617
Arnr6 19471

ALEXANDEN-TIPULIDAE

91

UNDESCRIBEDSPECIESOF TIPULIDAE FROM THE
V/ESTERNUNITED STATES(Diptera)PART IIIBY CHARLES P. ALEXANDER

Maasanhusette Statp College, Amherst

The preceding part under this title was published in the Pan.
Pacific Entomologist, 2I z9L-97;1945. At this time I am describing three new species of the genus Tipula from the San Bern.
ardino Mountains, California. In July, 1945, Mrs. Alexander
and I spent three days eamping at East Barton Flats, on the
East Fork of the Santa Ana River, at a general altitude of about
6250-6300 feet. This was close to the mountain cabins of the
Melander and Sperry families, of Riverside, California, and we
enjoyed and profited greatly by this visit with our good friends.
On one occasion, Nodl Crickmer, Melandero Sperry and I col.
lected up the slopes of Sugarloaf Mountain, to the Little Cienaga,
altitude about 72100feet. During the remainder of the summer
and fall, to the first snows about mid-October, the Sperrys continued to collect Tipulidae, chiefly at the lanterns operated at the
Sperry Cabin. I am vastly indebted to all my friends above indicated for their many kindnesses,including the large series ofTipulidae that has been presentedto me.

Tipula (Trichotipula) subapacheAlexander,new species
Allied tn a,pqnlrc,' general coloration of thorax EnLV, the praescutum with a central paler gray stripe, the posterior sclerites of
the notum with a yellow central line; antennae black, the scapechiefly yellow; femora brownish yellow, the tips narrowly blackened; wings with a strong blackish tinge, cell Sc and the stigma
darker brown; obliterative areas eonspicuous; macrotrichia inouter ends of cells .Es and M'; abdomen brownish black, the basal
tergal rings narrowly yellow; male hypopygrum with the notch of
the tergite V-shaped, provided with long cylindrical blackened
spines; outer dististyle nearly twice as long as broad, with long
black setae; inner dististyle with its main body nearly parallelsided, the beak short and stout, slightly upturned; outer basal lobe

92

TrrE PAN-PACTFICENToMOLoGIST [VOr,. XXIII, NO.2

large and conspicuous, flattened, exceeding one-half the length ofmain body of style.
Male. Length about 12.5-13 mm.; wing 11-11.5 mm.; antenna
about 4 mm.
Femnle. Length about t4-15 mm.; wing 14-15 mm.
Frontal prolongation of head blaek, more ehestnut brown basally
above; nasus elongate, black; palpi black. Antenna with scapeobscure yellow, narrowly darkened at apex; pedicel brownish
black, flagellum black; flagellar segments very gently incised.
Head alrove yellow, the sides of the posterior vertex and the genae
infuscated, pruinose.
Pronotum yellow, with a dark spot on either side of the midline.
Mesonotal praescutum with a broad light gray central stripe thatis more or less divided by a capillary darkened median vitta;
iateral stripes narrow, more brownish grZY, remainder of praescutum brownish black; posterior sclerites of notum blaek, more orless pruinose, with a continuous yellow stripe, broader and moretestaceous yellow on scutellum, very narrow on the mediotergite;
scutal lobes with the gray areas more or less distinctly ringed
with black. Pleura \Mith the propleura and mesepisternum more
blackened, pruinose, the mesepimeron, meron and pleurotergite
paler. Halteres with the stem blackened, its base and the apexof knob yellow. I-,egs with the coxae blackened, pruinose; trochanters brownish yellow; femora obscure brownish yellow, thebrownish yellow,
tips namowly blackened; tibiae arrd tarsi obscure'Wingswith a strong
the outer tarsal segments passing into black.
blackish tinge, cell Sc and the stigma darker brown; pale areas
before and beyond stigrna and as a disconnected band at cord, thelatter crossing the base of cell lst M,; further pale long:itudinal
streaks in several of the cells, especially on either side of vein M
and as a V-shaped area in cell lst A; less evident pale streaks insome of the outer cells; veins brown. Macrotrichia in outer ends
of cells Ro and,Mr. Venation: Sc' persistent; Rs arcuated, shorter
than nL-cui rzr longer than petiole of cell M,.
Abdominal tergites brcwnish black, at least on sirJes,in cases
paler brown medially, the basal rings narrowly but conspicuously
light yellow; basal sternites more uniformly brownish yellow, theouter segments, with the hypopygium, more infuscated. Ovipositor with both cerci and hypovalvae compressed-flattened,their tips
obtusely rounded. Male hypopygium having the posterior borderof the ninth tergite with a deep V-shaped notch that is provided
with several long eylindrieal biackened spines with blunt tips; atends of the obtuse lobes vrith a small group of more conical spines.
Outer dististyle moderately broad, the leneth nearly twice thebreadth, provided with long black setae, the longest on outermargin fully as long as the width of the style. Inner dististylewith the main body nearly parallel-sided, the beak short and stout,
slightly upturned; outer basal lobe large and conspicuous, flat-

Aenn, 19471

ALEXANDER_TIPI'LIDAE

93

tened, exceeding one-half the length of main body of style. Phal'
Iosome more or less rolled into an open tube.
Habita. C,u,lronnrl (S.rw BBnnmorNo CouNTy).
Holotype, 6, Elsr BenroN Fr..nrs, San Bernardino Mountains, altitude 6300 feet, September 22, 194,6 (John and Grace
Sperry) . Allotopotype, 9, with the type. Paratopotypes, LL 6 9,
August 19, September 27, 1946.
This fly is closest to TipuJa (Triehotipula) apach,e Alexander,of northern New Mexico, differing in details of coloration andin the structure of the male hypopygium, especially the tergite
and both dististyles, particularly the large outer basal lobe of
the inner style. The rather numerous California species oL TrL
chotipula have been revised recently by the writer (Bull. So.
California Acad. Sci., 45:l-16; L945).
Tipula(Oreomyza)

gracia Alexander, new species

Belongs to the mnrmorata (f ragi,li,s) group; general coloration{ray, the praescutum with four brownish gray sfuripes; femora
brownish yellow, the tips broadly and conspicuously blackened;wings whitish subhyaline, with a pale grayish brown marbled pattern, especially pale in the costal and apical portions heavier inthe medial and anal cells; abdomen brown the lateral and caudal
borders of segments yellow; male hypopygrum wtth the tergite
having a low V-shaped notch, the outer angles only sliightly produced; outer dististyle with the upper margin at base blackened
but very obtuse; inner dististyle with the apex blackened, relatively stout, simple; eighth sternite with the caudal border produced into a low median lobe that bears about a score of long
black setae.
MaIe. Length about L2-13 mm.; wing 13-15.5 mrn.; antenna
about 2.5-2.7 mm.
Frontal prolongation of head dark brown, heavily pruinose, especially above; nasus distinct; palpi black. Antennae short, scapeand pedicel yellow, flagellum black; flagellar segments simple, alittle longer than the verticils. Head gra;y, with indications of adelicate darker line on the low vertical tubercle.
Pronotum gr&V, the scutellum light yellow. Mesonotum gray,the praescutum with four brownish gray stripes, the intermediatepair more pruinose on anterior porbions; lateral margin below thehumeri blackened; posterior selerites of notum light gray, eachscutal lobe with two separate brownish gray areas; a delicate
broken central dark vitta on scutellum and mediotergite. Pleura
g:ra.y, vrith darker gray areas, most evident on the ventral sterno.
pleurite and meron; dorsopleural region yellow. Halteres yellow,

TrrE pAN-pACrF.rC ENT0MOLOGIST[VOl

XXIII, NO.2

knobs dark brown. Legs with the coxae gray pruinose; trochanters brownish yellowl femora obscure yellow basally, more darkened outwardly, the tips broadly and conspicuously blackened;
tarsi
tibiae brownish yellow, the tips more narrowly infuscated;'Wings
brownish yellow, passing into black; claws (male) toothed.
whitish subhyaline, with a pale grayish brom marbled pattern,
distributed about as in other members of the group, especially palein the costal and apical portions of wing, heavier in the medial
and anal cells, especially lst A; stigma paler brown; veins brown.
Venation: .Es about one-half to three-fourths longer than nx-cu;
basal section of Br*o short to very short; Mz+tYet;i' short to punctiform.
Abdomen brown, the lateral and caudal borders of the tergites
yellow,. the latter more distinct and becoming more extensive on
the outer segments; sternites generally similar, the bases of theeighth and ninth sternites extensively blackened. Male hypopygrum having the ninth tergite with a low V-shaped notch on
caudal margin, the outer angles only slightly produced, not spinous; margin of notch with a few small spinous points and setae.
Outer dististyle flattened, the upper margin at base blackened butvery obtuse, not at all toothed; surface of style with coarse black
setae. Inner dististyle heavily blackened at apex and along margins; apex relatively stout, simple; outer basal lobe a low cushion,
set with several erect stout black setae, the basal portion with asmall blaekenedlobule that is provided vrith microscopic blackened
setulae. Eighth sternite moderately sheathing, the caudal borderproduced into a low inconspicuous median lobe that bears abouta score of long black setae, the marginal setae of the remainderof sternite similarly long but yellow; center of disk glabrous.
Habitat.

C.lr-rroRut.q. (SaN BrnwanuNo CouNrY).

Holotype, 8, 1000 SpruNcs Mnloows, San Gorgonio Trail,
San Bernardino Mountains, altitude 7700 feet, September 2L,
1946 (John L. Sperry). Paratopotypes,l0 6 6, altitude 75007700 feet, September 20-2L, L946 (John and Grace Sperry and
NoEl Crickmer).
This very distinct fly is named for Mrs. John L. Sperry (Grace
Herreshoff Sperry), of Riverside, California, who specializeson
a study of the Lepidoptera. The Sperrys have camped and col'lected in hundredsof placesin the West and many entomologists
are greatly indebted to them for invaluable insect materials inmany orders. The discoveryof a member of this group oL Tipula
so far to the south was very surprising. The only other species
at present known from Western North America are Tipula(Oreomyza) lragiliru Alexander, Alaska to Colorado, along theAenlq 19471

ALEXANDER-TTPULTDAE

95

Rocky Mountains, and T. (O.) phoroctenia Alexander, from

British Columbia,eastwardto Maine, in the CanadianZone. All
three speciesare on the wing in the autumn. The speciesare
well distinguishedamong themselvesby hypopygial characters,
particularly the dististyles and eighth sternite.
Tipula(Lunatipula)fido Alexander, new species

Mesonotum chiefly gr&V, the praescutum with four entire reddish brown stripes, the intermediate pair narrow; antennae relatively short, basal three segments yellow, remainder of flagellum
black; flagellar segments only moderately incised; femora yellow,
weakly darkened at tips; wings with a weak brownish tinge, theobliterative area before cord conspicuous; abdomen, including hypopygium, dull yellow, scarcely patterned; ninth tergite almost
divided medially by pale membrane, the lower outer angle of eachlobe produced into a stout blackened spine; inner dististyle withbeak long and slender, widely separated from the small lower
beak; posterior crest of style produced into a triangular point;
phallosome consisting of four rods in pairs of two each; eighthsternite sheathing, each outer angle with a strong fasciculate
bristle; a broad median plate that suddenly narrows into a stout
fingerlike lobe, the whole outer porbion with a fringe of unusuallylong setae.
Male. Length about t6-L7 mm.; wing 15-15.5 mm.; antenna
about 3.5-4 mm.
Fem.ale. Length about 18 mm.; wing 17 mm.
Frontal prolongation of head yellow; nasus stout; palpi obscure brownish yellow, the outer half of terminal segment blackened. Antennae relatively short; basal three segments yellow, remainder of flagellum black; flagellar segments only moderately
incised, a trifle shorter than the longest verticils. Head yellow,
sparsely pruinose, especially on the scarcely developed vertical
tubercle and the posterior orbits.
Pronotum brownish yellow. Mesonotum chiefly gray, the praescutum with four entire reddish brown stripes, the intermediatepair narrow; posterior sclerites of notum more heavily pruinose.
Pleura yellow, sparsely pruinose; dorsopleural membrane clear
yellow. Halteres with stem dusky, pale at base, knob more darkened. Legs wth the coxae and trochanters yellow; femora brownish yellow, narrowly and weakly darkened at tips; tibiae and basitarsi yellowish brown, the remainder of tarsi passing into black;'Wing:sclaws (male) toothed.
with a weak brownish tinge, heavier
prearcularin theand costal fields; stigma small, darker brown;a diffuse brown cloud over rn4u,' obliterative area before cord conspicuous, extending to the basal third of cell M"i no post-stigmal
brightening; veins brown. Venation: Es arcuated to weakly angulated at origin, nearly twice mfrui Rr+zpr€s€rved; m and petiole

96

rHE pAN-pAcrFrc ENTouoLocrsT [VOI..XXIII, NO.2

of cell M, subequal in length; Mr- subequal to or a trifle longer
than the basal section of M,*,.
Abdomen dull yellow, scarcely patterned. Ovipositor with thecerci long and relatively slender, only gently upcurved to the subacute tips. Male hypopygium with the ninth tergite relatively
small, divided medially by pale membrane; lower outer angles
produced caudad and ventrad into stout blackened spines; posterior border mesad of these spines less heavily blackened. Ninthsternite with its appendage long and curved, more or less flab
tened. Basistyle entire, unarmed. Outer dististyle with outer half
dilated into a weak spatula- Inner dististyle with the beak longand slender, widely separated from the small more blackened
lower beak by a large oval notch; dorsal crest low, slightly corrugated, setae small; posterior crest produced into a triangularpoint, the entire region with numerous coarse reddish setae; outerbasal lobe elongate-triangular in outline, the posterior portion
fringed with long coarse reddish setae. Phallosome conspicuous,consisting of four sclerotized toothed rods, arranged in pairs of
two each. Eighth sternite distinctive, sheathing, with a semidetached lobe at each outer angle, this tipped with a single strong
faseiculate bristle; a broad median plate that suddenly narrows
at apex into a stout fingerlike lobe, the whole outer portion, including the lobe, with a fringe of unusually long setae that appear
as a flattened brush directed caudad.

Habitat. CnIronxr.e, (Sax BrnnenorNo CouNry).
Holotype, I , E.lsr B,mron Futs, San Bernardino Mountains, altitude 6300 feet, August 17, 1946 (John and Graco
Sperry) . Allotopotype, 9, at light, July L6, L946. (Sperry andAlexander). Paratopotypes,2 6 6, with the allotype.
This fly appears to belong to the unicircta group but is not
typical and its exact relationshipsare in doubt. Somepoints ofsimilarity with species such as Tipula (Lurwtipula) rnorrnon
Alexander seemto be indicated.
Ilvro rrrE SaN
A EunopEANWEEvTLNEwLy INTRoDUcEDFnaxcrsco Blv RBcroN
While collectingin the outskirtsof Mill Valley, Marin County,
on April 4, 1947,D. Giuliani, one of our youngerentomologists,
noticed numerous specimensof Barypi,thes pallucid,a Boh., crawling up the curbing along the roadside.
This European weevil has long been esetablished in t}e East-ern part of our country but this is, I believe,the firet time that it
has been noticed on the Pacific Coast.-EDwIN C. Vmv Dvrr.