, <-(lc</JJ

NBW OR I,ITTI,I'.KNOWN TIPUI,IDa: FROM TIII]
PHILIPI'INI'S (DIPTERA), PART IV

By CHenlns P. Ar,cxaNonn
Ol A m ILt'st,X'Iossut:hrrsclt.yS,'nlrrlttc ft'onr
TIIE

I'IIILIi'I'I\I.]I'JT-iI\ \I. ()1' S{--]I]'NCE

P u b i i - * h e d l t y t h e I l L r r t , i i , , tr , f S r . j r r r t , c, , 1 1 ! r t I ' h i l i t . r u i u e G o v e r . n m e n tllanilii,

1'. I.

Vor,ulra ii:,:],
No. J, Jutu, Lg27

MANILABUHDAU OF'PRINTING
1lJ:7

NEW OR LITTLE.KNOWN TIPULID.4EFROM THE
PHILIPPINES (DIPTERA), PART IV 'By Cn.o,nr,nsP. AT,nxANDER
Of Amh,erst, Massachusetts
TWO PLATES

Through the great kindness of Mr. Richard C. McGregor and
Dr. C. F. Baker, I have had for study a considerableamount ofadditional material in the Tipulida, some of which is discussed
in the present report. Through the kindness of the collectors,
I have been permitted to retain the types of the novelties discussedhere. Our knowledge of the crane flies of the Philippines,
although still very incomplete, has received its greatest impetus
through the kind interest of Mr. McGregor and Professor Baker.
A recent, very important paper on the Tipulide collectedby the late G. Boettcher has added greatly to our knowledgeof the subject. F. W. Edwards 2 has listed all of the papers
hitherto published that concern the Tipulide of the Islands
and has listed the speciesof the family hitherto recorded fromthe Philippines (about 56), to which are added 17 speciesfrom
the Boetteher material, making a total of about 73 species. The
exact number is uncertain, because of a question of correct
determination of some of the speciesrecorded and the possibilityof their being identical with earlier recorded forms. Edwards
has accomplisheda splendid piece of work in preparing this report and thus summarizing the earlier work. Doctor Baker
has called my attention to the fact that several of the localities
mentioned therein are misspelled and should be corrected; themistakes presumably were caused by poorly written labels.
A list of these follorvs:
Nos. 5 and 17. Leite - Island of Leyte.
Kalambugan - Kolambugan, in northwestern Mindanao.
No. 7. Bagnio = Baguio, Benguet Subprovince, Luzon.
Nos. 19, 28, and 30. Port Bauge - Port Banga, southwestern Mindanao.
'Contribution from the Department of Entomology, Massachusetts
Agri,cultural College.
'Philippine Nematocerous Diptera. I. Tipulidre, Notulae Entomologicae
6 (1926) 3344.
29L

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1927

No. 32. Heightpool (?) - Haight's Place, at 8,000 feet, in the mountains above Baguio, Benguet. This is the type locality for thisspecies (Pselliophora pu,milo Alex.).
No. 33. Calopan - Calapan, Mindoro.
No. 38. Antimonan - Atimonan, on the east coast of Luzon.

Keys to the larger and more involved genera (Eriocer,n and
Pselliopltora) are given. Bezzi' had earlier supplied keys to
Libnotes, Eriocera, and Psell;iophora. In the present report,
keys to the Philippine speciesof Scamboneuraand Trentepoh,tia
are supplied.
TIPULIN.IE
TIPULA

RMRAI

sp. nov.

Plate l, fige. I and 2.

General coloration dark fulvous; tibial spurs lacking; wings
subhyaline,the stigma dark brown; cell M, sessile; cell Zd Avery narrow.
Male.-Length about l3.b millimeters; wing, 15; fore leg,femur, 9.5; tibia, 11.6; basitarsus,16.8. Frontal prolongationof head relatively short and stout, brownishyellow; nasusstout,
concolorous;frontal prolongation meeting remainder of frons
at an acuteangle (when viewed laterally) ; palpi pale. Antenne
with the scapal segments yellowish brown; flagellum darkbrown; all flagellar segmentscylindrical with stout black sete
distributed along the outer face only, remainder of each segmentwith microscopicsetula; terminal segmentabruptly smaller andonly half the length of the penultimate. Head dark fulvous
brown, without a vertical tubercle; vertex relatively broad.
Mesonotal prrescutum dark fulvous with three, very illdefined,slightly more olive green stripes that are \:ery narrowly
margined with a reddish brorvn line, the median stripe further
split by a capillary vitta; remainderof mesonotumfulvous,scutellum slightly darker. Pleura dark fulvous brown, indistinctly
variegated with darker. Halteres relatively long, the knobs
darker brown. Legs with coxe and trochantersreddish brou'n;femora yellowish brown, the tips faintly darkened;tibire brorvn,
the tips very narrowly blackened; tibial spurs lacking; tarsi
broln, passing into brownish black; Iegs long; especially thetarsi, as shown by the measurementsgiven above. Wings (fig.
1) subhyaline,the stigma and cell Sc dark brorvn; wing baseand cell C more yellowish; wing apex vaguely more darkened;prearcular veins brown, the remaining veins black. Venation:
Rs of moderatelength, arcuatedi R z + e in alignment with the'Philip. Journ. Sci. g D 12 (191?)
108-11?

33,3

Alenander: Philippine Tipul;ida, IV

299

longer R, ; cell R, relatively small; cell M, broadly sessile;m-cu
at or immediately beyond the fork of M s + 4 ; Cu, extending
almost to wing margin; vein 2d A relatively short, cell 2d A
being very narrow.
Abdominal tergites dark brown, sternites obscure yellow;
fifth to eighth segmentsdarkened to form a subterminal ring;
hypopygium brown. Male hypopygium relatively small, thecomponent sclerites of ninth segment fused into a continuous
ring. Region of ninth tergite (fis. 2) terminating in two
slender points that are directed caudad, separated from one
another by a broad U-shaped notch which bears a small obtuse
median lobule at its base; the lateral lobes are blackenedandmicroscopicallyroughenedon their distal half. Basistyle separated from ninth sternite only by a curved ventral suture.
Ninth sternite extensive,membranousmedially and here producedinto a small, bilobed,fleshy structure.
LuzoN, Tayabas Province, Lucban, March, L926 (Franetsco
Riuera) ; a male. Named in honor of the collector, Mr. Francisco Rivera.
Tipula riuerai is very distinct from any of the speciesof thegenus known to me. The lack of tibial spurs, the peculiar
arrangement of setre on the antenna, and the sessilecell M,
are charactersthat ma)' be held as being sufficientto rvarranta new generic group for this fly.
Genus SCAMBONEURA Osten Sacken
Scumboneura OstsN SlcroN;
1 (1882) 95.

Berlin. Entomol. Zeitschr., 26, Heft

The genus Scamboneurawas proposedby Osten Sackenfor
the single speciesthen known to him, S. dotatcrOsten Sacken,
from the Philippines. Since that date additional specieshave
been described in and referred to this genus until now fivespeciesare recognized,to which number two more are addedin
the present paper. The great majority of the known forms arefrom Luzon.
Keg to the Philippine species of Scamboneura Osten Sacken.
1. Antennal flagellum bicolorous, the bases of the individual segments
black, the apices yellow...-..
-----...-.-.-.-..--2.
Antennal flagellum uniformly dark brown or black.---......-.-.-.-..-...-..-.....
3.
2. Mesonotal prascutum with three brown stripes, the margins of these
opaque, their centers shiny and with a slight metallic luster; legs
pale tawny, the tips of the femora and tibia darkened.
S. dotata Osten Sacken.

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The Philippine Journal of Seiettce$27

Mesonotal prascutum and scutum uniformly dark gray, without stripes;
Iegs obscure yellow, the tips of the femora and tibia not darkened.
S. psarophanes sp. noY.
B. General coloration of the head and thorax fulvous yellow to yellow,
S. unicolot Bezzi.
without stripes; pleura pale yellow--..--.-.---.Thoracic dorsum either plumbeous, without markings, or else yellow-""' 4'
with three black or dark brown stripes-----4. General coloration of the thoracic dorsum and pleura grayish plumbeous.
S. plumbea Alexander.
General coloration of the thoraeic dorsum yellow with three shiny black
stripes, the pleura uniformly light yellow.-.---..--.s. faceta sp. nov.

Besidesthe Philippine species,the only describedspeciesare
S. aittifrons (Walker) of Amboina, and S. quadrata de Meijereof Java. ScamboneuraplzwnbeaAlexander was omitted from
Edwards's tabulation of the Philippine species.
SCAMBONEURA PSAROPEANES BP. noY.

General coloration of head yellow, posterior portion gray witha median velvety black prolongation; prascutum and scutum
dark BI&y, without stripes; postnotal mediotergite whitish
yellow with a t-shaped darker marking ; pleura whitish EteY,
variegated with darker; abdominal segments obscure yellow,
the lateral margins of the tergites broadly blackened.
tr'etnnJe.-Length, 15.5 millimeters; wing, 11. Frontal proIongation of head light yellow, a trifle darker beneath; palpi
with the basal segrnentsyellow, the elongate terminal segment
infuscated, except at base. Antenna \Mith the scapal segments
yellow, the basal half of first segrnentblackened,the frons surrounding antennal fossa likewise blackened; flagellar segments
elongate, bicolorous, the basal trvo-thirds or more black, thedistal portion yello..v.the latter decreasingin amount and intensity distally, the terminal four or five segments being
uniformly darkened; flagellar segments with delicate, erectpubescenceand short, basal verticils. Anterior vertex rvhitish
yellow, the posterior orbits bright yellow; occipital region witha gray triangle, the anterior point extended cephalad into a
Iinear, velvety black line.
Pronotum dark, obscure yellow medially. Dorsum of themesonotalprescutum and scutum almost uniformly dark gray,rvith only the humeral triangles very slightly reddenedand theanteromedian portion of scutum yellowish; scutellum grayish
brown; postnotal mediotergite obscure whitish yellow with theposterior fourth dark Er&V, sending a median browu vitta
cephaladto the cephalicmargin of sclerite. Pleura whitish gray,3t,rAlenander: Philippine Tipulida, IV

295

variegat€d with dark gray on anepisternum, sternopleurite,
meron, and the postnotal pleuro-tergite; dorsopleural membranelight yellow. Halteres of moderate length only, pale brown,
the knobs dark brown. Legs with the coxe pale, the outerfaee of the fore coxa largely darkened, that of the posteriorcoxa a little darkened basally; trochanters, femora, and tibie
obscure yellow, the tarsi passing into darker. Wings subhyaline, base and cell Sc brownish yellow; stigrna small, darker
brown; wing veins black exceptthe basal and costal veins which
are bright brown. Venation: Tips of veins R, and R, both
atrophied; Rs preserved but pale; all forks of medial veins
deep; m-cu on M* at about one-third its length beyond forkof M.
Abdomen obscure yellow, the tergites broadly and conspicuously margined laterally with black. Ovipositor with thevalves relatively short and straight, reddish horn colored, thetips of the tergal valves obtusely rounded.
LuzoN, Laguna Province, Mount Maquiling (Baker); holotype, female; paratype, female.
SCAMBONEURA FACETA rp. nov.

General coloration of head obscure orange, the oceipitaltriangle darker; antennal flagellum black throughout; mesonotal prascutum obscure yellorv with three conspicuousblack
stripes; pleura light yellorv; legs largely brownish black; abdominal tergites obscureyellow, trivittate with brownish black,the median stripe very broad, interrupted at the caudal
margins of segments.
Male, length, about 12 millimeters; wing, L2. Female,length, about 15 millimeters; wing, 13. Frontal prolongationof head yellow, nasus black; palpi pale, the outer segment
passing into dark brown. Antenna of male elongate,if bent
backward extendingto about oppositebaseof secondabdominal
segment; scape obscure bror,vnishfulvous; flagellar segments
black. In the female, the antennre are much shorter. Headobscure dark orange, with a darker occipital triangle.
Pronotum narrow, obscureyellow, in the female, light orange
medially. Mesonotal prascutum obscure yellow rvith threeconspicuousblack stripes, the lateral stripes straight, the interspacesvery narrow; scutum obscure yellow, the lobes very
extensively blackened; scutellum shiny black, a little paler
caudally; postnotal mediotergite shiny brownish black, a little
paler on the lateral margins. Pleura light yellow, unmarked.

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The Philippine Journal of Science

t927

Halteres brownish black, only the extreme base of stem alittle paler. Legs with the coxre and trochanters yellow; femora brown, narrowly paler at base, the outer half or moreof the segmentsdarker brown; tibire and tarsi brownish black.
Wings subhyaline,stigma and subcostalcell brown; veins darkbrown to black. Venation: Rs as in the genus, short and simulating a crossvein;distal sectionsof veins R, and R" atrophied;m-cu on M* at about one-half its length beyond the fork of M.
Abdominal tergites obscure yellow, trivittate with brownish
black, the caudal margins of segments 2 to 7 narrowly andindistinctly obscureyellow; median stripe very broad, the sublateral stripes much less distinct; lateral margins of segments
pale; sternites light yellow, the outer segments darker; hypopygium small, dark brown. In the female the tergites arebrownish black, margined caudally with fulvous or brownish
orange, the lateral margins broadly of the same color; basal
sternites yellow, the outer segments duller in color; ovipositor
with the valves horn colored, the long tergal valves subacuteat tips.
In the paratype female the thoracic stripes are dark reddish
brown instead of black.
LuzoN, Tayabas Province, Alabat Island, September18 to 30,
1926 (Francisco Riaera) ; holotype, male; allotype, female;paratype, female.
Scambaneura faeeta is most closely allied to the Javanese
S. quadrata de Meijere, from which it differs especially in thedetails of coloration, notably of thorax, abdomen,and legs. The
general appearanceof the speciesis very much like certain
speciesof. Nephrotonta, and it is highly probable that the true
affinities of the genus lie rvith the Tipularia rather than with
the Dolichopezaria,despite the venation of the medial field of
the wing.
NESOPEZA CINCTITARSIS sp. nov. Plate l, figs. 3 and 4.

General coloration light cinnamon brown, antennal flagellum
black; legs dark brown, the tibial bases narrowly whitened;tarsi rvhite, the fore and middle basitarsi with the central half
blaekened or strongly infuscated; wings with a dusky tinge,
the small stigma darker brown ; cell 2d A narrow; male hypopygium large, ninth tergite conspicuouslydeveloped.
Male, length, about 11 millimeters; wing, 10 to 10.5. Female,length, about 11 millimeters; wing, 9. Frontal prolongationof head very short, light yellow; palpi yellow, passing into

Alerander: Philtppi'ne Tipukda, IV

33,8

297

brown. Antenna of male of moderate length only, if bent backward scarcely attaining base of abdomen; first scapal segment
pale brown; secondsegment yellow; flagellum black; antennaof female shorter, not attaining the wing root. Head lightcinnamon, frons passing into light yellow.
Mesonotal prrescutum and scutum cinnamon brown with theinterspacesa little darker; scutellum darker brown; postnotal
mediotergite 'darker brown, paler laterally. Pleura obscureyellow to brownish yellow, anepisternum and ventral portionsof sternopleurite and meron a little darker. Halteres elongate,
brownish black, the extreme base of stem a little paler. Legs
with coxe and trochanters pale yellow; femora dark brown,
paler basally; tibire dark brown, narrowly whitened basally;tarsi snowy white with about the central half of basitarsi infuscated; middle tarsi with the basal whitened portion more
obscured; terminal tarsal segments passing into light yellow
or whitish yellow. Wings (fig. 3) with a dusky tinge, thesmall stigma oval, dark brown; veins brownish black. Venation: Rs longer than the penultimate section of R, but usuallya little shorter than Rr*u, gently arcuated at origin; all medial
cells deep; m-cu close to fork of M; cell 2d A narrow.
Abdominal tergites dark brown with a transverse obscureorange ring just beyond base of tergites 3 to 5; subterminal
segments more uniformly blackened; male hypopygium conspicuouslyenlarged,basistyle brightened,tergite and dististyles
dark brown. Male hypopygium (fig. ) of very unusual form,
the ninth tergite (f) greatly producedcaudad and dorsad into
an elevated flattened plate that is deeply divided medially bya narrow split; lateral lobes relatively narrow, each shallowly
bifid on outer face near apex; ventral surface of the mesal
margin of each of these lobes at near midlength bearing aslender chitinized rod that is directed cephalad and slightly
laterad. Outer dististyle (o) an elongate,cylindrical lobe thatis provided with long erect setre. Inner dististyle (i) a small
flattened blade, the apex suddenly narrowed into a slender
point. Ovipositor with the valves chitinized, the tergal valves
gently upcurved at tips.
LuzoN, Tayabas Province, Lueban; at medium altitude on
Mount Banahao, May, L926; in dry forest, at base of tree, farfrom water (McGregor) ; holotype, male ; allotype, female;paratypes, both sexes.
Nesopezaci,nctitarsis is well distinguished by the remarkablemale hypopygium. It is plaeed in the genus in the broad usage
2,,8rtl-5

n8

The Philipptne Journal of Sciencetg27

of the name. The genus Nesopeza Alexander was proposed
for a ,group of Dolichopezaria that includes graei'Li'sde Meijere,
costa,IisBrunetti, and gerui,eulato
Alexander, all specieswith theradial sector very long, rectangularly bent, and spurred near
origin. To the genus a larger number of other species havebeen referred in which the sector is about as in the present
species,such forms being retained in Nesopezafot convenience
only.
LIMONIINEGERANOUYIAFLAVICOSTA

Brunetti.

Plate l, frg. 5.

Geronamgin flaui.costoBnuxnttt, Fauna Brit. India, Dipt. Nematoce ra (1 9 1 2 )3 8 9 -3 9 0 ,pI.8,fig.2 ( wing) ; pl. 11,fig.6 ( m esonotum) .

This interesting crane fly was described from a single imperfect female specimentaken from a light aboard a launch on
the Ganges Delta, India, August, 1909.
The male has not been described and the present specimen
is made the allotype.
Male.-Length (orcluding rostrum), about 6.5 millimeters ;wing, 6.7. Differs from the description of the female in thefollowing: Rostrum about as long as the combined head andthorax; black subterminal ring very narrow. Mesonotumshiny
ferruginous, without a dark pattern as described and figured
for the female. The wing pattern is very gaudy, in its general
pattern suggesting the pediciine genus Nipponuntyin ,lJexander
and the hexatomine genus Skuseomyia Alexander.
Male hypopygium (fig. 5) with the basistyle (b) relatively
small, the ventromesal lobe large, with long, conspicuoussetrc.
Ventral dististyle (u) very large, fleshy,the rostral prolongation
very short, sessile,provided with a single conspicuousspikelike
spine that is acute at tip. Dorsal dististyle (d,) relatively short,
only slightly curved, the tip suddenly narrowed into a slender
point. Gonapophyses(g) broadly flattened,apex slender,gently
curved to the acute tip, separatedfrom remainder of blade by anoval notch.
Luzox, Manila, March, L925 (McGregor) i allotype, male.
DICRANOMYIA
fg. 9.

(THRYPTICOMYIA) APICALIS(Wiedemann). Plate l, fis. 6; ptate 2,

Li,mnobia apicalis wrnourvraNN, Aussereur. zweifl. Insekt. 1 (1929)
551.

The crane fly described by wiedemann as apicalis has longremained in doubt. Through the kindness of Dr. Hans Zerny

33,3A[,erander:Philippi,neTipulida, IV

299

I was enabledto examine Wiedemann'stype and to settle finally
the identity of the fly, long suspected but never actually
proven. The fly belongsto the subgenusThrypticomyia Skuse.
There has been an unfortunate confusion regarding the correct
application of the specific names apicalis Wiedemann, saltens
Doleschall, and euneiformis de Meijere. The species determined by Brunetti a as saltens is now referred to Wical,is by
Edwards.o Edwards believesthat the name saltens Doleschall(saltans of authors) pertains to a species of the subgenus
Euglochirw Alexander rather than to TWpticomyin Skuse, afact that is borne out by Doleschall'sfigure, brief description,and measurements. De Meijere considered the name saltens
to refer to the subgenus Thrapticomyia. If Edwards's contention is correct, it is very probable that the name cuneiformis
de Meijere will fall in the synonymy of saltens (in the subgenus
Eugloclatnn). If de Meijere's original beliefs are well founded,
saltens will pertain to ThrypticomEin and will very possibly
be found to be a synonym of the present species. The chief
fact in the question that would tend to uphold de Meijere's contention lies in the curious dancing habit that Doleschall discussed for his saltens and which has been noted several times
by other collectors for different species of Thrypticomyia, butnot, to my knowledge, for species of Euglochino. Until Doleschall's material (if extant) can be examined, the problem will
still remain.
McGregor sent three genera of crane flies that he found aboutcacaotrees across the street from his house in Lucban, Tayabas
Province, Luzon, in May, L926. His interesting notes on theseflies are as follows:
The largest kind fConosin irt'orata'Wiedemann] rests on leaves with ahind leg extended on each side-mid and fore legs together, extended in
front-abdomen
elevated-no motion. This fly simulates certain spiders
and looks nothing like a fly.
The middle-sized kind fTrentepohlia trentepohlii,'Wiedemannl rests onleaves and dances rather slowly.
The smallest [D. (T.) api,calfs Wiedemann and D. (f .) arachnopltila sp.
nov.] rests the ends of the fore tarsi on a spider-web line and dances, threeor four flies sometimes resting close together on a line.

The last-mentioned habit has been discussed by earlier
authors, as Osten Sacken, Jacobson (through de Meijere),
Scott, Edwards, and others.
'Fauna Brit. Ind. Diptera, Nematocera 1 (1912) 373-374.
o Ree. Ind. Mus. 26 (1924) 295.

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The Philipptne Journal of Scimee

1927

The holotype of Wiedemann's species was redescribed by me
in l92L as follows:
The type has lost the apical half of the abdomen. Wings hyaline,
the cells beyond the level of the outer end of cell Jst M, distinctly infuscated; stigma elongate-oval, dark brown, clearly delimited; veins darkbrown, very distinct. Venation: ,Scr ending immediately beyond the origin of .Rs,,Scz sorre distance from its tip, Sc ' alone being a little shorter
than m-cz,' distal section of. R r pr€served, r being a little longer thannucu; distal section of B, atrophied beyond the distal margin of the stigma;
inner ends of cells B g &nd lst M z about on a level, lying more basad thancell R ri mrerr at near three-fourths the length of cell Ist M z; M in alignment with Msaa; distal section of Cur only a little longer than m-cu.
Ilind legs with the proximal third of the basitarsi blaekened; on middle
legs the darkening a little less extensivel fore legs lacking.

From these notes it can be seen that the absolutely criticalfeature, the structure of the male hypopygium, cannot be
described.
The Luzon material agrees very closely with the type, exceptthat Sc, ends shortly before the origin of Rs and m-cu is shorter
than Sc, alone.
Since there are rather numerous species of Thrypticomyinin the Oriental Region, it is deemed advisable to redescribe
and figure this species.
General appearanceas in the other members of the subgenus.
Verticils of the male antennre long and conspicuous. Mesonotal
prascutum very dark brown, somewhat shiny, the Iateral
margins of prrescutum somewhat paler. Pleura obscure testaceous yellow, the dorsopleural region darker. Legs black,tarsi snowy white, only the three terminal segments slightly
more yellowish white; basitarsi with about the proximal two
fifths darkened. Wings subhyaline, stigma large, oval, darkbrown. Wing tip strongly infumed, this including all of thecells from slightly beyond the cord outwardly. Venation as
discussedabove (fig. 9) : Cu, entirely lacking, as in the subgenus but not in Euglochina. Male hypopygium (fig. 6) withthe ventromesal lobe of basistyle (b) of moderate length andstoutness. Dorsal dististyle (d) strongly curved, the long tip
acutely pointed. Rostrum of the ventral dististyle (a) very
long and slender, the two spines widely separated from one
another, the distance between them approximately or nearly as
long as the length of a single spine; the more basal of thesespines arises from a short, hemispherical, enlarged base; ros-

,3,s

Alenond,er: Phi,l;i,ppineTipul;ida, IV

301

trum beyond the last spine from two to three times as longas the length of a single spine.
The species is most closely allied to D. (7.) fumida$cal;isAlexander, describedfrom North Queensland,and the two may
be merely geographical races. In the latter speciesthe distal
spur of R, is short, usually less than half the length of thevertical basal section; m-cu is usually far out toward the distal
end of cell lst M", though sometimesat about two-thirds thelength of the cell. The macrotrichia of the veins do not appear
so long or conspicuousas in apicalis. The male hypopygium(Plate 1, fig. 8) has the rostral spinesclosertogether,the morebasalarising from an elongatesubconicalbasethat is about twothirds as long as the spine itself ; the spinesare much less thantheir own leneth apart; apical beak of rostrum just beyond
spinesrelatively broacl,thencenarrowed strongly to tip.
(TERYPTICOMYH)DICRANOMYIA
ftg. 10.

ARACHNOPIIILA

ap. noy. Plete l, tg. 7; Plate 2,

Male.-Length, about 5 millimeters; wing, 5 to 5.5.
Generallysimilar to D. (T.) apicalis (Wiedemann),differingin the following:Thoracic pleura a little more variegated with brown, especially on the sternopleurite. Legs with the white more extensive,only the basalthird or lessof basitarsi being:blackened.
Wings (fig. 10) with tips slightly infumed, darkened back to
the level of the outer end of cell lst M-. Venation: Sc, ending
shortly before origin of Rs, Sc. some distance from its tip;
Rs angulatedand sometimesshort-spurredat origin; relatively
short, about as long as or a little longer than the distal spurof R, and in alig:nmentrvith it; distal sectionof R, equal to ora little longer than the basal section, provided with from six
to sevenmacrotrichire; m-cu at or before midlength of cell 1stM, and fully trvice its own length from tip of Cu,.
Male hypopygrumof the general type of apicalis. Basistylewith the ventromesa!Iobe short and stout. Dorsal dististylesiekle-shapedwith the apical spine relatively short. Ventral
dististyle with the rostrum shorter and stouter (fig. 7) withthe spinesrelatively short, placedclosetogether,the more basal
one from an enlargedtuberclethat is more than half the lengthof the spine, the more distal spine gently recurved.
Luzol, TayabasProvince,Lucban,on spider webs,May, 1926(}[cGregot'); holotype,male; paratype, male.

The Philippi,neJournol of Science

302

1921

The habits of this fly have been discussedunder the accountof the precedingspecies.
Genus TRENTEPOHLIA Bigot

The speciesof the genus in the Philippines may be separated
by the following key:'
Keg to Plvi,lippine spectes ot' Trentepohlin

Bigot.

1. Cell lst M, elosed, with three branches of media reaching margin(fig. 9). (Subgenus Mongonta Westwood.)--.-..2.
Cell lst Mz open by the atrophy of m and the two distal sections of
M'i only two branches of media reaching margin (figs. 12 to 16).
(Subgenus Trentepohli.a Bigot.)4.
-...-........
2. Tips of femora abruptly and conspicuously whitened; tibia white.
T. (M.) tenera Osten Sacken.
Femora brown, tips not whitened; tibia more or less infuscated--.- 3.
3. Femora uniformly infuscated, tips of all tibie white; (tips of midtibia
slishtly expanded and conspicuously fringed with long white setre).
T. (M.) pennipes Osten Sacken.
Femoral tips rather indistinctly darkened; tips of fore tibia broadly
darkened; (condition of midtibie not known).
T. (M.) luzonensis Edwards.
.... T. (T.) bakeri sp. nov.
4. Femora brown, tips abruptly whitened.-....-.
ofsegment or else darkened 5.
Tips of femora concolorous with remainder
5. Wings unmarked except for a narrow brown seam on Rri tips offemora and tibire conspicuously blackened.. T. (T.) mcgregori sp. nov.
Wings with a conspicuous brown pattern; femora not blaekened api- - - - - . - . . -6-.cally
6. Abdomen reddish, apex black; wings yellowish subhyaline, apex darkbrown; cord narrowly seamed with brown but not suffusing cell
. . . . . . . . ..... . T . ( T . ) t r e n t e p o h l i i ( W i e d e m a n n ) .
1st R,..............
Abdomen entirely black; a dark brorvn costal area at seetor and incell 1st R,, in addition to the darkened ape-'i..T. (T.) pictipennis Bezzi.
TRENTEPOIILIA(MONGOMA) TENERA Osten Sacken. Plate 2, fig. ll.

Mongorna tenera OsrpN Slcrprq;
1 (1882) 8e.

Berlin. Entom. Zeitschr. 26, Heft

LuzoN, Tayabas Province, Lucban, May, 1926; in small holes
in the shady bank of a creek; at base of large forest trees, farfrom water (McGregor) .
This common and widely distributed speciesis more variable
than are most speciesof the genus. Osten Sacken'stype (fromthe Philippines, collected by Semper) has the inner ends ofcells 2d M, and M, about on a line, but in the present series,
as rvell as in the material studied by Brunetti,G the inner end'Fauna Brit. Ind. Diptera, Nematocera (1912) 480-481.

rr, B

Alerander: Phi,Iippine Tipulida, IV

B0B

of cell M" lies far proximad of that of cell 2d M, (fig. 11), thebasal section of vein M,, being elongated and arcuated, much
longer than m. Brunetti places his pallidiaentris in thesynonymy of tenera but, unless his original description is very
erroneous,the identity of the two must be held in question.
TBENTEPOHLIA(MONGOMA) PENNIPES Osten Sacken.

Mongoma pennipes osroN(1887) 204.

slcxnN;

Berlin.

Entom.

Zeitschr. g1

LuzoN, Tayabas Province, Lucban, May, L1ZG (McGregor) ;Alabat Island, October 8, t926 (Francisco Riuera).
TRENTEPOIILIA(TRENTEPOHLIA) TRENTEPOELII(Wiedemann).

Limnobia trentepohJit WrnonrvrANN,Aussereur. zweifl. Insekt. 1 (1828)
551, pl. 6 b, fig. 12.

LuzoN, 'TayabasProvince, Lucban, in May, L9ZG(McGregor) ;numerous specimens,as discussedunder the account of Dicranomyia (ThrypticomEia) apicalis (Wiedemann).
There has been considerableconfusion concerning the identity of trentepohlii, but I have little doubt that the present
material is correctly determined. The specieshas a wide range
throughout the orient, extending eastward to northern Queensland, where it was describedas 7. (7.) meilio Alexander, whichname must be placed in the synonymy.
The speciesconsideredas being trente,poltlii,by Brunetti ? isgenerally similar, but has the u'ing pattern much paler andthe venation slightly different. I have material that was sent
to me by Brunetti. This speciesI had earlier described as ?.
(r.) doddi, from Melville Island, North Australia. Both ofthese closely allied species appear to have a very extensive
distribution in the Austro-Malayan Region.
TRENTEPOIILIA(TRENTEPOHLIA)MCGREGORI sp. nov. plate 2, ffs. tZ.

General coloration light orange yellow; head dark colored,
pruinose; legs yellow, tips of femora, basesand tips of tibie,
and the terminal tarsal segmentsblackened;wings light yellow;
vein R, narrowly seamedwith brown; vein R,,nearly transverse.
Female.-Length, 5.5 millimeters; wing, b.
Rostrum yellow; palpi pale at base,darker outwarcll5z. Antennre with the scapal segments obscure yellow; flagellum
broken. Head dark colored, heavily light gray pruinose.
'FaunaBrit. Ind. Diptera, Nematocera (1912) 482, pl. g, fig. 18, as
Mongomioides.

304

The Philipptne Journal of Science

General coloration of prothorax and mesothorax brightorange yellow, unmarked. Halteres relatively short, yellow, theknobs orange. Legs with the coxrc and trochanters yellow;
femora yellow, tips broadly and abruptly blackened;tibia yellow,
basesand apices conspicuouslyblackened,subequal in amount,
this about half as extensiveas the femoral tips; basitarsi yellow,
tips and remainder of tarsi dark brown. Wings (fig. 12) with alight yellow suffusion, the costal region more saturated; anarrow brown seam along vein Rr; membranehighly iridescent;veins yellow. Venation Sc, rather remote from tip of R,, thedistanee on costa about equal to Rs; Rs shorter than the basal
section of Rn*u; r on Rr*, just beyond midlenei"th;R, nearlytransverse, straight, relatively short.
Abdomen yellow. Ovipositor with the tergal valves horncolored, strongly upcurved.
LuzoN, Tayabas Province, Lucban, May, 1926 (McGregor) ;holotype, female.
This interesting crane fly is named in honor of Mr. Richard
C. McGregor, to whom I am vastly indebted for many rare
Tipulida from the Philippines. The speciesresembles?. Q.)
AIexnigroapi,calisBrunetti (India) and ?. Q.) septentrtoratls
ander (Japan) in the coloration of the legs. In all other regards
the present speciesis very distinct. Trentepohlia nigroapicalk(fig. 13) has the wings unusually long and narrow, the eells of
the radial field being correspondingly modified. Trentepohlia
septentri,onis (fig. L4) has the wing broader, almost as in thepresent species,but with vein R, long and oblique in position.
Trentepholin mcgregori (fie. 72) has vein R,, nearly perpendicular and cell R, relatively small.
TRENTEPOELIA(TBENTEPOELIA)

BAKERf ep. nov. Plate 2, 0s. 15.

General coloration reddish brown, mesonotum darker brown
medially; antenna black throughout; femora brown, tips
narrowly but abruptly whitened; fore tibie white; wings subhyaline, the oval stigma slightly darker; abdomen dark brown.
Fsmal,e.-Length, about 7 millimeters; wing, 5.
Rostrum brown, palpi concolorous. Antenna black throughout, the flagellar segments elongate-oval. Head dark brown.
Pronotum dark brown, paler laterally. Mesonotalprescutum
reddish brown, dark brown medially; remainder of mesonotum
dark brolvn, especially scutellum and postnotal mediotergite.
Pleura testaceous brown. Halteres relatively short, obseureyellow, the knobs a little darker. Legs with the coxrc and33,sAlerander: Pltili,ppi,ne Ti'pukda, IV

305

trochanters yellowish testaceous; femora brown, bases paler,tips narrowly but conspicuously whitened (about 0.5 millimeter) ; the only Ieg that is still attached is a fore leg; twoothers are detached but mounted with the type; fore tibia
white, those of the other legs very slightly more darkened;tarsi white, darkened outwardly. Wings (fig. 15) subhyaline,
the oval stigma slightly darker, poorly delimited; Cu and theposterior cord vaguely sullused with dusky; veins pale brown,
the costalregion abovestigma a little more yellowish. Venation:
Sc, remote from R, at margin, Sc, not far from tip of Sc,;Rs relatively short, nearly straight; distal ends of R, and r
relatively faint i R, a trifle longer than the seeond section of
Rr*r, oblique; basal section of Ra*u about equal to or a little
shorter than the fused Raar and M11r.
Abdomen dark broln, the genital segruent a little paler.
Ovipositor with the tergal valves very small, acutely pointed,
the sternal valves large.
LuzoN, Laguna Province, Mount Maquiling (Balcer'S; holotype, female.
This species is named in honor of the collector, Dr. C. F.
Baker, rvho has added very materially to our knowledge of thefauna and the flora of the Philippines. The fly is related to ?.
albogeniailataBrunetti (India), from rvhich it is distinguishedby the diagnostic features listed above.
CONOSIA

IRRORATA(Wiedemann).

Limnobw inorata
674.

Plate 2, 6g. 16.

WrcopuaNN, Aussereur. zweifl. Insekt. 1 (1828)

LuzoN, Tayabas Province, Lucban, May, L926 (McGregor) .
The occurrenceof this specieshas been discussedunder theaccottntof Dicranomyia (ThrUpticomyi,a)api,cali.s(Wiedemann).
The males of ircorata have the wings very greatly dilated,
Enderlein, the wing being widest
as in the genus Cludonocloztrs
opposite the second anal vein (fig. 16). In the female, thewings are narrower and normal in appearance.
The commonestrepresentative of Conosia throughout South
Africa is a small form, with the wings narrow in both sexes.
I had until now consideredthis as being i,rrorata, but the receipt of abundant material from several stations in the Orient
makes it clear that two distinct species have been confused
under this name.
The South African speciesis describedat this time.

306

The Phi,lippineJournal of Sei,ence

CONOSIA ANCUSTISSIUA!D. nov. Phto 2, fis. lT.

The wing is narrow in both sexesand of approximately equal
width for the entire central half of the length. fire irrorate
pattern is about the same in all three speciesof the g:enus,consisting of about four or five large costal blotches and abundantsmall irrorations at intervals along all the veins. The malehypopygium has the outer dististyle flattened, subcultriform,
broadest shortly before the tip, thence suddenly narrowed into
an acute apical point, the outer margin of the style with numerous subappressedspines. The g:onapophysesare extremely
Iong and slender.
Holotype, male, Pretoria, Transvaal, Januaty 26, 1919 (H.
K. Munro). Numerous other specimensfrom many parts of
Natal, Transvaal, and in Damaraland. M'fongosi, Zululand,
March, 1916 (W. E. Jones). Pietermaritzburg, Natal, January
5, 1911 (C. Fuller). Waterberg, Damaraland, Southwest CapeColony, February, L920 (R. W. Tucker) .

ILLUSTRATIONS
D, basiatyle; d, dorsal dististyle; g, gonapophysis; d, inner dististyle; o, outerll,egend:
dististyle; B, radius; r, radial crossvein ; Sc, subcosta; C, ninth tergite; o, ventral dististyle.
Yenationalterminology
used, Comstock-Needham-Tlllycrd.
Ifypopygial
terminology
used,
Crampton.lPr^a.rs 1
Frc.

L. Ti&ln riaerai sp. nov., wing.
2. firyla, ri,aerai sp. nov., ninth tergite, male hypopygium.
3. Nesopeza cinctitarsis sp. nov., wing.
4. Nesopeza chtntitarsrs sp. nov., male hypopygium.
5. Geranuntyin fl,avinoefo Brunetti; male hypopygium.
6. Dinranomyia (Thrgpticomyial api.calis (Wiedemann); male hypopygium.
(Th,wpticorwi,a, arachnnph,ib sp. nov.; rostral proDiqanomuia
7.
longation of ventral dististyle of rnale hypopygium.
8. Dicranotnyin (Thryptiaornui.e, fttmidnpicalis Alexanderl rostralprolongation of ventral dististyle of male hypopygium.
Pr,lrpFtc.

2

9. DioranomAia (Tlwgpticomyin) apicalis (Wiedemann); wing.
L0. Disanomryia (Thrypticorryin)a,rachnoph,ilo sp. nov., wing.
ll. Trentapohlin (Mongomal terura Osten Sacken, wing.
12. Trentepohlia (Trentepoh,lin) mngregori sp. nov., wing.
13. Trentepoh.Iia (Trentepolulio\ nigroapinalis Brunetti, wing.
14. Trentepohlin (Trentepohlial septentrionie Alerander, wing.
L5. Trentepohlia, (Trentepohlin) bokeri sp. nov., wing.
16. Cornsia irrorata (\Miedemann), wing, male.
L7. Conosia angustissima sp. nov., wing, male.
807

lPErLrP. Jounx. Scr., 83, No. 3.

ALExaNDEa:PntLrrrrxs Trrurrpp, IV.liI
PLATE I.

Ar,sxA.xoER: Pnnrpprrrtn Trrulroc,

IV.l

I

.r

Rl

lPErLrP. Jounw. Scr., 83, No. 3.

III
PLATE 2.