686
I,'rottt l/rr,.-\xN..rl,s .\Ntr Il .tcAzlNt,: ol. N,\.ntrlt,\t, IItsrolt't'

l i e r ' . 1 2 . r ' o l . 1 , , p . 4 1 i 5 ,M r t y , l t ) i r ! | .

il]t['ULIDAE (DTPTERA).
NEW OR I-,ITT[-,}I-KNO\\II{
X( JII.-ORIENTAL.AUSTRA
L,ASIANSPE('IES.
s . A r , n x A N D n nP,h . l ) . ,F . R . E . S . ,$ " y( l H a R L n P
University of lllassachusetts,
Amherst,Massaehusetts,
U.S.A.
A'r this time I anr tlescribir)grt pal't of' the new speciesdiscovered bv
Mr. Wilfred E. I-,ongin the vicinitv of'Hentlersonand at variousstations
otr the Waitakerc Range, North Auckland, New Zealand. Theseforests
occupy the subtropicalrain forest of the north island,with kauri (Agat,his
uu,.stralis
Salisbury) the dotttinatrttree of the lowlanclsand lower hills on
rlr"yersoils. Equall.ycharacteristicare three speciesof tree ferns belonging
to the generaCuatheaanrl I)icl;sonia,togetherwith a,considerablenumber.
of' speciesof trees ancl shrubs,with associatedherbs. A completelist of
Tipuloicleaof the \\'aitakere Rangewill be provided when thethe specie-.,ofsurvey reachescornpletiort. At the present time nearlv lii0 speciesof
these flies ltave been taken. r'irtuallv all through the tireless efforts ofillr. Long, to whonr f expressmy deepestthanks for this abundant mnteria,lanrl fbr the privilegeof retaining the t.vpe-speeirnens
of'the new species.
(Breuicerrt,)vrt,itakerensis,sp. n.
Lortg1tr,rJo

General coloration dark bt'own : a,ntennr (female) short, appa,rentl.V
l2-segmented,with the outer six flagellar segmentsvery reducecl; u'ings
infuscated,eellsC trntl.Scpar:ticularlyso: a whitenerlspot at proximal enrl
of'stignra: vein .R.relatively short : abdomenrelatively short.
tr'em,al,e.-I-,engtha,bortt l4 nln.l.: wing lll.ir mm.: antenna about
1 . 2m m .
Frontal prolongation of' heatl browrt, relatively short ; nasal region
short and broad : palpi dark brown. Antennre short, apparentl.v
l2-segnrented
but with the otrter six segnrentsso reduceclanclcrowdeclas to
be clifficult to differentiate : antennretestaceousbrown, the outer segments
only vaguely darker ; r'erticils numerous, shorter than the segments.
Hea,dbrown, the vertex with golden-yellowpollen.
'l'ltoracic dorsttm chiefly clark brown,
the nredian region of thellruesctttunrwith a, \-&gueobscnre brownish-yellow central stripe. Pleura,rlurk brown, the clorsopleuralnrenrbraneiu trifle paler. Halteres brou'n,t lre kttob a little darker'. I-,egs
with the coxtebrnwn: troehantersbrownislr.1'ellow:remainder of legs chiefly vellow, the extreme tips of the fernora
rlarkenecl: outer three tarsal segmentsdark brown. Wings fully developecl,
infttscatecl.the prearcttlar field and cells C' tr,nd^9cstrongl.y so : stigmaoval, darker brown: a whiteneclspot at proximal end of'stigma and a lesst-'videntobliterative streak aerossthe base of' cell lsl M, and adjacentcells: a,small pale spot irr the anterior prearcularfielcl: veinsbrown. Veins
ln.yonclcorcl withotrt lttttcrott'ichiaor virtuall.v so except fol' a serieson
anclbaseof /l*. \renatiotr:Ncnnt pro.rinraI trnrlof stigrnaor shortly
t'ein Rnnu

.1:|6

Cl.P. Alexander on Tipuliclae

lrelble truter.clclof'rfds; r'oin 1?,,rt,trdcc,ll11"sltorttlt'tlttlttitt u,enigtnat,ic,n;petiole of'cell M, ori.y u, littlc longer tltatt'lt,; t'ttill JIr*n tr,boutas longas m,
Abdomen clark brown, the genital shield a trifle brighter. Abdornen
rlistinctly shorter than in o,eniglnratica; vah'esof ovipositor,especiallvthe
cerci, long and slender,glabrous.
H ab. New Zealanc|(North Island).
Holotypte,?, Swanson,North Auckland, Station 9, January 13, 1951
(W.
E. Lons).
'
The only other describedmemberof the subgenushavingfully-develo_ped('liexantler),
aen'i,gm.atica
wings in ihe female sex is Longrtrio_(Breaicera\
likewise frorn the North Island. This differs in the coloration of t h eboctyanclwings, details of structure of the nasllsanclantennm,and I n theclistinctly longer abdomen.
Longurio (Ma,cromnstir)amissionis,sp. n.
Size mediurn (wing about lI mm.) ; general eolorat'ion brown, theprrescutum with four- poorly-indicated darker stripes ; antennre of malelroderately long, approximately one-half' the body; wings pale brown,
weakly prlt"rnea wifn pale ; cell JII,lackitg ; nrale_lyp.oplgiuq with thc
inner clististyle complex, produced into a cleaver-like beak and a lowet'
nrore slen4ei arm ; outer apical angle of style slightly prodttced, bearincabout five strong sPinous Points.
Male.-Length about 9 rnm. : wing LI'z rnm. ; antenna about 4'5 mm.
l'rontal prolongation of head obscure yellow, the nasus short anrl
stout : palpi ctarli brown. Antennre (male) relatively- long, &s showtr
anrlby the rir"u*orements, being approximately one-half the body ; s_gape
pldicel yellow, proximal two flagellar segments brgwnjsh yellow, thesucceeding ones black ; flagellar segments- long-cylindrieal, .with trrl
abundant-short white pubescence,the verticils very reducetl, uttilaterall.r4istribltecl, on the more proximal segments being searcelYone-seYentlr
to one-eighth the segment,. Head orange : vertieal tuberele reltrtivelv
la,rgeancl bulbous, entire.
fSronotum brown, variegatecl medially ancl on sitles with three llaler&reas. Mesonot,al prrescutum and scutum brown, with very-_sliglrtl,lrllrker llrown rnarkings, including four prrescutal stripes ; scutellum andpostnoturn obscure testaceous yellow ; mesonotal vestitule short ancl
,por.*", inconspicuous. Pleura, testaceous yello-*, - glabrous. Haltereswith stem yell,ow, knob dark brown. f-,egswith the coxre testacet)lls;
trochanters yellow ; remaincler of legs brown, the outet' tarsal segrrlents
still ctarker; claws simple. Wings with the ground pale brown. cells(: and. Nc, with the stigrna,-fronl clarker brown ; obliterative ilre& wltitenecl,
stigma - in cell R r', across thebefore th_e_colspic,r.rir*, extending
outei.end of'cell .E into the base of lst Mr: small, vagtle, more yellowerl
are&s befbre mid-length of vein M in both cells.E and M, ancl in tltt'bases of cells Cu, ls.f A and 2nd A : a barelv evident brightened slltlt
lle&r orrter. entl of cell l.sl A, r'ernote fl'ottt the tnargin ; veins lrl'o$'ll,

C. P. Alexandor on Tipulidae

437

Weak scatterett tnacrotrichia on vein R o* and outer end of M r*r.
\rettatiotr: ,fls gentlv alctta,tecl;/11a"p&le,almost in alignrnentwith lhebasal section of' ,fl" ; cell M , lackirg ; cell 2nd,.4 moderately broad.
Abdomen with the basal segment testaceous; succeedingtergites
shiny obscure yellow, t,heir posterior borders narrowly dalkenecl, thesecond tergite with a,n u,clclitionalconrparablering beyond rnid-length;
oll t,he irrtermediateautl outer segmentsthe rnid-dorsal region rnore ol.
le,ssinf'uscated; basal sternites yellow, the outer ones lnore tlarkenetl.
Male hypopygiunr u'ith the tergite tl'a,nsverse,its caudal rnargin withil, broad, shallow rnedian etnargination,t,he lobes broad ; lnore than theposterior half'of'tho tergite with very u,bundantdark-colourerlsetre,
tlirected outwardly. ISasistyleshort. Inner clististylecomplex, produced
into eucleaver-like beak and a lower more slender &rln that terrninates
iu a f'ewsmall black spicules; surfaceof'the cleavernear basewith a low
flange. Outer apical angle of' style rL little produced, bearing about
five strong spinouspoints : mesal f'aceof' style nerl,rerbaseproduced into
tt low lobe that is provicleclwith long clark setre. Outer dististyle short
aud stotrt, with coru'seseta:.
Hab. New Zeahtnrl(I{ort}r Islanc[).
Holoty,pe,3, Swa,nson,North Aucklancl, Station 10, NIay l, lg4g
(W. E. Long).
This very distinct fl.\' treedslittle colllparisotrwith other species. In
tlre lack of cell tr[ , it agrees with Longurio (Macromastir) submanco(Alexander), whereas in the tnoclerately elongate antennre it suggests(Alexander). lt differs from all known speciesin thcr
L. (M.) lnesocet'u
coloration o1'the wings and structure of'the male hypopygiurn.
glaucoca'pilla,
Longurio (M ac,romu,st'ir)
sp. n.
Size snrall (wing, rnrllc:.lo nrur.or less): mesorrotalprrescutrungreen withthree conspicuous orullge stripes : r'enrainclerof body with strong greenish
tints;wings wit,h Ilr+, oblique, in alignment with the shor-bRr, meetingvein -8, at about a right angle ; male hypopygium with the tergite very
narrowly trausverse, the lateral a,nglesproduced, separated b;'a very broa,clancl shallow emarginatiort ; lateral tergal lobes with relatively sparsesetre: inner ctististyle dilated at apex, the outer lnargin with a continuous
r'ow of' about a rlozen snrall black spines.
,fuIul,e.--_Length
tr,bout 7-8'5 lturr. ; wing ti-l0 lnln. ; antenna abnub
0.9- l .Onun.
-h'etnule.--Lengthtr,bout[) rnnr. : wing I I mnt.
l'r'ontal prol<.rttgtt,tiotr
of' heirtl brownish yellow, whitened above ; lrasuse,louga,te,
whitened ; palpi dark brown. Antennre short ; soa,peand pedicel
yellow, heavily white pruinose; flagellurn bltr,ck. Head above brownislr
yellow, heavily white pruitrose,especia,llyon the antericlr vertex, the latterbroatl, a,pproxitnately three tirnes the diameter of' the sca,pe.
Pronotunr olive-gr'(ren. Mesonotal prrescutum green, wit,h threeeottspicuousolrulgo strillt,s: in nrol'(' rlesiccttetl sllecirnens,the colours
le.ssl2rillia,nttlrt-rtrglrirrtlicrr,tcrl; postorior sclerites of' noturn green, tlrt-.

438

C. P. Alexander on TiPulidae

scutal l<.rbesconspicuously ora,nge; thorax glabrous. Pleura orallge'*iin greenish refrections. Halteres of moderate length, infuscated.
Legs #ittt the coxre testaceous, with green tints ; trochanters obscureyeliow ; femora obscure yellow, restrict-edly infuscated on outer face ati,p"" ; tibire and tarsi browrrish-yellow, with green-tints, outer tarsal
s6gments passing into black ; claws (male) strongly curved, simple.
WTngswith u *e-u,kbrownish tinge, the stigma somewhat darker browtr :yeinJ dark brown. Sparse macrotrichia on outer ends of veins beyond
cord. Venation : ni moderately long, straight, subequal to R'+s ioblique, in alignrnent with the sholt R?, meeting vein -Bt at about
Rr*,'iigtrtangle ; cell"M , a little longer -t_hanits petiole.; basal section of.r
M ri, *otJthan one-half longer than M-r*r; cell znd,..4,narrow.
.q,nao*"n greenish yellow, in casesthe green pattern much heavier ;hypopygium concolotbn.. In the holotype the -lateral -te_rgalborders
Ovipositor with the valves short and blunt. Male#"'*f.iy iarkened.
hypopygium with the ninth tergite very narrowly transverse, the lateral
nttgt"r"iroduced, separ.ated!V a very broad and shallow emarginatiotr'
ancl ldwer surface witti relatively sparse scattered bltr'ck
the" m#gins-setre.
Inner distislyle _narrow basally, -1o-t" dilated at al)ex.
spinousthe outer margin with a row of about a dozensmall black spines.
Hab. New Zealand(North Island).
Holoty,pe,6, Swanson, North Auckland, Station 9, June 2it, l95tl(W.E.iong)-. Paratoqtot'ypes,
JQ, Stat'ion9, March 16, 1950; Station l0'
April'Thu25, 1950.
(Alexander)
speciesis tr,llieclto Longurio _(Macromastir)cam'pbel{1ancl .t. iA.l tupteyi (Alexander), of the South Island of New Zealtrncl.
cliffering' in' the .tnuil size, details of coloration and venation. ir'trtl
in the strncture of the male hypopygium, particularly the tergite
especial"lya,ndinner dististYle.
sp. tr.
Limonia (Dicranornyia)waita,ket'cne,
Size small (wing, female, 5 mm.) ; mesonotumweakh'itt{usctltetl.thesiclesof the giUUoil.prrescutumlight yellow: antennreblack tltroughottt,
in" Uu*"I flalelar *"!tr.rnts with strort clark apical pedicels,the tertniutrl,egment eloigate : f,ead brownish gre), the auterior verter lelativelv
haltei6s infuscated ; legs brou'n ; rvings _with a, u'eak brownish
br."oacl;'thestigrna Yery_faintl;r indicated ; ^Scshort, S1, en{ing someiinge,
disfancebefore"originoi,Br, .Sc,very long, nearly equal to r?s; cell M ropenby" the atrophY of m.
Iemnle.-f,"ttgth about 5 mm. ; wing 5 mm. ; antenna about 0'9 mm.
Rostrum broinish yellow, clearer ]eilow at base, nearly one-half
as long as the remainder of head ; palpi black. Antennreblack throughout ;basal?agellar segmentsoval, the first one with a slender basal pedicel
ancl a stout shorier clarkenedapical one ; succeedingsegments becoming
rnore elongate, with \/ery short apical pedicels,. outer segmentspassing
especially-ihe tennitral otre which is tteitr'l.r'
into subcl:liptli.icul-elongir,te,
segrnentsconsiderablyctxt'et'tlittt
twic. tSo"penlltinrate;ierticils of btr,str,lC. P. Alexanrler on Tipulidae

439

t''hesegrttettts,on tltcr outer ones less conspicuclus. Head brownish grey ;ttnterior t'ertex relatively broad, rI little exceecling t,wice the clianreterof the scape.
Pronotum coucealecl beneath thc gibbous prrescutum, apparent[.yinfuscatetl, the sitles .\rellow. Mesonotal prrescuturn clear light .yellow
on sides, the rnediatr a,rea in front infuscated, continued back to thesuture as a capillar.ydark line: scutal lobes wit,h yellow centressurroundedby nlore duskl' rings : scutellum and mediot,ergite weakly inf uscated,
the pleurotergite and pleura light yellow, unpatternetl : propleura lnore
tlarkenecl. Halteres relatively long, inluscu,ted,the baserof stem narrowl.y
vcllow. Legs with the coxre .vellow, the fbre pair weakl.y inf'uscatetl ;trochanters yellow : r'etna,inderof legs mediunt brown : claws (female)with a single long, sletrclerspine. Wings with a weak brownish tinge, thestigrna very faintl.v indicated ; r'eins brown. Venation : ,5'c short,^Sc,ending a, tlistance befbre origin of ,Bs that slightly exceeds one-thirrlt,he length oI'the lattet. A'cof'ar fronr its tip. 6'c, alone nearly as long as/ds; basal sectiou of' Ra,s a, little shorter than Rs, both a,rcuateclin asirrrilar nta,nner; Rz and free tip oI' Er, in transverse alignment ;cell M, open b"y the atroph.y ol' nt,, Mr+a subequal to ol' a little shorter
than M r; nt-clt at or just be{ore the fork of' M : cell 2'nd,A rela,tivelylla,rrow,especiallyut the pointed outer end.
Abdornen broken.
Hab. New Zealand (North Island).
Ilol,ot'y,pe,?, Swanson. North Auckland, lSt,a,tiong, Muy 2:j, lg50
(W'. E. I'on17).
'Ihe present fl.r' is quite clifferent
ti'onr the nrost sirnilar regional species,
irrcltrclingI'itttt-tniu(l)icruttotrtryia)(Alexander),L. (D.) Ttiaticu,ne'ipenni,sth,ora,r(Alexanrler) a,trtl L. (|).) t(rsalhu (Alexancler). In the open cr,ll
M , it is urost like the otherwiser vel'.1rclistinct tarsulba, which is nrost
readilv clistinguishetlb.l' the snow.1'rvhite ta,rsi.

(Dicranont'yia)utiLtred,i,sp. lr.
L'irrwn'icr,
l'iize ttrecliutu (wing, ntale, ir'5 llllll. or less) ; general coloration
unifbrnrly pale gl'een or greenish yellow : antenna,l flagellum brownish
black, wings nearl.y h-yaline, unpatterned ; cell M, open by the atroph.yo{' the basal section of iVIs, ilL-crLat the fork of M ; nrale hypopygium withthe ventral tlististyle small, its &re&a litt'le lessthu,n that of the basistyle :rostral prolongation slencler. the two spines elongate, tlhnost as long as theprolongation itself'.
.ilIrile.--Length a,bout f i>-it llMr. ; rving 5-5.i-)lnrn.
Ilostrum green : palpi & litt,le darker. Antenna with the scape palegreen. the rettraintlel browttish bla,ck,w'ith a strong greenish tinge; basal
flagellar segnlentsshort-ovtr,l,thei outer olles nrore e,longate,the terminal
segment about otre-tltinl longer thtr,n the perrultirrrrlte,pointed a,t outerc'rtrl; longest l'rrrticils o{' tlre internrerliilte seigrrrentssubeclual to tlre
grtrtrtislr: rr,rrteriorvtrrtox rrrun'ow.
segrtrctrts. Htrrr,rl

440

C. P. Aloxandor on Tipulidae'I'hor.axunifbrmly pale greenor greenishyellow, the colclttrspresuntably
fading to yellow in olclermuseumspecimens. Halteres relatively-short,,
ptrle green. Legs pale green,the tips of the femora rather narrowly but
consplcuouslydark brown. Wings nearly hyaline, unpatterued;_veinsending tr'
pale-brown, more greenishin the costal field. Venation I 1Sc,_
ihort distancebeftrrethe origin of' ,Bs,the dist&ncevariable, longest inolle paratype where it is about equal t,o four-fifths the length of'Bs ; Sc, long, approxintittelv one-half to two-thirds Bs ; cell Mz open
lry tlre atrophy of the basalsectionof Ms, cell2nd M, slightly longer tharttits petiole rn-cu at'the fork of M .
Abclomen,including hypopygiunr,greenishyellow. Male hypopygiurnwith the caudal margin of the tergite wit,h a broad V-shapeclnotch, thekrbes low, with pale setre of rnoderate length. Basistyle with thc
ycrntromesallobe large, obtuse, with sonreof the outer setrelarge andstrong. I)orsal dististyle a strongly curt'ed sickle. narrowed graduall.v
into a long terminal spine. \rentral dististyle srnall, its are&tr,little lesstharr that of the basistyle;rostral prolongationslender,the two sllitles
elongate, almost tr.slong as the, prolongation itself, pla,cetllleirl' tlto btutt
of' t[e prolongation, slightll' separated. Gonapophysis witlt the tne,salapical lobe relativety slender,its tip narrowly obtttse, wea,klt tlttrlieuet[.
Hab. New Zealand (North Island).
Holotype, d, Swanson, North Auckland. Station +. Iltrrch | (i, l9irt)(W,. E. Long). Pura,types,Jd, Henclersoll, Station [, r\[tr,1'12, 1950(' W . E . I . , o n g ) .
I take great pleasurein rledicatingthis ver'.yclistinct fly to the collector,
nlr. Wil{r;d B. Long. The only other generally sirnilar greenish species
in tlre islantls are Lhnoniu (l,)iuanontyia) wl,lthuralis eltlorophylloirles(Alexander), both of lvhich differ in(Alexander) urrd .L. (l).\ su,bui'ridis
the closeclcell lsf X[, oI' the wings and in the strttctttre of the mttlc
h;rpopygiunr,partictrlarlv the veutral dististyle.
sp. rt.
u) ugath,'ictt1rt.
Austrolinmoplt'ilu(Austrolirnno'1thil
Belorrgsto the m,qrshulligt'oup: gcnet'ttl colot'tttiottot' -pt'it'sctttttttt
browu, ihc centra,lstripcrlraling to;'ellou'oll ceplraliclrtrlf: tltttetrtttr:
Iong, Ilagellar segnrentsbicolourecl,tirllv six tittres tts lortg as lrt'ottcl:
hearl grey pruinose, more blackeneclon ol'bits, llostet'iot'r'ertex rllol'e
itlottgthe posterior'f'ulvoul; wings yellow, patterrreclwith blorvrt,tlte a,r'etls
border ocellifbrrn ; ru&le hypopygium with the outer clististy'le relativelylla,rrow, at apex decurveclinto a hook ; itruer tlististyle u'eakly forked at
lobes near thea,pex: mesaf {ir,ceof b:r,sistylewith two pale, setulif'er'<ttts
end : interbase sinrple, its tip a trifle exptltrdetl ancl hooked.
proximal'Male.-Length trbout I l-l2 nun. ; wing 12-12'5 ntnt. ; antenna abclut
J-5.5 ntlll.

Il,ostrum alnd palpi da,r'kbrown. Atrtetruxl (nrale) relatively long, ttearlt'
three se.qttreutsyellow, the stteceerlittg
orre-half the length of' bocl.v: ba,str,ltltt, otttct' lt:rlf' or' less ycrllttu',the ottter
ollcs llicgl6urc,rl,|lttck ba,sit,ll.1',
sc'gmelts trnifbrrnl.1'rltrll<rncr[: {lrr,gelltrrsogttrotrtslong-cvlittth'ic,itl.tltc,'

C. P. Alexander on Tipulidae

441

length of the intermediate ones fullv six or seven tirrresthe diarneteir;segments with a dense erect pale pubescence,adrlitional to the longer
verticils. Head above gley llrtritrose,more blackened on olbits a,djoir'ring
the eyes, the posterior vertex more fulvous ; anterior vertex relativel.v
na,rrow,sornewhatwider than the diameter of the scape.
Pronotuttr Iulvous brown, patterned transversely with rla,rker brown.
Mcrsonotal prrescutum infuscated laterally, with three brown stripes,
thc tnotlitr,trone paling to obsctrreyellow on more than the cepha,licr
half ;
scuta,l lobes infuscated, the central region broacll.y yellow ; scutellurn
obscnre brownish yellow, the pa,rascutellamore darkened exeept on theyellow lateral portions : centru,l area of' mediotergite infuscated, thesitles, with the pleurotergites, obsctrre yellow. Pleura chieflv obscureyellow. the dorsopleural region and a,djoining ptr,rt of the anepisternum
nlore intuscrr,ted. Halteres with stenr yellow, base of knob dark brown,the' trpex lllol'e narrowl.y pale. Legs with the coxre and trocha,ntersobscure yellow : femora, obscure .yellow basally, passing into brownish
black, u'ith a clear cut yellow subterminal ring placed about its length
before the tip, the ertl'enle apex tr, little ilrore reddenecl: tibiar brown,
tarsi somewhat darker'. \Vings with the ground-colour vellow, conspicuousl;rpatterned with brown. including solid areas at rlrculus, origirr<rf.Bs, cord, stigma and fork of'-Br.,r*a, outer end of cell lst M 2 ancl forkof M y.,.2,with spots at ends of r,'eins.8, and fr, ; paler brown areas andclouds <ln posterior lralf' of wing, including ocellate ones at ends of trotha,nal veins and less evidently so ovel' (!u an<LneA,rotrter end of' vein M ;cruter ureclial cells chieflv clouderl, lsrtiM, clear : rreins brown, C more)'ellowecl. \'etratiott : ,\'r:o{i'om t wo to three times its length from tip
of'^Sc:,ueal'l.r,opllosite to be.vonrlthe {brk o1'fio*3rti nt-ctr,before mirllength ot' cell lsl ^lI ,: c't'll ^l[ , r-itr'.r'irts
fi'orn tlbout twice to firlly threetimes its petiole : squarrralsettr prese.nt.
Abclorninal tergites clrieflr' inftrscir,tecl,the extreme caudal nrargins.yellow; basa,lsternites clear'.yellou': hvpopygium brownish yellow. Male
hypopygiunr u'ith the tergite proctucedinto an elongate lobe, as in thegroup. Basistyle with the. interbase a simple sclerotizerl rod, narrowed
outwardlrt but, slightlv expandeclancl hooked at, tip ; nresa,lftr,ceof style
before rlpex proclucetl into a, snrall, flattenetl bla,de, neil,l' its base witht tt'o pale setuli{'erous lobes or bltr,cles,the rnore cephalic one narrower'.
Outer distist;zle relu,tively lla,rl'ow,at apex clecurt,edint,o a, slender point,
the cruter sut'faeewith couspicuolrssetre. fnner rlististyle weakly fbrkecl
trt apex : orrter ttrargin at rteiurrnicl-length with a conspicuoussetif'erous
tubercle. (lottrryoph.ysisa,ppearingas iL clll'\'ed rorl, much exceeding thesmall .ttlea,gus.
Ha,b. Neu' Ztaland"(North Island).
Holotype, C, Swanson, North Auckland, Station 9, March 21, I 950(W. E. Long). Pctrutopotypes,Cd, Station 10, March L4-29, r 9 5 l :Station 9, Ma,rch21, 195(t(W . E . Lon11).
The plescnt fl.f" :r,lrpit,t'tntlvis trtost nea,r'l.yrr,lliedto .4ustrol,'itn,rt,opfuilu,(Edwa,rtls), <[i{I'ering in the elongate(Aust,rol'hnttophilrt),slt'i,11itttur',u,lu,

442

C. P. Alexander on Tipulidae

antennr, details of coloration, and especially in the strttcture of thc
rnale hypopygium, particularly the dististyles and interbase. The
structure of the hypopygium, part,icularly the lobes of the basistyle antl
the inner dististyle somewhat suggest A. (A,) cyatheti (Edwards), an
otherwise entirely distinct fly.
Aust'rolimnopluila(Austrol,i'rrunolthila)
subinteraentaoriunrla, subsp. tt.
Male.-Length about lo rnm. ; wing 8'5x l'9 mnl. ; antenna about
9'5 mm.
(Edwards), differing as tbllows :-Oharactersas in typical su,bint,e.r'ue.nt,u,nresonotal prrescuttrm with the disk conspicuously dark brown, theposterior scleritesof the notum still darker. Wings relatively narrow, asshown by the measurements; ground-colour very slightly yellowed,
with a relatively heavy brown pattern that includes &reas at arculus,
origin of .Es,over the stigrnal region, cord, outer end of cell tsl Mrand forkof M *zi spots at tips o{'ft3, Cu, lst A a,nd 2nd A, the last large autl
conspicuous: r'eins uniformly darkened. Venation : cell Lsf M, longet'
and narrower than in the typical form, the basal section of M,lying {aI'
distad, transverse, &pproximatel.yone-third to one-fourth the oblique ?,r,.
Abdomen conspicuouslybicoloured, the basesof the segmentsbroadly
ringed with yellow, the tips more extensively blackened, the arnountof the latter increasing on the outer segments. Male hypopygiurn withthe tergal lobes relatively narrow, less than the broad U-shaped central
notch; lobes each with about 12 to 14 setre. Outer dististyle narrower
than in the typical forrn, the spine at base oI' outer margin conspicuous.
Inner dististyle conspicuously darkenecl. .:Otleagusmore slender, only
about as stout as the gonapophysis,with n grouping of slender spines er,tnear mid-length.
Hab. New Zealancl(North lslancl).
Holotylt€,6, Swanson,North Aucklanrl, Station 10, I)ecember 8, l95o
(W. E. Long).
sp. t'r.
u,i,!,f'redlorug1i,,
Au,strolinmophila (Austrol'i,mn,o'trfi,i,1a)
Mesonotal prrcscutunr vellou', with fottr brorvtr st'ripes: lllothol'tt,\
yellow ; pleura .vellow, chieflr' covered b.v a brownish black tratrsverse
stripe ; femora with three black rings, tibia black with a n&rrow subbasal
yellow ring ; wings yellow, heavily patterned with browu : tnale
hypopygiunr with the tergite proclucedinto two narrow lobes : interbase
a-slmple sinuous spine ; outer dististyle produced into a strong tlecurved
spine.
Male.--Length about I l-12 llllll. ; wing 13-13'5 mru. ; antenna
about 3 mm.
Rostrum dull black : palpi black. Antetrnre (male) relatively short :scapelight .y'ellou',pedicel more testaceousyellow : basal flagellar segments
brown, the remainder bla,ck, elongabe,a, little sltorter than the longestofver.ticils. Head with fi'ont a,nrlrnrrclroI'\'(rrtcx lrlaekenerl,tlto cerntt'tr
the anterior vertex restrictetllv recldenerl.

C. P. Alexanderon Tipulidae

143

l'r.gtlr9r'4x rr,ltI lrunrertr,lregion of t]re tuesontlta,lpt'tlrsctttttttt.yellou',t|e poster.ior part ot' the latter with lbur tlistinct brorvtr stt'i1les,tht:
int,erinediatepair paler but inclicatedon the anterior part o{'the scleritt' ;ll,terul prmcutal borders behintl the htttueri dark browtr : scuttttn yellow
nredirr,lly,the lobes brown, rlarker behind ; scutelluttr rlark browtt, withinrlicati,onsof a, recklish central vittrr, : posttrotuur dull bla,ck, the la,terallrpr.rlelsof'the nteclioterqitea,nd pleurotergite broadly;rellow. l)lettrtl
chiefl.1,occupiecl by & brownish-black trausverse striqe on the ttresepister.1unr, completely traversing the pleura,: centres of the ptercrPletrrittl
ilrrd pleuroteigite in.rre weakly darkened. Halteres brownish blaek. t h<lba,seof stenr tgtrt ;rellow. Legs with the fbre cox& light yellow, nrid-coxrc
br.ownish black, posterior coxre brownish yellow ; trochanters yellow :f'e1rgr.abrownish yellow basalht, with three black rings, the tla,rt'owe'st
apica,l. t,he subtei'minal and postmedial rings_ broader ancl subequa,l.
liuch exceeding the clear yellow interspaces; tibire black, with a, narl'ow
subbasal yellorn:ring;tarsi browuish black, the outer segment,spaling ttr
br.own, the terrninai seglnentsagain darker. Wings with the restrictecl
ground yellow, slightly rnore saturated in the prearcular and costal fields :inost, of the surtace covered by & uniform brown pattern, itt part'rcellifbrm, with nlore or less conrplete rings at arculus, nrigin of 1ls autl
oyer.the enclof'vein 2nd A; a secondextensive darkening at and near thcr
cord, separated frortr the basa,l &reas by a complete rlarrow yel.tglv
of -Es,this cla,rkenedarea nearlv parallelcr.oss-battaat about rnid-length-yellow
spots, especially in the costal fieltl ;sicle6, variegated by small
wing-tip unit subapical cells again darketrecl, including arn ocellate tlretu
at fdrk ol' M 1, o, this outer darkenetl tna,ssitr casesrnore or less fuseclwith
the central rltukening, in other c&sesentirel.y separated by A very rra,rrow
blt complete tr,r'ctratedvellow band at near fbur-fifths of the rving iveins br.owl i1 t|cr patterner[ fielrls. blowtrish .vellow in the itrterspaces,
clear. yellow in the sultcosttll gt'ouucl rll'eas. \"etratiotr : 1ls long' square,
ancl spt rrecl at origin : -Ilo*.r-,-,tr little less than twice tn-cu; petiole of'cell lVI, subequal to the cett ; cell lst Mo rectangular, with ?n-cu,a,bout,
one-half to two-thircls its length beyontl the fork.
Abclominal tergites vtr,r'iegatetlyellowish llrowtt, blackened laterally
iinrl on the ba,sali'ings: sternites unitbt'tnly .yellou': segttrenteight bltl,ck,
fgr.r1i1gA llilr'r'owrirg- : hvpopygium cltesttrut-br'<lrvtr.Nlale hypopygiunru'ith tle tergite ltrtirtuced rtredia,ll.yirtttl two llil,r't'owlobes that, ares('l)irr.irterlb.v ir, cleep U-s]rapetl ncltch tha,t is wider than either lobe.
Basistvle g.iih the inferbasea,long, sinrple.sinuousspine. Outer dististyler.elatirlelv llal'l'ow-,its a,pex produced into iL strong decurved.-spine ;i11er. rlististyle a,bout eclual in length but tnuch stouter basally, theouter hal{'with clelicatesetre,the apex obtttse.
Hult. Neu' Zeala,txl(North Island).
Holotyltr,.
,j. Nilrotupn, North Auekltrrrtl,October'2o, 1949(W.t. Long).
j, Octollerr'
22. l9-t9.
I)rtruto1ntry1x'.
'l'6is trrirr*rritll.r'rlistirrr:t c1ir,1e-11.f
is rlctlict,tctl t o tho colltrctot',
llr,. \\'ilfi.t,<llI. l.,ong,rls ru tirrtlrer rupPreciationof' his eflirrts in rrriuking

444

C. P. Alexander on Tipulidae'l'iptrlitl tauna o{' the Waitakere Prauge. The fly
lirowrr t,hc richn'igrocinckr,
ilu (Au,st,roL'hrunoplr''iln)srrperficially suggest,s Austrolint,noph(Eclwa,rds),lltrt is ontircly different in its coloration and structure of'thourale hypopygium.
Lhtr,noplfilaplnt'yruu,sp. lI.
iSizesnrall (wing, rnale, less than 4.5 rnrn.); getreral colot'atiott tla,rkblown, sparsely pruinose ; antenrtre short, black ; legs brownish yellow,
claws with a, weak spinule ol] inner rnargin beyond mid-length ; wings
greyish strbhyaline, t,he stigma clarker, the clisk vaguely patterrned withgreyish clouds ; cell IW, very small ; rn-cu a short distance beyond thc
fork ot' M ; male hypopygium with the outer dististyle stout, the u,pex
bidentate, the surface setigerous ; gonapophysis appearing as a strortg
slightly curved rod ; redeagusvery short.
Male.-Length about 4 mm. ; wing 4'3 mrn.
Rostrum and palpi black. Antennre short, if' bent backwartl cxtending
about to the posterior end of the pronotum, black ; flagella,rsegtueuts
oval. Head dark brown, grey pruinose.
Thorax almost unifrlrrnly dark brown, sparsely pruinose, the postttotunt
and pleura more heavily so. Halteres with stem pale, the knob very
weakly darkened. Legs with the fore cox& dark brown, pruinose, theremaining coxre paler, especiallv the posterior pair ; trochanters obscuretestaceous yellow ; remainder of legs brownish yellow, the outer tarsal
segments blackened ; claws with a weak spinule ou inner margin beyond
mid-length. Wings greyish subhyaline, the stigma darker ; broad vague
to scarcely indicated grey clouds over the cord and origin of -Es; wing-basea trifle more yellowed ; veins brown, a little paler in the ground areas.
Venation : ^Sc,ending nearly opposite the fork of .Es, ,Sc, a short distancefi'onr its tip ) Rz+*, a trifle longer than the basal sectionof .Eu;-Er*" atrtl
Rr*, subequal ; cells ,Il, and R, at, margin of nearly equal width : iutler'
endsof cells Ro, Rrand lsl Mrintransverse alignment;cell Mrvery stuall,a,bout one-third to oue-fburth as long as its petiole ; cell Lst M , nArrow,
trL r'€r'y short ; nl-c'tLat-routone-third to one-fifth its length beyoncl the
Ibrk of M.
Abclonren light brown, with el rlarker brown subteltuintr,l ring ; btr,sistyles of' male hypopygiurn conspicuously paler'. Male hypulr.ygitrnt
with the outer dististyle stc-rut,its apex bidentate, in addition trt theslender apical point there being a stoutel triangular l<lwerflange ; outer
third of stvle with nunlerous setre,those of outer margin stouter. Innerdististyle narrowecl on outer half. Gt;napophysis appearing &s tl strong,slightly curved rod, the tip subacute. ^lDdeagusvery short, long-oval
in outline.
Hab. New Zealand (North Island).
Holoty,lte, 6, Waittr,kere X'ilter Station, North Auckland, October l'1,1950(W. E. Lon11).
Tlrc prcsorrt fly is nrost sirnilar to speciessuch a,sLimnoplt'iln lrftistyln
Alexander',cliffering in the ventr,tionartd, especially,the structut'o of themale hypopygium.

P. AlexA,nderon Tipulidac

44lt

JI rt,rtli,ttr
ttolilt,ilrtIrtnlli, sl). lr.
( leneral colot'atiotr of' lte,rt,rlilrtrl t lrorrlx llrown ; antenna (male)exeeedingthree-fburthsthe lerrgth oft,he borl.yor wing, black throug}ront ;wirtgs weaklv tingerl with lll'owlr, the stignra anrl a, \-ilgue seanl over theatrteriot'corclslightly more rlarkenerl:rrrale h;zpop.ygiumwith the tergal
lobes utrttsttall;. short :ttttl inc;onslliertotts.ir,pperlringrls tlark tria,ngular'
points separilted b;r ir \.er.v broarl ilnrl shtr,llow notclr; appendage oflltr,sistvleelongat'e, tlilntetl ilt tip into rL subtriangula,r head; innerrlist,istylewith the apex exterrrlerlirrto il, slender sclerot,izecl
point, thelclwet' rnargin rvith rr, btrlbous llale lobe that bears about ir, rlozen strongllla,ck setrc, two or t,hree being of unusual length.
J/Ictlt.-l.,engtlr nbout ir nllr). . wing allout it.Z rnnr. ; nntenna about-[ mnt.
Rostrum clarli brorvn : palpi bluck. Antennre black throughont :flagellar segnlents elongate-cvlinrlrieal, nruch exce('cling the longest
vert,icils. Hearl rlark brown.
]\{esonotunr nrerlirrnr llrowrr, r}lore pnrirrose behind. Pleura with a<'onspicuous
tlark brou'n rlorsirlstrille, the ventrill scleritespaler. Halteres
llalt'. I.,egsobscure.vellow.tht'tips of the fenrora ancltibirc very narrowl.y
rtttrl r'aguely tlarkenecl. \\'ings with a weak brown t,inge,the stigma a,nrl
ft vague)seam over t,he lrnterior cord darkenerl. Wing tip broken lleyonrl
cefl lst l[ r, the ventltion evidently much as in mirilica and allies.
Abclornendark brou'n, the hypopygiurn a trifle paler. Nlale hypopygium
lvith the tergal lobes untrsuallv short anrl inconspicuous, appearing a,srlark triangular'lloirrts separateclb.\'a verv broad ancl shallow u-sha,perl
trotch. Basistvlt.u'itlr its appenrlagelong-producecl,
dilaterl at tip int<lir subtriirngulirr hc,ttrl,t he t'xtlenre enrl ternrinating in a small point :& group of trborrt five lrlng ereict st'tru on outer margin befrre the hearl.
the latter with abunrlant slrort rtertsesetullt'. Outer dististvle a long
eurved rocl, its tip rlarkenerl. Inner rlististyle clistinctive, the apex
extentlerl into il slenrler sclerotizerlpoint, before &pex on lorver tttargin
rvith ft bulbous pale lobe thrrt betrrs illlout a dozen strong black setru,
tlvo or three of these vcl')' long, alrotrt erlual to olre-lrtrlfthe leng^thof
tire entire style. Phallosomeu.it[r trvo elongirterorls irrrrlthe usual cltrsterof' llla,ekenedspines.
IIab. Neu' Zealanrl (North Tslanrl).
IIrlotryln, ir llroken j, Su'rrnson, North ^-\ucl<lancl,fitation l).
J l i r r c ' l rl ( i , t f ) 5 { )( W . f i . L o r w ) .
'l'lr
is rlistinct ,lletrtl'i,rtt,noplt.ilu
is clerlieatetl to llr. Wilfred II. L,ong.
Tlre elosest relrr,tives inclutle II etul,'tttnnph,'il,a,nt ir{fica (Alexander),(Alexander), especirrllythe
il . ltrorluctu (Alexanrter) anrl ]I . ryorA'cn;si,slast. --\ll threc. species differ in the structnre of the male hypopygium.
It nr&y lle notetl that uorA:en,si,,shas tl swollen glandular area at near
one-thirrl the length of the fernur in the nrale sex. Unfortunately, t,hellosterior legs ofthe present fl.y itro broken, anrl it eannot lle stirtc'rl
u'ltether rt corllparable it,roa,
is llreseut,.

.lI(i( ' . l t . ' \ l o x a t r r l t t ro t t ' f i p r r l i r l r t , t t( I t1tto1il
nsi.s,s[). n.
i,slirr ((I rynoplislirt) ruti,lrt,k'n'( leneral coloration of rrresonotrtttttlarli clttrstnut-brown, more or lesswith ten longpnrinose in lllaces: Alltenllr.' (nraler)lllack, l(i-seglnetttecl,llrirncherl segnrents: htrlteres vellon' : feurora yellow, lllackened a,t or
imrnediatel.v befrrre the tips : wings u'hitish subhya,line,pattemed withbrown, incluclingcells(I aurl,Sr,ittrdthe wing tip; cerll.8,,sessile:abdolllen
with brnwn; tnale hypopygiurnbrownish.\'r'llou',nlore or less r-trriegatecl
with t,he llasistyle tenninatirrg in two unequal flattened blactes: hothrlististyles sinrple : gollapoph.ysissletrtler, tr trifle expandecl at &pex.
-- Itength about f) mnr.: wiug S'(i ttrttr.: ittrtennaitbout 2'ti rnnr.
,,.|[ril(.
l)r.ostnrnrrlark brown: palpi brownish black. Antentttr (male) lllackt,lrrorrghout, l(i-segutente<t,the fortrtttla lleing 2+-2*8*a ; longest
bnrnch (about liagellar seglltentfive) u little lessthan one-half the entire: last bra,nchoul.y a little shorter thanorgan or ilbout htrlf the tla,ge'llutn
the first. HeratIrlat'li brown.
Pronotunr brou'n, sparsely pt'ttitrose. Ilesonotal prrtscrttttttrrvith therlisk dark chestnut-llr"own,polishecl,tlrtr htttneraI region nlore pt'uinose:
scuttrl lobes evr.ll clrrrher',the ttrecliattru'eit brotrdly prttinose : posteriot'
sclerites of notunr light brown, lltore or less pruinose, especially thc
c:ephalic errrl <lf' the nrecliotergite. l'lettra rnecliunt brou'n, heavilv
llruinose in pltrces,least so llear the suture of the anepisternunl trtrrl
sternopletrrite. Halteres .yellorv. Itegs with t-rllcoxr brownish yellow,
heayilv pprinose ; trochanters llrownish yellow ; fenrore yellow, blackened
at or irnnrerliatel.vbefore t,he tips : fore tibirt' ancl tarsi black, middle anrl
Iinrl tibirt'ancl birsitarsivellowish bt'ou'u,the tips antl renrainderof tarsi
lrlack: posterior fenrora rlilaterl a,t upices. \\rings whitish subhyaline,
stigtna,with tr conspicuousbrown patteru that incluclesall of cells (l and ^Se,a,nrlt-unrajor confluent seatnover the anterior cord : darkened areas at originof .,flsand ttt the wing tip, the inner edge of the latter about at the fork of',I[ rrr: flr'thel rlarkenings over the ottter end of cell l.sl f,I , antl its il
restricteclseam along vein ('rr, rttol'e expancletltr,tmargirt in cells ]/n anrl
( ' rui verv va,gue darkenings at ntirl-length of t'eitt I sf A , expanrle<
Iover.the sultapicttl part of cell lsf A to the marg-in at encl of 2n,d A ,llretrrcultr,rtielcl yellow, the veins concolorous, t!" remaining r.einsilark llrown. \'eins of otrter rtredial field virtuallv glabrous, t,here being\Ienation : ('ellirborrt 15 trichia on vein tll , aud tu'o or three on Mr.
R,t sessile,fir*rn, thus lacking : cell II-, about one-fottrth longer than'.
iti petiole : cell lst l[, short-rectangular nL-ctLjust }eyond mid-length
ct'll lst II ,'. r'ein 2rtrl A sintrous, not bent strongly into the rnargitr.
Ab{onrinal tergites brownish yellow, the posterior borders darkenetl,t[e arnount of rlark colortr increasing on the outer segrnents; basal
ter.gite clark',pnrinose : brrsul sternites lnore rtttiforntl.yllrownish yellow :slbterrtrinal segnlentsa littte clarker brown ; hypopygiunt obscure .yellou'.
I,l1lc,hypopygiunt with the basistylestout, the mesal face with abrtnrlttttt
11;19er('ct vellow setrc: ntcrsalapical blade relatively small, glaht'otts,
conspicttottslysetiferousexcept trt irpex trtrcl
t ltt. 1llictr,lltlacle rrrtrelrlarget'.'l'wottt
rlististylt,s,t,herrltttet' lllol'(' slettrlt,t',
il1-rpgorrter apical rtttlrgin.

('. ['. .'\lexrltttlet' on Tipulicla,tt

1+i

rrpexproducediuto a srnilll point, on lower rnargin belbret'his spine with
two strong setre; inner style strongly arcuated at base, the outer halfclusky, with abundant srnall punctures. Phallosomeconsistingof a pairof simple gonapophyses,slender,weakl.yexpanded and pointed at tips :mcleagus
simple, the openingspaired.
Hab. New Zealand(North lsland).
Ilolotype, 6 , Swanson,North Aucklantl, Sta,tion | 2, September 30,
l e50 (W. E. Long).
lrr its general appearancethe fly is sinrilar to Gynopl'istia(Gyno7il,'istia)
rlrrct'yl,o7th,ora
Alexander and G. (G.) pleuralis Alexander, differing in thecoloration, antennal formula and st,ructure, coloration of the body,wings and legs, a,ndespeciallyin the structure of the rnale hypopvgirrrrt.
Gynoplistia (G'ynolilistia) glauea uera,to'r,subsp. n.
Jfule. -Length a,bout I l] nrrn. : wing l2 mrn. ; a,ntennaabout i]'ir lnlll.
( rloseto the typical fornr, dift'ering in cletails of coloration ancl stnrct,rrreof the male hypopygiunr.
Antennue (rnale) apptr,rent[y only | (i-segrnented, the formula being2 )-2J- ff+ :1, the long termintr,l segruent fully one-half longer than thepentrltirnateand possibly forrned by the fusion of two segnrents. Mesonotal
prirscutum with the median st,ripe dark, strongly narrowecl orl the
cephalic third of the sclerite. [.,egswith the femora blackened, witha. nrlrrow but conspicuous yellow subterminal ring placed sorne distancelrefore the tip. Abclornen with the tergites brownish yellovi, without
pruinosit.r', the otrter four segments ntore darkenecl. Male hypopygiunr
with the stnrettrre generallv ils in the typical fornr, cliffering in details.
Ninth tergite with the nrerlitrnlobe nruch reduced, transverse, its marginsubtruncate. Basist.vle with t he interbasal lobes simple, blaekenetl,
il,bundantly spiculose: nresal face of' stvle merely protuberant, withotrt
rl clearly defined lobe, trs in the typical form. Outer dististvle wit,h thetrnls feebty divergent, separaterl lly a, deep lJ-shaped notch, the snrooth&rnl longer than the scabrousone, t,he latter shorter ancl storrter than in
typical glauca Edwards.
Hab. New Zealand (North Island).
Holotype, ,3, Su'anson, North Aucklancl, littttiorr 1), ,ltltttra,r.y5, I l)51

(ll'.n.I'ong).
AntphineLul.cs(Arnphineurus) te'nu,ipolIe.v,sp. n.
Size relatively snrtr,ll(wing, mAle, about i, nrnr.)-: general coloration
to yellou' ; legs obsctrre yellow,<lrrrk ]rown, the praescutunrca,st,Aneousrlarkenerl by abundant linear brown scales; wings greyish, tta,rkenedl,y rlense black setrt' on veins tr,ncl rnentbrane; r'ein 2ntl A strrlngly
sintrous; nrale hypupygiurn with the inner rlististyle bifict; interbases
a,nd gonapophysessirnple.
Ma,le.-l.,engt,habout 5-5'2 lnllr. : wing 4'lJ-5'2 llull. ; atrtennir illrout
l-t.l rnul.

448( l. P. Alexander on Tipulidaetliostrurtt ltrowtr; llalpi bluck. Arttcnn:rrr4ri11,
scirpr)tr,rrtlpctticelbrowtt,
flagellum dark brown, the segnrerntslong-ftrsifontr, with unilaterally
clistributed verticils that are consiclerably longer than the segments,
arlditional to long erect wlritened rnore clelicatesetre. Head above da,rli
brown, with conspieuousrlark setre.
Pronotum dark brown, the scutellumpale yellow. Mesonotalprrescutum
polished yellow rneditrlly, rnore castaneouson sides antl in front, theltumeral region broadly pale yellow : scuturn, scutellum and postnoturn
rlarker brown, the two latter with very long erect black setm; rnarginsof
both the rnediotergite and pleurotergite broa,dly nlore yellowed. Pleura,chiefly clark brown. Halteres short, clark brown, the base of stem pa,[eyellorv. Legswith the coxreand trochanterstestaeeousyellow ; remainder
of' legs obscure vellow, the appearancedarkened by abundant'linear pale
brown scales: outer tarsal segmentsclarkerbrown. Wings with the grounrl
greyish, the colour obscuredby abundant dense black setreon the rreinsand in all the cells: wing-base a trifle more brightened. \'enation :Rr*r*n shorter than Rr*r; cell lsf M z small, rL much short,erthan thebasalsectionof ,il[";'ttL-cuabout,one-fifthto two-fifths its length beyonrl
tlre fork of M ; r'ein 2nd.,l strongly sinuous.
Abdomen, including hypopygium, dark brown. Segments five toseven with transversemodifiectareas that are provicled with microscopic
pegs, these more dense than t,he scattered setrc on the surrounrling
integument ; caudal lateral angles of these same segmentswith a brushor pencil of exceedinglylong setre,these almost as long as the width of
the abclomen. Male hypopygium with the tergite provideclwith scatteredcoarse setrc, at apex procluced into a single slender spine. Basistyle\rery long and slender, particularlv the portion beyond the dististyles,
provided with abundant long retrorse setar. Outer dististyle expanded
at mid-length, thence n&rrowedinto a gently curveclrocl, its tip obtuse.
Inner dististyle smaller, curved to the acute tip, at near mid-length
bearinga smallerlateral spine. Interbasesor lateral apophysesappearing
as long, slenderstems,enlargerlat apex, ternrinatingin a slightlv recurved
acute spine. flonapophvses a,pproximatedat mid-line, rrppearing as
sinrple rods, sinuous, about the rlistal two-fifths directecllateracl as along, slender,slightly upcllrverlspine,sonlewhatsuggestingthe appeara,nceof the bill'of an avocet.
Hab. New Zealand (North Island).
Holotype, 3, HendersonDistrict, North Aucklancl,Station ?, February 3,
I {)5() (W . E. Lon{J). Parcttoltoty'pe,
6, Station 2, I)ecember 11, | 94!)
(W . E. Lons).
The present fly is quite distinct from the other describeclmembers of(Amphine'unus)
the enmpbelligroup of the genus, including Anr,'ph,irLellrus
bicorniger Alexander, A. (A.) campbelli Alexander, and A. (4.) ?to;tl
Alexander, in the structure of the rnale hypopygiurn. In the bispinotts
inner rlististyle it conresclosest t,o bicornige,r,differing conspicuouslvingroup
the very different interbases and gonapophyses. The cam,pbel,l,iof theis well clistinguishetlb.y certain abdominal and genitalic chara,eters

C. P. Alexander on Tipulidae

449

male sex. The transverseslightlv elevated areaswithout modified scales
on abdominal segments five to se\ren, as described under the present,
species,and the pencils of greatly elongatedsetreon thesesamesegments,
are not fotrnd in the other members of the genus. The male genitalia,
while being somewhat as in membersof the typical group, have the lobeof the basistyle unusuallv long and slender.
subsp.n.
Amphineurlrs (Antphi,neurus)hudsonicyathetanu,s,As in the typical form, differing in the details of structure of the malehypopygium. Ninth tergite producedinto two slenderblackenedpoints,
separatedby a small V-shapednotch that is less than one-half as deepas the median produced portion ; margins of the points microscopicallyroughened. fnner dististyle with the strong black spine of the typicalform reduced to a group of small spiculose points, entirely without abrush of setre. Phallosomewith both the redeagusand the spines of the
gonapophysesmuch smaller and less conspicuous.
In typical hudsoni Edwards, the ninth tergite of the hypopygium
terminatesin two blackenedblades,the surfacemicroscopicallyroughenedor spiculose,the blades separated by a deep U-shapeclnotch. fnner
dististyle with a strong black spine on face, bearing a brush of long setre
near its base. Phallosomewith the redeagusand the subtending spinesof
the gonapophyseselongate. Both subspecieshave the outer dististyleappearing as a slender rod that terminates in about six or sevenspinous
points, with a brush of long yellow setreon faceof style back from the apex.
Hab. New Zealand (North Island).
Holotype,3, Swanson,North Auckland, Station 9, November 16, 1950
(H'. E. Long). Paratll'pe,J, Kaitaia, North Auckland, September20,
1923(J . G. Muers).
(Tasiocera)longiarw,sP. h.
Ta,siocera
Sizerelatively large (wing, male, il'5 mm. or more) ; general coloration
dark brown, the pleura and pleurotergite yellow : macrotrichia of outerwing cells much reduced; m-cu at or close to the fork of M ; malehypopygium with the dististyle slender, gently ctrrved, narrowed to theslightly decurved tip, on lower margin just beyond mid-length with anacute slender spine : phallosomewith the lateral shouldersobtuse, notextending caudad beyond the central projection ; a pair of slenderrods
near base of phallosome,separatedby a U-shaped notch that, is wider
than either lobe.
Male.-Length about :]-3'5 mm. ; wing 3',\-*2 mm. ; antenna about
2-8-3.2mm.
Rostrum brownish yellow ; palpi brownish black. Anterure (male)elongate, as shown by the measurements,dark brown ; basal flagellar
segmentselongate-cylindrical,the outer ones more dilated on about theirproximal two-thirds, narrowed at tips ; besides the scattered elongate
verticils, there are further scattered delicate setrethat are much longer,

450

O. P. Alexandor on Tipulidae

exceedingthe segmentsin length, together with more abundant shorter
erect setre. Head dark grey.
Pronotum testaceous vellow. .llesonotum dark brown, the pleuraand pleurotergite yellow. Halteres dusky, the base of stem nariowly
yellow. Leg! with the coxa. testaceous yellow ; trochanters yellow-:remainder of legs pale, the colour darkened by abundant long setoid
scales. Wingsgreyish subhyaline: veins pale brown, rnacrot,richiadarker.
Macrotrichia in outer ends of cells lacking or reduced to a very few inthe extreme apices. Venation : -Er*r.'.,subequalto or a little longer thanthe basal section of fru ; inner end of cell fI " strongly arcuated, with
nl-cxt,closeto its base,in casesslightly befbrethe fbrk of llf.
Abdomen, including the hypopygium, dark brown. ltale hypopygiumwith the dististyle terrninal, appearing as a relatively slender,-gently
curved rod that narrows to the slightly decurved tip ; on lower margin
just beyond mid-length with an acute slender spine. Phallosome
rnoderatel;tcomplex: lateral shouldersof the central tlass obtuse, notextending caudad beyonclthe central projection. A pair of slenderrods
nearbaseof phallosome,separatedlly a cleeptl-shapednotch tlrat is wider
than either lobe.
Hab. New Zealancl(North lslanrl).
_Holntype,d, Swanson,North Aucklarrd, i:jtation t3, .Julv 2tJ, tgSo
(W'. E. Lorry). Parut,r-tltoty'pe,j, .Itrlr' 2(i, lgirO: purat'y,p€s,:j, Nihotupu,
August 26, l95o ; j, \\'aitakere l,'ilter lltation, Septenrber(i, t Sl+g
(W.E.Lons).
The present lly is still another of the interesting crane-fliesfrorn the\\Iaitakere Range that I arrl naming in honour of the collector. The
speciesis readily told from other regional membersof the genus by the
structure of the male hypopygitrnr, particularly the dististyle. In its
general appearance it suggests species such &s Tasiocerrl,(Tasioceru)
rli,oplmnaAlexander and T. (T.) tritort Alexander.
I'asiocera(Tasiocera)u,,produnta,
sp. n.
Size mediuur (wing, tnale, 3'5 mrn.) ; general coloration brown. thepleura paler ; wings with a weak brownish tinge '. nl-ctl lying verv far
distad, beyond mid-length of vein M r*, or rnore than four times its own
length beyondthe fork of M; male hypopygum with the dististyle slender,
with a few scatteredpunctures; phallosomewith the central mass notproduced medially, the lateral angles slightly so.
Male.-Length about 3 mrn. ; wing lJ.5mm.
ltostrunr and palpi brown. Antennrewith the scapeand pedicelbrown ;flagellum broken. Head brown.
Thorax brown, the pleura more yellowed. l{alteres broken. Legs
with the coxa' and trochanters brownish testaceous; remainder of legs
brownish yellow, a,ppearingdarker becauseof the abundant dark-coloured
vestiture. Wings with a weak brownish tinge, the veins a little darker ;macrotrichia brown. A series of macrotrichia in outer ends of cells
Ilz to znd A, inclusive. Venation : Rr*r*n exceeding twice the basal

C. P. Alexander on Tipulidae451

section of Ru; rn-cu lying very far distad, beyond mid-length of vein
Mr*nor more than four times its own length beyond the fork of M.
. Abdomen, including the hypopygium, dark brown. Male hypopygiunr
with the basistyle relatively long. Dististyle barely terminal in position,
slender, gently narrowed to the slightly decurved spinous tip ; surface
with a few scattered pale punctures, the more basal ones elongate.
Phallosome cornplex ; central mass not produced medially, the lateralanglesslightly so, with reflexed margins : a slight protuberance or shoulder
on side of phallosome beyond mid-length.
Hq,b. New Zealand (North Island).
Holotype, ,j, Swanson, North Auckland, Station I0, July b, l g50(W.E.Lons).

The regional specieshaving the male hypopygium most similar to that,trf' the present fly include Tasiocera, (Tasioceru) d,iapharm Alexander
and 7. (f .) sent,iermisAlexander, which differ in the venation and in thedetails of structure of the male hypopygium, particularly the phallosome.
Iloloplc'ilas (Iloloplt'ilus) coronurzru, sp. n.
-Belorrgsto the pLagiat'usgroup ; general coloration unifonnly yellow,
the head brown ; wings whitiih rrrntrfuline, macrotrichia of veins"utrdanallringe long : tnale hypopygium with the beak of the basistyle slender,
gently cun'ed : basal dististyle a relatively stout straight rod that bears
a,row of six strnng spines,forming a comb <lr partial crown ; outer surfaceof' distal third of style with microscopic appressedspinulre.
^llale.--I-,engthabotrt 3 rnm.; wing 3'5 rtun.; antenna about ().8mrn.
Rostrunr vellou', palpi a little clarker. Antenna, with the scape andpedicel reddish bro$'ll, flagellunr slightl.y darker : flagellar segments
subcyliudrical. sltorter than the long conspicuousverticils. Ffeaclbrown,
sparselypruinose.
Thorax almost uni{brrnly yellow, the rnid-dorsal region sornewhat
more reddened, the surface polished. Halteres yellow. Legs with thecoxre and trochanters yellow ; remainder of legs obscure yellow, thecolour slightly darkened by the vestiture. Wings whitish subhyaline,
the veins a trifle darker ; macrotrichia pale brown. Trichia long, theanal fringe especially so. Venation : tn-ca inevident in the unique type.
Abdomen, including hypopygium, yellow. Male hypopygium with thebeak of the basistyle slender, gently curved, blackened. Outer dististylcr
with its outer arm a blackened spine, the inner blade expanded a,ttip, the tnargitr microscopically erose. Basal dististyle distinctive,
appearing as a relatively stout straight rod, pale on basal half, theremainder darkened, at apex producecl into a l'ow of six strong powerfulspines that fornt a, cotnb or partial crown : back frclnr the apex with &setuliferouscollar': outer surface of distal third of style with rnicroscopic
appressed spinulte, the outer ones stronger. ;sdeagus slender, snrooth.
Hq,b. New Zealand (Itlorth lsland).
Holntypa, 6, Pukematekeo, Waitakere Range, North Auckland, altitudcr
900 feet, May 27, 1950( W. E. Long).

+52

C'. P. Alexandor on Tipulidae

The most similar regional speeiesis Molnphilus (Moloph,ilus\flauid'ulus
Alexander, which differs especially in the structure of the malehypopygium, particularly the basa,ldististvle, which is longer and more
slender,merely toothed at apex.
sp. n.
M olophi,lus(M olophil,us)cra,ssistylus,
Belongs to the plagiatus group ; general coloration yellow, the headdarker : antennre short ; halteres and legs yellow ; wings yellow,
unpatterned ; male hypopygiurn with the basal dististyle a powerfulsimple rod, its outer fifth expandedinto a head, the obtuse apex direeted
laterad : outer surfaceof head with strong appressedspinouspoints.
Male.-Length about 3 mm. ; wing 3'4 mm. ; antennaabout 0'6 mm.
Rostrum and palpi brownish yellow. Antennre (male) short' ; scapeand pedicel yellowish brown, the latter a trifle darker ; flagellum brownish
yellow ; flagellar segments long-oval to subcylindrical, with longconspicuousverticils, some on the more proximal segments unusuallylong and conspicuous. Head light brown.
Thora* alrnost uniformly shiny yellow. Halteres yellow. Legs yellow,
the terminal tarsal segmentsdarkened. Wings yellow, the veins a trifle
darker ; macrotrichia of veins light brown. Venation : Petiole of cellMsa little lessthan three times m,-c'u:vein 2nrl A endingnearlv opposite
the posterior end of rn-cu.
Abdomen yellow. Male hypopygiurn with the beak of the basistyleunusually slencler,especiallyon the outer half, the tip subacute. Outerdististyle with the two arms vory unequal, the flattened outer arm withmicroscopic roughenings on the surface. Basal dististyle a powerfulsimple rod that is expanded on outer fifth into a head that is directed
laterad, the apex obtuse ; outer margin of the head with several strong
appressedspines that are directed outward, these becoming smaller andrnore scattered on the more basal portions. ,lEdeagusrelatively short,
slender,entirely smooth.
Hab. New Zealand(North Island).
Holotype,,j, Swanson,North Auckland, Station 9, .fanuarY 6, 1950
(W.E.Long\.
There are a few other regional speciesof the plngiatus group that have
similar yellow body coloration and unpatterned wings, and in which thebasal dististyle of the male hvpopygium is expanded at its outer end intoa head. The most similar of these include Molophilus (Molophilus)
nronnrius, sp. n., and M . (M.) ,flauidulusAlexander, which differ in alldetails of the male hypopygium, particularly the basal dististyle.
M olophilus (M olo'ph'ilus)quinquespinosus,sP. tr.
group ; general coloration brownish black ;Belongs to the tn,a,crocerus
antennre (male) elongate, nodulose; male hypopygium with the apexof the basistyle produced into a weak point ; basal dististyle unusuallylong and slender,gradually narrowed to the long, acute tip, a short distancellack frorn the latter with a small brush of long delicate setre; surface

Cj.P. Alexanderon Tipulidae{53

of style at' and beyond mid-length with a linear series9f fi ve small blackspines.
Male.-Length about 3.b mm. : wing 4.2 mm. ; antennaabout 2.7 mm.
Rostrum and-palpiblack. A1te1nrelmale) elongate,exceed.ing
one-half_
the lengih of s?ry: brownis_hblack ; flagellat s"gtients strongly"nodulose,
the basaltwo-thirds oval, the apical thiid nu.roiuedinto a glibrous neck :dilated p-ortions o{ s9_Spentswittr \rery long outspreadin[ verticils, thelongest about one-halflonger than the-segm6nt. flead d.a-rkbrown.
Thorax almost uniformly brownish black, sparselypruinoseto produce
a plumbeous ppp-"u-t"tt.", lateral pretergitei pale ; humeral ,"gio., ofprrescutumreddish brown. Halteres dusky, the base of stem nirrowly
yellow. L,egswith the coxre testaceous6rown, the fore pair darker":trochantersobscurebrownishyellow ; remainder of legsbrown^,with d.arker
r-estiture; tarsi darkeningto black : clawssmall. Wings weakly d.arkened,t'he baserestrictedly more yellowed: veins and ntu.rolrichia dark brown.
Venatio! t RrJyilg basadof level of r-m; petioleof eell Msapproximately
one and one-half times m-cru: vein 2nd,,4 long, enclingr6ir" distanc!
beyond the level of m-au.
Abdomen, including l,ypopygiunr, br.ownishbtack. Male hypopygium"with tf" aqeT of basistyle produced into ?r weak point, iom-ewttut
suggestiveof the beak founcl in nrembersof the pl,agiatis
group but withsetreto the tip. Outer dististyle relatively smill, beyon-ilth^eenlarged
basenarrowed and curved to the small slightly e*pandedhead, the htler
further produced into a small point. Basal clislistyle unusually longand slender, gradually narrowed to the long acute tip, a short distanc[back from the latter with a small brush of long delicate setre; surfaceof style at and beyond mid-length with a linear seriesof fir,e small blackspines.
Hab. New Zealand(North Island).
__(o!otyg€,6 , Henderson,North Auckland, Station I , Decemberb, l gbo(W.E.Lons).
The present fly is entirely rlistinct frorn the other known members ofthe macrocerusqro_LlP.It is most similar to speciessuch as Molophilus(Molopthilus) sub_uliferusAlexander, differing in the quite dilstinct\ryopygilm.
Ih", five spinous points on tht basal clistislyle are quite
different frorn the slenderspinoussetrefounclin other speeiesbfthe group.
lI ololiltil us (futololth,iht,s)su,bscuber, sp. n.
Belongs to the t,errninans group ; general coloration brown, theposterior scleritesof the thorax and the pleura more blackened; halteresyellow ; -leg-sobscureyellow, the outer tarsal segments blackened; wings
with a dusky tinge, certain of the veins seamedwith still d.arker; mJe
hyp-opygiym with the outer dististyle relatively slender, the beak only
To-d"tltgly produced ; surface of head with scattered small spines, th;disk with a single retrorse spine ; inner style longer and. *oti slender,
the narrowed apex with microscopicspicules
Ma,le._--I,ength
about 4-4.2 mm. ; wing 4.G-4.8rnrl.

164

0. P. Alexander on liipulidae

Rostrum and palpi black. Antennre black, short ; basal flagellar
segmentssuboval, the verticils greatly exceeding the segments: outer
segmentsmore elongate. Head dark brown.
Thoracic dorsum chiefly brown, the pretergites obscure yellow ,prrescutumdarker brown at anterior end of the central stripe ; postnotumand pleura blackened. Halteres yellow, the knobs brighter. _Legs withthe coxre brownish testaceous; trochanters yellow ; remainder of legs
obscure yellow, the outer tarsal segments blackened. Wings with adusky tinge, the disk and cells beyond cord somewhat paler ; stigmal
region extensively infuscated; certain of the veins less conspicuously
seimed ; prearcular field obscure yellow ; veins brown. Venation :A, lyirg just beyond the level of r-rn; petiole of cell J}/, lessthan threetimes m-cu; vein Zncl -4 ending about opposite one-third the petioleof cell Mr.
Abdomen, including hypopygium, brownish black. Male hypopygiumwith the outer dististyle relatively slender, as compared with termirwns
ot gregensis,the beak only moderately produced; surface of head with
scattered small spines, apparently chiefly marginal in position ; disk
with a single retrorse spine. Inner dististyle longer and more slender,
its basal half more dilated ; apex with microscopicspinulre.
Hab. New Zealand(North Island).
Holntypa,6, Swanson,I{orth Auckland, Station 6, January 27, 1950
(W. E. Long). Paratopotypes,
dl, with the type ; dd, January 14, 1950.
The most nearly related speciesarc Mol,ophihrc(Molnphilus) greyensis
Alexander and A. W.) termi,nnnsAlexandei, especiallythe latter. Bothdiffer in the details of structure of the male hypopygium but are evidently
closelyallied.