6f 3

'

Frotn

THE PROCEEDINGS OF THE
ROYAL ENTOMOLOGICALSOCIETY OF LONDON
Series B. TAXONOMYVoI. Ig, Partsf6,

June t 5th, IgSo

The Royal Entomological Society o! London.
4L, Queents Gate, S.W.?

pRINrrD ar aDr,/$D & Bo!t, rrD., BABtEorrorEw EBrBg, Doarnto

85

NI'W OIT .I,ITTLE-KNOWNSPI]CIESOTTIJXOTIUTIPUTIDAU(DIPTHRA): VII.
tsy Pnol.ESsoR
CHenr,nsp. Ar,oxer.rDr4n,
F.R.U.S.
( U n,iL,et'.sityof trI ussu*husetl,;,,Amher.*t, II assac
ltu.selts.\

l'"" lrrecedingpart in this seriesof paperswas publishedin lg4,9,prrc. H. e*t.
Soc'Ltryul' (s) 18 : 153-159. The speciestreaied.herewith
arc fronr-ifrir",wlterr:thcy wcto ttrkcn chiefly by -i friend.s,the lateDrs. Jo1. A. niy.* ,"A
Jr-r,erb
A. Reis. The typer ui" p**tr*d in my collection.
Tipula (Acutipula)auspicissp. n.
siz. r*lat'ively slrall (wing, male, about lb mm.)
; antennae of male olongate, about
one-ha'lf as long as body ; general coloration of mesonotum
reddish b'own, the posteriorsclerit'esand llleura vellow, uumarked; legs obscureyellorv, tbc tips of the femora a.dtibiae trarrowly and very inconspicuously iarkened;wings with a very weak brow'ish
tirrge, cells c'and Bc sca,rcelytlarker, stigma longand narrow, pale brown ; Rsshort andweak, about two-thirds \n-cu; abdornir*t tu"gi6s reddish
bro-wn,sternites and hypopygiu[r yellow; tnale hlpoplgiunr rv.ith the tirgite
produced into & narrow clepressedflatterred bladc that splitt *t up"* into two *"luyaivergent ""rlik" lobes, unpr.ovidetl
lvith spines; itrner dististyle ,uith th" entire ouber
surfacJprovided with very abu'tla't
crect, spi'ous setae that are p'oduced at tips i'to long
hairlilie points.
Male.-r'ength about 14 mrn.; rving ib.5 ,.rrrr.;
antenna about 7 rnm.
Front'al prolongation of head trown]relatively
long; nasus small and inconspicu'us,rvith long black set-ae; palpi black. A'tennae (male)elongate, as shown by the me&surertteuts; scape' pedicel and extreme base of finstflagellar"seg-*tyellow, remainder ofolgan black ; flagellar segmentslong-cylindrical, withiery r*uiy a""aopun basal swelli'gs,
the verticils shorter than the segments, restricted.
to ine "irrg-"rrtr;
rernainder ofsegme'ts with an abundant erect white pubescence.
Head cinn-amon brown, the fro't
paler; posterior orbits narrowly rnore yellowed.
Protrotum reddish brown -"dituy, paling to yellorv
on the sides. Mesonoturn chiofly
rcddish brown, rvithout distinct ctarklrait"t.nltluicutellum
and lrcstnotum more yellowed.
Pleura yellow' Halteres pale brown, the baseof the stern restri^ctedly yellow. r,egs withall coxae and trochattters yellow ; reruaind.erof
legs obscure yellow, the tips of the femor.aand tibiae nan'ou'Iy and very inconspicuously da-rkened;
oirt"" tarsal segments scaroely
infuscated' wings with a- very wea,k brownish
tinge, cells c and Bc scarcely darker;
stigma long and na-l'lo'|4"pale brown, inconspicuous; veins brow', in the costal field. moreyellorvish-trrown' venation : 8s short andweak, about two-third s nx-arand three-fifths
the long Rr+r; Rr+z gently upcurved; petioleof cell Mland,na subequal; cell lst tr12
relatively lottg, the second section of M
1ja more tha' twice the bas€rlsection.
Abdrr,rirtal tcrgites redclish brown,-sternites lightyellow; hypopygiuur yellorv. .ulalc
hyPopygiu'r rvith t'he caudal bord.erof teryite prJau""ainto a ,rn-""o* dcprcsse4-llatte'ed
blade' at apex split into trvo weakly diverlenf,earlike
lobes that ar.eseparated by a dee'U-shaped notch ;virtually free frori setae but with sever.alrnicroscopic
punctures.
.lofesouter dististyle rcIatively
long and n&rrow, the tip obtuse. rnner dististylewith the beakand outer part of t'he style .truryd".ttely- coveled
bv a cushion of short errcct,spinous seta*t'hat are produceda't tips into very long hairlit" poiot,
Iower beak of style a low dar.ke'ed
flange t'hat is produced into two spittoi,s points,^one ;at eithel e1d of the flange.

Blotfpa, 6, Ceunnouu: Boopa,g.iii. lg3l (J. A. Reis).
Tipu,la(Ac'utiprtla-)
uuspiclsis.inost "gdt
to T. (A.) oryndexander,-related.
which differs evidently in coloration and.in"alld.etailsof structure of the malehypopygium.
pR',u' fi'' 'NT. suu. L,ND.
(i;) 19. prs. 5-6.

(.rurs$ Ig50.)

86

Prof. 0. P. Alexandet ott,rleut)or l,ittle-lm'owrt'

exsulsP.n.
Pseudolimnophila
Allied to fru,gi; general coloration of mesonotal praescuturn reddish brorvtt, with tlrreellal.row brolvn stripes; pseudosutural foveae blackened, tuberculate pits pale; antetrnae
relatively long, blick, the base of the first flagellar segment yellow; halteres darkened,
the base of stem yellow; legs brown, tarsi deepening into black; wings with a stt'ong
brownish tinge, very restrictedly patterned with darker brown, chiefly including the stigma'r-m a't'
arrd a cloud over the anterior cord ; 8s in direct longitudinal alignurent with -Et ;rrr just before the fork of rts i m-cu at, frorn one-t,hirclto one-fifth the length of cell lst
III z i alterior arculus preserved ; abdomen with tergites brown, their post'erior lateralborders obscure yellow, sternites uniformly yellow.
tr'enwle.-Length about 8.5 mm. ; wing 9 mm. ; antenna about 3 mm.
Il,ostrum blaJkeled, sparsely pruinose; palpi black. Antennae (fernale) relatively
long, as shown by the ,rr"uuor"orents, black throughout, with the exception o_fthe narrorv
y"lio* base of the first flagellar segment,; outer flagellar segments long-cylindrical, theiolgest verticils subequal in lengtli to the segments ; terminal segment,subequal to or
l.lnger than the penuitimate. Head gref, -or" brownish grey oll t,he ocntral part' of
1r"rl"* ; anterior vertex about twice the diameter of sca,pewith an oval irnpressed mediarral€&.

Prolotnm dark brownish grey. Mesonotunrchiefly reddish browrt, rvith three llal't'ow
but, relatively distilct brown rt,.ip"r, the central one most,distinct; pseudosut'ulalfoveae
long-oval, biackened; tuberculate pits rnuch paler, reddened, placed close to cephalicborder, separated frorn one anoth"" ny a distance equal to about, one-third their own
cliarleter ; scutal lobes and disk of scutellum weakly darkened ; mediotergite extensively
grey pruin<-rse,the lateral and posterior portions more brownish yellow; pleurotergit'e
clarkened, its cephalic part paler. Pleura brown, the pteropleurite behind and the meral&re& more yellowed. Halteres d,usky, base of stem yellow, knob dark brown. Legs withthe coxae testaceous yellow, the fore pair a trifle d.arker; trochanters obgcure yellow;
renrainder of legs brown, the femoral basls a little brighter ; tarsi passing into black' Wings
with a strong brownish tinge, very restrictedly patterned with darker browtt, the latterincluding the stigura, a cklud over the anterior cord, and more restricted areas over thetrrigin offir, posderiorcord ancl outer end of celllst Mr; prearcular field lllore brownish
y"ilonu ; veilJ brown. Venation : Sc1ending just beyorrtl the fork of -Rs,,Scanear its tip ;,,?, an,l Rr+, subequal or the latter somewhat shorter ; Rr+r+nrather strongly arcuate;
Ao in direci longitudinal alignment with .Ilu, the basal section of the latter lacking i r'nr
at <rrimmediat"iy b"for" the fork of -Bs; cell lst M, rectangular, parallel-sided, with m'-crt'
at from one-thircl to one-fifth its length ; vein znd A strongly arcuated near outer end ;anterior arculus preserved
Abdominal teigites brown, the posterior lateral bordert of the segmetrtsobscure yellorv ;tlorsopleural membrane darkened; sternites clear light yellow.

llolotypo, ?, SuoeN: Outlaw's village, Boma Plateau, altitudr: 37U0f1,.,
No. 10,503.
20.ii.1939(J.G.Myers); Collector's(1924,Atk. Zoul. 16
By *y key to tf,e African speciesof Pseud,ol'iuurtoplaiLa(Iti) :4-T), ttt. presentfly runs to fnqi (Bergroth),of $outh 'l{t'ica, whichappearcio be tLe nearuri d.rcribed.alty. The new form differs evidently in*ii.l.tuils of colorationof the body, wingsand legs,antl in the detailsof venation. X'romthe likewiseallied P.-senenAlexander,it differsin its surallersizc
and other characters.
bisatratasp. n.
PseudolimnoPhila
Generally similar to ourantiaca ; size medium (wing, female, bimru.) ; general coloratiorrof uresolot*l pr""r"otum and scutum almost uniform light fulvous yellow, without distinct
ruarkings; head dark brown, pruinose ; antennae with scape brownish yellow, sparselyspea'iesof euoti,aTiyrulidne(Diptera)

8?

pruittose, pedicel Lrrownishyellow ; pleural region yellow, on either side with two conspicuous
subcil'c'ular black Are&s,one oocupying rnost of the pleurotergite, the other rnuch of theartepistet'nurrl ; knob of haltere darkened ; legs light brown ; wings with a str.ong fulvous
brorvtr tinge, the prearcular and costal fields a little darker; stigma srnall, pale brown;
I'ein .82 very faint ; veins -8, and Bn strongly divergent at tips ; nl,-cujust beyond the forkof M; abdominal teryites dark brown, sternites obscure yellow.
Fem,ale.-Length about 8 nrrn. ; wing 8 mm.
Rostrum and palpi black. Antennae with sca,pe light brownish yellorv, sparsely
pruittosc ; pedicel brownish yellow; flagellurn broken. Head dark brown, pruin<xe.
Protrottrtu otrlS'utoclerately rrrassive,brownish black in front, tcstaoeousyellow behind.
illesonotal praescutunr atrd scuturrr almost uniforurly light fulvous yellow, without distincL,
rnarkings; praescutum rvith anterior end narrowed and slightly projecthg; scutellurrr
slightly more infuscated; mediotergite pale browrr, pleurotergite chiefly occupied by *rnajor subcircular black slmt, its cephalic part yellow. Pleura light yellow, the anepisteltrum chiefl-voccupied b.y another black spot, a trifle srnaller than t,he one on the pleurotcrgite; dorsopleural metnbrane weakly brownish yellorv. Halteres with stem yelkrw,
krrob darkened. L"gt with the ooxae and trochanters light yerllow; rcmainder of legs
light brown. Wiugs rvith a strong fulvous brown tirrge, the prearcular and costal fields
a, trifle darker; stigrna strrall, pale brown, only a little darker than the costal borders;
veiitts brownish yellorv. \renation : ,Sc1ending shortly beyond fork of .8s, ,So2a little
t'etnovedfrotn its tip; ,Il2 very faint, a,trifle shorter than 1lz+r; veins .Il, and r?nstrongly
clivelgent at tips so cell R" at rnargin is tnorB extensive than cell .Il2 and nearly fivo timcs
cell Rr i nl-cu just bcyond frlrk of M ; anterior ar.culuspreserved.
Abdominal tergites da,rk brorvn; sterrrites obscure yellow. Valvcs of ovilxtsitor
clotrgate, irorn-vellow.

Holotype,?, Suoerv-IITHIOpIA: Kende,AlouraPlateau,13.xii.1938(J. G.
ll'yers); Collector'sNo. L0,223.
By *y key to the African speciesof the genus(loc. cit. 16 (18): G7) the,lrlexander,of South Africa (Southernl)resentfly runs most nearly to a,u,rantaca
Rltodesia),whieh seemsto be thc closestdescribedally. The presentinsect
tliffers frorn all generallysinrilar speciesby the very oonspicuous
paired blaok
spotson the sirlt-,sof the thorax.

perpicturata(Trentepohlia)
Trentepohlia
sp. n.
(icnelal cololation yellow ; knobs of halteres darkened ; femora yellow, the tips paling
to yellorvish whil,e ; rvings whitish, u.ith t,hree brown crossbands,the most basal one n&rrow,the others very broad, r'ariegated by spots of the ground colour; abdomen bicoloured,dark blorvn, ringed rvith yellow, the subtenninal segmentsunifonnly blackened.
tr'enrule.-Length about 6 rnnr. ; wirrg ti mrn.
Il,ostrum a,rrclpalpi black. Antennae black t,hroughout, relatively elougate. Headchiefly tlark in colour.
'I'horax ahnost unifonnly yellow, tho
1-rleuraa triflc lllole tlarkened iu the dorsoplourallegion. Halteres pale, the knobs darkened. Legs ;rellow, the femoral tips bloadly but
inconspicuously paling to yellorvish white ; tibiae and tarsi dirty whitish. Wings with theglound colour whitened, the prearcular and costal portions more yellowish; a heavy
brown crossbanded pattenr that is more extensive than the ground ; basal band n&rrow,lying shortly beyond the arculus, extendirrg frorn ,ICto the anal angle, including the outerlralf of cellznd.,{, slightly interrupted in cell Cu ; secondband very broad, almost quadrate
in outline, lying chiefly before the cord, extending completely across the wing but paler,in the cubital and anal fields, r'ariegated by pale greyish areas in cell Bc1, beyond vein Bc2
on basal haH of cell .81 and near the outer end of cell r? ; outer band subequal in extent to
the last, invoh-ing the entire wing tip back to the level of the base of cell .Eu,variegatedtry an extensive ground arca in ccll 11, and a suraller one near the outer end of cell .Bu;

88

Prof. C. P. Alexandero'nneu)or little-hnwsn'

veins pale yellow, darkenedin the brown areas. Venation: -Rslong, arcuated at' origin,,*.."ding .h, bot, shorter than -Bn; veins Rr+r1a and /ls +r meet'ingat an angleat pointof departure of R2 ; m'cu shortly beforethe fork of M 'A-bdonenbicoloured,dark brown, segmentstwo to six, inolusive, rvit,h a conspiouousbasal yellow ring that likewise involves the posterior border of the precedingstenrite;
subternrinal segmentsuniformly blackened.

Holotype,?, ceupnouN: sakbayeme,1.viii.1931 (J, A. Reis).
is vgly d.iffcrent fronr the now
Trenti$ohlia (Trentepohtia)perpichmata-th6subgenus. The pattertr of the legs.andlrumerousEthiopian speciesof
especiallythc u.ry heavy wing pattern readily sufficeto separatethe fly fromall such allies.
sp. n.
malagasica
Erioptera(Podoneura)
Very silrilar to the type of the subgenus, untltrarcgr&nxrru(Ilergroth), differing csltcciallyin the structure of the tnale hypopygium.
Male hypopygiuur with the outer dististyle blackened, nearly straiglrt, tltc stettt stout,
tcrrni'atitlg i" u Jt"ong straight spine, with a smaller a,pprcsse(llatelal spine orl outer face ;'uter surface of style in the vicinity of the latcral spine rvith a ferv ttticroscoPicdenticles
and sparseerect seiulae. Inner dististyle shorter, at apex rrarrorvedinto a st'rotrgblackened
spirrei Gonapophysis appcaring a,s a curved blackened holn. lrr utttltrucognnn'trutthetl-ististyles *1a gori"pophyr"r arc slightly different, cspt'cially the ittttt'r distist'yle, rvhich isbroa<lly obt,useat aPex.

: Vinanitelo.
Holotype, $ on microscopeslide, MEneceSOAR
Toxorhina(Toxorhina)taeniomerasp. rr.
Size srnall (rving, rnale, less than 4 mm.) ; general coloration of b<ldy grelr t,he Praegcuturn and scuturn patterned rvith darker ; rostlum, lralteres and legs blackened; rvings
whitish subhyalile, unpatterned I veins blou'nish black, conspicuors i nl-ctl beforo ttre
fork of 17l; *bdom",,, including hypopy-liurn, uniftrrrnl.y dark browtt ; rnale hypopygiunr
with the dististyle profoundly divided to frtrur trvo sePalate arms or blanches; aedeagus
rvith the &rms .rn.,slo*ily long and slender, cach exceeding the total length of the courbined
btr,sistyleand dististYlc.
lI ale.-I'ength, ex<;luding ro8truut, about, 4 tttur. ; rvilg 3'tl urttt. I rostrutu about
9'6 mm.
Female.-I'ength, excluding rostrutlt, about 5'5 tttm.; wiug 4'5 tuttt'
Rostrurn bhJk, elongate, about two-thirds thc length of rving. Atrtenttac blackt,hethroughout; flagellar segrnentsoval to short-cyliudlical. Ilead dark leaden gre.v,
broad anterior vertex more silvery grey.
pronotum and cervical region brownish grey. Mes<lnotulngrey, the praescubumwith
three poorly defined darker stripes that, are conflue,nt,bchincl; scutal lobes similarly
tlarkeried. pleura uniformly dark grelr including the dolsopleural trtt'nrbrattt'. Haltertsbrightened. Legs rvith the coxae darkbrownish black, the base of rt"rtr-t.rtti"t"dly
theplumbeous grey; trochanters dark brown; r'emainder of legs brorvnish black, onlyveins
femor.al bases a trifle mone brightened. Wings whitish subhyalinc, unpattemed;brownish black, conspicuous. Venation : Cell lst M 2large, rn tnuch shorter than the basals e c t i o no f . M r; i n o ri e w i n g o f typ e, cell2nd,M ewithashor t,petioletofor r nattel em ent
m:cuin male almost its own length before the fork of M, in fernale much closer
Llr*,tati
to this fork, the distance slightly variable.
Abdomen, including hypopvg,um, uniforrnly dark brown. Male hypopygium rvith theprirnaryd.ististyle profoundly air.ia"a to?ot- a second branch or inner style at basc of the"r*, thu Ltt", ",r Llorrgo,te relatively slender blade, gradually narrou'ed to the obliqur:
about,*n"*, on basal ha11of oot"" margin with a high flange ; inner branch or style only
Aedeaoie-third the length of the primary one, appearing as a slender, gently curved rod.

speci,,esof erotic Tipulid,ae(Diptera)89

gus with the arms unusually long and slender, gradually narrowed and.sinuous,
the lengthof either a,rm exceedingthe total combined tength of ihe basistyle and dististyle.

i .Tolotf,pl, 6, FnnNcuEeuetoRrAl Arnrce : Brazzaville,'a.Z.ii.lg3g (J. A.
Reis), Alloty'pe,?, ceuERouN Moryjipome,20.i. lg3g (J .
nrir1.
,
present
Tlu
fly
closestistoTonorhiia(Tonorhina)
cisitlanticaSpuir*r,likewisefrom the C_ameroun,
which is still known to me only from tL originaldescription.- This differsfr9-- thefly in severalimportant regirds,
as in the colorationof the body and_present
legs,and in the whitenei h"lturur. The'elongatearmsof the aedeagus
ieem to 5e-acharacterof both of theseflies.
Styringomyiasubimmaculatasp n.
Size small (wing, male, about 4 mm.); genera,lcoloration obscure yellow,
without'distinct pattern ; legs obscure yellow, otrpu,iturned,
only the terminal tarsal segment
black; wings greyish yellow, the dark pattern very restricled, best indicatedby a darkening of t'he r-m crossvein; male hypopygium with the spine of ine basistyle tong
and black,'approximately twice the lobe; setae of outer dististyle threeor four in number, very
weak ; ninth tergite a broad cushion with a further .rurro.o median Iobe onouter portion.
Male.-Length about 6.b mm. ; wing 4.1 mm.
Rostrum and palpi pale brown. Antennae pale yellow, the basal segmentsa trifledarker ; verticils long and conspicuous,black. Head pale brown, with erect
black bristles.
Thorax above chiefly obscure yellow, without distinct pattern. pleura yellow.
Haltges weakly infuscated. I*g. with the coxae and trochunl"., yellow ,urnlir,der
of
legs
;
obscure yellow, without distinct pattern on any_of the segments,the terminal
tarsal segt:lt
abruptly black. Wings pale greyish yellow, the prearcular field a little clearer
yellow; no dark pattern excepting a va,guecloud ou", i-*; veins yellow, r-m,
fork ofM r * 2;rt?,-crl
and the tip of vein znd A a trifle darker. Venation : Cell ZnAU
,short-petiolate ;tn'cu about its own length beyond the fork of trI; vein znd,A curved grttity into
the anal
margin.
Abdomen pale brownish yellow, without distinct pattern ; hypopygium yellow, with
P*"tt of the styli and phallosome blackened. Male hyryopygium with the ninth tergito a
broad cushion, clothed with abundant recurved spinouss"tiu median portion of cushion;
further produced into a narrow median lobe. Uintn sternite a,narrow p"l" lobu thatbears
two stout blackenedsetaeat apex. Basistyle with the terminal spinesinigle,long and black,approximately t'wice as long as the lobe. Outer dististyle elongate, pal6, the ifex weakly
darkened, bearing a single powerful seta at apex ; setae alon! facJ of style #eak, threeor four in number. Middle dististyle much smaller than the inner one, bearing thrreo
concentrated groups of long peg-like spines, two of these marginal, the tlfrd ptaciealower on
the face of style, all groups with about a d.ozenspines "u.h. fnner dististyle a, very large
expanded blade, the outer apical angle produced.into a slenderarm that bearstwo strong
spines; margin of style, beginning at base of this arm, with a row of long spinous sebao,-rpu*
totalling about twenty, the more basa,lones much more slender and.weak"t ;of style
laterad of the arm with a further closecomb of about fifteen blackened spines iasal "rrgl,;
of style produced into a small acute point. Phallosomewith the apex consisting of two
blackenedcultrate blades,the upper margin with a single strong spine.

.Holotype,d, CeunnorrN: Momjipome,along stream, 20.1.lggg (J. A.
Reis\.
fly is readily distinguishedfrom all other similar regional
The _pres-ent
spe_cielsby the virtually unpatternedlegs,unusually restricted wing pattern,
and the structure of the male h_ypoplgium. fn somerespectsit a[rees with
Styr-ing-ornyia_impunctata
Edwards,of northern Ngeria, differing in tf,e patternof the legs,the venation,and in the structureof the hypopygium.