Frorn,fft,eArrrers ANDMeoeznrn or Nenm,er, IlrstonY,
1939.
Ser. ll, vol. iii. p. 186,I*rwry

NEW OR, LITTLE-KNOWN TIPULTD.&(DIPTERA) . r.VI. NEOTROPICAL SPECIES.

186

Dr. C. P. Alexander on

New or little-knownTipulid n (Diptera).-LW . NeotropicalSpecies.By Cnanl,ns P. Ar,nxeNDER,Ph.D., tr'.R,.E.S.,
MassachusettsState College,Amherst, Massachusetts,
U.S.A.
As was the casewith the two precedingparts under thisgeneral title, in the present instalment I am considering
only speciestaken in the Oriente of Ecuador. This riclr
material was secured by William Clarke-Macintyre.
assistedby native collectors. I am very much indebted
to the collectorsfor their interest in saving these fragile
flies, the_types of the novelties herein described being
preservedin my collection of Tipulidre.
Epiphragmn (Epiphragmn)subsolatrir, sp. n.
Mesonotal prrescutum chiefly dark brown, the setrtunr.
scutellum and mediotergite yellow pollinose ; haltereschiefly pale yellow ; legs yellow, each femur u'ith tu-o
black _rings on outer half ; wings whitish subhvalipe.
irregularly fasciate with yellowish brown or tau-nv areasthat are narlow-ly bordered by dark brorvn ; nrale h.r-popygium with the interbasal rods slender. the extrepre
tips bent mesad into short points.
Male.-Length about 8-g mm. ; u'ing I0-l I nrnr. :antennn 2.5-3 mm.
Female.-Length about 10-12 rnrn. ; rving lo-12 nrnr.
Rostrum yellowish pollinose above, darker on sides ;palpi black. Antennm with scape and pedicel brownto dark brown ; basal segment of flagellum yellow to
obscure brownish yellow, the succeeding segments pale,outu'ardly passinginto dark brorvn ; verticils conspicuous.
Head chiefly dark brown, the orbits more reddish brorvn.
Pronotum yellow. Mesonotal prrescutun almost uniformly {?rk brown, the central portion of disk usuallyoccupied by stripes being rnore reddish brou-n ; extreme
posterior border of prrescutum, scutunl, scutellum, andmediotergite more yellowish pollinose. Pleura chiefly
dark brown, variegated with brownish yellow on ventral
sternopleurite, the dark colour involving also the pleurotergite and lateral portions of mediotergite. Haltereschiefly pale yellow. L"gs with the coxre and trochanters
yellow ; femora yellow, each with two black rings, onenew or little-krwwnTipulidre.

187

placed close to mid-length, the_other subt'erminal, thesevariable in extent, the outer subequal to the
io-t*hatyellow interspace ; in some individuals the extreme
fr*otul tip G further darkened ; tibire and tqgi lightyellow. frittg* with the groun_d-colour whitish subiryaline, irregularly fasciate with - yellowish brown or
tawny areas [ttat are narrowly bordergd by dark brown ;bands before cord. interconnectedin the radial and anal
fields ; band at cord usually disconnectedfrom the basal
dark mass but, in cases,narrowly interconnectedin outerend of cell B; a narrow oblique area of the white groundbeyond the cord, almost completgly traversing the wing,*"pututed from the pale area before cord only in outer"na of cell ; beyond cord, the dark areas rnore or lessrestricted to the ends of the veins ; Inore extensive
clark &rea,sat stigma, anterior cord and outer end ofcell Rs; veins yellow, darker in t!" patterned areas.
Venatibn : Bs only weakly angulated but usually shortspurred.at origin ; n'L-cLLat' basal third or fourth of cell
Lst M r.
Abdominal tergites yellowish brown, the pro-rym-alrings of the motJ basaf segmentsmore brownish black,the" caud.al borders narrowly pale ; sternites rveakly
bicolourecl,brou'n. the caudal borders rlarrowly yellow ;hypopygium brou-n. Ilale hvpopygiurn *it-h the interbasai ioat slencler.feebly bent at near mid-length, theextreme tips bent rnesadinto short points.
Hab. Ecuador (Oriente).
Holotypa,
6, Abitagua, altitude 1200metres, sep_tember-(Macintyre).
Allotopotype,?, pinned with type.
z0,lg3f
Paratopotypes, d?, November 1936, August' 15 to
September20, 1937(MacintYre\.
itt its wing pattern , Epiphragma (Epiphrogry) subso'

latrir is geneialy similar7o E. (8.) fqsglnennis(Sry) utd

E. (8.) {olatrir (O.tttt Sacken),but differs conspicuously
in the coloration, especiallyof the femora.
(Epiphragmn) nigroplagi,ata,sp. n.
Epipthragnxq,
General coloration brown, the prescutum yellorv rvith
and two large polished black areas
three brown stripes-femorayellow with trvo narrow blackon either side ;remainder of legs yellow ; wings pale yellow withrings,"or"llatebrown patlern ; rru-c'uat ne&r one-third the
ur,

188

Dr. C. P. Alexanderon

length of cell ; abdominal tergites dark brown, thesternites yellow, ringed with darker.
Male.-Length about 11mm. ; wing 12 mm. ; antennre
about 4 mm.
Iemale.-Length about 13 mm. ; *irg I2.5 mm.
Rostrum yellow to brownish yellow ; palpi black.
Antennre moderately long ; scape brownish- yellow ;pedicel relatively long, dark brown ; first flagellar segment light yellow, remainder of flagellum black ; flagellar
segments long-cylindrical, with conspicuous verticils.
Head brownish yellow, the posterior portion of vertex
with a transyersedark brown area.
Pronotum dark brown. Mesonotal prrescutum rvith
the ground-colour obscure yellow, polished in front,
more pollinose near suture, the usual prrescutal stripes
dark brown, extensive; two major polished black areas
on either side of prrescutum, one at humeral area. thesecond larger, occupying the entire lateral nrargin ofsclerite between the pseudosutural fovere ancl strture ;scutal lobes similarly polished black, the mecliarl area.
together with the scutellum,pruinose; postnotunrblack.
heavily grey pruinose. Pleura black, variegatecl s'ith
obscure brownish yellow on the dorsal sternopletrriteand ventral pteropleurite ; ventral stentopleurite rlark
brown. Halteres elongate, brown, the knobs a little
paler. Legs with the coxre blackenecl,the distal ends offore coxre yellow ; trochanters yellorv ; fenrora .\'ellou'.
with two black rings, one beyond micl-length.the othersubapical, these rings relatively narrow, subequal inextent' to the alternating yellow rings ; tibire obscureyellow with &n extensive pale brown ring just beyoncl
base, this paling outwardly into the ground-colour;
tarsi obscure yellow. Wings pale yellow, the costal
portion more saturated ; a heavy brown ocellatepattem.
the area at origin of ^Esnot connectedwith the one atcord ; a seriesof dark dashesin cell Znil A ; veins bron'n.
yellow in the grouncl areas. Venation : supernumerarv
cross-vein in cell C perpendiculari Rz+s+E
subequal torn41r,, the latter at near one-third the length of celllst Mr; cell Mt a little longer than its petiole.
Abdominal tergites darl< brown, at, least the basal
sternites yellou' ; in female, the bases of stemites
narrowly but conspicuously ringed with brown. llale

new or little-lcnpwnTipulide.

r89

hypopygium with the outer dististyle stronglY curved
at apex ; interbaserelativelY stout, the tip curved.
Hth. Ecuador(Oriente).
Holotypo,3, Abitagua, altitude 1200metres,March 29,
1937 (Macintyre). AllotoPotYPe,?, September 2L, 1937
(M a,cintyre).
Epipi,rog*o (Epiphragma) nigroplagiata.ts readily told
from utt described species by the conspicuous patternof the mesonotum.
Epiphragm& (Eytiphragma)deliberata,sp' n'General coloration brownish black, the prrescutal
stripes reddish brown ; antennreshort ; basal t"{ apicalthe remainder black ; halteres
flagbttar segmentsyellow,"
dtrEky, th; knob yellow ; femora obscure brownish
yellow on basal haif, outer portion. yel-lgw,enclosing a
Lroacl black subterminal ring ; tibire black, the -baseyettow, most extensively.so
narrou'ly, the apex broadly-yellorv; wings yellow, withon pcrsierit,r legs ; tarsi
a hiavy bro.u-ri pattern.
^ot.or. including about eight larger
cells Br. M anclCu' more washedarrd daiker costal
with brown ; margin of 2ncl anal cell $'ith five dark
are&s; abtlomen b-lack, the iutermecliate sternites withbroad, pale yellow, posterior margins.
Mali.-Length about 6.5-8 '''m. ; rving 7-10 mm.
Fem,ale.-Length about 9-f l mm. ; *'ilg 8-11 mrn'
Rostrum brow-nish black ; palpi black. Antennre
short. black, basal fusion segment bright yellow ; scapeslightly pruinose ; terminal-flagellar segment paling -toy"ito*. Head clark brown, the posterior orbits broadly
obscureora,nge.
Mesonotal"pr*..utum with the gro-un(l-colour dull
black, most intense on the humeral and lateral -port'iorrs;
stripes more reddish brorvn, occupying virtually lll of
the disk, the posterior interspaces poorly delimited ;media,n stripe *ot" or less tlivided on anterior ^portion
by a capillary dark vitta ; posterior sclerites of notum
biack, piuinose, heaviest on the mediotergite. In somespecim"^tttthe Inesonotum is almost,unifotTty blackened'
plleura uniformly blackened. Halteres with stem,dusky,
knob pale yeliow. Legs with the cox& blackened ;trochatiters bbr.nt" yello* ; femora with basal half
obscure brownish yeltow, deepeningto a ll&rrow brownish

190

Dr. U. P. Alex anderon

black,rilg, iort-- beyond mid-length ; clistal portion of
femur bright yellow, enclosinga Sroad black ring that isnearly twice as wide as either the yellow apex or subterminal ring. ; tjlire with basenarrowly yelloir, th. intermediateportion black, the tips yellow, it "latter narrowest(about the distal sixth) on iore legs, widest on hindtibia (distal three-fourths or upproi'imately so) tarsi;
yellow. Wings with the groorra-colour yellow, with'lurg",abrown
pattern,including
about eight
fealxand darker-eoloured costal areas; areas uI urr,,li.r,
origin _of8s and fork of Bs ocellate ; stigmal area solicliground-colour of cells no,,Ig and ctc exteisively infumed,
the centres of remaining interspacesbasad oi cord lessdistinctly clouded; outer raclial cells u'ith the groundcolour clear y-ellow; yellow marginal spots in raclial
and medial cells ; cell lst A with tlree, 2nd,A rvith fivemargin-aldark &reas,excluding the spots at, ends of veins ;v_einsdark, a little brighter in the yellow ground or"ur.
venation : Rs relative"ly short,, square ancl spurrecl atorigin , Rz+?++a_bor{one-half the length of Es ; ,L-.ct(
at near mid-length of cell lst M r.
Abdominal _tergites and hypopygitnn clar.li br.,u-rr tobrownishblack ; sternitestiaik-brown. the caurlalborclersof - segmentsthree to seven broacllv ancl conspieuouslv
pale yellow.
Hab. Ecuador (Oriente).
_^4!lotg?t,.4, Abitag_ua,altitude 1200 metres, April 7,1937 (Ma'cirytyrQ. Atlotopotype,?. paratopotypei, I
several_9?,April_5-lB, September21,tgBT (Micintyre)."6,"airiirr-

is readilylg;plrrqgryo (lniphragma)d,eliberata

guishedby the diagnosticfeatureslisted above,
especially'abclorninat
the coloration of- the antennre, legs ancl
sternites.
Epiphrag?lla,(Epiplrra,gnte)cni.ru. sp. n.

General coloration of notum clark bron-n. u-ith gre.\r
stripes ; antennrewith scape ancl pecricelpale. terprinal
segment of flagellum paling to yellou' ; hilteres r-ellou,
with the base of knob daikened ; fernora vellorrl *-ith
lyo clearly-definedblack rings at and beyond nricl-lengt5.
the remainderof legsyellow ; wingspale yellou-.patterirecl
with brown; a seriesof about ten biowrlspotsin cell cu,;
abdominal tergites brown ; sternites pale or:lrreousrrithnew or little-lcnownTipulidre.

191

dark brown incisures ; male hypoygium with the interbasea slendercurved spinousrod.
Male.-Length about l0 mm. ; wing 11'5 mm. ;antennreabout 3 mm.
Rostrum brown, pruinose; palpi black. Antennre ofmoderate length, if bent backward extending about towing-root or slightly beyond ; scape obscure yellow ;pedicel yellowish brown ; first flagellar segment clear
yellow, remainder of flagellum black, the terminal
segments paling to yellow ; flagellar segments subcylindrical, with unusually long verticils. Head buffy
brown, the anterior vertex with a conspicuouschestnutbrown area ; posterior vertex with a dusky median line.
Pronotum brownish yellow, with a capillary dark line.
Mesonotal prrcscutum with grey stripes, the interspaces
interruptedly dark brown ; lateral margin of prrescutum
narrowly blackenecl; posterior scleritesof notum chiefly
pruinose, eaeh scutal lobe darkened medialiy ; a black
spot on side of scuturn behind suture ; scutellum black,thinly pruinose; rnediotergite black, more heavily
pruinose on sides. Pleura variegated brownish blackand obseure brownish yellow, the former including the
propleura, mesepisternum,pleurotergite and meron, thelatter colour including the mesepimeron and sclerites
ventrad of base of halteres. Halteres pale yellow, thebase of knoh darkened. Legs with the cox& obscurevellou'. the frrre anrl nrirlrlle eoxre restrictedly darkened
nrerliallr-: tnr<'ltatttersr-ellou': f'erlroraYello'v, with two
narro\\'. elertrlv tlefinetl lllack rings at and beyond midlength. these narro\\-er than the yellou'apex and subternrinal ring enclosedby them ; tibire and tarsi uniformly
yellow. Wings with the ground-colour pale yellow,
conspicuously patterned with brown, including a seriesof seven or eight darker brown costal areas ; areas at
origin of -Es,eord, forks of Rr*r*n and Mr*r, m and stigma
ocelliform ; a linear series of about ten brown spots incell Cu ; margin of cell lst A with two dark areas,of 2nil A
with six areas, adrlitional to those at ends of veins ; veins
yellow, clarker in the clouded areas. Venation : Rr*r*o
the latter before mid-length of cell
longer than n'L-c'u,,.lst Mr.
Abdominal tergites brown ; sternites pale ochreous,the incisures narrowly dark brown ; hypopygium more

t92

Dr. C. P. Alexandet on

yellowish brown. Male hypopygium with the interbasesappearirg as slender ,,imbiJ rods, the curved tips
acute.
Hab. Ecuador (Oriente).
- ^!:to!ype; 6, abitlgua, altitude 1200metres,March Zg,
l9?7 (Macintyre). Paratopotype,sex? septemberzl,lgBT(Mani,ntyre).
_ Ep:phr?g,y @piphragmn) enira is generally similar toE.- (8.) d,eliberqta, n., differing co-nspicuo"uslyin the-sp.
coloration of the body,und' wing^s.
"of Both ,p".i"*Irg!,
have at least the teiminat segment
the antennal
flagellum pale yellow.
Epiphragma (Epiphragma)subeniro,sp. n.
General coloration of mesonotum dark brown, theprescutum with four reddish orange stripes ; antennre
black, thesegment9f flagelluttiorr'gu , pleura and,basalpleu.rgtergitealmost uniform brownish yirffori, without
m.alkingsj - legs with coxreuniformly pale, femora yellow
with two bla.g|lings on outer half ; *irgs yellow, h'eavily
patterned with brown, the markings in part ocelliform.
cell Cu without dark areasexceptat outei encl; abclo-"tdark brown, the basal sternites variegatedrviih yellow;
mal.elypopygium with the interbasJ bent at ; righranglebeforeapex.
Male.-Length about g.b mm. ; rving l0.b mm. .
antennreabout 3 mm.
Rostrum reddishbrown ; palpi black. Antennremoderately long, if bent backwaia extendirg about to wingroot or shofly beyond; scapeand pedicel black ; basil
segment of^flagellum orange, witli white pubescence,remainder of flagellum black ; flagellarsegme'ts nearly
cylindrical, rvith
lotg consplcuous-verticilslHeacldeep
orange,the ceutral portion of r.ertexextensivelydarker.
Mesonotal prrescutum rvith the grouncl-.oioor darkbrown, with four more reddish otor.gi stripes, the intermediatepair separatedby u capillaty clatk iitta ; seutum
and mediotergite dark brorin, ihe surface sparsely
pruinose,the scutellum more testaeeousyellow. pleura,ttq pleurotergite almost uniformly brbwnish yello*,.
without markings. Halteresunifornily
infuscated. Legs
-yellowwith the coxa uniformly brou'nish
to obscuie
yellow ; trochanters yellow ; femora-yellow, rvith tu-oTipulidre.
newor little-lcnoum

193

narrow black rings, the nanower one just beyond midlength, the outer dark ring subterminal, nearly twice
as wide as the inner and subequal to either the yellow
apex or yellow subterminal ring ; tibire and tarsi uniformly yellow. Wings rvith the ground-colour yellow,
rnore saturated in the prearcular and costal fields ;a heavy brown pattern, including about eight darker costal
areas ; a distinct ocellus surrounding origin of -Es, with
more-broken and less-evidentocelli centreing at anterior
cord and fork of M s2 i stigmal area solidly darkened;ground-colour of cells R o, Ru, Crr,and Ist A suffusedwith
duskSr; dark spots in cell Cu lacking or confined to outer
end ; outer end of cell Lst Awith a singledark area between
veins, cell 2nd,A with about six such marginal areas;veins pale brown, yellow in the flavous areas. Venation :shorter than Rz+si.Bs nearly square at origin ) Rz+s+E
m-cu at mid-length of cell lst M r.
Abdominal tergites dark brown ; third and succeedingsternites u'ith posterior borders conspicuously yellow ;seconclsternite u'ith this ltand subapical; hypopygiumbrou,'nishyellou'. Male hypopygium u'ith the interbase
bent at a right-angle before apex.
Hab. Ecuador (Oriente).
Holoty,pa,
6, Abitagua, altitude I200 metres,September
20, I937 (Macintyre).
Epiphragnxa (Epiphragma,) subenira, while generally
similai to E. (8.) enira, sp. n., is entirely distinct in all
cletailsof colorationof body and wings,and in the structureof the interbasalrods of the male hypopygium.
(Epiphragma)diademo'sp. n.
Eptiphrag?n,a
Generalcolorationof mesouotumdark brown, the usual
prrescutalstripes tleep chestnut-brou'n,the intermediatepair clark brou-tt oll posterior half ; antennal flagellum
te1'oncl tlte first segrnent uniformly black; scutellum
and mecliarlarea of icutum pale ; pleura yellow, striped
longitudinally with dark brown, includilg a broad dorsal
strile from the cervical region to tl" pleurotergite,
steinopleurite and meron abruptly pale ; legs yellorv,
the femora with two broad black rings ; u'ings yellorv,
heavily patterned with brown, the areaspartly ocelliform ;abclominaltergites beyond the first brorvnishblack,

194

Dr. C. P. Alexanderon

Male,-,Length about 1l rnm. ; rving 12 mrn. ; antennre
about 3.3 mm.
Female.-Length about l2-I4 rnm. ; wing ll-12 mm.
Rostrum ochraceous; palpi black. Antennre of moderate length, in male if bent backwardendingshortlybefore
wing_-root; sc_apeyellow, pedicel dark brown, paler
basally ; first flagellar segment light orange, remainder offlagellum uniformly black, the segmentswith conspicuous
verticils. Head with front and anterior vertei pale ;remainder of head deep chestnut-brown, the central
portion of vertex more infuscated.
Pronotum dark brown medially and on ventral portions,the sublateral region broadlyyellow, confluent behind rvith
the pale dorsopleural region. Mesonotal prrescutum withmost of the usual stripes medium brown to chestnutbrown, vaguely separated by darker lines, the humeraland lateral portions brownish black ; posterior half ofintermediate prrescutal stripes darkened ; scutal lobes
extensively dark brown, the median are& pale ; scutellunr
pale, the parascutella dark ; mediotergite with eentral
portion grey pruinose, the borders darkened. Pleura
yellow, striped longitudinally with dark brown, inclucling
a dorsal stripe from the cervical region to the pleurotergite, the eentre of the latter pale ; a nlore ventral
dark stripe, involving chiefly the mid-coxre ; pale groundstripes occupying the dorsopleural region and the sternopleural and meral portions. Halteres pale, the base ofknob infuscated, the apex a little paler. Legs with thefore coxre yellow to more or less infuscated ; mid-coxre
blachened; posterior coxre obscure yellow ; trochanters
yellow ; femora yellow, with two broad black rings at
and beyond mid-length ; tibire and tarsi yellow. Wings
yellow, the costal portion more saturated ; a heavy
brown pattern, darker along the costal border ; ocelliform
areas centering at origin of -Bs, anterior cord and forkof Mr*r; dark areasin cell Cu, few and restricted to outerend of cell, the basal half u'ashed with darker ; dark
marginal spots at outer end of cell Ist A varying in number
from one to three, in cell Znd A folur or five ; veins yellou',darker in the clouded areas. Venation : Rr*r*n andRr*, subequal ; nr-cu at or before mid-length of celllst M r.
Abdominal tergites beyond the first brownish black ;basal sternites lnore uniformlv yellow ; hypop.ygiunr

neu or little-krwwnTipulidre.

195

obscure yellow. Male hypopygium with the tergal lobes
low and broad. fnterbasal rods relatively short, at apex
curved laterad into acute points.
Hab. Ecuador (Oriente).
Holot'ype,S,Abitagua, altitude 1200metres, August 15,
1937 (Macintyre). Allotoptotypa,?, April 4, 1937. Paratopotypa,?, September20, 1937 (Macintyre).
Epiphragnut (Epiphragmn) diadema is closest to E. (8.)
en'ira, sp. n., and E. (8.) subenira, sp. n., differing
especially in the pattern of the mesonotal prrescutum andpleura. It is certain that there are numerous generally
sirnilar but quite distinct species within this particularcomplex of forms.
Epiphragrna (Epiphragma) ranthomela, sp. n.
General coloration black, the prrescutum variegatedrvith dark brown ; antennre elongate, black, the first,
flagellar segment orange ; femora black, the base, apex
ancl a subterminal ring narrou'ly yellow ; tibire yellow
at extrellle base, follon'ed by a broad black ring, the tip
yellou' ; tarsi yellou' ; u'ings pale yellorv, rvith a heavy
brown pattern, including large ocelliform areas ; abdorninal tergites black, the caudal borders restrictedly
silvery ; sternites with posterior borders broadly yellow.
Femnle.-Length about l0 mm. ; wing 9 mm. ; antennre
about 3.3 mm
Rostrurn and palpi black. Antennre black, the basal
segment of flagellum abruptly orange ; antennre relatively
long, especially for the female sex. if'bent backward
extending to shortly beyond base of abdomen ; flagellar
segments cylindrical, u'ith conspicuousverticils. Headrlark brown. the front and ltarrow orbits somewhat
lnore reddish brou'n. the posterior vertex and genesliglrtly more pruinose.
Pronoturrr obscure yellow, the anterior notum andpleura rnore infuscated ; pretergites obscure yellow.
Mesonotal prrescutum with the disk chiefly tight brown,
variegated with black, including a conspicuous medianvitta and broad lateral and cephalic borders, slightlyinterrupted at the humeri ; scutum with the groundcolour brownish yellow pollinose, each lobe with disk
extensively darkened, the median line with a narrow
vitta ; scutelltrmand mediotergite black, sparselypruinose.
Pleura and pleurotergitechiefly black. ineluding the dorsol3*

196

Dr. C. P. Alexanderom

pleural region, the meral area a trifle paler. Halteresqo{era-tely_long, infuscated, the base of stem and apexof knob obscure yellow. Legs u'ith the coxre bla^ck,
the tips narrowly yellow, of the fore coxre the distai
Irltf more pieeous; trochanters brownish black ; femora
black, the basesnarrowly yellow, somewhat more extensively so on posterior legs ; a narrow ring at near twothirds to three-fourths the length, together *ith the apex,
yellow, the black subterminal ring very broad ; tibire
extensiyely black on proximal portion, the extreme
base yellow, the dark colour most extensive on the fore
and middle tibire where more than three-fourths the total
length is included, narrower on posterior legs u,here
about one-half the length is involvecl ; tips of iinirc andall tarsi yellow. Wings relatively ,rurroin,. pale yellou,,
the costal border more sat'rated yellou' ; a heavy"brownpattern, including about eight darker costal a"eas thatare more extensive than the interspaces, especially themore basal ones ; ocellate rings centieing at 6rigin of .Bs,
cord and folk of M 112,with a further incomplele oeellus
at arculus, the ringsl?rye, enclosingextensivd pale areas ;stigmal area not ocelliform ; cells Cu ancl lst-A clouclecl.
the former with only one or two darker areas: celllst A with four marginal darkenings. cell 2nd,A u,ith
about eigh_t_,
excluding the areas at enclsof veins ; veinsbrown, yellorv in the flavous areas. Venation : m-c,t(
at near two-thirds the length of cell lsl M 2.
Abdominal tergit_esblack, the eaudal margins of thesegments restrictedly silvery ; sternites similar, theposterior margins more broadly and eonspieuouslyyellow.
Hab. Eeuador (Oriente)H,ololu_pet?, Abitagua. altiturle 1200 nretres. April ir.
I 937 (Macintgre).
.-In its general appearanee.especialh-the blackelecl
tibire, Epipthra,g?n&(Epiphragr,ril .r.rtitltotttelrris prost
similar to E. (8.) deliberafa.sp. n.. but is entirelv clistipet.
The unusually long antennre of the fenrale iriclicate aneven longer organ in the male.
Epiphragma (Epiph,ragma)ory,phalltts. sp. n.
General coloration of prrescutum bright bron-n : antennre with basal three or four segments pale. theremainder black ; pronotum and dorsopleural rnelrbrane'ne'wor little-knownTipulidre.

1e7<lbscureyellow, the remainder of pleura black, including
the broacl bases of the coxre; halteres relatively long,infuscated ; legs yellow, the femora u'ith a narrowbrown subterminal ring ; rvings yellow, the grounclcolour lnore or less rvashedrvith brown, heavily patternedwith darker brou.tt, the rnarkings in part ocelliform ;a flattened cornpressedrnale hypopygium with the recleagus
blacle, narrowed into a long apical point, protruding
ventrad ancl cephalad from the genital chamber.
Male.-Length about 8-8'5 Inm. ; rving 9-9'5 mnr. ;antennre about' 2'8-B mrn.
Female.-Length about 9-10 llun. ; wing 9'5-10'5 mnl.
Rostrum brou'nish black ; palpi black. Antennre
with the basal three or four segments yellow, the scapesornetimes a trifle darker ; remainder of flagellum black.
Head brownish gre/, the posterior vertex with a darker
median line ; front and narrow posterior orbits moreyellou'.
Pronottrur obscure yellou'. u'ith a narrow median vitta.
Mesonotal prrescutunr u'itlt the stripes bright brown,
the interspaces clark brou'tr to brou'nish black, mostconspicuouson lateral nrargins behincl the pseudosutural
fovere and less evidently on tr,nterior portion of sclerite ;posterior interspaces and median clark vitta sometimes
restricted and virtually confluent with the ground ;scutum reddish browtt, the lobes darkened ; scutellum
clark ; mediotergite dark medially, the caudal and lateral
portions obscure yellou.. Pleura ancl pleurotergite black,
only the dorsopleural region abrtrptly yellorv. Halteres
relatively elongate. ittftrscatetl. I-,egs rt'ith the coxre
blackened, the tips abrtrptl.r' vellou-. involving about
the distal half' oll rnirlrlle unrI posterirlr legs, narrower
oll fore llair' ; tt'oc'ltatrters 1'ellou': f'ernora brownish
velkrn'. c.letrrer'.r'ellou-trt llase tltrcl tip, with a narrow
bron'n subtenninul ring ; tibire and tarsi yellow. Wings
u'ith the groult(l-colotrr yellow, more saturated in costal
portion ; tnuch of disk washed with brown but notforming distinct pale-margined centres to the interspacesas in many species,the rnost conspicuousdarkening
involving the outer radial field and cells M . C u. Lst A
and 2nil A ; a heavy brown pattern, including about
eight still darker brown costal areas ; ocellate areas
centreing at origin of Bs and corcl ; stigmal area, solid ;198

Dr. C. P. Alexanderon

an irregrrlar series of more or less confluent dark spots
in cell cu'; dark marginal spots at ends of longitudlinal
veins, in cases confluent along extreme outei margin
bo! leaving cent al pale droplets at ends of cells ; ooTut
end of cell rst A with two or three, of znd A withfrom six to eight dark spots, additional to areas on theveins themselves; veins pale brown, darker in theheavily patterned areas, yellower in the flavous portions.
Venation : _-Bsangulated and spurred at origin ; m-c,ubefore mid-length of cell LstM r.
Abdominal tergites brownish black, the incisures
somewhat brightened ; sternites more obscure yellow.
variegated with darker, the lateral portions blaci<enecl;h,rnonrgr-"T brownish yell'w. Male hypopygium u,itlrth"-.tergal lobes long and slender. Out,if ai*li-style witha distinct lote or shoulder on outer margin r""u,r base,the style gradually narrowed outwar.lli into a long
curved point. 'CIdeagusentirely different from the typE
normal in the genus, appearing as a flattened comprer*6a
blade that narrows to a slender tip, in a position^of rest
directed ventrad and cephalad, showing conspieuously
even with a hand lens.
Hab. Ecuador (Oriente).
Holot_upg,_C,.
Abitagua, altitude I 200 rnetres,september.
18, 1937(Mac'intyre). Allotopoty,pe,?, in copula*itn type.
_
Paratopotypes,several males and females, March zi'toApril ?9, 1937; August 20, lgBZ ; September 1g-20,
1937(Macintyre).
. rn the peculiar and somewhat remarkable red.eagus,
the present ny i! entirely_distinct from all regional spe-ciesof the genus so far described.
Epiphrcn{l?n&(Epipln agnm) hirtistylrttct.sp. n.
Getreralcoloration of' trotum obscure1'ellou'.pattenrecl
with brorvtr; tertninal segnrentsof' nraxillar.r'palpi ancl
antennre abruptly yellon-: antennre elongate. tne basal
flagellar segmentsbicolourecl; haltereselongate.obscureyellow ; __femorablack ; tibir black. the tipr vellou- :t3,rsi yelloy ; wings palg yellorv. all cells ibunclantly
dotted rvith brown, with about seven slightll' larger
areas that are costal in distribution ; ,,r-, u- shorllv
before mid-length of_cell lst Mr: nrale hypopygium rvitir
the outer face of basistyle u'ith unusuiily long andconspicuoussetre.

rle'wor littl,e-krwwrt,Tipulida:.

199

Male.-Length about,9'5-10 mm. ; wilg 9'5-10'5mm' ;antenna about 4'2 mm.
Rostrum black ; palpi black, the terminal lggTent
abruptly yellow. Antennre (male) elongate,, if bent
backward bxtending to about base of third abdominal
segment ; basal f'olrt segments yellow, the succeeding
thiee bicolorous, the base narrowly black, the apex
yellow, with the arnount of black becomittg-moreextensive
bn outer segments: outer half of flagellTm black, thesmall termiial segment abruptly pale yellow ; flagellar
segrnents long-cylinclrical. with conspicuous erect setre
ancl sparselorlger verticils. Hgad light orarlge.
Pronotottr y6llotr', narrowly darkened medially' Mesoobscure yello*',, restrictedly-.patterned
notal pr**.oirrwith birown, including the lateral and cephalic borders
and less disiinct discallmarkings ; an inverted V-shaped
brown markin g at mid-length of prrescutum,behind which
the median u]t"u is mot-e pruinose, bordered laterally
by browu ; scutal lobes rveakly darkened ; scutellum
utta rnecliotergite brou'tr. pruinose, especially-the latter'
Pleura obscuie yellon'. tt'ith & conspicuotlsdark brown
longituclinal stripe extencling f rorn the cervical region,
puriitrg above tie fore ro*b to base of halteres and- sharply defined.^ Halteresit.otoLrgite, its clorsal edge_
ilongate,- obscure yellow. Legs with the fore coxre
chiefry iirfuscated, iemaining cg{re -yellow ; . trochantersbrowi ; femora black : tibire black, the tip! _abruptly
yellow, including about the distal fourth or fifth, more
Lxtensive on po"steriorlegs ; tarsi yellow. - Wings p?1."
yellow, all ."ilt abunclantty dotted u'ith brown, witll
ibout sevenslightly larger arbasthat are costally arrauged,
the seeond at brigin of ,Bs, third at th9 supernumerarycross-vein irr cell O, fourth at fork of Sc ; last area at
end of vein R a 'tr[
; lnore restricted seamsalong pord, outerend of' cell Isir, fork of M t1o and ends of the longitudinal veins ; th-e only indicition of the usual ocellate
an area surroundi"gpattern colnlnon in the genus_includes
ittt origin of fis ; veiis yelloy, darker in the cloudeda,reas. venatiou t Rr*, and R2 subequal ; ,Bs nearlysquare at origin ; peliole of cell M r -ftolt about onefrif to two-thiids thd ceil ; nx-cushortlv before mid-lengthof cell Lst M r.
Abdominai tergites 'redium browu : sternites yellow.
ri'ged rvith brou,rion basalportiolls ; hvPoltygiurnbrowtt.

200

Dr. C. P. AlexanderonMale!.rngrrsium_ with the basistyle provided withunusually long and strong setre on outef face of distal
half, these hairs exceedi"gin length the basistyleitself;
shorter strong setre on proximaf hap of rn*ul face ofba,sistyle. Dististyles relatively small, the outer terminatitg ir ? simplehook. rnterbasewith the lobe relatively
short, the tip pppe_aring
as a short acutespine.
Hab. Ecuador(Oriente;.
__H-o!otyne_d,+bitagua, altitude 1200metres,september
z a6, Aprii t8-20,??,!93_1(Macintyre). Paratopotypes,
1937(Macintyre).
Epiphragma (Epiphragma) hirtistyrata is so distinct
from all other describedspeciestliat any comparison
with othersis unnecessary.
Austrolimnophi,la persessilis,sp. n.
General coloration of prrescutum grev, with threebrorvn stripes ; scape light yellow ; p6url'pale yellow;
yellowwings
subrryaline,;-clouds thi stigmi, .rcty palel"g*brown ; very pale brownat, origin-cellof Rsi uiorrg
cord and on outer end of cell lsl M z ;M L broadl!
sessile; nx-c1lat near one-third the length of celi LstM ;r'abdominal tergites weakly bicolorous, dark bro*.11,th;
caudal margins of the segmentsnarrowly obscureyelioro'.
Female.-L.lgth about 8.5 mm. ; wing g.b rnnr.
Rostrum yellg*' ; palpi brownish blaJk, the basal segrnent-yellow. Antennre rf ith scapelight yellow ; peclic6l
pale brown ;.flagellum dark brown ; flagellar segments
cylindrical, with conspicuous verticils tf,at "*."6d thesegments in length ;__segm_entsgradually increasing inlength out'wardly. Head browriish griy, the occTput
obscureyellorv.
Pronotum pale yellou'. Mesonotal pr'rescuttung,it[
lht ground-colottrgrey chiefly corrcealeiiby t hree clarkbrown stripes, tlte laterals ies clearly tlelinritecl, theinterspaces poorl,n-clefined; scutul loLes clarli brown,
the median_region a little paler i posterior sclerites ofnotum dark brorvn. Pleura pale ]eilow, the pleurotergite a little darker. Halteres very long, pui", the^andknobs weakly darkened. Legs with the .oi*
trochanters pale yellow ; rem-ainder of legs unifonlrl.\,yellow, the terminal segmentsrveakly infusiated ; clau,,small, simple. Wings subhyaline ; stigma long-oval,

new o'r little-lcnownTipulidre.

201

very pale brown ; scarcely evident clouds at -origin. ofRs, uiottg cord and al outer end of cell lsf M-, ; veinsclark br6wn. Macrotrichia of veins relatively short.
about opposite fork of
Venation : Sc long,-R*2,-at Srn ending
a little longer thanit; tip ;Rr*r*n, Sc, longer
R-r'i'hr*rln reLtively short, less than twice r-n1'; cell
M'r'broiaiy ttssile, rn-onvein Mrabout one-halfits length
beyond rork of M s2 i rn-cu at near one-third the lengthof cell lst M r.
bicoloured, dark brown,
Abdominai tergites weakly-segments
narrowly obscure
the caud.al murgi-tts of theyellow ; sternites obscure yellow ; cerci strongly upcurved, horn-yellow.
Hab. Ecuador (Oriente).
Holotype,g, e.bitagua,altitude 1200rnetres,September
2I, 1937(Ma,ci'ntyre).
Austrolimnopnita persessilisis very distinct from theother speciesio far d.escribed. In the sessile-cell M L it
resemblesA. pacifera Alexand.er, of Btazil, but in allother respectsis quite distinct.
sp. 11.
Austrolirnno'phila nernatoceru,,

General coloration brownish black ; antennre (rnale)equal in length to the wing ; flagellar segment's1o1g.ytittdtical, ivith numerous long- scattered verticils ;#trg* obscure yellorv, the colour almost obliterated_by
five-broad traniverse brown bands ; ,Sclong, Sc, ending
beyond fork of Rr*r*^, cell M 1 present ; tnale hypopygio- with the birter clistist,v-leterminating in trvo
small teeth.
Male.-Ilength abottt 6 l]1lll. ; u'ing ti mrn. ; antenn&
about 6 mtn.
Rostrunr uncl iralpi lllar:k. Antennre(rnale)of unusual
length. subecl.,olto the bocly or wing, b;o11 throughout
ot Iltt otrter segtnetltsa trifle paler ; flagellar-segments
long-cylindricall rvith numerous long scattered verticils
thal ure about two-thirds as long as the segments;
terminal segmentshorter. Head brownish black.
Pronoturi and mesonotumalmost,uniformly brownish
black, without clearly definedmarkings. Pleura similarly
darkened, the vential sclerites a trifle paler- Haltereswith stem pale, knob dark brown. Legs with the cox&

202

Dr. 0. P. Alex anderon

testaceous, the fore pair darker ; trochanters obscureyellow ; femora and tibia dark brown, the tarsi a little
p_aler. wings with_the ground-colour obscure yellow,
almost obliterated by five broad transverse "brownbands ; first band prearcular, almost confluent with thesecond ; third band ab level of origin of .Es and end of
2nd A, somewhat paler and broken in cell M ancl hereinterconnected with second band ; fourth band at cord :outermost.band- occupying the broad apex, a little
vgiegat.4_by pale spots; veins brown, witfr conspieuous
trichia. venation : 8c long, sr, ending beyond ^fork of
Rr*r*n, Bt, nea,.I.itr_ tip ; F-s long, ang"ulatLd at origin,in alignment, with Rr*i*n; basaliectiJn of R, and r-nx
in transverse alignmfu t- cell r[! subequal in"length il
its petiole_; cell lsl M z long u,rd tturroi, rvith nL-c,tlat
near two-fifths its length ; anterior arculus broken.
Abdomen,.in-cluding hypopygig-, black. Male hypop-ygium with the outer- dis[istyle small, subcylindiiial,
the^apex with two small teeth, the outer -ord slender;surface of style with numerous coarse setre. rnner
4lstistyle small, terminating in a short obtuse point.
,Ddeagus short.
Hab. Ecuador (Oriente).
4:tota_pe:dl, Abitagua, altitucle 1200rnetres,August 20,
1937(Macintyre).
r am referring
lhr present fly to Au,strolimnophita o'
the basisof the broken arculus,tlioughwith mucl
{uestion.
The. fly is qui_te different from uiy hitherto described
species,gspegia-llyin the elongate antenna and unusuallyheavily banded wings.
Pse'ud,olimnophila
plu,to,sp. n.
General col'r'ation brorvnish black ; antellna, black,the pedicel brou'nish y-ello*'; halteres obseure yellow;
tegj brown, the tarsi paling to .1,eJlou; u.ingsvery
slrongly"andsuffused u'ith brorvn, stigma aircl setlnls on-corcl
orrl"1.
end of cell lst M, darkei br-orvnI Bz+,]+oshort ; veins .R,
and n 4_extending generally poruilei iu one anotherl
so_cell Rz at margin is verf ttiae ; rn-ctc at from onehalf to two-thirds the lengtli of' cell LstM
r.
Male.-Length about 1-z.s rnm. ; rn'ii.g 7.b-g nrnr.
;
antennreabout 1.5 mm.
remale.-Length about g lrtt'. ; u,ing g.5 'u'.

nuw or little-lonown'l'ipulidre.

203

Rostrutn moderately long, about tlne-third the rernainclerof head, black throughout ; palpi black. Antennre
with scapeand flagellum black, pedicel brownish yellow-;
flagellar segments oval to long-oval, thg verticils
relatively tong and conspicuous; terminal segment
longer tlran the penultimate. Head brownish black ;anterior vertex about one-third the width of front.
Thorax uniformly brownish black. Halteres obscureyellou,. the base of stem a little tlarkeuecl. Legs-with
Ltre coxre blacli to brorvnish black ; trochanters yellow ;{'entora aucl tibire brolvrl, thc tarsi paling to yellow.
Wings ver.y strorrgly suffusetl rvith trrown, cell Sc above
the Jrculus restriciedly rllore yellorvish ; stigma elongate,
clarker brown ; darker seams along corcl and outer endof' cell ls/ M z i paler longitudinal streaks in cells .B and
M ; veins clark- brown, paler on basal third of' wing.
Macrotrichia of veins abundant, long ancl conspicuous.
Venation : ,Sc,ending just beyond fork of -Es, Sc, at lip ;Rr*r** in lonfitudinal alignrnent with fts, shorter thanRr*"'ancl about tu'ice the basal section of Rsl Rr!, lrom
otr"ancl one-half to ttt'o times Rz ) tip of vein Rs bent
strongly towarcl wing-tip, greatly rvidening-_cell Rr,
veins-.B, and .Eo thus extending generally parallel to one
another throughout their lengths ; cell M r aPProximatelV
twice its petiole ; 'm-cu front one-halt' to two-thirds
the length of the small cell l sf M , ; anterior arculus
present.
Abdomen brou,nish black, the sternites. and in cases.
the intermecliate tergites, a little paler ; hypopygium
brownish yellou.. Male hypopvgiunt rvith the outerdististyle ; simple r'ocl. its tip ir gently curved spine ;surface of style n'ith alrturdatrt setre oll tnesal face, theventral toutgitt befbre tlpex with several a,ppressedteeth.
Inner dististvle snrall.
Ha.b. Ecttaclor(Orieute).
Holotype. 6, Abitagu&, altitude 1200 nretres, April 20,
1937 @Iuciniyre). .q'ttotopotype,,?, April 7, 1937' Parutopotypa,-is3, rvith the allotype.
^
entirely distinct frotn all
Pidtd,oli*nophila pluto
bf the Limnophilaria in the Neotropicalother species-superficially
it most iesemblesLimnophila charort
fauna.
^ofsout[ern Brazil, but lnay be told readily
Alexander,[.y tfte venation anrl strtreture of tlte' tnttle hypopYgiuur.

20+

Dr. C. P. Alex andet on

Polymera(Polymera)clausa,sp. n.
General coloration of mesonotum testaceousbrown;yellow,withabroaclblack
dorsal stripe ; antennre
ryleu.ra(male) subequg,lto or longer than body, flaleilum black,tlr. segmentsbinodose; [albres and i.g, f,lack *ings;
with a uniform brown tinge, unpatt"rrrX i Rz+a+tshor''t.

about twice r-n't,; cell tJr M zilosed, ."il rt;";abdomenblack, hypopygiu- y6llowish brown. b;esent
Male.-Length about-6 mm. ; wing 7 mm. ; antenrrre
about 6 mm.
Femnle.-Length about 7 mm. ; wing 6.b-7 mm.
;
antennreabout,2.b-2.6mm.
Rostrum and.palpi black. Antennreblack throughout,in male applgximattly as long as or slightly lo'gei tharibody ; flagellar segments *"-ukly binoEosJ; in- female,antennre short a'd normal for this sex. Head d.ark
brownish grey.
Mesonotumuniformly light testaceousbrown. pleurawith a broad black dorcil stripe extending from thecervicalregion to the baseof abdomen; vential pleurites
pale yello_w. Halteres dark throughout. Legs *itt thecoxre and trochanters yellow ; femora clirk brown,
the basesnarrowly obscure yellorv ; tibire ancl all tarsi
black. witgs with a uniform brorvn tinge, unpatternecl,
the costal border a trifle ntore vellowi"sh; veins ancl
trichia brown. venation : sr, e'ding s6ortly before
lbrk of frr*1*n, Bc? near its trp ; firi, uppro"imately
two-thirds .Bs and from tu'o-thirds^ fo ihree-fourths
RZttj R_ryr*n_sho-rt,
about twice r-n't,; cell M, present;
cell lsf M z closed by-the retention of m, the cell longand narrow ; nx-cu about its own length, or slightl!
less,beyondthe fork of M.
Abdomenblack ; hypopygiuruyellou,ishbrou'n.
Ha,,b.Ecuador (Oriente).
, Holgtape,g, 4bitagua, altitude 1200rnetres,september
18.^ 1q37 (Macintyre). Attotoltoty1te, ?. paratopotypes,
3 ?3,-September
18-22,tgBZ(Maiintyre).
- Polyn'Le?'(r,_(Polymera)clausa is teaaity told fro'r all
known mernbersof the genus by
the closed cell lsf rI ,-blackenedof' the wings. The unifornrly
antenrrreor.,itarsi, in conjunction rvith the presenceof cell M L of' the-the
w1ngl, similarly distinguishes
fly from murt c,f its
relatives.

new or little-knoumTipulidre.

205

sp. n.
Polymera(Polyntera)albogenualis,
General coloration of mesonotum cinnatnon brown,
unmarkecl ; antennre (male) very loug, more than onehalf longer than body, the flagellar segments llinodose ;femora dark brown, the tips narrowly ancl abruptly
white ; tarsi dirty brownish white ; wings tingecl withgrey ; R2+s+4short, about one-fifth the length of veitr
Rr*, i R*o a little shorter than ,Esand about three tirnes
m-cu; abdomen,including hypopygium, black.
Male.-Length about 6 mnr. ; rving 7 nlm. ; antennre
about 10 mm.
Rostrum obscure yellou' ; palpi testaceous brown.
Antennre very long, more than one-half longer than the
body ; scape brou'n, pedicel brownish yellou' ; fl agellar
segments pale brown, the tips narrowly pale ; fl agellar
segmentselongate, binodose,with very long outspreacling
verticils on both nodes. Head dark brownish grey.
Mesonotumuniformly light cinnamonbrown, unmarked.
Pleura with a broad brorvnish black longitudinal stripe
on dorsal portiou, the sternopleurite and meron abruptly
light yellow. Halteres pale brownish yellow. Legs withthe fore coxredarkened basally, yellow at tips ; rema,ining
coxre and all trochanters light yellow ; femora darkbrown, the bases a little paler, the tips narrowly (about
0'3 mm.) but conspicuouslyrvhite, the amount subequalon all legs ; tibire brown, the extreme tips very narrowly
rrnd insensibly paler ; all tarsi paling to dirty brownish
u-lrite. Wings with a grevish tinge, utlltrarkecl; veinsarrrl trichia darker. Venation : Rz+a+.rottl.v a little
longer than basal section of' R-orrnclalrotrt orte-fifth thelength of vein Rr*, alone: frr+s illrottt three tirtres Rr*r*nartd onlv a little sltot'tet't ltrttt ,Rs: t'ell fI t present;
tn-crrat fbrli ot'JI .
black.
Alrrlonren.inc'ltrrlittgh.r-polx-gitutt.
Hab. Ecuaclot'(Oriente).
ilolotype. C, Rio Blanco, Itear Baflos, Tungurahua,
altitude 1700nretres.June 2, 1937(Macintyre).
The rnost sirnilar speciesis Polymera (Polymera) albo'
geniculataAlexander, likewise from Ecuador, which has
the femoral tips similarly whitened, cliffering in thepatterned prrescutum and seutum and in the venational
details, &s the long R r*r+n rvhielt is allottt two-thirds
as long as fr213,

206

Dr. C. P. Alexanderom

Atarba (Atarba) id,oneo,sp. n.
General coloration brown, the prrescutum more reddish
lrorvn ; ant3nna_(male) very lbng, nearly as long asbq{y ; flagellurn black throughout,- the longest verticils
solitary on. each.segment, unilaterally uttiroged ; legs
yellow ; wings with a weak brown tittge, *tig-u pai.
-cellbrown ; cell Rz at margin wider thany'ir, fr,-t ,,shortly before mid-length-of the rectangular celi lst Mr;abdominal tergites brownish black, hypopygium yeilo# ;maJe hypopysirlm with the redeagus ietativety short"and stout,_parallel-sided,
the apex nol expanded.
Male.-Length about 4'b mm. ; wing b mm. ; antennre
about 4 mm.
Rostrum brown ; palpi dark brown. Antennre (male)'scapeverJr lon$, nearly equal in length to the body ;"and pedicel palg brown, flagellum black ; flagellar
segments long-cylindrical, clothed u'ith abunclant longereet pale setre; each segment with a single 1lc,u'erfiil
seta, subequalin length to the segment,inserteclon basalh,alf of segment,unili,terally u"tu,iged ; terminal segment
about one-third the penultimate. Head clark browi.
Mesonotum almost uniform brown. the prascutum
somewhat more reddish brown. the posterior sclerites
darker. Pleura browtr, the sternopleuriteobscureyellou,.
Halteres clusky. the extreme base of stem pale. Legs
with the coxre ancl troehanters pale brown : relnaind6r
of legs yellow, the terminal tarsal segrnents-l\rings lnore infuscated ; no darkening of fernoraltips.
rvith a weakbrownish tinge, the prearcular aircl costal borders moreyellowish ; stigma pale brown, ill-defined ; veins brown.
Venation : ,Scshort, Sr, ending opposite origin of' Bs or.
virtually so, Stz eloseto tip ; .BJ irnusually--short, only
a little longer than basal section of Br; branchesof r?s
generally parallel to one another so cell R z at margin is
wider than cell Rs ; cell lst M , rectangular,nx-crlshortly
before mid-length. rn brunneicornistne branchesof .Bs
diverge at tips so cell R, at margin is considerablvmore
extensivethan Rz.
Abdominal tergites dark brown to brownish black.
the sternites a little more yellowish brown ; hypopygiurn
yellow. Sternal plate of rnale hypopygium rtitfi" s-hortlateral spines, the caudal margin truncate. Basistylewith a eonspieuousinterbasaf fobe, Outer rlististvle

Tipulidre.
neu)or lit,tl,e-lr.nown

207

rlilatecl beyondmid-length,thencenamowedinto a slendet'
point, the spinesof outer margin about eight in number,on the apical point rvith only a single spine before thebifid apex ; face of style with supplementaryroughenings
and spinulre. fnner distyle relatively stout, gently
curved. .a0deagusshort and stout. parallel-sided.theapex not expanded.
Hab. Ecuador (Oriente).
Holotype,6, Abitagua, altitude 1200 tnetres. April 5.
1937(Macintyre,).
The nearest ally is Atarba (Atarba) brunneicornis
Alexander, of Colombia, u'hich is readily told by thevenation ancl by the stmcture of the male hypopygium.
sp. n.
Atarba (Atarba) nod,u,losa,
Belongs to the cincticornisgroup ; general coloration
yellow ; antennre(male)elongate.
of prrescutumtestaeeous
subequalin length to trod.y.bicolourecl,the basesof' thesrvollen.producing
flagellarsegmentsblack, eonspicuously
a nodulose appearance; legs yellorv ; wings yellow,
with a restricted brou'n pattern : Sc, ending about
opposite one-third the length of' the relatively long Rs,
nx-cu closeto mid-length of the rectangularcell lst M 2;
male hypopygium with the redeagusangularly bent atnear two-thirds the length and here provided with aspinouspoint.
Male.-Length about 5'5 mm. ; wing 6 mm. ; anteunre
about 5'5 mm.
Rostrum brown ; palpi with basal segment yellow,
outer segmentsblack. Antennre(male)elongate,approximately as long as body ; scape,pedicel and first flagellar
segment pale yellow ; succeeding segrnents bicolored,yellow, the basal half black, conspicuously swolleu,
about twice as thick as the apical portion, with longconspicuousverticils. Head greyish brown ; eyes large ;anterior vertex greatly reduced, not exceedingone-halfthe diameter of the scape.
Mesonotal prescutum and scutum testaceousyellow,
the lateral margin narrowly darkened; posterior sclerites
of notum darker. Pleura testaceousyellow, the dorsopleural region more darkened. Halteres pale. Legs
with the cox& and trochanters yellorv : remainder oflegs vellow, the outer tarsal segtnentsrlarkened. Wings

208

Dr. C. P. Alex anderon

yellow, restrictedly patterned with brown, including
the stigma and a narrow vague seam along eord, the lattei
chiefly indicated by the darker veins ; veins yellow,
except as described. Venation : Sr, ending aboutgpposite one-third the length of .Bs, the latter relatively
Iong, nearly twiee nt-au ; cell lsf M z relatively large,
reetangular,with m-cu,at or eloseto mid-length.
Abdominal tergites chiefly dark brown, the sternites
more bicoloured, dark with paler caudal borders ; abdomen with conspieuous ereet pale setre; hypopygium
yellow. Male hypopygium with the caudal **gi"
ofster_nalplate only weakly emarginate. Outer disiistylemodera,tely rvide. the outer crest of spines extending
virtually to base of style, totalling about a dozenspines-,
the outer longer and more appressecl; apical spinesshort,stout, slightly decurved ; surfaee of style roughenedby other smaller scabrouspoints and spinulre. ,lDcleagusof moderate length and stoutness, about the clistal thlrcl
angularly bent ancl directecl lateracl, with a eonspicuous
spinous point at the angle.
Hab. Bcuador (Oriente).
Holotypa,6,Abitagua, altitude 1200metres, September
18. 1937(Macintyre).
Atarba (Atarba\ noilu,losais allied to A. (/.) cincticornis Alexander, of British Guiana ancl Ecuaclor. in thereversedpattern of the antennal flagellunr,u'ith tlre baseblack instead of vellou'. It is reaclilv tolcl frour the tu'ohitherto deseribeilmembersof'this gr"oopby the nodulose
antennre and by details of structure of the male hypopygium.
Trentepohli,a(I{eomongoma)suberecfA,sp. n.
Thoracic notum black. the pleura ancl pleurotergite
obscure yellow ; halteres blaekenecl, the base of sCem
ylllow ; legs dark brown, the tarsi a little paler ; wings
whitish subhyaline, the base and costal portion moie
yellowish, the outer radial field infumed ; stigma conspicuous, dark brorvn ; vein RB suberect ; petiole ofcell .Busubequal to rn.
Male.-Length about 8 mm. ; rving 7.b nrm.
Rostrum brown ; palpi small, black. Antennre black
throughout ; _ flagellar segments oval u'ith relatively
short verticils. Head grey ; anterior vertex reducerl

newor little-known,Tipulidre.

209

to a tiny line about, as wide as the diameter of a single
row of ommatidia.
Pronotum black. Mesonotalprrescut'umchiefly coveredby u black dorsal shield, the humeral and lateral portions
yellow ; scutal lobes black, the median area yellow ;scutellum and mediotergite black. Pleura and pleurotergite tight yellow, the ventral sternopleurite a trifle
infuscated. Halteres with baseof stem yellow, remainder
blackened. Legs with coxre and trochanters yellow ;femora dark brown, thb basesvery restrictedly brightened ;tibire brownish black ; tarsi paler brown or yellowish
brown. Wings whitish subhyaline, the basal and costal
portions a little more yellowish ; basal portion of cell Bcslightly darkened ; wing-tip in outer radial field more
infumed ; stigma conspicuous,ov&l, dark brown ; veinsdark brown, pale in the brightened portions. Venation :-8, suberect,.8, oblique ; petiole of cell .Busubequalto m ;rn-cu at fork of M ; cell Cu at' margin subequal to distal
sectionof Cur.
Abdominal tergites dark brown, the lateral borders
narrowly pale ; Iateral portions of intermediate sternites
rnore broadly yellow ; hypopygium black.
Hab. Ecuador (Oriente).
Holotypo,3, Abitagua, altitude 1200metres, September
20, 1937(Macintyre).
suberectais entirely distinct
Trentepohlia (I{ eomongonur)
from 7. (.nf.)ilisjuncta (Alexander), the only other speciesof the subgenushitherto described.