44?Soclutv,
Reprintedfrom Bur,r,nrrxoF TrrEBnoorr,yN ENtorroLoGICAt,
Vol. XXXIII, No. 2, pp. 71-78. April, 1938.

CRANE-FLIESNEW OR INSUFFICIENTLY-KNOWNFROM THE NEARCTIC REGION (TIPULIDAE,PART IV.
DIPTERA).
Bv Cnenrrs P. At BxaNDER,Amherst, Mass.*
The precedinginstalmentunder this generaltitle was publishedin r93r (Bull. Brooklyn Ent. Soc.,z6: ry7-r8+). Except wherestatedto the contrary, the types of the noveltiesherewith described
are preservedin my collection,through the friendly interest of thevarious collectors.
Tipula (Lunatipula) leechi n. sp.
Allied to rnacrolabis;ground-colorof mesonotumand pleura
light yellow, the praescutumwith four narrow bright brown
stripes; ground-colorof wings light yellow, with a restrictedbrown pattern; male hypopygium large, the lateral arms ofbasistylevery stout, with more than the outer half slightly
expanded,the dorsal face with a weak carina that extendsto
apex of arm, making it appearmore or lesspointed.
Male.-Length about20 mm.; wing 2r rnm.
Frontal prolongationof headbrownishochreous,nasusdistinct ; basal segmentof palpus obscureyellow, the remainderpassingto brownish black. Antennaewith the basalfour segsegmentsbicolorous,yellow, thementsyellow, the succeeding
basalenlargementbrown; outer segmentssomewhatmore uniformly darkened. Head light yellow in front, light brown on
posteriorvertex.
Mesonotalpraescutumwith the ground-colorlight yellow,
with four narrow, bright brown stripes,the intermediatepair
by a groundline nearlyas wide as either stripe; eachseparatedscutallobe with two small pale brown areas; posteriorsclerites
of notum and the pleurachieflyyellow pollinose,the scutellum
and mediotergite with a vague capillary median dark line.
Halteresyellow, the knobs dark brown. Legs with the coxae
and trochantersyellow; femora reddish yellow, the tips narrowly darkened; tibiae obscureyellow; tarsi passinginto darkbrown; tibial spur formula r-2-2; clawssimple. Wings withthe ground-colorlight yellow,the prearcularand costalregions
deeperyellow; a restrictedbrown and whitish pattern; thepale brown areasare evidentchiefly as cloudsin the apicalandcaudal portions of wing ; a seriesof four somewhatdarker* Contribution from the Department of Entomology, MassachusettsState College.

72 Btclletinof the BrooklgnEntomologicalSocietg Tot.xxxUlbrown areasin anterior field of wing, thesemuch smallerandless evident than in ncacrolabi,s;the whitish areas lie chiefly
alongboth sidesof cord and beyondthe stigma. Squamawith
a few setae.
Abdomenyellow, the tergitesvery narrowly trivittate withdark brown, these delicatelines more or less interrupted at
the caudalmarginsof the segments;extremecaudaland lateral
margins of tergites pale; hypopygiumlarge, chestnut-brown.
Male hypopygium generally as in rnacrolabrsbut the lateralarms of basistylevery stout,with slightly more than the distal
half a little dilatedinto a head,the dorsalfaceof the expanded
portion with a gently curved carina that extendsto the apexof the head,making it appearacuteor subacute.
Habitat; British Columbia.
Holotype: 3, Vancouver,July r r, r93o (Hugh B. Leech).
Tipula (Lunatipula) leechiis namedin honor of Dr. Hugh B.
Leech,of the University of California,to whom I am indebtedfor
numerousinterestingspeciesof Tipulidae. The fly is quite distinct
from T. (L.) macrolabisLoew and T. (L.) youngi,Alexander inthe unusuallystout lateralarm of the basistyle,with only the distal
half weakly expanded. In macrolabi,s,the outer third of the arm
is expandedinto a flattenedsmoothpaddle-likeblade; in youngi,,
this blade is produced into two acute spinous points. T. (L.)
spectabilrsDoane has long been regardedas being identical withmacrolabis, the structure of the male hypopygium being very
similar in both flies; however,the mesonotumof spectabilas
is moreyellowishthan in rnacrolabisand may be found to representstill
anothercloselyallied speciesin this group.
Pedicia (Pedicia) parvicellula n. sp.
Female.-Length about 29 mm.; wing 2r.S n'rm.
Size and generalappearance
about as in P. (P.) margari,taAlexander (EasternUnited States). Antennaedarker brown
throughout. Mesonotal praescutum with the ground-color
buffy, with three slightly darker stripes,the medianonevaguely
dividedby a capillaryvitta. Postnotalmediotergiteand pleurotergite dark brown. Femora obscureyellow, the tips narrowly and gradually infuscated,not abruptly blackened,as inrnargarita. Wings with the dark pattern as in margarita, thedark seamalong vein Cu continuedto margin; basalthird of
costalcell whitened. Venation: Cell M, very small, approximately one-thirdthe length of its petiole. Abdominaltergites
reddish brown, with a darker brown median stripe that is

7s38 Buttetin of the Brooklgn EntomologicalSocietg 73
Aptri,t,narrowly interrupted at the caudal margin of the segments;
sternitei uniformly reddish brown. Ovipositor with the cerci
much longer than in rnargarita, very gently upcurved. The
abdominal tergites entirely lack the contrasting pattern ofreddish brown and whitish gray characteristicof the three
speciesof Eastern North America, albi,ai,ttaWalker, conterrni,naWalker and rnargarita Alexander.
Habitat; British Columbia.
Holotype: CypressCreek, Vancouver,August 16, r93r (HughB. Leech).
I have little doubt but that this is the speciesthat has been determinedby Aldrich (Psyche,7| 2o2; 1895) "s obtusaOsten Sacken,
from material taken at Lake lJnion, Seattle,Washington, in lateAugust. His specimen,d male, agreesvery closely with the typefemale describedabove (Male, length about z8 mm.; wing aboutzz mm.). The specimenhas the pattern of the abdominaltergites
and the wing-markingsquite as describedabove. Aldrich describes
the male as having the styli of unusual length, being approximately
twice that of.albiztitta. The unusually long cerci of the female ofthe present fly would seemto be correlatedwith this elongationot
the male styli. Pedicia obtusa Osten Sacken (Bull. U. S. Geol.
Survey,3t 2c,5-zo6;ISTD was provisionallydefinedfrom a specimen taken near Saucelito,Marin County, California, in the Spring.
Later, Osten Sacken(Psyche,7: 22g, 23o; 1895) gave fuller noteson what would seemto representthis samespecies,taken in Siskiyou
County, California, Septemberz7 and October6; all of thesespecimens show the dark seamalong vein Cu endingat the level of thecord, quite as in the Eastern conterrni,na. The specieshere characterized as new has this seamcontinuedto the wing-margin. Although abnormal specimensof.Pedicia albiztitta and P. ri,ztosaLinnaeus are occasionallyfound where there is a difference in the
degreeof extensionof this seam,in large seriesof specimensit has
proved to be a very constantcharacter.
LruNopHrLA Macquart.
Trichephelia n. subgenus.
Charactersas in Elaeophila Rondani, differing especiallyin
the possessionof numerouscoarsemacrotrichiain apical ce]!
of wing, f rom R, to M * inclusive. Male hypopygium withthe apex of outer dististyleterminatingin a long slenderblackspine.
Typ. of subgenus.-Lirnnophila (Trichephelia) seti'cellulon. sp.
(Nearctic Region: AlleghenianSubregion).

74 Bulletin of the Brooklyn Entomological Society Tot.XXXIII
Limnophila (Trichephelia) seticellula n. sp.
Generalcolorationblackish,pruinose,the praescutalstripes
Tore yellowish brown ; antennal scapeand pedicel brownish
black, the. flagellum paler; pleura with a biack longitudinal
stripe; wings yellowish, with a heavy dark brown pattern;
apical cells of wing with conspicuous
macrotrichia; male hypopygiumwith the outer dististyleterrninatingin a long ".,t1.
spine, with a smaller lateral spine on outer margin "t nearfour-fifths the length.
Male.-Length about g mm.; wing g.g mm.
Rostrum and palpi black. Antennaeof moderatelength,ifbent backwardextendingaboutto wing-root; scapeand pedicel
brownish black, flagellumbrownish yellgrw,the olter segments
more uniformly darkened;verticilslong and very conspicuous.
Head blackish,sparselypruinose.
Ground-colorof mesonotumblackish,the praescutumalmost
coveredby three more yellowishbrown stripesthat are confluentin front, restrictingthe ground-colorto the posteriorinterspacesand narrow lateral margins behindthe pieudosutural
foveae; scutelluma little paler than the scutum or mediotergite. Pleura grayish brown, with a completedorsal longitudinal black stripe, extendingfrom the cervicalregion to thepostnotumand surroundingthe root of the halter; ventral
sternopleuriteblackened. Halteres obscureyellow, the knobs
infuscated. Legs with the coxae obscureyellow, the f ore
coxae darker; trochantersyellow; f emora testaceousyellow,
the tips blackened;tibiae similar,the tips more narrowly darkened; tarsal segmentschiefly obscureyellow, the tips of theindividual segmentsweakly infuscated; legs conspicuously
hairy. Wings tinged with yellow,the anterior half somewhat
cleareryellow; a heavy,dark brown pattern,including a seriesof sevencostalareas,the third at origin of R-r, the fifth stigmal ; other cloudsalong cord, outer end of cell rsf M, fort<ot
M, * zt dt supernumerarycrossveinin cellM , and with smaller
spotsat ends of all longitudinalveins with the exceptionof
Rn * ,, these becomingprogressivelylarger toward th; wingbase; a cloud on vein cu just beyondits origin; veins paG,
darker in the cloudedareas. Macrotrichiain apical cells of
wing, as definedunder the subgenus. Venation: Sc, ending
shortly before fork of Rs ; nu-cuat about one-third the lengthof the relatively small cell rst Mri cell M, subequaltolts
petiole.
Abdomen black, the hypopygium more brownish yellow.

Apri't,7938 Bulletin of the Brooklyn Entomol,ogicalSociety

75

Male hypopygium with the outer dististyle a heavily blackened
slenderstructure, strongly curved to the acuteapical spine; onlateral margin at near four-fifths the length of style with asmaller subappressed
acute lateral spine.
Habitat; South Carolina, North Carolina.
Holotype: 3, River Falls, near Greenville, South Carolina, altitude 3ooo feet, Jrly r, r93r (Henry K. Toum,es,Ir.). Allotopotype: f, with the type. Paratypes,severalof both sexes,in collections of Alexander, Rogers and Townes, with following data:
GreenvilleCo., ravine on north end of Paris Mt., GreenvilleCo.,
alt. 16oo ft., April 29, rg32 (Townes, No. ril. Ravine on northeast side Paris Mt., altitude r4oo ft., May 6, rg32 (Townes, No.
20). BetweenBoy Scout Camp at CalahanMt. and Saluda Falls,
altitude r2oo-r7oo ft., June 2, rg32 (Townes, No. 26). Big
Pisgah Mt., Haywood Co., North Carolina,streamby side of road,altitude4ooo ft., July rg, rg32 (Townes,No. 3).
The present fly is amply distinct frorn all known members of
the subgenusElaeophilain the setiferousouter cells of wing andin the structure of the outer dististyle of the male hypopygium.
Limnophila (Idioptera) mcclureana n. sp.
Mesonotal praescutumand scutum dark, the posterior sclerites of mesonotumpaler; pleura with a darker transverse
girdle of mesepisternum;wings with a restricteddark pattern,
the areas broadly pale in centers,margined with darker; noextensivedarkeningat arculus; band at origin of Rs narrowand interrupted; abdomenyellow, the subterminaltwo segments blackened.
Ma,Ie.-Length rr-r2 mm.; wing 8-8.5 mm.; antenna
4.8-5 mm.
Describedfrom alcoholicspecimens.
Rostrum and palpi black. Antennae black throughout, inmale less than one-half the length of body; flagellarsegments
subcylindrical,the longest verticils less than one-half the segments. Head dark.
Mesonotal praescutumand scutum dark, probably pruinose
in fresh specimens; posteriorscleritesof notum paler. Pleurapale, the propleura and mesepisternumdarker, especiallythe
latter, where a dark transversegirdle is produced; posteriorscleritesof pleura pale. Halteres pale, the knobs weakly darkened. Legs with the fore and middle coxae yellow; femora
yellow, the tips broadly blackened;tibiae yellow, the tips more

76 Bulletin of the Brooklgn EntomologicalSociety Vot.XXXIU
narrowly blackened;tarsi black. Wings with the groundcolor pale yellow; a restricted darker pattern, including anarrow broken fasciaat level of origin of Rs, including separate areasat origin of Rs, supernumerarycrossveinin cell M
and tip of vein znd A; a secondnarrow band at cord; fasciae
broadly pale buffy in centers,narrowly marginedwith dusky;
outer end of cell rsf M, and marginal spotson veins M n and
rst A; a tiny dark spot at arculus; stigma solidly darker
brown; wing-tip narrowly and solidly darkened; cells C and
St' especiallythe f ormer, unif ormly brownish yellow topale brown ; veins yellowishbrown. Venation: Rs squareandspurredat origin ; rn-cu at near midlengthof cell tst Mr.
Abdomen elongate,yellow, the subterminaltwo segments
blackened;pleural membranenarrowly darkened. Male hypopygium with the outer dististylemore acutelypointed thanin fosciatabut lessso than in f asciolata.
Habitat.' Manitoba.
_ H olotlPe: Alcoholic 3, Fort Churchill, July 20, 1936 (M tclure); coll. No. 4 ch 68. Paratopotypes,3alcoholicJ3, Julyr3-2o, 1936; Coll. Nos. 4 Ch 54, 68.
Limnophila (Idi,optera) mcclureanais named in honor of thecollector,Mr. H. Elliott McClure,of the Iowa StateCollege. The
fly is more closelyrelatedto the EuropeanL. (/.) fasciata (Linnaeus) than to the NearcticZ. (/.) fasci,olafa
Osten Sackenbut isamply distinct from both in the wing-patternand colorationof theabdomen.
Erioptera (Psiloconopa) lacustris n. sp.
General coloration dark gray; palpi, antennae and legs
black; halteres dusky; wings with a strong blackish tinge;
veins almostwithout macrotrichia;veins R" ind Rn upcurved
at margin; malehypopygiumwith a singlesimpledististyle,its
margin with peg-likespines;interbaseslender,needle-like.
Male.-Length about 3.5-3.8 mm.; wing 4-4.5 mm.
Rostrum gray; palpi black. Antennaeshort, black throughout. Head gray; anterior vertex broad.
Mesonotumdark gray, the central portion of praescutuma
little darker but without distinct stripes. Pleura dark gray.
Halteresdusky. Legs blackthroughout. Wings with a strong
blackishtinge, the stigmal region darker but not clearly delimited; veins darker brown. Macrotrichiaalmostlacking onveins,eventhe costalserieslimited in number. Venation: Sc

Apri'\,7938Bulletin of the Brooklgn Entomolog,i,colSociety 77
relatively short, Sc, ending about opposite the fork of Rs;
R, * , subperpendicular,shorter than Rri veins Ru and R4
rather strongly upcurved at margin i Rr+ s + 4 about twice R,
alone; outer medial veins deflectedrather strongly caudad; cellM s subequalin length to its petiole; m-cu shortly beyond
fork of.M; vein znd I straight.
Abdomen black, sparselygray pruinose; hypopygium large,
black. Male hypopygium with a single dististyle,this simple,
broad at base,narrowed outwardly, the apex and lower margin
with a single row of peg-like spines. Mesal face of basistylewith abundanterect coarsesetae;apex of mesalface produced
into a slenderfinger-like lobe. Interbasea slenderneedle-likespine. Aedeaguslong, slender,straight.
Habitat; Michigan (Northern Peninsula).
Holotype: 3, Raco, on shoresof Lake Superior, M"y 9, r93S(/. R. Gross); in U. S. N. M. Collection. Paratopotypes,
numer-

ous/J.
Erioptera (Psiloconopa)Iacustriswas included in material
kindly submittedto me for examinationby Dr. Alan Stone,of theUnited StatesNational Museum. The speciesis quite distinct fromall other generallysimilar forms in the venation,unusuallyreduced
trichiation of the wing-veinsand the single simple dististyle of themale hypopygium. The most similar speciesarc E. (P.) laticeps(Alexander) and E. (P.) pilipenni,s(Alexander), both of the
Pacific Coaststates.
Erioptera (Psiloconopa) churchillensis n. sp.
General coloration yellow, variegated with brown; headdark, the front and orbits pale; praescutumwith three brown
stripes, the median one more deeply colored in f ront; pleura
narrowly lined with brown i Rr+ B+ 4 short, about one-halflonger than the basalsectionof R, ; cell rst M, open.
Female.-Length about 8 mm.; wing 5 mm.
Describedfrom an alcoholicspecimen.
Rostrum and palpi black. Antennae r s-segmented,black
throughout; flagellar segmentsoval, the outer ones not abruptly smaller, as in Trimicra. Head dark brown, the front
and broad orbital region light gray, the latter decreasingin
width behind.
Pronotum yellow, narrowly dark brown medially. Mesonotal praescutumyellow, with three brown stripes,the median
one slightly darker and wider on anterior half, paler behind;78 Bulletin of the Brooklyn EntomologicalSocietg Tot.XXXmI
lateral stripespaler brown; scutumyellow, eachlobe with two
conspicuous,confluent,dark brown areas; scutellumyellow,
with a capillarybrown medianvitta that extendsforward onto
the medianarea of scutum; mediotergitedark brown, the antero-lateralportionsbroadly yellow. Pleura yellow, narrowly
lined longitudinally with brown, including a very restricted
line on the ventral anepisternumand a more extensiveventral
stripe, involving the prosternum,ventral sternopleuriteandmeron. Halteres pale,the knobs dusky. Legs with the coxae
and trochantersyellow; femorabrownishyellow,the tips passing into brown; tibiae similar, the tips more narrowly darkened;tarsi black; legswith relativelyshort setaeonly. Wings
with a yellowishtinge; veins brown, R, and rn-cu more conspicuouslyand narrowly seamedwith brown; a pale brownseambelow tip of R, * ,, demarkingthe outer end of the yellow stigmal area. Venation: Sc, ending just before level of
R* Sc, at near midlength of the distancebetweenorigin ofRs and tip of Sr, ; R, * , * n short, about one-halflonger thanthe basalsectionof R, ; cell rsf M, open,the distal sectionof
vein M " lyins f ree in the membranein the unique type, probably connectingwith M, * , in normal specimens; ltc-cu oblique; cell znd A wide, the distal third of the vein sinuous,quite as in angulari,s.
Abdomenchiefly brown, the incisurespaler.
Habitaf.' Manitoba.
Holotype:^illcoholict, Fort Churchill,July ro, r %6 (McClure) ;Coll. No. rz Ch 45.
The nearestally of the presentfly seemsundoubtedlyto be Erioptera (Psiloconopa)angularis (Alexander) of Utah, which has
the venationof the radial field of the wing quite distinct. I am now
consideringthesetwo speciesto belongto PsiloconopaZetterstedt
rather than to Trinicra Osten Sacken. It should be noted thatvery recently Dr. Fred W. Edwards has placed Tri,rnicra as asubgenusof Erioptera.