jNEW OR LITTLE.KNOWN TIPULID.?E FROM EASTERNASIA (DIPTERA), VI'
By Cn.mr,os P. ALD(aNDER
Of Amharst, Massachusetts
TWO PLATES

at this time are almostentirely based
The craneflies discussed
on extensivecollectionsmadeby Prof. Syuti Issiki on Mount Kirishima, in southern Kiushiu, Japan; in the rich native foreston the mountains of Yakushima Island, south of Kiushiu; andat Arisan and Chikurin, in the mountainsof Formosa. A few
additionalspecieswere taken in China,kindly submittedby Professor Jacot and Mr. H. S. Parish, and an interesting TipuJa wassent by Doctor U6no, taken above 8,500 feet in the JapaneseAlps, the highestaltitude at which craneflies havebeenreeorded
from the main island of Japan. I am very deeply indebtedto
all of the above,but especiallyto ProfessorIssiki, for their kindefforts in making known the rich tipulid fauna of easternAsia
and for the privilege of retaining the types in my collection.
TI PULI N E
TIPULA UfNOI rt. nov.

I
l'

*l

i'
I

General coloration of head and mesonotumdark, pruinose;wings light brownish, variegatedwith subhyalineand darker
brown areas; abdomenyellow, both tergites and sternites conspicuouslytrivittate with brown, the outer segmentsmore uniformly darkened;male hypopygiumwith the medianarea of theblackenedblade; eighth stertergite producedinto a compressednite with the caudal margin unequallytrilobed, the lobes setiferous.
Male.-Length, about 13 millimeters; wing, 14.5; antenna,
about 4.2. Describedfrom an alcoholic specimen.
Frontal prolongation of head dark brown, the nasus conspicuous; maxillary palpi black, the terminal segment exceeding
the remaining segments combined. Antenna with the scapal'Contribution from the Department of Entomology, MassachusettsAgricultural College.
59

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and first flagellar segmentsobscure yellow, the remainder ofthe organ black; flagellar segmentsstrongly incised. Head apparently dark gray in dry specimens;vertieal tubercle bifid.
Mesonotal prescutum apparently gray, the margins of thebroad median stripe brown; lateral stripes more broadly darkbrown; scutal lobesconspicuouslyblackened;posterior sclerites
of mesonotum dark colored in alcohol. Pleura chiefly darkbrown, possibly pruinosein fresh specimens. Halteres yellow,
the knobs infuscated. Legs with the coxe dark brown, theirtips paler; trochantersobscureyellow; femora brownish yellow
basally, passing into dark brown; tibire somewhatpaler, thetips dark brown; tarsi dark brown. Wings pale brownish, variegated with whitish subhyalineand darker brown; cells C and
Sc more yellow; the subhyaline areas are especially broad andconspicuous
before the cord, there forming an incompletecrossband; other more-restrietedpale areasbeyondthe stigma in cells
R, and R, and in the basesof the anal cells. Venation: Rs
relatively long and straighti Rr*rentire but without macrotrichia; cell lst M, very small, pentag:onal;petiole of cell M,
relatively long, approximately three times m.
Abdominal tergites yellow, trivittate with brown, the medianstripe being continuous and especially conspicuous;outer segments, including the hypopygium, more infuscated; sternites
more evidently trivittate with dark brown. Male hypopygium,(Plate 2, fig. 19) with the tergite, 9t, and sternitq gs, extensively fused; median portion of the tergal region (Plate 2,
fig. 20) producedcaudadinto a broad lobe,the apex blackened;viewed from above,the apex is weakly carinate; viewed laterally, the lobe is developedbeneath into a compressedblade.
Outer dististyle a flattened pale blade, a little dilated apically.
fnner dististyle, zd, bidentate, the outer margin at base again
toothed. Eighth sternite (Plate 2, fig.21) pale,the caudalmargin broadly notched,forming wide lateral lobesthat bear longer
sete; median region beneath further produced into a smaller
lobule.
Habi,tat.-trapan ( Honshiu).
Holotype, alcoholic male, Mount Kiso, Komagatake, Shinano,altitude 2,700meters, on snow, August, L928 (M. Udno),
I take great pleasure in naming: this interesting Tipula inhonor of the collector, Dr. MasuzoU6no. Ii,Wla udnoi is allied
and generallysimilar to T. shomfoAlexander (Japan), differingin the diagnostic features indicated, especiallythe coloration of
the body and wings.

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Aleuander: Ti,pulida from Eastern Asi,a,VI

61

LIMONIIN,,E
L IM ONIIN I
LIUONIA(LIBNOTES) RECURVINEBVIS sp. nov.

General coloration greenish yellow, the prascutum with a
median brownish black stripe; antennal scape black, the flagellum greenish; knobs of halteres dark brown; femora yellow,
the tips rather narrowly blackened; wings hyaline, with a restricted brown pattern; vein 2d A bent at a right angle into themargin; male hypopygium with the ventral dististyle large andfleshy, the rostral prolongation shorf blackened apically, the twospines unequal, divergent.
Male.-Length, about ?.5 millimeters; wing, 8.
Rostrum, and palpi black. Antenne with the scape black, theflagellar seg@ents green, long-oval. Head dark, pruinose.
Pronotum black, paling to yellow on the sides. Mesonotal
prescutum yellow with a broad brownish black median stripethat is a little constricted before midlength, thence widened tothe suture; scutal lobes conspicuously brownish black, the median area broadly pale yellow; scutellum dark brown, split bya narrow median yellow vitta; postnotum chiefly darkened.
Pleura, including the pleurotergite, pale yellow. Halteres ofmoderate length, pale yellow, the knobs dark brown. Legs withthe coxe and trochanters greenish yellow; femora yellow, thetips blackened; tibia obscure brownish yellow, the tips very
narrow,ly darkened; basitarsi brownish yellow, the distal third
and remainder of tarsi blackened; claws long and slender, witha single basal spine. Wings hyaline, iridescent, the prearcularand costal region more yellowish; a restricted dark brown pattern, as follows: At arculus; a narrow seam at origin of Rs;
Scr; a circular stigma; very narrow seams on the cord and outerend of cell lst Mr; the extreme wing tip in cells R, to Ru; narrowmarginal seams on the medial, cubital, and anal veins, large
and conspieuous on the 2d anal vein; a:rillary region in cell 2d A
narrowly darkened; veins yellow, darker in the infuscated areas.
Venation: Sc long, Sc, extending to some distance beyond thefork of Rs, Sc, just beyond this fork; Rs strongly arcuated tofeebly angulated; m-cu more than one-half its length beyond thefork of M; vein 2d A bent at a right angle into the margin.
Abdominal tergites dark brown, the caudal margins of thesegments very narrowly pale; sternites pale gteenish yellow; hypopygium pale. Male hypopyglum (Plate 2, fig. 22) with theterg:ite, 9f, parallel-sided, the caudal margin with a very broad

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1930

V-shaped notch. Basistyle, b, small, the ventromesal lobe low
and relatively inconspicuous. Ventral dististyle, ad,, large and,fleshy, much larger than the basistyle, the rostral prolongation
short and stout, blackenedapically and with a longitudinal blackened ridge the entire length; the two rostral spines are basal in
position, the outer stouter but only a little longer than the inner, the spines divergent. Dorsal dististyle a slightly curved
rod that terrninates in a long acute spine. Gonapophyses,gf,
with the mesal apical lobe slender, the tip acute, the margin
microscopieallyserrulate. .4Edeagus,
q terminating in trvo divergent flaps. Anal tube with a marginal series of long powerful sete almost to the apex.
H abitat.-W esternChina.
Holotype,male,Kwanhsien,Sze-chwan,June 7, 1928,throughMr. Herbert S. Parish.
I would have identified the present speciesas L. tri,maculata
Brunetti of the eastern Himalayas, except for the fact thatneither Brunetti nor Edwards mentions the recurved 2d analvein, nor is it figured by Bagchi. In any case,the name Limnobin trimaculnta Brunetti (1912) is preoccupiedby Limnobi,atri,mnculataZetterstedt(1838). The presentspeciesand related
forms, although most convenientlyplaced in Li,bnotes,show certain venational features of Li,montaand hypopygial and ungual
charactersthat cannot be distinguished from those of numerous
speciesof Dicrano?nAin.
LIUONIA(GERANOMYIA) RADIALIS sp. nov.

General coloration obscureyellow, the rostrum and antenne
black; head black with a median gray,line; legs yellow; wings
grayish yellow, with a very restricted brown pattern that isconfinedto the veins; Sc long, Sc, extending to just before thefork of Rs; male hypopygium with the ventral dististyle bearing two long divergent rostral spines that arise from enlarged
bases.
Mol,e.-Length, excluding rostrum, about 5.5 to 6.b millimeters; wing, 6.5 to 7.5; rostrum, 2.4 to 2.6.
Rostrum and palpi black. Antenne black throughout; flagellar segmentssuboval,gradually decreasingin size outwardly,
with short inconspicuousverticils. Head black, with a dorsomedian silvery gray line extendingthe entire length of the head.
Pronotum obscureyellow, restrictedly darkenedon sides. Mesonotal prescutum obscure yellow, sparsely pruinose, with anarrow brown median line; scutal lobes conspicuouslyand extensivelydark brown, weakly pruinose;medianregion of scutum

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Alerander:

Ti,pulida from Eastern Asia, VI

6g

and scutellum yellow, with a capillary dark line; postnotal mediotergite pruinose, darker posteriorly and laterally, each anterolateral angle wih a small yellow spot. Pleura testaceousyellow, the dorsal pleurites more infuscated to form an ill-definedbrown stripe. Halteres yel,low, the knobs weakly infuscated.
Legs with the coxe and trochanters yellow; femora obscureyellow, slightly darker toward tips; tibire brown, the tips andthe tarsi slightly paler, more yellowish brown. Wings (plate
1, fig. 1) grayish yellow; cells Sc and Cu, clearer yellow; a very
sparse brown pattern that is chiefly in the costal and radial
fields, including spots at the supernumerary crossvein in cell Sc,
origin of Rs, tip of Sc, the stigma, tip of vein R, and as narrow
seamsalong the cord, outer end of cell lst M, and as very vague
marg:inal clouds at ends of the anal veins; veins pale brown, veins
Cu, lst A, and 2d A more yellowish. Venation: Sc unusuallylong, Sc, ending just before the fork of Rs, Se, at its tip; asupernumerary crossvein in cell Sc; cell 1st M, closed; m-cu before the fork of M, the distance variable.
Abdominal tergites brownish yellow, indistinetly variegatedwith darker; sternites chiefly clear yellow. Male hypopygium(Plate 2, fig. 23) with the basistyle, b, relatively small. Ventral dististyle, ad, very ,large and fleshy, the rostral prolongation with two long divergent spines arising from enlarged basal
tubercles. Gonapophy5€s,9, with the mesal-apical lobe very
short. One dististyle of the holotype shows three rostral spines
instead of the apparently normal number of two.
Habitat.-Japan(Kiushiu) .
Holotype, male, Kosugidani, Yakushima, altitude 2,500 feet,
April 29, 1929 (S. Issiki). Paratopotype, a teneral male.
Limonia (GeranomEia) radinl;is is most olosely allied to Z.
(G.) aaocetfo (Alexander), differing especially in the unusually
Iong Sc, the more-restricted brown wing pattern, and the structure of the male hypopygium
LIMONIA(GERANOI,IYIA)ALPESTRTS

sp. nov.

Generalcolorationdark gray, the humeral region of the prescutumand the scutellummore yellowish;ventral thoracicpleura
pale; femora with a dark subterminalring; wings grayish, thecostalregion more yellowish; a heavy dark brown pattern thatis chiefly costalin position,the third and fourth areasaboveRs
very extensive,confluentor nearly so; abdominaltergites darkbrown, the ,sternitesobscure yellow.
Female.-Length, excluding rostrum, about b.b millimeters:
wing, 7.2; rostrum, about 2.

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Rostrum moderatelyelongate,black; palpi black. Antenne
black throughout; flagellar segments oval. Head dark, pruinose,the anterior vertex paler gray.
Pronotum dark brownish gray. Mesonotal prascutum darkgray, the humeral region restrictedly pale yellow, the disk withfour narrow darker lines, the more lateral pair representingthe
interspaces; scutal lobes dark gray, the median region and thescutellum conspicuouslypale yellow; postnotum dark, heavily
pruinose. Pleura dark gray, the sternopleurite, meron, andpteropleurite pale yellow. Halteres yellow at base, the remainder infuscated. Legs with the co:<eand trochanters paleyel,low; femora obscure yellow, brighter basally, deepeningto
a subterminal dark brown ring, the tips again narrowly yellow;
tibie and tarsi pale brown, the outer segments of the latter
passing into black. Wings (Plate 1, fig. 2) grayish, the prearcular and costalregions more yellowish; a heavy,chiefly costal
brown pattern, including sevenmajor areas,the third and fourth,placed abovethe origin of Rs and at end of Sc eonfluent in cell
G or nearly so; those portions of the secondand third areas incell R paler, with narrow darkenedmargins; stigmal area largest; a vague paler cloud betweenthe first and secondareas;interrupted seamsalong cord and outer end of cell lst M,; small
marginal areasat endsof veins Cu,, 1st A, and 2d A, the lastnamedlarger; veins pale yellow,thosein the oloudedareasmore
infuscated. Costal fringe (female) short. Venation: Se ilong,Sc, endingabout oppositemidle.ngthof Rs, Sc, at its tip; R, andfree tip of Sc, in alignment; cell lst M, long and narrow, rectangular, subequalto the long:estvein beyondit; m-cu at forkof M, shorter than the distal section of Cu,.
Abdominaltergites dark brown, the sternites obscureyellow;
ovipositor with the valves relatively short, yellowish horn color,the tergal valves unusual,lyslender and nearly straight, theirbasesdarkened.
Habitat.-Formosa.
Holotype, female, Heigansan,altitude 5,800 feet, October 24,
1928 (S. Issi,ki.).
Limonin (Geranom,gi,a)alpestds is apparently most closely
allied to L. (G.) septemnotalo(Edwards), which is similarly alate fall speciesfrom the higher mountains of Formosa. The
latter species differs in the reddish brown coloration of thethorax, uniformly brown legs, and the distribution of the wing
pattern.

4z,tAlerand,er: Tipulida from Eastern Asi,a, VI

LIDIONIA (TERYPTICOTYIA)

APICALIS UAJUSCULA

6b

subsD. nov.

Male.-Length, about 7 to 7.5 millimeters; wing, 6.5 to ?.8.
Femal,e.-Length, about 7 millimeters; wing, 7 to 7.2.
Generally similar to typical Limonta (Thrgpti,cornAln) aptcates (Wiedemann), differing especially in the larger size, whichapproximates that of. L. (7.) breai.anpi,s A,lexander. Legs withthe proximal fifth (forelegs) to third (hind legs) of the basitarsi
blackened. Wings nearly hyaline, with iridescent reflections,
the wing tip broadly and distinctly infumed; stigma elongateoval, dark brown ; veins black. Venation: Sc, long; R, relatively
long, more than two and one-half times R, alone; m-cu at near
two-thirds to three-fourths the length of cell lst M,. Male
hypopygium with the rostral prolongation of the ventral dististyle unusually long and slender, as in apicaJis; spines nearly
equal in size and slightly separated, the more proximal arisingfrom a basal tubercle.
Habitat.-Formosa.
Holotype, male, Toz,an, near Rato, November 5, 1928 (S. /ssiki,). Allotopotype, female. Paratopotypes, three males.
EELIUS (HELIUS) CEIKURINENSIS rp. nov.

General coloration dark brown; rostrum approximately as
long as the remainder of the head; wings with a brownish tinge;
male hypopygium with the outer dististy,le produced into a curved
apical point, with a small subapical tooth; gonapophysesappearing as pale flattened blades that are produced ,laterad into longacute points.
Male.-Length, about 4.8 millimeters; wing, 5.
Rostrum approximately as long as the remainder of head,
black; palpi black. Antenne black, the outer segments paling tobrown; basal flagellar segments short and crowded, the outer
segments more elongate, with long conspicuousverticils. Headdark brownish gray.
Thorax dark brown, the anterior lateral pretergites and adjoining regions more yellowish. Pleura dark brown, the dorsal
pleurotergite paler. Halteres dusky, the base of the stem narrowly yellow. Legs with the coxe and trochanters yellowish
testaceous; remainder of legs brownish yellow, especially thetarsi. Wings (Plate 1, fig. 3) with a brownish tinge, the stigmal region darker but ill-delimited; veins darker brown. Costal
fringe (male) relatively long. Venation: Sc, ending shortly before the fork of Rs, Sc, near its tip; veins R, and Raa6 strongly
250502-5

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The Phi,lippine Journal of Scienca

i:.taldiverging; inner end of cell lst M, more or,less pointed; m_cu
opposite r-m or nearly so.
Abdomen brownish black, the sternites paler. Male hypo_pygium (Plate 2, frg. 24) wittr the outer dististyle, od, a slender
blackened rod, narrowed to the curved acute apex, before thetip with a smaller denticle. Gonapophyses, gf, appearing as flat_
tened plates that are produced laterad into long apica,l spines.
.4Edeagus,q relatively short, straight, parallel-sided.
Habitat.-Formosa.
Ilolotype, male, Chikurin, altitude 8,000 feet, July g1, lg2g
(S. /sseka). Paratopotype, one male.
EELIUS(IIELIUS)CEIKURINENSIS MINUSCULUS subsp. nov.

Male.-Length, about 4 millimeters; wing, 4.
Generally similar to typical chi,ku,ri,nensrs,differing especiallyin the structure of the male hypopygium, where the gonapophy_ses (Plate 2, fir.25) are of quite different app"u"arcu, the apical point being bent across the blade.
Habi,tat.-Formosa.
male, Chikurin, altitude 8,000 feet, July 81, 1g2g
-Holotype,(5. Issi.lti,-y.
PEDIOITNTTRIcypEoNA

BAIKALICA .p. nov.

Large (wing, ma.le,lb millimeters); generalcolorationgray,the prescutum with three darker gray stripes; wings grayish,
with a heavy brown pattern distributed chiefly aloni ttr-ecostat
border and along the cord; cell lst M, present.
Male.-Length, about 1? millimetersf wing, lb.
Rostrum and palpi black. Antenne brack,the basarsegment
more pruinose; antennal segmentsapparenfly not .*.""dirrg
fourteen; flagellar segmentsshort and crowded,the basal ninesegmentsbroader than rong, the remainder smau and poorry
differentiated,provided with long conspicuousverticils. Headdark, pruinose; vertical tubercleconspicuous.
Mesonotal prescutum light gray, with ttrree darker graystripes; remainder of mesonotumdark gray, more or lessdiscoloredin the type. pleura dark gray, p"uirros"; dorsopieurat
region dark. Halteres yeilow. Legs with the coxa dark, pruinose; trochantensdark, the posterior trochanters paler; iu-o"ubrownish yellow, the tips broadry brackened;tibie darkbrown,
the tips blackened;tarsi black. Wings (plate l, fig. 4) grayish
with a heavy brown pattern that is chiefly costat, wiitr-"-rru"ybroad,irregular seamalongthe cord; the darkenedareasinclude
the prearcular region; cells c and sc; about the basal trri"o
ot

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Alenander: Tip.ulida from Eastern Asia, VI

67

cells R and M; a large area at origin of Rs, including most of cellRr; most of outer radial cells; seam along cord unusually wide,
including:all of cell lst Mr; anal cells less distinctly sutrusedwith
brown; veins brown. Venation: Sc, just basad of the originof Rs, the latter strongly angulated and spurred; r-m just before the fork of Rs ; R, transverse; Ra.'6shorter than the gently
sinuous r-m; cell 1st M, closed; m-cu before the fork of M.
AMomen dark, heavily pruinose; extreme margins of thesegments paler; hypopygium black. Male hypopygium (Plate2, fi4. 26) with the interbasal hook of the basistyle, b, a flattened cultriform blade. Dististyle, d, ,large and complex inform, roughly bilobed.
Habi,tat.-Siberia.
Holotype, male, Turan, Baikal, through Staudinger and BangHaas.
Tri,cEphona bai,lcali.cais very distinct from all described regional species.
TRICYPIIONA YAKUSEUTIANA sp, nov.

Size small (wing, male, not exceeding 6 millimeterp) ; general coloration brown; wings grayish yellow, clouded with pale
brown; R, oblique, sinuous, longer than R112i Rn captured by
Rz+g,Rz+a+rbeing nearly one-ttrird Rr*r; cell 1st M, closed.
Male.-Length, about 4.21o 4.5 millimeters; wing,5.6 to 6.
Rostrum and palpi brown. Antenne 16-segmented,the basal
segments dark brown, the outer half of the flagellum paling
to brownish yellow; antenne of moderate length, if bent backward extending approximately to the wing root; basal flagellar
segaents short and crowded, the outer segments elongate, with
more-conspicuous verticils. Head dark erayish brown.
Thorax high and gibbous. Mesonotum brown, very vaguelypruinose, without stripes. Pleura more brownish yellow, especially the anepisternum and sternopleurite in cases. Halteres
obscure yellow to dusky, the base of the stem restrictedly pale.
Legs with the eoxre and trochanters brown; remainder of legs
light brown, the tarsal segments somewhat darker. Wings(Plate 1, fig. 5) with a grayish yellow ground color, vaguely
clouded with pale brown, the stigmal region darker brown; thedusky olouds include all crossveins and deflections of veins. Venation: Sc, nearly midway between arculus and origin of Rs;
R, oblique, sinuous, longer than Rr* i Rr*r*n nearly one.third
Rz+si cell M, subequal to its petiole; cell lst M" closed; m-cu
about its own length beyond the fork of M; anal veins elongate.

The PhikWine'JournnL of Sci.ence

68

r980

Abdominal tergites dark brown, including the hypopygium;basal sternites paler, especially on the basal rings. Male hypopygium (Plate 2, fig.27) with caudal margin of tergite, gf, transverse, the lateral angles produced into very large conspicuous
hooks that terminate in a long straight spine. Basistyle, b,
bearing a fingerlike glabrous lobe on face at near two-thirds
the length; apex of style terminating in two low spinous lobes,
the small, ribbonlike dististyle, d, arising between these lobes.
(Kiushiu) .
Habitat.-Japan
Holotype, male, Kosugidani, Yakushima, altitude 2,b00 feet,
April 29, 1929 (S. Issi,hi.). Paratopotype, male.
Tri,cgphona Eakushi.mano is a very distinct species with no
known close allies. The venation of the radial field, especiallythe position and course of vein R,, is intermediate between thatof T. formosanfi Alexander and of the normal pediciine type,where R, is straight and has assumed a transverse or slighfly
obllque position.
HEXATOMINIuLAFLAVIDIBASIS

ep. nov.

General coloration of thorax dark gray, the mesonotal prascutum with three dark brown stripes; coxe blackened,pruinose;wings brownish yellow, the base and costal region ,light yellow;
a restricted dark pattern, chiefly along the cord; basal abdominal sternites light yel,low.
Femnle.-Length, about ? millimeters; wing, 8.4.
Rostrum and palpi black. Antennre black throughout, relatively elongate, the flagellar segments fusiform, with verticils that
only slightly exceed the segments in length. Head dark gray,the anterior vertex and narrow posterior orbits clearer gray.
Pronotum gray. Mesonotal prescutum gray with three darkbrown stripes that are dusted with a sparse yellow pollen; remainder of mesonotum chiefly dark gray. Pleura black, heavily
gray pruinose, the dorsop'leural membrane dusky. Halteres yel_low, the knobs infuscated. Legs with the coxe blackened,pruinose, the extreme tips and inner faces pale; trochantersinfuscated; femora black, the bases narrowly yellow, the amounton the forelegs including about the basal fifth; tibie and tarsi
dark brown. Wings (Plate 1, fig. 6) brownish yellow, the baseand costal region light yellow; a restricted brown pattern includes both ends of the light yellow stigma, the cord, origin ofRs, and outer end of cell lst Mr; veins brown, more yellowish
in the flavous areas. Venation: Rs angulated at origin; basal
section of R" reduced.

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Aleuander: Tigulida from Eastern Asia, VI

69

Abdominal tergites dark brown, blackenedlaterally, the terminal segmentsuniformly darkened; basal sternites uniformly
light yellow; ovipositorwith the valvesdark horn-brown.
Habi,tat.-Formosa.
Holotype,female,Arisan, a'ltitude6,500to 8,000feet, July ?,1929 (S. Isstki,).
Ula fl,annd,i,bosrls
is most closelyallied to U. perelegunsAlexander (northern Japan), differing especiallyin the colorationof
the thorax, legs, and wings. The almost uniformly blackened,heavily pruinose coxe of the present speciesare very conspicuous.
ADELPHOUYIAARIANA rD. nov.

General coloration black; antennal flagellum pale; wings witha strong brown suffusion; male hypopygium with the apex of
the basistyle produced into a slender point.
Male.-Length, about 4 millimeters; wing, 5.2.
Femnle.-Length, about 5 millimeters; wing, 5.
Rostrum and palpi brownish black. Antenna relatively longand slender for a member of the genus, if bent backward extending to slightly beyond the wing root; basal segments brownish black, the flagellum passing into pale yellow. Head black.
Thorax black, including the dorsopleural region. Halteresdusky, the base of the stem narrowly whitened. Legs with thecoxe dark; trochanters obscure whitish; remainder of legs pale
brown. Wings (Plate t, fi,g. 7) with a strong brown suffusion, the stigmal region slightly darker; veins dark brown.
Macrotrichia of cells relatively abundant, occupying the outerends of cells R, to lst A inclusive. Venation: Sc, relatively
long, exceeding m-cu; Rs strongly angulated at origin i Rz+
two to threetimes R, alone; cell M, present; m-cu at near midlength of cell lst Mr.
Abdomen black, including the genitalia of both sexes. Male
hypopygium (Plate 2, fig. 28) with the apex of the basistyle,b, produced into a slender point that is microscopically roughened. Outer dististyle, od, slender, dilated at apex, provided
with three slender denticles.
Habi,tat.-Formosa.
Holotype, male, Arisan, altitude 6,500 to 8,000 feet, July ?,1929 (S. Issi,ki). Allotopotype, female, altitude 7,300 feet.
Ailelphomyi.a ariona is readily separated from A. nipponensi,sAlexander (Japan: Honshiu) by the black body coloration anddarkened wings. The genus Ad,elphomyea,
is new to the fauna of
Formosa.

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The Phi,Wi,ne Journnl of Sci,ence

PSEUDOLIUNOPEILACEIKURINA

leao

ep. nov.

Generalcolorationbrownish gray; antenne black throughout;wings with a Sayish tinge, the stigrna vaguely darker; Rr*r*,
relatively short, about one-fourth longer than R1..2; m-cu shortly
beyondthe fork of M and before the,level of r-m; male hypopygium with the outer dististyle very slender and entirely blackened.
Male.-Length, about 4.5 millimeters; wing:, b.6.
Rostrum and palpi brown. Antenne (male) black throughout, of moderatelength, if bent backward extendingabout to theroot of the halteres; flagellar segments cylindrical, graduallyshortenedoutwardly, with long conspicuousverticils that muchexceedthe segments. Head brownish gray, the relatively broad
anterior vertex olearer gray.
Pronotum brownish gray. Mesonotal prescutum brownish
gray, the median stripe darker brown, the lateral stripes lessdistinct; pseudosuturalfovee black; tuberculate pits distinct;
pcsterior sclerites of mesonotumgrayish brown. Pleura brownish gray. Halteres obscure testaceous,the knobs dusky, thebase of the stem brighter. Legs with the coxe pale, slightly
pruinose, especially the fore coxa; trochanters testaceous,remainder of legs broken. Wings (Plate 1, fig. 8) with a grayish
tinge, the stigma only vaguely darker; veins pale brown. Costal fringe short. Venation: Sc, ending shortly before the forkof Rs, Sc, near its tip i Rz*r*o relatively short, only about onefourth longer than R112;Rs13 subequalto R, ; cell M, present,
longer than its petiole; m-cu shortly beyond the fork of M, be'fore the level of r-m; anterior arculus present.
Abdominal tergites dark brown, the sternites paler. Male
hypopygium with the outer dististyle very slender and entirely
blackened,gradually narrowed to the simple acute apex. Inner dististyle a little shorter, entirely pale, with abundant coarse
sete. Gonapophyses
bispinous.
Habitat.-Formosa.
Holotype, male, Chikurin, altitude 3,000 feet, July 31, 1928
(S. /ssi/ci).
Pseud,oli,mnophila
ehi,lartna is allied to P. i,nconcecso(Alexander), differing especiallyin the venationand the structure of
the outer dististyle of the male hypopygrum.
GenusNIPPOLIMNOPIIILA novum
Characters generally as in Limnophi,l,q,Macquart, diftering asfollows: Antenne ll-segmented (Plate 2, fig.29), both scapal
segments elongate and approximately equal in size; flagellar seg-

4e,rAleuander: Ti.Pulida from Eastern Asio, VI

7L

ments with relatively short verticils, not or only slightly longer
than the segmentsthemselves. Anterior vertex wide; head shortand broad, not narrowed behind. No tuberculate pits. Halteres very elongate, exceeding the length of the thorax. Tibial
spurs present. Wings (Plate 1, figs. 9, 10) with Sc relatively
short, Sc, ending before the fork of Rs, Sc, near its tip; cellM, lacking; anterior arculus preserved; anal angle lacking.
Veins pale, the macrotrichia much redueed in size and number.
Male hypopygium (Plate 2, fi$.30) with both dististyles simple;
tergal reg:ion evenly convex; eighth sternite produced ventrad,
the edeagus, o,, strongly deeurved, the tip recurved and directed
toward the notch of the eighth sternite.
Genotype, Nippolimnophi,la lciwiuensds sp. nov. (Palaarctic
Resion).
N i,ppol;i,mnoplt
i,Lag aku,shimensassp. nov. likewise belongs here.
The chief generic characters ,lie in the strueture of the antenne which are here reduced to an ll-segmented organ, withthe secondseapal segment (pedicel) of an unusual length. The
only other crane flies with an elongate pedicel are found in theeriopterine genus Chionea Dalman and allied forms. The factthat the entire series of specimens of the two species referredto this genus, totaling more than a score of individuals, included
only males may indieate that the female is subapterous.
NIPPOLIMNOPEILAKIUSIUENSIS

sp. nov.

Generalcolorationbrownish gray; antenne short; cell 2d A
relatively long:and narrow.
Male.-Length, about 3.5 to 4.5 millimeters; wing, 4.5 to 6.
Rostrum and palpi black. Antennre black throughout, short(Plate 2, fig. 29), if. bent backward not extending far beyond
the pronotum; flagellar segmentsoval, with short verticils thatslightly exceedthe segrnentsin leneth. Head brownish gray,the orbits clearer gray.
Mesonotumbrownish gray, the prescutal stripes lacking ori'll-defined. Pleura brownish gray. Halteres pale, the knobs
dusky. Legs with the corcreand trochanters brown, pruinose;
remainder of legs dark brown, the femoral basesvery restrictedly pale. Wings (P'late 1, fig. 9) milky white, the stigxnaand
vague seamsat origin of Rs, along cord and outer end of eelllst M, pale brown; veins brown. Venationi Rz+g+r
nearlytwice the basal section of Ru; veins issuing from cell 1st M,
somewhatdivergent; m-cu at near midlength of cell lst M,; vein
2d A relatively elongate,the cel,lnarrow.

The Phi,liWi,neJw,nwL of Seience

72

1es0

Abdomen, including the hypopygium, dark brown. Male
hypopygium (P,late2,fig.30) with the outer dististyle od.beav
ing a series of conspicuoussete along the outer face, the darkened apex simple, glabrous. fnner dististyle a shorter rod,heavily blackened,terminating in a slighfly dilated head.
Habitat.-Japan (Kiushiu).
Holotype, male, Mount Kirishima, altitude 2,b00 feet, May B,
1929 (S. Issilci,l. Paratopotypes,1 male; b additionalmalesat
3,000to 3,500feet, May 4,1929 (S. Issiki,).
NIPPOLI,INOPf,ILA

YAKUSEI}TENSIS

3p. nov.

Male.-Length, about 4 to 4.4 millimeters; wing, 4.d to 6.9.
Generally similar to N. ki,usiuemsassp. nov., differing especially in the elongate, anf,enna of the male rvhich are here approximately as long as the thorax, if bent backward extending about
to the root of the halteres; flagellar segments fusiform.
Wings(Plate 1, fig. 10) with cell 2d A somewhat shorter.
Habitot.-Japan (Kiushiu).
Holotype, male, Kosugidani, Yakushima, altitude 2,b00 feet,
April 29, 1929 (S. Issi,ki,). Paratopotypes, 20 males.
ULODIORPITAPOLYTRICEA

rp. noy.

General coloration polished black; halteres yellow; wings
tinged with brovrnish, scarcely variegated with darker; macrotrichia of rnembrane very numerous; cell M, lacking.
Male.-Length, about 7.5 millimeters; wing, ?.6.
Rostrum shiny black, the mouth parts paler; palpi brownish
black. Antenne relatively long and slender, the scape black,the flagellum pale brown; flagellar segments elongate-cylindrical, with verbicils that $rceed the segments. Head shiny black.
Pronotum and mesonotum shiny polished black. Pleura black,the sternopleurite paling: to brown. Halteres pale, the knobs
light yellow. Legs with the coxe obscure yellow, the fore coxe
more darkened basally; trochanters yellow; femora obscureyellow, the tips narrowly infuscated, more broadly so on thefore femora; tibie and tar:si brown; leg:s conspicuously hairy.
Wings (Plate 1, fig. 11) tingd with brownish, the base andcostal region clear yellow; a scarcely indicated darker clouding
along the cordi stigma small, darker than the ground color;veins brown, light yellow in the luteous areas. Conspicuous
macrotrichia in almost all cells of wing, lacking only in the extreme bases of the chief basal cells a,nd in the prearcularr andsubcostal cells. Venation: Rs angulated but scarcely spurrecl

4z,rAlerander: Tipuliila from Eastern Asia, VI

73

at origin i Rz+a+r shorter than the basal section of Ru, the veins
beyond it longer and less divergent than in ni,gNcolor; cell M,
lacking; m-cu at near midlength of cell lst M,.
Abdomen reddish brown, trivittate with black; hypopygium
black.
Habitot.-Japan (Kiushiu) .
Holotype, male, Kosugidani, Yakushima, altitude 2,500 feet,
April 29, 1929 (S. Iss'tki).
Ulomorpha polEtricha is readily distinguished from tJ. ni.gricolor Alexander (Japan: Honshiu) by the nearly immaculate
wings with more-abundant macrotrichia. In both wings of theunique type there is a short marginal spur in cell 2d M, that
presumably represents the partly atrophied vein M,.
ERIOCEBA PLATYSOMA aD. nov.

Abdomen of male dilated and flattened; general coloration
velvety black, the basal four segments of abdomen orange;
wings, legs, and halteres black.
Male.-Length, about 16 millimeters; wing, 14, width of abdomen, 4.5.
Rostrum, palpi, and antenna black, the last 7-segmented,,theflagellar segaents decreasing in size outwardly, the terminal
segment abruptly smaller, about one-fifth the penultimate.
Head velvety black.
Thorax deep velvety black, the scutellum dull orange. Halteres and legs entirely black. Wings blackish, the outer halfof the wing more suffused, the medial, cubital, and anal cells
more streaked with dusky; veins dark brown. Venation:
Rs long, weakly angulated at orig:in; Sc, elongate, exceeding
R +s+.i Re+asubequal to Rz+s+n;cell M, Iacking.
Abdomen broadly dilated and flattened, as shown by themeasurements; segments one to four dull orange, the caudal
margins of segments two to four narrowly and evenly blackened,
the lateral flanges of the segrnents slightly infuscated; remainder of abdomen deep velvety black, the hypopygium brown.
H abitat.-Western China.
Holotype, male, Kwanhsien, Sze-chwan, October 7, \g?g,through Mr. Herbert S. Parish.
Eriocera platysoma is allied to E. ubilom,inalis Alexander, likewise from western China, in the great lateral development of theabdomen of the male (though presumably not of the female).
It is readily told by the blackened wings and entirely black legs.

74

The PhikWi,ne Joumal, of Sci,ence

ELEPEANTOMyIA(ELEPEANTOMYTA)

rego

SDROTINA rD, nov.

Rostrum nearly as long as the remainder of the body; mesonotal prescutum obscureyellow, with a broad median darkbrown stripe that widens behind; a dark transversegirdle on thepleura; halteresdusky; femora extensivelyblackened,especiallythe fore femora; wings grayish yellow,the stigmalong_oval,darlbrown; abdominaltergites obscurebrownish yellow, darker medially, the caudal margins of the segmentsna*owly but completely borderedby dark brown.
Femnle.-Length, excluding rostrum, about g mil,limeters;
wing, 9.8; rostrum, about T.
Rostrum elongate,only a litile shorter than the remainderof
the body, black throughout. Antennreyellowish brown. Headbuffy brown; anterior vertex narro,w.
Mesonotalprescutum obscureyellow, with a broad mediandark brown stripe that widens behind; scutal lobes extensively
dark brown; median region of scutum and the scutellum moretestaceous-brown;
postnotalmediotergitebrownishblack. pleura yellow, the sternopleurite and anepisternum dark brown to
produce a more or less distinct girdle. Halteres dusky, the extreme base of the stem yellow. Legs with the coxe yellow,
the outer face of the middle coxa darke,r; trochanters yellow;
fefirora chiefly brownish black, the base,smore yellowi,sh, ,a"rowsst on the forelegs where about the basal fourth iS included,
hroadeston the posterior legs where only the tips are blackened; tibie dark brown, the tarsi paling to ob,scurebrownish
yellow. Wings with the ground color grayish yellow, ttre prearcular' and costal regions more yellowish; stigrna long-oval, darkbrown; distinct paler brown sea''x along the cord and oufurend of cell lst Mr; wing apex narrowly seamedwith brcxnrn;
veins dark brown. Venation: Cell lst M, largg subrectangular;
m-cu more than one-half its length beyond trhefork of M, longer
than the distal section of Cur; cell 2d A relative,ly narrow.
Abdominal tergites obscurebrownish yellow, t-hemedian region darker, including the dark brown caudal marginsof thesegments;sternites yellowish, the caudal marginp of the segments narrowly dark brown; ovipositor yelrowish horn eoror,
the tips of the slender upturned tergal valves narrowly darkened.
Habitat.-Formosa.
_Hglotype,female, Yrlsho, altitude 6,000feet, October 24, Lgzg
(5. Issilci). Paratype,teneral female,Shdrei,altitude Z,6OO
to8,000feet, October25, lgZS (S. Issa&a).

42,1

Ale*ander: Ti,pulida from Eastern Asia, VI

75

Elephantom,yia seroti.na is allied to E. hokkwid,ensisAlexander(northern Japan), differing especially in the coloration of the
body and legs.
ERIOPTERINICEBATOCIIEILUS

TINCTIPENNIS

sp. nov.

General coloration black, pruinose, the prescutal shield darkened; rostrum, antenne, and legs black; knobs of halteres infuscated; wings strongly tinged with brown.
Femnle.-Length, excluding rostrum, about 7.5 millimeters;
wing, 6.5; rostrum, about 4.5.
Rostrum black. Antenne black throughout. Head light gray,the posterior vertex darker gray; anterior vertex nearly twice
as wide as the diameter of the fusion segment of the antenna.
Mesonotum blackened, the humeral region of the prescutum
somewhat more pruinose; median region of the scutum and thescutellum a trifle more pruinose. Pleura black, pruinose. Halteres dusky, the knobs infuscated. Legs with the coxe obscurebrownish yellow, the outer face more infuscated; trochantersobscure yellow; remainder of legs black. Wings (Plate 1, fig.
L2) with a strong brown suffusion, the veins dark brown. Macrotrichia of veins relatively numerous, there being a series ofabout four or five on each of Rs and the basal section of Ru;
none on anterior branch of Rs. Venation: Anterior branch of
Rs relatively long and sinuous; m-cu before the fork of M.
Abdomen black, pruinose. Ovipositor brownish horn color.
Habitat.-Formosa.
Holotype, female, Arisan, altitude 6,500 to 8,000 feet, July 7,
1929 (S. Issiki).
Cero,tocheilus tinctipennis is readily distinguished from the
known reg'ional species by the coloration of the body and thestrongly tinted wings.
LIPSOTERIXYAKUSEIM.IE

sp. nov.

General coloration dark brown; halteres dusky, the knobs darkbrown; legs brown, the tarsi extensively whitened; wings witha strong dusky suffusion; Rs and Rr+r*nnearly equal in length,
pale, with few or no macrotrichia.
Femnle.-Lengi'th, about 8.5 millimeters; wing, 8.2.
Mouth parts much reduced. Antennre black throughout; flagellar segments oval, with long verticils; terminal segment reduced. Front yellowish testaceous, the posterior portion of thehead darker, the disk of the vertex dark brown.
Thorax dark brown, the median region of the scutum moreyellowish; dorsopleural region pale. Halteres dusky, the knobs

76

The PhikWine, lournal of Sciencer9'i 0

dark brown. Legs with the coxe and trochantersbrownishtestaceous;femora and tibie brown,
the genu,auno-nu""o* tip*of the tibie paler; tarsi with the out""
segmentsconspicuously
whitened. Wings (plate 1, fig. igt
witt * .trong-Au"iv suffu_
sion; a conspicuouswhitish .t"u"lin cell M, erossing m_cu
into the base of cell M,; veins t;;*".
Veins Rs and &+g*r
pale and without macrotrichia
or nearry so. v.nuuonr ns onrya trifle longer than R21s1o;
innerof ceil Lst M, slighily ar_cuated; m-cu a short distancebeyond""a thefork ofAbdomenbrownish brack,the genitarsegment m.
obseurebrowniBh yellow. Ovipositor with the-;Jves
blackened.
H_abitat.-Japan ( Ki ushiu)
Holotype,female,_Kosugidani,
yakushima,altitude,2,800feet,
Ap_ril 29, tg?g (5. Issikif.
..Lipsothrin garntshirna is most croseryrerated to L. taiwani,caA'lexander(southern Formosa), iin""ingin the rarge size anddetails of colorationand uun"tion.
ihe grabrousveins Rg and
Rz+g+lare conspicuous
features.
GONOMYIA(PTILOSTENA)CURTICELLULA

SP. NOV.

Generalcoloration_ofbody dark gray; rostrum
and antennal
scapedark brown; thoraeic pleura abrup'yyAlo*; *ing*
nafewith a grayish tinge, the stigma pale; cell R.
very small; ma,lehypopygiumwith the eedeagus
pale,not markedrychitinized;four
dististylesor branchesof the ,"rn..
Male.-Length, about 4 to 4.5 millimeters;
wing, 4.g to b.b.
Rostrum and palpi dark brown. Antenna*iif, tfr. scapebrownish black, the flagellum paler brown;
flagellar segments
elongate-fusiform,with a denseereci pubescence.
Head darkgray.
Pronotum and mesonotumdark gray, the prescutumwith theinterspaceseven darker brownisliiiu, pseudosuturalfovere
conspicuous;centers of the scutai lobes
darkened. preura
abruptly pale yelow, the dorsarscleritesmore
dusky. Harteresdusky' Legs with the coxe and trochanters parev.lio*; 1urno""
yellow, the tips narrowly dark brown;tibie obscureyellow, thetips_infuscated; tarsi passinginto darkbrown. Win"*-ipfut"
1, fig' 14) relativery narrow, with a grayish
tinge, the entire
costal margin to the apex Iight
vellow; ,iig-" u""v p"f", onfy
a little darker than-the ground coror; no distinct
dark crouds
on_membrane,but the-origin of Rs, cord,
and __.u a""t." incolor than the remaining veins; veins pale
brown, the costal
veins yellowish. Venation.. S",'ending
shorfly beyond originof Rs, Sc, far from its tip, iust before midway
betweenarculus

4z,rAlenander: Tipul,ida from Eastern Asia, VI

77

and origin of Rs; R, nearly perpendicular; cell R, very small;
m-cu more than its own length before the fork of M; Cu, extending to opposite or beyond m-cu.
Abdominal tergites brownish gray, the sternites obscure yellow; hypopygrurn yellow. Male hypopygium (plate 2, fig. P;L)with four arms to the dististyles, d, of which three are quite
glabrous; outermost a slender rod, its apex darkened;
Iongeststyle with the margin darkened, the apex obtuse; thiri branchbifid, one arm shorter and obtusely rounded. The fourth styleor branch is slender, terminating in a curved apical spine, thesurface of the style with about eight sete. .lBdeaeus, a, longand slender, entirely pale.
Habi,tat.-Formosa.
Holotype, male, Giran, November 2, Lg2g (S. /ssz/ci). para_
type, male, Taihoku, May 8, lg2g (S. Issiki)
Gonomgin ctnti,celluln differs rather conspicuously from G.
pruinosa Alexander (Formosa) in the nearly immaculate
wings(Plate 1, fig. 15) and unusually small cell Rr. The paratype
has the wings narrower, with correlated differences in venation,
as a shorter Rs and cell 2d M, and with m-cu a greater distance
before the fork of M. Despite these differen.er, I cannot separate the material from the type.
GONOMYIA(PTTLOSTENA)SEANTUNGENSIS

SP. NOV.

General coloration of notum brown, pruinose; basal segmentsof antenne pale, the outer gegments black; pleura yeUiv,
ttre
dorsal region darker; wings pale brownish yel,low, the stigmaalittle darker; Rs angulated at origin; abdominal tergites darkbrown, the caudal margin of the segments narrowry ringed
withyellow; sternites yellow; male hypopygium with theLd.ugu,entirely pale, straight, at apex curved into a genfle crook.
Male.-Length, about 4.b millimeters; wing, b.4 to 5.b.
Rostrum and palpi black. Antenna with the scape obscureyellow; basal segments of flagelum likewise pale, theoutei segments passing into brown. Head dark gray.
Pronotum dark brown, sparsely pruinose. Mesonotal prescutum with the ground color reddish, the humeral and lateralregions more yellowish, with a broad conspicuous dark
brown
stripe, the surface of the solerite a ritfle pruinose; pseudosutural
fovee large, reddish brown; scutum brown, the centersof thelobes vaguely darker; scutelum and poshrotum dark, sparsery
pruinose. Pleura pale yellow, the dorsal region
infuscated.
Halteres dusky. Legs with the coxre and trochanters paie yetIow, the fore coxa a litde darker; femora and tibie y.ttto*,
tt"

78

The PhiliWi,ne Journal of Seienca

1e30

tips of the latter narrowly darkened; basitarsi obscure yellow,
the tips narrowly darkened; terminal tarsal segments broken.
Wings very pale brownish yellow, the base, costal region, andouter ends of the radial cells c,leareryellow; stigma small, brown,
only a little darker than the ground color; veins brown, the cordand basal portion of Rs darker. Venation: Sc, ending shortly
beyond the origin of Rs, Sc, far from its tip; Rs ahgulated atorigin; petiole of cell R, angulated before midlength : R. about
one-third its length from R ,*, * margin; m-cu more than its
leneth before the fork of M.
Abdominal terg:ites dark brown, the segruents narrowly andindistinctly ringed caudally with pale yellow; sternites pale yellow. Male hypopygium (Plate 2, fir. 32) with the basistyle,b, lobed at apex, including a broad obtuse ventral lobe and asmall mesal lobuile. Three dististyles, d, the longest U-shaped,
both arms acute and glabrous, the shorter inner arm more blackened at apex; second dististyle broadest, the apex bifid, botharms blackened,the outer longer and slenderer, the inner obtuse;
third dististyle a short simple pale rod that is narrowed at apex
into a blackened spine. .iEdeagus, o, entirely pale, appearing
as a straight rod, the apex curved at more than a right ang,le.
H abi,tat.-Eastern China.
Holotype, male, Lau Stan, east of Tsingtao, Shantung, June
17, L927, taken in evening (4. P. Jacot),
Gonomgi,a shantungensis is closest to the Japanese G. subpruinosa Alexander and G. pallens Alexander, differing from all
regional species in the structure of the male hypopygium.
ERIOPTERA(DRIOPTERA)ENSIFERA

ap. nov.

General coloration yellow; thoracic pleura with a bluish sheen;
knobs of halteres infuscated; legs yellow with only the terminal
tarsal segments darkened; wings brownish yellow, with a restricted darkening along the eord; macrotrichia of veins longand conspicuous; male hypopygium with the gonapophyseslong
and ensiform, entirely pale in color.
Male.-Length, about 2.8 millimeters; wing, 3.8.
Femnle.-Length, about 3.5 to 4.5 millimeters; wing, 3.8 to
4.5.
,
Rostrum brownish yellow; palpi black. Antenne pale brown.
Head cinnamon brown.
Mesonotal prescutum light cinnamon brown, with scarcely indicated markings, the humeral region more yellowish; scutellum
and postnotum more yellowish, with a narrow brown medianline in the male, this not indicated in the female. Pleura pale,

42,tAlerander:

Tifiul,idu from Eastern Asia, VI

79

with a distinct bluish sheen, most evident when viewed fromabove and behind. Halteres pale, the knobs infuscated. Legs
yellow, the terminal two segments dark brown. Wings (plate
1, fig. 16) tinged with brownish yellow, the costal region clearer
yellow; a vague dusky seam along the cord; veins yellow, thecord infuscated. Macrotrichia of veins long and conspicuous.
Venation: Vein 2d A strongly sinuous.
Abdomen, inc,luding the hypopygium, yellow. Male hypopygium (Plate 2, fir. 33) with the median lobe of the terg:ite,
9t, quadrate, transverse. Outer dististyle, od, with the inner
blackened setiferous arm small, the outer arm unusually longand slender, subequal in length to the main axis of the style.
Gonapophyses appearing as long entirely pale, sword-shaped
blades that narrow to acute points.
Habi,tat.-Formosa.
Holotype, male, Taihoku, May 8, 1929 (S. /ssi/cf). Allotopotype, female. Paratopotype, female.
Eri,optera ensifera belongs to a large group of Oriental species
that are allied to E, notata de Meijere. It seems probable that
this is the species recorded by Riedel as E. flaao Brunefti,2from Taihoku. This species,which was ilater renamed E. bengalensis Alexander, is emphasized as having the halteres entirely
pale and I cannot accept Brunetti's later conception that it isidentical with E'. hnlterata Brunetti, a Himalayan species withdarkened halteres. The present species is certainly different
from all other regional forms from low altitudes in the Malayan
islands (8. iaaaneicsis de Meijere, E. notata de Meijere, E. Luzonl.caAlexander).
ERIOPTERA (ILISIA)TENUISDNTIS sD. roy.

Male.-Length, about 5 millimeters; wing, b.?.
Closely allied to E. (1., asymmetrtca,Alexander (Japan), differing especial,lyin the structure of the male hypopygium.
Antennae with the outer four or five segments infuscated. Mesonotal prascutum with the intermediate dark stripes narrow,entire. Dorsal pleural region more darkened. Legs with thefore and middle femora chiefly blackened, the subterminal yelIow ring much reduced. Wings (Plate 1, fig. 1T) with the pattern restricted to dark spots and dots along the veins. Male
hypopygium (Plate 2, fig.34) with the apical hooks of the ninth
tcrgite, 9f, very long and slender, strongly .recurved, very gradually narrowed into long acute points. Gonapophyses,gf,heavily'Archiv

fiir

Naturgeschichte 92, Abt. A(191?) 118.

80

The Phili,wine Journal of Science:,lao

blackened,graduallynarrowedto acutepointg,the mesalface notdilated.
Habi,tat,-Formosa.
Holotype,male, Arisan, altitude 7,300feet, July 7, 1929 (S.
Issi,lti,).
The subgenusllisia is new to the fauna of Formosa.
EBIOPTERA (EMPEDA) MICROTRICEIATA

sD. noy,

General coloration gray; halteres pale yellow throughout; legs
brown; wings tinged with gray, the diffuse stigmal region darker; Sc long, Sc, ending beyond midlength of Rs; macrotrichia
of veins relatively short; male hypopygium with both dististyles
pale.
Male.-Length, about 3 millimeters; wing, 3.4 to 3.7.
Female.-Length, about 3.5 to 3.8 millimeters; wing, 4 tn
4.2.
Rostrum and palpi brown. Antenna dark throughout, in themale with very long verticils, as usual in the subgenus. Headdark gray.
Mesonotum gray, the anterior lateral pretergites restrictedly
yellow. Halteres pale yellow. Legs with the coxe and trochanters obscure yellow; remainder of leg:sbrown, the terminal tarsal
segments darker. Wings with a grayish tinge, the large diffuse
stigma darker; veins brown. Macrotrichia of veins short andinconspicuous. Venation: Sc long, Sc, ending just beyond midlength of the relatively long Rs; Sc, subequal to r-m; R, subequal to or a little shorter than R213*n; cell R. relatively deep
for a member of the subgenus.
Abdomen dark brown, including the hypopygium. Male hypopygium (Plate 2, fig. 35) with the dististyles pale, the outer
oististyle, od, deeply bifld, the outer arm longest, curved to anacute point, the outer surface roughened; inner arm expanded
at apex, the outer angle produced into a point. Gonapophyses,
g, blackened.
H abitat.-Japan (Kiushiu) .
Holotype, male, Mount Kirishima, altitude 2,500 feet, May 3,
1929 (S. Issiki). Allotopotype, female; paratopotypes, 2 males,
females. Paratypes, 3 males, Kosugidani, Yakushima, altitude2,500 feet, April 29, 1929 (S. Issi,ki).
Erioptera (Empeda) nui,crotri,chinfo is distinguished from .8.
(8,) iaponi,co Alexander by the short macrotrichia of the wingveins and the uniformly pale halteres. It differs from E, (8.)
ungusti,sti,gma,sp.nov.in the grayish wings and paler ,legs.

42,rAlenunder: Tipulidp from Eastern Asia, VI

81

ERIOPTERA (EMPEDA) ANGUSTISTIGUA sp. nov.

Generalcolorationgray; mouth parts and antenna black; halteres pale yellow; legs black; wings light yellow with a narrowbrown stigma; Sc long, Sc, extending to beyond midlength of
Rs.
Female.-Length, about 3.8 millimeters; wing, 4.
Rostrum and palpi black. Antenn'a short, black throughout.
Head gray.
Anterior lateral pretergites dusky, the lateral margins clearer
yellow. Mesonotumgray, the humeral region of the prescutum
more reddish gray; pseudosuturalfovee reddish brown, the tuberculatepits darker. Pleura dark plumbeousgray, including
the dorsopleural region. Halteres pale yellow throughout,clothedwith silken golden sete. Legs with the coxa brown, thefore coxe darker brown; trochanters brownish testaceous;remainder of legs brownish black. Wings (Plate 1, fig. 18) lightyellow, with a narrow but well-definedbrownish stigma; veins
darker yellow than the ground color. Macrotrichia of veinsrelatively short and inconspicuous,
brown. Venation: Se long,Sc, ending beyond midlength of Rs, relatively short; Rz+a+r&
little longer than R, and approximatelythree times R21r; cellM, open by the atrophy of m; m-cu at fork of M.
Abdomen dark brown; tergal valves of ovipositor broken;
sternal valves blackened,long and straight.
Habi,tat.-Formosa.
Holotype, female, Arisan, altitude 7,300 feet, July ?, 1929
(S. /ssa&tl).
Eri,optera (Empeda) angustistigzaodiffers from the known
Formosan speciesin the coloration and wing venation. It is
more closely allied to a small group of speciesknown from themain islands of Japan, ditrering in the diagnostic features listed
above.
MOLOPUILUS NIGRITUS sD. rov.

General coloration black; halteres dark brown; wings strongly
tinged with blackish.
Female.-Length, about 4.5 to 5 millirneters; wing:, 4.b to b.
Rostrum, palpi, and antenne entirely black; flagellar segments
subcylindrical, passing into long-oval; terminal segment smaller
than the penultimate; verticils somewhat long:er than the segments. Head black, sparsely gray pruinose, especially in front
and on the posterior orbits.
2506,02-6

82

The Phil;i,Wi,ne Journal of Saience

Mesonotum black. Pleura black, sparsely pruinose. Halteresdark brown throughout. Legs blaek. Wings with a blackish
tinge, the base not brightened; veins and macrotrichia darker.
Venation: Vein 2d A gently sinuous, ending about opposite thecaudal end of m-cu.
Abdomen black, the bases of the valves of the ovipositor similarly colored, the remainder of the long slender valves yellowish
horn color.
Habitat.-Formosa.
Holotype, female, Arisan, altitude 7,300 feet, July 7, 1929 (S.
Issiki,). Paratopotypes, 1 female with type; 1 female, altitude
6,500 to 8,000 feet, July 7, 1929 (S. Issiki').
Molopltilus ni,gri,tu"sis readily distinguished from regional species by the coloration. It is most similar to M. trifi,tntus Alexander (Japan), differing: in the coloration of the wings andlegs.
MOLOPHILUS

NIGRITARSIS

aD. nov.

Female.-Length, about 4 millimeters; wing, 4.3.
Closely allied to M. costali,s Edwards (Formosa), differing
especially in the coloration of the body and legs.
Rostrum and palpi black. Basal segments of antenne yellow,
the remainder darkened. Head yellow, the center of the vertex with a conspicuous brown area. Anterior lateral pretergites
and extreme lateral margins of the prescutum sulphur yellow.
Mesonotum plumbeous brown. Pleura chiefly dark brown, thepleurotergite more yellow, bearing a group of long yellow sete.
Halteres yellow. Legs with the fore coxa darker than the
middle and hind coxe; forelegs brownish black; middle femora
and tibie yellow, the tips of the latter and all tarsi blackened;
posterior femora extensively infuscated, the narrow tips andbroader bases yellowish; tibiae bright yellow, the tips and allof tarsi blackened. Wings grayish, the prearcular and eostal
regions light yellow, this coloration including the veins andmacrotrichia; remaining veins and macrotrichia dark, excepting
vein Cu which is yellow. Abdomen brown, the caudal marginsof the segments narrowly and indistinctly more yellowish.
Habi,tat.-Formosa.
Holotype, female, Arisan, altitude 7,300 feet, July 7, 1929 (S.
Issilci.).
The pattern of the legs is distinctive of more than a single
species in the Formosan fauna.

ILLUSTRATIONS[L€er€nd; o, ed€agus; b, basistvle;4
oute! dististyle; s, sternlt€ i t, tergite;
crmPton'l

PT,ATEFrc.

g, gonapophyeis;

dististyle;od, ventral

dististyle.

fd, inner dististyle; od,
Hypopygialt€rminology u6ed,

1

1. Li,monia, (Gerarwm,gia) rad:inlis sp. nov., wing.
2, Linloni,a (Geranomyi.a) atpestris sp. nov., wing,
3. Heli,us ch,i,la,Lri,nmsissp. nov., wing.
4. Tricgphona baikal:ia, sp. nov., wing.
5. Tri,cyphonn gaktnhimiann sp. nov,, wing.
6. ULa flan:id.ibosis sp. nov., wing.
7. Ad,elphom4ia arfa,na sp. nov., wing:,
8. Pseudnl:imnnphila chiku,rtno sp. nov., wing.
9. Nippoli,mnoplui.la kiasiuensis sp. nov., wing.
10, Nippolimnoplr;i,la gakuslri,mpnsis sp. nov., wing.
71, Ulom.orpha polgtri.cha, sp. nov., wing.
12. Ceratocheilus tinntiperlrlis sp. nov., rfling:.
73. Lipsothria gakuslui,ma,esp. nov., wing.
L4. Gonomyin (Ptilostenn\ curti.cel,Iula sp. nov., wing.
75. Gartomgi.a (Ptil.ostenal ynti,noso Alexander, wing.
76. Erioptera (Erioptera)
ensif era sp. nov,, wing.
17. Eri.optera (Ilisi.a,) tetr,uisentie sp. nov., wing.
18. Erioptera (Empeila) angustistigma, sp. nov., wing.
Pr,lrp2

Flc. 19. Tiaula udnoi sp. nov., male hypopygium, lateral.
20. Tipila u6noi sp. nov., male hypopygium, ninth tergite.
21. Tipula udnoi sp. nov., male hypopygium, eighth sternite.
22, Litmotvin (Libnotes) reeuroinerais sp. nov., male hypopygium.
23. Lim,onia (Geranom4yia) rad:ialis sp. nov., male hypopygium.
24. Helitu,s chikurinercis
sp' nov., male hypopygiunr"
26, Eeli,us ch,i,latri,nerwisminu,scuhts subsp. nov., male hypopygium, gonapophysis.
26. Tricyphona baikali,ca sp. nov., male hypopygium.
27. Tricgphona yaktuhim&ne sp. nov., m,ale hypopygium.
28. Ad,elphnm4ia artana, sp. nov., male hypopygium.
29, Ni,p.polimnophila hiusi,uercsis sp. nov., antenna.
30. Nippoli.mnophila lciusiuezst's sp. nov., male hypopygium.
3L. Gtnom,gia (Ptilostmn)
curtiaellula sp. nov., male hypopygium.
32. Gonomyia (Pti,losterw) shantungensis sp. nov., male hypopygium.
(Erioptera)
33. Erioptera
erxi,fera sp. nov., male hypopygium.
34. Eri.optera (Ilisi.a) tenuisentis sp. nov., male hypopygium.
35. Erioptera (Empeda) nvicrotrichia,t@ sp. nov., male hypopygium.
83

ALEXANDER :TIPULIDAI

FRoM

EASTERN

ASIA,

VI.]

PLATE I.

lPHrLrP. JouRN. Scr., 42, No.

1.

ALDXANDER:TIPUIJD.E

I'RoM

EASTERN

AsI-{,

lPHrLrP. JouRN. Scr., 42, No.

VI.l

A

,/\
\/
\

9rl
-;

f----=z-\
8s\

I

21

{r\
/-t'\

"kK
"[fltr23'-@+e,P*..Fr .' .\<

I" . ',.\
i,.". :j\bt.f.',) ^"do..\./{I4frk'\
26

il-,v9Y{Jg\Gii..\\),'\'rtrytrN' /zJ\
30

"'/6t
,1,d
W

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4.t.

( .." " ',rML?ff,'[{cJ,'

I

or \

34
PLATE 2,1.