STIIDIES ON TEE CRAITE-T'IIESOF MEXICOPART Vt(ffier Diptera, SuperfamilyTipuloidea)BT

Cmnms P. Ar,Bxarvonn

REPRINITD

FROU

Axrers oF TEBExlorrorocrcer, SocIEly on Arenrcr
Vol. XXXII, No. l, March, 1939

STIIDIES ON THE CRANE-FIIES OF MEXICOPART VII(Order Diptera, Superfamily Tipuloidea)
Cnenlrs P. AuexlNDER,
Amherst, Massachusetts

The species of Tipulidae discussed in this report were allcollected by Dr. Alfons M. Dampf, Government Entomologistof Mexico, in the State of Chiapas. The localities and conditions
under which the various species were taken have been more
fully discussed under the preceding part under this general title(Annals Ent. Soc. America, 31 : 393 ; 1938). I am greatly
indebted to Dr. Dampf for the privilege of retaining the types of
the novelties described herewith.
Limonia Meigen
Limonia (Geranomna) civica sp. n.
yellow, the lateral margins darkened, !h" disk
Mesonotum buffy-brownstripes that are slightly wider than thewith three narrowinterspaces; rostrum relatively short, -a little more than one-third
the length'of the body; pleura virtually unmarked.i femora with arelatively narrow brown subterminal rittg; wings yellow, restrictedly
patterned with brown, inch,rding a-common area at fork of Sc andbrigin of Rs,. male hyloRlgium with the rostral lPines of the ventral
d.islistyle placed.on face of style and very. dissimilar .in form,_bothfrom conrpicoour basal tubercles; inner_spine reduced to a slendersetoid.strubture; gonapophyseswith abundant setulaeon their expandedbasal portion.
Aile.-Length, excludingrostrum, about 8 mm.; wing, 8.5 mm.;rostrum, about 3 mm.
Rostrum black, a trifle longer than the combined head and thorax.
Antennae black, relatively long; flagellar segments_orralto long-oval,
ihe verticils subequal in iengtfi to the segments. Head dark with apalecentral median vitta.
'
Pro1otum butry yellow, with three brown stripes. Mesonotal
praescutum buffy feilbw,- with three narrow brown stripes, alditional[o ttr" darkened fateral borders behind the yellow humeral {9gio1;
median dark vitta a little wider than the laterals; interspacesslightly
than the stripes; scutal lobes infuscated; median atea of
1"*o*"tr.ot,t* and. the scutellum pale; postnotum dark brownish plumbeous.
(Vol.
rThe- preceding- part under this general title.app*rtgd in these A:rnals{gg-+tZ, l'g38). Contribution from the Entomological Laboratory,
XXX];
Massachusetts State College.

701e3elAlexander: Mexican Crane-fies

7r

Pleura pale yellow, vaguely darkenedon thg anepisternum. Haltereswith th^esteh pale, especiallyat base,the knobs dark brown. Legs
with the fore cbxae dark brown, remaining coxaeand all trochanters
yellow; femora obscure browr_rishyellow, with.- a relatively narrowbrownishyellow,
6towtt ring about equalto the clearyellow apex;tib_i_?e
the tips nlarrowlydarkened;tarsi light brown.,, Wings with a yellow
tinge,-restrictedlypatterned with brown, as follorys:-1{common atea
at flik of Sc and-oiigin.ofRs; cord and outer end of cell lst Mzi stig_ma;
supernumerarycrossvelnln cell Sc; very sqal! spots near h and at
tif of vein nr; a paler cloud at end of vein 2nd A;veins ygllqw' slightly
dirkened,in the cloudedareas. Venation:,Scshort, both Scr and Scz
ending just beyond the origin of Rs,' Rs long, arcuated, nggly threeceli 1stMz shorterthan any of the veins beyon+it.
times"^Rz+gi
Abdominal tergites brown, sternites yellow; hypopygium "-biqfy
pale. Male hypopygiumwith the tergite-narrowlynotchedmedially,tt" lobes broidly obtuse and with numeroussetae. Basistyle vgry
much smallerthan the long-ovalventral dististyle; rostral prolongationof the latter long and slender,the usual spinespl.ac9d.onface of style
below the baseof ttte prolongation;spinesvery dissimilarin.fo*,_theouter stout and. straifht, from a balal tubercle that is subequal inlength and mergesgradually into the .spine;inner spine reducedto asleider seta that is nearly as long as its much stouter basal tubercle.
Dorsal dististyle a short, stout, stlongly cunred hook. - Ggnapophyses
with the expandedbasal portion denselyprovided with short setulae;
mesal apicai lobe darken-ed,narrowed gradually to the acute black
tip. Anal tube with few coarsesetae.
Holotype, d, Vergel, Chiapas, altitude 800 meters, above awaterfall in a damp valley, at light, June 1, 1935(A. M. Dampf);
M. F. 4384.
The most similar species are Limonia (Geranomyia) eurygrarnfna Alexander (N{exico-Panama) and L. (C.) recondita
Alexander (Peru), both of which differ in the wing-pattern, colorof thorax and structure of the male hypopygium. The femoraof the above-mentioned species are uniformly darkened, notannulate as in the present fly.
Limonia (Geranomyia)uberis sp.n.
Sizevery large (wing, d, 10 mm.); mesonotalpraescutumbrownish
yellow with thiee narrow brownish black striqes; femora obscureyellow, with a broad dark brown subterminal ring; ying-s with theground color light yellow, very heavily patterned with brown, the6eilsbeyond the cord almost uniformly darkened;a-commondark areaat origin of Rs and fork of Sc,'Scr ending shortly before midlength of,Rs,.male hypopygium with the ventral dististyle large, with. a single
weil-deneloied spine from a long curved base, the secondspine rudimentaryl anal tube denselyhairy.
Maie.-Length, excludingrostrum, about 9 mm.; wing, 10 mm.;rostrum, about 4 mm.

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Annals Entomological Society of America [Vol. XXXII,

Rostrum black. Antennae black throughout. Head dark blackish
gray_,with a narrow median gray line on veitex.
Pronotum black. Mesolgt"l praescutumobscurebrownish yellow,
with three narrow brownish blackltripes, the median one ending befote
the sutureI scutal lobes extensivelybiownish black; scutellumibr",tt"brownish yellow; postnotum brownish yellow,' darker behind. Pleuia
weakly infuscated, the sternopleurite paler. Halteres yellow. Legs
with the coxaeyellory,the fore coxaemore darkenedbasallir;trochante?syellow; feryog obscure yellow, with a broad. (0.g mm.i dart< brown
subterminal ring, preced.{ by a subequalclearer yellow iing, the apex
narrowly (about.O.2_mm.)of the latter color; tibiae brownjih yeilbw,
the -tips _narrowly darker;_tarsi obscure yellow, the outer selments
blackened. wings *ilh the ground coloi light yellow, chieflj, concealed by. -an unusually extensive brown patlernl ce[s' beyond cord
almost uniform_lyd_arlened,variegated by small ground "t""r in cells
R2,lst Mz and Me; bef.orethe-cord,the yellow ground is more extensive,
subequal in area to the dark, more evident near the wing-baseand
jult before the cord; a large common dark area involves"both thegtigi" of_Rsand fork of Scfenclosinga small central pale spot; darkband at level of .the :ypeglumerary.crossveinin cell ,.Sr,"otrsiderably
widened behind in cells M, Cu and lst A, the last-namedcell dart
exce?t at base;veins dark, yellow in the ground areas. Venation:,Screlatively_l9rg,scr.endingshortly beforemidlengthof Rs, sc2at its tip;cell /sl M2longerthan any of the veinsbeyondit.
Abdominal tergites br_own,the sternites somewhatpaler, yellowish
proyrn;.hypop-ygium_chieflypale yellow. Male hypopysium with thebasist_ylemuch smaller than the ventral dististyie-,i6J ventro-mesal
lobe dusky, extensive. Ventral dististyle with th-erostral prolongation
short and stout, with-a single well-developedspine, this irisingifrom
3 1o1Scurved swollenbasethat is about two-thiids as long as th6 spine
itself ; a secondrydjmgntar-yspine is present,so small as tlo be scar'celyevident, subequal in length to the smaller setae near it, arising froma small swollen 9?.9, the apex .of the spine truncated. Gonapolhyses
with mesal-apicallobe terminating in an-unusually slender,pale, iurved
point. Anal tube denselycoveredby denseblack hairs, a^aaitionalto
the usual coarsesetae.
Holotype, c/, vergel, chiapas, altitude 800 meters, at light,
1 0 P . M . t o 6 A . M . , M a y 1 3 , 1 9 3 b ( A . M . D a m p f ) ; N f. F . 4 1 b 8 .
Limon'ia (Geranomyia) uberis is so distinct from all otherdescribed species of the subgenus that comparison with otherforms is unnecessary. The combination of large size,unusually
heavy wing pattern, and the structure of the male hypopygium
furnish strong specific characters.
Limonia (Geranomyia)trichomera sp. n.
General coloration of mesonotum reddish brown, the praescutumwith four narrow silvery lines, the intermediate ones on eiiher side ofa slightly wider brownish black median vitta; femora with a narrow1eselAlexander: Mexican Crane-fl.ies

73

brown subterminal ring; wings subhyaline, sparsely patterned withdarker, including a series of four small costal areas; Sc long; malehypopygium with the gonapophyses blackened, their surface with
abundant short erect setulae.
Male.-Length, excluding rostrum, about 6.5 mm.; wing, 7 mm.;rostrum, about 3 mm.
Rostnrm black, nearly one-half the length of body; maxillary palpi
unusually long. Antennae black throughout; flagellar segments suboval, with truncated ends. Head black, the narrow anterior vertex
silvery, less distinctly prolonged backward onto the posterior vertex;
posterior orbits narrowly gray.
Pronotum reddish brown, darker laterally. Mesonotal praescutum
reddish brown, with four narrow silvery lines, the intermediate pairon either side of a slightly wider brownish black median vitta, thelateral stripes about mid-distance to the lateral margin; scutum andscutellum reddish brown, with a continuous median silvery line; postnotal mediotergite reddish brown. Pleura pale brown dorsally, paling
to yellow beneath. Halteres with stem pale, knob dusky. Legs withcoxae and trochanters yellow; femora yellow, with a narrow brown
subterminal ring, the slightly more extensive tip yellow; tibiae andtarsi brownl claws with a group of three or four teeth close to base.
Wings subhyaline, sparsely patterned with darker, including a seriesof four small costal areas, the largest stigmal; second area at originof Rs, third at fork of Sc, the latter two widely separated; very narrow
to scarcely evident seams along cord and outer end of cell l st Mzi
veins pale, darker in the clouded areas. Venation: Sc long, Sc1ending
shortly beyond midlength of Rs, 5c2at its tip; Rz a little shorter thanRr+zi cell .1st Mz subequal to vein Mpz beyond it; ,n-cxtclose to fork
of.M.
Abdomen dark brown, the hypopygium very slightly paler. Male
hypopygium with the ventro-mesal lobe of basistyle relatively long,simple. Ventral dististyle large and fleshy, much more extensivethan
the basistyle; rostral prolongation short but relatively stout, shorter
than the two rostral spines,theselatter arising from a small pale tubercle
close to base of prolongation; each spine nearly twice as long as theprolongation itself. Gonapophyseswith mesal-apical lobe blackened,high and erect, the surface with abundant short erect setulae, the tip
further produced into a long, gently curved spine.

Holotype, d, Finca Belem, Chiapas, altitude 850 meters,
J u n e 2 3 , 1 9 3 5( A . M . D a m p f ) ; M . F . 4 6 1 0 .
Limonia (Geranomyia) trichomera is entirely distinct fromother described species of the subgenus in Tropical America.
Elsewhere in this fauna, the setuliferous gonaphyses are foundonly in L. (G.) umbricolor Alexander, of.Brazil.
Limonia (Rhipidia) sejugatasp. n.
Allied to mul,tif,da,'antennae(male) with ten bipectinate flagellar
segments,the branchesshort; thorax.-d"t\ brown, the plg-uraand aconspicuouspale triangular area on side of praescutumyellow; wings

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Annals Entomologicol Society of America [\'o1. XXXII,

yellow, restrictedly pattetned with small brown spots; abdominal
tergites bicolored; male hypopygium with the ventraf dististyle small;rostral prolongation slender,with a single strong curved spine before
midlength.
Male.-Length about 5.5 mm.; wing, 6 mm.
Rostrum and palpi black. Antennae with scape,pedicel,terminal
segmentand all flagellar branchesblack, the pedicelsof the flagellar
segmentswhite; flagellar segmentsone to ten, inclusive, bipecfnate,
the longestbranchesabout one-halflonger than the segmentslterminai
segmentexceedingthe penultimatein length. Head dark gray; anteriorvertex very narrow.
Pronotum dark brown. Mesonotal praescutum dark brown infront and again on posterior third, the central portion paler, the entire
lateral region of the scleriteoccupiedby a major yellow pollinosearea
that also involves almost all the pleura, contrasting conspicuouslywith the dark color; posteriorscleritesof notum dark brown, the medianarea of scutum narrowly pale. Pleura as described,yellow, the propleura, ventral sternopleurite and pleurotergite dark biown. Halt-eresyellow. Legs with the fore coxaeand trochanters light yellow; remainltg coxae and trochanters conspicuously dark brown; remainder oflegs light yellow, the terminal tarsal segmentsblackened. Wings
with-brownthe ground color yellow, restrictedly patterned with small
spots, as follows: Origin of Rs,' tip of Sc,' outer end of the otherwise
chiefly pale stigma; cord and outer end of cell / st Mz; small marginal
spots at ends of veins Rg to ?nd A, inclusive, slightly larger on thelast-namedvein, smalleston R+ri proximal end of usual stigmal areawith a weak dait< cloud, the central portion pale, "r a"r"tifia; veinspale, darker in the clouded areas. Venation: Sc, ending opposite orjust beyondmidlength of Rs; cell lst Mz longerthan vein Mabeyondit;
m-cu at or shortly beforefork of.M .
Abdominal tergites bicolored, brownish yellow, the caudal bordersof the segmentsnarrowly dark brown; sternites more uniformly yellow,
the subterminal segmentsdarkened; hypopygium pale. Male hypopygium with the tergite rather deeply emarginate,eachlobe terminating
in four or five fasciculate setae, the margins with numerous normil
setae. Basistylewith the setaeof ventro-mesallobe of normal length;
face of style near base of lobe with a group of erect setae. Dorsal
dististyle a very strong, curved, slenderhook, at apex narrorvedinto along straight point. Ventral dististyle unusually small, subequal inarea to the basistyle; rostral prolongation long and slender, before
midlength with a singlestrong curved spine.
Ilolotype, C , Finca Victoria, Chiapas, altitude 900 meters,
June 29, 1935 (A. M. Dampf); M. F. 1710. Paratopotypes,2& & ; paratype, d , Vergel, altitude 800 meters, June 10, 1935;

M. F. 4503.
Most nearly related to Limonia (Rhipidia) mulrirtdaAlexander (Mexico), differing in the coloration, the short flagellar
branches,and in the detailsof structureof the malehypopygium,

1e3elAlexander: Mexican Crane-fliesID

especially the more shallowly emarginate tergite, with fewer,
more flattened setae on lobes, the different structure of the lobeof basistyle, and the small ventral dististyle, with strongly
curved rostral spine.
Limonia (Rhipidia) hirtilobata sp. n.
General coloration of mesonotum dark brown, the pleura and broad
lateral praescutaltriangles yellow; antennae(male) with nine bipectinate
segments,the branchesof moderate length; fore coxaepale, middle andposterior coxae black; wings with a strong brownish tinge, with agfeatly reduced darker pattern ; rn-cu at or shortly beyond the forkof M ; male hypopygium large and complex in structure; tergite andlobes of basistyles with unusual development of setae; rostral spines
two, unequal in length and thickness.
Male.-Length about 5-5.2 mm.; wing, 5.5-5.8 mm.; antenna
about 2 mm.
Rostrum and palpi black. Antennae relatively short, as shown by
the measurementi; basal three segmentsdark, flagellar segmentswithbasal enlargementsand branches black, apical pedicels pale, terminal
segment uniformly darkened; flagellar segmentstwo to ten, inclusive,
bipectinate, the first and eleventh merely short-produced; terminal
segment simple; longest branches short, approximately twice the segments. Head dark brownish gray; eyes contiguousor nearly so.
Pronotum infuscated. Mesonotal praescutum with the sidesextensively yellow, sparsely pruinose, the median ?rea chiefly 99vgre{.!,y_t
cential brown stripe that is most intense in front and behind, slightly
more reddish at midlength; posterior portion of praescutumand posteriorscleritesof notum almost uniformly dark brown. Pleura with propleuraand metapleura darkened, the extensive mesopleurapale yellow, confluent with the pale sides of praescutum. Halteres with stem pale,knob infuscated. Legs with the fore coxae pale, middle and posterior
coxae black; trochanters obscure yellow ; remainder of legs broken.
Wings with a strong brownish tinge, with a greatly reduced darker
pattern, including very small, inconspicuousareasat origin_of Rs, fork
bf ,Sc,cord and outer end of cell lst Mz; stigma bicolored,the proximal
portion pale, scarcely differentiated from the ground, the outer end
darkened at Rzi veins pale brown. Venation: Sc relatively short,
Scr ending opposite or just beyond one-third the length of Rs, Sc2at
its tip; cett 7il Mz closed, subequal in length to vein Ma beyond it;
rn-cu,at or shortly beyond the fork of.M .
Abdominal tergites dark brown, the caudal borders of the segments
narrowly more brownish black; sternites and hypopygium yellow.
Male hypopygium unusually large and very complex in structure.
Tergite shallowly notched medially, each lobe sfliquely truncated;caudal margin of lobes with modified setae, including a more mesalgroup of about eight long flattened setae,and an outer fascicqlate.group
Chatbecomestwisted at near one-third their length. Basistyle with theventro-mesal lobe conspicuous,at apex with a group of very long palesetae, on mesal face of basistyle just caudad of the lobe with a low
blackened flange. Ventral dististyle moderately large, the rostral

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Annals Entomological Society of America [Vol. XXXII,

prolongation stout at base,at near midlength narrowed and flattened;
two unequal rostral spines at near one-third the length of the prolongation, the more basal a short straight spine from a low base,thesecondspine placed at base of the first, more slender and more thanwith mesal-apicallobe slender,the tip
one-halflonger. Gonapophyses
slightly recurvedand narrowly blackened.
tlolotype, &, Finca Victoria, Chiapas, altitude 900 meters,
June 29, 1935 (A. M. Dampf); M. F. 4710. Paratopotypes,

2 & &, June 28-29, 1935;M. F. 4693, 4710.
Limonia (Rhipidia) hirtilobata is one of the most distinct
speciesof the subgenusso far made known, the structure of themale hypopygium being quite unique among the described
forms. In coloration, the fly comes closest to the otherwise
amply distinct L. (R.) sejugatosp. n.
Shannonomyia Alexander
Shannonomyia
dampfisp. n.
Generalcolorationgray, the praescutumwith three brown stripes;
antennae(male)elongate,nearlyas long as the body; flagellarsegments
cylindrical,with long coarseoutspreadingsetae;headclear light gray,the broad anterior vertex silvery; legs brownish yellow, the outertarsal segmentsdarkened; wings with a brownishtinge; macrotrichia

of cells very reducedin number; cell Ist Mz closed;male hypopygiumwith the gonapophysesmassive, blackened,each with from three tofive coarsespines.
Male.-Length about 6 mm.; wing,6 mm.;antennaabout 5 mm.
Rostrum pruinose;palpi black. Antennae(male) very elongate,asshown by the measurements;scapeand pedicelpale yellow, flagellum
black; flagellar segmentscylindrical, with long coarse erect setae
distributed throughout the length of the segment,the longest about
two-thirds the length of the segmentsI terminal segment about onethird the length of the penultimate,with setaeat apex. Head clear
light Eray, more silvery on the front and wide anterior vertex.
Pronotum pale, sparsely pruinose, darkened laterally. Mesonotal
praescutum gray, with three brown stripes; scutum dark brown;
scutellumand a basaltriangle on mediotergitegray,the posteriorlateralangles of latter darkened. Pleura and pleurotergite dark brown
dorsally,the sternopleuriteand meron paler. Halteresdusky, the baseof stem very restrictedly paler. Legs with the fore coxae weakly
darkened,the remaining coxae and all trochantersyellow; remainderof legs brownish yellow, the outer tarsal segmentsdarkened. Wings
with a brownish tinge, the small stigma a trifle darker; prearcularandcostal regionsa little darker than remainderof ground; veins brown.
About three macrotrichiain extreme outer end of cell Rc. Venation:
Ser ending a short distancebeforethe fork of Rs, Scza short distance
before tip; Rs relatively long, nearly three times Rz+a+riRz faint,
subequalto Rz+a,'Rr+, a little shorter than Rz+e+riveins Ra and .Rr

le3elAlexander: Mexican Crane-fies

77

divergent, so cell Ra at margin is very wide; cell Mr lackittg;-ffi-cu
variable in position, even in alingle specimen,on the left wing_of typebeing abouf two-thirds its length beyond the fork of M, on right wingclose to fork.
Abdomen dark brown, the discal portions of the more basal segments
a little brighter; hypopygium obscuie yellow. Male hlpopygium withthe outer dististyle-pate,darkened at apex, bidentatg, tfre ogtel spine
longer and more slender than the inner. fnner dististyle. palg' broadslender. Gonapophysesappearingas massiveblackened
bas6d,the apex
-eachwith about three to five coarse spines. Aedeagus
structures,
elongate,pale.
Holotype, & , Finca Germania, Chiapas, altitude 1190
meters, June 20, 1935 (A. M. Dampf) ; M. F. 4571. Paraty?e, &,Finca Victoria, altitude 900 meters, June 29, 1935; M. F.47108.
Shannonomyia dampfi,is named in honor of the collector, Dr.
Alfons M. Dampf, to whom our greatest knowledge of theinsects of Mexico is due. The species is so distinct that it
requires no comparison with others so far described. The only
other species with elongate antennae is S. myersiana .Plexander(Jamaica) which has abundant macrotrichia in all outer cellsof wing, cell l st Mz oPen by the atrophy of m, and the hypopygium, especially the gonapophyses and aedeagus, quite differently constructed. The relations of these various aberrant
species that have been referred to ShannonoftLyia remain inquestion, due to lack of material.
Hexatoma Latreille
Hexatoma (Eriocera) substolidasp. n.
Allied to macrocera;general coloration of notum brown, _sparsely
pruinose-setael on posterior sclerites;praescutalinterspaceswith pale sparseantennae (male) elongate,exceedingtlree times t-he length ofbody, flagellum blacki hatteies dark brown, the extreme base of stem
brown, the tips scarcely dalker; wings with a weak
pal6;'fem"orapale^bro*ntinge, including the costal region;. Rs strongly_arcuated tonearly squire at originfabdominal tergites almost uniformly brown, theincisures not darker, the lateral borders of the outer segmentsmore
yellowish.
'
Male.-Length about 6.5 mm.;wing,8 mm.;anle1naabout2Smm.
Rostrum greatly reduced,obscureyellow; palpi brown. Antennae
with r""p" an? pedicelobscuieyellow, hagellumblack; flagellum (male)
grogp; spinesof flagel.lal.segments
greatly elongated,as in the macrocer&
ielatively few and scattered, the row on basal segmentincluditg sevenor eight-spines. Head pale'brown, sparselypruinose;vertical tubercle
large and bulbous,entire.
idesottotum chiefly brown, sparsely pruinose,-darker especially .behind;stripes; vestiture ofpraescutum with thr6e barely indicated78

Annals Entomological Society of America [\rol. XXXII,

pra€scutal interspacespale and erect. Pleura dark brown. Halteresdark brown, the extreme base of stem restrictedly pale. Legs withthe coxae brownish yellow; trochanters yellow; femora pale brown,
the tips searcelydarker; tibiae and tarsi dark brown. Wings with aweak brown tinge, the costal and subcostalcells not brightened,as instoli,da;stigma oval, darker brown; cord and vein l?s darker brownthan the other veins. Venation: Rs moderatelylong, stronglyarcuatedto nearly squareat origin, slightly shorterthan in stolida.
Abdominal tergites almost uniformly brown, the incisures notdarkened,on outer segmentsthe lateral bordersof the segmentsslightly
more yellow; basal sternites brownish yellow with the incisuresdarkened;subterminalsternitesuniformly brown; hypopygiumdark.
Ifolotype, &, Finca Belem, Chiapas, altitude U00 meters,
June 22, 1935 (A. IU. Dampf) ; I\I. F. 4602.
The present fly is very closely allied to the larger llexatoma(Eriocera) stolida Alexander, of northern Panama, agreeing inmost details of coloration and venation, yet presenting somecharacters that make it impossible to consider the two flies as
being identical. The chief differences lie in the color of thehalteres, legs and wings, the lack of darkened incisures on theabdominal tergites, and in slight venational characters.
Hexatoma (Eriocera) aurantionotasp. n.
Mesonotum orange, unmarked; pleura pale brown; head withvertical tubercleand adjoining portions of posteriorvertex fiery orange,
the posterior portions of head darkened; legs chiefly brownish black;wings with a weak brown suffusion,the costal border darker brown;
abdomenwith basalfive tergitesyellow, the caudalmarginsdark brown,
the outer segmentsmore uniformly blackened.
Female.-Lengthabout 16 mm.; wing, 10.5mm.
Rostrum brown; palpi black. Antennae l0-segmented;basal threesegments yellow, succeedingsegments dark brown; terminal three
flagellar segmentssubequal. Head with vertical tubercle and that
portion of posterior vertex behind it fiery orange; sides of posteriorvertex and genae broadly grayish brown; vertical tubercle relatively
high, notched medially.
Pronotum brown. Mesonotum uniformly orange, without markings, parascutella a little darker. Pleura pale brown, the surface
weakly pruinose; a brown spot on membrane before wing-root.
Halteres black, the base of stem restrictedly pale. Legs with coxae
brownish yellow, pruinose; trochanters yellow; remainder of legs
brownish black, the proximal ends of femora, tibiae and basitarsi
slightly paler. Wings with a weak brown suffusion, the prearcularregionand cellsC and Scdarker brown; stigmalregionvaguelydarkened;veins along cord a little suffused;veins brown. Longitudinal veins
beyond cord without trichia, excepting a scatteredseries along R5.
Venation: Sc long, ,Sc1ending shortly beyond r-m; Rs one-halflonger

1e3elAlexander: Mexican Crane-flies

79

Rz about twice Rz+sand subequal to Rr+zi cell l st Mz
than Rz+a+qi
shorter than vein M+2 beyond it; m-cu at fork of.M.
Abdomen with basal five tergites yellow, the caudal margins ofsegmentstwo to five dark brown, the amount increasingon the outer
segments;terminal segmentsalmost uniformly darkened,the seventh
tergite obscure yellow in center; sternites yellow, the subterminal
segments weakly darkened; ovipositor with the genital shield andmoderatelyelongatecerci black.
IfobQpe, g , Huixtla, Chiapas, altitude 30 meters, November 21, 1930(A. M. Dampf); M. F. 1944.
Ilexatoma (Eriocera) aurantionota is closest to If . @.)obsoleta(Williston), of Honduras, differing in the small size andin many details of coloration, as the antennae, legs and wings.
Hexatoma (Eriocera) subgracilissp. n.
General coloration brown to brownish black; basal segments ofantennae yellow; head fiery orange, vertical tubercle entire; femora
and tibiae yellow, the tips narrowly blackened; wings with a strong
brownish tinge, the prearcular and costal border darker brown; trichia
of veins beyond cord reduced in number, Sczstronger than Scri Rz
yi rlt-cutbeyond fork of. M; abdomen
immediately before fork of Rz+a
black, the basal portions of tergites three and four very slightly paler;hypopygium fiery orange.
Male.-Length about 11 mm.; wing, 9 mm.
Rostrum yellow; first segment of palpus obscure yellow, outer
segmentsdark brown. Antennae (male) 7-segmented;scape, pedicel
and basal half of first flagellar segment yellow, succeedingsegments
passing into brownish black; terminal flagellar segmentselongate,alittle longerthan the penultimate. Head entirely fiery orange;vertical
tubercle moderatelyhigh, entire.
Pronotum brown, pruinose. Mesonotal praescutum with theground color brown, with four, du1l brownish black stripes that are
narrowly bordered by darker; posterior sclerites of notum brownish
black, mediotergite more pruinose, especially adjoining the sunken
areasat anterolateralangles. Pleura brownish black, sparselypruinose.
Halteres black, base of stem restrictedly obscure yellow. Legs withcoxae dark brown; trochanters brownish yellow; femora yellow, thetips narrowly blackened,the amount involving about the distal eighthor less and nearly equal on all legs; tibiae yellow, the tips narrowly
dark brown; basitarsibrownish yellow, the tips and remainderof tarsi
brownish black. Wings with a strong brownish tinge, the prearcularand costal regions darker brown; stigmal area not indicated; veinsbrown. No macrotrichiaon veins beyondcord exceptinga very sparseseries the entire length of the distal section of vein l?s. Venation:
Sc moderately long, Scr weak, erect, ending shortly before level ofr-ffi, Sczmuch stronger, subequal; Rz+s+nabout two-thirds Rs,' Rz
so a very short tosubequalto Rr+2,placed just before fork of Rz+e+apunctiform Rr+ois left; cell lst Mz shorter than vein M+z beyond it;,n-cut,nearly one-third its length beyond the fork of.M .

80Annals Entomological Society of America [Vol. XXXII,

AMomen black, the basal portions of tergites three and four veryslightly paler; hypopygium and preceding segmentfiery orange.
Holotype, &, Huixtla, Chiapas, altitude 30 meters, November 2L, 1930 (A. M. Dampf); M. f'. L944.
The nearest ally of the present fly "ppears to be Hexatorna(Eriocera) gracilis (Osten Sacken), from Onzaba, Mexico,
which differs in the large size and in details of coloration, especially the thorax and legs.
Atarba Osten Sacken
Atarba (Atarba) scutata sp. n.
General coloration yellow, the sides of the praescutum and thescutal lobes intensely blackened; antennae weakly bicolored, black, theextreme proximal ends of the more basal segmentspale; legs yellow,
the tips of the femora conspicuouslyblackened; wings brownish yellow,
stigma medium brown; Sc short; Rs and basal section of Ro subequal
and in oblique alignment; abdominal tergites bicolored;male hypopygiumwith apical point of outer dististyle slender; aedeagusweakly expanded
at tip.
M ale.-Length, 4.5-5 mm.; wing, 5-5.5 mm.; antenna about
2.2-3 mm.
Female.-Length, 6.5 mm.; wing, 6 mm.
Rostrum yellow; basal segment of palpus yellow, the remainder
black. Antennae (male) moderately long; scape and pedicel yellow;
flagellum weakly bicolored, black, the extreme proximal ends of themore basal segments yellow; beyond the third or fourth segments
uniformly blackened; flagellar segments long-cylindrical, the verticils
shorter than the segments. Head brownish yellow, the sides of theposterior vertex darkened; setae of vertex long and conspicuous,dark.
Thorax pale yellow, conspicuously variegated with black on theposterior lateral portions of the praescutum and on the scutal lobes,
the median area remaining of the ground color; dorsal anepisternum
weakly darkened. Halteres with stem pale yellow, knob darkened.
Legs with the coxae and trochanters yellow; femora yellow, the tips
abruptly blackened;remainder of legs yellow, the outer tarsal segments
weakly darkened. Wings with a brownish yellow tinge; stigma medium
brown; veins brownish yellow. Venation: Sc short, Sc1ending opposite
or just before the origin of Rs,' l?s and basal section of Rs short, nearlystraight and in oblique alignment; m-cu variable in position, from close
to fork of.M to near midlength of the cell.
Abdominal tergites bicolored,the basesof the segmentsyellow, theapical half brownish black; in male with a black subterminal ring;
hypopygium yellow. Male hypopygium with the apical point of theouter dististyle long _and slender, gently decurved at tip; subapicalspine appressed,nearly as long as the apex; two larger spinesand two
smaller denticles on outer margin, these progressivelylarger outwardly.
Aedeaguslarge, weakly expandedat tip. Appendageof ninth sternite
with caudal margin subtransverse to very shallowly notched; lateralhorns short.

rese]

Alexander: Mexican Crane-fl.ies

81

Holotype, d, Finca Victoria, Chiapas, altitude 900 meters,
June 29, 1935, 11 P. M. to morning (A. M. Dampf); M. F.
47108. Allotype, 9, Vergel, altitude 800 meters, July 2, 1935;
M. F. 4744. Paratyp€S,with the allotype, &, May 23, 1935,
M. F. 4268;I 9 , June 3, 1935;M. F. 4416.
The nearest relative of the present fly would seem to be
Atarba (Atarba) tatei Alexander, of Ecuador, which has theaedeagussimilarly constructed and agreesin other characters.
The pattern of the mesonotum and structure of the outerdististyle of the male hypopygium readily separatethe present
fly from other regional specieshaving the aedeagussimple.
Elephantomyia Osten Sacken
Elephantomyia(Elephantomyia)luteiannulatasp. n.
Generalcolorationpolishedfernrginousyellow; rostrum long, darkbrown; femora yellow basally, at near one-third the length passing intobrown, the tips blackened, preceded by . clear yellow ring of subequal
width; tibiae brown, tarsi black; wings gray, stigma long-oval, darkbrown; wings petiolate basally, cell ?nd .4 very narrow, reduced to alinear strip; abdomen yellow, the segments narrowly ringed caudallywith black; subterminal segment and genital shield black.
Female.-Length, excluding rostrum, about 8.5 mm.; wing, 7.5 mm.;rostrum about 6.5 mm.
Rostrum about three-fourths the length of the body, dark brown.
Antennae with scapepale brown, remainder of antennae brownish black.
Head brownish yellow, paler yellow behind; anterior vertex of female
subequal in width to diameter of scape.
Mesonotum uniform femrginous yellow, polished, the pleura clearer
yellow. Halteres dark brown, the extreme base of stem pale. Legs
with coxae and trochanters yellow; femora yellow basally, at near onethird the length passing into brown, the tips blackened,precededby aclear yellow ring of subequal width; tibiae brown to dark brown; tarsi
black; tibiae without true spurs but with a powerful fasciculate bristle
before tip. Wings petiolate basally, gray, the prearcular and basal
costal fields slightly more yellowish; stigma long-oval, dark brown,
conspicuous; veins dark brown. Venation: Sc moderately long, Sct
ending shortly before fork of Rs, Sczcloseto its tip; branchesof Rs only
slightly divergent; cell Rz at margin wider than cell Rs,' cell lst Mz
long-rectangular, exceedingvein M a beyond it, rlt-cu at near one-third
the length of the cell;cell ?nd A very narrow, reduced to a linear strip.
Abdomen yellow, the segmentsnarrowly ringed caudally with black;
subterminal segment and genital shield black; valves of ovipositor
horn-yellow, elongate.

Ilolotype, g, Finca Victoria, Chiapas,altitude 900 meters,
June 29, 1935(A. M. Dampf); M. F'.47104.

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Annals EntomologicalSocietyof America [Vol. XXXII,

Elephantomyia(Elephantomyia)luteiannulatais very distinct
from the other describedspeciesof the genus,being especiallycharacterizedby the pattern of the femora and the unusually
narrow cell 2nd A of the wings.
Teucholabis Osten Sacken
Teucholabis(Teucholabis)patenssp. n.
Generalcolorationof mesonotumdark brown; head polishedblack;pleura brown, with a broad black dorso-longitudinalstripe; halteresblack throughout; femora obscure brownish yellow, the tips rather
narrowly blackened; wings grayish subhyaline, the apical quarter
deepeninginto pale brown; cell lsl Mz openby the atrophy of the basal
sectionof M a;abdomenblack; malehypopygiumwith the outerdististyle
relatively slender,at near midlength with a nearly erect slenderspine.
Male.-Length about 6 mm.; wing, 5.8 mm.
Rostmm black, approximately as long as remainder of head; palpi
black. Antennae black throughout; flagellar segments oval. Headpolishedblack, the post-genaepaler.
Cervical sclerites black. Pronotum obscure brownish yellow.
Mesonotum dark brown to slightly piceous, the lateral portions ofpraescutumand the scutal lobes slightly more intense. Pleura brown,
the entire dorsal mesopleuraoccupied by a broad black longitudinal
stripel surface of pleura not or scarcely white pruinose, as in many
species of the genus. Halteres black throughout. Legs with thefore coxaebrownishblack, remainingcoxaeand all trochantersbrownish
yellow; femora obscurebrownish yellow, the tips rather narrowly but
conspicuouslyblackened;tibiae yellowish brown, the tips passinginto
black; tarsi black; posterior basitarsi weakly dilated. Wings grayish
subhyaline,the prearcular and basal portions more whitish, the apicalquarter deepening to pale brown; stigma short-oval, dark brown;veins brownishblack, thosein basalfifth paler. Venation: Sc relatively
long, Sc1ending about oppositemidlength of Rs, Scza short distancefrom its tip; fts long and unusually straight; Rz subequalto l?z+a+ri
branchesof Rs nearly parallel to one another for almost their whole
length; cell lst Mz open by atrophy of basalsectionof Ms, cell,?ndMz
about one-halflongerthan its petiole; m-cuabout two-thirds its length
beyond fork of M.
Abdomen, including hypopygium, black. Male hypopygium withthe spine of basistylesmall and slender,the tip acute, the surfacewith
numeroussetae;flange of mesalface of basistyleblackenedand irregularly roughened but not forming a single definite plate. Outerdististyle relatively slender, at near midlength bearing a nearly erect
slender spine; more basally, on mesal face, with two or three stout
retrorsetuberclesor spines;apexof style terminating in an acutepoint.
Aedeagusrelatively slender, terminating in a strong black spine andwith a shorter nearly apical lobe that bearsfour strong setae,the spine
and lobe separatedfrom one anotherby a narrow U-shapednotch.

1e3elAlexander: Mexican Crane-flies

83

Ilolotype, d, Vergel, Chiapas, altitude 800 meters, May 28,
1 9 3 5( A . M . D a m p f ) ; M . F . 4 2 6 8 .
Teuchol'abis(Teucholab'is)patens is entirely distinct from theother specieshaving cell lst M2 open, these including T. (f.)
gowdeyi Alexander, T . Q.) liponeura Alexander, T. ( r.)lipophleps Alexander, T. (r.) nigroterminalis Alexander, T. e.)om'issaAlexander and f . (7.) omissineraisAlexander. It is mostgenerally similar to T. Q.) furaa Alexander, which has celllst M closed and with a very distinct hypopygium.
Teucholabis(Teucholabis)perangustasp. n.
Allied to oteroi; general coloration yellow and black, the lateral
Pfaescutalstripes reaching the outer m-argins;basal pottion of icnob
of halteres darkened; femoral rings vefy -narrow; wing pattern very
pale brownt stiStr? small, subcircular;Sc long, Sci extendingto about
opposite two-thirds the length of fts; abdominal sternites"lined oneither side with dark brown; male hypopygium with spine of basistyle
relatively slort and stout, terminating in a short blacti lateral spine.
Male.-Length about 9 mm.; wing, 7 mm.
Female.-Lengthabout 9 mm.; wing, 6.b mm.
Rostrum dgk p-toqq; palpi brownish black. Antennaewith scapeyellow,. pedicel.yellowish brown,. flagellum black; flagellar segments
oval,.the verticils slightly exceedingthe segments. Head yellJw, theanterior vertex at narrowestpoint and a spot on cent". of post'eriorvertex dark brown.
Pronotum chiefly obscureyellow, variegatedwith darker. Mesonotal -praescutumyellow, with three black stripes,the medianone notreachingthe suture behind, at posterior end nearly confluentwith thelaterals which likewisedo not reachthe suture but are deflected.lateradto the 11argi1;scutumyellow,the lobesextensivelyblackened;scutellum
bj9ry"t-.h-yellow,p€rascutelladarker; mediotergiteyellow, the posterior
third darkened. Pleura chiefly pale,.the surfacewittr a'whiti bloom;
anegiste{numand p-leurotergite
extensive-ly
darkened. Halteresyellow,
the_basalportion of knob extensivelyinfuscated. Legs with th6 coxae
and trochantersyellow; femora yellow,-the tips narrowly dark brown,
the amount subequalon all legs; at or just beforemidlength of femora
with.a lvery narrow pale lroJvn rirg; tibiae yellow; tarJi yellow, theterminal tarsal segments{a.rkened;posterior-basitarsi(maie) slightly
e_nlarged-ang darkened,with an elongatesensory area;'clawssiirplti.
{igss relatively wide (male),.whitistrsubhyaline,restrictedlybanbed
with p.alebrown; stigma subcircular,dark brown-;band at c-ordmore
extensivethan others,.virtgallycomplete;basalband restricted,extend.ing from veT Y to-just .beyond.?nd A; wing tip-the
narrowly but conspicuously darkened; veins yellow, darker lnclouded. areas.
Venation: Sc long, {rt extending !o nearly opposite two-thirds Rs,
Sc, a short distancefrom its tip; cell lst Mi closed.
Abdominaltergitesyellow; sterniteswith a conspicuous
linear black
streak on either si-de;hypopygigm yellow. Male hypopygium withthe spine of basistylerelatively short and broad, the apex-inltrurrcate,

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Annals Entomological Society oI Arnerica [Vol. XXXII,

with one side produced into a short black spine; marginal flange ofbasistyle with relatively few coarseblackened teeth, the more cephalic
ones larger. Outer dististyle relatively slender, dusky, with conspicuoussetae almost to base, not terminating in a single acute spine.
Inner dististyle without a slender apical lobe, as in oteroi. Aedeagus
with setae abundant and conspicuous,exceedinga score in number;outer lateral anglenot producedinto a lobe, as in oteroi.
The female is generallysimilar to the male but darker, the scutellum
and pleura chiefly black, and with the abdominal tergites ringed withblack.
Holotype, &, Santa Isabel, Chiapas, altitude 800 meters,
November 16, 1930 (A. M. Dampf); M. F. 1879. Allotype, I ,Vergel, altitude 800 meters, May 25, 1935 (A. M. Dampf);
M. F. 4288.
The nearest relative of the present fly is undoubtedly
Teucholabis (Teucholabis) oteroi Alexander (Cuba), which, while
generally similar in the coloration of the body and legs, differs
conspicuously in the structure of the male hypopygium.
Gonomyia Meigen
Gonomyia (Lipophleps)subinermis sp. Ir.
Belongsto the rnanc&group; pleuralstripeslacking;malehypopygiumwith the dististyle fleshy, relatively small; phallosomewith two blackened points, the longest slender-stemmed,moderately dilated before
the apical point.
Male.-Length about 2.5-2.7 mm.; wing, 2.8-3.1 mm.
Rostrum testaceousyellow; palpi brownish black. Antennae black.
Head dark brown.
Pronotum and lateral pretergites yellow. Mesonotal praescutum
brown, the humeral region obscureyellow; scutum brown, the medianarea a trifle paler; scutellumyellow, the basemore darkenedmedially;
postnotum testaceous brown. Pleura brownish testaceous,.-paler
behind, without evident stripes. Halteres darkened. Legs with thecoxaeyellowish testaceous,the fore pair somewhatdarke_r;trochantersbrownish yellowish; remainder of legs dark brown. Wings !Io"4'strongly tinged with brownish gray; stigma pale brown, very faintly
indicited; veins brown. Venation: Sc short, Scr ending a distance
beforeorigin of Rs about equal to from one-halfto two-thirds the lengthof the latter; branchesof Rs strongly divergent; m'-cuat fork of.M.
Abdominal tergites brown, sternites paler; hypopygium yellow.
Male hypopygium-with the dististyle single, fleshy, smaller than ininermis. Ptiallosomewith two blackened points, as in inermis, butthese quite distinct in conformation; longest arm with a. very slender
stem, beforeapex moderatelyexpanded,thenceproducedinto a slender
black apical spine; secondpoint very nearly equal in length and size
to the first, longer than in inermis,arising from a moderatelyexpanded
pale basal plate, provided with marginal setae.

le3elAlexander: Mexican Crane-fl,ies

85

Holotype, d, Escuintla, Chiapas, altitude 80 meters, Novemb e r 1 1 , 1 9 3 0( A . M . D a m p f ) ; M . f ' . 1 8 1 4 . P a r a t o p o t y p e ,& .
Gonomyia (Li/o/hleps) subinermis is most closely related toG. (L.) inerm'is Alexander (British Guiana), differing mostevidently in the hypopygial structure, as described above.
Gonomyia(Lipophleps)orthomeroidessp. n.
Belongs to the rnancagroup; allied to orthomera;thoracic pleura
testaceous yellow, pruinose, without clearly defined stripes; malehypopygium with the basistyleproduceddirectly into a slenderstraightspine, the tip narrowly blackened;a singlevery small, fleshy dististyle;phallosomeconsistingof a long slenderaedeagusarising from a flattenedbasal plate that is further produced into a single blade with the tip
microscopicallytoothed.
Male.-Length about 2.8-3 mm.;wing,3-3.2 mm.
Female.-Lengthabout 4 mm.; wing, 3.5 mm.
Rostrum testaceous yellow; palpi brownish black. Antennae
brownish blackl verticils very long, especiallyin male. Head brown.
Mesonotum testaceousbrown, the lateral pretergites restrictedly
yellow; scutellum a little brightened. Pleura testaceous yellow,
pruinose, without clearly evident stripes. Halteres pale, the knobs
weakly darkened, dull yellow when rubbed. Legs with the coxae
pale;trochanterstestaceous;remainderof legsbrown. Wingsuniformly
suffusedwith brown, the prearcular and costal regionsa little paler;veins pale brown. Venation: 5-crelatively short, Sct endinga distance
before origin of Rs about equal to from one-half to two-thirds thelength of the latter; branchesof Rs strongly divergent; extreme tip of
Rr+zatrophied; m-cu at or a short distancebeforethe fork of.M.
Abdominal tergites brown, sternites paler; hypopygium yellow.
Male hypopygium with the basistyle narrowed outwardly and very
gradually produced into a slender straight spine that is tipped withblack, the darker portion set off by a weak constriction; at baseof spine
a single, unusually small, fleshy dististyle, bearing two long powerful
fasciculate setae. Phallosomeconsisting of a long slender aedeagus,
arising from a flattened basal plate; at base of aedeagus,the plate is
produced into a single blade, its tip microscopicallytoothed.
IfobQpe, d, Huixtla, Chiapas, altitude 30 meters, November 21, 1930 (A. M. Dampf); M. F. 1944. Allotopotype, 9.
Paratopotypes,d &; paratyPes, &, Escuintla, Chiapas, altitude
8 0 m e t e r s , N o v e m b e r I 1 , 1 9 3 0 ; M . F . 1 8 1 4; & , E l Z a p o t e ,Chiapas, altitude 450 meters, November 15, 1930; M. F. 1867.
Gonomyia (Lipophleps) orthomeroidesis closest to G. (L.) orthonlera Alexander (Puerto Rico), differing very conspicuously inthe details of structure of the male hypopygium, especially thenature of the spine of the basistyle, the unusually small fleshy
dististyle, and the very slender aedeagus.

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Annals EntomologicalSocietyof America [Vol. XXXII,
Erioptera Meigen

Erioptera (Empeda) divaricata sp. n.
General coloration pale reddish brown, the praescutum without
clearly defined stripes; legs yellowish brown, the terminal tarsal segments darker; femora with elongate flattened scales, in addition tonormal setae; wings with a slight brown tinge, stigma scarcely differentiated i Scr ending nearly opposite one-third .Rs,'male hypopygiumwith the outer dististyle a nearly straight pale rod, at neai midlength
bearing a long slendererect arrn that is strongly curved on outer portion.
Male.-Length about 2.5 mm.; wing, 2.8-3 mm.
Female.-Length about 3 mm.; wing, 3.3-3.5mm.
Rostrum ochreous; palpi dark brown. Antennae dark brown.
Head brown.
Mesothorax almost uniformly pale reddish brown, without clearly
defined praescutal stripes; pseudosutural foveae pale; pretergites moreyellow; pleura more testaceous. Halteres pale, the knobs weakly
darkened. Legs with the coxae and trochanters pale yellow; remainderof legs yellowish brown, the terminal tarsal segmentsdarker; legs withelongate flattened scales,in addition to the setae. Wings with a slightbrown tinge, cells C and Sc, with the prearcular field, more yellowish;
stigma scarcely differentiated; veins brown. Venation: Scr ending
nearly opposite one-third the length of lRs,' Rz a little shorter thanRz+a+riveins Rs and Ra moderately divergent, cell Rz at margin a little
lessthan cell Rg,'cellMz open; m-cu at fork of.M.
Abdomen pale brown; hypopygium large, more reddish brown.
Male hypopygium with the outer dististyle a nearly straight pale rod,the tip obtuse to obliquely truncated, at near midlength bearing along slendererect arrn that is only a little shorter than the apex of stylebeyond it, its tip strongly curved. Inner dististyle a little shorter
than the outer and of nearly the same conformation but unbranched.

Holotype, &, Vergel, altitude 1000 meters, June 3, 1935
(A. M. Dampf) ; N{. F. 1115. Al,lotopotype,? , with male.
Paratopotypes, several hundreds, altitude 800-1200 meters,
M " y 1 l - J u l y 2 , 1 9 3 5 ;N { . F . - 1 1 2 31, 1 2 1 , 4 1 3 2 , 4 1,511t ; 1 2 , 4 1 5 6 ,4172,4185,
4191,4192,41f)3,
4196,1203,1205,4206,
1207,4210,
4223,4239,1215,4259,4260,1267, 4269,1277, 4299,4320,132r,
4333,4334,4350,4351,4353, 4367, 4394,4396,4399,4399,4415,
4416,4417,4419,4430,1502,4503, 4524,4525,4633,464I, 4659,
4743,4744;paraty|es,d &, FincaAurora,June 18, 1935,M. F.
4552;Finca Belem,June23, 1935,M. F.4615;Finca Germania,
June 20, 1935,M. F. 1571;Finca Lubeca,June2L, 1935,M. F.
4582,4583; Finca San Cristobal,June 19, 1935,M. F. -1560;
Finca Victoria, M.y 29-June 29, 1935, N,I. F. 4342,4693,
46938, 4694,4709,47I0A and 47108.

1eselAlexander: Mexican Crane-fl,ies

87

Erioptera (Empeda)divaricatais readily distinguishedby the
structure of the male hypopygium, especially by the erectelongatearm of the outer dististyle.
Erioptera(Empeda)deludenssp. n.
General coloration dark brown, the pleura more testaceousbrown;
wings with a faint brown tinge, the stigma scarcely differentiated;.Scrending about opposite one-third the length of Rs,' &*r*n shorter
than Re+r, male hypopygium with the outer dististyle uniformly
blackened, profoundly bifid, the outer arrn a long simple blade, theshorter inner arrn expanded and irregularly lobed at apex; inner dististyle a pale simple rod, slightly narrowed on distal half, the apex
obtusely truncated.
Male.-Length about 2.5-2.6 mm.; wing, 3-3.2 mm.
Rostrum brownish yellow; palpi dark brown. Antennae darkbrown, the basal flagellar segmentspaler. Head gray.
Mesonotum almost uniformly dark brown, the anterior lateralpretergites restrictedly pale. Pleura testaceous brown. Halteresbrown, the base of stem restrictedly yellow. Legs with the coxae andtrochanters testaceous; remainder of legs brown, the outer segments
dark brown; legs with elongate scalesin addition to the normal setae.
Wings with a faint brown tinge, the stigma scarcely differentiated;prearcular and costal regions a little more yellowish; veins pale brown.
Venation: Sc moderately long, ending about opposite one-third thelength of Rs,'Rz+e+rshorter than Rr+ni Ra relatively long, longitudinalin position, lying generallyparallel to Rr,'cell M2open; rn-cuat fork of.M.
Abdominal tergites dark brown, sternites more brownish yellow;
hypopygium pale brown. Male hypopygium with the outer dististyleuniformly blackened, profoundly bifid, the outer arm a long simple
blade that gradually narrows to the subacute apex; inner arrn shorter,with a short stout stem, at apex expandedinto a very irregularly bilobed
or multilobed head. Inner dististyle subequal in length to the inner"arrn of outer style, pale, appearing as a simple rod, nearly parallel-sided
or slightly narrowed on distal half, the apex obtusely truncated.

IIolotype, d, Vergel, Chiapas,altitude 1200meters,June 3,
1935(A. M. Dampf); M. F. 4415. Paratopotypes,& &, altitudem e t e r sM
, a y l 3 - J u n e3 , 1 9 3 5 ; M .F . 4 1 5 3 ,4 L 9 2 , 4 4 1 5 .
800-1200
Erioptera (Em/eda) delctdensis entirely distinct from otherregional species in the structure of the male hypopygium,
especiallythe outer dististyle.
Molophilus Curtis
Molophilus(Molophilus)subsagaxsp. n.
Belongs to the plagiatus group; general coloration of mesonotum
pale brown; antennae(male) relatively long, about equal in length tothe combined head and thorax; male hypopygium with the basal

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Annals EntomologicalSocietyof America [Vol. XXXII,

dististyle a long, gently sinuous rod that terminates in an acute spine;
at near two-thirds the length on outer face with a small appressedspine;
surface of style with conspicuousminute spines and spinulae.
Male.-Length about 3.5 mm.;wing,4 mm.; antennaabout 1.4mm.
Rostrum and palpi black. Antennae black throughout, moderately
long, about equal in length to the combined head and thorax; flagellar
segmentsoval to long-oval, with very long conspicuousverticils. Headdark brown.
Mesonotum almost uniformly pale brown, the praescutal interspacesa trifle darker. Pleura dark brown. Halteres infuscated. Legs withthe coxae brownish testaceous;trochanters obscure yellow; remainderof legs dark brown to brownish black; tibiae and tarsi black. Wings
relatively wide, tinged with brown; veins and macrotrichia darker
brown. Venation: Rz lying some distance beyond the level of r-rn;
petiole of cell M s a little more than twice the oblietr€, sinuous m-cu;
vein 2nd A long, sinuous, ending about opposite midlength the petioleof. cell M a.
Abdomen, including hypopygium, dark brown. Male hypopygiumwith the apical beak of basistyle slender, curved. Basal dististyle along, gently sinuous rod, terminating in an acute spine; at near twothirds the length on outer face with a small appressedspine; surface ofstyle with conspicuousminute spinesand spinulae, including several on
basal half; just before tip, the style bears microscopicsetaeso arranged
as to produce a striate appearance. Phallosomicplate narrow, glabrous,
the distal end oval.

Holotype, &, Finca San Cristobal, Chiapas, altitude 1090
meters,June 19, 1935(A. M. Dampf); M. F. 4560.
Molophilus (Molophilus) subsagax is closest to M. (M.)
sagaxcAlexander, which differs especially in the short maleantennaeand in the details of structure of the male hypopygium,
especiallythe armature of the basal dististyle.
Molophilus(Molophilus)retrorsussp. n.
Belongsto the gracilisgroup and subgroup;generalcolorationdull
black; antennaeshort; wings strongly blackened;male hypopygiumwith the ventral lobe of basistyleproducedlaterad into a long straightspine; outer dististyle a flattened curved blade, the outer margin witha seriesof retrorse spines;phallosomicplate microscopicallysetulose.
Mole.-Length about 3.5 mm.; wing, 4.2 mm.
Rostrum and palpi black. Antennae short, if bent backrrard
extending about to wing-root, black throughout; flagellar segments
oval. Head dark gray, the anterior vertex and narrow orbits lighter
gray.
Anterior lateral pretergitesrestrictedly obscureyellow. Mesonotum
dull black, the medianregionof scutum,scutellumand postnotumvery
slightly more pruinose. Pleura black, sparsely pruinose. Halteresblackened,the knobs slightly brightened. Legs with the fore coxae
black, the remainingcoxaeand all trochantersobscureyellow; remainder

le3elAlexander: Mexican Crane-fl.ies

89

of legs black, the basesof the fore femora restrictedly pale. W'ings
strongly blackened,the veins vaguely seamedwith still darker; veinscoarse, dark brown; macrotrichia dark. Venation: Rz lying a shortdistance beyond the level of r-m; rn-cu about one-half the petiole ofcell M a; vein ?nd A relatively short, ending some distance before thelevel of m-cu.
Abdomen, including the hypopygium, black. Male hypopygiumwith the ventral lobe of basistyle at apex produced laterad into anacute straight spine; dorsal lobe longer,obtusely rounded at tip. Outerdististyle a broadly flattened sickle-shapedblade, the tip subacute;outer margin on distal half with a seriesof retrorse spines;inner margin
at near midlengthwith a seriesof erectdenticlesof varioussizes. Inner
style nearly as long, curved, slender,the outer margin on distal third
with microscopicspinules,thesebecominglarger and more conspicuous
just beforeapex. Phallosomicstructure oval, the surfacemicroscopically setulose.
Ifoh\pe, d, Vergel, Chiapas, altitude 800 meters, June 12,1935 (A. M. Dampf); M. F. 4525.
Molophilus (Molophilus) retrorsus is entirely different fromother members of the genus so f.ar described. It is the first
speciesof the gracilis group to be discovered in Middle America.
Cryptolabis Osten Sacken
Cryptolabis (Cryptolabis)longiradialis sp. n.
General coloration dark brown, the anterior lateral pretergites
conspicuouslylight yellow; femora and tibiae yellow, the tips narrowly
and gradually infuscated; wings with a uniform pale brown tinge;
macrotrichia in all cells beyond cord; Rs unusually long, on basal half
approximatedto Rr,' male hypopygiumwith two dististylesor lobesof asinglestyle, the outer a flattened pale lobe, the inner one complex.
Male.-Length about 3.5-4 mm.; wing, 3.5-4 mm.
Female.-Lengthabout 4 mm.; wing, 4.2 mm.
Rostrum and palpi brown. Antennae short, brown throughout.
Head dark.
Pronotum dark. Anterior lateral pretergites conspicuously paleyellow, becomingmore obscurebehind. Mesonotumalmost uniformly
dark brown, the scutal lobes more polished brownish black. Pleura
brown, more blackened dorsally; pleural membranespale. Haltereswith stem brown, paler basally, knob pale yellow. Legs with coxae
and trochanterspale; femora and tibiae
yellow, the tips narrowly and'Wingsgradually infuscated; tarsi brown.
with a uniform pale browntinge, the extreme base pale yellow; veins brown. Macrotrichia ofcells abundant, including all cells beyond cord except in their basal
portions; a few scattered trichia proximad of cord, in outer ends ofcells R, Cu and /sl A, and, in cases,cell R1; in cell R thesepunctures
follow a line or fold that is a direct prolongation of the base of.cell M2,
paralleling vein M . Venation: ,Sc moderately long, ending about
opposite four-fifths to five-sixths the length of the unusually long Rs,

90

Annals Entornological,
SocietyoI America [Vol. XXXII,

the latter approximating .Rr on basal half, thence diverging; Rr+r+r
subperpendicularat origin, a little longer than Rz+sirn-cu.opposite or
before midlength of M a+a.
Abdomen dark brown throughout. Male hypopygium with two
dististyles or lobes of a single style. Outer style a long pale lobe,
provided with abundant short setae. fnner style subequal in length
but more complex, narrowed at base, gently expanded outwardly, theouter angleproducedinto a curved blabfspine; just inside this black'ened
point, a dark flattened plate or flange; near outer mesal angle of style agroup of three stout spinous setae; surface of style near apex withseveralvery long, slendersetae. Aedeagusstout, long and straight.
Hol,otype, d, Vergel, Chiap&s, altitude 800 meters, May 11,

1935 (A. M. Dampf); M. F. 4123, Allotopotype, g . Paratopot y / e , & ; a s e c o n d d , M a y 3 0 , 1 9 3 5 ; M .F . 4 3 5 0 .
Cryptolabis (Cryptnlabis) l,ongirailialis is readily told fromthe other regional species by the elongate Rs, in conjunction
with the coloration of the body and wings. All other Mexican
specieshave Rs short and oblique. The male hypopygium of
the present fly is very different from that of all other described
species.