0~ new or little-lcnown Tilmlidte.

465

L X I I I . - - N e w , : little-lc~wwn Tipulidm ( D i p t e r a ) . ~ I i I .
Eth/opia,, SlJeeies. By CHa.~tLESP. ALEXaNDgI% Ph.D.,
Urbana, Illinois, U.S.A.
THIS paper is a contiuuation of the preceding parts under
this title. The mate,ial included herei, is based on collections
made in Cameroun br the Rev. J. A. Reis and in Eastern
Iransvaal by Mr. H. K. ~'[um'o. In ad(htlon, a few species
from the ~Paris Museum sent to me for determination by theCurator of the Diptera, Mons. Eug. Sdguy, have been
included. The holotyt)es are preserved in the collection of
the write U unless stated otherwise.
2~.MPHILIMNOBIA~gen. nov'.
[lead moderately large, tile eyes separated Loth above andbelow. Legs long and slender; tibiae unspurred; claws
small, simple, subterminal, empodia pres~mt. Wings with
wdn ~% very lo,g~ Scl endi,g opposil~e cross-vein r ; Se~ farbe/~,re the tip ot~ 8'q, the latter vein alone being about equal
to the basal deflection of/?4+,s ; r preset~t~ located far before
the tip of RI~ this portion of 1il beyond ~' being a little longer
thau the deflection of R4+s ; cell 1st M= open by the atrophyof tim outer deflection of Me; basal dettecdon of C'uI at thelbrk of M; anal veins long and straight. Mate hypopygiumwith the ninth tergite small, the caudal margin feebly concave ; pleurites moderately elongated, the proximal face witha cushion of numerous short spinules; pleur'd appendages
two, the outer appendage shortes b flatteued~ the apex obtuse ;inner ph'ural appendage larger and more elongated~ irregularly twisted, the apex with a few setigerous tubercles; nodistinct gonapophyses; penis-guard a subflattened lobe ofnearly mdform width for its entire length. Ovipositor withthe valves moderately elongat% powcrfu.l~ the tergal valvesmuch exceeding the rather weak, acutely pointed sternalvalves.
Gonotype.~Amphilimnobia leucopeza~ sp. n. (Western
Ethiopian liegio,).
The curious fly that is made the type of the above new
genus is represented in the writer's collection by three speclmens in poor condition, The fly possesses structural features
lhat preclude its disposition in any genus known at t~hepresent time~ Ahhough the gencrai appearance of the fly ismuch like a 2D&ranomgia, the details of structure seem to
A,m. & 3Iag. N. llgt. 8er. 9. Vol. v.
32

466

]Or. C. 13. Alexander onplace if: in a position much nearer JDicranaptyctsa and Rhamphi&'a. The principal generic characters are derived fromthe claws, the structure of the male and female genitalia, and,especially, the wing-venation. Unfortunately the tips of theantennm are broken and the number of segments cannot be
accurately determined.

Amphilimnobia leucopeza, sp. n.
General coloration dark brown ; legs dark brown, the tarsi
white ; wings with a strong brownish tinge; Sc long, cell
1st M~ open.
Male.--Wing 7"8 ram.
.Female.--Length abou~ 8 ram. ; wing about 6"4 mm.
Rostrum, palpi, antennae, and head dark brown.
Thorax dark brown. Head dark brown. Legs brown;fore tarsi, except the basal two-thirds of the metatarsus, purewhite ; the other legs are all detached ; in some the white ot~
the metatarsus is much more restricted, occupying only thenarrow apex, in others the entire tarsus is white excepting
only the base of the metatarsus. Wings with a strong
brownish tinge, the wing-tip more suffused; stigma a little
darkm" brown than the remainder of the wing~ elongate-oval ;veins dark brown. Venation as described under the genericcharacterization; Rs long', strongly arcuated or even angulaled and spurred at origin ; r on R2+8 nearly twice its length
beyond the fork of Rs ; r-m less than one-half the deflectionof Ra+5 ; petiole of cell 2nd J4~ approximately as long as thiscell ; the abortive vein behind Cu distinct.
Abdomen brown, indistinctly bicolorous ; posterior marginsof the segments narrowly darker.
I-lab. Cameroun.
tlolotype, c~, Olama, altitude 2000 feet, July 24, 1919

(J. A. Reis).
Allotopot2¢pe, 9, in copula with the type.
Paratopotype, a fragment.
The type is mounted in balsam.

.Erioptera (Erloptera) carior~ sp. n.
Antennm with the scapal segments brownish yellow;
flagellum dark brown; vertex brownish grey, broadly~hitish adjoining the inner margins of the eyes ; mesonotum
shiny dark brown ; legs yellow ; wings pale brown~ the costaland subcostal cells and a narrow band at the cord light
2yell°w; male hypopygiuru with the inner pleural appendage
unbranched (branched in carissima), tim head enlarged into

new or llttle-known Tipulldse.

467

a structure that resembles the head and beak of a crested
bird ; gonapophyses deeply bifid.
~I, le.--Length about 5 mm. ; wing 4'3 mm.
Rostrum and palpi dark brown. Anteroom short, the scapal
segments brownish yellow; flagellum dark brown, verticils
only moderately elongated. Vertex brownish grey, broadly
whitish adjoining the inner margins of the eyes ; eyes of themale very large as in this group (pdrlngue2/i group) of species,
broadly contiguous beneath.
Mesonotum shiny dark brown, the prsescutum wltll thethree usual stripes confluent, the humeral region and lateral
margins a lltt]e paler yellowish brown. Pleura dark brown.
Halteres brown~ the knobs dark brown. Legs with the coxse
and troehanters brown; remainder of the legs light yellow,
the distal three tarsal segments and the tips of the others(lark brown. Wings pale brown, sparsely marked with dull
yellow ; the costal and subcostal cells are light yellow ; thestigma dark brown, elongate-oval, and very distinct; thecells proximad of the cord are largely pale brown, thosedistad of the cord dull yellow, the veins broadly seamed withbrown ; the cord is narrowly margined with yellow. Venation as in E. carissima, but the deflection of Cul is more
perpendicular.
Abdomen dark brown. Male hypopygium very distine~from either E. pdringueyi or E. carissima ; outer appendage
slender, as in carissima; inner appendage with a short,
twisted stem that is inserted in the elongated, transverse headnear the middle of its length ; this head somewhat resembles
the head and beak of a bird with a long, curved crest ; ofthis structure, the compressed, obliquely truncated beak is
directed laterad; the curved crest is directed proximad andcephalad, covered with numerous short erect setse. Gonapophyses deeply bifid, the proximal arm longest, the tips
acute and blackened, subdeeussate with their fellow of the
opposite side; the lateral arm is a little shorter, largely
blackened, suddenly narrowed into the short, acute tip.
Hab. Eastern Transvaal.
tlolotype, (~, " Lot 30, De Kaap Block B," near Kaapmuiden, Ocgober 9, 1919 (H. K. Munro).

Erioptera nigrolatera brev@ilosa, subsp, n.
f'ernale.--Wing 5-5"1 mm.
Generally similar to typical aigcolatera~ Alexander (Nyasaland), differing as follows : Legs without the conspicuous erec~ pubescence found in
3:2~

Dr. C. P. Alexander on468n;grotatera; this may be a sexual character, however, as
only males of typical n~qrolatera and females of the now
variety have been made k,mwn. In the present form the
pubescence of the legs is very short, appressed, and not at allconspicuous. The wings are much paler, greyish subhyaline ; vein Sc considerably shorter, ending far before thefork of R2+~ ; S% is closer to the tip of Scl, SCl alone being
only about two-thirds R2+3; i n typical nigrolatera SCl islonger than R~+3 ; anal veins a little more divergent at thewing-margin. The valves of the ovipositor are short andfleshy as in the group.
flab. Eastern Transvaal.
Holotype, ?, "Lot 30, De Kaap Block B," near Kaapmuiden, October 11, 1919 (H. K. ~llunro).
Paratoio~,t~ype, ? , a badly injured specimen~ October 10,
1919.
Molophilus camerounensis, sp. n.
Rostrum and pa]pi black ; antenme of tile male elongate,
the basal segments yellowish ; general coloration dark brown,
the mesonotal pr~eseutum brighter brown; wings greyish,
tile veins clothed with long dark-coloured hairs ; male hypopygmm with three conspicuous chitinized appendages oneither side.
l]£ale.--Length 4"8 mm. ; wing 4q: ram.
Rostrum and palpi black. Antennee of the male elongated,almost as in the Nearctie M. pubipennis, O.-S., the basal
segments light yellow~ the distal segments a little morebrownish; flagellar segments elongate-eylindrieal~ clothed
wittl long. pale, outspreadin~ hairs Head dark brown.
Mesonotal prmseutum rather light brown, the remainder ofthe mesonotmn and the pleura dark brown. Halteres darkbrown. Legs with the eox~e and troehanters brown; remainder of the legs dark brown. Wings with a strong grey
ting% the costal and subcostal cells more brownish; veins
al~d pubesceuce dark brown, giving a dark eolour to thewings. Venation : R8+4+5 long, about one-half longer thanthe basal deflection of Cul ; fork of M far proximad of thefork of _Rs; inner ends of cells 22~ and i ~ in alignment;
petiole ot~cell M8 more than three times the basal deflectionof C,q or about two-fifths of vein CuI beyond i t ; basal
deflection of Cul perpendieular~ nearly twice as long as thebasal deflection of M3.
Abdominal segments brown, indistinctly bicolorous. Male
•

(' ~

•

new or little-known Tipulid~e.

469

hypopyglum with the armature unusually developed even fora member of this genus ; three powerful chitinized appendages
on either side, tt~-e longest and most powerful of which isenlarged apieally into a two-edged complicated head.
Hub. Cameroun.
Ilolotype, (~, Olam% altitude 2000 feet~ July 24, 1919

(3". A. Reis).

C,ERATOLIMNOBIA~gen. nov.
Vertex behind the antennal bases with a flattened whi~elobe or cornlele. Tibiae unspurred. Wings with So long',
8c2 close to the tip of Sc, ; tip of R l atrophied or nearly so ;cell 1st M~ closed ; basal deflection of Cul before mid-lengthof cell 1st M:. Male hypopygium small, with two pleuralappendages, tile longest recurred into a U, at the bend ofwhich is a cylindrical minutely branched arm.
Genotype.--Ceratolimnobia munroi, sp. n, (Southern Ethiopian :Region).
By means of all existing keys to the Tipulida~ this cranefly would run to the genus Gnophomyia, to which it isobviously not closely allied.
Gnophom.~ia is gradually
becoming a receptacle for many diverse Eriopterine elements,
and it seems advisable to remove the present fly from suchan incongruous gathering and erect a separate genus for it.
The presence of a snowy-white corniculus, much as in the
Ethiopian species of the genus Ceratocheilus, Weseb6, offers
the most ready means for distinguishing this genus fi'om
similar crane-flies.

Czratollmnobla munrol, sp. n.
Head light grey~ with a snowy-white corniele on thevertex; general coloration dark brown~ the mesonotal pr~escutum narrowly margined laterally with silvery white ; legs
dark brown, the tips of the femora and tibiae narrowly white,
the fore tarsi largely white ; wings dusky, tile costal region
more yellowish, with six large dark brown blotches; veins
beyond the cord broadly seamed with brown.
Male.--Length about 2"7 ram. ; wing 3"8 ram.
Rostrum an~ palpi black.
Antenua~ witlt the scapal
segments black ; flagellum bvokell. Head with a light grey
pubescence, the middle of the vertex a~,d tile postge,~m more
blackish ; on the lore part of the vertex immediately behilld
the autennai bases and between the anterior end of the eyes

470

Dr. C. P. Alexander on

is inserted a flattened snowy-white lobe or conllculus that is
directed cephalad, very much as in the African species of

Ceratochei/us.
Pronotum dark brown. Me~onotum dark brownish black,the lateral margins of the pr~escutum narrowly silvery white,
this beginning on the sides of the pronotum, continued backward to the wing-root; tuberculate pi~s a short distancebehind the anterior margin of the pr~escutum. Pleura andsternum dark brownish black. Haltores pale brown basally,
the knobs broken. Legs with the coxa~ dark brownish black ;~roehanters dark brown ; femora dark brown, darkest before
the tips, which are abruptly and narrowly white; tibimbrown, the tips narrowly white; fore tarsi with the basal
two-thirds of ttle metatarsus brown, tile remalndor of tile
tarsi white; the other legs are detached, but the middle andhind tarsi are presumably uniform brown, and what areprobably the hind logs lack the white tibial apices. Wings
with a strong brownish tinge, especially on the basal half ;beyond tile cord the membrane is subhyaline, with the veins
I,roadly seamed with brown ; six darker brown costal blotches,tile first at the areulus, the second at the origin of R 6 thethird at Scl, tile fourth at tile tip of R1 ; the fifth and sixth
at the ends of veins R: and R3; costal and subcostal cells
more yellowish; pale areas occur between the dark costal
areas, at tim ends of cells R, M, Cu, 1st A, and 2rid A ;veins dark brown, the pale spaces on veins C, Se~ and R
yellow. Venation: Sc long, ending about opposite twothirds the long Rs, Sc~ close to the tip of Scl, Scl equal toabout twice Sc2 ; tip of R1 atrophied ; r oblique, inserted at
the fork of R:+a; Rs longer than Ra; R~+a a ]itde shorter
than R:; inner ends of cells R3, R6, and 1st M~ in directalignment; r-m a little longer than the basal deflection of
R4+n; cell 1st M2 long and comparatively narrow, longer
than vein CuI beyond it; basal deflection of Cul just before
mid-length of cell 1st M2.
Abdomen dark brown. Male hypopyglum small; pleurites
rather short; and stout~ covered with microscopic hairs and a
few long setm ; two pleural appendages, the longest; strongly
recurred into the fl)rm of a tJ, the fl'ee end directed outward,
the tip narrowed and acute ; on the side at the bend of the tJ
with a conspicuous cylindrical arm that is minutely branched;
inner pleural appendage small and flattened, narrowed to theblunt apex, which bears several long setm; gonapophyses
small, flattened, triangular, directed proximad.

new or little-known Tiputidee.

471

Hub. Eastern Transvaal.
Holotype, g', " Lot 30, De Kaap Block B," neat' Kaapmuiden, October 8, 1919 (H. K. M,,nro).
It is with great pleasure that this interesting new species
is dedicated to its discoverer, Mr. H. K. Mu~aro.

Trentepohlia ( Trentepohlia) auvantla, sp. n.
Generally similar to T. speiserl; mesonotal prse~cutum
light orange-yellow, with a delicate reddish-brown mediau
line; legs yellow; posterior tibise with two strong black
bristles before the tip ; abdomen yellowish brown.
Male.--Length 7"6 ram. ; wing 7 ram.
Female.--Length 8"8 ram.
Described from alcoholic specimens.
Rostrum and palpi pale brownish yellow. Antennm withthe scape light brown ; flagellum broken. Head brownish.
3¢Iesonotal pr~escutum light orange-yeltow~ with a delicate
reddish-brown median line, tile lateral margins behind ttle
pseudo-sutural fove~e narrowly infuseated ; scutal lobes
reddish brown, the median area and the seutellum pale;postnotnm yellowish brown. Pleura brow% darker on thesternum. I-Ialteres pale, tile knobs white. Legs with tile
coxae yellow; trochanters dull yellow; femora and tibkeyellow, tlle tips not darkened ; tarsi broken ; posterior femorawith a series of six or seven short blackened spinules at thebase ; tibise with two widely separated black bristles before
the tip. Wings subhyaline, the costal margin strongly
yellowish, the anal cells dusky grey ; a rather heavy brownpattern, somewhat as in T. speiseri, bu~ more restricted;
the blotch at the origiu of Rs is entirely solid, bu~ comparatively small; the apical mark includes only the veins, thecentre of cell R2 yellowish subhyaline ; vein Cu seamed withdarker.
Abdomen yellowish brown, the apical segments darkened.
flab. Reunion.
Itolot~pe, ~, 1903 (Blanchard).
Allotopotj/pe, ?.
Type in the collection of the Paris Museum.
This species is~ perhaps, closest to T. spelseri, Edwards,
fi-om which it is told by the bright colour of the mesonotum
and the different wi~g-pattern.

Tipula ellenbergeri, sp. n.
Head indistinctly reddish ; thorax black ~ wings with the472

On new or llttle-known ].'ipulidm.

eostal margin and apical half brownish black, tile basal half
whitish hyaline; basal abdominal segments reddish, theposterior margins black.
~ex ? (probably a male).--Win z 14"6 mm.
Described fi'em an alcoholic specimen.
Front;al prolongation of head comparatively short, tlte sides
leddish brown, the dorsum narrowly darker ; nasus distinct ;palpi dark brown. .A.ntennte dalk brown, 1he flagellar segments more reddish basally ; the flagellum is broken, butdelaehed segments in the vial with the type presumably
belong to this speei.es ; in these the distal flagellar segments
are elongate-oval~ slightly constricted basally. Head reddish,dmker-coloured between tile eyes.
Thorax uniformly deep black, apparently shiny in dried
specimens. Halteres black. Legs blaek~ the femora andtibiae slightly paler basally ; legs slender, es?eeially the tarsi ;claws reddish, tile distal half blackened, at about mid-lengthwi~;h a small, acnt% ventral tooLh. All of the legs aredetached in the vial, and but few still remain. Wings wiih
the costal region and lhe apical half of the wings black, thebasal half conspicuously and abrut,tly whitish hyaline ; ~hedark apex includes all of the cells beyond the cord as well ,asthe outer ends of cells R, ~1[, Cu, and 1.~t A ; stigma small,dark brownish black ; veins dark brown. Wings broad.
Abdomen with the first segment black; second segment
reddisl b the caudal margin narrowly blackened ; third segment
reddish basally, the apex o~ the abdomen broken beyond thispoint.
Hab. French Congo.
Hol~tyt)e~ sex ~ ?, Lambs,dad, Ogoway R., 1911 (R. Ellen.
berger).
Type in the collection of the Paris Museum.
q'his beautiful crane-fly is readily told from all described
African q'ipulin~e by the b~'aulifitlty d:midiate wings. The
claws bear a smal]~ acute, ventral tooth that is only visible
when the foot is viewed fiom the side. The presence of thistooth would indicate that this fly is a male. This interesting
species is named in honour of its collector.