el5
Papeis Auulsos de Zoologia
P.rp6rs Avur,sos Zoot'. S. Plulq

vou 2L, anr. 8: 87-97

RECORDS AND DESCRIPTIONS OF TIPULIDAE
AMERICA (DIPTERA). II

5.IIr.1968

FROM SOUTH

Cnanr-us P. AlexaNDER
Assrnlcr
New speciesof Tipulidae, based mainly on material collected.
by Mr. L. E. Pefia in Chile, Peru and Ecuador, are described in(dif fustor
genera:torti,lis,andthe following
Paralimnophila(triatrata.
spp. n.), Neophilippiano, (breui,seta, sp. n.), Gnophomgia
sp. n.) , Teucholabi,s @olomelania, rectisp'ina, strdcti'spina, flauofintbria, lati,basalis, spp. n., and biclent.ifera mela,nophalhts, subsp. n.),Crgptolabis $thallostena, sp. y1.), Molophiltts (suberentpfros, sp. n.)
and Toaorhi,na (latamera, sp. n.).

The preceding paper under this title appeared in 1966. At
this time I again am considering materials that were taken by
Seflor Luis E. Pefla in Chile, Ecuaclor and Peru, with a few
further species from other sources that are mentioned under theindividual species. I again am greatly indebted to these friends
for making such materials available and for the privilege of retaining the type-especimens in my collection.
Paralimnophila diffnsiol', SP. l't.
General coloration of the entire body black; antennae black;femora and tibiae brown. tarsi yellorvish brown; wings with therestricted ground cream colored, extensively clouded with very
pale brown; nx-cu about one-fifth its length beyond the fork of M;
ovipositor with cerci very long ancl slender.
Female. Length about 9 mtn; rving 7 mm.
Antennae black throughout; fla'
Rostrum and palpi black.
outerones longer, with Jottg consgellar segments suboval, thefiicuous verticils that exceed the segments. Head black.
Entire thorax black, subniticlous, the pronotum and pleurasparsely and. vaguely pruinose, dorsopleural membrane dusky.
Fialterds pale, knobs weakly darkenecl. Legs with coxae brownish
pruinose; trochanters dark brown; femora and tibiae
Lrlack, sparsely-tarsi -paler
yellowish brown.
Wings with the restrictedbrowir,grouncl cream colored, with extensive very pale brown clouds thatclo not include the costal field, the most extensive' placed at near
one-third the length of cells R and M, in the latter cell confluent
with a comparable band at origin of .Rs, extending from R to theAmherst, Massachusetts,01002,U.S.A.

88

Departamentode Zoologia, S5o Pauloposterior.margin in cell 1st A; a broad bancl at cord, more expandedon anterior cord; outer cells extensively clouded, the distal endsof the radial cells paling to_ the ground color; stigma relatively
small, d_arker.brown; vgin,s -brown, slighily paler in the groundareas. venation: - R,." slightly longer than fi", the tip upiirrved;
cell M' a little shorter than its p-etiole; nl-cu about bne-iitttr jts
length beyond the fork of M.
Abdomen brownish black. ovipositor with cerci very longand slender, nearly straight, tips acute.
Holotype, 9, Contulmo, Palo Botado, Nahuelbtrta, Chile, Felrruary 1, 1953 (Luis E. Pefla)
P-aralimnophi,la diffusior is quite distinct from other regional
members of the genus by the subniticlous body coloration, unvariegated legs, and the very diffusely patternecl wings.
Paralimnollhila

tor.tilis,

sp. lt.

size medium (wing of male to 7.s mm ) ; general coloration
groy, the praescutum patterned with dark brown; knobs of halteres brown; femora brownish yellow, tips parker, preceecleclby avague more obscu_reyellow area; wings nairow, whitened, patlerned with dark 3nd paler brown, the former costal in distri6ution,
reaching vein C; a sparse series of brown spots in cell R ancl iribases of outer radial cells; male hypopygium with gor-rapophyses
elon_gat_e,
pale, twisted at near three-louiths their leigth.
Male. Length about 6.5-7mm; wing 6.s-7.smm; an-i,ennaabout
1.5 mm.
Rostrrrm dark brown, pruinose; palpi black. Antennae brownish black throughout; flagellar segments oval to elongate-oval,
shorter than their verticils. Head light gray, with a central darkbrown vitta.
Pronotum light_-gray. Mesonotal praescutum brownish gray,"dclearer gray laterally, patterned with clark brolvn, includirig
broad central stripe, on either sicle with two bro#n areas,'-theanterior one smaller; pseudosutural foveae large, black; -aposlieriorsclerites of notum brownish gray, each scutal lobe withdarkened center;- a continuous brown meclian vitta extending fromsuture to abdomen. Pleura gray, dorsopleuralregion dalkenecl.
Halteres with stem yellow, knob- brown. Legs wiltr coxae lightbrownish gray; trochanters obscure yellowf femora brownish
yellow. -tips darker brown, preceeded by vague scarcely indicatedmore obscure yellow area; tibiae and tarsi brown, the base andtlp of the former narrowly darkenecl. wings long and narrow.
the groqnd whitened, patterned with dark ancl paler-brown; clarker
areas- chiefly costal in distribution, reaching iosta and including
markings at near one-third R, origin of Rs, sc, ancl, in cases, thE
lip of $cr,' stigma qnd spots at ends of veins R, and Rn baler
prqwn; a sparse series of pale brown spots in cell R and others
in bases of cells Rr ond Rr- and near the outer end of cell 2nd. A:
cells y, cr.t, cw and the outer cells washed with pale brown;veins brown, yellow in the costal interspaces. venation: Sc, long,exceedin-gry-cq, the latter shortly beyoncl fork of M, in cases to
one-third the length of rt-cu.
Abdominal tergites dark brown, basal sternites yellowed, narrowly patterned with brown; outer segments clarker to form aPap6is Avulsos de Zoologia, vol. 21, 1968

subterminal ring; hypopygium brownish yellow. Male hypopygittmwith gonapophyses elongate, pale, narrowed gradually outwardly,at near three-fourths the length the blades twisted, their tips
acute.
Holotype, [i , Caramavida, Nahuelbuta, Arauco, Chile, 1,000
meters, February 5-10,1953 (Luis E. Pena). Paratopotypes,2 6 6.
Paralimnophi,Ia pi,rioni (Alexander) similarly has the darkened
areas of the wing reaching vein C and with sparse brown dots
in cell R but none in the medial cells but differs evidently inthe pattern of the legs.
Neophilippiana breviseta, sp. n.
Size small (wing of female 6 mm); mesonotum light brown;legs yellowish white, the vestiture relatively short; wings faintly
tinged with gray, veins yellow; macrotrichia of veins abundant,
relatively short; vein Sc. very long, only a little shorter than -Rs,'
Rs*ashort, less than twice Rr; ovipositor with cerci slender, curved,
hypovalvae much longer, straight.
Female. Length about 6 mm; wing 6 mm; antenna about
1.3 mm.
Rostrum obscure yellow; palpi brownish black. Antennae
black; pedicel enlarged, flagellar segments cylindrical,_ the verticils
very lohg, exceeding the segments. Head buffy yellow.
Mesonotum almost
Cervical region ancl pronotum yellow.
praescutumslightly
uniformly lighf brown, anterior border ofdarker, scutelluln more testaceous; tuberculate Plts and pseudogu'
tural foveae not evident. Pleura brown, more intense ventrally,
membrane light yellow. Halteres with stem whitened,
dorsopleural-infuscated.
Legs with coxae and trochanters yellow; remainknob
der of legs yellowish white; setae of legs relatively short andinconspicuous, on femora and tibiae the long_estslightly more than
one-half the diameter of the segment, in pilosipes the setae very
long, exceeding twice the diameter of femora and more than three
tim-es that of-the tibiae. Wings faintly tinted with grsY, prearcular and costal fields light yellow; veins darker yellow. Moderately long macrotrichia on all longitudinal veins beyond arculus
excepting ifre naruow base of Cu; Anal veins basad of arculus withtrichia. Venation: Sc long, Sc' ending nearly opposite Er, Sc. far
retracted, about opposite three-fourths Rs, Sc' being only a little
shorter than Rs; R**.n about twice R'.', the latter less than twice
Re,' cell lst M, long-subrectangular, gradually windened outwardly,
m.-curbeyond midlength; vein Cu' conspicuously removed from Ca".
Abdomen with subterminal segments brown, the intermediateones more yellowed; sternites pale yellow. Ovipositor with cerci
very slenddr, gently narrowed. and upcurved to the acute tips;
hypovalvae longer, appearing as powerful straight rods, their tips
subacute.
Holotype, 9, Pichinahuel, Nahuelbuta, Arauco, Chile, 1'600
meters, February 12-20,L953 (Luis E. Pefla).
breuiseta is quite distinct from the two other
Neophi.tippiana-species,
known
the genotype egregi,a (Alexander) and pi,losi,pes(Alexander). In venation iL is more as in the latter species,differing in the short leg vestiture. In egregi,a,Sc' is short and Ez*'

Departanrento de Zoologia, Sio Paulovery long. The species previously described were piaced in a
new genus Philippiana Alexander (1929) that later was found to
be preoccupiedand was re-namedNeaphilippianaAlexander (1964).
Gnophomyia (Gnophomyia) triatrata, sp. n.
General coloration of thoracic dorsum grayish brown, pleura
brorpn above, sternopleurite brorvnish yellow; legs yellowed; wings
yellow, stigma medium brown; male hypopygium with inner dististyle a straight blackened rod, tip broadly obtuse; phallosome
shield-shaped,the posterior margin rvith three separate-blackened
lobes.
Male. Length about 6.5 mm; wing 6.5 mm; anterrna about
1 . 5m m .
Female. Length about 8 lnm; rving 8 mm.
Rostrum obscure yellow; palpi dark brown. Antennae rvithscape brownish yellow, pedicel yellorv; proximal flagellar segments
obscure yellow, the outer ones brown, elongate, verticils relatively
short. Head brownish gray, paler anteriorly.
Pronotum brown, pretergites light yellow. Mesonotal praescutum grayish brown, humeral region yellowed, pseudosutural
foveae black; scutum grayish brown, scutellum obscure yellow;
postnotum darker. Pleura brorvn dorsally to form a broad diffuse
longitudinal stripe, ventral sternopleurite more brownish yellow,
metapleura and dorsopleural region light yellow. Halteres short,dark brown, knobs large. Legs with coxae obscure yellow, theposterior pair clearer yellow trochanters obscureyellow; remainderof legs yellowed, outer tarsal segmentsdark brown. Wings yellowed; stigma conspicuous,long and narrow, medium brown; veins
ligth brown, C, Sc and R, together with the prearcular field, clearer
yellow. Venation: Sc' ending opposite E, the latter close to forkof -Rr*s*.ivein Er arcu&ted before midlength, slightly widening thecell; r-m at fork of .Rs,'n1,-c1.0
subequal to or slightly less than its
own length beyond the fork of. M.
Abdominal tergites yellowish brown, sternites and hypopygium
yellowed. Male hypopygium with median region of tergal borderslightly produced into a broad low lobe, each outer lateral angle
farther producedinto a small obtuse lobe. Outer dististyle slender,
narrowed gradually to the subacute tip, inner margin membranous;
inner style a straight blackened rod, sides subparallel to slightly
divergent outwardly, tip very broadly obtuse, basal lobe of style
very reduced. Phallosome a shield-shaped plate, its posterior
margin with three separateblackenedlobes, the central one slightly
longer, gently arcuated, narrowed to the obtuse apex.
_ Holotype, 6 , RibeirSo Pires, S5o Paulo, Brazil, May 1gb4 (N.
L. H. Kraus). Allotopotype, g, pinned with type. Palatopotype,
6 , pinned with type.
T4e species is similar to various others such as Gnnphom.yia(Gnophomyia)_lata Alexander, G. (G.) tatissi.m.oAlexander,-G. (e.)
subhyalina_Alexander, ancl others, all differing among themselves
chiefly in hypopygial structure.
I an indebted to the collector, Mr. No€l L. H. Krauss, of Ho.
nolulu,_for this material and for many other series of crine flies
from all faunal regions of the rvorld.

Pap6is Avulsos de Zoologia, vol. 21, 1968

91

Teucholabis (Teueholabis) colomelania, sp. n.
General coloration of the entire body polished black; rostrum
elongate; halteres and legs blackened; wings whitened, the apex
Uroadly pale brown, with a narrower band at cord; male hypopygiurir witfr spine of basistyle stout, the outer fourth curved intoa point.
Male. Length about 7-7.5mm; vring 5.2-6.5mm; antenna about
7.2-I.4mm.
Rostrum black, elongate,only a little shorter than the remainder of head, palpi black. Antennae black throughout. Head polished black.
Thorax almost uniformly polished black, the pleura with a\rague ventral more silvery longittidinal stripe, more evident
tretrind; clorsopleuralregion paler. Halteres blackened. !.gt- black,femoral baseJ paler; proximal two-fifths of posterior basitarsi moreclilated. Winei whit6ned, the broad apex and a band at cord palebro'r,vn,the latter darker at the stigrna; a more restricted tr{-erbrown cloud in the basal cells excepting cell .R,'veins brou'n. Venation: Sc long, Sc, ending about opposite two-thirds l?s.
Abclomen black. sternal pocket on segruent five conspicuous,triangular in outline, narrowed anteriorly; central spines microscopic, abundant; lateral setae relatively few, directed mesadJ_o-n
selmbnt six with about 8 or I long slender black setae. Male
hypopygiurn rvith spine of basistyle a stout spine, the outer fourtha iuivecl black uoint; mesal face of style witir yellow setae, theflange relatively high. Outer clististyle with a strong lateral spiqe;
terminal spine-long with the more basal point very small, inner lobe
setae. Aedeagus terminating in a slender spine,with very
the expandecl base r,r'ith paired dorsal and ventral setae.
Holotype, 6 , Qttincemil, Cuzco, Peru, August 1962 (Luis E.
Pefla). Paratopotypes,4 66, October 20-30,1962.
Teuch,olabis (Teu,cholabis) colomelania appears to be quitedistinct from other polished black species of the genus that haveblackened halteres and iegs, particularly in the wing pattern andin details of the male hypopygium, especially the basistyle. T.
(f.) uolentis Alexander, while being generally similar is quitedistinct.
Teucholabis (Teucholabis) rectispina, sp. n.
Characters as in colontelania,differing especially in hypopygial structure. Wings rvith the basal darkened area very restricted,
inclucling a narrow wash near base of ceII Cu hut not involving
cell Cu or the Anals. Male hypopygium with spine of basistylelong and straight, only the apical seventh narrowed into a slender
blackened point, the remainder with strong setae that are moreconcentrated ol1 the concave margin. Blade of inner dististyle
narrow, slightly more widened outwardly.
Holotype, 6, Libertad, 6 km southeast of Tena, Napo-Pastaza,
Eeuador, I\{ay 10-12,1963 (Luis E. Pefla).

92

Departamentode Zoologia, 56o paulo

Teucholabis (Teuchotabis) strictispina, sp. n.
Characters as in colomelania, cliffering in the details of hypop-ygial_structure.- Ma-le hypgpygium witlr spine of basistyle straiffrt
throughout, the basal two-third unusually itout, the breidth acr6ss-probase about one-third the length of styki;- outer third oirecirv
cluced into a long straight point; setab abundant, restricted io'theinne_r_face, the outer margin entirely glabrous.
_ Holotype, I , Quincemil, Cuzco, peru, August 1962 (Luis E.
Pefla).
Teucholabis (Teueholabis) bidentifer.a melanophallus, subsp. 11.
characters gen-erally aq in typical bid,entifero Alexander (Hon9uras), differing in details or body coloratibn and especialiy in
Mesonotal -frae,*cutum uniforrnii polshecl
Itypopygial structure.
black, the pleura .with the meral and metapleurii- i"6io",rGrri
1'ellorr. Male hypgp.ygium-genela_lly- as in bidentifiio,-&*teiing"in
det3ils. -spine of basistyle with base yery small 'to virtu"allylacking, th_e long_.terminal_point shorter ttrair it ih; typicat sudspecies. Outer dististyle darkened, more slender, the i'fuo lateralpoints very small..- Aedeagus blackened throughout, the dilatedbase narrower, with _two 5et_ae;terminal spine" stotiter a"o ploportionately shorter, heavily blackened.
g, Balao chico, Rio Frio, Ecuador, April 26-80,
Ilolotyl",
1g6s
(Luis
E. Pefla)
Teucholabis (Teuehotabis) flavofimbria, sp. n.
size medium - (wing of male 6 mm); general coloration of body
black; rost-rum elong-ate,subequal to remlaindei of- h;;d;- monJ drhalteres obscure yellow; -femora obscure yelloiv, iipr' brownish
black, wings- browlish yellow, the relativet"y""iio*-ip"*
and a
-ancl
band at cord pal.e b-rown, darke-r at stigma
the ant'erior corcl,
sternites
f.t' ending shortly beyorid midlengttr-8i--hs; iua"minii
fiy.q through seven with weak
pockeis;male hypopygium-_se1alwi.th sqine of basistyle.longcr]rvect iirto i-toffi terminal_yellow,s-piqe,the concave margin wittr a fringe of long vetiow iEta"; outerdisti.qtyle with a slender lateral spin6.
Male. Length about 6.8-z mm; wing b.6-6mm; antenna about
7.1-1,.2mm.
Rostrum elongate, subsequal to reinainder of head, black, inclu,li".g the palpi. Antennae bralck; flagellar segments long-oval. Headdull brownish black.
Thorax blackened dorsally, the surface slightty polished, sic_lesof praescutum yellowed, the scutal lobes yellowed pbsterioriv r"aon sides. Pleura brownish black, variegated by paler,-kno6 ventral
sternopleurite black. Halteres with stem blackened.
obscureyellow. L.gr with coxae black; trochanters obscure yellow; femora
broadly obscure yellow basally, the narrower tips biownish black,narrowest on posterior legs where about the outer fifth is inclucied; tibiae light brown, tfo. darkened, posterior tibiae with basal
half more yellowed; tarsi brownish black, the llroximal forth ofposterior basitarsi slightly rtilatecl. \Vings rveakly brownish yellol,

Papdis Avulsos de Zoologia, vol. 21, 1968

with two clarkeneclbancls, the outer one pale brown, including
the distal sixth of wing, not involving the bases of the outermeaiit cells; central band much darker at stigma Sncl over theanterior cord, becomiug much paler behind, especially 1! end ofr,'ein 2nd A; ho darkening in basal cells; _veitls brown. Venation:
SC, ending shortly beyond midlength of Rs,' branches of- Rs gele'
rally parallel on more than the basal three-fourths, R- thence cleflecied caudad, cell R, at tnargin nearly three times as extensive
as cell R'
Abdomen brownish black, the hypopygium more intensely
black. Distinct sternal pockets on segments three through seven,
all with relatively sparse delicate setae, weakest on sternite seven.
Male hypopygium *itfr spine of basistyle yellow, long, gradually
narrowed. outwardly, the outer fourth curved into a slender moreclarkened spin6, the fringe of the concave margin comprised of
unusually long yellotv setae, in the paratype the spine somewhat
shorter and stouter; mesal flange of style long, irregularly roughened, the apical end rounded. Outer dististyle just beyond
midlength with a slender lateral spine; inner style with apicalblade unequally bidentate, basal lobe terminating in a short point,with thres long setae. Aecleagus with pairecl dorsal ventral setae,
the apical point slender.
Holotype, 6 , Quincemil, Crtzco, Peru, August 1962 (Luis E.
Pefla). Paratopotype, 6, October 20-30,1962.
From other blackened members of the genus having banded
wings the present fly is best distinguished by the lack of a darkened basal band on the wings and by hypopygial characters,
including the spine of the basistyle. The abdominal sternal pocketsof the male aie quite different from those of Teucholabis (Teucholabis) colontelania,sp. n., especially that of the fifth segment.
Teucholabis (Teueholabis) latibasalis, sp. n.
General coloration of bocty polished black; legs blackened;wings whitened, with two pale brown bands, the one at cord
narrow, apical band broad, no darkening in basal cells; male hypopygium with spine of basistyle dilated into a broad plate, mesal
flange with outer angle extended into a point; outer dististyle
relatlvely slender, inner margin with an acute spine; aedeagus
l:road, terminating in a slender curved rod.
Male. Length 5.5 mm; wing 4.8 mm.
Head broken. Pronotum black, scutellum and pretergites paler.
Mesonotum polished black, the humeral region of praescutum
restrictedly yellowed. Pleura black, the ventral sclerites with abroad grayish stripe, dorsopleural region restrictedly light yellow.
Halteres blackened, apex of knob very obscure yellow. Legs withcoxae black, midle and posterior pairs more grayish; trochantersobscure yellow, apices darker; femora brownish black, bases narrowly yellowecl; tibiae and tarsi brownish black; posterior basitarsi
with proximal third ctilated. Wings with the restricted gro'"tnd
whitened, with two pale brown bancls, the one at cord narrower,
the broad apical area including the cells beyond the general levelof outer end of cells lst IuI,; no darkening in basal cells; stigmaoval, darker bro'wn: veins tlror,vn. Venation: Sc long, ending
crppositethree-fifth Rs,' vein R, subequal to R.,-,.

94

Departamento de Zoologia, Sio paulo

Abdomen black, the hypopygium more intense. sternal pockets
co?spicuous,-on segmentfive-iomprised of numerous long setae on
either side of median area, segme-ntsix with about a doz"eninwardly directed stronge{
setae on- eithe" rio" i,i -.oil;-apicat;p;;". Matehypopygium withspine of basistyieoistincii"",
very
broad, the inner angle fariher extended into the ur""litiT"."i;gspine,
outwardly this narrowedinto.a long_point,margin witt rongsetae;
qlef.alflange_eTtensive,the outer an[te-"iextenoea"int""" p"irir. outerdististyle relatively"Lir two-thircti lhe tength
Jgtg and sleno.i,
on inner margin withan acute spine; i"n""- style with its outerblade extended into an acute qoiiit, #itrt a smaller- obtuse morebasal knob; basal lobe with-setaes'hort' ,toui setae. aeO"igus withbase yery broad, with paired
on both margrns,apex a slendercurved rod.
Holotype,
6 km southeast- of rena, Napo-pastaza,{. libertad,_'(Luis
Ecuador,_M_uy
1g63
E. penat.
.i.0-12,
Teucholabis (Teucholabis) tr.tteibasatis!s most readily told
trom other sma-ll polished black *..nn".:r_or trrJ-genus having
blackened legs by_ the lack of a basal oart<ened *iil'-dano
andln the sffucture of the hypopygiurn, particularly the Jpine of thebasistyle/6r/.-'
sp. n.
General coloration of thorax dark chestnut brown, pleura
obscure y_ellow,- the clorsal pleurites clarkened to form a stripe,
femora yellow, tips narroryly^brown,-tibiae- and basitarsi wfriien6d.

tips .narrowly brown; wingp broad, taintty- o"rk;ilil. macrotricliia
angulated and spurred. at oiigin;

i;.*i;E:in ceni"i's ofwing cells excepting the prdxirirat o""r; mate rrypopvgr"r" mostwithouter distist_yleprod_ucedlaterad into a bhckeneO spine, irtr"r stylea bro-ad pale spatula with long- setae; phallosot""'- ;iii--aeoeafrii
yery. long ald, slender, gonapophysis a harrow oirtenea spatfrta,
its tip rounded.
Male. Length about 4 mm; wing 4 mm.
Female. T.,engthabout 4 mm; rviirg 5 mm.
Rostrum ti_srr_tbrown, palpi black.
Antennae brown, scapeclarker; pedicel large, oval, basal flagellar segments short withlong verticils. Head dark brown.
Pronotum ?nd pretergites pale yellowish white. Mesonotum
dark chestnut brown, central region of scutum obscure yello.w,
pleura beneath obscure yetiow, - above
scutellum paler"vellow.
dark brown to form a conspicuous stripe on anepisternum ancl
dorsal pteropleu_rite, dorsopleural membiane whitened. Haltereswith stem weakly darkened, base narrowly yettoweo, rcnou white_neq. Legs with fore coxae weakly dari<ened, remaining coxae
and trochanters-yellow; femora yellow, tips narrowly brown, tibiae
and basitarsi wttite, tios narrowly oari<eneil, outer tarsal segments
pale brown.
wings l:roatl, raintly darkened; veins pir" brorvn.
Macrotrichia in centers of most celts lacking in bases
through 2nd, A, most extensive in M and,cu. venation: of cerls n
Rs rong,angulated at origin, with a long spur.
Abdome' light. brown.
uite-trypopygium rvith two undar.kened lobes on lnargin oi basistyle.
outer dististyle clilated out-Pap6is Avulsos de Zoologia, vol. 21, 196895

wardly, before apex with a-n ereCt blackened splne; inner st$-e a
nloaO"paie spatuia, provided lateralty with very long setae. Phal'losome unusually long, aedeagus slender; gonapophysis appearing-Hoiotype,narrow daikened spatula, the tip rounded.
as a
6, Contulmo, Palo -Botado, Nahuelbuta, Arauco,
Chila; Fe6iuiry- z, 19b3 (Lui's E. Pefla). Allo.typ_e,j,-Caramavida,
Arau6o, 1,000riretbrs, Februaqy 5-10,1953-(Luis F. Pef,a),
The most similar species-is CrAptolabis (Cryptolabis) spat-utata Alexander which differs especially in the coloration of thelegi anO in details of the male hipopygiury, particularl{ the basistyle, outer dististyte and gonapophysis (Alexander, 7929: 193-194).
Molophilus (ltfololthilus) subexertrptus, sp. n.
General coloration brorvnish blaek; halteres and legs -yellow;
u'ingi yellowed, axillary. region weakly darkened; male hypopygiufi glnerall as rn pretiosis in the heavily blackened-aedeagus,iii.af dististyte shortj phallosome a flattened glabrous blade thatis expanded outwardlY.
Male. Length aSout 4.5 mm; wing 5.1 mm; antenna about
1.4 mm.
Rostrum and palpi black. Antennae relatively- long, l-ight
nrowniiii Veffow; fiagbttar segmen-ts lengthened, with very longrvhite seta-e. Head dark grayish-brown, llrown.
Mesoscutellum light yellow'
dark
scutumPronotal
notum dirt<- brown to brownish black, humeral and lateral regionsof praescutum and mid-area of suture light brown. Pleura brown,,iit-f.-."-Ooirilty to form a diffuse stripe, sternopleurite extensively
Halter6s yellow. L_egswith cbxae^and trochanters yellow;
ill;i.
iemainder of legs yeflowed, outer ends of femora broadly darker,
iips of tibiae nlrrbwly br'own; outer .tqrpal segments darkened.
ffings yetto*, prearcrilar and costal fields hght yellow, axillary
regiJn in notfi anal cells weakly d-arkened;veins yellowish brown,y;iiow in the brightened fields, darker in the axillary_ region.
;i.;
Venition: R, shortly deyond level of r'nt; petiole- of c-ell Ms about
one-half longer thari tha oblique nL-cLL;vein 2nd A long, ending
about opposite midlength of the petiole of cell M".
Abdomen dark bro*n, basal sternites paler. Male hypopygium
generally as in pretiostts and etemyttus; apex of ventral lobe of
fiasistyle with an obtuse sclerotized blade. Outer dististyle agentli curved blackenecl arm, the concave margin with delicate
setaej basal dististyle relatively short, - expanded outwardly, theouter margin with a dense comb of black setae. Phallosome abroadly de-pressed-flattenedglabrous plate, broadest at near three'fourthi tha length, apex g-ently emarginate. Aedeagus heavily
blackenecl,the lower or concave margin with a series of about 20
nearlytruncate projections.
Iiolotype, 6,'Britamalal, Nahuelbuta, _Arauco, Chile, 1,400-1,600
meters, February 12-20,1.953(Luis E. Pefla).
The other regional members of the subgenu,sha.ving the aede'
agus heavily blackened include Molophilus (Molophilus) eaem.ptus
Alexander and M. (M.) pretiosus Alexander, both dift'ering eviden'
tly from the present fly in hypopygial characters, including espe'
cially the apiCal point of the basistyle, basal dististyle, phallosome

96

Departamento de Zoologia, Sio paulo

and armature of the aedeagus. In the last feature it is most likepretiosus but has the legs differently construted.
lbxorhina(Toxor.hina) latamera, sF. n.

General coloration of thorax fulvous brown; legs light brown;
wings brownish -yellow, prearcular and costal fieldl sriEniiv moreyellowed; male_hypnpygium with spine of basistyte striighi, slen4.r, beak of dististyle long and slender, narrowed. gradually tothe subacute tip; phallosome with gonapophysis very br"io, a-boutthree times as long as wide.
Male. Length, excluding rostrum, about z mm; wing 5.5 mm;
rostrum about 5 mm.
Female. Length, excluding rostrum, about 7.s mm; wing
6.6 mm; rostrum about 6.5 mm.
Rostrum brown, subequal to or a little shorter than the wing.
Antennae dark_ brown, pedicel and flagellar fusion segment enlarged. Head brown.
Pronotum brown.
Mesonotal praescutum almost uniform
fulvous brown, the humeral and lateral borders paler; scutal lobes
fulvous brown, median region of scutum, scutellum and postnotum
more testaceousyellow. Pleura fulvous. I{alteres yellowed, knobs
a trifle darker. Legs with coxae ancl trochanters yellow; remainder of legs light brown. wings brownish yellow, prearcular andc_ostal regions slightly more yellowed; veins brownish yellow.
venation: sc' ending almost opposite origin of Rs, sc, at its tip;celr lst Mz longer than veins beyond it, 1n-cu at the fork of M.
Abdominal tergites light brown, sternites more yellowed, subterminal segments slightly darker. In the female the tergites areslightly bicolored, obscure yellow, posterior borders narrowly pale
brown. Male hypopygium with spine of basistyle straight, slender,
narrowed gradually to an acute point. Dististyle with the bodyhigh, the terminal tubercle conspicuous,beak a long slender bladethat narrows gradually to the subacute tip. Phallosome with
gonapophysesvery broad, their length about three times the diameter;. _aedeagusr,vith arms short, their length about four times
the width across the tips, apices obtuse.
Holotype, 6, Huagra Yacu, Ecuador, altitude 900 meters, March
31, 1941 (William Clarke-Macintyre). Allotopotype, g, with thetype.
Among the regional members of the genus that have an apicar
spine on the basistyle, the present fly is most similar to Toxorhinn(Tororhina) pergrocilis Alexander, of peru, and T. (f.) phoracaena Alexander, of Ecuador, differing especially in the hypopygial
structure, including the dististyle and gonapophyses.
The collector of the present materials, William Clarke-Macintyre, was the one person who made the single greatest contribution to our knowledge of the vastly rich crane fly fauna ofEcuador. For an account of his life a paper by the writer (lgbg)
rnay be consulted.

Pap€is Avulsos de Zoologia, vol. 21, 1968

97

Rornnrncns
Ar,n:<etronn,Cnlruus P.
1929:. Dtptera of Patagonia and Soutlt, Clrdle 7: 19&194, fig. 230.
1953: Notes on the Tipulidae of Ecuador. Reo. Ecttot. Ent. Par.
I(1): 1-9.
1964: New or little-known Tipulidae from Eastern Asia (Diptera).
LfV. Plailnppine Jour. Scd. 93(2r: 2TI-324, I pls.
1966: Records and descriptions of Tipulidae from South America(Diptera). Papdis Auulsos Dep. Zool. S. Paulo 782 221-229.