-.,

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I

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AC/

1 * . . ?

ITEW SPECIES OF CRANE T'LIES FROM SOUTE
AMERICA. PART V.
(Famity Tipulide, Order Diptera.)

BY

Cgerr.rg

P. Ar.trxANDER.

l'

IEPl

l.lED

FBOII

Annets oF rsr Exmrf,or.ocrcer,Socrrrv or Aurnrce
Vol. XXIV,

No. 3.

:.
::

Cowraus, OEro,
Scptgbcr, r9ifr.

NEW SPECIESOF CRANE FTIES FROM SOUTH
AMERTCA. PART V.*t
(Family Tipulida, Order Diptera.)
CulRLss P. Ar,sxenpBn,
Amherst, Massachusetts.

The majority of the species herein described as new werecollected in Colombia by Mr. Fred W. Walker, chiefly at Vista
Nieve on Mount Santa Marta. A11such resulting types havebeen returned to the Museum of. Zoology of the University ofMichigan. One additional specieswas likewise taken on Mount
Santa Marta by my friend, Dr. George Salt. The other species
considered are from Brazil', collected by the veteran naturalist,
Mr. Friedrich Schade, and from Chile, where they were taken in
the Province of Malleco by Professor D. S. Bullock. Except
where indicated above, the types of the novelties are preservedin my own collection. I wish to thank the gentlemen above
mentioned very sincerely for this appreciated co-operation in
making known the rich Tipulid fauna of the Neotropics.
Macromastix Osten Sacken.
Macromastix (Araucomyia)bullocki sp. n.
General coloration shinv yellow, the praescutum withreddish stripes;antennaeof both sexesvery short, lO-segmented;
femora brownish yellow, the tips narrowly brownish black;wings grayish subhyaline, the stigma and a confluent seam on
the anterior cord dark brown; abdomen yellow, lined longitudinally with black, the latter areas very extensive on thesternites
Male.-Length, about 11 mm.; wing, L2 mm.
Female.-Length,about 15 mm.; wing, 14-74.5mm.
Frontal prolongationof head obscureyellorv,darker on sides,thenasuselongate;palpi black. Antenne l0-segmented,unusually shortin both sexes,a little longer than the head, excludingthe frontal prolongation; first scapal segment brownish-yellow, the second clearer*Contribution from the Entomological Laboratory, Massachusetts State
College.
tThe preceding part under this general title was published in these ANNnrs
in 1930 (XXIII: 72Lqn).

1e311

Alexander: South American Tipulid,a623

yellow; flagellum black; flagellar segments elongate-cylindrical, theverticils of the basal segments short, on the outer segments becoming
more elongate, exceeding the segments. Head pale brown above, theposterior orbits clear yellow.
Pronotum light yellow, restrictedly infuscated medially. Mesonotal praescutum nitidous, deep yellow, with four narrow dark reddish
stripes, the intermediate pair blackened on their anterior portions;median region of scutum yellow, the lobes more reddish; scutellum
yellowish-testaceous; postnotal mediotergite yellow with two reddish
brown longitudinal areas. Pleura nitidous, yellow, variegated withreddish areas. Halteres pale, the knobs infuscated. Legs with thefore coxe yellow, lined with reddish, the other coxa darker; trochantersbrownish yellow; femora brownish yellorv, the tips narrowly brownish
black; tibize and tarsi obscure yellow, the latter passing into darkbrown; claws (d) with a basal tooth. Wings grayish subhyaline,
cells C and ,Scmore yellowish; stigma and a connecting cloud on theanterior cord dark brown; wing-tip vaguely infumed; a dark seamin cell M alongvein Cu; veins black, more yellowish in the costal region.
Macrotrichia of veins small and inconspicuous. Venationi Rr+ediverging strongly from R3, paralleling the free tip of Scz; cell Mr withthe petiole longer to shorter than m.
Abdominal tergites obscure yellorv, u'ith a narrorv median black
vitta; sternites similar, trivittate rvith black, the median stripe very
wide, restricting the ground-color to narrow sublateral lines; hypopygium
obscure yellow. Male hypopygium with the lateral lobes of the tergite
very long and slender, separated by a deep notch; the fused basistyles
and sternites similarly long and slender. Ovipositor with all valvesvery long and slender, straight.

Haslrer: Chile. Holotype, d, Nahuelbuta, near Angol,
Ma1leco, among Chilean Pines, Araucaria imbricata Pav.,
altitude 800 meters, March 9, 1929 (D. S. Bullock). AllotopotyPe, 9, altitude 1,200meters,March 10, 1929. Paratopotype,a teneral g, with the allotype.
Macromastix (Araucomyia) bullocki is named in honor of
the collector, Professor Diltman S. Bullock. The speciesis
very different from M. (A .) paulseni (Phil.) in the short antennaeof the male and all details of coloration and structure. The
polished yellow and rusty coloratronpolrshedcoloration otof the thorax is verysuggestive of certain species of.Nephrotcma.

Tipula Linnaeus.
Tipula laterosetosasp. n.
Ground-color of mesonotum obscure yellow, the praescutumwith four brown stripes that are narrowly bordered by darker
brown; antennae short in both sexes, the flagellum bieolorous;

624

Annals EntomologicalSoc'ictyof Amer'i,ca[Vol. XXIV,

pleura variegated yellow and brown; femora with a broad
yellowish white subterminal ring; tibiae brown, the bases
narrowly pale yellow; wings grayish brown, variegated by darkbrown and cream-yellowareas,the latter including a W-shaped
mark acrossthe outer endsof cellsR anclM, and a seriesof small
marginal spots in the medial to anal fields; male hypopygiumwith a lateral pencil of yellow setae on either side of tergite;
ninth sternite bearing two approximated cylindrical lobes on
the mid-line.
Male.-Length, about 21 mm.; wing, 21.5 mm.
Fernale.-Length, about 25 mm.; wing, 24 mm.
Frontal prolongation of head brownish yellow, the nasus elongate,concolorous; palpi brownish black. Antenna relatively short in both
sexes;scapeyellow; flagellar segmentsbicolorous, the basal enlargement
brownish black, the remainder yellow; on outer segmentsthe coloration
becomesmore brownish yellow;verticils much longer than the segments.
Head buffy-yellow, on the posterior vertex trivittate with stightly
darker brown, including a median vitta and a similar line on the posterior
orbits; median region of anterior vertex produced into a small roundedpapilla.
-Mesonotal praescutum obscure yellow, with four light brown
stripes that are narrowly bordered by dark brown; humeral region
yellow, enclosing a small black marginal spot; posterior inter-spaces
appearing as elongate pale yellow triangles; a shallow crescent-shaped
brown area at suture; scutal lobes rvith centers brorvnish testaceous,encircled with dark brolvn, the median area testaceous,variegated bybrown behind; scutellum dark broln, the parascutellasimilar, rvith apale central spot; postnotal medio-tergite gray, rvith a capillary brown
median line that becomesmore expandedon the caudal margin. Pleura
light yellow, variegated by transverse brown areas on sternopleurite
and anepisternum, the latter broader and forming part of a more orless intermpted longitudinal stripe extending from the cervical sclerites
to the postnotum; a small velvety-black spot on the dorsal pteropleurite; pleurotergite dark brown, the elevated ventral portion densely
set with golden-yellow tomentum. Halteres dusky, the base yellow,
the knobs obscureyellorv. Legs with the coxrc and trochanters yellow;
femora yellow basally, passinginto black on the distal half, this darkening enclosing a pale yellow subterminal ring; tibia with the extreme
bases yellow, the remainder dark brown; tarsi dark reddish brown.
Wings with the ground-color grayish brown, variegated with darkbrown and cream-colored areas; prearcular region chiefly darkened,
separated from a more extensive post-arcular area by clear vellow at
arculus; no dark spots along cell Sc, as usual in the glaphyropteragroup;
stigma darkened; small dark areas along cord and at near mid-lengthof vein Cu; the chief creamy areas appear as a W-shaped mark lying
transversely across cells C, Sc, R and M before the cord; the distal
fourth of cell Rsi a narrow, irregular line beyond the stigma; small

1e311

Alexander: South American I'ipul,ide

625

F?rgtnal spots in outer ends of cells pnd.M2, Ms, Il[a and,Ist A, theteb"-t-"g-twoin the last-namedcell; and extensive'^r"i i"-t"r"f half ofcell cu, extending.into cell M; veins dark, pale in the cieamy areas.
veaation:T1a2 entire; Ra;.5llSually sinuous,deflectedcaua"E o" outer
half, cell .Rsat margin being unuiually wide.
Abdominal tergites chiefly blackened, the incisures of the basal
segmentsmore cinnamon-brown;outer segmentsmargined lateraitywith cinnamon-brown;basal sternites reddish Ito*",'r"gment four
chiefly darkenedl.ou.telsegmentsdarkened,except at marfins. Male
hypopygrum_relatively
large,the tergite fusedwith ttre sterni?e. Tergiie
with the medianregion prbducedinlo a depressedlobe, iir "p.* wifh anotchlextrememarginnarrowlycrriiintef andpro-

lLtg* atYTh,"p.9
duced
lateral angles-into small v-entral spinLs; lateral region of
tergite bearing a tuft of. long yellorv sete. ilegion of basistlle notdifferentiated from sternite ei_c-gnt
by a short st"raight."i"t"'Ulf"*iapex of.basistyletruncated.
Wtie! appeatsto be . Jotio.tsiy modifieciouter dististyle is borne at endof baiiityle, appearingas a iale membranous lobe that is prolongedventrad into'i ribb6n-iit" -extension.
Ninth sternite bearing two-conspicuoussubcylindticat-toles,closely
applied to one-anotherto appeaf almost ". " singlestructure. ovipositor rvith all r'al'u'esrvell-developed,the tergal ialves fr"iry except
on distal third; sternalI'alvescompressed,l,ellow.
Haetrar:Colombia. Ilototype, &, \rista Nieve, X,{t.Santa
Marta, altitude about b,000feet, August g, 1926 (F.w. Walker).
Allotopoty/e, g , August o, L926. Faratopotype, a damaged d,August 7, 1926.
Th9 general resemblance of this fly to various species ofthe.oblique-fascigta group is probably superficial orriy. The
species is well-distinguished by the peculiar structure of themale h.vpop1.gium.
Limonia lleigen.
Limonia (Rhipidia) tripectinatasp. n.
Belongs to the maculata group; allied to inaequipectinata;antennae (e) elongate; flagellar segments 2 to 11 unequallytripectinate, there being a very smalr median spur, in addition
to the elongate branches; wings with four dartene,C clouds inthe subcostal field, in addition to the stigma; male hypopygiumwith the rostral prolongation of the ventral dististyie long andslender, with two spines that arise close together.
Male.-Lgngth, about G mm.; wing, Z mm.
Female.-Length,.qlout2.5-Smm.; wing, 7.5.
Rostrum and palpi black. Antenna ('distancta ) iiery largeand conspicuous,
if bent backward extending to someU.-yona ttr"- l"r" of626

of Americo [Vol. XXIV'
Soc'i'ety
Annals Entomol,ogi,cal

abdomen;scapeblack, the flagellar segmentswhite, the basal_swelling
and all branclies black; terminal segmententirely black; branchesyery
long and slender, the longest approximately one-third the length of theentire organ; in addition to the long branches, there arises between
them a smali spur, forming a very unequal third branch; branch of firstflagellar segment single; segments2 to 11 unequally tripectinate, asdeJcribed, the branches of outer segments gradually decreasing inlength; terminal segmentelongate, a trifle shorter than the branchesofthJ penultimate segment. Antenna ( Q) neaTly s_1pqle,the -flagellar
segmentswith very short, inconspicuousapical pedicels;terminal segment elongate. Head dark gray.
Pronotum black. Mesonotal praescutumreddish brown, the color
broadened behind, the lateral and humeral regions extensively moreyellowish; median region of praescutumnarrowly darker-brown; scutumieddish brown; scutellum darker brown, paler medially; postnotum
dark brown. Pleura pale grayish brown, with two vague darker
longitudinal stripes, the more dorsal stripe ending at the pteropleurite,
thJventral strip-eextendingfrom the fore coxa to the baseof abdomen.
Halteres pale ye1low, the knobs weakly infuscated. -L.gt rvith thecoxe yeflbw, darkened basally; trochanters yellow; femora yellow,
narro*ly and vaguely darkened subapically; tibire brou'nish yellow;
into darker brown. Wings rvith the ground-colorcreamy,
tarsi paising^arestlricted dark pattern, this consisting of four diffuse darkwithclouds in the subcostalfie1d,in addition to the stigma; restricted pale
brown seamsalong cord and outer end of cell lst Mz; wing-tip, especiallyin the outer endslf the radial cells, darkened; veins yellow, darker inthe clouded areas. Venation: Sc long, Scr ending about opposite
two-thirds the length of Rs, Sczatits tip; Rs gently arcuatedat origin;
free tip of ,Sczand Rz in approximate transversealignment; m-cu at
f.ork of.M.
Abdominal segments obscure yellorv, the caudal margins -9onspicuously ringed with dark brott'n; hypop)'giuT obscure yellow.
Male hypopygium"itrd -riedianwith the tergite transverse, the caudal_m.argln
region tranwerse or feebly conca\-e. Basistyle
convex,to anmoderately large, the ventro-mesallobe stout basally,_narro'lved
dististyle
Ventralobtuse point tliat bears a singlelong powerful seta.
of mod6ratesize,a little larger than the basistyle; rostral prolongation
long and slender, at about two-thirds the length bearing two long
slenderspinesthat are placedclosetogether. Dorsaldististyle elongate,
curved and narrowed tb a long acute apical point. Gonapophysiswith
the mesal-apicallobe blackened, obliquely obtuse at !ip, separated
from the shorter outer lobe by a narrow U-shapednotch.
Colombia. Holotypr, d, Vista Nieve, \'It. Santa
Hesrrer:
Marta, altitude about 5,000 feet, August 7 , 1926 (F. Ur. Walker).
Allotopoty/e, Q, August 7, 1926. Paratopotypes,S e J, 10 I 9,
August 6-8, 1926.

1e311

Alexander: South American Tipulid,e627

Limonia (Rhipid'ia) tripectinata has an antennal structurethat is much like that of. L. (R.) ineguipectinata Alexander,
which diflers conspicuously in the smaller antennae, the wingpattern and venation, and in the structure of the -al"hypopygium.
Limonia (Rhipidia) mystica sp. n.
Belongs to the uniseriata group; general coloration yellow,
the mesonotal praescutum with a broad, median brown stripe;
antennae entirely black; flagellar segments 1 to 10 with asingle branch; thoracic pleura yellow, with two narrow black
stripes; halteres black; wings whitish, with a conspicuous
brown and gray clouded pattern; cell Mz tending to be open byatrophy of the basal section of. Ma; male hypopygium with th;spines of the rostral prolongation of the ventral dististvle
short and spike-like.
Male.-Length, about 5 mm.: n'ing,G mm.
Rostrumand palpi black, the former about as long as the remaind.erof head. Antenne black; basalflagellarsegmentsniih a singlebranch,
the formula being 2-+10+2; .longestbranch (about flagellai segments
7.t9 9) approxim?lelythree timeJ the segment;basalfligellar segmenr
with a stout basalbranchthat is shorterihan the segment;penult"imate
segment-simple, ,or _nearlyso; terminal segmentelongai6,narrowed
outwardly. Head black, sparselypruinose.Pronotum yellow, narrowly dark brown medially. Mesonotal
praescutumbuffy 1'e]low,with i broad dark brown medianstripe th;;is a trifle expandedlaterally at--nearmidlength; lateral stripesiacking
or representedonly by_" small circular dusky cloud at the suture;scutum_yellorv, each lobe rvith_-atriangular brou'n area, the point
directedcaudad onto the scutellum;scutellum]'ellorvishtestaceous;postnotalmediotergitechieffydark brorvn,pale1'eliorv
laterally. Pleura
yellow, with two narro\\' black longitudinal itripes, the dorsal one
extendingfrom the cervical scleritesacrossthe dorsal pleuritesto thehalteres;ventral stripe less-distinctand more broken, including thedorsal sternopleuriteand the meral region. Halteres short, b"lack,the extremebase-of the stem yellow, the knobs large. Legs with thecox& yellow, slighth' darkenedacrosstheir bases;tlochantersyellow;
femora brownishyellow, vaguelv darkenedoutwardly; tibia and tarsi
pale b_rown,the terminal tarsal segmentsblackened;posterior legs
lost. Wings whitish, more cream-coloredin the costai r^egion,with"aheavy brown costal pattern and abundant gray clouds"that covermost of the remainderof the wing; dark cloudd six in number, thefirst two confluent or nearly so; fourth area at origin of Rs; fitfh atfork of Sc, the sixth at stigma; the gray cloudsinclude the cord and amore or lesscompletesubapicalfascia; medial, cubital and anal c:l1s
more evenly cloudedwith gray; the ground-colorappearsat the outer628

Annals Entomological Society of America [Vol. XXIV,

end of cell M, basesof Anal cells and as a smaller area in cell Ist A at
end of vein ?nd, A; veins cream-yellow in the ground-areas, more
infuscated elsewhere. Venation: Scr ending about oppositethree-fifths
the length of Rs, Sczat its tip; Rs long, gently arcuated; R2faint, subequal to the free tip of Scz,'cell Mz open or tending to be open by theatrophy of the basal section of.7[s; m,-cucloseto fork of"M. In one*ittg of the type, the basal sectionof Ma is entirely atrophied; in theother wing, it is indicated by a vague trace.
Abdominal tergites yellowish brown, the caudal margins narrowly
blackened; sternites more uniformly yellow; hypopygium brownish
yellow. Male hypopygium with the tergite transverse, its caudal
margin straight or very gently concave. Basistylesmall, with a large
ventro-mesal 1obe. Ventral dististyle considerably larger than thebasistyle, the rostral prolongation slender,with two small spike-likespinesat near midlength, theseplacedclosetogether. Dorsal dististylea slightly curved darkened rod, slightly dilated outwardly, the tip
suddenly narrowed to a slenderpoint. Gonapophyseswith the rnesal
apical lobe darkened, the outer margin roughened,the apex a more
slenderblackenedpoint.
Colombia. Holotype, d, Vista Nieve, \It. Santa
Hasrret:Marta, altitude about 5,000 feet, August 7 , 1926 (F'. \\-. Walker't.
The closest relative of the present speciesis Z. (R.) pallatanga Alexander (Ecuador), which differs in the larger size,
details of coloration of the antennae, body and wings, and themuch shorter flagellar branches.
Gynoplistia Westwood.
Glmoplistia (Gynoplistia) vaiata sp. n.
Generally similar and most nearly allied to G. (G.) manicuta
Alexander (Chile), differing most conspicuously in the smaller
size and absence of blackened femoral tips.
Male.-Length, about 17-18mm.; wing, 14.5-15mm.
the formula being 2+2+8+6; scape
Antenna (c/) l8-segrnented,
yellow,
the branchesand basalenlargeflagellar
segments
and apicesof
ments of the segmentsdarkened,the outer segmentsmore uniformlv
darkened; longest branch (about flagellar segment7 or 8) about fivetimes the segmentand shorter than the combinedlength of the simple
terminal flagellar segments;last branch shorter than to approximatelrtwice the length of the segment. Head brown.
Mesonotum brown, with a yellowish pollen, the praescutalstripes
only feebly indicated. Pleura chiefly covered by_ a silvery-yellorv
bloom, the ventral sternopleuritemore glabrous. Halteres pale, theknobs infuscated. Legs with the femora brownish yellow, the tips
weakly darkenedbut not at all blackened,as in manicata;tibia. yellorv,
the tips more narrowly darkened. Wings pale yellow, the stigma a1e3lJ

Alerander: SouthAmerican Tipul,ide629

very little darker; in the holotype with a few grayish brown blotches
in cells M and Cu near the cord and larger onesin the ends of the Analcells, including a larger area in cell lst A that almost crossesthe cell.
\renation: Much as in manicata,with the anterior branch of Rs similarly
long and sinuous; Rs a little shorter; cell lst fufz slightly narrower,
especiallyon the basal half.
Abdominal tergites grayish brown, the basal sternites bilineate
rvith black; hypopygium yellowish brown. Male hypopygium withthe outer dististyle much as in manicata. Interbases appearing as
nearly simple chitinized horns, with a microscopicpoint on margin atnear midlength.
Hestrer:
Chile. Ilolotype, d, Angol, Mallecb, November
25, 1929 (D. S. Bullock). Paratopotype, d, April 30, lg2g.
Gynoplistia (Gynoplistia) biarmata sp. n.
Generally similar and allied to G. (G.) manicata Alexander,
dii',ering especially in the absence of black femoral tips and thestructure of the male hypopygium, especially of the interbasal
processes.
Itale.-Length. about 23 mrn.; rving, 19 mrn.
Antennrc l9-segmented,the formula being 2+2+9+6; black,the incisuresof the basal flagellar segmentrestrictedly yellow; tips offlagellar branchespaler than their bases;longest branch shorter thanthe combined terminal simple segments;last branch about one-halflonger than the segment;terminal segmentconstricted,as in the group.
Head gray.
Mesonotum gray, the praescutum with three brown stripes, themedian stripe darker in front, weakly divided behind; pseudosutural
fovea bicolorous,reddish r,r'iththe lateral portion blackened;medianregion of scutum and the scutellumlight gray; postnotalmediotergite
yellowishbro\r'n,sparselypminose. Pleuraheavilypruinose. Halteres
pale yellow, the knobs s'eaklv infuscated. Legs with the femora
dark brown, yellorv basally; tibie abruptly yellow, with narrowly
darkened tips; tarsi yellow, the outer segmentsa little darkened.
Wings pale yellow, the oval stigma pale brown; a small brown cloudat end of vein R3; veins brown, more yellowish in the costal region.
\renationas in manicalaand allies.
Abdomen black, pruinose, the segmentsvery narrowly marginedlaterally and caudally with yellow; hypopygium chiefly dark. Male
hypopygium as in rnanicataand allies, but the interbasal processes
very different in form, broad-based,with two large spines,the morebasal larger and bearing a microscopic spinule on outer margin atnear midlength.
Hesrrnt:
Bullock).

Chile.

IIol,otype, f , Angol, Malleco, 1928 (D. S.

630

Annals EntomologicalSocietyof America [vol. xxIV,

The two species described above, Gynoplistia (Gynopli'stia)
aariata sp. n. and G. (G.) biarmata sp. n., together with G- (G-)
gilaipennis Alexander and G. (G.) manicata Alexander, and very
possibly also G. (G.) flavipennis (Philippi), form a compact
group of large Gynoplistia in which the anterior branch of
Rs is long and conspicuously sinuous and the tibial spurs andtarsal claws of both sexesare microscopically toothed.
Eriocera Macquart.
Erioceratholopasp. n.
General coloration black, heavily pruinose; antennae shortin both sexes; head dull oraflB€, darkened behind ; mesouotal
praescutum with three shiny black stripes, the lateral margins
pruinose, the interspaces more yellowish pollinose; legs andhalteres black; wings dusky, the cord seamed with darker;
Re at or before the fork of Ra*5;cell Mrlacking; m-cu at fork of
M; ovipositor very long and slender.
Male.-Length, about 9 mm.; wing, 7 mm.
Female.-Length,about 72 mm.; wing, 8.5 mm.
Rostrum and palpi black. Antenne black throughout, short in(d), 9-segmented( 9 ) ; first flagellarsegment
both sexes,7-segm-ented
nearly aslong aJremainderof flagellum. Head dull orange,the occiput
infuscated;vertical tubercle weakly bifid.
Mesonotalpraescutumwith three shiny_b1+* stripes,the lateral
marglnsprulnose,the interspacesmore yellorvishpollinose;-posteriorscler-itesof mesonotumblack, sparsely pruinose, the scutellum andpostnotum more heavily so. Pleura black, heavily - pruinose, thedorso-pleuralregion slightly more buffy. Halteresblack; the extreme
black;
base of st"m pa1er. LEgswith the coxrepruinose;trocl-r-antersbrownishtibirebrown;
reddishmore
femora black,-the extremebases
black to black; tarsi black. Wings dusky, the costal region darker,
the wing-basesomewhatmore yellowish brown; xaffow _d3rke1brown
seams.Iotg cord and veins Rs and R5; veins black. Venation:Scr
ending opposite the fork of Rs, Scz at its tip; Rs elongate, weakly
angulatedat origini Rz at or just beforethe fork of Ra+r(c/) or morethin its own length beforethis fork ( I ); cell Mr lacking; tn-cuat or
the fork of.M,longer than the distal sectionof'Ca.
beyond-Abdomen
black, graypruinose,moreheavlly so ol the basalportionsof the segments;mJle hypopygiumblack. Ovipositor-with-thetergal
valves exieedinglylong and slender,horn-yellow,blackenedat bases.
He.srrnr : Brazil. Holotype, &, Jaragua, Santa Catharina,
October 10, 1929 (F. Schade). Allotopotype, 9 , October 9,1929.

1e311

Alexander: SouthAmerican Tipulide

631

Eriocera.tholopa is apparently closest to E. flaviceps (wied.),differing in the coloration of the body and wings. The latter
species is still known to me only from the very brief andinsufficient original description.
Elephantomyia Osten Sacken.
Elephantomyia (Elephantomyia) humilis sp. n.
Size sma1l (wing, not exceeding 6.5 mm.); thorax entirely
yellow, immaculate; legs pale brown; wings yellowish subhyaline.
Male.-Length, excludingrostrum, about 5.5 mm.; rving, b.8 mm.;rostrum, about 6 mm.
Female.-Length, excludingrostrum, about 6.5 mm.; wing, O.bmm.;
rostrum, about 5 mm.
Rostrum elongate,subequalto or longer than the body, dark brown,
including the palpi. Ant_ennre_darkbrown, the verticils very long andconspicuous,much exceedingthe segmentsin both sexes. Head very
light gray.
Thorax entirely yellorv,unmarked. Halterespale, the knobsweakly
infuscated. Legs rvith the coxre and trochanters .vellolv, the forecox€ slightly infuscated on outer face; remainder of legs pale brown;a weak tibial spur on hind legs of male, not developedon other legs,
at least of female. Wings yellowish subhyaline, a little darker towald
apex; stigma long-oval, brown; veins brown, those of basal half ofwing somewhat more yellowish brown. Venation: Scz ending shortly
before the fork of Rs, Sc1 atrophied or nearly so; Rs long, gently
arcuated to feebly angulated; branchesof Rs long, extending generally
parallel for their entire length; cell lst Mz short-rectangular, shorter
than any of the veins beyond it; m-cu subequal to or longer than the
distal section of Cuy at or before midlength of cell lst ltz; cell ?nd A
long and narrow.
Abdomen yellorvish brolvn, the sternites somervhatpaler; in thefemale, the basal tergites are weakll' bicolorous, the proximal halfbeing somewhat darker than the remainder; on the outer segmentsthe
color is more uniform brown. Male hypopygium with the mesal faceof the basistyle densely cushioned. Outer dististyle unusually shortand stout, broad-based,shorter than the strongly curved inner dististyle.

Hanner: Colombia. Holotype,d, Vista Nieve,Mt. Santa
Marta, altitude about 5,000 feet, August 8, 1926, at light,
4 A. M. (F. W. Walker). Al,lotopotype,9 , August 6, 1926.
Paratopotype,d, with the allotype.
Etrephantomyiahumilis is even smaller than E. boliz,iensis
Alexander, differing from the latter in the details of bodycoloration and wing-venation,such as the straighter anterior

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branch of Rs, small cell /sf Mz and'more elongate.veinhnd' A.
The paratype is very small and has lost the head, but seems
undoubtedly to representthe samespecies.
angustissimasp. n.
Elephantomyia(Elephantomyia)
Generalcoloration yellow; pronotum and praescutumwith abrown median stripe; femora obscureyellow, the tips narrowly
blackened, preceded by a wide clearer yellow annulus; malehypopygium with the inner dististyle relatively elongate, atrifle longer than the outer style.
I[ale.-Length, excludingrostrum,about 8 mm.; wing, 8.5 mm.;rostrum,about 8 mm.

Female.-Length, excluding rostrum, about I mm.; wing, 8 mm.;rostrum, about 8 mm.
Rostrum nearly as long as the body in both sexes, dark brown.
Antenne with the scapal segments dark brown, the flagellum paler;
flagellar segments becoming more slender and attenuate ou-twardly.
Head yellowish gray, the center of the posterior vertex more infuscated;
anterior vertex leduced to a strip that is a little narrower than the
diameter of the first scapal segment.
Pronotum and mesonotum-yellow, the anterior portion with a darkbrown median stripe that becomes obsolete at near midlength of thepraescutum. Pleuia yellow. Halteres pale, the knobs weakly infusLated, the extreme base of the stem yellow. Legs with the coxa andtrochanters yellow; femora obscure yellow, the extreme tips blackened,precededby-a wider, more diffuse, clearer yellow ring; ttlif and tarsi
yellowish brown to pale brown; tibial spurs present. Wings with ayellofish tinge, the apex more darkened, ceils C and Sc a little more
iuffused; stigma ova1, dark brown; veins dark brown. Venation:,Scrending about opposite the fork of Rs, Scz at its tip; Rf. relatively
short, weakly angul-ated at origin; branches of Rs extending nearlyparallel to one another for their entire length, the upper branch aiittle sinuous beneath the stigma; rn-cu at near midlength of cell lst tr[2,exceeding the distal section of Cur
Abdorninal tergites brownish yellow, with a more or less continuous
brown median line, more expanded on the caudal margins of the segments; subterminal segments more uniformly darBened; sternites
yellow, the caudal margins of the segmentsnarrowly.darkened. Male
hypopygium with the mesal face of the basistyle ]yitlt relatively. few
bilt i""g and conspicuous, erect sete. Outer dististyle relatively
elongatel bifid at ap-ex. Inner dististyle only a. trifle longer than theoutei, nearly straight. Gonapophyses appearing as 19ttg, slender,
sinuous spines. Aedeagus a penefilum of moderate length and stoutness.

Colombia. Ilolotypo, &, Vista Nieve, Mt. Santa
Hesrret:Marta, altitude 5,000 feet, August 6, 1926 (F. W. Walker).
Allotopoty|e, I .

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Elephantomyia angustissima is distinct from all describedregional species in the pattern of the legs and structure of themale hypopygium.
It is closest to E. westwoodi o. s. of the
Nearctic fauna, differing especially in the details of structureof the hypopygium, especially of the gonapophyses.
Teucholabis Osten Sacken.
Teucholabis (Teucholabis) marticola sp. n.
General coloration black; mesonotal praescutum yellow,
with three nearly confluent blackish stripes; scutellum yellow;
knobs of halteres infuscated; femora yellow, the tips blackened;wings subhyaline, the small stigma pale brown; malehypopygium with the apical spine of the basistyle very reduced;
outer dististyle simple, terminating in an acute blackened spine.
Male.-Length, about 4.5 mm.; wing, 4 mm.
Rostrum blac!, nearly as _long as the remainder of head; palpi
black. Antenne black throughouf; flagellar segmentsoval, decieasirig
in size outwardiv. Head black, sparsellrpruinose.
Pronotum obscurevellorv,narrowly bla'ckenedlaterally. Mesonotal
praescutum shiny yellorv, the disk chiefly covered bv three blackish
bloryn stripes that are nearl-vconfluent, th" median itrip. paler thanthe laterali; scutal lobes blaikened, the median region obr.nr" yellow;
rc_utellum p_aleyellow; postnotal mediotergite dirk reddish browr.
Pl9.1fq "ht94y black, the ventral sternopleurite and meral region more
reddish. Halteres pale, the knobs infuscated. Legs with the coxe
reddish b19wn, the trochanters more yellowish; femora yellow,-legs; thetips broadly blackened, the amount nearly equalon alltibia-Wingsyellowish bro\r'n, darker apically; tarsi broken.
subhyiline,
the-Venation:
small, subcircular stigma pale brown; veins pale brorvn.
Scr e-ndingabout opposite one-third the length of the long arcuatedR-s,,Scz-opposite
the origin of the latter; Rr lying just beyond the levelof the basal section of ,Rii Rr+zshort, the tip subobsolele;cell lst Mz
long, widened outwardly, nt longer and more arcuated than the basal
section of. Ma.
Abdominal tergites dark brown, the caudal margins of the segments
narrowly pale; sternites more uniformly pale; hypopygium yell,owish.
Sternal pocket relatively simple. Male hypopygium *iitr the basistyleshort-, the apical spine very tiny. outef distiityle a simple slenderrod that narrows to a slendersmooth blackenedspine. Inner dististylesmall, vgry strongly curved, the tip blackened, narrorrl'edto a slighily
decurved point.
Heener:Colombia. Ilolotype, &, Mt. Santa IVIarta,
November 1, 1926, at light (G. Salt).
By means of my key to the American species of. Teucholabis(Trans. Amer. Ent. Soc., 40: 235-239; 191a), Teucholabis

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marticola runs to couplet 24, disagreeingwith both included
species, tristis Alexander (Peru) and molesta Osten Sacken(Mexico), in the coloration of the thoracic dorsum, infuscatedhalteres,and details of venation.
Gonomyia Meigen.
lemniscatasp. n.
Gonomyia(Lipophleps)
group;sizesmall (wing, d, 3.5 mm.);
Belongsto the pleurali,smale hypopygium with the dististyle terminating in a short
blackenedspine; on outer face of style at near mid-length witha small spike-like spine that is surrounded by numerous erectsetae.
Male-Length about 3 mm.; wing, 3.5 mm.
Rostrum and palpi black. Antenne with the scape and basal
segmentsof the flagellum orange-yellow,the remainder of the flagellum
brown. Head chiefly pale yellow.
Pronotum and anterior lateral pretergites rvhitish. Mesonotal
praescutum fulvous-brown; scutal lobes similarly colored; medianregion of scutum and the scutellum more yellowish, with a continuous
capillary brown median line; postnotum yellow. Pleura yellow, with
two narrow brown longitudinal stripes. Halteres pale, the knobs
weakly darkened. Legs with the fore coxre darkened, the remaining
coxa yellow; trochanters yellow; femora light brown, darkening to anarrow subterminal ring, the extreme apex again pale; tibie and tarsi
pale brown. Wings with a faint brorvn tinge, the base and costal
region clear yellow; stigma ot'al, dark brorvn, as in the group; cells oneither side of the cord variegated by pale creamy areas;veins pale, thecord and outer end of cell 1st trfz darker. \'enation: ,Scrending a shortdistance before the origin of Rs, the distance on Costa being a little
more than one-half of Rs; anterior branch of sector pa1e, strongly
curved cephalad around the stigma; veins Rs and Mt+z approximatedat wing-margin.
Abdominal tergites pale yellow medially, margined caudally andlaterally with brown; sternites more uniformly pale, margined caudallywith brown. Male hypopygium of the general structure of recu.rvato,differing especially in details of the dististyle. Outer fleshy lobe ofbasistyle relatively stout; spinous lobe of basistyle pale basally, narrowed to an acute point. Dististyle extended into a short, blackened
spine, immediately before this with a small fleshy lobe bearing two
fasciculate sete; outer face of style at near midlength bearing a small
spikeJike spine that is surrounded by nrunerous erect seta. Gonapophysesof both sets almost as in recuraata,the elongate arrns d.elicately
fringed with sete along mesal edge; apical lobes of median phallosomic
structure pale, elongate, gently divergent.

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635

Hasrrer:
Colombia, Brazil. Holotype, & , Vista Nieve,
Mt. Santa Marta, Colombia, altitude b,000 feet, August 8,
1926,at light, 4 A. N{. (F'. W. Walker). Paraty/e, l, Jaragu&,
Santa Catharina, Brazil, September20, l92g (F . Schade).
Gonomyia(Lipophleps)lemniscatais allied to G. (L.) recuraata
Alexander, differing in the details of structure of the malehypopygium, more especiallyof the dististyle. The occurrenceof this speciesand Limonia (Geranomyia)walkeri Alexander on
Mt. Santa N{arta and again in Santa Catharina in southeastern
Rrazil is worthv of note.
Gonomyia(Gonomyia)methodicasp. n.
Belongs to the subcinereagroup; general coloration darkbrown; antennae black throughout; scutellum light yellow;
pleura yellow, the anepisternum more darkened; wings tinged
with brown; Sc short, Sc, ending opposite origin of Rs, Sczat
its tip i Rs Bently arcuated,not deflectedcephaladat outer end;
male hypopygium with the outer dististyle a sma1loval fleshy
lobe; phallosomebearing two strongly hooked lateral spines.
Male.-Length about 4.-1mm.; n'ing, 5.') mm.
Rostrum and palpi black. Antennie black throughout; outer
flagellar segments becoming long and slender, with relatively longdenseverticils. Head dark gray.
Pronotum dark brown. Mesonotal praescutum and scutal lobes
dark brown, very sparsely pruinose; anterior lateral pretergites andrestrictedhumeralregionvery pale yellow;pseudosuturalfovee elongate,
black; median region of scutum testaceous-yellow;scutellum lightyellow, the basemedially with a darkenedspot; postnotal mediotergite
pale brown, sparselypruinose,the cephaliclateral portions restrictedly
yellow. Posterior pleura and pleurotergite pale yellow, rvithout
clearly defineddark markings,the cephalicpleuritesrveaklyinfuscated.
Halteres dusky, the extreme base of the stem pale yellorv. Legs withthe coxrepale yellow, the fore coxa slightly more darkened;trochanterspale yellow; remainder of legs dark brown. Wings with a browntinge, the ill-defined stigma slightly darker brown; veins dark brown.
Costal fringe relatively long and conspicuous. Venation: .Sc short,
Scrending just beyondthe origin of Rs, Scrsubequalto Scz;Rs relatively
short, less than Rz+e+r;R3 gently arcuated, not deflectedcephalad attip, as in allied species;m,-cuat two-fifths the length of cell lst Mz.
Abdominal tergites dark brown, the sternites pale; hypopygium
yellowish brown. Male hypopygirrm with the basistyle relatively
short and stout, the outer lateral lobe longer than either dististyle;a small blunt tuberculatelobe on mesalface of proximal end of basistyle.
Outer dististyle a small, oval, fleshy lobe that is denselycoveredwith
delicate setule and a few long sete. Inner dististyle a little larger,

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producedinto a gently curvedblackenedspine,the margin of which,at near midlength,bearsa long seta;apexof style a short blunt lobe
bearingseveralseta, two of which are larger and fasciculate. Phallosomeasymmetrical,large and complex,bearingat nearmidlengtha
pale oval lobe; two lateral spinesthat are stronglyinterlocked.
Henrrar: Colombia. Holotype,d, Vista Nieve, Mt. Santa
Marta, altitude about 5,000feet, August 7,1926 (F. W. Walker).
Gonomyia(Gonomyia)methodicais readily distinguishedfrom
allied regional speciesby the structure of the male hypop-vgium.
NeognophomyiaAlexander.
Neognophomyia
colombicolasp. n.
General cotoration of mesonotal praescutum reddish; postnotum blackened; pleura pale yellow, with a conspicuous
blackened dorso-longitudinal stripe; wings whitish, with arelatively narrow brown seam along cord ; cell l st tr[z strongly
widened outwardly; rnale hypopygium with the lateral tergal
spinesblackened,microscopicallysetulosebeforethe actrtetips.
Male.-Length, about 6-6.5 mm.; wing, 6-6.5 mm.
Fernale.-Length, about 6.5-7 mm.; wing, 6-6.5 mm.
Rostrum yellow; palpi more darkened, especially the outer segments.
Antenna brownish black; flagellar segments oval, more elongate-oval
in male. Head yellow; vertex relatively narrow.
Anterior lateral pretergites lrslb\Mish white. Mesonotal praescutum
pale reddish or reddish brou'n, in caseswith vague indications of darker
stripes; scutal lobes dark brorvn or brorvnish black, the median region
paler, more silr'ery; scutellum pale; postnotal mediotergite chiefly
blackened, the lateral and cephalic portions narrorvly pale. Pleura
pale yellow, with a more or less entire, black, dorso-longitudinalstripe,
beginning on the propleura, extending to the postnotal mediotergite,
best defined as two large areas on the anepisternum and pleurotergite,
more nearly intermpted on the dorsal pteropleurite. Halteres pale.
Legs with the coxa and trochanters yellow; femora yellow, the tips
more or less infuscated; remainder of legs yellow, the outer tarsal
segmentspassing into brown. Wings whitish, with a relatively narrowbrown seam along the cord, narrowed behind, subobsolete on m-cu;
a scarcely indicated dark seam on outer end of cell /sl M2iva{te darkenings across the basal cells; veins brown, darker in the infuscated areas.
Venation: .Scr ending opposite or just beyond Rz, Sc2 pale, beyond
midlength of Rsi Rz about one-half RB+ricell lst Mz strongly widened
outwardly; a weak angulation or spur on ,n near its union with Mt;
cell ?nd A relatively wide.
Abdominal tergites dark brown, the basal segmentsyellow medially;sternites more uniformly pale yellow. Male hypopygium with thetergal spines appearing as blackened, gently curved rods that narrow1e31lAlexander: SouthAmerican Tipul'i'd'e

637

gradually to acute points, before their apices fringed with delicate
setule.

Colombia. Holotype, d, Vista Nieve. Mt. Santa
Hestrer:
Marta, altitude about 5,000feet, August 8, 1926(F. W. Walker).
Allotopoty/e, I , August 8, 1926,at light. 4 A. M.; Paratopotypes,
5 &9, August6-8, 1926.
The most nearly allied species seems to be l/. peraicax(Alexander) of Peru, which differs in the venation and structureof the male hypopygium, especiallythe tergal spines.
Gnophomyia Osten Sacken.
acriculasp. n.
Gnophomyia
General coloration of mesonotum black; thoracic pleura
striped longitudinally with dark and pale; haltereswith darkened
knobs; legs obscure yellow; wings with a weak brown tinge ;r-m connecting with Rs shortly before the fork; malehypopygium with the outer dististyle a simple, gently curved
rod that gradually narrows to a subacuteblackenedapex.
Male.-Length, about 7 mm.; rving,6 mm.
Female.-Length,about 7 mm.; rving,6 mm.
Rostrum and palpi black, paler in the female. Antenna black
throughout; flagellar segmentslong-oval, with verticils that are subequal to or a little longer than the segments. Head brown; eyes offemale apparently larger than in male, approximated or contiguous on
the vertex, in the male separatedby a strip of moderatewidth.
Pronotum brownish black. Anterior lateral pretergitesrestrictedly
pale yellow. Mesonotal praescutumblack or brownish black, sparsely
pruinose;humeral and lateral regionsrestrictedlybrightened;posteriorsclerites of mesonotum blackenerl. Pleura variegated, the dorsal
pleurites rvith a black longituclinalstripe, most intense on the anepisternum; beneath this stripe on the dorsai sternopleuritea moreyellowish area; ventral sternopleurite brorvn, rnore or less pruinose.
Halteres dark brown or brorvnish black, the extreme base of stem
brightened. Legs with the cox€ and trochanters yellow, the large
posterior coxa, together with the meral region, more conspicuouslyso;
trochanters yellow; remainder of legs obscureyellow, the outer tarsal
segmentsdarkened. Wings with a weak brownish suffusion,the stigma
scarcelyindicated; veins dark brown. Venation: .Scrending just before
the subevanescentRz,Sc2oppositer-m; r-rn connectingwith Rs a shortdistance before the fork of the latter; nt-cu opposite r-m and a shortdistancebeyond the fork of.M.
Abdominal tergitesblackened;sternitesmore yellowish;hypopygium
dark. In the female, the incisures are slightly darkened. Male
hypopygium with the basistyles short and stout. Outer dististyle asimple,gently curved rod, gradually narrowedto the subacuteblackened

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tip. Inner dististyle with the basal portion expanded,the apicalextensionobtuse, the whole style with severalcoarseseta. Whatappearsto be a tergal structureis a quadratemedianplate, the apex
decurvedinto two blunt blackenedlobesthat are separatedby a small
notch. Ovipositorwith the valvesblunt at tips, conspicuously
hairy,
the sternal valves very short.
HasrrA.r: Colombia. Holotype, c/, Vista Nieve, Mt. Santa
Marta, altitude about 5,000 feet, August 8, L926, at light,
4 A. M. (F. W. Walker) . Allotopotype, Q, August 8, 1926.
Gnophomyia acricula is allied to G. subhyalina Alexander,
differing especiallyin the body-coloration, details of venation,and structure of the male hypopygium.
Gnophomyia
vilis sp. n.
Generalcoloration pale reddish brown; pleura with a broken
blackish longitudinal stripe; femora yellow, the tips weakly
infuscated; wings tinged with grayish yellow, the veins yellowish
brown; veins Rg and Ra long, extendingnearly parallel for their
entire length; male hypopygium with the outer dististyle aflattened blade, the apex obtuse; ovipositor with the tergal
valves short and stout, truncated and setiferousat apex.
Male.-Length, about6-7 mm.; wing, 5.5-6mm.
Female.-Length, about 7-7.5 mm.; wing, 6-6.5 mm.
Rostrum pale brown; palpi darker brown. Antenna brownish
yellow, the outer flagellar segments somewhat darker; flagellar segments ova1, becoming more slender outrvardly, the verticils exceeding
the segments. Head obscure yellon', in cases dark grayish brown.
Mesonotum reddish brorvn, the humeral region and anterior lateralpretergites more yellowish; median area of scuturn rnore yellowish;
icuteliirm more teitaceous. Pleura reddish brou,n, rvith a conspicuous
blackish dorsal stripe, most intense on the anepisternum, paler tosubobsolete on the dorsal pteropleurite, Halteres brownish yellow,
the knobs infuscated. Legs with the coxa and trochanters yellow;
femora and tibie yellow, the tips weakly infuscated; tarsi yellow,
the outer segments passing into dark brown. Wings tinged withgrayish yellow; veins light yellowish brown. Venation: Scr ending
just before R2, Scg some distance from its tip i Rz a little shorter thanRs+si veins Re and Ra long, running nearly parallel for their entire
length; //t-cu at from one-third to two-fifths the length of cell lst Mz.
Abdominal tergites yellowish brown, darker laterally; sternites
more uniformly yellow; hypopygium reddish yellow. Male hypopygiumwith the outer dististyle a flattened blade, gently arcuated, the apex
broadly rounded. Inner dististyle short and stout, blackened, provided
with small black seta, the tip obtuse. Ovipositor with the valves
unusually short and stout, the tergal valves truncated at tips, darkened,
and bearing a few sete.

1e3uAlexand,er:SouthAmerican Tipul'id'a639

Hesnet:Colombia. Holotype, & , Orihueca, November
13, 1925 (F. W. Walker) . Allotopotype, I . Paratopotypes,
16d 9.
Gnophomyia ailis is closest to G. subhyal'ina Alexander,
differing in the coloration and venation of the wings and in
the structure of the male hypopygium, especially the outerdististyle.
Molophilus Curtis.
Molophilusflexilistylus sp. n.
Belongs to the plagiatu.sgroup; antennae short; mesonotal
praescutum reddish brown, with three more yellowish stripes;
knobs of halteres infuscated; male hypopygium with the basal
dististyle a long slender rod, expanded at apex, the margin at
this point with a series of appressedspines, together with asingle larger spine borne on the opposite margin.
Ittale.-Length, about 3.3-3.5nun.; wing, 4-4.2mm.
Fernole.-Length,about 3.5 mm.; wing, 4 mm.
Rostrum and palpi dark brown. Antennre short in both sexes,
if bent backn'ard not attaining the wing-root, dark brown throughout.
Head dark gray.
Mesonotal praescutumn'ith the ground-colorreddish brown withthree more yellorvishstripes; humeral region and anterior lateral pretergites obscure yellorv; scutum and scutellum reddish brorvn, thepostnotumbrorvn. Pleuratestaceous,latter somewhatmoretestaceous;the dorsal scleritesmore infuscated. Halteres dusky, the knobs infuscated, the color in large part due to abundant dark sete. Legs withthe coxa and trochanterstestaceous;femoraand tibire brownish yellow;
tarsi a little darker. Wings with a brownish tinge, the prearcular andcostalregionsa little more yellowish;veins pale brown, the macrotrichia
dark brown. Venation: Petiole of cell Ms abort one-half longer thanrn-cxt;vein ?nd A relatively short, ending opposite or just before thecaudal end of the oblique nt-cu.
Abdomen brou,n, including the hypopygium, the lateral region
darker brou'n. N[ale hypopl'gium rvith the apical beak ofsome'w-hat
the basistyle slender,gently curr-ed. Basal dististl'le a long, slenderrod, dilated at apex and here rvith a lateral seriesof acute appressed
spines,thesedecieasingin size outn'ardly; on opposite.face,at baseof
expandedportion of style, with a longer appressedspine. Oviposito_r
rvilh the valves long and slender,the tergal valves only gently curved.
Hesrrer : Colombia. Ilolotype, & , La Cumbre, Mt. Santa
Marta, altitude 8,000 feet, in deep cloud forest, August 8,
1926 (F. W. Walker) . Allotopotype, I . Paratopotypes, 3 d &,altitude 7 ,200-8,000 feet.

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The diagnostic features listed above, especially the structureof the male hypopygium, are distinctive of the species. One
paratype has the knobs of the halteres much paler, due todepilation.
Molophilus subtenebricosus
sp. n.
Belongs to the plagiatus group; allied to M. tenebricosus
Alexander (Colombia: Andes Occidental), differing especiallv
in the structure of the male hypopygium.
Male.-Length, about 4.3 mm.; wing, 5.2 mm.
Antenna elongate, strongly nodulose. General coloration ofmesonotalpraescutumblackishgray, with three reddishbrown stripes;
humeral and anterior lateral pretergiteswhitish; posterior scleritesof
mesonotum,togetherwith the pleura, blackish brown. Halterespale.
Male hypopygiumrvith the apical beak of the basistyleslender,as intenebricosrus,
but strongly curved. Basal dististylea long slenderrod,
the baseconstricted,beyondthis point flattened,rvith more than theapical third a slender,entirely smooth,black spine; outer aspectof
dilated portion of style with abundantappressedsetulre.
Haenar:Colombia. Ilolotype, d, \'ista Nieve, I\{t. Santa
Marta, altitude about 5,000feet, Arrgust7, 1926(F.W.Walker).
Molophilus walkeri sp. n.
Belongs to the plagiatu.t group; general coloration brownish
gray;'halteres yellow; wings obscure whitish, with two vague
darker crossbands,one at the cord, the other in the basal cells:
male hypopygium with the basal dististyle a powerful blackened
rod, the distal trvo-thirds roughened u'ith spinulae and spinous
setae, the tip suddenll' narrorved into a blackened spine, atnear one-third the length on outer margin rvith a powerfulerect spine.
tr[ale.-Length,about 3.6-3.timm.; wing, 1.5-4.7mm.
Rostrum and palpi black. Antennrc of moderatelength, if bent
backwardextendingabout to the wing-root,black throughout;flagellar
segmentsoval, gradually decreasingin size outwardly; verticils ofbasal segmentslong and conspicuous,becomingshorter and less conspicuouson the outer segments,on the penultimatebeing scarcelyas
long as the segment. Head black, sparselypruinose.
Mesonotum dark brownish gray. Anterior lateral pretergites
restrictedly pale yellow. Pleura black, sparsely pruinose. Halteresyellow. Legs with the coxe black; trochantersbrown; remainder oflegsblack, the femoralbasesrestrictedlypale. Wings obscurewhitish,with two vague darker crossbands,one at the cord, the secondmore
diffuse, crossingthe basal cells; veins pale, somewhatdarker in theinfuscated areas; trichia of veins pale brown, darker and forming

1e3llAl,exander: South American Tipulide

64r

conspicuous patches on those veins traversing the infuscated areas,
especially along the cord. Venation i rn-cu oblique, subequal in length
to the petiole of cell Ms; vein ?nd A ending shortly beyond the caudal
end of nt-cu.
Abdomen brownish black, including the hypopygium. Male
hypopygium with the apical beak of the basistyle decurved to anacute point. Outer dististyle relatively slender, bifid at apex. Basal
dististyle a powerful blackenedrod, the apex narrowed into a blackened
spine, the distal two-thirds of the style with abundant appressedspinuleon all faces,those of the ventral or inner face more slender and spinous;
on outer margin of style at near one-third the length with a powerfulerect spine.

Hanner: Colombia. IIolotypz,&, Vista Nieve,Mt. Santa
Marta, altitude 5,000feet, August 8, L926,at light,4 A.M.
(F. W. Walker).
Molophiluswalkeri is respectfully dedicatedto the collectorof this abundant series of Tipulida from Santa Marta, Mr.
Fred W. Walker. The structure of the male hypopygium isdistinctive.
Molophilus marth& sp. n.
Belongs to the plagiatu.s groutr); general coloration of mesonotal praescutum reddish brown, the postnotum and pleura
blackened; antennae relatively long, extending to beyond thewing-root; knobs of halteres infuscated; legs dark brown; wings
tinged with brown, the long abundant macrotrichia darkbrown; male hypopygium with the basal dististyle a powerfulrod, the distal half fringed with setae and appressed spines; basal
half of style on outer face with two powerful erect spines.
Male.-Length, about 3.5 mm.; wing, 4.4 mm.
Rostrum and palpi black. Antennrc black, relatively long, if bent
backward extending to beyond the rving-root; flagellar segmentsoval,
with long, unilaterally arranged verticils and additional long delicate
setula. Head dark brorvn.
Mesonotal praescutum reddish brown, the anterior lateral pretergites whitish; scutal lobes and scutellum darker, the postnotum
blackened. Pleura blackened. Halteres yellow, the knobs infuscated.
Legs with the coxe and trochanters yellow to testaceousyellow; remainder of legs dark brown. Wings with a faint brown tinge, moreevident as a seam along vein Cu; veins pale brown, the long abundant
macrotrichia dark brown; costal fringe long and dense. Venation:
m-cu about two-thirds the length of the petiole of cell Msi vein ?nd A
long, gently sinuous, ending beyond midlength of the petiole of.cell Ma.
Abdomen brownish black, the hypopygium more yellowish. Male
hypopygrum with the apical beak of basistyle slender and straight;
margin of basistyle cephalad of beak extensively blackened and642

Annals EntomologicalSocietyof America [Vol. XXIV,

microscopicallyroughened. Outer dististyle relatively stout, bifid,
the inner arrn more elongate. Basal dististyle a powerful rod, thedistal half or slightly less narrowedto a gently curved spine, theconcaveface provided with a densefringe of sete in addition to aspines;basalhalf of style more enlarged,with twoseriesof appressed
powerfulerectspines. Phallosomicstructuresmall,oval, glabrous.
Hastrer:
Colombia. Holotype,d, Vista Nieve, I\4t. Santa
Marta, altitude about 5,000feet, August 6, 1926(F. W. Walker).
Molophilus marthe is very distinct from all describedspeciesof the genusin the structure of the male hypopygium.
Molophilusbidigitatussp. n.
Belongs to the plagiatus group; general coloration reddish
brown, the head, pronotum and humeral regions of praescutum
light yellow; thoracic pleura dark brown; halteres yellow;
wings grayish yellow, the costal region clearer yellow; malehypopygium with the basal dististyle bifurcate at apex; phallosomic structure a pale setiferouscushion,its caudal end bifid.
Male.-Length, about 2.5 mm.; wing, 3.3 mm.
Rostrum and palpi brown. Antenne q'ith the scape yellow, theflagellum broken. Head pale yellow, more infuscated behind.
Pronotum and anterior lateral pretergites light sulphur-yellow.
Mesonotum reddish brown, the humeral region of praescutum lightsulphur-yellow; posterior sclerites of mesonotum somewhat darker
brown. Pleura dark brown, the dorso-pleuralregion chiefly darkened,
the posterior portion yellow. Halteres yellow, with golden setie.
Legs-with the fore coxueinfuscated,the remaining coxe and all trochanters yellow; remainder of legs broken. Wings grayish yellow, thepaie,the macrotrichiabrown.
baseand costalregioncleareryellorv;r,'eins
Venation: fn-cu about trvo-thirds the petiole of cell .41r;vein 2nd A
relatively elongate, extending to about opposite one-third the lengthof the petiole of cell ,1g.
Abdomen yellowish brown, darker laterally; hypopygium yellow'
with the apical beak of basistyle slender, gently
Male hypopygium-Oulerdististyle bifid, the inner afrn more slender. Basalcurved.
havingmuch the sameshapeas the apexof the outer dististyle,dististyle
being ionspicuouslybifurcate, both afins appearingas slenderblackened
spines,the outer arrn more curved and a little stouter than the innerspine; stem short. Phallosomicstructure a pale cushionthat is covered
niittr long conspicuousset.e,the apexdeeplynotchedmedially. Aedeagus
relatively short, subequalin length to the outer dististyle.
Hasrrar : Brazil. tlolotype, & , Jaragua, Santa Catharina,
August 2L, L92g (F. Schade).
Molophilus bidigitatus is very distinct from all described
Neotropical species in the structure of the basal dististyle ofthe male hypopygium.