ciT-)-;,r

IReprinted from Jourl.ar, or rrre Nnrv yonKENToMor.ocrcer. Sogrnry, Vel. xxlI.
No. s, September,r9r4.l

A

REVISION
OF' THE AMERICAN
SPECIES OFTANYPREMNA OSTEN SACKEN AND MEGISTOCERA WTEDEMANN.
(TTPULIDE,DIPTERA.)1
By Crrenrpsp. ArpxeNopn.
Iru.lce,

N. Y,

This paper is presentedin order to compretethe American speciesof the Dolichopezini that are allied to Megistocera wiedemann.
These gbnera are Brachyprewno osten Sackenwhich has been taken
up by the author in an earlier paper (Journ. N. y. Ent. Soc., vol.
XX, p. zz5-236, rgrz), TanyprernnaOsten Sacken and.Megistocera
wiedemann which will be consideredin the present paper in theorder named. A key to the Dolichopezinegenera of the worrd isgiven in Psyche,vol. XIX, p. 64, rgrz.
I am indebtedto Mr. Frederick Knab for the loan of the material
in the United StatesNational Museum; to Mr. E. T. Cresson,
Jr., forthe specimensin the PhiladelphiaAcademy of Sciences;to Mr. C. W.
Johnson fot a Tanypremno taken on the Mann expedition to Brazil;
to Dr. F. H. Lutz for the material in the American Museum of
Natural History; and to Mr. John Thomas Lloyd for the speciesof
Tanypremna herein described as new.
1 contribution

from the Entomological Laboratory of cornell university,

5f

206

[vol' XXrr'
Jounnar- Nuw Yonr Enrolrolocrcer, Socrrtv.
TANYPREMNA

Osten Sacken.

1 8 8 6 . T a n yp r e m n a Oste n Sa cke n ; Bio l. Ce nt' A m., D i pt., vol ' r, p. r9(opilio) "rgtz. Stegasnconotus Endetlein; Zoo1. Jahrb., vo1. 32, pt' r, p, 1r(longissimus),t g t z . P e h lh e a En d e r le in ;1 . c., p ' t5 ( co lu m b iana).

The genus Tonypremna was erected in 1886 by Osten Sacken to
receive the then unique species,opili'o, of Guatemala. The following
year the same author descrlbed T. manicata from Brazil. In a recent
paper Dr. Enderlein has erected two new genera which must be considered synonymous with Tanyprernna and the two types make thethird and fourth American species. The Tipula longipes of Fabricius
is now known to be a member of this genus, while the new form
herein described as Tanyprenxna regina is the sixth species to be madeknown. The species most closely allied to the genotype, opilio, arecolumbiana which Enderlein made the type of a new genas, Pehlleea,
and the new species,regina. T. columbiano Enderlein has a strongsupernumerary cross-vein in ce1l M, this latter character being also
found in regina, which, moreover, possessesa considerable series ofSuch veins and spurs of veins in both of the basal cells. This character of sllpernumerary cross-veins is one which has been overemphasizedin the past in the formation of genera and it is doubtful
whether even subgeneric rank should be given to the majority of suchforms. The extreme plasticity of ,these characters is shown in suchgenera as Cladura, Gnophorn'yia, Tricyphona, and others, in whichthese supernumerary cross-veins may be present or absent in the twowings of a single specimen. In the various subgenera of Limnophilasuch as Ephelia, Iilioptera, Dicronophragttl,o, etc., which possess thesesupernumerary cross-veins in all specimens, it is known and appreciated that these groups are scarcely of subgeneric value. Considering the very close relationship existing between opilio, colum,bianoand. regina, and taking into account. the plasticity of the charactersused in their diagnosis, I arn unwilling to consider the forms as
representing more .than very well-defined species of this genus,
Tanypremna.
The species of Tmypremna appear to be quite uncommon and notoften picked up by collectors. This is well shown by the fact thateach species was founded upon a single specimen and scarcely adozen specimens are known to be in existence in the various museums.

,
t.

trl

Sept.,r9r4.l Ar-Bxenonn: TexvpnBrrr.re erco Mccrsrocena.
Cn.lnacmns

207

oF THE GBNus.

Frontal prolongation of the head very short and stout, about as
deep as long; the nasus long and prominent, clothed at the tip withlong hairs; palpi with the last segment longer than any of the others'
Antenne very short, the basal segments larger, the flagellar segments
oval, more elongated toward the end. Front very broad between theeyes with no protuberence.
Prothorax viewed from the side very narrow' the scutellum closelv
applied to the mesonotal prascuturn and the head, in turn, closeiyapplied to this. Mesonotal praescutumvery gibbous, partly or almost
entirely concealing the head when viewed from above. Halteres longand slender. Legs very long and slender, the tarsi especially being
excessively elongated.
Wings with vein Sc long, Sc, persistent at the wing-margin andquite close to R, at its tip; Rs short, usually arcuated but sometimes
straight though never so square at the origin as in Brachypremna;R, nsually distinct, oblique, rarely vertical as in Brachyfremna.
Second anal vein usually long and not ending close in the anal angleof the wings. ln longipes Fabricius, R, is swung proximad at its tip
so that it is very close to R, at the wing-margin; Rs is oblique,straight and second anal is rather short. T. colurnbiano has a strongsupernumerary cross-vein in cell M, and, T. regina"has this crossvein and spurs of many other in the basal cells .
Abdomen elongated, in the fernales of some species, excessively
long and slender.
Coloration.-Usually light yellow, the thoracic dorsum dark brown.
the pleurae yellow with brown transverse bands' Legs brown, thesegments having more or less white. Wings usually subhyaline withthe veins indistinctly seamed with darker, rarely the whole' disk
marbled with darker.
A Krv ro trrr AunnrceN Spscrrs or T.tr'rvpnrrtNeOsrnN Secrrn.
. ' ' . . -.... -. '2
beforeor at its tip.
r. Tibie with more or less'r'tihite
Tibia with the baseabruptly white but the remainderof the segmentdarkc o l o r ed'...'.4
z. First tarsal segmentbroadly white medially,dark at both ends; remaining
tarsal segmentsentirely white; lradiai sectorof the wings short, straight,
oblique,forming a V with the deflectionof Rr*rl. (Guiana,Brazil')
Tarsal segmentsone and two tipped with white. . .

,.t.".".0.t::::1t'::t::

208

JounNlL

Nrw

Yonr< Enrouor-ocrcll

Socrrtv'[vol' XXIr'

Sept.,rer4.l Ar-rxeupnn:

apical veins seamed with brown; [abdomen very
3. .Wings hyaline with the(Brazil')ofthe body of the female over 5o mm'1'
lengthlong, slender;
longissima Enderlein.
less elongate' thewings pale brown with a darker brown stigma; [bodyis broken andspecimenknown
tip of the abdomen of the type and only
to be 14estimateditwasdetermined;
accuratelybenottie length can(Brazil')segmentl'
tarsalthirclmm. by Osten Sacken; some brown on the
manicata Osten Sacken'
in cell M'
4. No supernumerary cross-vein(Guatemala' Venezuela')oPilio C)sten Sacken'" " " " ' "5
A supernumerary cross-vein in cell ll4'
limited, confined to the cord and the super5. Wings with the seaming
male r8'5 mm' (Colombia')
numerary cross-vein in ce1l M; wing of thecolumbiana Enderlein.
a1l the cells; wing of the malealmost
in
brown
with
marbledheavily
Wings'"'"'regi nan'sp', ( Co lo m b ia .)""^Fabricius'
tongiPes
Tanypremna
p' z5' ftzr Tipula longipes
Tipula longibes Fabricius; Svst' Antl"fiq
fizB Tipulo longipes Wiedemann;
p'
t,
vol'
4r'
37,
exot.,
Dipt.
Wiedemann;
Tipula longipes Macquart; Suite i'
Aussereur, zweifl. Ins,, vol. r, p.-43. fi34
'frans' Am' Ent' Soc vol'Hunter;tongibes
Tifuta
p.
rgoo
82.
'
Bufion, vol. r,
ttol' z' p' 293' r9r2
Dipt"
Cat'
26, p. 286. tgoz Tipula longipes Kertesz;
vol ' 3z' pt' r' p' r3' 19r3
S t r g a r * o n o t u s lo n g ip e s En d e r le in ; Z o o l' Ja h r b ''Mus" vol' 44' p' 487'
Nat'
Tanyprentno longipes Alexander; Proc' U' S'
Female,length z4 mm.;wing'

r2'6*v'8

mm';

abdomen' 2Q'r-2o'2rnm'

tn;m; tarsus r' r5'5 mm';
Fore leg, femur, 9.6 mm'; tibia, r2'4
tarsus 2 to J, 8.r mm'tarsus r' 16'4 rnm'i
Hind leg, femur, 12 rntn'; tibia, 13 mm';tarsus 2 to 5, 10.6 mm'
yellow' the nasus
Frontal prolongatioh of the head short' pale dull
lower part of the rostrum
rather long with a brush of hairs at the tip;
segments pale' theintermediate
two
the
with
dark, blackish; palpi
segment very long'nanxedlatterthecolored,dark
first and last being
the first flagellar
of
base
the
and
segments
Antenne short, the scapal
dark brownish
antennetheofremainder
segment pale, yellowish,
into brown'
passing
vertexonthewhite,
Fiorrt p"1" yellowish
bl;k.
the occiput dark.
paler on the sides'
Pronotal scutum broadly dark brown medially'
extending to the
color
this
medially
Mesonotal prescutum dark brown,in front of the
space
the
darkest,isextreme cephalic margin whlch
pale' almost
very
is
below
described
dark mark on the pesudosuture

TaNvpnnntue AND MEGISTocERA'

209

the
white; the prascutum with deep impressed lines extending frombrownishadark
suture;thetransvefse
tofovea
caudadpseudosutural
onto
black mark at the humeral angle of the sclerite extending ventradandscutellum
medially;
paler
littleabrown,dark
the pleurze; sclltum
pale,
Pleuresides.
thepaler
onlittlelatterapostnotum brown, theyellowish white, with a broad brown stripe extending from thehumeral angle of the prascutum ventrad and caudad across the mesopleura, splitting near the base of the fore coxa, ihe anterior fork
thebroad suffusing the sides of the mesosterlla and the base ofknob
themesocoxa. Halteres long and slender, the stem pale brownr
dark. Legs with the coxa paie except the mesocoxe as describedabove; fore legs with the trochanters having a decided green caste'
pale'
femora dark brown, the base pale, tibia with the base broadly
white
broad
ratherring,aapical
black
the
to
width
about equal inasubterminal band; metatarsus dark brown, the white medial band
white.
tarsi
oftheremainder
fing,
basal
dark
little broader than theHinrl legs with the femora brown, the tibia with the basal threefifths clark brown, the remainder snowy white excepting a broad darkbrown tip; metatarsus white except the broad brown base and apex;
remainder o{ tarsi pufe white. wings usually with a light browntinge, in some more nearly hyaline, and the costal celFbrown; stigma
dark brown, the cord more indistinctiy seamedwith brown' VenationR'*"
as in fig. r : Rs oblique, short and straight; the deflection ofa r c u a t e r l ,f or r n i n g a V w i th th e e n d o f th e se cto r ;R , sh o r t a n d sh o w ing a tendency to atrophy; basal deflection of R",o lacking'Abdominal tergites 2 to 4 brown with a broad white basal blotch
middle'on the sides and a broad white biotch on either side beyond thebrown
blackishnarrowlyI
sclerite
theofmarginlateralthe extremetheyellow'
dull
Sternitesbrown.
clark
uniformly
tergites
apicalovipositor reddish brown.
Distri,buti,on.-British Guiana, Upper Potaro River, July 17' rgrr'(Crampton.) Specimen, a female, in the collection of the American
Specimen a
Btazil, Manaos (Mann)'Museum of Natural llistory.
female in the collection of Mr. C. W. Johnson'
Tanypremna longissima Enderlein.
longissimusEnderleini Zool' Jahrb'' vol' 3z' pt' r'
1912, Stegasm.onotus
p. rr, r2.

2IO

Enroltor'ocrcer'
Jounr.ar- Nrw- Yonn

Socrorv'[vol' XXrr'

mm'
Female, length, 55 mm.; wir.rg, 22'5 mm'; abdomen' 48'5
mm'
rr.5
femur,
Fore leg,tarsus r' 22 rnm'1'2t
Middle leg, femur, 16 mm.; tibia, I8'5 mm';ro.75 rnrn.; 3, 6 mm.
r' 20 mrn':' 2' lo'5
Ilincl leg, femur, 19 mm.; tibia, 22 rnrn'; tarsus
mm . ; 3, 5. 5 m m .
brown' vertex
Head whitish; occiput for the most part blackish
the head scarcely
pale brownish. Prolongation of the frontal part of
the head' Palpi
of
lengthrerrainingth"
one-quarter as long".
short (z'25 mm'greenish, the last segment greyish black' Antenne
the tips of the very
iong; ,r..y thin and delicate, r3-segmented,pale,segments entirely.l"rrder flageliar segments grey, the last five flageilar
flagellar segment
each
of
end
the
at
greenish;grey, basal segments
broad;
Foreheadverticillate'somewhathairs,
i"ttt., long fine
tubercle lacking.
over theThorax gibbous, extending far forward anrl projecting
latter'
of
the
seen
be
is
to
nothing
above
fromhead so that viewed
narrowerafront;
in
seam
broad
a
brown,
Thoracic dorlsum clark
white; the dark
seam on the sides up to the rving-root yellowish
broad
moderatelyaasmargincephalic
brown color extends to the

i

i:
(,

h
$
r

&

stripe .Pr as c ut um wit has onr ewhat elev a t e d b l a c k i s h - b r o w n m e d i a l r
long' brown' lvhitish
line. Scutum brown, very 1eve1' Postnotumon the micldle whichcrossline
impressed
yellow on the sicles with anand from thisfront)
in(concave{orms an obtuse angle medially
Metabackward'
line
longitudinalangle sends out an impressed
whitishn.rediaily'
brown
greyishpl"rrr* greyish brown. Prothorax
dark brown above'elsewhere. Ab<lomen extremely long and slender'
one to three there
tergites
theofochre-yellow beneath, on the middle
a light spot on
seven
to
four
is a sharp pale crossline, on tergites
one and onescarcely
short,
each sidp of the middle; eighth segment
moderately
thewithsegment
half tines as long as broad; genitalpolished'
brown'
reddishelongated and rather powerful ovipositor
with agreen
greyishpalesmooth, anil 6 mm' long. Fore femora
thebrown'
blackishlegs
narrow brown ring'before the tip' Middle
apicalthealmosttip;
tibia with a very broad white ring before thethird tarsal segment
half of tarsal segments one and two white, thefemora reddish
Hind
entirely white, remaining segments broken'
with a broad
brownyellow, darker brown at the tip; tibie blackish

Sept,r9r4.l Alsxer,{nRn: Ter'Ivpnptr{Ne eNn Mtcrsrocnna'

2l],

and two blackishwhite ring before the end; tarsal segments one
remainingbrown with somewhat more than the apical half white;
with
Halterestarsal segments white' Claws small, reddish brown'
hyaline'Wings
the peclicel long, yellowish brown, the knob brown'
quarter of the wings
apical
the
on
veins
A11
brown.
dark
veins
and deflections ofseamecl with pale brown, all of the cross-veinsand ends closeveins brown and more broadly seamed' R, very shortin punctiform conto R, (on the lelt wing R, is quite lacking) ; Cu,
Membrane verytact with cell rst M"; Rs short, not longer than Rr*,'smooth but with greenish reflections only at the tip'
D istribwtion.-Bt azil' Espiritu Santo'
Translated from the original description'
Tanypremna manicata Osten Sacken''vol 3r'r887. Tanyprennannnicata Osten Sacken;Berl' Ent Zeitschr''
Pt' 2, P. 24o.
r 9o 2 . Ta n y f re t n n a

ma n i c a t a

K e rt e s z ;Cat' Dipt''vol' z' p' 265'

confluent
Yellowish brown, thoracic dorsum with three almost
yellow'pald
Antennebrown stripes; brown spots on the metanotum'
yellowish
abdomenyellow;tlarker towarcls the tip; halteres brownish
paler; a small
brown (tip broken). Legs clark brown, but femora
white rings
successive
white rini before the tip of the tibiae; threesWingsa palewithat the encl of tarsal jcrints one, two anrl three'b l o w n ish ti n g e ; sti g m a b r o w tr '(discoverer, Sellow) ; a single specimen in theI{abitat.*Brazil
is broken
Berlin Museum; sex uncertain, as the tip of the abdomen
r4 mm'
about'
bemust
off. The length, when the abdomen is entire,T h e t - r u m ,|r e r o fj o i n tso fth e a r .r te n n r i sce r ta i n l ym o r e th a n e l e ve n
(this number I counted in T. opilio)'
descripThe above is taken verbatim from osten Sacken's original
taken'
fartion; the type specimen seems to be the only one so
Tanypremna oPilio Osten Sacken'
Dipt', vol' r,
I886' Tanyprelnnaopilio osten Sacken; Biol. Cent. Amer.,p. t9, pL.r, f. z.
vol; 3o' pt'
1882. Tanylremna opiLio Osten Sacken; Berl' Ent' Zeitsch''
2, p, 16 4 .
tgo2.

Tany!>remna

opilioKertesz;

Cat'

Dipt''

p' 265''tot' z'2o'4 mm'Male, length, 23.4 mm'; wing, 18 mm'; abdomen'
24'2 tnm'
Fore leg, femur, r2.8 mrr.'; tibia, r5 3 mm'; metatarsus'
Hind leg, femur, r4.5 mm'; tibia, r5 mm'

212

Jounrer. Nrw

Yonr< Ewtonror-ocrcer- Socrrrv.

[vol' XXrr"

sept.'r9r4.l Ar.nxawnrn: TANypREMNA AND MBcrsrocBna.

data; the note by Champion who collected the type, tell us that theinsect lives in the humid forest regions of the mountains (5,ooo feet).

Female, length, z6.z mm.; wing, r8.3 mm.; abdomen, 22.2 mrn.
Hind leg, femur, r3.9 mm,; tibia, r3.8 mm.; metatarsus, r8.7 mm.
Frontal prolongation of the head yellow, the nasus long, clothedwith dark hairs; the palpi and lips dark brown. Antennre with thebasal segments yellow, flagellum black, antennre with thirteen segments (not eleven as given by Osten Sacken). Head light yellow
except a linear brown blotch on the vertex along the inner margin ofeach eye.
Pronotal scutum brownish black, this color produced ventrad onto
the propleure and including the anterior face of the pro-coxa.
Mesonotal prascutum deep chocolate brown, darkest in front, withthree very narrow darker lines, one median, the other two lateral. A

Tanypremna columbiana Enderlein.
tgtz .
r5, fi g. B .

Hind leg, femur, r5.25 mm.; tibia, r3 mm.
Head yellow; eyes very large, black, almost semicircular; occiput
somewhat infumed. Front narrow, half as broad as the diameterI

I

and traverses the mesopleure, ending on the anterior face of themesocoxa, scuttlm, scutellum and postnotum dark chocolatebrown, thelatter with a pale, narrow median line. Pleura light yellow with theblack with the extreme base rather broadly white, tarsi white. Wings
with a rather uniform light brown suffusion, stigma dark brown; theveins seamed with a light greyish brown. Venation as in fig. z.

yellowish at the base; tibia clark brown, the basal eighth yellowish
white; first tarsal segment blaok, the second yellowish white; the remaining segments as well as those of the fore and middle legs
broken off. Abdornen long, brown, the tip somewhat enlarged andAbdominai tergites brown, the lateral margins of the sclerites

and Barber), Purula, Yera Paz,; altitude, 5,ooo feet
(Cha mpio n) . Venez uela, Car iaquit o, J an. z z , l g t r ( S . B r o w n ) .
The specimen from Cacao is a female in the U. S. National
Museum collection. The Purula specimen is Osten Sacken's type.
The Venezuela material consists of a fine pair in the collection of
the American Entomological Society, Philadelphia.
This is the only species concerning r,vhich we have any ecologicaiof the eyes. Antenne very small and short, about r.75 mm. long,dark brown, the basal segments yellow. Palpi black, the apical segnent with the exception of the base brownish yellow. Nasus long,pubescent.atthe end and somewhat propped.
Thoracic dorsnm clark browu, sternites, pleur:e and coxee brightbrownish yellow; humeral angles of the praescutumbright brownish
yeliow; a similar spot before the wing-base. Before this latter arather broad brown band extends from the margin of the thoracic
dorsum through the middle of the mesopleurre,ending on the mesocoxe which are thus infumed on their outer face. The sides of theprothorax brown, fore coxe browned on their outer face. Legs withthe trochanters bright yellowish, hind femorr dark brown, palevertical brown bands as described above; sides of the postnotum andthe caudal face o{ the metacoxaebrownish black. IIalteres very long,du1l yellow, the knob brown. Legs with the coxae as described above,trochanters greenish yellow, femora brownish black, tibia brownish(Schwarz

P ehl l z ea c ohrmbi qna E nderl ei n:' Zoo1. J ahrb., v ol . 32, pt. r, p.

Mal e, l ength, 27 mm.; w i ng, r8.5 mm.

yellow patch on the sides of the sclerite in front, before the pseudosutural fovea; a dark brownish black stripe begins at the pseudosuture

rather broaclly black which color runs across the dorsum as a subapical band; extreme apices of the basal tergites paler; the terminal
tergites uniformly dark brown. Sternites dull yellow with an apicalannulus of black and in the terminal segments a medial band also;
the eighth segment black; hypopygium reddish.
23
Distribwti,on.-Guatemala,Cacao, A1ta, Vera Paz, April

zLB

'lrl

darkened, the incisures rather brightened. Halteres greyish black,pedicel very long, brownish yellow and with yellowish pubescence.
Wings rather narrow at the base for a rather long space, very
narrow;bright brownish, a brown spot at the base of the raclial fork,a rounded blackish brown spot at the stigma, the caudal veins of cellrst L(e, the basal deflection of Cu, seamed with brown, the supernumerary cross-vein between M and Cu broadly seamed with brown;
proxima of the base of M a small hyaline spot. Membrane strongly
red to green iridescent.
Distributi'on.-Colombia, Hacienda Pehlkea.
Translated from the original description.
Enderlein's figure shows a typical Tanypremna with the exceptionthat the tip of R, is omitted in the figure; whether it is also lacking

274

Jounrvar, Nrw

Yonx ENroltot.ocrcar- Socrsrv.

[vol' XXIr'in the type is another question but if such is the case its disappearance
may be accounted for by atrophy. In any case there is a considerable error in Enderlein's interpretation of the veins in the radial fieldof the wing; the vein labelled R, should, of cowse, be R,, the part
spoken of as the radial cross-vein is the basal deflection of R" and thepart called Rr*, is R, alone. The wing is shown in fig. 3, taken from
Encierlein's original clescriptionof the species.
Tanypremnareginanew species.
wing heavily marked with
Large species, wing of the male, 22 171171.;brown; superntlmerary cross-veins in the basal cells of the wing; tibia whiteat the base.
M a l e , l e n g t h , z8 .z m n .; win g , zz m m .; a b d o r r en, 23.5 rnm.
F o r e l e g , f em tr , r z.7 m m .
H i n d l e g , f e m u r , r 5 .7 m m .; tib ia , r 5 .r m r n .; iletatarsus, 24.8 n7m.
Frontal prolongation of the head very short, the nasus long and slender,d a r k b r o l v n ; s i de s o f th e r o str u m m o r e ye llo wish ; pal pi dark brow ni sh bl ack.
Antennie with the two basal segments light yellow, the flagellurn broken.
Front light sulphttr-yellow; vertex and occipttt dark brou'tl, paler along thei n n e r m a r g i n o { th e e ye s a n d with a n a r r o w m e d ia n l i ne'
Pronotum with the scutum narrow and high, projecting dorsad as a sharpc o l l a r , t h e s c u tu n r d a r k b r o wn , r e r n a in d e r o f th e pronotum yel l ow Mesonotal prascutum light yellowish brown with fonr darker brown stripes, them i d d l e p a i r b e i n g lo n g e st, n a r r o we d in fr o n t a n d behi nd, very dark btow ni shb l a c k o n t h e e x tr e m e ce p h a lic m a r g in ; la te r a l str ipes shorter, begi nni ng at thepseudosutural fovee and running caudad to the sutttre, connected with the
middle stripes except behind; sctltun and scutellunl broken I postnotum darl<brown, more yellowish medially behind. Pleura yellowish with a green caste,a s m a l l o v a l b r o wn ish b la ck sp o t b e h in d th e h e a d at the end of the pronotals c u t u m ; a b r o ad e r d a r k b r o wn m a r k r u n n in g fr o m the base of the w i ng vent r a d t o t h e b a se o f th e m e so co xa wh o se p o ste r ior face i t suffuses. I{al teres
1 o n g ,s l e n d e r , g r e e n in co lo r . L e g s with th e co xa l i ght brow ni sh yel l ow except
t h e d a r k m a r k o n th e m e so co xa d e scr ib e d a b o ve ; t rochanters greeni shl femorab r o w n i s h y e l l o w, th e tip b r o a d ly d a r k b r o wn ; tibi e (onl v the posteri or pai r
remaining) with the extreme base white and having a decided green caste,
remainder of the tibia dark brown; the first tarsal segment pale, almost
whitish, basally soon darkening to a light brown that includes the remainderof the tarsi. Wings subhyaline in the radial and rnedial cells, cell C brighter,rnore yellowish, the anal cells greyish brown, a row of brown blotches in cells
R and. M, each one surrounding the spur of a cross-vein, there being about
five in cell R and three in cell M; the cord is seamed with darker brown as
are also most of the veins and deflections of veins; stigmal spot a little
darker brown. Venation as in figure 4; the presence o{ a supernumerarycross-vein in the basal cells is a feature that, in this subfamily of craneflies, occurs only in the Dolichopezini; the numerotls spurs of cross-veins in
the basal ce1ls is a very remarkable feature,

Sept.,rer4.l

Arnxawonn,:.T.Lxvpnrlrwe

AND \4EGrsrocERA'

215

narrowly
Abdominai tergites reddish brown, the apex of each segment
margin of thepaler, greenish white, the apical tergites darker brown; lateralyellow' the extreme apices oftergites narrowly dark brown. Sternites lightbrowr'
the sternites darker, the apical sternites unifornlly darker

Distribution.-Colombia, Cordillera Central'north o{Holotype, female, near LL Vega which is about I2 miles
(John
feet'
of
abotlt7,ooo
Almaguer, March 6, r9rz, at an altitude
Thomas Lloyd, coll.)
T hi ssp e ci e si sp a r to fth e L l o yd co l l e cti o n o fAn d e a n i n se cts;th epapercraneflies of tbis collection have been discussed in an earlier
sp
e
ci
m
e
n was
Th
i
s
r
9
r
3
)
'
( J o u rn . N . Y. En t. So c., vo l .2 r , Se p t',
butterflieswithpapersnot included in that article because it was inand so was overlooked until very recently'
T lr e r e ca n b e b u tl i ttl e c]o u b tb u tth a tth i sfl yi sq u i te g r e e n i sh i ntints on theits living state as shown by the presenceof decided green
thisstlggest
genus
halteres, legs and thorax. Other species of this
nrost
the
far
by
same condition to a mtlch less degree' This form isthat thebeautiful of al1 the species described and it is believed
nunlerousofthespecific name is not inappropriate' The presence
remarkable.
spurs of veins in the basal ce1ls of the wings is {uitecase' I
Since these spurs are surroundeciby dark malkings in ever'r
NeedhamI)r'believe these remnants to be constant or nearlyosorandsarlle opinion'
and Dr. Johannsen who examined the wing, are of theat first
This presence of a dark marking about a vein o{ spur seems
characterglance to be a trivial character but it is my belief that it is a
markdarkis,thatlaid,-thatbemuststressconsiderable
ipon which
veins and th(ings rvhen present on a wing nearly always surroundp r e s e n ce o fa b r o w n b l o tch i r r a ce l l m a yw e l l i r r d i ca te th e p o si ti o n o n ct
occupie(lbY a vein.
MEGISTOCERAWiedemann.
Wiedemann;Dipt' Exot', p' 4t'
t8zt. Macleistocero
t8 z8 .M e g i 'sto ce r a 'Wi e d e m a L r n ;Au sse r e u r 'zw e i fl 'In s''vo l 'I'p '5 5 'group of flie!
The genus Megistocera contains a small restricted
speciesirhaving a tropicopolitan distribution' There are about threet h e o i d Wo r l d a n d th e se a fe r e m a r ka b l e i n th e p o sse ssi o n o fe n o r 'singltmously elongated antenn& in the male sex so far as known; the
New World form has short antenne in both sexes'

Sept.'r9r4.lAr,rxeronn: Texypngrltr.raaNo Mnctsrocone.

218

Jounrver, Nnw Yonr Exrorr,ror,ocrcer,Socrury. tvol. XXrI..

Cnenactnns oF TrrE Gsxus.
Frontal prolongation of the head rather long, the nasus prominent,with a bunch of long hairs at the tip, Terminal segment of thepalpus elongate,whiplash-like. Antenne rather long, at least as longas the head, the scapal segments enlarged, the flagellar segments
six in number, elongate-cylindrical; in the New World species theantenn€ are about as long as the head, in the males of the old world
fornrs (this sex of M. bicauila speiser of Africa not yet described)
the antennze are ehormously elongated. Eyes with delicate ommatidia; the vertex rather approxirnated between the eyes.
Pronotum not produced dorsad into a narrow plate as.in Tantl_
Pretnna. Mesonotal prascutum not exceedingly gibbous or projecting
over the head. Halteres rather long, the knob small. Legs very longand slender, especially the tarsi. 'wings with the cross-vein m-cu
present and long; basal deflection of R,*, assuming a position thatisnearly vertical or perpe'dicular. The old world
fuscana wiedemann has a venation that is quite similar to that of our American
form except that the cell rst M, is much smaller and cell znd. A verymuch broader.
Abdomen rather short, the male hypopygium simple; the valvesof the ovipositor powerful, the lower valve arcuated basalrv. trre rower
valves appressed against the upper pair.
Megistoceralongipennis Macquart.
1838. Tifula longipennisMacqaart; Dipt. Exot., vol. r, pt. r, p.
57, pl.
5, fig. r.
1878. ?Longurio longipennis Osten Sacken; Cat. Dipt. N. Am., Ed.
2, p. 37,
i885. Tifula longipezris R6der; Stett, Ent. Zeit.,vol.46, p.
33g,
1885. Tipula tenuis v, d. Wulp; Notes Leyden Museum,vo1. p.
7, 7,
1885. Tipula tenuis v. tl. Wulp; Tijdschr, voor Entom., vol. _, p. g5,
pl: 4, fig. -.
1886. MegistoceralongipennisOsten Sacken; Berl. Ent. Zeit,, vol,3o,p. i6 r.
igo2. MegistoceralongipennisKerresz; Cat. Dipt., vol, z, p. 264.
1902. Tipula tenuis Kertesz; L c., p. 3o9.
rgo7, ?Ivfegistocera
tenuis Needham; Rept. Soc. Ent. N. y.,. p. zrz.
r9o9. MegistoceralongipennisJohnson; proc. Bost. Soc, Nat. Hist.. vol.
34, p. 123.

Male,length,ro mm.; wing, 15.6mm.
Middle leg, femur, rr.2 mm,; tibia, ro.6 mm.; metatarsusabout
z6 mm.

21\

Female,length, r4.3 to.r5 mm.; wing, 16 to 16.4mrn.
Fore leg, femur, 7.7 lr,m.; tibia, 8.6 mm.; metatarsus,8.r mm.
Middle leg, femur, rr.S mm.; tibia, ro.6 mm.; metatarsus,aboul
24 mrn.
Hind ieg, femur, ro.5 mm.; tibia, 8.8 mm.; metatarsus,aboul
25 mm.
Frontal prolongationof the head short, light brown, the nasuspalt
with the apical brush of hairs dark, palpi black. Antenne with thr
basal segmentslight yellow, the flageilar segmentsbrownish black
Front dull yellowish brown, the vertex and occiput grey.
Mesonoturnbrownish grey, the lateral margin of the prascutun
paler grey bordered interiorly by a narrow blackish line which is
waved at the pseudosuture;scutum brownish grey with a brown
blotch on each lobe; scutellum pale brown, on the sides dark brownand more shiny; postnoturn light brown, more fuscous laterally.
Pleura pale yellowish brown with a whitish bloom; a rounded shiny
blaok spot at the dorso-cephalic angle of the mesoepisternum
Halteres pale brown, the knob dark brown. Legs with the coxe pale,the outer faces somewhat browned. Wings whitish, veins brown;
stigma conspicuous,oval, brown. Venation as in fig. 5.
Abdominal tergites dark brown, the extreme lateral margins
broadly pale, the dorsum variegated with paler, the basal segments
being paler medially, the apical four or five segmentswith the palespotsin two rows one on either side of the middle line; sternitespale.
Distribution.-Florida, Lake Okeechobee;report by Mr. C. W.
Johnsonfrorn the Mus. Com. Zool., I have examinedthis same specimen on two different occasionsand there can be no doubt as to theidentity. Little River; Nov. 3o, tgl2, one male found in a spider's
web by Mr. Knab. Cuba, Macquart's type. Porto Rico (Rtider),Trinidad, June 13. Two males, three fem,ales,taken by August
Busck; U. S. National Museum. British Guiana, Bartica, Dec. g,
rgr2 to Jan. r8, r9r3. Mallali on the Demerara R., Mar. 20, rgr1
(Parish); col.lectionof the author. Dutch Guiana, Van der Wulp's
type of tenui.s. Brazil, female from Ceara in Museu Rocha. Pataguay, Piedro Bay. April, H. H. Smith, collector; part of the Williston collection in the American Museum of Natural History .in New
York City.

218

Ertoraor'octcer' Socrrrv'
Jounner- Nrw Yonr
ExPlexluox

or Plers

tvol' xxil'

V'

Fabricius'Fig. r. Wing of Tanypremtto longiSes
Qstea Sacken'
oqilioTan!4remna
of
Wing
,.
;i;
Enderlein
Fig. r. Wirg ot r"'iip'"**a"olumb"onan' sp'regina
TongptennaofWing
Fig. +.
longilennis Mgcquart'elit. Wf"s ol Megistocera

Vol. XXIL

Journ. N. Y. Ent. Soc,
(after Enderlein)'

Tipulide.
r l.

ii

PI' V.