AMERICAN MUSEUM NOVITATES
Number 491

Published by
THU AMURCAN MUisUm 0P NATURAL HISTORY
New York City

Sept. 24, 1931

59.57,71 T: 07 (74.7)

NEW OR LITTLE-KNOWN TIPULIDAE IN THE AMERICANMUSEUM OF NATURAL HISTORY (DIPTERA).-I
BY CHARLES P. ALEXANDER'

The crane-flies discussed in the present report were collected by Mr.
G. H. H. Tate on Mount Roraima, Brazil, and on Mount Duida, Venezuela. The material is of especial interest because of the geologicalstructure and isolation of these two mountains. The tipulid fauna of thetwo localities is distinct, but this may prove to be a matter of insufficientcollecting, since certain of the species are wide-spread in the Neotropics.
Most of the species described as new will probably be found to be restricted to the mountains in question. I am greatly indebted to Mr.
Curran for the opportunity of studying this very interesting series of
crane-flies. All types and uniques are preserved in the collection of The
American Museum of Natural History.
The conditions encountered by Mr. Tate in his ascent of Roraima
have been delightfully described in his article, 'Through Brazil to theSummit of Mount Roraima,' National Geographic Magazine, November,
1930, LVIII, pp. 585-605. The following paragraph is of interest.
"With nightfall and the lighting of the gasoline lantern came a host ofnight moths, daddy longlegs, and flying creatures that, despite oursearch, we seldom found by day."Besides the species recorded at this time, the series included a few
additional species in condition too poor to be determined.
The collection includes the following Tipulidae.
MT. RORAIMA, BRAZIL
Tipula arecuna, new species
Brachypremna breviventris (Wiedemann)
Limonia (Limonia) roraimme, new species
Limonia (Neolimnobia) diva (Schiner)
Limwnia (Dicranomyia) dolerosa, newspecies

Limonia (Geranomyia) tatei, new species
Limonia (Geranomyia) arecuna, newspeciesLimnophila roraima, new species
Limnophila roraimicola, new species'Massachusetts State College Amherst.

MT. DUIDA, VENEZUELA
Ozodicera duidensis, new species
Tipula duide, new species
Limonia (Rhipidia) conica turrifera, newsubspecies
Limonia (Geranomyia) rubiginosa, newspecies

Helius (Helius) albitarsis Osten Sacken
Limnophila kaieturana Alexander
Atarba (Atarba) unilateralis, new species
Teucholabis (Teucholabis) duidensis, newspecies

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AMERICAN MUSEUM NOVITATES[NO. 491

TIPuLiNS
Ozodicera (Ozodicera) duidensis, new species
Figures 1, 11 and 12
General coloration yellowish gray, the praescutum with four dark brown stripes,
the intermediate pair only narrowly separated; flagellar branches short; wings
brownish gray; male hypopygium with the apex of the sternite produced into aslender roughened point immediately dorsad of the point of insertion of the dististyles;
inner dististyle narrowed to a long chitinized point, with a smaller sublateral spine.
MALE.-Length, about 20-23 mm.; wing, 17.5-20 mm.; antenna, about 4.34.4 mm.
FEMALE.-Length, about 22 mm.; wing, 18 mm.
Frontal prolongation of head elongate, dark brown; nasus very small but distinct. Antenne with the basal segment dark brown, the second obscure yellow;
flagellum entirely black; flagellar branches (male) short, the longest (flagellar segment five) about two-thirds the segment; branches of flagellar segments two to sevensmall, triangular in outline; terminal flagellar segments elongate, slightly exceeding
the remainder of flagellum. Head brownish gray, somewhat clearer gray adjoining
the eyes; anterior vertex narrow, more buffy; front similarly colored, variegatedwith brown.
Mesonotal prmscutum yellowish gray, with four dark brown stripes, the intermediate pair only narrowly and vaguely divided; pseudosutural fovee small, blackened, nearly marginal in position; scutum gray, the centers of the lobes darker;
scutellum gray, with a vague capillary brown median vitta; postnotal mediotergite
gray. Pleura gray, indistinctly variegated with brown on the anepisternum, ventral
sternopleurite and dorsal pleurotergite. Halteres elongate, dark brown, the knobs
slightly paler, more yellowish brown. Legs with the coxe pruinose; trochantersreddish brown; femora and tibiae reddish brown, the tips narrowly dark brown or
black; tarsi reddish brown, the outer segments blackened. Wings (Fig. 1) with abrownish-ray tinge, the costal region and stigma more yellowish brown; a vague
pale brown seam on the anterior cord; veins brown. Venation: distal half of Ri +2
pale, subobliterated; cell MI broadly wseile.
Abdominal tergites reddish yellow, narrowly blackened laterally, the dorsoAiedian region dark brown, on the outer segments the areas becoming more extensive
and finally confluent; caudal margins of segments restrictedly pale yellow; sternites
obscure yellow, the extreme caudal margin yellow. Male hypopygium (Fig. 11)
with the sternite and basistyle extended far beyond the level of the tergite, the dististyles at extreme tip. Ninth tergite (Fig. 12) with a V-shaped median notch, thebroad lateral lobes feeblyemarginate, the median lobule a little more produced caudad
than the lateral shoulders. Apex of the fused basistyle and sternopleurite produced
into a slender blackened point, its surface set with abundant tubercles. Dististyles
about as figured; inner style narrowed to a chitinized point, with a smaller, more
obtuse subterminal point.

HABITAT.-Venezuela.
HOLoTYPE, c, Mount Duida, November 30, 1928 (G. H. H. Tate); Coll. No.
412. ALLOTOPOTYPE, 9, with the type. PARATOPOTYPES, two fragmentary specimens,
sex unknown, one collected with the types, the other on January 11, 1929; Coll. No.
651.

1931]
1NEW OR LITTLE-KNOWN TIPULIDE

3

Ozodicera (Ozodicera) duidensis is best defined by the combination ofgrayish coloration, short antennal pectinations, and the structure of themale hypopygium. Its closest ally would appear to be 0. (O.) bispinifera
Alexander (Brazil).
Tipula arecuna, new species
Figures 13 and 14
General coloration of thorax black, including three polished brownish-black
stripes on an obscure yellow ground; scutal lobes blackened; scutellum yellowish
testaceous, the parascutella abruptly blackened; thoracic pleura chiefly blackened;
antenna short, the basal fl4gellar segments bicolorous; wings yellow, with a heavy
tesselated brown and grayish-brown pattern; male hypopygium with the tergite
thin, feebly sclerotized, with a deep U-shaped median notch, the lateral lobes broad,obtusely rounded at tips; no appendages on the eighth and ninth sternites.
MALE.-Length, about 14 mm.; wing, 17.5 mm.
Frontal prolongation of head dark brown; nasus slender; palpi with the basal
segment obscure yellow, the second and third dark brown, more whitish at the incisures; outer segment black. Antennae short; scapal segments honey-yellow;
flagellum obscure yellow, the segments ringed just beyond base with brown, this ring
including the four verticils which exceed the segments in length; oufter flagellar segments more uniformly darkened. Head brown, the front more yellowish brown.
Posterior pronotum obscure yellow medially, passing into dark brown on thesides. Mesonotal praescutum with the ground-color polished obscure yellow, with-three polished dark brown or brownish-black stripes, the median stripe reaching thesuture, the broad lateral stripes including the entire lateral margins of the sclerite
behind the punctiform psuedosutural fovem; scutal lobes almost entirely blackened,
the median region very restrictedly obscure yellow; scutellum yellowish testaceous,the parascutella abruptly black; postnotal mediotergite polished black, vaguelybrightened along the lateral margins. Pleura chiefly brownish black, with vague
obscure yellow areas on the posterior sternopleurite and ventral pteropleurite; dorsopleural region buffy-yellow. Halteres obscure yellow. Legs with the fore and middle
coxe brownish black, the posterior coxs paler, more yellowish brown; trochanters
yellow; remainder of legs broken. Wings with the ground-color cream-yellow, cell Sc
bright yellow, cee C chiefly infumed; a very heavy brown and grayish brown tesselated pattern that is almost like that found in bigotiana, including four solid browndarkenings in cell Sc, these being narrower than the interspaces; veins yellow, infuscated in the clouded areas; obliterative areas extensive. Venation: Rs long;
R1 +2 entire; petiole of cell Ml a little longer than its petiole.
Abdominal tergites brownish yellow, margined laterally with brownish black,the bases of the segments somewhat darker than the apical portions; sternites andhypopygium more uniform yellow. Male hypopygium (Fig. 13) with the ninth tergite(Fig. 14) large, depressed, entirely pale and but feebly sclerotized, the caudal margin
with a deep U-shaped notch, the sublateral lobes thus formed broadly and obtusely
rounded at apex. Tergite and sternite fused into a ring. Basistyle complete. Innerdististyle as shown; outer dististyle of unique type apparently destroyed, not visible.
Eighth and ninth sternites without appendages.
HABrIAT.-Brazil.

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AMERICAN MUSEUM NOVITATES[No. 491

HOLOTYPE, c, Mount Roraima, summit, altitude 8600 feet, November, 1927
(G. H. H. Tate); Coll. No. 26.

The specific name, arecuna, is that of the tribe of Indians living
nearest the mountain. In its wing-pattern, Tipula arecuna much resembles species like T. bigotiana Alexander and T. mocoa Alexander,
differing in the coloration of the body, especially of the thorax. The
structure of the male hypopygium would seem to exclude the present
species from the glaphyroptera group, where, at first sight, it would appear
to belong.
Tipula duida, new species
Figure 15
Belongs to the glaphyroptera group; mesonotum light golden-yellow, with threedark chocolate-brown stripes; antennm short, bicolorous; femora blackened, with aconspicuous yellow subterminal ring; wings creamy-yellow, with a heavy brownpattern; male hypopygium with the median appendage of the eighth sternite small,
simple.
MAI.E.-Length, about 14-15 mm.; wing, 16.5-17.5 mm.
Frontal prolongation of head obscure yellow dorsally, passing into black beneath;nasus long and slender; palpi with the basal two segments light brown, the remainder
brownish black, the apex of the third pale. Antennae (male) short, if bent backward,
not attaining the wing-root; scape yellow, the base of the first segment darkened;flagellar segments bicolorous, blackened at base, the longer apical portion yellow;
outer segments more uniformly darkened. Head brownish yellow, more suffusedwith brown on anterior vertex and on gense, with an additional capillary dark brownvitta, the ground-color most evident on either side of the posterior vertex.
Pronotum dark brown medially, paler sublaterally. Mesonotal prsscutum lightgolden-yellow, with three dark chocolate-brown stripes that are nearly confluent
behind; scutum brownish gray, the centers of the lobes dark brown; scutellum gray,broadly dark brown medially; postnotal mediotergite yellowish gray, with a blackened 1-shaped marking. Pleura brown, the ventral pleurotergite more blackened;
dorso-pleural region buffy brown. Halteres long, pale brown, the base of stem
narrowly brightened, the knobs, in cases, brownish yellow. Legs with the coxin andtrochanters chiefly brown; femora dark brown to black outwardly, more yellowish atbase, with a narrow but conspicuous yellow subterminal ring, the extreme tip of thesegment paling from black to brown or yellow; tibiae light brown, the tips narrowly
darkened; tarsi light brown, the outer segments blackened. Wings with the groundcolor creamy yellow, the costal margin more saturated; a heavy brown pattern,including four solid darker brown areas in cell Sc; a heavy tesselate brown pattern,covering the entire wing-disk, being more extensive than the pale ground-color except
in cells R and M; in outer end of cell M a ring-like area, gray, margined with brown;veins dark, paler in the yellowish areas. Venation: R1 +2 entire, the distal two-thirds
pale yellow; petiole of cell M1 from one-third to two-thirds longer than m; fork of
M3+4 at near one-third the length of cell 1St M2.
Abdominal tergites brown, more blackened laterally, before caudal margin on
either side with a brighter yellow area; subterminal segments more uniformly dark

1931]NEW OR LITTLE-KNOWN TIPULID55

brown; sternites yellow, the caudal margins of the segments narrowly dark brown.
Male hypopygium (Fig. 15) small, the ninth tergite extensive, narrowed outwardly,
terminating in two small spinous points that are separated by a small U-shaped
median notch; when viewed from the side, these points appear as small flattened
blades; lateral margins of tergite fringed with setwe. Outer dististyle a slender,
weakly clavate lobe. Appendage of eighth sternite a small, simple, finger-like lobe
that is about three times as long as wide at base.
HABITAT.-Venezuela.
HOLOTYPE, c, Mount Duida, January 11, 1929 (G. H. H. Tate); Coll. No.
552. PARATOPOTYPE, c .

Tipula duida3 is best distinguished from the other members of theglaphyroptera group with tesselated wings by the combination of characters summarized above.
Brachypremna breviventris (Wiedemann)
Tipula breviventris WIEDEMANN, 1821, 'Dipt. Exot.,' I, p. 43.
Tipula breviventris WIEDEMANN, 1828, 'Aussereur. zweifl. Ins.,' I, p. 47.
Brachypremna breviventris OSTEN SACKEN, 1886, Berliner Ent. Zeitschr., XXX,
p. 161.
Brachypremna breviventris ALEXANDER, 1912, Journ. N. Y. Ent. Soc., XX, pp.
230-231.

Slopes of Mount Roraima, Brazil, altitude 4000 feet, November 5,
1927 (Paulo). This fly has a wide distribution in northern SouthAmerica.
LIMONIINZ

Limonia (Limonia) roraimm, new species
Figure 2
General coloration polished yellow; rostrum, palpi, and antennae black; head
blackish gray; halteres yellow; femora yellowish brown, darkest at base, with aconspicuous, obscure yellow, nearly terminal ring; wings yellow, more saturated in
the costal and apical portions, with conspicuous dark brown seams and paler brownwashes; basal section of R4+5 very short.
Sex(?).-Wing, 9-10.4 mm.
Rostrum and palpi black. Antennwe black throughout; basal flagellar segments
subtriangular to cordiform, with very short, thick, apical pedicels; verticils unilaterally arranged, not exceeding the segments. Head blackish gray; anterior vertex
reduced to a linear strip.
Pronotum dark brown medially, paling to yellow on sides. Mesonotal praescutum chiefly polished yellow, the interspaces in front a little darker; scutellum
medially and the cephalic portion of the postnotal mediotergite more darkened.
Pleura yellow. Halteres yellow. Legs with the coxe and trochanters yellow; femora
yellowish brown, darkest at extreme base, with a conspicuous obscure yellow, nearlyterminal ring; tibiie brownish yellow, the tips narrowly darkened; outer tarsal
segments blackened; claws with about three teeth, the outermost longest. Wings(Fig. 2) with a bright yellow ground-color, the prearcular, costal, and outer radial

I

3

If

I

11

7

8

9

10

77'...--.iFig. 1. Ozodicera (Ozodicera) duidensis, new species; wing.
" 2. Limonia (Limonia) roraimme, new species; wing.
3. Limonia (Dicranomyia) dolerosa, new species.; wing.
" 4. Limonia (Geranomyia) tatei, new species; wing.
5. Limonia (Geranomyia) arecuna, new species; wing.
6. Limonia (Geranomyia) rubiginosa, new species; wing.
7. Limnophila roraima, new species; wing.
8. Limnophila roraimicola, new species; wing.
9. Atarba (Atarba) unilateralis, new species; wing.
10. Teucholabis (Teucholabis) duidensis, new species; wing.

g~~

~

~ idi
~~~~~~~~~
b

9!
ofgt

12

14

9s

8s.

~~~~~~15

13
8s

11:.0.\s.*
SI

16
17
18
Fig. 11. Ozodicera (Ozodicera) duidensis, new species; male hypopygium, lateral
aspect.
12. Ozodicera (Ozodicera) duidensis, new species; male hypopygium, ninth
tergite, dorsal aspect.
13. Tipula arecuna, new species; male hypopygium, lateral aspect.
14. Tipula arecuna, new species; male hypopygium, ninth tergite, dorsal
aspect.
15. Tipula duid., new species; male hypopygium, details.
16. Limonia (Geranomyia) tatei, new species; male hypopygium.
17. Limonia (Geranomyia) arecuna, new species;. male hypopygium.
18. Atarba (Atarba) unilateralis, new species; male hypopygium.

[Symbols: b=basistyle; g=gonapophysis; id=inner dististyle; od=outerdististyle; s=sternite; t=tergite; vd=ventral dististyle.]
7

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AMERICAN MUSEUM NOVITATES[No. 491

regions more saturated yellow; a restricted dark brown pattern, including narrow
seams at origin of Rs; fork of Sc; R2 and R1 +2; along cord and outer end of cell 1st
M2; and as narrow marginal clouds at ends of all longitudinal veins; paler brownwashes in base of cell R; at stigma, the latter sending a conspicuous darker brown
extension basad over the anterior cord; posterior cells of wing, from M3 to 2nd A with
marginal brown washes, leaving the region of the vein clear yellow; veins yellow,
darker in the clouded areas. Venation: Scl ending about opposite two-thirds tothree-fourths the length of Rs, Sc2 near its tip; Rs angulated at origin; basal sectionof R4+5 very short, arcuated; cell 1st M2 relatively large, subequal in length to vein
Ml +2 beyond it; basal section of Ml long, arcuated; m-cu at or before the fork ofM, about one-half longer than the distal section of Cul.
Abdominal tergites obscure yellow, the organ broken beyond segment two.
HABITAT.-Brazil.
HOLOTYPE, sex(?), Mount Roraima, summit, altitude 8600 feet, November, 1927
(G. H. H. Tate). PARATOPOTYPE, Sex?

Limonia (Limonia) roraimae is most closely allied to L. (L.) sancta3
martze Alexander (Colombia) in the general features of wing-pattern,differing in the details of venation and coloration, such as the darkened
tips of the femora in the latter species.
Limonia (Neolimnobia) diva (Schiner)
Limnobia diva SCHINER, 1868, 'Novara Reise,' Diptera, p. 46.
Dicranomyia (Neolimnobia) diva ALEXANDER, 1928, Dept. Sci. and Agr. Jamaica,
Ent. Bull., IV, p. 20.

Three specimens, Mount Roraima, Brazil, summit, altitude 8600
feet, November, 1927 (G. H. H. Tate). The species has a wide range in
the northern Neotropical region.
Limonia (Rhipidia) conica turrifera, subspecies
Differs from typical conica Alexander (Trans. American Ent. Soc.,
1914, XL, pp. 227-228) of British Guiana, only in the more restricteddark wing-pattern. The dark areas of the membrane are small, clearly
delimited, and, for the most part, entirely surrounded by the pale yellow
ground-color. In the typical form, the darkened subcostal areas aretriangular in outline, widest at caudal ends and merging into the dark
ground-color of the medial and posterior regions of the wing.
HABITAT.-Venezuela.
HOLOTYPE, 9, Mount Duida, November 18, 1928 (G. H. H. Tate); Coll. No.
338. PARATOPOTYPE, a broken 9, November 4, 1928; Coll. No. 149.

Limonia (Dicranomyia) dolerosa, new species
Figure 3
General coloration black, including the disk of the mesonotal prescutum;
antennae black throughout; thoracic pleura brownish black, traversed by a narrow1931]NEW OR LITTLE-KNOWN TIPULIE99

transverse pale line on the pteropleurite; legs chiefly black, the extreme femoral tips
vaguely paler, especially on their flexor surface; wings with a strong blackish tinge,
the stigma and a diffuse seam along cord somewhat darker; free tip of Sc2 about its
own length before R2.
FEMALE.-Length, about 10-11 mm.; wing, 10.3-10.5 mm.
Rostrum relatively elongate, exceeding one-half the remainder of head, black;
palpi black. Antennre black throughout; flagellar segments long-oval, the verticils
shorter than the segments; terminal segment longer than the penultimate. Headblack, very sparsely pruinose; anterior vertex a little wider than the diameter of thescape.
Pronotum black, paler laterally. Mesonotal prsescutum chiefly covered by threenearly confluent black stripes, the humeral and lateral portions obscurely brightened;scutal lobes blackened, the median area pale; scutellum brownish testaceous; posts
notal mediotergite black, restrictedly paler on the anterolateral portions. Pleura
chiefly brownish black, interrupted by a narrow pale transverse line extending fromthe wing-root, crossing the pteropleurite to the meral region. Halteres dark brown,
the base of the stem restrictedly pale yellow. Legs with the fore cox2e black, the remaining coxse and all trochanters obscure yellow; femora dark brown, obscurely
brightened at extreme base and tip, the latter more especially so on the flexor surface;remainder of legs black. Wings (Fig. 3) with a strong blackish tinge, the oval stigma
darker; an ill-defined darkened cloud on the cord; paler longitudinal streaks incertain cells, more especially R, M, and M4, crossing the cord and outer end of cell
1st M2 and forming bullate areas; veins dark brown. Venation: Sci ending opposite
the origin of Rs, Sc2 some distance from its tip, Sci alone being more than one-half
Rs; free tip of Sc2 some distance before the level of R2; n-cu beyond the fork of M
about one-third longer than the distal section of Cul; cell 2nd A wide.
Abdominal tergites black, the basal sternites obscure yellow, the outer segments
more blackened, especially laterally. Ovipositor with the tergal valves slender,
relatively short and straight, narrowly blackened at bases; sternal valves blackened
at bases, their tips reddish horn-color.

HABITAT.-Brazil.
HOLOTYPE, 9, Mount Roraima, summit, altitude 8600 feet, November, 1927
(G. H. H. Tate). PARATOPOTYPE, 9.

In its strongly infumed wings, Limonia (Dicranomyia) dolerosa ismost similar to L. (D.) dampfi (Alexander), of Mexico, and L. (D.)
selkirki (Alexander), of the Islands of Juan Fernandez, differing in thedetails of coloration and venation, especially the wide second anal cell.
Limonia (Geranomyia) tatei, new species
Figures 4 and 16
General coloration reddish gray, the praescutum with five narrow liver-brown
stripes, the outermost pair on the lateral margins; knobs of halteres darkened; femorawith two dark brown and two yellow annuli; wings cream-yellow, with a very heavy
brown pattern, including the broad apex; Sci ending at near one-third the l,ength ofRe; male hypopygium with the rostral spines of the ventral dististyle arising from acommon tubercle.

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AMERICAN MUSEUM NOVITATES[No. 491

MALE.-Length, excluding rostrum, about 5.5 mm.; wing, 7.4 mm.; rostrum,
about 3.5 mm.
FEMALE.-Length, excluding rostrum, about 6 mm.; wing, 7.2 mm.; rostrum,
about 3.4 mm.
Rostrum relatively elongate, exceeding one-half the length of body, black, thedistal third of labial palpi paling to brown; maxillary palpi black. Antenne short,
black; flagellar segments subeylindrical, without pedicels. Head dark gray, theposterior vertex with a blackish area on either side of the median line.
Mesonotal prascutum with the ground-color reddish, sparsely pruinose, especiallylaterally behind, with five dark liver-brown stripes, the three intermediate ones
narrow, but still wider than the interspaces, the outermost stripes occupying thelateral margins of the sclerite; median area of scutum yellowish, the lobes gray,margined with liver-brown, most intense and best defined on the mesal margin of
lobe; scutellum pale brownish testaceous; postnotal mediotergite blackish. Pleura
blackened dorsally, the sternopleurite and meron paler, more brownish. Haltereswith base of stem yellow, the remainder, including knobs, dark brown. Legs with thecoxae brownish testaceous; trochanters obscure yellow; femora obscure yellow atbases, on distal half becoming dark brown, this area enclosing two yellow rings, one
apical, the other subterminal in position, the total effect being of two dark brown andtwo yellow rings; tibiae yellowish brown, the tips dark brown; tarsi light brown, theterminal segments blackened. Wings (Fig. 4) cream-yellow, more saturated along thecostal margin, with a very heavy brown pattern, distributed as a series of four costal
areas, broad seams to the cord, outer end of cell 1st M2, and the longitudinal veins of
the medial and cubital fields; area at stigma confluent posteriorly with the seaming along cord; wing-tip broadly and almost evenly darkened; cells M and Cu
extensively suffused with brown; extensive darkened clouds at ends of anal veins,sending paler washes basad into their respective cells; veins yellow in the ground-area,darker in the clouded portions. Venation: Scl ending about opposite one-third
the length of Rs, Sc 2 at its tip; h and a supernumerary crossvein in cell Sc present
but only faintly preserved; vein R4 +5 bending strongly caudad at r-rn, narrowing the latter; cell 1st M2 about as long as the longest vein beyond it; m-cu close
to fork of M.
Abdomen brownish black, the caudal margins of the segments narrowly and very
indistinctly paler; hypopygium dark. Male hypopygium (Fig. 16) with the tergite
transverse, each lateral lobe low, provided with about fifteen strong seta. Ventral
dististyle large and fleshy, the rostral prolongation small, slender, at base with acommon tubercle that bears two straight, subequal spines of moderate length. Dorsal
dististyle a chitinized sickle, widest just before the unusually long pointed apex.
Gonapophyses with the mesal apical lobe pale, blunt at apex, the lateral arms of theapophysis very naxrow.

HABITAT.-Brazil./

HOLOTYPE, e, Mount Roraima, summit, altitude 8600 feet, November, 1927
(G. H. H. Tate). ALLOTOPOTYPE, 9.

Limonia (Geranomyia) tatei is named in honor of the collector of thisunusually interesting series of Tipulidae, Mr. G. H. H. Tate. This fly isvery distinct in the heavy wing-pattern, which is approached in the
Neotropical fauna only by L. (G.) townsendi (Alexander) of Peru, anotherwise very different fly.

19311

11
NEW OR LITTLE-KNOWN TIPULIDE1

Limonia (Geranomyia) arecuna, new species
Figures 5 and 17
General coloration reddish brown, the prsescutum unmarked; legs brown;
wings subhyaline, unmarked except for the stigma; Sc short, Sci ending opposite theorigin of Rs; a supernumerary crossvein in cell Sc; male hypopygium with the ventral
dististyle large and fleshy, the rostral spines straight, separated at base, the innermost
from a slightly developed tubercle; gonapophyses acutely pointed at tips.
MALE.-Length, excluding rostrum, about 4.6 mm.; wing, 5.5 mm.; rostrum,
about 2.3 mm.
FEMALE.-Length, excluding rostrum, about 6 mm.; wing, 5.8-6.5 mm.;rostrum, about 2.4 mm.
Rostrum relatively elongate, black; maxillary palpi black. Antennie black
throughout. Head light gray, variegated with blackish areas on vertex.
Mesonotum reddish brown, without markings; scutellum more yellowish testaceous. Pleura yellowish testaceous. Halteres pale, the knobs infuscated. Legs withthe cox&e and trochanters yellow; femora yellowish at bases, passing into brown;
tibiie darker brown, the tarsi slightly paler, especially the posterior tarsi; fore tibine
not blackened at tips, as is the case in the tibialis group. Wings (Fig. 5) subhyaline,
unmarked except for the small oval brown stigma; veins darker brown. Venation:
Sci ending opposite or just beyond the origin of Rs, SC2 a short distance from its tip;a supernumerary crossvein in cell Sc beyond midlength; m-cu> close to fork of M.
Abdomen dark reddish brown, the extreme caudal margins of the segments paler;sternites brownish yellow. Male hypopygium (Fig. 17) with the tergite transverse,the very low lobes provided with numerous sete. Ventral dististyle large and fleshy,
its rostral prolongation small and slender, the two spines straight, obliquely acute attips, separated at base, the innermost arising from a small basal tubercle. Dorsal
dististyle a small curved sickle. Gonapophyses with the mesal-apical lobe an acutely
pointed blade.
HABITAT.-Brazil.
HOLOTYPE, e, Mount Roraima, summit, altitude 8600 feet, November, 1927
(G. H. H. Tate). ALLOTOPOTYPE, 9. PARATOPOTYPE, 9.

Limonia (Geranomyia) arecuna is named after the Arecuna Indians,
living near Mount Roraima. In its general appearance, the present fly
most closely resembles the Cuban L. (G.) myersiana Alexander, differingin the lack of blackened tips to the fore tibiae, and in the structure of themale hypopygium.
Limonia (Geranomyia) rubiginosa, new species
Figure 6
General coloration of thorax reddish, the prascutum darker medially but otherwise unmarked; antennae black throughout; halteres dusky; legs chiefly dark brown,
the femora not variegated with annuli at or near tips; wings with a brownish tinge,
the brown pattern very restricted in area; Sc short; m-cu close to fork of M.
FEMALE.-Length, excluding rostrum, about 5-5.5 mm.; wing, 6.2-6.8 mm.;rostrum, 2.6 mm.

AMERICAN MUSEUM NOVITATES

12[No. 491

Rostrum about one-half the length of body, black, the labial palpi paling at
3xtreme tips. Antennae black; flagellar segments subcylindrical, closely applied tome another; verticils relatively small and inconspicuous; terminal segment about)ne-half longer than the penultimate, pointed at apex. Head light gray, the posterior-ertex a trifle darkened on either side of median line.
Mesonotal prescutum reddish, more darkened medially; posterior sclerites ofmesonotum dark reddish brown to dark brown, pruinose. Pleura reddish yellow.
Halteres dusky. Legs with the cox= reddish yellow; trochanters yellow; femora
yellow basally, the fore femora almost uniformly yellow, the posterior femora
passing into dark brown shortly beyond base; remainder of legs brown. Wings(Fig. 6) with a brownish tinge; stigma subeircular, dark brown; very restricted toscarcely evident dark clouds at Sc2, origin of Rs, along cord and on outer end of cell
1st M2; veins light brown, darker in the clouded areas. Venation: Sei ending opposite
or just before origin of Rs, Sc2 at its tip; a supernumerary crossvein close to midlengthof cell Sc; Rs weakly angulated at origin; cell 1st M2 long, gently widened outwardly,
the second section of Ml +2 subequal to or a little longer than the outer section;
m-cu at or close to fork of M; vein 2nd A sinuous, the cell wide.
Abdomen dark brown, the caudal margins of the sternites very narrowly pale;
genital segment reddish yellow, the valves yellow, the tergal valves very gently
upcurved, their tips subobtuse; sternal valves short and stout, their tips pointed.
HABITAT.-Venezuela., HOLOTYPE, 9, Mount Duida, December 10, 1928
(G. H. H. Tate); Coll. No. 446. PARATOPOTYPE, 9, November 29, 1929; Coll.
No. 410.

Limonia (Geranomyia) rubiginosa is distinct from all describedregional species in the unvariegated wings, with Sc short, the uniformly
reddish thorax, the dusky halteres, and the unringed femora.
Helius (Helius) albitarsis (Osten Sacken)
Rhamphidia albitarsi.s OSTEN SACKEN, 1887, Berliner Entomolog. Zeitschr.,
XXXI, p. 184.

One broken specimen from Mount Duida, Venezuela, November 28,
1928 (G. H. H. Tate); Coll. No. 394. The species was described from
Porto Rico but is now known to have a wide range in the northern Neotropical region. It is more than possible that several closely allied species
have been confused under this specific name.

Limnophila kaieturana Alexander
Limnophila kaieturana ALEXANDER, 1930, Ann. Ent. Soc. America, XXIII,
p. 733.

One broken female, Mount Duida, Venezuela, November 28, 1928
(G. H. H. Tate); Coll. No. 394. Known previously only from the typelocality, Kaietur Falls, British Guiana.

1NEW OR LITTLE-KNOWN TIPULIDE113
1931]
Limnophila roraima, new species
Figure 7
General coloration pale ochreous, the pronotum and mesonotal prascutum witha capillary dark brown median vitta; antennae short in both sexes, the scape and extreme base of flagellum pale; legs yellow; wings cream-yellow, with a pale brownpattern that is mostly confined to the vicinity of the veins; m-cu at near midlengthof cell 1st M2.
MALE.-Length, about 8-8.5 mm.; wing, 8-8.6 mm.
FEMALE.-Length, about 8 mm.; wing, 8-8.5 mm.
Rostrum and palpi dark brown. Antennse short in both sexes; scape and basal
one or two of flagellar segments pale; remaining flagellar segments dark brown, thebasal and intermediate segments weakly bicolorous, the extreme base of each segment being pale; flagellar segments oval. Head yellowish gray.
Pronotum and mesonotal prEescutum buffy or ochreous, with a capillary mediandark brown line, the remainder of the usual three prsscutal stripes scarcely darker
than the ground-color; remainder of mesonotum ochreous; scutal lobes slightly
darkened; posterior portion of postnotal mediotergite with two confluent morebrownish areas. Pleura chiefly yellowish ochreous, in more heavily patterned specimens with brown areas on anepisternum and pteropleurite, to form a vague dorsal
darkening. Halteres pale yellow. Legs with the coxs, and trochanters yellow; remainder of legs yellow, the terminal tarsal segments slightly darker. Wings (Fig. 7)
cream-yellow, with a pale brown pattern, arranged as follows: extreme bases of cells
R and M, origin of Rs, fork of Sc, along cord and outer end of cell 1st M2, R2, tip of
R3, fork of M1 +2, m-cu, at about two-thirds the length of cell M, axilla, and as marginal
circular clouds at ends of veins M2 to 2nd A, inclusive; in more heavily patterned
individuals with additional clouds at near midlength of cells R4 and R5; veins yellow,
darker in the clouded areas. Venation: Sci ending just before the fork of Rs, Sc2,
longer, near its tip; inner ends of cells R4, R5, and 1st M2 in slightly oblique alignment; m-cu at or beyond midlength of cell lst M2; anterior arculus preserved.
Abdomen yellowish to dark brown, the sternites clearer yellow; hypopygium
yellow.
HABITAT.-Brazil.
HOLOTYPE, c, Mount Roraima, summit, altitude 8600 feet, November, 1927
(G. H. H. Tate); Coll. No. 40. ALLOTOPOTYPE, 9. PARATOPOTYPES, 10 e, 9, somein poor condition. PARATYPES, 2 fragments, altitude 6800 feet, November, 1927
(Gen. C. M. da S. Rondon).

Limnophila roraima was by far the most abundant crane-fly foundon the summit of Roraima. It is allied to L. kaieturana Alexander and
L. roraimicola, new species, but is well distinguished by the pale ochreouscoloration and the wing-pattern, which produces the effect of a very pale
brown spotting on a cream-yellow ground.
Limnophila roraimicola, new species
Figure 8

General coloration gray, the prescutum with three brown stripes; wings whitish
subhyaline, with a heavy brown pattern, including a series of nearly twenty more or

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AMERICAN MUSEUM NOVITATES[No. 491

less confluent brown spots in the costal cell; remainder of wing-disk with a heavy
brown pattern, some of the areas with pale centers to produce an ocellate appearance;
anterior arculus preserved; abdomen black.
FEMALE.-Length, about 8.5 mm.; wing, 9 mm.
Rostrum gray; palpi black. Antennse with the scapal segments dark brown, thefirst sparsely dusted with gray; flagellar segments broken. Head dark gray, somewhat clearer in front and on the orbits; two median, dark, more or less impressedspots on vertex.
Mesonotum gray, the prsescutum with a brown median stripe, and shorter, lessdistinct, lateral stripes; interspaces with brown setigerous punctures; no tuberculate
pits; pseudosutural fovea small, black, comma-shaped; posterior sclerites of mesonotum brown, gray pruinose. Pleura pale brown, pruinose, the anepisternum darker.
Halteres pale, the knobs weakly darkened. Legs with the coxwn and trochanters lightbrown, the former spasely pruinose; remainder of legs broken. Wings (Fig. 8)
whitish subhyaline, with a heavy dark brown pattern, including a series of nearlytwenty spots in cell C, these more or less confluent; areas at origin of Rs, at two-thirds
the length of cell M, at end of vein 2nd A; two additional areas in cell 2nd A are
brown with paler centers to produce a more or less ocellate effect; solid brown areas
appear as a seam from costa in the stigmal area, across the cord, narrowly interrupted
on M3 +4; outer end of cell 1st Ma2 darkened; several oval brown clouds in the cells
beyond the cord, these chiefly grouped at and near the wing-tip; veins pale, somewhat
darker in the infuscated areas. Venation: Scl ending opposite midlength of R2 +3 +4,Sc2 at its tip; Rs long, nearly straight; R2 less than one-half R1 +2; cell R3 deep;
inner ends of cells R4, Rs, and lst M2 in subtransverse to weakly oblique alignment;
m-cu at near one-third the length of cell lt M2; anterior arculus preserved.
Abdomen black. Ovipositor with the valves elongate, the cerci dark basally,
paler at tips.

HABITAT.-Brazil.
HOLOTYPE, 9, Mount Roraima, summit, altitude 8600 feet, November, 1927
(G. H. H. Tate).

Limnophila roraimicola is allied to L. kaieturana Alexander and L.
roraima, new species, but is very distinct from both species in the heavy
dark spotted pattern in the costal cell of the wings.
Atarba (Atarba) unilateralis, new species
Figures 9 and 18

General coloration yellow; antennae (male) elongate, exceeding one-half thebody; flagellar segments bicolorous, the base of each segment blackened, the apex
yellow; each flagellar segment with a very long, unilaterally arranged bristle; legs
yellow; wings yellow, the small diffuse stigma a trifle darker; Sc relatively long, Sci
extending to beyond one-third the length of the long Rs; abdominal tergites weakly
dimidiate, the bases yellow, the apices more broadly pale brown.
MME.-Length, about 6 mm.; wing, 6.3 mm.; antenna, about 4.5 mm.
Rostrum yellow; palpi dark brown. Antenna (male) elongate, as shown by themeasurements; scape and first flagellar segment yellow; succeeding flagellar segments yellow, the base blackened, on the proximal segments this including about the1NEW OR LITTLE-KNOWN TIPULID&E
19311

15

basal third, on the outer segments becoming more extensive but somewhat paler;
flagellar segments elongate-cylindrical, with an abundant erect elongate pubescenceand one very long bristle that considerably exceeds the pubescence on each segment,these bristles unilaterally arranged; on basal segments the bristle is placed beyond
midlength of the segment, on the outer segments becoming more basal in position;
terminal segment a little shorter than the penultimate. Head light gray, moreyellowish in front.
Mesonotum yellow, becoming more testaceous-yellow on the posterior sclerites.
Pleura yellow, the dorsal sclerites more darkened. Halteres dusky, paler at base ofstem. Legs with coxme and trochanters yellow; remainder of legs yellow, the terminal
tarsal segments infuscated. Wings (Fig. 9) broad, yellow, the small diffuse stigma alittle darker; veins yellowish brown. Venation: Sc relatively long, Sci ending beyond one-third the length of Rs, the latter elongate, about one-half longer than thebasal section of R3; cell 1st M2 long-pentagonal; m-cu at the fork of M.
Abdominal tergites weakly bicolorous, the bases of the segments light yellow,
the apices more broadly pale brown; subterminal segments blackened; sternites
and hypopygium more uniformly yellow. Male hypopygium (Fig. 18) with thesternal plate longer than broad, the lateral arms narrowly acute, glabrous. Outerdististyle slender, the apex bifid, the outer margin with three major spines andthree progressively smaller appressed denticles. JEdeagus relatively small andslender, in slide mounts not extending to beyond midlength of the basistyle.
HABITAT.-Venezuela.
HOLOTYPE, ce, Mount Duida, December 28, 1928 (G. H. H. Tate); Coll. No.
394.

Atarba (Atarba) unilateralis is most nearly allied to A. (A.) cincticornis Alexander (British Guiana) in the pattern of the elongate antennal
flagellum, which has the bases of the segments blackened, the apices
broadly pale. It differs most evidently in the wing-venation, in the long
Sc and Rs, and in the structure of the antennae.

Teucholabis (Teucholabis) duidensis, new species
Figure 10mesonotumcolorationofGeneralreddish, the prescutum with three black areas,each scutal lobe with two such areas; posterior legs entirely black, fore legs withfemoral bases yellow; wings whitish, the apex and a narrow seam at cord dark brown;
m long and arcuated, nearly twice the basal section of M3.
Sex(?).-Wing, 8 mm.
Rostrum and palpi black. Antenne black throughout; flagellar segments oval,
the basal six more enlarged, short-oval, the remaining segments smaller and more
crowded; terminal segment very small. Head black.
Pronotum reddish yellow. Mesonotal prascutum reddish, with three polished
black areas, one median on cephalic fifth of the sclerite, the others on lateral margins
behind the pseudosutural fovea; scutum reddish, each lobe with a large oval black
center and smaller lateral areas just beind the suture; scutellum reddish orange;
postnotal mediotergite similar, margined laterally with black. Pleura orange, with alarge central area occupying most of the mesopleura, including the dorsal sterno-

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AMERICAN MUSEUM NOVITATES[No. 491

pleurite and most of the anepisternum; meron black; pleurotergite orange, darkened
ventrally. Halteres black. Legs with the fore and middle coxse reddish yellow;
posterior coxie and trochanters black; fore legs black, the basal third of femora
yellow; posterior legs entirely black (middle legs broken). Wings (Fig. 10) whitish,
the apex and a narrow but conspicuous seam at cord dark brown; prearcular region
and cell Sc darkened; outer end of cell 1st M2 narrowly seamed with brown; stigmal
darkening larger than in decora; veins brownish black. Venation: Sc long, Sci
ending beyond five-sixths the length of Rs; vein R3 only gently curved; m unusuallylong, nearly twice the basal section of M3; m-cu close to the fork of M.
Abdomen broken.
HABITAT.-Venezuela.
HOLOTYPE, sex(?), Mount Duida, December 30, 1928 (G. H. H. Tate); Coll.
No. 497.

Closely allied to Teucholabis (Teucholabis) decora Alexander (Brazil:Para), differing especially in the details of venation, as the very long,arcuated m.