JUNE, 1947]173

ALEXANDER:CRANE-FLIESRECORDS AND DESCRIPTIONS
OF NEOTROPICALCRANE-FLIES(TIPULID2E, DIPTERA), XXIICHARLES P. ALEXANDERAMHERST,MASSACHUSETTS

By

inter
At this time I am continuing
the describingof various
not
Where
from South and Central America.
esting Tipulidae
indicated to the contrary the types of the novelties are preservedin my personalcollection of these flies.
Genus Tipula Linnaeus
wetmoreana
new species.
(Bellardina)

Tipula30 mm.

FEMALE.-Wing
to T.
Related(B.)

larga Alexander(Mexico:Michoacan).

Frontal prolongation of head light brown, narrowly dark brown above,
stout

the

includingpedicelbrown;

lightnasus;
basal

broken.

palpi

with

Antennaethethebicolored,

segments

flagellar

scape

and

scarcelydeveloped basal swellings brown, the remainder yellow; outer segments more
infuscated.
uniformly
and on sides
behind

above

Head
the

eyes,

brownishthreeformingyellow,
more

more
or

darkenedlessmedially

distinct

areas;

vertical tubercle narrow, relatively inconspicuous.
scutumPronotalnarrowly

produceddark

medially,brown.

Mesonotal

prsescutum with the ground fulvous, with four very slightly darker stripes,
thenarrow
darkenedlateralthe medianpair with
margins,lateralwithbrown
the broadevident;
stripesmergingof the praescutum
and scutum;
scutallobes chiefly
grayishwithscleritesofdarkerrestrictedly
brown;
variegatedposteriorintermediatelessdarkeninglateral margins
brown,

notum brown, the mediotergite more pruinose, more or less patterned withbrown.

and dark brown,
Pleurathe lattergraylight brownish
variegatedas a broad
and a less conspicuous
one over the ven
dorsalappearing
stripetral pleurites.
with
stem
Halteresobscureknobs
broken.
yellow,Legs
andlongerfemora;more

yellow

slender
ring

of

than
femur

in

very
narrowthesex

femalenarrow,

followedofthelarga,
especiallyofby a dark annulus
tarsi
tibe
and
light

the very
area,
equal
tip again
paler;
darkened.
the tips of the former
brown,
scarcely
Wings
the pattern
than in larga,
the darkdifferently
arranged,nearlymuch

more

extensive;

pale

area

in outerendofnarrowerslightlycolor being
groundcell R5 greatly
reduced;zigzag pale band before origin of Rs narrow, its cephalic end not expanded;
cell

2nd

R, morecell M,A

Rs
in larga,
Venation:thancelllongercell 1st M,
at its proximal
less pointed
shorter;end;
on the outer portion.
especiallyinfuscated.

uniformly

narrowed;
broader,

RI+2

JOURNAL NEW

174

YORK

ENTOMOLOGICAL[VOL. LVSOCIETY

Basal abdominal segments reddish brown, the lateral portions of thetergites darker; intermediate segments destroyed; ovipositor and the preceding segment preserved. Ovipositor with the cerci relatively strong, pro
jecting beyond the tips of the hypovalvae.
Habitat.-Guatemala.

Holotype, a badly damaged ?, Sierra Santa Elena, Tecpan,
altitude 9500 feet, November 25, 1936 (Wetmore); United States
National Museum.
I take pleasure in naming this species for the collector, Dr.
Alexander Wetmore, distinguished ornithologist and Secretaryof the Smithsonian Institution. The nearest relative is Tipula(Bellardina) largaAlexander, of Michoacan, Mexico, which dif
fers in several characteristics, as noted throughout the descrip
tion.
Genus Nephrotoma Meigen
Nephrotoma boliviana new species.
Allied to cacuminis; general coloration black and yellow, the prescutumwith three polished black stripes, the lateral pair curved outward to themargin; occipital brand broad and conspicuous; antenna black throughout;male hypopygium with the outer dististyle strongly narrowed outwardly,
the

tip

inner

subacute;near

dististyle

end with

its posteriorasinglestout

blackened flange or point; gonapophysis a little dilated, the tip narrowed
to a spinous

point;

MALE.-Lengthwithsterniteeighth
8.5 mm.;
about
wing

the

setae of normal

Frontal prolongation of head above weakly
distal

end,

clear

black

sets;

palpi

on
light yellow
black.
brownishsides

length.

9.5 mm.

and

Antennae

infuscated,beneath;nasus
brownishshort,

especially nearwithelongate,
black
throughfaceverticilsof outerincised;only moderately
segments
flagellar
short
of
inner
face
the
those
the
or
to
segments,
exceedingslightlysubequal
numerous
more
with
first
and
segment
subcylindrical,
spinous;flagellarout;spinousabout

ata small brownwithspot on orbits
orangeyellow
con
brand
and
of the vertex;
large
occipital
point
vertex wide,
into a point
in front;
narrowed
anterior
broad,of the scape.
times the diameter

setae.
theHeadnarrowest

black,
spicuous,
four
exceedingPronotumabove
scutumwith

thecentral

part

of

outer

prescutalthemedially,

clear

light

yellow,

groundlight yellow,athe medianstripebent
stripes
lobes patterned

lateradwith

infuscated
with

three

on

sides.

polished

trifle more
reddened;
scutumto the margin;ofthe vicinity
black,

Mesonotalpraethestripes,anterior
ends ofblack

extensively
the suture

yellow
exten

sively blackened; scutellum brownish black, parascutella extensively yellow;
mediotergite light yellow, with a blackened central stripe, expanded but
paler on the posterior portion; pleurotergite chiefly dark brown, partly en

JUNE,1947]

ALEXANDER:CRANE-FLIES

175

circling a large yellow mark on the katapleurotergite. Pleura yellow, ex
tensively variegated with dark brown, especially on the propleura, anepis
ternum, ventral sternopleurite and pteropleurite. Halteres obscure yellow,
bases of knobs weakly infuscated, the tips obscure yellow. Legs with coxas
more or less darkened on outer faces, the tips more yellowed; trochanters
yellow; femora obscure yellow, the tips narrowly brownish black; tibiae
brown, the tips and the tarsi blackened; claws large, toothed, conspicuously
hairy. Wings with a weak brownish tinge, cells C and Sc, together withthe prearcular field, more yellowed; stigma oval, brown; axilla and extreme
tip of wing vaguely darkened; veins brown, paler in the brightened costal
field. Stigma with about 6 or 7 trichia. Venation: Cell M1 narrowly sessile;
m-cu immediately before the fork of M.
Abdominal segments reddish yellow, the posterior borders, especially the
tergites, marked with brownish black; subterminal segments more uniformly
blackened to form a ring; hypopygium, excepting the eighth sternite andninth tergite, more yellowed. Male hypopygium with the lateral tergal
lobes broadly rounded, especially the outer shoulders, the median notch rela
tively narrow. Outer dististyle strongly narrowed on outer half, the apex
subacute.

Inner

dististyle

with

both

the

beak

and

lower

beak

obtuselyrounded, the latter blackened, the former pale; dorsal crest low; posterior
crest elevated into a single blackened point or narrow flange. Gonapophysis
half a little dilated
and narrowed
pale, on distal
withsternitebut not unusually
the setae abundant

to a spinous

point.

Eighthlengthened.

Habitat.-Bolivia.

Holotype, e, Cochabamba, Chapare, November 1930 (Francisco Steinbach).
Among the described regional species of the genus, the presentof southerncacumrinis Alexander,
fly is closest to Nephrotoma
in details of colorationof the body, antennae
Ecuador,differingof the male hypopygium,
and wings, and in the structure
espe
ciallythe tergite,

inner dististyle,gonapophysis

and eighth

ster

nite.
Genus
Gnophomyia
Generalwithofdarker;the basal

the

stem

propatula(Gnophomyia)
of mesonotum
colorationwings
sectionrelatively

head;phallosome
are weakly
bifid

at

gray;

eyes

new

Sackenspecies.

reddishbrown,
patterned
restrictedly
cell M2 open by the atrophy
tinge;
withthe inner dististyleof M,; male hypopygium
havingouterdilatedinto a long-oval
strongly
short,
portionarms
on eitherthatblackenedside by incurved
subtendedwitha weaktips,

the outerbrownishor lateral

5 mm.;
about
MALE.-Length
wing
brownishobscureRostrumyellow.
dark

Osten

Gnophomyia

large.

lobe microscopicallydenticulate.

5.3 mm.
Palpi

andantennae

broken.

Head[VOL.LV

JOURNALNEW YORK ENTOMOLOGICALSOCIETY

176

Pronotum yellow; pretergites light yellow. Mesonotum light reddish
brown, the priescutum and scutal lobes weakly darker; base of scutellum
and central portion of mediotergite somewhat paler. Halteres infuscated,base of stem restrictedly yellow. Legs with the coxe and trochanters paleyellow; remainder of legs pale brown (terminal tarsal segments broken).
Wings with a weak brownish tinge, the extreme base more yellowed; stigma
barely indicated as a narrow seam along R1+2; veins brown. Venation: Sc1
ending a short distance before the level of BR; Rs oblique, straight, in direct
with RB; r-m a short distance
beforethe fork of Rs; RB immedi
the fork of R2+3+,;
veins BR and R, long, extending
generally
beyondto one another;of the basal
sectionofcell M2 open by the atrophy
parallelm-cu
its petiole;cell 2nd M2 a little
than
about
one-third
its
Ms,
longeralignment

ately

length beyond the fork of M.
Abdominal tergites brown, sternites yellow; hypopygium dark brown.
Male hypopygium with the tergite produced into a median lobe that islonger than broad. Basistyle unarmed. Outer dististyle a slightly curved
blackened rod, relatively narrow, its tip subacute. Inner dististyle withthe base expanded, the stem relatively short, strongly dilated outwardly
into a long-oval head that is provided with several weak seta. Phallosome
with the central organ subtended on either side by incurved blackened arms
that

bifid

are weakly

at

tips,

or lateral

the outer

denticulobe microscopicallylate.
Habitat.-Costa

Rica.

Holotype, J, Pedregoso, altitude 2500 feet, January 1939
(Dean Rounds).
Gnophomyia (Gnophomyia) propatula is readily distinguishedfrom

all otherregional

species

Genus

Teucholabis

cell 1st M2

the open

by

of

the

wings.

Teucholabis

theblack,

polished

Sackennew species.

(Teucholabis) wilhelmine
coloration

GeneralOsten
ventralpleuriteswitha whitish

longitudinal stripe; pronotum yellow; knobs of halteres obscure yellow;
all

coxse

and

trochanters
middlefourthyellow;and hindfore

femorafemorabrown,
theyellow,thedistal

or

third
black

blackened,tips narrowlysubwhitishon proximal
basitarsi
dilatedwings
third;
posteriorbrown
diffusethan three broad
this color less extensiveSc,bands;
hyaline,abdominal
sternites
the lengthof Rs;
three-fifths
about
endingoppositeened;black,
bicolored,outerdististylespineelongatemuch

tubercle

MALE.-Lengththeincisuresa powerfulthanstrongerbroadly
blackened
the

axial

is tipped
6 mm.;
about

with

that

wing

male
yellow;bifurcate
rod,
one;
three

inner

elongate
5.5 mm.

hypopygium
near
apex,
with
dististylesetae.

with

the

the

lateralaslender

JUNE, 1947]177

ALEXANDER:CRANE-FLIES

Rostrum black; palpi brownish black. Antennee black throughout; flagel
lar segments oval, with conspicuous verticils. Head black.
Pronotum yellow. Mesonotum polished black. Dorsal pleurites similarly
blackened, including the dorsopleural membrane, anepisternum, dorsal ptero
pleurite and pleurotergite; ventral pleurites yellow, including the propleura,sternopleurite, ventral pteropleurite and the meral region, the more dorsal
portion of this latter with a silvery pruinose stripe. Halteres with stem
allknob obscure
black,
yellow.
Legs withfore femora
brown, moreyellowish
basally,brownishyellow;

coxae and trochanters
the distal
third or fourth

blackened; middle and posterior femora yellow, the tips blackened, narrower
on

the

where

legs

posterior

about

outer

the

is

tenth

fore

included;

tibia

light brown, the other tibiae brownish yellow, the tips narrowly darkened;tarsi black; posterior basitarsi dilated on about the proximal third. Wings
with the relatively restricted ground whitish subhyaline, with three broad
are morethat

diffuse

brown

bands

includestheprearcular
the cord;andcostalthan the ground;the palecolor
as wellas complete
bands
beforefields,band or cloud occupying
about
one-fourth
extensive

basal
dark
beyondits limits very ill-defined;band at cord narrower,
length of wing,withthe oval dark brownover the anterior
cord and confluent
stigma;anddarkerthedark

at wing

band

tip,

to

basad

extending

outer

the

end

cell

of

third
1stM2;

veins brown, a trifle darker in the patterned areas. Venation: Sc, ending
about opposite three-fifths the length of Rs, Sc2 some distance from its tip;of Rs

branchesnearlyto oneparallelforanotherthealmostentireof the anterior branch, the posterior one (R5) thence bending

lengthstronglycell
so cell R3 at margin
less than cell R2;
is very wide,
only a little
to the distal
about
and narrow,sectionof M,3; m-cu
subequal
within transverse
its length
the fork of M,
one-fourthalignment
beyondcaudad

1st M,

long

black,
black,

tergitetergites-Abdominal
Abdominal

termitesternitesthethebicolored,
bicolored,

broadly
broadly

insuresincisureselyellow, the intermediate portions brownish black; eighth segment and ninth
tergite chiefly yellow, the remainder of hypopygium darkened. Sternal
pocketonfivesegment

large

but

of

comprisedoriginating
spine

low down

on mesalcurvedto thestrongly

near

face
acute

seta.

Male

long and very slender,

lower

end

of

the mesal

point,

the

distalhalftheterminal

few

relatively

hypopygium with the spinous lobe of the basistyleflange;

with

long

setre; flange of basistyle heavily blackened, its outer margin irregularly
serrate, the outermost tooth more conspicuous. Outer dististyle a very
blackened
powerful
than
and weaker

the

near
the axial
apex,
rod, bifurcate
the latter
lateralarisingbranch,larly, very stout at base,the base on lower margin
seta;of mainouter

denticles

marginthat arethe usualtwoterminating
apex broken

continued

teeth
in about
beforenarrowedwithcurvedinto

blackened

teeth

and

a few
stem with
onto

conspicuous
axial
spine.
a broad U-shapedthebyseparated
three unusuallyabout
the tip with

long
three

strong
long

the
and

smallerspine much
almostperpendieu
long terminal
spine,
scatteredelongate

setae and
Inner

a few

appressed

dististyleelongate,lobe long,lateralnotch;
withits
setre.
AEdeagus

setee.

JOURNAL NEW

178

YORK

ENTOMOLOGICAL[VOL. LVSOCIETY

Habitat.-Surinam.

',Wilhelmina Mountains, Km 10, along stream,
Holotype,
August 23, 1943 (Geyskes).
The most similar species is Teucholabis (Teucholabis) stru
mosa Alexander, which has a somewhat similar general appear
ance, including the wing-pattern. The male hypopygium of theunique type of strumosa was defective in the loss of the outerdististyle, but the other hypopygial structures, including thespine of the basistyle and the inner dististyle, are quite distinct
from those of the present fly.
Teucholabis (Teucholabis) cuneiformis new species.
General coloration of thorax polished reddish yellow, variegated only bya cuneiform polished black median stripe on cephalic half of prescutum;
and

head

abdomenfemoralegs withdarkened;tibiaeblack,

and

tarsi

pale

yellow, the bases of the former narrowly blackened; wings with a strong
blackish tinge, the prearcular, costal and stigmal fields still darker; Sc long,
Sc ending about opposite five-sixths the length of Rs;,,,,,+ relatively
long, about
to or longer

two-fifths
than

Rs;
of

any

cell

1st M,

the

veins

and

long

unusuallyatit, n-cu

beyondnarrow,
subequal
near one-thirdits

length.
6 mm.;
5.5 mm.
about
FEMALE.-Length
wing
black
black.
Rostrumand palpi
Antennae
throughout;withelongate

relatively
Pronotalnarrow,
scutum

oval,

Head

verticils.

uniformly
than
wider

one-half

aboutbrownishscutellum

black,

polished reddish yellow, variegatedmedianonstripeknob

whitish,half

cephalic
blackened.

of

Legs

flagellarsegments
vertexanterior
brown;
the diameterof the scape.
Mesothorax
yellow.
uniformly
dark

only by a cuneiform polished blackwithstem obscure
Halteresprescutum.
allwithtrochanterscoxme and
abruptly

andblackened, contrasting with the conspicuously brightened mesopleura

a

with

yellow,

andstigma
Sc

long,

its

tip,
Rs andfifthsnarrow,tuallyblackishstrongdarkened;
Sc
tion:
from

the bases
tibiae pale yellow,
narrowly
black;
blackened;darkened.
the apex of the terminal
segment
Wings
weaklyfemorasternum;
tarsi paleprearcularthan the

about

two

beyondthe fork
vein

of M

gentlybrownish; R,+3+4

times R.;
one-halfthan any of thelongerof M. correspondingly
section
and

outer

of

end

or at near

the

cell;
one-third

m-cu
the

fieldscostalveinsevenbrown.

more
Vena

Rs, Sc2 some distancetwo
relatively
long, aboutcell 1st M2 unusually
long

to or

subequal

lacking,
the entire

andground;five-sixths

about
endingoppositeto mn-cm
Sc1 alone
subequalvirtually

acute tips.

darkerlong-oval,

the basal

thebroad,
Abdomenthetinge,

veins

lengthened
than its

more
lengthofit; m vir
to include

beyond

the

own

cell;

cell

length2nd A

sinuous.
black,

thecerci

horn-yellow,

long,

upcurved

totheJUNE. 1947]

ALEXANDER:CRANE-FLIES

179

Habitat.-British
Guiana.
Holotype, ?, Bartica, February 10, 1913 (H. S. Parish).
The most similar described species are Teucholabis (Teucho
labis) flavithorax (Wiedemann), of southeastern Brazil, which
differs in the much larger size, coloration of thorax, venationand coloration of the legs, and T. (T.) schineri Enderlein, differing in details of color, especially the legs.
(Teucholabis) dileuca new species.
Allied to schineri; mesothorax almost uniformly orange or reddish orange,
the praescutumwithout darkening; halteres with stem yellow, knob blackened;legs with all coxmeblackened; femora and tibiae black, with dense erect black

Teucholabis

tarsalsetae;

one

segments

two whitened,

and

their

tips

narrowly

infuscated;wings with a strong brownish suffusion throughout; R2+ +4 about one-halflonger than R2; abdomen black, the valves of the ovipositor light horn-yellow.
7 mm.;

about

FEMALE.-Length

6 mm.

wing

Rostrum and palpi black. Antennae black throughout;. flagellar seg
ments oval, with elongate verticils. Head above blackened.
Pronotum and propleura blackened. Mesonotum almost uniformly polishedorange, preescutum without darkening. Pleura orange yellow. Halteres withstem

yellow,knob

triflepaler;
black

trochanters

erectLegs withbrownishblack;

blackened.

tarsalsetse;

one

segmentsathe middle
paircoxee blackened,
and tibiae black, withdense
femoraand two whitened,
the tips narrowly

infuscated, a little more extensively so on the second; remaining tarsal seg
ments

no palerblack;

femoralrings

asin schineri.

Wings

proportionately

a little narrower than in schineri, with a strong brown suffusion throughout;stigma scarcely differentiated; veins darker brown. Venation: Sc endingof Rs;
just beyond
midlengthcellthan in schineri;elevated
the basal

third

sectionof M,;

R2+3+4
1st M,
m-cu

than R,,
longerless thanvery
short,
own length
thebeyondabout

one-half

lessnarrow,

m

one

its

about

fork

of M.
Abdomenblack,

horn-yellow,

light

includingthe former

thegenitalstronglyshield;upcurved

both

cerci

to the acuteandhypovalvae

tips.

Habitat.-Guatemala.

Holotype, ?, Quirigua, March 1915 (William Schaus); UnitedStatesNational

Museum.

ally of the present fly is Teucholabis(Teucholabis)which differs in the details of coloration of theschineri Enderlein,
and legs, and in slight details of venation.
A
thorax, halteresof the holotype specimen of schineri has been given
re-description
The nearestby

the present

writer(Encycl.

Ent.,

Diptera,4: 26-27;

1927).

JOURNAL NEW

180

YORK

ENTOMOLOGICAL[VOL. IV

SOCIETY

Genus Erioptera Meigen
Erioptera (Mesocyphona) surinamensis new species.
Allied to inmmaculata; size very small (wing, male, 2.5 mm.) ; generalcoloration of thoracic dorsum dark brown, the prwescutum faintly patterned;
thoracic pleura' conspicuously striped with brown and silvery white; knobsof halteres weakly infuscated; legs yellow, the femora with a pale brown
subterminal ring; wings with a moderately strong brown tinge, unpatterned;
abdomen, including hypopygium, black; male hypopygium with the dististyle
arm
outer
the stronger
two-branched,
bears
outer
of
which
the
angle
head,
a regularof head with

comb

of

at apexinto a subtriangularexpanded
two powerfulspines;appressed
apex
smallerteeth and with
innerlong erect sete;branch a very elongate paddle; gonapophyses single on either side, appearing

as flattened

dark-colored

blades,the distal

fourthor morewith

setee andappressed pale spinule.

dark,

flagellar

2.5 mm.;wing
black.
brownish

about

MALE.-LengthandRostrum

palpi

somewhatsegments

paler.

2.5 mm.
with

Antennae

and

scape

pedicel

dark.

Head

Pronotum brown, the posterior portions of the scutellum obscure yellow.
Mesonotum brownish gray, the prascutum faintly patterned with darker
on

brown

the

interspaces.
latterthewhite,
borderednarrowlybymembrane
pleuralinfuscated.
Legs

paler
with

silvery

mainderlegs yellow,
with
ring.
Wings
a
and
costalfieldsarcularmacrotrichia

darker.

of M,;

Abdomen,vein

Halteres

with

coxa

and

trochantersfemorawitha relatively

brown.
thetheofterminal

sectionandPleurawithbrown
stripedconspicuouslyisas a broadthatlongitudinalappearing
stripescleritesandthe dorso
darkerthe ventral
brown,a moderately
trifle more

Venation:

Cell M1

A

moderately

2nd

only

stem

broad,
brownishstrongveins

yellowed;
open

the

by

sinuous,knob

yellow,
testaceous
pale

re
sub

the

pre

tinge,
very

weaklyyellow;brown

brown,
paleof the basal

atrophyshort.

relatively

including hypopygium, black. Male hypopygium distinctive;

2-branched,dististyleatmoreconspicuous,

dark

coloredandstouterouterbranchthroughout;theinto a subtriangular
head,

some

outer

angle

apex
expanded
severallong erect black
apex withproduced
at least
two very powerfulwithof dilated
portionwhat

setae;

apex

appressed

of

black

spines,awiththe apex,
the intervening
towardapical margin
tips directed
at extreme
inner branchof smallerequal
nearlybase,
teeth;
arising
the apex
in length
to the outerarm,
expandedoutwardly,
very
gentlyas a flaton eitherobtuse.
side, appearing
singleobliquely
Gonapophysis
their

comb

more
narrowed
and gently
thirdslightly
fourthdistalsurfaceofmicroscop
truncated;
incurved,
apex obliquely
setme
abundantratherthird withof distalthe outer margin
icallyscabrous,
tened

dark-colored

blade,

the distal

the

and

a few

pale

spinule.

Habitat.-Surinam.
Holotype,

A, Paramaribo,March27, 1939, at light(Geyskes).

1947]JUNE,

ALEXANDER:

181

CRANE-FLIES

is one of the smallest
surinamensis
Erioptera(Mesocyphona)
It is readily told from the now
known membersof the subgenus.

relatively numerous allied forms by the structure of the malehypopygium, especially the dististyle and gonapophysis. I am
indebted

very muchfor many

to Dr. Geyskes

fromfine Tipulidae

Surinam.
Eriopteraintercepta new species.

(Mesocyphona)toAlliedimmaculate;

legs

all

yellow,

femorawith

tworings; wings with a brownish tinge, unpatterned;
blackened, including the dististyle and gonapophysis;rodcurved

that

a

bears

small

and

flangeannarrow

brown

male

hypopygium
dististyle a single

additional

acute

and

at

spine

gonapophysis single on either side, the subacute tip pale.
MALE.-Length about 2.5 mm.; wing 2.5 mm.
Rostrum and palpi black. Antennae with the basal segments dark, flagel
lum pale yellow, the outer segments not or scarcely darker; flagellar seg

before midlength;

with

ments

long

verticils.

Head

dark

brown,

theanterior

vertexandorbitsgray pruinose.
Pronotum dark; pretergites narrowly silvery. Mesonotal prascutum
medium brown, vaguely patterned with darker; posterior sclerites of notum
brown.

darkerPleuradark

brown,

witha silvery

longitudinalgraystripe;dorsopleural region pale yellow. Halteres with stem yellow, knob infuscated.
Legs with the coxse dark brown; trochanters obscure yellow; remainder oflegs

yellow,

midlength,
annulus.
slightly
R.+,:.4
atrophythefemorawithtwo narrowbrown

rings,

one

at or

just

beyondthe second subterminal, about as wide as the intervening pale

with
Wings
more
yellowed;
relativelyof basal

shortsectiona brownish

tinge,

veins

brown,paleandelevated,of M,.

andthe prearcularmacrotrichia

darker.

to R,+,;

cell M.

subequal

costalfieldsVenation:
open

by

the

Abdomen brownish black; hypopygium black, including the styli andapophyses. Male hypopygium with the dististyle single, unbranched or
virtually so, appearing as a long curved rod that is only a little shorter thanthe basistyle; on face of style, at or shortly before midlength, with two small
projections that are scarcely branches, the outer one a small subtriangulardarkened
Gonapophysis
pale

the

blade,

singlea moreacute
basal
secondslendermorespine.
slightlysubacute
narrowedtotheon either
at
broad
base,
side,

apex.

Habitat.-Surinam.

Holotype, c, Paramaribo, July 13, 1938 (Geyskes).
Thenearestthe present
fly is Erioptera
is very similar
which
Alexander,
inthedetailsof structure
differing

relativebicinctipes
cyphona)general appearance,ofmale hypopygium, particularly the dististyle.

(Meso
in itsof theIn this species,

JOURNAL NEW

182

YORK

ENTOMOLOGICAL[VOL. LVSOCIETY

the dististylewith the armature dif
is much paler horn-yellow,
flange at near mid
ferent, there being a low blunt triangular
or
no
with
other
lobes.
but
pointslengthGenus Molophilus Curtis
Molophilus (Molophilus) shannoninus new species.
Belongs to the gracilis group and subgroup; general coloration of thorax
fromvarying

reddishatobtusebasistyleto darkbrown

antennabrown;

three

apex;

the

dististyles,theshort,

arranged; male hypopygium withticils unilaterally

outermostverlongestthe ventral

lobe ofandslendervery

needlelike; intermediate style a more flattened curved sickle.
MALE.-Length about 4-4.2 mm.; wing 4.6-5 mmn.; antenna about 1.1
1.2 mm.
Rostrumshort;and palpi

black.

brownish

Antennae

segments

flagellar

long-oval,

with

black

truncatedthroughout,ends;

longestrelatively
verticils

unilaterally arranged, approximately twice as long as the segments. Headdark brownish gray.
Thorax varying in color from reddish brown to dark brown; pretergites
yellow; posterior sclerites of notum and the pleura, in cases darker thanthescutellum

praescutum;

narrowlyLegsyellow.

darkened.

with

the

coxe

Halteresandinfuscated,trochanters
yellow;

base

of

stenl

remainderoflegs brown, the femoral bases more yellowed, the outer tarsal segments
brown.

a weakwith
Wings
Venation:
2R about

cell M3

relatively

darker.

short,and macrotrichia
veinsdarkertinge;
withoftransverser-m;alignment
petioleto two times mn-cu;
vein
one and one-halfbrownish
inabout

2nd A long, sinuous, ending about opposite the posterior end of m-cu.
Abdomen, including hypopygium, dark brown to brownish black. Maleobtuseat apex; mesal
lobe
with
lobe of basistyle
the ventral
hypopygium
much
or profoundly
and morebroadlyobtuse.
Three
shorterdististyles

divided branches, the outermost a slender needle-like rod, very graduallysubto the acuteof the latterintermediatestyle or branch
tip;
in length,
as a more
the tip acute;flattened
curvedsickle,appearingfaceof distalhalf withnumerous
mesal
subappressed
spines;marginnarrowed
equal
outer
of

basal

half

setula;

inner

apex

darkened

about

with

8-10

scatteredasspicules.

style
shortest,
appearingofand pointed;
surface
Phallosone
narrow,
relatively

flattened,

the

tip

narrowedanddirecteda smalleach bearing
tubercles,
a pale moretheflattenedblade,a veryscatteredfew
blade with
densely

setuliferous.

.Edeaguslaterad.

Habitat.-Argentina.
Holotype,g, Tucuman, March3
types,
Sc.
I take great pleasurein naming
memory

of my

long

time friend

1926(Shannon).

this unusually
andco-worker,

Paratopo
distinctMr.

fly inRaymondJUNE, 1947]

ALEXANDER:CRANE-FLJES

C. Shannon.

is no

There

close

described

183

relative

the

among

comparatively few known Neotropical species of the gracilis subgroup. In some regards it comes closest toMolophilus (Molophilus) quadristylus Alexander, from an unknown locality in
Brazil. This latter.differs conspicuously in the structure of themale hypopygium, particularly of the armature of the basistyleand the presence

two dististylesof onlyor branchesof the same.

Molophilus (Molophilus) pacifer new species.
Belongs to the gracilis group and subgroup; size medium
about
black,
a weak

4 mm.);
antennaethe humeral
regionbrownishtinge;

mesonotal

short;(wing, male,almostpraescutum

uniformlywithreddened;legs brown;
about
cell M3
one-third

restrictedlyofpetiole

wings

than

longermr-cu; male hypopygium with the dorsal lobe of basistyle conspicuous, its
margin produced into three spinous points, ventral lobe relatively short
and

outer

slender;tonarrowedthetip.
3.5 mm.;

about

and

brownishpalpi

a bladeinto

expanded

apex

that

terminates

inner dististyle a very slender sinuous rod, graduallyacuteMALE.-Length
Rostrumatdististylein an apiculate point;

wing
black.

4.1 mm.;antennaabout

Antennae

short,

pale

1.1 mm.
outerbrown,

segments darker; flagellar segments oval, longest verticils about one-halfthanlongerPronotumthesegments.
thebrown,

Headdarklateralportions

brown,
ofstill

darkerthescutellum,

medially.
and

the

pre

tergites, paler. Mesonotum almost uniformly black, the humeral region
more reddened; posterior sclerites of notum chiefly blackened, more or lessmedianpruinose;byweaklysparselybrown.
brown.
ofscutuminfuscated,pruinose;trochantersa weakwith
Wings
Venation:R, lying
about

level

Abdomen,ventralofmoreblack,
obscureknob

cell M,

beforeregionPleurapinning.

lobeof

one-third
n-cu,

gentlyobscureor

lessa shortthanlongersinuous.

n-clu;

beyondvein

including hypopygium,

black. Male
short

relatively

and

level

of

2ncd A

black,
or

brown

legs
and macrotrichia

ofbasistyle

coxs

the

of

veins

tinge;
distance

destroyed
with
stem

Haltereswithyellow.
Legs
remainderbrown;

brownishscutellum

yellow;
pruinose.

lightdarkerr-m;

petioleendingshortly

hypopygium withslender;dorsal

lobe

the
very

as a flattened
its innerdark-colored
blade,
edge
appearing
conspicuous,
acute
into three
the outermost
surfaceproducedlongest;spinouspoints,numerous
basal punctures.
Outer
short sete from conspicuousof lobe withof distinctive
dististyleintotion constrictedconformation;a moreslenderbasal
sinuoushalfstraight,

thethe apicalneck,in a short apiculate

outer

por

fifth

again
the
point;

into a flattened
that ends
blade
expandedouterof theconcavewith microscopic
entireserrulations;
portionmargina few microscopic
and one more
blade withtubercles
spine.
apicaldeveloped
as a verysinuousrod
slenderInner
dististyleslightly
longer,appearingJOURNAL

184
that
with

surface

YORK

ENTOMOLOGICAL[VOL. LV

narrows

blunt

Habitat.-Costa

Rica.

Holotype, A, Higuito, San Mateo
StatesSOCIETY

on outer margin
to the acuteapex
just before
tip;
acute
ones at about
three
one or two morespine, with
Phallosomic
the apex
theplatenarrowly
obtuse;length.
oval,
with microscopic
delicate
setule.
apparently

graduallya smallfourthsNEW

National

The presentof the gracilis(Pablo Schild); United

Museum.
few species
fly is very different from the relatively
so far discovered
in Tropical America,
subgroup

differing especially in the structure of the male hypopygium.
Among

such

ph ii.s)procaxregional

forms

Alexander,

it is closestof Mexico,

yet

to Molophilusabundantly(Molodistinct.