o f r P
ltretoReprinted lrom Tnn Pex.Pecrrrc Etror,ror.ocrsr
Vol. 45, No. 4, 8 Deeember1969
pp. 29t-293
Made in United Statesol Anterica

A New Species of Crane Fly Asgociated with the Plant Genue
Lopezia in Mexico(Diptera: Tipulidae)

Cnlnr.rs P. Ar,nxlxouR
Amherst, Massachusetts

During the course of his studies on the pollination of plants of the
onagraceousgenusLopeziain Mexico, Dr. Dennis E. Breedlovecollected
about a dozenspeciesof Tipulidae of which one proved to representa
new species.In order to make the name availablefor his use the fly is
describedherewith. The type of the novelty, as well as the other craneflies collected,are preservedin the California Academy of Sciences.I
am indebtedto Dr. Paul H. .Arnaudand Dr. Breedlovefor the privilegeof examiningthis interestingmaterial.
Tipula(Trichotipula)

breedlovei Alexander, new species

Mesonotal praescutum with four nearly confluent grayish brown stripes, posteriorsclerites of notum variegated brown and yellow, pleura chiefly light gray; wings
light brown, stigma and costal border slightly darker; outer wing cells with strong
TUB Prx-Pecrprc Exrouor.ocrsr 45:

2gI-ZgJ. October l!b9

292

THE PAN-PACIFIC ENTOMOLOGIST [vol.

45, NO' 4hypopygFrc. I. Tipula (Trichotipula) breedloaefAlexander. Details of maleium. Symbols: d-dististyles; r-ninth tergite'
black trichia, abundantin cell Ru; abdominaltergitesbrown, basesmore yellowed,sternites brown, yellow laterally; male hypopygium with tergal lobes bearing
abundantspinoid setae;outer margin of beak of inner dististylewith about a dozenrod
long powerful yellow bristles,facs of style near basewith a slender darkened
blackenedplate that narrowsinto an acutespine.
and a broad-based
Mlr,s.-Length about13 mm; wing 13 mm.
Frontal prolongationof head obscureyellow,nasusshort; palpi broken' Antennae
gray above,with proximal three segmentsyellow,remainderbroken. Head brownish
front, orbits and sidesof the low verticaltubercleobscureyellow'
Pronotal scutum brownish graY, scutellum and pretergites yellow' Mesonotal
praescutumwith four nearly confluentgrayish'brownstripes,the very narrow inter'yellow, para',pu.., darker brown; scutal lobesbrownish gray; scutellumbrownish
pleu'
scutella clearer yellow; mediotergitelight yellowish brown, yellowed laterally,light EraY'rotergite clear light yellow, its ventral border light brown. Pleura chiefly
yellow'propleura restrictediy brornr,; dorsopleural membrane broadly clear lightyellow, base narmetapleuralregion more obscured. Halfereswith stem brownish
yellow;,oruly yellowed,knobs broken. Legs with coxaegrayishyellow; trochantersparts and the tarsi
femora yellow, tips infuscated; tibiae obscureyellow, the outer
Sc slightly
broken. Wings ridtt brown, stigma, prearcular field and cells C andof cells Rt to Mq'
darker; veins brown. Strong black macrotrichiain outer ends
abundantincellRsiafewsimilartrichiainproximalhalfofstigma.Venation:
petiole of cell Mt nearlyScr ending about oppositeone'third Rs, Sc, near its tip;
subequalin extent
twice m,l rn-cuon vein ll4r somedistancebeyondbase,the latterto Ms*$
Abdominal

brown' yellow
tergites chiefly brown, bases more yellowed; sternites(fig'1) with posteriorborder
laterally; hypopylium light yellow. Male hypopygium
black spinoid setae'of tergiie, r, deeJy notched,the resulting lobeswith abundantocroBER L9691

ALEXANDER-A NEW CRANEFLy293

those at midline long and slender,the most lateral ones small and peglike. Outerdististyle,d,relatively small, about equal in length to the inner style, Betaepale;
inner style on outer margin of the beak with about 2Olong powerful yellow spinoid
bristles,the remaining setaeshorter,black; face of style in the position of the lower
beak bearing a long slenderdarkenedrod, abovethis with a broad blackenedplatethat narrows to an acutesprne. Aedeagusvery long and stout, as in the subgenus.

Holotype rna,Ie(broken), taken ABour I0 nom MrLEsNoRrHEAsroF
Mlonnl, CntnuAHUA, Mcxtco, dt 7300 feet, 29 SeptemberL966, on
flowers of Lopeziagracilrc (Onagraceae)by D. E. Breedlove.
I am pleasedto name this distinct fly for the collector, Dr. Dennis E.
Breedlove. Approximately 35 regional speciesof the subgenusnow areknown, as discussedin the Crane llics ol California" (Alexander, C. P.,
Bulletin of the California Insect Survey,volume B: 5G60, 1967) and invarious other papers that concern the speciesof Mexico. All of thesediffer from the present fly in hypopygial characters,most evidently inthe long powerful bristles on the beak of the inner dististyle, a uniquecharacter in this subgenusbut occurring in various membersof the subgenusInd,otipulaBdwards in the Oriental and easternPalaearcticregions.