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NEw oR LITTLE.KNowN.
TIPULID"&Fnom rrln
PHILIPPINES (DIPTERA), PART III

By Cnnnr,os P. ALD:(ANDEEOf Amherst, Massachusetts

Separate from:THE PHILIPPINE JOURNAL OF SCIENCE
Published by the Bureau of Science of the Philippine Government.

Manila. P. I."
.
J-

.

,.

:'

VoLUME28, No.3, NovaMcrn,1925-.

zizzs6-,iliANILABUNEI|U oF PRTNT'NG
1 925

,

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il
NEW OR LITTLE-KNOWN TIPULID& FROM THE
PHILIPPINES (DIPTERA), PART III
By Cnanr,ns p, ATEXANDEn
Of Amherst, Massachttssetts1
ONE PLATE

The present instalment is based primarily upon someinteresting crane-fly material taken i' samar bv i\{r. Richard
C. ru.Gregor. A few additionar specimens are from thevicinity of
Manila. The series of specimens labered "Loquilocon,i*"*ocollected 11 kilometers east of wright, samar, near
the setflement called Loquilocon. I am, g:reaily indebted
to Mr. Mc_
Gregor for his kindness in collecting and submitting:these
specimens. The types are preserved in my collection.
LI M O NI I N. €GeranomyiacomigeraAlexander.

plate 1, figs. 1, ?, and g.

Geranomyia cornigera ALEXANDEa,
fnsec. Inscit. Menst, 1 (1918)
137-139.
This remarkablecrane fly was described from a small
seriesof specimenstaken october zz-zg, rgLZ, at pettit
Barracks. Mr.
McGregor's materiar includes two maresthat were taken at
Manila, May 7, 1924. The peculiar charactersof *-n".nv
tubercle, t,, on the vertex urrd , small mediantubercle on the
cephalic margin of the mesonotal prescutum
are shown infic. z.
The venation and wing pattern are shownin fig. 1. These
specimens are better preserved.than the type
series and showthat the vein Sc, isbut faint, placed a short distance
before the origin of^present
Rs, near the center of a dark brotch.
Moreover, there is a similar rveak supernumerarycross vein in
the dark subcostal brotch at middistance
between arcurus andthe origin of Rs.
.
'Contribution
from
Agricultural
College.

theDepartmentofEntomology,

Massachusetts

970

The phitippine Journatof Science

The structure

discussedir,of fha hAr^ L-lu of the male hypopy5iu3(fig. 8) had not

re26

been

iliil:;u*'L:l;::#*'*;:##L{J:n*}'ifi
Basistyre,
r*:_#.ijf;la,rerati.iJ;.;_r,;';ilf,"il't,
iffi"rlTffl

i:1;"T covered
9lt:tttu,--i,,""*".andfleshy,
witrrj"J*l,i:1t",J:,f;j,Ty*:ffiii:,:"?j".,ii$."H""X,'",ffi*T1*"ffffi:ii:n*illl;
fr*'*i#Tli""
::,1,
{fr
una*ir,Ji

ately curvea,ifr"-

lT'1""*""?'"
conuooolr.fi:T}lf

L:ll:"orsaldististyle,d,,moder_lngurarryl,iu"1o*u,
mesal apical angle
slendeldirected .uuaud, thetip subacute und directedsliehfly laterad.
trimonia trigonia sanarensis
subsp. nov.

, yd:._tnnsth,':f;"TlfllL'X;,*:1,?:f,#'lliif;";i,Hi:;'*i*ty,-ff
9 miilimetersl v.

Size,larger, as shownby the_prescutum'hhr,du"k.r'.'ii,
t' i?"ffnarrowryblackish,.are-

iil:i:rTi,,T"Tffi".rijrate"ai;;;".
the *"i""ii" "n"ouary
dark brown; scutalr"n.,9!
1""r.'fr"o=*rr,the medianarea con_;ilff :'*,rfl "'j" ;:'^:t"*-a""r.l'oostaotar
mediotergite
dark:u,""T*iT.6f;;'f:'h#;161;i.""n'i.fl"ff:';l*"tmi:*:"ry*ri"""'.,*!iil$"$i{ii:,fiHaltereserongate,ou n"oin "ilt_""t rjf":r:?
;;:il:conspicuously- "rothe f.r"n,?flfocherous,
iliactisfr. W.ings*iif, H:ri.
tf,"
r.::
interspaces*1: ;at;'"fiti""lX""t"''.,*,
being
t"
1"1"
"
l;;;

iater
oiii'*'##i'i# Jl"^,ot1u'";;J$'+
Fl'.sternites
basatfour
of
at
*'
reasr
the
c;;.;,;,;:,::i: :_1,:39.bases--'=oosAMAR,
r,"n"a".i,Tffi'3l"rTjl
ifo,!#;n"*,.
0rimargula philippina

Alexander. plate! fiL. Z,
Orimargula phitippino,e"on*oo,for". tntttt' Menst'
5 (7917) 6-.7,
unique tvpu or thisspeeie. ,u--Ttu

Mr.McGregor
cou"ctuaoruiu]"

22, lg,4, that is i".,u"i"ilnl,,Hf,tilil"ffi,#
T_u.t, netter state of preservationthan
the type and is describedfr"""*i"r,f,"r"ufiotypi."l. The indicated'Journ. Fed. Malay StatesMus. 8 (1919) 1E_16.

2a,sAlenand,er: Tipulid,a, IIIg?1

differences between the two sexes are probably due in large partto the relative condition of the two types.
Femo,Ie.-Length, about 2.5 millimeters: wing, 8.6. Rostrum
and head dark brown. Cervical sclerites elongate, as in Ton_orhi,na and allied genera. Mesonotum dark brown, the scu_
tellum conspicuously pale: Pleura dark brown, with a paletransverse stripe extending from the wing root ventrad acrossthe cephalic margin of the pteropleurite, hypoepimeron, andmeron, onto the posterior coxre which are thus conspicuously
paler than the others. Halteres pale brown, the base of thestem conspicuously paler. Legs rather dark brovrn. Wings
as in the type; venation as in fig. 2.
SAMAR, Loquilocon, June 22, lg24 (McGregor); allotype,female.
Erioceralativentris Bezzi.
Eriocera lutiaentris Bozzr, philip. Journ. Sci.
$ D 12 (191?) 118.

SAuAn, Loquilocon, June 21, lg}4 (McGregor), a male.
Two females that are referred to this species were taken at
the same station on May 80, L924. However, these female
specimensdo not exhibit any dilation of the abdominal segments
as does the male and may not be corecily associated.
Xrioceraspatulatasp. nov.

Plate 1, fig. b.

Abdomen with the basal half slender, the segments cylindrical, the apical segments dilated into a conspicuous blackened
spatula; general coloration gray; legs black; wings strongly
suffused with blackish; a discal blotch and bases of the analcells dirty whitish.
Male.-I*ngth, 1g to 14 millimeters I wing, l1^.Zto I2.5. Rostrum arld palpi black, the former dusted with gray. Antenne
relatively short; scapal segments dark colored, dusted withgtayi flagellum pale yellowish brown, the terminal segments
darker; flagellar segments decreasing in length and diameter tothe end. Head light gray, with a small vertical tubercle.
Pronotum black, pruinose. Mesonotal prescutum graS theinterspaces with conspicuous erect black seta; three smooth,darker gray stripes, the lateral ones narrowly margined ivith
the median stripe broadty divirded meOlatty by a!fac_t<isf,blackish line; remainder of mesonotum dark gray, pruinose.
Pleura black, sparsely pruinose. Halteres short, stout, black.
Legs with the coxe and trochanters black, the fore coxa atrifle paler; remainder of legs blaek, the fore femora a very

372

The philippine

Jou,rnal of Science

TszE

little paler. Wings (fig. S) with a strong blackish tinge,
thebase and costal margin darker; a dirty white band crosses cells
1st R,, R, and M inrmediately before the cord; bases of cells
lst
A and 2d A extensivery dirty white; veins black. venation:sc,
ending opposite midlength of the basal section of R,, Sc,
beforethe. fork of
Rs elong:ate, feebly angulated ut o"igirr, u|,.,;little longer than R; r more than its length beyond thefork of
Rr*r; basal section-of Rn,u about equal to Rr.r; cell M,
verysmall, asymmetrical, M" being in alignment"'wittr lt,_";cell]st_M, relatively small, the proximar end weakry angutated s'horttv
before r-m; m-cu varying in position from before to beyond
midlength of cell lst M,, about one-half longer than the
distarsection of Cur; vein 2d A long, sinuous.
Abdomen unique in structure, so far as the family Tipulideisconcerned, in that the terminal segments are
dilated into aspatula, producing an appearance that is suggestiveof certain
Conopide and aculeate Hymenoptera; basal four
segments
reddish brown, blackened laterally, more narrowly
darkenedcaudally, very slender; the apex of the fourth
segment begins
to widen out and is blackened; segments b to Z
dilated, black;segments 8 and 9 gradually narrowed and shortened,
black.
fn one paratype, in both wings vein M, is deflected cephaladand unites with the tip of vein Ro,o, closing cell Ro.
SAMAR,Loquilocon, June 29, lg24 (McGregor) fou,males.
;
Elephantomyia(Elephantomyotles)
samarensis
sp. nov. plate 1, flC. 4.
General coloration of the thorax orange-yellow; legs brack,
the tarsi extensively light yellow; wings subhyaline, cel-il
sc andthe stigma infuscated; cer 2d A very narrow; abrdomen
brack,
the base of each segment narrowly yellowish.
Femnle.-Length, excludirig rostrum, 10.5 millimeters; wing,
7.4; rostrum alone, about 8. Rostrum long and slender, tro*rr_
ish black. Antenna brownish brack, the scapal segmentsatrifle paler; flagellar segments oval; basat flagellar segment
about equal in Iength to the following two taken togethe-r,themore distal segments becom,ing fusiform, all but the basal
fla_gellar segment with long delicate verticils. Headdark urolrnistr
gray; vertex between the eyes very narrow.
Pronotum yellowish brown. Mesonotum orange ororange_yellow, the scutellum and postnotal mediotergite
more infuscated. Pleura testaceousyellow to orange. Hartereslong andslender, black. Legs with the coxa and trochanters
coicoto-2s,s

Alenand,er: Tipulid,a, Iil373

rous with the pleura; femora and tibie blaek, the extreme basesof the former narrowly paler; basitarsi black, the tips abruptly
light yellow, this including a little more than one-fourth thesegment; segments 2 and 3 concolorous; segments 4 and 5 darkbrown. Wings subhyaline, cell Sc and the stigmal region, which
includes cells Sc, and 2d R,, dark brown; veins dark brown.
Venation (fic. 4) : Sc, ending shortly before the fork of Rs,
Sc, at its tip; Rs very strongly arcuated to weakly angulated
at origin, cell Lst R, being very wide; cell 2d R, abruptly reduced to a linear strip occupied by the stigma, gently widened
outrvardly; Rrr, perpendicular at origin, in alignment with
r-m,, beyond.the base bent at a right angle, with what seems
to represent r as a spur at the bend, this delimiting the proximal end of the stigma i Rn*o in direct alignment with Rs,
the vein,s at the end of the sector thus forming a cross; cell lst
M. rectangular; m-cu a little more than one-half its length
beyond the fork of M; distal section of Cu, shorter than m-cu;
distance on margin between Cu, and lst A about equal to the
distal section of Cu,; cell 2d A reduced to a narrow linear strip.
Abdomen black, the basal third or less of each segment conspicuously bright yellow. Ovipositor with the long straightvalves rusty horn colored.
SAMAR,Loquilocon, July 2, L924 (McGregor) ; a female.
The subgenas Elephantomyodes 3 was erected by me for the
Formosan E. major Alexander. The group includes a numberof Oriental and Australasian species. The present fly is distinguished from the other known species by the very narrow eell
2d A, in conjunction with the cruciform arrangement of theveins at the fork of the seetor.
Styringomyiamcgregorisp. nov. Plate 1, figs. 3 and 9.
Mesonotal prescutum ashy gray with linear black interspaces
posteriorly; legs uniformly dark brown; wings yellowish subhyaline, immaculate; vein 2d A elongate, not incurved to theanal margin.
Male.-T,ength, 6 to 6.5 millimeters; wing:, 4 to 4.8.
Female.-Length, about 4 millimeters; wing, about 4.
Rostrum and palpi brown. Antenna brown. Head brorvn,
the orbits and anterior part of the front broadly and eonspicuously whitish; gena and postgene darkened.
"Ann. Ent . Soc . Am . 16 ( 19 2 3 ) 6 4 .

374

The Phi.Iippi,ne Jou,rnal of Science

1e26

Pronotum light yellow, indistincily darkened mediaily. Me!onotal prescutum light ashy gray, narrowly lined posteriorly
with darker, these lines representing the usual interspaces; lateral margins of prescutum narrowly but conspicuously moreyellowish; scutal lobes darkened; scutellum conspicuouslyyellow
medially, the sides and the parascutella darker; postnotat mediotergite gray with a narrow yellowish median line. pleura
dark, t*re base of the wing more yeilowish. Harteres parebrown, the knobs somewhat darker. Legs dark brown, without
marking:s. Wings yellowish subhyaline, immaculate; veins a
trifle darker colored. venation (fig. B): vein 2d A erongate,
running straight to the margin, ending some distance beyond
the origin of Rs.
Abdomen pale yellowish brown, the lateral margins narrowly
darker colored; sternites more yellowish; hypopygium brownish
testaceous. Male hypopygrum (fiS. g) with the basistyles, b,
produced caudad into a long fingerlike lobe which bears several
elongate ,setre,the two that are more nearly apical in position
somewhat stouter, the subterminal sete on mesal face long andslender, curved. Dististyles, d, complex, the mesal lobe very
extensive, densely set with powerful, recurved brisfles, the styleterminating in a powerful, blackened, genfly curved spine.
What seems to be an interbasal process, i, appears as a genfly
curved, cylindrical, blackened rod.
LuzoN, Manila, October 1,lg%4 (McGregor) ; a male. A pairin copula at light on October Z,lg%4.
StEringomyi,a mcgregori is very distinct from all described
species in the cinereous mesonotum, the uniformly dark brownlegs, and the subhyaline, unmarked wings, with vein 2d A longand running straight back to the anal margin. The only otherknown species having the latter clharacter are S. aenustu Loew
(fossil in copal) and S. bi,punctata Edwards (Queensland).
The present speciesis unusually primitive in the scarcely modified
apices of the basistyles of the male hypopygium. This very
interesting fly is named in honor of the collector, Mr. Richard
C. McGregor, to whom I am indebted for many favors in thepast.
TIPULIN/E?selliophoraprafica fenestrellasubsp. nov.
Female.-Length, about 20 millimeters; wing, 20.8. Agreeing with typical prafica Bezzi in its general black coloration,differing in the following respects:

zs,sAlerander:

Tipul;ida, III

375

Branches of the flagellar segments relatively elongate, stout.
Wings dark brown, all of the cells of the wing with the exception of Sc and 2d R, with conspicuous pale gray centers, restricting the ground color to uniform eonspicuous searns to all
the veins and as a margin of equal width that completely encircles the wing.
SAMAR,Loquilocon, June 18, L924 (McGregor) ; a female.
Pselliophoratigriventris sp. nov. Plate L, fig. 6.
Allied to P. gaud,ens (Walker); general coloration orange;head scarcely variegated with darker; mesonotal prescutum withthree brownish blaek stripes; halteres orangd with brownish
blaek knobs; legs black, the femoral bases conspicuously orange,
broadest on the posterior legs; all tibia with white subbasal
rings; wings black, the base broadly and eonspicuously orangeyellow; a triangular paler yellow discal blotch; abdomen orang:e,the segments conspicuously cross-banded with black.
Female.-Length, about 20 millirneters; wing, L7.5. Frontal
prolongation of head yellow, blackened laterally; palpi darkbrown, the third segment more yellowish. Antenna with thescapal segments yellow; basal segment of flagellum elongate,obscure yellow, the ventral face infuscated; the terminal ten
flagellar segments dark brown, gradually becoming shorter andmore crowded, the terminal segment very small. Head orangeyellow, the occiput darkened behind.
Pronotum orange, the sides narrowly blackened. Mesonotal
prescutum orange, with three conspicuous brownish black stripesthat are contiguous near the suture, which is conspicuously
blackened medially; scutum orange, each lobe with two brownish
black spots on anterior [ralf; scutellum orange, the parascutella
black; postnotal mediotergite orange, narrowly margined anteriorly and posteriorly with brownish black, the latter marking
extending onto the postnotal pleurotergite and completely surrounding the base of the halteres.
Pleura orange-yellow,
conspicuously variegated with brownish black markings, theseincluding spots on the propleura, dorsopleural membrane, aspot on pteropleurite immediately ventrad of the wing root,and some smaller ventral markings. Halteres conspicuously
orange, the knobs brownish black. Legs with the fore andmiddle coxa extensively infuscated, the posterior coxe largely
obscure orange; trochanters obscure orange; fore.and middle
femora black, the bases conspicuously orange, this narrowest
on the fore femora, a little more extensive on the middle femora

376

Th,e Phili,ppi,ne Journal of Science

where about the basarthird is included; posterior femoraorangewith less than the apical third blackened; tibie
nack, eachwith a conspicuouswhite subbasal ring, appro*i*utuiy
in extent on all the legs or that of the tore tilia"qoufa trifle"wider;tarsi black. Wings (fig. 6) blackened,the basen ouafv u"Oconspicuouslybright orange-yelow; a roughly triangulardiscalarea of a paler shadeof yelrow, this including portionsof ceils
lst R' R, M, l,st M, an{dthe extreme proximal
ends of ceils
Rul{r, and Mn; pale brownish white washesin cells Cu andlst A on either side of vein 1st A; veins darkbrowrr, *o""orange-yellowin the flavous basal and discar areas.
venation:sc, preserved;del M, narrowly sessile;m-cu immediately
beyond the fork of M ..
Abdomentergite with a broad, conspicuous,-black""""g,'-'"u:h_ring acrossthe middre,reavingthe baseand apex broadryof the ground color; on tergite 7, the pale apexd.."u".", lotent, on tergites 8 and 9 being replacedby b1ack.
"*_
BasarJi"ta,of ovipositor conspicuousry blackened. oviposito"
st"uigrrt,
brownish black, the tips of the valves narrowly brightened. ,.
SAMaR,Loquilocon, June Lg, ]rg}4 (McGregort; a f"*af".

lPHrLrP. JouRN. Sd.,28,

ALEXANDER: TrPUr,rDE, IILIPLATE I.

No. 8.