508

Tke Pki.l,ippirreloumd, of Sci,eflce

Unonia lLi atuie) retuiasd Alet,6,000 to 8,000 fet, October 28-24.
Lintuid (Ubnot6, ha$enaw sp. aoa., 4,600 to 6,000 Jet, Augrst 30.
(Alex.),3,500 feet, Octobe! 24; 6,000
Liwnia @hipidid) ldtuoso
to ?,000 feet, Ausust 30.
Atuaiasp. no1t.,4,500 tt 6,000 feei, Auslst 30.
l&hipid,itr) triMt'
Uwnitt <Dicrdhonyia, depdrperatd l\lex.l,3,500 feet, October 21.
L;hohid (Diff@omyi4) M|otuorir (t\16x.),6,500 to ?,800feot, OctG
be" 26.
Linotuid (DidfuotuAiot ./did4 (Brunetti),6,000 to 7,500 iet, Au8!!t 30-31.
Unonia. lDicranorrwia) srbpMctuldta sp. nov., 3,500feet, October21.
Linotuie (Gewnorwi.r, dlpestris Alea.,5,500 to ?,800 feet Octobe! 24.
Anto.ha lAktoch@)btfdo Alex., 3,600feet, Au$st 29.
Antacha \Anto.ht) st!'r sp, nov., 3,500 Jeet Oclober 24.
HeUaa lHelius) eLtetuatts 41ea.,5,600teet, Octob€r 22.
Eeti6 (Elchmph lnt, p.telagars st. nov., 6,000 t6 ?,000 feet, August 30.
Th&tuM9toptere lT@i|oMita, istihiMd Alex., 5,600 fet, October 22.
Diqanaptscha i$ihind .p. nov., 3,500 to 5,500 f@t, Ociober 22.
Nippotomlrio ssnlphatetes (Alex.), 5,600 feet, Octobe! 22.
rricgphow ait@
Aiex., 6,600 feet, October 22; 6,000 to ?,000 feet,
Tdauphona lorwsq@ Alex., 4,600to ?,500feet, Ausust 30 .hd 31.
Ditanota <Ah6lopina) delectdta6p. nov., 6,000to 8,000feei, O.iober
23.
Dioanotd (AmdlopiM) 9i66eld (AIex.), v.r., 4,600to 6,000feet, AuAd.e|pllonuia issikiea sp. nov., 6,60{ fet, October 22.
Epiphra.sM .ti:tri.d Alex., 3,500 to 5,500 feet, October 22.
Pseudolimaphita oLtm@lis Llex., 5,600 to ?,800 fet, Ocrober 22 to
24.
Lin@phil@ (Priotulab.is) setm,tutatd sp. nov., 3,500 to 8,000 feet,
Octobe. 22 to 24.
Li@ophila lDiqamphrasw)
lomosd Lle\., 6,000 to ?,000 Jeet, August 30.
Lintuopbikt \Diaranoptdstfu)
tdiatuehsis Alex., 6,500 to ?,500 fet,'

At@b@ lAtarbodes) fBca@mis EdMrds, 3,600 leet, October 21.
Atwbd (Atbbodes, leptarqath@Nex.,6,000 to 7,000feet August 30_
Atqrba \Atarbod@) poilt.rt.omtu Ed{ards,4,500 to 6,000 feet, Aug8t 30; 3,500to 5,600feet, Octoter 22.
Elephanto'ryia lElphdntomvia) serotind Alex., 6,500 to ?,800 feet,
ELphtutottqid(Eleph^ntotluod6)

afuilomri, Alei., 4,500 to 6,000 feet,

Neolituiophakt dlticola Llet,,6,000 to 8,000 feet, Octoter 21.
aforohim (Catuto.heilus) tai@@i.olo (Alex.), 3,500 to ?,000 fet,
Auglst 30 and 31.
T"efiepoh\o (Monsona\ tuntina .p. tuv., 4,500to 6,000feet, Aueurt
30.

Aler.lNraLet:Tipulide fron' Eastern Asi'', VIII
Gonomlttd \Proso@1nsi\,.onfruentt(Llex,t'

609

3,500 to 5'500 leet, Oc_

Goionaid (Conattuaie)dnse, sp. nov" 3,C00leet, Aueust 29; 2,500to
5,600 feet, October 21 to 26.
Gonmyia \Lipotthlaps) tuowbulosa sp. nov., 3,600leet' Ausust 29'
DdsufuIlottuyie siotuta Brunetti, 3,5'00 let, Octobe! 21'(Enpetl\\ ,tlt@rizo sp. @v., 3,600 feet' Augnst 29
EriateraAlex , 3,600 fet Ausust 29; 3,600 fe€t,
E'!.Jo;turd \EqJ.da) mtn*qie(Ilitid, teitlsentis Llex., 6,600 fet, Octob€r 22'
Eirptetu
Ovrtusia MuLra.oPoald sp. !1o!., 6,500 to ?,800 f@t' Octobe! 20'
Molopl,il@ onsdnB Alqt, 6,600to ?,600leet' August 31
Molophiltus nisritB Alex, 6,600 feet, Ociobe. 22
Stu;rtsotwia ,i&@i3 sp. nov., 2,500 to 3,500 l€eti October 21 to 25'
S&tir;omrio tuibownsis sp. nov, 2,500 to 5,500 f@t, octole! 21
to 25.
TIPULIN,€
DOl,rclIoPEz(NE6oPEzt)

I^RSAI-D^.D rd.

Generalcolorationblack; antenna shod; terminal tarsal segmentssnowy white; wings tinged with bla,ckish,the obliterative
areas restricted; male hypopygium 1fiith the ninth tergite terminating in three blackenedlobes, the median lobe long andslenaler; gonapopllYsesappearing as long' yellow, beaklike structures; eighth sternite ody modemtelyenlarged,the caudalmargin notched and beadng two pale lobes.
Male.-Lerl€:th, dbout 11 millimeters; ving, 12.
Femdle.-Le1|,gth.about 14 millimeters; wing' 13.
Frontal prolongation of head and palpi black. Antenne ofmale much shoder than ia tMstiis; scapal seg:rnentsyellow,
flagellum black; flagellar segmentscylinddcal, segmentsfour to
twelve of nearly equal length, the last selirnent about one_third
the leneth of the penultimate; verticils shorter than the segments. Head yellowish brown, brighter in front, darkening
to bro&'n oD p@terior vertex and occiput.
Thora-\ chiefly brownish black, in the nale with a paler meilian pr€scutal stripe; antedor dorsopleuFl region somewhat
paler. I]altercs obscure yellow, the knobs blackened. Legs
with the coxe blackened, their apices and the trochanters ob_
scure yellow; femora, tibi€e, and basitarsi black, the femoral
bases restrictedly pale; tips of basitarsi and remarninli tarsal
segments white. Wings with a strong blackish tinge, the oval
stigDa darker; a dark seamon anterior cord; obliterativeareas
very restricted, appearing as small areas before the stigma arld
across the fork of M; veirrc black, pale in the obliterative ereaa.
Venation: Medial forks shallow.

510

Tke Ptuiliwitle Journal ol Science

Abdominal tergit€s brownish black, the basal sternites variegated with a broad, subapical, yellow annulus. Male hypopygium (Plate 2, frg, 25) relatively small. Ninth tergite (Plate2, fig, 26) terminating in three blaclelled lobes, the median lobe
slender,a little longer tllan the laterals. Gonapophyses(Platejutting from the genital cham2, fig.27) long and conspicuoLls,
ber as yellow beaklike structures. Eighth stelnite (Plate 2,
fig. 28) with a deep U-shapedemargination that bears fivo small
lobes.
apa]f,(Honshiu).
E a,bite.t.--J
Holotype, male, Shirahone Hot Spdngs, Shinano, July 24,
1929 (M. aeno.) Allotopotype, female, with the type.
(Nesopeza)tars&lba is most closelyallied to D.
Dol,ichdpezo.
(N.\ tarscLis (Alexander), ditredng most corupicuouslyin tbe
short ant€nne of ih€ male and details of structure of the msle
hypopygium.
DOLiCSOPEZ(ORO?EZA)

General coloration brown, the pr€escutumwitll three slightly
darke! subdtidous stripes; head blark; wings tinged with
bro'wn. with three creamy areas, the largest beyond the stigma;
male hypopygiumwith the caudalmargin of the ninth tergite aweakly chitinized pale flange, the median portion further prG
ducedinto a quadrateplate.
Male.-Lenglh, about 12 millimeters; wing, 13.6.
Frontal prolongation of head dark brown, paler laterally; palpi black. Antenne with the scapal segments yellow, flagellar
seg:nentspassing into dark brown; segxnentssubcylilldrical,
the verticils relatively short and unilaterally arranged j terminal
segment about two-thirds the penultimate. I{ead black, spaNely pruinose.
Mesonotum brown, the prescutum with three slightly darker
brcwn, faintly shiny brown stripes; scutal lobesand scutellum
dtrrk brown; postnotum paler, with short yellow set€. Pleura
brownish testaceous. Halteres tinged with dusky. Legs withthe cox@and trochanterstestaceous;remainder of legs paBsing
into brown, the tarsi slightly paler yellow WingE broad, tilged
with brown, the stigma darker brown; cream-coloredobliterative areasbefore and beyondthe stiSrne,the latter very large;a thild obliterative area acloas the base of cell 1st M"; veins
pale brown. Velation: Cell 1st M" relatively long and narrow.
Abdominal segnerts brown, indistinctly variegated withbrownish yellow. Male hypopygium with the ninth tergite

Alexander: TiIru.lidv frorn Edster"n Asiq, VIII

511

(Plate 2, fig. 29) broad, the caudalmargin appeariBgas a pale,
weakly clitinized, narrow border, with a median quadrateextension that is produced into short lateral points; sublateralpoints slenderer;lateral arms nnusuallybroad, the obtusetips
microscopicallyroughened. outer dististyle a relatively shortclavatelobe. Inner dististvle (Plate z, fig. 30) distinctlv bilobed
at apex, the out€r angle being produced into a slender rod, theremainder a broad, smootl! darkened bla.de.
H ebitat.-J apar' ( Honshiu).
Holotype,male,Chichibu,Saitama,Ma.v29, 1919 (R Takltho'slri).
is closestto the species(Oropeza\ saitd.lrrcnsis
Doli,chopezcl''(O.) bi,spi.nala(Alexander), differing in the structure of themale hypopygium, espe{ially the armature of the ni[th tergite'
LI M O NII N , €I-I ONIA (DIACOBOLA)

Allieil to dJ.glrs;wings with scattered brown dots in cells
R and M, in addition to the o€ellatepattern; male hvpopygiumwith the caudalmargirnof the tergite producedinto two blacker]edsetiferouslobes that are separatedby a quadratenotch;
lobe.
ventral dististyle an elongate-oval
Mdle.-Leng|h, 6.5 to ? millimetels; wing, 9 tb 9.5.
Female.-Length, 7.5 to 8 millimeters; wing, 9 to 10.
Rostrum and palpi black, the former about one-half the re_mainaler of head. Antenn€ black, the segmerts with short pale
apical pealicels. Head gray, variegatedwitlt blackish,the antedor vertex more silvery gray.
Pronotum greenish yellow, narrowly darkened laterally. Mesonotal prescutum olive yellow to yellow, the lateral marglns
brown, the median re€ion more infuscated;scutal lobesbrown;
scutellum a.llalpostnotum yellow, sparsely pollinose. Pleura yellow, $'hitish pruinose, with two nallow brown stripes, the more
clo$al exteraling from behind the fore coxe, completely suffusing
the pleurotergite; ventral stripe occupying the ventral sterno_
pleurite; dorsopleuralregion dark, connectedwith the areas oII
the propleura anallateral rnargitnsof prescutum. Halteres black,the extreme base of stem and apical half of knob whitish. Legs
with the cox€eand trochantersyellow; femom bro\\'nishyellow,
the distal end light yellow, inclosinga narrow black subterminal
ring that is nearly equal in width to the yellow apex; in somespecimens,the basal poftion of the femora is clearer yellow;

572

Tke Pknl,iwirre Jounnl ol Science

remainder of legs brovnish yellow, the terminal tarsal segments
passing into brownish black. Wings (Plate 1, fig. 1) yellortr'
ir addiwith a brown ocellate pattern that is much as in 019?1s,'
R
and
M,
these
iackingtion, with scatteredbrown dots in cells
ln argua,
Abdominal tergites brown, the extreme caudalmargins pale,in caseBwith a more yellow basal and medial fing on each segment; sternitesgxeenishyellow, the caudalmargins of the segments narrowly dark brown; hypopygiumyellow. Male hypopygium (Plate 2, 69. 31) with the tergite, 9t, narrowed apically, each laieral angle produced into a dusky setiferous lobe,
these separated by a quadrate notch, Ventromesal lobe ofbasiswle,b, very stout, occupyingslmost the entire face of thestyle. Ventral dististyle, t,d, an elongate-oval pale lobe that is
approximately twice as long as the dorsal dististyle; rostralprolongationslender,darkened,the two pale peglikespinesarisingi from face of style above. Dorsal dististyle lvith the surface microscopicallyroughened, Gonapophyses,g, with the
apex of mesal apical lobe bluntly obtuse.
Habitat,-Formosa.
Holotype,male, Ilassensan,altitude 6,500to 7,500feet, Aue]lst 31, 1929 (S. lss't,tt). Allotopotype,female, altitude ?,500
feet, October 24, 1929 (S. lssifti). Paratopotypes,2 males;paratype,1 fernale,Shdrci, altitude ?,00010 8,000feet, October
25, 1928 (S. lssiAi;).
Limonia (Discobol&) taiaanetrl&i.E separatedfrom t. (r.)
argra (Say) chiefly by the vely difierent structure of the malehypopygium. The female from Shorei had earlier b€en recorded as o?g?,s.: The Arisan record for orgfl.s in the preceding part under this general title 3 is correct, and there are unquestionablythree speci€sol Discoboklinhabiting the higher
mountainsof Formosa.
(LIrIONIA)',IMON'A

Gen€ral coloration ol thorax reddish yellow, unmarked; head
blackish gray; halteres relatively short, brown; legs yellow, thefemoral and tibial tips narrowly blackened; wings gray, darker
basally, stigma brown; rlarrow'dark seams along cord and ou'uer
end of cell 1st M"; male llvpopygium lvith the tergite deeply
notched; dolsal dististyle lacking; rostml prolongation of ven'Philir. Joun. Sci. 40 (1929) 626.
" Pbilip. Jouh. Sci.42 (1930) 509.

Al,extndet: Tipulil.e lrom Edstehr Asia, VIII

513

tral dististyle long and slender; a spine arising from a long
basal tubercle on face of dististyle near base; tergal valves ofovipositor very small.
Mdle.-Lerrgth, about 7.5 to 8 millimeters; \\inq, 7 lo 7.4.
Female.-Lengtb about 8.5 to 9 millimeters; wing, 7.5 toRostrum shiny black; palpi brownish black. Antennrelightbrown; flagellar septnentsoval, becorningmore slender andelongateoutt'ardly, the vedicils a little exceedingthe segments;
terminal segmenta trifle shoder than the penultimate, Headblackish gtay.
Pronotum reddish yellow, more blackenedanteriorly above.
Thorax uniformly reddish yellow, the surface nitidous, thepleura cleareryellow. Halteresrelatively short,brown, the baseof the stem yellorr. Legs yellow, the femoral tips narrowly but
conspicuouslyblackened;tibize more narrowly darkened; tarsi
gradually darkened. Wings (Plate 1, fig, 2) slightly tinged
with gray, the basal cells somewhatmore strongly so; cells C
and Sc more yellowisb;stigma brown; narrow dark seamsalong
cord and outer end of cell 1st M,; \'eins black. Venation: Sc,
ending oppositeor just beyondmidlength of Rs, Sc" at its tip;
free tip of Sc, snd R. about in alignment; m-cu at or closeto
the fork ol M.
Abdominaltergites dalk brown, the caudalmargins narrowly
pale; sternites and hypopygium paler, Male hypopygium(Plat€ 2, fig. 32) witl the tergite, 9t, deeply notched medially, ihe conspicuous lateral lobes with long conspicuous s€te.
Basistyle, b, with the ventromesallobe conspicuous, Ventral
dististyle, ?d, large and ieshy, much larger tban the basistyle;rostral prolongationlong and slender,cur.ved;a single powerJulapinearfuesfrom the face of the ventral dististyle near baseof
prolongation,this spine from a longer basal tubercle. No dorg, with the mesal apical angle
sal disiistyle. Gonapophyses,
elongate,gradually expandedat tip into s weak spatula. Ovipositor with the tergal valves (cerci) very small.
Eabitat.-Fotmosa,Holotype,male, Ilassensan,a.ltitude4,500to 6,000feet, August 30, 1929 (S. lssti"r). AllotopotlTe,female. Paratopotype€1 male, 1 female,
Litnoni& kor.inga is named from Koxinga, piratic lord of
Taiwan in the seventeenth century. The species is very different from L. (L.\ alticokL (Edwards) in the details of struc-

514

Tke Pkili.ppi,ne Journal of Science

ture though very similar in the reducedtergal valves of theovipositor.
LIl[ol{IA(LIDNOTE6) SASSENINA@ noY.

Femal,e,-Lengt}',about 7 millineters; wing, 7.5.
Generallysimilar and closelyallied to ,. (L ) ti1)erai Alex'
ander (Luzon), difierilrg especiallyin the details of venation.
General coloration dark blackish gray. Halteres darkened,
the baseof the stem bdghtened. Legs with the femora obscureyellow, scarcelydarkenedat tips. Wings (Plate 1, fig. 3) rtith
Rs long and gently arcuated; Sc ending oppositer-m analthus
appea ng very long, the distancebetweenotigin of Rs and Sc,
Bubequalto Rr+e and e{ceedingR, alonei free tip of Sc, somedistance before R,, the latter arcuate. In the absence of themale, the length of ihe costal fringe in this sex cannot be stated.
Ilabitar.-F'ormo,sa.
HolotlTe, female, Ilassensan,altitude 4,500 to 6,000 feet,
Ausust 30, 1929 (S. lssi&i).
LIMONI^(NEPIDIA)

General coloration gray; artenne with eight flagellar segments, eachb€aring two branches; terminal segment oval ; wings
gray, variegated with white spots and dots, the cubital andanal fields more uniformly gray; Sc, ending about opposite onethird the length of Rs; male hypopygium with three long spines
on rostral prolongation of ventral dististyle.
Male.-Lengih, about 6 millimeters; wing, 5.5.
Rostrum and palpi black. Antenne with flagellar seglnent on€
merely produced beneath, segme[ts two to nine inclusive withtwo branches, the lo[gest about twice the segments; dagella,rsegxnentsten and eleven merek produc€d; terminal Begment
relatively short, oval; basal enlargementsand branchesdark,
the long glabrous apica.l necks pale, these necks shortening onouter segments, on the penultimate and antepenultimate very
short. Head brownish gray.
I'hora-x g?ey, the prescutum with a median brcwn stripe, thelateral stripes lacking or nearly so. Ealteres pale, the knobs
weskly infuscated. LegF brown, the segxnentsnot conspicuously
variegated. Wirgs (Plate 1, fig.4) gray, variegated with whitespots and dots, most evident in the ra.dial and medial fields, thecubital and anal fieldsalmost uniformly darkened;veins brownish black. Venation: Sq ending about oppositeone-third thelength of R9, Sc. not far Jrom its tip; m-cu before the fork of M.

Aleratul.er: Tipll.lide f.ronl Edatern Asid, VIII

515

Male hypopygium (Plate 2, fig. 33) much ss in m.e,culo,td;
rostra.l prolongation oJ ventral dististyle, ,d, unusually iongand slender,at near midlength with three long reddish Bpine,sfrom a restricted point, these spines a little shoder than the
prolongation.
Habitat.-Formosa,HolotlTe, male, flassensan,altitude 4,500to 6,000feet, August 30, 1929 (S. lssi/d).
Li"nonia (Rhipid,i&) triltnt1&ta is evidently closely allied toL, (R,) ltulcuklta (Meigen) and may prove to be a geogTaphic
race of the latter, I have seena closelyallied speciesor raeefrom Szechwan,China.
LrlroNra (DrcRixo[Yr^)3

Allied to ptnctulatq,; wings without spots in costal and subcostal cells exceptat ends of cells; male hypopygiumwith therostral prolongation of the ventral dbtistyle elongate, with two
small basal spines; gonapophyses
simple at tips.
Male,-Length, about 6 to 5.3 millimeters; wing, 6.
Female---"langt\ about 6.6 to 7 millimeters; wing, 6.5.
Ro,strumand palpi black. Antennreblack; flagellar segments
oval, gradually decreasingin size outwardly. Head dark gmy;
anterior vertex narrow.
General coloration of thorax gray, the pr€scutum with abroad median brown stripe and less distinct lateral stripes;
scutellum light gray. I{alteres dusky, the knobs dirty white.
Legs with the cox€edark brown, pruinose; trochanters brown;
femora gradually deepeningto dark brown, the extreme tips
narrowly obscureyellow; tibire and tarsi brownish yellow, thelatter blackenedat tips. Wings (Plate 1, fig. 5) cream-colored,'with a restdcted brown pattern, including spots at arculus; tipof Sc; R ; at intervals along cord ; outer end of cell lst M, andtip of R";" small spotsat two-thirds the length of R{+6,midlengthof distal sectionof Ml+r, midlength of M, and tw'o spots in celllst A adjoining vein 2d A; cells C a.ndSc without darkening
except at each end of cells; veins yellow, dark browl in theinfuscatedareas. Venation: Sc, ending oppositeorigin of Bs,
Sc, at tip; Rs straight, oblique; cell lst M, elongale; m-cu atfork of M.
Abdomen dark grey, the caudal margins of the segments
paler; basal sternites obscureyellow; hypopygiumdark. Male
hypopygium (Plate 2, fig. 34) with the lateral lobesof the ter-

516

lke Pkilippine Journatr ol Science

gite, 9t, low and obtuse, with numeious set€. Basistyle, b,
relatii.ely small, the ventromesallobe large. Ventral dististyle,r)d,a large fleshylobe,the rostral prolongationlong and slender,at base iyith two small subequalspites that are Iess than onehalf as long as the prolongation. Dorsal dististyle a gentlv
curved chitinized rod, narowed to the acute tip. Gonapophyses,g, with the mesalapical argle a simpleacutespine.
Habitat.-I'ormosa.
altitude
Holotype,male,Meizi Hot Spdngs,foot of Hassensan,
2,600feet, October25, 1929 (S. Issii:i). Allotopotype,female,
October26, 1929 (S. lssi/rd). Paratopotypes,20 males and females, with the type; paratypes, 1 male, Hassensan,altitude3,500feet, October21, 1929 (s. lssd/ci); 1 mele, Noka, altitude
8,000feet, June 26, 7927 (5. I$iki\.
Lirlonial (Dicranon4Jia)subptlnchllatabelongsto the p?rzctukfta. grotrp and has been confused \r iih punctulata (de Meijerc). The latter is ngured by de Meijere as having a single
rostral spine $ith its tip shongly curved; L, (D,\ fltllolo(Iyi(Alexander) has the rostral spine single, entirely straight, aswithlong as the prolongationitself or loDger. Gonapophyses
the mesalapical angle blackened,broad, more or lesstoothed.
Generalcoloration reddish brown; rostrum black, the apicalfoudh yellow; halteres dusky; legs yellow; wings whitish hyaiine with a heavy brown, chiefly costal pattern, the outermost
area a completefascia; male hypopygiumwith the two rostralspinesadsing from a long slendercommontubercle.
Md,Ie,-Length,excludingrostrum, about 6 millimeters; wing,
6.8: rostrum. about3.
Rostrumblack,the apicalfourth paling to yellow; palpi bleck.
Antenne with the scapa.lsegmentsblack; flagellumpale bro\r'n;ffagellar segmentswith sholt inconspicuousvedicils. Headgray, the antedor vertex more silvery, the posterior vertex more
blackish.
Mesonotalprescutum brown, with four reddishbro\,i'nstripes,
the lateral podions more pruinose; scutum and scutellumprui'nose,eachscufal lobe with an elongate-tdangularreddish brown
area; postnotumpale brown, the surface pruinose. Pleura testaceousyellow. Halteres dusky, the base of the stem restrictedly yellow. Legs with the coxeeand trochantersyellowishtestaceous; remainder of leg:syellow, only the terminal tarsal

Alem,nd.er:TipltlidtElrom Ed.sternAsia, VnI

El7

segmentsinfumed. Wings (Plate 1, fig. 6) whitish hyaline,
the costal region pale yellow; a heavy brol!,n, chiefly costal
pattern; six major costalar.easthat extendinto the cellsbehind.
the ihird ar origin ot Rs,lherast a comple{e.
lj"ansverne.
nearlyapical fascia; betweenthe major areas in cells C and Sc aresmallerdark spotsthat restrict the gaoundcolor to small areas:
additionalresrriciedbrown areasalongcord and outer cnil ofcell lst M" and as marSjnal cloudsat ends of longitudinal veins
and at midlength of cell 2d A; veins yellowish brotvn, alarkel
in the infuscated areas. Venation: Sc long, Sc, oppositethe
fork of Rs, Sq at its tip; a supernumerarycrossveinin cell Sc;
In-cu at fork of M; vein 2d A short, strongly culvealto nurgin,
the cell nide.
Abdominal tergites reddish brown, more blackenealmecliallv
and subapically.the caudal margin narro$ly pale; srFrnites
more uniformly yellow; hypopygium brownish vello\y. Male
hypopygium (Plate 2, fig. 35) with the tergite, 9r, transverse.
the caudalmargin verl' gently emarginat€. Basistyle,b, muchsmaller than the ventral disfistyle, the ventromesallobe larce.
Ventral dististyle,/d. ovat. the rostral regionIarge.prod.rcoa
into a very loDg tuberclethat bearstwo slightly longer redtlish
spines. Dorsal dististyle a strongly cur-v.eal
sickle. Go[aDo_physes,g. wil h rhemesalapicalanglea sfronglycurvedsubacutc
spine.
Ilarir&r.-Formosa.
Holotype,male, Shinten, December3, 1928 (S. Lesi&i).
LimonirL (GemnomlJia)&picifa.scidtais very difierent fromallied regionalspeciesof the subgenus.
ANTOCIII(ANTOCHA)

General coloration dark gyay; halteres ancl legs blackeneal;wings tinged with blackish; mate hypopygiumblack, the outerdististyle pointed at apex; €edeagus
broad.
Mal,e.-Leagth, about 4.5 to 6 millimeters; win& b to 5.b.
Rostrum and palpi black. Antennarblack; flagellar segments
oval, gradually decreasingin size outwarally,the terminil ses_rnen{aboul onerhird longer Ihan lhp penultimate;verliciL
very short, more conspicuousort basal analtwo out€r segments.
Ilead dark gray.
Mesonotumdark gray, the prescutum almost coveredby still
darker conflueni stripes. Halteres infuscateal,especiallytheknobs. Legs black, Wings (plate 1, fig. ?) with a sirong

Tke Pkilippine Journal of Sci,ence518

blackishtinge; stigma s little darker; veins still darker brown.
Venation: Sc, ending a short distancebefore the fork of thelong Rs; cell 1st M" closed.
Abdomenblackish gray, the hypopygiumblack. Malc hypopygium (Plate 2, fig. 36) with the cauda.lmargin of the tergite,
9r, gently emarginate. Surface of basistyle, b, with very
numerous course sete. Outer dististyle chitinized, curved
gently to an acutepoint, the face carinate. Inner distbtyle velT
strongly curved. .Adeagus,q broad.
l1abiror.-tr'ofmosa.
Holotype,male,Meizi Hot Springs,foot of Hassensan,
altitude2,500 feet, October 26, 1929 (S. lsst,ti). Paratopotypes,3males,October25, 1929 (S. lssi/rd); paratypes,2 males,Hassensan, altitude 3,500feet, October24, 1929 (S. lssiki).
Antochast1fi ia readilytold from all describedregionalspecies
by the diaglostic features listed above.
IIDiIUS(IUREAPEIDIA)

F.'nnle.-Length, about5.5 millimeters; $iDg, 5.5.
Alexander,diffedng especiallyin
Allied to ti. (8,, inetregans
the venational details and pattern of the legs.
Mesonotalpr:escutumwith the disk ulmost coveredby three
confluent browtl stripes, restricting the ground color to thehumeral and lateral podions; scutal lobesdark brown, Femorawith the tips rather broadly and conspicuouslystowy white;tibial basesnarrowly pale, the tips very broadly white; tarsi
white, the terminal tarsal segmentsdalkened. Wings (Plate 1,
fig. 8) with Sc short, Se, ending oppositer-m, Sq a little longer
than Sc, ; Rs short and sJraight; m-cu before the fork of M.
Iloritof .-Formosa.
Holotype, female, Hassensan,altitude 6,000 to 7,000 feet,
Ausust 30, 1929 (S. Isst4t).
Generalcolorationof thorax brown, the prcescutumlvith fourns,rrow shiny black st pes, the scutal lobesfurther variegatedwith similar areas; legs yellow; wings gray, the prearcular andcostal regions clear yellow; stigma elongate,brown; veins yellow, narrowly bordered on the membrane by the same color,
Male.-Length, 8.5 to 9.5 miilimeters; wing, 9 to 10.5.
Rostrum brownish gaay; palpi dark brown. Antenne reddish brown, the fiIst segmentdarker at baBe; basal flagellar
segmentswith long conspicuousverticils, thsse sholter on theouter segiments. Heed dark brownish gaay,Aleiand,er: Tipltl le lrom Eoatern Asin, V I

b1g

Mesonotal pfescutum grayish brown to brown, more grayish
laterally, lffitt four narrow and incompleteshiny black stripes,
the intermediate pair longer and broader; scutum brownish
gray, eachlobe with two shiny black areas; scutellumbrownish
gray, the postnotal mediotergite clearer gray. Pleura gr.ay.
Halteres yellow, the knobs a little dusky. Legs with the coxe
and trochantersbrownish yellow; remainderof legs clearer yellovr, with dark sete, the outer talsal segmentsdarke4ed. Wings(Plate 1, fig. 9) gray, the prearculsr and costal reg:ionsclear
yellow; stigma elongate,derk brown; veins yellow, narrowly
bordered on membraneby clear yellow. Venation: R,s angulated and short-spurredat origin, longer than cell lst M,; m-cu
at before midlength of the latter.
Abdominaltergites obscureyellow,with a dorsomedianbrownline; sternites clearer yellon; a conspicuoussubterminalblackish gray ring involving segmentssix to eight; hypopygium
fulvous. Male hypopygium (Plate 2, fig. 3?) with the tergite
transverse,the Yentrolateralangl€s producedcaudad into spatulate pale blades (lateral processes). Outer diBtiswle, od,
terminating in a slenderblack point, the suface with abundant
erect yellow setul€, the disk and inner margin with longer
recurved se&e. Inrer dististyle, id, longer, set with spinouR
seta. Gonapophysessmali, slender, shorter then the bifid€edeagus,@.
H@bitat.-F ormosa.
Holotype,male Hassensan,altifude 3,500to 6,500feet, October 22, \929 (S. lssi&i). ParatopotlTe, male.
I ta.ke great pleasure in naming this bea]ulifnl Dicraneptltchcl
in honor of the collector,Prof. Syuti Issiki, distinguishedstudentof the Mecopteraof easternAsia. The speciesis very distinct
from all other membersof the genusso far discovereal
in eastern
Asia.
DICE-{NOTA (AMTLOPINTI)

General coloration pale yellow, including the legs a4alhalteres;$ings cream yellov, with a conspicuousbrown pattern that
includesthe prearcular cells and darkenedcosta as far alistsal
as the origin of Rs; supernumerarycrossveinsin cells R ancl
R : cell lqt N. closed.
Male.-Length, about 6 millimeters; wing, ?.
Rostrum and palpi brown. Antenne l4-segmented; basal
segmentblack, the remaining segmentspale testsceousbrown.
Head ochefous, the vertex a little darkened.

520

The Pkitri,ppineJournnJ of Science

Mesonotal pr.:escutumpa.le yellow with a whitish bloom;
scutal lobesmore darkened; scutellumand postnotumpale yellow. with a whitish bloom. Pleura pale yellolY. Halteresyellow. Legs yello*-, the two terminal tarsal segEents brown.
Wings (Plate l, fig. 10) pale creamyellow, with a conspicuous
brown pattern; prearcular region and cells C end Sc darkened
to approximately opposite the origin of R,s; base of cell R
darkened to opposite Sc,; stigmal regirondifiusely darkened;
nanow but conspicuous
dark brown seamsat origin of Rs, along
cord and outer end of cell 1st M", on Rz and the supernumerarycrossveinin cell R", at the tips of veilrs Rr1, and Rr, snd atfork of Mr. r; veinsyellow,brown ia the infuscatedareas. Venacrossveinin cell R,; Rs stroruily arcuatedtion: A supernumerarx.
crossveinin cell Rs more tl1an its
asupernumeraryat origin;
1st
M" closed.
beyondR,;cell
own Iengthyellow, ringed caudallywith
thesegments
Abdomenrvith
basal
on
the
outer segments;outer segbrown, the amountincreasing
more
uniformly dark brownments, including the hypopygium,
Habdto"t.-Forlr.'oaa.
Holotype, male, Hassensan,altitude 6,000 to 8,000 feet, October 23, 1929 (S. Issikd).
Dicranota (Amarlopina,d.electdtais most nearly allied to ,.
(A.) dioanotoides (Alexandel) and ,. (A.) sibidca (Alexander), difiering in the wing pattern and venation.
Generalcoloration of notum ocherous.marked 1^'ithbrown;knobs of halteresinfuseated;legs yellow, the femoral tips conspicuouslyblackened;wings $'ith the costalthird cream-colored,
darkened;a sparsebuf heavy wing
the remainderconspicuously
pattern; abundant macrotrichia in cells of wing beyond cord:R, far before fork of Rsra;cell M, present.
Femal..-Length, about 5.5 millimeters; wing, 6.
Rostrumreddishbro{'n; palpi black. Antenne 16-segmented,
black, the flagellar segmentsa little paler; flagellar segments
becoming more slender and elongated; \'erticils conspicuous.
Head ltght brown.
Pronotum dark br'own, palef behind. Mesonotumocherous
to reddish brown, the praescutumwith a more or less distinct
mcdianbrolvn st pe; scutallobesdarkened;scutellumand postnotum dark bro$n. Pleura chiefly dark bro\'!"lLthe sternopleurite, meral region, and pleurotergitemore yellowish. Hd-41.1

Alerander: TipuUd,e fron, Eo,stetn Asia, VIII

52L

teres pale, the knobs infuscated. Legs with the fore cox@infuscated,the remaining coxreand irochante$ yellow; femora
yellow, the tips narrowly blackened; tibia yellow, the tips very
narrowly blackened;tarsi yellow, passing into brown; tibial
spurs distinct. Wings (Plate 1, flg. 11) with the costal third
cream-colored, the central and posterior thirds darkened, theanal cells again somewhat more yello\irlsh; a heavy dark brownpattern, srrsnged as follows: Origin of Rs; stigma; along cordand outer end of cell 1st Mr; veins yellow, more irfuscated ill
the darkenedregions. Abundant macrotrichia in the cells oft}le wing beyond the cord, Venation: Sc, ending nearly oppositethe fork of Rs, Sc, someilistancefrom its tip; Rs angulated and spurred at origin; R3 more than one-half its length
beJorethe fork of Ri+a; cell M, present; m-cu at near midlength of cell 1st M,; vein 2d A nearly stmight.
Abdominal tergites dark brown; sternitesbrown basally,thecaudalhalf obscureyellow, the amount of the latter decreasing
on the outer segments. Ovipositor with the basal shields blackened; valves yellow, the sternal valvesblackenedventrally.
Eabir,rr.-Fomosa.
Holotype,female, Hassensan,altitude 5,600feet, October22,
1929 (S. lssi&d). Paratopotype,female.
Adelpkowia issikilw is another of the very distinct speciesof
crane flies discovered by the collector in the mountains of Formosa, I take great pleasure in dedicating the present fly to
Professor Issiki who has done so much toward making known
the dch tipulid fauna of the island. The presentspeciesis very
distinct from all regional forms. I w'ouldbelievethat On?d?.ec{s de MeijcrA is identicalwith AdetphonA;n,despitcjh; implied lack of tibial spurs,a highly variable characterin tbis, aswell as other groups of Tipulide.
LII(NOPHTLI(PIUONOL]|BIS) SERRTDENTTIiI D, nut.

Generalcolorationblack, the surface opaqueby a sparsepruinosity; wings grayish with vagueseamson the crossveinsand
deflections;Sc, much longer thsn Sc,; Rr-3 from one-half to
two-thirds R.; cell M, lacking; m short and straight, less thanthe basal sectionof M,; male hypopygiumwith the gonapophy6esserrate along outer margin.
' Tijd. c hr . \ oor Er ionologje 5 6 ( l D t 3 , 3 5 0 3 6 1 .

522

The Phitriwi,ne Journal, of Science

Ma.Ie.-Lengl}].,about4.3 to 5 millimeters; wing, 5 to 6.2.
Female,-Length, about 4.8 to 6.2 millimeters; wing, 5.5 to 6.
Rostrum and palpi black. Antenne blerk throughout; flagellar segEents o!'al, gredually decreasirg in size outwardly, theterminal se€tnent little larger thsn the penultimate; longest
verticils unilatemlly arranged, exceedingthe segmentsin length.
Headblack,pruinGe.
Thor&x blsck, the surface pruinose,least so on the medianlegion of prEscutum. Halteres pal€ yellow, in caseswith tle
k[obs weakly infuscated. Legs vith the fore coxe darkened,
the remaining coxe and all bochanters yellow; femora yellow,
the tips blackened;tibie and basitarsi similar, the tips morenarowly blackened;remainder of tersi black; Iegt conspictrously hsir?. Wings (Plate 1, fig. 12) gaeyish,the stigma andvague seamsat origin of Rs, along cord and outer end of cell
1st M" slightly darker; veins light brown. Venation: Scj ending shortly before fork of Rs, Sc" somedistsnce from its tip;
R, subequalto Rr*r; Rr*, approximatelyone-half to two-thirds
R. alone; cell I{, lacking; m straight, transverse,shorter thanl,he arcuatedbasal sectionof M.; m-cu closeto or before midlength of cell 1st M..
AHomen black, including the hypopygium,the surface moreor less pruinose. Male hypopygium (Plate 3, fig. 38) with theinner dististyle, i{t, dilated at base, the outer surface with numerouserect sete, the apex suddenlynarowed into a blackened
point. Gonapophyses,
g, appearingas slenderblackenedplates,
the outer margin conspicuouslyserrate, -ZEdeaels,o, narrow,gently curved.
Ho,bitoit,-Formosa,Holotype,male, Hassensan,altitude 6,500to 7,800 feet, October 24, 1929 (S, /ssifti). Allotopotrue, female. Paratopotypes,16 malesaDdfemales,altitude 3,500to 8,000feet, October
22 to 24, 7929 (S. Issiftt).
LinoLophikt (PrionolaLbb) sen tl.entat& is obviously closely
allied to ,. (P., IdponeuraAlexander and L. (P,) tripopkleps
Alexander, ol Kiushiu, Japan, difiedng most evidently in the
structure of the male hypopygium. The somewhatsimilar Z.
ni.gronitidnEd$ards, likewise from the high mountainsof Formosa,difiers in the polishedblack thoracienotum and in a number of important venationalcharacters,as the position of Sc,,the subequalR2€, and the long obliquem.

Alerdnd,el': Titpa.l,id,alrom Eastern Asid, VIII

523

(MONCOIIA) MONTTN^ 3p. nov,

General coloration dark brown; legs black, the tips ol thetibite and the tarsi paling to yellow; wings tinged with dusky;
R. shortly beforefork of R a; fusion of Cu, and 1st A slight.
Mal,e,*Lengtb, about 6 millimeters; wing, 6.5.
Femdle.-Length, about ? millimeters; wing, 6.4.
Rostrum and palpi dark brown. Antenne black; flagellar
segmentswith vedicils of moderateleDgth. Head black, very
spalsely Pruinose.
Mesonotumdark brown, the posterior margin of th€ scutellum
and posteriorhalf of the postnotalmediotergitemore yellowish.
Pleura yellowish brown, the dorsopleuralregion more blackish.
Halteresbrownish black, the baseof the stem rest ctedly pale.
Legs with the fore coxe dark brown, the remaining coxe moreyellowish brown; trochanters obscure yellow; femora black;tibie black,the tips paling to dirty yello.lv;tarsi yellow. Wings(Plate 1, fig. 13) with a strong dusky tinge, the stigma darker
but small and ill-defined;veins broNnish btack. Venation: R"
shortly before fork of R3ra ; m-cu at or just before the fork of
M; fusion of Cu, and 1st A slight.
Abdomen blownish black, the sternites paler, especiallyin
the male.
I{abirnr.*Formosa.
Holotype,ma1e,Hassensan,altitude 4,500to 6,000feet, August 30, 1929 (S. lssift). Allotopotype,female.
It is probable that the preseBtspecieswill be found to bea characteristicmountain form. It was associatedwith typicalPal€arctic crane flies, as Tricaf)lLon&
fofmosa?roAlexanderandDi,cranota(Amalopine) gi,bbera(Alexanaler),var.
coNoMYIA (PnOCONOrlYrl) CONFLUENIA (lr*.nd..),Gnophotwia conflLenta AtExANDm, Ann. Ent. Soc. Ahedca
69 .

17 1192!r)Two malesfrom Hassensan,altitude 3,500to 5,500feet. October 22, 1929 (S. Issifti). The venation (Plate 1, fig. 14)
showsa long Sc, Sc, ending opposjtethe fork of Rs or nearlyso, Sc, at near middistancebetweenorigin of Rs and tip ofSc,; cell R. relatively d€ep; m-cu at or closeto the fork of M.
The male hypopygium (Ptate 3, fig. 39) has the outer lobeof the basistyle,b, stout. Three dististyles,the outer a slenaler
curved rod from an enlar.gedbase; seconddististyle bifid at

524

Tke Pki.lippdke JoutnaL of Scianceapex,the stem with erect conspicuoussete; irner style simple,o, compressed.
stouterthan the first, the apexobtuse. .4Edeagus,
The speciesis very diffelent from G. (P.) scutellxlltu-dlbun
Alexander, likewise from ihe Formosan mountaiBs, in the uniformly black coloraiion and very different male hypopygium.
CONOMYIA (CONOIIYIA)

Generalcolorationdark brown; rostrum and antenne black;headdark gray; pleura yellow,more or lessdistiBctlyvadegatedwith browr; halteresdusky; legsbrown; wings $ay, the stigrnal
region more infuscated; male hypopygilum'with the outer dististyle a slendersetiferousrod; phallosomewith three spinous
points.
Male,-Length, about 3.5 to 4 millimeters; wirg, 4.5 to 5.
Feualle.-Length, about 4.5 to 4.8 millimeters; wing, 5 to 5.4.
Rostrum and palpi black. Antenne black throughout, theouter flageller segmentsvery slender. Head dark gray.
Pronotum and anterior latelal pretergites light sulphur yelIow, Mesonotumdark brown, very sparselypruinose,the lateral and humeral rcgions of the prescutum yellow; scutellum
light sulphur yellow; postnotalmediotergitegray. Pleura yellow, with more or less distinct darkenedareas on the anepisternum and on ventral sternopleurite. Haltereselongate,dusky,
the extremebaseof stem yello{'. Legs lvith the coxE}brownish
trochanters yeilow; remainder of legs pale brown,te,staceous;the outer tarsal selimentsdeepeningto black. Wings (Plate 1,
fig. 15) eaay, the stig]nal region more infuscated; veins darker
brown. Venation: Sc ending just beyond origin of Rs, Scrcloseto its tip; eell 1st M, closed;m-cu at or closeto fork of M.
Abdominal tergites brownish black, the sternites more yellowish. Xlalehypopygium (Plate 3, fig. 40) \vith the outer dististyle a slendersetiferousrod. Inner dististyle, iir, a flatteneal
chiiinized plate, the outer lateral abgle producedinto a curved
blackenedspine, the inner lateral angle less produced; mesal
margin pr-oducedinto a fleshy setiferouslobe. Phallosome,p,with three blackenedspines,two apparently borne by an api>
physalstructure,the third at apexof adeagus.
I1dblrat.-Formosa.
Holotype, male, Hassensan,altitude 5,600 feet, October 22,
1929 (S. lsstki:). Allotopotype,female. Paratopotypes,1 male,
altitude 3,600feet, August 29, 1929; 4 males and fenales, altitude 2,500to 5,600feet, Oetober21 to 25, 1929 (S. lsstkt).

Al,eno,kiLer:TipulidE tronx EestenL Asin, VII|

625

The specific lratne, nMLsei,is bhat of a local Formosan tribe,
The fly fu distinguishedfrom all similar regional speciesby the
structure of the male hypopygium.
coNoMYIl(LFOPELEPB)SAUTEFI 3!. noy.

Generalcolorationdark brown; rostrum, palpi, and antennE
black; headgra.y;pleura dark, with a silvery white longitudinal
stripe; halteres d$lry; wings unmarked except for a ditruse
stigma; Sc long; male hypopygium with two dististyles, theouter terminating in an acute blackened point.
Male.-LengLh, about 3.3 millimeters; wing, 4.1,
Rostrum and palpi brori'nishblack. Antenne black throughout; flagellar segmentselongate, clothed with a long erect whitepubescence,in addition to the longer unilaterally arranged verticils. Head dark gray, with a small yellow area on the posterior veftex.
Pronotum and antedor lateral pretergit€s light sulphur
yellow. Mesonotalprrescutumand scutal lobes uniformly darkgaayish brown; median region of scutum snd the scutellum e).cept for a basal darkeniDg obscure yellow; postnotum dark,heavily pruinose, the arterolateral portions more yellowish.
Pleura dark, with a broad silvery white longitudinal stripe,
extending from and including the fore cox€e,reaching the abdomen; pleurotergite chiefly yellow. Halteres dusky, including the knobs. Legs with the fore coxzeas described,t}le othercoxe darker, their tips pa.le;troclEnters testaceousyellow; femora brownish yellow, passinginto darker brown at tips; tibia
and tarsi uniform brown. Wings (Plate 1, fig. 16) with abro{'nish tinge, the prearcular and costal regions more yellowish; the very difiuse stigma a little darker than ihe groundcolor; veins brown. yenation : Sc long, Sq extendingto shodly
before midlengthof the long Rs, Sc, somedistancefrom its tip;branchesof Rs strongly divergent; R, and Mr+, deflectedtoward
one another at margin, narrowing the cell; cell 1st M, narrowedat base.
Abdominal tergites unifonnly dark brown, the hypopygium
more yellowish. Male h]?opygium (Plate 3, fig. 41) {,ith two
dististyles, tbe oLrter,ad, a slender rod that gra.dually narrows
to an acuteblackenedpoint, the baseof the latter with rumer.ous
sete. Inner dististyle a small pale lobe ihat is a little shoderthan the apical lobeof the basistyle,terminating in a fasciculate
bristle and with a.very long slendersetaon outer margin beyond

526

Th? Ph;Iippinc Journat of Sc;Pncr

midlength. Phallosome,p, appearing as a broad pale plate,the surface laterally with abundant microscopic setulE, the tip
proaluceal
into a glabrouspoftion; two additional elongaterods,
the shorter from an enlarged baae.
I{abr:ror.-Fo}mosa.
Holotype,ma1e,Daitotei, April 1914 (H. Sarterr 'MuType in the collectionof the DeutschesEntomologisches
seum.
Gonomyia(Lipophleps) satderi is named in honor of Mr' H'
Sauter. well-knorivn collector of Formosan insects. The species
is very tlistinct from G. (r.) LongiradialisAlexander (Luzon)
and G. (.L.) s&us?iAlexander (eastem Australia) in the structure of the male hypopygium. This is very probably the spe_
ciesrecordedas .s/cziei(gracildsSkuse,preoccupied)by Riedel'from Macuyama,Formosa.
coNo

Ylr\ (LIPoPEiEPa)

Generalcolorationclark brownish gray;pleura dark, with ayellowish white longitudinal stripe; femora browrish black;wings gray, variegateilwith brownish gray; costal region paleyellow; abalomengrayish black, the caudal margins of the seg_ments narrowly Yellow.
Femd,Ie,-Lengtb,about 4.5 millimeteN; $ing' 4.1.
Rostrum and palpi black. Antenne with the scapal segmenta
obscureorange; flagellumbtack. Head orange,tle disk ol thevertex darkened.
Pronotum anal arltedor lateral pretergites light yellow' theformer alarkeneallaterally. Mesonotumdark brownish g?ay, thecaualalmargin of the scutellumbroadly yellow; postnoial mediotergite pale, with a little less than the distat half darkened
Pleum alark, with a broad, yellowish white, longitudinal stdpe
exienaling from anal including the fore coxe, passing beneaththe halteresto the abalomen;dolsal pleurites brown, the ventral
sclerites more bluish gray. Halteres with the stem dusky atbase,lhe distalhalf yellow;knobsdu{ky.the endsconspicuously
yellow. Legs with ihe fore coxe yellow; remaining coxresimi_
femora brownishblack beyond
iar, the basesnarrowly alsrkeneal;
the base, the odreme tips on lower sur{ate a little brightened;tibiE and tarsi bro\,!'nishblack. Wings (I'late 1, fig. 17) *'ith
the grounalcolor gray, variegatedwith darker brownish grayclouJs; cel18C and Sc pale yeilow; stigrra not darker than the(191?) 112'
"Archiv lnr Naiuqeschichte s2 for 1916,Abteil A

4.,1

Alennd.er: Tipuli.de trom Eastetn Asia, VI(I527

remainingdark areas; darkenedcloudsoccupyingmost of radial
field, exceptan area in outer end of cell R; other fields of wingmerely skeaked with dark; veins brown, Sc yellow. Venation:
Sc, ending some distancebefore origin of Rs, the latter veinsinuousat end; anterior branchof Rs relativelylong and straight,
cell R, being nearly parallel; m-cu closeto fork of M.
Abdomengrayish black,tle caudalmargins of the segments
narrowly yellow.
H abitet.-Fotmosa.
Holotype,f€maie,Xleizi Hot Springs, foot of HasseNan,altitude 2,500 feet, October 25, 1929 (S. lssikd). Paratopotype,female; paratype, 1 sex?, Hassensan,altitude 8,600 Jeet, Ausust 29, 1929 (S. lssikt).
By Edwards's key to the orientsl spef,jd ol Lipophleps I thepresentspeciesruns to G. (L., robinsoni Edwards, of the Federated Malay Stat€s,diffefing in slight details of colorationandcoNo{r

{LrPoPsr,a?s)

Generalcolorationbrown; basalsegmentsof antenne orange;head yelloq the center of the vertex blackened;a conspicuous
silvery lonlltudinel st pe on pleura; legs brownish yellow, thetsrsi passing into black; rvings brownish yellow, the stigrnal
and axillary regions a little darkened; anterior branch of Rs
sinuouson distal third.
Femo,le. Len'gt}r,about 5 millimeters; wing, 4.5.
Rostrum and palpi blaek. Antenne lfiith scapal segmentsorange, the flagellum black. Head yellow, the center of the vertex blackened.
Pronotumand $nterior lateral pretergiteslight sulphur yellow,
the former dark brown on sides. Mesonotum light brown,slightly darker medially; scutal lobesbrown, the median region
yellow; scutellum brown, narrowly margined caudally withpaler; postnotumpale, sparselypruinose. Pleura Iight brown,with a conspicuoussilvery white longitudinal stripe extending
from behindthe fore coxreto the baseof the abdomen,this passing beneaththe halteres;the stdpe is marginedboth aboveand
belowby a nanower infuscatedline; ventral stemopleuriteanddorsal pleurites, including the pleurotergite,more bufiy yellorv.
Halteresyellow, the knobsa little more obscure. Legs with thecoxarand trochantersyellow; femora obscurebrownish yellow,"Joun.

Fed. Malay St. Mtrs- 14 (1928) 104 106.

528

fke Phililipine Journal of Sci.ence

tle tibi€egradually darkening; tarsi passinginto black. Wings(Plate 1, lig. 18) $'ith a faint brownish yellow tinge; costal
region cleareryellow; stigma appeadngas a longitudinal dusky
streak in cell R,",the remainder of the region yellow; axillary
region a little darkened; outer ends of the radial cells slightly
darkened;veins pale brown, Sc yellower. Venationi Sc short,
Se, ending some distanc€ before the o gin of Rs, the latter
angulatedat origin; anterior bmnch of Rs sinuous,the distai
third arcuateand evidently marking the point of depadure ofa small cephalicbra[ch, R3,normally present in the sulphurella
gToup; m-cu shortly before the fork of M.
Abdominal tergites brown, the segmentsnarroivly malgined
caudallywith pale yellow; sternites more yellowish,the caudal
margins narrowly paler yellow, the lateral margins narrov.ly
infuscated,most distinct on the basal segnents.
Habito,t.-Formosa.
Holotype,female,Meizi Hot Springs,foot of Hassensan,altitude 2,500feet October25, 1929 (S. lsst/ci).
Gmorwia (Lipophleps) sikuosa is well distinguishedfrom
other regiolla.lspeciesby the details of co]orationand venation.
The courseof the anterior branch of Rs is verv Deculiarandsuggestive.
CRYPToI,ADIS (BADoURA) TnIcEoPoDA

r!, .o1.

General coloration blark; head gray; halteres with yellorv
knobs; legs brown, conspicuouslyhairy; wings strongly tinged'lvith blackish,streakedlongitudinallywith pale; cell 2d A rarrow; male hypopygiumwith the dististyle a simplecurvedblade,
the tip acuie.
Male,-Lengtb about 3.8 millimeters; wing, 4.5.
Rostrum and palpi dark brown. Antenn€ black throughout;secondscapal segimentoval; flagellar segmentsgradually decreasingin size outwardly; verticils elongate. Head dull gray,the orbits paler.
Pronotum black. Mesonotumblack, the very restricted anterior and posterior lateral pretergites, together with the humeral region of ihe prEescutumyellow; scutellum orange, ihebase blackenedmedially. Pleura black. Halteres blach, theknobs and extreme base of stem yellow. Legs with the co).:€eblack; trochanters brownish yellow; femora and tibie brown,
the tips darker; tarsi more blackened;segmentsof legs withvery long conspicuouserect sete. Wings (Plate 1, fig. 19)

Aleiand,el': Tipulid,e lrom Eo$tern ABio, VIII

529

strongly tinged with blackish, the elongate stigmal regionslightly darker; pale longitudinal streaks in costal region, andalong veins M, Ms+a,and 1st A; veins dark brorrn. Macrotrichia of veins long and conspicuous. V€nation: Sc, ending
oppositefork of Rs, Sc" some distance from tip of Sc1; cell
2d A narrow.
Abdomen dark brown, the hypopygium blackened,the disiistylesyellow. Male hypopygium (Plete 3, 69. 42) with the dististyle 4 a curved blade that terminates in an acute point, thedistal half $rith Dumerous small set€. What may be a tergal
structure appeaN as two pale tmncated cushions,denselyset
with short setule, eachlobewith a single longer bristle. Thereis a possibility that these structures are really gonapophyses,
but if so they are very dilTerent from those of other allied
species.
-Iilbirat.-Formosa {south).
Holotype, male, Keinensan,near Heito, altitude 5,000 feet,
March 13. 1929 (S, /ssi,ti).
(AlexCrlptol&bis trickopoda is very distinct from tl. aA_???ogenus.
Formosanander), the only otherspeciesof the
In general appearanceit is more like the Indian C. funebris (Alexander), diffe ng conspicllouslyin the structure of the dististylesof the msle hypopygium.
ERIOPTEN^ (MEDA)

Genelal coloration gaay; scapal segmentsof antenn.eblack;legs light brown; halteres with conspicuoussulphur yellow
knobs; wings gray, the Btigmalregion scarcelydarker; Sc long;anal veins strongly divergent; male hypopygiumwith the dististyles urblackened.
Mole.-Length, about 2.8 millimeters; wing, 3.5.
Rostrum gray; palpi black. AntennE with th€ scapal segments black, the flagellumpale brownish yellow, the outer segments darker. Head light gray.
Mesonotumdark browtish gray, the lateml pretergites lightsulphur yellow, the posterior scledtes of notum clearer gTay.
Pleura dark gray, the dorsopleumlregion pale yellowish white.
llalteres pale, the knobs light sulphur yellorir. Legs rvith thefore coxE dark brown, the remaining corneand trochantersbrowrish yellow; remainder ol legs light brorfin, the terminal
tarsal segmentsdarker. Wings (Plate 1, fig.20) broad, iddescent gaay, the presrcular and costal portions more yellowish;

530

The Pkilippine Journ1l of Science

Ihose in the costal
stiqma scarcelydarker: \eins pale brown'
ypllowish. Macrotriehia "l \"ins rPlati\elv short'
;*"
;;;;;
Rs' Sc" av"-"uilo", s" fo"g, Sc' enaling bevond midlength olf""- ita tip; R, of moderatelength' longer than"r't""i-al"ir"""lork of M: \ein 2d A shorr and nearlv"traishl';i" :; ;-;;;,
margin'rhe
inal voinsstronglvdi\ ergent:cell lsi A \ erv $ ideal"--l'bd;-""pale;
hvpopJowlynaincisures
dark browa, theMalp hvpopvgiumrrlate 3 fis 43r with rhe.t;;-;;il"*.
verv longiui"i iot" oi tu"l"tvle. D.terrninatingin two or lhree
Dististyles
sete
erect
denselyset withit.". f"t""tttter'""t*; broa.d,not blackened,
the out€r style bifid
both
H abitat.-Ct,i}.'s"(Szechwan).
14'
Holotype,male, Mount Omei, altitude 4,500leet' August(er Pariskl .
1929"
is rcadilv separated(Erwedd) sutfureocLro@to
nrloir"rif.o- ait tegio"at specieswith long subcostaby the coloration'
yellow knobsof the halteres'and theu" ttt" lo""ii"uooa"olphurand courseof vein R3 and the strong,renation,as the lengthveins.
ilivergenceof ihe anal
DNIO"IERA(E}IPEDA)

generalcoloraSizevery small (1ving,male, 2.6 millimeters) ;yellow;
obscureknobs
ihetion graylsh black; halteres dusky,
cell
long;
Sc
blackish;with
i"g" lf""t; $'ings stronglv suiluseal
1st M, closed.
nIaie.-Leng\h, about 2 millimetels; wing, 2 6'
no"t"o* utl'A palpi bro'n'nish black AntennEeblack thrcughout. Head dark brownish gray.
Pronotum brownish black Anterior lateral pretergites very
restricteallyobscureyellow. Mesonotumdull grayish black' the
fovee anal tuberculatepits more intense black;p"".a"-.t.i*fscutellum somewhat more pruinose Pleura grayish black'
Halteres alusky,the knobs obscureyellow Legs with the co*e
blackish; trocharters blowDishyellorfi;remainderof legsbrownish black. Wings (Plate 1, fig. 2t) with a strong blackish suffusion, the veins alarker. Venation: Sc long, Sc' extendingto
beyoni one-thirdthe len$h of Rs, Sc. a short distancefrom its
tip; Rs relatively long ar'}alstraight; vein R, straight, the cell
justthus pointed at base; cell 1st M" closed;m-cu erect, placed
beyond the fork of M.
AMomen black, including the hypopvgium Male hypopvqium with the outer dististyle blackened,the dististyle bifid'
Ihe outer arm denselycoveredwith microscopicsetuls lnner

43,{

Aler:a der: Tipund,e lron, Ed"stern Asi&, VIII

691

dististyle s straight slenderrod, the apex obtuse. Gonapophysesappearingas pale flaftenedblades.
Eabitat.-Fotmosa.
Holotype, male, Hassensan,altitude 8,600 feet, August 29,
1929 (S. l,ssiri).
Erioptera (Empeda\ liiputiwt is readily told from all de_scribedregional speciesby the small size and venation,notably
the closedcell 1st M?.
oaxoara aNTERAcoPoDt 4. nor,

Generalcoloration gray; halteres yellow; legs black; wings
cream-colored,with a sparsebrown pattern; rDalehypopygiumwith the apex of the basistyleproducedinfo a small spine,
Male.-Length, about 6 miUimeters;wing, ?.
Rost rm and palpi black. Antenn€ black throughout, of moderate length; velticils of the basal flagellar segments long, becomitg shorter on the outer segmenLs. Head gaay,
Mesonotalpr-ascutumlight g:ray with four alarker brownish
gray stripes that are inconspicuousagainst the ground color;pseudosuturalfoves and tuberculate pits black; scutal lobes
brownish gray; scutellumand postnotumclearer g?ay. pleura
clear gray. Halteres yellow. Legs with the coxe anal trochanters brownish yellow; remainder of legs black. Wings (plate
1, 69. 22) cream-colored,*ith e slight brownish tinge; stigma
brown; restricted darker bro$rn clouds at origin of Rs, Sc,,
along cord, m, and tips of most longitualinalveias; retatively
incompicuouswhitish areasbeforeand beyondthe stigma; veinsbrown, darker in the cloudedareas. Venation: Sq enalingopposit R.; Sc, about optr)ositeone-flfth the lengdr of Es; ii,
just beyondrhe lork o[ Rr.r_r: tips of vpins R and R,
de_nectedcephalad:point ot unionot disralse(tionot ll. *iitr rn
angulaledand weakly spurred: m-cu shortly before fo;k oI M;
vein 2d A sinuouson distal half.
Abdomen bromish gray, the hypopygium dark. Male hypo_pygium (Plate 3, fig. 44) with the tip of the basistyte,b, pro_alucedinto a small acut€ spine. Two dististyles, one a strongly
curved sickle-shaped
spine, ertendedinto a long straight blackenedpoint, the margin with very long erect sete; seconcldisti_styl€ nearly straight, the basal half more dilateal. conapophyses
small, in generaloutline nearly like the inner dististyle,the apical spine more acute.
Ed,bita;t.-Formosa.

532

Tke PlLiwpine Journal of Scie ce

Holotype,male, Hassensan,altitude 6,500to ?,800feet, October 26, 1929 (S. Issi,!t?).
Ormosiaanthracopoddis very distinct from the other speciesof the gerus describedfrom Formosa. It belongsto the ,rcrle{tta gro].Jp,
havinr: the apex of the basistyleof the male hypopygium producedinto an acute spine.
Generalcolorationpale yellow; legs uniformly yellow or withscarcelyindicated pale brown bands on femora; wings yello\ry,
the veinsalongcord and outer endsof cell 1st M" a little darker;
vein 2d A gently cuNed to the margin; male hypopygium{iththree apical spineson basistyle,one more isolatedand terminating in an obtuse kllob that is fudher abruptly narrowed intoa hairlike spine.
M&Ie.-Length, about 7 millimeters; wing, 5 to 5.5.
Femal,e.-Lelj,gtrh,
about 5 to 5.5 millimeters; rting, 5 to 5.4,
palpiyellow;
Rostrum
alternately yellow and dark brown.
paleyellow,
the scapal seSments scarcely
Antennal flagellum
darker. Ilead light sulphur yellow.
Pronotum sulphur yellow. M$onotum more testac€ousyellow, without dark markings. Pleura testaceousyello,F. HaIteres pale, the knobs light yellow. Legs uniformly yellow, illcases the femora with scarcely indicated pa.le brown bands;
terminal tarsal seglnentsblack. Wings (Plate 1, fig. 23) yellow, with a sparse brown clouding along the cord and outer mdof cell 1st M,, most evident on r-m; veins yellow, darker in theinfuscated areas. Venation: Vein 2d A curved gently to themargrn,Abdomenyellow, the caudalmargns of the intermediatetergites narrowly dark brolr1r. Male hypopyg:ium(Plate 3, lig.
45) with the niDth sternite, 9s, broad, the apical spineswidely
separated.Basistylctprminatingin three conspicuous
spines.
two being long and extendedinto acutepoints, the third arisingfrom a separate apical lobe of the basistyle, knobbed at apc\,
thence further producedinto a hairlike point. Dististyle, d,complicatedin structure, the long slender outer arm bearing
a very elongate subterminal seta; margin of arm near base witha series of peglike spines. Spines of intermediate arms of
dististyle unusually long and slender.
Aabit&t.-Formosa.
Holotype, male, Hassensan,altitude 3,500 feet, October 21,
1929 (S./sst,&t). Allotopotype,female,in copulawith the type.

Aleraniler: Tipulii,E from E6te1"n ABitL,VIII

533

Paratopotypes,6 malesand females,altitude 2,500to 5,500feet,
October22 to 25, 1929 (,s.Issdtrd); paratype,1 female,Funkiko,April 21, 1917 (7. Slriraki\.
St'ltrinftor Uia tailoanensis has been confused with ld?., Brunetti and the paraiype wa"5 earlier recorded as being thatspecies.r The prssentform differs in the colorationof the wingsand de[ai]sol slruc{ureol Lhemaleh}?opygium.
Generalcolorationyellow, the mesonotumlargely black, moreor less vadegated with pale at the suture; halteres yellow;
femora and fibiar with narrow brownish black rings; ivirlgs
yellow, with a sparse dark pattern, including the vicinity of
t-m, outer end of cell 1st M" and end of vein 2d A; small dark
marginal spots at ends of medial and cubital veins; veir 2d A
strongly angulatedand weakly spurred at margin; male hypopygium lvith the dististyle very Iarge and complex,fourlobed,
the two intermediatelobeswith combsof sfines.
Me,tre.-Length,about ? to 8 millimeters; wing, 5.2 to 6.
Fam,ala.-Ler].gt}i,
about 6 to 6.5 millimeters; wing, 4.6 to 5.
Antenn€ewith the scapeblack; flagellumyellow. Headchiefly
ocherous;a dark spot touchiEginner margin of eye at narowest point of vertex.
Mesonotumextensivelyto almost entirely blackened,more orless pruinose, in certain of the specimensbrightened at theauture; the amountof pale coloring variable,in somespecimens
lnvolving the posterior third of the prescutum and the scutal
lobes, the anterior portion of prpscutum always blackened.
Pleura abruptly aDduniformly yellow,. Halterespale,the knobs
bright yellow. Legs with the cox€ and trochanten yellow;
femora yellorr, with two narrow dark rings that are interrupted
beneath;tibi,D yellow, the tips narrowly bla4ken€d,with a narrower ring at just before midlength, this obsoleteon lorrer
surface; tarsi yellow, the tips of tarsal segaents one and two
darkened; terminal segment dark broivll. Wings (plat€ 1,
fig. 24) yellow,the veins a little darker yellow; a restrieteddark
pattern, arranged as follows: At r-m, involving the endsof all
surrounding veins; outer €nd of cell 1st M"; on m-cu and its
junction with Cu,; tip of 2d A; small darkenedmarginal a}6as
at endsof veins Ri to Cu,, inclusive. Venation: 2d A angularly' -{ n n . E x t. So c .Am€ ri c a 1 3 (1t20) 253.

554

The Phdwpine Jownal of Science

bent into the margin, this curvature rectangular or acute, usuallywith a sholt spur at the bend.
Abdomen yellow, the tergites with geminate brown spota at
caudalmargin, with vagueindicatiols of paler brown markingBon the bgsal ring ; on segmentsevenwith a median browlt stripe ;hypopygium pale. Male hypopygium (Plate 3, fig. 46) $rith
the distiBtyle, d, fourlobed, the outer lobe long and slender,
beadng the usual very long apical seta; eecondarm bilob€dat
apex,the margin with groups of long black spinesto producea
cornblike appearauce; third lobe flattened, with a ^ -shaped
seriesof short€rpeglikespine,s;innermostlobebearingan apicalsedes of very long setoid Bpines. Ninth sternite, 9s, slender,
with two apical se@ that s,re placed close together. Ninth tergite without lateral shoulders, as fol.ltld.in makensia.
Eabitat.-W e9letrr Chi[a and Formosa.
China, altitude 4,500
Holotype,male, Mount Omei, Szech$ran,
feet, Auglst 4, 1929 (et Pd,rbh\. Allotopotype, female. Paratopotlaes, 15 males and females,August 2 to 19, 1929;paratyp€s, 8 malesand Jemales,Hassensan,Formosa,altitude 2,500
to 3,600fe€t, October21 to 26, 1929 (S. lssiftd).
By Edwards'skey to the speciesof Stltringomaia" the presEdwafds, s.n otherwise very difierentent fly runs to S.
'l?dr€?xsis
fly. The structureof the male hypopygium separates S. sizet sis from all regionalforms.
"Tranr. Ent. Sd, tondon (191a1)210-212.

ILLUSTRATIONS
Il,aerdr o cdau

j r, bsLirj.j

4 dt turyle; r, son.lolLstu; at jn.€r disti.t le: od,

pr,^rD I
Frc. .1. Litwnia lDiscobold, taitdna\d sp. nov., venauor.
2, Iritunia lLiwnia,/rortngo sp. Dov., venauon.
3. Liwtuio (Libtuor.sl i'@seeM sp. nov., ven.rioa.
1. Irinoni@ (Rhtpid:ia) tridnhnta sp, nov., venation.
6. Linontd (Diqenom!)it, subpan t tab sp. nov., venauo!.
6. Irinonia (e.r@M,ryia) epi.it'Bci4ta sp. nov., venation.
'1, AnNo.h4 (Antocha, rrrr sp. nov., venarion.
A. E.l'iua (Ethdanphid,id\ p.rebsaha sp, nov., venation.
9. DiiLtuptU.hs i:rrd<irasp, ,w., veDarioD,\0. Difianota (AmabpiMt i|Iectdta .p. nov., venetion,
11. Adelphohyia issihihd sp. nov., venation.
72. Untuphi.La (Pfionolabis) s"ni.lentdta sp. !ov., v€Mtio!.
r3, Trent paw,it, \Monsorna, m@tika sp. t
14. Gonmuia \Prosnorngi4,
cozJ!?@ta Ateiandd, venation.
16. Goaohvia (Gnawia,) nansai sp. aov., aenanon,
16. Gan rnAia (Upophleps) sduiat sp. iFv., veBuon.
17. Gonornt t (Lipopklepa, netuehtlosd Bp. trov., vestion,
18. ctuorwta (Lipopkleps) sirwosa sp. nov., venarioh.
79. Cryptolan. lBa.ourdl tttchapodd s!. noy., veDaEon.
20. Eriot'tatu \Emt eda)
sp. Doy,, ven&uon.
21. Erioptetu (Emp.d^, 'ullureodaaata
Iitipttiaa sp. nov., vemrion.
22, Ornbei^ antkracopod.a sp. nov., vensriol
24. Sfuringomltio taiwo,netub rp, nov., vonafion.
24, Slsrinsonuiq si@Mis sp. nov., venrtioD.
PI,Am 2
Trc. 25. DoUchopedo (N.sope2al rdrcalba s!. nov,, msle hwolygium,

late"

25, Dolithops.a lNesope|e, tarsalbd .p. nov., male hr?opysium, ninth
27. DohckopezalN@op..d, ta,satba sp. nov.,mal€ hylopygilm, g@._popnyrB,
28. Dolichopetd (Nesopa.a, tatsalbe sp. ttoa., m.le hylopygiuh, oishth
25. Doli.hopeza loftpetal

saitam^ntis

dp. nor.,30, Dolbhope.a loropa.a)i[ne! disttutyle.

saitane8is

sp. nor.,
585

636

TlLe Phitrinpine Joururl' of Science

Frc, AL Lirmia (Ditcobola) tai"tuelld 3p. nov., oale hr?opvgium
32. LiMtuia \LituoLia) ktuinsa sp. [oi, Eale hvlopvgiun
38. Litunta (Rlupid.ie) triumata sp. n@a role hlTopyg,urrl
Ea. Limonia (Diodnamsi,J subltMtulata 3p. nov, Dale hvpopvgium
36. LiMnia (Gerdnona t) aptcildsDidtd 3p nov., hale hvpopvgium
36. Atut cha \Antocha) st?, sp. novl nale hypopygiuh.
37. D;mnopt!.hu t53i/.i/ sp. nov.. mak n!?opvsiun
Pr.ArD 3
Frc. 3a. Limhophila (Priowlab;s, se1t l,eht4lL sp. tuv, msle hJaopygium
79. G@muia lPtostuotusia) confluentd <l\lexar'detr, dale hvpolv_
a0. Go@n'rio (Gotutnsia) Mwei sp. nov., male h$opvgim
47. Gonotuu;e (Lipophlatu) stuteri sp. nov., ma.le hylopysium
42. CtsptaLdbis (Baeoura, trichopodd 6p. nov., m4le hypolvglum.
43. Enoptetu \Empeda, sulfweo.I@eta sp. !oL, rule hrTopvsium.
11. Ormositu dnthrdcopodd sp. nov, Mle hypopE:m.
15. Stad,ngoevtt tditudnends Ep. rov" mle hypopygiu.
46, Stuinsohrrt sineBis sp, nov. Mle bypoptsiut

STERN

!1!']
^s:A'

Ir\.uP. JorR), S[, rr. No.1.

9TnN As',, v]rl ]LI!'LD. ro*N.

gr-N/i!'.\a26

Ir:--t-ffi

scr.,43, Nn.1,

FRoU xiSTERl

As|r,

vIIl

l

IPsLf.

JouN. S.r., !r, No. 1,fi