'

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67 t.'.

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NUW ON INSUFFICIEITTLY-KIYOWNCBANE.FLIES FBOM CEII.S',GATEILYTrplrunaE, oBDnn ormEBA)(Apartado de Agricultura T6cnica y Lito.

Inp.

"STANLEY'

Junio de 1951-

Afio XI i

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Bv
GEABLES P. AI,EXANDEB'

Univcroity of Marirchurettc, Amhcrrt, Macr.

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ilIrISTEBIOITDE AGBICULTT'BA

DINbCOION ,GENEBAL DE AGBICULTUBA
SAIfIIACO-.'CEILE../

NEW OR INSUFFICIENTLY.KNOWN CRANE-FLIES FROMCHILE(FAMILY TIPULIDAE, ORDER DIPTERA)Part III (*)
By
CHARLES P. ALEXANDER(**)

Since the publication of the two preceding papers under
thisgeneral title, our knowledgeof the distribution of
tfre Tipulid"" l" Chile
has advancedmaterially. Important seriesof specimenshave
been taken
particularly by Messrs.Rairl Cort6s P., Guilleimo
Kuschel and Luis E.
Pefra G', to whom I am
greatlyindebted for the opportunity ofYelycontinuing the study of Chilean
crane-flies. A further """y importantcollectionof theseflies was taken by the California Academ;of bcience
Expedition, 1950-1951,
under the leadership of Dr. Edwards s. Ross,
All of these materials will be stud.ied "rrd the results publishedinsucceedingreports. In the present paper I am describing:a series of
speciesbelonging to the Eriopterine genus Motophitus -Curtis,
richlydeveloped in the South Chilean fauna. The types Lt the novelties
arepreserved in my collection, with paratypes and duplicatesof certain
speciesto be returned for the National Collectionof Chile.
In continuing this study it seemsadvisableto record thespecies
describedor included in the preceding parts and to give these
consecutive numbers, which may be used.when referring t; such species
inthe future. This system of consecutivenumbering;ri1 be followedinthe present paper and in subsequentreports under this same title.
(t) Part I of this seriesof papercwaspublishedin Bol. san. veg. cbile 3 (2): ll7-ltg.
ll figs.,1944.Part II in Agric.T6cn.cbile 5 (l): 5-23, 12 fig-s.,
lgqs.
(**) Universityof Massachusetts,
Amherst,Mass.

Agr. Tic. Chile.-

6

Afio Xl NP 1

TIPULINAE

1 Macromastir (Macromastit) atrirostris Alexander; II: 5-6'
2 M. M. nahuelbutaeAlexander; II: 7-8.
3 M. (Araucomyr,a) butlockr'Alexander; I: 118'
4 M. A. pent'taAlexander; I: 118-119.
5 Holorusia problematicaAlexander; II: 8-10.
6 Tiputa (Eimicrotipula) angotensis,Alexander;I: 119-121'
7 T . E. antarctica Alexander; I : LzL.
8 T. E. anthongmPhaAlexander; I: LZl.
I:119.
I T. E. aroucanioAlexander;
1 0 T. E. bigotiana Alexander; II: 10.
1 1 T. E. brethesianaAlexander; I: LZL.
t 2 T. E. breuicoma Alexander; I : LZL-L2?'
1 3 T. E. chitlanico Alexander; II: 10-1-2.
L4 T. E. dr'morphaAlexander; I: L22.
' 1 5
T. E. duseni Alexander; I : L22.
1 6 T. E. lonigera tau Alexander;I: L22-L24'
L 7 T . E. Iatif otia Alexander; I : t24'L26.
1 B T . E. microspilota Alexander; I : L26.
1 9 T. E. pattiifiiignata salutatorio Alexander; II: L2-L3.
20 T. E. paruilobo Alexander; I: L26.
2 L T. E. pirionr,Alexander; I: L26.
22 T. E. spatutifera Alexander; I : L26'
23 T. E. subandinoPhiliPPi; I: L26.
24 T. E. subligulataAlexander; I: .126;II: 13'
25 T. E. tersoidesAlexander; II : 13-15'
26 T. E. triemargr'nataAlexander; I : L27'
CYLINDROTOMINAE

Stibadocerina chilensis Alexander; II: 15'
LIMONIINAE
LIMONIINI'28
Limona (Dicranomyia) butlockiana Alexander; I: t2l '29 L. D. infumata (PhiliPPi); II: 15'
30 L. D. pinodesAlexander; II: 15'
3 1 Helius (Hetius\ araucariaeAlexander; II: 15-17'

CRANE-FLIES FROM CHILE. -ChorlesP. Alexander

7

PEDICIINI

32 Pedicia (Tricgphona) agsenensisAlexander; I: t27-t29.
33 P. T. chilota Alexander;I: L29.
34 P. T. serrimargo Alexander; I : 129-130( Patagonia) .
HEXATOMINI

35
36
37
38
39
40
4L
42
43
44

Austrolimnophila (Limnophilella) patagonicaAlexander; II: L7.
A. (Austrolimnophila) eelestissimaAlexander; II: 1?-19.
A. A. hazelaeAlexander;I: 130.
A. A. merklet,Alexander; II: 19.
A.' A. microsticta Alexander; I: 130.
A. A. aaritarsis Alexander; II: 20.
A. A. nanthopteracayutuensisAlexander; I: 130-131.
Limnophila armigera Alexander; Il: 20-2L.
Ggnoplistia (Ggnoplistia) bicolor ( Philippi) ; il : 21.
G. G. aariicalcarataAlexander; II: 2L.
ERIOPTERINI

45
46
47
48
49

Sigmatornera(Austrolimnobia\ bullocki ( Alexander) ; I : 131.
Aphrophila aurantiocoAlexander;I: 131-132
A. coronatoAlexander; I: L32-L34.
A. airidinerais Alexander; I: 134.
Alexander; I: 134-135.
Molophilus (Molophilus) dr,ceros

50 Molophilus (Molophilus) monostgloides,sp. n.
Belongs to the monostglus group; general coloration grayish
brown; antennaeshort; male hypopygium with the basistyle termindting in a stout oval lobe that is provided with about eight long stout setae;
a single unequally trilobed dististyle, the inner lobe placed far basadon lower margin.
Male. Length about 5 - 5.2 mm.; wing 5.5 - 5.8 mm.; antenna
about L.4 - 1.5 mm.
Rostrum and palpi brown. Antennae brownish black, relatively
short, as shown by the measurements;flagellar segmentslong-suboval,
the longest verticils approximately one-half longer than the segments,
which are provided with a short densewhite pubescence.Head grayish
brown.

Asr. Tic. Cnite.-

Afio XI No I

Pronotum brownish gray; pretergites obscure yellow. Mesonotum
grayish brown, the praescutum with two poorly indicated intermediate
brown stripes; pseudosutural foveae elongate, blackened; posteriorsclerites more pruinose. Pleura grayish pruinose, with long yellow setae.
Halteres obscure yellow, the knobs clearer yellow. Legs with the coxae
brown; trochanters brownish yellow; femora brown, stightly morebrightened basally; tibiae brownish yellow, the tips narrowly darkened,
the posterior tibiae clearer yellow; tarsi brownish black. Wings with adusky tinge, the prearcular and costal fields more yellowed; veins brown,
the macrotrichia still darker. Venation: R2 lying slightly distad of levelof r-m; petiole of cell Ma relatively short, about one-third longer than,n-cu; vein 2nd A long but only slightly sinuous.
Abdomen dark brown, the hypopygium slightly more yellowish
brown. Male hypopygium ( Fig. 1 ) with the basistyle, b, terminating ina stout oval lobe that is provided with about eight long stout setae;
surface of lobe microscopically scabrous. A single dististyle, d, that is
unequally trilobed; longest lobe a narrow central blade, its tip obtuse;outer lobe at near two-thirds the central blade on outer margin, appearing as a strong blackened spine; inner lobe a strong black spine, placed
far basad on lower margin, its base and axil with delicate setae.
Aedeagus elongate, its basal part consisting of two narrow blackened
rods.
Holotype, male, Curaeautin, Malleco, altitude 400 meters, December 18, 1950 (Pefra). Paratopotypes,6 3 .
The present fly is readily distinguished from the only otherdescribed member of the monostglus group, Molophilus (Molophilus)
monostglus Alexander, by the structure of the male hypopygium. The
hypopygium of this latter specieshas been figured in another report by
the writer (Rev. de Entomologia,18: 339; 1947).

51

Molophilus (Molophilus) pefi'ai, sp. n.

Belongs to the gracilis group, ruficollis subgroup; general coloration
fulvous yellow; halteres and wings pale yellow; male hypopygium withthe outer dististyle having both arms shorts, obliquely truncated attips; basal dististyte a long simple spine; phallosomic plate glabrous,
gently widened outwardly, the apex subtruncate.
MaIe. Length about 4.2 mm.; wing 4.8 mm.; antenna about L.2 mm.
Rostrum yellow; palpi brownish black. Antennae short; scapeyellow, flagellum dark brown; flageltar segments subcylindrical totruncated long-oval, the verticils very long. Head yellow, somewhat
paler behind.

cRANE-Ft/Es FRolv[?HILE. -p. Arexander
Charres

Pronotum and pretergites pale yellow. Mesonotum fulvous,thelateral praescutal margin and humeral triangles
clearer yellow. pleurafulvous; dorsopleural membrane yellow. Halteres paleyellow. Legs
with the coxae and trochanters yltto*; remainder of legs yellow,theouter tarsal segments brownish black; modified subbasall"irrg of fore
tibia of male conspicuous,dark brown, a little more extensive
than thepaler base. Wings pale yellow, the veins and maerotrichia
darker yellow.
venation: Rz lying slightly distad of the level of r-m; cell Mg
deep, its
petiole correspondingly shortened, approximately twice rn-cu;
vein Znd,Along, sinuous,ending some distanceblyona the leveI of m-cu.
Abdomen yellow, the hypopygium somewhat more fulvous. Male
hypopygium (Fig. 2) with the veniral lobe of the basistyle,b,longproduced; the mesal lobe much shorter. Both dististyles placedat baseof ventral lobe, their points of insertion close together. Outerdististylewith both outer arms short, obliquely truncated at tips, separated
by asmall oval notch. Basal dististyle, bd, a long simple spine, its
basal third
more twisted. Phallosomic plate, p, glabrous, gutrtty widened
outwardly,
the apex subtruncate.
Holotgpe, male, Termas de Rio Blanco, Malleco, altitude
l0b0
meters, March 20, 19b1, ( pefla ) .
I take great pleasure in naming this very distinct fly forthecollector, Sefror Luis E. Pefra Guzm6n, who ."pir.rrud all of thespecies
discussed in the present report. By my key t; the Chilean species-SouthofMolophilus ( Diptera of Patagonia andChile, 1 : 200-20i; Lg2g),
the present fly runs to Molophitus (Molophilus) pirioni Alexander,anentirely different fly. In its general appearance and coloration,
it moreresembles M- (M.) flaaidus Alexander and allies, which belong
to anentirely different group of the genus.
52

Molophilus (Molophilus) perertens?rs, sp. n.

Belong to the plagiatus group, flaaidus subgroup; general coloration dark brown, sparsely pruinose; halteres uniformty yettow;
wings
broad, with a strong dusky tinge; male hypopygium with the outerlobeof the basistyle provided with unusually l;;;setae; outer dististylestrongly widened outwardly; basal dististyle proloundly bifid, theouterarm long and slender, very gradually narrowed. into " lorrg apicalspine,
beyond midlength of mesal face with a compact group of "Uo.rt adozen
appressed spines; inner arm likewise narrowed to a simpte acute spine,
near base with two or three strong spines.
Male. Length about 4.b mm.; wing b mm.; antenna about 1.8 mm.
Rostrum and palpi black. Antennae black, approximately one-

10

Asr. Tdc. Chite.-

Afio X/ NP I

fourth the tength of wing; flagellar segments oval, with very long
verticils, especiatty on the more proximal segments,and with a shorterd.ensewhite pubescence. Head dark gray.
Pronotum dark brown, the anterior pretergites narrowly yellow.
Mesonotum dark brown, sparsely pruinose; humeral region very restrictedly obscure yellow. Plu,rr" dark brownish gray, the dorsopleuralregion more yellowed. Halteres uniformly yellow. Legs with the coxae
testaceousyellow; trochanters yellow; remainder of legs brown, the outertarsal segments somewhat darker; fore legs broken. Wings broad, witha strong dusky tinge, the prearcular and costal regions a little morebrightened; ,r"irrr pale brown, the macrotrichia darker. Venation: Rg'm'cu;
Iying just beyond the level of r-rn; petiole of cell M.g about twicevein Znd"A moderately sinuous, ending beyond one-third the length ofthe petiole of cell M.r.
Abdomen, including hypopygium, brownish black. Male hypopygium (Fig. 3) with the beak of the basistyle, b, long and slender,rito"gty curved, blackened; surface of outer lobe of basistyle with un,rr,r"lly long setae. Outer dististyle strongly widened outwardly, thetwo arms very dissimilar, the outer with an oblique blackened ridge
before the obluse tip. Basal d.ististyle, bd,, distinctive, profoundly bifid
as in the subgroup; the glabrous outer arm long and slender, very
gradually narrowed into a long apical spine; on mesal face just beyond*iat"ttgitt *ittt a compact group of about a dozen appressed spines but
no setae; inner arm of style about two-thirds as long, Iikewise narrowed
to a simple acute spine, near its base with two or three strong spines.
Holotgpe, male, Termas de Rio Blanco, Malleco, altitude 1050
meters, March 20, 1951 (Pefra).
The present fly is most similar to species such as Molophilus(Molophrlus ) semulatus Alexander, differing in the coloration and,especially, the structure of the male hypopygium'

53

Motophilus (Molophilus) dilatus, sp. n.

Belongs to the plagiatus group, llauidus subgroup; general coloration yellow, including the antennae, halteres and legs; male hypopygium with the curved apical spine of the basistyle and both disiiriyf"r heavily blackened; basal dististyle profoundly bifid, the outerarm longer, its stem slender,scabrous,the outer fourth a little expanded,
the inner margin with about 10 to L2 appressedblackened teeth, thesetae lacking * ,r"ry reduced.in number; inner arm terminating in anacute spine, the outer margin at near midlength produced into a broad
flange or crest, its margin microscopicallytoothed.

CRANE-FLIES FROIT CHILE. -QhsTl$P. Alexander

11

Male. Length about 4.2 - 4.5 mm.; wing 4.G - b mm.; antenna
about L.4 - 1.5 mm.
Rostnrm yellow; palpi brown. Antennae (male) relatively long,nearly one-third as long as the wing or body, obscure yellow to brownish
yellow; flagellar segments long-oval, with very long verticils. Head paleyellow.
Thoracic dorsum reddish yellow, the pleura clearer yellow. Halteres pale yellow throughout. Legs yellow, the outer tarsal segments
more infuscated. Wings subhyaline, the veins pale brownish yellow,
macrotrichia darker. Venation: Petiole of cell Ms about one-half longer
than rn-cu; vein 2nd A long, slightly sinuous, ending just beyond thelevel of m-cu
Abdomen, including hypopygium, obscure yellow. Male hypopygium (Fig. 4) with the basistyle, b, terminating in a strong curved
blackened hook. Outer dististyle very unequally bifid at apex, the outerarm a gently curved blackened rod, the inner one broadly flattened, its
apex subtruncate. Basal dististyle, bd, profoundly bifid, as in the flauidus subgroup, both arms entirely blackened; outer arm longer, its
stem slender, scabrous,with about the outer fourth a little expanded,glabrous, its inner edge with a series of about 10 to 72 appressedblackened teeth but with few or no setae, the apex a strong straight spine;
inner arm shorter, terminating in an acute spine, the margin at near
midlength produced into a broad flange, its margin micros,copically
toothed or crenate.
Holotgpe, male, Termas de Rio Blanco, Malleco, altitude 400
meters, March 5-27, 1951 (Per1a). Paratopotgpes,66. Paratgpes,
6 6 , Curacautin, Malleco, altitude 400 meters, December B-18, 1950

(Pefra).
Among the rather numerousspeciesof the flaaidus subgroup,

the present fly seems most nearly related to Molophilus (Molophilus)
infleribilis Alexander, of Patagonia, especially in the expanded innerarm of the basal dististyle. It differs evidently in the details of structureof the remainder of the male hypopygium.
54

Molophilus (Molophilus) metuendus, sp. n.

Belongs to the plagiatus group; general coloration reddish brownto darker brown; antennae (male) short; halteres pale; legs brown; malehypopygium with the beak of the basistyle long and slender, curved;
basal dististyle profoundly branched near base, the axial arm expanded
at d.istalend, its outer margin with a series of about 10 to 12 obtuse
teeth, the apex a long slender spine; on lower face a short distance fromthe base with a long powerful simple rod that terminates in a strong
spine.

L2

Asr. Tdc. Chite.-

Afio XI NP I

Male. Length about 3.8 - 4 mm.; wing 4.5 - 5 mm.; antenna about
1.0 - 1.2 mm.
Female. Length about 4.5 - 4.6 mm.; wing 5 mm.
Rostrum and palpi brownish black. Antennae brown, the flagellum
somewhat paler than the basal segments; flagellar segments oval, withlong conspicuousverticils. Head brownish gray to clear gray.
Pronotum brown, the scutellum and pretergites narrowly whitened. Mesonotum reddish brown to brown, the humeral region of thepraescutum restrictedly brightened. Pleura dark brown; dorsopleural
membrane pale. Halteres yellow, in cases the knobs appearing darker
because of setae. Legs with the coxae brownish testaceous; trochantersobscure yellow; remainder of legs brown. Wings tinged with brownish
gray, the base and costal region more yellowed; veins and macrotrichia
brown, the latter darker. Venation: Rz lying distad of the level of r-m;
petiole of cell Ms relatively long, nearly three times rn-cu; vein Znd A
long, very gently sinuous, ending about opposite one-third the lengthof the petiole of cell Me.
Abdomen, including hypopygium, dark brown. Male hypopygium(FiS. 5) with the beak of the basistyle, b, long and slender, strongly
decurved to an acute point. Outer dististyle with the two arms very
unequal, the inner slender, curved. Basal dististyle, bd, distinctive,
profoundly branched near base, the axial expanded at distal end, theouter margin of the expanded part with a series of from 10 to 12 obtuse
teeth; apex of style thence produced into a long slender, slightly upcurved spine; basal arm of style a powerful simple rod that extends
virtually to the apex of the main arm, at tip narrowed into'a long spine.
Holotgpe, male, Curacautin, Malleco, altitude 400 meters, December 18, 1950 (Pefra). Allotopotgpe, g, pinned with type.Paratopotgpes, 6 9, December B-18,1950 (Pefi,a).
In some respects, the present fly suggests certain dark-coloredspecies of the flauidus subgroup, but from the structure of the basal
dististyle of the male hypopygium scarcely seems to be placeable withthem. In the present fly, the common base of the style is much longer
than in virtually all the species being more as in the otherwise quitedistinct Molophilus (Molophilus) gytnnocladus Alexander.

55 Molophilus (Molophilus) fortidens, sp. tl.
Belongs to the plagiatus group; general coloration of mesonotal
praescutumand scutum reddish, the postnotum and pleura dark plumbeous brown; legs obscurebrownish yellow, the subbasalfore tibial ringof male conspicuouslydarkened; wings broad; male hypopygium withthe basal dististyle a lrcwerful three-spinesstructure, one spine being

CRANE-FI/ES F ROM CHILE.

CharlesP. Alexander

13

on the outer margin before midlength, the remaining pair on outer halfof a broad flattened plate; phallosomea strongly bilobed cushion,provided with long densesetae.
Male. Length about 4.3 - 4.8 mm.; wing 4.5 - 5.5 mm.; antenna
about L.2 - 1.4 mm.
Female.Length about 4 - 4.5 mm.; wing 4.5 - 5.5 mm.
Rostrum and palpi brown. Antennae relatively long, approximately one-fourth as long as wing, dark brown, the more proximal segmentsa trifle paler; flagellar segmentssuboval, with truncated ends; verticils very long, especially on the more proximal segments,the longest
nearly three times the segments;a pale dense pubescence.Head gray.
Pronotum and pretergites whitened. Mesonotal praescutum andscutum reddish, V€rY weakly pruinose, the praescutum more heavily
darkened medially in front; scutellum yellow; postnotum and pleura
dark plumbeousbrown. Halteres yellow, the knobs very slightly darker.
Legs with the coxae yellow, the fore pair slightly darker; trochanters
yellow; remainder of legs obscure brownish yellow to pale brown, theouter , tarsal segmentsdark brown; subbasal fore tibial ring of maledarkened, conspicuous.Wings broad, yellow, the outer half a trifle
more darkened. Venation: R2 lying immediately distad of level of r-m;
petiole of cell Ms short, only about one-third to one-half longer thanrn-cu; vein Znd A sinuous,ending shortly beyond level of m-cu.
Abdomen brown, the hypopygium more brightened, in cases
yellow. MaIe hypopygium (Fig. 6) with the beak of the basistyle,b,slender, only gently curved. Outer dististyle with the arms very
unequal, the outer one flattened, its apex truncated, with the innerangle farther produced into a short point. Basal dististyle, bd, appearing as a powerful three-spinedstructure, including a long spine on outermargin before midlength; beyond this point the style is broad, forking
on outer half or less into two stout spines. Phallosome,p, a strongly
bilobed cushion,the surface with denselong setae. Aedeagus,c, weakly
dilated at near midlength.
Holotgpe, 6, Termas de Rio Blanco, Malleco, altitude 1050
meters,March 5, 1951 (Pefra). Allotopotgpe, 9, pinned with type. Paratopotgp€s,3 I .
Molophilus (Molophilus) fortidens is most nearly related to
speciessuch as M. (M.) appressusAlexander and M. (M.) subappressus
Alexander, differing most evidently in the structure of the male hypopygium, particularly the basal dististyle.

L4
56

Aqr. Tic. Chile. -Afio XI No I

Molophilus (Molophilus) furcus, sp. n.

Belongs to the plagiatus group; general coloration brownish yellow
to darker brown; male hypopygium with the basal dististyle a powerful
blackened rod, at midlength forking into two subequal spines, the inner
one a little shorter; phallosomean oval setiferousplate, the apex entire,
truncate.
Male. Length about 3.5 - 4 mm.; wing 3.9 - 4.6 mm.; antennaab,out1.3 - 1.5 mm.
Rostrum obscure yellow; palpi black. Antennae with the basal
segmentsyellow, remainder of flagellum brownish black; basal segments
enlarged; flagellar segments oval, the unilaterally distributed verticils
greatly exceeding the segments. Head light gray.
Thorax almost uniformly brownish yellow to ochreous, thesurface sparsely pruinose. Halteres whitened. Legs with the coxae andtrochanters yellow; remainder of legs pale brown. Wings with a grayish
tinge, the prearcular and costal fields more yellowed; veins brownish
yellow, the macrotrichia dark brown. Venation: R: in approximatetransverse alignment with r-rn; 'rn-cu oblique, varying from about twothirds the petiole of cell Ms to subequal in length to it; vein 2nd A
sinuous, ending about opposite midlength of n1,-cr.L.
Abdomen, including hypopygium. dark brown. Male hypopygium( Figs. 7, 7A) with the beak of the basistyle, b, slender, decurved. Outerdististyle with the arms unequal, the inner one slender, cylindrical.
Basal dististyle, bd, appearing as a powerful blackened rod, at midlength
forking into two unequal spines, the inner one a little shorter. Phallosome, p, an oval plate, covered with abundant relatively short setae,
the apex entire, truncate. Aedeagus, o, long and slender, a trifle dilated
shortly before midlength.
The paratype is smaller ( the smallest measurements given ) anddarker, especially the rostrum and mesonotum. There is a slight difference in the shape of the basal dististyle but the two flies must be
regarded as being conspecific.
Holotgpe, d, Termas de Rio Blanco, Malleco, altitude 1050 meters,
March 5-27, 1951 ( Pefra) . Paratgpe, 6 , Curacautin, Malleco, altitude
400 meters, December 6-24, 1950 (Peffa).
In the nature of the phallosomic plate of the male hypopygium,
the present fly is most like Molophilus (Molophilus) rubidithorl"r Alexander, which differs very evidently in the structure of the basal
dististyle. Most other allied specieshave the outer end of the phallosome
deeply emarginate.

CRANF-IL/ES

57

FROM CHILE.

Chsrles P. Alexander

15

Molophilus (Molophilus) monoctenrls,sp. n.

Belongs to the plagtatus group; general coloration brownish gray;antennae of moderate length, approximately one-fourth the wing; malehypopygium with the basal dististyle appearing as a powerful blackened
rod, strongly pectinate on outer margin, including a series of about sevenbranches, the outer three spines much reduced; phallosome a bilobed
setiferous cushion.
Male. Length about 4.5 - 4.7 mm.; wing 5.1 - 5.5 mm.; antenna
about 1.3 - 1.4 mm.
Rostrum and palpi brownish black. Antennae moderately long,
approximately one-fourth the length of the wing; scapeyellow, remainderof organ brownish black; flagellar segments long-oval, with truncatedends; verticils of the more proximal segmentsvery long, approximately
two and one-half times the segments themselves; an additional dense
white pubescence.Head dark gray.
Pronotum brown, the pronotum obscure yellow; anterior pretergites more whitened. Mesonotal praescutum virtually covered by three
confluent brownish gray stripes, the humeral triangles and narrow lateralborders yellow; most of remainder of notum brownish gray, the scutellum
obscuretestaceousyellow. Pleura brownish gray, the dorsal sternopleurite
a trifle paler; dorsopleural membrane yellow. Halteres whitened. Legs
with the coxae and trochanters yellow; remainder of legs yellow, theouter tarsal segments darker; fore tibia of male with the subbasal ringbrown, relatively broad. Wings broad, strongly tinged with yellow;
veins darker yellow, macrotrichia brown. Venation: Rz virtually intransverse alignment with r-rn, R:r*s*+ thus unusually short, arcuated;petiole of cell Mg about one-fourth longer than 'm-cu; vein Znd A strongly
sinuous, ending about opposite the cephalic end of rn-clt'.
Abd.omen dark brown, the hypopygium conspicuously yellow.
Male hypopygium ( Fig. B) with the beak of the basistyle, b, slender,
darkened, with conspicuoussetae surrounding its base. Outer dististylewith the two arms only slightty unequal, the outer one broadest. Basal
dististyle, bd, distinctive, appearing as a powerful blackened rod,strongly pectinate on the outer margin, including a series of about sevenbranches, the most basal before midlength, shorter than the subequal
second and third, the latter at near two-thirds the style; fourth spine
about one-half the size of the basal one; outer three spines progressively
more reduced, the last placed close to the short apical point. Phallosome,p, d bilobed cushion that is covered with unusually long and conspicuous
setae and with scattered fleshy setiferous protuberances. Aedeagus, a,long and slender, rnore dilated at near midlength'

16Asr. Tic. Chile. -Afio XI No I

Holotgpe, male, Termas de Rio Blanco, altitude 1050 meters,
March 5-27,1951 (Pefra).Paratopotgpe,6 .
The most similar describedspecies is Molophtlus (Molophilus)
perfidus Alexander, which is well-distinguishedby the structure of themale hypopygium, particularly the basal dististyle. The hypopygium ofM. (M.) peetinatusAlexander also is generally similar but quite distinct
in the various details.
58

Molophilus (Molophilus)

apicidens, sp. n.

Belongs to the plagiatus group; general coloration brownish yellow;
antennae short; halteres and legs yellow, the outer tarsal segments
brownish black; wings brownish yellow; male hypopygium with theventral lobe of the basistyle unusually slender; outer dististyle withboth apical arms appearing as slender spines; basal dististyle a long
slender rod, at apex expanded into a blade, each outer apical angle farther
produced into a curved black spine; phallosorhe truncated at apex, thesurface microscopically setuliferous.
Male. Length about 3 mm.; wing 3.7 mm.
Female. Length about 4.5 mm.; wing 5.2 mm.
Rostrum yellow; palpi brownish black. Antennae short, the outer
segments broken; seape and pedicel yellow, flagellum a trifle darker;
flagellar segmentsoval, about one-half as long as the longest verticils.
Head gray.
Thorax almost uniformly brownish yellow, the pronotum andhumeral triangles of the praescutum clearer yellow. Halteres yellow.
Legs yellow, the outer tarsal segments brownish black; fore tibial darkened ring of male narrow but distinct. Wings weakly brownish yellow,
the prearcular and costal portions clear light yellow; veins slightly darker
than the ground; macrotrichia dark brown. Venation: Rz and r-m virtually in transverse alignment; petiole of cell M3 approximately three .
times rn-cu; vein Znd A gently sinuous, ending a short distance beyond
ITL-CU,

Abdomen brown, the hypopygium a trifle more brightened. Male
hypopygium ( Fig. 9 ) with the lobe of the basistyl€, b, unusually slender,
the beak stout and scarcely curved; mesal face of lobe more setuliferous.
Outer dististyle slender-stemmed,the two apical arms blackened, appearing as slender spines, the outer one longest, curved at tip. Basal dististyle, bd, a long slender rod, at apex expanded into a blade, each outerapical angle of which is farther produced into a curved black spine.
Phallosome, p, gently widened outwardly, the apex truncated, surface
microscopically setuliferous. Aedeagus, a, relatively small and slender.

C R A N E -F L IE S I:R OM CIIILE.

Char lesP. Alcxander

L7

Holotgpe, male, Curacautin, Malleco, altitude 400 meters, December 19, 1950 ( Pefra) .
This fly is most similar to speciessuch as Molophitus (Molophilus)
oppressusAlexander, and allies, differing particularly in the struetureof the male hypopygium, especially of the styli.

59 Molophr.lus (Molophilus) leonurus, sp. n.
Belongs to the plagiatus group; general coloration of thorax
reddish yellow, the dorsal pleural region slightly darker; head yellow;
male hypopygium with the basistyle elongate, at apex produced into adecurved double beak; outer dististyle primarily bilobed, the outer arm
emarginate to form two lobules; basal dististyle a long ,curved simple
rod, gently expanded on outer end which splits into approximately ascore of points or fimbriations; phallosome glabrous; aedeagusconspicuously blackened, its lower face with microscopic spinulae.
Male. Length about 4 - 4.2 mm.; wing 4.5 - 4.8 mm.; antenna
about 1 mm.
Female. Length about b mm.; wing 5 - i.Z mm.
Rostrum obscure yellow; palpi dark brown. Antennae relatively
short, obscure testaceous yellow, the outer segments slightly more d,arkened; flagellar segments long-oval with truncated ends; verticils, especially of the more proximal segments, very long. Head yellow, theposterior vertex more or less darkened medially.
Pronotum yellow. Mesonotum chiefly reddish yellow, the pretergites and humeral region of praescutum clearer yellow. Pleura with apale brown dorsal longitudinal stripe, the ventral portions and dorsopleural region yellow. Halteres pale yellow. Legs with the coxae andtrochanters yellow; remainder of legs yellow, the outer tarsal segments
more infuscated; darkened ring of fore tibia of male narrow but conspicuous, darkest in color on the outer surface. Wings pale yellow, thg
prearcular and costal regions somewhat more saturated; veins yellow,
macrotrichia brown. Venation: Rz lying a short distance beyond thele'reI of r-rn; petiole of cell Ms relatively long, a little less than threetimes rn-cu; vein 2nd A sinuous, ending some distance beyond m-cu.
Abdomen pale brown, the hypopygium darker. Ovipositor withthe valves, especially the cerci, long and slender, only gently upcurved.
Male hypopygium (Fig. 10) distinctive in all details. Basistyle, b, elongate, narrowed outwardly, the apex a decurved double beak. Bothdististyles arising close together at near midlength of the style. Outerdististyle entirely blackened, primarily bitobed, the outer lobe again
deeply emarginate to form two lobules; basal lobe longest, cylindrical,

1B

tlqr. Tcc. Chile. -Aiio X/ Np I

simple; outer bifid lobe conspicuously setiferous. Basal dististyle, bd,
a long curved simple rod, gently expanded at outer end which is conspicuously fimbriate with nearly a scofe of slender spinous points, someof rvhich again are split apically. Phallosome, p, glabrous, narrowed
outwardly, the apex truncate. Aedeagus, a, conspicuously blackened,
curved on basal third, the lower face for much of the length with microscopicspinulae.
Holotgpe, male, Curacautin, Malleco, altitude 400 meters, December 6-24, 1950 (Pefra). Allotopotgpe, female, pinned with type. Paratopotype, I male.
The present fly is so distinct from all other regional members of
the genus that no clcse relatives can be indicated. The double beak of
the basistyle, trilobed outer dististyle, and very peculiar structure of thebasal dististyle all mark the fly as being entirely distinct. The mostsimilar speciesis perhaps Molophilus (Molophilus) pretiosus Alexander.

SUMARIO

El autor continira en este trabajo sus anteriores publicaciones sobre Tipulidae ( Diptera ) de Chile que con el mismo tftulo han aparecido
en esta Revista. En el presentc trabajo se describe diez nuevas especies
chilenas del g6n ero Molophilus Curtis, todas procedentes de Curacautin,
provincia de Malleco, Chile.

q
:[d

I,'7
;p

l0
EXI'LANATIONOTI FIGURES

m,ottostyFig. 6. trIoloph,llus (IlI olopltihts .fortitlctt.s,_ - - Fig. 1. trIoloTtltiltts (Illololtltiltts)
loitles, sp. n.
sp. ll.
Irig. 2. Ifiolopltiltts (Molopltilu,s) peiiai, sp. n.
(tr[olophiltts)
IriS.
7. IloloTtlr,iltts!tu'ctts,
Fig. 3. trIolo1thilu,s (trIol<ttthilus) pet eitensp. n.
8tt.s, sp. n.
tttttu,octeIrig. 8. trIoloph,ihr,s (III clottltiltts\
FiS.
4. Molopltdhr,s (Illolophiltts)
dilatu,s, , t ? { s . .s p . n .
sp. 11.
}'ig. 9. Illolopltilnts (IlI oloplriltts) apicidens,l-ig. 5. trtolophilus (IVolottltilus) metu,end,u,s, s p . n .
sp. n.
Fig. 10. trIolopltiltts ( LI olottltihts ) leonuru,s,
sp. n.

(Explanationof symbols: a, aedeagus;b, basistyle;bd, basaldististyle;d, dististyle;p, phallosome;ub, ventrallobe of basistyle).