,

'

1

t

'2

,

43

New or little-knownspeciesof exotic Tipulidae(Diptera). XV
By Cunnlrs P. AIexANDER
Amherst,Massachusetts
Universityof Massachusetts,
SvNopsrs
craneflies,two belonging
of Pediciine
are givenof threenewspecies
Descriptions
to the genusU/a and one to Dicranota,all from Assam,and of five further speciesof Limoniini,all in the genusAntocha. Thematerialswerecollectedin the Himalayan
regionfromKumaonin thewestto SikkimandAssam.
THe preceding part in this seriesof papers was published in 1967(Proc. R. ent. Soc.
Lond. (B) 36 (5-6):61-68). At this time I am discussingthree further speciesin thetribe Pediciini and am beginning the treatment of the genus Antocha Osten Sacken,
vastly developedin the Himalayan region and throughout easternAsia. The materials
being considered are from the Himalayan region of India from Kumaon to Sikkim
and Assam, where they were taken by Dr. Fernand Schmid. Through the appreciatedinterest of the collector, the types are preservedin my personal collection.
ProrcrrNr
Ula (Ula) parabidens sp.n.
about2'7 mm.
Male: length.about6 mm.; wing,6'5 mm.; antenna,
Alexander,differingin hypopygialcharacters.
Generallyas in bidensAlexanderand subbidens(fig.3) with theposteriorborderof tergite(l) truncateto feeblyconvex,thesetae
Malehypopygium(p)(b) withouta setiferous
tubercleat innerapicalangle. Phallosome
relativelysparse.Basistyle
position,
points;inapophysis
lateralinner
unequaltwo
slenderstightly
with lateralarmsdividedintoasa stronghorn that narrowsgradually,the apexdirectedmesad.
appearingln bidens(fig. 2) the posteriorborderof tergite(l) is gentlyconcave,the setaemore numerous
and conspicuous.Basistyle(b) with innerapicalangleproducedinto a lobe with severallong(p) with lateralarmsappearing
theapicalspines
asnearlystraightblades.
yellowsetae. Phallosome
placedfar out on thephallosomic
complex,directedmesadto appearvirtuallysmall,innerapophysis
acrossthe mid-line.
decussate
Holotype j on microscopeslidc, INota: Sirhoi Kashong, Manipur, Assam, 7500
feet, l3.vii. 1960(F. Schmid).
UIa (Ula) bidensAlexander is known to me only from the type locality in Northeastern Burma (Kambaiti, 2000 m., taken 16.v. 1934,by Rene Malaise). UIa (Ula)subbidensAlexander is known from Nepal and Sikkim.
Ula (UIa) unidens sp.n.
thewing;thoraxdark
long,nearlyone-half
relatively
Sizesmall(wingof male5.5mm.);antennae
patterned
malehypopy'
withdarkerbrown;
restrictedly
with
wings
tingedbrown,
chestnut-brown;
gium with lateralarm of phallosome
a long flattenedyellowblade,beyondmid-lengthbent andslender,stronglysinuous.
into a longstraightspine,innerapophyses
extended
about2'5mm.
Male:length.
about5'5mm.;wing,5'5mm.;antenna,
elongate,dark brown; flagellarsegments
Rostrumand palpi brown. Antennae17-segmented,
and with a singleverticilon each,placedat nearwith short densepubescence
long,subcylindrical,
aboutone-halflongerthanthepenultimate.
terminalsegment
the basalthird,shorterthansegment;
paler
anteriorly.
Headbrown,
notal vestituresparse,erect,dark coloured.
Thorax almostuniformlydark chestnut-brown;
yellow;
obscureHaltereswith stembrown,knob brownish-black.Legswith coxaeandtrochantersremainderof legslight brown,femoralbasesyellowed. Wings(fig. l) tingedwith brown,restrictedly
patternedwith darkerbrownon anteriorcord andmorenarrowlyon remainderof cord,originof Rs,
portions. Venation:
andouterendof cell lsr M2; veinspalebrown,slightlydarkerin thepatternedbasalsectionof Ralacking,r-m joiningRsat the fork;m-cu aboutone-thirdits lengthbeyondforkof M.
Proc. R. ent. Soc.Lond. (B). 37 (3-4). Pp.43-49, 13 figs. 1968.

44

C. P. Alexanderon exotic Tipulidae

Abdominal tergitesand hypopygiumdark brown, sternitespaler. Male hypopygium(fig. 4) with
dististyle (d) with elongateblack spinoid setae. Phallosome(p) with lateral apophysisor interbasea
long flattened yellow blade, at near two-thirds its length bent at nearly a right angle and extended
into a long slenderspine;gonapophyses
appearas slenderstronglysinuousrods that narrow graduallyto the subacutetip; aedeagusstraight.

HolotypeJ, [Nole: Jhum La, Kameng,North East Frontier Agency,7800 feet,
14.v.196l(F. Schmid).
UIa (UIa) unidensis similar in generalappearanceto U. (U.) bidensAlexanderand
U. (U.) subbidensAlexander, all three differing among themselvesin hypopygial
characters,particularly of the phallosome.

3

Frcs. l-6.-(l) Venation of Ula (Ula\ unidenrsp.n. (2-4) Male hypopygiurnof: (2) Ula (Ula) bidens
Afexander; (3') UIa (Ula) parabiderusp. n.; (4) UIa (Ula) unidenssp.n. (5) Venation of Dicranota(Rhaphidolabis)subtumidosasp.n. (6) Male hypopygium of Dicranota (Rhaphidolabis)subtumidosa
sp.n. (6, basistyle;4 dististyle;r, interbase;p, phallosome;r, tergite.)

Dicranota (Rhaphidolabis)subtumidosasp.n.
General coloration dark brown; wings uniformly suffusedwith light brown, veins pale brown,
Rzns*qsubequalto basalsectionof vein R5; male hypopygiumgenerallyas in tumidosabut the shapeand proportions of the marginal and subterminallobesdifferent.
Male: lengthabout 5'5 mm.; wing, 5 mm.
Rostrum and palpi dark brown. Antennaedark brown; in unique type only 8 segmentsremain,
proximal flagellarsegmentsoval, longer than their very small verticils. Head dark brown.
Thorax uniformly dark brown, the usual praescutalstripesnot differentiated. Haltereswith stem
light brown, base yellowed, knob dark brown. Legs with coxae and trochanterslight brown, remainderdarker brown. Wings (fig. 5) uniformly suffusedwith light brown, stigmaonly vaguelymore
darkened;veins pale brown. Veins beyondcord with abundantsmall trichia, basal veinsbehind R
glabrous. Venation: Ralaaasubequalto basal sectionof vein Ri; nt-cu nearly one-half its length
beyond the fork of M.
Abdomen dark brown. Male hypopygium (fig. 6) with posterior border of tergite(l) producedat mid-regioninto a transverselyrectangularlobe that is broaderthan long, the margin with a continu-

C. P. Alexanderon exotic Tipulidae

45

ous row of long setaethat are subequalin length to the lobes; lateral tergal lobes conspicuous,rounded; just cephalad of the mid-tergal lobe a low blackened transverseprotuberance, more than
twice as wide as marginal lobe. Other hypopygial details as shown. ln tumidosa the medianrectangular lobe is much broader, being more extensivethan the compact subterminal darkened lobe.

Holotype J, Iuntl: Kujjalong, Kameng, North East Frontier Agency,4500 feet,
28.vi. 1961(F. Schmid).
Dicranota (Rhaphidolabis)subtumidosais closely related to D. (R.) tumidosa
Alexander,of Pakistanand Kashmir, differing in the hypopygial structure,particularly the tergite and interbase.
LrruoNrrNt
AntochaOsten Sacken
AntochaOsten Sacken,1859,Proc. Acad. Nat. Sci.Philad.l859: 219. Name in continuous use
from 1859to date.
Taphrophila Rondani, 1856, Dipter. Italicae prodr. I : 185. A ttonten obliturtt between 1856
and 1938.

The genusAntochais greatly developedthroughout the Himalayan region and ineasternAsia, in contrast to the condition found in a correspondingarea in western
North America,wherethe singlespeciesso far discovered, Antocha(Antocha)monticola
Alexander,rangesfrom the Yukon Territory (Whitehorse)south to Mexico (Morelos).
The charactersto be found in the male hypopygiumin this genusare of extreme
importancein classificationand have not beenemphasisedproperly. The phallosomic region is very complex,including not only the aedeagusand gonapophysesbut
also the interbase,which is a direct outgrowth from the basistyle;all thesestructures
are interconnectedto form an intricate scaffolding,as shown in the various species
here considered(figs. 8-13). Similarly,the dististylesvary greatlyin conformationin the different species,as shown. It has been believedthat the genital tube within
the aedeaguspossesseda singleterminal opening,but in some speciesthere are two
closelyapproximatedtubes,with separateopenings(fig.'13).
In the presentpaper I am discussingsix speciesand expectto treat many further
speciesin succeedingreports.
Antocha (Antocha)biobtusasp.n.
Generalcoloration of mesonotumbrownish-grey,the lateral and humeral regionsof the praescutum yellowed;antennaewith scapeyellow,remainderblack,flagellarsegmentsoval; knobs of halteres
infuscated;legslight brown; wingsmilky-grey,prearcularfield whitened;male hypopygiumwith both
dististylesyellowed, unusually broad, tips obtuse, outer gonapophyseslong and slender, inner
apophysesbroader,especiallybasally.
Male: length,about 5.2 mm.; wing. 6.4 mnr.; antenna,about 1.0mm.
Rostrum yellow, palpi slightly darker. Antennae with scapeyellow, remainderblack; flagellar
segmentsoval, slightly longer than their verticils. Head brownish-grey; anterior vertex broad,about three times the diameter of scape.
Pronotum obscureyellow, darkenedmedially. Mesonotalpraescutumwith lateral and humeral
regions broadly yellow, disk with 3 brown stripes,the broad central area vaguely divided behind,the interspacesonly a little paler than the stripes; remainder of notum brownish-grey. Pleura
obscureyellow. Haltereswith stem whitened,knob infuscated. Legs with coxae and trochanters
yellowed,remainderof legs brownish-yellowto light brown. Wings (fig. 7) milky-grey, prearcularfield whitened; veins dark yellow, poorly differentiatedagainstthe ground. Macrotrichia on distal
sectionsof veins Raas,Mrazand Ms. Venation: m-cu about one-third its length before forkof M.
Abdominal tergitesbrown medially, paler on sides,subterminalsegmentsmore uniformly darkened; sternitesobscureyellow, hypopygiumcleareryellow. Male hypopygium(ng.8) with thetergite (t) narrowly transverse,with both cephalicand posterior bordersvery shallowly emarginate.
Both dististyles(d) yellowed, unusually broad, tips obtuse. Phallosome(p) with the interbase
long-oval; outer gonapophysislong and slender,simple, much more slenderthan the inner apophysis,
the latter with tip obtuse. Margins of both the inner apophysesand the aedeagusvery pale anddifficult to delimit.

46

C. P. Alexanderon exotic Tipulidae

HolotypeJ, INota: Teng,Sikkim,4600feet,l.viii.1959 (F. Schmid).
Antocha (Antocha)biobtusais most readily distinguishedfrom other generally
similar regionalspeciesby the structureof the male hypopygium,especiallythe paledististyleswith obtusetips, and by the details of the phallosome. The pale yellow
outer dististyleis broaderthan in other regionalmembersof the genus.
Antocha(Antocha)hyperlatasp.n.
Sizesmall(wing of maleto about 4 mm.); antennaeof male moderatelylong,about one-halfthe
body or wing, scapeyellowed,remainderdark brown; mesonotumyellowed,praescutumwith 3 brown
stripes,the central one long, pleura yellow, patternedwith dark brown, especiallyon the ventral
sternopleurite;knobs of halteresbrown; legslight brown, femoral tips not darkened;wings greyish,veinsRals and Ms with trichia; abdomenbrown, posteriorbordersof segmentsmore yellowed; malehypopygium with the phallosomedistinctive, the apex of central region trilobed, gonapophyses
slender,needle-like.

f.R'
I

.t

I'

ll

r0

\)

t3

t2

Frcs. 7-13.{7) Venation of Antocha (Anrccha)biobtusasp.n. (8-13) Male hypopygium of: (8)Antocha(Antocha)biobtusasp.n.; (9) Antocha(Antocha)hyperlatasp.n.; (10) Antocha(Antocha)
incurva sp.n.; (ll) Antocha (Antocha) ophioglossodes
sp.n.; (12) Antocha (Antocha) scapularis
sp.n.; (13) Antocha (Antocha)turkestanicade Meijere (a, aedeagus;D, basistyle;4 dististyle;g, gonapophysis;i, interbase;p,phallosome;s, ninth sternite;t, ninth tergite.)C. P. Alexanderon exotic Tipulidae

47

Male: length,about 3'8-4 mm.; wing, 3'7-4 mm.; antenna.about l'8-2'0 mm.
Rostrum yellow to brownish-yellow,palpi darker brown. Antennae of male moderatelylong,
approximatelyone-half the body or wing; scapeyellow, remainderdark brown; flagellarsegments
elongate,with denseerectsetae,the sparseverticilsshorterthan the segments. Head brown, heavily
grey pruinose,cleareston orbits and the broad anterior vertex.
Pronotum buffy-yellow,cleareryellow on sides. Mesonotalpraescutumyellowed,with 3 brown
stripes,the central one long and more conspicuous;posterior scleritesof notum testaceousyellow.
Pleuraobscureyellow, conspicuouslypatternedwith dark brown on the anepisternumand extensively
on the ventral sternopleurite. Halteres with stem yellow, knob brown. Legs with coxae andtrochantersyellow, fore coxae darker; remainder of legs light brown, femoral tips and talsi notdarker. Wings greyish, prearcular and costal fields more yellowed; veins light brown. Macrotrichia on veins Rq*s,M t*z and M4 rarely at tip of Rr. Venation: vein Rz very faint, opposite toshortly beyond level of r-m;cell lst Mz normally closed,rarely open by atrophy of m; nrcu a shortdistancebeforefork of M, in casesnearlyat the fork.
Abdominal tergites brown. their posterior borders more yellowed, sternitesless evidently bicolored; hypopygiumyellow to brownish-yellow.Male hypopygium(fig. 9) with tergite(l) transverse,
central third of posterior border truncate. Dististyles (d) subterminal,outer style shaped like aboomerang, smaller and more slender than the inner style. Phallosomic scaffolding large andconspicuous,at outer end the parts united and appearingtrilobed, the centrallobe obtuselyrounded,
the lateral bladesmore produced;the scaffoldingthat surroundsthe centralarea includesthe slenderneedle-likeouter gonapophysesand the shorter interbase.

HolotypeJ, INotn: LuanglongKhunou, ManipuroAssam,2500feet, 28.v.1960(F. Schmid). Paratypes:| $ (paratopotype),with the type; severalJ, Apong, Manipur,
2430feet,l.viii.1960; 2 J, BongbaKhulen, Manipur, 2500feet,28.vii.1960;I i,
H u i a h u ,Ma n i p u r,5 0 0 0fe e t,3 .v ii.1960;I J, Mapum, Manipur ,5000feet, l2.vi .
1960; severald, Ninghti, Manipur, 2500 feet, 30.vii.1960; I dl, Sihai Khulen,
M a n i p u r,4 7 0 0 fe e t,2 5 .vi .1 9 6 0;2J, Sr inagi,Sikkim,4920feet, 6.iv.1959; I d,Dulahar,PauriGarhwal,Kumaon,4500feet,2 . vi . 1958; I d, Pulwara,PauriGarhwal,
4300feet,3l . viii. 1958(all r. Schmid).
In its generalappearance,especiallythe pattern of the praescutum,the present
fly agreeswith the descriptionof Antocha(Antocha)triangularis(Brunetti), differingin the darkenedknobs of the halteresand the femoral tips, and in the conspicuous
darkenedpattern of the thoracic pleura. The types of triangularisand its apparent
synonym,A. (A.) unilineqtaBrunetti,both from the westernHimalaya,arefemales.
Antocha (Antocha)incurva sP.n.
General coloration of mesonotunrand pleura plumbeous;antennaeblack throughout; halteres
infuscated; legs dark brown; wings brownish-grey,unpatterned; male hypopygium with ninth
sternite broad, its posterior border rounded; phallosomewith inner gonapophysesappearing as
short sclerotisedhooks. the tip incurved into a strong point.
Male: lenglh,about 5 mm.; wing, 5'7 mm.; antenna,about l'4 mrn.
Rostrum yellowish-brown;palpi brownish-black. Antennae black throughout, relatively long;
flagellar segmentsoval, with denseerect dark setaethat are somewhatshorter than the verticils.
Head brown; anterior vertexvery broad, exceeding5 timesthe diameterof scape.
Pronotum light brown. Mesonotum dark plumbeous grey, praescutum of the unique typediscolouredand it is uncertainas to whetherthere are differentiatedmarkings. Pleura dark plumbeous,dorsopleuralmembraneobscureyellow. Halteresinfuscated. Legs with coxae and trochantersbrownish-yellow;remainderof legsdark brown; clawsof male relativelyshort for membersof
the subgenus. Wings brownish-grey,unpatterned; veins light brown. Macrotrichia on distal
section of vein Ra1r, lrrore sparseon M1n2, M:t and Mq. Venation: m-cu about three-fourthsits
length before fork of M.
Abdomen dark brown. Male hypopygium (fig. l0) with tergite (r) narrowly transverse,thebreadth about 4 times the length; posterior border nearly truncate,with abundant setaeover muchof the surface. Ninth sternite(r) broad, posteriormargin gently rounded, nearly the outer half with
abundantsetae,thosenear midJine smaller. Basistyle(b) with unusuallylong coarsesetae. Outerdististyle (d) blackened,outer third narrowed, tip obtuse; inner style longer, setaeconspicuous.
Phallosome(p) with interbaseand outer gonapophysisyellowed, the latter longer, with a small
marginal point before apex. Inner apophysissubtendingthe long-oval aedeagus,each appearing
as a short sclerotisedhook, its apex incurvedat a right angleinto a strong point.

48

C. P. Alexanderon exotic Tipulidae

Holotype J, INote: Tsomgo, Sikkim, 12,500feet, in Rhododendron
association,(F.
26. viii. 1959
Schmid).
ln its generalappearancethe presentfly resemblesAntocha(Antocha\anrblystyla
Alexander,differing conspicuouslyin the hypopygialstructure,including virtually
allparts.
A ntocha (Antocha)ophioglossodes sp.n.
General coloration of thorax dark grey, praescutumwith a very broad brown central stripe;
antennaeof male relativelylong; legs brownish-yellow,claws with a singlelong nearly basalspine;
wings greyish-white,unpatterned,prearcularfield whitened; male hypopygium with outer dististyle
blackened;terminal filamentsof aedeaguslong, only slightly shorter than the base.
Male: length,about 5.5 mm.; wing, 6 mm.; antenna,about 2 mm.
Rostrum obscureyellow; palpi black. Antennaeof male relativelylong, about one-thirdthe wing,
black throughout; flagellar segmentslong-cylindrical, with densc erect whitened setae; terminal
segmentabout one-half the penultimate. Head dark brown.
Thoracic dorsum dark gley, the praescutumwith a very broad median stripe that ends abruptly
some distancebefore the suture, lateral stripes more greyish,crossingthe suture on to the scutal
lobes. Pleura dark grey, the ventral sternopleurite more blackened; dorsopleural membrane dusky.
Halteres yellow, apex of knob more darkened. Legs with coxae and trochantersobcure yellow;
remainderof legs brownish-yellow,terminal tarsal segmentsslightly darker; terminal tarsal segment
sinuous,lower face with a tuberclenear outer end; claw with a very long nearly basalspine. Wings
greyish-white,unpatterned,prearcular field whitened, basesof both anal cells weakly darkened;veins pale. Numerous macrotrichia on distal sectionsof veins Ra+sand Mr*2, with fewer on tr/s.
Venation: m-cu before fork of M.
Abdomen, including hypopygium, dark brown. Male hypopygium (fig. ll) generally as inophioglossa. Ninth tergite(r) transverse,median region of posterior border slightly convex. Dististyles(d) terminal, outer style blackened,curved on distal half to the subacutetip, inner style slightly
longer and stouter, tip obtuse. Phallosome(p) generallyas in ophioglossabut with the terminal
filamentsmuch longer, sinuous,only slightly shorter than the base.

Holotype 6f, INotn: Yagtang, Sikkim, I 1,650feet, in Rhododendrorassociation,
l7.vi. 1959(F. Schmid).
Antocha (Antocha)ophioglossodes
is related to A. (A.) ophioglossaAlexander, ofnortheasternBurma, differing evidently in the male hypopyglum, particularly theblackenedouterdististyleand the elongatesnake-tongue-likefilamentsof the aedeagus.
Antocha (Antocha)scapularissp.n.
Generalcoloration of thorax grey; antennaeblack, scapepaler; legsbrown; wings brownish-grey,
prearcular field whitened, the small stigma pale brown; male hypopygium with tergite transversc,
the outer lateral anglesproduced,conspicuouslysetiferous,median area with two slenderlobules,
interbasea small pale blade,apex obtuse;outer gonapophyses
long and very slender,about twice theinterbase;aedeagusoutwardly relativelylong and slender.
Male: length,about 5-6'5 mm.; wing, 6'5-8 mm.; antenna,about l'4-1.8 mm.
Rostrum obscure yellow; palpi dark brown. Antennae with scape yellowish-brown,clearer
yellow beneath, remainder of organ brownish-black to black; flagellar segmentsoval, exceeding
theirverticils. Head brownish-grey.
Pronotal scutum brownish-grey,scutellum and pretergitesyellowish-brown. Mesonotal praescutum light grey with 3 dark brown stripes, the broad median area ending some distance before
suture, lateral stripes obscured by pruinosity; scutal lobes dark brown, median area and base of
scutellum light grey, remainder of the latter, with the postnotum, darker grey. Pleura gtrey,dorsopleural membrane more yellowed. Halteres with stem yellow, knob brown. Legs with coxae
brownish-grey,trochantersyellow; remainder of legs brown; claws long, near base with a single
major spine. Wings brownish-grey,prearcularfield whitened;stigmasmall,pale brown; veinsbrown.
Sparsemacrotrichiaon outer ends of distal sectionsof veins Rrar ond Mt+2, and usually with a few
on Ms. Venation: Rz distinct, opposite or slightly before level of r-m; m-cu before fork of M, incasesabout to one-half its length.
Abdomen dark brown. Male hypopygium (fig. l2) with tergite (t) transverse, outer lateralanglesof posterior border produced into subtriangular lobes that are provided with long conspicuous
setae; central emargination broad, near the mid-region further produced into 2 slender lobules, thespace between these more or less convexly rounded; thickening of cephalic border of tergite andcentral area conspicuous. Ninth sternite large, with abundant long coarse setae,apex truncate or
virtually so. Outer dististyle (d) blackened, apex narrowly obtuse; inner style sliehtly longer, withlong conspicuoussetae,apex obtuse. Phallosomic scaffolding(p) with the interbasea short flattened

C. P. Alexanderon exotic Tipulidae

49

paleblade,its lengthabout 3 timesthe diameter,occasionallyonly about twicethis length,tip obtuse;outer gonapophysisdilated at base,long and very slender,about twice the interbase,the narrowed
outer end paler; central area of phallosomea flattenedplate, aedeagusrelativelylong and slender.

Holotype d, INprn: Dhar, Pauri Garhwal, Kumaon, 7220 feet, l7.viii.l958(F. Schmid). Paratypes:26, Kanol, Pauri Garhwal,8530feet, 19.viii.1958; ld,
Kulara, PauriGarhwal,12,000feet,4.viii.l958; I 6f,Pana,PauriGarhwal,8200feet,
l0.viii. 19581'2
6, Sutol,PauriGarhwal,7250feet,l8.viii.l958 (all F. Schmid).
The most similar regional speciesis Antocha(Antocha)turkestanicade Meijere,
the hypopygiumof which is shown for comparison(fig. 13); there are conspicuousin the ninth tergiteand outer gonapophysis.
differences
Antocha(Antocha)turkestanicade Meijere
Antochoturkestanicade Meijere, 1921,Tijdschr.Ent. 64 : lO7, frg. 147(d hypopygium). Fergana,Turkestan: Kirghiz S.S.R.
Antocha(Antocha)libitina Alexander,1959,Philipp../.Sci. 88 : 159-160;syn.n. Afghanistan.
Taphrophilaalbhana Nielsen, 1962, Vidensk.Meddr. dansk. naturh. Foren. lA : 167-169,fig. 3(?), fig. 4 (d hypopygium);1963,Stuttg.Beitr.Naturk.no.ll8: l-2; syn.n. Afghanistan.
Male hypopygium(fie. l3) with outer lateral anglesof tergite (r) producedinto low obtuselobes
provided with pale inconspicuoussetae,medianregion of emarginationtruncate,not producedas inappearingas a whitenedmembranousplate with
scapularis. Ninth sternitesmall and inconspicuous,abundantmicroscopicsetulaethat are arrangedin small groupsand lines. Both dististyles(d) long,outer styleslender,microscopicallynotchedat apex to appearbidentate;inner style shorter,its setae
small and inconspicuous. Phallosomicscaffolding(p) with the interbasea flattened oval blade.
Outer gonapophysisonly a little longer than the interbase,the proximal two-thirds dilated, thence
narrowedinto a spine. Aedeagus(a) terminatingin two divergentlobesthat bear the paired genitalorifices.
Antocha(Antocha\turkestanicadiffers trom scapularisin the male hypopygium,particularlyin the(figs.12, l3).
and apexof aedeagus
tergite,outerdististyle,outer gonapophysis,(Manuscriptreceived25th August, 1967)

rBof

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