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l!"ir":UNDESCRIBED SPECIES OF TIPULA FROM WESTERN NORTH AMERICA (DIPTERA,TIPULIDAE). PART Ir.
Bv CuaRLESP. ATBxANDER,
Amherst, Mass.*
The precedingpart under this generaltitle was publishedin 1945
(Btrll. Brooklyn Ent. Soc., XL: 33/37). At this tirne I am
consideringthree further speciesof the genus Tipula, all from thedesert section of u'estern Arizona and southern California where
they were collectedby my friends, Messrs.GeorgeF. Knowlton andJohn L. Sperry, to whom l am very much indebted for many appreciatedfavors. The types are preservedin my personalcollectionof theseflies.

Tipula (Lunatipula) boregoensis,n. sp.
Sizevery small (lving, male,8 mm.) ; mesonotalpraescutum
gray, with four entire brown stripes, the intermediate pair
separatedby a broad gray line ; antennaewith scapeand pedicel
yellow, flagellumblack ; flagellar segrnentssimple,without basal
swellings; femora and tibiae yellow, the tips narrowly blackened; clawsvery weakly toothed; wings with a weak brownish
tinge, stigma brown ; a very conspicuousobliterative band
before cord, virtually crossingthe wing along vein Ma,i malehypopygiurn having the posterior border of tergite very shallowly emarginate;outer dististyleu'idely dilated on outer part;inner dististyle with beak long-extended,the lower beak lacking ; a conspicnousblackenedtooth on posterior portion of
crest; outer basallobe narrowly attachedto body of style, appearing as a long narrow pale blade,its apex pointed and nearly
glabrous; eighth sternitenarrowed outwardly, the tip truncate,provided with two groups of relatively few long yellow setae,
without lateral lobesor projections.
Male.-Length about 7 n'nn.; wing 8 mm.
Frontal prolongation of head shorter than the remainder,
gray above,obscureyellorvon sidesand beneath;nasusstout;
palpi brownish black, the tip of the third segmentnarrowly
paler. Antennae rvith scapeand pedicel light yellow ; basal
two flagellar segnrents black, sirnple; remainder of organ
broken but evidently short. Head light gray, still paler onfront ; vertex with a very slightly darker median line that ismore expandedbeforemidlength.
* Contribution from the Entomological Laboratorl, Massachusetts State College.

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Societg vot' xLr
Buttetinof theBrooklynEntomological
Pronotal scutum gray,vaguelypatternedwith brown, scutellum yellow. Mesonotal praesctltum gray with four entire
especiallythebrown stripesthat are relatively inconspicuous,lateral pair ; intermediate stripes separatedby a broad grayline that is only a little narrower than the stripe; humeral region restrictedly yellow ; remainder of mesonotum gtaY, eachscutallobe with two brown marks; parascutellayellow; pleurotergite chiefly light yellow. Pleura light gfaY, the ventral
sternopleuriteshowing a darker color beneaththe pruinosity;
dorsopleuralregion yellow. Halteres with stem obscureyellow, knob infuscated. Legs rvith the coxae light gtay; trochantersyellow; femora and tibiae yellow, the tips narrowly
but conspicuouslyblackened,the amount subequalon all legs;
tarsi obscureyellow, passing into black; claws very weakly
toothed. Wings with a weak brownish tinge, slightly darker
beyond the cord ; stigma brown ; small dark spots at end of Sc
and over the anterior cord ; a very conspictlouswhitish obliterative band before cord, virtually crossingthe wing alongvein M+i poststigmalbrightening much more restricted andcells Sc, and R-,; veins brown, paler
inconspicuous,occupying-ofc-ord,especiallyin the prearcular andin the areas basad
costal fields., Squarnalsetaeconspicuous;numerousmacrotrichia on most veins beyondcord. Venation: Rs more than
twice as long as ux-clt,
i R, * , entire, its distal third without
trichia; nc and petiole of cell M, subequal; In-cLt on M, * n
in length toshortly beforefork ; basalsectionof M, + n SUbequalr-rrc; cell 2nd A rclatively narrow.
Abdominal tergites obscureyellow, more or less distinctly
trivittate with dark brown, most evidenton the outer segments,
the lateral seriesbroken into spots; central stripe more nearlycontinuous,interrupted by the yellow posteriorborders of thesegments; sternitesmore brou'nishyellow ; hypopygiumchiefly
Male hypopygium with the ninth tergite with they.llo*.
posterior border very shallowly emarginate,the lateral lobes
ihur formed very broad; at baseof notch with a microscopic
lobule; ventral face of lobes with an obtuse blackenedtooth
directedmesad. Ninth sternitewith its appendagesmall andinconspicuotts,subcylindrical,the basal portion more dilatedon the inner face, the whole appendage'rn'ithnumerous setae,
those of the expandedportion longer and more conspicuous.
Basistyle entirg its dorsal end much narrorved to pointed;
caudai margin oppositethe point of insertion of the dististyles

Aprit,1e46 Bulletin of the Brooklyn EntomologicalSociety 47
with a very small sclerotizedpoint. Outer dististyle with its
outer portion broadly expanded. f nner dististyle distinctive;
beaklong-extended,the lower beaklacking; dorsalcrest microscopicallytoothed,separatedfrom a more blackenedposterior
lobe or crest by " U-shapednotch; the entire face of style unusually glabrous, with small and inconspicuoussetae; outerbasal lobe attachedto body of style only by a narrow basal
connection,appearingas a long narrow pale blade, its apex
pointedand more nearly glabrous. Eighth sternitemoderately
sheathing,narrowed outwardly, the apex truncate, provided
with a triangular grorlp of about 18 to 20 long yellow setaeon either side of the midline ; no lateral lobes or other modifications; setaedirectedcaudad,not or scarcelydecussate.
Habitat; California.
H olotype: 3 , Borego Desert State Park, California Painted
Desert, San Diego Co., altitude about 450 feet, April 23, L944

g. L. Sperry).
This very distinct fly has no very close relativesknown to me.
The unusually simple eighth sternite of the male, with no laterallobes,somewhatsuggestsspeciesthat are allied to Tipula (Lu,natipeila) usitata Doane,which are entirely distinct flies. The species
more resemblesT. (L.) buenoi Alexander and allies which havethe lateral lobesof the eighth sternite well developed.
Tipula (Lunatiputa) mohavensis,n. sp.
brown,
Allied to buenoi,'generalcolorationlight testaceous
more or lesspruinose; pleurapatternedwith brown and yellow ;antennaewith basal three segmentsyellow, flagellum black;wings rvith a strong brownish gray tinge, the prearcular andcoastal regions even darker brownish yellow ; stigma pale
brown, small; obliterativeareasvery reduced; veins delicate;male hypopygiurnwith the tergite large, its caudalborder produced into four blackenedpoints, the outer pair appearingas
spines,the shorter inner pair more obtuse; outer dististyleconspicuouslyexpandedat apex; inner dististylewith the beakmassive,heavily blackened;dorsal crest producedbackward
into a conspicuousblade; outer basallobea large flattenedplate'margins
obtuse; eighth sternite with the lateral andwith allmedian lobes detachedfrom the body of the sclerite by pale
membrane; lateral lobes two-armed,the outer one a glabroussclerotizedhorn, the inner arm with severalstrong setaeon its
basalpart ; rnedianlobe transverse,with two large groups oflong crinkly setae.

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Tol. XLI

Male.-Length about 15 rnm.; wing 15.5 mm.; antenna
about4.7 mm.
Frontal prolongationof headobscureorangeyellow; nasus
distinct; palpi light brorvnor brownishyellow. Antennaewith
scape,pediceland most of first flagellar segmentyellow, theouter end of the latter more darkened; remainderof flagellurn
black,the segmentsmoderatelyincised. Head obscureyellow,
discolored;verticaltuberclevery lor,r'to scarcelyevident.
Pronotal scutum brorvn, rnore or less darkenedmedially;
scutellumand adjoining regionsof praescuturnbright yellow.
Mesonotumdiscolored,evidentlylight testaceous
brorvn,moreor less pruinose, the praescutumapparently with stripes.
Pleura chiefly brown, pruinose; dorsopleuralregion and extensiveareason the pleurotergiteand posteriorpleuritesbrightyellow,includinga very narrow transverseline extendingfrom
the propleura to behind the fore coxa. Halteres with stem
obscureyellow, knob darkeneclon basal portion, the apex
obscureyellow. Legs with the coxaeobscurebrownishyellow,
gray pruinose; trochanters)'ellorv;remainderof legs broken.
Wings with a strong brownishgray tinge, the prearcularandcostalregionsevendarkerbrownishyellow; stigmapalebrown,
small; obliterativeareasverv reduced,only the prestigmalonepresent, extending from before the stigma along the cord,barely entering cell M s i veins light brown, fflore yellowish
brown in the saturatedportions. Squamalsetaestrong andconspicuous;trichia of veins beyondcord numerousand welldistributed,lackingon distalthird of Rr * 2i v€insrnoredelicate
than in bttenoi. Venation: Rs abouttwice nx-cxt,R, * 2 entire;
petiole of. cell M, nearly twice rn; nrcu on M n immediately
beyondorigin.
Abdominal tergiteschieflyfulvous yellow, the posteriormargins narrowly, the lateralbordersmore broadlygrayish yellow;
sternitesand hypopygiumyellow. Male hypopygiumwith theninth tergite large,its caudalborder producedinto four blackenedpoints,the outermostpair appearingas strong blackened
spines,the inner pair as somewhatshorter blackenedknobs,
the variouspointsspacedaboutequidistantlyfrom oneanother;a conspicuousdorsal furrow. Ninth sternite with its appendagesmall and inconspicuous,provided with long palesetae. From the notch of the sternite,visiblein profile, juts asmall sclerotizedstructure that terminates in two divergent
spinouspoints. Basistyleentire, its dorsal portion produced

Aprit,1e46 Bulletin of the Brooklyn BntomologicalSociety 49
into a slenderarm; caudal margin unmodified. Outer dististyle expandedat apex into a flattened oblique head. fnner
dististylewith its beak massive,heavily blackened;lower beakmuch smaller, likewise heavily blackened; dorsal crest produced caudad into a conspicuousblade; outer basal lobe alarge flattenedplate, in the type slide bent backwardover the
body of the style and its exact outlinesdifficult to delimit but
the marginsevidentlyall obtuse;basalportion of lobe with anextensivegroup of long yellow setae; sensoryarea long-oval,
including about ten facets. Phallosomeconsisting of two
united plates,eachwith a strong recurvedreddishspineand alonger, more nearly straight outer blade. Eighth sternite
with three isolatedlobes,detachedfrom the rnain bo_dyof thescleriteby pale membrane;lateral lobestwo-arrned,the outerarm a glabrous sclerotizedhorn, the inner one with several
strong setaeon its basalportion; medianlobe pale,its cephalicmargin transverseand more thickened,on either side withlarge groups of setae that bend toward the midline but are
scarcely decussate,these setae conspicuouslyfirnbriate anci
tangledat their tips.
H abitat.'Arizona.
Holotype: 3, Kingman, Mohave Co., altitude 3340 ft., M"y 9,1945 (G. F . Knou,lton).
The most similar describedspeciesinclude Tiltula (Lunatipula)buenoiAlexanderand T. (L.) stolagm,ites
Alexander,which differ
very evidentlyin all detailsof structureof the rnalehypopygium.
Tipula (Lunatipula) macracantha, n. sp.
Belongsto the irnpudicogroup ; generalcoloration of mesonotum gray,includingthe praescutalstripes,the rnedianstripe
with brown borders; antennaewith scapeand pedicelyellow,
flagellum brownish black; flagellar segmentsstrongly incised;legs brown, tarsi blackened; wings rather conspicuouslypatterned with brownish gray and whitish subhyaline,the lattercolor includinga major prestigmalarea that reachesthe wing
tip in cell Rr ; abdorninaltergitesyellow with three dark brown
stripes, the lateral pair much broken, virtually restricted tolinear dashesnear the basesof the intermediatetergites; malehypopygiurnwith the tergal canthi appearingas long triangularblades,the tips acute; median lobe conspicuous,compressedalong tlpper margin ; lateral processesappearingas small, very
acutespinouspoints; outer dististylewith apex very expanded,50

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Vol. XLI

roughly triangular in outline; inner dististyle with both thebeak and lower beak heavily blackened,posterior crest small,outer basal lobe short and stout, its apex truncate; eighthsternite with a small dark-coloredlobule in the membraneof
the following sternite.
Male.-Length about 15 mm.; wing 13.5 mm.; antenna
about4.8 mm.
Frontal prolongationof head brown, pruinose; nasusvery
small; palpi brownish black. Antennae (male) moderately
long; scape and pedicel yellow, flagellum brownish black;flagellar segmentswith the basal swellings a trifle more intenselycoloredthan the sterns,strongly incised; longestverticils subequalin length to the segments. Head light gray, witha narrow but conspicuousblack median vitta extendingfrom
the low verticaltubercleonto the occiput.
Pronotal scutum brown, darker on lateral parts; scutellum
light yellow. Mesonotal praescutum with the interspaces
buffy, the stripesand lateral borders light gray, the latter moreyellowed on the humeral region; median stripe with conspicnousbrown borders that do not quite reach the suture;lateral stripes narrow, dark brown, only a little wider thantlre dark margins of the central stripe; scutum gray, eachlobe
weakly patterned with brown, median area more yellowed;
posteriorscleritesof notum obscureyellow, gray pruinose,witha continuousdark brown median line. Pleura light brown,
gray pruinose,vaguely patterttedwith brown. Halteres withstem testaceous,yellow at base,knob infuscated,restrictedly
more brightenedon dorsal edge. Legs with the coxae obscureyellow, sparselypruinose; trochantersyellow; remainder oflegsbrown, the tarsi passinginto black; claws (male) toothed.
Wings rather conspicuouslypatternedwith brownish gray andwhitish subhyaline,the latter color including the prearcularfield and basalcells; a major prestigmalarea,continuedto thewing tip in cell Rr ; poststigmalbrightening rnore restricted;
other pale areasin outer end of cell Rr, most of.M r, basesof
Znd M z, M, and M n, and as conspicuousborders to both Analveins, leaving conspicuousmarginal darkeningsin the centersof the Anal cells; cellsC and Sc a trifle more yellowedthan theremainder of ground ; somewhatdarker clouds in cells R, andR, and again in the outer end of cell M at nt-cw; veins brown,
paler in the prearcular and costal fields. Venation: Rs relatively long, about two and oue-halftitnes nx-cLc;R, * 2 entire;rn longer than petiole of cell M L.

Aprit,7e46 Bulletin of the Brooklyn EntomologicalSocietg 51
Abdominal tergites yellow, with three dark brown stripes,
the medianone rnorenearly entire, interruptedat the incisures.
particularly at the posterior borders of the segments; lateral
stripes much interrupted, appearingas a linear spot near thebaseof tergites three to seven,less evidently on tergite trvo;
lateral tergal borders broadly grayish yellow ; sternitesyellow,
the outer segmentsmore patterned, especiallymedially onsternites seven and eight; hypopygium yellow. Male hypopygium having the ninth tergite with the canthi unusuallyelongate,appearingas long-triangularcurved bladesthat narrow to acute points ; setae of margin very srnall and inconspicuous,thoseof the basaland lateral portions long and stout;
median lobe (subtergalprocess) only a little shorter than the
canthi, the dorsal edgecompressed-flattened,
the lower portion
more expanded; lateral processesappearing as small, very
acute spinouspoints. Ninth sternite having the margin provided with coarse setae, those nearest the midline arisingfrom conspicuorlsbasal tubercles; lobe of sternite small, suboval, with a smaller lateral lobule or flange, the main body ofthe lobe provided with coarseyellow setae. Basistylewith its
dorsal part narrowed into a slightly sclerotizedspinouspoint,
the ventral end also narrowed but more obtuse. Outer dististyle with the outer end very expanded,roughly triangular inoutline; setaeelongate,restrictedto outer half. Inner dististylewith both the beakand lower beak heavily blackened,the lattervery stout ; posteriorcrest small,its dorsal margin microscopically toothed; outer basal lobe short and stout, its apex truncated,on the proximal portion lvith the usual area of delicate
appressedsetae; sensoryarea comprisedof only about 8 or 9
facets. Eighth sternite with a small dark-colored medianlobule in the membraneof
the following sternite, this srnaller'structure
than the correspondingin ucitrata, provided withmicroscopicscabrouspoints; central portion of sternite rvith
severalrows of long setae,the lateral parts produced into lorv
lobesthat are provided with a few still longer bristles.
Habitat.' Arizona.
Holotype: 3, PeachSprings, Mohave Co., altitude 4790ft., May
8, 1945(G. F. Knozulton).
The only speciesof the impudico group with which the presentfly
requires comparisonare Tipula (Lunatipula) rmitrata Dietz and
T. (L.) u,tahicoloAlexander, both of which differ conspicuouslyin
the details of structure of the male hypopygium, particularly of thetergite and dististyles.