6n:
Reprinted from ANNn s oF TrrEExrouotocrcAt- Socrerv or AMenrce.
Vol. 45, No. 1, March, 1952.

NEW SPECIESOF CRANE-FLIESFROM SOUTH
AMERICA. PART XIV1
(Diptera: Tipulidae)
CHARLES P. ALBXANDER
Amherst, Massachusetts

Most of the speciesconsideredherewith are from Ecuador, Peru
and Bolivia, with a few further forms from Guatemala and Paraguay.
Individual acknowledgementsare provided under the various species.

GenusAustrolimnophilaAlexander
Austrolimnophila (Mediophragma) paraguayana sp. n.
Mesonotal praescutum grayish yellow behind, the anterior half
prow.1; femora yellow, the tips dark brown; wings whitish subhyaline,
heavily patterned with brown; male hypopygium with the polteriorborder of the nlnth tergite truncate or virtually so; outer dististylebulbo_uson proximal half, the apex narrowed to a slender curved spine.
M ale.-Length, about 12 mm. ; wing, 11 mm. ; antenna, about
2.2 mm.
Rostrum and palpi black. Antennae with the scape- black, grayish
pruinoser;pedicel brownish black; flagellum much paler, brownish
yellow; flagellarsegmentssubcylindrical,with long conspicuousverticils,
the longestabout one-half longer than the segments. Head dark gray,the orbits narrowly clearer gray; anterior vertex narrow, less than onehalf the diameter of scape;setaeof head unusually long and conspicuous.
Pronotum broadly dark brown medially, the sides narrowly
testaceous yellow. Mesonotal praescutum obscure grayish yellow
behind, the anterior half brown, including also a clearly defined medianstripe and vague indications of the anterior ends of the lateral pair;scutal lobes extensively inftrscated, the mediarr region somewhat p-aler;scutellum and a conspicuoustriangular areaon the mediotergite obscureyellow, the sides and posterior border of the latter broadly brown;
Pleurotergite extensively pale. Pleura chiefly brown, the propleura,dorsopleuralmembrane,meral region and adjoining part of the iternofleurite yellowed. Halteres elongate, pale yellow, the knob weakly
infuscated. Legs with the coxae and trochanters yellow, the fore
coxae somewhat darker; femora yellow, the tips rather broadly and
c_o_nspicuqqsly
dark brown; remainderof legs pale yellow; clarvssimple.
Wings whitish subhyaline, heavily patterned with brown, the areasin outer radial field, along cord and at origin of Rs larger and more solid;
elsewherethe cells with abundant spots and dots, in placesso denseas
to becomeconfluent;outer medial field more nearly free from darkening;rContribution from the lintornological Laboratorl', Uni'u'ersity of Massachusetts.
The prec-edingpart under this general title rvas published in the Annols of theEnlomologtcalSocietyo.fAmerica, 42:101-119,1949.
119

I20

Annal,sEntomologicolSocietyoI Arnerica[Vol. 45

veins brown, yellow in the restrictedground areas. Venation: Sc long,
Scznear its tip;Sc, ending a itro* d.istancebeyond the fork of R2a311,
cofd
Ri long, t"qo"t" at origin; Rz ohly ."boq-tone-sixthas long-as--Rraz;
oftiqo6; r,rp"ttrumerarycrossveinin cell Mt at near two-thirds the cell;p"tiot" irt .'.U Mr shorier than m; m-cu at near two-fifths the length ofcell / st,Mz.
Abdominal tergites dark brown, the posterior margins -narrorrly
pale; sternites moie brownish yellow, vaguely patterned with darker,
irp".i"tty at base and subterminally; hypopygium.brownish black.
with the ninth tergite 1arg9, slightly narrowed
M"t. hypopygium^th;bord.ertruncate or virtually so, the margil llti.kened
port.tio;tt,'anddarker in color. Outer dististyle bulbouson proximal half beyond
the narrowed.base,providedwith numeroussetae;aPexof style narrowed
into a slendercurvedblack spine. Inner dististyle about eq.ualin length,
rpp""ti"g as a sinuouslytwisted yellow blade, lhe apex obtuse. Eachgotr"popfrysisbent at altrong angle_- beyond midlength, the tig qpute..
tioiU,it.-Central Paraguiy. Holoty\e, &, CarlosPfannel,October
20, 1919(Ex Juan Foerster).
The ottr.i described members of the subgenus M ediophragma
bi-fidariaAlexander,(M ediophragma.)
Alexanderinclude Austrotimnophi,ta
Alexander(subAlexander,A. (M,)-delectissima
A. (M.) caparaoensis'andA. (M ,) microspilotaAlexander, all of southeastern
genotype),h,r^"ii.' Att speciesdiffer from- one another in the coloration andthat is most like
especiallyin the pattern of the ryittgt. .The_species
itri pt"sent fly is^microspilota,differing in relatively-slight featuresof
coloiation of the body, antennaeand wings. The male sexof the latter
fly is still unknown.

GenusHexatoma Lalreille
Hexatoma (Eriocera) esmeralda sp. n.
Size medium (wing, male, 9.5 mm.) ; head fiery orange; vertical
tubercle very large; mlsonotum chiefly dark brown, the center of thedisk yellowish orange,this including the-posteriorp?+ of the praescutum,.."i,it of the scuturi and the scutelfum; halteresand legschiefly brownish
black; wings broad, brown, the costal border and a major confluent
areabefore"cordyellow; veins unusually glabrous; cell /sl Mzrectangular,
basal section of Mz greatly rcduced, m coffespondingly long; basal
abdominal segments obscure orange, the subterminal four segments
brownish black; hypopygium orange.
Male.-Length,^ ab6ut 10 mm.; wing, 9.5 mm.; antenna, about
1 . 8m m .
Rostrum brown; palpi black. Antennae (male) seven-segmented;
scape fiery orange, pedicel obscure yellow, basal tryo lagellar segments
ofrior" yellow," the outer ones infuscated; terminll segment about
one-half ihe ^penultimate and a little longer than flagellar segments
two or three. Head fiery orange; vertical tubercle very large, in part
cephalad over the antennal bases.
projecting'?ro1oto* dark brown. Mesonotal praescutum and scutum with thelateral portions extensively dqt\ brown, the disk yellowish orange,
including the posterior pait of the praescuttlffi, most of the scutum,

L9521 Alexander: SouthAmerican Crane-Flies. X IV.

L2I

and the scutellum, parascutella dark; mediotergite brownish yellow,
darker laterally. Pleura and pleurotergite brownish black; pteropleurite vaguely obscureyellow on ventral portion; dorsopleuralmembrane brownish black. Halteres brownish black. Legs with the coxae
and trochanters brownish black; femora brown at either end, theintermediate portion vaguely paler; tibiae and tarsi brownish black.
Wings broad, brown, conspicuouslypatterned with yellow, the latterincluding the costal border and a confluent quadrate yellow areabefore
the cord, this extendingcaudadalmost to vein Cu in cells Rt, R and M ;veins brown, pale in the brightened areas, particularly in the costal
field. Veins unusually glabrous,with a few scatteredtrichia on veins Rr
and the distal section of Ro. Venation: Vein Rz subobsolete,shorter
than Rz+a;cell lst Mz rectangular; basal section of.Ma greatly reduced,,n coffespondinglylong; rn-cu about one-third its length beyond thefork of.M.
Basal abdominal segmentsobscure orange, the first tergite morechestnut; segmentsfive to eight brownish black; hypopygium orange.
Habitat.-Ecuador (Esmeraldas). Holotyqe, &, Rio Salimas, at
sea-levet,February 15, 1950(William Clarke-Macintyre).
Readily told from other regional members of the subgenusby thecoloration,especiallyof the thorax and wings. It is closestto species
such as Heratoma(Eriocera)laticostataAlexanderand I/. (E.) magistra
Alexander,yet quite distinct.

GenusAtarba OstenSacken
Atarba (Atarba)subdentatasp. n.
Generalcolorationmedium brown, the headdarker; antennae(male)elongate,the flagellar segmentswith long outspreadingverticils; legs
yellow; wings brownish yellow, vaguely patterned with brown clouds
and seams;Sc long, nr-cu,about two-thirds its own length beyond thefork of. M ; abdominal tergites dark brown, hypopygium yellow; malehypopygium with the outer dististyle blackenedat tip, the usual spinesof the outer surface much reduced, pale, the style merely scabrous;
inner dististyle conspicuouslydilated on outer end, the swollen partwith a concentrationof setae; aedeaguslarge, darkened, forming an
obtusemass.
Male.-Length, about 7 mm.; wing, 6 mm.; antenna,about 3.8 mm.
Female.-Length,about 7 mm.; wing, 6 mm.
Rostrum brownish black; palpi black. Antennae (male) elongate,
exceedingone-half the body; scape and pedicel testaceousyellow;
basal flagellar segmentsdark brown, the outer ones paling to obscureyellow; segmentselongate-cylindrical,with very long outspreading
verticils scatteredover the entire segment. Head brownish black.
Pronotumconcealed. Mesonotumalmostuniformly mediumbrown.
Pleura infuscated dorsally, the ventral portion yellow. Halteres paleyellow. Legs with the coxae and trochanters yellow; renrainder oflegs yellow, the outermost tarsal segmentsvaguely more darkened;claws small, simple. Wings browrrish yellow, vaguely washed withdarker clouds,the most distinct beingin the axillary region,at midlengthof cells R and M, and somewhatdarker areasat origin of Rs, cord andt22

Annals EntomologicalSocietyof Arnerica[Vol. 45

outer end of cell /sl M2i prearcular and costal fields somewhat clearer
yellow; veins yellow, infuscated in the patterned areas. Venation:.Sc long, ,Sc, ending about opposite four-fifths Rs, Sczweak to subevanescent,placed a short distance from tip of Scr; Rs long, weakty
angulated at origin, about two and one-half times m-cu; branches of
Rs elongate, very gradually diverging at outer ends; cell lst Mz small,subequal to vein M Ei rr-cut,about two-thirds its own length beyond
the fork of.M.
Abdominal tergites dark brown, the sternites vaguely more yellowed;
hypopygium yellow. Male hypopygium with the basistyles slender,
not produced into lobes,with someof the outer setaevery stout. Outerdististyle slender,widest at near midlength, the apex a curved blackened
spine, with two or three sma1lblack spineson outer face back from thetip, the remaining outer surface with very weak pale rougheningson1y.
Inner dististyle with the outer margin of distal end dilated, provided
with abundant pale setae,the apex virtually glabrous. Gonapophyses
obtuse, with smooth margins. Aedeagusa very large darkened mass,
its outer end enlarged and more or less curved, the parts difficult tohomologize. An appendageto the ninth sternite cannot be noted in
the unique type male.
Habitat.-Bo1ivia. Holotype, d, Cristal Mayu, Chapare, altitude1200 meters, October 15, 1949 (Luis E. Pefra). Allotopotype, I , incopula with the type.
Atarba (Atarba) subdentatais entirely distinct from all other species
that have vein .Scelongate and with long outspreading verticils on themale antennae, including A. (/.) dasyceraAlexander and A. (A.)
religiosa Alexander. The patterned wings and structure of the malehypopygium, especially the dististyles and aedeagus,are quite distinct
from all other regional species.

Genus Elephantomyia Osten Sacken
Elephantomyiadistinctior sp. n.
Sizesmall (wing, male, 5 mm.) ; rostrum exceedingone-halfthe body,
black throughout; thorax black, the pleura with a broad pale longitudinal
stripe; antennae and legs black; tibial spurs lacking; wings grayish,
conspicuously patterned with brown, including a partially complete
marginal seami r-rn some distance before the fork of Rs,' abdominal
tergites brownish black, basal sternites weakly bicolored; male hypopygium with the aedeagus relatively stout, moderately sinuous, notforming a hairlike penefilum.
Male.-Length, excluding rostrum, about 5.5 mm.; wing, 5 mm.;rostrum alone, about 3 mm.
Rostrum exceedingone-half the length of bodl', black throughout,including palpi. Antennae (male) about two-fifths the length ofrostrum, black throughout; verticils long and conspicuous,even on
the fusion-segment,the latter exceedingthe next two segmentscombined.
Head dark gray; anterior vertex narrow, about one-half the diameterof scape.
Thorax above almost uniformly blackened, sparsely pruinose, more
so behind. Pleura black, with a narrow pale line across the dorsal

19521 Al,exander:SouthAmerican Crane-Flies.XIV.

123

propleura and a broad conspicuousone on the sternopleura,reaching
the halteres. Halteres with base of stem yellow, the outer ha1f,withthe knobs, brownish black. Legs with the coxae light brown; trochanters brownish yellow; legs black, the femoral basesnarrowly moreyellowed; tibial spurs lacking; claws small, simple. Wings gray,conspicuouslypatterned with brown, including the narrow costalborder,
involving cells C and Sc, continued as a marginal seamvirtually aroundthe wing, widest in the outer radial field, broken in cells Cu, and Ist A
and again in the axillary region; other dark seams at origin of Rs,
cord and outer end of cell / st Mz; a lessevident seamalong vein Cu;
veins brown. Venation: Scr ending just beyond r-ffi, Sc, near its tip;r-tn nearly its own length beforethe fork of Rs, the branchesof the latter
virtually parallel to one an(lther throughout their lengths; cell /sl Mz
rectangular, with m-cu shortly before midlength, subequal in length
to the distal sectionof.Cur; cell ?nd A narrow.
Abdominaltergitesbrownishblack; basalsternitesweakly bicolored,
pale brown basally, the apicesbroadly yellow, the latter color becoming
more obscure outwardly; outer segmentsuniformly darkened. Male
hypopygium with the outer dististyle a slender blackened glabrousrod, gently curved and unequally bidentate at tip, the lower or axial
tooth stouter;inner dististyle a little longer,narrowedto the obtusetip.
Gonapophyses appearing as narrow flattened blades. Aedeagus
relatively stout and only moderately sinuous, not forming a hairlike
penefilum,as in somespecies.
Habitat.-Bolivia. Holotype, d , Cristal Mayu, Chapare, altitude
1200meters,September26, 1949(Luis E. Pefla).
In its conspicuouswing pattern and the venation, as the positionof r-m somedistancebeforethe fork of Rs, the presentfly is more likethe only describedspeciesof the subgenusElephantomyinaAlexander,
supernumerariaAlexander, which differs in the subgenericcharacterof a supernumerarycrossveinin cell R, and in all detailsof wing pattern.
The speciesis entirely distinct from all other Neotropical membersof the genusthat have lost the tibial spurs.

GenusTeucholabisOstenSacken
Teucholabis(Paratropesa)
fumidapicalislaetabilissubsp.n.
Male.-Length, about 7 mm.; wing, 6.6 mm.
Rostrum orange; palpi black. Antennae with the scape orarlgeyellow, the remainder brown; flagellar segmentsoval. Head orange.
Pronotum uniformly orange. Mesonotal praescutum orange, witha complete transverse black saddle shortly before the suture, the cephalicthird of the sclerite similarly polished black; scutum orange, each lobe
polishedblack; scutellum black, parascutellaobscureorange; postnotum,
including both the mediotergite and pleurotergite, black, only theventral edge of the latter restrictedly orange. Pleura orange, thedorsal pteropleurite black, confluent with the major mass on the pleurotergite. Halteres with stem brown, knob light yellow. Legs with thecoxaeand trochanters orange; fore legsblack, the femoral basesnarrowly
yellow; middle and posterior femora yellow, the tips narrowly brownish
black, including about the distal sixth; tibiae darkened at base and tip,t24

Annals EntomologicalSoc'ietyof Arnerica[Vol. 45the central portion broadly obscure yellow; tarsi black, the middle
basitarsi vaguely more brightened; posterior basitarsi with about theproximal third moderately dilated. Wings with the ground subhyaline,
patterned with dark and paler brown; the darkest color includes thearculus and origin of Rs, with most of the intervening cells C and Sc,'a narrow seam over the cord, ending at Cu, in alignment with a muchpaler cloud at end of vein ?nd A,' a secondmore broken band at level oforigin of Rs, ending at vein ?nd A, a trifle more expandedin cell Cu;
wing tip-Mr+r;
broadly paler brown, the inner margin almost al the fork
veins pale brown, darker in the patterned areas,C, 5c andof veinRs before the stigma and the subbasal portions of veins Cu, l st A
and ?nd A clear light yellow. Venation: Rs ending about opposite
two-fifths the length of Rs,' Rz immediately beyond the fork of Rs,
ftr+r+nthus extremely short or lacking.
Abdominal tergites black; sternites yellow, the extensive but illdelimited pocket of the fifth segmentweakly darkened; ninth segment
yellow, the styli abruptly black. Male hypopygium with the outerappendageof the basistyle longer and narrower than in lindneri. Innerdististyle with the outer lobe unusually slender,terminating in an acuteblackened spine.
Habitat.-Bo1ivia. Holotype, &, Cristal Mayu, Chapare, altitude
1200meters, October 4, 1949 (Luis E. Pefra).
Close to the typical form (from Amazonian Peru) which is still
known only from the female sex. The two races are separated by thedetails of color.
Teucholabis (Teucholabis) egens sp. n.
group ; mesonotalpraescutumwith theBelongsto the melanocephaladisk polishedblack, the humeral, posterior laterat and posterior medianareas yellow; posterior sclerites of notum with a broad yellow central
area; halteres infuscated; femora black, with an obscure yellow ring at
near two-thirds the length, the fore pair brightened basally; wings
whitish subhyaline, unpatterned except for the small brown stigma;
abd,omenof male with ihe sternal pocket of segment five with six rowsof sensorypits; male hypopygium with the basistyle narrowed outwardly,
the appendage placed at or before midlength of the mesal face; outerdististyle a straight rod, its apex vaguely bidentate; lower surface of
aedeaguswith a group of about sixteen setae.
Male.-Length, about 8.5 mm.; wing, 7.5 mm.
Rostrum moderately long, black; palpi black. Antennae black
throughout; flagellar segments oval, shorter than the longest verticils.
Eyes- very large; head above reddish brown.
Pronotum brown. Mesonotal praescutum with the humeral region
on either side before the suture polished yellow, the remainderand a spot
-occupied
by a polished black shield that reaches the laterat
chiefly
margin, the median area before the suture broadly yellow; posteriorsclerites of notum broadly yellow, including the wide central regionof scutum, the outer parts black; parascutella darkened; mediotergite
yellow on cephalic half, darkened behind. Pleura attq pleurotergite
cnieny- blackened, the pteropleurite and a ventral longitudinal stripe
more silvery pruinose, the color produced by microscopic scales.

L9521 Alexander: SouthAmericanCrane-Flies.XIV.

l2s

Halteres infuscated. Legs with the coxae reddish brown to brown;
trochantersbrownish yellow; femora black, with very vague indicationsof a yellow ring at near two-thirds the length, leist distinct on theposterior legs; fore femora yellowed basally-and with the ring more
distinct; remainderof legs brownish blacki posterior basitarsititatedon about the proximal third. Wings whitisli subhyaline,unpatterned.
exceptfor the small brown stigma; veins brown, more browniih yellow
in.the prearcularand costalareas. Venation:Screndingabout opposite
midlength of lts, Sc, at near one-fourth this length; branchesbf Rs
generally parallel to one another for virtually their entire lengths;
rn-cuabout one-third its length beyond the fork of.M .
Abdomen dark brown, the sternites and hypopygium somewhat
more brightened. Sternal pockets generally as in the melanocephalagro_Tp,-witha conspicuouspocket on sternite five, including six longitudinal rows of sensorypits, and a longitudinal row of about six setaeon either side of the midline of sternite six. Male hypopygium withthe basistyle narrowed apically, the tip more obtuse; lbp-endageat or
before midlength of the mesal face of-style, relatively narrow, curved.
into the- long black apical spine, the concavemargin-its with a fringe oflong yellow setae; mesal flange of style darkerred, margin coarsely
crenate. Outer dististyle a straight rod, the apex vaguely notched orbidentate. fnner dististyle irregular in outline, its outer blade simple.
Aedeaguswith two setaeon upper surfaceand a group of some 16 to 18
on the lower portion.
Habitat.-Bolivia. Holotype,& , Cristal Mayu, Chapare,altitude
1200meters,September24, 1949(Luis E. Pefla).
. _ The 9n1yother allied specieshaving unpatternedwingsis Teucholabis(Te,ucholabis)
jiaaro Alexander,which hasthe coloration,iternalpockets,and the male hypopygiumquite distinct.
Teucholabis (Teucholabis)volentis sp. n.
General coloration of thorax black, the praescutum with threeb-lackstripes; head black; rostrum relatively long; halteresinfuscated,
the knob yellow; femora brownish yellow, the tips brownish black;posteriortibiae with the basalfourth moderatelydilated; wings whitish,
vaguely patterned, including the narrow tip and a narrow but moreevident seamover the anterior cord; Sc long; male hypopygium withthe appendageof the basistyle a flattened y-ellowblade, ai apex bent?t a right angleinto a long black spine; outer dististyle bearingan acutelateral spine; lower lobe of inner dististyle extended into a point ;aedeagusterminating in a sinuousreddish spine.
Male.-Length, about 6.5 mm.; wing, 5.8 mm.
Rostrum relatively long, nearly equal to the remainder of head,
black throughout; palpi black. Antennae black; flagellar segments
oval, shorterthan the verticils. Head black, narrowedbehind.
Pronotum above obscure yellow, narrowly darkened laterally.
Mesonotal praescutumwith the ground light biown, with three blaCk
stripes, the lateral pair crossing the suture onto the scutal lobes;
posterior scleritesof notum chiefly black, the parascutellaand suture
between the mediotergite and pleurotergite- pale. Pleura chiefly
blackened,the ventral pteropleuriteand meral regionpaler but scarcelyr26

Societyol Arnerica
Annals Entomological,[Vol. 45

pruinose; dorsopleural reg_ion"L"gr restrictedly pale. .Halteres with stemwith the coiae infuscatg{, restricted-ly
intuscated,knob yellow.
pt"i"or.; trochanlers browniih -yellow; femora brownish yellow, the;ipr b;;#"irtt biack; tibiae yellowish brown, !h.- tips and the tarsi
darker brown; posterior'Wingr tibia6 with more than the basal fourth moderwhitish, clearerin the basal and costal portio.ns;;6ly dilated.'u "i-ow and very loorly indicated dark seam over the cord, darker
oo ift" anterior .oia, paler and.more diffuse at end of vein l st A; wingbrownish black.
n"*o*ty aqd poorly darkened;__veins.
;ip il"*ft"tCbrt"f fringe (male) ielatively consplcuous. Venation: Sc long, Scr.trdi"g aboirt opporite three-fourths Rs, Sczfar from its tip,- at nearbranchesof Rs,'Rzabout one-halflonge-rthlunl?r+r+n;
one-thirclthe te"gtfr-parallel
ioi most of their- lenfth, Rs on outer fourth;f Rr generally
deflectedstrongly caudad,terminating at the wing tip ; m-cuabout twothe fork of-M .
fifths to one-hatfits length beyond-brownish
black, the posterior and lateral
Abdomen dark bro'iln toborders darkei; hypopygium black. Pocket of sternite five of malei*toai"g relativety t"*" and scattered setae, these directed caudad.
Male hypopygium"with the appe-n{ageof the basistyle ."b.t..t-ina1,
upp.uti"i ;i i flutt"ned yellow-blade, narrowed at aPex and bent at"'figf.t a?rgleinto a strorli black -spine;inner margin fringed ryilh lottg
yeil6w set?e; marginal fl"angeblackened, its edge- coarsely lobed orarm,. at beyond midlengthioothed. Outer d-ististyle ; powerful^rod.
that terminates in an acute black
nurro*.d. ilto a strong curved.
;-pi";, ui poi"t of na#owing wi-th a small slender acute black -spine;rlrf".. of^style with abundint long coarsesetae; near base of lateral;pi"; with "".or,."ntration of setulie. Inner _dististylewith the outer"i pri".iput nUde bidentate; lower lobe blackened,extendedinto an"."t" poitrt,'inthe surface with about five strong- setae. Aedeagus
a sinuousreddish spinouspoint, before the spine with
f.i*i"uting
a pair of loig setaeaboveand a singleoneon lower edge.
Habitat.-Bolivia. Holotype,d, Cristal Mayu, Chapare,altitude
1200meters,August 16,' 1949(Luis E. Pefra)'
The .nori- Jimilar regional species appears to be Teucholabisd,enud.aAleiander, wtrich differs conspicuouslyin theeriiiotiiltj
structureof the male hypopygium.
Teucholabis (Teucholabis)rectangularis sp. n.
Mesonotumblack, the lateral and humeralmarginsof th.9praescutuma"ff r.aaisfr; pflrr.tu-'b1r"k,striped longitudinally .*i!h silvery; knobs
of t "tt.res yel-low;femora'yellow, the Iips abruptly blackened;wings;bhy"iine, Lnmarked. e*cept for the^tlig.*q lttd .a paler brown seamoir the anterior cord; sternai pocket of fifih abdominalsegmentof maleruig. and colspi.oonr, consisiing9f abundant spinousPoints surroundi;g ; .""ttut Jt""; male hypggigiuq with the spine of th.e appgndqs-e
oitt " basistylestrong, at iiigttt"u"gle to the axis; outer dististyle witha strong rat6tar-;pi;r;; inner?irtirtJ'le with a single developedapicalspine.
'
Mole.-Length, about5 mm.; wing, 5'2 mm'
Rostrum i"i"ti""ly 1ong,slightly'i"ss than the remainderof head,1952] Alexander: SouthAmerican Crane-Flies.XIV.

I27

black; palpi black. Antennaeblack throughout;flagellarsegmentsoval,
shorterthan the verticils. Eyes large; headblack.
Pronotum above dark reddish brown, blackenedon sides. Mesonotum black, the humeral and lateral margins of the praescutumdull
reddish. Pleura black, with a broad silvery longitudinal stripe, more
expandedon the posteriorsclerites. Haltereswith stem black, restrictedly obscureyellow at base,knob obscureyellow. Legs with the fore
coxaedarkened,the remainingcoxaeobscureyellow; trochantersyellow;
femora yellow, the tips rather narrowly but abruptly blackened,
including about the distal sixth or seventh;tibiae obscureyellow, thetips narrowly darkened; tarsi passinginto black; posterior basitarsi
narrowly dilated at proximal end. Wings subhyaline,very restrictedly
patternedwith darker, includingthe oval dark brown stigma and a vague
paler brown seamover the anterior cord; prearcularfie1dyellow; veinsbrown, the basal ones more yellowed. Venation: Sc long, Sc, ending
approximately oppositemidlength of Rs, .Sczabout oppositeone-fourth
this length; branchesof Rs generallyparallel to one another, Ru near
apex bent caudad to the wing tip, cell Rr being approximately fourtimes as wide at margin as cell Rr i m.-cuabout one-fourth its length
beyond the fork of.M.
Abdomen,includinghypopygium,black. Sternalpocketof segment
five large and conspicuous,including an extensive area of blackened
spinous points directed chiefly mesad, the center of the area an
elongate patch of microscopic scalelike points. Male hypopygiumwith the apex of the basistylebroken; appendageon mesalfaceremotefrom tip, stout, at apex bent into a right angle into a long straightspine,the inner edgeof the rod with a fringe of long yellow setae. Outerdististyle stout on proximal two-thirds, thence narrowed into a more
slender arm, at point of narrowing with a powerful erect spine; outermargin and the narrowed arm microscopicallyscabrous;setaeof style
relatively sparse. Inner dististyle with a singledevelopedapical spine.
Aedeaguswith a pair of strong setaeon upper surfaceand about three
on the lower; apex of aedeagusnaffowed into a slenderspinousportion(the tip apparentlybroken).
Habitat.-Bolivia. Holotype, d , Cristal Mayu, Chapare,altitude
1200meters,December9, 1949(Luis E. Pefla).
The male hypopygium is generally similar to that of Teucholabis(Teucholabis)
aolentissp. n., but all colorationalfeaturesand the abdominal sternalpocketsof the male are entirety differentin the two flies.
Teucholabis (Teucholabis) tactilis sp. n.
Head black; generalcoloration of mesonotumreddish yellow, thepraescutumwith three major black areas,the scutal lobes blackened;
pleura obscureyellow, with a broad black longitudinal stripe; knobs ofhaltereslight yellow; femora yellow, the tips blackened;wings with avague yellow tinge, the tip narrowly infuscated, stigma oval, darkbrown; abdomenblack,the posteriorbordersof the segmentsrestrictedly
obscure yellow; male hypopygium with the outer dististyle a simple
rod, narrowedinto a spineat apex; inner dististyle with its outer blade
terminating in two widely separatedteeth; aedeagusterminating in astrong black spine.

128

Soc'ictyoI Arnerica
Annals Entomological,[Vol. a5

Male.-Length, about 6.5 mm.; wing, 6.5 mm.
Rostrum blick, about two-thirds as long as remainder of head;palpi black. Antennae black throughout; flagellar. segments oval,
weli constricted, with a dense dark pubescenceadditional to the long
verticils. Head black, more pruinose behind.
Pronotum obscureyellow, blackenedlateratly. Mesonotum polishedwith ihree major black areas, including the cephalicreddish yellow,-theusual median stripe and conspicuous lateral marks thatthird ofvirtually reach the lateral bord-er; posterior sclerites of notum reddish
yellow, each scutal lobe with a large polished black spot. Pleuraobs.ute yellow, with a broad black longitudinal stripe over the dorsal
sclerites,-extending from the propleura onto the pleurotergite, passing
above the root of the halteres;-ventradof this stripe an extensivesilvery
lorrgitudinal area; ventral sternopleurite weakly dartened. Halteresbrownish black, the apex of knob light yellow. Legs with the fore coxae
weakly darkened, the middle and hind coxae, with all trochanters,
yellow; a single (detached) leg remains, femora yeltow, with about
lhe outer fifth blackened; tibiae and tarsi black; sincethere is no modification of the basitarsus and from its relative length, it is presumedthat
this is a fore leg. Wings with a vague yellowish tinge, the Prearcularand. costal fields somewhat clearer yellow; stigma oval, dark brown;
wing tip narrowly and vaguely infuscated; veins dark brown, brightenedin tle yellow netas. Venation: ^Scrending about opposite midleggthof Rs, sc, just before one-third this length; branches gf_4r slightly
divergenf ai outer ends; rr-cu shortly beyon4 the fork of. M.
A6domen black, the posterior borders of the third and succeeding
segments very narrowly and inconspicuously obscure -yellow, more
two sternites yellow; segment
extensive on ihe sternites, the proximal-Malehypopygium with .the basistylenine yellow, the styli black.
blunt-at tip, the appendageslightly subterminal in position, narrowed
into a long'terminai spine, fringed wittr long setae; mesal flange o{ style
black, irregularly and coarselyloothed. Outer dististylg a simple -tod,
at apex narrowed into a spine. Inner dististyle with the outer blade,r"ry broad, flattened.,terminating in two widely separated teeth; lobeof ityt" reiatively small and inc-onspicuous,terminating in a. curved
black spine, with about four long setae. Aedeagus terminating in astrong black spinouslobe, its tip acute.
nlU;tat.-Guatemala. Holotype,d, Chicacao,El Naranjo, altitude
3800 feet, July 12, Ig49 (T. H. Farr).
such as Teucholabis(Teucholabis)
This fl1i is"most similar to species-alldetails of structure of the malefurva Aleiand.er, differing in
hypopygium.
Teucholabis (Teucholabis) podagra sp. n.
Head. behind the antennal bases black, front and rostrum yellow;
mesonotum polished black, covering virtually the.whole praescutlm;
postnotum ahd pleura black, the latter with conspicuoussilvery whiterreas above the middle and hind coxae; knobs of halteres light yellow;
legs chiefly darkened; posterior tibia of male with a swollen area at neart#o-thirai tne length;-wingswhitish subhyaline,patterned with brown;
abdomen black, tfie incisuies yellow; male hypopygium with the outer1952] Alexander: SouthAmericanCrane-Flies.xIV.

Lzg

dististyle bearing an acute spine beforemidlength; teeth of outer bladeof inner dististyle unequal.
Male.-Length, about7.5mm.; wing, 0.8mm.
Rostrum short and stout, yellow; palpi brownishblack. Antennae
black; flagell?rsegmentswell constricted,especiallythe enlargedbasal
ones; verticils longer lhan the segments. Head brownisii yellow
surroundingthe antennalbases,the remainderblack.
Pronotum light yellow. Mesonotalpraescutumblack, the extreme
posterior border adjoining the suture r-eddish;scutum and scutelium$tq the ground reddish,the scutal lobesand basal part of scutellum
blackened;postnotum and pleura black, the latter with conspicuous
silvery white pubescentareasabovethe middle and hind coxaei dorsopleural-membraneand region of the wing-root yellow. Halteres withstem black, knob conspicuouslylight yellow. Legs with all coxaeand
trochanters yello*; fore femora obscureyellow bisally, the outer half
more infuscated,the tip brownishblack, libiae and tarsi black; middle
and hind femora.yellow,tibiae brownish yellow, tarsi brownish black;posterior tibiae (rya-le). qt about two-thiids the length conspicuously
swollenand providedwith denseblack setae,additionalto the stronger
ones that cover_!h. 9egry_e_nt
basitarsi with the proxirial; posterior
third strongly dilated. Wings whitish subhyaline, patterned withbrown; stigmaoval, darker brown, confluentwiih a seamover the cord,reachingthe margin at-end of vein lst A but paler; wing tip similarly
palebrown,the inner edgevirtually at the level-ofm;bas{dol cord witlr
a darkenedcloud at near midlength of veins Cu, lst A and pnd,A;veins brown, those at extreme 6ase more yellowed. \,renation:Sc
moderately lolq, {rt ending just before midlength of Rs, Sc, nearly
oppositeone-fifth lltjr length; ftz+g+rabout one-tfiird Rz; cell Ragently
widened before midlength and strongly so at apex by the dowiwari
deflectionof vgi1_Rsi cell lst Mz narrow, gently wid-enedoutwardly;
distal section of.M +z arched; n -cu at the fofu of-M .
Abdomen black, the incisures yellow, very narrowly so on theouter segments,!h" pale color finally obliterated outwardly; eighth
segmentyellow; hypopygium black. Sternal pocket of segment-fivetriangular in outline, the point directed cephilad, the cetiter of thearea Y-ery_.
densely let with sma1l sensory points, the margin withinwardly directedb,ristles;sternite six with a group of six to eight strong
bristleson eithersideof the midline,directedmesad. Male hypopygium
wit-h the-appendageof the basistyle subterminal in position,-flittEned
and setiferous,at apex narrowed into a slenderspin-e;flange of mesal
face.of style compact, the margin vaguely crenuiate or er5se. Outerdististyle a slenderrod, at apexabruptly narrowedinto a spine,on lower
m.algiq before midlength with a longer-theslenderspine. Inner dististylewith the apex unequally bidentate,outer spine long and slend'er,
ge-n!1y-curved,
the inner spineshorter; basal lobe of style producedintoa blackenedspinelike beak, the surfacewith more than a dozenstout
setae. Aedeaguswith the apical spineslender,only gently curved.
Habitat.-Guatemala. Holotype,d, Chicacao,ilfNarinjo, altitude
5100feet,July 11, 1949(T. H. Farr).
Readily distinguishedfrom the other members of the subgenus
that have a modified area on the posterior tibia of the male bv tt e

130Annals EntomologicalSocietyof Arrerica[Vol. a5coloration,wing pattern, venation, and structure of the male lrVRonVgium. Such otlier species include Teucholabis(Teucholabi,s)elissa
Alexander,T. (f.) stadelmanniAlexander,T. (f.) wighti Alexander,and some others, all being further differentiated among themselvesin the detailsof structureof the dilated tibial area.

GenusGnophomyiaOstenSacken
Gnophomyla (Gnophomyia) certa sp. n.
Belongsto the tristissimogroup; generalcolorationblack, including
the antennae,halteresand legi; thoracic pleura heavily pruinose;wings
with a strong blackish tinge, more suffusedalong vein Cu and above
the anterior cbrd; male hypopygium with the ninth tergite nearly square,
the outer angles produCedinto stout lobes, the entire caudal borderprovided with abundant short black spines; mesal face of basistylewittr two unequal rods, the basal one stout, terminating in .an_agutespinouspoint; outer spine small, its base narrowed and encircledby
hyaline Collarlikemembrane;outer dististyle a 1ong.b1"*enedglabrous
r6a; inner dististyle small, the apex shapedlike a birds head and beak.
M ale.-Length, about 5.5mm. ;wing,5.7mm. ;antennaabout2.3mm.
Rostrum an? palpi-byblack. Antennae black throughout, relatively
the measurements;flagellar- segments longelongate, &S shown
subrylindrical, more narrowed outwardly, shorter than the longest
verticils. Head black, sparselypruinose.
Thorax black, the notum sparselypruinose, the pleura heavily so,
particularly- behind. Halteres and leg! black, the coxae sparsely
pruinose. Wings with a strong blackish tinge, more suffused alongvein Cu in cellM and in cell R1,'stigma long and narrow, darker brown;veins brownishblack. Venation: Rs virtually straight, r-m at its fork;
veins Raand ^Rrvirtually parallel to one another, a little more separated
at outer end; rn-cuat near midlengthof cell / st Mz.
Abd,omen,including hypopygium, black. Male hypopygitp with
the ninth tergite nearly siliraie in outline, the caudal margin broadly
emarginate,rliith the outei lateral anglesproduced; the entire margin
is pr"ovided with abundant short black spines. Basistyle with themeial face producedinto two very unequal spines,the_largerone basal,
at apex narrowed into a strong spinouJpoini, the surfaceof outer halfwith at least two powerful setae; secondspine just before midlengthof mesal face of style, its base abruptly-q narrowed and encircled bylong blackened glabrous rod,hyaline membrane. Outer dististyl"
giadually narrowed to the blunt tip. Inner dististyle small, narrowed
it apex'into a structure that is shapedlike a bird's head and beak,the ityle with severalcoarsesetae. Phallosomewith a deep U-shaped
central emargination.
Habitat.jPeru (Junin). Holotyqe, &, Chanchamayo, altitude
1100meters, January27, 1949(J. M. SchunLe)-,
The closeit allies include Gnophomyia (Gnophomyia)permagico
Alexander,
and-G.(G.) podacantha
Alexander,G. (G.) petentisAlexander,
all readily distinguishedby the structure of the male hypopygium.

lg52lAlexander: SouthAmerican Crane-Flies. XIV.

181

Genus NeognophomyiaAlexander
Neognophomyia
obtusilaminasp. n.
Mesonotumreddishbrown, the postnotumdark brown; pleura

yellow with a broad dorsal dark brown stripe ; femora brownish y-ellow;
wings subhyaline, very slightly patterned along the cord; male hypopygium with the teryal spines very strong and powerful, blackened;phallosome consisting of two large blackened obtuse plates that are
not producedinto spinouspoints.
Male.-Length, about 5.5 mm.; wing, 5.8 mm.
Female.-Length, about 7.5 mm.; wing, 7 mm.
Rostrum yellow; palpi brown. Antennae dark brown; flagellar
segmentsoval, a little shorter than the verticils. Head yellow.
Pronotum infuscated medially, light yellow on the sides and on
pretergites. Mesonotal praescutum chiefly reddish brown, darker
brown in front, the humeral region more yellowed; a vaguely indicatedobscureyellow median vitta; scutal lobes and scutellum medium brown,parascutella paler; mediotergite and dorsal pleurotergite dark brown,
this being the posterior extension of a nearly unbroken broad dorsopleural stripe; remainder of pleura yellow. Halteres with stem yellow,
knob infuscated. Legs with the coxae and trochanters light yellow;
femora brownish yellow, tibiae and tarsi a trifle darker; outer tarsal
segmentsbrown. Wings subhyaline, very slightly patterned, the only
indication of darker color being a weak suffusion along cord and a
deepeningin color of the veins, at the cephalicpart including the vague
stigmal darkening; veins brownish yellow, darker in the clouded
portions. Venation: Scr ending about opposite the slightly oblique R2,
long, being more than one-third Rs,'vein J?aver1rdecurved, ending justbefore the wing tip; m-cu nearly its own length beyond the fork of.M .
Abdominal tergites dark brown; basal sternites yellow, the outer
onesmore infuscated; hypopygium brownish black. Male hypopygiumwith the tergal spines very strong and powerful, blackened. Outerdististyle darkened, provided with about five strong setae,the outermost
terminal in position. Inner dististyle a blackenedflat plate. Phallosome
consisting of two large obtuse blackened plates, not produced into
spinouspoints as in related forms.
Habitat.-Peru (Huanuco). Holotype, &, Fundo Sinchono, Cordillera Azal, altitude 1500meters,August 23, 1947(Schunke). Allotopoty|e, I .
The closest allies of the present fly include Neognophomyiaadara
Alexander, /f. cochlearisAlexander, ^|f. consociafu.eilexander,and /f.
schildi Alexander, all of which differ from it and among themselvesin
the structure of the male hypopygium.
Neognophomyia scaphoides sp. n.
Allied to scapho,'mesonotum
pale brownish yellow, vaguelypatternedwith pale brown, especiallyon the scutal lobes and scutellum; pleurawith a broad dorsal stripe beginning on the propleura, passing above
the halteres onto the pleurotergite and mediotergite; tips of femora
infuscated; wings whitish subhyaline,very restrictedly patterned withbrown, most evident as a seam along the cord; abdominal tergites
brown, paler along the midline; male hypopygiurn with the tergal spinesr32

Annals EntomologicalSocietyof America[Vol. a5long and powerful, blackened;inner dististyle a broadly flattened blade,its apex a more or less distinctly bifid blackened beak; phallosomea
narrow central structure, at apex on either sidewith a recurvedblackened
arm that terminates in from two to four irregular teeth.
Male.-Length, about 5.5mm.;wing, 5.5mm.
Rostrum testaceousyellow; palpi light brown. Antennae black
throughout; flagellar segments long-oval, shorter than the verticils.
Head yellow.
Pronotum yellow above, darker on sides. Mesonotum almost
uniformly pale brownish yellow, clearer yellow on sides,the disk vaguely
patterned with very pale brown, most evident on the scutal lobes andscutellum. Pleura and pleurotergite yellow ventrally, the dorsal
portion chiefly occupied by a broad brownish black longitudinal stripe
beginning on the propleura, deepest in color on the anepisternunt,passingabovethe root of the halteresonto the pleurotergite and posterior
lateral parts of the mediotergite. Halteres obscure brownish yellow,
the base of stem narrowly more brightened. Legs with the coxae andtrochanters yellow; femora and tibiae obscureyellow to brownish yellow,
the tips rather conspicuouslyinfuscated;tarsi yellow, the outer segments
passinginto brown. Wings whitish hyaline, the prearcular and costal
fields slightly more yellowed; a restricted brown pattern along the cord,most distinct at the cephalic or stigmal end; veins obscure yellow,
darker in the clouded portions, including the outer end of cell / st M*
Venation: Vein Rr relatively short and oblique, as usual in the genus,
cell Rz at margin thus very narrow; vein Ra greatly decurved, endingat wing tip.
Abdominal tergites chiefly dark brown, paler along the midline
to produce a rather evident stripe, most conspicuous on the basal
segments;basal sternites chiefly yellow, the outer ones weakly infuscated; hypopygium darkened. Male hypopygium with the tergal
spinesstrong and powerful, blackened. Outer dististyle a narrow yellow
blade, gradually narrowed outwardly, at and near apex with three strong
yellow setae, with one further isolated bristle at near two-thirds thelength. Inner dististyle a broadly flattened blade, its apex a more orless bifid blackened beak; basal flange a broadly triangular blackened
point, provided with severallong strong blackened setae; setaeof outermargin of style about seven in number. Phallosome somewhat asin scapha, appearing as a central narrow depressedstructure, at apex
on either side with a recurved narrowly blackened arm that terminatesin from two to four small irregular teeth.
Hobitat.-Peru (Huanuco). Holotype, 6/, Fundo Sinchono,Cordillera Azul, altitude 1500meters,August 14, 1947 (Schunke).
While somewhat similar to Neognophomyiascapha Alexander, ofEcuador, the present fly is quite distinct in all details of structure of
the male hypopygium, especiallyof the phallosome.

GenusGonomyia Meigen
sp. n.
Gonomyia(Gonomyta)platymeroides
Mesonotum brown, sparsely pruinose, scutellum pale yellow;
antennae black throughout; wings with a strong brownish tinge, thestigma and a seam behind vein Cz slightty more darkened; Scr just19521 Alexander: SouthAmerican Crane-Flies. XIV.

lBB

beyond the origin of Rs, the latter subequalin length to vein Rr+r+nand
lessthan twice R3;male hypopygium with the apical lobe of the basistyleve-rYsmall,_itsapical setaeexceedingthe lobe in length; inner dististylea broadly flattened disk, on outer margin near apei with a blackened
tooth; phallosomeconsisting of two pairs of gonapophyses,the outerpair terminating in a strong spine, the inner ones expandedoutwardly
into weak blades.
M ale.-Length, about 5 mm. ; wing, 5 mm.
Rostrum and palpi black. Antennae black throughout; flagellar
segmentsrelatively long, cylindrical, eachwith a singleelongateverticil,
unilaterally arranged, additional to a densewhite pubescence. Head
dar! blue gray, with orangemarkings on posterior portion.
Pronotum and pretergites whitened. Mesonotum brown, thesurface more or less pruinose, especiallyon the median region of the
scutum and on the mediotergite where a blue-greensheenis evident;
scutellum pale yellow. Pleura and pleurotergltechiefly pale, the dorsal
pleurites and ventral sternopleuriteweakly more infuscated,leaving apale longitudinal stripe. Halteres infuscated,the baseof stem narro*ly
yellow. Legs with the coxae and trochanters pale; remainder of legs
dark brown. Wings with a strong brownish tinge, the stigma and aseam behind vein Cu slightly more darkened; prearcular and costal
fields a trifle more whitened; veins brownish black. ' Venation: Sc,
ending just beyond the origin of Rs, Sczbefore this origin; Rs subequalin length to R +r+nand less than twice R3; cell l st Mzlelatively small,closedi rn-cuat fork of.M .
Abdomen dark brown, the hypopygium a trifle paler. Male
hypopygium with the apical lobe of basistyle very small, the terminal
setae longer than the lobe itself. Two dististyles, the outer a long
slender fleshy lobe, gently curved and very gradually narrowed toan acute needlelikepoint. fnner dististyle a broadly flattened disk,
the outer margin near apex with a blackenedtooth or projection; inner
margin of disk evenly rounded, with relatively short setaebut with nodifferentiated fasciculate ones. Phallosomeconsisting of a relatively
small pale blade, subtended by two pairs of gonapophyses;outerapophysesslender,terminating in a strong spine that is subtendedby
paler membraneextendingbeyond the point of insertion of the spine;
inner apophysesblack, & little shorter, expandedoutwardly into weak
blades,the apex produced into a spine; surfaceof the expandedouterportion with abundant setae.
Habitat.-Peru (Junin). Holoty?e, d, Chanchamayo, altitude
L200 meters, January 20, 1949 (Schunke).
Most similar to Gonomyia(Gonomyia)platymera Alexander,which
has the inner dististyle of the male hypopygium somewhat the samebut differs in all other details of structure of the organ.
Gonomyia (Neolipophleps)topoensis sp. n.
Belongs to the f alcifer group; male hypopygium with the outerdististyle a strong blackenedrod, its apex extendedinto a long glabrouspoint, the base more expanded, its surface microscopicallyscabrous;
inner style producedinto two unequal blackenedspines,the pale outerportion with three fasciculatesetae.

134

Annals Entomologr,cal Society of America[Vol. a5

Male.-Length, about 4 mm.; wing, 4.3 mm.
Rostrum and palpi black. Antennae black, the basal segments
variegated with obscure yellow; flagellar verticils of male very long.
Head above chiefly obscure yellow, variegated with brown.
Pronotum and pretergites light yellow. Mesonotum brownish gray,the central area of the scutum more reddish brown; posterior border ofscutellum narrowly yellowed. Pleura ventrally dark brown, sparsely
pruinose; a conspicuouswhite longitudinal stripe extending from andincluding the fore coxae,reachingthe baseof abdomen,passingbeneaththe halteres, bordered above by a narrow dark brown line; dorsal
pleurites and dorsopleural membrane chiefly pale. Halteres with stem
yellow, knob infuscated. Legs with the coxae and trochanters yellow;
only the posterior leg preserved,femora yellow with a vague to scarcely
indicated narrow brown subterminal ring; tibiae pale ye1low, the tip
narrowly dark brown; tarsi dark brown, the proximal part of basitarsi
pale. Wings grayish subhyaline, the prearcular and costal fields morewhitened, including more extensive areas before and beyond the pale
brown oval stigma; veins pale brown, whitened in the costal field, thosecomprising the cord a trifle darker. Venation: .lc short, Sc, ending a
distance before origin of Rs that is approximately equal to the latter
vein; cell M2 openby the atrophy of the basal sectionof vein Msi m-cu
at or close to fork of.M .
Abdomen chiefly dark brown; hypopygium chiefly infuscated, thebasistyles and inner dististyles extensively more yellowed. Male
hypopygium with the outer dististyle a strong blackenedrod, expanded
at beyond one-third the length and thence narrowed into a long glabrouspoint; inner margin and surfaceof the expandedportion microscopicallyscabrous,the teeth larger and more conspicuousat the outer angle ofthe expanded part; a long straight spine at near midlength of basal
part of stem. fnner dististyle produced into two unequal blackened
spines, the paler outer part with three fasciculate bristles and a few
other setae. Basistyle at apex produced into a small blackened conicalpoint.
Habitat.-Ecuador (Tungurahua). Holotype, d, Upper Rio Topo,
altitude 1500meters,February 1949(S. Velastegui,through Macintyre).
The closest relatives of the present fly include Gonomyia (N eolipophleps)falciJer Alexander, G. (If.) neofalciferAlexander, and G.
(ff.) subfalciferAlexander, all of which differ from the present fly andamong themselvesin the details of structure of the male hypopygium.

Genus Toxorhina Loew
vulsa sp. n.
Toxorhina(Ceratocheilus)
General coloration gray, the praescutum with three virtuallyconfluent brown stripes; legs black, including the tarsi; wings with aweak lrrownish tinge, the extreme base yellowed; veins dark brown;,Screlatively long, ending oppositeone-third the length of Rs, the lattera trifle shorter than the basal section of Rs; anterior branch of Rs longand sinuous, at least twice as long as Rs,' male hypopygium with theouter dististyle stout on the proximal two-thirds, the apex cylindrical,
narrowed; inner dististyle with base enlarged,the outer half or more

lg52lAlexander: SouthAmerican Crane-Flies.XIV.

135

narrowed; each arm of aedeagusflattened and with a strong carina onouter half.
Male.-Length, excludingrostrum, about 6.5 mm.; wing, 6.2 mm.;rostrum, about 5 mm.
Rostrum black, elongate,as shownby the measurements. Antennae
black throughout. Head gray, the center of vertex extensively
blackened.
Pronotum and cervical region dark gray. Mesonotum light gray,the praescutumwith three virtually confluent ill-defined brown stripes,
the scutal lobes similarly darkened. Pleura gray, variegated withd.arker, especially on the anepisternum and ventral sternopleurite;
dorsopleuralmembranedusky. Halteres with stem yellow, knob darkbrown. Legs with the coxae gray; trochanters brown; remainder oflegs black, including the tarsi, only the femoral bases restrictedly
brightened. Wings with a weak brown tinge, the extremebaseyellowed;
veins dark brown. Venation: Scr ending about opposite one-third thelength of Rs, the latter a trifle shorter than the basal sectionof vein Ro;
anterior branch of Rs long and sinuous,two or more times Rs itself;
Ronot as stronglydecurvedat tip as in superstes,'innerend of cell lst Mz
more pointedthan in superstes.
Abdomendark gray. Male hypopygiumwith the basistyleproduced
at apex into an elongate spine. Outer dististyle with the proximal
two-thirds stout, thence suddenly narrowed into a more slendercylindrical portion, the tip obtuse and blackened. fnner dististyle with thebase enlarged, setiferous,the outer half or more narrowed, the apex
with a few microscopic setulae. Aedeaguswith the arms flattened
and with a strong carina running back from the acute tips for about
one-halfthe length.
Habitat-Peru (Junin). Holotyqe, d, Chanchamayo, altitude
1200meters,April 29, 1948 (Schunke).
The only similar described speciesis Tororhina (Ceratochei,lus)superstesAlexander, which differs in the venation and in the structureof the male hypopygium.