*'

5'f{}i

NE1V OR LITTLE-K]'{OWN ?IPULID.€ F"ROM EASTERNASIA {DTPTERA},iV

Bi- C}I,{-:a.rrs 1)- ,+-l.E:,{N}Ec
A i A:::l;.:;:t,:l.I,,:.ssuci.
t:stlts

Sepal*te fron
TIIE P}iII,tr?FiNEJOURN,dL OF' SCIEI{CE
Fublishr:ci bi' iire B'.ric:,r,1cf Scie::c': cf i.hc Pltiiiap::re Gc.,:ct::*rcnt
fI::.r:l;i. ir" i::-ul -jl:ir-i.'- l',':,

3. l'lit"or\:E::i,}IANItA
BLTREAU OF PRINTINg
13l s2,16.16?

1S23

NEW OR LITTLE-KNOWN TIPULID,IE FROM EASTERNA S IA (DIP T E RA ), I V 'By Cxlnr,os P. Ar,ux.a.Nnon
Of Amherst, Massachusetts
THREE

PLAfES

The crane flies discussed at this time are from various partsof eastern and southern Asia and were received from the following entomolgists and collectors: China, Messrs. A. P. Jacot'
E. Suenson, and T. Yokoyama; Japan, Messrs. Teiso Esaki, C.
Harukawa, S. Kuwayama, Jiro Machida, Akio Nohira, S. Sakaguchi, and E. Suenson; Formosa, Messrs. S. T. Issiki and K.
Takeuchi; Philippines, Messrs. Duyag, McGregor, and Rivera;
and India, Mr. C. F. C. Beeson and the authorities of the IndianMuseum. I wish to express my deepest thanks to all of theabove-named gentlemen for the opportunity of studying thegreat collections of crane flies upon which the following descriptions are based. Except where stated to the contrary, the typesof the novelties are preserved in my collection.
The crane-fly fauna of the Oriental Region is f,remendously
large and varied, more especially so on the slopes and summitsof the higher mountains where a characteristic Palearctic element is found. There can be little question that this region,
with its abundant isolated mountain ranges, major islands, andarchipelagoes,separated for past ages and having had an abundance of tirne for the formation of species,will be found to support
a greater number of species of Tipulide than any other regionof the World.
TIPULIN4
TIPULIN I
BRITHURASANCTA

6P. noY.

General coloration dark clove-brown, the prmscutum with threepaler reddish brown stripes; pleura light brown with two narrower dark brown longitudinal stripes; femora yellowish brown,
the tips narrowly blackened, preceded by a clearer yellow subAgri'Contribution from the Departmentof Entomology,Massachusetts
culturalCollege.
Bl7

318

The Phili,ppine Journol of Sciertce

rszs

terminal ring; wings with Sc, present; Rs subequal to R,, thelatter angulated and short-spurred near outer end; male hypo_pygium large, the sternal region only moderately produced.
Male.-Length, excluding head, about 26 millimeters; wing,
23 by 6. Head broken.
Pronoturn yellow, the posterior notum with two approximatedmedian tubercles that are dark brown, the anteiior notum
darkened laterally. Mesonotal prrescutum dark clove-brownwith three nearly confluent reddish browrr stripes, the dark
ground color restricted to the humeral and lateral regions; median stripe broad; scutum chiefly reddish brown; scutellum dark
reddish brown, more blackened medially, the parascutella black;
postnotal mediotergite dark brown, the posterior portion
withan obscureyellow triangle. pleura light brown with two narrowdarker brown longitudinal stripes, the more dorsal lying justbeneath the dorsopleural membrane, including the propieura;
ventral stripe including the fore coxe, continued across thesternopleurite to the metapreura; preurotergite erevated, theanterior face with an appressed yellow pubescence. Haltereslight reddish brown, the base of each knob darker. Legs withthe coxa dark brown; trochanters paler, especially the fore
trochanters; femora yelrowish brown, the base and a subapicar
ring clearer }'ellon', the tip narrowry brackened; tibia brown,
the base yellow; tarsi brown. Wings (plate 1, fig. 1) as in thegenus; costal region opposite the stigma dilated but notbroken;ground color yellow, with a sparse dark pattern, incruding
thebases of cells R and M; origin of Rs; a large area extending
from the stigrna across the cord; outer ends of all outer cells
from R, to 2d A inclusive infumed, most of them inclosing small
marginal yellow areas; medial veins beyond the cord narrowly
bordered by yellow, which in turn has a dusky margin,
darker than the centers of the cells; a conspicuous darkened
area on either side of vein lst A at near midlength; veins yellow,
the costal vein more reddish brown. venation: sc, present; Rs
strongly arcuated, subequal to R'the latter angulated and shortspurred just before apex; cell Lst M, higher than long.
Abdominal tergites brownish black ; hypopygiurn conspicuousry
reddish; sternites reddish brown. Male hypopygium -(pratez,fig. 19) relatively large but ninth sternite, 9s, not g*""iry d"rr"loped, as is the case in B. crassa Edwards. Ninih te"gite gt,
narrowed outrvardly, the caudal margin with a deep u-shaped
emargination that is fringed rvith conspicuouserect }-ellorr sete,
the lateral lobes thus formed relativell. conspicuous. Basistl-le,

4o,sAlenand,er: Ti,pulid'n from Eastent Asia, IV

319

b, produced, the apex flattened, provided with long yellow set're'Dististyle, d, shiny reddish brown, very large and complex instructure. Ninth sternite, 9s, produced into a median tubercleof moderate size.
Habitut.-China (Chi li).
Holotype, male, Ton Che ssu, a ternple in hills west of Peking,August 27, t92]- (A. P. Jacot) .
Bri,thura sancta differs from B. nymphi,cu Alexander, which
seemsto be its nearest relative, in the details of coloration andvenation, especially the retention of Sc' and the arrangementof veins in the radial field. The venation of B. coni,frons Edwards, the genotype, is shown for comparison (Plate L, fig. 2).
TIPULASUBFUTILIS

3p. noY'

Allied to T. futi'Iis Alexander; antenne with the basal segments of the flagellum bicolorous; wings with a complete whitecrossband beyond the cord; distal end of Rr*, atrophied; malehypopygium with the lateral lobes of the ninth tergite very low
and obtuse, the caudal margin notched, with two small submedian ridges; gonapophyses blackened, simply bispinous.
Male.-Length, about 13 millimeters; wing, 16 by 3.7.
Dbscribed from an alcoholic specimen.
Frontal prolongation of head relatively long, brownish yellow;
palpi pale. Antennee of moderate length, if bent backward extending to ,shortly beyond the base of the abdomen; basal segment yellow; second scapal segment brown; first flagellar segment yellow; succeedingsegpents brown, the distal end yellow,
the amount of the latter decreasing on the outer segments, on
the second flagellar including only the distal fourth; beyond thefifth flagellar, the segments are entirely brown; terminal flagellar
segrnent very small, oval. Head brownish gray; vertical tubercle low and obtuse, entire.
Mesonotal prascutum pale brown, with three paler' morebrownish yellow stripes that are ill-defined against this background; scutellum and postnotum yellowish brown, with a delicate brown median vitta. Pleura uniformly brownish yellow'
possibly pruinose in dry specimens. Halteres chiefly pale. Legs
with the femora brownish yellow, clearer basally, darker at tips;
tibie and tarsi passing into dark brown' Wings relatively longand narrow, as shown by the measurements; pattern almost as
in T. futi,tis, the ground color brownish, variegated with whitish
subhyaline, this latter including a complete crossband beyond
the cord; smaller pale areas in cell R, before the stigma; nearouter end of cell M; at outer end of cells Cu and lst A, crossing

320

The Phikppine Journal of Sci,ence

1e2g

the latter vein; and at midlength of cell Lst A, not touching theveins; darker areas than the ground color at origin of Rs. anterior cord and as a conspicuousseam along the distal half of Cur.
Venation, Rr+, pale at base, scarcely visible, the outer end
entirely atrophied; R, short, perpendicular.
Abdominal tergites obscure yellow with a narrow but conspicuous median brown line that is nearly continuous; sternites
more uniformly yellow. Male hypopygium (plate 2, fig. Z0)
with the ninth tergite (Plate 2, fig. 2L) extensive, gradually
narrowed outwardly, the caudal margin with a very shallow
emargination, the lateral lobes thus formed very low and obruse;a very small rounded tooth on either side of the median line,
these produced back onto the dorsal surface of the tergite asshort ridges that inclose a narrow linear furrow. Outer dististyle widely expanded outrvardly. Gonapophyss, g, appearing
as heavily blackened structures that lie on either side of theredeagus, each apophysis bispinous, the more dorsal spine alittle longer.
H abitat.-Japan ( Kiushiu ) .
Holotype, alcoholic male, Unzen Park, Hizen, May, Ig26 (8.
Suenson), received through Doctor Crampton.
Tipu,l,afutili.s Alexander (Japan) differs from the present species in the details of structure of the male hypopygium. In T.
futilis, the caudal margin of the ninth tergite (plate Z, fig. 22)
is more deeply and evidently notched, the margin heavily blackened, the ]ateral lobes more conspicuousi the base of the notchbears a small black median tooth, in addition to two smaller
and blunter sublateral knobs; dorsal surface not evidenfly ridged
and furrowed as in subfutili,s. Outer dististyle (plate 2, fig.
23) quite as in T. wbfutilis.
Inner dististyle, fd, with theventral margin produced into a stout lobe that ends in a nearlyterminal black spine. Gonapophyses,gf, with the dorsal arm
bifid, the ventral arm longer and slenderer than in T. subfuti,ti,s.
TIPULA

I-USOUOIDES

sp. nov.

Male.-Length, about 14 millimeters; wing, 1?.
Female.-Length, about 20 millimeters; wing, 19.
Described from alcoholic specimens.
Closely allied and generally similar lo T. yusoar Alexander(Japan), differing chiefly in the structure of the male hypopyglum.
Male hypopygium (Plate 2, fig. 24) wit]n the caudal marginof the ninth tergite arched and heavily blackened, as in ?. Auaou,

4o,s

I

Alexand,er: Tipul,idp from Eastetn Asia, IV

321

the dentation of the margin quite different, the most ventral
tooth on either side being much larger and produced into anacute spine. Outer dististyle smaller, more dilated before apex.
Inner dististyle, id, with the blade much smaller, the bilobed apex
heavily blackenedand relatively small. Eighth sternite, 8s, withthe caudal lobe longer than broad, clothed rvith abundant long
golden sete.
H abi,tat.-Japan (Kiushiu) .
Holotype, male, Unzen Park, Hizen, May, L926 (8. Suenson).
Allotopotype, female.
In Tipula Ulsou Alexander' the caudal margin of the tergite
is highly arched, narrowly blackened, with numerous small teeth
and lobules, none of which is acute. Outer dististyle elongate'
gradually widened outwar'dly. Inner dististyle large, the blades
conspicuous. Eighth sternite with the caudal lobe transverse,
the sete chiefly marginal.
TIPULASUBYUSOU sp. noY.

Male.-I'ength, about 16 millimeters; wing, 18.
Closely allied to T. gusou Alexander (Japan), differing especially in the structure of the male hypopygium.
Ninth tergite with the caudal margin narrowly darkened or
blackened but without distinct denticles. Outer dististyle nearly
as in 7. ynsouoides, being rather conspicuously dilated just beyond midlength. Inner dististyle with the outer marginal lobe
intensely blackened, obtusely rounded, the caudolateral margin
bearing a pale lateral flange. Eighth sternite with the caudal
lobe transverse, its margin with a dense brush of long golden
sete. Gonapophyses (Plate 2, fig.25) trifid, as in 7. gttsou,bnt
the arms much longer and slenderer, arising from a long, heavily
blackened base, the arms gently curved and slightly decurved.
In T, yusou (Plate 2, fig.26), the gonapophysesare paler andnearly straight, the loneler median spine subtended by the lateralspines.
H abi,tat.-Japan ( Honshiu ) .
Holotype, male, Mount Takao, altitude 1,000 to 2,000 feet,
May 7,]^922 (Tei,so Esaki). Paratype, male, Nakano, Tokyo,
May 1, t922 (Teiso Esaki).
TIPULARANTAICOLA

sp. nov.

General coloration ocherous yellow, the prescutum with illdefined darker stripes ; antenne bicolorous; wings grayish brown,'Canadian Entomolog:ist 46 (1914) 240-241, pl. 19, fig. 1.

322

The Phi,kppine Jownal of Sci,ence

7s2s

certain of the veins seamedwith darker; cell lst M, very small;male hypopygium with the median region of the tergite produced
into a decurved setiferous lobe.
Male.-Length, about 9 to 10 millimeters; wing, 9 to 10.
Female.-Length, about 11 millimeters; wing, 11.
Frontal prolongation of head brownish yellow; palpi darkbrown. Antennre with the scape yellow; flagellum weakly tostrongly bicolorous, the basal enlargement of each segment
brownish black, the remainder brownish yellow to yellow; antennre (male) relatively elongate, if bent backward extending
to some distance beyond base of abdomen; antennrc (female)
shorter, distinctly bicolorous. Head brownish gray, clearer grayin front; a narrow dusky median line on anterior vertex.
Mesonotal prascutum ocherous yellow, the three usual stripes
only a trifle darker, the median stripe split by a capillary, slightly
darker vitta; scutal lobes extensively darkened; scutellum andpostnotum brown. Pleura brorvn. Halteres yellow, the knobs
infuscated, their apices pale to distinctly yellow. LeSs lrith thecoxe and troehanters yellow; femora yellow, passing into darkbrown; tibie and tarsi dark brown. Wings (Plate 1, fig. 3)
with a grayish brown suffusion, the base and costal region moreyellowish; cell C a trifle infumed; stigma conspicuous, darkbrown ; outer end of cell R, darkened; conspicuousbrown seams
on Cu, m-cu, and more naryowly on most other longitudinal veins;
extensive whitish areas before and beyond the stigma and acrosscell 1st M,; veins dark brown, the obliterative areas whitish, theveins in the yellowish areas more flavous. Macrotrichia of veinsvery long and abundant. Venation: Cell lst M, very small,diamond-shaped to pentagonal.
Abdominal tergites obscure yellow, the lateral and caudal
margins of the segments blackened, the amount increasing on theouter segments; sternites yellow, the caudal margins of the segments a little darkened; eighth sternite blackened; hypopygium
chiefly pale. Male hypopygrum small, the median region of thetergite produced into a conspicuous decurved lobe, the sides ofwhich are densely clothed with conspicuousyellow setre. Innerdististyle large, jutting caudad as large, compressedto slightly
tumid lobes, the apex of each with a small group of black spinous sete, this lobe much larger than the flattened outer dististyle. Eighth sternite unarmed.
Habi,tat.-Formosa.
Holotype, male, Rantaizan, altitude 7,000 feet, J:une 2, 1927
(S.
Issiki). Allotopotype, female. Paratopotypes, 4 males.
".

Alenand,er:Ti,puli,'dpfrom Eastern Asia, IV

40,3

323

TittuLa rantaioola bears a marked resemblance to ?. microcellula Alexander (Plate 1, fig. 4) in the general appearanceandvery small cell 1st M,, differing especially in the details of coloration, wing venation, and the structure of the male hypopygium,
especially the ninth tergite which here bears a setiferous medianlobe.
NESOPEZAEANTAIZANA

sP. nov.

General coloration ocherous, the mesonotal prescutum with a
median darker line; antenna (male) relatively long; legs brown,
the tibiee and tarsi extensively white; wings nearly hyaline, thestigma brown; free tip of Sc, nearly atrophied, indicated only
by a pale line; basal section of Rn*u and r-m pale, without
macrotrichia, in transverse alignment; forks of medial field
relatively shallow, male hypopygium with the tergite produced
laterad into slender blackened rods, the median area of thecaudal region further produced into a decurved point.
Male.-Length, about 8 millimeters; rving, 10.
Rostrum testaceousyellorn'; palpi pale brown. Antenne withthe scapal segments yellorn',the flagellum dark brown; antennm
l2-segmented, the terminal segment small; flagellar segmentswith dense short seta and longer unilaterally arranged black
verticils that are conspicuousbut scarcely one-half the length of
the segment. Head ocherous yellow.
Thorax almost uniformly ocherous,the prrescutum with a median darker line; pleura more testaceous. Halteres very longand slender, pale brown, the knobs darker. Legs with the coxre
and trochanters yellowish testaceous; femora brown; posterior
tibire dusky, the extreme tips white; tarsi dusky, the tips paling
to rvhite; on the other legs the amount of rvhite on the tibim
and tarsi appears to be mot'e extensive' \Yings (Plate 1' fig. 5)
relatively broad, nearll' h1'aline,the oval stigma brown; veinsbrown, unusually slender, rvith long conspicrrousmacrotrichia.
Macrotrichia on the distal third of vein ilI, distal half of basal
section of Cu,, and outer ends of both anal veins. Venation:sc, ending opposite four-fifths the length of Rs; free tip of
6c, atrophied or barely visible; basal section of Rn*u and r-m
pale, without macrotrichia, in transverse alignment; forks of
medial veins relatively shallo\\'; m-cu subtransverse; cell 2d A
narrow.
Abdomen yellowish brown, darker laterally. Male hypopygium with the lateral margins of the tergal region produced
caudad and slightly mesad into slender blackened arms, their
246466-3

The phitippine Journal of
Science324

rszs
tips microscopicalry spicurose;
median reg:ion of tergite withabroad-based acutepoint
itrat is directed ventrad, the.medianmargin of the tersite
dorsad of this_with a fringeof long erectsete; tergite pare except ror
ttre nuckened ,aterar andcaudarmargins. outer dististyre relativeiy
short and--ilr.#.pi.uou*,cylindrical, the apex truncate.
lnner dististyle deeply bifid.
Habitat._Formosa.
Holotype,slighily

Rantaizan, altitude, 2,000-teneral,feet,Junez, tg27
-ufu,.(,S.i. tisttci\.--..''NDSOpDZAIDIOPEALLUS

aD. nov.

Generalcolorationdark brown;
antenna with the scapalseg_
ments ye'ow, the flagetum oartrrrown; front and anterior vertex conspicuously
yeltow; mesonotum.r,i.nr
Ju"t;";;;;
medianarea of the prescu}ls,culumnaferl pleura pale brownish
yellow, the anepisternum
IrU
and sternopleuriteconspicuously
dark

s3gwq
white;
mare
ffix'J#iffi*fi'p*";ir'"L"'i'"t"'.r*rr
subtendedon eithersideby a flattenedtHfiffi-"h:onspicuous,(ale.7Length,aboutro .liii**ers; wing,11.b.

Female.-r*ngth, about l5.;luil;ers;
yi1g, 19.
Frontal protonsationor hlJ;;;,
entirely without nasus;
palpi dark brown. .l,ntenne
1_"f"1ofmoderate length, if
bent backward exrendingaboui-;;'th"
bu."
;i;;;
uiuo_"r,scaparsegmentspare.yetow,
tne nagertumdark browni flags11sr
segmentselongate-cylindrical,
the ba'salenf*rg"*."t* oi ifru ."*_
ments inconspicuous,the*.attered, shorter than thesegments. Front and anterior"""ti"it,vertexr n,qre
pale yellow;
Jsrruw; rem
remainderof head dark brown\
Pronotumbrown, the posterior
notum more yellowish. Meso_
notat prescutum dark brown,
trre muaiun
paler;
scutumdark brown, the media;"d";;;;;;t;.;r; paler;posterior sclerites
of mesonotumtestaceousbrown.
irf"":t pale brownirf,
with a broadtransversedark brownv"ffo*,
ti"ar" that incrudestiu ane_pisternum and sternopleuritet.i-ilarly darkened. Hal_
teres pale,the knobsdark brown.
-uronkg. with the coxa yellow,
little darkenedat base;trochrnt""*llitow;afemora arlf.Lo*rr,
their basesparer; tibizeand tu";i;u;'brown, the distar fifthofthe basitarsi and remaining tu.r"isegments snowy white.
Wings with a strong brownisil .ufuriorr,-on' the oval stigma darker
brown; ilr-delimited dark
unterior .o;J-;;
io_.u,
whitish obliterative areas "roua*
before and.O"_-r; irr" .rrJ*j.raacrossthe fork of M. Venation: Rso, moderatelength, about

The Phili,ppine Jourrwl of Sci,ence

326

rczs

a little shorter than the segments; antenne of moderate length,
if bent backward extending about to the root of the halteres.
Head light brown.
Mesonotal prrescutum dark brown, subnitidous, with threebrownish black stripes; scutum, scutellum, and postnotum uniformly dark brown. Pleura dark liver brown, the dorsopleural
membrane pale. Halteres pale, the knobs brown. Legs withthe coxa concolorouswith the pleura; trochanters yellow; femora
dark brown, the tips broadly more yellowish; tibia and tarsi
brown. Wings (Plate 1, fig. 6) with a pale brown suffusion,
the base and costal region a little more saturated; stigma oval,
dark brown; conspicuous pale yellow antestigmal and poststigmal areas,the latter including more than the basal half of cellSc,; veins dark. Venation: Rs transverse,opposite Scr; R, andR, in nearly transverse alignmenti R,*o entirely atrophied.
Abdomen with the basal tergites obscuie brownish yellow, beyond the second segment reddish brown, gradually deepening toblack; sternites beyond the basal segment obscure yellow withnarrow black cross lines near base and at margins; outer sternites uniformly darkened; male hypopygium with the inner dististyle, id, yellowish, the remainder chiefly dark. Male hypopygium (Plate 2, fi,g. 30) u'ith the caudal margin of the ninth
tergite, 9t, nearly transverse, the median area blackened,chitinized, produced into two slender, needlelike points, one oneither side of the median line (Plate 2, flg. 31).
(Honshiu) .
Habitat.-Japan
Holotype, male, Lake Ozenuma, on boundary between Iwashirono-kuni and Kotsuke-no-kuni, altitude 5,410 feet, July 25,lg2}(Tei.soEsaki).
OROPEZASAUTERI

Riedel-Oropezasauteri,RIEonL,Archiv ftir Naturgeschichte82 (7917')IL4\
115.
Riedel's types, both sexes, were from Okaseki, Taihoku District, Formosa, collected in June, I9I4, by Sauter. I have seen
additional males and females from Taihoku, taken Apnl20, 7g22,
by K. Takeuchi.
These latter specimens agree very closely with Riedel's description except that the incisures of the abdomen are not darkened but the narrox' transverse dark marking of each segmentlies at or before midlength of the segment, the base and apexbeing broadly yellowish. The sternities are extensively brownat the base and very narrowly so at the apex, l'ith a conspicuous40 , 3

Alerand.er: Tiputid,a from Eastern Asia, IV

327

yellowish ring before the apex; subterminal segpents more uniyellow'
iormty darkened; hypopygium small, chiefly obscure
Malehypopygiumwiththemediannotchofthetergitesmall,
U-shaped with no distinct notches or incisions laterad of this;ventrolateral arms of tergite terminating in large oval heads.
LIMONIIN,lE
LIM ONIINI
LIMONIA(LIMONIA)RANTAIENSIS

aD' nov'

Mesothoraxshinyblack;halteresyellow;legsblack,thebroad
femoral bases,a subterminal femoral ring, and the narrow basesof the tibia yellow; wings yellowish, the veins seamed withbrown; Sc shorti Rr+, elongate.
Female.-Length, about 9 millimeters; rving, 9'3'
Rostrum and palpi black. Antenne rvith the scape black, thebasal flagellar segments black rvith their tips somervhat morebrownish; flagellar segments oval, the verticils considerably
longer than the segments. Head brorrnish black, not nitidous;
anterior vertex a little wider than the first scapal segment.
pronotum black, more pruinose laterally. Mesonotum black,nitidous, only the dorsopleural region restrictedly paler' Pleura
black. Halteres yellow, the knobs orange-yellow. Legs with thefore coxa black; middle coxa darkened basally; hind coxa andall trochanters yellow; femora with more than the basal halfyellow, the remainder black, inclosing a conspicuousyellow ring
about its own length before the apex; tibire black, the extreme
base yellow; tarsi black. Wings (Plate 1, fig. 7) yellow, the baseand costal region somewhat brighter; an extensive brown pattern
distributed as follows: Broad seams along Rs, Rz+s, and Ra,
confluent with the small and ill-defined stigrna; cord, outer endof cells lst M,, and longitudinal veins beyond cord seamed withbrown; broad seams on Cu', interrupted at base and before
end of basal section of vein; broad seams at ends of, both analveins and in the axilla; veins brown; yellow in the basal andcostal regions. venation: h indistinct; sc relatively short, sc,extending to about opposite one-fourth the length of the strongly
arcuated Rs, Sc, close to its tip; R, only about one-third to onefourth of Rr*r; cell 1st M, closed; m-cu shortly before fork of M'
Abrtominai'tergites brownish black, the segments variegatedmedially and caudally with yellow, the ground color more or lessrestricted to large basolateral areas; genital segment and ovipositor pale reddish yellow; sternites obscure yellow; base of

328

The Phi,kppine Journal of Sci,encerezesternal valves of ovipositor blackened. Ovipositor with thetergal valves unusually slender, acicular, nearly straight.
Habttat.-Formosa.
Holotype, female, Rantaizan, altitude 6,000 feet, June 3, L927
(S.
Issiki.).
".
Limoniarantaiensis is well distinguished from allied regional
speciesby the shining black mesothorax, short Rs, long Rr*r, andthe wing and leg pattern.
LIMONIA(DICRANOMYIA)

TRISTOIDES

sp. nov.

Allied to L. (D.) tristis Schummel; general coloration gray, thepr€scutum with three brown stripes; legs chiefly brownish black,the femoral bases yellow; wings subhyaline, the stigma reducedor lacking; veins pale brown; male hypopygium with the spinesof the rostral prolongation of the ventral dististyle united for ashort distance at base, arising from long basal tubercles.
Male.-Length, about 5.2 millimeters; wing, 6.1.
Rostrum and palpi dark brown. Antenne dark brown, thesecond,segmentmore reddish apically; flagellum black; flagellar
segments oval, the outer segments more elongate. Head lightgray.
Ilesonotal prascutum 1'ellos'ishgral'rvith three brorrn stripes,
the median stripe longer and broader; scuteilum brown, paler
behind ; postnotum gral-. Pleura gra), vaguely marked withbrorrn. Halteres long, entirely pale. Legs with the coxe reddish brorvn; trochanters yellow; femora yellow at base,passing
lnto black, most extensively blackened on fore femora, only theapices of middle and hind femora blackened; tibie and tarsi
brownish black; legs long and relatively stout. Wings (Plate
1, flg. 8) subhyaline, the stigmal area greaUy reduced or lacking;veins pale brown. Venation: Sc, ending opposite origin of Rs,
the latter relatively long, about two and one-half times the basal
section of Rrlu ; cells lst M, closed; m-cu oblique, at fork of M,
shorter than the distal section of Cu,.
Abdomen dark brown, the hypopygium chiefly obscure yellow.
Male hypopygium (Plate 2, fig.32) with the tergite, 9f, transverse, the caudal edge narro*'ly margined with chitin and provided with sete, including a small median group; caudal marginof tergite gently emarginate. Basistyle, b, relatively long, witha single fingerlike lobe shorty beyond midlength, this tuftedwith setre; mesal apical angle of basistyle with about four setigerous punctures but not produced into a. tubercle; ventromesal
lobe of basistyle large, setiferous. Dorsal dististyle. dd, much

4o,sAlerand,er: Tipulidp from Ea,stern Asia, IV

329

stouter than in L. subtristis, curved to the acute apex' Ventral
dististyle, i'd. unusualll' lorv, pale, produced into the blackened
rostral prolongation. the latter bearing two long blaek spines atnear midlength. the-.s arising from a very short common base,both from etrongateba-ral tubercles: inner spine a little shorter
than the ouiter. Gonapophl'ses.g. very broad, the mesal apicalangtreproduced into a short blackened hook.
H a'liitat .-]Ianchuria.
Holo4ae, male, Koshurei, Kongchuling, altitude 625 feet, July25, 1924 (T. Yokoyama).
Limonia trtstoi.des is most closely alied to L. (D.) subtristis
Alexander (northern Japan), differing especiallyin the details of
structure of the male hypopygium, as the transverse emarginate
ninth tergite, the short stout dorsal dististyle, and the low oval
ventral dististyle with the long black rostral spines quite distinct in their structure and origin.
LIDIONIA(GERANOMYIA)TENUISPINOSA

sp. nov.

General coloration of mesonotum reddish brown, the prascutum with three narrow brownish black stripes; pleura yellow
with a broad dark brown dorsal stripe; legs yellow; wings grayish yellow with a heary, chiefly costal pattern; Sc long; m-cu
far before the fork of M; abdominal tergites dark brown; malehypopygium with the spines of the rostral prolongation very
long and slender, the outermost arising frorn a large swollen
tubercle that is subequal in size to the prolongation beyond it.
Male.-Length, excluding rostrum, about 6 millimeters; wing,
7; rostrum, ahoat 2.5.
Female.-Length, excluding rostrum, about 7.5 millimeters;
wing:, 7.5; rostrum, about 3.
Rostrum black; maxillary palpi black, 2-segxnented. Antenneewith the basal segment black; second segment dark brown;
flagellum obscure yellow, the outer segments darkened; flagellar
segments short-oval, the outer segments more elongate. Headdusky gray, the narro\ry anterior t'ertex clearer gray.
Mesonotal prescutum reddish brown with three conspicuous
brownish black stripes, the median stripe narrowed to a point at
the suture; humeral and lateral regions more yellowish; scutal
lobes extensively dark brown; median region of scutum and thescutellum reddish, slightly pruinose, with a continuous capillarybrown vitta; postnotum dark plumbeous. Pleura yellow, thedorsal pleurites and membrane occupied by a broad dark brown
longitudinal stripe that extends from the pronotum to the ab.

330

The Phikppine Journal of Sciencers2s

domen, including the preurotergite. Hartereschiefly pare, thekn-obs weakly darkened. Legs wittr
the coxe and trochanters
yellow; remainder of legs yetow,
onry the outer tarsal segments
dark brown. Wings grayish yet'tow, witha dark b;";;; --it chiefly
costal pattern; a small, scarcely evident
Ou"f. *ul_t ,similar area beyond this in cet sc; a conspicuous^
trianguLr areasurrounding the subcostalcrossvein; extensive
areas at- origin ofRs and tip of sc, these latter confluent
or nearry so, especiaty incell C; stigmal area very large, long_rectangular;a ,arge area
at outer end of vein R, and a small
circular cloud at eno ofvein Rn*u; narrow and only slightly conspicuous
seams at forkof Rs, anterior cord, m-cu, und oui""of end ce' fst Vtr; veirrs
pale, darker in the infuscated areas.
venation: A sup-6"nume_rary crossvein in cell Sc at near midlength;
Sc long, S.] *t."a_ing to about two-thirds the length ofRs,'S", ui-iL-lio,__.o- more than one-half its length before thefork of ffn.
Abdominar tergites uniformry dark brown,
the sternites obsc-urebrownish yetow, the caudar margins
or ine."r*"rl*somewhat paler, the lateral margins nu""orily
aartenJihypoivgiurndark. Male hypopygium
lFtate B, fig. BB) with the ,,initie"gitetransverse, the caudar margin with a sma,u-shaped emargination; seta mosfly marginal but continued
to the cephalolaterar
portions of the tergite. Basistvre,
b, relativery ;;-ii-d;;entro_
mesal lobe small. Ventral dististyie,ud,, large, exceeding thebasistyle in size; rostral prolongation slenJer,
U.u"i"s t*o'fleshyspines, the outermost arising froma very largs
iuU"""f"that is subequar to the apei of the rostrar prorongation
itserf,the spine very long and slender; the second
spine is close to
the base of the rostrum, about'two-thirds
tne re"gtrr-ot tt"
outer spine, very srender and genly curved.
Dorsaidististyre,
dd, a strongly curved sickre, the extreme
tip acute and upturned.
$gnanophyses, gr, pale, the lateral margjn of the mesal apicallobe with microscopic roughenings. .CI.*gu,
ilr;t;;;;trictea
at near midlength.
Habitat.-China(Che.kiang).
Holotype, male, hills south oJ Ning_po, May
1, j.gZE (8. Suen_son_).. Allotopotype, female. pa"aiopotype,male.
(G.) aaocetta r(Alexlimontn tinuispinosa resembles i.
ander) and allied forms, being most readity
separated ry tnu
somewhat remarkable structure of themale hypopygiu*. 'LIMONIA(GERANOMYIA)SUENSONIANA

sp. nov.

General coloration of mesonotum obscure yellow,
the prre_scutum with three brown stripes; pleura yellow,
with a natrrow40,3

Alerand,er: Tipulid'a from Eastern Asia, IV

331

and relatively ill-defined dorsal brown stripe; legs yellow, theterminal tarsal segments darkened; wings subhyaline, with ahearf' costal brown pattern, the areas at origin of Rs and end of
Sc widely separate; Sc long, Sc' ending nearly opposite thefork of Rs; m-cu far before the fork of M; abdominal tergites
bicolorous, their apices broadly darkened; male hypopygiumwith the rostral prolongation of the ventral dististyle short'the two spines close together, relatively short.
Mole.-Length, excluding rostrum, about 6 millimeters; wing,
6; rostrum, about 2.
Rostrum and palpi black, the former slightly paler at tips.
Antennae dark brown; flagellar segments oval to long-oval.
Head gray.
Mesonotal prascutum obscure yellow with three brown stripes,
the median stripe considerably narrowed behind; lateral stripes
narrow, crossing the suture onto the scutal lobes; humeral andlateral regions of prescutum extensively light yellow; scutellum
yellow with a brown spot medially at base; postnotum pruinose.
Pleura yellow with a relatively narrow and ill-delimited dorsal
brown stripe extending from the propleura to the postnotum,
including the dorsal pleurotergite. Halteres pale, the knobs anda narrow subbasal ring on stem dark brow:r. Legs with thecoxre and trochanters yellow; remainder of legs obscure yellow,
the terminal two tarsal segments dark brown. Wings (Plate 1,
fig. 9) subhyaline, with a fairly heavy, chiefly costal brownpattern that is arranged much as in tenuispinosa sp. nov.; theareas at Rs and end of Sc are widely separated; vring tip more
extensively darkened, inclosing a pale circular spot in outer endof cell Ru; cord and outer end of cell lst M, interruptedly andnarrowly seamed with brown. Venation: Sc very long, Sc'
ending shortly before the fork of Rs, Sc, not far from its tip;
m-cu nearly its own length before the fork of M.
Abdominal tergites bicolorous, the bases of the segments
obscure yellow, the caudal margins broadly infuscated; subterminal segmentsmore uniformly darkened; hypopygium chiefly
dark; sternites pale yellow. Male hypopygium with the ninth
tergite tralrsverse, the caudal margin notched, the seta chiefly
confined to the lobes, not reaching the cephalic lateral portionsof the tergite. Basistyle small, the ventromesal lobe low. Ventral dististyle very large and fleshy, the rostral prolongation
short and blunt, bearing two spines that are placed close together
at near midlmgth of the prolongation, the outer spine a little
longer than the inner, both of moderate length, acute at tips.

332

The Pldlippine Journal of Sci,ence

Ls2o

Dorsal dististyle a strongly curved sickle, the tip suddenly narrowed into an acute point. Gonapophyses with the mesal apicallobe long and slender, the tips narrowed into slender acutepoints.
Habitat.-China (Che-kiang).
Ifolotype, male, hills south of Ning-po, May 1, t925 (8. Suenson). Paratopotype, male.
I take gfeat pleasure in naming this interesting erane fly
in honor of the collector, Mr. E. Suenson,who has collected numerous Tipulide in China and Japan. It is allied to L. (G.)
semi.strtata (Brunetti) and similar species of the Oriental
fauna, differing especially in the venation and details of the malehypopygium. The acutely pointed gonapophyses present anunusual character in this involved genus.
LIMONIA(ALEXANDRIARIA)

ARGYRATA

sp. noy.

Head black, the broad front and posterior orbits silvery;
mesonotal prescutum shiny castaneous,the lateral regions yellow; scutal lobes blackened; wings hyaline or nearly so, thestigma a little darker; Sc short, Sc, and 2d A long.
Female.-Length, about 5.5 millimeters; wing, 5.4 to 5.5.
Rostrum reddish brown, the basal segment of the maxillary
palpus pale, the outer segments black. Antennre black throughout; flagellar segments oval. Head black, the broad front andwide posterior orbits silvery.
Pronotum shinl' I'ellorv. Mesonotal prrescutum shiny castaneous, the outer ends of the usual lateral stripes darker; lateraland humeral regions clear 1-ellor-; scutal lobes conspicuously
blackened, the median region yellorv; scutellum testaceous yellow; posnotal mediotergite dark plumbeous medially, margind
with paler. Pleura yellow, the sternopleurite and anepisternum
more purplish. Halteres short, pale, the knobs a little infuscated. Legs with the coxre reddish yellow; trochanters obscureyellow; remainder of legs obscure yellow, the outer tarsal segments passing into black. Wings (Plate 1, fig. 10) hyaline or
nearly so; stigma pale, scarcely darker than the ground color;veins brown. Venation: Sc short, Sc, ending far before theorigin of Rs; Sc, long, about one-third longer than m-cu; Rs
short, arcuated, about one-half the basal section of R r_u; free
tip of Sc" before the level of R,; m-cu just before the fork of
M; vein 2d A long, the cell wide.
Abdominal tergites dark brown, the sternites obscure yellow,
the basal segments a little darker; genital segment yellow.
Ovipositor wtith the terg:al valves slender, gently upcurrred.

r,3 :

Alerartd,er: Ti,pulida from Eastern Asia, IV

333

Habttat.-Formosa.
Holoty'pe, fernale, Urai, altitude about 1,500 feet, April 20,
L922 (K. Takeuchi). Paratopotype, female. Paratype, female,
Hokuto, 1912 (H. Sauter); Deutschen Entomologischen Institute, through Dr. Walther Horn.
LIMONIA(TERYPTICOMYIA)

BREVICUSPIS

sp. nov.

Size large (wing, male,7 rnillimeters or more) ; general coloration dark brown, the pleura and sternum paler; wings whitish
subhyaline; stigma small, dark brown; free tip of Sc, not far
before R, R, shortened; apical spur of Rr*, elongate; malc
hypopygium with the spines of the rostral prolongation of theventral dististyle short and spikelike, placed close together atbase of the prolongation.
Male.-Length, about 7 to 7.5 millimeters; wing, 7 to 5.
Rostrum obscure yellow, darker above; palpi brownish black.
Antennre brownish black' the flagellar segrnents elongate-oval,
with pale glabrous pedicels to produce a moniliform appearance.
Head dark brown.
Mesonotal prescutum brown, the humeral region paler; posterior sclerites obscure testaceous yellow. Pleura obscure yellow. Halteres dark brown, the stem paler. Legs with the coxe
and trochanters yellowish testaceous;femora brown, paler basally; tibire dark brown; basitarsi with about the basal two-fifths
dark broWh, the remainder of the segment and all remainder oftarsi except the outer ssgment snowy white; terminal segrnent
darkened. Wings (Plate L, flg. 11) whitish subhyaline, iridescent; stigma oval, relatively small, dark brown; veins nearly
black, with conspicuous macrotrichia. Venation: Sc' ending
opposite origin of Rs, Sc, some distance from its tip; free tip ofSc. far distad, R, being only about one-half the length of thelong apical spur of R,or; cell 1st M, relatively large, m-cu
shortly before midlength. fn one specimen, the free tip of Sc'
and R, are almost in alignment.
Abdominal tergites brown, the outer segments darker; sternites pale. Male hypopygium (Plate 3, fig. 34) with the rostralprolongation of the ventral dististyle, ad, of moderate length,
broad at base, the apex narrowed, provided with a conspieuous
seta; two rostral spines, placed at the base of the prolongation,
only slightly separated; both spines unusually small and spikelike, subequal, the innermost forms a slightly enlarged basal
swelling.
Habi,tat.-Japan (Riukiu Islands) .

334

The Phi,lippine Jownal, of Sci,encets2s

Holotype, male, Nago, altitude 30 feet, March 18, lgZB (S.
Sakaguchi). Paratopotypes, 4 males, mostly in poor condition.
Limonin breuicuspi.s is the largest species of the subgenus
Thrgpticomya, so far described.
LIMONIA(EUGLOCHINA)ARACENOBIA

sp. nov.

General coloration dark brown, the pleura paler; legs black,
including the proximal ends of all basitarsi; wings strongly infumed, the stigma darker; Rs very short, eneircling the proximal
end of the stigma; cell 1st M, present.
Male.-Length, about b.8 to 6 millimeters; wing, 6.b to 6.7.
Rostrum testaceous yellow; palpi black. Antenna black
throughout. Head dark brown.
Mesonotum shiny dark brown, the lateral margins of theprascutum and median region of the scutum paler. pleura morebrownish yellow, shiny. Halteres obscure yellow at base, passing into brownish black. Legs black, the coxa and trochantersconcolorous with the pleura; porximal ends of basitarsi extensively blackened,this including from one-third to approximately
one-half the length of the segment. Wings (plate 1, fig. L2)
with a strong dusky suffusion, the large oval stigma still darker;
veins dark brown. Venation: Sc of moderate length for amember of this subgenus,the distance on Sc2f R, between Sc:
and origin of Rs more than one-half longer than Rr; Rs very
short, arcuated, bordering the proximal end of the stigma,
shorter than the basal section of Rr*, i cell lst M, relatively
small, rectangular; m-cu beyond one-third the length of the cell;Cu, preserved to opposite one-third the length of the basal section of Cu,.
Abdomen black, the sternites obscure yellou', except laterally
and at apex.
H abitat.-Philippines.
Holotype, male, Majayjay, Lagtna Province, Luzon, altitude
above 500 meters, June 3, 1928 (R. C. McGregor). paratopotype, male, May 28, 1928 (n. C.'McCrregor).
Limonta wachnobia is a very different species from L. (8.)
okinawensis (A'lexander), the only other regional species withsimilarly patterned legs.
The following notes of occurrence by Mr. McGregor are ofunusual interest: "Hanging from a spider line by fore feet, inIight forest." and "On horizontal line of spider lysf-sams
habitat as Limoni,a (Thrgpti,cornAta) arachnophilz Alexander
and other members of the subgenus,at Lucban. Only three or

&r

Aierander: Tipulida from Eastern Asia, IV

335: -;:r .etn here, in rather light hillside forest, away from any::r.arr-.' The great value in the above observations lies in the:a;: :irat they are the first concerning this habit in any member:j ;he subgenus Euglochina. Doleschall, in 1857, described asa doubtful member of the genus Limnobia, a species that hecalled saltens,that had the habit of hanging on horizontal strandsof spider's webs. The arguments that have been made to settle
n'hether the species was a TltrApti,comgia or a Euglochfur,ahave
been discussedby the writer.s The above notes by Mr. McGregor and a study of a copy of Doleschall's figure of saltens sentme by Mr. Edwards have convinced me that saltens certainlypertains to the subgenus Euglochi,na, as long contended by Mr.
Edwards. In the light of our present knowledge of the fact that
there are numerous, closely allied species of Euglochi,na in the
Oriental and Ethiopian Regions, it would be very unwise toattempt to place L. cunei,formis de Meijere as a synonym ofsaltens without an examination of the type of the latter, if itis still extant.
LIMONIA(PSEUDOGLOCHINA)RIUKIUENSIS

sp. nov.

Mesonotum dark brown, the prescutum yellowish in front;
tibie snowy white, with trvo narrolv brownish black rings;
wings grayish, the stigma dark brorm; Rs relatively long, angulated at origin; r'ein 2d A unusually long, exceeding the
distal section of R,1u; abdominal tergites dark brown, the basal
segments with a yellowish median line; sternites yellow, s€gments four to seven, broadly dark brown at base.
Male.-Length, about 7 millimeters; wing, 6.5.
Female.-Length, about 7 to7.5 millimeters; wing, 6.5.
Rostrum, palpi, and antenna black. Head yellow.
Pronotum yellow. Mesonotal prescutum yellow in front andon the humeral region, the remainder dark brown; remainderof the mesonotum dark, paler medially. Pleura yellow, the sternopleurite extensively dark brown. Halteres infuscated. Legs
with the fore coxe and trochanters dark brown, the remaining
coxe and trochanters yellow; femora yellow, the tips infuscated,
more broadly so on the posterior legs; tibia snowy white, including the sete, the two rings brownish black, the first placed atnear one-fourth the length of the segment, the second justbeyond midlength; tarsi snowy white. Wings (Plate 1, fig. 13)
with the ground color grayish, the costal region a little darker;'Philip. Journ. Sci. 33 (1927\ 299.

336

The Phi,kppine Journal of Sciencetszc

stigma dark brown, long-oval, the proximal end sometimes morepointed; a very narrow brown seam at origin of Rs; veins darkbrown. Venation: Sc, ending beyond midlength of Rs, Sc,
opposite or just beyond the origin of Rs; Rs elongate and angularly bent near origin; petiole of cell 2d M, nearly one-halfthe cell; vein 2d A of unusual length for a mernber of this subgenus, on basal fifth approximating the margin, thence swinging
cephalad, the cell relatively long and wide, the vein beyond
the anal crossvein fully one-fourth longer than the distal sectionof R, ..
Abdominal tergites dark brown, the basal segments with anarrow continuous yellow median vitta; incisures and lateral
margins pale; sternites extensively pale yellow, the bases ofsegments four to seven broadly dark brown.
(Riukiu Islands).
Habitat.-Japan
Holotype, male, Shuri, Okinawa, altitude about 300 feet, March
8, 1923 (5. Sakaguchi). Allotopotype, female. Paratopotypes,
3 females.
Closely allied to L. (P.) bicinctipes (Brunetti) differing in
the larger size and details of coloration of the wings and abdomen. Vein 2d A is of unusual length for a member ofPseudoglocltina.
L IIIO\IA 'P S E L 'DOGL OCEI\- { )L- \- ICI\CTIPES

sp. nov .

General coloration of mesonotum dark brown, the median area
behind 1-ellori'ish;all tibire white with a single narrow brownish
black ring at near midlength; wing nearly hyaline, the stigma
conspicuous, short-oval, dark brown; vein 2d A short, approximately three.fifths to two-thirds the length of the distal sectionof Ro*r; abdominal sternites uniformly pale.
Male.-Length, about 4.4 millimeters; wing, 4.7.
Femnle.-T.ength, about 5 millirneters; wing, 5.2.
Rostrum and palpi dark. Altennre black throughout. Headpale, darker on posterior vertex, the latter area narrorvly dit'idedby a pale line.
Mesonotum dark brown, paler on the posterior portion of theprrescutum,this color increasing in amount behind, on the scutellum and postnotal mediotergite including the entire sclerite except laterally. Pleura yellow, the sternopleurite darker. Halteres darkened. Legs with the coxe pale yellorv, including thefore coxe; trochanters dark; femora slightly darkened basally,
the posterior femora broadly infuscated, the fore and middle
more narrowly so; tibia snowy white with a very narrow brownish black ring at near midlength; tarsi snowy n'hite. Wings

io,3Alerand.er: Tipulidu from Eastern Asia, IV

337(Plate 1, fig. 14) nearly hyaline, the stigma small, short-oval,
dark brown i reins black. Venation : Sc, ending beyond midlength of R**,Sc. opposite this origin; Rs short, a ]ittle exceeding
the ba*qaldeflction of Rn*u; m-cu at fork of M; vein 2d A short,the ba-salthird approximating the anal margin and parallel withit, the remainder arched and thence extending straight to themarg:in, the entire vein beyond the anal crossvein from threeftfths to two-thirds the distal section of Rn*r.
AMominal tergites brown, the basal two segments paler medially; sternites rnore uniformly pale brownish yellow. In themale the tergites are more darkened laterally at base, themedian and caudal portions extensively pale.
H abitat.-Phi I i ppines.
Ilolotype, male, Badajoz, Tablas Island, Romblon Province,
August 20, 1928 (Francisco Riaera and, A. C. DWag). Allotopotype, female.
In the single narrow dark tibial ring, Li,monin uni,cinctipes
agrees most closely with the larger L. (P.) kobusi, (de Meijere)
and,L. (P.) pulchri,pes (Alexander). It is difficult to know how
accurate is the figure given by de Meijere' of the wing of
L. kobusi. The features shown of a very short petiole to cell
2d M, and the position of m-cu at nearly its own length beyond
the fork of M are distinctive. From L. puJchri,pes the present
species differs in the short Sc,, the more oblique alignment of
the elements of the anterior cord, together with the shorter Rs
and the much shorter and more-arcuated vein 2d A.
EELIUS(HELIUS)TENUISTYLUS

sp. troy.

General coloration of mesonotum dark brown, the pleura morechestnut brown; rostrum black; legs dark brown, the tarsi paling
to yellow; wings with a faint brownish tinge, the stigma andcostal margin slightly darkened i Rr*, long, diverging gently
from Rn*u; m-cu at the fork of M; male hypopygium with thedististyles long and slender.
excluding rostrum, about 5.5 to 6.5 milliMald.-Length,
meters; wing, 6.5 to 8; rostrum, 0.7 to 0.8.
o<cludingi rostrum, about 9 rnillimeters;
Female.-I'ength,
wing, 8; rostrum, 0.8 to 0.9.
Rostrum black. Antennre black, the outer segments more
attenuated and slightly paler. Head black.
Pronotum blaek medially above, paler laterally. Mesonotum
dark brown, the humeral and lateral regions of the prescutum'Bijd.

tot de Dierkunde 18 (1904) pl. 8, fig. 6.

338

The Phi,l,i,ppi,neJou,rnnl of Sci,ence$zsbrighter brown; extreme posteromedian reg:ion of prrescutum
and median portion of scutum more yellowish; remainder of notum dark brown. Pleura chestnut brown, the dorsopleuralregion darker. Halteres dark brown, the base of the stem narrowly yellow. Legs with the coxre and trochanters brownish
yellow; remainder of legs dark brown, the tarsi paling to yellow.
Wings (Plate 1, fig. 15) with a faint brownish tinge, the stigma
and costal region a little darkened; veins brownish black. Venation: Rr*, long, gently sinuous, diverging gently from Rr*r; celllst M, widest at base; m a little longer and more oblique thanthe basal section of Mr; m-cu at fork of M.
Abdomen brownish black. Male hypopygrum (Plate B, fig.
35) with the dististyles long and slender, the outer style, od,chitinized, the apex obtuse; inner style, id,, longer, subangularly
bent. Mesal face of basistyle, b, produced into a weakly spinous
lobe.
Habi,tat.-Formosa.
Holotype, male, Kirakei, altitude 4,000 feet, June, 28, lgZZ (S.
T.Issiki).
Allotype, female, Chipon, July 4, LgZ8 (S. ?. Issi,ki,);paratypes, 3 males, with the allotype.
Heli,us tenuistylus belongs to a group of closely allied forms
that center about H. ruigriceps (Edwards).
The species differsfrom I/. nigriceps in the longer, more gently sinuous Rr*, andthe structure of the male hypopygium.
EELIUS (HELruS) ATTENUATUS sI'. noy.

General coloration of thorax brownish black, the pleura morebrownish yellow; rostrum black; legs brownish blaek, the tarsi
paling to brownish yellow; wings with a dusky tinge, the stigma
and costal region darker brown; Sc long, Sc, ending opposite
or beyond the fork of M; Rr*, long, gently sinuous; m-cu atfork of M; male hypopygium with the outer dististyle truncatedto weakly bidentate at apex; inner dististyle prolonged intoa slender yellowish apical portion.
Male.-Length, excluding rostrum, about b.b millimeters;
wing, 5.5 to 5.8; rostrum, about 0.5.
Female.-Length, excluding rostrum, about 6 millimeters;
wing, 5.5; rostrum, about 0.5 to 0.6.
Rostrum dark brown, palpi black. Antenne black throug:hout; basal four flagellar segments larger, the remainder narrowed. Head black, the ffont more opaque and slightly gayish.
Pronotum and mesonotum brownish black, the median areaof the scutum paler. Pleura brownish yellow, the dorsopleural

Aleuander: Tipulida from Eastern Asia, IV

40, 3

ggg

region dusky. Halteres dusky, the base of the stem very restrictedly paler. Legs with the coxe and trochanters brownish
yellow; femora and tibia brownish black, the tarsi paling tobrownish yellow. Wings with a strong dusky tinge, the stigma
and costal region darker brown; wing apex and a seam alongvein Cu vaguely infumed; veins dark brown. Venation: Sc
long, Sc, ending opposite or shortly beyond the fork of Rs,
longer than Sc, i Rz+s long and gently sinuous; cell lst M,
generally parallel-sided, the proximal end not or but slighfly
widened; m-cu at fork of M.
Abdomen, including the hypopygium, black. Male hypopygium (Plate 3, fig. 36) with the outer dististyle, od,, much shorter
than the inner, the apex irregularly truncate to weakly bidentate, in the latter case,the inner tooth more acute. Inner dististyle, id, long, the apical portion elongate, pale yellow; baseof style with numerous sete, including a fringe of four or five
major bristles along the mesal edge. Basistyle, b, without alobe.
Habitat.-Formosa.
Ilolotype, male, Chipon, July 4,1927 (S.
Issi,lci). Alloropotype, female. Paratopotypes, 3 males.
".
Hel;tns attenuntus is closely allied to H, nigriceps (Edwards),
differing especially in the darker coloration of the thorax andwings, the details of venation, as the long: Rr+g, and the detailsof the male hypopygium, as the very long apical portion of theinner dististyle.
HE) <ATOMINIPEYLLOLABISBEESONI

ap. noy.

General coloration pale testaeeousyellow; legs pale yellow, theterminal tarsal segments darkened; wings with a pale brownish
yellow suffusion, the base and costal region clearer yellow;
cell lst M, closed,m-cu at the fork of Mr*n; abdomen dark brown,
the genitalia brighter.
Female.-Length, about 6.b to Z millimeters; wing, 7.2 to g.
Rostrum pale, the palpi brown. Antenna of moderate length,
if bent backward extending to shortly beyond the base of thehalteres; antennre pale, only the outer segments a trifle darker;
flagellar segments elongate, a little enlarged at bases. Headlight grayish brown.
Pronotum and mesonotum testaceous yellow, the pleura alitle more yellowish. Halteres pale, the knobs a trifle darker.
Legs pale yellow, the terminal tarsal segments darkened. Wings
246465_4

The Phili,ppine Jout'nal of Science

340

tezs

(Plate 1, fig. 16) with a pale brownish yellow suffusion, thebase and costal region clearer yellow, veins brown, the prearcularand costal veins brighter. Venation: Cell 1st M, closed; m-cu
at the extreme outer end of the cell, opposite the fork of Mr*n.
Abdomen dark brown, the genital segment and ovipositor yellowish horn-color; tergal valves of ovipositor elongate and only
gently upcurved.
H abitat.-British India.
Holotype, female, Mussoorie, Himalayas, altitude 6,500 feet,
August, 1927 (C. F. C. Beeson). Paratopotype, female.
I take great pleasure in naming this interesting crane fly
in honor of the collector, the Forest Entomologist, Dr. C. F. C.
Beeson. Phyllolabi,s beesoruiis very different from the only
other known Himalayan member of the genus, P. confl,uenta
Alexander.s
LIMNOPIIILA

AINO

slr. nov.

Male.-Length, about 5.8 to 6 millimeters I wing, 6 to 6.5.
Closely allied to L. dicranophragmoid'es Alexander (Riukiu
Islands), differing chiefly in the arrangement of the color pattern.
Antenne n'ith the first segment of scape black, the second obscure brourrish 1-ellorv; flagellum chiefly yellow, the outer segments a little darkened. Mesonotal prascutum grayish brown,
the three usual stripes entire, the rnedian stripe wide, narrowly
divided by a more grayish median vitta; lateral stripes narrow;pseudosutural fovee and tuberculate pits black, conspicuous,
the latter some distance back of the cephalic margin of the sclerite; centers of scutal lobes and the median area brown, yellowish
pollinose; scutellum and postnotum brown. Pleura chiefly darkbrown. Halteres yellow, the knobs conspicuously dark brown.
Legs with the coxa dark brown; remainder of legs yellow, theouter tarsal segments more infuscated; legs rvith conspicuous
semierect pale sete. Wings with the ground-color more whitish
subhyaline, the pattern arranged in similar manner to L. d:icranophragmoid,esbut the individual bands more separated, especiallythe one at origin of Rs and the one along the cord; the latter
two are separated by a wide, generally clear band that is approximately as wide as the first-mentioned dark band; dark bandsof wing more compact, the pale spots and dashes within them
being much reduced; along the costal margin the four or fiveuRec. Indian Mus. 29 (7927) 206-207,fig. L7.

40.3

Ale*ander: Tipulidp from Eastem, Asia, IV

B4l

outer areas are solidly darkened, not with a pale central spot
as in Z. d;icranophragmoides. Venation: Sc short, Sc, ending
opposite or shortly before the fork of Rs; inner ends of cells
R.,
R' and 1st M, in oblique alignment, the last most basad; m_cu
not far beyond the fork of M. Abdomen chiefly brown, the tergites darker laterally; a narrow subterminal darkring; hypo_pygium reddish brown; basal sternites more reddish
brown,
darkened laterally and caudally, the outer segments more uniformly darkened. Male hypopygrum (plate B, fig. BT) withthe outer dististyle, od, appearing as a blackened rod, ifr" tipstrongly curved into a spine, the outer margin quite smooth,the inner margin, in the axil of the terminal spine, with a few
microscopic denticles. rnner dististyle, id,, very broad, mittenshaped. Gonapophyses,gf, appearing as pale rods, the apex ofeach dilated into an oval head.
Habitat.-Japan(Hokkaido) .
Holotype, male, Jozankei, altitude about 1,000 feet, July 29,
1923 (S. Kuwagamn). paratopotypes, 9 males, mosfly in poor
condition.
ERIoPTERTNT.LADURA

ALprcoLA

sp. nov.

General coloration pale yeilow; legs yellow, the extreme tipsof the femora and tibia infuscated; tarsi extensively brownish
black; wings pale yellow; a tiny black spot at extreme base ofwing; male hypopygium large and complex, the basistyle with alarge mesal lobe that is unequally bilobulate near tip; inner dististyle with a conspicuous basal shoulder.
Male.-Length,7.5 to 8 millimeters; wing, g.b to 9.
Head, thorax, and abdomen with long conspicuous pale setre.
Rostrum brownish yellow, palpi darker. Antenna wiitr ttre basal segment dark brown, in some cases so colored only basally,
the remainder yellowish; outer flagelrar segments darker; verticils elongate. Head pale brownish yeilow, more rarely darker,
grayish brown.
General coloration of prothorax and mesothorax pale yellow,
immaculate. Halteres pale, the knobs weakly darkened. Legs
with the coxa and trochanters pale yellow; femora yellow, theextreme tips infuscated, tibire yellow, the tips narrowly darkened; tarsi extensively brownish black; legs conspicuouslyhairy.
wings (Plate 1, fig. 1T) pare velrowish; a single tiny bracir dot at
extreme wing baseabove the squama; veins brown, the prearcularand costal veins more yellowish. Macrotrichia of veins long

The Phi'lippi'ne Journal of Sci,ence

342

r'eze

and conspicuous. Venation: Sc, relatively short, only a little
longer than Sc,1 Rs long, gently arcuated i Rz+e+ashorter thancell 1st Mr; petiole of cell M' subequal to or considerably exceeding m; m-cu from one-half to nearly its own length beyond
the fork of M; vein 2d A long, very gently sinuous.
Abdomen yellorv, the tergites with narrow' interrupted median and lateral dusky stripes, in cases including the subcaudal
portions of the segments; sternites similar, their caudal margins
darkened; hypopygium pale. Male hypopygium (Plate 3, fig.
38) very large and complicated in structure, much as in C.
megacaud,a, differing especially in the structure of the tergite,
basistyle, and dististyle. Basistyle, b, with the mesal lobe large'
flattened, at apex somewhaf expanded, on outer margin before
apex with a small fingerlike lobe terminating in a small pencil
of seta. Oute'r dististyle, od, very long and pale, slender' considerably longer than the inner style, setiferous, with larger
seta at apex. Inner dististyle, id, distinetly bilobed, there being
a stout basal shoulder, the more slender inner arm truncate
at apex. Gonapophyses,gf, taken together appearing lyriform,
relativell- slender, the obliquely truncated apex weakly toothed.
Habitat.-Japan (Honshiu).
Holotlpe, male, Yarigatake, Japanese Alps, August 24, L925
(Jiro llachitla) . Paratopotl-pes,4 males; paratypes, 3 males,
Tsubakuro, JapaneseAlps, August 23, 1925 (Jiro Machida).
Clnud.ura,alpicola is most closely allied to C. megacauda Alexander (Hokkaido, Japan), differing especially in the detailsof struc,ture of the male hypopygium. These two species belong
to a distinct group of the genus that can scarcely be referred tothe subgenas Neoclnduro Alexander, the only other divisionof the genus in which the male hypopygium possessestwo
dististyles.
GYUNASTESCATAGRAPIIA

ap. noY.

? Teucholabis sp. nov. BnuNETTt, Rec. Indian Mus' 15 (1918) 306.

Head orange; mesonotum yellowish, the prescutum with a
median black stripe; knobs of halteres orange'yellow; femorawith a yellow subterminal ring; wings whitish subhyaline,with three brown crossbands; abdominal tergites dimidiateorange and dark brown.
Mal,e.-I'ength, about 6.5 millimeters; rving, 5.7.
Rosf,rum shiny reddish yellow; palpi brownish black. Antennre with the basal segment orange; second segment brown, paler
at base; flagellum brownish black ; flagellar segments elongate-,ro.3

Alerander: Ti,pulida lrom Eastern Asi,a,IV

343

oval to subcylindrical, with elongate verticils. Head broad,obscure orange.
Pronotum obscure orange-yellow. Mesonotal prrescutum yelIow, the median stripe shiny black, the lateral stripes reddish,poorly delimited; scutum shiny yellow medially, each lobe with ablackenedspot near mesal portion, the remainder reddish; scutellum brownish blaek, brighter basally; postnotum blackened,
paler laterally. Pleura badly injured by the pin, the propleura
reddish brown, the mesopleura apparently darker brown, variegated with paler. Halteres dusky, the knobs orange-yellow.
Legs with the coxre and trochanters reddish yellow; femora
obscure yellow, the distal half passing into dark brown, witha conspicuous subterminal yellow ring that is a little narrower
than the darkened tip; tibiae brown, becoming darker browntoward the tip; tarsi brown, the distal segments darker. Wings(Plate 1, fig. 18) whitish subhyaline, the prearcular and costal
region more yellowish; a heavy brorvn pattern, appearing as
three crossbands;basal band narrorv, almost parallel-sided,extending from R at the origin of Rs to the anal margin at theend of vein 2d A, verl'narrowly interrupted in cell R adjoining
vein M; the second band includes ttre cord, widened out at thestigmal region, narrower along the cord, again widened out
but much paler in the outer end of cell Cu; the third band
includes the wing-tip in cells R, Rn, R, 2d M' and M., the inner
margin nearly straight; a broad additional seam at the outerend of cell 1st Mr; veins pale brown, a little darker in the infuscated areas. Venation: Sc, ending shortly before the forkof Rs, Sc, some distance from the tip of S",l Rr*r*, a little
longer than R, alone; Rr*, punctiform; R, directed a litile basad, about one-half Rr*, i veins R, and Rn widely divergent,
the former about one-half the latter; cells R, R., and 1st M,
in slightly oblique alignmont; cell 1st M, rectangular, a litile
widened outwardly; m-cu less than its length beyond the forhof M.
Abdominal tergites conspicuonsly dimidiate, the basal two_
thirds of each segment dark brown, the remainder orange; sternites similar but the dimidiate appearance not so conspicuous,
especially on the basal segments where the ground color is pale
brown, the incisures broadly orange; on the subterminal seg_ments the colors are more contrasted; hypopygium obscure ful_vous; no sternal pouch. Male hypopygium (plate B, fig. Bg) verysmall; caudal margin of the tergite convex. Basistyle, b, rela_

344

The Phi,lippi,ne Jownnl of Sci,ence:g,2s

tively large, the outer angle produced, the dististyles being placedon the mesal face of the style at near midlengfh; two small blackened combs on the style. Outer dististyle a flattened blade,
the outer angle at base darkened and microscopically roughened;
apex obtuse, darkened, sparsely setiferous, the margin with microscopic serrulations. Inner dististyle smaller, the margins
smooth, before the apex with a powerful brisile. ,lEdeagus,o,very large, appearing as a conspicuouspale blade that juts cau_dad beyond the other elements of the hypopygium, the apex
obtuse but extended into a short point on the ventroapical portion; distal portion of adeagus with conspicuoussete.
H abi.tat.-Southwest India.
Holotype, male, Castle Rock, North Canara District, OctoberlL to 26,1916 (S. Kemp). Type in the collectionof the IndianMuseum.
GEmnastes catagra'pha is allied to G. d,emeijerei, (Riedel),6
differing in the chiefly yellow body coloration, as well as in the
differently colored halteres and legs.
STYRINGOMYIATABLASENSIS

sp. nov.

General coloration yellow, variegated with brownish blaek;head gray; femoral dark rings incomplete; wings yellow, withfour separate dark areas; vein 2d A simple, the apex genily
curved; male hypopygium with the ninth sternite deepiy bilobed at apex; outer arm of dististyle a long rod that nu"ro*,gradually to the acute simple apex.
Male.-I'ength, about 4 millimeters; rving, 4.2.
Antennal scapedark, the flagellum conspicuouslypaler. Headgray.
Pronotum brownish black, narrowly paler medially; posterior
notum paler laterally. Mesonotal prrescutum with the median
portion brownish black, the posterior twethirds fulvous-ye,llow
with a sinuous black line on either side of the more cinerous
median area; scutal lobes and median area chiefly pale, com_pletely margined laterally and caudally with black: scutellum
black, the median area restrictedly pale; postnotal mediotergite
brownish black. Pleura pale, yellowish, the color including thepleurotergite. Halteres pale yellow. Legs with the coxeandtrochanters yellow; femora yellow with two narrow incomplete
dark rings, as usual in the genus, the ventral surface of thefemora immaculate; tibie yellow, the tips darkened; tarsi yellow,
uAnn.

Mus. Nat. Hung. 19 (1921) 1gb-186, new name tor Gnophomgio
fascipennis de Meijere, Bijd. tot de Dierkunde 21 (1919) 15, preoccupied.

,ro,B

Alerander:

Ti,puliila from Eastern Asia, IVUE

the last segment black. Wings with a yellowish suffusion; four
dark brown areas, as follows: A large oval area centering about
r-m, ineluding the extreme basesof the adjoining veins; fusion of
Mr*, and M,; m-cu; distal third of vein 2d A; veins yellow,
brownish black in the infuscated areas. Venation: A short fusion of M, with Mr*, beyond cell lst M,; vein 2d A gently
curved to the margin.
Abdominal terg:ites yellow, each segment with two brown spots
at caudal margin, on the seventh segment becoming larger andconfluent; eighth segment with a longitudinal brown medianstripe; hypopygum chiefly pale; sternites more uniformly paleyellow. Male hypopygium (Plate 3, fig. 40) with the ninth
tergite, 9f, a broad pale setiferous lobe, the apex very insensibly
bilobed. Ninth sternite, 9s, very deeply bilobed, each lobe bearing a very long seta. Basistyle, b, with the outer apical angle
a slender pale lobe that bears a slender seta that is longer thanthe lobe itself. Dististyle, d, very complex, the outer arm along rod, its base wide and pale, the outer end narrowed to theacute gently curved apex, entirely without a long apical seta,
as in many species of the genus. Two arms, gently curved andmore or less opposed to one another, bearing series of peglikespines, those of the outer arm (about 15) more crowded, of
the inner or cephalic arm (about 9) more scattered; other lobesof the complex dististyle include a simple boomerang-shaped
structure and s1broad-based lobe that is tipped with one or two
powerful black spines and bears a long powerful seta beforeapex.
H abitat.-Philippines.
Holotype, male, Badajoz, Tablas Island, Romblon Province,
August 27, t928 (Framci,sco Riaera and A. C. DWag).
STYRINGOIIYIANIPPONENSIS

sp. nov.

Male.-Length, about 5.51millimeters; wing, 5.2.
Closely allied to S. formosano Edwards, differing especiallyin the structure of the dististyle of the male hypopygtum.
Second segment of scape dark brown, the base paler. Largest
seta of head black, a few smaller ones yellow. Menosotal prascutum dark brown, with longitudinal paler stripes, the darkareas of the interspaces interrupted at the level of the pseudosutural fovee; posterior median reg:ion of prrescutum, medianarea of scutum, and central portion of scutellum yellow, scutal
lobes reddish brown, encircled by darker, each with an erect setanear mesal edge; disk of scutellum with an erect seta on eachside of the yellowish median area; postnotum darkened. Pleura

346

The Plr,i,lippine Journnl of Sci,ence

pale yel,low. Halteres pale, the knobs weakly infuscated. Legs
with the dark areas on femora narrow and restricted to thedorsal portion; medial tibial ring similarly incomplete, the apicalring narrow but entire; basai four tarsal segmentsalmost white,
incisures weakly darkened; terminal segment black. Wings witha yellowish suffusion, the base and costal region brighter; acircular dark spot on r-m; Iess distinct dark seams on m-cu,
outer end of cell 1st ]I" ancl clistal third of 2d A; medial veins
beyond cord chiefly darkenecl; radial, costal, and cubital veins
pale. Venation: Rz-s-r oblique; m-cu nearly its own length
beyond fork of II; vein 2d A ltent strongly at end, subangulate
but unspurred. Abclominal tergites obscure yellow; basal segment dark; succeedingsegmentss'ith pailed dark spots on caudal
margins and less distinct oneson basal ring of segment; segment
7 with a median darkening; hypopygium chiefly pale. Male hypopygium with the dorsal apical lobe of basistyle rvith a longer
and more hairlike apical bristle. Dististyle with the outer lobe
long and slender, the apical seta long, as in f ormosana; anterior
lobes of dististyle (Plate 3, fig. 41) broad 4t base, one narrowed
apically into an acute pale point, the apical margin with two or
three groups of blackened pegs, the more proximal of thesearranged more or less in the form of a crescent; innermost lobeof dististyle with the apex obtuse, the outer caudal angle rri'ith
numerous black peglike spines that decreasein size and become
more separated outwardly; surface of style basad of these pegs
with very long, basally directed seta. Ninth sternite very
narrow, entire, with the usual two apical seta.
(Honshou) .
Habttat.-Japan
Holotype, male, Asahara, June 7, L925 (C. Harukawa).
It may be noted that Stgringomyiu f ormosana Edwards has
been taken as far north as Tokyo (May 14, 1919, R. Takahs,shi,\and is thus the most northerly record for the genus.

ILLUSTRATIONS
lLegend:o, adeagus; b, basistyle; d, dististyle;dd, dorcal rtististyte; g, sonapopyhysis;
id' inner distiBtyle; od, outer dististylc;r, tergite;& sternite;
od, ventral dististvre,lPr,lroFrc.

1

7. BNthura scnxctq,sp. nov., wing.
2, Brithura coni,frons Edwards, wing,
3. Tipula ranto,icola sp. nov., .wing.
4. Tipula ruicrocellula Alexander, wing.
5, Nesopeza rd,nta,iaa,nn sp. nov., wing.
6. Oropeza bispinula sp. nov., wing.
7. Limoni,a (Itimonia) ro,ntaiensis sp. nov., .riling.
8. Li,monia (Di,cranomy,ia) tri,stoid,es sp. nov., wing.
9. Limonia (Geranomyin) suensoni,ano sp. nov., wing.
10. Li,monia (Alenand,riaria)
orgyrata sp. nov., wing.
ll. Limorui,a (Th:rgpticomgia)
breaicttspi,s sp. nov., wing.
12. Limonin (Euglochina) q,rd,cllnob,is,sp. nov., wing.
!3, Lim"onia (Pseuilogloch,ina) riukiuensas sp. nov., wing.
14. Limnnia (Pseudoglochina) unicinctipes sp. nov., wing.
75, Helius (Helius) tenuistElus sp. nov., wing.
16. Ph,gllolabis beesoni sp. nov., wing.
17. Cladura alpi.cola sp. nov., wing.
t8, Ggrrnnostes cd,ta,grapho sp. nov., wing.
Pr,,c.tp 2

FrG. 19, Brithu,ra sancta sp. nov., male hypopygium.
20. Ti,ptla *ubfutili,s sp. nov., male hypopygium.
27. Tipula subfutilis sp. nov., male hypopygium, ninth tergite.
22. TiTtu.la futi.Iis .liexander, male hypopygium, ninth tergite.
23. Tipu.la futilis .Llexander, male hypopygium.
24. Tipuln Ausouoid,es sp. nov., male hypopygium.
26. Tipulfr subgtusou sp, nov., male hypopygium, gonapophysis.
26, Ti,pula gusou .lJexander, male hypopygium, gonapophysis.
27, Nesopeza iiliopha,llus sp. nov., male hypopygium, adeagus,
28. Oropeza satsumu Alexander, male hypopygium.
29. Oropeza satsurna, Alexander, male hypopygium, ninth terg,ite.
30. Oropeza bispinula sp. nov., male hypopygium.
31, Oropeza bispi:nula sp. nov., male hypopygium, ninth tergite.
32. Lixnonia (Dicranomgia) tristoiiles sp. nov., male hypopygium.
Pr,lrnB

Frc. 83. Limnni,a (Geronamryia) tenuispinos@ sp. nov., male hypopygium.
34. Limonia, (Thrgpti.co,rngia,) breaicuspis sp. nov., mate hypopyg:ium.
847

348

The Phttippine Jourywl of Sei,enceFte. 35. Eelfus (Eelias\ tenui,stglus sp. nov., mple hypopygium.
36. Halias (Hetius) attenuatus sp. nov., rrale hypopygium.
37. LhrnnophiJa aino sp. nov., male hypopygium.
88, Claihna a,lpi,cola sp. nov., male hypopygium.
39. Ggmnastas catagrapha sp. nov., male hypopygium.
40. Stgringomgi,a tablusensrs sp. nov., male hypopygium.
41. Stgringomgia nipponensi.s sp. nov., male hypopygium, dististyles.

t
-i
',t

TTPULTD.€ FROM EASTERN

AsrA,

IV.l

PL AT E 1 .

IPH r l ,r p. J ounw . Sc r ., 40, N o. B.

ALEXANDER: TrpuLrD_& FRou EASTERN AsrA, IV.llPHrLrp. JouRN. Scr., 40, No. B.

/\.,4

/\

/l

qq

n"
25

>

26

fa-^^q29:
I30@-^--h\31

PLATE 2.

ALEXANDERTTrpuHDE FBoM EASTERN AsrA,
IV.l

0d

.'

[pllrl,rp.JouBN.Scr.,40,No.B.

.f\

38

PLATE 3.

fi