Studia Ent., vol. 16, fasc. 1-4, outubro de 1973

261

New or ingufficientlyknown AustralasianCrane Flies. II(Diptera, Tipulidae)

Ch a rl e s P . A l e x a n d e r
Emeritus, University of MassachusettqDepartment of Entomology,
Amherst, Massachusetts( With 7 6 Iig u r e s)

The precedingpaper under this title was publishedin Studia
Entomologica,volume 74, in November 1971. In continuing thisstudy I now am treating various speciesthat were taken in New
Guinea,the SolomonIslands,and Fiji, the New Cuineanmaterials
having beentaken chiefly by Evelyn,Cheesmannin 1938and l93g;
Linsley
Cressittin
1955; Lambertus Toxopeus in 1938-193g,
J.
and Harry Hoogstraal in 1945. The Cheesmancollectionshad
been treated in earlier papers that are listed under the Selected
Referenceslater in this report but here are figured for the firsttime. The materialsfrom the SolomonIslandswere taken in lg44
by two former students in the MassachusettsState College ofthat period (now the University), Ashley B. Gurney, now in
the SystematicEntomologicalLaboratory of the United States,
Department of Agriculture,'Washington,and Robert Staples, ot
the Departmentof Entomology,University of Nebraska,Lincoln.
Materials from Fiji were taken by Mr. H. Greenwood,Elwood
Zimmerman in 1938, and Nodl L. H. Krauss in lg7l. I am
greatly indebtedto all of the above for ihe privilege of studying
these important series of specimens.In a subsequentpaper now
being preparedfurther spbciesfrom New Caledoniaare discussed
and earlier papers by the writer on this subject are includedin the list of selectedpapers.
Selectedlist of references
Alexander, C. P. New or little-known
Tipulidaefrom EasternAsia(Diptera),XX. PhilippineJour. Sci., 54: 433-471,35figs.; 1934-

(New Caledonia).
New or little-knownTipulidae (Diptera). LXXVIL Oriental-Australasianspecies.
- Ann. Mag. Nat. Hist. (11) 13:74A-767;1946(Bibliography).
The Same,LXXVIII. - Ibidem, (11) 14: 256-2&O;1947.
The Same,LXXIX. - lbidem, (11) 14: 388-414;1947.
The Same,LXXX. - Ibidem, (ll) 14: 552-577;1947.
The Same,LXXII. - Ibidem, (12) l: 391-417;1948.

26 2Alex ander ,Aus t r alas i a n C r a n e l l i e s I I

Th e Sa m e, LXXXVI I I . - I bidem , ( 12) 3 : 9 3 5 - 9 5 9 ; 1 9 5 0 .
Th e Sa m e, LXXXI X. - lbidem , ( 12) 4 : 5 7 6 - 6 0 6 ; 1 9 5 1 .
New or little-known Tipulidae from New Caledonia (Diptera). Proc.
Hawaiian Ent. Soc., 12: 235-244; 1945.
- Undescribed species of crane-flies from New Guinea in the South
Australian Museum (Diptera; Tipulidae). Records South Australian
Mus.. 8: 585-606: 1947.
- New or insufficiently-known crane-flies from New Caledonia (Diptera:
Tipu lida e) . Par t I . Ann. Ent . Soc . Am e r i c a , 4 l : 1 3 7 - 1 4 8 ; 1 9 4 8 .
- The crane-flies of New Caledonia (Diptera, Tipulidae). Trans. Rl
Ent. Soc . London, 99: 361- 393, 36 f ig s . ; 1 9 4 8 .
- Und escr ibeds pec iesof Nem at oc er ousD i p t e r a . P a r t X l l l . B u l [ . B r o o k l y n
En t. So c . , 58: 133- 138; 1963 ( New Ca l e d o n i a ) .
- New or ins uf f ic ient ly k nown Aus t r alas i a n c r a n e f l i e s ( D i p t . T i p u l i d a e ) .
Stud ia Ent om ok r gic a. 11 267- lll\ , 50 f i g s . ; 1 9 7 1 .
C h e e s m a n, Ev ely n. The land of t he R e d B i r d , p p . 1 - 3 0 0 , 2 2 f i g s . ;date unknown.
- Six-legged snakes in New Cuinea, pp. l-2ft1, 3l plates, 1l text-figs.
(Waigeu and Japen Islands, Northwest New Guinea; Cyclops andTorricelli Ranges, New Guinea, collections 1938-l939) .

-TIPULINAE
l. Holorusia illex (Alexander)Cte n a cro sce l i s iller

Alexander ;Rec. South

Austr al i an

M us .,8:588- 590; 1947.

Holotype, 6l, Torricelli Mountains,NortheastNew Guinea,
200-1,000 feet, January 1939 (Cheesman).Male hypopygium(Fig. 1).

2. Nephrotoma (Nephrotoma) atrohirsuta sp. n.
Ceneral coloration of thorax orange yellow, praescutum with threebroad black stripes, pleura yellowed, patterned with black; th,orax andcoxae with groups of unusually long black setae, as described; legs
appearing brownish black from abundant black setae, outer tarsal segmentsslightly darker; wings subhyaline, stigma slightly darker, with about 25
trichia; cell Ml short-petiolate,m-cu beyond base of cell 1sl Mr; Proximal
abdominal segments dull orange, their posterior borders restrictedly
darkened, subterminal segments brownish black, hypopygium chiefly
blackened; male hypopygium with posterior border of tergite produced
into two oval blackened plates; outer dististyle very long, inner stylewith dorsal crest very low; eighth sternite with posterior border extended
into a yellow blade that is fringed with unusually long dark colored setae.

M a l e. Length, about l3 mnt.; wing, 10.5 mm.; antenna,
al'rorrt4 nrrrr.

Studia Ent., vo,l. 16, fasc. l-4, outubro de lg73

263

Frontal prolongationof head orange, nasus darkened,withlong black setae; palpi long, proximal three segmentsyellowish
brown, the long terminal segmentbrownish black. Antennaewith
scapeand pedicelbrownishblack,flagellumpalerbrown; segments
longer than their verticils,basal enlargements
poorly developed.
Head deep orange,sides of posteriorvertex darkened;occipitalbrand dull brownishblack.
Pronotal scutum medially obscureyellow, sides blackened,with densebrushesof long black setae;scutellumblack; pretergites
darkened above, yellow laterally. Mesonotal praescutumobscureyellow, with three broad black stripes, central area narrowed
and reachingthe suturebehind,interspaces
nearly parallelsided;
humeral and lateral regions of praescutumwith groups of long
chiefly blackenedsetae; scutum obscure yellow, each lobe witha singlemajor blackened
area;scutellum,
includingthe parascutella,brownishblack, the former convex,with abundantsetae;mediotergite obscure yellow, at midlength with very abundantlong
black porrect setae, posterior third glabrous, more depressed,
pleurotergitechiefly yellowed. Pleura yellow, propleura andanterior border of pteropleuriteand meron narrowly blackened,
the broad mesepisternum
chiefly obscureyellow, including also
the middle coxae; propleuraand meron with conspicuous
setae.
Halteres short, dark brown, knob large, obscure yellow. Legs
with fore and hind coxae brownish black, mid-coxae paler, as
described,all coxae with long black setae; trochantersobscureyellow; remainder of legs appearing dark brown to brownish
black from the very abundantblackenedsetae;claw of male witha slenderspine at near midlength.Wings (Fig. 2) subhyaline,
stigma slightly darker; veins light brown, the more anteriorones
darker; stigmal trichia about 25 in number.Venation: Cell M1
short-petiolate,about one-fourth m; m-cu beyond base of celllst Mr, placedofl M4, this sectionshort, subeqvalto MB_4.
Abdomenwith proximalsegmentsdull orange,their posteriorbordersrestrictedlydarkened;subterminalsegmentsbrownishblack
to form a broad ring; hypopygiumchieflyblackened,eighth tergite
slightly reddened basally, outer dististyle yellowed. Male
hypopygium (Fig. 3) with ninth tergite, /, transverse,posteriorborder convex,on either side of midline with a blackenedoval
plate,the outer half with numerousspinoid setae,more laterally
the plate paler, with a secondrow of about l0 to 12 smaller
spinoidsctac or pcgs. Outer dististylc,d, vcry long, broadcstat

264

Alexander, AustralasianII
Craneflies

Studia Ent., vol. 16, fasc. 1-4, outubro de 1973

265

near one-third the length, thencegradually narrowedl inner stylewith beak slender,lower beak short and very broad, dorsal crest
low. Eighth sternite,8 s, large, posteriorborder convexwith a
median yellow prolongationthat is fringed on one margin withnumerousdark coloredsetae,the outer ones long and conspicuous.
Habitat: NetherlandsNew Guinea (West lrian).
Holotype,d, Station15,AraucariaCamp,altitude800 meters,
March 4, 1939 (Toxopeus); Archbold III Expedition; type inauthor's collection,through Lieftinck.
This distinci fly is most readily told Jrom regional allies by theunusually conspicuous long black setae on the thorax and coxae, as
described. The ninth tergite of the hypopygium is most as in Nephrotoma
boyesi Alexander, of eastern New Guinea, but all other hypopygial
structures are quite distinct. In this latter species vein Mo originates
before cell lst M, (see Entomological News, 80: 260; 1960).

3. Nephrotoma(Nephrotoma)augustanasp. n.

al-{1

v{

\

rc$^V
et
{i,fu

,fi
$h, \#)

Mesonotal praescutum yeltow with three light brown stripes; headyellow, occipital brand small and poorly delimited; wings subhyaline,
prearcular and costal fields slightly more yellowed; abdominal tergites
brownish yellow, posterior borders very narrwly darkene'd,sternites more
uniformly yellow; male hypopygium with apices of intermediate bladesof tergite obtuse; inner dististyle with dorsal crest low and rounded,
beak relatively short and stout; eighth sternite with posterior bordernearly transverse, median region without setae.

M a le. Length,about l1-12 mm.; wing, 9.5-10mm.; antenna,
about 3.5 mm.
Frontal prolongationof head clear light yellow, outer end
dorsally very slightly more darkened,including the nasus which
is tipped with long black setae; palpi brownish yellow, outer
segments,especiallythe last, darker. Antennaewith scapeyellow,
pedicel slightly darker, without setaeI f lagellar segments veryvaguely bicolored, brownish yellow, basal enlargementsslightly
darker. Head clear light yellow, sidesof posteriorvertex narrowly
more darkened; vertical tubercle broad, shallowly emarginate;
occipitalbrand unusuallysmall and poorly delimited,pale brown;
central area of posterior vertex virtually without setae, thesenumerousand conspicuouslaterally.
Pronotal scutum broadly yellow medially, sides infuscated;scutellumvery reduced,pale brown. Mesonotalpraescutumyellow
with three light brown stripes,Iateralsstraight, not outcurved;

I" !g. l . H ol or us i o(A l e x a n d e r) ; ma l e h y p o p y g i u m. illex
Fi g . 2 . N e p h ro t o ma
( N aphr otom a)
atr oh i rs u t a s p . n . ; v e n a t i o n . (N e p h ro t o ma )Fi g . 3 . N e p h ro t o ma
utr ohi r s uta s p. n.; ma l e h y p o p y g i u m. (N e p h ro t o ma ) a u g u s t a n a
Fi g . , t . N e i h ro t o mas p. n.; m al e hy popy g i u m. Fi g . 5 . N e p h ro t o ma (N e p h ro t o ma , s o l o mo n i s g u a d a l c a n a ra n a
Al c \i l nder ; m al e hy p o p y g i u m. Fi g . 6 . N e p h ro t o mA \ N e p h ro t o ma l s o l o mo n i s s o l o mo n i s
Afc .\ander ; m al e hy p o p y g i u m. Fi g . 7 .
N e p h ro t o mo (N e p h ro t o ms ) b y e rs i n a s p . n . ;r r i l l c hy nony gi um . S y mb o l s : b , b a s i s t y l e ; d , d i s t i s t y l e ; p , p h a l l o s o me ; s , S t h
s t e rn i t e ; t , g t h t e rg i t e

Alex ander ,2 66

At r s t r alas i a n C r a n e f l i e s I lscutumyellow,eachlobe with two broadlyconfluentbrown areas;
scutellumbrownishyellow; postnotuntclear light yellow,narrowlv
withvirtuallyglabrous,mediotergite
brown posteriorly;mesonotum
behind;sides
onvery sparsescatteredblack setae,more numerous
surrounding
pleurotergite
light yellow,vaguelydarkenedposteriorly,
andthe bases of halteres.Pleura yellow, ventral sternopleurite
light brown. Haltereswith stem brownish
most of anepisternum
yellow, with abundantblack setae,base narrowly orange' apexof knob more yellowed. Legs with coxae obscure yellow'
trochanterscleareryellow; remainderof legs light brown, apicesof femora with denseblack setae; claws long and slender,witha short spine at micllength.Wings subhyaline'prearcularand
pale
costal fields slightly nrore yellowcd; stigma inconspicuous,generalbeyondlightbrown,
veinstrichia;
eight
with
about
brown,
level of cord with abundantblack trichia. Venation:Cell R2 atnrargin slightly less extensivethan cell R3; cell M1 broadly
sessile,cell 1.s1M, small; m-cu at base of Ma, the latter shortly
Lref
ore cord, leaving a very short to punctiformelententMt-r-".
Abdonren(ntountedon slide) with tergitesbrownishyellow,
the poslerior borders very narrowly darkened,sternitesmore
yellow; hypopygiumbrownishyellow.Male hypopygiuntruni{ormly(Fig. a) with ninth tergite,/, producedinto two flattenedmedianblades,tl-reirapicesobtuse,providedwith abundantspinoidsetae,
stouterand ntorecrowdednearbases;lateralarnlaturewith similar
stout setaearrangedin two rows. Outer dististyle,d, slightlynore
than twice as long as the greatestwidth; inner style with beakrclativelysirort ancl stout, lower beak obtuselyrounded; dorsal
crest low, outer basal lobe unntodiiied,the margin with aboLttp, with apophysessrtbatrophied,trvelvestrong setae.Phallosonle,
larQe,the subbasalimpresscd
Bs,sternite,Eighthvery reduced.
line reachingthe lateral border or virtually so; posteriorbordcr
nearly transvcrse,ntedian region without setae.
Habitat: SolomonIslands.
Holotype, 8, Empress Augusta Bay, Bougainvillelsland,
APO 716, July 10, 1944 (A. B. Gurney). Paratype,61, Entpress
Augusta Bay, KorontokinaRiver, September29, 1944 ( Robert
Staples).
The most similar regional species having th,e male hypopygium
generally as in the present fly is Nephrotonta solomonis solomonis
Alexander (Fig. 6), and its subspecies,N. s. guadalcanaran.oAlexander( Fig . 5), t 5e ir y popv gia being s hor v n f o r c o m p a r i s o n .A t t e n t i o n i s c a l l e d

S l rrr l i ;r l .ttl

v ol .

l ti .

l ;rs t.

I l.

ttl tl l e cl tttr:rtl t'rs ol l l rt. l t,t l ,rl t, r.l ,' rrr,,rl r. l ,t;rL ,,1 l l rr. l rrr,,r ,l r l r.t\ 1,.
and tl te ent;trqi tt:tl t' l ){ )s 1(fror l rrrrrl t' r rrl l l rt,t.rp,trl l r..1(.i l t!tr,tl r,rt r' . Ir,,rr,l r,lw i th l ong setl tt. -l ' l tt, s l )(,(r(' s r)l l l rr. t,(,n1., l \n,,\\ll r,,nl tl rr. ..,,l orn,,rr
l sl ancl s r.l 'cre l <c )' c tl i rr ;rrt t.;rrl rr.r' |;tl )(.t l r\ l l r(. \\rtr.t(1,r,,, (.rl rl ,,ti l r,lA cad. S ci . (l rour th S eri c s ) 2' J , no. l : .l ; Irr.i r' )4. Nephrotoma (Nephrotoma) byersina sl). n.
G e n e r a l c o l or a ti o n o f h e a d a n ci th o r a x o r a n g e a n tl ye l l o w , i l i ttr o r r t
c o n s p i c u o u s p a tte r n ; o cci p i ta l b r a n d r e d u ce cl ; r vi n g s b r o w n i sh ycl l o r v,
p r e a r c u l a r a n d co sta l fi e l d s m o r e ye l l o w e d ; a b d o m e n o f m a l e w i th b a sa lf ive segments orange, of female with borders conspicuously blackenecl,
outer f cur segments blackened in both sexes; male hypopygium withbeak of inner clististyle slender, dorsal crest produced caudad; regionof outer basal lobe a strong blackened rod that terminates in an upcurveds p i n e ; e i g h t h ste r n i te w i th p o ste r i o r b o r d e r d e e p l y e m a r g i n a te , w i thconspicuous black setae.

M a le. Length, about 14 mm.; wing, I1.5 nrm.; antenna,
about 4 ntnr.
F e m a le. Length,about 16 mm.; wing, 12.5 mm.; antenna,
about 3.3 mm.
Fronlal prolongationof head yellow above, with a morc
fulvous line or.reither side of the nridarea;nasuswith vcry long
black setae;palpi with basal segmentyellow, intermediatepair
light brorvn, terurinalsegmentbrownish black. Antennaewith
scapeand pedicclyellow,first flagellarsegmentbrownishyellow,
remainder dark brown; flagellar segntentsrnoderatelyincised,longestverticilssubequalto the segntents.Head orange; vertical
tuberclc low, virtualiy entire, without setae; sides of posterior
vei'iexand genaervith abundantlong black setae;occipitalbrandvcry surall to subobsolete,
light brown.
Pronotunt and pretergitesoraltge. Mesonotal praescutulr
orange yellow with three scarcelydiffercntiatedurore polishedstripes,in the type male variegatedby darker f lecks; posterior
sclcritcsof notumcleareryellow; setaeof scutellunr
very smalland
sparse, of riieclioterqite
longer. Pleura Iight yellorv,variegatedby orangeon ventralsternopleurite,
ancpisternum
and dorsopleuralrcgion.Haltereswith stem brownishycllow, knob dalker brown,
apcx yellowed.Legs with fore coxae orange, remainingcoxae
rrrorcyellowed; trochantersyellow; rentainderoi legs obscureycllow, tarsi passinginto black; claws oi male long and slender,lvith a conspicr-rous
spur at near midlength;ventral surtacewith
Iong arrclconspicuous
lisht ltrown setac.Wings brownishyellow,
prcarcrrlar;urrlcostallickls and vcin Cl yellriwecl; stignrabrown
tvillr ;rlrr)utllJ lriclrilr;vt,inslrrorvlrt,xccJtt
irr thc ltrightcncrliiclrl:;.

Al e x a n d e r.

Studia Ent., vol. 16, fasc. l-4, outubro de 1973

Au s tra l asi anC ranel l i esII

sessilein female;vein
Venation:Cell M1 in male short-petiolate,beyond its base.
with
cord,
m-cu
shortlyatthe
Ma forking
Abdomenof male with basal five segmentsorange,succeeding
segmentsdeep yellow, the borders more orange, segmentssix
to nine brownish black; in female proximal five segmentswith
tergites orange, sternitesyellowed, posterior and lateral borders
narrowly but conspicuouslydull black; succeedingsegmentsblack,basesbroadly polishedto nacreous,margins similarly dull black,pleural membranebrownish black. Male hypopygium (Fig. 7)
with lobes of ninth tergite, l, about as shown, the intermediatepair with apicesobtuse,basal spinoidsetaelarger. Outer dististyle,d, short and broad, outgr margin at near midlengthexpandedintoa darkenedlobe; inner style with beak slender,lower beak broadly
produced caudad; region of
obtuse; dorsal crest conspicuous,
outer basal lobe with a strong blackened rod that terminates
in an upcurvedspine, basally the structureextendedto the lower
beak; vestiture about as shown, includingfour modified setaewith gonapophyses
on outermarginnear baseof style.Phallosonre
atrophied or virtually so. Eighth sternite, 8 s, large, posteriorof long black setae
border deeplyemarginate,with a concentration
relativelyshort,
impressions
subbasaltransverseat base of notch;
ending some distancebefore margin.

1A

f >J
WW
,4 .
Tf

..

I

,C5*A\)

Holotype, 6l, Empress Augusta Buy, BougainvilleIsland,along Koromokina River, altitude 45 feet, October I 1, 1944
(RobertStaples)- In thick junglebeneathlogs on spiderweb StaplesAllotopotype,I , with type.

5. Dolichopeza (Dolichopeza\ jobiensis Alexander
Do l i ch o p e za ( Dotichopeza)
3 9 2 -3 93;1947.

jobiensis

Alexander;

Ann.

M ag.

N at

H i s t.

(ll)

14:

,

,rYt<1

Habitat. SolomonIslands.

I dedicate this species to my esteemed friend, Dr. Ceorge W Byers,
outstanding student of the Tipulidae, Mecoptera, and other groups ol
insects. From other generally similar and allied species from the Solomons,
including Nephrotoma opima Alexander and N. solomonis Alexander, thepresent fly is readily told by the hypopygial structure, including especiallythe inner dististyle.The dorsal crest oI the latter is much as in Nephrotoma
boyesi Alexander, of New Cuinea, an otherwise entirely dillerent fly'

269

]//t
L

\b

\_

tq

1a

L6

Fi g .
F i g.8.
D ol i c hopez a (D o l i c h o p e z a ) j o b i e n s i s A l e x a n d e r; ma l e h y p o p y g i u m. Fi g . 1 0 .
9. D ol i c hopez a ( D o l i c h o p e z a ) d i s s e mi n a t a A l e x a n d e r; ma l e h y p o p y g i u m. (
D
ol
i
c
ho
p
e
z
a
)
Fi
g
.
1
1
.
D
o
l
i
c
h
o
p
eza
D ol i c hopez an e p h a t i a A l e x a n d e r; ma l e h y p o p y g i u m. ( D ol i c hopez a) nok en i i s A l e x a n d e r; ma l e h y p o p y g i u m. Fi g . 1 2 . Ti p u l a (I n d o t i p u l a \(P a p u a t i p u l a ).tuer el l a Al ex ander ;1 3 . Ti p u l aa rt i f e x
ma l e h y p o p y g i u m. Fi g .
Al ex ander ; m al e hyp o p y g i u m. Fi g . 1 1 . Ti p u l o (P a p u a t i p u l a ) c y c l o p i c a A l e x a n d e r;
(P a p u a t i p u l a )ma l ental e hy popy gi um .
nokicoloA l e x a n d e r;
Fi g .
1 5 . Ti p u l ahy popy gi um . F i g . 1 0 . Ti p u l a (P a p u o t i p u l a ) p e d i c o i d e s A l e x a n d e r; ma l e h y p o p y g i u m.
b,
S y mb o l s :F i g.
17. Sti ba d o c e ra p a p u a n a A l e i a n d e r;
f e ma l e o v i p o s i t o r. bas i s ty l e; d, d i s t i s t y l e ; o rl , o v i p o s i t o r; s , 8 t h s t e rn i t e ; f , g t h t e rg i t e .

Holotype,6l , Mount Oud, JapenIsland, Northern Netherlands
New Guinea, altitude 3,500 feet, November1938 (Cheesman).
Male hypopygium(Fig. 8)
s l r ,l l ,r I r t

l l J l {J

lx

27 0

Alex ander ,StudiaEnt.,vo'I.16, Iasc. 1-4,outubrode 1973

Aus t r ala s i a n C r a n e f l i e s l l

271

12. Tipala (Papuatipula) nokicola Alexander

6. Dolichopeza (Dolichopeza> disseminata Alexander
D o l i ch o p cza ( Dolichope2a.) . disseminats. Alexander ; R ec . Indi an M us ., 50: 326; 1952.
R e -n aming ol Dolichoppza lDolichop( za)fhy s he Al ex ander : Ann. M as . N at.
Hi st.. l1 ( 14) : 395- 397; 1947. pr eoccupie d by D ol i c hopez a ( D oti c hopez al thi s bt
A 'e xa nder , 1938.

Holotype, 61, Mount Lina, Cyclops Mts. NetherlandsNew
Guinea,March 1936 (Cheesman).Male hypopygium (Fig. 9).

T i putan o k t c o t a A l e x a n t l e r; A n n . Ma g '( Papuati pul a )NatHist',(1 2 ) 6 : ? 4 7 -7 4 8 ; 1 9 5 3 'Holotype,61,Mount Nok, WaigeuIsland,NorthernNetherlands
Male hypopygium(Fig. 15)'
New Guinea,May 1938 (Cheesman).

13. Tipula (Papuatipula) pedicioides Alexander
T i pul a ( Ac uti pul a)

7. Dolichopeza (Dolichapeza) nephalia Alexander
Do l i ch o p e za ( Dqltchopeza)
3 9 ? -398; 19,17.

nephalia

Alexander;

Ann.

M ag.

N at.

H i s t.,11( 14) ;

p e d i c i o i d e s A l e x a n d e r;

A n n . Ma g . N a t

Hist,(1 2 )

1 : 3 9 3 -3 9 6 ; 1 9 4 8 'MaleHolotype,61,Mafulu,Papua,January1934 (Cheesman).
hypopygium(Fig. 16). .

Holotype, d, Mafulu, Papua, lanuary 1934 (Cheesman).
Male hypopygium (Fig. l0).
CYLINDROTOMINAE

14. Stibadocerapapuana Alexander

8. Dolichopeza (Dolichopeza) nokensis Alexander
Do l i ch o p e za ( Dolichopeza)
3 9 3 -395; 194?.

nolr cnsis

Alexander;

Ann.

M ag.

N at.

H i s t.,ll( 14) :

Holotype,/, CampNok, WaigeuIsland,NorthernNetherlands
New Guinea (Cheesman).Male hypopygium(Fig. l1).

Sti badoc er a papuan aA l e x a n d e r;Ann.

Ma g .

Nat.

Hist.,(1 1 )

l4:

2 5 6 -2 5 a ; 1 9 4 7 'Holotype, I , Mondo, Papua, January-February 1934
(Cheesman).Femaleovipositor (Fig. 17).

LI M O NI I NAE

9. Tipula (Indotipula) querella Alexander
Tipuld(l n dotipulo)

quer ella Alexander ;

Ann.

Mag.

N at.

H i s t.,

( 12)

l:

Limoniini
398- 399; 1948.

Holotype,61,Kokoda,Papua,September
6, 1933(Cheesman).
Male hypopygium(Fig. 12).

10. Tipula (Papuatipula) artifex Alexander
Tipula(P a puatipula)ar tilex

Alexander ; Ann. Mag. N at. H i s t.,

( 11)

l 4:

400- 102;1947.

Holotype,61, Mount Tafa, Papua,March 1934 (Cheesman).
Male hypopygium(Fig. l3).
I 1. Tipula (Papuatipula) cyclopica Alexander
Tipula(A cutipula)

cyclopica Alexander ;Ann. M ag.

N at.

H i s t.,

( 12)

l:

396- 397; l 9J ti .

Holotype,6l , Mount Cyclops,North NetherlandsNew Cuinca,
March 1936 (Cheesman).Male hypopygium(Fig. 1 )15. Limonia (Atypophthalmus)parvapiculatasp' n.
Ceneral coloration of thorax reddish yellow, pleura with a narrow
pale brown longitudinal stripe; legs yellow; wings brownish . yellow,
itigma subcirculai, brown; vein Sc, ending about opposite two-thirds Rs;
ouier radial veins gently decurvecl;'abdominalsegments weakly bicolored,yellow, posterior borders broadly light brown; male hypopygium withiergite very large, posterior border truncate, medially produced into
an acute pbint; basistyle with ventromesal lobe large, outer part witha retrorse brush of long setae; dorsal dististyle a short stout darkened
spike, the surface with abundant delicate setulae, ventral style withrostral portion elongate, subequal to remainder of style'M a f c. Lcngth,about 6-6.2 mm.; wing, 6.7-7 mm.; antenna,;rllorrtL2- 1.3 rnrrr.
l(ostrrrrrroltscrrrt'yclkrw,palpi ligltt ltrowtt. Antcnnacwiths c ; t p t ' y t 'l l o w , r 't'ttt;ti tttl t'r l i g l tl l r r o l vr r ; l ) r {) xi l tl ;l l l l ;tg t'l l ;tr scg l l l e l l ts
l l l ( '0 i l 1 ( 't o i l ( 's ttto tt'l tttIl l tU ttt'r l , r vi l l t !i l ) ;l l sr '\'( 'l - ]'l ( ]l l K l ) l ;l ( 'k'l \ '. 't l ,27 2Alex ander ,Studia Ent., vol. 16, fasc. 1-4, outubro de 1973

Aus t r alas i a n C r a n e f l i e s I I

setae.Headof malewith eyesenlarged,as commonin the subgenus,
the reducedvertex dark brown.
Pronotumvery reducedobscureyellow. Mesonotalpraescutum
and scutal lobes reddishyellow, praescutalinterspaceswith sparsevery long erect setae;scutellumand mediotergitecleareryellow.
Pleura yellow, with a narrow dorsal pale brown longitudinal
stripe, more expanded anteriorly, becoming obsolete on thepteropleurite.Halteres yellow, knob weakly darkened.Legs withcoxae and trochanterslight yellow, fore coxae darker; remainderof legs yellow, the outer tarsal segmentsscarcelymore darkened;claws long and slender,with a single very long hairlike spine
beyond base. Wings (Fig. 18) brownish yellow, stigma brown,
subcircular; veins brown, those of base yellowed. Longitudinalveins beyond generallevel of origin of Rs with trichia. Venation:
Sc1 ending about opposite two-thirds Rs, Sc2 slightly retracted;
outer radial veins gently decurved,Ra-5 ending shortly beyond
wing tip; m-cu shortly beyond Iork of M.
Abdominal segments weakly bicolored, yellow, posteriorborders broadly light brown; hypopygium chiefly yellowed. Male
hypopygium(Fig. 2l) with ninth tergite,t, very large,as common
in the subgenus,all borders nearly truncate, posterior margin
mediafly produced into a small acute point. Basistyle, b, verylarge, ventromesallobe larger and longer than body of style,apical part with a retrorse pencil or brush of long setae,morebasallywith a darkenedlobe or tlange. What appearsto represent
the dorsal dististyle,d, is a short stout darkenedspike, its surface
with abundant delicatesetulae;ventral style with rostral portion
subequal to body of style, enlarged on basal half, outer end
slender,gently curved. Gonapophysis,g, with mesal-apicallobe
slender, apex narrowly obtuse; apex of aedeagus,a, about asshown.
Habitat: SolomonIslands.
Holotype, 6l, Empress Augusta Bay, BougainvilleIsland,along Koromokina River, altitude 45 teet, October I l, 1944
(Robert Staples)- Beneathlogs on spider webs in thick jungle.
Paratopotype,61, Hill 700, altitude 650 feet, December6, 1944
(Staples)- Under logs on banks ol a cascadingsmall stream
in jungle - Staples.
The most similar regional species include Limonia (Atypophthalmus)
goodenoughensis Alexander, oI Goodenough lsland, Papua; L. (4.)
ta ntb orin a Alex anc ler ,of Nor t h Q r r eens l a n c al ,n t l t h e s t r h g c n o t y p t , L . ( A . )ztJ

Wf,,hWffi\'j<tr&"
?l

lt

/-{.

AF
r 28
1S'
Fi g .
(A t y p o p h t h a l mu s )p a rv a p i c u l a t d
sp. n.; venation.
l l {. Li m oni al 'i g.
Fi g . 2 0 . L i mo n i a . (D-=, i c ra n o my i a )t.i i noni u ( N eatex a n d ri a ri a i i u l v i c o t o r i p . ' n . ; v e n a t i o n . p a rv d p i c u l a t a s p . . n . ;Fi g . 2 1 -- L i mo n i a (A t y p o p h t h a l mu s l
dni s ol d s p. n.; v e n a t i o n . Fi g . 2 2 . L i mo n i a (D i c ra n o my i a ) a l i e n a t a s p n . ; . ma l e n y p o p y g l u m
ffr afe ffy popy gi unr . Fj q . 2 4 .
. f.i c ,'.' i :.i . ti m o n i a(' b i c ra n o m)' i a ) d n i s o t a s p . i . ;ma l e . h y p o p y g i u m.
25
niale liynopygium--. Fi g . .
l .i non'i r .t l l ) i tr un!) n t J ' i a l g rt f u l a i A l e ra n t l e r):-Li moh y" p o p y g i u m. -, . Fi g . 2 6 . --L i mo n i a .
\ L i mon tnai o)r /i /r i na r A l c x i i n t l e r; mrl el l .i m oni ul(D i c ra n o my i a )l o rp i d a
Fi C . 2 7 . L i mo n i ar r s r l l l r r AIc r ;i r tr fe r; rt t ; t l c l ry p o p y g i t t n r.
I ' i A . 2 8 . L i n t o n i a l N (a l rx A n d ri A ri A . \ f u l v i c o l o r s p . n ;Al r r r r r r r fc r ; l r f;l l r l rV' l )r)l )y g i t l l\ v i rrl ro l s : rr, ; rc i l e i , g t t s ; b . b i rs i s t y l e ; r/ , d i s t i s t y l e ; { , g o n a p o p h y s i s ;r r r ;r l r . l r v pr i pv ai r r r rrt . { l t h t c rs i t c .

2 74

Alex ander ,ttmbrata (de Meijere), widespread in the Palaeotropics and evidenily
by means of commerce in various parts of Tropical America., All olthese differ among themselves in hypopygial structure and in detailsof body and wing coloration.

16. Limonia (Limonia) alienata Alexander
L i mo n i a(L i m onia)

olienota Alexander ;

Ann.

Mag.

N at.

H i s t.,

( 12)

275

Studia Ent., vol. 16, fasc. 1-4, outttbro de 1973

Aus t r alas i a n C r a n e f l i e s I I

l:.t06- 40?; 1948.

Holotype, 6r, Hollandia,NetherlandsNew Guinea, January
21, 1945 (Hoogstraal).Male hypopygium(Fig. 22).

Abdominal tergites and hypopygiumbrown, sternitesmore
yellowed. Male hypopygium (Fig. 23) with ninth tergite, t,transverse, posterior border truncate, anterior lateral angles
produced,united with the basistyle.Basistyle,b, relativelysmall,its area about one-third the ventral dististyle, ventromesallobe
small, setaelong, on mesal face of lobe' Dorsal dististyle,d, astrongly curved rod, outer third narrowed into an acute spine;
ventral style with rostral prolongationblunt at apex' an enlarged
spineon outer margin longer than the prolongationitself, narrowed
into an acute point; inner bristle long, at base of prolongation'g, with mesal-apicallobe a slenderstraight rod'
Gonapophysis,
Aedeaguswith each apical lobe extendedinto a long pale point'
Habitat: Fiji Islands.

17. Limonia (Dicranomyia) anisota sp. n.
Allied to pectinunguis; mesonotal praescutum obscure yellow withthree light brown stripes, pleura yellow with a narrow brown longitudinal
stripe extending from cervical region to abdomen; legs dark yellow;
wings light brown, veins darker brown; male hypopygium with ventromesal
lobe of basistyle small; rostral prolongation of ventral clististyle terminating in an obtuse lobe, the enlarged spine on outer margin longer thanthe prolongation, narrowed into an acute point.

Holotype,6l, Maikeleyaga,Kambara,altitude0-100 meters'
February22, l97l (Noel Krauss). Allotopotype,9, in copula,pinned with type.
Closest to Limonia (Dicranomyia) pectinunguis Tokunaga, ofMicronesia (Caroline and Marshall Islands), differing especially inhypopygial detaits, including the smaller ventral dististyle with its rostralprolongation and major spine.

M a le. Length,about 4.5 mm.; wing, 4 mm.
F e m a le. Length,about 5.5 mm.; wing 5 mm.
Rostrumbrown, trunklike,elongate,togetherwith the mouthparts only slightly less than remainderof head; palpi black.
Antennaelight brown; proximal flagellar segmentssubglobular,outer ones more oval, verticilssmall. Head with anteriorvertex
pale brownish yellow, about three times the diameter of scape,posteriorvertex slightly darker,with conspicuoussetae.
Pronotal scutum brownish yellow, scutellum clear yellow.
Mesonotalpraescutumobscure yellow with three light brown
stripes;scutallobeslight brown, medianarea and scutellumlightyellow; postnotumpale brown. Pleura yellow with a narrowconspicuousbrown stripe extending f rom cervical region toabdomen,passingbelow the halteres,ventral sternopleurite
palerbrown. Halteresyellowed.Legs with coxaeand trochanterslightyellow; remainderof legs darker yellow, dark brown outwardly.
Wings (Fig. 20) light brown, veins darker brown. Longitudinalveins beyond general level of origin of Rs with trichia, including
also the outer ends of Anal veins, especially2nd A. Venation:
Sc1 ending shortly before origin of Rs; cell M, open by atrophyof m; m-cu variablein position,in holotypeabout one-thirdits
Iength before fork of M, in allotypeclose to the fork.

18. Limonia (Dicranomyia) guttula (Alexander)
1915.
'nl"':,?9b
I 950.
"'*;Jx;'5*4'}r"?!';::i{i'3,,f;iii{;,"i'';l"u,^'iJ*":i:ll'f;"":\!"&:'h*:?lType oi guttula from LourencoMarques,Mozambique(PortugueseEast Africa); lype of sus f rom Hollanda,Netherlands
New Guinea, Ianuary 21, 1945 (Hoogstraal).Male hypopygium(type of guttula) (Fig.2a).

19. Limonia (Dicranomyia) torpida Alexander
I'i m oni a ( D i c r anomf i s )t o rp i d a A l e x a n d e r; A n n . Ma g . N a t . H i s t ,(l l )

l4:

4 0 7 -' 1 0 8 ; l 9 ' 1 7 'Holotype, d , Mafulu, Papua, January 1934 (Cheesman).
Malc hypopygium(Fig. 27).

20. Limonia (Geranomyia) longicrinita sp. n.
( i t'rrt.r;rl ( ol or:rl i rtt ol l l l orl tt i t tl (ttrl l l l or;l l l gt' , pl c ttra ntore y el l ow ed;l l tt' l t' rrgl l l of w i ttH ; l t' gs l l row tti s l t;rpptl rttttl tl t' 11' nttt' l l ttrtl
tl \1111
l
l
tl
||;tl
l { ' l ttt' rl t' ttt' P l l rrr l l l r' l tl rrtrtt * l rK ttt:t,
l
rttl
l
rl
,
l
,tttl
l
l
t
rrrttl"'1r'l l ,,rr.

Alex ander ,Aus t r ala s i a n C r a n e f l i e s I I

Studia Ent., vol. 16, fasc. 1-4, outubro de 1973277

central section of costa with a fringe of long erect setae, vein Sc long,Sc, ending about opposite three-fourths Rs, cell I st M, lo{rg, malehypopygium with rostral prolongation of ventral dististyle large andcomplex; gonapophysis with mesal-apical lobe long and nearly straight.

M a le. Length, excludingrostrum,about 6 mm.; wing, 6.5
2 mm.
mm.: rostrum.about
Rostrumof moderatelength,as indicated,dark brown. Headdark brown, sparselypruinose,posteriorlymore yellowed.
Pronotumsmall, scutumobscureyellow, darkenedanteriorly,
scutellum and pretergites light yellow. Mesonotal praescutumalmost uniformly brownish orange of rutous, humeral region
restrictedlycleareryellow; remainderof notum orange,scutellum
more yellowed, especially posteriorly. Pleura yellow. Haltereswith stem yellow, knob vaguely more darkened.Legs with coxae
orange, trochantersyellow; remainderof legs brownish yellow,
tarsi slightly darker. Wings (Fig. 29) subhyalineto very faintly
tinted; stigma short-oval,brown; veins brown. Costal fringe ofcentralhalf of wing very long and conspicuous,
longestat andabove Rs, the longest setae in length exceedingthe width of the
combinedcells C and Sc (as shown in subfigure).Longitudinalveins beyond general level of origin of Rs with short black
trichia,includingalso a few on distal fourth of 2nd A. Venation:
Sc long, Scr about opposite three-fourthsthe long Rs; cell 1s/
M2 long, subequal to distal section of M1-2, gently widened
outwardly; m-cu at or just beyond tork of M.
Abdominal tergites light brown, sternites more yellowed;
hypopygiumpale. Male hypopygium (Fig. 33) with ninth tergite,l, broadly transverse,with both posterior and anterior borders
nearly straight. What appearsto be the ninth sternite,s, consistsof a centralplate, its apex deeplydivided into two lobes provided
with abundant black setae. Basistyle,b, slightly more than onehalf the size of body of ventral dististyle,ventromesallobe small,placed far distad. Dorsal dististyle,d, gently curved to an acutepoint; ventral style large and fleshy, rostral prolongation large
and complex,as shownl rostrum relativelyslender,nearly parallelsided,with two large unequalspines,the outermoststrongly curved
and narrowed to a slender point, inner spine about one-half as
long and more slender(in figure shownseparatelyin two aspects).
Gonapophysis,g, with mesal-apicallobe long and nearly straight.
Aedeagusnarrow, apical lobes very small.
Habitat: SolomonIslands.

k

. $ft

!Y

)5
(G e ra n o my i a )t o n g i c ri n i t a s p . n . ; v e n a t i o n . Fi g . 3 0 . H e t i u sl('jRsl_tam29,
mnohi
oni .a.
pl tol,iima)
Fi g . 3 1 . Ta i w a i o my i al e n c s t ra t u s s p . n . l v e n a t i o -n .
talana( Al c \andc r ) : v enal i o n . 32.
l
E
t
e
p
h
t
"
a
n
t
o
my
o
d
e
s
l p c rru n c a t a s p .
lig.
e pnhoamyn t ioomy
r f.: .v eni l ti on. e lra
l i lao n g i c ri h i t a s p . n . ; ma i e h y p o p y g i u ri r.
F i g . 3 3 . L i mo n i a (G-E
.l ;i g.. i l J . Aus tr .ot i mn o p h i t a (A u s t ro l i mn o p h i t a \ c y t l o p i c a A l e x ' a n d e r; ma t e i ri b o b y E i u m.
(A u s t ro l i mn o f h i t a r'l r i A: 35.
Aus t ro l i mn o p h i l a
discoboloides
A l e ra n i i e rj --ma l e.
l r v l r opy gi r r m . .- ......F ig .
(A u s t ro t i mn o p h i t a )j o p e n e n s i s A l 6 x a n d e r;
A u s t ro l i mn o p h i l a
^
3
6
.
.
r Lr l c l l y popy gi unr . S y mb o l s : a , a e d e a g u s ; b , b a s i s t y l e ; i t , rt i s t i s i y i e ; g , g o n a p o p h y s i s ;i , i n t e rb a s e ; s , s t e rn i t e ; l , 9 t h t e rg i t e .

Alex ander ,2 78StudiaEnt.,vol. 16, fasc. l-4, outubrode 1973

Aus t r ala s i a n C r a n e i l i e s I I

Holotype, 6, Empress Augusta Bay, BougainvilleIsland,
jungle,inluly 27, 1944 (AshleyB. Curney).
Other regional species that have the male hypopygium somewhat
comparably modified include Limonia (Ceranomyia) samoona (fdwards),of Samoa; L. (G.) tanytrichiata Alexander, New Cuinea; L. (G.) vitiella
Alexander, Fiji, and probably L. (G.) vanikorensis Alexander, of the
Santa Cruz Islands, still known only lrom the lemale. OI the above theonly species with a somewhat comparable costal fringe is tanytrichiata
which has the wings patterned and showing important differences in thedetails ol hypopygial structure.

21. Limonia (Limonia) dipinax Alexander
L i mo n i a( Lim onia\

dipinax

Alexander ;

Ann.

M ag

N at.

H i s t.,( 12)

1:'108- 410;

l 9'18'

Holotype,i, Hollandia,NetherlandsNew Guinea,April 18'
1945 (Hoogstraal).Male hypopygium(Fig. 25).

22. Limonia (Limonia) insitiva Alexander
L i mo n i ainsitiva

Alexander ;

Ann.

Mag.

Nat.

H i s t.,( 12)l 95l '
'1: 581- 583;

Holotype, 6l , Kokoda, Papua,June 1933 (Cheesman).Malehypopygium(Fig. 26).

23. Limonia (Nealexandriaria) fulvicolor sp' n.
Size small (wing to about 4 mm.); thorax almost unilormly fulvous;
head polished brownish btack to black; wings tinged with light brown,veins relatively indistinct, Sc short, Sc, tar before origin of the shortRs, cell 2nd-.4 narrow; male hypopygium with tergite conspicuously
emarginate, lobes low; ventral dististyle with rostral spines long, subequal,
placei ctose together; gonapophysis small, mesal-apical lobe long andslender, aedeagus broad.

M a le. Length,about3-3.3 mm'; wing 3.8-4.2mm.; antenna'
about 0.6-0.65mm.
F e m a le. Length, about 3.5-3.8 mm.; wing, 4'4.2 mm'
Rostrumbrownish black to black, palpi black' Antennaewith
scape yellow, remainder black; proximal flagellar segments
short-oval, outer ones progressivelylengthened,verticils shorter
Head abovebrownishblack to polishedblack,than the segments.
beneathand on sidcs more yellowed.

279

Thorax almostunitormlyfulvous,humeralregionof praescutum
more yellowed. Halteres with stem yellow to brownish yellow,
knob blackened. Legs with coxae and trochanters yellowed;
remainderof legsmediumto darkerbrown.Wings (Fig. 19) tinged
with light brown, veins relatively indistinct, darker brown.
Longitudinalveinsbeyondgenerallevelof cord with trichia,lacking
on Cu1 Venation: Sc short, Sc1 ending far before origin of Rs,
Sc2 not evident; Rs very short, slightly exceedingone-half thebasal sectionof Ra-5; m-cu unusuallyvariablein position,from
oppositetork of M to some distancebetore this, in casesto onehalf its own length; cell 2nd A relatively narrow.
Abdominaltergites brown, sternitesand hypopygium usuallymore yellowed. Male hypopygium (Fig. 28) with ninth tergite,
I transverse,posterior border with a deep U or V-shaped
emargination,the lobes Iow. Basistyle,b, small, its area about
one-thirdthat of the ventraldististyle;ventromesal
lobe conspicuous.
Dorsal dististyle,d, a gently curved sickle, narrowed gradually
outwardly,apex a long spine;ventralstyle with rostral prolongation
short, tip obtuse; rostral spines long and straight, subequal,
inserted close together but not contiguous. Gonapophysis,g,with body unusuallysmall,mesal-apicallobe and slender.Aedeagus
broad.
Habitat: SolomonIslands.
Holotype, d, EmpressAugusta Buy, BougainvilleIsland,along KoromokinaRiver, September23, 1944 (Robert Staples)- beneathfallen log on side of gully. Allotopotype, 9 , altitude
40 feet, September21, 1944 - clinging to roots of banyan.
I)aratopotypes,
several d g, September19-27, 1944, commonlyin vicinityof banyans;one ol, August 1944 (Ashley B. Gurney).
In its general coloration the present fly resembles Limonia(Ntolcxandriaria) carneotincta (Alexander), of the Malay Peninsula andSingapore, differing in the small size, with darker wings and pale veins,with Rs short and cell 2nd A narrow.
'l'ltc subgenus Nealexandriaria Alexander was proposed in 1966
( l 'h i l i p p i n e J o u r n a l o f Sci e n ce ,9 5 : 1 0 7 - 1 0 8 ) ,w i th 1 8 sp e ci e s w i d e - sp r e a d
i r r t h c O r i e n t a l a nd Au str a l a si a n r e g i o n s. Th e va r i o u s sp e ci e skn o w n fr o m
l l t e v a r i o r r s P a ci fi c i sl a n cl s i n cl u d e L . ( N .) a tr o m a cu l a ta ( Ed w a r d s) ,o f S i t n r r r i r ; / . . ( N .) o ch r i co p i l l tt Al e xa n d e r , Fi j i ; a n tl L . ( N .) se m i r u fa
( l : t fw i r r t fs ) , N c w l l e b r i r l cs. L . ( N .\ sco l o l ti u Al cxa n d e r , o f To n g a , i s
c t t l i r c l y t l i s l i r r c l fr o r r r l l l l h c;r b o vc i r r h yp o p yg i a l str r r ttr r r c.

Alex ander ,2 80

Aus t r a l a s i a n C r a n e f l i e s I l

Stuclia Ent., vol. 16, fasc. l-4, outubro cle 1973

24. Limonia (subgenus incertum) dactylolabis (Alexander)
L i mn o b i d( ?)dactylotabis Alexander ;Ann.

l !{ag. N at.

H i s t.,

( 9)

8:

552- 553; 1921'

The unique type of this fly was a nale, taken at Lautoka,Fiji, April 11, 1920,by H. Greenwood,preservedin the British
Museum (Natural History). More recentlytwo female specimens
have been secured,both from Viti Levu, Fiji, one at Tholo-i-suva,
500-600 feet, July 31, 1938, by Elwood Zimmerman,the otherfrom the Nausori Highlands,Fiji, at 500-600 meters, Octoberl, 1970,by Noel Krauss.
The holotypewas leglessand the distinctivecolorationof
the legs as shown by the females,is described.Femora andtibiae dark brown, with about the outer tenth of the latter snowy
white, this color also including tarsal segmentsone and two,
togetherwith the extreme base of the third segment'torming avery broad white ring, that oi the fore leg being about subequal
to the darkenedremainderof the tibia, on the posterior legs
about two-thirds to three-fifths of this darkenedpart.
In somerespects,especiallythe venationand leg coloration,
the fly suggests speciesof the subgenus Thrypticomyla Skusebut the hypopygial structure is nore as in typical Limonia. The
in the original
positionof this fly was questioned
strict subgeneric
doubt.
instill
remains
descriotionandtltc interspacesblack or brownish black; posterior scleritesof
rtotum,includingscutellum,dark brown. Pleura brownish black.
llaltereswith stemyellow,knob brownishblack. Legs with coxae
and trochantersbrownishblack; femorayellow basally,apex darkllrown, enclosinga silvery white ring that is broader than thesubterminaldarkening,beyond the white annulus with a veryllarrowblackenedring, the extremetip pale; tibiaeyellow,extreme
tips more whitened;tarsi yellow.Wings (Fig.30) with the groundwhitened,prearcularand costalfieldsmorewhitened;a very heavy
and conspicuous
brown reticulatedpattern that is only slightly
lcss extensivethan the ground; a completeband at cord, witha narrower nearly completeone before origin oi Rs, interrupted
in cell Cu, two very heavy solidly darkenedareas in cell Rs,
lacking in cell R1 before the stigmal band; dark areas beyond
cord heavy,especiallynear wing tip, leavingoval whitenedgroundspots; centerof cell lst M2 with a similar relativelysmall oval
pale area; basad of cord and excludingthe proximal band thetransverselines narrow; a prearculardarkeningbehind ft,' veinsbrown, yellowedin the costaland prearcularinterspaces.
Macrotrichia on veins Ra-5,outer sectionof Mr-" and M3. Venation:
Sc long, Sc' ending nearly oppositer-m; distal sectionof Rs
nbout two-thirdsr-m; branchesof Rs stronglydivergent,cell R3
at rnargin very extensive;cell lst M2 small, the veins beyond
crrrrespondingly
lengthened;
m-cu about one-halfits lengthbefore
irrrk of M; Anal veins curved strongly into the wing margin.
Abclonrendark brown, without clearly evident pattern.
Habitat, NctherlandsNew Guinea.

25. Helius (Rhampholimnobia) fenestratus sp. n.
Belongs to the papuonus group; size small (wing of lemale. 4'2mm.) ; mesonotal praescuium with three chocolate brown stripes, remainderof th,orax dark brown to brownish black; femora yellow, apices darkbrown, enclosing a relatively broad white ring that is followed. by anarrow darkening; wings whitened, with a very heavy reticulated dark
pattern, beyond ihe cord and in cell 1sl M, appearing as whitened oval
areas surrounded by dark; abdomen dark brown'

F e m a 1e. Length,includingrostrum,about 6.5 mm.; wing,
4.2 mm.; rostrum alone about 0.6 mm.
Rostrum moderatelylong, brownish black, mouthpartspale.
Antennae black, the proximal two flagellar segmentstestaceous
yellow. Head dark gray.
Cervical region black. Pronotum light brown, sparsely
pruinose. Mesonotal praescutum with three chocolatc llrown
stripcs,in front separatedby line.sof light gray p0llcrr,bclrintl

281

Holotype,9, Hollandia,lanuary21, 1945(Harry Hoogstraal).
'l 'h e o t h e r r e g i o n a l sp e ci e s h a vi n g a n u n u su a l l y h e a vy d a r ke n e d w i n g
l r i r l l ( 'r n r r r r t l l a c k i n g a cl a r k a r e a i n ce l l R i s H e l i u s ( R h a m p h o l i m n o h i a )t , i t : t 'r r t i r t u t r tAsl e xa n d e r w h i ch d i ffe r s";Jf,? l r tt i n th ,el e g a n d w i n g p a tte r n s.

'l'aiwalt onr yia t alana ( ; \ lt 'xr r r r t lt ,)rlli
It'tt"ttJttltt/rrl,rrr,r,\1,'.rrrrl|r| 1 , , 1 , , t 1 , p r., i ,t( l r ,,!i l .i l 1).\rrrr\l.rr'\.rlllr'.t

III)

II

ir,l

ir,:'.

l'lll'.\l r r r r r r l | ,r l ,r , l ';r l r r r ,r , S,.r o ( ) [t.r '1 , A1 ;r r rl t\'in,llt,'ll

t l rl 'll)l 1 ) .1 .1

StucliaEnt., voi. 16, fasc. 1-'1,outubro tle l97l.i

Aus t r a l a s i a n C r a n e i l i e s l lAlex ander ,28 2

27. Asstrolimnophila (Austrolimnophila) cyclopica AlexanderA u stro l i mnophila ( Austr olimnophila\l 3 : 713- 7- 15; l9' 16.

cJ' ctopic a Al ex ander ;Ann'\'IagN at(II)Hist 'New Guinea,7,500
Holotype,6l, Mount Cyclops,Netherlands
teet, March 1936 (Cheesman).Male hypopygiun (Fig' 34)'

2g. Austrolimnophila(Austrolimnophila)discoboloidesAlexanderA u stro l i mnophilo ( Austr olim nophila)(1 1) 13: 7- ll- 7' 13; 19' 16

discobo tol dos Al e\ander ;Ann'\l ag

Hist

N at

'

Holotype, d1, Mount Tafa, Papua, 8,500 feet, March lg34
(Cheesman).Male hypopygiun (Fig. 35).

29. Austrolimnophila (Austrolimnophila) japenensis AlcxanderA u stro l i m notlhila ( Austr olim nophila)
tl l:7.17- 7ltl;19J( i.

iaptilnsls

Al ex andc r ;Ann'N l ag N at

Hist

'

(l l )New Guinea,
Holotype,6l, Mount Oud, NorthernNetherlands
hypopygiuttt
Male(Cheesnran).
1938
November
3,500 f eet,(Fig.36).

3U. Elephantomyia(Elephantornyodes)percufleatasl). n.

pattern i ncl uding a cloud at bend of Rs, base of R2- 3- r and t hc
cord; vague b r own seanls behind vein Cu, and in cell 2nd A:
prcarcul ar f i eld concolor ous r vit h r em ainder of gr ound; veinsbrow n. Longi t udinal veins beyond cor d wit h t r ichia, lacking on
tl tc crcct ba:e of R2- 3- aand M 3- . r ; basad oi cor d wit h scat t er ed
l ri chi a on R s and wit h t hr ee or f our on out er hal{ of Cu, ; asi ngl c tri chi r-rr non 2nd A, none on lst A. Wings cuneif or m ly
narrow cd at b ase, anal angle lacking. Venat ion: Sc1, ending
shortl y before f or k of Rs, Ac, at it s t ip; R2_3_aper pendicular
at ori gi n, bcyo nd base ext ended par allel and ver y close t o R1- 2;
basal secti on o f R5 in longit udinal alignm ent wit h it s dist al sect ion
arrd w i th R s; cell I st M 2 long, about equal t o dist al sect ion of
Mij; m-cu at or beyond one-third the length of M13-a;cell 2nd A
rtarrow , thc vein long.
A bdonren conspicuously bicolor ed, bases of int er nediat es(' gnl entsbroadly yellow, appr oxint at cly t he out er halves br ownish
bl ack,:evcnth and eight h segr r ient s black; hypopygiunr chief lyrl arl < brow n.
H abi tat: New G uinea.
Fl ol otype , d, O kait adi, Nor t heast New G uinea, alt it ude 1760
nl eters, A r" rgust7, 1955 ( J. Linsley G r essit t ) , Bishop M useunr .
Distinguislrecl
from th,e rjtlrer resional membersof the subgenusbyl l tc venati onand t he snowy whit e int er m ediat et ar sal segm ent s.

Ce ner al c olor at ion of t hor ax y e l l o w , t h e m e s o p l e u r a a n c l s t c r n u mexten s iv ely t iar k br own t o f or m a br o a d g i r c l l e ; l e g s d a r k b r o w n , t a t s a lsegmentsiwo an4 three white; wings weakly s1f [uset1,restrictecilypatterned
rvith d ar k er br own, c uneif or m ly nar r o w e d a t b a s e , a n a l a n g l e l a c k i l l g ,cell 2rtci A long and narrow; intermeciiateabdominal segments bicolorecl,ca ci, yelk r w wit h out er half blac k .
M a lc .

Lengt h, wit houthc ac l , a b o u t 7 m n l . ; w i n g ,8 '5 r l l t r t .

Head lost. Mesonotuntobscureyellow. Pleura yellow postgirdle on nlesepisternum
eriorly,propleuraand a broad transverse
brown, the latter extendingfront the dorsopleuralregionventrad,
includingthe sternalregionwith the exceptionof a narrow yellow
midline. Halteresblack, stent long and very slender.Legs withcoxaeand trochantersyellow; femora dark brown, basesbroadly
paler; tibiae and basitarsidark brown, secondand third tarsal
segnlentswhite, outer segntentsblack; iemora and tibiac with afew erect black setaescatteredover the entile length,on tibiae
iotall ingabout sevenor eight in number.Wings (Fig.32) wcakly
palcr llrowlt
sr.rffusi:cl,cells C and Sc clarkerbrown; a rc'strietecl

2lJ:J

Eriopterini:i l. Teuchclabis (Teucholabis) dimelanocycla sp. n.
. \ l t 's r t l r o r a x b l a cl <,I a r i e g a te d b y ye l l o w o n p r a e scu tu m a n d scu te l l u m ;r , , l l r n l r r r r t l l r a l te r e s b l a cl <; fe m o r a ye l l o w w i th tw o b r o a d b l a cl < r i n g s
I l r , r l : r : 1 . n l o i c e xte n si ve th a n th e i n te r m e d i a te g r o u n cl a n n u l u s; w i n g s'. r r l r l r \ '; r l i r r ts',t g r n a a n r l a n a r r o w m a r g i n a l se a m i n o u te r r a d i a l fi e l clr l . r r k l r r o r v n ; c n sl l r l {r 'i i r g e l o n g , Sc, e n r l i n g a b o u t o p p o si te tw o - th i r cl s
/ , ' , . t n ; t l t ' l t t 'p o p1 ,g i 1 1$n 1i th b o th cl i sti styl e sb l a cke n e cl ,i n n e r styl e sl e n cl e r .
l . t.r r g tl r , a l r o r r l ( i l l l n l .; w i n g , 5 .4 n tr n .
I t ' r r r ; r l t ' . l .t'r r g l l t, a l r o r r l fr r r r r r r .; w i n g , 5 .6 r r r n t.

A 1 ; r l t '.

t( rr,ttn(llrlr l ;r ek, l o r r ! l n tl sl t'n r l t,r ., ( ) n l y ;r l i ttl e sl r o r tr ,r l l r ;r r r, r l lr r ';ti l ; |;r l Pr l r l ;tr 'l i . ,,\n l l r r r r ;r r . l r l ;r tl i l l l o r r q l r o ttl ; l i r sl

l l ,r r ,,, !'l ttr

l i 's l I r r r r r
l l .i l: 'r ,

tll

l ,,r r t , . rl ,l , l , ' l rt t l , i l ,, , r, rl .

, , l l ( ( { ( rl rt t i ,rr rl r,

tl,

, rl

oll,

' . r' i , t i l r' i l 1 , ,
t

.,{

t,llr

. , l t rrt l o \ ' ; t l ,rri l l (' tr l,,t|',tlr

l l , . r, l

rt l . ,28 4Alex ander ,Studia Ent., vol. 16, fasc. 1-4, outubro de 1973

Aus t r al a s i a n C r a n e l l i e s I I

polishedblack,front and anteriorvertexlight gray pruinose,sidesof occiput and posterior vertex more brownish yellow'
Pronotumchestnut,blackenedin front. Mesonotalpraescutum
polishedblack,humeralregionand a smallersublateralareabefore
suture more yellowed; scutal lobes uniformly black, central areain front with a light yellow spot that crossesthe suture into theblack; postpraescutum;scutellumbrownishyellow, parascutella
yellow,
regiondorsopleural
black,
inmale
notum black. Pleura
ventralinfemale,anteriorly extendedventrad to the fore coxae;
pleuritespaler. Halteresblack, in male includingthe knobs except
for a vague paler apical line, in femalemore extensivelyyellowed.
Legs with fore coxae black, middle and hind coxae brownish
black, apicesbroadly yellowed; trochantersbrownish yellow; fore
and middletemorayellow with two broad black rings, one apical,
the other beyond midlength of segment,the intermediategroundring narrower,posteriorf emora similar, basal third blackened;
tibiae and tarsi black; claws in both sexeswith a lateral spine'
Wings subhyaline,the very small stigma and a narrow marginal
seamin outer radial field dark brown; veinsbrown. Costalfringeof male long and conspicuous;trichia of veins sparse'beyond
cord includingshort seriesat outer end of R+ and virtually allof Rr and M*2, M3 with a single trichium.Venation: Sc long
Scl ending at near two-thirdsRs, Sc2 about oppositeone-fourth
the length; R1-2 and Rr-r-o subequal;cell I st M2 subequalin
length to distal section ol M;2; m-cu about one-half its length
beyond fork of M.
Basal abdominaltergitesdark brown, outer segmentsvague'ly
bicolored,their proximalparts slightly paler, the conditionbetter
shown on sternites;hypopygiumblack. Sternal pocket of fifth
segmentnot developed.
Male hypopygium(Fig. 0) with posteriorborder of ninth tergite, l, convexly rounded. Dististyle, d,subterminal,outer style larger, appearingas a darkencdblaclc,
with a smaller subbasalpoint; inner style slender,apex acute.
Habitat: New Guinea.
Holotype, 61, Itouda NortheastNew Guinea, altitude 1500
meters,August 13, 1955 (J. Linsley Gressitt).Allotopotype'9,
August 14, 1955: Bishop Museum.

285

45
fr l x :J 7. 7'tuLl tol abi s (Te u c h o l a b i s ) s e t i g e ra s p . n . ; v e n a t i o n . Fi g . 3 8 . G o n o my i agor ok anu s p . n . ; v e n a t i o n . l l r l i oc r r u)
Fi g . 3 9 - G o n o my i a (C o n o my i a )ma l i t i a
r
f
.
\'c
l
r
i
l
ti
or
.
Fi
g
.
J
{
).
(Te
u
c
h
o
l
a
b
i
s
)
;
Tru
t
h
o
l
o
h
i
s
d
i
me
l
a
n
o
c
y c l a s p . n . ; ma l e:l r
fr y l r r r l r y gi r r r r r -.
l ;i g. t l , 7 ' ? u (h o l a h i s (Tt u t h o l a b i s ) g o ra n d A l e x a n d e r; - ma l e h y p o p y g i u m.
l 'l A. .12. 'l 'tuc h o l a h i s (Tt u c h o l a b i s \s ? t i c o s t a A l e x a n d e r; ma d e h y p o p y g i u m. -.
.
l :l 'l 'tutht,l uhi s (7 ' ru (h o l a h i s \s r/ f g rra s p . n . ; ma t e i ry p o p y g i u m. ' --" Fi g .
++.
! ln
l i t'i l t,ti l r i tt tl r l i i l r tr ut
i n t p u y i d u A I c r; t n d e r; l l a l e l l y p o p y g i u n r. --Fi g .
J 5 . G o i o ml , rc
l l l l 'ul tl tl r t,\t {r ,( r r r , A I c \ rrr(l c r; rra l e l )y p o l )y A i u [ r. l ; i g . { (i . G o n o m)' i a (L i p o p h f t p s )'h:isistylc';
I'r r \/r 'tr r l ,r s AI.r ;r r r r l c r;
. ri l l f
l t v l )o l )y A i l u i l . ' S y rl b . l s :c , l c t l c a A t ri ;D,r /, r l i s l i s t y l u ; / , ; rl r; rl l 0 s o rrrt ; s , l ' rl l r s l rrrri t t : / , 1 [ l l l c rrl l r.

The species is most like Teucholabis (Teucholabis) biarmillata
Alexander (Koroda, Papua, 1200 feet, July 1933), differing in the cletailsof coloration of the body, legs and halteres.The structure of the hypopygiumof this latter srrecies has not been clescribed.
llrr,lt.r

I rrl

|

='J l-r

ltl

2 86

Alex ander ,S t u di a En t., vo l . 1 6 , fa sc. l - ,1 , o tttu b r o cl e l !) 7 .i

Aus t r a l a s i a n C r a n e f l i e s I I

32. Teucholabis (Teucholabis) gorana Alexander( Teucholohis)
T cu th o l a bis
2 7 1- 273; 1917.

gor ana

Alexander ;Ann.

Al ag.

N at.

H i s t.,

( 11)

1.1:

The type was frc)nrGora, Papua.Mesothoraxreddishyellow,
posteriorhalf of praescutumwith two major black areasthat are
separatedby a narrow ground vitta; halteresobscure yellow;
legs with fore fenrorayellow,outer fourth black, middle and hindfemora yellow, each with a narrow brown beyond midlength;
wings obscureyellow,pattcrncdwith brown, includingthe broad
apex; abdomen orange, hypopygiumabruptly blackencd.Malehypopygium(Fig. 4l) with basistyle,b, obtuseat apex.Dististyles,d, subterminal
outer
style
a
simplecurved blackenedhook,;narrowedinto a long acute spine; inner style larger, strongly
curved,terminatingin a spinouspoint, near base with a large
lateral arrn that is approxinrately
as long as the style itself.

33. Teucholabis (Teucholabis) metamelaniasp. n.
General coloration polished yellow ancl black, praescutum with threenearly conlluent black stripes, postnotum and anepisternum black; rostrum
u nu su ally s hor t , obs c ur e y ellow; k no b o I h a l t e r e s y e l l o w e d ; l e g s r v : t 't
fore coxae blackened; wings pale yellow, restricteclly patterned withpale brown, Sc long, cell 1.sf M, short; abdominal tergites uniformly
brcrwnish black, sternites yellow.

Female. Length,about 6.5 nrnr.;wing,5.4 mnt.
Rostrumunusuallyshort,only aboutone-fourththe remainder
blackened;palpi brownish
of head,obscureyellow,apexrestrictedly
black. Antennaedark brown; basal flagellar segmentsoval, tlrc
outer onesmore elongate,shorterthan their verticils.Head black,behind and beneathbrownish yellow, sparselypruinose.
blackenedin
Pronotalscutum obscureyellow, conspicuously
front and on sides,scutellumnroreyellowed.Mesonotalpraescutunt
chietly coveredby three highly polishedblack stripes,the paleinterspaces
vaguelyindicated,centralstripe ending sone clistancc
beforesuture,lateralstripesvery large,their anteriorendsreaching
the margin, the yellow ground restrictedto the narrow humeri
and laterallybeforethe suture;scutunrobscureyellow, each lobe
polishedblack area; scutelluntobscureyellow,
with a transverse
narrowly brownishblack nrediallyat base, setaeof scutal lolres
cntircly
postnoturttand scutellumsparseand weak,inconspicuous,with long black st't;tt.
blackcned,cephalicborder of plcurotcrgite

2S 7

l'lcura with propleura and anepisternuntblackencd,tltc lattt'r';trea isolated by the yellow dorsopleuralregion, silvery ycllorv
ptcropleurite
and a narrow line behindthe propleura;nletaplcura
anclmeralregionlight yellow.Haltereswith stctttweaklydarkenccl,
remainingcoxac
knob light yellow.Legs with fore coxaedarkenccl,
lrrrclall trochanterslight yellow; remainderof legs broken.Wings
pale yellow, stigntasntall,dark brown; three much paler brownbands,the most evidentone at cord, secondarea interrupted,atorigin of Rs and more diffusely in cubital and anal cells, thirdrfarkeningat outer cnd of cell /s/ Mr; ccll Sc nlore yellowcd;
vcins light brown, scarcelydarker in the patternedarcas,C, Sc;rnd R more yellowed.Weak trichia on all outer veins beyond
crrrd and at outer end of l st A. Vcnation:Sc long, Scr ending
about oppositcthree-fourthsto {our-fifthsthc long p.s, Sc, only
short;
slightlyrenroved,
Sc1 beingsubequalto R2 or to r-nt; R2-3-aabout
one-half
ccll /.sf M, shorter than veins beyond it; nt-cu
its lcngthbeyondfork oi M.
Abdominaltergitesunifornilybrownishblack,sternitcsyellow.
( )vipositor with ccrci horn yellow, strongly upcurved to the;rcutctips.
Habitat: New Guinea.
llolotype, 9, Tsenga,NorthcastNew Guinea,July 14, 1955(.f. Linslcy Grcssitt); Bishop Museum.
'l 'h e p r e s e n t fl y i s d i sti n g u i sh e dfr o m o th ,e r r e g i o n a l sp e ci e s b y th er r r r r r s r r a l l sy h o r t a n d sto u t ye l l o w r o str u m a n d th e d i sti n cti ve p a tte r n o f
l l r t , t l r o r a x a n r l a b d o m e n . Oth e r sp e ci e s h a vi n g th e kn o b s o f th e h a l te r e s
v t . l l o w c t l h a v e th ,e r o str u m b l a ck a n d m o r e e l o n g a te .

.t,1. Teucholabis (Teucholabis) nigrircstris Alexandcr|'

tt, lt',lrtttt:
I I I ut lr)lul)i\)
ll., il I.
l!ili

Ale\ander;

llilrlfoslris

Ann.

Nat.

llag.

Hist.,(ll)

1'l:l )t' st ri l rl r l f r or r r var ious st at ions in Papua, collect ed by M iss( l ri r' !nr;ur. l ' rr r 'llr t 'r t 'r lcnsions oJ r angc include t he f ollowing:
Kor ip, Jr r ly 12, l( ) 55; M ir at t t ar ,N , rrl l rt' ;rs l Nt 'r r ' ( it t ir t t 'r t(
irt'ssilt
).
(
|
.l
7,
l
(l
.r
r
I
ir
r
slr
'1
l rrrr,

Lrt,
| '

;

l r.

rl ,rl tt\,rtl l t

l ',r1't;.1',l .rl r,'r

.rl ltl tt,l ,l i trr|!'rrrrr.

/l )O( rrr,i l rrr\rr,,rr I

llol

rttr l l t,,
l .' tr, rl rr,l,\l ottl tll );tv ttt;ttt,l rrl t'' \l .rl r,ri lt(r,,,l l t,\

A ' 1;tl tt' l ttt Ii l rrtgr' ,l ' l r.i((i t' rrl l ttIi,,tl l tl tt,l ,I rl l\1

l ,rl ,

I l .' O,t11tttt l t t' ,,28 8Alex ander ,StudiaEnt.,vol. 16, fasc. 1-4,outubrode 1973

Aus t r a l a s i a n C r a n e f l i e s I I

35. Teucholabis (Teucholabis) seticosta Alexander
T e u ch o l abis ( Teuchotabis\
4 1 3- 111: 1947.

seticosts

Alexanc l er ;

Ann.

M ag.

N at.

H i s t.,

( 11)

l 4:

Type from Mount Nok, Waigeu Island, NetherlandsNew
Guinea, taken by Miss Evelyn Cheesman.
Mesonotumyellow, praescutumwith three confluentpolished
blackstripes,scutellumblack,sidesof postnotumyellowed;halteres
infuscated;middle femorayellow basally,outer three-fifthsblack,posteriorfemorauniformlyblackened;
restrictedly
wingssubhyaline,
patterned with darker, costal fringe of male long; abdominal
tergites darkened, sternitesand hypopygium yellow; male withsternal pocket of fitth abdominal segment(Fig. 42, s) distinct,
subcircularin outline. Male hypopygium (Fig. 42) with outerdististyle,d, slender,curvedinto a long axial spine,with a smaller
straight lateral branch; inner style proloundlydivided near base,the more slenderarm graduallynarrowedinto a slightly curved
point, some distancebefore apex with two approximatedelongate
setae,inner arm a more flattenedblade that narrowsto an elongate
curved noint.

36. Teucholabis (Teucholabis) setigera sp. n.
General coloration yellow, patterned with black, on thorax including
three confluent praescutal stripes, scutal lobes, scutellum and mediotergite;
pleura reddish brown with a silvery gray ventral stripe; halteres andposterior lemora black; wings subhyaline,restrictedly patterned with brown,
macrotrichia of veins sparse; sternal pocket of male abdomen lunate;male hypopygium with outer dististyle and basal arm of inner style bifid.

M a I e. Length,about 7 mm.; wing, 6 mm.
F e m a le. Length, about 7 nm.; wing, 5.6 mm.
Rostrum and palpi black. Antennaedark brown; flagellar
segmentsoval; normal vestitureshort, yellow, verticilsexceeding
the segmentsin length. Head black.
Cervicalregion black. Pronotumpolishedorangeyellow, moredarkenedanteriorly, medially and behind. Mesonotal praescutum
polishedyellowwith threeblack stripesthat are virtuallyconfluent,
the medianarea widenedand slightlybifid behindto leavea Iarge
yellow area betore suture, humeral region and Iateral borders
yellow; scutum yellow, each lobe with a major black area andwith severalIong erectto porrectsetae;scutellumbrownishblack,brownish black,with scveral lons black setac; me'cliotergite289

abruptlyreddishyellow,with Iongyellowsetae.Pleura
pleurotergite
slightly more reddish brown, the meron,yellow, mesepisternum
ventralpteropleuriteand dorsal sternipleuritewith a narrow silvery
gray longitudinal stripe. Halteres black, knobs large' Legs withcoxae reddishyellow; trochantersyellow; a single (posterior) leg
remains;femora black, extremebase yellow; tibiae and basitarsi
yellow, tips very narrowly blackened,remainderof tarsi broken;
Wingvcstitureof leg (female)relativelyshort and inconspicuous.
outer
male
in
yellow,(Fig. 37) subhyaline,costal border lightvein
asrnargin of radial field narrowly darkenedas far distad
Rs; stigmasmall,oval, dark brown; a vague darkenedseamover
cord, best indicated by a deepeningin color of the veins which
elsewhereare light brown. Costal fringe of male relatively long'
Macrotrichiaof veins relativelysparse,basad of cord lacking,includingSc and both Anals;both outerbranchesof Rs with trichia
virtually throughouttheir lengths.Venation:Scr ending opposite
or shortly betore midlength of Rs, R2-3-alacking or unusuallyshort, less than one-half Rzi cell 1sl M, long, subequal to or
fongerthan distal sectionof My2; m-cu abouttwo-thirds to threefourths its length beyond fork of M.
Abdomen chiefly yellow, basal tergites with a weak central
Setigerouspocket on sternitefive (Fig' 43, s) aboutclarkening.
as figured,appearinglunate,gentlyconcave,with numeroussetaeon cephalicand lateral parts, these directedcaudad and mesad,
Male hypopygium(Fig' 43) with outerdististyle,chieflydecussate.
r/, snrall,strongly curved to the acute tip, on outer margin with
an appressedspine; inner style including a narrow blackened
lrcak, the apex obliquely truncate, with a conspicuousbasal
lrispinousarm, the long axial spine gently curved'Aedeagus,a,lelatively narrow, central area at apex farther produced; twolrrge setaeat ncar midlength,with two pairs of smallerbristles
lrt'[ort'apex .
ll;rbitat: Ncw Guinea.
llololypc, i, Busu, NortheastNew Guinea,September15,(i. l-irrslcyGressitt); Bishop Museum.Allotopotype,I.
11155'! 'r r t t l t r t l t t l ti s ( 'l t'ttr h ttl u tti s) st'ti g e r a i s d i sti n g u i sh e d fr o m o th e r
r : t 'r r c t ; r l l ys i t t t i l;tr r t'g i o r tr tlsl tcci e sth a t l ta ve th e r o str ttm , scu te l l u m ,h a l te r e s; r r t r l p l s l t 'r i o t l ( 'tl to r ;t l r l r ttl <ettctl h y th c h yp o p yg i a l str ttctttr e , p a r ti ctr l a r l y
l l r r l r ; t s ; r li t n n ol l l t( .i r r r r e l tl i sti sl yl t'.'l 'l r e sctttc o f tl tc tl to r ;tx i tr c tl n ttstta l l y
l ,'ttl :;r tt,l| i l i l \l rt r i l i l i l \ ,\ i l 1 ' H (' \ l rrt t 'lltt's|( ( ill(l l i l l l l (' .

2 SOAlex anc ler ,Aus t r a l a s i a n C r a n e f l i e s I I

29rStudia Ent., vol. 16, Iasc. 1-4, outubro de 1973

37. Gonomyia (Gonomyia) malitia sp. n.
Mesonotum brownish black, scutellum yellow, pleura conspicuously
patterned with brownish black and yellow; rostrum tight yetlow; antennae
black throughout; legs with coxae dark brown; wings subhyaline, Sc
short, Sc, ending opposite origin of Rs; abdominal tergites dark brown,sternites slightty paler.

F e m a le. Length, about 5-5.2 mm.; wing, 4.8-5 mm.
Rostrumlight yellow, contrastingwith the black palpi. Antennae
black throughout;flagellarsegmentslong-oval,with a short dense
white pubescence,verticilsshort. Head gray.
light
Pronotumdarkenedmedially,yellow on sides,pretergites
black,
yellow. Mesonotalpraescutunrand scutal lobes brownish
sparsely pruinose, pseudosuturalfoveae black; scutellum andanteroblackened,
medianregionof scutumyellowed;mediotergite
lateral angles yellowed.Pleura dark brown to brownish blackand meron,theon propleura,anepisternum,
ventralsternopleurite
remainderlight yellow, forming a broad stripe on dorsal sternopleurite, pteropleuriteand meron. Halteres infuscated,base ofstem narrowly yellowed.Legs with coxae dark brown, fore pairstill darker, trochantersbrown; remainderof legs broken.Wings(Fig. 39) subhyaline,
stigma barely indicated;veinsbrown. Veinsof about the outer four-fifthsof wing with long trichia.Venation:
Sc short,Sc1 ending oppositeorigin of Rs, Sc2 shortly removed,
Scl about as long as basalsectionof M1-2; m-cu aI fork of M.
Abdominaltergitesdark brown to brownishblack, including
the genital segment,sternitespaler brown. Ovipositorwith cerci
elongate,slender,gently upcurved.

ffv

Jho

fl?
47

k

A

./'

p'\

5Z

Habitat: New Guinea.
Holotype, I , Goroka, NortheastNew Guinea,altitude 1550
meters,June 25, 1955 (J. Linsley Gressitt); Bishop Museum.
Paratopotype,I.
The present fly is quite distinct from the only other regional memberof th,e typical subgenus so far described, Gonomyia (Gonomyia) irianensis
Alexander, from higher altitudes in the Snow Mountains of Netherlands
New Guinea. The coloration of the body and wings, together with theshort vein Sc. readilv separate the two flies.

Fi g '( ) i l i l tui l ttt t (l -i p o l rh t t P s lb i s . ' rp e n t i g e ra A l e x a n d e r: - m. a l e h y p o p y g i u t '
li
I lg
-;
ti t'i l i l t,tt'l tr tl . t l , , t l t h l ' t f ' s l ' -h i u g a i n v i f i r: a e -A l e x a -n d e r; ma l e h y p o p y g i u m
lx
-^
^
Fi
g
:
-1
--9
:_
u
^
2
1
9
ry
!',o
( i i l ttnnl i l
ma
l
e
h
y
p
o
p
y
g
i
u
m.
--. F-i 9 . . ? utl .i l topll ' 1 y 1 1 " ,i1i t ri h c s i . r: A l e x a n d e r;
Fi g
ma l e . h y p o p y g i u m
A l e x a n t l e r;
It l y r tpl l 1v 1t.1 di y lu : o p h o ru
. 5 l , , u o -n o my l a-^^
Fi g . 5 2 . C o n o my i a . (L i p o p h l e p s )n ra l e ' h y-:p o p y-tg"i u
r t ti ',i y tttr i ,'i l s thy ri u ' A l t \ rn (l c r;i cri. r. , 5 3 . A o n o my i a,,, 1l ,,i ,i f .r .r r r r l r r . t t t ; rl r l ry t l (' l )v A i rrrrr.
\(L+ ii .ppoopphl ll ,ec.pp:-s\ .l ?
op. pl lhi oi onn
ff.
A o n o my i a,,r l i t,,r t Atr \,ur r l r | .
rrrrl r l rv 1 i , , 1 l y [ 1 , " t ' . --- f i g .
Fl t { . 5 5 . G o n o my . i a (L i p o p h l e p s )i i i , i t " " i v 1 t , ' 1 ' y , . i rrt rr.
i '| ,,,1,'':r ,,.',,
ni , i " ' ; , i " ' .
l i i g i rl i . O o n rrn r)' i a , . (* i P o P h l (P s ), t ! ! , t ! ! l o l l "tttr tt:t\l tnr
Al l r ,r r rrl rr. rrr. rl r t rv g ' , ' 1 ' v g l ri rn .
rl , t l i s t i s t v l c ; / ) p n f l l l o s o me ;StJ,'lllil|l
Al r r ,r nr l Ir ,r r r ,r l r , l rv l rrr| v g rrt t r,,.,,,i,,11:,t'*tt'"'

Al e x a n d e r, Au s tr al asi anC ranefl i esII

Stud,iaEnt., vo,l. 16, fasc. 1-4, outubro de lg7ll

38. Gonomyia (Idiocera) gorokana sp. n:narrowly brownishblack,_ Mesonotalpraescutumgray, interspaces
includinga centralvitta; rostrum,palpi and antennae
black; femora
yellow,tips narrowlybut conspicuously
black; wings brownishyellow,
very _restrictedly
patternedwith darker; cel.lRr. relativelysmall, Rr_,
a1d.Rr_on costasep_arated
by a distanceneaily equalio the lenjtfi
of the latter; veinsR, and 1sl A withouttrichia; abdomenbrownish
black,posteriorborderiof tergitesnarrowlypale.
Female. Length, about 6.5 mm.; wing, 6 mm. Rostrum
and palpi black. Antennaewith scapeand pedicelblack; flagellum
chiefly black, proximal segmentsslightly paler; basal segments
long-oval, outer ones smaller. Head dark gray.
Pronotum brownish gray. Mesonotalpraescutumgray, interpacesnarrowly brownishblack, includinga centralvitta on anterior
haff, posterior sclerites of notum brownish gray, pleurotergite
and pleura yellow. Halteresdusky, base of stem restrictedlypaler.
Legs with fore coxae dark brown, remaining coxae and alltrochantersyellow; femora yellow, tips narrowly but conspicuously
black, the amount subequal on all legs; tibiae obscure yellow,
tips very narrowly blackened;tarsi with basitarsiobscureyellow,
passing into black. Wings (Fig. 38) brownish yellow, prearcularand costal f ields clearer yellow; stigma pale brown; a vague
darkening in outer end of cell R+; veins pale brown, the cord,
including m-cu, darker brown to brownish black. Veins beyond
generallevel of origin of Rs with trichia, lacking of R3 and lst A,
a few at outer end of 2nd A. Venation:Sc moderatelylong, Sc1
ending about opposite one-sevenththe length of Rs, Sc far
removed; cell R3 relatively small, vein R3 nearly as long as the
distanceon costa betweenit and tip of R1_2icell 2nd M, about
one-half longer than its petiole.
Abdomen brownish black, posterior borders of tergites
narrowly pale.
Habitat: New Guinea.
Holotype, 9, Goroka, NortheastNew Guinea,1550 meters,
June 25, 1955 (J. Linsley Gressitt); Bishop Museum.Paratopotype, I .
Gonomyia(ldiocera) gorokana is quite distinct from the otherpresentlyknown membersof the subgenus,
G. (1.) impavidaAlexander
and G. (1.) punctipennls Edwards, differing especial,ly in the colorationof the body, legs and wings, and in the venation and trichiation of
the veins. In both of these latter species cell R., of the wings is close<Iby the touching of veins R,-, and R. at margin.

39. Gonomyia (Idiocera)( l t,nom ) 'i a ( l di oc er a)2{r : J

impavida Alexanderi mp a v i d a A l e x a n d e r; A n n . Ma g . N a t . H i s t . , (l t )! 4 : 5 5 2 -5 5 i J ;t0{7.

Type from Hollandia, NetherlandsNew Guinea, April 24,
l{f45 (Harry Hoogstraal).Male hypopygium(Fig. a ).

40. Gonomyia (Lipophleps) aegina Alexander( l onom y i s( Li pophl e p s ) a e g i n a A l e x a n d e r; A n n . Ma g . N a t . H i s t . , (l t )1 4 : 5 6 0 -5 6 1 ;1947.

Type trom Hollandia,NetherlandsNew Guinea, January77,l{)45 (Harry Hoogstraal).Male hypopygium (Fig. 45).

41. Gonomyia (Lipophleps) basicuspis Alexander""n"^{;t-r,j*,'o?'nX+:"\

basicuspisAlexander;Ann' Mas' Nat

Hist', (11)Type from Hollandia,NetherlandsNew Guinea,January 18,l{)45 (Harry Hoogstraal).Male hypopygium (Fig. O).

42. Gonomyia (Lipophleps) biserpentigera Alexander
al ononr y i s ,( Li pophtep s )5lilt-564; I9,17.

b i s c rp e n t i g e ra

A l e x a n d e r;Ann.

Ma g .

Nat.

(i l )

Hist.,

Type from Hollandia, NetherlandsNew Guinea, January 18,1045 (Harry Hoogstraal).Male hypopygium (Fig. aZ).

43. Gonomyia (Lipophleps) bougainvilleae Alexander
al t'i l t,nty i d . ( l .i l r ottl tlt l r$ )!l .l ti - ( .1.17;105 { ).

b o u g a i n v i l l e a e . A l e x a n d e r;Ann.

Ma g .

Nat.

(1 2 )

Hist.,

3:'lypc frorrr BougainvilleIsland, SolomonIslands,December:ll, l{)44 (llobcrt Staples).Male hypopygium(Fig. a8).

44. Gonomyla (Lipophleps) citribasis Alexander( l .l po1t1 1 l 1 1 1 x 1r' i l ri 0 c s l s
l l nni i l u'ttt.
1l X:r l r l r . l !1.1 7' l yP r.

A l e rrn t t c r;Ann.

Ma g .

Nat.

Hist.,(l l )ll:l ror r r lr r r r 111l;l';1,r 1lr r ; r( ( 'olk, r . lol.r r r r kr r own) .M ale hypogrypl rrrrrr{ l ' i pi , 1()l ,1StudiaEnt.,vol. 16, fasc. l-4, outubrode 1973

Aus t r a l a s i a n C r a n e l l i e s II

Alex ander ,2 54

50. Gonomyia (Lipophleps) tenuipollex Alexander

45. Gonomyia (Lipophleps) dipterophora A l exander
C o n o myi a ( Lipophleps)
5 6 9- 570; 1947.

dipter ophor aAlexan der ;

Ann.

M ag.

N at. rl i st., (l l )

295

14:

( Li pophl e p s )Oonom y i a
555- 55?;1947.

tenuipollex

A l e x a n d e r;Ann.

Ma g .

Nat.

Hist.,(l I )

l,l:

Holotype, d, Mafulu, PaPua, 4000 feet, December 1933
(Cheesman).Male hypopygium(Fig. 50).

Type from Sairopi,Papua (Collectorunknown).Male hypopygium (Fig. 56).

46. Gonomyia (Lipophleps) ischyria Alexander

51. Trentepohlia(Trentepohlia)abronia sp. n.

Co n o myi a ( Lipophleps\ iscftyr ia Alexander ; Ann. M ag. N at. H i s t., ( l t)

l 4: 557- 558;1947-

Type from Araibo, Papua (Collectorunknown). Male hypopygium (Fig. 51).

47. Gonomyia (Lipophleps) ocypete Alexander
Go n o myia ( Lipophteps) ocypete Alexander ; An n. M ag. N at. H i s t., ( tt)

l 4:570- 572;1947'

Type from Hollandia,NetherlandsNew Guinea,October24,
1944 (Harry Hoogstraal).Male hypopygiLrm(Fig. 52).

48. Gonomyia (Lipophleps) ophion ophion Alexander
A o n o myia

( Lipophteps) ophion

Alexander ; Ann. M ag. N at. H i s t',(ll)l 4:566- 507;1947'

Type from Cyclops Mountains, Netherlands New Guinea,
March 1945 (Jean Laifoon). Male hypopygium (Fig. 53)'

48A. Gonomyia (Lipophleps) ophion brachyglossa Alexander
Go n o myia ( Lipophleps\
l 4:567:1947.

ophionbr achygloss a Al ex ander ;

Ann

M ag

N at'

H i s t'

( !1)

1933
Type from Kokoda,Papua, 1200feet, August-September(Cheesman).Male hypopygium(Fig. 5a).

49. Gonomyia (Lipophleps) recurvispina Alexander( Lipophleps\
Co n o myia
561- 563; 1947.

r ecur vispina

Alex ander ;

Ann'

M ag

N at'

H i s t',( 11)

General coloration oJ thorax Iulvous yellow, abdomen orange, terminal
lwo segments in ma'le black; antennae black throughout; halteres andlegs yellow, in cases outer ends of femora weakly darkened; wings
whitish subhyaline, prearcular and costal fields light yellow, cord, R3,
rtrrter medial fork, Cu and axilla narrowly seamed with brown, wing
npex paler brown.

M a le. Length, about 5.5-6 mm.; wing, 5.5-6 mm.
F e m a I e. Length, about 6-6.5 mm.; wing, 5.8-6 mm.
llostrum and palpi brownish black. Antennaeblack throughout;f fagellar segments elongate. Head dark gray; vertex strongly
carinate.
Cervicalscleritesbrown. Thorax fulvous yellow, unpatterned
cxccptfor a vaguecentraldarkeningat anteriorend of praescutum;
pracscutaland scutalsetaesmall,pale,on anteriorparts; pronotal
bristleselongate.Halteresyellow.Legswith coxaepolishedyellow;
runainderof legs yellow,outer ends of femora in casesslightly
rttoreclarkened;dark brown. Wings (Fig.
terminaltarsalsegmentsl'r7) whitish subhyaline,prearcularand costal fields light yellow;;t restrictedbrown patternthat includesnarrow seamsover cord,/?.1ancl outer medial fork; vein Ca and axilla darkened;wing;t1tcxin ccllsR3 to R5 narrowlypaler brown, cell Rs chieflypale;vt'ins ycllow, clarkerin the patternedareas,clear yellow in the
hriglttcrrctlf iclcls. Veins chiefly glabrous; distal sectionof Rs
witlr ;rllorrteight to ten scatteredtrichia, on elementSc2 - R1
vr'ry sparsc. Vcnation: Rs very gently arcuated,shorter thanlr;ts;rlst'ctiorrof R;,; R2-3-aand R3-a subequal; apical fusion of
vt'irts('rr1 arrd /.s/ A short.
Altrlorrrt'nlight orange,in male terminaltwo segmentsblack,
Ityltopygirrrnntorc intcnscly so. Ovipositorwith cerci slender,
ttturlt'r;rlt,lyrrllcrrrvetl
to thc acrrtctips.

l 4:

Type from Hollandia,NetherlandsNew Guinea, April 17,1945 (Harry Hoogstraal).Male hypopygium (Fig.55)'

I l;rlrit;rl: Nt,u, ( irrilrt','r.
ll,rlr'1y11,',,1,
llorrrl;r,Nt,llrr,r'l;rrrtls
Nt,w (irrint,;r,;rltilrrtle
1500
r r r ,l i l, \ r r p 'r r .i t l .l ,l tl !1(|| r r r sh ,y, ( i r |ssi i l ) ; l l i sl r o g r M r r ser i l tr .

Alex ander ,Aus t r a l a s i a n C r a n e f l i e s l lStudia Ent., vol. 16, Jasc. 1-4, outubro de 1973297

8 d 9, August 12-14,1955.Paratype,d' Obano,Paratopotypes,
NetherlandsNew Guidea,August 9, 1955.
The most similar species is Trentepohlia (Trentepohlia) doddi
Alexander, of Melville Island, North Australia. This differs in colorationof the wings and in slight details of venation, including the straighter
Rs, longer hr-r-n and shorter R4, which is only about twice the lengtho{ vein Rr.

52. Trentepohlia(Trentepohlia)angustilineasp. n'
Mesonotal praescutum light yeltow with a very narrow brownish
black line, scutal lobes yellow, posterior sclerites of notum and dorsal
two-thirds oI pleura dark brown, ventral sternopleurite abruptly yellow;
wings pale yellow, prearcular and costal fields more saturated yellow;
u ."onspi.uous pale brown pattern that includes a solid band at cordwith separate areas at R, and medial fork, th,e wing tip and a broad
seam over vein Cur.

FemaIe.

-Length,about 5.8-6 mm.; wing' 5'8-6'2 mm'

Rostrum testaceousyellow, palpi black' Antennaewith scapedark brown, succeedingsegmentsyellow, outer flagellar segments
passinginto light brownl tlagellar segmentsrelativelylong withshort verticils.Head black, sparselygray pruinose;vertex carinate.
Cervical sclerites and pronotum uniformly brownish black'
Mesonotalpraescutumlight yellow, with a very narrow brownish
black line on anterior half, most distinct in front, narrowed toa point behind, becomingnearly obsoleteat near two-thirds thelength, lateral praescutalborders less evidently darkened; scutal
lobes yellow, central area dusky; scutellum and postnotumdark
brown, margins restrictedly yellowed. Pleura with dorsal twothirds dark brown, ventral sternopleuriteabruptly yellow. Haltereswith stem whitened, knob light yeltow. Legs with coxae andtrochantersyellow; remainderof legs broken. Wings pale yellow'
prearcularand costal fields more saturatedyellow; a conspicuous
brown pattern that includesa solid band at cord, a broken areaat near three-fourthsthe wings length, including broad seams
over vein R3 and outer medial tork; wing apex narrowly darkened,
involving cells R3 to Rr; a conspicuousdarkened seam alongvein Ca extending from base of cell 2nd. A to the cord' Veins
unusuallyglabrous,outer sectionof Rs with about sevenor eight
trichia. Venation:Sc short, Sc1 ending beyond fork of R's,Sc2
near its tip; Rs nearly straight,short, about one-half thc basal
sectionof vein R:,.

(Tr(n t rt ro h l i a \I l g tll tr i l l tl tt,l tl i u
a b ro n i a s p . n . ; v e n a t i o n . Fi g . 5 8 . Tre n t e p o h l i a
l l tti l l .tnhl i ul
i ,r ,r U o l n y l l l f n s i s s p . n . ; v e n a t i o n . Fi g . 5 9 . Tre n t e p o h l i a (Tre n t e i o h l i a )i f r ,/'r .r c t'
fl ; v c i l i rl i o n . -(Tre n t e p o h l i a )Fi g . 6 0 . Tre n t e p o h l i a
l i ma t a
sp. n.;\ Gr ,l fl ,l l f
t i l . ' l ' r(n t t ' p o h l i a (I I o n g o md . ) o c a n t h o p h o ra s p . n . ; v e n a t i o n . l 'i A
I r B f',f l r r ul tt,r tl tl it t t A I t )t rt ! t )n t u l l i n t h ri t o s t a s p . n . ; v e n a t i o n . Fi g . 6 3 . Tre n t e p o h l i at,l l ',,f !f ,r r 'r I /r r n1i . , ' r, , . , , A l c \ i rn d e r;
venation. Fi g . 6 . 1 . Tre n t e p o h l i a (Mo n g o ma )l ri g . t i 5 . Tr? n t rp o h l i a (Mo n g o ma ) p a rv i c e l l u l a s p . n . ;I'r r rl t l r c i f r r , r' err; rl i o n .
l i l i . ' l t ri l l (t ] o h l i u
i c r fnl for fl 'l gp a rv i s t i g ma
sp. n.; venation. \ l l l o n g o ma )I tg tt;
I ttnl r f'hl ti
sp. n.; venation.
l l l t )n { )n t a l
Fi g . 6 8 . Tre n t e p o h l i al )(r(rr. l a*t ]t1,r 119,,111111
rr ; v c n l t i o n .
Fi g . t i 9 . Trc n t c p o h l i a (Mo n g o ma )v i t ri n a
I,r r r r tl l u s l ts t ). n . : v c n a l l 0 n .

298

Aus t r a l a s i a n C r a n e f l i e s l lAlex ander ,Sturiia Ent., vol. 16, fasc. l-'1, outubro de 1973

Abdomcn dark brown; ovipositoryellowishhoru color.
Holotype,?, Nondugl,NortheastNew Guinea,July 8, 1955
(J. Linsley Gressitt); Bishop Museum.Paratopotype,I .
Trentepohlia (Trentepohlia) angLtstilineahas the wing pattern muchas in ?. (7.) Iepida sp. n., but the coloration of the thorax in thetwo f lies is entirely cliff erent.

53. Trentepohlia (Trentepohlia) bougainvillensis sp. n.

Holotype, 9 , EmpressAugusta Bay, Bougainvillelslands,;tlrrngKoroniokinaRiver, Septentber19, 1944 (Robert Staples)clingingto washedout roots of banyantree.Allotype, I , Gizo
( iizo, altitude 0-50 rneters,November 10, 1970 (Noel Krauss .
)l':rratypes,l1 d 9, with the allotype.
The most similar species is Trentepohlia (Trentepohlia) pictipennis
llczzi, of th,e Philippines. This differs in cietails of coloration of the
Irolly 6nd wings, including very evidently the entirely clarkened vein R^
rvhich in the present fly in the outer ground crossband is light yellow. In
the Oriental region there are several other members of the subgenushaving
tlte wings attractively patterned, all differing from the present fly.

Head black; mesonotal praescutum light brown, posterior sclerites
d arker br own, pleur a blac k ened; ha l t e r e s a n d l e g s l i g h , t y e l l o w ; w i n g slight yellow with two major complete brown crossbands, one including
the broacl apex, the other chiefly at ancl basacl of cord, extencleciovervein Cu as a broad seam; veins in the outer ground crossband entirely
lig ht y ellow.
M a I e.

Lengt h,F e nr a I e.

about

Lc ngt h,

5. 2- 5 . 5

about

nlm.;

6-6.5

wil1g, 5-5.5

mnr.;

wit.tg,liut.l.
5-6.5

ntnl.

Ros tru n r s h o rt, b l a c k ; l a b i a l p a l pi el ongate, hotn yel l ow ; ntaxi l l ary
palpi b l a c k . A n te n n a e b l a c k . I-l ead pol i shed bl ack.
C e rv i c a l re g i o u b l a c k . P ro n otum brow ni sh bl ack. Mesonotal
pr acs c u tu m a l n ro s t u n i fo rml y l i g h t brow n, posteri or scl eri tesdarker
br or , v n . P l e u ra a n d s te rn u m chi ci l y bl ackened, very sparsel y
pr uin o s e . Il a l te re s l i g h t y e l l o rv . Legs w i th fore and tni ddl e coxae
blac k , p o s te ri o r p a i r y c l l o w ; f ore trochatrters brow ni sh bl ack,r em a i n i n g tro c h a n te rsy e l l o w ; rel nai nder of l egs cl ear l i ght yel l ow ,t c r nr i n a l ta rs a l s e g n re n t b ro w n . W i ngs ( Fi g. 58 ) l i ght ycl l orv
r , v it htl o l rra j o r c o u rp l e tcb ro w u bands that cross the w i ttgs, pal cr
on p o s te ri o r h a l v e s ; b a s a l b a n d at and near cord, extendi ttg frottt
l sl A , sol i dl y darkencd i n cel lc os t a to a p c x o f v e i n s C u , R1, p o s te ri o rl y b e h i n d th e ra d i al f i el d {ormi ng darkenecl seanl s
oll c o rd a n d o v e r v e i n s M a n d C u cxtended basal al ntost t()
ar c ul u s ; o u te r b a n d i n c l u d i n g the broad w i ng apex, i ts i nneredge s tra i g h t, a t l e v c l o f i n n er ends of cel l s R 13and R 5, theint erv e n i n g p a rts o f l v i n g c l e a r l i ght yel l ow , the band separati ng
t he t w o d a rk e n e d a re a s c l e a r yel l ow , i ncl udi ng the vei ns; vei ns
in da rk e n c d a re a s b ro w n , e l s e w here yel l ow . V ery sparse tri chi aon d i s ta l s e c ti o n o f v e i n R 5 . Venati on: R s and basal secti on ofv ein R 5 s u b e q u a l ; R s p l u s M1 -2 l onger than basal secti on oi R r.
A b d o m e n d a rk b ro w n , m o re bl ackened l ateral l y; val vcs ot
ov ipo s i to r h o rn y e l l o w .
F l a b i ta t: So l o n ro n Is l a n d s .

299

54. Trentepohlia (Trentepohlia) fuscomedia sp. n.
( i e n c r a l c o l o r a ti o n o f th o r a x l i ve r b r o w n ; h e a d d a r k g r a y; h a l te r e s; r r r t l l e g s y e l l o w ; w i n g s co n sp i cu o u sl y p a tte r n e d w i th p a l e b r o w n a n dr t , l l o w , t h e f o r m e r i n cl u d i n g a b r o a d l o n g i tu d i n a t a r e a l i n ce n tr a l th i r d .

N 1a l c. Lcngt h, about 5 nt nr . ; wing, 4. 6 m nr . Rost r um andp;rl l ri dark br own. Ant ennae wit h scape and pedicel br own,l l .;rscl l Lrrn
pal e r br own; f lageliar segnr ent slong, subequal t o t heirl rr1l oqs1vcrti c ils, t cr nr inal segm cnt elongat e. Head dar k gr ayl
vesl i turc of ver t ex light yellow.
' l ' l torax a lnr ost unifor r nly liver
br own; vest it ur e spar se, pale
yt' l l rrrv. H al teres pale yellow. Legs wit h coxae br ownish yellow,
l rrl 1' pai r darker ; t r ochant er s yellow; r em aiuder of legs yellow,
It' rrror;l lti ps nar r owly and vaguely dar kened, out er t ar sal segm eut stttl rrse;rtcrlW. ir r gs dist inct ively pat t er ned; pale yellow wit h anl xl crrsi vc browu longit udinal seant in m edial f ield ext ending f r onr
l r;rst' to trpcx oI wing, including nr or c t han one- t hir d t he widt h,rr rl l r ;rrr cxl cnsior r t o cost a over t he ant er ior cor d and a f ur t her,,rrt' ;rl nrr)st l () nlar llin at vein Rai a post er ior seam along vein
( r/ l ,r nri ugi rt ; t lr c pale yellow gr ound ar eas on cost al t hir d include
l l rl l rrol rrl l rl rst ',r vit lr 1wo snr aller br ight enings bef or e and beyoncl/,' 1; posl tri or. llr ir t l of wing less conspicuouslyvar iegat ed by pale;
l r' rr\ v(' ry l i t 1lt l llr own, r r r or e yellowed in cost al f ield. Spar serr,r(ri )l rrtl ri ;r r r r r t list ; r l scct ion of vein Rs. Venat ion: Rs st r aight ,,rl r.nt l ,rrr l i l llt s llr t . lxr sal scct ion <t f Rsi vein R3 oblique, about,' rrr rl rrr,l /r' 1, / i, ' , ; 1 sliglr lly lor r gcr t han Rs_+i m - cr close t ol ,' rl r,,l .11,;r Pr r ; r l lr r sior rof ( 'r r , ant l / . sl A shor t .
\1,,1,,111,, ypopygir r lr rslight ly dar ker .
1 1r r r r r l, 'r r r r lIr l; r r li I r r . , r vr r lrll

r l 'r tr l

" , r\ \(ri l 1 i l { , 1IlCraneflies
Alexander, AustralasianStudiaEnt.,vol. 16, fasc. 1-4,outubrode 1973

Holotype, qr Obano, Wisselmerenarea' Netherlands New'
Guinea,August 9, 1955 (J. LinsleyGressitt); Bishop Museum'

Abdominal tergites black, sternites slightly paler, genital
mgrttentsbrown. Ovipositor with cerci horn-yellow,broad-based,
outcr half very slender,upcurved to the acute tip.
Habitat: New Guinea.
Holotype,I , Nondugl,NortheastNew Guinea,July 8, 1955
(J, Linsley Gressitt); Bishop Museum.Paratopotypes,3 9 I .
l)aratype,1 9, Korip, July 12, 1955.

300

The distinctive wing pattern of the present fly readily. separates
it 1rom other regional mimbers of the typical subgenus. The liver -browncoloration oi tlie thorax is much as in the otherwise quite distinct
Trentepohlia (Trentepohlia) pictipennis Bezzi, ol Mindanao, eastward to
New Cuinea and satellite islands.

301

This attractivelly is generallysimilarto speciessuchas Trentepohlia
55. Trentepohlia (Trentepohlia) lepida sp. n.
General coloration of thorax dark brown, abdomen darker; halteresyellow; legs white; wings pale yellow, extensively patterned with yellow,
including t'ransverse bands at cord, at near three-fourth the wings length,-wingtip, with a further seam along vein Cu,' Rs short, slightly
and atarcuated.

Female. Length, about 5-5.3 mm.; wing, 4.8-5.2 mm.
Rostrumand palpi black. Antennaewith scapedark brown, pedicel
and proximal tlagellar segmentspaler, more testaceousyellow,
outer flagellar segmentsdarkened; flagellar segmentslong-oval,
exceedingtheir verticils.Head dark brown; anteriorvertexnarrow,posteriorvertex carinate.
Pronotumand cervicalregion dark brown. Mesonotumalmost
uniformly dark brown, humeral region of praescutum slightly
paler; sparse setae on praescutum,longer on pronotum andHaltereswith stem
posteriorborderof the triangularpteropleurite.
whitened, knob pale yellow. Legs with coxae light yellow orslightly darkened,especiallythe anteriorpair; trochanterslightyellow; remainderof legswhite.Wings (Fig. 59) with the groundpale yellow, more saturated along costal border; a conspicuous
darkenedpattern that includestransversebands at cord, a broken
area at near three-fourthsthe wing length, more intense over
the radial fork, and the narrow wing apex, the latter two areasin cell Rs; a further longitudinaldarkened
narrowly interconnected
streak chiefly along vein Cu, in cell M extending from bases ofAnal cells outwardly, joining the band at cord; darkeningin cell
R1 almost filling basal two-thirds of the cell; veins yellow, darker
yellow in the more saturated fields, infuscatedin the darkened
areas. Veins with very sparse trichia, including about eight toten on outer section of vein R5. Venation: Sc2 subobsoleteor
faintly indicated,a short distancefrom tip of Sc1,'Rs short,slightly
arcuated;R1-2 and R2 very faint; apical fusion of veins Cu'
and lsl A short.

('l'rcntepohlia) pictipennis Bezzi, differing in the coloration of the antennae
nrrd thoracic dorsum and in the pattern and venation of the wings.

56. Trentepohlia(Trentepohlia) limata sp. n.
Ceneral coloration oJ thorax fulvous yellow, unpatterned; head light
grly; halteres and legs yellow; wings whitish yellow, costal borderclcarer yellow, virtually without pattern; abdomen brownish yellow, tergites
with a narrow darker central line.

F e m a I e. Length, about 5.8-6 mm.; wing, 5.6-6.2 mm.
Rostrum pale yellow, palpi testaceousyellow, apex of terminal
scgrnentdarker. Antennaeyellow basally,outer tlagellar segments
palc brown. Head light gray.
Cervical region, prothorax and mesonotumfulvous yellow,
pleura slightly clearer yellow. Halteres pale yellow.
runpatterned,
l.egs pale yellow, outer tarsal segmentsweakly darkened.Wings(l;ig. 60) pale whitish yellow, costal border very light yellow;
v('ry narrow to scarcelyapparent darkenedseamsover cord, Cu
itnd /?;1,best indicatedby the slightly darker veins which elsewhereflr(' pale yellow. A few scattered trichia over the whole lengthof tlistalscctionof vein R5. Venation:Rs oblique,nearlystraight,
rrlltively long, about two-thirds the basal sectionof vein R5;
apical fusion of Cu1 and /s/ 'A in casesrelativelylong, to aboutrlllr'-l{)rrrtll
thc length oI m-cu.
r^rlrtlorttinal
tergitesbrownish yellow with a narrow darker
hlown central Iine, in cases poorly indicated;sternitesclearer
yt'lhrw;srrllternrinal
segmentsslightly darker, especiallyin male;gerritalshicld yellow; cerci less stronglyupcurvedthan
irt ft'rrr;rle
Itt v;rriorrs;rllicrl spccies.
ll;rlrit,;rt: Ncw (iuinca.
,,,, ltouda, Wisse|nrcrenarea, NetherlandsNew
Iloloty'11t,,( lrrtrrt',;r,1,5(X)rrrclt'rs,Attgust 14, 1955 (J. Linsley Gressitt);
several,Allrtopolylx',(t, witll typc.[)aratopotypes,
lllilr,rI Alrrslurrr,,,' ,,,, ( ) l r ;tr to , Wi sst'l r ttt'r e t1 a r ca , Au g u stI t r r r ! r l l t l r r r r i t l r , r . I'l r r l r l - r y,;r t,s,

ru

3 02

Alex ander ,Aus t r a l a s i a n C r a n e f l i e s I i

9, 1955; Okaitadi, 1,760 meters,August 5, 1955 (Gressitt); I
g , Baliam River,NetherlandsNerv Guinea,Archbold Expedition
III, Station9, 1,600nteters,December18, 1938 (Toxopeus).
The present fly is most readily distinguished from other regional
members of the subgenus by the clear yellow thorax and virtuallyunpatterned wings.

57. Trentepohlia (Trentepohlia) luteicosta sp. n.
Thorax fulvous; rostrum yellow; antennaewith scape brownish yellow,
remainder black, flagellar segments elongate; halteres and legs yellow;
wings whitish subhyaline, costal border light yellow, including the veins;a restricted brown pattern that does not attain the costal margin, cellR" c hief ly pale; v ein Rs s t r aight , s u b e q u a l t t - r b a s a l s e c t i o n o f R " ; c e l l
R" fr om about t wo and one- half t o n e a r l y t h r e e t i m e s i t s p e t i o l e .

Female. Length,about 6 mm.; wing,6 mm. Rostrumandmouthparts light yellow; palpi brownish black. Antennaewith
scapebrownishyellow,tip narrowlydarkened,remainderof organ
black; flagellarsegmentselongate,exceedingthe verticils.Headgray; vertex carinate.
Cervicalregion and pronotun'lweakly darkened.Mesonotum
light f ulvous, unpatterned, posterior sclerites darker. Pleura
brownishfulvous.Halteresyellow.Legswith coxaeand trochanters
yellow; remainderof legs yellow,outer tarsalsegmentsinfuscated.
Wings whitish subhyaline,costal border light yellow, including
tlre veins,involvingbasal halt of Rs, outer two-thirds of Rr-"-*and all of R r-+i all posterior veins brownI a restrictedbut
conspicuous
brown patternthat includesnarrow seamsover cord,
Cu, and especiallyvein Ra, this restrictingthe pale centralareaof cell R3 to the outer two-thirds of cell; paler brown washes
in outer parts of cells Ra and M2 and virtually all of Rs. Distal
sectionof vein R5 with about six scatteredtrichia.Venation;Rs
straight, subequalto basal sectionof Rsi cell R5 from about
two and one-half to nearly three times its petiole; vein 2nd A
arched.
Abdomenbrown, posteriorbordersof tergitesslightly darker,
subternrinalsegmentsglackened.Ovipositor with cerci slender,
especiallyon outer halves,upcurvedto the acute tips.
Habitat: New Guinea.
Flolotype,9 , Goroka,NortheastNew Guinea,June 25, 1955
(J. Linsley Gressitt). Paratopotype,?.

Studia En't., vol. 16, fasc. 1-4, outubro de 1973

303

The present fty is most nearly allied to Trentepohlia, (Trentepohlia)
uhronio sp. n. and T. (7.) doddi Alexancler,differing therefrom chiefly
lrr details of coloration of the wings'

58. Trentepohlia (Mongoma) acanthophora sp' n'Size large (wing of male to 9 mm.) ; general coloration of thorax
lrr,rwn;-treaO-frowniitr black; pleura with a compact tuft oI blackened
xctae on ventral pteropleurite; legs brown, feniora with spinoid setae
nt tase, posterior iiUiae ."itt.t two long subterminal spinoid setae; wings
outwcakly'darkened, costal border narrowly more infuscated, continuedA;
warOfy to apex, two elongate setae on prearcular section oI vein 2nd
radialouter
beforevein
extent,R,incells R" una' R. at margii subequalf ork, l'eaving f short element Rr-n.

Male. Length, about 7-8 mm.; rving,8-9 mm' Rostrum
black, labial palpi light brorvn, paler outwardly,maxillary palpi
brownishblack.Antennaeblack, relativelylong, if bent backward
cxtcnding about to wing root. Head brownish black; posterior
vcrtex with strong black setae.
Pronotal scutum pale brown, nlore yellowed laterally,with
frcct black setae;pretergitesyellow. Mesonotalpraescutumlightbrown, with three darker brown stripes, the intermediatepair
regionllalcr bchind,endingsomedistancebeforesuture'humeralrcstrictedlyyellowed;scutallobes brown, median area and outer
postcriurangles of lobes yellowed;posteriorscleritesof notum
tl:rrkcr brown medially,more yellowedon sides. Pleura chietly
more yellowed;ventral pteropleurite
llrown, clorsalsternopleurite
with a cotltpacttuft of about a dozenblack setae.Halteresbrown,lllrsr,of stcnr yellow. Legs with coxae and trochantersyellow;
of legs medium brown, ollter tarsal segmentspalingrlrrr:rirrclcr
femoraat basewith irom six to ten spinoidsetae,
yt,lhrw;
frlrc
lrr
willr ;rlrottt the sante number of sntaller ones; middle f emora
sirrril;rrlrtrt thc larger spines,from about 10 to l2 in number;
Itirrtl l(.rrror;lwith thc spines smaller; posteriortibiae with twosetae,with an additionalsmallerbristle.wings
Lilrg srrlrlt,rrrrinal( liig. lil ) wcaklyclarkened,costalbordernarrowlymoreinfuscated,darker brown, appearingas astignaapcx;
r'orrlirrrrctl
lo thevi.r'y1,iI1()wsgirprt6 vcin R; veins brown. PrearCularsection,rt vlirr Jrnl A witlt lwtt cftrngateblack setae.Venation:R2 before
rrtrllr ltrli;tl folk, lt'avittga shrlrtvein Rg-+ivein R3 long, oblique;t r'llr /,'1 ;tttrl/|r :tt rtt;trgittsttllctlttalin extent; basal sectionot, l l , . l r t t ; t l 1 r l , ;tl r o ttl r r ttt'- l t;tl l l 1 l l l g cr th a tt n t- ctl ,' a p i ca l fu si o n
I r r , , r r i r l / r / ,'l ;tl r ,r tl l l tvo l l ti ttl s tl t- r 'tt.

of

304

II
Craneflies
Alexander, Australasian

Proximal abdominal tergites light brown, darker laterally,remaining tergites dark brown; sternites yellowed, posteriorborders of subterminalsegmentsnarrowly more darkened.
Habitat: SolomonIslands.
Holotype, 61, Empress Augusta Buy, Bougainville Island,along Koromokina River, altitude 50 feet, September28, 1944
(Robert Staples) - Along sides of dugout in jungle - Staples.
2 dd, mountedwith type; one d, September
Paratopotypes,
13, 1944. Paratypes,.;:6:, Hill 700, altitude 650 feet, December
6 and 31, 1944 - In thick jungle, under logs and banks alongcascadingclear rivulet - Staples.
There are several other Australasianspeciesthat are generally
similar to the present fty, all differing among themselves in size, detailsof venation and in the armature of the legs, such species including
Trentepohlia (Mongoma) brunnea Edwards, f . (M.) guamensis (Alexander),T. (M.\ ibelensis Alexander, f. @.) macrotrichiata Alexander, and T.
(M.) spinaspersa Alexander. It should be noted that macrotrichiata and,some other species have from one to three modified setae at the proximal
end of the basitarsus of the hind leg instead oI on the tibia as in
the present fly.

StudiaEnt., vol. 16, fasc. 1-4, outubrode 1973

305

Iottr short black spinoid setae; basitarsi at proximal end with
tltrcc rnodifiedsetae,the largest black, the others yellow. Wings(Fig.62) weakly suffused,cells C and Sc darker brown; stigma
narrow,mediumbrown; veins brown. Costal fringe (male) long;rlutcr section of vein Rs with scatteredtrichia. Venation: R2
alrout twice R3-af vein R3 oblique; cell 1sl M2 longer than vein
Ma; basal sectionof M" arcuatedlm-cu shortly before lork of M;
apicalfusion of Ca1 and 1sl ,4 relativelylong, about one-halfm-cu'
Abdominal tergites, including hypopygium, dark brown,
nlcrnitesobscureyellow.
Habitat: New Guinea.
Holotype,6l,Korip, NortheastNew Guinea,July 12, 1955
(J. Linsley Gressitt); Bishop Museum'
The most similar regional species include Trentepohlia (Mongoma)costofimbriata Alexander 1nd T. (M.) tongisetosa Alexancter, which havefltc riostal fringe of wing of male even longer and with -cell lst M,
rhort. Both of these species have the spinoid setae of the femora more
nlrmerous, eight or more, in costolemorata more elongate than in theprcscnt fly.

60. Trentepohlia (Mongoma) longisetosa Alexander
59. Trentepohlia (Mongoma) fimbricosta sp. n.
General coloration of thorax liver brown, pleura more yellowish
brown; antennae black throughout; Iemora dark brown, posterior pairwith four short black spinoid setae, remainder of legs brown, outertarsal segments paling to yellow; wings weakly suffused, costal regionand stigma darker brown; costal fringe of male long; vein R, about
twice Rr-n; cell 1st M, longer than vein M.n; abdominal tergites andhypopygiuin dark brown, sternites obscure yellow'

M a le. Length,about 7 mm.; wing, 6.3 mm' Rostrumdark
brown, labial palpi brownishyellow,maxillarypalpi black.Antennae
black throughout; flagellar segmentselongate-oval,longer outwardly; segmentswith a short densewhite pubescence'verticils
short. Head blackened; posterior vertex carinate'
Cervical region and pronotum brownish black, anterior
pretergitesobscureyellow. Mesonotumliver brown, scutellumand
central region of scutum more testaceous; cephalic part ot
praescutumwith a few relatively long erect to slightly retrorse
black setae. Pleura and pleurotergiteyellowish brown' Halteresdark brown, base of stem restrictedlybrightened.Legs with coxac
yellow, fore pair slightly darker, trochantersyellow; femora darkbrown, basal fourth paler; tibiae and basitarsi mediunt brown,
outer tarsal segmentspaling to yellow; posteriorfent<jrawitlt

T 'r ti l l tD ohl i a ( M on e o ma \ l o n l i s ? t o s a A l e x a n d e r;i l t, l 2n- tz ?," f i g .
0 (v e n a t i o n ); 1 9 3 6 .

P ro c . L i n n e a n S o c

New South Wales'

The types were from Maini and Kone, Papua, taken by K'
J. Clirrton.
A further specimenfrom Daulo Pass, NortheastNew Guinea,
24(X)rnetcrs,collectedby Gressitt.Venation (Fig. 63).

61. Trentepohlia (Mongoma) parvella sp' n.
sirc snrall (wing to 5 mm.); general coloration of thorax liver
Irr'wrr; legs 5rrwn, iarsi and tips oi tibiae whitened; posterior femora
ill rrrllt, willt ilbout five spinoid setae; wings slightly dusky, apex more
16frrrir'trletl
i /?, :rbetrt,,ne and one-half to two times R3-0, cell /sf M, short'

M a le . l.t'ngth,allout 4.5-5 mm.; wing, 4-5 mm.
I't' rttI I r'. l.ength, about 5.5-6 mm.; wing, 4.8-5 mm'f{orlrrrrrrlrrowrrto brownishblack; palpi black. Antennaeblack;the outer ones longer; verticils
eIrrrgatc-oval,llagcll;rrs(.1{1t(,llls
llcad lllack; posteriorvertex carinate.
rhullt,r llt;rrrtltt, st'gtttt'ttts.
('t,rvtr,;rlrr.giorrllrowrrisI Illack. [)rrlth6raxand nresothorax
with long
ligltl lrrnwrtlo livt'r l)rowll; prrlnrltttttt
6l111nef
rrllfrrrrrrly

StudiaEnt.,vol. 16, fasc' 1-4,outubrode 1973

erect setae,mesonotumwith setaeshorter,sparsebut conspicuous.
Halteres dusky, knob a little darker. Legs with coxae andtrochantersliver brown; femora dark brown; tibiae brown, thetips and tarsi paler, more whitened; posterior femora at basewith about spinoid setae; in female posterior basitarsuswith astrong black seta at base. Wings (Fig. 64) with a weak dusky
tinge,stigma and costalregionslightlydarker,wing apex weakly
more infuscated;veins light brown. Costal fringe relativelyshort;
distal sectionof vein R5 with scatteredmacrotrichiathroughout
the length. Venation: R2 about one and one-half to two times
Rg-ai cell lst M, short, not exceedingvein Ma; apical fusion ofveins Ca1 and 1sl A about one-fourthm-cu.
Abdomen dark brown, basal sternitesmore brownish yellow.
Ovipositor rvith cerci slender, gently upcurved.

in holotype'the paratype specimenhas cell lst M2 shorter than(cephalic
wfth veinsM1-2 and the combinedelementR+-s * M1-2
vein |onger
brrrc|erot cell l st M) Subequalin length, the lattervein M1-2'
twice
than
more
In the holotype,,, .ho,tn, slightly
AbdomenuniformlY light Yellow'

Habitat: New Guinea.
Holotype, 6t, Korip, NortheastNew Guinea,July 12, lgsb
(J. Linsley Cressitt); Bishop Museum.Allotopotype,9 , Tsenga,
July 14, 1955. Paratopotypes,2 broken 6 d, paratypes,2 6t 9 ,Busu, September15, 1955.
The species is most similar to Trentepohlia (Mongoma) brevipes
Alexander, which differs in details of coloration, especially of the wings,
as the conspicuously yellowed costal border, including the veins, themore numerous trichia on vein R^, and commonly with few to several
trichia on Rr-n and adjoining velns.

62. Trentepohlia (Mongoma) parvicellula sp. n.
Size small (wing to about 4.5 mm.); general coloration of body,legs and halteres light yellow; antennal ilagellum pale brown; wings
with vein R, shortly be{ore the outer radial fork; cell 1sl Mo small,the inner eniis of cells Ru and M" virtually in alignment.

.il

307

Alexander, AustralasianII
Craneflies306

M a le. Length, about 5-5.5 mm.; wing, 4-5 mm. Rostrum
light yellow, palpi slightly darker. Antennae with scape andpedicelyellow,flagellumpale brown; flagellarsegmentslong-oval,slightly exceedingtheir verticils.Head light brown, anteriorvertex
very narrow.
Thorax, togetherwith the halteresand legs uniformly lightyellow. Legs with abundantshort black setae,nonemodified.Wings(Fig.65) yellow,veinspale brown, thosein prearcularand costal
fieldsmoreyellowed.Venation:Vein R2 shortlybeforeouterradialfork, leavinga short elementR3-aicell Ist M2 small,rectangular,
innerendsof cellsR5 and M3 slightlyretracted,
nearlyin alignrrrt'nr;

Habitat: Fiji.
Holotype, 6, Lami, Viti Levu, 100-350 meters,March l,
l97t (Noel Krauss). Paratype,a broken i, Tholo-i-cuva,Viti
Levu, July 27, 1938 (Elwood Zimmerman)'
Other regional uniformly yellow members of the subgenus oI small
and T'
rlze include Trentepohlia iMongoma) poliocephala Alexander
themselves by theamongsepirated
readily
mosf
all
n.,
sp.
vitrinalM.)
vtnational characters as described.

63. Trentepohlia(Mongoma)parvistigmasp' n'(ieneral coloration of thorax yellow, the mesonotumweakly patterned,
hcnd light yellow; wings subhyaline, prearcular and costal fields lightycllrrw,'.tigti.tu very small, dark brown; vein R, shorter lhan Rr-n, m-cu
nl fork of M.

Male. Length,about 5 mm'; wing' 5.5 mnl' Rostrumand
palpi light yellow. Antennae with scape yellow, remainder dark
br0wn; flagellar segmentssubcylindricalto long-oval,exceedingtlteir verticils. Head light yellow, the narrow anterior vertex andposteriororbits light gray; posteriorvertex carinate'
Cervical region and pronotum light yellow' Mesonotal
prfl(.scutunland scutal lobes fulvous, the remainder of notum
palt'r ycl6w, lined longitudinallywith pale brown' Pleuri andand scutallobes
light yellow; vestitureof praescutum;rlrrrrotergitc
coxae andwithLegs
vr.ry sltort ancl sparse. Halteres yellow.
lroi'ltattlersycllow; remainderof legs broken.Wings (Fig' 66)
prcarcularand costalfieldslight yellow;stigmadistinct
xrrlrhy;rline,
Itttl vt'ry stttall,clark brown; veins pale brown, more brownish
yclLrw in thc brightenedlields. Distal section of vein Ro withelrorrltt,rrsrrrallscatteredtrichia.Venation:R2 shorterthan Rs-a
errtl srrltt.tlralto basal sectionof Rri cell 1sl M2 subequalto
tffnlalst'cliottol M;;; nt-cu at fork of M; apical {usion of Cu1
nttrl /.sf I sltorl; cell 2nd A relativelynarrow.
tergitt'svaguclybicolored,yellow, the lateral and
AlrtLrrrrirr;rl
hypo;trulr'riorlrorrlersntrr0wly rrroreinfuscated;sternitesand;tyglttrrreltitr('r Yt'lLrw.
llolrllnl: Nlr\' (ittittt'lt,

308

Alexander, Australasian
Craneflies
It

StudiaEnt., vol. 16, fasc. 1-4, outubrode 1973

Holotype, 6f, Kabebe, NortheastNew Guinea, 2100 meters,
June 21, 1955 (J. Linsley Gressitt); Bishop Museum.

(Robert Staples)- in jungle beneathlogs on spider webs Staples.Allotopotype,9, pinnedwith type. Paratopotypes,
5 6 9,
on three pins, September28 - October 11, 1944. Paratypes,2
d 6, 2 I 9 , EmpressAugusta Bay, APO 716, June 6, lg44
(Ashley B. Gurney).

-

The species is most simirar to Trentepohlia (Mongoma) poliocephala
Alexander, widespread in the oriental and Australasiai tugionr, differingin the coloration of the body, especially the head, and "in the patternand venation of the winss.

309

The presentfly is readirlytold from other regionalspeciesby the

64. Trentepohlia(Mongoma)pererectasp. n.
Siz'emedium(wing about 6 mm.); generalcolorationof thorax
fulvous ; antennaeof male relativelylong, about one-thirdthe bodyor wing; knobs of halteresdarkened;tegl tigtrt brown, outer tarsals_egments
brownishyellow;wingswhitened,aper weaklyinfuscated;vein
erect,vein R* unusuallyshort, obliqu-e;ceil Ro small,lJz-s..preserved,
tne drstanceon cosfaaboutone-sixth
to one_eighth,that of cell?r.
M a le. Length, about 5.8-6 mm.; wing, 5.5-6.3 mm.;
antenna, about 2 mm. or more.
F e m a I e. Length, about 6.5 mm.; wing, about 6 nim.
Rostrum brownish black, labial palpi paler brown, maxillary
palpi black. Antennae black, unusually long, as shown by themeasurements;flagellar segmentssubcylindrical,much exceeding
the veiticils or the shorter pale erect setulae. Head brownish
black.
Cervicalregion brownish black. Mesothoraxalmost uniformly
fulvous, vaguely patternedwith yellow on praescutalhumeri ancl
pleura. Halteres yellow, clearer at base, knob light brown tobrownish black. Legs with coxaefulvous; trochanterslight yellow;
femora light brown, bases narrowly more yellowed; tibiae lightbrown, tarsi outwardly paling to brownish yellow. Wings (Fig.
67) whitish, stigma small and narrow, inconspicuous,
pliced far
cephalad;wing apex in cells Rs, Ra and R5 weakly infuscated;veins browfl, Rr-z relatively faint. Sparse scatteredtrichia over
the whole length of distal section of Rs. Venation: Vein R2-3
preserved, erect, about one-third to one-fourth Rzi vein Re
unusually short, oblique, cell R2 ver! small, its length on costa
onfy about one-sixth to one-eighththat of cell Rsi m-cu shortly
before fork of M.
Abdominal tergites dark brown, basal sternites brownish
yellow, outer segmentsslightly darker.
Habitat:, Solomon Islands.
Holotype, 61, Empress Augusta Bay, BougainvilleIslancl,
along Koromokina River, altitude 45 feet, October I l. l\t44clongate antennae in the male and in the venation of the radial fieldof the wing, with vein R2_e preserved as a distinct element, cell Ro
treing small, its length on co-stabeing only about one-sixth to one-eightfi
fhat of cell Re. Other somewhatsimilar regional speciesincludeTrentepohlia (Mongoma) brunnea Edwards and T. (M.) guamensis(Alexander)
which have vein R._o presentand cell R, at margin nearly as extensivear cell Rn. f . @:)- brevicellulaAlexandtr has cel,l R, more as in theprcsentfly but with vein R,-u present.

65. Trentepohlia(Mongoma) pumilla sp. n.
Size small (wing about 5 mm.) ; general coloration of thoracic notum
llght brown; legs brown, tarsi light yellow; wings very weakly darkened,vein R. oblique, cell R, on costa about two-thirds cell R3, cell 2nd A
llnrrow.

M a le. Length,about4.5-4.7 mm.; wing, 5-6 mm.; antenna,nltorrt1.5-1.8mm. 'Rostrumunusuallyreduced,brown, labial palpi
lttediuntbrown, maxillary palpi darker brown. Antennaerelatively
kltg, lnore than one-fourth the wing, dark brown; flagellar
rt'grrrcntslong-cylindrical,much exceeding their verticils. Head
Itrownish black; anterior vertex very narrow.
Ccrvical region and pronotum light brown. Thoracic dorsum&lrrrosluniformly light brown, without pattern; pleura yellow.
llaltt'rt,swith stem obscureyellow, knob brown. Legs with coxae
attrl trochantcrslight yellow; femoraand tibiae pale brown; tarsi
llghl yt,lkrw,slightlydarkerbasally.Wings (Fig. 68) very weakly
tlttrkt'rtr,rl,
stignralackingor virtuallyso; veinsbrown.Veinsbehindcosllrvirlrrallyglabrous,with very sparsesmall trichia on R andlls ;rrrrlllrorrt six on distal sectionof vein R5. Venation: Sc,
r.'ntfirtg ol)posilcjunction of veins R1 and Rzi Rz_+slightly morelhan lwo-llrirdsRe; R1-2 subequalto or somewhatshorter thanolllirprc,about one-halfR+; cell Rz on costa aboutlll; vt'in /?11
fwrr-flrirrlslo llrrer'-fourths
Rq; cell 1s/ M, relativelysmall; inner
cttrf rrf ct,ll M;1 slightly basaclof cell 2nd M2; m-cu close toof M; cr,ll Jt t d A ll; lr r ow.
A l rrftrrtri rt lr l lt , r git t , s l) r own, stcrnitcs ycllow; outer fourl tsgl nr,nl q,i rrt'lr r r lir r ghypr r pygir r r r r ,Itl;tcke
rrctl.
l l nl ttt* tt S, r l, r nt , r t tlsl; r r r r ls,
l rtl

3 10

Alex ander ,Studia Ent., vo'[. 16, fasc. 1-4, outubro de 1973

Aus t r a l a s i a n C r a n e f l i e s I I

311

Holotype, 6, Empress Augusta Bay, Bougainville.lsland,
June6, 1944 (AshleyB. Gurney). Paratype,i, Hill 700, altitude
600 feet,December28, 1944 (RobertStaples)- along cascading
streamin thick jungle - StaPles.
In its general appearance the present ily is much like Trentepohlia(Mongoma) brevicellula Alexander, differing especially in the venationof thi radial Iield, including the relative proportions of cells R, andR* at the margin.

66. Trentepohlia (Mongoma) vitrina sp. n.
Body whitish yellow without pattern; wings vitreous, veins very
pale, difficult to see, especially in balsam slide mounts; vein Rr-n long,subequal to Ro; celt 1sf M, long, from about one-half (as figured) tonearty two-thir'ds vein R. belond it, basal section of M. slightly retracted.

Male. Length,about 6 mm.; wing,5.5 nlm.; antenna,about
1.5 mm.
Female. Length, about 6 mm.; wing,5 mm.; antenna,
about 2 mm. Generalcolorationof body very pale whitish yellow,
unpatterned,including also the halteres and legs; head morewhitened,the black eyes strongly contrasting;anterior vertex
narrow, about one-thirdthe antennalscape.Antennaewith scapeand pedicel whitened, f lagellum light brown, segmentssubcylindricalwith short verticils.
. Legs with severalstrong black setaeon all femora.Wings(Fig.69) vitreous,veins very pale and difficult to see,especiallyin balsamslide mounts.Trichia of veins reducedin number andsize, including especiallyveins R, Rr and distal sectionof Rs.
Venation: R:]-+ long, subequalto R2; vein Rr oblique,about
one-halfR+i cell lst M2long, from about one-halfto nearly twothirds as long as vein R5 beyond it; basal sectiouof vein M3
slightly retracted,m-cu shortly before fork of M.
Habitat: Samoa.
Holotype,d, Vailima,Upolu, March 6, 1971 (Noel Krauss).
Allotopotype, 9 , on pin with type. Paratopotype,d, ofl pin
with types.
As presently known there are four yellow regional species of tht
subgenus Mongoma Westwood, best distinguished among themselves b1'
the body size and venation. The unusually large Trentepohlia (Mongomu\
spectralis (Edwards), of Samoa, is readily told by the size (rving'com m only m or e t han 7 m m . , in c a s e s t o 9 m m . ) . T h e m o s t s i n l i l a r r r t
th es e s pec ies is T. ( M . ) polic ep h a l o A l e x a n c l e r , u 'i t l t 's p r e a t l f r o t t t l l t t '

72

vn p
E '\ V '+//t i \ \

t\1 ,

n
AI
t./

$/4,

/\

F

Toxorhina
venation.
Fig.
71.
Flf
t,o.solis Alexander;llf
I
I I t,\t)thinal't|trhrut(Psiloconopa)
nnlulensis
rtrrl'lr\riI
Eriopterar,v('ilirllor.
-.1:ig.72.
l_firilthlilnt(Molophilus)
sp. n.i
flErrnrfFf.
rf,rl',
lr\t,otrvAlrrr
lilt.
subvinnulus?ii. IlololthilusH€13 f rlf ,l,l,t l'trrrrr
male
hypopygium.
Stt,rin|Jtnriu
Alexander;
1l,il(lanopinax
I tg
Fig.
hypopygium.
i rC
li
Alc.rander;malel,,r,,tl|Irt
I l.'\t)tltlilill
ttillt,inario\lra, aedeagus;iFt,'r,,r/r/tr,tttttttnhlnilt\firl'lr'\lrr; rrr;rlc hyt)opygilrtrr.
SymbolS:,- l,i.i!ltl'
g.rr;rlrrrqrltlrir,
tergite,,lletlbtylf,1'
ittlcrlrase:
t,gth
,t
s,9th
sternite;
t,

?6

#--

312

Al e x a n d e r,

StudiaEnt.,vo;l.16, fasc. 1-4,outubrode 1973

A u stral asi anC ranel l i esII

eas te rn o ri e n ta l to n o rth e rn A u s tral i a,w hi chdi l fersfromthe.present
the small cell.,JsfM, which is subquadrate
fly in the venation,i".i;;l;;to one-fifth as l6ng as vein R.u beyond
one-iourtft
iri ouline and scarcely-Frtiiip'"pi".
40: 266, plate 1'- fig' 14:..re2e)'
Sci',
Jour.
ii' i;T;;;"d.i,the wing veins are darker yellow toin '"Lf of these simiiar species'readi'ly seen a[ainst the yellow ground'Uio*ni.ft yellow and are313

lt :lrrrwrr;basal style profoundlybifid, its outer arm a slenderhfagkcncdspine,inner arm a flattenedglabrousblade that narrows
Inkr a slutttlcracute spine. Phallosomicplate pale, depressed,theapl(al nlargin, produced into f imbriations, on central area
long and slender,setoid.Aedeagusshort,the basestout.
hecurrrirtg
llabitat: New Guinea.

67. Erioptera (Psiloconopa)malulensisAlexander( Psitoconopa)
E ri o ptera'572-573; 1947'

m alutensis

Alex ander ;

Ann

M ag'

N at

Hist

'

(ll)

14:

1933
Type 6 from Mafulu, Papua,4000 feet, December
Male hypopygium(Fig' 72)'
(Cheeiman).

lfirkrtypc, 6, Itouda, Wisselmerenarea, NetherlandsNew
0ulrtca, 1,500 meters,August 14, 1955 (J. Linsley Gressitt);
Bhlt,r;rMuscum.
'f 'hc nrost nearly related species are Molophilus (Molophilus)
grtrxlllionus Alexander and M. (M.) vinnulus Alexander, both of New[lulnen, all most readily totd by the distinctive hypopygial characters.

68. Molophilus(Molophilus) subvinnulussp' n'yel]gwl dorsal thoracic
Allied to vinnulus; rnesothorax brownish,
black; legs brownish
pleura brown, unt.nnui tft"?, ptOit"l enlarged'
vaguelyyellow'brownishblack, genua u.ry nuiio*ty- *tr,iteneO; wingJ
bifid'
dististyles
both
with
hypopygiurir
patterned with pale brown;" male -ipicbi,"';i[
inner branch of basal style
unO acute atili".f..".0;t

69. Styringomyia melanopinax Alexander,ttr tntum r l u

nr r l an o p i n a x

A l e x a n d e r;Ann.

Ma g .

Nat.

hist.,(1 1 )

l4:

5 7 5 -5 7 7 ; 1 9 1 7 ,'f'y1re from Kokoda, Papua, 1200 feet, August-SeptemberdMale hypopygium(Fig. 7a).
l9ilil (Cltcesnran).

a slender acute sPine.

andM a l e. Length, about 3'5 mm'; wing, 4 mm' Rostrum
large'
pedicelyellow'
palpi black. Antennaeshort; scapeobscure
to long-oval'
ttu.t, flagellum brown; flagellar segments oval
dark
shorter than the verticils. Fiont yellow, head posteriorly
brown.
pleuraPronotum dark brown. Mesonotumbrownish yellow'
Halteresstripe'ot
area
dorsalcleareryellow, with a broad brownremainder ot
yellow. Legs with coxae and trochantersyellow;
tibialwhitened'
iegs brownish black, temoral tips very narrowly
patternedbasesless evidentlyso' Wings brownish yellow' vaguely
costal fieldwith pale brown, especiallyover the cord, base ofmoreslightly
base
wing
A;
and across out.l, .nd of cell 2nd
R2
Venation:brown'
yellowed; veins pale brown, trichia dark
anci
two
about
just distad of level ol r-m; petiole ol cell M3
posterior
one-half times rn-cu,'vein 2nd A ending nearly opposite

70. Toxorhina (Toxorhina) basalis Alexanderl r ttr hl r tr tl l r t.r r tr hin u )b c s o l i s A l e x a n d e rl

A n n . Ma g . N a t . H i s t . , (1 2 ) 4 : 6 0 1 -6 0 1 ; 1 9 5 1 .

'l'ypr' 3 f ronr the Cyclops Mountains, Netherlands New(lttirrt'it,i1500fcct, March 1936 (Cheesman).Venation(Fig. 70).

7l . Toxorhina (Toxorhina) pulvinaria AlexanderI r ttthtnl
I I t\l ttlt i n u l{t1x 0i r r , l 1t5 t l

p u l v i n a ri a

A l e x a n d e r;Ann.

Ma g .

Nat.

Hist.,(1 2 )

3:'l'ypt' tl, llollanclia,Netherlands
New Cuinea,April 18, 1945(llarry llrrogslr;r;rl).
Malc hypopygium(Fig. 75).

72. Toxorhina (Toxorhina) staplesi sp. n.
end ol m-cu.
hypoMalebrown'
dark
Abdomen, including hypopygium,( lerreral lolunrliorr of thorax light brown, pleura more yellowed/' witlr
pygium (Fig. 73) geneialty ut in vinnulus'Ninth tergite'f E n l t a l l y , r r ) s l n u n sl i g h l l y l o n g e r th a n th e b o d y; m a l e h yp o p yg i u m w i th
setac' I t€ticr lf rrvcn powcrful black spinoid setae on mesal face of basistyle;pfJt.tio.'Ooider convexlyrounded,provided with numero.ust l l € t n f i t t t r r l r l r ,r . o l r l r r su ; l l o th d i sti styl e s w i th a str o n g sp i n e a t o rshallott'with a marginal concenirationon either side of a small
thickent'tl i r p y l r r r l n r i r l l r . r r g l l r
median emargination,cephalicborder of plate heavily
A l a l r ' l r .r r g l l t, r .x( 'l r r ( l i n g r o sl r u n l , a l l o u t 5 r n r r r .; w i n g , 4 .2 - 5base clivitlt''land sclerotized.Outer iirtittylt, d, bifid, beyond
nrrl , tlalllrlt,
i tl r o tr l i r ,5 - ( i r ttr tt.
intotwoflattencclblac|esthathavethesantegt.ncralc()nf()rlllati{)||,

rc"

31 4

Alex ander ,Aus t r ala- s i a nC r a n e f l i e s I I

F e m a le. Length, about 6-6.5 nrm.; wing, 5-5.2 nrm.;
rostrum,about 6-6.5 mm.
Rostrumin male slightly longer than body, light t,r clarker
brown. Antennaebrown. Head clear light gray.
Pronotum and cervical region clark brown. Jlcsonotal
praescutumlight brown, margins light gray pruinosc. lateralpraescutal
stripesslightlydarkerthan thc ground;centers,rf scutal
lobes darker brown, scutellumyellowishbrown to brorvn; p()stnotum brown, conspicuouslygray pruinose.Pleura chicflt' lightbrown above,cleareryellowbelow.Haltereswith stemlight br,irvn,
yellowed at base, knob darker brown. Legs with coxa( andtrochanters
yellow; remainderof legs yellowishbrown, thc darker
appearance
producedby the abundantbifid setae.Wings t Fig.
7l) weakly darkened,veins dark brown. Trichia on Rs anrl thelongitudinalveinsbeyondcord,lackingon M3-a andMa. \rcnalioll:
Cell M, open by atrophy of m,' cell 2nd A narrow.
Abdominaltergitesslightly bicolored,light brown, p,,st(ri()rborders dark brown; sterniteslight yellow, posterior ritargirts
narrowly brown; subterminalsegmentsmore uniformlyblaikrnccl
to form a ring, more extensiveon tergites; hypopygiunrliqht
brown. Male hypopygium(Fig. 76) with basistyle,b, on tttesal
face with a linear seriesof sevenmodifiedblack setae,tltc,rttter
ones progressivelysmaller.Interbase,t, a relativelystotrt palcd, arnitcl
blade,gently curvedto the obtuseapex.Both dististyles,
with a strong spine,the outer style with this at near two-thircls
the length, the inner style about twice as large with the spitte
at or closeto middlength.Arms of aedeagusshort.
Ilabitat: SolomonIslands.
Flclctype,6l, Empress Augusta Bay, Bougainvillelslarttl,
along KoronrokinaRiver, Septcmber25, 1944 (Robert Staplcs)- anrongrvashedout roots of banyau,associatedwith Treticpoltl:n(Pleuontottgoma')novaebrittanicae
Alexander.Allotopotypc. ?.
Paratopotypes,d9, Septenber ll-28, 1944. Paratypes.d9,
Bougainvilic,July-August 1944 (Ashley B. Gurney).
Th is inter es t ing f ly is dedic at ed t o t he c o l l e c t o r , D r . R o b e r t S t a p l e s ,my former student at the University oI Massachusetts, now on th,eEntomologi,:Llfaculty of the University of Nebraska. I am deeply intlebtecl
to him fo r,r . r . nU new and r ar e s pec ies o f t h e s e f l i e s t h a t w e r e t a k e nwhile he r s stationecl at Bougainville during the war period. Other
regional r, rbers of the genus that possess modified spinoicl setae onof the hypopygium include Toxorhina (Toxorhina) rtoelianath,e basistri 7. (7.) perproducta Alexander, both of Fiji, differing
Alexander,pygial characters, especially the dististyles.
in otheri