4a7

From

THE PROCEEDINGS OF THE
ROYAL ENTOMOLOGICALSOCIETY OF LONDON
SedesB. TAXONOMY
Vol. tS, Parts rr-r2, Decemberz1rd, 1946

The Royal Entomological Society of London,
4r, Queents Gate, S.W.7

132

NEW OR LITTLE.KNOWN SPECIES OF' EXOTICTIPULIDAE (DIPTERA) IV
By ProfessorCharlesP. Ar,nxmvopn,F.R.E.S.
Statc,College,Amher st, M assachusetts')(M assachusetts

Tnn precedingpart under this generaltitle_was published in 1945,Proc. R. ent.
Soc.Lond,.tgj i* : g5-102. At this time I am discurliog further new speciesof crane-fliesthat were collectedby *y friend, the late Mr. Alexander Cuthbertson, in Southern Rhodesia and the nearby western section of PortugueseEast
Africa (i\tozambique). The types are preserved.in my collectiorr of these flies,
and the secondspecimenof avi,ilable speciesis being sent to the British Museum(Natural History).
insinoerasp. n.
Dolichopeza(Trichoiloliohopeza)
Size small (wing, female, less than I mm.)1 mesonotal praescutum with four ill-definedbrown stripes; femora and tibiae testaceousbrown, the tips very vaguely more darkened,tarsi dirty yellow; wings with a brownish tinge, cell So and the stigma darker brown;
three isolated yellowish-white obliterative spots, these being prestigmal, poststigmal and&crossthe fork of LI; macrotrichia of wing cells relatively numerous in eells Sc, to Cu,
inclusive; stigmal trichia very few; outer forks of JI deep, cell J/1 more than four times
its petiolei rn-cu only moderately oblique in position.
FemaIe. Length about 8 mm. ; wing 8'{ mm.
Frontal prolongation of head testaceousyellorv above, darker on ventral half; nasus
fistinct; palpi dark brown, the terminal segment somewhat paler. Head above brownishgrey.
Pronotum brown. Mesonotal praescutum with the restricted ground brown or slightly
reddish.brown, with four very slightly darker brown stripes, the intermediate pair separated
by a very vaguely indicated darker vitta; posterior sclerites of notum, including pleurot€rglte, rather dark brown. Pleura obscure brownish-yellow, variegated beneath thewing root and less evidently on the ventral portions b5r slightly darker brown; dorsopleural membrane dusky. Halteres wibh stem yellow, knob broken. Legs with cox&e
obr"o* yellow, the bases of the middle and hind pairs narrowly infuscated; trochanters
yellow, with a small brown spot on inner face near apex; femora and tibiae testaceousbrown, the tips very va,guelymore darkened; tarsi dirty white. Wings with a brownish
tinge, cell Bc and the stigma darker brolvn; relatively broad but inconspicuouspaler brown
se&msover anterior cord and.m-ar; three yellowish-white spots, prestigmal, poststigmal
and across the fork of M, all disconnected; veins brown, except in the whitened obliter'
ative portions. lVlacrotrichia of cells relatively numerous, in all cells beyond cord, restricted
to two or three in the stigma; in cells Sc, to Jlzl1,inclusive, they occupy all but the basesof the cells though chiefly restricted to the centres; in cells 2nil Mrlo Cu they occur inthe extreme outer end, becoming still more sparse in cells MnandCu; veins unusually'
delicate. Venation : -Bsrelatively long, straight and oblique, more than one-half longer
than the basal section of Rn1 5; vein r?, meeting Rn at almost a right angle, the latter
perylendicular; 8, * , evidently preserved as a whitish element that forms the outer end
outer forks of M deep, cel! M, being more than four times its.petiole i m'cu
of .tig-";

pRoc. R. ENr. soc. LoNo. 1n; tf.

Prs. 11-12. (ouc. 1946.)

Nqw u little-knutn speciesaf erntic Tipuliilae (Diptero) IV.

r33

more than one-half its length b'eforethe fork of M, less oblique than in fauifrons; cell
Znd A relatively wide.
Abdomen, including the pleural membrane, dark brown to brownish-black, not or
only vaguely patterned with obscureyellow (abdomenof type filled with eggsand badly
shrunken); genital shield fulvous brown.

Rnoonsre: Yumba Mountains,March rgSb(Cuth- Holotype,Q, SouTHERN
bertson\.
(Tri'clrcdol'inhopeza)
insirwerais very different from its regional.-.Dol'i'chopeza
allies in the'unbrighten-edlips of the femora and tibiae. ft is more iearly
related to various species_
from the Cape District, perhaps being closest tb
D. lf .) fl,auifronsAlexa-nder,of the Calidon Division. If differJfrom thesevariousspeciesin the colorationof the thorax, and in the venationand trichiation of the_wings. The South African speciesof Tri,clwd,ol,i,clnpeza
Alexander
havebeendiscussed
in an earlierpaperby the author (L925,S. Afr. J. nnt. H'ist.
5 :39-48).
Dolichopeza(Trichodoliehopeza)
vumbicolasp. n.
Allied to albogeniculata; mesonotum brown, without pattern, the humeral region ofpraescutum broadly obscure yellow; legs brown, the genua narrowly but conspicuously
snorvy-white; wings with a brownish tinge, the dark brorvn stigma preceded and followedby very conspicuous whitish obliterative areas; wing-tip, anterior cord and m-cu seamedwith brown; abundant macrotrichia in outer cells of wing; cells of outer medial field relatively shallou', -|lf, about three times its petiole; abdominal segmentsdark brown, each witha central transverse yellow ring; male hypopygium with the outer dististyle glabrous anddilated at base, the outer two-thirds very slender; a,pex of inner dististyle with three orfour strong scattered teeth.
Male. Length about g mm.; wing 10.5 mm.; antenna about 2.6 mm.
tr'rontal prolongation of head pale yellow above, more infuscated on ventral half;nasus long and conspicuous; basal three segments of palpi testaceous yellow, terminal
segment broken. Antennae with scape and pedicel light yellow, flagellum dark brown;
antennae relatively long, broken beyond the ninth segment, when entire being approximately one-fourth the length of wing; longest verticils unilaterally distributed on uppersurface, on the intcrrnediate segrnents being a little shorter than the segment; a single
ehort verticil on lower face near base; remirinder of segment rvith a short dense pale
pubescence. Front and anterior vcrtex clear light yellorv, the pcsterior vertex somewhat
more darkened, especially on sides; vertical tubercle low and entire.
Pronotum testaceous, the central and lateral portions slightly darker brown. Mesonotum brown, without pattern, the humeral region of praescutum broadly obscure yellow,
the lateral border restrictedly grey. Pleura testbceous yellow, the propleura, ventral
anepisternum, ventral sternopleurite, and an area below the wing-root slightly infuscated.
Halteres brown, base of stem restrictedly yellow, knob dark brown. Legs with the coxao
chiefly testaceous yellow, the fore pair vaguely darkened on outer faces ; trochanters yellow;
femora pale brown, the bases narrowly yellow, the tips narrowiy but conspicuously snowy
white, preceded by a subequal, more brownish-black ring; tibiae brown, the extreme bases
whitened, the amount a little less than on the femora, the tips weakly more darkened;tarsi obscure pale yellow. Wings with a brownish tinge, variegated by darker brown andwhitish subhyaline; stigma dark brown, conspicuous I paler brown areas in the costal
field, at wing apex, and as se&msover the antbrior cord and rn-cui whitish obliterative areas
yery conspicuous, especially a larger one before cord, extending from C across Mr.
*r*"into the extreme basesof cells.Bu arld M"; a smaller but equalty well-defined poststigmal
area just beyond E2i veine brown, except in the whitened obliterative &reas. Numerous
macrotrichia in the stigma and in the outer cells, including the distal ends of cells Bcg

134

ProfessorC. P. Alexandet on nsw or liuln-knfrwn

tn Ms,inclusive, in cellsR, and 8u occupyingabout the outer two-thirds of cells. Venation :.Bsshort, oblique, subequal tn r-m, originating opposite Sc2; Rz meeting .8, at an angle,
with indications of persistenceof vein R, *, along the extreme outer end of the stigma to
the costal border; cells of outer medial field shallowerthan in olbogeniculata,cell M, being
about three times its petiole; vein M, strongly arcuated on its basal half.
Abdomen ringed with dark brown and yellow, the latter colour occupying the broadcentral portion of each segment; subterminal segments more uniformly infuscated ;hypopygrum obscurebrownish-yellow. Male hypopygium with the median area of tergite
blackenedbut virtually smooth-margined; lateral angles of this flange a trifle produced
and bearing about ten shor,t peg-like spines,chiefly at apex. Outer dististyle dilated on
the glabrous basal third, thence slender, gradually narrowed to the tip, the surface with
scattered setaeof moderate length. Inner dististyle a little longer, more strongly curved.
at apex narrowly blackenedand provided with three or four strong scattered t€€th.
Holotype, $, SouTIIERN Ruoonsre:

Vumba Mountains, March 1935 (Cufh-

bertson\.
(Tri,chodglinhopem,)
The most similar described species is Dol'i,clwpez.a
the female sex.
only-intome
ahogeninulataNexand,er,of lJganda,still know!
ThiJ difrersfrom the presentfly in the details of coloration,in the distributionof macrotrichiain theiring celG,and in the venation,asthe deeperouter medial
cells.
Llmonia (Ltmonla) ilttlor sp. n.
Allied to su,bapim,lis;eizemedium (wing, male, 12 mm.); general coloration of thorax
greenish-yellow,the praescutumwith a central black stripe that is split by a capillary grey
vitta; thoracic pleura restrictedly patterned with brown; legsblack, the tips of the femora
narrowly obscure yellow, enclosing a still na,rrower dark subterminal ring; wings lightyellow, with a restrictedbrown pattern, including a ferv major spotsand seams,with somespotsin variouscells; cell -r?,narrowedout*'ardly, cell ls/ JI, irregularin outline; abdomen
uniformly brownish-yellow; male hypopygium with four long rostral spines.
Male. Length about 9 mm.; wing 12 mm.
Rostrum and palpi black. Antennae with the sca,peand pedicel black; firet two
flagellar segments brownish-black, the succeedingones paling to greenish'yellow, theelongate terminal segmentagain darkened; basal flagellar segmentshort'oval, with very
short glabrous apical necks; terminal segmentelongate, nearly one-half longer than the
penultimate. Head above brownish-grey; anterior vertex very na,rrow, reduced to alinear strip.
Pronotum light yellow. Mesonotalpraescutum with the ground greenish'yellow,witha conspicuousmedian black stripe that is split by a capillary ground vitta, the stripe wider
behind; no lateral praescutalstripes; scutal lobes,scutellumand central portion of mediotergtte blackened,the latter &re&more expanded behind; parascutella and pleurotergite
greenish-yellow. Pleura greenish-yellow,with an incomplete and relatively inconspicuous
brown stripe extend.ingfrom the cervical region &crossthe propleura and ventral anepisternum, ending at po-terior portion of latter I ventral sternopleurite even more vaguelydarkened. Halteres with st€m and base of knob pale yellow, the apex of latter broken.
Legs with all coxae and trochanters greenish-yellow; femora black with the extreme baseand wider apex obscureyellow, the latter enclosinga subterminal dark brown ring thatis mornethan twice as wide as the actual pale apex; tibiae and tarsi black. Wings lightyellow, the prearcular and costal fields more saturated; a relatively restricted brown
pattcrn, inctuding somew,hat,larger marks beyond arculus in bases of cells R and' il[;o"ig of -Es; fork of Sc; stigma and a confluent seamalong cord, this area very irregularin outline, enclosingtwo paie spots in the stigma and sending spurs outward along thespec;esof motic Tipuliiloe (Diptua) IV.

136

various veins that enter the cord; outer ond of cell leJ M* this further connected.
with
se&mBover the outer medial veins and more or less connected across the outerradialfield with a comparableseamover tho outer half of vein .Br; elsewhereon disk a few
additional ecattered spots, more or less confluent, not evident in the cubital and analcells;
in cell C a series of smaller, darker brown dots, lacking in cell Bc; veins light yellow,
darkened in the brown markings. Venation: Bc1 "trditrg about opposite
six-sevenths
the length of the long arcuated 8s, Bc, a short distance from the tip; B, intransversealignment with the free tip of Bc, ; branchesof r?sextending generallyparallel io
orruanother,more approximated on the outer third, so cell J?,at margin is only a-little
more than twice
its width near base; cell lst Mr long, irregular in ooiliou, widened outwardly,
m onlyabout one-third as long as the angulated basal section of M"; rlln shorter
thun Mr*n;rn'c'usvbequalto the distal sectionof Cu1,placedshortly beforethe forkof M ; vein2nil A
sinuous.
Abdomen, including hypopygium, uniformly brownish-yellow. Male hypopygium
withthe caudal border of tergite strongly convexly round.ed,with a fringe
oit.rg pale setae
just back from the border. Basistyle relatively large,
its area "*"u"d.irrg that of the ventral
dististyle; ventromesallobe simple. Dorsal dististyle a slender curved
rod, the tip decurved into a spinouspoint. Ventral dististyle long-oval, the main
body naryowedoutwardly, its apex obtuse; rostrum long and slender,with a close group
Lf foo" elongate
spinesat base,these from a slightly developedcommon tubercle; "nurny setiferouslobe
on face of style rrearbageof rostrum is tipped with two strong setae.
Gonapophysiswith
the mesal-apicallobe elongate, very gradually narrowed.to the slenderiip. Apex of
aedeagusbilobed.

Rnoonsre: vumba Mountains,March lg}b (cuth, .HoPty?!,J,-sourHERN
bertson);No. 4133.
Although lt-it evidently related.to several-other African speciesof thesubgenus,incl
including Limonin (Limonia) subapinalis
Al.;;d;;;thl-pr.runt " fly
corNtpicuousiy
inthecoloratioo
of
thr
b6dy,
and
and.rp;;ili
especiallyof"rin[l.e"
the legs. iiIir$S"tt

groupof formsthereoccurboth"p..i.r i"ih ti.""ii;;;il;r";;l$_lrlli:o9,rwingsandwrngs
andothers
otheir "Tr\with *J"gqwings impacutatr
immaculateo.
or "irfi;ily-J.-virtirally so. T;;;;]i
There is iil;#;
iikewisea

great rangein physicalsizelbut from the structo.u o'f the male
male nypol
hypopygiumit
seemscertain that all of these speciesare allied among themselves.

Oxyrliscus(Oxydiscus)
vumbensis
sp. n.
Size relatively large (*itg, female, over 7 mm.); general coloration ofbody black;halteres darkened, the base of stem restrictedly yellow f"-om obscurebrownish-yellow,fwith a nearly terminal brownish-black ring; tiliat spurs elongate;wings obscureyellow,
heavily patterned with brown cloud.s,including a marginal seriesat ends ofthe Iongitudinalveins; outer cells of wing with macrotrichia, including a stigmal series;
Bc, ending before
level of fork of?s, 8c, long, about four-fifths R,* ,* nf vein ,B, atrophied ; cell M lack,ng;
tip of vein 2ruL4 strongly curved into the -*gi";
oviposito" *ith valves blackened
basally, the tips of the cerci paling to yellow
Female. Length about Z mm.; wing 7.2 mm.
liostrum and palpi black. Antennae broken. Head blackened.
Thorax black, including the pleura, the surface slightly opeque.
Halteres dusky, thebasc of stem restrictedly yellow. Legs with the coxae testaceousyellow,
very narrowly
more darkened at bases; trochanters obscureyellow; femora obscurebrownish-yellow,
r*'ith a brownish-black subterminal ring, this more than three times thenarrow yellow&pex; tibiae light brown, the tips narrowly infuscated; tarsi light brown,
the outer segments a.trifle darker; tibial spurs long and conspicuous,their length about
two-thirds tho136

ProfessorC. P. Alexanderon nan)otlittle-knunn

diameter of the tibia opposite their insertion; 'claws nearly straight. Wings with therelatively restricted ground obscure yellow, heavily patterned with brown, including conspicuous solid clouds at stigma, origin of .Es, Scs, cord and outer end of cell lsl Mw andas marginal areas at ends of tn" longitudinal veins; other brown washes along veins ClL
and .Bu; veins brownish-yellow, darker in the patterned areas. Ilaerotrichia in all outercells frtm Rrto Jlnf',inclusive, chiefly restricted to the outer ends of the cells, including the
distal third of cells 8n and -Ru; stigmal trichia numerous; squama with two or three setaeof unusual length, these subequal in length to the width of the wing opposite their origrn'
Venation: ,Sc1ending just before level of fork of -Bs, Sc, some distance from its tip soRsangulatedatorigin; Rz+r*nitlongitudinal
thatBcraloneisabout.four-ffths Rr* r+{alignment with 82 + $ Rznot preserved; inner ends of cells Xa,I{u and lst Mrin approxim'"ull Mt lacking; m-cu from about one'fourth to more thanate transv"rru "ligrr-".ri;
one-third i;s length beyond the fork of M ; tip of vein 2n'tl '{ curved into the anal margin
so as to be perpendicular at border; anterior arculus preserved.
Abdomen with proximal and outer segmentsbrownish-black, the intermediate segments
destroyed. Ovipositor with the shield brown, the valves more blackened, the tips of theslender cerci paling to yellow.

Holotype, Q, SoutnERN RII9DESIA: Umtali, Vumba Mountains, January

No. 4028.
1935(Cuthbertson);polysticta(Edw"t{t),is2r3lil'iscus(Oryd-iscus.)
speciesregional
Themostsimiiir
the coloration
diffbrJinof the islandor saotn?-c infhe GuIfof Guinea. inir
;l ;h;-ilJn nritrt;r ;;d *"g1, and in_the.detailsof venationnp"lr::_{TY
the strongly'decurvedtips of v-einsCu1, lst A andznil A.
only Znd-A is so curved.to the margin.

In the present fly,sp. n.
Austrolimnophilaephippigera
discalGeneralcoloration of thorax yellorv, the mesonotal praesc'ututnrvith a blackened
and the scutelarea that includes the posterior three-fourths of the praescutum,the scutum
proxilum ; rostrum and..ti" yellow ; halteresyellow ; femora black, yellorv on about theyellowish
strongmal fifth; tibiae and tarsi light browr,, ih" tips darker; wings with atinge, restrictedly patternea *itn large pale brown clouds; cell Isl Jll, short'subrectangular,
two towlih m-cu beyond midlength; abd,omen yellow, the dorsal margins of sternites
a very narrowseven, inclusive, with a t "rro* blackened anea; subterminal segmentswith
blackened ring.
Femole. Length about l0 mm.; wing l0'5 mm'
pedicelIl,ostrum yellow; palpi brownish-black. Antennae rvith scape light yellow;
flagellar seg'and first flagellar ."g-"nt yellow at bases,darkenccl outrvardlv, remaininga'little shorter
verticils
ments passinginto black; flagellar segmentslong-fusifomr ; longest
onc-h'rlf $'ider than
than the segments. Head,yellow; anterior vertex narrow. onlv about
the diameter of the scape; head not particularly narrorved posteriorlv'
sides' thePronotum yellow. Mesonotal praescutum light yellou' in front and on
to weakly
posterior three-fourths of d.isk with three black stripes, the surface subnitidous
blackened,
opa,queby a grey pruinosity, the interspacesfaintly paler; scutal lobessimilarly
parascutella
the median are& heavily grey pruinolse; scut€llum clark brown, pruinose,
dorsopleuraltheobscure yellow. Postnotum and pldura abruptly golden-yello'rv,including
Halteressaddle'discalmembrane, the colour contrasting conspicuootty with the blackened
black' the bases
obscure yellow. Legs.with ailloxae and trochanters yellovi'; femora
the proximal
about
rather narrowly yellow, the amount subequal on all legs, including
the terminal
fifth; tibiae light brown, the tips narrowly dark brown; tarsi pale brown,
portions
costalsegmentsdarker. Wings with a strong yeliowish tinge, the prearcular andarranged as follows :somewhat clearer yellow; a restrictealtf" brown pattern, the clouds

speciesof erotic Tipiliilae (Diptera) IV.

137

very restrictedly above the posterior arculus; large are&s at origin of.Bs, cord and outerend of cell lsl Mz; a smaller spot over ,8, and adjacent veins ; apical border of wing veryslightly darkened, more evident in the typu ; veins brown, those of the prearcular field moreyellowed. Venation : ,Sc, ending about opposite one-third R, * "+ lr Bc, longer, near its
tip; Rrlongerthan -8, + z; Rz+s + 4 from one-third to about one-half.Bs+ s, gently arcuated.;petiole of cell 111,relatively short, from one-half longer than to twice m; cell lst Mrshortsubrectangular, slightly widened outwardly, with m-cu at near two-thirds the lower face;anterior arculus atrophied.
Abdomen yellow, sternites two to seven, inclusive, with a n&rrow but conspicuous
blackened a,reaon dorsal margin, closer to anterior border than to the posterior margin;
succeedingsegments and base of ovipositor blackened to form a narrow ring.

Holotype, Q, SournERN RHoDESTA: Umtali, Yumba Mountains, altitude
5400 feet, October 1940 (Cuth,bertson).Paratoptotype,L 9; British Museum(Natural History).
Most nearly allied. to speciessuch as Austrolimnophila diffusa (Alexander)
and A. ilist'igma (Alexand.er),both of British Tropical Africa, difrering conspicuouslyin the coloration of the body, legs and wings, and in the details ofvenation. Yarious African speciesthat have been consideredto fall in thegenus PseudolimnophilaAlexander seem more properly referable to AustroI'imnolth'ilnAlexander, as re-definedby Edwards (1938, Trans. Soc.Brit. Ent.
5 : 67-8). It may be observed that the important character of a broken
arculus, as discussedby Edwards, has not been correctly defined; accorfing
to him, the missing element of the arculus in the present genusis interpreted
as being a cross-vein; actually it is the basalsectionof the main stem of vein
tr[, irnrnediately after its departure from the basal fusion with vein R. The
so-calledanterior arculus in the Diptera is this section of M; the posterior
arculus is a supermrmerary cross-veinconnecting vein M wrth vein Cu.
Limnophila (Elaeophila)subannulatasp. n.
General coloration of mesonotum grey, restrictedly patterned with brown; thoracic
pleura restrictedly grey, conspicuously striped longitudinally rvith brownish-black, themost dorsal area r*'idest,passing above the root of the halteres, the latter dark brown;
coxae bandedwith dark bro*-n; legs vello*', the ft'nrorir,rvith a narro\y pale brown subterminal ring; wings pale yellow, heavily spotted and dotted rvith dark brown, including aseriesof about five larger costal areas, the stigmal one largest; a pale spot at tip of vein
.8, eneircled by a lunate darkening extending from the tip of vein ,Bnto Mr; vein znd A
bent into the anal margin at a right angle.
Ser ?-Wing 7 mm.
Head broken.
Mesonotum gtey, restrictedly patterned. with brown, including small are&s on thepraescutum that represent remnants of the usual stripes; lateral and sublateral praescutal
stripes behind the pseudosutural foveae larger and more conspicuous. Pleura restrictedly
grey, conspicuously striped longitudinally with brorvnish-black, the dark colour much
more extensive than the ground, including a wider more dorsal stripe extending from thecervical region a,crossthe dorsal pleurites to the abdomen, crossing the pleurotergite andsides of mediotergite; a second narrower, more ventral stripe includes a ring on the fore
cox&, thence caudad acrossthe dorsal sternopleurite, passing beneath the halteres; ventral
aternopleurite dark brown; dorsopleural membrane chieflv pale. Halteres dark brown,
the base of stem restrictedly pale. Legs with the coxae light brownish-grey at ends, thecentral portion of outer face patterned with darker; trochanters obscure yellow, darker
beneath; remainder of legs light yellow, all femora with & n&rrow, pale brown subterminal

138

Professor C. P. Alexander on nau ar little-knul)nring that is about equal in width to the pale apex; setaeof legs long and conspicuous,pale;tibial Bpursblack, conspicuous; claws simple. Wings with the ground colour pale yellow,
heavily spotted and dotted with dark brown, including a series of about five larger costal&re&s,the stigmal marking largest, produced caudad over the fork of Rr*"*n and intoor &crosscell 8n; dark area at origin of 8s n&rrow, extending from C almost to vein J}/,
not confluent with the narrow se&m over the supernumera,rycross-vein; cord, outer endof cell Lst M2, fork of Mr*r, and a seam over the supernumerary cross-veinnarrowly
brown; remainder of cells with almost circular or subcircular brown spots that are inpart confluent; longitudinal veins with dark marginal spots, lacking on fis, the area at.Bnconfluent with the one on M y to form a lunate darkening that enclosesa pale spotat the tip of the wing; veins yellow, darker in the patterned &re&s. Venation : Sc.
ending immediratelybeforc fork of ,Es,Bc, near its tip; ,Bs angulated at origin, in longitudinal alignment with Rr* "* n, the latter about twice the basal section of .Eu; supernumerarycross-veinin cell M lylr;lg opposite or basad of origin of .Esi nx-cuabout one-half its length
beyond the fork of M ; vein 2nil / before apex strongly arched, slightly widening the cell,the tip entering the posterior border at a right angle.
Abdomen brownish-black on basal portion, the outer segmentsbroken.

Holotype,Sex? Sourrrnnr.rRnoopsre: Vumba Mountains,March'1935(Cutl$ertson); No. 2397.
The only similar described.speciesis Limnophiln (Elneophiln) ma,rnwratoeformis (Riedel), of British Easf Africa. This fiffers in the coloration of thebody, the uniformly yellow legs,and in the details of wing pattern and venation,
as described.
Limnophila sumhta sp. n.
General coloration dark brown, pruinose, the praescutum u-ith three darker brown
stripes, the median one divided in front by a paler line; haltcres vellorv; s'ings relatively
n&rrow,weakly tinged with brorvn,the oval stigma a trifle darker bro*-n; male hypopygiumwith the outer dististyle nearly parallel-sided,the outer apical angle produced into astrong curved spine, with a smaller spinule on its outer margin ; outer surface of style withconspicuoussetae; inner dististyle with its distal portion very na,rrow, terminating in asingle seta; gonapophysis small, terminating in two or three strong spinous points;
aedeagusunusually small and weak.
Male. Length about 5.5 rnm.; wing 6 mm.
Rostrum and palpi brownish-black. Antennae with scape and pedicel dark brown;
flagellum broken. Head dark brown; anterior vertex broad, approximately four times
the diameter of scape
Pronotum dark brown, sparsely pruinose. l'fesonoturn dark greyish-brown, thepraescutumwith three darker brorvnstripes,the median one a little paler on central portion,
especiallyin front; pseudosuturalfoveae large, reddish-brou'n; tuberculate pits black,
relatively large but separate,the distancebetweenthern about equal to the narrow diameterof either, not confluent as in.flafo; posterior scleritesof notum chiefl.vdark grey. Pleura
dark brown, sparsely pruinose; dorsopleural membrane & little brightened. Halteresyellow. Legs with the coxae dark brown, sparselypruinose; trochanters obscureyellow;
remainder of legs broken. Wings relatively narrow, weakly tinged rvith brown, the stigmaoval, a trifle darker brown; prearcular cells, including the veins, yellorv; remaining veins
pale brown. Veins beyond cord, excepting Rr*"+e, with numerous long trichia. Venation : Sc, ending immediately before fork of Rs, Bc, much longer, close to extreme tip of
Bc, ; R, a 2 fully twice as long as .B2,the latter faintly indicated; .Bselongate, beyond its
base nearly straight; Rz+r*n subequalin length to the long cell lst Mri Rz+r a little
longerthan.Br; cellMrevidentlylacking,if presentabnormallyshort;celllsf Mrrect-

spwiesof motin Tipulid,oe(Apbra) IV.

139a,ngular, slightly widened outwardly, both m and basal section of M, subequol and intransverse alignment i rn-cu at midlengtb of the cell; cell znd A n&rrower than in filatn,
tho vein gently sinuous on its distal third.
Abdomen, including hypopygium, brownish-black. Male hypopygum with the outerdististyle nearly parallel-sided, tho outer apical angle produced into a strong curved spine,with a smaller spinule on the outer margin of the latter; outer surfaco of style with conspicuous setae over most of tho length. Inner dististyle very narrow on outer third,
terminating in a single very strong seta. Gonapophysis small, terminating in two or threo
strong spinous points. Aedeagus unusually small and weak.
.

Holotgpe,6f,PonrueuEsEEasr Annrcl (MozeuarauE): Border X'am, Rio
Jardin, April 1929(Cuthbertson\;No. 2386.
The closestrelative of the presentfly is Limrnphilnf,lan (Walker), of northern Europe; L. batnaa Edwards, likewise from northern Europe, is moredistantly- related. The unique type of the present fly"Infi,lnn,
has only a single *iog
preservedbut in this, cell U, iJ'e"identlyiacking.
tn. cJIl whil6
normally present is frequently lacking in one or both wings of an individual
fly. The present speciesis distinguishedfrom filon by the details of venationand the structure of the male hypopygium, especially the outer dististyle,gonapophysesand aed.eagus.

c3