r4a>(lr UISTII{DIAIIIIEW(lRLITTLE.TilOWN
SPECIES(DIPTERA).
T|PUUDAItV

By:

GhuluP. Alcnnder

Reprinted fromfheJournrl
of Agiculturc
ol thcUnircritfol PlcrtoRico
Vol, XXfff, No. z-Pagcs gr to ryo.
Afril, rg7g.

NEW OR,LITTLE_KNOWN SPECIES OT WEST INDIAN
TTPUT.IDAE(DIPTERA) IV
By Crlnr,ns

P. ALn'ANDER, *The third part under the above title appeared in this Journal,
vol. XXr, No. 4, Oetober 1937. The present report is baseclalmost
entirely on extensive collections of erane-flies that were taken on
various'West Indian islands, ehiefly by Mr. P. J. Darlington, Jr., andby Mr. Marston Bates, and are now preserved in the Museum of
Comparative Zoology.Cambridge, l\[ass. The speeiesthat were takenon the higher mountains of Haiti and the Dominican Republic proved
to be of especialinterest ancl value.
fn a preliminarl' section.I am cliscnssinga small but very interesting eollectionof theseflies that rr'astalien in the island of Dominica,
I:esser Antilles, by 1\Ir. Walter II. Hodge, at the present time a
student at the Gray Herbarium, Harvard University. The specimens
in this latter series are preserved.in my personal eolleetion. I wish
to expressmy sincere thanks and indebtednessto l\fessrs.Bates. Darlington and rlodge, and to Dr. Nathan Banks of the Museum of
Comparative Zoology, for the privilege of studying this exceptionallyinteresting series of Tipulidae.
I

Tipulidae from the fshndof Dominica

The speeiesherewith recorded appear to be the first that have beentaken in Dominica, although rather numerous speciesare known from
St- Vincent. A most interesting general aecount of eonditions in theisland is given by F rederiek A. ober, camps in the caribbees (Edinburgh; David Douglas), pp. 1-860; 1880. Mr. rrodge, colleetor of
the present material has supplied the following notes eoncerning the
localities where the more interesting speeieswere seeured.
Morne Trois Pitons.. A voleanic peak situated roughly in thecenter of the island and a peak whieh has at times been claimed tobe higher than the generally acceptedtallest peak, Morne Diablotin.
According to Dr. Paul Griswold rrowes, of Greenwieh's Bruce Museum, Trois Pitons is 4,600 feet. The erane-flieswere eollected on
one of the two lesser summits, altitude 4,b00 feet, in typical winddwarfed, rain-swept cloud forest, the branches of the trees dripping. M"*".fh*etts

State College, Amherst, Maesachusetts.
9l

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On AGRI(:LTLTUREOF' THE UNMRSITY

OF P. R.

with mossesand hepatics ancl laclenrvith other epiphytes, particularlywater-containing Bromeliads. Here the eane-flies (Tangprem'n('hoilgei (sp. n.) occurred in dozens,many mating. The few I managed
to pick up I had. to stalk after they had alighted beneath the shelterof the leaves. During the sgmmer months, part of the rainy season'
the summits of all of Dominica's mountain peaks are almost continually enshroudedby rain clouds. Rainfall near the summits must
exceed300 inches per Year.
IIattort, Garden Trail' An old Carib footpath running throughvirgin tropical rain forest from Sylvania Estate, llear Trois Pitons,
to llatton Garclen Estate on the northeast (windward) coast. Craneflies were'secured by sweeping the vegetation along the Layou Rivernear its source in the neighborhood of the old abandoned Riversdale
estate. Altitucle approximately 2,000 feet; rainfall about 200 inches.
River bed rocky and boulder-fiIled. Stream turbulent.
Syluania Estate.. Near the base of Trois Pitons (west). Altitucle
1,800feer; rainfall betrveen1?5 ancl 200 inches. Surrounding forests
can be called true ''rain-forests".
Tanypremna (Tanyprem,n'u) hodgei sp. n.
General coloration of mesonotum dark brown; pleura med.ium
brown, unmarked; halters relatively short, dark brown throughout;legs brownish black, passing into black on all tarsi; wings with astrong brown suffusion, cell Sc and the stigma darker brown; cell
7st XIz short ancl wide; abdominal tergites dull black, before midlength ringed, with yellow, the outer segments and hypopygium uniformly black; male hypopygium with the dististyle at base on outermargin proclucedinto a pou'erful spine.
I\Iale.-Length about 14-15 mm. ; wing LI.5-12.5 mm. ; antennae
1.4 mm.
Female.-I-iength about 17-l-9 mm. ; wing 13-14 mm. ; antennae
1.4 mm.
X'rontal prolongation of head very short, reddish brown; palpi
black. Antennae shorter than palpi; basal segments brownish yellow, the outer onesmore darkened; basal flagellar segmentsenlarged,
the outer ones linear, with long conspicuous verticils. Head. reddish
fulvous, variegated. with d.arker on vertex, expeclally across theanterior vertex as a more or less continuous band.
Pronotum yellowish brown. Mesonotal praescutum chiefly covered.
by four dark brown, subnitidous stripes, the intermediate pair separated by a eapillary darker vitta, the posterior interspaees searcelyindicated; posterior sclerites of notum dark brown, the sides ofNEW OR, LITTLEKNOWN

TIPULIDAE

93

mediotergite a little paler. Pleura medium brown, unmarked.. Halteres short, dark brown throughout. Legs with the eoxtre atrtl tlochanters pale brolvn; femora and tibiae brownish black, the tips
someu'hat more blaekenecl; all tarsi blaek. Wings (Fig. 1) with astrong brown suffusion, cell ,Sc and the stigma darker brown; paler
streaks in centers of some of the cells. espeeially in female; veinsbrownish blaek. Yenation: ,Sc1,free tip of Sc2 and .81 * , all relatively close together at wing-margin, the distance between them lessthan vein R1 -F o alone; eell 7st M2 short and wide, nl-c'tt,at near twothirds its lorver faee; petiole of eell Mr variable in length, but approximately equal to m.
Abdomen elongated; tergites ringecl rvith yellorv and dull blaek,
the latter incrluclinglnore than the outer' half of the segment,as well
as a ver-y narro\\- darliening at base. restrieting the 1'ellorvto theouter portion of llastrl rings; outel' segntetrtsancl h1-llopr-gittrnunithe posteriot'lrordersof thefonull' blacli; slet'llitesrllolt: 1-ellou'islr.
more basal segments broadll' hrou'n. passing into blacli on outersegrnents. In the fernale, the l ellol' r'olol' is nrol'e obseurecland
pruinose. IIale h;'pop;'gium rvith the outer basal portion of clististyle (Fig. 10, d) proclueeclinto a strong blackenerlslrine; beakof st-vleon outer margin before apex rvith one or two small blaekened
pegs. Ovipositor rvith valves sclerotizecl,relatively slender.
IIabitat.-Dominica.
Holotype, 6 , llorne Trois Pitons, altitude 4,500 feet, August 15,
1 9 3 8 ( H o d g e ) . A l l o t o p o t - v p e ,9 . P a r a t o p o t y ' p e sA, A , 1 9 .
To,nyprenlnil (Tangprentna) hod,gei is named in honor ofl }tr.
'WalterII. Hoclge who eolleeted this important series of Dominican
Tipulidae. It is yer)' different from the other speciesof the typicalsnbgentrsthat have darkened tarsi, as 1. (1.) carbonipcs Alexancler(Eenador), I. (1.) fusci,tarsisAlexander (Colombia), T. (T.) k,udeni
Alexander (\renezuela T. (T.) inuaripes Alexander (Brazil), espeeially in the eoloration of the thoraeic notum and pleura, the venation,and the strueture of the male hypop)'gium. The genns hitherto had
not been detected in any of the'West Indian islands, on the mainland
extending from southwestern Mexieo southward to southeastern
Brazil.

Limonia (Lim,onia) apicata dominicensis subsp.n.
l'Iale.-Length about 5 mm. ; rving 5.5 mm.
Female.-Length about 7.5 mm. ; wing 7 mm.
Charactersas in typieal apicata Alexander (ranging from northern
Florida, subspecies ntbapica,ta Alexander, to Peru, subspeciesna-

94

TEE JouRNArr oF aGRrcurJTUREoF TrrE uNrvERsrry oF p. R.

poensisAlexander), differing only in slight hypopygial eharacters.
lVlalehypopygium with the gonapophyses(Fig. 13) very broadly expanded and compressed,before apex bearing a small spinous point.
Habitat.-Dominiea.
Holotype, 6 , Layou River, on Hatton Garden Trail, altitude2,000feet, August 16, 1938 (Hodge). Allotype, 9 , Sylvania Estate,
altitude 1,800 feet, August 29, 1938 ('Hodge).
Limonin (nhifidia)

domestica(Osten Saeken).

Dominica: Marigot, July 1938 (Hoclge). Lisdara Estate, altitude1,800feet, August L937 (Hodge).
Limonia ( Rh,ipid,in) costal,is(Williston).
Dominica: Sylvania Estate, altitude 1,800feet, August 29, 1938
(Hodge).
L,i,mon,ia( Geranorny,in)cinereinota (Alexander).
Dominica: Layou River, on Hatton Garden Trail, altitud.e2,000
feet, August 16, 1938 (Ilodge). Hitherto from Greater Antilles;
northern South America, south to Paraguay.
Limonira ( Geranomy,in) dominicana sp. n.
Mesonotal praescutum pale greX, with a broad, yellowish brown,
median stripe, the lateral bordersof the seleritebroad.lydark brown;knobs of halteres dark brown; femora yellow, with a narrow brown
subterminalring; wings yellow, variegatedwith pale brown, the darkareasat origin of Rs and fork of ,Scbarely connected; Sc short, ,Scr
ending about opposite one-fourth the length of Rs; abdominal segments yellow, narrowly ringed caudally .rvith brown; male hypopygium with the rostral prolongation of ventral dististyle short, witha powerful common tubercle bedring two unusually long curved.
spines; aedeagusat apex with two conspicuouslobes.
Male.-Length, excluding rostrum, about 7 mm. ; wing 6.8 mm.;rostrum about 3 mm.
Rostrum and palpi black, the former elongate. Antennae withscape and pedicel black, flagellum pale brown; flagellar segments
oval. Ifead dark gray, the anterior vertex and a central line backto occiput light gray.
Mesonotal praeseutum pale grayt rilith a broad. yellowish brown
median stripe that is a little darker in front, on posterior half narrowly delimited by delicate dark margins; lateral praescutal borders
broadly and conspicuouslydark brown; scutellum and median region

NEW OR LITTI,E

KNOWN

TIPI'I,IDAE

of scutum pale testaeeous,the scutal lobes d.arker,delimited internallyby a narrow dark line; postnotum pale. Pleura yellow, variegatedwith brown, especially on dorsal portion. Halteres with stem paleyellow, knob dark brown. Legs with the coxae yellow, fore pair a
trifle darker; trochanters yellow; femora obscure yellow, with anarrow, dark brown, subterminal ring, about equal in extent to theclear yellow apex; remainder of legs obscure yellow, the terminal
tarsal segmentsa little darker. Wings yellow, the eostal border moresaturated; a conspicuouspale brown pattern, as follows: A seriesof five costal darkenings, the first at arculus, second.at supernumerarycrossveinin cell ,Sc,both small; third area larger; at origin of Rs
and fork of ,Sc,barely interconnectedand.forming an irregular area;
stigma relatively small; fifth darkening small, at end of vein ,Rs;cord and.outer end of cell lsf ,412seamedwith brorvn; small marginal
dark clouds at ends of veins 1113
ancl J14,\\'ith larger and more conspicuous ones on veins Cu, 1st /. ancl 2ncl A: veins yellow, slightly
darker in the clouded areas. Venation: ,Screlatively short, Sc1 ending about opposite one-fourth the length of Rs, Scz at its tip; cell
7st Mz about as long as vein Mt * 2 beyond it; nl-cu at fork of trI.
Abdominal segmentsyellow, narrowly ringed caudally with brown;
hypopygium yellow brown Male hypopygium with the caudal
margin of tergite nearly truncate or very feebly emarginate. Yentral dististyle much more extensive than basistyle; rostral prolongation short and stout, more than one-half of its outer margin involvedin a large tubercle that bears the two long spines; these latter borne
at tip of tubercle, subequal in length, strongly curved., exceedingin length the dorsal dististyle; base of tubercle expanded proximad
into a flange; apex of rostrum beyond tubercle very short, only
about one-third the length of tubercle. Gonapophyseswith nesalapical lobe blackened, expanded, at apex narrowed. into a very shortdecurved beak. Aedeagus short, at apex with two conspicuouslobes.
Habitat.-Dominiea.
Holotype, 3 , Sylvania Estate, altitude 1,800 feet, August 30,
1938 (Hodge).
The most similar described speciesinclude Limonia (Geranomg,in)carunculota Alexander and L. (G.) deliciosa Alexander, both ofwhich are readily distinguished. by the structure of the male hypopygium.
Linonia( Geranomg'tn) microphaea sp. n.

Size very small (wing, d, less than 5 mm.) ; general eoloration
brownish black, unmarked; rostrum elongate, about twothirds the

96

THE JOURNAIJ OF AGRICUI-,TURE OF TIIEUNMRSITY

OF P. R.

length of body; halteres brownish black, the base of stem obscureyellow; wings strongly tinged with brown, stigma darker brown;
Sc relatively short, Scr ending about opposite one-forth to one-fifth
the length of Rs; cell 2nd A narrow, espeeially on outer half ; abdominal tergites brownish black, basal stertrites obseure 'r'ellow;
hypopygium uniformly blackened; male hypopygiurn with the ventral
dististyle much more extensive than the basistyle; rostral prolongation long and. slender, at base with a low common tubercle from thesummit of which arise two straight spines of moderate length, thesespirresshorter than the prolongation; gonapophl'sesrvith mesal-apicallobe a short, curved black spine.
Male.-Length, excluding rostrum, about 4 mm. ; wing 4.8 mm.;rostrum about 2.8 mm.
Rostrum very long, about two-thirds the length of body, black,the distal thirtl of paraglossaepaling to brown. Antennae broken.
Head dark.
I,Iesonotum almost uniformly dark bron'n to brorvnish black, thesurface polished, the humeral region of praescutum a little brightened. I'leura dark brown, the ventral portion of sternopleurite paler.
Halteres brorvnish black, the base of stern obscru'eyellol'. fregs with(:oxae brownish yellorv; trochanters yellorv: remainder of legs
hrokeu. Wings strongly tingecl u'ith brorvn. cells (i ancl 8c a trifle
rlarker; stigma oval, darkel brornr ; veins darrk brorur. Venation:.Sc relatively short. Sr,r encling about oplrosite one-fourth to onetifth of .R.s,Ncs at its tip; Fs relativell. long, fulll- three times thearcuated basal section of 1?a+5 ; cell 7sf i12 lleorl;' as long as vein
Mt* 2 beyond it; nr,-cucshortly before fork of llI: eell 2nd r{ narrow,especially or outer half.
Abdorninal tergites brownish black; basal sternites obscure yellou'1 hypopygium uniformly blackened. Male hypopygium (Fig. 15)
l'ith the eaudal margin of tergite with a broacl U-shaped notch, thelateral lobes broad. I)orsal clististyle only gently curved, the tip aslender spine. \rentral dististyle large and fleshy, mueh more extensive than basistyle; rostral prolongation long and slender, at baseon outer face rvith a lorv common tubercle, from summit of whicharise trvo relatively short, straight spines, these shorter than the
prolorrgation distad of tuberele. Gonapopyses'with mesal-apicallobe
a short cnn'ecl black spine, the tip acute.
Habitat.-Dominiea.
Holotype. 6 , La.vou River, on llatton Garden, altitude 2,000 feet,

August16, 1938(Hottge).

NEW OR LITTLEKNOWNTIPUI,IDAE

Limonia ( Geranomg'ia) microplw,ea is somewhat similar to L.
(G.) c'erberusAlexander and -L. (G.) fl,una Alexander, differing especially in the small size, uniformly dark brown mesonotnm, andstructure of the male hypopygium.
Limon'tn ( Geranomg'in) pltcmbeipleura Llexander.
Dominica: Sylvania Estate, altitude 1,800 feet, August 28-30,
1938 (Hodge). F ormerly known from northern South America andTrinidad.
Limonia ( Geranomyin) sylvania sp. n.
Allied to plumb.eipleu,ra; size small (wing, 6 , under 6 mm.) ;mesonotum buffy brown, with three brown praescutal stripes; femora
obscureyellow, rvith a narrow brorvn subterminal ring; wings yellow,
patterned rvith brown, the clark area at origin of Rs also involving
the fork of ,Sc; abdominal segmentsdimidiate, the bases darkened,
the apices yellorv; male hypopygium l'ith the spines of the rostralprolongation of ventral dististyle arising from tubercles that aremuch shorter than the spines themselves; mesal-apicallobe of gonapophysis a short blackened horn.
Male.-Length, exeluding rostrum, about 4.9-b mm. ; wing 5.45.8 mm.; rostrum about 2 mm.
Rostrum black, relatively short. Antennae black throughout..
Head black, the median region of vertex lined with light gray.
lVfesonotum obscure buffy brown, rvith three distinct but somewhat poorly delimited brorvn stripes; scutum buffy brown, the lobes.
variegated l'ith darker brorvu; posterior sclerites of notum ehiefly
darkened. Pleura with propleura obscureyellow; dorsal mesopleuraand pleurotergite rveakly darkened, the sternopleurite and meron yellou'. Halteres obseureyellorv, the baseof stem a little brighter. Legs
rvith the coxae and trochanters yellon': femora obscure yellow, witha narrow brorvn subterminal ring, the yellou' apex beyond slightly
wider; in cases,the femora are darker, more brorvnish yellow, and.
the subterminal dark ring is preeeded by a narrow, clearer yellow
annulus; tibiae and tarsi yellow, the outer tarsal segmentsdarkened.
Wings yellow, more saturated in the costal and prearcular fields, the.
apex, in cases,weakly darkened; a sparse brown pattern, including
four subcostal areas that are inore restricted. than the interspaces;
first area above arculus, tending to becomeevanescent; second.area
at supernumerar;r crossvein in cell Sc,. third area large, irregUlar
in outline, common to the origin of Rs and fork of Sc,. fourth arealargest, at stigma; narrorver brown seams along cord, outer end of

98

TIIE

JOURNAIJ OF AGR,ICIIIJTURE OF TIIE

UNIVERSITY

OF P. R.

cell lsf ]Iz and,on distal portion of vein 2nil a; veins yellow, darker
in the infuseated areas. Venation: Sc relatively short, ,Sc1endinga short distance beyond. origin of ,Es, at about opposite one-fifth thelength of this vein; nL-cILat fork of M.
Abdomen more or less d.imidiate, the bases of the segmentsnarrowly darkened, the apices more extensively yellorv; subterminal
segments uniformly darkened; hypopygium yellow. Ifale hypopygium (Fig. 16) 'with the caudal margin of tergite broadly and' conspicuously emarginate, the margins of the lobes broadly thickened
and. sclerotized; tergal setae coarser and. more sparse than in plumbeiqtleura. Dorsal dististyle a curved sickle. Ventral dististyle large
ancl fleshy, the rostral prolongation. and spines much as in Ttlumbeipleura; rostrum nearly parallel-sided, scarcely expand.edoutwardly ;lpir.es equal in size, the tubercle of the lolver or more cephalic spine
slorter, less than half the spine; both tubercles much shorter than
the spines they bear, the outer being about two-thirds its spine.
Gonapophyseswith rnesal-apical lobe appearing as a short blackened
horn.
Habitat.-Dominica.
Holotype , 6 , Sylvania Estate, altitud.e 1,800 feet, August 28,
1938 (Hodge). ParatoPotYPe,6 .
Although allied to Limonia (Geranom'yia) plunfteipleura Alexander (Dominica and Trinidad; northern South America) and to
L. (G.) mctrtlaaeAlexancler (llt. Santa nlarta, Colombia), the present
fly is distinct in the characters listed in the diagnosis. The size
ancl the general structure of the rnale hypopygium is more as inmarthae but the coloration of the legs and. details of the hypopygium
in the latter, especiallythe very large ventral dististyle and its rostralspines, indicate a distinct species. AII three forms constitute a groupof closely interrelated' flies.
Limonia (Geranomgia) tibinlis (Loew).
Dominica: Sylvania Estate, altitude 1,800 feet, August 30, 1938
(Hodge).
Polymera (Potymer'u) albitarsis dominicae subsp. n.
Male.-Length about 4.5 mm.; wing 5.2 mm. ; antennae about
6 mm.
Differs from typical olbitaris \Yilliston (St. Vincent), as follows:
Size small. trIesonotum uuiformly dark brown, only the humeral
region and. pretergites restrictedly obscure yellow. Pleura chiefly

NEW

OR I,ITTI,E

KNOWN

TIPII,IDAE

99

covered.by a very broad., black stripe, only the stenopleurite antl
ventral meron abruptly yellow. Legs with fore coxae blackened' on
basal two-thirds, the apex and remaining coxae light yellow; femora
brown, passing into dark brown, the tips abruptly whitened; tibiae
dark brown; outer tarsal segmentswhitened, in degree less than in
typical albitarsis, the proximal half of the second tarsal segment of
posterior leg clarkened.; in atbitarsis, on the posterior legs, only thebasitarsi are darkened. Wings very strongly suffused with brown;veins darker. Venation: ,Sc short, ,Scr ending opposite fork of thelong, nearly straight, fully as long as Rs andshort Rs; Rz*r*r
nearly twice Rz* s; Rt * 2 and Rr* s subequal.
Habitat.-Dominica.
Holotype, 6 , Sylvania Estate, altitude 1,800feet, August 30, 1938
(Hodge).
Bhabdomastix fumipennis sp. n.
General coloration brorvnish black, the thorax unpatterned; antennae black; halteres and legs brorvnish black; wings with a strong
brown suffusion; stigma ill-defined, very slightly darker than the
ground; Es approximately twice R, * r * n; abdomenblack.
Female.-I-:ength about 5.5-6.5 mm. ; wing 4.5-5 mm.
Rostrum and palpi brownish black. Antennae black throughout,of moderate length (female), if bent backward.scarcely attaining thewing-root; flagellar segments subcylindrical. Head blackish.
Thorax uniformly brorvnish black, without markings. Halteresbrownish black, the extreme base of stem pale. Legs with the coxae
brownishbrownish black; trochanters testaceous; r'emainder of legs
-Wings(Fig.
black, the subterminal tarsal segmentsa little paler.
9) broacl, rvith a strong brorvn suffusion; stigma ill-defined, very
stightly darker brorvn than the glottncl; veins brorvn. Wing-veinschiefly naked; a series of trichia on outer ends of veins .Es, Mr * z
and,Il[s; in cases,a single or very sparse trichia on Ma and distal
section of Cut. Venation: Rs considerably longer than in paruula,
a littte more than twice R, * r * n ; R+ slihtly variable in length,
subequal to or shorter than Rz* e * n.
Abdomen black. Yalves of ovipositor, especially the cerci, elongate and very slender.
Habitat.-Dominica.
Holotype, 3 , Sylvania Estate, altitud.e 1,800 feet, August 29,
1938 (Ilodge). Paratopotypes,7 I9.

100

oxt P' R'
TrrE JOrrRNArr oF AGRICITTTUREoF TEE UNTSERSITY

to be closed to R'
Rhabil,omastir fumipentc'is seems undoubtedly
whieh belongs
paruulo Alexander (parua Alexand.er, preoecupied),of the maleto the subgenus Sananila,ga.Llexand,er. In the abserice
refer it to a given
sex of the present fly, it seems inadvisable toin both of thesubgenus since the antennae are short in the females
s' s', have theregional subgenera; the males of Rhabdomasti"rSkuse,short as in theantennae much longer than the body, of sacanrlaga,evidently in thefemales. F rom paru%la, the present fly differs mostthe long Rs'
broacl rvings with the venational details distinct, notably
Erioqttera ( lllesoclrlTthonn)caloptera Say
29, 1938
Dominica: sylvania Estate, altitude 1,800 feet, August( Hodse ) .
' Toxorhina (Tororhina) fumipennis sp' n'lateral borders
General coloration of mesonotum dark brotTn, thebrown;of praescutum broadly buffy gray; halteres and' Iegs darkveins I][s + 4
wings with a strong brown sufrusion, the veins darker;
of .'ll'and,M+ subequal; rm-c%about one-third its length before fork
5'2 mm'Femaie.-Length, exclucling rostrum, about 7 mrn.; \vingRostrum black, its apex broken. Head dark gray.
praescutum
cervical region and pronotum blackenetl. ]Iesonotal
gray; posterior*.it1 cLisl<browl, the lateral margins broadly buffy
centralsclerites of noturn d,ark, sparsely pruinose' Pleura withportion chiefly infuscated, paler dorsally and ventrally' Halteres
suftlarli brorvn. I-regsrlark brolvtr. Wings rvith a strong brown
fusiorr. uripattertrecl; veins darker ihan the ground' Venation: rScr
of M1* 'ending opposite origin of Rs, ,Scznear its tip ; basal section
y'n-ct'L
abont two-thirds the second.; trIs * a subequal to II + alone i
nbout tlue-thilcl its leugth before fork of XI'
Abclominal tergites brownish black, sparsely pruinose; stmnites
paler.
IIabitat.-Dominica.
.-olype, 6 , sylvania Estate, altitude 1,800 feet, August 2i,
1938 (Hodge).
The strongly infumed wings of the present species read'ily distinguish it from all other members of the genus so far describedin the Neotropical fauna.

NEW OR LITTLE KNOWN TIPULTDAE

101

II. Tipulidae from the Greater Antillean fslands. Descriptions of
new speciesand recordsof distribution from Cuba,
Hispaniola,Jamaicaand Puerto Rico.
Tipula darlingtoniana sp.n.
Size large (wing, 6, over 25 mm.) ; mesonotumbuffy, withfour entire brown stripes, the interspaces rvith brown setigerous
punctures; antennae yellow; legs with femora obscure brownish
yellol,, the tips blackened; rvings whitish subhyaline,patterned'withof brown; ns long, about three times rL-cIL; abdominal
various shacles
tergitesbuffy, quadrivittate with brown; cerci long, slender,straight'
Female.-Length about 31 mm. ; wing 26 mm.
Frontal prolongation of head brorvnish gray above, brown on,sidesand beneath; nasrlslong autl slenclet':palpi black, the apicesof the intermerliatesegntentsnarrorvll- 1-ellorv. '\tttt'ltttite1-ell,rrv:{irstflagellar segrnentlonger than the scape; r'eriicils of intermediatesegmelts exeeeding the segments in length. Ileacl graf ish brorvn,
rvith a narro\\', clark brorvn, meclianvitta on posteriol vertex; vertical
tubercle relatively low.
Pronotum b1ffy, rvith three brown areas, the interspaces withbrolvn setigerous punctures. Mesonotal praescutum buffy, rvith four
entire clarli broln stripes, the intermediate pair eontiguous or narrorvly- confluent at ends, the median portion a little infuscated and
Iined eentrally with darker, humera region of praescutum extensively clark brown; intersl)aceswith conspicuousbrown setigerous
punetures; scutal lobes brownish groX, narrorvly darker brown on
mesal eclge: meclian region of praeseutum gr&y, with a eapillarydarker vitta; sttntellum rvhitish groy, rvith a meclian brown vitta,
parascltella cltrrk; mediotergite Sray, the central fourth rvhitish
groy, borclereclsublaterally by dark brown. Pleura brownish gray,clearer gray on ventral sclerites, sparsely variegated with browtr on
dorsal anepisternum and sternopleurite, and, especiallY,'or the pleurotergite. Halteres rvith stem yellow, knob infuscated. I-regs withthe coxae gray, rvith brown punctures; trochanters obscure yellow;
femora obscure brownish yellow, the tips rather narrowly blackened;
tibiae obscure yellow, the tips passing into black; first and second.
tarsal segments obscure yellow, the tips darkened; remainder oftarsi blaek. Wings whitish subhyaline, conspicuously patterned withvarious shadesof brown; prearcular field. chiefly darkened, its outerend. white; eell C uniformly brorvn exeept at outer end; cell ,Sc
brownish yellow, with four major areas that extend. into cell R, theseL02

TrrE JoURNAIJ oF AGRrcurrTUREoF THE uNrvnnsrrY oF P. B.

pale brown, bordered posteriorly by darker brown; outer radial
field pale brown, bordered. behind by a dark brown seam on outer
two-thirds of vein Re* a; basal two-thirds and outer end of ceILM
brown, the latter area paler, the cubital veins narrowly bordered bydark brown; outer portions of Anal cells pale brown; cell Cu andbasal third of cell 7st A washed with darker brown; restrictecl
darker brown seams at outer end of cell lst M, * 2, flI'-cv.land veinsat base of cell 1st M2, and on vern 2nd, A; veins brown, a little palerin the ground areas. Yenation: Distal half of vein R1 * 2 pale but
lrreserved; ns very long, about three times as long as m-cu; petioleof cell M1 longer than nr,; basal section IlIa ercet, exceeding trIs * a.
Abdominal tergites rvith the ground color buffy, variegated by.two submedian dark stripes that are separatedby a capillary medianground vitta; additional sublateral brorvn stripes, the interspaces
u'itlr. brolvn setigerous punctures; sternites chiefly dark brown, variegatecl by buffy. Ovipositorwith long slender straight cerci; hypovalvae stouter.
Habitat.-Dominican Republic.
Holotype, 6 , Valle Nuevo, southeastof Constanza,altitude about
7,000 feet, August 1938 (Darlington).
I take very great pleasure in naming this striking and. distinet
fly in honor of the collector,Dr'. P. J. Dtrllington. Jr., l'ho has added
so materially to our linol'leclgeof the Tipulidae of llispaniola. The
speciesappears to belong to the glaphyroptcro group of the genus,
including a vast aggregation of South and llidclle American forms,
but not previously knorvn from the West Iudian islands. It is amply
distinct from all other knorvn species, especially in the colorationof the antennae, mesonotum,legs ancl 'rviugs.
Tipula (YamatotiTtu,lu) ludouic'inna Alexander.
Cuba: Santiago de las \regas, I\Iarch 31, 1931 (A. Otero) ; April
20, 1931 (A. Cremata).
N epltrotoma circumscripta (Iroew) .
Cuba: Pico Turquino, Oriente altitude 5,000 feet, June 1936 (Juli6n Acuna ) .
Nephrotoma glabricristata sp. n.
Allied to fercugineal basal three antennal segmentsyellow, thesueeeeding segments varying from uniformly blaek to weakly bicolorous; mesonotal praescutum yellow with three more nacreoru;
stripes that are very narrowly bordered in part by dark brown;

NEW OR LITTLE KNOWN TIPULIDAE

103

mediotergite with a brown eentral stripe on anterior half ; wings narro.!v,with a faint brown tinge, the stigma and costal border darker;
wing-tip and seams along veins darkened; a few macrotrichia iu
outer end,of eell R3; abdomenorange, sparsely patterne,ilwith darker;
male hypopygium with the dorsal crest of the inner dististyle highand entirely glabrous.
i\tale.-L,ength about 10 mm.; wing 9.5-10 mm.; antennae about
3.3 mm.
tr'emale.-Length about 15 mm. ; wing 11-11.5 mm'
Frontal prolongation of head polished. yellow; nasus d'istinet;
palpi yellow, the terminal segment a little darker. Antennae relatively short; basal three segments yello'w, sueceeding segments
varying from unifonn blacli to rvealily bicolorecl, rvith the basal
enlargement of the segrnentblackenecl,the apex paler (this lattercondition in holotype) ; flagellar segnrentsralher strongly incised.
Heacl orange; occipital braucl of rnocleratesize, elongate oval, withthe anterior end pointed, poorly defined.
Pronotum darkened laterally, clear light yellow medially above.
Mesonotal praescutum yellow, with three more nacreotls stripes thatare very narrowly bordered by dark brown; on the lateral stripes
this border is restricted to inner margin; lateral stripes straight,
without clearly defined, marginal marking opposite anterior end';
scutum yellow, each lobe entirely traversed by posterior extensionsof the lateral praescutal stripes, these not margined with darker;
scutellum darkenecl, parascutella pale; med.iotergiteyellow, with abrown central stripe that is narrowed or obliterated on posteriorhalf, posterior border of sclerite narrowly darkened. Pleura yellow,
variegatecl with reddish brolvn, especially on propleura and on ventral anepisternum, sternopleurite, meron and pleurotergite. Haltereswith stem obscure yellow, knob infuscated. Legs with the coxae
reddish yellow; trochanters yellow; femora yellow, the tips very
narrowly blackened; tibiae pale yellorvish brown, passing into darkbrown; tarsi black. Wings narrow, with a faint brown tinge; cell
Sc and stigma brown, cell C and prearcular field more yellowish
brown; wing-tip and very narrow seamsalong eord brown; a brown
seam along vein Cu; veins dark brown, narrowly and vaguely seamedwith brown. A ferv macrotrichia in outer end of ceII Rr, rvith a few
others at proximal end of stigma. Venation: rscz ending opposite
origin of the short oblique Rs, the latter in virtual alignment' u'ith
Rt * e; eell Mr narrowlY sessile.

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TEE JOURNAIJ Or AGIiIOUIJTUIiE OF TFIE UNMRSITY

OF P. R.

Abdomen orange, both tergites and sternites with a faint dusky
median spot shortly before apex; hypopygium orange. Male hypopygrum of the general type of fercuginea; inner dististyle with thed.orsalcrest unusually high and.entirely glabrons; in f errugineo, theentire margin is provided with abundant short setae.
Ilabitat.-Cuba, Ilispaniola.
Holotype, I , I-,lomadel Gato, Sierra clel Cobre, Oriente, Cuba,
altitucle about 3,000 feet, July 3-7, 1936 (Darlington). Allotopotype, g, Buenos Aires, Santa Clara, Cuba, July 19, 1932 (Bates &Fairchild). Paratopotype,6; paratypes,16, 2 g 9, with allotype, JuIy 19-20, L932; 6 I , Kenskoff,near Port-an-Prince,Haiti,
altitude 4,000-6,000feet, September2, 1934 (Darlington) ; La Visite
ancl vicinity, La Selle Range, Haiti, altitude 5,000-7,000feet, September 76-23, 1934 (Bates); Loma Rucilla, Dominiean Republic,
altitude 5,000-10,000
feet, June 8, 1938 (Darlington).
I{ephrotomaglabricristatais closestto /[. f erruginea (Fabricius)ancl allies, especiallyto the southernNearctictrf. ferruginea suturalis(I-,oew). It differs especiallyin the coloration,narrow wings, andstructure of the male hypopygium,as abovedescribed. I had.earlier(Journ. Dept. Agr. Puerto Rico, 16: 378; 7932) recordeclthis species
as a variety of f erru,gineab:ut u,ith the presentabundantmaterial itappearsto be quite distinet. In the }lispaniolanmaterial,there isa slight modificationof the praeseulalpattern as abovedescribed.
The margins of the central praescutal stripe ancl the mesal edgeof eachlateral stripe in caseswith anterior euclsontcnlvccltorvards
margin but entirely polished and not differentiated in color fromremainder of stripe.
trIegistoceralongiTtennis(llacqtrart).
Cuba: Santiago de las Vegas, IIay 22, 1931, October 3, L929
(Otero); July 12, 1930 (Bruner).
Brachyprem,n&unicolor Osten Sacken.
Cuba: San Blas, Santa Clara, Trinidad n'Iountains,altitud.e?00
feet, July 26; December23, 1931 (Rowe).
Puerto Rico: one 6 , without further data, ex 'winthem Colleetion in the Vienna Museum.
D olichopeza( Megistomastir)deueroAlexander.
Cuba: North side of the Pieo Turquino, Sierra Maestra,Oriente,
altitude 4,500-6,000feet, June 18-20, 1936 (Darlington).

NEW OB I,ITTI,E KNOWN TIPUI,IDAE

105

Dolichopeza( nlegistomastir)darlingtoni sp.n.
General eoloration dark brown, the praescutum more redd.ishbrown; antennae (male) short, less than one-half the length of body'verticils short; legs black, the femoral basesnarrowly yellow; wings
strongly darkened, the stigma dark brown; abundant macrotrichia
in cells of outer three-fourths of wing; abdominal segmentsweakly
bieolored, yellowish brown, the incisures narrowly blaekened; malehl'popygium with eighth sternite produced into a long median lobe
that terminates in a trident of black spines; median area of ninth
tergite somervhatsimilarly produced into a long lobe bearirig coarse
setae; outer dististyle lvith the outer lobe narrow.
1\[ale.-Length about 10 mm.; lving 9.5 mm.; antennae about
4 mm.
Frontal prolongation of head short, blacli; palpi ltlael<. Antennae blacli, the pedicel obscure ;,-ellorv; flagellum lelatir-el1-sliort,less than one-half the length of body; segnents c;'linclrieal, rvith
verticils that are shorter than the segrnents: pnbescenceof segments
short and inconspicuous. Heacl brorvnish black, tire anterior vertex
paler, relatively narrow, less than trvo times the cliameter of scape.
tr'Iesonotalpraescutum almost uniformly clark recldish brown, withindications of a median and shorter lateral brorvn stripes; humeraland lateral portions of praescutummore blackened; posterior sclerites
of noi irm dark brorvn, the median region of scutum and. scutellum
a little paler; mediotergite ancl pleurotergite more reddish brown.
Pleura brownish black. Ilalteres with stem and apex of knob obscureyellorv, base of knob wealily darkened. Legs with coxae dark brown;
troehanters rrellou'. remainder of legs black, the femoral basesvery-Wingsnarrorvly yellow.
strongly darliened, especially the costal
borcler ancl outer radial field; stigma darker brown; veins ancl
trichia brorvnish black. I\facrotrichia of cells abunclant, including
all but the basal fourth of u'ing and involving the distal half of cell
2nd A. Venation : R1* 2 preserved as a weak spur; petiole ofcell JI1 a little shorter than m; cell 7sf IlI2Iong and narrow, a trifle
widened outwardly but still approximately parallel-sided.
Abdominal segments rveakly bicolored, yellowish brown, the incisures narrowly blackened, involving both the apices and. bases of
the individual segments; hypopygium darkened. MaIe hypopygium(Fig. 11) with the eighth sternite, Bs, produced into an elongate
median lobe that is tipped with three conspicuous black spinous
setae. Ninth tergite, 9t, somewhat similarly produced. into a long
mediau lobe that is a little expanded at distal end and provid,ed with

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TIIE JOURNAL OF AGRIUULTUIiE Or TIIE UNIVERSITY OF P. R.

long coarsesetae; lateral tergal lobes low, with a close group of abbuta dozen setae. Basistyle, b, produced into a eonspieuouslobe thatbears numerous short black spines at apex, with a few longer spines
along mesal faee. Outer dististyle, od more elongate than in det'era,
especially the narrou' outer lobe, the inner mesal lobe slend'er,provided with blaekened setae. Beak of inner dististl']s, fd, small.
Habitat.-Cuba (Oriente).
Holotype, 6 , Pico Turquino, Sierra Maestra, north side, altitude4,500-6,000feet, June 18-20, 1936 (Darlington).
Nambd in honor of the collector, Dr. P. J. Darlington, Jr. The(Ilegistomastir) deuero Alexander, which
nearest atly is Dolich.oTteza
is well-distinguished.by the elongate antennae that are more than
twice as long as in the present fly. The structure of the male hypopygium differs in all details. The somewhat remarkable lobe of
the eighth sternite of the present fly is represented in d'eaeraby asmaller pale unarmed. lobe.
Dolichopeza (nlegistomastir) domingensis sp. n.
General coloration dark gray; antennae relatively short, black,with inconspicuousverticils and pubescence; pleura variegated yel'
low and brorvnish bla.ck; halteres black; legs black, the femoral
bases restricteclly obscure .vellorv; 'lvings strongly infuscated., thestigma darkenecl; certain of the veins seameclrvith brown; obliteratiye areasat corclconspicuous;macrotrichiaof cells abundant,present
on distal three-fourthsof rvingi Rr*2 cntire; cell Ur sessile;abdominal tergites black, the lateral margins obscure yellow; malehypopygium with the inner dististyle unusualll'broad, its apex obtuse;eighth sternite with the median lobe lorv and obtuse, rvithout mod.i'
fied spines or setae.
1\Iale.-I-iength about 9 mm.; rving 10.8 mm.; antennae about
3.6 mm.
Rostrum obscure brownish yellow; palpi black. Antennae relatively short, blaek, the pedieel a little brightened; flagellar segments
subcylindrical, the verticils much shorter than the segments, Pubeseeneeshort. Head dark brown.
Pronotum dark brown. Pretergites and humeral region of praescutum restrictedly pale. Disk of praescutum covered.by confluent
dark gray stripes, obliterating the interspaces and restricting thebrownish black ground. to the lateral borders; scutal lobes dark {taYtslightly variegated by blackish, the median area narrowly pale;scutellum polished blaek, the parascutella pale, slightly marked with

NEW OB LIITLEKNOWN TIPULIDAE

107

darker; mediotergitebrown, more blackenedlaterally. Pleura yellow, variegatedwith brownishblack areas,especiallyon the propleuraand anepisternumand again on the ventral sternopleuriteand meronI
a dark spot on ventral pleurotergite. Halteres black, the base o-tstem restrictedly yellow. Legs with coxaeand trochantersyellowish
testaceous;remaind,erof legs black, the femoral basesrestrictedly
obscure yellow. Wings (Fig. 2) strongly infuscated.,the stigma
a little darker
darker; cells C and ,Sc,together with a seamoL nl,.c'u,,
rvhitish obliterativeareasbeforestigua
than the grountl; conspicuousand especiallyacrossbasesof cells 1st Mz and Ms; veins beyond.
cord and tip of 2ncl ,4 rather insensiblyseamedwith darker; veins
and trichia dark brou'n. n{acrotriehiaof cells abundant, including
virtually the whole u'ing rvith the exceptionof the proximal fourth(representedin figure by stippling) ; in the outer cells where theveins are seamedn'ith cLarker,the tricltia are more restrictedto thecenters of the cells. Venation: R1-l-', entire; Rs short, olllique,subequalto basal section of Ra * 5; eeII //1 }roaclly sessile.
Abdomenwith tergites black, paling to obscurevellorv on sides;
sternites ehiefly yellow, the incisures darkened,more especiallyinvolving the basesof the segments. Male hypopygium (Fig. 12) withlobe of eighth sternite, 8s, very low and broacl,provided only withnormal setae. Caudal margin of ninth tergite, 9t, generally transverse to very feebly emarginate,with abundant coarseblack setae.
Region of basistyle,b, with coarseblack spines that are continued
based on the face of style. Outer dististyle, od, scarcelymodified..
Inner dististyle, id, with the blade very broad, the apex obtuse.
Habitat.-Dominican Republic.
feet, June 1938
Holotype, 3 , Loma Rucilla, altitude 5,000-10,000(Darlington).
DolichoTteza(Megistomastir) dom'ingeruzsis so distinct from theother described speciesof the subgenusthat it requires little comparison. It is the only specieshaving vein R t 12 entire and cell
-I/r sessile. The structure of the male hypopygium is somewhatunusually generalized. The subgenusMegistomastir had not hitherto
been recorded from Hispaniola though represertted.by somewhat
numerousspeciesin Puerto Rico and Cuba.
Limonin ( D'fscobola)gowdeyi (Alexand.er).
Cuba: San Blas, Santa Clara, Trinidad. Mountains, altitude 700
feet, July 1926, December4, 1931 (Rowe). Pico Turquino, Sierra
Ilaestra, Oriente, south side, altitude 3,000-5,000feet, June 1936
( Darlington) .

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Limonia( Limonia)

OF P' R'

luoffmani 'lrLexander'

Puerto Rico: El Semille, Villalba, altitude 1,600 feet, at light,
Janirary 26, L935 (W. A. Hoffman).
Li.m,onia(Limonia) umbrato' (de Meijere)'
Dicranomg,[n umbrata de Meijere, Tijd. voor Eut., 54; 25-26;
1911.
Limonia fissitisAlexander, Ann. Ent. soc. America, 19: 159 ; t926'
widely distributed in eastern Asia. Apparently introdueecl byeommerceinto the Neotropiear Region. The type-Ioeality of. fissilis
is Vera Cruz, lVlexico.
Cuba: Sierra Rangel, Pinar del Rio, altitude 1,500 feet, 1936
(Brother Chrysogone,Colegio de la SaIIe,Havana) 'Limonia ( Litnonia) domballah sp. n.
General coloration of mesonotum black, the praescutum more
obscure yellow and castaneous; pleura black; knobs of halteresyellorv; femora yellorv, narrowly ringed with black before tips;
wings yellow, patternecl with brown; Rs angllated and spurred' atorigin; abdominal segments black, the eaudal borclers narrowly
yellow.
Female.-I-.rength about 10 mm. ; wing 10 mm'
Rostrum and palpi blaek. Antennae black throughout; flagellar
segments oval, with very short apical neclis; longest verticils a
Iittle exeeeding the segments. Ifeacl rvith the front and' narrowanterior vertex gray; remaincler of vertex ancl the occiput black.
the orbits weakly dusted with gray.
Pronotlm blaek meclially above,obscureyellow on sides. Mesonotal praescutlm polished.obscureyellorv, more castaneousbehind, withthree black stripes, the meclian one intense only on eephalic portion,
becoming obsoleteat or before micllength of sclerite; lateral stripes
appearing as major areas on sides of praescutum, crossing the dorsopleural region and becoming eonfluent with the blackened pleura;scutal lobes black, the median area brownish yellow; seutellum ancl
postnotum shiny black. Pleura and. pleurotergite uniformly blaek.
Ifalteres with base of stem and the knob obscure yellow, the distal
portion of stem infuscated. I-iegs with the coxae yellow, the fore
coxae a trifle darkened on anterior faee; trochanters yellow; femora
yellow, with a narrow black subterminal ring that is a little wid'er
than the yellorv apex; tibiae yellowish brown, the tips narrowly

NEW OR I,ITTI,E

KNOWN

TIPI'LIDAE

109

darliened; remainder of legs black. Wings (FiS. 3) yellow, witha conspicuousbrorvn pattern, as follows: Extensive washesin basal
cells from ,Sc to margin in cell 2nd. A, at origin of Rs, eord andouter end of cell 1st ilIzi stigma oval, darker brown; wing-tip
rveaklyinfumecl; veins brown. Venation: ,Sc1ending about opposite
four-fifths the length of Rs, Scz aI its tip; a short spur of .Rr * z;
Es angulated and spurred. at origin; free tip of Scz and Rz in transverse alignment; inner end of cell R3 lying far proximad. of either
eeII R5 or 7st tr12; cell lst M2 subequalin length to vein Jll3beyond
it; m-c,u a short distance before fork of. M; ceLI2nil /. wide.
Abdominal tergites black, the caudal borders of the segments
narrowly ringed with yellow, on. the subterminal segmentsuniformly
blackened.; basal sternites obscure yellow, the remaining segments
patterned as on the tergites. Ovipositor l'ith the cerci relatively
small and slender, gently upcurvecl to the acute tips.
Habitat.-Dominican Republic.
Holotype, 6 , Loma Rucilla and rnouutaius to north, altitude
5,000-8,000feet, June 1938 (Darlington).
The name, Dunballnh, is that of the Father of the Gods of the
Vood.oosin Hispaniola. The speciesis very distinct from all described regional species,especiallyin the uiriformly blackened.pleuraancl in the partern of the wings.
Limonia (Dicranomplia) calliergon sp. n.
General coloratiou gray, r'ariegated.lvith black; cephalic half ofmesonotal praescutum uniformly brownish gray, the posterior half
clearergray, rvith four distinct black stripes; knobsof halteresblack;legs black, the femoral basesyellow, broadest on fore legs; tips of
all femora narrorvly rvhitish yellow; wings whitish subhyaline, reticulately patterned u,ith dark brorvnI a complete 'vvhitecrossband
before cord 1 vn-c'tLoblique, before the fork of trI, longer than the
distal sectionof Clur; abdomenyellow; rnale hypopygium with tworostral spines from a single common tubercle.
llale.-Length about 6-6.5 mm.; wing 6.4-7 mm.
Female.-Length about 7-7.5 mm. ; 'wing 7.5-8 mm.
Rostrum and palpi black, the former a little pruinose. Antennae
rvith seapeblack, pruinose; pedicel obscureyellow; basal flagellar
segrnentsbrownish yellow, the outer segments passing into black;flagellar segmentsoval. Head black, variegated with silvery, on theanterior vertex a little more golden pollinose.

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THE JOURNAL OF AGRICULTURE OF THE UNIVERSITY OF P' R'

l\lesonotal praescutum gta1, the cephalic half more uniformly
brownish graY, unstripecl; on posterior half with four clearly defined'blach stripes, the intermediate pair not reaching the suture, thelaterals flsecl with the clark lateral margin, crossing the suture andsuffusing most of the scutal lobes; median area of scutlm and thescltellum broadly light SraL parascutella dark; mediotergite chicfly
blackened.,variegated with gray on sid.es; Iateral borders of pronotumand praescutum blackened. Pleura greY, variegated. with velvety
black, especially on anepisternum, dorsal pteropleurite ancl pleurotergite. Halteres with stem pale yellorv, knob black. Legs l'ith fore
coxaeblackened.,remaining coxae more pruinose ; trochanters brorvn;
femora black, the baseslight yellow, more extensively so ori fore legs
where about the basal third. is included, Iess so on the middle andposterior legs where only about the proximal eighth is involved; tipsof all femora narrowly but very conspicuously whitish yellow, theamount subequal on all legs; remainder of legs black. \Yings withthe ground color rvhitish subhyaline, handsomely patterned rvith darkbrown on cephalic portion and more grayish brown behind; patternmore or less reticulate, with a complete white crossbandbefore corcl;
before the cord with from nine to twelve costal clark rays, cell Sc
uniformly darkened; in cell R at base,before mid-distance to originof Rs and at the origin of the latter with dark lines that reach vein
M behind or virtually so; beyond the cord and stigma a similarmuseoid pattern that sendsabout four or five rays to costal border atstigma and about three or four others to above the termination ofvein R3, these latter areas extensively darhened and fused behind in
the proximal portion of the outer radial field; posterior cells of wing
more sparsely reticulate with grayish broln, ehiefl1-evident as narginal rays in ait the cells; eells adjoining vein Cu slightly suffusedwith brown; prearcular field slightly more 1-ellorv; veins pale, clarker
in the patterned areas. Venation: Rs angulatecl and more or lessspuned at origin; ,Scr extending a short clistancebeyond origin of
Rs to near one-fourth the length, rScgclose to tip; cell 7st Mz relatively large, longer than any of the veins be;'ond it; rn-cu oblique,before the fork of trI,longer than the distal section of Cu1; eell 2nd A
rvide.
Abdomen conspicuously pale yellorv; in cases, the subterminal
segments more weakly infuscated; hypopygium yellow. I\fale hypopygium with the tergite transverse, the eaudal margin gently
emarginate. Dorsal dististyle a gently curved sichle, the long drawn
tip acute. Yentral dististyle fleshy, a little larger than the basistyle;NEW OB LITTLE KNOWN TTPUTJTOAN

lllrostral prolongation stout, on outer margin beyond midlength witha single tubercle that bears two subequal spines,these latter about
trvice as long as the tubercle. Gonapophyseswith mesal-apicallobe
stout, with a coarsely and inegularly serrated lateral flange.
Habitat.-I{aiti.
Holotype, 6 , I-.'aVisite and vicinity, La Selle Range, altitude
5,000-7,000feet, September76-23, 1934 (Bates). Allotopotype, g.
Paratopotypes,10 3 g.
A very distinct species,superficially resembling speciesIike Li.
monia (Dicranonr,yia)reticulata (Alexander) or L. (D.) pampoecila(Alexander), yet entirely distinet. I am placing it in the subgenus
Dicranomyia despite the fact that rscr ends a short distance beyond
the origin of .Es, as in L,imonia, s.s.
Limonia (Dicranomyia) calliergon polygrapha subsp.n.
Male.-Length about 6 mm.; lving 7 mm.
Closeto typieal call,iergon,difr.eringas follorvs:
Scutellum with darkened extensionsfrom the scutal lobes, oneither side of median line. tr'emora more extensively yellorv, withmoderately broad, black, nearly terminal rings; tibiae yellow, thebasesnarrowly black, the tips and all tarsi broken. All legs of theunique type are detachedand thus eannot be safely associatedwith-Wingsany given trochanter.
with the dark reticulate pattern much
more extensive,the costal rays narrow and entire; cells R and,Mwith darkened areas, as in the other cells; white crossband.before
cord narrower and less evident; veins dark, including those in the
ground areas,and therefore much more distinct. Ifale hypopyg:um
with the ventral dististyle smaller than in the typical form, therostral spines arising from a similar conspieuousbasal tubercle but
uneqrralin lengih, the outer about three-fourthsthe length of theinner and closely applied,to it.
Ilabitat.-Dominican Republic.
Irolot.r'pe, 3 , Valle Nuevo, southeastof Constanza,altitude about
7,000feet, August 1938 (Darlington).
.\lthough the coloration of ihe thorax and general pattern of thewiugs is ver;' sirnilar in the trvo forms, the differencesin the colorof the legs ancl rvings, and the nature of the rostral spines of themale h1-pop1'gium,in the present fly, ineline me to believe that itma]' be fouutl to represent a valid speciesrather than a subspecies.

ILZ

THE JOURNAL OF AGRICULTURE OF THE UNIvERSITY

OF P' R'

Litrtonia ( Dicranomyin) reti,cttlato (Alexander) .
san Blas, santa clara, Trinidad l\lountains, August 12,
clba:
Sierra Rangel, Pinar clel Rio, January
Ig32 (Bates & Fairchild).
27-90, 1931 (Acufla & Otero).
timonia(Dicranomyia) indefensa sp' n'General coloration medium brown, the praescutum unpatterned;
yellow;antennae black throughout ; halteres black, the base of stemwith alegs dark bro$,n, the femoral tips not brightenecl; rvings
alongb"ow,. tinge, the oval stigma darker brown; vague blorvn seams
thecord and outer end,of cell 1st M2; abdominal tergites dalk brolvn,
thesternites weakly bicolored, reddish brown, the outer half ofsegments darker.
Female.-Length about 9 mm.; wing 9'3 mm'
Rostrum ancL palpi black throughout; flagellar segments longoval. Head blaek; anterior vertex about equal in rvidth to thediameter of the scape.
pr.onotum brown. Mesothorax almost uniformly meclium brown,
without markings, the pleura scarcely paler. Halteres blaek, base ofgreenishstenr pale. Legs rvith the coxae pale brorvn; trochanters
lestaceous; remaincler of legs dark brorvn, the femortrl tips notbrightelecl. wings tyith a brorvn tinge, the oval stigma darker brown;
paler l)ro\\,n, poorly clefineclseams or cord and outer end of cell
1sttr12; veins pale brown. Venation: ,Scrending just llel-oncloriginof lls, Sc, shortly before this point; free till of 5c, atrcl R: intransverse alignment; cell 1st M2 subequal in lengtll to the lonliest
veins beyond it; ttl-curclose to fork of III'-.\bclorninaltergites clark brottn; sternites l'eakll' bicolored, redclisl brorvn basally, the outer half of each segtneut some'whatdarker.
ovipositor with cerci slenrler, verv gelltl1' upcttlYecl.
Habitat.-Haiti.
Ilolotype, g , T-,aVisite, L,a" Selle Range. altitude 5,000-7,000
feet, September 1G-23, 1934 (Bates).
is cpritetlistinct from the other(Dicrrntonryta,) indeltnsr.r,
L1,mottiu,
regional speciesof the subgenus. It is apparently closestto L. (D')(Osten Sacken) ancl allies, cliffering in the coloration ofhu,,m,iclicola
the bocly ancl rvings, and in the uniformly darhened femora.
Limonia (Dicranom,yia) diuisa Alexander.
Haiti z T-taVisiteand vicin ity,I-ta Selle Range, altitud'e 5,000-7,000
feet, September 1&-23, 1934 (Bates).

NEW OR LITTLE KNOWNTIPULIDAS

113

Puerto Rico: El Semille, Villalba, altitude 1,600 feet, at light
January 26, 7935 (W. A. Hoffman).
L'imon,in ( N eolimnobin) il,iua ( Schiner), var.
Cuba: Sierra Rangel, Pinar del Rio, January 27-30,1931 (Acufla& Otero).
Limon,i,o,( RlLipidin) schwarzi (Alexander).
Cuba: Sierra Rangel, Pinar del Rio, January 27-30, 1931 (Aeufla& Otero).
Jamaica: Balaclava (A. E. Wight).
I-imonin (Rhipidin) d,omestica(Osten Sacken).
Cuba: San Jos6,July 7 (at light). Santiagode las Vegas,April7, 1930 (Otero). San Blas, Santa Clara, Trinidad lfts., altitude 200
feet, January 1, 7932 (Rowe). Sierra Rangel, Pinar del Rio, January 27-30, 1931 (Aeufla & Otero).
Puerto Rico: El Semille, villalba, altitude 1,G00feet, at light,
January 26, 1935 (Ifoffman).
Jamaica: Balaclo,ys.;Moneague,tr'ebruary 2-B (w. s. Brooks).
'Wight).
Port Antonio, (A. E.
Limonia ( Geranomyia) antillarum .Llexand.er.
cuba: Buenos Aires, Santa Clara, Trinidad Mountains, May 14,7932 (Aeufia).
Puerto Rieo: El semille, villalba, altitude 1,600 feet, at light,
January 26, 1935 (Hoffman).
Limonin (Geranomaia) mgers,innoAlexander.
Cuba: San Blas, Santa Clara, Trinidad l\{ountains,altitude ?00'
feet, Deeember4, 1931 (Rowe). Sierra Rangel, pinar del Rio, Januarl' 27-30, 1931 (Aeufia & Otero).
Puerto Rieo. El Semille, Villalba, altitude 1,600 feet, at lighf
January 26. 1935 (Hoffman).

Limon,ia( Geranomgia)tibintis (Loew).
Cuba: San BIas, Santa Clara, July 20, 1932 (Bates& F'airehild)..
Sierra Rangel. Pinar del Rfo, January 27-30,1931(Aeufra& Otero).
Puerto Rieo: El Sernille,Viltalba, altitude 1,600feet, at light,
January 26. I935 (IIoffman).
Jamaiea: Balaclava (4. E. 'Wight). Port Antonio (A. E,
Wight).

114

THE JOURNAL Ox' AGRICULTURE On' THE UNMRSITY

OF P' R'

In the Port Antonio specimenthe praescutal stripes are obsoleteisor nearly so, but the iclentity seemscertain. whether the speciesinthe same as the earlier L. (G.) intermed,ia (Walker) remains
question yet is entirely possibleLimonia ( GeranomAi'a)banksiana sp' n.
Alliecl to canaclensis;general coloration of praescutummedium
paler
brorvn, lvithout stripes; scutellumbrolvn, r'vitha vague central
area; mediotergite dark brown, the apical portion yellol'; legns
brorvn; rvings subhyaline,ullmarked' except for the oval, pale brorvn
greatly
stigma; scr ending about opposite midlength of Bs; r-nl
redueedby the approximationof veins R+* 5 and' tr[1* 2; male hypopygium with G lobes of ninth tergite roundecl,provided withconspicuoussetae,the extensivemeclianarea glabrous; ventral dististyle large and fleshy; rostral prolongation short, rvith trvo subequal spines arising from very unequal tubercles, the shortest one
close to apex of prolongation; mesal-apicallobe of gonapoilysisentirely pale, simple, nearly straight.
i\Iale.-Length, excluding rostrum, about 6 mm. ; rving 6.4 mm. ;rostrum approximatelY 2.6 mm.
Rostrum relatively long, brown, the tips of paraglossaedarker
brorvn. Antennae clark brown, the scapea little paler brorvn. Ilead
grayish brorvn; anterior vertex narrow' subequal to diameter ofscape.
i\lesonotal praescltum medium brown, without stripes; humeral
region a little more yellolvish; scutal lobes pale bros'n, the meclian
ur.o pale yellow; scutellum brown, rvith a vague cetrtral paler aveai
mecliotergiteclark bro.vvn,wirh less than tiie apical half yellow.
Pleura br.orvnishyellorv, unmarliecl. I-Ialteres pale. Legs rvith theeoxaealcl trochantersobscureyellorv,the fole coxaea trifle darker;
remaincler of legs pale brorvn, the terminal talsal segmcnts darh
brorvn. \vings, subhyaline,the oval stigma pale blown; vcins blown.
Nnmer,ousmacrotrichia on veins be;'ond eot'd, as rvell as a cornplete
scrics on /ls. Venation: Sc relartivelylotrg, Scr erlding about opposite micllengthof .[is,,Sc2a short clistaneclrom its tip 1 r-?11greatly
rcclneetlby the approximation of veins llo * 5 and Mt * z; n1'-c1'L
closeto fgr.k of.trI; ccll 2nd A rclatively narrow.
liasal abclgrnilal segrnentsblol'tr, the outer segmentssomewhat
more bicolorecl,clarker brorvn at apex than at base; hypopygiurnclark. nlale hypopygiunr (FiS. 14) rvith the outer angles of ninth

NEW OB LITTLE

KNOWN

TIPI'LIDAE

115

tergite, 9f, produced.into rounded lobes that are provided with couspicuous setae, the extensive intermediate area glabrous. Ventromesallobe of basistylelarge. Ventral dististyle,ud,,largeand fleshy,much more extensivethan the basistyle; rostral prolongation short,with trvo spines arising from unequal tubercles; outer spine placecl
very close to apex of prolongation on a low tubercle; second.spine
subequal in length, arising a little farther basad and from a much
longer tubercle that exceedsone-third.the length of spine. l\[esalapical lobe of gonapophysis,g, entirely pale, simple, nearly straight.
Habitat.-Cuba.
Holotype, 6, Soledad,near Cienfuegos,August 6, 1920 (Banks).
I take very great pleasure in naming this fly in honor of thecollector, Dr. Nathan Banks, to whom I am greatly inclebted formany appreciatedfavors. Although generally similar in its generalappearanceto Limonia (Geranomgia) canaclensis(Westrvood),thepresent fly is very distinct in the structure of the male hypopygium.
Alexander.
Limonia ( Geranomyia) su.buirescens
Cuba: I:oma de la Sierra, Santiago de las Vegas, March 5, 1931
(Acufla & Otero).'Limonia ( GeranornVin) subvirescens jamaicae subsp.n.
1\[ale.-I-.rength,excluding rostrum, about 5 mm. ; wing 5 mm.;rostrum about 3 mm.
Female.-Length, exclud.ingrostrum, about 6 mm.; wing 5.8 mm. ;rostrum about 3 mm.
Rostrum pale yellon'ishbrown, the tips of the paraglossaestrongly
recurved. Antennae dark brorvn throughout; flagellar segmentsoval.
Ilead clark brou'n.
Thorax uniform orange yellorv, unmarl<ed. Ifalteres pale. I-regs
yellorv throughout. Wings pale yellon', the subcircular stigma pale
brorvn; veins brownish yellorv. Venation: Sc moderatelylong, Sct
encling about opposite trvo-thirclsthe length of lls, ,Scr at its tip;
Bs abor.rttrviee the basal seetionof Ra * ri In-cul,at fork of-trI; cell
2nd I relatively narrow.
Abclomen yellorv. IIale hypopy'gium mueh as in typical srtbAlexancler (Cuba) but apex of rostral prolongation ofv*irescctts
ventral clististylemuch more slencler,the cliameterat this point only
llte
about one-third the length of the rostral spines. In subuirescens,transverse diameter be.vonclthe rostral spines is more than one-halfthe length of the spines.

116

THE JOURNAL OF AGRICULTURE Ox' THE UNMR'SITY

OX' P' R'

Habitat.-Jamaica.
Holotype, 9 , Port Antonio (A. E. Wight). Allotopotype' I , hpoor condition.
the wings are subhyaline,with the brown
In typieal subui,rescens,veins much more distinct.
Limon'in ( Geranowain) uirescens (I'roew)'
Puerto Rieo: EI Semille, Villalba, altitude 1,600 feet, at light,
January 26, 1935 (Hoffman).
(Osten Sacken)'
Orimarga ( Diotrephn) m'i,rob'tl''t's
cuba: santo Tom6s,Province of zapata, May 5-9, 1927 (Aeufla& Bruner).
Hetius (Hetius) atbitarsis (Osten Sacken).
Cuba: Sierra Rangel, Pinar del Rio, altitude 1,500 feet, April
26, 1933 (Bruner & Otero); January 27-30,1931(Aeufra& Otero).
Eph,iphrogrna( Epi'ph"regma)cubensisAlexandercuba: sierra Rangel,Pinar del Rio, January 27-30, 1931 (Acufia& Otero).
Epiphragma (Epiphragma) auricosta sp. n.
General coloration brown and yellow I antennae (male) short,blaek; first flagellar segmentyellow; femora yellon', with three blackrings; tibiae chiefly dark brown to black, the tarsi paling to obseure
yelow; wings yellow, the eephalic third more saturated, butter-yellow, with an ocelliform pattern.
Male.-Length about 9-10 mm.; wing 11-12 mm.; antennae
about 2.3-2.4mm.
Female.-I-:ength about 12-13 mm.; wing 11.5-12 mm.
Rostrum dark brown; palpi black. Antennae short, blaek, thebasal segmentof flagellum yellow; flagellar verticils exceedingthe
segmentsin length. Ilead ehestnutbrown, with a d.ark brown transverse band aerossanterior vertex and a less distinct median darkening on posterior vertex; posterior portions of vertex more or lesswhite pruinose.
Pronotum clark brown, more yellowish on sides. Mesonotalpraescutum with the ground color reddish brown, with three eonspicuous,
dark brown stripes on cephalic half, on posterior half beeomingobscured and confluent, more uniformly dark brown; humeral regions

I

F

NEW OB I,ITTI,E KNOWNTIPULIDAE

fi7

of praescutum restrictedly pale, more extensively bordered.internallyby dark brown, the lateral margins behind the pseudosuturalfoveae
broadly dark brown; in still other specimens,the color of the praeseutum varies from uniform dark brown to striped with dark throughout the length; scutum chiefly brownish yellow, eontrasting abruptly
with the praescutum; scutellum and. postnotum dark colored, lightgray pruinose, the mediotergite and pleurotergite with their posteriorborders broadly dark brown. Pleura with the propleura and pleurotergite chiefly dark brown, the mesopleura almost uniformly pale.
Ifalteres elongate,stem obscure yellow, knob brown. Legs with theeoxae obscure brownish yellow; trochanters yellow; femora yellow'
with three black rings, the last terminal, the meclian one about twice
as wide as the yellow ring on either side; basal darli ring narrowand somewhatpaler; tibiae dark brorvn to black, in easesthe extreme
tips reddish; basitarsi clark bro'wu, remainder of tarsi paling to
obscurebrorvnish yellorv. \\-ings l-ith the grouncl color ;'ellorv, theeephalic third more saturated butter-yellol'; a conspicuous.chiefly
ocelliform, clark pattern, including oeelli be;'ond areulus, origin ofBs, cord., outer end of cell 7st Mz and fork of M; in region of
stigma. several small, more blaekish areas; a solidly darkenedmark
in cells Cu, 1st ^4"and 2nd A, in alignment with the ocellate area atorigin of Rs; areasin cell 2nd A about three in number, the inter'mecliateones in casesinterconnected; veins brown, more yellowish
not more than its own
in tlre costal interspaces. Yenation : ft7-Cu,clength beyond.the fork of M.
Abdominal tergites dark brou'n, the caudal margins broadly moregrayish; sternites and hypopygium more yellow to brownish yellow.
Ilale hypopygium with the lobes of the tergite obtusely round'ed,
eeparatedby a deep linear notch. Interbases with short triangularapical points.
Habitat.-Dominican Republic.
Holotype, 6 , Valle Nuevo, southeastof Constanza,altitude about?,000 feet, August 1938 (Darlington). Allotopotype, I . Paratopot54rc, 1 I ; paratypes, 1 sex?, Foothills of the Cordillera Central,south of Santiago,June 1938; 1 6 , Loma Rucilla, altitude 8,00010,000feet, June 1938 (Darlington).
Epi,phragma (Epi,phragma) auricosta is very different from thetlrree other speciesknown from the Greater Antilles, especiallyin thertrongll' yellorv wings and the pattern of the legs, as the three-bandecl
femora and darkeil.ed.tibiae.

118

rHE JOURNALoT'AGRICULTUREoFTHEUNIVERSITYox,P.R.
,( Epiphragma) buscki'Alexander'
ETtip'hragma
IIt. La Hotte, altitudeabout4,000feet, octoH a i t i : Desbarriere,ber 12-14, 1934 (Darlington).
Epiphragma (Eytiphragma,)inornatipes sp' n.
General coloration of pr.aescutumbrorvnish yellorv, the cephalicthe rernaitlclerrvith narrow brorvn stripes; pleura
folrtlt darl<enecl,gr.a) pruinose, the clorsopleuralregion and membraue dalli brorvu;
feutora alnost urriforrnll,'bron'u, the tips narrorvly pa1e,tibiae andtarsi yellorv; wings rvhitish subhyaline,patterned with brorvn, muchas in sol.utt.ir; darli areas in outer radial ficld large ancl full, clark
areas in cell llcl .,1 very extensive,involving the central portion ofcell; nale h;.popygiurntlith the inner dististyle relatively narrow'u'ith conspicuoussetae.
]Iale.-Lengtlt about 11.5 n m . ; r v i n g 1 2 . 3 m m .
liostrunt brorvn; palPi black. Antennae rvith scape ancl ped'icel
6laeli, flagelllm lrrolien. I-Icaclchiefly brorvn,the posterior orbits moregrayish; anterior vertex behind antennaemore blackened'
Plonoturn liglit ltrgn n, narrolvly,-blaclieneclmedialll', more broadly
l'ith cephalicfourth almost uniso or1sicles. Ilesouotal przrescututn
forynly clarkenecl,the remaincler nrol'e brorvnish yellolv rvith four
nar.tow bro.lvnstripes that do not reach the suture behincl,the lateralpair much reduced; lateral borders of praescutum Jlehitrr-lpseudos11tgral fovea more lniformly d.arkened,the suturc llorclcred bybrorvl; scutlm rvith cephalic haif buff, the posterior half almost
covereclhy a trausr-elsebroivn band; scutellum buffy; mediotergite
pale, the poster.iorthircl ancl a nalro\v transverseline at near midiength darher brorvn. Pleura ehiefly gray prttinose,variegated withbrolynish black, especialll'oll clorsalpleulites ancl the dorsopleuralregion. Halieres rvith stern yellorv,knob more or less darkened,theapex narro.lvlypale. Legs rvith the coxaegray, the extremebaseof
fore pair d,arkenecl;trochanters yellow; femora almost uniform
brown, paler basally,the apex narrorvly pale; tibiae and tarsi yellow.
Wings rv|itish subhyaline, heavily patterned.with brown, the areasof cephalicancl apical portions paler brorvn with narrow d.ark bord€rSI in Anal field, the areasuniformly dark brown; stigma narrow'almost uniformly clarl<brorvn; wing pattern somelvhatas in solatrir
yet cliffering in eertain important points; dark areasin cell C more
extensivethan the interspaces; marginal areas in outer radial fieldlarge a1d fuII; clark areas in cell ,2nclA very extensive,involving
the ceniral portion of cell.

NEW OR LITTLE KNOWN TIPULIDAE

It9Abdominal segmentsvariegateddark and paler brorm, the lateralbordersbuffy. Ilale hypopygium rvith the outer dististyle relatively
narrow, the tip deeurved; inner dististyle naruowerthan in solatrin,
provided with eonspicuoussetae. Interbases rvith apical spines reeurved.
IIabitat.-Cuba.
Ilolotype, 6 , Pico Turquino, Sierra }laestra, Oriente,north side,
altitude 4,500-6,000feet, June 18-20, 1936 (Darlington).
Epiplwagma (Epiphragma) inornatiTtesis apparently elosestto
the srnaller E. (8.) solatrir (Osten Sacken), having the pattern of
the femoravirtually'the same. It differs in the eolorationof the lri;,lX
and rvings,and in the details of structure of the male lrypopygium.

Shannonomyiabatesi sp.n.
Belongsto the mcsophragmegroul); rlresonotLim
glaJ'; halteres
pale yellos'; fenrorayeliou-u'ith a lrrou-uishl-rlaclisubtclnrirralring,on fore and midclle femora rvith tire l-rasalthircl ot' morc sinrriuli.v
blacliened; s'ings yellorv,spottecland clottecln'ith ltrorvn; cell 7st Mz
closed,elongate,rvith m,-clLat near one-thircltlie length.
1\Iale.-Length about 5.5-6 mm. ; rving 6-6.5 mm.
Female.-Length about 7.5-8 mm. ; rving 6.5-7 mm.
Rostrum and palpi black. Antennae yellow, the outer flarleliar
segmentsinfuseated; in darker individuals the antennae are rhorc
uniformly brorvn throughout; flagellar segmentsoval. Head gray.
nlesonotum grsX, the praescutum rvith a slightly darker graymedian stripe, the scutal lobes similarly darkened. Pleura black,gray pruinose. Halteres pale yellorv throughout. Legs with thecoxae black, pruinose; trochanters yellow; femora yellorv, rvith aconstrricuous,
dark brou'n to brol'nish black ring before the subequal
or narro\\'er yellon' apex; fore ancl middle femora rvith the basal
third or more similarly blaekened,leaving only a relativcly narlorv,
obscureyellou' ring betrveenthe two dark bands; posterior femora
less distin'ctly or not at all darkened at base; tibiae and tarsi yellow,
the tips of the latter darkened. The females seem to have clar.ker
legs than the males and at least one specimenhas the fore and middle
femora uniformly blaekened'rvith the exception of the narrow yellow
bases; hind femora less evidently blackeneclexcept at tips; in alleasesthe fore and midclle femora are more extensively blackened
than are the posterior femora. \\rings (Fig. 4) v'ith the srouncl
color yellorv, the prearcular and costal fielclsa little deeper yellow
to bron'nish yellorv; a heavl' spotterl aucl dotteclbrorvn pattern, the120

ox' THE uNIvERSrry ox'P. R.
TrrE JOURNALon AGRTOULTURE

largest areas at stigma, more or less eonfluent rvith a seam alonq
cord; smaller brorvn spots at origin of Rs, supernumerary erossveiu
in eell n, nt-ut ancl outer ends of veins Rs and R+i small brownat enclsof longituclinal veins, usually more extensive
marginal spcl-rs
at 2ncl A; numerousto somewhatsparsebrown dots in most cells of
wing; veins yellon', darl<er in the cloudeclareas. Venation: ,Scr
ending at near ts'o-thirds the length of Rs, rSc2near its tip; a supernumerary crossyeinin cell R connectingrvith Rs about its or,vnlength
beyond origin 1 r-nl strongly bowed; eell 1st 1I, elosec'I,elonsate,
longer than any of the veins beyonclit; nx-cllat near one-third thelength ; basal section of.Ms variable in length, from short ancl nearlystraight to areuateclor even feebly angulated.
Abclomenblaelr, sparsely pruinose; hypopygium reddish brown.
I-Iabitat.-IIaiti.
I{olotl'pe, d . La \risite ancl vieinity, T-.taSelle Range, altitude
5.000-7.000
feet. September16-23, 1934 (Bates). Allotopotype, I .
P a r a t o p o t y p e s , 96 g .
f take great pleasurein naming this very distinet fly in honor of
the collector.llr. ]Iarston Bates. The speciesancl the two next
in Hiscleseribedare the fir'st speeiesof the genus to be cliseoverecl
paniola. Bp- my kev to the Crrbanrnernbersof the genus (Journ.
Agrj lTniv. Pnerto Rieo, 21,:l>25-rt27;1937). the 1l'esentfl1' l'un.s
phragntophoraAlexander. It cliffers from all
to Shnnnonontyia.
thi'ee Cnl,an inenrlrersof the g]"oupin the numerousdots in the cellsof thelvings ancl jn the hear'.n:clark pattern of the femora.
Shannonomyia haitensis sp. n.
General coloration gl'a)-. 1he lrlaeseutum rvith vaguely darker
stripes; basal three antennal segurents),ellol', tlte remainclerblack;heacl ancl pronotum clear light gray; legs yellow; wings whitish
subh,valine,eonspicuouslypatterned with brorvn, the areas confi:rncl
to the vieinity of the veins; Rs relatively short, angulatedat origin;
R2 somedistaneebefore fork of ecll l?3; vein Ra strongly upcurved
at tip 1 t'-I't?,
stronglv borvecl; cell 7st ll[z elongate.
1\[a]e.-I-.,engthabout 5-5.5 mm. ; wing 5-6 mm.
Female.-Length about 6.5-7 mm.; wing 5.5-6.5 mm.
Rostrum gray; palpi black. Antennae short; basal three segments yellorv, the remainder of flagellum black; flageller segments
oval, mueh longer than the segments. Head light gray; antcrior.
vertex relatively wide, exceedingtwo times the diameter of scupe.

NEW OB IJTTIJ

KNOWN

TIPI]LIDAE

L?rPronotum clear light gray; lateral pretergites similarly clear3n]'. ]Iesonotal praescutum light gray to brownish gray, with three&rLer gray stripes; scutal lobes similarly darkened.; scutellum, rvith a capillary brown median vitta; mediotergite gray
. Pleura grey, somewhat variegated with darker gray, esially dorsally and on ventral sternopleurite. Iralteres pale yel. Legs with the coxae yellow, fore coxae a trifle d.arker; troters yellow; remainder of legs yellow, the outer tarsal segments'wingsi trifle darker.
(Fig. b) whitish subhyaline, with a conqlieuous pattern that is confinedto the vicinity of the veins, arranged.
es follorvs: Arculus; origin of Rs; fork of Sc,. stigma; cord andouter end of cell 7st IlI2; marginal spots at ends of all longitudinalveins, snallest on R5, largest on 2nil A; in cases,dark cloudsat forkof n2 * s * a and at intervals along distal section of IlIl * 2; veins
yellorv, darker in the clouded areas. Venation: Bs relatively short,engulated and, in cases,short-spnrredat origin; Scr enclingat or
beyond.three-fourthsthe length of .Es,' .Eesom€clistancebefore forkof eell ,Es; vein Ra, and usually R3 also, strongly upcurveclat tip;
J-rn very strongly bou'ed; cell 1st Mz elongate,subequalto or longer
than vein llh + 2 beyond it; In=cvtat from one-thir.dto ncar one-halftihe length of the cell.
^\lrdomendark brown, including the hypopygium.
IIabitat.-Haiti.
Holotype, 6 , La Yisite and vicinity, La selre Range, altitude
5,000-7,000feet, september 16-28, rg}4 (Bates). Allotopotype, 9 .
P a r a t o p o t y p e s , 66 g .
Shannonontyiahaitensisis very different from the describedCuban
and Puerto Rican speciesof the genus. rt is perhaps closestto B.
bretricula Alexander, of Cuba, differing notably in the pattern andvenation of the wings.
Shannonomyia septem-punctatasp. n.
General coloration pale yellow, including the antennal flagellum,
halteresand legs; wings yellorv,rvith sevenconspicuousbrown spots;
Bs unusually long, exceedingvein Ra, angulated.and short-spurred.
rt origin ) Rz shortly before fork of cell Rs; rn-cu shortly before
nidlength of the long eell 7st Mz.
Female.-Length about 6.5 mm. ; wing 6.2 mm.
Rostmm ancl palpi l-rrown. Antennae short; scopebrown; succeeding segmentspale, the outer ones a little darker; flagellar segnmts oval. Head brownish yellow to pale brown, darker anteriorly.

122

TrrE JOURNAITOx' AGRICULTUREOx' TIIE UNIVERSITY OI' P' R'

Thorax uniformly pale yellow, virtually unmarked'; central portion of scutum and adjoining meclian portion of praescutum somewhat clearer yellow; scutellum and.med.iotergitewith a vague darker
yellorv. Legs yellow, the terminal
median vitta. Halteres pale
'Wings(Fig. 6) yellow, the prearculartarsal segments darkened.
and.costal portions a trifle more saturated yellow; seven conspicuous
brorvn spots, as follou's: At arculus; origin of Rsr' stigma; basal
section of Rr; remainder of anterior cord at end. of vein Mr' outerend of eell kt M2; m-c\e; veins yellow, darker in the clouded areas.
Venation: Rs unusually lo1g, exeeedingvein Ra, on$ulated and shortspurred. at origin ; Rz subequal to Rr * z, placed just before forkof ceII Rs; r-nt, arcuated, lying proximad of vein .Er; cell lsf IIz
elongate, subequal to the veins beyoncl it, with nx-cu shortly before
midlength.
Abdomen obscureyellorv, the incisures of the more basal tergites
a little darker. Ovipositor with unusually long and slend'ervalves,especially the cerci.
Ilabitat.-Dominican RePublic.
Holotype, 6, Yalle Nuevo, southeastof Constanza,altitude about
7,000 feet, August 1938 (Darlington).
is quite distinct from all other
Shannonomyiaseptem-pu,nctata,speeieslinown from the Neotropics. The almost uniformly yellow.olo", eonspicuously patternecl rvings and unusually long Rs will
suffice for the recognition of the fly.
Henatoma (Eriocera) bruneri (Alexander).
Cuba: Coast below Pico Turquino, Oriente,. June 26-30, 1936
(Darlington) . Near Santiago, September 4, 1928 (Bruner).
The first specimenlisted, as well as the holotype specimen,showsa crossveinin cell rScbeyond the origin of Rs, this being lacking inthe second.speeimenabove listed and eviclently not entirely constant.
Heratomo (Eriocera) cubensis (Alexander).
The female had not been deseribed.
Female.-Length about 18 mm. ; wing 15 mm. ; antennae about
3 mm.
Characters as in male, differing in the sexual characters. Antennae 9-segmented.Ovipositor with the valvesvery long and.slend.er,
espeeiallythe cerci.
Allotype , I , Buenos Aires, Trinidad Mountains, Cuba, altitude2,500-3,500feet, May 8-t4, 1936 (Darlington).

NEW OB I,IBTT,E KNOWN

TIPI'LIDA3

t23

The type maleshad been taken in the sameloeality at a somewhat lower level (altitude 1,600feet, March 24, t925, J. G. l\Iyers).
Heratama (Eriocera) iul,iana Alexander.
The allotopotype and paratopotype were added to the original
definition (Journ. Agr. Univ. Puerto Rieo, 21-:529-530; 1937) while
the paper was in proof but no further data eould be supplied at thattime. The paratype were taken on the north side of the Pico Turquino, altitude 4,500-6,000feet, June 18-20, 1936; the allotype atthe summit, altitude 6,000 feet, June L6-27, 1936, all secured by Mr.
Darlington. These speeimensare smaller than the type (I\Iale.Length about 12 mm. ; wing L2.fu73.5 mm. F emale.-Irength about
1? mm. ; wing 15 mm.) but otherwise eonform in all essential features. The antennae of the male are 8-segmented.
Ilexatoma ( Eriocera) otnaticornis sp. n.
Mesonotal praeseutum reddish brown with four elearly definecl
dark brown stripes; antennae?-segmented,basal four segmentsand
apical third of last segmentblack, the remainder yellow; posteriorsclerites of notum and the pleura dark brown to brownish black;halteres with brownish blaek knobs; femora yellow, the tips blackened; wings whitish hyaline, heavily patterned with dark brown,
the areas in part oeelliform; margin of wing narrowly darkened;
Bs in alignment with Ra, the basal section of the latter short;Bz* 3 short, only a little more than one-half Rr* r*n and subcqual to R1l 2; rtu-cutat near one-third the length of eell 7st Mz;
basal abdominal segments yellow, darkened outwardly, the distalrguents, ineluding hypopygium, brownish black.
Male.-I-rength about 14 mm. ; wing 11.7 mm.; antennae about
l.4 mm.
Bostrum blaek, sparsely pruinose; palpi black. Antennae 7-segncnted; scape, pedicel, basal two flagellar segments exeept at incigures and distal third of terminal segment blaek, the remainder
ptly yellow; basal two flagellar segmentsmore incrassated than
three outer ones; three outer flagellar segmentslong-eylind.rical,
ual in length. Head gray; vertieal tuberele moderately derdoped, low, on its eentral portion further produeed into a small
hDule.
Pronotum dark brown. Mesonotal praescutum reddish brown,tith four elearly defined but nanow dark brown stripes, the interncdiate pair ending before the suture, the lateral pair a trifle wider,

I24

THE JoURNAL ox' AcRTcULTUREox' THE uNrvERsrTY ox'P. R.

crossing the suture onto the scutal lobes; median region of seutum
paler; scutellum testaeeousbrolvn, the parascutella darker; mediotergite testaceous brorvn, darlier laterally, the pleurotergite more
uniformly darkened. Pleura brownish blaek. Halteres witlt stem
yellorv, knob brownish black. Legs 'with coxae brou'nish blaekened
troehanters yellow; femora yellow, the tips conspicuosly blaekened(about 1 mm. or a little more) ; tibiae yellow, the tips brownish
blaek; basitarsi brorvnish yellou', the tips and remainder of tarsi
blaeliened. Wings (Fig. 7) rvith the ground eolor whitish hyaline,
heavily patternecl rvith clark brown, the pattern with more or lessdistinct ocelli at origin of -Es,cord and outer end,of cell 1st lllz; two
dar'li costal areas at end of Sc ancl at stigma, converging behindand rrniting rvith clarliening at cord; cell C dark brown, alternating
l'ith forir smaller more yellorv spots; a narrorv but conspicuousbrownmargin frorn bey'onclstigma almost to rving-basein eelL2nd A; asemicircular brorvn alea acrossbasesof cells R and III beyond.arculus;
brorvtr rvashesin basal portions of outer radial fielcl ancl in cells M,
Crt, 1st ,l ancl .2ttd,A: veins brol'n. Numerous macrotrichia oll
veins be;'ond eorcl. Yenation: l?s longer than in ,autfiai, in alignment u'ith R; the basal section of latter l,'et'y short to virtually
laeliing; Rz*s short, only a little more than one-half Rr* u*nancl snbequal to Rt * z; eell R3 much shorter than in acuiiai; cell
7st nI2 relatively large, vein l/1 * 2 beyond it altout one-half longer
than the cell; nl-ctt.at near one-thircl the lcngth of cell 7sf IIe, alittle longer than the clistal section of Cut. No supernumerary crossvein in cell Ra, as in acuiiai.
Abclomen elongate; first tergite clalk brorvn; second segment
yellorv, narro'wly darkened laterally, the following segments more
obseure; outer segmeuts passing tirrough dark Jtrorvn to Jn'or','nish
blaek.
Habitat.-Cuba.
Holotype, 3 , Loma del Gato, Sierra del Cobre, Oriente, altitude
abont 3,000feet, July 3-7, 1935 (Darlington).
The most similar species is Heratoma (Eriocera) acuiia,i,Alexander, of western Cuba, rrhieh, rvhile having the wing-pattern somewhat similar, differs in all cletails of bodv coloration and venation.

Hexatoma (Eriocero) multiguttula sp. n.
l\'[esonotumdark broln, the ]ateral borders of praeseutum very
narrou'ly yellorv; heacl gray, the posterior vertex with an extensive
brown area; legs black, the tips of all tibiae and the entire tarsi
paling to brownish yellow; wings saturated, with a conspicuous pat-NEW OB IJTTLE

KNOWN

TIPI'LIDAE

t25

tern of sparse brown spots and abundant dots in all the eells;
R, * s * n shorter than Rz * s; rn-cu far distad., its caudal end
more proximad.than its cephalic end.
Female.-Length about 17 mm.; wing 18 mm.
Rostrum blaek, gray pruinose; palpi black. Antennae (female)
9-segmented; scape and pedieel obscure yellow, sparsely pruinose
above; basal two flagellar segmentsclear light yellow; succeeding
four segmentsmore infuseated, with their apices clearer yellow;
terminal segmentpale basally,the outer half darkened; first flagellar
segmentabout one-halflonger than the seeond;terminal segmentonethird longer than the penultimate. Head.light Sray, the posteriorvertex with an extensivebrown central area; vertieal tubercle low.
Pronotum light yellow, the eolor continued baekward onto theextremelateral border of praescutumand the d.orsopleuralmembrane
to the wing-root. Mesonotalpraescutum,seutum and seutellumuniformly dark brown; mediotergite dark brown, the posterior andlateral bordersa trifle pale, with a delicatepale median vitta. Pleura
obscureyellow, the propleura and mesopleurasomewhatinfumed but
not forming a stripe; ventral pleurites and pleurotergite clearer
yellow. Halteres with stem obscureyellow, knob dark brown. I-regs
with the coxaeand troehanterslight yellow; femora entirely blaek;'Wingstibiae black, the tips paling to brown; tarsi brownish yellow.
saturated yellow, patterned.with brown, includ.ing major areas at
origin of Rs, R2, anterior eord,and tip of vein Cu; all cells with
abundantsmall brown dots; veinsyellow,darker in the major clouded
areas. Venation: ,Sc1ending oppositefork of Rz * s * n; .82nearlf
as long as Er * zi Ez* s * n shorter than Rr* s but longer thanthe basal section of Rr; eell 7st M2 long, m only about one-third aslong as the basal section of. Ms; rn-cu far distad.,nearly twice its
length beyond the fork of M, about one-third longer than the distal
seetionof Cu,r.
Abdominal tergites obscureyellow, more darkened medially, theextremecaudal bordersof the intermediatesegmentsyellow; sternites
obseureyellow. Ovipositorwith long slendereerei.
Habitat.-Dominiean Republie.
Holotype, 9 , Valle Nuevo, southeastof Constanza,altitude about?,000feet, August 1938 (Darlington).
The only generally similar speeiesis Heratanxa(Eriocera) cramptoni (Alexander), known only from the Blue Mountains,Jamaica.
This difrers most evidently in the pattern of the thorax, unbrightened.
tarsi and in the details of venation, especiallyin the medial field,

126

TIIE JOURNAL Ox'AGRICULTURE OX'Tt{E UNI\TERSITY OX'P, R.

where ra is only a little shorter than the basal sectionof Ms and'm'cu
has a different course,with the caudal end lying more distad.thanthe cephalicportion.
Henatoma ( Eriocera) trifasciata (Riider)'Puerto Rico: El Yunque, Lruquillo National Forest, June 7-10t
1935 (Garcia Diaz). I-.,asMesas,near Mayagiiez, over quiet pool insmall stream, altitude 900 feet, November3, 1935 (A. H. I\Iadden)'Teucholabis (Teuch'olabis)mgersi Alexander.
cuba: sierra Rangel,Pinar del Rio, January 27-30,1931 (Acufla& Otero).
T eucholabis( T eucholabis) nigrosignata .Llexander.
Cuba: Buenos Aires, Trinidad. l\fountains, altitude 2,350-2,800
feet, May 3, lg32 (Bruner & otero). Baraeoa,Oriente, April 2130, 1929 (Bruner & Leon Boucl6).
is ( T eucholab'is
T euch'olab) pmt wicana AJ.exander'
Puerto Rieo: el Semille, Villalba, altitude 1,600 feet, at light,
January 26, 1935 (Hoffman).
Teucholabis (Teucholabis) wighti sp. n.
Size small (wing, 6, under 5 mm.) ; generaleolorationyellow, thepraescutum with three major polished black areas, pleura with asilvery longitudinal stripe; posterior tibiae of male with a narrowdilated ring at near three-fourths the length and a similar dilationnear proximal end of posterior basitarsi; wings rvhitish subhyaline,thestigma and a confluentseamof cord.brown; Sc shorti Rz interstitialwith fork of Rs,. branchesof Rs strongly divergent; male hypopygium with apieal spine of basistyle very small and weak; outerdististyle with a slenderspine on mesaledge.
Itale.-I-rength about 4.5 mm.; wing 4.8 mm.
Rostrum and. palpi yellow. Antennae with scape and petlieel
obscurebrownish yellow, flagellum black; flagellar segmentsoval,
decreasing in size outrvardly; segments with short apical necks.
Head yellow.
Pronotum yellow. l\[esonotalpraeseutumwith the ground reddish
brorvn,with three major black areas,involving the cephalicancllateral
portionsof the sclerite,the latter reachingthe lateral margin; scutumwith lolrcsextensivelyblackened,the broad median area obscureyellow; scutellum broad, yellow, parascutella darkened; mediotergite

NEW OR LITTLE KNOWN TIPI'LIDA3

127

black, the central portion of anterior half a little paler. Pleurawith the dorsal sclerites black, including the anepisternum, dorsal
pteropleurite and pleurotergite; a conspicuoussilvery longitudinal
stripe extending from the fore coxaeacrossthe dorsal sternopleurite,
ventral pteropleurite and meral region; ventral sternopleurite anddorsopleural membrane more yellorvish. Halteres with stem darkened, knob light yellorv. I-.iegsrvith coxae and trochanters yellow;
femora yellow, rvith a narrow brorvn subterminal ring, the narrowapex a trifle narrower, the darhened ring subequal in rvidth on all
legs; tibiae obscureyellorv, in male 'with a slightly enlarged brown
ring at near three-fourths the length, this provided with a group of
settre; tarsi ;'ellorv,the outer segmentsmore clarkened,in male rvitha slightly enlarged and darlieneclring near proximal end of basitarsi.
Wings (Fig. 8) whitish subhyaline, the stigma and a narrow confluent seam on cord dark brou'n; veins pale brorvn. Venation: Sc
relatively short, ,Scrending about oppositeone-fifth the lengtlr.of Rs;
Rz interstitial with cord, at end of Rs; anterior braneh of Rs indireet alignment with the stem, the braneheswidely divergent so eellR+ at margin is approximately twice as wide as cell R2; eeIL7st Mz
closed, gently widened outwardly, shorter than vein Mt * 2 beyond
it; m-cu shortly beyond fork of. M; vein Cu2 very short and pale,indicated only at base.
Abdorninal tergites black, the incisures a little more reddish pieeousI basal sternites yellow, transversely ringed at near midlengthwith darli broln ; snbterrninal segments whitened; hypopygium
abruptly black. Ilale hypopygium (Fig. 7) with the apical spineof basistyle,b, \,ery short, weali and slender; dorsal flange long but
only feebly sclerotized. Outer dististyle, 0d, rvith apical half a flah
tened blade, glabrous exeept on outer margin, eonstrieted at its
baseancl at this point on mesal edge produced into a slenderspine.
Crest of inner dististyle, id, high, the apex obtuse,the beali biclentate.
IIabitat.-Jamaica.
Ifolotype, 6, Balaclava (A. E. Wight).
Tcttcltolubis(Teuclrclabis)rvights is named in honor of the eollector, rvho has talien many of the JamaieanTipuliclacnorv presen'etlin
the lluseau of ComparativeZoijlogy. It is most nearly allied to the
Ctrban T. (7.) nigrosignata Alexancler,differing especiallyin the
naruow subterminal femoral rings, the rving pattern and venation,and the stmcture of the male h;-pop1'gium,especiallythe small weakapieal spine of basistyleand the spine on mesal edge of the outerdististyle.

128

THE JOURNAL OX'AGRICULTUREOx' THE UNwER'SITY OX'P' R'

Gonomg,in(Iiliocera) ongustissimaAlexand.er.
Cuba: Soledad,April Lg26 (Bequaert). San Blas, Santa Clara'
Trinidad Mountains,altitude ?00 feet, Deeember4, 1931 (Rowe)'
Gonomy'ia( Lipophleps) bif'li'gera Alexander'
Puerto Rieo: EI Semille,Villalba, altitude 1,600 feet, at lightt
January 26, 1935 (Hofrman).
Alexand'er'
Gonom,g'in(Lipophteps) san'ders'i''Wojay,Ilavana, tr'ebruarY L4, 1931
Cuba: I-ragUnaCastellanos,(Otero). The speeimen,a male, has eell M2 of both wings open by
the atrophy of basal sectionof Mz.
Gonurng'in(Lipophleps) pleural'is ('Williston)'cuba: santiago de las vegas, March 3, 193L (Otero) ; July 12'
1930 (Bruner).
Gonomyia (Lipophteps) monacanthaplatymera subsp.n.
Male.-I-iength about 3.5 mm.; wing 3.8 mm'
Female.-I-rengthabout 5 mm.; wing 4.5 mm'
Differs from the typieal form in the structure of the male hypopygium (Fig. 1s). outer spine of outer dististyle, od, muchstouter than in either rnonacanthinor helophilaAlexander,the apex
blackened and microscopicallyroughened by appressed'spinulae,smooth in the others; basal spine of outer dististyle a strongly
blackenedflange that extendsinto a strong spine, the outer marginof flangemore or lessirregularly serrate,the surfaceslightly seabrous.
Inner dististyle,id, simPle.
Ilabitat.-Cuba.
Irolotype, 6 , soledad,near cienfuegos,August 6-20 (Banks).
Allotype, I , San Blas, Santa Clara, August L2, 1932 (Bates &Fairchild).
When more mat'erial beeomesavailable, this fly will probably Lre
aceordedfull specific ranking.
Erioptera (Mesocyphona)costal'i'sAlexander.
Cuba: San Blas, Santa Clara, July Z0-August 12, 1932 (Bates& F airehild).

NEW OB LITIII/D KIIOWN TIPIEIDAE

l2g

Ertoptera (Trimi'cra) pili,pesatwmn'l,a(Osten Sacken).
Bermud.a: Cavendish,January (W. S. Brools).
Ifamilton, Deeember(W. S. Brools).

Swamp nea,rAlexand,er.
T oaorhi,na( T oaorhiia) il,omingens'ds
Cuba: San Blas, Santa Clara, August 20, 1932 (Bates & Fairchild).

EXPTANATION OF FIGURES( Symbols: Male hypopygiumi o, aedeagus; b, basistyle; d, dis"tistyle i g, gonapophysis
i M, inner dististyle; od, outer dististyle;s, sternite; f, tergite i ad, ventral dististyle.)
Fig. l.-Tanypremna ( T anyprenxnn,)hodgei sp. tr., venation.
Fia. 2.-Dolichopeza (Megistomastir)domingensissp. D., venation.
Fig. 3.-Limonia (Limon'in) domballah sp.tr., venation.
Fig. 4.-Shannonomyia batesi sp.tr., venation.
Fig. 5.-shannonomyia haitensis sp. tr., venation.
Fig. 6.-shannonomyia septem-punctatasp. n., venation.
Fig. ?.-Hexatoma ( Eriocera) ornaticornis sp. tr., venation.
Fig. 8.-Teucholabis ( T eucholabis) wighti sP.tr., venation.
Fig. 9.-Rhabdomastix fumipennis sp. tr., venatiou.
Fig. l0.-Tanypremna ( T anypremno) hodgei SP.tr., male hypt pygruni
Fig. ll.-Dolichopeza (IVegi"stomostir)darlingtoni sp. n., male hypopygrum.
Fig. l2.-Dolichopeza (Ilegistomastir) domingensisSp.n., male hypopygrum.
Fig. l3.-Limonia (Limonin) apicata dominicensissubsp. n., malehypopygium.
banhsianasp. n., male ltl popygium.
Fig. l4.-Limonia ( Gera,nomAi,a)Fig. 15.-Limonia ( Gcranomyia) microphaea sp.L., rnale hypopy'' grum.
sylvania sp.n., male hypopygium.
Fig. 16.-timonia ( GeranomAin,)
Fig. l7.-Teucholabis ( T eucholabk)wighti sp. n., male hypopygium.
Fig. l8.-Gonomyia (Lipoph,leps)monacanthaplatymera subsp.n.'male hypopygium.
r30

PLATEIPLATEII

9r.9r#.',:LI

8st2