NEW OR LITTLE-KNOWN TIPULID,ZE FROM EASTERNA S IA (DIP TE RA ), I X 'By CnEnr.ns P. AT,EXANDER
Of Amherst, Massachusetts
TWO PI,ATES

The crane flies discussed herein are chiefly from Szechwan
Province, western China, where two important lots of material
are included, one from Mount Omei, co,llectedby a Canadian
missionary and received through the kind interest of my friend
Mr. Herbert S. Parish, the second collected by Reverend David
C. Graham and now preserved in the United States National
Museum. An exceptionally interesting series of crane flies taken in the Japanese Alps, on and near Mount Norikura, weresent to me by Prof. Jiro Machida, to whom I express my deep
thanks for continued coiiperation in making known the rich tipulid fauna of the Japanese mountains. One further species
from Formosa was included in rich collections taken by myfriend, Prof. Syuti Issiki. Except where noted to the contrary,
the types of the specieshere discussedare preserved in my co,llection.
The following records of speciestaken in July, 1929, on andnear Norikuradake, Shinano, Japan, by Prof. Jiro Machida, areof interest.
Norilnn'cLdctke,July 26, 1929 (J. Mct,ch,id,a).
Tipula depressd sp. nov.
Tipulo. latzu.ensis Alex.
Tipula nippoalphta sp. nov.
Tipula politostriata sp. nov.
Ti.pulo. sachalinensis Alex,
Tipula uenoi Alex.
N epht'otomct sag hcr,IiensisAlex.
Cylindt"otoma japonica Alex,
PhalacroceramegacatLd,asp. nov.
(Lintonia)
Limontttm,achidai(Ale x . ) .

Di,cranoptgch,a cesia pallid,ibttsis subsp. nov.
Tr'icyphona pectinata sp. nov.
Tricyphona seticauda sp. nov.
Heterangaus laticincta sp. nov.
Limnophila (Tricholimnopltila)saitama Alex.
Limnophila mttndellcL sp. nov.
Mol.ophihts /alor sp. nov.

'Contribution from the Department of Entomology, Massachusetts Agricultural College.
oaoThe Phili,Wi,ne Jowtnl

840

of Sciencer0s1

Shi,rah,onaHot Spri,ngs, nea'r Norilatroda,ke, JuIy 25, 1929 (J,, Maoh'idal ,Gtuph,orWin trietio !Jex.
Tipu,la bubo Alex.
Hiro,gu, Norilcura'ilake, Julg 27, 7929 (J. Macldd'al.
Tipu.la ma,twntmimw Alex,
Tipu,la,politostri.ata sp. nov.
Nephrotoma eoghdieneie Llex.
Li,monia (Iri'mnnia,, subnubecu&oso (Alex.).

Heteran4eua I'atictwto' sl'. nov.
Limnnphila (Eplwlial il;iatziona
Alex.
Erinptero, (Ilisia, manhffia,i sp.
nov.

TIPTILIN,IE
DOLICEOPEZA(NESOPEZA) SUBGENICULATA .D. noY.

Closelyallied to D. (N.) geni,culq'ta(Alexander) but details ofma,lehypopygium quite distinct; outer dististyle slender, constricted and bent at nea,r midlength; lateral portions of ninth
sternite strongly produced eaudad.
Male.-Length, about 6.5 millimeters; wing, 8.
Fema,le.-I*ngth, about 8 millimeters; wing, 9.
Antennal flagellum yellow, the outer segxnentsdarkened.
Mesonotalprascutum brown with four darker brown stripesthat are little evident against the ground color; scutellum andpostnotal mediotergite more testaceous. Pleura chiefly 'darkbrown, the dorsopleural region and pteropleurite abruptly pale.
Halteres yellow, the knobs blackened. Legs with the fore co:rebrownish black, the remaining eoxe pale; troehanters yellow;
femoral tips broadly blackened,the tibial basesnarrowly whitened, the tips more broadly blackened; both femora and tibie
with a more whitish subterminal ring; tarsi white. Wings withthe pattern as in genfuulato,'ground color gray, the costal margin with four brown a;reas,the largest basal,the third along thecord to M, the fourth apical, interrupted by hyaline dropletsin outer endsof cellsRuand Ro; a more whitish border delimiting
the brown costal border behind; no conspicuousdarkening caudad of the main costal pattern excepting a small cloud at endof Cu,. Venation: Medial forks short.
Abdomendark brown, the tergites variegated with giray areas
on lateral margins of basal rings; outer sternites more blackened. Male hypopygium with the caudal margin of the ninth
tergite (Plate 2, frg. 21) tri'lobed, the median lobe shortest,
obtuse. Outer dististyle (Plate 2, fig. 22\ slender, strongly
constricted and strongly bent at just beyond midlength. Inner dististyle (Plate 2, fi9.23) broad, near base on outer face

4,t
i
I
I

I

Aleuand,er: Ti,pulid,s from Eostern Asin', IX

841

with several strong sete, in addition to the usual more delicate
ones. Lateral lobes of ninth sternite strongly developed,stouter
than in oberon.
Habi,tat.-China (Szechwan).
Holotype, male, Mount Omei, altitude 4,500 feet, July 81,
1929 (ex Parish). Alilotopotype,female, Augiust L4, 1929.
Very closely related to D. (N.) geni'culataof Japan, differing especially in the structure of the male hypopygium. The
production of the lobes of the ninth sternite is much as in D.
(N.) oberon (Alexander) of Luzon, but the other features of
the hypopygium are distinct.
TIPULA IXUSTA

rD. noY.

Belongs to the ci,nerei,frons group; thorax and abdomen entirely reddish orange; head gray; antenne and leglsblack; R112
atrophied; male hypopygium with only a single complicated dististyle.
Male.-Length, about 12 mil,limeters; wing, 11.
Frontal prolongation of head gray, the nasus short and stout;
palpi black. Antenne (male) of moderate length, if bent backward scarcely attaining the wing root; black, the second scapal
segment a little paler apically; flagellar segments only slightly
enlarg:ed basally. Head dark gray.
Thorax entirely reddish orange, without markings. Halterespale, the knobs dark brown. Legs with the coxe and trochanters ora,nge; legs black, the femora narrowly obscure yellow atbase. Wings (Plate l, fig. 1) gray, the stigrna brown; obliterative areas very restricted; veins black. Yenation ! Rr*, entirely
atrophied; cell 1st M, long-pentagonal; petiole of cell !{, about
one-half the cell; cell 2d A relatively wide.
Abdomen, including the hypopygium, reddish orange. Male
hypopygium (Plate 2, fig. 24) with the tergite (Plate 2, fls.
25) pale, the caudal margin with a broad, shallow, V-shaped
notch, the margins medially narrowly blackened, beneath further produced on either side of the median line into a smooth
flattened black plate that is directed eaudad and slightly ventrad,
the margin obtuse. Basisty,le produced caudad into a small
obtuse lobule. A single complex flattened dististyle (Plate 2,
fig. 26) shaped as in the figure. Notch of the ninth sternite
relatively shallow, with a smal,l more or less bifid fleshy lobe
jutting caudad, plaeed close to the suture of the basistyle.
Eighth sternite, 8s, unarmed.

342

The Phili,ppine Journal of Sciencergsr

Habitat"-China(Szechwan).
Holotype, male, Mupin, altitude 8,b00 feet, June 80, tg2;g (D.
C. Graham,). Type in the collection of the United States National Museum.
Tipula euusta is distinct from regional small species of theeinereifrons group in the structure of the male hypopyglum.
TIPULANIPPOALPINA

sp. nov.

General coloration gray:' nasus lacking; head with a medianbrown vitta; prrescutum with four brown stripes; legs red_dish, the tips of the femora and tibi,e narrowly b,lackened;wings whitish yellow, with a gray and brown clouded pattern;
Rr*, in direct alignment with the straight R,; abdomen reddish yellow, segments five to nine black; male hypopygium withthe tergite with a u-shaped median notch, and having a smaller
quadrate median incision at its base; eighth sternite unarmed.
Male.-Length, about 1b millimeters; wing, lb.b.
Frontal prolongation of head black, sparsely pruinose; nasus
lacking; palpi black. Antenne with the scapal segments andall but the extreme apex of the first flageilar segment reddish
yellow; remainder of antenne black; flag:ellar segments moderately incised, the longest verticils a trifle shorter than thesegments. Head gray, with a pale brown median vitta.
Mesonotal pnescutum grar, with four scarcely indicatedbrown stripes, the intermediate pair obsolete on anterior portions; scutum gray, the lobes very slighily darkened; scutellum
and postnotum gray with a nearly continuous median darker
vitta. Pleura gray, the dorsopleural region more buffy. Hal_
teres pale, the knobs infuscated. Legs with the coxre brownish
gray; trochanters reddish; femora and tibie reddish yellow,
the tips narrowly but conspicuously blackened; tarsi passing
into b'lack. Wings (Plate l, fig. 2) with the ground color
whitish yellow, the base and costal region clearer yellow; ahandsome clouded brown and gray pattern; the ground color
includes areas before and beyond the stigma; at beyond midIength of cell R; an oblique area beyond the cord, extending fromcosta into cell Ru and a second area extending from the outerend of cell R obliquely across cell 1st M, almost to the margin
in cell Mn; other pale areas in bases of cubital and anal cells.
Venation: Rs long;.lr+r- subobsoleteon outer half ; R, elongate;
Rr*, ir alignment with the straight Ru.
Abdomen reddish yeilow, the basa,rtergite more pruinose;
lateral and caudal portions of segments insensibly darkened:

t_ -

44,8

Al,exo,ttiler: Tipu,lifu frotn, Eastem' Asi'a, IX

843

posterior half of fifth, and all succeedingsegments,black. Male
hypopygium (Pilate 2, fig. 27) with the tergite, sternite, andbasistyleall distinct. Ninth tergtte (Plate 2, fi9.28) moderately chitinized, not heavily blackened,the caudal margin with a
U-shapednotch, at the baseof which is a smaller quadrate median incision. Outer dististyle a dusky subcy,lindricallobe, theapex obtuse. Inner dististyle .(Plate 2, fi$. 27, il') flattened,
compressed,the base with two small lobes, the apex bidentate.
.iEdeagus,a, jutting from g:enitalchamber. Ninth sternite, 9s,
with a V-shaped incision that extends to the base of the segment, the surface unarmed with lobesor tufts. Eighth sternite
unarmed.
Habi,tat.-Japan (Honshiu) .
Holotype; male, Norikuradake, JapaneseAlps, Shinano, July
26,1929 (J. Maehida).
Ti'puln nf,ppoalpfuwis distinct from all similar reg:ionalspecies in the lack of a nasus, details of venation, and structureof the male hypopyglum.
TIPULA DDPRESSA Brt, noY.

' General coloration light brown, the prrescutum with almost
concolorousstripes that are narrowly margined with darker
brown; wing:sbrown and gpay,variegatedwith extensivecreamcolored areas; male hypopygium with the ninth tergite tridentate; eighth sternite with a conspicuousmedian tonguelike lobe.
Mole.-Length, about 12 mi'llimeters; wing, 14. Deseribedfrom a somewhat teneral type specimen.
Frontal prolongation of head brownish testaceous. Antenne with the first scapalsegxnentdark brown, the tip and second
scapal segxnentmore reddish brown; flagellum apparently almost uniformly infuscated, the basal enlarg:ementsof the segments little if any darker. Head brown.
Mesonotalpre.scutum light brown, the four usual stripes almost concolorous,the intermediate stripes margined laterallywith a conspicuousdarker brown line, the mesal edge of thelateral stripes similarly darkened;posterior scleritesof mesonotum light brown, the seutellum more testaceous. Pleura lightbrown, the ventral pleurotergite darker. Halteres pale, theknobs broken. Legs with the coxa light brown, trochanters yellow; remainderof legsbrown, the femoralbasesmorebrightened.
Wings relatively broad, the ground color brown or gray, variegated by extensive cream-coloredareas, including a broadcrossbandbeyond the stigma and cord i other areas before stig-

gA

The PhiliWi,ne Jowrntr of Sciencerrrlma, at base and outer end of cell M, outer ends of radial cells,
as a broad seam along vein lst A in cells Cu and lst A andin the anal eells; veins brown, F4ler in the creamy areas. Venation: \*, entire but with macrotrichia confinedto basal twothirds; R, elongate,gently arcuated and thus not in alignment
with R2r.s;cell lst M, small, pentagonal.
Abdomen chiefly reddish yellow. Male hypopygium with thecaudal margin of the ninth tergite (Plate 2, fig. 29) conspicuouslytrifid, the lobesblackened;lateral lobes flattened, slightly incurved, the tips truncated, the outer anglesa little decurved;
median lobe narrowed outwardln the ventral surface with a
median keel that juts caudad slightly beyond the level of thelobe. Outer dististyle (Plate 2, fig.30) with tbe stemnarrowed,
the apex expandedinto a setiferous foot-shapedstructure. Inner dististyle (Plate 2, fig. 31) of unusual form, being deeply
bi,lobed,the anterior or cephalic portion being separated fromthe basal lobe by a conspicuousrounded notch. Eighth sternite(Plate 2, fig.32) with a conspicuous
median depressedlobe,thisbearing a small lateral lobule or shoulder on either side; themedian tonguelike lobe beyond the lateral lobules is providetl
with abundant sete that are directed mesad, these becoming
progressivelysmaller and more delicate away from the margin.
(Plate 2, fig.33) bifid, the outer arms appearing
Gonapophyses
as acute blackenedhooks.
-Habitat.-Japan (Honshiu).
Holotype, male, teneral, Norikuradake, JapaneseAlps, Shinano, July 26, L929 (J, Machi.d'a).
The strueture of the male hypopygium serv€sto separatethis
alpine speciesfrom similar forms.
TIPULA

POLfTOSTRIATA eP. nov.

Allied tn T, sh,omi,oAlexander; mesonotal prescutum andscutum gray, variegated with polished black areas; blackened
femoral tips narrow, occupying about the distal fourth of thesegment,the amount subequalon all legs; wings yellow, especially the baseand costal margin, the dark pattern restrieted; malehypopygium with the outer dististyle relatively small.
Male.-Length, about 13.5 to 14 millimeters; wing, 15.
FemnLe.-Length,about 17 to 18 millimeters; wing, 15 to 16.
Frontal prolongation of head black, sparsely pruinose, parler
laterally; nasus distinct; palpi black. Antenna (male) of moderate length, if bent baekward extending: about to the wingroot; scapeand basa,ltwo flagellar segmentsyellow; succeeding

44,rAl,enaniler: Tipttl,ita frsm' Eustern Asin, IX

945

littlesegxnentsalmost uniformly darkened or with the basesa
aa-rker;outerverticilsmuchlongerthantheventralpair.An.
tenne of female more uniformly pale' Head gray'
gray, thePronotum blackish gray. Mesonotal prescutum
threeusualstripespolishedblack,distinctlyseparatedbylinespol'oi trt" ground colori scutum gray, the centers of the lobes
ished biack; scutellumbrownish blaek; postnotum,blaek,sparseiy pruinose. Pleura gray. Halteres pale, the knobs -weakly
infuscated. Legs with the coxe black, pruinose; trochanters
the amount;;iilt*; iemora yellow, the tips broadlv blackened'
fourthdistaltheapproximately
subequalon all i"gB, o".opying
b'lack'tarsi
blackened;
tips
y"1l'o*,theof tft. segxnent;tibie
yelloqpale'
Winq
the proximal ends of basitarsi restrictedly
the base and costal region clearer yellow; a restricted brorrn
the stigma and a seam on the anterior eord;il1||;;inctuaing
nafrower brown seams on posterior cord and along vein cu;
a triangular dusky area in outer end of cell lst A; wing apex
* o ""o "le .ssdarkened,incaseslea v in g a p o o rly in d ic a t e d a re aoi tt. ground color distad of the stigma and cord' Venation:t'han
R,*o preserved; petiole of cell M, subequalto or shorters&rintl section of M,*o.
Abdominal tergit6J'.yellow,with three nearly entire blackish
whichstripes, the lateral stripes wider tlan the median one'
seg@ents;
theofmargins
caudaltheatis rnoreor lessinterrupted
subterminal segmentsblackened,this including the entire eig:hthoutersternite; sternites pale, with a median black stripe' thehypopypale'Malechieflysegrnentsblackened; hypopygium
giim with the median lobe of the tergite a nearly cireular comwith 1 short'[ressed blade; lateral margins of tergrte beneath
fig._34) rel(Plate2,
slender,blackenedpoint. outer dististyle
dististvle
Inner
ativety'small, the head only slightly dilated.
(Plate2'
fig' 36)
as figured (Plate 2, fig.35). Gonapophyses
appearing as bisPinous Plates.
Habi,tat.-JaPan (Honshiu).
Holotype, male, Norikuradake, JapaneseAlps, Shinano' July
26, Lgz';,'d. Machiila). Allotype, female, Hirayu, July 27' 1929(. m*niao1. Paraiopotvpes, 3 females; paratypes' 4 males
and females, with the allotYPe.
Tipula poli'tostriatu is most closely allied to T' shomio Alexandei rrrO f. strzr Alexander, differing especially in the morenearly unicolorous wings, without infuscation in the costal region, in the mqre narrowly darkened femoral tips, and in thedetails of structure of the male hypopygtum'

346
TIPULA

The Philippine Journnl of Science
ANGUSTILIGULA

real

sp. nov.

Belongs to the continuata group; antennal flagellum entirely
black, the segments with a single well-developed verticil; pr*scutum yellow with three brown stripes, the median stripe prolonged backward onto the median region of the scutum andscutellum; pleura entirely yellow; tips of femora blackened:male hypopygium with the ninth tergite unblackened, the caudal margin with a broad u-shaped emargination; ninth sternite
with a narrow median liguliform lobe.
Male.-Length, about 9.b millimeters; wing, 10.
Frontal prolongation of head relatively short, light yellow on
dorsal half, more infuscated laterally; nasus conspicuous; palpi
black. Antennre with the first scapal segment yellow, the second tinged with green ; flag'ellum entirely black; verticils offlagellar segments reduced to a single long basal bristle on outerface. Head yellow, the vertex with a linear more-darkened
atea.
Pronotum darkened medially, more yellowish laterally. Mesonotal prrescutum yellow with three brown stripes, the medianstripe narrowed behind and traversing both the scutum andscutellum; scutal lobes and parascutel,ladarkened; postnotal mediotergite brownish yellow, with a clearer yelrow median line.
P,leura yellow. Halteres dusky, the knobs yellowish. Legs
with the coxa and trochanters yellow; femora obscure yellow
basally, passing into dark brown; tibie black, the extreme basea little brightened; tarsi black. wings with the ground color
whitish, the base and prearcular region more yellowish; stigmaoval, dark brown; conspicuous brown washes and seams along
most of the veins, more extensive in the outer radial field andin cells R and M, the latter diversified by a single area of theground color at about the distal fifth; veins black, the basa,land
costal regions light brown or yellowish brown. venation: Rs
relatively short, genUy arcuatedi Rr*, entire; cell M, very deep,barely sessile to very short-petioldte; second section of Mr*orelatively extensive, subequal to or longer than r-m.
Abdomen yellow, becoming more brownish yellow on the outer
segments; subterminal segments more uniformly infuscated.
Male hypopygium (Plate 2, fig. 3?) with the tergite, 9t, extensive, not blackened, the caudal margin with a broadly
rounded U-shaped notch (Plate 2, fig.88) ; on ventral margin
at near midwidth of the lobes with a small blackened point.
Outer dististyle a long cy,lindrical rod, the base thicker, thesurface with long conspicuous sete. Inner dististyre with the

44,8Al,eranl,er: Ti,pulid,a from Eostern Asi,a, IX

U7

apex simple. Median notch of the ninth sternite at base witha slender pale lobe (Plate 2, fig.39), slightly broader at base,gradually narrowed to the blunt tip.
(Szechwan).
Habitat.-China
Holotype, male, Mount Omei, altitude 4,500 feet, August 10,
1929 (ex Parish).
Tipula angustiligulo is closest to ?. ranthoplew.a Edwards(Tibet to Sikkim) ; it differs in numerous details, as the entirely
blackened antennal flagellum and structure of the male hypopygium, especially the emarg:ination of the tergite and the elongate
liguliform ,lobeof the ninth sternite.
TIPULALATIFLAVA

sD. nov,

Belongs to the mntila group; general coloration gray, the prrescutum with four broken brown stripes, the interspaces with
abundant brown setigerous punctures; knobs of halteres infuscated; legs black, all femora with a very broad yellow subterminal ring; wings whitish subhyaline, with a brown patternthat appears as four interrupted crossbands.
Female.-Length, about 15 millimeters; wing, 13.5.
Frontal prolongation of head gray, more b,lackenedlaterally;
palpi black. Antenne with the scape and basal four flagellar
segments yellow, the remaining segments more bicolorous, thebase being darkened, the apex pale. Head gray, the centra,lregion of the vertex with a more or less distinct dusky line.
Pronotum gray, with three brown lines. Mesonotal pr€scutum gray with four broken brown stripes, the intermediate pairentire on front half, becoming broken behind, separated fromone another by a capillary brown median vitta; lateral stripes
nearly obsolete, best represented by delicate margins; interspaceswith abundant brown setigerous punctures; scutum gray,each lobe with two brown areas; scutellum gray, with a faintbrown median vitta; postnotal mediotergite gray with a darkbrown median line. Pleura yellowish gray, more or less variegated with darker, espeeial,lyon the ventral sternopleurite
and meron. Halteres yellow, the knobs infuscated. Legs withthe cox,regray; trochanters brownish yellow; femora black, witha very broad yellowish ring before the tips, this ring approximately three times the black tip; tibie and tarsi b,lack. Wings(Plate 1, fig. 3) relatively narrow, whitish subhyaline, with abrown pattern that is chiefly distributed as four broken crossbands; cell C dark brown; Sc more yellowish brown; the mostextensive of the crossbands is that at origin of Rs, becoming

S4S

The Phil'iVpi'ne Jau'rwl of Saietwe

ltst

wider and more diffus€ in the anal eells; apical band confinedto the radial field; narrow brown searnsat margin and alongveins Cu and 2d A; veins yellow, brown in the olouded&reag.
Venation! R,*n atrophied; Rs about one-third longer than $1s.
cell lst M, iiiiall, irregularly pentagonal, the shortest elernen{
beingm.
Abdominal tergites obscure brownish yellow, with a conspicuous dorcomedian black stripe that is narrortrly interrupted at
the caudal margin of each segment; sternites similar, the median stripe broad. Ovipositor with the valves yellowish horncolor, the basesof all valvesb,lackened;tergal valvesstraight andslender.
H abita,t.-China ( Szechwan).
Holotype, femde, Mount Omei, altitude 4,600 feet, August 7,
1929 (ex Parish).
Tipula tali,fl,aaais readily told from the two similar regional
species, T. stri,atipeflroisBrunetti and f. quqd,ri'fasci'a'toMatsumura (afuto Alexander), by the unusually broad yellow ringson all femora.
CYLINDROTOMIN.€CYLINDROTOXA NIGRIPES 3D, rqY.

General coloration black, the head and thorax more pruinose;
dorsopleuralregion yellow; halteres and legs black; wings witha faint dusky tinge, the small stigma dark brown.
Male?-Length, about 8 millimeters; wings, 9.5.
Mouth parts destroyed. Antenne short, if bent backward not
attaining the wing root; flagellar segmentsshort-cylindrical, theterminal segmentilongerthan the penultimate, pointed at apex;
flagellumpalebrown, the scapalsegrnentsdestroyed. Head gray.
Thorax blackened,sparsely gray pruinose, the three prescutal stripes and centers of the seutal lobes more blackened,theinterspaces gray pruinose. P,leura black, pruinose, the dorsopleural region eonspicuouslylight yellow; dorsal sternopleurite
and ventral pteropleurite a little paler than the remaining pleurites. Halteres blackened. Legs with the coxe blackish g:ray,
their tips paling to yellow; trochanters obscureyellow; remainder of 'legs black, only the femoral bases restrictedly obseureyellow. Wings (Plate 1, fig. 4) with a faint dusky tinge, thesmall oval stigma dark brown; veins dark brown. Venation:
Rs long, gently angulatedat origin; cell M, short-petiotrate;m-cu
at about two-fifths the lower face of the large cell lst M,; cell
2d A relatively narrow

&,8

Alerander:

Tipuli.d,a. from Eastern Asta, IX

349

AMomen, including the hypopygium, black.
( Szechwan).
Habitat.-China
Holotype, sex male?, Mupin, altitude 3,500 feet, June 30, 1929
(D. C. Grahom'). Type in the collection of the United States
National Museum.
The antenna are detached and the tip of the abdomen socrushed that only fragments remain but the sex eertainly seems
to be male. The black coloration and unusually short antenna
well suffice to distinguish the present species from other known
members of the genus.
PHALACROCERA IIECACAUDA

rD. nov.

General coloration pale yellow, the mesonotum and pleuravariegated with polished black; wings tinged with yellow, thestigma obsolete; basal section of Rr+s longer than m-cu, thelatter at near two-fifths the length of cell lst M,; male hypopygium unusually larg:e, the basistyles apically produced into black
earlike plates.
Male.-Length, about 11 millimeters; wing, 10.5.
Rostrum testaceous; palpi pale, the outer segments darkened.
Antenne pale, the outer segments passing into brown. Headbrownish black, the anterior vertex silvery.
Pronotum yellow, Mesonotal prrascutum yellow with threenearly confluent polished black stripes, the median stripe broadest in front; lateral stripes crossing the sutnre onto the scutal
lobes; remainder of mesonotum yellow, the posterior margin of
the postnotal mediotergite with two confluent black areas,
Pleura testaceous yellow, the sternopleurite extensively blackened; a smaller black area on the ventral pleurotergite. Halteres chiefly pale. Legs with the coxe and trochanters yellow;
femora obscure yellow, the tips weakly darkened; tibie brownish
yellow; tarsi passing into dark brown. Wings (Plate 1, fig. 5)
tinged with yellow, the stigma obsolete; veins brown. Venation:
Sc, not reaching costa; free tip of Sc, nearly perpendicular, thetip obsolete or nearly so; basal section d &*u longer than m-cu,
the latter gently sinuous, placed at near two-fifths the lengthof cell lst M,.
Abdominal tergites light brown, the outer segments moredarkened; hypopygium chiefly pale; sternites obscure yellow,
the subterminal segments more infuscated. Male hypopyglum
unusuaLly larg:e for a member of the genus, the basistyles produced into conspicuous black earlike plates that are conspicuously
fringed with yellow sete.
206956_?

350

Ihe Phi,li,ppi,neJowtnJ of Sci,einee

resr

Habi,tat.-trapan (Honshiu) .
Holotype, male, Norikuradakg JapaneseAlps, Shinano, July
26, I92g (J. Machid,a).
Plmlawoeera megacantd,ais generally similar ta p. ,ntkddn
Alexande&differing especiallyin the unusually large male hypopyglumLIMONIIN.E

LrroNra (Drc*ANolryr^rr"""JH,J'tiX;
re. noy.
Differs from Z. (D.) pttlchri,pennisinthe darkenedwing apex
and complete white subapical crossband; darkened areas of
costal field extending caudadto vein M.
Male,-T,ength, about 6.b millimeters; wing, Z.b.
Female.-LenErth, about g.b millimeters; wing, g.b.
Rostrum black, pruinose; palpi black. Antenne with thescapal segmentsblack, the flagellum yellow, only the outer seg_ments more infuscated; flagellar segmentsoval, becomingmore
elongate outwardly, the verticils short. Head black, yellowish
gray pruinose.
Pronotum black medially, more grayish pruinose laterally.
'Mesonotalprascutum with the ground color brownish gray withthree black stripes, the median one broad and complete, thelateral stripes remotefrom the suture; scutal lobesblack; medianregion of scutum and the scutellum more grayish; postnotum
biack, sparsely pruinose. Pleura heavily gray pruinose, striped
longitudinally with a narrow blackish area on the anepisternum.
Halteres yellow, the knobs black. Legs with the coxe and troehanters blaclq more or less pruinose; femora yellow, the tips
broadly black; tibie yellow, the basesand tips narrowly blackened; basitarsi brownish yellow, the outer segmentsblackened.
Wings (Plate 1, fig. 6) whitish, with a heavy brown patternof the general type of pulchri,pennis,. areas of costal margin
continuedback to vein M and only slightly narrowed posteriorly; wing apex uniformly darkened, not intermpted by a palestreak in cell Ro,as is the casein pulcltdpennis,.a completewhite
subterminal crossband. Venation: Cell 1st 1\{, open or tending
to be open by the atrophy or thinning out of the distal sectionof M; (figured as present, though faint).
Abdomen black, the hypopygium obscureyellow. Male hypopyCrum witJr the rostral spine shorter than in pul,chrtpenni,s;appressedto the margin of style.

,r,r,rAl,eaanl,er: Tipafid"p from Eastern Asi,a, IX

351

H abi,tat.-China ( Szechwan).
Holotype, male, Mount Omei altitude 4,500feet, July 31, 1929
(ex Parish). Allotopotype, female.
Closelyallied to pulohri,pennis(Brunetti), differing.esDeciallyin the wing pattern and details of structure of the male hypopygrum.
LIIIOMA(EUGLOCEINA)DIGN]IOSA!D. nov.

General coloration dark brown; tarsi entirely snowy white;wings with Rs long, angulated at origin; inner end of stigxna
not passing far basad of fork of Rs; Cu, present as a distinct
elementthat is equal in length to Sc or 2d A.
Male:-Length, about 7.5 to 8 millimeters; wing, 8 to g.
Ferwl,e.--Length, about 7 to 7.5 millimeters; wing, 7 to 8.
Rostrum and palpi brown. Antenne black, relatively long fora member of the subgenus;flagellar segmentslong-oval,with adusky glabrousapical neck; eac-hflagellar segxnentwith one very
long verticil, these unilaterally arranged. Head black.
testaceous,the knobs
Thorax dark brown. Halteres elongi'ate,
dark brown. Legs with the coxre and trochanters brownish
testaceous;femora and tibie black; tarsi entirely snowy white.
Wings (Plate 1, fig. 7) with a strong brownish tinge, somewhat
clearer basally; stigma oval, darker brown; veins brownish
black. Venation: Sc relatively long, the distance behreen the
tip oJ Sc, and origin of Rs not exceedingtwice Rs; Sc, variable
in length; Rs long, arcuated or angulatedat origin, remote fromthe inner end of the stigma; cell lst M, elosed;Cu, present as a
distinct elementto about oppositethe end of veins Sc, and 2d A
Abtlomen brownish black, the basal sternites paler.
Habi,tat.-China (Szechwan).
Holotypq male, Mount Omei, altitude 6,000feet, July 24, 1929
(ex Parish). Allotopotype, female. Paratopofpes, 1 male, 1female.
Li,monia, (Erylnch,i,rw) ili.gnitosa is distinct from all known
speciesof the subgenus. The length of Rs and its relation to
the proximal end of the stigma is approachedby L. (8.) olci'nn(Alexander), which differs in the darkened.proximal ends
usensi,sof the basiiarsi, the loss of vein Cu, and the shorter Sc.
ANTOCf,A(ANTOCEA)

ilINUTICOR.NIE

lrr. nov.

Generql coloration ocherous' the prescutum with four brpwn
stripes; antenna small; wings grayish white, the stigltnalacking;852

The Phi,li,Wi,neJournal of ScienceroElmale hypopygium with the outer dististyle blackened,glabrous,
here the apex narrowed to an obtuse point.
Male.-Length, about 4.b millimeters; wing, b.2.
Fema,Ie,-Length, about 8.5 millimeters; wing, b.6.
Rostrum brownish yellow; palpi brownish black. Antenne
small in both sexes,if bent backward ending some distancebefore the wing root; basal segmentspale, the outer segments
passing:into dark brown; pubescenceand verticils of segments
short. Head brownish gray.
Mesonotalprescutum ocherous,with four brown stripes, theapproximated intermediate pair darker and better delimited;scutum and postnotum gray, the scutellum more oeherous.
Pleura obscurebrownish ocherous,more or less variegated withdarker. Halteres dirty white. Legs with the coxe ocherous,
the fore co)r@a Iittle darker; trochanters obscure yellow; remainder of legs brownish yellow, the tarsi passing into browu.
Wings (Plate 1, fig. 8) grayish white, the costal region moreyellowish; no indication of a stigma; veins pale. Anal angleof wing prominent. Venation: Rs of moderate length only;
m-cu before fork of M.
Abdominal tergites brown, the sternites and hypopygium yellow; a conspicuousdark brown subterminal ring in male, occupying segmentsseven and eight. Male hypopyglum (Plate2, fi$. 40) with the caudal margin of tergite, 9f, very genfly
emarginate. Outer dististyle, od, blackened,glabrous,near apex
gradually narrowed to an obtuse point. fnner dististyle, id, a
Iittle longer, pale, with conspicuoussete. .iEdeagusslender,
subtendedby several pale bladesand points, as figured.
Habi,tat.-China (Szechwan).
Holotype, male, Mount Omei, altitude 4,500 feet, August 2,
1929 (ex Parish). Allotopotype,female,in eopulawith type.
Antoclw minuti,cornishas a hypopygium that suggestsA. iw
di,caBrunetti (Himalayas, western China, Malay States) and A.
satsuma Alexander (Japan). The species differs from ind,i,ca
in the small antenne of both sexes and from satwltnn in theimmaculatewings and details of the hypopyfium.
ANTOCIIA(A!{[OCEA)NEBULIPENMS

gD, nov.

Size large (wing, male, 8 millimeters); general coloration
yellowish gray; wings whitish subhyaline, handsomely cloudedwith gray; male hypopygium with the caudal margin of tergite
with a rounded lobe on either side of a small rounded notch;

{,tAleuand,er: Ti,pldiile from Ea^ster"nAsin, IX

853

basistyle with the outer apical angle produeed into an obtuse
lobe; outer dististyle blackenedand acute at tip.
Male.-Length, about 7 millimeters; wing:,8.
Rostrum and palpi destroyed. Antenne relatively short,
black; flagellar segmenh oval, with short verticils. Head yellowish gray.
Mesonotum yellowish {tayt.crushed in the unique type, thepreseutum apparently without stripes. Pleura yellowish gray.
Legs with the coxe brownish gray; legs relatively long, reddish
brown, the outer tarsal segmentsmore darkened. TV'ings(Plate
1, fig. 9) whitish subhyaline,with a cloudedgray pattern, including areasin eell R at near midlength of Rs, along cord and outerend of cell lst Mr; outer end of vein R, and as marginal seams
along the longitudinal veins beyond the cord; stigma elongate,
brownish grayi veins brown, darker in the infuscated areas.
Venation: Rs long; inner end of cell R, far basad of cell Ro;
cell lst M, closed;m-cu shortly before the fork of M.
Abdomen gray. Male hypopygium (Plate 2, fi9.41) with thetergitq 9t, transverse, the caudal margin with a small rounded
lobe on either side of a similarly formed median notch. Basistylq b, with the outer apical angle produced caudad into an
obtuselobe, the dististyles thus subterminal in position. Outerdististyle, o4 strongly bent before apex, the tip blackened,narrowed into an acute spine. Inner dististyle, 'td,, arcual'ed,theapex more dilated into a weak spatula that is denselyset withshort curved spinous seta. Eaeh gonapophysisappearing as aslender yellow rod.
Habi,tat.-China (Szechwan).
HoloWpe,male, Mupin, altitude 3,500feet, June 30, 1929 (D.
C. Grallmrt). Type in the collection of the United States National Museum.
Antocha nebulipenwisis very distinct from regional speciesin
the, large sizq wing pattern, and structure of the male hypopygrum. Antoclwnebulosa Edwards (Pahang) has a somewhat
similarly cloudedwing but is a very different fly.
DICRANOPTICSA

C.lsSIA PALLIDIBASIS

subcrt. nov.

Male.J.ength, about 7.5 millimeters; wing:, 8.6.
Female.-Length, about 8 millimeters; wing, 9.5.
Generally similar to typical D. casia' Alexander, differing asfollows:

854

The Phi,lippine Jmrnal of Science

resr

Scapeof antenne pale. Abdominal tergites dark brown, thebasal sternites paler. Male hypopygium with the outer dististyle gently curved,the outer marg:inwith conspicuousappressed
serrations, the apical spine smooth. Inner dististyle stout, angularly bent near midlength, the apex subtruncate. Lateralarms of tergite gently curved,the concavemargin microscopically
serrulate.
Habitat.-Japan (Honshiu) .
Holotype, male, Norikuradake, JapaneseAlps, Shinano, July
26, 1929 (J. Mach,i,ilal. Allotopotype, female.
PEDICIINIIBICYPIIONAPECTINATA

srr. Doy.

General coloration reddish brown, the prrescutumwith three
ill-defined blaekish stripes; wings faintly suffused with brown;
cell R* petiolate; medial field of wing beyond cord tending tobe pectinate by the atrophy of the basal section of Mr.
Male.-Length, about 5.5 to 6 millimeters; wing, 6.5 to 8.
Rostrum and palpi black. Antennre brown, l5-segmented;
basal flagellar segmentssmall and crowdednthe outer segments
becoming more attenuate. Head gray; anterior vertex wide,with a circular median depressionsurroundedby an elevatedrim.
Pronotum brownish gray. Mesonotum reddish brown, thepr"ascutum with three ill-defined blackish stripes, the lateral
stripes barely indicated; scutellumreddish brown. Pleura blackish gray. Halteres pale, the knobs dusky. Legs with the coxe
and trochanters obscureyellow; femora yellow, the tips passing
into brown; tibie and tarsi brown. Wings (Plate 1, fig. 10)
faintly suffusedwith brown, the stigrnal region diffusely darker
brown; veins pale brown. Venation: Cell R. petiolate, the petiole subequalto r-m; medial field beyond cord tending to be
pectinate by the atrophy of the basal section of Mu, in a few
caseswith cell lst M, closedor with a mere spur of Mr.
Abdomen black, the hypopygium paler. Male hypopyglum
about as in ?. insulana Alexander.
Habi,tat.-Japan (Honshiu) .
Holofpe, male, Norikuradake, JapaneseAlps, Shinano, July
26, L929 (J. Maclai,iln). Paratopotypes,3 males.
Tri,cgphonapectirnta is obviously allied to T. insulano Alexander, differing most evidently in the smaller sizg brownish bodycoloration, tinted wings, and the normaUy pectinate venationof the medial field of the wing.

u,sAleuand,er: Tipulida fram Eastern Asi'a, IX

TBICYPEONA

SEflPENNIS

355

rp. noY.

Allied to T, aetusfo Alexander; general coloration gray, thepraeseutumwith four brown stripes; apical cells of wing, fromSc, to 2d M", inclusive, with macrotrichia; male hypopygrum
with the tergite narrowed outwardly, setiferous; mesal lobe of
dististyle narrowed outwardly.
Male.-I-ength, about 9 millimeters; wing, 12.
Rostrum black, a little paler ventrally; palpi black. Antenne
black, the first scapal segmenta little paler. Head dark gray,the anterior vertex paler gray.
Pronotum light gray. Mesonotal prescutum gray with fourbrown stripes, the intermediate pair long, more confluent infront; scutum gray, the lobes more blackened; scutellum andpostnotum pale brown, sparsely pruinose. Pleura brown,
sparsely pruinose, the ventral sternopleurite darker. Halterespale, the knobs weakly infuscated. Legs with the cox pale,sparsely pruinose; trochanters yellow; femora yellow, passing
into brownish lolack;tibie brown, the tips darker; tarsi brownish
black. Wings (Plate 1, fig. 11) yellow, with a sparse brownpattern arranged about as in aetusta. Sparse macrotrichia inouter ends of eells Sc, to 2d,M", inclusive. Venation: Cell R,
petiolate, the petiole subperpendicular,in alignment with r-m.
Abdominal tergites light brown, paler laterally; outer segments
and hypopygium brownish black. Male hypopygium with thegeneral structure of T. seticaudoAlexander. Ninth tergite extensive,gradually narrowed outwardly, the surface conspic'uously
setiferous. Mesal lobe of dististyle narrowed outwardly, thethe mesal
blunt apex with a group of spinoussete. In seti,caud'a,
its marflatteneddisk,
circularlobe of the dististyle is a nearlygin evenly rounded.
H abi,tat.-Japan ( Honshiu).
Holotype, male, Norikuradake, JapaneseAlps, Shinano, July
26, 1929 (,1.Machidn).
Trtcuphona setipenni,sis most closely related to T. uehwta
Alexander, differing most evidently
Alexander and T. seti,cotuda
in the macrotrichia in the extreme outer ends of cells Sc, to
2d M", and the structure of the male hypopygrum, especiallyof
the tergite and mesal lobe of the dististyle.
EETERANGIEUS LATICINCTA

rP. noY.

Size large (wing over 10 millimeters) ; antennal flagellum
yellow; thoraeis pleura with the anepisternumand sternopleurite
darkened;legs yellow, the tips of the femora and tibia darkened;

356

The Philippine Journnt of Scienceresrwings whitish with a heavy, chiefly cross-banded pattern, including three crossbands; cell C uniformly darkenld; cell 2d
A wide, the margin crenulate; supernumerary crossveins as in
the genus, that in cell R, placed far distad, in cell R, about inalignment with R, in cell M, at two-thirds the length bf cell.
Male.-Length, about 9.b millimeters; wing, 10.b.
Female.-Length, about 10 millimeters; wing, 11.5.
Rostrum and palpi black. Antenne with the basal segment
black, the second segment and flagellum yellow. Head gray;vertical tubercle conspicuous.
Pronotum brown, paler laterally. Mesonotal pr:rescutumgrayish brown, the humeral region paler; scutellum and postnotum obscure yellow Pleura yellow, the sternopleurite and anepisternum darker. Halteres of moderate length, yellow, theknobs only weakly darkened. Legs with the coxa and trochanters pale yellow; femora yellow, the tips broadly and conspieuously dark brown; tibire yellowish white, the tips conspicuously
blackened, the amount a little less than the femoral tips; tarsi
yellow, the tips of segments one to three and all of the remaining segments black. Wings (Plate 1, fig. 12) whitish, with an
unusually heavy, chiefly cross-banded pattern, pale yellowish
brown, margined with darker brown; cells C and Se uniformly
dark brown, cell Sc, variegated with the ground color; thechief fascie are those at origin of Rs, including cells R and M;
the cord; and a more irregular fascia at the level of the outerend of cell Lst M, sending a ray to the apex in cell Ro and connected with the central fascia in cells Ru and Cu; fascia nearlyparallel-sided and unusually wide; a more or less confluent seriesof spots in cell Cu; a large confluent area in cells lst A and 2d
A, inclosing a circular pale area near the outer end of cell 2d
A; veins yellow, darker in the infuscated areas. Wings relatively broad, especially the anal cells; cell 2d A with the margin strongly crenulate to bilobed before the vein. Venation: Rs
square and spurred at origini &+s+a short, less than the perpendicular basal section of R2..s; supernumerary crossvein incell R, Iying far distad, that of cell R, nearly in alignment tojust beyond the level of R,, that of cell M, lnng beyond twothirds the length of the cell and in alignment with the crossveinin cell Rr; cell 2d A much wider than in H. ,japoni,cus.
Abdominal tergites brownish black, a litfle paler laterally;sternites brighter; male hypopygium dark. Male hypopygium
almost as in the other. speciesof the genus, the arm of the dististyle unispinous, the redeagus capitate.

,r+,E

Aleraniler: Tipuliite from Eastern Asin, IX

957

Habitat.-Japan ('Honshiu) .

Holotype, male, Hirayu, JapaneseAlps, Shinauo,July ZT,I:g1g(J. Machidn). Allotype, female, Norikuradake, JapaneseAlps,
July 26, L929 (1. Machi,iln).
Heterangaotslati,cincta is evidently most closely allied to H.
japoni,cus (Alexander) from which it differs in the large size,
darkenedareas on pleura, and details of venation and wing pattern. The allotype female has a weak adventitious crossveinin cell R, immediately distad of the angulation of &+s aud another in cell Ro just beyond the fork of M1*r.
HEXATOMINILIINOPEILAUUNDELLA

rp. nov.

General-colorationblack, the mesonotumsubnitidous; pleura
heavily pruinose; halteres yellow; wings brownish yellow, thebase and costal region brighter yellow; cell M, about twice its
petiole; m-cu beyondmidlength of cell 1st M,.
Female.-Lenggh, about 7.5 millimeters; wing, 6.b.
Rostrum and palpi black. Antenne with the scapalsegments
black, the flagellum brown; flagellar segmentsoval, gradually
decreasing in size outwardly. Head black, sparsely pruinose,
especiallyin front.
Pronotum black, prulnose. Mesonotumblaclc,subnitidous,flresurface only vaguely pruinose. Pleura heavily pruinose. Halteresyellow. Legs with the co:re and trochanters gray pruinose;
remainder of legsbroken. Wings (Plate 1, fig. lB) with a strong
brownish yellow tinge, the costal and prearcular regions
brighter yellow; stigrna oval, pale brown, a little deeper than
the ground color; veins brown, brighter in the flavous areas.
Venation: Sc, ending shortly before fork of Rs, Sc, near its tip;
R, a trifle longer than R1a2;cell M, about twice its petiole;m-cu beyond midlength of cell 1st M,
Abdomen with the tergites black, sparsely pruinose; sternites
brownish gray. ovipositor with the valves elongate,blackened.
Habi,tat.-Japan (Honshiu) .
Holotype,female, Norikuradake, JapaneseAlps, Shinano,July
26, lg29 (J. Machida).
Li,mnophi,tammd,elle is generally similar to the Nearctic Z.
wundn osten sacken. In the absenceof the male sex r hesitate
to refer the species to the subgenus Pri,onolnbis where it willprobably be found to belong. It is readily told from all otherJapanesespecie$of Pri,onolabds
with eell M, present by the small
size and unpatterned wings.

358
LIIINOPIIILAThe Philippi'ne Journnl' of Science

ut1

NESONEMORALIS rD. rov.

Belongs to the nemoral;i,sgroup; wings gray, in male not evidently widest opposite vein 1st A; male hypopygium with the
gonapophysesappearing as spatulate blades.
Male.-Length, about 5 millimeters; wing, 6 to 6.5.
Femate.-Lencth, about 5.5 to 6 millimeters; wing' 5.6 to 6.5.
Rostrum dark brownish gray; palpi black. Antenne with thebasal segmentblack, the flagellum brown. Head gray, the orbits broadly more whitish graY.
Mesonotumbrownish gray, the prescutum with an intermediate pair of darker brown stripes, the lateral stripes scarcely
evident. Pleura gray. Halteres relatively elongate, yellow, theknobs very slightly darkened. Legs with the coxa and trochanters yellow; femora yellow, the tips slightly darkened; remainder of legs yellow, the tarsi deepeningto brown; segmentsof
legs rather conspicuouslyhairy. Wings (Plate 1, fig. 14) ofmale not noticeablywidest oppositevein 1st A, as is the casein L.
subnemoralds,'brownish gray, the stigma a little darker; veins
pale brown. Venation: Se' ending about opposite the fork of
Re+a+r,Sc, a variable distance from its tip; R, faint; cell M'
small, as in the group.
Abdomen brown, the subterminal segmentsof male not suddenly blackened,as is the case in subnemorali,s, Male hypopysiwn (Plate 2, fig. 42) with the outer dististyle, od, conspicuously bifid at apex. Gonapophyses,9, appearing as flattened
spatulate blades, the tips obtusely rounded.
Habitat.-Formosa.
Holotype, male, Shinten, December3, 1928 (S. /-ssd/ri). A1lotopotype, female. Paratopotypes, 6 males and females.
is very similar to L.innequnlfs AlexLimnophilnnesarLqrnora,lis
ander (Formosa) but has the gonapophysesvery different in
structure.
EBIOCSRA CIESAREA rP. aov.

Head black; thorax entirely dark brown' opaque;halteresdark
brown; wings suffusedwith brown, deepestalong the outer costal margin; cell M, deep; cell lst M, small and nearly square;
abdominal segirnentsone and five to nine inclusive black, segments
two to four orange-Yellow.
Mate.-T'ength, about 25 millimeters; win& 28'5; antenna'
about 6.
..
Rostrum and palpi black. Antenna ?-segmented,relatively
short, ai lhoivn:by the'measuiemerts; flagpllar segmentsde-

A,E

Aleronder: Tipuliils from Euste,m,Asia, IX

g5g

creasing in length and diameter outwardly, densely set with semierect sete; scapalsegmentsblack; flagellum yellowish brown,
the outer segmentsdarker. Head dull black; vertical tubercle
entire.
Thorax entirely dull chocolate brown to dark brov'rn, theprascutum with four glabrous stripes thbt are slightly brighterin color. Halteres dark brown. Legs with the coxe and tro.
chanters dark brown; remainder of legs brownish black (onlya single, posterior, leg remains, tarsi broken). Wings (Plate
1, fig. 15) with a strong brown suffusion,the costal region darker brown, esperially on the distal half ; extensive paler discal
streaks before cord; anal cells pale, especially basally; veinsbrownish yellow. Venation: Sc, angulated and weakly spurredat extreme tip; Rs very long, about onethird loneier than R;
Rr*r*. subequal to basal section of Roi Rr*, about two-fifths
Rr*r; cell M, deep,nearly three times its petiole; cell lst M, small
and nearly square; m-cu about one-third longer than the distal
section of Cu,, some distance beyond the fork of M.
Abdomen with the basal segment brownish black, its caudal
margin narrowly yellow; segmentstwo to four bright orangeyellow; segmentfive black,the disk suffusedwith orang:e;remainder of abdomen, including the hypopygrum, velvety black.
Eabi.tat.-China (Szechwan).
Holotypg male, Kwanshieq altitude 1,800to 3,500feet, 1924
(D. C. Grolw,nu). Tlpe in the collection of the United States
National Museum.
By Edwards's key' the present speciesruns to couplet 41,
aereeing most closely with E'. ni,grtaedni"sde Meijerg a quitedistinct species. The fly appearsto be most nearly allied to thelater-describedE'. pralata Alexanfler, differing in the pattern ofthe body, wings, and legs.
NEoLnINoPHTLA Prcrunar^*.E:::P"ERINI

Generalcoloration black, pruinose; halteresyellow; legs black;wings pale cream color, with a heavy dark browrr pattern, including broad seamaalong cord and some of the longitudinalveins, and large rounded clouds at origin of Rs, fork of Rr*.
and fork of Mr*ri r-m gonnecting with Rs hfore forkj d,nearly twice its own length before the fork of Br*n.
14qls.;T-e\gtfr, @out 6.6 millimeters; wing, d.b'.
I Ann. & Mag. Nat. Eist IX 8 (1921) ?0-?8.

360

Th,e-Phi,tippi,neJournnl of Sci,encet*L

Rostrum and palpi black. Antenna black throughout; fusion
segment involving four segmentsand a partial fifth; outer flagellar segmentselongate-oval,the verticile shorter than the segments. Head dark gray.
Thoracic dorsum black, with a very sparse gray pollen.
Pleura more heavily pruinose. Halteres yellow. Legs withthe co:re pruinose; remainder of legs black. Wings (Plate 1, fig.
16) pale cream eolor,with a heavy dark brown pattern, arranged
as follows: Cell C; cell Sc exceptbaseand apex; stigma; broad
seamsalong cord and outer end of cell lst Mr; continuousbrown
seamsalong veins Ro, outer sections of veins M' M., Cu, and
2d A; large rounded clouds at origin of Rs, fork of \*n, crossing the adjoining cells to vein R, and fork of Mr*r; the following veins are not seamedwith dark, except as noted above:Rs, R' R*, M and its branches, with the exception of the outer
sections of M" and M*, and lst A; prea/cular region moreyellowish; veins pale yellow, darker in the infuscated areas.
Venation: Sc, ending about opposite r-m, Se2 at its tip; Rsstrongtly arcuated at origin; r-m connectingwith Rs before thefork of latter; R, only about onehalf \*n; cell R, relatively
small, vein R, being a little more than one.half R.; cell 1st M,
elongate; m-cu about two-thirds its own length beyond the forkof M.
Abdomen, including the hypopygrum, black. Male hypop5'gium with the basistylesbroken beyondbase.
Habi,tat. hina (Szechwan).
Holotype, male, ,Mupin, altitude 3,500 feet, June 30, 1929
(D. C. Graham). Type in the collection of the United States
National Museum.
The only allied speciesis N. fusci,nerai,sEdwards (Yunnan),which differs in the wing pattern and venation. In addition to
the seamed veins of fusci,nerui,s,the present species has large
rounded clouds at origin of Rs, and at forks of Rr*. and Mr*ri
r-m connectswith Rs before the fork and R, is ifan before thb
fork of Rr*n.
TEUCEOLABIS (TEUCEOLABIS) SCITAIIENTA

sD. !ov.

General coloration yellow and blue-black; three nearly confluent black stripes on prescutum, the lateral pair reaehing themargins of the sclerite; wings yellowish with a restricted dark
pattern; male hypopyglum very simple, the styli terminal in
position, the outer dististyle a pale ribbonlike blade.

4,tAlerander:

Tipulid,a from Eastern Asia, IX

861

Male.-Length, about 6 millimeters; wing, b.g to 6.
FemaLe.-Length, about 6.b millimeters; wing 6 to 6.9.
Rostrum brown, a little longer than the remainder of thehead; palpi black. Antenne with the scape yellowish brown;
flagellum light brown, the outer segments darker; flagellar segments short-oval, with a dense white pubescence. Head dull
black, heavily pruinosg especially in front.
Pronotum orange-yellow. Mesonotal prescutum yellow withthree extensive blue.black stripes that are confluent behind or
nearly so; lateral stripes reaching the lateral margin of prescutum, leaving a narrow yellow area behind at the suture;
humeral region more orange-yellow; median region of scutum
and the scutellum testaceous-yellow; scutal lobes blackened;
postnotum black. Pleura yellow, the anepisternum, dorsal pteropleurite and the pleurotergite blue-black, the sternopleurite more
reddish brown, the two areas separated by a narrorr/ yellow
longitudinal stripe that is densely white pubescent. Halteresblack, the outer half of knob orange.yellow. Legs with thecoxre and trochanters orange-yellow; fore femora with morethan the distal half black, the base yellow; middle femora yellow; posterior femora yellow, the tips narrowly brownish black;fore tibia and tarsi black; middle and hind tibie yellow, narrowly blackened at bases; tarsi obscure yellorv, the apices of segments one and two and all of outer segments blackened. Wings(Plate 1, fig. 17) broad, yellowish, the costal region brighter
yellow; stigma brown; a narrow brown seam on anterior cord;a very broad and diffuse slightly darker clouding occupies thecentral part of the region before the cord; veins yellow, darker
in the infuscated areas. Venation: Sc, ending at near midlengthof Rs, Sc, far from its tip; Rs nearly straight on basal half;$*r*n subequal to or shorter than Rr; vein 2d A curved strongly
into the anal margin of wing.
Abdomen black, the bases of the second and succeeding segments in male yellow, the black of the outer tergites sending
cephalad a median spur that becomes larger on outer segments
so as finally to isolate the yellow to lateral areas on the subterminal segments; hypopygium small, dark brown. Abdominal
tergites of female uniformly blackened, only the genital segment
orange, the sternites with more yellow. Male hypopygium(Plate 2, fig. 43) unusually small and simple. Basistyle, b,
short and broad. Outer dististyle, od, a pale ribbonlike blade.
Inner dististyle, id., a flattcned blade that terminates in a black-

962

The Phi'kppine Jomttal' of Sci'ence

retr

ened beak, the outer angle with three long setr. Phallosome inslide mounts large.
H abitat.-China ( Szechwan).
Holotype, male, Mount Omei, altitude 4,500 feet' August 12'
rg2g (ei Parish). AllotoPotvpq femalq Augrrst 18' L929'
Paratopotype, female, August 22, L929.
Tewhol,abi,ssci,tamentais distinct from all regional species
in the featursg above listed. The structure of the male hypopygium is unusually simplg more so than in any reg:ional species
known to me.
EBK)PTERA(ERIOPTEBA)BREVIRAUA!D. nov.

General coloration reddish brown; head black, the orbits narrowly marked with silver; pleura iridescent blue; knobs ofhalteres darkened; wings brownish yellow; male hypoPygtum
with the tergrte emarginate; inner dististyle with two divergent arms.
Male.-Length, about 4.6 to 6 millimeters; wing, 4'5 to 5'
Female.-Length, about 5.5 to 6 millimeters; wing, 6'5 to
6.8.
Rostrum and basal segf,nentsof palpi yellow, the outer segments darker. Antenne brownish black, the basal segmentsa very little paler. Head black, the orbits with a narrow silvery
line.
Mesonotum reddish brown, the median region darker' thehumeri brighter; scutellum dark brown, narrowly bordered by
pale; postnotum dark gray' Pleura almost entirely covered byin iridescent blue pruinosity. Halteres palg the knobs infuscated. Legs relatively long and stout; co:re and trochbnters
yellow; remainder of legs yellow, the terminal tarsal segments
infuscated. Tyings (plate 1, fig. 1g) with a strong brownish
yellow tinge, the costal region clearer yellow; veins light brown,
more yeltowish in the costal region. venation: Distal sectionof Cg, and fi A strongly sinuoug ol distal portion'- Abiominal segtnents ehiefly dark brown, the caudal margins
somewhat brightened; hypopygrum obscure yellow' Male hypopyglum (pdte 2, fig.44) witb the tergrte shallowly €rnafproducedas in 'o' (E')
ii"ii", the median regxonnot convexly
Dististyles almost as ln ornati,froms, entirely glaE r*teiro*,brous, the branches of the inner dististyle, 'dd,shorter and more
egual,t.$eouter arm not subtendedon its basal third.by,a""""rypate ilanle bui with a eonspicuousflattened structure near thefork of the style.

4,E

Aleaaniler: Tipldida from Eastern Asto,, IX

g6g

H abi,tat.-China ( Szechwan) .
Ilolotype, male, Mount Omei, altitude 4,500feet, July 81, 19Zg
(ex Parish). Allotopotype, female, August 11, 1929. paratopoffpes, 4 males and females, August 10 to 20, lg2g.
Most clo,selyallied to E. (8.) ornati,frons Edwards (Ceylon), differing most evidently in the structure of the malehypopygium.
EBIOPTER-{ (ILISH)}IACEIDAI

sD. !ov.

General coloration brown; antennal flagellum brownish yellour; wings yellowish gray, with a very sparse pale brownpattern, including vague clouds at Sc' R, tip of R,.o and along
cord; Sc, longer than Rs; eell 1st M, moderately el,oigate; analveins gradually diverging.
Female.-Length, about 4.b millimeters; wing, 4.6.
Rostrum and palpi brown. Antenne with the basal segments
dark brown, the flagellum paling to yellow or brownish yellow.
Head grayish brown.
Pronotum brown, the anterior lateral pretergites light yellow.
Mesonotumbrown, the prascutum a little paler laterally. pleura
light brown, the dorsopleuralregion pale yellow. Halteres paleyellow, the knobs with golden-yellowsete. Legs with the coxe
and trochanters yellow; femora brownish yellow, the tips a litile
darkened; tibie and tarsi obscure yellow, the latter passing
into brown. Wings (Plate l, fig. 19) yellowish gray, the baseand costal region a trifle more yellowish; a very sparse andvague pale brown pattern, including clouds at Sc, R, and tipof Sc,, and tip of \*r, the latter two areas delimiting the ends
of a yellowish stigmal area; cord vaguely seamedwith dusky;
veins very pale brown, the macrotrichia darker. Venation:
Sc, elongate, exceeding Rs; Sc, about opposite one.sixth thelength of Rs; cell lst M, moderately elongate; m-cu before thefork of M; vein 2d A nearly straight the anal veins gradually
diverging.
Abdomen dark brown. Ovipositor with the valves very longand slender, pale yellow.
Habitat.-J apan (Honshiu) .
Holotype, female, Hirayu, .IapaneseAlps, Shinano, July 2?,
L929 (J. Machida).
I take great pleasurein naming this very distinct speciesin
honor of my friend Dr. Jiro Machida, to whom I am indebted
for many JapaneSeTipulidce. Eri,optera ,mnchiil.ai,is very different from the.remaining species of the subgenus ltisia de-

364

The PhiliWine Journal of Science

LsEr

scribed from eastern Asia. It is more closely allied to E. muculata and allies than to the speciescentering about E. oteolata
Siebke.
OBDIOSIA GRAEAIII

sD. noy.

Size very large (wing, female, 10 millimeters) ; general coloration gray, the prescutum with three confluent black stripes;
halteres light yellow; legs black; wings white, with a heavy darkbrown pattern that appears chiefly as seams along the veins;
veins R, and Rn deflected strongly cephalad at outer ends; aweak spur at near two-thirds the length of vein Ro, directed intocell Rn; cell lst M, closed; vein 2d A very strongly sinuous.
Femnle.-Length, about 8.5 millimeters; wing, 10.
Rostrum and palpi black. Antenne black throughout; flagellar segments oval, with relatively short verticils, these becoming smaller and less conspicuous on the outer segxnents. Headblackish, the orbits gray.
Mesonotal prascutum with the disk chiefly covered by three
confluent black stripes, the ground color restricted to the humeraland lateral portions; pseudosutural fovee black; scutum gray,the lobes chiefly blackened; posterior sclerites of mesonotum
black, sparsely pruinose. Pleura black, pruinose. Halteres entirely light yellow. Legs with the coxe black, pruinose; remainder of legs black. Wings (Plate 1, fig. 20) white, with aheavy dark brown pattern; prearcular region and an oval area
in cells C and Sc before the stigma light yellow; the brown areas
include most of cells C and Sc and very conspicuous seams along
most of the longitudinal veins, in cell M, so broad as to restrict
the ground color to a long-oval atea; a cross-fascia in cell 1st A
at about two-thirds the leneth; veins brown, more flavous in
the yellow areas. Macrotrichia of cells corresponding in color
to the areas occupied, being white, brown, and golden. Venation: R, just before fork of R. and Rn, the two latter veins deflected strongly cephalad at their outer ends; vein R, strongly
looped on distal half ; Rn with a spur at two-thirds the length,
directed into cell R*; cell lst M, closed; vein 2d A very strong:lysinuous.
Abdomen grayish black, with sparse yellow seta. Ovipositor
with the valves horn yellow, the tergal valves blackened basally.
(Szeehwan) .
Habi,tnt.-China
Holotype, femalg Mupin, altitude 3,500 feet, June 30' 1929
(D. C. Graham). Type in the collection of the United States
Nationa,l Museum.

+4,sAlerand,er: Tipukila lram Eastern Asia, IX

365

Ormosia grahami is named in honor of the Reverend David
C. Graham, who has added materially to our knowledgeof thefauna and flora of Szechwanand eastern Tibet. It is a strikingly distinct fly that requires no comparison with any known
patternedform
species,being the largest and most conspicuously
so far discovered.
IIOLOFUILUS

FEROX

sp. noy.

Belongs to the gracilis group; general coloration light brown;head gray; male hypopygrum with unusually complicated armature, including a spinous rod arising from the outer portionof the basistylg together with three dististyles, one of which
is bifid.
Male.-Length, about 4.6 millimeters; wing, 5.
Rostrum and palpi black. Antenne broken. Head gray.
Pronotum blackish. Mesonotum light brown, the pleura somewhat darker, the ventral sternopleurite and meron more testaceous brown. Halteres chiefly pale, the base of the stem
restrictedly yellow, the stem dusky, the large knobs yellowish
brown. Legs with the coxe and trochanters yellowish testaceous; remainder of legs pale brown, the tarsi passing intobrownish black. Wings grayish, the veins darker; macrotrichia
long, dark brown. Venation: R" lying distad of r-m; m-eu alittle less than one-half the petiole of cell M,; vein 2d A ofmoderate length.
Abdomen brown, the sternites somewhat paler, the conspicuous
genitalic styli black. Male hypopygium (Plate 1, fig. 45) withthe outer lobe of the basistyle, b, produced into a long sinuous
rod that is subequal o longer than the dististyle, on outer margln at near one-third the length with an aeute yellow spine;distal two-thirds of the rod narrower, the surface more roughened; ventral surface of basistyle with a slender flngerlike lobe.
Three dististyles, d,, t}:re longest a curved sinuous smooth rod,the distal portion bent at about a right angle into a long straightspine. Second dististyle conspicuously bifid. Third dististylemore basal in position, inserted in the notch of the basistyle,a simple blackened rod, the basal half a little thicker, the gently
curved distal half narrower. ,?Edeagus long, narrow, yellow.
H abi,tat.-Japan (Honshiu) .
Holotype, malg Norikuradake, Japanese Alps, Shinano, July
26, L929 (J. Mach.ilIa).
Molophi.Ius f eroo is very distinct from all other reg:ional speciesof the genus in the structure of the male hypopygrum.
266956_6

ILLUSTRATIONS
t:*tjil,?,:TT"*"Li?'&Ti;
rrscnd: d' Ddea'8u8;

ff' rnn.rdietrstvre;
od'

PI,ATE 1
Frc. 1. Tipula eousta sp. nov., venation.
2. Tipula nippoalpina sp. nov., venation.
3. Tiwla latiflnou Bp. nova venation.
4. Culhdrotoma ni.gdpes sp. nov., venation.
6. Phalac,roceromngacau.dagp. nov., venation.
6, Lilmoiia (Di.crannmgia) subpu.Ichripennis ap. nov., yenation.
7. Lirnonia (Euglnchhwl ilignitoaa sp. nov., veuation.
8. Antoclw (Antochal mi,nuticonaissp. nov., venation.
9. Antocho (Antocho) nebulipenni.ssp. nov., venation.
t0. Trirllphonm pectinata sp. nov., venation.
lL. Tricgphono setipennis Bp. uov., venatio{r.
t2. Heterangeue Loti,cinctorsp. nov., venation.
L8. Lbnnophila, rw.rd,ella, sp. nov., venation.
14. Iti,mnnplvilo nnsonemaralie sp. nov., venation.
16. Eri,ocero c&Borea,sp. nov., venation.
L6. Neolimnoplvila picturoto sp. Dov., venation
L7. Taucholabis (Taucholabisl gcito,menta sp. nov., venation.
L8. Eriopteru (Erbpteralbreoiramn sp. nov., venation.
t9. Erioptera (Ilisiul mo,chiilai sp. nov., venation.
20. Ormaeia grahmni, sp. nova venation.
Prars 2
FIc.21. Doliahopeza (Neaopezal subgenioulato sp. nov., male hypopygium,
nintb tere'ite.
22. Doliahopeza (Nesopezal wbgeni,adato sp. nov., male bypopygium,outer dististyle.
23. Dol,ichopezo(Nesopeza) wbgeniaild'to sp. nov., rtrale hy?opygium,
inner dististyle.
24. Tipu,la eausto sp. nov., male hypopygium, lateral aspect
26. Tipul,o erusta sp. nov., male hypopygirrn, ninth tergite.
26. Tiryln, eonsta sp. nov., male hypopygium, dististyle.
27. Tipu,lo nipoolpbw sp. nov., male hypopygium, lateral aspect.
28. TiNJa nippoolpi,no sp. nov., male hypopygium, ninth tcrg:ite.
29. Tipu,la d,epressosp. nov., male hypopygium, ninth tergite.
,80. Tipulo d,agresaasp. nov., male hypopygium, outer'dististyle.
81. Tipulo d,epreasosp. nov., male hypopygium, inner dististyle.
82. Ti,pu,lod,eyreeeosp. nov., male hypopygiurn, lobe of eighth stemite.
88. ?ipu,ln ilepreeso sp. nov., uale hypopygium, gonapophysis.
36?

368

The Phtl,iWine Jou'rqal of Scienceouter-- dististfle'
I'rc. 34. Ti'pttto, politoetri.ata sp. nov.' male hypopygium,
politostriata .p. ,orr., male hypopygium, inner dististyle.
ss. rhulagonapophysis'
36. Tipulo potitostriata, sp. nov', male hypopygium,
g. fipul'u angustiligula
sp. nov., male hypopygium, lateral aspect'
sp. non., urale hypopygium, ninth tergite'
38, Tiputa ongustiligula
lobe of ninth
sp. nov" male hypopygium'
39. Tipu,ta angusti'Iigula
sternite.
40. Antocha (Antoahal mht'uticornis sp' nov" rnale hypopygium'
41. Antocho (Antocha) nebul:i,pennis sp. nov" male hypopygium'
42. Limnophila nescnemoralis sp. nov', male hypopygium'
43. Teucholabis (Teucholabisl scitamentu sp' nov', male hypopygiunu
44. Eri,optera (Eri.optera) breoirama, sp. nov" male hypopygium'
46. Molophilus feron sp. nov'' male hypopygium'

ALEXANDER:

TIPULIDA

FR0M EAsrmN

Asrl,

IX.l

PL AT E 1,l PH r Ll P. J ouR N . Sc r .. 4{, N o. 3.

ALEXANDER:Trpulma

FRoM EASTERN AsrA, IX.J

[pHrLF. JouRN. Scr.,44, No. B.