IndianolWestspcieslfew0rlittleknown(Drptera)
IIl.
Iipulidae

GhulaP,Aloranda

Reprintedfrotn

Ricool Pucrto
ol thcUaiversity
ThcJounelol Agriculh,rel/ol. XXI,

No.4-Pages 521 to 534.
Octobcr, rg77.

agAN JUAN, P. R.
Bunneu or Suppr,tns, PntNTrNG,exo fhnxsPoararroN
198?

NEW OR LITTTE-KNOWNSPECIESOF WEST INDIAN
TIPUTIDAE (DTPTERA).rrr.
By Cnenr,us P. Ar,nxelronn,llfassachusettsState College,Amherst, Massachusetts

The seeondpart unrlertlris g'cueri,ltit e appcurcrlin this Jotirnal.
vol. XXI, No. 2, pages 179-190. At this time I rvish to discuss
some unusually interesting eolleetionstaken in Oriente, Cuba, andsent to me through the eontinuedfriendly interest of Director Juli6n
Aeufla ancl l\[r. LawreneeC. Bruner. Through the hindnessof thetwo latter seientists,I am privileged to retain the noveltiesherein
diseussedin my eolleetionof these flies. The speeimensrecordetl
herervithrveretaken chiefly on trvo eolleetingexcursionsto the mountainoussectionsof Oriente: Loma clelGato,Sierra clelCobre,altitude
2600-3325feet, September25-30, 1935 (Acufla, Bruner and Scaramuzza). Pieo Turquino, Sierra l\'Iaestra,altitude 1650-6000feet,in June 1936 (JuliSn Aeufia and P. J. Darlington,J".). The majori[' of the latter were taken at light, Cuevadel Aura, a]titude 3750
feet, on the night of June 72, 1936.
Dolichopeza (Megistomastix) devexa sp. n.
Size large (rving, 10 mm. or more); general coloration clark brown; antennae (male) nearly as long as bocly, the pubescenceand verticils of the flagellar segments short; wings with a brown tinge; stigma oval, dark brown;
abunrlant trichia in cells of outer half of *iog; male hypopygum with lateral
portions of tergite not produced.; basistyle extencleclinto a slender, pale am,
the tip set rvith strong blaek spines.
ftale.-Length about 10-11 mm.; wing 10-11 mm.; antenna g-10 mm.
Female.-Length about 10 mm.; wing 10 mm.
Frontal prolongatiou of heacl, together with the palpi, black. A:rtonnae(male) nearl"vas long as the body; seapedark brown; pedicel yellow; flagellar
segmentsblaek, the more basal ones with the ineizures very restricteclly pale;segments loug'cylindrical, l'ith a short erect pale pubescencel vertieils relatively short and inconspicuous,about two or two antl one-half times the lengtb
of the pubeseence.Head grayish brown.
l\fesonotal praescutum and seutum dark brown, the former with a capillarydarker median vitta; posterior sclerites of notum paler brown. Pleura chiefly
d.ark brown, the posterior sclerites and pleurotergite paler. Halteres d.arkened,
the base of stem restrictedly paler. fregs with the coxae dark I troehantersobseureyellowI remainder of legs black, the femoral bases restrictetlly obscure'W'ings
yellow.
(Fig. 1) with a brown tinge; stigma oval, dark brown; restricted dark seams on anterior cord rn-aut; veins and macrotrichia dark.
Abund.ant trichia in all outer cells of wing (intlicatetl in ffgure by stippling).
5n

62L

OF TIIE UNMRSITY Of'' P. R.
TIIE JOURNAITOF AGRICUIJTURE

Venation: ^Sc, entling oppositeorigiu of Rsi R 'a , atrophietl or (as figuretl)
representedby a basal spur.
Abdominal segments obscure brorvnish yellow, the caudal portions of tlre
segnelts darkerI seeoud.tergite lvith a dark ring at near mictlength; hypop;gium chiefly pale. Ilale hypopygiuur (Fig. 9) rvith the caudal margin ofi.tgitu, qt, subtransverseor onl;. gently eonvex, the lateral encls not procluced'
into lobes. Basist;'le, b, prod.ueeclinto a slenclerpale armr the apex set with
abundant blaekeueclspines. Outer clististyle, otl, elongate, strongly angulatecl
at near nrirJlength and here prod.ueeclinto a setiferous lobe. Eighth sternite
n ith eauclal rnargin prod.ueerlinto a tongue-like metlian lobe.

Habitat.-Cuba (Oriente).
Ilolotype, d , Pieo Turcluino, Sierra llaestra, alt'itude 6,000 feet,
Jnne 20, 1936 (Acufla). Allotopotype, I altitude 5100-5200 feet,
June 12, 1936 (Acuf,a). Paratopotypes, 1 A , altitude 6000 feet,
June 12, 1936; 1 A , altitude 3750 feet, June 12, 1936 (Acufla).
Dolichopeza (Xlegistomastir,) cleuera is ver1- clifferent from theother regional specles in its large size and structure of the malehypopygium. Both of the Cuban specieshitherto made known arey€ryr small, u-hereas the present species rivals in size the largest
Puerto Riean memlter of the subgenuscliscoveredto this clate.
L'imonio ( Discobola)gowcleyiAlexander.
Pieo Turclrtino,altitucle 3750 feet, June L2, 1936 (Acufla).
This specieshas tltree tlark femoral rings.
t\lexander.
Linronio ( Discobola) glotccleyz
Pieo Turquino, A. Naranjo, altitude 1650 feet, June 12, 1936
( Acufla ) . Loma clel Gato. Sierra d.el Cobre, altitutle 2600-3325
feet, September 2ir-110.1935 ( Acufla, Brttner & Scaram\zza).
Lintonio ( I'intont,u) eari'baeaAlexander.
Loma clel Gato, Sierra clel Cobre, altitude 2600-3325 feet, Seplember 25-30, 19:i; (Acuita. Bruner & Scatamrtzza).
Limoniu, ( I.,i'ntonia) hoffntani Alexancler.
Loma clel Gato, Sierra del Cobre. altltucle 2600-3325 f eet, September 25_-90,19;i5 (Aeuiia, Brttner & Scaramtzza). The species
hacl not been recorcleclfrom Cuba.
(Alexancler).
Lintontu ( Rhipidiu) schwa.r"z'i
Pieo Tlrquino. Sien"a l\Iaestra, altitucle 1650-3750feet, June 12,1936 (Aeuiia). Lorna clel Gato. Sierra clel Cobre, altitucle 2600-3325
feet, September 25-:10, 1935 (Acufia. Brttner & Scaramuzza).

NEW OR LITTLE KNOWNTIPULIDAE III

626';,.
Limonia ( Dicranomyia,)retiauluta (Alexancler)
Pico Turquino, Sierualfaestra. altitude 3?50 feet, June 12, 1936
(Acuiia) . Loma del Gato, Sierra clel Cobre, altitude 2600-3325.
feet, September 25-30, 1935 ( Acufla, Brttner & Scaramuzza)
f ,futonia ( DtcranomA'ta) trinitatis AlexanderPico Turquino, Sierra llaestra, altitude 3750 feet, June L2, 1935
(Acufla). Loma del Gato, Sierra del Cohre, altitude 2600-3325
feet, September 25-30, 1935 (Acuiia, Rrttner & Scaramuzza)
Alexander.
Linr,onia ( Geranomgia.)myersia,na.
Pico Turquino, Sierra }Iaestra, altitucle 1650-3750feet, June 12,1936 (Acufla).
Or[ntargu( Dtotrcphu) fluricostu(Alexander).
Pico Turcluino,Sierra Jlaestra,altitutle 3750feet, June 12, 1936
(Acuira). Loma clel Gato, Sierra clel Cobre, altitnde 2600-:3325.
feet, September 25-30, 1935 (Acuffa, Bruner & Scaramuzza)
Thesespecimensare all smaller than the type series(BIue Mountains, Jarnaica.)but agree in all other essentialsancl surely appear
to be eonspecific.
E Ttiltln'ugnte( tr pi ltltrag mu) cubcnsis A lexander.

Pico Turclttino,A. Naranjo, Sierra Jlaestra, altitucle 1650 feet,
Jture 12, 1936 (Acufia). Loma del Gato, Sierra de1Cobre,altitudc
2600-332irfeet. September25-30, 1935 (Acuira, Bruner & Scar*ntrrzza). Los Llanos, Oriente }Iaisi, Februarl' 5, L929 (Acufla).
BuenosAires, Trinitlacl llountains, altitucle 2350-2800feet, May 3,
f 9 3 2 ( B r u n e r& O t e r o ) .
Shannonomyia phragmophora sp. n.
Gerreral eoloration clark brorvnish gra)'1 haltercs ltale yellow throughout I
legs lellou', the tenninal tarsal segments clarkenecl; w-irrgs Iight yellol', witha heavl'brorvn pattem that is restrictetl to the vieinity of the veins; .Rs lot,eonrreetedrvith vein M by a supernurnerary crosst'ein in eell Rr' -8, shortly b+
beyonc l i t;
ce l l 7 st M, elongate, exceeclingvein 14 r *,f o r e f o r k o f l l , *n ibasal seetion of M, not in transverse alignment with m., areuatecl to rveakly engulateil l abtlomen clark brorvn, hypopygiurn brighter.
IVlale.-Length about 5-5.5 nrm.; rving 5.5-6 mnr.
Female.-Length about 6.5-7 mllr.; rving 6-6.5 nlnl.
Rostrum and palpi blaek. Antennae short; basal segments yellow, the outen
basal flagellar segrnents subglobular tohalf of flagellurn passing into brol'n;
short-oval, the onter segrnents s)ightly longer. Heacl brorvnish gray.

626

TIIE JOUNNAIJ OF AGRICULTURE OF TIIE UNIVEBSITY OF P. R.

Mesottotum dark bron'nish grol the praescutum somewhat clearer graydorgo-metlially,the hunreral portions slightly more brightenecl. Pleura blackish,pluinose. Halteres pale )'€llorv throughout. Iregs with the coxae tlark brown,
pruinose; troehanters obscure yellow I femora brownish yellow; tibiae ancl
tarsi clearer yellow, the outer tarsal segments darkenerl. 'Wings (Fig. Z) lightyellow, with a eonspieuousbrorvn pattern, as follows: Arculus; origin of Be
and supernumerary crossvein in cell E; So 2i eord. antl outer encl of cell lst
M zi margiual spots at ends of all longitutlinal veins, smallest on I s1 veins
yellorv, darker in the cloucleclareas. Venation: rSo1 ending about opposite three.
fourths the length of the long J?sr' a supernumerary crossvein in cell B, conneeting vein M rvith the basal portion of Rsr. E z a short clistancebefore forkof I , * .; eord oblique, r-nl mueh arcuated; cell 7st M 2 unusually elongate,considerablyexeeedingin leng'th vein Mr*2
beyond it; basal section of Mttriore than twice nt, areuated to feebly angulatetl at near mictlength; qn-su
about its orvn length beyond fork of M or a litile less.
Abrlourcn tlark brorvn, the surface sparsely pruinose; hypopygrum obscureyellorv to yellorvish brown.

Ilabitat.-Cuba (Oliente).
Holot;'pe. 6 . Pico Turquino, Sierra Maestra,altitude 3750 feet,at light, Jnne 12, 1936 (Acuiia). Allotopotype, g h. Paratopotypes,
L2 6 ?.
The present fly is very different from the two other known species
of rSftannononryiahaving a supernumerarycrossveinin cell R (mesophragnra Alexander, mesopltragmoid,es
sp. n.) . The elongate cell
7st II2, rvith the elementsclosingits outer end not lying in transversealignment,is distinetiveof the species.
Tlre I'itriousspeciesof N/ron,tLor?,orny,ia
now known from Cuba may
be separatedb.y the follon'iug key :1. A supcl'nuluerurycrossvciuin trell R, connecting the long -Eswith vein M."
No supernunlcrary crossveirrirr. cell .8, -Es relatively short.n
2. Cell Lst M e elongate, longer than any of the veins beyond it; basal section
of. trI, not in transverscalignment rviilr nr. (Fig. Z.)
S, phragmoythorasp. n.
CeIl l st M z shorter than veirr M ' *,beyoncl it; veins closing eetl 7st
M, in approximate transvcrse alignment.r.
.
3. General coloration of mosonotum brownish yellow; wings yellowish, with aheavy brorvn pattern, inclucling marginal areas at ends of veins n s antl
B +i a dark cloud at arculus.
S. mesogthragmaAlexancler
General eoloration of mesonoturn clark brown; wings with a weak browntinge, restrietedly patterned with brown; marginal clark - clouds sparse,
small or lacking at ends of yeins -Es ancl B ti no dark cloucl at arculug.
(Fig.3).
S. tnesopthrag,moiitressp. n.

NEW

4.

'WingsOR LITTIJE

KNOWN

TIPUIJIDAE

III

627

unpattelned, except for the stigma and a darkened.area at origin ofBs.
S. s@,ratnuazai, .Llexancler
Wings rvith a clark pattern, including a series of marginal eloutls at endsof all longitudinal veins excepting R o
5. Cell B s very shorti R , far before fork of E, * ., the latter element exceeding vein -Rr alone; rnesonotumand pleura light reclclishbrown. (Fig.5).
S, breaicala Alexancler.
Cell n s long, R 2 lying at or elose to fork of B, *, * ., shortening or
obliterating 1?, * n ; mesonotumand dorsal pleura clark brown. (Fig. 4.)(bruneriarr,oand race).
6. Antennae (rnale) very small, scarcely longer than the combineclhead andpronotttm.
S. bruneriana bruneriono sp. n.
Antennae (tuale) Iarger antl more powerfully eonstructed,if bent backwartl
extending to shortly before wing-root.
S, bruneriana forti,cornis subsp.n.

Shannonomyiamesophragmoidessp.n.
Alliecl to mesopltragma,' general coloration of thorax and abtlominal tergites
dark brown I rvings with a weak brown tinge, restrictetlly patterned with brown;a supernumerary crossvein in eell 8r' eell lst M z of moderate length, the elemente elosing its outer entl in transverse alignment, the eell shorter than vein
lf,r*2 beyonilit.
Male.-Length about 5-6.5 mm.; wiog 5.5-7 mm.
Female.-Length about 6.5 mm.; wing 6.5 mm.
Bostrum ancl palpi blaek. Antennae short; seapebrown; peclieeland flagel.
lun blaek; flagellar segments short-oval, the verticils exceeding the segments.
Eead dark gray.
Thorax almost uniformly dark brown, the praescutum with a single metlian
darker vitta I humeral region of praescutum not brightenecl. Ealteres withd€m pale, knob weakly darkened. Legs with the eoxae yellowish brown totestaeeousl trochanters yellowl femora brownish yellow; tibiae and tarsi darkbrown. Wings (FiS. 3) rvith a weak brown tinge, the costal and prearcularportions a trifle norc yellowish; a restrictetl brown pattem, including tho
origin of -Bs and supernumerarycrossveinin eell Rr' Sazi stigmal cord antl
outer end of cell lst M zi very small to seareelyevident darker marginal cloutls
at ends of longitudinal veins, from -Er to ?nd A, Iacking or virtually so on
B ti veins brorvn. Venation: .Bs long, exeeeclingtwiee B , *, * .; a supemumeraly erossvein in eell n, eonnecting base of -Bs with about the mitlclle of
X; B, at or just beyonclfork of .Er*r*.;ecll 7st M, of mod.eratelength,
chorter thau vein M , * , beyonclit; elements elosing outer entl of eell 7st M z
in approximate transverse alignment, m shorter than the basal section of M s;
Hu about tlo-thirds its length beyontl the fork of M.
Abdominal tergites and subterminal sternites dark brown; basal sternites
oDecure yellow; hypopygrum yellowish brown.

628

OF P. R.
TIIE JOTTRNAIJOF AGRICULTURE OF THE UNIVERSITY

Ilabitat.-Cuba (Ot'iente).
Holotype, 6 . Pico Turcluino,Sierra llaestra, altitucle3750 feet'"'w'ithtype'at light, June, 1936 (Acuira). Ailotopotype, ? ' pinnecl
5 3 6.
Paratopotypes,rnes(bThe relationship of the present fl;' u'ith Sh,annon(rnxyia,phrugnw Alexancler is best shown by the hey $iven with the aG-'
eount of the Precedingspecies.

ShannonomYiabruneriana sP.n.
General coloration 4ark grayish brorvn, the pretergites conspicuous, eleaf
graJrtsilvery-white; halteres pale, the knobs weakly darkenecl; wings yellowish
E , Lt orwith a relatively heavy brown pattern; Rs of mod.erate length ;, beyond !t.utt Itt M, shorter than vein Mrf
shortly before fork of Es*ri
the elements elosing its outer end in tratrsverse alignment"
Male.-Length about 4.8-5 mm.; wing 5-5'3 mm'
Female.-Length about 5.8-6 mur.; rviug 5.8-6 tttm.
ligr4Rostruur rlark brorvn; palpi black. Antennae with scape ancl peclicel
longu
yellorv; flagellum blaek I antelnae relatively small, in the male scarcely
thau the combinetl head aud pronotum. Heacl ashy gray.
clear
Pronotum ash"v gla)'. Mesonotum tlark grayish browu; pretergites
paler. Ilalterersilverv-rvhite, conspicuous. Pleura dark brolvn, the sternopleurite
than thepale, the knobs iveakly darkened. Legs rvith the fore eoxae darker
darker''trochanters yellolv; legs yellorv, the ternrinal talsal segments
otlerslyellow;\yings (l.ig. a) I'ellorvish gray, the prearcular and costal t"egions eleat'er
origin of,a relatively heavy brorvn pattern, tlistributetl :rs follorvs: Al'culusI
series of/is; Sc,ri stiglnal eortl anrl outer elrl of eell lst II't: a marginal'd o and' grd
spots on all longitu4inal veins rvith the exception of -Rn , largest on
opveins pale, tlarker i' t1e cloutlerl ureas. Venation: Sc , ending abouti;or shortly
positc tlree-fourths the iength of th'l relativc'ly slrort /is; ll " atti p o f vein - 8, r atlter str ongly upcur vedi r - m r noc l er atel yl r e f o r e f o r k o f xr*.i
it, the eleto strongly arcuatetl l eell lst 11 , shorter than vein '7[I, * , beyoncl
beyondrrrents elosing itr outel entl in tlansverse alignment I m-cu' some distanee
fork of ilt.
.\btlonren tlark brorvn; hvpopygiunr pa.lel.brorvn.

Ilabitat.-Cuba (Oriente).
Ilolot.r-pe. d , Pico Turrluino, Sierra }Iaestra, altitucle 3750 feet,at light, Jlue 12. 1936 ( Acuira) . Allotopotl'pe, I , pinned witht)'pe. Paratolrotl'pes.10 d I .

Shannonomyia bruneriana forticornis sub sp. n.
C)Saracter.sas in the typical form, cliffering in the nature of the antenna'e""s4ric6 in the mzrle sex are larger and rnore powerfully constructecl than in the
typical for.m, if bent bAeku'arrl extending to shortly bef<l'e the rving-root..

NEW OR LITTLE KNOWNTIPULIDIE

III

529

Ilabitat.-Cuba (Ofiente).
Holotype, A , Pieo Turquino, Sierra Maestra,altitude 6000 feet,
Jnne 17, 1936 (Acufla). Allotopotype, I . Paratopotype,8 6 I ,rvith the t;'pe; 1 I , altitud.e3750 feet, June 12, 1936 (Acufla).
I take great pleasure in naming this interesting new Shan'in honor of I\[r. StephenC. Bruner, to whom I am indebted?tonontAia
for much friendly co-operationin studying the Tiputid.ae of Cuba.
Tlre relation of brtmeriana and its high-altitude form, f ort'i,contis,isinclieatedin the key provided rvith the deseriptionof Shq,nnonornAia
sp.n.
phragmophora

Hexatoma (Eriocera) juliana sp.n.
thorax and abtlouten oranger theSize large (rving, female, 77.5 mm.);sixtl abdominal segment blackened; head. dull blaek; legs ancl halteres black,the bases of the fore femora restrictedly yellorv I rving's rvhitish subhyaline,
leavilv patterned rvith clark brorvn, inc:luding a broacl nrargin that almost en'eircles the rving, together rvith broacl, entit'e, transverse seams at level of originof -Rs, cortl and outer end of cell I.st -U ,; basal section of vein -E u longer thanr-m; veins issuing flom eell 7st M 2 relativelv sltort, vein I[, being about a'3
long as th,e cell.
Feurale.-Length

about 19 mm.;wing 17.5 mm.

palpi black. Antennae black
Itostrurl black, sparsely gray pruinose;
tlrouglout, g-segmented; flagellar segments two and three subequal in length,
euec,eetling segurents gradually clecreasing in length I terminal segment about afift5 longer. than the penultimate. Head dull black, very sparsely pruinosel asmall tl6erele immediately behind each antennal fossa I median vertical tubercle
snrall anrl very poorly developed.
Tlorax entirely fiery orange, unmarked except for a more brownish suffusion and rveak pruinosity on the outer half of the mesonotal scutellum.
Leg'3 with the coxae and trochanters orange; reHalteres black throug[out.
ruaintler of legs black, the fore femora lvith nearly the basal fourth yellorv.
rvith the ground-color rvhitish subhyaline, h,eavily patternecl\\'ings (Fig.6)
rvitl rlar.k 5r.oyn, so as to leave extensive fenestrate areas of the grouncl before
origin of .R.s.,before and beyond th,e cord; additional areas of the ground inouter entl of cell Cw and involving most of cell 1st A; posterior border ofr.ell 7nd .1 narrorvly grayish; the brorvn pattern includes the prearcular ande<r.;t:rlbortlels, the latter continued almost arouncl the wing-margin to the end'of vcin Znd A, thence as a seam the length of the latter veinl broad dark
seiyrr.rlrevonrl arc:ulus, at level of origin of -Bs, along cord and on outer end of
e3ll 7st l/ ,,; ull veins rJark-eolored, seamed with brown, very narrolvly andinsensibll' -so in the interspaces. All longitudinal veins beyond eortl rvith abuntlant, eoprlllete ser.ies <lf triehia. Venation: Basal 'section of vein 1J, longer
tlran r.-rll,. R. just beyond fork of R, *, * n, the latter a little longer thanIt , lrut slorter t[an .RnI veins issuing from eell 1st M, relatively short, veinl,[ , allout 1s long as the eell, Il r rrruch shorter ; m-cu a little Irlore than onetlrirtl its length bevontl the fork of M.

630

TEE JOURNAITOF AGRICUITTUREoF THE UNWERSITY OF P. R.

Abdomen fiery orange, the sixth segrnent blackenecl excepting the restrictecl
nrargins: genitul segmeut orangel eerci elongate, horn-yellorv, except for thedarkened basal portion.

Habitat.-Cuba (Oriente).
Holotype, I , Pieo Turquino, Sierra Maestra,altitude 5100-5200
feet, June 12, 1936 (Acufla).
Allotopotype, 3 ,, Pico Turquino, north side, altitude 4500-6000
1 6,1 9,feet. June 18-20, l-936 (Darlington). Paratopotypes,
with the allotype; Museum of CornparativeZoiilogy, Cambridge.
Ttris striking speciesof erane-fly is named in honor of the collector, Director Juli6,u Acuira, of the Estaei6n'ExperimentalAgron6mica,Santiagode las Vegas,Cuba, to whom I expressmy sincere
thanks for nuulerousspecimensof Cuban Tipulidae. The only regional speciesthat at all resemblesthe present fly is Henatsrna(Eriocera) cubensisAlexancler,which is much smaller and has thewing-patternand.venationquite distinct, the veins beyond cell 7st Mz
being mueh longer than the cell.
Atarba ( Atarba) angust'ipennisAlexand.er.
Pico Turquino, Sierra Maestra,altitude 3750 feet, June 12, 1936
feet,
(Acufia). Loma del Gato, Sierra del Cobre,altitude 260G-3325
September25-30, 1935 (Acufla, Bruner & Scaramvzza).
( T eu,cholabis')bruneri Alexander.
T euch,ol,abis
Pico Turquino, Sierra l\'Iaestra,altitude 3750 feet, at light, Junet2, 1936 (Acufla).
( T eu,cholabis
T eu,ehola.bis) myersi Alexander.
Pieo Turquino, Sierra Maestra,altitude 3750 feet, June 12, 1936
(Acnfla). Loma del Gato, Sieruadel Oobre,altitude 260f3325 feet,
September25-30, 1935 (Acufla, Bruner & Scaramuzza).
oteroi Alexand.er.
Teucholabis(Teu,'chol.abis)
One male, Lorna cle Cunagua,June 24, 1936 (8. T. Barreto &Ir. C. Scaramuzza). The median dark annulus is clearly definedon
the fore and rniddle femora but is lacking or nearly so on the posterior legs.
Gnophomyia (Gnophomyia) darlingtoni sp. n.
Belongs to the lu,ctwosagroup; allied to flagran"s; size relatively small(wing, male, 6 mm.); heacl black, sparsely pruinosel rostrum, palpi, antennae,
halteres and legs blackl thorax fiery orange, unmarked.l wings with a strong

NEW

OIi I,ITTI,E

KNOWN

TIPUIJIDAE III

631

Br*"
blackish tinge, relatively narrow; B, just before fork of Er*r*.,thus being preserved a'r a short elementl abdomen blackl male hypopygium
rvith the basistyle narrowed. antl produced at apex.
Male.-Length about 5.5 mm.; rving 6 mm.
Female.-Length about 8 mm.; lving 7 rnut.
Rostrum ancl palpi black. Antennae black throughoutl basal flagellar seg'
nrents short-oval, the outer ones & little more elongate; verticils dense, longer
than the segments. Head black, sparsely gray pruinose.
Prothorax and rnesothorax entirely fiery orirnge, unmarked. Halteres black.
Legs rvith the coxae orange-yellolvI trochanters nrore brownish. yellow I remainderof legs black. Wings (Fig. 7) relatively narrorv, strongly tinged with blackish;posterior cells of rviug from M to caudal margin paler in color I veins andtrichia black. Abundant eoarse maerotrichia in outer cells of wing (intlicatedin figure by stippling). Venation: ^Sc, ending just beyond the fork of Bs,
Br*t
S c ' a s h o r t d i s t a n c ef r o m i t s t i p i R " j u s t b e y o n df o r k o f R r * ' * ^ ,tlrus being present but very short to punctiform 1 rm-cwshortly beyond the forkof .lI.
lbclornen in both sexes black throughout. I\{ale hypopygium (}.ig. 10)with the basistyle, b, narrorved at tip, the apex produced into a flattened yellorv obtuse lobe, its inner or rnesal erlge rvith a linear series of about six spinous
setaeI remainder of apieal lobe glabrous but the darkenetl portion of stylewith abundant setue. Outer dististyle, ocl, gently turved, the apex narrow. In'
ner dististyle, itl, nrore or less c,lavater provided with setae and short spines.
Plrallosor\e, p, broadly d'epressecl.

Habitat.-Cuba (Oriente).
Ifolotp-pe, A , Pico Turquino, Sierra l\{aestra,altitude 3750 feet,at light, June 12, 1936 (Acufla). Allotopotype, I .
:\t the suggestionof Direetor Aeufla, I take pleasurein naming
this interestingspeciesin honor of Dr. P. J. Darlington, Jr. The
nearestally is Gnophamyia(Gnophomyia)fl,agransAlexand.er(Costa
Rica) , rvhiehhas much the sameeoloration. The latter fly is larger
and bulliier, rvith broader wings and \Mith the venational details
distinct, having Rz beforethe fork of Ra*+.
Gonomyia (Progonomyia) bifasciolata sp. n.
Antennae weakly bicolorous, the flagellar segments brownish black, theirproximal ends yellorvI nresonotal praescutum with three polished black areas;legs I'ellorv, the tips of the tibiae and the outer tarsal segments clarkened;wings pale 1-ellorv,rvith trvo pale brown crossbandsI Sc , ending shortly before
fork of }is,' nrale hvpopvgium with the basistyle protluced at tip into a po\ryerful, gentlv eurved spine.
llale.-Length about 5 mm.; wing 5 mm.
Female.-Length about 8 mm.; wing 6.2 mm.
Rostrum brorvnish black; palpi black. Antennae with scape and pedicel
light I'ellorv; flagellar segments bicolorous, brownish black to black, with theproxinral enrls of the segments narrorvly yellolv, on the outer segments becoming

632

TrIE JOURNAL Ox' AGRICULTUREOF THE UNMRSITYOx' P. R.

unifornrlv darkenerl; flagellar segments passing tlrrough oval, long-oval to cylindrieall segrnents (nrale) rvith a dense rvhite pubescence and relatively shortverticils; in the fenra.le, the verticils are relatively longer, exc'eeding the
pubescence. Heatl light yellorv to brownish yellow.
Pronotum and lateral pretergites clear light yellorv. Mesonotal praeseututn
rvith the ground.-color obscure yellorv, with polishecl black areas on the cephalicportion of the otherrvise brorvn median stripe, and rvrth broad lateral areasthat nearlv r.each the nrurgins of th,e sclerite: scutunl blackenecl, the meclian&rea a little puler lrntl nrore pruinose; scutellum brorrn, darker medially at base;rlediotergite bla.ekenecl,sparsely pruinose, pale laterallyl pleurotergite pale on
rlorsal half, rrlol'e blackenecl ventrally. Pleura yellon', rvith & Inore blackened,
pr.uinose area on unepist,ernunl, and a reddish brown pruinose area on ventral
sternopleurite. In the female, the tlorsal are& is nlore continuous behind, involving the dorsal pteropleurite. Halteres obscure brorvnish yellorv, the knobs
6ar.ker, rvith yellorv apices. Legs rvith the fore coxae darkened, the remaining
coxae r.ecldishy.ellorv; trochanters yellorv I fenrora and tibiae yellon', the tips oft[e ]atter narrorvly but eonspicuously clark brolvn; tarsi yellorv, the basitarsi
darker apieally; renraining tarsal segments brorvn. Wings (Fig. 8) pale yellorv, 5'ith trvo pale brorvn crossbuncls, the tnore basal lying before origin of-Es, ,extending from eell l? to posterior ntargin in outer cnd of cell ?nd A : seconrl banrl at corcl, completely traversing the rving, clarker in the stigmal field;
veins ehiefly pale, clarker in the cloudecl area'J. Venation: Sc relatively long,
Sc , ending shortly before fork of -Es, 8c, At near mid-clistance betrveen origin
the latter subequal
shorter than Br*",of Xs ancl tip of Sc,; Rr*r*n
e e l l ? n tl , l 1 , a b o u t one- thir d longer than its petiolei m - cu s hor tl y
to n r *n ibeyond fork of ,I[.
Abclonrinal tergites of male brorvn, variegate,l by paler are&s, the sternites
1ro1e unifolnrly vellou'; in the fentale, the abdornen more uniformly blackenerll
hvpopygiunr pak' vellorv. Male hypopygium rvith the basistyle procluced into a
porvcr.ful, gentle eurved sitine. Outer dististyle a flattenerl, nearly hyaline blacle,
the urargin neat' apex rveakly toothed.

tlabitat.-Cuba, Panama, I\fexieo.
llolotype. A . Potrerillos, Chiriqui, Panama, altitude 3500 feet,llay 72. 1934 (D. V. Brou'n). Allotype, I , A. Naranjo, Pleo
Turcluino. Sierra }Iaestra, Oriente, Cuba, altitude 1650 feet, June
1 2 , 1 9 3 6( A c u i r a ) .
Parat,vpes,nulnerolls 6 L Vergel. Chiapas, Ilexieo, IIa;' 18Jurre 25. 1935. altitucle 800-1200meters (Damf) Nos. 4192,4193,
4203, 4207, 4239, 4259, 4260, 4267, 4268, 4320, 432r, 4350, 4353.
.1398,4416,4430.4503 and 4641; 6 I , Eseuintla, Chiapas,altitucle
80 meters,November11, 1930 (Damf), No. 1814; EI Zapote,Chiapas,
altitude 4-D0meters,November15, 1930 (Damf), I{o. 1867; Santa
Isabel, Chiapas,altitude 800 meters, November 16, 1930 (Damf ),Nos. 1879, 1880. 1881; Esmeralda, Chiapas. altitude 500 meters,
November18, 1930 (Damf), No. L924.

NEW OR LITTI,E

KNOWN

533

TIPUL,IDAE III

Gonrtnryiu(Progon,ontyia.)bifasciolatais entirely distinct fromthe other clescribeclspeciesof the subgenus,differing especiallyin
the llit.oloronsantennae,bifasciate u'ings, and the structure of theespeeiallyof the basistyles.
male ltvpo1l.1'giurn,
Erioptera (Mesocyphona) subdulcis sp. n.
l;. iolg to the tlulcis group; genelal eoloration clark brown; legs yellorv,
the fr.rnolil rvith a nArrow, seareely evident, subterminel, brorvn ring; wings
dark lrr.tr\\'ll, hanclsonrely patternecl rvith rvhitish areas, including a series ofspots at ends of longituclinal veins, and a nearly transverse and parallel'
mrgintlsieled rvhite crossbantl at eortl.
Irt'rrritle.-Length about li.5 nrm.; rving i-j.(inuu.
Hostrurn and palpi black. Antennae black throughout;dongate. Ilead chi'efly dtrrk brorvn.

flagellar vertieilsI)rotiotunr ltrorvnish black. IVfesonotal praescututtr tlttrk l)t'o$'tl, indistinctly;jriperl longitutlinally u'ith brorvni,sh 1'cllorv; posterior selerites of trtesonotuttr
Dorc unifornrly clarkened. Pleura blaek, n'ith I tlirty rvhite longitudiual stripe,
extentling fi.onr behind the fore coxue to the btrse of ttbtlottten,passing at'ountl
the bntse of h:rlteres. Halteles p:rle, the knol,s tlark brorvn. Legs n'itlt theeoxac lrron'nish bhl.h; tlochirnters obscure yellotv I feurora yellorv, rvith a paler
brorvn to y,ellorvish l:rorvn subternrinal ring that is very poorly evident; remainrlcr of legs yellon'. \\:ings dark brown, handsomely patterned with rvhitish
areas tha.t are confinecl to the vicinity of the veins I costal alea sliglrtly more
-Bsr' Sc, antl
5rellorvish; rvhite area,J arranged as follows: Arculus; origin offork of U ., * oi a nearly transverse band at cord, extencling from costa to
Cit, ahnost parallel-sided; large, conspicuous white areas at ends of allrcillongitutlinal veins exc'epting -Er, largest on -Rr *,, and Rrl veins pale blorvn,
pale yclloiv rvhelc traversing the rvhite &reas. Venation: Elements of corcl inalmost tlansverse alignnrent, as shorvn by th,e white crossband that lies along
the eorrl.
.,\lrrlonren dark brorvn, not or scr"cely variegate,l by briglrter.

If abitat.-Cuba ( Oriente) .
Ilolotp-pe, g, Pieo Turquino, Sierra I'Iaestra,altitude 5100-5200
feet, Jtttre 12, 1936 (Acufla).
is generally similar to certain
Erioptera (Illesocyqthona)su,bdu,Leisregional members of the dulcis group, differing especially in thessarcel.vevirlent femoral rings and in the nature of the wing-pattern,espeeilll.v the conspieuouswhite band at the cord.
r orhi,na ( T a.rorltina) uiolaceipennis Alexander.
7'cr
Pieo Turquino, Sierra Maestra,altitude 3750 feet, June 12, 1936
(Aeufla). Loma del Gato, Sierra del Cobre,altitude 2600-3325feet,
September 25-30, 1935 (Acufla, Bruner & Scaramvzza)

63+

TrrE JoUBNAL oF' AGRI0ULTURE
oF' THE uNtvFrRsITv oF p. R.

IJxpr,lNATroNoF Xtraunus( Symbols: Male hypopygium,-b, basistyle; 'ad,inner clististyle;od, outer dististyle; p, phallosomei t, tergite.)Fig. L.-Dolich,opeza (Megistornastir) deaera sp. n., venationFiS. 2.-Shan?tonorn?l'iaphragm,ophoro,sp. tr., venation.
FiS. 3.-Shnnnononttj'ifrnxesophragntoides
sp. n., venation.
Fig. 4.-ShonnonornAia bruneriana sp. r., venation.
Fig. 5.-Bhannonornyio brea'i,culoAlexander, venation.
Fig. 6.-He'fiaton?,a( Eriocero) ju,hianasp. tr., venation.
Fig. 7.-Gnoph,omgia ( GnopltornAia) dnrlingtoni sp. r., venafiorr.
Fig. 8.-Gonomy,i& ( Progonorny,in)bifasciolatasp. r., venationFiS. 9.-Dolich,opeza (Megistom,astir) cleaenasp.r., male hypopygium
Fig. L0.-Gnoplr,ormyia(GnophornAin)darlingtoni sp.r., male hypopygitrm.

PLATE XVI