රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-9➨ 㸷 ❶            ࢽ࣍ࣥ࢖ࣀࢩࢩࡢෆ㒊ᐤ⏕⹸బ⸨ ᏹ 1࣭ᡞ⏣ṇᯞ 1࣭M. A. Omar1࣭㎷ ▱㤶 2࣭ᶓᒣ┿ᘪ 2,3
せ Ⅼ࣭ රᗜ┴ୗ࡛཰㞟ࡋࡓࢽ࣍ࣥ࢖ࣀࢩࢩ 132 㢌ࢆᑐ㇟࡜ࡋ࡚ࠊࡑࡢෆ㒊ᐤ⏕⹸㸦⽸⹸㸧ࢆ⥙⨶ⓗ࡟ㄪ࡭ࡓࠋ࣭ ᑐ㇟ື≀࡟⥺⹸ 22 ✀࡜᮲⹸ 1 ✀ࠊ྾⹸ 1 ✀ࡢᐤ⏕ࡀ☜ㄆࡉࢀࡓࠋ࿴ḷᒣ┴⏘ࢽ࣍ࣥ࢖ࣀࢩࢩࢆㄪ࡭ࡓሗ࿌㸦Sato et al. 2008㸧࡛ࡣ⥺⹸ 18 ✀࡜᮲⹸ 1 ✀ࡀグ㘓ࡉࢀ࡚࠸ࡿࡀࠊ⥺⹸࡟ࡘ࠸࡚ࡣ᭷ṑ㇜⫶⹸ Ascarops dentataࠊ⣚Ⰽẟᵝ⥺⹸Hyostrongylus rubidusࠊ⨾㯇㣗㐨⹸ Gongylonema pulchrum ࡀᮏᐟ୺࠿ࡽึグ㘓࡜࡞ࡗࡓࡇ࡜ࠊࡲࡓࠊ㸯✀࡜⪃࠼࡚ࡁࡓẟ⣽⥺⹸ࡀ 3 ✀࡟෌ศ㢮ࡉࢀࡓࡇ࡜ࡀ
୧ᆅᇦ࡛ࡢᐤ⏕⹸┦ࡢ㐪࠸ࡢ኱ࡁ࡞せᅉ࡛࠶ࡿࠋࡲࡓࠊ྾⹸࡟ࡘ࠸࡚ࡣ༢Ⅽ⏕Ṫᆺ⫢⻄㸦࠸ࢃࡺࡿ᪥ᮏ⏘⫢⻄ aspermic Fasciola sp.㸧ࡢᐤ⏕ࡶᅜෆ࡛ึࡵ࡚☜ㄆࡉࢀࡓࠋ࣭ ㇜⭠⤖⠇⹸ Oesophagostomum dentatum ࡣ࿴ḷᒣ┴⏘࢖ࣀࢩࢩ࡛ࡣ㧗࠸ᐤ⏕⋡
㸦55%㸧࡛࠶ࡗࡓࡀࠊරᗜ┴⏘ಶయ࡛ࡣࡈࡃ⛥࡞ᐤ⏕࡛࠶ࡗࡓࡇ࡜㸦3.8%㸧ࠊ࿴ḷᒣ┴⏘࢖ࣀࢩࢩ࡛ࡣᐤ⏕ࡀ☜ㄆࡉࢀ࡞࠿ࡗࡓ᭷ṑ㇜⫶⹸ࡢឤᰁಶయࡀ㜰⚄ᆅᇦ࡟㞟୰ࡋ࡚࠸ࡿࡇ࡜࡞࡝ࠊ࢖ࣀࢩࢩಶయ⩌࡟ࡼࡾ≉᭷ࡢᐤ⏕⹸┦ࢆࡶࡘࡇ࡜ࡀ♧၀ࡉࢀࡓࠋ࣭ ே⋇ඹ㏻ᐤ⏕⹸⑕ࡢཎᅉ࡜࡞ࡿࢻࣟࣞࢫ㢡ཱྀ⹸ࡢᐤ⏕⋡ࡣරᗜ┴ᮏᕞ㒊࡛ࡣᆅᇦࢆၥࢃࡎ࡟㧗࠿ࡗࡓࡀࠊಶయ࠶ࡓࡾࡢᐤ⏕ᩘࡣᴫࡋ࡚ᑡ࡞࠿ࡗࡓࠋ࢖ࣀࢩࢩ࠿ࡽ
┤᥋ឤᰁࡍࡿࡇ࡜ࡣ࡞࠸ࡀࠊᮏ✀ࡢศᕸᆅ࡛࠶ࡿࡇ࡜ࢆ⌮ゎࡋࠊຍ⇕୙༑ศ࡞⊦⫗࠶ࡿ࠸ࡣῐỈ㨶ࡢ⏕㣗➼࡛ឤᰁࡋ࡚⓶⭵∐⾜⑕࡜࡞ࡿ༴㝤ᛶ࡟␃ពࡍࡿᚲせࡀ࠶ࡿࠋKey words㸸ᐤ⏕⽸⹸┦ࠊරᗜ┴ࠊ⏕≀ከᵝᛶࠊ␿Ꮫࡣࡌࡵ࡟
ᮏ㑥࡟ࡣࢽ࣍ࣥ࢖ࣀࢩࢩ㸦Sus scrofa leucomystax㸧࡜ࣜࣗ࢘࢟ࣗ࢘࢖ࣀࢩࢩ㸦Sus

scrofa riukiuanus㸧ࡀศᕸࡋ㸦Watanobe et al. 1999㸧ࠊࡑࡢ␗࡞ࡿ⏕≀ᆅ⌮Ꮫⓗ఩⨨࡙ࡅࡶ࠶ࡗ࡚ࠊࡑࢀࡒࢀࡢᐤ⏕⹸┦࡟ࡣ≉Ⰽࡀ࠶ࡿ㸦Shoho & Machida 1979; ෆ⏣࡯ᒣཱྀ኱Ꮫඹྠ⋇་Ꮫ㒊⮫ᗋ⋇་Ꮫㅮᗙ⋇་ᐤ⏕⹸⑓Ꮫศ㔝࣭ රᗜ┴᳃ᯘື≀◊✲ࢭࣥࢱ࣮࣭ රᗜ┴❧኱Ꮫ⮬↛࣭⎔ቃ⛉Ꮫ◊✲ᡤ 

රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-9

࠿ 1984; Sato et al. 2008㸧ࠋ୧⪅࡟ඹ㏻ࡍࡿᐤ⏕⹸࡜ࡋ࡚ࠊ㇜⫵⹸ Metastrongylus spp. 
3 ✀ࠊ㢮෇㇜⫶⹸ Ascarops strongylinaࠊࢻࣟࣞࢫ㢡ཱྀ⹸ Gnathostoma doloresi㺂Globo-

cephalus samoensisࠊ࢖ࣀࢩࢩ⭠⤖⠇⹸ Oesophagostomum watanabei ࡀ࠶ࡆࡽࢀࡿ㸦Yamaguti 1961; Noda 1973; Chinose et al. 1984; ෆ⏣࡯࠿ 1984; Morita et al.
2007㸧ࠋḢᕞ࡛ࡣࠊ㏆ᖺࠊே⋇ඹ㏻ᐤ⏕⹸⑕ࡢཎᅉ࡜࡞ࡿ᪕ẟ⹸ Trichinella spp.ࡢಖ
⹸ᐟ୺࡜ࡋ࡚࣮ࣚࣟࢵࣃ࢖ࣀࢩࢩ㸦Sus scrofa subspp.㸧ࡢ㔜せᛶࡀㄆ㆑ࡉࢀࠊຍ⇕୙༑ศ࡞⊦⫗ࡢႚ㣗ࢆ௓ࡋࡓ᪕ẟ⹸⑕ࡢ㞟ᅋឤᰁ஦౛ࡢⓎ⏕ࡀ▱ࡽࢀ࡚࠸ࡿ
㸦Pérez-Martín et al. 2000; Gari-Toussaint et al. 2005㸧ࠋ୍᪉ࠊᅜෆศᕸࡢ࢖ࣀࢩࢩ
࠿ࡽࡢ᪕ẟ⹸ឤᰁࡢグ㘓ࡣࡇࢀࡲ࡛ࡢ࡜ࡇࢁ࡞࠸ࡀࠊࢻࣟࣞࢫ㢡ཱྀ⹸ࠊ Onchocerca

dewittei ࠊ ࢘ ࢙ ࢫ ࢸ ࣝ ࣐ ࣥ ⫵ ྾ ⹸ Paragonimus westermani ࠊ ኱ 㖋 㢌 ⹸
Macracanthorhynchus hirudinaceus ࡢ ಖ ⹸ ᐟ ୺ ࡜ ࡋ ࡚ ࠊ ࡲ ࡓ ࠊ ࢺ ࢟ ࢯ ࣉ ࣛ ࢬ ࣐
Toxoplasma gondii ➼ࡢே⋇ඹ㏻ឤᰁᛶཎ⹸ࡢಖ⹸ᐟ୺࡜ࡋ࡚ࠊබ⾗⾨⏕ୖࡢὀពࡀᡶ
ࢃࢀࡡࡤ࡞ࡽ࡞࠸㸦Miyazaki & Habe 1976; Kamiya et al. 1977; Miyazaki et al. 1978;ᮧ⏣ 1988; Nawa & Imai 1989; Uni et al. 2001; Takaoka et al. 2005㸧
ࠋSato et al. 㸦2008㸧ࡣࠊ2005㹼2006 ᖺࡢ⊁⊟ᮇ࡟࿴ḷᒣ┴ෆ࡛཰㞟ࡉࢀࡓ 20 㢌ࠊරᗜ┴ෆ࡛཰㞟ࡉࢀࡓ 9 㢌ࢆ᳨ᰝᑐ㇟࡜ࡋ࡚ࠊࢽ࣍ࣥ࢖ࣀࢩࢩࡢෆ㒊ᐤ⏕⽸⹸ࢆヲ⣽࡟ㄪ࡭ࠊ⥺⹸ 18 ✀࡜᮲⹸㸯✀ࠊィ 19 ✀ࢆሗ࿌ࡋ࡚࠸ࡿࠋ୧ᆅᇦ㛫࡛ᐤ⏕⹸┦࡟኱ࡁ࡞㐪࠸ࡣ࡞࠸ࡀࠊ኱⭠ᐤ⏕ࡢ㇜⭠⤖⠇⹸ Oesophagostomum dentatum ࡀරᗜ┴⏘ಶయ࠿ࡽࡣ☜ㄆࡀ࡞ࡃࠊ࿴ḷᒣ┴⏘ಶయ࡛ࡣ 55㸣ࡢಶయ࡛☜ㄆࡉࢀࡓࡇ࡜࠿ࡽࠊ㏆㞄ᆅᇦ࡟ࡶ㛵ࢃࡽࡎ୍㒊࡛ࡣ␗࡞ࡿᐤ⏕⹸ศᕸࡀࡳࡽࢀࡿྍ⬟ᛶࡀ♧၀ࡉࢀ࡚࠸ࡿࠋᮏ◊✲࡛ࡣࠊරᗜ┴ୗ࠿ࡽࡢ࢖ࣀࢩࢩ᳨ᰝᩘࢆ඘ᐇࡉࡏࠊ࿴ḷᒣ┴ୗࡢ࢖ࣀࢩࢩࡢᐤ⏕≧ἣ࡜ࡢẚ㍑ࠊ୪ࡧ࡟ྠ୍┴ෆ࡛␗࡞ࡿᆅᇦ࡟⏕ᜥࡍࡿಶయ⩌࡛ࡢᐤ⏕≧ἣࡢ≉ᚩ࡙ࡅࢆヨࡳࡓࠋᮦᩱ࡜᪉ἲ
᳨ᰝᮦᩱ
2007 ᖺ 8 ᭶ 8 ᪥࠿ࡽ 2010 ᖺ 12 ᭶ 8 ᪥࡟ࢃࡓࡾࠊරᗜ┴ୗ࡛୺࡜ࡋ࡚᭷ᐖᤕ⋓ࡉࢀࡓࢽ࣍ࣥ࢖ࣀࢩࢩ 132 㢌ࢆ཰㞟ࡋࡓࠋణࡋࠊ2007 ᖺࡣ 1 㢌ࠊ2008 ᖺࡣ 6 㢌࡛࠶ࡾࠊ୺యࡣ 2009 ᖺ 5 ᭶࠿ࡽ 2010 ᖺ 12 ᭶࡟࠿ࡅ࡚ᤕ⋓ࡉࢀࡓಶయ࡛࠶ࡿࠋ᭶ู࡟ࡳࡿ࡜ࠊ
1 ᭶ 9 㢌ࠊ2 ᭶ 11 㢌ࠊ3 ᭶ 2 㢌ࠊ4 ᭶ 0 㢌ࠊ5 ᭶ 11 㢌ࠊ6 ᭶ 12 㢌ࠊ7 ᭶ 8 㢌ࠊ8 ᭶ 10
㢌ࠊ9 ᭶ 23 㢌ࠊ10 ᭶ 12 㢌ࠊ11 ᭶ 19 㢌ࠊ12 ᭶ 13 㢌࡛࠶ࡗࡓࠋᆅᇦẚ㍑ࢆ⾜࠺ࡓࡵࠊᤕ⋓ࡉࢀࡓᆅᇦࡈ࡜࡟㜰⚄ᆅᇦࠊ୹Ἴᆅᇦࠊణ㤿ᆅᇦࡢ 3 ༊ศ࡟ᆅᇦศࡅࢆ⾜ࡗࡓ㸦⾲㸯㸧ࠋᤕ⋓ࡉࢀࡓ࢖ࣀࢩࢩࡣẅฎศᚋࡍࡳࡸ࠿࡟ゎయࡋࠊႃဗ㢌㒊࠿ࡽ┤⭠࡟࠿ࡅ࡚ࡢ
࿧྾ჾ⣔⮚ჾ㸦Ẽ⟶࣭⫵㸧ࠊᚰ⮚ࠊᾘ໬⟶㸦㣗㐨ࠥ┤⭠㸧ࠊ⫢⮚ࢆྲྀࡾฟࡋࡓࠋࡇࢀࡽෆ
⮚ࡣරᗜ┴᳃ᯘື≀◊✲ࢭࣥࢱ࣮࡟ᣢࡕᖐࡾࠊ㸫20C ࡛෭෾ಖᏑࡋࠊ㡰ḟࠊᒣཱྀ኱Ꮫ
ᐤ⏕⹸Ꮫ◊✲ᐊ࡟෭෾≧ែ࡛⛣㏦ࡋࡓࠋ࡞࠾ࠊ⭈⮚࡞ࡽࡧ࡟ࡑࡢ࿘ᅖ⬡⫫⤌⧊ࡣ࢖ࣀࢩ
ࢩࡢᰤ㣴Ꮫⓗศᯒ࡟౪ࡋࡓࡓࡵࠊ┤᥋࡟ࡣᐤ⏕⹸Ꮫⓗ᳨ᰝࡣ⾜ࡗ࡚࠸࡞࠸ࠋ 

රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-9

ᐤ⏕⹸Ꮫⓗ᳨ᰝ
෭෾ಖᏑࡉࢀࡓෆ⮚ࡣᐊ࡟࡚ゎ෾ࡋࠊ

⮚ჾู࡟ᐤ⏕⹸Ꮫⓗ᳨ᰝࢆᐇ᪋ࡋࡓࠋࡍ
࡞ࢃࡕࠊᾘ໬⟶ࡣ⦪࡟㛤ࡁࠊࢢ࣮ࣟࣈࢆ
ࡣࡵࡓᡭᣦ࡛⢓⭷㠃ࢆࡼࡃ᧿㐣ࡋ࡚ࠊỈ㐨Ỉ࡟ෆᐜ≀࡜⢓⭷ࢆࡼࡃὙ࠸ⴠ࡜ࡋࡓࠋ
┠㛤ࡁ 5.6mm ࡢ⢒࠸ࢫࢸࣥࣞࢫ〇⠠࠸㸦ᮾிࢫࢡ࣮ࣜࣥᰴᘧ఍♫ࠊᮾி㒔༓௦⏣༊㘫෬⏫㸧ࢆ㏻ࡋ࡚኱ᆺࡢࢦ࣑ࢆ㝖ཤࡋࠊ ṧࡾࡢᠱ⃮ᾮࢆ┠㛤ࡁ 300μm ࡢࢫࢸࣥࣞࢫ⠠࠸㸦ᮾிࢫࢡ࣮ࣜࣥᰴᘧ఍♫㸧ୖ࡛Ỉ㐨Ỉ࡟࡚ࡼࡃὙ࠸ࠊᚤ⣽࡞ࢦ࣑ࢆ㝖ཤࡋࡓࠋ⠠࠸ୖ࡟ṧࡗࡓෆᐜ≀ࢆỈ㐨Ỉ࡟ᡠࡋࠊࡇࢀࢆᐇయ㢧ᚤ㙾ୗ࡛⢭ᰝࡋࡓࠋࡲࡓࠊ〄㢌᮲⹸ࠊᅇ⹸ࡸ㢡ཱྀ⹸ࠊ኱⭠ᐤ⏕ྛ✀⥺⹸➼ࠊ⫗║ⓗ࡟☜ㄆ࡛ࡁࡿᐤ⏕⹸ࡣࠊ☜ㄆࡢ㒔ᗘࠊ㐺ᐅᅇ཰ࡋࡓࠋ
᳨ฟࡉࢀࡓᐤ⏕⹸ࡣ⏕⌮㣗ሷỈ࡟ࡲࡎᅇ཰ࡋࠊࡑࡢᚋࠊ10%୰ᛶ⦆⾪࣐࣍ࣝࣜࣥᾮ
࠶ࡿ࠸ࡣ 70%࢚ࢱࣀ࣮ࣝ࡟࡚ᅛᐃ࣭ಖᏑࡋࡓࠋࡲࡓࠊ୍㒊ࡢ⹸య࡟ࡘ࠸࡚ࡣࠊ㑇ఏᏊ᳨ᰝ࡟౪ࡍࡿࡓࡵ࡟㸫20C ࡛෭෾ಖᏑࡋࡓࠋᙧែᏛⓗゎᯒࡣࠊඛࡢሗ࿌㸦Sato et al.
2008㸧࡜ྠᵝ࡟ࠊᖖἲ࡟ᚑ࠸ᐇ᪋ࡋࡓࠋィ್ࡣᖹᆒ್±ᶆ‽೫ᕪ࡛♧ࡋࡓࠋ࡞࠾ࠊ

୍㒊ࡢᐤ⏕⹸ࡢᙧែᏛⓗ≉ᚩ࡟ࡘ࠸࡚ࡣబ⸨㸦2007㸧࡟ᅗゎࡋ࡚࠸ࡿࠋ
ศᏊ㑇ఏᏛⓗゎᯒ࢚ࢵ࣌ࣥࢻࣝࢳ࣮ࣗࣈ࡟ධࢀ㸫20C ࡛෭෾ಖᏑࡋ࡚࠸ࡓᐤ⏕⹸࡟ࡘ࠸࡚ࠊIllustrae

tissue & cells genomicPrep Mini Spin Kit 㸦GE Healthcare UK, Buckinghamshire, UK㸧ࢆ⏝࠸࡚ DNA ᢳฟࢆ⾜ࡗࡓࠋᡭᢏ࡟ࡘ࠸࡚ࡣࠊ࢟ࢵࢺ࡟௜ᖏࡍࡿㄝ᫂᭩
࡟ࡋࡓࡀࡗࡓࠋ᰾ DNA ࡢࣜ࣎ࢯ࣮࣒ RNA 㑇ఏᏊ㸦rDNA㸧࡞ࡽࡧ࡟࣑ࢺࢥࣥࢻࣜ࢔
DNA ࡢࢳࢺࢡ࣮࣒ࣟ c ࢜࢟ࢩࢲ࣮ࢮ࣭ࢧࣈࣘࢽࢵࢺ I 㸦 cox-1 㸧ࡢ PCR ቑᖜࡣ
Makouloutou et al.㸦2013㸧ࡢ㏻ࡾ࡟ᐇ᪋ࡋࡓࠋ࡞࠾ࠊ⏝࠸ࡓ DNA ࣏࣓࣮ࣜࣛࢮࡣ
BlendTaq-Plus-㸦TOYOBO; ኱㜰ᕷ໭༊㸧࡛ࠊ཯ᛂࡣ㸶㐃 PCR ࢳ࣮ࣗࣈ࡟ DNA ᢳฟᾮ 1μl ࢆྵࡴ཯ᛂᾮ 20μl ࢆධࢀ࡚⾜ࡗࡓࠋPCR ⏘≀ࡣ Ethidium bromide ྵ᭷ 1.3%
࢔࣮࢞ࣟࢫࢤࣝ࡟࡚㟁ẼὋືࡋࠊࢺࣛࣥࢫ࢖࣑ࣝࢿ࣮ࢱ࡛ྍど໬ࡋࠊࣂࣥࢻ㛗ࢆࢳ࢙ࢵࢡࡋࡓࠋ┠ⓗ࡟㐺࠺ PCR ⏘≀ࡣ FastGene Gel/PCR Extraction Kit㸦NIPPON
Genetics Co., Tokyo, Japan㸧ࢆ⏝࠸࡚⢭〇ࡋࢩ࣮ࢡ࢚ࣥࢫࢆ⾜ࡗࡓࠋ㔜」ࡍࡿࢭࢢ࣓
ࣥࢺ࡜ࡋ࡚ࢩ࣮ࢡ࢚ࣥࢫࡉࢀࡓ rDNA ࢩ࣮ࢡ࢚ࣥࢫࡣࠊCLUSTAL W multiplealignment programme㸦Thompson et al. 1994㸧ࢆ᥼⏝ࡋ࡚⹸యẖ࡟ 1 ᮏ࡟ࡋࡓࠋ 

රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-9

⤖ᯝරᗜ┴ୗ࡛཰㞟ࡋࡓ࢖ࣀࢩࢩ 132 㢌ࡢ⬚⭍࡞ࡽࡧ࡟⭡⭍⮚ჾ࠿ࡽ 24 ✀ࡢᐤ⏕⽸⹸ࢆ
᳨ฟࡋࡓ㸦⾲㸰㸧ࠋᐟ୺ࡢᤕ⋓᫬ᮇࡸᛶู࡟ࡼࡾឤᰁ⋡ࡀ㢧ⴭ࡟␗࡞ࡿᐤ⏕⹸✀ࡣ࡞࠸ࡀࠊ⏘ᆅࢆణ㤿ᆅᇦ㸦17 㢌㸧ࠊ୹Ἴᆅᇦ㸦47 㢌㸧ࠊ㜰⚄ᆅᇦ㸦53 㢌㸧࡜༊ษࡿ࡜ࠊᆅᇦ
࡟ࡼࡾ᳨ฟ⋡࡟኱ࡁ࡞ᕪࡀࡳࡽࢀࡿ✀ࡀ࠶ࡗࡓࠋࡍ࡞ࢃࡕࠊ୹Ἴᆅᇦࡢ࢖ࣀࢩࢩ࡛ࡣ௒
ᅇ᳨ฟࡉࢀࡓࡍ࡭࡚ࡢ⽸⹸✀ࡢᐤ⏕ࡀ࠶ࡗࡓࡀࠊ༢Ⅽ⏕Ṫᆺ⫢⻄ aspermic Fasciola sp.
㸦ᅗ㸯; ᕷᕝ࣭ᯈᇉ 2013㸧ࠊ㇜⭈⹸ Stephanurus dentatusࠊ⨾㯇㣗㐨⹸ Gongylonema

pulchrum㸦ᅗ㸰㸧ࡣణ㤿ᆅᇦࠊ㜰⚄ᆅᇦ࠿ࡽࡢ᳨ฟࡣ࡞࠸࠿ࠊ࠶ࡿ࠸ࡣࡈࡃ⛥࡛࠶ࡗࡓࠋ୍᪉ࠊ᭷ṑ㇜⫶⹸ Ascarops dentata㸦ᅗ㸱㸧ࡣ୺࡟㜰⚄ᆅᇦ࡛ᤕ⋓ࡉࢀࡓ࢖ࣀࢩ
ࢩ࠿ࡽ㧗⋡࡟᳨ฟࡉࢀࠊ௚ᆅᇦ࡛ࡢ᳨ฟࡣࡈࡃ㝈ࡽࢀࡓࠋ㜰⚄ᆅᇦ࡛ࡣࠊ᪥ᮏ〄㢌᮲⹸

Pseudanoplocephala nipponensis㸦ᅗ㸲㸧ࡢឤᰁ⋡ࡶ㧗ࡃࠊࡲࡓࠊ௚ᆅᇦ࡛ࡣ 64 㢌୰
1 㢌࠿ࡽࡢࡳ᳨ฟࡉࢀࡓ㇜⭠⤖⠇⹸ࡀࠊ㜰⚄ᆅᇦ 53 㢌୰ 4 㢌࠿ࡽ☜ㄆࡉࢀࡓࠋࡲࡓࠊ
ᓥᕋ㒊ࡢῐ㊰ᓥ࠿ࡽࡢ᳨ᰝಶయᩘࡣ 4 㢌࡟㝈ࡽࢀࠊᐤ⏕⹸᳨ฟࡣ 5 ✀ࡢࡳ࡛࠶ࡗࡓࠋࡲ
ࡓࠊ㔝⏕ಶయࢆᤕ⋓ࡋࠊ1 ᖺ௨ୖ࡟ࢃࡓࡾ㣫⫱ࡋ࡚࠸ࡓ 11 㢌࡛ࡶࠊ㔝⏕ಶయ࡛ࡣ㧗⋡࡟ᐤ⏕ࡍࡿ⫵⹸ࡢᐤ⏕ࡀࡳࡽࢀࡎࠊࢻࣟࣞࢫ㢡ཱྀ⹸ࢆࡣࡌࡵ඲⯡ⓗ࡟ᐤ⏕⋡ࡶప࠸ഴྥ
ࡀࡳࡽࢀࡓࠋ
⨾㯇㣗㐨⹸ࡢᅜෆ࢖ࣀࢩࢩ࡛ࡢᐤ⏕ࡀ☜ㄆࡉࢀࡓࡢࡣ௒ᅇࡢරᗜ┴࡛ࡢㄪᰝࡀึࡵ
࡚࡛࠶ࡗࡓࡇ࡜࠿ࡽࠊ18S rDNA ࡞ࡽࡧ࡟ cox-1 ሷᇶ㓄ิ࡟ࡘ࠸᳨࡚ウࡋࡓࠋ࣍ࣥࢩࣗ
࢘ࢪ࢝㸦Cervus nippon㸧ࡸࢽ࣍ࣥࢨࣝ㸦Macaca fuscata㸧࡞࡝ᅜෆ㔝⏕ື≀࡟ᐤ⏕ࡋ࡚࠸ࡿ⣔⤫࡜ᐙ␆࡛࠶ࡿ࢘ࢩᐤ⏕ࡢ⣔⤫࡜ࡀ༊ูࡉࢀ࡚࠸ࡿࡀ㸦బ⸨ 2009㸧ࠊ௒ᅇ࢖ࣀࢩࢩ࠿ࡽᚓࡓ⣔⤫ࡣ๓⪅࡟ᒓࡋ࡚࠸ࡓࠋヲࡋࡃࡣࠊูࡢㄽᩥ࡛ሗ࿌ࡋ࡚࠸ࡿ
㸦Makouloutou et al. 2013㸧ࠋ⾲㸰࡟࠾࠸࡚ࠊẼ⟶ᨭෆᐤ⏕ࡢ⫵⹸ Metastrongylus spp.ࡣ୍ᣓࡋࡓࡀࠊࡇࢀ࡟ࡣ

M. elongatusࠊM. salmiࠊM. asymmetricusࠊM. pudendotechus ࡢ 4 ✀ࡀྵࡲࢀ࡚࠸
ࡿࠋࡲࡓࠊ⫶⢓⭷ࡸ⢓ᾮ୰࡟ᐤ⏕ࡋ࡚࠸ࡿẟ⣽⥺⹸⛉⥺⹸ Capillarid spp.࡟ࡘ࠸࡚ࡶ୍
ᣓࡋࡓࡀࠊࡇࢀ࡟ࡣ Aonchotheca suisࠊAonchoteca riukiuensis䚸Aonchotheca sp. ࡢ
3 ✀ࡀ༊ูࡉࢀࡓ㸦ᡞ⏣࣭బ⸨ 2010㸧ࠋ௒ᅇࡢ᳨ᰝᑐ㇟ಶయࡢ୰࠿ࡽẟ⣽⥺⹸ࡀከᩘᐤ
⏕ࡋ࡚࠸ࡓ 14 㢌ࢆ㑅ࡧࠊA. suisࠊA. riukiuensis䚸Aonchotheca sp.ࡢᐤ⏕⋡ࢆࡳࡿ࡜ࡑࢀࡒࢀ 64.3%ࠊ78.6%ࠊ14.3%࡛࠶ࡗࡓࠋࡇࢀࡽ 3 ✀࡟ࡘ࠸࡚ࠊ௨ୗ࡟ᙧែᏛⓗ≉ᚩࢆグࡍࠋ  㻝㻡㻥㻟㻥㻞㻣㻻㼑㼟㼛㼜㼔㼍㼓㼛㼟㼠㼛㼙㼡㼙㻌㼐㼑㼚㼠㼍㼠㼡㼙㼀㼞㼕㼏㼔㼡㼞㼕㼟㻌㼟㼡㼕㼟㻝㻜㻥㻼㼟㼑㼡㼐㼍㼚㼛㼜㼘㼛㼏㼑㼜㼔㼍㼘㼡㼟㻌㼚㼕㼜㼜㼛㼚㼑㼚㼟㼕㼟㻟㻥㻤㻢㻡㻿㼠㼞㼛㼚㼓㼥㼘㼛㼕㼐㼑㼟㻌㼞㼍㼚㼟㼛㼙㼕㻻㼑㼟㼛㼜㼔㼍㼓㼛㼟㼠㼛㼙㼡㼙㻌㼣㼍㼠㼍㼚㼍㼎㼑㼕㻞㻠㻳㼘㼛㼎㼛㼏㼑㼜㼔㼍㼘㼡㼟㻌㼘㼛㼚㼓㼑㼙㼡㼏㼞㼛㼚㼍㼠㼡㼟㻮㼡㼞㼓㼑㼘㼍㼠㼕㼍㻌㼐㼕㼐㼡㼏㼠㼍㻝㻞㻤㻡㻣㻳㼘㼛㼎㼛㼏㼑㼜㼔㼍㼘㼡㼟㻌㼟㼍㼙㼛㼑㼚㼟㼕㼟㻞㻝㻠㻭㼟㼏㼍㼞㼕㼟㻌㼟㼡㼡㼙㻯㼍㼜㼕㼘㼘㼍㼞㼕㼐㻌㼟㼜㼜㻚㻜㻜㻟㻌㻔㻢㻜㻚㻜㻕

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㻟㻌㻔㻢㻜㻚㻜㻕㻜㻜㻜㻟㻌㻔㻢㻜㻚㻜㻕

䗢㻌㻔㻡㻕㻝㻌㻔㻠㻚㻤㻕
㻣㻌㻔㻟㻟㻚㻟㻕

㻝㻌㻔㻤㻚㻟㻕

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㻝㻡㻌㻔㻡㻣㻚㻣㻕㻜㻞㻞㻌㻔㻤㻠㻚㻢㻕㻞㻡㻌㻔㻥㻢㻚㻞㻕㻝㻌㻔㻟㻚㻤㻕

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㻝㻢㻌㻔㻢㻝㻚㻡㻕㻞㻝㻌㻔㻤㻜㻚㻤㻕㻝㻤㻌㻔㻢㻥㻚㻞㻕㻠㻌㻔㻝㻡㻚㻠㻕

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┴୰㒊㻜㻣㻌㻔㻡㻤㻚㻟㻕

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䗠㻌㻔㻝㻞㻕

┴໭㒊㻖㻌ឤᰁಶయᩘ㻔ឤᰁ䠂㻕䜢䛭䜜䛮䜜♧䛩㻚㻖㻖㻌㻹㻚㻌㼑㼘㼛㼚㼓㼍㼠㼡㼟㻘㻌㻹㻚㻌㼟㼍㼘㼙㼕㻘㻌㻹㻚㻌㼍㼟㼥㼙㼙㼑㼠㼞㼕㼏㼡㼟㻘㻌㻹㻚㻌㼜㼡㼐㼑㼚㼐㼛㼠㼑㼏㼔㼡㼟 㻌䠐✀䜢ಶู䛻༊ู䛧䛶䛔䛺䛔㻚

኱⭠

ᑠ⭠

㻡㻝㻞

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㻠

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᭱኱ᅇ཰
⹸యᩘ

㻳㼚㼍㼠㼔㼛㼟㼠㼛㼙㼍㻌㼐㼛㼘㼛㼞㼑㼟㼕㻳㼛㼚㼓㼥㼘㼛㼚㼑㼙㼍㻌㼜㼡㼘㼏㼔㼞㼡㼙㣗㐨㻌㻔⢓⭷ෆ㻕㻿㼠㼑㼜㼔㼍㼚㼡㼞㼡㼟㻌㼐㼑㼚㼠㼍㼠㼡㼟⭈⮚䞉࿘ᅖ⬡⫫⤌⧊㻭㼟㼜㼑㼞㼙㼕㼏㻌㻲㼍㼟㼏㼕㼛㼘㼍 㻌㼟㼜㻚⫹⟶㻔⫢⮚㻕

⫶✀ྡ㻹㼑㼠㼍㼟㼠㼞㼛㼚㼓㼥㼘㼡㼟 㻌㼟㼜㼜㻚㻖㻖Ẽ⟶ᨭ㻌㻔⫵㻕⮚ჾ㻟㻌㻔㻥㻚㻣㻕

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┴༡㒊㻝㻌㻔㻞㻡㻚㻜㻕

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䗠㻌㻔㻡㻕

㣫⫱ಶయ㻌㻔┴୰㒊㻕⏕ά⎔ቃࡢởᰁࡸ୰㛫ᐟ୺࡜࡞ࡿື᳜≀ࡢႚ㣗ࢆ௓ࡋ࡚ே⋇ඹ㏻ᐤ⏕⹸ࡢཎᅉ࡜࡞ࡿྍ⬟ᛶࢆᣢࡘᐤ⏕⹸࡟ࡘ࠸࡚ࡣୗ⥺ࢆᘬ࠸ࡓࠋ⾲㸰 රᗜ┴⏘ࢽ࣍ࣥ࢖ࣀࢩࢩ࠿ࡽ᳨ฟࡉࢀࡓᐤ⏕⽸⹸ රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-9ᅗ㸯㸬࢖ࣀࢩࢩ⫹⟶࡟ᐤ⏕ࡍ ࡿ ༢ Ⅽ ⏕ Ṫ ᆺ ⫢ ⻄
DVSHUPLF)DVFLRODVS㸦ⓑ▮༳㸧௒ᅇࡢㄪᰝ࡛ࡣࡶࡗࡥࡽ୹Ἴ
ᆅᇦࡢ࢖ࣀࢩࢩ㸦 㢌୰  㢌㸧᳨࡛ฟࡉࢀࡓࠋ⫹⟶ቨࡢ⫧ཌࡀ㢧ⴭ࡛࠶ࡿࠋ࣐࣍ࣝࣜࣥᅛ
ᐃᶆᮏ㸦:㸧ࠋᅗ㸰㸬࢖ࣀࢩࢩ㣗㐨⢓⭷ෆ࡟ࢪࢢࢨࢢࢆᥥ࠸࡚ᐤ⏕ࡍ ࡿ ⨾ 㯇 㣗 㐨 ⹸*RQJ\ORQHPDSXOFKUXP 㞘㸦 ࡘࡢ▮༳࡛  ⹸యࢆ♧ࡍ㸧௒ᅇࡢㄪᰝ࡛ࡣ୹Ἴᆅᇦࡢ࢖
ࣀࢩࢩ࡛㧗⋡࡟᳨ฟࡉࢀࡓ㸦  㢌 ୰  㢌 㸧ࠋ ⏕ ᶆ ᮏ
㸦:㸧ࠋᅗ㸱㸬࢖ࣀࢩࢩ⫶ෆ࡟ᐤ⏕ࡍࡿ᭷ṑ㇜⫶⹸ $VFDURSVGHQWDWD௒ᅇࡢㄪᰝ࡛ࡣࠊ㜰⚄ᆅᇦ࡛㧗⋡࡟᳨ฟࡉࢀ㸦 㢌୰ 㢌㸧ࠊణ㤿ᆅᇦ㸦 㢌㸧࡛ࡣឤᰁಶయ࡞ࡋࠊ୹Ἴᆅᇦ࡛ࡣ 㢌୰  㢌ࡢࡳ࡟࡜࡝ࡲࡗ
ࡓࠋᮏ౛࡛ࡣᐤ⏕ᩘ  㞘ࢆᩘ࠼ࡓࠋ⏕ᶆᮏ㸦:㸧ࠋ 

රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-9ᅗ㸲㸬࢖ࣀࢩࢩᑠ⭠࡟ᐤ⏕ࡍࡿ᪥ᮏഇ〄㢌᮲⹸

3VHXGDQRSORFHSKDODQLSSRQHQVLV ୗࡢ෗┿ࡣ 㞘ࡢ㢌⠇㸦▮༳㸧࠿ࡽཷ⫾∦⠇࡟⮳ࡿࢫࢺࣟࣅࣛࢆ♧ࡍࠋ௒ᅇࡢㄪᰝ࡛ࡣࠊ㜰⚄ᆅᇦ࡛㢖⦾࡟᳨ฟࡉࢀ㸦 㢌୰ 㢌㸧ࠊణ㤿࣭୹Ἴᆅᇦ࡛ࡣ 㢌୰  㢌ࡢࡳ࡟࡜࡝ࡲࡗࡓࠋ
ᮏ౛࡛ࡣᐤ⏕ᩘ  㞘ࢆᩘ࠼ࡓࠋ⏕ᶆᮏ㸦:㸧ࠋ Aonchotheca suis㸦Yamaguti 1943㸧 㞝⹸㸦n=8㸧㸸య㛗 11.89㸫13.53㸦13.02 ± 0.01㸧mmࠊ㣗㐨㒊ࡢ㛗ࡉ 4.60㸫5.89㸦5.16± 0.43㸧mmࠊ᭱኱యᖜ 0.054㸫0.074㸦0.067 ± 0.007㸧mmࠋ㣗㐨㒊㸸⭠⟶㒊ࡢẚࡣ 1㸸
1.30㸫1.69㸦1.53 ± 0.14㸧ࠋᑿ➃ࡢᕥྑ୧ഃ⭡㠃ᐤࡾ࡟ࡣ๓ᚋ 2 ᑐࡢ✺㉳ࡀ⏕ࡌࠊࡸࡸ๓᪉࡬ྥ࠿࠺㸦ᅗ㸳㸧ࠋ๓ഃࡢ 1 ᑐࡣ༢⣧࡞ᣦ≧࡛ࠊᚋࢁഃࡢ 1 ᑐࡣඛ➃ࡀ๓᪉࡬ᒅ᭤
ࡋ࡚ࣁ࣐࣮ࣥ≧ࢆ࿊ࡍࡿࠋࡇࡢ✺㉳࡟ࠊࡼࡃⓎ㐩ࡋࡓᑿഃ⩼ࡢ㐲఩➃࠾ࡼࡧᑿ➃⫼ഃࡢᑿᄞࡀ௜╔ࡋ࡚࠸ࡿࠋ஺᥋่ࡣ⥺≧࡛୰➼ᗘ࡟◳໬ࡋࠊ㛗ࡉࡣ 0.75㸫0.82㸦0.79 ± 0.02㸧
mmࠊ㐲఩➃ࡣࣛࣥࢭࢵࢺ≧ࢆ࿊ࡍࡿࠋᑿഃ⩼࡜஺᥋่㠧࡟ࡣᶓ࡟⣽ࡃ⥺᮲ࡀධࡾࠊ஺
᥋่㠧ࡣ↓Ჲ࡛࠶ࡿࠋ 㞤⹸㸦n=23㸧㸸య㛗 13.32㸫22.77㸦18.65 ± 2.33㸧mmࠊ㣗㐨㒊ࡢ㛗ࡉ 4.88㸫6.44㸦5.87± 0.37㸧mmࠊ᭱኱యᖜ 0.063㸫0.100㸦0.083 ± 0.010㸧mmࠋ㣗㐨㒊㸸⭠⟶㒊ࡢẚࡣ 1㸸
1.64㸫2.93㸦2.18 ±0.34㸧ࠋᑿ➃ࡢᙧ≧ࡣ㕌෇≧㹼෇㗹≧࡛࠶ࡿࠋ㝜㛛ࡣ㣗㐨࣭⭠⟶⤖ྜ㒊ࡢ 0.044㸫0.203㸦0.139 ± 0.041㸧mm ᚋ᪉࡟఩⨨ࡋࠊ㝜㛛㒊࡟┠❧ࡗࡓ✺㉳ࡸ௜ᒓ≀ࡣ࡞࠸ࡀࠊ㛤ཱྀ㒊๓ᚋࡢ⭡㠃య⾲࡟ᐇ㉁ᛶ࠾ࡼࡧࢡࢳࢡࣛᛶࡢ⣽࠿࡞พฝࡀ࠶ࡾࠊ⢒⣁࡞⾲㠃ᵓ㐀ࢆ࿊ࡍࡿ㸦ᅗ㸳㸧ࠋ⹸༸ࡣ㛗ᚄࡀ 0.050㸫0.070㸦0.063 ± 0.003㸧mmࠊ▷ᚄࡀ 0.024㸫0.033㸦0.028 ± 0.002㸧mm ࡛ࠊ༸Ẇࡀࡸࡸཌࡃ୧ᴟࡢ⵹ࡣ✺ฟࡋࠊ༸Ẇ⾲㠃ࡣᖹ⁥࡛≉Ṧ࡞ᵓ㐀ࡸ⣠ᵝࡣぢࡽࢀ࡞࠸ࠋ 

රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-9ᅗ㸳㸬࢖ࣀࢩࢩࡢ⫶࡟ᐤ⏕ࡍࡿẟ⣽⥺ᅗ㸴㸬࢖ࣀࢩࢩࡢ⫶࡟ᐤ⏕ࡍࡿẟ⣽⥺⹸ $RQFKRWKHFDVXLV⹸ $RQFKRWKHFDULXNLXHQVLV㸦$㸧㞝⹸ᑿ㒊ࡢഃ㠃ീ࡛ࠊᑿ➃㏆ࡃࡢᕥྑ㸦$㸧㞝⹸ᑿ㒊ࡢഃ㠃ീ࡛ࠊᑿ➃ࡀࣁ࣐࣮ࣥ

ࡢᣦ≧✺㉳㸦▮༳㸧࡜ᑿഃ⩼㸦 㸧ࢆ♧ࡍࠋ≧✺㉳࡜࡞ࡗ࡚㢌ഃ࡟ᒅ᭤ࡍࡿ㸦 㸧ࠋ▮༳

㸦%㸧ᑿഃ⩼ࢆḞࡃ㞝⹸ࡢᑿ㒊ഃ㠃ീࠋ㸦&㸧ࡣ⥲᤼ἥཱྀ๓⦕࠿ࡽᑿഃ࡟ྥ࠿ࡗ࡚⣽㛗࠸㞤⹸ࡢ㝜㛛㸦▮㢌㸧࿘ᅖࡢഃ㠃ീࠋ㣗㐨ᮎᣦ≧✺㉳㸦▮༳㸧ࡀࡳࡽࢀࡿࠋ㸦%㸧㞝⹸ᑿ➃㒊㸦 㸧࡟㏆ࡃࠊ㝜㛛ࡀ㛤ཱྀࡍࡿࡀࠊࡑࡢ

㒊ࡢ⭡㠃ീ࡛ࠊᕥྑ࡟ࡳࡽࢀࡿ⣽㛗࠸ᣦ≧࿘ᅖࡢ⭡㠃ࡣࡁࢃࡵ࡚⢒㐀࡛ࠊ୙つ๎࡞ฝ✺㉳ࢆ♧ࡍࠋ㸦&㸧㞤⹸ࡢ㝜㛛㸦▮㢌㸧࿘ᅖพ≧࡛࠶ࡿࠋ෗┿ $ࠥ&ࠊࡍ࡭࡚ྠ୍ಸ⋡ࠋࡢഃ㠃ീࠋ㣗㐨ᮎ➃㒊㸦 㸧࡟㏆ࡃࠊ㝜㛛ࡀ㛤ཱྀࡋࠊ࣋ࣝ≧ࡢ௜ᒓ≀ࡀ࠶ࡿࠋ෗┿ $ࠥ&ࠊࡍ࡭࡚ྠ୍ಸ⋡ࠋ

㸦2㸧Aonchotheca riukiuensis㸦Shoho & Machida 1979㸧 㞝⹸㸦n=9㸧㸸య㛗 7.29㸫13.75㸦11.47 ± 2.04㸧mmࠊ㣗㐨㒊ࡢ㛗ࡉ 3.59㸫5.18㸦4.59± 0.50㸧mmࠊ᭱኱యᖜ 0.038㸫0.067㸦0.054 ± 0.009㸧mmࠋ㣗㐨㒊㸸⭠⟶㒊ࡢẚࡣ 1㸸
1.03㸫1.82㸦1.49 ± 0.30㸧ࠋయᚋ➃ࡣ 1 ᑐࡢࣁ࣐࣮ࣥ≧✺㉳࡜࡞ࡗ࡚⭡ഃ࡟ྥ࠿࠸ࠊࡑ
ࡢ㛫࡟ࡣὸ࠸⿣ࡀධࡿࠋᑿ➃⫼ഃ࡟ࡣᣦ≧✺㉳࡟ᨭᣢࡉࢀ࡚ࡼࡃⓎ㐩ࡋࡓᑿᄞࡀ࠶ࡾࠊ
ࡇࡢ㎶⦕ࡣ⢒࡛ࠊ⾲㠃࡟ࡣ⣽࠸⥺ࡀධࡗ࡚⦰⦉ᵝࢆ࿊ࡍࡿࡶࡢࡀ࠶ࡿࠋࡲࡓࠊ⫠㛛㛤ཱྀ㒊๓⦕࠿ࡽ 1 ᑐࡢ⣽࠸✺㉳ࡀ⏕ࡌ࡚ᚋ᪉࡬ྥ࠿࠺ࠋᑿ㒊ࡢᕥྑ୧ഃ࡟ࡣࠊࡈࡃᖜࡢ⊃࠸ࢡࢳࢡࣛᛶࡢᑿഃ⩼ࡀᏑᅾࡍࡿ㸦ᅗ㸴㸧ࠋ஺᥋่ࡣ⥺≧࡛ࡑࡢ㐲఩㒊ࡣ༢ㄪ࡟⣽ࡃ࡞ࡾࠊ 

රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-9

㛗ࡉࡣ 0.71㸫1.01㸦0.86 ± 0.09㸧mmࠋ஺᥋่㠧ࡣ↓Ჲ࡛ᶓ࡟⥺᮲ࡀධࡿࠋ 㞤⹸㸦n=34㸧
㸸య㛗 13.92㸫36.11㸦27.83 ± 4.70㸧mmࠊ㣗㐨㒊ࡢ㛗ࡉ 4.38㸫6.27 㸦5.24± 0.45㸧mmࠊ᭱኱యᖜ 0.044㸫0.085㸦0.069 ± 0.009㸧mmࠋ㣗㐨㒊㸸⭠⟶㒊ࡢẚࡣ 1㸸
2.18㸫5.14㸦4.29 ± 0.69㸧ࠋᑿ➃ࡢᙧ≧ࡣ㕌෇≧㹼෇㗹≧࡛࠶ࡿࠋ㝜㛛ࡣ㣗㐨࣭⭠⟶⤖ྜ㒊ࡢ 0㸫0.236㸦0.104 ± 0.041㸧mm ᚋ᪉࡟㛤ཱྀࡍࡿࠋ㝜㛛㛤ཱྀ㒊࡟ࡣ࣋ࣝ≧ࡢࢡࢳࢡࣛᛶࡢ⟄ࡀ௜ᒓࡋࠊ๓᪉ࢆྥࡃ㸦ᅗ㸴㸧ࠋ⹸༸ࡣ㛗ᚄࡀ 0.049㸫0.064㸦0.059 ± 0.002㸧
mmࠊ▷ᚄࡀ 0.020㸫0.033㸦0.027 ± 0.002㸧mm ࡛ࠊ༸Ẇ⾲㠃࡟ࡣ࣐ࢫࢡ࣓ࣟࣥᵝࡢ⥙┠ᵓ㐀ࢆㄆࡵࡿࠋ
㸦3㸧Aonchotheca sp.
 㞝⹸㸦n=2㸧
㸸య㛗 6.90㸫7.59mmࠊ㣗㐨㒊ࡢ㛗ࡉ 3.21㸫3.70mmࠊ᭱኱యᖜ 0.050㸫
0.061mmࠋ㣗㐨㒊㸸⭠⟶㒊ࡢẚࡣ 1㸸1.05㸫1.15ࠋయᚋ➃ࡣᕥྑ 2 ࡘࡢ✺㉳࡜࡞ࡗ࡚⭡ഃ࡬ᒅ᭤ࡋࠊࡑࡢඛ➃ࡣᑠ⤖⠇≧ࡲࡓࡣ㕌࡞ᣦ≧✺㉳࡟⤊ࢃࡿࠋᑿ➃⫼ഃ࡟ࡣⓎ㐩ࡢᝏ࠸ᑿᄞࡀࠊᑿ㒊ࡢᕥྑ୧ഃ࡟ࡣⓎ㐩ࡋࡓᑿഃ⩼ࡀᏑᅾࡋࠊᣦ≧✺㉳ࡀࡇࢀࡽࢆᨭᣢࡍࡿ
㸦ᅗ㸵㸧ࠋ஺᥋่ࡣ୰➼ᗘ࡟◳໬ࡋ㐲఩㒊ࡣࣛࣥࢭࢵࢺ≧ࠊ㛗ࡉࡣ 0.39㸫0.46mmࠋ஺᥋่㠧ࡣ↓Ჲ࡛ᶓ࡟⥺᮲ࡀධࡿࠋ 㞤⹸㸦n=6㸧
㸸య㛗 8.96㸫9.73㸦9.22 ± 0.35㸧mmࠊ㣗㐨㒊ࡢ㛗ࡉ 2.66㸫3.12㸦2.90 ±0.20㸧mmࠊ᭱኱యᖜ 0.053㸫0.061㸦0.056 ± 0.003㸧mmࠋ㣗㐨㒊㸸⭠⟶㒊ࡢẚࡣ 1㸸
1.90㸫2.39㸦2.18 ± 0.18㸧ࠋᑿ➃ࡢᙧ≧ࡣ෇㗹≧㸦ᅗ㸵㸧ࠋ㝜㛛ࡣ㣗㐨࣭⭠⟶⤖ྜ㒊ࡢ
0.082㸫0.132㸦0.110 ± 0.018㸧mm ᚋ᪉࡟㛤ཱྀࡍࡿࠋ㝜㛛┤๓࡛ᐇ㉁ࡢ㝯㉳㒊ࢆࢡࢳࢡࣛࡀ⦕ྲྀࡾࠊ㝜㛛㛤ཱྀ㒊࡟ࡣࢡࢳࢡࣛᛶࡢ෇࠸௜ᒓ≀ࢆㄆࡵࡿ㸦ᅗ㸵㸧ࠋ⹸༸ࡣ㛗ᚄࡀ
0.054㸫0.066㸦0.059 ± 0.003㸧mmࠊ▷ᚄࡀ 0.024㸫0.027㸦0.025 ± 0.001㸧mm ࡛ࠊ༸Ẇ⾲㠃࡟ࡣ㛗㍈᪉ྥ࡟㉮ࡿ⥙┠≧⥺᮲ࢆㄆࡵࡿ㸦ᅗ㸵㸧ࠋ㸺ഛ⪃㸼 Aonchotheca suis㸦Yamaguti 1943㸧ࡣࠊᒣཱྀ┴࠾ࡼࡧ࿴ḷᒣ┴ࡢ㔝⏕ࡢࢽ࣍ࣥ࢖ࣀ
ࢩࢩࡢ⫶࠿ࡽศ㞳ࡉࢀࠊCapillaria suis ࡜ࡋ࡚✀グ㍕ࡉࢀࡓ㸦Yamaguti 1943㸧ࠋ㞝⹸ࡢᑿ✺㉳ࡢ≉ᚩⓗ࡞ᙧ≧ࡸ㞤⹸ࡢ㝜㛛㒊࡟≉ู࡞௜ᒓᵓ㐀≀ࡀ࡞࠸ࡇ࡜ࠊ⹸༸ࡢ༸Ẇ⾲㠃ࡀᖹ⁥࡛࠶ࡿࡇ࡜࡞࡝ࠊ௒ᅇほᐹࡉࢀࡓ≉ᚩࡣཎグ㍕࡜ࡼࡃ୍⮴ࡍࡿࠋణࡋࠊ㞝⹸ࡢᑿഃ⩼࡟㛵ࡋ࡚ࡣཎグ㍕࡟୍ษグ㏙ࡀ࡞ࡃࠊཎグ㍕ࡢ௜ᅗ࡟ࡶᑿഃ⩼ࡣᥥ࠿ࢀ࡚࠸࡞࠸ࠋ
௒ᅇ᳨ฟࡋࡓ⹸య࡛ࡣṤ࡝ࡍ࡭࡚ࡀࡼࡃⓎ㐩ࡋࡓᑿഃ⩼ࢆ᭷ࡋ࡚࠸ࡓࠋᑿഃ⩼ࢆకࢃ࡞࠸⹸యࡣ࿴ḷᒣ┴⏘ࢽ࣍ࣥ࢖ࣀࢩࢩ 1 㢌࠿ࡽᅇ཰ࡉࢀࡓ㞝⹸ 21 㞘ࡢ࠺ࡕࡢ 2 㞘ࡢࡳ࡛࠶ࡗࡓ㸦ᅗ㸳B㸧ࠋᑿഃ⩼ࡢ᭷↓௨እࡢᙧែⓗ≉ᚩࡸィ್࡟ࡘ࠸࡚ࡣࠊࡇࡢ 2 ࢱ࢖ࣉࡢ㞝⹸ࡢ㛫࡛ᕪ␗ࡣぢࡽࢀ࡞࠿ࡗࡓࡇ࡜ࠊྠ᫬࡟ᅇ཰ࡋࡓ㞤⹸࡟ᙧែᏛⓗ࡞ኚ␗ࢆ≉࡟ᣦ᦬ࡍࡿࡇ࡜ࡣ࡛ࡁ࡞࠿ࡗࡓࡇ࡜࠿ࡽࠊᑿഃ⩼ࡢ᭷↓࡟㛵ࢃࡽࡎྠ୍✀ A. suis ࡜⪃࠼ࡽࢀࡓࠋ 

රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-9ᅗ㸵㸬࢖ࣀࢩࢩࡢ⫶࡟ᐤ⏕ࡍࡿẟ⣽⥺⹸ᮍグ㍕✀ $RQFKRWKHFDVS
㸦$㸧㞝⹸ᑿ㒊ࡢഃ㠃ീ࡛ࠊ஺᥋่㠧ࡀ⥲᤼ἥཱྀࡼࡾ⩻㌿⬺ฟࡋ࡚࠸ࡿࡇ࡜㸦▮༳㸧ࢆ♧ࡍࠋ㸦%㸧㞤⹸ࡢ㝜㛛㸦▮㢌㸧࿘ᅖࡢഃ㠃ീࠋ㣗㐨ᮎ➃㒊㸦 㸧࡜㝜㛛ࡢ୰㛫ࡢ⭡㠃࡟ࢡࢳࢡ࡛ࣛ⦕ྲྀࡽࢀࡓ㢧ⴭ࡞㝯㉳㸦▮༳㸧ࡀ࠶ࡾࠊࡲࡓࠊ㝜㛛ࡣ኱ࡁ࡞ࢻ࣮࣒ᆺࡢ✺ฟ≀ࢆక࠺ࠋ㸦&㸧㞤⹸ᑿ㒊ࡢഃ㠃ീࠋᑿ➃ࡣ෇㗹≧ࢆ࿊ࡋࠊ⫠㛛㸦▮㢌㸧ࡣᮎ➃㏆ࡃ࡟㛤ཱྀࡍࡿࠋ㸦'㸧㝜㛛㏆ࡃ࡛⏘ฟࡉࢀࡿ┤๓ࡢ⹸༸࡟ࡣࠊ㛗㍈᪉ྥ࡟㉮ࡿ⥺᮲ࡀ࠾஫࠸࡟஺ཫࡋࠊ୙つ๎࡞⥙┠ᶍᵝࢆぢࡏࡿࠋ෗┿ $ࠥ'ࠊࡍ࡭࡚ྠ୍ಸ⋡ࠋ Aonchotheca riukiuensis ࡣࠊす⾲ᓥࡢࣜࣗ࢘࢟ࣗ࢘࢖ࣀࢩࢩࡢ⫶࠿ࡽ᳨ฟࡉࢀ✀グ㍕ࡉࢀࡓ㸦Shoho & Machida 1979㸧ࠋཎグ㍕࡜௒ᅇࡢᅇ཰⹸యࡢᙧែᏛⓗ≉ᚩࡣࡼࡃ୍⮴ࡋࡓࠋ࡞࠾ཎグ㍕࡛ࡣ㞝⹸ࡢ⫠㛛㛤ཱྀ㒊┤๓ࡢ✺㉳ࢆṇ୰࡟୍ᮏ⏕ࡌࡿ࡜グ㘓ࡋ࡚࠸ࡿࡀࠊࡇࡢ✺㉳ࡀᐇ㝿࡟ࡣᕥྑᑐ㇟࡟ 1 ᑐᏑᅾࡍࡿࡇ࡜ࡀ௒ᅇࡢほᐹ࡛☜ㄆࡉࢀࡓࠋ ௒ᅇ✀ᮍྠᐃ࡜ࡋࡓ Aonchotheca sp.ࡣ㞝⹸ࡀⓎ㐩ࡋࡓᑿഃ⩼࡜ᑿᄞࠊ୰➼ᗘ࡟◳໬ࡋࡓ஺᥋่࡜↓Ჲࡢ஺᥋่㠧ࢆ᭷ࡋࠊ඾ᆺⓗ࡞ Aonchotheca ᒓࡢ≉ᚩࢆ♧ࡋࡓࠋᅇ཰
ࡋࡓ⹸యࡢከࡃࡀኚᛶࡂࡳ࡛ࠊ㞝⹸ࠊ㞤⹸࡜ࡶ࡟ほᐹ࡛ࡁࡓᶆᮏᩘࡀᑡ࡞࠿ࡗࡓࡀࠊග
Ꮫ㢧ᚤ㙾ୗ࡛ࡢᙧែほᐹࡢ⤖ᯝࠊࡇࡢ✀ࡣ㔝⏕⫗㣗ື≀ࡢ⫶࣭ᑠ⭠࠿ࡽ᳨ฟࡉࢀࡓᮍグ㍕ Aonchotheca ᒓẟ⣽⥺⹸࡟㓞ఝࡋ࡚࠸ࡓ㸦ᡞ⏣࣭బ⸨ 2011㸧ࠋᮏ✀࡜㔝⏕⫗㣗ື≀ࡢ Aonchotheca ᒓẟ⣽⥺⹸ᶆᮏࡢィ್ࡣࡼࡃ୍⮴ࡍࡿ࡜࡜ࡶ࡟ࠊ18S rDNA ሷᇶ㓄ิࡢ☜ㄆ࠿ࡽࡶ୧⪅ࡣྠ୍✀࡛࠶ࡾࠊࡲࡓࠊᙧែᏛⓗ࡟㢮ఝࡋ⫗㣗ື≀୍࡛⯡ⓗ࡞

Aonchotheca putorii ࡜ࡣ༊ูࡉࢀࡿࡇ࡜ࡀ☜ㄆࡉࢀ࡚࠸ࡿ㸦ᡞ⏣࣭బ⸨ 2011㸧ࠋ⪃ᐹ௒ᅇࠊරᗜ┴ෆ࡜࠸࠺㝈ࡽࢀࡓᆅᇦ࡞ࡀࡽࡶࠊ⏕ᜥ⎔ቃࡢ␗࡞ࡿᆅ༊࡛ᤕ⋓ࡉࢀࡓࢽ࣍ࣥ࢖ࣀࢩࢩ࡟ࡘ࠸࡚ᐤ⏕⹸ࡢฟ⌧㢖ᗘࢆẚ㍑ࡋࡓࠋణ㤿ᆅᇦࡣἙཱྀ࡟㏆࠸ᖹ㔝㒊ࡀᗈࡀࡾࠊࡑࡇ࡟ᒣ㔝ࡀ㏕ࡗ࡚࠸ࡿࠋ୹Ἴᆅᇦࡣࠊᒣ㔝࡟ᐩࢇࡔ⎔ቃ࡛࠶ࡾࠊ㜰⚄ᆅᇦࡣ㒔ᕷ໬ࡀ㐍ࡴ୰࡟ṧࡉࢀࡓ⮬↛⎔ቃࡀ࠶ࡿ࡞࡝⎔ቃ᮲௳ࡀ␗࡞ࡗ࡚࠸ࡿࠋࡲࡓࠊᓥᕋ㒊࡛ 

රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-9

࠶ࡿῐ㊰ᓥࡢ᳨ᰝ㢌ᩘࡣ௒ᅇ 4 㢌࡜㝈ࡽࢀ࡚࠸ࡿࡀࠊᮏᕞ㒊࡟ẚ࡭࡚ᐤ⏕⹸✀ࡀ㝈ࡽࢀࡿ࡜࠸࠺ண

㏻ࡾࠊᮏᕞ㒊࡛ࡣィ 24 ✀ࡢᐤ⏕⽸⹸ࡀ☜ㄆࡉࢀࡓࡢ࡟ᑐࡋ࡚ࠊῐ㊰ᓥ࡛ࡣ 5 ✀࡟࡜࡝ࡲࡗࡓࠋࡇࡢⅬࡣࠊ௒ᚋ࡟᳨యᩘࢆቑࡍࡇ࡜࡛ࠊࡼࡾṇ☜࡞ᐇែࡀᥗࡵࡿ
ࡇ࡜ࡀᮇᚅࡉࢀࡿࠋ㣫⫱ಶయ 11 㢌ࢆ௒ᅇࡢ᳨ウ࡟ຍ࠼ࡓࡢࡣࠊ㝈ᐃࡉࢀࡓ㣫⫱⎔ቃ࡛ᐤ⏕⽸⹸࡟ࡼࡿ⃰ཌởᰁࡀ㉳ࡇࡗ࡚࠸ࡿྍ⬟ᛶࢆ᳨ドࡍࡿࡓࡵ࡛࠶ࡗࡓࠋᐇ㝿࡟ࡣࠊ᳨ฟࡉࢀࡿᐤ⏕⹸✀ࡣ⮬↛ಶయ⩌࡟ẚ࡭࡚㝈ࡽࢀࠊࡲࡓࠊᐤ⏕ᩘࡶ≉ẁ࡟ከ࠸ࢃࡅ࡛ࡣ࡞
࠿ࡗࡓࠋᤕ⋓ᚋࡢ㣫⫱࡟࠶ࡓࡗ࡚ࠊ㥑⹸๣ࡢᢞ୚➼࡝ࡢࡼ࠺࡞⟶⌮ࡀ⾜ࢃࢀࡓࡢ࠿ࡣ୙࡛᫂࠶ࡿࠋす⾲ᓥ࠶ࡿ࠸ࡣ዆⨾኱ᓥࡢࣜࣗ࢘࢟ࣗ࢘࢖ࣀࢩࢩ࡟ࡘ࠸࡚ࡣ⽸⹸඲⯡ࡢᐤ⏕≧ἣ࡟ࡘ࠸࡚ሗ࿌ࡉࢀ࡚࠸ࡿࡀ㸦Shoho & Machida 1979; ෆ⏣࡯࠿ 1984㸧ࠊࢽ࣍ࣥ࢖ࣀࢩࢩ
࡟ࡘ࠸࡚ࡣ≉ᐃ✀࡟ὀ┠ࡋࡓᐤ⏕≧ἣㄪᰝࡀ࡯࡜ࢇ࡝࡛࠶ࡾࠊᮏᕞᆅᇦࡢ㔝⏕ಶయ⩌㛫ẚ㍑࡟ࡼࡿᐤ⏕⹸┦ࡢグ㘓ࡣ᭱㏆ࡲ࡛࡞࠿ࡗࡓࠋSato et al.㸦2008㸧ࡣࠊ࿴ḷᒣ┴⏘
20 㢌ࠊරᗜ┴⏘ 9 㢌࡟ࡘ࠸᳨࡚ウࡋࠊ⥺⹸ 17 ✀ࠊ᮲⹸ 1 ✀ࠊᶓ㝸⭷➽⫗࡟⿕ᄞࡍࡿናᆺ྾⹸ Pharyngostomum cordatum ࣓ࢱࢭࣝ࢝ࣜ࢔ࢆሗ࿌ࡋ࡚࠸ࡿࠋ࿴ḷᒣ┴⏘࢖ࣀ
ࢩࢩ 1 㢌ࡢ኱⭠࠿ࡽ᪂✀グ㍕ࡉࢀࡓ Morgascaridia kugii ࡣู࡜ࡋ࡚ࡶࠊ㇜⭠⤖⠇⹸ࡢᐤ⏕ࡀࠊ࿴ḷᒣ┴⏘࢖ࣀࢩࢩ࡛ࡣ 20 㢌୰ 11 㢌㸦55%㸧࡛࠶ࡗࡓࡢ࡟ᑐࡋࠊරᗜ┴⏘࢖
ࣀࢩࢩ 9 㢌࡛ࡣ᳨ฟࡉࢀ࡞࠿ࡗࡓࠋࡓࡔࡋරᗜ┴⏘࢖ࣀࢩࢩࡢᶆᮏᩘࡀᑡ࡞࠿ࡗࡓࡓࡵࠊ
ᙜ᫬ࡣᆅᇦᕪࡢ᳨ウࡀ༑ศ࡟࡛ࡁ࡞࠿ࡗࡓࠋ௒ᅇࠊරᗜ┴ࡢణ㤿ᆅᇦ㸦17 㢌㸧ࠊ୹Ἴᆅᇦ㸦47 㢌㸧
ࠊ㜰⚄ᆅᇦ㸦53 㢌㸧࡜඘ᐇࡋࡓᶆᮏᩘ᳨࡛ᰝࢆ㐍ࡵࡿࡇ࡜࡛ࠊᆅᇦᛶࢆ᳨
ウ࡛ࡁࡓࠋ㇜⭠⤖⠇⹸ࡣࠊ
ణ㤿ᆅᇦ࡛ࡣᐤ⏕ಶయࡀ࡞ࡃࠊ
୹Ἴᆅᇦ࡛ 47 㢌୰ 1 㢌
㸦2.1%㸧ࠊ㜰⚄ᆅᇦ࡛ࡶ 53 㢌୰ 4 㢌㸦7.5%㸧࡟࡜࡝ࡲࡾࠊරᗜ┴ୗࡢ࢖ࣀࢩࢩ࡛ࡣࡈࡃ⛥࡞ᐤ⏕⹸✀࡛࠶ࡿࡇ࡜ࡀ෌☜ㄆࡉࢀࡓࠋ୍᪉ࠊ๓ᅇࡢㄪᰝ㸦Sato et al. 2008㸧ࢆྵࡵࢽ࣍ࣥ
࢖ࣀࢩࢩ࠿ࡽࡣᮍ☜ㄆ࡛࠶ࡗࡓ᭷ṑ㇜⫶⹸ࡀࠊ㜰⚄ᆅᇦ࡟ศᕸࡍࡿಶయ࡛㧗⋡㸦60.4%㸧
࡟᳨ฟࡉࢀࡓࠋྠࡌ┴ෆ࡛ࡶࠊ୹Ἴᆅᇦ 8.5%ࠊణ㤿ᆅᇦ 0%࡛࠶ࡿࡇ࡜࠿ࡽࠊᴟࡵ࡚㝈ᒁⓗ࡞ศᕸ≧ἣ࡛࠶ࡿࠋ᭱㏆࡟࡞ࡗ࡚ࠊᒣཱྀ┴࡛ᤕ⋓ࡉࢀࡓ࢖ࣀࢩࢩ࠿ࡽࡶ᳨ฟ౛ࡀ」ᩘグ㘓ࡉࢀ࡚࠾ࡾ㸦ᮍබ⾲ࢹ࣮ࢱ㸧ࠊࢽ࣍ࣥ࢖ࣀࢩࢩ⮬యࡀ㐣ཤ࠿ࡽࡶࡗ࡚࠸ࡓᐤ⏕⹸✀࡛࠶ࡿྍ⬟ᛶࡶฟ࡚ࡁࡓࡀࠊ࡝ࡢࡼ࠺࡞⤒⦋࡛ࠊᆅ⌮ⓗ㐃⥆ᛶࢆḞ࠸ࡓ⌧ᅾࡢศᕸ࡜
࡞ࡗ࡚࠸ࡿࡢ࠿ࠊරᗜ┴ࠊᒣཱྀ┴௨እࡢᆅᇦ࡛ࡶㄪᰝࡀ㐍ࡴࡇ࡜࡟ࡼࡗ࡚ࠊࡑࡢ≉᭷࡞ศᕸࡢ⣒ཱྀࡀぢࡘ࠿ࡿࡇ࡜ࢆᮇᚅࡋࡓ࠸ࠋ
᭷ṑ㇜⫶⹸࡟ຍ࠼ࠊ௒ᅇࠊᅜෆࡢ࢖ࣀࢩࢩ࠿ࡽึࡵ࡚⨾㯇㣗㐨⹸ࡀ᳨ฟࡉࢀࡓࠋ࢖ࣛ
ࣥ⏘࢖ࣀࢩࢩ࡛ࡣ㧗⋡࡞ᐤ⏕ࡀሗ࿌ࡉࢀ㸦Eslami & Farsad-Hamdl 1992㸧ࠊࡲࡓࠊྜ
⾗ᅜࡢᐙ␆࡜ࡋ࡚㣫㣴ࡉࢀࡿࣈࢱࡸ㔝⏕໬ࣈࢱ࡛ࡶ㧗࠸ᐤ⏕⋡ࡀሗ࿌ࡉࢀ࡚࠸ࡿࡇ࡜
࠿ࡽ㸦Zinter & Migaki 1970; Smith et al. 1982㸧ࠊື≀✀࡜ࡋ࡚㧗࠸ឤᰁឤཷᛶࢆࡶࡘ
ࡇ࡜ࡀ᥎ࡉࢀࡿࠋࡋ࠿ࡋ࡞ࡀࡽࠊ௒ᅇࡢㄪᰝ࡛ࡣᮏᕞ㒊 117 㢌୰ 5 㢌㸦4.3%㸧࡟࡜࡝ࡲࡾࠊῐ㊰ᓥ⏘ 4 㢌࠿ࡽࡢ᳨ฟࡣ࡞࠿ࡗࡓࠋྜ⾗ᅜࡢ㔝⏕ࣈࢱࢆㄪ࡭ࡓ Coombs &Springer㸦1974㸧ࡣࠊ⯉࠿ࡽࡢ᳨ฟ⋡ࡀ㧗ࡃ㸦6 / 9㸧ࠊ㣗㐨࠿ࡽࡢ᳨ฟ⋡ࡣప࠸㸦1 / 10㸧ࡇ࡜ࢆሗ࿌ࡋ࡚࠸ࡿࠋ௒ᅇࡢ᳨ᰝࡣ㣗㐨⢓⭷࡟㝈ࡽࢀ࡚࠸ࡓࡇ࡜࠿ࡽࠊᐇ㝿ࡢᐤ⏕⋡ࡣ 

රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-9௒ᅇࡢ⤖ᯝ௨ୖ࡟㧗࠸ྍ⬟ᛶࡶ༑ศ࡟࠶ࡿࠋ࠶ࡿ࠸ࡣࠊ௒ᅇࡢ࢖ࣀࢩࢩᐤ⏕⹸ㄪᰝ࡜ྠ
᫬ᮇ࡟ᐇ᪋ࡋࡓ࣍ࣥࢩࣗ࢘ࢪ࢝ࡢㄪᰝ࡛ࡶಶయ⩌ࡶࡋࡃࡣ⏕ᜥᆅᇦ࡟ࡼࡾឤᰁ⋡࡟኱ࡁ࡞㐪࠸ࡀ࠶ࡗࡓࡇ࡜࠿ࡽ㸦బ⸨ 2009; Makouloutou et al. 2013㸧ࠊྠᵝࡢせᅉ࡛ࠊ௒ᅇ᳨ᰝࡋࡓ࢖ࣀࢩࢩ࡛ࡢឤᰁ⋡ࡣᐇ㝿࡟ప࠿ࡗࡓྍ⬟ᛶࡶ࠶ࡿࠋ⤖ᯝ࡛グࡋࡓࡼ࠺࡟ࠊ
Makouloutou et al.㸦2013㸧࡟࠾࠸࡚ศᏊ⣔⤫Ꮫⓗゎᯒࡢヲ⣽ࢆሗ࿌ࡋ࡚࠸ࡿࠋᅜෆࡢ
⨾㯇㣗㐨⹸ࡣ㣗⣅ᛶ⏥⹸ࢆ୰㛫ᐟ୺࡜ࡋ࡚ࠊ㔝⏕ື≀࡛࠶ࡿࢩ࢝ࡸ࢖ࣀࢩࢩࢆ⤊ᐟ୺࡜ࡍࡿ᳃ᯘᆺࡢ⏕ά⎔ࢆྲྀࡿ⣔⤫࡜ࠊ࢘ࢩࢆ⤊ᐟ୺࡜ࡍࡿᐙ␆ᆺࡢ⏕ά⎔ࢆྲྀࡿ⣔⤫ࡀ࠶ࡿࡇ࡜ࡀุ᫂ࡋ࡚ࡁ࡚࠸ࡿ㸦బ⸨ 2009㸧ࠋරᗜ┴ୗ࡛ࡢ㇜⭈⹸ࡢឤᰁ≧ἣ࡟ࡘ࠸࡚ࡣࠊ๓ሗ㸦Sato et al. 2008㸧࡛ࡣࡓࡲࡓࡲ
ឤᰁࡋ࡚࠸ࡓ࢖ࣀࢩࢩ 1 㢌ࢆ☜ㄆࡋࡓࡀࠊឤᰁ⋡࡜ࡋ࡚࡝ࡢࡼ࠺࡞≧ἣ࡟࠶ࡿࡢ࠿ࡣ᫂
☜࡟࡛ࡁ࡚࠸࡞࠿ࡗࡓࠋ௒ᅇࡶᐊෆ࡛ࡢ᳨ᰝࡣᐇ᪋ࡉࢀ࡚࠸࡞࠸ࡀࠊゎయ⌧ሙ࡛ࡢ⫗║࣭ゐデ࡛ࡢ☜ㄆ࡛ឤᰁࡢ᭷↓ࢆグ㘓ࡋࠊឤᰁಶయ࠿ࡽ⹸యࡢ཰㞟ࢆ⾜ࡗࡓࠋࡑࡢ⤖ᯝࠊ⾲㸰࡟♧ࡍࡼ࠺࡟୹Ἴᆅᇦ࡛ࡣ 47 㢌୰ 9 㢌㸦19.1%㸧ࡢᐤ⏕ࡀ࠶ࡗࡓࡶࡢࡢࠊ┴ෆࡢ௚ᆅᇦ࡟⏕ᜥࡍࡿಶయ⩌࡛ࡣ࡯࡜ࢇ࡝ឤᰁࡀ࡞࠿ࡗࡓࠋ࡞࠾ࠊ࿴ḷᒣ┴⏘ࡢ࢖ࣀࢩࢩ࡛
ࡣࡈࡃ୍⯡ⓗ࡞ᐤ⏕⹸࡛࠶ࡗࡓ㸦ឤᰁ⋡ 55%: Sato et al. 2008㸧ࠋᮏ✀ᗂ⹸ࡣయෆ࡟ධ
ࡗ࡚ 3 ᅇ⬺⓶ࡋࡓᚋ࡟⫢⮚࡟ධࡗ࡚ࠊࡑࡢᐇ㉁ෆ࡛Ⓨ⫱ࡋࠊ෌ࡧ⭡⭍ෆ࡟ฟ࡚࠿ࡽ᭱⤊ᐤ⏕㒊఩࡛࠶ࡿ⭈⮚ࡶࡋࡃࡣࡑࡢ࿘ᅖࡢ⬡⫫⤌⧊࡟⛣ືࡍࡿࠋ࿴ḷᒣ┴⏘࢖ࣀࢩࢩ࡛ࡣࠊ
⫢⮚࡟⹸㐨࡟㉳ᅉࡍࡿ⤖ྜ⤌⧊ቑ⏕ࡸ⭡⭷⅖࡜㛵㐃ࡋࡓ⥺⥔⣲ᛶ⫢⿕⭷⅖ࡀࡋࡤࡋࡤほᐹࡉࢀࡓࡀࠊරᗜ┴⏘࢖ࣀࢩࢩ࡟࠾࠸࡚ࡣࠊ㇜⭈⹸ᐤ⏕࡟క࠺⑓ኚࡣ࡯࡜ࢇ࡝☜ㄆࡉࢀ࡚࠸࡞࠸ࠋప࠸ឤᰁ⋡ࢆ཯ᫎࡋࡓゎ๗⫗║ᡤぢ࡛࠶ࡿ࡜⪃࠼ࡿ࡜ࠊ௒ᅇࡢᐤ⏕⹸ᅇ཰≧ἣ࡜ࡼࡃ➢ྜࡍࡿࠋ࢖ࣀࢩࢩࡢゎ๗࡟࠾࠸࡚ࠊ⫶ቨ࡟✸Ꮝࡋᐤ⏕㒊఩ࢆ⫧ཌࡉࡏࡿࢻࣟࣞࢫ㢡ཱྀ⹸ࡣ┠❧ࡘࠋᮏ✀ࡣࣈࢱ࣭࢖ࣀࢩࢩࢆᅛ᭷ᐟ୺࡜ࡋ࡚⫶ቨ࡟ᡂ⹸ࡀᐤ⏕ࡋࠊ➨ 1 ୰㛫ᐟ୺ࡣࢣࣥ
࣑ࢪࣥࢥࠊ➨ 2 ୰㛫ᐟ୺ࡣ୧⏕㢮㸦ࢧࣥࢩࣙ࢘࢘࢜ࠊ࢚࢝ࣝࠊ࢖ࣔࣜ➼㸧ࡸ㨶㢮ࠊᚅᶵᐟ୺ࡀ࣊ࣅ㢮࡛࠶ࡿ㸦ᐑᓮ࣭⸨ 1988㸧ࠋᐑᓮ┴ࢆ୰ᚰ࡟஑ᕞᆅ᪉࡛ࡣ῱ὶ㨶࣐࣓ࣖᩱ⌮ࢆཎᅉ࡜ࡋࡓே࡬ࡢឤᰁ౛㸦⓶⭵∐⾜⑕㸧ࡀሗ࿌ࡉࢀ࡚࠸ࡿ㸦㉥⩚ 1999㸧ࠋ௒ᅇࠊරᗜ┴ࡢᮏᕞ㒊࡛ࡣᆅᇦࢆၥࢃࡎ࡟㧗࠸ឤᰁ⋡ࡀ☜ㄆࡉࢀࡓࡇ࡜࠿ࡽࠊ࢖ࣀࢩࢩࡢ⏕ᜥᆅ࡟ࡣᗈࡃࢻࣟࣞࢫ㢡ཱྀ⹸ࡀศᕸࡋ࡚࠸ࡿࡇ࡜ࡀ᫂ࡽ࠿࡜࡞ࡗࡓࠋ➨ 2 ୰㛫ᐟ୺ࡸᚅᶵ
ᐟ୺ࢆႚ㣗ࡍࡿᶵ఍ࡀ࠶ࡿሙྜ࡟ࡣࠊࡑࡢຍ⇕ฎ⌮ࡀ୙༑ศ࡛࠶ࡗࡓሙྜࠊ⓶⭵∐⾜⑕ࡀᘬࡁ㉳ࡇࡉࢀࡿ༴㝤ᛶࡀ࠶ࡿࡇ࡜࡟␃ពࡍࡿᚲせᛶࡀ࠶ࡿࠋ
ᅜෆ㎰ᐙ࡛ࡢᐙ␆࡜ࡋ࡚ࡢࣈࢱࡢ㣫㣴ᙧែࡣ㏆ᖺ኱ࡁࡃኚ໬ࡋࠊᮏ◊✲࡛ᢅࡗࡓᐤ⏕⽸⹸ࡢከࡃࡀ㣴㇜⌧ሙ࡛┳㐣࡛ࡁ࡞࠸ၥ㢟࡛࠶ࡗࡓ㐣ཤࡀ㐲࠸᫇ヰ࡟࡞ࡗ࡚࠸ࡿࠋ⟶⌮ࡉࢀࡓࢥࣥࢡ࣮ࣜࢺᗋࡢ㣫⫱⯋࡛ࡢ㣫㣴ࡣࠊࡑࡢ⏕ά⎔⥔ᣢ࡟୰㛫ᐟ୺ࢆᚲせ࡜ࡍࡿᐤ⏕⽸⹸ࡸእ⏺࡛ࡢ⹸༸Ⓨ⫱ࡀᚲせ࡞ᐤ⏕⽸⹸ࡢឤᰁ㜵Ṇ⟇࡜࡞ࡗ࡚ࡁࡓࠋ࢜࢞ࢡࢬⓎ㓝㇜⯋ࡢᬑཬ࡟ࡼࡾ㇜㠴⹸ࡀ⮴࿨ⓗ࡞ឤᰁࢆᘬࡁ㉳ࡇࡋࡓࡼ࠺࡟㸦ᖹ 1989㸧ࠊ≉ᐃ✀࡟࠾࠸࡚ࡣேᕤ⎔ቃࡀ㏫࡟ᐤ⏕⽸⹸࡟ዲ㐺࡞⏕⫱࣭ఏ᧛ᶵ఍ࢆ๰ฟࡍࡿࡇ࡜ࡶ࠶ࡿࠋ㣗⫗࡜ࡋ࡚ࡢ௜ຍ౯್ࢆࡘࡅࡿࡓࡵ࡟ࠊ⮬↛ࡢ୰࡛⫱ࡗࡓ㇜ࡢ㣫⫱㸦ᨺ㣴㸧ࡶᅜෆ࡛ヨࡳࡽ 

රᗜ࣡࢖ࣝࢻࣛ࢖ࣇࣔࣀࢢࣛࣇ 6-9

ࢀ࡚࠸ࡿࠋࡋ࠿ࡋ㇜࡛ࡣ㐣ཤࡢ⑓Ẽ࡛࠶ࡗࡓࡣࡎࡢᐤ⏕⽸⹸⑕ࡣࠊ㔝⏕࢖ࣀࢩࢩ࡛ࡣࡋ
ࡗ࠿ࡾ࡜ᅜෆ࡛ࡶ⏕ά⎔ࡀ⥔ᣢࡉࢀ࡚࠸ࡿࠋࡑࡢࡓࡵ࢖ࣀࢩࢩࢆಖ⹸ᐟ୺࡜ࡋ࡚༑ศ࡟
⪃៖ࡋࠊᐙ␆㣫⫱࢚ࣜ࢔࡜࢖ࣀࢩࢩ⏕ᜥᇦ࡜ࡢ᥋Ⅼࡀ࡞࠸ࡼ࠺࡟ὀពࡍࡿࡇ࡜ࡶᛀࢀ࡚
ࡣ࡞ࡽ࡞࠸ࠋㅰ㎡ᮏ◊✲ࢆ㐙⾜ࡍࡿ࡟࠶ࡓࡾࠊࡈ༠ຊ࠸ࡓࡔ࠸ࡓ⊟཭఍ࡢⓙᵝ࡟ཌࡃᚚ♩⏦ࡋୖࡆࡿࠋ
ࡲࡓࠊᮏ◊✲ࡢ୍㒊ࡣ JSPS ⛉◊㈝ 22580349ࠊ25450478 ࡢຓᡂࢆཷࡅࡓࠋ

ᘬ⏝ᩥ⊩㉥⩚ၨᰤ. 1999. 㢡ཱྀ⹸࣭๛Ჲ㢡ཱྀ⹸࣭ࢻࣟࣞࢫ㢡ཱྀ⹸. ᪥ᮏ࡟࠾ࡅࡿᐤ⏕⹸Ꮫࡢ◊✲ 7.
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