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෗┿㸯 ᳃ᯘື≀◊✲ࢭࣥࢱ࣮᧜ᙳ  ᖺ  ᭶  ᪥ ୹Ἴᕷ㟷ᇉ⏫ἑ㔝᳃ᯘື≀◊✲ࢭࣥࢱ࣮ࡢᩜᆅෆ࡟ฟἐࡋࡓ࢖ࣀࢩࢩ
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෗┿㸰 ᳃ᯘື≀◊✲ࢭࣥࢱ࣮᧜ᙳ  ᖺ  ᭶  ᪥ ⚄ᡞᕷℿ༊ᕷ⾤ᆅࡢࢦ࣑ࢫࢸ࣮ࢩࣙࣥ࡟࠾࠸࡚ࠊኪ㛫࡟ฟἐࡋࡓಶయ
ཱྀ⤮㸯  රᗜ┴ෆ࡛ほᐹࡉࢀࡓ࢖ࣀࢩࢩ

෗┿㸱 ᫛࿴ึᮇࡢ໭ᦤᆅᇦ࡟࠾ࡅࡿ࢖ࣀࢩࢩ⊟ࡢᵝᏊ㸦෗┿ᥦ౪㸸ᯘ ┿⌮Ặ㸧෗┿㸲 ᫛࿴  ᖺ௦࡟࠾ࡅࡿභ⏥ᒣ࢖ࣀࢩࢩࡢ㥑㝖ࡢᵝᏊ㸦෗┿ᥦ౪㸸Ọᓥ᫛஧㑻Ặ㸧ཱྀ⤮   රᗜ┴ෆ࡟࠾ࡅࡿ᫛࿴ࡢ⊁⊟࡜㥑㝖ࡢᵝᏊ

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ཱྀ⤮㸱  ࢖ࣀࢩࢩࡢ⏕ᜥᐦᗘࡢ⤒ᖺኚ໬ࡣࡌࡵ࡟
රᗜ┴᳃ᯘື≀◊✲ࢭࣥࢱ࣮ࡣࠊ࣡࢖ࣝࢻࣛ࢖ࣇ࣐ࢿࢪ࣓ࣥࢺ࡟ಀࢃࡿ◊✲ᡂᯝࢆᗈࡃᕷẸࡢ᪉ࠎ࡟▱ࡗ࡚࠸ࡓࡔࡃࡇ࡜ࢆ┠ⓗ࡜ࡋ࡚ࠊᖹᡂ  ᖺᗘ࠿ࡽẖᖺࠕරᗜ࣡࢖ࣝ
ࢻࣛ࢖ࣇࣔࣀࢢࣛࣇࠖࢆห⾜ࡋ࡚࠸ࡲࡍࠋ
ᮏᖺᗘࡣࠕරᗜ┴࡟࠾ࡅࡿ࢖ࣀࢩࢩࡢ≉ᐃ㫽⋇ಖㆤ⟶⌮ィ⏬ࡢ⌧≧࡜ㄢ㢟ࠖࢆห⾜࠸ࡓࡋࡲࡍࠋ
ከࡃࡢேࡧ࡜ࡀㄆ㆑ࡋ࡚࠸ࡿࡼ࠺࡟ࠊරᗜ┴ࡢࡳ࡞ࡽࡎ᪥ᮏྛᆅ࡟࠾࠸࡚ࠊ࢖ࣀࢩࢩ
࡟ࡼࡿ㎰ᮧ㒊࡟࠾ࡅࡿ㎰ᴗ⿕ᐖࡸࠊ㒔ᕷ㒊࡟࠾ࡅࡿ⏕ά⿕ᐖࡣ┳㐣࡛ࡁ࡞࠸≧ἣ࡜࡞ࡗ࡚࠾ࡾࡲࡍࠋࢃࡓࡋࡓࡕ࣡࢖ࣝࢻࣛ࢖ࣇ࣐ࢿࢪ࣓ࣥࢺ࡟ᚑ஦ࡋ࡚࠸ࡿ⪅࡟࡜ࡗ࡚ࠊ࢖ࣀ
ࢩࢩ࡟ࡼࡿ⿕ᐖࢆ㍍ῶࡉࡏࡘࡘࠊࡑࡢᣢ⥆ⓗά⏝ࢆ᥎㐍ࡍࡿࡇ࡜ࡣࠊ᪥ᮏ࡟࠾ࡅࡿ᭱㔜せㄢ㢟ࡢࡦ࡜ࡘ࡜࡞ࡗ࡚࠸ࡲࡍࠋ
⛉Ꮫࡢᙺ๭ࡢࡦ࡜ࡘࡣࠊᗈ⠊࡞ᆅᇦ࡬ࡢ㐺⏝ྍ⬟ᛶࡢ㧗࠸▱㆑య⣔ࢆᵓ⠏ࡍࡿࡇ࡜࡛
ࡍࡀࠊᆅᇦ࡟㈉⊩ࡍࡿ▱㆑య⣔ࢆᵓ⠏ࡍࡿࡇ࡜ࡶ㔜せ࡞ᙺ๭ࡢࡦ࡜ࡘ࡛ࡍࠋ࢖ࣀࢩࢩ࡟
ࡼࡿ⿕ᐖࡣከᵝ࡛ࠊᆅᇦᕪࡀ኱ࡁ࠸ࡓࡵࠊࡲࡎࠕරᗜ┴ෆ࡟࠾ࡅࡿࢽ࣍ࣥ࢖ࣀࢩࢩࡢ⏕ᜥศᕸ࡜㎰ᴗ⿕ᐖࡢ㛵ಀࠖࢆ᫂ࡽ࠿࡟ࡍࡿ࡜ྠ᫬࡟ࠊ㒔ᕷ㒊࡟㏆᥋ࡋࡓࠕභ⏥ᒣ࢖ࣀࢩࢩࡢ⿕ᐖࡢᐇែ࡜ฟἐせᅉࠖࡸࠕභ⏥ᒣ⣔࡟࠾ࡅࡿᤕ⋓ಶయࡢ≉ᚩࡢᐇែࠖࢆ᫂ࡽ࠿࡟
ࡋࠊࡑࢀࡽࡢ⤖ᯝ࡟ᇶ࡙࠸࡚ࠕරᗜ┴࡟࠾ࡅࡿࢽ࣍ࣥ࢖ࣀࢩࢩࡢಶయᩘ᥎ᐃࠖࢆ࠾ࡇ࡞
࠺࡜ྠ᫬࡟ࠊࠕරᗜ┴࡟࠾ࡅࡿຍᐖಶయࡢ⾜ື≉ᛶࠖ࡟ࡘ࠸࡚ࡶ✚ᴟⓗ࡟ㄪᰝࡋࠊࡑࡢ
ᡂᯝࢆᮏࣔࣀࢢࣛࣇ࡟ࡲ࡜ࡵࡲࡋࡓࠋ
࡞࠾ࠊ࢖ࣀࢩࢩࡣࠊ໭ࡣᮾ໭ᆅ᪉࠿ࡽ༡ࡣす⾲ᓥࡲ࡛ᗈࡃ⏕ᜥࡋ࡚࠾ࡾࡲࡍࡀࠊᆅᇦ
࡟ࡼࡗ࡚⦾Ṫ≉ᛶࡀ␗࡞ࡾࡲࡍࡢ࡛ࠊᮏࣔࣀࢢࣛࣇ࡛ࡣࠕරᗜ┴࡟࠾ࡅࡿࢽ࣍ࣥ࢖ࣀࢩ
ࢩࡢ⦾Ṫ≉ᛶࠖࢆ᫂ࡽ࠿࡟ࡋࡲࡋࡓࠋࡇ࠺ࡋࡓරᗜ┴࡟࠾ࡅࡿ࢖ࣀࢩࢩࡢ≉ᛶ࡟㛵ࡍࡿㄪᰝ⤖ᯝࡣࠊරᗜ┴ࡢࡳ࡞ࡽࡎ௚ࡢᆅᇦ࡟㐺⏝ࡍࡿࡇ࡜ࡀྍ⬟࡛ࡍࠋࡉࡽ࡟ࠕᖺ㱋ᰝᐃ᪉ἲࠖࠊࡸࠕዷፎ⋡⟬ฟ᪉ἲࠖࡢ㛤Ⓨࡸࠕ࢘࢖ࣝࢫ࣭ᐤ⏕⹸ࡢឤᰁ≧ἣࠖࡢㄪᰝ⤖ᯝࡣࠊࢽ࣍ࣥ࢖ࣀࢩࢩ඲య࡟㐺⏝ྍ⬟࡞ㄪᰝ⤖ᯝ࡛ࡍࠋ
ࡇࡢࡼ࠺࡟ᮏࣔࣀࢢࣛࣇࡣࠊරᗜ┴࡛ᚓࡽࢀࡓ◊✲ᡂᯝࢆࡼࡾᗈᇦࡢᆅᇦ࡟㐺⏝ࡍࡿࡇ࡜ࡀྍ⬟࡛ࡍࠋࡍ࡞ࢃࡕࠊරᗜ┴࡛ᚓࡽࢀࡓㄪᰝ⤖ᯝࡸ㛤Ⓨᡭἲࡣࠊᆅᇦ≉ᛶࢆຍ࿡ࡍࡿ࡞ࡽࡤࠊ᪥ᮏ඲య࡛㐺⏝࡛ࡁࡿࡇ࡜ࡀ࡛ࡁࡲࡍࠋ
᭱ᚋ࡟࡞ࡾࡲࡋࡓࡀࠊ
ࠕරᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇࠖࡣࠊ⦅㞟ጤဨ఍ࡀẖᖺタᐃࡍࡿࢸ࣮࣐࡟ἢࡗ࡚ᇳ➹ࡉࢀࡓㄽᩥ➼ࢆࣔࣀࢢࣛࣇ࡜ࡋ࡚⦅㞟ࡋ࡚࠾ࡾࡲࡍࠋヲ⣽࡟ࡘࡁࡲࡋ࡚ࡣࠊᢞ✏つᐃࢆࡈཧ↷ࡃࡔࡉ࠸ࠋࡳ࡞ࡉࡲࡢࡈᢞ✏ࢆ࠾ᚅࡕࡋ࡚࠾ࡾࡲࡍࠋ
⦅㞟ጤဨ㛗 ᯘⰋ༤┠  ḟ➨ 㸯 ❶  රᗜ┴࡟࠾ࡅࡿࢽ࣍ࣥ࢖ࣀࢩࢩࡢಖㆤ⟶⌮ࡢ⌧≧࡜ㄢ㢟        ᶓᒣ┿ᘪ➨ 㸰 ❶  රᗜ┴࡟࠾ࡅࡿࢽ࣍ࣥ࢖ࣀࢩࢩࡢ⏕ᜥศᕸ࡜㎰ᴗ⿕ᐖࡢ㛵ಀᯇᮏ ᓫ࣭ᆏ⏣ᏹᚿ࣭ᓊᮏᗣ㄃➨ 㸱 ❶  රᗜ┴࡟࠾ࡅࡿࢽ࣍ࣥ࢖ࣀࢩࢩࡢಶయ⩌ືែࡢ᥎ᐃᯇᮏ ᓫ࣭ᆏ⏣ᏹᚿ࣭ᓊᮏᗣ㄃➨ 㸲 ❶  ㎰ᆅ࡟㞄᥋ࡋ࡚⏕ᜥࡍࡿࢽ࣍ࣥ࢖ࣀࢩࢩࡢຍᐖ⾜ືࡢゎᯒ
ᶓᒣ┿ᘪ࣭Ụ⸨බಇ࣭ᮌୗ⿱⨾Ꮚ➨ 㸳 ❶  ࢽ࣍ࣥ࢖ࣀࢩࢩࡢᖺ㱋ᰝᐃ᪉ἲ
㎷ ▱㤶࣭ᶓᒣ┿ᘪ➨ 㸴 ❶  ࢽ࣍ࣥ࢖ࣀࢩࢩࡢዷፎ⋡⟬ฟ᪉ἲࡢ㛤Ⓨ
㎷ ▱㤶࣭ᶓᒣ┿ᘪ➨ 㸵 ❶  රᗜ┴࡟࠾ࡅࡿࢽ࣍ࣥ࢖ࣀࢩࢩࡢᇶᮏⓗ⦾Ṫ≉ᛶ
㎷ ▱㤶࣭ᶓᒣ┿ᘪ➨ 㸶 ❶  ࢽ࣍ࣥ࢖ࣀࢩࢩࡢ࢘࢖ࣝࢫឤᰁ⑕
⡿‶◊୕࣭᭹㒊ᚿಖ࣭㕥ᮌ⤢Ꮚ࣭὾ᓮ༓⳯⨾࣭ୗ⏣ᐂ࣭๓⏣೺➨ 㸷 ❶  ࢽ࣍ࣥ࢖ࣀࢩࢩࡢෆ㒊ᐤ⏕⹸బ⸨ ᏹ࣭ᡞ⏣ṇᯞ࣭M. A. Omar࣭㎷ ▱㤶࣭ᶓᒣ┿ᘪ➨ 㸯㸮 ❶ භ⏥ᒣ࢖ࣀࢩࢩၥ㢟ࡢ⌧≧࡜ㄢ㢟
㎷ ▱㤶࣭ᶓᒣ┿ᘪ➨ 㸯㸯 ❶ භ⏥ᒣ࢖ࣀࢩࢩࡢၥ㢟ಶయࡢ≉ᚩ
㎷ ▱㤶࣭ᶓᒣ┿ᘪරᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-1➨ 㸯 ❶            රᗜ┴࡟࠾ࡅࡿࢽ࣍ࣥ࢖ࣀࢩࢩࡢ
ಖㆤ⟶⌮ࡢ⌧≧࡜ㄢ㢟ᶓᒣ┿ᘪ 1,2

せ Ⅼ࣭ රᗜ┴࡟࠾ࡅࡿே࡜ࢽ࣍ࣥ࢖ࣀࢩࢩࡢ㛵ࢃࡾࡢṔྐࢆᴫほࡋࡓࠋỤᡞ᫬௦ࡣ῝้࡞㎰ᴗ⿕ᐖࡢࡓࡵࠊ⊦ᇉࡀྛᆅ࡛タ⨨ࡉࢀࡓࠋ᫂἞௨㝆ࡣࠊᤕ⋓ᅽࡀᙉࡲࡾࠊ᫛
࿴୰ᮇࡲ࡛㈨※ⓗ౯್ࡀ㧗ࡃࠊ௚ᆅᇦࡼࡾᤕ⋓ᩘࡀከ࠿ࡗࡓ࡞࡝ࡢ≉ᚩࡀ࠶ࡿࠋ࣭ ≉ᐃ㫽⋇ಖㆤ⟶⌮ィ⏬ࡢ⟇ᐃࡢ⫼ᬒ࡜┠ⓗࢆ♧ࡋࡓࠋィ⏬ࡣࠊ㎰ᴗ⿕ᐖࠊ⏕ά⿕
ᐖࠊே㌟஦ᨾࡢ㍍ῶ࡜ゎᾘ࠾ࡼࡧࠊᆅᇦಶయ⩌ࡢ೺඲࡞⥔ᣢࢆ┠ⓗ࡜ࡋ࡚࠸ࡿࠋ࣭ ┠ᶆタᐃ࡟ࡣࠊ㎰ᴗ⿕ᐖࢆ༙ῶࡍࡿࡇ࡜࡜ࡋࠊࡑࡢ㐩ᡂࡢࡓࡵࠊ⊁⊟ᮇ㛫ࡢᘏ㛗ࠊࢶ࢟ࣀ࣡ࢢ࣐ࡢ㘒ㄗᤕ⋓ࡢྍ⬟ᛶࡢ࡞࠸ῐ㊰ᆅᇦ࡟࠾࠸࡚ࠊࡃࡃࡾ⨜ࡢ┤ᚄ 12੉௨ୖࡢไ㝈ࢆゎ㝖ࡍࡿ࡞࡝ࡢᤕ⋓つไࡢ⦆࿴࡟ྲྀࡾ⤌ࢇ࡛࠸ࡿࠋ
࣭ ㏆ᖺࡢᤕ⋓ᩘࡢቑຍࡣࠊ⊁⊟つไࡢ⦆࿴࡟ࡼࡿࡶࡢࡔࡅ࡛࡞ࡃ⏕ᜥᩘࡢቑຍࡶ཯
ᫎࡋ࡚࠸ࡿ࡜⪃࠼ࡽࢀࡿࠋ㐺ษ࡞ಶయᩘ⟶⌮ࡢࡓࡵࡢᤕ⋓ᙉ໬ࡣ௒ᚋࡶᚲせ࡛࠶ࡿࠋ㎰ᴗ⿕ᐖ㜵Ṇ࡟ࡣࠊ≀⌮ⓗ࡞㜵⾨ࡢయไࢆᩚ࠼ࡿࡇ࡜ࡀ㔜せ࡛࠶ࡿࠋKey words㸸≉ᐃ㫽⋇ಖㆤ⟶⌮ィ⏬ࠊ⊦ᇉࠊ⊁⊟㈨※ࠊ᭷ᐖᤕ⋓ࠊ㎰ᴗ⿕ᐖࡣࡌࡵ࡟
රᗜ┴࡟࠾ࡅࡿࢽ࣍ࣥ࢖ࣀࢩࢩ㸦Sus scrofa leucomystax㸹௨ୗ࢖ࣀࢩࢩ࡜⾲グ㸧ࡢ⏕ᜥࡣࠊྂ௦ࡢฟᅵရ࡟ᥥ࠿ࢀࡓ⤮ࡸ㑇㊧ࠊ᪥ᮏ᭩⣖࡞࡝ࡢṔྐⓗ㈨ᩱ࠿ࡽ▱ࡽࢀ࡚࠸ࡿ㸦すᮧ 2003㸧ࠋ⦖ᩥᅵჾ࡟ࡣࠊேࡀ࢖ࣀࢩࢩࢆᘪ▮ࡸ≟ࢆ⏝࠸࡚ᤕ⋓ࡍࡿሙ㠃ࡀᥥ࠿ࢀ࡚࠸ࡓࡾࠊ㈅ሯࡢฟᅵရ࡞࡝࠿ࡽࠊᙜ᫬࠿ࡽ㈗㔜࡞㣗ᩱ㈨※࡛࠶ࡿࡇ࡜ࡀ࠺࠿ࡀ࠼ࡿ
㸦ෆᒣ 2005㸧ࠋࡑࡢᚋࠊᘺ⏕᫬௦࡟✄సࡀⓎᒎࡍࡿ࡜ࠊࡴࡋࢁᐖ⋇࡜ࡋ࡚ࡢ఩⨨࡙ࡅࡶᙉࡃ࡞ࡗࡓ࡜⪃࠼ࡽࢀ࡚࠸ࡿ㸦⏣ཱྀ 2000㸧ࠋỤᡞ୰ᮇ࡟ࡣࠊ᪥ᮏྛᆅ࡛⋇ᐖࡀ῝้໬ࡋࡓࡇ࡜ࡀᵝࠎ࡞Ṕྐ᭩࡞࡝࡟グ㘓ࡉࢀ࡚࠾ࡾࠊ⌧௦ࡢ῝้࡞⿕ᐖࡢ≧ἣ࡜㢮ఝࡋ࡚࠸ࡿⅬࡀ࠶ࡿ࡜࠸ࢃࢀ࡚࠸ࡿ㸦⏣ཱྀ 2000㸧ࠋࡇࡢࡇࢁ἞Ᏻ⥔ᣢࡢࡓࡵࠊᖥᗓ࡟ࡼࡿ㖠つไࡀᙉ໬ࡉࢀ㸦Ṋ஭ 2010㸧
ࠊ㎰ᆅࢆᏲࡿࡓࡵ࡟
⊦ᇉ࡜࿧ࡤࢀࡿ▼✚ࡳࡢ⋇㢮ࡢ౵ධ㜵Ṇቨࡀ᪥ᮏྛᆅ࡛సࡽࢀ࡚࠸ࡓ㸦▮ࣨᓮ 2001㸧ࠋරᗜ┴᳃ᯘື≀◊✲ࢭࣥࢱ࣮࣭ රᗜ┴❧኱Ꮫ⮬↛࣭⎔ቃ⛉Ꮫ◊✲ᡤ රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-1

රᗜ┴ෆ࡛ࡣࠊ⌧ᅾࡢす⬥ᕷ㯮⏣ᗉ⏫ࠊྠ୰⏫ࠊከྍ⏫ຍ⨾⏫ࠊ୹Ἴᕷᒣ༡⏫ࠊ⚄ᡞᕷ໭༊ῐἙ⏫ࠊྠᒣ⏣⏫ࠊ┦⏕ᕷࠊ㉥✑ᕷࠊὪᮏᕷࠊ༡࠶ࢃࡌ⏫࡟⊦ᇉࡀ☜ㄆࡉࢀ࡚࠸ࡿ
㸦ᒸᮏ 1989㸧ࠋす⬥㯮⏣ᗉ⏫ࠊຍ⨾⏫࡛Ⓨぢࡉࢀࡓ⊦ᇉࡣࠊ㧗ࡉࡀ 2m ࡯࡝࠶ࡿࡇ࡜࠿
ࡽ࢖ࣀࢩࢩࡔࡅ࡛࡞ࡃࢽ࣍ࣥࢪ࢝㸦Cervus nippon㸧࡟ࡶᑐᛂࡋ࡚࠸ࡓ࡜⪃࠼ࡽࢀ࡚࠸
ࡿࠋࡲࡓࠊỤᡞ᫬௦࡟⫗㣗ࡀ⚗Ṇࡉࢀ࡚࠸ࡿ࡜࠸ࢃࢀ࡚࠸ࡓ᫬௦࡟ࡶᐇ㝿࡟ࡣࠊᒣ㪒࡜ࡋ࡚㣗࡭⥆ࡅࡽࢀࠊ⏕ᴗ࡜ࡋ࡚ࡢ⊟ᖌࡶᏑᅾࡋ࡚࠸ࡓࠋ
᫂἞࡟࡞ࡿ࡜ࠊᨻᗓࡀ㔝⏕ື≀ࡢᤕ⋓ࢆゎ⚗ࡋࠊศᕸᇦࡣ㏆␥௨す࡟㝈ࡽࢀࡿࡼ࠺࡟
࡞ࡗࡓ㸦㧗ᶫ 2006㸧ࠋරᗜ┴ෆ࡛ࡣࠊ⌧ᅾࡢ⠛ᒣᕷ࡟㝣㌷ࡀ㥔ᒡࡋࡓࡇ࡜࡟ࡼࡾࠊ㌷㝲ࡢ⦎⩦࡜ࡋ࡚࢖ࣀࢩࢩ⊟ࡀ⾜ࢃࢀ࡚࠸ࡓグ㘓ࡀ࠶ࡾࠊࡲࡓ㌷㝲㣗࡜ࡋ࡚࢖ࣀࢩࢩ⫗ࡀዲࡲࢀ࡚࠸ࡓࡇ࡜࡞࡝࠿ࡽ㸦⚄ᓮ 2001㸧ࠊ᫂἞୰ᮇ࡟ࡶᮏᆅᇦ࡟ࡣẚ㍑ⓗከࡃ࢖ࣀࢩࢩࡀ
⏕ᜥࡋ࡚࠸ࡓࡇ࡜ࡀ࠺࠿ࡀ࠼ࡿࠋࡓࡔࡋࠊ᫂἞᫬௦ࡣࠊ᳃ᯘ㈨※࡟㐣ᗘ࡟౫Ꮡࡋࡓ⏕άࡸ⏘ᴗᙧែࡀ㐍ࡳࠊ⸄Ⅳᯘࡣ⏕⏘ຊࢆኻ࠺≧ἣ࡜࡞ࡗ࡚Ⲩᆅ࡜࡞ࡗࡓᆅᇦࡀቑຍࡋࡓ
㸦ᅗ㸯㸹኱⏣ 2012㸧ࠋࡑࡢࡓࡵࠊ࢖ࣀࢩࢩࡢ⏕ᜥ⎔ቃࡶᝏ໬ࡋࠊ㧗࠸⊁⊟ᅽࡀ࠿࠿ࡿ࡜
ಶయᩘࢆᅇ᚟ࡋ࡟ࡃ࠸≧ἣࡀⓎ⏕ࡋ࡚࠸ࡓ࡜⪃࠼ࡽࢀࡿ㸦኱᮰̺ఀ⸨࣭⚄ᓮ 1998㸧ࠋ ᖺ㡭 ᖺ㡭ᅗ  රᗜ┴࡟࠾ࡅࡿᅵᆅ฼⏝ࡢኚ㑄 ᖺ௦ࡣࠊ➨  ᅇ⮬↛⎔ቃಖ඲ᇶ♏ㄪᰝࢆ⤫ྜࠊ ᖺ௦ࡣᅜᅵᆅ⌮㝔Ⓨ⾜ࡢ  ୓ศࡢ㸯ࡢᆅᙧᅗ㸦᫛࿴ 㸫 ᖺ㸧ࢆࢹࢪࢱࣝ໬ࡋࡓࠋ  රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-1

኱ṇ࣭᫛࿴ࡢ᫬௦ࡣࠊ඲ᅜⓗ࡟ࡶ⊁⊟ࡀ┒ࢇ࡟⾜ࢃࢀࡿࡼ࠺࡟࡞ࡾࠊ㧗࠸⊁⊟ᅽࡀ࠿࠿ࡗ࡚࠸ࡓ࡜⪃࠼ࡽࢀࡿࡀ㸦⏣ཱྀ 2000㸧ࠊࡇࡢ㡭ࡢ┴ෆࡢ᝟ሗࡣᑡ࡞࠸ࡇ࡜࠿ࡽࠊಶయ
ᩘࡣῶᑡࡋ࡚࠸ࡓ࡜⪃࠼ࡽࢀࡿ㸦኱᮰̺ఀ⸨࣭⚄ᓮ 1998㸧ࠋࡑࡢᚋࠊ᫛࿴ 30 ᖺ௦ᚋ༙࠿ࡽ 50 ᖺ௦୰┙ࡲ࡛ࡣࠊරᗜ┴ෆ࡛ࡣ 4000 㢌࠿ࡽ 8000 㢌࡯࡝ࡢᤕ⋓ᩘ࡛᥎⛣ࡋ࡚࠾ࡾࠊ඲ᅜⓗ࡟ࡶ㧗࠸ᤕ⋓ᩘࡀ⥔ᣢࡉࢀ⥆ࡅ࡚࠸ࡿࠋࡓ࡜࠼ࡤࠊ᫛࿴ 43 ᖺࡣࠊරᗜ┴ࡢᤕ⋓ᩘࡣ 4854 㢌࡛࠶ࡾࠊ୕㔜┴ࠊᐑᓮ┴࡟ḟ࠸࡛ 3 ␒┠ࡢᤕ⋓ᩘ࡛࠶ࡗࡓ㸦⚄ᡞ᪂⪺♫ 1974㸧ࠋࡇࡢ㡭ࠊ⇞ᩱ㠉࿨࡟ࡼࡾ⸄Ⅳᯘ࡜ࡋ࡚฼⏝ࡉࢀ࡚ࡁࡓ㔛ᒣᯘࡀ୍ᩧ࡟ᨺᲠࡉ
ࢀࡿࡼ࠺࡟࡞ࡾࠊ࢔࣐࢝ࢶ-ࢥࢼࣛᯘ࡞࡝ࡢᗈⴥᶞᯘࡀ෌⏕ࡋࡓࠋ1950 ᖺ௦࡜ 1980 ᖺ௦ࡢ᳜⏕ᅗࢆẚ㍑ࡍࡿ࡜᳃ᯘࡢ෌⏕ࡢᵝᏊࡀఛ࠼ࡿ㸦ᅗ㸯㸧ࠋ
᫛࿴ 50 ᖺ௦௨㝆ࡣࠊ᪤࡟㈨※ⓗ౯್ࡀ㧗࠿ࡗࡓ࢖ࣀࢩࢩ⫗ࡣࠊࠕࡰࡓࢇ㘠ࠖ࡞࡝ࡢࣈ
࣮࣒࡟ࡢࡗ࡚ࡉࡽ࡟ᾘ㈝ࡀ┒ࢇ࡟࡞ࡗࡓ࡜⪃࠼ࡽࢀࡿ㸦ᮅ᪥ 1985㸹኱᮰̺ఀ⸨࣭⚄ᓮ
1998㸹㉥ᫍ 2001㸧ࠋࡋ࠿ࡋࠊ50 ᖺ௦ᚋ༙࠿ࡽᖹᡂ 9 ᖺࡲ࡛ࡢᤕ⋓ᩘࡣ 4000 㢌௨ୗ࡜ῶᑡࡍࡿࠋ㈨※ⓗ࡞౯್ࡢ㧗ࡉ࠿ࡽ㧗࠸⊁⊟ᅽࡀ࠿࠿ࡗ࡚࠸ࡓࡇ࡜ࡀせᅉ࡜⪃࠼ࡽࢀࡿ㸦኱᮰̺ఀ⸨࣭⚄ᓮ 1998㸧ࠋࡑࡢᚋᖹᡂ 10 ᖺ࠿ࡽࡣᤕ⋓ᩘࡀቑຍഴྥ࡟࠶ࡾࠊᖹᡂ 16ᖺ௨㝆ࠊᤕ⋓ᩘࡀ 1 ୓㢌ࢆ㉸࠼ࡿࡇ࡜ࡶከࡃ࡞ࡗ࡚ࡁࡓ㸦ᅗ㸰㸧ࠋ 
20,00018,00016,00014,00012,00010,000
8,000
6,000
4,000
2,000

H24

H22

H20

H18

H16

H14

H12

H10

H8

H6

H4

H2

S63

S61

S59

S57

S55

S53

S51

S49

S47

S45

S43

S41

S39

S37

0

   ᅗ  රᗜ┴࡟࠾ࡅࡿ࢖ࣀࢩࢩࡢᤕ⋓ᩘࡢ᥎⛣㸦᫛࿴  ᖺ㹼ᖹᡂ  ᖺ㸧ᖹᡂ 22 ᖺ࡟ࡣࠊᤕ⋓ᩘ㸦⊁⊟࡜᭷ᐖᤕ⋓㸧ࡣ 1 ୓ 8 ༓㢌࡟㐩ࡋࠊ㐣ཤ 40 ᖺࢆࡉ࠿ࡢࡰࡗ࡚ࡶ≉ฟࡋࡓ್࡜࡞ࡗࡓ㸦රᗜ┴ 2013㸧ࠋᖹᡂ 20 ᖺ㹼24 ᖺࡢᤕ⋓ᩘࢆᖹᆒࡍࡿ
࡜ 12,957 㢌/ᖺ࡜࡞ࡿࠋࡇࢀࢆ᳃ᯘ㠃✚ᙜࡓࡾࡢᤕ⋓ᩘ࡟᥮⟬ࡍࡿ࡜ᖹᡂ 22 ᖺࡣ 36.5
㢌/੎ࠊᖹᡂ 20 ᖺ㹼24 ᖺࡢᤕ⋓ᩘࡢᖹᆒࡣ 23.1 㢌/੎࡜࡞ࡿࠋྠࡌࡃ 40 ᖺ௦ࡣࠊ10.0
㢌/੎ࠊ50 ᖺ௦ࡣ 9.2 㢌/੎ࠊ60 ᖺ௦࠿ࡽᖹᡂ 9 ᖺࡲ࡛ࡣ 6.1 㢌/੎࡜ẚ㍑ࡍࡿ࡜ࡇࡢᩘ
ᖺࡣࠊᛴ⃭࡟ᤕ⋓ᩘࡀቑຍࡋ࡚࠸ࡿࠋ රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-1

ᖹᡂ 10 ᖺ௨㝆ࡣࠊ᭷ᐖᤕ⋓ࡶቑຍഴྥ࡟࠶ࡾࠊ⌧ᅾ࡛ࡣࠊ⊁⊟࡜᭷ᐖᤕ⋓ࡢ๭ྜࡣ࡯ࡰ 1:1 ࡜࡞ࡾࠊ㎰ᴗ⿕ᐖࡢ῝้ࡉ࡜௨ୗ࡟㏙࡭ࡿ⊁⊟つไ⦆࿴࡟ࡼࡿᤕ⋓ᙉ໬ࡢྲྀࡾ
⤌ࡳࢆ཯ᫎࡋ࡚࠸ࡿ࡜⪃࠼ࡽࢀࡿࠋࡋ࠿ࡋつไ⦆࿴࡟ࡼࡿᤕ⋓ᩘࡢቑຍ࡜࡜ࡶ࡟⪃៖ࡍ࡭ࡁࡣࠊ㏆ᖺࡢᛴ⃭࡞⊁⊟⪅ࡢῶᑡ࡛࠶ࡿ㸦㕥ᮌ࡯࠿ 2003㸧ࠋࡇࢀࡽࢆ຺᱌ࡍࡿ࡜ 2010ᖺ௦ࡣࠊᤕ⋓ᩘࡀᛴ⃭࡞ቑຍࢆぢࡏ࡚࠾ࡾࠊಶయᩘࡢቑຍࢆ཯ᫎࡋ࡚࠸ࡿྍ⬟ᛶࡀᙉࡃ♧၀ࡉࢀࡓࠋ
රᗜ┴࡟࠾ࡅࡿ≉ᐃ㫽⋇ಖㆤ⟶⌮ィ⏬ࡢ⌧≧රᗜ┴࡛ࡣࠊࡇࢀࡲ࡛♧ࡋࡓࡼ࠺࡟ே࡜࢖ࣀ
ࢩࢩࡢ㛵ࢃࡾࡣ᪥ᮏேࡀ᪥ᮏิᓥ࡟Ώ᮶ࡋ࡚࠿
ࡽࠊ࡯ࡰࠊ㏵ษࢀࡿࡇ࡜࡞ࡃ⥆࠸࡚࠾ࡾࠊ኱ṇ᫛࿴ึᮇ࡟ᮾ᪥ᮏ࡛ࡢศᕸࡀ㏵⤯࠼ࡓ࡜ࡁ࡟ࡶ
ẚ㍑ⓗከࡃ⏕ᜥࡋ࡚࠸ࡓ࡜⪃࠼ࡽࢀ࡚࠸ࡿ㸦⚄ᡞ᪂⪺♫ 1974㸧ࠋࡑࡢࡓࡵࠊ㏆ᖺࡲ࡛ࠊࢺࢱࣥ
ᯈ࡟ࡼࡿ㜵ㆤᰙ࡞࡝㎰ᴗ⿕ᐖࡢᑐᛂ࡜㈨※ⓗ฼⏝ࢆ┠ⓗ࡜ࡋࡓ⊁⊟ࡀ⥅⥆ࡋ࡚⾜ࢃࢀ࡚࠸ࡓࠋ
⊁⊟ࡣࠊ඲┴ᇦ࡛࣡ࢼ⊟ࡀࠊ㜰⚄㛫ࡢ㖠⊟⚗Ṇ༊ᇦࢆ㝖ࡃᗈ࠸ᆅᇦ࡛㖠⊟ࡀ⾜ࢃࢀ࡚࠾ࡾ㸦ᅗ 㸧ᖹᆅ࠿ࡽᒣ㛫㒊ࡲ࡛࡯ࡰ඲┴࡟⏕ᜥࡋ࡚࠸ࡿࠋරᗜ┴࡛ࡣ࢖ࣀࢩࢩ࡟ᑐࡋ࡚ࠊ㛗ᮇⓗ࡟㜵ㆤ   ᅗ  රᗜ┴࡟࠾ࡅࡿ࢖ࣀࢩࢩࡢ
࡜ᤕ⋓ࡢᑐ⟇ࢆ⥅⥆ࡋ࡚ࡁࡓṔྐࡀ࠶ࡾࠊᨵࡵ     ┠ᧁຠ⋡ࡢศᕸ࡚ಖㆤ⟶⌮ィ⏬ࢆ⟇ᐃࡍࡿᚲせᛶࡣపࡃࡃࠊ≉    ᐃ㫽⋇ಖㆤ⟶⌮ィ⏬ࡣࠊไᗘⓎ㊊ᚋࡶࡋࡤࡽࡃ⟇ᐃࡉࢀ࡞࠿ࡗࡓࠋ
ࡋ࠿ࡋࠊ๓㏙ࡋࡓࡼ࠺࡟ࠊᖹᡂ 16 ᖺ㡭࠿ࡽ᭷ᐖᤕ⋓ࡢ๭ྜࡀᛴ⃭࡟ቑຍࡋ㸦ᅗ㸰㸹ᯇᮏ࡯࠿ 2011㸧ࠊࡲࡓ௚ࡢ⋇✀ࠊ≉࡟ྠᵝࡢ⿕ᐖࢆᘬࡁ㉳ࡇࡋ࡚࠸ࡿࢽ࣍ࣥࢪ࢝ࡢ⊁⊟ᮇ㛫ࡀᘏ㛗ࡉࢀࡿࡇ࡜࡟క࠺ၥ㢟ࡸ࢖ࣀࢩࢩࡢ᭷ᐖᤕ⋓⨜࡟ࢶ࢟ࣀ࣡ࢢ࣐ࡀ㘒ㄗᤕ⋓ࡉࢀࡿࡇ࡜ࡀከⓎࡋࡓࡇ࡜㸦✄ⴥ 2011㸧ࠊῐ㊰ᓥࡢ࢖ࣀࣈࢱ⏤᮶ࡢ࢖ࣀࢩࢩࡢᛴቑࠊභ
⏥ᒣ࢖ࣀࢩࢩࡢᕷ⾤ᆅ࡬ࡢฟἐ࡜㣵௜ࡅၥ㢟㸦㎷࣭ᶓᒣ 2014a㸹㎷࣭ᶓᒣ 2014b㸧࡞࡝ಶయᩘࡢቑຍ࡟క࠺ྛ✀ၥ㢟ࡀቑຍࡋ࡚ࡁࡓࠋ㎰ᴗ⿕ᐖ࡟ࡘ࠸࡚ࡶᖺ㛫㸰൨෇ࢆ㉸࠼ࡿ≧ἣࡀ⥆࠸࡚࠾ࡾࠊࡉࡽ࡞ࡿᑐ⟇ࡢᙉ໬ࡀᚲせ࡜࡞ࡗ࡚ࡁࡓ㸦ᅗ 4㸧ࠋࡇࢀࡽࡢ⫼ᬒ࠿ࡽࠊ࢖ࣀࢩࢩ࡟࠾࠸࡚ࡶ⟶⌮ࡢࡓࡵࡢ┠ᶆタᐃ࡜᪉⟇ࢆ᫂☜໬ࡉࡏࡿࡇ࡜ࡀᚲせ࡜࡞ࡾࠊᖹᡂ 21 ᖺ࡟➨ 1 ᮇࡢィ⏬ࡀ⟇ᐃࡉࢀࡓ㸦රᗜ┴ 2009㸧ࠋィ⏬⟇ᐃ࡟࠾ࡅࡿ┠ⓗࡣࠊ㎰ᴗ⿕ᐖࠊ⏕ά⿕ᐖࠊே㌟஦ᨾࡢ㍍ῶ࡜ゎᾘ࠾ࡼࡧࠊᆅᇦ රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-1

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㻴㻝 㻴㻞 㻴㻟 㻴㻠 㻴㻡 㻴㻢 㻴㻣 㻴㻤 㻴㻥 㻴㻝㻜 㻴㻝㻝 㻴㻝㻞 㻴㻝㻟 㻴㻝㻠 㻴㻝㻡 㻴㻝㻢 㻴㻝㻣 㻴㻝㻤 㻴㻝㻥 㻴㻞㻜 㻴㻞㻝 㻴㻞㻞 㻴㻞㻟ᖺḟᅗ  රᗜ┴࡟࠾ࡅࡿ࢖ࣀࢩࢩࡢ㎰ᴗ⿕ᐖ㔠㢠ࡢ᥎⛣ಶయ⩌ࡢ೺඲࡞⥔ᣢ࡜ࡋ࡚࠸ࡿ㸦රᗜ┴ 2009㸧ࠋ┠ᶆタᐃ࡟࠾࠸࡚ࡣࠊ㎰ᴗ⿕ᐖ࡜ಶయᩘࡢᐦᗘᣦᩘ࡛࠶ࡿ┠ᧁຠ⋡࡜ࡢ㛵ಀᛶࡀぢࡽࢀࡓ㸦ᆏ⏣࡯࠿ 2008㸧ࡇ࡜࠿ࡽࠊ┠ᶆ
ࡣ㎰ᴗ⿕ᐖࡢ༙ῶࢆ┠ᣦࡍࡇ࡜࡜ࡋ࡚࠸ࡿࠋලయⓗ࡟ࡣࠕ῝้ࠖ࡞㞟ⴠࡢ๭ྜ㸲㸣௨ୗࠊࠕ኱ࡁ࠸ࠖ㞟ⴠࡢ๭ྜ 10㸣௨ୗ࡜ࡍࡿࡇ࡜࡜ࡋ࡚࠸ࡿ㸦ᅗ 5㸹රᗜ┴ࠊ2013㸧ࠋࡇࢀࡽࡢ┠ᶆࢆ㐩ᡂࡍࡿࡓࡵࡢ᪉⟇ࡣࠊ⊁⊟ᮇ㛫ࡢᘏ㛗㸦2 ᮇ࡛ࡣ 3 ᭶ 15 ᪥ࡲ࡛ࡢᘏ㛗㸧ࠊࢶ࢟ࣀ࣡ࢢ࣐ࡢ㘒ㄗᤕ⋓ࡢྍ⬟ᛶࡢ࡞࠸ῐ㊰ᆅᇦ࡟࠾࠸࡚ࡣࠊࡃࡃࡾ⨜ࡢ┤ᚄ 12 ੉௨ୖࡢไ㝈ゎ㝖࡞࡝୺࡟ᤕ⋓ಁ㐍ࡢࡓࡵࡢྲྀࡾ⤌ࡳ࡛࠶ࡿࠋ⊁⊟ᮇ㛫ࢆᘏ㛗ࡋ࡚࠿ࡽࡣࠊᤕ
⋓ᩘࡀᛴ⃭࡟ቑຍࡋ࡚࠾ࡾࠊ⊁⊟ᮇ㛫ࡢᘏ㛗ࡢຠᯝࡀ㧗࠸ࡇ࡜ࡀ࠺࠿ࡀ࠼ࡿ㸦ᅗ 2㸹ᯇᮏ࡯࠿ 2014㸧ࠋࡓࡔࡋࠊ࢖ࣀࢩࢩࡢ㏆ᖺࡢᤕ⋓ᩘࡣࠊᖺኚືࡀ኱ࡁࡃࠊᤕ⋓ᅽࢆቑ኱ࡉ
ࡏࡿ࡞࡝ࡢ⊁⊟つไ⦆࿴௨እࡢせᅉࡀࠊ࢖ࣀࢩࢩࡢᤕ⋓ᩘ࡟ᙳ㡪ࢆ୚࠼࡚࠸ࡿࡇ࡜ࡶㄞ
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䛺䛔 䜋䛸䜣䛹䛺䛔 ㍍ᚤ ኱䛝䛔䠄⏕⏘㔞䛾䠏䠌䠂ᮍ‶䠅 ῝้䠄⏕⏘㔞䛾䠏䠌䠂௨ୖ䠅ᅗ  ㎰ᴗ⿕ᐖࡢ⛬ᗘࡢᖺኚ໬㸦➨㸰ᮇ࢖ࣀࢩࢩಖㆤ⟶⌮ィ⏬ࡼࡾ㸧 රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-1

ಖㆤ⟶⌮ィ⏬ࡢㄢ㢟
㎰ᮧ㒊࡟࠾ࡅࡿ࢖ࣀࢩࢩࡢ⟶⌮࡟࠾࠸࡚ࠊ௒ᚋࡶᤕ⋓ᅽࡢቑຍ࡜㐺ษ࡞㜵ㆤᰙࡢタ⨨࡜⟶⌮ࡀ㔜せ࡛࠶ࡿࠋ⌧ᅾࡢಖㆤ⟶⌮ィ⏬࡟࠾࠸࡚ࡣࠊ୺࡟ᤕ⋓つไࡢ⦆࿴ࡀ୰ᚰ
࡜࡞ࡗ࡚࠸ࡿࡀࠊ௒ᚋࡶࡑࡢഴྥࡣ⥆ࡃ࡛࠶ࢁ࠺ࠋ
≉࡟ࠊ࢖ࣀࢩࢩࡣከ⫾ື≀࡛ࠊᡂ⋇ࡢ⏘Ꮚᩘࡣᖹᆒ㸲㢌࡛࠶ࡾ㸦Tsuji, et al. 2013;㎷࣭ᶓᒣ 2014c㸧ࠊ⌧ᅾࡢරᗜ┴࡟࠾ࡅࡿ⮬↛ቑຍ⋡ࡣࠊ18.8㸣࠿ࡽ 100.3㸣ࡲ࡛ᗈ࠸ኚືᖜࢆᣢࡘ࡜᥎ᐃࡉࢀ࡚࠸ࡿࡇ࡜࠿ࡽ㸦ᯇᮏ࡯࠿ 2014㸧ࠊ⿕ᐖࡸᤕ⋓ᩘࡶᖺኚືࡀ
኱ࡁ࠸ࠋಶయᩘࡢኚືࡸᖺ࡟ࡼࡿ㈨※ኚື㸦⸨ᮌ࡯࠿ 2011㸧ࡢᙳ㡪ࢆཷࡅ࡚ࠊᤕ⋓ࡋࡸࡍࡉ࡞࡝ࡀ኱ࡁࡃ᣺ࢀࡿྍ⬟ᛶࡀ࠶ࡿࠋࡉࡽ࡟௒ᚋࠊ⊁⊟⪅ࡢῶᑡࡸ㧗㱋໬࡟ࡼࡾᤕ⋓ᅽࡀῶᑡࡋ࡚࠸ࡃྍ⬟ᛶࡀ㧗࠸㸦㕥ᮌ࡯࠿ 2003㸧
ࠋࡇࢀࡽࢆ⪃៖ࡋ࡚ࠊ௒ᚋࡶᙉ
࠸ᤕ⋓ᅽࢆ࠿ࡅ⥆ࡅ࡚࠸ࡃࡇ࡜ࡀᚲせ࡛࠶ࡿࠋῐ㊰ᓥ࡟ࡣࠊ༡㒊ࡢㅍ㭯⩚ᒣ⣔࡟ᅾ᮶ࡢ࢖ࣀࢩࢩࡀ⏕ᜥࡋ࡚࠸ࡿࡀࠊ⏕ᜥࡀ㛗࠸㛫☜ㄆࡉࢀ࡚ࡇ࡞࠿ࡗࡓ໭ῐᆅᇦ࡟ࠊᛴ⃭࡟⏕ᜥࡀᣑ኱ࡋ࡚࠸ࡿၥ㢟ࡀ࠶ࡿ㸦ᅗ 3㸹ᯇᮏ࡯࠿ 2014㸧ࠋࡇࢀ࡟ࡘ࠸࡚ࡣࠊ㣫⫱ࡢ࢖ࣀࣈࢱࡀ㏨ࡆฟࡋࠊ஺㓄ࢆ⧞ࡾ㏉ࡋࡓ࡜࠸࠺ㄝࡀ࠶ࡗࡓࡀࠊࡇࡢᆅᇦࡢ࢖ࣀࢩࢩࡢ㑇ఏᏊࢆㄪ࡭ࡓ࡜ࡇࢁࠊࣈࢱࡢ㑇ఏᏊࡀ☜ㄆࡉࢀ࡚࠸ࡿ㸦රᗜ┴ 2009㸧ࠋ࡯࠿࡟ࡶ࢖ࣀࢩࢩࡢᨺ㏲ࡢშࡣ⪏࠼࡞࠸ࡓࡵࠊ௚ᆅᇦ࠿
ࡽ࢖ࣀࢩࢩࢆᣢࡕ㎸ࡳᨺ㏲ࡍࡿࡇ࡜ࡣࠊ⮬↛ಶయ⩌ࡢ㐺ษ࡞⟶⌮ࡢほⅬ࠿ࡽ୙㐺ษ࡞
⾜Ⅽ࡛࠶ࡿࡇ࡜ࢆᬑཬ࣭ᾐ㏱ࡉࡏࡿᚲせࡀ࠶ࡿࠋࡉࡽ࡟රᗜ┴ࡢ࢖ࣀࢩࢩ⟶⌮࡟࠾ࡅࡿㄢ㢟ࡣࠊ㎰ᴗᆅᇦࡢၥ㢟ࡔࡅ࡛ࡣ࡞࠸ࠋභ⏥ᒣᆅᇦ࡟࠾ࡅࡿ㣵௜ࡅၥ㢟ࡸ㒔ᕷ㒊࡬ࡢฟἐࠊࡑࢀࡽ࡟క࠺ே㌟஦ᨾࡢⓎ⏕࡞࡝ࡀ㠀ᖖ࡟኱ࡁ࡞ㄢ㢟࡜࡞ࡗ࡚࠸ࡿ㸦㎷࣭ᶓᒣ 2014ab㸧ࠋࡇࢀ࡟ࡘ࠸࡚ࡣ 10 ❶㸦㎷࣭ᶓᒣ
2014a㸧ࠊ11 ❶㸦㎷࣭ᶓᒣ 2014b㸧࡟ヲࡋࡃࡲ࡜ࡵࡓࠋࡇࡢ࡯࠿ࠊ㏆ᖺࡇࢀࡲ࡛࢖ࣀ
ࢩࢩࡢ⏕ᜥ᝟ሗࡢ࡞࠿ࡗࡓᖹ㔝㒊࡟✺Ⓨⓗ࡟࢖ࣀࢩࢩࡀฟἐࡋࠊே㌟஦ᨾࢆ㉳ࡇࡍ஦
౛ࡶⓎ⏕ࡋ࡚࠸ࡿ㸦ኴᏊ⏫ p://www.town.taishi.hyogo.jp/dd.aspx?menuid=4623㸧ࠋ௚ᗓ┴࡟࠾࠸࡚ࡶ㒔ᕷ㒊࡟࠾ࡅࡿ࢖ࣀࢩࢩࡢฟἐ⿕ᐖࡀከⓎࡋ࡚࠾ࡾ㸦㤶ᕝ┴ 2011㸧ࠊ௒ᚋ඲ᅜⓗ࡟࢖ࣀࢩࢩࡢ㒔ᕷࡸᕷ⾤ᆅ࡬ࡢฟἐࡢ༴㝤ᛶࡶ㧗ࡲࡗ࡚ࡁ࡚࠸ࡿࠋᕷ⾤ᆅ࡟࠾ࡅࡿ࢖ࣀࢩࢩࡢ⏕ᜥࣔࢽࢱࣜࣥࢢ࡜⥭ᛴฟἐయไࡢᵓ⠏ࡶ㔜せ࡞ㄢ㢟࡛࠶ࡿࠋ࢖ࣀࢩࢩ࡟࠾ࡅࡿಶయᩘࡢ㐺ษ࡞⟶⌮ࡣࠊࢫࢱ࣮ࢺࡋࡓࡤ࠿ࡾ࡛࠶ࡾࠊࣔࢽࢱࣜࣥ
ࢢࢆ㔜ࡡ࡞ࡀࡽࠊヨ⾜㘒ㄗ࡟ࡼࡾ㐺ṇ࡞ᤕ⋓ᅽࡢ࠿ࡅ᪉࡟ࡼࡿᤕ⋓ࡢຠᯝࢆ᳨ドࡋ࡚࠸ࡃᚲせࡀ࠶ࡿࠋ㎰ᴗ⿕ᐖࡢ㜵Ṇ࡟ࡣࠊᤕ⋓つไ⦆࿴࡟㢗ࡿࡔࡅ࡛ࡣ୙༑ศ࡛࠶ࡾࠊ
㐺ษ࡞㞟ⴠᰙࡢタ⨨࡜ࡑࡢ⟶⌮ࡢ௙⤌ࡳ࡙ࡃࡾࡀồࡵࡽࢀࡿࠋ≉࡟㎰ᆅ࡟㞄᥋ࡍࡿ࢖ࣀࢩࢩࡢ⾜ືᅪࡣ⊃ࡃࠊ㜵㝖ࡀ୙༑ศ࡞ሙᡤࢆᏛ⩦ࡋฟἐࡋ࡚࠸ࡿ㸦ᶓᒣ࡯࠿ 2014㸧
ࡓࡵࠊ㎰స≀⿕ᐖࡢ㜵Ṇ࡟ࡣࠊ≀⌮ⓗ㜵⾨ࡀ୙ྍḞ࡛࠶ࡿࠋ㐣ཤࡢṔྐ࠿ࡽぢ࡚ࡶ≉࡟㎰ᴗࢆႠࡴୖ࡛࢖ࣀࢩࢩࡢ㜵㝖ࡣỌ⥆ⓗ࡟ྲྀࡾ⤌ࡴㄢ㢟࡛࠶ࡿࡓࡵࠊே㛫♫఍ࡢయไࢆᵓ⠏ࡍࡿࡇ࡜ࡀ᭱ࡶ㔜せ࡛࠶ࡿ࡜⪃࠼ࡽࢀࡿࠋ රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-1

ᘬ⏝ᩥ⊩㉥ᫍᚰ㸬2001㸬࢖ࣀࢩࢩࡢࡲࡕ㸫୹Ἴ⠛ᒣ 㧗ᶫ᫓ᡂ⦅ࠕ࢖ࣀࢩࢩ࡜ே㛫Ɇඹ࡟⏕ࡁࡿࠖ㸬
pp.290-313㸬ྂ௒᭩㝔㸪ᮾி㸬ᮅ᪥⛱㸬1985㸬㈨※࡜ࡋ࡚ࡢ࢖ࣀࢩࢩ㸬့ங㢮⛉Ꮫ㸬50㸸27-30㸬
⸨ᮌ኱௓࣭ᶓᒣ┿ᘪ࣭ᆏ⏣ᏹᚿ㸬2011㸬රᗜ┴ෆ࡟࠾ࡅࡿࣈࢼ⛉ᶞᮌ㸱✀ࡢሀᯝࡢ㇏ป࡜
ࢶ࢟ࣀ࣡ࢢ࣐ࡢ㣵㈨※࡜ࡋ࡚ࡢホ౯㸬ࠕරᗜ┴࡟࠾ࡅࡿࢶ࢟ࣀ࣡ࢢ࣐ಖㆤ⟶⌮ࡢ⌧≧࡜ㄢ㢟࣭ࠖරᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 3 ྕ࣭pp39-49㸬 රᗜ┴᳃ᯘື≀◊✲ࢭࣥࢱ࣮㸬රᗜ┴㸬2013㸬➨ 1 ᮇ࢖ࣀࢩࢩಖㆤ⟶⌮ィ⏬㸬රᗜ┴㸪⚄ᡞ㸪23 pp.
රᗜ┴㸬2013㸬➨ 2 ᮇ࢖ࣀࢩࢩಖㆤ⟶⌮ィ⏬㸬රᗜ┴㸪⚄ᡞ㸪18 pp.
✄ⴥ୍᫂㸬2011㸬රᗜ┴ࡢࢶ࢟ࣀ࣡ࢢ࣐ࡢฟἐ≧ἣ࡜ᑐ⟇㸬ࠕරᗜ┴࡟࠾ࡅࡿࢶ࢟ࣀ࣡ࢢ࣐
ಖㆤ⟶⌮ࡢ⌧≧࡜ㄢ㢟ࠖ㸪රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 3 ྕ㸬pp.1-17㸬 රᗜ┴᳃ᯘື≀◊✲ࢭࣥࢱ࣮㸬㤶ᕝ┴㸬2011㸬࢖ࣀࢩࢩ➼ࡀฟἐࡋࡓ࡜ࡁࡢᑐᛂ࣐ࢽࣗ࢔ࣝ㸬㤶ᕝ┴࢖ࣀࢩࢩ➼ᑐ⟇㐃⤡఍㆟㸪15pphttp://www.pref.kagawa.lg.jp/kankyo/data/topics/pdf/manual.pdf㸬2014. 2.
12 accessed.
⚄ᓮఙኵ㸬2001㸬࢖ࣀࢩࢩࡢၟရ໬࡜ಶయ⩌⟶⌮㸬㧗ᶫ᫓ᡂ⦅ࠕ࢖ࣀࢩࢩ࡜ே㛫Ɇඹ࡟⏕ࡁࡿࠖ㸬pp.258-288㸬ྂ௒᭩㝔㸪ᮾி㸬⚄ᡞ᪂⪺♫Ꮫⱁ㒊රᗜ᥈᳨⮬↛⦅ྲྀᮦ⌜㸬1974㸬රᗜ᥈᳨࣭⮬↛⦅㸬⚄ᡞ᪂⪺♫㸪⚄ᡞ㸬ෆᒣ⣧ⶶ㸬2005㸬⦖ᩥேࡢ⛣ື⏕ά㸫⦖ᩥ᫬௦ࡢ⏕άࣃࢱ࣮ࣥࡢኚ㑄࡜ືᅉ㸫㸬ụ㇂࿴ಙ࣭㛗㇂ᕝᨻ⨾⦅ࠕ᪥ᮏࡢ⊁⊟᥇㞟ᩥ໬㸫㔝⏕⏕≀࡜࡜ࡶ࡟⏕ࡁࡿ㸫ࠖ㸬pp.73-97. ୡ⏺ᛮ᝿♫㸪ி㒔㸬
኱⏣⊛ᙪ㸬2012㸬᳃ᯘ㣬࿴㸫ᅜᅵࡢኚㇺࢆ⪃࠼ࡿ㸫㸬pp.260. NHK ฟ∧㸪ᮾி㸬ᒸᮏ኱஧㑻㸬1989㸬රᗜࡢ⊦ᇉɆ⋇ᐖ㜵㝖ྐࡢ㸯㈨ᩱ࡜ࡋ࡚Ɇ㸬ᏘหἙ̺ᖖẸ◊✲ྐ㸬ຍྂᕝὶᇦྐᏛ఍㸪➨ 14 ᕳ㸦2㸧42㸸5㸫11㸬
すᮧ୕㑻㸬2003㸬ẟ⓶࡜ே㛫ࡢṔྐ㸬pp. 388. ⣖ఀᅧᒇ᭩ᗑ㸪ᮾி㸬኱᮰㸫ఀ⸨⤮㔛Ꮚ࣭⚄ᓮఙኵ㸬 1998㸬 ㏆࣭⌧௦ࡢࢽ࣍ࣥ࢖ࣀࢩࢩࡢಶయ⩌ࢺࣞࣥࢻ㸬 㔝⏕⏕≀ಖㆤ㸪 3: 95-105㸬ᆏ⏣ᏹᚿ࣭㩪ᓥᘯග࣭ᶓᒣ┿ᘪ㸬2008㸬┠ᧁຠ⋡࠿ࡽࡳࡓ࢖ࣀࢩࢩࡢ⏕ᜥ≧ἣ࡜✚㞷㸪᳜⏕㸪ࢽ࣍ࣥࢪ࢝㸪⊁⊟㸪㎰ᴗ⿕ᐖ࡜ࡢ㛵ಀ㸬 ့ங㢮⛉Ꮫ㸬48㸸245-253㸬㕥ᮌ∾࣭ᆏ⏣ᏹᚿ࣭⏣୰ဴኵ㸬2003㸬රᗜ┴࡟࠾ࡅࡿ⊁⊟⪅ேཱྀࡢືែ㸬ே࡜⮬↛㸪14㸸
33-41㸬
㧗ᶫ᫓ᡂ㸬2006㸬ே࡜⏕ࡁ≀ࡢᆅ⌮㸬ྂ௒᭩㝔㸪ᮾி㸪120 pp.
⏣ཱྀὒ⨾㸬2000㸬ิᓥ㛤ᣅ࡜⊁⊟ࡢ࠶ࡺࡳ㸬ᮾ໭Ꮫ㸪3㸸67-102㸬Ṋ஭ᘯ୍㸬2010㸬㕲◙ࢆᡭᨺࡉ࡞࠿ࡗࡓⓒጣࡓࡕ㸬ᮅ᪥᪂⪺ฟ∧♫㸪ᮾி㸪256 pp.
Tsuji, T㸬, Yokoyama, M㸬, Asano, M㸬, Suzuki, M㸬2013㸬 Estimation of the fertility ratesof Japanese wild boars (Sus scrofa leucomystax) using fetuses and corpora albicans㸬  රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-1

  Acta Theriol 58(3): 315-323㸬
㎷▱㤶࣭ᶓᒣ┿ᘪ㸬2014a㸬භ⏥ᒣ࢖ࣀࢩࢩၥ㢟ࡢ⌧≧࡜ㄢ㢟㸬ࠕරᗜ┴࡟࠾ࡅࡿࢽ࣍ࣥ࢖
ࣀࢩࢩࡢಖㆤ⟶⌮ࡢ⌧≧࡜ㄢ㢟㸬ࠖ㸪࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6 ྕ㸪pp121-134㸪රᗜ┴᳃ᯘື≀◊✲ࢭࣥࢱ࣮㸬㎷▱㤶࣭ᶓᒣ┿ᘪ㸬2014b㸬භ⏥ᒣࡢࢽ࣍ࣥ࢖ࣀࢩࢩࡢၥ㢟ಶయࡢ≉ᚩ㸬ࠕරᗜ┴࡟࠾ࡅࡿࢽ࣍ࣥ࢖ࣀࢩࢩࡢಖㆤ⟶⌮ࡢ⌧≧࡜ㄢ㢟㸬ࠖරᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6 ྕ㸪pp135-142㸬රᗜ┴᳃ᯘື≀◊✲ࢭࣥࢱ࣮㸬㎷▱㤶࣭ᶓᒣ┿ᘪ㸬2014c㸬රᗜ┴࡟࠾ࡅࡿࢽ࣍ࣥ࢖ࣀࢩࢩࡢᇶᮏⓗ⦾Ṫ≉ᛶ㸬ࠕරᗜ┴࡟   ࠾ࡅࡿࢽ࣍ࣥ࢖ࣀࢩࢩࡢಖㆤ⟶⌮ࡢ⌧≧࡜ㄢ㢟㸬ࠖරᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6 ྕ㸪pp84-92㸪රᗜ┴᳃ᯘື≀◊✲ࢭࣥࢱ࣮㸬ᯇᮏᓫ࣭ᆏ⏣ᏹᚿ࣭ᓊᮏᗣ㄃㸬2014㸬රᗜ┴࡟࠾ࡅࡿࢽ࣍ࣥ࢖ࣀࢩࢩࡢ⏕ᜥศᕸ࡜㎰ᴗ⿕ᐖࡢ㛵ಀ㸬ࠕරᗜ┴ࡢࢽ࣍ࣥ࢖ࣀࢩࢩࡢಖㆤ⟶⌮ࡢ⌧≧࡜ㄢ㢟ࠖ, රᗜ࣡࢖ࣝࢻࣛ࢖ࣇ
ࣔࣀࢢࣛࣇ 6 ྕ, pp9-20㸬රᗜ┴᳃ᯘື≀◊✲ࢭࣥࢱ࣮㸬▮ࣨᓮᏕ㞝㸬2001㸬ࢩࢩᇉࡢศᕸ࡜ᵓ㐀㸬㧗ᶫ᫓⦅ࠕ᪥ᮏࡢࢩࢩᇉࠖ
㸪pp.2-26㸬ྂ௒᭩㝔㸪  ᮾி㸬ᶓᒣ┿ᘪ࣭Ụ⸨බಇ࣭ᮌୗ⿱⨾Ꮚ㸬2014㸬㎰ᆅ࡟㞄᥋ࡋ࡚⏕ᜥࡍࡿࢽ࣍ࣥ࢖ࣀࢩࢩࡢຍᐖ⾜ືࡢゎᯒ㸬ࠕරᗜ┴࡟࠾ࡅࡿࢽ࣍ࣥ࢖ࣀࢩࢩࡢಖㆤ⟶⌮ࡢ⌧≧࡜ㄢ㢟㸬ࠖරᗜ࣡࢖ࣝ
ࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6 ྕ㸪pp43-58㸪රᗜ┴᳃ᯘື≀◊✲ࢭࣥࢱ࣮㸬



 

රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-2➨ 㸰 ❶            රᗜ┴࡟࠾ࡅࡿࢽ࣍ࣥ࢖ࣀࢩࢩࡢ
⏕ᜥศᕸ࡜㎰ᴗ⿕ᐖࡢ㛵ಀᯇᮏ ᓫ 1,2࣭ᆏ⏣ᏹᚿ 1,2࣭ᓊᮏᗣ㄃ 3
せ Ⅼ
࣭ 2002 ᖺ࠿ࡽ 2012 ᖺࡲ࡛ࡢ 11 ᖺ㛫ࡢරᗜ┴࡟࠾ࡅࡿ࢖ࣀࢩࢩࡢ⏕ᜥศᕸࠊ㎰ᴗ⿕ᐖࠊ⿕ᐖᑐ⟇ࠊ࠾ࡼࡧᑐ⟇ࡢຠᯝ࡟ࡘ࠸࡚ࠊරᗜ┴ෆࡢ 4187 㞟ⴠ࡬ࡢ࢔ࣥࢣ࣮ࢺ
ㄪᰝࠊ⊁⊟⪅࢔ࣥࢣ࣮ࢺࠊ࠾ࡼࡧ᭷ᐖᤕ⋓ᩘࢹ࣮ࢱ࡞࡝ࢆ⏝࠸࡚ㄪ࡭ࡓࠋ࣭ ࢖ࣀࢩࢩࡣࠊ㜰⚄ἢᓊ㒊ࡢ୍㒊ࢆ㝖ࡁ┴ෆ඲ᇦ࡟⏕ᜥࡋ࡚࠸ࡓࠋす᧛☻ࠊᮾ᧛☻ࠊ⚄ᡞࠊ㜰⚄ࠊῐ㊰ᓥ࡛ࡣ⏕ᜥᐦᗘࡀ 10 㢌㸭km2 ࢆ㉸࠼ࡿ㧗ᐦᗘᆅᇦࡀከ࠿ࡗࡓࠋࡉࡽ࡟ࠊす᧛☻ࠊᮾ᧛☻ࠊ⚄ᡞࠊῐ㊰ᓥ༡㒊࡛ࡣࠊ2008 ᖺ࠿ࡽ 2012 ᖺࡢ 5 ᖺ㛫࡟⏕ᜥᐦᗘࡀቑຍࡋ࡚࠸ࡿᆅᇦࡀከ࠿ࡗࡓࠋ≉࡟ࠊῐ㊰ᓥ༡㒊࡛ࡣ࡯࡜ࢇ࡝ࡢᆅᇦ࡛
20㸣௨ୖቑຍࡋ࡚࠸ࡓࠋ࣭ ࢖ࣀࢩࢩ࡟ࡼࡿ㎰ᴗ⿕ᐖࡣࠊㄪᰝᮇ㛫ࢆ㏻ࡌ࡚⥅⥆ⓗ࣭ᗈ⠊ᅖ࡟Ώࡗ࡚࠸ࡓࠋ≉࡟ࠊణ㤿ࠊ᧛☻໭㒊ࠊῐ㊰ᓥ࡛ࡣ῝้࡞⿕ᐖࢆཬࡰࡋ࡚࠸ࡓࠋࡲࡓࠊ2008 ᖺ࠿ࡽ 2012ᖺ࡟࠿ࡅ࡚ࠊරᗜ┴ᮏᕞ㒊ࡢ༡ഃࡢᆅᇦࠊ࠾ࡼࡧῐ㊰ᓥࡢ࡯࡜ࢇ࡝ࡢᆅᇦ࡛⿕ᐖ⛬ᗘࡀ῝้໬ࡋ࡚࠸ࡓࠋ

࣭ ㎰ᴗ⿕ᐖࡀ῝้࡜ᅇ⟅ࡋࡓ㞟ⴠ࡯࡝ᰙࠊᤕ⋓࡞࡝ࡢᑐ⟇ࢆ⾜ࡗ࡚࠸ࡓࠋ⿕ᐖࡀ῝้࡜ᅇ⟅ࡋࡓ㞟ⴠ࡛ࡣࠊ90㸣௨ୖࡀᰙࠊࡲࡓࡣᤕ⋓ࢆ⾜ࡗ࡚࠾ࡾࠊ4 ๭௨ୖࡢ㞟ⴠࡀᰙ࡜ᤕ⋓ࡢ୧᪉ࡢᑐ⟇ࢆ⾜ࡗ࡚࠸ࡓࠋ

࣭ 2010 ᖺ௨㝆ࠊᮏᕞ㒊ࠊῐ㊰ᓥ࡜ࡶ࡟ᤕ⋓ᩘࡢቑຍࡋࡓᆅᇦࡀከ࠿ࡗࡓࠋ≉࡟ࠊῐ㊰ᓥ໭㒊ࠊ୹Ἴࠊణ㤿ࠊᮾ᧛ᆅᇦ࡛ࡣᤕ⋓ᩘࡀቑຍࡋࡓᆅᇦࡀከ࠿ࡗࡓࠋ࣭ 㜵ㆤᰙࡣࠊ࡯࡜ࢇ࡝ࡢ㞟ⴠ㸦73.2㸣㹼87.9㸣௨ୖ㸧࡛ຠᯝࡀ࠶ࡿ࡜ᅇ⟅ࡋࡓࠋࡓࡔ
ࡋࠊຠᯝࡀ࠶ࡿ࡜⟅࠼ࡓ㞟ⴠࡢ๭ྜࡣ 2006 ᖺ௨㝆ᚎࠎ࡟పୗഴྥ࡟࠶ࡗࡓࠋ୍᪉ࠊ
ᤕ⋓ࡢຠᯝࡀ࠶ࡿ࡜ᅇ⟅ࡋࡓ㞟ⴠࡣࠊ2006 ᖺ࡟ࡣ 3 ศࡢ 1 ⛬ᗘ㸦35.4㸣㸧ࡔࡗࡓࡀࠊ
2011 ᖺ௨㝆ቑຍࡋࠊ2012 ᖺ࡟ࡣ࡯ࡰ༙ᩘ㸦47.0㸣㸧ࡢ㞟ⴠ࡛ຠᯝࡀ࠶ࡿ࡜ᅇ⟅ࡋࡓࠋ

Key words 㸸㔝⏕ື≀⟶⌮ࠊ⋇ᐖࠊ⿕ᐖᑐ⟇ࠊಖㆤ⟶⌮ィ⏬ࠊ⿕ᐖ㜵Ṇィ⏬ࠊ Sus scrofa
leucomystaxරᗜ┴᳃ᯘື≀◊✲ࢭࣥࢱ࣮࣭ රᗜ┴❧኱Ꮫ⮬↛࣭⎔ቃ⛉Ꮫ◊✲ᡤ࣭ ᰴ㸧㔝⏕ື≀ಖㆤ⟶⌮஦ົᡤ රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-2

ࡣࡌࡵ࡟
ࢽ࣍ࣥ࢖ࣀࢩࢩ㸦Sus scrofa leucomystax㸹௨ୗ࢖ࣀࢩࢩ࡜⾲グ㸧࡟ࡼࡿ㎰ᴗ⿕ᐖࡣࠊᖺࠎ῝้໬ࡋ࡚࠾ࡾࠊ2010 ᖺᗘ࡟ࡣ඲ᅜ࡛࠾ࡼࡑ 68 ൨෇࡟ࡶୖࡗ࡚࠸ࡿ
㸦㎰Ỉ┬ 2012㸧ࠋරᗜ┴࡟࠾࠸࡚ࡶࠊ࢖ࣀࢩࢩ࡟ࡼࡿ㎰ᴗ⿕ᐖ㢠ࡣࠊ㧗࠸Ỉ‽࡛᥎⛣ࡋ㸦රᗜ┴ 2012㸧ࠊ
2012 ᖺᗘ࡟ࡣ 2 ൨ 3 ༓୓෇௨ୖ࡟ୖࡾࠊ㎰స≀࡟ࡶࡗ࡜ࡶ῝้࡞⿕ᐖࢆཬࡰࡍ㔝⏕ື≀࡜࡞ࡗ࡚࠸ࡿࠋ࢖ࣀࢩࢩ࡟ࡼࡿ㎰ᴗ⿕ᐖ࡟ᑐࡋ࡚ຠ⋡ⓗ࡞ᑐ⟇ࢆ࡜ࡿࡓࡵ࡟ࡣࠊ⿕ᐖ≧ἣࡸᑐ⟇ᐇ᪋≧ἣࠊ࠾ࡼࡧᐇ᪋ࡋࡓᑐ⟇ࡢຠᯝ࡞࡝ࢆ┴ෆ඲ᇦ࠿ࡽ⥙⨶ⓗ࡟ᢕᥱࡍࡿᚲせࡀ࠶ࡿࠋࡉࡽ࡟ࠊ࢖ࣀࢩࢩࡢ⏕ᜥ≧ἣࡣࠊྠࡌᕷ⏫ᮧෆ࡛ࡶ኱ࡁࡃ␗࡞ࡿୖ࡟ࠊ㜵ㆤᰙࢆࡣࡌࡵ࡜ࡋࡓ⿕ᐖᑐ⟇ࡣ㞟ⴠ࡟ࡼࡗ࡚ࡶ␗࡞ࡿࠋࡑࡢࡓࡵ⿕ᐖ≧ἣࡣ㞟ⴠ࡛኱ࡁࡃ␗࡞ࡿሙྜࡀ࠶ࡿࠋ㒔㐨ᗓ┴ࡸᕷ⏫ᮧ࡛ࣞ࣋ࣝࠊ⿕ᐖᑐ⟇ࡸಶయᩘ⟶⌮ࡢィ⏬ࢆ⟇ᐃࡍࡿ࠺࠼࡛ࡶࠊ㞟ⴠ༢఩࡛ࡢ᝟ሗࢆ㞟⣙ࡍࡿࡇ࡜ࡀ࡛ࡁࢀࡤࠊࡼࡾຠᯝⓗ࡞⌧≧ᢕᥱࡀ࡛ࡁࡿࠋරᗜ┴࡛ࡣࠊ⿕ᐖᑐ⟇ࡸ㔝⏕ື≀⟶⌮࡟ᚲせ࡞᝟ሗࢆ㞟ⴠ༢఩࡛཰㞟ࡍࡿࡓࡵࠊ2003
ᖺᗘࡼࡾ㎰ᴗ㞟ⴠࡈ࡜࡟ࠊ㔝⏕㫽⋇࡟ࡼࡿ㎰ᴗ⿕ᐖࡢ⛬ᗘࡸኚ໬ࠊᑐ⟇ࡢᐇ᪋≧ἣ࡞࡝ࢆᅇ⟅ࡋ࡚ࡶࡽ࠺࢔ࣥࢣ࣮ࢺㄪᰝࢆᐇ᪋ࡋ࡚ࡁࡓࠋᮏㄽᩥ࡛ࡣࠊරᗜ┴ୗࡢ࢖ࣀࢩࢩࡢ
⏕ᜥ࣭⿕ᐖ≧ἣࢆ㞟ⴠ༢఩࡛ᢕᥱࡋࠊຠ⋡ⓗ࡞㔝⏕ື≀⟶⌮⟇ࡢ⟇ᐃ࡜ពᛮỴᐃ࡟㈨ࡍ
ࡿࡓࡵ࡟ࠊ࢖ࣀࢩࢩࡢ⏕ᜥศᕸࠊ㎰ᴗ⿕ᐖ࣭ᑐ⟇≧ἣࠊ࠾ࡼࡧᑐ⟇ࡢຠᯝࢆࠊරᗜ┴ෆ
ࡢ 4187 㞟ⴠࢆᑐ㇟࡟ࡋࡓ࢔ࣥࢣ࣮ࢺㄪᰝࠊ⊁⊟⪅࢔ࣥࢣ࣮ࢺࠊ࠾ࡼࡧරᗜ┴ࡀ཰㞟ࡋࡓᤕ⋓ᩘࢹ࣮ࢱ࡟ࡼࡾ᫂ࡽ࠿࡟ࡋࡓ⤖ᯝ࡟ࡘ࠸࡚㏙࡭ࡿࠋ᪉ἲ㸧 රᗜ┴ࡢ NP ࣓ࢵࢩࣗᙜࡓࡾࡢ࢖ࣀࢩࢩ⏕ᜥᐦᗘ᥎ᐃ࡜ᐦᗘኚ໬
⏝࠸ࡓࢹ࣮ࢱ
᭷ᐖᤕ⋓ᩘ㸦ᖺᗘ㸧 i_yugai[i,j]㸸i ᖺᗘ j ࣓ࢵࢩ࡛ࣗࡢ᭷ᐖチྍ࡟ࡼࡿᤕ⋓ᩘࠋಶయᩘ
ࢆ཯ᫎࡍࡿᣦᶆ࡜ࡋ࡚⏝࠸ࡿࠋ᭷ᐖᤕ⋓ᩘࡣᕷ⏫ᮧࡈ࡜࡟ሗ࿌ࡉࢀࡿࡓࡵࠊྛᕷ⏫
ᮧࡢ᭷ᐖᤕ⋓ᩘࢆ࣓ࢵࢩࣗෆࡢ᳃ᯘ㠃✚࡟ᛂࡌ࡚ྛ࣓ࢵࢩࣗ࡟㓄ศࡋࠊྛ࣓ࢵࢩࣗ
ᙜࡓࡾࡢ᭷ᐖᤕ⋓ᩘ࡜ࡋࡓࠋ
⊁⊟ᤕ⋓ᩘ r_ca[i,j]㸸i ᖺᗘ j ࣓ࢵࢩࣗ࡟࠾ࡅࡿ⊁⊟࡟ࡼࡿᤕ⋓ᩘࠋරᗜ┴࡛ࡣ⊁⊟⪅Ⓩ㘓⪅඲ဨࢆᑐ㇟࡜ࡋࡓ࢔ࣥࢣ࣮ࢺㄪᰝࢆᐇ᪋ࡋ࡚࠸ࡿࠋࢩ࣭࢝࢖ࣀࢩࢩ⊟ࢆᑐ㇟࡜ࡋࡓ㖠⊟ฟ⊟ࡢᗘࠊฟ⊟᪥ࠊ࣓ࢵࢩࣗ␒ྕࠊྠ⾜⪅ᩘࠊ┠ᧁᩘࠊᤕ⋓ᩘࢆࠊࢃ࡞
⊟ࡢሙྜࡣࠊᤕ⋓᪥ࠊ࣓ࢵࢩࣗ␒ྕࠊᤕ⋓ᩘࢆ࢔ࣥࢣ࣮ࢺ⏝⣬࡟グ㘓ࡋ࡚ࡶࡽࡗ࡚
࠸ࡿࠋ⊁⊟ᤕ⋓ᩘࡣࠊ⊁⊟⪅࢔ࣥࢣ࣮ࢺࢆ㞟ィࡋࡓࠋಶయᩘࢆ཯ᫎࡍࡿᣦᶆ࡜ࡋ࡚
⏝࠸ࡿࠋ┠ᧁຠ⋡ spue[i,j]㸸i ᖺᗘࡢ⊁⊟ᮇ㛫୰࡟ j ࣓ࢵࢩࣗ࡟࠾࠸࡚ࠊ⊁⊟⪅Ⓩ㘓⪅࠿ࡽᚓࡽࢀࡓ㖠⊟᫬ࡢ┠ᧁຠ⋡ࠋಶయᩘࢆ཯ᫎࡍࡿᣦᶆ࡜ࡋ࡚⏝࠸ࡿࠋ රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-2

᳃ᯘ㠃✚ f_area[j] 㸸j ࣓ࢵࢩࣗࡢ᳃ᯘ㠃✚ࠋྛᕷ⏫ࡢ᭷ᐖᤕ⋓ᩘࢆ࣓ࢵࢩࣗᙜࡓࡾࡢ
᭷ᐖᤕ⋓ᩘ࡟ኚ᥮ࡍࡿ㝿ࠊ࠾ࡼࡧ⏕ᜥᐦᗘࡢᮇᚅ್ࢆィ⟬ࡍࡿ㝿࡟⏝࠸ࡿࠋ᥎ᐃ᪉ἲ➨㸱❶࡛᥎ᐃࡋࡓරᗜ┴ᮏᕞ㒊࣭ῐ㊰ᓥࡢಶయᩘࠊ࠾ࡼࡧ 5km ࣓ࢵࢩࣗ༢఩ࡢ┠ᧁຠ⋡ࠊ᭷ᐖ࣭⊁⊟ᤕ⋓ᩘࡢ 3 ࡘࡢᣦᶆࢆᇶ࡟ࠊ2002 ᖺ࠿ࡽ 2012 ᖺࡲ࡛ࡢ 5km ࣓ࢵࢩࣗ༢఩ࡢ࢖ࣀࢩࢩᐦᗘࢆ᥎ᐃࡋࡓࠋᣦᶆࡈ࡜࡟ࠊᮏᕞ㒊࣭ῐ㊰ᓥࡢྜィ್࡟ᑐࡍࡿྛ࣓
ࢵࢩࣗࡢ್ࡢẚࢆ⟬ฟࡋࠊྛᣦᶆࡢ࣓ࢵࢩࣗ༢఩ࡢኚືಀᩘ࡛⿵ṇࡋࡓࠋරᗜ┴ᮏᕞ㒊࣭ῐ㊰ᓥࡢྜィ್࡟ᑐࡍࡿྛ࣓ࢵࢩࣗࡢ್ࡢẚࢆ࣓ࢵࢩࣗࡈ࡜࡟ྜィࡋࠊ3 ࡘࡢᣦᶆࡢྜィ್㸦Ai㸧࡜࣓ࢵࢩࣗᙜࡓࡾࡢಶయᩘࡀẚ౛ࡍࡿࡼ࠺࡟ࠊරᗜ┴ᮏᕞ㒊࣭ῐ㊰ᓥࡢಶయᩘࢆྛ࣓ࢵࢩࣗ࡟㓄ศࡋࡓࠋ⏕ᜥᐦᗘࡣࠊ࣓ࢵࢩࣗࡈ࡜ࡢಶయᩘࢆྛ࣓ࢵࢩࣗᙜࡓ
ࡾࡢ᳃ᯘ㠃✚࡛๭ࡗ࡚⟬ฟࡋࡓࠋ
Ai = i ࣓ࢵࢩ࡛ࣗࡢ┠ᧁຠ⋡/㸦඲࣓ࢵࢩ࡛ࣗࡢ┠ᧁຠ⋡ࡢྜィ್㸨┠ᧁຠ⋡ࡢኚືಀᩘ㸧+ i ࣓ࢵࢩ࡛ࣗࡢ⊁⊟ᤕ⋓ᩘ/㸦඲࣓ࢵࢩ࡛ࣗࡢ⊁⊟ᤕ⋓ᩘࡢྜィ್㸨⊁⊟ᤕ⋓ᩘࡢኚືಀᩘ㸧+i ࣓ࢵࢩ࡛ࣗࡢ᭷ᐖᤕ⋓ᩘ/㸦඲࣓ࢵࢩ࡛ࣗࡢ᭷ᐖᤕ⋓ᩘࡢྜィ್㸨᭷ᐖᤕ⋓ᩘࡢኚືಀᩘ㸧Ai㸸ྛ࣓ࢵࢩࣗ࡟ᮏᕞ㒊࣭ῐ㊰ᓥࡢ࢖ࣀࢩࢩࡢ᥎ᐃಶయᩘࢆ㓄ศࡍࡿࡓࡵࡢᣦᩘ㸧 ࢖ࣀࢩࢩ࡟ࡼࡿ㎰ᴗ⿕ᐖࡢ⛬ᗘ࡜ࡑࡢኚ໬
㎰ᴗ㞟ⴠ࿘㎶࡟࠾ࡅࡿ㫽⋇ᐖࠊ㫽⋇ᐖᑐ⟇ࡢ≧ἣࠊ࠾ࡼࡧᑐ⟇ࡢຠᯝ࡟ࡘ࠸࡚ࠊࠕ㎰ᯘᴗࢭࣥࢧࢫࠖࢆᇶᮏ࡜ࡋࡓ 4187㸦2012 ᖺᗘ㸧ࡢ㎰ᴗ㞟ⴠࢆᑐ㇟࡟ࠊ2003 ᖺ࠿ࡽ 2012ᖺࡢ 10 ᖺ㛫࡟ࢃࡓࡗ࡚࢔ࣥࢣ࣮ࢺㄪᰝ㸦௨ୗࠊ㫽⋇ᐖ࢔ࣥࢣ࣮ࢺ㸧ࢆ⾜ࡗࡓࠋ࢖ࣀࢩ
ࢩࡢ⿕ᐖ≧ἣ࡟㛵ࡍࡿㄪᰝ㡯┠ࡣ௨ୗࡢ࡜࠾ࡾ࡛࠶ࡿࠋ
ᑐ㇟ື≀㸦࢖ࣀࢩࢩ㸧‫ࡿ࠸ڧ‬Ѝ㸦‫ڧ‬㏻ᖺ ‫ڧ‬᫓㹼⛅ࡔࡅ ‫ڧ‬୙᫂㸧‫ ࠸࡞࠸ڧ‬㸦ୗグࡣグධ୙せ㸧㎰ᴗ⿕ᐖࡣ‫࠸࡞࡝ࢇ࡜࡯ڧ‬‫ڧ‬㍍ᚤ
‫ڧ‬኱ࡁ࠸‫้῝ڧ‬
 ㎰ᴗ⿕ᐖ⛬ᗘࡢศᕸ≧ἣࢆ✵㛫ⓗ࡟ᢕᥱࡍࡿࡓࡵ࡟ࠊ⿕ᐖᆅ⛬ᗘࡣ㞟ⴠ༢఩࡛ᚓࡽࢀ
ࡓ⏕ᜥࡢ᭷↓࡜㎰ᴗ⿕ᐖࡢ⛬ᗘ࠿ࡽ 0 ࠿ࡽ 4 ࡲ࡛ࡢᩘ್㸦0㸸࠸࡞࠸ࠊ1㸸࡯࡜ࢇ࡝࡞
࠸ࠊ2㸸㍍ᚤࠊ3㸸኱ࡁ࠸ࠊ4㸸῝้㸧࡟ኚ᥮ࡋࠊArcGIS㸦Esri ♫㸧ࡢ Spatial Analyst
ࡢ IDW㸦㏫㊥㞳ຍ㔜ἲ㸧࡟ࡼࡾ᥎ᐃࡋࡓࠋ࡞࠾ࠊෆᤄฎ⌮࡟฼⏝ࡍࡿ࣏࢖ࣥࢺࡲ࡛ࡢ
㊥㞳ࡣ 5km ࡜ࡋࡓࠋࡉࡽ࡟ࠊ2008 ᖺ࡜ 2012 ᖺࡢ㎰ᴗ⿕ᐖ⛬ᗘࡢᩘ್ࢹ࣮ࢱࢆ⏝࠸࡚ රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-2

᭱㏆ 5 ᖺ㛫࡛ࡢ㎰ᴗ⿕ᐖ⛬ᗘࡢኚ໬ࢆ⟬ฟࡋࠊArcGIS㸦Esri ♫㸧ࡢ Spatial Analyst
ࡢ IDW㸦㏫㊥㞳ຍ㔜ἲ㸧࡟ࡼࡾ㎰ᴗ⿕ᐖ⛬ᗘࡢศᕸ≧ἣࢆ᥎ᐃࡋࡓࠋ㎰ᴗ⿕ᐖ⛬ᗘࡢኚ໬࡟ࡘ࠸࡚ࡶෆᤄฎ⌮࡟฼⏝ࡍࡿ࣏࢖ࣥࢺࡲ࡛ࡢ㊥㞳ࡣ 5km ࡜ࡋࡓࠋ㸧 㞟ⴠ࡟࠾ࡅࡿ⿕ᐖᑐ⟇࡜ࡑࡢຠᯝ
㞟ⴠ࡟࠾ࡅࡿ⿕ᐖࡢ⛬ᗘ࡜ᑐ⟇ᐇ᪋≧ἣࡢ㛵ಀࠊ࠾ࡼࡧ⿕ᐖᑐ⟇࡜ࡑࡢຠᯝࢆ᫂ࡽ࠿
࡟ࡍࡿࡓࡵࠊୖグࡢ㫽⋇ᐖ࢔ࣥࢣ࣮ࢺ࡟࠾࠸࡚ࡇࢀࡲ࡛⋇ᐖᑐ⟇࡜ࡋ࡚ᐇ᪋ࡋࡓᑐ⟇ࢆᑜࡡࡓࠋᑐ⟇ᐇ᪋≧ἣ࡟㛵ࡍࡿㄪᰝ㡯┠ࡣ௨ୗࡢ࡜࠾ࡾ࡛࠶ࡿࠋ
ᤕ⋓    ‫ڧ ↓ڧ‬᭷㸦‫ڧڧڧ‬㢌ࡄࡽ࠸ ‫ڧ‬㢌ᩘ୙᫂㸧            Ѝຠᯝ㸦‫ڧ ↓ڧ‬᭷ ‫ڧ‬୙᫂㸧㜵ㆤᰙ‫ڧ ↓ڧ‬᭷㸦‫ڧ ࢺࢵࢿڧ‬㟁Ẽᰙ      ‫ڧ‬㔠⥙ᰙ ‫ࣥࢱࢺڧ‬ᰙ㸧   Ѝຠᯝ㸦‫ڧ ↓ڧ‬᭷ ‫ڧ‬୙᫂㸧㸧 ⊁⊟ᮇ㛫ᘏ㛗ࡢຠᯝ⿕ᐖᑐ⟇࡜ࡋ࡚ࡢᤕ⋓ࢆᙉ໬ࡍࡿࡓࡵࠊ2010 ᖺ࠿ࡽ⊁⊟ᮇ㛫ࢆ㸯࠿᭶㛫ᘏ㛗ࡋࠊ11
᭶ 15 ᪥࠿ࡽ 3 ᭶ 15 ᪥ࡲ࡛࡜ࡋࡓࠋ
⊟ᮇࡢᘏ㛗ࡀᤕ⋓ᣑ኱࡟ཬࡰࡍᙳ㡪ࢆㄪ࡭ࡿࡓࡵࠊ
⊟ᮇࡢᤕ⋓ᩘ㸦㖠⊟ࡢࡳ㸧ࢆ㏻ᖖᮇ㛫࡜ᘏ㛗ᮇ㛫࡟ࢃࡅ࡚㞟ィࡋࡓࠋࡉࡽ࡟㏻ᖖᮇ㛫࡜ᘏ㛗ᮇ㛫ࡢ㖠⊟࡟ࡼࡿᤕ⋓ຠ⋡ࢆㄪ࡭ࡓࠋᤕ⋓ᮇ㛫ูࡢᤕ⋓ᩘࠊ࠾ࡼࡧᤕ⋓ຠ⋡ࡣࠊ⊁⊟⪅࢔ࣥࢣ࣮ࢺ࠿ࡽ㞟ィࡋࡓࠋᤕ⋓ຠ⋡ࡣᤕ⋓ᩘࢆᘏ࡭ྠ⾜⪅ᩘ࣭ฟ⊟᪥ᩘ࡛㝖ࡋ࡚⟬ฟࡋࡓࠋ㸧 ᤕ⋓≧ἣරᗜ┴࡟࠾ࡅࡿ࢖ࣀࢩࢩࡢᤕ⋓≧ἣࢆ✵㛫ⓗࠊ᫬㛫ⓗ࡟ᢕᥱࡍࡿࡓࡵࠊ1㸧5km ࣓ࢵࢩࣗᙜࡓࡾࡢ⏕ᜥᐦᗘ᥎ᐃࠊ࡛⏝࠸ࡓ 2002 ᖺ࠿ࡽ 2013 ᖺࡲ࡛ࡢ࣓ࢵࢩࣗᙜࡓࡾࡢᤕ
⋓ᩘ㸦⊁⊟࣭᭷ᐖู㸧ࡢࢹ࣮ࢱࢆ⏝࠸࡚ࠊ࣓ࢵࢩࣗᙜࡓࡾᤕ⋓ᩘࡢศᕸ࡜᫬㛫ⓗኚືࢆㄪ࡭ࡓࠋࡉࡽ࡟ࠊරᗜ┴඲యࡢᤕ⋓ືྥᢕᥱࡢࡓࡵࠊරᗜ┴඲యࡢᤕ⋓ᩘ㸦᭷ᐖ࣭⊁⊟ู㸧ࡢᖺḟኚືࡶㄪ࡭ࡓࠋ
⤖ᯝ࡜⪃ᐹ㸧 රᗜ┴ࡢ NP ࣓ࢵࢩࣗᙜࡓࡾࡢ࢖ࣀࢩࢩ⏕ᜥᐦᗘ᥎ᐃ࡜ᐦᗘኚ໬
㜰⚄ἢᓊࡢࡈࡃ୍㒊ࢆ㝖ࡁࠊ࢖ࣀࢩࢩࡣරᗜ┴඲ᇦ࡟⏕ᜥࡋ࡚࠸ࡓ㸦ᅗ㸯㸧
ࠋ10 㢌㸭
ᖹ᪉ km ௨ୖࡢ㧗ᐦᗘᆅᇦࡣࠊ2002 ᖺࡢ᫬Ⅼ࡛ࡣῐ㊰ᓥࡢ༡໭➃࡜㜰⚄ࠊᮾ᧛ࠊణ㤿ᆅᇦࡢࡈࡃ୍㒊࡟㝈ࡽࢀ࡚࠸ࡓࡀࠊᚎࠎ࡟ᗈࡀࡾ 2010 ᖺ࡟ࡣࠊῐ㊰ᓥࡢ኱༙ࠊణ㤿ࠊ᧛☻ࠊ୹Ἴࠊ㜰⚄ࠊ⚄ᡞᆅᇦ࡟ࡶᗈࡀࡗࡓࠋ2011 ᖺ௨㝆㧗ᐦᗘᆅᇦࡣᚤῶഴྥࡔࡗࡓࠋ රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-2

2008 ᖺ࠿ࡽ 2012 ᖺࡢኚ໬⋡ࡣࠊᮏᕞ㒊ࡢ୰㒊ࡸ໭㒊ࡣῶᑡࡋ࡚࠸ࡿᆅᇦࡀከ࠿ࡗࡓ
ࡀࠊᮏᕞ㒊༡㒊ࡸῐ㊰ᓥࡣቑຍࡋ࡚࠸ࡿᆅᇦࡀ࡯࡜ࢇ࡝࡛࠶ࡗࡓ㸦ᅗ㸰㸧ࠋ㸧 ࢖ࣀࢩࢩ࡟ࡼࡿ㎰ᴗ⿕ᐖࡢ⛬ᗘ࡜ࡑࡢኚ໬
࢖ࣀࢩࢩ࡟ࡼࡿ㎰ᴗ⿕ᐖࡀ࡞࠸࡜ᅇ⟅ࡋࡓ㞟ⴠࡣ 28.9㸣㸦2003 ᖺ㸧㹼35.2㸣㸦2007
ᖺ㸧࡛ࠊ㎰ᆅࡢ࡯࡜ࢇ࡝࡞࠸㜰⚄ᆅᇦࢆ㝖ࡁ┴ୗࡢ኱༙ࡢᆅᇦ࡛࢖ࣀࢩࢩ࡟ࡼࡿ⿕ᐖࡀሗ࿌ࡉࢀࡓ㸦ᅗ㸱㸧ࠋ2008 ᖺ࠿ࡽ 2012 ᖺࡢ⿕ᐖ⛬ᗘࡀ῝้໬ࡋ࡚࠸ࡿᆅᇦࡣࠊ࡯ࡰᐦ
ᗘࡀቑຍࡋ࡚࠸ࡿᆅᇦ࡜㔜࡞ࡗ࡚࠾ࡾࠊῐ㊰ᓥ඲ᇦࠊᮏᕞ㒊༡㒊ࠊす᧛໭㒊ࠊ୰᧛໭㒊࡛ࡣ≉࡟⿕ᐖࡀ῝้໬ࡋ࡚࠸ࡿᆅᇦࡀከ࠿ࡗࡓ㸦ᅗ㸲㸧ࠋ㸧 㞟ⴠ࡟࠾ࡅࡿ⿕ᐖᑐ⟇࡜ࡑࡢຠᯝ
⿕ᐖᑐ⟇࡜ࡋ࡚㜵ㆤᰙࡢタ⨨࡜ᤕ⋓ࡢ≧ἣࢆ⿕ᐖ⛬ᗘ࡟ࡼࡗ࡚ศ㢮ࡋࠊ㞟ィࡋࡓ㸦ᅗ㸳㸧ࠋᆏ⏣㸦2012㸧ྠᵝࠊ⿕ᐖ⛬ᗘࡀ῝้࡞㞟ⴠ࡯࡝ఱࡽ࠿ࡢᑐ⟇ࢆᐇ᪋ࡋ࡚࠸ࡓࠋ⿕ᐖࡀ῝้࡜ᅇ⟅ࡋࡓ㞟ⴠ࡛ࡣࠊ70.0㸣ࡀᰙࠊࡲࡓࡣᤕ⋓ࢆ⾜࠸ࠊ32.3㸣ࡀᰙ࡜ᤕ⋓ࡢ୧᪉ࡢᑐ⟇ࢆ⾜ࡗ࡚࠸ࡓࠋ୍᪉ࠊ⿕ᐖࡀ࡯࡜ࢇ࡝࡞࠸࡜ᅇ⟅ࡋࡓ㞟ⴠ࡛ࡣࠊᰙࠊࡲࡓࡣᤕ
⋓ࢆ⾜ࡗ࡚࠸ࡿ㞟ⴠࡣ 41.1㸣࡟࡜࡝ࡲࡗࡓࠋᑐ⟇ᡭẁࡢ࠺ࡕࠊ㜵ㆤᰙࢆタ⨨ࡋ࡚࠸ࡿ㞟ⴠࡣࠊ⿕ᐖࡀ῝้ࠊࡲࡓࡣ኱ࡁ࠸࡜ᅇ⟅ࡋࡓ㞟ⴠࡢ࠺ࡕ 60㸣௨ୖࢆ㉸࠼ࡓࠋ୍᪉ࠊ
ᤕ⋓ࢆᐇ᪋ࡋ࡚࠸ࡿ㞟ⴠࡣࠊ⿕ᐖࡀ῝้࡜ᅇ⟅ࡋࡓ㞟ⴠࡢ࡞࠿࡛ࡶ 37.3㸣࡟࡜࡝ࡲࡗ
ࡓࠋ㜵ㆤᰙࡢຠᯝࡣࠊ77.2㸣ࡢ㞟ⴠ࡛ㄆࡵࡽࢀ࡚࠸ࡓ㸦ᅗ㸴㸧ࠋࡓࡔࡋࠊᰙࡢຠᯝࡀ࠶ࡿ
࡜ᅇ⟅ࡋࡓ㞟ⴠࡢ๭ྜࡣࠊ2006 ᖺࡢ 87.9㸣࠿ࡽ 2012 ᖺࡢ 75.8㸣ࡲ࡛ᚎࠎ࡟ῶᑡഴྥ
࡟࠶ࡗࡓࠋຠᯝࡀ࠶ࡿ࡜ᅇ⟅ࡋࡓ㞟ⴠᩘࡣ࠶ࡲࡾኚࢃࡽ࡞࠿ࡗࡓࡀࠊຠᯝࡀ࡞࠸࡜ᅇ⟅ࡋࡓ㞟ⴠᩘࡀࡸࡸቑຍࡋࡓࡓࡵࠊຠᯝࡀ࠶ࡿ࡜ᅇ⟅ࡋࡓ㞟ⴠࡢ๭ྜࡀࡸࡸῶᑡࡋࡓ࡜⪃࠼ࡽࢀࡿࠋ୍᪉ࠊᤕ⋓ࡢຠᯝࡣࠊᰙ࡟ẚ࡭࠶ࡲࡾㄆࡵࡽࢀ࡚࠸࡞࠸ࠋࡋ࠿ࡋࠊᤕ⋓ࡢຠᯝࢆㄆࡵࡿ㞟ⴠᩘࡣᚎࠎ࡟ቑຍࡋࠊ2006 ᖺࡢ 35.4㸣࠿ࡽ 2012 ᖺࡢ 47.0㸣࡜ቑຍഴྥ
࡟࠶ࡗࡓ㸦ᅗ㸵㸧ࠋ≉࡟ࠊ2011 ᖺ௨㝆ࡢᤕ⋓ᣑ኱௨㝆ࠊᤕ⋓ࡢຠᯝࢆㄆࡵࡿ㞟ⴠᩘࡀᛴ㏿࡟ቑຍࡋࡓࠋ㸧 ⊁⊟ᮇ㛫ᘏ㛗ࡢຠᯝᘏ㛗ᮇ㛫ࡢᤕ⋓ຠ⋡ࡣ 3 ᖺ࡜ࡶ㏻ᖖᮇ㛫ࡢᤕ⋓ຠ⋡ࡼࡾࡶࡸࡸప࠸ഴྥ࡟࠶ࡗࡓ㸦ᅗ㸶㸧ࠋࡉࡽ࡟ࠊᤕ⋓ᩘࡣࠊ2010 ᖺ࡟ࡣቑຍࡋࡓࡀ௨㝆ῶᑡࡋࡓࠋࡋ࠿ࡋࠊ㏻ᖖᮇ㛫ࠊᘏ㛗ᮇ㛫࡜ࡶ࡟ᤕ⋓ຠ⋡ࡣ 2010 ᖺ௨㝆ῶᑡࡋ࡚࠾ࡾࠊ⊟ᮇࡢᘏ㛗ࡀ࡞ࡅࢀࡤᤕ⋓ᩘࡣࠊῶᑡࡋ࡚࠸ࡓ࡜⪃࠼ࡽࢀࡿࠋࡋࡓࡀࡗ࡚ࠊ⊟ᮇࡢᘏ㛗ࡣࠊᤕ⋓ຠ⋡ࡢపୗ࡟క࠺ᤕ⋓ᩘࡢῶᑡ⛬ᗘࢆᢚ࠼ࡿ࡜࠸࠺ព࿡࡛ຠᯝࡀ࠶ࡗࡓ࡜⪃࠼ࡽࢀࡿࠋ㸧 ᤕ⋓≧ἣ᭷ᐖ㥑㝖࡜⊁⊟ࢆྜࢃࡏࡓ⥲ᤕ⋓ᩘࡣࠊᖺ࡟ࡼࡾኚືࡀ࠶ࡿࡶࡢࡢ 2010 ᖺࡲ࡛ࡣቑ රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-2

ຍഴྥ࡟࠶ࡗࡓ㸦ᅗ㸷㸧ࠋ࡜ࡃ࡟ࠊ᭷ᐖᤕ⋓ᩘࡣࠊ2005 ᖺࢆ㝖ࡁ༢ㄪ࡟ቑຍࡋࡓࠋ⥲ᤕ⋓ᩘࡢ࠺ࡕ᭷ᐖᤕ⋓ᩘࡀ༨ࡵࡿ๭ྜࡣࠊ2002 ᖺ࡟ࡣ 25.0㸣ࡔࡗࡓࡢࡀ 2012 ᖺ࡟ࡣ
51.3㸣ࡲ࡛ୖ᪼ࡋࡓࠋ2010 ᖺ௨㝆ᤕ⋓ᩘࡣῶᑡࡋࡓࠋᤕ⋓ᩘࡣࠊ2010 ᖺ௨㝆ࠊ⏕ᜥᐦᗘࡢ㧗࠸ణ㤿ࠊᮾ᧛ࠊ୹Ἴࠊῐ㊰ᓥᆅᇦ࡛ከࡃ࡞ࡗ࡚࠸ࡓ㸦ᅗ㸯㸮㸧
ࠋ





















 



 

ᅗ  රᗜ┴ෆࡢ⊁⊟࣓ࢵࢩࣗ࠶ࡓࡾࡢ࢖ࣀࢩࢩࡢ⏕ᜥᐦᗘࡢศᕸ රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-2

ᅗ㸰 ⊁⊟࣓ࢵࢩࣗ࠶ࡓࡾࡢ࢖ࣀࢩࢩࡢ⏕ᜥᐦᗘࡢ᭱㏆㸳ᖺ㛫㸦 ᖺ㹼 ᖺ㸧ࡢኚ໬⋡ࡢศᕸ රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-2























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ᅗ㸱 㞟ⴠ࡬ࡢ࢔ࣥࢣ࣮ࢺㄪᰝ࡛᫂ࡽ࠿࡟࡞ࡗࡓ࢖ࣀࢩࢩ࡟ࡼࡿ㎰ᴗ⿕ᐖࡢศᕸ


 

රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-2

ᅗ㸲 ࢖ࣀࢩࢩ࡟ࡼࡿ㎰ᴗ⿕ᐖࡢ᭱㏆㸳ᖺ㛫㸦 ᖺ㹼 ᖺ㸧ࡢኚ໬⋡ࡢศᕸᅗ㸳 ࢖ࣀࢩࢩ࡟ࡼࡿ⿕ᐖ⛬ᗘูࡢᑐ⟇ᐇ᪋㞟ⴠࡢ๭ྜ
 ᖺ࠿ࡽ  ᖺࡲ࡛ࡢ  ᅇࡢㄪᰝ࡛⿕ᐖࡀ࠶ࡿ࡜ᅇ⟅ࡋࡓ㞟ⴠࢆࡍ࡭࡚ྵࡵ࡚㞟ィࡋࡓࠋ රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-2

ᅗ㸴 㞟ⴠ࠿ࡽࡳࡓ㜵ㆤᰙࡢຠᯝ
 ᖺ࠿ࡽ  ᖺࡲ࡛ࡢ㞟ⴠ࢔ࣥࢣ࣮ࢺㄪᰝ࡛㜵ㆤᰙࡢຠᯝ࡟ࡘ࠸࡚ᅇ⟅ࡋࡓ㞟ⴠᩘࢆ♧ࡋࡓࠋᅗ㸵 㞟ⴠ࠿ࡽࡳࡓᤕ⋓ࡢຠᯝ ᖺ࠿ࡽ  ᖺࡲ࡛ࡢ㞟ⴠ࢔ࣥࢣ࣮ࢺㄪᰝ࡛ᤕ⋓ࡢຠᯝ࡟ࡘ࠸࡚ᅇ⟅ࡋࡓ㞟ⴠᩘࢆ♧ࡋࡓࠋ රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-2

ᅗ㸶 ᖺᗘูࡢᤕ⋓ᩘ࡜ᤕ⋓ຠ⋡ ᖺ௨㝆ࡣࠊ㏻ᖖᮇ㛫㸦 ᭶  ᪥࠿ࡽ  ᭶  ᪥㸧࡜ᘏ㛗ᮇ㛫㸦 ᭶  ᪥࠿ࡽ  ᭶  ᪥㸧ࡈ࡜࡟♧ࡋࡓࠋ
⊁⊟ᤕ⋓ᩘ
᭷ᐖᤕ⋓ᩘᖺᗘᅗ㸷 ᤕ⋓ᩘ㸦⊁⊟࣭᭷ᐖู㸧ࡢᖺḟኚ໬

 



රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-2





















 





 



ᅗ㸯㸮 රᗜ┴ෆࡢ⊁⊟࣓ࢵࢩࣗ࠶ࡓࡾࡢᤕ⋓ᩘࡢศᕸ
ᘬ⏝ᩥ⊩රᗜ┴㸬2012㸬➨ 2 ᮇ࢖ࣀࢩࢩಖㆤ⟶⌮ィ⏬. රᗜ┴㸪⚄ᡞ, 18pp㸬
ᓊᮏᗣ㄃㸬2010㸬ࢩ࣭࢝࢖ࣀࢩࢩ࡟ࡼࡿ⿕ᐖࡢ≧ἣ. ࠕ㎰ᴗ㞟ⴠ࢔ࣥࢣ࣮ࢺ࠿ࡽࡳࡿࢽ࣍
ࣥࢪ࣭࢝࢖ࣀࢩࢩࡢ⿕ᐖ࡜ᑐ⟇ࡢ⌧≧ࠖ㸪 රᗜ࣡࢖ࣝࢻࣛ࢖ࣇ࣏࣮ࣞࢺ 2 ྕ㸪pp.5-15. රᗜ┴᳃ᯘື≀◊✲ࢭࣥࢱ࣮㸬ᆏ⏣ᏹᚿ㸬2010㸬ࢩ࣭࢝࢖ࣀࢩࢩࡢ⿕ᐖᑐ⟇ࡢ≧ἣ. ࠕ㎰ᴗ㞟ⴠ࢔ࣥࢣ࣮ࢺ࠿ࡽࡳࡿࢽ࣍
ࣥࢪ࣭࢝࢖ࣀࢩࢩࡢ⿕ᐖ࡜ᑐ⟇ࡢ⌧≧ࠖ㸪 රᗜ࣡࢖ࣝࢻࣛ࢖ࣇ࣏࣮ࣞࢺ 2 ྕ㸪pp.16-28. රᗜ┴᳃ᯘື≀◊✲ࢭࣥࢱ࣮㸬SAS Institute Inc. 2011. SAS/STAT® 9.3 User’s Guide. SAS Institute Inc., Cary, NC.

 රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-3➨ 㸱 ❶රᗜ┴࡟࠾ࡅࡿࢽ࣍ࣥ࢖ࣀࢩࢩࡢಶయ⩌ືែࡢ᥎ᐃ
ᯇᮏ ᓫ 1,2࣭ᆏ⏣ᏹᚿ 1,2࣭ᓊᮏᗣ㄃ 3
せ Ⅼ
࣭ 2012 ᖺࡲ࡛࡟ධᡭࡉࢀࡓࢹ࣮ࢱ࠿ࡽࠊරᗜ┴࡟⏕ᜥࡍࡿ࢖ࣀࢩࢩࡢ⮬↛ቑຍ⋡ࡸಶయᩘ࡟ࡘ࠸࡚ࠊ㝵ᒙ࣋࢖ࢬࣔࢹࣝࢆᵓ⠏ࡋࠊ࣐ࣝࢥࣇ㐃㙐ࣔࣥࢸ࢝ࣝࣟἲ࡟ࡼࡗ
࡚᥎ᐃࡋࡓࠋ࣭ ಶయᩘࡢ᥎ᐃࡣරᗜ┴ᮏᕞ㒊࡜ῐ㊰ᓥ࡟ࢃࡅ࡚⾜ࡗࡓࠋ࣭ ᥎ᐃ࡟࠶ࡓࡗ࡚ࡣࠊ㖠⊟᫬ࡢ┠ᧁຠ⋡ࠊ⊁⊟ᤕ⋓ᩘࠊ᭷ᐖᤕ⋓ᩘࡢࢹ࣮ࢱࢆᇶ࡟ࠊ
ࡑࢀࡒࢀࢹ࣮ࢱࡢㄗᕪኚືࢆ⤌ࡳ㎸ࢇࡔࣔࢹࣝࢆᵓ⠏ࡋࡓࠋ࣭ ⮬↛ቑຍ⋡ࡣࠊ୰ኸ್࡛ 18.8㸣࠿ࡽ 91.5㸣㸦ᮏᕞ㒊㸧ࠊ24.0㸣࠿ࡽ 100.3㸣㸦ῐ㊰ᓥ㸧ࡢ್ࢆ࡜ࡗࡓࠋ࣭ ᥎ᐃಶయᩘࡢ୰ኸ್ࡣࠊ2012 ᖺࡢᖺᮎࡢẁ㝵࡛ࠊᮏᕞ㒊࡛ࡣ 16,478 㢌㸦90%ಙ㢗㝈⏺࡛ࡣ 8,143㹼47,213 㢌⛬ᗘ㸧ࠊῐ㊰ᓥ࡛ࡣ 8,237 㢌㸦90%ಙ㢗㝈⏺࡛ࡣ 3,871
㹼19,161 㢌⛬ᗘ㸧ࠊ࡜᥎ᐃࡉࢀࡓࠋ

key words㸸ಶయᩘ⟶⌮ࠊ⮬↛ቑຍ⋡ࠊ࣋࢖ࢬ᥎ᐃࠊ࣐ࣝࢥࣇ㐃㙐ࣔࣥࢸ࢝ࣝࣟἲࠊಶయᩘ᥎ᐃ

㸬ࡣࡌࡵ࡟
රᗜ┴࡛ࡣࠊ≉ᐃ㫽⋇ಖㆤ⟶⌮ィ⏬࡟ᇶ࡙࠸࡚ࠊࢽ࣍ࣥ࢖ࣀࢩࢩ㸦 Sus scrofa

leucomystax㸹௨ୗ࢖ࣀࢩࢩ࡜⾲グ㸧ࡢ⟶⌮࡟ྲྀࡾ⤌ࢇ࡛࠸ࡿࡀ㸦රᗜ 2010㸧ࠊ㐺ษ࡞ಶయᩘ⟶⌮ࢆ⾜࠺ࡓࡵ࡟ࡣࠊಶయ⩌ືែࢆ᥎ᐃࡋࠊᤕ⋓ࡢຠᯝࡸᙳ㡪ࡢ᳨ドࡍࡿᚲせࡀ࠶ࡿࠋࡇࡢㄽᩥ࡛ࡣࠊරᗜ┴࡟࠾ࡅࡿ࢖ࣀࢩࢩࡢ⮬↛ቑຍ⋡ࡸ⏕ᜥಶయᩘ࡞࡝ࢆ᥎ᐃࡋࠊ
ಶయᩘ⟶⌮ࡢ⌧≧࡜ࠊ᥎ᐃᡭἲࡢ㛗ᡤ࣭▷ᡤࢆ⪃ᐹࡍࡿࠋ⮬↛ቑຍ⋡ࡸಶయᩘࡢ᥎ᐃࡣࠊᤕ⋓ᩘࡢᖺḟኚືࢆ୺せ࡞᝟ሗ※࡜ࡋࡓ
Harvest-based model 㸦 Yamamura et al. 2008 㸧 ࢆ ᇶ ᮏ ࣔ ࢹ ࣝ ࡜ ᥇ ⏝ ࡍ ࡿ ࠋHarvested-based model ࢆᇶᮏࣔࢹࣝ࡜ࡋࡓಶయᩘ᥎ᐃࡣࠊ1㸧ᤕ⋓ᩘ࡜㐃ືࡋࡓ᥎ᐃ
࡛࠶ࡿࡓࡵ㔝⏕ື≀ࡢಶయᩘ⟶⌮ࢆ⾜࠺ୖ࡛㔜せ࡞ᤕ⋓┠ᶆࡢタᐃ࡟㐺ࡋ࡚࠸ࡿࠊ2㸧⊁⊟ᤕ⋓ᩘࠊ᭷ᐖᤕ⋓ᩘ࡜ࡶ࡟⾜ᨻࡀ㏻ᖖᴗົ࡜ࡋ࡚㞟ィࡋ࡚࠸ࡿࡢ࡛ࠊ᥎ᐃࡢࡓࡵ࡟㈨㔠࡜ປຊࢆ࠿ࡅ࡚᪂ࡓ࡞ㄪᰝࢆ⾜࠺ᚲせࡀ࡞࠸ࠊ3㸧඲┴ୗ࠿ࡽ⥙⨶ⓗ࡞ࢹ࣮ࢱࡀᚓ
රᗜ┴᳃ᯘື≀◊✲ࢭࣥࢱ࣮࣭ රᗜ┴❧኱Ꮫ⮬↛࣭⎔ቃ⛉Ꮫ◊✲ᡤ࣭㸦ᰴ㸧㔝⏕ື≀ಖㆤ

⟶⌮஦ົᡤ

 

රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-3

ࡽࢀࡿࠊ4㸧⮬἞యࡢ⋇ᐖᢸᙜ⪅ࡀ㞟ィ࣭ᢕᥱࡋࠊఫẸ࡟ࡶඹ᭷ࡉࢀ࡚࠸ࡿᩘ್ࢆᇶ࡟
᥎ᐃࢆ⾜࠺ࡢ࡛ࠊ฼ᐖ㛵ಀ⪅ࡢឤぬ࡜᥎ᐃ್ࡢ஋㞳ࡀᑡ࡞ࡃࠊྜពᙧᡂࡀࢫ࣒࣮ࢬ࡛࠶
ࡿࠊ࡞࡝ࡢ฼Ⅼࡀ࠶ࡿࠋ௒ᅇࡢ᥎ᐃ࡟ࡣࠊරᗜ┴࡛య⣔ⓗ࡟཰㞟ࡋ࡚࠸ࡿ 2002 ᖺ࠿ࡽ 2012 ᖺࡲ࡛ࡢࢹ࣮ࢱ
ࢆ⏝࠸ࡿࠋලయⓗ࡟ࡣࠊරᗜ┴᳃ᯘື≀◊✲ࢭࣥࢱ࣮ࡀ཰㞟ࡋ࡚࠸ࡿ⊁⊟Ⓩ㘓⪅ࡢሗ࿌࡟ᇶ࡙ࡃ㖠⊟᫬ࡢᖹᆒ┠ᧁᩘ㸦┠ᧁຠ⋡㸧ࠊ⊁⊟࡟ࡼࡿᤕ⋓ᩘࠊ᭷ᐖᤕ⋓チྍ࡟ࡼࡿᤕ
⋓ᩘ࡛࠶ࡿࠋ ᥎ᐃࡣࠊᤕ⋓ᩘࡣࡌࡵࠊୖグࡢࢹ࣮ࢱ࡜᫬⣔ิⓗ࡞㛵ಀࢆグ㏙ࡍࡿ㝵ᒙ࣋࢖ࢬࣔࢹࣝ
ࢆᵓ⠏ࡋࠊ࣐ࣝࢥࣇ㐃㙐ࣔࣥࢸ࢝ࣝࣟἲࢆ⏝࠸࡚⾜ࡗࡓࠋࡇࢀࡽࡢసᴗࡢ୰࡛ࡣࠊ⮬↛ቑຍ⋡ࠊಶయᩘࡢ௚࡟ࠊᤕ⋓⋡ࡸ┠ᧁຠ⋡࡜ಶయᩘࡢ㛵ಀࢆ⾲ࡍಀᩘ࡟ຍ࠼ࠊほࢹ࣮ࢱ࡟ྵࡲࢀࡿㄗᕪኚືࡢ኱ࡁࡉ࡞࡝ࢆᵓ⠏ࡋࡓࣔࢹࣝࡢ୰࡛᥎ᐃࡍࡿࠋ
⊁⊟ࡸ᭷ᐖ㥑㝖࡟࠾ࡅࡿᤕ⋓⋡ࡣࠊᕷ⏫ᮧࡸ㒔㐨ᗓ┴ࡀタᐃࡍࡿᤕ⋓ᩘࡸಶయ⩌⟶⌮஦ᴗࡢෆᐜ࡟ࡼࡗ࡚኱ࡁࡃᕥྑࡉࢀࡿࠋࡇࢀࡽࡢኚືࡣࠊ┠ᶆᤕ⋓ᩘࡢኚ᭦ࡸᤕ⋓ᢏ⾡ࡢྥୖࠊ⊁⊟⪅ᩘ࡞࡝ࡢ♫఍᝟ໃࡢኚ໬࡟క࠸ࠊᖺࢆ㏣࠺࡟ࡘࢀ࡚ࠊᚎࠎ࡟ኚືࡍࡿྍ
⬟ᛶࡀ㧗࠸ࠋྠᵝ࡟ࠊ⊁⊟᫬ࡢ┠ᧁຠ⋡࡟ࡘ࠸࡚ࡶࠊ⟶⌮஦ᴗࡢෆᐜࡢኚ᭦ࡸㄪᰝ࡬ࡢ
༠ຊ≧ἣ࡟ᛂࡌ࡚ࠊ⤒ᖺⓗ࡟ኚ໬ࡍࡿྍ⬟ᛶࡀ࠶ࡿࠋࡇࢀࡽࡢၥ㢟ࢆゎỴࡍࡿࡓࡵࠊᖺ
࡟ࡼࡗ࡚ಶࠎ࡟ኚືࡍࡿㄗᕪ࡛ࡣ࡞ࡃࠊᖺࢆ⤒ࡿ࡟ࡘࢀ࡚ᚎࠎ࡟ኚືࡍࡿㄗᕪࢆ⤌ࡳ㎸ࡳࠊ⟶⌮ィ⏬ࡢ㐍ᤖ≧ἣ࡞࡝ࡢ♫఍ⓗせᅉࡢኚ໬ࡀ᥎ᐃ࡟ཬࡰࡍᙳ㡪ࢆ⪃៖ࡋࡓࠋࡲࡓࠊᤕ⋓⋡ࡢኚືࡣࠊ♫఍ⓗ࡞せᅉࡔࡅ࡛ࡣ࡞ࡃࠊ㣗≀㈨※㔞ࡢኚື࡟௦⾲ࡉࢀࡿ⮬↛⎔ቃࡢኚ໬࡟ࡼࡗ࡚ࡶᕥྑࡉࢀࡿࠋࣈࢼ⛉ࡢሀᯝ㢮ࡣࠊࡑࡢ㇏ปࡀࢶ࢟ࣀ࣡ࢢ࣐㸦⸨
ᮌ࡯࠿ 2011㸧ࢆࡣࡌࡵࠊ㔝⏕ື≀ࡢฟἐࡸᤕ⋓≧ἣࢆᕥྑࡍࡿ௦⾲ⓗ࡞㣗≀㈨※࡛࠶ࡿࠋᮏ✀࡟ࡘ࠸࡚ࡶࠊ⛅Ꮨ࡟ࡣ୺せ࡞㣗≀㈨※࡜ࡋ࡚ሀᯝ㢮ࢆ฼⏝ࡋ࡚࠸ࡿ㸦ᑠᑎ࣭⚄ᓮ 2001㸧ࡇ࡜࠿ࡽࠊ㇏ปࡢ⛬ᗘ࡟ࡼࡗ࡚ᤕ⋓⋡ࡀኚືࡍࡿྍ⬟ᛶࡣ㧗࠸㸦ୖ⏣࡯࠿
2013㸧ࠋࡑࡢࡓࡵࠊࣈࢼ⛉ሀᯝ㢮ࡢ㇏ปㄪᰝࢆ⾜ࡗ࡚࠸ࡿᮏᕞ㒊ࡢ᥎ᐃ࡛ࡣࠊ⊁⊟࡜᭷ᐖᤕ⋓᫬ࡢᤕ⋓⋡࡟ࡘ࠸࡚ࠊ㇏ปㄪᰝࢹ࣮ࢱࢆ౑⏝ࡋ࡚ࠊ㇏ปࡢᙳ㡪ࢆ⿵ṇࡍࡿࣔࢹࣝ
ࢆᵓ⠏ࡋࡓࠋῐ㊰ᓥ࡛ࡣࠊ㇏ปㄪᰝࢆ⾜ࡗ࡚࠸࡞࠸ࡓࡵࠊ㇏ปࡢᙳ㡪ࡣ⿵ṇ࡛ࡁ࡞࠿ࡗ
ࡓࠋࡉࡽ࡟ࠊᮏ✀ࡣࠊቑຍ⋡ࡀᖺ࡟ࡼࡗ࡚኱ࡁࡃኚືࡍࡿࡇ࡜ࡀሗ࿌ࡉࢀ࡚࠸ࡿࡇ࡜࠿ࡽࠊ
ࡑࡢᖺḟኚື࡟ࡘ࠸࡚ࡶ᥎ᐃࡍࡿࡼ࠺࡟ࠊࣔࢹࣝࢆᵓ⠏ࡋࡓࠋᮏㄽ࡛ࡣࠊ௨ୖࡢࡼ࠺࡞ᇶᮏⓗ࡞⪃࠼᪉࡟ᇶ࡙ࡁ᥎ᐃࢆ⾜ࡗ࡚⤖ᯝࢆ᳨ウࡋࠊරᗜ┴ࡢ࢖ࣀࢩࢩࡢᤕ⋓࡜ಶయ⩌ࡢ≧ἣࢆ⪃ᐹࡍࡿࠋ㸬᪉ἲ
ࠕ3-1㸬ࡣࡌࡵ࡟࡛ࠖ㏙࡭ࡓᇶᮏⓗ࡞⪃࠼᪉࡟ἢࡗ࡚ࠊලయⓗ࡞ Harvested-basedmodel ࡟ᇶ࡙࠸ࡓ㝵ᒙ࣋࢖ࢬࣔࢹࣝ࡜࣐ࣝࢥࣇ㐃㙐ࣔࣥࢸ࢝ࣝࣟἲ࡟ࡼࡿ᥎ᐃࢆࠊ௨ୗ
ࡢࡼ࠺࡞ࢹ࣮ࢱ࡜᪉ἲ࡛⾜ࡗࡓࠋ රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-3

ᑐ㇟࡜ࡍࡿᮇ㛫࡜ᆅᇦᑐ㇟ᮇ㛫ࡣࠊ2002 ᖺ࠿ࡽ 2012 ᖺࡲ࡛࡜ࡋࡓࠋᑐ㇟ᆅᇦࡣරᗜ┴ᮏᕞ㒊ࠊ࠾ࡼࡧῐ
㊰ᓥ࡜ࡍࡿࠋ
⏝࠸ࡓࢹ࣮ࢱ ᥎ᐃ࡟ࡣ௨ୗࡢࢹ࣮ࢱࢆ౑⏝ࡋࡓࠋ᭷ᐖᤕ⋓ᩘ㸦ᖺᗘ㸧 i_yugai[i]㸸i ᖺᗘࡢ᭷ᐖチྍ࡟ࡼࡿᤕ⋓ᩘࠋಶయᩘࢆ཯ᫎࡍࡿᣦ
ᶆ࡜ࡋ࡚⏝࠸ࡿࠋ
┠ᧁຠ⋡ spue[i]㸸i ᖺᗘࡢ⊁⊟ᮇ㛫୰࡟ࠊ⊁⊟⪅Ⓩ㘓⪅࠿ࡽᚓࡽࢀࡓ㖠⊟᫬ࡢ┠ᧁຠ⋡ࠋಶయᩘࢆ཯ᫎࡍࡿᣦᶆ࡜ࡋ࡚⏝࠸ࡿࠋ⊁⊟ᤕ⋓ᩘ r_ca[i]㸸i ᖺᗘࡢ⊁⊟࡟ࡼࡿᤕ⋓ᩘࠋಶయᩘࢆ཯ᫎࡍࡿᣦᶆ࡜ࡋ࡚⏝࠸ࡿࠋ᭷ᐖᤕ⋓ᩘ㸦ᖺ㸧 y_ca[i]㸸i ᖺᗘࡢ 1 ᭶࠿ࡽ i+1 ᖺᗘࡢ 12 ᭶ࡲ࡛ࡢ᭷ᐖチྍ࡟ࡼࡿᤕ ⋓ᩘࠋ᳃ᯘ㠃✚ f_area㸸රᗜ┴ᮏᕞ㒊ࡲࡓࡣῐ㊰ᓥࡢ᳃ᯘ㠃✚ࠋ⏕ᜥᐦᗘࡢᮇᚅ್ࢆィ⟬ࡍࡿ㝿࡟⏝࠸ࡿࠋ
ࣈࢼ⛉ሀᯝ㢮ࡢ㇏ปᣦᩘ nut[i]㸸i ᖺࡢ㇏ปᣦᩘࠋே࡜⮬↛ࡢ༤≀㤋ཬࡧ᳃ᯘື≀◊✲ࢭࣥࢱ࣮ࡢㄪᰝ⤖ᯝ࠿ࡽ㞟ィࡋࡓࢶ࢟ࣀ࣡ࢢ࣐ࡢฟἐᆅᇦ࡛ࡢ㇏ปᣦᩘ㸦⸨ᮌ࡯࠿2011㸧ࠋᐇ㝿ࡢ᥎ᐃ࡟ࡣࠊ⮬↛ᑐᩘࢆ࡜ࡗ࡚ᖹᆒ 0ࠊศᩓ 1 ࡟ᶆ‽໬ࡋ࡚⏝࠸ࡿࠋῐ㊰ᓥ࡛ࡣ㇏ปᣦᩘࡢࢹ࣮ࢱࢆ࡜ࡗ࡚࠸࡞࠸ࡢ࡛ᮏᕞ㒊ࡢ᥎ᐃ࡛ࡢࡳ⏝࠸ࡓࠋ௨ୖࡢ᪉ἲ࡛཰㞟ࡋࡓࢹ࣮ࢱࢭࢵࢺࢆ⾲㸯࡟♧ࡍࠋ᥎ᐃࡍࡿࣃ࣓࣮ࣛࢱ
௨ୗࡢ⪃࠼᪉࡟ᇶ࡙࠸࡚ࠊᮏᕞ㒊࡛ࡣࠊ lire2002-2011ࠊlr_spueࠊpre2002ࠊpyp2002ࠊ
lnNinsࠊlkࠊprrࠊpyrࠊnutx2002-2004ࠊv_spueࠊv_ryoࠊv_yugaiࠊran_ryo2002-2011ࠊ
ran_yugai2002-2011ࠊran_s2002-2011ࠊv_ran_ryoࠊv_ran_yugaiࠊv_ran_s ࡢ 56 ࡢ
ኚᩘࠊῐ㊰ᓥ࡛ࡣ lire2002-2011ࠊlr_spueࠊpre2002ࠊpyp2002ࠊlnNinsࠊlkࠊv_spueࠊ
v_ryoࠊv_yugaiࠊran_ryo2002-2011ࠊran_yugai2002-2011ࠊran_s2002-2011ࠊv_ran_ryoࠊ
v_ran_yugaiࠊv_ran_s ࡢ 51 ኚᩘ࡟ࡘ࠸࡚᥎ᐃࡋࠊ┠ⓗ࡛࠶ࡿ⮬↛ቑຍ⋡ࡸಶయᩘࢆ
᥎ᐃࡍࡿࠋ᥎ᐃኚᩘࡢึᮇ್࡜஦๓ศᕸࡣ⾲㸰ࡢ࡜࠾ࡾ࡛ࠊྛ᥎ᐃኚᩘࡢᐃ⩏࡜஦๓ศᕸタᐃࡢ㝿ࡢ⪃࠼᪉ࡣࠊ௨ୗࡢ࡜࠾ࡾ࡛࠶ࡿࠋ
1.

⮬↛ቑῶ⋡ࡢᑐᩘ್ lire[i](i = 2002, 2003,࣭࣭࣭2011) 㸸ฟ⏕࡜⮬↛Ṛஸࡢ⤖ᯝ࡜ࡋ࡚ࡢ㞤㞝ྜࢃࡏࡓ඲ಶయᩘ࡟ᑐࡍࡿቑῶࡢẚ⋡࡜ࡍࡿࠋlire ࡟ࡘ࠸࡚ࡣࠊ⎔ቃ┬ࡢ≉ᐃ့ங㢮⏕ᜥືྥㄪᰝࡢಶయᩘ᥎ᐃ㸦⎔ቃ┬⏕≀ከᵝᛶࢭࣥࢱ࣮ 2011㸧࡟᥇⏝ࡉࢀࡓ஦๓ศᕸࢆ⏝࠸ࡿࠋࡲࡓࠊexp(lire)ࢆ⮬↛ቑຍ⋡ ire ࡜ࡍࡿࠋ2.

⎔ቃ཰ᐜຊࡢᑐᩘ್ lk㸸ᖺ࡟ࡼࡿኚືࡣ࡞ࡃࠊ᥎ᐃᮇ㛫୰ࠊ୍ᐃ࡛࠶ࡿ࡜௬ᐃࡍ
ࡿࠋlk ࡢ஦๓ศᕸࡣṇつศᕸࢆ௬ᐃࡋࠊ஦๓ࡢ᝟ሗࡣ࡞࠸ࡓࡵࠊศᩓࡣ኱ࡁࡵ࡟タ
ᐃࡍࡿࠋ

 

රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-3

⾲㸯㸫㸯 ධຊࢹ࣮ࢱࢭࢵࢺ㸦ᮏᕞ㒊㸧᭷ᐖᤕ⋓ᩘ

ᖺ

㸦ḟᖺ㸧

⊁⊟ᤕ⋓ᩘ

᳃ᯘ㠃✚

᭷ᐖᤕ⋓ᩘ

┠ᧁ

㇏ป



㸦NP 

㸦ᖺᗘ㸧

ຠ⋡

ᣦᩘ













 











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 

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

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

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

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

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

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

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 

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
⾲㸯㸫㸰 ධຊࢹ࣮ࢱࢭࢵࢺ㸦ῐ㊰ᓥ㸧ᖺ㻌᭷ᐖᤕ⋓ᩘ㻌䠄ḟᖺ䠅㻌⊁⊟ᤕ⋓ᩘ㻌᳃ᯘ㠃✚㻌᭷ᐖᤕ⋓ᩘ㻌

┠ᧁ㻌䠄㼗㼙㻞㻕㻌

䠄ᖺᗘ䠅㻌

ຠ⋡㻌

㻞㻜㻜㻞㻌

㻞㻝㻝㻌

㻞㻞㻥㻌

㻟㻟㻢㻚㻤㻢㻌 㻌㻞㻝㻞㻌㻜㻚㻠㻜㻌 㻌

㻞㻜㻜㻟㻌

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㻟㻟㻢㻚㻤㻢㻌 㻌㻞㻜㻠㻌㻜㻚㻡㻜㻌 㻌

㻞㻜㻜㻠㻌

㻞㻞㻜㻌

㻟㻥㻣㻌

㻟㻟㻢㻚㻤㻢㻌 㻌㻟㻝㻡㻌㻜㻚㻡㻤㻌 㻌

㻞㻜㻜㻡㻌

㻟㻠㻢㻌

㻟㻣㻟㻌

㻟㻟㻢㻚㻤㻢㻌 㻌㻞㻟㻣㻌㻜㻚㻣㻠㻌 㻌

㻞㻜㻜㻢㻌

㻢㻠㻢㻌

㻟㻤㻟㻌

㻟㻟㻢㻚㻤㻢㻌 㻌㻟㻣㻞㻌㻜㻚㻠㻢㻌 㻌

㻞㻜㻜㻣㻌

㻥㻣㻢㻌

㻠㻠㻝㻌

㻟㻟㻢㻚㻤㻢㻌 㻌㻢㻣㻜㻌㻜㻚㻡㻞㻌 㻌

㻞㻜㻜㻤㻌

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㻟㻟㻢㻚㻤㻢㻌 㻌㻝㻜㻢㻥㻌㻜㻚㻡㻞㻌 㻌

㻞㻜㻜㻥㻌

㻝㻡㻜㻡㻌

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㻟㻟㻢㻚㻤㻢㻌 㻌㻝㻝㻝㻜㻌㻜㻚㻠㻣㻌 㻌

㻞㻜㻝㻜㻌

㻝㻣㻢㻣㻌

㻝㻥㻟㻣㻌

㻟㻟㻢㻚㻤㻢㻌 㻌㻝㻠㻡㻥㻌㻜㻚㻡㻞㻌 㻌

㻞㻜㻝㻝㻌

㻝㻢㻜㻜㻌

㻝㻡㻡㻞㻌

㻟㻟㻢㻚㻤㻢㻌 㻌㻝㻢㻥㻜㻌㻜㻚㻟㻥㻌 㻌

㻞㻜㻝㻞㻌

㻌㻌

㻝㻠㻥㻢㻌

㻟㻟㻢㻚㻤㻢㻌 㻌㻝㻢㻢㻞㻌㻜㻚㻠㻟㻌 㻌3.

⏕ ᜥ ᐦ ᗘ ࡜ ┠ ᧁ ຠ ⋡ ࡢ ẚ ⋡ ࢆ ♧ ࡍ ಀ ᩘ ࡢ ⮬ ↛ ᑐ ᩘ ್  lr_spue[i] (i =2002,2003,࣭࣭࣭2012)㸸 lr_spue[i]ࡣ๓ᖺࡢ್࠿ࡽࣛࣥࢲ࣒࡟ኚື㸦ࣛࣥࢲ࣒࢘
࢛࣮ࢡ㸧ࡍࡿ࡜⪃࠼ࠊlr_spue[i+1] = lr_spue[i]+ran_s[i]࡜ࡍࡿࠋlr_spue 2002 ࡢ஦๓ࡢ᝟ሗࡣ༑ศ࡟࡞࠸ࡓࡵࠊ஦๓ศᕸࡢศᩓࡣ኱ࡁࡵ࡟タᐃࡍࡿࠋran_s ࡣࠊᖹᆒ 0ࠊศᩓ v_ran_s ࡢṇつศᕸ࡟ᚑ࠺㝵ᒙⓗ࡞஦๓ศᕸࢆタᐃࡍࡿࠋࡇࡢศᩓ v_raࡢ஦๓ศᕸࡣᙧ≧ẕᩘࠊᑻᗘẕᩘ࡜ࡶ࡟ 0.01 ࡢ㏫࣐࢞ࣥศᕸࢆ⏝࠸ࡓࠋࡲࡓࠊ
exp(lr_spue[i])ࢆ rs[i]࡜ࡋࡓࠋ රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-3

4.

⊁⊟᫬ࡢᤕ⋓⋡ pr[i] (i = 2002, 2003,࣭࣭࣭2012)㸸⊁⊟ࡢᤕ⋓ᩘࡢ⏕ᜥಶయᩘ࡟
ᑐࡍࡿẚ⋡ࢆ⾲ࡍࠋࡇࡢಀᩘࡣࠊ0 ࠿ࡽ 1 ࡢ㛫࡛ኚືࡍࡿ࡜⪃࠼ࠊ
pr[i]=1/(1+exp(-(pre[i]+prrnut[i])))࡜ࡋࠊpre2002 ࡜ prr ࢆ᥎ᐃࡍࡿࠋpre[i]ࡣ๓ᖺ ࡢ ್ ࠿ ࡽ ࣛ ࣥ ࢲ ࣒ ࡟ ኚ ື 㸦 ࣛ ࣥ ࢲ ࣒ ࢘ ࢛ ࣮ ࢡ 㸧 ࡍ ࡿ ࡜ ⪃ ࠼ ࠊ pre[i+1]=
pre[i]+ran_ryo[i]࡜ࡍࡿࠋpre2002ࠊprr ࡜ࡶ࡟஦๓ࡢ᝟ሗࡣ༑ศ࡟࡞࠸ࡓࡵࠊ஦๓
ศᕸࡢศᩓࡣ኱ࡁࡵ࡟タᐃࡍࡿࠋࡇࡢ ran_ryo ࡣࠊᖹᆒ 0 ࡛ศᩓ v_ran_ryo ࡢṇつศᕸ࡟ᚑ࠺㝵ᒙⓗ࡞஦๓ศᕸࢆタᐃࡋࡓࠋࡇࡢศᩓ v_ran_ryo ࡢ஦๓ศᕸࡣᙧ≧ẕᩘࠊᑻᗘẕᩘ࡜ࡶ࡟ 0.01 ࡢ㏫࣐࢞ࣥศᕸࢆ⏝࠸ࡓࠋ

5.

᭷ᐖ᫬ࡢᤕ⋓⋡ py[i] (i = 2002, 2003,࣭࣭࣭2012)㸸᭷ᐖࡢᤕ⋓ᩘࡢ⏕ᜥಶయᩘ࡟ ᑐ ࡍࡿẚ⋡ࢆ ⾲ࡍࠋࡇࡢ ಀᩘࡣࠊ0 ࠿ࡽ 1㸫 pr[i]ࡢ㛫 ࡛ኚືࡍࡿ ࡜⪃࠼ ࠊ
py[i]=(1-pr[i])/(1+exp(-(pyp[i]+pyrnut[i])))࡜ࡋࠊpyp2002 ࡜ pyr ࢆ᥎ᐃࡍࡿࠋ
ࡲࡓࠊpyp[i]ࡣ᫬㛫ⓗ࡞ኚ໬࡟క࠸ࠊ๓ᖺࡢ್࠿ࡽࣛࣥࢲ࣒࡟ኚື㸦ࣛࣥࢲ࣒࢛࢘
࣮ࢡ㸧ࡍࡿ࡜⪃࠼ࠊpyp[i+1] = pyp[i]+ran_yugai[i]࡜ࡍࡿࠋpyp2002ࠊpyr ࡜ࡶ࡟஦๓ࡢ᝟ሗࡣ༑ศ࡟࡞࠸ࡓࡵࠊ஦๓ศᕸࡢศᩓࡣ኱ࡁࡵ࡟タᐃࡍࡿࠋࡇࡢ
ran_yugai ࡣࠊᖹᆒ 0 ࡛ศᩓ v_ran_yugai ࡢṇつศᕸ࡟ᚑ࠺㝵ᒙⓗ࡞஦๓ศᕸࢆタᐃࡋࡓࠋࡇࡢศᩓ v_ran_yugai ࡢ஦๓ศᕸࡣᙧ≧ẕᩘࠊᑻᗘẕᩘ࡜ࡶ࡟ 0.01 ࡢ㏫࣐࢞ࣥศᕸࢆ⏝࠸ࡓࠋ

6.

4ࠊ5 ࡢ nut[i]࡟ࡘ࠸࡚ࠊ2005 ᖺ࠿ࡽ 2012 ᖺࡣࠊᐇ್ࡢᑐᩘࢆᖹᆒ 0ࠊศᩓ 1 ࡟ᶆ‽໬ࡋࡓ್㸦nut[i]㸧ࢆ⏝࠸ࡿࠋㄪᰝࢆ⾜ࡗ࡚࠸࡞࠸ᖺࡢ㇏ปᣦᩘ nut[i]࡟ࡘ࠸
࡚ࡣࠊ᥎ᐃ್࡛࠶ࡿ nutx[i]ࢆ௦ධࡍࡿࠋࡇࡢ nutx[i]ࡢ஦๓ศᕸࡣᖹᆒ 0ࠊศᩓ 1ࡢṇつศᕸ࡟ᚑ࠺࡜ࡋࡓࠋᮏᕞ㒊࡛ࡢࡳ᥎ᐃࡋࡓࠋ
7.

2011 ᖺࡢ⏕ᜥಶయᩘࡢ⮬↛ᑐᩘ್ lnNins㸸஦๓ศᕸࡣṇつศᕸ࡜ࡋࠊᖹᆒ್࡟
ࡣරᗜ┴ࡀ๓ᖺ࡟⾜ࡗࡓಶయᩘࡢ᥎ᐃ್㸦ᮏᕞ㒊㸹ᆏ⏣࡯࠿ 2012ࠊῐ㊰ᓥ㸹㛵࡯࠿ 2012㸧ࢆ⏝࠸ࡓࠋࡓࡔࡋࠊ2011 ᖺᗘࡢಶయᩘࡢಙ㢗༊㛫࡟㛵ࡍࡿ஦๓ࡢ᝟ሗࡣ
ᑡ࡞࠸ࡓࡵࠊศᩓࡣ኱ࡁࡵ࡟タᐃࡋࡓࠋ2002 ᖺ࠿ࡽ 2011 ᖺࡲ࡛ࡢࢹ࣮ࢱࡣ๓ᅇࡢ
᥎ᐃ࡜௒ᅇࡢ᥎ᐃ࡜࡛ 2 ᅇ⏝࠸ࡿࡇ࡜࡟࡞ࡿࠋࡋ࠿ࡋ๓ᅇ࡜᥎ᐃ್ࡀⴭࡋࡃ␗࡞ࡾࠊ
᥎ᐃ್ࡢಙ㢗ࡀኻࢃࢀࡿࡇ࡜ࢆ㜵ࡄࡓࡵࠊ๓ᅇࡢ᥎ᐃ್ࡢᖹᆒ್ࡢࡳࢆ஦๓ศᕸࡢ
ᖹᆒ࡜ࡋ࡚⏝࠸ࡓࠋ᏶඲࡟ࢹ࣮ࢱࡢ஧ᗘ౑࠸࡜࡞ࡿࡇ࡜ࢆ㜵ࡄࡓࡵࠊ஦๓ศᕸࡢศ
ᩓࡣᗈࡃタᐃࡋࡓࠋ

8.

┠ᧁຠ⋡ࠊ⊁⊟ᤕ⋓ᩘࠊ᭷ᐖᤕ⋓ᩘࡢᮇᚅ್࠿ࡽࡢㄗᕪศᩓ v_spue v_ryo
v_yugai㸸ࡑࢀࡒࢀࠊほࣔࢹ࡛ࣝ♧ࡍ☜⋡ศᕸࡢㄗᕪศᩓ࡜ࡋ࡚ほࢹ࣮ࢱ࠿ࡽ᥎ᐃࡍࡿࠋࡇࢀࡽࡢㄗᕪศᩓࡢ஦๓ศᕸࡣࠊᙧ≧ẕᩘ 0.01ࠊᑻᗘẕᩘ 0.01 ࡢ㏫࣐࢞ࣥศᕸࢆ⏝࠸ࡓࠋ
9.

ྛ᥎ᐃኚᩘࡢึᮇ್ࡣࠊ஦๓ศᕸࡢᮇᚅ್࡜ࡋࡓࠋv_spueࠊv_ryoࠊv_yugaiࠊv_ran_sࠊ
v_ran_ryoࠊv_ran_yugai ࡟ࡘ࠸࡚ࡣࠊࡑࢀࡒࢀึᮇ್ࢆ 0.01 ࡜ࡋࡓࠋ රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-3

⾲㸰㸫㸯 ᥎ᐃࡋࡓኚᩘ࡜ࡑࡢึᮇ್ࠊ࠾ࡼࡧ஦๓ศᕸ㸦රᗜ┴ᮏᕞ㒊㸧
᥎ᐃኚᩘ

ࣈࣟࢵࢡ

ึᮇ್

஦๓ศᕸ

OLUH



ṇつศᕸ

ON



ṇつศᕸ ORJ   YDU  

ORJ    YDU  

OUBVSXH

ṇつศᕸSUH

ORJ 

YDU  ࣮࣋ࢱศᕸ  

SU\

OQ1LQV

࣮࣋ࢱศᕸ  

YBUDQBU\R





㏫࣐࢞ࣥศᕸ VFDOH  

YBUDQB\XJDL





㏫࣐࢞ࣥศᕸ VFDOH  

YBUDQBV

㏫࣐࢞ࣥศᕸ VFDOH  YBVSXH

㏫࣐࢞ࣥศᕸ VFDOH  

YBU\R



㏫࣐࢞ࣥศᕸ VFDOH  

YB\XJDL

㏫࣐࢞ࣥศᕸ VFDOH  

UDQBVṇつศᕸ Y YBUDQBV 

UDQBU\Rṇつศᕸ Y YBUDQBU\R 

UDQB\XJDLṇつศᕸ Y YBUDQB\XJDL 

QXW[ṇつศᕸ YDU  

SUU





ṇつศᕸ YDU  

S\U





ṇつศᕸ YDU  ṇつศᕸ YDU  ஦๓ศᕸࡢᘬᩘࡣࠊṇつศᕸ㸦ᮇᚅ್ࠊศᩓ㸧
ࠊ㏫࣐࢞ࣥศᕸ㸦ᙧ≧ẕᩘࠊᑻᗘẕᩘ㸧ࠊ࣮࣋ࢱศᕸ㸦ᙧ≧ẕᩘȘࠊᙧ≧ẕᩘș㸧ࢆ♧ࡍࠋ⾲㸰㸫㸰 ᥎ᐃࡋࡓኚᩘ࡜ࡑࡢึᮇ್ࠊ࠾ࡼࡧ஦๓ศᕸ㸦රᗜ┴ῐ㊰ᓥ㸧
᥎ᐃኚᩘ



ࣈࣟࢵࢡ

ึᮇ್

஦๓ศᕸ

OLUH



ṇつศᕸ

ON



ṇつศᕸ ORJ   YDU  

OUBVSXH

ṇつศᕸ

SUH

࣮࣋ࢱศᕸ  

SU\

࣮࣋ࢱศᕸ  

OQ1LQVYBUDQBU\R





㏫࣐࢞ࣥศᕸ VFDOH  

YBUDQB\XJDL





㏫࣐࢞ࣥศᕸ VFDOH  

YBUDQBV

㏫࣐࢞ࣥศᕸ VFDOH  YBVSXH

㏫࣐࢞ࣥศᕸ VFDOH  

YBU\R



㏫࣐࢞ࣥศᕸ VFDOH  

YB\XJDL

㏫࣐࢞ࣥศᕸ VFDOH  

UDQBVṇつศᕸ Y YBUDQBV 

UDQBU\Rṇつศᕸ Y YBUDQBU\R 

UDQB\XJDLṇつศᕸ Y YBUDQB\XJDL 












ORJ    YDU  

ORJ 

YDU  ṇつศᕸ YDU  ஦๓ศᕸࡢᘬᩘࡣࠊṇつศᕸ㸦ᮇᚅ್ࠊศᩓ㸧
ࠊ㏫࣐࢞ࣥศᕸ㸦ᙧ≧ẕᩘࠊᑻᗘẕᩘ㸧ࠊ࣮࣋ࢱศᕸ㸦ᙧ≧ẕᩘȘࠊᙧ≧ẕᩘș㸧ࢆ♧ࡍࠋಶయ⩌ືែࡢ㐣⛬ࣔࢹࣝಶయ⩌ືែࡢ㐣⛬ࣔࢹࣝࡣࠊBeverton-holt ࣔࢹࣝࢆ᥇⏝ࡋࡓࠋ඲⏕ᜥಶయᩘࡣ 2011ᖺࢆ㉳Ⅼ࡜ࡋࠊ
⩣ᖺࡢ 2012 ᖺࡲ࡛ࡢኚ໬ࢆ
ß[2011] = (ire[2011]㸫1)/(kf_area[2011])
N[2012] = (ire[2011]N[2011]/(1㸩ß[2011]N[2011]))㸫caa[2011] රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-3

2002 ᖺࡲ࡛ࡢኚ໬ࢆ
ß[i] = (ire[i]㸫1)/(kf_area[i])N[i] = (caa[i]㸩N[i+1])/(ire[i]㸫ß[i](caa[i]㸩N[i+1]))ࡢࡼ࠺࡟ኚ໬ࡍࡿࡶࡢ࡜௬ᐃࡍࡿࠋ
ࡇࡇ࡛ࠊN[i]ࡣࠊi ᖺࡢ⏕ᜥಶయᩘࢆ♧ࡍࠋࡲࡓࠊcaa[i]ࡣࠊi ᖺࡢᤕ⋓ᩘ࡛࠶ࡾࠊiᖺࡢ⊁⊟ᤕ⋓ᩘ r_ca[i]࡜᭷ᐖᤕ⋓ᩘ y_ca[i]ࡢྜィ್࡛࠶ࡿࠋ
2011 ᖺࡢಶయᩘࡣ N2011 = round(exp(lnNins))࡜ࡋࡓࠋ࡞࠾ࠊ⏕ᜥಶయᩘࡣࠊᖺᮎ
᫬Ⅼ࡛ࡢಶయᩘࢆ᝿ᐃࡋ࡚࠸ࡿࠋ

ほ

ࣔࢹࣝ
᥎ᐃࡍࡿಶయᩘ࡜ほࡉࢀࡿࢹ࣮ࢱ࡜ࡢ㛵ಀࢆ♧ࡍほࣔࢹࣝࡣ௨ୗࡢ࡜࠾ࡾ࡜ࡍ

ࡿࠋ
1.

┠ᧁຠ⋡࡟㛵ࡍࡿほࣔࢹࣝlog(SPUE[i]) = log(rs[i]N[i]/f_area) -0.5v_spue + e_spue[i@2.

⊁⊟ᤕ⋓ᩘ࡟㛵ࡍࡿほࣔࢹࣝ

log(r_ca [i]) = log(pr[i]N[i]) -0.5v_ryo + e_ryo[i]3.

᭷ᐖᤕ⋓ᩘ࡟㛵ࡍࡿほࣔࢹࣝ

log(i_yugai [i]) = log(py[i]N[i]) -0.5v_yugai + e_yugai [i]e_spue[i]ࠊe_ryo[i]ࠊe_yugai [i]ࡣࠊㄗᕪኚືࢆ♧ࡋࠊࡑࢀࡒࢀᮇᚅ್ 0ࠊศᩓࡀ v_spue ࠊv_ryo ࠊv_yugai ࡢṇつศᕸ࡟ᚑ࠺ࡶࡢ࡜ࡍࡿࠋ
࣐ࣝࢥࣇ㐃㙐ࣔࣥࢸ࢝ࣝࣟἲ
ࡇࢀࡲ࡛㏙࡭ࡓࢹ࣮ࢱ࡜ࣔࢹࣝ࠾ࡼࡧ஦๓ศᕸࡢタᐃ࡟ࡶ࡜࡙࠸࡚ࠊ࣐ࣝࢥࣇ㐃㙐ࣔࣥࢸ࢝ࣝࣟἲ㸦Gilks et al. 1996㸧࡟ࡼࡿ᥎ᐃࢆ⾜ࡗࡓࠋࡇࡢ᥎ᐃࡣ SAS/STAT9.3 ࡢ
MCMC Procedure ࢆ⏝࠸ࡓ㸦SAS Institute Inc. 2011㸧ࠋࢧࣥࣉࣜࣥࢢ
᥎ᐃኚᩘࢆ⾲㸰ࡢ࡜࠾ࡾᮏᕞ㒊ࡢ᥎ᐃ࡛ࡣ 7 ࡘࠊῐ㊰ᓥ࡛ࡣ 6 ࡘࡢࣈࣟࢵࢡ࡟ศࡅ࡚ࠊ
࣓ࢺ࣏ࣟࣜࢫἲ࡜ conjugate ࢧࣥࣉࣜࣥࢢ࡟ࡼࡿ⊂❧ࢧࣥࣉ࣮ࣛࢆ⏝࠸࡚஦ᚋศᕸࢆ
ࢧࣥࣉࣜࣥࢢࡋࡓࠋࢧࣥࣉࣜࣥࢢᅇᩘ࡟ࡘ࠸࡚ࡣࠊ᭱ึࡢ 100 ୓ᅇࡣࢧࣥࣉࣜࣥࢢࡏ
ࡎࠊᮏᕞ㒊ࡢ᥎ᐃ࡛ࡣḟࡢ 5000 ୓ᅇࡢ࠺ࡕ 5,000 ᅇ࡟ 1 ᅇࢧࣥࣉࣜࣥࢢࡋࠊῐ㊰ᓥࡢ᥎ᐃ࡛ࡣḟࡢ 2000 ୓ᅇࡢ࠺ࡕ 2,000 ᅇ࡟ 1 ᅇࢧࣥࣉࣜࣥࢢࡋࠊィ 1 ୓ᅇࡢࢧࣥࣉࣜࣥ
ࢢࢆ⾜ࡗࡓࠋ
ᥦ᱌ศᕸࡣࠊṇつศᕸ࡜ࡋࠊᐇ㝿ࡢࢧࣥࣉࣜࣥࢢᅇᩘ࡟ྜࢃࡏ࡚ 5 ୓ᅇࡢࢧࣥࣉࣜࣥ
ࢢ࡟ࡼࡿ஦ᚋศᕸ࡟ࡶ࡜࡙࠸࡚ࠊRoberts et al.㸦1997㸧ࡢ♧ࡋࡓ᭱㐺࡞᥇ᢥ⋡ 23.4㸣ࢆ┠ᶆ࡟±7.5㸣ࡢ⠊ᅖࡢ᥇ᢥ⋡࡟࡞ࡿࡼ࠺࡟ࠊࢫࢣ࣮ࣝ࡜ඹศᩓ⾜ิࡢࢳ࣮ࣗࢽࣥࢢࢆ
⾜ࡗࡓࠋ

 

රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-3

཰᮰ุᐃ ཰᮰ุᐃࡣࠊ᭷ຠࢧࣥࣉࣝࢧ࢖ࢬ㸦Kass et al. 1998㸧࡜ Geweke ᳨ᐃ㸦Geweke 1992㸧
ࡢ 2 ࡘࡢᇶ‽࡛☜ㄆࡋࡓࠋ᭷ຠࢧࣥࣉࣝࢧ࢖ࢬ࡟ࡼࡿุᐃ࡛ࡣࠊࡇࢀࡀ 500 ௨ୖ࡛࠶ࡿࡇ࡜ࢆᇶ‽࡜ࡋࡓࠋGeweke ἲ࡛ࡣࠊࢧࣥࣉࣜࣥࢢࡉࢀࡓࢹ࣮ࢱࡢ࠺ࡕࠊ᭱ึࡢ 1,000ᅇ࡜᭱ᚋࡢ 5,000 ᅇࡢᮇᚅ್ࡢᕪࢆ᳨ᐃࡋࡓࠋᲠ༷Ỉ‽ࡣ 0.05 ࡜ࡋࡓࠋ㸬⤖ᯝ
཰᮰≧ἣ࠸ࡎࢀࡢ᥎ᐃኚᩘ࡟ࡘ࠸࡚ࡶࢧࣥࣉࣜࣥࢢࡢ㝿ࡢ⮬ᕫ┦㛵ࡣ࡯࡜ࢇ࡝࡞ࡃࠊ᭷ຠࢧࣥ
ࣉࣝᩘࡣ 500 ࢆ㉸࠼ࠊⰋዲ࡞ࢧࣥࣉࣜࣥࢢࡀ࡛ࡁࡓ࡜ุ᩿ࡉࢀࡓࠋGeweke ᳨ᐃࡢ⤖ᯝࡶᮏᕞ㒊ࡢ᥎ᐃࡢ lire2007ࠊ࡜ lire2009 ௨እࡣ᭷ពᕪࡀ᳨ฟࡉࢀࡓኚᩘࡣ࡞࠿ࡗࡓࠋ
ᮏᕞ㒊ࡢ᥎ᐃ࡛ࡣྜィ 56 ࡢኚᩘࢆ᥎ᐃࡋࡓࡓࡵࠊ5㸣Ỉ‽࡛ࡣഅ↛࡟ࡼࡗ࡚ࡶ 3 ኚᩘ⛬ᗘ࡟᭷ពᕪࡀ࡛࡚ࡋࡲ࠺ࡓࡵࠊ2 ࡘࡢኚᩘ࡛᭷ពᕪࡀ࡛ࡿࡇ࡜ࡣチᐜ⠊ᅖ࡜ุ᩿ࡋࡓࠋ
᥎ᐃ್᥎ᐃࡋࡓኚᩘࡢ஦ᚋศᕸࡣ⾲㸱ࡢ㏻ࡾ࡛࠶ࡗࡓࠋࡲࡓࠊ஦๓ศᕸ࡜஦ᚋศᕸࡢᙧ≧ࢆᅗ㸯࡟♧ࡋࡓࠋ⾲ 3 ࡢ⤖ᯝ࡟ᇶ࡙࠸࡚ィ⟬ࡋࡓ⮬↛ቑຍ⋡(ir)ࠊ┠ᧁຠ⋡ࡢಀᩘ(rs)ࠊ⊁⊟ᤕ⋓⋡(pr)ࠊ᭷ᐖᤕ⋓⋡(py)ࡣ⾲㸲ࡢ࡜࠾ࡾ࡛࠶ࡗࡓࠋࡲࡓࠊᚓࡽࢀࡓࢹ࣮ࢱࡢほ್࡜ᮇᚅ್࡜ࡢ㛵ಀࢆᅗ㸰࡟♧ࡋࡓࠋᮏᕞ㒊ࡢ⎔ቃ཰ᐜຊ௨እ࡛஦ᚋศᕸࡢᖜࡣ஦๓ศᕸࡢᖜࡼࡾ⊃ࡲࡗࡓࠋᮏᕞ㒊ࡢ⎔ቃ཰ᐜຊࢆ⾲ࡍኚᩘࡣࠊ⤠ࡽࢀࡿᖜࡀᑡ࡞ࡃࠊ஦๓ศᕸࡢタᐃࡀ᥎ᐃ࡟ᙳ㡪ࢆ୚࠼࡚࠸ࡓ
㸦ᅗ㸯㸫㸰㹟㸧ࠋᮏᕞ㒊ࡢ᥎ᐃ࡟࠾࠸࡚ࠊ⊁⊟࡜᭷ᐖࡢᤕ⋓⋡ࢆㄝ᫂ࡍࡿሀᯝ㢮ࡢ㇏ปࡢಀᩘ㸦prrࠊpyr㸧ࡣࠊ࠸ࡎࢀࡶ 95%ಙ㢗༊㛫࡛࣐࢖ࢼࢫ࡜࡞ࡾࠊ㇏ปᣦᩘࡢቑຍ࡟
ࡼࡿᤕ⋓⋡ࡢపୗࢆㄝ᫂ࡋ࡚࠸ࡓ(⾲㸱㸫㸯)ࠋ
⮬↛ቑຍ⋡ࡣᖺ࡟ࡼࡿኚືࡀ኱ࡁࡃࠊ୰ኸ್࡛ࡳࡿ࡜ࠊᮏᕞ㒊࡛ࡣࠊ᭱ࡶప࠸ᖺ࡛
18.8%㸦2005 ᖺ㸧࠿ࡽ᭱ࡶ㧗࠸ᖺ࡛ 91.5%㸦2009 ᖺ㸧ࠊῐ㊰ᓥ࡛ࡣࠊ24.0㸣㸦2004 ᖺ㸧࠿ࡽ 100.3㸣㸦2009 ᖺ㸧࡜᥎ᐃࡉࢀࡓ㸦⾲㸲㸧ࠋࡲࡓࠊ᥎ᐃᖜ࡟㛵ࡋ࡚ࡶࠊ90%ಙ㢗㝈⏺࡛᭱ࡶᗈ࠸ 2009 ᖺ࡟ࡣࠊ28.1㹼202.7%㸦ᮏᕞ㒊㸧
ࠊ27.0㹼197.2%㸦ῐ㊰ᓥ㸧࡜࡞ࡾࠊ࠿࡞ࡾᗈ࠿ࡗࡓ㸦⾲㸲㸧ࠋᤕ⋓⋡ࡶ୰ኸ್࡛ࠊ⊁⊟ᤕ⋓⋡ࡀ 21.8%࠿ࡽ 37.9%㸦ᮏᕞ㒊㸧ࠊ
12.8㸣࠿ࡽ 18.6%㸦ῐ㊰ᓥ㸧ࠊ᭷ᐖᤕ⋓⋡ࡀ 9.8%࠿ࡽ 23.0%㸦ᮏᕞ㒊㸧
ࠊ10.9㸣࠿ࡽ 20.3%
㸦ῐ㊰ᓥ㸧࡜ᖺ࡟ࡼࡿኚືࡀ኱ࡁ࠿ࡗࡓ㸦⾲㸲㸧ࠋࡉࡽ࡟ࠊࡇࢀࡽࡢ⤖ᯝ࡟ᇶ࡙࠸࡚ィ⟬ࡋࡓಶయᩘ࡜ቑຍಶయᩘࢆ⾲㸳ࠊࡑࢀࡽࡢືྥ
ࢆᅗ㸱ࠊᅗ㸲࡟♧ࡍࠋಶయᩘࡣࠊᖺ࡟ࡼࡿኚືࡣ࠶ࡿࡶࡢࡢࠊᮏᕞ㒊ࠊῐ㊰ᓥ࡜ࡶ࡟2002 ᖺ࠿ࡽ 2010 ᖺ࡟࠿ࡅ࡚ࡣቑຍഴྥ࡟࠶ࡾࠊ2011 ᖺ௨㝆ࡣࠊᮏᕞ㒊࡛ࡣῶᑡࠊῐ
㊰ᓥ࡛ࡣ࡯ࡰᶓࡤ࠸࡜᥎ᐃࡉࢀࡓࠋ2012 ᖺࡢಶయᩘࡣࠊ୰ኸ್࡛ᮏᕞ㒊 16477.8 㢌
㸦90%ಙ㢗㝈⏺࡛ࡣ 8142.8㹼47213.2 㢌⛬ᗘ㸧ࠊῐ㊰ᓥ 8236.8 㢌㸦90%ಙ㢗㝈⏺࡛ࡣ
3870.6㹼19161.4 㢌⛬ᗘ㸧࡜᥎ᐃࡉࢀࡓࠋ රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-3

⾲㸱㸫㸯 ஦ᚋศᕸࡢ⤫ィ㔞㸦ᮏᕞ㒊㸧
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⾲㸱㸫㸰 ஦ᚋศᕸࡢ⤫ィ㔞㸦ῐ㊰ᓥ㸧
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 㻞㻜㻜㻞 ᖺ㻌㻞㻜㻜㻟 ᖺ㻌㻞㻜㻜㻠 ᖺ㻌㻞㻜㻜㻡 ᖺ㻌㻞㻜㻜㻢 ᖺ㻌㻞㻜㻜㻣 ᖺ㻌㻞㻜㻜㻤 ᖺ㻌㻞㻜㻜㻥 ᖺ㻌㻞㻜㻝㻜 ᖺ㻌㻞㻜㻝㻝 ᖺ㻌ᅗ㸯㸫㸯D ⮬↛ቑຍ⋡ࡢ஦๓ศᕸ࡜஦ᚋศᕸ࡜ࡢ㛵ಀ㸦ᮏᕞ㒊㸧ᐇ⥺ࡣ஦ᚋศᕸࢆ◚⥺ࡣ஦๓ศᕸࢆࡑࢀࡒࢀ♧ࡍࠋ
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රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-3

㻞㻜㻜㻞 ᖺ㻌㻞㻜㻜㻟 ᖺ㻌㻞㻜㻜㻠 ᖺ㻌㻞㻜㻜㻡 ᖺ㻌㻞㻜㻜㻢 ᖺ㻌㻞㻜㻜㻣 ᖺ㻌㻞㻜㻜㻤 ᖺ㻌㻞㻜㻜㻥 ᖺ㻌㻞㻜㻝㻜 ᖺ㻌㻞㻜㻝㻝 ᖺ㻌    ᅗ㸯㸫㸯E ⮬↛ቑຍ⋡ࡢ஦๓ศᕸ࡜஦ᚋศᕸ࡜ࡢ㛵ಀ㸦ῐ㊰ᓥ㸧ᐇ⥺ࡣ஦ᚋศᕸࢆ◚⥺ࡣ஦๓ศᕸࢆࡑࢀࡒࢀ♧ࡍࠋ රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-3

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ᅗ㸯㸫㸰D ࣃ࣓࣮ࣛࢱࡢ஦๓ศᕸ࡜஦ᚋศᕸ࡜ࡢ㛵ಀ ᮏᕞ㒊 ᕥୖᅗ㸸⏕ᜥᐦᗘ࡜┠ᧁຠ⋡ࡢẚ⋡ࢆ♧ࡍಀᩘࡢ⮬↛ᑐᩘ್ྑୖᅗ㸸⊁⊟࡟ࡼࡿᤕ⋓⋡ ࣟࢪࢵࢺኚ᥮್ ᕥ୰ᅗ㸸᭷ᐖ࡟ࡼࡿᤕ⋓⋡ ࣟࢪࢵࢺኚ᥮್ ྑ୰ᅗ㸸 ᖺ๓  ᖺࡢ⏕ᜥᩘಶయᩘࡢ⮬↛ᑐᩘ್ᕥୗᅗ㸸⎔ቃ཰ᐜຊࡢ⮬↛ᑐᩘ್ᐇ⥺ࡣ஦ᚋศᕸࢆ◚⥺ࡣ஦๓ศᕸࢆࡑࢀࡒࢀ♧ࡍࠋ රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-3

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






































⾲㸲㸯E ᥎ᐃࡉࢀࡓ⮬↛ቑຍ⋡ LU>L@ ࠊ⎔ቃ཰ᐜຊ㸦N㸧ࠊ┠ᧁຠ⋡ࡢಀᩘ UV>L@ 
ῐ㊰ᓥ㸧
᥎ᐃኚᩘ
LU
LU
LU
LU
LU
LU
LU
LULU
LU
N
UVB
UVB
UVB
UVB
UVB
UVB
UVB
UVB
UVB
UVB
UVB

ᖹᆒᶆ‽೫ᕪ
















 









































 

୰ኸ್










































රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-3

⾲㸲㸰D ᥎ᐃࡉࢀࡓ⊁⊟ᤕ⋓⋡ SU>L@ ࡜᭷ᐖᤕ⋓⋡ S\>L@  ᮏᕞ㒊㸧᥎ᐃኚᩘ
SUB
SUB
SUB
SUB
SUB
SUB
SUB
SUB
SUB
SUB
SUB
S\B
S\B
S\B
S\B
S\B
S\B
S\B
S\B
S\B
S\B
S\B

ᖹᆒ






















ᶆ‽೫ᕪ











































୰ኸ್













































⾲㸲㸰E ᥎ᐃࡉࢀࡓ⊁⊟ᤕ⋓⋡ SU>L@ ࡜᭷ᐖᤕ⋓⋡ S\>L@  ῐ㊰ᓥ㸧᥎ᐃኚᩘSUB
SUB
SUB
SUB
SUB
SUB
SUB
SUB
SUB
SUB
SUB
S\B
S\B
S\B
S\B
S\B
S\B
S\B
S\B
S\B
S\B
S\B

ᖹᆒ






















ᶆ‽೫ᕪ










































 ୰ኸ್










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
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









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

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

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








┠ᧁຠ⋡

⊁⊟ᤕ⋓

᭷ᐖᤕ⋓

ᅗ㸰㸫㸯 ほ್࡜ᮇᚅ್࡜ࡢ㛵ಀ㸦ᮏᕞ㒊㸧ୖᅗ㸸┠ᧁຠ⋡ࡢほ ್࡜ᮇᚅ್ࠊ୰ᅗ㸸⊁⊟ᤕ⋓ࡢほୗᅗ㸸᭷ᐖᤕ⋓ࡢほ ್࡜ᮇᚅ್
୰ኸ್࡜ ಙ㢗㝈⏺ࠊ㸣ಙ㢗㝈⏺ࢆ♧ࡍࠋ



 

್࡜ᮇᚅ್

රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-3



┠ᧁຠ⋡

⊁⊟ᤕ⋓

᭷ᐖᤕ⋓


ᅗ㸰㸫㸰 ほ್࡜ᮇᚅ್࡜ࡢ㛵ಀ㸦ῐ㊰ᓥ㸧ୖᅗ㸸┠ᧁຠ⋡ࡢほ ್࡜ᮇᚅ್ࠊ୰ᅗ㸸⊁⊟ᤕ⋓ࡢほୗᅗ㸸᭷ᐖᤕ⋓ࡢほ ್࡜ᮇᚅ್
୰ኸ್࡜ ಙ㢗㝈⏺ࠊ㸣ಙ㢗㝈⏺ࢆ♧ࡍࠋ



 

್࡜ᮇᚅ್

රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-3ᅗ㸱㸫㸯 රᗜ┴ࡢ࢖ࣀࢩࢩࡢ᥎ᐃ⏕ᜥಶయᩘࡢືྥ㸦ᮏᕞ㒊㸧୰ኸ್࡜ ಙ㢗㝈⏺ࠊ㸣ಙ㢗㝈⏺ࢆ♧ࡍࠋ






ᅗ㸱㸫㸰 රᗜ┴ࡢ࢖ࣀࢩࢩࡢ᥎ᐃ⏕ᜥಶయᩘࡢືྥ㸦ῐ㊰ᓥ㸧୰ኸ್࡜ ಙ㢗㝈⏺ࠊ㸣ಙ㢗㝈⏺ࢆ♧ࡍࠋ


 

රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-3ᅗ㸲㸫㸯 රᗜ┴ࡢ࢖ࣀࢩࢩࡢ᥎ᐃቑຍಶయᩘࡢືྥ㸦ᮏᕞ㒊㸧୰ኸ್࡜ ಙ㢗㝈⏺ࠊ㸣ಙ㢗㝈⏺ࢆ♧ࡍࠋ






ᅗ㸲㸫㸰 රᗜ┴ࡢ࢖ࣀࢩࢩࡢ᥎ᐃቑຍಶయᩘࡢືྥ㸦ῐ㊰ᓥ㸧୰ኸ್࡜ ಙ㢗㝈⏺ࠊ㸣ಙ㢗㝈⏺ࢆ♧ࡍࠋ





 

රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-3

 ࠊቑຍಶయᩘ LQF>L@㸦ᮏᕞ㒊㸧⾲㸳㸯 ᥎ᐃࡉࢀࡓ⏕ᜥಶయᩘ㹌㹙㹧㹛᥎ᐃኚᩘ1
1
1
1
1
1
1
1
1
1
1
LQF
LQF
LQF
LQF
LQF
LQF
LQF
LQF
LQF
LQF
LQF

ᖹᆒ




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
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
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
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


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



ᶆ‽೫ᕪ





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
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







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


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
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
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୰ኸ್

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


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
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






⾲㸳㸫㸰 ᥎ᐃࡉࢀࡓ⏕ᜥಶయᩘ 1>L@ࠊቑຍಶయᩘ LQF>L@㸦ῐ㊰ᓥ㸧᥎ᐃኚᩘ1
1
1
1
1
1
1
1
1
1
1
LQF
LQF
LQF
LQF
LQF
LQF
LQF
LQF
LQF
LQF
LQF

ᖹᆒ










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









ᶆ‽೫ᕪ

























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


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









 

୰ኸ್


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රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-3

⪃ᐹ࢖ࣀࢩࢩࡣࠊ⮬↛ቑຍ⋡ࡸ⏕ᜥಶయᩘࡢᖺḟኚືࡀ኱ࡁࡃࠊ୰ኸ್࡛ࡳࡿ࡜᭱኱್࡜
᭱ᑠ್ࡢᕪࡣᮏᕞ㒊࡛ 72.7㸣ࠊῐ㊰ᓥ࡛ࡣ 76.3㸣࠶ࡿࠋ᥎ᐃಶయᩘࡢഴྥ࡜ࡋ࡚ࡣࠊ
ᮏᕞ㒊ࠊῐ㊰ᓥ࡜ࡶ࡟ࠊ2010 ᖺࡲ࡛ࡣቑຍഴྥ࡟࠶ࡗࡓࡀࠊ2010 ᖺ௨㝆ࡣࠊᮏᕞ㒊࡛ࡣῶᑡഴྥࠊῐ㊰ᓥ࡛ࡣ࡯ࡰᶓࡤ࠸≧ែ࡛࠶ࡿ࡜᥎ᐃࡉࢀࡓࠋࡇࡢ⤖ᯝࡣࠊᮏᕞ㒊ࠊῐ㊰ᓥ࡜ࡶ 2010 ᖺ࠿ࡽࡢᤕ⋓ᩘࡢቑຍ࡟ࡼࡾᑡ࡞ࡃ࡜ࡶಶయᩘࡢቑຍࡣ㜵ࡆ࡚࠸ࡿࡇ࡜
ࢆ♧ࡋ࡚࠸ࡿࠋࡓࡔࡋࠊ㎰ᴗ⿕ᐖࡢ⛬ᗘࡣࠊᮏᕞ㒊࣭ῐ㊰ᓥ࡜ࡶ࡟ࠊ2010 ᖺ௨㝆ࠊῶᑡࡋ࡚࠸࡞࠸㸦ᯇᮏ࡯࠿ 2014㸧ࠊ⿕ᐖࡀ῝้࡞ᆅᇦ࡛ࡣࠊࡉࡽ࡞ࡿᤕ⋓ᙉ໬ࡸ㜵ㆤᰙ➼ࡢᑐ⟇ࡢ඘ᐇࡀᚲせ࡛࠶ࡿࠋẖᖺࡢ࢖ࣀࢩࢩࡢቑຍ㢌ᩘࢆぢࡿ࡜ࠊ2011 ᖺࡢ᥎ᐃቑຍಶయᩘࡢ୰ኸ್ࡣࠊᮏᕞ㒊
࡛ 7408.8 㢌㸦90㸣ಙ㢗㝈⏺࡛ࡣ 1268.4 㢌㹼12995.9 㢌㸧ࠊῐ㊰ᓥ࡛ࡣ 3158.2 㢌㸦90㸣ಙ㢗㝈⏺࡛ࡣ 712.0 㢌㹼6149.0 㢌㸧࡛࠶ࡿࠋ2011 ᖺࡢᤕ⋓ᩘࡣᮏᕞ㒊 9147 㢌ࠊῐ㊰
ᓥ 3152 㢌࡞ࡢ࡛ࠊ⌧ᅾࡢ᥎ᐃಶయᩘ࡟ᑐࡍࡿᤕ⋓⋡ࢆ⥅⥆ࡍࡿ㝈ࡾࠊ୰ኸ್࡛ぢࡿ㝈ࡾࠊಶయᩘࡢᛴ㏿࡟ῶᑡࡍࡿྍ⬟ᛶࡣప࠸ࠋࡓࡔࡋࠊ90㸣ಙ㢗㝈⏺ࢆࡳࡿ࡜᥎ᐃㄗᕪࡶ
ぢ㎸ࢇ࡛࠾ࡃᚲせࡀ࠶ࡿࡓࡵࠊ௒ᚋࡢᤕ⋓ᩘࡸ┠ᧁຠ⋡ࠊ⿕ᐖ≧ἣ࡞࡝ࢆぢ࡞ࡀࡽࠊẖᖺࡢ᥎ᐃࢆ⧞ࡾ㏉ࡋࠊ㡰ᛂⓗ࡟᪉㔪ࢆタᐃࡋ࡚࠸ࡃᚲせࡀ࠶ࡿࠋ ᮏㄽᩥ࡛⏝࠸ࡓ Harvested-based model ࢆᇶᮏࣔࢹࣝ࡜ࡋࡓ࣋࢖ࢬ᥎ᐃ࡟ࡼࡿಶయ
ᩘ᥎ᐃἲࡣࠊ㸧ᤕ⋓ᩘࢆࡣࡌࡵࠊಶయᩘ᥎ᐃࡢᣦᶆ࡜࡞ࡾ࠺ࡿ」ᩘࡢᣦᶆࢆᰂ㌾࡟᥎ᐃࣔࢹࣝ࡟⤌ࡳ㎸ࡳࡸࡍ࠸ࠊ㸧⮬↛ቑຍ⋡ࠊᤕ⋓⋡࡞࡝ࡢ᥎ᐃኚᩘ࡟᪤Ꮡࡢ▱ぢࢆ஦๓ศᕸ࡜ࡋ࡚཯࡛ᫎࡁࡿࠊ㸧ಙ㢗༊㛫ࢆకࡗ࡚ኚᩘࢆ᥎ᐃ࡛ࡁࡿࡢ࡛ྜពᙧᡂ࡟฼⏝ࡋࡸࡍ࠸ࠊ㸧ಶయᩘࠊቑຍ⋡࡞࡝ࡢಶయ⩌ࣃ࣓࣮ࣛࢱࡔࡅ࡛࡞ࡃࠊᤕ⋓⋡࡞࡝ࡢಶయ
⩌⟶⌮஦ᴗࢆ⾜࠺ୖ࡛㔜せ࡞ኚᩘࠊ᥎ᐃ್ࡢศᩓࡸࢹ࣮ࢱࡢほㄗᕪࡶ᥎ᐃ࡛ࡁࡿࠊ࡜

࠸࠺฼Ⅼࡀ࠶ࡿࠋ௚᪉ࠊẖᖺ᪂ࡓ࡞ࢹ࣮ࢱࢆ㏣ຍࡍࡿࡇ࡜࡛ࠊ㐣ᖺᗘࡢ᥎ᐃ್࡟㛵ࡋ࡚ࡶࠊ๓ᖺ࡟ฟࡋࡓ᥎ᐃ್࡜␗࡞ࡿ್࡟࡞ࡿࡇ࡜ࡶ࠶ࡿࠋ㐣ཤ࡟ࡉ࠿ࡢࡰࡗ࡚᥎ᐃ್ࢆぢ┤ࡍࡇ࡜ࡣࠊ୍⯡ⓗ࡞⌮ゎࢆᅔ㞴࡟ࡍࡿᜍࢀࡶ࠶ࡾࠊࡇࡢᡭἲࡢࢹ࣓ࣜࢵࢺ࡜ࡶ࠸࠼ࡿࠋ
ࡓࡔࡋࠊࡑࡢ᫬Ⅼ࡛ᚓࡽࢀࡿࢹ࣮ࢱ࡟ᛂࡌ࡚☜⋡ㄽⓗ࡟᭱ࡶጇᙜ࡞☜⋡ศᕸࢆ᥎ᐃࡍࡿ
ࡢࡣ࣋࢖ࢬ᥎ᐃࡢ㛗ᡤࡢ୍ࡘ࡛࠶ࡿࠋ≉࡟ࠊ㔝⏕ື≀ࡢ⟶⌮࡟ᚲせ࡞㛗ᮇⓗ࡞どⅬ࡛ぢࡿ࡜ࠊࡇࡢࡼ࠺࡞᥎ᐃ್ࡢ㡰ᛂⓗ࡞㏲ḟಟṇࡣࠊ⌧ᐇ࡜᥎ᐃࡢ஋㞳ࢆ㜵ࡄࡓࡵ࡟ᚲせ୙ྍḞ࡞సᴗ࡛࠶ࡿࠋ ࡇࡢ᥎ᐃ᪉ἲ࡛ࡣࠊู㏵ㄪᰝࢆ⾜ࡗ࡚ಶయᩘࡸ⏕ᜥᐦᗘࡢᣦᶆࡀᚓࡽࢀࢀࡤࠊࡑࢀࢆ
ࣔࢹࣝ࡟⤌ࡳ㎸ࡴࡇ࡜ࡀ࡛ࡁࡿࡋࠊ⦾Ṫ⋡࡞࡝⮬↛ቑຍ⋡࡟㛵ࡍࡿ᝟ሗࡀᚓࡽࢀࢀࡤࠊ
ࡑࢀࢆ஦๓ศᕸ࡟⤌ࡳ㎸ࡳ᥎ᐃ⢭ᗘࢆୖࡆࡿࡇ࡜ࡶྍ⬟࡛࠶ࡿࠋࡇࡢࡼ࠺࡞㛵㐃ࡍࡿㄪᰝ◊✲ࡢ㐍ᒎࢆཧ↷ࡋ࡞ࡀࡽࠊ᥎ᐃᢏ⾡ࡢᨵၿࢆ⥆ࡅࡿࡇ࡜ࡀồࡵࡽࢀࡿࠋࡲࡓࠊ⌧᫬
Ⅼ࡛ࡣ᭱ၿࡢດຊࢆࡋࡓୖ࡛ࡢ᥎ᐃ್ࢆ✚ᴟⓗ࡟බ⾲ࡋࠊ㛵ಀ⪅࡟ࡼࡿ᳨ドࡸ㏲ḟಟṇࢆཷࡅ࡞ࡀࡽࠊ࢖ࣀࢩࢩࡢಶయ⩌ືែ࡟㛵ࡍࡿ⌧≧ㄆ㆑ࢆඹ᭷ࡋ࡚࠸ࡃᚲせࡀ࠶ࡿࠋ රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-3ᘬ⏝ᩥ⊩⸨ᮌ኱௓࣭ᶓᒣ┿ᘪ࣭ᆏ⏣ᏹᚿ. 2011. රᗜ┴ෆ࡟࠾ࡅࡿࣈࢼ⛉ᶞᮌ 3 ✀ࡢሀᯝࡢ㇏ป
࡜ࢶ࢟ࣀ࣡ࢢ࣐ࡢ㣵㈨※࡜ࡋ࡚ࡢホ౯.ࠕරᗜ┴࡟࠾ࡅࡿࢶ࢟ࣀ࣡ࢢ࣐ࡢಖㆤ⟶⌮ࡢ⌧≧࡜ㄢ㢟ࠖ㸪රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 3 ྕ㸪pp.39-49. රᗜ┴᳃ᯘື≀◊✲ࢭ
ࣥࢱ࣮.
Geweke, J. 1992. Evaluating the Accuracy of Sampling-Based Approaches to the
Calculation of Posterior Moments. In Bayesian Statistics 4 (Bernardo, J. M., Berger,
J. O., Dawid, A. P. and Smith, A. F. M. eds), pp.169-193, Oxford Univ Press, Oxford.
Gilks, W. R., Richardson, S. and Spiegelhalter, D. 1996. Markov Chain Monte Carlo in
Practice. Chapman & Hall/CRC, New York, USA, 512pp.
රᗜ┴㸬2012㸬➨ 2 ᮇ࢖ࣀࢩࢩಖㆤ⟶⌮ィ⏬㸬රᗜ┴㸪⚄ᡞ, 18pp㸬
⎔ቃ┬⮬↛⎔ቃᒁ⏕≀ከᵝᛶࢭࣥࢱ࣮㸬2011. ᖹᡂ 22 ᖺᗘ⮬↛⎔ቃಖ඲ᇶ♏ㄪᰝ≉ᐃ့ங㢮⏕ᜥ≧ἣㄪᰝཬࡧㄪᰝయไᵓ⠏᳨ウᴗົሗ࿌᭩㸬411pp㸬
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ᑠᑎ♸஧࣭⚄ᓮఙኵ. 2001. ᓥ᰿┴▼ぢᆅ᪉࡟࠾ࡅࡿࢽ࣍ࣥ࢖ࣀࢩࢩࡢ㣗ᛶ࠾ࡼࡧᰤ㣴≧ែࡢᏘ⠇ⓗኚ໬㸪㔝⏕⏕≀ಖㆤ 6(2)㸸109-117.
ᯇᮏᓫ࣭ᆏ⏣ᏹᚿ࣭ᓊᮏᗣ㄃㸬2014㸬රᗜ┴࡟࠾ࡅࡿࢽ࣍ࣥ࢖ࣀࢩࢩࡢ⏕ᜥศᕸ࡜㎰ᴗ⿕ᐖࡢ㛵ಀ㸬ࠕරᗜ┴࡟࠾ࡅࡿࢽ࣍ࣥ࢖ࣀࢩࢩࡢ⟶⌮ࡢ⌧≧࡜ㄢ㢟ࠖ, රᗜ࣡࢖ࣝࢻࣛ࢖ࣇ
ࣔࣀࢢࣛࣇ 6 ྕ㸪pp.9-20㸬රᗜ┴᳃ᯘື≀◊✲ࢭࣥࢱ࣮㸬
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ᆏ⏣ᏹᚿ㸬2010㸬㎰ᴗ⿕ᐖࡢ≧ἣᢕᥱ࡜㎰ᴗ㞟ⴠ࢔ࣥࢣ࣮ࢺ㸬ࠕ㎰ᴗ㞟ⴠ࢔ࣥࢣ࣮ࢺ࠿ࡽࡳ
ࡿࢽ࣍ࣥࢪ࣭࢝࢖ࣀࢩࢩࡢ⿕ᐖ࡜ᑐ⟇ࡢ⌧≧ࠖ㸪රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 2 ྕ㸪
pp.1-4㸬රᗜ┴᳃ᯘື≀◊✲ࢭࣥࢱ࣮㸬ᆏ⏣ᏹᚿ࣭ᓊᮏᗣ㄃࣭㛵㤶ዉᏊ. 2012. ࢖ࣀࢩࢩࡢಶయ⩌ືែࡢ᥎ᐃ㸦රᗜ┴ᮏᕞ㒊 2011
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ୖ⏣๛ᖹ࣭ຍ⸨❳┿࣭ఀ࿃⏣ᏹṇ࣭⃝⏣ㄔ. 2013. ⊁⊟⪅ࡢࢽ࣍ࣥ࢖ࣀࢩࢩ㸦Sus scrofa

leucomystax㸧ࡢᤕ⋓ᩘ࡟ᙳ㡪ࡍࡿせᅉ. ့ங㢮⛉Ꮫ 53(1): 17-29.
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Saitoh T. and Hirakawa H. 2008. Harvest-based Bayesian estimation of sika deer
populations using state-space models. Population Ecology 50: 131-144.

 රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-4➨ 㸲 ❶            ㎰ᆅ࡟㞄᥋ࡋ࡚⏕ᜥࡍࡿࢽ࣍ࣥ࢖ࣀࢩࢩࡢ   ຍᐖ⾜ືࡢゎᯒᶓᒣ┿ᘪ 1,2࣭Ụ⸨බಇ 3࣭ᮌୗ⿱⨾Ꮚ 3
せ Ⅼ࣭ ࢖ࣀࢩࢩ࡟ࡼࡿ㎰ᴗ⿕ᐖࡀ῝้࡞ᆅᇦ࡛࠶ࡿ⠛ᒣᕷ࡜す⬥ᕷ࡟࠾࠸࡚ࠊ㎰ᆅ࡟㞄᥋ࡋ࡚⏕ᜥࡍࡿ࢖ࣀࢩࢩࡢ⾜ືࣃࢱ࣮ࣥࢆ᫂ࡽ࠿࡟ࡋࡓࠋ
࣭ 9 㢌ࡢ࢖ࣀࢩࢩ࡟ GPS 㤳㍯ࢆ⿦╔ࡋࠊ✵㛫ⓗ࡞⾜ືࢆ㏣㊧ࡋࡓࠋ㏣㊧ᮇ㛫୰ࡢ⾜ືᅪࡢᖹᆒࡣࠊ3.39±2.89km2 ࡜኱ᆺ⋇࡜ࡋ࡚ࡣẚ㍑ⓗ⊃࠸⾜ືᅪ࡛࠶ࡗࡓࠋ
࣭ ㎰ᆅ㏆ࡃ࡛ᤕ⋓ࡉࢀࡓ࢖ࣀࢩࢩࡣࠊ㎰ᆅ࡟㞄᥋ࡋࡓᗈⴥᶞᯘࢆ୺࡞άືᇦ࡜ࡋ࡚
࠾ࡾࠊ㎰ᆅ࡬ࡢฟἐࡣࠊ୺࡟ኪ㛫ࠊ㞟ⴠᰙࡢ࡞࠸ሙᡤ࡟㞟୰ࡋ࡚ฟἐࡋ࡚࠸ࡓࠋ
ఫᏯᆅ㏆ࡃࡢ㫽⋇ಖㆤ༊ෆ࡛άືࡋࡓಶయࡣࠊ᪥୰࡛ࡶᕷ⾤ᆅ࡟ฟἐࡋ࡚࠸ࡓࠋ
࣭ ㎰ᆅ࡬ࡢฟἐࣃࢱ࣮ࣥࡣࠊྠࡌሙᡤ࡟⧞ࡾ㏉ࡋฟἐࡍࡿࣃࢱ࣮ࣥ࡜࠸ࡃࡘ࠿ࡢሙ
ᡤࢆኚ࠼࡞ࡀࡽฟἐࡍࡿࣃࢱ࣮ࣥࡀ࠶ࡗࡓࠋ࣭ 㞟ⴠ༢఩࡛ࡢ㜵ㆤᰙࡢタ⨨ࡸ࣓ࣥࢸࢼࣥࢫࡀ⾜ࡁᒆ࠸࡚࠸ࡿ㟁Ẽᰙ࡛ࡣࠊຠᯝⓗ࡟ฟἐࢆᢚไ࡛ࡁ࡚࠸ࡿࡀࠊ㏆㞄ࡢಶேᑐᛂࡢ㎰ᆅ࡟⿕ᐖࡀ㞟୰ࡋ࡚࠾ࡾࠊ1 ಶయࡀ⧞ࡾ㏉ࡋฟἐࡋ࡚࠸ࡿ౛ࡶ࠶ࡾࠊ㎰ᆅ࡬ࡢᇳ╔⾜ືࡀ♧၀ࡉࢀࡓࠋ࣭ ࢖ࣀࢩࢩࡣ㎰ᆅ࡟㞄᥋ࡋࡓᗈⴥᶞᯘ࡛ẚ㍑ⓗ⊃࠸⾜ືᅪෆ࡟⏕ᜥࡍࡿࡇ࡜ࡀ࡛ࡁ
ࡿࠋ᳃ᯘ࡟㞄᥋ࡋࡓ㎰ᆅ࡛ࡣࠊᗈᇦⓗ࡞㜵㝖ࢹࢨ࢖ࣥࡀᚲせ࡛࠶ࡿࠋ

Key words㸸⾜
⾜ືᅪࠊGPS 㤳㍯ࠊฟἐࣃࢱ࣮ࣥࠊ㞟ⴠᰙࠊ㏣࠸ᡶ࠸

ࡣࡌࡵ࡟
㔝⏕ື≀ࡢಖㆤ⟶⌮ࢆ㐍ࡵࡿ࠺࠼࡛ࠊಶయᩘ⟶⌮ࠊ⏕ᜥᆅ⟶⌮ࠊ⿕ᐖ⟶⌮ࡢ 3 ࡘࡀ㔜せ࡞ഃ㠃࡛࠶ࡿࡀࠊࢽ࣍ࣥ࢖ࣀࢩࢩ㸦Sus scrofa leucomystax㸹௨ୗ࢖ࣀࢩࢩ࡜⾲グ㸧ࡢ⿕ᐖᑐ⟇ࡢ⌧≧ࡣࠊ᭷ᐖ㥑㝖ᤕ⋓࡟ࡼࡿಶయᩘㄪᩚࠊ㜵ㆤᰙ࡞࡝࡟ࡼࡿ⿕ᐖ㜵㝖ࡀ୰ᚰⓗ࡞᪉⟇࡜࡞ࡗ࡚࠸ࡿࠋࡋ࠿ࡋࠊࡇࢀࡽࡢᑐ⟇ࡣࠊ⌧≧ࡢᡭἲࡢࡳ࡛ࡣࠊ඘ศ࡞ຠᯝ
ࡀⓎ᥹ࡉࢀ࡚࠸࡞࠸ࠋ᪥ᮏྛᆅ࡛࢖ࣀࢩࢩ࡟ࡼࡿ㎰ᴗ⿕ᐖࡀ῝้໬ࡋ࡚࠸ࡿ㸦㎰ᯘỈ⏘┬⏕⏘ᒁ 㸧ࠋරᗜ┴ࡢሙྜࠊྂࡃ࠿ࡽ࢖ࣀࢩࢩࡣ⊁⊟㈨※࡜ࡋ࡚ࡢ౯್ࡀ㧗࠸ࡇ࡜࠿ࡽ(኱᮰㸫ఀ⸨࣭⚄ᓮ 1998㸹ᮅ᪥⛱ 1985)ࠊ≉࡟⊁⊟ᮇ࡟ᙉ࠸ᤕ⋓ᅽࡀ࠿࠿ࡿࡶࡢࡢࠊ
රᗜ┴᳃ᯘື≀◊✲ࢭࣥࢱ࣮࣭ රᗜ┴❧኱Ꮫ⮬↛࣭⎔ቃ⛉Ꮫ◊✲ᡤ࣭//3 㔛ᒣ㫽⋇◊✲ᡤ රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-4㎰స≀ࡢ⿕ᐖࡣᖺ㛫㸰൨෇ࢆ㉸࠼ࠊࢽ࣍ࣥࢪ࢝㸦Cervus nippon㸧 ࡟ḟ࠸࡛㸰␒┠࡟ከ࠸⿕ᐖ࡛࠶ࡾࠊ౫↛࡜ࡋ࡚῝้࡞ၥ㢟࡜࡞ࡗ࡚࠸ࡿ㸦රᗜ┴ 2013㸧ࠋࡋࡓࡀࡗ࡚ࠊ⌧≧ࡢᑐ⟇ࡢၥ㢟Ⅼࢆ᫂ࡽ࠿࡟ࡋࠊࡼࡾຠᯝⓗ࡞⿕ᐖ㜵Ṇᑐ⟇ࡀᚲせ࡞ẁ㝵࡜࠸࠼ࡿࠋ
ಶయᩘ⟶⌮࡟ࡼࡿ⿕ᐖᢚไࡀຠᯝⓗ࡟ᶵ⬟ࡋ࡞࠸せᅉ࡜ࡋ࡚ࠊࢶ࢟ࣀ࣡ࢢ࣐㸦Ursus

thibetanus㸧㸦ᶓᒣ࡯࠿ 2011㸧ࡸࢽ࣍ࣥࢨࣝ㸦Macaca fuscata㸧ࡢࡼ࠺࡟࢖ࣀࢩࢩ࡟࠾
࠸࡚ࡶຍᐖಶయ࡜㠀ຍᐖಶయࡢ⾜ື≉ᛶࡢಶయᕪࡀ኱ࡁࡃࠊຍᐖಶయ࡬ࡢᤕ⋓ᅽࡀప࠸ࠊ
࠶ࡿ࠸ࡣ㐺ษ࡞ㄏᘬ≀㝖ཤࡸ㜵ㆤᰙタ⨨࡞࡝ࡢ⿕ᐖᑐ⟇ࡀຍᐖಶయࡢ⾜ືࢆᢚไ࡛ࡁ
࡚࠸࡞࠸࡞࡝ࡢྍ⬟ᛶࡀ⪃࠼ࡽࢀࡿࠋᐇ㝿ࠊ᭷ᐖᤕ⋓ࡢᐇ᪋≧ἣࢆࡳࡿ࡜ࠊᤕ⋓ࡋࡸࡍ࠸ሙᡤ࡛ࡢᤕ⋓άືࡀከࡃࠊຍᐖಶయࢆᤕ⋓࡛ࡁ࡚࠸࡞࠸ྍ⬟ᛶࡀ⪃࠼ࡽࢀ࡚࠸ࡿࠋࡲ
ࡓࠊ࢖ࣀࢩࢩࡣᏛ⩦⬟ຊࡀ㧗ࡃ㎰ᆅ࡬ࡢ౫Ꮡᗘࡀ㧗ࡲࡿ࡜⧞ࡾ㏉ࡋ⿕ᐖࢆⓎ⏕ࡉࡏࡿ࡞࡝ࡢ⾜ືࡀ▱ࡽࢀ࡚࠸ࡿ㸦Ụཱྀ 2003㸹2008㸧ࠋࡉࡽ࡟ࠊ࢖ࣀࢩࢩࡢ୺࡞άືᇦ࡜㎰⪔ᆅ࡜ࡢ㞄᥋≧ἣࡀ࡝ࡢࡼ࠺࡞㓄⨨࡟࠶ࡿࡢ࠿ࠊఇᜥሙࡸ㌟ࢆ₯ࡵࡿሙᡤࡀ㎰ᆅ㏆ࡃ࡟࡞࠸࠿࡞࡝ࡢⅬࡀᮍ᳨ウ࡛ࠊ✵㛫ⓗ࡞ᵓ㐀࡟ࡼࡾ࢖ࣀࢩࢩࡀ㎰ᆅ࡟౵ධࡋࡸࡍ࠸≧ἣࡀᨺ⨨ࡉࢀ࡚࠸ࡿࢣ࣮ࢫࡀከ࠸㸦Ụཱྀ 2008㸧ࠋරᗜ┴ෆ࡟࠾࠸࡚ࡶ࢖ࣀࢩࢩࡢಶయᩘᐦᗘᣦᶆࡀప࠸ᆅᇦ࡛ࠊ⿕ᐖࡀ῝้࡜࡞ࡿᆅᇦ
ࡀⓎ⏕ࡋ࡚࠸ࡿࡇ࡜࠿ࡽ㸦ᆏ⏣࡯࠿ࠊ2008㸧ࠊ≉ᐃࡢຍᐖಶయ࡟ࡼࡾ῝้࡞⿕ᐖࡀⓎ⏕ࡋ࡚࠸ࡿᆅᇦࡀ࠶ࡿ࡜⪃࠼ࡽࢀࡿࠋ⿕ᐖࡀ῝้࡞ᆅᇦ࡟࠾࠸࡚ࡣࠊᨺᲠ㎰⪔ᆅࡶቑຍࡋ
࡚࠾ࡾࠊ࢖ࣀࢩࢩ࡟࡜ࡗ࡚฼⏝ࡋࡸࡍ࠸✵㛫ᵓ㐀ࡀቑຍࡋ࡚࠸ࡿ≧ἣࡶ⿕ᐖࢆຍ㏿ࡋ࡚࠸ࡿ࡜⪃࠼ࡽࢀࡿࠋ
ࡋ࠿ࡋࠊ࢖ࣀࢩࢩ࡟ࡘ࠸࡚ࡣࠊ⾜ືᅪࡸ㑅ዲࡍࡿ⎔ቃࠊຍᐖಶయࡢ㎰ᆅ࡬ࡢ౵ධ≧ἣ࡞࡝⾜ື≉ᛶࡸ⏕ᜥᆅ⎔ቃ࡞࡝⿕ᐖࢆ㍍ῶࡍࡿࡓࡵࡢᑐ⟇࡟ᚲせ࡞ᇶᮏⓗ᝟ሗࡀ㝈ࡽࢀ࡚࠾ࡾ㸦ᮏ⏣࡯࠿ 2008㸹㔝ᮏ࡯࠿ 2010㸧ࠊຠᯝⓗ࡞⿕ᐖ㜵㝖᪉ἲ࡞࡝ࡢ᳨ウࡀ୙༑ศ࡛࠶ࡿࠋࡇࢀࡽࢆゎỴࡍࡿࡓࡵ࡟ࡣࠊ࢖ࣀࢩࢩࡢຍᐖ⾜ືࣃࢱ࣮ࣥࡸ࢖ࣀࢩࢩࡢ⏕ᜥᆅ฼⏝ࢆゎᯒࡋࠊ⿕ᐖⓎ⏕࣓࢝ࢽࢬ࣒ࢆ᫂ࡽ࠿࡟ࡋࡓ࠺࠼࡛ࠊ㎰⪔ᆅ࡬ฟἐࡋ࡟ࡃ࠸⎔ቃࡢ๰ฟࡸຠᯝⓗ࡞⿕ᐖ⟶⌮࡟╔ᡭࡍࡿࡇ࡜ࡀ㔜せ࡟࡞ࡗ࡚ࡃࡿࠋࡲࡓࠊᤕ⋓࡟ᡂຌࡋ࡞ࡃ࡚ࡶ
⊁⊟άື⮬యࡀ࢖ࣀࢩࢩࡢ⏕ᜥᆅ฼⏝ࣃࢱ࣮ࣥࢆኚ໬ࡉࡏࠊຍᐖ⾜ືࡢపῶ࡟ຠᯝࡀ࠶
ࡿࡢ࠿ࢆᢕᥱࡍࡿࡇ࡜ࡶࠊ㐺ษ࡞㜵㝖ἲࢆ㑅ᢥࡍࡿୖ࡛㔜せ࡛࠶ࡿࠋᑗ᮶ⓗ࡟ࡣ㜵㝖≟ࢆ⏝࠸ࡓ㜵㝖ἲ࡞࡝ࢆ☜❧ࡍࡿࡇ࡜ࡀ᳨ウࡉࢀ࡚࠸ࡿࡀࠊ࢖ࢾ࡟ࡼࡿ㏣࠸ᡶ࠸࡟ࡼࡗ࡚ࠊ⾜ື≉ᛶࡸຍᐖ⾜ືࡀ࡝ࡢ⛬ᗘಟṇࡉࢀࡿࡢ࠿࡞࡝ࢆ᫂ࡽ࠿࡟ࡍࡿࡇ࡜ࡶᚲせ࡜࡞ࡿࠋ௨ୖࢆࡩࡲ࠼ࠊᮏ◊✲࡛ࡣ㎰ᆅ࡟㞄᥋ࡋ࡚⏕ᜥࡍࡿ࢖ࣀࢩࢩࢆᑐ㇟࡟⾜ືᅪࡸ⎔ቃ㑅ᢥࠊ㎰ᆅฟἐ᫬้➼ࠊຍᐖಶయࡢ⾜ື≉ᛶ࡞࡝ࢆヲ⣽࡟ᢕᥱࡍࡿࡇ࡜ࢆ┠ⓗ࡜ࡋࡓࠋࡲ
ࡓࠊ࢖ࣀࢩࢩࡀᐇ㝿࡟ฟἐࡋࡓ㎰ᆅࡢ⿕ᐖᑐ⟇≧ἣࢆㄪ࡭ࠊฟἐᆅ࡜㠀ฟἐᆅ࡟࠾ࡅࡿ
㜵㝖ᑐ⟇ࡢᕪ␗ࢆ᳨ドࡋࡓࠋࡉࡽ࡟࢖ࣀࢩࢩࡀ⏕ᜥࡍࡿ࢚ࣜ࢔࡟࠾࠸࡚ࠊ᭷ᐖᤕ⋓άືࠊ⊁⊟άືࡀ࢖ࣀࢩࢩࡢ⾜ື࡟ཬࡰࡍᙳ㡪ࢆ᳨ドࡋࡓࠋ රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-4

㸬ㄪᰝᆅ
ㄪᰝᆅㄪᰝࡣࠊරᗜ┴す⬥ᕷ㸦ᮾ⤒ 134°58’ࠊ໭⦋ 35°00’㸧ཬࡧ⠛ᒣᕷ㸦ᮾ⤒ 135°12‘ࠊ໭⦋ 35°04’㸧࡛⾜ࡗࡓ㸦ᅗ㸯㸧ࠋㄪᰝᆅࡣࠊ࠸ࡎࢀࡶேཱྀつᶍࡣ㸲୓ேࢆ㉸࠼࡚࠸ࡿᆅᇦ
࡛࠶ࡿࡀࠊᕷ⾤ᆅ㏆ࡃ࠿ࡽ⏣ᅬᆅᖏࡀᗈࡀࡾࠊẚ㍑ⓗ࡞ࡔࡽ࠿࡞ᒣ㛫ᆅᖏ࡟ᅖࡲࢀࠊ࢖
ࣀࢩࢩ࡟ࡼࡿ⿕ᐖࡀከ࠸ᆅᇦ࡛࠶ࡿ㸦ᅗ㸰㸧ࠋẼೃࡣ࠸ࡎࢀࡶ℩ᡞෆẼೃ༊࡟ᒓࡋࠊᖺᖹᆒẼࡣ 14Υ ࡛ ࠶ ࡿ ࠋ ᳜ ⏕ ࡣ ࢥ ࢼ ࣛ

Quercus serrata ࢆ୺య࡜ࡋࡓᗈⴥᶞᯘࡀከ ࡃ ࠊ ࡑ ࡢ ෆ 㒊 ࡟ ࢫ ࢠ 㸦 Cryptomeria

japonica 㸧࣭ ࣄ ࣀ ࢟ 㸦 Chamaecyparis
obtuse㸧ࡢ᳜ᯘᆅࡀࣃࢵࢳୖ࡟㓄⨨ࡉࢀ࡚࠸ࡿࠋす⬥ᕷ࡟࠾ࡅࡿ࢖ࣀࢩࢩ⿕ᐖࡣᒁᡤⓗ࡛࠶ࡿࡶࡢࡢᕷ⾤ᆅ㏆ࡃ࡟࢖ࣀࢩࢩࡀ⏕ᜥࡋࠊ࠿ࡘ᭷ᐖᤕ⋓άືࡀᐃᮇⓗ࡟ᐇ᪋ࡉ

⠛ᒣᕷ
す⬥ᕷ

ࢀ࡚࠸ࡓᆅᇦ࡛࠶ࡿࠋᕷ⾤ᆅࡢ୰ᚰᆅ࡟ࡣࠊ
す⬥ᕷすᯘᑎᒣ㫽⋇ಖㆤ༊ࡀ఩⨨ࡋ࡚࠾ࡾࠊ
ࢽ࣍ࣥࢪ࢝ࡸ࢖ࣀࢩࢩࡀ⏕ᜥࡋ࡚࠸ࡿࠋ⠛ᒣᕷࡣࠊỈ✄ࡸ኱㇋⏕⏘ᆅ࡜ࡋ࡚ࠕ୹Ἴ⠛ᒣࠖࣈࣛࣥࢻࢆᣢࡘᆅᇦ࡛࠶ࡿࡓࡵࠊస௜
㠃✚࠶ࡓࡾࡢ༢౯ࡀ㧗࠸ᆅᇦ࡜ࡋ࡚▱ࡽࢀ࡚࠸ࡿࠋࡲࡓࠊ
ࠕࡰࡓࢇ㘠ࠖ࡜ࡋ࡚୺࡟⊁   ⊟ᮇࡢ࢖ࣀࢩࢩ⫗ࡀࣈࣛࣥࢻ໬ࡋ࡚࠸ࡿࡓ
ࡵ㸦㉥ᫍࠊ2001㸧ࠊ࢖ࣀࢩࢩࡢ᭷ᐖᤕ⋓ࡢせㄳࡔࡅ࡛࡞ࡃࠊ⊁⊟ᮇ࡟࠾ࡅࡿ⊁⊟ᅽࡀ    ᅗ㸯 ⾜ືㄪᰝࢆ⾜ࡗࡓㄪᰝᆅᇦ
࡜ࡶ࡟㧗࠸ࠋ       ᅗ  ㄪᰝᆅࡢᴫせ㸦㸯㸧ㄪᰝಶయ 1ࠊ1 ࡢ⏕ᜥᆅ࿘㎶   㸦㸰㸧ㄪᰝಶయ 1 ࡢ⏕ᜥᆅ࿘㎶ රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-4㸫㸬᪉ἲ
⾜ືㄪᰝGPS ࢸ࣓ࣞࢺ࣮ࣜἲࡣࠊணࡵࣉࣟࢢ࣒ࣛタᐃࡋࡓ GPS఩㛫㝸࡛఩⨨᝟ሗࢆᚓࡿࡇ

࡜ࡀ࡛ࡁࡿࡓࡵࠊ㠀ᖖ࡟㧗㢖ᗘࠊ࠿ࡘ㧗⢭ᗘ࡞ື≀ࡢ఩⨨᝟ሗࡀᚓࡽࢀࡿࠋࡇࡢࡓࡵࠊ
㏆ᖺ࡛ࡣࠊࢽ࣍ࣥࢪ࢝ࡸࢶ࢟ࣀ࣡ࢢ࣐࡞࡝ࡢ኱ᆺ⋇ࡢ⾜ື㏣㊧࡟ GPS 㤳㍯ࡀ㢖⦾࡟౑ࢃࢀࡿࡼ࠺࡟࡞ࡗ࡚࠸ࡿࠋ࢖ࣀࢩࢩࡢሙྜࡣయᆺⓗ࡟㤳㍯⿦╔ࡀ㞴ࡋࡃࠊࡲࡓἾᾎࡧࡸ
⫼ࡇࡍࡾ࡞࡝ࡢ⾜ື≉ᛶ࡟ࡼࡗ࡚ࡶ㤳㍯ࡢᦆയࡀⴭࡋ࠸࡜⪃࠼ࡽࢀࠊGPS 㤳㍯࡟ࡼࡿ
ㄪᰝࡀ࡯࡜ࢇ࡝⾜ࢃࢀ࡚ࡇ࡞࠿ࡗࡓࠋࡋ࠿ࡋࠊኪ㛫ࡢ⾜ືࡸ㎰⪔ᆅ࡬ࡢฟἐ᫬㛫ᖏࠊ㎰⪔ᆅ࡬ࡢ⛣ື⤒㊰ࠊ⊁⊟άື࡟ࡼࡿ⾜ືࣃࢱ࣮ࣥࡢኚ໬࡞࡝ࡀヲ⣽࡟グ㘓ࡉࢀࡿ࡜ᮇᚅ
࡛ࡁࡿࡓࡵࠊᮏㄪᰝ࡛ࡣࠊGPS ࢸ࣓ࣞࢺ࣮ࣜἲࢆ᥇⏝ࡋࡓࠋ࢖ࣀࢩࢩࡢ⏕యᤕ⋓ࡣࠊ2005 ᖺ 4 ᭶࠿ࡽ 2006 ᖺ 10 ᭶࡟࠿ࡅ࡚ࠊᏛ⾡◊✲ᤕ⋓࡟ࡼࡾ⟽⨜ࢆ⏝࠸࡚⾜ࡗࡓ㸦ᅗ㸱㸧ࠋ⟽⨜ࡢタ⨨ሙᡤࡣࠊ㎰ᴗ⿕ᐖࡀⓎ⏕ࡋ࡚࠸ࡿᆅᇦࢆᑐ
㇟࡜ࡋࡓࠋᤕ⋓ࡉࢀࡓಶయࡣࠊ୙ື໬ᚋࠊGPS ෆ⮚㤳㍯㸦GPS3300 ࠾ࡼࡧ GPS4400ࠊ
Lotek ♫㸧ࢆ⿦╔ࡋࠊᤕ⋓ሙᡤ࡟࡚ᨺ⋇ࡋࡓࠋ GPS఩ࢫࢣࢪ࣮ࣗࣝࡣࠊ30 ศ㛫㝸ࡶࡋࡃࡣࠊ1 ᫬㛫㛫㝸ࢆタᐃࡋࠊ5 㐌࠿ࡽ 30 㐌㛫࡟ࢃࡓࡾࠊGPS ࡟ࡼࡿ࢖ࣀࢩࢩࡢ఩⨨᝟ሗࡢ⵳✚ࢆヨࡳࡓࠋࡲࡓࠊ⿦╔ᮇ㛫୰࡟࠾࠾ࡼࡑࡢಶయࡢ఩⨨ࢆᢕᥱ࡛ࡁࡓ୍㒊ࡢಶయ࡟ᑐࡋ࡚ࠊ᭷ᐖᤕ⋓ࡸ⊁⊟άືᮇ㛫୰࡟⊁⊟≟࡟ᑐࡍࡿ⾜ືࡢ཯ᛂࢆぢࡓࠋࡑࡢᚋ
GPS 㤳㍯ࢆᅇ཰ࡋ࡚ᚓࡽࢀࡓ఩⨨᝟ሗࢆ GIS ゎᯒࢯࣇࢺ㸦ArcGIS10.2ࠊ ESRI ♫㸧࡟ࡼࡾゎᯒࢆ⾜ࡗࡓࠋ࢖ࣀࢩࢩࡢ⎔ቃ฼⏝ゎᯒ࡟ࡣ➨㸱ࠊ㸲ࠊ㸳ᅇ⮬↛⎔ቃಖ඲ᇶ♏ㄪᰝ㸦⎔ቃ┬ http://wwwࠋbiodic.go.jp/J-IBIS.html㸧ࡢ᳜⏕ᅗࢆ⏝࠸ࡓࠋ㸱㹂ࢹ࣮ࢱ࡟ࡘ࠸࡚ࡣࠊࢹ࢕ࣇ࢓ࣞࣥࢩࣕࣝ⿵ṇࢆ⾜ࡗࡓࠋᅗ  Ꮫ⾡◊✲ᤕ⋓࡟ࡼࡿ࢖ࣀࢩࢩࡢ⏕యᤕ⋓࡜ *36 㤳㍯⿦╔ಶయ⿕ᐖᑐ⟇⌧≧ㄪᰝ
⾜ື㏣㊧ㄪᰝ⤊஢ᚋࠊ࢖ࣀࢩࢩࡀ㎰ᆅ࡟ฟἐࡋ࡚࠸ࡓ㞟ⴠࢆ୰ᚰ࡟ࠊ୺࡟㜵ㆤᰙࡢタ⨨≧ἣࡸ⟶⌮≧ἣࠊ࠾ࡼࡧఇ⪔ᆅࡢ㓄⨨➼࡟ࡘ࠸࡚⌧ᆅㄪᰝࢆ⾜࠸ࠊGIS ୖ࡟సᅗࡋࡓࠋ
ࡲࡓࠊ⪺ࡁྲྀࡾㄪᰝࢆ⾜࠸ࠊ⾜ືㄪᰝᮇ㛫୰ࡢ㜵㝖≧ἣ࡟ࡘ࠸࡚ࡶ☜ㄆࢆ⾜ࡗࡓࠋ රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-4

ࡉࡽ࡟ࠊ⠛ᒣᕷ࡟ࡘ࠸࡚ࡣࠊ⾜ᨻⓗ࡟⿵ຓࢆ⾜ࡗࡓ㞟ⴠ༢఩ࡢ኱つᶍᰙࡢタ⨨≧ἣ㸦௨ୗࠊ㞟ⴠ㜵ㆤᰙ㸧࡟ࡘ࠸࡚↷఍ࡋࠊGIS ୖ࡟సᅗࡋࡓࠋ

㸬⤖ᯝ*36఩⋡GPS 㤳㍯ࢆ⿦╔ࡋࡓಶయࡢ୍ぴࢆ⾲ 1 ࡟♧ࡋࡓࠋす⬥ᕷ࡛㸴㢌ࠊරᗜ┴⠛ᒣᕷ㸳㢌ࠊྜィ㸯㸯㢌ࡢ࢖ࣀࢩࢩ࡟⿦╔ࡋࡓ㸦⾲ 1㸧ࠋࡇࡢ࠺ࡕ N4ࠊS5 ࡟ࡘ࠸࡚ࡣࠊᨺ⋇┤ᚋ࡟Ⓨಙᶵࡢ୙ලྜࡀ⏕ࡌࠊ ఩ࢹ࣮ࢱࡀྲྀᚓࡉࢀ࡞࠿ࡗࡓࠋࡋࡓࡀࡗ࡚ࠊࡇࡢ㸰㢌࡟ࡘ࠸࡚ࡣゎᯒ࠿ࡽ㝖እࡋࠊ9 㢌࡟ࡘ࠸࡚ゎᯒࢆ⾜ࡗࡓࠋGPS఩⋡ࢆࠊ⾲ 2 ࡟♧ࡍࠋ㸰㹂ࠊ㸱㹂ࢆ࠶ࢃࡏࡓࢹ࣮ࢱࡢ

఩ᡂຌ⋡ࡣࠊ11㸣㹼62㸣఩⋡ࡀపୗࡋࠊ2㸣㹼30㸣࡜࡞ࡗࡓࠋࡋ࠿ࡋࠊ

࡛࠶ࡗࡓࠋ㸱㹂ࢹ࣮ࢱࡢࡳ࡛ࡣࠊ኱ᖜ࡟

఩ᩘ࡜ࡋ࡚ࡳࡿ࡜ 61㹼833 ᆅⅬࡀᚓࡽࢀࠊᚑ᮶ࡢࣛࢪ࢜ࢸ࣓ࣞࢺ࣮ࣜἲ࡟ẚ㍑ࡍࡿ
࡜኱㔞ࡢ᝟ሗࡀᚓࡽࢀࡓࠋ⾜ືゎᯒ࡟ࡣࠊ఩⨨⢭ᗘࡀ㧗࠸㸱㹂ࢹ࣮ࢱࡢࡳ࡛༑ศ࡞᝟ሗ
㔞࡜ุ᩿ࡋࠊ௨ᚋࡢゎᯒ࡟ࡣ㸱㹂ࢹ࣮ࢱࡢࡳࢆᢅࡗࡓࠋ
⾲㸯㸬ᤕ⋓ಶయࡢᇶᮏ᝟ሗ࡜ GPS సື᪥ᩘ䉰 ಶయ㻵㻰ᤕ⋓ሙᡤᛶู ᖺ㱋 య㔜ᤕ⋓᪥㻳㻼㻿సື᭱⤊᪥ 㻳㻼㻿సື᪥ᩘ

㻝

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す⬥ᕷ⯪⏫

䗠

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㻞㻜㻜㻡ᖺ㻠᭶㻝㻤᪥ 㻞㻜㻜㻡ᖺ㻡᭶㻞㻟᪥

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す⬥ᕷ᪥㔝⏫ 䗢㻝㻟㻞㻚㻡㻞㻜㻜㻡ᖺ㻥᭶㻝㻣᪥ 㻞㻜㻜㻢ᖺ㻝᭶㻝㻟᪥

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㻞㻜㻜㻢ᖺ㻣᭶㻣᪥ 㻞㻜㻜㻢ᖺ㻣᭶㻞㻥᪥

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㻞㻜㻜㻢ᖺ㻝㻜᭶㻝㻟᪥ 㻞㻜㻜㻢ᖺ㻝㻜᭶㻞㻤᪥

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㻝㻟㻚㻤 㻞㻜㻜㻢ᖺ㻝㻜᭶㻝㻟᪥ 㻞㻜㻜㻢ᖺ㻝᭶㻝㻡᪥

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⠛ᒣᕷᆶỈ

䗠

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㻞㻠

㻞㻜㻜㻢ᖺ㻟᭶㻝㻟᪥㻞㻜㻜㻢ᖺ㻢᭶㻢᪥

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⠛ᒣᕷᏳ⏣

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㻥

㻿㻙㻟

⠛ᒣᕷ௒⚟

䗢

㻝

㻡㻣

㻞㻜㻜㻢ᖺ㻣᭶㻠᪥ 㻞㻜㻜㻢ᖺ㻝㻞᭶㻝㻜᪥

㻝㻡㻥

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㻿㻙㻠

⠛ᒣᕷ௒⚟

䗢

㻞㻗

㻢㻣

㻞㻜㻜㻢ᖺ㻤᭶㻝㻢᪥ 㻞㻜㻜㻢ᖺ㻝㻜᭶㻞᪥

㻠㻣

㻝㻝

㻿㻙㻡

⠛ᒣᕷ᪂Ⲯ

䗠

㻞㻗

㻡㻤

㻞㻜㻜㻢ᖺ㻥᭶㻣᪥ 㻞㻜㻜㻢ᖺ㻥᭶㻝㻣᪥㻥㻜⾜ືᅪࡢ≉ᚩ
⿕ᐖᆅ࿘㎶࡛ᤕ⋓ࡉࢀࡓ࢖ࣀࢩࢩࡣࠊ㎰ᆅࡀ㏆᥋ࡋ࡚࠸ࡿ᳃ᯘࢆ୰ᚰ࡜ࡋࡓ⾜ືᅪࢆᣢࡕ㸦ᅗ 4㸧ࠊ᭱እ㒌ἲ࡟ࡼࡾồࡵࡓ⾜ື⠊ᅖࡣ 3.39±2.89km2㸦ᅗ 4ࠊ⾲ 3㸧࡛࠶ࡗࡓࠋ
ᑐ㇟࡜࡞ࡗࡓ࢖ࣀࢩࢩࡢ⾜ືᇦࡣࠊධࡾ⤌ࢇࡔ㇂ᆅᙧ࡟㎰⪔ᆅࡀ㞄᥋ࡋ࡚࠸ࡿ᳃ᯘෆࡀ රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-4

ከ࠿ࡗࡓࠋࡲࡓࠊ⾜ືᅪ௜㏆ࡢ᳃ᯘ᳜⏕ࡣࢥࢼࣛ஧ḟᯘࢆ୰ᚰ࡜ࡋࡓᗈⴥᶞᯘࡀከࡃࠊ
ࣃࢵࢳ≧࡟ࢫࢠ࣭ࣄࣀ࢟ࡢ᳜ᯘᆅࡀ㓄⨨ࡉࢀ࡚࠸ࡿ⎔ቃ࡛࠶ࡗࡓ㸦ᅗ 5ࠊ6㸧ࠋ࢖ࣀࢩࢩࡢ⾜ືᅪෆࡢ฼⏝᳜⏕ࡣࠊᗈⴥᶞᯘࡀከࡃ᳜ᯘᆅࡢ๭ྜࡣప࠿ࡗࡓ㸦ᅗ 7㸧ࠋࡲࡓࠊỈ⏣ࡸᕷ⾤ᆅ࡞࡝ࡢேⅭⓗ⎔ቃࡢ฼⏝㢖ᗘࡢ㧗࠸ಶయࡶㄆࡵࡽࢀࡓࠋ
௒ᅇࡢㄪᰝ࡛ࡣࠊྠᡤⓗ࡟」ᩘಶయࡀ⏕ᜥࡋ࡚࠾ࡾࠊ⾜ືᅪࡣ୍㒊㔜࡞ࡗ࡚࠸ࡓ㸦N
㸯ʐ࡜ N3ʐࠊN2ʑ࡜ N5ʑ࡜ N6ʑࠊS3ʑ࡜ S4ʑࡢ 3 ᆅᇦ㸧㸦ᅗ 4㸧ࠋᡂ⋇࢜ࢫࡢ⾜ືᅪࡀ㔜࡞ࡿ஦౛ࡀㄆࡵࡽࢀࡓࡇ࡜࠿ࡽࠊ࢖ࣀࢩࢩࡣ㞤㞝࡟㛵ಀ࡞ࡃࠊ」ᩘಶయࡀྠᡤⓗ࡟⏕ᜥࡍࡿࡇ࡜ࡀ᫂ࡽ࠿࡜࡞ࡗࡓࠋ
㎰ᆅࡢ฼⏝ࣃࢱ࣮ࣥ
㎰ᆅ࡬ࡢฟἐ᫬㛫ᖏࡣࠊኪ㛫 18 ᫬௨㝆ࠊ᪩ᮅ 6 ᫬ࡲ࡛࡟㞟୰ࡋ࡚࠸ࡓ㸦ᅗ 8㸧ࡀࠊಶయ࡟ࡼࡗ࡚ࡣࠊ᫨㛫࡟㎰ᆅ࡟ฟἐࡍࡿಶయࡶ࠸ࡓࠋࡲࡓࠊఫᏯ⾤࡟ฟἐࡋ࡚࠸ࡓ N6 ࡣࠊ௚ࡢಶయ࡟ẚ࡭᫨㛫ࡢάື᫬㛫ࡀከ࠿ࡗࡓࠋ⾲ 㸬ྛಶయࡢ *36

ಶయ㻵㻰

඲

఩ᩘ࡜

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఩⋡

఩ᩘ

⾲ 㸬ྛಶయࡢ⾜ືᅪ

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ಶయ㻵㻰 ⾜ືᅪ䠄㼗㼙 㻞 䠅

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ᖹᆒ
ᶆ‽೫ᕪ

㻟㻚㻟㻥

㎰ᆅ࡬ࡢฟἐࣃࢱ࣮ࣥࡣࠊS3ࠊS4 ࡢࡼ࠺࡟▷ᮇ㛫ࡢ࠺ࡕ࡟࠸ࡃࡘ࠿ࡢ㎰ᆅ࡟ሙᡤࢆኚ࠼࡞ࡀࡽฟἐࡍࡿࣃ

㻞㻚㻥ࢱ࣮ࣥ㸦ฟἐࣃࢱ࣮ࣥ㸯㸧㸦ᅗ 6㸧࡜ N2ࠊN6ࠊS1ࠊS2ࡢࡼ࠺࡟≉ᐃࡢሙᡤ࡟㞟୰ࡋ࡚ฟἐࡍࡿ㸦ฟἐࣃࢱ࣮ࣥ㸰㸧㸦ᅗ 5ࠊ6㸧ࡢ㸰ࡘࡢࢱ࢖ࣉ
ࡀㄆࡵࡽࢀࡓࠋ
N2ࠊN6 ࡣࡑࢀࡒࢀࠊ≉ᐃࡢሙᡤ࡟⧞ࡾ㏉ࡋฟἐࡋ࡚࠸ࡿࡀࠊ࡜ࡶ࡟ᕷ⾤ᆅ࡟ᅖࡲࢀ
ࡓ㫽⋇ಖㆤ༊࡟୺せ࡞⾜ືᅪࢆᣢࡕࠊฟἐሙᡤࡣࠊఇ⪔ᆅ࠾ࡼࡧࢦ࣑⟽➼ࡀ࠶ࡿఫᏯᆅ රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-4

࡛࠶ࡗࡓࠋN1ࠊN3ࠊS1ࠊS4 ࡟ࡘ࠸࡚ࡣࠊ≉ᐃࡢ㎰ᆅ࡟ࡢࡳฟἐࡋࠊࡑࢀ௨እࡢ㎰ᆅ࡟ࡣ౵ධࡋ࡞࠿ࡗࡓ㸦ᅗ 5ࠊ6㸧ࠋN5 ࡟ࡘ࠸࡚ࡣࠊᨺ⋇ 15 ᪥ᚋ࡟᭷ᐖᤕ⋓άື࡟ࡼࡗ࡚
ᤕ⋓ࡉࢀࡓࠋ㏣㊧ᮇ㛫୰ࡣ㎰ᆅࢆ฼⏝ࡍࡿࡇ࡜ࡣ࡞࠿ࡗࡓࠋ
ฟἐ≧ἣ࡜⿕ᐖ㜵Ṇᰙࡢタ⨨࡜ࡢ㛵ಀ
9 㢌ࡢ࢖ࣀࢩࢩࡢ⾜ືᅪ࿘㎶ࡢ㎰ᆅࢆ୰ᚰ࡟⿕ᐖ㜵Ṇᰙࡢタ⨨≧ἣࠊ㞟ⴠ࿘㎶ࡢ⎔ቃ࡟ࡘ࠸࡚⾲ 4 ࡟ࡲ࡜ࡵࡓࠋࡲࡓࠊฟἐሙᡤ࡟࠾ࡅࡿ㞟ⴠࡢᰙࡢタ⨨≧ἣ࡜࢖ࣀࢩࢩࡢά
ືⅬࢆ㔜ࡡྜࢃࡏࡓ㸦ᅗ 9ࠊ ᅗ 10㸧ࠋฟἐࣃࢱ࣮ࣥ㸯ࡢಶయࡀฟἐࡋࡓ㎰ᆅࡣࠊ㞟ⴠ㜵ㆤᰙࡀタ⨨ࡉࢀ࡚࠸࡞࠸ࠊ࠶ࡿ࠸ࡣ୍㒊࡟タ⨨ࡉࢀ࡚࠸ࡿ≧ἣ࡜࡞ࡗ࡚࠸ࡿ㞟ⴠ࡛࠶ࡗࡓࠋ≉࡟ࠊS2 ࡢάືᆅᇦ࿘㎶ࡢ㞟ⴠ࡛ࡣࠊࡲࡗࡓࡃ㞟ⴠ㜵ㆤᰙࡀタ⨨ࡉࢀ࡚࠸࡞࠿
ࡗࡓࠋ࢖ࣀࢩࢩࡢฟἐᆅⅬࢆ☜ㄆࡍࡿ࡜ಶே㜵ㆤᰙࡀタ⨨ࡉࢀ࡚࠸ࡿ㎰ᆅ࡛࠶ࡗࡓ㸦ᅗ
9ࠊ⾲ 4㸧ࠋ౵ධࢆ࠺ࡅࡓᰙࡢ✀㢮ࡣࢺࢱࣥࡸᮌᮦࠊࡢࡾ⥙࡞࡝ࢆ฼⏝ࡋ࡚࠸ࡿࡶࡢ࡛࠶
ࡗࡓࠋS3 ࡢάືᆅᇦ࿘㎶ࡣࠊᯘ⦕㒊࡟㞟ⴠ㜵ㆤᰙࡀタ⨨ࡉࢀ࡚࠸ࡿ࢚ࣜ࢔ࡀከࡃ࠶ࡗ
ࡓࡀࠊฟἐᆅ࡟㞟ⴠᰙࡣ࡞ࡃࠊಶே㜵ㆤᰙࡀᩘከࡃタ⨨ࡉࢀ࡚࠸ࡓ㸦ᅗ 9㸧ࠋ౵ධࢆཷ
ࡅࡓᰙࡣࡢࡾ⥙➼ࢆ౑ࡗࡓᰙࡀከࡃࡳࡽࢀࡓࠋ
ḟ࡟ࠊฟἐࣃࢱ࣮ࣥ㸰ࡢಶయࡀฟἐࡋ࡚࠸ࡿᆅᇦࡢ⿕ᐖ㜵㝖࡟ࡘ࠸࡚♧ࡍࠋS4 ࡣࠊฟἐࡋ࡚࠸࡞࠸㞟ⴠ࡟ࡣᯘ⦕㒊࡟㟁Ẽᰙࡀタ⨨ࡉࢀࠊ㇂ዟ࡟ࡣ㏉ࡋᰙࡶసࡽࢀࡿ࡜࡜ࡶ
࡟ࠊ₃㟁ࢆ㑊ࡅࡿࡓࡵࡢୗⲡࡢ⟶⌮➼ࡀ⾜ࡁᒆ࠸࡚࠸ࡓ㸦ᅗ 10㸧ࠋฟἐࡋ࡚࠸ࡓ㎰ᆅ࡛ࡣ㞟ⴠ㜵ㆤᰙࡣタ⨨ࡉࢀ࡚࠾ࡽࡎࠊ⬤ᙅ࡞ಶே㜵ㆤᰙࡢࡳ࡛࠶ࡗࡓࠋࡇࡢ㎰ᆅ࡟ࡣఇ⪔ᆅࡶከ࠿ࡗࡓࠋ
S1 ࡟ࡘ࠸࡚ࡣࠊ⾜ືᅪࡀ S4 ࡜㔜࡞ࡿ㒊ศࡀከ࠿ࡗࡓࡀࠊS4 ࡜ࡣ␗࡞ࡿ㎰ᆅ࡟ฟἐࡋ࡚࠸ࡓࠋS1 ࡢฟἐ㎰ᆅ࡟ࡘ࠸࡚ࡶࠊಶே㜵ㆤᰙࡢࡳࡀタ⨨ࡉࢀ࡚࠸ࡿ㎰ᆅ࡛࠶ࡗࡓ
㸦⾲ 4㸧
ࠋS4 ࡜ྠᵝ࡟㞟ⴠ㜵ㆤᰙࡀタ⨨ࡉࢀ࡚࠸ࡿ㎰ᆅ࡟ࡣ㏆࡙࠸࡚࠸࡞࠸ࠋ
 N1 ࡜ N3 ࡣ⾜ືᅪࡀ㔜࡞ࡾࠊ࡜ࡶ࡟㞄᥋ࡍࡿ㸰ࡘࡢ㎰ᆅ aࠊb ࡢ࠺ࡕࠊ㎰ᆅ a ࡢࡳࢆ฼⏝ࡋ࡚࠸ࡓ㸦ᅗ 11㸧ࠋ㎰ᆅ㹠ࡣࠊ㏻ᖺ࡛㟁Ẽᰙࢆタ⨨ࡋࠊᯘ⦕㒊ࡢぢ㏻ࡋࡀࡼࡃฟ
ἐᢚไᑐ⟇ࡀ⾜ࢃࢀ࡚࠸ࡓࠋ୍᪉ࠊ㎰ᆅ a ࡛ࡣࠊ⪔సᮇ㛫ࡢࡳ࡟㟁Ẽᰙࡀタ⨨ࡉࢀࠊᯘ⦕㒊࡟ࡣ㧗ⱼⲡᮏࡸἙ␁ᯘࡀⓎ㐩ࡋ࡚࠾ࡾࠊぢ㏻ࡋࡢᝏ࠸⎔ቃ࡛࠶ࡗࡓࠋN1ࠊN3 ࡜ࡶ࡟㏣㊧ᮇ㛫ࡣ㎰㛩ᮇ࡟࠶ࡓࡾࠊ㎰ᆅ a ࡢ㟁Ẽᰙタ⨨ࡉࢀ࡚࠸࡞࠿ࡗࡓࠋ
N2ࠊN6 ࡣ୺࡞⾜ືᅪ࡟ᙜࡓࡿ᳃ᯘࡣ㫽⋇ಖㆤ༊࡟ᣦᐃࡉࢀ࡚࠸ࡿᆅᇦ࡛࠶ࡗࡓ㸦ᅗ
5㸧ࠋN2 ࡟ࡘ࠸࡚ࡣࠊᕷ⾤ᆅ㏆ࡃࡢఇ⪔ᆅࢆ୺࡟ኪ㛫฼⏝ࡋ࡚࠸ࡓࡀࠊᰙ࡜ࡢ⿕ᐖᑐ⟇ࡣ⾜ࢃࢀ࡚࠸࡞࠿ࡗࡓࠋN6 ࡟ࡘ࠸࡚ࡣࠊᕷ⾤ᆅࢆ฼⏝ࡋ࡚࠸ࡓࡀࠊ᫨㛫౑࠺ሙᡤ࡜ኪ
㛫౑࠺ሙᡤࡀ࠶ࡗࡓ㸦ᅗ 5㸧ࠋ௨ୖࠊ࢖ࣀࢩࢩࡣ㞟ⴠ㜵ㆤᰙࡀタ⨨ࡉࢀ࡚࠸࡞࠸ࠊಶே㜵ㆤᰙᑐᛂࡢࡳࡢ⬤ᙅ࡞㜵㝖ࡢ㎰ᆅ࡟౵ධࡋ࡚࠸ࡿࡇ࡜ࡀ᫂ࡽ࠿࡜࡞ࡗࡓࠋࡲࡓࠊᑐ⟇ࡀ⾜ࢃࢀ࡞࠸㎰㛩ᮇ࡟࠾࠸࡚
ࡶ㎰ᆅࢆ฼⏝ࡋ࡚࠸ࡓ㸦ᅗ 11㸧
ࠋ

 

රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-4

㸦㸯㸧す⬥ᕷす⬥ᕷ࡟࠾ࡅࡿ
ㄪᰝಶయࡢ⾜ືᅪ1 ‫ڧ‬1 ‫ڧ‬1 ‫ڧ‬1 ‫ڧ‬1 ‫ڧ‬㸦㸧⠛ᒣᕷ

⠛ᒣᕷ࡟࠾ࡅࡿ
ㄪᰝಶయࡢ⾜ືᅪ6 ‫ڧ‬6 ‫ڧ‬6 ‫ڧ‬ᅗ  ㄪᰝಶయࡢ᭱እ㒌⾜ືᅪ㸦㸯㸧す⬥ᕷ࡟࠾࠸࡚ᤕ⋓ࡉࢀࡓ㸳ಶయཬࡧ㸦㸰㸧⠛ᒣᕷ࡟࠾࠸࡚ᤕ⋓ࡉࢀࡓ  ಶయ රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-41

䕔㻌 ᳜ᯘᆅ㻌 㻌1䕔㻌 ᗈⴥᶞᯘ㻌 㻌
䕔㻌 ㎰⪔ᆅ㻌 㻌䕔㻌 ᕷ⾤ᆅ㻌 㻌

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1

1

ᅗ රᗜ┴す⬥ᕷ࡟࠾ࡅࡿ *36 ⿦╔ಶయࡢ

఩ᆅⅬ

఩ᆅⅬࡣࠊ᫬㛫ࡈ࡜࡟♧ࡋࡓࠋ‫ە‬ኪ㛫㸦 ᫬ࠊ ᫬㸧ࠊ‫ە‬᫨㛫㸦 ᫬㸧ࠋᩳ⥺ࡣ㫽⋇ಖ
ㆤ༊ࠋ

 

රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-4

6

6





 NP

6

6

䕔㻌 ᳜ᯘᆅ㻌 㻌䕔㻌 ᗈⴥᶞᯘ㻌 㻌
䕔㻌 ㎰⪔ᆅ㻌 㻌ᅗ  ⠛ᒣᕷ࡟࠾ࡅࡿ *36 ⿦╔ಶయࡢ



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䕔㻌 ᕷ⾤ᆅ㻌 㻌

఩ᆅⅬ

఩ᆅⅬࡣࠊ᫬㛫ࡈ࡜࡟♧ࡋࡓࠋ‫ە‬ኪ㛫㸦 ᫬ࠊ ᫬㸧ࠊ‫ە‬᫨㛫㸦 ᫬㸧ࠋ⊁⊟άື➼࡟ࡼࡿ㏣࠸ᡶ࠸
N1 ࡣᤕ⋓ᨺ⋇┤ᚋ࡟ࡶ኱ࡁࡃ⛣ືࡋࡓࡀࠊᩘ᪥ᚋ࡟ᤕ⋓ᆅ࿘㎶ᅇᖐࡋࠊࡑࡢᚋࡶᤕ
⋓ᆅⅬ࿘㎶࡛⾜ືࡋࡓ㸦ᅗ 5㸧ࠋᤕ⋓࠿ࡽ⣙㸯ࣨ᭶ᚋ࡟ N1 ࡢ⏕ᜥࡍࡿ᳃ᯘ࡛᭷ᐖᤕ⋓άືࡀ⾜ࢃࢀࡓ㝿࡟ࡣࠊ⊟≟ࡀ N1 ࡟᥋㏆ࡋࡓᚋࠊ1km ࡯࡝⛣ືࡋࡓࠋࡋ࠿ࡋࠊࡑࡢ 4᫬㛫ᚋ࡟ࡣ⊁⊟๓ࡢ⁫ᅾᆅⅬ௜㏆࡟ᡠࡗ࡚࠸ࡓ㸦ᅗ 5㸧ࠋ රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-4

S4 ࡢሙྜࠊ⊁⊟⪅㺃⊟≟ࡀ㏆ࡃ࡟ࡁ࡚ࡶ⛣ືࡋ࡞࠿ࡗࡓࠋ⊁⊟⪅ࡀ❧ࡕཤࡗࡓࡢࡕ
2.5km 㞳ࢀࡓሙᡤ࡟⛣ືࡋࡓࡀࠊ⩣᪥࡟ࡣ⛣ື๓࡜ྠࡌ㎰⪔ᆅ࡟ฟἐࡋ࡚࠸ࡓࠋࡑࡢᚋ
ࡶ⊁⊟๓࡜ྠᵝ࡟ࠊ࡯ࡰẖ᪥㎰⪔ᆅ࡬ฟἐࡋ࡚࠸ࡓ(ᅗ 12)ࠋูࡢ᪥࡟ࠊྠᵝࡢ㏣࠸ᡶ࠸άືࢆᐇ᪋ࡋࡓࡀࠊS4 ࡟ࡘ࠸࡚ࡣࠊྠᵝࡢ⾜ືࢆ࡜ࡾࠊ⾜ືᇦࢆኚ࠼ࡿࡇ࡜ࡣ࡞࠿ࡗ
ࡓࠋࡇࢀࡽࡢ౛࠿ࡽࠊ࢖ࣀࢩࢩࡣ⊁⊟ࡸ㏣࠸ᡶ࠸࡞࡝࡟ࡼࡗ୍࡚᫬ⓗ࡟኱ࡁࡃ⛣ືࡍࡿࡇ࡜ࡣ࠶ࡗ࡚ࡶࠊࡍࡄ࡟ࡶ࡜ࡢ⾜ືᇦ࡟ᅇᖐࡋࠊ㎰ᆅ࡬ฟἐࡍࡿ࡜࠸࠺⾜ື࡟ኚ໬ࢆ୚࠼
ࡿࡇ࡜ࡀ㞴ࡋ࠸ࡇ࡜ࡀ᫂ࡽ࠿࡜࡞ࡗࡓࠋ

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ᅗ  ⾜ືᅪෆࡢ᳜⏕๭ྜ࡜⾜ືᆅⅬࡢ᳜⏕๭ྜ Წࢢࣛࣇࡣ⾜ືᅪෆࡢ᳜⏕๭ྜࠊᢡࢀ⥺ࢢࣛࣇࡣ ఩ᆅⅬࡢ᳜⏕๭ྜࢆᆅⅬᩘ࡛♧ࡋࡓࠋ㸬⪃ᐹ⾜ື≉ᛶ㎰ᆅ࡟㞄᥋ࡋ࡚⏕ᜥࡍࡿ࢖ࣀࢩࢩࡢ⾜ືᅪࡣࠊ኱ᆺ⋇࡜ࡋ࡚ࡣࠊẚ㍑ⓗ⊃࠸ࡶࡢ࡛࠶ රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-4

ࡗࡓࠋ㎰ᆅ࡟㞄᥋ࡋ࡚⏕ᜥࡍࡿ࢖ࣀࢩࢩ࡛ࡣࠊྠᵝ࡟⾜ືᅪࡀ⊃࠸ࡇ࡜ࡀᓥ᰿┴㸦ᑠᑎ2001)࡛ࡶሗ࿌ࡉࢀ࡚࠸ࡿࠋ≉࡟⿕ᐖࢆ㜵ࡄࡓࡵ࡟⤥㣵ࢆ⾜࠺࡜⾜ືᅪࡣࡉࡽ࡟⦰ᑠࡍ
ࡿࡇ࡜࡞࡝࠿ࡽ㸦ᑠᑎ࡯࠿ 2006㸧
ࠊ࢖ࣀࢩࢩࡣ㈨※ࡀ㇏ᐩ࡟࠶ࢀࡤ⊃࠸⠊ᅖ࡛⾜ືࡍࡿ≉ᛶࡀ࠶ࡿ࡜᥎ᐃࡉࢀࡿࠋᮏㄪᰝ࡛ࡣ㏣㊧ᮇ㛫୰࡟ᡂ⋇㞝ࢆྵࡵ㛗⛣ືࡍࡿࡇ࡜ࡣ࡞࠿
ࡗࡓࠋ᪥ᖖⓗ࡟ࡶ᭷ᐖᤕ⋓άືࡀ⾜ࢃࢀ࡚࠸ࡿᆅᇦ࡛ࡶ࠶ࡿࡇ࡜࠿ࡽࠊ⟬ฟࡉࢀࡓ⾜ືᅪࢧ࢖ࢬࡣࠊࡇࢀࡽࡢ㏣࠸ᡶ࠸άື࡟ࡼࡗ࡚࢖ࣀࢩࢩࡢ⾜ືࡀ㢖⦾࡟ேⅭⓗᨩ஘ࡢᙳᙳ 

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ᅗ  ࢖ࣀࢩࢩࡢ㎰ᆅ࣭ᕷ⾤ᆅ࡬ࡢฟἐ᫬้
㡪ࢆཷࡅࠊ୍᫬ⓗ࡟⛣ືࡍࡿ⾜ືࢆྵࡵࡓࡶࡢ࡛࠶ࡿ࡜⪃࠼ࡽࢀࡿࠋࡑࡢࡓࡵࠊ㏣࠸ᡶ࠸ࡸᤕ⋓άືࡀ࡞࠿ࡗࡓሙྜࠊࡉࡽ࡟⾜ືᅪࢧ࢖ࢬࡣ⊃࠸ࡶࡢ࡜⪃࠼ࡽࢀࡿࠋ௨ୖ࠿ࡽࠊ
࢖ࣀࢩࢩࡣᐃ╔ᛶࡀ㠀ᖖ࡟ᙉࡃࠊ⾜ືᅪࡣ㏣࠸ᡶ࠸➼ࡢ᧠஘ࡢᙳ㡪ࢆཷࡅ࡟ࡃ࠸࡜⪃࠼ࡽࢀࡿࠋ⾜ືᅪෆࡢ᳜⏕ࡣࢥࢼࣛࢆ୺య࡜ࡍࡿᗈⴥᶞᯘࡀከࡃࠊࡉࡽ࡟⾜ືᆅⅬࡣᗈⴥᶞࡀ㑅ᢥⓗ࡟฼⏝ࡉࢀ࡚࠸ࡓࠋࡑࡋ࡚࢖ࣀࢩࢩࡣࠊ᳜ᯘᆅࢆᚷ㑊ࡍࡿഴྥ࡟࠶ࡗࡓࠋࡇ
ࡢ⤖ᯝࡣࡇࢀࡲ࡛ࡢ࢖ࣀࢩࢩࡢሗ࿌ࢆᨭᣢࡍࡿࡶࡢ࡛࠶ࡿ㸦ᑠᑎ࡯࠿ 2001㸹ᮏ⏣࡯࠿
2008㸧ࠋࡲࡓࠊ」ᩘಶయࡀྠᡤⓗ࡟⏕ᜥࡋ࡚࠸ࡿሙᡤࡶ☜ㄆࡉࢀࡓࠋ㎰ᆅ࡟᥋ࡋࡓᗈⴥ
ᶞᯘࡀከ࠸⎔ቃࡣࠊ㞧㣗࡛࠶ࡿ࢖ࣀࢩࢩ࡟࡜ࡗ࡚ࡢ㈨※㔞ࡢከᵝᛶࡀಖࡓࢀ࡚࠸ࡿ࡜⪃࠼ࡽࢀࠊ」ᩘಶయࡀ⏕ᜥ࡛ࡁࡿⰋዲ࡞⏕ᜥ⎔ቃ࡟࠶ࡿࡇ࡜ࡀ࠺࠿ࡀ࠼ࡿࠋ
㎰ᆅ࡬ࡢฟἐ࡜⿕ᐖᑐ⟇ ㎰ᆅ࡟ฟἐࡋ࡚࠸ࡓಶయࡣኪ㛫ࠊ≉࡟ 21 ᫬࠿ࡽ 5 ᫬ࡲ࡛࡟㞟୰ࡋ࡚࠸ࡓࠋࡇࢀࡣࠊ
ே㛫άືࢆ㑊ࡅኪ㛫࡟㎰ᆅࢆ฼⏝ࡋ࡚࠸ࡿࡓࡵ࡜⪃࠼ࡽࢀࡿࠋࡋ࠿ࡋࠊᕷ⾤ᆅࢆ฼⏝ࡋ࡚࠸ࡓಶయࡣࠊ᫨㛫࡟ࡶ㧗࠸㢖ᗘ࡛ฟἐࡋ࡚࠸ࡿࡇ࡜ࡀ᫂ࡽ࠿࡜࡞ࡗࡓࠋ࣮ࣚࣟࢵࣃ࢖
ࣀࢩࢩ࡛ࡣࠊே㛫ࡢᙳ㡪ࡀᑡ࡞࠸࡜᪥୰࡟άືࡋࠊኪ㛫࡟ࡣάືࡀⴠࡕࡿ㸦Kurz &Marchinton 1972㸧ࡇ࡜ࡀሗ࿌ࡉࢀ࡚࠸ࡿࡀࠊ⊁⊟ᅽࡀ㧗࠸ᆅ఩ᆅᇦ࡛ࡣࠊኪ㛫ࡢάື රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-4

ࡀ㧗ࡲࡿ㸦Hanson & Karstad 1959; Boitani et al 1994㸧ࡇ࡜ࡀሗ࿌ࡉࢀ࡚࠾ࡾࠊᮏ◊✲ࡢෆᐜࡶྠᵝࡢ⤖ᯝ࡜࡞ࡗࡓࠋ㣵࡙ࡅ࡟ࡼࡿฟἐࡀၥ㢟どࡉࢀ࡚࠸ࡿභ⏥ᒣ⣔࡟࠾࠸࡚ࡶ᪥୰ࡢάືࡀከࡃሗ࿌ࡉࢀ࡚࠸ࡿ㸦㎷࣭ᶓᒣ 2014㸧ࠋヱᙜࡢಶయࡣ୺࡞⾜ືᇦࢆ㫽⋇ಖㆤ༊ෆࡢ᳃ᯘ࡟ᣢࡘࡓࡵࠊ≉࡟㏣࠸ᡶࢃࢀࡿࡇ࡜࡞ࡃேࡢ⏕άᅪࢆᐜ᫆࡟฼⏝࡛ࡁࡿࡼ࠺࡟࡞ࡾࠊ⾜ືࡀ࢚ࢫ࣮࢝ࣞࢺࡋ࡚࠸ࡿྍ⬟ᛶࡀ࠶ࡿࠋ㒔ᕷࡸᕷ⾤ᆅ࡟㞄᥋ࡋࠊ࢖ࣀࢩࢩࡀ⏕ᜥࡍࡿ㫽⋇ಖㆤ༊࿘㎶࡛ࡣ㎰ᆅࡔࡅ࡛࡞ࡃࠊᕷ⾤ᆅ࡟࠾ࡅࡿฟἐ㜵Ṇᑐ⟇࡞࡝⿕ᐖ⟶⌮యไࡀᚲせ࡛࠶ࡿࡇ࡜ࢆ♧၀ࡋ࡚࠸ࡿࠋ
௒ᅇࠊ㎰ᆅ࡬ࡢฟἐࣃࢱ࣮ࣥ࡜ࡋ࡚௒ᅇ㸰ࡘࡢࣃࢱ࣮ࣥࡀㄆࡵࡽࢀࡓࠋฟἐሙᡤࢆኚ
࠼ࡿࣃࢱ࣮ࣥࢆ♧ࡋࡓಶయࡀ⾜ືࡋ࡚࠸ࡓᆅᇦ࡛ࡣࠊ㞟ⴠ㜵ㆤᰙࡀタ⨨ࡉࢀ࡚࠸࡞࠸ࠊ
࠶ࡿ࠸ࡣタ⨨ࡀ୍㒊࡟㝈ࡽࢀ࡚࠸ࡿᆅᇦ࡛࠶ࡾࠊ࢖ࣀࢩࢩࡣࠊᑐ⟇ࡀ⬤ᙅ࡞㎰ᆅࢆ㑅ᢥ
6

6

ᅗ  ฟἐࣃࢱ࣮ࣥ  ࡢ㎰ᆅ࡬ࡢฟἐ≧ἣ࡜㜵ㆤᰙࡢタ⨨≧ἣ

6

6

ᅗ  ฟἐࣃࢱ࣮ࣥ  ࡢ㎰ᆅ࡬ࡢฟἐ≧ἣ࡜㜵ㆤᰙࡢタ⨨≧ἣ ̿ 㞟ⴠ㜵ㆤᰙ   㸦タ⨨῭ࡳ㸧̿ 㞟ⴠ㜵ㆤᰙ   㸦タ⨨ணᐃ㸧̿ ಶே㜵ㆤᰙරᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-4

㎰ᆅ D

㎰ᆅ E
ᅗ  ㏣࠸ᡶ࠸άື࡟ࡼࡿ࢖ࣀࢩࢩࡢ⾜ືࡢ㌶㊧㸦ᶳⰍ▮༳㸧ᅗ  1ࠊ1 ಶయࡢ⾜ືᅪ࿘㎶ࡢ㎰ᆅ⾲  ࢖ࣀࢩࢩࡢฟἐ㎰ᆅ࡟࠾ࡅࡿ⿕ᐖ㜵㝖ᑐ⟇࡜㞟ⴠ⎔ቃಶయ㻵㻰⿕ᐖ䝺䝧䝹䠆

㻺㻝

㻺㻞

㻺㻟

㻺㻢

㻿㻝

㻿㻞

㻿㻟

㻿㻠

䠄⪺䛝ྲྀ䜚䠅

୰

኱

୰

୰

ᑡ

኱

䛺䛧

୰

㞟ⴠ༢఩䛾㜵㝖

୍㒊

䛺䛧

୍㒊

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୍㒊

䛺䛧

୍㒊

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ᰙ䛾✀㢮

㔠⥙ᰙ

䝛䝑䝖

㔠⥙ᰙ

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䝛䝑䝖

㟁Ẽᰙ䞉
䝛䝑䝖䞉䝖䝍
䞁䝛䝑䝖䝛䝑䝖䞉䝖䝍
䞁ᰙ䛾⟶⌮≧ἣ

⟶⌮

⟶⌮

⟶⌮

ᨺ⨨

⟶⌮

⟶⌮

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ᗫᲠ㎰స≀➼

ฎ⌮

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ᝏ

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ᝏ

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㜵㝖ᑐ⟇

㞟ⴠ⎔ቃ

ᶞᯘෆୗᒙ≧ែ䠄ぢ㏻䛧䠅㞟ⴠ࿘㎶ୗᒙ≧ែ
䠄ぢ㏻䛧䠅䠆⿕ᐖ䝺䝧䝹䠖⏒኱㻔⏕⏘㔞䛾㻣㻡㻑௨ୖ㻕䞉኱㻔㻣㻡䡚㻡㻜㻑㻕䞉୰㻔㻡㻜䡚㻞㻡㻑㻕䞉ᑡ㻔㻞㻡㻑ᮍ‶㻕䞉䛺䛧ࡋࠊ㞟୰ⓗ࡟⿕ᐖࢆ࠾ࡼࡰࡍࡇ࡜ࡀ᫂☜࡜࡞ࡗࡓࠋ≉ᐃࡢሙᡤ࡟ฟἐࡍࡿࣃࢱ࣮ࣥࢆ♧ࡋࡓಶయࡢ⾜ືᅪ࡛ࡣࠊࡑࢀ௨እࡢሙᡤ࡟㞟ⴠ㜵ㆤᰙࡀタ⨨ࡉࢀ࡚࠾ࡾࠊຠᯝⓗ࡟ฟἐࡀᢚไࡉࢀ࡚࠸ࡓࠋ㜵㝖యไࡢ⬤ᙅ࡞ሙᡤࡣࠊ≉ᐃࡢ㎰ᆅ࡟㝈ࡽࢀ࡚࠸ࡿࡓࡵ࡟㞟୰฼⏝ࡋ࡚࠸ࡿࡶࡢ࡜⪃࠼ࡽࢀࡓࠋ࢖ࣀࢩࢩࡣᚭᗏࡋࡓᑐ⟇ࡀ⾜ࢃࢀࡓሙྜࠊࡑࡢ㎰ᆅ௜㏆
࡟㏆࡙ࡃࡇ࡜ࡶ࡞ࡃࠊᑐ⟇ࡀ⬤ᙅ࡞㎰ᆅࢆ᥈⣴ࡋࠊ㞟୰ࡋ࡚฼⏝ࡍࡿ࡜࠸࠺⾜ືࢆᣢࡗ࡚࠸ࡿ࡜࠸࠼ࡿࠋ ௨ୖࠊ࢖ࣀࢩࢩࡢ㎰ᆅฟἐࡣࠊ⿕ᐖᑐ⟇ࡢ᭷↓ࡢᙳ㡪ࢆ኱ࡁࡃཷࡅࠊ≉࡟㞟ⴠ඲యࢆ࢝ࣂ࣮ࡍࡿ㞟ⴠᰙࡀຠᯝⓗ࡛࠶ࡿࡀࠊࡑࡢศ㞟ⴠᰙࡀ࡞ࡃᑐ⟇ࡢᙅ࠸ᆅᇦ࡬ࡢฟἐࢆຍ
㏿ࡉࡏ࡚ࡋࡲ࠺ྍ⬟ᛶࡀ♧၀ࡉࢀࡓࠋ࢖ࣀࢩࢩ࡟ࡼࡿ⿕ᐖ㜵Ṇᑐ⟇ࡣࠊ⿕ᐖᆅࡢ✵㛫ⓗ රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-4≉ᛶࢆ༑ศ࡟᳨ウࡋࠊ≉ᐃࡢ㎰ᆅ࡬࡟⿕ᐖࡀཬࡤ࡞࠸ࡼ࠺࡞㜵ㆤᰙࡢタ⨨ࢆィ⏬ࡍࡿ࡞࡝ࠊᗈᇦⓗ࡞どⅬ࡛ࡢ᳨ウࡶ㔜せ࡛࠶ࡿࠋࡲࡓ௒ᅇࡢㄪᰝᮇ㛫ࡣ㎰⦾ᮇࠊ㎰㛩ᮇ࡜ࡶ࡟㏣㊧ᮇ㛫࡜࡞ࡗ࡚࠸ࡓࡀࠊ࢖ࣀࢩࢩ࡟ࡼࡿ㎰ᆅ࡬ࡢฟἐࡣࠊ㎰㛩ᮇ࡟ࡶㄆࡵࡽࢀ࡚࠸ࡿࠋ
㎰㛩ᮇ࡟࠾࠸࡚ࠊ࢖ࣀࢩࢩ࡟ࡼࡿ㎰ᆅ฼⏝ࢆᨺ⨨ࡍࡿࡇ࡜ࡣࠊ࢖ࣀࢩࢩࡢຍᐖ⾜ືࢆ࢚ࢫ࣮࢝ࣞࢺࡉࡏࡿྍ⬟ᛶࡶ࠶ࡿࡓࡵࠊ௒ᚋࡢ⿕ᐖ⟶⌮࡛ࡣࠊ㎰㛩ᮇࡢ㎰ᆅ⟶⌮࡟ࡘ࠸࡚ࡶ᳨ウࡍࡿࡇ࡜ࡀ㔜せ࡛࠶ࡿࠋຍᐖಶయ࡟ᑐࡍࡿ㏣࠸ᡶ࠸ࡢຠᯝ ⊁⊟άື➼࡟ࡼࡿ㏣࠸ᡶ࠸ࡀࠊ࢖ࣀࢩࢩࡢ㎰⪔ᆅ࡬ࡢฟἐࡢᢚไ࡟ຠᯝࡀ࠶ࡿ࡜ண᝿
ࡋ࡚㏣࠸ᡶ࠸ࡢ཯ᛂࢆ㏣㊧ࡋࡓࡀࠊ㠀ᖖ࡟ᐃ╔ᛶࡀᙉࡃࠊᖖ࡟⾜ືᅪෆ࡟ᅇᖐࡍࡿ⾜ືࢆ࡜ࡗࡓࡓࡵࠊ኱ࡁࡃ⾜ືࢆኚ࠼ࡿࡇ࡜ࡣᅔ㞴࡛࠶ࡿࡇ࡜ࡀ♧ࡉࢀࡓࠋฟἐࢆᢚไࡍࡿ
ࡓࡵࡢ㏣࠸ᡶ࠸࡟‽ࡌࡓᑐ⟇ࡣࠊࡉࡽ࡟ᕤኵ࡜᳨ドసᴗࡀᚲせ࡛࠶ࡿࠋ౛࠼ࡤࠊ࣓ࣥࢸ
ࢼࣥࢫࡀ⾜ࡁᒆ࠸ࡓ㞟ⴠ༢఩ࡢ኱つᶍᰙ࡟ຠᯝࡀ࠶ࡗࡓࡇ࡜ࠊ㏣࠸ᡶ࠸άື୰࡟ࡣᩄឤ
࡞཯ᛂࢆ♧ࡋ༶ᗙ࡟㏨ࡆࡿሙྜࡸࠊேࡸ≟ࡀ㏻㐣ࡍࡿࡢࢆ₯ࢇ࡛ᚅࡘࡼ࠺࡞⾜ືࢆ࡜ࡿ࡞࡝ࠊ㠀ᖖ࡟⮫ᶵᛂኚ࡞཯ᛂࢆ♧ࡋ࡚࠸ࡓࠋࡑࡢࡓࡵ౛࠼ࡤࠊ᪥ᖖⓗ࡟ᯘ⦕㒊➼࡟࠾࠸࡚ࠊேࡸ≟࡞࡝ࡀάືࡍࡿࡇ࡜࡛ࠊ㎰ᆅ࡬ࡢ᥋㏆ࢆ࠶ࡁࡽࡵࡉࡏࡿ࡞࡝ฟἐࡀᢚไࡉࢀ
ࡿྍ⬟ᛶࡣ㧗࠸ࠋࡲࡓࠊ᭷ᐖᤕ⋓࡟ࡼࡿຠᯝࢆ㧗ࡵࡿࡓࡵ࡟ࡣࠊ࢖ࣀࢩࢩ࡟ࡼࡿ⿕ᐖⓎ⏕ᆅ࡟࠾࠸࡚ຍ
ᐖಶయࢆᤕ⋓ࡋ࡞ࡅࢀࡤࠊ⿕ᐖ㍍ῶࡢຠᯝࡀぢ㎸ࡵ࡞࠸ࡓࡵࠊᤕ⋓ᑐ㇟ಶయࢆᤕ⋓ࡍࡿయไࡀᚲせ࡜࡞ࡿࠋ
௨ୖࠊ⿕ᐖࢆཬࡰࡍ࢖ࣀࢩࢩࡢ⾜ືࢆ㏣㊧ࡋࡓ⤖ᯝࠊ㎰ᆅ࡟㞄᥋ࡋ࡚⏕ᜥࡍࡿ࢖ࣀࢩ
ࢩࡣࠊ⊃࠸⾜ືᅪ࡛ᐃ╔ⓗ࡛࠶ࡾࠊ⿕ᐖᑐ⟇ࡢ⬤ᙅ࡞ሙᡤࡸᑐ⟇ࡉࢀ࡚࠸࡞࠸ሙᡤࠊ࠶
ࡿ࠸ࡣ᫬ᮇ࡟㎰ᆅ࡟㞟୰ࡋ࡚ฟἐࡍࡿࡇ࡜ࡀ᫂ࡽ࠿࡜࡞ࡗࡓࠋࡲࡓࠊ⊁⊟άື࡟ࡼࡿ㏣࠸ᡶ࠸࡛ࡣࠊ࢖ࣀࢩࢩࡢ⾜ື࡟ኚ໬ࢆ୚࠼ࡿࡇ࡜ࡣᅔ㞴࡛࠶ࡗࡓࠋ⿕ᐖᆅ࿘㎶࡛ࡢᐃᮇⓗ࡞ಶయᩘㄪᩚ࡜㜵㝖ᰙ⟶⌮ࠊ㌟ࢆ₯ࡵࡿሙᡤࡸἩࡲࡾሙ࡜࡞ࡿ⸭࡞࡝ࡢ⎔ቃࡢ᤼㝖࡞࡝ࡀ㔜せ࡛࠶ࡿࡇ࡜ࡀ᫂ࡽ࠿࡜࡞ࡗࡓࠋㅰ㎡ ᤕ⋓ㄪᰝ࡟ᙜࡓࡾࠊරᗜ┴⊟཭఍す⬥ከྍᨭ㒊ࠊ⠛ᒣᨭ㒊ࡉࡘࡁ఍ࠊᰴᘧ఍♫㔝⏕ື≀ಖㆤ⟶⌮஦ົᡤ࠿ࡽከࡃࡢࡈ༠ຊࢆ࠸ࡓࡔࡁࡲࡋࡓࠋ㜵ㆤᰙࡢタ⨨≧ἣ࡟㛵ࡋ࡚ࡣ⠛ᒣᕷ࡟᝟ሗᥦ౪㡬࠸ࡓࠋ㈈ᅋἲே⮬↛⎔ቃ◊✲ࢭࣥࢱ࣮ࡢᖖ⏣㑥ᙪẶ࡟ࡣ◊✲ࡢ❧᱌ࡸᐇ᪋࡟
ࡈᨭ᥼ࢆ࠸ࡓࡔࡁࡲࡋࡓࠋ࡞࠾ࠊᮏ◊✲ࡣࠊᖹᡂ 17 ᖺᗘ㹼19 ᖺᗘ⎔ቃ┬ᆅ⌫⎔ቃಖ඲➼ヨ㦂◊✲㈝ཬࡧᖹᡂ 25㹼27 ᖺᗘ JSPS ⛉◊㈝㸦25450478㸧ࡢຓᡂࢆཷࡅࡲࡋࡓࠋࡇࡇ࡟ཌ
ࡃᚚ♩⏦ࡋୖࡆࡲࡍࠋ

ᘬ⏝ᩥ⊩㉥ᫍᚰ㸬2001㸬࢖ࣀࢩࢩࡢࡲࡕ㸫୹Ἴ⠛ᒣ㸬㧗ᶫ᫓ᡂ⦅ࠕ࢖ࣀࢩࢩ࡜ே㛫Ɇඹ࡟⏕ࡁࡿࠖ㸬 රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-4

pp.290-313㸬ྂ௒᭩㝔㸪ᮾி㸬ᮅ᪥⛱㸬1985㸬㈨※࡜ࡋ࡚ࡢ࢖ࣀࢩࢩ㸬့ங㢮⛉Ꮫ㸪25㸸27-30㸬
Boitani, L㸬, L㸬 Mattei, D㸬 Nonis and F㸬Corsi㸬1994㸬 Spatial and activity patternsof wild boars in Tuscay, Italy㸬Journal of Mammalogy, 75: 600-312㸬Ụཱྀ♸㍜㸬2003㸬࢖ࣀࢩࢩ࠿ࡽ⏣⏿ࢆᏲࡿ ࠾ࡶࡋࢁ⏕ែ࡜࠿ࡋࡇ࠸㜵ࡂ᪉㸬㎰ᒣ⁺ᮧᩥ໬༠఍㸪ᮾி㸪147pp㸬
Ụཱྀ♸㍜㸬2008㸬 ㎰స≀⿕ᐖᑐ⟇㸫࢖ࣀࢩࢩࡢ⿕ᐖ⟶⌮㸫㸬㧗ᵳᡂ⣖࣭ᒣᴟᑑ୍⦅ࠕ᪥ᮏ
ࡢ့ங㢮Ꮫ㸬➨஧ᕳ㸬୰኱ᆺ့ங㢮࣭㟋㛗㢮ࠖ
㸪pp.401-426㸪ᮾி኱Ꮫฟ∧㸪ᮾி㸬
Hanson, R㸬P㸬 and L㸬Karstad㸬 1959㸬 Feral swine in the southeastern United States㸬
Journal of Wildlife management, 23: 64-74㸬ᮏ⏣๛࣭ᯘ㞝୍࣭బ⸨ၿ࿴㸬2008㸬ᯘ⦕࿘㎶࡛ᤕ⋓ࡉࢀࡓ࢖ࣀࢩࢩࡢ⎔ቃ㑅ᢥ㸬့ங㢮⛉Ꮫ㸪48㸦㸯㸧
㸸11-16㸬රᗜ┴㸬2013㸬➨ 2 ᮇ࢖ࣀࢩࢩಖㆤ⟶⌮ィ⏬㸬 රᗜ┴㸪⚄ᡞ㸪18pp㸬
⎔ቃ┬⮬↛⎔ቃᒁ㸬2013㸬ᖹᡂ 24 ᖺᗘ≉ᐃ㫽⋇࡟㛵ࢃࡿಖㆤ⟶⌮᪋⟇᥎㐍ࡢࡓࡵࡢ᳨ウㄪᰝᴗົሗ࿌᭩㸪164 pp㸬
Kurz, J㸬C㸬and R㸬L㸬Marchinton㸬1972㸬 Radiotelemetry studies of feral hogs in South
Carolina㸬Journal of Wildlife management㸬36:1240-1548㸬
⚄ᓮఙኵ㸬2001㸬࢖ࣀࢩࢩࡢၟရ໬࡜ಶయ⩌⟶⌮㸬㧗ᶫ᫓ᡂ⦅ࠕ࢖ࣀࢩࢩ࡜ே㛫Ɇඹ࡟⏕ࡁࡿࠖ㸬pp.258-288㸬ྂ௒᭩㝔㸪ᮾி㸬ᑠᑎ♸஧࣭⚄ᓮఙኵ㸬2001㸬ᓥ᰿┴▼ぢᆅ᪉࡟࠾ࡅࡿࢽ࣍ࣥ࢖ࣀࢩࢩࡢ㣗ᛶ࠾ࡼࡧᰤ㣴≧ែࡢᏘ⠇ⓗኚ໬㸬 㔝⏕⏕≀ಖㆤ 6 (2): 109-117㸬ᑠ ᑎ ♸ ஧ 㸬 2001 㸬 ᓥ ᰿ ┴ ▼ ぢ ᆅ ᪉ ࡟ ࠾ ࡅ ࡿ ࢽ ࣍ ࣥ ࢖ ࣀ ࢩ ࢩ ࡢ ⎔ ቃ 㑅 ᢥ 㸬 Wildlife
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Conservation Japan 3:95-105㸬ᆏ⏣ᏹᚿ࣭㩪ᓥᘯග࣭ᶓᒣ┿ᘪ㸬2008㸬┠ᧁຠ⋡࠿ࡽࡳࡓ࢖ࣀࢩࢩࡢ⏕ᜥ≧ἣ࡜✚㞷㸪᳜⏕㸪ࢽ࣍ࣥࢪ࢝㸪⊁⊟㸪㎰ᴗ⿕ᐖ࡜ࡢ㛵ಀ㸬 ့ங㢮⛉Ꮫ 48㸸245-253㸬
㎷▱㤶࣭ᶓᒣ┿ᘪ㸬2014㸬භ⏥ᒣ࢖ࣀࢩࢩၥ㢟ࡢ⌧≧࡜ㄢ㢟㸬ࠕරᗜ┴࡟࠾ࡅࡿࢽ࣍ࣥ࢖ࣀ
ࢩࢩࡢಖㆤ⟶⌮ࡢ⌧≧࡜ㄢ㢟㸬ࠖ㸪࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6 ྕ㸪pp121-134㸪රᗜ┴᳃ᯘື≀◊✲ࢭࣥࢱ࣮㸬ᶓᒣ┿ᘪ࣭ᩪ⏣ᰤ㔛ዉ࣭Ụ⸨බಇ࣭୰ᮧᖾᏊ࣭᳃ග⏤ᶞ㸬2011㸬රᗜ┴࡟࠾ࡅࡿࢶ࢟ࣀ࣡
ࢢ࣐ࡢ⾜ືᅪࡢኚ␗࡜ࡑࡢせᅉ㸬ࠕරᗜ┴࡟࠾ࡅࡿࢶ࢟ࣀ࣡ࢢ࣐ࡢಖㆤ⟶⌮ࡢ⌧≧࡜ㄢ㢟ࠖ㸪 රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 3 ྕ㸪pp.62-73㸬 රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-5➨ 㸳 ❶            ࢽ࣍ࣥ࢖ࣀࢩࢩࡢᖺ㱋ᰝᐃ᪉ἲ
㎷ ▱㤶 1࣭ᶓᒣ┿ᘪ 1,2せ Ⅼ࣭ 2003 ᖺ࠿ࡽ 2013 ᖺ࡟⊁⊟ࡸ᭷ᐖᤕ⋓࡟ࡼࡾᤕ⋓ࡉࢀࡓಶయ➼ࠊィ 517 㢌ࢆ⏝࠸࡚ࠊୗ㢡ࡢṑࡢⴌฟ࡜஺᥮≧ἣࡢほᐹ࠾ࡼࡧ➨୍ᚋ⮻ṑࡢࢭ࣓ࣥࢺ㉁ᒙࡢᖺ㍯ᩘ
ࡢ࢝࢘ࣥࢺ࡟ࡼࡗ࡚ᖺ㱋ᰝᐃࢆᐇ᪋ࡋࠊරᗜ┴࢖ࣀࢩࢩࡢᖺ㱋ᵓᡂࢆ᫂ࡽ࠿࡟ࡋࡓࠋ࣭ ᖺ㛫㏻ࡌ࡚཰㞟ࡉࢀࡓಶయࡢṑࡢⴌฟ࡜஺᥮≧ἣࢆほᐹࡍࡿࡇ࡜࡟ࡼࡾࠊරᗜ┴࢖ࣀࢩࢩࡢ 3 ṓࡲ࡛ࡢᖺ㱋ᰝᐃ⾲ࢆసᡂࡋࡓࠋ࣭ ࡯࡜ࢇ࡝ࡢṑࡢⴌฟ࡜஺᥮࡟࠾࠸࡚ 7㹼8 ࠿᭶ࡢኚ␗ᖜࡀ☜ㄆࡉࢀࡓࡇ࡜࠿ࡽࠊ
࢖ࣀࢩࢩ࡛ࡣṑࡢⴌฟ࡜஺᥮ࡢಶయᕪࡀ኱ࡁ࠸ࡇ࡜ࡀ♧၀ࡉࢀࡓࠋࡋࡓࡀࡗ࡚ࠊ
ࡑࢀࡽࡢほᐹ࡟ࡼࡿ࢖ࣀࢩࢩࡢ᭶㱋ࡸ㐌㱋ࡢᰝᐃࡣᅔ㞴࡛࠶ࡿ࡜⪃࠼ࡽࢀࡓࠋ࣭ ศᯒಶయࡢᖺ㱋ࡣ 0 ṓ࠿ࡽ 11 ṓࡲ࡛☜ㄆࡉࢀࡓࡀࠊᖺ㱋ᵓᡂࡣ 0 ṓ࡜ 1 ṓࡀ඲యࡢ 56.3㸣ࢆ༨ࡵࡿࠊⱝ㱋ᒙࡀከ࠸ࣆ࣑ࣛࢵࢻᆺ࡛࠶ࡗࡓࠋࡇࢀࡼࡾරᗜ┴ࡢ࢖ࣀࢩࢩࡣቑຍᆺࢆ♧ࡍ୍᪉࡛ࠊẚ㍑ⓗᙉ࠸ᤕ⋓ᅽࡀ࠿࠿ࡗ࡚࠸ࡿࡇ࡜ࡀ⪃࠼ࡽࢀࡓࠋ㞤㞝ู࡟ࡳࡿ࡜ࠊⱝ㱋࡛ࡣ࢜ࢫࡢ᪉ࡀከࡃࠊ㧗㱋࡛ࡣ࣓ࢫࡢ᪉ࡀከ࠿ࡗࡓࡇ
࡜࠿ࡽࠊ࣓ࢫࡢ᪉ࡀ࢜ࢫࡼࡾࡶ㧗㱋ࡲ࡛⏕Ꮡࡋࡸࡍ࠸ࡇ࡜ࡀ⪃࠼ࡽࢀࡓࠋ

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Key words㸸ᖺ㱋ᰝᐃࠊṑࡢⴌฟ࣭஺᥮ࠊᖺ㱋ᵓᡂࠊቑຍᆺ

ࡣࡌࡵ࡟
㔝⏕ື≀࡟࠾࠸࡚ಶయࡢᖺ㱋᝟ሗࡣࠊಶయࣞ࣋ࣝ࠶ࡿ࠸ࡣ✀࡛ࣞ࣋ࣝࡢ⦾Ṫ≧ἣࡸ⾜ື≉ᛶ➼ࢆ᫂ࡽ࠿࡟ࡍࡿࡓࡵ࡟୙ྍḞ࡛࠶ࡿࠋࡲࡓಶయ⩌ࡢᖺ㱋ᵓᡂࡣࠊಶయ⩌ືែࡢᢕᥱࡸᑗ᮶ண

ࡢࡓࡵࡢᇶ┙᝟ሗ࡛࠶ࡿ㸦ᩪ⏣࡯࠿ 2011㸹ῦ㔝 2012㸧ࠋಶయ⩌ࡢ㐺ษ࡞⟶⌮ࢆ⾜࠺ࡓࡵ࡟ࡣࠊᤕ⋓ಶయࡢᖺ㱋ᵓᡂࢆᢕᥱࡋࠊᤕ⋓ࡢᙳ㡪ࢆ᳨ウࡍࡿࡇ࡜ࡶồࡵࡽࢀࡿ㸦Milkowski & Wójcik 1984; ᩪ⏣࡯࠿ 2011㸧ࠋࡘࡲࡾࠊᤕ⋓᫬ᮇࡸᤕ⋓ᡭἲࡢ㐪࠸࡟క࠺ᤕ⋓ಶయࡢᖺ㱋ࡢ೫ࡾࡢ᭷↓ࢆᢕᥱࡍࡿࡇ࡜࡛ࠊቑ࠼ࡍࡂࡓಶయ⩌࡟ᑐࡍࡿಶయᩘ๐ῶຠᯝࡸ⤯⁛ࡀ༴᝹ࡉࢀࡿಶయ⩌࡛ࡢ㑅ᢥᤕ⋓ࡢ᫝㠀ࢆ᳨ウࡍࡿࡇ࡜ࡀ㔜せ࡜࡞ࡿ㸦Apollonio et al. 2011㸧ࠋ௨ୖࡢࡼ࠺࡟ࠊಶయ⩌ࡢᖺ㱋ᵓᡂࡣࠊᴟࡵ࡚㔜せ࡞᝟ሗ࡛࠶ࡿࡇ࡜࠿ࡽࠊࡇࢀࡲ࡛ከ

1 රᗜ┴᳃ᯘື≀◊✲ࢭࣥࢱ࣮࣭2 රᗜ┴❧኱Ꮫ⮬↛࣭⎔ቃ⛉Ꮫ◊✲ᡤ රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-5ࡃࡢ㔝⏕ື≀࡛ᖺ㱋ᰝᐃࡀᐇ᪋ࡉࢀ࡚ࡁࡓ㸦ᯘ࡯࠿ 1977㸹㡲⸨ 2001㸹ᩪ⏣࡯࠿ 2011㸧ࠋ㔝⏕့ங㢮ື≀ࡢ᭱ࡶ୍⯡ⓗ࡞ᖺ㱋ᰝᐃ᪉ἲࡣࠊṑࡢⴌฟ࡜ஙṑ࠿ࡽỌஂṑ࡬ࡢ஺᥮≧ἣࡢほᐹ࡟ࡼࡿุᐃ࡛࠶ࡿ㸦㡲⸨ 2001㸹ῦ㔝 2012㸧ࠋṑࡀ඲࡚ⴌฟࡋࠊỌஂṑ࡟⨨ࡁ᥮ࢃࡗࡓಶయ࡟ࡘ࠸࡚ࡣࠊṑࡢࢭ࣓ࣥࢺ㉁ᒙ࡟ᙧᡂࡉࢀࡿᖺ㍯ᩘࡢ࢝࢘ࣥࢺࡀ⾜ࢃࢀ࡚ࡁࡓ㸦ᯘ࡯࠿ 1977㸹㡲⸨ 2001㸧ࠋ඲ࡃูࡢ᪉ἲ࡜ࡋ࡚ࠊỈᬗయࡢ㔜㔞ࡢィ࡟ࡼࡿᖺ㱋ᰝᐃࡶ⾜ࢃࢀ࡚ࡁࡓ㸦㡲⸨ 2001㸧ࠋ࢖ࣀࢩࢩ㸦Sus scrofa㸧࡛ࡣࠊ࣮ࣚࣟࢵࣃࡢ࢖ࣀࢩࢩ࡟࠾࠸࡚ࠊ㇏ᐩ࡞㢌ᩘࡢ㣫⫱ಶయࢆᑐ㇟࡟ࡋࡓṑࡢⴌฟ࡜஺᥮ࡢほᐹ࡟ࡼࡾࠊヲ⣽࡞ᖺ㱋ᰝᐃ᪉ἲࡀ☜❧ࡉࢀ࡚࠸ࡿ
㸦Matschcke 1967; Boitani & Mattei 1992㸧ࠋ୍᪉ࠊࢽ࣍ࣥ࢖ࣀࢩࢩ㸦 Sus scrofa

leucomystax㸹௨ୗ࢖ࣀࢩࢩ࡜⾲グ㸧࡛ࡣࠊᯘ࡯࠿㸦1977㸧ࡀࠊ㣫⫱ಶయ࡜෤Ꮨ࡟ᤕ⋓ࡉࢀࡓ㔝⏕ಶయࢆ⏝࠸࡚ᩚ⌮ࡋ࡚࠸ࡿࠋ᭱㏆ࠊAnezaki㸦2008㸧ࡀࠊ෤Ꮨ௨እࡢಶయࡶ
⏝࠸࡚ṑࡢⴌฟࡸ஺᥮≧ἣࢆሗ࿌ࡋ࡚࠸ࡿࠋࡋ࠿ࡋࠊᶆᮏᩘࡀ㝈ࡽࢀ࡚࠸ࡿࡇ࡜ࡸⴌฟࠊ
஺᥮ࡢ㐍⾜≧ἣ࡟ಶయᕪࡀ☜ㄆࡉࢀ࡚࠸ࡿࡇ࡜࠿ࡽࠊ෤Ꮨ௨እࡢ㔝⏕ಶయࡢᖺ㱋ᰝᐃ᪉ἲࡢᩚ⌮ࡣ༑ศ࡛࡞࠸ࠋ
㏆ᖺࠊ࢖ࣀࢩࢩࡢ⏕ᜥᇦࡣᛴ㏿࡟ᗈࡀࡾ㸦⎔ቃ┬⮬↛ಖㆤᒁ⏕≀ከᵝᛶࢭࣥࢱ࣮
2004㸧
ࠊᤕ⋓ᩘࡸ㎰స≀⿕ᐖ㔠㢠ࡶቑຍࡋ࡚࠸ࡿ㸦⎔ቃ┬ 2010ࠊ㎰Ỉ┬ 2010㸧ࡇ࡜࠿ࡽࠊ࢖ࣀࢩࢩࡢಶయᩘࡣቑຍഴྥ࡟࠶ࡿ࡜᥎

ࡉࢀ࡚࠸ࡿࠋ⌧ᅾᐇ᪋ࡋ࡚࠸ࡿ࢖ࣀࢩࢩ

ࡢಶయᩘ᥎ᐃࡣࠊᤕ⋓ᩘࡸ⊁⊟⪅࠿ࡽᐤࡏࡽࢀࡿሗ࿌࡟ᇶ࡙࠸࡚⾜ࢃࢀ࡚࠾ࡾࠊ⏕≀Ꮫⓗ࡞᝟ሗࡣ஦๓ศᕸ࡟࠾࠸࡚⪃៖ࡉࢀ࡚࠸ࡿࡀࠊࢹ࣮ࢱ࡜ࡋ࡚ࡣά⏝ࡉࢀ࡚࠸࡞࠸㸦ᆏ⏣࡯࠿ 2012㸹ᯇᮏ࡯࠿ 2014㸧ࠋ࢖ࣀࢩࢩࡣಶయᩘࡢᖺḟኚືࡶᖺෆኚືࡶ኱ࡁ࠸ࡇ࡜
㸦ᆏ⏣࡯࠿ 2012㸧ࠊࢽ࣍ࣥࢪ࢝㸦Cervus nippon㸧ࡢ⣅ሢᐦᗘㄪᰝ࡟ᑐᛂࡍࡿࡼ࠺࡞⏕ᜥᐦᗘㄪᰝࡀᏑᅾࡋ࡞࠸ࡇ࡜࠿ࡽࠊ⌧≧࡛ࡣࠊᑗ᮶ண

࡞࡝ࢆྵࡵ⢭ᗘࡢ㧗࠸᥎ᐃࡀ㞴ࡋ࠸㸦ᆏ⏣࡯࠿ 2012㸧ࠋࡇࡢࡼ࠺࡞࢖ࣀࢩࢩ≉᭷ࡢㄢ㢟ゎỴ࡟ྥࡅ࡚ࠊಶయ⩌ࡢᖺ㱋ᵓᡂ࠾ࡼࡧᖺ㱋࡟౫Ꮡࡍࡿ⦾Ṫࢹ࣮ࢱ࡞࡝ࡢ⏕≀Ꮫⓗᇶ┙᝟ሗࢆ᫂ࡽ࠿࡟ࡍࡿࡇ࡜࡛ࠊಶ
యᩘ᥎ᐃࡢไᗘࢆ㧗ࡵࡽࢀࡿྍ⬟ᛶࡀ࠶ࡿࠋ
ࡑࡇ࡛ࠊᮏ◊✲࡛ࡣࠊᖺ㛫㏻ࡌ࡚཰㞟ࡉࢀࡓಶయࢆ⏝࠸࡚ࠊ㏻ᖺ࡛ྍ⬟࡜࡞ࡿᖺ㱋ᰝᐃ᪉ἲࡢᩚ⌮ࢆヨࡳࠊරᗜ┴ࡢ࢖ࣀࢩࢩࡢᖺ㱋ᵓᡂࢆ᫂ࡽ࠿࡟ࡍࡿࡇ࡜ࢆ┠ⓗ࡜ࡋࡓࠋᮦᩱ࡜᪉ἲ
ᮦᩱ
 2003 ᖺ࠿ࡽ 2013 ᖺ࡟࠾࠸࡚ࠊ⊁⊟࠾ࡼࡧ᭷ᐖᤕ⋓࡟ࡼࡾᤕ⋓ࡉࢀࡓಶయࡸ஺㏻஦ᨾṚ➼࡟ࡼࡾᅇ཰ࡉࢀࡓಶయࠊྜィ 517 㢌ࢆᑐ㇟࡜ࡋࠊᮦᩱ࡜ࡋ࡚㢌㦵ࢆ᥇ྲྀࡋࡓࠋ
᪉ἲ ࢖ࣀࢩࢩࡢṑᘧࡣࠊୖୗᕥྑࡑࢀࡒࢀษṑ㸦I㸧3 ᮏࠊ≟ṑ㸦C㸧1 ᮏࠊ๓⮻ṑ㸦P㸧4ᮏࠊᚋ⮻ṑ㸦M㸧3 ᮏࡢィ 44 ᮏ࡛࠶ࡾ㸦Matschcke 1967; ᯘ࡯࠿ 1977; Boitani & Mattei රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-5

1992; Anezaki 2009㸧ࠊࡇࢀࡽࡢỌஂṑࡀ⏕࠼ࡑࢁ࠺ࡢࡀ 38 ࠿᭶௨ෆ࡜ሗ࿌ࡉࢀ࡚࠸ࡿ
㸦ᯘ࡯࠿ 1977㸧ࠋࡲࡓࠊ࢖ࣀࢩࢩࡢṑࡢⴌฟ࡜஺᥮ࡣࠊୖ㢡ࡢ᪉ࡀୗ㢡ࡼࡾࡶಶయᕪࡀ
኱ࡁ࠸࡜ሗ࿌ࡉࢀ࡚࠸ࡿ㸦Matschcke 1967; ᯘ࡯࠿ 1977㸧ࠋࡑࡇ࡛ᮏ◊✲࡛ࡣࠊ3 ṓ௨ୗࡢಶయࡢᖺ㱋ᰝᐃ࡜ࡋ࡚ࠊୗ㢡ࡢṑࡢⴌฟ࡜஺᥮≧ἣࢆほᐹࡋࡓ㸦Matschcke
1967; ᯘ࡯࠿ 1977; Boitani & Mattei 1992; Anezaki 2009㸧ࠋࡓࡔࡋ➨୍๓⮻ṑࢆḞኻࡋ࡚࠸ࡿಶయࡀከ࠿ࡗࡓࡓࡵࠊ➨୍๓⮻ṑࡣᖺ㱋ᰝᐃࡢᣦᶆ࠿ࡽ㝖እࡋࡓࠋḟ࡟඲࡚ࡢṑࡢⴌฟ࡜஺᥮ࡀ᏶஢ࡋ࡚࠸ࡓಶయࠊࡘࡲࡾ 3 ṓࡼࡾࡶ㧗㱋ࡢಶయࡢᖺ㱋ᰝᐃ࡜ࡋ࡚ࠊ
➨୍ᚋ⮻ṑࡢࢭ࣓ࣥࢺ㉁ᒙࡢᖺ㍯ᩘࢆ࢝࢘ࣥࢺࡋࡓ㸦ᯘ࡯࠿ 1977㸧ࠋᖺ㍯ࡢほᐹࡢࡓࡵ࡟ࠊ➨୍ᚋ⮻ṑࢆᢤṑࡋ࡚⬺⅊ฎ⌮ࢆ᪋ࡋࡓᚋ 35㹼40ȣm ࡟ⷧษࡋࠊࡑࡢษ∦࡟࣐࣊ࢺ࢟ࢩࣜࣥᰁⰍࢆ⾜ࡗࡓ㸦㡲⸨ 2001㸧ࠋ ᮏ◊✲࡛ࡣࠊᯘ࡯࠿㸦1977㸧࡟ᇶ࡙࠸࡚ࠊ➨୍ᚋ⮻ṑ㸦M1㸧ࠊ➨஧ᚋ⮻ṑ㸦M2㸧ࠊ➨୕ᚋ⮻ṑ㸦M3㸧ࡢⴌฟ≧ែ࡟ࡼࡾࠊᑐ㇟ಶయࢆ 5 ࡘࡢ㱋ࢢ࣮ࣝࣉ࡟༊ศࡋࡓ㸦⾲㸯㸧ࠋ⾲㱋ࢢ࣮ࣝࣉࡢ༊ศ᪉ἲ
㱋ࢢ࣮ࣝࣉ

ᚋ⮻ṑࡢⴌฟ≧ែ,0 ⴌฟ᏶஢ࡲ࡛ϩ0 ⴌฟ㛤ጞ㹼᏶஢Ϫ0 ⴌฟ㛤ጞ㹼0 ➨  ည㢌ⴌฟϫ0 ➨  ည㢌ⴌฟ㛤ጞ㹼᏶඲ⴌฟࠊᦶ⁛࡞ࡋϬ0 ᏶඲ⴌฟࠊᦶ⁛࠶ࡾୖグࡢ 5 ࢢ࣮ࣝࣉ࡟༊ศᚋࠊࡲࡎࠊࡑࢀࡒࢀࡢࢢ࣮ࣝࣉࡢᚋ⮻ṑࡢⴌฟ≧ἣࢆ☜ㄆࡋࡓࠋ➨୍ࠊ➨஧ᚋ⮻ṑ࡛ࡣࠊⴌฟࡋ࡚࠸࡞࠸ሙྜࡣ NP㸦Non-Eruption㸧ࠊⴌฟ୰ࡣ EP
㸦Erupting㸧ࠊⴌฟ᏶஢ࡢሙྜࡣ E㸦Erupted㸧࡜⾲グࡋࡓࠋ➨୕ᚋ⮻ṑ࡛ࡣࠊည㢌ࡢⴌฟ≧ἣ࡛ࠊ➨୍ࠊ஧ည㢌ࡲ࡛ࡢሙྜ M3-1ࠊ➨୕ࠊᅄည㢌ࡲ࡛ࡢሙྜ M3-2ࠊ➨஬ࠊභည㢌ࡲ࡛ࡢሙྜ M3-3ࠊ᭱ᚋࡢ➨୐ည㢌ࡲ࡛☜ㄆ࡛ࡁࡓሙྜ M3-L ࡜グ㍕ࡋࡓࠋḟ࡟ษṑࠊ≟ṑࠊ๓⮻ṑࡢஙṑ࠿ࡽỌஂṑ࡬ࡢ஺᥮≧ἣࢆ☜ㄆࡋࠊஙṑࡢሙྜࡣࠕᑠᩥᏐࠖࠊ
஺᥮୰ࡣࠕᑠᩥᏐЍ኱ᩥᏐࠖࠊỌஂṑࡢሙྜࡣࠕ኱ᩥᏐ࡛ࠖ࢔ࣝࣇ࢓࣋ࢵࢺ⾲グࡋࡓࠋ௨ୖࡢṑࡢⴌฟ࡜஺᥮≧ἣࡢほᐹ⤖ᯝࢆࡶ࡜࡟ᖺ㱋ᰝᐃ⾲ࢆసᡂࡋࡓࠋ
సᡂࡋࡓᖺ㱋ᰝᐃ⾲࡜ࢭ࣓ࣥࢺ㉁ᒙࡢᖺ㍯ᩘ࡟ࡼࡾࠊ඲࡚ࡢಶయࡢᖺ㱋ࢆỴᐃࡋࡓࠋࡇࡢ㝿ࠊ౽ᐅୖࡢ௬ࡢฟ⏕᪥ࢆ 5 ᭶ 1 ᪥࡜ࡋ࡚㸦ᯘ࡯࠿ 1977㸹す 2003㸧ࠊ5 ᭶ 1 ᪥ࢆ㐣ࡂࡓሙྜ࡟ࡣᖺ㱋ࢆ୍ࡘ⧞ࡾୖࡆࡓࠋ රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-5

⤖ᯝ㸬ᖺ㱋ᰝᐃ㱋ࢢ࣮ࣝࣉࡢ༊ศ ศᯒಶయࡣࠊϨࢢ࣮ࣝࣉ࡟ 145 㢌ࠊϩࢢ࣮ࣝࣉ࡟ 158 㢌ࠊϪࢢ࣮ࣝࣉ࡟ 78 㢌ࠊϫࢢ࣮ࣝࣉ࡟ 19 㢌ࠊϬࢢ࣮ࣝࣉ࡟ 117 㢌ࡀ༊ศࡉࢀࠊ࡝ࡢࢢ࣮ࣝࣉࡶ⊁⊟ᮇ㛫ࡢ 11㹼2 ᭶࡟ከ࠿ࡗࡓ㸦ᅗ㸯㸧ࠋᅗ ᤕ⋓᭶ࡈ࡜ࡢྛ㱋ࢢ࣮ࣝࣉࡢ☜ㄆᩘϨࢢ࣮ࣝࣉ
➨୍ᚋ⮻ṑࡀᮍⴌฟࡢಶయ㸦M1NE㸧ࡣ 6㹼12 ᭶ࠊⴌฟ୰࡛࠶ࡗࡓಶయ㸦M1EP㸧ࡣ
7 ᭶࡜ 9㹼2 ᭶ࠊⴌฟ᏶஢ಶయ㸦M1E㸧ࡣ 7 ᭶࡜ 9㹼5 ᭶࡟☜ㄆࡉࢀࡓ㸦ᅗ㸰.㹟㸧ࠋஙṑ࠿ࡽỌஂṑ࡬ࡢ஺᥮ࡀ᭱ึ࡟ぢࡽࢀࡓࡢࡣ➨୕ษṑ࡜≟ṑ࡛࠶ࡗࡓࠋ஺᥮୰࡛࠶ࡗࡓಶ
య㸦c,i3ЍC,I3㸧ࡣ 10㹼5 ᭶࡟☜ㄆࡉࢀࡓࡀࠊỌஂṑ࡟⨨ࡁ᥮ࢃࡗࡓಶయ㸦C,I3㸧ࡣϨࢢ࣮ࣝࣉࡢ୰࡛ࡣ☜ㄆࡉࢀ࡞࠿ࡗࡓ㸦ᅗ㸰.㹠㸧ࠋϩࢢ࣮ࣝࣉ
➨஧ᚋ⮻ṑࡀⴌฟ୰࡛࠶ࡗࡓಶయ㸦M2EP㸧ࡣ 4㹼8 ᭶ࠊ10㹼11 ᭶ࠊ1 ᭶ࠊⴌฟࡀ᏶
஢ࡋࡓಶయ㸦M2E㸧ࡣ 5㹼3 ᭶࡟☜ㄆࡉࢀࡓ㸦ᅗ㸱.㹟㸧
ࠋஙṑ࠿ࡽỌஂṑ࡬ࡢ஺᥮≧࡟ࡘ࠸࡚ࡣࠊ➨୕ษṑ࡜≟ṑࡀஙṑ࡛࠶ࡗࡓಶయ㸦c,i3㸧ࡣ 4ࠊ6ࠊ7ࠊ11 ᭶ࠊ஺᥮୰࡛࠶
ࡗࡓಶయ㸦c,i3ЍC,I3㸧ࡣ 4㹼6 ᭶࡟ຍ࠼ࠊ኱ࡁࡃእࢀ࡚ 10 ᭶࡜ 1 ᭶࡟ほᐹࡉࢀࡓࠋỌஂṑ࡛࠶ࡗࡓಶయ㸦C,I3㸧ࡣ඲ᮇ㛫୰࡟☜ㄆࡉࢀࡓࠋ㸦ᅗ㸱.㹠㸧ࠋ➨୍ษṑࡀஙṑ࡛࠶ࡗࡓಶయ㸦i1㸧ࡣ 4㹼1 ᭶ࠊ஺᥮୰࡛࠶ࡗࡓಶయ㸦i1ЍI1㸧ࡣ 5㹼10 ᭶࡟ຍ࠼ࠊ኱ࡁࡃ
እࢀ࡚ 2 ᭶࡟☜ㄆࡉࢀࡓࠋỌஂṑ࡛࠶ࡗࡓಶయ㸦I1㸧ࡣ 5㹼3 ᭶࡟☜ㄆࡉࢀࡓࠋ
㸦ᅗ㸱.

 

රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-5

㹠 

㹟 

ᅗ Ϩࢢ࣮ࣝࣉ࡟࠾ࡅࡿࠊᤕ⋓᭶ࡈ࡜ࡢⴌฟࠊ஺᥮☜ㄆᩘ
㸦D㸧ࡣ➨୍ᚋ⮻ṑࡢⴌฟ≧ἣࢆ♧ࡋࠊ01( ࡣᮍⴌฟࠊ0(3 ࡣⴌฟ୰ࠊ0( ࡣⴌฟ᏶஢ಶ
యࢆ⾲ࡍࠋ㸦E㸧ࡣ➨୕ษṑ࡜≟ṑࡢஙṑ࠿ࡽỌஂṑ࡬ࡢ஺᥮≧ἣࢆ♧ࡋࠊFL ࡣஙṑࠊ
FLЍ&, ࡣ஺᥮୰ࢆ⾲ࡍࠋ㹡㸧ࠋ➨஧ษṑࡀஙṑ࡛࠶ࡗࡓಶయ㸦i2㸧ࡣ 4㹼2 ᭶ࠊ஺᥮୰࡛࠶ࡗࡓಶయ㸦i2ЍI2㸧ࡣ
5㹼6 ᭶ࠊ11㹼3 ᭶ࠊỌஂṑ࡛࠶ࡗࡓಶయ㸦I2㸧ࡣ 5㹼6 ᭶ࠊ8㹼9 ᭶ࠊ11㹼3 ᭶࡟☜ㄆࡉࢀࡓࠋ㸦ᅗ㸱.㹢㸧๓⮻ṑࡀஙṑ࡛࠶ࡗࡓಶయ㸦p㸧ࡣ 4㹼11 ᭶ࠊ1㹼2 ᭶ࠊ஺᥮୰࡛࠶ࡗ
ࡓಶయ㸦pЍP㸧ࡣ 6㹼1 ᭶ࠊỌஂṑ࡛࠶ࡗࡓಶయ㸦P㸧ࡣ 5㹼6 ᭶ࠊ8㹼9 ᭶ࠊ11㹼3 ᭶࡟☜ㄆࡉࢀࡓࠋ
㸦ᅗ㸱.㹣㸧ࠋ ϩࢢ࣮ࣝࣉ࡟࡚ࠊ඲࡚ࡢஙṑࡀỌஂṑ࡟⨨ࡁ᥮ࢃࡗࡓࠋࡑࢀࡒࢀࡢ⏕࠼᥮ࢃࡾᮇ㛫࡟ኚ␗ᖜࡣ࠶ࡿࡶࡢࡢࠊ஺᥮ࡢ㡰␒࡜ࡋ࡚ࡣࠊ➨୕ษṑ࡜≟ṑࠊ➨୍ษṑࠊ๓⮻ṑࠊ➨஧ษṑࡢ㡰࡛࠶ࡗࡓ㸦ᅗ㸰㸪㸱㸧ࠋϪࢢ࣮ࣝࣉ ➨୕ᚋ⮻ṑࡢ➨୍ࠊ஧ည㢌ࡀⴌฟࡋࡓಶయ㸦M3-1㸧ࡣ 10 ᭶࡜ 1㹼8 ᭶ࠊ➨୕ࠊᅄည㢌ࡀⴌฟࡋࡓಶయ㸦M3-2㸧ࡣ 7㹼8 ᭶ࠊ11㹼12 ᭶ࠊ2 ᭶ࠊ➨஬ࠊභည㢌ࡀⴌฟ㏵୰ࡢ
ಶయ㸦M3-2ЍM3-3㸧ࡣ 7㹼4 ᭶ࠊ➨஬ࠊභည㢌ࡀⴌฟࡋࡓಶయ㸦M3-3㸧ࡣ 9 ᭶࡜ 11
㹼6 ᭶࡟☜ㄆࡉࢀࡓ㸦ᅗ㸲㸧ࠋ࡝ࡢⴌฟ≧ἣ࡟࠾࠸࡚ࡶࠊ☜ㄆࡉࢀࡿᮇ㛫ࡀ㛗ࡃࡤࡽࡘ
࠸࡚࠸ࡓࠋ

 

රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-5

㹟 

㹠 


㸦㹡㸧

㸦㹢㸧


㸦㹣㸧

ᅗ ϩࢢ࣮ࣝࣉ࡟࠾ࡅࡿࠊᤕ⋓᭶ࡈ࡜ࡢⴌฟࠊ஺᥮☜ㄆᩘ
㸦D㸧ࡣ➨஧ᚋ⮻ṑࡢⴌฟ≧ἣࢆ♧ࡋࠊ0(3 ࡣⴌฟ୰ࠊ0( ࡣⴌฟ᏶஢ಶయࢆ⾲ࡍࠋ㸦E㸧ࡣ≟ṑ࡜➨୕ษṑࡢஙṑ࠿ࡽỌஂṑ࡬ࡢ஺᥮≧ἣࢆ♧ࡋࠊFL ࡣஙṑࠊFLЍ&, ࡣ஺
᥮୰ࠊ&, ࡣỌஂṑࢆ⾲ࡍࠋ㸦&㸧ࡣ➨୍ษṑࡢஙṑ࠿ࡽỌஂṑ࡬ࡢ஺᥮≧ἣࢆ♧ࡋࠊLࡣஙṑࠊLЍ, ࡣ஺᥮୰ࠊ, ࡣỌஂṑࢆ⾲ࡍࠋ㸦G㸧➨஧ษṑࡢஙṑ࠿ࡽỌஂṑ࡬ࡢ஺᥮≧ἣࢆ♧ࡋࠊL ࡣஙṑࠊLЍ, ࡣ஺᥮୰ࠊ, ࡣỌஂṑࢆ⾲ࡍࠋ
㸦H㸧ࡣ๓⮻ṑࡢஙṑ࠿ࡽỌஂṑ࡬ࡢ஺᥮≧ἣࢆ♧ࡋࠊS ࡣஙṑࠊSЍ3 ࡣ஺᥮୰ࠊ3 ࡣỌஂṑࢆ⾲ࡍࠋ



 

රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-5

ᅗ Ϫࢢ࣮ࣝࣉ࡟࠾ࡅࡿࠊᤕ⋓᭶ࡈ࡜ࡢⴌฟ☜ㄆᩘ➨୕ᚋ⮻ṑࡢⴌฟ≧ἣࢆ♧ࡋࠊ0 ࡣ➨୍ࠊ஧ည㢌ࡀⴌฟࡋࡓ≧ែࠊ0 ࡣ➨୕ࠊᅄည㢌ࡀⴌฟࡋࡓ≧ែ 0 ࡣ➨஬ࠊභည㢌ࡀⴌฟࡋࡓ≧ែࢆ⾲ࡍࠋ0Ѝ0ࡣࠊ➨஬ࠊභည㢌ࡀⴌฟ୰ࡢ≧ែࢆ⾲ࡍࠋϫࠊϬࢢ࣮ࣝࣉ ➨୕ᚋ⮻ṑࡀ➨୐ည㢌ࡲ࡛඲࡚ⴌฟࡋࡓಶయ࡟࠾࠸࡚ࠊṑࡢᦶ⁛ࡀ☜ㄆࡉࢀ࡞࠿ࡗࡓ
ϫࢢ࣮ࣝࣉࡣ 8 ᭶࡜ 10㹼3 ᭶࡟☜ㄆࡉࢀࡓ㸦ᅗ㸳㹟㸧ࠋࡲࡓࠊᦶ⁛ࡀ☜ㄆࡉࢀࡓϬࢢ࣮ࣝࣉࡢಶయࡣ඲࡚ࡢᮇ㛫୰࡟☜ㄆࡉࢀࡓ㸦ᅗ㸳㹠㸧ࠋ

㸦㹟㸧

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ᅗ ϫࠊϬࢢ࣮ࣝࣉ࡟࠾ࡅࡿⴌฟ࡜ᦶ⁛≧ἣ
㸦D㸧ࡣ➨୕ᚋ⮻ṑࡢⴌฟࡀ᏶஢ࡋṑࡢᦶ⁛ࡀ☜ㄆࡉࢀ࡞࠿ࡗࡓಶయࠊ㸦E㸧ࡣᦶ⁛ࡀ☜ㄆࡉࢀࡓಶయࢆ♧ࡍࠋṑࡢⴌฟ࡜஺᥮ࡣࠊ඲యⓗ࡟ኚ␗ᖜࡀ㢧ⴭ࡟኱ࡁ࠿ࡗࡓࡀࠊࡑࡢኚ␗ᖜࢆຍ࿡ࡋ࡚ࡲ
࡜ࡵࡿ࡜௨ୗࡢ㏻ࡾ࡜࡞ࡗࡓࠋࡲࡎ 0 ṓ࡛ࡣ෤࡟➨୍ᚋ⮻ṑ㸦M1㸧ࡢⴌฟࡀ᏶஢ࡋ㸦ᅗ
㸰.㹟㸧
ࠊ෤࠿ࡽ⩣᫓࡟࠿ࡅ࡚ࠊ➨୕ษṑ㸦I3㸧࡜≟ṑ㸦C㸧ࡀ⏕࠼ኚࢃࡗ࡚࠸ࡓ㸦ᅗ㸰.
㹠, ᅗ㸱.㹠㸧
ࠋḟ࡟ 1 ṓࢆ㏄࠼ࡿ᫓࠿ࡽึኟ࡟࠿ࡅ࡚➨஧ᚋ⮻ṑ㸦M2㸧ࡀⴌฟࡋ࡚㸦ᅗ
㸱.㹟㸧ࠊ➨୍ษṑ㸦I1㸧ࡀ⏕࠼ኚࢃࡗࡓ㸦ᅗ㸱.㹡㸧ࠋ1 ṓࡢኟ㹼⛅࡟࠿ࡅ࡚๓⮻ṑ㸦P㸧
ࡀ⏕࠼ኚࢃࡗࡓ㸦ᅗ㸱.㹣㸧
ࠋࡑࡋ࡚㸯ṓࡢ⛅࠿ࡽ෤࡟࠿ࡅ࡚ࠊ➨஧ษṑ㸦I2㸧ࡢ⏕࠼ኚ රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-5ࢃࡾ㸦ᅗ㸱.㹢㸧࡜➨୕ᚋ⮻ṑࡢⴌฟࡀ㛤ጞࡋࡓࠋࡑࡢᚋࠊ2 ṓࡢ⛅࠿ࡽ෤࡟࠿ࡅ࡚➨୕ᚋ⮻ṑࡢ➨୕ࠊᅄည㢌࠾ࡼࡧ➨஬ࠊභည㢌ࡀⴌฟࡋࠊ3 ṓࡢ⛅࠿ࡽ෤࡟࠿ࡅ࡚᭱ᚋࡢ➨୐ည㢌ࡀⴌฟࡋࡓ㸦ᅗ㸲㸧ࠋ௨ୖࢆࡲ࡜ࡵࡓරᗜ┴ࡢ࢖ࣀࢩࢩࡢ࠾࠾ࡼࡑࡢᖺ㱋ᰝᐃ⾲ࢆᅗ㸴࡟♧ࡍࠋ

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ᅗ රᗜ┴࢖ࣀࢩࢩࡢᖺ㱋ᰝᐃ⾲, ࡣ➨୍ษṑࠊ, ࡣ➨஧ษṑࠊ, ࡣ➨୕ษṑࠊF ࡣ≟ṑࠊ3 ࡣ๓⮻ṑࠊ0 ࡣ➨୍ᚋ⮻ṑࠊ0 ࡣ➨஧ᚋ⮻ṑࠊ0 ࡣ➨୕ᚋ⮻ṑࢆ♧ࡍࠋࡲࡓ➨୕ᚋ⮻ṑ࡟࠾࠸࡚ࠊ0 ࡣ➨୍ࠊ஧ည㢌ࡲ࡛ࠊ0 ࡣ➨୕ࠊᅄည㢌ࡲ࡛ࠊ0 ࡣ➨஬ࠊභည㢌ࡲ࡛ⴌฟࡋ࡚࠸ࡿ≧ែࢆ♧ࡍࠋ㸬රᗜ┴ࡢ࢖ࣀࢩࢩࡢᖺ㱋ᵓᡂ㞤㞝ࡀุᐃ࡛ࡁࡓ 451 㢌ࡢᖺ㱋ࡣ 0㹼11 ṓࡲ࡛☜ㄆࡉࢀࠊᖺ㱋ᵓᡂࡣࣆ࣑ࣛࢵࢻᆺ
ࢆ♧ࡋࡓ㸦ᅗ㸵㸧ࠋ඲య࡟ᑐࡍࡿ 0 ṓࡢ๭ྜࡣࠊ࢜ࢫ࡛ࡣ 34.6㸣㸦N=81㸧ࠊ࣓ࢫ࡛ࡣ 24.4%
㸦N=67㸧ࠊ1 ṓࡢ๭ྜࡣࠊ࢜ࢫ࡛ࡣ 31.2%㸦N=73㸧ࠊ࣓ࢫ࡛ࡣ 25.5%㸦N=70㸧࡜࡞ࡾࠊ0 ṓ࡜ 1 ṓ࡛඲యࡢ༙ᩘࢆ༨ࡵࡓࠋ㞤㞝ࡢ೫ࡾ࡟ࡘ࠸࡚ࡣࠊ0㹼1 ṓ࡛ࡣ࢜ࢫࡢ᪉ࡀከ
࠿ࡗࡓࡀࠊ2 ṓ௨㝆ࡣ࣓ࢫࡢ᪉ࡀከ࠿ࡗࡓࠋ
᭱㧗ᖺ㱋ࡣ࢜ࢫ࡛ࡣ 8 ṓ࡛࠶ࡗࡓࡢ࡟ᑐࡋࠊ
࣓ࢫ࡛ࡣ 11 ṓ࡛࠶ࡗࡓࠋ රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-5

ᅗ රᗜ┴࢖ࣀࢩࢩࡢᖺ㱋ᵓᡂ㸬⪃ᐹ
㸬ᖺ㱋ᰝᐃ᪉ἲ ⴌฟࠊ஺᥮᫬ᮇࡢኚ␗ᖜ
ᮏ◊✲ࡼࡾࠊ࢖ࣀࢩࢩࡢୗ㢡ࡢṑࡢⴌฟ࡜஺᥮᫬ᮇ࡟ࡣኚ␗ᖜࡀ኱ࡁ࠸ࡇ࡜ࡀ᫂ࡽ࠿
࡜࡞ࡗࡓࠋⴌฟ୰ࡸ஺᥮୰࡛࠶ࡗࡓಶయ࡟㝈ᐃࡋ࡚ࡶほᐹࡉࢀࡿኚ␗ࡢᮇ㛫ࡣ 6㹼8 ࠿᭶࡟ཬࢇࡔ㸦ᅗ㸰ࠊ㸱㸧ࠋࡇࡢኚ␗ᖜࡀ⏕ࡌࡿせᅉࡣ 2 ࡘᣲࡆࡽࢀࡿࠋ୍ࡘࡣࠊࡶ࡜ࡶ
࡜ࡢฟ⏕᫬ᮇࡢኚ␗ᖜ࡟㉳ᅉࡍࡿࡶࡢ࡛ࠊࡶ࠺୍ࡘࡣࠊฟ⏕ᚋࡢṑࡢⴌฟ࡜஺᥮ࡢ㐍⾜≧ἣࡢಶయᕪ࡟㉳ᅉࡍࡿࡶࡢ࡛࠶ࡿࠋࡇࡢ୰࡛๓⪅࡟ࡘ࠸࡚ࠊරᗜ┴ࡢ࢖ࣀࢩࢩ࡛ࡣࠊ
⫾㱋᥎ᐃࡼࡾ⟬ฟࡉࢀࡓฟ⏘᫬ᮇࡢኚ␗ᖜࡣࠊ⣙ 5 ࠿᭶࡜ሗ࿌ࡉࢀ࡚࠸ࡿ㸦㎷࣭ᶓᒣ
2014㸧ࠋᮏ◊✲ࡢ⤖ᯝࡣࠊฟ⏘᫬ᮇࡢኚ␗ᖜࡼࡾࡶ㛗࠿ࡗࡓࡇ࡜࠿ࡽࠊฟ⏕ᚋࡢṑࡢⴌฟ࡜஺᥮≧ἣ࡟࠾࠸࡚ࡶಶయᕪࡀ⏕ࡌ࡚࠸ࡿࡇ࡜ࡀ♧၀ࡉࢀࡓࠋ㐣ཤࡢ◊✲࡟࠾࠸࡚ࡶ
ṑࡢⴌฟ࡜஺᥮᫬ᮇࡣࠊಶయࡢᰤ㣴≧ែࡸ⑌⑓ࠊ⦾Ṫ≧ἣ࡞࡝ᵝࠎ࡞せᅉࡀᙳ㡪ࡋ࡚ࠊᖜࡀ⏕ࡌࡿࡇ࡜ࡀሗ࿌ࡉࢀ࡚࠸ࡿ㸦Boitani & Mattei 1992; Bridault et al. 2000;
Mangnel & Carter 2007; Anezaki 2009㸧ࠋࡉࡽ࡟⏕⫱⎔ቃࡀ⤫୍ࡉࢀ࡚࠸ࡿ㣫⫱ಶయ
࡛ࡶ 0.3㹼4.5 ࠿᭶ࡢಶయᕪࡀሗ࿌ࡉࢀ࡚࠸ࡿ㸦Matschke 1967㸧ࠋࡋࡓࡀࡗ࡚ࠊ࢖ࣀࢩ
ࢩࡢṑࡢⴌฟ࡜஺᥮᫬ᮇࡢኚ␗ᖜࡣࠊ༢࡟ฟ⏕᫬ᮇࡢࡤࡽࡘࡁࡔࡅ࡛ࡣ࡞ࡃࠊᡂ㛗ẁ㝵࡟࠾ࡅࡿಶయᕪࡀᙉࡃᙳ㡪ࡍࡿࡇ࡜ࡀ♧၀ࡉࢀࡿࡓࡵࠊ᭶㱋ᰝᐃࡸ㐌㱋ᰝᐃࡣ㞴ࡋ࠸࡜⪃࠼ࡽࢀࡓࠋ රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-5

࢖ࣀࢩࢩࡢᖺ㱋ᰝᐃ ᮏ◊✲࡛ࡣࠊ๓㏙ࡢ࡜࠾ࡾṑࡢⴌฟ࡜஺᥮࡟ኚ␗ᖜࡣ࠶ࡿࡶࡢࡢࠊ࠾࠾ࡼࡑࡢᖺ㱋ᰝᐃࡢ┠Ᏻࢆసᡂࡋࡓ㸦ᅗ㸴㸧ࠋୗ㢡ࡢṑࡢⴌฟ࡜஺᥮ࡀほᐹࡉࢀࡿ㡰␒࡟ࡘ࠸࡚ࡣ
Matschcke㸦1967㸧ࡢሗ࿌࡜୍⮴ࡋࡓࡀࠊBoitani & Mattei㸦1992㸧ࡸ Anezaki 㸦2009㸧ࡣࠊ➨஧ษṑࡢⴌฟࡼࡾࡶඛ࡟➨୍ษṑࡢ஺᥮ࡀඛ࡟㉳ࡇࡿ࡜ሗ࿌ࡋ࡚࠾ࡾ┦㐪ࡀ࠶ࡗࡓࠋṑࡢⴌฟ࡜஺᥮᫬ᮇ࡟࠾࠸࡚ࡶࠊ㐣ཤࡢሗ࿌㸦Matschcke 1967; Boitani & Mattei
1992; Anezaki 2009㸧࡜୍⮴ࡍࡿ㒊ศࠊ๓ᚋࡍࡿ㒊ศࡀࡑࢀࡒࢀ☜ㄆࡉࢀࡓࠋࡋࡓࡀࡗ
࡚ࠊ࢖ࣀࢩࢩࡢṑࡢⴌฟ࡜஺᥮ࡢ㡰␒ࡸ᫬ᮇ࡟ࡘ࠸࡚ࡣࠊಶయᕪ࠾ࡼࡧᑐ㇟ಶయ⩌ࡸᑐ㇟ᆅᇦ࡛ࡢኚ␗ࡶ኱ࡁ࠸࡜⪃࠼ࡽࢀࠊ඲ᅜ୍ᚊ࡜⪃࠼ࡿ࡜ࡉࡲࡊࡲ࡞ㄗᕪࢆ⏕ࡳฟࡍᜍࢀࡀ࠶ࡿࠋࡑࡢࡓࡵࠊᮏ◊✲ࡢᖺ㱋ᰝᐃ⾲ࡶ࠾࠾ࡼࡑࡢ┠Ᏻ࡜ࡋ࡚౑⏝ࡍࡿࡇ࡜ࡀᮃࡲ
ࢀࡿࠋࡲࡓࠊ➨୕ᚋ⮻ṑࡢⴌฟ≧ἣࡣ࡜ࡃ࡟ኚ␗ᖜࡀ኱ࡁࡃ࡞ࡿࡇ࡜࠿ࡽ㸦ᅗ㸲㸹
Matschcke 1967; Boitani & Mattei 1992; Anezaki 2009㸧ࠊほᐹࡉࢀࡿ᫬ᮇ࡟ࡼࡗ࡚ࡣࠊ
ⴌฟࡔࡅ࡛ᖺ㱋ᰝᐃࢆࡍࡿࡢ࡛ࡣ࡞ࡃࠊ➨୍ᚋ⮻ṑࡢࢭ࣓ࣥࢺ㉁ᒙࡢᖺ㍯ᩘࡢ࢝࢘ࣥࢺ
ࡶ⾜ࡗ࡚ࠊ཮᪉ࡢ⤖ᯝࢆࡶ࡜࡟ᖺ㱋ࢆỴᐃࡍࡿࡇ࡜ࡀᚲせ࡜⪃࠼ࡽࢀࡓࠋ
➨୍ษṑ࡜➨୕ษṑࠊ≟ṑࡢ஺᥮≧ἣ➼࡟࠾࠸࡚ࠊ᫂ࡽ࠿࡟᫬ᮇࡀࡎࢀ࡚࠸ࡿಶయࡀ
1㹼2 㢌☜ㄆࡉࢀࡓ㸦ᅗ㸱㸧ࠋࡇࢀࡽࡣࠊ᫓௨እ࡟⏕ࡲࢀࡓಶయࠊࡘࡲࡾ⛅࡟⏕ࡲࢀࡓಶయ࡜⪃࠼ࡽࢀࡓࠋࡋࡓࡀࡗ࡚ࠊṑࡢⴌฟࠊ஺᥮≧ἣࡢ☜ㄆ࡟࡚ࠊ⛅Ꮚࡢ๭ྜࡸࡑࡢ⏕Ꮡ⋡ࢆ᥎ᐃ࡛ࡁࡿྍ⬟ᛶࡀᣲࡆࡽࢀࠊ௒ᚋࡉࡽ࡞ࡿ᳨ウࡀᚲせ࡜⪃࠼ࡿࠋ㸬රᗜ┴࢖ࣀࢩࢩࡢᖺ㱋ᵓᡂᤕ⋓ಶయࡢᖺ㱋ᵓᡂࡣࠊ㔝⏕ࡢಶయ⩌ࢆᚲࡎࡋࡶ཯ᫎࡍࡿ࡜ࡣ㝈ࡽ࡞࠸ࡇ࡜ࡀᣦ᦬ࡉ
ࢀ࡚࠸ࡿ㸦ᩪ⏣ 2011㸧ࡀࠊᮏ◊✲ࡣ㏻ᖺ࡛཰㞟ࡋࡓ㇏ᐩ࡞ᶆᮏᩘ࡟᳨࡚ウࡋࡓࡇ࡜࡛ࠊ
࠶ࡿ⛬ᗘࡢุ᩿ࡀ࡛ࡁࡿ࡜⪃࠼ࡿࠋࡲࡎරᗜ┴ࡢ࢖ࣀࢩࢩࡢᖺ㱋ᵓᡂࡣࠊⱝ㱋ᒙࡢከ࠸ࣆ࣑ࣛࢵࢻᆺࢆ♧ࡋࡓ㸦ᅗ㸵㸧ࡇ࡜࠿ࡽࠊಶయᩘࡣቑຍഴྥ࡟࠶ࡿࡇ࡜ࡀ♧၀ࡉࢀࡓࠋ
࢖ࣀࢩࢩࡣ⟽ࢃ࡞ࡸᅖ࠸ࢃ࡞࡛ᤕ⋓ࡋࡓሙྜࠊ0 ṓࡀྠ᫬࡟」ᩘ㢌ᤕ⋓ࡉࢀࡓࡾࠊ0 ṓࡔࡅࡀᤕ⋓ࡉࢀࡿࡇ࡜࡛ᡂ⋇ಶయࡀᤕ⋓ࡋ࡟ࡃࡃ࡞ࡿࡇ࡜ࡀᣦ᦬ࡉࢀ࡚࠸ࡿ㸦Ụཱྀ
2003㸹す 2003㸧ࠋࡋ࠿ࡋࠊᮏ◊✲ࡢᑐ㇟ಶయ࡟ࡘ࠸࡚ࡣࠊ⟽ࢃ࡞ࡸᅖ࠸ࢃ࡞࡛ᤕ⋓ࡉࢀࡓಶయࡶࡃࡃࡾࢃ࡞࡛ᤕ⋓ࡉࢀࡓಶయࡶࠊ୍ᵝ࡟ 0 ṓ࡜ 1 ṓࡀ඲యࡢ༙ᩘ௨ୖࢆ༨ࡵ
ࡓࡇ࡜㸦㎷ ᮍⓎ⾲㸧࠿ࡽࠊ⌧᫬Ⅼ࡛ࡣࠊᤕ⋓᪉ἲ࡟ࡼࡿ኱ࡁ࡞ᖺ㱋ࡢ೫ࡾࡣぢࡽࢀ࡚࠸࡞࠸ࠋ࣮ࣚࣟࢵࣃࡢ࢖ࣀࢩࢩࡢሗ࿌࡛ࡣࠊᤕ⋓ᅽࡀ㧗࠸ಶయ⩌࡯࡝ᖺ㱋ᵓᡂࡀⱝ㱋࡟೫ࡿࡇ࡜ࡀ☜ㄆࡉࢀ࡚࠸ࡿ㸦Milkowski & Wójcik 1984; Fernandez-Llari et al. 2003㸧ࠋරᗜ┴ࡣࠊྂࡃ࠿ࡽ࢖ࣀࢩࢩࡢ㈨※ⓗ౯್ࡀᴟࡵ࡚㧗ࡃࠊ᫛࿴ 30 ᖺ௦㸦ᶓᒣ 2014㸧ࡼࡾ⊁⊟ᮇ࡟ࡣ㧗࠸ᤕ⋓ᅽࡀ࠿ࡅࡽࢀ࡚࠸ࡿ㸦ᶓᒣ 2014㸧ࠋࡋࡓࡀࡗ࡚ࠊᮏ◊✲࡟࠾࠸࡚0 ṓ࡜ 1 ṓࡢಶయࡢ๭ྜࡀከࡃ࡞ࡗࡓࡢࡣࠊරᗜ┴≉᭷ࡢ࢖ࣀࢩࢩ࡟ᑐࡍࡿ㧗࠸ᤕ⋓ᅽࡢᙳ㡪ࢆ཯ᫎࡋ࡚࠸ࡿ࡜⪃࠼ࡽࢀࡿࠋ ḟ࡟ࠊ㞤㞝ࡢ೫ࡾ࡟ࡘ࠸࡚ࡣࠊ௒ᅇࠊ0 ṓ࡜ 1 ṓࡣ࢜ࢫࡀከࡃࠊ2 ṓ௨㝆ࡣ࣓ࢫࡢ᪉ රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-5

ࡀከࡃ࡞ࡗࡓࠋ࡜ࡃ࡟ 5 ṓ௨㝆࡟ࡘ࠸࡚ࡣ࣓ࢫࡀ༨ࡵࡿ๭ྜࡀᴟ➃࡟ከࡃ࡞ࡾࠊ᭱㧗ᖺ㱋ࡶ࣓ࢫࡢ᪉ࡀ 3 ṓୖᅇࡗࡓ㸦ᅗ㸵㸧ࠋᅜෆࡢ௚ᆅᇦࡢ㱋ᵓᡂ㸦す 2003㸧ࡸ࣮ࣚࣟࢵࣃ
ࡢಶయ⩌ࡢ㱋ᵓᡂ㸦Fernandez-Llari et al. 2003㸧࡛ࡶྠᵝࡢഴྥࡀ࠶ࡿࡇ࡜࠿ࡽࠊ࢖
ࣀࢩࢩࡢ≉᭷ࡢ⩌ࢀᵓᡂࡸ⾜ື≉ᛶ࡟ࡼࡾࠊ㞤㞝࡛ᑑ࿨ࡀ␗࡞ࡾࠊ࣓ࢫࡢ᪉ࡀ࢜ࢫࡼࡾ
ࡶ㧗㱋ࡲ࡛⏕Ꮡࡋࡸࡍ࠸ࡇ࡜ࡀ⪃࠼ࡽࢀࡿࠋࡓࡔࡋࡇࢀࡽࡢሗ࿌ࡣࠊ඲࡚ᤕ⋓ಶయࡀᑐ㇟࡛࠶ࡿࡇ࡜࠿ࡽࠊ㞤㞝ࡸᖺ㱋࡟ࡼࡿᤕ⋓ࡢࡉࢀࡸࡍࡉ࡟ࡘ࠸࡚ࡶ௒ᚋࡉࡽ࡞ࡿ᳨ウࡀᚲせ࡜⪃࠼ࡽࢀࡿࠋㅰ㎡
 ᮏ◊✲ࢆᐇ᪋ࡍࡿ࡟࠶ࡓࡾࠊ2003 ᖺ࠿ࡽ 2013 ᖺࡢ 10 ᖺ㛫࡟රᗜ┴⊟཭఍ࡢከࡃࡢ
ᨭ㒊ࡢⓙࡉࡲ࡟ᶆᮏࢆࡈᥦ౪㡬ࡁࡲࡋࡓࠋⓙࡉࡲࡢከ኱࡞ࡿࡈ༠ຊ࡞ࡋ࡛ࡣᮏ◊✲ࡣ㐙⾜࡛ࡁࡲࡏࢇ࡛ࡋࡓࠋࡇࡢሙࢆ೉ࡾ࡚ࠊ῝ࡃᚚ♩⏦ࡋୖࡆࡲࡍࠋ

ᘬ⏝ᩥ⊩
Anezaki, T. 2009. Estimating age at death in Jomon Japanese wild boar (Sus scrofa

leucomystax) based on the timing of molar eruption in recent comparative samples.
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Apollonio, M., Putman, R., Grignolio S. and Bartos L. 2011. Hunting seasons in relation
to biological breeding seasons and the implications for the control or regulation of
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ῦ㔝⋞㸬2012㸬ᛶุูἲ࡜㱋ᰝᐃἲ㸬⩚ᒣఙ୍࣭୕ᾆៅᝅ࣭Ლග୍࣭㕥ᮌṇႹ⦅㸬㔝⏕ື≀⟶⌮Ɇ⌮ㄽ࡜ᢏ⾡Ɇ㸬pp247-255㸬ᩥỌᇽฟ∧㸪ᮾி㸬
Boitani, L. and Mattei L. 1992. Aging wild boar (Sus scrofa) by tooth eruption. In: Spitz,
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Ụཱྀ♸㍜㸬2003㸬࢖ࣀࢩࢩ࠿ࡽ⏣⏿ࢆᏲࡿ ࠾ࡶࡋࢁ࠸⏕ែ࡜࠿ࡋࡇ࠸㜵ࡂ᪉㸬pp.152㸬㸦♫㸧㎰ᮧ⁺ᮧᩥ໬༠఍㸪ᮾி㸬
Fernandez-Llario, P., Mateos-Quesada, P., Silvério, A., and Santos, P. 2003. Habitat
effects and shooting techniques on two wild boar (Sus scrofa) populations in Spain රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-5

and Portugal. Z Jagdwiss 49: 120-129.
ᯘⰋ༤࣭ す⏣㝯㞝࣭ ᮃ᭶බᏊ࣭ ℩⏣Ꮨⱱ㸬1977㸬᪥ᮏ⏘࢖ࣀࢩࢩࡢṑ∳࡟ࡼࡿᖺ௧࡜ᛶ
ࡢุᐃ. ᪥ᮏ⋖㓾Ꮵ㞯ㄅ㸪39 (2)㸹165-174㸬⎔ቃ┬⮬↛ಖㆤᒁ⏕≀ከᵝᛶࢭࣥࢱ࣮㸬2004㸬➨ 6 ᅇ⮬↛⎔ቃಖ඲ᇶ♏ㄪᰝ ✀ࡢከᵝᛶㄪᰝ ့ங㢮ศᕸㄪᰝሗ࿌᭩㸬pp.60-63㸬⎔ቃ┬⮬↛ಖㆤᒁ⏕≀ከᵝᛶࢭࣥࢱ࣮㸪ᒣ᲍┴㸬⎔ ቃ ┬ 㸬 2010 㸬 ⊁ ⊟ ཬ ࡧ ᭷ ᐖ ᤕ ⋓ ➼ ࡟ ࡼ ࡿ ୺ ࡞ 㫽 ⋇ ࡢ ᤕ ⋓ ᩘ . ⎔ ቃ ┬ 㸪 ᮾ ி 㸬
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ᯇᮏᓫ࣭ᆏ⏣ᏹᚿ࣭ᓊᮏᗣ㄃㸬2014㸬රᗜ┴࡟࠾ࡅࡿࢽ࣍ࣥ࢖ࣀࢩࢩࡢಶయ⩌ືែࡢ᥎ᐃ
㸦රᗜ┴ 2013 ᖺ㸧㸬ࠕරᗜ┴࡟࠾ࡅࡿࢽ࣍ࣥ࢖ࣀࢩࢩࡢ⟶⌮ࡢ⌧≧࡜ㄢ㢟㸬ࠖ㸪රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6 ྕ㸪pp.21-42㸪රᗜ┴᳃ᯘື≀◊✲ࢭࣥࢱ࣮㸬
Milkowski, L. and Wójcik, J. 1984. Structure of wild boar harvest in the Bialowieza
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㎰ᯘỈ⏘┬㸬2010㸬㔝⏕㫽⋇࡟ࡼࡿ㎰స≀⿕ᐖ≧ἣࡢ᥎⛣. ㎰ᯘỈ⏘┬㸪
http://www.maff.go.jp/j/seisan/tyozyu/higai/h_zyokyo2/h22/index.html.2012.6.14
accessed.
すಙ௓㸬2003㸬࢖ࣀࢩࢩࡢ⏕ᜥᐦᗘ࡜㱋ᵓᡂɆ㫽ྲྀ┴ඵ㢌㒆࡛ࡢㄪᰝ஦౛Ɇ㸬ᅜ㝿࢖ࣀࢩࢩࣇ࢛࣮࣒ࣛㅮ₇せ᪨㞟㸹79-80.
ᩪ⏣ᰤ㔛ዉ࣭ᶓᒣ┿ᘪ࣭୰ᮧᖾᏊ࣭᳃ග⏤ᶞ㸬2011㸬රᗜ┴࡟࠾࠸࡚ᤕ⋓ࡉࢀࡓࢶ࢟ࣀ࣡
ࢢ࣐ࡢᛶ࣭ᖺ㱋ᵓᡂࡢ≉ᚩ㸬ࠕරᗜ┴࡟࠾ࡅࡿࢶ࢟ࣀ࣡ࢢ࣐ࡢಖㆤ⟶⌮ࡢ⌧≧࡜ㄢ㢟㸬ࠖ㸪රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 3 ྕ㸪pp.94-101㸬රᗜ┴᳃ᯘື≀◊✲ࢭࣥࢱ࣮㸬ᆏ⏣ᏹᚿ࣭ᓊᮏᗣ㄃࣭㛵㤶⳯Ꮚ㸬2012㸬࢖ࣀࢩࢩࡢಶయ⩌ືែࡢ᥎ᐃ㸦රᗜ┴ᮏᕞ㒊 2011ᖺ㸧㸬රᗜ࣡࢖ࣝࢻࣛ࢖ࣇ࣏࣮ࣞࢺ 1 ྕ㸪pp.44-55㸬රᗜ┴᳃ᯘື≀◊✲ࢭࣥࢱ࣮㸬㡲⸨㸦ᒣ㊰㸧᫂Ꮚヂ㸬2001㸬ᛶุู࡜㱋ᰝᐃ㸬᪥ᮏ㔝⏕ື≀་Ꮫ఍࣭㔝⏕⏕≀ಖㆤᏛ఍┘ಟ㸪㕥ᮌṇႹ⦅ヂ㸪㔝⏕ື≀ࡢ◊✲࡜⟶⌮ᢏ⾡㸬ᩥỌᇽฟ∧㸪ᮾி㸪pp224-256㸬
㎷▱㤶࣭ᶓᒣ┿ᘪ㸬2014㸬රᗜ┴࡟࠾ࡅࡿࢽ࣍ࣥ࢖ࣀࢩࢩࡢᇶᮏⓗ⦾Ṫ≉ᛶ㸬ࠕරᗜ┴࡟࠾
ࡅࡿࢽ࣍ࣥ࢖ࣀࢩࢩࡢ⟶⌮ࡢ⌧≧࡜ㄢ㢟㸬ࠖ㸪රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6 ྕ㸪
pp.84-92㸪රᗜ┴᳃ᯘື≀◊✲ࢭࣥࢱ࣮㸬ᶓᒣ┿ᘪ㸬2014㸬රᗜ┴࡟࠾ࡅࡿࢽ࣍ࣥ࢖ࣀࢩࢩࡢಖㆤ⟶⌮ࡢ⌧≧࡜ㄢ㢟㸬ࠕරᗜ┴࡟࠾ࡅ
ࡿࢽ࣍ࣥ࢖ࣀࢩࢩࡢ⟶⌮ࡢ⌧≧࡜ㄢ㢟㸬ࠖ㸪රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6 ྕ㸪pp.1-8㸪රᗜ┴᳃ᯘື≀◊✲ࢭࣥࢱ࣮㸬 රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-6➨ 㸴 ❶            ࢽ࣍ࣥ࢖ࣀࢩࢩࡢዷፎ⋡⟬ฟ᪉ἲࡢ㛤Ⓨ㎷ ▱㤶 1࣭ᶓᒣ┿ᘪ 1,2
せ Ⅼ࣭ ࢖ࣀࢩࢩࡢዷፎ⋡ࢆ⟬ฟࡍࡿࡓࡵࡢ᪂ࡓ࡞ዷፎᣦᶆ࡜ࡋ࡚ࠊዷፎ㯤య㏥⦰≀࡟╔┠ࡋࠊ2004 ᖺ࠿ࡽ 2013 ᖺ࡟཰㞟ࡋࡓᤕ⋓ಶయࢆ⏝࠸࡚ࠊࡑࡢุูྍྰ࡜᭷⏝ᛶࢆ᳨ウࡋࡓࠋ࣭ ࢖ࣀࢩࢩ࡛ࡣཷ⫾࣭ฟ⏘᫬ᮇࡢࡤࡽࡘࡁࡀ኱ࡁ࠸ࡓࡵࠊ⫾Ꮚ☜ㄆࡢࡳ࡛ࡣዷፎ⋡ࢆ㐣ᑠ⟬ฟࡍࡿࡇ࡜ࡀ᫂ࡽ࠿࡜࡞ࡗࡓࠋ࣭ ዷፎ㯤య㏥⦰≀ࡣࠊࡑࡢ⏤᮶࠾ࡼࡧ᪂ᪧࡢุูࡀྍ⬟࡛࠶ࡾࠊ☜ᐇ࡟☜ㄆ࡛ࡁࡿᮇ㛫ࡀἪங☜ㄆࡼࡾࡶ㛗࠿ࡗࡓࡇ࡜࠿ࡽ㸪ฟ⏘ᚋಶయࡢᢕᥱ࡟᭷ຠ࡞ᣦᶆ࡛࠶ࡗ
ࡓࠋ࣭ ዷፎ⋡ࡣࠊ4㹼11 ᭶࡟⫾Ꮚ☜ㄆ࡟ࡼࡿዷፎಶయࡢ᳨ฟ࡜ዷፎ㯤య㏥⦰≀ TypeϨ☜ㄆ࡟ࡼࡿฟ⏘ᚋಶయࡢ᳨ฟࢆ⤌ࡳྜࢃࡏࡿࡇ࡜࡛㸪ࡼࡾṇ☜࡞ዷፎ⋡ࢆ⟬ฟࡍࡿࡇ࡜ࡀ࡛ࡁࡓࠋ

Key words 㸸ዷፎ⋡ࠊዷፎ㯤య㏥⦰≀ࠊ⫾Ꮚ☜ㄆࠊཷ⫾࣭ฟ⏘᫬ᮇࡢኚ␗ᖜࠊᤕ⋓ಶయศᯒ㸬ࡣࡌࡵ࡟
ື≀ࡢዷፎ⋡ࡣࠊಶయᩘࡢቑῶ࡟┤᥋ⓗ࡟㛵ࢃࡿࡓࡵࠊಶయ⩌ືែࢆᢕᥱࡍࡿୖ࡛ࡢ
ᇶ┙᝟ሗ࡛࠶ࡿ㸦Bailey 1984; Read 1990; Fonseca et al. 2004㸧ࠋࡲࡓዷፎ⋡ࡣࠊಶయ⩌ࡢᰤ㣴≧ែ࡟ᛂࡌ࡚ኚືࡍࡿࡓࡵࠊ⏕ᜥ⎔ቃࡢ㉁ⓗຎ໬ࡢᙳ㡪࡞࡝ࡑࡢ⩌ࢀࡢ೺඲ᛶࢆホ౯ࡍࡿୖ࡛ࡶ㔜せ࡞ᣦᶆ࡜࡞ࡿ㸦Matschke 1964; Kaji 1988㸧ࠋ㔝⏕ື≀࡛ࡣࠊ୍⯡ⓗ࡟஺ᑿࡸฟ⏘ࡢ┤᥋☜ㄆࡀ㞴ࡋ࠸ࡓࡵࠊዷፎ⋡ࡣࠊ⦾Ṫ᫬ᮇ࡟ᤕ⋓ࡉࢀࡓ㞤ࡢ⫾Ꮚ᭷↓ࡢ☜ㄆ࡟࡚⟬ฟࡉࢀࡿ㸦Ahmad et al. 1995; Gethöffer et al. 2007; Herrero et al.
2008㸧ࠋ෤╀࡞࡝Ꮨ⠇ⓗ࡞⾜ື࡟ࡼࡾዷፎಶయࡢᶆᮏࡀᚓ࡟ࡃ࠸ሙྜࠊ⫾┙⑞ࡸἪஙࡢ
᭷↓ࡢ☜ㄆࡀ௦⏝ࡉࢀ࡚࠸ࡿ㸦Tsubota et al. 1990; Asano et al. 2003㸧ࠋࡲࡓ㠀౵くⓗ࡟ᐇ᪋ࡍࡿሙྜ࡟ࡣࠊ⾑୰ࠊ⣅୰ࠊᒀ୰ࡢ࣍ࣝࣔࣥືែࡢ☜ㄆ࡞࡝ࡶ⏝࠸ࡽࢀ࡚࠸ࡿ
㸦Macchi et al. 2010㸧ࠋ ᅜෆ࡟⏕ᜥࡍࡿ኱ᆺ᭷㋟㢮ࡢ୰࡛ࠊࢽ࣍ࣥࢪ࢝㸦Cervus nippon㸧ࡸࢽ࣍ࣥ࢝ࣔࢩ࢝
㸦Capricornis crispus㸧࡛ࡣࠊ20 ᖺ௨ୖ๓࡟ዷፎ⋡ࡀ⟬ฟࡉࢀࠊ⌧ᅾࡶࡑࡢືែᢕᥱ

රᗜ┴᳃ᯘື≀◊✲ࢭࣥࢱ࣮࣭ රᗜ┴❧኱Ꮫ⮬↛࣭⎔ቃ⛉Ꮫ◊✲ᡤ රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-6

ࠋࡇࢀ࡟ᑐࡋ࡚ࠊ
ࡸᆅᇦẚ㍑ࡀ⾜ࢃࢀ࡚࠸ࡿ㸦Kita et al. 1987; Koizumi et al. 2009㸧ࢽ࣍ࣥ࢖ࣀࢩࢩ㸦Sus scrofa leucomystax㸹௨ୗ࢖ࣀࢩࢩ࡜⾲グ㸧ࡢዷፎ⋡ࡣࠊሗ࿌ࡀ
ᴟࡵ࡚ᑡ࡞࠸ࠋࡲࡓࠊ⛉Ꮫⓗ᰿ᣐ࡟ᇶ࡙࠸ࡓಙ㢗ᛶࡢ㧗࠸್࠿࡝࠺࠿ࡢ᳨ドࡶ༑ศ࡛࡞
࠸ࠋࡑࡢࡓࡵࠊ࢖ࣀࢩࢩࡢዷፎ⋡ࡣࠊಶయᩘࡢ᥎ᐃ࡟ࡶά⏝ࡉࢀ࡚࠸࡞࠸ࠋࡉࡽ࡟ࠊ⏕ᜥ⎔ቃࡢኚ໬ࡢᙳ㡪ࠊ࢖ࣀࣈࢱ࡜ࡢ஺㞧࡟ࡼࡿᙳ㡪࡞࡝ࡢ᳨ウࡶ⾜ࢃࢀ࡚࠸࡞࠸ࡢࡀ⌧≧࡛࠶ࡿࠋዷፎ⋡ࡢ⟬ฟࡀ㞴ࡋ࠸➨୍ࡢཎᅉࡣࠊ࢖ࣀࢩࢩ≉᭷ࡢ⊁⊟ᙧែ࡟ࡼࡾࠊዷፎಶయࡀᚓ
࡟ࡃ࠿ࡗࡓࡇ࡜࡟࠶ࡿ㸦Tsuji et al. 2013㸧ࠋ㣗⫗㈨※࡜ࡋ࡚ࡢ౯್ࡀᴟࡵ࡚㧗࠸࢖ࣀࢩࢩ࡛ࡣࠊࡑࡢᤕ⋓ᩘࡣ㈨※ⓗ౯್ࡢ㧗ࡲࡿ 11㹼12 ᭶࡟ᛴቑࡋࠊ౯್ࡀୗࡀࡿ 1 ᭶௨㝆࡟⃭ῶࡍࡿ࡜࠸࠺ࣃࢱ࣮ࣥࢆࡶࡘࠋࡋ࠿ࡋ᭱ࡶᤕ⋓ᩘࡢከ࠸᫬ᮇ࡟ࡣዷፎಶయࡀ࡯࡜ࢇ
࡝☜ㄆࡉࢀ࡞࠸ࠋࡘࡲࡾࠊᤕ⋓ᩘࡢከ࠸᫬ᮇࡣࠊዷፎ☜ㄆࡀ୙ྍ⬟࡞᫬ᮇ࡜࡞ࡿࡓࡵࠊ
ࡇࢀࡲ࡛⫾Ꮚ☜ㄆ࡟ࡼࡿዷፎ⋡ࡢ⟬ฟ࡟࠶ࡓࡗ࡚ࠊ⤫ィⓗศᯒ࡟㈨ࡍࡿ࡯࡝ࡢᶆᮏᩘࢆ
☜ಖ࡛ࡁ࡚ࡇ࡞࠿ࡗࡓࠋ➨஧ࡢཎᅉࡣࠊ࢖ࣀࢩࢩࡢዷፎ⋡ࢆⓗ☜࡟⟬ฟࡍࡿ᪉ἲࡀᩚ⌮ࡉࢀ࡚࠸࡞࠸ࡇ࡜࡛࠶ࡿࠋ㏆ᖺࠊ⫾Ꮚ☜ㄆ࡟࡚⟬ฟࡉࢀࡓ࢖ࣀࢩࢩࡢዷፎ⋡ࡀ 32%㸦ጜᓮ࣭ᆏᗞ 2010㸧࡜ࠊ࣮ࣚࣟࢵࣃࡢ࢖ࣀࢩࢩࡢሗ࿌㸦63㹼100%㸧㸦Fonseca et al. 2004;
Gethöffer et al. 2007; Fonseca et al. 2011㸧࡟ẚ࡭࡚ⴭࡋࡃప࠸⤖ᯝࡀᚓࡽࢀ࡚࠸ࡿࠋ
ᚑ᮶ࡢ⟬ฟ᪉ἲ࡛࠶ࡿ⫾Ꮚ☜ㄆ࡛ࡣࠊዷፎ⋡ࢆⓗ☜࡟⟬ฟ࡛ࡁ࡚࠸࡞࠸ྍ⬟ᛶࡀ᥎ࡉࢀࠊ᪥ᮏࡢ࢖ࣀࢩࢩࡢ≉ᛶ࡟࠶ࡗࡓ⊂⮬ࡢዷፎ⋡⟬ฟ᪉ἲࡀồࡵࡽࢀࡿࠋ ୖグࡢၥ㢟ࡢゎỴ࡟ྥࡅ࡚ࠊᮏ◊✲࡛ࡣࠊዷፎಶయࢆ⫾Ꮚ☜ㄆ࡟᳨࡚ฟࡍࡿࡇ࡜࡟ຍ
࠼ࠊฟ⏘ᚋಶయࢆ㐣ཤࡢዷፎࡢ᭷↓ࢆ☜ㄆࡍࡿࡇ࡜᳨࡛ฟࡍࡿࡇ࡜࡜ࡋࠊ᪂ࡓ࡞ዷፎ⋡⟬ฟᣦᶆ࡜ࡋ࡚ࠊዷፎ㯤య㏥⦰≀࡟╔┠ࡋࡓࠋࡇࢀࡣࠊዷፎ⥔ᣢ࡟ᶵ⬟ࡋ࡚࠸ࡓዷፎ㯤యࡀࠊฟ⏘ᚋ༸ᕢෆ࡟ᇙἐࡋࡓࡶࡢ࡛࠶ࡿ㸦㧗ᶫ࡯࠿ 1999㸧ࠋฟ⏘ᚋࡶ୍ᐃࡢᮇ㛫ṧᏑࡍࡿࡓࡵࠊዷፎ㯤య㏥⦰≀ࡢ᭷↓ࢆ☜ㄆࡍࡿࡇ࡜࡛ࠊฟ⏘ᚋࡢಶయࡸᤵஙࢆ⤊࠼ࡓಶయ࡛ࡶ㐣ཤࡢዷፎṔࢆᢕᥱ࡛ࡁࡿ 㸦Yamashita 1962; Kita et al. 1995㸧ࠋࡲࡓዷፎ㯤య㏥
⦰≀ࡣࠊ࡯࡜ࢇ࡝඲࡚ࡢ့ங㢮ື≀࡛సࡽࢀࡿࡓࡵࠊ⫾┙⑞ࡀṧࡽ࡞࠸✀࡟ࡶ㐺⏝ྍ⬟࡛࠶ࡿࠋᅜෆ࡛ࡣࢽ࣍ࣥࢪ࢝ࡸࢽ࣍ࣥ࢝ࣔࢩ࢝㸦Sugimura et al. 1984; Kita et al.
1995㸧ࠊᾏእ࡛ࡣ᭷㋟㢮ࡔࡅ࡛࡞ࡃ㪒㢮㸦Born 1982㸧࡟࠾࠸࡚ࡶࠊዷፎ㯤య㏥⦰≀ࢆά⏝ࡋ࡚ࠊዷፎᒚṔࡀᢕᥱࡉࢀ࡚࠸ࡿࠋࡓࡔࡋ࢖ࣀࢩࢩ࡛ࡣࠊࡇࢀࡲ࡛࡟ዷፎ㯤య㏥⦰≀ࢆᣦᶆ࡜ࡋ࡚ዷፎṔࢆุᐃࡋࡓሗ࿌ࡣ࡞࠸ࠋ
௨ୖࡢ⫼ᬒࡼࡾࠊᮏ◊✲࡛ࡣࠊ࢖ࣀࢩࢩࡢዷፎ⋡ࢆ㐺ษ࡟⟬ฟࡍࡿࡇ࡜ࢆ┠ⓗ࡜ࡋ࡚ࠊ⫾Ꮚ࠾ࡼࡧዷፎ㯤య㏥⦰≀ࢆే⏝ࡍࡿ᪂ࡓ࡞ዷፎ⋡⟬ฟ᪉ἲࢆ᳨ウࡋࡓࠋࡲࡎࠊዷፎ⋡ࡢ⟬ฟ࡟㐺ษ࡞᫬ᮇࡢᢕᥱࠊḟ࡟ዷፎ㯤య㏥⦰≀ࡢุูࡢྍྰࢆ᳨ウࡋࡓࠋࡑࡢᚋࠊዷ
ፎ㯤య㏥⦰≀ࡢ᭷⏝ᛶࢆホ౯ࡍࡿࡓࡵ࡟ࠊྠࡌࡃฟ⏘ᚋಶయࡢᣦᶆ࡜࡞ࡿἪங࡜ࡢẚ㍑
ࢆ⾜ࡗࡓࠋ

 

රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-6

ᮦᩱ࡜᪉ἲᮦᩱ ᮏ◊✲࡛ࡣࠊ㐺ษ࡞ዷፎ⋡⟬ฟ᪉ἲࢆ᳨ウࡍࡿࡓࡵ࡟ࠊᛶᡂ⇍࡟㐩ࡋࡓ⦾Ṫྍ⬟ಶయࢆศᯒࡍࡿᚲせࡀ࠶ࡗࡓࠋරᗜ┴ࡢ࢖ࣀࢩࢩࡢ࡯࡜ࢇ࡝ࡣࠊ1 ṓࡢ 12 ᭶௨㝆࡟ᛶᡂ⇍࡟㐩ࡍࡿ㸦㎷ 2013㸧ࠋࡋࡓࡀࡗ࡚ᮏ◊✲࡛ࡣࠊ⦾Ṫྍ⬟ಶయ࡜ࡋ࡚ࠊ12㹼4 ᭶࡟ᤕ⋓ࡉ
ࢀࡓ 1 ṓ࡜ᖺ㛫㏻ࡌ࡚ᤕ⋓ࡉࢀࡓ 2 ṓ௨ୖࡢ࣓ࢫࢆᑐ㇟࡜ࡋࡓࠋ2004 ᖺ࠿ࡽ 2011 ᖺ࡟⊁⊟࠾ࡼࡧ᭷ᐖᤕ⋓࡟࡚ᤕ⋓ࡉࢀࡓ⦾Ṫྍ⬟ಶయ 142 㢌ࢆゎ๗ࡋࠊ༸ᕢ࡜⬇࠾ࡼࡧ⫾Ꮚࢆ᥇ྲྀࡋࡓࠋ
ዷፎ⋡⟬ฟᣦᶆࡢほᐹᚑ᮶ࡢᣦᶆ࡛࠶ࡿ⬇࠾ࡼࡧ⫾Ꮚࡢ᭷↓ࡣࠊ⫗║ほᐹ࡟࡚⾜ࡗࡓࠋࡇࢀࡽࡀ☜ㄆࡉࢀࡓሙྜࡣࠕ⫾Ꮚ+ࠖࠊ☜ㄆࡉࢀ࡞࠿ࡗࡓሙྜࡣࠕ⫾Ꮚ㸫ࠖ࡜༊ูࡋࡓࠋ
ዷፎ㯤య㏥⦰≀ࡣ⤌⧊ほᐹ࡟ࡼࡾุูࡋࡓࠋ⤌⧊ᶆᮏࡢస〇ᡭ㡰࡜ࡋ࡚ࠊࡲࡎ༸ᕢࢆ
10%࣐࣍ࣝࣜࣥᾮ࡛ᅛᐃᚋࠊ⣙ 4mm 㛫㝸࡟ࢫࣛ࢖ࢫࡋࡓࠋࡑࡢᶆᮏࢆ⬺Ỉ㏱ᚭᚋࣃࣛ
ࣇ࢕ࣥໟᇙࡋࠊ100ȣm 㛫㝸࡛ཌࡉ 4ȣm ࡟ⷧษࡋࡓࠋⷧษࡋࡓ⤌⧊ษ∦࡟ࡣࠊ࡬࣐ࢺ
࢟ࢩ࣭࢚ࣜࣥ࢜ࢪࣥᰁⰍࠊᙎᛶ⥺⥔ࢆᰁࡵศࡅࡿࡓࡵࡢ࢚ࣛࢫࢳ࣭࢝࣡ࣥࢠ̿ࢯࣥᰁⰍࠊ
⭺ཎ⥺⥔ࢆᰁࡵศࡅࡿࡓࡵࡢ࣐ࢵࢯ࣭ࣥࢺࣜࢡ࣮࣒ࣟᰁⰍࢆ⾜ࡗࡓࠋḟ࡟ዷፎ㯤య㏥⦰≀ࡢุู࡟ࡣࠊ࢖ࣀࢩࢩࡢள✀ࡢ୍ࡘࡢࣈࢱ㸦 Sus scrofa domesticus 㸧ࡢ༸ᕢᡤぢ㸦Yamashita 1959; Yamashita 1960a; Yamashita 1960b; Yamashita 1962㸧ࢆᇶ‽࡜ࡋࡓࠋࣈࢱ࡛ࡣࠊዷፎ㯤య㏥⦰≀ࡣ㏥⾜ẁ㝵࡟ᛂࡌ࡚ TypeϨ࠿ࡽϫࡲ࡛ 4 ༊ศࡉࢀ࡚
࠾ࡾࠊ᭱㛗ṧᏑᮇ㛫ࡣ 3 ᖺ࡛࠶ࡿࠋࡇࡢ୰࡛᭱ึࡢ㏥⾜ẁ㝵 TypeϨࡀほᐹࡉࢀࡿࡢࡣࠊ
ฟ⏘ᚋ⣙ 1 ᖺ㛫࡛࠶ࡿࠋࡇࢀࡼࡾᮏ◊✲࡛ࡣࠊ㐣ཤ 1 ᖺ௨ෆࡢ⦾Ṫ᫬ᮇ࡟ዷፎࡋࡓ࠿࡝
࠺࠿ࢆᢕᥱࡍࡿࡓࡵ࡟ TypeϨࡢ᭷↓ࢆ☜ㄆࡋࠊࡑࡢ᭷↓࡟࡚ࠕ㏥⦰≀+ࠖࠊ
ࠕ㏥⦰≀㸫ࠖ
࡜༊ูࡋࡓࠋ
ẚ㍑ᣦᶆ࡜ࡋࡓἪஙࡢ᭷↓ࡣಶయࡢᤕ⋓⌧ሙ࡟࡚㝶᫬⾜࠸ࠊࡑࡢ☜ㄆྍྰ࡟࡚ࠕἪங
+ࠖࠊࠕἪங㸫ࠖ࡜༊ูࡋࡓࠋ
ዷፎ⋡⟬ฟྍ⬟᫬ᮇࡢᢕᥱ
ዷፎ⋡ࢆⓗ☜࡟⟬ฟ࡛ࡁࡿ᫬ᮇࢆ᳨ウࡍࡿࡓࡵ࡟ࠊࡲࡎࠊዷፎಶయ࠿ࡽᚓࡽࢀࡓ⫾Ꮚࡢయ㔜࠿ࡽ⫾㱋ࢆ᥎ᐃࡋࠊཷ⫾᫬ᮇ࡜ฟ⏘᫬ᮇࢆ⟬ฟࡋࡓࠋ⫾㱋᥎ᐃ࡟ࡣࠊ࢖ࣀࢩࢩࡢ
ள✀ࡢ୍ࡘ࡛࠶ࡿ Iberian wild boar㸦Sus scrofa baeticus㸧ࡢ㣫⫱ಶయ࠿ࡽసᡂࡉࢀࡓ
᥎ᐃᘧࢆᘬ⏝ࡋࡓ㸦Vericad 1983㸧ࠋࡓࡔࡋࠊዷፎᮇ㛫࡜ฟ⏕య㔜ࢆ᪥ᮏࡢ࢖ࣀࢩࢩࡢ
್࡜ࡍࡿࡓࡵ࡟᥎ᐃᘧࢆᨵኚࡋࡓࠋᘧࡢኚ᥮࡟ᙜࡓࡗ࡚ࠊዷፎᮇ㛫ࡣ᪥ᮏࡢ࢖ࣀࢩࢩࡢ
㣫⫱ಶయࡢሗ࿌㸦වග࡯࠿ 1988㸧ࡼࡾ 117.1s0.85 ᪥㸦N=7㸧ࠊฟ⏕య㔜ࡣ 804g㸦Tsuji

et. al 2013㸧࡜ࡋࡓࠋ රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-6

Iberian wild boar ࡢ⫾㱋᥎ᐃᘧ㸸T㸻Ps1/3㸭0.097+24.1ᮏ◊✲࡛⏝࠸ࡓ⫾㱋᥎ᐃᘧ㸸T㸻Ps1/3㸭0.097+21.1379
T ࡀ୍⭡ෆࡢᖹᆒ⫾㱋ࠊPs ࡀ୍⭡ෆࡢᖹᆒ⫾Ꮚయ㔜ࢆ♧ࡍࠋ⫾㱋᥎ᐃ࡟࡚⟬ฟࡉࢀࡓཷ⫾᫬ᮇ࡜ฟ⏘᫬ᮇࢆࡶ࡜࡟ࠊ᭱ࡶ᪩࠸ཷ⫾᭶㹼᭱ࡶ㐜࠸ฟ⏘᭶ࢆዷፎᮇࠊࡑࢀ௨እࡢ᭶ࢆ㠀ዷፎᮇ࡜༊ูࡋࡓࠋ
ḟ࡟༸ᕢ୰ࡢ㯤యࡢ᭷↓ࢆ⫗║ほᐹ࡟࡚☜ㄆࡋࡓࠋ㯤యࡣࠊ௚ࡢື≀࡟࠾࠸࡚ࠊ⬇ࡀ⫗║࡛☜ㄆ࡛ࡁ࡞࠸ዷፎึᮇ࡟࠾ࡅࡿዷፎᣦᶆ࡜ࡋ࡚⏝࠸ࡽࢀ࡚࠸ࡿ㸦Born 1982;
Gethöffer et al. 2007㸧ࠋࡇࡢ㝿ࠊ஺ᑿ᤼༸࡛ࡣ࡞࠸ື≀࡛ࡣࠊዷፎ㯤య࡜Ⓨ᝟㯤యࡢุูࡀᚲせ࡜࡞ࡿ㸦Mansell 1971; Kita et al. 1994㸧ࠋዷፎࡋ࡚࠸ࡿಶయࡣዷፎ㯤యࢆಖ᭷ࡍࡿࡀࠊዷፎࡋ࡚࠾ࡽࡎⓎ᝟ࡢࡳࡢಶయࡣⓎ᝟㯤యࢆಖ᭷ࡋ࡚࠸ࡿࠋࡋࡓࡀࡗ࡚ࠊࡇ
ࢀࡽࢆุูࡋ࡞࠸ሙྜࠊⓎ᝟ಶయࡶࠕዷፎ࠶ࡾࠖ࡜ุᐃࡋ࡚ࡋࡲ࠸ࠊዷፎ⋡ࢆ㐣኱⟬ฟࡍࡿࠋ࢖ࣀࢩࢩ࡛ࡣࠊ⌧᫬Ⅼ࡛ࡣࠊࡇࡢุูࡀ୙ྍ⬟࡛࠶ࡾࠊ͆⬇ࡣ☜ㄆ࡛ࡁ࡞࠸ࡀ㯤యࢆಖ᭷ࡍࡿ͇࡜࠸࠺࣓ࢫࡢዷፎุᐃࡀ࡛ࡁ࡞࠸㸦Tsuji et al. 2013㸧ࠋࡋࡓࡀࡗ࡚ᮏ◊✲࡛ࡣࠊࡇࡢࡼ࠺࡞ዷፎุᐃࡀ࡛ࡁ࡞࠸ಶయࡀᏑᅾࡍࡿ᫬ᮇࡣࠊዷፎ⋡⟬ฟ࡟୙㐺ษ࡞ᮇ㛫࡜ࡋ࡚㝖እࡋࡓࠋዷፎ⋡⟬ฟ
ዷፎ⋡⟬ฟ࡟୙㐺ษ࡞ᮇ㛫ࢆ㝖እࡋࡓᚋࠊዷፎᮇ࡜㠀ዷፎᮇࡢዷፎ⋡ࢆ⟬ฟࡋࡓࠋዷ
ፎ⋡⟬ฟ࡟࠶ࡓࡗ࡚ࡣࠊ⫾Ꮚ☜ㄆࡢࡳࡢ᪉ἲ㸦ጜᓮ࣭ᆏᗞ 2010㸧࡜ᮏ◊✲࡛᪂ࡓ࡟᳨
ウࡍࡿ⫾Ꮚ࡜㯤య㏥⦰≀ࠊ⫾Ꮚ࡜Ἢஙࡢ☜ㄆࢆ⤌ࡳྜࢃࡏࡓ᪉ἲࢆ⏝࠸ࠊ⟬ฟ᪉ἲࡢ㐺ṇ࡜ྛᣦᶆࡢ᭷⏝ᛶࢆホ౯ࡋࡓࠋ⤫ィฎ⌮ཷ⫾࡜ฟ⏘᫬ᮇࡢእࢀ್ࡢ᳨ฟ࡟ࡣࠊSmirnov-Grubbs ᳨ᐃࢆ⏝࠸ࠊ༴㝤⋡ࡣ 0.01%
࡜ࡋࡓࠋ⤖ᯝዷፎ⋡⟬ฟ࡟㐺ࡋࡓ᫬ᮇ
ཷ⫾᫬ᮇࡣ 12 ᭶ୖ᪪࠿ࡽ 3 ᭶୰᪪ࡢᮇ㛫࡟࠶ࡾ 1 ᭶ୗ᪪ࢆࣆ࣮ࢡ࡜ࡋࠊ1 ౛ࡢࡳ 5᭶ୖ᪪࡜⟬ฟࡉࢀࡓࠋฟ⏘᫬ᮇࡣ 4 ᭶ୖ᪪࠿ࡽ 7 ᭶୰᪪ࡢᮇ㛫࡟࠶ࡾ 6 ᭶ୖ᪪ࢆࣆ࣮
ࢡ࡜ࡋࠊ1 ౛ࡢࡳ 9 ᭶୰᪪࡜⟬ฟࡉࢀࡓࠋ 5 ᭶୰᪪࡟ཷ⫾ࡋࠊ9 ᭶୰᪪࡟ฟ⏘ࡍࡿ࡜᥎ᐃࡉࢀࡓ 1 ౛ࡣࠊ እࢀ್࡜ࡋุ࡚ᐃࡉࢀࡓ㸦p < 0.01㸧ࠋරᗜ┴ࡢ࢖ࣀࢩࢩಶయ⩌ࡢ⦾Ṫࢧ࢖ࢡࣝࡣ 12 ᭶࠿ࡽ⩣ᖺࡢ 11 ᭶ࢆ 1 ࢧ࢖ࢡࣝ࡜ࡋࠊዷፎᮇࡣ 12㹼7 ᭶ࠊ㠀ዷፎᮇࡣ 8㹼11 ᭶࡛࠶ࡗࡓ㸦ᅗ㸯㸧ࠋ රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-6

ᅗ ࢖ࣀࢩࢩࡢཷ⫾᫬ᮇ࡜ฟ⏘᫬ᮇ⬇ࡸ⫾Ꮚࠊ㯤యࡀ☜ㄆࡉࢀࡓ᫬ᮇ࡟ࡣᏘ⠇ᛶࡀ࠶ࡗࡓࠋ㯤యಖ᭷ಶయࡣ 12 ᭶࠿ࡽ☜ㄆࡉࢀࡓࠋࡓࡔࡋ 12㹼3 ᭶࡟ࡣࠊ㯤యࢆಖ᭷ࡍࡿࡀࠊ⬇ࡀ☜ㄆ࡛ࡁ࡞࠸㸦⫾Ꮚ㸫㺃㯤య+㸧ࠊࡘࡲࡾⓎ᝟୰ࡶࡋࡃࡣዷፎึᮇࡢ࣓ࢫࡀ 21 㢌☜ㄆࡉࢀࡓࠋ2㹼6 ᭶࡜ 8 ᭶࡟ࡣࠊ㯤య࡜⬇ࡸ⫾Ꮚࢆಖ᭷ࡋࡓዷፎ࣓ࢫ㸦⫾Ꮚ+㺃㯤య+㸧ࡀ 38 㢌☜ㄆࡉࢀࡓࠋ12㹼3 ᭶࡜ 5
㹼11 ᭶࡟ࡣࠊ㯤యࡶ⬇ࡸ⫾Ꮚࡶ☜ㄆࡉࢀ࡞࠿ࡗࡓ࣓ࢫ㸦⫾Ꮚ㸫㺃㯤య㸫㸧ࡀィ 83 㢌☜ㄆࡉࢀࡓࠋ12 ᭶࠿ࡽ⩣ᖺࡢ 11 ᭶ࡢ୍⦾Ṫࢧ࢖ࢡࣝ୰࡛ࠊ12㹼3 ᭶ࡣዷፎุᐃࡀ࡛ࡁ࡞
࠸ಶయࡀᏑᅾࡋࡓࡇ࡜࠿ࡽࠊዷፎ⋡⟬ฟ࡟୙㐺ษ࡞ᮇ㛫࡜ุ᩿ࡋࡓ㸦ᅗ㸰㸧ࠋᅗ ⫾Ꮚ࡜㯤యࡀ☜ㄆࡉࢀࡓ᫬ᮇ රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-6ዷፎ㯤య㏥⦰≀ࡢุูࡢྍྰ
༸ᕢࡢ⤌⧊ほᐹ࡟࡚ࠊࣈࢱࡢሗ࿌࡜ྠᵝࡢ⤌⧊ᡤぢ࡜࡞ࡿዷፎ㯤య㏥⦰≀ࡀほᐹࡉࢀࡓࠋ1 ࡘࡢ༸ᕢ࠿ࡽᵝࠎ࡞ࢱ࢖ࣉࡢዷፎ㯤య㏥⦰≀ࡀ☜ㄆࡉࢀࡓಶయࡶ࠶ࢀࡤࠊ1 ࢱ࢖ࣉࡢࡳ☜ㄆࡉࢀࡓಶయࡶᏑᅾࡋࡓࠋ☜ㄆࡉࢀࡓዷፎ㯤య㏥⦰≀ࡢ୰࡛ࠊ㏥⦰≀ࡢෆ㒊࡟
㏥⾜㯤య⣽⬊ࡀᏑᅾࡋࠊ㏥⦰≀࿘ᅖࡸ⾑⟶ቨ࡟ᙎᛶ⥺⥔ࡢ㞟✚ࡀほᐹࡉࢀࡓࡶࡢࢆ
Type Ϩ࡜ุ᩿ࡋࡓ㸦ᅗ㸱. A ࡜ B㸧ࠋࡇࢀ࡟ᑐࡋ࡚㏥⾜㯤య⣽⬊ࡀᾘኻࡋࠊ㏥⦰≀ෆࡢᅗ ዷፎ㯤య㏥⦰≀ࡢุู $ ࡜ % ࡣࠊዷፎ㯤య㏥⦰≀ 7\SHϨࠊ& ࡜ ' ࡣ 7\SHϨࡼࡾࡶྂ࠸ዷፎ㯤య㏥⦰≀ࠊ( ࡜ ) ࡣⓎ᝟㯤యࡢ㏥⦰≀ࡢ⤌⧊ീࠋ$㹼' ࡢዷፎ㯤య㏥⦰≀࠿ࡽࡣࠊ⾑⟶ቨࡸࡑࡢ࿘ᅖ࡟ᙎᛶ⥺⥔ࡢ㞟✚ࡀ☜ㄆࡉࢀࡓ㸦▮༳㸧ࠋዷፎ㯤య㏥⦰≀ 7\SHϨ࡟ࡣ㏥⾜ࡋࡓ㯤య⣽⬊ࡀ☜ㄆࡉࢀࡓ㸦▮㢌㸧ࠋ࢚ࣛࢫࢳ࣭࣮࢝࣡ࣥࢠࣥࢯࣥᰁⰍࠋ6FDOHEDU ȣP $&( ࠊȣP %') ࠋ රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-6

⾑⟶ࡢ኱ࡁࡉࡸᩘࡀቑຍࡋࡓࡶࡢࡣࠊTypeϩ௨㝆ࡢዷፎ㯤య㏥⦰≀ࡔ࡜ุ᩿ࡉࢀࡓ㸦ᅗ
㸱. C ࡜ D㸧ࠋࡉࡽ࡟ᙎᛶ⥺⥔ࡢ㞟✚ࡀほᐹࡉࢀࡎࠊ㏥⦰≀ෆࡀࣄ࢔ࣜࣥ≀㉁࡛༨ࡵࡽࢀ
࡚࠸ࡓࡶࡢࡣࠊⓎ᝟㯤య㏥⦰≀࡜ุ᩿ࡉࢀࡓ㸦ᅗ㸱. E ࡜ F㸧ࠋ௨ୖࡼࡾ㯤య㏥⦰≀ TypeϨࡢ㆑ูࡣྍ⬟࡛࠶ࡗࡓ.
Ἢங࡜ዷፎ㯤య㏥⦰≀ࡢ᭷⏝ᛶࡢẚ㍑
ฟ⏘ᚋࡢಶయࢆ᳨ฟࡍࡿᣦᶆ࡜ࡋ࡚ࠊዷፎ㯤య㏥⦰≀ TypeϨ࡜Ἢஙࡢ᭷⏝ᛶࡢẚ㍑
ࢆ⾜ࡗࡓ㸦⾲ 1㸧
ࠋ4㹼11 ᭶࡟⬇ࡸ⫾Ꮚࡀ☜ㄆࡉࢀ࡞࠿ࡗࡓ࣓ࢫ 45 㢌୰ࠊዷፎ㯤య㏥⦰
≀ TypeϨࡣ 5㹼11 ᭶࡟ 40 㢌࠿ࡽ☜ㄆࡉࢀࡓࠋࡇࢀ࡟ᑐࡋ࡚Ἢஙࡣ 5㹼9 ᭶࡜ 11 ᭶࡟
15 㢌࠿ࡽ☜ㄆࡉࢀࡓࠋ୧ᣦᶆࡣ 5㹼6 ᭶ࡣྠᩘࡀ☜ㄆࡉࢀࡓࡀࠊ7 ᭶௨㝆ࡣἪஙࡢ☜ㄆᩘࡀῶᑡࡋࡓࠋࡲࡓἪஙࡀ☜ㄆࡉࢀࡓࡍ࡭࡚ࡢಶయ࠿ࡽ㯤య㏥⦰≀ typeϨࡀ☜ㄆࡉࢀࡓࠋ௨ୖࡼࡾࠊฟ⏘ᚋࡢಶయࢆ᳨ฟࡍࡿᣦᶆ࡜ࡋ࡚ࡣࠊዷፎ㯤య㏥⦰≀ TypeϨࡢ᪉ࡀἪஙࡼࡾࡶ☜ㄆྍ⬟ᮇ㛫ࡀ㛗࠸ࡇ࡜ࡀ᫂ࡽ࠿࡜࡞ࡗࡓࠋ
⾲ 㯤య㏥⦰≀࡜Ἢஙࡢ☜ㄆᩘࡢẚ㍑
 ᭶

 ᭶

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

␗࡞ࡿᣦᶆࡢ⤌ࡳྜࢃࡏ࡟࡚⟬ฟࡉࢀࡓዷፎ⋡ࡢẚ㍑
 ዷፎ⋡ࡢ⟬ฟࡣࠊዷፎᮇ㸦4㹼7 ᭶㸧࡜㠀ዷፎᮇ㸦8㹼11 ᭶㸧ࡢᶆᮏࢆ⏝࠸࡚ࠊ᪉ἲ
࡛♧ࡋࡓ 3 ࣃࢱ࣮ࣥࡢᣦᶆࢆ⤌ࡳྜࢃࡏ࡚⾜ࡗࡓࠋࡲࡎ⫾Ꮚ࡜ዷፎ㯤య㏥⦰≀ TypeϨ☜ㄆࡢే⏝࡟ࡼࡿዷፎ⋡ࡣࠊዷፎᮇࡀ 93.94㸣ࠊ㠀ዷፎᮇࡀ 90.91㸣࡜࡞ࡾࠊ඲య࡜ࡋ࡚ 92.42㸣࡜⟬ฟࡉࢀࡓ㸦⾲㸰. 㹟㸧ࠋḟ࡟⫾Ꮚ☜
ㄆࡢࡳ࡛ࡣࠊዷፎᮇࡀ 60.61㸣ࠊ㠀ዷፎᮇࡀ 3.03㸣࡜࡞ࡾࠊ඲య࡜ࡋ࡚ 31.82㸣࡜⟬ฟࡉࢀࡓ㸦⾲㸰. 㹠㸧ࠋ᭱ᚋ࡟⫾Ꮚ࡜Ἢங☜ㄆࡢే⏝࡛ࡣࠊዷፎᮇࡀ 87.88㸣ࠊ㠀ዷፎᮇࡀ21.21㸣࡜࡞ࡾࠊ඲య࡜ࡋ࡚ࠊ54.55%࡜⟬ฟࡉࢀࡓ㸦⾲㸰. 㹡㸧ࠋ௨ୖࡼࡾࠊዷፎᮇ࡜㠀ዷፎᮇ࡟ฟ⏘ᚋಶయࢆぢ㏨ࡋ࡞ࡃ᳨ฟࡋࡓ⫾Ꮚ࡜ዷፎ㯤య㏥⦰≀ TypeϨࡢే⏝࡟ࡼࡿዷፎ⋡ࡀ᭱ࡶ㧗ࡃ⟬ฟࡉࢀࡓࠋ

⪃ᐹዷፎ㯤య㏥⦰≀ࡢุู࡜ࡑࡢ᭷⏝ᛶ ᮏ◊✲ࡣࠊ࢖ࣀࢩࢩࡢ༸ᕢࡢ⤌⧊ほᐹࢆࡍࡿࡇ࡜࡛ࠊዷፎ㯤య㏥⦰≀ࡢุูࡀྍ⬟࡛࠶ࡿࡇ࡜ࢆ᫂ࡽ࠿࡟ࡋࡓ㸦ᅗ㸱㸧ࠋ୺࡞ุู࣏࢖ࣥࢺࢆࡲ࡜ࡵࡿ࡜ࠊ㐣ཤ 1 ᖺ௨ෆ࡟ᙧ
ᡂࡉࢀࡓዷፎ㯤య㏥⦰≀㸦TypeϨ㸧࡜ࡑࢀࡼࡾࡶ๓࡟ᙧᡂࡉࢀࡓྂ࠸ዷፎ㯤య㏥⦰≀㸦Typeϩ㹼ϫ㸧ࡢ᪂ᪧุᐃ࡟ࡣࠊ⤌⧊ษ∦࡟࢚ࣛࢫࢳ࣭࢝࣡ࣥࢠ࣮ࢯࣥᰁⰍ᪋ࡋࠊኚ
ᛶ㯤య⣽⬊ࡸᙎᛶ⥺⥔ࡢ㞟✚ࡢ᭷↓ࢆ☜ㄆࡍࡿᡭἲࡀ᭷ຠ࡛࠶ࡗࡓࠋⓎ᝟㯤య㏥⦰≀࡜ රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-6

⾲ ྛዷፎᣦᶆ࡜ࡑࡢ⤌ࡳྜࢃࡏ࡟ࡼࡗ࡚⟬ฟࡉࢀࡓዷፎ⋡㸦㹟㸧⫾Ꮚ࡜㏥⦰≀ࢆᣦᶆ࡜ࡋࡓሙྜ
⫾ᏊѸ⫾ᏊѸ

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1⫾Ꮚ ᭶㸦ዷፎᮇ㸧 ᭶㸦㠀ዷፎᮇ㸧ྜィ㸦㹠㸧⫾Ꮚࡢࡳࢆᣦᶆ࡜ࡋࡓሙྜ


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1

⫾Ꮚ

 ᭶㸦ዷፎᮇ㸧

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

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ࡢุู࡟ࡣࠊ࣐ࢵࢯ࣭ࣥࢺࣜࢡ࣮࣒ࣟᰁⰍ࡟࡚ࠊࣄ࢔ࣜࣥ≀㉁ࡢ᭷↓ࢆ☜ㄆࡍࡿࡇ࡜࡛
ྍ⬟࡜࡞ࡗࡓࠋ
ḟ࡟ᮏ◊✲࡛ࡣࠊዷፎ㯤య㏥⦰≀ TypeϨࡀዷፎ⋡ࡢ⟬ฟᣦᶆ࡜ࡋ࡚᭷⏝࡛࠶ࡿ࠿࡝࠺࠿ࢆホ౯ࡍࡿࡓࡵ࡟ࠊἪங☜ㄆ㸦Fonseca et al. 2004㸧࡜ẚ㍑ࡋࡓࠋ୧⪅ࡢ☜ㄆᩘࡢ㐪࠸ࡣ 7㹼11 ᭶࡟ㄆࡵࡽࢀࠊἪங☜ㄆࡢ᪉ࡀᑡ࡞࠿ࡗࡓ㸦⾲㸯㸧ࠋ࢖ࣀࢩࢩࡢᤵஙᮇ㛫ࡣ⣙ 3 ࠿᭶㛫࡛࠶ࡿࡇ࡜࠿ࡽ㸦Ụཱྀ 2003㸧ࠊරᗜ┴ࡢ࢖ࣀࢩࢩ࡛ࡣࠊ7 ᭶௨㝆࡟ࡣᤵஙࢆ⤊࠼ࡓಶయࡀᏑᅾࡋࡓ࡜⪃࠼ࡽࢀࡿࠋࡋࡓࡀࡗ࡚ࠊ7 ᭶௨㝆ࡶἪங☜ㄆࢆࡶࡗ࡚ዷፎ
⋡ࢆ⟬ฟࡍࡿ࡜ࠊฟ⏘ᚋಶయࡢぢⴠ࡜ࡋࡀⓎ⏕ࡋࠊዷፎ⋡ࢆ㐣ᑠ⟬ฟࡍࡿ࡜ุ᩿ࡉࢀࡓ㸦⾲㸰㹡㸧ࠋ㐣ᑡ⟬ฟࢆ㑊ࡅࡿ࡟ࡣࠊዷፎ⋡⟬ฟࡢᮇ㛫ࢆἪங☜ㄆࡀ☜ᐇ࡟⾜࠼ࡿ 4㹼6᭶࡟㝈ᐃࡋ࡞ࡅࢀࡤ࡞ࡽࡎࠊ▷ᮇ㛫࡛༑ศ࡞ᶆᮏᩘࢆ㞟ࡵࡿࡇ࡜ࡀ㞴ࡋ࠸࡜⪃࠼ࡽࢀࡓࠋ
ࡇࢀ࡟ᑐࡋ࡚ࠊዷፎ㯤య㏥⦰≀ TypeϨࡣࠊ5㹼11 ᭶ࡢᮇ㛫࡟⫾Ꮚࡀ☜ㄆࡉࢀ࡞࠿ࡗࡓ
ಶయࡢ 92.1%࠿ࡽ☜ㄆࡉࢀࠊ11 ᭶ࡲ࡛ࡑࡢᩘࡣῶᑡࡋ࡞࠿ࡗࡓ㸦⾲㸯㸧ࠋࡉࡽ࡟ዷፎ㯤
య㏥⦰≀ TypeϨࡣࠊἪஙಶయࡢࡍ࡭࡚࠿ࡽ☜ㄆࡉࢀࡓࡇ࡜ࡼࡾࠊἪங☜ㄆࡼࡾ⢭ᗘࡢ㧗࠸ᣦᶆ࡛࠶ࡿ࡜⪃࠼ࡽࢀࡓࠋ
௚ࡢື≀࡛ࡣࠊฟ⏘ᚋಶయࢆ᳨ฟࡍࡿᣦᶆ࡜ࡋ࡚ࠊ⫾┙⑞ࡀ฼⏝ࡉࢀ࡚࠸ࡿ㸦Tsubota රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-6

et al. 1990; Mowat & Slough 1996㸧ࠋࡋ࠿ࡋ࢖ࣀࢩࢩ⛉࡟ࡘ࠸࡚ࡣࠊẕయ࡜⫾Ꮚࡢ⤖ྜᛶࡀప࠸ୖ⓶⤧ẟᛶ⫾┙࡛࠶ࡿࡓࡵ࡟⫾┙⑞ࡣṧࡽ࡞࠸㸦Friess et al. 1980㸧
ࠋࡑࡢࡓࡵ࢖ࣀࢩࢩ࡛ࡣࠊ⫾┙⑞ࢆฟ⏘ᚋಶయࡢᣦᶆ࡜ࡋ࡚⏝࠸ࡿࡇ࡜ࡣ࡛ࡁ࡞࠸ࠋ
௨ୖࡼࡾࠊᮏ◊✲࡛ࡣࠊぢⴠ࡜ࡋ࡞ࡃฟ⏘ᚋಶయࢆ㛗ᮇ㛫᳨ฟ࡛ࡁࡿዷፎ㯤య㏥⦰≀TypeϨࡀࠊ⫾Ꮚ☜ㄆࢆ⿵࠺ዷፎ⋡⟬ฟᣦᶆ࡜ࡋ࡚᭷⏝ᛶࡀ㧗࠸࡜ุ᩿ࡋࡓࠋዷፎ㯤య
㏥⦰≀ TypeϨࡢ☜ㄆ࡟ࡣࠊ㈝ࡸࡍ᫬㛫࡜ᢏ⾡ࡀồࡵࡽࢀࡿࡶࡢࡢࠊᶆᮏ཰㞟ࡢᅔ㞴ࡉࡀ኱ࡁ࡞ㄢ㢟࡛࠶ࡿ࢖ࣀࢩࢩ࡟ࡘ࠸࡚ࡣࠊ࡛ࡁࡿ㝈ࡾ㛗ᮇ㛫☜ㄆྍ⬟࡞ᣦᶆࢆ᥇⏝ࡋ࡚ࠊࡼࡾከࡃࡢᤕ⋓ಶయࢆά⏝ࡍࡿࡇ࡜ࡀ᭱ࡶඃඛᗘࡀ㧗࠸࡜⪃࠼ࡽࢀࡿࠋࡑࡢࡓࡵዷፎᮇ
௨እࡢᶆᮏ࠿ࡽ☜ᐇ࡟ุᐃྍ⬟࡞㯤య㏥⦰≀ࢆᰝᐃࡍࡿᡭἲࢆ᥇⏝ࡍࡿࡇ࡜ࡀ᭱⤊ⓗ࡟ࡣຠ⋡ࡀⰋ࠸࡜ุ᩿ࡉࢀࡿࠋ
ዷፎ⋡⟬ฟ࡟㐺ࡋࡓ᫬ᮇ රᗜ┴ࡢ࢖ࣀࢩࢩࡢཷ⫾࣭ฟ⏘᫬ᮇ࡟ࡣࠊዷፎᮇ㛫࡜࡯ࡰྠࡌ⣙ 4 ࠿᭶ࡢኚ␗ᖜࡀ࠶
ࡿࡇ࡜ࡀ᫂ࡽ࠿࡜࡞ࡗࡓ㸦ᅗ㸯㸧ࠋࡇࡢཷ⫾࣭ฟ⏘᫬ᮇࡢኚ␗ᖜࢆᅗ♧ࡋࡓ㸦ᅗ㸲ୖ㸧ࠋ
ᅗࡢ㏻ࡾࠊኚ␗ᖜࡀ 4 ࠿᭶࠶ࡿ࡜࠸࠺ࡇ࡜ࡣࠊ12 ᭶࡟ཷ⫾ࡋ࡚ 4 ᭶࡟ฟ⏘ࡍࡿಶయࡀ
࠸ࡿ୍᪉࡛ࠊ3 ᭶࡟ཷ⫾ࡋ 7 ᭶࡟ฟ⏘ࡍࡿಶయࡶᏑᅾࡍࡿࡇ࡜ࢆព࿡ࡍࡿࠋࡉࡽ࡟ࠊ࢖
ࣀࢩࢩ⛉ࡣཷ⫾ᚋ⣙ 20 ᪥㛫㸦ᅗ㸲ࡢዷፎึᮇ㸧ࡣ⬇ࡀ⫗║࡛ࡣ☜ㄆ࡛ࡁ࡞࠸኱ࡁࡉ࡛
࠶ࡿ㸦Perry & Rowlands 1962㸧ࡓࡵࠊ12㹼4 ᭶ࡲ࡛ࡣࠊ⬇ࡀ☜ㄆ࡛ࡁ࡞࠸ዷፎಶయࡀ
Ꮡᅾࡍࡿྍ⬟ᛶࡀ࠶ࡿࠋࡋࡓࡀࡗ࡚ࠊ࡝ࡢ᫬ᮇࡶࠊ⫾Ꮚ᭷↓ࡀุᐃ࡛ࡁ࡞࠸ዷፎึᮇಶ
యࡸฟ⏘ᚋಶయࡀᏑᅾࡍࡿࡇ࡜ࡀ᥎

ࡉࢀࡿࠋẚ㍑࡜ࡋ࡚ࠊࢽ࣍ࣥࢪ࢝ࡢཷ⫾࣭ฟ⏘᫬ᮇࡢኚ␗ᖜࢆ Suzuki et al.㸦1996㸧ࡢሗ࿌ࢆඖ࡟ᅗ♧ࡋࡓ㸦ᅗ㸲ୗ㸧ࠋࢽ࣍ࣥࢪ࢝࡟ࡶ
ኚ␗ᖜࡣ⣙ 3 ࠿᭶Ꮡᅾࡍࡿࡀࠊዷፎᮇ㛫ࡀ⣙ 230 ᪥࡜㛗࠸ࡓࡵࠊ2㹼4 ᭶ࡢ 3 ࠿᭶㛫ࡣࠊ࢖ࣀࢩࢩࡢࡼ࠺࡞ぢⴠ࡜ࡋ࡞ࡃࠊ⫾Ꮚ᭷↓࡟࡚ዷፎุᐃࡀྍ⬟࡛࠶ࡿࠋ௨ୖࡢཷ⫾࣭ฟ⏘᫬ᮇࡢ᥎ᐃ⤖ᯝ࠿ࡽࠊ࢖ࣀࢩࢩ࡛ࡣࠊዷፎᮇ㛫࡜࡯ࡰྠࡌ⣙ 4 ࠿᭶ࡢኚ␗ᖜࡀ࠶ࡿ࡜
࠸࠺≉ᚩࡀཎᅉ࡛ࠊ⫾Ꮚ☜ㄆࡔࡅ࡛ࡣዷፎ⋡ࡀ㐣ᑠ⟬ฟࡉࢀࡿྍ⬟ᛶࡀ♧၀ࡉࢀࡓࠋ
ዷፎ⋡⟬ฟ ᮏ◊✲࡛ࡣࠊ4㹼11 ᭶࡟⫾Ꮚࡀ☜ㄆࡉࢀࡓዷፎಶయ࡜ዷፎ㯤య㏥⦰≀ TypeϨࡀ☜ㄆࡉࢀࡓฟ⏘ᚋಶయࢆ⤌ࡳྜࢃࡏࡿࡇ࡜࡛ዷፎ⋡ࢆ⟬ฟࡍࡿࡇ࡜ࡀ࡛ࡁࡓ 㸦⾲㸰㹟㸧ࠋࡇࡢᡭἲࡣࠊ࢖ࣀࢩࢩࡢዷፎ⋡ࢆ⟬ฟࡍࡿ࡟࠶ࡓࡗ࡚⏕ࡌ࡚࠸ࡓ 2 ࡘࡢၥ㢟ࢆゎỴࡋࡓࠋ୍ࡘ┠ࡢၥ㢟ࡣࠊ⫾Ꮚ☜ㄆࡢࡳ࡟ࡼࡿ⟬ฟ᪉ἲ࡛ࡣዷፎ⋡ࢆ㐣ᑠ⟬ฟࡍࡿྍ⬟ᛶࡀ࠶ࡗࡓࡇ࡜࡛࠶ࡿࠋ㐣ཤ࡟၏୍ሗ࿌ࡉࢀࡓࢽ࣍ࣥ࢖ࣀࢩࢩࡢዷፎ⋡ 32㸣ࡣࠊᖺ㛫ࢆ㏻ࡌ࡚
ᚓࡽࢀࡓಶయࡢ⫾Ꮚ☜ㄆࡢࡳ࡛⟬ฟࡉࢀ࡚࠸ࡓ㸦ጜᓮ࣭ᆏᗞ 2010㸧ࠋᮏ◊✲࡛ࡶྠࡌࡼ
࠺࡟⫾Ꮚ☜ㄆࡢࡳࡢሙྜࡢዷፎ⋡ࡣ 31.82㸣࡜పࡃ⟬ฟࡉࢀࡓ㸦⾲㸰㹠㸧ࠋࡇࢀ࡟ᑐࡋ
࡚⫾Ꮚ࡜ዷፎ㯤య㏥⦰≀ TypeϨࡢ☜ㄆࡣ 92.42%࡜⟬ฟࡉࢀࡓ㸦⾲㸰㹟㸧ࠋࡇࡢ㐪࠸ࡣࠊ
ዷፎ㯤య㏥⦰≀ࡢ☜ㄆࢆ⾜࠺ࡇ࡜࡛ࠊฟ⏘ᚋࡢಶయࢆ᳨ฟ࡛ࡁࡓ࠿ࡽ࡛࠶ࡿࠋࡋࡓࡀࡗ රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-6

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ዷፎึᮇ㸦⃰࠸⅊Ⰽࡢ㒊ศ㸧࡛ࡣࠊ⬇ࡀ⫗║ほᐹ࡛ࡁࡿࡲ࡛ᡂ㛗ࡋ࡚࠸࡞࠸᫬ᮇࠊዷፎ
୰㹼ᚋᮇ㸦ⷧ࠸⅊Ⰽࡢ㒊ศ㸧ࡣ⬇ࡀ⫾Ꮚ࡟ᡂ㛗ࡋࠊ⫗║ほᐹࡀྍ⬟࡞᫬ᮇ࡛࠶ࡿࠋ

࡚ࠊ⫾Ꮚ☜ㄆࡢࡳ࡛ࡣࠊࡑࡢᖺࡢ⦾Ṫᮇ࡟ዷፎࡋࡓࡀㄪᰝ᫬࡟ࡣ⫾Ꮚࡀ☜ㄆ࡛ࡁ࡞࠸ዷ
ፎึᮇಶయ࡜ฟ⏘ᚋಶయࢆぢ㏨ࡋࠊ࢖ࣀࢩࢩࡢዷፎ⋡ࢆ㐣ᑠ⟬ฟࡍࡿࡇ࡜ࡀ᫂ࡽ࠿࡜࡞
ࡗࡓࠋᮏ◊✲࡟࡚ࠊዷፎุᐃࡀ࡛ࡁ࡞࠸ᮇ㛫ࢆ㝖እࡋࠊ☜ᐇ࡟ุᐃࡀ࡛ࡁࡿ 4㹼11 ᭶࡟࠾࠸࡚ࠊ⫾Ꮚ☜ㄆ࡟ຍ࠼࡚ࠊዷፎ㯤య㏥⦰≀ TypeϨࡢ☜ㄆࢆ⾜࠺ࡇ࡜࡛ࡼࡾṇ☜࡞ዷፎ⋡ࡀ⟬ฟ࡛ࡁࡿࡇ࡜ࡀ♧ࡉࢀࡓࠋ
ࡶ࠺୍Ⅼࡢၥ㢟ࡣࠊ⊁⊟ᅽࡢ㧗࠸᫬ᮇ࡜࢖ࣀࢩࢩࡢዷፎᮇࡢࡎࢀ࡟ࡼࡾዷፎಶయࡀᚓ
࡟ࡃࡃዷፎ⋡ࡀ⟬ฟ࡛ࡁ࡞࠿ࡗࡓࡇ࡜࡛࠶ࡿࠋᮏ◊✲ࡢ᪂ࡓ࡞⟬ฟ᪉ἲ࡛ࡣࠊዷፎಶయ
ࡀᚓ࡟ࡃ࠸࡜࠸࠺ၥ㢟Ⅼࢆዷፎ㯤య㏥⦰≀ TypeϨࡢ☜ㄆ࡟ࡼࡗ࡚ฟ⏘ᚋಶయࢆከࡃ᳨
ฟࡍࡿࡇ࡜࡛ᨵၿࡋࡓࠋ࡜ࡃ࡟⊁⊟ᅽࡢ㧗࠸ 11 ᭶࡟ᤕ⋓ࡉࢀࡓ㠀ዷፎಶయ࡛ࡶዷፎ㯤
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ࡓዷፎ⋡⟬ฟ࡟᭷ຠ࡞ᮇ㛫ࢆ 4㹼11 ᭶ࡢ 8 ࠿᭶㛫࡜ࡍࡿࡇ࡜࡛ࡶࠊከࡃࡢᤕ⋓ಶయࠊ࡜ࡃ࡟᭷ᐖᤕ⋓࡟࡚㥑㝖ࡉࢀࡓಶయࢆࣔࢽࢱࣜࣥࢢᮦᩱ࡜ࡋ࡚฼⏝࡛ࡁࡿࡼ࠺࡟࡞ࡗ
ࡓࠋ௨ୖࡼࡾࠊᮏ◊✲ࡢ᪂ࡓ࡞ዷፎ⋡⟬ฟ᪉ἲࡣࠊᶆᮏ཰㞟ࡢ㞴ࡋࡉ࡜࠸࠺ၥ㢟ࢆゎỴࡋࡓⅬ࡛ࠊỗ⏝ᛶࡀᴟࡵ࡚㧗࠸࡜⪃࠼ࡿࠋ௨ୖࢆࡲ࡜ࡵ࡚࢖ࣀࢩࢩࡢዷፎ⋡ࡢ⟬ฟ࡟㐺ࡋࡓᮇ㛫࡜ᣦᶆࢆḟ࡟♧ࡍࠋ රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-6

ㄪᰝᮇ㛫㸸4㹼11 ᭶⟬ฟᣦᶆ㸸⬇ࡸ⫾Ꮚ࡜ዷፎ㯤య㏥⦰≀ TypeϨ⟬ฟᘧ㸸⫾Ꮚ☜ㄆಶయᩘ + ዷፎ㯤య㏥⦰≀ TypeϨ☜ㄆಶయᩘዷፎ⋡㸦%㸧㸻                 ඲యࡢ࣓ࢫࡢᩘ

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 ᮏ◊✲ࢆᐇ᪋ࡍࡿ࡟࠶ࡓࡾࠊ2003 ᖺ࠿ࡽ 2011 ᖺ࡟රᗜ┴⊟཭఍ࡢከࡃࡢᨭ㒊ࡢⓙࡉࡲ࡟ᶆᮏࢆࡈᥦ౪㡬ࡁࡲࡋࡓࠋⓙࡉࡲࡢከ኱࡞ࡿࡈ༠ຊ࡞ࡋ࡛ࡣᮏ◊✲ࡣ㐙⾜࡛ࡁࡲࡏࢇ࡛ࡋࡓࠋࡇࡢሙࢆ೉ࡾ࡚ࠊ῝ࡃឤㅰ⏦ࡋୖࡆࡲࡍࠋ

ᘬ⏝ᩥ⊩
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࣭ 2004 ᖺ࠿ࡽ 2013 ᖺ࡟཰㞟ࡋࡓᤕ⋓ಶయࢆ⏝࠸࡚ࠊዷፎ⋡ࠊึᅇዷፎᖺ㱋ࠊᖹᆒ⫾Ꮚᩘࠊ⦾Ṫ᫬ᮇࢆ᫂ࡽ࠿࡟ࡋࡓࠋ࣭ ዷፎ⋡ࡣࠊ0 ṓ⩌ 8.8%ࠊ 1 ṓ⩌ 84.2%ࠊ2+ṓ⩌ 96.1%࡜⟬ฟࡉࢀࠊ࢖ࣀࢩࢩ࡛
ࡣ 0 ṓ࡛ࡶዷፎྍ⬟࡛࠶ࡿࡇ࡜ࠊ2 ṓ௨ୖ࡛ࡣ࡯ࡰẖᖺዷፎࡋ࡚࠸ࡿࡇ࡜ࡀ᫂ࡽ
࠿࡜࡞ࡗࡓࠋ࣭ ᖹᆒ⫾Ꮚᩘࡣࠊ0㹼1 ṓ⩌࡛ࡣ 2.40±1.14 㢌ࠊ2+ṓ⩌࡛ࡣࠊ4.26±1.41 㢌࡜⟬ฟࡉࢀࠊ㱋༊ศ㛫࡛᭷ព࡟␗࡞ࡗࡓࠋ࣭ ⦾Ṫ᫬ᮇࡣࠊཷ⫾᫬ᮇࡢࣆ࣮ࢡࢆ 1 ᭶ୗ᪪ࠊฟ⏘᫬ᮇࡢࣆ࣮ࢡࢆ 6 ᭶ୖ᪪ࡍࡿࠊ
1 ᖺ 1 ⏘ࡢ᫂░࡞Ꮨ⠇ᛶࢆ♧ࡍࡇ࡜ࡀ᫂ࡽ࠿࡜࡞ࡗࡓࠋࡓࡔࡋཷ⫾࡜ฟ⏘᫬ᮇ࡟ࡣࠊ⣙ 5 ࠿᭶ࡢኚ␗ᖜࡀ☜ㄆࡉࢀࡓࠋ࣭ රᗜ┴࡟⏕ᜥࡍࡿ࢖ࣀࢩࢩࡣࠊ࡯࡜ࢇ࡝ࡢಶయࡢึᅇዷፎᖺ㱋ࡀ 1 ṓ࡛࠶ࡿࡇ࡜ࠊ
2 ṓ௨ୖࡢ࣓ࢫ࡟ࡘ࠸࡚ࡣẖᖺᖹᆒ 4 㢌ࡢᏊࢆዷፎࡍࡿࡇ࡜࠿ࡽࠊࡁࢃࡵ࡚㧗࠸⦾Ṫຊࢆᣢࡘࡇ࡜ࡀ᫂ࡽ࠿࡜࡞ࡗࡓࠋ

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Key words 㸸 ዷፎ⋡ࠊึᅇዷፎᖺ㱋ࠊᖹᆒ⫾Ꮚᩘࠊཷ⫾᫬ᮇ࣭ฟ⏘᫬ᮇࠊᤕ⋓ಶయศᯒ㸬ࡣࡌࡵ࡟
㔝⏕ື≀ࡢዷፎ⋡ࠊึᅇዷፎᖺ㱋ࠊ⫾Ꮚᩘ࡜࠸࠺ᇶᮏⓗ࡞⦾Ṫ≉ᛶࡣࠊಶయ⩌ືែࡢᢕᥱ࡟୙ྍḞ࡞᝟ሗ࡛࠶ࡿ㸦Bailey 1984; ㎷ 2013㸧ࠋࡲࡓ⦾Ṫ᫬ᮇࡣࠊຠᯝⓗ࡟ಶయᩘ⟶⌮ࢆᐇ᪋ࡋ࡚࠸ࡃࡓࡵࡢᤕ⋓᫬ᮇࡢỴᐃࠊ㐺ษ࡞ㄪᰝ᫬ᮇࡢタᐃ࡟᭷⏝࡞᝟ሗ࡛࠶
ࡿ㸦Apollonio et al. 2011; Fonceca et al. 2011㸧ࠋࡉࡽ࡟ࡇࢀࡽ 4 ࡘࡢ⦾Ṫ≉ᛶࡣࠊಶయ⩌ࡢᰤ㣴≧ែࡸ⏕ᜥ⎔ቃࡢኚືࡢᙳ㡪ࢆཷࡅࡿࡓࡵࠊಶయ⩌ࡢ೺඲ᛶࢆホ౯ࡍࡿୖ࡛ࡶ㔜せ࡞ᣦᶆ࡜࡞ࡿ㸦Kaji 1988; ୰ᮧ࡯࠿ 2010㸧ࠋࢽ࣍ࣥ࢖ࣀࢩࢩ㸦Sus scrofa leucomystax㸹௨ୗ࢖ࣀࢩࢩ࡜⾲グ㸧ࡣࠊ᭷ྐ௨๓࠿ࡽ
᪥ᮏ࡟⏕ᜥࡋࠊே࡜ࡢ㛵ࢃࡾࡀ῝࠸ື≀࡛࠶ࡿ㸦㎷ 2013㸹ᶓᒣ 2014㸧ࠋࡑࢀ࡟ࡶ࠿࠿
ࢃࡽࡎࠊ⏕ែࡢゎ᫂ࡣᮍࡔ࡟༑ศ࡛࡞࠸ࠋ࡜ࡃ࡟⦾Ṫ᝟ሗࡣࠊᅜෆࡢ௚ࡢ⋇✀࠾ࡼࡧㅖእᅜࡢ࢖ࣀࢩࢩ࡟ẚ࡭࡚ᴟࡵ࡚ஈࡋ࠸ࠋࡇࢀࡲ࡛ࡢ⦾Ṫ᝟ሗࡣࠊ㣫⫱ಶయ㸦Ụཱྀ 2003㸧රᗜ┴᳃ᯘື≀◊✲ࢭࣥࢱ࣮࣭ රᗜ┴❧኱Ꮫ⮬↛࣭⎔ቃ⛉Ꮫ◊✲ᡤ රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-7

ࡸࣈࢱ㸦Sus scrofa domesticus㸧ࡢ▱ぢࠊࡲࡓࡣ⊁⊟⪅㛫࡛ཷࡅ⥅ࡀࢀ࡚ࡁࡓ⤒㦂▱࡟ࡼࡿ࡜ࡇࢁࡀከ࠿ࡗࡓࠋࡑࡢࡓࡵ㔝⏕ࡢ࢖ࣀࢩࢩࡢ⏕⌮Ꮫⓗ≉ᛶࢆ༑ศ࡟཯ᫎࡋ࡚ࡇ࡞
࠿ࡗࡓࠋ㏆ᖺࠊ⏕ᜥᇦࡢᣑ኱㸦⎔ቃ┬⮬↛ಖㆤᒁ⏕≀ከᵝᛶࢭࣥࢱ࣮ 2004㸧ࠊᤕ⋓ᩘࡸ㎰స≀⿕ᐖ㔠㢠ࡢቑຍ㸦⎔ቃ┬ 2010㸹㎰Ỉ┬ 2010㸧࡞࡝࠿ࡽࠊ࢖ࣀࢩࢩࡢಶయᩘ
ࡀ඲ᅜⓗ࡟ᛴቑࡋ࡚࠸ࡿࡇ࡜ࡀ᥎ᐃࡉࢀ࡚࠸ࡿࠋࡇࡢࡇ࡜࠿ࡽ௒ᚋࡢ⏕ᜥ࡜⿕ᐖࢆணࡍࡿୖ࡛ࠊ⦾Ṫ᝟ሗࢆᢕᥱࡍࡿࡇ࡜ࡀồࡵࡽࢀࡿࠋ࡜ࡃ࡟⟶⌮ࡢᇶ┙࡜࡞ࡿ⦾Ṫ᝟ሗ࡜ࡋ࡚ࠊዷፎ⋡ࠊึᅇዷፎᖺ㱋ࠊ⫾Ꮚᩘࠊ⦾Ṫ᫬ᮇࡢ᝟ሗᩚ⌮ࡀᚲせ࡛࠶ࡿࠋࡑࡢ⌮⏤࡜ࡋ࡚ࠊ࢖ࣀࢩࢩࡣࡇࢀࡽࡢ᝟ሗࡀஈࡋ࠸ࡔࡅ࡛࡞ࡃࠊྛ㡯┠࡟ࡘ࠸࡚௨ୗࡢㄢ㢟ࡀᣲࡆࡽࢀࡿ࠿ࡽ࡛࠶ࡿࠋࡲࡎዷፎ⋡࡟ࡘ࠸࡚ࡣࠊ㐣ཤ࡟୙㐺ษ࡞᪉ἲ࡛㐣ᑠ⟬ฟࡉࢀ㸦ጜᓮ࣭
ᆏᗞ 2010㸧ࠊⓗ☜࡟⟬ฟࡉࢀࡓሗ࿌ࡀ 1 ౛㸦Tsuji et al. 2013㸧ࡋ࠿࡞࠸ࡇ࡜࡛࠶ࡿࠋ
ḟ࡟⫾Ꮚᩘ࡟ࡘ࠸࡚ࡣࠊ1 ⭡࠶ࡓࡾᖹᆒ 4 㢌࡜࠸࠺ሗ࿌ࡀከ࠸ࡀ㸦Ụཱྀ 2003㸹ጜᓮ࡯࠿ 2009㸧ࠊ⫾Ꮚᩘࡢኚ␗ᖜࡀ⏕ࡌࡿせᅉࡣ୙࡛᫂࠶ࡾࠊᤕ⋓ࡢ⌧ሙ࡛ࡣࠊ౫↛࡜ࡋ࡚ 1
ᅇ࡟ 10 㢌⏘ࡴ࡜࠸࠺ㄆ㆑ࡀṧࡗ࡚࠸ࡿࡇ࡜࡛࠶ࡿࠋ᭱ᚋ࡟⦾Ṫ᫬ᮇ࡟ࡘ࠸࡚ࡣࠊ࢖ࣀ
ࢩࢩࡣᇶᮏⓗ࡟෤࡟ዷፎࡋ᫓࡟ฟ⏘ࡍࡿ㸦Ụཱྀ 2003㸧࡜࠸ࢃࢀ࡚࠸ࡿࡀࠊ⛅࡟ฟ⏕Ꮚ㸦࠸ࢃࡺࡿ࢘ࣜ࣎࢘㸧ࡀ┠ᧁࡉࢀࡿࡇ࡜ࡀ࠶ࡿࡓࡵ࡟ࠊฟ⏘ࡣ᫓࡜⛅ࡢ 2 ᅇ⾜ࢃࢀࡿ࡜࠸࠺ㄆ㆑ࡀ࠶ࡿࠋࡉࡽ࡟ฟ⏕᫬ᮇࡀ୍ᖺ୰Ꮡᅾࡍࡿࡇ࡜ࢆ♧၀ࡍࡿሗ࿌㸦ᑠᑎ࡯࠿
2012㸧ࡀ࠶ࡾࠊ࡜ࡃ࡟᝟ሗࡀΰ஘ࡋ࡚࠸ࡿࠋ
௨ୖࢆࡩࡲ࠼ࠊྛㄢ㢟ࡢゎỴ࡟ྥࡅ࡚ᶆᮏᩘࡀ඘ᐇࡋࡓරᗜ┴ࡢ࢖ࣀࢩࢩ࡟ࡘ࠸࡚ࠊ
ዷፎ⋡ࠊึᅇዷፎᖺ㱋ࠊ⫾Ꮚᩘࠊ⦾Ṫ᫬᫬ᮇࢆ᫂ࡽ࠿࡟ࡍࡿࡇ࡜ࢆ┠ⓗ࡜ࡋࡓࠋᮦᩱ࡜᪉ἲ
ᮦᩱ࡜᪉ἲ
 2004 ᖺ࠿ࡽ 2013 ᖺ࡟⊁⊟࠾ࡼࡧ᭷ᐖᤕ⋓࡟࡚ᤕ⋓ࡉࢀࡓ࣓ࢫ 107 㢌ࢆᑐ㇟࡜ࡋࡓࠋ
ᑐ㇟ಶయ࠿ࡽࡣࠊ༸ᕢࠊ⬇࠾ࡼࡧ⫾Ꮚࠊୗ㢡ࢆ᥇ྲྀࡋࡓࠋᑐ㇟ಶయࡢᖺ㱋ࢆୗ㢡ࡢṑࡢⴌฟ࡜஺᥮࠾ࡼࡧࠊ➨୍ᚋ⮻ṑࡢࢭ࣓ࣥࢺ㉁ᒙࡢᖺ㍯ᩘࡢ࢝࢘ࣥࢺ㸦Matschcke 1967;ᯘ࡯࠿ 1977; Boitani & Mattei 1992㸧࡟࡚Ỵᐃࡋࡓᚋࠊዷፎᖺ㱋ࡈ࡜࡟ 0 ṓ⩌ࠊ1 ṓ⩌ࠊ2㸩ṓ⩌ࡢ 3 ࡘࡢ⩌࡟ศ㢮ࡋࡓࠋዷፎ⋡࡜ึᅇዷፎᖺ㱋ࠊ⫾Ꮚᩘࠊ⦾Ṫ᫬ᮇࡣࠊ㐣ཤࡢሗ࿌ࢆඖ࡟ࡑࢀࡒࢀ௨ୗࡢ᪉ἲ࡛⟬ฟࡋࡓࠋ࣭ዷፎ⋡࡜ึᅇዷፎᖺ㱋͐༸ᕢෆࡢዷፎ㯤య㏥⦰≀ TypeϨ࡜Ꮚᐑෆࡢ⬇࠾ࡼࡧ⫾Ꮚࡢ☜ㄆ࡟࡚⟬ฟࡋࡓ㸦Tsuji et al. 2013㸧ࠋ࣭⫾Ꮚᩘ͐Ꮚᐑෆࡢ⬇࠾ࡼࡧ⫾Ꮚࡢᩘࢆ☜ㄆࡋࡓࠋ⫾Ꮚࡢᛶูࡣࠊእ㒊⏕Ṫჾࢆほᐹࡋุ࡚ᐃࡋࡓ㸦Inomata et al.1993㸧ࠋ࣭⦾Ṫ᫬ᮇ͐⫾Ꮚయ㔜ࢆ⏝࠸ࡓ⫾㱋᥎ᐃᘧ㸦Tsuji et al. 2013㸧࡟࡚⫾㱋ࢆ⟬ฟࡋࠊಶయࡢṚஸ᪥࠿ࡽཷ⫾᫬ᮇ࡜ฟ⏘᫬ᮇࢆ⟬ฟࡋࡓࠋ රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-7⤫ィฎ⌮ዷፎᖺ㱋⩌㛫ࡢዷፎ⋡ࡢᕪࡢ᭷↓ࡣࠊȮ2 ᳨ᐃᚋࠊBonferroni ἲ࡟ࡼࡾከ㔜ᛶࢆㄪᩚࡋุ࡚ᐃࡋࡓࠋዷፎᖺ㱋⩌㛫ࡢᖹᆒ⫾Ꮚᩘࡢẚ㍑࡟ࡣࠊMann-Whiteney ࡢ U ᳨ᐃࢆ⾜ࡗࡓࠋ⫾Ꮚᛶẚࡢᕪࡢ᭷↓࡟ࡘ࠸࡚ࡣࠊȮ2 ᳨ᐃࢆ⾜ࡗࡓࠋཷ⫾᫬ᮇ࡜ฟ⏘᫬ᮇࡢእࢀ
್ࡢ᳨ฟࡣࠊSmirnov-Grubbs ᳨ᐃࢆ⏝࠸ࡓࠋ඲࡚ࡢ᳨ᐃࡢ༴㝤⋡ࡣ 5%௨ୗ㸦p < 0.05㸧࡜ࡋࡓࠋ⤖ᯝ࡜⪃ᐹዷፎᖺ㱋⩌ࡈ࡜ᶆᮏᩘ
 0 ṓ⩌ࡣ 34 㢌ࠊ1 ṓ⩌ࡣ 19 㢌ࠊ2+ṓ⩌ࡣ 51 㢌ࠊୗ㢡ࡢᶆᮏࡀ࡞ࡃᖺ㱋୙᫂࡜࡞ࡗ
ࡓࡢࡀ 3 㢌࡛࠶ࡗࡓࠋ2+ṓ⩌ࡢዷፎᖺ㱋ᖜࡣࠊ2㹼11 ṓࡲ࡛☜ㄆࡉࢀࡓࠋዷፎ⋡ ዷፎ⋡ࡣࠊ0 ṓ⩌࡛ࡣ 8.8%ࠊ 1 ṓ⩌࡛ࡣ 84.2%ࠊ2+ṓ⩌࡛ࡣ 96.1%࡜⟬ฟࡉࢀࡓ㸦ᅗ㸯㸧ࠋ࡞࠾ 1 ṓ⩌࡜ 2+ṓ⩌ࡢዷፎ⋡࡟᭷ព࡞ᕪࡣ࡞࠿ࡗࡓ㸦p > 0.05㸧ࠋᅗ  ዷፎᖺ㱋㒆ูࡢዷፎ⋡௒ᅇࡢ⤖ᯝࡼࡾࠊ࢖ࣀࢩࢩࡣ 0 ṓ࡛ࡶዷፎࡍࡿࡇ࡜ࡀ᫂ࡽ࠿࡜࡞ࡗࡓࠋࡇࢀࡣᮏ◊✲࡟㝈ࡽࡎࠊ࣮ࣚࣟࢵࣃ࡞࡝ᾏእࡢ࢖ࣀࢩࢩ࡟࠾࠸࡚ࡶ☜ㄆࡉࢀ࡚࠸ࡿࡓࡵ㸦Fonseca etal. 2011㸧ࠊ࢖ࣀࢩࢩ࡜࠸࠺✀ࡢ≉ᚩ࡜⪃࠼ࡽࢀࡿࠋࡓࡔࡋࡑࡢ๭ྜࡣࠊᮏ◊✲ࡶ㐣ཤࡢሗ࿌ࡶ୍ᵝ࡟ప࠿ࡗࡓࡇ࡜࠿ࡽ㸦Fonseca et al. 2011; Gethöffer et al. 2007㸧
ࠊ0 ṓ࡛ࡢ
ዷፎࡣ୍⯡ⓗ࡟ࡣ㉳ࡇࡾ࡟ࡃ࠸࡜⪃࠼ࡽࢀࡿࠋ௒ᅇࡢ 0 ṓࡢዷፎಶయࡢయࢧ࢖ࢬࡣࠊྠ රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-7

᫬ᮇ࡟ᤕ⋓ࡉࢀࡓ 0 ṓࡢ㠀ዷፎಶయࡼࡾࡶ኱ࡁ࠿ࡗࡓࠋᾏእࡢ࢖ࣀࢩࢩࡢ◊✲࡛ࡣࠊึᮇᡂ㛗ࡢᮇ㛫࡟࠾ࡅࡿయ㔜ቑຍࡢ㐪࠸࡟ࡼࡗ࡚ࠊᛶᡂ⇍᫬ᮇ࡟ኚ␗ࡀ⏕ࡌࡿ㸦Gaillard

et al. 1992㸧ࡇ࡜ࡀ☜ㄆࡉࢀࡓࠋࡲࡓࠊᛶᡂ⇍࡟⮳ࡿ࡟ࡣ୍ᐃࡢయ㔜࡟㐩ࡍࡿᚲせᛶࡀ࠶ࡿ࡜ࡢሗ࿌ࡶ࠶ࡿ㸦Gethöffer et al. 2007㸧ࠋࡇࢀࡽࢆ⪃៖ࡍࡿ࡜ࠊ௒ᅇࡢ 0 ṓዷፎࡢせᅉࡢ 1 ࡘ࡜ࡋ࡚ࠊᙜヱಶయࡀ㏻ᖖࡼࡾࡶึᮇᡂ㛗ࡀⰋዲ࡛࠶ࡗࡓࡓࡵ࡟ࠊᛶᡂ⇍ࡀ᪩ࡲࡾዷፎ࡟⮳ࡗࡓࡇ࡜ࡀ⪃࠼ࡽࢀࡿࠋึᮇᡂ㛗᫬ࡢయ㔜ቑຍ࡟ᙳ㡪ࢆ୚࠼ࡿせᅉ࡜ࡋ࡚ࠊ㣗≀㈨※㔞ࠊ࡜ࡃ࡟⛅Ꮨࡢ୺せ࡞㣗≀࡛࠶ࡿሀᯝ㢮㸦ᑠᑎ࡯࠿ 2001㸧ࡢ⏕⏘㔞ࡀᣲࡆ
ࡽࢀࡿ㸦Matschke 1964; Mauget 1982㸧ࠋරᗜ┴࡛ࡣࣈࢼ⛉ሀᯝ㢮ࡢ㇏ปࡀ☜ㄆࡉࢀ࡚࠾ࡾ㸦⸨ᮌ࡯࠿ 2011㸧ࠊ㇏ป࡜ 0 ṓዷፎಶయࡢฟ⌧࡜ࡢ㛵ಀ࡟ࡘ࠸࡚௒ᚋ᳨ウࡍࡿᚲせࡀ࠶ࡿ࡜⪃࠼ࡿࠋ
0 ṓࡢዷፎ⋡࡟ẚ࡭࡚ 1 ṓࡢዷፎ⋡ࡣᛴ⃭࡟㧗ࡃ࡞ࡗࡓࡇ࡜࠿ࡽ㸦ᅗ㸯㸧ࠊከࡃࡢ࢖
ࣀࢩࢩࡢึᅇዷፎᖺ㱋ࡣ 1 ṓ࡛࠶ࡿ࡜ุ᩿ࡉࢀࡓࠋࡇࡢ⤖ᯝࡣࠊ࢖ࣀࢩࢩ࣓ࢫࡢ࡯࡜ࢇ
࡝ࡢᛶᡂ⇍᫬ᮇࡀ 1 ṓࡢ෤࡛࠶ࡿࡇ࡜㸦㎷ 2013㸧࡟ࡶ୍⮴ࡋࡓࠋࡘࡲࡾࠊᛶᡂ⇍࡟㐩ࡋࡓᖺࡢ⦾Ṫ᫬ᮇ࡟ึᅇዷፎࡍࡿࡇ࡜ࡀ᫂ࡽ࠿࡜࡞ࡗࡓࠋ
2 ṓ௨ୖࡢಶయࡢዷፎ⋡ࡀ 100%࡟㏆࠿ࡗࡓࡇ࡜ࠊࡇࡢ୰࡟ࡣ 2㹼11 ṓࡲ࡛ࡢዷፎಶ
యࡀྵࡲࢀ࡚࠸ࡿࡇ࡜ࠊᮏ◊✲ࡢศᯒಶయࡣᩘᖺ㛫⥅⥆ࡋ࡚཰㞟ࡉࢀࡓಶయ࡛࠶ࡿࡇ࡜
ࢆ⪃៖ࡍࡿ࡜ࠊ2 ṓ௨ୖࡢಶయࡢ࡯࡜ࢇ࡝ࡀẖᖺዷፎࡋ࡚࠸ࡿ࡜♧၀ࡉࢀࡓࠋࡇࡢዷፎ
⋡ࡢ㧗ࡉࡣࠊ࣮ࣚࣟࢵࣃࡢ࢖ࣀࢩࢩ࡜ࡶ୍⮴ࡋ㸦Fonseca et al. 2004; Fonseca et al.
2011㸧ࠊ᪥ᮏࡢ࢖ࣀࢩࢩ࡟ࡘ࠸࡚ࡶᡂ⋇࣓ࢫࡣࡁࢃࡵ࡚㧗࠸⦾Ṫຊࢆᣢࡘࡇ࡜ࡀ᫂ࡽ࠿
࡜࡞ࡗࡓࠋ᭱኱ዷፎᖺ㱋ࡣ 11 ṓ࡛࠶ࡗࡓࠋ㔝እࡢ࢖ࣀࢩࢩࡢᑑ࿨ࡀ 12㹼13 ṓ㸦Jezierski 1977㸧࡜ሗ࿌ࡉࢀ࡚࠸ࡿࡇ࡜࠿ࡽࠊᑑ࿨㏆ࡃࡲ࡛ዷፎྍ⬟࡛࠶ࡿ࡜⪃࠼ࡽࢀࡓࠋ⫾Ꮚᩘ࡜ᛶẚศᯒಶయ 107 㢌ࡢ࠺ࡕዷፎಶయࡣ 46 㢌࡛࠶ࡾࠊᚓࡽࢀࡓ⬇ࡶࡋࡃࡣ⫾Ꮚࡣィ 180 㢌࡛࠶ࡗࡓࠋ
1 ⭡࠶ࡓࡾࡢ⫾Ꮚᩘࡣ 1㹼7 㢌࡛ࠊ4 㢌ࡀ᭱ࡶከ࠿ࡗࡓࠋዷፎᖺ㱋⩌ࡈ࡜࡟ࡳࡿ࡜ࠊ0㹼1 ṓ⩌ࡢ⫾Ꮚᩘࡣ 1㹼4 㢌ࡢ⠊ᅖ࡟཰ࡲࡗࡓࠋ୍᪉ࠊ2+ṓ⩌࡛ࡣ 2㹼7 㢌࡜ኚ␗ᖜࡀ
኱ࡁࡃࠊ⫾Ꮚᩘࡶከ࠿ࡗࡓ㸦ᅗ㸰㸧ࠋ
0㹼1 ṓ⩌ࡢᖹᆒ⫾Ꮚᩘࡣ 2.40±1.14 㢌㸦N=5㸧
ࠊ2+ṓ⩌ࡢᖹᆒ⫾Ꮚᩘࡣࠊ4.26±1.41
㢌㸦N=38㸧࡛࠶ࡾࠊ2 ⩌㛫࡟᭷ព࡞ᕪࡀ࠶ࡗࡓ㸦p < 0.05㸧ࠋ඲యࡢᖹᆒ⫾Ꮚᩘࡣࠊ
4.07±1.45 㢌࡜⟬ฟࡉࢀࡓ㸦⾲㸯㸧
ࠋ

 

රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-7

ᅗ ୍⭡࠶ࡓࡾࡢ⫾Ꮚᩘࡢ㢖ᗘศᕸ⾲  ዷፎᖺ㱋ูࡢᖹᆒ⫾Ꮚᩘ
ዷፎᖺ㱋

1

ᖹᆒ⫾Ꮚᩘ ṓ

s

ṓ



s

ᖺ㱋୙᫂



s

ྜィ



s

රᗜ┴ࡢ࢖ࣀࢩࢩࡢ⫾Ꮚᩘࡣ᭱኱ 7 㢌࡛࠶ࡗࡓ㸦ᅗ㸰㸧ࠋᮏ◊✲ࡢᑐ㇟ಶయ࡛ࡣࠊ᤼
༸ᩘ⮬యࡶ౛እࢆ㝖ࡃ࡜᭱኱ 8 ಶ࡛࠶ࡗࡓࡇ࡜ࡀ☜ㄆࡉࢀ࡚࠸ࡿ㸦㎷ ᮍⓎ⾲㸧ࠋࡲࡓࠊᅜෆࡢ௚ᆅᇦ࡛ࡢሗ࿌࡛ࡶ⫾Ꮚᩘࡣ᭱኱ 9 㢌࡛࠶ࡾ㸦ጜᓮ࡯࠿ 2009㸧ࠊ࣮ࣚࣟࢵࣃࡢሗ
࿌࡛ࡶ 10 㢌௨ୖࡢ౛ࡣ☜ㄆࡉࢀ࡚࠸࡞࠸㸦Fonseca et al. 2004; Bywater et al. 2010㸧ࠋ௨ୖࡢ᰿ᣐ࠿ࡽࠊ࢖ࣀࢩࢩࡢ 1 ⭡࠶ࡓࡾࡢ⏘Ꮚᩘࡀ 10 㢌ࢆ㉸࠼ࡿྍ⬟ᛶࡣᴟࡵ࡚పࡃࠊ
⌧ሙ࡛ࡢㄆ㆑࡜ᮏ᮶ࡢ࢖ࣀࢩࢩࡢ⦾Ṫ≉ᛶ࡟ࡣ┦㐪ࡀ࠶ࡗࡓ࡜ุ᩿ࡉࢀࡓࠋ
ᖹᆒ⫾Ꮚᩘࡣ඲య࡛ࡣ 4.07±1.45 㢌࡜࡞ࡾࠊࡇࢀࡲ࡛࠸ࢃࢀ࡚࠸ࡓ 4㹼5 㢌㸦Ụཱྀ
2003㸹ጜᓮ࡯࠿ 2009㸧ࡢ⠊ᅖ࡟཰ࡲࡗࡓࠋࡋ࠿ࡋࠊዷፎᖺ㱋⩌࡛༊ศࡍࡿ࡜ 0㹼1 ṓ⩌࡜ 2+ṓ⩌㛫࡛᭷ព࡟␗࡞ࡿࡇ࡜ࡀ᫂ࡽ࠿࡜࡞ࡗࡓ㸦⾲㸯㸧ࠋࡇࡢ⤖ᯝࡼࡾࠊᖺ㱋ࢆ༊ูࡏࡎ࡟⫾Ꮚᩘࢆ⟬ฟࡋࡓሙྜࠊศᯒಶయࡢᖺ㱋ࡢ೫ࡾࡢ⛬ᗘ࡛඲యࡢᖹᆒ್ࡀኚືࡍ
ࡿࡇ࡜ࡀ♧ࡉࢀࡓࠋᐇ㝿࡟ Fernandez-Llario & Mateos-Quesada㸦1998㸧ࡣࠊศᯒࡍ
ࡿᶆᮏࡀⱝ㱋࡟೫ࡗࡓࡓࡵ࡟ࠊ඲యࡢᖹᆒ⫾Ꮚᩘࡀᑠࡉࡃ࡞ࡗࡓࡇ࡜ࢆሗ࿌ࡋ࡚࠸ࡿࠋ
ࡋࡓࡀࡗ࡚ࠊࡼࡾṇ☜࡟࢖ࣀࢩࢩࡢ⦾Ṫ≉ᛶࢆ཯ᫎࡋࡓᖹᆒ⫾Ꮚᩘࢆ⟬ฟࡍࡿࡓࡵ࡟ࡣࠊ
᭱ప㝈࡛ࡶึ⏘ಶయࡢከ࠸ 0㹼1 ṓ⩌࡜ 2+ṓ⩌࡟ᖺ㱋ࢆ༊ูࡋ࡚⟬ฟࡍࡿᚲせࡀ࠶ࡿࠋᮏ◊✲ࡼࡾࠊዷፎᖺ㱋ࡀࠊ⫾Ꮚᩘࡢኚ␗ᖜࡢⓎ⏕࡟኱ࡁࡃᙳ㡪ࡍࡿࡇ࡜ࡀ᫂ࡽ࠿࡜࡞ රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-7

ࡗࡓࠋࡓࡔࡋᶆᮏᩘࡢ୙㊊࡟ࡼࡾࠊຍ㱋࡜⫾Ꮚᩘࡢ㛵ಀࡣ᳨ウ࡛ࡁ࡞࠿ࡗࡓࡓࡵࠊ௒ᚋ
ྲྀࡾ⤌ࡴᚲせࡀ࠶ࡿࠋ௚ࡢせᅉ࡜ࡋ࡚ࠊ࣮ࣚࣟࢵࣃ࡛ࡣࠊ⫾Ꮚᩘࡀᆅᇦ࡟ࡼࡗ࡚␗࡞ࡾࠊ
⏕ᜥᆅࡢ⦋ᗘࡀ㧗࠸࡯࡝⫾Ꮚᩘࡀከࡃ࡞ࡿࡇ࡜㸦Bywater et al. 2010㸧࡞࡝ࡶ♧ࡉࢀ࡚࠸ࡿࠋ⌧ᅾࠊ᪥ᮏᅜෆࡢ࢖ࣀࢩࢩࡢ⏕ᜥᇦࡣ໭ୖࡋ࡚࠸ࡿ㸦⎔ቃ┬⮬↛ಖㆤᒁ⏕≀ከᵝᛶࢭࣥࢱ࣮ 2004㸧ࠋࡇࢀࡽࡢᆅᇦࡢ࢖ࣀࢩࢩࡢศᕸࡣࠊᒣ㛫㒊࠿ࡽ㒔ᕷ㒊ࡲ࡛ከᵝ࡞⎔ቃ࡟ᗈࡀࡗ࡚࠸ࡿࠋࡋࡓࡀࡗ࡚ࠊࡇࡢࡼ࠺࡞ศᕸᣑ኱ᆅᇦࡢ࢖ࣀࢩࢩࡢ⫾Ꮚᩘࢆẚ㍑
ࡋࠊᆅᇦⓗ࡞ኚ␗ᖜࡢ᭷↓ࢆ᳨ドࡍࡿࡇ࡜࡛ࠊቑຍ⋡ࡢホ౯ࡀྍ⬟࡟࡞ࡿ࡜⪃࠼ࡽࢀࡿࠋ
㞤㞝ุูࡀ࡛ࡁࡓ⫾Ꮚࡣࠊ180 㢌୰ 164 㢌࡛ࠊࡑࡢẚ⋡ࡣ࢜ࢫ㸸࣓ࢫ㸻1㸸1.22㸦࢜ࢫ 74 㢌ࠊ࣓ࢫ 90 㢌㸧࡜࡞ࡾࠊ㞤㞝ᕪࡣㄆࡵࡽࢀ࡞࠿ࡗࡓ 㸦df=1ࠊȮ2=0.78ࠊp > 0.05㸧ࠋࡇࢀࡼࡾ࢖ࣀࢩࢩࡣࠊฟ⏕᫬Ⅼ࡛ࡣ㞤㞝ࡢ๭ྜࡣᆒ➼࡛࠶ࡿࡇ࡜ࡀ♧၀ࡉࢀࡓࠋ
ཷ⫾ᮇ࣭ฟ⏘ᮇࡢ⟬ฟཷ⫾᫬ᮇࡣ 11 ᭶୰᪪࠿ࡽ 3 ᭶୰᪪ࡢᮇ㛫࡟࠶ࡾࠊ1 ᭶ୗ᪪ࢆࣆ࣮ࢡ࡜ࡋࡓࠋࡓࡔࡋ
1 ౛ࡢࡳ 5 ᭶୰᪪࡜⟬ฟࡉࢀࡓ㸦ᅗ㸱㸧ࠋฟ⏘᫬ᮇࡣ 3 ᭶୰᪪࠿ࡽ 7 ᭶୰᪪ࡢᮇ㛫࡟࠶
ࡾࠊ6 ᭶ୖ᪪ࢆࣆ࣮ࢡ࡜ࡋࡓࠋྠᵝ࡟ 1 ౛ࡢࡳ 9 ᭶୰᪪࡜⟬ฟࡉࢀࡓࠋ5 ᭶୰᪪࡟ཷ⫾ࡋ 9 ᭶୰᪪࡟ฟ⏘ࡍࡿ࡜᥎ᐃࡉࢀࡓ 1 ౛ࡣࠊ እࢀ್࡜ุᐃࡉࢀࡓ㸦p < 0.05㸧ࠋࡇࡢ 1౛ࢆ㝖ࡁࠊཷ⫾᪥୰ኸ್ࡣ 1 ᭶ 28 ᪥ࠊฟ⏘᪥୰ኸ್ࡣ 5 ᭶ 25 ᪥࡜⟬ฟࡉࢀࡓ㸦ᅗ㸱㸧ࠋᅗ  ཷ⫾᫬ᮇ࡜ฟ⏘᫬ᮇࡢศᕸ
රᗜ┴ࡢ࢖ࣀࢩࢩࡣࠊᇶᮏⓗ࡟෤࡟ዷፎࡋࠊ᫓࡟ฟ⏘ࡍࡿ 1 ᖺ࡟ 1 ⏘ࡢ᫂░࡞Ꮨ⠇⦾Ṫᛶࢆ♧ࡍࡇ࡜ࡀ᫂ࡽ࠿࡜࡞ࡗࡓࠋࡇࢀࡲ࡛᪥ᮏࡢ࢖ࣀࢩࢩࡢཷ⫾᫬ᮇࡣࠊ12㹼1 ᭶㸦⚄ᓮ 1993㸧ࠊฟ⏘᫬ᮇࡣ 4㹼5 ᭶ 㸦ᯘ࡯࠿ 1977㸧࡜࠸ࢃࢀ࡚࠸ࡓࡀࠊ⫾㱋᥎ᐃ࡟ᇶ࡙࠸࡚ศᯒࡋࡓᮏ◊✲࡟࡚ࠊරᗜ┴ࡢ࢖ࣀࢩࢩࡣࡑࢀࡼࡾࡶ 1 ࠿᭶⛬ᗘ㐜࠸ࡇ࡜ࡀ᫂ࡽ
࠿࡜࡞ࡗࡓࠋཷ⫾࣭ฟ⏘᫬ᮇ࡟ࡣࠊ᫂░࡞ࣆ࣮ࢡࡀ࠶ࡗࡓࡶࡢࡢࠊ⣙ 5 ࢝᭶㛫ࡢࡤࡽࡘࡁࡀ⏕ࡌ࡚࠸ࡓࠋྠᵝࡢ⌧㇟ࡣࠊ࣮ࣚࣟࢵࣃࡢ࢖ࣀࢩࢩ࡛ࡶሗ࿌ࡉࢀ࡚࠾ࡾ 㸦Mauget රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-7

1982; Fonseca et al. 2011㸧ࠊ࢖ࣀࢩࢩࡢ≉ᚩ࡜ࡋ࡚ࠊ⦾Ṫ᫬ᮇࡢኚ␗ᖜࡀ኱ࡁ࠸ࡇ࡜ࡀ
♧၀ࡉࢀࡓࠋ࣮ࣚࣟࢵࣃࡢ࢖ࣀࢩࢩ࡛ࡣࠊዷፎᖺ㱋ࡀప࠸࡯࡝ཷ⫾࣭ฟ⏘᫬ᮇࡀ㐜࠸ࡇ࡜ࠊ⛅ࡢ㣗≀㈨※㔞ࡀᑡ࡞࠸ࠊ࠶ࡿ࠸ࡣ⏕ᜥᆅࡢ⦋ᗘࡀప࠸⎔ቃ࡯࡝ኚ␗ᖜࡀ኱ࡁࡃ࡞
ࡿࡇ࡜ࡀሗ࿌ࡉࢀ࡚࠸ࡿࠋ
㸦Matschke 1964; Mauget 1982; Fonseca et al. 2004㸧ࡘࡲࡾࠊ࢖ࣀࢩࢩࡣᵝࠎ࡞せᅉ࡟ᛂࡌ࡚ࠊ⦾Ṫ᫬ᮇࡢኚ␗ᖜࢆኚ࠼ࡿᰂ㌾ᛶࡀ࠶ࡿ࡜᥎ᐹࡉ
ࢀࡿࠋࡑࡢࡓࡵࠊᅜෆࡢ࢖ࣀࢩࢩ࡟ࡘ࠸࡚ࡶ௒ᚋᆅᇦࡈ࡜࡟᳨ウࡍࡿᚲせࡀ࠶ࡿ࡜⪃࠼ࡿࠋ ᮏ◊✲࡛ࡣࠊዷፎಶయࡢ୰࡛ 1 ౛ࡢࡳࠊ5 ᭶ࡢ୰᪪࡟ཷ⫾ࡋ 9 ᭶ࡢ୰᪪࡟ฟ⏘ࡍࡿ࡜᥎ᐃࡉࢀࡓ㸦ᅗ㸱㸧ࠋࡇࢀࡣࠊ㐣ཤࡢᩥ⊩࡟࡚㆟ㄽࡉࢀ࡚࠸ࡿ⛅ฟ⏘࡟ヱᙜࡍࡿ 㸦⚄ᓮ
1993㸹Ụཱྀ 2003㸧
ࠋࡋ࠿ࡋࡑࡢ๭ྜࡣࠊ107 㢌୰ࡢ 1 㢌࡜ᴟࡵ࡚ప࠸ࡇ࡜࠿ࡽࠊරᗜ
┴ࡢ࢖ࣀࢩࢩ࡛ࡣ⛅ฟ⏘ࡣ⛥࡞౛࡛࠶ࡾࠊಶయᩘቑຍ࡬ࡢᙳ㡪ࡣప࠸࡜⪃࠼ࡽࢀࡓࠋࣚ
࣮ࣟࢵࣃࡢሗ࿌࡟࠾࠸࡚ࡶ⛅ฟ⏘ࡣ☜ㄆࡉࢀ࡚࠸ࡿࡀࠊࡇࢀࡣࠊᛶᡂ⇍ࡀ㐜ࢀࡓಶయ࠶
ࡿ࠸ࡣዷፎ୰ࡢὶ⏘ࡸฟ⏘ᚋࡢᏊࡢᾘኻ࡟ࡼࡾⓎ᝟ᅇᖐࡋࡓಶయ࡟ࡼࡿࡶࡢࡔ࡜⪃࠼ࡽࢀ࡚࠾ࡾࠊᮏ◊✲࡜ྠᵝ࡟≉ู࡞౛࡜ࡋ࡚ᢅࢃࢀ࡚࠸ࡿ 㸦Mauget 1982㸧ࠋࡲࡓ࢖ࣀ
ࢩࢩࡣ▷᪥Ꮨ⠇⦾Ṫᛶ࡛࠶ࡿࡓࡵ㸦Delcroix et al. 1990㸧ࠊ᫓࠿ࡽኟ࡬᪥㛗ࡀ㛗ࡃ࡞ࡿ
࡟ᚑࡗ࡚Ⓨ᝟ᅇᖐࡢⓎ⏕๭ྜࡣపࡃ࡞ࡾࠊኟᏘࡢཷ⫾⋡ࡣᴟࡵ࡚ప࠸࡜ሗ࿌ࡉࢀ࡚࠸ࡿ㸦Mauget 1982㸧ࠋࡋࡓࡀࡗ࡚ࠊ࢖ࣀࢩࢩࡀᖺ୰⦾Ṫࡍࡿ࡜࠸࠺஦㇟ࡣࠊᮏ᮶ࡢ࢖ࣀࢩ
ࢩࡢ⦾Ṫ⏕⌮ࢆ㐓⬺ࡋ࡚࠾ࡾࠊ⮬↛࡟ࡣ㉳ࡇࡾ࠼࡞࠸࡜⪃࠼ࡿࠋࡓࡔࡋවගࡽ㸦1988㸧
ࡣࠊ㣫⫱ୗ࡟࠾࠸࡚ࠊฟ⏘ࡋࡓ࣓ࢫࢆ 1 ࣨ᭶࡛Ꮚ࠿ࡽ㞳ࡋࠊࡑࡢ 1 㐌㛫௨ෆ࡟࢜ࢫ࡜ྠ
ᒃࡉࡏࡿࠊ࡜࠸࠺ேⅭⓗ࡞᧯సࢆຍ࠼ࡿࡇ࡜࡛ࠊ࣓ࢫࢆⓎ᝟ᅇᖐࡉࡏࠊ᫓࡜⛅ࡢᖺ 2 ᅇࡢฟ⏘ࢆྍ⬟࡜ࡋࡓࠋࡘࡲࡾ࢖ࣀࢩࢩࡣࠊఱࡽ࠿ࡢせᅉࡀຍࢃࡿࡇ࡜࡛ᖺ 2 ⏘ࢆྍ⬟࡜ࡍࡿື≀࡛࠶ࡿࡓࡵࠊ௒ᚋࡶ⛅ฟ⏘ࡢฟ⌧᭷↓ࢆᢕᥱࡍࡿࡇ࡜ࡣ㔜せ࡛࠶ࡿࠋࡲࡓࡑࢀ
࡜ྠ᫬࡟ࠊ⏕ࡲࢀࡓ⛅Ꮚࡢ⏕Ꮡ⋡࡜ಶయ⩌࡟୚࠼ࡿᙳ㡪࡟ࡘ࠸࡚ࡶ᳨ウࡀᚲせ࡜⪃࠼ࡿࠋㅰ㎡
 ᮏ◊✲ࢆᐇ᪋ࡍࡿ࡟࠶ࡓࡾࠊ2004㹼2013 ᖺ࡟රᗜ┴⊟཭఍ࡢከࡃࡢᨭ㒊ࡢⓙࡉࡲ࡟ᶆᮏࢆࡈᥦ౪㡬ࡁࡲࡋࡓࠋࡇࡢሙࢆ೉ࡾ࡚ࠊ῝ࡃឤㅰ⏦ࡋୖࡆࡲࡍࠋ

ᘬ⏝ᩥ⊩ጜᓮᬛᏊ࣭ᆏᗞᾈஅ࣭ᑠ㔝㔛ග࣭ᡞሯṇᖾ࣭୰ᔱ ⸅࣭➉ෆᛅ⩏࣭ᐩ⏣බ๎࣭ᮌ⁥኱௓㸬2009㸬⩌㤿┴࡟࠾ࡅࡿ࢖ࣀࢩࢩࡢศᕸᣑ኱࡜⦾Ṫ≧ἣ㸬⩌㤿┴❧⮬↛ྐ༤≀㤋◊✲ሗ࿌㸪13㸸
119-128㸬ጜᓮᬛᏊ࣭ᆏᗞᾈஅ㸬2010㸬࢖ࣀࢩࢩ࡟࠾ࡅࡿಶయ⩌ࣃ࣓࣮ࣛࢱ࣮ࡢゎ᫂㸬ⓒ℩ ᾈ ⦅㸬Ⴀ㎰⟶⌮ⓗ࢔ࣉ࣮ࣟࢳ࡟ࡼࡿ㫽⋇ᐖ㜵Ṇᢏ⾡ࡢ㛤Ⓨᡂᯝሗ࿌᭩㸬pp.55-60.㸦⊂㸧㎰◊ᶵᵓ ୰ኸ㎰ᴗ⥲ྜ◊✲ࢭࣥࢱ࣮㸪⟃Ἴ㸬 රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-7

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ࡿࢽ࣍ࣥ࢖ࣀࢩࢩࡢ⟶⌮ࡢ⌧≧࡜ㄢ㢟ࠖ㸪රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6 ྕ㸪pp.1-8㸬රᗜ┴᳃ᯘື≀◊✲ࢭࣥࢱ࣮㸬 රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-8➨ 㸶 ❶            ࢽ࣍ࣥ࢖ࣀࢩࢩࡢ࢘࢖ࣝࢫឤᰁ⑕
⡿‶◊୕ 1࣭᭹㒊ᚿಖ 1࣭㕥ᮌ⤢Ꮚ 1࣭὾ᓮ༓⳯⨾ 1࣭ୗ⏣ ᐂ 1࣭๓⏣ ೺ 1
せ Ⅼ࢖ࣀࢩࢩ࡟ឤᰁࡍࡿ࢘࢖ࣝࢫឤᰁ⑕ࡢ୰࡛ࠊࣄࢺࡸ⏕⏘ື≀㸦≉࡟ࣈࢱ㸧࡟ᑐࡋ࡚ၥ㢟࡜࡞ࡿ E ᆺ⫢⅖࢘࢖ࣝࢫࠊ㔜⑕⇕ᛶ⾑ᑠᯈῶᑡ⑕ೃ⩌࢘࢖ࣝࢫࠊ࣮࢚࢜ࢫ࣮࢟
⑓࢘࢖ࣝࢫࠊ᪥ᮏ⬻⅖࢘࢖ࣝࢫࡢ␿Ꮫㄪᰝࢆᐇ᪋ࡋࠊࣄࢺࡸື≀࡬ࡢࣜࢫࢡ࡟ࡘ࠸࡚
⪃ᐹࡋࡓࠋ

Key words㸸࢖ࣀࢩࢩࠊE ᆺ⫢⅖࢘࢖ࣝࢫࠊ᪥ᮏ⬻⅖࢘࢖ࣝࢫࠊ㔜⑕⇕ᛶ⾑ᑠᯈῶᑡ⑕ೃ⩌࢘࢖ࣝࢫࠊ࣮࢚࢜ࢫ࣮࢟⑓࢘࢖ࣝࢫ

㸫ࡣࡌࡵ࡟
ࢽ࣍ࣥ࢖ࣀࢩࢩ㸦Sus scrofa leucomystax㸹௨ୗ࢖ࣀࢩࢩ࡜⾲グ㸧ࡣ㞧㣗ᛶ࡛࠶ࡾࠊ
㏆ᖺ㎰స≀➼ࢆồࡵ࡚ࣄࢺࡢ⏕άᅪ࡟౵ධࡍࡿࡼ࠺࡟࡞ࡗ࡚ࡁࡓࠋࡲࡓࠊྂ᮶ࡼࡾ࢖ࣀ
ࢩࢩࡢ⫗ࡣࠊ㣗⏝࡜ࡋ࡚ᾘ㈝ࡉࢀ࡚࠸ࡿࠋ࢖ࣀࢩࢩ࡟ឤᰁࡋࠊྠ᫬࡟ࣄࢺࡸࡑࡢ௚ࡢື
≀ࠊ≉࡟ࣈࢱ࡟ࡶឤᰁࡋ⑓Ẽࢆᘬࡁ㉳ࡇࡍ࢘࢖ࣝࢫ࡟ࡣࠊᅜෆ࡛ࡣ E ᆺ⫢⅖࢘࢖ࣝࢫࠊ
㔜⑕⇕ᛶ⾑ᑠᯈῶᑡ⑕ೃ⩌࢘࢖ࣝࢫࠊ࣮࢚࢜ࢫ࣮࢟⑓࢘࢖ࣝࢫࠊ᪥ᮏ⬻⅖࢘࢖ࣝࢫࠊ㇜
࢖ࣥࣇ࢚ࣝࣥࢨ࢘࢖ࣝࢫࠊ㇜ࣃࣝ࣎࢘࢖ࣝࢫࠊ㇜⦾Ṫ࣭࿧྾ჾ㞀ᐖ⑕ೃ⩌࢘࢖ࣝࢫࠊ㇜
ࢧ࣮ࢥ࢘࢖ࣝࢫ 2 ࡞࡝ࡀ࠶ࡿ㸦⾲㸯㸧ࠋࡋ࠿ࡋࠊ࢖ࣀࢩࢩ࡟࠾ࡅࡿឤᰁ⑕◊✲ࡢṔྐࡣὸࡃࠊ◊✲ࡀ୙㊊ࡋ࡚࠸ࡿࡓࡵࠊ⾜ᨻࡸ⊁⊟⪅ࠊ୍⯡ᕷẸ࡞࡝⌧ሙࣞ࣋ࣝ࡟ࡲ࡛༑ศ࡞
᝟ሗࡀఏࢃࡗ࡚࠸ࡿ࡜ࡣゝ࠸㞴࠸ࠋࡑࡇ࡛ᮏㄽ࡛ࡣࠊᡃࠎࡢ᭱᪂ࡢ◊✲ᡂᯝ㸦Hara et al.
In press; Takahashi et al. In press; ๓⏣ 2014; Mahmoud et al. 2011; Shimoda et al.
2013; Shimojima et al. 2011; Shimoda et al.; Ohno et al. 2009㸧࡟ࡼࡾุ᫂ࡋࡘࡘ࠶
ࡿࠊ࢖ࣀࢩࢩ࡟࠾ࡅࡿ E ᆺ⫢⅖࢘࢖ࣝࢫࠊ᪥ᮏ⬻⅖࢘࢖ࣝࢫࠊ㔜⑕⇕ᛶ⾑ᑠᯈῶᑡ⑕
ೃ⩌࢘࢖ࣝࢫࠊ࠾ࡼࡧ࣮࢚࢜ࢫ࣮࢟⑓࢘࢖ࣝࢫ࡟㛵ࡍࡿឤᰁ≧ἣࡸឤᰁ⤒㊰ࡢ≉ᚩࢆ⾜ᨻᢸᙜ⪅ࡸ⊁⊟⪅࡞࡝ࡢᑐ⟇⪅ྥࡅ࡟⥲ㄽⓗ࡟⤂௓ࡋࠊࣄࢺࡸື≀࡬ࡢឤᰁࣜࢫࢡ࡟ࡘ
࠸࡚⪃ᐹࡋࠊࡲࡓឤᰁ㜵ᚚ᪉ἲ࡟ࡘ࠸࡚ᥦゝࡍࡿࠋ࡞࠾ࠊᮏ◊✲࡛ࡣᢠཎ㸦࢘࢖ࣝࢫࡢ
㑇ఏᏊ㸧᳨ฟ᳨ᰝ࡜ᢠయ᳨ฟ᳨ᰝࢆ⾜ࡗࡓࠋᢠཎ᳨ᰝࡣࢧࣥࣉࣜࣥࢢࡉࢀࡓᙜ᫬ࡢ࢘࢖
ࣝࢫࡢឤᰁ᭷↓ࢆ᳨⣴࡛ࡁࡿ᳨ᰝ࡛࠶ࡿࠋ୍᪉ࠊᢠయ᳨ᰝࡣࠊ㐣ཤࡢឤᰁṔࢆ᳨⣴ࡍࡿࡇ࡜ࡀ࡛ࡁࡿ᳨ᰝ࡛࠶ࡿࠋ
ᒣཱྀ኱Ꮫඹྠ⋇་Ꮫ㒊⑓ែไᚚᏛㅮᗙ⋇་ᚤ⏕≀Ꮫศ㔝 රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-8

⾲㸯㸬࢖ࣀࢩࢩ⏤᮶ࡢࣄࢺ࠾ࡼࡧື≀࡟ឤᰁࡍࡿྍ⬟ᛶࡢ࠶ࡿ୺せ࡞࢘࢖ࣝࢫឤᰁ⑕

( ᆺ⫢⅖( ᆺ⫢⅖ࡢ␿Ꮫ࡜୍⯡ᛶ≧E ᆺ⫢⅖ࡣࠊE ᆺ⫢⅖࢘࢖ࣝࢫ㸦௨ୗࠊHEV㸧ឤᰁ࡟ࡼࡗ࡚ᘬࡁ㉳ࡇࡉࢀࡿே⋇ඹ
㏻ឤᰁ⑕࡛࠶ࡿࠋⓎᒎ㏵ୖᅜ࡟࠾࠸࡚ࡣỈࢆ௓ࡋࡓ⣅ཱྀឤᰁ࡟ࡼࡾឤᰁࡍࡿ࡜࠸ࢃࢀ࡚
࠾ࡾࠊ௨๓ࡣࢃࡀᅜ࡛ࡣὶ⾜ᆅ࡟Ώ⯟ࡋࡓࣄࢺࡀឤᰁࡍࡿࠊ㍺ධឤᰁ⑕࡜ࡋ࡚ㄆ㆑ࡉࢀ
࡚࠸ࡓࠋࡋ࠿ࡋࠊ2003 ᖺ 4 ᭶࡟රᗜ┴࡛ࢽ࣍ࣥࢪ࢝㸦Cervus Nippon㸹௨ୗࢩ࢝࡜⾲グ㸧ࡢ⫗ࡢ⏕㣗࡟ࡼࡿឤᰁ஦౛ࡀⓎ⏕ࡋࠊࡇࢀࡀ㣗ရ࡟ࡼࡿឤᰁࡢึࡵ࡚ࡢ☜ㄆ஦౛࡜࡞ࡗࡓ㸦Tei et al. 2003㸧ࠋࡑࢀ௨㝆ࠊE ᆺ⫢⅖ࡣ㣗ရ፹௓ᛶឤᰁ⑕࡜ࡋ࡚ㄆ㆑ࡉࢀࡿࡼ
࠺࡟࡞ࡗࡓࠋࡲࡓࠊᕷ㈍ࡉࢀ࡚࠸ࡿ㇜ࡢ⏕ࣞࣂ࣮࠿ࡽ HEV ࡀ᳨ฟࡉࢀࡓࡇ࡜ࢆࡣࡌࡵ
㸦Yazaki et al. 2003㸧ࠊ⌧ᅾࠊ㇜⫗ࠊ࢖ࣀࢩࢩ⫗ࠊࢩ࢝⫗ࡢ⏕⫗ࡢႚ㣗ࡀᅜෆ࡛ࡢࣄࢺ
࡬ࡢឤᰁ※࡜ࡋ࡚㔜せどࡉࢀ࡚࠸ࡿࠋࡲࡓࠊE ᆺ⫢⅖࢘࢖ࣝࢫ࡟ࡣ 4 ✀㢮ࡢ㑇ఏᏊᆺࡀᏑᅾࡋࠊ1 ᆺࠊ2 ᆺࡀᾏእ࡛ὶ⾜ࡋ࡚࠸ࡿࡢ࡟ᑐࡋ࡚ࠊ3 ᆺࠊ4 ᆺࡣᅜෆ࡛ື≀ࡢ⫗࠿ࡽឤᰁࡋ࡚࠸ࡿࡇ࡜ࡶ▱ࡽࢀ࡚࠸ࡿࠋ⌧ᅾࠊឤᰁ⑕ࡢண㜵ཬࡧឤᰁ⑕ࡢᝈ⪅࡟ᑐࡍࡿ་⒪࡟㛵ࡍࡿἲᚊ㸦௨ୗࠊࠕឤᰁ⑕ἲࠖ
㸧࡟ᇶ࡙ࡁࠊE ᆺ⫢⅖ࡣឤᰁ⑕ἲࡢ➨ᅄ㢮ឤᰁ⑕࡟ᣦᐃࡉࢀ࡚࠾ࡾࠊᝈ⪅ࢆデ᩿ࡋࡓ་ᖌࡣಖ೺ᡤ࡟┤ࡕ࡟ᒆฟࡿࡇ࡜ࡀ⩏ົ࡙ࡅࡽࢀ࡚࠸
ࡿࠋᮏ◊✲࡛ࡣࠊ᪥ᮏࡢ 3 ᆅᇦ࡛ᤕ⋓ࡉࢀࡓ࢖ࣀࢩࢩ࠾ࡼࡧࢩ࢝࠿ࡽᚓࡽࢀࡓ⾑ᾮࢆศᯒࡋࠊᢠయ࠾ࡼࡧᢠཎ㸦࢘࢖ࣝࢫࡢ㑇ఏᏊ㸧ࡢ᳨ฟࡢ୧ᐇ㦂࡟ࡼࡾࠊHEV ࡢឤᰁ≧ἣࢆ
ศᯒࡋࡓࠋ
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රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-8

ᮦᩱ࠾ࡼࡧ᪉ἲ
᳨ᰝᑐ㇟ᆅᇦ࠾ࡼࡧࢧࣥࣉࣝᩘ
୰ᅜ࣭஑ᕞ࣭㛵ᮾᆅ᪉࡟࡚ 2009 ᖺ࠿ࡽ 2013 ᖺ࡟ᤕ⋓ࡉࢀࡓ࢖ࣀࢩࢩࡼࡾᚓࡽࢀࡓ
⾑Ύࢆ⏝࠸ࡓࠋ࢖ࣀࢩࢩ⾑Ύࡢࢧࣥࣉࣝᩘࡣ୰ᅜᆅ᪉ 167ࠊ஑ᕞᆅ᪉ 46ࠊ㛵ᮾᆅ᪉ 152࡛࠶ࡗࡓࠋࡲࡓẚ㍑ᑐ㇟࡜ࡋ࡚ࠊ࢖ࣀࢩࢩ⾑Ύࢆ᥇ྲྀࡋࡓࡢ࡜ྠᆅᇦ࡛࠶ࡿࠊ୰ᅜᆅ᪉
࡛ᤕ⋓ࡉࢀࡓࢩ࢝ࡼࡾᚓࡽࢀࡓ⾑Ύࡶศᯒ࡟⏝࠸ࡓࠋࢩ࢝⾑Ύࡢࢧࣥࣉࣝᩘࡣ 209 ࡛࠶ࡗࡓࠋ᳨ᰝ┠ⓗࡀ␗࡞ࡿࡓࡵࢧࣥࣉࣝᩘ࡟ࡣᕪࡀ࠶ࡿࠋ
+(9 ࡟ᑐࡍࡿᢠయࡢ᳨ฟELISA ἲ࡟ࡼࡾ⾑Ύ୰ࡢᢠయࢆ᳨ฟࡋࡓࠋELISA ᢠཎࡣࠊୗ㛵ࡢ HEV ᝈ⪅ࡢ㑇ఏ
Ꮚ㸦JTF-Yamagu11 ᰴ㸧ࢆᇶ࡟ N ᮎ➃㡿ᇦࢆḞᦆࡋࡓ ORF2 ࢆࢥ࣮ࢻࡍࡿ㑇ఏᏊࡣࣉ
ࣛ࢖࣐࣮Yamagu11 ORF2 112F (ClaI)(5’-GTA TCG ATC ACC ATG GCT GTG GCT
CCG GCC CCT-3’)࡜ Yamagu11 ORF2 660R-His (5’-GTA GAT CTT CAG TGA TGGTGA TGG TGA TGG TAC TCC CGG GTT TTA CCC A-3’)࡛ቑᖜࡋࡓࠋቑᖜࡋࡓ㑇ఏ
Ꮚࡣ ClaI ࡜ BglII ࡛ษ᩿ࡋࡓᚋࠊpCAGGS/MCS ࡢ ClaI ࡜ BglII ࢧ࢖ࢺ࡟ᤄධࡋࡓࠋᚓࡽࢀࡓⓎ⌧ࣉࣛࢫ࣑ࢻࢆ 293T ⣽⬊࡟࣏࢚ࣜࢳࣞࣥ࢖࣑ࣥࢆ⏝࠸࡚ࢺࣛࣥࢫࣇ࢙ࢡࢩ
ࣙࣥࡋࡓࠋࢺࣛࣥࢫࣇ࢙ࢡࢩࣙࣥ⣽⬊ࡣ RIPA ࡟ࡼࡗ࡚ 4Υ1 ᫬㛫ฎ⌮ࡋࡓᚋࠊ15000
ᅇ㌿ 4Υ30 ศ㛫㐲ᚰࡋ࡚ୖΎࢆᅇ཰ࡋࡓࠋᢳฟᢠཎࢆ 5ȣg/mε࡟ᕼ㔘ࡋࡓᚋࠊ100ȣεࢆྛ࢙࢘ࣝ࡟᥋✀ࡋ࡚ ELISA ࢆ⾜ࡗࡓࠋࣈࣟࢵ࢟ࣥࢢᾮ࠾ࡼࡧᢠཎᕼ㔘ᾮ࡟ࡣࣈࣟࢵ
ࢡ࢚࣮ࢫࢆ⏝࠸ࡓࠋ⾑Ύࡣ 1㸸100 ࡟ᕼ㔘ࡋࠊ஧ḟᢠయ࡟ࡣ࣌ࣟ࢟ࢩࢱ࣮ࢮᶆ㆑
ProteinA/G ࢆ 1㸸10000 ᕼ㔘ࡋ࡚⏝࠸ࡓࠋⓎⰍ࡟ࡣ Bio-Rad ࡢࣃ࣮࢜࢟ࢩࢲ࣮ࢮᇶ㉁࢟ࢵࢺࢆ⏝࠸ࡓࠋ
+(9 㑇ఏᏊ᳨ฟ
⾑Ύ࠿ࡽ QIAamp Viral RNA Mini Kit ࢆ⏝࠸࡚ RNA ࢆᢳฟࡋࠊHEV-F1 ࣉࣛ࢖࣐࣮࡜ HEV-R2 ࣉࣛ࢖࣐࣮ࢆ⏝࠸࡚ RT-PCR ࢆᐇ᪋ࠊ᭦࡟ RT-PCR ⏘≀ࢆࠊHEV-F2 ࣉࣛ࢖࣐࣮࡜ HEV-R1 ࣉࣛ࢖࣐࣮ࢆ⏝࠸࡚ Nested PCR ࢆ⾜࠸ࠊ㑇ఏᏊࡢ᳨ฟࢆヨࡳࡓࠋ
HEV 㑇ఏᏊࡢ᳨ฟࢆヨࡳࡓࠋ㝧ᛶࡀ␲ࢃࢀࡿࢧࣥࣉࣝ࡟ᑐࡋ࡚ࡣࠊሷᇶ㓄ิࢆỴᐃࡋࠊ᭱⤊ⓗ࡟ุᐃࡍࡿ࡜࡜ࡶ࡟ࠊ㑇ఏᏊᆺࡢゎᯒࢆ⾜ࡗࡓࠋ⤖ᯝ࠾ࡼࡧ⪃ᐹ࢖ࣀࢩࢩ࡛ࡣ୰ᅜᆅ᪉࡟࠾࠸࡚ 75 㢌୰ 23 㢌㸦31%㸧ࠊ஑ᕞᆅ᪉࡟࠾࠸࡚ 46 㢌୰ 10
㢌㸦22%㸧
ࠊ㛵ᮾᆅ᪉࡟࠾࠸࡚ 152 㢌୰ 12 㢌㸦8%㸧ࠊࢩ࡛࢝ࡣ୰ᅜᆅ᪉࡟࠾࠸࡚ 209
㢌୰ 1 㢌㸦0.5㸣㸧ࡀࠊHEV ࡟ᑐࡍࡿᢠయࢆಖ᭷ࡋ࡚࠸ࡓ㸦⾲㸰㸧ࠋᚑࡗ࡚ࠊHEV ࡟ᑐࡍࡿឤᰁ⋡ࡣᆅᇦᕪࡀ࠶ࡿࡶࡢࡢࠊ3 ᆅᇦ࡛ࡣࠊ࢖ࣀࢩࢩࢆឤᰁᐟ୺࡜ࡋ࡚ HEV ࡀศ රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-8

ᕸࡋ࡚࠸ࡿࡇ࡜ࡀ♧၀ࡉࢀࡓࠋࡲࡓࠊྠ୍ᆅᇦ࡛࠶ࡿ୰ᅜᆅ᪉࡟࠾࠸࡚ࡣࠊ࢖ࣀࢩࢩ࡜
ࢩ࢝ࡢ HEV ឤᰁ⋡ࡀ኱ࡁࡃ␗࡞ࡾࠊࢩ࢝ࡢឤᰁ⋡ࡣ㠀ᖖ࡟ప࠿ࡗࡓࡇ࡜࠿ࡽࠊࢩ࢝࡟
ࡘ࠸࡚ࡣࡇࢀࡲ࡛ࡢሗ࿌࡜ྠᵝ࡟ࠊHEV ࡢឤᰁࡣ㠀ᖖ࡟ࡲࢀ࡛࠶ࡿࡇ࡜ࡀ♧၀ࡉࢀࡓࠋ
HEV 㑇ఏᏊ᳨ᰝ࡟ࡼࡾࠊ࢘࢖ࣝࢫࡢ㑇ఏᏊࡢಖ᭷ࡢ᭷↓ࢆㄪ࡭ࡓࠋ࢖ࣀࢩࢩ࡛ࡣ୰
ᅜᆅ᪉ࡢಶయ 167 㢌୰ 6 㢌㸦4%㸧࠿ࡽࠊࢩ࢝࡟࠾࠸࡚ࡣ୰ᅜᆅ᪉ࡢಶయ 201 㢌୰ 1 㢌
㸦0.5%㸧ࡢ⾑Ύ࠿ࡽ HEV 㑇ఏᏊࡀ᳨ฟࡉࢀࡓࡀࠊ஑ᕞ࠾ࡼࡧ㛵ᮾᆅ᪉ࡢ࢖ࣀࢩࢩ࠿ࡽ
ࡣ HEV 㑇ఏᏊࡣ᳨ฟࡉࢀ࡞࠿ࡗࡓ㸦⾲㸱㸧ࠋHEV 㑇ఏᏊࡀ☜ㄆࡉࢀࡓ୰ᅜᆅ᪉ࡢ࢖ࣀ
ࢩࢩ 6 㢌࡜ࢩ࢝ 1 㢌࠿ࡽ᳨ฟࡉࢀࡓ HEV 㑇ఏᏊᆺࢆゎᯒࡋࡓ⤖ᯝࠊࡇࢀࡽࡢ㑇ఏᏊࡣ㠀ᖖ࡟㏆⦕࡛࠶ࡗࡓࠋ᭦࡟ࠊྠᆅᇦ࡛Ⓨ⑓ࡋࡓࣄࢺࡢ E ᆺ⫢⅖ᝈ⪅࠿ࡽ᳨ฟࡉࢀࡓ㑇ఏᏊࡀࡇࡢࢡࣛࢫࢱ࣮࡟ྵࡲࢀࡓࠋࡲࡓࠊࡇࡢᆅᇦࡢ HEV ࡣࠊ᪥ᮏࢆྵࡴ௚ࡢᆅ᪉࠿
ࡽ᳨ฟࡉࢀࡓ HEV ࡜ࡣ␗࡞ࡿ⊂❧ࡋࡓࢡࣛࢫࢱ࣮ࢆᙧᡂࡋࡓ㸦ᅗ㸯㸧ࠋ௨ୖࡼࡾࠊ୰
ᅜᆅ᪉࡛ࡣࠊ࠶ࡿ HEV ࡢ㑇ఏᏊᆺ࡟ࡼࡿ࢖ࣀࢩࢩࠊࢩ࢝ࠊࣄࢺ㛫࡛ឤᰁ⎔ࡀᡂ❧ࡋ࡚࠸ࡓࡇ࡜ࡀ♧၀ࡉࢀࡓࠋ⾲ 㸬+(9 ࡟ᑐࡍࡿᢠయಖ᭷⋡
࢖ࣀࢩࢩ

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⾲ 㸬⾑Ύ୰࡟࠾ࡅࡿ +(9 㑇ఏᏊ᳨ฟ⋡
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ᮏ◊✲ࢆྵࡵࠊ⾑ᾮ࠿ࡽ E ᆺ⫢⅖࢘࢖ࣝࢫ㸦HEV㸧ࡢ㑇ఏᏊࡀ᳨ฟࡉࢀ࡚࠸ࡿࠋࡍ
࡞ࢃࡕࠊ࢖ࣀࢩࢩ࠾ࡼࡧࢩ࢝࡟࠾࠸࡚ࡣ඲㌟ࡢ➽⫗ෆ࠾ࡼࡧྛ⮚ჾ࡟ HEV ࡀᏑᅾࡍࡿ
ࡇ࡜ࢆ♧ࡋ࡚࠾ࡾࠊ࢖ࣀࢩࢩ࠾ࡼࡧࢩ࢝ࢆႚ㣗ࡍࡿሙྜ࡟ࡣࠊ࡝ࡢ㒊఩࡟㛵ࡋ࡚ࡶ༑ศ
࡟ຍ⇕ࢆࡍࡿᚲせᛶࡀ࠶ࡿࠋࡲࡓㄪ⌮࡛౑⏝ࡋࡓჾලࡶ༑ศ࡟Ὑίࡍࡿᚲせࡀ࠶ࡿ㸦ᅗ
㸰㸧
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රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-8ᅗ  ࢖ࣀࢩࢩ࣭ࢩ࣭࢝ࣄࢺ࠿ࡽ᳨ฟࡉࢀࡓ +(9 㑇ఏᏊࡢ⣔⤫ゎᯒ࢖ࣀࢩࢩ⏤᮶ᰴࡣ㉥Ꮠ࣭⥳Ꮠࠊࢩ࢝⏤᮶ᰴࡣ㟷Ꮠࠊࣄࢺ⏤᮶ᰴࡣ⣸࡛♧ࡍࠋ
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ᅗ  ( ᆺ⫢⅖࢘࢖ࣝࢫࡢឤᰁࡢྍ⬟ᛶ㔜⑕⇕ᛶ⾑ᑠᯈῶᑡ⑕ೃ⩌ 6)76 ࢘࢖ࣝࢫ
6)76 ࡢ␿Ꮫ࡜୍⯡ᛶ≧㔜⑕⇕ᛶ⾑ᑠᯈῶᑡ⑕ೃ⩌㸦SFTS㸧ࡣ SFTS ࢘࢖ࣝࢫ㸦௨ୗࠊSFTSV㸧ࡢឤᰁ࡟ࡼ
ࡾ㸪ࣄࢺ࡟Ⓨ⇕ࠊⓑ⾑⌫ῶᑡࠊ⾑ᑠᯈῶᑡࠊ⫢㓝⣲ࡢୖ᪼࡞࡝ࢆᘬࡁ㉳ࡇࡍࠊ⮴Ṛᛶࡢ㧗࠸ே⋇ඹ㏻ឤᰁ⑕࡛࠶ࡿࠋ2011 ᖺ࡟୰ᅜ࡟࠾࠸࡚ึࡵ࡚ሗ࿌ࡉࢀࡓ⑓Ẽ࡛࠶ࡾࠊ୰ රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-8

ᅜ࡛ࡣࢲࢽࡀᮏ࢘࢖ࣝࢫࢆ፹௓ࡋࠊࡵࢇ⨺ࠊࣖࢠࠊ࢘ࢩࠊ࢖ࢾࠊࣈࢱࠊࢽ࣡ࢺࣜ࡞࡝ከ
ࡃࡢື≀ࡀ SFTSV ࡟ᑐࡍࡿᢠయࢆಖ᭷ࡋ࡚࠸ࡿ࡜ሗ࿌ࡉࢀ࡚࠸ࡿ㸦Yu et al. 2011ࠊᅗ
㸱㸧ࠋᅜෆ࡛ࡣ 2013 ᖺ 1 ᭶࡟ᒣཱྀ┴࡛ᅜෆึࡢ SFTS ᝈ⪅ࡀሗ࿌ࡉࢀࠊࡑࡢᚋࡢㄪᰝ
࡟ࡼࡾࠊ2005 ᖺ࠿ࡽ SFTS ᝈ⪅ࡀⓎ⏕ࡋ࡚࠸ࡓࡇ࡜ࡀุ᫂ࡋ࡚࠸ࡿ㸦Takahashi et al.
In press㸧ࠋࡑࡢ௚ SFTS ᝈ⪅ࡢⓎ⏕᫬ᮇࡣ 4 ᭶࠿ࡽ 10 ᭶࡛ࢲࢽࡢάື᫬ᮇ࡟୍⮴ࡍࡿࡇ࡜ࠊᖺ㱋ู࡛ࡣ 50 ṓ௨ୖࠊᆅᇦู࡛ࡣ୰ᅜࠊᅄᅜࠊ஑ᕞᆅ᪉࡟Ⓨ⏕ࡀከ࠸ࡇ࡜ࡀࠊ
≉ᚩ࡜ࡋ࡚ሗ࿌ࡉࢀ࡚࠸ࡿ㸦Takahashi et al. In press㸧ࠋ⌧ᅾࠊឤᰁ⑕ἲ࡟ᇶ࡙ࡁࠊᝈ
⪅ࢆデ᩿ࡋࡓ་ᖌࡣಖ೺ᡤ࡟┤ࡕ࡟ᒆฟࡿࡇ࡜ࡀ⩏ົ࡙ࡅࡽࢀ࡚࠸ࡿࡀࠊᅜෆ࡟࠾ࡅ
ࡿ SFTS ࡟㛵ࡍࡿ᝟ሗࡣᑡ࡞࠸ࠋᮏ◊✲࡛ࡣࠊ୰ᅜᆅ᪉ࡢ࢖ࣀࢩࢩ࠾ࡼࡧࢩ࢝࡟࠾ࡅࡿ
SFTSV ࡢឤᰁ≧ἣࢆศᯒࡋࡓࠋ
༸

⏘༸

Ꮴ໬
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ᡂ䝎䝙

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㻿㻲㼀㻿㼂

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㻿㻲㼀㻿㼂

྾⾑

྾⾑

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ᅗ  6)76 ࢘࢖ࣝࢫࡢឤᰁ⎔࡜㔝⏕ື≀࡜ࡢ㛵ಀᮦᩱ࠾ࡼࡧ᪉ἲ
᳨ᰝᑐ㇟ᆅᇦ࠾ࡼࡧࢧࣥࣉࣝᩘ
୰ᅜᆅ᪉࡛ 2009 ᖺ࠿ࡽ 2013 ᖺ࡟ᤕ⋓ࡉࢀࡓ࢖ࣀࢩࢩ 115 㢌࠾ࡼࡧࢩ࢝ 130 㢌ࡢ⾑Ύࢆ⏝࠸ࡓࠋ රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-8

6)769 ࡟ᑐࡍࡿᢠయࡢ᳨ฟ
SFTSV㸦HB29 ᰴ㸧ឤᰁ⣽⬊࠾ࡼࡧ mock ឤᰁ⣽⬊ࢆᢠཎ࡜ࡋࡓ ELISA ἲࢆ 100ࠊ
400ࠊ1600ࠊ6400 ಸᕼ㔘ࡋࡓ࢖ࢾ⾑Ύࢆ⏝࠸࡚ᐇ᪋ࡋࡓࠋ஧ḟᢠయ࡟ࡣ HRP ᶆ㆑
Protein A/G ࢆ⏝࠸ࡓࠋ⤖ᯝ࠾ࡼࡧ⪃ᐹ ୰ᅜᆅ᪉࡛ᤕ⋓ࡉࢀࡓ࢖ࣀࢩࢩ 115 㢌୰ 9 㢌㸦8%㸧
ࠊࢩ࢝ 130 㢌୰ 65 㢌㸦50%㸧ࡀSFTSV ࡟ᑐࡍࡿᢠయࢆಖ᭷ࡋ࡚࠸ࡓ㸦⾲㸲㸧ࠋࡲࡓࠊ᳃ᕝ࡯࠿㸦2013㸧ࡢ⤖ᯝ࠿ࡽࠊ㣫⫱≟ࡼࡾࡶ⊁⊟≟ࡢ᪉ࡀ SFTS ࢘࢖ࣝࢫᢠయࡢ㝧ᛶ⋡ࡀ㧗࠸ࡇ࡜ࡀሗ࿌ࡉࢀ࡚࠾ࡾࠊ
ᒣᯘ࡟ධࡿᶵ఍ࡀከ࠸࡜ឤᰁ⋡ࡀ㧗ࡲࡿྍ⬟ᛶࡀ♧၀ࡉࢀࡓࠋ௨ୖࡼࡾࠊᅜෆึࡢ
SFTS ᝈ⪅ࡀ☜ㄆࡉࢀࡓᆅᇦ࿘㎶࡟⏕ᜥࡍࡿ࢖ࣀࢩࢩ࠾ࡼࡧࢩ࢝࡟࠾࠸࡚ࠊSFTSV ឤᰁṔࡀ࠶ࡿࡇ࡜ࠊ᪥ᮏ࡟࠾࠸࡚ࡶ㔝⏕ື≀࡟ SFTSV ࡢឤᰁࡀᡂ❧ࡍࡿࡇ࡜ࡀ♧ࡉࢀࡓࠋࡇࢀࡲ࡛ࡢሗ࿌࠿ࡽ SFTSV ࡣࢲࢽ࡟ࡼࡾ፹௓ࡉࢀࡿࡓࡵࠊᮏ◊✲⤖ᯝࡣࠊࡇࡢᆅᇦ࡟
⏕ᜥࡍࡿࢲࢽࡣ SFTS ࢘࢖ࣝࢫࢆಖ᭷ࡍࡿࡋ࡚࠸ࡿྍ⬟ᛶࡀ࠶ࡿࡇ࡜ࢆ♧၀ࡋ࡚࠸ࡿࠋ
ࡇࡢྍ⬟ᛶࢆ⌮ゎࡋࠊࢲࢽ࡟ࡼࡿ่ညࢆ㜵ᚚࡍࡿᡭẁࢆྲྀࡿ࡜ඹ࡟ࠊSFTS Ⓨ⑕᫬ࡢ⑕≧ࡢ≉ᚩࢆࡼࡃ⌮ゎࡋࠊSFTSV ឤᰁࡀ␲ࢃࢀࡿሙྜ࡟ࡣ⮳ᛴ⑓㝔࡟⾜ࡁࠊࢲࢽ࡟่ည
ࡉࢀࡓࡇ࡜ࢆ࿌ࡆ࡚㎿㏿࡟デ᩿ࡋ࡚ࡶࡽ࠺ᚲせࡀ࠶ࡿࠋຍ࠼࡚ࠊࢲࢽ࡟ࡼࡾ፹௓ࡉࢀࡿ
ឤᰁ⑕ࡣࠊSFTS ௨እ࡟ࡶ᪥ᮏ⣚ᩬ⇕ࡸࣛ࢖࣒⑓ࠊࡘࡘࡀࡴࡋ⑓࡞࡝ᵝࠎ࠶ࡿࡓࡵࠊࢲ
ࢽ࡟ࡼࡿ่ည࡟ࡣᬑẁ࠿ࡽ༑ศ࡟ὀពࡍࡿᚲせࡀ࠶ࡿࠋ⾲ 㸬6)769 ࡟ᑐࡍࡿᢠయಖ᭷⋡
࢖ࣀࢩࢩ

ࢩ࢝

ㄪᰝᩘ





ᢠయ㝧ᛶᩘᢠయ㝧ᛶ⋡  ᮏ◊✲࡛࢖ࣀࢩࢩࡢᢠయ㝧ᛶ⋡ࡀࢩ࢝࡟ẚ࡭࡚ప࠿ࡗࡓ⤖ᯝ࡟㛵ࡋ࡚ࡣࠊཎᅉࡣ⌧ᅾ᫂☜࡟ࡣ࡛ࡁ࡞࠸ࡀࠊSFTSV ࢆ፹௓ࡍࡿࢲࢽࡢ྾⾑ື≀࡟ᑐࡍࡿႴዲᛶࡀ㛵୚ࡋ࡚࠸ࡿྍ⬟ᛶࡀ⪃࠼ࡽࢀࡿࡓࡵࠊ௒ᚋࡢ◊✲ࡀᮇᚅࡉࢀࡿࠋ
࣮࢚࢜ࢫ࣮࢟⑓࢘࢖ࣝࢫ
࣮࢚࢜ࢫ࣮࢟⑓ࡢ␿Ꮫ࡜୍⯡ᛶ≧࣮࢚࢜ࢫ࣮࢟⑓ࡣ㇜࣊ࣝ࣌ࢫ࢘࢖ࣝࢫ 1㸦௨ୗࠊPRV㸧࡟ࡼࡗ࡚Ⓨ⑕ࡍࡿࠊࣈࢱ࠾ࡼ
ࡧ࢖ࣀࢩࢩࡀᑐ㇟࡜࡞ࡿᒆฟఏᰁ⑓࡛࠶ࡾࠊᗂ㇜ࡀⓎ⑕ࡍࡿ࡜⮴Ṛ⋡ࡀ㧗࠸ࡓࡵࠊ㣴㇜ රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-8

ᴗ࡟ከ኱࡞⤒῭ⓗ⿕ᐖࢆཬࡰࡍࠋឤᰁࡣࠊ㰯Ồࡢ㣕ἓࠊᦤ㣗ࠊയཱྀ࠿ࡽࡢឤᰁࡸởᰁ≀ࡢᦤྲྀࡀཎᅉ࡜࡞ࡾࠊᡂ㇜࡛ࡣṚஸ⋡ࡣ㧗ࡃ࡞ࡃࠊከࡃࡢሙྜࡣ୙㢧ᛶឤᰁ㸨࡟⤊ࢃࡿࡇ࡜ࡀከ࠸ࠋࡋ࠿ࡋࠊࣈࢱࡸ࢖ࣀࢩࢩ௨እࡢື≀࡟ឤᰁࡋࡓሙྜࡣࠊ⚄⤒⑕≧࡛࠶ࡿᥙ⑛⑕ࢆᘬࡁ㉳ࡇࡋࠊ࡯ࡰ 100%Ṛஸࡍࡿࠋឤᰁᚋ࡟ PRV ࡟⪏㐣ࡋࡓࣈࢱ࡛ࡣࠊ࢘࢖ࣝ
ࢫࡀ₯అឤᰁ㸨㸨ࡍࡿ≉ᚩࡀ࠶ࡾࠊᅜෆ࡛ࡣࠊ
ࠕ࣮࢚࢜ࢫ࣮࢟⑓㜵␿ᑐ⟇せ㡿ࠖ࡟ᇶ࡙ࡁࠊ
ࣈࢱ࡟࠾࠸࡚ࡣ࣡ࢡࢳࣥࢥࣥࢺ࣮ࣟࣝ࡟ࡼࡾ 36 ┴࡛ PRV ࡢΎί໬࡟ᡂຌࡋ࡚࠾ࡾࠊ
11 ┴࡛Ὑί໬ᑐ⟇ࡀᐇ᪋ࡉࢀ࡚࠸ࡿ㸦㎰ᯘỈ⏘┬ᾘ㈝࣭Ᏻ඲ᒁື≀⾨⏕ㄢ㸧ࠋࡋ࠿ࡋࠊ
⮫ᗋ⑕≧ࢆ♧ࡋ࡚࠸ࡿࣈࢱࡢ᦬Ⓨ࣭Ὲử௨እ࡟ࡶࠊ₯అឤᰁࡋ࡚࠸ࡿ㔝እ࢘࢖ࣝࢫᢠయ
㝧ᛶࣈࢱࢆ᪩ᮇ࡟Ⓨぢࡍࡿᚲせࡀ࠶ࡿࠋ
㏆␥ᆅ᪉ࡣࠊ᏶඲࡟࣮࢚࢜ࢫ࣮࢟⑓ࡢΎί໬࡟ᡂຌࡋ࡚࠸ࡓࡀࠊዉⰋ┴࡟࠾࠸࡚ࠊ
1997 ᖺ࡟࢖ࣀࢩࢩࡢ⏕⫗ࢆ㣗࡭ࡓ⊟≟ࡀ࣮࢚࢜ࢫ࣮࢟⑓࡟ࡼࡾከᩘṚஸࡋࡓࡇ࡜ࡀሗ
࿌ࡉࢀ࡚࠸ࡿ㸦ᖾ⏣࡯࠿ 2000㸧
ࠋᚑࡗ࡚ࠊ࢖ࣀࢩࢩ࡟ࡣ₯ᅾⓗ࡟࣮࢚࢜ࢫ࣮࢟⑓࢘࢖ࣝ
ࢫࡀឤᰁࡋ࡚࠸ࡿྍ⬟ᛶࡀ㧗࠸࡜⪃࠼ࡽࢀࡿࡀࠊᅜෆࡢ࢖ࣀࢩࢩ࡟࠾ࡅࡿ࣮࢚࢜ࢫ࣮࢟
⑓࢘࢖ࣝࢫࡢឤᰁ≧ἣ࡟ࡘ࠸࡚ࡣሗ࿌ࡀ࡞࠸ࠋᮏ◊✲࡛ࡣࠊ࣮࢚࢜ࢫ࣮࢟⑓ࡢ⾑Ύ␿Ꮫㄪᰝࢆᐇ᪋ࡋࡓࠋ
㸨୙㢧ᛶឤᰁ㸸ឤᰁࡣࡋ࡚࠸ࡿࡀࠊⓎ⑓ࢆకࢃ࡞࠸ឤᰁࠋ

㸨㸨₯అឤᰁ㸸࢘࢖ࣝࢫࡣ᳨ฟࡉࢀ࡟ࡃ࠸ࡀࠊᣢ⥆ⓗ࡟ឤᰁࡀᡂ❧ࡋ࡚࠸ࡿ≧ἣࠋᮦᩱ࠾ࡼࡧ᪉ἲ
᳨ᰝᑐ㇟ᆅᇦ࠾ࡼࡧࢧࣥࣉࣝᩘ
ࣈࢱ࡛ࡢ࣮࢚࢜ࢫ࣮࢟⑓ࡢΎί໬࡟ᡂຌࡋ࡚࠸ࡿ 3 ┴ࠊ㏆␥ᆅ᪉ࡢ A ┴࠾ࡼࡧ B ┴ࠊ୰ᅜᆅ᪉ࡢ C ┴࡛ࠊ2007 ᖺ࠿ࡽ 2010 ᖺ࡟ᤕ⋓ࡉࢀࡓ࢖ࣀࢩࢩࡑࢀࡒࢀ 50 㢌ࠊ71 㢌ࠊ
52 㢌࠿ࡽᚓࡽࢀࡓ⾑Ύࢆ⏝࠸ࡓࠋ
359 ࡟ᑐࡍࡿᢠయࡢ᳨ฟ
࢘࢖ࣝࢫࡣ PRV Indiana ᰴࠊ⣽⬊ࡣࣈࢱ⏤᮶ CPK ⣽⬊㸦ື≀⾨⏕◊✲ᡤࡼࡾศ୚㸧ࢆ⏝࠸࡚୰࿴ヨ㦂࡟ࡼࡾࠊ⾑Ύ୰ࡢ࢘࢖ࣝࢫᢠయࢆ᳨ฟࡋࡓࠋPRV ࡟ᑐࡍࡿᢠయ㝧ᛶಶయࢆ᳨ฟࡍࡿࡓࡵ࡟ࠊ1㸸10 ᕼ㔘⾑Ύࢆ⏝࠸ࡓ 80㸣ࣉ࣮ࣛࢡῶᩘヨ㦂ࢆᐇ᪋ࡋࡓࠋ
㝧ᛶಶయ࡟㛵ࡋ࡚ࡣࠊ⾑Ύࢆ஧ಸ㝵ẁᕼ㔘ࡋ࡚ᢠయ౯ࢆồࡵࡓࠋ࣡ࢡࢳࣥᰴ࡜㔝እᰴࡢ㆑ู
㔝እᰴឤᰁಶయ࡟ࡢࡳฟ⌧ࡍࡿ gE ᢠయࡢ᳨ฟࢆࠊIDEXX ♫ࡢ g1 Antibody ELISAࢆ⏝࠸࡚ᐇ᪋ࡋࡓࠋ᪉ἲࡣ〇ရῧ௜ࡢࣉࣟࢺࢥ࣮ࣝ࡟ᚑࡗࡓࠋ
⤖ᯝ࠾ࡼࡧ⪃ᐹ㏆␥ᆅ᪉ࡢ 2 ┴࡛ᤕ⋓ࡉࢀࡓ 6 㢌㸦5%㸧ࡢ࢖ࣀࢩࢩ࡟࠾࠸࡚ࠊPRV ࡟ᑐࡍࡿ࢘࢖ࣝ
ࢫ୰࿴ᢠయ㝧ᛶࡀ☜ㄆࡉࢀࡓࠋ୍᪉ࠊ୰ᅜᆅ᪉ࡢ㸯┴࡛ᤕ⋓ࡉࢀࡓ࢖ࣀࢩࢩ࡟࠾࠸࡚ࡣࠊ 

රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-8

㝧ᛶಶయࡣ☜ㄆࡉࢀ࡞࠿ࡗࡓ㸦⾲ 5㸧ࠋ࢘࢖ࣝࢫ୰࿴ᢠయಖ᭷࡛࠶ࡗࡓ࢖ࣀࢩࢩ 6 㢌ࡣࠊࡍ࡭࡚ gE ࡟ᑐࡍࡿᢠయࢆಖ᭷ࡋ࡚࠸ࡓࠋᚑࡗ࡚࢖ࣀࢩࢩࡣ࣮࢚࢜ࢫ࣮࢟⑓࢘࢖ࣝࢫࡢ
㔝⏕ᰴ࡟ឤᰁࡋ࡚࠾ࡾࠊࣈࢱ࡬ࡢឤᰁ※࡜࡞ࡾᚓࡿࡇ࡜ࡀࡀ☜ㄆࡉࢀࡓ㸦ᅗ 4㸧ࠋ㎰ሙ
ෆ࡛ࡢ PRV Ύί໬ࡀᡂ❧ࡋ࡚࠸ࡿሙྜ࡟࠾࠸࡚ࡶࠊ≉࡟࢖ࣀࢩࢩ࠿ࡽࡢ㔝እᰴឤᰁࡢ
ྍ⬟ᛶࢆ⪃៖ࡋࠊᑐ⟇ࢆᐇ᪋ࡍࡿࡇ࡜ࡀᚲせ࡛࠶ࡿࠋ⾲ 㸬࣮࢚࢜ࢫ࣮࢟⑓࢘࢖ࣝࢫ㸦359㸧࡟ᑐࡍࡿᢠయಖ᭷⋡
㏆␥ᆅ᪉

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

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㣴㇜
㻟㻢㒔㐨ᗓ┴䛷䝽䜽䝏䞁䝁䞁䝖䝻䞊䝹䛻䜘䜚Ύί໬

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㔝እᰴ䜴䜲䝹䝇䜢ᣢ䛴ྍ⬟ᛶ䛜㧗䛔ᅗ  ࣮࢚࢜ࢫ࣮࢟⑓࢘࢖ࣝࢫ㸦359㸧ࡢឤᰁࡢྍ⬟ᛶ᪥ᮏ⬻⅖᪥ᮏ⬻⅖ࡢ␿Ꮫ࡜୍⯡ᛶ≧᪥ᮏ⬻⅖ࡣࠊ᪥ᮏ⬻⅖࢘࢖ࣝࢫ㸦௨ୗࠊJEV㸧ࡀཎᅉࡢឤᰁ⑕࡛ࠊ့ங㢮࠿ࡽ㫽㢮ࡲ࡛ᗈ⠊࡞ື≀✀࡟ឤᰁࡍࡿࡀࠊ኱㒊ศࡣ୙㢧ᛶឤᰁࢆ♧ࡍࠋࣄࢺ࠿ࡽࣄࢺ࡬ࡢឤᰁࡣ࡞
ࡃࠊቑᖜື≀㸨㸦ࣈࢱ㸧ࡢయෆ࡛࠸ࡗࡓࢇቑ࠼࡚⾑ᾮ୰࡟ฟ࡚ࡁࡓ࢘࢖ࣝࢫࢆࠊ⺅ࡀ྾⾑ࡋࠊࡑࡢୖ࡛ࣄࢺ࡞࡝ࢆ่ࡋࡓ᫬࡟ឤᰁࡍࡿࠋࣄࢺࡸ㤿ࡣឤᰁࡍࡿ࡜㔜⠜࡞⬻⅖ࢆⓎ⑕ࡍࡿࡇ࡜ࡀ࠶ࡿࡀࠊᅜෆ࡛ࡣ㏆ᖺ࣡ࢡࢳࣥ᥋✀࡟ࡼࡾࠊࣄࢺࡸ㤿࡛ࡢⓎ⏕ࡣ⃭ῶࡋ࡚࠸ࡿࠋࣈࢱࡣዷፎ㇜࡛ὶ⏘ࢆᘬࡁ㉳ࡇࡍࡇ࡜ࡀ࠶ࡾࠊࡲࡓ㏆ᖺࠊ࢘ࢩ࡛ࡢ JEV ࡟ࡼࡿ⬻⅖ࡢⓎ⏕ࡀᐑᓮ┴ࡸឡ▱┴࡛ሗ࿌ࡉࢀ࡚࠸ࡿ㸦ຍྂ࡯࠿ 2011; ∦ᒣ 2013㸧ࠋ᪥ᮏ⬻ 

රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-8

⅖ࡣࠊឤᰁ⑕ἲ࡟ᇶ࡙ࡁࠊᝈ⪅ࢆデ᩿ࡋࡓ་ᖌࡣಖ೺ᡤ࡟┤ࡕ࡟ᒆฟࡿࡇ࡜ࡀ⩏ົ࡙ࡅࡽࢀ࡚࠸ࡿࠋ
2008 ᖺ 12 ᭶࡟ࠊරᗜ┴ෆ࡛ᤕ⋓ࡉࢀࡓ࢖ࣀࢩࢩࡢ⾑ᾮ୰࠿ࡽ࢘࢖ࣝࢫࡀศ㞳ࡉࢀࠊ
᭦࡟ 2009 ᖺ 5 ᭶࡟රᗜ┴࡟࠾࠸࡚ᤕ⋓ࡉࢀࡓ࢖ࣀࢩࢩࡢ⾑ᾮ࠿ࡽࡣ࢘࢖ࣝࢫ㑇ఏᏊࡀ᳨ฟࡉࢀࡓ㸦㧗ᓮ࡯࠿ 2009㸧ࠋࡑࡢࡓࡵࠊ᪥ᮏ⬻⅖࢘࢖ࣝࢫࡢឤᰁ⎔࡟࠾࠸࡚ࠊ࢖ࣀࢩ
ࢩࡶቑᖜື≀࡜࡞ࡿྍ⬟ᛶࡀᣦ᦬ࡉࢀ࡚࠸ࡿࠋᮏ◊✲࡛ࡣࠊ㏆␥ᆅ᪉࠾ࡼࡧ୰ᅜᆅ᪉ࡢ
࢖ࣀࢩࢩ࠾ࡼࡧࢩ࢝࡟࠾ࡅࡿ JEV ࡢឤᰁ≧ἣࢆศᯒࡋࡓࠋ
㸨ቑᖜື≀㸸࢘࢖ࣝࢫࢆయෆ࡛ቑᖜࡋࠊ௚ࡢᐟ୺࡟౪⤥ࡍࡿࠋ ᮦᩱ࠾ࡼࡧ᪉ἲ
᳨ᰝᑐ㇟ᆅᇦ࠾ࡼࡧࢧࣥࣉࣝᩘ
㏆␥ᆅ᪉࡛ 2009 ᖺ࠿ࡽ 2013 ᖺ࡟ᤕ⋓ࡉࢀࡓ࢖ࣀࢩࢩ 33 㢌ࠊࢩ࢝ 25 㢌ࠊ࠾ࡼࡧ୰
ᅜᆅ᪉࡛ᤕ⋓ࡉࢀࡓ࢖ࣀࢩࢩ 63 㢌ࡼࡾᚓࡓ⾑Ύࢆ⏝࠸ࡓࠋ-(9 ࡟ᑐࡍࡿᢠయࡢ᳨ฟ
JEV/sw/Chiba/88/2002 ᰴࢆ⏝࠸࡚ 80㸣ࣉ࣮ࣛࢡῶᩘヨ㦂࡟ࡼࡾ୰࿴ᢠయ౯ࢆ

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ࡋࡓࠋ10 ಸ௨ୖࡢᢠయ౯ࢆᬻᐃⓗ࡟㝧ᛶ࡜ุᐃࡋࡓࠋ
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ࠊ୰ᅜᆅ᪉ࡢ
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ᅗ  ᪥ᮏ⬻⅖࢘࢖ࣝࢫ㸦-(9㸧ࡢឤᰁ⎔࡟࠾ࡅࡿ㔝⏕ື≀ࡢᙺ๭ࡲ࡜ࡵ
ᡃࠎࡣࠊ㔝⏕ື≀࡛ࡢ⾑Ύ␿Ꮫㄪᰝࢆᐜ᫆࠿ࡘ≉␗ⓗ࡟ᐇ᪋ࡍࡿࡇ࡜ࢆ┠ⓗ࡜ࡋ࡚ࠊᇵ㣴⣽⬊࡟࡚Ⓨ⌧ࡋࡓࢱࣥࣃࢡ㉁ࢆᢠཎ࡜ࡋ࡚ࠊProteinA/G ࢆ⏝࠸ࡓ ELISA ⣔ࡢᵓ⠏࡟ᡂຌࡋࡓࠋࡉࡽ࡟ᡃࠎࡢ␿Ꮫㄪᰝ࡟ࡼࡾࠊE ᆺ⫢⅖ࠊSFTSࠊ࣮࢚࢜ࢫ࣮࢟⑓ࠊ᪥ᮏ
⬻⅖ࡢཎᅉ࢘࢖ࣝࢫࡣࠊ࢖ࣀࢩࢩࡀಖ᭷ࡋ࡚࠸ࡿྍ⬟ᛶࡀ㧗ࡃࠊࣄࢺ࠾ࡼࡧࣈࢱࢆྵࡴ
ᵝࠎ࡞ື≀࡟ᑐࡋ࡚ࠊ࢖ࣀࢩࢩࡀࡇࢀࡽࡢឤᰁ⑕ࡢឤᰁ※࡜࡞ࡿࡇ࡜ࡀ♧၀ࡉࢀࡓࠋࡲ
ࡓࠊࡇࢀࡽࡢឤᰁ⑕ࡢឤᰁ⤒㊰ࡣࠊᦤ㣗ࠊ྾⾑᪻⹸࡟ࡼࡿ፹௓ࠊ㣕ἓ࡞࡝ᵝࠎ࡛࠶ࡿࠋ
ࡇࢀࡽࡢ⤖ᯝ࠿ࡽࠊ㔝⏕ື≀࡜᥋ゐࡍࡿᶵ఍ࡀከ࠸ࠊ⊁⊟ࢆࡍࡿ᪉ࠎࠊ㔝⏕ື≀ࢆ㣗⫗࡜ࡋ࡚฼⏝ࡉࢀࡿ᪉ࠎࠊ㣴㇜ᴗ⪅ࡢ᪉ࠎ࡟ᑐࡋࠊ௒ᅇㄪᰝࡋࡓឤᰁ⑕ࡢឤᰁࢆண㜵ࡍࡿ᪉ἲ࡜ࡋ࡚ࠊ௨ୗࡢὀព஦㡯ࢆᥦゝࡍࡿࠋ㔝⏕ື≀⟶⌮࡟㛵ࢃࡿ᪉ཬࡧ࢖ࣀࢩࢩࡀ⏕ᜥࡍࡿᆅᇦ࡟࠾ఫ࠸ࡢ᪉࡬
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‫ࡓࡋ⏝౑ۑ‬ໟ୎ࡸࡲ࡞ᯈ࡞࡝ࡢ㐨ල㢮ࡶࠊᚲࡎ୎ᑀ࡟Ὑίࡍࡿ
‫⏕ࠊࡵྵࡶ≟⊟ۑ‬㣗ࡣཝ⚗࡛࠶ࡿࠋႚ㣗᫬࡟ࡣᚲࡎ༑ศຍ⇕ࡍࡿ‫ۑ‬㔝ᒣ࡟⾜ࡃ㝿ࡣࠊ⺅ࡸࢲࢽ࡞࡝ࡢ྾⾑᪻⹸࡟ࡼࡿ྾⾑ࢆண㜵ࡍࡿࡓࡵ࡟ࠊ㛗⿇㺃㛗ࢬ࣎ࣥࡢ╔⏝ࠊ⹸㝖ࡅ࡞࡝ࢆᐇ᪋ࡍࡿ
‫࡜ࢱࣈࡣࢩࢩࣀ࢖ۑ‬ඹ㏻ឤᰁࡍࡿ⑓ཎయࢆಖ᭷ࡋ࡚࠸ࡿࡓࡵࠊ㣫⫱ሙෆ࡬ࡢ࢖
ࣀࢩࢩࡢ౵ධࢆ㜵Ṇࡍࡿ 

රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-8

ࡲࡓࠊ௒ᅇㄪᰝࡋࡓ࢘࢖ࣝࢫឤᰁ⑕௨እ࡟ࡶࠊ࢖ࣀࢩࢩ➼㔝⏕ື≀ࡀឤᰁ※࡜࡞ࡿឤᰁ⑕ࡀᏑᅾࡍࡿࡇ࡜ࡸࠊᮍࡔⓎぢࡉࢀ࡞࠸࢘࢖ࣝࢫࢆ㔝⏕ື≀ࡀಖ᭷ࡋ࡚࠸ࡿྍ⬟ᛶࡀ࠶ࡿࡇ࡜࠿ࡽࡶࠊୖグࡢ஦㡯࡟ࡣࠊᖖ࡟Ẽࢆ௜ࡅ࡚㡬ࡁࡓ࠸ࠋᘬ⏝ᩥ⊩
Hara, Y., Terada, Y., Yonemitsu, K., Shimoda, H., Noguchi, K., Suzuki, K. and Maeda,
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ຍ ྂ ዉ ⥴ ⨾ ࣭ 㕥 ᮌ Ύ ♧ ࣭ ᮡ Ụ ඾ ᫎ 㸬 2011 㸬 ⦾ Ṫ ࿴ ∵ ࡟ ࠾ ࡅ ࡿ ᪥ ᮏ ⬻ ⅖ Ⓨ ⏕ ஦ 㸬
http://www.pref.aichi.jp/cmsfiles/contents/0000049/49901/13.pdf
2014.3.10 accessed.
∦ᒣ㈗ᚿ㸬2013㸬Ꮚ∵ࡢ኱⬻࡟࠾ࡅࡿ᪥ᮏ⬻⅖࢘࢖ࣝࢫ࡟ࡼࡿ㠀໬⮋ᛶ㧊⭷⬻⅖. ື⾨◊◊✲ሗ࿌. 119:37-45.
ᖾ⏣▱Ꮚ࣭ᰗᮏ῟Ꮚ࣭᯽ཎ ⿱࣭௚㸬2000㸬≉㞟 㔝⏕ື≀ࡢឤᰁ⑕㸦2㸧㔝⏕࢖ࣀࢩࢩࡢ
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᳃ᕝ ⱱ࣭Ᏹ⏣ᬗᙪ࣭ຍ᮶⩏ᾈ࣭ᮌᮧᫀఙ࣭௒ᒸᾈ୍࣭⚟ኈ⚽ᝋ࣭ྜྷἙᬛᇛ࣭㇂ⱥᶞ࣭ୗ
ᓥᫀᖾ࣭Ᏻ⸨⚽஧࣭すᲄᨻᖾ࣭⃝㎶ிᏊ࣭ᕝ➃ᐶᶞ࣭᪂಴ ⥤࣭๓⏣ ೺࣭㧗㔝 ឡ࣭ᰗ஭ᚨ☻࣭⸨⏣༤ᕫ࣭㧗⏣ఙᘯ㸬2013㸬㸺㏿ሗ㸼㔜⑕⇕ᛶ⾑ᑠᯈῶᑡ⑕ೃ⩌㸦SFTS㸧
࢘࢖ࣝࢫࡢᅜෆศᕸㄪᰝ⤖ᯝ㸦➨୍ሗ㸧㸬ᅜ❧ឤᰁ⑕◊✲ᡤ㸪⑓ཎᚤ⏕≀᳨ฟ᝟ሗ㸬
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රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-9➨ 㸷 ❶            ࢽ࣍ࣥ࢖ࣀࢩࢩࡢෆ㒊ᐤ⏕⹸బ⸨ ᏹ 1࣭ᡞ⏣ṇᯞ 1࣭M. A. Omar1࣭㎷ ▱㤶 2࣭ᶓᒣ┿ᘪ 2,3
せ Ⅼ࣭ රᗜ┴ୗ࡛཰㞟ࡋࡓࢽ࣍ࣥ࢖ࣀࢩࢩ 132 㢌ࢆᑐ㇟࡜ࡋ࡚ࠊࡑࡢෆ㒊ᐤ⏕⹸㸦⽸⹸㸧ࢆ⥙⨶ⓗ࡟ㄪ࡭ࡓࠋ࣭ ᑐ㇟ື≀࡟⥺⹸ 22 ✀࡜᮲⹸ 1 ✀ࠊ྾⹸ 1 ✀ࡢᐤ⏕ࡀ☜ㄆࡉࢀࡓࠋ࿴ḷᒣ┴⏘ࢽ࣍ࣥ࢖ࣀࢩࢩࢆㄪ࡭ࡓሗ࿌㸦Sato et al. 2008㸧࡛ࡣ⥺⹸ 18 ✀࡜᮲⹸ 1 ✀ࡀグ㘓ࡉࢀ࡚࠸ࡿࡀࠊ⥺⹸࡟ࡘ࠸࡚ࡣ᭷ṑ㇜⫶⹸ Ascarops dentataࠊ⣚Ⰽẟᵝ⥺⹸Hyostrongylus rubidusࠊ⨾㯇㣗㐨⹸ Gongylonema pulchrum ࡀᮏᐟ୺࠿ࡽึグ㘓࡜࡞ࡗࡓࡇ࡜ࠊࡲࡓࠊ㸯✀࡜⪃࠼࡚ࡁࡓẟ⣽⥺⹸ࡀ 3 ✀࡟෌ศ㢮ࡉࢀࡓࡇ࡜ࡀ
୧ᆅᇦ࡛ࡢᐤ⏕⹸┦ࡢ㐪࠸ࡢ኱ࡁ࡞せᅉ࡛࠶ࡿࠋࡲࡓࠊ྾⹸࡟ࡘ࠸࡚ࡣ༢Ⅽ⏕Ṫᆺ⫢⻄㸦࠸ࢃࡺࡿ᪥ᮏ⏘⫢⻄ aspermic Fasciola sp.㸧ࡢᐤ⏕ࡶᅜෆ࡛ึࡵ࡚☜ㄆࡉࢀࡓࠋ࣭ ㇜⭠⤖⠇⹸ Oesophagostomum dentatum ࡣ࿴ḷᒣ┴⏘࢖ࣀࢩࢩ࡛ࡣ㧗࠸ᐤ⏕⋡
㸦55%㸧࡛࠶ࡗࡓࡀࠊරᗜ┴⏘ಶయ࡛ࡣࡈࡃ⛥࡞ᐤ⏕࡛࠶ࡗࡓࡇ࡜㸦3.8%㸧ࠊ࿴ḷᒣ┴⏘࢖ࣀࢩࢩ࡛ࡣᐤ⏕ࡀ☜ㄆࡉࢀ࡞࠿ࡗࡓ᭷ṑ㇜⫶⹸ࡢឤᰁಶయࡀ㜰⚄ᆅᇦ࡟㞟୰ࡋ࡚࠸ࡿࡇ࡜࡞࡝ࠊ࢖ࣀࢩࢩಶయ⩌࡟ࡼࡾ≉᭷ࡢᐤ⏕⹸┦ࢆࡶࡘࡇ࡜ࡀ♧၀ࡉࢀࡓࠋ࣭ ே⋇ඹ㏻ᐤ⏕⹸⑕ࡢཎᅉ࡜࡞ࡿࢻࣟࣞࢫ㢡ཱྀ⹸ࡢᐤ⏕⋡ࡣරᗜ┴ᮏᕞ㒊࡛ࡣᆅᇦࢆၥࢃࡎ࡟㧗࠿ࡗࡓࡀࠊಶయ࠶ࡓࡾࡢᐤ⏕ᩘࡣᴫࡋ࡚ᑡ࡞࠿ࡗࡓࠋ࢖ࣀࢩࢩ࠿ࡽ
┤᥋ឤᰁࡍࡿࡇ࡜ࡣ࡞࠸ࡀࠊᮏ✀ࡢศᕸᆅ࡛࠶ࡿࡇ࡜ࢆ⌮ゎࡋࠊຍ⇕୙༑ศ࡞⊦⫗࠶ࡿ࠸ࡣῐỈ㨶ࡢ⏕㣗➼࡛ឤᰁࡋ࡚⓶⭵∐⾜⑕࡜࡞ࡿ༴㝤ᛶ࡟␃ពࡍࡿᚲせࡀ࠶ࡿࠋKey words㸸ᐤ⏕⽸⹸┦ࠊරᗜ┴ࠊ⏕≀ከᵝᛶࠊ␿Ꮫࡣࡌࡵ࡟
ᮏ㑥࡟ࡣࢽ࣍ࣥ࢖ࣀࢩࢩ㸦Sus scrofa leucomystax㸧࡜ࣜࣗ࢘࢟ࣗ࢘࢖ࣀࢩࢩ㸦Sus

scrofa riukiuanus㸧ࡀศᕸࡋ㸦Watanobe et al. 1999㸧ࠊࡑࡢ␗࡞ࡿ⏕≀ᆅ⌮Ꮫⓗ఩⨨࡙ࡅࡶ࠶ࡗ࡚ࠊࡑࢀࡒࢀࡢᐤ⏕⹸┦࡟ࡣ≉Ⰽࡀ࠶ࡿ㸦Shoho & Machida 1979; ෆ⏣࡯ᒣཱྀ኱Ꮫඹྠ⋇་Ꮫ㒊⮫ᗋ⋇་Ꮫㅮᗙ⋇་ᐤ⏕⹸⑓Ꮫศ㔝࣭ රᗜ┴᳃ᯘື≀◊✲ࢭࣥࢱ࣮࣭ රᗜ┴❧኱Ꮫ⮬↛࣭⎔ቃ⛉Ꮫ◊✲ᡤ 

රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-9

࠿ 1984; Sato et al. 2008㸧ࠋ୧⪅࡟ඹ㏻ࡍࡿᐤ⏕⹸࡜ࡋ࡚ࠊ㇜⫵⹸ Metastrongylus spp. 
3 ✀ࠊ㢮෇㇜⫶⹸ Ascarops strongylinaࠊࢻࣟࣞࢫ㢡ཱྀ⹸ Gnathostoma doloresi㺂Globo-

cephalus samoensisࠊ࢖ࣀࢩࢩ⭠⤖⠇⹸ Oesophagostomum watanabei ࡀ࠶ࡆࡽࢀࡿ㸦Yamaguti 1961; Noda 1973; Chinose et al. 1984; ෆ⏣࡯࠿ 1984; Morita et al.
2007㸧ࠋḢᕞ࡛ࡣࠊ㏆ᖺࠊே⋇ඹ㏻ᐤ⏕⹸⑕ࡢཎᅉ࡜࡞ࡿ᪕ẟ⹸ Trichinella spp.ࡢಖ
⹸ᐟ୺࡜ࡋ࡚࣮ࣚࣟࢵࣃ࢖ࣀࢩࢩ㸦Sus scrofa subspp.㸧ࡢ㔜せᛶࡀㄆ㆑ࡉࢀࠊຍ⇕୙༑ศ࡞⊦⫗ࡢႚ㣗ࢆ௓ࡋࡓ᪕ẟ⹸⑕ࡢ㞟ᅋឤᰁ஦౛ࡢⓎ⏕ࡀ▱ࡽࢀ࡚࠸ࡿ
㸦Pérez-Martín et al. 2000; Gari-Toussaint et al. 2005㸧ࠋ୍᪉ࠊᅜෆศᕸࡢ࢖ࣀࢩࢩ
࠿ࡽࡢ᪕ẟ⹸ឤᰁࡢグ㘓ࡣࡇࢀࡲ࡛ࡢ࡜ࡇࢁ࡞࠸ࡀࠊࢻࣟࣞࢫ㢡ཱྀ⹸ࠊ Onchocerca

dewittei ࠊ ࢘ ࢙ ࢫ ࢸ ࣝ ࣐ ࣥ ⫵ ྾ ⹸ Paragonimus westermani ࠊ ኱ 㖋 㢌 ⹸
Macracanthorhynchus hirudinaceus ࡢ ಖ ⹸ ᐟ ୺ ࡜ ࡋ ࡚ ࠊ ࡲ ࡓ ࠊ ࢺ ࢟ ࢯ ࣉ ࣛ ࢬ ࣐
Toxoplasma gondii ➼ࡢே⋇ඹ㏻ឤᰁᛶཎ⹸ࡢಖ⹸ᐟ୺࡜ࡋ࡚ࠊබ⾗⾨⏕ୖࡢὀពࡀᡶ
ࢃࢀࡡࡤ࡞ࡽ࡞࠸㸦Miyazaki & Habe 1976; Kamiya et al. 1977; Miyazaki et al. 1978;ᮧ⏣ 1988; Nawa & Imai 1989; Uni et al. 2001; Takaoka et al. 2005㸧
ࠋSato et al. 㸦2008㸧ࡣࠊ2005㹼2006 ᖺࡢ⊁⊟ᮇ࡟࿴ḷᒣ┴ෆ࡛཰㞟ࡉࢀࡓ 20 㢌ࠊරᗜ┴ෆ࡛཰㞟ࡉࢀࡓ 9 㢌ࢆ᳨ᰝᑐ㇟࡜ࡋ࡚ࠊࢽ࣍ࣥ࢖ࣀࢩࢩࡢෆ㒊ᐤ⏕⽸⹸ࢆヲ⣽࡟ㄪ࡭ࠊ⥺⹸ 18 ✀࡜᮲⹸㸯✀ࠊィ 19 ✀ࢆሗ࿌ࡋ࡚࠸ࡿࠋ୧ᆅᇦ㛫࡛ᐤ⏕⹸┦࡟኱ࡁ࡞㐪࠸ࡣ࡞࠸ࡀࠊ኱⭠ᐤ⏕ࡢ㇜⭠⤖⠇⹸ Oesophagostomum dentatum ࡀරᗜ┴⏘ಶయ࠿ࡽࡣ☜ㄆࡀ࡞ࡃࠊ࿴ḷᒣ┴⏘ಶయ࡛ࡣ 55㸣ࡢಶయ࡛☜ㄆࡉࢀࡓࡇ࡜࠿ࡽࠊ㏆㞄ᆅᇦ࡟ࡶ㛵ࢃࡽࡎ୍㒊࡛ࡣ␗࡞ࡿᐤ⏕⹸ศᕸࡀࡳࡽࢀࡿྍ⬟ᛶࡀ♧၀ࡉࢀ࡚࠸ࡿࠋᮏ◊✲࡛ࡣࠊරᗜ┴ୗ࠿ࡽࡢ࢖ࣀࢩࢩ᳨ᰝᩘࢆ඘ᐇࡉࡏࠊ࿴ḷᒣ┴ୗࡢ࢖ࣀࢩࢩࡢᐤ⏕≧ἣ࡜ࡢẚ㍑ࠊ୪ࡧ࡟ྠ୍┴ෆ࡛␗࡞ࡿᆅᇦ࡟⏕ᜥࡍࡿಶయ⩌࡛ࡢᐤ⏕≧ἣࡢ≉ᚩ࡙ࡅࢆヨࡳࡓࠋᮦᩱ࡜᪉ἲ
᳨ᰝᮦᩱ
2007 ᖺ 8 ᭶ 8 ᪥࠿ࡽ 2010 ᖺ 12 ᭶ 8 ᪥࡟ࢃࡓࡾࠊරᗜ┴ୗ࡛୺࡜ࡋ࡚᭷ᐖᤕ⋓ࡉࢀࡓࢽ࣍ࣥ࢖ࣀࢩࢩ 132 㢌ࢆ཰㞟ࡋࡓࠋణࡋࠊ2007 ᖺࡣ 1 㢌ࠊ2008 ᖺࡣ 6 㢌࡛࠶ࡾࠊ୺యࡣ 2009 ᖺ 5 ᭶࠿ࡽ 2010 ᖺ 12 ᭶࡟࠿ࡅ࡚ᤕ⋓ࡉࢀࡓಶయ࡛࠶ࡿࠋ᭶ู࡟ࡳࡿ࡜ࠊ
1 ᭶ 9 㢌ࠊ2 ᭶ 11 㢌ࠊ3 ᭶ 2 㢌ࠊ4 ᭶ 0 㢌ࠊ5 ᭶ 11 㢌ࠊ6 ᭶ 12 㢌ࠊ7 ᭶ 8 㢌ࠊ8 ᭶ 10
㢌ࠊ9 ᭶ 23 㢌ࠊ10 ᭶ 12 㢌ࠊ11 ᭶ 19 㢌ࠊ12 ᭶ 13 㢌࡛࠶ࡗࡓࠋᆅᇦẚ㍑ࢆ⾜࠺ࡓࡵࠊᤕ⋓ࡉࢀࡓᆅᇦࡈ࡜࡟㜰⚄ᆅᇦࠊ୹Ἴᆅᇦࠊణ㤿ᆅᇦࡢ 3 ༊ศ࡟ᆅᇦศࡅࢆ⾜ࡗࡓ㸦⾲㸯㸧ࠋᤕ⋓ࡉࢀࡓ࢖ࣀࢩࢩࡣẅฎศᚋࡍࡳࡸ࠿࡟ゎయࡋࠊႃဗ㢌㒊࠿ࡽ┤⭠࡟࠿ࡅ࡚ࡢ
࿧྾ჾ⣔⮚ჾ㸦Ẽ⟶࣭⫵㸧ࠊᚰ⮚ࠊᾘ໬⟶㸦㣗㐨ࠥ┤⭠㸧ࠊ⫢⮚ࢆྲྀࡾฟࡋࡓࠋࡇࢀࡽෆ
⮚ࡣරᗜ┴᳃ᯘື≀◊✲ࢭࣥࢱ࣮࡟ᣢࡕᖐࡾࠊ㸫20C ࡛෭෾ಖᏑࡋࠊ㡰ḟࠊᒣཱྀ኱Ꮫ
ᐤ⏕⹸Ꮫ◊✲ᐊ࡟෭෾≧ែ࡛⛣㏦ࡋࡓࠋ࡞࠾ࠊ⭈⮚࡞ࡽࡧ࡟ࡑࡢ࿘ᅖ⬡⫫⤌⧊ࡣ࢖ࣀࢩ
ࢩࡢᰤ㣴Ꮫⓗศᯒ࡟౪ࡋࡓࡓࡵࠊ┤᥋࡟ࡣᐤ⏕⹸Ꮫⓗ᳨ᰝࡣ⾜ࡗ࡚࠸࡞࠸ࠋ 

රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-9

ᐤ⏕⹸Ꮫⓗ᳨ᰝ
෭෾ಖᏑࡉࢀࡓෆ⮚ࡣᐊ࡟࡚ゎ෾ࡋࠊ

⮚ჾู࡟ᐤ⏕⹸Ꮫⓗ᳨ᰝࢆᐇ᪋ࡋࡓࠋࡍ
࡞ࢃࡕࠊᾘ໬⟶ࡣ⦪࡟㛤ࡁࠊࢢ࣮ࣟࣈࢆ
ࡣࡵࡓᡭᣦ࡛⢓⭷㠃ࢆࡼࡃ᧿㐣ࡋ࡚ࠊỈ㐨Ỉ࡟ෆᐜ≀࡜⢓⭷ࢆࡼࡃὙ࠸ⴠ࡜ࡋࡓࠋ
┠㛤ࡁ 5.6mm ࡢ⢒࠸ࢫࢸࣥࣞࢫ〇⠠࠸㸦ᮾிࢫࢡ࣮ࣜࣥᰴᘧ఍♫ࠊᮾி㒔༓௦⏣༊㘫෬⏫㸧ࢆ㏻ࡋ࡚኱ᆺࡢࢦ࣑ࢆ㝖ཤࡋࠊ ṧࡾࡢᠱ⃮ᾮࢆ┠㛤ࡁ 300μm ࡢࢫࢸࣥࣞࢫ⠠࠸㸦ᮾிࢫࢡ࣮ࣜࣥᰴᘧ఍♫㸧ୖ࡛Ỉ㐨Ỉ࡟࡚ࡼࡃὙ࠸ࠊᚤ⣽࡞ࢦ࣑ࢆ㝖ཤࡋࡓࠋ⠠࠸ୖ࡟ṧࡗࡓෆᐜ≀ࢆỈ㐨Ỉ࡟ᡠࡋࠊࡇࢀࢆᐇయ㢧ᚤ㙾ୗ࡛⢭ᰝࡋࡓࠋࡲࡓࠊ〄㢌᮲⹸ࠊᅇ⹸ࡸ㢡ཱྀ⹸ࠊ኱⭠ᐤ⏕ྛ✀⥺⹸➼ࠊ⫗║ⓗ࡟☜ㄆ࡛ࡁࡿᐤ⏕⹸ࡣࠊ☜ㄆࡢ㒔ᗘࠊ㐺ᐅᅇ཰ࡋࡓࠋ
᳨ฟࡉࢀࡓᐤ⏕⹸ࡣ⏕⌮㣗ሷỈ࡟ࡲࡎᅇ཰ࡋࠊࡑࡢᚋࠊ10%୰ᛶ⦆⾪࣐࣍ࣝࣜࣥᾮ
࠶ࡿ࠸ࡣ 70%࢚ࢱࣀ࣮ࣝ࡟࡚ᅛᐃ࣭ಖᏑࡋࡓࠋࡲࡓࠊ୍㒊ࡢ⹸య࡟ࡘ࠸࡚ࡣࠊ㑇ఏᏊ᳨ᰝ࡟౪ࡍࡿࡓࡵ࡟㸫20C ࡛෭෾ಖᏑࡋࡓࠋᙧែᏛⓗゎᯒࡣࠊඛࡢሗ࿌㸦Sato et al.
2008㸧࡜ྠᵝ࡟ࠊᖖἲ࡟ᚑ࠸ᐇ᪋ࡋࡓࠋィ್ࡣᖹᆒ್±ᶆ‽೫ᕪ࡛♧ࡋࡓࠋ࡞࠾ࠊ

୍㒊ࡢᐤ⏕⹸ࡢᙧែᏛⓗ≉ᚩ࡟ࡘ࠸࡚ࡣబ⸨㸦2007㸧࡟ᅗゎࡋ࡚࠸ࡿࠋ
ศᏊ㑇ఏᏛⓗゎᯒ࢚ࢵ࣌ࣥࢻࣝࢳ࣮ࣗࣈ࡟ධࢀ㸫20C ࡛෭෾ಖᏑࡋ࡚࠸ࡓᐤ⏕⹸࡟ࡘ࠸࡚ࠊIllustrae

tissue & cells genomicPrep Mini Spin Kit 㸦GE Healthcare UK, Buckinghamshire, UK㸧ࢆ⏝࠸࡚ DNA ᢳฟࢆ⾜ࡗࡓࠋᡭᢏ࡟ࡘ࠸࡚ࡣࠊ࢟ࢵࢺ࡟௜ᖏࡍࡿㄝ᫂᭩
࡟ࡋࡓࡀࡗࡓࠋ᰾ DNA ࡢࣜ࣎ࢯ࣮࣒ RNA 㑇ఏᏊ㸦rDNA㸧࡞ࡽࡧ࡟࣑ࢺࢥࣥࢻࣜ࢔
DNA ࡢࢳࢺࢡ࣮࣒ࣟ c ࢜࢟ࢩࢲ࣮ࢮ࣭ࢧࣈࣘࢽࢵࢺ I 㸦 cox-1 㸧ࡢ PCR ቑᖜࡣ
Makouloutou et al.㸦2013㸧ࡢ㏻ࡾ࡟ᐇ᪋ࡋࡓࠋ࡞࠾ࠊ⏝࠸ࡓ DNA ࣏࣓࣮ࣜࣛࢮࡣ
BlendTaq-Plus-㸦TOYOBO; ኱㜰ᕷ໭༊㸧࡛ࠊ཯ᛂࡣ㸶㐃 PCR ࢳ࣮ࣗࣈ࡟ DNA ᢳฟᾮ 1μl ࢆྵࡴ཯ᛂᾮ 20μl ࢆධࢀ࡚⾜ࡗࡓࠋPCR ⏘≀ࡣ Ethidium bromide ྵ᭷ 1.3%
࢔࣮࢞ࣟࢫࢤࣝ࡟࡚㟁ẼὋືࡋࠊࢺࣛࣥࢫ࢖࣑ࣝࢿ࣮ࢱ࡛ྍど໬ࡋࠊࣂࣥࢻ㛗ࢆࢳ࢙ࢵࢡࡋࡓࠋ┠ⓗ࡟㐺࠺ PCR ⏘≀ࡣ FastGene Gel/PCR Extraction Kit㸦NIPPON
Genetics Co., Tokyo, Japan㸧ࢆ⏝࠸࡚⢭〇ࡋࢩ࣮ࢡ࢚ࣥࢫࢆ⾜ࡗࡓࠋ㔜」ࡍࡿࢭࢢ࣓
ࣥࢺ࡜ࡋ࡚ࢩ࣮ࢡ࢚ࣥࢫࡉࢀࡓ rDNA ࢩ࣮ࢡ࢚ࣥࢫࡣࠊCLUSTAL W multiplealignment programme㸦Thompson et al. 1994㸧ࢆ᥼⏝ࡋ࡚⹸యẖ࡟ 1 ᮏ࡟ࡋࡓࠋ 

රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-9

⤖ᯝරᗜ┴ୗ࡛཰㞟ࡋࡓ࢖ࣀࢩࢩ 132 㢌ࡢ⬚⭍࡞ࡽࡧ࡟⭡⭍⮚ჾ࠿ࡽ 24 ✀ࡢᐤ⏕⽸⹸ࢆ
᳨ฟࡋࡓ㸦⾲㸰㸧ࠋᐟ୺ࡢᤕ⋓᫬ᮇࡸᛶู࡟ࡼࡾឤᰁ⋡ࡀ㢧ⴭ࡟␗࡞ࡿᐤ⏕⹸✀ࡣ࡞࠸ࡀࠊ⏘ᆅࢆణ㤿ᆅᇦ㸦17 㢌㸧ࠊ୹Ἴᆅᇦ㸦47 㢌㸧ࠊ㜰⚄ᆅᇦ㸦53 㢌㸧࡜༊ษࡿ࡜ࠊᆅᇦ
࡟ࡼࡾ᳨ฟ⋡࡟኱ࡁ࡞ᕪࡀࡳࡽࢀࡿ✀ࡀ࠶ࡗࡓࠋࡍ࡞ࢃࡕࠊ୹Ἴᆅᇦࡢ࢖ࣀࢩࢩ࡛ࡣ௒
ᅇ᳨ฟࡉࢀࡓࡍ࡭࡚ࡢ⽸⹸✀ࡢᐤ⏕ࡀ࠶ࡗࡓࡀࠊ༢Ⅽ⏕Ṫᆺ⫢⻄ aspermic Fasciola sp.
㸦ᅗ㸯; ᕷᕝ࣭ᯈᇉ 2013㸧ࠊ㇜⭈⹸ Stephanurus dentatusࠊ⨾㯇㣗㐨⹸ Gongylonema

pulchrum㸦ᅗ㸰㸧ࡣణ㤿ᆅᇦࠊ㜰⚄ᆅᇦ࠿ࡽࡢ᳨ฟࡣ࡞࠸࠿ࠊ࠶ࡿ࠸ࡣࡈࡃ⛥࡛࠶ࡗࡓࠋ୍᪉ࠊ᭷ṑ㇜⫶⹸ Ascarops dentata㸦ᅗ㸱㸧ࡣ୺࡟㜰⚄ᆅᇦ࡛ᤕ⋓ࡉࢀࡓ࢖ࣀࢩ
ࢩ࠿ࡽ㧗⋡࡟᳨ฟࡉࢀࠊ௚ᆅᇦ࡛ࡢ᳨ฟࡣࡈࡃ㝈ࡽࢀࡓࠋ㜰⚄ᆅᇦ࡛ࡣࠊ᪥ᮏ〄㢌᮲⹸

Pseudanoplocephala nipponensis㸦ᅗ㸲㸧ࡢឤᰁ⋡ࡶ㧗ࡃࠊࡲࡓࠊ௚ᆅᇦ࡛ࡣ 64 㢌୰
1 㢌࠿ࡽࡢࡳ᳨ฟࡉࢀࡓ㇜⭠⤖⠇⹸ࡀࠊ㜰⚄ᆅᇦ 53 㢌୰ 4 㢌࠿ࡽ☜ㄆࡉࢀࡓࠋࡲࡓࠊ
ᓥᕋ㒊ࡢῐ㊰ᓥ࠿ࡽࡢ᳨ᰝಶయᩘࡣ 4 㢌࡟㝈ࡽࢀࠊᐤ⏕⹸᳨ฟࡣ 5 ✀ࡢࡳ࡛࠶ࡗࡓࠋࡲ
ࡓࠊ㔝⏕ಶయࢆᤕ⋓ࡋࠊ1 ᖺ௨ୖ࡟ࢃࡓࡾ㣫⫱ࡋ࡚࠸ࡓ 11 㢌࡛ࡶࠊ㔝⏕ಶయ࡛ࡣ㧗⋡࡟ᐤ⏕ࡍࡿ⫵⹸ࡢᐤ⏕ࡀࡳࡽࢀࡎࠊࢻࣟࣞࢫ㢡ཱྀ⹸ࢆࡣࡌࡵ඲⯡ⓗ࡟ᐤ⏕⋡ࡶప࠸ഴྥ
ࡀࡳࡽࢀࡓࠋ
⨾㯇㣗㐨⹸ࡢᅜෆ࢖ࣀࢩࢩ࡛ࡢᐤ⏕ࡀ☜ㄆࡉࢀࡓࡢࡣ௒ᅇࡢරᗜ┴࡛ࡢㄪᰝࡀึࡵ
࡚࡛࠶ࡗࡓࡇ࡜࠿ࡽࠊ18S rDNA ࡞ࡽࡧ࡟ cox-1 ሷᇶ㓄ิ࡟ࡘ࠸᳨࡚ウࡋࡓࠋ࣍ࣥࢩࣗ
࢘ࢪ࢝㸦Cervus nippon㸧ࡸࢽ࣍ࣥࢨࣝ㸦Macaca fuscata㸧࡞࡝ᅜෆ㔝⏕ື≀࡟ᐤ⏕ࡋ࡚࠸ࡿ⣔⤫࡜ᐙ␆࡛࠶ࡿ࢘ࢩᐤ⏕ࡢ⣔⤫࡜ࡀ༊ูࡉࢀ࡚࠸ࡿࡀ㸦బ⸨ 2009㸧ࠊ௒ᅇ࢖ࣀࢩࢩ࠿ࡽᚓࡓ⣔⤫ࡣ๓⪅࡟ᒓࡋ࡚࠸ࡓࠋヲࡋࡃࡣࠊูࡢㄽᩥ࡛ሗ࿌ࡋ࡚࠸ࡿ
㸦Makouloutou et al. 2013㸧ࠋ⾲㸰࡟࠾࠸࡚ࠊẼ⟶ᨭෆᐤ⏕ࡢ⫵⹸ Metastrongylus spp.ࡣ୍ᣓࡋࡓࡀࠊࡇࢀ࡟ࡣ

M. elongatusࠊM. salmiࠊM. asymmetricusࠊM. pudendotechus ࡢ 4 ✀ࡀྵࡲࢀ࡚࠸
ࡿࠋࡲࡓࠊ⫶⢓⭷ࡸ⢓ᾮ୰࡟ᐤ⏕ࡋ࡚࠸ࡿẟ⣽⥺⹸⛉⥺⹸ Capillarid spp.࡟ࡘ࠸࡚ࡶ୍
ᣓࡋࡓࡀࠊࡇࢀ࡟ࡣ Aonchotheca suisࠊAonchoteca riukiuensis䚸Aonchotheca sp. ࡢ
3 ✀ࡀ༊ูࡉࢀࡓ㸦ᡞ⏣࣭బ⸨ 2010㸧ࠋ௒ᅇࡢ᳨ᰝᑐ㇟ಶయࡢ୰࠿ࡽẟ⣽⥺⹸ࡀከᩘᐤ
⏕ࡋ࡚࠸ࡓ 14 㢌ࢆ㑅ࡧࠊA. suisࠊA. riukiuensis䚸Aonchotheca sp.ࡢᐤ⏕⋡ࢆࡳࡿ࡜ࡑࢀࡒࢀ 64.3%ࠊ78.6%ࠊ14.3%࡛࠶ࡗࡓࠋࡇࢀࡽ 3 ✀࡟ࡘ࠸࡚ࠊ௨ୗ࡟ᙧែᏛⓗ≉ᚩࢆグࡍࠋ  㻝㻡㻥㻟㻥㻞㻣㻻㼑㼟㼛㼜㼔㼍㼓㼛㼟㼠㼛㼙㼡㼙㻌㼐㼑㼚㼠㼍㼠㼡㼙㼀㼞㼕㼏㼔㼡㼞㼕㼟㻌㼟㼡㼕㼟㻝㻜㻥㻼㼟㼑㼡㼐㼍㼚㼛㼜㼘㼛㼏㼑㼜㼔㼍㼘㼡㼟㻌㼚㼕㼜㼜㼛㼚㼑㼚㼟㼕㼟㻟㻥㻤㻢㻡㻿㼠㼞㼛㼚㼓㼥㼘㼛㼕㼐㼑㼟㻌㼞㼍㼚㼟㼛㼙㼕㻻㼑㼟㼛㼜㼔㼍㼓㼛㼟㼠㼛㼙㼡㼙㻌㼣㼍㼠㼍㼚㼍㼎㼑㼕㻞㻠㻳㼘㼛㼎㼛㼏㼑㼜㼔㼍㼘㼡㼟㻌㼘㼛㼚㼓㼑㼙㼡㼏㼞㼛㼚㼍㼠㼡㼟㻮㼡㼞㼓㼑㼘㼍㼠㼕㼍㻌㼐㼕㼐㼡㼏㼠㼍㻝㻞㻤㻡㻣㻳㼘㼛㼎㼛㼏㼑㼜㼔㼍㼘㼡㼟㻌㼟㼍㼙㼛㼑㼚㼟㼕㼟㻞㻝㻠㻭㼟㼏㼍㼞㼕㼟㻌㼟㼡㼡㼙㻯㼍㼜㼕㼘㼘㼍㼞㼕㼐㻌㼟㼜㼜㻚㻜㻜㻟㻌㻔㻢㻜㻚㻜㻕

㻠㻌㻔㻤㻜㻚㻜㻕

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㻟㻌㻔㻢㻜㻚㻜㻕㻜㻜㻜㻟㻌㻔㻢㻜㻚㻜㻕

䗢㻌㻔㻡㻕㻝㻌㻔㻠㻚㻤㻕
㻣㻌㻔㻟㻟㻚㻟㻕

㻝㻌㻔㻤㻚㻟㻕

㻝㻡㻌㻔㻣㻝㻚㻠㻕㻝㻤㻌㻔㻤㻡㻚㻣

㻝㻌㻔㻠㻚㻤㻕

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㻝㻣㻌㻔㻤㻝㻚㻜㻕㻣㻌㻔㻟㻟㻚㻟㻕

㻥㻌㻔㻠㻞㻚㻥㻕

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㻝㻌㻔㻠㻚㻤㻕

㻝㻞㻌㻔㻡㻣㻚㻝㻕㻝㻡㻌㻔㻣㻝㻚㻠㻕㻝㻜㻌㻔㻠㻣㻚㻢㻕㻝㻌㻔㻠㻚㻤㻕

㻡㻌㻔㻞㻟㻚㻤㻕

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㻝㻡㻌㻔㻣㻝㻚㻠㻕䗢㻌㻔㻞㻝㻕

㻝㻡㻌㻔㻡㻣㻚㻣㻕㻜㻞㻞㻌㻔㻤㻠㻚㻢㻕㻞㻡㻌㻔㻥㻢㻚㻞㻕㻝㻌㻔㻟㻚㻤㻕

㻞㻌㻔㻣㻚㻣㻕

㻢㻌㻔㻞㻟㻚㻝㻕

㻞㻢㻌㻔㻝㻜㻜㻕

㻝㻡㻌㻔㻡㻣㻚㻣㻕㻝㻢㻌㻔㻢㻝㻚㻡㻕㻜㻟㻌㻔㻝㻝㻚㻡㻕

㻝㻢㻌㻔㻢㻝㻚㻡㻕㻞㻝㻌㻔㻤㻜㻚㻤㻕㻝㻤㻌㻔㻢㻥㻚㻞㻕㻠㻌㻔㻝㻡㻚㻠㻕

㻠㻌㻔㻝㻡㻚㻠㻕

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㻞㻠㻌㻔㻥㻞㻚㻟㻕䗠㻌㻔㻞㻢㻕

┴୰㒊㻜㻣㻌㻔㻡㻤㻚㻟㻕

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㻝㻜㻌㻔㻤㻟㻚㻟㻕㻣㻌㻔㻡㻤㻚㻟㻕㻜㻜㻜㻢㻌㻔㻡㻜㻚㻜㻕

䗠㻌㻔㻝㻞㻕

┴໭㒊㻖㻌ឤᰁಶయᩘ㻔ឤᰁ䠂㻕䜢䛭䜜䛮䜜♧䛩㻚㻖㻖㻌㻹㻚㻌㼑㼘㼛㼚㼓㼍㼠㼡㼟㻘㻌㻹㻚㻌㼟㼍㼘㼙㼕㻘㻌㻹㻚㻌㼍㼟㼥㼙㼙㼑㼠㼞㼕㼏㼡㼟㻘㻌㻹㻚㻌㼜㼡㼐㼑㼚㼐㼛㼠㼑㼏㼔㼡㼟 㻌䠐✀䜢ಶู䛻༊ู䛧䛶䛔䛺䛔㻚

኱⭠

ᑠ⭠

㻡㻝㻞

㻭㼟㼏㼍㼏㼞㼛㼜㼟㻌㼐㼑㼚㼠㼍㼠㼍㻝㻠㻞㻥㻭㼟㼏㼍㼞㼛㼜㼟㻌㼟㼠㼞㼛㼚㼓㼥㼘㼕㼚㼍㻴㼥㼛㼟㼠㼞㼛㼚㼓㼥㼘㼡㼟㻌㼞㼡㼎㼕㼐㼡㼟㻝㻝㻝㻢㻤㻼㼔㼥㼟㼛㼏㼑㼜㼔㼍㼘㼡㼟㻌㼟㼑㼤㼍㼘㼡㼠㼡㼟

㻠

㻞㻜

㻞

㻞㻠㻣

᭱኱ᅇ཰
⹸యᩘ

㻳㼚㼍㼠㼔㼛㼟㼠㼛㼙㼍㻌㼐㼛㼘㼛㼞㼑㼟㼕㻳㼛㼚㼓㼥㼘㼛㼚㼑㼙㼍㻌㼜㼡㼘㼏㼔㼞㼡㼙㣗㐨㻌㻔⢓⭷ෆ㻕㻿㼠㼑㼜㼔㼍㼚㼡㼞㼡㼟㻌㼐㼑㼚㼠㼍㼠㼡㼟⭈⮚䞉࿘ᅖ⬡⫫⤌⧊㻭㼟㼜㼑㼞㼙㼕㼏㻌 㻲㼍㼟㼏㼕㼛㼘㼍 㻌㼟㼜㻚⫹⟶㻔⫢⮚㻕

⫶✀ྡ㻹㼑㼠㼍㼟㼠㼞㼛㼚㼓㼥㼘㼡㼟 㻌㼟㼜㼜㻚㻖㻖Ẽ⟶ᨭ㻌㻔⫵㻕⮚ჾ㻟㻌㻔㻥㻚㻣㻕

㻟㻌㻔㻥㻚㻣㻕

㻞㻝㻌㻔㻢㻣㻚㻣㻕㻞㻢㻌㻔㻤㻟㻚㻥㻕㻥㻌㻔㻞㻥㻚㻜㻕

㻠㻌㻔㻝㻞㻚㻥㻕

㻝㻢㻌㻔㻡㻝㻚㻢㻕㻞㻤㻌㻔㻥㻜㻚㻟㻕㻤㻌㻔㻞㻡㻚㻤㻕

㻞㻡㻌㻔㻤㻜㻚㻢㻕㻜㻞㻝㻌㻔㻢㻣㻚㻣㻕㻞㻥㻌㻔㻥㻟㻚㻡㻕㻝㻣㻌㻔㻡㻠㻚㻤㻕㻝㻤㻌㻔㻡㻤㻚㻝㻕㻝㻌㻔㻟㻚㻞㻕

㻜㻜㻞㻝㻌㻔㻢㻣㻚㻣㻕䗢㻌㻔㻟㻝㻕

㻞㻌㻔㻥㻚㻝㻕

㻝㻌㻔㻠㻚㻡㻕

㻝㻡㻌㻔㻢㻤㻚㻞㻕㻝㻣㻌㻔㻣㻣㻚㻟㻕㻤㻌㻔㻟㻢㻚㻠㻕

㻟㻌㻔㻝㻟㻚㻢㻕

㻣㻌㻔㻟㻝㻚㻤㻕

㻝㻠㻌㻔㻢㻟㻚㻢㻕㻞㻌㻔㻥㻚㻝㻕

㻝㻟㻌㻔㻡㻥㻚㻝㻕㻝㻌㻔㻠㻚㻡㻕

㻝㻝㻌㻔㻡㻜㻚㻜㻕㻝㻢㻌㻔㻣㻞㻚㻣㻕㻥㻌㻔㻠㻜㻚㻥㻕

㻝㻞㻌㻔㻡㻠㻚㻡㻕㻜㻝㻌㻔㻠㻚㻡㻕

㻜㻝㻞㻌㻔㻡㻠㻚㻡㻕䗠㻌㻔㻞㻞㻕

┴༡㒊㻝㻌㻔㻞㻡㻚㻜㻕

㻜㻞㻌㻔㻡㻜㻚㻜㻕

㻟㻌㻔㻣㻡㻚㻜㻕

㻜㻜㻝㻌㻔㻞㻡㻚㻜㻕

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㻜

䗢㻔㻟㻕㻛䗠㻔㻝㻕ῐ㊰ᓥ㻝㻌㻔㻝㻢㻚㻣㻕

㻜㻞㻌㻔㻟㻟㻚㻟㻕

㻞㻌㻔㻟㻟㻚㻟㻕

㻟㻌㻔㻡㻜㻚㻜㻕

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㻜㻠㻌㻔㻢㻢㻚㻣㻕

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䗠㻌㻔㻡㻕

㣫⫱ಶయ㻌㻔┴୰㒊㻕⏕ά⎔ቃࡢởᰁࡸ୰㛫ᐟ୺࡜࡞ࡿື᳜≀ࡢႚ㣗ࢆ௓ࡋ࡚ே⋇ඹ㏻ᐤ⏕⹸ࡢཎᅉ࡜࡞ࡿྍ⬟ᛶࢆᣢࡘᐤ⏕⹸࡟ࡘ࠸࡚ࡣୗ⥺ࢆᘬ࠸ࡓࠋ⾲㸰 රᗜ┴⏘ࢽ࣍ࣥ࢖ࣀࢩࢩ࠿ࡽ᳨ฟࡉࢀࡓᐤ⏕⽸⹸ රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-9ᅗ㸯㸬࢖ࣀࢩࢩ⫹⟶࡟ᐤ⏕ࡍ ࡿ ༢ Ⅽ ⏕ Ṫ ᆺ ⫢ ⻄
DVSHUPLF)DVFLRODVS㸦ⓑ▮༳㸧௒ᅇࡢㄪᰝ࡛ࡣࡶࡗࡥࡽ୹Ἴ
ᆅᇦࡢ࢖ࣀࢩࢩ㸦 㢌୰  㢌㸧᳨࡛ฟࡉࢀࡓࠋ⫹⟶ቨࡢ⫧ཌࡀ㢧ⴭ࡛࠶ࡿࠋ࣐࣍ࣝࣜࣥᅛ
ᐃᶆᮏ㸦:㸧ࠋᅗ㸰㸬࢖ࣀࢩࢩ㣗㐨⢓⭷ෆ࡟ࢪࢢࢨࢢࢆᥥ࠸࡚ᐤ⏕ࡍ ࡿ ⨾ 㯇 㣗 㐨 ⹸*RQJ\ORQHPDSXOFKUXP 㞘㸦 ࡘࡢ▮༳࡛  ⹸యࢆ♧ࡍ㸧௒ᅇࡢㄪᰝ࡛ࡣ୹Ἴᆅᇦࡢ࢖
ࣀࢩࢩ࡛㧗⋡࡟᳨ฟࡉࢀࡓ㸦  㢌 ୰  㢌 㸧ࠋ ⏕ ᶆ ᮏ
㸦:㸧ࠋᅗ㸱㸬࢖ࣀࢩࢩ⫶ෆ࡟ᐤ⏕ࡍࡿ᭷ṑ㇜⫶⹸ $VFDURSVGHQWDWD௒ᅇࡢㄪᰝ࡛ࡣࠊ㜰⚄ᆅᇦ࡛㧗⋡࡟᳨ฟࡉࢀ㸦 㢌୰ 㢌㸧ࠊణ㤿ᆅᇦ㸦 㢌㸧࡛ࡣឤᰁಶయ࡞ࡋࠊ୹Ἴᆅᇦ࡛ࡣ 㢌୰  㢌ࡢࡳ࡟࡜࡝ࡲࡗ
ࡓࠋᮏ౛࡛ࡣᐤ⏕ᩘ  㞘ࢆᩘ࠼ࡓࠋ⏕ᶆᮏ㸦:㸧ࠋ 

රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-9ᅗ㸲㸬࢖ࣀࢩࢩᑠ⭠࡟ᐤ⏕ࡍࡿ᪥ᮏഇ〄㢌᮲⹸

3VHXGDQRSORFHSKDODQLSSRQHQVLV ୗࡢ෗┿ࡣ 㞘ࡢ㢌⠇㸦▮༳㸧࠿ࡽཷ⫾∦⠇࡟⮳ࡿࢫࢺࣟࣅࣛࢆ♧ࡍࠋ௒ᅇࡢㄪᰝ࡛ࡣࠊ㜰⚄ᆅᇦ࡛㢖⦾࡟᳨ฟࡉࢀ㸦 㢌୰ 㢌㸧ࠊణ㤿࣭୹Ἴᆅᇦ࡛ࡣ 㢌୰  㢌ࡢࡳ࡟࡜࡝ࡲࡗࡓࠋ
ᮏ౛࡛ࡣᐤ⏕ᩘ  㞘ࢆᩘ࠼ࡓࠋ⏕ᶆᮏ㸦:㸧ࠋ Aonchotheca suis㸦Yamaguti 1943㸧 㞝⹸㸦n=8㸧㸸య㛗 11.89㸫13.53㸦13.02 ± 0.01㸧mmࠊ㣗㐨㒊ࡢ㛗ࡉ 4.60㸫5.89㸦5.16± 0.43㸧mmࠊ᭱኱యᖜ 0.054㸫0.074㸦0.067 ± 0.007㸧mmࠋ㣗㐨㒊㸸⭠⟶㒊ࡢẚࡣ 1㸸
1.30㸫1.69㸦1.53 ± 0.14㸧ࠋᑿ➃ࡢᕥྑ୧ഃ⭡㠃ᐤࡾ࡟ࡣ๓ᚋ 2 ᑐࡢ✺㉳ࡀ⏕ࡌࠊࡸࡸ๓᪉࡬ྥ࠿࠺㸦ᅗ㸳㸧ࠋ๓ഃࡢ 1 ᑐࡣ༢⣧࡞ᣦ≧࡛ࠊᚋࢁഃࡢ 1 ᑐࡣඛ➃ࡀ๓᪉࡬ᒅ᭤
ࡋ࡚ࣁ࣐࣮ࣥ≧ࢆ࿊ࡍࡿࠋࡇࡢ✺㉳࡟ࠊࡼࡃⓎ㐩ࡋࡓᑿഃ⩼ࡢ㐲఩➃࠾ࡼࡧᑿ➃⫼ഃࡢᑿᄞࡀ௜╔ࡋ࡚࠸ࡿࠋ஺᥋่ࡣ⥺≧࡛୰➼ᗘ࡟◳໬ࡋࠊ㛗ࡉࡣ 0.75㸫0.82㸦0.79 ± 0.02㸧
mmࠊ㐲఩➃ࡣࣛࣥࢭࢵࢺ≧ࢆ࿊ࡍࡿࠋᑿഃ⩼࡜஺᥋่㠧࡟ࡣᶓ࡟⣽ࡃ⥺᮲ࡀධࡾࠊ஺
᥋่㠧ࡣ↓Ჲ࡛࠶ࡿࠋ 㞤⹸㸦n=23㸧㸸య㛗 13.32㸫22.77㸦18.65 ± 2.33㸧mmࠊ㣗㐨㒊ࡢ㛗ࡉ 4.88㸫6.44㸦5.87± 0.37㸧mmࠊ᭱኱యᖜ 0.063㸫0.100㸦0.083 ± 0.010㸧mmࠋ㣗㐨㒊㸸⭠⟶㒊ࡢẚࡣ 1㸸
1.64㸫2.93㸦2.18 ±0.34㸧ࠋᑿ➃ࡢᙧ≧ࡣ㕌෇≧㹼෇㗹≧࡛࠶ࡿࠋ㝜㛛ࡣ㣗㐨࣭⭠⟶⤖ྜ㒊ࡢ 0.044㸫0.203㸦0.139 ± 0.041㸧mm ᚋ᪉࡟఩⨨ࡋࠊ㝜㛛㒊࡟┠❧ࡗࡓ✺㉳ࡸ௜ᒓ≀ࡣ࡞࠸ࡀࠊ㛤ཱྀ㒊๓ᚋࡢ⭡㠃య⾲࡟ᐇ㉁ᛶ࠾ࡼࡧࢡࢳࢡࣛᛶࡢ⣽࠿࡞พฝࡀ࠶ࡾࠊ⢒⣁࡞⾲㠃ᵓ㐀ࢆ࿊ࡍࡿ㸦ᅗ㸳㸧ࠋ⹸༸ࡣ㛗ᚄࡀ 0.050㸫0.070㸦0.063 ± 0.003㸧mmࠊ▷ᚄࡀ 0.024㸫0.033㸦0.028 ± 0.002㸧mm ࡛ࠊ༸Ẇࡀࡸࡸཌࡃ୧ᴟࡢ⵹ࡣ✺ฟࡋࠊ༸Ẇ⾲㠃ࡣᖹ⁥࡛≉Ṧ࡞ᵓ㐀ࡸ⣠ᵝࡣぢࡽࢀ࡞࠸ࠋ 

රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-9ᅗ㸳㸬࢖ࣀࢩࢩࡢ⫶࡟ᐤ⏕ࡍࡿẟ⣽⥺ᅗ㸴㸬࢖ࣀࢩࢩࡢ⫶࡟ᐤ⏕ࡍࡿẟ⣽⥺⹸ $RQFKRWKHFDVXLV⹸ $RQFKRWKHFDULXNLXHQVLV㸦$㸧㞝⹸ᑿ㒊ࡢഃ㠃ീ࡛ࠊᑿ➃㏆ࡃࡢᕥྑ㸦$㸧㞝⹸ᑿ㒊ࡢഃ㠃ീ࡛ࠊᑿ➃ࡀࣁ࣐࣮ࣥ

ࡢᣦ≧✺㉳㸦▮༳㸧࡜ᑿഃ⩼㸦 㸧ࢆ♧ࡍࠋ≧✺㉳࡜࡞ࡗ࡚㢌ഃ࡟ᒅ᭤ࡍࡿ㸦 㸧ࠋ▮༳

㸦%㸧ᑿഃ⩼ࢆḞࡃ㞝⹸ࡢᑿ㒊ഃ㠃ീࠋ㸦&㸧ࡣ⥲᤼ἥཱྀ๓⦕࠿ࡽᑿഃ࡟ྥ࠿ࡗ࡚⣽㛗࠸㞤⹸ࡢ㝜㛛㸦▮㢌㸧࿘ᅖࡢഃ㠃ീࠋ㣗㐨ᮎᣦ≧✺㉳㸦▮༳㸧ࡀࡳࡽࢀࡿࠋ㸦%㸧㞝⹸ᑿ➃㒊㸦 㸧࡟㏆ࡃࠊ㝜㛛ࡀ㛤ཱྀࡍࡿࡀࠊࡑࡢ

㒊ࡢ⭡㠃ീ࡛ࠊᕥྑ࡟ࡳࡽࢀࡿ⣽㛗࠸ᣦ≧࿘ᅖࡢ⭡㠃ࡣࡁࢃࡵ࡚⢒㐀࡛ࠊ୙つ๎࡞ฝ✺㉳ࢆ♧ࡍࠋ㸦&㸧㞤⹸ࡢ㝜㛛㸦▮㢌㸧࿘ᅖพ≧࡛࠶ࡿࠋ෗┿ $ࠥ&ࠊࡍ࡭࡚ྠ୍ಸ⋡ࠋࡢഃ㠃ീࠋ㣗㐨ᮎ➃㒊㸦 㸧࡟㏆ࡃࠊ㝜㛛ࡀ㛤ཱྀࡋࠊ࣋ࣝ≧ࡢ௜ᒓ≀ࡀ࠶ࡿࠋ෗┿ $ࠥ&ࠊࡍ࡭࡚ྠ୍ಸ⋡ࠋ

㸦2㸧Aonchotheca riukiuensis㸦Shoho & Machida 1979㸧 㞝⹸㸦n=9㸧㸸య㛗 7.29㸫13.75㸦11.47 ± 2.04㸧mmࠊ㣗㐨㒊ࡢ㛗ࡉ 3.59㸫5.18㸦4.59± 0.50㸧mmࠊ᭱኱యᖜ 0.038㸫0.067㸦0.054 ± 0.009㸧mmࠋ㣗㐨㒊㸸⭠⟶㒊ࡢẚࡣ 1㸸
1.03㸫1.82㸦1.49 ± 0.30㸧ࠋయᚋ➃ࡣ 1 ᑐࡢࣁ࣐࣮ࣥ≧✺㉳࡜࡞ࡗ࡚⭡ഃ࡟ྥ࠿࠸ࠊࡑ
ࡢ㛫࡟ࡣὸ࠸⿣ࡀධࡿࠋᑿ➃⫼ഃ࡟ࡣᣦ≧✺㉳࡟ᨭᣢࡉࢀ࡚ࡼࡃⓎ㐩ࡋࡓᑿᄞࡀ࠶ࡾࠊ
ࡇࡢ㎶⦕ࡣ⢒࡛ࠊ⾲㠃࡟ࡣ⣽࠸⥺ࡀධࡗ࡚⦰⦉ᵝࢆ࿊ࡍࡿࡶࡢࡀ࠶ࡿࠋࡲࡓࠊ⫠㛛㛤ཱྀ㒊๓⦕࠿ࡽ 1 ᑐࡢ⣽࠸✺㉳ࡀ⏕ࡌ࡚ᚋ᪉࡬ྥ࠿࠺ࠋᑿ㒊ࡢᕥྑ୧ഃ࡟ࡣࠊࡈࡃᖜࡢ⊃࠸ࢡࢳࢡࣛᛶࡢᑿഃ⩼ࡀᏑᅾࡍࡿ㸦ᅗ㸴㸧ࠋ஺᥋่ࡣ⥺≧࡛ࡑࡢ㐲఩㒊ࡣ༢ㄪ࡟⣽ࡃ࡞ࡾࠊ 

රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-9

㛗ࡉࡣ 0.71㸫1.01㸦0.86 ± 0.09㸧mmࠋ஺᥋่㠧ࡣ↓Ჲ࡛ᶓ࡟⥺᮲ࡀධࡿࠋ 㞤⹸㸦n=34㸧
㸸య㛗 13.92㸫36.11㸦27.83 ± 4.70㸧mmࠊ㣗㐨㒊ࡢ㛗ࡉ 4.38㸫6.27 㸦5.24± 0.45㸧mmࠊ᭱኱యᖜ 0.044㸫0.085㸦0.069 ± 0.009㸧mmࠋ㣗㐨㒊㸸⭠⟶㒊ࡢẚࡣ 1㸸
2.18㸫5.14㸦4.29 ± 0.69㸧ࠋᑿ➃ࡢᙧ≧ࡣ㕌෇≧㹼෇㗹≧࡛࠶ࡿࠋ㝜㛛ࡣ㣗㐨࣭⭠⟶⤖ྜ㒊ࡢ 0㸫0.236㸦0.104 ± 0.041㸧mm ᚋ᪉࡟㛤ཱྀࡍࡿࠋ㝜㛛㛤ཱྀ㒊࡟ࡣ࣋ࣝ≧ࡢࢡࢳࢡࣛᛶࡢ⟄ࡀ௜ᒓࡋࠊ๓᪉ࢆྥࡃ㸦ᅗ㸴㸧ࠋ⹸༸ࡣ㛗ᚄࡀ 0.049㸫0.064㸦0.059 ± 0.002㸧
mmࠊ▷ᚄࡀ 0.020㸫0.033㸦0.027 ± 0.002㸧mm ࡛ࠊ༸Ẇ⾲㠃࡟ࡣ࣐ࢫࢡ࣓ࣟࣥᵝࡢ⥙┠ᵓ㐀ࢆㄆࡵࡿࠋ
㸦3㸧Aonchotheca sp.
 㞝⹸㸦n=2㸧
㸸య㛗 6.90㸫7.59mmࠊ㣗㐨㒊ࡢ㛗ࡉ 3.21㸫3.70mmࠊ᭱኱యᖜ 0.050㸫
0.061mmࠋ㣗㐨㒊㸸⭠⟶㒊ࡢẚࡣ 1㸸1.05㸫1.15ࠋయᚋ➃ࡣᕥྑ 2 ࡘࡢ✺㉳࡜࡞ࡗ࡚⭡ഃ࡬ᒅ᭤ࡋࠊࡑࡢඛ➃ࡣᑠ⤖⠇≧ࡲࡓࡣ㕌࡞ᣦ≧✺㉳࡟⤊ࢃࡿࠋᑿ➃⫼ഃ࡟ࡣⓎ㐩ࡢᝏ࠸ᑿᄞࡀࠊᑿ㒊ࡢᕥྑ୧ഃ࡟ࡣⓎ㐩ࡋࡓᑿഃ⩼ࡀᏑᅾࡋࠊᣦ≧✺㉳ࡀࡇࢀࡽࢆᨭᣢࡍࡿ
㸦ᅗ㸵㸧ࠋ஺᥋่ࡣ୰➼ᗘ࡟◳໬ࡋ㐲఩㒊ࡣࣛࣥࢭࢵࢺ≧ࠊ㛗ࡉࡣ 0.39㸫0.46mmࠋ஺᥋่㠧ࡣ↓Ჲ࡛ᶓ࡟⥺᮲ࡀධࡿࠋ 㞤⹸㸦n=6㸧
㸸య㛗 8.96㸫9.73㸦9.22 ± 0.35㸧mmࠊ㣗㐨㒊ࡢ㛗ࡉ 2.66㸫3.12㸦2.90 ±0.20㸧mmࠊ᭱኱యᖜ 0.053㸫0.061㸦0.056 ± 0.003㸧mmࠋ㣗㐨㒊㸸⭠⟶㒊ࡢẚࡣ 1㸸
1.90㸫2.39㸦2.18 ± 0.18㸧ࠋᑿ➃ࡢᙧ≧ࡣ෇㗹≧㸦ᅗ㸵㸧ࠋ㝜㛛ࡣ㣗㐨࣭⭠⟶⤖ྜ㒊ࡢ
0.082㸫0.132㸦0.110 ± 0.018㸧mm ᚋ᪉࡟㛤ཱྀࡍࡿࠋ㝜㛛┤๓࡛ᐇ㉁ࡢ㝯㉳㒊ࢆࢡࢳࢡࣛࡀ⦕ྲྀࡾࠊ㝜㛛㛤ཱྀ㒊࡟ࡣࢡࢳࢡࣛᛶࡢ෇࠸௜ᒓ≀ࢆㄆࡵࡿ㸦ᅗ㸵㸧ࠋ⹸༸ࡣ㛗ᚄࡀ
0.054㸫0.066㸦0.059 ± 0.003㸧mmࠊ▷ᚄࡀ 0.024㸫0.027㸦0.025 ± 0.001㸧mm ࡛ࠊ༸Ẇ⾲㠃࡟ࡣ㛗㍈᪉ྥ࡟㉮ࡿ⥙┠≧⥺᮲ࢆㄆࡵࡿ㸦ᅗ㸵㸧ࠋ㸺ഛ⪃㸼 Aonchotheca suis㸦Yamaguti 1943㸧ࡣࠊᒣཱྀ┴࠾ࡼࡧ࿴ḷᒣ┴ࡢ㔝⏕ࡢࢽ࣍ࣥ࢖ࣀ
ࢩࢩࡢ⫶࠿ࡽศ㞳ࡉࢀࠊCapillaria suis ࡜ࡋ࡚✀グ㍕ࡉࢀࡓ㸦Yamaguti 1943㸧ࠋ㞝⹸ࡢᑿ✺㉳ࡢ≉ᚩⓗ࡞ᙧ≧ࡸ㞤⹸ࡢ㝜㛛㒊࡟≉ู࡞௜ᒓᵓ㐀≀ࡀ࡞࠸ࡇ࡜ࠊ⹸༸ࡢ༸Ẇ⾲㠃ࡀᖹ⁥࡛࠶ࡿࡇ࡜࡞࡝ࠊ௒ᅇほᐹࡉࢀࡓ≉ᚩࡣཎグ㍕࡜ࡼࡃ୍⮴ࡍࡿࠋణࡋࠊ㞝⹸ࡢᑿഃ⩼࡟㛵ࡋ࡚ࡣཎグ㍕࡟୍ษグ㏙ࡀ࡞ࡃࠊཎグ㍕ࡢ௜ᅗ࡟ࡶᑿഃ⩼ࡣᥥ࠿ࢀ࡚࠸࡞࠸ࠋ
௒ᅇ᳨ฟࡋࡓ⹸య࡛ࡣṤ࡝ࡍ࡭࡚ࡀࡼࡃⓎ㐩ࡋࡓᑿഃ⩼ࢆ᭷ࡋ࡚࠸ࡓࠋᑿഃ⩼ࢆకࢃ࡞࠸⹸యࡣ࿴ḷᒣ┴⏘ࢽ࣍ࣥ࢖ࣀࢩࢩ 1 㢌࠿ࡽᅇ཰ࡉࢀࡓ㞝⹸ 21 㞘ࡢ࠺ࡕࡢ 2 㞘ࡢࡳ࡛࠶ࡗࡓ㸦ᅗ㸳B㸧ࠋᑿഃ⩼ࡢ᭷↓௨እࡢᙧែⓗ≉ᚩࡸィ್࡟ࡘ࠸࡚ࡣࠊࡇࡢ 2 ࢱ࢖ࣉࡢ㞝⹸ࡢ㛫࡛ᕪ␗ࡣぢࡽࢀ࡞࠿ࡗࡓࡇ࡜ࠊྠ᫬࡟ᅇ཰ࡋࡓ㞤⹸࡟ᙧែᏛⓗ࡞ኚ␗ࢆ≉࡟ᣦ᦬ࡍࡿࡇ࡜ࡣ࡛ࡁ࡞࠿ࡗࡓࡇ࡜࠿ࡽࠊᑿഃ⩼ࡢ᭷↓࡟㛵ࢃࡽࡎྠ୍✀ A. suis ࡜⪃࠼ࡽࢀࡓࠋ 

රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-9ᅗ㸵㸬࢖ࣀࢩࢩࡢ⫶࡟ᐤ⏕ࡍࡿẟ⣽⥺⹸ᮍグ㍕✀ $RQFKRWKHFDVS
㸦$㸧㞝⹸ᑿ㒊ࡢഃ㠃ീ࡛ࠊ஺᥋่㠧ࡀ⥲᤼ἥཱྀࡼࡾ⩻㌿⬺ฟࡋ࡚࠸ࡿࡇ࡜㸦▮༳㸧ࢆ♧ࡍࠋ㸦%㸧㞤⹸ࡢ㝜㛛㸦▮㢌㸧࿘ᅖࡢഃ㠃ീࠋ㣗㐨ᮎ➃㒊㸦 㸧࡜㝜㛛ࡢ୰㛫ࡢ⭡㠃࡟ࢡࢳࢡ࡛ࣛ⦕ྲྀࡽࢀࡓ㢧ⴭ࡞㝯㉳㸦▮༳㸧ࡀ࠶ࡾࠊࡲࡓࠊ㝜㛛ࡣ኱ࡁ࡞ࢻ࣮࣒ᆺࡢ✺ฟ≀ࢆక࠺ࠋ㸦&㸧㞤⹸ᑿ㒊ࡢഃ㠃ീࠋᑿ➃ࡣ෇㗹≧ࢆ࿊ࡋࠊ⫠㛛㸦▮㢌㸧ࡣᮎ➃㏆ࡃ࡟㛤ཱྀࡍࡿࠋ㸦'㸧㝜㛛㏆ࡃ࡛⏘ฟࡉࢀࡿ┤๓ࡢ⹸༸࡟ࡣࠊ㛗㍈᪉ྥ࡟㉮ࡿ⥺᮲ࡀ࠾஫࠸࡟஺ཫࡋࠊ୙つ๎࡞⥙┠ᶍᵝࢆぢࡏࡿࠋ෗┿ $ࠥ'ࠊࡍ࡭࡚ྠ୍ಸ⋡ࠋ Aonchotheca riukiuensis ࡣࠊす⾲ᓥࡢࣜࣗ࢘࢟ࣗ࢘࢖ࣀࢩࢩࡢ⫶࠿ࡽ᳨ฟࡉࢀ✀グ㍕ࡉࢀࡓ㸦Shoho & Machida 1979㸧ࠋཎグ㍕࡜௒ᅇࡢᅇ཰⹸యࡢᙧែᏛⓗ≉ᚩࡣࡼࡃ୍⮴ࡋࡓࠋ࡞࠾ཎグ㍕࡛ࡣ㞝⹸ࡢ⫠㛛㛤ཱྀ㒊┤๓ࡢ✺㉳ࢆṇ୰࡟୍ᮏ⏕ࡌࡿ࡜グ㘓ࡋ࡚࠸ࡿࡀࠊࡇࡢ✺㉳ࡀᐇ㝿࡟ࡣᕥྑᑐ㇟࡟ 1 ᑐᏑᅾࡍࡿࡇ࡜ࡀ௒ᅇࡢほᐹ࡛☜ㄆࡉࢀࡓࠋ ௒ᅇ✀ᮍྠᐃ࡜ࡋࡓ Aonchotheca sp.ࡣ㞝⹸ࡀⓎ㐩ࡋࡓᑿഃ⩼࡜ᑿᄞࠊ୰➼ᗘ࡟◳໬ࡋࡓ஺᥋่࡜↓Ჲࡢ஺᥋่㠧ࢆ᭷ࡋࠊ඾ᆺⓗ࡞ Aonchotheca ᒓࡢ≉ᚩࢆ♧ࡋࡓࠋᅇ཰
ࡋࡓ⹸యࡢከࡃࡀኚᛶࡂࡳ࡛ࠊ㞝⹸ࠊ㞤⹸࡜ࡶ࡟ほᐹ࡛ࡁࡓᶆᮏᩘࡀᑡ࡞࠿ࡗࡓࡀࠊග
Ꮫ㢧ᚤ㙾ୗ࡛ࡢᙧែほᐹࡢ⤖ᯝࠊࡇࡢ✀ࡣ㔝⏕⫗㣗ື≀ࡢ⫶࣭ᑠ⭠࠿ࡽ᳨ฟࡉࢀࡓᮍグ㍕ Aonchotheca ᒓẟ⣽⥺⹸࡟㓞ఝࡋ࡚࠸ࡓ㸦ᡞ⏣࣭బ⸨ 2011㸧ࠋᮏ✀࡜㔝⏕⫗㣗ື≀ࡢ Aonchotheca ᒓẟ⣽⥺⹸ᶆᮏࡢィ್ࡣࡼࡃ୍⮴ࡍࡿ࡜࡜ࡶ࡟ࠊ18S rDNA ሷᇶ㓄ิࡢ☜ㄆ࠿ࡽࡶ୧⪅ࡣྠ୍✀࡛࠶ࡾࠊࡲࡓࠊᙧែᏛⓗ࡟㢮ఝࡋ⫗㣗ື≀୍࡛⯡ⓗ࡞

Aonchotheca putorii ࡜ࡣ༊ูࡉࢀࡿࡇ࡜ࡀ☜ㄆࡉࢀ࡚࠸ࡿ㸦ᡞ⏣࣭బ⸨ 2011㸧ࠋ⪃ᐹ௒ᅇࠊරᗜ┴ෆ࡜࠸࠺㝈ࡽࢀࡓᆅᇦ࡞ࡀࡽࡶࠊ⏕ᜥ⎔ቃࡢ␗࡞ࡿᆅ༊࡛ᤕ⋓ࡉࢀࡓࢽ࣍ࣥ࢖ࣀࢩࢩ࡟ࡘ࠸࡚ᐤ⏕⹸ࡢฟ⌧㢖ᗘࢆẚ㍑ࡋࡓࠋణ㤿ᆅᇦࡣἙཱྀ࡟㏆࠸ᖹ㔝㒊ࡀᗈࡀࡾࠊࡑࡇ࡟ᒣ㔝ࡀ㏕ࡗ࡚࠸ࡿࠋ୹Ἴᆅᇦࡣࠊᒣ㔝࡟ᐩࢇࡔ⎔ቃ࡛࠶ࡾࠊ㜰⚄ᆅᇦࡣ㒔ᕷ໬ࡀ㐍ࡴ୰࡟ṧࡉࢀࡓ⮬↛⎔ቃࡀ࠶ࡿ࡞࡝⎔ቃ᮲௳ࡀ␗࡞ࡗ࡚࠸ࡿࠋࡲࡓࠊᓥᕋ㒊࡛ 

රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-9

࠶ࡿῐ㊰ᓥࡢ᳨ᰝ㢌ᩘࡣ௒ᅇ 4 㢌࡜㝈ࡽࢀ࡚࠸ࡿࡀࠊᮏᕞ㒊࡟ẚ࡭࡚ᐤ⏕⹸✀ࡀ㝈ࡽࢀࡿ࡜࠸࠺ண

㏻ࡾࠊᮏᕞ㒊࡛ࡣィ 24 ✀ࡢᐤ⏕⽸⹸ࡀ☜ㄆࡉࢀࡓࡢ࡟ᑐࡋ࡚ࠊῐ㊰ᓥ࡛ࡣ 5 ✀࡟࡜࡝ࡲࡗࡓࠋࡇࡢⅬࡣࠊ௒ᚋ࡟᳨యᩘࢆቑࡍࡇ࡜࡛ࠊࡼࡾṇ☜࡞ᐇែࡀᥗࡵࡿ
ࡇ࡜ࡀᮇᚅࡉࢀࡿࠋ㣫⫱ಶయ 11 㢌ࢆ௒ᅇࡢ᳨ウ࡟ຍ࠼ࡓࡢࡣࠊ㝈ᐃࡉࢀࡓ㣫⫱⎔ቃ࡛ᐤ⏕⽸⹸࡟ࡼࡿ⃰ཌởᰁࡀ㉳ࡇࡗ࡚࠸ࡿྍ⬟ᛶࢆ᳨ドࡍࡿࡓࡵ࡛࠶ࡗࡓࠋᐇ㝿࡟ࡣࠊ᳨ฟࡉࢀࡿᐤ⏕⹸✀ࡣ⮬↛ಶయ⩌࡟ẚ࡭࡚㝈ࡽࢀࠊࡲࡓࠊᐤ⏕ᩘࡶ≉ẁ࡟ከ࠸ࢃࡅ࡛ࡣ࡞
࠿ࡗࡓࠋᤕ⋓ᚋࡢ㣫⫱࡟࠶ࡓࡗ࡚ࠊ㥑⹸๣ࡢᢞ୚➼࡝ࡢࡼ࠺࡞⟶⌮ࡀ⾜ࢃࢀࡓࡢ࠿ࡣ୙࡛᫂࠶ࡿࠋす⾲ᓥ࠶ࡿ࠸ࡣ዆⨾኱ᓥࡢࣜࣗ࢘࢟ࣗ࢘࢖ࣀࢩࢩ࡟ࡘ࠸࡚ࡣ⽸⹸඲⯡ࡢᐤ⏕≧ἣ࡟ࡘ࠸࡚ሗ࿌ࡉࢀ࡚࠸ࡿࡀ㸦Shoho & Machida 1979; ෆ⏣࡯࠿ 1984㸧ࠊࢽ࣍ࣥ࢖ࣀࢩࢩ
࡟ࡘ࠸࡚ࡣ≉ᐃ✀࡟ὀ┠ࡋࡓᐤ⏕≧ἣㄪᰝࡀ࡯࡜ࢇ࡝࡛࠶ࡾࠊᮏᕞᆅᇦࡢ㔝⏕ಶయ⩌㛫ẚ㍑࡟ࡼࡿᐤ⏕⹸┦ࡢグ㘓ࡣ᭱㏆ࡲ࡛࡞࠿ࡗࡓࠋSato et al.㸦2008㸧ࡣࠊ࿴ḷᒣ┴⏘
20 㢌ࠊරᗜ┴⏘ 9 㢌࡟ࡘ࠸᳨࡚ウࡋࠊ⥺⹸ 17 ✀ࠊ᮲⹸ 1 ✀ࠊᶓ㝸⭷➽⫗࡟⿕ᄞࡍࡿናᆺ྾⹸ Pharyngostomum cordatum ࣓ࢱࢭࣝ࢝ࣜ࢔ࢆሗ࿌ࡋ࡚࠸ࡿࠋ࿴ḷᒣ┴⏘࢖ࣀ
ࢩࢩ 1 㢌ࡢ኱⭠࠿ࡽ᪂✀グ㍕ࡉࢀࡓ Morgascaridia kugii ࡣู࡜ࡋ࡚ࡶࠊ㇜⭠⤖⠇⹸ࡢᐤ⏕ࡀࠊ࿴ḷᒣ┴⏘࢖ࣀࢩࢩ࡛ࡣ 20 㢌୰ 11 㢌㸦55%㸧࡛࠶ࡗࡓࡢ࡟ᑐࡋࠊරᗜ┴⏘࢖
ࣀࢩࢩ 9 㢌࡛ࡣ᳨ฟࡉࢀ࡞࠿ࡗࡓࠋࡓࡔࡋරᗜ┴⏘࢖ࣀࢩࢩࡢᶆᮏᩘࡀᑡ࡞࠿ࡗࡓࡓࡵࠊ
ᙜ᫬ࡣᆅᇦᕪࡢ᳨ウࡀ༑ศ࡟࡛ࡁ࡞࠿ࡗࡓࠋ௒ᅇࠊරᗜ┴ࡢణ㤿ᆅᇦ㸦17 㢌㸧ࠊ୹Ἴᆅᇦ㸦47 㢌㸧
ࠊ㜰⚄ᆅᇦ㸦53 㢌㸧࡜඘ᐇࡋࡓᶆᮏᩘ᳨࡛ᰝࢆ㐍ࡵࡿࡇ࡜࡛ࠊᆅᇦᛶࢆ᳨
ウ࡛ࡁࡓࠋ㇜⭠⤖⠇⹸ࡣࠊ
ణ㤿ᆅᇦ࡛ࡣᐤ⏕ಶయࡀ࡞ࡃࠊ
୹Ἴᆅᇦ࡛ 47 㢌୰ 1 㢌
㸦2.1%㸧ࠊ㜰⚄ᆅᇦ࡛ࡶ 53 㢌୰ 4 㢌㸦7.5%㸧࡟࡜࡝ࡲࡾࠊරᗜ┴ୗࡢ࢖ࣀࢩࢩ࡛ࡣࡈࡃ⛥࡞ᐤ⏕⹸✀࡛࠶ࡿࡇ࡜ࡀ෌☜ㄆࡉࢀࡓࠋ୍᪉ࠊ๓ᅇࡢㄪᰝ㸦Sato et al. 2008㸧ࢆྵࡵࢽ࣍ࣥ
࢖ࣀࢩࢩ࠿ࡽࡣᮍ☜ㄆ࡛࠶ࡗࡓ᭷ṑ㇜⫶⹸ࡀࠊ㜰⚄ᆅᇦ࡟ศᕸࡍࡿಶయ࡛㧗⋡㸦60.4%㸧
࡟᳨ฟࡉࢀࡓࠋྠࡌ┴ෆ࡛ࡶࠊ୹Ἴᆅᇦ 8.5%ࠊణ㤿ᆅᇦ 0%࡛࠶ࡿࡇ࡜࠿ࡽࠊᴟࡵ࡚㝈ᒁⓗ࡞ศᕸ≧ἣ࡛࠶ࡿࠋ᭱㏆࡟࡞ࡗ࡚ࠊᒣཱྀ┴࡛ᤕ⋓ࡉࢀࡓ࢖ࣀࢩࢩ࠿ࡽࡶ᳨ฟ౛ࡀ」ᩘグ㘓ࡉࢀ࡚࠾ࡾ㸦ᮍබ⾲ࢹ࣮ࢱ㸧ࠊࢽ࣍ࣥ࢖ࣀࢩࢩ⮬యࡀ㐣ཤ࠿ࡽࡶࡗ࡚࠸ࡓᐤ⏕⹸✀࡛࠶ࡿྍ⬟ᛶࡶฟ࡚ࡁࡓࡀࠊ࡝ࡢࡼ࠺࡞⤒⦋࡛ࠊᆅ⌮ⓗ㐃⥆ᛶࢆḞ࠸ࡓ⌧ᅾࡢศᕸ࡜
࡞ࡗ࡚࠸ࡿࡢ࠿ࠊරᗜ┴ࠊᒣཱྀ┴௨እࡢᆅᇦ࡛ࡶㄪᰝࡀ㐍ࡴࡇ࡜࡟ࡼࡗ࡚ࠊࡑࡢ≉᭷࡞ศᕸࡢ⣒ཱྀࡀぢࡘ࠿ࡿࡇ࡜ࢆᮇᚅࡋࡓ࠸ࠋ
᭷ṑ㇜⫶⹸࡟ຍ࠼ࠊ௒ᅇࠊᅜෆࡢ࢖ࣀࢩࢩ࠿ࡽึࡵ࡚⨾㯇㣗㐨⹸ࡀ᳨ฟࡉࢀࡓࠋ࢖ࣛ
ࣥ⏘࢖ࣀࢩࢩ࡛ࡣ㧗⋡࡞ᐤ⏕ࡀሗ࿌ࡉࢀ㸦Eslami & Farsad-Hamdl 1992㸧ࠊࡲࡓࠊྜ
⾗ᅜࡢᐙ␆࡜ࡋ࡚㣫㣴ࡉࢀࡿࣈࢱࡸ㔝⏕໬ࣈࢱ࡛ࡶ㧗࠸ᐤ⏕⋡ࡀሗ࿌ࡉࢀ࡚࠸ࡿࡇ࡜
࠿ࡽ㸦Zinter & Migaki 1970; Smith et al. 1982㸧ࠊື≀✀࡜ࡋ࡚㧗࠸ឤᰁឤཷᛶࢆࡶࡘ
ࡇ࡜ࡀ᥎ࡉࢀࡿࠋࡋ࠿ࡋ࡞ࡀࡽࠊ௒ᅇࡢㄪᰝ࡛ࡣᮏᕞ㒊 117 㢌୰ 5 㢌㸦4.3%㸧࡟࡜࡝ࡲࡾࠊῐ㊰ᓥ⏘ 4 㢌࠿ࡽࡢ᳨ฟࡣ࡞࠿ࡗࡓࠋྜ⾗ᅜࡢ㔝⏕ࣈࢱࢆㄪ࡭ࡓ Coombs &Springer㸦1974㸧ࡣࠊ⯉࠿ࡽࡢ᳨ฟ⋡ࡀ㧗ࡃ㸦6 / 9㸧ࠊ㣗㐨࠿ࡽࡢ᳨ฟ⋡ࡣప࠸㸦1 / 10㸧ࡇ࡜ࢆሗ࿌ࡋ࡚࠸ࡿࠋ௒ᅇࡢ᳨ᰝࡣ㣗㐨⢓⭷࡟㝈ࡽࢀ࡚࠸ࡓࡇ࡜࠿ࡽࠊᐇ㝿ࡢᐤ⏕⋡ࡣ 

රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-9௒ᅇࡢ⤖ᯝ௨ୖ࡟㧗࠸ྍ⬟ᛶࡶ༑ศ࡟࠶ࡿࠋ࠶ࡿ࠸ࡣࠊ௒ᅇࡢ࢖ࣀࢩࢩᐤ⏕⹸ㄪᰝ࡜ྠ
᫬ᮇ࡟ᐇ᪋ࡋࡓ࣍ࣥࢩࣗ࢘ࢪ࢝ࡢㄪᰝ࡛ࡶಶయ⩌ࡶࡋࡃࡣ⏕ᜥᆅᇦ࡟ࡼࡾឤᰁ⋡࡟኱ࡁ࡞㐪࠸ࡀ࠶ࡗࡓࡇ࡜࠿ࡽ㸦బ⸨ 2009; Makouloutou et al. 2013㸧ࠊྠᵝࡢせᅉ࡛ࠊ௒ᅇ᳨ᰝࡋࡓ࢖ࣀࢩࢩ࡛ࡢឤᰁ⋡ࡣᐇ㝿࡟ప࠿ࡗࡓྍ⬟ᛶࡶ࠶ࡿࠋ⤖ᯝ࡛グࡋࡓࡼ࠺࡟ࠊ
Makouloutou et al.㸦2013㸧࡟࠾࠸࡚ศᏊ⣔⤫Ꮫⓗゎᯒࡢヲ⣽ࢆሗ࿌ࡋ࡚࠸ࡿࠋᅜෆࡢ
⨾㯇㣗㐨⹸ࡣ㣗⣅ᛶ⏥⹸ࢆ୰㛫ᐟ୺࡜ࡋ࡚ࠊ㔝⏕ື≀࡛࠶ࡿࢩ࢝ࡸ࢖ࣀࢩࢩࢆ⤊ᐟ୺࡜ࡍࡿ᳃ᯘᆺࡢ⏕ά⎔ࢆྲྀࡿ⣔⤫࡜ࠊ࢘ࢩࢆ⤊ᐟ୺࡜ࡍࡿᐙ␆ᆺࡢ⏕ά⎔ࢆྲྀࡿ⣔⤫ࡀ࠶ࡿࡇ࡜ࡀุ᫂ࡋ࡚ࡁ࡚࠸ࡿ㸦బ⸨ 2009㸧ࠋරᗜ┴ୗ࡛ࡢ㇜⭈⹸ࡢឤᰁ≧ἣ࡟ࡘ࠸࡚ࡣࠊ๓ሗ㸦Sato et al. 2008㸧࡛ࡣࡓࡲࡓࡲ
ឤᰁࡋ࡚࠸ࡓ࢖ࣀࢩࢩ 1 㢌ࢆ☜ㄆࡋࡓࡀࠊឤᰁ⋡࡜ࡋ࡚࡝ࡢࡼ࠺࡞≧ἣ࡟࠶ࡿࡢ࠿ࡣ᫂
☜࡟࡛ࡁ࡚࠸࡞࠿ࡗࡓࠋ௒ᅇࡶᐊෆ࡛ࡢ᳨ᰝࡣᐇ᪋ࡉࢀ࡚࠸࡞࠸ࡀࠊゎయ⌧ሙ࡛ࡢ⫗║࣭ゐデ࡛ࡢ☜ㄆ࡛ឤᰁࡢ᭷↓ࢆグ㘓ࡋࠊឤᰁಶయ࠿ࡽ⹸యࡢ཰㞟ࢆ⾜ࡗࡓࠋࡑࡢ⤖ᯝࠊ⾲㸰࡟♧ࡍࡼ࠺࡟୹Ἴᆅᇦ࡛ࡣ 47 㢌୰ 9 㢌㸦19.1%㸧ࡢᐤ⏕ࡀ࠶ࡗࡓࡶࡢࡢࠊ┴ෆࡢ௚ᆅᇦ࡟⏕ᜥࡍࡿಶయ⩌࡛ࡣ࡯࡜ࢇ࡝ឤᰁࡀ࡞࠿ࡗࡓࠋ࡞࠾ࠊ࿴ḷᒣ┴⏘ࡢ࢖ࣀࢩࢩ࡛
ࡣࡈࡃ୍⯡ⓗ࡞ᐤ⏕⹸࡛࠶ࡗࡓ㸦ឤᰁ⋡ 55%: Sato et al. 2008㸧ࠋᮏ✀ᗂ⹸ࡣయෆ࡟ධ
ࡗ࡚ 3 ᅇ⬺⓶ࡋࡓᚋ࡟⫢⮚࡟ධࡗ࡚ࠊࡑࡢᐇ㉁ෆ࡛Ⓨ⫱ࡋࠊ෌ࡧ⭡⭍ෆ࡟ฟ࡚࠿ࡽ᭱⤊ᐤ⏕㒊఩࡛࠶ࡿ⭈⮚ࡶࡋࡃࡣࡑࡢ࿘ᅖࡢ⬡⫫⤌⧊࡟⛣ືࡍࡿࠋ࿴ḷᒣ┴⏘࢖ࣀࢩࢩ࡛ࡣࠊ
⫢⮚࡟⹸㐨࡟㉳ᅉࡍࡿ⤖ྜ⤌⧊ቑ⏕ࡸ⭡⭷⅖࡜㛵㐃ࡋࡓ⥺⥔⣲ᛶ⫢⿕⭷⅖ࡀࡋࡤࡋࡤほᐹࡉࢀࡓࡀࠊරᗜ┴⏘࢖ࣀࢩࢩ࡟࠾࠸࡚ࡣࠊ㇜⭈⹸ᐤ⏕࡟క࠺⑓ኚࡣ࡯࡜ࢇ࡝☜ㄆࡉࢀ࡚࠸࡞࠸ࠋప࠸ឤᰁ⋡ࢆ཯ᫎࡋࡓゎ๗⫗║ᡤぢ࡛࠶ࡿ࡜⪃࠼ࡿ࡜ࠊ௒ᅇࡢᐤ⏕⹸ᅇ཰≧ἣ࡜ࡼࡃ➢ྜࡍࡿࠋ࢖ࣀࢩࢩࡢゎ๗࡟࠾࠸࡚ࠊ⫶ቨ࡟✸Ꮝࡋᐤ⏕㒊఩ࢆ⫧ཌࡉࡏࡿࢻࣟࣞࢫ㢡ཱྀ⹸ࡣ┠❧ࡘࠋᮏ✀ࡣࣈࢱ࣭࢖ࣀࢩࢩࢆᅛ᭷ᐟ୺࡜ࡋ࡚⫶ቨ࡟ᡂ⹸ࡀᐤ⏕ࡋࠊ➨ 1 ୰㛫ᐟ୺ࡣࢣࣥ
࣑ࢪࣥࢥࠊ➨ 2 ୰㛫ᐟ୺ࡣ୧⏕㢮㸦ࢧࣥࢩࣙ࢘࢘࢜ࠊ࢚࢝ࣝࠊ࢖ࣔࣜ➼㸧ࡸ㨶㢮ࠊᚅᶵᐟ୺ࡀ࣊ࣅ㢮࡛࠶ࡿ㸦ᐑᓮ࣭⸨ 1988㸧ࠋᐑᓮ┴ࢆ୰ᚰ࡟஑ᕞᆅ᪉࡛ࡣ῱ὶ㨶࣐࣓ࣖᩱ⌮ࢆཎᅉ࡜ࡋࡓே࡬ࡢឤᰁ౛㸦⓶⭵∐⾜⑕㸧ࡀሗ࿌ࡉࢀ࡚࠸ࡿ㸦㉥⩚ 1999㸧ࠋ௒ᅇࠊරᗜ┴ࡢᮏᕞ㒊࡛ࡣᆅᇦࢆၥࢃࡎ࡟㧗࠸ឤᰁ⋡ࡀ☜ㄆࡉࢀࡓࡇ࡜࠿ࡽࠊ࢖ࣀࢩࢩࡢ⏕ᜥᆅ࡟ࡣᗈࡃࢻࣟࣞࢫ㢡ཱྀ⹸ࡀศᕸࡋ࡚࠸ࡿࡇ࡜ࡀ᫂ࡽ࠿࡜࡞ࡗࡓࠋ➨ 2 ୰㛫ᐟ୺ࡸᚅᶵ
ᐟ୺ࢆႚ㣗ࡍࡿᶵ఍ࡀ࠶ࡿሙྜ࡟ࡣࠊࡑࡢຍ⇕ฎ⌮ࡀ୙༑ศ࡛࠶ࡗࡓሙྜࠊ⓶⭵∐⾜⑕ࡀᘬࡁ㉳ࡇࡉࢀࡿ༴㝤ᛶࡀ࠶ࡿࡇ࡜࡟␃ពࡍࡿᚲせᛶࡀ࠶ࡿࠋ
ᅜෆ㎰ᐙ࡛ࡢᐙ␆࡜ࡋ࡚ࡢࣈࢱࡢ㣫㣴ᙧែࡣ㏆ᖺ኱ࡁࡃኚ໬ࡋࠊᮏ◊✲࡛ᢅࡗࡓᐤ⏕⽸⹸ࡢከࡃࡀ㣴㇜⌧ሙ࡛┳㐣࡛ࡁ࡞࠸ၥ㢟࡛࠶ࡗࡓ㐣ཤࡀ㐲࠸᫇ヰ࡟࡞ࡗ࡚࠸ࡿࠋ⟶⌮ࡉࢀࡓࢥࣥࢡ࣮ࣜࢺᗋࡢ㣫⫱⯋࡛ࡢ㣫㣴ࡣࠊࡑࡢ⏕ά⎔⥔ᣢ࡟୰㛫ᐟ୺ࢆᚲせ࡜ࡍࡿᐤ⏕⽸⹸ࡸእ⏺࡛ࡢ⹸༸Ⓨ⫱ࡀᚲせ࡞ᐤ⏕⽸⹸ࡢឤᰁ㜵Ṇ⟇࡜࡞ࡗ࡚ࡁࡓࠋ࢜࢞ࢡࢬⓎ㓝㇜⯋ࡢᬑཬ࡟ࡼࡾ㇜㠴⹸ࡀ⮴࿨ⓗ࡞ឤᰁࢆᘬࡁ㉳ࡇࡋࡓࡼ࠺࡟㸦ᖹ 1989㸧ࠊ≉ᐃ✀࡟࠾࠸࡚ࡣேᕤ⎔ቃࡀ㏫࡟ᐤ⏕⽸⹸࡟ዲ㐺࡞⏕⫱࣭ఏ᧛ᶵ఍ࢆ๰ฟࡍࡿࡇ࡜ࡶ࠶ࡿࠋ㣗⫗࡜ࡋ࡚ࡢ௜ຍ౯್ࢆࡘࡅࡿࡓࡵ࡟ࠊ⮬↛ࡢ୰࡛⫱ࡗࡓ㇜ࡢ㣫⫱㸦ᨺ㣴㸧ࡶᅜෆ࡛ヨࡳࡽ 

රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-9

ࢀ࡚࠸ࡿࠋࡋ࠿ࡋ㇜࡛ࡣ㐣ཤࡢ⑓Ẽ࡛࠶ࡗࡓࡣࡎࡢᐤ⏕⽸⹸⑕ࡣࠊ㔝⏕࢖ࣀࢩࢩ࡛ࡣࡋ
ࡗ࠿ࡾ࡜ᅜෆ࡛ࡶ⏕ά⎔ࡀ⥔ᣢࡉࢀ࡚࠸ࡿࠋࡑࡢࡓࡵ࢖ࣀࢩࢩࢆಖ⹸ᐟ୺࡜ࡋ࡚༑ศ࡟
⪃៖ࡋࠊᐙ␆㣫⫱࢚ࣜ࢔࡜࢖ࣀࢩࢩ⏕ᜥᇦ࡜ࡢ᥋Ⅼࡀ࡞࠸ࡼ࠺࡟ὀពࡍࡿࡇ࡜ࡶᛀࢀ࡚
ࡣ࡞ࡽ࡞࠸ࠋㅰ㎡ᮏ◊✲ࢆ㐙⾜ࡍࡿ࡟࠶ࡓࡾࠊࡈ༠ຊ࠸ࡓࡔ࠸ࡓ⊟཭఍ࡢⓙᵝ࡟ཌࡃᚚ♩⏦ࡋୖࡆࡿࠋ
ࡲࡓࠊᮏ◊✲ࡢ୍㒊ࡣ JSPS ⛉◊㈝ 22580349ࠊ25450478 ࡢຓᡂࢆཷࡅࡓࠋ

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රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-10

➨ 㸯㸮 ❶            භ⏥ᒣ࢖ࣀࢩࢩၥ㢟ࡢ⌧≧࡜ㄢ㢟㎷ ▱㤶 1࣭ᶓᒣ┿ᘪ 1,2
せ Ⅼ࣭ භ⏥ᒣ⣔༡ഃࡢᕷ⾤ᆅ࡛Ⓨ⏕ࡋ࡚࠸ࡿ࢖ࣀࢩࢩၥ㢟ࡢᑐ⟇ࡢ᳨ウ࡟ྥࡅ࡚ࠊභ⏥ᒣ࢖ࣀࢩࢩ࡟㛵ࢃࡿ㐣ཤࡢ㈨ᩱࠊࣄ࢔ࣜࣥࢢࠊ᳃ᯘື≀◊✲ࢭࣥࢱ࣮ࡢ⵳✚ࢹ࣮ࢱࢆࡲ࡜ࡵࡓࠋ࣭ ฟἐ࣭⿕ᐖⓎ⏕ࡢ᭱኱ࡢཎᅉ࡛࠶ࡿ࢖ࣀࢩࢩࡢ㣵௜ࡅ⾜Ⅽࡣࠊ1965 ᖺ࡟ࡣ☜ㄆࡉࢀ࡚࠸ࡓࠋᙜ᫬࠿ࡽࡍ࡛࡟ே័ࢀࡋࡓ࢖ࣀࢩࢩ࡟ࡼࡿ⿕ᐖࡀⓎ⏕ࡋ࡚࠾ࡾࠊ⿕ᐖ㜵Ṇࡢࡓࡵࡢ᭷ᐖᤕ⋓ࡶᐇ᪋ࡉࢀ࡚࠸ࡓࠋࡘࡲࡾභ⏥ᒣ࢖ࣀࢩࢩၥ㢟ࡣ⣙ 50ᖺ㛫⥆࠸࡚࠸ࡿࠋ࣭ ࠿ࡘ࡚ࡢභ⏥ᒣࡣࠊ㐣๫࡞ᮦᮌ฼⏝࡟ࡼࡗ࡚ࡣࡆᒣ࡜࡞ࡾࠊ㔝⏕ື≀ࡀ⏕ᜥ࡛ࡁࡿ⎔ቃ࡛ࡣ࡞࠿ࡗࡓࠋ1900 ᖺ௦ࡢ✚ᴟⓗ࡞᳜ᯘ஦ᴗ࡟ࡼࡾࠊ⌧ᅾࡣᗈⴥᶞࡀඃ
༨ࡍࡿࠊ࢖ࣀࢩࢩࡢዲ㐺࡞⏕ᜥ⎔ቃ࡬࡜ኚ໬ࡋ࡚࠸ࡿࠋ࣭ ⚄ᡞᕷࡣ 2002 ᖺ࡟࢖ࣀࢩࢩࡢ㣵௜ࡅ⚗Ṇ᮲౛ࢆ᪋⾜ࡋࠊ2004 ᖺ࡟㣵௜ࡅ⚗Ṇࡢつไ༊ᇦࢆタ⨨ࡋࡓࠋࡇࢀࡽࡢ᪋⟇ࡢຠᯝࡣࡳࡽࢀࡓࡶࡢࡢࠊ㣵௜ࡅ⾜Ⅽࡢ᰿⤯ࡀ㞴ࡋࡃࠊ㏆ᖺࠊⱞ᝟࣭㏻ሗ௳ᩘࡸ᭷ᐖᤕ⋓ᩘࡣቑຍഴྥ࡟࠶ࡿࠋ࣭ ࢖ࣀࢩࢩ࡟࡜ࡗ࡚ዲ㐺࡞⏕ᜥ⎔ቃ࡟ኚ໬ࡋࡓභ⏥ᒣ࡛ࡣࠊ⏕ᜥᩘࡀῶᑡࡍࡿせᅉࡣᴟࡵ࡚ᑡ࡞ࡃࠊࡲࡓࠊ㣵௜ࡅ⾜Ⅽࡀ⥆ࡃ㝈ࡾ࢖ࣀࢩࢩࡢᕷ⾤ᆅฟἐ࡜⿕ᐖⓎ⏕ࡣ࡞ࡃ࡞ࡽ࡞࠸ࡇ࡜ࡀ⪃࠼ࡽࢀࡿࠋ௒ᚋࡶ㒔ᕷఫẸ࡬ࡢᬑཬၨⓎࡢ᥎㐍࡜࡜ࡶ࡟࢖ࣀࢩࢩࡢᤕ⋓యไࡢ☜ಖࡀồࡵࡽࢀࡿࠋ

Key words㸸භ⏥ᒣࠊ㣵௜ࡅ⾜Ⅽࠊ᳜⏕ኚ㑄ࠊⱞ᝟࣭㏻ሗࠊᤕ⋓ᑐᛂ㸬ࡣࡌࡵ࡟ 㔝⏕ື≀࡜ே࡜ࡢ㌵㎚ၥ㢟ࡣࠊ㏆ᖺࠊࡇࢀࡲ࡛Ⓨ⏕ࡢ࡞࠿ࡗࡓ㒔ᕷ⎔ቃ࡛ࡶຨⓎࡋ࡚࠸ࡿࠋ㒔ᕷ⎔ቃ࡟࠾ࡅࡿ㔝⏕ື≀ࡢฟἐࡸ⿕ᐖࡢⓎ⏕௳ᩘࡣ඲ᅜⓗ࡟ቑຍഴྥ࡟࠶ࡿࡇ
࡜࠿ࡽࠊၥ㢟ࡣ୍᫬ⓗ࠶ࡿ࠸ࡣ౛እⓗ࡞⌧㇟࡛ࡣ࡞ࡃࠊ☜ᐇ࡟㒔ᕷ⎔ቃ࡬Ἴཬࡋ࡚ࡁ࡚
࠸ࡿ࡜࠸࠼ࡿࠋேཱྀᐦᗘࡀ㧗ࡃ஺㏻㔞ࡢከ࠸㒔ᕷ⎔ቃ࡛ࡣࠊ୰ᒣ㛫ᆅᇦࡢ≧ἣ࡜ࡣ඲ࡃ
␗࡞ࡿ஦ែࡀⓎ⏕ࡋ࡚࠸ࡿࠋࡓ࡜࠼ࡤࠊ㔝⏕ື≀ࡀࠊண᝿ࡋᚓ࡞࠸஺㏻㔞ࡢከ࠸ሙᡤࡸ
⥳ᆅ⎔ቃࡢ࡞࠸ሙᡤ࡟✺Ⓨⓗ࡟ฟἐࡋ࡚࠸ࡿ㸦ᮐᖠᕷ 2010㸹㎷࣭ᶓᒣ 2012㸧ࠋࡲࡓࠊேཱྀࡀᐦ㞟ࡋ࡚࠸ࡿࡓࡵࠊே㌟⿕ᐖࡸ஺㏻஦ᨾ࡞࡝ேࡢ⏕࿨࡟┤᥋㛵ࢃࡿ⥭ᛴⓗ࠿ࡘ῝රᗜ┴᳃ᯘື≀◊✲ࢭࣥࢱ࣮࣭ රᗜ┴❧኱Ꮫ⮬↛࣭⎔ቃ⛉Ꮫ◊✲ᡤ 

රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-10

้࡞⿕ᐖࡢ༴㝤ᛶࡀᣲࡆࡽࢀ࡚࠸ࡿ㸦ᮐᖠᕷ 2010㸹㤶ᕝ┴ 2011㸧ࠋ㒔ᕷ⎔ቃ࡞ࡽ࡛ࡣ
ࡢၥ㢟ࡀ᝿ᐃࡉࢀࠊࡑࡢሙ㝈ࡾࡢᑐᛂ࡛ࡣ࡞ࡃࠊయ⣔❧࡚ࡓ㔝⏕ື≀࡬ࡢᑐᛂ࣭ᑐ⟇ࡢ௙⤌ࡳࢆᵓ⠏ࡍࡿࡇ࡜ࡀᛴົ࡛࠶ࡿ㸦㎷࣭ᶓᒣ 2012㸧ࠋ රᗜ┴࡛ࡣࠊ㜰⚄ᆅᇦࡢᕷ⾤ᆅ࡟㔝⏕ື≀ࡀฟἐࡋࠊ⿕ᐖࡀⓎ⏕ࡋ࡚࠸ࡿࠋ࡜ࡃ࡟⚄ᡞᕷࠊⰱᒇᕷࠊすᐑᕷࡢභ⏥ᒣ⣔༡ഃᩳ㠃ࡢᆅᇦ࡟࠾ࡅࡿࢽ࣍ࣥ࢖ࣀࢩࢩ㸦Sus scrofa

leucomystax㸹௨ୗ࢖ࣀࢩࢩ㸧ࡢၥ㢟ࡣ῝้࡛࠶ࡿࠋභ⏥ᒣࡢ࢖ࣀࢩࢩၥ㢟ࡢⓎ➃ࡣࠊ࢖ࣀࢩࢩ࡬ࡢ㣵௜ࡅ⾜Ⅽ࡛࠶ࡿ㸦⚄ᡞᕷ 2002㸧ࠋ㣵௜ࡅ⾜Ⅽ࡟ࡼࡾே័ࢀࡋࡓ࢖ࣀࢩࢩ
ࡀࠊேᕤ≀࡟ᇳ╔ࡋࠊࢦ࣑ࢆⲨࡽࡋࡓࡾࠊேࢆくࡗࡓࡾࡋ࡚࠸ࡿࠋࡇࢀࡲ࡛⚄ᡞᕷࢆ୰ᚰ࡜ࡋ࡚ᑐ⟇ࡀᐇ⾜ࡉࢀ࡚ࡁ࡚࠸ࡿࡀ㸦⚄ᡞᕷ 2002㸹⚄ᡞᕷ 2005㸧
ࠊ᰿ᮏⓗ࡞ཎᅉ࡛
࠶ࡿ㣵௜ࡅ⾜Ⅽࡢ᰿⤯ࡀ㞴ࡋ࠸ࡓࡵࠊ⿕ᐖࡀ࡞ࡃ࡞ࡽ࡞࠸ࠋே័ࢀࡋࡓၥ㢟ಶయࡀᤕ⋓ࡉࢀ୍࡚᫬ⓗ࡟ฟἐࡸ⿕ᐖࡀ཰ࡲࡗ࡚ࡶࠊ㣵௜ࡅ⾜Ⅽ࡟ࡼࡾࠊ෌ࡧูࡢே័ࢀၥ㢟ಶయࡀฟ⌧ࡋ࡚⿕ᐖࡀ෌Ⓨࡍࡿ࡜࠸࠺⧞ࡾ㏉ࡋ࡛࠶ࡿࠋ ࡇࢀࡲ࡛භ⏥ᒣ࠾ࡼࡧ࢖ࣀࢩࢩ࡟㛵ࡋ࡚ࡣࠊከࡃࡢ᭩⡠࡟グ㍕ࡉࢀ࡚࠸ࡿࠋࡑࡇ࡛ᮏ
✏࡛ࡣ࡜ࡃ࡟ࠊᕷ⾤ᆅ࡟ฟἐࡍࡿ࢖ࣀࢩࢩࡢ⿕ᐖၥ㢟࡟↔Ⅼࢆ࠶࡚ࠊභ⏥ᒣ࢖ࣀࢩࢩࡢ
ฟἐ࣭⿕ᐖࡢపῶ࡟ྥࡅࡓᢤᮏⓗ࡞ᑐ⟇ࡢ᳨ウࢆ⾜࠺ࡇ࡜ࢆ┠ⓗ࡜ࡍࡿࠋ୰࡛ࡶࡇࢀࡲ
࡛ࡢ࢖ࣀࢩࢩ࡬ࡢ㣵௜ࡅ⾜Ⅽ⮬యࡢ≧ἣ࡟ຍ࠼ࠊ⿕ᐖ࡜ᤕ⋓➼ࡢᑐ⟇≧ἣࠊฟἐಶయࡀ
⏕ᜥࡋ࡚࠸ࡿභ⏥ᒣ⣔ࡢ᳜⏕ࡢṔྐⓗ࡞ኚ㑄ࡢ≧ἣ࡟╔┠ࡍࡿࠋ㸬㞟⣙㈨ᩱ
 ᮏ✏࡛ࡣࠊභ⏥ᒣ࢖ࣀࢩࢩ࡟㛵ࢃࡿ᭩⡠ࡸ㈨ᩱࠊ㛵ಀ⪅࡬ࡢࣄ࢔ࣜࣥࢢࠊ2009 ᖺ௨㝆ࡢ᳃ᯘື≀◊✲ࢭࣥࢱ࣮ࡢᑐᛂグ㘓ࢆඖ࡟ࡲ࡜ࡵࡓࠋྛ㡯┠࡛฼⏝ࡋࡓ㈨ᩱࢆ௨ୗ࡟♧ࡍࠋ ࠕ10㸫3㸬⚄ᡞᕷࡢ༊ᇦࠊ㫽⋇ಖㆤ༊ࡢศᕸ࡜࢖ࣀࢩࢩࡢᤕ⋓ᙧែ࡛ࠖࡣࠊ➨㸯㸯ḟ㫽⋇ಖㆤ஦ᴗィ⏬᭩㸦රᗜ┴ 2012㸧࡜᳃ᯘື≀◊✲ࢭࣥࢱ࣮࡟⵳✚ࡉࢀ࡚࠸ࡿරᗜ┴ࡢ࣮࣋ࢫ࣐ࢵࣉࢆ฼⏝ࡋࡓࠋࠕ10㸫4㸬භ⏥ᒣ࢖ࣀࢩࢩࡢ㣵௜ࡅ⾜Ⅽࠊ⿕ᐖࠊᤕ⋓ࡢṔྐ࡛ࠖࡣࠊභ⏥ᒣ࢖ࣀࢩࢩࡢ㣵௜ࡅ⾜Ⅽ࡟㛵ࡍࡿ᭩⡠㸦⏣୰ 1986㸧ࠊ⚄ᡞᕷࡢ᮲౛࡟㛵
ࡍࡿ㈨ᩱ㸦⚄ᡞᕷ 2001㸹⚄ᡞᕷ 2002㸹⚄ᡞᕷ 2005㸧ࠊ㛵ಀ⪅ࡢࣄ࢔ࣜࣥࢢࢆࡲ࡜ࡵࡓࠋ
භ⏥ᒣࡢ᳜⏕࡟㛵ࡋ࡚ࡣࠊ2012 ᖺ 4 ᭶࡟⚄ᡞᕷᘓタᒁබᅬ◁㜵㒊භ⏥ᒣᩚഛᐊࡀⓎ⾜ࡋࡓࠕභ⏥ᒣ᳃ᯘᩚഛᡓ␎ࠖ࡟ヲࡋ࠸㸦⚄ᡞᕷ 2012㸧ࠋࡑࡇ࡛ࠕ10㸫5㸬භ⏥ᒣ⣔ࡢ᳜⏕ኚ㑄࡛ࠖࡣࠊࡇࡢ㈨ᩱࢆཧ↷ࡋࠊ࢖ࣀࢩࢩ⟶⌮࡟㛵ࢃࡿ㒊ศ࡟⤠ࡗ࡚せ⣙ࡋࡓࠋࡲࡓ
᳜⏕ᅗࡢసᡂ࡟ࡣࠊ኱᪥ᮏᖇᅧ㝣ᆅ

㔞㒊Ⓨหࡢ 5 ୓ศࡢ 1 ᆅᙧᅗ㸦1900 ᖺ௦ࡢ᳜⏕ᅗ㸧ࠊᅜᅵᆅ⌮㝔Ⓨหࡢ 5 ୓ศࡢ 1 ᆅᙧᅗ㸦1950 ᖺ௦ࡢ᳜⏕ᅗ㸧ࠊ⎔ቃ┬⏕≀ከᵝᛶࢭࣥࢱ࣮Ⓨ⾜ࡢ⮬↛⎔ቃಖ඲ᇶ♏ㄪᰝࠕ➨ 2-5 ᅇ࡜➨ 6-7 ᅇ᳜⏕ㄪᰝ㔜ࡡྜࢃࡏ᳜⏕ᅗ㸦5
୓ศࡢ 1 ࡜ 2 ୓ 5 ༓ศࡢ 1㸧ࠖ㸦1980 ᖺ௦࡜ 2000 ᖺ௦ࡢ᳜⏕ᅗ㸧ࢆ౑⏝ࡋࡓࠋ
ࠕ10㸫6㸬
㣵௜ࡅ⾜Ⅽ࡜⿕ᐖࡢᐇែ࡛ࠖࡣࠊ2009 ᖺ௨㝆࡟᳃ᯘື≀◊✲ࢭࣥࢱ࣮ࡀㄪᰝࠊศᯒࡋ
ࡓグ㘓ࢆࡲ࡜ࡵࡓࠋࠕ10㸫7㸬ఫẸࡢⱞ᝟࣭㏻ሗ࡜᭷ᐖᤕ⋓≧ἣ࡛ࠖࡣࠊ➨㸰ᮇ࢖ࣀࢩ 

රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-10

ࢩಖㆤ⟶⌮ィ⏬ࡢ㈨ᩱ㸦රᗜ┴ 2012㸧࡜᳃ᯘື≀◊✲ࢭࣥࢱ࣮࡟⵳✚ࡉࢀ࡚࠸ࡿ᭷ᐖ
ᤕ⋓᝟ሗࢆࡲ࡜ࡵࡓࠋࠕ10㸫8㸬ᑐ⟇≧ἣ࡛ࠖࡣࠊ⚄ᡞ㎰ᯘỈ⏘஦ົᡤࡀᐇ᪋ࡋࡓභ⏥ᒣ࢖ࣀࢩࢩᑐ⟇஦ᴗࡢሗ࿌᭩ࢆᘬ⏝ࡋࡓࠋ㸬⚄ᡞᕷࡢ༊ᇦࠊ㫽⋇ಖㆤ༊ࡢศᕸ࡜࢖ࣀࢩࢩࡢᤕ⋓ᙧែ ⚄ᡞᕷࡣࠊᨻ௧ᣦᐃ㒔ᕷ࡛࠶ࡾ 9 ࡘࡢ༊࡟ศ࠿ࢀࡿ㸦ᅗ㸯㸧ࠋභ⏥ᒣ⣔ࡣࠊ⚄ᡞᕷࡢ୰ኸࢆᮾす࡟ᶓ᩿ࡋ࡚࠸ࡿࠋභ⏥ᒣ⣔ࡼࡾ༡ഃࡢ℩ᡞෆᾏ࡟㠃ࡍࡿᮾℿ༊ࠊℿ༊ࠊ୰ኸ༊ࠊරᗜ༊ࠊ㛗⏣༊ࠊ㡲☻༊ࠊᆶỈ༊࡛ࡣࠊఫᏯᆅࡸၟᴗ᪋タࡀ❧ࡕ୪ࡧேཱྀᐦᗘࡀ㧗࠸ࠋࡇࡢ୰࡛ࠊᕷ⾤ᆅ࡟࠾ࡅࡿ࢖ࣀࢩࢩ⿕ᐖࡀ࡜ࡃ࡟῝้࡞ࡢࡣࠊᮾℿ༊ࠊℿ༊ࠊ୰ኸ༊ࡢ 3 ༊࡛࠶ࡿࠋ୍᪉࡛ࠊභ⏥ᒣ⣔ࡼࡾ໭ഃࡢ໭༊ࠊす༊࡟ࡣ㎰ᆅࡸᒣᯘࡀᗈࡀࡾࠊ༡
ഃ࡟ẚ࡭࡚ேཱྀᐦᗘࡣప࠸ࠋࡇࡢ 2 ༊࡛ࡣ୺࡟࢖ࣀࢩࢩ࡟ࡼࡿ㎰ᴗ⿕ᐖࡀⓎ⏕ࡋ࡚࠸ࡿࠋ
භ⏥ᒣ⣔ࡣࠊ1956 ᖺ࡟℩ᡞෆᾏᅜ❧බᅬ࡟⦅ධࡉࢀࡓࠋ⌧ᅾࠊ⣙ 9,000ha
ࡢ㠃✚ࡢ࠺ࡕ 5,440ha ࡀ㫽⋇ಖㆤ༊ࠊࡑࡢ࠺ࡕ 138ha ࡀ㫽⋇ಖㆤ༊≉ูᆅᇦ࡟ᣦᐃࡉࢀ࡚࠸ࡿ㸦ᅗ㸯㸹රᗜ┴ 2012㸧ࠋභ⏥ᒣ⣔༡ഃ࡛ࡣࠊ᳃ᯘࡢ࡯ࡰ඲࡚ࡀ
㫽⋇ಖㆤ༊࡟࡞ࡿࡓࡵࠊ⊁⊟ࡀ⚗Ṇࡉࢀ࡚࠾ࡾࠊᤕ⋓ࡣᖺ㛫ࢆ㏻ࡌ࡚ࡍ࡭࡚
᭷ᐖᤕ⋓࡜࡞ࡿࠋࡲࡓ㖠ჾࡢ౑⏝ࡀ⚗Ṇࡉࢀ࡚࠸ࡿࡓࡵࠊᤕ⋓᪉ἲࡣࢃ࡞࡟
㝈ࡽࢀࡿࠋࡇࢀ࡟ᑐࡋ࡚භ⏥ᒣ⣔໭ഃ࡟ࡘ࠸࡚ࡣࠊ୍㒊㫽⋇ಖㆤ༊ࡀධࡿࡶᅗ  ⚄ᡞᕷྛ༊ࡢ❧ᆅ࡜㫽⋇ಖㆤ༊ࠊ㖠⊟⚗Ṇ༊ᇦࡢศᕸ

ࡢࡢ࡯࡜ࢇ࡝ࡀ⊁⊟ྍ⬟ᆅᇦ࡛࠶ࡾࠊ
㖠⊟ࡀྍ⬟࡞ᆅᇦࡶ࠶ࡿ㸦ᅗ㸯㸧ࠋ

௨ୖࡢ㏻ࡾࠊ⚄ᡞᕷෆ࡟࠾࠸࡚ࡶභ⏥ᒣ⣔༡ഃࡢᆅᇦ࡛ࡣᕷ⾤ᆅ࡟ฟἐࡍࡿ࢖ࣀࢩࢩ
⿕ᐖࡢ㜵Ṇࡢࡓࡵࡢࢃ࡞࡟ࡼࡿ᭷ᐖᤕ⋓ࠊභ⏥ᒣ⣔໭ഃࡢᆅᇦ࡛ࡣ㎰స≀⿕ᐖᑐ⟇ࡢࡓࡵࡢ᭷ᐖᤕ⋓࠾ࡼࡧ⊁⊟࡜ࠊᆅᇦ࡟ࡼࡗ࡚⿕ᐖࡢࢱ࢖ࣉࠊᤕ⋓┠ⓗࡸᤕ⋓᪉ἲࡀ␗࡞ࡿࠋ

㸬භ⏥ᒣ࢖ࣀࢩࢩࡢ㣵௜ࡅ⾜Ⅽࠊ⿕ᐖࠊᤕ⋓ࡢṔྐ භ⏥ᒣ࡟࠾ࡅࡿ࢖ࣀࢩࢩ࡬ࡢ㣵௜ࡅ⾜Ⅽࠊ࢖ࣀࢩࢩࡢ⿕ᐖ࡜᭷ᐖᤕ⋓࡟㛵ࢃࡿฟ᮶஦
ࢆᅗ㸲ࡢᖺ⾲࡟ࡲ࡜ࡵࡓࠋ
㹼 ᖺ௦࢖ࣀࢩࢩ࡬ࡢ㣵௜ࡅ⾜Ⅽࡀ᭱ึ࡟ጞࡲࡗࡓ᫬ᮇࡣᐃ࠿࡛ࡣ࡞࠸ࡀࠊ௒࠿ࡽ⣙ 50 ᖺ๓
ࡢ 1965 ᖺ࡟ࡣ⾜ࢃࢀ࡚࠸ࡓࠋᙜ᫬࠿ࡽࡍ࡛࡟㣵௜ࡅ⾜Ⅽࡢᙳ㡪࡛ࠊ࢖ࣀࢩࢩࡀᕷ⾤ᆅ 

රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-10

࡟ฟἐࡋ⿕ᐖࢆⓎ⏕ࡉࡏ࡚࠸ࡓ㸦⚄ᡞ
ᕷ 2001㸧ࠋࡲࡓ 1979 ᖺ࡟ࡣࠊⰱᒇᕷ
ࡢ㧗ᗙࡢ⁪ࡸ௜㏆ࡢ◁㜵ሖሐ࡟࡚ᆅ
ඖࡢⓏᒣ⪅࡟ࡼࡿ㣵௜ࡅ⾜Ⅽࡀጞࡲࡗࡓࠋࡇࢀࡽࡢሙᡤࡀࠊ኱つᶍ࡞㣵௜ࡅሙᡤ࡜ࡋ࡚ၥ㢟࡜࡞ࡿⰱᒇ࢖ࣀࢩ
ࢩᮧ࡛࠶ࡿ㸦ᅗ㸰㸹⏣୰ 1986㸧
ࠋ1960
㹼70 ᖺ௦ࡣࠊ㔝⏕ື≀ࡀᕼᑡ࡞Ꮡᅾ
࡛࠶ࡗࡓࡇ࡜࠿ࡽ㸦ᶓᒣ 2014㸧ࠊ㛫㏆࡛࢖ࣀࢩࢩࢆぢࡿࡇ࡜ࡀ࡛ࡁࡿሙᡤᅗ  ࢖ࣀࢩࢩᮧࡀᏑᅾࡋࡓሙᡤ

࡜ࡋ࡚ࠊභ⏥ᒣ࡛ࡢ࢖ࣀࢩࢩࡢ㣵௜ࡅ
⾜Ⅽࡣᗈࡲࡗࡓ࡜⪃࠼ࡽࢀࡿࠋᙜ᫬ࡣࠊ
ே័ࢀ࢖ࣀࢩࢩࢆ◊✲ᑐ㇟࡜ࡋ࡚࠸
ࡓሗ࿌ࡶ࠶ࡾ㸦Hirotani & Nakatani
1987; Nakatani & Ono 1994㸧ࠊ㔝⏕ື≀࡟ᑐࡍࡿ㣵௜ࡅ⾜Ⅽࡀබ↛࡜ၿព࡜ࡋ࡚⾜ࢃࢀ࡚࠸ࡓ࡜᥎ᐹࡉࢀࡿࠋ
ࡑࡢ୍᪉࡛ࠊ࢖ࣀࢩࢩ⿕ᐖࡢ㜵Ṇࡢࡓࡵ࡟රᗜ┴⊟཭఍ࡢᮾℿࠊℿࠊᦶ⪨ࠊⵌྜࡢᅄᨭ㒊࠿ࡽ࡞ࡿࠕ⾲භ⏥᭷ᐖ㫽ᅗ  ⾲භ⏥᭷ᐖ㫽⋇㥑㝖⌜ࡢάື⋇㥑㝖⌜ࠖࡀ⦅ᡂࡉࢀࠊභ⏥ᒣ⣔࡛࢖ࣀࢩࢩᤕ⋓ࡀᐇ᪋ࡉࢀ࡚࠸ࡓ㸦ᅗ㸱㸧ࠋᙜ᫬ࡣ≟ࢆ౑ࡗࡓ㖠ჾ࡟ࡼࡿᤕ⋓ࡀ୺࡛࠶ࡗࡓࠋ
㹼 ᖺ௦ᗘ㔜࡞ࡿ࢖ࣀࢩࢩ⿕ᐖࡢⓎ⏕ࢆཷࡅࠊ⾜ᨻⓗ࡟㣵௜ࡅ⾜Ⅽࡢὀពႏ㉳ࡀ⾜ࢃࢀࡓࠋࡋ
࠿ࡋⰱᒇ࢖ࣀࢩࢩᮧ࡞࡝භ⏥ᒣ⣔ྛᆅ࡛ࠊ⾜ᨻࡢὀពࢆ↓どࡋࡋࡓ㣵௜ࡅ⾜Ⅽࡀ⥆ࡅࡽࢀࡓࠋࡑࡢᙳ㡪࡛࢖ࣀࢩࢩࡢ⿕ᐖࡸⱞ᝟ࡢ௳ᩘࡣᛴቑࡋࠊ⿕ᐖᆅᇦࡶᣑ኱ࡋࡓࠋ᭷ᐖᤕ
⋓࡟ࡘ࠸࡚ࡣࠊ⊟≟࡟ࡼࡿ஦ᨾࡢⓎ⏕ࡸఫẸ࠿ࡽࡢ㖠⊟࡟ᑐࡍࡿ㠀㞴ࡢቑຍࢆཷࡅ࡚ࠊ
1980 ᖺ௦࡟ᤕ⋓ᡭẁࡀ㖠ჾ࠿ࡽࢃ࡞࡟ษࡾ᭰ࢃࡗࡓࠋ1980 ᖺ௦ᚋ༙ࡢ࢖ࣀࢩࢩ⿕ᐖࡢ῝้໬࡜㣵௜ࡅ⾜Ⅽࡢ⥅⥆ࢆཷࡅ࡚ࠊ1992 ᖺ࡟⚄ᡞᕷࡀࠊ࢖ࣀࢩࢩ࡬ࡢ㣵௜ࡅࡢ⚗Ṇࢆᐃࡵࡿ᮲౛ไᐃ࡟ྥࡅ࡚ㄪᩚࢆጞࡵࡓࠋ
㹼 ᖺ௦
2002 ᖺ 5 ᭶ 1 ᪥࡟࢖ࣀࢩࢩ࡬ࡢ㣵௜ࡅࢆ⚗Ṇࡍࡿ඲ᅜึࡢ᮲౛࡜ࡋ࡚ࠊ
ࠕ⚄ᡞᕷ࠸ࡢ
ࡋࡋࡢฟἐཬࡧ࠸ࡢࡋࡋ࠿ࡽࡢ༴ᐖࡢ㜵Ṇ࡟㛵ࡍࡿ᮲౛ࠖࠊ࠸ࢃࡺࡿࠕ࢖ࣀࢩࢩ㣵௜ࡅ⚗Ṇ᮲౛ࠖࡀ᪋⾜ࡉࢀࡓࠋ᮲౛࡛ࡣࠊ࢖ࣀࢩࢩ࡬ࡢ㣵௜ࡅ⾜Ⅽ࡜㣵࡜࡞ࡾ࠺ࡿࡈࡳࡢᨺ
Რࡢ⚗Ṇࡀᐃࡵࡽࢀࡓࠋ᮲౛㐪཯⪅࡬ࡢᥐ⨨࡜ࡋ࡚ࡣࠊᕷ࠿ࡽࡢᣦᑟ່࣭࿌ࡀᐃࡵࡽࢀࠊ⨩๎ࡣタࡅࡽࢀ࡞࠿ࡗࡓ㸦⚄ᡞᕷ 2002㸧ࠋ᪋⾜ᙜึࡣࠊ᮲౛ࡢ㐺⏝⠊ᅖࢆ඲ᕷᑐ㇟࡜ࡋ 

රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-10

࡚࠸ࡓࡀࠊ2004 ᖺ࡟᮲౛㐪཯࡜⿕ᐖࡀ῝้࡞ᆅᇦ࡟ⓗࢆ⤠ࡗ࡚ᑐ⟇ࢆᙉ໬ࡍࡿࡓࡵ࡟ࠊつไ༊ᇦࡀタᐃࡉࢀࡓࠋつไ༊ᇦ࡟ࡣᮾℿ༊࡜ℿ༊ࡢ୍㒊ᆅᇦࡀᣦᐃࡉࢀࡓ㸦⚄ᡞᕷ
2005㸧ࠋつไ༊ᇦእ࡛࠶ࡗࡓ୰ኸ༊࡟࡚ࠊ2011 ᖺ࡟㣵௜ࡅ࡟ࡼࡾே័ࢀࡋࡓ࢖ࣀࢩࢩࡀ
ே㌟⿕ᐖࢆከⓎࡉࡏࡓࠋࡇࢀࢆཷࡅ࡚ࠊ2012 ᖺ࡟୰ኸ༊ࡢ୍㒊ᆅᇦࡶつไ༊ᇦ࡟ຍ࠼ࡽࢀࡓࠋすᐑᕷ࡟࠾࠸࡚ࡶࠊ2013 ᖺ 4 ᭶ 1 ᪥࡟ࠊࠕすᐑᕷ࠸ࡢࡋࡋ㣵ࡸࡾ⚗Ṇ᮲౛ࠖࡀ᪋⾜ࡉࢀࡓࠋ⚄ᡞᕷࡢ᮲౛࡜ྠᵝ࡟㣵ࡸࡾࢆ⚗Ṇࡍࡿ༊ᇦࢆタᐃࡋࠊࡑࡢ༊ᇦෆ࡛ࡢ᮲౛㐪཯⪅࡟ᑐࡋ࡚ࡣࠊᕷ࠿ࡽࡢ່࿌ᥐ⨨ࡀᐃࡵࡽࢀࡓ㸦すᐑᕷ 2013㸧ࠋ㏆ᖺᐇ᪋ࡉࢀ࡚࠸ࡿභ⏥ᒣ࢖ࣀࢩࢩ≉᭷ࡢᑐ⟇࡜ࡋ࡚ࠊ࢖ࣀࢩࢩ⥭ᛴᑐ⟇༠ຊဨ㸦රᗜ┴㸧࡜࢖ࣀࢩࢩ㣵௜ࡅ┘どဨ㸦⚄ᡞᕷ㸧ࡢタ⨨ࡀᣲࡆࡽࢀࡿࠋ2005 ᖺ࠿ࡽጞࡲࡗࡓ
࢖ࣀࢩࢩ⥭ᛴᑐ⟇༠ຊဨไᗘࡣࠊ࢖ࣀࢩࢩࡀఫẸ࡟༴ᐖࢆ୚࠼ࡿྍ⬟ᛶࡀᴟࡵ࡚㧗ࡃ࡞ࡿ⥭ᛴ஦ែ࡟ᑐฎࡍࡿࢩࢫࢸ࣒࡛࠶ࡿࠋ࢖ࣀࢩࢩ⥭ᛴᑐ⟇༠ຊဨࡣࠊྛ༊ࡢ⊟཭఍ᨭ㒊࠿ࡽᩘྡ㑅ฟࡉࢀࠊࡑࢀࡒࢀࡢ༊ࢆ⟶㎄ᇦ࡜ࡋ࡚άືࡋ࡚࠸ࡿࠋ࢖ࣀࢩࢩ㣵௜ࡅ┘どဨࡣࠊ2011 ᖺ࡜ 2012 ᖺ࡟⚄ᡞᕷ࡟ࡼࡿ࢖ࣀࢩࢩ㣵௜ࡅ⚗Ṇᑐ⟇஦ᴗ㸦⥭ᛴ㞠⏝ᑵᴗᶵ఍๰ฟ஦ᴗ࡜ࡋ࡚ᐇ᪋㸧ࡢ୍ࡘࡋ࡚タ⨨ࡉࢀࡓࠋ┘どဨࡣࠊྛ༊ࡀᢕᥱࡍࡿ㣵௜ࡅሙᡤ࿘
㎶ࢆࣃࢺ࣮ࣟࣝࡋࠊ㣵௜ࡅ⪅࡬ࡢὀព࣭㆙࿌࡜࿘㎶ఫẸ࡬ࡢၨⓎάືࢆᐇ᪋ࡋࡓࠋᅗ  භ⏥ᒣ࢖ࣀࢩࢩࡢ㣵௜ࡅࠊ⿕ᐖࠊᤕ⋓࡟㛵ࡍࡿᖺ⾲ 

රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-10

 ௨ୖࡢ࡜࠾ࡾࠊභ⏥ᒣ࢖ࣀࢩࢩࡢ㣵௜ࡅ⾜Ⅽ࡜⿕ᐖࠊ᭷ᐖᤕ⋓ࡣ⣙ 50 ᖺ࡟Ώࡗ࡚⥆࠸࡚࠸ࡿࠋ㣵௜ࡅ⾜Ⅽࡸࢦ࣑ࡢᨺᲠၥ㢟࡟ᑐࡋ࡚ࠊ⚄ᡞᕷࢆ୰ᚰ࡜ࡋࡓ᰿ᙉ࠸ὀពႏ㉳ࡢᐇ᪋࡜㣵௜ࡅ⚗Ṇ᮲౛ࡢ᪋⾜࡟ࡼࡾࠊⰱᒇ࢖ࣀࢩࢩᮧࡢᾘኻࡸࢦ࣑࣐ࢼ࣮ࡢᨵၿ࡞࡝
ࡢຠᯝࡣㄆࡵࡽࢀࡓࠋࡋ࠿ࡋ୍㒊ࡢఫẸࡸⓏᒣ⪅ࡀ㣵௜ࡅ⾜Ⅽࢆ⥆ࡅ࡚࠸ࡿࡓࡵࠊ⌧ᅾࡲ࡛⿕ᐖࡀ࡞ࡃ࡞ࡽࡎࠊ⿕ᐖ㜵Ṇࡢࡓࡵࡢ᭷ᐖᤕ⋓ࡸ⥭ᛴᑐᛂࡶ⥆ࡅࡽࢀ࡚࠸ࡿ≧ἣ࡛࠶ࡿࠋ㸬භ⏥ᒣ⣔ࡢ᳜⏕ኚ㑄 භ⏥ᒣ⣔ࡣࠊྂࡃ࠿ࡽ⇞ᩱᮦࡸ⫧ᩱࠊᘓ⠏≀ࡢᮦᮌࢆ☜ಖࡍࡿࡓࡵ࡟ᶞᮌࡢఆ᥇ࡀ⧞
ࡾ㏉ࡉࢀ࡚࠸ࡓ㸦⋢㉳ 1997㸹⚄ᡞᕷ 2012㸧ࠋ࡜ࡃ࡟ேཱྀࡀቑຍࡋࡓỤᡞ᫬௦ᚋᮇ㹼᫂἞᫬௦ࡢ㐣ᗘࡢఆ᥇࡟ࡼࡾࠊභ⏥ᒣࡣࡣࡆᒣ࡜໬ࡋࡓ㸦⋢㉳ 1997㸹࿴⏣ 1997㸹⚄ᡞᕷ
2012㸧ࠋ1900 ᖺ௦ࡢ᳜⏕ᅗ࡛ࡣࠊᒣࡢ୰⭡࡟ᗈ኱࡞Ⲩᆅࡀㄆࡵࡽࢀ㸦ᅗ㸳㹟㸧ࠊ⥳ᆅ㒊ศࡶ₤ᮌࡸⲡᆅࡀከ࠿ࡗࡓࠋࡋࡓࡀࡗ࡚ᙜ᫬ࡢභ⏥ᒣࡣ㔝⏕ື≀ࡀ⏕Ꮡࡋ࡚࠸ࡃ࡟ࡣ୙
㐺࡞⎔ቃ࡛࠶ࡗࡓ࡜᥎

ࡉࢀࠊᐇ㝿ࠊ࢖ࣀࢩࢩࡣ☜ㄆࡉࢀ࡚࠸࡞࠿ࡗࡓ࡜࠸ࢃࢀ࡚࠸ࡿ

㸦୕㇂ 2001㸧ࠋࡣࡆᒣ࡜࡞ࡗࡓභ⏥ᒣࡣỈ※ᾰ㣴ຊࡀᴟᗘ࡟పୗࡋࠊỈᐖࡀ㢖Ⓨࡋࡓ㸦⚄ᡞᕷ 2012㸧
ࠋ1892 ᖺࡢ኱Ỉᐖࢆዎᶵ࡟ࠊỈᐖ㜵Ṇ⟇࡜ࡋ࡚භ⏥ᒣࡢ◁㜵ࡸ἞ᒣᕤ஦ࡀ
╔ᕤࡋࠊ1902 ᖺ࡟᳜ᯘ஦ᴗࡀጞࡲࡗࡓࠋࡸࡏࡓᅵᆅ࡛࠶ࡗࡓࡓࡵࠊᅵተᅇ᚟࡜ࡋ࡚᰿
⢏⳦ࢆࡶࡘࣖࢩࣕࣈࢩࠊ◁㜵⏝࡜ࡋ࡚ࢽࢭ࢔࢝ࢩ࢔ࠊᐃ╔ࡋࡸࡍ࠸࢔࣐࢝ࢶ࡞࡝ࡀ᳜ᯘ
ࡉࢀࡓ㸦⋢㉳ 1997㸹⚄ᡞᕷ 2012㸧
ࠋ1950 ᖺ௦࡟ࡣࠊභ⏥ᒣࡣ࢔࣐࢝ࢶ࡜ࢥࢼࣛࡀඃ༨
ࡍࡿΰ஺ᯘ࡬࡜ኚ໬ࡋࡓ㸦ᅗ㸳㹠㸹ᑠ⯓ 2001㸧ࠋࡇࡢ㡭࠿ࡽࠊ࢖ࣀࢩࢩࡀ☜ㄆࡉࢀࡿࡼ
࠺࡟࡞ࡗࡓ࡜ࡢグ㏙ࡣከ࠸㸦⋢㉳ 1997㸹୕㇂ 2001㸧
ࠋ1980 ᖺ௦࡟ࡣࠊⲨᆅࡀᾘኻࡋ࢔
࣐࢝ࢶ࡜ࢥࢼࣛࡢΰ஺ᯘࡀᗈࡀࡗࡓ㸦ᅗ㸳㹡㸧
ࠋ2000 ᖺ௦௨㝆ࠊ㑄⛣ࡀ㐍ࢇ࡛࢔࣐࢝ࢶ
ࡣῶᑡࡋࠊࢥࢼࣛ࡞࡝ࡢⴠⴥᗈⴥᶞᯘࡀඃ༨ࡍࡿࡼ࠺࡟࡞ࡗࡓ㸦ᑠ⯓ 2001㸹⚄ᡞᕷ
2012㸧ࠋࡉࡽ࡟ᒣ㝿࡛ࡣ࢔ࣛ࢝ࢩࢆ୰ᚰ࡜ࡋࡓᖖ⥳ᗈⴥᶞࡀᗈࡀࡗ࡚ࡁ࡚࠸ࡿ㸦ᑠ⯓
2001㸹ᅗ㸳㹢㸧ࠋࢥࢼࣛࡸ࢝ࢩ࡞࡝ࡢሀᯝ㢮ࡣࠊ࢖ࣀࢩࢩࡢ⛅ࡢ୺せ࡞㣗≀࡛࠶ࡿ㸦ᑠ
ᑎࡽ 2013㸧ࠋ࢖ࣀࢩࢩࡣࠊᰤ㣴౯ࡢ㧗࠸ሀᯝ㢮ࢆ㣗࡭ࡿࡇ࡜࡛㣵㈨※ࡀஈࡋ࠸෤Ꮨ࡟ഛ
࠼ࡿࠋᗈⴥᶞࡀඃඛࡍࡿ⌧ᅾࡢභ⏥ᒣࡣࠊ࢖ࣀࢩࢩ࡟࡜ࡗ࡚㇏ᐩ࡞㣵⎔ቃ࡛࠶ࡿ࡜࠸࠼
ࡿࠋ 

රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-10

D

E

F

G

ᅗ  භ⏥ᒣ࿘㎶ࡢ᳜⏕ᅗ
D㸬 ᖺ௦ࠊE㸬 ᖺ௦ࠊF㸬 ᖺ௦ࠊG㸬 ᖺ௨㝆ࡢ᳜⏕ᅗ㸬㣵௜ࡅ⾜Ⅽ࡜⿕ᐖࡢᐇែ
㣵௜ࡅ⾜Ⅽ
㣵௜ࡅ⾜Ⅽࡀ࢖ࣀࢩࢩࡢᕷ⾤ᆅฟἐ࣭⿕ᐖࡢཎᅉ࡜࡞ࡿ⤒⦋࡟ࡣࠊ࢖ࣀࢩࢩࡢᮏ᮶ࡢ
㣗ᛶ࡜㧗࠸Ꮫ⩦⬟ຊࡀ㛵ಀࡍࡿࠋ࢖ࣀࢩࢩࡣ㞧㣗ᛶ࡛࠶ࡿࡀࠊ୺࡟ࡣⲡᮏ㢮ࡸᆅୗⱼ࡞࡝᳜≀㉁ࢆ᥇㣗ࡋࠊ⿵㊊ⓗ࡟᪻⹸㢮࡞࡝ࡢື≀㉁ࠊᏘ⠇ⓗ࡟ሀᯝ㢮ࡸࢱࢣࣀࢥ࡞࡝ᰤ㣴౯ࡢ㧗࠸ࡶࡢࢆዲࡴ㸦ᮅ᪥ 1975㸹ᑠᑎ࡯࠿ 2013㸧ࠋࡑࡢࡓࡵࠊே㛫ࡀ୚࠼ࡿ㣵ࡸࢦ࣑ࡣࠊ࢖ࣀࢩࢩ࡟࡜ࡗ࡚ࡣᴟ➃࡟ᰤ㣴౯ࡀ㧗࠸࡜⪃࠼ࡽࢀࡿࠋࡋࡓࡀࡗ࡚࢖ࣀࢩࢩࡣࠊே
࠿ࡽ͆⡆༢࡟͇ࠊ
͆⨾࿡ࡋ࠸͇㣵ࢆࡶࡽ࠼ࡿࡇ࡜ࢆᏛ⩦ࡍࡿ࡜ࠊᒣᯘෆ࡛⮬↛ࡢࡶࡢࢆ⮬ࡽ᥈ࡋ࡚㣗࡭ࡿࡇ࡜ࢆṆࡵࠊ㣵ࢆᚓࡿࡓࡵ࡟㣵ሙ࡟ᇳ╔ࡍࡿࠋࡸࡀ࡚ேࢆᜍࢀ࡞ࡃ࡞ࡾࠊ
ᕷ⾤ᆅࡀᏳ඲࡛࠶ࡿ࡜Ꮫ⩦ࡍࡿࠋ᭱⤊ⓗ࡟ேࢆく࠺ࡇ࡜࡛㣵ࢆᚓࡿࡇ࡜ࢆぬ࠼ࡿࠋ࢖ࣀ
ࢩࢩࡣẁ㝵ⓗ࡟Ꮫ⩦ࢆ㐍ࡵࠊ⾜ືࢆ࢚ࢫ࣮࢝ࣞࢺࡉࡏࡿࠋࡇࡢࡼ࠺࡟ࡋ࡚Ᏻ᫆࡞㣵௜ࡅ
⾜Ⅽࡀࠊே័ࢀၥ㢟࢖ࣀࢩࢩࢆసฟࡋ࡚࠸ࡿࠋ
࢖ࣀࢩࢩ࡬ࡢ㣵௜ࡅ⾜Ⅽ࡟ᑐࡍࡿၥ㢟ㄆ㆑ࡣᗈࡲࡗ࡚࠸ࡿࡶࡢࡢ㸦⚄ᡞᕷ 2005㸧ࠊ⌧ 

රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-10

ᅾࡶ୍㒊ࡢఫẸࡸⓏᒣ⪅ࡀ㣵௜ࡅ⾜Ⅽࢆ⥆ࡅ࡚࠸ࡿࠋ࢖ࣀࢩࢩࡣࠊᮏ᮶ࡣ㆙ᡄᚰࡀᙉࡃ
ே๓࡟ฟἐࡋ࡞࠸ື≀࡛࠶ࡿࡀ㸦Ụཱྀ 2003㸧ࠊ㣵௜ࡅ⾜Ⅽࡀᖖ⩦ⓗ࡟⾜ࢃࢀ࡚࠸ࡿᆅᇦ
࡛ࡣࠊ㣵ࢆࡶࡽ࠼ࡿ᪥୰࡟ே๓࡟ᇽࠎ࡜ฟἐࡍࡿࠋᅗ㸴ࡢࡼ࠺࡟ẕ࢖ࣀࢩࢩࡀ㣵௜ࡅࡉࢀࡿࡇ࡜࡛ࠊᏊࡶே࠿ࡽ
㣵ࢆࡶࡽ࠺ࡇ࡜ࢆᏛ⩦ࡋ࡚࠸ࡿ࡜⪃࠼ࡽࢀࡿࠋ㣵
௜ࡅ⾜Ⅽࡢ⥅⥆࡟ࡼࡾࠊఱ௦࡟ࡶࢃࡓࡗ࡚ே័ࢀࠊ
ၥ㢟࢖ࣀࢩࢩࡀᖖ࡟⏕ࡳฟࡉࢀ࡚࠸ࡿ≧ἣ࡟࠶ࡿࠋ
ᅗ㣵௜ࡅࡉࢀࡓே័ࢀ࢖ࣀࢩࢩぶᏊ⿕ᐖ≧ἣභ⏥ᒣ⣔༡ഃࡢᕷ⾤ᆅ࡛ࡣࠊ࢖ࣀࢩࢩ࡟ࡼࡿே㌟⿕ᐖࠊ஺㏻஦ᨾ㸦ᅗ㸵㸧ࡸิ㌴஦ᨾࠊ
ࢦ࣑Ⲩࡽࡋ㸦ᅗ㸶㸧ࡸᩜᆅෆ౵ධࠊ⣅ᐖ㸦ᅗ㸷㸧
࡞࡝ࡀⓎ⏕ࡋ࡚࠸ࡿࠋ⿕ᐖࡣࠊᴟᗘ࡟ே័ࢀࡋ
ࡓ≉ᐃಶయ࡟ࡼࡿሙྜ࡜୙≉ᐃಶయ࡟ࡼࡿሙྜ
࡟኱ูࡉࢀࡿࠋ๓⪅ࡢሙྜࡣࠊၥ㢟ಶయࡀ≉ᐃࡢᆅᇦ࡟㧗㢖ᗘ࡟ฟἐࢆ⧞ࡾ㏉ࡋࠊே㌟⿕ᐖ࡞࡝ࡢ῝้࡞⿕ᐖࢆⓎ⏕ࡉࡏࡿഴྥ࡟࠶ࡿࠋᚋ⪅ࡢሙྜࡣࠊண᝿ࡋᚓ࡞࠸ሙᡤ࡟✺Ⓨⓗ࡟ฟἐࡋࠊ
஺㏻஦ᨾࡸิ㌴஦ᨾࢆ㉳ࡇࡋࡓࡾࠊኪ㛫࡟ẸᐙࡢᩜᆅෆࢆⲨࡽࡋࡓࡾࡍࡿࠋ࡝ࡕࡽࡢ≧ἣ࡛ࡶᬑẁ㔝⏕ື≀࡜ࡢ㛵ࢃࡾࡀᕼ࡛ⷧ࠶ࡿ㒔ᕷఫẸࡢከࡃࡣࠊฟἐ᫬ࡢᑐᛂࡸ⿕ᐖᑐ⟇ࡢ᪉ἲࢆ▱ࡽ࡞࠸ࡓࡵࠊ⿕ᐖࡀ⥆Ⓨࡋࡓࡾᣑᩓࡋࡓࡾࡋࡸᅗ  ஺㏻஦ᨾᑐᛂ⌧ሙࡍ࠸ࠋᅗ  ࢦ࣑ࢆⲨࡽࡍ࢖ࣀࢩࢩᅗ  ⣅㸦㯤୸㒊ศ㸧ࡀⓎぢࡉࢀࡓᕷ⾤ᆅ 

රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-10 ᕷ⾤ᆅ࡛ࡢ⿕ᐖ࡜ࡣู࡟ࠊභ⏥ᒣᒣ㯄࡛ࡣࠊ㣵௜ࡅ⾜Ⅽ࡟ࡼࡾே័ࢀࡋࡓ࢖ࣀࢩࢩࡀ
Ⓩᒣ⪅ࢆく࠺⿕ᐖࡀᩓⓎࡋ࡚࠸ࡿࠋ

㸬ఫẸࡢⱞ᝟࣭㏻ሗ࡜᭷ᐖᤕ⋓≧ἣ
1999 ᖺ࠿ࡽ 2011 ᖺ࡟࠾࠸࡚ࠊ⚄ᡞᕷࡢྛ༊ᙺᡤ࡟ᐤࡏࡽࢀࡓ࢖ࣀࢩࢩࡢ⿕ᐖ࡟㛵ࡍ
ࡿ㏻ሗ࣭ⱞ᝟௳ᩘ࡜⚄ᡞᕷෆ࡛᭷ᐖᤕ⋓ࡉࢀࡓ࢖ࣀࢩࢩࡢ㢌ᩘ㸦රᗜ┴ 2012㸧ࢆᅗ㸯㸮࡟♧ࡋࡓࠋ㏻ሗ࣭ⱞ᝟௳ᩘࡣࠊ2004 ᖺࢆࣆ࣮ࢡ࡜ࡋ࡚ࠊ2007 ᖺࡲ࡛ࡣῶᑡഴྥ࡟࠶
ࡗࡓࡀࠊ2008 ᖺ௨㝆࡟෌ࡧቑຍࡋ࡚࠸ࡿࠋ࡜ࡃ࡟ 2011 ᖺࡣࠊ㏻ሗ࣭ⱞ᝟௳ᩘࡀ㐣ཤ᭱ከ࡛࠶ࡗࡓࠋࡓࡔࡋ 2010 ᖺ࡜ 2011 ᖺࡣࠊ୍㒊ࡢᆅᇦ࡛ၥ㢟ಶయࡀฟ⌧ࡋࠊฟἐ࡜⿕
ᐖࢆ⧞ࡾ㏉ࡋࡓࡓࡵ࡟ⱞ᝟࣭㏻ሗ௳ᩘࡀከࡃ࡞ࡗࡓࡇ࡜࡟␃ពࡍࡿᚲせࡀ࠶ࡿࠋ⚄ᡞᕷ඲యࡢ᭷ᐖᤕ⋓ᩘࡣࠊ2007 ᖺࡲ࡛ 300㹼400 㢌ࢆ᥎⛣ࡋ࡚࠸ࡓࡀࠊ2008 ᖺ࡟⣙ 700 㢌࡟ᛴቑࡋࡓᚋࠊከ࠸ࡲࡲ᥎⛣ࡋ࡚࠸ࡿࠋභ⏥ᒣ⣔༡ഃࡢᆅᇦࢆᢤࡁฟࡋࡓᕷ
⾤ᆅᤕ⋓ࢆぢࡿ࡜ࠊ2007 ᖺࡲ࡛ࡣ 200 㢌๓ᚋࢆ᥎⛣ࡋ࡚࠸ࡓࡀࠊ2008 ᖺ௨㝆 300 㢌๓ᚋ࡟ቑຍࡋ࡚࠸ࡿࠋ㐣ཤ᭱኱㢌ᩘ࡜࡞ࡗࡓ 2010 ᖺ࡛ࡣᕷ⾤ᆅᤕ⋓ࡔࡅ࡛⣙ 400 㢌࡟㐩ࡋ࡚࠸ࡿࠋ
භ⏥ᒣ⣔ࡢ࢖ࣀࢩࢩࡢ᥎ᐃ⏕ᜥᩘࡸ⏕ᜥᐦᗘࡣศ࠿ࡗ࡚࠸࡞࠸ࡀࠊ㏆ᖺࡢⱞ᝟࣭㏻ሗ
௳ᩘࠊ᭷ᐖᤕ⋓ᩘࡢቑຍഴྥ࠿ࡽุ᩿ࡍࡿ࡜ࠊಶయᩘࡣቑຍࡋ࡚࠸ࡿ࡜᥎

ࡉࢀࡿࠋᅗ ⚄ᡞᕷෆࡢ࢖ࣀࢩࢩ⿕ᐖ࡟㛵ࡍࡿⱞ᝟࣭㏻ሗ௳ᩘࡣᕷ⾤ᆅᤕ⋓࡜ࡋ࡚ࠊᮾℿ༊ࠊℿ༊ࠊ୰ኸ༊ࠊරᗜ༊ࠊ㡲☻༊࡛᭷ᐖᤕ⋓ࡉࢀࡓ㢌ᩘࠊ ࡣ໭༊ࠊす༊࡛᭷ᐖᤕ⋓ࡉࢀࡓ㢌ᩘࢆ♧ࡍࠋ㸦㛗⏣༊ࠊᆶỈ༊࡛ࡣᙜヱᮇ㛫࡟࢖ࣀࢩࢩ᭷ᐖᤕ⋓ࡣᐇ᪋ࡉࢀ࡚࠸࡞࠸ࠋ
㸧 

රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-10

㸬ᑐ⟇≧ἣ㣵௜ࡅ⚗Ṇ࡜⿕ᐖ㜵Ṇࡢὀពႏ㉳ ࢖ࣀࢩࢩ࡬ࡢ㣵௜ࡅ⚗ṆࡢၨⓎࡸ⿕ᐖ㜵Ṇࡢὀពႏ㉳ࡀࠊ⚄ᡞᕷࢆ୺య࡟ᗈሗㄅࡸ
HP ࡞࡝ࢆ㏻ࡌ࡚ᐇ᪋ࡉࢀ࡚࠸ࡿࠋࡲࡓ㣵௜ࡅ⾜Ⅽࡸ⿕ᐖࡀ῝้࡞ᆅᇦ࡟ࡣࠊ┳ᯈࡢタ⨨ࡀ㐍ࡵࡽࢀ࡚࠸ࡿࠋ㣵௜ࡅ⪅࡟ᑐࡋ࡚ࡣࠊྛ༊ࡢᢸᙜㄢ࡟ࡼࡿὀពࡀᆅ㐨࡟⥆ࡅࡽࢀ࡚࠸ࡿࠋࡇࡢࡼ࠺࡞ὀពႏ㉳࡜ᬑཬၨⓎ࡟ࡼࡾࠊᑡࡋࡎࡘ࢖ࣀࢩࢩၥ㢟ࡢㄆ㆑ࡣᗈࡲࡗ
࡚࠸ࡿࡀࠊ㣵௜ࡅ⾜Ⅽࡢ᰿⤯ࡣ㞴ࡋ࠸≧ἣ࡟࠶ࡿࠋ2011 ᖺ࡜ 2012 ᖺࡢ㣵௜ࡅ┘どဨࡢタ⨨࡛ࡣࠊ㣵௜ࡅࡸ⿕ᐖࡀ㍍ῶࡋࡓᆅᇦࡀㄆࡵࡽࢀࡿ࡞࡝ࡢຠᯝࡀ࠶ࡗࡓࠋ
᭷ᐖᤕ⋓࡜⥭ᛴᑐᛂ ⚄ᡞᕷ࡛ࡣࠊ࢖ࣀࢩࢩ⿕ᐖࡢ┤᥋ⓗ࡞ᑐ⟇࡜ࡋ࡚ࠊ᭷ᐖᤕ⋓࡜࢖ࣀࢩࢩ⥭ᛴᑐ⟇༠ຊဨ࡟ࡼࡿ⥭ᛴᑐᛂࡀᐇ᪋ࡉࢀ࡚࠸ࡿࠋࡇࡢᑐᛂ᫬࡟୰ᒣ㛫ᆅᇦ࡜ࡣ␗࡞ࡿ㒔ᕷ⎔ቃ࡞ࡽ࡛ࡣࡢᘢᐖࡀⓎ⏕ࡋ࡚࠸ࡿࠋ᭷ᐖᤕ⋓࡟ࡘ࠸࡚ࡣࠊ㖠ჾࡀ౑⏝࡛ࡁ࡞࠸ࡔࡅ࡛࡞ࡃࠊఫ
Ꮿࡀ❧ࡕ୪ࢇ࡛ᅵᆅᡤ᭷ࡀ」㞧࡛࠶ࡿࡓࡵࠊࢃ࡞ࡀタ⨨࡛ࡁࡿሙᡤࡶ㝈ࡽࢀ࡚࠸ࡿࡇ࡜
ࡀᤕ⋓άືࢆᅔ㞴࡟ࡋ࡚࠸ࡿࠋࡉࡽ࡟㒔ᕷఫẸࡢᤕ⋓࡟ᑐࡍࡿ᎘ᝏឤࡶ࠶ࡾࠊ⿕ᐖࡀⓎ⏕ࡋ࡚࠸࡚ࡶఫẸࡢ⌮ゎࡀᚓࡽࢀ࡞࠸ሙᡤ࡛ࡣᤕ⋓ࡀ㞴ࡋ࠸≧ἣ࡟࠶ࡿࠋࡲࡓ㣵௜ࡅ⪅࡞࡝࡟ࡼࡾࠊࢃ࡞ࡀቯࡉࢀ࡚ࠊᤕ⋓ࢆጉᐖࡉࢀࡿࡇ࡜ࡶ࠶ࡿࠋࡋࡓࡀࡗ࡚ࠊᤕ⋓ᚑ஦⪅
࡟ࡣᵝࠎ࡞㠃࡛ࡢ㓄៖࡜㧗࠸ᤕ⋓ᢏ⾡ࡀồࡵࡽࢀࡿ≧ἣ࡟࠶ࡿࠋ ⥭ᛴᑐ⟇༠ຊဨࡣࠊ✺Ⓨⓗ࡞࢖ࣀࢩࢩࡢฟἐࡸ⿕ᐖ࡬ࡢᑐᛂࢆᐇ᪋ࡋ࡚࠸ࡿࠋ2008
㹼2011 ᖺ࡟⥭ᛴᑐ⟇༠ຊဨࡀᑐᛂࡋࡓ 149 ௳ࡢฟື᫬㛫ࡣ඲య࡛ࡣ᪥୰ࡀከ࠿ࡗࡓࠋ
ࡋ࠿ࡋே㌟⿕ᐖࠊ஺㏻஦ᨾࠊิ㌴஦ᨾ࡞࡝㏿ࡸ࠿࡞ᑐᛂࡀᚲせ࡜࡞ࡿ⥭ᛴᛶࡢ㧗࠸ฟື࡟㝈ࡗ࡚ࡳࡿ࡜ࠊኪ㛫࣭῝ኪࡢฟືࡀከࡃ࡞ࡗࡓ㸦ᅗ㸯㸯㸧ࠋ⥭ᛴ࡟ᑐᛂࡋ࡞ࡅࢀࡤ࡞
ࡽ࡞࠸᱌௳࡯࡝ࠊᑐᛂ᫬ࡢ㈇ᢸࡀ኱ࡁ࠸ࡇ࡜ࡀ♧၀ࡉࢀࡓࠋᅗ ࢖ࣀࢩࢩ⥭ᛴᑐ⟇༠ຊဨࡢฟື᫬㛫ᕥࡣ඲࡚ࡢᑐᛂࡢฟື᫬㛫ࠊྑࡣ⥭ᛴᛶࡢ㧗࠸ࡶࡢ࡜ࡋ࡚ࠊே㌟⿕ᐖࠊ஺㏻஦ᨾࠊ
ิ㌴஦ᨾࡢᑐᛂ᫬ࡢฟື᫬㛫ࢆ♧ࡍࠋ㸦⚄ᡞ㎰ᯘỈ⏘஦ົᡤᥦ౪㸧 

රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-10

㸬ࡲ࡜ࡵ
㣵௜ࡅၥ㢟 㔝⏕ື≀࡬ࡢ㣵௜ࡅၥ㢟ࡣࠊභ⏥ᒣ࢖ࣀࢩࢩࡔࡅ࡛࡞ࡃࠊ▱ᗋᅜ❧බᅬࡢࣄࢢ࣐
㸦Ursus arctos㸧ࠊ᪥ගᕷࡸ⟪㠃ᕷࡢࢽ࣍ࣥࢨࣝ㸦Macaca fuscata㸧࡞࡝࡛ࡶ῝้࡛࠶ࡿ㸦⟪㠃ᕷ 2006㸹᪥ගᕷ 2009㸹▱ᗋ㈈ᅋ HP㸧ࠋၥ㢟ࡢඹ㏻Ⅼࡢ 1 ࡘ┠ࡣࠊᬑẁ㔝⏕ື≀࡜ࡢ㛵ࢃࡾࡀᑡ࡞ࡃ௜ࡁྜ࠸᪉ࢆ▱ࡽ࡞࠸ேࠎࡀᏳ᫆࡞⪃࠼࡛㣵௜ࡅࡍࡿࡇ࡜࡟
࠶ࡿࠋ࡜ࡃ࡟㒔ᕷఫẸࡣࠊ㔝⏕ື≀࡜≟ࡸ⊧࡞࡝ࡢక౶ື≀࡜ࡢ㐪࠸ࡀ⌮ゎ࡛ࡁࡎࠊྠ
ᵝࡢឤ᝟⛣ධࡸᑐᛂࢆ⾜࠸ࡸࡍ࠸ࡇ࡜ࡀᣦ᦬ࡉࢀ࡚࠸ࡿ
㸦ᓊᮏ 2001㸹York and Andreas
2008㸧ࠋඹ㏻Ⅼࡢ 2 ࡘ┠ࡣࠊ㣵௜ࡅ⪅ᮏேࡣ㔝⏕ື≀࠿ࡽࡢ┤᥋ⓗ࡞⿕ᐖࢆཷࡅ࡟ࡃ࠸
ࡓࡵࠊ㣵௜ࡅ⾜Ⅽࡢၥ㢟ㄆ㆑ࡀప࠸ࡇ࡜࡛࠶ࡿࠋ㒔ᕷఫẸ࡛ࡶ⿕ᐖࢆཷࡅࡓேࡣ✚ᴟⓗ࡞ᑐ⟇࡬ࡢ⌮ゎࢆ♧ࡍࡀࠊ⿕ᐖࢆཷࡅ࡞࠸ேࡢ⌮ゎࡣప࠸㸦ᓊᮏ 2001㸹York andAndreas 2008㸧ࠋࡋࡓࡀࡗ࡚ࠊ㣵௜ࡅ⪅ᮏேࡀ⿕ᐖࢆཷࡅ࡞࠸㝈ࡾࠊ㣵௜ࡅࡢ᰿⤯ࡀ㞴ࡋ࠸ࡢࡀ⌧≧࡛࠶ࡿࠋேཱྀࡀᐦ㞟ࡍࡿ㒔ᕷ⎔ቃࡣࠊ୰ᒣ㛫ᆅᇦ࡜ࡣ␗࡞ࡾࠊఫẸ඲యࡢ୰࡛ࡣ⿕ᐖ⪅ࡶ㣵௜ࡅ⪅ࡶ୍㒊ศ࡟㝈ࡽࢀࠊ኱༙ࡣၥ㢟࡟↓㛵ಀࠊ↓㛵ᚰ࡞ே࡜࡞ࡿࠋ
㒔ᕷ⎔ቃ࡞ࡽ࡛ࡣࡢ」㞧࡞♫఍ᵓ㐀ࡀࠊఫẸࡀ୍୸࡜࡞ࡗࡓᑐ⟇ࡢ᥎㐍ࢆ㞴ࡋࡃࡋ࡚࠸ࡿせᅉࡢ୍ࡘ࡛࠶ࡿࠋࡇࢀࡽࢆᨵၿࡋ࡚࠸ࡃࡓࡵ࡟ࡣࠊ㔝⏕ື≀࡜ࡢ㐺ษ࡞㛵ࢃࡾ᪉ࡸ㊥㞳ࡢ⨨ࡁ᪉࡟ࡘ࠸࡚ࠊᬑཬၨⓎάືࢆᆅ㐨࡟⥆ࡅ࡚࠸ࡃࡇ࡜ࡀ㔜せ࡛࠶ࡿ࡜⪃࠼ࡿࠋ 㔝⏕ື≀࡬ࡢ㣵௜ࡅ⚗Ṇ᮲౛ࡣࠊ⚄ᡞᕷࡸすᐑᕷ௨እ࡛ࡶ᪋⾜ࡉࢀ࡚࠸ࡿࠋ᮲౛㐪཯⪅࡟ᑐࡍࡿ⨩๎つᐃ࡟╔┠ࡍࡿ࡜ࠊ⟪㠃ᕷࢧࣝ㣵ࡸࡾ⚗Ṇ᮲౛㸦⟪㠃ᕷ 2009㸧࡛ࡣ 1୓෇௨ୗࠊ⟪㠃ᕷ࢝ࣛࢫ࡟ࡼࡿ⿕ᐖࡢ㜵Ṇཬࡧ⏕ά⎔ቃࢆᏲࡿ᮲౛㸦⟪㠃ᕷ 2011㸧࡛ࡣ 10 ୓෇௨ୗࡢ⨩㔠ࢆࠊ᪥ගᕷࢧࣝ㣵௜ࡅ⚗Ṇ᮲౛㸦᪥ගᕷ 2006㸧࡛ࡣẶྡබ⾲ࢆᐃࡵ࡚࠸ࡿࠋ⌧ᅾࠊභ⏥ᒣ࢖ࣀࢩࢩ࡟ࡼࡿ⿕ᐖ࡛ࡣࠊ஺㏻஦ᨾࡸධ㝔ࢆせࡍࡿ῝้࡞ே㌟
⿕ᐖࡀⓎ⏕ࡋ࡚࠸ࡿ㸦㎷࣭ᶓᒣ 2014㸧ࠋࡇࡢࡼ࠺࡞≧ἣ࡟ᑐࡋ࡚ࠊッゴၥ㢟࡟࡞ࡗ࡚ࡶ
࠾࠿ࡋࡃ࡞࠸࡜㆙㚝ࢆ㬆ࡽࡍኌࡸ⨩๎つᐃ࡞࡝ࢆᮃࡴኌࡶከࡃ⪺࠿ࢀ࡚࠸ࡿࠋ࢖ࣀࢩࢩ
ࡣయ㔜ࡀ 100 ਻ࢆ㉸࠼ࡿࡇ࡜ࡶᑡ࡞ࡃ࡞ࡃ㸦㎷࣭ᶓᒣ 2014㸧ࠊࡲࡓࠊᡂ⋇ࡢ࢜ࢫ࡞࡝≟ṑࡀⓎ㐩ࡋࡓಶయ࡜㖊ྜࢃࡏ࡚Ṛஸ࡟⮳ࡿ஦౛ࡀ௚ᆅᇦ࡛ࡣⓎ⏕ࡋ࡚࠸ࡿࠋ㛵ಀᶵ㛵ࡢᵝࠎ࡞ດຊ࡟ࡼࡗ࡚ࡶ⣙ 50 ᖺ࡟ࡶࢃࡓࡿ㣵௜ࡅ⾜Ⅽࡣ࡞ࡃ࡞ࡿẼ㓄ࡀ࡞࠸ࡇ࡜ࡸ῝้࡞஦ᨾࢆண㜵ࡍࡿほⅬ࠿ࡽࡶࠊࡉࡽ࡟㋃ࡳ㎸ࢇࡔᑐ⟇ࡀᚲせ࡜࡞ࡗ࡚࠸ࡿࠋ
භ⏥ᒣࡢ᳜⏕ኚ໬࡜࢖ࣀࢩࢩࡢ⏕ᜥᩘࡢ㛵ಀ 㐣ཤ 100 ᖺ㛫ࡢභ⏥ᒣࡢ᳜⏕⎔ቃࡣࠊேࡢᡭࡀຍࢃࡗࡓࡇ࡜࡟ࡼࡾࠊ⮬↛ࡢ᳜⏕㑄⛣ࡼࡾࡶᴟࡵ࡚㏿࠸ࢫࣆ࣮ࢻ࡛๻ⓗ࡞ኚ໬ࢆ㐙ࡆࡓ㸦ᅗ㸳㸹⚄ᡞᕷ 2012㸧ࠋ࢖ࣀࢩࢩࢆ
ࡣࡌࡵ㔝⏕ື≀ࡀ⏕ᜥ࡛ࡁࡿࡼ࠺࡞⎔ቃ࡛ࡣ࡞࠿ࡗࡓභ⏥ᒣࡣࠊ⌧ᅾࡣከࡃࡢ㔝⏕ື≀ࢆ㣴࠼ࡿዲ㐺࡞⏕ᜥ⎔ቃ࡬࡜ኚㇺࡋ࡚࠸ࡿࠋᮏ᮶࢖ࣀࢩࢩࡣࠊ⦾Ṫ⋡ࡀ㧗ࡃ㸦㎷࣭ᶓᒣ
2014㸧ᤕ⋓ࡀ⾜ࢃࢀ࡞ࡅࢀࡤࠊᛴ⃭࡟ቑຍࡍࡿື≀✀࡛࠶ࡿࠋ⌧ᅾࡢරᗜ┴඲యࡢഴྥ࡜ࡋ࡚ࡶ㧗࠸ᤕ⋓ᅽ࡟ࡶ࠿࠿ࢃࡽࡎࠊಶయᩘࡢᛴ⃭࡞ቑຍࡀ♧၀ࡉࢀ㸦ᶓᒣ࣭㎷ 2014㸧ࠊ 

රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-10㎰ᴗ⿕ᐖࡶᣑ኱㸦ᯇᮏ࡯࠿ 2014㸧ࡋ࡚࠸ࡿࠋࡇࢀࡽࡢ≧ἣࢆ㋃ࡲ࠼ࡿ࡜ࠊ⌧ᅾࡢභ⏥ᒣ࡟ࡣ࢖ࣀࢩࢩࡢಶయᩘࡀῶࡿせᅉࡀぢᙜࡓࡽࡎࠊ௒ᚋࡶ⏕ᜥᩘࡣቑຍࡍࡿ࡜ண᝿ࡉࢀ
ࡿࠋࡉࡽ࡟㣵௜ࡅ⾜Ⅽ࡟ࡼࡾࠊᮏ᮶ࡢ⮬↛ࡢᦤ⌮࡛࠶ࢀࡤῈửࡉࢀࡿಶయࡶ⏕ࡁṧࡿྍ⬟ᛶࡀᣲࡆࡽࢀࡿࠋࡇࡢࡼ࠺࡟ࡋ࡚භ⏥ᒣࡢ࢖ࣀࢩࢩࡢ⏕ᜥᩘࡀቑ࠼ࡓሙྜࠊ୍ᐃ๭ྜ
ࡀᕷ⾤ᆅ࡟ฟἐࡍࡿྍ⬟ᛶࡀ㧗࠸ࠋ⌧࡟㏻ሗ࣭ⱞ᝟௳ᩘࡸ᭷ᐖᤕ⋓ᩘࡣቑຍഴྥ࡟࠶ࡿ
㸦ᅗ㸯㸮㸧ࠋ௒ᚋࡢᑐ⟇࡜ࡋ࡚ࠊၥ㢟ಶయࡢ᭷ᐖᤕ⋓ࡔࡅ࡛࡞ࡃࠊవ๫ศࡢ㛫ᘬࡁࠊࡘࡲࡾභ⏥ᒣ࡟࠾ࡅࡿ࢖ࣀࢩࢩࡢಶయᩘ⟶⌮ࡀᚲせ࡞᫬ᮇ࡟ࡁ࡚࠸ࡿ࡜⪃࠼ࡿࠋฟἐ࣭⿕ᐖ᫬ࡢᑐᛂᑐ⟇ᕷ⾤ᆅ࡟ฟἐࡋࡓಶయࡸே㌟஦ᨾࢆ㉳ࡇࡋࡓಶయ࡬ࡢᑐᛂࡣࠊ᭷ᐖᤕ⋓ᚑ஦⪅ࡸ࢖ࣀࢩࢩ⥭ᛴᑐ⟇༠ຊဨࡀᢸࡗ࡚࠸ࡿࠋࡓࡔࡋᕷ⾤ᆅ࡛ᑐᛂࡍࡿሙྜ࡟ࡣࠊ஧ḟ⿕ᐖࢆ㜵ࡄ
ࡓࡵࡢ஺㏻ᩚ⌮ࠊఫẸࡢΰ஘ࢆ㜵ࡄࡓࡵࡢㄝ᫂ࠊ㐣๫࡞࣐ࢫࢥ࣑ሗ㐨ࡢᅇ㑊࡞࡝ࠊ⾜ᨻࡸ㆙ᐹࡢ༠ຊࡀ୙ྍḞ࡛࠶ࡿࠋ㐃ᦠయไࢆ☜ಖࡍࡿࡓࡵ࡟ࠊࡓ࡜࠼ࡤࠊᮐᖠᕷࡸ㤶ᕝ┴࡛ࡣࠊ㔝⏕ື≀ࡢฟἐ᫬ࡢᑐᛂ࣐ࢽࣗ࢔ࣝࡸᏳ඲ᑐ⟇ࡢᡭᘬࡁࢆసᡂࡋ࡚࠸ࡿ㸦ᮐᖠᕷ
2010㸹㤶ᕝ┴ 2011㸧ࠋࡇࡢ୰࡛ࡣࠊྛᶵ㛵ࡢᙺ๭ศᢸ࡜㈐௵ࡢᡤᅾࢆ᫂ࡽ࠿࡟ࡋࠊฟἐ
᫬ࡢ㐃⤡యไࢆᩚ࠼࡚࠸ࡿࠋභ⏥ᒣࡢ࢖ࣀࢩࢩࡢᕷ⾤ᆅฟἐࡸ⿕ᐖࡣࠊ௒ᚋࡶⓎ⏕ࡀ⥆ࡃ࡜ணࡉࢀࡿࡇ࡜࠿ࡽࠊฟἐ᫬࡟㎿㏿࠿ࡘ㐺ษ࡞ᑐᛂయไࢆᩚഛࡍࡿࡇ࡜ࡀ㔜せ࡛࠶ࡿ࡜⪃࠼ࡿࠋㅰ㎡ ᮏ✏ᇳ➹࡟࠶ࡓࡾࠊከ኱࡞ࡿࡈ༠ຊࢆ㡬ࡁࡲࡋࡓ⚄ᡞᕷ⏘ᴗ᣺⯆ᒁ㎰ᨻ㒊ィ⏬ㄢࠊྛ༊ࡢࡲࡕ࡙ࡃࡾㄢ࠾ࡼࡧࡲࡕ࡙ࡃࡾ᥎㐍ㄢࠊ⚄ᡞ㎰ᯘỈ⏘᣺⯆஦ົᡤࡢࡳ࡞ࡉࡲ࡟῝ࡃᚚ♩⏦ࡋୖࡆࡲࡍࠋࡲࡓࠊ㐣ཤࡢ࢖ࣀࢩࢩ⿕ᐖࡸ᭷ᐖᤕ⋓ࠊභ⏥ᒣࡢ᳜⏕࡟㛵ࡋ࡚ࡢ᝟ሗ࠾ࡼࡧ෗┿ࢆࡈᥦ౪㡬ࡁࡲࡋࡓỌᓥ᫛஧㑻Ặࠊᒣ᰿ㅬḟẶ࡟῝ࡃឤㅰ⮴ࡋࡲࡍࠋᮏ◊✲ࡣࠊᖹᡂ 25㹼27 ᖺᗘ JSPS ⛉◊㈝㸦25450478㸧ࠕ㒔ᕷ⎔ቃ࡟౵ධࡍࡿ࢖ࣀࢩࢩࡢࣜ
ࢫࢡศᯒ࡜ࣜࢫࢡᅇ㑊ᡭἲࡢ᳨ウࠖࡢຓᡂࢆཷࡅࡲࡋࡓࠋ

ᘬ⏝ᩥ⊩ᮅ᪥⛱㸬1975㸬⊁⊟ᮇ࡟࠾ࡅࡿ࢖ࣀࢩࢩࡢ⫶ෆᐜ㸬့ங㢮ື≀Ꮫ㞧ㄅ㸪6(3)㸸115-120㸬Ụཱྀ♸㍜㸬2003㸬࢖ࣀࢩࢩ࠿ࡽ⏣⏿ࢆᏲࡿ ࠾ࡶࡋࢁ⏕ែ࡜࠿ࡋࡇ࠸㜵ࡂ᪉㸬㎰ᒣ⁺ᮧᩥ໬༠఍㸪147pp㸪ᮾி㸬
Hirotani,A.

andNakatani,

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andinter-grouprelationships of Japanese wild boars (Sus scrofa leucomystax) in the Rokko
mountain area. Ecol res. 2(1): 77-84.
රᗜ┴㸬2012㸬➨㸰ᮇ࢖ࣀࢩࢩಖㆤ⟶⌮ィ⏬㸬රᗜ┴㸪18pp㸬

 

රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-10

රᗜ┴㸬2012㸬➨㸯㸯ḟ㫽⋇ಖㆤ஦ᴗィ⏬᭩㸬රᗜ┴㸪66pp㸬㤶ᕝ┴㸬2011㸬࢖ࣀࢩࢩ➼ࡀฟἐࡋࡓ࡜ࡁࡢᑐᛂ࣐ࢽࣗ࢔ࣝ㸬㤶ᕝ┴࢖ࣀࢩࢩ➼ᑐ⟇㐃⤡఍㆟㸪15pp㸬http://www.pref.kagawa.lg.jp/kankyo/data/topics/pdf/manual.pdf㸬2014.
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ᓊᮏ┿ᘪヂ㸬2001㸬19 ❶㒔ᕷ㔝⏕ື≀ࡢ⟶⌮㸬㕥ᮌṇႹ⦅ヂ㸪᪥ᮏ㔝⏕ື≀་Ꮫ఍࣭㔝⏕⏕≀ಖㆤᏛ఍┘ಟ㸪㔝⏕ື≀ࡢ◊✲࡜⟶⌮ᢏ⾡㸬pp.603-634㸪ᩥỌືฟ∧㸪ᮾி㸬ᑠ⯓ㄋ἞㸬2001㸬භ⏥ᒣᆅࡢ᳜≀࡜᳜ᯘ᳜⏕㸬⮬↛⎔ቃ࢛࢘ࢵࢳࣥࢢࠕභ⏥ᒣࠖ㸪රᗜ┴❧ே࡜⮬↛ࡢ༤≀㤋ࠕභ⏥ࠖ◊✲ࢢ࣮ࣝࣉ⦅㸪pp.44-48㸬⚄ᡞ᪂⪺⥲ྜฟ∧ࢭࣥࢱ࣮㸪⚄ᡞ㸬ᑠᑎ♸஧࣭⚄ᓮఙኵ࣭▼ᕝᑦே࣭ⓙᕝᬗᏊ㸬2013㸬ᓥ᰿┴▼ぢᆅ᪉࡟࠾ࡅࡿ࢖ࣀࢩࢩ㸦 Sus

scrofa㸧ࡢ㣗ᛶ࡜ᰤ㣴≧ែ㸬့ங㢮⛉Ꮫ㸪53(2)㸹279-288㸬⚄ᡞᕷ㸬2001㸬2001 ᖺᥦゝ᭩㸬⚄ᡞᕷ㸪http://www.city.kobe.lg.jp/information/proj+6ect/industry/boar/teigen.html㸬2014.2.5 accessed.
⚄ᡞᕷ㸬2002㸬⚄ᡞᕷ࠸ ࡢ ࡋ ࡋ ࡢ ฟ ἐ ཬ ࡧ ࠸ ࡢ ࡋ ࡋ ࠿ ࡽ ࡢ ༴ ᐖ ࡢ 㜵 Ṇ ࡟ 㛵 ࡍ ࡿ ᮲౛㸬⚄ᡞᕷ㸪http://www.city.kobe.lg.jp/information/project/industry/boar/joreihonbun.
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⚄ᡞᕷ㸬2005㸬2004 ᖺᥦゝ᭩㸬⚄ᡞᕷ㸪http://www.city.kobe.lg.jp/information/project/
industry/boar/H16teigen.html㸬2014.2.5 accessed.
⚄ᡞᕷ㸬2012㸬භ⏥ᒣ᳃ᯘᩚഛᡓ␎ࠕ㒔ᕷᒣࠖභ⏥ᒣ࡜ேࡢᬽࡽࡋ࡜ࡢ᪂ࡓ࡞㛵ࢃࡾ࡙ࡃࡾ㸬122pp㸬⚄ᡞᕷᘓタᒁබᅬ◁㜵㒊භ⏥ᒣᩚഛᐊ㸬ᯇᮏᓫ࣭ᆏ⏣ᏹᚿ࣭ᓊᮏᗣ㄃㸬2014㸬රᗜ┴࡟࠾ࡅࡿࢽ࣍ࣥ࢖ࣀࢩࢩࡢ⏕ᜥศᕸ࡜㎰ᴗ⿕ᐖࡢ㛵ಀ㸬ࠕරᗜ┴࡟࠾ࡅࡿࢽ࣍ࣥ࢖ࣀࢩࢩࡢ⟶⌮ࡢ⌧≧࡜ㄢ㢟㸬ࠖ㸪රᗜ࣡࢖ࣝࢻࣛ࢖
ࣇࣔࣀࢢࣛࣇ 6 ྕ㸪pp.9-20㸬රᗜ┴᳃ᯘື≀◊✲ࢭࣥࢱ࣮㸬⟪㠃ᕷ㸬2013㸬⟪㠃ᕷ࢝ࣛࢫ࡟ࡼࡿ⿕ᐖࡢ㜵Ṇཬࡧ⏕ά⎔ቃࢆᏲࡿ᮲౛㸬⟪㠃ᕷ㸪http://www.
city. minoh.lg.jp/animal/karasu/documents/karasujyourei.pdfࠊ2014.2.11 accessed㸬⟪㠃ᕷ㸬2009㸬⟪㠃ᕷࢧࣝ㣵ࡸࡾ⚗Ṇ᮲౛㸬⟪㠃ᕷ㸪http://www1.g-reiki.net/minoh/reiki_honbun/at00008741.htmlࠊ2014.2.11 accessed㸬୕㇂㞞⣧㸬2001㸬භ⏥ᒣࡢ࡯ங㢮㸬⮬↛⎔ቃ࢛࢘ࢵࢳࣥࢢࠕභ⏥ᒣࠖ㸬රᗜ┴❧ே࡜⮬↛ࡢ༤≀㤋ࠕභ⏥ࠖ◊✲ࢢ࣮ࣝࣉ⦅㸪pp.85-94, 105-109㸬⚄ᡞ᪂⪺⥲ྜฟ∧ࢭࣥࢱ࣮㸪⚄ᡞ㸬
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inosisiesayarikinsijyourei.pdfࠊ2014.2.5 accessed.

 

රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-10⋢㉳ᙲ୕㸬1997㸬භ⏥ᒣ༤≀ㄅ㸬246pp㸪ࡢࡌࡂࡃᩥᗜ㸪⚄ᡞ᪂⪺⥲ྜฟ∧ࢭࣥࢱ࣮㸪⚄ᡞ㸬⏣୰ᫍᒣ㸬1986㸬භ⏥ᒣࡢ⊦࡜Ἑ㮵㸬ࢨ࣭භ⏥ᒣ Ⓩࡾࡲࡃࡿャ࠸ࡲࡃࡿ㸳㸪40pp㸪⚄ᡞᕷ❧ᅗ᭩㤋㒓ᅵཧ⪃㈨ᩱ㸪⚄ᡞ㸬㎷▱㤶࣭ᶓᒣ┿ᘪ㸬2012㸬㒔ᕷ࡟ᾐධࡍࡿ㔝⏕ື≀࡟ᑐᛂࡍࡿࡓࡵࡢయไ࡙ࡃࡾࢆ⪃࠼ࡿ(ࢸ࣮࣐ࢭࢵࢩࣙࣥ㸪<≉㞟 2>➨ 17 ᅇ㔝⏕⏕≀ಖㆤᏛ఍໭ᾏ㐨኱఍ሗ࿌)㸬࣡࢖ࣝࢻࣛ࢖

ࣇ࣭ࣇ࢛࣮࣒ࣛ㸪16(2)㸸16-17㎷▱㤶࣭ᶓᒣ┿ᘪ㸬2014㸬භ⏥ᒣ࢖ࣀࢩࢩࡢၥ㢟ಶయࡢ≉ᚩ㸬ࠕරᗜ┴࡟࠾ࡅࡿࢽ࣍ࣥ࢖ࣀ
ࢩࢩࡢ⟶⌮ࡢ⌧≧࡜ㄢ㢟㸬ࠖ㸪රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6 ྕ㸪pp.135-142㸬රᗜ┴᳃ᯘື≀◊✲ࢭࣥࢱ࣮㸬ᮐᖠᕷ㸬2010㸬ᖹᡂ 22 ᖺᗘ ᮐᖠᕷ⥭ᛴ㞠⏝๰ฟ᥎㐍஦ᴗ㔝⏕ື≀࡟ࡼࡿᕷ⾤ᆅ➼࡬ࡢ౵ධ⤒㊰ㄪᰝ࠾ࡼࡧ౵ධ㜵Ṇ⟇ࡢㄪᰝ࣭◊✲ᴗົሗ࿌᭩㸬ᮐᖠᕷ㸪http://www.city.
sapporo.jp /kurashi/animal/choju/shika/kenkyu/㸬2014.2.11 accessed.
▱ᗋ㈈ᅋ㸬▱ᗋࡢㄢ㢟㸬http://www.shiretoko.or.jp/shiretoko/㸬2014.3.14 accessed.
ᶓᒣ┿ᘪ㸬2014㸬රᗜ┴࡟࠾ࡅࡿࢽ࣍ࣥ࢖ࣀࢩࢩࡢಖㆤ⟶⌮ࡢ⌧≧࡜ㄢ㢟㸬ࠕරᗜ┴࡟࠾ࡅ
ࡿࢽ࣍ࣥ࢖ࣀࢩࢩࡢ⟶⌮ࡢ⌧≧࡜ㄢ㢟㸬ࠖ㸪රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6 ྕ㸪pp.1-8㸬රᗜ┴᳃ᯘື≀◊✲ࢭࣥࢱ࣮㸬York, K. and Andreas, K. 2008. Conflicts, crises and challenges: Wild boar in the Berlincity – a social empirical and statistical survey. Nat. Croat. 17(4): 233-246.
࿴⏣ඞᕫ㸬1997㸬ࡴ࠿ࡋࡢ⚄ᡞ ⤮ࡣࡀࡁ࡟ぢࡿ᫂♧࣭኱ṇ࣭᫛࿴ึᮇ㸬269pp㸪⚄ᡞ᪂⪺⥲ྜฟ∧ࢭࣥࢱ࣮㸪⚄ᡞ㸬 

රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-11

➨ 㸯㸯 ❶            භ⏥ᒣ࢖ࣀࢩࢩࡢၥ㢟ಶయࡢ≉ᚩ㎷ ▱㤶 1࣭ᶓᒣ┿ᘪ 1,2
せ Ⅼ࣭ ၥ㢟ಶయࡢ≉ᚩࢆᢕᥱࡍࡿࡓࡵ࡟ࠊ2009 ᖺ࠿ࡽ 2011 ᖺ࡟⚄ᡞᕷ࡟࡚᭷ᐖᤕ⋓ࡉࢀࡓၥ㢟ಶయ 2 㢌ࠊิ㌴࣭஺㏻஦ᨾಶయ 5 㢌ࠊィ 7 㢌ࢆศᯒࡋࡓࠋ࣭ ศᯒಶయ 7 㢌୰ 6 㢌ࡢ⫶ෆᐜ≀ࡼࡾேᕤ≀ࡀ᳨ฟࡉࢀࡓࡇ࡜࠿ࡽࠊࡇࢀࡽࡢಶయ
ࡣேᕤ≀ࢆ㣵࡜ㄆ㆑ࡋᇳ╔ࡋ࡚࠸ࡓಶయ࡛࠶ࡗࡓ࡜᥎ᐹࡉࢀࡓࠋ࣭ ศᯒಶయ࡟ᖺ㱋ࠊᛶูࡢ೫ࡾࡣㄆࡵࡽࢀ࡞࠿ࡗࡓࠋ⌧᫬Ⅼ࡛ࡣࠊఱࡽ࠿ࡢᶵ఍࡟
㣵௜ࡅࡉࢀࡓࡾࠊࢦ࣑ࢆ㣵࡜ㄆ㆑ࡋࡓࡾࡋࡓಶయࡀࠊᛶࠊᖺ㱋࡟㛵ಀ࡞ࡃၥ㢟ಶయ࡟࡞ࡾ࠺ࡿ࡜⪃࠼ࡽࢀࡓࠋ࣭ 㣵௜ࡅ⾜Ⅽ࡟ࡼࡾே័ࢀࡋࡓ⤖ᯝࠊே㌟⿕ᐖࢆከⓎࡉࡏࡓ 2 㢌ࡣࠊ4 ṓ࡜ 5 ṓࡢᡂ⋇࡛࠶ࡗࡓࠋࡇࡢࡇ࡜࠿ࡽᏊ࡝ࡶࡢ㡭࠿ࡽ㣵௜ࡅࡉࢀ࡚࠸ࡓಶయࡣࠊᡂ⋇࡟࡞ࡿ࡟ࡘࢀࠊᏛ⩦ࡀ㐍ࡳ⾜ືࡀ࢚ࢫ࣮࢝ࣞࢺࡉࡏ࡚῝้࡞⿕ᐖࢆ㉳ࡇࡍၥ㢟ಶయ࡟࡞ࡿྍ⬟ᛶࡀ♧၀ࡉࢀࡓࠋ


Key words㸸භ⏥ᒣࠊே㌟⿕ᐖࠊิ㌴࣭஺㏻஦ᨾࠊၥ㢟ಶయ㸬ࡣࡌࡵ࡟
 ㏆ᖺࠊ඲ᅜྛᆅ࡛㔝⏕ື≀ࡀᕷ⾤ᆅ࡟ฟἐࡋ࡚⿕ᐖࢆⓎ⏕ࡉࡏ࡚࠸ࡿࠋࡑࡢከࡃࡀ✺Ⓨⓗ࡞ฟἐ࡛࠶ࡾࠊ⥭ᛴᑐᛂ࡜ࡋ࡚ࠊ㆙ᐹࠊ⾜ᨻࠊ⊟཭఍࡞࡝ࡀᑐฎࡋ࡚࠸ࡿ㸦ᮐᖠᕷ
2010㸹㤶ᕝ┴ 2011㸧ࠋᤕࡲࡗࡓ㔝⏕ື≀ࡢ኱༙ࡣࠊ௜㏆ࡢᒣ࡟ᨺ⋇ࡉࢀ࡚࠸ࡿࠋࡓ࡜࠼
ࡤᮐᖠᕷࡣࠊᕷ⾤ᆅ࡟ฟἐࡋࡓ࢚ࢰࢩ࢝㸦Cervus nippon yesoensis㸧࡟ᑐࡋ࡚ࠊ
ࠕ↓஦࡟ࡶ࡜ࡢᒣᯘ࡬ᡠࡍࠖࡇ࡜ࢆ➨୍࡟యไࡀྲྀࡽࢀ࡚࠸ࡿ㸦ᮐᖠᕷ 2010㸧ࠋࡇࡢࡼ࠺࡞⫼ᬒ࠿ࡽࠊᕷ⾤ᆅฟἐಶయࡀẅฎศࡉࢀࡿᶵ఍ࡣᑡ࡞ࡃࠊᖺ㱋ࡸ㣗ᛶࠊ⦾Ṫ≧ἣࢆ┤᥋ほᐹࡋ࡚ศᯒࡍࡿࡇ࡜࡛ࠊࡑࡢ≉ᚩࢆ᥈ࡿࡇ࡜ࡣ㞴ࡋ࠸ࠋ⚄ ᡞ ᕷ ࡢ භ ⏥ ᒣ ⣔ ༡ ഃ ᩳ 㠃 ࡢ ᕷ ⾤ ᆅ ࡛ ࡣ ࠊ ࢽ ࣍ ࣥ ࢖ ࣀ ࢩ ࢩ 㸦 Sus scrofa

leucomystax㸹௨ୗ࢖ࣀࢩࢩ㸧࡟ࡼࡿ⿕ᐖࡀከⓎࡋ࡚࠸ࡿࠋ⿕ᐖⓎ⏕ࡢ᰿ᮏⓗ࡞ཎᅉࡣࠊ
࢖ࣀࢩࢩ࡬ࡢ㣵௜ࡅ⾜Ⅽ࡛࠶ࡿ㸦⚄ᡞᕷ 2002㸹㎷࣭ᶓᒣ 2014㸧ࠋ㣵௜ࡅ࡟ࡼࡾே័ࢀ
ࡋࡓ࢖ࣀࢩࢩࡣ⧞ࡾ㏉ࡋᕷ⾤ᆅ࡟ฟἐࡋࠊࢦ࣑ࢆⲨࡽࡋࡓࡾࠊேࢆくࡗࡓࡾࡍࡿࠋ࢖ࣀ
ࢩࢩࡣ኱ኚᏛ⩦⬟ຊࡀ㧗࠸ື≀࡛࠶ࡾ㸦Ụཱྀ 2003㸧ࠊࠕே࠿ࡽ㣵ࢆ⡆༢࡟ᚓࡿࡇ࡜ࡀ࡛
රᗜ┴᳃ᯘື≀◊✲ࢭࣥࢱ࣮࣭ රᗜ┴❧኱Ꮫ⮬↛࣭⎔ቃ⛉Ꮫ◊✲ᡤ 

රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-11ࡁࡿࠖࠊࠕᕷ⾤ᆅ࡟ฟἐࡍࢀࡤᰤ㣴౯ࡢ㧗࠸㣗≀ࢆᚓࡿࡇ࡜ࡀ࡛ࡁࡿࠖࠊࠕேࢆく࠺ࡇ࡜࡛
㣵ࢆᚓࡿࡇ࡜ࡀ࡛ࡁࡿࠖ࡜ẁ㝵ⓗ࡟Ꮫ⩦ࡋࠊ⾜ືࢆ࢚ࢫ࣮࢝ࣞࢺࡉࡏࡿ㸦㎷࣭ᶓᒣ 2014㸧ࠋ୍᪦ࠊேᕤ≀ࡸᕷ⾤ᆅ࡟ᇳ╔ࡋ࡚ࡋࡲࡗࡓಶయࡢ⾜ືࢆᢚไࡍࡿࡇ࡜ࡣࡁࢃࡵ࡚㞴ࡋ࠸ࠋ௨ୖࡢ࢖ࣀࢩࢩࡢ≉ᛶ࠿ࡽࠊ⿕ᐖࢆ࡞ࡃࡍᑐ⟇࡜ࡋ࡚ࡣࠊ᭷ᐖᤕ⋓ࡉࢀࡓၥ㢟ಶయ࠾ࡼ
ࡧ஺㏻஦ᨾࡸิ㌴஦ᨾಶయࡣẅฎศࡍࡿ࡜࠸࠺㑅ᢥ⫥ࡋ࠿ṧࡽ࡞࠸ࠋࡋ࠿ࡋࠊࡇࡢࡼ࠺
࡞⌮⏤࡛ẅฎศࡉࢀࡓಶయࡢ≉ᚩࡣࡇࢀࡲ࡛୙࡛᫂࠶ࡗࡓࠋ
ࡑࡇ࡛ᮏ◊✲࡛ࡣࠊ࡝ࡢࡼ࠺࡞ಶయࡀᏛ⩦ࡋၥ㢟ࢆ㉳ࡇࡋ࡚࠸ࡿࡢ࠿ࢆᢕᥱࡍࡿࡓࡵ
࡟ࠊ≉ᐃࡢၥ㢟ಶయ࡜ࡋ࡚᭷ᐖᤕ⋓ࡉࢀࡓಶయ࡜ᕷ⾤ᆅ࡟࡚஺㏻஦ᨾ➼࡛Ṛஸࡋࡓಶయ
ࢆᅇ཰ࡋศᯒࢆ⾜ࡗࡓࠋ㸬ᮦᩱ࡜᪉ἲ
ᮏ◊✲ࡣࠊ࢖ࣀࢩࢩ⥭ᛴᑐ⟇༠ຊဨ㸦㎷࣭ᶓᒣ 2014㸧ࡀ⥭ᛴᑐᛂࡋ࡚ᤕ⋓ࡋࡓಶయࢆᑐ㇟࡜ࡋࡓࠋ஦๓࡟⥭ᛴᑐ⟇༠ຊဨ࡟༠ຊ౫㢗ࡋࠊヱᙜࡍࡿᑐᛂࡀ࠶ࡗࡓሙྜࡢ㐃⤡ㄪᩚࢆ⾜ࡗࡓࠋ2009 ᖺ࠿ࡽ 2011 ᖺ࡟⚄ᡞᕷ࡟࡚ࠊၥ㢟ಶయ࡜ࡋ࡚᭷ᐖᤕ⋓ࡉࢀࡓಶయ
2 㢌ࠊ஺㏻஦ᨾ࣭ิ㌴஦ᨾಶయ 5 㢌ࠊィ 7 㢌ࢆ⏝࠸ࡓࠋࡇࢀࡽࡣ඲࡚ࠊಶయࡢẅฎศࡀ᏶஢ࡋࡓ᫬Ⅼ࡛እ㒊ィࢆ⾜࠸ࠊෆ⮚࡜ୗ㢡ࢆྲྀࡾฟࡋࡓࠋෆ⮚࠿ࡽࡣࠊ⫶ෆᐜ≀ࠊ⏕Ṫჾࢆ᥇ྲྀࡋࡓࠋ⫶ෆᐜ≀ࡣ⫗║ほᐹ࡟࡚☜ㄆࡋࠊ⏕Ṫჾࡢ⫗║࠾ࡼࡧ⤌⧊ほᐹ࡟࡚ᛶᡂ⇍ࡸዷፎࡢ᭷↓ࢆุ᩿ࡋࡓࠋᖺ㱋ࡣࠊୗ㢡ࡢṑࡢⴌฟ࡜஺᥮࠾ࡼࡧ➨୍ᚋ⮻ṑࡢࢭ࣓
ࣥࢺ㉁ᒙࡢᖺ㍯ᩘࡢ࢝࢘ࣥࢺ࡟࡚Ỵᐃࡋࡓ㸦Matschcke 1967; ᯘ࡯࠿ 1977; Boitani &Mattei 1992㸧ࠋ ࡲࡓࠊᮏ◊✲ࡢᑐ㇟ಶయ࡜௚ᆅᇦ࡛ᤕ⋓ࡉࢀࡓಶయࡢయࢧ࢖ࢬࠊᰤ㣴≧ែࡢ㐪࠸ࡢ᭷↓ࢆ᳨ウࡍࡿࡓࡵ࡟ࠊ2003 ᖺ࠿ࡽ 2012 ᖺ࡟රᗜ┴ᮏᕞ㒊࡛⊁⊟࠾ࡼࡧ᭷ᐖᤕ⋓࡟࡚ᤕ⋓ࡉࢀࡓಶయ 195 㢌ࡢయ㔜ィ್ࢆ⏝࠸ࡓࠋ

㸬⤖ᯝ࡜⪃ᐹ
1R
 No.1 ࡣࠊิ㌴஦ᨾ࡟㐼ࡗ࡚ᶓࡓࢃࡗ࡚࠸ࡿ࡜ࡇࢁࢆ῝ኪ࡟Ⓨぢࡉࢀࠊ㥑ࡅࡘࡅࡓ⥭ᛴᑐ⟇༠ຊဨ࡟ẅฎศࡉࢀࡓಶయ࡛࠶ࡿࠋNo.1 ࡣ 2 ṓࡢᡂ⋇ࡢ࢜ࢫ࡛࠶ࡗࡓ㸦⾲㸯㸧ࠋ⫶ෆᐜ࡟ࢧࣛࣥࣛࢵࣉࡸ⣬ࠊ࢝ࢵࢺࡉࢀࡓࢩ࢖ࢱࢣࡸࣈࢻ࢘ࡢ⓶࡞࡝ࡀྵࡲࢀ࡚࠸ࡓࡇ
࡜࠿ࡽ㸦ᅗ㸯㸧
ࠊࢦ࣑ࢆ࠶ࡉࡾ࡟ᕷ⾤ᆅ࡟ฟἐࡋࠊิ㌴஦ᨾ࡟㐼ࡗࡓ࡜᥎ᐹࡉࢀࡓࠋ 

රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-11

⾲ 1R ࡢศᯒ⤖ᯝᅇ཰᪥

Ṛஸཎᅉ

ิ㌴஦ᨾ

ᛶู

࢜ࢫ

ᖺ㱋

 ṓ

య㔜

NJ

య㛗

FP

⫶ෆᐜ≀ ேᕤ≀࠶ࡾ
ᛶᡂ⇍

࠶ࡾ
ᅗ  1R ࠿ࡽ᳨ฟࡉࢀࡓ⫶ෆᐜ≀

1R
No.2 ࡣࠊභ⏥ᒣᒣ㯄࡛Ⓩᒣ⪅ࢆく࠺⿕ᐖࢆከⓎࡉࡏࡓၥ㢟ಶయ࡛࠶ࡿࠋ▷ᮇ㛫࡛⿕ᐖࡀࡁࢃࡵ࡚῝้࡟࡞ࡗࡓࡇ࡜࠿ࡽࠊ᭷ᐖᤕ⋓ࡍࡿᚲせᛶࡀ࠶ࡿ࡜ุ᩿ࡉࢀࠊᤕ⋓ࡀᐇ⾜ࡉࢀࡓࠋNo.2 ࡣࠊ4 ṓࡢᡂ⋇࢜ࢫ࡛࠶ࡗࡓ㸦⾲㸰㸧ࠋ⫶ෆᐜ࠿ࡽࡣ⡿⢏ࠊ࠾ⳫᏊࡢໟࡳ⿄ࠊࢧࣛࣥࣛࢵࣉ࡞࡝኱㔞ࡢேᕤ≀ࡀ☜ㄆࡉࢀ㸦ᅗ㸰㸧ࠊேⅭⓗ࡞㣗࡭≀࡟ᙉࡃ౫Ꮡ
ࡋ࡚࠸ࡓࡇ࡜ࡀ᥎ᐹࡉࢀࡓࠋ
⾲ 1R ࡢศᯒ⤖ᯝᅇ཰᪥

Ṛஸཎᅉ

᭷ᐖᤕ⋓

ᛶู

࢜ࢫ

ᖺ㱋

 ṓ

య㔜

NJయ㛗FP⫶ෆᐜ≀ ேᕤ≀࠶ࡾ
ᛶᡂ⇍

࠶ࡾ

ᅗ  1R ࠿ࡽ᳨ฟࡉࢀࡓ⫶ෆᐜ≀

No.2 ࡀẅฎศࡉࢀࡓ 5 ᪥ᚋ࡟᳃ᯘື≀◊✲ࢭࣥࢱ࣮ࡀ⾜ࡗࡓᤕ⋓ᚋㄪᰝ࡟࡚ࠊᤕ⋓⌧ሙ࡟⊩ⰼࢆⓎぢࡋࡓ㸦ᅗ㸱㸧ࠋ⊩ⰼ࡟ῧ࠼
ࡽࢀࡓ࣓ࢵࢭ࣮ࢪ࡟ࡣࠊಶయࡢ⏕ᖺ᭶ࠊṚஸ᪥ࡀ᭩࠿ࢀ࡚࠾ࡾࠊ⏕ࡲࢀᖺࡣศᯒ⤖ᯝ࡜୍⮴ࡋࡓࠋࡇࡢࡇ࡜࠿ࡽࠊNo.2 ࡀ㛗ᖺⓏᒣ⪅࡟ほᐹࡉࢀ࡚࠸ࡓே័ࢀಶయ࡛࠶ࡗࡓ
࡜ุ᩿ࡉࢀࡓࠋᅗ  1R ࡢᤕ⋓⌧ሙ࡟࠶ࡗࡓ⊩ⰼ 

රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-11

1R2010 ᖺ࠿ࡽ 2011 ᖺ࡟≉ᐃࡢᆅᇦ࡟࠾࠸࡚ࠊ࢖ࣀࢩࢩ」ᩘ㢌ࡀ㐃⥆ⓗ࡟ே㌟⿕ᐖࢆ㉳ࡇࡋࡓࠋၥ㢟ಶయࢢ࣮ࣝࣉࡣࠊ୍㒊ࡢఫẸࡢ㣵௜ࡅ⾜Ⅽ࡟ࡼࡾே័ࢀࡋ࡚࠸ࡓࠋࡇࢀ
ࡽࡢ࢖ࣀࢩࢩࡢ⾜ືࡣ࢚ࢫ࣮࢝ࣞࢺࡋࠊ᭱⤊ⓗ࡟ࡣẖ᪥ࡢࡼ࠺࡟㈙࠸≀⿄ࢆᣢࡗࡓዪᛶ
ࡸ㧗㱋⪅ࢆ㑅ᢥⓗ࡟くࡗ࡚࠸ࡓࠋ⿕ᐖࡢᣑ኱࡜῝้໬࠿ࡽࠊၥ㢟ಶయࢢ࣮ࣝࣉࡢᤕ⋓ࡀ㡰ḟ⾜ࢃࢀࠊNo.3 ࡀ᭱ᚋࡢ 1 㢌࡛࠶ࡗࡓࠋNo.3 ࡢᤕ⋓ᚋࠊ⿕ᐖࡣ཰᮰ࡋࡓࠋNo.3 ࡣࠊ
5 ṓࡢᡂ⋇࣓ࢫ࡛࠶ࡾࠊయ㔜ࡀ 100kg ࡛࠶ࡗࡓ㸦⾲㸱㸧
ࠋࡲࡓ 6 㢌ࡢ⫾Ꮚࢆዷፎࡋ࡚࠾
ࡾࠊ⫾Ꮚࡢᡂ㛗ẁ㝵࠿ࡽฟ⏘┤๓࡛࠶ࡗࡓ࡜♧၀ࡉࢀࡓ㸦ᅗ㸲㸧ࠋ⾲ 1R ࡢศᯒ⤖ᯝ㻌ᅇ཰᪥

Ṛஸཎᅉ

᭷ᐖᤕ⋓

ᛶู

࣓ࢫ

ᖺ㱋

 ṓ

య㔜

NJ

య㛗FP⫶ෆᐜ≀ ேᕤ≀࠶ࡾ
⦾Ṫ≧ἣ

ዷፎ࠶ࡾ

ᅗ  1R ࠿ࡽ☜ㄆࡉࢀࡓ⫾Ꮚ

1R㹼
 No.4㹼6 ࡣࠊ஺㏻஦ᨾ࡛Ṛஸࡋࡓಶయ࡛࠶ࡿࠋNo.7 ࡶྠᵝ࡟஺㏻஦ᨾ࡛࠶ࡗࡓࡀࠊ
஦ᨾᚋࡶࡲࡔ⏕ࡁ࡚࠸ࡓࡓࡵࠊ㆙ᐹ࡟ᅄ⫥ࢆ⦖࡛ಖᐃࡉࢀࠊ࢖ࣀࢩࢩᑐ⟇༠ຊဨ࡟ࡼࡾ
ࡑࡢሙ࡛Ᏻᴦẅࡉࢀࡓ㸦ᅗ㸳㸧
ࠋ஦ᨾⓎ⏕᫬ࡣࠊ༗๓ 8 ᫬๓ᚋ࡛࠶ࡗࡓࡓࡵࠊ㏻໅࣭㏻
Ꮫࡢேࡢ㏻ࡾࡸ஺㏻㔞ࡀከ࠿ࡗࡓࠋ㎿㏿࠿ࡘ㐺ษ࡞ᑐᛂࡀ࡞ࡉࢀ࡞ࡅࢀࡤࠊಶయࡀᭀࢀ
ࡓࡾࠊ஺㏻࡟ΰ஘ࡀ⏕ࡌࡓࡾ࡜஧ḟ⿕ᐖࡀⓎ⏕ࡍࡿ༴㝤ᛶࡢ㧗࠸஦ᨾ࡛࠶ࡗࡓࠋ
No.4㹼6 ࡢ஦ᨾࡀⓎ⏕ࡋࡓ 4 ᪥ᚋ࡟ No.7 ࡢ஦ᨾⓎ⏕ࡀⓎ⏕ࡋࡓࠋࡲࡓ஦ᨾ⌧ሙࡣࠊྠࡌ㐨㊰ୖࡢ㞄᥋ࡍࡿ஺ᕪⅬ௜㏆࡛࠶ࡾࠊ⣙ 60㹼70㹫ࡢ⠊ᅖ࡛࠶ࡗࡓࠋ
 No.4~6 ࡣ 0 ṓࡢ࣓ࢫࠊNo.7 ࡣ 10 ṓࡢ㧗㱋ᡂ⋇࣓ࢫ࡛࠶ࡗࡓ㸦⾲㸲㸧ࠋゎ๗ࡢ⤖ᯝࠊ
No.4 ࡣ⭜᳝ᦆയࠊNo.5 ࡣ⭜᳝࣭㦵┙࣭ෆ⮚ᦆയࠊᕥ๓⫥࣭ᚋ⫥㦵ᢡࠊNo.6 ࡣ඲㌟ᡴ᧞ࠊ
ෆ⮚◚⿣ࠊNo.㸵ࡣ⭜᳝ࠊ㦵┙ᦆയ࡛࠶ࡗࡓࠋ
࡝ࡢಶయࡶᦆയࡀࡦ࡝ࡃࠊ㌴࡜ᙉࡃ⾪✺ࡋࡓ࡜⪃࠼ࡽࢀࡓࠋNo.5 ࡜ 6 ࡣෆ⮚ࡢᦆയࡢ⛬ᗘ࠿ࡽ༶ṚࠊNo.4 ࡜ 7 ࡶṚ࡟┤⤖ࡍࡿ኱᛹ᡃ࡛࠶ࡗࡓ࡜᥎ᐹࡉࢀࡓࠋNo.4㹼6 ࡢ⫶ෆᐜ≀ࡼࡾࠊᘚᙜࡢໟࡳ⣬ࠊࢧࣛࣥࣛࢵࣉ࡞࡝ࡢ ᅗ  1R ࡢ஺㏻஦ᨾ⌧ሙරᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-11

ேᕤ≀ࡀ᳨ฟࡉࢀࡇ࡜࠿ࡽ㸦ᅗ㸴㸧ࠊࡇࡢ 3 㢌ࡣࠊࢦ࣑ࢆ࠶ࡉࡾ࡟ᕷ⾤ᆅ࡟ฟἐࡋࡓ㝿
࡟஺㏻஦ᨾ࡟㐼ࡗࡓ࡜⪃࠼ࡽࢀࡓࠋࡲࡓ No.6 ࡢྑ๓⫥࡟ࡣࠊࣉࣛࢫࢳࢵࢡࡢ㍯ࡀ⫥࡟㣗࠸㎸ࡳࠊ㋟ࡀኚᙧࡋ࡚࠸ࡓ㸦ᅗ㸵㸧ࠋᗂ࠸᫬ᮇ࡟ㄗࡗ࡚㍯ࡀࡣࡲࡗ࡚ࡋࡲ࠸ࠊࡑࡢࡲ
ࡲᡂ㛗ࡋࡓࡓࡵ࡟㋟ࡀኚᙧࡋࡓ࡜᥎

ࡉࢀࡓࠋࡇࡢࡼ࠺࡞≧ἣ࠿ࡽࠊNo.4㹼6 ࡣᗂ࠸᫬ᮇ࠿ࡽᕷ⾤ᆅࢆ⾜ືᅪ࡜ࡋ࡚࠸ࡓࡇ࡜ࡀ⪃࠼ࡽࢀࡓࠋ⾲  1R㹼 ࡢศᯒ⤖ᯝ
1R

1R

1R

1R

ᅇ཰᪥









Ṛஸཎᅉ

஺㏻஦ᨾ

஺㏻஦ᨾ

஺㏻஦ᨾ

஺㏻஦ᨾ

ᛶู

࣓ࢫ

࣓ࢫ

࣓ࢫ

࣓ࢫ

ᖺ㱋

 ṓ

 ṓ

 ṓ

 ṓ

య㔜

NJNJ

NJ

NJ

య㛗

FP

FP

FP

FP

⫶ෆᐜ≀

ேᕤ≀࠶ࡾ

ேᕤ≀࠶ࡾ

ேᕤ≀࠶ࡾ

ேᕤ≀࡞ࡋ

⦾Ṫ≧ἣ

ᮍᡂ⇍

ᮍᡂ⇍

ᮍᡂ⇍

ዷፎṔ࠶ࡾᅗ  1R ࠿ࡽ᳨ฟࡉࢀࡓ⫶ෆᐜ≀ᅗ  ㍯ࡀ㣗࠸㎸ࢇࡔ 1R ࡢ⫥

No.4㹼7 ࡢ஺㏻஦ᨾࡣࠊྠ୍㐨㊰ୖࡢ㏆᥋ࡋࡓሙᡤ࡛㐃⥆ⓗ࡟㉳ࡇࡗࡓࡇ࡜ࠊಶయࡢ
ෆヂࡀᡂ⋇࣓ࢫ 1 㢌ࠊ0 ṓ 3 㢌࡛࠶ࡗࡓࡇ࡜ࠊࡇࡢ᫬ᮇࡣᡂ⋇࣓ࢫ࡜ 0 ṓࡢᏊࡀ⾜ືࢆඹ࡟ࡍࡿ᫬ᮇ࡛࠶ࡿࡇ࡜࠿ࡽࠊࡇࢀࡽࡢಶయࡣぶᏊ࡛࠶ࡗࡓྍ⬟ᛶࡀ࠶ࡿࠋࡋࡓࡀࡗ࡚ࠊ
ࡇࡢ୍㐃ࡢ஺㏻஦ᨾࡣࠊẕ࢖ࣀࢩࢩ࠿ࡽᏊ࡬࡜ᕷ⾤ᆅࡀ⾜ືᅪ࡛࠶ࡿࡇ࡜ࠊ㣗࡭≀ࡀࢦ࣑࡞࡝ࡢேᕤ≀࡛࠶ࡿࡇ࡜ࡀఏ࠼ࡽࢀ࡚࠸ࡿࡇ࡜ࢆ♧ࡍ஦౛࡛࠶ࡗࡓ࡜⪃࠼ࡿࠋ௚ᆅᇦಶయ࡜ࡢẚ㍑
 No.1 ࡜ No.2 ࡢయ㔜ࢆරᗜ┴ෆࡢ௚ᆅᇦ࡟࡚ᤕ⋓ࡉࢀࡓᡂ⋇࢜ࢫ㸦N=64㸧ࡢయ㔜࡜ẚ㍑ࡋࡓࠋࡑࡢ⤖ᯝࠊ୧⪅࡜ࡶ 9 ᭶ࡢᖹᆒయ㔜ࡼࡾࢃࡎ࠿࡟㔜࠿ࡗࡓࡀࠊ኱ࡁࡃࡣኚࢃ 

රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-11

ࡽ࡞࠿ࡗࡓ㸦ᅗ㸶㸧ࠋḟ࡟ No.3 ࡜ No.7 ࡢయ㔜ࢆᡂ⋇࣓ࢫ㸦N=102㸧ࡢయ㔜࡜ẚ㍑ࡋࡓ
࡜ࡇࢁࠊ࡝ࡕࡽࡶᖹᆒ್ࢆୖᅇࡗࡓࠋ࡜ࡃ࡟ No.3 ࡣᴟࡵ࡚㔜࠿ࡗࡓ㸦ᅗ㸷㸧ࠋ᭱ᚋ࡟
No.4㹼6 ࡢయ㔜࡜ 0 ṓಶయ㸦N=129㸧ࡢయ㔜࡜ࢆẚ㍑ࡋࡓࠋࡑࡢ⤖ᯝࠊNo.4 ࡣᖹᆒࡼࡾࡸࡸ㔜ࡃࠊNo.5 ᖹᆒ್࡜ྠ⛬ᗘࠊNo.6 ࡣᖹᆒࡼࡾᑡࡉࡃ࡞ࡗࡓ㸦ᅗ㸯㸮㸧ࠋ ௒ᅇࡢ⤖ᯝ࠿ࡽࠊNo.3 ࡸ No.7 ࡣ㣵ࡸࢦ࣑࡞࡝ࡢேᕤ≀࡟౫Ꮡࡋࡓ⤖ᯝࠊయ㔜ࡀ㔜ࡃ࡞ࡾࠊᰤ㣴≧ែࡶྥୖࡋ࡚࠸ࡓࡇ࡜ࡀ♧၀ࡉࢀࡓࠋ௚ࡢಶయ࡟ࡘ࠸࡚ࡣࠊ㣵ࡸேᕤ≀࡬ࡢᙳ㡪ࡀయࢧ࢖ࢬࡸᰤ㣴≧ែ࡟ࡲ࡛࡟ࡣ཯ᫎࡉࢀ࡚࠸࡞࠸࡜ุ᩿ࡉࢀࡓࠋࡋࡓࡀࡗ࡚ࠊ
ᕷ⾤ᆅ࡟ฟἐࡋࡓࡾࠊேࢆくࡗࡓࡾࡍࡿၥ㢟ಶయ࡛ࡶࠊேᕤ≀ࢆᚓࡿ⛬ᗘ࡟ࡼࡗ࡚ࠊ⏕⌮ⓗ࡞ᙳ㡪࡟ࡣࡤࡽࡘࡁࡀ⏕ࡌࡿࡇ࡜ࡀ⪃࠼ࡽࢀࡓࠋᅗ  ᡂ⋇࢜ࢫࡢᏘ⠇ูయ㔜᥎⛣ᅗ  ᡂ⋇࣓ࢫࡢᏘ⠇ูయ㔜᥎⛣ 

රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-11

ᅗ   ṓࡢᏘ⠇ูయ㔜᥎⛣㸬ࡲ࡜ࡵ ᮏ◊✲ࡣᶆᮏᩘࡀᑡ࡞࠿ࡗࡓࡀࠊ⌧᫬Ⅼ࡛♧၀ࡉࢀࡓၥ㢟ಶయࠊ஦ᨾಶయࡢ≉ᚩࢆ௨ୗ࡟ᣲࡆࡿࠋ
ձேᕤ≀࡟ᇳ╔ᛶࢆᣢࡘࠋ
ศᯒಶయ 7 㢌୰ 6 㢌ࡢ⫶ෆᐜ࠿ࡽேᕤ≀ࡀ☜ㄆࡉࢀࡓࠋேᕤ≀ࢆ㣵࡜ㄆ㆑ࡋᇳ╔ࡋࡓಶయࡀࠊᕷ⾤ᆅࡢฟἐࡸேࢆく࠺⾜ືࢆ㉳ࡇࡍࡇ࡜ࡀࠊ௒ᅇࡢ⤖ᯝ࡛⿬௜ࡅࡽࢀࡓࠋ
ղၥ㢟ಶయࡸ஦ᨾಶయ࡬࡜Ⓨᒎࡍࡿྍ⬟ᛶ࡟ࡣᛶᕪࡸᖺ㱋ᕪࡣ࡞ࡃࠊ㣵௜ࡅࡉࢀࡓಶయࠊ
ࢦ࣑࡟ᇳ╔ࡋࡓಶయࡀၥ㢟ಶయ࡜࡞ࡿࠋ  ศᯒࡋࡓᡂ⋇ಶయ࡟ࡣࠊ㞤㞝ࡢ೫ࡾࡀㄆࡵࡽࢀ࡞࠿ࡗࡓࠋ0 ṓಶయࡣ࠸ࡎࢀࡶ࣓ࢫ
࡛࠶ࡗࡓࡀࠊ0 ṓࡣᇶᮏⓗ࡟⾜ືࢆඹ࡟ࡍࡿࡓࡵࠊฟἐ᫬ࡢᛶᕪ࡛ࡣ࡞ࡃ⏕ࡲࢀࡓ㝿ࡢᛶᕪࢆ཯ᫎࡋ࡚࠸ࡿ࡜⪃࠼ࡽࢀࡓࠋࡋࡓࡀࡗ࡚⌧ẁ㝵࡛ࡣࠊၥ㢟ಶయ࡟ᛶᕪࡣ࡞࠸
࡜⪃࠼ࡿࠋࡲࡓศᯒಶయࡣ 0㹼10 ṓ࡟ࢃࡓࡾᖺ㱋ࡢ೫ࡾࡣ☜ㄆࡉࢀ࡞࠿ࡗࡓࠋ⌧᫬
Ⅼ࡛ࡣࠊၥ㢟ಶయ࡟࡞ࡾࡸࡍ࠸ᖺ㱋ࡀ࠶ࡿࡢ࡛ࡣ࡞ࡃࠊ㣵௜ࡅࡉࢀࡓࡾࠊࢦ࣑࡟ᇳ╔ࡋࡓࡾࡍࡿࢱ࢖࣑ࣥࢢ࡟౫Ꮡࡍࡿ࡜⪃࠼ࡿࠋ
ճே㌟⿕ᐖ࡞࡝ᝏ㉁࡞⿕ᐖࢆ㉳ࡇࡍಶయࡣࠊ㛗ᮇࡢ㣵௜ࡅ⾜Ⅽ࡟ࡼࡾே័ࢀࡋ࡚ᡂ㛗ࡋࡓಶయ࡛࠶ࡿࠋ
 ே㌟⿕ᐖࢆከⓎࡉࡏࡓ࡜ࡋ࡚᭷ᐖᤕ⋓ࡉࢀࡓ No.2 ࡜ No.3 ࡣࠊ࡝ࡕࡽࡶ㛗ᮇ㛫㣵௜ࡅࡉࢀࡓᡂ⋇ಶయ࡛࠶ࡗࡓࠋࡋࡓࡀࡗ࡚ࠊᏊ࡝ࡶࡢ㡭࠿ࡽ㣵௜ࡅࡉࢀ࡚࠸ࡓಶయࡀࠊ
ᡂ⋇࡟࡞ࡿ࡟ࡘࢀ࡚Ꮫ⩦ࡀ㐍ࢇ࡛ࠊ⾜ືࡀ࢚ࢫ࣮࢝ࣞࢺࡋࠊே㌟⿕ᐖࢆ㉳ࡇࡍၥ㢟ಶయ࡟࡞ࡿࡇ࡜ࡀ⪃࠼ࡽࢀࡿࠋմぶࡀᕷ⾤ᆅ࡟ᇳ╔ࡋ࡚࠸ࡓሙྜࠊᏊࡶᕷ⾤ᆅ࡟ᇳ╔ࡍࡿࠋ  0 ṓಶయࡣࠊᇶᮏⓗ࡟ẕ࢖ࣀࢩࢩ࡜⾜ືࢆඹ࡟ࡍࡿࠋࡋࡓࡀࡗ࡚ No.4~6 ࡣࠊぶ࡟㐃ࢀࡽࢀ࡚ᕷ⾤ᆅ࡟ฟἐࡋࡓ࡜᥎

ࡉࢀࡿࠋၥ㢟ಶయࡀẕ࢖ࣀࢩࢩࡢሙྜࠊᏊࡶྠᵝ࡟ၥ㢟ಶయࠊᕷ⾤ᆅฟἐಶయ࡟࡞ࡿྍ⬟ᛶࡀ㧗࠸ࡇ࡜ࡀ♧၀ࡉࢀࡓࠋ 

රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-11ㅰ㎡ศᯒಶయࡢᅇ཰࡟࠶ࡓࡾࠊከ኱࡞ࡿࡈ༠ຊࢆ㡬ࡁࡲࡋࡓ⚄ᡞᕷ⏘ᴗ᣺⯆ᒁ㎰ᨻ㒊ィ⏬
ㄢࠊྛ༊ࡢࡲࡕ࡙ࡃࡾㄢ࠾ࡼࡧࡲࡕ࡙ࡃࡾ᥎㐍ㄢࠊ⚄ᡞ㎰ᯘỈ⏘᣺⯆஦ົᡤࠊ࢖ࣀࢩࢩ
⥭ᛴᑐ⟇༠ຊဨࡢⓙࡉࡲ࡟῝ࡃᚚ♩⏦ࡋୖࡆࡲࡍࠋᮏ◊✲ࡣࠊᖹᡂ 25㹼27 ᖺᗘ JSPS⛉◊㈝㸦25450478㸧
ࠕ㒔ᕷ⎔ቃ࡟౵ධࡍࡿ࢖ࣀࢩࢩࡢࣜࢫࢡศᯒ࡜ࣜࢫࢡᅇ㑊ᡭἲࡢ᳨ウࠖࡢຓᡂࢆཷࡅࡲࡋࡓࠋ

ᘬ⏝ᩥ⊩
Boitani, L. and Mattei, L. 1992. Aging wild boar (Sus scrofa) by tooth eruption. In: Spitz,
F., Janeau, G., Gonzales, G. and Aulagnier, S. editors. pp.419-421. Ongules/
Ungulates 91, Toulouse-Paris.
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ᮐᖠᕷ㸬2010㸬ᖹᡂ 22 ᖺᗘᮐᖠᕷ⥭ᛴ㞠⏝๰ฟ᥎㐍஦ᴗ㔝⏕ື≀࡟ࡼࡿᕷ⾤ᆅ➼࡬ࡢ౵ධ⤒ ㊰ ㄪ ᰝ ࠾ ࡼ ࡧ ౵ ධ 㜵 Ṇ ⟇ ࡢ ㄪ ᰝ ࣭ ◊ ✲ ᴗ ົ ሗ ࿌ ᭩ 㸬 ᮐ ᖠ ᕷ 㸪 http://www.city.
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㎷▱㤶࣭ᶓᒣ┿ᘪ㸬2014㸬භ⏥ᒣ࢖ࣀࢩࢩၥ㢟ࡢ⌧≧࡜ㄢ㢟㸬ࠕරᗜ┴࡟࠾ࡅࡿࢽ࣍ࣥ࢖ࣀ
ࢩࢩࡢ⟶⌮ࡢ⌧≧࡜ㄢ㢟㸬ࠖ㸪࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6 ྕ㸪pp.121-134㸪රᗜ┴᳃ᯘື≀◊✲ࢭࣥࢱ࣮㸬 

࠾ࢃࡾ࡟
රᗜ┴Ẹ࡟࡜ࡗ࡚ࠊ࢖ࣀࢩࢩࡣẚ㍑ⓗ㌟㏆࡟⏕ᜥࡋ࡚࠸ࡿ㔝⏕ື≀࡛ࡍࠋ㌟㏆࡟࠸ࡿ࠿ࡽࡇࡑࠊ㎰ᴗ⿕ᐖ࡟ࡶᖖ࡟ᝎࡲࡉࢀ࡚ࡶࡁࡲࡋࡓࡋࠊ㈨※࡜ࡋ࡚ࡢ౯್ࡶ㧗ࡃㄆࡵ࡚ࡁࡲࡋࡓࠋ㒔ᕷ࡜භ⏥ᒣ࡜࠸࠺࠸ࢃࡤே࡜㔝⏕ື≀ࡢධ఍ᆅ࡛ࡢᵝࠎ࡞ฟ᮶஦ࡸၥ㢟ࡶ
Ⓨ⏕ࡋ࡚࠸ࡲࡍࠋⰋࡃࡶᝏࡃࡶྂࡃ࠿ࡽࠊᵝࠎ࡞ࡸࡾྲྀࡾࢆ㔜ࡡ࡚ࡁࡓ㥆ᰁࡳࡢ࠶ࡿ㔝⏕ື≀࡜࠸࠼ࡲࡍࠋ࡜ࡇࢁࡀࠊ⛉Ꮫⓗ࡛ィ⏬ⓗ࡞⟶⌮ࢆ⾜࠾࠺࡜ࡍࡿ࡜ࠊࡑࢀࡽ࡟ᚲせ࡞᝟ሗࡣࠊ௚ࡢ㔝⏕ື≀࡟ẚ࡭ࠊᴟ➃࡟ᑡ࡞࠸≧ἣ࡟࠶ࡾࡲࡋࡓࠋ㌟㏆ࡍࡂ࡚ࠊ࠶࠼࡚
ㄪ࡭࡞ࡃ࡚ࡶࠊ࠾࠾ࡼࡑࡢ≧ἣࡣศ࠿ࡗ࡚࠸ࡿື≀ࠊ࡜࠸࠺ࡇ࡜ࡔࡗࡓࡢ࡛ࡋࡻ࠺ࠋ
ᐇ㝿࡟ㄪᰝࢆጞࡵࡿ࡜ࠊࡸࡣࡾࠊ᝟ሗࡀᚓࡽࢀ࡞࠸ࡇ࡜ࡀከࡃࠊࢹ࣮ࢱ㞟ࡵ࡟ⱞປࡍ
ࡿࢃࡾ࡟ࡣࠊ㛗࠸㛫ࡘ࠿ࡳ࡝ࡇࢁࡢ࡞࠸୙ᛮ㆟࡞┦ᡭ࡛ࡋࡓࠋᑐ⟇ࡶฟ᮶ࡿࡇ࡜ࡣ㝈ࡽࢀࠊࡍ࡛࡟⾜ࡗ࡚࠸ࡿࡇ࡜ࡶከ࠿ࡗࡓࢃࡅ࡛ࡍࡀࠊ㎰ᴗ⿕ᐖࡸே㌟஦ᨾࡣῶࡿẼ㓄ࢆぢࡏࡲࡏࢇࠋࡑࢀ࡝ࡇࢁ࠿ቑຍഴྥ࡟㌿ࡌጞࡵ࡚ࡁࡲࡋࡓࠋே㛫ഃࡢᑐ⟇ࢆ㏫ᡭ࡟࡜ࡗ࡚ࠊ
࢖ࣀࢩࢩࡣࠊ╔ᐇ࡟ໃຊࢆᣑ኱ࡋ࡚࠸ࡿࡼ࠺࡟ࡉ࠼ឤࡌࡿ≧ἣ࡜࡞ࡾࡲࡋࡓࠋ
ࡇ࠺ࡋࡓヨ⾜㘒ㄗࡢ୰ࠊ᝟ሗࡣ㐜ࠎ࡜ࡋ࡚㞟ࡲࡾࡲࡏࢇ࡛ࡋࡓࡀࠊ㔝⏕ື≀ࡢ㐺ṇ࡞⟶⌮࡟ྥࡅࡓྲྀࡾ⤌ࡳࢆ⌮ゎࡋࠊ᝟ሗ཰㞟࡟ࡈ༠ຊࢆ࠸ࡓࡔࡅࡿᑐ⟇⪅ࠊ㛵ಀ⪅ࡢ᪉ࠎࡀቑ࠼࡚ࡁࡲࡋࡓࠋ࢖ࣀࢩࢩࡢ⏕ែࡸ⿕ᐖ㜵Ṇᑐ⟇࡟㛵ᚰࢆᐤࡏ࡚ࡃࡔࡉࡿேࡀቑ࠼ࡿ
࡜ࠊḟ➨࡟ࡑࡢ᝟ሗࢆ᳃ᯘື≀◊✲ࢭࣥࢱ࣮࡟㞟⣙ࡋ࡚ࡃࡔࡉࡿࡼ࠺࡟࡞ࡾࠊ⭾኱࡞ᤕ⋓ࡸ⿕ᐖ࡟㛵ࡍࡿ᝟ሗࡀ㞟ࡵࡽࢀࠊ⤫ィⓗ࡞ゎᯒࢆ⾜࠺ࡇ࡜ࡀ࡛ࡁࡿࡼ࠺࡟࡞ࡾࡲࡋࡓࠋ
ࡲࡓࠊ඲ᅜⓗ࡟ࡶ㞟ࡵ࡟ࡃ࠿ࡗࡓ⦾Ṫࡢ᝟ሗࡸឤᰁ⑕ࡢ᝟ሗࠊ⾜ືࡢ᝟ሗ࡞࡝ࢆ⵳✚ࡍࡿࡇ࡜ࡀ࡛ࡁࡿࡼ࠺࡟࡞ࡾࡲࡋࡓࠋ
ࡇࡢࣔࣀࢢࣛࣇ࡛ࡣࠊ௒ࡢẁ㝵࡛᫂ࡽ࠿࡟࡞ࡗࡓ࢖ࣀࢩࢩࡢጼ࡜ࠊே࡜࢖ࣀࢩࢩࡢࡸ
ࡾྲྀࡾࡢጼࢆᑡࡋ࡛ࡶᙧ࡟ࡋࡓ࠸࡜࠸࠺ᛮ࠸࡛⦅㞟࠸ࡓࡋࡲࡋࡓࠋ࢖ࣀࢩࢩࡢ⿕ᐖᑐ⟇ࡸ㐺ษ࡞⟶⌮࡟ࡣ㐨ࡢࡾࡀከ࠸ࡇ࡜ࡶྠ᫬࡟⾲ࡋ࡚ࡋࡲࡗ࡚࠸ࡿ࠿ࡶࡋࢀࡲࡏࢇࠋ≧ἣࢆᢕᥱࡋ࡞ࡀࡽࠊヨ⾜㘒ㄗࢆ⧞ࡾ㏉ࡋࠊࡼࡾⰋ࠸࢖ࣀࢩࢩࡢ⟶⌮࡬࡜ྥ࠿࠺ࡓࡵࠊ୍ࡘ
ࡢࢫࢸࢵࣉ࡜ࡋ࡚ࡇࡢ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇࡢෆᐜࡀ㛵ಀ⪅ࡢⓙᵝྡࡢ୍ຓ࡟࡞
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᭱ᚋ࡟࡞ࡾࡲࡋࡓࡀࠊᆅඖࡢఫẸࡢⓙࡉࡲࠊᆅඖ⾜ᨻࡢⓙࡉࡲࢆࡣࡌࡵࠊරᗜ┴ࡢ࢖ࣀࢩࢩࡢಖㆤ⟶⌮ཬࡧ᝟ሗ⵳✚࡟ࡈ༠ຊ࣭ࡈᑾຊࢆ㡬ࡁࡲࡋࡓⓙࡉࡲ࡟ཌࡃᚚ♩⏦ࡋୖࡆࡲࡍࠋ㈐௵⦅㞟⪅ ᶓᒣ┿ᘪරᗜ┴᳃ᯘື≀◊✲ࢭࣥࢱ࣮රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6 ྕරᗜ┴࡟࠾ࡅࡿࢽ࣍ࣥ࢖ࣀࢩࢩࡢ⟶⌮ࡢ⌧≧࡜ㄢ㢟
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