93 2

Popdis Auulsos de Zoologia
PlpErs Avulsos

Zoor. S. Plulo.

vol,. 22, ART.

231-243

1 5 .V r r r . 1 9 6 9ITECORDSAND DESCRIPTIONS OF TIPULIDAE FROM SOUTH
AMERICA (DIPTERA). III1
Csanlns P. Ar,rxaNnrn2

Aesrnacr
New species of crane flies belonging to the genus Teu,cholabis
Osten Sacken are described, collected in Ecuador and Peru by
Mr. L. E. Pefia. The novelties include Teucholabis (Teucholabis)
cariosw, T. Q ) inermls, T. Q.) m'egaspatha, T. Q.) perproducta,f. Q) sentosa, T. g.) wirthiano (all from Libertad, 6 km southeastof Tena, Napo-Pastaza, Ecuador), T. Q.) cinderella (from Avispas,
Peru), ?. g.) felicita, T. Q.) lipacantha, T. (f .) metatibiata andT. Q.) schisfostyla @ll from Quincemil, Cuzco, Peru).

The preceding-depart under this title appeared in March 1968
(Pap6is Avulsos
Zoologi4 S5o Paulo, vol. 2I, art. 8: 87-97).
In fhe present report I again am considering speciesbelonging to
the Eriopterine genus Teucholabis Osten Sacken, vastly developed
throughout Tropical America where more than 200 species havebeen described and unquestionably many more await discovery.
1. "Records and descriptions of Tipulidae from South America
II" was published on March 5, 1968, as article 8, pages 87-97
tDiptera).
of volum e 2L of this _iournal. The list of species contained in the article
is correctly cited in the paper's abstract, p&Be 87. I{owever, after thelast proof had been returned to the printers, in part of the issue, aline (page 92), containing the heading Teucholabis (Teu,cholabis) flauo'
was dropped and inadvertedly substituted by another one,
fimbria
reading Cruptolabis (Craptotabis) plnllostena, a name correctly appliedin page 94 to another species; in another part of the issue, a line
(page 94), containing the heading CrAptolabts (Cryptolabis) phallostena
was dropped and also inadvertedly substituted by another one, reading
Teu,cholabis (Teucholabis\ flauofimbria, a name correctly applied on
page 92 of this part of the issue.
This note is to ensure that the name Teu,cholabis (Teu'cholabis)
pageflauofi,rnbria Alexander is to be applied to the species described on
92 of Pap6is Avulsos de. Zoologia, vol. 2L (type from Quincemil, Cuzeo,
Peru, August !962, Luis E. Pefra), and that the name Cryptolabis(Cwptolabts) phallostena Alexand.er is to be applied to the species
described on page 94 of the same paper (type from Contulmo, Palo
Botado, Nahuelbuta, Arauco, Chile, February 2, 1953, Luis E. Pefia).
The Editors.
2.

Amherst,Ma,ssachusetts, 01002, U.S.A.

232

Departamento de Zoologia, 56o Paulo

The genus Teucholabi,swas proposedby Osten Sacken in 1859for asingle Nearctic species, T. complexa, a new species, occurring
throughout eastern North America. Only seven speciesof the genus
presently are known from north of the Mexican border. In the
Old World rather numerous speciesoccur, particularly in the Oriental fauna. At this time I am describing a series of species from
Ecuador and Peru that were collected by Seflor Luis E. Pef,a towhom I am greatly indebted for the privilege of retaining thematerials in the Alexander Collection of Tipulidae.
Teueholabis (Teueholabis) eariosa, sp. n.
Size medium (wing of male about 6.5 mm) ; thorax almost
ttniformly reddish yellow, abdomen black; rostrum, antennae, halteres and legs black; wings strongly darkened, more uniformly incells beyond cord; abdominal sternal pocket on segment five blackened, with numerous darkened tubercles, lateral setae relatively
small; male hypopygium with spine of basistyle yellowed, curved
into an acute blackened spine; outer dististyle a long simple rod
with about eight long scattered setae, inner style with beak bidentate; aedeagusblackened, rvith four setae, terminal spine slender.
Male
Length about 6.5 mm; wing 6 mm; antenna about 1.5 mm.
Rostrum polished black, about one-half as long as remainder ofhead, palpi black. Antennae black throughout. Head apparently
reddish yellorv.
Thorax almost uniformly reddish yellow; praescutum andscutum with long erect setae. Halteres black. Legs with coxae
and trochanters yellowed, fore coxae slightly darker; remainder ofiegs black, femoral bases very restrictedly yellowed, on fore legs
including about the basal sixth, on middle and hind pairs more
restricted; posterior tibiae near outer end with an elongate glandular area provided with abundant short erect setae, posterior
basitarsi with more than proximal third dilated. Wings strongly
darkened, more uniformly so in cells beyond cord, basal cells
vaguely streaked with paler; a narrow searcely evident darker seamover cord, stigma still darker; veins brownish black. Venation:
Sc long, Sc, ending about opposite three-fifths Rs; Rz+:r+.r about
two-thirds Rzt m-cu close to fork of M.
Abdomen, including hypopygium, black. Sternal pocket on
segment five placed far distad, subtriangular in outline, appearing
blackened from very numerous short dark tubercles, surrounding
setae relatively small; pocket on segment six with only five or six
short setae. Male hypopygium with spine of basistyle a yellow
horn that curves into an acute blackened point, the latter withlong yellow setae at base; mesal flange blackened,margin virtuallysmooth, outer angle produced into a blackened spine. Outer dististyle a long simple rod, more than one-half longer than the spineof basistyle, narrowed gradually to a microscopic point, surface
with about eight long setae arranged in pairs along the surface;
inner style with beak bidentate. Aedeagusblackened,terminal spine
slender; base with four setae, two near end of expanded partshorter, two longer setae before midlength of the narrowed portion,exceeding the latter in length.

Pap6is Avulsos de Zoolcgia,vol. 22, L969

233

Holotype, 3 , Libertad, 6 km southeast of Tena, Napo-Pastaza,
Ecuador, May 10-72,1963,Luis E. Pefia col.
Other speciesthat resemble the present fly in certain features
include T eucholabis ( T eucholabis) atripennis Alexander and T .
(f.) lara Alexander, differing in the details of the male hypopygium
and of the modified sternal abclominalpocket.
Teueholabis (Teueholabis)einderella, sp. n.
General coloration of thorax polished black, variegated withyellow; frontal prolongation of head relatively short, reddened;knobs of halteres obscure yellow; fore legs chiefly blackened,basal
lourth of femora yellowed, remaining femora and tibiae yellow, theformer with a vague pale brown subterminal ring, posterior tibia
with a small darkened glandular spot before apex; wings whitened,very restrictedly clouded with darker, more evident as a narro\t'
seam at cord; male hypopygium with spine of basistyle slender,
glabrous; outer dististyle with apical fourth abruptly narrowed intoa spine.
Male
Length about 6.5 mm; wing 5.2 mm; antenna about 1.5 mm.
Frontal prolongation of heacl reddened, short, about one-third
the remainder of head, palpi black. Antennae with scape andpedicel reddened, flagellum black; intermediate flagellar segmentswith lower face slightly produced, obtuse, outer segments shorter
than verticils. Head with vertex polished black, more reddened
beneath and on front and anterior vertex.
Pronotum yellowed, anterior end narrowly blackened. Mesonotum chiefly polished black, yellowed at suture; scutellum andanterior third of mediotergite yellowed; pleurotergite black, withvery long setae. Pleura reddened,with conspicuouslight silvery
areas on dorsal sternopleurite and metapleura. Halteres with stem
clusky, knob obscure yellow. Legs with all coxae and trochantersreddish yellow; fore legs blackened, basal fourth of femora
yellowed, middle and hind legs with femora and tibiae yellow witha vague subterminal darkening on femora; tarsi black, middle legswith proximal half of basitarsi vaguely yellowed; posterior tibiae
with a small darkened glandular area at near six-seventh thelength, proximal third of basitarsi dilated. Wings whitened, restrictedly to scarcely evidently patterned with very pale brown
along cord and over the broad wing apex, stigma dark brown;veini brown, more intensely so in the darkened areas, wing baseyellow. Venation: Sc relatively short, Sc, about opposite one-fourth
the arcuated Rs, R, at fork of Rs; vein R' deflectedstrongly caudad
to wing tip, narrowing the cell, cell R. at margin subequal tocell R,i m-cu at fork of M.
Abdominal tergites black, basal sternites yellow, variegatedwith
black, outer segmentspaler, hypopygium black. Sternal pocket on
segment five transverse,with abundant relatively short setae on allparts excepting the posterior margin; no pocket on segment six.
Male hypopygium with spine of basistyle elongate,slender,subequalin length to the outer dististyle, glabrous, narrowed gradually tothe acute tip; mesal flange reduced to a small blackened triangular234

Deparlamento de Zoologia, S5o Paulopoint, margins smooth. Outer dististyle expanded at midlength,
thence abruptly narrowed into a slender spine, tip acute, marginof expanded part with microscopic denticles; inner style with beakmicroscopically bidentate, the teeth approximated. Aedeagus withe conspicuousobtuse flange on outer half.
Holotype, 3 , Avispas, Peru, September 20-30, 1962, Luis E.
Pefia col.
Teucholabis (Teucholabis) cinderel;Ia is generally similar tovarious other J.[eotropical species, including T. (f.) azuagensis
Alexander, T. (f .) hilaris Alexander, T. (7.) laterospinosaAlexander,'I'. ( f .) neoleridensis Alexander, T. ( f .) nigrosignata Alexancler, and some others, all differing among themselves chiefly inhypopygial characters,including the spine of the basistyle ancl both
dististyles.
Teueholabis (Teueholabis) felicita, sp. n.
Size medium (wing of male 7 mm); general colorationof thorax
and abdomenblack, head paler; halteresblack, base of stem obscure
1'ellow; legs black, bases of fore femora yellow; wings whitened,
rn'ith three conspicuousbrown crossbandsancl a darkened spot at
arculus, Sc long; male hypopygium with apex of basistyle atransverse bar, its mesal end produced outwardly into a strong
spine, outer margin with numerous setae;outer clististylea simple
slenclerrod, beak of inner style bidentate.
Male
Length about 7.5 mm; wing 7 mm; antenna about 1.8 mm.
Rostrum black, relatively short, less than one-halfthe remainclerof head, palpi black. Aniennae black; flagellar ssgments oval, alit,tle shorter than the longest verticils. Head brownish black above,
lront and anterior vertex brown, beneath more reddened; vertex
with abundant erect black setae.
Pronotum obscurereddish yellow. Thorax polished black, including the pleura, dorsopleural membrane light bro'wn. Halteresblack, extreme base of stem obscure yellow. Legs with fore coxae
and trochanters reddish brown, remainder of leg black excepting
more than proximal third of femur which is yellow, proximal halfof basitarsus brourn; middle coxae black, trochanters broln,remainder broken; posterior legs entirely black, proximal third ofbasitarsusslightly dilated. Wings whitened, with three conspicuous
brown bands and a darkened spot at arculus, reaching costa at h;basal dark band extending from R to tip of vein lst A, generally
parallelsided,a little expandeddistad in cell M; band at cord broader
at stigma, narrowed on posterior cord, slightly disconnectedfrom apaler brown cloud at end of vein lst A; outer dark band including
the broad apex beyond the general level of outer end of cell lst Mr,
leaving only a small whitened spot in base of cell M", prearcularfield yellowed, including the veins; veins yellowish brown, darker
in the patterned areas, darkest at cord. Venation: Sc long, Sc'
endittg just beyonci midlength of Rs; R, *, * * about one-third Rei
m-cu one-third its length beyond fork of M.
Abdomen black throughout. Sternal pocket on seg,mentfive
heavily darkened, marginal setae numerous, of moderate length;
segment six with about ten similar setae on either side of the broadPap6is Avulsos de Zoologia, vol. 22, 1969

235

central area. Male hypopygium with apex of basistyle a transverse
bar, its mesal end produced outwardly into a strong spine, outermargin with abundant long yellow setae; mesal I'lange unusuallylarge, margin irregularly toothed, outer angle with long setae.
Outer dististyle a simple slender rod with coarse setae; inner stylewith beak bidentate. Phallosome blackened, apex of aedeagus
extended into a narrow rod, the tip slightly dilated.
Holotype, 3 , Quincemil, Cuzco, Peru, August 1962, Luis E.
Pefla col.
In its wing pattern this conspicuous fly bears a superficial
resemblanceto several species,including T eucholabis ( T eucholabis)
cybele Alexander, T. (f .) form,osissimaAlexander, T. (f .) Longis7ti.naAlexander, T. (f.) ruigrocorporis Alexander, T. (f .) rostrsta
Enderlein, T. ( f .) subpulchella Alexander, T . ( f .) tartorus Alexander, and some others, differing in details of coloration of the body,legs and wings, and especially in hypopygial structure. In thelatter regard it is more like T. (f .) bi,ramosaAlexander, especiallyin the apical spine of the basistyle, but in other respects the latter
fly is entirely different.
Teueholabis (Teueholabis) inermis, sP. n.
General coloration of thorax polished black, variegated withyellow on mesonotum and with a broad silvery pleural stripe;
knobs of halteres light yellow; fore legs dif fering in color fromthe other pairs; wings whitish hyaline, very restrictedly patternedwith brown, including the very pale apex, Sc short; male hypopygium without a terminal spine on basistyle; outer distist.vle
unequally bifid at near midlength.
Male
Length about 4.5 mm; rving 1.5 mm.
Rostrum brown, palpi black. Antennae black; proximal flagellar
segments short-oval, u'ith short abrupt apical pedicels, outer segments more elongate. Head black, heavily light gray pruinose
anteriorly, less evident behind.
Prothorax, including pleura, light yellow. Mesonotum chiefly
intense polished black, center of suture, including parts of both thepraescutum and scutum, and the scutellum light ;r-ellow. Mesopleura
broadly black above, sternopleurite similarly blackened, the twoareas separateclby a conspicuous silvery longitudinal stripe e-xtending from behind the fore coxae to base of abdomen,_widened
behind to inclucle the metapleura; dorsopleural region darkened,
only the extreme praescutal border brighter. Halteres with stem
black, knob light yteltow. Legs with coxae and trochanters yellcw;
fore femora b1ack, basal sixlh yellowed, remainder of fore legs
black; middle and hind femora yellow, tips narrowly black, including about the outer sixth or less, tibiae more obscure yellow,
tips brownish black, tarsi black, proximal end of posterior basitarsus
slightly dilated. Wings whitish hyaline, very restrictedly patternedwith brown, including the large short-oval stigma and a narrower
seam over cord to vein Cu; wing apex beyond general level of celllst M, very pale brown, scarcely apbarent; veins brownish black to
black, thole at wing base yellowed-.-Venaticn: Sc short, Sc' endiug

236

Departamento de Zoologia, S5o Paulovirtually opposite origin of Rs, branches of the latter stronely
divergent, especially R' which ends at or close to wing tip, cell R,
at margin about one-half cell R'i R, at or close to end of Rs;
m-cu immediately before fork of M.
Abdominal tergites brownish black, incisures narrowly yellowed,
especially on outer segments, hypopygium black; sternites withposterior borders broadly yellow. Sternal pockets on segmentsfive,six ancl seven, largest on five, on the others clearly indicatedlaterally. Male hypopygium without a spine on basistyle. Outerdististyle a strong rod that is unequally bifid at near midlength,
axial spine subequal in length to the base, lateral spine aboqt
one-third as long; inner style with blade long and narrow, tip
decurved. Aedeagus broad, blackened, apex a small curved blackspine and several longer setae.
Flolotype, 6 , Libeltad, 6 km southeast of Telta, Napo-Pastaza,
Ecuador, May I0-I2, 1963,Luis E. Pefla eol.
Other species that have the body and lving coloration muchas in the piesent fly and with vein Sc short include Teuch"olabis( Teucholabzs) biaci,fera Alexancler, T . ( f .) cinereiceps Alexander,and some others, all differing from the present fly in hypopygial
structure. T. ( f .) desdemona Alexander likewise is generally
similar, differing in the open cell M, of the wings and in hypopygial
structure, especially the dististyles and aedeagus.
Teucholabis( Teucholabis)

liltacantha, sp. n.

MaleLength about 6.5 mm; wing 6 mm; antenna about 1 .6 mm.
Characters generally as in perprodu,cta,sp. n., differing chiefly
in hypopygial characters.
Wing with the darkened bands narrower, the white, groundareas correspondingly widened, more extensive than the darkened
bands, dark marking at cord virtually interrupted at base of cell
ist M,. Venation: Sc, ending opposite midlength of Rs, Sc' alone
much longer, exceeding vein R, in length. Abdomen with sternal
pocket of segment five with lateral setae fewer. Llale hypopygium
wittr apical spine of basistyle relatively short, less than the outerdististyle, appearing as a slightly arcuated rod that narrows
gradually inio an acute spine, margin of basal half with abundantietae. No seconclaryspihe or spinule, as found in bigladia andperproducta.
Holotype, 6 , Quincemil, Cuzco, Peru, August 7962, Luis E.
Pefla col.
Teucholabis (Teucholabis) megaspatha, sp. n.
Allied to hilaris and rubriceps; general coloration of head andthorax orange, praescutum with a large oval black area on eitherside behind; knbbs of halteres light yellow; posterior femora withbase and apex black, posterior tibia with a modified glandular -areanear outer end; wings whitened, apex vaguely darkened, stigma
and an area at arculus darkenecl, Sc relatively long; abdominal
segments black, ringed with yellow; male hypopygium with phallo'Pap6is Avulsos de Zoologia, vol. 22, 1969237

some bearing a very large oval clarkened blade that extends someclistancebeybnd apex of the small aedeagus.
Maleantenna about 1 .5 mm.
Length about 6.5 mm; wing 6 mm;__
Head, including rostrum, orange yellow, center of posterior
black. Antennae with scape andvertex siigtrtty claiker, palpi-black;flagellar segments long'oval,
pedicel biowir, flagellum
exceeding their verticils.
Thor"ax polished orange, conspicuously patterned with black,on either side behindon the praesiutum including a large oval area'with
two contiguousthat atlains the border; each scutal lobe
brownish black marks; median region of notum, including_scutellum,
light yellow. Halteres with stem black, !.lob light _yellow. Legs
wlttr ioxae and trochanters of fore and midclle legs yellow, posteriorlegs with trochanters and tips of coxae black; fore l"gt broken;
mlOdte femora yellow, apical- fourth or more black, tibiae yellow,
tips more nariowly'blackened, tarsi brownish black; _posteriorfe^mora with basal seventh and apical fourth black, the -b-roadc"nt.ai part light yellotv, remainder bf leg brownish black to black;tibia wiltr a rioOiiieA giandular area about twice its length fromtip, proximal fifth of baiitarsus slightly dilated. Wings whitened, -tip
yiiriefy darkened, extending from vein Rr to M*i .gtiSm-adarker
fiiSwn, subcirculai', cord very narrowly - and insensibly darkenecl;
a conspicuousbro*n prearcuiar spot;-analveins brown, darker at cot'd,fielcl. Venation: Sc' ending
motJ fellowed in the prearcuiar
opposiie two-fifths Rs, in hil,aris much shorter, at about one-fourth Rs.
outer segmentswith posterior borders
Abdominal tergites black,'bicolored,bases brownish black, apicalsternites
1uriowly yelloweclj'uniformly
yellowed on outer segments, styli off,iff yetioiv, more
frypopygium^ -more black. Sternil pocket on segment five subcircular,
whitened circula-r central spot. alt setae on cephalicwittr imargin, ielatively short and abundant; ngcket of sixth sternite withaborit L5 strong- setae on either side of the broad central space.
with spine of basistyle broad ancl yellow at base,Mafu hypopysidm^nalrowed
into a long darliened point, inner margin of
nntwaritiybase witlr abundant appressedyellow setae, outer margin of spinewith microscopic tpinuies; outer mesal angle of^-style at base ofspine procluceOinto an acute black point; mesal-flange very small,fliicleireO, margin smooth. Outer dististyle a .l9ng- simple gently
tip an acute spine, surface with abundant setae,.i""o". rod, id-marginerect; inner style -stout,-terminaling in a
iho.e of innerstrong recurvecl spiie, without a distinct basal lobe. Phallosome
witft 6 ,rery- large darkeneclblade that extends some distance beyoncl
the small stenter aedeagus, the latter with a pair of setae, theuipa"OeO ntuO" with foui longer setae near center of the disk.
Holotype, 3 , Libertacl, 6 km southeast of Tena, Napo-Pastaza,
Ecuador, May 70'12,1963,Luis E. Pefla col.
The closest relatives of this species are Teucholabis (Teucho'
rubricevts Alexander, -ancl.?' (r')'differingUai{f-iiarts- llexander, r: (i:iin details of coloration andiiUr"Oir-icepi AtexinOer, alleJpecially in frvpopvgiai Jtiucture, iiclucling the phallosome. The
frototype-male oi nit7 fi formerly'was preserved in the Hungarian

238

Departamento de Zoologia, S5o Paulo

National Museum but was destroyed in 1956 during the riots in
Budapest. A paratype male is in the author's collection.
Teueholabis (Teueholabis) metatibiata, sp. n.
General coloration of entire body polishecl black; rostrum
elongate, subequal to remainder of head; knobs of halteres obscureyellow; tegF with femora yellow, tips narrowly brownish black, theamount subequal on all legs, posterior tibiae with outer two-fifths
darker and slightly mors enlarged; wings very weakly tinted,
unpatterngd except for the small light brown stigma; male hypopygium with outer dististyle slender, strongly dilaied at midlength,*itrr a- strong marginal spine, inner style #i[tr blac]gelongate, #itda) single apical spine; terminal spine of aedeagusslenderl
Male
Length about 6 mm; wing 5 mm; antenna about 0.8 mm.
Rostrum elongate, subequal to remainder of head, black, palpi
lllack. Antennae black, flagellar segments oval. Head potisheO
black.
Pronotal scutum black, scutellum yellow. Mesonotum black,hu-meral_region of praescutum and ends of suture slightly more.l'ellowed. Pleura brownish black, with a vague longitudinat paler
stripe. Halteres brownish black, knob elongate, obscure yellow.
Legs with fore coxae brown, remaining coxae ancl all trochanters
yellow; femora yellow, tips narrowly brownish black, subequal inamount on all legs, on posterior pair including about the outereighth; tibiae brown, the posterior pair yellowed, [he outer two-fifths
darker and slightly more enlarged, tarsi black. Wings very weakly
tinted, prearcular field light yellow; stigma small, short-oval, lightbrown; veins brown, slightly darker along cord, more yellowed-inthe brightened base. Venation: Sc' ending about opposite two-fifths
the length of Rs; cell lst M, elongate, nearly equal to vein M, *,beyond it.
Abdomen, including hypopygium, black. sternal pockets weak,on segment five larger but indistinct, with small setae; segment six
with a similar pale center with few weak marginal setae. Male
hypopygium of holotype with spine of basistyle broad-based,
narrowed outwardly into a blackened point, inner margin with afringe of long yellow setae, in the parbtype the spine longer ancl
trore uniformly slender; mesal flange with margin only weakly
crenulate. Outer dististyle with basal third slender, thence strongly
dilated, outer fourth suddenly narrowed, inner margin with astrong spine; inner style with blade long, with a single apical spine,
basal lobe small, with three setae. Apical spin-e of aedeagus
slender.
Holotype, 6, Quincemil, Cuzco,Peru, August 1962,Luis E. Pefia
col. Paratopotype, 6 , with the type.
The most similar species include Teucholabis ( Teucholabis)
rectangularis Alexander and T. (f .) unicingulata Alexander, which
cliffer in details of the sternal pockets and hypopygia. The modification of the posterior tibia should be noted.

Pap6is Avulsos de Zoologia,vol. 22, 1969

299

Teucholabis (Teucholabis) perprodueta, sp. n.
Allied to bigl,adia,'size medium (wing of male over G mm ) ;general coloration of thorax and abdomen black; rostrum, antennaeand halteres black; fore femora yellow on proximal half, remainder
black; wings whitened, very heavily bandeo with dark brown,
including also a conspicuous spot at arculus; abdomen with sternal
pocket of segment five triangular in outline; male hypopygium withapex of basistyle produced directly caudad into a blade that is
farther extended into a spine; outer dististyle a simple rod, innerstyle with beak large, bidentate.
Male
Length about 8 mm; wing 6.5 mm; antenna about 1.5 mm.
Rostrum black, about one-third the remainder of head, palpi
lrlack. Antennae black; flagellar segments long-oval, terminal- segment about two-thirds the penultimate. Head dark brown, paler
on genae and beneath.
Pronotum above obscure reddish brown, sides blackened. Mesothorax polished black, pleura with a small pruinose area above
the mid-coxa. Halteres black, extreme base of stem vaguely paler.
Legs with all coxae black, trochanters brown, posterior pair black;a single leg (fore) remains, femora with proximal half yellow,
remainder of leg black, proximal part of basitarsus paler. Wings
whitened, with three very conspicuous dark brown bands additional
to a large spot at arculus; basal band broad, from origin of Rs to
tip of vein 2nd A, slightly widened in cell M, second bancl at cord,
the cephalic end more intense, stigma not further differentiated, theband constricted at M; wing tip very broadly brown, including allcells beyond level of outer end of cell lst Mr, including also aslightly darker seam at outer encl of the latter; prearcular and costal
fields slightly more yellowed; veins brown, darker along cord,
prearcular veins yellowed. Venation: Sc long, Sc, ending about
opposite midlength of 'Rs, R:: just beyond the fork of Rs.
Abdomen black throughout. Sternal pocket of segment fivelarge, triangular in outline, the point directed cephalad, center withvery abundant microscopic black points, lateral setae numerous, ofmoderate size; sternite six with about a dozen scarcely modified
setae on either side of median area. Male hypopygium withbasistyle narrowed outwardly, produceclinto a blade that is extencled
into a long spine, margin of blade before the spine with relatively
few yellow setae, margin of blade at base with a microscopic erect
black spine; mesal flange conspicuous,appearing as a projecting
knob with several strong setae. Outer dististyle a slender simple
rod, narrowed outwardly, terminating abruptly in a microscopic
spine, setae long but relatively sparse; inner style with beak large,
bidentate, the teeth spinelike. Aedeagus blackened, with four setae,
the abruptly narrowed apex cylindrical.
Holotype, 6 , Libertad, 6 km southeast of Tena, Napo-Pastaza,
Ecuador, May 11, 1963,Luis E. Pefla col.
The closest relative of the present fly is Teuch,olabis(Teucholabis) bigladia Alexander which is distinguished chiefly byhypopygial structure, including the apical blade of the basistyle;240

Departamentode Zoologia, 56o Pauloin the present fly the second inner . spine. foy.tt,l in- big_ladia.is-again A third related_species
reduced to a micrbscopic blackened point.
has the apical blade ofir- ". (f .l ipacantha, sp. n., whicht,he style quite distinct, as described.
Teucholabis (Teucholabis) schistostyla' sp. n.
Allied to nocturna; general coloration of body black, thoracic
pleura more pruinose; rostrum long, subequal to remainder of head;^haltereslegs black; witlg-s
black, knob vaguely moie yellowed;-long;
mal-e hypopygium with*ftit"""d, the tip narrowly blown, Sc
rpirr" of 6asistyl6 long and slender, outer dististyle broad, profoundly
divided into tfuo undqual arms, the outer one slender, inner styleunusually narrow, without a lower beak.
Male
Length about 7 mm; wing 6 mm. _Rostlum black, unusually-long, subequal to remainder of head,-Iralpi- black. Antennae black. HeqQ _polishedblack.
Pronotum black, pretergites tight yellow. Mesonotum black,by- a faint bloom. Pleura -c-hiefly-graysurface slightly opaquepruinose, ieniral sternopleurite and anepisterlunq black, dors_oirleural membrane light y-eltow. Halteres black, knob Yery vaguelywith coxae and trochanters black, r-emai_nder
inore yellowed. Legs
-lacking)
black; posterior basitarsi dilated onof legs (fore pairnearl! the proiimal third.- Wings whitened, prearcular_field lightbrown, extending -from cell R, into M,,,
yello'iv; wing tip narrowly-veinsbrownish black. Venation: Sc long,itigma'oval,-dafk brown;
Sc," ending' about opposite three-fifths Rs; m-cu shortly beyond
fork of M.
Abdomen black throughout, surface subnitidous. Sternal pocket
the center darkened- by microscgpic setae,
on segment five large,'conspicuous,surrounding the pocket except
margiial setae black-,
at p6sterior end, segment six witlr about six or seven strong Ptt.-tsetie on either' sicle. Male hypopygium with spine of basistylelong anci slender, tip acute, blackened; mesal flange , conspicuous.
divided, out_erarm slender, inner
Outir dististyle large, profoundly-styleunusually narrow,- the lower
arm broad, its tip aiule; innerbeak lackiirg; body of style with a low lobe that terminates in awith a fey_ strong black -setae.
small blackened point, near apex
-gently
curved black hook, with- two
Aedeagus with aficat point-and atwo longer setae on back of the basal
long sltae at its baseenlargement.
Holotype, 6 , Quincemil, Crtzco, Peru, August 1962, Luis E.
Pefla col.
The only closely related species is Teucholabis (Teucholabis)which differs elpecial]V ilt
noctirna Aiexandei, of VeneZuela,'spineof the basistyle and bothtrypopygial structure, including thedististyles.
Teucholabis (Teueholabis) sentosa, sp. n.
General coloration of thorax yellowish brown above, light yellow
9n ventral pleura, above with three major black areas placed on

Pap6is Avulsos de Zoologia, vol. 22, 196924L

sides of praescutum ancl on clorsal pleurites; wings witlt the
restrictecl ground obscure yellow, disk with extensive pale brownmarkings; male hypopygium witir apical spine of basistyle a broad
blade, the outer margin serruiate; outer dististyle a slender simple
spine.
Male
Length about 6.3- 6.5 mm; wing 5.8- 6.2 mm.
Rostrum yellow, about one-half the remainder of head; palpi
and antennae black, flagellar segments oval, smaller and more
slender outwardly. Head above dark brown, beneath more brightened.
Pronotum brownish yellow. . Mesonotal praescutum mediallypolished yellowish brown, blackened in front and before suture,sides with a major polished black area; posterior sclerites of notum
obscure yellow, scutal lobes slightly more darkened, posteriorborder of mediotergite broadly black. Pleura yellow on ventral half,with a more whitened longitudinal stripe, dorsal pleura brownish
yellow with two major black circular areas. Halteres dark brown.
Legs with coxae and trochanters yellow; fore femora dark brown,basal third yellowed, tibiae and tarsi dark brorvn to brownish black;
middle and hinct femora and tibiae yellow, the latter clearer, tips
uarroWly and vaguely darkened; tarsi black, proximal end of
posterior basitarsus slightly dilated. Wings with the restricted
ground obscure yellow, including the basal fifth and a band before
corcl together with much of the costal border to beyond stigma;
remainclerof wing paler brown, including virtually all cells beyond
cord and a broad band extending from Rs to Cu across the basal
cells, cells Cu and the Anals with the exception of their bases paler
brown; stigma dark brown, conspicuous; veins brown. Venation:
Sc long, SE' ending about opposite two-thirds Rs; branches of Rs
generally parallel to one another throughout their lengths.
Abdominal tergites brownish black, basal sternites light yellow,
patternecl with brown, outer sternites darkened, eighth and ninth
segmentsyellow, styli brownish black. Sternal pocket o! segment
five longitudinally oval, with numerous setae on all sides _except
posterioi end, center of pocket without major setae; pocket ofsegment six with abotrt five or six very weak and inconspicuous
setae on either sicle directeclmesad. Male hypopygium with spineof basistyle a broacl conspicuouslydarkened blade, outer margin of
distal hait with microscopicserrulations, apex narrowed into s long
black spine; margin at base of spine with a pencil of long yellow
setae, the longest virtually equal to the spine, the row extencled
basad on face-of blacle for about one-half its length; mesal flange
clarkened,margins smooth; setae at apex of bas_istyleblack, -very
long, near:ly equal in length to the blade. Outer dististyle aslencler simple yellow spine that is nearly as long as the blade ofstightly ditated at near midlength, thence narrowed
the basistyl-e,.slenderpoint, surface with very sparse small setae;
into a long
tip of inner style cbnspicuously bidentate. Apex of aedeagus
narrowed to a cylindrical blackened rod.
Holotype, 6 'Luis, Libertad, 6 km southeast of Tena, Napo-PastaVa,
E. Pef,a col.. Paratopotypes,3 6 6, with the
May 10-li,-1963,

242

Departamento de Zoologia, 56o Paulotype. Paratypes, 2 6 6 , Quineemil, Cuzco, Peru, August 18 andOctober 15-20,7962,Luis E. Pefla col..
In its general appearance Teucholabis ( Teucholabis) sentosa
suggestsspecies such as T. (f.) atripennis Alexander and T. (f.)
tnEthica Alexander, but is quite distinct in hypopygial structure,
particularly the outer dististyle and armature of the basistyle.
Teucholabis (Teueholabis) rvirthiana, sp. n.
General coloration of body yellow, including the short rostrum;
mesonotal praescutum with a conspicuous subcircular black spoton either side; halteres and legs yellow; wings light yellow,
prearcular and costal fields more saturated, stigma brown, vein Sc
long; male hypopygium with spine of basistyle distinctive, apicallyrvith two long points; outer dististyle with a small lateral spine,
inner style without a lower beak.
Male
Length about 6 mm; wing 5.3 mm; antenna about 0.9 mm.
Rostrum light yellow, relatively short, slightly more than
one-third the remainder of head, palpi obscure yellow, indistinctly
patterned with light brown. Antennae short; scape and pedicel
light yellow, flagellum light broln, incisures of the more proximal
segments narrowly pale, outer segments darker brown; flagellar
verticils much exceedingthe segments,especiallythe more proximal
ones. Head light yellow, the broad vertex light gray, its center
weakly more darkened.
Prothorax yellow. Mesonotum fulvous yellow, praescutum oneither side with a single subcircular black spot at more than its
own diameter from margin, central region of praescutum vaguely
more fuscous. Pleura almost uniformly yellow. Halteres paleyellow, knobs more saturated yellow. Legs with coxae and trochanters yellow; remainder of legs yellow, fore femora with extreme tip
light brown, outer tarsal segments more infuscated; posterior
basitarsi with proximal sixth slightly dilated. Wings light yellow,
prearcular and costal fields somewhat more saturated, stigma
small, oval, brown; veins yellow, m and veins comprising the cclrd
brown. Venation: Sc relatively long, Sc' ending about opposite
two-thirds Rs, cells R:r and R+ s{, margin subequal in extent.
Abdomen yellow, the segments vaguely more darkened basally,t he pattern produced by dense microscopic black setulae. Sternal
pocket of segment five indicated by a grouping of about a dozen
long setae on either sicieof a narrow central unmodified area, setae
darkened basally, outer ends pale, all such setae contiguous or
decussateat midline; no setal modifications on sixth sternite. Male
hypopygium with spine of basistyle distinctive, glabrous, broacily
flattened, outer end slightly expanded, terminating in two longacute black points, the axial one slightty longer; mesal flange
undeveloped. Outer dististyle a pale rod, at near two-thirds thelength with an acute subappressedspine; inner style with beakblackened, elongate, cultriform, without a lower beak or clearly
defined basal lobe. Aedeagus with base dilated, with four setae,
apex extended,tip blackened,obtuse and weakly bilobed.

Pap6isAvulsos de Zoologia,vol. 22, 1969

243

Holotype, I , Libertad, 6 km southeast of Tena, NapoPastaza,
Ecuador, May L0-L2,1963, Luis E. Pefla col..
I dedicate this species to Willis W. Wirth, outstanding studentof the Diptera, co-author of the invaluable Catalog of the Di.ptera of
Amenca north of Mertco (Washington, 1965). In its generalappearaneeTe'rrtholabi,s(Teucholnbis) wirthi,ono is similar to f . (f .)jocosa Alexander and some allied species, differing in all detailsof the male hypopygium, particularly the basistyle and outerdististyle, as well as in the very different sternal pocket. The
bispinous terminal spine of the basistyle suggests the condition inf.(f.)
bi,ramn'saAlexander, ?. (f.) scabrosoAlexander, and ?. (f.)
spica Alexander, all quite distinct in other features.