F r 4, j + t,D

Reprinted from Bur,r,ETrNoF THE BBooKLyN ENTotrror,ocrcAlSocrnrv,
VoI. LI, No. 3, pp. 75-81. June, 1956.

UNDESCRIBED SPECIES OF NEMATOCEROUS
DIPTERA.
PART IV.
By Crr.lnI.EsP. AlaxeNoen, Arnherst, Mass.
The preceding part under this title appearedin 1953 (Bull.
Brooklyn Ent. Soc., 48: 97-103). In the present report I am
describingthree new speciesof Ptvchopteridaeand one Blepharoceridaeand am providing further notes on the remarkableTrichocerid fly Kau,asentyiahnanishii,(Tokunaga).

PTYCHOPTERIDAE
Ptychoptera chalybeata n. sp.
Size small (wing of male 6 mm.) ; gen.t"l coloration of head andthorax black with bluish reflections; femora yellow, the tips narrowly brownish black, tibiae and basitarsiobscureyellow, the tips
narrowly darkened; wings whitish, the broad tip and a narrow
band at cord brown; abdomenyellow, the posterior borders of theintermediate segmentsblack, the subterminal segmentsuniformly
blackenedto form a ring.
MaIe: Length about 6 mm.; wing 6 mm.; antennaabout 3 mm.
Mouthparts, including palpi, light yellow. Front and clypeus
brownish black. Antennae moderatelylong; scapebrown, pedicel
and first flagellar segment obscure yellow, succeeding segments
dark brown, cylindrical, with short verticils. Head black, withbluish reflections.
Prothorax chiefly dark brown. Mesonotum black, with steel
blue reflections,most evident on the praescutum. Pleura black,

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vot'Ltthe dorsopleuralregion pale. Halteres with stem yellow, knob
broken. Legs with all coxae and trochanterspale yellow; feurora
yellow, the tips narrowly but conspicuouslybrownish black; tibiae
and basitarsi obscure yellow, the tips narrowly more darkened,remainderof tarsi black. Wings whitish, with an unusuallyheavy
brown pattern, including the apex basadto just beyond the levelof the forks, the wing tip a trifle paler; a brown band at cord, narrowed posteriorly,extendingto the bendof vein Cu; a small postarcular spot in basesof cellsR and M; cells C and Sc urore yellowed
than the ground ; veins brownish black. Venation: Rs very shortand straight, about twice the basal sectionof Ru; cell R, a trifle
deeperthan cellZndM u.
Abdornenchieflyyellow,the posteriorbordersof the intermediatesegmentsblack, subtertninalsegmentsuniformly black to form aring; hypopygium obscure yellow. Male hypopygium with thetergal lobesrelativelylong and narrow, their tips weakly expanded,on inner surface provided with strong retrorse setae. Dististylesrnall,pale, shallowlyand unequallybilobed at tip, the lower lobe
longer. Phallosomeheavily blackened.
Habitat: Thailand.
Holotype: {, Huey Kao, June 14, 1953 (Manop Ruttanapradith).
Typ. in my collection.
The only generallysimilar regionalspeciesis the larger Ptychoptera di'stinctaRrunetti,of the easternHimalayas. This has the wing
pattern and venationvery tnttch as in the presentfly, differing in
the coloration of the body and legs. The male sex of distincta
is still unknown.
Ptychoptera schoutedeni n. sp.
Mesonotum orange, the praescutumchiefly black, the cephalicfourth pale; posterior scleritesof notum black, the posterior twothirds of the mediotergitefulvous yellow; basal palpal segments
yellow; antennal scapefulvous; haltereswith stem fulvous, knob
infuscated;femora and tibiae obscureyellow, the tips weakly darkened; wings pale yellow, with a brown seamover cord, the wing
tip paler brown; vein R, oblique,basalsectionof Rr conspicuous;
male hypopygium with the elongatelobesof the tergite terminating
in slenderglabrousspines;dististyletrilobed, the basallobe small,
capitate.
Male; I-ength about9 mm. ; wing 8 mm. ; antennaabout 4.8 mm.
Rostrum blackened,pale basally; palpi yellow basally,the outerJune,1e56 Bu,lletinof the Brooklyn Entomologica,I
Society77

segnrentsbrownish black. Antennae relatively long, a little tnore
than one-half the body; scapeand pedicel fulvous, flagellum black,the proxinral third of the first segment obscure yellorv; flagellar
segmentselongate-cylindrical,
with short verticils. Front reddish;head behind the antennal basesrnetallicblue.
Mesonotal praescutum chiefly black, the cephalic fourth, withthe pronotum, obscureyellow; posterior scleritesof notum black,the posterior two-thirds of the mediotergite and all of the pleurotergite fulvous yellow. Pleura fulvous yellow. Halteres with stem
fulvous,knob infuscated. Legs with all coxaeand trochantersfulvous yellow; fenrora obscure yellou', the tips weakly darkened,
tibiae sirnilar with more narrowly darkenecltips; tarsi yellowish
brolvn basally,outer segmentsdark brown. Wings with the groundpale yellou', the prearcular and costal regions more saturated yellow; a narrow but conspicuousbrorvn crossbandat cord (presumably crossingthe wing but the posterior part darnagedin the uniquetype) ; wing tip darkened as far back as the level of fork of Rr*r,'veins brown, rllore yellowish brown in the brightenedparts. Abundant ntacrotrichiain the outer cells, in R, occupying all but thebase,in cell Ru less than the outer half. Venation: Rs subequalin length to r-nt, straight; basal section of Ru conspicuous;veinR, oblique, as in hophinsi,,lully as long as R,*r; cell R, about twothirds its petiole, cell M, about one-third this length.
Abdonrinalsegrnentsyellow, their posteriorbordersdark brown;
fifth and succeedingsegrnentsdark brown, including the hypopygiurn and appenclages. Male hypopygium with the tergite profoundly divided into two slender arms that are directed dorsad
and thence caudad, broad at base, thence nrore narrowed, at tips
suddenly produced into long straight spines, these provided withweak setae,the extreme tips glabrous. Dististyle produced intoa long slender arm, near base with a second smaller branch thatis slightly more than one-third as long, its tip obtuse,with conspicuous setae; at extreme base of style with a short-stemmedclub,with long coarse setae. Sternal lobes in profile appearing subtriangular, the lower and outer points obtuse.
Habitat : I3elgianCongo.
Holotylte ; Volcan Karissimbi, Ny" Muzinga, January 1926 (H.
Schouteden); Museum of the Belgian Congo, Tervuren.
I take great pleasurein dedicatingthis fly to Dr. Henri Schouteden. The speciesis closestto Ptychol>terahophi,nsiEdwards, ofUganda, which agreesin the oblique vein R, of the wings, disagreeing in the coloration of the palpi, antennae,thorax, legs and wings.
The male hypopygium of hophinsi has not been described.

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VoI. LI

Ptychoptera stuckenbergi n. sp.
Anterior scleritesof mesonotumchieflyblack,posteriorpart andthe pleura yellow; wings grayish, conspicuouslypatterned withbrown, including an irregular band along cord and broad seams
over the veins beyond the cord; vein /1, transverse;abdominal
tergites black, the basesof the fourth and fifth segmentsobscureyellow,genital segmentand ovipositororange.
Female;Length about9 mnr.; wing 8.5 mm.
Rostrum fulvous yellow; palpi black,basalsegmentpaler. Antennaewith scapeand pedicellight brown, first flagellar segment
obscureyellow, succeedingsegmentspassinginto black; flagellar
segmentssubcylindrical,longer than the verticils. Head withfront fulvous; vertex and occiput polishedblack.
Pronoturrr fulvous. Mesonotal praescutum chiefly polished
black, the anterior third fulvous, isolating a small black median
triangle on cephalicend; scutal lobes chiefly fulvous, the rnedianarea narrowly black; scutellumfulvous, parascutellablack; postnotum and pleura uniformly yellow. Halteres brownish black,
only the baseof stem pale. Legs with all coxae and trochanters
yellow; femora obscureyellow to brownish yellow, the tips narrowly brownishblack; tibiae dark brown, tarsi passinginto black.
Wings with the ground grayish, conspicuouslypatterned with
dark; prearcular and costal regions light brown; a dark brown
seamover cord,its marginsvery irregular or erosedtteto palespots
seamedwith
in the abuttingcells; veinsbeyondcord conspicuously
dark brown, leaving the centerspale, involving about the outerhalf of the includedveins; a faint dusky suffusionover urostof cellalong vein Cu; veins brown.
R and in the baseof M ; a dusky sean-lMacrotrichia of outer cells extending basadto lrcyond the radial
fork. Venation: R, transverse,Rr*, unusuallylong, approximately
two-thirds to three-fourthsRs,' basal sectionof R., very short topunctiform ; cell Ra about as long as its petiole, cell Znd M t about
one-third its petiole.
Abdominal tergitesblack, the basesof the fourth and fifth segments obscureyellow; genital segment and ovipositor orange;
basal sterniteschiefly darkened.
Habitat: Southern Rhodesia.
H olotype: J, Rhodes-lnyangaNational Park, near Inyanga,& P. Graham);Natal Museum,
January14, 1955(8. Stuckenberg
Pietermaritzburg.
This interestingfly is named in honor of Dr. Brian Stuckenberg, who hasadcledmost rnateriallyto our knowledgeof the Dip-

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79

tera of South Africa. The speciesis quite distinct from Ptychoptera capensisAlexander and P. matongoensisAlexander, the only
other known regional speciesof the genus, in the coloration of thethorax and wings. The irregular darkened band along the cordof the wing is very different from the condition found in the above
nrentioned speciesthough similar to that of. P. ghesquierei (Collart), of the Belgian Congo, an otherwise quite different fly.
TRICHOCERIDAE
yia imani,shii (Tokunaga)
Karutasent
Alfredia imanishii Tokunaga; Annot. Zool. Japon., 15: 98, pl. 11,
fis. 2 (adult Q) ; 1935.
Trichocera i,tna,nishiiTokunaga; Annot. Zool. Japon., 15: 468;
1936.
Trichoceraitnanishii E saki et al., Icon. Insect. Japonicorum,Ed. 2:1512,fig. 4337 (adult t) ; 1950.
KazuasernY,ia
imanishii Alexander; Trans. Shikoku Ent. Soc., 3:
14-16, fig. (venation,d) ; 1952.
The type, a female, was taken on Mount Sasagamine,Echigo,
Honshu, Japan, on March L7, 1932, by Kinji Imanishi. Later,
both sexes were discoveredby Eiji Kawase, also in Echigo, andnot far from the type locality. In January 1955, Kintaro Baba
found still further specimensat Kurokawa, Echigo. To my knowledge, the fly still is known only from the Province of Echigo.
A very distinct fly, very different fronr all other speciesin the
fanrily by the nearly winglessfemalesand by the males having thewing shape and venation quite unlike that of any Trichocerid
hitherto made known. The remaining generain the farnily, including Trichocera Meigen (Holarctic, introduced by accidentin various placesin the southernhemisphere),DiazosmaBersroth (Holarctic) , ParacladuroBrunetti (Eastern Palaearctic,western Nearctic, Maorian, Chilean),I{othotrichoceraAlexander (Australasian),and PolaeopetauristaS6guy (Kerguelen Island, Indian Ocean),all have a uniform type of venation that differs but slightly in thevariotrs genera. Kawasernyia, on the other hand, has the wing
shapeand venation so different that the latter cannot be homologized with any of the above.
The figure of the male provided at the time I proposedthe genus,
as allove cited, shows the wing shapeand trichiation satisfactorily
but the venation as there shown and describedis evidently variable
within certain limits. In the specimens now available, there
are two outer forks on the wing disk, interpreted as being R and80

Bullet'in of the Brooklyn Entomolngi,calSociety

vot.I'lM, connectedby what seemsto be the r-m crossvein. As in theother specimen,there are no veins at all in the anal field or behindthe single vein interpreted as being Cr. In the new material, Sc
is present, ending beyond the level of r-tn, Sc, only moderately
retracted; no crossveinin the radial field, as in the earlier specimen; r-nx connectingat fork of.M but far before the fork of Rs.
The wing veins are very pale againstthe nrilky white ground.
BLEPHAROCERIDAE
Paulianina robinsoni n. sp.
Generalcoloration dull black, the thoracic pleura clear pale yellow; antennae,halteresand legs chiefly black; wings nearly hy"line, stigmal area narrowly blackened,veins black; abundant macrotrichia in outer wing cells; 1ts before level of.r-rn; basal sectionof M r*o about its own length before r-w, not fused with C'r,.
F etnale: Length about 4 nrnr.; wing 6.2 ntm.
Mouthparts castaneous;palpi brownish black, 4-segrnented,thesegurentsprogressivelyshortened. Antennae lS-segmented;scapeand pedicelbrown, flagellurnblack; flagellar segmentssubequalin
length, gradually decreasingin diameter, covered with a delicate
white pubescence.Head brownish gray; a compressedtubercle
irrrntediately
behind each antenna; ocellar tubercleconspicuous.
Thoracic dorsum dull black, subnitidous. the surface sparsely
pruinose, more heavily so laterally and on posterior sclerites.
Pleura clear pale yellow. Halteres black,the baseof stem yellowed.
Legs with the coxaeyellow, trochantersa little darker; remainderof legs dark brown, the bases of the fore and middle femora
slightly paler; legs, especially the femora, conspicuouslyrnore
slenderthan in hoaa. Wings nearly hyaline; stignral area narrowly blackened; veins black, conspicuous. Abundant macrotrichia in all outer cells, especiallynumerous in the radial andmedial fielcls but including the entire rrrargin back to the anal
lobe; rows of trichia in cell5 Rr, M u, M, and M o. Venation: Rs
before level oL r-rn, the spur about as long as either sectionof Rs,'
basal section of M r*o about its olvn length before r-nt, not fusedwith Cu,.
Abdomen dark brown, sternites with paler posterior llorders.
Genitalia appearing as two compressedblades, their inner faces
closely applied.
H abitat: Madagascar.
Holotype: 9, km 57, Route Anosibe-Tananarive,
Februarv 1955
(A. Robinson); Institute Scientifiquede Madagascar.

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81

I am pleasedto namethis fly for the collector,Mr. Andria Robinson, Preparator at the Institute Scientifique de Madagascar,towhom, and to Dr. Renaud Paulian, I am indebtedfor great assistance in making known the crane-fliesand some other groups ofDiptera of Madagascar. The only other species of Paulianina
known to this date is the genotype, P. hoztaAlexander, which differs evidently from the present fly in coloration and in details of
wing venation.
It is of great interest to know that Paulian discoveredno fewer
than sevendistinct types of larvae in the mountain streamsof Madagascarthat are referrable to the subfamily Edwardsiinae and presurnablyto the genus Paulianina Alexander. The following papers
bear on this subject.
Alexander, C. P. A new genus and speciesof net winged midge
from Madagascar. Mem. Inst. Scient. Madagascar, E. I :227-230, I fig. ; 1952 (descriptionof Pauli,aninaand P. hova).
Paulian, R. Fauna des Eaux douces de Madagascar. Ibid., E,

IV: 431441,15 figs.; 1953.
The fly herewith described may well be the speciesof which
Paulian describedthe larva from material taken in the forest northof Anosibein January 1951 (Paulian, 1.c.,p. 435, fig. 10).