From the Proceedings ol the Linnean societg ,of New Boufit, wales,
Vol. lai, Parts 3-4, 1936.

THE

DIPTERAOF THE
F,TMILYTERRITORYTIPULIDAE.

OF' NEWPARI

GUINEA.

IV.

II.'

By crrenr,usP. Ar,nxennrn,Massachusettsstate college, Amherst, Mass.,u.s.A.
(Comntunicated,by Frank H. Taylor, F.R.^E.S.,
F.Z.S.)(Seventeen Text-figures.)[Read 26th August, 1996.]
The materials included in this instalment are chiefly from the mountains
of north-eastern New Guinea, the majority from Edie Creek, where they werecollected by 1\Ir. Frank H. Taylor. Additional materials were sent to me by Mr.
Taylor and Professor Harvey Sutton, collected by Mr. Norman Ferguson and
Dr. E. A. Holland, and forming a valuable addition to our knowledge of the
Tipulidae of this little-known area. All types and uniques are preserved in thecollection of the School of Public Health and Tlopical Medicine, ilre University
of Sydney. As before, I expt'ess my very sincele thanks to the Director of the
School of Public Health, Professor Harvey Sutton, and to my very good friendand co-worker, Mr. Frank H. Taylor, for this continued co-operation in studying
the Tipulidae of the Territory of New Guinea.
The rich collections studied herewith add no fewer than nine generic andsubgeneric groups to those hitherto known from the Territory, of which
Dolichopezo, s.s., Heliu,s, s.s., and Eurhamphid,ia, are included here. A numberof the large and conspicuous autochthonous Australian genera of the subfamily
Tipulinae that might reasonably be expected to extend their range into New
Guinea remain undetected and, in the light of the collecting done by Mr. Taylor,it may be expected that most of these, at least, clo not occur within the Territory.
Such genera, as Clytocosmus, Platyphasia, Plusiomgia and,Ptilogyna, with branchedantennae, and Leptotarsus and ,.sern.mofes.
with the antennae simple and reduced,
include the largest and most conspicuous crane-flies wiilrin the Australian faunal
limits.
Trpur_rs,rr:.
( P-lruerrrur.,l ) LEUcosrr(,.I..r.
Alesander.
Tipula leucosticta Alexander, philippine Journ. sci., liv., 1984, 444-446.Tipulo (Papuatipula) leucosticta Alexander, ibitl., lvii, lggb, 114-118.
Known hitherto only from Bogadjim (Stephansort), Astrolabe Bay, northeastern New Guinea, collected B0th March, 1g00, by Bird. Two females, Kavieng,
New Ireland, 31 January-14 February, 1gB4 (F. H. Taylor).
Trpur.r

NnpRnorouA FlAvoposrrcarA, n. sp. Text_flgs. 1, ?.
Mesonotal praescutum yellow, with velvety-black stripes, the median one
plumbeous on central portion; scutellum black; pleuraorange, unmarked; legs
black, the posterior tibiae (d) or middle and hind tibiae (?) broaaty yellowon. Continued from These pnocgporNcs, lx,
1988, b1.

170

DIPTERA OF TIIETEBBITOBY OF NEW GI'INEA.

IV,

central portion; wings strongly tinged with blackish, cell Sc and stigma darker;
abdomen (6), including terminalia, almost uniformly blue-black.
12-13 mm.; wing, 10-11 mm. t.-Length,13.5-14 mm.; wing,
J.-Length,11.5-12 mm.
Sexes feebly dimorphic in colour.
3. Frontal prolongation of head orange, the dorsal surface and conspicuous
nasus black. Antennae black, the apex of scape a little reddened; antennae ofmoderate length, if bent backward extending to just beyond root of halteres;
flagellar segments moderately incised. Head orange, without occipital brand.
Pronotum orange. Mesonotal praescutum yellow, with three velvety-black
stripes, the median one with posterior half and central portion of anterior half
more plumbeous in colour; lateral stripes straight; scutum yellow, the lobes chiefly
velvety-black; scutellum blackish-plumbeous, margined laterally with more velvetyblack, the parascutella yellow; mediotergite yellow, on posterior third with aplumbeous area that is narrowly bordered with black. Pleura orange, the pleurotergite more yellowish-orange. Halteres black. Legs with the coxae orange;
trochanters yellow; femora black; fore tibiae blaek; mid-tibiae light brown, thebase and apex blackened; posterior tibiae bright yellow, blackened at both ends;
all tarsi black. Wings (Fig. 1) with a strong blackish tinge, cell Sc and stigma
darker brown; veins brown. Stigma with only flve or six trichia. Venation:
Sc, ending shortly beyond origin of Rs; cell I\I, very short-petiolate to narrowly
sessile.
Abdomen uniformly black, with bluish reflexions; the paratype from Mainishows restricted yellowish areas on bases of tergites two to four. Male terminalia(Fig. 7) relatively small. Ninth tergite, 9f, produced medially into two flattened
divergent ear-like points; a blackened stub at each outer lateral portion of tergite.
Both dististyles darkened; inner style, id, with a very low dorsal crest; apicalbeak unusually slender. Gonapophyses appearing as tiny oval pale plates,
subtending the base of aedeagus.
9. Similar to male, differing as follows: Antennae sholter; scape and pedicel
brown. Halteres somewhat paler in colour'. Legs with both middle and hindtibiae yellow on central portions. Abdomen u.ith basal two tergites and basal
four sternites obscure yellow, the outer segments more blackened. In the Mainiparatypes, the amount of pale colour is somewhat more restricted.
Holotype, 3, Edie Creek, New Guinea, altitude 6,550 feet, February, 1935
(F'. H. Taylor). Allotopotype, ?, in copulation and carded with type. Paratopotypes, 1d, 19; paratypes, L 3,2 Q, Maini, altitude about 6,300feet, July, 1935
(K. J. Clinton).
Neplr,rotoma flauoposticata is readily distinguished from the other describedregional species of the genus by the coloration of the posterior tibiae. The
nearest allies are various members of t}:.e melanura group, especially f'. cl,imid,iata(de Meijere) and -N,'.fttmiscutellata Alexander.
NrpnRoroMA KAvrnNGENSrs,
n. sp. Text-fig. 8.
General coloration yellow; occipital brand not indicated; pronotum orangeyellow; mesonotal praescutum with three black stripes that are confluent or
virtually so, the median stripe velvety-black on cephalic portion, the posterior
three-fourths nacreous, polished; lateral stripes velvety-black, extended lateradto margin of sclerite; scutellum black; wings greyish; stigma small, dark brown;
abdomen with basal six segments orange, the posterior borders of the segments
narrowly and evenly darkened; subterminal segments black; male terminalia
with beak of inner dististyle slender, spinous; gonapophyses bispinous at tips;

By c. p. ALDx-INDER.

L?L

caudal margin of eighth sternite produced medially into a narrow compressed
point.
3. Length about 12 mm.; wing, 10.b mm.
tr'rontal prolongation of head fulvous, unmarked; palpi pale brown. Antennae(d) of moderate length, if bent backward extending to the wing-roor orapproximately so; scape and pedicel orange; flagellum brownish-black; flagellar
segments moderately incised, with verticils that are shorter than the segments.
Head fulvous-orange, the occipital brand not differentiated; a vague dusky cloud
on anterior vertex adjoining the inner margin of eye; vertical tubercle entire or
virtually so.
Pronotum orange-yellow. Mesonotal praescutum with three black stripes,
the polished black laterals virtually confluent with the broad median vitta, thelatter velvety'black on anterior fourth, the remainder of stripe polished nacreous;outer stripes continued laterad to margin, restricting the yellow ground-colour tohumeral triangles and narrow lines adjoining the suture; scutum yellow, eachlobe with a velvety-black area; scutellum velvety-black, parascutella yellon';
mediotergite yellow, with an oval nacreous area at posterior border. pleura andpleurotergite yellow, the anepisternum and sternopleurite vaguely marked withpale reddish-yellow. Halteres yellow, the base of knob infumed. Legs withthecoxae and trochanters yellow; femora obscure brownish-yellow; tibiae brown; tarsi
black. Wings with a greyish tinge; prearcular region and cells C and Sc moreyellow, the latter cell slightly deeper in colour; stigma small, oval,6ark brown;veins dark. Stigmal trichia about fifteen in number'. Venation: Cell M, very
short-petiolate.
Abdomen with basal six segments orange, the posterior bolders very narrowly
ringed u-ith brou'nish-black, the bands uniform in width or virtually so; outer
segments black; terminalia more brownish at tip. Male terminalia (Fig. g) withthe tergite, 9f, extended caudad into two submedian spinous points. Outer dististyle,od, with the apical point relatively short. Inner dististyle, id, with the terminal
beak a slender sclerotized spine; a low dorsal crest. Gonapophyses,gt, terminatingin two aeute spinous points. Eighth sternite, 8s, sheathing, the median portion
produced caudad into a narrow point, on ventral portion strongly
compressed;
surface of sternite with long coarse black setae.
Holotype, J, Kavieng, New Ireland,4 Feb.uary, 1gg4 (F. H. Taylor'). paratopotype, d, 11 February, 1934.
Tlris fly is allied to species such as l'r'plr rotona rlimirliata (de Meijere),N. melanuro (Osten Sacken) and f. s;rccrrlnla (de nleijere). It differs from all
described fol'ms in the pattern of tlie praescutum and the structure of the maleterminalia, notably of the inner dististyles, gonapophysesand eighth sternite.
Dor,rcuopnze ( Dor,rcrrornz.r) T,\l-LonrAt{.\,n. sp. Text_figs. 2, g.
General coloration of mesonotal praescutum dark brown, the humeral region
yellow; a transverse, dark brown girdle on mesepisternum; legs with genua,
tips
of tibiae, and all but proximal portions of basitarsi, white; wings with a weakbrown tinge, conspicuously patterned with darker brown on cord and outerlongitudinal veins; anterior cord oblique, with Rs lying most distad; Sc, opposite
Rs; cell 2nd A narrow; male terminalia with the tergite produced laterad into
triangular darkened lobes.
9.5-10 mm.; wiug, 11-11.b mm. !._Length,
J.-Length,
11_12 mm.; wing,
L2-12.5 mm.
tr'rontal prolongation of head yellow above, darkened laterally; palpi darkbrown. Antennae with the scape yellow, pedicel white; flagellum black; antennae

L72

DIPTERA OF TIIETEBBITOBI

OF NEW GUINE]\'IV'( d ) o f m o d e r a t e l e n g t h , i f b e n t b a c k w a r d e x t e n d i n g a b o u t t o b a s e o foutwardly'
abdomen;gradually decreasing in length
flagellar segments long-cylindrical,
the penultimate; verticils of inter'the terminal segment oval, about one'third
yellow;
the segments themselves' Front
mediate segmenE much shorter thanvertex light brown.
Mesonotal
pronotum darkened medially, paling to yellow on sides.
that become darker and nearly
praescutum with an intermediate pair of stripes
the very extensive lateral darkenings;
confluent behind, similarly fusing withyellow; scutal lobes extensively dark brown' thehumeral region extensively lisht

9Text-figs. 1-10'
fl'oaoposticoto' D' SP'' venation'
1. Nephrotorno '<ooti"noptezo)
taglorian'a' D' sP'' venation'
po\a"nop"ii
2.
venation'tt'
percune@to"
iboucnopezo):p"
3. pou"noiiii
venation'D'sP"
h'olland'i'(Libnotes\
4. Limond,consolilo' n' sp" venation'
5. Limonio (Libtwtes)
procella' tr' sp'' venation'
6. Lilmonti ii"i"aogtochino)male terminalia' details'
7. Nepnro'tomo fl'oaoposticoto' [' sP''
terminalia' details'
maletcoui'zngerosis'n' sp"
8. Neehrol;;in' sp'' male terminalia'
touloriono'(iottciopezo)
9. ooucniiiii
D' sD" male terminalia'
poucnoileZi
r'ercun'eoto'ibotlcnriipeza)
10.
id, inner dististy|e: od, outer dististyle;Symbols: D, basisty|eI g, gonapophysis;
tergite')t,sternite;s,tsY C. P. .ILEXANDER.

1?3

cephalic portion, adjoining the suture, paler; scutellum testaceous-yellow in middle,
darker posteriorly and on sides; mediotergite infuscated. Pleura yellow, with atransverse dark brown girdle occupying the anepisternum and sternopleurite,
being a continuous extension of the lateral praescutal stripes; pleurotergite
similarly darkened. Halteres moderately long, the stem dusky, the extreme basemore greenish; knobs blackened. Legs with the coxae and trochanters yellow;
femora brown to brownish-black, the tips narrowly white to greenish-white,
preceded by a more intensely blackened area; tibiae black, the bases narrowly
white, the amount subequal to the femoral brightening, the tips more extensively
whitened, very narrowly so on fore tibiae, more extensive on posterior tibiae where
nearly the distal sixth is involved; fore and middle basitarsi blackened on proximal
portion, the outer half or more yellowish-white; posterior basitarsi almost entirely
white, the proximal fourth or fifth infuscated; outer tarsal segments white or
greenish-white. Wings (f ig. 2) with a weak brown tinge, patterned with darker
brown, more conspicuously so at the stigma and as broad seams on both anterior
and posterior cords; outer radial and medial veins more narrowly seamed withbrown; veins brownish-black. Holotype with a series of about thirty macrotrichia
in cell Ro, not evident in the remainder of type series. Venation: Anterior cord
oblique, but not as markedly so as in percuneata, n. sp.; Sc, ending opposite Rs;
outer medial forks of moderate depth; m-cu gently arcuated, about one and onehalf times its length before the fork of 1\[; cell 2nd A narrow.
Abdominal tergites obscure yellow, the bases and apices of the segments darkbrown, the latter more broadly so; sixth tergite more uniformly darkened; sternites
and terminalia more uniformly obscure yellow. Male terminalia (Fig. 9) with thetergite, 9f, produced laterad into triangular darkened lobes, the caudal margin
entire, gently emarginate. Both dististyles only feebly sclerotized, the apex of
inner style, id, densely hairy.
Holotype, J, Edie Creek, New Guinea, altitude 6,550 feet, February,(F. H. Taylor).
Allotopotype, t. Paratopotypes, 1 d, 1 ?.

193t

I take very great pleasure in dedicating this handsome crane-fly to Mr.
Frank H. Taylor, who collected the majority of the new and rare Tipulidae
discussed in the present report. The nearest ally is Dolichopeza (Dolichopeza)
percuneata, D. sD., which differs conspicuously in the venation and pattern of thewings, and in the structure of the male terminalia. Both species find their only
near described allies in the eastern Australian fauna, since no member of thetypical subgenus had been discovered n'est of Papua in the Austro-N{alayan islands,
their place being taken by other subgeneric groups of. Dolichopeza that haveevidently been derived from the Oriental faunal region, as Eunesopeza .Llexander,
Mitopeza Edwards and .l/esopeeoAlexander. The presence of macrotrichia in cell
Ru of the wing of the holotype specimen of the present fly and not in the remainderof the type-series is distinctly puzzling, since macrotrichia in the wing-cells ofother species have been found to be relatively constant both in number anddistribution.
(Dor,rcrrornze) pERcuNEArA, n. sp.

Text-fi9s. 3, 10.
General coloration of mesonotum brownish-black; pleura yellow, with atransverse brownish-black girdle covering the anepisternum and sternopleurite;
legs black, the genua very narrowly white, the tips of all tibiae somewhat more
broadly so; tips of basitarsi and succeeding tarsal segments snowy-white; wings
unusually narrowed at base; .Rs very short, lying far distad of the other elementsof the anterior cord; medial forks deep; cell Znd A very long and narrow; male
Dor,rcgoprzl

774

DIPTERA OI' THE

TERRITORI

OF NE1V GUINEA.

IV,

hypopygium with the outer dististyle expanded at apex into an obtuse spatulate
head.
J.-Length about 8 mm.; wing, 9 mm.
Frontal prolongation of head brownish-yellow; palpi brownish-black. Antennae
black, the scape and pedicel a trifle paler; flagellar segments cylindrical, theverticils shorter than the segments. Head with the front yellow, the vertex morecastaneous,the posterior orbits darker.
pronotum brownish-black above, yellow on sides. Mesonotal praescutum
brownish-black, the humeral region of the yellow ground-colour; median regionof praescutum a little paler behind; scutal lobes brownish-black posteriorly, muchpaler in front, adjoining the suture; posterior sclerites of notum brownish-black,the scutellum paler on anterior portion. Pleura yellow, the anepistet'num andsternopleurite brownish-black, a direct continuation of the lateral portions of thepraescutum, forming a conspicuous transverse girdle; pleurotergite dark brown.
Halteres very long and slender, black. Legs with the coxae yellow; trochanterstestaceous; femora black, the tips very narrowly snowy-white; tibiae black, theextreme bases white, the tips more broadly so, the amount subequal on all legs orsomewhat narrolryer on the mid-tibiae, including one-eighth or less of the segment;
tarsi dusky, the outer ends of basi-tarsi and the remaining tarsal segments snowywhite. Wings (f''ig.3) unusually narrowed at base into a petiole; strongly tinged
with brown, the prearcular field and cells C and Sc mol'e yellowish-brown; stigma
small, darker brown; a very narrow and restricted brown seam on anterior cord;veins brownish-black. Venation: Rs unusually short, transverse, lying far distad,some distance beyond Scr, delimiting the proximal end of stigma; cord unusually
oblique, the fork of M lying most basad; outer medial forks deep; m'cu about
one-half its length before fork of M; cell 2nd A long but reduced to a narrow
strip.
Abdominal tergites black, with a yellow ring at near midlength of theindividual segments, this subequal to or a trifle narrower than the blackenedbase and apex of the segment; subterminal sclerites and terminalia more uniformly
blackened; sternites more extensively light yellow, the incisures narrowly darkened.
Male terminalia (Fig. 10) small and with few distinctive features, othel than the
dististyles. Ninth tergite, 9t, with the caudal margin narrowly blackened. Outerdististyle, ,od, expanded at apex into an obtuse spatulate head. Inner style, id,
shorter, more narlowed outwardly, the tip obtuse, the basal three-fourths withlong, coarse setae.
Holotype, 3, Edie Creek, New Guinea, altitude 6,550 feet, February, 1935
(F. H. Taylor).
By Skuse,s table to the Australian species of Dolicltopeza (Pnoc. Lrxx. Soc.
) nnulipes
N . S . W . , ( 2 ) 5 , 1 g 9 0 , 6 0 ) , t h i s f l y r u n s t o D o l i c h o p e z a( D o l i c l r ' o p e z aa
andAlexander(D.)
dorrigensis
Skuse. It is somewhat more closely allied to D.
The nearest ally is
D. (D.) kurand,ensis Alexander, of eastern Australia.
underthat species. The
undoubtedly D. (D.) tagloriana, n. sp., as discussed
venation, with Rs lying far distad, and with the cord unusuall] oblique, is somewhat suggestive of that found in the genus Scantbonerro Osten Sacken, and marks
the extreme condition in this venational peculiarity so far discovered in the genus
Dolichopeza.
MeorstocnnAruscANe (Wiedemann).
Nematocera luscana Wiedemann, Dipt. Exot., 1, 1821,29. 1 J, Salamaua, New
Guinea, 16 December,1933 (F. H- Taylor).

BY C. P. ALEXANDER.

L75

Lruoxrnr.te.
Lrlronrrxr.
Lnror.rre (Lruonrl)
ExpEDrrA,n. sp. Text-flg. 11.
General coloration of mesonotal praescutum reddish-brown, with a dark brown
median stripe; pleura reddish-brown, with a narrow blackish longitudinal stripe;
halteres chiefly blackened; legs black, the femoral bases brightened; wings with abrownish tinge; stigma oval, darker brown; costal fringe(d)long and
conspicuous; cell M, open by the atrophy of m; abdominal tergites black; maleterminalia deeply notched medially; dististyle single, produced into a curved
blackened spinous point.
6. Length about 4.2 mm.; wing, b mm.
Rostrum obscure yellow; palpi black. Antennae black throughout; flagellar
segments oval, with short glabrous apical pedicels. Head grey, lighter on front.
Pronotum dark brown above, paler laterally.
Mesonotal praescutum high andgibbous, reddish-brown, with a dark brown median stripe, the lateral stripes paler
and ill-defined; scutum reddish-brown, the centres of the lobes darkened; scutellum
brown, darker posteriorly; mediotergite dark brown. Pleura reddish-brown, witha narrow blackish longitudinal stripe extending from the fore coxae to theabdomen, passing beneath the root of halteres. Halteres pale basally, the distal
portion of stem and the knobs blackened. Legs with the fore coxae blackened,
the remaining coxae and all trochanters reddish-yellow; femora obscure yellow
basally, passing into black at near midlength; tibiae and tarsi black. Wings
with a brownish tinge; stigma oval, darker brown; veins black. Costal fringe(d) long and conspicuous, though sparse. Venation: Sc long, Sc, ending about
opposite three-fourths the length of Rs, Sc, some distance from the tip of Sc'
the lattter about one-half longer than r-m; free tip of Sc, and R, both pale andin transverse alignment; basal section of Rn*ulong, exceeding one-half the lengthof Rs; cell M, open by the atrophy of m; m-cu just before fork of M; anal veins
gently convergent at bases.
Abdomen with tergites and terminalia black; sternites obscure yellow, theincisures dusky; terminal segments more uniformly darkened. Male terminalia(Fig. 11) with the tergite, 9f, profoundly notched medially, the lateral lobes
glabrous, darkened, their tips obtuse. Basistyle, D, with the ventro-mesal lobe
very low and stout. A single dististyle, d, from a swollen pale base, the distal
half narrowed into a curved blackened spine, the tip acute, the surface with
scattered setae and setulae, including one spinous bristle at near one-fourth thelength that may be homologous with the usual spines of the rostral prolongation
found throughout the genus. Gonapophyses, g, ll-ith the mesal-apical lobe verybroad and flat, pale, the apex irregularly obtuse.
Holotype, 3, Edie Creek, New Guinea, altitude 6,550 feet, February, 193b
(F. H. Taylor).
The type-specimen is in very indifrerent eondition, both wings being badly
torn. The species is so distinct that its recognition will be simple. It is the only
species of the subgenus Limonia so far described from the Australasian regionin which cell NI, of the wings is open by the atrophy of m, rather than by that of
the basal section of vein Mr. In the Oriental fauna, various members of the pacatagroup, such as pucata Alexander, pacatella Alexander, and others, possess thischaracter.
LruoNre (Lrnxorns) Hor.LANDr,n. qp. Text-figs. 4, LZ.
General coloration obscure yellow, conspicuously patterned with brownishblack and blaek; antennae black throughout; tips of femora narrowly blackened;

176

DIPTER.\ O[' TIIE TERRITORY OF NEW GUINE.\.

I\',

wings weakly tinged with brown, spalsely patterned with darker brown; Rs verystrongly arcuated, perpendicular at origin or nearly so; cell 1st M, elongate, withm-cu at near midlength; m and basal section of M. subequal; anal veins convergent
at bases; male terminalia with the dorsal dististyle present as an acute rod;
rostral prolongation of ventral dististyle broadly flattened, produced into a slenderapical point; gonapophyses with mesal-apical lobe slender, the lnargin withmicroscopic denticles.
J.-Length
wing, ? mm.

about 6'5-6.7 mm.; wing, 6-6.5 mm.

Q.-Length

about 7 mm.;

Rostrum and palpi black. Antennae black throughout; flagellar segments (J)short-oval, the first subglobular, the outermost more elongate; terminal segmeut
a little exceeding the penultimate, pointed at tip; apices of flagellar segments
glabrous but not constricted into necks; longest verticils subequal in length to theintermediate segments. Head dark grey, the occiput and posterior vertex with avelvety-black area; anterior vertex reduced to a linear strip in both sexes.
Pronotum dark brown, more or less pruinose. Mesonotal praescutum brownishyellow, with a broken brownish-black pattern, the usual three stripes narrow,especially the median one, the laterals confluent at their anterior ends with themedian vitta, isolating two linear strips of the ground-colour on posterior half ofsclerite before the suture; lateral and humeral portions of praescutum similarly
darkened; scutum pale medially, the lobes darker, more intensely blackened
along their mesal edges; scutellum black, the parascutella a little paler; postnotum
almost uniformly dalkened. Pleura brownish-yellow, striped longitudinally withblackish, including a more dorsal alea across propleura and anepisternum, and ascarcely separated line across the dorsal sternopleurite and ventral meron; ventral
sternopleurite clearer yellow. Halteres obscule yellow at base, the outer portion
dusky, the extreme tip more yellowish. Legs with the coxae pale, the fore coxae
a trifle darkened; trochanters yellow; femora obscure brownish-yellow, the tips
narrowly but conspicuously blackened, the amount subequal on all the legs, thedark apex preceded by a very nal'row, scarcely evident, clearer yellow ring; tibiae
brownish-yellow, the tips very narrou'ly darkened; tarsi brownish-yellow, passing
into black. Wings (Fig. a) weakly tinged with brown, restrictedly patterned withslightly darker brown; cells C and Sc more brownish-yellow; the dark aleas
include arculus, origin of Rs, cord, outer end of cell lst M., and the small stigma;
wing-tip and longitudinal veins very narrowly and insensibly seamed with brown;veins dark, paler in costal region. Venation: Sc moderately long, Sc, ending
opposite r-m, Sc, at tip; Rs short and very strongly arcuated at origin, being
perpendicular or virtually so; free tip of Sc, and R, in transverse alignment;m<u at near midlength of the long cell 1st M,; m and outer deflection of M"
subequal; anal veins slightly convergent at origin.
Abdomen bicolorous, the tergites brownish-black, the bases of the individual
segments narrowly yellow; sternites obscure yellow, the dark colour vaguely
shown on the outer segments; pleura variegated with darker; eighth segment (J)obscure yellow; terminalia brownish-yellow. Male terminalia (Fie. 12) with thecaudal margin of tergite, 9t, gently emarginate, the lateral lobes low and obtuse,
with strong setae. Ventromesal lobe of basistyle, D, simple. Dorsal dististyle, dd,
a strong chitinized rod, the tip acute. Ventral dististyle, z*d, deeply bilobed, theouter lobe dusky, with conspicuous setae; inner lobe consisting of the broadly
flattened rostral prolongation which is extended into a long slender point, theentire prolongation pale yellow. Gonapophyses,g, with mesal-apical angle long777

BY C. P. .ILEXANDER.

and straight, the inner edge with very acute microscopic denticles.
with cerci very slender, the tips acute.

Ovipositor

Holotype,6f, Kavieng, New Ireland, 16 February, 1984 (Dr. E. A. Holland).
Allotopotyp€, 9, carded with type. paratopotype, d.
This very distinct erane-fly is named in honour of the collector of the type
series, Dr. E. A. Holland. The species is most generally similar to Li.rnonia(Libnotes) perkinsi (Grimshaw), widely distributed in the Pacific Islands,
differing notably in the coloration, venation, and structure of the male terminalia.
Lruowra (Lrnlvorns) oBLreuA (Alexander), var.
Libnotes obliqua Alexander, Rec. south australian lItts., ii, tgzz, zgz.
L 3, I t, Bulolo, New Guinea, altitude 2,200 feet, 26 December, 1g33 (F. H.
Taylor).
The problem of specific distinction is well shown in the present pair ofindividuals (vide papers by the writer: Pnoc. Lrwrv. soc. N.s.w., lx, 1g3b, 60-61;
Reoue Srrisse de Zoologie, 43, 1936, 89).
Lr*roNre (Lrnxorrs)

sor,olroltrs (Alexander).

Libnotes solomonis Alexander, Ann. Mag. Nat. Hist., (9) xiii, 1924, 39.
Recorded from several Paciflc Islands (Solomons, New Britain, Santa Cruz,
Reef Islands). Pondo, New Britain, 28 November, 1933 (F. H. Taylor).
Lruonr.r (Lrnworns) coNsoNl\,n. sp.

'

Text-figs. 5, 13.

General coloration of thorax brownish-black, the humeral region of praescutum
obscure yellow; femora black, the tips narrowly and abruptly yellow; tarsi paling
to light brown; wings greyish-yellow, the ground almost concealed by an extensive
brown pattern that appears as crossbands aud seams to the veins; basal sectiou of
R.*u about one-half Rs; cell 1st M, relatively small, about one-half the length ofcell 2nd M,; m-cu beyond midlength of cell 1st Mr; male hypopygium with theventral dististyle large and fleshy, the rostral prolongation with two very unequal
spines, the inner one reduced to a seta.
about 9'5 mm.; wing, 11 mm.
J.-Length
Antennae black; flagellar
Rostrum dark brown; palpi brownish-black.
segments long-oval; verticils shorter than the segments. Front dusky; posterior
portions of head grey; anterior vertex reduced to a linear strip.
Pronotum brownish-black. Mesonotum brownish-black, the humeral regionof praescutum obscure yellow, the remainder of the sclerite entilely occupied bya dorsal black shield; median region of scutum and scutellum obscure yellow.
Pleura brownish-black, including the dorso-pleural membrane. Halteres dusky,
the knobs more infuscated. Legs with the coxae dark brown; trochanters obscuretestaceous yellow; femora black, the tips narrowly and abruptly yellow, the amountsubequal on all legs; tibiae black; tarsi paling to light brown; claws'with a long
slender spine at near midlength. Wings (Fig. 5) with the ground-colour greyishyellow, almost covered by extensive brownish clouds and cross-bands, including
a broad area crossing the basal cells beyond arculus; a second band at level oforigin of Rs, not involving cells C or Sc; broad areas along cold and outer endof cell 1st Mr; most of cells beyond cord suffused, variegated only by small difruse
areas of the ground; cells C and Sc chiefly yellow; stigma small, short-oval,
darker brown; veins brown. Venation: Sc, ending opposite or just beyond r-m,
Sc, a short distance from tip; free tip of Sc, in virtual alignment with R,; basal
section of Rr*e about one-half Rs; cell 1st 1\I, relatively small, about one-half cell

178

DTPTEBA OF TIIETERBITORY OI' NEW

GUINDA.

ff,

2nd Mr; m-cu beyond midlength of cell 1st Mr, longer than distal section of Cur;
anal veins gently divergent.
Abdominal tergites, including terminalia, brownish-black, the ertreme bases of
the intermediate segments pale; sternites a little brightened. Male terminalia(Fig. 13) with the tergite,9f, moderately notched, the setae of the lateral lobes

tl

t2

t7
Text-figs.11-17.
11. I'dmonio (Limonio) erped'ita, r. sP., male terminalia'
12. Limonio (Libnotes) hollandi, D. sP., male terminalia.
13. Limonio (Libnotes) conson&, D. sp.r male terminalia.
14. Limonio (ThrApticomuio) spathulilero, r. sD', male terminalia.
15. Orimorga (Orimarga) papuicolo, n. sp., venation.
ouronticolor, D. sD., male terrninalia.
16. Helius (Eurhomphidio)
17. Heliu,s (Eurhornphidia) melanosorncL,n. sp., male terminalia.
( Symbols : o, aedea8ius; D, basistyle ; d, dististyle; dd' dorsal dististyle ;g, gonapophysis; i, interbase ; id, inner dististyle ; oil, outer dististyle ;t, terBite ; od, ventral dististYle. )Basistyle, D, much smaller than the ventral dististyle, withalnrost all marginal.
lobe. Ventral dististyle, o*d', large and fleshy, its rostral
nornral small ventromesal
prolongation
stout, with two spines of very unequal size, the outer one a strong
Gonapo'black spike, the inner spine basal in Bosition, reduced to a mere seta.
serrulate.
physes, g, with the inner margin of the mesal-apical lobe irregularly

Holotype, J, Edie Creek, New Guinea, altitude 6,550 feet, February, 1935
(F. H. Taylor).
Limonia (Libnotes) consona is a large, striking species that requires noconrparison with any of the numerous regional forms so far described. By

1?9

BY C. P. ALNVANDER,

Edwards' key to the species of. Libnotes (Journ. Feitr. Malay St. Mus., 14, 1928,
74-80), the fly runs to Z. (L.),nonti'v*agans (Alerander), of Java, an entirely
difierent species.
Lrltoxre(Drcnewourre)

Dicranornyia sord,ida Brunetti,
pp. 382-383.

sonoroe (Brunetti).

Fauna Brit.

India, Dipt. Nematocera, 1912,

Widespread throughout the Oriental Region. The present material is deflnitely(Alexander), characreferable to sorilida rather than to the allied illinoworthi
teristic of the Pacific Islands.
Edie Creek, New Guinea, altitude 6,550 feet, February, 1935 (F. H. Taylor).
M a i n i , P a p u a , a l t i t u d e a b o u t 6 , 3 0 0f e e t , J u l y , 1 9 3 5 ( K . J . C l i n t o n ) .
Lruoxrr(Trrnnrrcourre)

ABAcrrNopErLA (Alexander).

Dicranomyia (Thrypticontyia) arachnophitro Alexand,er, Philippine Journ Sci.,
xxxiii, 1927, 301.
Numerous specimens ofRecorded from the Philippines and New Britain.
both sexes, Salam'aua, New Guinea, 21 August, 1935 (Norman Ferguson).
N{r. Ferguson furnislred notes of unusual interest relating to the habit ofthese flies of resting and dancing on spiders' webs and long strands of spiders'silk. Similar habits in this same species had been discussed by the writer in an
earlier paper (1.c.,xxxiii, t927,299), but in the present instance, instead of only
a few individuals being involved, Mr. Ferguson states that many hundreds ofindividuals were encountered.
In this material of arachnophila, the wing-tips are slightly infumed, but not
so markedly so as in apicalis (Wiedemann), lumidapicalis (Alexander) andspatlrutifero, n. sp. The male terminalia lacks the two especially modified elongate
setae at the outer lateral angle of the ninth tergite, and the conformation of therostral prolongation and its spines is distinctive of the present fly.
n. sp. Text'fig. 14.
sPATHULTI'ERA,
General coloration dalk plumbeous-grey; ploximal fifth of basitarsi blackened;wings hyaline, the cells beyond cold very strongly infumed; Sc' ending a shortdistance before origin of Rs; m-cu at near three-fifths the length of cell lst M";
abdomen black, the basal segment brown; male terminalia with the aedeagus
expanded at apex into a subcircular oval spatula, the surface provided withnumerous setae.
5-5'5 mm.; wing,
5-5'5 mm.; wing, 6-6'3 mm.
i.-Length,
J.-Length,6-6'2 mm.
Rostrum and palpi black. Antennae relatively elongate, black; flagellar
segments long-oval, with short pedicels and vely long verticils. Head dark grey.
Pronotum and mesonotum almost unifolmly dark plumbeous-grey. Pleura
heavily grey pruinose, the sternites more testaceous. Halteres with base of stem
yellow, the outer end and knob blackened. Legs with the coxae and trochanterstestaceous-yellow; femora black, the bases restrictedly paler; tibiae black; tarsi
snowy-white, the proximal fifth of basitarsi black, the amount of the latter subequalon all legs. Wings hyaline, the cells beyond cord very strongly infumed; stigma
long-oval, even darker brown; veins black. Venation: Sc relatively long, Sct
ending a short distance before origin of Rs, the distance on costa about as longas the basal section of vein Mr*r; fre€ tip of Sc, far before Rr, R, alone exceeding
m-cu; m-cu at near three-fifths the length of cell 1st M'.
Lrrtoxre(Trrnrnrrcorrtrl)

180

DIPTERA OI' TIIETERRITOITY OF NEW CUINE,\.

IV'

Abdomen black, only the basal tergite and sternite brown. Male terminalia(Fig. 14) with the tergite, 9t, transverse, the caudal margin gently emarginate,each side with five setae, of which two are much stronger and placed close
together. Basistyle, b, with ventro-mesal lobe unmodifled. Ventral dististyle, od,with the spines separated, the more basal from a low tubercle, the outer sessile.
Aedeagus, o, at apex expanded into a subcircular to transversely oval spatula ofthin membrane, this set u'ith numerous setae, all of which are directed mesadand caudad.
Holotype, J, Admiralty Group, Lombrum, Manus Island, 19 February, 1934
(F.H.Taylor).
Allotopotype, Q, earded with type. Paratopotypes,2 6,2 9,
on cards.
The most similar described species is Limonia (ThrgpticomUia) fumidapicalis(Alexander), of north-eastern Australia. This has the wing-tip much less strongly
darkened, Sc longer, with Sq ending opposite the origin of Rs, and with thedetails of structure of the male terminalia quite distinct. The peculiarly dilated
apex of the aedeagus is different from that of all other species of the subgenus
so far discovered.
PRocELLA,n. sp. Text-fig. 6.
General coloration of mesonotum dark brown, with a more brownish-yellow
median line; a broad, pale yellow, longitudinal stripe across pleura, the ventral
sternopleurite and meral region dark; tibiae white, with a single, relatively narrow,dark ring at near midlength; wings whitish subhyaline, the stigma conspicuous;
Sc relatively long, Sq ending distinctly beyond fork of Rs; medial fork deep;
m-cu approximately its own length beyond fork of M; cell 2nd A narrow; abdominal
Lruoxre(PsnuoocrocnrNe)

segments bicoloured.
6-6'5 mm'; *'ing,
5'5-6'5 mm.; wing, 5'8-6'8 mm. i.-Length,
J.-Length,6-6'5 mm.
Rostrum and palpi brownish-black. Antennae black; flagellar segments oval,
with very short, glabrous, apical pedicels; verticils about as long as the segments.
Head light brown.
Pronotum pale brownish-yellow. Mesonotum brownish-yellow down the central
portion, the sides darker, the praescutum a little more intensely coloured in
Pleura chiefly pale brownish-yellow, the ventral sternofront and laterally.
pleurite and meral region brownish-black. Halteres dusky, the knobs even darker.
Legs with the coxae darkened; trochanters obscure yellow, the fore pair darkened;
fore femora yellow, very narrowly darkened at base, the tip a little more extensively so; middle femora dusky on basal half, outwardly paling to whitish, thetips narrowly blackened; posterior femora almost uniformly blackened, the tip
a litle more intense; all tibiae snowy-white, with a single, relatively narrow,darkened ring at near midlength; tarsi white. Wings (FiS. 6) whitish subhyaline,
unmarked except for the oval, dark brown stigma; veins brown. Venation:
Sc relatively long, Sc, ending some distance beyond fork of Rs, in cases extending
to just before the level of r-m; Sc, opposite origin of Rs or nearly so; medial fork
deep; m-cu beyond the fork of M, the distance variable, usually approximately
equal to the length of the vein itself; distal section of Cu, a little longer thanm-cu; cell 2nd A narrow.
Abdominal tergites bicolorous, dark brown, the caudal portions of the segments
obscure yellow; lateral borders of seeond tergite darkened; sternites obscureyellow, the outer segments restrictedly darkened subapically; terminalia small,dusky.

BY C. P. .\LEXANDER.

181

Holotype, J, Kavieng, New Ireland, 14 February, 1934 (F.H.Taylor).
Allotopotype, t, carded with type. Paratopotypes, 12 d, 9, 10-16 February, 1934 (Taylorand Holland).
The nearest ally is Limonia (Pseud,ogloch,ina)kobusi (de Meijere) of Javaand Sumatra, which differs in the coloration of the thorax and in the wingvenation, especially the less oblique cord, longer Sc, and slightly narrower cell
2nd A. Both species agree in having a single narrow tibial ring and with m-cu
some distance beyond the fork of M.
(Psnuoocr,ocurrve) EvANEscENs,n. sp.
Lrlroxn
Antennae (J) relatively long; flagellar segments with glabrous apical necks,
the main body of the segments with long dark pubescence; tibiae white, with asingle, nearly evanescent, dark ring at near midlength; wings u'ith Sq ending
shortly before the fork of M; m-cu at or close to the fork of M; abdominal
tergites bicolorous; sternites and terminalia uniformly yellow.
J.-Length about 5.5 mm.; wing, 5 mm.
Rostrum and palpi testaceous brown. Antennae relatively elongate, much
Ionger than in procella, n. sp.; flagellar segments oval to long-oval, with longconspicuous glabrous apical pedicels; surface of segments with a dense conspicuous
erect pubescence that is only a little shorter than the verticils; in Ttrocella, t};repubesceneeof the flagellar segments is short and relatively inconspicuous. Head
dark.
Pattern of mesonotum not readily discernible in unique type because of methodof mounting, but apparently much as in procella. Pleura chiefly pale, dark brorvn
on the ventral sternopleurite. Halteres dusky, the knobs darker. Legs with thefore femora white, narrowly dark brown at both ends; mid-femora dusky on basal
half, the outer half paling to white, the tips narrowly darkened; posterior femora
infumed, the central area somewhat more brightened; tibiae white, with a very
narrow, scarcely indicated, pale brown ring at near midlength, this being so faintas to be virtually evanescent; tarsi white. Wings subhyaline, unmarked except
for the small, oval, dark brown stigma; veins brown. Venation: Se relatively
short, Sq ending just before the level of the fork of M; medial fork of moderate
depth; m-cu at or very close to fork of M, shorter than the distal seetion of veinCu,; cell 2nd A relatively narrow but long.
Abdominal tergites dark brown, the caudal borders narrowly brownish-yellow;
sternites and terminalia uniformly yellow.
Holotype, J, Admiralty Group, Lombrum, Manus Island, 1g February, 1934
(F'. H. Taylor).
The nearest regional ally is the type of the subgenus, Limonia (Pseudoglochina)pulchripes (Alexander) of northern Queensland, distinguished by the leg-patternand details of venation. In the present fly, the dark tibial ring is so pale andnarro\ry as to be nearly lacking.
Onru,rno-1,(Onrlrenoe) rerurcor,A, n. sp. Text-flg. lb.
General coloration light grey, the praescutum and scutum searcely variegatedby darker grey; knobs of halteres moderately infuscated; wings subhyaline,
unmarked ; macrotrichia of outer radial and medial veins abuudant; free tip ofSe, far before Rr; Rs and R*, subequal; Rr,, about twice R, alone; abdomen
brownish-black.
about 7 mm.; wing, 5.8 mm.
t.-Length
Rostrum grey; palpi brownish-black. Antennae black, the scape pruinose:
flagellar segments oval, gradually decreasing in length outwardly. rread grey.

182

DIPTERA OF TIIETERRITORY OF NEW GUINEA.

IY,

Pronotum and mesonotum uniformly light grey, the praescutum and scutum
scarcely variegated by darker grey. Pleura dark grey. Halteres white, the knobs
moderately infuscated. Legs with the coxae dark, heavily pruinose; trochantersp'ale brown; remainder of legs broken. Wings (f ig. 15) subhyaline, unmarked;
veins pale. Macrotrichia of veins relatively abundant, including complete series
on outer radial and medial veins, with about 25 on R.; 30 on distal section of
R.*o; 40 on distal section of Mr*r; and 35 on distal section of Mr; no trichia on Rs,
main stems of M or Cu, or either anal vein. Venation: X'ree tip of Sc, far before
Rr, the distance approximately one-half longer than the latter vein; Rs and Rr*,
subequali Rr*, about twice R, alone; Mr*. B little shorter than Mr; m-cu about
opposite two-thirds to three-fourths the length of Rs; cell 2nd A relatively long
and wide.
Abdomen brownish-black, including the bases of the ovipositor; cerci darkbrown.
Holotype, ?, Edie creek, New Guinea, altitude 6,bb0 feet, February, lg3b
(F'. H. Taylor).
The only described regional species of. Orimargo with which this fly may be
profitably compared is orimarga (orimarga) hapopugiolis Alexander (celebes).
The latter differs in coloration and, especially,in the details of venation, including
the strongly angulated Rs, loss of the free tip of Scr, longel. Rr*r, deeper cell lI.
and position of m-cu.
Hnr.rus (Hu.rus) ANAElrrrcusAlexander.
Helitts (Helius) anaemicus Alexander, philippine J,ourn. sci., xlix, 1gg2,
254-255.
Hitherto known from the Loochoo Islands, tr'ormosa, Luzon and l\Iindanao,
in all cases at low altitudes. Two male specimens, Kavieng, New Ireland, 16
February, 1934 (Dr. E. A. Holland). A wing of one of the specimensshowsabnormal venation. By existing keys, the fly runs to Helius (Helius) unicolor(Brunetti), a very different species.
Hnr,rus (Eunrr,rrmrrrora) ,\L:R.\NTrcoLoR,
n. sp. Text-fig. 16.
General coloration liSht orange, the terminal abdominal segrnents black;head grey; halteres pale throughout; legs obscure yellor',',the outer ends of thetibiae and tarsi paling to white; wings tinged with yellow, the veins very pale;a conspicuouspale brown cloud extends from the stigma along the anterior cord
to the fork of M; male terminalia with the dististyles apical in position.
J.-Length about 4 mm.; wing, 4.6 mm.
Rostrum about equal in length to remainder of head, brown; palpi darker
brown. Antennae short, pale brown throughout; flagellar segments long-oval;
verticils much exceeding the segments. Head light grey; anterior vertex about
as wide as the rostrum.
Pronotum and mesonotum entirely light orange, unmarked. Halteres palethroughout. Legs with the coxae and trochanters pale orange-yellow; femora
obscure yellow; tibiae a trifle darker, especially the posterior legs, the outer ends
and tarsi paling to white; genua not brightened. Wings with a yellowish tinge,
variegated only by an oblique, pale brorvn cloud extending from the stigma along
the anterior cord to the fork of M; veins very pale yellow, a trifle darker in theclouded area. A few macrotrichia on outer ends of veins Rn*u,Mr*, and M3.
Venation: r-m a little less than its own length before the fork of Rs; cell 1st M,
large, its inner end lying proximad of other elements of cord; m shortened: m-cu
about its own length beyond fork of M.

BI

C. P. ALEXANDER.

183

Abdomen obscure yellow, the seventh to ninth segments, inclusive, black; morebasal tergites vaguely darkened medially on basal ring. Male terminalia (f ig. 16)
with the basistyle, b, slender, terminating in a small glabrous blackened point;
dististyles apical in position; outer style, od, extended into a long point; innerstyle, id, with a conspicuousincision at near midlength, delimiting a conspicuousbasal lobe.
Holotype, J, Edie Creek, New Guinea, altitude 6,550 feet, February, 1935
(F. H. Taylor).
Helius (Eurharnphidia) auranticol^oris very different from the other described
species of the subgenus in the light orange colour of the thorax, in conjunction
with the patterned wings.
Hnr,rus (EtrnuelmHrnra) trELANosolrA,n. sp. Text-fig. 17.
General coloration polished black; mesonotal praeseutum strongly arched;knobs of halteres darkened; legs dirty white, the tarsi clearer; snowy-white;wings strongly tinged with grey, the stigma dark brown, conspicuous; abdomeu
bicoloured, black, the posterior borders of the intermediate segmentsbroadly pale.
J.-Length, 4.8-5 mm.; wing, 5-5.2 mm.
Rostrum longer than remainder of head, black; palpi black. Antennae black
throughout, about one-half longer than the rostrum. Head black, sparsely pruinose.
Pronotum and mesonotum polished black, the praescutum very strongly arched.
Pleura polished black, the dorso-pleuralmembrane pale. Halteres white, the knobs
infuscated. Legs with the fore coxae blackened, mid-coxae infuscated, posterior
coxae yellow; trochanters obscure yellow; remainder of legs chiefly dirty white,
the femora and tibiae somewhat more obscurethan the snowy-white tarsi; terminal
tarsal segmentsdarkened; genua vaguely brighter than the adjoining parts of thefemora and tibiae. Wings rather strongly tinged with grey, cells C and Sc weakly
infumed; prearcular region whitish; stigma oval, darker brown, conspicuous;veins brown. Costal fringe relatively long and conspicuous. Venation: Sc, ending
just before level of r-m, Sc, at its tip; r-m variable, from about its own length tonearly twice this distance before the fork of Rs; m-cu beyond midlength of celllst N{r.
Abdomen conspicuouslybicoloured, black, the caudal borders of segments two
to six, inclusive, broadly pale; subterminal segments and terminalia black. Maleterminalia (Fig. 17) with the sternite bearing blackened cushions on either sideof midline, these densely set with microscopic setulae. Basistyle, b, slender.
Outer dististyle, od, terminating in a simple, gently curved point; inner style, id,longer. Interbasal plates, i, produced into long straight spines.
Holotype, 6, Edie Creek, New Guinea, altitude 6,550 feet, February, 1935
(F. H. Taylor). Paratopotype, d.
Helius (Eurhamph,id,ia) ntelanosomo is readily distinguished from all otherdescribed regional allies by the polished black colour of the body and by thewing-pattern.