3 86

Entomolog, Mittoilungen XV, 1926, Nr. 5/6.

in the collection
Newor little-known
Tipulidaeof the(Dipt.).
Deutsches
Institut
Entomologisches
By Charle s P. Aiexander, Amherst(trfassachusetts).
(With 2 fig.)
Through the kindnessof Dr".\Valther Horn, I havebeenprivileged
to examine the undeterminedTipulidae in the collectionof the Deutsches
EntomologischesInsti',ut. Several of the speciesincluded.in this collection provetl to be of unusualinterest, In this paper, I am discussing
cei'tainTipuline forms fi'om llrazil and Chile, together with a few criticalnotes on tlro genera of Tiytul'inaedesclibetlby Euclerlein in 1912.
I wish to extend ny very sinceretlranks to Dr. Horn for the opportunity of examining this rich series of crane-flies.All rnaterial has beenreturneclto Dr. Horn.
Oaodicera (Ozod,i,cera)epiconna n. sp.
General coloration of the thorax buffy yellow, thc plaescutum with

Entomolog, Mittoilungon XV, 1926, Nr. 5/6.

56t

foul blown stripes that are confluetrtbehind; wings with a ltantlsottte
patlern, the costal regiou broatlly dark brown, the posteriormargiu paler
brown, the metlian portion cream-coloretl.
Male. - Length 23 mm; wing 18 mml antennaabout 6 mm'
Frontal prolongation of the head elongate, reildish brown, ldarker
laterally antl beneath, the nasus short ancl stout; palpi clark brown.
Antennao with the scapeancl pectinated segmentsbrownish yellow, thebranchesancl terrninal simple segmentsdarker; pectinationsa little longer
than the segmentsthat bear them, with tho exceptionof the last which
is a little shorterl simple flagellar segmentselongate, together about afourth longer than the combiuedpectinatesegments.Heacl brownl vertex
at its narrowest point only a trifle wider than the flrst scapalsegment.
Pronotum brown. Mesonotalpraescutumobscure buffy yellow, withfour brown stripes that aro confluentbehinil, the intermecliatepair separaterl from one another ouly on the anterior half; seutal lobes brown,
tho posterior Iateral thirtl more buffy; scutellum aud postnotum brown,
the color concealecl
by a heavy buffy-yetlow pollen. Pleura brown, thocolor about concealeclby a heavy whitish to yellowish pollen; tlorso'
pleural membraueclark brown, to proclucoa tlorsal longitudinal stripe'
Halteres tlark brown, tho base of the stem a littlo paler. Legs withthe coxae brown, sparsely pruinoso; trochanters ilarker bqownl femora
brownish yellow, the bases brighter, immediately before the tips with avague clarker ring; tibiae brown, the tips narrowly tlarkenetl; tarsi
brownish black. Wings hanclsomelypatternecl,the costal margin being
the posteriorthirtl
broadly ilark brown, the rretlial portion cream-colorecl,
the prearcularborcler
incluclesdarkanterior
more greyish brownl the
B and' M; a
of
cells
bases
broaclcells; cells C, Sc, Scr, 2nd, R' theyery
the anterior
on
seam
broad
large oval area at origin of lls, antl apaleon
the basalsnffusion
corcl, cell lst R, being tlarkeneclexcept for a
of
cells
ctt,,
thirtl; the cream-coloreilcenter includes the broacl bases
lst A and 2nd A; the portions of cells I? and M not' clarkenetl,asabove, and an extensivearea beyonfl the corcl, inclutling mostcliscussealof cells lst M, anrl Q antl the ailjoining portions of cells -I?, anil theextreme base of cell Mr; posterior cortl anil outer end of cell 1st M,
broadly seamedwith darker brown; tho dark posterior borcler inclndes
all of tlre remaitring cells of the wing, colls 2nd, Mz, Ms anil Mn being
uniformly infumed, t[e infuscation being connecteclwith the anteriorclarkenetlregiou around the wing-tip; veins brown, clark brown in thetlarkest are&s,luore yollowisf brown i1 the creamy portions. Veuation:
Cell M, broadly sessile,the proximal end being longer than r-zl.
Abtlomenreddish browu, the lateral margins of the tergites tlarker
brown, becorrringbroatler behinl; distal sternites antl the hypopygium

3 88 A lex ande r,N c w o r l i ttl ek n o w n T i p u l i d ao
i rr the col l .oftheD .rl rrt.r,st.

Entomolog.
Mitteilungerr
XV, 1926,Nr. 5/6.

rtarker blolvu. llalc hypopygiumsnrall. Ninth tergite leiatively shor.t,
the cauda,lnrargiu with a narro\\r v-shaped urediirrrnotch, the laterallobes thus iirrnreil broadly obtnge to stibtruncate. The combinedninth
stelnito aud basistyleelongate. Caudaimalgin of the dististyle produceil
into a sleniler',geltly cnrveil, r'eddishhook, the tip of wtrich is weakly
anil unequallybilid.
Hab. Brazil (Espir.itoSanto).
Hol, ot y pe,d , Po rto d o C a d i , IIa rc h 1 g2l (F. H offman); A rp.
d eilic .1921.
ozod,iceraepicosm&is u'ell-distiriguishedby thc body-colorationancl
the handsomely
patternedwings. The only altied speciesso far. clescribeil
is O. b,imaatlata Etde,-lein, likewise frorn Espirito Santo, in whichtlre tholax is bright rust-yellowantl the highly contrasteilwing-patternis less conspicuonslyilevelopecl.
Holoru,s,iaocellata (Onderlein).
1912. Icri,omastarocellataEnilerlein; Zool. Jahrb., SJ,st.,32: 9 - 10.
7917. fcr,iunastar ocellataEntlerleinl ZooL Anzeig.,,1g161- 62; tig. 6.
(wing), flg. ? (palpus).
A femalein the DeutschesEntomolog.Instit. collectionfrom Joinville, Santa Catharina,Brazil, August-October, 1g22, taken at electric
light' Entierleiu's type was from Espirito santo. From the present
specimen,it can be clenonstratedthat the gennsrcr,iomasta,riE invalid
arrrl must be placed.in the synonynryof Holorus,iu. The spurs of crossveins in cells-R and lsf ,4, as shorvnin Enderlein's type anil offering
priniary cha,ractersfor the separationof the group from ailied ripulineg€ilera, are quite iackiDg in the present specimen,anil tlreso must be
consialeredas being ailventitiousrather thau superlu,mealy. other species of Holorusia have the maxillary palpus with the terminal segm€nt
relatively short, anil thig cliaractermust be heLilas boing of very doubtful
volue. r cannot clearly detect the bifid nature of the nasusin the present specimen. As iudicateil by Edwards for the genus Macrontusti,r(Trans. N. Z. Inst., b4: 333; 1gZ3), the uature of the nasus varies
greatly even within closelyallied groups of species.
The writer's oarlier opinions(proc. U. S. Nat. Ifus., 44: 4g6-lg?;
1 913) of E nder l e i n ' s T i p u l i n e g e n e ram a y n ow be somew hatmodi fi ei l
as a result of several yoars experiencewith the group, during which
period represeutativesof all of these genela aucl supposedgeriera havebeen examined.
ctcnacroscelisEudorlein is l'ery close to Hol,orusi,aLoew. anil las
more lcceutly been placedin the synonymyof the lalter by Ellilerieiu
hirnreif (Zool. Anzeig.,4g: 6o; 1g1?). rlolvever,bo{h Eilu'arclsanti thewritel slill feel inclirredto recognizethe gronp as valid, although real-izirrg tlre extrenteclose tffinitics n'ith llolorrtsi.u,, ,\s so lestrictod, thegentrs incluiles nlltllel'0ttsPtlireottopictl species.I[olorusict,is essentilll.y
Neotlopical,rvith olly the gerrot;'pe,rtibigirtosaLoer,v,occurliug in tho
Nearctic region, aud a suligenusof d.oubifnl position (ZclandotQtula
Alexander)in Nerv Zealancl.
Icrionusta,r Itrlderlein must be placerlirr tlio synouyny of llol,orusia,,
as i udi cat edabove. Eilwar t ls ( Tr aus.N. Z. I nst . , b4: 333; t g2B) hailsuspecterlthat this geuus, as well as PhymatoltsisSlirse trnrl Pt:,h,|/tcq,
Enderleirr,uright prove to be syrronymsof Mncromctstirosten sacken.
Phytrtatopsis
is uow utuch better'huowrriu ths adult couclitiotr,
aud theiurnature stagesof the genotypehirve recerrtlybeol iliseovereil.It hasbeel fouuil tlrat memhersof the geuus have stbaptelous fernales u'ith
long, chitinizeclvalves to the ovipositor,a charactelthat at ouceexcludes
tlre gronp from Macromastir, The true affinities c$ Pehl/tca al'e oolrsiderealbelow.
Stegus'tnonotus
Encler'leinatl Pekl/rcaEndellein, are, in the opiniou,
of tlre writer, Bynoxlrns of Tanypramzo Osten Sacken. 'Iire reasonsfor sur:han assignrrentiravebeeugiveu in an ealliel paper(Journ.N. y.
E rrt. S oc. ,22: 206; 191a) .
Malpigh,ia Enderlein is very doubtfully distinct frotln Ctenoythora
1\[eigen.
PltacelodocenrEnderlein shoulclbe maintaineclas ilistinct, at least
until tlre riumerous species of tlie Australian genus Plaszontyi,askttse
are better kuown. The genera Ptilogyna \\restwooil,Plu,s,iomy,ia
S\<nse,PlatyphasictSkuse, and tho present gtoup are all verl' closely relateil
to olle another,Jifferingcliefly in various moililicationsof the anteulae.
DiheraclorutsEnilerleiu is considereilby the writer as constitutilq
a valid subgenusof OzodiccrctMacquart.
Tiyttilodi'naEnderlei', placed by its author i' trre Limonii'e pediciitti, is now represerrled
by nulnerousEthiopian and orieltal spe.ics,
a few of the latter occulring fnlther northr,r'arcl
in tho south.easter.nPalaearcticregiol. Although the structure of the adult ollers vely few
satisfactory charactersfor separating the gloup fron Ti,1tu,laLinuaeus,
the prrpa showsfeaturesthat are mnch more like those of Prianocera
I:oew. Tito grotp may be held as valid unless more cvidetce to the'coutrary is forthconring.
AlcJrouandi'aEntleriein, placed by its autho. i' the Limoniine
Eriopteriui, is certainly a valiil genus. 'rhe venationsuggestsaffinities
witlr- oaorlicu'o }lacqnart, but the rvholelrabitus of the fly points strolgly
towarrl'Ianypremrta Osten Sacken.
Ilolontsi,tt, lrcrti n. sp.
Auterior half of mesonoturn
cla,r'kbrolvn, the posteriolhalf ab.tptly

3gg

3g0

Alexancler, New or littlc-knorvnTiIulirlao in tho coll. of the D. Ent. Inst.

r l i i l 'k n i t l r t h e l l i c e s o f t l l e
Irule ; ullte llllne bic r . r lor uus t, Le f lr lgt ' llt t lr r
individual segments nan.owly pale ; plerlr'a yellou" the clorsal portioll of
the pleurotergite dark; wiDgs with tlre costal rcgion narlowly darkthcbrown, follolvecl by a bload subhyaline area,, tltrl posterior half ofp:r]':t'wiDg ilfuscated; cell .r11,sessileI abdominal tergi'-es ilark brolvn,
medially.
Female. -Lerrgth about 21 mm; wing 23 rttm'

Frontal prolongatiou of the head light brown, tlte nasus elotlgate,tlre
clark brown, tnfted. wilh yellolv selae; palpi much longer thanantenllae, ilark brown, the terminal segnent passing into obsoule ofallge.
Anteunae with the scapal segrnents obscnre brownish yellow, the apex
theof the tirst segment clarker; flagellar segmelits black, the apices ofsegment
flageilar
lirst
the
of
base
pale;
extremesegments narrowly
restriotedly paler; terrninal three antenlal segments more uniformly
clarkeneil. Head lich ftlvous brown, with a capillary ilatk brown vitta
theand a paler brown cloud on either side of the meclian line ofpostetiol' vertex.
Pronotum brown. Mesonotal praescutum almost uniforlnly opaquebrown, the usual iuterspaces sc&rcely of a different color, with a sparseyellow;clothing ofyellow setae; huneral region very lestrictetlly obscureposteriortheblown,
dark
half
anteliorthescutum abruptly dimidiate,half buffy-yellow; scutellum pale browr , indistinctly variegateil witlt
paler on either side of a capillary dark vitta; anterior ritlge of theparascutella dark brown; postnotal rneiliotelgite light blorvn, paling into
Pleurawhito on the cauclal portion, the cephalic lateral corners clusky.
postnotal
the
of
ha]f
yellow, the clorso-pleural regiorr darkerl dorsal
pleurotergite tlark brown, confluent with the abdominal stripes. Haltereswithclark brown, the extreme base of the stem restrictetlly pale. Legs
femorathe coxae anil t|oclranters yeilow; legs very lolg a[tl slencler;
and tibiae brorvuish yellolv, the tips nart'owly infuscatecl; tarsi obscu|ebrownish ycllorv, passing iuto clarker at tips I basitarsi much longer tharr
the tibiae. WiDgs with the costal margin narrolvly dark brown, followedofby a broad whitisli subhyaline longitntlilal stripe, tho posterior balf
cells
all
of
iuclucles
margincostaldark
thedusky;
the wing entirely
C, Sc, ,Sc' the stigma in cells 1 st R, and 2 nd fir, antl the extrents
cephalic nrargin of cell 1l; a brown spot at extremo base of cell M;
a small clearly-cleliDeil brolvn spot on vein' Cu near mitllength of cell M;
A,tlre pale stlipe includes the narrow bases of cells Cu,, lst A and,2nil,
R,
2ncl
celis
of
po'tions
alniost all oi cells B a'd M, atd adjoini.g
llu1
before tho stig-ma, crossing thc co|tl into cells Ra, Rs aDil base ofthD
brou'tt,dalk
Oeulel of cell .R, arrd apex of E, dzrrkenedl vtins

MittcilungonXY, 1926,Nr. 5i6.
li)rrl,ornolog.

391

near
oblittrliLtivrraleas extensive. Vetra,tiou:(1r'll/i:r vely cctttstlictecl
rrridlergtlr; tt:ll Jl[, narrowly sessile.
Abtlorrrinal tergites broadly dark blown sublatelally, the basal
seqrnentsbroadly pale medially, this color becomingnarrower behinilI
yellotv.
ltteral rnalgius ofthe tergitcs larrowly pale; sternitestestacoous
Ovipositorwith the valves pale, the tergal valves very cotnptessetl.
Hab. Brazil (Sao Paulo).
Holotylte, Q, Butantal, August 9, 1920, (R. Fischer).
This very iuterestilg crane-fly is nameclin honor of my friencl,
D r. W al ther Hor n, t o whom I am gr eat ly indebt edf or m any kint lnessesin the past. Holorws'ictltorni' is reaclily distinguished by thestrihing body- and wing-coloration,the bicolorous antenuae anil thesessilocell lZr.
Ilolortrsia hontot:)trou,n. sp.
Generalcoloratiol yellorvishto fnlvous; mesonotalpraesctltumwith
four brolvn stlipes, the broatler lateral stripes darker'1 wings almost
with light brolvn, the stigmatlarker; malehypopygium
uniformly suffusecl
with the region of the ninth sternite proclucecistrongly ventrail, thenarrorv apices tufted with long yellow setae.
Ilale, - frength about 22 ntm.I wing 25 mm.
X'rontal prolongation of the head brownislt fulvous, the nasus
elongate; palpi black, the last segnert a trifle longer than segments
tlo aud tlrree taken together. A[tennal scape dark reddish browt.tI
flagellum brokel. Heail fulvous, witir a lralrow anti rolatively indistittct
mecliauline posteriorly; veltical tubercle low.
Pronotum light bror,vn,the latcral margins ntrrl'owly litred withfonr brown stripes,the ilternediate pa,irpaler brou'n. relatively narro\\'t
separatedb1' a lirre of the gt outrrl-cololther,tis about ole-thircl fls
huntet':rl
wiilc as the stripe; later.nlstr.ipesdalker btorvn, coltspir:uousl
region alil t|e sublatr:ralpoltions of the plaescltunt vaguely darkened;
scntum obsclrreyellow, eacli lobe largely covereclb5' a slightly oblique
brolvn marking, the mealianline at the suture less ilistincily darkeuedI
scltellum broacl,yellow; postnotumsimilar, a trifle ciarkeneilposteriorly.
Pleura light browuish fulvous, the dorso-pleuralmembraneclearer ili
colol'. Halteres iight brown, the hnobs darker. Legs with the coxae
brownish fulvons; trochanterslulvousl femora reddishbrown, the tips
scarceiydarkened;tibiae light brolvn, the tips very narrowly infuscatefll
tarsi brown, the outer segntetltspassing into black. Wings almost
lnifornily snffusefl with light brown, the siigma darker brown; veins
somelvhatmore ilistinctly seameclwith brown, Ieaving the centers of thecells a trifle paler; veins pale, the radial vein between tho wing'root
and arcnlus incrassateil and blackenetl. Venation: Vein 11, stronglyu32

Entomolog. Mittellungen XV, 1926, Nr. 6/6.

afcuatetll at ln th€ genrts; cell 4 relatively narrrw rt its inner ond,strongly uarroweclbefore ancl at miil-length, the outer end wldely expahal€d,the cell at the wlng.margin hs Wlde &i5 or wider then, cell fir; petioleof cell M, rubtqual to or shorter than ri,
Abdonen rtildish brown, brighter fnlvous basally, the eighth stertrite
darker at bane. Male hypopygirtm (Flg. l) rnassive, the I th tergite (t)producerlmedially into a lobe (Fig. 2) thet splitr into tWo tllvergent
flatteneil blades, the etrrface auil mernl tnargiur of which aft pfovitletl
with short cette. Combined I th sternite anil baristyle extenrive, theforilef region (s) proilnced itrongly ventrad, narroweel towartl apex ehd
h0re tuftetl with long yellow setae. Yiewod cauilally, the rt'ernite isieon to have a ileep V.shapetl notch. Region of the basistyle (b) witlt
relatively long anil conspicuousyellow setae. Outer distlttyle (il) strort
clavate.
fJrb. Chtle (Malleco).
Holotgpe, dr Victoriq (von B od e rn oyc r)-

Fig. l.

Eolorusia hom:orhroan. sp.
Ifele hypopygium; lateral e8p€ct.b : regiod of the basistyle;tergito.
a l: out €r dis t is t y le; s : gt hs t er n i i e ; t : 9 t hFig. 2. Male hypopygium. Dorsal asirectof 9th tergite.