$'-ig

Reprinted from Bur,r,nrrNoF run BnooKLyNEnrouor,ocrcAr,Socrnnv,
Vol. XLY, No. 2r pp. 4l-47. April, 1950.

NEW OR INSUFFICIENTLY-KNOWNCRANEFLIES FROM THE NEARCTIC REGION(DIPTERA, TIPULIDAE).
PART XII.1
By C. P. ATExANDER,
Amherst, Mass.
The precedingpart under this generalseriesof paperswas publishedin 1949 (Bull. Brooklyn Ent. Soc.,44: I52-I57). At thistime I am describingfour noveltiesfrom the mountainsof southeastern Arizona, collectedby my friends Messrs. Owen Bryant,Willianr Nutting and Floyd Werner, to whom I am greatly indebtedfor the privilege of retaining the types in my personalcollectionof theseflies.
Tipula (Bellardina) catalinensis n. sp.
Mesonotalpraescutumbuffy gray, with four brown stripes,themedian interspacemore reddened,the lateral ones clearer gray ;scutellumbrownish yellow, with a narrow brown central line ; antenna with scapeand pedicelyellow, flagellum dark brown ; femora
obscureyellow, the tips narrowly brownish black; wings marbled,brownish yellow, variegatedwith crearn-yellowand sparsedarker
brown areas; abdorninaltergites yellow, vaguely trivittate withpale brown, the outer segmentsbrownish black; male hypopygiumwith the caudal border of the tergite very shallowly emarginate,with a further more ventral median lobe; ninth sternite with asmall lyriform medianappendage;outer dististylea large flattened
curved blade; inner dististylesmall but complex,the beak slender.
Male; Length about 18 nun.; wing 19 mm.; antennaabout 3.5mm.
Frontal prolongationof head obscurebrownish yellow, slightly
pruinose dorsally; nasus elongate; palpi brown, the incisuresrestrictedlypale. Antennaerelativelyshort; scapebrownish yellow,
corrugated; pedicelcleareryellow, flagellunrdark brown ; flagellar
feeblyindisegrnentssubcylindricalor with the basalenlargernents
cated; verticils longer than the segments. Head above brownishgra)-, the sides of the occipital region more yellowed, the vertex
with a brown central stripe; vertical tuberclevery low.
Pronotal scuturn weakly infuscatedrnedially, more yellowed on
1 Contribution from the Department of Entomology, Universityof Massachusetts.

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sides; scutellurnand pretergitesclearer yellow. Mesonotal praescutum buffy gray with four brorvn stripes, the rnedian interspace
more reddened,slightly broader at and beyond nridlength; lateralinterspacesclearer gray, the reddish lateral stripes narrowly bordered internally by brown ; scutal lobesdark gray, the central area
obscureyellow; scutellumbrolvnish yellorv,with a narrorv brou'ncentral line, parascutellabror,vnishtestaceous;postnotutn graypruinose,the cephalichalf less so. Pleura and pleurotergitepale
brown, sparsely pruinose, the propleura and dorsal mesopleuraslightly darker to form a weak stripe, the ventral sternopleurite
and meron similarly darkened; dorsopleural membrane broadly
yellow. Halteres with stetn yellorv, knobs broken. Legs rvith
the coxae yellow, sparsely pruinose, the bases of the fore pairvaguelydarkened; trochantersyellow; fernora obscureyellolv, thetips narrowly brownish black; tibiae yellowishbrown to brorvn,thetips narrowly more infuscated;tarsi brownishblack; claws (male)weakly toothed. Wings marbled,brorvnishyellow variegatedwith
cream-yellowand sparsedarker brown areas; prearcularand costal
fields a trifle more yellowed; the darkest markings include thestigrna,origin of Rs, anterior cord and a spot before midlength ofcellCu,,the last being precededand followed by more whitish markings; tips of cells R, to Il[", inclusive,with pale spots; cell lst A
with tlvo larger similar areas; irregular, more cream-yellowmarkings before and beyond stigma and in cells R and M at about thelevelof the origin of Rs,'veins brown. Venation: R1*z €ntire; Rs
about one-halflonger than rn-cu; petiole of cell M, and rz subequal.
Abdominal tergites yellow, vaguely trivittate with pale brown,
the central stripe broader and more diffuse, the narrow sublateral
stripes barely interrupted at the sutures; lateral tergal borders
narrowly light gray; sternitesobscureyellow, darkenedon sides;
outer segffrentsbrownish black. Male hypopygiurnrelatively large ;ninth tergite separatedfrom the sternite by " virtually entire suture ; basistyleand ninth sternite entirely fused. Ninth tergite
large, when viewed from aboveappearingnarrowed outwardly, theapex with a very shallowemargination; more ventrally with a smill
median lobe that is denselyprovided with blackenedspinules,thelobe subtendedon either side by a small pale blade. Ninth sternite
with a small lyriform medianappendage,the arms slender,glabrous.
Outer dististyle large, appearingas a flattenedgently curved blade,at base on either side with a pale membranouslobe that bears agroup of setae; outer end of style more expanded,on the concave
lo*ei surface with numerous setaeand a small lobule before theAprit,1e50 Bulletin of the Brooltlyn EntomologicalSociety

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obliquely truncated apex. Inner dististyle small, complex; beaksmall, slender,the correspondingposterior end of style a trifle
larger and stouter but having the sarne general shape. Eighthsterniteunarrned.
Habitat; Arizona (Pima County).
Holotypc: 3, Santa CatalinaN{ountains,June 26, 1935 (Owen
Bryant).
This interestingcrane-flysuperficiallyresemblesTipula (Bellardina) gothicanoAlexanderand someallied forrns,differingfrom allparticularly in the structure of the rnale hypopygium.
Tipula (Eumicrotipula) werneri n. sp.
Belongsto the gloplry,yoptera.
group; generalcolorationgray, thepraescutuurwith three darker brown stripes, their anterior ends.
with four polished black areas, the intermediatepair more extensive; antennalflagellum bicolored; femora obscureyellow, thetips blackened,precededby a clearer yellow ring ; wings whitish
subhyaline,
patternedwith dark and paler browni Rr+z ?trophied;abdcmenyellorv; ovipositor with the cerci slender,curved gently
downwardsto the tips.
Fernale;Length about 13 mm.; wing 13 mm.
Frontal prolongationof head polished yellow, narrowly linedwith brown on the sides; nasuslong and slender; basal two segments of palpi brown, the outer ones black. Antennae with thescapeand pedicel yellow, flagellum bicolored, the segmentsdark
brorvnor brownishblackat base,yellow apically,on the rnoreproximal segmentsthe amount of 1'ellor,vslightly exceedingthe black, theouter segrnentsmore uniformly darkened. Head above gray,vaguelypatternedwith a reddishline on either side of the posteriorvertex and with a capillary dark brown rnedian vitta; vertical
tuberclemore yellolved.
Pronotum gray, with three narro\,v brown markings. Mesonotum light gray, patternedwith brown; praescutumwith threedarker brown stripes,the anterior ends of the lateral pair and theanterolateral parts of the median vitta conspicuouslypolished
black, the latter areas larger ; median praescutalstripe narrowed
and darker at posterior end; scutum with a central darkening and
two isolated brown areas on each lobe; mediotergite with vague
indicationsof an incompletecapillary darkenedarea. Pleura andpleurotergitelight gray; dorsopleuralregion paleyellow. Haltereswith stem yellow, clearer at base,knob dark brown. Legs withthe coxae grayish pruinose; trochantersyellow; femora obscttre

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Tot.XLv

yellow, the tips blackened,precededby a clearer yellow ring ofabout the same width ; tibiae brownish yellow, the tips narrowly
brorvnishblack; tarsi passinginto black. Wings with the groundwhitish subhyaline,patterned with darker and lighter brown, theformer including much of the prearcular field and the vicinity ofarculus and h, with three further dark areas in cell Sc, the one atouter end smallest; interspacesof cell Sc paleyellow ; cell C medium
brown, variegatedwith pale yellow beyond lt, and again at the outer
end ; stigma brown ; paler brown washesover most of the remaining cells, more expandedand confluent near the wing tip, due tobroad marginal seall1sto the veins ; other slightly darker washes
acrossthe outer radial field and over the outer ends of cells R and
M ; still other pale washesin outer ends of the Anal cells; over .mostof the surface,the pale ground exceedsthe dark color in area; veinschiefly brown. Venation: Rr * 2 atrophiedi Rr nearly three times
Rr*r,' Rs more than twice nl-cut,,the latter oblique, placed on Ma
just beyondthe origin; cell Znd A relativelynarrow.
Abdomen yellow, the outer segmentssomewhat darker, possibly
causedby internal discoloration;caudal margins of segmentsvery
narrowly cleareryellow. Ovipositor with the cerci relatively slender, curved gently downward so that the concaveedgeis the ventral
one, instead of the reverse,as corllrnonin the genus.
H abitat: Arizona ( Pima County) .
Holotype: 9, Madera Canyon, Santa Rita Mountains, altitude5,600feet,August 14, 1949 (Floyd Werner and Williarn Nutting).
I take great pleasurein naming this distinct fly for Mr. Floyd
Werner, at this time a graduate student at Harvard University.
This is the first memberof the glapltyropteragroup of the subgenus
to be found in the United States. The very numerousNeotropical
speciesof the subgenusEu,nticrotipulaAlexander have been discussedby the writer in another paper ( Rev. de Enton'rologia, 17 :
172-201; 1946). There are severalspeciesin South America havittg vein Rr * 2 atrophied, as in the present fly, including Tipttla(Euncicrotipula)clticanoAlexander, T. (8.) co?rsonafo
Alexander,
T. (E.) efficar Alexander, T. (E.) flaztiduloAlexander, T. (E.)
fraudu,lentaAlexander, and others, all being quite distinct in thecoloration of the body, legs and wings, and in the structure of theovipositor. A conformationof the cerci such as occrlrsin the present fly is very uncommonin the entire family Tipulidae.
Wings fully developed (danmla) or greatly reduced (aestigipennis) when presentwith Sc unusually short, Sct ending far before the fork of Rs,' tips of outer radial veins gently upturned, R,

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GenusDactylolabis OstenSacken.
SubgenusEudactylolabis n. subgen.
more strongly so. iVlalehypopygium with two dististyles,the outert long pale blade; inner style unequally bifid, the lower arm apowerful blackenedclttb, its surfacewith abundant erect spinous
setae,the upper or inner arr11a pale lobe lying in the spacebetween
the two major styles. Ovipositor with the basal shield elongate,
dark brown; cerci very large, forming a dorsal sheathfor the hypovalvae,fused basally, the separatetips short and divergent; onlower outer rnargin of each cercus lvith a secondbroader acutetooth. In the elongatedconcavelorver surfaceof the cerci lie thepale,denselyhairy hypovalvae,their tips obtuse.
Type of subgenus: Doctl,lolobisdornwla(osten Sacken). The
new speciesherer,vithdescribedas Dactylolobis(Eudactilolabis)
ztestigiltennislikewise belongshere. The group seemsto be characteristicof the serniaridregionsof the southu'esternUnited Statesand it seernsprobablethat still further specieswill be discovered
here. Osten Sacken (LI/esternDiptero, Bull. U. S. Geol. Survey,
3 : 202; 1877) has describedthe very peculiar ovipositor, statingthat "it differs from that of any Tipulid I know of."Dactylolabis (Eudactylolabis) vestigipennis n. sp.
Generalcolorationlight gray; wings and halteresvery reduced;
male hypopygiumwith the lower arm of the inner dististyleunusually stout, its surfacewith abundantshort spinoussetae.
Male; Length about6 mm.; wing 0.85 mm.
The abovemeasureurents
are thoseof the paratypespecirnen; theholotype is larger but is not sufficientlywell preservedto add tothese measurements.
Rostrum gray pruinose; palpi brownish black. Antennae withthe scape and pedicel brorvn, sparsely pruinose; flagellum darkbrown, the segmentsoval, becorningmore elongate outwardly;
verticils short, subspinous,placed at near rnidlength of the segnrents. Head clear light gray.
Thoracic dorsum chiefly light gray, the praescuturnvaguely patterned with darker ; pleura clearer gray. Halteres very small,shorter than the wings, entirely pale. Legs with the elongatecoxae
pale brown, sparsely pruinose; trochanters obscure yellow; remainder of legs light brown, the outer tarsal segmentsa little
darker; tibial spurs distinct. Wings greatly reduced,as shqwn by
the measurements,
whitened,only the costalbolder a trifle darker
providedwith dark setae. Venation entirely atrophied.

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Abdorninal tergites conspicuouslypatterned, dark gray, with asubmedianpair of broken dark brown stripes, the median spaceclearer silvery gray, the dark stripes narrowly interrupted by thepale caudal borders of the segments; basal sternites more uniforrnly brown, the outer segmentsand hypopygittrnobscttrebrownish yellow. Male hypopygiurr with the tergal region thickened,
provided with abundant setae that are directed chiefly caudad.
Interbaseswith proximal ends stout, the long apical blades directed mesad. Outer dististyle a long narrow pale blade, setifercuson aboutthe proximal third. Inner dististl'lervith the lower
arm unusually stout, blackened,the tip obtuse, the entire outersurfacewith abundantshort spinoussetaethat are erect or retrorse,
thosenearerbaselonger and more slender;upper or axillary arm apale flattened blade, provided with scatteredsetae. Phallosome
complex,the terminal elernenta slenderacutespine.
Habitat: Arizona (Pima County).
Holotype: 3, Tucson Mountains, altitude 4,500 feet, February
2I, 1937 (Owen Bryant). Parotopotype:,{, altitude 4,000 feet,
Nlarch 4, 1937 (Owen Bryant).
I am indebtedto Mr. Bryant for further data concerningtheseflies and their habitat. The type was found on the under side ofa stick, the paratypebeneathstones. With the latter it was indiwere taken but one of thesewas apparently
catedthat two specirnens
lost or mislaid. Bryant stressesthe fact that the Tucson Mountains, lying to the west of Tucson, are entirely different from theSanta Catalinas, lying to the east. These tnountains are lower
and dryer and it is very difficult to find insect specimensthereon
except under stonesand the like. It is indicatedthat the range
probably has an endemicfauna that appearsonly in February andMarch, following the February rains. Bryant further states thatthe insect fauna hides beneathstonesand fragments of lvood during the day and is active at night. It is to be hoped that moremiterial of this unusuallyinterestingfly may be found in the future.
Rhabdomastix (Rhabdomastix) nuttingi n. sp.
Generalcolorationgray, the abdomen,including the hypopygium,
dark brown; antennaeof male approximatelyfour times the lengthof the body; wings with a weak grayish tinge, the stigma palebrou,n; veins virtually glabrous; Sc, ending approximatelyopposite
four-fifths the length of Rs.
Male: Length a-bout6 mm. ; wing 6.3 mm. ; antennaabout 24 mm.
Rostrum short,brown; palpi black. Antennae (male) very long,

Aprit,1e50 Bulletin of the Brooklyn EntomologicalSociety47

about four times the body; scapedark brown, greatly enlarged,
pedicelvery small; flageliurny.ilo*, the outer ig-.nt,
p"rJirrg
into brown, the segrnentsprogressivelylengthenedand rnoi. slen-der outwardly, beyondthe first glabrous. Head abovedark gray,with three oi"ttg. yellow spots,i medianone behind the antennalbases,the others adjoining the eyes at the narrowestpoint of thevertex; anterior vertex very broad.
Pronotum gray, the pretergitespale yellow. Mesonotum gray,the stripesnot or scarcelydifferentiated;pseudosutural
fovea. ""atuberculatepits black, the latter large. Pleura gray, the dorsopleural membranedusky. Halteres pale throughout. Legs rvith
the coxae testaceous,the fore pair darker; trochanterseLngate,yellow; remainderof legs generallylight brown, the femoral b"r.tmore yellowed,the tips of fernoraand tibiae narrowly and vaguelydarker ; outer tarsal segmentsinfuscated. Wings rvith a *"ulgrayishtinge,the prearcularand costalfieldsa trifle more yellowed;
9ligmaoval, palebrown; veinsbrown, more yellorvedat wing base.
Veinsbehindcostavirtually glabrous,lackingon R and its brinches,with the exceptionof a few unusuallysmall trichia at outer endsof
veinsRs and Mt*r. Venation: Sc relativelylong, Sc, endingabout
oppositefour-fifths the length of Rs, Sc, not far from its tip, Sc,
aloneabout one-halfr-nci R, suboblique,a little shorter thin the
distanceon costabetweenthe tips of veins Rr*, and R, ; nr, long,transverse; outer sectionof vein M t *2 not conspicuously
archedat
origin, as in many allied species; vein Znd A stronglysinuous,thecell broad.
Abdomen,includinghypopygium,dark brown.
Habitat.' Arizona (Santa Cruz County).
H olotype: 3, Bear valley, Tumacacori Mountains, at light,
August 22, 1949(Floyd Werner and William Nutting).
This interestingaddition to the United States fauna is namedfor Mr. William Nutting, to whom I expressmy appreciationfor
various Western North American Tipulidae. This is the firstrecord of any speciesof the typical subgenu
s RhabdomastirSkuse
from north of Mexico. The distributionof the relativelynumerous
speciesis chieflyantipodal,with a smallnumberof speciesoccurring
north of the equatorin both the New and the Old Worlds. The
most northerly American speciesto this date were various ones insouthern Mexico, including Rhabdornasti,r(Rhabdornastir) isobella Alexander, R. (R.) longiterebrataAlexander, and R. (R.)
ncericanoAlexander, all of which differ from the present species
and among themselvesin the details of venation and trichiation of
the wings and in featuresof sizeand coloration.