,7/
Rcprinted fronr ExrouoloclcAI- NEws, Vol. LVII, No. 7, July, 1946
Printed in U. S. A.
a,UndescribedSpeciesof Crane-Fliesfrom the Western United Statesand Canada(Dipt.: Tipulidae). Part VI
Bv CseRLEsP. Ar,rxANDER,Nlassachusetts
State College,
Amherst, Nfassachusetts
The precedingpart under this title was publishedin ENrorror,oclcer,NEws,57:65-71,1946. At this time I am characterizing three further speciesfrom California arrd Idaho, all belonging to the major gellus Tiltula. Acknowledgementfor these
specimensis given under the individual species.
Tipula (Oreomyza) inyoensis nerv species
Belongsto the borealis (u,nca)group; mesonotalpraescutum
vellorv,rvith four more reddish brown stripes.the intermediatepair approximatedor confluent in front; posterior scleritesof
notunt chiefly yellorvi pleura and pleurotergiteentirely yellow;
femora vellorv, the tips narrorvly and rveakly clarkened; wings
u'ith a strong brownish tinge, lllore yellowedon basalthird, without clearly defined pattern; basal abdominal segmentschiefly
yellorv, the outer ones more darkened; ninth tergite with thernedian portion produced, its margin gently emarginate, on
either sidebeneathwith a slenderblackenedrod; inner dististylervith the beak unusually slender, blackened,extended straightbackrvardinto the dorsal crest; lateral appendagevery reduced,
sonren'lratas in ntadero,withorrt a penclulouslower processor
blackened:r clarv-likenriddle one; upper processa conspicuor.ls
t'lavate lobe; gonapaphysisappearingas a flattened blade.
3. Length about 15 mnr.; wing 13 mut.; antenna abolt 5.2mm.
9. Length about 15 mm.; wing I2.2 mm.
Frontal prolongation of head obscure yellow, paler beneath;nasus very long, tufted with long black setae; palpi obscuretestaceousyellow, the terminal segmentlnore infuscated. Antennae (male) long; scape,pedicel and extreme b.aseof firstflagellar segmentobscure yellow, remainder of flagellum black,

174

E N TO\{ OLOGICAL NEW S'46
IJtrly,the extrelnebasesof the nlore proximal segmentspaler; flagellar
segmentselongate,gently incised.longer than arry of the verticils. Head abovebrownish gr&)', clearer gra)' in front, lllore
yellowedbehind; posteriorvertex nroreinfuscated,rvith a vague
darker medianvitta.
Pronotuminfuscatedmediallyabove,obscureyellorvon sides.
Mesonotalpraescutumrvith the ground color yellor,v,rvith four
more reddishbrown stripes,the intermediatepair approximatedor confluentin front, obliteratingthe central pale vitta; outerbordersof both intermediateand lateralstripesvaguelyborderedby darker; scutumyellow uredially,the lobesextensivelygrayish
brown; posteriorscleritesof notum chieflyyellorv,the scutellurnwith a capillary brown nredianvitta, this less evident on the
scutum; mediotergitemore redclenedon sidesof posteriorthird.
Pleura and pleurotergiteyellow, unpatterned. Halteres withstemyellowishbrorvn,the extremebaseyellolr-,knob infuscated.
Legs with the coxaeand trochantersyellon'; femoravellow,thetips narrowly and u'eakly darkened,most evidently so ol1 theouter face; tibiae obscurebrorvnishyellow, the tips narrowly
darkened;tarsi black, basitarsirestrictedlypaler or1proximal
portions; claws hairy, simple. Wings with a strong brownish
tinge,more yellowedon basalthird, rvithoutclearlydefinedpattern, the prearcularand costalfields most evidentlyso; stigma
brownish yellow, vaguely borderedby darker; a scarcelyevident pattern in the basalcells,particularlyIl[ and Cu; oliiterative areasrestricted;veins brown, paler in the tltore brightened
fields. Venation: Rs loug, about three-fourthslonger than ltlcu; Rr*, entire; ln abouttwice the petioleof cell /11,.
Abdomen chiefly yellolv on basal segments,the outer orles
more suffused,or the subterminalsegmentsthe sublateralportions more darkened; hypopygium chiefly yellowish brown on
tergite and styli, the eighth sternite pale. Male hypopygiurnwith the median portion of the ninth tergite slightly produced,
its caudalmargin gently emarginate;from the lower surfaceon
either side arises a slender blackenedrod. Outer dististyle
flattened,itp apex truncate. Inner dististyle with the beak un-lvii, '461

ENTOMOLOGICAL NEWS175ausually slender,blackened,extendedstraight backwardinto thedorsal crest; lower beak very deep,its upper outer angle more
blackenedand producedinto a short point; dorsal crest with
nutneroussetae,on the outer face more emarginateand produced into a conspicuouslobe that is tufted with a few longsetae. Lateral appendage
very reduced,somewhatas in mad,era,
without a pendulouslower processor a claw-like middle one;upper processa conspicuouslyblackenedclavate lobe, its tip
obtuse;lower anglemerelytriangular,not at all produced,entirely pale. Gonapophysisappearingas a flattenedblade, thetip narrowed abruptly into a small obtuselobule. Eighth sternite very shallowlyincisedto form three low lobesthat are only
sparselyhairy, the centralone especiallyso.
Hobitat.-Cnr,rroRNrA. Holotylte: 3, Camp Manzanar,Inyo
Countl', altitucleabout 37C/0.
feet, July l-6, 1945 (Jo" M. Mayeda) ; tlrroughDr. John A. Conrstock. Allotoltotylre:J; paratofotyfc, | 3.
The presentflf is one of the interestingspeciescollectedby
Mr. I{ayeda irr the vicinitv of Camp Manzanarand kindly presentedto me by Dr. Comstock. This is a very distinct species
in an unusuallvdifficultgroup, in its almostunpatternedwings
sonren'hatresernblingTiltula (Oreom,yza)appendiculataLoew(darelicta Dietz, stala.ctoides
Doane), differing very evidently
in the structureof the malehypopygium,particularlythe reduced
lateral appendage. In this last respect it most resemblesT.
(o.) maderaDoane,which is one of the largestspeciesin thegroup, u'ith the wing pattern especiallyheavyand distinct.
Tipula (Lunatipula) alcestis new species
Belongsto the barbatagroup; allied to accurato;wings witha brownish tinge, the obliterative area before cord extensive,
virtually crossingthe wing; remainingcells not conspicuously
streakedrvith whitish; male hypopygiumwith the lobesof theninth tergite obliquelytruncated;beak of inner dististylestout,
outer basallobe broad; gonapophysisdark-colored,terminating
in an acute spine; eighth sternite with the median plate small,
conlpressed-flattened.
\

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ENTOI\TOLOGICAL NEWS

IJuly''6

J. Lengthabout 11-11.5nlrn.; n,ing 13-13.5rnrn.;atrtentta
about3.5 mm.
9. Lengthabout 1+15 rnm.; rving 13 mur.
Describedfrom alcoholicspecimens.
Frontal prolongation of head browtt, darker above; tlastls
elongate; palpi dark brorvtt, terminal segrnentsomervhatpaler.
Antennaewith scapeand pedicelyellow, the fortner a little darker
at proximal end ; basaltwo flagellar segmentsweakly bicolored,
the remainderuniforurly brownish black; flagellar segmentsonly
feebly incised,longer than the verticils. Head brorvtt.
Thorax brown, variegatedwith darker brown, includingbroadpraescutalstripes and a single major darkening on either sctttal
lobe; postnotumchiefly darkened; scutum and scutelltttnu'ith adarkened capillary median vitta. Pleura rvith the tuesepisterllum dark brorvn, the mesepitneronand uretaplettrapaler, \'ariegated with brorvn. It is probable that in dty speciuretrsthe
surfaceis more or less pruinose. Halteres rvith stetn yellorv,
knob bror,vn,its apex paler. Legs u'ith the coxae llron'u; trochanters yellolv; femora brownish t'ellou', the tips narrowlv
blackened;tibiae 1'ellorv,still more narrorvly darkenedat tip;
tarsi obscureyellon', passinginto black. Wings rvith a brorvuish tinge, the prearcularand costalfields a trifle more I'ellorved;
stigma darker brown ; cells bevond cord more strongly inftrscated, especiallyin the ottter radial field ; small brown spotsat origin of Rs and over the anterior cord; obliterative area
before cord extensiveand conspicuous,virtually crossing thewing along vein LI n, slightly broken at end of Rs; poststigmal pale area likewise conspicttous,including parts of cells
Sc, Ro and R, ; pale streaksalong veins virtually lacking,mostevident as vague lines in cells Cu, and Ist A; veins brorvn.
Venation: Rs relatively long, from about one and one-half tonearly two times nbctt, longer than in acurata. In this latter
species,the wings are filore conspicttouslyvariegatedby paler,
involving virtually the entire wing and appearingchiefly as broadstreaksalong the veins,exceptingin the outer radial field.

lvii. '-t6l

ENTO\I OLOGICAL NEWSt77

aAbdomen u'ith the segmentschiefly yellow, the tergites trivittate u'ith llro\\'11,
the lateralpair nlore broken; on the fourthand succeeding
segments
the color n-roreuniformly dark brown;
h1'popygiumchieflv1'ellorv. \[ale hl'popygiumhaving the ninth
tergite rvith broad lobes,obliquely truncated and less pointed
tlrarr in accut'ata:rnediannotch a trifle u'ider. Appendageof
ninth sternitesmaller,oval, rvith long dark-coloredsetae. Outerdististvlelong and slender. Inner dististylervith the beakmuch
stotrterthan in accurato,sloping upn'ard to the dorsal crest;
darkotrterbasallobe broaderthan in occurato. Gonapophysis
colored,relativelylong, producedirrto a slenderspine on sidelrear apex. Eighth sternite s'ith the median plate small, comllt-rtrectangtrlar.as irt accurata.
pressed-flattened,
H abitat.-[pass .

H olotJ'fr': 6f. alcoholic. Httrnphrey, Cen-

terurial\[ts.. Clarl<Co.. alorrg Rotrte 9I, altitude 6,500 feet,
Atrgrrst20, I94l (StanlevB. \Iulaik). Allotopotype:alcoholic? Paratoltot\'ltcs:| &, 1 $. alco6olic.
I am greatlyindebtedto \Ir. Stanlel'B. Mulaik for the present
materialand other interestingTipulidae from Colorado,Idaho
is Tipula (Lunaspecies
arrdUtah. The mostsimilarclescribedin thetifula) accut'ofoAlexander.n'hich differs conspicuouslyu'ing pattern and in severalinrportarrtdetailsof strttctureof themale hypopl'gium;including the tergite, inner dististylergon&pophysesand eighthsternite.
Tipula (Lunatipula) zelotypa new species
Size relatively small (wing, male, 12 mm.) ; mesonotalpraescutum gray, with four entire brown stripes; antennaewith
scapeand pedicel yellow, basal flagellar segmentsweakly bicolored;femora obscureyellow, the tips narrowly and weakly
infuscated; claws (male) simple; wings with a weak browntinge, sparselyspottedwith darker brown; a virtually complete
obliterativebandat arrdbeforecord; no trichia on squamaor in
rving cells; male hypopygium with the ninth tergite having abroadshallowV-shapednotch,the low lobesheavilyblackened;basistyleproducedcaudadinto a very thin scooplikeblade; dististyle compact,beak obtttse;two spinelikepoints on style' one

r78

ENTOMOLOGICAL NEWS'M
IJuly,

behind the beakl tn. other in the region of the posterior crest;
outer basal lobe a strong curved ann, unequally bifid at tip;eighth sternite with its caudal border very slightly produced,
provided with numerouspale setaethat are directed caudad,notarranged in brushes or pencils.
J. Length about 12 mn.; wing 12 mm.; antennaabout 4 mm.
Frontal prolongationof head brownish yellow, the dorsal surface sparselydusted; nasuselongate;palpi with basalthree segrnentsbrownish yellon', terminal one blackened. Antennae withthe scape and pedicel yellow; first flagellar segment obscurebrownish r*ellorv; succeedingsegments weakly bicolored, thesrnall basal swellings brorvn. the apices paler, the outer segments lnore uniformly dark browrr. Head above light gray,more yellowed on the occipital region; a short brown medianvitta on the posterior vertex; vertical tubercle low, entire.
Pronotum brorvnish gral'. \{esonotal praescutumgray, rvith
four narrow entire brorvn stripes, the irrtermediatepair separated by u ground vitta of more than one-half their rvidth;
lateral stripes slightly broader; humeral region narrowly bordered by dark brown; posterior scleritesof notum testaceousbrown, the scutal lobes and rnedian region of scutellum patterned with somewhat darker brorvn; pleurotergite pruinose.
Pleura pruinose behind, the anterior portion somewhat morevariegatedwith darker, especiallyon the anepisternumand ventral sternopleurite; dorsopleural membrane yellow. Halteresyellow, knob weakly darkened. Legs with the coxae yellow,
sparselypruinose, the fore pair a little darker; trochanters yellow; femora and tibiae obscure yellow, the tips narrowly andweakly infuscated, the latter more narrowly so; tarsi lightbrown, passing into black outwardly; claws (male) simple.
Wings with a weak brownish tinge, heavier along the veins
beyond cord; stigma small, pale brown, inconspicuous;obliterative band before cord large, extending from before stigma to
the posterior border along vein M n, involving lessthan the proximal half of cell /sf M,; veins brown. No squamalsetae; notrichia in wing cells, including the stigma; veins beyond cordwith relatively abundant long trichia. Venation: Rs a little

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ENTO}IOLOGICAL NEWS179

less tlran trvice nFcu i Rr*" entire; nt, oblieu€,longer than the
petiole of cell LIri tr[r*, little nlore than one-half the basal
sectionof.tr[r* " ; cell Znd A broacl.
Abdominal tergites obscureyellow, trivittate lvith brown, themedian stripe broad and distinct, narrowly interrupted at theposterior borders of the segments;lateral areas much broken,
becomingmore distinct on the outer segments;basal sternites
yellow, the outer segmentsdarker but this apparently causedby discoloration; hypopygium chiefly pale. Male hypopygium
rvith the tergite having a very broad and shallow V-shaped
emargination,the lorv lobes heavily blackened; rnedian region
rvith a microscopiclobule; outer apical portion of lobe with an
indistinct blackenedpoint or carina. Appendageof ninth sternite a small lobe provided with nulxerous short pale setae.
Basistyleproducedcaudadinto a flattened,very thin, scooplikeblade. Dististyle with the beakvery stout and obtuse,the lower
beak lacking; dorsalcrest scarcelyelevated; region of posterior
crest producedinto i short spinelikelobe; on face of style behind the beak r,r'ithall everlsrnallerreddish spine; what appears
to be the outer basallobe is a curved arm, unequallybilobed attip, the axial spine larger, the subapicalone evidently variable
in size; apical portion of arm with numeroussetae. Aedeagus
stout but sirrrple;gonapophyses
reduced. Eighth sternitetransverse,the central area of the posterior border very slightly produced, provided with numerous long pale setae that are notarrangedin brushesor pencils.
IIabitat.-CerrroRNrA. H olotypc: 3, Brawley, Imperial
C'outrty,altitucle109 feet below sea-level,April 5, 1935 (A. L.
Nfelancler); Alexander Collection, through kindness.of Dr.
Melander.
The presentisolatedfly showssolnepoints of resemblance
to
Tipula atrisurtumoDoane,which I have referredto the subgenus
Lrmatipula Edwards. In all other respects,the hypopygiumand
generalappearanceof the two flies is quite distinct. It may befotrnd that zelotyPais more properly to be consideredas being a
Tri,chotipulabut, if so, it deviatesin many respectsfrom thehitherto acceptedcharactersof the subgenus.