!r7q
.
+
1.,

7Y8

J. Ent. (B) q (t), pp. r-ro, r5 figs. 1973

New or little-known speciesof exotic Tipulidae (Diptera). XIX
CHA RLE S P . A LE X A NDE R
Unio ersity of Massachusetts, Amher st, M assachusetts,U. S. A.

SYNOPSIS

Four new species of.Antocha, one new speciesof Gonornyia, two new speciesof Trentepohlia and one new species of Erioptera are described from stationsin Sikkim and Assam. Additional illustrations of previously described
species of Antocha and Trmtepohlia are included.
The preceding part in this series of papers was published in r97r (J. Ent. (B) +o : 1637z). Lt this time I am describing further novelties from various stations in Sikkim andAssam, the materials having been collected by Dr Fernand Schmid in 1959, 196o and
196r. In addition to the new species, I have provided figures for a few further species
that had not been shown previously, mostly based on materials taken by Dr Schmid andby Mr P. Susai Nathan of Coimbatore, South India. I again express my deep thanks to theabove for the privilege of retaining the materials in my personal collection of these flies.
Antocha(Antocha)decussata sp. n.

General coloration of thoracic dorsum brown, pleura yellow; antennae short, black,flagellar segments short-oval; legs dark brown; wings light grey, veins R2, R213 andR112 subequal; male hypopygium with apex of outer dististyle shallowly bifid; gonapophyses very long and slender, about twice the length of the aedeagus,decussate.
Male. Length, about 4.5 mm.; wing, 5.5 mm.; antenna, about o.9 mm.
Rostrum and palpi black. Antennae black, short; flagellar segments short-oval,
subequal in length to the longest verticils. Head medium brown, extensively light grey
pruinose on front and orbits; anterior vertex very broad, about twice the exposed widthof eye.
Pronotal scutum light brown, scutellum and pretergites light yellow. Mesonotal
praescutum uniformly brown, very sparsely pruinose, lateral borders yellow; scutum
and scutellum brownish-grey, parascutella light yellow; postnotum light brown, internotal suture yellowed. Pleura virtually uniformly yellow, sternopleurite vaguely darker;
dorsopleural membrane clearer yellow. Halteres with stem yellow, apex of knob brown.
Legs with coxae and trochanters yellow; remainder of legs dark brown, femoral bases
narrowly paler; claw of male with a single powerful nearly basal spine. Wings light grey,
extreme base more yellowed; veins brown, relatively distinct; costal fringe short.
Macrotrichiu ot *or. than outer third of distal section of Ra15. Venation: R2"subequal
to R112 and R213, some distance before level of r-m: m-cu atfork of M.
Abdomen brown. Male hypopygium (fig. t) with outer dististyle (d) relatively slender, strongly curved at near mid-length, apex bifid into two strong spine-like points.

z

C.P.Alexander

Phallosome(2) with gonapophyses(g) very long, slender, decussate,narrowed very
gradually into a long slender spine, lower margin of its outer fourth appearing as anarrow hyalineflange.Aedeagusslender,relatively short, lessthan one-halfgonapophysis.
HolotypeJ, INore: Bakkim, Sikkim, 689o feet, tz.iv.rg5g (F.Schmid).
Only a few regionalmembersof the genushave the apex of the outer dististyle of the
malehypopygium shallowly bifid as in the presentfly, theseincluding lztocha (Antocha)
bifida Alexander, A. (A.) dafla Alexander and A. (4.) turkestanicade Meijere. The
specieshere describedis most readily told by the unusually long sword-like gonapophyses,asdescribedand figured.
Antocha (Antocha) globulosa sp. n.
Size small (wing of male about 4'5 mm.); generalcolorationof mesonotumlightgrey,
praescutumwith three light brown stripes; antennaeshort, intermediate flagellar segmentsvirtually globular; legsbrown; wings pale grey, veinsvery indistinct, Rs very long,subequalto its anterior branch, vein R2 in transversealignment with r-m; male hypopygium with a concentration of strong black setae at outer end of basistyle; outerdististyle narrowed into a slender spine, apex subacute; outer gonapophysisa long
slender pale blade, narrowed very gradually into a spine.
Malz. Lenglh, about 5 mm.; wing, 4'5 mm.; antenna,about o'8 mm.
Rostrum and palpi light brown; terminal segmentof latter aboutone-third longer thanthe penultimate.Antennae(fig. z) short; scapebrownish-yellow,remainderdark brown;intermediate flagellar segmentsvirtually globular, outer ones slightly longer, terminal
segmentabout one-half longer than the penultimate; verticils short. Head dark brown.
Pronotum with a narrow brown central stripe, sides brownish-yellow. Mesonotal
praescutum light grey with three pale brown stripes, central one broader, darker atanterior end, humeral and lateral borders broadly yellow; posterior scleritesof notum
light grey, scutal lobes slightly more darkened.Pleura yellowed, vaguely more pruinose
behind. Haltereswith stem yellow, knob light brown. Legs with coxaeand trochantersbrownish-yellow,remainder brown; claws long and slenderwith a single elongatebasal
spine. Wings pale grey, basenarrowly brightened; veins very pale brown, difficult tosee in balsam slide mount. Macrotrichia on outer third to half of distal sections ofveins Raa5and M*2. Venation: Rs very long and straight, subequal to the anterior
branch or nearly twice R; vein R2 very faint, in nearly transversealignment withr-m;
Rz+aand basal section of Raa5subequal; m-cu shortly before f.ork of.M.
Abdomen with proximal tergites medium brown, outer four segmentsdarker brown.
Male hypopygrum (fig. 3) with ninth tergite narrow, transverse,posterior border nearlytruncate; vestiture pale and inconspicuous, some lateral setae elongate. Basistyle (b)with a concentrationof strong black setaeon outer end. Dististyles (d) closelyapplied,
subequalin length; outer style broadest beyond base,thence narrowed into a slenderspine, apex subacute; inner style stouter, with numerous setae,apex obtuse. Interbase(r) with the free outer part reduced to subatrophied.Outer gonapophysis(g) appearing
as a long slender pale blade, narrowed very gradually into an acute spine. Aedeagus
weaklycoiledat apex.
Holotype $, Ixore: Domkho, Kameng, North East Frontier Agency, Assam, 65oo
feet, 8. vi. ry6t (F.Schmid).
In hypopygial structure the presentfly is closestto Antocha (Antocha)scutellasp.n.,Exotic Tipulidae

3
differs
in certain characters,including the normal setaeof the basisryleand thewhichobtuse apex of the outer dististyle. The antennalflagellar segmentsin scuteilaare oval
not subglobularasin the presenifly.
Antocha (Antocha) scutella sp. n.
Mesonotum grey, praescutumwith three darker greyish-brown stripes,in casespraescut]lT r-norebrightened on sides; rostrum yellow; legs pale brown, outer tarsal segmentsslightly..darker; wings milky white, stigma lacking;-male hypopygium with ip"x ofo]lt_er_dististyleobtuse; outer gonapophysisdistinctive, .rtt"*"ity-tong and sLnder,
slightly expandednear outer end, tlrenceextendedinto a long straig-ht.pii.; phallosome
with central area long-subrectangular,at mid-portion of apex witl a smali
iale brownshield-shapedstructure at termination of aedeagus.
Male. Length, about 4-4.S
1nm.; wing, 5-6 mm.; antenna,about o.75_o.g5mm.
Rostrum relatively_long, yellow; palpr dark brown. Antennae short,"as slio*r, bymeasurements,black; flagellar segmentsoval, the outer two longer. Head dark grey.
Pronotum yellowed, scutum darker medially. Mesonotum!r.y, pru.."utum withthree slightly darker greyish-brown stripes, the lateral pair tior. ob."rr.., cro"rirr!
suture to include scutal lobes;,in certain specimensthat are consideredto be conspecifi{
sides of Praescutummore yellowed; posterior scleritesof notum grey, parascutellaandpleurotergitepaler. Pleurayellowed,patternedwith brownish-grey,esp".i"lty the sternopleurite. Haltereswith stempale yellow, knob weakly infuscatel. i-egs with co"u. y.llo*,basesof fore and middle pairs darkerl remainder of t"g. pale brown, outer tarsal segi
t1"1t. slightly darker; claw of male with a single tong s"b-basal spine. Wings
milfy
white, especiallyat base; no stigmal darkening; veins,jc and R
fale brownish'-yellow,
darker,_especiallyoutwardly. Macrotrichia on outer haif of Ra15and "moresparselyon
M*2. venation: R2 opposite or shortly beforelevel of r-m; m-cu before fork of fr, jn
casesnearlyto one-halfits length.
Abdomen dark brown, hypopygium slightly parer. Male hypopygium (fig.
withninth tergite (l) transverse,posterior bordei very shallowly .t""tgi""t-., urrte.i]o,4)i11urgilwidely thickened. Outer dististyle (d) obtuse at apex, innlr style*slightly longer.
O"i"igonapophysis(g) pale, unusually long and slendir, slightly .*pattaia near iuterend,thence extendedinto a long straight spine. Central area-of(p)
Iong-subrectl
ih"tL*orn"a.ngglar.in outline, apex nearly truncite, at its mid-portion with a .*utt pit" brownshield-shapedstructure at termination of the slightly ,ir.ou" aedeagus.
Holotype6f, INore: Hkayam Boum, Manipu-r,Arr"-, g5oo feet, zt.vi.196o. para_
topotypesJQ, with holotyp-e.paratypes. gQ, zema, sikliim, ggoofeet,ri.vi.r959;
r J, Sutol, Pauri Garhwal,Kumaon,
TzSofeet, 18. viii. r95g; r i, ialung Drong, f<ii_eng, North East Frontier Agency,
Tooofeet, lz.v.196r. (Alt i.Srlrma\(Antocha)scutelh is most readily told from other related regional speciesby,- _n!,*?
hypopygial
characters,including the very long outer gonapophysesand the central
phallosomic structure, includinf the smail striJa-snapi a..u'thut hassuggestedthe
specificname.
Antocha (Antocha) scutifera sp. n.
Mesonotal praescutumwith a brown central stripe and nearly confluent greylaterals,
outer margins broadly buffy yellow, concolorouswith the pleura, ventral
rl"rrroplerrrite

+

C.P.Alexander

weakly darkened; antennaewith scape brownish-yellow, remaining segmentsblack;notouter end of halteres darkened; legs brownish-yellow; wings whitened, stigma
darkbrown,
abdomenindicated, veins delicate,brown; m-cu shottly before fork of M;
basal sternites weakly yellowed; male hypopygium with outer gonapophysisa yellow
blade with up"* rrrr.qrr"lly bispinous; apex of central phallosomic masswith incurved
armsto form a small shield-shapedstructure'
Male. Lengrh about 5.8-6 mm., wing, 7-7' 5 mm.; antenna'about r'5-r'6 mm.
Female.Length, aboutT mm.; wing, 8 mm'
Rostrum ligilt yellow; palpi black. Antennae with scapebrownish-yellow,remainder
black; basal fligelia, segrrre.ttslong-oval, the outer ones shorter; normal vestiture pale
but abundant and conspicuous,verticils short. Head greyish-brown.
Pronotum yellowed. Mesonotal praescutumwith a brown central stripe and nearly
confluent grey laterals, humeral and lateral margins broadly buffy yellow; posteriorsclerites oi rrot r- grey, scutellum yellowed posteriorly. Pleura buffy yellow, ventral
sternopleuriteslightl-yiarker. Halteris with stem light yellow, outer half of knob brown'I,egs with fore co*ue more or less dark brown, remaining coxae and all trochantersouter tarsal segmentsslightly darker. Ii"g'V"tt*t remainder of legs brownish-yellow,
lighti"hiterr.d, especiallythe prearcular ield, stigma not differentiated; veins delicate,transverse
approximate
R2
into darker brown, extremebasalveins whitened. Venation:
itsalignment with r-m; m-cu shotlly before fork of M, in casesto about two-thirds
length.
in male dark brown, basal sternitesweakly yellowed, hypopygium brownibdo-.r,
ish-yellow; in female sternitesalmost uniformly yellow. Male-hypopygiuT (fig. 5) with
fo1
ceniral puit of posterior border of ninth tergite (t) very shallowly emarginate!o
to
the
fifth
narrowedoutertheoutwardly,
1o* lobes.buter dististyle (d) darkened
theblade,'.ry
yellow
narrowalonggonapophysis
obtl.e tip. Phallosome(?) rvitft in. outer
to
arms
with
incurved
structurephallosomic
tip unequatly bispinou{ apex of central
form a small shield-shaPedarean\laoamdron association'
Hototype 6, INnti: Tsomgo, Sikkim, r2,5oo feet,
F'Schmid)'
(All
d ?,
z6.viii. tg5g. AllotopotypeQ. Paratopotypes,'.nOther iegional me'ribers of the ginus that have the outer gonapophyses-ofthe
hypopygium forked at the apex include Antocha (Antocha) I'ibanoticaLackschewitz,
North Libanon to North Afghanistan; A. (A.) quafuifttrcaAlexander,Assam,and'A' (A')
spiralis Alexander, Kumaon eastward into western China. All such speciesare most
readily told by details of hypopygial structure'
Antocha (Antocha)studiosaAlexander
Alexander,rg5t, Ann. Mag. nat. Hist.$z) 4 : ro98-ro99_.
Antocha(Antocha)stud.iosa
Types fro- Singara,Nilgiri Hills, South India,34oo feet,3-29.v.ry48 (P.Susai'
Nathan). Male hYPoPYgium(fig. 6).
Gonomyia (Protogonomyia) cobelura sp. n.
Sizelarge (wing of male about 8 mm.; female Io mm.); generalcolorationof body anddui black; male hypopygium with outer dististyle an elongatesinuousspine,"pp.rau'g".
intermed"iatestyle with t*o snbiquai branchesat near two-thirds the length, inner stylewith distal half a long-ovalblade.

Exotic Tipulidaer€41:dffF€o.F"d"<O@coFigs. r-6. (t, :-6). Male.hypopygiumof: (t) Antocha(Antocha)decussata
sp. n.; (3) l.
(4.) globulosa
sp. n.; (4) G) A. @.) scutellasp. n.; (5) A. (A.) scutiferasp. n.; (6) A. (A.)
studiosaAlexander. (z) Male antenna of A,(A.)globulosa Alexander a, aedeagus;b, basistyle;d, dististyles;g, gonapophysis;/, interbase;p, phallosome;t, ninth tergite.

MaIe.Length, about8 mm.; wing,8 mm.
Femalc. Length, aboutT mm.;wing, ro mm.
[Iead dull black. Antennae black; proximal four flagellar segments long-oval, without
setae on ventral face, outer segments progressively lengthened, shorter than their longest
verticils.
Thorax uniformly dull black. Halteres and legs black. Wings strongly infuscated,

6

C.P.Alexanderprearcular field, cells C and ,Sc and stigma more darkened; veins brownish-black.
Venation: Rz+a+qabout twice the basal section of R5; cell m2 open by atrophy of basal
section of.rna; m-cu closeto fork of M.
Abdomen black. Male hypopygium (fiS. r+) with apicallobe of basistyle(6) short andstout, with long setae.Dististyles (d) as shown, outei style an elongatesinuous spine,
narrowed very gradually to the acutetip; intermediatestyle with ttvo subequalbranches
at near two-thirds the style, axial one somewhatstouter, tip more obtuse, lower marginof outer two-thirds of style with abundant delicatesetae;inner style darkened,narrow-est
at near mid-length, distal half expandedinto a long-ovalblade.
Holotype J, Ilore: Talung Dzong, Kameng, North East Frontier Agency, Assam,
78oo feet, 6.vi.196r. (F.schmid). Altotopotypi Q, pinned with holotyp.. t" narure,associatedwith the very similar Gonomyia(Protogonomyia)
gtabristylaAlexander lg67a).
In the genusGonomyiathere are severallarge-sizedspeciesin the subgenui nongonomyiacomparableto the present fly, all most readily distinguished by hypopygial
structure, particularly the dististyles.In the referencecited above I had describedand
figured the hypopygium of fourteen such speciesin the presentsubgenus.
TrmtEohlia (Mongoma)amphiniphaAlexander
Trmtepohlia(Mongoma)amphiniphaAlexander,rg67bJ. N.Y. ent. Soc. :24.
75
Type, Lingtham, Sikkim, 46oofeet, z.ix. rg5g (F.Schmid).Yenation(hg. 7).
Trentepohlia (Mongoma) eurystigma sp. n.
Mesonotum chestnut brown, pleura yellow; halteres brown, base of stem yellowed;
legsyellow, femoral tips narrowly brownish-black,no modified spinoid setaeon femora;
wings whitish subhyaline,prearcular and costal fields clear light yellow, including theveins, remaining veins dark brown; a restricted dark brown putt"tn, including the unusually large stigma and narrow seamsover certain veins especiallyvein Cu in cell M,
wing tip paler brown; a short elementR213,vein Rs subvertical,cell R2 at margin very
narrow; abdominaltergitesdark brown, sterniteslight yellow, posteriorbordersnarrowly
blackened,outer three segmentsblack.
Male.Length, about rr mm.; wing,9.3 mm.;antenna,abouta.r mm.
Rostrum light brown, palpi dark brown. Antennae with scapeand pedicel medium
brown, pedicel slightly paler; flagellum brown, segmentslong-cylindrical, exceedingthe
verticils.Head obscurebrownish-yellow:anteriorvertexnarrow.carinate.
Cervical region and pronotum yellow. Mesonotal praescutumchestnut,with indications of darker lateral stripes; posterior scleritesof notum darker chestnutbrown, restrictedly more yellowed medially. Pleura yellow. Halteres brown, base of stem narrowly
yellowed.I 'egswith coxaeand trochantersyellow; femorayellow,tips narrowly brownishblack, including about the outer fifteenth of segment; femoral baseswithout spinoid
setaesuch as occur in hempi; tibiae and tarsi brownish-yellow. Wings (fig. 8) whitish
subhyaline,prearcularand costalfields clear light yellow; stigma darklrown, unusuallylarge, filling cell R2 except for the costal fifth, reachingRs behind and including partsof adjoining cells Sc1and R1; narrow dark brown seamsat origin of Rs and over entire
length of vein Cz in cell M; wing apex extensivelypaler brown, extending from cellRs to znd M2; veins dark brown, clear light yellow in the prearcular and costalfields.

Exotic Tipulidae

7

Abundant costal setae,very sparseand scatteredon outer ends of vein R1 and distal
sectionof R5. Venation: a short elementRz+g,vein R3 subvertical; cell R2 at costaveryn^rrow, about one-twelfth cell R3 or less.
Abdominal tergites dark brown, posterior borders narrowly black; sternites yellow,
posteriormarginsand all of outer three segmentsblack.
HolotypeJ, INnre: Nizong, Kameng, North East Frontier Agency, Assam, Coofeet,z7 .vi. rg6t (F.Schmid).
Trentepohlia(Mongoma) ewystigma is generally similar to certain other regional
speciesincluding T.(M.)butlui Alexander, T.(M.)flaaicosta Edwads and f.(M.)
kernpi(Brunetti), all with the darkenedwing stigma small and with the venationaldetails
distinct. The most similar of theseis butleri, with the stigmal areamuch smaller and withvein R3 oblique, cell Rz ai costabeing about one-fourth to one-fifth as extensiveas cellR3. In butleri the wing veins posterior to R1 are more extensivelyyellowed, the darkened
band at mid-width of wing narrow. f.(M.) hempi difrers in the coloration of the abdomen, halteresand wings, in the presenceof spinules on basesof the fore and middle
femora, and in the venation, with vein R2 before the radial fork to leavea short element
Re+a.
Attention is called to Trentepohlia(Plesiomongoma)oenosa(Brunetti), type of thesubgenus,which in most features is very similar to speciesof the presentgroup in the
typical subgenus,differing almost solely in the principal subgenericcharacterof loss ofvein Rs by atrophy. I am showing the venation of this speciesfor comparison(fig. rr,
with enlargementof the radial field).

Trentepohlia (Mongoma) flavoides sp. n.
Allied to flaaa; size very large (in female,body and wings eachabout ro **.); mesonotal praescutumyellow, patterned with orange; legs yellow, apicesof femora and bases
and tips of tibiae narrowly blackened;wings deep yellow, veins darker yellow; abdomenlight yellow, tergites medially broadly brown; ovipositor with cerci long and slender.
Female.Length, about romm.; wing, romm.; antenna,about z.zmm.
Rostrum, palpi and antennaelight yellow; flagellar segmentssubcylindrical, slightly
narrowed at incisures; verticils short, longest about one-half of segment or slightly
more. Head yellow.
Mesonotal praescutum and scutum yellow, former with three more orange stripes,
scutal lobes orange; scutellum light brown, parascutella light yellow; mediotergite
light brown, slightly darker behind, pleurotergite and pleura orange yellow, dorsopleural membranelight yellow. Halteres broken. Legs with coxaeand trochanterslightyellow; femora light yellow, tips narrowly brownish-black; tibiae yellow, basesand tips
narrowly brownish-black, the amount less than half the femoral darkening; tibiae
brownish-yellow, tarsi similar, outer segmentsslightly darker, claws smooth. Wings(ng. g) uniformly deepyellow, veins darker yellow, evident againstthe ground. Venation:
R2 ver/ oblique, subequal to R3, leaving a very short element R21s; cell Rz at costaslightly more than one-half cell Rg; cell rsl M2large, rn-cu atforkof M; cell Cu narrowly
openat margin; cell znd Abroad.
Abdomen light yellow, tergites broadly brown medially to form a broad longitudinal
stripe. Ovipositor with valves, especiallycerci, long and slender.

C.P.Alexander

tc' R,,,-\____FR.
<---nL=.-

- d.

r'#

,'
Wd'\ffrm
TUI

WFb{"
ll

I

\u

\-a

t

\,rr\*
l4

.

p'

-$l

15

Figs. 7-r5. (7-r3) venation of: (7) Trmtepohlia (Mongorna)ry.qhinioha Alexander;(s) ?. (M.)flaooidessp. n.; (ro) T: (M) patms 'Plexandet:'<e;T.(M.r;yttig*o"p'n.;(Brunetti); (tz) T.(Trmtepohlia) infernaksAlexander;o*ota(irlesiomoigomilf.
itrl
sp. n. (r4, rs) Malehypop-ygiumof: (r4) Gononryia
ittj n Uptrro (Eri-optera)scoloph,ora(r5)Erioptera (Erioptaa) scolophorasp. n. Venation:n.;sp.
cobetwa
iFioneoi""ryiaj
it, ,"&,rr; ,3c, subcosta.Male hypopygium: a, aedeagus;b, basistyle;d, dististyles;p'phallosome;l, ninth tergite.

Holotype J, INore: Talung Dzong, Kameng, North East Frontier Agency, Assam,
78oo feet, 6.vi. 196r (F.Schnid).

Exotic Tipulidae
The present fly is generally similar to Trentepohlia(Mongoma)flaoa (Brunetti) but
is much larger, with the wings deep yellow with darker yellow veins and the abdomenlight yellow with a broad brown median stripe on the tergites.
Tr entepohlia(M ongoma)patms Alexander
Trentepohlia(Mongoma)patms Alexander, rg67b J. N.Y. ent. Soc.75 : 24.
Type-Pynter, Khasi-Jaintia, Assam,rToo feet, zo.r.tg6o (F.Schmid). Venation(fig. ro).
Tr entEohlia (Trentepohlia)infnnalis Alexander
Trentepohlia(Trentepohlia)infernalisAlexander, ry67b J. N.Y. mt. Soc.75 : 25.
Type-Lingtham, Sikkim,65oofeet,4-ro . viii. r959 (F. Schmid).Venation(fig. rz).
Erioptera (Erioptera) scolophora sp. n.
Sizesmall (wing of male about 4 mm.); headwhitened; generalcolorationof mesonotal
praescutumand scutal lobes obscureyellow, lateral borders more whitened, pleura
yellow, with a broad pale brown longitudinal stripe; knobs of halteresblackened;legs
yellow; wings light yellow, veins more saturatedyellow; male hypopygiumwith outerdististyle a slender straight spine, inner style a curved blade, on outer margin bearing
an appressedstraight black spine.
Male. Length, about 3.5 mm.i wing, 4 mm.; antenna,about r mm.
Rostrum yellow, palpi black. Antennaewith proximal six or sevensegmentsyellowed,
remaining segments light brown; flagellar segments subcylindrical, verticils slightly
exceedingthe segments.Headwhitened.
Pronotum and pretergites whitened. Mesonotal praescutum with disk obscureyellow, humeral and lateral bordersnarrowly more whitened, scutallobesobscureyellow,
posterior scleritesof notum paler yellow. Pleura with a broad pale brown longitudinal
stripe extending from fore coxaebackwardover the dorsal sternopleurite,ventral pteropleuriteand katapleurotergite;
dorsalpleura,with the dorsopleuralmembrane,whitened;
ventral pleura, including the lower sternopleurite, broad meron, and metapleura lightyellow. Halteres with stem yellow, knob brownish-black. Legs with fore coxae pale
brown, remaining coxae and all trochanters yellow; remainder of legs yellow, outer
two tarsal segmentsslightly darker. Wings (fig. t:) light yellow, veins more saturated
yellow, trichia slightly darker. Venation as in the typical subgenus,vein R213 slightly
i shorter than R2; vein znd A very strongly sinuous, the outer half nearly parallel to the! posteriorborderof wing.
Abdomen obscureyellow, the hypopygial dististyles darker. Male hypopygium (fig.
II5)
with ninth tergite (r) slightly narrowed outwardly, apex with two lobes that areI
separatedby a subequalmedian emargination;proctiger projectingbeyond the tergalImarginas
two slender lobes, each tipped with a single strong seta. Basistyle (D) withI
apicallobe
relatively long, narrowly cylindrical. Outer dististyle (d) a slender straightI
spine;
inper
style subequalin length, appearingas a gently curved blade that is shapedI
likeaboomerang,
at near mid-length of outer margin with a subappressed
slenderstraightI
I spinethat is about one-half aslong as the outer style. Phallosome(p) with gonapophyses
f yellow, apicesdarkened,slightly expanded; aedeagusconspicuouslybifurcate as in thei subgenus,the arms long and slender, strongly recurved near outer ends.

i

t

II

T

ro

C. P. Alexander: Exotic Tipulidae

HolotypeJ, Ixore: Kelang, Kameng, North East Frontier Agency, Assam,6ooofeet,r-z.vii.rg6t. Paratopotype,$, pinned with holotype.PmatypeJ, Dirang Dzong, Kameng, 53oo-65oofeet, fi-zz.vii.196r. (All F.Schmid).
Other regional speciesof the subgenusthat have the inner dististyle of the malehypopygium bifid as in the present fly but of different conformation include Erioptua(Erioptera)orbitalis Alexander (ornatifronsEdwards), widely distributed in southeastern
Asia, E. (8.) himalayaeAlexander, North India, E. (E.) lunicolaAlexander, Philippines,
E.(8.) nigripalpis de Meijere, Java and E. (E.) pila Alexander, North India, all quitedistinct in hypopygialcharacters.
I.lt

REF E R E N C E S)

Ar,sxANoon C.P. r95r. New or little-known Tipulidae (Diptera). Ann. Mag. nat. Hist. (rz) 4:
roTr-rro2.
Ar,xexonn C.P. tg67a. New or little-known Tipulidae from eastern Asia (Diptera).62. Philipp.
J. Sci.96 : zg4g.
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Ar,nxeNorn C.P. r97r. New or little-known species of exotic Tipulidae (Diptera) XUII. J. Ent.
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lVlanuscript received zrst February, rgTz

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