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NE.w OR LITTLE KNOvIIN TIPULIDAE
FROM
NEv\I CALEDONIA AND SAMOA
by Carnr,rsP. ATEXANDER(Amherst, Massaohusetts).

The present report is based on a small collection of crane-frios
taken in New Caledoniain late May and June, {.g28, by profossor
T. D. A. and Mrs. Cockerell,togethorwith a singlo species from
Samoa receivedfrom Mr. E. II. Bryan, Jr. The typesof the novelties
from New calodonia are prosorved in my colleotion,through ilre kindinterest of Professor Cockerell. The type of tho Samoan Limonia
bryaniana has been returnod to Mr. Bryan and is proservedin the
Bishop Museum, Honolulu. I extend my sincerethanks to professor
and Mrs. Cockerell, and to Mr. Bryan, for their kind iutorost in
making known the interesting Tipulid fauna of tho pacific Isranals.
Macromastix

novocaledonica,

sp. n.

Goneral coloralion fulvqus yellow, the praescutum with four
more or less distinct brown stripes ; frontal prolongation of heod
long, exceedingtho remaindor of head ; postnotal mediotergite darkbrown, witha nedian yellow vitta; wings with a strong yellowish
tinge, l,hostigma dark brown, conspicuous; abdominaltergitesobscureyellow, the caudal margins of the segments infuscated, the aruountdecreasingon the outer segments.
Male. - Leng[h 6-9 mm. ; wing &11 mm.
Ftmale. - Length about 8 mm. ; wing tj. mm.
Frontal prolongabionof head long and slondor, oxceeding theremainder of the hoad, obscuro brownish yellow, darker brown
abovo ; nasus lacking ; palpi black. Antennao short, not oxtending
far beyond tho end of the frontal prolongation ; first scapal segment
pale at base, the remainder of l,he organ brownish black. Heacl deep
tulvous orango, most intense on tho vertox. In the fomale, the vertex is moro infuscated,with the orbits broadly yellow.
Mesonotalpraescutum orange-yellow to obscure yollow, in cases
Dipbra, t. Y, feec. r, p. 83; Parir, r5. XI. rgrg.

TIP U LID A E

84with four bcown stripes, the posterior ends better defined, in still
othoe cases tho stripes obliterated ; scutum conspicuously darkbrown ; scutellum testaceousyellow ; postnotal mediotergite dorkbrt)wn, divided medially by a yollow vitta that widens out behind,
not or scarcely reaching the onterior margin of the sclerite. Pleura
chiofly pale yellow, the pteropleurite paler yellow ; dorsal pleurotergito chiefly dark brown ; dorso-plourril region infuscated anteriorly,
tho posterior half palo. Ilalteres infuscated, the extreme base of thostom yollow. Legs with the coxae and trochanters yellow I femoro
obscureyellow, the tips very narrowly blackenetl ; tibiae brown, thetips passioginto brownish black ; tarsi black. Wings with a stroug

C A LE D ON IA

85stroight,
ringto bo atnearmidlength of cell lst X[21 vein 2nd' A
the cell of moderote width.
Abdominal tergites bicolorous, the segments obscuro yollow, theamountcaudel margins of the segments broadly infuscatod, tho
basal
the
with
tergiteseconilsegmenl,s;decreasingon the outerdark'hypopygium
ring entirely palo ; sternites yellowish ;IIab. Naw Caledonia.
Ilototype ;f, PlumFarm. May 30, lg28(T. D' A' Cockerelt)'
AtlotipotgPe Q, Juno7, l'928.
j.928(f 'D'A'and' W' P' Cocke'
Parutipolypes, 6 $ 9, Juno 3'?,goo"uil, lvest coast, May 23, L928 (CocherelQ,paiatgprt,2 dd,rell).
The presont species is closely alliecl to Macromastio tenuilrons
Alexander, of northern Now Zealand'
Macromastix

figr

tFlt 3Fig.{

FR OM N X W

C. P. ALEXANDiN

Fig.5

2, Wing of Mauo'
I e 5. - {, Wing of Macromastio nouocaledonica, sP' n. Fis.
"mastir cockerellac,sp.
n. - 3, Wing of Ltmonia (Limonia) brganiana, sp. n. - 4'
5, Wing of Gonomgia
noaocaledonica, sp. n. Wing of Limonia (Idioglochina)(Pti-Ioslena) cockerelli, sP. n.

yellowish tingc, iridescent, cell C moro suffusod : cell Sc brownish
yellow ; stigma conspicuous,dark brown ; Yague' scarcely evidentsea,mson anterior cord; extremo wing-tip narrowly darkened ; veinsderk brown ; costa in the stigmal region incrassated. v.-einselmost
without rnacrotrichiao except on C, there being a fow on tho basal
portion of Sc and others in the stigmal region on 8c2t r?r. Yenation(Fig. t) r ^Rs moderatoly long, gently ercuated at origin I frr+g+rolongate, exceeding rn-cu ; cell Mr subsessilo to shorl, petiolato ;Ms+r short, subequal to tho basal sectiosot M, zr-co thus spPos'

cockerellae,sP' n'

Goneral coloration brownish yellow, the praoscutum with fourdarker lrrown stripes ; postnotum whitish cinoreous,darkenedbehind ;frontal prolongatioir of head short ; wings whitish subhyoline, orwilh a iaint duskv tingo ; veins almost without macrotrichiae ;r?z48elongale:.m cuin alignment with the short basa! seotion ofMa, the latter approrimately ono-half Mt+a'
MaIe. - Length about 6. 5 mm. ; wing 9 mm'
Femule. - Length aborit 7. 5 mm' ; wing l'L'5 mm'
Frontal prolongatio4ot hesd relatively shortand stout, about onohalf the remainder of the head, obscure yellow, without n&sus;
palpibrown. Antennaewibh the scapalsegmentsbrownish [estaceousI
ftagellum broken. Ileait broad, Iighb brown'
Nlesonotalpraescutumbrownishyellow,withfourdarkerbrowndarksl,ripes,the iniermodiate pair confluont in front ; scutal lobes
eneil medially ; scutum infuscated ; postnotum whitish cinoreous,the caudalpo*tioo darkenod.Ploura chiotly whitish,,,the anepisternum, sternopleurite and meron slightly darkenod' Ilalteres chiefly
knobs
put", tfr" stel a [ttle darkened outwardly, tho apices of thedaribr"oru yellow. Legs with the coxae obscureyellow, somewhot
browker basaily ; trochanters obscuro yellow ; femora and tibiae
brown.
into
passingtarsi
infuscated;uish testaceous,the tips weakly
wingswhitishsurrhyalineorwithafaintduskytinge,collscandthe Jma[ stigma pale brown; veins dark brown. Voins without macroe fow
trichiao except on costa, the prearcular portion of R'M' and86c. p. ALExar\D:n

microscopic punctures on the basal half of gc.
venation (Fig. 2) :t?s relatively short, arcuated to subangulatedArarr.ry ioig, uUout;
one-half longer tban rn'cu; fiz in alignment withni+s ;ie' M1 deep,
its potiole nearly twice m nx-cuin arignment
with the short boser;
section of Ms, al, extreme outer end of cell lst M; rfZr*ruppro_i*utely twice the basal section of Ms ; yoin 2nd A
short ana straignt, cell
znd A relatively wido.
Abdomen with the bas-altergite yo'owish testaceous
l secondsegrnent obscureyellow,mediaily, dark brownlaterally ;turgites, including the hypopygium, dark brown"u,oJoiog; fusut ,to"oit", yutthe outer segmentsmoro bicolorous,lhe bases
obscureyellow,
fgw,the-tips broadly darkened ; subterminal segments
unifomry darkoued.
l/aD. New Caledonia.
Holotype 6i'.Bouroil, west coast,May 26, rgzg (7.
D. A. cockerettl.
A.llotopotype g, May ZZ, LgZg(W. p. CockeritQ.
very distinct speciesof Mau'omasrrois respectfu[ylli*dodicatedto Mrs. T. D. A. Cockerell.
r sinco the above description was preparedI havo receive, twomales, that are consideredas being pu""typ..,from professor cockerell. These wero taken at Bourait'oott{uy lgrh. A notablo featuroof tho male lies in the fact that the antennae
sre yery rong and f iform, exceeding the body in length. r

Limonia (Limonia) bryaniana, sp. tr.
Gonerolcolorationyellow; antennaomoniliform,
black,the basal
segmentochreous; mesonotarpraescutum'Jthbuffy yelrow,,h.uubrownishfulvousstripos knobs of halteres;
aurt n"o*o j iostu*io,tarsi sxtensivelywhito wings wil,h astrong brownish ti.ngo,the;veinsdarkerir-rn mirchshortenedto obtiteratld; ;"|;l;pyg,um
with the ventraldististyresmat, theapexheavily brackened,bidentate.
("1:, - LengthaboutT mm. ; wine g.T mm.
Rostium short,,brownishtrtack;.pafi concolorous.Antennaewith
the first scapalsegmentoc,hreous,the ietainuer of the organbrack;scaparsegme.nte ritile ronger than thefirst fragoilar; tagel::-""-"dlarsegmentsmon'iform, nearry
iloburar, with short'.";i*i necks,
graduallydecreasine-i4
rir,euna rJ.";G
moreelongateoutwardry;
terminalsegmont,iq
slondor,ourrt*.a into an apicalpoint;"iq
eachflagellars"gmgolwillr
a ooiiqt;llv aprangedsetaof unusual

TIP U LID A E

FN OM N E W

C A LE D ON IA

87

length. Anterior v6rtex and front narrow, obscureochreous, the pos'terior .veriex more brownish ochreous, darkor medially.
Pronotum yollow, v€ry narrowly darker. medially. Mesonotal
praescutum buffy yellow with threo broad brownish fulvons stripes,
the lateral stripes vory broad, extending to the laterol margins of
the sclerito I scul,umyellowish fulvous, the median area and scutellum more testaceousbrown ; postnotum palo testaceous brown'
Pleura obscure testaceousyellow, the dorso-pleural rogion a little
d.arkor. Ilaltores of modorate length, obscure yellow, the knobs
dark brown. Legs with the coxae and trochanters obscure yellow ifemora (posterior) brownish yellow to light brown; tibiae somowhat
darkor brown ; tarsi brownish yellow, the terminal segments moro

Fig.E

Fig.6

- 7, M Je
of limonia (Limonia) brganiana,-s-P'l'sordi.da (Brunetti). - 8, MAle h-y.popygium: basrstyle.; dd : oorsar(Di$anomgia) illingworthi.(Alexandet).
. Dv entral ds trs ty l e'th. tergi te; v d:
t:9g:go napophy s i s ;6 e 8. - 6, MAle hypopygiumFis.
-'l""oouii"ttt'oi-Iimoiki"'(biqanomgia)
oi:timinta
di sti styl e;

infuscuted I posterior l,arsi with the basal two sogmenLswhite, including the."iu", th"'terminal segmentsinfuscated; posteriorlegs muchstouter than tho others ; claws lo'ng and slender, bearing an acute
Iateralspine at near midlength,in addition to the stouter basal spine.
wings with a strong brownish tinge, somewhat more suffused in
the Jpical and anal t"tl* ; baial half of cell 8c darker ; wing aper incell ^ice and r?3narrowly infumed ; stigma long-oval, slightly darker browo I veins dark brown. costal fringo very long- antl conspicuous, longest on the basal halt otwingrthence decreasingin length
to the rving-tip. venaiion (Fig.3) : Sc long, sca endiugat midlengthof the long l?s, scs at its tip ; free tip of sc2 pale, wiihout macrotrichie, subequal to Rt ; Rt i e ontirely atrophied, its position indi-

88

TIP U LID A E

C. P. AL EXANDER

cated by a weak angulation of the combinpd voin r?-and .Pz; l?rlong, only gently arcuated; veins frsrand rft .1slong, extending gonerally parallel fro one anothor to the wing-margin ; coll Ist Mz vorv
broad at bose, obliterating or greatly reducing r-rn ; m strougly
arcuatod, tho basal soction ol Mz straight, transverse; m-cu closo to
tho fork of {f.
Abdominal torgitos dark brown, the caudal margins of tho seg_ments very narro'\Mlyand vaguely palo ; basal sternitos obscuro yellow, tho outer segmontsdarker. Male hypopygium (Fig. O) with the
9th tergite (9t) having the lateral lobes vory broad, their mosalcaudal margins with numerous powerful setaethat are slighily docusBate i what appeors to be a median lobe from benoath the tergito isa very large, oval, dopressed plate, dark-colored, the margin and e
median lino pale, the surfacedensely sot with microscopicsetm; therois a possibility that this is a modified anal tube but this seemslsgs
probable. Basistyles(b) very long and slender,the mesallobe relatively small, sel,iferous.Dorsal distisiyle (dd) a noarly straight, weak
rod, the apex darkened and slightly roughened. Ventral disl.islyle(vd) much smallor than tho basistyle but longer than the dorsal dististyle, the basal portion fleshy, with long setae,thoso on tho outerface poworful and curved, those on the inner face long and straight ;apex of stylo heavilv chitinized,bidentate, with a single conspicuous
sota from between tho two teeth. Gonapophyses(g) with the mesal
apical angle producedinto a long, nearly straight horn, the apex alittle curved and darkened.
.fy'aD.Samoa..
Holotype d, Yaeo, Upolu, altitude {100 feet, April 25, lg}4(8. II.
Eryan, Jr.).
I tako great pleasuroin naming this very distinct Linonia in honorof the.collector,Mr. Edwin H. Bryan, Jr.
Limonia(Thrypticomyia)

subsaltens(Alexander).

One d, PIum Farm, New Caledonia,June 7,lg28 (Cockeretl).
Edwards (Insectsof Samoa, Diptera Nematocera,pt. vI, Fasc. 2 :77; 1928) has discussed the structure of the male hypopygiurn.
Tho rostral prolongation of the ventral dististyle is very slenirer, alittlo expanded before the tip and provided with two long strong
setao. Tho rostral spinesaro placed on the stylebasadof therostralprolongation, nearly equal in size and curvature, being long andslonder, the outormost from a larger basal tuborclo than the innerspino.

Lirnonia(Dicranomyia)

FN O} IN E .w

C A LX D ON IA

illingworthi

89( Alexander)'New Caledonia:Noumea, May 16, L928 ; Bourai.l,west coast,May
27, Lg28 (Cockerell).
I cannot agree with myfriend Edwards (Ann. Mag. Nat. Hist-'
ser. 9, vcl. 20 : 237 : 1927.- Insects of Samoa, Diptera Nematocera'
Pt. VI, Fasc. 2 : 76 ; {928) that the present species is identical withL. (D.) so:.dida(Brunetti). The two speciesaro closely ollieclbut thedistinctions botween the two are apparently constant'hypopygial
sordida (Brunetti) is hers described from
(Dimanomyia)
Limoniamateriirl taken at cherrapunji, Assam, Octobor ,.8, {920, by SeniorWhite, determinedby Edwards. The structuro of the hypopygium
1fig. 7) is as follows :Basistyle (b) with the ventro-mesallobe long, narrowed outwardly,
the tip boing narrowly obtuse ; the finger liko lobe on the face ofthe basistyle is long and slender, the length more than four times thediameter at mitilength ; in addition to this tubercle, thero is a short,
low tubercle neaf the caudo-mesal angle of the basistyle. Rostralspines slighily longer. Dorsal dististyle narrowed gradually to thelong acute tip.
Limonia (Dicranom'yi'a)iltingworthi is now ropresontedin my collection by specimensfrom Fiji, New caledoniaand Northern Queensland. The structure of l,he hypopygium is as follows (Fig. 8) :Basistyle(b) with tho ventro-mesal lobo vory broad, not or soarcely
narrowed outvrardly, tho apex subtruncatetl ; finger-liko lobe on faceof basistyle short, tho length scarcelv twice the dianleter at midlength ; tho socondtubercle is.lacking, but replaced by a group of
powerful setaeat this point. Rostral spinesa little smaller and shori.r. Dorsal dististyle (dd) more suddenly namowed to the slendor
acutoapex, a little eXpandedjust before this point.
The condition obtaining between sordida and illingtnorli.i thus isentirely analogousto that oxisting between pseudomorro Aloxander'of Eastern Asia, and occidua Edwards, of Europe- Both couplets
include vicarious represontabivesof speciosthat aro distinguishable
chiefly by slight but apparently constant differences in the structureof the male hypopygium. The question of tho distinctness or idontity of such closely allied pairs of species hecomes of more thanordinary inrportanco,due to the vast range of the insocts ooncorned.
Limonia (Dicranomyia) rapa Alexander is still known only fromfemalo specimonsfrom the Austral Islandsl Thero is a possibility
thet these represont largo, doeply-colorodindividuaLsol illingworthi

90c. p. ALExANDER

but theof the malo sex wil be required to fina[y setfle.discgvery
tho rolationship. Basedon tho female sex alono,thertwo forms certainlv
oppear distinct, rapa having strongly tinted wiugs, with lhe potternunusually heavy for a member of rhe liberta group ; the stigmal
spot is oval, instead of being a relatively narro.w seam [o Rc or subcircular in outlino i m-cu is immediately bofore the fork of M andthe distal end of .r7lhas from eight to ten macrotrichiae.
Limonia(Idiogtochina)

novocaledoniea,
sp. n.
General coloration dark brown, the praescutum with four still
darker brown stripes I antonnae dark-brown ; flageilar stigments
strongly produced, almost sessils; wings yollowishgrey, the veins
beyond ihecord dark brown ; abdomen dark brown, the caudal margins of the segments narrowly paler, nore evidenily so on the sternites.
Male. - Length 5-S.Smm. ; wing Z-8 mm.
Female. - Longth about $ mm. ; wing 6 mm.
Antonnae dark brown, darker than tho rostrum flagollar segmetrts;of the male strongly produced, as in tusitala Alexandoi, uila Edwards
and othor species; individual flagoilar segmentsalmost without necks.
Head infuscated,a little brighter on tho orbits.
Mesonotalpraescutum dark castaneous,paler latera[y, with four
dark brown stripes, the intermediate pair not reaching the suture, incsses more or less confluont on the anterior half ; in somo specimens,
tho lateral praoscutal stripos are distincily darker in color'than theintormsdiato pair ; scutal lobes dark brown, the median area pale;scutollum and posl,notum palo, slighily pruinose. pleura reddish
brown, tho surfaco weakly pruinose. Halteres pale throughout. Legs
wil,h thecoxao obscure yellow, the fore coxae slighily infusoated;remainder of the legs yellow, the terminal tarsal segments darkenod.
Wings yollowish grey, the base and costal region poro yellowish ;prearcular and costal voins and the longitudinal veins of besal thirdof wing yollowish, the remsining veins passing into brown. costal
vestituro short and spinous; longitudinal veins beyond the cordwith regular series of small setes. Yenahion (fig. a) : sc relotively
short, sc ending nearly throe-fourths the longth of Rs beforo theorigin of tho latter ; sce ver-v far from the tip of scr, the latter alonslonger ihan r? : r?s straight, without macrotrichiae, nearly equal tollear and nearlytwice the basal section of Fr+ r ; coll r?l large; cell
7st Mz relatively narrow i m-cu before the fork of If.
Abdomon dark. brown, tho caudal margins of tho segmentspaler,

TTPuLTDAE FRoM NEw

oALEDoNIA.

9t

more conspicuouslyso on the sternites ; hypopygium pale. Male hypopygium with the ninth tergite large, tho caudal margin with a cons
picuous V-shaped median notch, the sidesmore or less sinuous I
laterallobesbroad, with stout spinoussetae.
The female differs in the usual sexual characters, especially t[evenation and slightly less developed flagellar segments.
I/a6. New Caledonia.
Holotype, d, Ngo Bay, atlight, May L4, lg28(T,D.A. Cockerell).
Allotopotype, S, pinned with the type.
Paratopotypes,4 $ Q.
Limonia (Idioglochina) nouocaledonicaseems.to bo most closely
allied to L. (1.) ailae Edwards, of New Hebrides, differing especiallyin the coloration of the body and wings.
Gonomyia(Ptilostena)

cockerelli,

sp. n.

General coloration brownish grey ; antennae black throughout;pleura dark brown, stripedj longitudinally with yellow ; wings witha faint bror,vnishyellow suffusion,the stigma slightly darkor ; r?, distinctly preserved.
Female. - Length abouti6.2mm. ; wing 6.2 mm.
Rostrum and palpi black. Antennae black throughout. Headbrownish grey, somewhat brighter in front.
Pronotumdark browuish grey I antorior lateral pretergitesrestrictodly obscure yellow. Mesonotal praescutum brownish grey, with astill darker median stripo I humeral rogion and lateral margins obscure buffy; scutal lobos brown, each lobe variegatecl with darker
brown ; scutollum light brown ; postnotum grey. Ploura dorsallydark brown, striped longitudinally with yellow ; dorso-plour.alregion
yellow ; a yellowish stripe bogins behind the fore coxa, broadoning
on the meron and metapleura ;ventral sternopleurite more reddish
brown. Ilalteres dusky, the knobs infuscated. Legs with thecoxae and trochanters yellow ; remainder of legs broken. Wings
with a faintbrownish yollow tingo, the base and costal region somewhat brighter ; stigma a littte darker than the ground-color, lying' entirely between veins l?, and E3; veins dark brown, the prearoularvoins, 8c, l? and Cu paler. Venation (Fig. 5) : .P, preserved, apptoximately as loug as r?r+s+l ; r?rnr about sne-half longer than
nx-cu ; rrl- cu, more than its own length before the fork of M.
Abdomendark brown, the basai sternites obscureyellow; caudal
margins of the sternitesnarrowly yellow. Ovipositor with the tergal
valves slender, reddish horn,color, strongly upcurved.

H.Hfir'
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ii

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92

C. P. AL EX A N D E N

lTaD.New Caledonia.
HolotEpe ?, Dge, Ilo Ouen,June6, 1928(T. D. A. Cockerell\.
G, (P.) cockerelliis namod in honor of Prolessor T. D. A. Cockorell, to whom I am very greatly indebted for specimons of Tipuli_
dao from all parts of the World. It is reaclily told from alt altod
speciesby l,hevenation, especiallythe retention of r?r.

Descriptiond'un Atherix

du Japonpar n. SEGUY(Paris).

Atherix"l l

Galloisin. sp.

t. Allong6, d'un noir brun ldgdremont luisant, couvert d,uno
pruinositd grise plus dense sur les pleures. Yeux s6par6s par un
espace6gal d deux fois Ia largeur ilu triangle ocellaire. Bande frontale d'un noir velout6 au milieu, grise en haut et en bas. Occiput
gr!s, d pilositdnoire en haut, grise et blancheen bas; Trompe 6poisso
et longue, uoire; palpes bruns. Antennes courtes : articles I et II
6gaux,noirs; Ill rdniforme aussi long que les deux premiers r6unis,roux ; chbtelong, noir, d puboscencemicroscopique.- Ddpression
noto-pleuraleb longuessoiesnoires;hypopleuro,m6sopleureethanches
i soies blanches.Pattos brunes : fdmurs jaunes; tibia II cl'un brunclair. Balanciersjaunesi renflement noir. Ailes jaunies : une bands
tra,nsversalo
brune,diffuse en arridre; cellule discalelongue; fourehe
MAt * l?++ uitrds longue. - Abdomen d'un noir veloul,6: vu d'&rribre en avant on observerrne ortrementationd'un gris bleu : deuxtacheslat6ralessur les tergitesI et II, tergito III & tachesdiffuses;ly
et V avec une large bande posl,6rieure; VI et VII avec une tachemddiane,sterniLeset cerquesbruns. Pilosit6 blanchdtreen avant. uoiro
en arribreetsur les cdtds.- Long.{.0,5 mm.
Japon:Tokyo, 27.VI.{908 (EdmeGallois).

Diptera,t. V, fasc. r, rb. XI. rgrg
(Paris, P, Lechevalier).