lec
Reprinted from. Bur,r,nrrwoF TrrE Bnooxr,yu Eurouot,oclcAr, Socrnry,Vol. XLI, No. 3, pp. 65-71. June, 1946

UNDESCRIBED SPECIES OF TIPULA FROM WESTERN NORTH AMERICA (DIPTERA, TIPULIDAE).
PART III.
Bv CnIRLES P. AT,BxANDER,Amherst, Mass.

The precedingpart under this generaltitle was publishedin 1946
(Bull. BrooklynEnt. Soc.,XLI:45). At this time I am describing
three further speciesof.Tipula, chiefly basedon the materials containedin the great MelanderCollection. As before,my deepthanks
are extendedto Dr. Melander for the opportunity and privilegeof naming these flies and for his kindness in permitting me toretain the type of one of the speciesthat is representedby a single
specimen.
Tipula (Oreomyza) yellowstonensis,n. sp.
Belongsto the borealisor xrncagroup ; size unusually small(wing, male,11 mrn.or less); rnesonotal
praescutumgray withthree darker gray stripesthat are borderedby brown ; femora
brorvnish yellow, the tips narrowly brownish black; claws(male) toothed; wings pale brown and yellowishsubhyaline,
the pattern relatively ill-defined; abdominal tergites obscureyellow,trivittate with brown; male hypopygiumwith the beakof the inner dististyle short but slender,sloping directly into
the long crest that is fringed With delicatesetae; lateral appendagewith two processes,
the lower a long slendercurved
spine; gonapophysiswith its apex obtusely rounded; eighthsterniteweakly trilobed, the setaeshort and abundant.
Male.-Length about 10-11 mm.; wing 10.5-11mm.; antennaabout4 1.4mm.
Fernale.-Length about 13.5 mm.; wing 12 mm.
Frontal prolongation of head light brown, slightly darker
above,including the long nasus; palpi dark brown. Antennae(male) relatively long; scape and pedicel yellow, flagellum
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Buttetin of the BrooklynEntomologicalSociety Tot'xLr
black; flagellarsegmentslong, only feeblyincised. Front andanterior vertex light yellow; posterior portion of head lightgray,with a medianbrown stripe.
Pronotal scutum gray, variegated with brownish gray ;scutellumand pretergitesyellow. Mesonotalpraescutumwith
the ground color gray, with three darker gray stripesthat are
narrowly bordered by brown, the broad rnedian stripe on
cephalicthird enclosingtwo similar brown lines, thesemoreor less connectedwith the outer borders behind; posteriorscleritesof notum gray,variegatedwith darker brownishgray,includingtwo areason eachscutallobe; centralarea of scutellum and mediotergitenarrowly infuscated;pleurotergiteyelgray, vaguelypatterned
low. Pleura with the mesepisternum
lvith darker brownishgray ; mesepimeronand metapletlramore
uniformly yellowed. Halteres brown, the base of stem restrictedly yellow, apex of knob vaguely pale. Legs with thecoxae yellow, sparselypruinose; trochantersyellolv; fetnora
and tibiae brownishyellow, the tips narrowly brownish black;tarsi black,the proximal portionsof the basitarsipaler; claws(male) toothed. Wings pale brown and yellowishsubhyaline,
the pattern relativelyindistinct; prearcularfield and costalregion more saturatedyellow; stigmaand a smallspotover originof Rs somewhatdarker brown ; the pale areas are best developedas a band beyondthe stignla,extendingbackwardto
veinM t * 2,rfloreor lessconnectedwith a similar areacentering
at cell 1stM z; anotherlarge pale spot beforeouter end of cell
M and including the adjoining cells; other areas still morebasadin cellsCu,and lst A; veinsbrown, more yellowedin thesaturatedportions. Venation: Rs about twice nc-c14.
Basal abdominal tergites chiefly gray ; succeedingtergites
grayishbrown medially,obscureyellow sublaterally,with conspicuousbroken darker brown sublateralstripes; lateral margins broadly, posterior borders more narrowly pale yellow;
outer segments,including most of hypopygiurn,dark brown;
mediandark tergal stripe narrowedposteriorlyand becoming
obsoletebeyondthe sixth segment; basal sternitesyellow, theouter onesslightly darker. Ovipositorwith the cerci long andslender, virtually straight, considerablyexceeding the compressed-flattenedhypovalvae. Male hypopygium having theninth tergite with its apical portion broadly sessile,the lateralangles produced laterad into long slender blackenedpoints.
Outer dististylebroad, its apex very obtuse. Inner dististyleJune,7946Bu,lletinof the Brooklyn EntomologicalSociety

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with the beak short but slender,sloping gradually upward into
the dorsal crest, the latter with a long row of setae. Lateralappendagewith two processes,
the upper a pale,nearly parallelsided blade,the lower one a long slendercurved spine. Gonapophysis slender, strongly bent on proximal half, the apex
narrorvly obtuse. Eighth sternite weakly trilobed, the setae
abundantbut short.
Hesrrer : Wyoming, Utah.
Hororvpn: 3, Yellowstone National Park, Wyoming, south
entrance, July 24, 1934 (M elond,er); in Melander Collection.
Allotoltotype,J. Poratopotypes,663; paratyfe, 3, SpanishFork,
Utah Co., Utah, altitude 4550feet (D. Elrno Hardy).
Tlris fly is most similar to the eastern North American Tipula(Orcontj,sa) grata Loew, differing in the very small size, detailsof colorationand, especially,the structure of the male hypopygium,including the tergite, inner dististyle, gonapophysesand eighthsternite. The speciesis lesslike ?. ( O.) ingrata Dietz and T. ( O.)roltzs,eriDoane,rvhich have the lateral appendageof the hypopygium
sornervhatthe same.
Tipula (Arctotipula) loganensis, n. sp.
General coloration gray; praescutum light gray with fourverv slightly differentiateddarker brownish gray stripes thatare narron'ly bordered by blackish, most intensely so as amedian vitta separating the intermediate stripes; antennae
short, the basalswellingsof the flagellar segmentsvery poorly
developed; legs dark brown, the femoral basesrestrictedlyobsctlreyellorv; claws (male) toothed; wings grayish subhyaline,
variegatedwith pale brown, appearingas a seamover cord andat lving tip ; stigma somewhatdarker brown; abdominal tergites brown, the caudal borders broadly gray, the median area
more vaguely of this latter color; male hypopygium relatively
large ; tergite extensive,its caudalmargin bilobed; outer dististyle long and slender; inner dististyle with the beak a shortcultriform blade, the margin behind this with a slender acutespine; gonapophysisappearingas a conspicuousdark-colored
oval blade, the stem short; ninth sternite with a conspicuous
protruded pale membraneat baseof the median emargination.
Mole.-Length about 11 mm. ; wing 14.8 mm. ; antenna
about 2.8 mm.
Frontal prolongation of head brown, gray pruinose, clear
above; nasus reduced to a small stub, with a concentrationof

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black setaeon and near it; palpi medium brown. Antennae
short ; scape gray pruinose, pedicel light brown, flagellum
brownish black; flagellar segmentsshort, basal enlargements
very poorly developed; verticils shorter than the segments.
Head brownish gray, the front and anterior vertex clear lightgray, including somewhatdepressedareason either side of theanterior vertex ; a vague darkening from the summit of thevery low vertical tubercle backward; occipital region restrictedly obscureorange.
Pronotum brownish gray, scutellumyellow, the central depressiondarker. Mesonotalpraescutumlight gray, with fourvery slightly differentiated darker brownish gray stripes thatare narrowly bordered by blackish, most intense as a medianvitta separating the intermediate stripes; outer margins oflateral stripes undarkened; scutum with median region lightgray, the lobes chiefly occupied by two brownish gray areasthat are narrowly borderedby more blackish lines, especiallyon their mesal edges; scutellum brownish gray, parascutella
obscurebrownish yellow ; postnotumclear light gray. Pleura
light gray, the anepisternuma trifle darker ; dorsopleuralregion broadly light yellorv. Both head and thorax unusuallyglabrous,the vestiture reducedto virtually microscopicblack
setulae. Halteres obscureyellow, the knob weakly darkened.
Legs with the coxae light gray; trochantersobscurebrownish
yellow; femora dark brown, the basesrestrictedlyobscureyellow; tibiae and tarsi dark brown ; claws (male) with a toothon proximal third. Wings grayish subhyaline,variegatedwith
pale brown, the latter appearing chiefly as a seam over thecord and at wing tip ; stigma somewhat darker brown ; areasthat are somewhatpaler than the ground before and beyond
stigma and in outer end of cell M ; veins brown. Macrotrichia
on veins beyond cord, including Rt + z to Znd A, very sparseon Mn and Lst A,' squarnanaked. Venation: Rs long, exceeding three times nr-cu,ci R, * , entire ; petiole of cell M t about
two and one-half times nc; cell 1st M z relatively small, pentagshortly
onal ; rn shorter than the basal sectionof M , * 2| 'nc-cxc
before the fork of M, * n.
Abdominal tergites brown, the caudal borders broadly gray,the median area of the segmentsmore vaguely of this lattercolor ; sternites clear gray, with yellow posterior borders,broadest on segmentsthree and four and again on sternite
eight; hypopygium dark. Male hypopygium with the ninth

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tergite extensive,the outer portion narrowed into a yellowed
apexthat is bilobedby a shallowV-shapednotch; lobesdensely
providedwith black setae;no ventral armature (as in besselsi,
protrudedpale
and allies). Ninth sternitewith a conspicuous
membraneat baseof the median emargination. Outer dististyle long and slender,gradually narrowed to the obtusetip,
the length about four times the greatestbreadth. Inner dististyle with the beak a short cultriform blade; sensorygroupplacedat baseof beak,consistingof relativelyfew areas; outermargin of style behindthe beakbearinga slenderacutespine.
Gonapophysisappearing as a conspicuousoval dark-colored
blade,the stem short. Eighth sternitetransverse,the caudal
margin pale,unarmed.
Henrrer: Montana.
HororypE : {,Logan Pass,GlacierNational Park, altitude6,655
feet, Jtrly 18, 1935 (M elander); Alexander Collection,throughMelander.
Tipula (Arctotipula) loganensisis quite distinct from the othermembersof the subgenusoccurring in the northern and central
Rocky Mountain Region. It comesclosestto speciessuch as T.
Alexander, differing(A.) socra Alexander and T. (A.) send,dea
in the colorationof the body and wings and in theconspicuously
structureof the malehypopygium. The body is unusuallyglabrous
for a memberof this subgenusbut the assignrnentseemsunquestionablyto be correct.
Tipula (Lunatipula) rainiericola, n. sp.
Belongsto the unicinctagroup, allied to wnicincta;generalcolorationof mesonotumgray,the praescutumwith four entirerich brown stripes,the intermediatepair narrowed and outbowed behind; antennae relatively short, flagellum black;fernorayellow,the tips narrowly blackened;wings with a weak
palebrown and whitish subhyalinepattern,the latter including
a major prestigmalarea; malehypopygiumwith the appendage
of the ninth sternite narrowed into a cylinder that is tippedwith a brush of reddish setae;inner dististylenarrow ; gonapophysessymmetrical.
Male.-Length about 13 mm.; wing 17-18 mm.; antenna
about4 mm.
Frontal prolongationof head light brown, sparselypruinose
above; nasus short and stout to virtually lacking; palpi withsegmentsdark brown,basalsegmentobscureyellow, succeeding70

Bu,lletinof the Brooklyn EntomologicalSocietg1vot.XLr
the incisurespale. Antennaewith scapeand pedicelyellow;
basal flagellar segmentpale brown, outer ones brownish black
to black; flagellar segmentswith small basal enlargements;
longestverticils slightly longer than the segments. Head above
brownish gray, with a more or less distinct brolvn line onvertex, best indicatedon the vertical tubercle.
Pronotum brown, pruinose. Mesonotalpraescuturnwith theground gray, with four entire rich brorvn stripes, the intermediate pair narrowed and outborved behind so the central
ground areais widenedon posteriorhalf ; posterior scleritesof
notum gray, each scutal lobe rvith two paler brown areas.
Pleura light brown, sparsely pruinose. Halteres pale, thebase of knob a little darker. Legs with the coxae and trochanters pale, yellow pollinose; trochanters yellow; femora
and tibiae yellow, the tips narrorvly blackened; tarsi black, thebasitarsipaler at proximal ends; clarvs(male) toothed. Wings
with a very pale brownish tinge, restrictedly patterned lvith
whitish subhyaline; stigrna slightly darker brown than the
ground, relatively srnall; obliterative band before cord broadand conspicuous,extending into cells M B and M +i no poststigmalbrightenirg;trvo isolatedsmallpalespotsin cell Lst A;
veins pale brown. VenatiorT:ltt,longer than the petioleof cell
Mr.
Abdominal tergitespale, more or lesstrivittate rvith brorvn,
somewhatmore extensiveand intenseon the fourth and fifth
segments,the marginsyellow ; outer segments,including hypopygium paler brown. Male hypopygium having the ninth
tergite with conspicuous,relatively broad lobes, their outerangle acute; a broad dorsal furrow ; lobes separatedby a relatively narrow U-shaped notch. Ninth sternite with its appendagea broad-basedlobe that narrows into a short cylinderthat is tipped with a brush of reddishsetae. Basistylewith theventral angle produced into a flattened sclerotized point or
blade. Outer dististyle broadly expanded on outer portion,
provided with very long setae. fnner dististyle with the beakrelatively slender; lower beak stout; posterior crest narrow,the dorsal portion pale to subhyaline,the margin microscopically serrulate; outer basal lobe broad, with numerous long
pale setaeand a few scatteredblackenedshort bristles. Gonapophysespaired, the major bladeappearingas a nearly straight
reddish spine, narrowed very gradually to the acute tip; atbase,eachspinebearsa smaller flattenedyellow blade. Eighthsternite with the lateral lobes very low, each terminating inJune,7e46Bulletin of the BrooklgnEntomological,
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77

a strong fasciculatebristle, with one or two much smallersetae
at its base; mediancushionrelatively small, the relatively few
setaeexpandedand branchedat their outer ends.
Hesrrer: Washington.
Hor,otypE: 3, Berkeley Park, Mount Rainier, August 23, 1934
(M elander); in MelanderCollection. Paratopotypes,
S'33, August23-27, L934 (M elander).
Among the relatedspeciesof the unici,nctagroup,the presentfly
is most similar to Tipula (Lunatipula) uni,cinctaDoane, which
differs in slight details of sizeand colorationand, especially,in thedetails of structure of the male hypopygium, particularly the appendageof the ninth sternite and the inner dististyle.