Reprinted from Bur,r,rrru oF THE Bnooxr,yN ExrouoLocrcAl, Socrnry,Vol. XXXfV, No. 2, pp. 92-100. April, 1939.

NEW OR INSUFFICIENTLY-KNOWN CRANE-FLIESFROM THE NEARCTIC REGION (TIPULIDAE,DIPTERA). PART V.
Bv CHeRLESP. ATExANDER,Amherst, Mass.*

The precedingpart under this title was publishedin r938 (Bull.
Brooklyn Ent. Soc. 33: Zr-78). Except where indicated to thecontrary, the types of the new_species
herein defined are preservedin my personal collection. I am greatly indebted to Messrs.
Brower, Hanson, Ide, Macnab, Procter and Ting for the privilegeof studying this material.
Tipula (Lunatipula) macnabi n. sp.
General coloration yellow, the praescutumwith four moreorange-brownstripes; antennaelong ; pleura yellow with asparsewhitish bloom; femora black, with an obscureyellow
subterminal ring, the tibiae and tarsi black; wings grayish
brown, the costal border, with the stigrna, darker brown ; aconspicuouswhite obliterative area before and across cord;
abdomenyellow, the tergite with three narrow brown stripes;
rnale hypopygium with the tergite produced into two strongly
divergent, subrnedian,black spines; inner dististyle with thebeak very slender, the dorsal crest conspicuouslyserrate;eighth sternite elongate,projecting,the apex with a fringe of
unusuallylong setae.
Male.-Length about r7-r8 mm.; wing l,6-17.5 mrlr.; antenna 6 mm.
Frontal prolongation of head obscure brownish yellow,
sparselypruinoseabove; nasusdistinct ; palpi dark. Antennae
relativelylong, as shownby the measurements; scapeand ped. icel light yellow, first flagellar segment brownish yellow, remainder of flagellum black; flagellar segments strongly incised; longestverticils subequalto the segments. Head gray,lighter on anterior portion and on the narrow orbits; a moreor lessdistinct darker medianvitta.
Praescutumgrayish yellow, with four more orange brolvn
stripesthat are scarcelyapparentagainstthe groundi posteriorsclerites of notum buffy-yellow; scutal lobes variegated byslightly darker areas. Pleura yellow, with a sparservhitish
bloom. Haltereswith stemyellow,knob weaklydarkened.Legsrvith coxaeyellow, sparselypruinose;trochantersyellow; fem* Contribution f rom the EntomologicalLaborator! t Massachusetts State College.

Aprdt,7s3eBu,lletin of the Broolilyn EntomologicalSociety 93
ora narrowly yellow basally,the remainderblack, with an obscureyellow subterminalring ; tibiae and tarsi black. Wings
with the ground-colorgrayish brown ; cells C and Sc, with thestigma darker brown ; a small brown spot at origin of Rs; aconspicuouswhitish obliterativearea before stigma and cord,crossingcell rst Mz into baseof cell Msi a less distinct poststigmal brightening; veins dark. Venation: Rs about two andone-half times the length of ntrcui R, long, nearly one-halfthe short R1; rn-cu a short distancebefore fork of M r*n.
Abdomen yellow, the tergites with a narrow brown medianvitta that is slightly interrupted at the sutures; less distinct
sublateralbrown lines, beginningon the posterior portion of
secondtergite; hypopygiumbrown. Male hypopygiumwith
the caudal margin of tergite produced into two black submedian spinesthat are strongly divergent, separatedby a Vshapednotch; ventral surfaceof tergite on either side of midline with a small blackenedpoint. Basistyleunproduced,butwith a detachedouter sclerite that is extended into a longsinuous spine. Outer dististyle at apex much enlargedintoa head. fnner dististyle a compressedblade, the beak very
long and slender,the high dorsal crest conspicuously
serrate;base of style produced into two small sclerotizedlobes, thenarrower one glabrous. Eighth sternite elongate,narrowed
outwardly, projecting, the slightly concaveapex on either side
bearing a fringe of unusually long setae,the submedianones
shorter; on ventral surfaceof sternitea deeplyforked mediansclerotizedplate.
Habitat; Oregon.
Holotype: 3, Boyer, September25, 1937 (Macnab). Paratopo-

types,r 3, April rS,rg37; r 3, July 30, 1936.

Ti,pula (Lunatipula) macnabais namedin honor of the collector,
ProfessorJamesA. Macnab. The fly is very different from othersimilar yellow species,especiallyin the colorationof the legs andwings, and in the structure of the male hypopygium. It rnost resemblesspeciessuch as T. splendens Doane and T. Iancellata
Doane,but is entirely distinct.
Tipula (Lunatipula) tingi n. sp.
Mesonotalpraescutumochreous,with four grayish stripesthat are margined with dark brown ; nasus lacking; antennae
with basalthree segmentsyellolv, the remainderdark; f emora
brownish yellow, restrictedlymore darkenedat tips; wings
with a strong brown tinge, the stigma darker; a broad, verv

94 Bulletin of the Brooklyn Entonr,ologicalSociety Vot.XXXIT
conspicuous,white band before cord, extending f rorn costa
into baseof cell Msi abdominaltergitesyellow, the outer segments more uniformly darkened,conspicuouslytrivittate withdarker brown, the rnedian vitta especiallybroad and conspicuous; ninth tergite with a very broad and deep V-shaped
notch; eighth sternite with conspicuouslateral lobes that are
tufted with setae,the medianarea with a brush of long setae.
Male.-Length about tGrT mm.; wing rGrT ffirr1.;antenna
about g mm.
Fenoale.-Lengthabout 24 mrn.; wing r9 mm.
t'rontal prolongation
Frontalprolongationof head obscureyellow ; nasuslacking;palpi with basal segmentsobscureyellow, the two outer segments passing into black. Antennae with basal three segrnentsyellow; renrainingsegmentsweakly bicolored,the basal
enlargementsblack, the apex of stem paling to reddish brown;
outer segmentsmore unifornrly blackened;flagellar segments
moderately incised; verticils conspicuous,subequalin length
to the segments;terminal segmentvery reduced. Head withfront obscureyellow, posterior scleritesbrown, more grayish
on posterior orbits; vertex with a linear dusky median vitta.
Mesonotal praescutum ochreous, with f our gray stripesthat are narrowly bordered by darker brown, somewhatmore
conspicuousalong the mesal edgesof the intermediatestripes,
the anterior ends of the latter clearer gray ; scutum with
Iobes gray, the rnedianarea yellow ; scutellumyellow, sparsely
pruinose,with a capillary brown median vitta, the parascutellamore dusky; mediotergiteyellowish gray medially, more ashygray on posterior third, the basal lateral portions broadly inf uscated. Pleura yellowish gray, the mesepisternumsomewhat clearer gray. Halteres yellow, the knobs dark brown.
Legs with the coxaeand trochantersyellow ; femora brownish
yellow, restrictedlymore darkenedat tips; tibiae and basitarsi
obscureyellow, the tips narrowly darkened; outer tarsal segments uniformly black. Wings with a strong brown tinge,
the prearcular field and costal border clearer yellow ; stigma
and a spot at tip of Scz darker brown ; very restrictedand lessevident dark seamsat origin of Rs and on posterior cord ; avery conspicuouswhite obliterativeband before cord, extending f rom vein C to the basal f ourth of cell M, i no distinct
post-stigmal brightening; veins brown. Squama with fivestrong setae. Venation: Rr+z pr€sent;Rs about two and onehalf times as long as nt,- cui Mra+ fl€?rly as long as the basal
section of.M t.

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95

Abdominal tergiteswith the ground color yellow, cleareron
the more basalsegments,passinginto brown on the fifth andsucceedingtergites; three brown stripes,the medianone broadand conspicuous;lateral stripes beginning on posterior halfof secondtergite, widenedbehind,the extremelateral borders
gray ; sternitesbrownish yellow; hypopygiumpale brown. In
and virthe female, the median tergal stripe is conspicuous.
tually continuousbut the lateral pair is broken into spots on
segmentstwo to five, inclusive. Male hypopygium with thetergite transverse,with an unusually large V-shapednotch, thelateral lobes subacuteat tips. Outer dististyle spatulate on
distal two-thirds or more. fnner dististyle with apical beakdarkened, obtusely rounded; a conspicuouslobe on outermargin at base; outer edge of style with unusuallyabundant,
long, coarsesetae. Each gonapophysisterminating in an acutepoint, before apex with a conspicuouslateral arm. Eighth
iternite with conspicuouslateral lobes that are tuf ted withlong yellow setae;medianregion notched,with a brush of longconspicuous
setae.
Habitat; California.
Hototype: 3, Marin County, bred from larvae in decayingwood
and leaves,collected March Io, 1935, ernergedApril I I, 1935
(Ting). Attotopoty,pe,
9, Paratopoty\e,S,larva collectedMarchIo, 1935, emerged April 8, 1935. Types in collection of the
U.S.N.M
Tipula (Lunatipula) tingi is named in honor of the collector.
Mr. P. C. Ting. The speciesis allied to several others in thewesternUnited States,apparentlybeing closestto T. (L.) Iygropis
Alexander, of Santa Cruz Island. The structure of the malehypopygiumis distinctive.
Limonia (Limonia) pemetica n. sp.
Allied to shnulans; general coloration gray, the praescutumwith three conspicuous,dark brown stripes; femora obscureyellow, the tips blackened,precededby " clearer yellow ring ;wings subhyaline,spotted and dotted with brown, including
a seriesof about sevenspotsalong vein Cu; a singledark area
involving the tip of Sc and origin of Rs; Sc schort,Sc' ending
about oppositeone-fifth the length of Rs; nx-cu a short distance before fork of M ; male hypopygium with the rostralspines blackened,conspicuous,widely separated; just caudadof the rostral prolongation on face of ventral style a small

96 Bulletin of the Brooklyn EntomologicalSocietgvot.XXXIT
blackened lobe covered with appressedspines; rnesalapicallobe of gonapophysislong and slender.
Male.-Length about 7 mm.; wing 8 mm.
Rostrum light brown; palpi darker. Antennae brown, theflagellarincisuresa little paler. Head gray.
Pronotum brown, pruinose. Mesonotal praescutum graypruinose, with three conspicuousdark brown stripes, themedian one becoming obsolete before suture; scutal lobes
darkened,the median area gray; posterior scleritesof notum
brownish testaceous. Pleura pale, the surface gray pruinose.
Halteres pale, the knobs dark brown. Legs with the coxae
and trochanters pale; femora obscure yellow, the tips
blackened,precededby a clearer yellow ring ; tibiae obscureyellow, the tips narrowly and weakly darkened; tarsi pale
brown, the outer segmentsdarker. Wings relatively narrow,subhyaline,spottedand dotted with brown, including a seriesof four costal areas,the third involving both the fork of Sc
and the origin of Rs; stigmal area confluentwith a cloud atf ork of Rs ; a seriesof about sevenspotsalong vein Cu; cordand outer end of cell rst M2 narrowly seamedwith brown.
abundant brown dots in cells of wing; veins pale bro*rrj
darker in the clouded areas. Venation: Sc relatively short,sc' ending about oppositeone-fifth the length of Rs, sc, beyond this origin ; a supernulnerarycrossveinin cell Sc at near
two-thirds the length-as ; m-cu a short distancebefore f ork of.M .
cell rst M z abor:t long as vein M t *, beyondit, its inne, .rrd
arcuated.
Abdominal tergites dark brown, the basal sternites moreyellowish; hypopygium pale. Male hypopygium with thecaudal margin of tergite emarginate,the lateral lobes broadly
obtuse. Basistyle with ventrornesallobe rounded. Ventril
dististylelarger than the basistyle,the rostral prolongationrelatively slender,with two black spinesthat are widely separated,the outer shortly before tip of prolongation,the inner oblique,subbasalin position; on face of style, caudadof baseof prolongation, with a small curved darkened lobe covered withtnicroscopicappressedspines; vestiture of mesal face of styleconsisting of abundant delicate setae. Dorsal dististyle agently curved pale rod, the tip subobtuse. Gonapophyses
withmesal-apicallobe long and slender,gently curved.
Habitat: Maine.
H olotype: 3, Hunter's Beach, Mount Desert, September24,
r935 (Brower).

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Limonia (Linoonia) perneticais generallysimilar to L. (L.)
sirnulans(Walker), yet very different in the structureof the malehypopygium. It may well have been confusedin collectionswith
si,rnulans.It is similarly relatedto the Rocky Mountain L. (.L.)
nelliana(Alexander), which hasSc evenshorter,Sc, endingopposite the origin of Rs. The specificname,pernetica,is derivedf rom
the Indian nameof Mount DesertIsland.
Pedicia (Pedicia) procteriana n. sp.
Generalcolorationgral, the praescutumwith four brown
stripes,the intermediatepair darker than the laterals; halteres
pale throughout; femora brown, the tips somewhatdarker;
wings yellowishsubhyaline,with the usual dark pattern of thesubgenus;dark seamalongvein Cu variable,in the type reaching wing-margin,in the paratypeendingat level of cord; twoisolatedbrown spotsbeyondcord at outer end of cell rst Mr;
cord very oblique; cell M, short, subequalto its petiole; cell
rst M, relatively long, its length about twice the maximum
breadth; cell M n deep,its length nearly three times the width
at base; abdominaltergites gray, with a conspicuous,darker
gray area that is narrowly marginedwith brown, theseareas
only moderately narrowed behind; male hypopygium withventromesallobe of basistylerelatively short; ninth tergite
without modifiedgroups of setaeon basal-lateralportions of
dorsal face; lobe of mesalface of basistylefringed with longsetae.
Male.-Length about24 mm.; wing 22 mm.
Rostrum dark, pruinose; palpi black. Antennae rvith thescapeand pedicelblack; flagellumbroken. Head light gray;vertical tuberclesmall but distinct.
Mesonotalpraescutumgray, with four brown stripes, theintermediatepair separatedby a capillaryground vitta, darker
than the laterals; scutum gray, eachlobe with two dark areas;
medianregion of the transversesuture darker brown; scutellum more reddishgray; mediotergitegray, the posteriorhalf
more reddish. Pleura pale, sparselypruinose; dorsopleuralregion dark brown. Halteres pale throughout. Legs withthe coxae light gray; trochantersbrown; femora brown, thetips somewhatdarker; tibiae brown, the basesnarrowly, thetips more broadly, blackened;tarsi black. Wings yellowish
subhyaline,rvith the usual pattern of the subgenus; costalborder pale, especiallythe basalhalf of costalcell; dark seamalong vein Cu reachingmargin as a narrow seamalong distal

98 Bulletin of the Brooklyn Entomologica.lSociety Tot'XXXIT
sectionof Cur; two isolateddark spotsbeyond the sealnat' cord, one at m, the other at fork of.Xtlt*ai veins brownish yellow, darker in the cloudedareas. Trichia of veins of moderate length. In the paratype, the dark searnalong Cu ends
oppositenl-cxcor nearly so. Venation: Cord very oblique;cell Mr short, subequalto its petiole; cell rst M, relatively
long, its length about twice the maximum breadth; cell M 'deep,its length nearly three times the width at base.
Abdominal tergites light gray, each segmentwith a conspicuous,darker gray, brown marginedarea,narrowedbehind.
the width at apex about one-half the greatestdiameterof thearea; incisuresnarrowly pale; sternitesgray laterally, with anearly continuous median brown stripe; hypopygium dark.
Male hypopygium having ninth tergite without modified
portionsof dorsalsurface,asbrushesof setaeon basal-lateralin contern'tina;median area of tergite producedinto a depressedlobe, the caudal margin gently emarginate,subtended
on either side by " flattenedplate, the apex truncate. Basistyle with ventro-mesallobe of moderatelength, when viewed
from beneathnot extending caudadbeyond the level of the
dististyles,broad-based,
stout, the apex with abundantspinous
setae; viewed from above,the apex of this lobe is visible
oppositethe inner arm of dististyle; outer apical region ofbasistylewith a brush of long, dark-coloredsetae. Interbasal
rod relatively small, with a broader flattenedlobe immediately'
caudad,the apex of latter fringed with long pale setae. Dististyle trifid, the outer blademore expanded,more or lesscultriform; centrallobe more slender,pointedat apex and without
blackenedspines,as in conterntina;innermostlobe very low
and obtuse,denselyset with short blackenedspines.
Habitat; Ontario, Maine.
Holotypet 3, Corfield,l\Iount Desert,Maine, July, 1937 (Proc'
ter).
probablya female,Horning's Mills'
Paratype,abrokenspecimen,
It is presumedthat the adult
Ontario,June rz, r9z8 (F.P.Ide).
of the holotypeflew from one of the rocky mountainstreamsabove
Corfield,as perhapsDuck Brook.
I take unusualpleasurein naming this striking crane-flyin honorof my friend, Dr. William Procter, whose recentlypublishedlistof the insectsof Mount Desertmust long remain our chief sourceof information concerningthis fascinatingregion. The speciesis
yet is amply
most similar to Pedicia (Pedicia) conterncinaWalker,

Aprit,TeseBu,lletin of the Broolilyn EntomologicalSociety

99

distinct. The reduced ventro-mesallobe of the basistyle of thetnale hypopygium is more as in P. (P.) olbiztittaWalker, which inother regards is entirely different. In contenrcina,besides thecharacters above mentioned, the median lobe of the tergite isshorter and broader, with the margin more deeply emarginate,thesubtending plates reduced. Ventro-mesal lobe of basistyle very
iarge and greatly produced,extending caudad beyond the level ofthe dististyle in both dorsal and ventral views, the mesaledgewitha brush of blackenedsetae,additional to more normal yello'rvones.
Body of basistyleshort and stout, with a rounded lobe on dorsaltnesal aspect; mesal lobe caudad of interbasentore flattened,notconspicuouslytufted with setae.
Rhabdomastix (Sacandaga) hansoni n. sp.
Generalcoloration of thorax almost uniformly dark brown,
sparselypruinose; halterespale yellow ; legs brownish yellow ;rvings subhyaline,sparselypatternedwith brown, including
tlre stigmaand sealnson anteriqrcord and alongvein Cu; prearcular field more whitened;numerousmacrotrichiaon veins
beyondcord; Sc relativelyshort, Sc, ending shortly beyond
midlengthof Rs; R, slightly oblique; abdomenblack.
F enmle.-Lengthabout8 mm.; wing 6.5 mm.
Rostrum dark brown, pruinose; palpi black. Antennae
rvith the scapeand pedicelbrown, flagellumobscureyellow;
flagellarsegmentswith long verticils,the longestonessecund.
Head brownishgray.
Mesonotum almost unif ormly dark brown, the surface
sparsely dusted with gra!t thinner on median region ofpraescutunt,leaving the surfacesubnitidous;anterior lateralpretergites
paleyellow; pseudosutural
foveaedark; medianarea
of scutum a little paler. Pleura dark brownishgray. Halterespale yellow. Legs with the coxaepale,sparselypruinose; trochantersyellow; remainderof legspale brownishyel ,low. Wings subhyaline,sparselypatternedwith brown, includingthe stigmaand seamson anteriorcord and alongvein
Cu; prearcularfield more whitish; veinsbrown. Macrotrichia
abundanton veinsbeyondcord. Venation:Sc relativelyshort,
Scr endingjust beyondmidlengthof Rs, Sc2closeto its tip,
about oppositernidlengthi R, slightly oblique,about equal tothe distanceon costabetweentips of veinsRr*z and Rs; Rnlonger than Rr*s+ai cell rst Mz small.
Abdomen black, including the genital shield and valves ofovipositor.

100 Buttetin of the Broolilyn EntomologicolSociety Tot'XXXIT
H obitat; Massachusetts.
Holotype: ?, ParadiseTrail, Mount Toby, Franklin Co., altitude6oo feet, July 19, 1938 (Hanson).
Rhabdomasti,r(Socandaga)hansoniis namedin honor of thecollector,Mr. John Francis Hanson, student of the Plecoptera.
The speciesis very different from all those previouslydescribedfrorn northeasternNorth America, especiallyin the body coloration and sparselypatternedwings. It is closestto R. (.S.) subfasciger Alexander,of Alberta, differing in detailsof colorationand venation,as the shorterSc and more obliqueRr.
Ormosia broweri n. sp.
Relatedto arcuata,differing especiallyin the structureof themale hypopygium.
Male.-Length about5.2-5.5mm.; wing ffi.s mm.
Generalcolorationof body gray pruinose. Antennaeblack
throughout. Legs dark brown. Wings with a strong brownsuffusion,cell C and the stigma darker brown. Venation:
Veins R, and Rn slightlyupturnedat tips ; vein zrtd I strongly
sinuous.
Male hypopygiumwith the lateral apophysesof the phallosome entirely different from arcuato,appearingas relatively
straight, broad-basedrods, the distal f ourth narrowed anddeflectedlaterad into acute points. In arcuata,thesehorns
subtendthe central platesof the phallosome,each appearing
as a slenderspine, strongly arcuatedat midlength,the longstraight apicalpoint directedcaudadand slightly mesad. The
paratypesdiffer very slightly in having the lateral apophyses
appearingas straight,long-triangularpoints that are gradually'
narrowedto the acutetiPs.
Habitat; Maine
H olotype: 6, Mount Desert, near EntomologicalLaboratory,M"y r5, 1935(Brower). Paratyp€s,333, April 28, 1936;May
16, 1937.
I take great pleasurein namingthis interestingOrmosiain honorof Dr. A. E. Brower, whoseintensiveeffortshave addedvastly to
our knowledgeof the insectfauna of N'IountDesert.