Vol.

42, No.

1969

1, January,

83. . . ," as do the discussion and figures by Hennig
to Notomyza(1958, p.
630, figs. 231, 234, 238, 239, 242). Thus, I am now of the opinion that
Cinderella must be considered a member of the Heleomyzidae
in thebroader sense in which Malloch
treated that family(1933, p. 178-198)
when he described Notomyza.

LITERATURE CITED
der Diptera
und ihre phylogenetischen
1958. Die Familien
W.
Schizophora
Beitr. Entomol.
8: 505-688.
Verwandtschaftsbeziehungen.
British Museumof(Natural History),
Malloch,J. R. 1933. Acalyptrata.In
Dipteraand South Chile, Part VI, Fasc. 4: 177-391,
Patagonia
pis. 2-7.
G. C. 1949. A new anomalous
Bull. Brooklyn
Steyskal,fly (Diptera).
acalyptrate
Entomol.
Soc. 44: 134-137.
of the Dipteraof America
north of Mexico.
U. S.
Stone, A., et al. 1965. A catalog
276: i-iv,
1-1696.
Dept. Agr., Agr. Res. Serv., Agr. Handbook
Hennig,NEW SPECIES OF CRANE FLIESFROMTROPICALAMERICA.
1
Tipulidae)(Diptera:
P.

Charles

Alexanderof Massachusetts,

University

V.

Amherst,

01002

Mass.

Abstractof craneflies are described,
fromspecieschiefly
Panama,including
Orimarga(Diotrepha)
Polymeraluteipleura,(Polymera)
cingulata,
AtarbaAtarba(Atarba)fieldiana,(Atarba)panamensis,
Gonomyia
andstenorhabda,
Gonomyia(Gonomyia)(Gonomyia)
stylacantha,
fromandGonomyia
uncinata,(Paralipophleps)
Panama,Gonomyia
from Grenada,Lesser Antilles.
(Lipophleps)
minutistyla
New

Virtually all of the included species are from Panama, where they
were collected by my friend and former student, Lt. Col. Gordon Fieldof Rhode Island, Kingston.
of the University
One further species isfrom the Lesser Antilles,
taken by Noel L. H. Krauss of Honolulu.
I
express my continued appreciation to the collectors for cooperation in thesurvey of the crane flies of Tropical America. The types are preservedin the Alexander Collection of Crane Flies.
Orimarga
Thoraxclear

weaklyinfuscated;

1
Accepted
Journalyellow,offor publication
theKansasespeciallyantennal
AugustEntomologicalnewluteipleura,(Diotrepha)thepleura,

dark

scape

centralspeciesarea

brown,

remainder

42:83-90.

January,

of

notum

obscure1, 1968.
Society

1969.

84

of

JournalKansasEntomologicalSocietyyellow; legs whitened,tips of femora and bases and tips of tibiae blackened; wings yellowed, prearcular and costal fields clear light yellow,
veins very pale brown; macrotrichia
of veins excepting costa virtuallylacking, vein R, + 2 atrophied; abdomen long, brownish yellow, extreme
posterior borders of segments light brown.
Female.

LengthRostrumbrownish

scape

dark

Head

light brown.
andregionyellow

7.8-8

black,

remainderbrown,

Cervical
clear

about

laterally,

mm;

wing

palpi

black.

obscuremm.

4.8-5.2

Antennaebrown.
dark
pronotummedianregionweaklythe elongate

withsegments

flagellar

yellow;

Mesonotaloval.

praescutumremainder

infuscated,ofnotum pale brown, the sides broadly yellow. Pleura uniformly clear lightyellow. Halteres with stem light yellow, knob weakly darkened. Coxae
and trochanters light yellow; remainder of legs white, the femoral tips
tibial bases and tips more narrowly black, about one-third
blackened,
the femoral darkening. Wings
yellowed,
prearcular and costal fields
clear light yellow; veins very pale brown, in the costal region more yelof veins
against the ground. Marcrotrichia
lowed, poorly differentiatedin numbers,

reducedvery

a costal

includingseries

and

one

or

two widely

Sc long, Sc?
separated trichia on distal section of M1 +2. Venation:of the very long Rs; R2 curved into
ending about opposite midlength
R2 + 3
short.

r-m.

opposite

Abdomen
ders

of

large

and

obscurelong,

segments

+2

R1

light

conspicuous,

cells

atrophied;thebrownishyellow,
subterminal

brown,
especially

the

cord

beyondextremerelatively

posteriordarker.
segment
cerci
small
hypovalvae,borOvipositor
slender.
andHolotype
female, Fort Gulick, Panama, January 21, 1954 (Gordon
1 broken
2 females, January
1954;
13-21,
Field).
Paratopotypes,
specimenofunknown

sex,

Barracks,

TabogaPanama,December

10,1953 (Gordon Field).
Other members of the subgenus having the leg pattern somewhat asofin the present fly include Orimargaarawak Alexander,(Diotrepha)of northern South America;
Jamaica; O. (D.) atribasis(Alexander),O. (D.) mirabilis
and the type of the subgenus, the Nearctic(OstenSacken), none of which have the clear light yellow thoracic pleura andall differing further among themselves in venation and trichiation of thewingveins.

new species(Polymera)
cingulata,
Polymeraantennae of male relatively
Rostrumshort,
brown, palpi yellow;
slightly exceeding in length the body or wing; yellow, central parts of
the intermediate flagellar segments brownish black to produce a bicoloredappearance;mesonotalandpraescutum

rites of notum and the pleura brownish
conspicuous

fused withbrown

subterminal

ring,

scutumblack;
tarsi

brown,

legs yellow,

yellow;brown, Sc short, Scx ending opposite

wings

posteriorscle

femora with astrongly

fork of Rs;

suf

R^+o andVol.
i?2 +3 subequal;

42, No.

abdomen1969

1, January,85
region more

its pleurallight brown,

black

ened.
Male.

about

LengthRostrumbrown,

4.5

palpi

4.6
wing
Antennaemm;
yellow.

about

5 mm.

relatively

short,

annenna

mm;
of

male

first
slightly exceeding body or wing; scape and pedicel light yellow;
flagellar segment obscure yellow, slightly darker outwardly;
succeeding
segments bicolored, brownish black, the narrow base and slightly broader
outeramountend whitened,oflatter

color

more

on

reducedintermediate

intermediate part of flagellar
segments, outer ones uniformly darkened;
segments slightly binodose, each with two separate groups of very longsetae that are slightly longer than the segments. Headoutspreadingdarklarge,
Mesonotalbrown.

notum

the

and

praescutum

andbrownish

pleurascutumbrown,
Halteresblack.

sclerites

posteriorofinfuscated.

moderately

Coxae and trochanters light yellow; all femora yellow with a conspicu
ous subterminal brown ring that is slightly wider than yellow apex;
tibiaeobscuredarkeryellow,

suffusedstrongly

over

cloud

very

tips

with
anterior

brown.

Wings

yellow.

of a reduced

indications

veins

cord;

tarsi

darkened,

narrowlybrown, with

Venation:Scslightly
Sci

short,

ending opposite fork of Rs, Sc2 near its tip; Ri +2 and R2 +s subequal;
7^2+3+4 subequal to nearly twice the basal section of R5; cell M\
m-cu

small;
ments

to

close

Abdomen

fork

of M.

light brown,

region more

pleuralblackened,

outer

seg

broken.

19, 1954 (Gordon Field).
Holotype male, Pina, Panama, Jaunary
a broken male, April 20, 1954.
Paratopotype,
Polymera{Polymera) cingulata is quite distinct from other regional
members of the subgenus in the combination of bicolored antennae, pat
tern

of

the

and

femora,

the

short

vein

Sc.

Other

such

species

that

have

the tips of the femora yellowed or whitened
included P. (P.) albogenua
lis Alexander
and P. (P.) geniculata Alexander, which otherwise arequite different flies.
AtarbaThorax

fulvous(Atarba)headbrown,

newfieldiana,
darker;speciesantennaeofmaleelongate,

extreme tips offlagellar segments with short vestiture;legs yellow,
femora blackened; wings brownish yellow, stigma pale brown; abdomen
subterminal
with outer
segments brown; male hypopygium
yellowed,
a
the
outer
half
with
terminal
long
dististyle blackened,spine and two
separatedsmall,

pointsnarrowed

on

outer

gradually

margin,to slender

inner

style

slender;aedeagus

very

apex.

Male.
Length about 6 mm; wing 5 mm; antenna about 3.7 mm.
Female. Length about 5.5 mm; wing 5.5 mm.
Rostrum yellow, proximal segments of palpi light brown, outer seg
ments

darker,Antennaewithterminalone

scape yellow,

subequal

to preceding

flagellum black,in male

long,

two

combined.

incisures of proxi

of

86

Journalmalthree or four segments

ance

more

KansasEntomologicalrestrictedly

Societyin female bicoloredpale,

appearin malesix or sevensegments;
proximal
and
incon
but
vestiture
dense
short
segments
long-cylindrical,asas
to one-sevenththeone-sixth
seg
long
approximately
involvingevident,

flagellar

spicuous,ments.
Headbrown.

Thoraxalmostbrown.

fulvous

uniformly

stem

with

Halteresobscureyellow, knob dark brown. Legs with coxae fulvous, trochanters yellow;
outer tarremainder of legs yellow, extreme tips of femora blackened;sal segments dark brown. Wings brownish yellow, prearcular and costal
fieldspaleyellow,

stigma

pale

veins

elongate;brown,

brown.

pale

Longi

tudinal veins beyond general level of cord with numerous trichia, Vena
tion: Sc short, Sct ending before origin of Rs, the latter short andstraight, subequal to basal section of R5, slightly longer in female; m-cu
at

from

about

to one-sixth

one-fourthM^+4.

Abdomen of male yellowed, proximal five segments restrictedly pat
terned laterally and subterminally with pale brown, segments 7 and 8with posterior half of 6 darker brown to form a ring; hypopygium
light
In

yellow.

female,abdominal

tergites

clearersternitesbrown,

yellowish

yellow.
hypopygium with tergal plate subquadrate, outer lateralangles produced laterad into acute points. Basistyle on mesal face with alobe. Outer dististyle
very small subterminal
blackened,distinctive,
outer half slightly expanded, the terminal beak very slender, outer apicalof style on outerangle with a single strong spine, at about midlength
margin with one or two small points; inner style slightly longer, appear
Male

ing as a slender
rowed
graduallyitsstraight
rod,
to slender
apex.

tip

obtuse.

very

Aedeagussmall,

nar

23, 1953
2,000 feet, December
Holotype male, El Valle, Panama,(Gordon Field). Allotopotype
female, pinned with type.
I take pleasure in dedicating this fly to Dr. Gordon Field, to whom
I am indebted for many new and rare Tipulidae
from Central America.
is no

There

closely

relatedThe

species.

dististyle of the hypopygium
ospina Alexander, of Bolivia,
distinct.

generalofappearance

the

outer

is somewhat as in Atarba (Atarba) laterfeatures are quite
but the other hypopygial

new species
Atarba(Atarba)panamensis,
Small (wing of male nearly 5 mm);general coloration fulvous yel
low, abdomen with a broad subterminal darker ring; antennae of malesetae; legs brownish
elongate, flagellar segments with long outspreading
fields clearer yel
andcostalbrownishyellow; wings
yellow, prearcularwith
low; Sc^ ending nearly opposite origin of Rs; male hypopgyium
outer dististyle dilated beyond midlength,
thence narrowed into a slender
rod,

outer

beyondmarginwithspines;aedeagus

simple,very

mm;antennalong,toextendingthe outer ends of the styli.

Male.

RostrumLengthaboutbrownish

5 mm;yellow,wing

4.8

palpi brown.

Antennae

aboutof male3.2

mm.

elongate,Vol.

setae,
longestof segments

bases of segments
Thoracic
coxae

and

about

two-thirds

reducedto smalllengthdelicateslightly darker. HeaddorsumHalteresyellow.

1969

1, January,

87

three-fourths
length of wing;
flagellar segments long,or moreslightly
long out
enlarged, provided withapproximately
proximal half
spreading
outer
ends

42, No.

almosttrochanters

fulvousknobs

yellow,remainder

yellow;at

vestiture

segment;

setae;

brown,

flagellum

brownish yellow.

uniformly

brownishofyellow,oflegs

clearer

pleuradarker.

slightlybrownish

with

Legs

outer

yellow,segments slightly darker. Wings brownish yellow, prearcular and costal
fields clearer yellow; veins pale brown, yellowed in the brightened fields.
on longitudinal veins beyond general level of origin of Rs
Macrotrichia
a further

with

on

series

outer

half

of

Sc

A. Venation:

2nd

end

Sc?

short,

ing just beyond origin of Rs, the latter very short, subequal to basal sec
tion of R5; branches of Rs diverging very gradually, cell R2 at marginabout two-thirds cell R?] cell 1stM2 closed but basal section of M3 very
faint;

m-cu

to

close

fork

of M.

Abdomen brownish yellow, subterminal segments brown to form abroad ring, hypopygiumwith tergal plateyellow. Male hypopygium
in outline, the outer lateral angles only slightly
small, subrectangular
produced. Dististyles
terminal; outer style dilated beyond midlength,with a few blackened spines on outer margin, the slender outer fifth ofstyle with similar, more appressed spines; inner style a slender gently
curvedrod,

tip

obtuse.

Aedeagus

and

long

extendedconspicuous,

caudadbeyond the level of the styli, appearing as a simple broadly
blade that is strongly sinuous before midlength.
Holotype male, El Valle, Panama,
2,000 feet, December(Gordon Field).
Other

similarsomewhat
Atarbaincludeelongateligiosa Alexander,
structureofthespeciesthehavingmexicana(Atarba)of Mexico,that differ

antennae

andhypopygium

aedeagus

A.

in

the

re

(A.)

fly infrom the present
and

1953

23,

and
and

Alexander

both

male

antennae

flattened

venation,

especially Sc, Rs, and the position of m-cu. Still other Neotropicalof Atarba have the aedeagus similarly lengthened but profoundly

species
bifur

cate.

Gonomyia
Generalyellow;colorationofrostrum yellow;

about
endingm-cu
atRs,
slenderrodlikethetened,
Male.

outer
LengthRostrum
broadlyandscutallobes

legs light brown; wings

faintlypraescutum

originoppositenear
one-fourth

terminal, apex of basistylenewstenorhabda,(Gonomyia)

of Rs,
jR2 + 3+4 subequal+ 4; male
M3
hypopygium

not produced;

gonapophyses;
more
two-thirds,
4-5 mm;about

aedeagus
widened.
wing

phallosome
nearly4-6

astospeciesbrown,

pleurainfuscated,

Sci

or

than

with

longerdististyles

including very long
long,

depressed-flatmm.

yellow, palpi brownish black. Antennae with scape brown,
remainder of organ dark brown; proximal
yellowed basally,

88

of

Journalsegments

flagellar

Kansas

with

three

remainderbrown,

ofbrown

stripes,notum

yellowed.

more

and

longerscutellum

brown,

confluentSociety

ones

outer

the

long-oval,

Head dark brownish gray.
Pronotal scutum yellowish
praescutum
scutallobes

Entomologicalslender.

Mesonotal

light yellow.
sides

narrowly
yellowed,Hal
Pleurayellowed.

teres with stem light brown, knob brownish black. Coxae and trochanters
faintly infuscated, stigma paleyellow, remainder light brown. Wings
brown,

poorlydelimited;thanlongerveinsbrown.

origin of Rs, Sc2 nearjust beyond

m-cu

Rs;

at near

Venation:Sc-lendingone-fourth

cell

M3+4;

2nd

A

or

opposite

its tip; #2 +3+4 long, subequal

to or

narrow.

with
Malebrown, hypopygium
hypopygium
yellowed.
dististyles
terminal, basistyle not produced; outer style long-oval, apex
longest setae nearly equal in length to style; inner style inpointed,cluding a slender blackened spine curving gently to the acute tip, rostral
includingportion shorter, with two long fasciculate setae. Phallosomethat are subequal in length to orvery long slender rodlike gonapophyses
slightly exceed the central aedeagus, each rod very slightly thickened
Abdomen

beforeoutwardly,

depressed-flattened,
membranous.

anarrowed
into
apex
outer
two-thirds

longspine.

Aedeaguselongate,nearlytruncate,

apex

broader,

23, 1953
2,000 feet, December
Holotype male, El Valle, Panama,
several males, December
13-22, 1953.
(Gordon Field).
Paratopotypes,
is
In its hypopygial
stenorhabda(Gonomyia)
characters, Gonomyia
most similar to G. (G.) mexicana Alexanderamong the described reincluding the very
gional species, differing especially in the phallosome,
slenderlonggonapophyses.

andpleurabrown,

third M3 +4; malecurvedwithphallosome
and

the

horns,

newstylacantha,

Gonomyia(Gonomyia)
to quaesita; mesonotal
Alliedandpraescutum

pleurotergite

yellowed;Sc

hypopygium

withdististyles

larger

inner

horn

gonapophyses

both

acute

two

with

relatively

short,

species
scutal lobes dark

m-cu

spinesshort, nearly

at

near

including
and

equal

one

simplelong

setae;in shape

size.
Male.

Length

about

3 mm;

wing

3.6 mm.

obscure yellow, palpi brownish black. Antennae with scapebasedarkened,yellowed, pedicel dark brown, flagellum paler brown.
Head yellow, vertex slightly darkened medially.
Pronotum and pretergites yellowed. Mesonotal
praescutum with disk
dark
sides
scutallobes
brown, posterior sclerites
brown,
broadly yellow;
and posterior border of scutellum yellow.
paler brown, pleurotergite
Rostrum

Pleurayellowed,

cleareronposteriorsclerites.

Halteres

with

stem

whit

trochanters
ened, knob infuscated. Legs with coxae chiefly yellowed,
remainder of legs light brown. Wings
clear yellow;
faintly darkened,veins pale brown.
prearcular and costal fields slightly more yellowed;
outer
with
of
three-fourthsoftrichia. Venation:
veins
wing
Longitudinal

Vol.

42, No.

1969

1, January,89

Sc short, Sc1 ending some distance before origin of Rs, the distancea little less than the latter vein, Sc2 slightly removed; #2+3+4
twice

nearly
outer

R3

Rs;

Abdomena simpledististyle

face with a setiferousation

curvedlargera

withmidlength,

short,
gentlycurved

horn,

near

at

hypopygium.

one-third

?3+4.

Male

hypopygium

withacute,

near

outer

tip

on

base

inner style bifid, the outer sclerotized por

tubercle;

two

horn

bearingthesetae,

few

m-cu

oblique,includingbrown,

only
long,

acute

outermostonspines
very

face

long;

at

rostral

and

below

prolongation of style relatively long, terminating in two unequal fasciculate setae.
Phallosome with gonapophyses
relatively short, nearly equal in size andshape; aedeagus with a small slender blackened
spine, the structure
beyond it pale and membranous.
22, 1953
2,000 feet, December
Holotype male, El Valle, Panama,(Gordon Field). Paratopotype male, pinned with type.
is related to various other Neo
Gonomyia(Gonomyia)
stylacantha
tropical species that center about G. (G.) quaesita Alexander,
includingalso G. (G.) anduzei Alexander, G. (G.) animula Alexander,
G. (G.)
birama Alexander,
and G. (G.) dominicana Alexander,
all being mostcharacters.
among themselves by hypopygial
readily distinguishednew species
uncinata,
Gonomyia(Paralipophleps)
Allied to pleuralis; basistyle of male hypopygium with a single long
slender spine, fleshy lobe relatively
short; dististyle with a strongly
curved black spine; phallosome with gonapophyses
long and slender;
ventral plate terminating in a strong curved blackened spine.
Male.
Length about 5mm; wing 3.2 mm.
Female. Length about 6 mm; wing 4.6 mm.
Coloration generally as in pleuralis and allies. Rostrum
and palpi
black. Antennae with scape and pedicel yellow, the former darkened
beneath; flagellum black. Head yellow, posterior vertex darkened.
Pronotumscutumandchestnutpretergitesbrown,

lightscutellum

yellow.
in

Mesonotal
type

and

praescutum

blackened;

postnotum

blackened, with a yellowed area at cephalic end of the interpostnotal
suture. Pleura brownish black, with a conspicuous whitened
longitudinal
stripe that includes the fore coxae, extended caudad to include base of
abdomen.

Halteres

with

stem

pale

brown,

knob

obscureyellow.

Legs

coxae and trochanters yellowed,
asthe fore pair more whitened,femora dark brown, tibiae and tarsi slightly paler brown.
described;
Wings weakly darkened; stigma dark brown, preceded and followed by
with

more

yellowed

Abdomenareas.

brown, restrictedly variegated with paler, posterior bordersof segments narrowly more darkened; hypopygium brownish yellow. Male
hypopygium with basistyle bearing a single long slender spine, fleshy
lobe relatively short, about three times as long as broad. Dististylewitha single strongly curved blackened spine at base of the subrectangular
beak, the latter with the usual pair of strong setae, at base of style with a

90

of

JournalKansaslarge flattened

yellow

plate.

Phallosomeasspines,

their

outer

long

slenderSociety

Entomological

ends

with

gonapophyses

appearing

ventralblackened;

plate

termi

nating in a single strongly curved black spine, in the unique slide mountthis structure appearing to be single, not paired as in pleuralis.
13, 1954 (Gordon
Holotype
male, Fort Gulick, Panama, January
Field).
In the strong blackened spine of the ventral plate of the phallosome,the present fly agrees most closely with Gonomyia(Paralipophleps)
in all other hypopygial
structures. As
pleuralisdiffering(Williston),indicated on the slide mount of this organ, the spine appears to be single
and not paired, as in pleuralis, but this condition may prove to be ap
ratherparentthan

theactualstructure.

new species
Gonomyia(Lipophleps)
minutistyla,
Mesonotaland scutal lobes brownish black, posteriorpraescutumsclerites of notum variegated brown and yellow; halteres light brown,
wings tinged with brown; male hypopygium with dististyle terminal, verysetae;
small, subequal in length and breadth, shorter than the modified
phallosome with two major elements, the longer one spearlike with theouter slightly more thickened.
Male.

about

Length

3.5 mm;wing

3.7 mm.

black, palpi black. An

Rostrumbrownishlight yellow, mouthpartstennae black throughout. Head dark gray.
Pronotumbrownishblack,

a darkenedoflateralMesonotal
yellow.
centralregionandpraescutum
scutumyellow;brownishbasal spot; mediotergite
border

and

the

Legs

with

coxae

yellow.

pleurotergite

ral region and a conspicuous
brown.

ofventralyellowed;

stripetrochanters

lobes

scutalscutellum

uniformly
withyellow,

black, anterior

two-thirds

Pleuradorsopleu

brown,

light yellow.
palebrown;

Halteresremainder

light
of

legs dark brown. Wings tinged with brown, without stigma; veins brown.
Venation: Scx ending just before origin of Rs; m-cu at fork of M.
brownish yellow. Male hypopy
Abdomen dark brown, hypopygium
gium with spinous armature of tergite abundant but relatively slender.
terminal, very small, its length and breadth subequal, shorter
Dististylesetae. Phallosome with two major elements,
than the terminal modified
the longer one spearlike, its outer half slightly thickened, tip acute; sec
ond element with basal half dilated, the outer half a slender spine; atbase of phallosome a small triangular plate terminating in a darkened
knob.

Holotype
L. H.

male,

Grand Etang,

Grenada,September

21, 1967(Noel

Krauss).

The most similar regional species include Gonomyia(Lipophleps)
haploa Alexander and G. (L.) maya Alexander, which differ especiallystructure. The dististyle of the present fly isin details of hypopygial
the smallest of any of the known members of the group.