STUDIES ON THE CRANB-FLIES OF MEXICO
Penr II
Order Diptera, SuperfamilyTipuloid.ea

Cnenr,Es P. Ar.ExaNnnn

REPRINTEDFROMANNALS oF aHE ENrollor.ocrcer,
Vo l. XIX,

Socretv

No . z.

Cor.uMBUs,OEro,
June, 1926.

oF AMERTCA,

19261

Alexander: Crane-Flies of Mexico

159

STI]DIES ON THE CRANE-FLIES OF MEXICO.
PART II.
(Order Diptera, SuperfamilyTipuloidea).
Cn,rnrBs P. AlBx.rxor,:n,Amherst, Massachusctts.

The present paper is a continuation of the studies outlined
in part I of this seriesof papers (Annals Ent. Soc. America, 1E:
341 - 362; 1925). As before, the species discussedwere all
included in very large and important series of crane-flies thatwere collected and sent to me for dctcrmination by my friend,
Dr. Alfons Dampf, State Entomologist of Mexico. A few
brief notes on certain additional Mexican Biocenosis (M.8.)
and Mexican Fauna (M. F ) numbers are given hereu'ith:
Hac'iend,a San Franc'isco, near C6rdoba: Swept from thewalls of a passage way, cut three to four mctcrs deep acrossan uncultivated area covered with woods and bushes, theperpendicular walls of the pass overgrown with sma1l ferns
and other plants. M.F. 335, 490, etc.
Barranca of the R'io San Antonio, near C6rd,oba, altitude
870 meters, swept from ferns and other plants beneath overhanging trees. The Rio consists of small clear ponds connected by faint water-courses with very stony beds. f socoenosisto NI.8.73 (seePart I, p.3a1). M. B. 102.
Cerro de Hongos: Southern prolongation of the Desierto
mountain range; swept from grass in open pine forest at analtitude of from 3500 to 3700 mcters. M. B. 117.
Ajusco: The highest and most easterly part of the mountain chain south of Mexico City, of which the Desierto forms
the western part. Dense Ab'ies forest, altitude 3500 meters,
swept from weeds along the course of a small brooklet.Rhaphid,olab'is(Plectromyia) tergata sp.n. In pine forests between 3500 and 4100 metets.-T'ipula oreomyzoitlessp.n.
I express my deepest thanks to Dr. Dampf for the privilegeof studying this splendid seriesof Mexican Tipulidae, the typesof which are retained in my collection through the kindnessof the collector.
*Contribution from the Entomological Laboratory of the Massachusetts
Agricultural College,
r58

Limonia Meigen.
n.
sp.
fissilis
Limonia
General coloration brown, the pleura with a conspicuous longitudinal black stripe; rostrum and antennae black throughout; wings witha dusky tinge, ihe stigma darker; Sc and Rs long; male hypopygium
complicated in structure.
Male. Length about 4.8 mm;rving, 5 mm.
Rostrum and pa$i b1ack. Antennae black, the flagellar segments
oval. Head dark brorvn.
\llesonotal praescutum clark brorvn 1ateral1y and more broadly somediallv behind, thc remainder of the disk more reddish brown; scutal
lobes dirk brolvn, the median arca and the scutellum more testaceous;postnotum shiny brorvn. Pleura light brown with a broad black longiiuclinal stripe extcnding from the cervical sclerites to the base of theabclonten, passing bencath the root of the halteres; ventral.pleurites
paler. Halteres d-ark brown, the base of the stem.narrowly paler. Legs
fore coxae slightly infuscated; remaining coxae. and the*ith tt"yellor,vish teitaceous; femora dark . brown,. -the bases
trochanterspaler; tibiae'and tarsi clarker brown. \Vings (FiS' 1) with a dusky
i;t g", th" stigma ova1, darker brorvn; slightly paler- brown clouds atorigiin of Rs, tip of Sc and as \iague seams.along the cord and outer
en,Tof ce117il ,[4r;veins dark brown. Venation: Sc long, Sc1extending
to oirposite two-thilds the length of fts,.Sc2 at its tip;-Rs 1ong,.gently
arcu'aied; ce11-1sl M2 large, ividened distally; m-cu close to the forkol M.
Abdor-ninal tergitcs dark brorvn, thc sternites vaguely bicolorous.,
the bases being ialer than the tips. N{a1e h1'popygium- (Fig. 1.2)
w i rh t he nint h'r eigit e ( t ) near ly it r aight acr osst he caudal m ar gin,with a large patch of setae on eithet side. Basistyle-(b) relatively
1arge, the ,r"ttt.o-t-tt"t"1 lobe very large and complicated in structure;
on'it.s outer sicle near base bearing a smai1, densely hairy, finger-like
1obu1e;lobe dilatecl outwarclly, the innet margin b9f9r9 apex dilated
into a ledge that is microscopicalll' setulose: apex of lobe terminating
in a smali- conical papil1a. Dorsal dististyle (dd) boomerang-shaped'
ventral dististyle (vd) very complex, the rostral prolongation large.
Phallosome (p) forming a large central mass.
Holotype, c/, Tres Valles, Yeta Cttz,
in hotel (A. M. Dampf).

Septembet 26, t924;

Rhipidia Meigen
Rhipidia (Rhipidia) multifida sp. n.
Male with ten branched flage11arsegments; flagel1um bicolorous;
thorax anteriorly obscure ye11ow,the posterior sclerites of the nolutlt
and pleura darkened; all iarsi infuscated; wings. tingcd. with c1usl<y,with-three additional smaller dark spots and inconsllicttortsst'ltttts
rvitlr tlrt'
along the cotd; m-cu before thc forl< of ,lf; rtralgh1,1l6p.v*giurrr't ',lr ot t t 'ltt 't t t 'r 't 'rtlt t 'r t l't t t ir llt 't r t '1ll.
drl rsil dist isi. , r '1r . lor; tr r! rl r 1sr lt 'nr 1t sr

1e261160

Annals Entomological Society of America[Vo1' XIX'

Mole. Length about 'i-rmm.;wing, 5'8 mm'segments
Rostrum and palpi black. Antennae with the scapal
and the.branches
segments
flagel1ar.
theofenlargements
black;-basa1
whitish; tenbi;;t; the elongafe pedicel of the individual segments
three times
about
branchlo-ngest
the."g-.tttt,
flageliar[;;;A"dproawithsegment
lirst flagcllar_conicallylfr" i""efh of-the segrie.t; 'Headfi";;:;i";i;;;1";?i;".

clarkbrownishgrav' e191Ja) at-anteriot I'ertex being reduced to a narrorv strrp'
most-'-" touching. thebrown. Mesonoial praescutum obscure. vellow,
dark
Fro"oi"--'near midlength;*ith . bro*n median stripe that is slightly interrupted
dark brown;
entirelyscutumin.coloi;
paler
ancl
;tr;;
i;ili
yellow'
obscure"";_o:..not,Pleurabrown'.""i"ii"nt and poitnotum slightll'paiersclerites caudad
alland(1-rteiopleuiite,meron;;;";;;tkti; 't,;frr'"utfi.
iteurites
obscure yellorv, the l<nobs infuscated.
ft.tt"t".
;;id'""r)yellowi middle and hjnd coxae dark
otr*"ore
fore
Less tvith thc"n*n"
ntorvnish teslaceous;. femora brown' -the bases;t;;;,'";;;'iut,t!ttand tarsi bror'vn, the terminal tarsal segments
tibiae;;;r"'h.;rrul"t;r,vagt (Fig.2) with a slrong dusky tinge, the stigma
ir"*"i.rr-lir"t.'ti.o*",'t16 outer end darker than the remainder
aurt".
;;i,'.tfihtlyr; smal1 brown clouds at origin of Rs andas a *""r,',rra-'.pfruri'"g"tdusky seams along cord and outervagl1eancl,larrow
tip of^Sc; veiypale;veins dark brown' Venation:end. of ce11-/sl Mr; ptiatcttlar cE11s
justmidlength of Rs' Scz at its
before.S, t"f"i-"1"o.'rttoit,'St,""Ji"gshortly before the fork
m-cu
origin;Ltiio, nl f""s, weakly titg"f"t.8
lacking.
M; h appirentiyoi"' 'cta"rii""ithe caudal
i"rgit", <1aik bro*n; sternites obscure vg1_1oy.,
black' Maie
brownishwithnarrowly'ringed
it..g-"ttheofmarcinswith 1 d"ggu-shapedii;;3ffiil'iFil: "theii, i;'"i"g tt'" ninih.tergite
iateral iobes conspicuous, fringed with powerfullobe"6i"fi-"&"au1Ly,=";no". bristles. Basistvle tb) relativeiy sma1l,the ventro-mcsal
na rro w c dout war dll' , p ro v i d c d rv i th n trme ro u s tu b e r cl estnatD earconDorsal distispicuous selae, some 5i *rri"t are exceedinglv elongate.
a.ri
ght angl e'
a
boul
a
t
te
n
t
mi
rl
l
e
n
g
th
(dd)
t
t
.
"
d
"
.,
and
lonq
stvl e"tspine.
blackened
acure,
long,ainto,nrrowed;,d;; i;;;;;."."aJ*iv
the. rostrumbasistyle,.
thethanlargerfleshy,V;;i;i-dtt.t'f"t.,ai(its tip broken
spi""s two, iire.outer one much stouterslender, setiform.
more
spine
"l;ft;;;;";rit
thelength."?i"tJ_inner(s)"ot-"etfai"),conspicuously blackened' Aedeagus (a) at apex
Contpopitytd
split into two lobes.
(A' M' Dampf);
Holotype, d, C6rdoba, September 30, L924

M . F. No.38 5 .
Rhipidia (Arhipidia) agglomerata sp' n'Male. Length 4.5-4.7 mm; wing, 6'2-6'3 mm'
Guatemala' to
Closely a11iedto R. (,4.) multigtrttota,Alexander, of
World
key
to
author's
ofihewhich ipdcies it runs b1.'-""trrlh"-{:X'of Rhihittia-(Eiull. Brooklyn Ent. Soc., 8: 7-8; i912)- The*ii,,lii,
a s fo l l tx v s :lr t ' t : ic s ' < lif f crs
1,n 's,'r rs1.

Alexander: Crane-Fl'iesof Mexico

161

Head brou,'nish gray. Xtlesonotal praescutum rich reddish brorvn
i n fr ont , t he t hr ce br or vnishblack st r ipcs indicat ed behind, cr ossing
the suture and suffusing thc scutal lcibes;scutellum pruinose; postnotal
mediotergitc baso-media11vbrownish black. Ventral hrown pleuralstripe broader and darker, suffusing the basal halr.es of the middle
and posterior coxae. Legs with the femora obscure }.eliow, and tips alittle clearer yel1ow but thc segments without distinct darker subtetminal rings. Wings with the brorvn dots in the celis of the wings
rel a t ively f eu'ir r num her . lending t o m assint o ir r egularblot ches. Venation: Scr ending opposite or beyoncl midlength of Rs.
Ilolotype, d, C6rdoba, Rio San Antonio, after sunset,
S ept em ber 30, 1924 ( A. N{ Dam pf ) ; M . F. 385.
Par at opot ypes, 2 & d.
The hypopygial details in these two species are very similar
to one another.
The same general type is found not only inother species of Arhipidia,
but in the typical subgenus (R.
Iong ispina Alexander . ) O ne of t he par at ypes shows a r em ar kabie deformity of the male hypopygium, the dististyles of theright side being duplicated; the basistyle of this side is larger
than usua1. The gonapophyses and aedeagus likewise appear
to be asymmetrically repeated.
Ilelius St. Fargeau.
quadrifidus
Heliussp. n.
Allied to H. albitarsis (O.S.); size sma1l (wing, d,4 mm.)l rvings
with a dusky tinge, veins Ra15 and L[yr2 ver)r extensively fused, obliterating r-m and giving the appearance of four veins issuing fromcell lst AIz.
M ole. Lengt h, J m m ; wing, 4 m m .
Rostrum and palpi black. Antennae black. Head black.
Thorax dark brown, the mesopleura paler brown. Halteres darkbrown. Le{rs with the coxae and trochanters dark brown; femora
and tibiae black, the extreme bases of the former somewhat paler;
tarsi black, the end of the basitarsus and all the remaining tarsal segments except the last pure white, clothed with snorvy white setae.
Wings (Fig. 3) with a dusky 6ings, the oval stigma somewhat darker;
veins dark brown. Venation: Sc1 ending opposite three-fourths thc
length of Rs, Sc2 at its tip; R,snearly straight; R213sinuous; Ra*5rrcry
extensively fused with Mp"2to form the cephalic margin of ce111st tr[2,giving the appearance of four veins is-"uing from the ce11;basal dcflcction of Ra15short; ce117sl M2vet! Targe;m-cu shortly beyond midlcnglhof the ceil.
Abdomen black, the basal sternites paler; hypopygium black.
Ho:l,olype, d, At light on train between La Favorita lrrrl
Tapachula, St at e of Chiapas, June 74, 7\ 125 ( A. M . l) nr r Jr f ) ;M. F. No. 55S.

162

Annals Entomolog'icalSoc'ietyof Amer'i'ca[Vo1. XIX'

Orimarga Osten Sacken.
Orimarga dampfi sp. n.
General coloration gray, the mesonotal praescutum with.a medianbrown stripe; thoracic pteura with a conspicuous silvery_white stripellegs pale brown; wings with a pale grayish tinge, the base whitish;
ndmacrorichiae on distal sectionof vein ,41,r.
Male. Length, 7.5-8 mm.; wing, 5.5 mm.
Female. Length about 8.5 mm.; wing,- 6-6.2 mm'
Rostrum and"palpi brownish b1ack. Antennae dark brown, pruinose; flage11arsegmentsglobular to short-ova1. Head light gray'
Mesonotum dark grJy, the praescutum with a median brown stripe,
the extreme lateral margns bt ttre sclerite pa1e. Pleura brownish
gray, with a conspicuouJsilvery while- stripe extending from behindih""for" co"a, acrois the dorsal edge of the sternopleurite, ending above
the posterior coxa. Halteres short, pale yeliow..- Leg-swith the coxae
brownish testaceous, the trochzLntersobscure yellowish .brown; femora
pale brown, the remainder of the legs somewhat darker brorvn, the tips
bt th" individual segments narrorvly darker, the terminal tarsal segments more uniformly darkened. Wings (trig. a) with a very pale
gray tinge, the prearcirlar region whitish; veins-pale brown.. Macrolrict iae 6tr veins rather numelous, there being about 20 on the distal
section of vein M3, abortt 24 on the distal section of R:r+si no macfotrichiae on Ma. Venation: Sc ending opposite one-third the lengthof Rs, Scz at its extreme tip; Rs gently angulated at origin; r abottto.r" ut d one-half times its length from the tip of Rr and about itslength
bevond the fork of Rs,. r rlear midlength of the basal section of .Ra15;
M"^ a IitLIe more than one-half the distal section of Mi m-cu obTique,near midlength of .Rs.
Abdomen elongate, dark grayish brown'
Ilolotype, &,Yera Cruz, November 10, 1923 (A. M. Dampf)'
AIIotoPotYPe, I .
Pa ra topot y pes , S 0 &Q.
This interesting species is named in honor of the collector,
Dr. Alfons Dampf, to whom we are indebted for much of ourknowledge of the crane-flies of Mexico.
Erioptera Meigen.
Erioptera (Mesocyphona) venustipes sp. n.
Belongs to the d.ulcis grottp; head variegated;^fore and middle*ith three broid biack rings; posterior femora black withfemoraye11owsubterminal ring; tips of the tibiae narrowly brownish
o ,1o.ro*"ibh
black.
4.8 mm.
Iicmale. Length about 3.5 mm.;"ving, with the scapal segments
I{ostrunt and.-palpi b1ack. Antennae.
lrt.,rrvrrisltlrlac1<,thb basal flage11arsegments pale yel1ow, the terminal

1e261

Alexander: Crane-Fl'iesof Mex'ico

r63

segments passing into pale brown. Head dark brown, the orbits
narrowly but conspicuouslyyellowish.
Lateral pretergites narrowly pale sulphur-yellow. Mesonotal praescutum brownish gray with two conspicuous dark brown stripes thatare subequal in width to the space separating them, these dark stripes
continued caudad onto the scutal lobes; lateral margins of the praescutum passing into light brown, the extreme lateral edge dark-brown;
pseudosutural lor.'eae pale; scutellum with the caudal margin broadly
ye1low. Pleura dark brown and silvery, altcrnated, the clark brown
including the dorsal half of the dorso-pleural region; a conspicuous
stripe exlending from and including the fore coxa, passing above thehalferes to the postnotum; a narrow ventral stripe extending from thesternopleurite passing above the middle coxa and below the halteresto the abdom-en; a-conspicuous sulphur-ye11owstripe occupies theventral portion of the dorso-p1eural membrane, extending from above
the fore coxa to the wing-root; the silvery stripes remaining are two
in number, the dorsal one narrowed in front' widening behind andincluding the base of the halteres; the ventral silvery stripe. includes
the broad meron and the katepisternum. Halteres white, the knobs
infuscated. Legs with the fore coxae dark brown; middle and hindcoxae pale, the apices of each with a dark spot; troehanters paJel iorg
and middle femora ye11orv,each t'ith three conspicuous brownish black
annuli that are mu-ch more extensive than the ground-color, the firstpre-medial, the second, a little wider, post-media1,_thethird-broadest,
subapical; posterior femora entirely brorvnish.black except for a conspicuous yellowish rvhite ring far before the tip and the extreme apex
oi the segment rvhich is pale; tibiae yellowish rn'hite' the tips narrowly
brorn'nish-b1ack; two basal segments of tarsi yellowish white, the tips
and the remainder of the tarsal segments dark brorvn. \Arings witha brown suffusion, mole intense apically and in the radial fieid; basal
half of cel1sC and Sc more yellowish; prearcular cel1sdark; disk withtr,venty large rvhite spots arranged about as in the other members ofalong the cord and fork
the ditcis itotp, including spots at h, Sc2,.Sc1,of tr't+zi a series of nine marginal spots between veins Rr and lst A;
the remaining spots are located at arculus, origin of Rs, on Rs just before the fork; on ,4f1-'2shortly beyond m; and two areas along vein
1st A, one near the base, the second near midlength of the.vein; anal
angle'restrictedly pale; veins brown, r'n the pale areas whitish; vein 1llargely black, R1 and Cur largely yellow. Venation: Ce11-lsl M2 opcr'
by- the atrophy of the outer deflection of Mv
Abdominal tergites brorvn, the lateral margins of the segments
narrolr'ly blackened; sternites more yeilowish brown, with numerous
brou'n setigerous punctures.
Ifolotype, I , Coscomatepec, altitude 1580 meters, flying
over the surface of the Rio Tlacuapan, a swift-flowing strcrLm
with sandy bottom and very clear water, November 29, 1\124(A . M . Dam pf ) ; M . F. No. 463'

L64

Annals Entomological Soc'ietyof America[Vol. XIX,

Pa rat opot y P e, I ; M . F . N o .4 6 1 .
E. venustipes belongs to the d,ulc'is group of the subgenus,
which now includes dulc'is Osten Sacken, eisen'i Alexander,
needham'i Alexander, and the present species.

1e261

Alexand,er: Crane-Fl,iesof Mex,ico

165

Gonomyia remota departs from the normal characters oftlre subcinerea grovp in the ye11ow antennal scape and short
subcosta. The detached Scz is unique in the genus, as far as
known.
Gonomyia (Lipophleps) strigilis sp. n.

Gonomyia Ntleigen.
Gonomyia (Gonomyia) remota sp. n.
Belongs to the subcinerea group; antennal scape largely yellorv;
rostrum black; head ye11owwith a dark spot on vertex; pleura yel1ow
with two dark brown stripes; wings with Sc short, Sc1ending far before
the origin of Rs, Scz lying detached, some distance distad of Sc1; abdominal tergites dark brown, with conspicuous caudo-1ateral yellorv
triansles.
Sir? Length about 3.8 mm.;rn'ing,4.6 mm.
Rostmm and palpi b1ack. Antennal scape conspicuously yellow
with r.ague darker rnarkings; flagellum dark brown; flage1lar segments
oval, passinginto elorrgate-oval. Head light yel1ow,the center oi thevertex with a dark.brown spot.
Pronotum light sulphur ye1low. Laterai pretergites similar. Nlesonotal praescutum brown with three nearly confluent darker brown
stripes, the surface weakly pruinose; lateral margins of sclerite and thehumeral triangle obscure brownish yeilow; pseudosutural foveae elongate, reddish brown; scutum obscure yellow, the lobes extensively
dark brown, confluent across the suture with the lateral praescutal
stripes; scutellum testaceous yellow; postnotal mediotergite ye11ow,
darkened posteriorly. Pleura yellow, with trvo conspicuous darkbrown stripes, the dorsal one extending from the cervical sclerites,
passing above the root of the halteres and including the posterior
margin of the postnotum; ventral stripe including the stenopleurite
and meron. Halteres slender, dark-colored. Legs with the coxae
and trochanters light yellow; remainder of legs dark brown. Wings(Fig. 5) with a faint brown tinge, the stigma slightly darkerl veins
darker brown. Venation: Sc very short for a member of the subcinerea
group, Sc1 ending a distance before the origin of Rs that is nearly as
long as the latter; the detached ,Sczlies considerably beyond the end of
Scr at near two-fifths the distance between Scr and the origin of Rs; Rsstrongly arcuated, about equal to the slightly less arcuated R2a31basal
deflection of Ra15 short; m-cu immediately beyond the fork of ,44.
Abdominal tergites dark brown; tergites two to four with conspicuous light ye11owcaudo-lateral triangles; abdomen broken beyond
the fourth segment; sternites light ye11ow.
Holotype, Sex? , Barranca of the Rio San Antonio, nearC6rdoba, aititude 870 meters, November 20, 1924 (A. M.
Da mp f); M . B . No. 1 0 2 .

Xfale. Length about 3.2 mm.; wing, 3.8 mm.
. Belongs to the cinerea group; most closely allied to G. (L.) cinerea(Doane), differing chiefly in the structure of the male hypopygium.
Rostrum and palpi black. Scapal segments of antennae'brorn'nish
ye1low, biackened beneath. Head ye11ow,the center of the vertex
infuscated. I,Iesonotal praescutum brown, the lateral margins paler;pseudosuturai foveae conspicuous, reddish brou'n; posterior sclerites
of mesonotum weakly pruinose; a capiliary median vitta extending
from the suture to the abdomen. Pleura with the whitish lonsitudinai
stripe fu11ytwice as wide as the brown stripes that enclose it. Wings
with the costal margin china-while as in the group. Male hypopygium
lFio
l z t \/ , \ , 1, 1
+r h'L- .,' ^.-, , t r al or out cr dist ist lle
idt a'long blade,ihe- apicat-" ' 1 L
half infuscatgd, ol the mesal face at near the basal third with a small,
subappressedblack spine, the margins of the style not otherwise armed;
intermediate dististyle a short blackened spine, narrowed to the acute
apex;_dorsal or inner style a pale fleshy 1obe, the caudal margin produced caudad into a blackened spine.
Holotype, d, Between Coscomatepec and Tomatlan,
altitude neady 1500 meters, swept from the train, from grassand bushes, November 30, 7924 (A. M. Dampf); IVI. F. No. 466.
The chief differences between the present species and G.
c,inerea are the small spine on the ventral dististyle and theshort, heavily blackened intermediate dististyle.
Gonomyia (Lipophleps) latistyla sp. n.
Relongs to the pleuralis group; size sma1l (wing, d, less than 3.5mm.); male hypopirgium with the dististyle appearing as a flattencd
blade, the lateral apical angle produced into a small spinous 1obe, thc
apex of the style with numerous setae and a conical spine.
Male. Length about 3.5 mm. l wing, 3.2-3.3 mm.
_ Rostrum lnd palpi black. Antennal scape obscure orange; flagcllum black. Head pale orange-yellow.
Praescutum pale brown, the lateral margins broadly light yel1ow,
margined internally by a narrow dark brown line; scutal-lobes lxrlc
trrovnl scutellum darker brownl postnotal mediotergite brownislr
yellow, indistinctly .variegated medially with darker. Pleura l)rowrr,
wlth a conspicuous white longitudinal stripe extending from thc lxrscsof the fore coxae, across the anepisternum and ventral nrargin o[ llrc
pteropleurite to the base of the abdomen; this stripc is lrrargirrt'tllrotlr
dorsally and ventrally by a slightly narrowcr rla.rl<lrrorvn lint,; stt,r.rr,r-166

oleurite and meron pa1e. Halteres pale brown, the knobs a iittle iniuicated. Legs with the coxae pa1e, the apical_half of the.fore coxaa"tf. fto*"; f"emorabrown, the iips a little darker;.remainder.of legs
dark brown. Witrgr tinged with duskiz, the oval stig-ma dark brown;it" t"giotr bbfore aid beyond the stigma paler; base of wing atd costal
iegion-slightly more yellorvish; veins pale, the cord darker' Venation:
Sishort,'Sc1 entling a distance before the origin of Rs that is about
iwo-thirds the lengt"hof the latter vein; Rzar short and arcuated, bendthe sti"gma;cel1,Isl M2 strongly narrowed at proximal end;
lng"io""dbefore fork of ,41.
shortlym-cu
Abdomen dark brown. Male hypopgium (Fig' 15) of the generaltvp" 6i C- (L.) pteurali.r (Wi11.); chitinlzed spines at apex, of basistvle(6) shorter,'the'longer slender,.curved, the tiP acute,.the base notchiiinizecl,; tlie shorter spine is very sma11and pale throughoutiil'."ilv
birt#t t" (d) a bioad flattened blade, the outer apical angle produced
into a imaii spit ous 1obe,the terminal surface of the style with.numeru conspicuous conical spine at the inner apical angle;
ous setae"trdofstyie with a pale flange.
marginouter
Holotype, 8,(A.M.D am pf ) .

Vera

Cruz,

Parato'potype, &; paratype,
January 18, 1921 (A. Alfaro).

Mexico,
&,

10,

t923

Costa

Rica,

NovemberTempisque,

Gonomyia (Lipophleps) spinistyla sp. n.
Male. Length about 3 mm.;wing, 3 mm'-n"tong,(L.). lotislyla
Lo ihe pleuralisgroup; most closelyallieclto G.
hypofygl
um'
mal
e
th
e
o
l
sp .n ., dif ler ing c s pec i a l l yi n th c s tru c tu re.-''M"-.onotum rjct-l t"ddi.h brown, the scutal lobes darker; medianwith a narrow dusky. r-itta that .",.t:lqt, caudadt"gio" L} .""tum yellow"area
the scuteilum, the latter broadly iight yeiiowofmedianthe
on-tou"H"a; portnotum dark at base, the.remainder more yellowish. Pleural
Abclomen with the disk of the segments pa1e,.ttio"-ii""-r.-white.
broadly darker. Male hypopYSlum (Fig' .16) with
ifr"'frt"tui"*rgins(b) relatively short and stout, c1.lindrical,
basistyie.
ofrpine;pi";a;h;ifr" l';p t"aaenly narrow,la i"to an acute polnt;-ventral "fl".th{ lobe ofbasistvle moderitelv stout; the second chitinized spine of the basistyle
ir apiut"ntly lacking in the species.. Dististyle (d) pale, the outer*;t6 Jn acute black spine, the apex oj the style beyond-rhir-*;;"ir"dO-"rlinnarro*ed and provided with _erecLsetae)the Lwo apical ones
much'larger. Phallosome asymmetrical, as figured'
Holotype, d, At light on train, between La Favorita andand Tapachula, State o1 Chiapas, June 14, L925 (A' M' Dampf);
M. F. No. 558.

1e261

[Vo1' XIX,

Annals Entomolog'icalSociety of America

I

Alexand.er: Crane-Fl'ies oJ Mex'ico

t67

Gonomyia (Lipophleps) bifida sp. n.
Male. Length about 4 mm.;wing, 3.6 mm'
Female. Length about 4.5 mm.; wing, 4 mm.
Belongs to thi pleuralls group; most closely allied to G. (L') recurvata
A1ex.
Rostrum and palpi black. Basai antennal segments orange, thiscoloration including the first segment of flagellum; remainder of .organ
dark brown. Head light yellow, the center of the vertex broadly infuscated.
Pronotum sulphur-yellorv. N'Iesonotal praescutum rich reddish
brown, the lateral margin narrowly light yellow,- the-co1or marginedinternally by a narrow brownish black line that ends before the cephalicend of the sclerite which remains of a rich reddish brown; a mediandark stripe that is better indicated posteriodl'; scutal_lobes reddish
brown, the meciian area broadly pale with a capi1lary.brown medianvitta that passes caudad onto the testaceous ye11owscutellum; postnotum pale, dark at the cephalic portion, the caudal end slightly darkened. Pleura light ye11owwith two narrow bror'vnish black stripes,
the space between light ye11owand a little wider; dorso-pleural area
more chestnut brown.
Abdominal tergites obscure 1'e11ow,the caudal and lateral marginsof the segments darker, each segment with a natrow'-dark, -longitudinal
median stripe that is more expanded anteriorly. Male hypopygiuP(Fig. 17) rvifh the outer angle of the basistyle (b) prolonged into a gently
curved acicular spine; outer fleshy lobe moderately stout. Dististl'le(d) profoundty bifid, the lateral arm a slender, gcntly cutv-ed spine,ih" jt t-r"t arm pa1er, the apex terminating in two powerful bristles,
the surface of this arm with numerous smaller setae; at extreme basechitinized peg or spine. Gonapophyses (g) with theof dististyle a tiny-acicular,
divergent; ventral. gonapophyses (g) I'ery
blades eiongate,
short, aboui- as'broad as long, the apex terminating in two sma11,divergent spines.
Ilolotype, d, Tospan, near C6rdoba, swept from wa1ls of
a way, December 75, 1925 (A. M. Dampf); M. F. No. 490.
Attotopotype, d, with the type and pinned on same pin.
Paratype, 9, San tr'rancisco, near C6rdoba, December 10,

1924(A. M. Dampf); M. F. a74.
Gonomyia (Lipophleps) haploa sp. n.
If ale. Length about 3.3. mm.; wing, 3.7 mm'
Belongs to the ,nanca group; resembling,G. (L.).inermis AIex',
differing dspecially in the structure of the male h1'popy.gi"P. .
Rosirum obscure orange; palpi b1ack. Antennae black throughout,the verticils very elongate. Head biack, the center of the vertex rvitl'r
an orange-yellow spot.
Pronotum dark. Mesonotal praescutum shiny, rather dark lrrowtt,
the pretergites conspicuously pale whitish yellow, this coloral,i()r1('x-168

Annals Entomological Soc'ietyof America

1e261[Vo1. XIX,

169

legs; tibiae brorvnish ye1lot, the tips passing into black; basal half ofbasitarsi pa1e. the rcmainder of the 1,arsipassing into black. WilSs(trig. 6) hvalinc, the sl,igma lacking: veins dark brorvn. i\'Iacrotrichiaeof veins rllatir.ely sparse, there bcing on11tthree on the distal section
scattered along the vein. Venation: .Sc1ending shortlyu,'ir1e11'
of -Re13,
be-vondthc origin of Rs; r almost at the tip of 1i1,thelatter being aboub
one-thircl thc r.cin; ce1l Rs at wing-malgin nearly trvice as wide as cell
2nd R1;nt a Lrifle shorter than the outer deflection of M:.
Abr'lomerr brown, the sternites paler; hypopygium dark brolvn,
Nlalcr lrylrcrpvgium of the general structute ol 7'. pobulatoria Alexancler,
bul lhe short distist-v1esof difierent shape, the outer arm not extendecl
into a long point.

tending to the rving-root; pseudosutural foveae shiny bron'n; scutellum
broad11.yellow, rvith a dark brorvn median basal spot. Pleura duskl-,
sparsely pruinose, rvith a broad white longitudinal stripe extending
from the forc coxa, beneath the root of the halteres, to the abdomer.r.
Halteres with the knobs dark. Legs rvith the coxae and trochantersbrorvnish testaceous; remainder of legs darker brown. Wings u'ith agrayish tinge, the stigma lacking; r-eins dark brou'n.
Abdominal tergites dark brorvn. Male hypopl.gium (FiS. 18)
with the basistylc (b) long and slcnder, narron'ed distal1v, the surface
with setae, inciuding a longitudinal series of stouter setae alon;1 themesal face. Dististylc (d) single, simple, entirely pa1e, appearing asa direct apical prolongation of the basistyle, the apex r,r'ith two polverfulsetae, the surface rvith about six smaller setae, the outer surface rvithabundant erect setulac. Phallosome (p) pale, as1'mmetrical, the longest element (aedeagus,a) long and slender, projecting far beyond theo th er elem ent sof th e h y p o p l g i u tn , th e g o n a p ophysessmal l and on11'
weakly chitinized.

I lolot ype, &, Yer a Cr uz, Novem ber 10, 1923 ( A. M . Dam pf ) ;along margins of a ditch.
T. minuta is the smallest species of the genus yet described.
Its nearest a1ly is T. pabulatoria -Piexander, of Guatemala.

Ilolotype, d, Barranca of the Rio San Anbonio, near C6rdoba, altitude 870 meters, November 20, 1921(A. M. Dampf);
M. B . No. 102.
The members of tlne manca group in which the male hypopygium has but a single, entirely fleshy dististyle arei manca
O.5., ,inertnes Alex., phoroctenia Alex. and the present form.
The species are readily separated from one another by the.details of structure of the phallosome.

Pilaria Sintenis.

Teucholabis Osten Sacken.
Teucholabis minuta sp. n.
General coloration yellow, the praescutum with three reddish bron'nstripes, the scutal lobes darker; pleura yellow with a darker dorsal
longitudinal stripe; knobs of halteres dark brown; wings h]'aline' the:stigma lacking; Sc short; ceIT2nd R1 at wing-margin much natrot'ver
than ce1l,Re.
Male. Length about 3 mm.;wing, 3.1 mm.
Rostrum and palpi dark, the latter slender. Antennae dark brown
throughout, the flage11arsegmentswith relatively long verticils. Headlight
" gray.
Pionotum ye11ow. Mesonotal praescutum sulphur-yel1ow, lvith
three reddish brown stripes, the median stripe darker anteriorly, thelateral stripes crossing the suture and suffusing the scutal lobes; scutellum yellow; postnotum with the mediotergite infuscate, the posterior
third brighter. Pleura whitish ye11orv, including the pleurotergite,
the dorsai sclerites suffused rvith brown to produce a longitudinal
stripe. Haiteres testaceous, the knobs dark blown. - Legs with thecoxie and trochanters yellow; femora obscure ye11ow,the tips conspicruously though narrowl.v blackened, the amount subequal on al1 theAlexonder: Crane-Fl'iesof Mexico,l,4

Pilaria rubella sp. n.
Antennae of male clongate; general coloration of thorax shiny reddish brorvn;wings with a strong brownish tinge, the oval stigma slightly
darker brown.
Male. Length, 7-7.5 mm.;lving,8 8.8 mm.
Rostrum ferruginous, the terminal segments of the palpi dark brown.
Antennae with the scapal segments orange-brown; flagellum brownish
b1ack, the segments rvith a dense erect pubescence,in addition.to theelongate verticils. Head rvith the center of the vertex dark brown,
the color passinginto reddish brown on the genae.
Thorax shiny reddish brown to ferruginous, without evident darker
markings, only- the scutal lobes somewhat darker, the pale medianarea of the scutum continued cephalad acrossthe suture onto the praescutum. Halteres brown, the knobs darker. Legs with the coxae
and trochanters reddish brown; femora )'e1lowish brown with darker
brown setae; tibiae brown, the tips and the tarsi passing into brownish
b1ack. Wings (Fig. 7) with a strong brown tinge, the oval stigma
setiferous and slightly darker brown; veins dark brown. Venation:
Sc1 ending about opposite tu'o-thirds the length of Rs, Scz at its tip;r immediately beyond the fork of ,R213and a little more than twice its
length from the tip of R1,'R2 relatively short, arched near midlength;
inner ends of ce1lsRr, Ru and -/sl M2 in oblique alignment, ceIT1st tr'[2lying more basad than the other elements; m-cu just before to iust
midlength of ce1l,1slM* ceTl MlTonget than its petioie' be1.6n6-Abdominai'tergites
dark brown, indistinctly variegated with palcr;
sternites obscure brownish yel1ow; hypopygium reddish yellow.
Ilolotype,d, C6rdoba, Hacienda
900 meters; swept from perpendicular

San tr'rancisco, altitrrtlt:
walls of a passagc wily

t7O

Annals Entomological Soc'ietyof Amer'ica[Vol. XIX,

that was covered with smal1 ferns and other vegetation; September 13,1924 (A. M. Dampf); M. F. No. 335.
ParatopotyPe,d.
P. rubella differs from P. tenu'ipes (Say) in the colorationof the thorax and the strongly infumed wings.
Atarba Osten Sacken.
Atarba mexicana sp. n.
General coloration shiny brownish yel1ow; antennae bicolorous,
the flagellar segments b1ack, the apices of the segments narrowly paleclosed;abdomen
yel1ow;wings with veins Sc and Rs elongate;cell 7sl ,11,6
bicoiorous, the basal portion of the segments b1ack, the apical portion
broadly brownish ye11orv.
M ale. Length about ,1.5mm.; n'ing, 6 mm.
Rostrum obscure brorvnish yellorv; palpi b1ack. Antennae elongate,
if bent backward extending about to the third abdominal segment; scapal
segments dark brorvn: fiagellar scgments black, thc apex of the segments light ye11ow,that of the first segment including about one-sixth
the total length, of the second and third segments including approximately one-third the length; on the succeedingflage11arsegments theamount of 1tellorv becomes gradually iess, the terminal four or fivesegments uniformly dark. Head dark bror'lrr.
N{esonotum shiny brownish ye11ow, without distinct markings.
Pleura somewhat more yellowish, the dorso-pleurai region darker brown.
Halteres obscure ye11ow,the knobs dark brown. Legs r'vith the coxae
and trochanters yellorvish testaceous; femora yellow' the tips very
narrowly and vaguely darkened; tibiae obscure yellow; tarsi passing
i n to dar k br own. Wi n g s rF i g . 8 ) w i th a s tro n g brorvni shyel l ow ti nge,
the costal region somewhat 6righter ye11ow,e.speciallyin the stigmal
region; veins-brorvn, those in the costal region paler, especially Sc.
Venation: Sc long, Sc1 ending be1'ond four-fifths the length of Rs, Scg
some distance from its tip; Rs strongll. angulated and weakly spunedat origin; cel1 Rr at wing-inargin nearly three times as \^'ide as ce11R3;
cell 1sl M2 closed,rectangular, m-ut beyond midJength.
Abdomen bicolorous,-the basal half or less of the segments blackened, the apical portion broadly bror,vnishyellow; hypopygirrm-obscureyellow; sternites somewhat less distinctly bicolorous' lvJ'alehypopygium r,vith the aedeagus very elongate, decurved.
Holotype, d. C6rdoba, near the Rio San Antonio,
8 , 1 9 24 ( A . M . Da mp f); M.F . N o . 4 0 1 .

1e261

Holotype, 9, Heights of Penna Blanca, a mountain chain
north of C6rdoba, white limestone cliffs in dense forest, swept
from sunny places, altitude 1200 meters, December 75, 1924

(A . M . D a m p f ) ; M . B . 1 1 0 .
A Key to the American Species of Atarba.
1.
2.

3.
4.

5.

6.

7.

8.

Female. Length about 5 mm.;wing, 5.6 mm.
Generally similar to A. mexicana sp.n., differing especially in thevcnation, trichiation and the coloration of the antennae and abdomen'

T7T

Antennae broken near midlength, if complete extending approximately to the base of the abdomen; flage11arsegments uniformly paiebrown. Headbrown.
Pleura s1ightl1.pruinose. Wings (Fig. 9) with cel1 1st l[t open by
the atrophy of the outer deflection of Ms; Sc1shorter than in mericatt.a.
Macrotrichiae of wing-veins less abundant than in mericana, the basesof the longitudinal veins largely without trichiae, this glabrous area
including the basal fourth of Sc, the basal three-fourths of ,1,f and thebasal two-thirds of Czr. In meri,cana,the trichiae on Sc extend basadalmost to arculus; occupy the distal four-fifths of M;are much more
numerous on the Anal veins, especially .|sl ,4.
Abdominal tergites dark brown, at most vaguely bicolorous; basal
sternites obscure ye11ow. Genital segrnent obscure ye1low, the valvesof the ovipositor horn-yel1orv.

October

Atarba apeda sp. n.

Alexand.er: Crane-Fl,iesof Mexico

9.

2.........
S c 1ong, S c r endi ng bc y ond mi dl ength ol the l ong R s .....
S c s hort, ,S c rendi ng approx i matel y oppos i te the ori gi n of the s hort R .r.... 3
conlincd to distal fourth.
Ce1l /sl M2 openi vein .41 with macrotrichie(Mex i c o)....a/erta s p. n.
Celi -/sl Mz closed; vein M with macrotrichie almost to base. (N{exico),nlexlcana sp. n,
Femora with a brown subapical ring; wings with a vague darker marking......4
al ongthec ord...
Femora without a subapical ring; wings unmarked, except for the stigma. . . 5
Mesonotum shiny yellow, the praescutum r,vith a brorvn median stripe,
each scutal lobe with two dark brown spots; wings with a narrolv brolvn
seam along cord and outer end of ceII 1st M2. (Paraguay), punctiscuta AIex.
Nlesonotum yellowish, unmarked; wings rvith a dusky spot on anterior cord.
(C ol ombi a). .. . ..c ol ,umbi ana A l ex .
(Colombia),
Antennal flage11um uniformly brownish black.
brunneicornis A1ex.
Antennal flage11um bicolorous; (in megaphall,us very indistinctly so, only
6........
the i nc i s ures betrv een the s egments bei ng pal er)
Flagel1ar segmerits dark brorvn, with both ends very narrowly and indistinctly pale; male hypopygium with the acdaegus greatly dilated.
(Brazii).megaphallus Alex.
Flagellar segments with one end.broadly pale; male hypopygium with the.......7
aedeagus of moderate s i z e on1y .........
Flagellar segments dark brown or black with the apicai portion con(British Guiana)
spicuously yellow.
. . . .cinrlicornis Alex.
Flagellar scgments dark brown or black with the basal portion con.....8s pi c uous l y y el l ow .
(Paraguay) . . . . . .Jiebrigi AIex.
Femora with the tips narrowly blackened.
.....9Femorauni forml y y el l ow i s h.
Abdominal tergites du11 ye11ow; size larger (wing, L over 5.5 mm.)
(E as tern N orth A meri c a)....
...pi c ti c orni s O. S .
Abdominal tergites dark brown; size sma11 (wing, L under 5 mm.). (Pcru),. _aartcornt's Alt:x.

L72

Annals Entomolog'icalSoc'ietyof America[Vol. XIX,

Eriocera \{accluart.
Eriocera pallidipes sp. n.
G ener al c olo ra ti o n b r, )\\' n , th e 1 ,1 c u ral argcl y bl ack; ti ps of theti biae and all bu t th c a l ri c c so l th e tc rs i o b s curcbrorvni shrvhi te; rvi rgs
with a pale browrr tinge, the costal regir.rnand ntost oi the longitudinalveins clouded r'vith darker brorvn; Rs iong, ce11,111lacking.
tr[ale. Length, 1! mrn.; u.ing, 10-10..1mm.
Femalc. Length, 1-t mm.;rving, 10.2 10.5mm.
Rostrum and palpi b1ack. Antennae sltort, t'Lre scapal segments
obscure brorvnish ye11ow;flage11umb1ack, the first segment paler atbase; flagel1ar segments elongate, gradually shortening outwardly.
Head dark plumbeous; vcrtical tubercle small.
N{esonotum broln, the pracscutum behind, the scutal lobes and thescutellum more plumbeous. Pleura largcly black, the coloration including t he pos tn o ta l p l e u ro te rg i l e :s fe rn o rll euri te and meron pal i ng
into shiny brownish ye1low. Halteres relatively short, the knobs
dark brown. Legs with the coxae shiny obscure yeliow, the extreme
bases of the fore and middle coxae blackened; trochanters ye11ow;
femora obscure ye11ow,the tips narrowly blackened; tibiae brown, theti ps nar r owll- u ' h i te n e d ; ta rs i o b s c u rc b ro w ni sh w hi te; thc termi nal
segments passing into brown. Wings (l'ig. tU) rvith a pale brownish
tinge, the stigma scarcely indicated, the base, costal region and seams
along most of the veins darker brorvn; veins dark brown. Venation:,Sc1ending about opposite trvo-thirds -R2a3,Sc2 just beyond the forkof Rs,' Rs very long, from two to two and one-half times Rs13;r fromone and one-half to twice its or,vnlength from the tip of R1 and about
twice its length beyond the fork of R2*3; ceITMr lacking; ce71.
lst I[zrectangular; m-cu about its own length beyond the fork of ,4{.
Abdomen brown, with a more or less distinct median dark vittaon the tergites; pleural region darker brown. Male h1'popygium relatively smal1. Ovipositor rvith the valves long and slender, acute,
blackened basally, the tips paler.
Ilolotype, d, Vera Cruz, November 10, 1923 (A. M. Dampf).

Allotopotype, g .
Paratopotypes, 6J g i l, Laboratorio Howard B. Cross,
JaIy, 1924.
E. palli.d.iprsstil1 further eliminates the characters available
for separating Penthoptera frorn Eriocera.
RhaphidolabisOsten Sacken.
Rhaphidolabis (Plectromyia) tergata sp. n.
Cell & sessile;vein M very delicate,in alignmentwith M4; fn-cu
shortiy beyond the fork of M; male hypopygium with the tergite conspicuous,the lateral anglesproduced into earlike lobes; mesal faceof llasistylcwith a conspicuous
pencilof long yel1owsetae.

1e261

Alexand,er: Crane-Fl'iesof Mexico17;imn.; rving, 5.3 mm.
Male. Length about ,1..1
Rostrum and palprib1ack. Antennae relatively short, black throughout; flagellal segmct.ttsova1. Head dark gray.
X{csbnotum discolorecl;from slight patches of the normal coloratiorlremaining, probably bronnish gray, the pracscutum presurnabll' strilled.
Pleura brorvnish grar.. Ifalteles short, brorvn, the base of the stenr
narrot'1.v pa1e. Legs l'ith the coxae light gral' pminose; trochantersbrorvnish .vcilorv; rcmainder of legs bro',vn, the femorai bases pa1er,
the ternrinal tarsal segmentspassing into dark brorvn. \Vings (Fig. 11)
rn'ith a pzrle bror,rnish tinge; stigma vaguel-v delimited, slightly darkcr
bror,ln; an i1l-definec1brorvn cloud on the anterior cord; veins darker
bronnr. Veins ,11 and -lsl / rrery slender and pa1e. Venation: Scr
extending to be-vond one-third the length of R213,^Sc2far before thc
origin of -Rs,the distance being nearly one-half longer than that portionof Sc1bel,ond the origin of Rs; Rs gently arcuated; R213clongate; R2
about equal to R1aR2;R; a 1itt1eshorter than Rr,3, gentlv arcuated:basal scction of Ra15very short, about equal to the basal deflectionof R; ; At in alignm er r tl'it h, 4/ r . : , bot h ver y pa1e,t he lat t cr beconr inx
stronger beyond the fork of L[+i m-ut tmmediately beyond the forkof ,rtf; cel1M aTongand narrou., gently rvidened outwardly.
brorvn.
Abdomen dark brorn'n, the hypopygium more ye11or,r'ish
Male hypopygium (Pig. lt)) rvith the ninth tergite (t) conspicuous,
the caudolateral angles produced into ear-1ikeiobes, the surface of thesclerite densely set with setiferous tubercles. Basist-vle (b) produced
into a slender lateral lobe that is spinous at apex, with a sma1l sessile
dorsal lobe that bears a lesser number of spinulae. Mesal face ofbasistyle rvith a conspicuous pencil of long ye11owsetae; siightly morc
basad of this, an oval cushion denselv set with smal1erect setae. Dististyle (d) broad at base, the apex narrowed, the style extended be1'on,1
the apical lobe of the basistyle, the surface with a few large and numerous microscopic setulae. Interbases (i) slender, more or less footshaped. Aedeagus (a) broad.
IIolotype, i, Ajusco, altitude( A. M . Dam pf ) ; M . F'. No. 549.

3500 meters, May

10, 1925

Tipula Linnaeus.
Tipula bellardiana sp. n.
General coloration grav; antennae with the basal flage11arsegments
bicolorous, black, with the tips paler; head with a natrow brown vitt:r.;praescutum with three clearer gral' stripes that are narrowll' margincrl
with brou'n, the median stripe further split b]'a capillary dark vitta;
femora with a vague .vellowish subterminal ring; rvings variegal,r:rl
whitish subhyaline and brorvn, including a complete lunate band lrt'yond the cord; tip of Re atrophied; abdomen obscure ye11orv,the sr.tlrterminal sesments blackened.
M ale. Lengt h, 10- 11m m . ; wing, 13- 14m m .

174

Annals Entornolog'icalSoc,ietyof Amer,ica[Vol. XIX,

. Frontal prolongation of head brown, the nasusslcnder; palpi darker
brown. Antennae relatively short, if benL backward extending aborlt
to the root of the haltere; scape and first flagellat s"gr.,"ti-y"tto*;
intermediate-flagel1ar segments more or less b'icolorouJ,-ni""r., *itr,
the apex of the segment narrowly and so'retimes indistinctly brownish
yeliow, more extensive on the basal segment, the amount becoming
more restricted outwardly, the terminal flagellar segments uniform
scgmenls wirh rhe basal enlar!e-.nt ,ind"i"i"-in ,ir",
F,11"!illg:ilar
the \/ertlcrls conspicuous. Hea,d gray, the vertex with a median dark
stflpe.
.Mesonotal praescu.tuq Vellowish gray with three clearer graystripes, the median stripe further s.ptit me<iiatty and marginea r.t"iirrywith brown to form..three_approximated bro#n
poorly indicated, usually ap_pearing "itt;;ti";;ri*stripes
gr"y area margined:?T",i-:r". onto\4r1thdark brown, the. color crossing the suture"
the scutal lobes;
scuLtallobes gray, each with the antero-latera1 marking just describecl
and a much larger posterior spot that nearly covers thelobe; ,".rt"ltrrgray, the median area somewha! d-1r\er; poslnotum dark brown, heavily
pruinose. Pleura dark gray,^indistinctiy variegated with paier,
thisincluding the dorsal portibn of the sternopleurite-andnfcroor'tn"
pleurite and meron.j dorso-prcuralmembrane light -oit
yeliow: Hii;;;,obscure yellow, the knobs dark brown. Legs with the fore and middle
coxae dark gral',_the. posterior coxae_paler;"trochanters reddish yellow;
femora brown,.the tips blackened,thls coior preccc,led
bV a vai,ue ob_
scure y.ellow ring of approximatel_vequal width; tibiae light i.u*rr,
gradually passing into_.browniqh bl-ack,-the tarsi concolorou?;-alltarsi
longer. than tibiae. \tr/inqs.whitish s_ubhyaline,variegated *idh-bru*rr,
the stigma darkest; cel1 c infuscated except near the outer end where
there.rs a clear.spot; cell sc yellow, alternating with four dark brownoL sllghilv greater extent; wing__apcx
broadly darkcned; all.lTlnF.
runaLe Drown mark before the. cord, extending from the stigma to thc
wing-m_argin at c,:t; an irregular pale brorvn iross-band u".lo., cells R,
M p-l,A,,near two-fifths the length of the former; bases of cells R
the chief pale areas thus remaining include a cur.red?dband"M-darkened;that completely clolsei the.wing b91'o.ndthe Jord,;trci"ai"g tlr"
basesof cells R2, R3 ind R5 and aimost arf"of
I J" a', i"i-iir;V_shape{ band with the origin of fts in the fork""fis"of the y, the-l.o'airt"tpale blotches in cel1
,sc folmlng. the arms; conspicuous p.t"- rr"". inbasesof cells R and M and including much of the Anal celld.- venation:
Distal section of Rz entirely atrophied; petiole of cerrMt about one-nau
longer than m.
Basal abdominal tergites obscure ye11ow,the subterminal segments
pas_singinto dark brown, the segmenti with the iateral tr."tg"and indistinctly blackened, the mid-dorsal region with a -black"r?o*ty
strip!
that becomesbroader behind; .",lqll margins o'f the segmentsnarrowly
ringed n'ith pale; hypopygium reddish brdwn; sternites-brown;strvetto#
with an interrupted brown median vitta, the brown of the inai'viauat
scgnrents broadest behind, the color interrupted at the base of eachscgrrcnt. \,Iale hypopvgium with the ninfh tergite large, re.ddish,

1e261

Alexand.er: Crane-Fl,ies of Mex,ico

175

the caudal margin rvith a deep V-shaped notch, the lateral ears thus
formed being very conspicuous, over-shadowing the dististyles. Basistyles very sma1l. Ninth sternite with a U-shaped median notch.
Eighth sternite unarmed.
Holotype, 8, Chapingo, luly
23, 1924 (A. M. Dampf);
F. No. 254.
Par at opot yPes, &, JuJy 14, 1924, M . F. No. 235; &, July 21,
1924, M . F. No. 249; &, J, aly 25, 7924, NI . F. No. 2bb: C,
A u gust 8, 1921, \ 4. F. No. 285i d, August 13, 1924, lVI . F.
N o. 291.
M.

This interesting species of Tipula is named in honor
Professor Luigi Be11ardi, pioneer student of the Diptera
Mexico.

ofof

Tipula oreomyzoides sp. n.
Antennae 1_6-segmented,elongate, the flagel1ar segments b1ack,
strongly incised beneath; general coloration of head and thorax grrLy,
the praescutum with four. darker gray stripes; wings tinged with brown,
longitudinally _vittate _with whitish subhyaline, this color including
cel1slsl M z and R5; abdomen yellow, the tergites tri-vittate with brown,
the sternites with a median interrupted brown stripe.
fuIale. Length, 72.5-14 mm.; wing, 15-18.5 mm.; antenna, 7.5-8mm.
Frontal prolongation of head obscure yellow, the sides with a narrow dark brown line; nasus conspicuous; palpi dark brown. Antennae
16-segmented,elongate, if bent backward extending to opposite midlength of the abdomen; scape and basal portion of the fiist flage11ar
segrhent obscure .ye11ow,the remainder of the organ black; flagellar
segments deeply incised beneath. Head light gray, the center of thevertex a little darker gray.
_ Mesonotal. praescutunr light gray with four darker gray stripes,
the narrower intermediate pair briefly connected at the cipialic end;
scutal lobes gray, each with two darker gray spots; scutellum pale ashy
gray; postnotum light gral'. Pleura gray, vaguely variegated wjth
darker, especially on the sternopleurite and meron; postnotal pleurotergite whitish gray; dorso-pleural membrane obscure ).e11orv. Halterespale, the knobs dark brown. Legs rvith the coxae obscure yellow, thc
fore coxae.slightly pruinose; trochanters )'e11ow;femora brown, paler
at base, _the tips passing into darker broum; tibiae brown, the- tips
broadly darker brown; tarsi brownish black; all tarsi longer than tibiae.
Wings wfth a faint brownish tinge, the costal region more yellorvish
brown; disk of wings with a iongitudinal vittate pattern of whitish subh1.ali're, this including the broad bases of the Anal ce11s;the narrowbase and broader apex of ce77Cu; bases of ce11sR and Mjapical thir-rl
of ce1lM,'al1 of cel1s1st Mz and,R5,and the baseof ce1l,41r,.
loirgilrrrlirriLl
vcins and the cord narrowly seamed lvith somewhat darl<r:r-lrrorvrr.

L76

Annals Entomological Society of America[Vo1. XIX,

In.a flw cases,.thewings.aresti1l paler than described,with the basal
cel1salmost entirely whitish subhyaline;veins brolvn, p"ier in the costal
region.;obliterativeareasincludingthe cord and outei sectionof Ma+t.
venation: Distal section-ofR2pale but complete;cel1.1srM2ftregulirry
pentagonal;m andpetioleof ce11
,lfr subequal.
of the tergitesbroadlybr,rvn,. Abdomenyellow,.thelareral_margins
the-line becoming_widerbehind, or the sixth to e;gntn segmentsincluding the caudal margins of the segments;an intemrpTed dorsomedianstripe, the vitta broadly broken-at the fosterior maigin of eachsegment,less_so.at the base;hypopygiumyellowish,the ap]cesof theinner di,qti51y1es
hear.ily blackened;siernites-yellow, ihe caudal marginsof the basal segments_narrowl)'
infuscated,-on sternitessix to eight
more extensively so. Male hypopygium very similar in structure tothat of T. rrtoctezumae
Alexancer. Ninttr tergite with the lateral lobes
less conspicuouslytruncated at apr'ces,the s?na1ler
median notch lessabrupt. Eighth sternitervith a smallmedianliguliform1obe.
Flolotype, f , Ajusco, altitude 3800 meters, in pine forest,
May 10, 1925(A. M. Dampf).
Paratopotypes, 2 & d, altitude 8500-4100 meters, in pine
forest, May 10, 1925. Paratypes, Z e g, Cerro de Hongos,
altitude 3500-3700 meters, swept from grass in open pine
forest,April 5, 1925 (A. M. Dampf);M. n. No. 117.
T. oreomyzo,ides
ts obviously closely allied to T. moctezu.n'ae
Alex. and r. azleca Alex., which are likewise Mexican soecies
from the higher mountains near Nlexico City. The presenceof sixteen antennal segments tn T. yreomyzoidesindicates thatthe range of antennal segments in Tipula, as now known, lies
between 13 and 16. The supposed genus Oreomyza pokorny
is based upon three species from the higher Alps in which themales have 15-segmented antennae, the femalei l4-segmented
antennae. The genus Oreomyza has no validity and must
be placed in the synonymy of T,ipula.

1e261

Alexand,er: Crane-Flies of Mexico

EXPLANATION177

OF PLATES.

Symbols: a-aedeagus; b-basistyle; d-dististyle; dd-dorsal dististyle; g-gonapophysis; i-interbasal process; p-phallosome; t-9th tergite; vd-ventral
dististyle.
Prern XI.
Fig. 1. Wing of Limoniaf,ssilis sp. n.
Fig. 2. Wing of Rhipidilr (Rhipidia) multif.da sp. n.
Fig. 3. Wing of Hel'ius quadrifidus sp. n,
Fig. 4. Wing of Orimargatlampf,sp. n.
Fig. 5. Wing of Gonomyia(Gonomyia)remota sp. n.
Fig. 6. Wing of Teucholabisminuta sp. n.
Fig. 7. Wing of Pilaria rubel.l'osp, n.
Fig. 8. Wing of Atarba mexicanosp. n.
Fig. 9. Wing of Atarba apertrasp, n.
sp. n.
Fig. 10. Wing of Eriocera Pal'l.id.ipes
Fig. 11. Wing of Rhaphidolabis(Plectromlia) tergatasp. n.
ParB XII.
Fig. 12. Male hypopygirm of.Limonia f.ssilis sp. n.
Fig. 13. Male hypopygium ol Rhipdd.a(RhdPida)mul.tifiila sp. n.
Fig. 14. Male hypopyghtm ol Gonomyia (Lipophleps) strigilis sp. n.
Fig. 15. Male hypopygium of G. (L.) latistyla sp. n.
Fig. 16. Male hypdpygium of C. (L.) spinistyla sp. n.
Fig. 17. Male hypopygium of G. (L.) bifida sp. n.
Fig. 18. Male hypopygium of G. (L.) haploa sp. n.
(Plectromyia)tergatasp. n.
Fig. 19. Male hypopygium of Rhaphid'olabis

AxNns E. S. Al,'ol. XIN. Preru Xi.

Fig.4

Fig,6

-t

ANN,rls E. S, A,

nlKl

s'",9

Fig.18C. P. Alcrandq

179