[Reprinted from Insecutor Inscitie Menstruus, Vol. VIII, Nos.7-9,19201

NEW SPECTES OF JAPANESE CRANE-FLIESPARTI(Diptera, Tipulide)
Bv CHARLES P. ALEXANDER
The following undescribed species of Japanese Tipulide
were sent to me by Mr. Ryoichi Takahashi, who collected themin the vicinity of Tokyo. I wish to express my thanks to Mr.
Takahashi for this and other very valuable crane-fly material.
The types are preserved in the collection of the writer.
Genus ERIOPTERA

MeigenSubgenus
EMPEDA OstenSacken
Erioptera (Empeda) japonica, new species.
General coloration gray; antenne brownish black throughout ; wings light gray, the stigma a little darker; Sc, unusuallylong for an Empeda; R, generally parallel to R.; cell rst M,open.
Fernale.-Length, 5 mm.; wing, 5.2 mm.
Rostrum and palpi dark brown. Antenna brownish blackthroughout. Head light gray.
. Pronotum gray. Mesonotum gray, the praescutum withthree ill-defined brownish gray stripes, the lateral and anterior
margins indistinctly flesh-colored. Pleura light gray pruinose.
Halteres light brown, the knobs darker. Legs with the coxegray; trochanters light trrown ; remainder of the legs brown.
Wings light gray, the stigma indistinctly darker; veins darkbrown; veins with rather long and conspicuous macrotrichia.
Venation: Sc, ending a little beyond two-thirds the long andnearly straight Rs,'Sc, unusually far from the tip of Sc, fora member of this subgenus, Sc, alone being but little shorter
than the deflection ol Cu, and longer than that section ofRrf,
before r; r on Rr-1-, at about one-third its length;
R, running generally parallel to R, and but little shorter than
this vein, so that iell znd, R, at the wing-margin is about equal
to cell Rr; cell rst M2 open; basal deflection ol Cu, at thefork of M.

I NSECUT OR INSCIT IE

M ENST RUU S

135

Abdomen dark brown. Ovipositor with the valves horncolored, the tergal valves rather strongly upcurved, the tips
acute,Ha.bitat.-Japan.
Holotype, 9, Tokyo, December3, 1919 (R. Takahashi).
Genus PSILOCONOPA

Zeterste<lt

Psiloconopa verna, new species.
General coloration black; lateral margins of the pronotal
scutellum and the posterior margin of the mesonotal scutellum
yellowish; knobs of the halteres conspicuously bright yellow;
wings grayish subhyaline; cell Ist M2 open by the atrophy ofm; abdomen black, the caudal margins of the segmentsyellow.
Mqle.-Length, about 3.5 mm.; wing, 4 mm.
Rostrum and palpi black. Antenna black, the flagellar segments oval. Head black.
Pronotum black, the lateral margins of the scutellum brightyellow. Mesonotum black, sparsely brownish pollinose; posterior margin of the scutellum obscure yellow; postnotum deep
black. Pleura black; a tiny yellow spot at the wing-root. Legs
entirely black. Halteres black, the base of the stem morebrownish, the knobs conspicuouslylight yellow. Wings grayish
subhyaline; veins dark brown. Venation: Sc long, Sc, extending to opposite midlength of Rrfu; Sc, far before the tip ofSc' the latter vein alone being but little shorter than'Rs,'Rslong, straight; Rr*, short, a little longer than the deflectionof Rn-1u; r on R, about its own length beyond the fork ofRr*ri cell rsf M" open by the atrophy of m; basal deflectionof Cu, subsinuous, located at the fork of M,' fusion ol Cu,and,Mu about equal to the basal deflection of the latter; analveins clivergent.
Abdomen black, the posterior margins of the tergites yellow,narrow and less distinct on the basal tergites, very distinct on
segments 6 to 8; sternites similar but with the yellow margins
very narrow and indistinct; hypopygium small, black.
Holotype, 6, Tamagawa, Tokyo, Japan, March 20, 1920
(Ryoichi Takahashi).

I ;i (iIN S N C U TOR IN S C ]TI,' I]

Genus CLADURA]\{ I.:N S I' R U LTS

Osten Sacken

Cladura nipponensis, new species.
Head dark gray; antennal flagellum dark brown; legs palebrown, the tips of the femora ancl tibia scarcely darkened;
abdomen dark brown; male hypopygium with the pleurites
long and slender, nearly twice as long as the small pleuralappendage.
Male.-Leneth, about 5.3-5.5 mm.; wing, 6.5-6.? mm.
Rostrum and palpi pale brown. Antenne with the scapal
segments pale brown, the long-oval flagellar segments darkbrown; flagellum with thirteen segments, the basal f usionsegment composedof two segments,equaling in lengtl-rthe second and third taken together. Head dark gray.
Mesonotum pale yellowish testaceous,the praescutum withthree confluent, darker brown stripes. Pleura pale yellowish
testaceous. In the paratype the praescutum is not so dark incolor. Halteres pale. Legs with the coxe and trochanters
pale; femora brownish yellow, the bases more yellowish, thetips scarcely darker; tibie pale brown, the tips narrowly andindistinctly darkened; tarsi dark brown ; pubescenceof the legs
not conspicuous. Wings nearly hyaline, the base and cells C
and Sc' more yellowish; veins brown; macrotrichia on theveins of moderate length only. Venation: Sc, ending beyond
the fork of Rr1r, Sc, a very little before this fork; r somewhat indistinct; Rrf, about equal to that section of R, beforer,'fusion of M"and, Cu, about equal to the outer deflectionof
Mu alone; petiole of cell M, about equal to the basal deflectionof Cur; znd Anal vein ending about opposite the base of Rs.
Abdomen dark brown throughout. Male hypopygium withthe pleurites very long, cylindrical, the single pleural appendage
only a little more than half the length of the pleurite, slender,
tapering gradually to the apex; gonapophyses in the form ofslender, chitinized hooks.
Habitat.-Japan.
Holotype, 6, Tokyo, November 16, 1919 (R. Takahashi).
Allotopotype, 9, December9, 1919.
Paratopotype, 6, November 16, 1919.

INSI' :CU' I' OR INSCIT I,4 l

M ENSTR U U S

137

Cladura autumna, new species.
.

Head and antennre yellow; general coloration yellow, themesonotum without darker markings; f emora and tibia yellowish, narrowly tipped with dark brown; abdomen obscureyellow, the tergites indistinctly ringed basally with brown;
male hypopygium with the pleurites and appendage stout.
Male.-Length, 5.5-6 mm.; wing, ?-8 r.nm.
Femnle.-Length, about 6 mrn.; wing, ?.8 mm.
Generally similar to C. nipfonensis, differing as follows:
Larger. Antennre pale, with long, black verticils, the terminal three or four segmentsdarker. Head obscure yellowish.
Thorax shiny yellow, the mesonotum without distinct darkermarkings. Legs with the femora and tibie yellow, the tips narrowly but conspicuously dark brown; legs clothed with rather
conspicuous, semi-erect hairs. Wings subhyaline; veins darkbrown, with long, conspicuous macrotrichie. Venation: Sc,
far before the fork of Rrar; basal deflection of. Cu, usuallyat or before the fork ol M,, the fusion of M" and Cz, beingextensive.
Abdomen obscure yellow, the tergites indistinctly ringed
basally with brown. Male hypopygium with the pleurites shortand stout, the single pleural appendage very short and stout,
the rounded apex densely set with numerous short spinous
sete. Outer gonapophysesshaped like broad, flattened wings,
the outer margin fimbriate, the two apophysestogether appearing somewhat like a bird in flight.
Habi.tat.-Japan.
Holotype, 3, Meguro, Tokyo, October 22, L979 (R. Takahas hi) .
Allotopotype, 9.
Paratopotype, 6.
This species is also very close to Crypteria jaf onico Alex.,
which is very probably likewise a Cladura. This latter species
is still known only from the type females and the male will benecessary to secure an exact definition of the characters.
C. japonica (Alex.) has the wings distinctly although faintly
tinged with yellowish; the macrotrichire on the veins are shorter

138

IN S TC U I' OR IN S C ITIII

I\{ E N S TR U U S

and relatively inconspicuous; Sc, is almost opposite the f orkof Rrlr;
cell rsf M" broader and the basal deflection ofMr*, consequently longer; basal deflection of. Cu, some distance beyond the fork of M as in C. ni.pponensis. In bothC. auturnna and C. japonicaa,the second anal vein adds between
one-fourth and one-half the length of the long sector.
Genus TIPULA

I,innaeus

Tipula tokionis, new specics.
Ferutle.-Length, 25 mru.; wing, 20.5-21.5nrm.
Superficiallyvery similar to 7. sa,itamae
Alexander (Japan)
from which it differs as follows:
Head much darker brownish gray. Pronotum not yellow
but distinctly suffused with brown. Mesonotal praescutum
without the grayish cast in T. saitarnae. Legs with the tips ofihe femora and tibia narrowly but distinctly darkened. Wings
with the ground-color more grayish than yellowish. Abdominal tergites with a broad, blackish sublateral stripe on eitherside. Ovipositor with the tergal valves slender, almost
straight; sternal valves very broad and compressed, the tips
evenly rounded.
Habitat.-Japan.
Holotype, 9, Tokyo, September 15, 1919 (R. Takahashi).
Paratopotype, 9, August 16, 1919.