6 7s
Reprinteclfrom Bur,r,nrrr.roF THE Bnoorr,yx ENrouor,ocICALSocrnrv,
Vol. XLVI, No. 4, pp. 85-91. October,1951.

NEW OR INSUFFICIENTLY-KNOWN CRANE.FLIESFROM THE NEARCTIC REGION (DIPTERA,TIPULIDAE) PART XIV.1
By C. P. ATBxANDER,
Amherst, Mass.
The preceding part under this general title was published in
December,1950 (Bull. Brooklyn Ent. Soc., 45: 15G160). At
this time I arn characterizing a few further speciesof the genus
Tipula, all from California and chiefly from the Hastings Reservation, in the Santa Lucia Mountains, Monterey County, where they
were taken by the Director of the Reservation,Dr. Jean M. Linsdale. I wish to expressmy deepestthanks and appreciationto Dr.
Linsdale for the privilege of studying exceedingly large series of
crane-fliesfrom the Reservation which have provided an exceptionally completelist of the species. Representativesof the species
are contained in the collections of the Frances Simes Hastings
Natural History Nluseum and in rny personal collection. Further
acknowledgmentsare made later in the paper under the discussionof Tipula (Lwratipula) hastingsae.
Tipula (Hesperotipula) linsdalei n. sp.
Male; Length about l2-I3 mm.; wing I4-L5 mm.; antennaabout
3.5 mm.
Fenmle; Length about 13 mm.; wing 13.5-14mm.
Frontal prolongation of head brown ; nasus relatively short ;palpi black. Antennae (male) with basal three segmentsyellow,
rernaining ones black. Head black, gray pruinose.
Pronoturn dark brorvn rnedially above, paling to yellow on thesides. Mesonoturn yellou', the praescutum with rnore reddish
stripes; pleura yellow. Halteres rvith stern brownish yellow, knob
1 Contribution from the Department of Entornology, Universityof Massachusetts.
85

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Bulletin of the Brooklyn EntomologicalSociety Vot.XLVI

darker brown, the base of stem clearer yellow. L.g with thecoxaeand trochantersyellorv; femora yellow, the tips blackened;
tibiaeobscureyellow,the tips lllore narrowly darkened;tarsi passing into brownishblack;clarvs(rnale) toothed. Wings whitish subhyaline,the centersof cellsbeyondcord, as u-ellas cell R, slightll'
more darkened,the veinsthus appearingto be seamedwith the paleground; stigma and a small spot at origin of Rs darker brown;
prearcularand costalfieldsyellowed;veins brown, lnore brownish
yellow in the brightenedportions. Venation: Rs fully three times
the short nrcu.
Abdominal tergites reddish yellow, with three broken black
stripes,the medianone interruptedby the gray posteriorbordersof the segments,the lateral stripes reducedto major spots on thebasal parts of the individual segments;sternitesclearer yellow;
hypopygiumcastaneous.Male hypopygiumrvith the ninth tergite
distinctive,appearingas a flattenedplate on either side of a narrow
mediannotch,the marginsrolled downwardto form a partial cylinder. Horn of the basistyleshort and ttnttsuallywidenedat base,scarcelylonger than the inner dististyle. Inner dististyle with the
beakshort and blunt, not blackened; outer basallobe large,virtuallyon a levelwith the main body of the style from which it is separatedby " srnallnotch; basallobesof style of rnoderatesizeonly. Eighthsternite with the setaeof both the lateral and rnedian lobes very
long and nutllerous.
Habitat; California(Monterey County).
Holotype: 6f, HastingsReservation,June 1, L943 (J. M. Linsdale) ; returnedto Dr. Linsdale. Allotopotype;t, pinnedwith type.
Paratopotypes:7 3?,Muy 6-June l, t943 (Linsdale).
I am very pleasedto name this fly for the collector,Dr. Jean M.
Linsdale,Director of the FrancesSimesHastingsNatural History
Reservation. The most similar describedspeciesis Tipula (H esperotiputa) nti,clteneriAlexander, which differs chiefly in the details of structure of the male hypopygiutn.
Tipula (Hesperotipula) ovalis n. sp.
Male: Length about 14 mm.; wing I4.5 mm.; antennaabout
4.6 mm.
Fetnale; Length about 13 mm.; wing 16 mm.
Frontal prolongation of head obscureyellow; nasus elongate;
palpi with basal three segmentsdark brown, the terminal one
fading to pale brown. Antennae (male) of moderate length, asbasal three segmentsyellow, reindicatedlry the measttrements;

oct',7e57 Bultetin of the Brooklyn Entom,ologicat
society87

mainder of flagellum black; flagellar segmentsonly gently incised;longest verticils subequal to the segments. Head black, graypruinose.
Thoracic dorsum reddish yellow, without distinct pattern, thepleura somervhatclearer yellow. Halteres dark brown, the baseof stem narrowly yellow. Legs with the coxae and trochanters
yellow; fetnora obscureyellow, the tips brownish black; tibiae andtarsi brown; clarvs (rnale) toothed. Wings with a brownish
ground, more or less distinctly streaked with paler, especiallyevident along the veins beyond the cord; prearcular and costal
fields yellorv; stigrnamedium brorvn; veins bro.lvn; sparsemacrotrichia in cell Rr.
Abdomen (male) with the tergitesyellow, rvith a broken medianltror,r-nstripe and conspicuousbrownish black sublateralspots on
segutentstn'o to fir'e; sternites and hypopygiunt lltore uniformly
pale. Itr female,the lateral tergal borders are distinctly infuscated,the median stripe best indicated on the more proximal segments.
Male hypopygium with the upper plate of the ninth tergite producedinto tlvo oval blackenedlobes,rvith srnoothrnargins,separatecl
by a narro\\: notch; lou'er plate terminating in tr,volorv transverse
blackenedledges,separatedat the nridline by a very small notch.
Basistyleproducedinto a long, unllsually slenderhorn that narrows
gradually to the acute tip. Outer dististyle dark-colored,gently
sinuousin outline,provided rvith coarseblack setae. Inner dististyle with the beak slender,blackened; clorsalcrest lorv, the outerrnost lobe large and flattened,provided with abundanterect yellorv
setae,the snraller outer lobe behind the dorsal crest small andseparatedfrom the crest onlv by a small notch. Eighth sternitc
with the setaeof the lateral lobeslong and delicate,very abundant;
rnedian lobe rvith only about a dozen much stouter bristles.
Habitat; California (Monterey County).
Holotype: 6f, Hastings Reservation,along Finch Creek,May 2I,
1943 (J. N{. Linsdale); Alexander Collection. Allotopotype; t,rvith the type; returned to Dr. Linsdale.
The most sirnilar describedspeciesare Tipula (Hesperotipula)
circulari.sAlexander and T. (H.) streptoceraDoane, which differ
in the details of structure of the male hypopygium, particularly of
the ninth tergite, basistyle,inner dististyle and eighth sternite.
Tipula (Lunatipula) hastingsae n. sp.

Male; Length about 15-16 mm.; wing lGLg mrn.; antenna
about4.5-5 mm.

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Bultetin of the Brookl,ynEntomologicalSociety Tot"XLYL

Fernole;Length about 18-22mm. ; wing lLzl mm.
Charactersmostly as in polycanthaAlexander ( SouthernRocky
Mountain Region), differing in detailsof rving pattern and structure of the malehypopygium.
Head light gray. Antennaewith the scapeand pedicelbrown,
flagellumbrownish black to black; basalswellingsof the segments
moderatelydeveloped.
Mesonotalpraescuturnwith all dark stripes confluent,the posterior end of the medianstripe much paler. Wings with the dark
and pale pattern very contrasted,the former being slightly nlore
extensivethan the latter; costalborder uniformly darkened;whiteareasbeforecord very irregular,not forming a narrow transverse
bandat near one-thirdthe rving length,as in polycantlm;palepattern in cellsIl[ and,lst A particularly extensive; beyondthe cord,
the patterndistributionmore as in polycantlmbut rnuchlnore contrasted.
Male hypopygiummore as in polyconthathan in vittatipennis,
especiallyi" the armatureof the eighthsternite. Ninth tergite withthe tergil lobesbroaderand more rounded. Outer dististyleexpandedat outer end, the outer angle obtuse,not produced-as inpotycantha. Inner dististyle narrower and rnore produced,theouter basallobe larger. Ninth sternitewith the horns slenderandappearthe tips pointed. Gonapophyses
relativelyinconspicuous,
ing as ,r.ty po*erful divergentcurved.artns. Eighth sternitewith
about five or six majot n-dified setaeon either side,with a few
more reducedones it the base of the series. Tipula polycantlta
differs especiallyin the detailsof the ninth tergite,both dististyles,
and the eighthsternite.
gonapophyses
Habitot; California.
HolotyPe:3, Mount Diablo,Contra CostaCounty,altitude.2860'Cazier). AUotopotype: pinned
feet, M;t n: Ig40 (Mont A.
_?,
Sanctuary,Birds: 3,San GabrielRiver
with the type. Para'type-Comstock)3?, Del Mar, San
M"y 2I, tg+S (J. A.
; nun-Ierous
Diego County, April 29-May 13, 1945,June.10, 1945,late June
195d (J. A.-Comstock); 3?,San JacintoRiver canyon,above
H.,rl.i, RiversideCountY,ln-R.r.rtchapaital,May !4, 1939 (T' H' G'"tion, Monterey County, June.13,
Aitken) ; 3?, Hastings
1
1938,Aptil'26,^Spti"gs,Vta1i9, M"y 23, 1943 (J, M. Linsdale); 9,
El Dorado County, May 27, 1939 (Mont A' CaShinglezier) .
This very attractive speciesis named for Mrs. Frances Sirnes
Hastings, through whose thoughtfulnessand generositythe Has-

oct.,1e57 Bulletin of th,eBrooklyn EntomologicalSociety

89tings Natural History Reservationhas been establishedand maintained. As indicatedin the abovediagnosis,the closestrelative isTipula (Lunatipula) polycantha Alexander, which has been compared throughout the description.
Tipula (Lunatipula)sancta-lucia n. sp.

Size rnediurn (wing, rnale,about 15 mm.) ; mesonotalpraescutunl yellow, with three black stripes,the whole surfaceobscuredbya sparse gray pruinosity ; legs yellow, the tips of the femora andtibiae narrorvly brownish black; rvings with the ground creamy
white, the centers of virtually all cells slightly infuscated,leaving
broad borders of the ground along the veins; stigma brown ; abdorninaltergiteswith a conspicuousblack median stripe, the usual
lateral stripes representedby isolated dark spots on the sublateral
portions of tergites two to seven; male hypopygiurnwith the tergite deeply notched.medially, eachlobe again with a very deep andnarrow U-shapedemargination,the inner lobulethus formed lying
almost on edge,the flattened face against the midline ; outer dististyle appearing as a glabrous polished scoop,the entire inner orconcaveface with an abundanceof long yellow setae; eighth sternite sheathing, strongly narrowed outwardly, bearing two apicallobes,eachwith a densebrush of relatively short yellow setae.
wing 15.5 mm.; antennaabout
Male: Length about 14 m111.;
4 mm.
Frontal prolongation of head castaneous,slightly darker on dorsum ; nasus long and slender; palpi black, the long terminal segment much paler. Antennae with the basal three segmentslightyellow, the remaining segmentsbrownish black, the basal enlargements of the lnore proxirnal ones still darker ; verticils subequalin
length to the segments. Anterior vertex reddish,posterior vertex
and occiput black, both partly concealedby a gray prttinosity, particularly in front; no vertical tubercle.
Pronotal scutum protuberant and blackenedmedially, the sides
brownishyellow; scutellumand pretergitesobscureyellow. Mesonotal praescutumwith the restricted ground color yellow, the disk
chiefly coveredby three black stripes, the whole surface rendered
opaqueby a sparsegray pruinosity ; lateral and humeral portions of
the ground more or less patterned with reddish; scutum obscureyellow, each lobe with a single major blackenedsparselypruinose
area; scutellum and mediotergitereddish yellow, both with sparse
scattered erect setae; pleurotergite rnore or less darkened, lightbrown. Pleura chiefly obscureyellow. Halteres with stem ob-90Bulletin of the Brooklyn,EntontologicalSociety vot'.xLvI

scure yellow, knob infuscated. Legs tvith the coxae and trochanters yellow; femora and tibiae yellow, the tips narrowly brownish
black,the amountsubequalon all legs; basitarsiobscureyellow, theapex and remainderof tarsi black; clarvs (male) toothed. Wings
relatively broad; ground color creamy rvhite, the centers of virtually all the cells slightly infuscated,leavingbroad bordersof theground along the veins; cell C pale bror,r'n,prearcular field and cell
Sc clearyellow; stigmabrown, relativelyconspicuous,with several
trichia; a small brown spot at origin of Rs,' vein brown, paler in thebrightenedfields. Venation: Rs about three tirnesthe short ut-mt,;
lretiole of cell AI, abotrtone-half longer than m.
Abdomenyellow,rnoreor lesspolishecl; tergitesrn'itha conspicttous blackmedianstripe,on the subterminalsegmentsinterruptectb1'
pale yellorv borders; the usual lateral stripes replacedby isolated
llrou'nishblack spotson strblateralportionsof tergitestrvo to seven,
inclusive; sterniteschiefly obscureyellorv,the posteriorborclersof
the segrnents
vaguelylllore infuscated; hvpopygiumrelativelylarge,
castaneous. Male hypopygiumrvith the tergite small, set off fromthe sternite by impresseclsutures; basistyleentire, large and conslticuous. Ninth tergite of peculiar confortnation,strongly elevated and corrugated,alnrostdivided by " cleepand narrow notchand groove,the lateral lobesthus formed again deeplydivided bya narrow U-shapenotch,the mesallobulelying aluroston edgewith
the flattenedface directedto the midline; the broaclerlateral lobes
narrowly rirnrnedwith black edge. Ninth sterniterelativelyinconspicuous,its appendagesrnall,rvith uroderatelylong yellorv setae,
the posterior
contiguousat the miclline. Basisty'lecorrspicuotls,
edge sharply rnargined and at its ventral angle produceclinto anacutespinouspoint ; caudalnargin of the stvle back from this spine
flattenedand sclerotizedon the lower portion, near the dististylewith a roundednotch that is filled r,r'ithrvhitish tnetnbrane,the dorsal angle of this notch producedinto a second,inwardly directedchitinizedpoint; dorsal apical part of style with long black setae.
Outer dististyle unique, appearingas a polishedscoop,the outerconvex face glabrous, the concaveinner surface provided with
abundantlong yellorvsetae. Inner dististl'lenot clearly evident in
the unique dry type. Eighth sternite sheathing,strongly narrowed
outwardly, the tip more or less circular; viewed from below theapex appearstruncated and filled with pale membrane; viewed
from behind, the sides of the shallow aperture are produced into
short polished lobes that bear dense brushes of relatively shortyellor,r'setae,the rnore ventral ones longer and stouter, decussate.

oct.,7e57 Buttetin of the Brooklyn Entornological,
Society

91

Habi,tat: California (Monterey County).
__Hol9lyfe: 3; H-agtingsReservation,"iorg Finch creek, May 21,
1943(J. M. Linsdal.) l AlexandercollectiJn.
f am unableat this time to indicatethe exact relationshipsof this'Hesperofly. fn someregardsit suggestsspeciesof the subgenus
tipula Alexander rather thJn Luiatipula Edward!, where I am
placingit. In the latter subgenusit suggestsspeciessuchas Tiputa(Lunatipula) splendezsDoine, and.alli-es,bui the actual relationship seemsdistant. It should be noted that the male hypopygium
was describedfrom the unique dty type, without mounting^oidissection.