35t/

SOCtEtv,
fronr BUT,LETINOF THE BnOOrr,YN EuTOUOI,OGICAL
Reprinted
Vol. XXVI, No. 4, PP. 177-184. October, 1931.

NEW OR INSUFFICIENTLY_KNOWN CRANE_FLIESFROM THE NEARCTIC REGION (TIPULIDAE,DIPTERA). PART III.
B v C H IR L E S P . A l .r xANDER, Am her st, Mass.*

The precedingparts under this generaltitle were publisheclin
r93o ( Bur-r-.Bnoorrvx E,xr. Soc., z5 : 7r-77 ; 276-z8z). The
types of the speciescollectedby Dr. G. C. Crampton and thein my collection. The type of.Litnonio swb*iit.t are preservecl
apicatahas beenreturnedto Dr. J. SpeedRogers. The very interesting seriesof Tipulidae collectedby Mr. Owen Bryant in
British Columbiaand Alberta has been returned to Mr. Bryant.
entomoloMy sincerethanksare extendedto the above-mentioned
gists for their continuedco-operationin making known the cranefly fauna of the Nearcticregion.
Dolichopeza (Oropeza) tridenticulata n. sp.
trlale.-Length about ro n-lffI.;wing rI lrllrl.
Describedfrom alcoholicspecimens.
Closelyrelatedto obscotra,differing especilllf in the structure of the rnalehypopygium. Antennaedark brown. Mesothe pleura still darker. Legs with
notum dark reddish-bt-or"tt,
the tarsi a little paler than the tibiae. Wings suffusedwith
brown, the oval stigma slightly darker brown; pal-erareas
before'andbeyondthe stigmaaud acrossthe baseof cell rst
M.. Abdominal segmenft brownish yellow, conspicuously
ringed with dark brown on the incisures,oo_thesternitestheof the segmentsmore broadly darkenedthan the apices.
bas"esMale hypopyglumwith the central portion of the tergal margin produied-into a small rectangular area that bears three
imall chitinizeclpoints; lateral arms of tergite evenlyroundedat tips. Inner dististyle very broad, weakly bidentateat tip,
otr. bf the points being a small blackenedspine. Outer dismuch as in obscura.
tistyle and gonapophyses
H abitot.'Massachusetts.
Holotype; alcoholicJ, Goshen,HampshireCo., July r' r93r
(G. C. Crant,pton).
Dolichopeza(Orope;a) tridurticulata is most closelyallied toin theD. (O.) obscrro (Johnson),clifferingvery conspicuously
detailsof structureof the rnalehypopygium,notably the medianx Contribution from the EntomologicalLaboratory, MassachusettsState College.

178 Bull,etin of the Brooklyn EntomologicalSociettl Tot.xXTI
toothing and obtuselateral arms of the tergite,and the very broacl
inner dististyle. Besidesthe type, a few additionalfemaleswere
tal<en. Dr. Crampton reports the speciesas being verv cornmonina rlarkeneclshecl.
Limonia (Limonia) subapicata n. sp.
Belongs to the apicata group; mesonotal praescutum polished yellow, with a broad black median stripe; lateral margiqs of praescutumblackened;wings grayish subhyaline,thestigma long-oval,pale brown.
Male.-Length about 5--5.5mm.; wing 5.2-6 mm.
Antennae black throughout ; flagellar segmentsoval, withverticils that slightly exceedthe segments. Head black, very
sparselypruinose.
Pronotum dark medially, paler laterally. Mesonoturnpolished yellow, with a conspicuousmedian black stripe that iswidenedat the suture; lateral margins of praescutumbehindthe pseudosutural foveae conspicuouslyblackened; scutal
lobesblackened;median region of scutum and the scutellum
rnore brownish testaceous; postnotal mediotergite darkbrown. Pleura testaceousbrown, darkest on the anepisternuln, and, in some cases,the ventral sternopleurite;in othercases,the latter region is distinctly yellow. Halteres pale,theknobs weakly darkened. Legs with the coxae and trochanters yellow; femora black, the basesyellowish; remainderoflegs black. Wings grayish subhyaline; stigma long-oval, pale
brown; no darkening in outer radial field; veins dark brown.
Venation: Sc, ending about oppositethree-fourthsthe lengthof Rs, Sc, closeto its tip; Rs weakly angulatedat origin; f ree
tip of Sc, anclR, in nearly transversealignment; m-cu at orcloseto fork of M.
Abdominal tergites brown; basal sternites yellow, theouter segments dark brown; lateral line narrowly darkbrown; hypopygium brownish yellow. Male hypopygiumvery much as in apicata, differing only in details. Lobules
at proximal enclof basistylevery similar in both species;subapical lobeson same face having setaeand spinesmuch lessdeveloped,the elongate paired spinous setae of. apicata here
being a single, small, gently curved spine. Outer margin of
dististyle with the crest of blackeneclspines low and depressed.
H abitat: Floricla.
Holotype: 3, Gainesville,Alachua Co., May ro, r93o (1. S.
Roqers); Collector'sNo. 6++. Porattpr.t.'several33.

Oct.,7e31Bultetin of the Brooklyn Entomological,Society 179
The closestally in the apicata group is l. (L.) apicata (Alexander) which differs in the pattern of the tnesonotum,the strongly
infumed wing-tips, and the details of structure of the male hypopygium. The fly is entirely different from the only other regional
member of the group, .L. (L.) rogersiana Alexancler.

Tricyphona rubiginosa n. sp.
General coloration reddish brown, sparsely pruinose; antennae black throughout, r6-segmented; wings wilh a -{e9nfulvous tinge; venation of radial field very variable, cell R-,
shortlpetiolate to broadly sessile; cell M, 1t1,sent; cell
being
"open
IlI,
by atrophy of m'; male hypopygium with two dististyles.
iIale.-Length GZ.S mm.; wing 6.5-Z.Smm.
Female.-Length
Z.S mm.; wing about 7 mm.
Rostrum and palpi black. Antennae black throughout,t6-segmented,the segmentsshort-oval, with short verticils.
Head brownish gral'.
Mesonotum reddish brown, sparsely pruinose, the praescuturu with a slightlv darker median brown stripe ; a deep
pit on parascutellJon either side of scutellum. Pleura reditith brbwn, overcastby gral'. Halteres pale yellow, the large
knobs weakly inf uscatecl. Legs u'ith the coxae reddish S?y itrochanters yellow; remainder of legs brown, the terminal
segrnentspaising into dark brow-n. \\'ings relatively narrow'rviih a cleip fulvous suffusion, iridescent,the stigmal region
lnore suffusecl; veins deep yellow. Venation : St. ending
shortly bevonclthe level of- cord, Sq, a distancebefore originof Ri gr.it.r than the length of the latter vein; forking ofIis varlaltle, in some caseswith a short Rr*r*nthat is a little
shorter than the basal section of Rs, in other cases,even on
the two rvings of a single specimen, cell Ru lryrowly topresent;
broadly sessiiebv the obliteration of Rr*"*n;-cell /'1u1,
cell lll" open b1-the atrophy of nr,; nx-!Nt,oblique, more than
one-half its length be1'onclthe fork of,IW.
Abdominal iergite! brou'nish gra!, the sternites morebrownish yellow, the caudal margins of 1!re segmentsinfuscated, ott lhe subterminal segmenisincluding the entire sclerite ; hypopygium obscure fellorv. Male hypopygium relatively large, the tergite broacl, its caud-allobe transverse tot'.ty feebly'emarginite; lateral arms of tergite pale, at ends
irregularty Uina, the principal arm a po\\'erful_curved point.
Basistyl. "t "p.* ptohuced laterad into a slender lobe. Dististyles two, ; longer foot-shaped_lobe, in addition to theusual oval lobe set with abundant short spinous setae.

180 Bulletin of the Broolilyn Entonr,ologicalSociety Tot'xxrr
Habitat : British Columbia,Alberta.
Holotypt: 3, Laggan,Alberta, Jtly t6, rgzS (Owen Bryant) ;Collector's No. B-zB-5. zlllotype: l, Ptarmigan Pass, Laggan,
Alberta, altitude 6ooo-7ooofeet, July 23, r9z8 (OzuenBryant).
Paratopotypes:2 33, with the type ; paratype: r {, Hector, British Coltrmbia,on dry recl rock, July r 5, ry28 (Ozuen, Brvant) 'Collector'sNo. 8-28-1.
T'ricyplt,ona
nftiginosa is very clifferentfrorn all tlescribedspecies of the genus. The venation of the raclial fielcl is ttnusuallv
plastic, and, if this were neecled,furnishes additional evidence
that the interpretationof this fielcl as given b)' the writer in recent papersis the correctone.
The very excellent fielclnotes taken b1' x{r' Brvant are given
pertainlikew.iseto Ornithodesharherewith. Theseobservations
rimani Coquilletand Ph4tllolabisbryontianan. sp.
" B-28-t. Hector, B. C.,.|uly 15, t928. On face of rock-cut
beside road betweenWapta Carnp (and Lake) ancl high bridgeover outlet to lake. Mostly in hollows in rock and where rock
was actually wet. Many actually in behind falling water so that
one had to reacha tube in through the water to get them. Speckled-wingedonesmostly on dry mossor on clry rock. This species(Limonia sci,ophila)and the redrlish ones u-ith copper)/wings(fricyphona ntbigino.fon. sp. anclPln'llolabisbryontianan. sp.)rocl<acrossthe brirlge.u'ith springs
occttrredmore colnlnonl)'ot-t
trickling clownover it. The three large spottedones (Ornithodes
harrinruni Coq.) rvereon this rock u,herethis u-asoverhttngseveral inches by moss ll'ith n'ater dripping frorn it. A feu- aboutpier-head of bridge. The redtlish ones occurre(l on this rockwhere this was dry and showeclan especialfondnessfor rock u'ith
a good deal of red color in it, which made them very hard to see.
All thoseon wet rock, with the three Ornith,odesmentioned,\;erequite easily caught by putting the tube over them. The red ones
on clry rocks were more Scary. About 4 P. M. rnostlv in sharleand some in quite dark recesses,the reddish ones more in thelight. Some on vegetationabout pier-head,perhaps after being
clisturbed." Other Tipulidae under this number include<lLintonia( Dicranom,yia..'
morioides ( O.S.), Elliptero astiqnwtica Alex.,
Tricyphona aperta Coq., Dicranota (Rhaphidolobis\ subsessilis(Alex.), Molophilus colonusBergr., anrl Rhabdomastir(Sacandop1a)subcau,dataAlex.
" 8-28-.5. Laggan,Alberta,
July 16, rqz8. Tipulids, etc., on

oct.,7e37 Bulletin ol' the Brooklyn Entomological Society 181
face of rock and in large holes in rock face. Clitr at southwest
enclof Lalie Louise, altitude 57aofeet. Part of thesetaken roo
feet abovelal<eat top of slide where water is flowing down and alarge cavern is made by " large slice of rock which has separated
from the main body and leansagainstit. Others lessthan ro feetabove lake. Rocks quartzite, red, purple and various colors,
stainedin placesrvith iron (and copper?). Ornith,odesseemto
be foncl of getting into dark places,all being found in big holes,incomparativedarkness,4-6 P. M., dull day, sun behind mountain."
Besiclesthe records for Ornitlt,odesharrimoni and Tricyphonanftiginoso, Mr. Rryant had included under this numbet Limonio(I-imonio\ scioplilo (O. S.), and Elh,pteraastiqmaticaAlex.
Dactylolabis hudsonica n. sp.
General coloration brown, the praescutal stripes very illdefinedand inconspicuousagainst the ground-color; femora
light brown, the tips paling to yellowish; wings with a heavy
giayish brown pattern, arranged as in the rnontana gro]rq;
beyondthe forkno supernumerarycrossveinin cell R ; nx-cul,of Ii, in most iares the distance more than one-half thecrossveinitself ; male hypopygium with the basistylesdark
brown.
Atale.-Length about 6.5 mm.; wing 8-8.S mm.
Fenrale.-Length about 6.5-Z ffiffi:; wing 7.54 mm.
Heacl dark, coveredwith a yellowishgray pollinosity.
Mesonotal praescutum-ill-defined, brown, sparsely p-ollinose,-theespeciallythe lateral Ppir; interbrown stripes
nrecliatestiipes becoming divergent.to nearly obsolete on
cephalic poition. Pleura more. pruinose, !9gt .with thef"nlo.a tight brown, the tips paling to yellowish; tibiae andtarsi brorvir. \\iings"in with a heavy brownish -gray,pattern,thethe montana group. Cos!1l fringe andareas arrangeclas
vestitureof'veins consistingof deliiate setae. Venation: No
supernullteraf\:crossveinin cell R"; m,-cabeyondthe.fork of
I,l', in most specitnensthe distance more than one-half thelength of the crossvein.
Abdo-.n dark brown, the caudal margins of the sternites
pale. Male hypop)'giur1with the basistylesdark brown.
H abitat: Quebec ( Gasp6) .
Holotypa:-Perc6,June 28, rc:.3r(C. P. Alerander). Allotopot\tPe:i, June 30, r93r. Paratopotypes:4 3?, June z8-3o, r93r.
The specimenswere not especiallylithophilous,as is the casein other membersof the nrontanagrotlp. Most of the individuals

L82 Btclletinof th,eBrooklyn EntomologicalSociety vot'xxvl
were swept from rank herbagegrowing in the partly dry bed of amountain stream.
Phyllolabis bryantiana n. sp.
General coloration reddish ; antennae black ; wings with astrong brownish yellow suffusion, the oval stigma- a trifle
darker; male hypopygium with the appendageof the ninth
flattened, profoundly bifid ; inner dististernite very bio-ad-,style a simple clavate rod; gonapophysesYery elongate,
cbrci of ovipositor slender,with entire margins.
stylet-like;
Il'Iale.-Length about 6 mm.; wing 7.5-8 mm.
Female.-Length about 7.5 mm.; wing 8 mm.
Rostrum and palpi black. Antennae dark brown to black
throughout; flagittar segmentsoval, gradually decreasingin
sire o"utwardly.-'Head iiark gray; anterior vertex relatively
narrow.
Pronotum testaceousbrown. Mesonotum brown, the praescutum darker brown medially in front; no tuberculate-pits;
yellowish. Pleura
pseudosuturalfoveae pale; scutellu1 m_oreHalteres pale, the knobs dusky. Legs
itti.tty reddish yellow.
-andtrochantersyellow ; femora yellow' passwith ih. .o*"eing into brown at tips; tibiae and tarsi passingthrough brownto"clarkerbrown. iVittgs with a stronf brownish yellow suffusion, the oval stigma i ttifle darker ; veins yellowish brown.
Venation: 5'c, endlng about opposite one-third the length of'trx-cxt
at or
Rr*r*o,Sc" lear its tip; Rs shorter than Rr*r*o;
ltefore the forl< of /11.,* o.
Abclominal tergitei clark brown ; hypopygium black, theyellow; sternitestestastyli and appendageof ninth.sternite_.Jo.t, yellorv. Mile hypop)'gium with the outer apical angleof basistyleproduced into a stout lobe that is about as long as
the outei cliitistyle. Outer dististyle a bilobed structure,-theouter lobe e*p"nd.d and tmncate at apex. Inner dististyle asimple pale cjavate rod, with abundant erect setae over theentire surface except the inner or cephalic margin. 4gqqqclageof ninth sternite very broad_,flattened,profou".dly bifid.
long and slender,
Stylet-like spinesof the gonapophyses-yery-slendeinessin thisall other sp-ecies
exceedingi; length andfaunal rEgion. bvipositor with the tergal- valves (-cerci)
relatively lottg and slender, nearly straight, the ventral margin untoothecl.
Habitat; British Columbia.
H olotype: Hector, on dty red rock, Jrly r 5, - ry2? (Owen
Bryant) ; Collector'sNo. R-28-r . Allotopotype: J. Parotopo'
types:q d?.

oct.,1e31Bulletin of the Brooldyn EntomologicalSocietg 183
PlryJls\abisbryantiana is named in honor of the collector, M..
Owen Bryant. The fly is very different from all other described
speciesin the Nearctic Region. The conditions under which the
rubiginosan. sp.
lly rvas tal<enare discussedunder T'ricS,pltona
Phyllolabis lagganensis n. sp.
General coloration dark brown ; wings tinged with gta! i
costal region of male dilated before stigma i Rr*"*nshort, notexceeding one-half of R, alone ; male hypopygiurn with theappendage of the ninth sternite very large, prow-shap-e.d!
bisistylei complicated by an unusual development of apicaland mesal lobes; inner dististyle much larger than the outer;
gonapophyseslong and slender.
Xlale.-Length about 6 mm. ; wing 6.8-Z mm.
Rostrum an-d palpi black. Antennae of moderate length,
if bent backwarcl-notattaining the wing-root, brown throughout; flagellar segmentsoval to long-oval,with numerousverticils that slightly'exceedthe segments. Head gray.
Mesonotum dark brown, with a sparse gray pruinosity, thepleura paler, especially the dorsal portions and the ptelopleurite. Halteres vellow, the l<nobsinfuscated. Legs withthe coxae ancf trochanters veffow; remathdel ot' fegs fitght
brown, the terminal tarsal segmentsdarkened. Wings tinged
with gra! t the oval stigma slightly darker ; veins brown.
Costaf region just before stigma strongly dilat_ed,this presurnablyJ tp.ii"l character of the male sex. Venation r Sc,
encling-aboutopposite midlength of Rr*r*o unusually short,about'bne-halfR-, alone; inner end of the relatively short andu'irle cell rs/ ll1. arcuated.
Rasal ahclominaltergites blackened,the intermediate tergites ltron'n, the subteiminal segmentswith a conspicuous
irlacl<subternrinalring. Male hypopygium large and unus]lally, complicateclin siructure. -Appendage of ninth sternite.'.ty large, in rlriecl specinrensforming-a-prow-*"pgd structrlre, on slides appearirlgas flattenedblades,each with a lateral lobule as base. Rasistyleproduced into lobes,the mostconspicuousbeing three on apical and lateral -portions,alq
three others on niesal face. Outer dististyle relatively small,beal<-like. Inner dististyle a very large flattened blade, theapex long-produced. Gonapophysesvery long and slender,much "r itr bryantiano but stightly shorter and more slender.
Habitot: Alberta.
Holotlpe: 3, I-aggan, July r(1, rg28 ( OzvcnBryran|\. Paratop o t y p e: J .

184 Bul,letin of the Broolilyn Entomologi,colSocietg Tot'xxvr
Phyllolabis lagganenns is very different from the other known
speciesof the genus (eight recent,one fossil in Baltic Amber), allof which are availableto me for comparison. The charactersof
the male hypopygium in the present speciesare unusually complex and difficult of description. The shortnessof Rr*r*ndistinguishesthis speciesfrom all other describedNearctic species,but
is approached by mocrura ( Siebke) of northern Europe and by
two Himalayan speciesdescribed by the writer. Whether the
curious dilation of the costal region is a sexual characteronly remains to be discovered. The genus Phyllolabis is new to the
Canadian fauna.