l -1.3
Reprintedlront theoF
THE
K.rNs.csENrolror-ocIc,rrSocrEty
JourN..rr
V o l . 4 3 , A p r i l , 1 9 7 0 ,N o . 2
pp. ll5-l2l
trtadein United Statesol America

NEW SPECIESOF CRANE FLIES FROMTROPICAL AMERICA. VI.
(Diptera: Tipulidae)'CHenr.ns P. Ar.nxaNDER
Amherst,Mass.01002
Universityof Massachusetts,

Assrnlcr
Seven new species of crane flies are described from Panama: Conomgia(Gonomgia) idiostyla, G. (G.) sejuneta, Cheilotrichia (Emped"a) appressa,
C. (.E.) brevifida, C. (E.) luteivena, Erioptera (Erioptera) interrita, andE. (Mesocgphona) latilimbata.

In the presentreport, I am continuing the discussionof eriopterine
crane flies that were taken in northern Panama by Lt. Col. Gordon
Field of the University of Rhode Island, Kingston, and his associates.
I am greatly indebted to Dr. Field for the privilege of retaining thetypes of the specieshere considered."El Volcan" is the 11,410-footVolcan de Chiriqui in northwesternChiriqui Province,westernPanama.
Gonomyi,a(Gonomyia) idiostyla, new species
General coloration of mesonotalpraescutumand scutum brownish
black, restrictedly patterned with yellowi pleura conspicuouslypatterned, brownish black, rvith a broad whitened longitudinal stripe;
male hypopygium with the outer dististyle a very long slender rod,
inner style profoundly bifid, including a long slenderouter spine,bodyof style broader, terminating in a long spine; phallosomewith apex of
aedeagusan oval pale lobe to appeargaleate.
Mer.n. Length about S-5.2 mm; wing 5.5-5.8 mm.
Rostrum and palpi blackened.Antennaewith scapeblack, remainderof organ brownish black. Head dull orange,center of posterior vertex
slightly darker.
Pronotum and pretergiteslight yellow. Mesonotalpraescutumdark
brown to brownish black, humeri and lateral margins narrowly andabruptly light yellow; scutal lobes brownish black, median area andposteriorcalli brownishyellow; scutellumobscureyellow, basebrownish
black, parascutellayellowed: mediotergiteyellow, posterior half and al Acceptedfor publication August 7, 1969.
Jounrvnr,oF rHE KnNsesENroIrrorocrcAlSocmtv 43:115-121. April, 1970.

116

JounNer,oF THEKaNsas ENrouolocrcAl Socrcry

central anterior area dark brown, pleurotergite obscure yellow, thelower margin darkened.Pleuradark brown to brownishblack, variegatedby a broad whitened longitudinal stripe extending from behind thefore coxa to the metapleura. Halteres brown throughout. Legs withall coxaebrown, trochantersobscureyellow; remainderof legs brown.
Wings weakly darkened,stigmal area scarcelyindicated,prearcularandcostal fields slightly more yellowed; veins brown. Venation: Sc short,
Scr endingjust beforeorigin of Rs, Scgshortly removed;anteriorbranchof Rs about two-thirdSRr*:rr+i m-cu closeto fork of,M.
Abdominal tergitesbrown, sternitesand hypopygiumyellowed.Malehypopygium with apical lobe of basistyle fleshy, apparently unequalin length on the two sides. Dististylesdistinctive; outer style a very
long slenderrod, its length exceedingten times the diameter,the setae
more concentrated
at outer end; inner style profoundlybifid, including
longaslenderouter spine that is nearly as long as the outer style,terminatingin a darkenedpoint, body of style much broader,extended
into a longer darkenedspine, the modified setaenear its base.Phallosomewith outer end of aedeagus
an oval pale lobe to appeargaleate;
apophysesunequal,onea long, very slenderspinethat narrowsgraduallyto the acute tip, secondapophysisshort and stout, subangularlybent
at near midlength,apex suddenlynarrowedinto a short spine.
Holotype, male, El Volcan, Chiriqui, Panama,4,200 feet, April 4,1954 (Gordon Field). Allotype, female,Cerro Punta, Chiriqui, 6,200
feet, April 20, 1954, at light over stream. Paratopotypes: 2 males,
May I l, 1954. Paratype: 1 male, with allotype.
The species'generalappearance
is much as in Gonomyia(Gonomyia)
remotaAlexander,G. (G.) salmani Alexander,and someothers,but it
differs in the male hypopygium, including both dististyles and the
galeateaedeagus.The closestrelativeis G. (G.) conniaensAlexander,of Chiapas,Mexico,with all detailsof the hypopygiumdistinct.
Gonomyia (Gonomyia) sejuncta, new species
Size medium (wing of male 5 mm) ; thoracic pleura brown witha broad whitenedlongitudinalstripe; halteresand legs brown; wings
suffusedwith brown,Sc short,Scr endinga short distancebeforeoriginof Rs, Scg at its extreme tip, cell Mz closed; male hypopygium withouter dististyle long and slender,simple; outer arm of inner style adarkenedrod, its tip obtuse; phallosomewith apex subtriangular,outerapical angle a short blackenedspine.
M^lrn. Length about4.5 mm; wing 5 mm.
Rostrum dark brown, palpi black. Antennaewith scape brown,
pedicel more yellowed; flagellum broken. Head dark brown.
Pronotum and pretergiteslight yellow. Mesonotalpraescutumuniformly light chestnutbrown, humeri and lateral bordersvery narrowly
light yellow; scutal lobes slightly darker brown, central area andVor,.43. No. 2. Apnrr,,L970

r17

posterior callositiesyellowed; scutellum brown, parascutellaand anterior half or more of postnotumyellow, the posteriorpart of the latterdark brown being the end of a conspicuouspleural stripe extending
backwardfrom the cervicalregion,ventral pleura slightly paler brown,
the two darkenedpleural areas separatedby a broad whitish yellow
longitudinalband. Halteresentirely dark brown. Legs with fore coxae
yellow, middle coxaeyellow, basesnarrowly darkened,posterior coxae
pale brown; trochantersyellow; remainderof legsbrown. Wings witha relativelystrongbrown suffusion,prearcularand costal fields slightly
more yellowed; a vague narrow darkeningalong cord, best evidenced
by the slightly darker veins. Venation: Sc short, Scr ending a shortdistancebeforeorigin of Rs, Scgat the extremetip; vein R3 only about
one-thirdthe long Rri cell Me closed;m-cu closeto fork of.M.
Abdominaltergitesdark brown, incisurespale,sterniteslight yellow,
hypopygium paler brown. Male hypopygium with apical lobe ofbasistyle slightly less than one-half as long as the outer dististyle,the latter a long slenderrod narrowinggraduallyoutwardly,apexobtuse,inner margin with a row of long setae;inner style bilobed,outer arm
or lobe a darkenedrod, the tip obtuse,lower arm terminatingin astrongdarkenedblade extendingsomedistancebeyondthe apex,apex
bearing two strong modified setae. Phallosomewith apex beyond thenarrowedneck subtriangular,the outer apical angle a short blackened
spine; apophysesvery unequal,the very long outer one a slendercurved
darkenedspine,basalapophysisvery small.
Holotype,male, El Volcan,Chiriqui, Panama,4,2OOfeet, April 4,1954 ( GordonField) .
The most similar speciesis Gonomyia (Gonomyia) salmani Alexander,describedfrom El Salvador.G.(G.) sejunctohas certainof the
hypopygial parts about the same, including the dististyles, differing
most evidently in the details of structure of both dististyles and especially the phallosome.Other generallysimilar regionalspecieshaving
a marginal spine on the outer dististyle are more distinct.
Cheilotrichi,a(Empeda) appresEa, new species
Sizesmall (wing of male about 4 mm); antennaeand palpi black;mesonotalpraescutumand scutal lobeschiefly dark brown, pleura lightbrownishyellow; knobs of halteresinfuscated;legsbrown; wingsvery
weakly darkened; male hypopygium with dististyles yellowed, outerstyle with arms appressed,the inner one a flattenedblade.
Mar.r. Length about 3.5-3.6mm; wing 3.8-4.2mm.
Fnuer-r. Length about 4.5 mm; wing 4.5-4.6mm.
Rostrum obscureyellow, palpi black. Antennae brownish black;flagellar segmentsoval. Head brownishyellow, center of vertex more
infuscated.

118

Socrnry
JounNer,oF THEKeNsnsENrovror,ocrcAr,

Pronotum brorvn, pretergiteslight yellow. Mesonotal praescutum
dark brown, humeralregionextensivelyyellowed,scutal lobessimilarly
dark brown; medianregionof scutumand posteriorscleritesof notum
more yellowed. Pleura light brownish yellow. Halteres with stem
yellow, knob infuscated. Legs with coxae and trochantersyellow;
remainderof legs light brown, outer tarsal segmentsblackened.Wings
subhyalineto very weakly darkened,stigma not or scarcelyindicated;veinslight brown. Venation: Rg slightly oblique,a little shorterthan
Rz*g*+.

Abdominal tergites dark brown, sternites yellowed, hypopygium
light brown. Male hypopygium yellowed,including the dististyles;
outer style forking at near midlength,the arms appressed,
outer arm
slender,gently curved, inner arm a broader flattened blade, broadest
at midlength; inner style nearly as long, appearingas a parallel-sided
rod, apex obtuse.
Holotype,male,El Volcan,Chiriqui, Panama,4,200 feet, May 11,
1954,at light (GordonField). Allotopotype,female,pinnedwith type.
Paratopotypes,males and females,April 7-20, May ll-IZ, 1954.
Paratypes: males,Cerro Punta, Chiriqui, 6,200 feet, April 19, 1954.
Ch'ei,lotrichi'a(Empeda) appressais generallysimilar to certainotherspecies,includingC. (8.) abi,taguaa
Alexander,differing evidently inhypopygial structure.
Chei,lotrichin(Empeda) brevifida, new species
Size small (wing of male about 3.3 mm); generalcolorationof
thorax and abdomenbrownish yellow, thoracic pleura clearer yellow;
knobs of halteres infuscated; legs yellowish brown; wings weakly
darkened,unpatterned,Sc endingabout oppositeone-fourthRs; malehypopygiumwith outer dististylehorn yellow, bearinga curved lateralspineat a right angle,apexof style shallowlynotched.
MerB. Lengthabout 2.5 mm; wing 3.3 mm.
Head lacking. Thoracicdorsumbrownishyellow,praescutumwithout well-definedstripes. Pleura clearer yellow. Halteres with stem
whitened,knob infuscated. Legs with coxae and trochantersyellow;
remainderof legs yellowishbrown, setaedarker. Wings weakly tinged
with brown, unpatterned;veins light brown. Venation: Scr ending
about oppositeone-fourthRs; vein R:r slightly oblique, about twothirdsRs*s*-r.
Abdomen, including hypopygium, brownish yellow. Male hypopygium with outer dististyle distinctive,horn yellow, including themain axis, with a more slenderbranch arising at a right angle at near
midlengthof inner margin,apexof axis very shallowlynotched,branch
gently curvedto an acutepoint.
Holotype,male,El Volcan,Chiriqui, Panama,4,200 feet, May 13,
1954,at redlight (GordonField).

Vor. 43, No. 2, Arnrr, 1970

119

Cheilotrichia (Empeda) in,strenua(Alexander), of peru, has theot1t9rdististyle of the hypopygium generally as in the present fly butwith the branchesof the inner style distinctive,the wing venationwith
Sc longer. The Mexican C. (E.) diaaricato(Alexander)is more distinct
in the hypopygialstructure.
Cheilotri,chi,a(Empeda) luteivena, new species
Ilead, includingrostrumand antennae,yellow; mesonotalpraescutumwith threebroad brown stripes,pleurachiefly yellow; wingspale yellow,
v9i1s only slightly darker, Sc long, Scr ending shortly before midlengthof Rs; abdomenbrownishyellow, tergiteswith a broad brown central
stripe; male hypopygium with outer dististyle forking into two broadslightly unequalarms, the outer broadly obtuse,inner lirtirtyle narrow.
Marn. Length about 3.5 mm; wing 4.5 mm.
- Rgstrum light yellow, palpi darker. Antennaewith basal segments
clear light yellow, outer four or five darker. Head yellow.
Prothorax and pretergitesclear light yellow. N{esonotalpraescutum
yellow with three broad brown stripes,poiterior scleritesmore uniformly
brown, posterior border of scutellum, parascutella,pleurotergiteand
Iateral borders of mediotergiteyellowed, Pleura chiefly yellor,v,with avague pale brown dorsal cloud, ventral sternopleuriteand meron more
reddened. Halteres with stem light yellorv, outer half of knob weakly
infuscated. Legs with coxaeand trochantersyellow; remainderof legs
broken. Wings pale yellow, prearcular and coltal fields clearer yellow;
veins yellow, only slightly darker than the ground. Veins beyond
generallevel of origin of Rs with long dark trichia, lacking on basalveins. Venation: Sc relativelylong, Scr endingbeforemidlengthof Rs:
vein Rs slightly oblique, nearly equal to Rs*3*a
i m-cu at fork of M.
Abdomen brownish yellow, tergites with a broad brown central
stripe; hypopygium brownish yellow, styli darker but not blackened.
Male hypopygiumwith stemof outer dististyleshort and stout, forking
into two broad slightly unequalarms, the inner a more slendercurved
horn narrowinggraduallyto the subacutetip, inner arm broadly obtuse;inner dististylenarrow throughout.
Holotype, male, El Volcan, Chiriqui, Panama,4,200 feet, April 13,
1954(GordonField).
The most similar speciesis Cheilotrichia (Empeda) ni,grolineata(Enderlein), describedfrom Costa Rica. C. (8.) appressaagreesin
the darkened pattern of the abdomen but has darker wing veins andwith the hypopygialstructuredistinct, particularly the outer dististyle.
The generallysimilar C. (8.) pubescens(Alexander) differs from allother known membersof the subgenusin the relatively numerouslong
trichia in the wing cells, thesebeing virtually as long as those on theveins.

l2O

SocmrvJounNar,oF rHE KeNses ENroNaoLocrcAL

Erioptera (Erioptera) interrita, new species
Head and pronotumwhitened,mesonotumlight cinnamonbrown,
pleurabrownishblack with a conspicuous
whitenedlongitudinalstripe;
haltereswhite; wingslight fulvousbrown,veinsand trichia pale; malehypopygiumwith both dististylessubequalin length, the inner style
expandedon outer third, thence extendedinto a long straight spine.
Mem. Lengthabout 2.5-2.7mm; wing 2.7-3.2mm.
Fuuer.n. Lengthabout4 mm: wing4.5mm.
Rostrumdark brown,palpi black. Antennaewith scapeand pedicel
black; flagellumbroken. Head whitened.
Pronotumwhitened. Mesonotumlight cinnamonbrown, postnotum
darker brown,lateral bordersof praescutumand scutumnarrowly more
infuscated.Pleurabrownishblack,dorsopleuralregionbroadlywhitened;a conspicuouswhite longitudinalstripe from behind the fore coxaeto
beneaththe halteres,slightly more widened behind. Halteres white.
Legs with fore coxae brownish black, remaining coxae and all trocrantersbrownish yellow; femora white, with three black rings, thesubbasalone narrowest,the other two broad, much wider than theinterveningpale ring; tibiae yellowed,apex narrowly white, precededby a subequalblackenedring: tarsi yellow, extremeproximal ends ofbasitarsidarkened. Wings light fulvous brown, prearcularand costal
regions more yellowed; veins yellowed, inconspicuous,trichia pale.
Venationr Rg*g**subequalto basal section of Rr; cell Mz open byatrophyof m; rn-cuerect,at fork of M; vein 2nd A sinuous.
Abdomen dark brown, hypopygium slightly darker. Male hypopygiumwith both dististylessubequalin length,outer style a flattened
blade,expandedat near midlength,thencenarrowedto an acutepoint;
inner style with basalhalf smooth,outer third more expanded,provided
with numeroussetaeand a few microscopicmarginal points, apex ofa slender,pale, gently
style a long straight spine. Each gonapophysis
curvedblade,tip subacute.
Holotype,male, Camp Pina, Panama,100 feet, January 14, 1954
(GordonField). Allotype,female,El Volcan,Chiriqui, Panama,4,200
feet,April 13, 1954. Paratypes,males,with the allotype,April 4, 1954.
polydonta AlexThe most similar speciesis Eri,optera(Eri.optera)ander,of Peru, which differs in leg colorationand in hypopygialstructure, especiallythat of the dististyles.
Erioptera (Mesocyphona)Iatilimbata, new species
Mesonotalpraescutummediumbrown, pleura darker,with a conspicuoussilvery longitudinal stripe, dorsopleuralmembranelikewise
silvery; knobs of halteresdark brown; femora obscureyellow, outerbrownishblack rings, the outer one broader,half with two conspicuous
apexwhite, tibiae obscureyellow,tips white, tarsi pale yellow,proximal
ends of basitarsi narrowly infuscated; wings strongly darkened,utr-

Vor. 43, No. 2, Annn, 1970t2lpatterned,cell Mz open by atrophy of basal section of Msi malehypopygium with dististyle tripartite, the major central arm with arelaconspicuouspale flange on lower concavemargin; gonapophyses
tively short,stronglyincurved.
Mern. Lengthabout3 mm; wing 3.2 mm.
Rostrum and palpi dark brown. Antennaedark brown; flagellar
verticilslong. Head light silvery.
Pronotumbrown. Mesonotalpraescutummedium brown, without
a distinct pattern. Pleura darker brown with a conspicuoussilvery
longitudinal stripe extendingfrom behind the fore coxae to base of
abdomen,dorsopleuralmembranesimilarly silvery white presentingthe
appearanceof two silvery pleural stripes. Haltereswith stem whitened,knob dark brown. Legs with fore and middle coxaedark brown, hindcoxaeand all trochanterspale; femora obscureyellow on basal half,
outwardly with two conspicuousbrorvnishblack rings, the outer one
broader,tip more narrowly white; tibiae obscureyellow, tips white;tarsi pale yellow,proximalendsof basitarsinarrowly infuscated.Wings
stronglydarkened,unpatterned;veinsslightly darker than the ground.
Venationl Rg*s*+lessthan twice the basalsectionof R5; cell M2 open
by atrophyof basalsectionof M.z; m-ca at fork oLM.
Abdomen,includinghypopygium,brown. Male hypopygtumwith asingletripartite dististyle,the outer arm a slenderacutespine that is
morethan one-halfthe major branchbeyondit, this with a conspicuous
broad pale flange on lower concavemargin; third arm a slenderpalerod closeto base,about one-halfas long as the style itself. Phallosome
relativelyshort,outer half stronglyincurved,directedwith gonapophyses
towardthe armsof the aedeagus.
Holotype,male,El Volcan,Chiriqui, Panama,4,200 feet, May 15,
1954 ( GordonField) .
In its hypopygial structure, Erioptera (Mesocyphona)turrialbae
Alexanderis most similar to the presentfly, differing in the details of
and in the coloration of the body,the dististyle and gonapophyses,
halteres,andlegs.