26f
UNDESCRIBEDCRANEFLIESFROM THE HOLARCTICREGION IN THE
UNITED STRIESNAIIONAL MUSEUM

BY

CHARLESP. ALEXANDER
Of the Mtrechurcur A3riculturrl Collcae,Anbcnt

No. 2698.-From th€ koceedingsof thc United StatesNational Murann
Yol.72, Pu.t.2,pp. l-17, with pl. I

UNITED STATESGOVERNMINT PRINTING OFFICEWASHINGTON
t927

UNDI]SCRIBED CRAI{E FLIES FROM THE HOLARCT'ICREGION IN THE UNITED STATES NAI''IONAL MUSBUM
By Crranr.nsP. Ar,nx.rwonn
Of t,he MassachresettsAgricul,tural, Col'l'ege,Amluerst

The craneflies (Tipulidae) describeclat this time were included inlarge seriessent to me for determination by Dr. J. M. Aldrich and
Dr. H. G. Dyar, the majority having been collectedby them in the'W'esternStates and Canada. The material from western China wascollectedby the Rev. D. C. Graham and presentedby him to thenational collection. I wish to expressmy thanks to thesegentlemen
for their kind cooperationin maliing knourn these neglectedflies.
The types are preselvedin the collectionof the National }{useurn,with tlre exceptionof that of I)icrctnoptycltaoccklentttlis,
ne\y species.
P.TLAE^{RCTIC SPECIESBRITHURA ^IYMPHICA, new sp€cies

General eoloration rich brown I antennal flagellum yellow; dorsopleural region of thorax conspicuouslyochreousI ventral portion of
the pleurotergite producedinto a flattened lobe, the.cephalicside ofwhich is densely velvetyI femora yellow, the tips conspicuously
blackened;wings brown, sparselyvariegated with yellow i 8c, lackitg; Htn reduced; ntrctr long and sinuous,longer than the distal
section of Cw'
Fernale.-Length about 38 mm.1 wing, 24 mm. I abclornenalone,25
mm. ; its greatestwidth, 6.4 mrn.
Frontal prolongationof the heucllonc, clark chestnutbrorvn, thenasuslong and powerfully der-eloped; basalsegmentsof palpi darkbrown, the third segmentpaler, the last segmentblack with theextreme tip brightened. .\ntennae with the first scapal segment
dark reddish brown. the seeondsegment fulvous; flagellum paleyellow, the basal enlargernentof the individual segmentsscarcely
darker; verticils all basal in position, r'ery long and conspicuous,
including thoseof the redueeclterminal segment. Head dark brown,
the orbits paler; vertieal tubercle conspicuous,the tip slightly
decurved.
N o . 2 6 9 8 .-P R o ce e D tN Gs
U. S. Nnr toNALMuseuM ,Yot.72. ART.2
55413-27-1
1

PROCEEDINGS OF TEENATIONAIJ

MUSEUIIT

vOt+ 72

Pronotum dark clove brorvn, the lateral margins paler, the scutellum brighter, narrorvly margined anteriorly with blaek. Mesonotal
praescutumbrorvn with four barely evident brighter brorvn stripes,
the posterior scleritesof the mesonotumgenerally of this samebrightbrown. Pleura brorvn,the dorso-pleuralregion extensivelyand conspicuously pale ochreousI ventral sclerite of the pleurotergite producedlaterad into a conspicuoustriangular flattenedlobe,the cephalicface of rvhich is denselycoveredwith a cream-coloredvelvety pile.
Halteres brorvnish yellorv, the hnobs clark brown. Legs with thecoxaeand trochantersdark cinnamonbrown; fernorayellow, the tips
rather broadly (2 mm.) and conspicuouslyblackenedI tibiae brownish yellorv, the tips narrorvly infuscated; tarsi light brown, the outer
segmentsdarker. \f ings of the usual Britlwa type, the membraneyellorvish, extensively variegated with darker, or brorvn, sparselyrariegated with yellow; prearcular region largely pale; cell C
darlrel brorvn, Ec more yellorvish; stigma relatively pale, withbrighter vellow markings before and beyondit I a srnall darker brorvn
spot at origin of Bs, precededand followed by the vellow markings
desclibedI anterior eord a little seamedwith darker; a small yellow
area near outer end of cell ^Br1 similar yellow marginal areasin cells.B, to .ilfn,inclusive, rvith tlvo similar rnarkings in the first anal andanother in the secondanal cells,theselatter areaslying not far fromthe veins; a small brown r:loudin cell Cu not far from midlength; upale yellorv, gently arcuatedtransverseare& acrossthe outer end ofcell JII; r'ein Cu, beyond this rnark, as well as rlt/-clcconspicuously
seamed.
with dusky I outer end of cell f,rst M * fork of M and marginal
spots at the ends of the veins, as well as the wing apex in the radial
field, and the posterior margin of the wing in the anal cells, moredusky. Venation (fig. 1) : rSc,lacking j Sc, ending just before r, thelatter closeto the fork of Rr*r; r-ln reduced;cell first M, pentagonal;
petiole of cell IlI, a little longer than nr; mrcu strongly arcuated,
longer than the distal sectionof Cu, insertedon M n,shortly beyondits
origin j Cu, attaining the margin at the end of Cwr; cell secondA'wide.
Abdomen rich cinnamon brown, the lateral region of the tergites
broadly darlier brown I caudal margins of the segments narrowly
and obscurely, the extreme caudo-lateral angles more brightly yellowish, most conspicuous on segments two and. three I sternites
somewhatdarlrer brown, the caudal margins more sooty brown, theextreme lateral angles of the segmentsagain with yellow triangles.
Dorsal shield of ovipositor obscure yellow, the conspicuoustergal
valves dark chestnut horn color.
Habi,tat.-China (Szechuen). Holotype, female, Shin Kai Si,

10,L922(D. C. Graham).
Mount Omei,altitude4,400feet,September
Type.-Female, Cat. No. 403L7,U.S.N.M.

enr.2N"EW UNDESCRIBED CRANE FLIES-ALEXANDER,

3

The speciesis readily distinguishedfrom all describedforms by
the colorationof the antennaeand legs.
ERIOCERAGRAEAMI,new opeclel

Head and thorax velvety black, the scutellum orange; abdomenwith tergites one to four orange, all except the first with a narrow
black border along the caudal margin I remaining tergites black;wings blackish; cell ^lll,present.
Female.-Length, 20 mm.; wing, L7 Dr.r the greatest width,
5.4 mm.
Rostrum and palpi black. Antennae brownish black throughout,
if bent backward not attaining the wing root. Thorax velvety black,the scutellum orange. Halteres and legs black. lYings relatively
broad, blackish, the base and costal region clarkerI veins & little
darker than the ground color. Venation (fig. 2) : Rs long, about onethird longer than R; Sc, ending beyondthe fork of Rr*"*nthe latter
equal to the basal sectionof Bu; basal sectionof R, transverse,Rr*,equal to one-half llr*r; cell 11, present,shorter than its petiolel m-cu
at about two-fifths the lower face of cell first M, straight, about onethird longer than the gently alcuated distal sectionof Cur; cell Cu
at n'ing mnrgin about trvo-fifths as wide as cell ^try'n.
Abclominal tergites 1 to 4 orange, the lateral margins weakly
infuscated, segments2 to 4 with a very narrow but conspicuous
black line across the caudal margin, the segmentswithout shiny
bands; tergites 5 to 8, velvety black I sternitessimilar, but the black
caudal marsins of the basal segmentslacking. Ovipositor with theclorsal shield brownish black dorsally; tergal valves of ovipositor
broken.
Eabitat.-Ohina (Szechuen). Holotype, female, Shin Kai Si,
Mount Omei, altitude 4.400feet, July 1-30, Lg25(D. C. Graham).
Type.-Female, Clat.No. 40318,U.S.N.DI.
This interesting crane-fly is named in honor of the collector, theRev. David C. Graham, who has added greatly to our knowledgeof
the zoology of western China. By Edwards's key to the Old World
speciesof Eri'ocer"aLthis speciesruns to E. umbrigtennis Edwards(Penang), an entirely distinct species.
ERIOCERAFIIMIDIPENNIS,

new rpeclel

General coloration dull gral: the praescutum with three blackish
stripes; legs black, elongateI wings relatively long and narrow,strongly blackish, the costal margin darker i Rr*, about one-halflonger than Er', Er*, and.E, subequaliRr*r*nshorter than the basal
section of Bu; cell .i|y',present.
Male.-Length about 17.5-18mm.1 wing, 16.5-18mm.
lAnn. Mag. Nat. Hist., ser. 9, vol. 8, pp. 6?-?8, 1921.

4

PROCEEDINGS OF TEE NATIONAI, MUSEUI\Ivou.72

Rostrum and palpi brownish black. Antennae relatively short,brorvnish blacli throughout, if bent backward, scarcelyattaining thewing root. Head dull brownish gray: the vertical tubercle low ancl
relatively inconspicuous,blackish. Mesonotumblackish, with a dull
brownish gray pruinosity, the praescutumwith three blackish stripes.
Pleura g*yn variegated with blackish areas. Halteres brownish
blaek, the base narrowly brightened. Legs with the coxae gray;trochanters brownish griayI remainder of legs black, elongate (hinclleg, femur, 13 mm.;tibia, 14 mm.; tarsus,11.6mm.). wings relativefy long and.narrow, strongly blackish, the costal margin tlarker I
stigma ttouil, oval, darker brown than the ground color I veins dark'
Venation: 8c, ending about opposite three-fourths .Er*r, St" solne
distance from its tip ; Es long, strongly angulated at origin 1 Rr*r*n
shorter than basal sectionof .R,i R,.*, about one-half longer than thebasal section of .8,, or a little lessl Rr*, subequal to Rri r-4tf inalignment with the inner end of cell first M "; cell M, present,a little
sh,Jrte1.than its petio\el nvcu not far beyoriclthe fork of. n[, about
one-third longer than the distal sectionof Cu*
Abd.omenblackish,sparselypruinose,the hypopygium concolorous.
Habitat.-China (Szechuen). Holotype-,male, Suifu, I\Iay-June,
L92L (D. C. Graham). Paratopotype, rnale'Type.-Male, Cat. No. 40319' U'S'N'M.'
ny na.wardsiskey to the old worlcl Eriocera2 this fly would run
walker (Borneo), a very difrerent species.
to E. r,u,brescens
ERIOCERA CYBELE, new sPecies

Velvety black; abdominal tergites 1, 2, 4, and' 5 with basal glab'with arous rings; genital segmentof female fulvous I wings black
long, naruow, white, transverseband before the cord i R , oblique'Male.-Length, about 13 mm.1 wing, 11'5 mm'
Femntre.-Length, 19 mm.; wing, 16 mm'
Head and appendagesvelvety black.
Thorax entirely velvety black, including the halteres and legs'
Wings black, the anal cells sornewhatpaler I a long, narrow, transo.r..] discal, white arear extending from vein -E ' before the cord,nearly to the posterior margin of the wing in cell cu. venation:in position, lessobliqtre
Basai sectionof n, obliqueto subtransverse
than in E. obti,qtn; cell .R, Ionger and narrower than in the lastnamed species.
Abdomen velvety black, the segmentsrelati'sely short; tergites 1
and 2 with shiny glabrous basal rings of silvery gray; segments
4 and b similar but the rings opaque;tergite 3 uniformly black except
for a very narrow glabrous basal ring. Male hypopygium entirell'
2 Ann. Mag. Nat. Ilist., ser. 9, vol. 8' 1921.

enr.2

NEW UNDESCRIBED CRANE FI,IES-AI,EXANDEB

5

black. Female with the genital segmentbright fulvous, the elongate
valves of tlie ovipositor rnore horn-colored.
(Szechuen). Holotype, male, Shin Kai Sit
Habitat.-China
Mount Omei, altitude 4,400 feet, July 1-30, L92L (D. C. Graham).
Allotopotype, female. Paratopotype, female.
Type.-Male, Cat. No. 40320,allotype, female,U.S.N.M.
Eriocera cybele is another of the numerous recently discovered
speciesof the genus that run to E. hi,lpa Walker, by Edwards's key
to the Old lYorld speciesof the genus.s It is most closely allied. to
E. obligua .Llexander, of Macao,adifrering in the longer and n&rrower white crossbandof the wing and the slightly difrerent venation. The ovipositor has the genital segment bright fulvous, thevalves only a little duller in eolor. Tn E. obliqua, the genital segmentis black, the valves dark horn color.
ERIOCERA ARROGANS, new tpeclcr

Head black I antennae short I thorax dull blaek with three shiny
btackish gray stripes; legs dark brown; wings grayr the veins before
the cord broadly margined with rich yellowish brown, beyond thecord similarly margined with darker brown; cell M, present, d.eepI
abdomenblack, including the genital shield.
Fernale.-Length, about 12 mm. I wing, 11 mm.
Rostrum and palpi black. Antennre brownish black, the outer
flagellar segmentssomewhat paler I antennreshort, if bent backward.
not attaining the wing root. Head dull black, the vertical tubercle
not conspicuous.
Mesonotnm intense dull black, the praescutum with three broad
shin5' blackish gray stripes that are virtually confluent behind, restricting the ground color to the humeral triangles and broad lateral
margins I rernaindel of notnm of the same shiny blackish gray as
the praescutalstripes. Pleura shiny blacliish gray. Halteres black,the knobs broken. I-,,egsrvith the coxae and trochanters black; remaincler of legs dark brorvn, the femoral basesa trifle paler; legs
relatively short (fore legofemur, 5.6 mm.; tibia, 7 mm.1 tarsus about
5.8 nun.). Wings with the very restricted ground-color gray, thebase and costal region darker; all veins before the cord broadly
margined rvith lich yellowish brown, restricting the ground color tomedian streaks in the principal cells; beyond the cord the colorehangesabnrptly to clarlier brown, with similar pale centersto thecells; anal sells lnore extensively grayish; vein secand'A conspicuorrslv searnedwith brown I veins dalk brown, C and 8c paler brown.
\renation (fig. 3) : Ec, ending opposite the fork of Er*r*n,lSc, just
3 A n n . f f r r g . N n t . Hist., ser. 9, vol. 8, 1921.
l Proc. Ilawaiian Ent. Soc.,vol. 5, pp. 255-256, 1923.

PROCEEDINGS OF TEE NATIONAL

MUSEUI\Ivot^ 72

beyondthe fork of Rs; l?s angulatedand weakly spurred at origin;
Rz*s*na little longer than Rr*") l?r*, twice the length of R, or a little
morei Br," about one-half longer than Eri En deflecteclrather conspicuouslytoward the wing-tip at outer end; cells .8, and /?n with
inner ends acute; cell .i|y',present, relatively deep, nearly twice its
petiole ; n>ctl beyond one-third the length of cell frst M r, il little
shorter than the distal sectionof Cur.
Abclomendull black, including the genital shieldI valves of theovipositor horn colored;tergal valvesslender,gently upcurvedto theacute tips.
Ilabi,tat.-China (Szechuen). Holotype, female,Mount Omei (D.
C. Graham).
Type.-Female Cat. No. 4032L,U.S.N.M.
A very distinct speciesof the genus,without any very closeallies
among the describedspecies.
NEARCTIC

LIMONIA

SPECIESNITIDIUSCIILI\,

new specier

Antennae dark brown throughout, the flagellar segments withelongate verticils; last flagellar segment very long, nearly as longas the eombined twelfth and thirteenth segments; praescutum obseureyellow, with a broad dark brown median stripe I femora brown,
the tips narrowly yellowish; wings with a strong brownish tinge, thestigma darker, with a paler spot before and beyond it I male hypopygium with the dorsal dististyle a powerful chitinized rod, the apex
with a flattened black plate that terminates in a cornb of about nine
blunt teeth.
Male.-Length about 5 mm.1 wing, 6.5 mm.
Rostrum and palpi black. Antennae dark brown throughoutI
basal flagellar segmentsshorter, gradually increasingin length anddecreasingin diameter outwardly, the segmentswith a short apicai
and similar basalglabrousportion, forming a short pedicel; all segments with verticils of unusual length, much exceedingthe segments,
except those of the unusually long terminal segment; this segment
very long and slender, only a little shorter than the two preceding
segmentstaken together. Ifead dark colored,the anterior vertex moresilvery.
Mesonotum obscure yellow, the praescutum with u broacl clarkbrown median stripe and lessdistinct lateral stripes,the lateral margins and humeral region shiny yellow I scutal lobes yellorv, extensively marked with brown I scutellum and postnotum largely dark.
Pleura obscureshiny yellow with a very broad, conspicuous,longitudinal brown stripe and a less distinct brown mark on the sternopleurite, the remainder of the sternum of the ground color. Halteres

enT.2NDW UNDESCRIBBD CRANE FLIES-ALEXANDER,

7

with the unusually long lmobs dark brown, the baseof the stem narrowly paler. Legs with the coxaeand trochantersobscureyellow I
femora brown, paler basallS',the color deepeningoutwardly, the tips
tight yellow; tibiae and tarsi dark brown.
narrowly but conspicuously
Wings with a strong brownish tinge, the stigma oval, darker brown I
a paler spot before and beyond the stigma I veins darker brorvn, theobliterative areasat end of ^Rsand acrosscell f,rst M, conspicuous;vein .Rn*odark and equally developedthroughout its length. Venation: ,Sc' ending about oppositeone-third to one-fourth the length ofRs, Se, at its tip; l?s gently arcuated; basal sectionof Rn*, strongly
arctrated; eell f,rst M " relatively wide, only about one-half longer
than wide i nFal at the fork of M.
Abdominal tergites dark brown, the basal sternitesobscurebrownish yellow, the basesof the segmentsdarker; segmentseight andnine abruptly paler yellow I remainder of hypopygium dark. Male
hypopygium (fig. a) with the ninth tergite (t) gently emarginate
posteriorly,the lateral portions provided with a few very long setae.
Basistyle (D) relatively elongate,the mesallobe large, denselysetiferous. Dorsal dististyle a powerful, nearly straight, chitinized rod,at the apex with a blaekened flattened plate that terminates in a
comb of about nine blunt teeth. Ventral dististyle (d) small, fleshy,
denselysetiferous,the rostlal prolongation a long, sinuousrod, atnear midlength bearing two long black spinesfrom elongateswollen
papillae. Gonapophyses(g) with the mesal apical angle greatly
produced into a long flattened blade, the apex very shallowly bifid.
Habi.tat.-Oregon. Holotype, Marshfield (J. M. Aldrich).
Type.-Male. Cat. No. 40322,U.S.N.M.
The present speciesappears to be closely allied to L. adiecta(Doane), but I can not reconeilethe descriptionsof the two. The
present specieshas the anterior vertex silverv, the praescutumwitha broad median stripe and the rvings strongly infumed, with pale&re&sbefore and beyond the stigma. Vein .8, doesnot behaveat all
as describedand figured by Doane for acllecta,but bends at a rightangle into the costa, immediately opposite 8r. The very elongate
last segmentof the antenna,the long flagellar verticils, the conspicuous yellow terminal ring of the femora and the very peculiar structure of the hypopygium are all noteworthy features of the present
species'

DTcRANoMyH 'ENT.ILLATA, new rpecler

Related to D. lwpreti,ca Osten Saclien, from which it is distinguished especiallyby the structure of the male hypopygium; dorsal
dististyle slender, sickle shapedI base of ventral dististyle on inner
pencils of long yellow
face with a low lobe bearing two eonspicuous
setae.
Male.-Length about 6.2 mm.1 wing, 6.8 mm.

PROCEEDINGS OF TEE NATIONAL

MUSDUM

votu 72

Rostrum and palpi dark brown. Antennae brownish black
throughout I flagellar segmentsoval. Head dark.
Pronotum obscure yellow, with a, narrow median brown line.
l,fesonotal praescutum with three brown stripes, the ground color
well concealed.by yellowish gray pollen I scutal lobes dark, themedian area obscure yellow; scutellum obscure yellow I postnotal
mediotergite dark brown, sparselypruinose, the lateral margins narrowlv paler. Color of the pleura obscuredby glue but apparently
largely pale. Halteres short, pale, the knobs infuscated. Legs withthe coxae and trochanters pale brown, the latter more yellowish;
femora obscureyellow, the tips infuscated,especiallythe fore femora I
tibiae brownish yellow, the tips becoming darker I tarsi passing into
dark brown. Wings with a strong yellowish tinge, the stigma oval,
very slightly darker than the ground color I veins dark brown. Vena'
tion: Ec short,8c, ending just beyond the origin of Bs;8c, notvisible in the type, due to the flexing of the wings at this point, but
presumably not far removed from the tip of ,Sc' as in the related
Itnereti,ca;Bs about one and one-half times as long as the basal sectionof En*r; mrca before the fork of M.
Abdominal tergites dark brown, the sternites yellow. Male
hypopygium (fig. f ) relatively large and complicated in structure.
Basistyle (b) darkened outwardly, the ventro-mesal lobe elongate,
the apex truncated and provided with setiferoustubercles,with a few
other setae on the mesal edge of the lobeI mesal face of basistylenear apex with a low squat tubercle densely set with setae. Dorsal
dististyle (d) il slender curved hook, the apex acute. Yentral
dististyle (d) large, widened distally, sparsely provided with longcoarse setaeI rostral prolongation stout, the two spines relatively
short, separated from one another by a distance about equal to onehalf the length of one, placed more than their own length from theapex of the prolongation; basad and ventrad of the rostrum a conspicuous low lobe, each outer angle of which bears a conspit:uous
pencil of Iong yellow setae. Gonapophyses(g) with the mesal apical
prolongation slender. Aedeagus (") subtendedon either side by aflattened, shell-like apophysis.
Dakota. Eol'otype, male, Minot, July 16, I92L
IIabitat.-North(H. G. Dyar).
Type.-Male, Cat. No. 40323,U.S.N.M.
DICRANOMYIANEGLIGENS,

new species

General coloration gray, the praescutumwith three brown stripes,
the median stripe narrowly split by u capillary pale line; halteres
pale yellow, the knobs dark brown; femora yellow, the tips broadly
dark brown, precededby u slightly n&rrower ring of clearer yellow;

rnr.2N]IWUNDIISCRIBED CRANE FLIES-^I,EXANDER

9rvings whitish subhyaline, with four conspicrionsbrorvn costal aleasancl sparsemarkings elsewhereon the rving.
,Fenrule.-Length, 6.5-7.5mrn.; n'ins, 8.2-10mnl.
Rostrum ancl palpi blacli. -t\ntennaeirlacli. the basal half of thefirst flagellar segment yelloiv; fiageii:rl sey,lments
olal., becoming
srnaliei' outn'nrdly. I-Iead glay, thc center of the veltex nru'lied'rvith blaclr.
nfesonotal praescutum gray with three conspicuousbrown stripes,
the median stripe nau'orvly split lry a capillary palc line, thecephzrlicend of tlte stripe entile I scutum gra]', the lobes variegated'n'ith brown scutellum grav postnotum brown, sparsely pruinose.
I
I
Pleula gray. Halteres relatively short, pale yellow, the knobs darkbrown. Legs with the coxae brorvn, darker basally I trochanters
yellow I femora yellow, a little darker outwarclly, the tips broadly
dark brorvn, prececledby u slightly narrower clearer yellorv ring;tibiae ligirt brorvn, the extreme basesand the siightly broader tips
infuseated; basitarsi light blorvn, passing into darker brorvn, theremaining tarsal segmentsdark brown. \\rings whitish to whitish
subh;raline, sparsely variegated with brown, including four larger
costal blotches,the secondat the supernumerary crossvein in cell 8c,
the third at the tip of ,Sc, and origin of Rs, thc fourth the stigmal
area, barely confluent with a circular spot at the fork of Es; cord.
and outer end of cell first tr[, narrowly seamedwith darker brownl
paler gray marginal clouds on vein M" and all veins beyond, largest
and most conspicuouson the anal veins; smaller but darker m&rginal spots at ends of veins 8, and Rn*uia few scatteredbrown dots
in cells ilr, Cu, ancl the anal cells, not more than' four or five in asingle cell, none in the radial field; veins clar.kbrown, C, Sc, and l?
more yellou'ish in the interspaces. Venation: rSd relatively short,Ec, ending about opposite one-fourth the length of .8.s,Ec" Iacking,unless consideredas being lepresenteclby the so-called supernumerary cross vein at midlength of cell 8t; .Bs nearly straight; cellfirst M" gently widened outwardly, longer than vein M" beyond
itl tn-an before the fork of M, approximately equal to or shorter
tlran the clistal section of Cur. One paratype has an adr.entitions
cross vein in cell /?, of both wings.
Abdominal tergites brorvnish yellow, r'ariegateclwith clark brown.
rn other speeimensthe segments are dark brown, narrowlt' mareined cauclally rvith obscure yellorv. Ovipositor with the [asaihalf of the genital segment dark, the apical half obscure yellolv.
Valves horn color, the tergal.vatvcs relatively small, slender,genfly
upcurvecl,the tips acute.
Habito.f.-Colorado. EIolotypa, female, TennesseePass, July 24,
19LT (J. lf. ,\ldrich). Paratopotypes,2 females.
Type.-Female, Cat. No. 40324,U.S.N.I{.
55413_27_2

1tl

Ptioililr'lDl}]GS

iill

Tl:tIq l{--\'|IONAL

I\IUSEUM

vou.72

is most ciosely allied to D. nelliana
11sr1l'ir1t'ns
Dicrct,otttyia-i'Uos'!se'fhe rving-ptrttclir is much{r'orn Colorado.
Al.:xaut[,.i.,trixli.e re:;trictecl in the present species anrl Sc is slightty longer'
to urembelsof tlie sint,.;lotw
L:ear.a gleat resemblt,nee
Jir iir s,pet.ies
group crf the genusL'itnoruta
DICRANOPTYCHAOCCIDENTALIS,

new species

Gcner.aily similtrr to .D. sobt^itiaOsten Saehen; costal fringe ofmaie shor.t; mtle hyglopygiumrvith the ternrinal spine of the outerdististyie lons trncl straight.
trIcie.-l',trngth about 8 mm.1 wing, 9 mm.
Rostrurn darlr grey: the palpi brownish black. Antennae withthe scapal segmentsyellow, the flagellum drrk brorvn, with long
verticils. Head brorvnisb gray.
I\{esonoturnclark Sray: the praescutuin with a sc&rcelyindicatedd.arker gray meclian area, the surface of the clislc u'ith a sllarse
yellow pollen, thc lateral margins clearer gray. Pleura gl'ay'
tlre linobs rveakly infuscateci. Legs with the coxaei{aiteres pale,^yeilow:trochanters yellow I remainder of legs brownish
brownishyellow, the tips of the femora and tibiae & tittle clarkenedI terminal
iarsal ruguots clark brorvn. lVings with the costal fringe (male)
short, the membrane stronglv sufruseclwith yellowish; Rs ancl cellfirst M, relatively short.
Abdomen generally dark brorvn, the subterminal segrnentssiolnewhat rlarkei. I'Iale hypopygium about as in D. guaclii'u'itttta'
Alexander. but the outei ciististyle longer ancl mol'e slencler.withthe tern:inal spine lonq ancl straight, the spines on the_outer faceof the style rrbr. erect,ancl conspicuotts. Inner dististyie with thetip broa,il;t ro.,trclecl. Aecleagusblunt, not tenttinating in trvo rocls,
th; sribtcnciing plates less broadly flattenecl apir:alh". f-aterul
slender.eachterminating in a rnorc ehitinizerll:cali.rvitirt-iut
processes
apical cltnticles.
Ifabitaf.-Crrlifornia. IIotot.tlpe,rntle, Aipine" April 10, 1915 (M'C. \tarr l)lzee ). PtrutyTtes,a bloiien tnttle. lluir \Yoocls,IIarin Co.,
IIny 19. 1915 (l'I. C. Van Du:zce)I a broken tnale BerkeleyrNlav 14,
fgf.; (l,L C. Yan Duzee). The type is preserveclin the vriter''s collection.through ihe kindnessof I'{r. }I. C. Van Duzee.
Tirs sDeciesmentioneclaboveas l)'icrctnoptllr:ltaq'urtdri'ultirfto'\iex'
6 as a variety of sobrina Osten Sacken,
ancler *o. first clescribecl
but is nolv linoii'n to r.reamplv clistinct. It has the follor-;itlg clistribution. as lrnori'lt:
Peacef6i Vn]}ey. Al6lttst 2ir. 1917 (Coelierell): type.
Cirlori-..;'[r:I Proc. r\i:iltl. Nat. sci. I'hiiadelphia,
6('an. Ent., r'nl. !'r1, p. 191, 1910.

1914, pp. 579-5ti0, pl. 27, fig. 22.

,Ienr. 2NI'\Y

UNDDSCI.:IBED CRANE FLTES-ALDXANDER

11

Iclaho: Sandpoint,July ;], 191?(H. G. Dyar) ; Lake Pend Oreille,
July 4,,L9LT(I{. G. D1'ar).
},lontana: Glacier Park Station, altitucle4.800feet, July 24r (J. ilI.
Aldrich).
Alberta: ISanil,July 10.,1918(H. G. I)yar).
The speciesis told bv the four usually distinct plaescutalstripes.
The male hlzpopy$iumhas the outer dististyle unusually short andon the distal half, the trpicalspine
stout.strongiy arcuated,blackenecl. relatively short,both the upDerand lorverfaeesof the style roughened
but rrrorc especirill.ythe clutel ol conyex faee rvhere the sprnes arelarge ancl conspicuousbut strongly appressetl. fnner dististylc shortand stout, the apex bhint. Aedeagus terminating in trvo sruall parallei rods. subtenileclon either side by \:erv broad flattenecl plates.
Lateial processslender, carinate, the apex rvith micloscopic clenticles
aud roughenings.
LIMNOPIIILA,l

I(EPHDLIA)

ALDRfCIIf,

new specics

Gcneral coloration clarlr bron'nish gray, the praeseutttm lvith fottr'
sligh',ly clarker brorvn stripesI haltcres light yelloir; fetnora andtibiae yellowish, the tips darkenecl: wings subhyalinc, rvith a heavy
brorvn pattern that is restrictecl to the vicinity of the veins I malehypop;'gium with the outer clististyle bearing a smail triangtilar'
wing on outer margin at near tn'o-thirds the length.
Mala.-I'ength about 5.8-6.4mm. I wing, 7.5-I.8 mnr.
Fe'nml.e.-Length about 7 mm.1 l'ing, 8.5 mm.
Ilostrum and palpi dark. Antennae with the seapal segments
dark; basal segments of the flage'llum vellow. the outer segments
passing into brorvn. Head brownish gray.
Mesonotal praescutnm dark brorvnish gray with four slightly
darker brown stlipes, the interrnecliatepair narrowly separateclfrom
one another I p:eudo-*utural foveae conspicuous, subcireular, black I
remainder of mesonotum clalk, heavily pnrinose. Pleura dark bro'wn,
scarcely variegated n'ith brishter. Ifalter:es light yellorv. I-,egs
with the coxae brownish yellow, the midclle coxae dalker brown I
trochaniel's obscure vellow I femora yeilcn', the tips rather broaclly
but not abruptly infuscatedI tibiae yellow, the tips narro'n'ly infuseatedI tarsal segments obscure yellorv. their tips narrowl-t' clarkenecl, the terminai tarsal segments uniformly darkenecl. lYings
subhyaiine, with a heavl, brorvn pattei'n, as in the group I a seriesof sevencostal marks, the thild at the origin of -F,s,tire fifth, largest,at the stigma, the sixth ancl seventh at the enclsof vcins fr, and Bn;con:;pir:uousbrolrn seams along thc corrl ancl outcr encl of cell fi'.sf
M ri il broacl s:elrr ol1 the sullcrnllmerulv clossr-ein in cell M ; aclorrciat the folli of Xf r*,1 marginal clouclson nll the longitudinalr.einsearrciadof Il^,r, becoming iarger posteriorly. that at the sccond

12

pRocEEDrNcs oF THE N.lrroNAL MUsEUrlr

vor,.72

anal vein large, subcir:cular;that of the first anal vein very small lveinsyellorv,brorrn in the infuscatedat'eas.\ienation: Supernumeru"y .io.svein in cell M lying just bevond the level of the encl ofso the marlis are not in alignment.
vein secctnd,./-,
Abdominal tergites dark brou'nish bJack, the basesof the segments broaclly paler. I\fale hypopygiurn with the outcr dististyle(fig. O) relatively short and broad, the onter margin at near trvothirds the length with a small triangular flattenecl wing that is
microscopicallytoothed; the end of the style terminatesin a slenderchitinized pointl before this point, on the outer margin with numcrspinesI inner or lon'er margin of the style in theous subappressed
angle of the apical spine microscopicall;' denticulate.
Habi,tat.-Alberta, Montana. Ilolotype, mule, Glacier Park Station, I\fontana, altitude 4,800feet, July 24 (J. M. Aldrich). Allotype, female, Banff, Alberta, July 7, L922 (C. B. Garrc'tt) - Paratypes, two males with allotype, July 28-30, L922. Allotype in the
Canadian National Collection.
Type.-Male, Cat. No. 40325'U.S.N.M.
This interesting Li,mnophi'la is named in honor of Dr. J. M.
Aldrich, to whom I am greatly indebted for very many kindnesses
in the past.
LIMNOPUIL.I\(PEYIIDOREA)

COLUMBIANA,

new rpccies

Male.-Length about 9 mm.1 wing, 9.5 mm.
Closely allied to L. (P.) adusla Osten Sacken,differing especiallyin the coloration and strttcture of the male hypopS'gium.
Rostrum and palpi dark brorvn. Antennae with the basal segrnent dark brown, the succeeclingthree segmentsobscure brownish
vellorv, the remainder of the flagellum passing into brown. Headdark, grayish pruinose.
Pronotum and mesonotnmclark brorvn, the sides of the sclerites
paler, the surface with a sparse pollenI scutellum atld postnotum
paler with a narrow dark bron'n median line. Pleura brown, with
iI distinct microscopic pruinosity. Halteres pale, the knobs dark
brorvn. Legs rvith the coxae pale brown, sparsely pollinoseI trochanters obscure yellow I femora obscure yellow, the tips rather
narrowly ancl vaguely infuscatedI tibiae obscure veliow. the tips
vaguely darkenedI tarsi pale brown, the tips of the inclividual -.egments narrowly darker I terminal segmentuniformlv chlk. lYings
subhvaline,the eostal region slightly more vellowish: -.tigrna oval,
dark brown I cord narro.wly and lery intlistinctly seamecl withdarker; wing apex not darlienedI veins cla,rkbrorvu. Venation:
Ec, much longer, ending^Sc,ending shortly before the folk of ./?.s.
just beyond this fork ; Rs angulated and short spurred at origin;tnr,2

NEW UNDESCBIBED CRANE FLIES-ATJEXANDER

13cell M, longer than its petiole. The right wing of the type showsan adventitiouscrossvein in the axil of ^Es.
Abdominal tergites dark brown, the sternites somervhat paler I
hypopygium with the basistylesdark brown, the dististyles yellow,
Male hypopygium of the general type of L. adusta. Outer dististyles appearing as flattened blades, the outer apical angle ofeach being further produced into a, long, gently curved., flrg..like point that lies along the longitudinal axis of the style. Innerdististyle broad based, the distal two-thirds strongly narrowed,
thencereducedin diameter to the tip, without a conspicuousshoulder
on the outer margin near midlength, as in ad,usta. Aedeagus andt'he subtending gonapophysesvery long and slend.er,as in the group.
E abi,tat-British Columbia.

Eolotypa.-Male, Prince Rupert, June L7, 1919 (H. G. Dyar).
Type.-Male, Cat. No. 40326,U. S. N. M.
LIUNOPUILA(PEYLIDOREA) MICROPEALLUS, new rpecier

General eoloration shiny ferruginous, the abdomen of the malewithout a dark subterminal ring I femora almost uniformly yellow,
the tips very vaguely darkenedI male hypopygium with the aedeagus
and subtending apophysessmall and slender.
Male.-Length about 8 mm.1 wing, 9.5 mm.
Rostrum and palpi brown. Antennae with the basal segmentdark
brown, the secondsegmentbrorvnish yellow; basal segmentsof theflagellurn yellow, the terminal segments slightly more infuscated.
Ifead dark, presumably gray pruinose in fresh specimens.
Thorax shiny ferruginous, without rnarkings, the surface very
sparsely pruinose. Halteres pale, the knobs slightly infuscated.
Legs with the coxaeand troehantersleddish ferruginousI remainderof legs yellow, the tips of the fernora ancl tibiae very insensibly
darkenedI terminal tarsal segmcnts darlr brorvn. Wings with ayellowish tinge, the stigma elongateoval, dark brorvn; distal threefourths of cell C infumed I rving apex distinctly darkened; narrowand ill-defincdseamsalong the cord and vein Cur; veins brown, palerin the costal legion. Venation: Cell Il[, about equal to its petiole;cell first i11"elongate.
Abdomen yellorvish ferruginous, without darker markings. Male
hypopygium of the same general structure as L. fumidi,costa Alexander. Outer dististyle bloadly flattened,the apical portion suddenly
naruowedand blackened,the tip weakly bifid. Inner dististyle withthe curvedapical portion short. Bifid gonapophyses
with the lateralspine long and slender, acute, the a.xial spine a little l<;nser but
slender, the blackenedtip acutely pointed. Aedeagus and subtend-

L4

PROCEEDINGS OF TEENATIONAI,MUSEUM

VOtz.72

ing apophysesrelatively short and very slender,the former a little
the longer.
Habi,tat-lVlontana.
(H. G.Dyar).
Hol,ot'y'pe.-Male,Big Timber, July L4rLg1-T
Type.-Male, Cat. No. 40327,U.S.N.M.
LIMNOPEIII\

NICROFEMORATA, new rpecier

Male.-Length about 10.5mm.1 wing, 10.8mm.
Closely allied to L. mcdunnoughi,Alexander (Alberta), difiering
as follows:
Tuberculatepits appearing as barely evident linear impressionsat
the cephalicmargin of the praescutum,one eachon the inner marginof the two intermediatepraescutalstripes. Dorso-pleuralmembrane
brown,buffy surroundingthe spiracles. Legs with the eoxaesearcely
paler apicalty; fore femorablack,with aboutthe basalfourth yellow;
middle femora blaek, with approximately the basal third yellow;
fore and middle tibiae almost
posterior femora as in mcd,unnouglui,'black, the posterior tibiae paler. lYings with Rs angulated.and
spurred at origin.
Abdominal tergites dark brown, variegated with obscureyellow
immediately beyooa tn. clarh basal ring and transverseimpressionsof eachsegment;a blackishsubterminalring; hypopygiumbrownish
yellow. Male hypopygiumwith the basistyle(fig. ?) relatively stout,
the dorsal lobe short and broad, with conspicuoussetiferoustubercles.
Outer dististyle flattened but not as broadly so as in nrcd,wr'nough'i,,the distal half narrowed into a blackenedapex that terminatesin
teeth,the subapicalone shorter and straighter; surtwo conspicuous
face of style with abundant microscopicsetae. Inner dististyle averv small, stout fleshy lobe. Gonapophysesand aedeagusrelatively small and ineonspicuous.
Habitat -Montana. H olotype, male, Summit Station, altitude
5,200feet, July 25 (J. I\{. Aldrich).
T.t1pe.-\fale,Cat. No. 40328,U.S.N.M.
li- f . mcdunnouglui,,the gray of the mesonotumis duller anddarker. The fore femur has the darkenedapex of about the same
degree as in the other legs, but when specimensare fully colored
there is a d.istinctdarkenedring beyondthe midlength of the segment, followed by an ill-defined dull yellow annulus. Abdomen
uniformly darkened. Male hypopygium with the outer dististyle
evenbroader than in n'i,grofemnrata,greatly flattened; gonapophyses
more elongate,much exceedingthe aedeagus. L. mndunnpughi'is
still known only from Alberta (Nordegg, July L4, L92L,Mc DunnoughI Banfr, July L8-25,L922,Garrett).

ABT.2

NEW UNDESCRIBED CRANE IILIES-ALEXANDDR

15ULOMORPIIA ARIDEII\, new rpecier

Head dark, pruinose; mesonotumobscrlrebrown with three shiny
black stripesI scutal lobessimilarly blackened;wings fulvous-yellow,
the stigma barely indicated; cell M,. presentl abdorninal tergites
brown, the sternites obscureyeliow.
F'emale.-Length, L4 mm.; wing, L2 mm.
Rostrum obscureyellow, the palpi dark brown. Antennae very
long and slender (for this sex), if bent backward extending to beyoni
the wing rootl basal segmentsbrown, paler ventrally; flagellar segments beyond the secouduniform brownish black; flagellar segments
cylindrical, becomingvery elongatecylinclrictrlon the outer segments,
with the limits betweensegnents difficult to determine; all flagellar
segments with yery long verticils, as in the genus. Heacl clark,heavily gray pruinose, brighter adjoining the eyes, the anteriorvertex and front more yellowish.
Pronotum dark brown, yellow laterally. Mesonotum obscurebrown, the praescutum with three shiny black stripes that are virtually confluent,the interspacesbeing a trifle paler, sparselypruinoseI
median stripe more intensely black than the lateral stripesf humeral
yery small brighter yellow spot
leqion more yellowish, including abehind the pseuclosuturalfoveaeI scutum reddish brown, the lobes
largely shiny blnck, the posterior lateral regions obscure yellow;
scutellum and postnotal mediotergite black, the latter dulled basally,
the extremelateral margins of the cephalichalf yellow; pleura mostiyyellow, indistinctly variegated with reddish brown ott [he anepisternum and sternopleurite, producing ill-defined longitudinal stripesI
dorso-pleural region more yellowish. Halteres brown, the extreme
base of the stem narrowly yellowish. Legs with the coxae andtrochanters brownish yellow I femora brorvn, the basesyellow, thetips insensibly
passinginto clarkcrbrorvn; tibiae brorvn,the tips nAr'brorvn
rowly darker
I tarsi darh brorvn. \Yings with a strong fulvous-vellowtinge, the stigma barely evidentI veins brown. Macrotrichiae of cells abundant, continued basad beyond the level of theorigin of Rs, in cells C, R, M, and first /. considerablybasad of thislevel; cell ,Scdevoid of trichiae except at extreme outer end. Vena.tion; 8c,, ending shortly before the fork of 8s, 8c, at its tip; cell E,
sessilei Er*, nearly three times E, alonei Er*, and R, subequal; cell11, present, a little more than twice its petiole; cell f,rst Mrlarge,
pentagonal,m--cu not far from midlength.
Abdominal tergites brown, with a na,rrow darker dorso-medianline; sternites obscure yellow. Ovipositor with the tergal valvesvery long and slender, gently upcurved to the acute tips, horncolored, darker basally.

16

PROCEEDINGS OF TEE NATIONAI, MUSEUM

vot^ 72

Habi,tat.-Oregon. Holotypa, female,Marshfield, June 27 (J. M.
Aldrich).
Type.-Female, Cat. No. 40329,U.S.N.M.
(Jlomorpha ari,d,elais the seconclspeciesof the genus to be discoveredhaving ceILM, present. ft is readily told. from U. quinque'cellula Alexander by the large size ancl coloration of the mesonotum.
TRICYPEONA STENOPTERA, new specier

Antennae l5-segmented; legs long and slenderI wings reduced tolong ribbonlike strips.
Male.-Length about 6 mm.; wing, 3.2 mm., its greatest widtht
0.2 mm.
Rostrum obscure yellow, the palpi darker. Antennae brownish
black throughout, 15 segmented,the second scapal segment large,
as long as or longer than the first flagellar segment; flagellar segments beyond the first decreasingin length and diameter outwardly,
passing into short oval and then subglobular. Head light gray.
Eyes denselyhairy.
Pronotum bufry, a, Iittle infuscated medially. Mesonotum uniformly pale bufry yellow, without distinct markings. Pleura yellow, the anepisternum and sternopleurite a little more grayish.
Halteres pale yellow, the knobs infuscated. Legs unusually longand slender for such extreme stenoptery (fore leg, femur, 5 mm.;tibia, 5.4 mm.; tarsus, 7.9 mm.) ; cox&elong and slender, obscureyellow, sparsely pruinoseI fenora obscureyellow, the tips vaguelydarkened; tibiae light brown, the tips narrowly dalk brown I tarsi
dark brown. Wings reduced to long, ribbonlike strips, strongly
tinged with brownish yellow, clearer yellow basallyI veins pale, themacrotrichiae yery well preserved. Although the rving is so degenerate,the venation of the radial field is well preseryedand shows
frs arising at shortly beyond two-thirds the wing length, pale, without macrotrichiae; upper fork of the sector,Ilz*z*n,bears two long
branches, both with conspicuousmacrotrichiae for almost their
entire length; the basal sectionof E, is not preserved,there being noconnection between n, and the branches of the sectori Bu withmacrotrichiae for its entire length.
Abdominal tergites light yellowish brown with a vagruecllrker
brown dorso-medianvitta I sternitesmore uniformly eolored,redclish
brown, darker outwardly, the caudal margins of the segmentsnarrowly obscureyellow I subterminal segmentsuniformly darkenedI
hypopygium obscureyellow. Dlale hypopygium (fig. 8) with thetergite (t) greatly produced medially into a conspicuousbase that
divides further into two divergent setiferousarms, the setaeon thesurface erect, on the ventral surface more spinous and recurved.

enr.2

NEW UNDESCRIBED CBANE FLIES-ALEXANDEB

Lz

Basistyle (A) relatively large, the mesalmargin with a densepatchof short spines and longer setre,and a smaller group of very long
slendersetae. Outer dististyle (d) broadly flattened,dilated distally,
the outer angle a blunt point, the inner angle producedinto a slender
fleshy lobe that is set with small black spines, as common in thetribe. Inner dististyle longer and more slender,gently curved, thelower margin with small spinous setae. IVhat appear to be theinterbasesare broad, flattened curved.plates, narrowed distally, terminating in two powerful, unequal spines.
Habi,tat-Colorado. Eolotype,male, Tennessee
Pass,July 23,L9LI(J. M.Aldrich).
Type.-Male, Cat. No. 40330,U.S.N.M.
EXPL/ANATION

O['' PLATE(Svrasols: The venatlon used is the Comstock-Needham system, with the
eubital field as modified by Tillyard and the radial fleld as modifled by Alexander.-/.:Analveins ; C:Costa:' Cu:Qubitus ; J/:Media ; .R:Radius ; ,Sc:
Subcosta.
The terminology of the hypopygium is that of Cramton (1923).-4:ssfleagus;
D:basistyle ; d:dististyle ; g:gon'apophysls ; t:9th tergite.)Frc. 1. Britluura n'ymyth,ica,'wing.
2. Eriocera grahamri,' wing.
3. Eriocera amogans,' rving.
4. Limonia nitidiuseula,' male hypopygium.
5. Dicranomyio penicillato,' male hypopygium.
6. Limnoghi,lo (Ephetria) alil,rich,i; outer dististyle.
7. Limnoph,ilo nigrofemorata ; styli.
8. Tricgpluona ilewptero,' male hypopyglum.

cU . S . N A T I O N A LM U S E U M

PROCEEDTNGV
SO, L.72, ART. 2 PL. I

7