V ol.6(3):35 1-3 58

ORIUNI'AL INSEC'I'S

July 1972

NEWAND LITTLE-KNOWNINDIANCRANEFTIES(DIPTERA: TIPT]LIDAEFVCharlesP. Alexaniler
39, OId TownRoad,Amherst,Massachusetts,(I.S.A.
Ansrnacr' Three new speciesof Indian Tipulidae are described,these being Limonia(Melanolimonia) latemarginata, L, (sivalimnobia) clavula and, pilaria tiro all fromKameng, Assam. In addition, figuresof the venation and male hypopygium of several
Indian speciesof thc Hcxatomine genusParadelphomyra
Alexander, not previouslyillustrated, are providcd.

The precedingpart under this generaltitle was published in oriental Ins.,
5 : 73-82, January 1971. The noveltiesdescribedare from Kameng,Assamwhercthcy werecollectedby Dr. Fernandschmid, to whom I am indebted fornumerousspeciesof these flies. continuing the policy previouslyadoptedI
am providingfiguresof severalspeciesof Indian
ntt previously illust"rutrifli.r
rated' all of thesebelongingto the Hexatomine
genusparadeiphomyiaAlexander.
Sunneutly LntoNlN,tr
1. Limonia (Melanolimonia)latemarginata,sp. nov. (Figs. l, 4)Generalcolorationof thoracicdorsum brown, sparsely pollinose; antennae
with scape and pedicel obscureyellow,flagellumbtackjhattereselongate;legs
brown, tarsi black; wings light brown, stigma oval, darker brown; abdomen
brownish-black;male hypopygiumhaving posteriorborder of tergitewith small
laterallobes,the emarginationbroad;dorsal dististylelong and slender.
Male : Rostrum obscureyellow, silvery pruinose, palpi brack. Antennae
with scape and pedicel obscureyellow,flagellumblack;proximalflagellarsegmentsoval, outer onesprogressivelymore elongate, shorter than the longest
verticils; terminal segmenton oi.rterthird about one-halflongerthan the penuttimate. Front obscureyellow,anteriorvertex broad, silvery, posterior vertex
brownishblack.
Pronotumbrown, gray pruinose. Mesonotalprescutumdull brown, sparsely
yellow pollinose, with indications of darker stripes, humeral region moreyellowed;scutallobesyellow pollinose;posteriorscleritesgray pruinose. pleura
blackened,heavily pruinose. Halteres elongate,stemdark biown with yellow
base,knob black. Legs with fore coxae blackened, remaining coxae and alltrochanters obscure yellow; femora and tibiae brown, tarsi black;claw with a
Published November 1972352( ) RIIiNl' AL INSL CT S

Vol. 6, No. 3

long slcndcrncarlybasalspine. wing (Fig. 1) light brown, stigmaoval, darker
brown; vcins light brown. Longitudinalveins beyondgenerallevelof origin
endingshortly beforeorigin of Rs, sc, notof Rs with trichia. venation : ,Sc1apparent;cell 1sl M, shorterthan vein Ma; m-cu shortly beforefork of M.
ventraldististyleof hypopygiummore yellowed.
Abdomenbrownish-black,slightlynarrowed(Fig.
ninth tergite,/, transvcrsc,a)withMale hypopygium
a very broad
byseparatedsmall,lobes
lateralwithposteriorborderoutwardly,Proctiger
base.
atlobeofthediameterthetwice
about
that
is
emargination
6, withBasistyle,innigrithorax'
than
less
conspicuous
withietae feweranddististyle,Dorsalconspicuous.
setae
delicate
long
tbe
ventromesallobe low,d, long and slender,curvedgentlyto the acute tip; ventral style with a single
stout pale rostral spine from a membranousbasallobe' the spinesubequaling, with mesal-apicallobe
lengthto the prolongationbeyondit. Gonapophysis,q,,size,lateral margins
ofmoderatelobewith apicalsmall, acute. Aedeagus,
setulae'of basewith abundantmicroscopic
Length.'if, about6.5 mm, wing7.2mm, antennaabout 1.3mm.
Holotype $, INnta. : Assnu : Shergaon,Kameng, North-East Frontierfeet,8. v. 1961,Coll. Schmid'
Agency,6400
Thereare rather numerousregionalspeciesin the subgettusMelanolimonia
Alexander,the one that is most similar to the presentfly beingLimonia(Melano'limonia) nigrithorax (Brunetti) which differs in the coloration of the thorax
and abdomenand in hypopygialstructure,especiallythe tergitewherethe outer
lateral lobesare much largerand more approximated.
clavula,sp. nov. (Figs. 2, 5)
2. Limonia(Sivalimnobia)
Sizelarge(wing approximately10 mm); generalcolorationof thorax yellow,
prescutummore or less darkenedanteriorly;wingsstronglytingedwith brown'
unpatternedexceptfor the darker brown stigma, Sc long, Sc, ending beyond
midlength of Rs; male hypopygium with tergallobesconspicuous'separated
by a v-shaped emargination;gonapophysesclavate, enlarged distally, tipsobtuse.
Rostrumbrown, palpi black. Antennae with scape black, pedicel obscureshortyellow, flagellum brown, darker outwardly;proximal flagellarsegments
verticils
very
subequal;
threetbe
outerlengthened,
ones
more
oval, succeeding
sparsely
darkbrown,
Headthesegments.
thanlonger
one-half
long, aboutpruinose;anteriorvertexbroad,nearly twice the diameterof scape.
Pronotalscutumdark brown, yellowedlaterally, scutellum usually concolorous, in casesyellowed. Mesonotalprescutumobscureyellow,in cases(including type) with anterior end darkenedto form a short central stripe; remainderof notum and the pleurapaleyellow. Haltereswith stem obscureyellow,knob
darkened. Legswith coxae and trochanters yellow; femora brownish-black'bases narrowly yellowed, remainder of legsblack; claw long and slender,with

1912

A l .l :X A N I)l :R : ()N IN I)l A N

fl P t,Li l )A l ,353

a l.rrg spincat nearmidlength,with few smaller more basal denticles. wings(l;ig. 2) stronglytinged with brown, stigma oval, darker brown; veins darkb'.wn. Longitudinalveins beyond generallevet oforigin of Rs with trichia,
itcluding a few at tips of both analveins,morenumerouson 2nd l. venation :.Sclong, ,Sc, ending some distancebevond origin of Rs, in casesto about
two-thirds the length of vein, scs near its tip; free tip of sc2slightlybefore
Rr; m-cuat or slightly beforefork of M.
Abdominal tergitesdark brown to brownish-black,sterniteslight yellow,
incasesthe basesof intermediatesegments
narrowly darkened,hypopygiumblack.
Male hypopygium(Fig. 5) wirh laterallobesof ninth tergite,l,'clnrpic.rous,
theenrarginationbroadly v-shapcd.Dististyles,d, generallyasinfortis. Gonapophysis,g, diagnostic,clavatc,outerendsdilated,apexobtuse. Length.',1,about 7.5-8mm, wing 9-10mm, antennaabout 2-2.2 mm;e,about8-9mnr,wing l0-ll mm, antenna2.1-2.2mm.
Holot.ypt,.j,luom : Assau : Shergaon,Kameng,North_EastFrontier
Agency,64(x) t'cct, 8. v.1961, coll. Schmid. Artotype Tarung Dozong,Q,
Kameng,7800I'eet,6.vi.1961. paratopotypes
2gg,l Q, with type. paratypes6 g ],with allotype.
The most similar,regio'al speciesappears to be Limonia (sivatimnobia)/orlls (Brunetti) which is smailer, having the wingspaler, with sc short,sc1e.ding shortlybeyondorigin of R.r,and with the hypopygialdetairs
distinct,
includingthe tergiteand gonapophyses.
It may be notedtbat the typesof lbrtis
are females,
not malesas indicatedand describedby Brunetti, as was correctedby Edwards (RecordsIndian Museum, 26 :2964 1[124). This latterspeciesisthe type of the subgenusSivalimnobiaAlexander(phittpp,tneJ. Sci.,9Z :iZO_ZZl;r963).

3. Pilaria tiro, sp. nov. (Figs. 3, 6)
Generalcoloratibnof thorax and abdomen fulvous and yellow; antennae
with scape light yellow, remainder brownish-black; anterior vertex
broad,polishedbrown;Iegsbrownish-black,femoralbasesyellowed;
wingslight brown,
stigmaldarkeningvery smalr, veins R, and Rrls iubequal, veii
R, straight,
nearly parallel to Rr12, ceil R, at margin about on.-hfth as extensive
as celr
R' cellMr subequalin lengthto its petiole;marehypopygiumwith gonapophysis extendedinto a sle'der spine,the posteriotuogl. ;it[ u comparablecurved
narrow blade.
Rostrum brown,palpi black.Antennae with scape short and stout, lightyellow, remainder of organ brownish-black;proximal flagellar
segmentsshort,beyondthe third becoming progressivelylonger and moie slender, thoseatmidlength with long
thelongest
nearlythreetimes
thesegment,on,verticils,
outer segments
much shorter,lackingon outer three. Front yellowed,iemainderof headbrownish-black,anteriorvertexbroad,polishedbrown.

354

ORIENTAL INSICTS

Vol.6, No.3

Pronotumwith scutumnarrowly blackenedmedially,sidesand the scutellum
yellow. Mesonotalprescutumand scutallobesdark fulvous, posterior sclerites
and the pleuraecleareryellow. Haltereswith stemyellow,knob brownish-black.
Legs with coxae and trochanterslight yellow; femora dark brown, bases
yellowed,tibiae and tarsi brownish-black;clawslong and slender.Wings(Fig. 3)
light brown, prearcularand costalfieldslight yellow,stigmavery small,brown,veinsdarker brown. Venation: ,Sc1
endingabout opposite four-fifths Rs, .9c,near its itip; Rs nearly twice R21s1a;R, subequalto Rr*r; vein Rs short andcell R, at margin about one-fifth as extensive
straight,nearlyparallelto R1a2,
as cell Rs; cell M, subequalin length to its petiole; m-cu shortly beyondmidlengthof Msaa.
Abdomenlight brown, subterminal segments,especiallythe eighth, darkbrown, hypopygiumobscureyellow. Male hypopygium(Fig. 6) as in the genus,
outerdististyle,d, slender,outer apical angle extendedinto a slender spine,g, outwardlyextendedinto a long narrowly acute spine posterior
gonapophysis,
anglewith a similar slendercurvednarrow blade.
Length.' J, about 8 mm, wing 7.5 mm, antennaabout 1.8mm.
Holotyped, INpta: Nrnl: Shergaon,Kameng, 6400 feet, 8.v.1961,Coll.
Schmid.
Other regional species,as Pilaria amica (Alexander) and P. coorgensis
Alexander,havecell Ml of the wingslacking. Other speciesthat have this cellretaineddiffer from the presentfly in the venationof the radial field, including
the lossof vein Rra3and the broadercell R, in comparisonwith cell Rr.
(Oxyrhiza)dichromataAlexander(Figs. 7, 10)
4. Paradelphomyia
Paradelphomyia(Oxyrhiza')
dichromataAlexander, 1965,Ent. News.,76 : 41.

Typed, Chingsao,Manipur,Assam, 3800feet, 13.vi.1960.Venation (Fig.
7), malehypopygium(Fig. 20).
(Oxyrhiza)flavescens(Brunetti)(Fig. I l)
5. Paradelphomyia
Cladura flavescarsBrunetti, 1911, Rec.Indian Mus., 6 : 284.
Gnophomyiafurcata Br:unetti, 1912,Fauna British India Diptera, Nematocera, p' 491.

Male hypopygium(Fig. 11).
(Oxyrhiza)hkayamensis
Alexander(Figs.8, 12)
6. Paradelphomyia
Paradelphomyia(Oxyrhiza) hkayamensisAlexander, 1965, Ent' News., 76 :43.

Type d, Hkayam Boum,Assam, 7500feet, 20.vi.1960. Venation (Fig. 8),
male hypopygium(Fig. l2).
7. Paradelphomyia(Oxyrhiza)myriacanthaAlexander(Figs.9, l5)Paradelphomyia (Oxyrhiza\ myriacantha Alexander, 1965,Ent. News., 76 : 44'

Type d, ChumzomoiChoka,Sikkim, 11,800feet,8.vii.1959.
Venation(Fig. 9), malehypopygium(Fig. l5).

t972

ALLXANDIR : ON INDIAN TIPULIDAL,

3-55

ll. Paradelphomyia(Oxyrhlza)newrr Alcxantlcr(Fig. l6)
Paradelphomyia(Oxyrhiza)newar AlcxanrJcr,1965,J. Ntw York Ent. Soc.,65 : 156.

Type d,SimbhanjangPass,Nepal,8190feet, 1.x.1956. Malc hypopygium(Fig. l6).
9. Paradelphomyia(Oxyrhiza) ruficolor Alexander(Figs. 13, lZ)
Paradelphomyia (Oxyrhiza) ruJtcolor Alexander, 1965, Ent. News.,76 : 46.

Type d, Bourzil R.H., NorthwestFrontier Province, Pakistan, 11,150 feet,10.xi.1953.Venation(Fig. 13), male hypopygium(Fig. l7).
10. Paradelphomyia(Oxyrhiza)tritumutaAlexander(Figs. 14, 18)
Paradelphomyia (Oxyrhiza) tritumula Alexander, 1965, Ent, News.,76 : 47.

Typc d, Mangu Chatti, Pauri Garhwal,Kumaon, Uttar Pradesh, 9000-9270
fcct, 31.v.1958.Venation(Fig. l4), matehypopygium(Fig. l8).

356Vol. 6, No. 3

ORIENTAL INSECTSt972

3s7

Al .ttxAN t) l l R : o N tN D tAN 'l tl ,t,t.tl ) Al :((,d //H,u,, \'.

$iii"\tr/\;'kv$53)1t . "\\l 'l \J /#//

Wl/r///,tlnIllll\\4(f"lW.

lllll;' l11,/r r " ',///'\l>(K;#re.r ff'l .r \l6

II4(6;fr
Figs. l-6, Venation of :1, Limonia(Melanolimonia) latemarginata,sp. nov.; 2, Limonia (Sivalimnobia)clavula,sp. nov.; 3, Pilaria tiro,
sp. nov. I\{ale hypopygirtm of : 4, Limonia (Melanolimonia)
latemarginata,sp. nov.; 5, Limonia (Sivalimnobia) clavula, sp' nov';
6, Pilaria tiro, sp. nov. (Symbols, venation, R Radius; malehypopygium, a, aedeagus;D, basistyle;d, dististyle;g, gonapophysis;t, 9th tergite).

aFigs. 7-12. Venation of :7, Paradelphomyia(Oxyrhiza) dichromata
Alexander; 8, Paradelphomyia (Oxyrhiza) hkayamensis Alexander:
9, Paradelphomyia (Oxyrhiza) myriacantlra Alexander. Male hv_popygium of : 10, Paradelphomyia(Oxyrhiza) tlichromata Alexander:ll, Paradelphomyia (Oxyrhiza) flavescens (Brunetti); 12, paradel_phomyia (Oxyrhiza) hkayamensis Alexander. (Symbols 6, basistyle;d, dististyle style; g, gonapophysis; r, 9th tergite; yjf, ventral fork);

358

ORIENTAL INSECTS

fr''-rll7

Vol.6,No. 3

drurtcolor
Figs: 13-18.Venationof z 13,Paradelphomyia(Oxyrhiza)
Albxander'tritumula(Oxyrhiza)
Paradelphomyia'!4,Atex-anOer;(oxvrhizal mvriacantha
ira"i" ivp"'pvgiitm of : 15; Paradei'lphomvia
newar Atexander;17'(oxvrhiza)
Paradelihomvia
Ai;;;ili;,'18, Paradelphomvia
Alexander;(Oxvrhiza) raficolor
iioartpni.yio
D' basistvle;(Svmbolsa, aedeagus;
Alexander'
tiitunuli@;y;l;t4fork)'
ventraluf'p,phallosome,a, iittlt,yf.; g, gonapophysis;