/'i "/10

INSECUTORINSCITIE MENSTRUUS

UNDESCRIBED SPECIES OF ANISOPODIDAEFROM NEW ZEALAND_PART II(Diptera)
BV CHARLES P' ALBXANDER
The first part under this general title was published in 1923
The new speciesdescribedat(Ins. Ins. Mens., I7,73-7+).
tiris time were collected by Mr- T. R' Harris in Westland, andat Taumarunui in the North Island, and by Mr' Leon Curtis
in Otago. Recently, Mr. Edwards has made the interesting
discovery that all of the New Zealand crane-flies hitherto described as species ol Trichocera should be referred to Para'cladura Brunetti, which was based upon a species from northern
India. The genus is distinguished from Trichocero chiefly by
the venation and the much abbreviated basitarsi' Besides thetwelve species now known from the Maorian subregion, Poracladwralncludes four species from India and Formosa, andP. trichoptera (Osten Sacken) of western North America'
The types of the new species described forthwith are preserved ln the writer's collection through the kindness of thecollectors, Messrs. Harris and CurtisParacladura harrisi, new sPecies.
Male.-Length about 3.? mm.; wing 4'4 mm'Rostrum, palpi, antennae and head dark brown' Mesonotumand pleura uniformly fuscous. Halteres pale' the knobs brown'
legs
Legs with the coxae yellowish brown; remainder of thelittleasegments
tarsaltheterminal
only
with
p"l" bro*r,darker. Wings subhyaline, the veins dark brown' Venation:
Sc, ending opposite r, Sc" at about midlength of R'i,' Rr*u alittle longer than the basal section of Rr; r-rn oblique, inserted
at the fork of.M ;cell rst M, closed but rruweak; cell M' about
twice its petiole ; rn-cu about two-thirds m'
Abdomen dark brown throughout. Male hypopygium withthe disti-styles moderately elongate, the mesal face near thebasal third proclucecl into an obtuse lobe which is entirely
covered with microscopic setaeas is the remainder of the mesal
face. Each gonapophyse appearing as a flattened plate, theINSECUTOR INSCITI.4

MENSTRUUS

IIcaudal margin of which is produced into four or five conspicuous spines, in the type there being five on the right side
und fout on the left; beneath this apophyseand possibly a partof it, appears a complex structure which is densely set withappressed,feebly curved spines.
Habitat.-New Zealand ( South Island).
Holotype, /, Rewanui, Paparoa Range, Westland, altitude
600-800 feet, February 15, 1923 (T. R. Harris).
Paraclad.ura herrisi is named in honor of the collector, Mr'
Thomas R. Harris, to whom the writer is vastly indebted forinvaluable collections of New Zealand crane-flies. It is mostnearly related to the larger P. antipodeum (Mik.) of theAuckland Islands; the latter species differs in the obliterationof m-cu, and in the details of structure of the hypopygium, as
the short, broad-based aedeagus, the differently constructedgonapophyses,and other details.
Paracladura curtisi, new species.
Male.-Length 2.6-2.8 mm.; wing 3.4-3.6 mm.
Female.-Length 3-3.3 mm.; wing 3.64.2 mm.
Head and appendagesdark brown. Mesonotum dark brown,
the lateral ends of the suture and the scutellum a little paler.
Pleura brown. Halteres dark brown, only the extreme base of
the stem paler. Legs dark brown throughout. Wings subhyaline, the veins dark brown. Macrotrichiae of veins longand conspicuous; base of cell rst A with macrotrichiae.
Venation: Sc, ending some distance beyond r, Scrbelore midlength of Rs,' r on R, a little more than its own length beyond
the end of Rs; r-m jast beyond the fork of M on Mrrr;cell M,
comparatively shallow, subequal to its petiole; cell rst M,
open by the atrophy of rn; m-cu longer than r in the J, subequal in the t, in alignment with the basal section ol M"';cell znd, A natow.
Abdomen dark brown, including the hypopygium. Malehypopygium of the same simple structure as P. ruacrotrichiata(Alexander) ; dististyles cylindrical, not at all produced onmesal face. Aedeagus and gonapophyses of very simple
structure, the latter subtending and exceeding in length the12INS4CUTOR INSCITI4MENSTRUUS

former, appearing as slender, siiaigtt rods directed caudadand lying'lr"ry .lot" together along the median line of thebody. Just beyond mid-length of the aedeagus'on either side,
appearsa small oval lobe, gently divergent, directed caudad'
Ovipositor obscureyellow, the tip darker.
Habitat.-New Zealatd (South Island).
Holotype,J, Ben Lomond, Otago,altitude 2,500ieet, April't,1923 (L. Curtti).
AllotoltotyPe,].
ParatopotyPe.t,15 f$a
Paraclod.uracurtisn isi named in honor of the collector, MrLeon Curtis, who has collectedmuch valuable material in thevicinity of Lake Wakatipu and on Stewart Island. It is mostcloselyallied to the larger P. marrotnchiata (Alexander) fromwhich it differs chiefly in the details of structure of the malehypopygium and in the venation, especiallyin the relatively
shallow ceTlM* The macrotrichiaeare neither so numerous
nor so extensively distributed as in macrotrichi,ata. In thelatter species,the aedeagusextendsfar beyondthe gonapophyses
which are recurvedand decussateacrossit.
Paracladura decussata,nerv species.
Male.-Length about 2.8 mm.; wing 3 mm'
Generally similar to, and a close relative ol, P. obtusi,cornis(Alexander), from which it difiers chiefly in the structureof
the male hypopygium. Head dark grey. Mesonotdm shiny
brown. Wings paler. Venation: Sc, ending shortly beyond
the fork of Rr*, Sc, about oppositeone-fourth the length ofRs; rn faint, beyondmidlengthof the petioleof cell Mr; cellznd,A long and narrow.
Male hypopygium with the dististyles relatively short, cylindrical, feebly arcuated, entirely without lobes. Gonapophyses
very conspicuous,appearingas powerful chitinized arms, directed caudad and thence mesad so as to be decussateacrossthe genital chamber,the extreme tips suddenly narrowed andacute. ln P. obtusicornis,besidesthe flattened, obtuse plates,
ihere are slender and very delicate,strongly curved horns directed cephaladand thencemesad.

INSECUTOR INSCITI,{MENSTRUUS

13Zealand ( North Island).
H abitaf.-New
Taumarunui,
April 30, 1923 (T. R' Harris)'
Holotype, J,
Paracladura complicata, new species'
Male.-Length about 3.2 mm.; wing 3.7-3.8 mm.
Female.-Length about 3.4 mm.; wing 4 mm.
Generally similar to and a close ally of P' lobifera (Alexander), fror.n which it difiers in the darker coloration and thevery distinci structure of the gonapophyses of the hypopygium'
The geneial coloration is much darker than in lobi'fera but
paler than'"in maori, The anterior part of the mesonotal
praescutum is darker than the sclerites behind the suture.
Wings pale brown, the veins darker. Venation: rn lar out on
the petiole of cell M' usually less than its own length fromthe fork, due chiefly to the great depth of cell Mr.
Male hypbpygium with the dististyle bearing a conspicuous,
subbasal lobe on mesal face, as in lobifera, the apex of thislobe subacute and glabrous. The armature of the genital chamber is very intricate; the lateral pair of plates bear one or two
apical spines'that are directed caudad, and two or three longspines on the outer or lateral edge that are directed laterad.
A transverse slender bar lying across the chamber has themesal end lferminating in two powerful spines, the mesal one
small, the lateral one about twice as large, directed chiefly
caudad; lateral or outer ends of this bar extend laterad andterminate in numerous powerful ctrrved chitinized hooks, theterminal ones largest, becoming smaller mesally and obsolete
near midlen{th of each arm; the arms of either side lie almost
in a straight line across the genital chamber. What seems tobe the aedeagus is a massive, central ptate, each lateral angle
produced into a slender arm directed caudad, the tips strongly
mesad, to produce a somewhat lyriform appearance.
Habitat.-New
Zealand, (North Island).
Holotype, J, Taumarunui, April 30,l99g (T. R. Harris).
Allotopotype, J,
Paratopotypes, 5 {{;paratypes, Z 33, Ohakune, May 10,
1923 (T. R. Harris).