45L
Reprinted from ANNels oF THE ENrouot ocIcel SoctsrY on Aunntce,
Vol. XXXI, No.3, September,1938.

STIIDIES ON THE CRANE-FLIES OF MEXICOPART VI(Order Diptera, Superfamily Tipuloidea)
Cn.q,nI.rsP. AlnxlNopn
Amherst, Massachusetts

The presentreport is basedentirely on someextensiveseriesof Tipulidae collectedin the State of Chiapasby Dr. Alfons M.
Dampf, chiefly in 1935. Dr. Dampf writes as follows concerning"The material is the result of a twothis very rich collection:months collectingin the virgin forestsof the SierraMadre during
the year 1935. I arrived at the coffeeplantation Vergelwith thefirst rains of the wet seasonand had the opportunity to witness
the suddenchangeof the fauna after the dry time. The country
is characteized by deep woodland valleys, sometimesin theform of gorgesand by mountain rangescovered with remarkable virgin forest. The collectionsafford a good opportunity
to study the distribution of the speciesaccordingto altitude."Certain of the Mexican Fauna (NI. F.) numbers of Dr.
Dampf used in the presentreport are summaized herewith.
Finca Vergel,:Situated on the right steepslopeof the Valleyof the Huixtla River, in coffeeplantations that have changed
the virgin forest, making it much more clear. The vegetationisvery rich and varied. Besidesa permanent brook, there are in
the rainy seasonmany small rivulets. Altitude 800 meters.
Many numbers from M. F. 4123 (May 11) to 4714 (July 2).
Additional collectionsweremade at stationshigher up the slope,
altitude 1000 meters, overlooking the woodclad slopesof the
Huixtla Valley; M.F.4191, and still higheron the crestof thewatershedbetween Rio Huixtla and Rio Despoblado,altitude
1200meters;M. F. 1792.
F'inca Victorio: The region is much more virgin than Vergel,full of brooks, rivulets, barrancasand the like-a paradiseforthe entomologist. Altitude about 900 meters. M. F. Nos. 4342,
4693,4694, 4709,1710.
Finca Aurora: Trap light coveringa valley with brooks androcks, densevegetation. Altitude 200 meters.M. F.4552.
lContribution from the Department of Entomology, Massachusetts State
College.
The preceding part under this general title appeared in these Annals (Vol. 21,
101-119,1928).

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Annals EntomologicalSocietyof America [Vol. XXXI,Finca German'ia: Trap lamp, on veranda overlooking valleys,
rocks and many brooks. Altitude 1190 meters. M. F. 457L.
The great majority of all the specimens secured were taken
at trap lights set in a variety of locations throughout the region.
In the present paper a portion of the tribe Eriopterini is considered. Through the continued friendly interest of Dr. Dampf,
I have been permitted to retain the types in my collection but
duplicates and paratypes of many of the species have beenreturned to Dr. Dampf for the Government Collection. As
before, I express my very deepest thanks to the collector forthe privilege of examining this remarkable series of Mexican
Tipulidae.
Gonomyia Meigen
Gonomyia (Progonomyia) atroapicata sp. n.
General coloration yellow; mesonotal praescutum and scutum
conspicuouslypatterned with polished black; antennaeblack, the scapeand pedicel yellow; femora yellow, the tips conspicuouslyblack; wings
with a weak brownish yellow tinge, the stigma slightly indicated; Sc
long; petiole of cell -R3long, nearly three times vein R3alone; cell ?nd M
subequal to its petiole; abdominal tergites bicolored, the basal half
darkened; male hypopygium with basistyle unarmed at apex; innerdististyle a flattened blade.
MaIe. Length about 4 mm.; wing, 4.6 mm.
Rostrum and palpi brown. Antennae with scape and pedicel lightyellow, flagellum black; flagellar segments long-oval, the verticils
exceedingthe segmentsin length. Head light yellow.
Pronotum and pretergites sulphur-yellow. Mesonotal praescutum
obscureyellow, conspicuouslypatterned with three polishedblack areas,
including two oval lateral markings and a similar one on antero-median
portion of sclerite; remainder of median praescutal stripe much paler;scutal lobes polished black, the median area pale; scutellum blackened,
the posterior border yellow, parascutella yellow; mediotergite chiefly
brownish black, the antero-lateral portions broadly yellow. Pleura
chiefly yellow. Halteres yellow. Legs with the coxae and trochanters
yellow; femora yellow, the tips conspicuouslyand abruptly black, theamount subequal on all legs; tibiae yellow, the tips narrowly darkbrown; tarsi brownish yellow, the terminal two segments darker.
Wings with a weak brownish yellow tinge, the small stigma slightly
indicated, pale brown; veins brown, the prearcular area and veins Cand ,Scyellow. Venation: Sc long, Scr ending about opposite four-fifths
the length of Rs, Sc2some distance from its tip, about opposite twofifths the length of R-S;petiole of cell R3long, nearly three times vein R3:alone; eell ?nd M2sabequalto its petiole; rn-cuat fork of.M.
Abdominal tergites bicolored, dark brown, the caudal half of thesegmentsyellow; sternites more uniformly yellow; hypopygium yellow.

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Alexander: Crane Flies of Mexico

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Male hypopygiumwith the basistylenot producedinto an ap.19a1_spine,
blade, theas in biiasciotata. Outer dististyle a palg, lo:rg-rectln-9119r
outerflattened.blade,_t_heapex truncate. Inner dististyle a broadly
angle producedinto a curved spine, the inner_angle into-a blunt lobetraiing a microscopicallysermlated margin, the two anglesseparated
by a roundednotch.
Holotype, d, Huixtla, Chiapas, altitude 30 meters, November 21, 1930 (A. M. Dampf); M. F. 1944.
Gonomyia (Progonomyia) atroapicata is most generally similar to G. (P.) b,ifasc'iolotaAlexander (Cuba; southern Mexico to
Panama), both specieshaving the same peculiar pattern of themesonotum. The present fly is readily told by the blackened
femoral tips, unbanded wings, long petiole of cell Rs and,especially, the structure of the male hypopygium. The distinctions between the three allied subgenera of GonomyiaProgonomyia Alexand er, Ell'ipteroidesBecker and Protogonomy'ia
Alexander-have been given in a separate paper by the writer(Philippine Jour. Sci. 62 : 177-178; 1937).
Gonomyia(Euptilostena)dampfianasp.n.
General coloration gray, the praescutumwith four brown stripes;
basal flagellar segmenti bicolored; femora yellow, the tips brownish
black; wittgs grayish subhyaline,patterned with brown;_.outerradial
cells infumidf enclosingpale margin-a!dropletsi Scr ending opposite
midlength of Rs, Sc2before origin of latter; supernumerary_crossveinin cell ht at near three-fourthsthe length of cell; m-cumore than twice
its length beforef.ork of.M.
Male. Length about 6 mm.; wing, 6 mm.
Female.Length about 7 mm.; wing, 6 mm.
Rostrum and palpi black. Antennaewith scapeand pediceldark;basal flagellarsegmentsbicolored,dark basally,_thedistal endsyellow;
outer flagellar segments-Head uniformly darkened; flagellar-segm.entslongabove graY,narrowly darkenedmedially; sides
oval to fisiform.
of posteriorvertex weakly infumed.
Pronotum gray, lined on sideswith yellow. Mesonotalpraescutum
light tray, wit*h fbur narrow brown stripes,the intermediatepair narrowed before the level of the transverseblack pseudosuturalfoveae,
wider behind; scutum #aY, the median line narrowly darkened,eachscutal lobe with two brown areas;scutellum and mediotergitebrown,with a narrow paler medianvitta. Pleura grayt with three ver.ynarrow
black longitudinal stripes, the most dorsal crossingthe anepisternum,
the most ventral on tlie sternopleuriteand beneaththe halteres. Haltereswith stem yellow, knob black. Legswith the cox?epale,narrowly
darkened at bas-e,mosi evidently so on fore legs; trochanters brownish
yellow; femora yellow, the tipi rather broadly brownish black, ttt"
i-ont i subequal on all legsf tibiae yellow, the tips .very nagoruly
d.arkened;tarii black, the ploximal portion of basitarsi paler. Wings

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Annals Entomological Society of Arnerica[Vol. XXXI,

grayish subhyaline,the prearcular and costal portions more luteous; aconspicuousbrown pattern, arranged as follows: Spots at arculus;
origin of Rs and Sc2;Scr;forks of l?s and M4z j supernumerarycrossveinin cell Rainl-cu; stigma dark brown; paler brown washesin outer ends
of cells Rz to ?nd' Ra,inclusive, these enclosingpale marginal droplets;veins pale in the luteous areas,darker outwardly. Venation: Scr ending
opposite midlength of Rr, Scza short distance before origin of fts;
petiole of cell R3long. a little shorter than .Rs;distanceon costabetween
veins R112and fts a little exceedingthe length of the latter vein; supernumerary crossveinin cell Rn at near three-fourths the length of cell;
rn-cumore than twice its length before f.ork of.M.
Abdomen dark brown. the caudal borders of the intermediate segments very narrowly pale; hypopygium dark. Male hypopygium withthe ventral lobe of basistyleproducedcaudad into a long slenderfleshy
appendage.Both dististylesfrom the baseof this lobe,the outer with along slenderstem, at apex unequally bifid; inner dististyle subequalin
length, a simple,flattened,yellow blade, the distal third narrowedintoa slenderblack spine. Cerci darkenedbasally,the outer endspaler.
Holotype, ?, Vergel, Chiapos, altitude 800 meters, at light,
June 12, 1935 (A. M. Dampf) ; M. F. 4521. Allotype, d, F'inca
Germania, altitude 1190 meters, June 20, 1935 (A. N{. Dampf);

M. F. 157r.

Gonomyia(Euptilostena)dampfiana is named in honor of thecollector,Dr. Alfons N,{.Dampf, to whom I am greatly indebted
for many appreciated favors in the past. The present fly
requires no comparison with any described species since thethree known members of the subgenusare all restricted to the
Palaearcticand Oriental Regions. The discovery of Euptilostena
in the New World is thus of considerableinterest. The structure of the male hypopygium is more generalized than that ofspeciesof the subgenusld,ioceraDaTe(PtilostenaBergroth) andsomewhat curiously resemblesthat of certain Australian speciesof the genus Moloph'ilus belonging to the ruJicoll,issubgroup of
the grac'il'isgroup.
Gonomyia(Lipophteps)
aequidenssp. n.
Belongsto the cinereagroup; wings with cell Mz open; male hypopygium with the outer dististyle present,appearingas a slenderpale
spine; middle dististyle darkened, the surface squamose,near basebearing a lobe that forks into two approximately equal blackened
spines;inner dististyle fleshy, pale.
Male. Length about 3.5-4 mm.; wing, 4-4.5 mm.
Female.Length about 5.5 mm.; wing, 6 mm.
Rostrum and palpi black. Antennae with scapeand pedicel yellow
above, dark beneath; flagellum yellow. Head pale yellow, the center ofvertex darkened.

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Alexander: Crane Flies oJ Mexoco

397

Pronotum yellow, narrowly darkened medially; lateral pretergites
pale yellow. Mesonotal praescutum gray, with four slightly darker
brownish gray stripes; pseudosutural foveae pale brown; scutal lobes
dark brown, the median areapaler Fay,with a narrow dark median vitta;
scutellum dark basally, obscureyellow on margin; mediotergite brown,
pruinose. Pleura narrowly lined longitudinally with brown and silvery,
including further a broader yellowish white ventral stripe extending
from the fore coxae to base of abdomen; sternopleurite {ay, bordereddorsally by brown. Halteres with stem pale yellow, knob dark brown.
Legs with fore coxae dark, variegated with white; remaining coxaeand
trochanters yellow; fore femora black; mid-femora yellow, the tips
(about distal fifth) black; posterior femora yellow, with a scarcely
indicated, narrow, brown, subterminal ring; tibiae snowy-\Mhite,thenarrow basesand slightly wider tips of fore and mid-tibiae blackened,
the posterior pair with the ends still more narrowly infumed; tarsi
black. Wings with a gray tinge, the costal border white, more extensivein the radial field before and beyond the oval, pale brown stigma; veinsbrown. Venation: Scr ending a distance before origin of the short .Rs
greater than the length of the latter; rn-cu close to fork of. M; cell Mz
open.
Abdominal tergites dark brown, the caudal borders of the segments
narrowly pale; hypopygium yellow. Male hypopygium with the outerdististyle a slenderpale spine,shorter than the middle dististyle. Middle
dististyle complex, the outer portion dark, sinuous, the outer surface
with microscopic spinous points to produce a squamoseappearance;
beyond midlength with a small darkened lobe; near base with a conspicuouslobe that forks at apex into two nearly equal black spines,one
subglabrous,the other with microscopicappressedspines. Inner dististyle an entirely pale fleshy lobe.

IIolotype, d, Vergel, Chiapas, altitude li00 meters, IIay 11,
1935 (A. M. Dampf) ; M. F. 4126. Allotopoty/e, I , ParatopotyPe, d &; paratypes, d &, \rergel, altitude 1000 meters, \[ay1 9 , 1 9 3 5 ;N { . F . 4 2 0 5 .
Gonomyia (Lipophleps) cequidensis very distinct from theother members of the group in the structure of the male hypopygium. In the unarmed inner dististyle and retention of theouter dististyle, it is closest to G. (L.) heloph'ilaAlexander but
the middle dististvle is entirelv different.
Gonomyia(Lipophleps)haploidessp. n.
Belongsto the rnancagroup; rostrum obscureyellow; antennae
black throughout; pleura heavily striped, black, with a broad silvery
band; knobsof halteresobscureyellow; Screndingsomedistancebefore
origin of Rs; male hypopygium with the dististyle simple, fleshy;
phallosomewith no spinous points other than the slender straightaedeagus.
MaIe. Length about 3.4-3.6mm.; wing, 4-4.2 mm.

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Annals Entomolog,ical Society of America[Vol. XXXI,

Rostrum obscure yellow; palpi black. Antennae black throughout;basal segmentenlarged;verticils very long. Head gray; center of vertex
a little darkened; eyes large.
Pronotum above bright yellow, darker on sides; pretergites similarly
yellow. Mesonotal praescutum dark brownish $ay; pseudosutural
foveae black; scutal lobes brownish Eray, the median area clear lightyellow; scutellum broadly obscure yellow, the central portion at basedarkened, parascutella dark; mediotergite dark, the lateral portions
obscure yellow; pleurotergite chiefly obscure yellow, bordered beneathby black. Pleura heavily striped, black, with a broad silvery band
extending from behind the fore coxae to base of abdomen; dorsopleuralregion restrictedly obscure yellow. Halteres dusky, the knobs obscureyellow. Legs with coxae and trochanters medium brown; remainder oflegs black. Wings broad, with a brownish tinge; stigma scarcely differentiated; veins brown. Venation: Sc short, Sc1 ending a distance
before origin of Rs nearly equal to length of latter; branches of Rs
strongly divergent; m-cu close to fork of.M.
Abdomen dark brown, the lateral and posterior borders of the segments narrowly paler; hypopygium brownish yellow. Male hypopygiumwith the dististyle simple, ffeshy, unarmed except for two fasciculate
setaeat apex. Phallosomeconsisting of a quadrate basal structure, theaedeagus produced from this point to beyond the level of the styli;broad at base, at near midlength abruptly narrowed into a straight
slender point, the apex further narrowed into a hair-like spine.

Ilolotype, J, Finca Victoria, Chiapas,altitude 900 meters,d 9,June 29, L935(A. M. Dampf) ; N4.F. 4709. Paratopotypes,
paratypes,& &, Vergel,
June 28-29, 1935;M. F'. 4(i93A, 17t0;
altitude 800 meters May 13, 1935,M. F..1153;May 19, 1935,
M. F. 4205;June 1, 1935,M. F. 438-1.
Gonomyia (Lipophleps) haploidesis most nearly allied toG. (L.) haploa Alexander (Mexico), from which it differs
especially in the heavily striped thoracic pleura and in the
structure of the male hypopygium, notably of the phallosome
which has no spinouspoints other than the slenderaedeagus.
Gonomyia(Gonomyia)triaculeatasp. n.
group; generalcolorationof mesonotum
Belongsto the subcinereo
yellow;antennae
dark brown,the pleurapale;rostrumobscurebrownishblack throughout;legsblack; Scr endingabout oppositeorigin of ^Rs,
rn-cu about two-thirds its length beyond
the latter subequal to Rz+s+ai
the fork of. M; male hypopygium with the aedeagusbearing three longpowerful spines, two near base, one subapical.
Male. Length about 3 mm.; wing, 3.6 mm.
Rostrum obscure yellow; palpi brownish black. Antennae brownish
black throughout. Head dark gray.
Mesonotum dark brown, the median region of scutum restrictedly
paler; posterior sclerites of notum hidden by the mounting medium.

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Alexander: Crane Flies of Mexico

399

Pleura testaceousyellow, unmarked. Halteres dusky. Legs with thecoxae and trochanters yellow; remainder of legs black. Wings subhyaline; stigma faintly darkened;veins dark brown. Costal fringe relatively long and conspicuous.Venation: Scr ending opposite origin ofRs, Sc2at its tip; Rs subequalto R213sand shorter than Raalone;vein
Rs oblique; rn-cuabout two-thirds its length beyond the f.ork of.M .
Abdomen dark brown; hypopygium paler. Male hypopygium withthe outer lobe of basistyleof moderatelength, extendingcaudad about
to level of outer dististyle. Outer dististyle a simple, strongly curved
hook, the tip acute. fnner dististyle broad, the outer margin with anacute appressedblack spine,the apex of style with two fasciculatesetae.
Aedeaguswith unusual armature, consistingof three powerful spines,
additional to the shorter curved apical one; most basal spine slender,
gently curved, about one-third the length of aedeagus,directed laterad;secondspine arising close to first, directed caudad, long and slender,
only a little shorter than the aedeagusitself; third spine subapical,
directed cephalad, about two-thirds the length of aedeagus,the tip
blackened.
Holotype, & , Finca Victoria, Chiapas, altitude 900 meters,
June 29, 1935 (A. M. Dampf) ; M. F. 4710 A.
Gonomyia (Gonomyia) triaculeata is readily told from all
described species of the genus by the structure of the malehypopygium, especially of the aedeagus.
Gonomyia (Gonomyia) microserrata n. sp.
Belongs to the subcineredgroup; mesonotumdark brownish gray,scutellum conspicuouslyyellow; rostrum and antennae black; pleuraslightly variegated with brown and yellow; halteres darkened; legs
black; wings with a faint brown tingei Screndingoppositeorigin of .Rs,
the latter about two-thirds as long ?,SRz+a+a;,n-cumore than one-half
its length beyond fork of M; male hypopygium rvith the aedeaguslong,near apex bearing a blackened double crest of microscopicretrorse
spinules;apophysessymmetrical,each bearing a long appressedspine
on mesaledgenear base.
Male. Length about 5-5.2 mm.; wing, 5.7-6 mm.
Femole.Length about 6 mm.; wing, 6 mm.
Rostrum and palpi black. Antennae black throughout. Head darkgray.
Pronotum yellowish brown; anterior lateral pretergites yellow.
Mesonotal praescutum and scutum almost uniformly dark brownish
ffay, the humeral region not brightened; pseudosuturalfoveae black;scutellum yellow, restrictedly darkened at base, parascutella dark;
mediotergite dark, more or less pruinose.the cephalic lateral portions
yellow. Pleura slightly variegatedbrown and yellow, the pleurotergite
chiefly pale. Halteresdarkened,the baseof stem pale. Legs with thecoxae and trochanters yellowish testaceous;remainder of legs black.
Wings with a faint brown tinge, the diffuse stigma darker; veins darkbrown. Costal fringe short. Venation: Scr ending oppositeorigin of400

Annals Entomological Society of Arnerica [Vol. XXXI,

Rs, ,Sc2near its tip; Rs relatively short, about two-thirds -R2.'qs,thelatter subequalto or somewhatshorter than Ra alone; rn-cumore than
one-half its length beyond the fork of.M.
Abdominal tergites dark brown, sternites obscurebrownish yellow;
hypopygium yellow, conspicuous. Male hypopygium with the outer
lobe of basistylepale, shorter than the dististyle, at outer apical angle
produced into a short spinuous point; lobe clothed with long coarse
setae. Dististyle with two long fasciculatesetae,near apex further producedinto a powerful blackenedcurved spine,this latter at bend with asingle strong seta. Phallosomecomplex,the long aedeagusexpanded
near outer end and provided with a blackened flange that bears adouble crest of microscopieretrorse spinules; apophysessymmetrical,
eachconsistingof a pale bladethat is expanded
subtendingthe aedeagus,
and twisted at apex, on mesal edge near base bearing a long slenderappressedpale spine that is only a little shorter than the apophysis
itself, the slendertip acute and blackened.
Holotype, & , Finca Germania, Chiapas, altitude 1190
meters, June 20, 1935 (A. M. Dampf); M. F. 4571. Allotopot y P e , g . P a r a t o p o t y p e s1, 2 & 9 .
This large species is readily told from all approximately
similar regional forms by the very distinct hypopygial characters, especially of the phallosome.
Gonomyia(Gonomyla)quaesitasp. n.
group; generalcolorationof notum brown,
Belongsto the subcinereo
unmarked;
the praesiutum darker medially; pleura testaceous-yellow,
kno6s of halteresdusky; legsand antennaeblack; wings subhyaline;Sc
more than one-half
n -cv,cshort, Rs about three-fifthsthe length of l?2-.31a,withthree dististyles,
hypopygiumM;
male
its lengthbeyondthe fork of
the outer pioduced into a long recurved spine, on outer margin nearbase with a slightly smaller similar appressedspine; phallosomeconsistingof paired straight apophyses,eachterminatingin a strongspine.
Male. Length about 3.5 mm.; wing, 4 mm.
Female.Length about 4-4.5 mm.; wing, '1.5-5mm.
Rostrum testiceousyellow; palpi brownishblack. Antennaeblack
throughout; flagellarsegmentsoval, more elongatein female;in male,verticils. Head brown.
with long conspicuous
Pronotum pale brown; pretergites yellow. Mesonotal praescutum
brown, the median area still darker, the lateral margins paling_toyellow; pseudosutural foveae reddish; scutum darkened; scutellum
dark, the posterior margin yellowish; mediotergite.dark. Pleura,
including pl-eurotergite,uniformly pale testaceousyellow-,unmarked.
HaltereJpble, the knobs dusky. Legs with the coxaeand trochanterstestaceousyellow; remainder of legs black. Wingq subhyaline,.thestigma a litlle darker, diffuse: veins and trichia dark brown. Venation:
Sc short, Scr ending a distancebefore origin of Rs equal to. from onethird to one-halfthe length of the latter; Rs short, nearly straight. aboutcell R3 small, vein R3 short, straight,.three-fifthsthe length of Rz+g+n;1e381

Alexander: Crane Fl,rcsot' Mexico

40t

oblique; distance on margin between veins R1a2and fts about twice thelength of latter; basal section of R5 reduced; ril,-cumore than one-half
its length beyond f.ork of.M.
Abdomen dark brown; hypopygium reddish yellow. Male hypopygium with three dististyles; outer style blackened, produced into along straight recurved spine, on outer margin near base with a slightly
smaller similar appressedspine; intermediate style blackened, at apex
produced into two subequal, slightly divergent spines,the base of stylewith two long setae; inner style pale, fleshy, simple, at apex with two
pale fasciculatesetae. Phallosomeconsistingof two darkened apophyses,
each a simple rod that terminates in an acute spine, the surface of rod
with numerous delicate setulae; central portion of phallosome elongate,
pale. extending caudad beyond the general level of the apophyses.

Ifobgpe, d, Vergel,Chiapas,altitude E00meters,May 30,
1935(A. M. Dampf); M. F. 4351. Allotopotype,I , with type.
Paratopotypes,2d &, I g, NIay 11, 1935;M. F. 4123.
Gonomyia (Gonomy'ia)guoesitais entirely distinct from allother regional speciesof the subgenus,the chief distinctions
lying in the venation and in the structure of the male hypopygium.
Gonomyia(Gonomyia)subremotasp.n.
group; generalcoloration Cark brown andBelongsto the sctbcinereo
yellow; rostrum black; scapeand pedicel of antennaeyellow, flagellum
black; head, including e):es,enlarged in male; dorsopleuralregion conspicuously yellow: u'ings rvith a u'cak gray tinge; ^Scshort, Rr*,,+narcuated,cell lst Mzclosed,tn-c'uat orvervclose to fork of M:abdominal
tergites dark brorvn, the posterior lateral angles of segments paleyellow; hypopygium yellow; male ii1'popygiunrn'ith the outer dististylebearing an erect spine on outer margin beyond midlength; intermediatestyle a slender glabrous blade; phallosome bearing three powerful
blackened spines,the basal pair symmetrical, strongly curved.
Male. Length about 4.5 mm.; wing, 5 mm.
Female. Length about 5 mm.; wing, 5 mm.
Rostrum and palpi black. Antennae with scape and pedicel clear
light yellow above, a trifle darker on ventral portion; f;agellum black.
Head light yellow, the central portion of vertex with a brown area; headof male large, including eyes;vertex'wide.
Pronotum light yellow abol'e, darkened on sides;lateral pretergites
and dorsopleural membrane, together with lateral borders of praescutum, broadly sulphur-yellow, conspicuous,the color continued caudad
across the mediotergite; disk of praescutum ancl the scutal lobes darkbrown, sparsely pruinose; pseudosutural foveae black; median area of
scutum paler, darkened behind; scutellum dark; mediotergite weakly
darkened at base and more strongly so on posterior border, the central
portion conspicuously yellow, as above described. Pleura yellow, with
two narrow brown stripes, the more dorsal extending from the cervicalregion acrossthe dorsal sclerites,crossingthe pleurotergite and including

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Annals Entomological Society of Amer'ica [Vol. XXXI,

the posterior border of the mediotergite; ventral stripe still narrower,
occupying the sternopleurite. Halteres dark brown. Legs with thecoxaeyellow, the basesof posterior pair weakly darkened; trochanters
yellow; remainder of legs black. Wings with a weak gray tinge; stigmaoval, pale brown; veins dark. Costal fringe short. Venation: Sc short,
Scr ending a distance before origin of lRsabout equal to one-third thelength of the latter, Sczcloseto its tip in both sexes;l?s long, at leastbasal section of Rs short but
one-half longer than the arcuated R2.,.3r.n;
evident; cell lst Mz closed;tn-cuat or very closeto fork of.M.
Abdominal tergites dark brown, the posterior lateral angles of thesegmentspale yellow; sternites pale brownish yellow medially, clearer
yellow on sides;hypopygium yellow. Male hypopygium with the lobeof basistyle small and pale. Outer dististyle a little longer than theothers, weakly clavate, the apex terminating in a tiny obtuse point,
the style at near two-thirds the length bearing a long acute erect spine.
Two inner styles subequalin length, the intermediatea slenderglabrous
blade from a dilated base,bearinga strong seta on outer margin at near
midlength; innermost style a little longer than the last and nearly as
long as the outer style, moderately slender,narrowed to the cultriform
apex, before which are borne the usual two long pale fasciculatesetae,
with other coarsenormal setaescatteredalong the face of style. Phallosome bearing three powerful blackenedspines,the basal pair approximately equal in size and conformation, very strongly curved; apicalspine much smaller but arising from an expandedpale baseso that the
total length is subequalto either of the basalspines.
Holotype, d, Vergel, ChiapoS,altitude 1200 meters, May 16,
1935 (A. M. Dampf) ; M. F. .1192. Allotopotype, I .
Gonomyia (Gonomyia) subremota is most nearly allied to
G. (G.) remota Alexander (Mexico, Salvador), differing especially in the structure of the male hypopygium, notably of the
dististyles.
Gonomyia(Gonomyia)multispicatasp. n.
group; generalcoloration of mesonotum
Belongsto the subcinereo
brownish gray; pleura striped longitudinally with whitish yellow andbrownishblack; legsblack; wingsgrayish,the small oval stigmadarker;
Sc short, Rr+r+astrongly arcuated,cell lst Mz closed,rn-cubeforeforkof M; male hypopygium with three simple dististyles, all apical in
position; phallosomebearing three black spines, the terminal one
smaller.
Male. Lengthabout 4 mm.;wing, 4.4mm.
Rostrum and palpi black. Antennae with scapeand pedicel yellow,
flagellumblack. Head yellow, the central portion restrictedly darkened.
Pronotum obscure yellow above, darkened on sides. Mesonotal
praescutum brownish gray; scutal lobes darkened, the median area
obscureyellow; scutellumreddishbrown; med.iotergiteyellow, darkened
posteriorly. Pleura whitish yellow, striped longitudinally with brownish black, including a more dorsal stripe acrossthe anepisternumand

1e381

Alerander: CraneFl,iesof Mex'ico

403

central pteropleurite and a ventral line on sternopleurite. -1e4 withcoxae teddirh; trochanters testaceousbrownl remainder of legs black.
Wings with a'grayish tinge, the small oval.stigma-darker; yeins dark
ending a distance before origin about
brow=n. Venation i Sc shoit, ^Sc1
equal to one-half the length of Rs, Sczat extreme.tip of ,Sc1;R2^as
Lrstiongly arcuated; cell IslM2 closed;rn-cua short distancebeforeforkof. M.
Abdomen with tergites dark brown, variegatedwith paler; sternites
and hypopygium pale. Male hypopygiuq ryith_apical prolongation of
all.simple and subbasistirlesfrort and stout. Three apical dististyles_,equal-in length, not very different from one another in conformation;
ooter style terminating in an acute spine,widest at near midlength,.thesurfacewith abundant setae;middle style narrow, especiallyon distal
portion, with a singlestrong seta on face at near midlength; inner stylei relatively broad plate, the outer apical angle produced into a stout
flattened ipinous pbint, the surface with several coarsesetae; before
apex with iwo pale fasciculatesetae. Phallosomewith three blackened.pin"t, including two strongly curved onesnear baseand a smaller andmore slenderapical spine from an expandedbase.
Holotype, d, Vergel, Chiapas, altitude 725 meters, May 21,
1935 (A. M. Dampf) ; M. F. 4259.
The nearest a1ly of the present fly is the closely related
Gonomyia (Gonomyia) subremota sp. n., which, while similar ingeneral appearance, differs conspicuously in the structure of themale hypopygium, especially the dististyles.
Gonomyia (Gonomyia)expansasp. n.
gfoup; generalcolorationdark brown andBelongsto the subcinerea
pleurawith two narrorv brorvn stripes; rostrum black;vellow. tfiel,ntennaewith scapeand pedicel yellorv, flagellum black; wings subhyaline, stigma pale brown; va-gue_
Plle brown washeson cord, outer"r,d of cell /-sl Mi and,in basesof cells Cu and lst A; .Scshort; cell lst Mz
closed; ,n-cushortly beforefork of M; maTehypopygium with the outerdististyle a pale blide, expandedat near midlength and here produced
into a darkenedmarginal tubercle.
Male. Length about 4.3-4.5mm.; wing, 4.8-5 mm.
Female.Length about 5 mm.; wing, 5 mm.
Rostrum and palpi black. Antennaewith scapeand pedicely-e!ow,
flagellum black; int-ennae (male) .relatively_long and slender,.if bent
balkwards extending to beyond wing-root; flagellar segmentselongateoval to subcylindricil, wittr,moderatelylong verticils. Head yellow, thecentral portion of vertex marked with dark brown.
Pronotum yellow and brown; lateral pretergite.s-lightyellow. Mesonotal pttesc,tiom and scutal lobes dark brownish EraY, the central
region^ofscutum a little paler;lateral border of praescutumrestrict_edly
y"ilo*; a yellow spot on posteriorportion of.eachscutal lobe.;scutellum
obr.nt" brown; medioteigite testaceousyellow, the posterior portion
darkened,being a continuition of the dorsal dark pleural stripe. Pleura404

Annals Entomological
soc,ietyof Arner,ica[vol. xxxl,

yellow,-with two narrow dark br_ownstripes, the more dorsal extending
from above the fore coxae to the mediotergite, as described; ventra'i
stripe occupying the ventral sternopleurite and meron. Halteres brownish black. Le_gsyit_h the coxae and trochanters yellow, the posterior
coxae r_r.arrowlydarkened at .bases; remainder of legs black. Wings
subhyaline;^stigm?oyql, pale brown; vague pale brown clouds on cord,
outer end of cell /sl Mz, and as a wash in basesof cells Cu and lst Ai
veins dark brown.. lost-al_fringe short. Venation: Sc short, Sc1ending a
distance before origin of l?s about equal to two-thirds the'length of ihelatter, Sczat its extreme tip in male, some distance removed ftom tip infemale; Rz+,a+o
nearly- as long as Rs, moderately arcuated; r-m at orclose to fork of Rs, basal section of R5 thus obliterated; cell lst Mz
closed; rn-cu shortly before f.ork of.M.
Abdominal tergites dark brown, the caudallateral angles of theindividual segments ]i_g!t -yellow; sternites more uniformly yellow;
hypgpygium yellow. Male hypopygium with the lobe of basijtyie pale,moderately.elongate. Outer dististyle an elongate pale blade, at near
midlength- slightly expande4 and at margin at this point produced intoa low darkened tubercle. Intermediate style a slender, gently curved
pale blade, parallel-sic1_ed,
the tip obtusely rounded; shortly beiore apex
*itl, a powerful seta. Inner dististyle nearly as long as the interrnediate,with two fasciculatesetaeat apex, additional to nbrmal setae. Fhallosome conspicuous,consistingchiefiy of a strongly sinuousslenderblackened rod or spine, the tip acute, superimposedover the pale aedeagus.
Ilolotype, d, Finca Lubeca, Chiapas, altitude 850 meters,

June 2I, 1935(A. X'I.Dampf); M. F. 4582. Allotype, 9, \rergel,
altitude 800 meters,June 10, 1935; N4.F. 4502. Paratype, &,F'incaGermania,altitude 1190meters,June24, rg}s; N{.F. 4bzr.
The structure of the outer dististyle of the male hypopygium
is much as in Gonomy'ia(Gonamyia)tuberculataAlexander(\{exico), which has cell L[z of.the wings open and with the otherdististylesquite dif;'erentlyconstructed.
RhabdcmastixSkuse(Rhabdomastix)
Rhabdomastix
mexicanasp. n.
Generalcoloration of notum opaquelight {ay, the praescutumwith
four brown stripes; antennae with basal segmentsyellow, the outer ones
darker; pseudosutural foveae black; knobs of halteres dark brown:
wings with a weak brown tinge, stigma darker; Rz+r+oshorter than thestraight Ra; vein R3 oblique, about equal in length to the distance on
costa between the tips of veins Rr+zand l?s.
Female. Length about 7.5 mm.; wing, 6.3 mm.
_$o9trury pale; palpi black. Antennae with the scapeobscureyellow;
pedicel and basal two or three flagellar segments pale yellow;
outer
flagellar segments passing into brown; basal segment of flagellum
nearly as long as segments two and three combined. Head brownish
gray, the front yellow.

1e381

Al,exander:CraneFlies of Mexico405

Pronotum brownish graY,- variegated with yellow; pretergites lightyellow on lateral portions_. Mesonot_alpraescutum opaque-light gray,*itfr four brown siripes, the intermediate pair confluent in front; pseudosutural foveae subcircular in outline, black; posterior sclerites ofnotum brownish BraY, the scutal lobes darker. Pleura brown, very
sparsely pruinose.- H-aitereswith stem pale.yellow, klob dark brown.
t "g. *ittr the coxae and trochanters_paleyellow, th._fore coxae a trifle
darier; remainder of legs broken. Wings with a weak brown tinge,-theoval stigma darker brown; indicationq of ? vefy weak darkenedcloud on
cord; vii.ts brown. Macrotrichia.of veins beyond cor{. vgry spafse'
there being about five or six on distal fourth of each of veins Rr, andM+2. Veiation: Sc long, Sc, ending ."!o"l oqposite four.-fifths theshortei than tle straigh! R4i Rsoblique, about egr1al
tengih of ,Rs;Rz+e+e
in lingth to ifre d.istanceon costa betweenth-e-tipsof veins Rr+zand Ra;
cell /s"l Mz smal| the secondsection of vein Mr+, only about one-fourth
as long as the outer section.
Ab"clomendark brown, the caudal borders of the segmentsnarrowly
light gray.
Holotype, 9, Vergel, Chiapas, altitude 800 meters, June 3,
1935 (A. M. DamPf) ; M. F. 4417.
The nearest described relative of the present fly is Rhabd,omastix (Rhabdomast'ix) isabella Alexander, likewise from
Chiapas. It is readily told by the diagnostic features listed,"rp""i"lly the opaque gray coloration of the body, and in the
delails of venation, especially of the radial field.
Rhabdomastix (Rhabdomastix)longiterebrata sp' n'Femal,e.Length about 8 mm.; wing, 6.5 mm.
Closely relatEd to R. (R.) mesicarta.sp.n., differing especiallyin
the venation and structure of the ovipositor'
Head broken. General coloration of thorax much as in me*icanalegs almost uniformly yellowish.brorvn,the outer tarsal segmentssomeWings more grayish; stigma darker brown, clea-rlydelim-Macrotrichii*,fi"i a"tt.r.
of ,r.Ir,r more numerous, involving _almost theitil.
."tii" i"ngtfr of distal section of Rs and outer ends of distal sectionsofb;ih Arl,*7and.Ms. Venation: ,Scsh-ort,Sc1ending-oppositethree-fifths
it" t.ngih of Rs; R3 nearl] perpendicular, short, the distance on costa
U"i*".i veins Rr+zand R, ne-arlytwice its length; Raarcuated,subequal
cell rRa-atmargin ottly a little -more than one-half that of;-R;;;*r;."U-nri'{n *r*itino, cell R, is approximately two-thirds as extensiveas
Mz small, relatively ihort and broad, the distal section of
R;;"li-lst-'b"ipotitot four times the secondsection'
Ar*rnearlywith the cerci unusually ]9"g and. very. slender, morethan one-third longer than the correspondingcondition in mericana.
Holotype, 9 , vergel, Chiapas, above a waterfall in a damp

valley, utiitoa" 800 meters, June 1, 1935 (A. M. Dampf);
M. F'. 4384.

406

Annals EntomologicalSoc'ietyof Arnerica [Vol. XXXI,
Erioptera Meigen

Erioptera(Erioptera)laetipleurasp. n.
Mesonotalpraescutum
brown,the cephalicthird andhumeralregion
light yellow; dorsal pleura and pronotum with a conspicuous black
stripe, ventrad of which is a slightly wider white stripe; knobs of halteres
more or less darkened; femora pale, with dark vestiture, including flattened scalesas well as setae; terminal tarsal segmentsyellowish; wings
with a strong dark brown tinge; vein ?nd .4 moderately sinuous; malehypopygium with two slender dististyles.
MaIe. Length about 2.8 mm.; wing, 3.2 mm.
Female. Length about 3-4 mm.; wing, 3.5-4.4 mm.
Rostrum and palpi brown. Antennae brownish black throughout;flagellar segments oval. Head with central portion of vertex darkbrown, the front and orbits pale.
Pronotum black, the color continued longitudinally and obliquely
acrossthe dorsal pleurites as a broad stripe, passingbeneath the halteres,
reaching the abdomen. Mesonotal praescutum with the cephalic third
and the humeral margin light yellow, the remainder of praescutum
brown, the usual interspacesnarrowly darker brown, the median stripe
feebly indicated on the yellow anterior portion of the sclerite but
becoming obsolete behind at near midlength of the sclerite; posteriorsclerites of notum more uniformly dark brown. Pleura with a dorsal
black stripe, as above described; ventrad of this, a slightly wider white
stripe extending from the cervical region to above the hind legs; ventral .
pleurites brownish black. Halteres with stem light yellow, knob more or
lessinfuscated,least-soin allotype-male. L.gf with the fore coxaepale,remaining coxae dark brou'n; trochanters yellow; legs chiefly dark, thecolor producedby abundant setaeand elongateflattened scales;extreme
tip of femur whitened, precededby a \-aguesubterminal darkening; tipof tibia and the tarsi passinginto more yellon'ish. \Vings with a strong
dark brown tinge, the veins and macrotrichia dark. \/enation: Vein
?nd A moderately sinuous.
Abdomen brownish black, with conspicuous erect setae. Male
hypopygium with two slender dististyles, the outer one terminating inan acute decurved spine, with a smaller similar spine on outer marginclose to apex. Inner style simple, a little expanded on outer third.
before apex on lower margin with a small fleshy lobule. Ovipositor withthe cerci reddish horn colored, strongly upcurved.

Holotype, g, Vergel, Chiapas,altitude 800 meters,June 3,
1935 (A. M. Dampf) ; M. F. 4416. Allotype, I , Santa Isabel,
Chiapas,altitude 800 meters,November 16, 1930;M. F. 1880.
Paratypes, 1 I with allotype; 1 broken individual, apparently
9 , Finca Germania, altitude 1190 meters, June 20, 1935;
M. F. 457TA,
Erioptera (Erioptera) Ie,t'ipleurais very different from all
speciesof the subgenushitherto made known. The presenceof

1e381

Alexander: Crane Fl'iesof Mexico407

scales on the legs had not been described in the subgenus
Erioptera but had been detected in Empeda and in the allied
genus GymnaslesBrunetti.
sp. n.
fuscodiscalis
Erioptera(Mesoc5ryhona)
Femora with a single dark subterminal ring; wings brownish yellow;
anterior cord with a cbnspicuousdark brown blotch; male hypopygiumwith two slenderdististyles, the outer one stouter, at apex with numerous
coarse setae; gonapophyses appearing as slender black spines.
Male. Length about 3 mm.; wing, 3.4-3.5 mm.
Female. Length about 3.5 mm.; wing, 4mm.
Rostrum and palpi black. Antennae black throughout; flagellar
verticils long. Head brown, the anterior vertex light gray.
Pronotum pale medially, dark brown sublaterally. Mesonotal
praescutum with three pale yellowish brown stripes, clearly delimitedby dark brown interspaces and dark lateral margins; scutum p".t. Xtlowish brown, each lobe bordered externally and internally by slightly
d.arker brown lines; scutellum pale; mediotergite variegated with
obscure yellow and brown. Pleura conspicuously striped with dark
the latter color including a stripe above the midbrown and silvery,-behind;a broad ventral band on the mesal region.
coxae, expanded
Halteres yellow, knobs dgrk brown. Legs with the fore coxae darkeled,remalnlng coxae and all trochanters yellow; femora yellow, with abrown subterminal ring; tibiae and tarsi yellow. Wings with a brownish
yellow tinge, darkest before cord and in outer radial field;_prearcular
ind costal regions, as well as a blotch near cord, clearer yellow; a conspicuous darli brown discal area on anterior cord; veins pale, darker intlre clouded areas. Venation: Cell ?nd. Mz open by atrophy of basal
section of Ms, as in subgenus.
Abdomen,' including hypopygium, brown; pleural membrane still
darker. Male hypopygium with both dististyles apical, slender; outera densestyle with a stou[ stem, at apex a little expanded and beari.nS_
brush of coarsesetae,with additional longer, more delicate bristles intermixed; inner dististyle fused with the outer on about the basal fourth,slightly shorter than the outer, pale; stem slender, at apex weakly
ex[anded into an oval head. Gonapophysesappearing as slender black
splnes.

Holotype, & , Finca Germania, Chiapas, altitude 1190
meters,June20, 1935(A. M. Dampf); M. F.457I. All,otopotype,g. Paratopotypes,
several d9;paratypes, &9, Vergel,altitude 1000 meters,June 3, 1935, M. F. 44L6; June 26, 1935'
M. F. 4643; July 2, 1935,M. F. 4744;Finca Victoria, altitude
900 meters,June 29, 1935;M. F. 4709,4710.
Erioptera (Mesocyphona)fuscodiscal'isis amply distinct fromall describedmembersof the genus. The structure of the malehypopygium is entirely diflerent from anything hitherto foundin Mesocyphona.

408

Annals EntomologicalSoc'ietyof Arnerica [Vol. XXXI,

Molophilus Curtis
Molophilus (Molophilus) miraculus sp. n.
Belongsto the plagiatusgroup; mesonotalpraescutumand scutum
pale brown, the posterior sclerites and pleura dark brown; antennae
short; male hypopygium with the basal dististyle profoundly bifid nearbase, the outer arm a slender curved rod, the inner arm much more
powerful, before apex producedmesadinto a strong spine.
Male. Length about 3.5 mm.; wing, 4 mm.
Rostrum and palpi black. Antennae short, dark throughout;flagellar segmentsoval. Head dark.
Pronotum dark. Mesonotalpraescutumand scutum uniformly pale
brown or testaceousbrown, the posterior scleritesdarker. Pleura darkbrown. Halteres pale yellow. Legs with the coxae and trochantersbrownish yellow; remainderof the legsbrown, the outer tarsal segments
somewhatdarker. Wings with a gray tinge, the prearcularfield a littlebrightened; veins pale brown, the macrotrichia dark. Venation: Rz
lying shortly beyond level of.r-m; petiole of cell Mt about twice n -cu;
vein ?nd A ending oppositern-cu,.
Abdomen dark brown; hypopygium more yellowish brown. Male
hypopygium with the ventral lobe of basistyle relatively small, theapical beak slender. Outer dististyle with arms slender,the inner one
longest. Basal dististyle profoundly bifid near base,the outer arm aslendercurved rod; inner arm much more powerful, before apex producedmesadinto a strongstraight spike-likespinethat is about one-halfthe length of the arm itself ; apexof arm beyondthe spike producedinto
two or three smallerspinouspoints.
Holotype, d, Vergel, ChiapoS, altitude 1000 meters, June 3,
1935(A. M. Dampf); M. F. 4416.
Molophilus miraculus is entirely distinct from all other
Neotropical speciesof the genus, the basal dististyle of the malehypopygium being quite different from any other known form.
Molophilus (Molophilus)ductilis sp. n.
Belongsto the plagiatusgroup; sizesmall (wing, d , 3 mm. or less);
generalcoloration dark brorvn; antennaerelatively short; male hypopygium with the basaldististylean unusuallylong, slender,simplerod,
the basalportion with numerousappressedspinulae.
Male. Length about 2.3-2.5mm.; wing, 2.8-Bmm.
Fernale.Length about 3 mm.; wing, 3-3.2 mm.
Rostrum dark brown, palpi black. Antennaemoderatelylong, darkbrown throughout; flagellar segmentsoval, with very long verticils.
Head dark brown.
Mesonotum almost uniformly dark brown, the posterior scleritesa
little darker. Pleura dark brown. Halteresinfuscated,the baseof stem
restrictedly pale. Legswith the coxaeand trochantersbrownishyellow;
remainderof legsbrown, the outer segmentspassinginto black. Wings
with a weak brownish tinge, the basal portions a little more yellowish:

1e381

Alexander: Crane Fl'iesof Mexico409

veins pale brown; trichiae, including the long densecostal fringe, darkbrown. Venationr Rz n€arly opposite r-m; petiole of cell Ms about
one-half to three-fourthslonger than n -cu; vein ?nd ,4 ending about
oppositeor shortly beyond rn-cu.
Abdomen, including hypopygium, dark brown. Male hypopygiumwith the apical beak of the ventral lobe of basistyle slender. Outerdististyle with the outer blade expanded,the inner longer and more
slender. Basal dististyle an unusually long and slender simple rod,
exceedingin length the outer style; numerousappressedspinulae on
subbasalhalf and more scatteredsetigerouspunctureson distal third.
Phallosomicplate short-oval, the apex bluntly obtuse to subtruncate.
Holotype, d, Finca Victoria, Chiapas, altitude 900 meters,
June 1, 1935 (A. M. Dampf); M. F. 4391. Paratopotypes,& 9,
J u n e 2 9 , 1 9 3 5 ,M . F . 4 7 0 9 , 4 7 L 0 , 4 7 1 0A ; p a r a t y p e s ,& , 9 , F i n c a
San Cristobal, altitude 1090meters, June 19, 1935, M. F. 45C0;
Finca Germania, altitude 1190 meters, June 20, 1935, M. F.

457t.
This small speciesis readily told from all regionalforms by
the combination of small size, relatively short antennae, andthe structure of the male hypopygium, especially of the basal
dististyle.
Molophilus (Molophilus) sagax sp. n.
Belongs to the plagiatu.sgroup; antennae (A) short; generalcoloration of thorax dark brown, the praescutum a little brighter; malehypopygium with the basal dististyle a long slender rod that terminatesin a slender straight spine, at near two-thirds the length on outermargin with a small acute spine.
Male. Length about 3 mm.; wing, 3.8 mm.
Rostrum dark brown, palpi black. Antennae short, dark brown
throughout. Head dark brown.
Mesonotal praescutum brorvn, the anterior interspaces a trifle
darker; posterior sclerites of notum and the pleura darker brown.
Halteres dusky, the large knobs a little paler. Legs with the fore coxae
darkened, the remaining coxae and all trochanters obscure yellow;
remainder of legs brown, the terminal tarsal segments darker. Wings
relatively broad, strongly tinged with brown, the stigmal region extensively but diffusely darker brown; veins and trichia brown. Venation:
R2 lying some distance beyond level of r-m; petiole of cell Ms relatively
short, about one-half longer than m'-cu;vein ?nd ,{ long, extending toabout opposite midlength of petiole of cell Ma.
Abdomen, including hypopygium, brown. Male hypopygium withthe beak of ventral lobe of basistyle slender, gently curved. Outerdististyle slender, the outer arm a broadly dilated, obtuse blade, theinner arm long and slender. Basal dististyle a long slender rod, terminating in a slender straight spine; at near two-thirds the length onouter margin a small acute subappressedspine, the style beyond thispoint a little dilated.

410

Annals EntomologicalSocietyof Arnerica [Vol. XXXI,Holotype, d, Vergel, Chiap&s,altitude 1000meters,June B,
1935(A. M. Dampf); N{. F. 4416.
Moloph'ilussagar is readily told from all related speciesof
the genusby the structure of the male hypopygium.
Molophilus(Molophilus)falx sp. n.
Belongsto the plagiatusgroup; antennae(d) long; generalcolorationof thorax dark grayish brown; halteres yellow; male hypopygium withthe basal dististyle a strongly curved sickle, at near midlength bearing
a low obtuse flange, the surface of style with microscopicappressed
spinulae and scattered setigerouspunctures.
Male. Length about 3.8-4 mm.; wing, 4.3-4.5mm.; antennaabout
1.8-2mm.
Rostrum and palpi black. Antennae brownish black throughout,elongate in male, as shown by the measurements;flagellar segments
more or lessfusiform or flask-shaped,the expandedbasal portion withvery long verticils that exceed the segmentsin length. Head darkgrayish brown.
Mesonotum almost uniformly dark grayish brown, the anterior
lateral pretergitesrestrictedly brightened. Pleura dark. Halteres paleyellow. L"gs with the coxaedarkened;trochantersyellow; remainderoflegs dark brown, the femoral basesrestrictedly yellow. Wings tinged
with brown, the extreme base brighter; veins and macrotrichia dark.
Venation: R2 lying some distance beyond level of r-m so that .R2.qis
approximately three times ftr+si rn-cuoblique and sinuous,about twothirds as long as petiole of cell Ms; vein ?nd A long, ending about
opposite one-third the length of the petiole of. cell Ms.
Abdomen, including hypopygium, dark brown. Male hypopygiumwith the apical beak of basistylevarying from relatively stout to moderately slender. Basal dististyle a strongly curved hook, more or lessresemblinga sickle, at near midlength bearing a low obtuse flange;
beyond this latter point, the style narrows gradually to the acuteblackened point; surface with microscopic appressedspinulae andscatteredSetigerouspunctures.
Holotype, d, Vergel, Chiap&S, altitude 800 meters, May 29,
1935 (A. M. Dampf) ; M. F. 4334. Paratopotype, d, altitude1000 meters, May 19, 1935, M. F. 4205.
Molophilus f alx is quite distinct from all other regional
species in the combination of elongate antennae and structureof the male hypopygium, especially the falciform basal
dististyle.
Toxorhina Loew
Toxorhina (Ceratocheilus)chiapasensissp. n.
Sizesmall (wing, cy',lessthan 4 mm.); mesonotumalmost uniformly
brown; pleura obscureyellow, variegatedwith brown on anepisternum

1e381

Alexander: Crane Flies of Mexico

4TT

and ventral sternopleurite; legs brown, the terminal tarsal segment
paling to yellow;wings faintly tinged with brown, immaculate; Sc ending
about opposite four-fifths the length of Rs; anterior branch of Rs longer
than Rs itself ; cell /sl M2 roughly pentagonal, r-rn opposite midlength,
rn-cu opposite one-third the length; basal section of M+z longer thanthe other elements comprising cell lst M2; cell ?nd A narcow.
Male. Wing, 3.7 mm.
Rostrum black, shorter than the body.
Mesonotum almost uniformly brown, the scutal lobes a little darker
than the praescutum. Pleura obscureyellow, variegated with brown on
the anepisternum and ventral sternopleurite. Legs with the coxae andtrochanters testaceous yellow; remainder of legs pale brown, the lasttarsal segmentpaling to yellow; claws simple. Wings with a faint browntinge, immaculate; veins dark brown. Macrotrichia of veins numerous,
including extensive series on distal three-fourths of Rs, all but base ofanterior branch of Rs, both sections of vein l?5, and all outer medial
veins. Venation: Sc relatively long, ,Scrending about opposite four-fifths
the length of Rs, ,Scsatrophied; anterior branch of Rs long, exceeding
Rs itself; cell Rz at wing-margin about one-half cell l?a; cell lst Mz
roughly pentagonal in outline, the longest element being the obliquebasal section of M42; cell lst Mz shorter than any of the veins beyond
it; r-m opposite midlength of cell /sl M2, ffi,-cuat near one-third thelength; basal section of Ms",aa little shorter than the distal section ofvein Cur; cell ?nd A narrow; vein -1slA nearly straight, its basal portion
not approximated to Cu as is common in the genus. Abdomen broken.

Holotype, d , F'inca Belem, Chiapas, altitude 800 meters,
June 22, L935 (A. N{. Dampf); M. F. 4596.
Toxorhina (Ceratocheilus) chiapasensis is very distinct fromthe remaining Neotropical speciesof the subgenus so far madeknown. In the venational characters of long Sc, position of
rn-cu beyond the fork of. M and slight basal approximation ofveins Cu and lst A, the fly comes closest to the white-footed
species, T. (C.) leucostenaAlexander (Dutch Guiana) andT. ( C.) nizteitarsis Alexander (Panama-Colombia). It differs
conspicuously in the sma1l size, darkened legs, and, especially,
the details of venation, as in the medial field where the basal
section of vein Mt+, exceedsin length the second section.
Toxorhina (Toxorhina) trilobata sp. n.
Generalcoloration Bray, the praescutumwith three brown stripes,
the median one divided behind; halteresyellow; legs yellow, the tipsof all tibiae conspicuouslyblackened;male hypopygiumwith the apicalspine of basistyleslender;outer blackenedlobe of dististyle trilobate;arrns of aedeagusof moderatelength, subtendedand slightly exceededby the bladesof the gonapophyses.
Mal,e. Length, excludingrostrum, 8-10 mm.; wing, 5.5-6.5mm.;
rostrum,4.5-5.5mm.

4L2

Annals Entomological Society of Arnerica[Vol. XXXI,

Female. Length, excluding rostrum, 10-11 mm.; wing, 6-2 mm.;rostrum, 5-5.5 mm.
Rostrum pale brown, the base and apex narrowly more blackened.
Antennae black throughout. Head Eray, the anterior vertex moreyellowish gray.
Pronotum and cervical region black. Mesonotum grayr the lateral
margins lighter; praescutum with three conspicuousbrown stripes, themedian one very naffowly divided on posterior portion; scutal lobes
darkened. Pleura gray, sparsely variegated with darker on ventral
anepisternum and ventral sternopleurite. Halteres light yellow. Legs
with coxae gray; trochanters obscure yellow; remainder of legs yellow,
the tips of all tibiae conspicuously blackened; outer tarsal segments
weakly darkened. Wings subhyaline, the prearcular and costal portions
a little more yellowish; veins dark brown, the prearcular ones paler.
Venation: Sc ending opposite or before origin of Rs; nt-cu before fork
of.M.
Abdomen elongate; basal tergites brown, the caudal margins a little
darker; subterminal segmentsmore uniformly darkened; basal sternites
more uniformly bicolorous; hypopygium reddish brown. Male hypopygium with the apical spine of basistyle very slender. Dististyle complex in structure, the darkened basal portion trilobed, the more basal
lobe obtusely rounded, the outer lobes slender; outer blade of style witha triangular flange on outer margin. Arms of aedeagusof moderate
Iength, slightly exceededby the subtending blades of the gonapophyses.

Holotype, d, Vergel, Chiapas,altitude 800 meters,June 3,
1935(A. M. Dampf); M. F. 4417. Allotopotype,I , pinnedwith
type. Paratopotypes,
severalof both sexes,May l3-June t2,
1935, M. F. 4151, 4268, 4334,4385, 4398, 4502,4503, 4524.
ParatyPes, I I , San Cristobal, June 19, 1935, M. F. 4560;
Finca Belem,altitude 800 meters,June 23, 1935,M. F. 1602.
The nearestally of the presentfly is Toxorloina(Toxorhina)
rnendosaAlexander (southern Brazil) rn'hich differs chiefly inthe details of structure of the male hypopygium, as the stout
apical spine of the basistyle and the differently constructed
dististyle.