum
passing into black. Head light yellow with a small dark
spot
on the disk of vertex.
Pronotum and anterior lateral pretergites light yellow. Mesonotum brownish g'ray, the caudal margin of the scutellum a
trifle paler; postnotal mediotergite extensi'ely pale, darker
posteriorly. Pleura black, pruinose, the pleurotergite pale
brownish yellow; a narrow but conspicuous light yellow longitudinal stripq extending from the fore coxe, passing above theremaining coxe to the base of the abdomen. Halteres broken.
Legs with the coxe chiefly pale, especially the fore coxe, theothers narrowly darkened at base; trochanters yellowish testaceous; femora brownish Vellow, with a narrow dark brown
subterminal ring, the extreme tip again narrowly pale; tibie
and basitarsi yellow, the tips darkened; terminal tarsat segments
infuscated. Wings (Plate 1, fig. 18) with a strong brown suffusion, the costal region yellowish white, more extensive
beforeand beyond the stigma; stigma oval, brown; conspicuousgrayish

42,1Al,eranf,er: Ti,pulid,a from Eastent, Asia, VII

531

white spots and flecks on the membrane, the most evident appearing as an incomplete crossband before the cord; remaining
cells of wing spiinkled with pale; veins dark, the costal veins
pale. Venation: Sc short, Sc, ending a distance before theorigin of Rs about equal to the total length of the latter; anterior
branch of Rs unusually short and oblique, the cell at margin
being very wide; m-cu at or just before the fork of M.
Abdominal segmentsbrownish black, narrowly ring:ed caudallywith yellow; hypopygium dark. Male hypopygium (Plate 2,
fig. 31) with three dististyles, d, two very long and filiform;
longest style appearing as a very long curved rod that ends in anacute blackened point; second style more than one-half the lengthof the first, dusky, gently arcuated, the tip ob'liquely obtuse;
third dististyle, or braneh, small and fleshy. Phallosome, p,large, pale, terminating in two expanded subtriangular dusky
blades that lie parallel to one another.
Habitat.-Japan (Kiushiu) .
Holotype, male, Yasakagawa, Oita-ken, May 10, 1929 (7.

uve).
Gonomuin quad,rifi,ln is most closely related to G. fl,auocostalis
Alexander (Japan), differing very conspicuouslyin the structureof the male hypopygrum.
EBIOPTERA(ERIOPIERA)

LUTEICORNIS

Ep. noy.

General coloration pale yellow; palpi black; basal segments ofantenna pale yellow; head pale; pleura yellow with a relatively narrow dusky longitudinal stripe; knobs of halteresblackened; wings pale yellow, the veins yellow, those of theanterior cord slightly infuscated; male hypopygium with thedististyle single but profoundly divided, the two principle arms
bearing two unequal black spines in the axil.
Male.-Length, about 3.8 to 4 millimeters; wing, 4.2 to 4.5.
Female.-Length, about 4.5 to 5 millimeters; wing, 5.5 to 6.
Rostrum pale yellow; palpi black. Antennre with the scapeand basal six or seven flagellar segments pale yellow, the outer
segments infuscated. Head whitish, the center of the vertex
more suffused with yellow.
Pronotum yellowish white, infuscated medially. Mesonotal
prrescutum light reddish brown, more infuscated medially, thelateral margins paling to yellowish white; pseudosutural fovee
and tuberculate pits reddish brown; scutal lobes light reddish
brown; scutellum chiefly pale; postnotal mediotergite reddish
brown, more whitish laterally, more darkened behind. Pleura

532

The Philippine Jornnnl of Sciencelesoyellow, with a relatively narrow dusky longitudinal stripe, extending from the fore coxe to the posterior portion of thepostnotum, the pleura dorsad of this stripe very pale, yellowish
white, the ventral sclerites somewhat deeper yellow. Halterespale, the knobs blackened. Legs with the fore coxe darkened;remaining coxe and all trochanters yellow; remainder of legs
yellow, the outer tarsal segments infuscated. Wings paleyellow, the veins dark yellow; a scarcely evident darkening involving the veins of the anterior cord. Venation: Vein 2d A
unusually long and sinuous, at near midlength a little closer tothe anal margin.
Abdomen pale brownish yellow, the lateral margins darker,
the caudal margins pale. Male hypopygium (Plate 2, fig. 32'ywith the dististyle, d, apparently simple but profoundly branched;outer arm a simple straight rod, the tip acute; inner arm anarcuated flattened blade, provided with marginal seta; twoblackened spines in the axil, one being very small. Gonapophyses, g, appearing as flattened pale blades, the acutetips narrowly blackened, the outer margin microscopicallyroughened.
Habita,t.-China (Szechwan).
Holotype, male, Mount Omei, altitude 4,500 feet, August 14,
L929 (ex Parish). Allotopotype, female. Paratopotypes, bmales and females, July 31 to August 22, 1929.
Erinptera Luteicorni,s is evidently allied to the Nearetic .8.
septemtrionis Osten Sacken.
ERIOPTERA (ERIOPTERA) HIMALAY.aE sp, noy.

Male.-Length, about 4.5 millimeters; wing, 4.5.
Characters generally as in E. subti,ncta Brunetti, differing
especially in the str,ucture of the male hypopygium.
Wings pale yellow, the veins and macrotrichia darker yellow;
a faint but distinct clouding on the anterior cord. Venation:
Vein 2d A very strongly sinuous, on its distal third extending
parallel to the margin or nearly so. Male hypopyglum (Plate2, fig. 33) with the outer dististyle, od, a simple flattened rod,
gently narrowed and curved to the acute tip, the outer facemicroscopically roughened. Inner dististyle, id, a larger,
broadly triangular structure, the outer apical angle produced
into a long slender spine, the inner margin microscopically serrulate; inner mesal region of the trianglq extended into a dusky
flange, weakly and irregularly tridentate on margin. Gonapo-

42,1

Alenand,er: Tipul;ida from Eastern Asia, VII533

physes, g, appearing as straight slender rods, pale yellow, thedusky tips obtuse.
Habi,tat.-India.
Holotype, male, Ghumti, Darjiling District, Eastern Himalayas, altitude 4,000 feet, July 1911 (f'. H. Graaelg).
I received this species in excha-nge with Mr. Brunetti as
being E. (8.) subtincta Brunetti and considered the determination as being correct until Edwards n published further notes on
the type of subti,ncta. From this it is very evident that thepresent species has nothing in common with subti'ncta nor can
I identify it vrith any of the described reg:ional species. The
structure of the male hypopygtum is distinctive.
MOLOPHILUS

ABICOLA

ap. nov.

General coloration brown to reddish brown; antenne relatively
short in both sexes; legs dark brown; halteres yellow; wings
yellowish g:ray, the costal region clearer yellow; abdomen black'the hypopygium reddish brown; lobes of the basistyle not produced into spinous points; surface of inner dististyle glabrous.
Male.-Length, about 3 to 3.2 millimeters; wing, 3.8 to 4.
Female.-Length, about 3.5 to 3.7 millimeters; wing, 4.8 to 5.
Rostrum and palpi black. Antennre with the scape and basal
segments of flagellum yellow, the remaining flagellar segments
passing into black; antenne relatively short, when compared
witln pegastzs, with long conspicuous verticils, the outer series
longer. Head gray.
Mesonotal prrescutum reddish brown to brown, with a faintgray pruinosity; lateral pretergites and narrow lateral marginsof the prescutum pale yellow; posterior sclerites of mesonotum
more grayish. Pleura dark brown. Halteres pale yellow'
Legs with the eoxe and trochanters dark yellow; remainderof legs dark brown, the femoral bases slightly paler. Wings
yellowish gray, the prearcular and costal regions clearer yellow;
veins darker than the ground color; macrotrichia brown, thecostal trichia light golden yellow. Venation: Vein 2d A ending
about opposite one-third the length of the petiole of cell M,'
Abdomen black, the hypopygium conspicuously reddish brown'
Male hypopygrum with the apical lobes of the basistyle not at
all chitinized. outer dististyle a slender rod, the apex slightly
expanded into an oval head that is provided with a few darkened'Rec. Indian Mus. 26(1924) 300.

534

The Phi,Ii,ppi,neJournnl of Sciencetubercles. Inner dististyle glabrous, except for microscopic
appressedspinula at apex and along outer margin.
Habitat.-Formosa.
Holotype, male, Arisan, altitude 6,500 to 8,000 feet, July 7,
L929 (S. Issiki). Allotopotype, female. Paratopotypes, 2males and females.
Molophi.lusaricola differs from M. ni,gr.r,pes
Edwards, which
has a somewhat similar hypopygium, by the short antenna ofboth sexes. It is even more similar to the JapaneseM. pegasus
Alexander, differing in details of coloration, the shorter antenneof the male, and slight details of the male hypopygrum.

ILLUSTRATIONS
dististyle; g, gonapophyrfu;[Legend: o, edeagus; D, basistyle;4id, inlter dietigtyte;outer dististyle ; p, phallmome ; s, stemit€ ; t, t€rgite
ventral dististyle.l"d,Pr,.lrs
FIG.

o4

1

L. Umonin (Dicranom,gial ka,nsunnsis sp. nov., venetion.
2, Limonia (Litnanie') gakushimtnsis sp. nov., venation.
3. Li,monin (Limonin'1 unieinctifera sp. nov,, venation.
4. Heliu,s (HeXinnl costofimbriat?ra sp. nov., venation.
5. Heliu.s (IIeli,u,s) pallid,issimas sp. nov., venation.
6, Antocha (Antochal subconfluenta sp. nov., venation.
7. Orimarga cruciforrrui,s sp. nov., venation.
8. Ori,ma,rga gakwhi.mana sp. nov., venation.
9. Orimarga omph.a sp. nov., venation.
10. Ori,ma,rga seticoeta sp. nov., venation.
LI-. Dicrunota (Amalopina) gibbera korafutonis Bubsp. nov., venation.
(Amalopina'1 gibbera gibbera Alexander, venation.
12. Dicrarcta
13. Limnophilo (Di.craruphragmal
d,orsolineoto ap. nov,, venation.
14. ILimnophiln (Prienolnbis) lipoph)eps sp. nov., venation.
L5. Atarbo (Atarboiles') beikiann, sp. noy., venation.
76. Gonnm,gin, (Progonam,yia,) porturbata
sp. nov., venation.
L7. Gonomvia (Gorcmyial omp,i,ensissp. nov., venation.
18. Gonomgia, (Lipophlepsl qw,ilrif,ln sp. nov., venation.
PLAm 2

FIG. 19. Tipu,la latil;i.gula sp. nov., male hypopygium, lateral view.
20. Tipula latilti,gula sp. nov., male hypopygium, ninth tergite.
21, Tiptila lati.ligula sp. nov., male hypopygium, eighth sternite.
22. Li,nwnie (Dicranomyia) Icansunnsis sp, nov., male hypopygium.
23. Li.mmai.a (L&moni,a) galcuslvi.menstssp. nov., male hypopygium.
24. Limonin (Limonin) unieinctifera sp. nov., male hypopygium.
25, Antochn (Antochal subconfl,u.enta,sp. nov., male hypopygium.
26. Lfunnophila, (Dicra,nophragma,) d,orsol'ineata sp. nov., male hypopygtum.
27. Limnophila (Prinnolabis\ Ii,pophleps sp. nov., male hypopygium.
28. Ltmnopltila (Pri,onolabis\ Iiponnwa sp. nov., male hypopygium.
29. Gonom,yia, (Progonoruyia,) perttnbata sp. nov., male hypopygium.
30. Gonom,yia (GonoWia)
omei,ensis sp. nov., male hypopygium.
31. Gonom.gin (LipophXeps) qlnilri.fila sp. nov., male hypopygium.
32, Eri,optera (Erioptera)luteicom:is sp. nov., male hypopygium.
(Eriopteral
hima,Iage sp. nov., male hypopygium.
33. Erioptera
535

ALEXANDER:TrpuLrDA FRoM EasrERN ASIA, VII.I

PLATE 1.

[PErLrP. JounN. Scr.,42, No.4.

ALDXANDER:TrpuLrDA FnoM EASTERNAsrA, VII.I

/l

/,s
20

lPHrLrP. JouRN. ScI.,42, No. 4.