JOURNALOF

THE

New York EntomologicalVOL. LVI

Society
No. 3

1948
SEPTEMBER,

RECORDS AND DESCRIPTIONS OF NEOPTROPICALCRANE-FLIES(TIPULIDIE, DIPTERA), XXIIIBY CHARLES P. ALEXANDERAMHERST,MASSACHUSETTS

The preceding
title was publishedinpart under this general
this JOURNAL (vol. 55: 173-184; June 1947). The types of the nov
elties described herewith
in my collection
are preserved
through
the appreciatedof the various collectors.
Two species
kindnessare based on materials
in the UnitedStates National
Museum,
sent

to me

for determination

by Dr. Alan

Stone.

Genus Austrolimnophila Alexander
Austrolimnophila
Generalcoloration(Austrolimnophila)ofthoraxyellow,bulbulifera,
the pleura

new species.
withatransversebrown

girdle occupying the mesepisternum; femora obscure yellow, the tips nar
rowly blackened, remainder of legs brown, the tarsi extensively whitened;wings subhyaline, restrictedly patterned with brown; m-cu about threefourths its length beyond the fork of M; male hypopygium with themedian region of tergite produced into a narrow lobe, its tip shallowly
emarginate; dististyle complex, the main body with long coarse nodulose
sets;

face

of

style

with

a

low

blackenedflange.

MALE.-Length about 11 mm.; wing 10 mm.; antenna about 1.9 mm.
Rostrum and palpi black. Antenne short; scape and pedicel obscure yellow, flagellum black; flagellar segments subcylindrical to cylindrical, thelongest verticils unilaterally arranged, much longer than the segments.
Head brown.
Pronotum testaceous, paling to yellow on the sides and on the propleura.
Mesonotal prescutum obscure yellow, the stripes not or scarcely indicated;
lateral prescutal borders slightly darkened, representing the upper end of
the pleural girdle described below; scutum and scutellum chiefly pale;[VOL.LVINEW YORK ENTOMOLOGICALSOCIETY

138

mediotergite dark brown; pleurotergite yellow, its posterior border moredarkened. Pleura pale yellow, with a narrow dark brown transverse girdle
extending from the prescutum, as described, across the entire mesepisternum
and sternum. Halteres elongate, stem obscure yellow, knob dark brown.
Legs with the coxse and trochanters pale yellow; femora obscure yellow,
the tips narrowly blackened; tibiae and proximal half (hind) to three-fourths(fore) of basitarsi medium brown, the remainder of basitarsi and segments
two to four white, the terminal segment infuscated. Wings with the groundyellow on cephalic third, more grayish on posterior portions; a restrictedbrown pattern, including spots at origin of Rs, cord, outer end of cell
1st M,, R2 and adjoining veins, fork of M1+2, and near outer end of cellR3; a faintly indicated dusky wash in center of cell R; veins dark brown,
conspicuous. Venation: Sc, ending shortly beyond the fork of R,+3+4,Sc2 a
short distance from its tip; R2+,+4about one-half longer than m-cu; R,1+
shorterthancell M,

about

fork

cord

R,+3+4; anterior
twice mn; m-cu

aboutcell BR mostoblique,proximad;its own length
three-fourths

petioleofbeyondtheof M.

Abdominal

tergites dark brown, the caudal margins of the more basal
segments narrowly yellow, the outer segments more uniformly darkened;hypopygium chiefly black; sternites bicolored, chiefly yellow, with a narrow
blackened ring before midlength. Male hypopygium with the tergite pro
duced caudad into a medium depressed-flattened lobe, its apex shallowly
emarginate.

Basistyleon mesal

near

face

apex witha smalltubercletuftedwith long yellow setra. Interbase a small curved yellow structure, the tip
acute. Dististyle complex, the main body more or less triangular, extended
into a long slender rostrum, the concave mesal face of style with several
strong yellow setae that are provided for most of their length with globular
excrescences; on face of style a low blackened lobe or flange, its surface
with dense microscopic setulae.
Habitat.-Southeastern

Brazil.
Sao Paulo,(, Boracea,

Holotype,

altitude

800 meters,

August1947 (John Lane).
similargenerally

Althoughto species

such

as Austrolimno

phila (Austrolimnophila) candiditarsis Alexander, A. (A.)
pallidistyla Alexander, and other regional forms, the type ofmultipicta
crossvein.

is more as in the various
the male hypopygium
as A.
the subgenus(L.)
Limnophilella
Alexander,
m-cu
retracted
all
the
much
of
which
have
Alexander,
I am deeply indebted to Dr. John Lane for many fine

Tipulidae

from Sao Paulostructure
speciesofof

and Riode Janeiro.

Genus Pseudolimnophila Alexander
Pseudolimnophila
Alliedto pluto;

plutoides, new species.
size

large

(wing,

male,

over

9 mm.);

generalcoloration

139

ALEXANDER:CRANEFLIESSEPT., 1948]

black, the premscutum and scutal lobes more reddish brown; legs and wings
strongly blackened; R,2+3+from two to three times as long as R?; abdomen
dark brown, the hypopygium more yellowish brown.
MALE.-Length about 9-10 mm.; wing 9-10 mm.; antenna about 1.5 mm.
Rostrum and palpi black. Antennae black, the pedicel a very little paler;
flagellar segments passing through long-oval to subcylindrical, the verticils,ofespeciallythe outersegments,

long

and

Head

conspicuous.

black;

anter

ior vertex relatively broad, about two and one-half times the diameter of thescape.

Pronotum black. Mesonotal

prescutum and scutal lobes dark reddish
brown, virtually unpatterned; posterior sclerites of notum more brownish
black. Pleura, including the dorsopleural membrane, blackened. Halteres
infuscated. Legs with the coxse brownish black; trochanters obscure yellow;
remainder of legs dark brown, the femoral bases restrictedly obscure yellow.
Wings with a strong blackish tinge, especially in the outer radial field;
stigma and narrow seams over cord and outer cell 1st i21 still darker brown;
veins

Rs

Venation:brown.

long,

subequal

to veinR4;

R2+3+4 from

two

to

three times R?; veins ,R and R4 extending generally parallel to one another,
the former arched at near midlength, slightly widening the cell at thispoint; cell M1 more than three times its petiole; m-cu at near two-fifths tothree-fifths

thelengthofcell

1st M,.

Abdomen dark brown, the hypopygium a trifle brighter, more yellowish
brown. Male hypopygium with the ninth tergite bilobed, the lobes narrower
than the U-shaped median notch. Outer dististyle with the entire mesal
face densely covered with erect setae. Gonapophyses small, bispinous.
Habitat.-Peru.

Holotype, dd,Carpish, Huanuco, in dwarf fog forest, altitude
2800meters (9100 feet), November 4, 1946 (Woytkowski). Para
topotype, 3,November 14, 1946.
The nearest ally of the present fly is Pseudolimnophila pluto
Alexander, from the Tropical Zone of Amazonian Ecuador, asmaller and much paler fly. The male hypopygia of the two
species are very similar and indicate a close relationship.
Genus
Teucholabis
Size

small

(Euparatropesa)(wing,

female,Teucholabis

Osten

Sackennewlaetfica,
species.
5 mm.);
alliedto lindneri;head,

pronotumandpleura orange; mesonotum shiny black, restrictedly patterned with yellow;
femora yellow, the tips blackened, broadest on the fore femora; wings
whitish subhyaline, restrictedly patterned with brown; dark areas at arculus
and origin of Rs interconnected in the costal field; Sc ending about opposite
one-third the length of Rs; R2 nearly twice R2+?+4;abdominal tergites darkbrown, sternites obscure yellow.

140

NEW

about

FEMALE.-Length

YORK

ENTOMOLOGICAL

5 mm.;[VOL. LVISOCIETY

5 mm.

wing

Rostrum reddish yellow, shorter than remainder of head; palpi brown.
Antenna with scape and pedicel yellow, flagellum brown; flagellar segments
much

oval,

than

shortertheverticils.

Headorange.

Pronotum orange. Mesonotum shiny black, restrictedly patterned withyellow, including the suture and most of the scutum, the lobes of the latterwith polished black centers; humeral region of prsescutum extensively yellow; pleurotergite black. Pleura uniformly orange yellow. Halteres withstem weakly darkened, knobs light yellow. Legs with all coxse and tro
chanters orange or orange-yellow; femora yellow, the tips broadly black,on the fore legs including the outer three-fourths, on middle and hind legs
about

the distalor sixth;

fifth

fore

and

tibiaetarsi

middleblack;

andhindtibia dirty white, blackened at base and tip; middle basitarsi obscure yel
low at proximal end, remainder of tarsi black. Wings whitish subhyaline,
restrictedly patterned with brown; dark areas at arculus and origin of Rs,
interconnected without a break in cells C and Sc, the latter area completely
crossing the wing, widest in cell Cu; a narrower but darker band at cord, ex:tending from C to Cu, deepest in color at the stigma, nearly parallel-sided;
no darkening at wing-tip; veins brownish yellow, darker in the patternedareas. Venation: Sc ending about opposite one-third the length of Rs, Sc,
a

short

distance

beforetheoriginofthelatter;RB nearly

twice

R2+3+4 orR1+2; cell R, small, vein R4 subequal to R3,+4; cell M2 open by atrophy ofbasal

sectionof M,3, cell 2nd M2

subequalin length

to its petiole;

m-cu

about

two-thirds its length beyond the fork of M; vein 2nd A nearly straight formost of its length. In lindneri, the darkened area at and beyond arculus
and that at origin of Bs broadly interrupted in cells C and Sc; vein Sc
longer, ending beyond one-third the length of Bs; R2 at fork of Bs; m-cu
at

or

close

to

the

fork

of M.

Abdominal

tergites dark brown, sternites obscure yellow; subterminal
segment orange; genital shield extensively darkened. Ovipositor with thevalves orange; cerci strongly upcurved.
Habitat.-Peru.

Holotype, ?,Yarinacocha, Loreto, near Pucallpa, on theUpper
Ucayali River, altitude 180 meters (Schunke).
Teucholabis (Euparatropesa) laetifica is closest to T. (E.)
lindneri Alexander, of the Bolivian Chaco (Konowia, 12: 42-44,
figs. 1-2; 1933), differing in the coloration of the body, especiallythe mesonotumand venation,
Teucholabis

and pleura,and in the details
as compared above.

of wing

pattern(Teucholabis) atrolata, new species.
dark
Antennaebrown throughout; anterior vertex black, elevated into asmall tubercle; prescutum yellow, with three chestnut brown stripes, darker

SEPT., 1948]

141

ALEXANDER:CRANEFLIES

behind; scutellum broadly yellow behind; pleura brown, heavily pruinosewith silvery; femora yellow, the tips and a medial band broadly blackened,
mid-femur with the base similarly darkened; wings subhyaline, with threedark

the apical
one narrow;bands,
two to six withsternites
black
broad

costalfringelateralareas;

short;
male

abdomenhypopygium

yellow,withthe spine of the basistyle relatively large; strongly curved; outer dististylelong and slender, unequally bidentate at tip; inner dististyle conspicuously
bifid, both arms terminating in points.
MALE.-Length about 9.5-10 mm.; wing 7.5-8 mm.
Rostrum obscure yellow medially above, dark brown on sides; palpi
black. Antenna dark brown; flagellar segments passing through oval, longoval to elongate-cylindrical, the verticils long and conspicuous. Front yel
low; anterior vertex narrow, black, raised into a small tubercle near its
anterior end; posterior vertex paler brown or yellowish brown.
Pronotum chiefly yellow, variegated with chestnut brown. Mesonotal
prescutum with three chestnut brown stripes, paler in front, darker andalmost contiguous behind, the humeral triangles yellow, lateral preescutal
borders more obscure; median region of prsescutum before suture and thecentral portion of scutum more grayish yellow; scutal lobes extensively
darkened, almost black, their posterior portions yellow; scutellum broadly
yellow behind, the base more testaceous yellow; mediotergite testaceous
yellow, with almost the posterior half dark brown. Propleura chestnut;
remainder of pleura darker brown, heavily silvery pruinose; mid-meso
sternum narrowly yellow, the posterior thoracic sternites entirely yellow.
Halteres yellow, knob brown, the apex paler. Legs with coxye orange-yellow,
the middle and posterior pairs more pruinose; trochanters chestnut-yellow;
femora yellow, the tip broadly and conspicuously blackened; middle femorawith a basal and a median dark ring, alternating with yellow annuli, theblack tip a little narrower than the subterminal yellow ring; darkened basal
annulus wider than the subbasal yellow one; hind femora similar but withthe basal darkening less evident; tibiae yellow, the tip narrowly to scarcely
darkened; tarsi yellow, the outer two segments blackened. Wings subhyaline, trivittate with brown, including a broad complete band at cord, darkest
at stigma; wing-tip narrowly darkened; subbasal band at level of originof Rs and across vein 2nd A; veins yellow, darker in the patterned areas.
Costal fringe short. Venation: Sc ending about opposite midlength of BRs,
Sc, at near one-fifth the length; RB slightly oblique, in cases weakly sinuous;
m-cu about one-third to one-half its length beyond the fork of M.
Abdomen conspicuous hairy; tergites and hypopygium yellow; sternites
yellow,

darkening.
segment,

two

segments

to sixeach witha conspicuous

brownishblack

lateralSternal pocket of segment five close to the posterior end of

more

or

less

triangularin outline,

thelongest

setae

at

the

sides;

pocket of segment six consisting of six or seven strong curved sete in a
vertical row on either side, directed mesad. Male hypopygium with theapical spine of the basistyle relatively large, strongly curved, the apical142[VOL.LVINEW YORK ENTOMOLOGICALSOCIETY

third narrowed into a long straight spine; mesal flange coarsely and ir
regularly toothed. Outer dististyle long and slender, unequally bidentate atapex, the surface with long scattered seta. Inner dististyle conspicuously
bifid, the larger beak terminating in a strong spine; lateral lobe terminating
in a short conical point.
Habitat.-Panama(Canal Zone).

Holotype, <, Barro Colorado Island, January-March 1944
(Zetek No. 5126); United States National Museum. Paratopo
types, 2 3<; one of these is much paler, including the leg pat
tern and may not prove conspecific.
The allied species include Teucholabis (Teucholabis) angust
apicalis Alexander, T. (T.) catharinensis Alexander, T. (T.)
melanocephala (Fabricius), T. (T.) neosalva Alexander, T. (T.)
perangusta Alexander, and T. (T.) salva Alexander, all having
the legs and wings banded and with the sternal pockets of thefifth and sixth abdominal segments of approximately the same
type. The present fly differs in coloration of the body, legs andwings, especially the unusually heavy leg pattern. There arefurther differences in the male hypopygium, especially the basistyle and inner dististyle.
Teucholabis(Teucholabis) hypomela, new species.
Allied to flavithorax; size small (wing, female, 6 mm.); pronotum black;mesonotum and dorsal part of pleura orange, the ventral pleura and sternum
legs withblack;

allcoxe

trochantersandremainder

black;

oflegs

black,

the basal two tarsal segments obscure yellow; wings with a strong brownish
suffusion; Sc ending about opposite three-fifths Rs; vein BR transverse,
about one-half R2+3+4;ovipositor with the cerci yellow.
6 mm.;

about

FEMALE.-Length
segments

thesubglobular,

6 mm.

wing

Antennae black throughout; basal flagellar

Rostrum and palpi black.

ones

outer

oval.

more

Head

black.

Pronotum black, the sides of the scutellum orange. Mesonotum
the

third

cephalicofthepleura,anepisterum
sternopleurite,
mesonotum
and the pleurotergite
coxa and trochanters
blackened;
obscuresegments

brownishstrongtion:Scnear

relatively

one-third

than R,-+;yellow,
suffusion,

cell

the

long, Sc

black

triangularorange,Proofthe remainder
region black,allinfuscated.
Halteres
Legs with
two tarsal
the basal
and tibiae black,orange.
femoratarsala triflesegments
darker;black.
veins

dark

Wings
brown.

with

a

Vena

ending about opposite three-fifths Rs, Sc, atabout one-halfvein R, transverse,
length;m slightly
1st M2 slightly
widenedoutwardly,
the

area.

and meralouterstigmaawithprsescutum

R,2++4 and longer
longer and more

SEPT., 1948]

143

ALEXANDER:CRANEFLIES

arcuated than the basal section of Mi3; m-cu about one-half its length be
yond

the

fork

of M.

Abdomen black; cerci slender, yellow strongly upcurved.
Habitat.-Peru.

Holotype, ?, Yahuarmayo, February 8, 1910 (C.H. T. Townsend); United States National Museum.
The most

similar

speciesare Teucholabis(Teucholabis)

cunei

formis Alexander, T. (T.) dileuca Alexander, T. (T.) rufithorax(Wiedemann), T. (T.) rufula Alexander, and T. (T.) schineri
Enderlein, occurring from Mexico to southeastern Brazil, alldiffering
venation.

in detailsof coloration

The blackand mesosternum,of the bodyand narrowly
in conjunction
thewithprothorax

andlegs and in theblackened
pleura

small

size,

shouldbe emphasized.
Teucholabis (Teucholabis) subargentea, new species.
Allied to argentea; general coloration polished black, the mesonotum
variegated with yellow; pronotum pale yellow; pleura black, with a broad
silvery white stripe; femora yellow, the tips of the fore pair blackened;wings yellowed, crossbanded with brown; sternal pockets of male strongly
developed; male hypopygium with the inner dististyle very unequally bident
ate, with two approximated setae on disk of style; aedeagus a moderately
compressed blade, the lower edge with two long setaa.
MALE.-Length about 6.5 mm.; wing 7-7.1 mm.
Rostrum orange; palpi black. Antennae black throughout; flagellar
segments passing through subglobular, short-oval to long-oval, the verticils
longer than the segments. Head dull black.
Pronotum and pretergites pale yellow. Mesonotum polished black, theposterior portions of the scutal lobes, posterior border of scutellum andlateral margins of mediotergite obscure brownish yellow. Dorsal thoracic
pleura chiefly polished black, the dorsopleural region yellow; a broad silvery
white longitudinal stripe extending from behind the fore coxa to base of
abdomen, widened behind; ventral sternopleurite blackened, the remainderof venter pale. Halteres black, the apices of the knobs obscure yellow. Legs
with the fore coxse blackened the remaining coxse yellow; trochanters
yellow; femora yellow, the tips of the fore pair blackened, of the remaining
femora scarcely darkened; fore and middle tibias and tarsi black; hind tibiae
obscure yellow, the tips narrowly blackened; tarsi black; posterior basitarsi
dilated on about the proximal fourth. Wings with a weakly yellowed ground,the prearcular and costal fieldsmore whitened; a medium brown crossbanded
pattern, including a narrow band at cord, crossing the wing from the darkbrown oval stigma to the posterior margin; wing tip broadly darkened, ex
tending basad about to the level of the outer end of cell 1st M2, the posterior

NEW YORK ENTOMOLOGICALSOCIETY

144[VOL.LVI

margin of cell M. pale; a third band at one-third the wing length, extending
fromR

to the anal margin

at vein2nd A;

veins

brown,

lightindarkerthepatterned areas. Venation: Sce ending just beyond midlength of Rs, Sc2 amoderate distance from its tip; R2 about twice R,2++4; branches of Rs ex
tending generally parallel to one another, cell 12 correspondingly wide atcell B4 slightly

margin;at basalwidenedm-cu

third;

close

to forkof M.

Abdominal tergites dark brown, sternites paler; sternal pockets darkcolored and conspicuous; hypopygium black. Sternal pocket of segment fivelarge and conspicuous, with very numerous setse; sternite six with a narrowlytransverse

oval

area on posterior
six or seven very

on

half;

anterior

verticalrow ofwidelyseparated. Male hypopygium within

subterminal

stout

position,

sete

long
at

Outer

dististyleasinuousspine,

thesurfaceterminal

halfof

directedwithsternitethemesad,arows

the appendage of the basistyle

the

base,

straight spine; two darkened marginal

are

that

outer

narrowed

third

into

a

lobes nearer mesal part of style.

rod thatelongatewithlong coarse

intonarrowed

is graduallysetee fromscabrousa

tubercles.

Inner dististyle with beak very unequally bidentate, the outer spine elongate;
two approximated elongate bristles on disk of style. Aedeagus a moderately
compressed flattened blade, at apex produced into a blackened beak or spine;
surface just back from tip with a few microscopic scabrous points; lower
edge

of

with

aedeagus

two

long

setae.

Brazil.

Habitat.-Southeastern

Holotype, 3 Boracea, Sao Paulo, altitude 800 meters, August
1947 (John Lane). Paratopotypes, 4 S.
The most similar described species is the smaller Teucholabis(Teucholabis) argentea Alexander, which differs in the detailsof colorationandin the structureof the malehypopygium.

Teucholabis(Teucholabis) elissa, new species.
Allied to laterospinosa; mesonotum chiefly blackened, prothorax reddish
yellow; rostrum reddish; halteres dusky, the knobs yellow; wings subhyaline
on proximal half, very weakly infuscated beyond the cord; stigma darkbrown,

thecordnarrowly

seamed;apicallobeofbasistyleof malehypopygium tumid, terminating in a straight spine; outer dististyle unequallybispinous

at apex;surfaceofedeagus

with

about

15 strong

sete.

MALE.-Length about 4.5-5 mm.; wing 4.8-5.5 mm.
Rostrum reddish; palpi brown. Antenna with the scape and pedicel
obscure yellow, flagellum brownish black; basal flagellar segments sub
globular, the outer ones passing into oval; proximal segments abruptly
narrowed into very short apical necks, producing a moniliform effect. Headreddish.
Prothorax reddish yellow. Mesonotum chiefly black, the humeral regionof praescutum, suture, scutellum, and, in cases, about the cephalic fourthof the mediotergite reddish to yellow; pleurotergite black. Pleura reddish

SEPT., 1948]

ALEXANDER:CRANEFLIES

145

yellow, the ventral sternopleurite more infuscated; posterior pleurites somewhat brighter and more or less pruinose. Halteres dusky, knobs yellow.
with

Legs

coxie

the

trochanters

and

fore

yellow;

femorachieflytheblack,

basal fifth yellow; tibiae dark brown, the tips and the tarsi black; middle
and

hind

femoratibse andyellow;basitarsi

obscureyellow,

the outertarsalsegments black; posterior tibiae at near five-sixth the length with a slightly
swollenarea.

glandular

Wings

on proximal

subhyaline

very

half,

to

weaklyscarcely infuscated beyond the cord; stigma short-oval, dark brown, sending
a muchnarrower

seam

back

over

the

cord;

cell

Sc

above

the

arculusrestrictedly darkened; veins brown. Venation: Sc short, Sc1 ending just beyond
origin of Rs, Sc2 some distance before this origin; R2 at or just beyond
the

fork

of Rs;of Rs

branchesonanother

to oneparallel

the

about

basal

half, R5 thence bent strongly caudad to the wing tip, cell I4 very wide at
margin,
of M.

than

greatercell R2;cell 1st M2

closed;m-cu

at or close

to thefork

Abdominal tergites and the hypopygium black; sternites weakly bicolored,
black, the narrow apices yellow, the amount of pale color decreasing on theouter segments. Sternal pocket of segment five comprised of a transverse
semicircular row of relatively short setae; sternite six without a well-defined
pocket. Male hypopygium with the apical lobe of basistyle a tumid structurethat terminates abruptly in a straight spine; mesal flange of basistyle
blackened but smooth. Outer dististyle a strong darkened club, graduallywidened beyond the base, the apex truncate, the outer angle thence pro
duced

into

a

strong

spine

that

is about

one-halfthelength

of

the

base,

the inner apical angle with a much smaller spine; inner margin of basal
part of style with a series of strong serrations or spines. Inner dististylewith the apex unequally bifid, the lower arm or flange with marginal micro
scopic serrulations; a strong basal lobe bearing three very long setee.
_Edeagus conspicuously compressed into an obtuse blade, the apical parta low darkened knob; surface of aedeagus with about 15 strong setee.
Habitat.-Peru.

Holotype, A, Chanchamayo, Junin, altitude 1300 meters, Feb
ruary 10, 1941 (Schunke). Paratopotypes, 5 ', August 15,
1941-February 10, 1942 (Schunke).
The most similar species include Teucholabis (Teucholabis)
laterospinosa Alexander and T. (T.) tullochi Alexander, which
differin the detailsof colorationand structureof the malehypopygium.

Genus Gnophomyia Osten Sacken
Gnophomyia (Eugnophomyia) glabripennis, new species.
Size large (wing, male, over 8 mm.); general coloration of body, antenna,
halteres and legs black; wings with a strong blackish ground, the centersof mostofthecells morewhitened

to produce

a streakedappearance;

cells

NEW

146

YORK

ENTOMOLOGICAL[VOL. LVISOCIETY

of wings glabrous; male hypopygium with the dististyle elongate, its tip
narrowly obtuse, the lower surface with abundant very long black setre; on
outer

face

of

at

style

base

with

a

obtuse

small

lobe;

stronglyedeagusrecurved at midlength.
about

MALE.-Length

7.5 mm.;

8.3 mm.

wing

Rostrum and palpi black. Antenna black throughout; flagellar segments
terminalsubcylindrical;

segmentmuchlarger

the

than

Head

penultimate.

dull black.
Thorax uniformly dull black. Halteres and legs black. Wings with astrong blackish ground, the centers of most of the cells beyond wing basewhitened to produce a streaked appearance; veins brownish black. Cells of
wing without macrotrichia. Venation: Scl ending a short distance before
level of fork of Rs, Sc1 alone longer than r-m; Rs long; vein RB oblique, its
cephalicend

portion;cell 1st M2

entireto subatrophieda little shorter

and
than

basad
than
lying moreat nearvein M4; m-cu

the posterior
theone-third

length of cell 1st M, or about one-half its length beyond the fork of M;
vein 2nd A sinuous.
Abdomen, including hypopygium, black. Male hypopygium generally asin the subgenus; apex of basistyle produced beyond the point of insertionof the dististyle into an elongate conical point, the tip subacute. Dististylea little longer and larger than the apex of the basistyle, its tip narrowly
obtuse;lower

surfacewithabundant

very

long

black

setae;

on

outer

face

at base with a small obtuse lobe. ZEdeagus strongly recurved at midlength,
the basal part more expanded.
Habitat.-Peru.

Holotype, A, Chanchamayo, Junin, altitude 1350 meters,
March 10, 1941 (Schunke).
Gnophomyia (Eugnophomyia) glabripennis is readily told
from all other regional species by the glabrous wings. All
otherspecieshave numerous

in the outermacrotrichia

cells.

In

general appearance, the present fly is most like Gnophomyia(Eugnophomyia) funerea Alexander, from which it is further
distinguishedbythe streaked

wings

and

of

the structure

themale hypopygium.
Genus Gonomyia Meigen
Gonomyia
Allied(Gonomyia) subunicolor, new species.
to unicolor;

generalcolorationof mesonotal

praescutum

andscutumdark brown, the posterior sclerites of the notum more yellowed; antennae
black, the proximal half of scape yellow; legs blackened; wings with a weakbrownish tinge; Sc1 ending a short distance beyond origin of Rs; R2+s+,
long; cell 1st M, closed; male hypopygium having much the structure of

SEPT., 1948]

ALEXANDER:CRANEFLIES

147

unicolor but the gonapophyses very reduced, appearing as small flattened
black blades.
Rostrum
Antenna

4 mm.;

about

MALE.-Length

narrowly

with

the

yellow

half

proximal

4.5 mm.

wing

the palpi

basally,

black;
segments
flagellar
long-oval,
Pronotumbrown.
Mesonotal
yellowish

scutumandprEesutum

the sides paler;scutellumandthe propleuraand mesepisternum

brown,
yellow;and mouthparts

the remainder
scape yellow,
withverticils.
Headelongatemore
postnotum
moreinfuscated.

black.

of

organ

dark

gray.
dark

of

chiefly

Pleurayellowed.
Halteresbrown

ish black. Legs with the coxmeyellow, the fore pair slightly more darkened;trochantersofremainder

infuscated;blackened.

legs

with

Wings

a weakbrownish tinge, the long-oval stigma slightly darker brown; veins dark brown.
Venation: Sc, ending a short distance beyond origin of Rs, Sc2 opposite
thisin direct

Bs

origin;withalignment

the usual

Rs,

basal

deflection

of

the

latter lacking; R2+,+4long, nearly twice vein R.; cell 1st Mf rectangular with
m-cu

at

near

one-third

thelength.

Abdomen with tergites brown, the sternites more yellowed; hypopygium
brownishMalewiththe styli muchas in unicolor
andyellow.
hypopygium
the phallosome
much
as in this species
in an importantbut differingwithregard, the length and shape of the gonapophyses;
appear
of
the

as

small

apicesblack

flattenedofthelateralbladesthatplatesofextendthein the present fly thesecaudadto theonly
in unicolor

phallosome;

level
the

apophyses appear as long blackened rods, much exceeding the aedeagus inlength, the tips acutely pointed. The outer portion of the phallosome, pre
sumed

to be

the

shows

edeagus,twonarrow

blackened

lines

or

sclerotizedstrips.
Habitat.-Panama.

Holotype, I, Cerro Pena, near El Valle, November 1946 (NoelL. H. Krauss).
The present fly is undoubtedly close to Gonomyia (G6onomyia)
unicolor Alexander (Mexico, Salvador, Guatemala) but differs in
the important genitalic character discussed above.
Gonomyia (Lipophleps) spiniterga, new species.
Size large (wing, male, over 5 mm.); mesonotum dark brown, the posteriorborder of scutellum broadly yellow; pleura with a whitish stripe; legs darkbrown; wings with a strong dusky tinge; Sc short, Sc1 ending some distance
beforeoriginof Rs;

male

about

MALE.-Lengthwithhypopygium

spicuous blackened spinous seta;bilobed.
5 mm.;

dististyle

wing

the

tergiteprovided

with

con

single, terminal in position,

5.2 mm.

Rostrum orange; palpi black. Antennae relatively long, blackened; flagel
lar segments elongate, especially the outer ones; verticils of the more proxi
malsegments

very

long.

Head

above

chieflydark

colored.

148

NEW YORK ENTOMOLOGICALSOCIETY[VOL.LVI

Pronotum and pretergites light yellow. Mesonotum dark brown, the sur
face pruinose, more heavily so behind; scutellum with posterior borderbroadly yellow. Pleura black, heavily pruinose, with a broad more whitish
longitudinal stripe extending from behind the fore coxae to the base of abdomen, slightly widened behind. Halteres with stem pale, knob dusky. Legs
with the coxa yellow, more or less darkened basally, the surface pruinose;
trochanters dusky; remainder of legs dark brown. Wings with a strong
dusky tinge, the prearcular and costal fields more whitened; stigma veryslightly darker, very diffuse; veins brown, paler in the brightened portions.
Venation: Sc short, Sc, ending a distance before origin of Rs only a little
less than the length of the latter; anterior branch of Rs curved strongly
cephalad, nearly parallel to R,+2; cell 1st M2 closed.
Abdominal tergites brown; sternites obscure yellow. Male hypopygiumwith the tergite conspicuously armed with blackened spinous setue, including
a marginal row of six or seven on either side, the more basal ones larger andstouter. Basistyle elongate, slender. Dististyle single, terminal in position,bilobed, the outer lobe a sinuous rod that terminates in a blackened spine,
the inner lobe fleshy, tipped with a single very strong fasciculate bristle; afew other setm on this lobe. Phallosome consisting essentially of a single longstraight spine, the tip blackened and acute; at base of spine more dilated
and with two smaller pale acute points.

Habitat.-Southeastern Brazil.
Holotype, A, Boracea, Sao Paulo, altitude 800 meters, August
1947 (John Lane).
In the strongly developed spinous armature of the tergite of
the male hypopygium, the present fly agrees most closely with
Gonomyia (Lipophleps) ctenophora Alexander and G. (L.) ju
quiana Alexander, differing in the large size and in all detailsof structure of the male hypopygium.