NEW OR LITTLE-KNOWN TIPULIDA FROM EASTERNASIA (DIPTERA), V 'By Cnlnr,ns P. Ar,sxlxPm'
Of Amherst, Massaclw.setts
TWO PI,ATES

The crane flies discussed at the present time are almost entirely
from the mountains of Formosa, where they were collected by
my friend Prof. Syuti Issiki. The most interesting series arethose taken at Shorei (Syorei) at an altitude of 7,000 to 8'000
feet, in late October, representing a tipulid fauna that is quitecharacteristic of the Palearctic Region in autumn. I am very
greatly indebted to Professor Issiki for the privilege of retaining
the types of the novelties described at this time.
In the present report, nine genera and subgenera of Tipulide
are added to the Formosan fauna; namely, Sti,badocerella, CEttarom,g'ia, Discobola, Thauma^stoptera, Ula, Troglophi'Ia, Pila'ria, Cladu,ra, and Neolimnoplt'i'ln. This brings the total of 8eneric and subgeneric groups for the island to more than seventg
a number that is greater than that of Europe or of America
north of Mexico. The additional groups found in the majorislands of Japan bring the number for the Japanese Empire toapproximately one hundred, with several additional generaand subgenera that are regional and will almost certainly betaken as a result of future collecting.
TIPULIN.€NESOPEZABASISTYLATA

sP. nov.

General coloration yellowish brown, the mesonotal prrescutum with a narrow brown median vitta; antenne relatively long,the basal segments yellow, the flagellum black; verticils of flagellar segments short; femora ancl tibia yellowish brown, the tarsi
chiefly snowy white; wings suffused with brown, the stigma
darker; abdomen dark brown, the segments ringed with yellow,
especially the sternites; male hypopygium with the basistyles
Agri'Contribution from the Departmentof Entomology,Massachusetts
culturalCollege.
b19

520

The Philippine Journal of Science

reze

greaUy produced into slender arms that are tufted with black
setre.
Male.-Length, about 8.5 millimeters; wing, 9.5; antenna,
about 3.3.
Femnle.*Length, about 8.5 millirneters; wing, 9.3.
Frontal prolongation of head pale brown, darker mediallyabove; palpi dark brown. Antennre (male) relatively long, thebasal segments obscure yellow, the flagellar segments beyond
the first passing into brownish black; flagellar segments'elongate-cylindrical, gradually decreasing in length outwardly, clothedwith a short, erect, pale pubescence and slightly longer black
Verticils that are more or less unilaterally arranged and shorter
than the segments themselves. Head light brown.
Pronotum dark brown. Mesonotal prrescutum rather lightyellowish brown, with a narrow brown median line; posteriorsclerites of mesonotum somewhat darker, especially the postnotal mediotergite; humeral region of prescutum restrictedly
pale yellow. Pleura pale, the sternopleurite and meron darker,
producing a more or less distinct longitudinally striped appearance. Halteres elongate, dark brown, the base of the stem
restrictedly yellow. Legs with the coxre yellowish testaceous;
trochanters pale yellow; femora and tibire yellowish brown, thetarsi chiefly snowy white. Wings (Plate 1, fig. 1) with a strong
brownish suffusion, the costal and apical regions somewhat
darker; stigma conspicuous, dark brown, irregular in outline,
oval, with a basal extension in cell Sc, almost to the fork of
Sc; eord vaguely seamed lvith darker; a feebly indicated paleantestigmal spot; veins dark brown, the obliterative areas restricted. Venation: Sc, lackingi Sc, at near three-fourths thelength of Rs, the latter arcuated; forks of medial field relatively
shallow; m-cu at less than one-half its length before the forkof M.
Abdominal tergites chieflly dark brown, the basal segmentswith a yellowish triangle on either side at near midlength; sternites even more conspicuously banded, dark brown, the extreme
base narrowly yellow, with an additional wider yellow ring at
near midlength of the sclerite; subterminal segments with thebasal pale ring lacking, becoming more uniformly darkened;hypopygium chiefly yellow. In the female, the banded abdominal pattern is much less distinct. Male hypopygium (Plate 2,
fis. 25) with the ninth tergite, 9t, having the caudal margin
blackened, trilobed, the lateral lobes larger and wider, the me-r,o,4

Alenander: Ti,pulida from Eastern As'ia, V

52Ldian lobe very lorv; on ventral surface back from each laterallobe extendsa blackenedserratedbar. Basistyle, b,very greatly
produced caudad. before midlength on mesal face with a tumid
swelling set rvith abundant microscopic setulre; beyond this enlargement.the st1'leis siender.culved gently mesad,at apex witha tuft of black set*. Outer dististl-le, od, Iong and slender.
Habttat.-F orrnosa.
Holotlpe. rnale. Rantaizan, altitude 1,000 to 6,000 feet, May
20. 1928 (S. lssiki). Allotopotype, female.
CYLINDROTOMIN.E
Genus STIBADOCERELLA

Brunetti

Stibailocerella BRUNETTI,Rec. Indian Mus. 15 (1918) 283.
Agastnm,Uia Dn MEtJnnE, Bijd. tot d,e Dierkunde 21 (1919) 17,

The genus Sti,badocerella has been represented only by two
closely allied species, S. presti.na Brunetti, from the Garo Hills,
Assam, altitude 3,500 to 3,900 feet, collected in July by Kemp;
and S. albitarsis (de Meijere) from Korinchi, Sumatra, taken in
September by Jacobson. The discovery of a third closely allied
species in Formosa is thus of great interest.
STIBADOCERELLAFORUOSENSIS

gp. nov.

Thorax polished reddish brown, without distinct stripes; boclywith strong greenish tints ; tips of fore tibie broadly white;
rvings with the second section of Rs longer than R.*u, cell R, being
relatively small; cell 1st M, large, exceeding the longest veinbeyond it.
Female.-Length, about 12 millimeters; wing, 9.2.
Rostrum and palpi greenish brown. Antennae u'ith the scapal
segments green, the flagellum black; antennre about as long as
the combined head and thorax. Head broad, greenish brown.
Mesonotum polished reddish brown, without distinct stripes,
the margins and posterior sclerites with greenish tints. Halteresbrownish black. Legs with the coxa and trochanters green;
femora brownish black; tibie black, the base narrowly ringed
with white, more broadly so on the fore tibia; tips of fore tibire
broadly white (2 millimeters) ; fore basitarsi much shorter than
the tibia, black, the extreme tip pale; remainder of tarsi snowy
white; posterior legs stouter, the tips of the tibia darkened;basitarsi with about the distal fourth white. Wings (Plate 1,
fis.2) with the coloration as in the other species,nearly hyaline,with very distinct black veins; stigma lacking. Venation: Basal
section of Rs nearly straight, subequal to the second section;

522

The Phi'tippi,ne Jownal of Sci'ence

1e2e

than the second
Ro-osubperpendicular I Ro-,.udistinctly shorter

i,s, cell R, retiivelv short; cell lst M' very larse'of M.r*,"r
Lxceeding in iength t"he cells beyond, the second section

rl?ii""

equaltotheoutersection;m-culongerthanthedistalsection
to nearlv;i C;'; vein 2d A represenied by a marg:inal thickening
midlength of cell lst A'
oPpositevalvesibdo*"r, chiefly greenish brown' Ovipositor with thetheunusually developei for a member of the Cylindrotomina'
conspicuous
i;"g"I urtu"* especially well-formed, margined withhalf paling
outer
the
blackened,
valve
tt
of
half
*Ji, 6r. basal"
to green.
Habitat.-Formosa.
20' 1928
Holotype, female, Rantaizan,4,O00 to 6,000 feet' May(S. /sszkz).
vein rvas
It has generally been assumed that the second anallackingintfrisgenus ,butthisisnotexactlytrue'Itisrepresentedbyurru*illutymarginalthickening,thecellitselfbeing
the arculus.
reduced io a linear stiip tying proximad of the level ofbeen ablb
I
have
Pruthi'Through the kindness of Dr' Singh
toexamineaparatypeofs.pristi'nnBranettiandhaveincluded
a few supplementary notes concerning this species:
antennre'
Mute.-Length, about 10 millimeters; wing, 9'2iabout 13.
. Antenne (male) much longer than the body, as shown by theflagellummeasurements, pale yellow, the outer segments of the
Prescutal
darker. Body Lolorution u* described by Brunetti'*trip"*distinctlyseparated.Paleringatbaseoftibiredistinct;) slightlv
pufu up"" of fore tibire relatively wide (1'? millim*f'basitarsi
it i.Lurrua, the sete similarly white; fore and middle
tips
about as long as the respective tibia, brown, only the extreme
hyaline' theof each a little whitenld. Wings (Plate 1, fig' 3)
its tip; secveins black. Venation: Sc. ending before r-m, Sc, at
Rat,
sectiotl
basal
the
than
shorter
littleond section of Rs a
M'
1st
cell
atrophied;
Rr*rentirely
andoblique; free tip of Sc,
ofsectiondistalthethan*-du'tottger
large, roughly rectangular;
nearly
A
long'
1st
vein
m-cu;
Cui vein Cu, evideni to opposite
straieht;vein2dAveryshort,oppositethearcu]usbecoming
confluent with the anal margin of the wing'
Genus CYI"IAROMYIA

Scudder

Surv' Terr' 3 (1877)
Cyttaromgia ScuDDEB,Bull. U. S' Geol' Geogr'(1894)
190-194'7111; Proc. Am' Phil. Soc. 32

40,4

Alerander: Tipulidu from Eastern Asta,

V

62B
The g:enuscgttaromyi,a has been known..
only from the Tercharacteristic of the Frorissantliaiy, having been especiaily
bedsof Colorado(Miocene). f am reterring
toglnus withsome questiona remarkable cylindrotom,ine tni.
Ry r"ori-no"*oruthat is certainry distinct trom" citind,rotomm.
one wing of the unique type of the present The venation of
speciesconforms
closely to the essentialfealures of itu known speciesof cgtta_
romyia' but the oppositewing has rost
the rup""nu-*J"]vein in celr Ru that furnistre-sttre ctrief""o*.character of the senus.
The most notabrefeature_sof the-present
fly are the very large,
nearly contiguous.eyes,
the coarseiypuncturedthoracic dorsum
and pleura (which reminds one strongryof the otherwise distinct Stibadocera),and the venation,
cell M. being present,con_nectedwith vein Rn*uby a supernumerarT,or adventitiouscross_vein, forming an acCdssory
discalcell immediatelyabovethe true
cell lst M (Plate 1, fig. 4), the last feature possibry
not constant.
It shouldbe noted that the p""runfny
disagreeswith scudder,sdefinition of cgttaromuia ii the rossor s","unJ i' tt u-f"".urr."of tibial spurs.
CYTTAROMYIATAIWANIA

sp. nov.

I

Generalcoloration black, the thorax coarsely
punctured; an_tenna (male) elongate,.lgu"rv u* torg
as the body, the flagellar
segmentscylindrical, with short u.*ti.il.;ii"ri"i^"r*".t
contiguouson anterior vertex.
"y",Male-Length, about ? millimeters; wing,?; antenne, about^
6.5.
Rostrum and palpi black. Antenne
dark brown, in malenearly as long as the body; flagellar
segmentselongate_cylin_
drical, with scattered pubesc"""u una
verticils that are much
shorter than the segments;antennal
segmentsbroken beyond
the twelfth; flagellar segments
Sraduallylrr."uurirrg irr l"rirti, totheor sixth, thence *tro"t""ing outwardly.
Head dull-fifthblack,
impunctate; eyes large, with ioarse
ommatidia, in themale alrnost contiguouson the anteriorvertex.
Mesonotumblack, the surface *iit,
tures, the three prascutar stripes and "orr.picuouscoarse punc_
centei; ;inearly smooth' prer'1'sbrack,the extensive il" ;;;ii, ,onu*
dorsopreurarmembrane paleyellow; pleura,,l""ropt"u"ite, with numer_
ous coarsepunctures. Halteres""."p1ifru
erongate,infuscaiei, lrr.'nuru
of the stem narrowly pale yellow.
Le? wittr
brownish black; trochanters obscurJ-yellow;the coxe smootir,
femora obscureyellow, the tips sligh,y mo"u it iur.ut"a;tibie brownish yerow,

624

TIut I'lt;il;i'pp'nrc'tournnL ol r)Qlt:tt<:t:yellow' passing
the tips very narrowly darkened; tarsi obscureintoblack.Wings(-ptate1,fig.4)withauniformgrayish
brown' Venabrown suffusion, witirout a stigma; veins darkRs; Rs long'
of
just
fork
the
beforetion : Sc' lacking, Sc, endingwithinterstitial
Rr*,
Rn*ui
u".out"O at origin, in alignment withofdistal-end,_-, urisirrg at end ;i d free tip of Scl'presi:ived;ofcell-Rucrossvein,inir,-i"ti""rv atrophied; a supernumerary
in
as
field
medialwing;
wing oi type, lacting in the leftii"
m-cu
of-.{r1ri
justfork"igfttbeyond them netore to
CiUnaroto*cr'
subequal to the distal section
M,,
tsl
cell
of
midlengthl"vottaof Cur.
except for
Abdomen elongate, brownish black, nearly smooththe transverse impressed areas; hypopygium blac'k'
Habitat.-Formosa.
feet' May
Holotype, matre, Rantaizan, altitude 4,000 to 6'000
20, L928 (,S. /ssiki).
LIMONIIN/E
LIMONIINILIMONIA(LIBNOTES)

CLITELLIGEBA

ap' nov'

pleura, pleurotergite,
Thorasic dorsum chiefly polished black;andsoutellumreddishy"tto*;femoraobscureyellow,thetips
beyond the cord
weakly darkened; wings grayish, the cellswith brown; Sc
rt"""gfy darkened; *o.I u"iot of wing seamed
m-cu beio"c;-Ii, and free tip of Sc, in transverse alignment;
divergent'
Vo"a to"t of M; anal veinsmillimeters; wing' 9'
8
about
Female.-Length,
terminal segtnent of maxbrown;
paipi
dark
Rostrum andflagellar segillary palpi smail. Antennn black throughout;very long' arranged
ments with the verticils of upper surface
elongate' Headsegments
ffagellar
pairs;
outer
utifal""affv inlarge' nearly contigdull black, the restricted frons gray; eyes
uousonthedorsum,restrictingtheanteriorvertexatthispoint
to a linear striP.
lobes polished
Pronotum, mesonotal prascutum, and scutal
restrictedly
veryprescutum
black, the humeral region of thebrown;
reddishscutellum
pale; median region of scutum andunipleurotergite
andpostnotal mediotergite black' Pleura
membrane'
Io"-fv reddish y"llo*, including the dorsopleural
Legs with theHalteres obscure yellow, the knobs dark brown'
yellow'
obscurecoxa and tronchanters reddish yellow; femora
thetipsweaklydarkened;remainderoflegsobscureyellow'theterminal tarsalr segrnents black; legs relatively long and slender.

Arcnorlctc,r:T'tpulitl,l from ilastern
Asia, VEZs
Wings_ (Plate l,"fig: 5) dimidiate, thecells beyond the cord
strongly darkened, the basal cells
more grayish; costal region,and especially cell Sc, more darkened;
stigm;;h;;_*"Lbrown;
prearcurar ceils and axilrary
region aarkenea;
atorigin of Rs; narrow but conspicuou*"-Lrg.'"roudb"owrr-ru"**"ulongCu,
the cord and outer end of cell ist
M,; other longitudinal veins
less distincty seamed-with n"o*r;veins dark brown. venation: Sc very long, ending some distancebeyond r_m; R, and free
tip of Sc, pale, in transverse alignment;Rs ur"aatuili origin,more than three times the basal
section of R"*^; ;;a]at veirrs
long,.generalrv
parailerce' r.st u,,*frii ri-lu a ritile
lessthan its length beyondtfre iort
"ututiu"ry
of M;
gen,y
divergent at base.
"rrrt'u.irr*

Abdominal tergites.dark brown, the outer segments more
reddish brown; sternites
yeilowistr brown. Ovipositor-morewith the genital
obscurev*lto*; t."g"t uutu"i'*t"rrau",
legmeatsently curved; sternalvarves stouter, their bases darkened.
Habitat.-Formosa.
Holotype, female, Rantaizan, altitude
4,000 to 6,000 feet, May
20, l92B (5. Issitci)
The peculiar body coloration
urrq wing pattern readily
separate the present species fromother d;."i;;-;;""iesorLi,bnotes.
LIIIONIA(LTMONIA) TENUICULA
sD. nov.

General coloration yellowish brown,

the prescutum

median dark brown. discal u"""; untun"*'*oniriiJrl,' with anru.t
throughout; wings wi,th a pafu U"o*nthestigmadarker;Sc, ending beyondmidlengthot nr; linse,
inner u"d;i:;;istM,strongly arcuated; m_cubeyondthe forkof lf ; ,naf" fopop"Sio*with the ventral dististyle .-uif, if,elost"af
prolongation small,without spines.
Male.-Length, about 4.b millimeters;
wing, b.6.
Rost-rumand parpi brownish bt"k. Antenne brownish brack,,throughout; flagellar segmentsstrongly
nodulose,the segments
short-oval with glabrous apical ;;;ffi;
turminal
segment elon_
gate, narrowed to a point.
Head dark blac.kish*;;.-*'
Mesonotalprescutum brownish;;il;;,the disk potisheddark
brown to brownish.black,the ground color to thehumeral and lateral portions;""-i"i.ifrg
of mesonotumdark
brown. pleura yeltowish b";;;:-'"",,,"?na.r
Hrlteres relatively short,dark brown, the base of the *t",ra"rowly yellow. Legs withthe coxa and trochanters yellowi*tiurta""ous; femora pale
brown, passingto darker ut iip.;
tilia ana tarsi pale brownish

The Phit'iytpilt'eJourn'al ol Scicrce

526

1020

(Plate 1'
yellow, the terminal tarsal segments darker' Wings"fig.
shortthetinge'
pale
brownishO) relatively narrow' with aSc
Venation:brown'
oial stiexna darker brown; veins dark
Sc'
Rs'
of
midlength
relativeft long, Sc, ending shortly beyond
inner end' of cellat its tip; free tip of Sc, and R, in alignment;
M' elongate;l-st
irt lt, si"ongty arcuated; veins beyond cellM'ofm-cu about one-third its length beyond the fork
Abdominaltergitesdarkbrown,thesternitessomewhatpaler'
9f' transMale hypopygium (Plate 2, fig'26) with the terg:ite'
whatu"t.", Lu"ft-lo* lateral lobe with about four long sete;is proappears to be the median ventral porbion of the tergite
ducedintoarectangularplatethatisdenselysetwithshortevensmall'
seta. Basistyle, U, tong, the ventromesal lobe relatively
nearlysimpleabyrepresented
Dorsal dististyle apparently
vety small'
Jraight rod, ihe tip acute. Ventral dististyle, ud'of the
lobe
the
of
size
the
about
tt u *uirr body of th" o"gutt
proportion
apical
mesal
the
seta,
few long
n"*i*tyt", *itlprolongation"
duced inio a small slender spinous beak; rostral
other thanarmature
evident
very large and flattened, without
set€.
Habitat.-Formosa.
Holotype, male, Shinten, April 15, 1928 (S' /ssz/ci)'
LIMONIA(DISCOBOLA)

MARGARITA(Alerander)'24 (1924) 535Discobolo-margarita ALExANoEn,Philip. Journ. sci'
640.
Hokkaido' northern
Known hitherto only from Karafuto and

7'000
Japan. A female, Mtunt Rantaizan, Formosa' altitude(S.
/ssikz)'feet, June 3,1927
LDIONIA(DISCOBOLA)

ARGUS(SaY)'(1824) 358'
Limnobia argus SAY, Long's Exped" Append'

Awide-rangingspeciesinnorthernNorthAmericaandnorth.
main islandeastern Asia, not before recordedfrom south of the
to 8'000
7'000
of Japan. A female, Shorei, Formosa, altitude
feet, October25,1928 (S. Issiki't'
LIMONIA(DICRANOMYIA)

AURITA

ap' nov'(wing:, ma'le, over 7
Belongs to the mori,o group; size large
the caudolateralmillimeters) ; Sc' long; male hypopygium with
anglesofthetergiteproducedintolongslenderarrns;-ventrodististylemesal lobe of basistyle long, boomerang-shaped; outerbifid at aPex.
Male.-Length, about ?'5 mil'limeters; wing, 7'4'

40,{

Alerander: Tipulidu from Eastern Asia, V

527

Rostrum and palpi black. Antenne brack throughout; flagellar segments oval to long-oval. Head black, tht frons moresilvery pruinose.
Mesonotum polished black, the median region of the scurum,
the scutellum, and the cephalic portion of the postnotal mediotergite slightly more pruinose. pleura brack, witrt a gray pruinescence. Halteres yellow, the knobs brownish black. Legs
with the fore and middle coxe porished brack, the apices moreyellowish; posterior coxa obscure yellow, trochanters yelow;
remainder of legs black, the femora yellow basally, narrowest on
the fore femora where about the basal fourth is included, least
evident on the posterior femora where onry the tips are darkened.
Wings (Plate l, fig. Z) with a brownish tinge, the oval stigma
darker brown; veins dark brown. Venation: Sc short, Sc, end_
ing shortly before the_origin of Rs, Sc, far before the tip, Sc,
alone being nearly as ,long as Rs; free lip of Sc, shorfly proxi_
mad of the level of R,; m-cu shorily before the iork of M, sub_equal to the distal section of cu,; vein 2d A relatively long.
Abdomen black, the caudal margins of the basar sternites obscure yellow; hypopygium chiefly dark. Male hypopygium(Plate 2, fig.27) with the ninth tergite, 9r, transverse,-the'outerlateral angles produced caudad and slighfly mesad into rong
slender arms that are tipped with short sete. Basistyle, b,
relatively smal,l, the vgntromesal lobe large and of very
unusualform, being more or less boomerang-shaped, the outer marginatthe angulation with a few ronger seta; a smail tubercle on
mesal
face of basistyle toward apex; a conspicuous pencil of longseta that are curved at tips, rocated near base oi styre. outerdististyle, od, bifid and brackened at apex. rnner dististyre,rid, fleshy, irregular in shape, the rostral prolongation
very
stout, with a sing:le pale spine. Gonapophyses with the mesal
apical angle slender.
Habitat.-Formosa.
Holotype, male, Ritozan, altitade b,000 feet, August 1,
1g2g(5. Issi,lti) .
Limonia (D.) atr'ri'ta is very distinct from the other regional
members of the morio group. The peculi ar charactu"*
of ttbasistyle and outer dististyle of the male hypopygium are"sug_gested by L. (D.) pseudomorio Alexander (Japani.
LIMONIA(DICRTINOMYIA)

MONTIUM

SI,. NOY.

General coloration brownish gray, the pronotum and meso_
notal prascutum with a median dark brown stripe; antennre
entirely black; wings grayish, the oval stigma darker; 3c, ending

r 0pu,t,t,(r,tL
Ir,o?tl ltoute?,n A:;i,0,

shortly beyond, Sc, nearly opposite, the origin of Rs; cell lst M,
relatively large; m-cu at the fork of M; male hypopygium witha single spine on the rostral prolongation of the ventral dististyle.
Male.-Length, about 6.5 millimeters; wing, 8.2.
Rostrum black, sparsely pruinose; palpi black. Antenne black
throughout; flagellar segments subglobular, passing into oval,
the terminal segment longer than the penultimate; segments
clothed with a dense white pubescence. Head grayish brown;
anterior vertex of moderate width, with an impressed medianline.
Pronotum gray, black medially. Mesonotal prrescutum brownish gray with a median dark brown stripe that is slightly constricted before midlength, somewhat wider behind; posteriorsclerites of the mesonotum dark gray, the centers of the scutal
lobes dark brown. Pleura dark brown, heavily pruinose. Halteres yellow, the knobs weakly infuscated. Legs with the fore
and middle coxa dark, pruinose; posterior coxe and all trochanters yellow; femora obscure brownish yellow at base, passing
into dark brown at tips; remainder of legs black. Wings (Plate
1, fig. 8) with a grayish tinge, the oval stigma darker; veins darkbrown, with long conspicuous black macrotrichia. Venation:
Sc, ending shortly beyond the origin of Rs, Sc, immediately
before this origin; Rs about two and one-half times the morearcuated basal section of Rn*u;free tip of Sc, and R, in approximate alignment; cell 1st M, large, longer than the veins beyond
it; basal section of M" arcuated to feebly angulated ; m-cu at forkof M.
Abdomen brownish black, sparsely pruinose. Male hypopyCrum (Plate 2, fiq.28) with the ninth tergite, 9f, transverse,
the caudal margin gently emarginate, the lateral lobes low. Basistyle, b, relatively small, the ventromesal lobe large, obtuse.
Outer dististyle, od, only moderately curved, the tip suddenly
narrowed into a slightly decurved black point. Ventral dististyle, ed, large and fleshy, the rostral prolong:ation flattened, thetip twisted and pendulous; a single rostral spine of moderate
size. Gonapophyses, gf, with the mesal apical lobe stout, its
lateral margin microscopically serrulate.
Habi,tat.-Formosa.
Holotype, male, Shdrei, altitude 7,000 to 8,000 feet, October
25, 1928 (,S. /ssi/ci).

LIMONIA(ALEXANDRIARIA)

ATAYALv

52g

ap. nov.

General coloration brown; prescutum
obscure yellow wilh
three brown stripes, tfre faierai-.t"ip.,broad, reaching theIateral margins or,o; *i,'g* with a strong brownis'-nearly
suffusion; sc long, sc, ending'armo.t
opposite the origin of Rs,
Sc, far from its
n,ro"i""
ir,untrr" nu*ar.e"uo-r,of Rr*u
Female.-Length,tip;about 42 millimeters;
wing:, b.

parpi black. Antenne macr
troiugJrout. Head'"ff*;H:nd
Mesonotarprescutum obscure brownish
yellow, with threebrown stripes, the median stripe
lateralstripes broadand diffuse; humer,alregion ot-sc,te"ite-;ril#"rrti"",;r1ff, scutal
lobesdark brown, the medianpa,rer;remainder of mesonotumdark"""r-l"o oistiot-scu;iil;
preura ;il;;;brown.
yerow,
striped longitudinally wittr
Ou"t i"o*n, includin* the ventral
anepisternumand
ventrar sternopreurite. Harteres-theinfuscated,the baseof the,tu* nu""o*lv or*"u"u y"il"#. "'iTgs
withthe fore coxa infuscated, irt"
ru-lin ng coxe and trochantersobscureyellow; remainder l"cJ;;ken._Wings (plate 1, fis.
9) with a strong"f,ufr*iJf-t;;;ti"r,
ifr"
orrrt
stigmal"q*nvery slighgy
darker; veins darkbrownSc^unusuallyiong, Sc,
ending just before the origin;i *u Sc, far from lts tip;
Rs
Ionger than the basal sectiJnO,_^,,free tip of Sc, and R,
inalignment,both pale; *_.u ut "if;#itf M, longer ttran ttre
distalsectionof Cur.
Abdominal tergites.
.dllk brown; sternites paler brownish

with thetu"guilongandstraight,theilitl"Xt;rr3]ipositor"ur"esHabitat.-Formosa.
Holotype, female, Shinten, Aprillb, tg2g (,S. Issi,ki),

ORIMARGAFUSCIVENOSA

sp. nov.

Generarcororationof head andthorax dark prumbeous;harteresand legsdark; wings *itf,a fuin"tbrownishtinge,
the veinsbrownishblack,,"r{R, an{ Rr*, subequal;basal:ollpicuous.
sectionof_Ronu
angulatednufor" _iajlr"rn.
Mate.-Leirth, a!o.u1b *il;;;teirl
ring, s.

o,ffTulTo#f,o;lf;.,'"uo"l;;nra.['tr,"oughout.
Head
Mesonotum and pleura entirely
dark plumbeous gray.
teres infuscated. L-egs
HaI_
th:
;;;;
and trochanrers darkryith'#r""r.(platebrown;remainder:_f_tsi"urr"t

1, fls. 10) witha faint brownishtinge,-the.rr"in"t*o-irrrish
black,
very conspi_
cuous; wing apex in radial
nefOna"rowly marginedwith brown:

b:JU

L',t,v r u Lt't lu*o r
I ruA I' ILLL',Lp',pcostal vein incrassated; costal fringe relatively long and conspicuous. Macrotrichia of medial field beyond cord long and conrpi"ooo.. Venation: Sc, ending about opposite midlength of Rs,
Sc, near its tip; free tip of Sc, nearly three times its length before
andR, i R, subequal to Rr*r; basal section of Rn*u?lsluted
R,
level
of
of
r-m
distal;midlengtn;
befor6
wlakly spurred
petiole of cell M, short; m-cu about opposite one-fifth the lengthof Rs; vein 2d A produced, the cell narrowed and acute at outer
end.
Abdominal tergites dark brown, the sternites a little paler;hypopygium black.
Habitat.-Formosa.
Holotype, male, Shinten, April 15, L928 (S. /sstka).
Ori,marga fusat'aenosa is most closely allied to O. taiwanensis
Alexander, likewise from Formosa, differing in the uniform dark
plumbeous eray coloration, including the thoracic pleura, andthe very dark wing veins, which are provided with long conspicuous macrotrichia. The details of venation are similarly distinct, including the position of Sc,, the distinctness of the free
tip of Sc,, the longer Rs, with cell R, relatively longer and narrower, and the more-pointed 2d anal cell.
Mik
GenusTHAUMASTOPTERA
ThaurnastepteraMrK,

Verh' zool.-bot. Ges. Wien 16 (1866) 302.

The ,genus Thaumnstoptera was proposed for the single species,calceatoMik, now known to be widely distributed in Europe.
The only other described species are the Oligocene T. electra
Alexander, from the Baltic amber, and' T. undntlnta (Cockerelland Haines), from the Gurnet Bay deposits. It is thus a matterof great interest to record a species of the genus from the highmountains of Formosa. The presence of three supernumerarycrossveinsin the wings of the new speciesrequires the formationof a new subgeneric group that is described below as Taitoanita.
This venational condition would appear closely to parallel thatfound in Limonin and its subgenus Di'scobola Osten Sacken. The
supernumerary and other transverse elements in the wing of
Tai,wanita (Plate 1, fig. 11) form an irregular cordlike arrangement lying proximad of the true cord, these crossveins and deflections being Sc, the basal section of Rs, the supernumerarycrossvein in cell R, the m-cu crossvein, and the supernumerarycrossveinsin cells Cu and 1st A. It should be noted that a super"
numerary crossvein in cell Cu is not known in any other crane fly.
The members of the genus Tha'rtmastoptera are peculiar in the40,I

Alerander: Tiputid,afrom Eastern Asia, V

i:ffit:tl*.,'u""*'characte",u,,, oJon;';;:##;
i:rrf

531

whichl'ivein coldspringsandconsrructa:;;:;eciarry
venation. Sup"".rrr,."u#Tror*ueinsdiffering esp
in celtsR, Cu, andType of the subg:enus,Thaum,astoptera(Taiwaruita) issirciana
sp. nov. (Eastern palearctic
Regiont.
TEAUIIASTOPTERA(TAtrWANITA) ISSIKIANA

nov.

"p.
General coloratioa grayish*friiu; antenna with thesegmentsinfuscated, theoutercuS*1rt*-g"ling to yellow;basal
yeltow, the femorat tips
Iegs.;;pi"r';;stybtackened; winss patevetlow' variegated *iti t;;;-ii.u,
deflectionsand numerousgray_on the crossveins anddots in
the',"n"*lr*"""ocrossveinsin cell R, Cu,""rfJ,
i.l.q.
Male.-Leryrth, about a""amifiimeters;
Rostrum obscure U"o*"irt -l;ilL.w, wing, b.
maxillary palpi btack.
Antenna with the--scap"tr.g*"ot*-brownish
testaceous;basal
segmentsof flagetum infuscated,
the
outer
.ug-"nt.
passingrnto ye'ow; basal flagetar*u*"nir
ovar,
the
outer
segments
more elongated;all segments;ith i;;; powerful seta thatthe segmentsin leneth. i;r;-;;iJ'l"uvi.r,
exceedwhite.
Mesonotumpare s:ravish *hi6;;itnout
markingg the lateral
portions of the prascltum*o".'yuioyi*h; posterior
gite a little infus.3lld_
medioter_yellow, with a dusky
lil_dt"on the dorsopleural region.
ctoua
H;il;;,
white. l,egs n"ittr th"
coxa and trochanterspale yellow,-i"_o"u
spicuouslyblackened,,o".cejJ;;'u^iu""o_ yellow, the tips conmore whitish ring;remainder of tess._n1G.""1,9l.-"d;q_ .(pJate1, fis. lij p"tu
ye'ow' handsomeryvarilgatedwitrr-'larr< brown seams
veins and numerouson thenareg.""1;r;;the cells; the darker seams
include the arculus, orisi_n
R"; uiiand deflectionsof veins, tips of Sc,, and-alf""f fonlituAinal"*"ru"insveinsi the grsy 4.ts
occur in all ce's; veins
nareygi;[,-r]"r."".in the seamedtreas.
Venation: Sc long, sc,uioi,t*lopo^.itethree-fourths
rength of Rs, S", ju:J before;il;;i_t"""ainsthethe latter; ns_rneu_
lated' with a supemum-erary
9jcrossveinat the angulation,connecting posteriorly with.{, n,t; ;'"".;;"
urignii.i;;j#;rnner end of cell R. lying
r-,r,fitUu p"oiimaO_otcell
tst M,; m_cu
nearry twice its length n.r""""lrt"-to'fr'ot M; a supernu.uru".," Lenz,Fr.,

Thaumasonfpra

rrrr^q1.66+u'^"uriaenra".,,L,
Ar;#;;'irTi,::;:,""f,h"eti#f;*
ripu-' -v e46v D L.ruuue oa' A D t' A .' S"l.uu.etragente
H eft
11{t-1 36, fi gs . 1-2g.

4 for 1g19 (1920)

The Philippine Journal of Science

5gZ

rszs

crossvein in cell cu at near midlength, this completely traversing
vein Cur; a supernumerary crossvein in cell 1st A, connectingwith vein 2d at near two-thifds its length'
Male
Abdomen pale yellow, the pleural region more darkened'd,single'dististyle,hypopygiurn (Plate 2, fig. 29) with theproas in the genus, appearing as a flattened blade, the apex
small
a
with
Juced into a slendei acute point, the outer margin
appressed tooth.
Hubi,fu't.-Formosa.
Holotype, male, Sh6rei, altitude ?,000 to 8,000 feet, October
25,t928 (S. /ssokz).
HELIUS(HELIUS)NAWAIANUS

glt' nov'

prrescutum
Mesonotum reddish yellow, the cephalic hal'f of thewith a dark brown median s,tripe; antennre (male) relatively
long,abouttwi ceaslongasthecombinedheadandrostrum;
abdimen dark brown, the hypopygium yellow; male hypopygiumthewith the outer dististyle terminating in two obtuse spines,
powerfulsete'longouter margin with three
Male.-Length, about 6.5 millimeters; wing, 7'5'
Female.-Length, about 8'5 millimeters; wing' 7'8'
Rostrum and palpi dark brown. Antenne relatively elongated,
as in .I/. wi'pponensis, about twice as long as the combined head
androstrum,darkbrown;flagellarsegmentscylindrical,witha dense whiie pubescence and short scattered black verticils.
Head gray.
Mesono'tum
Pronotum dark brown medially, paler laterally'a darkwithprrescutum
reddish yellow, the cephalic half of theunmarked.
notum
thebrown median stripe, the remainder ofPleura reddish yellow. Halteres pale. Legs with the coxn andyellow,trochanters concolorous with the pleura; femora obscureyellow,
thetips
the tips narrowly infuscated; tibire brownish
beyonduue.r"iy darkened; tarsi passing into brownish black
only
bales. Wings with a very pale brownish tinge, the stigma
justslightly darker; veins dark brown. Venation: Sc, ending
beiore the fork of Rs, Sc, at its tip ; Rs nearly straight; cell lst
M, large, the proximal end wider; m-cu close to the fork of M'
paler;
Abdomen dark brown, the basal sternites a very little
hypopygium and subterminal segments yellow' Male hypopygi;; *itft tft" outer dististyle, od (Plate 2, fig'30), ending in
produced
Iwo sub"qual blackened knobs, or obtuse spines, not(Platefrg'31)
2,
H.
ni'pponensis
; on
in
as
into a long apical spine
powerful
that
seta
long
three
with
apex
beforeootl" **gin40,4Aletander: Tiputidn from Eastern Asia, V

bBB
are entirely lacking in ni,pponens,i,s,breuioricorni,s,and all,ietlrorlns.
Habitat.-Japan(Honshiu).
Holotype, male, Oi, Gifu, June 6, 1g2B (n. Talcahashi).
.
Allo_
topotype, female.
Helius nnwuianus is named in honor of
the great pioneer
entomolog:istof Japan. In H. nipponensis
Alexander the pra_scutum has three distinct stripes, the centersof the scutar robes
are blackened, and the abdomen is much paler.
EELIUS (trELIUS)

LILIPUTANUS

sp. nov.

Size very small (wing, male, under B.b millimeters);
rostrum
elongated; thorax ocherous yellow; wings grayish
subtrvafirre,
without a distinct stigma; Sc short; cell M,
opu" UV tt e at"ophyof m; abdominal segments conspicuously bicolorous.
Male.-Length, excluding rostrum, about B mllimeters;
wing,
3.2;
rostrum, about 1.1.
Rostrum elongated, dark brown, exceeding
the combined head
and thorax. Antenna brownish black, with
only fifteen appar_
ent segments; first flagellar segment large;
terminal- segment
reduced to a tiny button; verticirs rong andconspicuous, muctr
exceeding the segments, more elongate on theouter segxnerrts.
Head pale, the anterior vertex ,u""o*.
Thorax pale ocherous yellow, the mesonotum
somewhat darker
behind; sternopleurite slighfly infuscated.
Halteres du*k b"o*n,the base of the stem narrowry pale. Legs
with the coxe andtrochanters pale yellow, those otlne fore legs
somewhat darker;
remainder of legs broken. Wings (plate
1, fig. 12) grayish
subhyaline, the stigma scarcely darkened;*ingi_u"grr-lr,
dial field narrowly and vaguely seamed with",u_brown; veins pale
brown. Venation: Sc short, Sc, ending about
opposite midlengthd.
short Rs, Sc, at its-tip; veins R, and nr*u at*ing *"rginth:widely divergent, cell R, heing urru*oully
widllcell fU,
nvthe, atrophy of m; m-cu nearly its own length"p""fork
beyond" th-eof M, Ionger than the distal section of
Cu,;Lff Zd e n"""o*.
Abdominal segmentsconspicuousrybicororous,
the basesbroaoly obscure yellow, the slighfly ,ru""o*u"apical portions darkbrown; subterminal segments uniformry aarr.e"ei,-io-rriirr,
ring; hypopygrum chiefly pale. Male hypopygium"
filiru z,
fig. 32) with a conspicuous interbasal rod,-i.- outer diJstyte,od, slender, bifid at apex, the outer toothbroken near base in
the unique type.
Habitat.-Forrnosa.
216827_8

534

The Pluilippine Jourrml' of Sci'ence:gzs

Holotype, male, Michishita, altitude 4,000 feet, July 31, 1928
(S. /ssi/ci).
The structure of the outer dististyle is generally similar to
Tui,wani,nn pando*a Alexander, and it is very possible that thelatter genus is more correctly placed near Heli,tts despite thereduced palpi, peculiar antennal structure, and shape of thethorax.
HEXATOM INIULASUPERELEGANS

sp. nov.

General coloration black, pruinose; antennre elong:ate; wings
yellowish, with a heavy brown pattern, including a large areasurrounding Scr; stigma yellow, both ends darkened; abdominal
terg:ites black, the caudal margins narrowly ringed with obscureyellcrw; sternites dark, the incisures broadly yellow.
Female.-Length, about 6.5 to 7 millimeters; wing, 7 to 8.
Rostrum and palpi black. Antenna elongate, in female, if
bent backward, extending to some distance beyond the wingroot; scapal segments obscure yellow, the flagetrlum black; flag:ellar segmentsfusiform, with long conspicuousverticils. Headbrownish gray.
Mesonotum black with a conspicuous brownish yellow pollen,
the scutellum and postnotum somewhat more pruinose. Pleura
black, sparsely pruinose; dorsopleural membrane dusky. Halteres dusky, the basal half of the stem yellow, the knobs more
infuscated. Legs with the coxa brownish yellow, paler apically;trochanters obscure brownish yellow I femora brownish black,the bases obscure yellow, more broadly so on the posterior legs;
tibie yellowish brown to brownish black, the tips still darker;
tarsi passing into black. Wings with a yellowish color, varieg:ated with a very heavy brown pattern, including extensive
seams at Sc, origin of Rs, along the cord from the costa at Sc,
to Cu; outer end of stigma surrounding R, and Rr*r; outer endof cell 1st Mr; narrower dusky clouds at wing apex, as marginalareas at ends of radial veins and as a conspicuous seam for almost the whole length of the distal section of Cu,; stigma chiefly yellow, brighter than the ground color; veins dark brown'
The holotype has an additional small linear dark seam in cell 1stA. Macrotrichia of cells nearly lacking in the proximal ends ofthe basal cells, including nearly the basal half of 2d A. Venation: Rs angulated at origin, forking at r-m; m-cu nearly inalignment with r-m.
Abdominal tergites black, the segrnents narrowly ringed caudally with obscure testaceousyellow; sternites similar, the inci-

40,,tALcnu,rttlcr:Tiputid,n lrom Eastern
Asia, Vb$bsures broadrv an{ cons'icuorlsry
obscure yeilow, this incruding
both the basesand apices_ofthe"segments;
subterminal,segments
more uniformry brackened. ovipositor
with the bur" of tt s"nital segment infuscated, the remainder"
brownish yellow; tergal
valves strongly upcurved. to the very
acute tips, the dorsal margin
at base with a low obtuse darkened
tooth.
Habitat.-Formosa.
Holotype, female, Shori, altitude
7,000 to g,000 feet, October
25, 1928 (5. Issiki). Paratopotypu,
female.
Ula superelegans sp. nov. andU. fuscistigma sp. nov. are
distinct from the two species known
from north ern Japan, Ula
perelegans Alexander and,U. cincta Alexander.
ULA

FUSCISTIGMA

sp. nov.

General coloration black, pruinose;
antenne elongate; wings
grayish, the stigma uniformly
brown; Iarge subcircurar brownof Rs and on anterior cord; abdo-"r,unito"rr,ty;i""*r:l."rigin
Female.-Length, about 7 millimeters;
wing ?.2.
Rostrum and palpi black. e"tu""*(female) elongate; scapedark brown, the flagellum black.---tfeaa
dark gray.
Thorax black, the mesonotrm *pair"lypruinose, the scutellum
and postnotum more heavily so. pleura.";;i.;;;livp"ri_
nose' especiary the ventral preurites;
dorso pruo"ui""eio'au"t.
Halteres pale, the knobs dark brown.
f,"s* with the coxa darkbrown, the trochante,rs more f""#rirf,
yellow; femora brownish
yellow, passing into black;of Iegs black. Wings(Plate 1, fig. 18) grayish,
the"u-"irrjur
stigma uniformly darkened; con_spicuous subcircular brownorigin of Rs and on anterior cord; narrower-seams "roua?
"t
on posterior
cord and outer end ofcell lst M"r a tiny brown *.ui"-onsc,; veins dark brown.
Macrotrichia of cells- llgkine
exrreme wing base, in_cludingaboutthebasarthird of .ufzJ-rly;t e. venatior,:-n.
ii"r.irrg
just beyond r-mi Rr+zat"ine fo"gu"*tt"r, R,
Lv2 @tvtLv
alone;
cell
cerl
Lst
M,
-L
felatively
small.
'
Abdominal tergites black, the sternites
a lit'e paler, the pleu_
ral membrane dark; ovipositor
v"ifo*i.f, horn_color, the tergal
valves relatively broad, ,p"o",o.dto il" acute tips.
Habitat.-Formosa.
Holotype, female, Shori, altitude2,000 to g,000 feet, October
25, 1929 (5. Issiki).
PSEUDOLIMNOPEILAAUTUMNALIS

sp. nov.

General coloration gray, the prrescutumwith four narrow,rrwn stripes; antenna black,tne nase'ot the first flage,ar
seg-

536

Th,e PhiLi'ppine Jo't't,rn'alof Scicncc

1020

ment yellow; legs chiefly brown, the tarsi passing into black;wings with a grayish brown suffusion, the pearcular and costal,"giorr. more yellowish; stigma and a vague pattern on the disk
dait<er; distai section of Cu, straight; male hypopygium withthe edeagus short.
Male.-Length, about 8 millimeters; wing, 9'3'
Female.-Length, about 7 millimeters; wing, 8'2'
Rostrum brown, the palpi darker brown' Antenne (male)
relatively short, if bent backward, extending to just beyond thewing root; antenna black, the basal third of the first flagellar
segment yellow; flagellar segments subcylindrical, with verticils
that exceed the segments. Head brownish gray, clearer gray infront.
Pronotum gray, the lateral portions of the scutellum obscureyellow. Mesonotal prescutum gray with four brown stripes,
ihu intur-"diate pair separated by a capillary gray line; pseudosutural fovea small, black; no tuberculate pits; remainder ofmesonotum gray, the scutal lobes variegated with brown' Pleuragray. Halteres dusky, the base of the stem narrowly pale;
itt iitu paratype, the apices of the knobs slightly paler' Legs
with the coxe brownish gray basally, the apices pale; trochantersobscure yellow, margined with brown; femora brownish yellow,
paler basally, the tips passing into dark brown; tibie and tarsi
iark brown, th" tutt"" passing into black. Wings (Plate 1, fig'
14) with a grayish brown suffusion, the prearcular and costal
regions *o". y"llo*ish; stigma darker; very vague and diffuse
clouas at origin of Rs, along cord and outer end of cell 1st Mr;
axillary region darkened; veins dark brown. Macrotrichia ofveins unusually long and conspicuous; costal fringe short in both
sexes. Venation: Sc, ending about opposite r-m, Sc, at its tip;
Rs angulated at origini Rz+s+rrelatively long, gently arcuated'
longer than Rr*r; R, subeqriaitb_!r+r; cell 1st M, short, m-cu atnear two-ttrirdi" its length; cell M, deep, its petiole subequal toor shorter than m-cu; distal section of Cu, straight; vein 2d A
long, gently sinuous.
Abdominal tergites dark brown, the central portion moreyellowish to prodube a more or less distinct dorsomedian vitta;
subterminal segments and hypopgium entirely blackened; sternites infuscated, the intermediate segments with a large yellowish triangle on posterior half, the point directed cephalad' Male
hypopygium (Plate 2, fig. 33) with the tergite, 9t, deeply
notched. Basistyle short and broad. outer dististyle, od, narrowed to the simple apex, the outer surface setiferous' Inner

40..lAlo,t:utttlu.: TiTtulidu from Dastern Asiu, VbB?
dististvle with the basal harf enrarged, theouter margin withnumerous conspicuous erect seta, the apical point
Jongate.
.4Edeagusshort. What appears to represent
an interbasal pro_
cess appears as a trispinous structure,
as figured.
Habitat.-Formosa.
. j{o_ld-{qu, male, Mareppa, altitude 6,000 feet, October 28, Lgzg
(S. lssz'ki). Allotype, female,Sh6rei, altitude 2,000 to 8,000
feet, October 2E,1g2g (S. Issiki). paratopotype,male.
The onlv sim'ar grayish pseiiotimnophiia
from this generar
region is P. h,orii' Alexander (Japan),which differs in the general coloration of the body and wings,
the yellow harleres, thevenation, as the strongly curved
disial section of Cu,,lna tfre
structure of the male hypopygium.
TROGLOPHILARITOZANENSIS

sp. nov.

General coloration brown, the humeral regionof the prrescu_
tum more yellowish; head brownish ,gray;wings with a browntinge;.Rs rn{ Rr*r*onlbgcuat, the forrier
slighfly morearcuated;
m-cu just before'ihi fork of M.
Female.-Length, about B.b millimeters;
wing:, 4.6.
Rostrum and palpi pale brown. Antenna(female) of mod_erate length onry, if bent backward extending
about- to-- ttre---l wingroot; antenna dark brown. Headbrownish gray.
Mesonotum brown, the humeral regionof prescutum moreyellowish. pleura brown. Halteresdusky, the base of the stem
narrowly yelrow' Legs with the coxa
brownish tu.tu.uou., trr"
posterior coxre paler; remainderof legs pale brown, it*.r*darl<. Wings (plate 1, fig. 1b) with" tf,"
a iuint n"o*.,i*fr'tirrg",
stigma scarcerv darker; veins anJ
Lacrotrichia dark brown.
Venation: Sc relatively long, S., unaing-justbeyond r-m, Sc, a
short distance from.the tip of S.r;n,short, arcuated,
subequal to or slightly Ionger tiian "Aatively
the more gen'y arcuatedRrfaint, Rr*, nearly ,. f""e as R,*r,
fr*rnnithe latter Iess"-.Iy_
tha.n
one-halfR,*

sto"ttvne"t";;if

fil #*.
Abdomenuniforml.,i,i,_.obrown, ttre sternitls a litfle paler.
Ovipos_itor with the valves fleshy,'terminatin*
t' u srenderpale setoid
structure.
The left wing of th:.l.rq: has cell M, present
as a small areotethat is lessthan one_fifth1helength oftfr" petiole.
Habitat.-Formosa.
Holotype,female,Ritozan,altitude b,000
feet, August 1, Lgzg
(S. /ssi&i).
Trogloph,a ritozanensisis most closeryarlied to T. articora(Edwards), of Borneo. It cannot
ne stateawhetherthe condi_

538

Th'e Plr,itippine Jourrut'Lof Sciencetttzllof thetion of having cell M, preserved is normal, the venation
unlike'two wings of the type being conspicuously
LTMNOPIIILA(TRICIIOLIMNOPUILA)

C'|ESIELLA

sp' noY'

Male.-Length, 7.5 to 8 millimeters; wing, 7 lo 7 '5'
pi'Li'ferAlexanClosely allied and generally similar to L' (T')
of colordetails
andder, differing in the Jigntty iottg"" antenne

ation.
in 1''
Rostrum and head light grav' Antennr longer than
root
to
the
Itilif er, if bent backward extending approximately
pragray,the
It trr" halteres, pale brown. Mesothorax entirely
the entire
scutal stripes scarcely evident, the pruinosity includingp"*..rto-.Malehypopygiumwiththelobesofthetergitelong
more orand *ore expanded at apices, the notch inclosed thus
less narrowed at outer end.
(Kiushiu) .
Habitat.-Japan
Hill, near Fukuoka, April 5' 1924 (H'
Hirao
Holotype, male,
Fukuoka'Hori,). Paratopotypes, 1- male, with type; 2 males'(H.
Hori).
April 13,1924
pili'fer
The specimens of the rather extensive series of L'
beforemeallshowthecephalicportionofthemedianprrescutal
stripe highly polished and shiny black'
LIMNOPHILARANTAIZANA

aP' nov'(male) short;
General coloration brownish yellow; antenna
macrotrichia
pattern;
dark
weak
wing. brownish yellow, with a
Ru, and 1st
Rn,
cells
of
ends
of veins long and conspicuous; innermedianthewithi{, i; oblique alignment; male hvpopvgiumrod'bilobedaintoof the tergite produced caudad"Mate.-Length,""gionabout 5'5 millimeters I wing' 6'8'
(male) short'
Rostrum yellow; palpi dark brown' Antennre
the wing
beyondto opposite or iust
if bent backward"*t"ttdittgput" brownish yellow, the outer segments somewhat darker;
pale pubescenceand"oof;nug"tL" segments llng-oval, with a dense
grayish ocher,r"iti"it. ttrat stigtrtty exceedthe segrnents' Headous, the center of the broad vertex a little darkened'yellow, subnitidotts'
ieneral coloration of the thorax brownish
withoutevidentmarkings;pseudosuturalfoveaconcolorouswith
theprescutum;notuberculatepits.Pleuratestaceousyellow.
yellow' the knobs
Halteres relatively long and slender, obscurepale yelmore darkened. l,egs wittr the coxa and trochanters
low;femorabrownishyellow,alittled.arkenedoutwardly;tibire
and.tarsipalebrown,theoutertarsalsegxnentssomewhatdarker;
fis' 16) with a strong
i.g. .o".picuously hairy. Wings (Plate 1'

40,.r

Alctt;urxlcr: Tipulid,"ef rom Eastern Asia, V

SBg

lrrownish yellow tinge; stigma oval, d.arker brown, very
vague
:lnd restricted; dark seams on anterior cord; veins brown,themacrotrichia long and conspicuous. Venation: Sc, ending justbefore the fork of Rs, sc, a short distance from its tip; Rs longand g:ently arcuatedi Rr*r+rshort, less than the basal sectionof R,
sub-eq"dilto Rr*r; inner ends of cells Rn,l: l{:, .Rr.,_r_
H=, and lst M, in oblique arignmentj the rast-named most proximad; cell M, deep, about one-half longer than its petiole;cell
lst M, small, m-cu beyond midlength; anterior u""uio* present.
Abdomen pale brown, the sternites somewhat paler. Male
hypopygium (Plate 2, fig. 84) with the tergite, gt, produced
medially into a narrow rod, the apex bilobed.' gasistyre,b,relatively short and stout, the interbasal process small, pale,ap_'a'narrow
pearing as an obtuse blade. Outer dististyle,od,,rod, the apex unequally bidentate. Inner ii.ii*tyi",id,, verysmall, pale, setiferous, the lower or cephalic margin
nefore miO_
Iength bearing two or three more powerful setafrom
tubercu_late bases. .?Edeaguslong and slender, pale, the apex apparenfly
broken in the unique type. GoSapophy*u., g,-"pp.u"ingu,slender hooks, narrowed to the acute tips.
Habitat.-Formosa.
Holotype, male, Rantaizan, altitude 6,000 to 2,000
feet, May21, 7928 (5. Issiki,).
The structure of the ninth tergite and outer dististylesomewhat suglgests mernbers of the subgenus Tricholi^oophilo,b"tin other respects there is ritile in common between the groups.
PILARIAFORMOSICOLA

sp. nov.

General coloration shiny dark brown, the preura more b ownish yellow; wings with a strong brown tinge, the stigma
darker;R, just before the fork of R"*n; cell M, present; m_cu at about
two-thirds the length of cell l;t-M,; abdominal tergites uniformly
blackened, the sternites yellowish brown.
Femnle.-Length, about 9 millimeters; wing, g.5.
_ Rostrum and palpi dark brown. Antenna (female) rerativery
elongate, if bent backward extending about to theof trru
halteres; antenna brack throughout; flagelrar segments"ootelongate,with long conspicuousverticils, the dorsal, unilaleralry u""urg"aseries much exceeding the segments in length. Headdarkbrown.
, Mesonotum shiny dark brown, the preura more brownish yerlow. Halteres relatively elongate, dusky, especially the knobs.
[,cgs with the coxre and trochanters yelro*; -t"-uinderoi tegs
blnck, the femoral bases vaguely brightened. Wings (plate
1.

540

Thc l'hil'ippine Journul o,f Scicnce,,,!rrfig. 17) with a strong brownish tinge, the oval stigma darker
brown; veins dark brown. Venation: Sc, ending some distance
before the fork of Rs, Sc, at its tip; Rs ang:ulated and weakly
spurred at origin; R, oblique, justbefore the fork of Rr*r; innerends of cells R,, R,, and 1st M, in oblique alignmentl ieli IVr.
longer than its petiole; m:cu at about two-thirds the length ofcell 1st M,; vein 2d A elongate.
Abdominal tergites black, the sternites yellowish brown.
Ovipositor with the valves very elongate; tergal valves darkbasally, the tips paling to yellow; sternal valves yellow, the bases
more narrowly blackened.
Habitat.-Formosa.
Holotype, female, Sorempi, near Giran, altitude 1,b00 feet,
November 4, L928 (5. Issi,lti,).
rcLADURADEcEM-NorAToo.r"""ulloPTERrN
Cloilurad,ecem*notata
Ar,nxaNoon,,Ann. Ent. Soc.America1Z (1924)
43U437.
This interesting species was described from various stations
in Kiushiu, Japan. In the present series of Tipulida from
Professor Issiki were a few males of what appears to be thesame species from Shorei, Formosa, altitude 2,000 to g,000 feet,
October 25, L928. The type series was represented only by females, while the present material ineludes only males. The Formosan material shows slight differences, chiefly in the size andintensity of the black body markings, and it is possiblethat they
may represent a distinct but close,lyallied species.
NEOLIMNOPHILA

ALTICOLA

sD. noy.

General coloration gray; mesonotal prrescutum with the cepha_lic ends of the intermediate and lateral stripes shiny polished
black; legs black; wings grayish, the prearcular region yellowish;
stigma a little darker than the ground color; maie hypopygiumwith the basistyle having a single curved spinous brisfle nearbase.
Male.-Length, 6.5 to 7 millimeters; wing:, Z to g.
Female.-Length, 7 to 7.5 millimeters; wing, T to 8.
Rostrum dark gray; palpi black. Antenne black throughout;fusion segment elongate-conical,composedof four subsegments,
the sutures between indicated; segmentsbeyond the fusion subcylindrical, gradually increasing in length to the antepenultimate,
the remainder shorter. Head brownish gray, the front and re_stricted posterior orbits clearer gray.

40,rAlerander: Tipulid.n rom EasternfAsia, V

547

Mesonotalprescutum yellowishgray
the cephalicend of the-inie"*uafitu with four brown stripes,
pairstripe polishedbrack; port""io" -."r"rites and of each lateralof mesonotumcrearer
gray. pleura gray. Halterespaiu, tt u t"on, *.ui.L
dusky.
Legs with the coxe-gray; trochanters
yellowishbrown; remain_
der of ,egs brack, tr,e remorai;;very narrowly and restrictedly obscureyellow. WinsslH"i9r, fig. 18) with a srayish
suffusion,the oval stigrna
ilr;;tt'edarker than the groundcolor; prearcular.region
yellow; veins brown.
venation: sc, ending,u""i-"o;#i;"orrrpi"uorsly
thefork of Rs, sc, at its
tiP; Rr*r*, elongate;n, ,ru"iu^niJfi'-position,
in most instances
about its own Iength buyonatf,"-.

small;
cenvr,.ur"qrurtoor;n"h;;,:X}.ffr*,iil,;"r,"tll'*-y;variable in posi'tion,trom
ctosellirru*"
r*r. of M ito oppositetwofifth5
leneth of ceil lJ M,.""-the
Abdomen, including tfru irypopvgium,
MatehypopyciyTetiie-T,?*. black, sparsely prui_:o:..
ilsl *iir,"iilu"[u*irtyru,o, rong'and slender,at basewith a single.long-"u"-iujrpinou,bristle. Outer dististyle,oa,lo"i Jnd slender,curved,
thenarrowedand arched,theouie'iu"gi" with appressed apex
denticresthat are somewhatmore erectand ionspicuousjust before
apex. fnner dististyle wittr
theconspicuous
sete.
Hubitat._Formosa.
Holotype, male. Shirasetsu,
altitude1e28(5. rssiki).6,000 feet, October 24,
Sh6rei,altitude2,000to_?ffi;,
8,000feet, October-2b,^jtr""lry;$2dparatopotypes,b mates,
1 female; paratrngl, 1 male,-iS.lrJki).
t teiale, rvith the allotype.
Neolimnophilnatticolo i* u_pfJ^Oi*tin.t
from the other species known from easte"r,eriu; namely, N. uttima(osten sacken)' N'"nJ.-,i#r,J*
snni'tiiii
rri"r""tttr (equarssimpler Brunetti), and N. fuscinerais pA*""Ai.'"'LIPSOTERIX

pLUTO

ep. nov.

General coloration shiny coal_black;antennre (male) elongate;
Ie'gs brack' only the extreme
bases of ihu fu*o", obscureyerow;
wings with a strong, dusky
fid;, th";val stigma darker.

aboutz.s ,iiifi.ltl"r,"rmrr;;:engrh,
Rostrum andwing, 8.5; antennre,

nalpi black. Antenne (male)elongate, black
throughout; flagetar segments
cytndricar with vertic's
shorter than the segments;
tirat areflagellar segments g:radually.r.lsins inde_tengrhoutwardly,tr,"i""*ii;isnrttllcr,subglobular. Head
,irirrnf""ir.'G;;#Jij"or,,/Lt(:,t.,,.(t,t:r. .l,t,.1ttt[,[,tl;.
l.t.ottt It)ost,t:r,rt, Asi,u,. V

rv, ,'

Thorax shiny coal-black, the anterior lateral pretergites very
restrictedly obscure yellow; dorsopleural membrane black. Halteres dirty white, the knobs infumed. Legs with the coxe andtrochanters yellow; femora black, the extreme bases obscureyellow; remainder of legs black. Wings (Plate 1, fig. 19) witha strong dusky tinge, the oval stigma darker brown; veins black,with long conspicuous macrotrichia, especially on the veins
beyond origin of Rs. Venation: Sc' ending about opposite
one-third the length of Rr*r-ro, Sc, at its tip; Rs long, nearlystraight; Rrur*, a little shorter than Rr*r; R, equal to Rr*r;m-cu
about one-half its length beyond the fork of M.
Abdomen black, the hypopygium a very little paler. Male hypopygium with the basistyles relatively stout, the interbasal process pale yellow, bladelike. narrowed to the acute tip. Outerdististyle entirely blackened, the apical spine much larger than
the subterminal appressedtooth. Inner dististyle more elongate,
the distal half slender, tufted apically with long stout setre.
Hubi,tat.-Formosa.
Holotype, male, Sh6rei, altitude 7,000 to 8,000 feet, October
25, L928 (,S. /sszfti)
Li.psothri,r plu.to is a striking species of the g:enus, very distinct from the only other regional member of the group.
GNOPHOMYIADEJECTAsP. nov.

General coloration plumbeous black; antenna relatively elongate, flagellar segments cylindrical; halteres and legs darkbrown; wings with a faint brown suffusion, the stigrna scarcely
darker; Rs in alignment with Ru; cell lst M, relatively elongate,
exceeding vein Mn beyond it; m-cu shortly before midlength ofthe cell; male hypopygium with the tergite subquadrate, its
caudal margin with a U-shaped emargination; outer dististyle asimple glabrous rod.
Male.-Length, about 5.5 millimeters; wing, 6.
Female.-Length, about 6.5 millimeters; wing, 7.
Rostrum and palpi black. Antenna relatively elongate, black
throughout; flagellar segments (male) cylindrical, the outer segments gradually decreasing in length, the terminal segment
about two-thirds the penultimate; verticils shorter than thesegments. Head brownish black.
Thorax plumbeous black, the posterior sclerites of the pleura
somewhat paler; dorsopleural region paler; anterior lateral pretergites restrictedly light yellow. Halteres dark brown, thebase of the stem narrowly paler. Legs dark brown. Wings(Plate 1, fig. 20) with a faint brown suffusion, the stigma

54rr

scarcely darker; veins dark brown.
Macrotrichia of veins .fmoderate length only. Venation:
Sc, ending just before R,, St.
about opposite one_third the length
L Ornr*r; Rs in alignment
withR,; r-m at fork of Rs; ce' ist M,

ion?

l".u"aingvein Mn beyond it, m-cu shorily before".iltiia,
midlength.
Abdomen dark br^own, the hypopygium
blaci
Male hypopy_
gium (Plate Z, fig.86) small. "Nt"ti, tergite 9f, small, subqua_drate, the caudal margin with a conspicuous
U_shaped emar_
gination, the subrateral lobes
thus formed acute. Basistyte, b,with the mesar margin produced intoa short acute rtacto spine.
outer dististyle, od,.a simpr",
entirery grabrous rod aris_
ing from an expanded base.'"r""4"",lnner'aististyle shorter, strongly
arcuated, the surface with several
stout spinous setre,the apex a
Iittle expanded, blackened.,glabrou.. Orripo.it;" ;;ir;hetergatvalves relatively short and high,th" basal half wider, the distal
half slightly upturned, narroried
61;" subacute tip.
Habi.tat.-Formosa.
Holotype, male, Taihoku, April
20., 1_gZZ(K. I,akeuchi).
Allotype, female, Hokuto, near
Taihoku,rFebruary
svru'r'rv ru-*-^'v*s'
10, l92g(5. Issiki,)

Gnophomaio d,eiactais a'ied to
G. oNentaris deMeijere, differing in the larger size and a"tuifs
ol-"oloration.
GNOPEOMYIA"";f:ffi:"

BREYICELLULA

Alexender.

breoicellulaAr,nxanorn,ptriiip. Journ.
sci. 22 (rszl)

The type, a unique male, was taken
at Tattaka, Formosa, alti_
tude.?,400-feet,August 16-,rgzr,
ny i"i.o Esaki. A secondmare
specimen is now on hand, co'ected
al Cnim"in, Formosa, alti_
tude 8,000 feet, July g, igzg,
ly Svuti Issiki.
The wing venation has not nu"" ngu""dand is shown herewith(Plate 1, fig. 2t).
The male f,"O-*r*iom
exhibits a rather
remarkabtestructureof what-gt, r" nintr, tu"litl ?prrtu
2, fis. 37). This supposedtergite,""*"iL
appearsas a somewhat
lyriform structure,, the long alr"i,g*t
arms being graduallywidenedto shor'y before ttre tips,
lfrJrr". ,ru""owed to the acutepoints. In the holotype,u, ngu*"A,ifr.'""*.
are sligh,yshorter"Jnu.i_"rr.
and broader than in the *o""air.olr""Ja
Outer dististyte, od,.a tong:lg"d;l "L.""tfy;;;; the rip obtuselyrounded.
Inner dististyle, zd, expandeoat
uu.e,-'--' trre
- curved apical,FOrtion
with flattenedpeglike sete.

b44.
TRENTEPOHLIAI ILC I' ILLLLI) pLtt,((MONGOMA)ATAYALJ tr llI

ILUI' U!

\><i tutLl tl t

ap. noY.

Size smali (wing, male, 5 millimeters) ; antennal verticils
short; mesonotum dark brown, the prascutum brownish yellow
with a conspicuousdark brown median stripe; legs pa'le; wings
with a grayish tinge, almost unmarked except for the stigma andan apical clouding; apical fusion of Cu, and 1st A slight.
Male.-Length, about 5 millimeters; wing, 5.
Rostrum and palpi dark. Antenne brown throughout, ofmoderate length; flagellar segrnentscylindrical, with short verticils. Head blackish gray, carinate medially.
Mesonotal p rescutum brownish yellow, with a conspicuous
dark brown median stripe that is slightly wider behind; scutum
similarly pale, the centers of the lobes extensively dark brown;
scutellum and postnotum dark brown. Pleura uniformly darkbrown. Halteres short, dark brown, the base of the stem restrictedly pale. Legs with the fore coxre darkened, the remaining coxae paler; trochanters yellow; remainder of leg:s pale
brownish yellow, without clearly defined pattern, the tarsi passing into darker; no especial armature on legs other than two
elongate setreat tips of femora. Wings (Plate l, fie.22) with agrayish tinge, the prearcular region and cells C and Sc moreyellowish; stigma elongate-oval,brown; wing apex narrowly anddiffusely darkened; restricted brown seams along vein Cu and incell 1st A at angulation of vein 2d A; veins brown. Venation:
Rs subequalto Rr*r*n; cell R, Iarge; inner end of cell M, lying
proximad of othirf cells beyond cell lst M,; m-cu at fork of
M; fusion of Cu' and 1st A slight, about one-half m-cu; vein 2d
A subangulate before midlength.
Abdomen brownish black, including the hypopygium, the central portions of the intermediate tergites paler.
Habitat.-Formosa.
Holotype, male, Inzan' near Giran, November 2, 1928 (S.
Issiki) .
Trentepohlie (M.) ataEal is distinguished from allied small
species of the subgenus by the combination of characters, as
diagnosed above.
TRENTEPOELIA(MONGOMA)TARSALBA

8p' nov'

General coloration dark brown, the posterior sclerites of themesonotum and the pleura paler; legs black, the tarsi chiefly
snowy white; wings gFayish, variegated with brovi'n seams'
including vein Cu and the fused portion of Cu' and lst A; veins
Cu, and 1st A extensively fused before margin.
Female.-Length, about 5.5 millimeters; wing, 5.2.

40,.lALcrander: Tipulitln from Eastern Asia,

V

545

Rostrum and palpi dark, the tips ofthe labial palpi yellow.
.
Antenna black throughout" Headbtack.
Mesonotunordark brown, the lateral marginsof the prascu_
tum and posterior screrites of thenotum more testaceous. preura testaceous, the dorsal pleurites
darker. galte"es short,dusky' Legs with the coxa and trochantersconcolorous withthe pleura, the fore coxa somewhat
darker; femora and tibire
black; basitarsi brown, paling to white
at iips; ,u*uirra"" oftarsi snowy white, the terminal segment
darkened. Wings(Plate l, fig.28) grayish, sparsely
variegated with brown, in_g]udins the elongate stigma; a longitudinal brown washon veinil cell M, beyond the fork or "trre vein
continued arong the
9'o{used Cu, * 1st A to the wing margin; axillary region somewhat
darkened; wing apex a lit'e sutrused;veins brown. Venation:
Rs longer than the basal section of *a,jJr",*r_*.'u"",u"ceeding R, i R, Ion_gerthanvein
powerful,
ti,
the
\*o;
medialveins beyond the fork of M "s6ak
and semievurr.*."n1; innu"
end of cell M, lying proximad of the others;
fusion oi-Cr, arra
lst A extensive, more than twice
m-cu.
.Abdomen black, the sternites somewhat paler. Ovipositor
with relatively large and conspicuous
varves, the tergar varvesd,ark, strongly upcurved, the sternalvalves high, yellow, dark
at base.
Habi,tat.-Formosa.
Holotype, female, Giran, November
5, lg}g (5. Issi,ki).
The only described species of thesubgenus Mongoia witha similar venation of the cubital andanal fields ils i. fA.l
retracta Edwards.
ORMOSIASHOREANAaD, nov.

Size relatively Iarge (wing, male,
6 millimeters) ; generalcoloration gray, the prascutum witha brown median -.t"ip";femora obscure yetow with a broad
brownish black subterminarring;_wings dusky, variegated witrr
aarrrer brown and pale areas;
cell M, open bv the atrophy of theuusur sectionni;- t"i" za
A strongly sinuous i male hypopygrurr"iwith the dististyles
sub_terminal in position, three inall simple.
Male.-Length, about b millimeters;"urnn"",
wing, 6.
Ro,strum and palpi brownish black.
Antenne of moderate.
length; basal segments pale,brown;
flagellum black, the inci_sures restrictedly pale-;-flagellar
segments gradually decreasing
outwardly, with long, uniiJeraffv
l: "iro,arranged verticils.
llotrldnrk gray.

546

The Phi,Iittpine Journa,l of Sci'ence

MePronotum brownish gray, the lateral pretergites yellow'
stripemediansonotal prascutum brownish gray, with a browntheand less evident lateral stripes; scutum gray, the centers of
postnotum
heavily
lobes variegated with darker; scutellum andpruinose.Pleuragray.Haltereswiththebasalhalfofthe
stem yellow, the distal half infuscated, the knobs paling to obscureyellow.Legswiththecoxregrayitrochantersobscureyellowl femora ob."or" yellow, with a broad brownish black
subterminal ring, the tips narrowly yellow; tibia black, the extreme bases pale; tarsi black. Wings (Plate 1, fig' 24) with astrong dusky suffusion, the costal and stigmal regions darker
brown; extensive paler areas, as follows: Before and beyond theorigin of Rs, the latter extending cephalad into cell Sc'; outerend of cell R and across the fork of M into the adjoining cells;
justaxillary region pale; veins brown. Venation : Sc, ending
beyond R,, Sc, opposite one-fifth the length of Rs; R, a trifle
longer tfr# nr*r; cell M, open by the atrophy of the basal sectionof fo"; m-cu d-fittt" less than its own length before the fork of
M; vein 2d A very stronglY sinuous.
Abdomen, including the hypopygium, black, more or less pruinose. Male hypopygium (Plate 2, fig. 38) with the ninth tergite, 9f, apparently double as in this group of species, there
being a paler and more fleshy structure beneath the true tergite,
the Ltter profoundly bifid. Basistyle, b, produced slightly beyond the level of insertion of the dististyles. Dististyles, d,lh"*, the outermost a glabrous earlike portion; longest stylegently curved to the obtuse apex, the ventral margin with pale
set with tubercles; innermost style a slender curved-"*b.urr"rod, the tip acute or nearlY so.
Habitat.-Formosa.
Holotype, male, Shdrei, altitude ?,000 to 8,000 feet, October
25,1928 (S. lsszki).
All of the Formosan species of Ormosia known to me havethis apparent duplication of the ninth tergite, there being ather}aler and thicker flattened structure imrnediately beneathplate.
outer ter,gal

ILLUSTRATIONS
llegend;procea ; i4b, basistyle;4
dististyle; dd, donal dististyle; g, gonapophysis;ri, interbasal'Oi.ri"r"r".t
inner dististyle ; 04 outer dististyle; t, ninth tergite ; ad,, wtral

Pr.a,r:o 1
Frc. 1. Nesopeza basistylato sp. nov., venation.
2, Stibad,ocerella formoseasis sp. nov., venation.
3. Stibadocerella pri,stina Brunetti, venation.
4, Cyttaromgia taiuania sp. nov., venation.
5. Limonia (Libnotes) clitelligera sp. nov,, venation.
6. Limonia (Limunia)
tenuicula sp. nov., venation,
7. Limonia (Dicranomqia) aurita sp. nov., venation.
8, Limotyia (Dicranomgi,a) montium sp. nov., venation.
9. Li,monia (Aleuandri,aria) atagal sp. nov., venation.
1-0. Orimarga fusciaenosa sp. nov,, venation.
17. Thaurnastoptera(Taiwaruita)
issikiana sp. nov., venation.
L2. Heliu,s (Helius) Liliyutanus sp. nov., venation.
13. Ula fuscistigma sp, nov,, venation,
14. Pseudolimnophila autumnalis sp. nov.,
venation.
I5. Troglophila ritozanensfs sp. nov., venation.
16. Limnophila rantaizano sp. nov., venatioh
17. Pilaria formosicola sp. nov., venation.
18. Neolimnophi,Ia alticot, sp. nov., venation.
79. Li,psothrir pluto sp. nov., venation.
20. Gnophonzgia d,ejecta sp. nov., venation.
21. Gnophomyi,a breuicellula Alexander, venation.
22. Trentepohlia (Mungoma) atayal sp. nov., venation.
23. Trentepohlia (4l[ongoma) tarsalba sp. nov,, venation.
24. Ormosia shdreana sp. nov., venation.
Pr.aru 2
Frc.25. Nesopeza basi,stglnta sp. nov., male hypopygium.
26. Limonia (Limoruia) tenuicula sp. nov., male hypopygium.
27. Limonia (Dicradomgia) aurita sp. nov., mate irypopygium.
28. Limonia (Dicrano.tttyia) montium sp. nov.,
male hypopygium.
29. Thaumastoptera (Taiwarvita) issikiina sp. nov.,
male hypopygium.
30. Helius (Helius). nawaianus sp. nov., male hypopygium,
dististyle.
3L. Helius (Helius) nipponensis Alexander, _ui"
typopygium, disti_
style.
32. Helius (Helius)- tiliputomus sp. nov., malehypopygium.
33, Pseudolimnophila autumna,lis sp. nov.,
male hypo-pyeium.
34. Limnophila rantaizan, ,p. ,rorr., mut"' t ypopygiu_. til6. Neolimnophila alticola sp. nov., malehypopygium,
116.Cnophomy'ia ilejecta sp. nov., malehypopygium.
il7. Gnophomltria breuicelluln Alexander,
rnale hypopygium.
ilt|. Onn.osia shdreana sp. nov., malehypopygium,

547

ALEXANDER:

TrpuLrDA

FR0M

EASTERN

Asrl,

V=l

PLATE I.

IPH ILT P. J ouR N . Sc r .. 40, N o.

4.

ALEXANDER:

TrpuLrDA

FR0M EASTERN

Asrl,lPHrLrP. JouRN, Scr., 40, No.

Y.l.<'d_J--$/-N
\b -

so

;>
31

al.