J. Ent. (B) +Z (t), pp. r-ro, ro figs. 1974

New or little-known speciesof exoticTipulidae (Diptera). XX
CHARLES P. ALEXANDER
Uniz;ersity of M assachusetts,Amher st, Massachusetts, U. S.A.

SYNO PS I S

Eight new species of the Limoniine genus Antocha Osten Sacken are
described, from Kumaon, Sikkim and Assam, in north India, and from
Kerala, Madras and Mysore in South India.

The preceding part in this series of papers was published in 1973 (J. Ent. (B)
4z (r) :t ro). All speciesdescribed at this time belong in the genus Antocha Osten Sacken andwere collected in various parts of India by Dr Fernand Schmid, to whom I express mv
sincere thanks for the privilege of retaining the type materials.
Antocha (Antocha) aciculifera

sp. n.

General coloration of thorax yellorv, praescutum with three nearly confluent pale brown
stripes, pleura light yellow with a light brown longitudinal stripe; apex of knob ofhalteres light brown; legs brownish-yellow; wings very faintly darkened, unpatterned,
cellfirst M2 long and narrow, exceeding the veins beyond it; male hypopygium withouter dististyle blackened, terminating in an acute point, outer gonapophysis a straight
narrow blade, its apex broadly rounded, inner gonapophysis with a strong basal spine,outer end extended into a needlelike point, aedeagusnarrow, outer end short.
Male. Length, about 5 mm.; wing, 5.5 mm.; antenna,about r.z mm.
Rostrum light yellow, palpi light brown. Antennae of male long, as shown by themeasurements; flagellar segments long-oval, progressively smaller outwardly, terminal
segment about two-thirds the penultimate; vestiture of segments very short and inconspicuous.Head brownish-yellow.
Pronotum pale brown medially, yellow on sides. Mesonotal praescutum with threenearly confluent pale brown stripes, sides narrowly yellowed; scutal lobes, scutellum, andcentral area of mediotergite pale bror,r'n,remainder of mesonotum light yellow. Pleura
yellow, with a light brown longitudinal stripe from the propleura backwards across thedorsal sternopleurite, behind becoming more restricted to obsolete. Halteres with stem
yellow, apex of knob light brown. Legs with fore coxae pale brown, remaining coxae andall trochanters yellow; remainder of legs brownish-yellow, outer tarsal segments slightly
darker brou'n; terminal segment unmodified, claw w'ith a single long basal .pi.r.. Wi.tg.

zCharles P.Alexander

very faintly darkened, unpatterned; veins pale brown, Cu darker' Outer longitudinalveins with .pu.r" trichia, most numerous on distal half of outer sectionof Ra ;5. Venation:
R2 in transverse alignment with r-m; cellfrst M2long and narron', exceeding the veins
beyond it; basal section of Ms only slightly less than the distal section and subequalto
M v a'
Abdominal tergites light brown, sternites yellor,ved,bases vaguely darkened; subterminal segments darker brou''n, hypopygium bror'vnish-yellorn'.Male hypopygium(fig. :) with tergite (r) narrou,ly transverse, both the anterior and posterior borders
ri.i"rify straightl Ouier dististyle (d) blackencd, gently curved at midlength, narrowed
outwarily, teininating in an acute point; inner style slightly longer, outer half broader'Interbase (i) long-oval, apex rounded. Outer gonapophysis(g) straight and narrow' apex
broadly .oundeJ; inner apophysis subequal to or slightly longer, the basal two-thirds,ru..o*, r,vith a strong basal spur, outer end extended into a needlelike point. Aedeagus(a) narrou,, the pointed outer end simple, relativcly short'
Holotype ;f, INore: Chumzomoi Choka, Sikkim, in Rhododendron association,r r , 8oo f eet , 8. v i i . r9 5 9 (F .Sc h m i d ).
Other allied regionalspeciesinclude Antocha (Antocha)attenuataAlexander, Assam;
A.(A.)exilostyla Alexandei. Assam; A.(A)gladiaul sp. n.' Kumaon, and A.(A.)stenophallus
sp. n.; South India, all di{Tering among themselves chiefly in hypopygial structure.
Antocha (Antocha) brevifurca

sp. n.

Belongs to the nigribasisgroup; general coloration of head and thorax yellow, sides of,r"gn"ly more darkened; abdominal tergites slightly bicoloured, reddish-..oritrr-basally, distal half yellowed, sternites ancl hypopygium light yellow; halteresbrown
yellow; legsyellou,,tips of femora and tibiae narrorvlYblack; wings greyish-yellow,costal
yeilbiv, including thc veins; Rs longer than R, vein R2 far basad,to.d..'.tu"r.o*ly
subequal to Rzt-.';,m-ca far before fork of M, in casesto about tu'o and one-half times its
l.ngth; male hypopygium r,vith outer dististyle broadly flattened, slightly expanded
outir,,ardly,apex shallowly notched, the lou'er point smallcr, triangular; interbase abroadly flattened oval Pale blade'Male. Length, about 5-5'2 mm.; wing, 5'5 6 mm'; antcnna, about o'9-r'o mm'
Female.Length, about 5-5'5 mm.; wing, 5'5 6'5 mm'
Rostrum and palpi light yellorv. Antennae lvith scape and pedicel light yellow,
flagellum slightly darker; flagellar segmentsoval, tcrrninal shorter than the penultimate;v.st-titrlr. pale and inconspicuous shorter than the segments. Head yellolv
Pronotrrrn and pretergites clear light yellow. N{esonotum yellow to more fulvous, insome paratypes wiih indications of a slightly darker patt€rn, most evident laterally onpru...rrtrr.n and postnotum. Pleura light yellor'v,in casesthe sternopleurite slightly more
iarkened. Halteies light yellow. Legs yellow, tips of femora narrowly and abruptly
blackened, including about the outer twentieth of segment; tibiae similarly yellow, tips
more narrowly blackened; tarsi yellor,v,extreme tips of proximal segmentsand all of outer
three slightlf more darkened; terminal tarsal segment r,r'ith a slightly developed epicondyle, cla* lo.tg, curved, with a single well-developed basal spine. Wings (fig. t)
greyish-yellow, costal border narrou.ly clearer yellow, including veins C, Sc and R; smallExotic Tipulidae

3brown clouds at origin of Rs, R2, cord, m and m-cu, the included veins darker than theremainder. Costal setae short and inconspicuous; trichia on distal sections of veins
Rs+s, MHz and Ms. Venation: Rs long, about one-half longer than R, R2far basad,subequal to R213; m-cu far before fork of .44,in casesto about two and one-half times its
length.
Abdominal tergites bicoloured, reddish-brown basally, with about the distal half lightyellou'; sternites and hypopygium light yellow. Male hypopygium (fig.
+) rvith dististyles(d) neaily terminal, outer style glabrous, broadly flattened, out$,ardly gently curved andslightly expanded, apex unequally emarginate by a shallow notch, outer lobe rounded,
lower point smaller, triangular; inner style subequal in length, apex slightly enlarged,
with long pale setae from conspicuous tubercles. Interbase (i) a broadly flattened oval
pale blade. Aedeagus (a) very slender outwardly, angularly bent before apex.
Holotype;f,Ixore: swamp Hill, Madras,73oofeet, ro.xii. ry$ (F.schmid). Allotopot2tPe?, pinned rvith type. Paratopotypes,5 4 Q, with types, ro.xii.t96r, z.ii.ry6z (allF.Schnid\.
Other related regional specieshaving the leg pattern, venation and hypopygial structure
generally as in the present fly include Antocha (Antocha) hhasiensisAlexander andA.(4.)prolixistyla Alexander, both of Assam. The most important charactersfor species
separation are in the venation, as the position of m-cu, and in hypopygial characters,
including the outer dististyle. A further species,A.(A.)nysorezsrs sp. n., is described in
the present paper.
The nigriba.re's
group is based on A.(A.)nigribasis Alexander, from Mount Omei,
Szechwan, China (Phil'ippineJ. sci. 49:3go_l, r93z). For a basic discussion of the
Oriental members of the group, a further paper by the writer may be consulted (Phitippine
J. Sci.6r ; t79-8r, 1936).The members of the group are usually small yellow flies witha characteristic venation, vein R2 lying some distance basad of the level of r-m and withm-cuabout its own length or more before the fork of M.The regional members of thegroup are keyed herewith.
r-Legs pale, femoral tips narrowly to more broadly black ...............
)
Legs uniform in colour, brownish-yellow to brown7) Male hypopygium with apex of outer dististyle obtuseaJ
Apex of outer dististyle more or less notched or emarginate.........
)a-) Darkened apex of femur very narrow, subequal in extent to diameter ofsegment. Tibet, Assam.........................yatungensis
Alexander
Darkened apex of femur more extensive, exceeding three times the diameter
4

6

wing pattern darker and more extensive, especially the stigma and cell sc.
Darkened wing pattern very sparse and poorly indicated, especially thestigma. 4ssam...............
. .............
........sparsipunctataAlexander
Fork of outer dististyle very asymmetrical, the upper lobe a very long slencierrod, lower lobe a small subbasalpoint. Assam..............................prolixistyIa
Alexander
Apical fork of outer dististyle shallow, the outer lobe a short obtuse blade,ventral lobe more acute.................
Wings with m-cu more retracted, about tr,viceits leneth before the fork of.M .
brevifurca sp. n.

Charles P.Alexander
Wings withm-cu about its own length befnre fork of M. South India: Mysore
Wings with a weak brown pattern additional to the stigmal darkening,
including an areaat origin of Rs (male hypopygium with outer dististyleforked, the lobes very stout, especially the triangular lou'er lobe). Assam.........
........latifurcaAlexander
Wings without darkened pattefn other than the stigma when this is present ....... ..8,9
8 Thorax with mesonotal mediotergite bror,r'n
IO
Thorax posteriorly more uniformly yellowed,
light brown, scutum yellow,
9 Mesonofal praescutum with disk yellow, margins
sides of lobes brown, mediotergite brown; wings with m-cu nearly its own
postnotalis sp. n.
length before fork of M. South India: Madras...........
Mesonotal praescutum pale brown, lateral margins darker brown, remainderof mesonotum chiefly yellow, mediotergite slightly darker; wings withm-cu less retracted, slightly more than one-half its length before fork of M.
IO

II

t2

r3

.. ... "" " r rMale hypopygium with apex of outer dististyle simple....
""" "" " 12Male hypopygium with apex of outer dististyle unequally forkedagonapophysis
outer dististyle of hypopygium a simple blade, apex obtuse;
pale
entirely
narrow flattened hyaline blade, outer end twisted; thorax
basivena Alexander..................
yellow. Assam
Outer dististyle decurved outwardly, apex pointed; gonapophysis slender,
slightly darkened, the short apex acute; thorax yellow, praescutum with a
Alexander...........scelesta
brown central stripe. Assamevident;
scarcely
Wings virtually unpatterned, the usual stigmal darkening
hypopygium with gonapophysis extended into a long needlelike point'
Wings with stigmal darkening usually more conspicuous; hypopygium with13"" "gonapophysisnot extended into a needle-likepoint ............. . '
lobe
lower
forked,Male hypopygium with outer dististyle very unequallylong, the diameter at base about one-fifth the length of lobe. Assamlongispina Alexander
shallow,
the two lobes
dististyle
Male hypopygium with apical fork of outer
Alexander.............................s1udiosasubequal.South India, Ceylon
Antocha (Antocha) gladiata sP' n'Thorax almost uniformly light orange-yellow; antennae of male relatively long, flagellar
segments long-oval; legs dark yellow; wings pale cream coloured, veins only slightly
daiker, veins Rz and r-m in transversealignment; abdomen yellow, subterminal segments
weakly darkened; male hypopygium with inner dististyle unusually narrow outwardly;interbase slender; inner gonapophysis blade-like; outer end of aedeagusnarrowed.
Male.Length, about 5 mm.; wing, 5'5 mm.;antenna' about r'3 mm.
Rostrum yellow, palpi slightly darker. Antennae relatively long; scape and pedicel
yellow, flagellum dark brown to brownish-black, segments long-oval, vestiture delicate,pale, verticils single, long and dark' Head light brown'

Exotic Tipulidae

5
Pronotum yellow' Mesonotum and pleura almost.uniform
light orange, mediotergite
with a vague slightly darker central line. Halteres.
light y.lloir, apex of knob weakly
darkened' Legs with fore coxae light brown, remaining"coxae
and all trochanters yellow;
remainder of legs darker yellow;ierminal
tarsal segmlnt without epicondyle, claw long,
gently curved, with a long straight basal spine..
wi-ng. p"r;;;;;colour, veins srightiy
darker yellow. sparse trichia on-outer longiudinar*l^'r.onl to M4,more numerous
on distal sectionsof R++s,Ms2 and,Ms.\enation:
R2 and ,_*intransverse alignment;
m-cu at fork of M.
Abdomen yellow, subterminar segments weakly
darkened, most conspicuous onseventh segment' hypopygium yelowed. Nlale
hypoiygi"1ng. 5) with ."ii. Jrr,i.ryi"(d)^narrow, gently..,ru.d, outer harf darkened,
apex subacute; rnner style with outerhalf unusually narrow. Interbase (i) long,
about five times the'diameter, apex obtuse.
Phallosome with outersonayo.lrr.l:s (gj srender, at base with an acute point, inner
apophysis longer and more ulaae-i;ke. ,[.d"ug.r.
(a) with outer end narrowed.
Holotype 6f, INote: Trijugi, Pauri Garhwal,
Kumaon , in Rhoclodendron@/u/'4DruLrdLruil'assocjation
Tooo feet, 26.v.t958 (F.schmid).
Allied. regional speciesinclude Antocha (Antocha)
acicurifera sp. n., of sikkim, andcertain other specieslisted under the latter name.
All oi these have the inner gonapophysisof.the hypopygium with a strong basal spur,
differing u-o.rg rn.-selves in the details ofother structures.
Antocha (Antocha) microcera sp. n.
Size small (wing of maremesonotar praescutum light brown, slightly
4'z-*.);
moredarkened anteriorly, pleura light'yellow;.-ull, outer two segments.antennae
legs yellow; wings pare grey, veins scarcery-r..ydarker, difficult to see, trichia ofl:lglnr"ro;very restricted, m-cu about one-third its rengthb.fo.. fori of M;'erns
t ypopygi.r;*'ith internal organs virtually transparent
ir, .fid.
-rt.
;;;h dististyles
with tips
verylongandslender,aedeagus-o""roflatteneJ,*i,r,""iii.ri;;;;t#;:lt_11-g""rpophysisorpolnts.
-llale. Length, about 4 mm.; wing, +.2 mm.; antenna, about o.g mrn.
Rostrum very light yellow, palpi dark brown,
terminar segment slightly shorter than,the preccding two segments combinecr. Antennae(fig. 6) vJy short, as shown by themeasurements,brown throughout; flagellar
segments"lorrg-orrul, outer two longer, theterminal slightly exceeding the penu"ltimate,more than twice the antepenultimate
segment,both with three.strong setae; proximar
six flageila,
at near midlength*'ith strong black setae that are slightry".g-".rt.
shorter than tEe ..g;-.;;;,
three in number on
the more proximal, two on the outlr pair; flagellar
r.g-.ri."rr"rr
ro twerve, inclusive,,r'ith shorter inconspicuous pale setae.
Head grey.
Pronotum yellow, narrowly darkened
Mesonotar praescutum light brown,-.iirtty.
restrictedrymoredarkenei,,;";;i l;"i^rigl,t y.uo'"; posrerior::::::::
lr:t-fTediallyiclerltesoi
notum
and the pleura light yellow. Halteres yellow,
knlbs weakly'a"rr..".a.
-egs with coxae and tro_chantersiello*, remaining segments slightly darker
yelrow;erminal tarsal segment srightly thickenei
basaily; iu*iorg,
g#ly.rrru.d,
near
base
a-singlelong straight"snile wings pare grey,veins scarcely darker, on sride mounts'ithlifficult to see. Trichia of veins u.r1ir..tri.ied
and scattered, occurring on R beyond

6Charles P.Alexander

origin of Rs and on distal ends of Ra15 and M1a2. Yenation: Rs very long, exceeding R;
R2 and r-min transverse alignment; m-cu about one-third its length before fork of M.
Abdomen brownish-yellow, including hypopygium. Male hypopygium (fig. Z)
mounted on slide and certain of the structures virtually transparent and difficult toanalyse except by manipulation of light under high magnifications. Dististyles (d),interbase (i), gonapophysis (g) and aedeagus(a) as shown. Outer dististyle stout, apex
obliquely truncated; interbase with apex subacute, margin with three small setae.
Gonapophysis sinuous, very long and slender, outer end needle-like. Aedeagus virtuallytransparent.
Holotype 6f, INue: Bhairabkunda, Kameng, North-East Frontier Agency, Manipur,
7oo-rooo feet, 5 . iii. r96r (F.Schmid).
The present fly is best distinguished by the very small antennae, coloration of thervings and their veins, and in hypopygial details. I'he most similar regional species
include Antocha (Antocha) mara Alexander and A.(A.)perobtzsa Alexander.
The speciesdescribed by Brunetti as A.(A.)triangularis and A.(A.)unilineata appear
to represent a single species,each name having been based on a single female. Both flies
are from the western Flimalayas (triangularis, Barogh, near Simla, 5ooo feet; unilineata,
Mundali, Dehra Dun District, gooo feet), each supposed specieshaving been taken on
the same date, roth May, r9ro. I am indebted to Dr A.N.T.Joseph, of the Zoological
Survey of India, Calcutta, for providing excellent figures of the male hypopygium of
w h a t he c ons ider st o b e th i s s p e c i e s(D w a l i , A l mora, K umaon,32oom., r6.i x.1958).
This has the outer dististyle much as in the present fly but with the aedeagusdifferent.
Antocha (Antocha) mysorensis sp. n.
Belongs to the nigriba.szs
group; size small (wing to about 4'5
body and appendages
yellow, thoracic dorsum posteriorly patterned with brown; -*.);
legs yellow, tips of femora
narrowly black; wings whitened, faintly darkened basally, stigma and narrovl'seams atorigin of Rs, cord and m-cu pale brown, including the veins; vein R2 far basad,subequal
to R213, m-culess than its own length before fork of M; abdominal tergites bicoloured,
pale brown, posterior borders narrowly yellowed; male hypopygium with outer dististylelarge, apex conspicuously forked, lower point acute.
Male.Length, about 3'S +'z mm.;wing, 3'8 +'z mm.; antenna,about o'g r'o mm.
Rostrum and palpi light yellow. Antennae with proximal segmentsyellow, outer halfof flagellum very slightly darker; proximal flagellar segments oval, outer ones moreelongate, terminal segment larger than the penultimate; flagellar vestiture very shortand delicate, terminal segment with four strong darkened setae on apical half. Headyellow anteriorly, posterior vertex more ferruginous.
Pronotum and anterior half of praescutum light yellow, the latter with two palebrou'n intermediate stripes that are continued posteriorly on to the uniformly pale brown
posterior half of segment; scutum more strongly darkened, including the central portion;scutellum light yellorv, parascutella slightly darker; mediotergite obscure yellor,v on
central part, sides slightly darker; pleurotergite and pleura clear light yellow. Haltereswith stem light yellow, the large knob weakly darkened. Legs with coxae and trochanterslight yellow; femora yellow, tips narrowly black, on fore legs including about the outereighth, on middle and posterior pairs slightly narrower; tibiae light yellow, tips very

Exotic Tipulidae
Znarrowlypalebrown; basitarsilight
ye'ow, outer tarsalsegmentsparebrown;epicondyleof terminal sesmentpresent,craiv
with u single.pil.-.iii..'i"i"r. wings (fig. z) withbasalcetlsfaintrv darkened,"*",;;;; ;d;abrown; very narrowdarkeningsat origin or Rr,^.orJ-u-o.. -r,i*.,.db.yond*-'),'i"ai"ir"a .ii"ny jyru'aurr..ungof the veins;prearcularfierd srightryinfuscated;"aj.'uuorr.
*i"r'ri;ht-;;"*il;ili
indicated; costa
more yellowed'especiallybefore
and beyona_thestigma.nau"roiri"niaon outer sectionsof veins Ra*s,Mrlz, I[s and,M^.
i;;;i;;:.
Rs long, slightly exceeding
reolngR, R2tar
R2 far basad,sufequal Rr*r; m-culessttrr.r
basad
it. i..rgth b"for. forkof M.
1oAbdominal tergitespale brown, p;.;:ri;r.borders.rurro*ry yellowed,srernitesand
outer two tergites cr:.::e.:hvpopvgi;-;';;;.'n#,j.,r\4are hypopygium(fig' 8) generallyasin bre.z'ifurca,]'elror.r',differingespeciallyi" tr,. l"rir"rlarion"narrow'of the dististyles

tY*.1'r'jffi;r:r::"'stvle

therarge"ilp!' ";rurg.i u'J .*.fi.'"rrry forked,

Holotypei, INore: yo.dpai,Mysore,
2ooo,o,paratopotypes,\. rySg@.Schmid).
ontwopins'nu'utvp.'j, l:.j:

Mulbii",nry.Ji{'soJ-r3oo
reet,r8.r.rese

Lfi,;.[kflr:;'

Among the regional,..y"::1.:-"f.he nigribasisClg.rp,the presentfly is most similar to
Antocha(Antochi) breaifurcasp.
D., difering.i,;a".rtty;rr'"oto*tio.,of the body andwtngs and in detailsof venation
and'hypopygialstructure.
Antocha (Antocha) platystylis
sp. n.
Generallysimilar t"
generarcororarionof praescutum
dark brown, paler
Iatera'y; antennae !Tb?uy!?;oval; knobs'or r,nrt.r., brown; tegs
j:gall;:;;;;,.
brown; wings brownish-grey,
tight
::*
prearcurarn"ra rigl,il.il;,rnconspicuous,
pale brown; veinsR2 undr-*i";i.;;;itransverse
very"k-"m-cubefore fork
arignment,
abdominaltergitesor:*:,sternitesof M:
yelrow,outer segmentsdarker brown;
pygium with outer dististyre
rrypJlbru.k;";l, uper broJdly out.rr.;-ut. a rons
gonapophysis;$iff;:11"i",t#:'thetippla;-'"rterminud.,g"."i"'-.Jr".'a;.'g
ifl".a""g,.pare,Male. Length, about 6.5 mm.;wing, 7 mm.; antenna, about r.z
rnm.
Rostrum and palpi brown, terminal
segment of latter about one-halflonger than the
penultimate, its apex pale.
Antennu. u.-i,u.r; fl_a_geilar
segments oval, the outer ones
smaller, exceeding the sparseverticils in length. Head brown,
grey pruinose.
Pronotum dark brown above,
tight t.o*,r, on sides arrd bzneuth.
Mesonotal prae_
scutum with disk virtualry
by trrree confluent dark brown
stripes, the broadcentral one darker anterioriy, "orr...jsid.. bro',irri.h-gr.y, posteriorscleritesof notum brownish_
grey' parascuteta and anterior
pleurorergitelgiri !.o*r,. pi;;;;'g..yish_brown,
pleurite brownish-ve[ow. Harte;;;il.",;prero_y.ilo*, knob d-kbrown. Legs
wirh coxae
and trochanters ve-lrow,.remainder
or t.g. iight brown,
segments brownish_
black;clawlonga.rasre.,a..,"r,"r'iurrur;ih;'t?;J;r"r"i.pr""."wings brownish_grey,
prearcularfieldlight yelrow,stigmuin.offi.,rous,
irj", pale
brown;vernsbrown,prearcularvetnsand baseof M parer.Trichia

outer sectionof Ms. Venation:illilf;:*'1":{":,frx:

on distil seclionsof Ra15 and,'M'2,more
sparseonRs about one_ha.lflorrg.i inr" n,Rz pale, in virtual'ho't"ihu; ;;section;ii^;;;";'cu about
one-thirdits8Charles P.Alexander

Abdominal tergites brown, subterminal three segments darker brown, sternites andhypopygium more yellowed. Male hypopygium (fig. q) with tergite (l) narrowly transverse, posterior border very shallowly emarginate, internal scaffolding of pale conspicuous. Outer dististyle (d) blackened, boomerang-shaped, tip broadly obtuse. Interbase(i) long-oval. Gonapophysis a long slender arcuated spine, tip acute. Phallosome(p) withapex of aedeaguscurved, the subtending plate pale to nearly hyaline outwardly, outerangles extended into acute pale points.
Holotype J, INoIe: Swamp Hill, Madras,73oo feet, ro.xii. ry6r (F.Schmid).
The most similar regional species are Antocha (Antocha) amblystyla Alexander andA.(4.)perobtusa Alexander, which have the outer dististyle of the hypopygium generally
similar in conformation, differing among themselves in other structures, including thephallosome.
Antocha (Antocha) postnotalis sp. n.
Belongs to the nigribasisgroup; size medium (ri,ing to 5.5 mm.); general coloration ofbody light yellow, mesonotum characteristically patterned with darker, including thedark brown postnotal mediotergite; legs yellow; wings nearly hyaline, stigma darkbrown, vein R2 basad of level of r-m, about two-thirds R21s,nt-cu about its own length
before fork of M; abdomen light yellow.
Female.Length, about 5.2 5'5 mm.; wing, 5.+ 5'5 mm.
Head, including rostrum, light yellow, palpi pale brown. Antennae with scape andpedicel yellow, flagellum pale to darker brown.
Cervical region and pronotum pale yellorn'.Mesonotal praescutum lr'ith disk yellow,
bordered in front and on sides by light brown, humeral region clear yellovi'; scutum
yellow, lateral third of each lobe brown, this being a posterior extension of the praescutal
darkening; scutellum pale yellow, parascutella with two small brown spots; postnotal
mediotergite conspicuously dark brown, pleurotergite yellow. Pleura uniformly clear
light yellow. Legs yellow, outer tarsal segmentslight brown. Wings nearly hyaline, baseand costal region light yellow; stigma dark brown, conspicuous; veins of anterior border,
including Rs and R3 yellowed, remaining veins brown. Outer section of Ra15 and outer
medial veins with trichia. Venation: Rs longer than R; R2 basad of level of r-m, about
two-thirds R21s:,m-cu about its own length before fork of M.
Abdomen clear light yellow, especially the basal sternites. Ovipositor r,vith cerci very
long and slender.
Holotype Q, INore: Senbaganur,Madras, 58oo feet, 8.xii. r96r (F.Schmid).
The related and generally similar species have been included in the key to theregional members of the nigribasis group provided earlier under the description of
Antocha (Antocha) brevifurca sp. n. The most similar such species is A.(A)perstudiosa
Alexander, of Nepal.
Antocha (Antocha) stenophallus sp. n.
General coloration of thorax brownish-grey, anterior half of praescutum with a slightly
darker central stripe; legs brown, posterior coxae yellow; wings faintly tintecl, basenarrowly yellow, stigma dark brown; male hypopygium with outer dististyle narrowly

Exotic T'ipulidae

O*PtrA
t------\

\i.

r 'b

/

lts A

D) / I ' \)'' '/'r'
,7

'%\

r0
Figs. r-ro. (r, z) Venationof: (r) Antocha (,;#3';i,'f;t;'ijnT:i:tii,
T,_,"i+,1,,::;1i,;,"1,,;i"lii,ff

i##y:"i^ti,:i?"'::i:!!.:?":*:#";*i:l:l:"'ii':;,Ji

loCharles P.Alexander : Exotic Tipulidae

obtuse at apex; both gonapophysesslender, needle-like, inner apophysis with a strong
basal spur; aedeagusvery narrow, apex obtuse'
Male. Length, about 4'5 5 mm.; wing, 5-6 mm. ; antenna,about r'z-r'3 mm.
Female.Length, about 5'5 mm.; wing, 6 mm.
Rostrum yellow, palpi black, terminal segment elongate, subequal to the preceding
two combined. Antennae brownish-black to black; flagellar segments oval, verticils
small and inconspicuous. Head brownish-grey.
Pronotal scutum brown above, yellowed on sides, scutellum yellow. Mesonotal
praescutum almost uniformly brownish-grey, anterior half with a slightly darker central
stripe; scutal lobes dark brown, median areayellowed; scutellum light brown, parascutella
yellowed; mediotergite brown, anterolateral parts and the pleurotergite more yellowed.
Pleura chiefly brown, yellowed posteriorly. Halteres with stem yellow, knob dark brown.
Legs with fore coxae brown, mid-coxae paler brown, posterior pair large, yellow;
trochanters yellou'; remainder of legs brown; claw with a long basal spine. Wings very
faintly tinted, base narrowly yellow, stigma small, pale brown; veins brown. Longitudinal veins beyond level of stigma with trichia, including all outer radial and medial
veins. Venation: Rz in nearly transverse alignment with r-m; m-cu shortly before forkof M, in extreme casesto about one-half its length'
Abdominal tergites and hypopygium dark brown, basal sternites more brownisor
yellow. Male hypopygium (fig. ro) with tergite (/) transverse,unusually narrow, poster-hi
border very slightly emarginate. Ninth sternite (not figured) extensive, semicircular inoutline. Outer dististyle (d) darkened, slender, curved gently to the narrowly obtuse apex;
inner style slightly longer. Interbase (f) relatively long. Both gonapophyses(g) slender,
needle-like, the longer inner pair with a strong basal spur.
Holotype 6f, INnte: Bodimettu, Madras, 2ooo-3ooo feet, z4.i.tg6z (F.Schmid).
Allotype Q, Munnar, Kerala, 35oo feet, r5.xii. ry6t (F'Schmid)' Paratyper d ?, with theallotype; j, Swamp Hill, Madras, 73oo feet, rr,xii. r96r (F.Schmid).
T'he most similar regional species are Antocha (Antocha) attenuata Alexander andA.(A.)exilistyla Alexander, both from Manipur, all three species differing in details ofhypopygial structure.
Manuscript received zSth August, r973