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NEW SPECIES OF JAPANESE
Penr III(Aptera, Tipditlu)

CRANF-F.LIES

Bv CHARLESP. ALEXANDER
The species of Tipulidae described in this paper were collected by Dr. Kichizo Takeuchi in the vicinity of Gifu and on
Mounts Ibuki and Minomo, Central Honshiu. I am very greatly
indebtedto Dr. Takeuchi for this seriesof Japanesecrane-flies.
The types are preserved in the writer's collection.
Geranomyia multipuncta, new species.
Male.-Length, excluding roitrum, about 6.p mm.; wing,
8.1 mm.; rostrum, about 2.5 mm.

1?8

INSECUTORINSCITId

INSECUTORINSCITIE MENSTRUUS

MENSTRUUS

Fernale.-Length, excluding rostrum, about ?'5 mm'; wing,
9 mm.; rostrum about 2.6 mm'
Rostrum and palpi dark brown. Antennae dark brown
throughout. Head dark gray with a narrow' longitudinal, lightgray median line on vertex; vertex between eyes silvery white'
Mesonotal praescutum obscure yellow, the dorsum occupiedby three completely confluent brownish gray stripes, leaving thehumeral region and the lateral margins of the ground-color;scutum gray, the mesal margin of each lobe with a narrow
blackish longitudinal line; scutellum light gray with a capillary
black median line; postnotum brown, sparsely dusted with gray'
Ventral pleurites obscure reddish yellow, the dorsal pleurites
infuscated, pruinose. Halteres yellow, the knobs da'k brqwn
on the distal half . Legs with the coxae and trochanters yellow;
femora and tibiae obscure brownish yellow, the terminal tarsal
segments darkened. Wings with a faint brownish tinge, the baseand costal region yellow; a conspicuousbrown pattern, including
seven larger costal areas, the first in the basesof cells Sc and R,'
the second with a pale center in cell R; third at origin of Rs,
extending from C almost to M; stigmal spot large, connectedwith a spot in the basal third of cell Ro; terminal spots at ends
of veins Rrf, and Rn*ri smaller spots in cell C,' a series ofspots and blotches along vein Cu in cell M and caudad of thevein in cell Cu; cord and outer end of cell tst M" seamedwith
brown; small brown spots at ends of anal veins, that on znd, Alarger; brown dots near base of cell rsf A and, in cases, inbases of cells R and end A; veins brown, bright yellow in the
costal region except where they traverse dark areas. Venation:
Sc long, Sc, ending opposite midlength of Rs, Sc" at tip ofScr; Rs angulated and spurred at origin; cell rst M" elongate,
gently widened distally, longer than any of the veins beyond it;
basal deflection of Cu, beyond the fork of M'
Abdominal tergites brown, the sternites yellow. Mesal faceof pleurites of hypopygium produced into a long, very slender,
curved spine.
Holotype, male, Gifu, May 1, 1921 (K' Takeuchi)'Allotopotype, female.

179

curtivena' new species'
Male.-Length, 7 mm.; wing, 6.8 mm'
Rostrum obscure yellow, the palpi dark brown' Antennae
dark brown. Head dark brown, the anterior part of the vertex
Dicranomyia(Euglochina)

pale.
Thorax pale brown, the pleura indistinctly pruinose' Halteresvery long, pale, the knobs brown. Legs with the coxae andtrochanters concolorous with the pleura; f ermora and tibiae
black; tarsi white. Wings faintly infuscated, especially the apex
beyond the cord; stigma barely indicated, very pale; veins
Venation: Sc short, the distance between the tip of
brt*r,.
theSc, and the origin of Rs about one and one-half times
Rrf"
vein
the
distance from the origin of Rs to the tip of
justSc, close to the tip of Scr; Rs angulated and spurred
littlebeiond midlength, r at tip of Rr; deflection of Rnfu ashorter than Rs,' cell 'rst Mralittle longer than vein Cunbeyond
its
it; basal deflection of Cu' from one-third to one-fourth
anal
in
the
end
I
vein
of
originM;olfork
length beyond theangle immediately distad of the level of arculus'
Abdominal tergites dark brown; sternites obscure yellow'
Holotyye, male, Gifu, September3,7921 (K' Takeuchi)'Dicranirnyia (Euglochina) curtivena is very distinct fromthe type of the subgenus, D. (E') saltens (Dol') (syn''n. ,rrni|ormris Meij.) in the details of venation' The specimen
determined by Brunetti (Fauna British India, Diptera Nematocera, p. 3?3, 1912) as being this species is obviously notsaltens and is undescribed. For this species with infuscatedbasesto the metatarsi (Dawna Hills, India, altituile 2,000-3'000
fusci'basis'
feet, March 2-3, 1908), I propose the name D'(E')
new species.
Dicranomyia takeuchii, new species.
Male.-Length, ? mm.; wing, ?.5 mm.
Rostrum and palpi dark brown. Anteunal scape black; basal
half of flagellum obscure yellow, the terminal flagellar segments passing into dark brown; flagellar segments oval' Headshinv black.

180

TNSECUTOR INSCITI.4

MDNSTRUUS

Thorax entirely shiny coal-black, the praescutum with vague
metallic reflexions; a small silvery patch on the meospleura
ventrad of the wing-root. Halteres yellow, the knobs conspicuously dark brown. Wings subhyaline with a very heavy brownpattern, the baseand cells C and Sc yellow I conspicuousbrownareas at base of cells R and M, reaching costa; at tip of Sc
and origin of Rs, extending from C to M,'mid-distance between
the above two, extending from R to M; a very large stigmal
area, connected with a circular area at end of Rs and a conspicuous seam along the cord; outer end of cell zst M" seamedwith irown; wing-tip very broadly dark brown; cells M, Cw,
rst A and znd. A more grayish; veins brown, yellow in theflavous areas. Venation: Sc short, Sc, ending just beyond
origin of Rs, Sc, exactly opposite this origin which is angulated
and spurred; r at extreme tip of Rr; deflection of .Rn{o very
long, about equal to the first section of Rrf. ; cell rst M"
closed; basal deflection ol Cw, about two-thirds its length befor'e
the fork of M.
Abdomen shiny black, including the hypopygium, the segments narrowly but conspicuously ringed caudally with silvery white.
Holotype, male, Mount Ibuki, June 4, 1921 (K. Takeuchi).
This exquisite crane-fly is named in honor of the collector,
Dr. Kichizo Takeuchi.
Limnobia anthracina, new species.
Male.-Length, 8 mm.; wing, 8.6 mm.
Female.-Length, 8.8-9 mm.; wing, 8-9 mm.
Rostrum and palpi brownish black. Antennae with the scapeobscure yellow, the flagellum dark brownish black. Head gray.
Thorax shiny black. Halteres yellow, the knobs faintly darkened. Legs with the coxae yellow, the bases of the fore andmiddle coxae infuscated basally; trochanters yellow; femora
yellow, the tips broadly blackened; tibiae dark brown, the tips
blackened; tarsi black. Wings with a yellow tinge, the base andcostal region clearer yellow; stigma large, oval, brown; cord

INSECUTORINSCITI,4 MENSTRUUS

181

and outer end of cell rsi M" narrowly seamed with brown;
Cw broadly seamedwith brown; wing-tip indistinctly darkened;veins dark brown, those in the costal region clear yellow.
Venation: Sc long, Sc, ending about opposite two-fifths thelength of Rs, Sc, much longer than Sc, and at its tip; Rs long;r less than its own length from the tip of Rr; inner ends ofcells R, and rsl M" arcuated; basal deflection ol Cut close to
the fork of M.
Abdomen brownish black. Ovipositor with the valves longand slender, tergal valves very slender.
Holotype, male Mount Minomo, May 4,1921 (K. Takeuchi).
Allotopotype, female.
Paratofotypes, I male,5 females.
Limnobia amatrix, new species.
Male.-Length about ?.5 mm.; wing, 10 mm.
Fewale._Length about 8.5 mm.; wing, 9.5 mm.
Rostrum and palpi brownish black. Antennae with the basal
segment of scape dark brown; remainder of antenna light yellow, the terminal segments scarcely darkened. Head obscureyellow, the center of the vertex darkened.
Pronotum greenish yellow. Mesonotal praescutum greenish
yellow with four pale brown stripes, the lateral pair less distinct; scutal lobes conspicuouslymarked with brown, the medianarea greenish yellow; scutellum dark brown, the median portion obscure greenish yellow; median sclerite of postnotum darkbrown, the sides paler. Pleura brownish yellow with a narrow,dorsal, longitudinal, brown stripe extending from the cervicalsclerites to beneath the wing-root. Halteres yellow, the knobs
dark brown. Legs with the coxae and trochanters yellow;
remainder o{ the legs black, the tips of the femora narrowly
but very conspicuously light yellow. Wings tinged with paleyellow, handsomely variegated with brown; costal cell brown;
subcostal cell brown at base, over origin of Rs and at tip; abrown seam at base of cells R and M, continued as a conspicuous seam along vein Cu to the wing-margin, narrowly

L82

rNsEcuToR rNScrTrE MENSTRUUS

interrupted near three-fourths the length of vein Cu; a conspicuous seam along the whole length of Rs,' a brown cloud
extending from the tip of Sc across cells rsf R, and R, intocell Ru; narrow seamsalong cord and outer end of cell rst M";
stigma large, U-shaped; ro'ws oI circular brown spots on theveins beyond the cord; wing-tip in cells znd R' znd, M, andM, darkened; conspicuousseams on outer ends of anal veins;narrow but conspicuousmarginal seams in ends of all the posterior and anal cells; veins pale, dark brown in the infuscatedareas. Venation: Sc very long, ending far beyond the end ofRs, Sc, at tip of Sc, ; Rs extremely arcuated i r morc than twice
its length from the tip of R, which is arcuated at this point
as in Teucl'rolabis; cell rst Mr rcry long, widened distally; basal
deflection of Ctc, about two-fifths its length beyond the forkof. M.
Abdomen pale brownish yellow. In the female, the base of
the abdomen is tinted with green. Ovipositor with the tergal
valves bidentate at tips.
Holotntpe, male, Mount Ibuki, June 4, 1921 (K. Takeuchi).
Allotopotype, female.
Li,rnnobia o,matrir is a beautiful fly that is very isolated in
its relationships.
Ormosia confluenta, new species.
Fernale.-Length about 3-3.3 mm.; wing, 3.6-4.3 mm.
Rostrum obscure yellow, the palpi brown. Antennae withthe scapal segments brownish yellow, the flagellar segments
pale brownish testaceous to brown. Head obscure brownish
yellow, the center of the vertex sometimes infuscated.
Pronotal scutum light brown, the dorso-lateral angles sometimes darkened; scutellum yellow. Mesonotum shiny fulvous
brown, unmarked. Pleura yellow. Halteres yellow, the knobs
faintly darkened. Legs with the coxae pale, sparsely pruinose;
trochanters and femora obscureyellow; tibiae yellowish brown;
tarsi dark brown: Wings yellowish fulvous, veins a little darker.
Venation: Sc, ending beyond the fork of Rs, Sc, atrophied ;TNS4CUTORINSCITI4

MENSTRUUS

183

r on R, about its own length beyond the fork; cell rst M,
open by the atrophy oI m; basal deflection of Cu, a short distance beyond the fork of. M; anal veins divergent.
Abdominal tergites brown, the lateral margins of the segments paler; sternites brownish yellow. Ovipositor with very
long valves.
Holotype, female, Mount Minomo, May 4, 1921 (K. Takeuchi).
Paratopotype, female.
Orrnosia confl.uenta is allied to the Alaskan O. flazteola(Coquillett).
Dactylolabis diluta, new species.
Male.-Length about 6 mm.; wing, 9 mm.
Rostrum and palpi dark brown..Antennae dark brown. Headdark gray.
Pronotum large, brownish gray, the lateral angles of thescutellum obscure yellow. Mesonotum shiny black with a rather
sparse microscopic yellowish pubescence that appears like apollen. Pleura and sternum light gray. Halteres yellow. Legs
with the coxae black, microscopically pubescent; trochanters
yellow; remainder of the legs dark brown, the femoral bases
somewhat paler. Wings with a strong brownish yellow tinge,
the base clear yellow; very diffuse brown clouds, distributed asfollows :. Origin of Rs, tip of Sc. continued caudad along theless discord; outer end of cell rsl M"; at r; f.ork of Mrf";
tinct clouds at the ends of veins R, Cu,r, Cu, and in cell znd' A;
veins dark brown, more yellowish at the wing-base. Venation:
Sc, ending just before the fork of Rs, Sc, about twice its
length from the tip of Sc, ; Rs long i Rr*u one-half longer
than the deflection of Rnf u; petiole of cell M, aboat one-halflonger than the cell; basal deflection of. Cu, before the fork ofM, the distance about equal to the basal deflection of Rn*u.
Abdominal tergites dark brown, the basal sternites obscurebrownish yellow.
Holotype, male, Gifu, April 20, 1921 (K. Takeuchi).

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INSECUTORINSCITI,4 MENSTRUUS

Dactylolabi5 flavidipennis, new species.
Feynale.-Length, 6.3 mm.; wing, l.? mm.
Rostrum and palpi dark brown. Antennae with the firstscapal segment gray; remainder of antenna black. Head clear
gray pruinose.
Pronotum light gray. Mesonotal praescutum gray with four
conspicuousdark brown stripes; remainder of mesonotum darkgray, the scutal lobes a little infuscated. Pleura light gray.
Halteres yellow. Legs with the coxae yellow, the outer faces
pruinose; trochanters yellow; femora and tibiae light brown,
the tarsi dark brown. Wings light yellow; very indistinct brown
clouds at r, along the cord and outer end of cell rsf Mr;veins
yellow, darker beyond the cord. Venation: Sc, ending opposite
the fork of Rs, ,Sc, not far from the tip of Scr; Rrf , about
twice the length of r-m;Rs gently arcuated; petiole of cell M.
short, about three-fifths the cell; basal deflection ol Cu, at thelork of M.
Abdominal tergites light brown, sternites obscureyellow, witha capillary sublateral black line. Ovipositor very peculiar in
structure, the valves very high and compressed,notched at tipsHolotype, female, Gifu, May 5, 7921 (K. Takeuchi).
Dactylolabis longicauda, new species.
Male.-Length about 6-6.5 mm.; wing, 6.5-?.5 mm.
Rostrum and palpi black. Antennae with the basal segment
dusted with gray; remainder of the antenna black. Head gray.
Pronotum light gray with a narrow brown median line.
Mesonotal praescutum light brown with four dark brown
stripes, the extreme cephalic margin of the sclerite narrowly
grayish; scutum grayish brown, the lobes conspicuously darkbrown; scutellum and postnotum brownish gray. Pleura gray.
Halteres yellow. Legs with the coxae dusted with gray;trochanters obscure yellow; remainder of the legs dark brownish black, the femoral bases narrowly paler. Wings with auniform browr' suffusion, the base indistinctly yellowish;
stigma barely indicated; veins dark brown. Venation: Sc,
ending before the fork of Rs, ,Sc, at its tip; r near the extreme

INSECUTORINSCITIE MENSTRUUS

185

tip of ,Rr; petiole of cell 11, elongate,varying from a little lessthan the cell to one-half longer than the cell; in the paratype
cell M, is narrowed at the wing-margin ; basal deflection ofCu, at or before the fork of M.
Abdominal tergites dark brown, the sternites and hypopygium paler brown. Male hypopygium with the pleurites very
long, the pleural appendage appearing as a continuation of thepleurite, fleshy, on the mesal face at base bearing a conspicuous,curved, semi-chitinized spine.
Holotype, male, Mount Minomo, May 4, 1921 (K. Takeuchi ).
Paratopotylte, male.
Dactylolabis longicaud,a differs from the related D. flavidipennis chiefly in the coloration of the wings. The two species
form a very peculiar group of the genus.
Pilaria melanota, new species.
Male.-Length, 5 mm.; wing, 5.8 mm.
Rostrum brown, the palpi black. Antennae elongate, if bent
backward extending to just before midlength of the abdomenI
dark brown throughout. Head shiny black.
Mesonotum shiny black, the median area of the scutum andthe postnotum obscure yellow. Pleura obscure yellow. Halterespale, the knobs dark brown. Legs with the coxae and trochanters yellow; remainder of the legs dark brown, the femoral
bases narrowly obscure yellow. Wings tinged with brown;
stigma not evident; veins darker brown. Venation: Sc, almost
lacking, ending just beyond two-thirds the length of .Rq Sc,
at its extreme tip; Rs long, angulated at origin; r practicallyatrophied, on R, shortly beyond its origin; cell M, present,
a little longer than its petiole; basal deflection ol Cu, jast
beyond midlength of cell rst Mr.
Abdominal tergites dark brown, sternites yellow.
Holotype, male, Gifu, September 75,7921 (K. Takeuchi).
Hexatoma japonica, new species.
Mole.-Length about 5 mm.; wing, 6.2 mm.
Female.-Length, 4.8 mm.; wing, 6 mm.

186

INSECUTOR INSCIII&

rNsEcuToR rNscrTu€ MENSTRUUS

MENSTRUUS

Rostrum and palpi brown. Antennae short, pale brown.
Head dark brown, sparsely dusted with gray, more conspicuous
adjoining the margins of the eyes; vertical tubercle low.
Mesonotum black, very sparsely pollinose, the scutellum andpostnotum with an appressed microscopic pubescence. Pleura
dark brown with a sparse microscopic pubescence. Halteresbrown, the knobs darker. Legs with the coxae concolorous withthe pleura; remainder of the legs dark brown, the femora
slightly paler at base. Wings with a strong blackish suffusion;veins dark brown. Venation: Sc long, Sc, ending opposite thefork of Rs, Sc, a little more than its length from the tip of
Sc, ; r on R, at or just beyond the base, sometimes atrophied;
Rrf,
about two-thirds the length of. Rr; basal deflection ofCu, at the fork of M.
Abdomen brownish black.
Holotype, male, Gifu, September 5, 1921 (K. Takeuchi).
Allotopotype, female.
Paratottotype, male.

and along the cord; veins dark brown, paler in the costal
region. Venation: Sc, far before the origin of Rq Rs strongly
arcuated at origin i r-rn connecting directly with Rs about onehalf iti length before the distal encl of the sector; R, connecting with R, almost at the extreme tip of the latter; Rnlu about
twice r-m; cell rsf M" open by the atrophy of m; cell M, only
a little shorter than cell Mr;basal deflection ol Cu, about twothirds its length beybnd the fork of M, the fusion extensive.
Abdomen obscure yellow, the tergites with a continuous,
relatively narrow, dark brown median line.
Holotype, female, Gifu, May 10, 1921 (K. Takeuchi).

Tricyphona confluens, new species.
Female.-Length, 19 mm.; wing, 16.5 mm.
Rostrum and palpi dark brown. Antennae with the scapeand first flagellar segment obscure yellow, the remainder ofthe flagellum brown; fourteen flagellar segments, the basal
ones crowded. Head black, dusted with gray.
Mesonotal praescutum obscure yellow with a very broad andconspicuous median brown stripe and much less conspicuous
lateral stripes; scutum yellow, the lobes conspicuouslydarkened;
median area of scutum at suture and posteriorly with narrow'darker brown marks; scutellum and postnotum pale brown,
sparsely dusted. Pleura pale tawny, the sternites slightly infuscated. Halteres elongate, yellow. Legs with the coxae andtrochanters yellow; femora dark brown, the basesnarrowly but
conspicuously yellow ; tibiae pale brown, the tips darkened;tarsi dark brown. Wings with a strong yellowish tinge, cells
C and ,Sc more saturated; stigma large, yellow, faintly outtined with brown; narrow brown seams at Scr, origin of Rs

187

I

I

Nesopeza albitibia, new species.
Male.-Length about 9 mm,; wing, g.B mm.
Rostrum yellow, the palpi brown. Antennae relatively elongate, if bent backward extending beyond the base of the abdomen; scape light yellow; first flagellar segment obscure yellow;
remainder of flagellum dark brown. Head brown, the anterior
part of vertex yellow.
Thorax brown without distinct markings, the pleural region
crushed. Halteres yellow, the knobs conspicuouslydark brown.
Legs with the coxae brownish yellow; trochanters yellowish
testaceous,the tips narrowly dark brown; tibiae white, the tipsof the posterior tibiae narrowly black; anterior tibiae slightly
infuscated, the extreme base and a broad subterminal area
white, the tips narrowly black; tarsi white. Wings with a strong
brown tinge, the stigma dark brown; a conspicuouswhife spot
before and beyond the stigma; veins dark brown. Venation:
Generally as in N. taiutani,aAlex.; Sc longer; petiole of. cell M,
only about one-half the length of the petiole of cell Mr; fusion
of. M and Cz slight; cell znd, A narrow.
Abdomen dark brown, the tergites indistinctly ringed with
pale; sternites more distinctly bicolorous, the apical half ofthe segments distinctly obscure yellow, the basal half darkbrown. Hypopygium chestnut-brown.
Holotype, male, Mount lbuki, June 4, 1921 (K. Takeuchi).

188
TipulaINSECUAOR

annulicornuta,

INSCITI.&MENSTRUUS

new species.

Male.-Length, 15 mm.; wing, 18 mm.
Frontal prolongation of the head brownish yellow; palpi pale
brown. Antennae of moderate length, obscure yellow, basal
flagellar segments a little darker basally, the terminal segments
more uniformiy dusky. Head brownish yellow.
Mesonotal praescutum obscure yellow with three brownish
yellow stripes, the median stripe slightly darker anteriorly; remainder of mesonotum yellow. Pleura whitish yellow. Halterespale, the knobs brown with yellow apices. Legs with the coxae
concolorous with the pleura; trochanters yellow; femora andtibiae yellowish brown, the tarsal segments dark brown; legs
long, metatarsi only a little shorter than the tibiae. Wings
tinged with grayish yellow, the base and cells C and Sc moresaturated, yellow; stigma bicolorous, the base yellow, the distal
third brown; a very conspicuous obliterative area before thestigma, extending from Sc, into the base of cell M", almost
filling cell rst Mr; veins brown. Venation: R, preserved; cell
rst M" small, pentagonal.
Abdomen brownish yellow. Male hypopygium with the ninth
tergite small, the caudal margin with two long, widely separated
spines. Outer pleural appendage very small, cylindrical. Dorsomesal angle of ninth sternite with a dense brush of short yellow setae, immediately basad of which arises a powerful chitinized arm, bent laterad for a short distance and thence caudad
and dorsad. Eighth sternite with a median brush of setae, on
either side of which arises a chitinized spine that bears numerous, parallel, ridges for its whole length, the whole appearing
like the horns of certain antelopes, these spines decussate.
Holotype, male, Gifu, May 10, 1921 (K. Takeuchi).