Flora of China 12: 210–222. 2007.

8. VITIS Linnaeus, Sp. Pl. 1: 202. 1753.
葡萄属 pu tao shu
Ren Hui (任辉); Jun Wen (文军)Ampelovitis Carrière; Spinovitis Romanet du Caillaud.
Lianas, woody, usually polygamo-dioecious, rarely hermaphroditic. Bark usually shedding; tendrils leaf-opposed, usuallybifurcate. Leaves simple, often lobed, sometimes palmately compound; stipules usually caducous. Inflorescence a thyrse. Flowers
5-merous. Calyx saucer-shaped; sepals minute. Petals united at apex and shed as a cap (calyptra) at anthesis. Stamens opposite topetals, undeveloped and abortive in female flowers. Disk conspicuous, 5-lobed or ring-shaped. Pistil 1; style slender; stigma
slightly expanded. Berry globose, 2–4-seeded. Seeds obovoid or obovoid-elliptic, base rostrate, abaxially 1-furrowed with arounded or suborbicular or elliptic chalazal knot, adaxially 2-furrowed; endosperm M-shaped in cross-section.
About 60 species: mostly in temperate regions, extending to the subtropics with the species diversity primarily in China and E North America;
37 species (30 endemic, one introduced) in China.
Vitis vinifera Linnaeus, the well-known source plant for grapes and wine, has a long history of cultivation all over the world. The wild grape
species are primarily distributed in three centers of diversity: East Asia, North to Central America, and Europe to Central Asia. In China, Vitis occurs
naturally in all provinces except Xinjiang, Qinghai, Nei Mongol, and Ningxia. The roots, stems, and leaves of several wild species are used as
medicine; the berries are edible or used to make wine; and the seeds are sometimes used for oil.
Vitis chrysobotrys H. Léveillé & Vaniot (Repert. Spec. Nov. Regni Veg. 3: 350. 1907) is a synonym of Iodes vitiginea (Hance) Hemsley, and V.
seguinii H. Léveillé (Repert. Spec. Nov. Regni Veg. 4: 331. 1907, “seguini”) is the basionym of I. seguinii (H. Léveillé) Rehder, both in the Icacinaceae (see Fl. China 11).
Vitis mairei H. Léveillé (Repert. Spec. Nov. Regni Veg. 11: 299. 1912, not 7: 338. 1909) and V. martini H. Léveillé & Vaniot (Bull. Soc.
Agric. Sarthe 40: 41. 1905) are synonyms of Gynostemma pentaphyllum (Thunberg) Makino in the Cucurbitaceae (see Fl. China 19).
The following species were described from China but could not be treated here because no material was seen by the authors:
Vitis adenoclada Handel-Mazzetti (Anz. Akad. Wiss. Wien, Math.-Naturwiss. Kl. 62: 145. 1925), described from Hunan.
Vitis jinzhainensis X. S. Shen (Acta Phytotax. Sin. 27: 304. 1989), described from Anhui.
Vitis shifunensis Hayata (Icon. Pl. Formosan. 5: 31. 1915; V. triphylla Hayata, J. Coll. Sci. Imp. Univ. Tokyo 30(1): 63. 1911, not Noronha,1790), described from Taiwan.
Vitis yuenlingensis W. T. Wang (Bull. Bot. Res., Harbin 9(1): 75. 1989), described from Hunan.

1a. Leaves palmately compound with 3–5 leaflets.
2a. Branchlets and inflorescence axes pilose; leaflets abaxially sparsely pubescent, or with woolly hairs when
young, becoming sparsely so when mature ........................................................................................................... 10. V. piasezkii
2b. Branchlets and inflorescence axes with arachnoid tomentum; leaflets abaxially with persistent
rubiginous arachnoid tomentum ............................................................................................................... 37. V. lanceolatifoliosa
1b. Leaves simple.
3a. Branchlets with prickles, becoming tuberculate on old branches .............................................................................. 1. V. davidii
3b. Branchlets and old branches without prickles or tubercles.
4a. Branchlets and petioles with setose and glandular hairs, glands with or without a stalk.
5a. Branchlets and petioles with dense glandular as well as simple hairs; leaves inconspicuously
3–5-lobed or unlobed, margin with sharp teeth ....................................................................................... 2. V. romanetii
5b. Branchlets and petioles with sparse glandular hairs; leaves conspicuously 3–5-lobed, margin withobtuse teeth ............................................................................................................................................ 3. V. shenxiensis
4b. Branchlets and petioles pilose or with arachnoid tomentum.
6a. Leaves abaxially with dense white or rubiginous arachnoid or woolly tomentum.
7a. Leaves 3–5-lobed or heterophyllous and mixed with undivided ones.
8a. Leaves 3–5-lobed, lobes broad.
9a. Branches with white woolly tomentum, erect hairs absent; tendrils bifurcate .................. 28. V. heyneana
9b. Branches pilose; tendrils unbranched or mixed with bifurcate ones .............................. 35. V. sinocinerea
8b. Leaves 3–5-cleft or deeply parted, mixed with undivided ones.
10a. Leaves of two types, 3–5-cleft and undivided ......................................................... 31. V. longquanensis
10b. Leaves all divided deeply or at least to middle, sometimes deep lobes pinnatifid,
some lobes broad and undivided .................................................................................. 36. V. bryoniifolia
7b. Leaves undivided or inconspicuously 3–5-lobed.
11a. Leaves basally deeply cordate, notch rounded or acutely angular, sinuses close together
or overlapping.

Flora of China 12: 210–222. 2007.

12a. Leaves oval or ovate-elliptic, abaxial veins with woolly tomentum ................................ 29. V. retordii
12b. Leaves suborbicular or quinquangularly rounded, abaxial veins densely
pubescent with appressed soft hairs ........................................................................ 30. V. menghaiensis
11b. Leaves basally slightly cordate or subtruncate, sometimes mixed with conspicuously
cordate ones.
13a. Branchlets and inflorescence axis ± with arachnoid tomentum ................................... 28. V. heyneana
13b. Branchlets and inflorescence axis with variable hairiness (including arachnoid
tomentum).
14a. Leaves adaxially with sparse arachnoid tomentum when young, then glabrescent;branchlets and inflorescence axis with sparse arachnoid tomentum ....................... 32. V. bellula
14b. Leaves adaxially or only veins with dense, short, appressed hairs; inflorescenceaxis densely pubescent.
15a. Branchlets sparsely pilose; leaves slightly lobed, base truncate, only
adaxial veins sparsely pubescent ............................................................... 33. V. bashanica
15b. Branchlets densely pubescent; leaves undivided, base subcordate,adaxially densely pubescent .................................................................................. 34. V. hui
6b. Leaves abaxially glabrous, or pilose, or with sparse arachnoid tomentum.
16a. Leaves abaxially glabrous or only axils of veins with hairs, or young leaves with tomentum thenfalling off.
17a. Leaves sagittate or deltoid-sagittate, 3–5-lobed or cleft, or mixed with undivided ones;
inflorescence narrow, small, cylindrical, with basal branches poorly developed.
18a. Leaf margin 12–16-toothed on each side, conspicuously ciliate, abaxial
veinlets raised ...................................................................................................... 17. V. wenchowensis
18b. Leaf margin 7–10-toothed on each side, only upper part of margin with very
short and sparse hairs, not ciliate, abaxial veinlets conspicuous, but not raised ... 20. V. ruyuanensis
17b. Leaves variously shaped but neither sagittate nor deltoid-sagittate, undivided.
19a. Leaves ovate, oval, elliptic, or ovate-lanceolate, base slightly subcordate or
subtruncate.
20a. Leaves ovate or oval, not glaucous, veinlets inconspicuous .............................. 25. V. flexuosa
20b. Leaves elliptic or ovate-lanceolate, usually glaucous, veinlets conspicuous andraised ........................................................................................................................ 8. V. chungii
19b. Leaves oval, broadly ovate, or triangular-ovate, base cordate or deeply so.
21a. Basal sinuses close together or partly overlaid.
22a. Leaves subleathery, margin serrate, apex sharply acute, abaxiallyusually glaucous, rarely glaucescent ................................................... 6. V. chunganensis
22b. Leaves papery, margin coarsely dentate, apex cuspidate, abaxiallynot glaucous ......................................................................................... 21. V. mengziensis
21b. Basal sinuses forming an obtuse angle without any overlapping.
23a. Peduncles with sparse arachnoid tomentum when young, then graduallyfalling off ................................................................................................... 4. V. balansana
23b. Peduncles pilose.
24a. Leaves oval, apex acute or acuminate, veinlets slightly raised ... 5. V. yunnanensis
24b. Leaves ovate-oblong or triangular-ovate, apex acuminate,veinlets conspicuously raised .................................................... 7. V. luochengensis
16b. Leaves abaxially pilose or at least abaxial veins pilose or with arachnoid tomentum.
25a. Leaves conspicuously 3–5-divided or mixed with undivided leaves.
26a. Leaves basally deeply cordate, basal sinuses attaching or partly overlapping,margin coarsely toothed (cultivated) ............................................................................. 27. V. vinifera
26b. Leaves basally cordate, basal notch obtuse or rounded, margin shallowly toothed.
27a. Tendrils unbranched, rarely bifurcate.
28a. Branchlets and petioles sparsely brown villous; leaves undivided or
mixed with 2- or 3-cleft leaves, margin with long ciliate hairs .............. 23. V. hancockii
28b. Branchlets, petioles, and peduncles sparsely pilose; leaves usually
3–5-lobed or cleft, margin not ciliate or only slightly so with shortciliate hairs.
29a. Leaves abaxially greenish or lavender, veinlets slightly
raised ........................................................................................ 18. V. jinggangensis
29b. Leaves abaxially reddish purple, veinlets conspicuously
raised on both sides .................................................................. 19. V. erythrophylla
27b. Tendrils 2(or 3)-branched.

Flora of China 12: 210–222. 2007.

30a. Leaves unlobed or 3–5-lobed, lobes broad.
31a. Leaves oval or obelliptic, basal sinuses forming an obtuse
angle ................................................................................................. 9. V. betulifolia
31b. Leaves broadly oval, basal sinuses rounded, rarely forming an
obtuse angle ................................................................................... 26. V. amurensis
30b. Leaves 3–5-cleft to middle or deeply parted, lobes narrow, rarely lobes
pinnatifid, or sometimes mixed with shallowly lobed leaves.
32a. Plants small, stems slender; leaf margin 5–9-toothed on eachside .................................................................................................... 15. V. silvestrii
32b. Plants strong, stems robust; leaf margin 16–25-toothedon each side ...................................................................... 14. V. zhejiang-adstricta
25b. Leaves undivided, rarely inconspicuously 3–5-lobed.
33a. Leaves abaxially with at least veins with arachnoid tomentum, rarely glabrous,
never with erect hairs.
34a. Inflorescence axis pilose and with arachnoid tomentum .............................. 22. V. fengqinensis
34b. Inflorescence axis with arachnoid tomentum, but not erect hairs.
35a. Leaves 7–16 × 5–12 cm, margin 16–20-toothed on each side ................ 12. V. wilsoniae
35b. Leaves 3–7 × 2.5–6 cm, margin 5–13-toothed on each side ................ 16. V. wuhanensis
33b. Leaves abaxially with at least veins with erect hairs, or sometimes mixed witharachnoid tomentum.
36a. Leaves rhombic-ovate or rhombic-elliptic, base broadly cuneate or
suborbicular ....................................................................................................... 23. V. hancockii
36b. Leaves ovate or oval, base cordate, subcordate, or subtruncate, rarely rounded.
37a. Leaves basally conspicuously cordate.
38a. Branchlets glabrous; abaxial veins of leaves pilose withspreading brown hairs ................................................................. 11. V. pilosonerva
38b. Branchlets with sparse woolly tomentum; abaxial veins ofleaves with white hairs and with sparse arachnoid
tomentum ............................................................................. 13. V. pseudoreticulata
37b. Leaves basally subcordate or subtruncate, rarely rounded.
39a. Tendrils bifurcate; leaves oval or ovate-elliptic .............................. 9. V. betulifolia
39b. Tendrils unbranched; leaves ovate or oval-lanceolate ............................. 24. V. tsoi
1. Vitis davidii (Romanet du Caillaud) Föex, Cours Compl.
Vitic. 44. 1886.
刺葡萄 ci pu tao
Plants polygamo-dioecious. Branchlets prickly or tuberculately so, terete, with longitudinal ridges when young, glabrescent; tendrils bifurcate. Stipules greenish brown, ovate-lanceolate, 2–3 × 1–2 mm, nearly herbaceous, glabrous, caducous;
leaf blade oval or oval-elliptic, undivided or shallowly 3-lobed,
5–12 × 4–16 cm, basal veins 5, lateral veins 4 or 5 pairs, veinlets conspicuous and raised, base deeply cordate, margin with
12–33 teeth on each side, teeth sharp, apex acute or mucronate.
Panicle leaf-opposed, 7–24 cm, with well-developed basal
branches; peduncle 1–2.5 cm, glabrous. Pedicel 1–2 mm, glabrous. Buds obovoid, 1.2–1.5 mm, apex rounded. Sepals minute. Filaments filiform; anthers yellow, elliptic, 0.6–0.7 mm.
Ovary conical; style short; stigma expanded. Berry globose,
1.2–2.5 cm in diam. Seeds obovoid-elliptic, apex rounded andobtuse, ventral holes furrowed upward 3/4 from base. Fl. Apr–
Jun, fr. Jul–Oct.
● Forests, shrublands, hillsides, valleys; 500–2300 m. Anhui,
Chongqing, Fujian, Gansu, Guangdong, Guangxi, Guizhou, Hubei,
Hunan, Jiangsu, Jiangxi, Shaanxi, Sichuan, Yunnan, Zhejiang.
Vitis davidii var. hispida X. D. Wang & S. C. Chen (Acta Phytotax. Sin. 37: 603. 1999), described from Fujian, could not be treatedhere because no material was seen by the authors.

1a. Leaves abaxially ferruginous pubescent................................................................ 1c. var. ferruginea
1b. Leaves abaxially glabrous.
2a. Berry purple at maturity ..................... 1a. var. davidii
2b. Berry blue-black at maturity ...... 1b. var. cyanocarpa
1a. Vitis davidii var. davidii
刺葡萄(原变种) ci pu tao (yuan bian zhong)Spinovitis davidii Romanet du Caillaud, Rev. Hort. (Paris)
55: 53. 1883 [“davidi”]; Ampelovitis davidii (Romanet duCaillaud) Carrière; Vitis armata Diels & Gilg; V. prunisapida
H. Léveillé & Vaniot.
Branchlets with prickles. Leaves abaxially glabrous. Berry
purple at maturity. Fl. Apr–Jun, fr. Jul–Oct.
● Shrublands, hillsides, valleys; 600–1800 m. Anhui, Chongqing,
Fujian, Gansu, Guangdong, Guangxi, Guizhou, Hubei, Hunan, Jiangsu,
Jiangxi, Shaanxi, Sichuan, Yunnan, Zhejiang.

1b. Vitis davidii var. cyanocarpa (Gagnepain) Sargent, Pl. Wilson. 1: 104. 1911.
蓝果刺葡萄 lan guo ci pu tao
Vitis armata var. cyanocarpa Gagnepain in Sargent, Pl.

Flora of China 12: 210–222. 2007.

Wilson. 1: 104. 1911.
Old branches tuberculate; young branchlets without
prickles or only sparsely prickly. Leaves abaxially glabrous.
Berry blue-black at maturity. Fl. Apr–May, fr. Aug–Sep.
● Forests, shrublands; 600–2300 m. Anhui, Hubei, Yunnan.

1c. Vitis davidii var. ferruginea Merrill & Chun, Sunyatsenia
1: 69. 1930.
锈毛刺葡萄 xiu mao ci pu tao
Leaves abaxially ferruginous pubescent on veins. Fl. Apr–Jul.
● Shrublands, hillsides; 500–1200 m. Fujian, Guangdong, Hubei,
Jiangxi.

2. Vitis romanetii Romanet du Caillaud, Rev. Hort. (Paris) 55:
53. 1883 [“romaneti”].
秋葡萄 qiu pu tao
Ampelovitis romanetii (Romanet du Caillaud) Carrière;
Vitis pentagona Diels & Gilg var. laotica Gagnepain; V. romanetii var. arachnoidea Y. L. Cao & Y. H. He; V. rutilans
Carrière.
Plants polygamo-dioecious. Branchlets terete, with conspicuous longitudinal ridges, pubescent with simple and glandular hairs; tendrils usually bifurcate or trifurcate. Leaves
simple; stipules brown, ovate-lanceolate, 7–12(–14) × 3–5 mm,
membranous, glabrous, entire, apex acuminate; petiole 2–6.5
cm, with glandular and simple hairs; leaf blade oval or broadly
so, 5.5–16 × 5–13.5 cm, slightly 5-lobed or undivided, abaxially tomentose with long and short hairs, adaxially pubescent,basal veins 5, lateral veins 4 or 5 pairs, veinlets conspicuously
raised abaxially, slightly raised adaxially, base deeply cordate,notch acute, rarely obtuse, sometimes two sides overlapping,margin with obtuse teeth, teeth sharp. Panicle loose, leafopposed, 5–13 cm, basal branches well developed, sometimes
with a tendril below basal branch; peduncle 1.5–3.5 cm,
densely pubescent with simple and glandular hairs. Pedicel 1.6–
2 mm, glabrous. Buds obovoid, 1.5–2 mm, apex rounded. Calyx cupular, ca. 2 mm, glabrous, subentire. Petals galeate. Filaments filiform, 1.4–1.8 mm; anthers yellow, elliptic, ca. 0.5
mm. Ovary conical; style short; stigma expanded. Berry globose, 7–8 mm in diam. Seeds obovoid, apex rounded and retuse, chalazal knot ovate-elliptic, ventral holes obelliptic. Fl.
Apr–Jun, fr. Jul–Sep. 2n = 38.
Forests, shrublands, hillsides; 100–1500 m. Anhui, Gansu, Henan,
Hubei, Hunan, Jiangsu, Shaanxi, Sichuan [Laos].
The berries are edible and used for making wine.

3. Vitis shenxiensis C. L. Li, Chin. J. Appl. Environ. Biol. 2:239. 1996.
陕西葡萄 shan xi pu tao
Plants polygamo-dioecious. Branchlets terete, with longitudinal ridges, sparsely pubescent with simple and stalked
glandular hairs, glands 0.8–1 mm; tendrils usually bifurcate.

Leaves simple; petiole 4–5.5 cm, sparsely pubescent withsimple and glandular hairs; leaf blade oval, conspicuously 3–5lobed, rarely tripartite or 3-divided, notch obtuse, rarely acute,
11–16 × 9–14 cm, abaxially sparsely pilose, then glabrescent,adaxially nearly glabrous, basal veins 5, lateral veins 5 or 6
pairs, pilose with spreading hairs, rarely glabrous, base cordate,margin irregularly 25–32-toothed, teeth large and obtuse, apex
acuminate or mucronulate, rarely acute. Panicle leaf-opposed,
loose, 8–9 cm, with well-developed basal branches; peduncle
1.5–2 cm, nearly glabrous. Pedicel 3–4 mm, glabrous. Buds
obovoid, 2–2.5 mm, apex rounded. Calyx cupular, glabrous,
entire. Filaments filiform, 1.7–2 mm; anthers yellow, elliptic,
ca. 0.5 mm. Pistil abortive in male flowers. Berry purple-black
at maturity, globose, ca. 1.2 cm in diam. Seeds obovoid, apex
retuse, surface smooth. Fl. Jun, fr. Oct.
● Shrublands, hillsides, valleys; 1100–1400 m. Shaanxi.

4. Vitis balansana Planchon in A. Candolle & C. Candolle,
Monogr. Phan. 5: 612. 1887 [“balansaeana”].
小果葡萄 xiao guo pu tao
Branchlets terete, with longitudinal ridges; tendrils bifurcate, leaf-opposed. Stipules brown, oval to oblong, 2–4 × 1.5–3mm; petiole 2–5 cm; leaf blade adaxially green, cordate-oval or
broadly ovate, 4–14 × 3.5–9.5 cm, basal veins 5, lateral veins
4–6 pairs, veinlets conspicuous and raised, base cordate, notchobtuse, margin with 16–22 fine teeth on each side, slightly
undulate, apex acute or mucronulate. Panicle leaf-opposed, 4–
13 cm. Pedicel 1–1.5 mm. Buds obovoid, 1–1.4 mm, apex
rounded. Calyx cupular, entire. Petals galeate. Filaments filiform in male flowers, 0.6–1 mm; anthers yellow, elliptic, ca.
0.4 mm; stamens shorter than pistils in female flowers, abortive.
Disk 0.3–0.4 mm. Ovary conical; style short; stigma slightly
expanded. Berry purple-black at maturity, globose, 5–8 mm indiam. Seeds obovoid, apex rounded, chalazal knot elliptic. Fl.
Feb–Aug, fr. Jun–Nov.
Forests, shrublands, valleys; 200–800 m. Guangdong, Guangxi,
Hainan [Vietnam].

1a. Leaves 3-cleft ...................................... 4c. var. ficifolioides
1b. Leaves undivided.
2a. Branchlets, leaves, and petioles withsparse arachnoid tomentum when
young, becoming glabrescent ....... 4a. var. balansana
2b. Branchlets, leaves, and petioles with
dense, persistent, white arachnoid
tomentum ...................................... 4b. var. tomentosa
4a. Vitis balansana var. balansana
小果葡萄(原变种) xiao guo pu tao (yuan bian zhong)Vitis flexuosa Thunberg var. gaudichaudii Planchon.
Branchlets, petioles, adaxial leaf surface, inflorescenceaxes, and peduncles with sparse arachnoid tomentum when
young, becoming glabrescent. Stipules glabrous or with arachnoid tomentum. Pedicel and calyx glabrous.
Forests, shrublands in valleys, sunny valleys; 200–800 m. Guangdong, Guangxi, Hainan [Vietnam].

Flora of China 12: 210–222. 2007.

4b. Vitis balansana var. tomentosa C. L. Li, Chin. J. Appl.
Environ. Biol. 2: 240. 1996.
绒毛小果葡萄 rong mao xiao guo pu tao
Branchlets, petioles, abaxial leaf surface, and inflorescences with dense, persistent, white arachnoid tomentum.
● Guangxi.

4c. Vitis balansana var. ficifolioides (W. T. Wang) C. L. Li,
Chin. J. Appl. Environ. Biol. 2: 240. 1996.
龙州葡萄 long zhou pu tao
Vitis ficifolioides W. T. Wang, Acta Phytotax. Sin. 17(3):75. 1979.
Leaves 3-cleft, notch rounded.
● Guangxi (Longzhou).

5. Vitis yunnanensis C. L. Li, Acta Bot. Yunnan. 19: 217.
1997.

Branchlets usually purple, terete, with longitudinal ridges,
glabrous; tendrils bifurcate. Leaves simple; stipules caducous;
petiole 2.5–5 cm, glabrous; leaf blade ovate-oblong or ovate,
12–18 × 4–12 cm, basal veins 5, lateral veins 5 or 6 pairs, veinlets conspicuously raised, base cordate, notch obtuse, margin
with 15–21 fine teeth on each side, apex acuminate. Panicle
leaf-opposed, ca. 17 cm at fruiting stage, with well-developed
branches; peduncle ca. 2.5 cm. Pedicel 4–7 mm, nearly glabrous. Berry globose, ca. 0.5 mm in diam. Fr. May–Jul.
● Forests, shrublands on rocky hills, valleys; 400–700 m. Guangdong, Guangxi.

1a. Leaves glabrous ............................... 7a. var. luochengensis
1b. Abaxial veins of leaves with dense
white tomentum ............................ 7b. var. tomentosonerva
7a. Vitis luochengensis var. luochengensis
罗城葡萄(原变种) luo cheng pu tao (yuan bian zhong)

云南葡萄 yun nan pu tao

Leaves glabrous. Fr. Jul.

Branchlets terete, with longitudinal ridges, glabrous; tendrils bifurcate. Leaves with caducous stipules; petiole 1.5–4 cm,
glabrous; leaf blade oval, 5–10 × 4–7.5 cm, abaxially nearly
glabrous, basal veins 5, lateral veins 4 pairs, veinlets slightly
raised, base cordate, margin 15–17-toothed on each side, teeth
acute, apex acute or mucronulate. Panicle leaf-opposed, loose,base with well-developed branches, rachis densely pubescent.
Pedicel glabrous. Berry globose, ca. 1 cm in diam. Seeds obovoid, apex rounded, surface smooth, chalazal knot elliptic, ventral holes furrowed from base to apex. Fr. Aug.

● Shrublands on rocky hills. Guangxi.

● Forests; 500–1800 m. Yunnan (Jingdong, Jinghong).

6. Vitis chunganensis Hu, J. Arnold Arbor. 6: 143. 1925.
东南葡萄 dong nan pu tao
Plants polygamo-dioecious. Branchlets terete; old branches with conspicuous longitudinal ridges, glabrous; tendrils bifurcate. Stipules ovate-elliptic or lanceolate, 1.5–3 × 1–1.5 cm,
glabrous, apex obtuse, caducous; petiole 2–6.5 cm, glabrous;
leaf blade ovate or ovate-elliptic, 6.5–22.5 × 4.5–13.5 cm,
abaxially glaucous, rarely glaucescent, basal veins 5–7, lateralveins 5–7 pairs, veinlets inconspicuous, base cordate, margin
12–22-toothed on each side, teeth small, apex acute, acuminate,
or caudate. Panicle leaf-opposed, loose, 5–9 cm, base with welldeveloped branches, occasionally basal branches reduced totendrils; peduncle 1–2 cm, pubescent or nearly glabrous. Pedicel 1.2–2 mm, glabrous. Buds globose or elliptic, 1–1.5 mm,
glabrous. Calyx ca. 2 mm, glabrous. Filaments filiform, 0.5–0.7mm; anthers yellow, elliptic, ca. 0.4 mm. Ovary oval; styleshort, slender; stigma expanded. Berry purple-black at maturity,
globose, 0.8–1.2 cm in diam. Seeds obovoid, apex retuse, chalazal knot elliptic. Fl. Apr–Jun, fr. Jun–Aug.
● Forests, shrublands, hillsides, streams in valleys; 500–1400 m.
Anhui, Fujian, Guangdong, Guangxi, Hunan, Jiangxi, Zhejiang.

7b. Vitis luochengensis var. tomentosonerva C. L. Li, Chin. J.
Appl. Environ. Biol. 2: 241. 1996 [“tomentoso-nerva”].
连山葡萄 lian shan pu tao
Branchlets, petioles, and abaxial veins of leaves with
dense white tomentum. Fr. May.
● Forests, valleys; 400–700 m. Guangdong.

8. Vitis chungii F. P. Metcalf, Lingnan Sci. J. 11: 102. 1932.
闽赣葡萄 min gan pu tao
Plants polygamo-dioecious. Branchlets terete, with longitudinal ridges, glabrous; tendrils bifurcate. Leaves simple; stipules brownish, lanceolate, 2.5–3 × 1–1.2 mm, membranous,
glabrous, caducous; petiole 1–3.5 cm, glabrous; leaf blade
ovate-elliptic or ovate-lanceolate, 4–15 × 2–8 cm, abaxially glabrous, glaucous, basal veins 3, lateral veins 4 or 5 pairs, veinlets
raised, glabrous, base truncate, rounded, or subrounded, rarely
subcordate, margin sparsely 7–9-toothed on each side, teeth
sharp, apex acuminate or caudate, rarely acute. Panicles leafopposed, terete, 3.5–10 cm, basal branches poorly developed;
peduncle 1.5–2.5 cm, pubescent when young, then glabrescent.
Pedicel 1–2.5 mm, glabrous. Buds obovoid, 1–1.4 mm, apex
rounded. Calyx entire. Filaments filiform, 0.8–1.2 mm; anthers
yellow, elliptic, 0.4–0.5 mm. Ovary oval; style short; stigma
expanded. Berry purple-red at maturity, globose, 8–10 mm indiam. Seeds obelliptic, apex rounded, chalazal knot elliptic,
ventral holes obelliptic, furrowed upward 3/4 from base.
● Forests in valleys, shrublands, hillsides; 200–1000 m. Fujian,Guangdong, Guangxi, Jiangxi.

7. Vitis luochengensis W. T. Wang, Guihaia 8: 110. 1988.

9. Vitis betulifolia Diels & Gilg, Bot. Jahrb. Syst. 29: 461.
1900.

罗城葡萄 luo cheng pu tao桦叶葡萄 hua ye pu tao
Vitis hexamera Gagnepain; V. shimenensis W. T. Wang;

Flora of China 12: 210–222. 2007.

V. trichoclada Diels & Gilg.
Branchlets terete, with conspicuous longitudinal ridges
and sparse lanate tomentum when young, then becoming glabrescent; tendrils bifurcate, leaf-opposed. Leaves simple; stipules brownish, lanceolate, 2.5–6 × 1.5–3 mm, membranous,
glabrous, margin entire, apex acute or obtuse; petiole 2–6.5 cm,
with arachnoid tomentum when young, then glabrescent; leaf
blade oval or ovate-elliptic, undivided or 3-lobed, 4–12 × 3.5–9cm, abaxially densely tomentose when young, then hairs falling
off and only veins pubescent or nearly glabrous, adaxially pubescent and with sparse arachnoid tomentum when young, thenglabrescent, basal veins 5, lateral veins 4–6 pairs, veinlets
slightly raised abaxially, base cordate or subtruncate, rarely
subrounded on upper leaves, margin 15–25-toothed on eachside, teeth sharp, apex acute or acuminate. Panicle leaf-opposed, with well-developed basal branches, loose, 4–15 cm,
with arachnoid tomentum when young, then nearly glabrescent.
Pedicel 1.5–3 mm, glabrous. Buds obovoid, 1.5–2 mm, apex
rounded. Calyx ca. 0.2 mm, margin membranous, entire. Filaments filiform, 1–1.5 mm; anthers yellow, elliptic, ca. 0.4 mm.
Ovary oval; style short. Berry purple-black at maturity, globose,
8–10 mm in diam. Seeds obovoid, apex rounded, chalazal knot
rounded or elliptic, ventral holes furrowed upward 2/3–3/4 frombase. Fl. Mar–Jun, fr. Jun–Nov.

● Forests, forest margins, shrublands, hillsides, riversides, usuallyin sunny and disturbed areas; 900–2100 m. Chongqing, Gansu, Hebei,
Henan, Shaanxi, Shanxi, Sichuan, Zhejiang.

11. Vitis pilosonerva F. P. Metcalf, Lingnan Sci. J. 11: 14.
1932 [“piloso-nerva”].
毛脉葡萄 mao mai pu tao
Vitis davidii (Romanet du Caillaud) Föex var. brachytricha Merrill.
Plants polygamo-dioecious. Branchlets terete, with conspicuous longitudinal ridges, glabrous; tendrils bifurcate, leafopposed. Leaves simple; stipules caducous; petiole 3.5–11 cm,
sparsely pilose; leaf blade broadly oval, inconspicuously 3lobed, 10–16 × 9–14 cm, abaxially glaucous, densely pubescenton veins, basal veins 5, lateral veins 5–8 pairs, veinlets raised,base cordate, apex of notch acute, margin 28–36-toothed oneach side, teeth large and sharp, apex acute or mucronulate.
Panicle leaf-opposed, loose, 15–34 cm, with well-developed
branches; peduncle 5–9 cm, subglabrate. Pedicel 1.5–2 mm,
nearly glabrous. Buds obovoid, ca. 1.5 mm, apex rounded. Calyx glabrous, subentire, teeth 0.1–0.2 mm. Filaments filiform,ca. 1 mm; anthers yellow, elliptic, ca. 0.4 mm. Ovary abortivein male flowers. Fl. Jun.

● Forests in valleys, shrublands, hillsides; 600–3600 m. SE Gansu, Henan, Hubei, Hunan, Shaanxi, Sichuan, Yunnan.

● Forests, hillsides, valleys; 700–800 m. Fujian, Guangdong,
Jiangxi.

10. Vitis piasezkii Maximowicz, Bull. Acad. Imp. Sci. SaintPétersbourg 27: 461. 1881.

12. Vitis wilsoniae H. J. Veitch, Gard. Chron. 46: 236. 1909[“wilsonae”].

变叶葡萄 bian ye pu tao网脉葡萄 wang mai pu tao

Parthenocissus sinensis Diels & Gilg; Vitis baihensis P. C.
He; V. pagnuccii Romanet du Caillaud; V. piasezkii var.
baroniana Diels & Gilg; V. piasezkii var. pagnuccii (Romanetdu Caillaud) Rehder; V. tiubaensis X. L. Niu.

Vitis marchandii H. Léveillé; V. reticulata Pampanini(1910) and Gagnepain (1911), not M. A. Lawson (1875).

Branchlets terete, with longitudinal ridges; tendrils bifurcate, leaf-opposed. Leaves highly variable in shape, 3–5-foliolate, or mixed with simple and unlobed, or simple and variously lobed leaves; stipules caducous; petiole 2.5–6 cm; leaf
and leaflet blades adaxially nearly glabrous, or pilose andlanate, basal veins 5, lateral veins 4–6 pairs, pubescent to sometimes slightly so or nearly glabrescent, veinlets slightly raised
abaxially, inconspicuous adaxially; central leaflet rhombicelliptic or lanceolate, 5–12 × 2.5–5 cm, base cuneate, apex
acute or mucronulate; lateral leaflets ovate-elliptic or ovatelanceolate, 3.5–9 × 3–5 cm, base asymmetric, subrounded or
broadly truncate, margin with 5–20 sharp teeth on each side,
apex acute or acuminate; simple leaves oval or ovate-elliptic,
5–12 × 4–8 cm, base cordate, notch obtuse, margin serratewith 21–31 teeth, apex acute. Panicle leaf-opposed, loose, 5–
12 cm, basal branches well developed; peduncle 1–2.5 cm.
Pedicel 1.5–2.5 mm, glabrous. Buds obovoid, 1–2.5 mm, apex
rounded. Calyx glabrous, margin undulate. Filaments filiform,
0.7–1 mm. Ovary oval; style short; stigma expanded. Berryblack-purple, globose, 0.8–1.3 in diam., glaucous. Seeds obovoid, apex retuse, chalazal knot oval, ventral holes furrowed
upward 1/4 from base. Fl. Jun, fr. Jul–Sep.

Branchlets terete, with longitudinal ridges, with sparse,
brownish arachnoid tomentum; tendrils bifurcate, leaf-opposed.
Leaves simple; stipules caducous; petiole 4–8 cm, nearly glabrous; leaf blade cordate to ovate-elliptic, 7–16 × 5–12 cm,
abaxially with brownish arachnoid tomentum on veins, adaxially glabrous or nearly so, basal veins 5, lateral veins 4 or 5
pairs, veinlets raised, base cordate, notch obtuse, margin 16–20toothed on each side, apex acute or acuminate. Panicle leafopposed, loose, 4–16 cm, with well-developed basal branches;peduncle 1.5–3.5 cm, with sparse arachnoid tomentum. Pedicel
2–3 mm, glabrous. Buds obelliptic, 1.5–3 mm, apex subtruncate. Calyx undulately lobed. Filaments filiform, 1.2–1.6 mm;anthers yellow, ovoid-elliptic, 0.8–1.2 mm. Ovary oval; styleshort; stigma expanded. Berry globose, 0.7–1.5 cm in diam.
Seeds obovoid, apex globose, chalazal knot oblong, rapheslightly raised, surface smooth, ventral holes furrowed frombase to nearly 1/4 near apex. Fl. May–Jul, fr. Jun–next Jan.
● Forests, shrublands, valleys, streamsides; 400–2000 m. Anhui,
Chongqing, Fujian, Gansu, Guizhou, Henan, Hubei, Hunan, Shaanxi,
Sichuan, Yunnan, Zhejiang.

13. Vitis pseudoreticulata W. T. Wang, Acta Phytotax. Sin.
17(3): 73. 1979.
华东葡萄 hua dong pu tao

Flora of China 12: 210–222. 2007.

Plants polygamo-dioecious. Branchlets terete, with conspicuous longitudinal ridges, sparsely lanate when young, becoming glabrescent; tendrils bifurcate, leaf-opposed. Leaves
simple; stipules caducous; petiole 3–6 cm, with long villous
arachnoid tomentum when young, then falling off and pubescent; leaf blade oval or oblong, 6–13 × 5–11 cm, both surfaces
with villous tomentum when young, then becoming sparsely so,
abaxially whitish pubescent on lateral veins, basal veins 5,
lateral veins 3–5 pairs, veinlets conspicuous abaxially, basedeeply cordate to cordate, notch rounded or obtuse, margin 16–
25-toothed on each side, teeth sharp and slightly irregular, apex
acute or acuminate, rarely rounded. Panicle leaf-opposed, loose,
5–11 cm, basal branches well developed, sometimes with atendril below basal branch, with sparse arachnoid tomentum
when young, then falling off. Pedicel 1–1.5 mm, glabrous. Buds
obovoid, 2–2.5 mm, apex rounded. Calyx glabrous, teeth inconspicuous. Filaments filiform, ca. 1 mm; anthers yellow, elliptic,
ca. 0.2 × 0.1 mm. Ovary conical; style short. Berry purple-black
at maturity, globose, 8–10 mm in diam. Seeds obovoid, apex
retuse, chalazal knot elliptic, ventral mid-raphe slightly raised.
Fl. Apr–Jun, fr. Jun–Oct.
Forests, shrublands, hillsides, grasslands, riversides; 100–300 m.
Anhui, Fujian, Guangdong, Guangxi, Henan, Hubei, Hunan, Jiangsu,
Jiangxi, Zhejiang [Korea].

14. Vitis zhejiang-adstricta P. L. Chiu, Bull. Bot. Res., Harbin10(3): 39. 1990.
浙江蘡薁 zhe jiang ying yu
Branchlets terete, slender, with thin longitudinal ridges,
with sparse arachnoid tomentum when young, eventually subglabrate; tendrils bifurcate, leaf-opposed. Leaves simple, 3–5lobed to parted, usually mixed with undivided leaves; petiole 2–
4 cm, sparsely pubescent; leaf blade ovate or ovate-pentagonal,
3–6 × 3–5 cm, both surfaces with sparse very short fine hairs
and midvein with fine stiff hairs, basal veins 5, lateral veins 3–5
pairs, veinlets slightly raised abaxially, base cordate, notch
rounded to acute, teeth obtuse, apex acute or acuminate; midlobes rhombic-ovate, base constricted, notch rounded. Panicle
3.5–8 cm, glabrous or subglabrate; peduncle 1–3 cm; bract
narrowly triangular, margin ciliate, then cilia falling off. Pedicel
2–3 mm, glabrous. Berry globose, 6–8 mm in diam.
● By streams in valleys; 600–700 m. Zhejiang.

15. Vitis silvestrii Pampanini, Nuovo Giorn. Bot. Ital., n.s., 17:430. 1910.
湖北葡萄 hu bei pu tao
Plants polygamo-dioecious. Branchlets terete, slender,
with longitudinal ridges, densely pubescent, eventually glabrescent; tendrils bifurcate, leaf-opposed. Leaves simple; stipules
brownish, lanceolate, 1.5–2 × 0.5–1 mm, membranous, sparsely
pilose or glabrate; petiole 1–3 cm, pubescent; leaf blade oval,
3–5 × 2–3 cm, 3–5-lobed or parted, abaxially pubescent, adaxially sparsely pubescent when young, then glabrescent, basalveins 5, lateral veins 3 or 4 pairs, base subcordate or subtruncate, notch retuse to obtuse, rarely acute or rounded, margin
5–9-toothed on each side, teeth obtuse, apex acute or acu-

minate. Panicle leaf-opposed, 2–4.5 cm, narrow, basal branches
well developed; peduncle 1–1.5 cm, pubescent or subglabrate.
Pedicel 2–3 mm, glabrous. Buds ovoid-elliptic, 1.5–2 mm, apex
rounded. Calyx subentire. Filaments filiform, 1.3–1.5 mm; anthers yellow, elliptic, ca. 0.4 mm. Pistil abortive in male flowers. Fl. May.
● Forests, forest margins, hillsides; 300–1200 m. W Hubei, S
Shaanxi.

16. Vitis wuhanensis C. L. Li, Chin. J. Appl. Environ. Biol. 2:
243. 1996.
武汉葡萄 wu han pu tao
Vitis wuhanensis var. arachnoidea X. D. Wang & C. L. Li.
Plants polygamo-dioecious. Branchlets terete, slender, withlongitudinal ridges, with sparse arachnoid tomentum; tendrils
bifurcate, leaf-opposed. Leaves simple; stipules brownish, elliptic-lanceolate, 2.5–3.5 × 1–1.5 mm, membranous, glabrous,
entire, apex obtuse or acute; petiole 1.5–4 cm, with arachnoid
tomentum when young, then glabrate; leaf blade ovate or oval,
3-lobed or inconspicuously 3–5-divided, 3–7 × 2.5–6 cm,
abaxially white with sparse arachnoid tomentum when young,then glabrate, adaxially glabrous, basal veins 5, lateral veins 4
or 5 pairs, veinlets conspicuous at fruiting stage, base cordate,margin 5–13-toothed on each side, teeth obtuse, apex acute or
acuminate. Panicle leaf-opposed, 2.5–4 cm, narrow, basal
branches undeveloped; peduncle 1.5–2.5 cm, with sparse arachnoid tomentum. Pedicel 2–3 mm, glabrous. Buds obovoid-elliptic, 1.5–2.5 mm, apex rounded. Calyx ca. 2 mm, subentire. Filaments filiform, 1.2–1.8 mm; anthers yellow, elliptic, ca. 0.5
mm. Pistil abortive in male flowers. Berry globose, 6–7 mm indiam. Seeds obovoid, apex retuse, chalazal knot elliptic, ventral
holes furrowed from base upward nearly to apex. Fl. Apr–May,
fr. May–Jul.
● Forests, shrublands in valleys, hillsides; 300–700 m. Henan(Xinyang), Hubei, Jiangxi.

17. Vitis wenchowensis C. Ling ex W. T. Wang, Acta Phytotax. Sin. 17(3): 74. 1979.
温州葡萄 wen zhou pu tao
Branchlets slender, with longitudinal ridges, glabrous; tendrils unbranched. Leaves simple; stipules caducous; petiole
1.8–3.2 cm, glabrous; leaf blade halberd-shaped or triangularovate, undivided or 3–5-lobed, divided leaves with obtuse or
acute notch, 4–9.3 × 2.8–4.5 cm, abaxially glabrous, adaxiallyvery shortly strigose on veins, basal veins 5, lateral veins 4 or 5
pairs, veinlets conspicuously raised, base cordate, margin irregularly 7–9-toothed on each side and shortly ciliate, apex longacuminate. Panicles leaf-opposed, 3.8–6 cm; peduncle andpedicels glabrous. Berry globose, 5–8 mm in diam. Seeds obelliptic, chalazal knot elliptic. Fr. Jun–Jul.
● Broad-leaved evergreen forests by streams in valleys. Zhejiang.

18. Vitis jinggangensis W. T. Wang, Bull. Bot. Res., Harbin
1(1–2): 167. 1981.
井冈葡萄 jing gang pu tao

Flora of China 12: 210–222. 2007.

Branchlets terete, with longitudinal ridges, densely pubescent; tendrils unbranched, leaf-opposed. Leaves simple; stipules
caducous; petiole 3–4 cm, sparsely pubescent; leaf blade oval
or ovate-elliptic, usually 3-lobed, rarely 3–5-cleft, 7–9 × 6–7
cm, adaxially pubescent on veins at lower part, basal veins 5,
lateral veins 3 or 4 pairs, veinlets conspicuously raised abaxially, inconspicuous adaxially, base slightly cordate or subtruncate, lobes with base cordate, notch obtuse or acute, rarely
rounded, margin undulately 8–12-toothed on each side, apex
acuminate or mucronulate. Panicle leaf-opposed, subcylindric,
4–4.5 cm, narrow, basal branches well developed; peduncle 2–
2.7 cm, pubescent. Pedicel 2–2.5 mm, glabrous. Buds obovoid,
1.8–2 mm, apex rounded. Calyx ca. 1.5 mm, subentire. Filaments filiform. Disk well developed, slightly 5-lobed. Pistil
abortive in male flowers. Fl. Apr.
● Shrublands on hillsides; ca. 1000 m. Hunan (Yongshun), Jiangxi.

19. Vitis erythrophylla W. T. Wang, Bull. Bot. Res., Harbin
1(1–2): 168. 1981.
红叶葡萄 hong ye pu tao
Plants polygamo-dioecious. Branchlets terete, slender,
with longitudinal ridges, sparsely pubescent when young, thensubglabrate; tendrils unbranched, leaf-opposed. Leaves simple,
3–5-lobed to -cleft; stipules brownish, lanceolate, 2–3 × 1–1.5
mm, pubescent, entire, apex obtuse; petiole 4–6 cm, pubescent;leaf blade abaxially purplish, adaxially purplish when young,becoming purplish green when old, ovate or ovate-lanceolate,
7–12 × 5–7 cm, abaxially pubescent on veins, adaxially pubescent on veins, basal veins 5, lateral veins 5 or 6 pairs, veinlets
usually raised, base cordate, notch obtuse, rarely acute, base of
midlobes broad, rarely constricted, margin irregularly 12–16toothed on each side, teeth sharp, apex acute or acuminate.
Panicle leaf-opposed, 6–8 cm, basal branches poorly developed;
peduncle 1–2 cm, pubescent. Pedicel 2–3 mm, subglabrate.
Buds obovoid, 1.5–2 cm, apex rounded. Calyx ca. 2 mm, entire.
Filaments filiform, ca. 0.8 mm; anthers yellow, elliptic, ca. 0.4
mm. Fl. Apr–May.
● Forests, shrublands, hillsides, grasslands, field margins; ca.
1000 m. Jiangxi, Zhejiang.

20. Vitis ruyuanensis C. L. Li, Chin. J. Appl. Environ. Biol. 2:
244. 1996.

Ovary conical; style slender. Berry globose, 6–8 mm in diam.
Seeds obovoid, base rostrate, apex retuse, chalazal knot rounded, ventral holes furrowed from base nearly to apex. Fl. Apr–
May, fr. Jun–Jul.
● Shrublands on hillsides; ca. 200 m. Guangdong.

21. Vitis mengziensis C. L. Li, Chin. J. Appl. Environ. Biol. 2:245. 1996.
蒙自葡萄 meng zi pu tao
Branchlets terete, glabrous, glaucous; tendrils unbranched.
Leaves simple; stipules caducous; petiole 4.5–10 cm, glabrous;
leaf blade oval, 11–13 × 7–11 cm, glabrous, basal veins 6 or 7,
lateral veins 5 or 6 pairs, veinlets slightly raised abaxially, basedeeply cordate, notch narrow to acute, two sides close together,margin irregularly 31–39-toothed on each side, apex mucronulate. Panicle leaf-opposed, ca. 12 cm; peduncle subglabrate.
Pedicel 2–2.5 mm, glabrous. Buds obovoid, 2–2.5 mm. Calyx
glabrous, margin undulate. Petals oblong, 1.5–2 mm. Filaments
filiform, slightly flat, 1–1.5 mm; anthers yellow, oblong, ca. 0.8
mm. Ovary conical; style short.
● Forests; ca. 1600 m. Yunnan.

22. Vitis fengqinensis C. L. Li, Chin. J. Appl. Environ. Biol. 2:245. 1996.
凤庆葡萄 feng qing pu tao
Branchlets terete, with sparse, white arachnoid tomentum.
Leaves simple; stipules brownish, oblong, 2–2.5 × 1.8–2 mm,
membranous, with sparse white tomentum, apex acuminate;petiole 2–6 cm, with sparse, white arachnoid tomentum; leaf
blade oval or ovate-elliptic, 4–9.5 × 3.5–7 cm, with sparse,
white arachnoid tomentum on veins, basal veins 5, lateral veins
4 or 5 pairs, veinlets raised, base cordate or subcordate, notchobtuse, margin irregularly 17–29(–32)-toothed on each side,
apex acuminate or mucronulate. Panicle sparse, basal branches
well developed, axis sparsely lanate and pubescent; peduncle 2–
6 cm, with sparse, white arachnoid tomentum. Pedicel 2–3 mm,
glabrous. Buds obovoid, 1.8–2 mm, apex rounded. Calyx shallow and saucer-shaped, margin undulate. Filaments filiform,
1.3–1.5 mm; anthers yellow, oblong. Pistil entirely abortive inmale flowers. Fr. Jun.
● Shrublands on hillsides; ca. 2000 m. Yunnan.

乳源葡萄 ru yuan pu tao

23. Vitis hancockii Hance, J. Bot. 20: 4. 1882.

Plants polygamo-dioecious. Branchlets terete, slender,
with inconspicuous longitudinal ridges, glabrous; tendrils
unbranched. Leaves simple, undivided or 3–5-lobed; stipules
caducous; petiole 1.5–2 cm, glabrous; leaf blade abaxially purplish, sagittate or ovate-hastate, 5–8 × 3–4 cm, abaxially pubescent on veins, adaxially glabrous or with sparse very short hairs,
basal veins 5, lateral veins 5 or 6 pairs, veinlets conspicuous but
not raised, base cordate, notch acute or obtuse, margin irregularly finely 7–10-toothed on each side, apex acuminate. Panicle
leaf-opposed, 7–9 cm, basal branches undeveloped; peduncle
1.5–2 cm, subglabrate. Pedicel 2–3 mm, subglabrate. Buds obovoid, 1.5–2 mm, apex rounded. Calyx ca. 2 mm, entire. Filaments filiform, ca. 1 mm; anthers yellow, elliptic, ca. 0.5 mm.

菱叶葡萄 ling ye pu tao
Vitis fagifolia Hu; V. wentsaiana P. L. Chiu.
Plants polygamo-dioecious. Branchlets terete, with longitudinal ridges, densely brown villous; tendrils bifurcate or unbranched, sparsely brown pilose. Leaves simple, undivided,
rarely 3-lobed; stipules brown, triangular-lanceolate, 2–4 × 1–
1.5 mm, membranous, glabrous or base sparsely ciliate, margin
entire, apex acuminate; petiole 0.2–3 cm, brown villous; leaf
blade rhombic-ovate or elliptic, 3.5–13 × 2–7 cm, abaxiallysparsely brownish pilose, adaxially sparsely pubescent only on
midvein, basal veins 3, lateral veins 3–5 pairs, veinlets slightly
raised abaxially, inconspicuous adaxially, base usually asym-

Flora of China 12: 210–222. 2007.

metric, truncate or broadly so, base of lower leaves rarely subrounded, margin 6–12-toothed on each side, teeth large andsharp, rarely obtuse, apex acute. Panicle leaf-opposed, loose,
2.5–5.5 cm, basal branches undeveloped; peduncle 1–2 cm,
densely brownish villous. Pedicel 1.5–2 mm, glabrous. Buds
oval, 1.2–1.5 mm, apex rounded. Calyx 0.1–0.2 mm, glabrous,
entire. Filaments filiform, 0.5–1 mm; anthers yellow, oval orovate-elliptic, 0.5–0.6 mm. Ovary rounded; style short; stigma
expanded. Berry globose, 6–8 mm in diam. Seeds obovoid,
apex retuse, chalazal knot elliptic, ventral holes elliptic andbroad, furrowed upward 1/4 from base. Fl. Apr–May, fr. May–
Jun.
● Forests, shrublands, hillsides; 100–600 m. Anhui, Fujian, Jiangxi, Zhejiang.

24. Vitis tsoi Merrill, Lingnan Sci. J. 11: 101. 1932 [“tsoii”].
狭叶葡萄 xia ye pu tao
Vitis embergeri Galet.
Branchlets terete, with longitudinal ridges, densely pubescent; tendrils unbranched. Leaves simple; stipules brown, lanceolate, 1–2 × 0.3–0.5 mm, membranous, glabrous; petiole 1–2
cm, densely pubescent; leaf blade ovate-lanceolate or triangular-ovate, 3.5–9 × 1.5–4 cm, abaxially pubescent on veins,
adaxially sparsely pubescent on veins, basal veins 5, outermost
pair of veins usually on a line, lateral veins 5 or 6 pairs, basesubtruncate or rounded, margin 10–15-toothed on each side,
teeth sharp and fine, apex acute. Panicle leaf-opposed, 2–6 cm,
narrow, basal branches undeveloped. Pedicel 1.5–2.5 mm, subglabrate. Buds obelliptic, ca. 1.5 mm, apex rounded. Calyx subentire. Filaments filiform, ca. 0.8 mm; anthers yellow, elliptic,
ca. 0.5 mm. Ovary conical; style short. Berry purple-black atmaturity, globose, 5–8 mm in diam. Seeds obelliptic, apex
rounded, chalazal knot elliptic, ventral holes obelliptic andbroad, furrowed upward from base to apex. Fl. Apr–May, fr.
Jun–Sep.
● Forests, shrublands, hillsides; 300–700 m. Fujian, Guangdong,
Guangxi.

25. Vitis flexuosa Thunberg, Trans. Linn. Soc. London 2: 332.
1794.
葛藟葡萄 ge lei pu tao
Vitis cavaleriei H. Léveillé & Vaniot; V. flexuosa var.
chinensis Veitch; V. flexuosa f. parvifolia (Roxburgh) Planchon; V. flexuosa var. parvifolia (Roxburgh) Gagnepain; V.
flexuosa var. wallichii (Candolle) F. S. Wang; V. parvifolia
Roxburgh; V. purani Buchanan-Hamilton ex D. Don; V. vulpina Linnaeus var. parvifolia (Roxburgh) Regel; V. wallichii
Candolle.
Branchlets terete, with longitudinal ridges, with sparsearachnoid tomentum when young, becoming glabrescent; tendrils bifurcate. Leaves simple; stipules caducous; petiole 1.5–7
cm, with sparse arachnoid tomentum or subglabrate; leaf blade
ovate, triangular-ovate, oval, or ovate-elliptic, 2.5–12 × 2.3–10
cm, abaxially with sparse arachnoid tomentum when young,
eventually glabrescent, basal veins 5, lateral veins 4 or 5 pairs,
veinlets inconspicuous, base slightly subcordate or subtruncate,rarely cordate, often asymmetrical, leaves with cordate baseswith obtusely angled notch, margin with 5–12 slightly irregularteeth on each side, apex acute or acuminate. Panicle leaf-opposed, loose, 4–12 cm, basal branches well developed, or shortand slender; peduncle 2–5 cm, with sparse arachnoid tomentum
or nearly glabrescent. Pedicel 1.1–2.5 mm, glabrous. Buds obovoid, 2–3 mm, apex rounded and subtruncate. Calyx glabrous,
undulately lobed. Filaments filiform, 0.7–1.3 mm; anthers
yellow, oval, 0.4–0.6 mm. Pistil abortive in male flowers; ovary
oval; style short. Berry globose, 8–10 mm in diam. Seeds
obovoid, apex subrounded, chalazal knot elliptic, raphe slightly
raised and surface smooth, ventral holes furrowed upward 1/4
from base. Fl. Mar–May, fr. Jul–Nov. 2n = 38.
Forests, shrublands, hillsides, valleys, meadows, fields; 100–2300.
Anhui, Fujian, Gansu, Guangdong, Guangxi, Guizhou, Henan, Hunan,
Jiangsu, Jiangxi, Shaanxi, Shandong, Sichuan, Taiwan, Yunnan, Zhejiang [India, Japan, Laos, Nepal, Philippines, Thailand, Vietnam].

26. Vitis amurensis Ruprecht, Bull. Cl. Phys.-Math. Acad.
Imp. Sci. Saint-Pétersbourg 15: 266. 1857.
山葡萄 shan pu tao
Branchlets terete, glabrous, with sparse arachnoid tomentum when young; tendrils 2- or 3-branched. Leaves simple;
stipules brown, 4–8 × 3–5 mm, membranous, entire, apex obtuse; petiole 4–14 cm, with sparse arachnoid tomentum when
young, eventually glabrate; leaf blade broadly oval, 6–24 × 5–
21 cm, adaxially with sparse arachnoid tomentum when young,then glabrescent, basal veins 5, lateral veins 5 or 6 pairs, veinsraised abaxially, conspicuous or slightly impressed adaxially,
abaxial veinlets conspicuous, ± raised, usually pubescent or
glabrescent, base cordate, notch rounded to obtuse, base oflobes usually constricted or mixed with broad ones, notch
rounded, rarely acute or obtuse, margin 28–36-toothed on eachside, teeth sharp, slightly irregular, apex acute or acuminate.
Panicle leaf-opposed, loose, 5–13 cm, basal branches welldeveloped, with sparse arachnoid tomentum when young,
eventually glabrescent. Pedicel 2–6 mm, glabrous. Buds obovoid, 1.5–3 mm, apex rounded. Calyx 0.2–0.3 mm, glabrous,
subentire. Petals calyptrate. Filaments filiform, 0.9–2 mm; anthers yellow, ovoid-elliptic, 0.4–0.6 mm, conspicuously shortand abortive in female flowers. Disk 0.3–0.5 mm. Ovary
conical; style obvious, slightly thick at base. Seeds obovoid,
apex retuse, chalazal knot elliptic, ventral holes furrowed upward to middle or near apex from base. Fl. May–Jun, fr. Jul–
Sep.
● Forests, shrublands, hillsides, valleys; 100–2100 m. Anhui, Hebei, Heilongjiang, Jilin, Liaoning, Shandong, Shanxi, Zhejiang.
Vitis amurensis var. funiushanensis F. S. Wang (J. Trop. Subtrop.
Bot. 8: 5. 2000), described from Henan, could not be treated here because no material was seen by the authors.

1a. Berry 1–1.5 cm in diam. ...................... 26a. var. amurensis
1b. Berry 0.8–1 cm in diam. ......................... 26b. var. dissecta
26a. Vitis amurensis var. amurensis
山葡萄(原变种) shan pu tao (yuan bian zhong)Flora of China 12: 210–222. 2007.

Vitis vinifera Linnaeus var. amurensis (Ruprecht) Regel.
Leaves 3(or 5)-lobed or cleft, or undivided. Berry 1–1.5
cm in diam. Fl. May–Jun, fr. Jul–Sep. 2n = 38*.
● Forests or shrublands in valleys, hillsides; 200–2100 m. Anhui(Jinzhai), Hebei, Heilongjiang, Jilin, Liaoning, Shandong, Shanxi, Zhejiang.

26b. Vitis amurensis var. dissecta Skvortsov, Chin. J. Sci.
Arts. 15: 200. 1931.
深裂山葡萄 shen lie shan pu tao
Vitis amurensis var. yanshanensis D. Z. Lu & H. P. Liang;
V. baihuashanensis M. S. Kang & D. Z. Lu.
Leaves 3–5-parted. Berry 0.8–1 cm in diam. Fl. May–Jun,
fr. Jul–Sep.
● 100–200 m. Hebei, Heilongjiang, Jilin, Liaoning.

rarely acute, margin 9–19-toothed on each side, teeth sharp,
apex acute or acuminate. Panicle leaf-opposed, 4–14 cm, basal
branches developed; peduncle 1–2 cm, with gray or brownarachnoid tomentum. Pedicel 1–3 mm, glabrous. Buds obovoid
or elliptic, 1.5–2 mm, apex rounded. Calyx ca. 0.1 mm, subentire. Filaments filiform, 1–1.2 mm; anthers yellow, elliptic or
broadly so, ca. 0.5 mm. Ovary oval; style short. Berry purpleblack at maturity, globose, 1–1.3 cm in diam. Seeds obovoid,
apex rounded, chalazal knot rounded, ventral holes furrowed
upward ca. 1/4 from base. Fl. Apr–Jul, fr. Jun–Oct.
Forests, shrublands, hillsides, valleys; 100–3200 m. Anhui,
Chongqing, Fujian, Gansu, Guangdong, Guangxi, Guizhou, Hebei, Henan, Hubei, Hunan, Jiangsu, Jiangxi, Shaanxi, Shandong, Shanxi, Sichuan, Xizang, Yunnan, Zhejiang [Bhutan, India, Nepal].

1a. Leaves oval, ovate-oblong, or ovatequinquangular ................................... 28a. subsp. heyneana
1b. Leaves usually 3-lobed to cleft ........... 28b. subsp. ficifolia

27. Vitis vinifera Linnaeus, Sp. Pl. 1: 202. 1753.

28a. Vitis heyneana subsp. heyneana

葡萄 pu tao

毛葡萄(原亚种) mao pu tao (yuan ya zhong)Branchlets terete, with longitudinal ridges, glabrous orsparsely pilose; tendrils bifurcate. Leaves simple, conspicuously
3–5-lobed or cleft; stipules caducous; petiole 4–9 cm, nearly
glabrous; leaf blade oval, 7–18 × 6–16 cm, basal veins 5, lateralveins 4 or 5 pairs, veinlets inconspicuously raised, base deeply
cordate, notch rounded, 2 sides usually overlapping to nearlyso, margin 22–27-toothed on each side, teeth large, irregular,
and sharp, glabrous or sparsely pilose, apex of midlobes acute.
Panicle sparse or dense, opposite to leaves, basal branches welldeveloped, 10–20 cm; peduncle 2–4 cm, glabrous or withsparse arachnoid tomentum. Pedicel 1.5–2.5 mm, glabrous.
Buds obovoid, 2–3 mm, apex rounded. Calyx glabrous, undulate. Filaments filiform, 0.6–1 mm; anthers yellow, oval, 0.4–
0.8 mm. Pistil entirely abortive in male flowers; ovary oval;
style short; stigma expanded. Berry globose or elliptic, 1.5–2
cm in diam. Seeds obovoid to obelliptic, apex subrounded,chalazal knot elliptic, raphe slightly raised, ventral holes
broadly furrowed upward ca. 1/4 from base. Fl. Apr–May, fr.
Aug–Sep. 2n = 38*.

Vitis ficifolia Bunge var. pentagona Pampanini; V.
kelungensis Momiyama; V. lanata Roxburgh; V. pentagona
Diels & Gilg; V. pentagona var. honanensis Rehder; V. quinquangularis Rehder; V. thunbergii Siebold & Zuccarini var.
yunnanensis Planchon.

Cultivated throughout China for grapes and wine-making [nativeto SW Asia and SE Europe].

28. Vitis heyneana Roemer & Schultes, Syst. Veg. 5: 318.
1819.
毛葡萄 mao pu tao
Plants polygamo-dioecious. Branchlets terete, with longitudinal ridges, gray or brown lanate; tendrils bifurcate. Leaves
simple; stipules brown, ovate-lanceolate, 3–5 × 2–3 mm, membranous, entire, apex acuminate, rarely obtuse; petiole 2.5–6
cm, with dense arachnoid tomentum; leaf blade sometimes 3lobed to -cleft, 4–12 × 3–8 cm, abaxially densely grayish or
brown tomentose, gradually becoming less so, abaxial veinstomentose, sometimes pilose or lanate, adaxially with sparsearachnoid tomentum when young, then glabrate, adaxial veinsglabrous or sometimes sparsely pubescent, basal veins 3–5,
lateral veins 4–6 pairs, base cordate to subcordate, notch obtuse,

Leaves oval, ovate-oblong, or ovate-quinquangular. Fl.
Apr–Jun, fr. Jun–Oct.
Forests, shrublands, hillsides, valleys; 100–3200 m. Anhui,
Chongqing, Fujian, Gansu, Guangdong, Guangxi, Guizhou, Henan,
Hubei, Hunan, Jiangxi, Shaanxi, Shandong, Shanxi, Sichuan, Xizang,
Yunnan, Zhejiang [Bhutan, India, Nepal].

28b. Vitis heyneana subsp. ficifolia (Bunge) C. L. Li, Chin.
J. Appl. Environ. Biol. 2: 250. 1996.
桑叶葡萄 sang ye pu tao
Vitis ficifolia Bunge, Enum. Pl. China Bor. 12. 1833; V.
labrusca Linnaeus var. ficifolia (Bunge) Regel.
Leaves usually 3-lobed to cleft, and mixed with undivided
ones. Fl. May–Jul, fr. Jul–Sep. 2n = 38*.
● Forests, shrublands, hillsides; 100–1300 m. Hebei, Henan,
Jiangsu, Shaanxi, Shandong, Shanxi.

29. Vitis retordii Romanet du Caillaud ex Planchon in A. Candolle & C. Candolle, Monogr. Phan. 5: 613. 1887.
绵毛葡萄 mian mao pu tao
Vitis hekouensis C. L. Li.
Plants polygamo-dioecious. Branchlets terete, with longitudinal ridges, densely brownish villous when young, eventually glabrescent; tendrils bifurcate. Leaves simple; stipules
brown, ovate-lanceolate, 3–5 × 2–3 mm, membranous, subglabrate, apex acuminate, caducous; petiole 1.5–9 cm, densely
pubescent with brown and arachnoid hairs; leaf blade oval orovate-elliptic, 6–15 × 4–11 cm, abaxially tomentose with brownand villous hairs, adaxially densely pubescent, basal veins 5,

Flora of China 12: 210–222. 2007.

lateral veins 4 or 5 pairs, veinlets tomentose and raised abaxially, raised and pubescent adaxially, base deeply cordate, notchwith an acute angle, rarely overlapping, margin 19–43-toothedon each side, teeth sharp. Panicle leaf-opposed, loose, 6–10 cm,
basal branches well developed; peduncle 1.2–2.5 cm, usuallybrown tomentose. Pedicel 1–1.5 mm, glabrous. Buds obelliptic,
1.2–1.5 mm, apex rounded. Calyx cupular, ca. 1.5 mm, glabrous, subentire. Filaments filiform, 1–1.2 mm; anthers yellow,
oblong, ca. 0.5 mm. Ovary oval; style short. Berry globose, ca.
8 mm in diam. Seeds obelliptic, apex rounded, chalazal knot
ovate-elliptic, ventral holes furrowed upward ca. 1/3 from base.
Fl. May, fr. Jun–Jul.
Forests in valleys, shrublands, hillsides; 200–1000. Guangdong,
Guangxi, Guizhou, Hainan [Laos, Vietnam].

30. Vitis menghaiensis C. L. Li, Chin. J. Appl. Environ. Biol.
2: 250. 1996.
勐海葡萄 meng hai pu tao
Branchlets terete, densely ferruginous pilose. Leaves simple; stipules brown, ovate-elliptic, 4–6 × 2–3 mm, membranous,
apex obtuse; petiole 3–5.5 cm, densely ferruginous pilose; leaf
blade suborbicular or quinquangularly orbicular, 9–11 × 8–10
cm, abaxially with dense rubiginous arachnoid tomentum,
adaxially densely pubescent, basal veins 5–7, lateral veins 5 or
6 pairs, veins densely pubescent with spreading hairs, basedeeply cordate, notch narrow, 2 sides close together or overlapping, margin 28–47-toothed on each side, teeth fine andsharp, apex acute or rounded, rarely retuse. Panicle loose, ca. 12cm, basal branches undeveloped; peduncle ca. 2.5 cm, densely
ferruginous pilose. Pedicel 1–2 mm, subglabrate. Buds obovoid, 1.5–2 mm, apex rounded. Calyx ca. 0.2 mm, undulate.
Filaments filiform, slightly flat, 1–1.2 mm; anthers yellow,
ovoid-oblong. Ovary abortive in male flowers. Fl. May.
● Mixed forests; 1500–1600 m. Yunnan.

31. Vitis longquanensis P. L. Chiu, Bull. Bot. Res., Harbin10(3): 41. 1990.
龙泉葡萄 long quan pu tao
Branchlets terete, with longitudinal ridges, white lanate
when young, then glabrescent. Leaves of two types, those of
upper branches sometimes 3–5-divided, notch rounded, mostleaves of lower branches undivided, simple; stipules caducous;
petiole 2–5.5 cm, densely light brown lanate; leaf blade adaxially greenish, oval, triangular-ovate, or triangular-lanceolate,
4–12 × 4–10 cm, abaxially densely light brown lanate, rarely
hairs falling off and only veins tomentose, adaxially with sparsearachnoid tomentum when young, then glabrate, basal veins 5,
outermost pair of veins on a line and bifurcate at 5–9 mm frombase, lateral veins 3 or 4 pairs, veinlets tomentose and raised
abaxially, conspicuously raised adaxially, base subcordate or
subtruncate, notch very short, two sides nearly overlapping,margin with fine teeth, apex acute or acuminate. Panicle leafopposed, loose, basal branches well developed; peduncle 3–9
cm, with sparse arachnoid tomentum. Pedicel 1–4 mm, glabrous. Berry purple-black and glaucous at maturity, globose, 5–
6 mm in diam. Seeds obovoid, apex retuse, chalazal knot
rounded, ventral holes narrow and oblong. Fr. Jul–Sep.

● Forests and shrublands in valleys, roadsides; 700–1300 m. Fujian, Jiangxi, Zhejiang.

32. Vitis bellula (Rehder) W. T. Wang, Acta Phytotax. Sin.
17(3): 74. 1979.
美丽葡萄 mei li pu tao
Branchlets terete, slender, with longitudinal ridges,
sparsely white lanate; tendrils unbranched or mixed with bifurcate ones. Leaves simple; stipules green-brown, ca. 0.8 × 0.7mm, submembranous, glabrous, apex obtuse; petiole 1–3 cm,
with sparse arachnoid tomentum; leaf blade 3–7 × 2–4 cm,
abaxially with dense gray-white or gray-brown arachnoid tomentum, adaxially subglabrate, basal veins 3, lateral veins 4 or
5 pairs, veinlets tomentose and raised abaxially, not raised
adaxially, base subcordate, subtruncate, or subrounded, notch
very short, margin 7–10-toothed on each side, teeth fine andsharp, apex acute or acuminate. Panicle leaf-opposed, terete,basal branches undeveloped; peduncle 0.5–1.2 cm. Pedicels 2–3mm, slender, glabrous. Buds elliptic or obelliptic, 1.5–1.8 mm,
apex rounded. Calyx glabrous, teeth inconspicuous. Petals
ovate-elliptic, 1.2–1.6 mm. Filaments filiform, ca. 1.2 mm;anthers yellow, oblong, ca. 0.5 mm. Disk well developed andpistil entirely abortive in male flowers. Berry purple-black andglaucous at maturity, globose, 6–10 mm in diam. Seeds obovoid, apex rounded and retuse, chalazal knot rounded, ventral
holes furrowed from base upward nearly to tip. Fl. May–Jun, fr.
Jun–Aug.
● Forests, shrublands, hillsides, cliffs; 400–1600 m. Guangdong,
Guangxi, Hubei, Hunan, Sichuan.

1a. Leaves oval or ovate-elliptic .................... 32a. var. bellula
1b. Leaves mostly ovate-triangular ............. 32b. var. pubigera
32a. Vitis bellula var. bellula
美丽葡萄(原变种) mei li pu tao (yuan bian zhong)Vitis pentagona Diels & Gilg var. bellula Rehder in Sargent, Pl. Wilson. 3: 428. 1917; V. quinquangularis Rehder var.
bellula (Rehder) Rehder.
Leaves oval or ovate-elliptic. Peduncle with sparse arachnoid tomentum. Fl. May–Jun, fr. Jul–Aug.
● Forests, shrublands, hillsides; 1300–1600 m. Hubei, Sichuan.

32b. Vitis bellula var. pubigera C. L. Li, Chin. J. Appl. Environ. Biol. 2: 251. 1996.
华南美丽葡萄 hua nan mei li pu tao
Leaves mostly ovate-triangular. Inflorescence axis pilose.
Fl. May, fr. Jun–Aug.
● Forests, shrublands, cliffs; 400–1500 m. Guangdong, Guangxi,
Hunan.

33. Vitis bashanica P. C. He, Acta Univ. Agric. Bor.-Occid.
23(5): 121. 1995.
麦黄葡萄 mai huang pu tao
Branchlets terete, slender, with longitudinal ridges, pilose
and lanate with white hairs; tendrils unbranched or bifurcate.
Leaves simple; stipules caducous; petiole 1.6–3 cm, pilose andFlora of China 12: 210–222. 2007.

lanate; leaf blade oval or ovate-elliptic, 5.5–6.9 × 3.6–4.8 cm,
abaxially densely gray-white tomentose, adaxially sparsely pubescent on veins, basal veins 5, lateral veins 3 or 4 pairs, veinlets inconspicuously raised, base truncate, margin 7–12-toothedon each side, teeth fine and sharp, apex acuminate. Panicle ca.
3.6 cm; peduncle ca. 1.2 cm. Berry purple-black and glaucous
at maturity, globose, ca. 7 mm in diam. Fr. Aug.
● About 300 m. Shaanxi (Baiheyinfang).

34. Vitis hui W. C. Cheng, Contr. Biol. Lab. Sci. Soc. China,
Bot. Ser. 10: 77. 1935.
庐山葡萄 lu shan pu tao
Plants polygamo-dioecious. Branchlets terete, with longitudinal ridges, densely pubescent and sparsely lanate; tendrils
unbranched or mixed with bifurcate ones. Leaves simple; stipules ovate-elliptic, 2–3 × 1.5–2 mm, subglabrate, subentire,
apex obtuse; petiole 5–10 mm, densely pubescent and lanate;leaf blade oval or ovate-elliptic, 3–5.5 × 2–4 cm, abaxiallydensely lanate with gray hairs and pubescent with short hairs,
adaxially densely pubescent and sparsely lanate, basal veins 5,
lateral veins 4 or 5 pairs, veins densely pubescent and withsparse arachnoid tomentum, veinlets densely tomentose and pubescent abaxially, inconspicuous adaxially, base cordate, notch
acute, margin with sharp teeth, apex acute or acuminate. Panicle
leaf-opposed, 2–8.5 cm, basal branches short; peduncle 0.5–1.5
cm, densely pubescent. Pedicels 1–1.5 mm, densely pubescent.
Buds obovoid, 1.5–2 mm, apex rounded. Calyx ca. 0.2 mm,
glabrous, 5-lobed. Filaments filiform, 1–1.2 mm; anthers
yellow, oblong, ca. 0.5 mm. Ovary abortive in male flowers. Fl.
May, fr. Jul.
● Shrublands, fields; 100–200 m. Jiangxi, Zhejiang.

35. Vitis sinocinerea W. T. Wang, Acta Phytotax. Sin. 17(3):75. 1979.
小叶葡萄 xiao ye pu tao
Vitis thunbergii Siebold & Zuccarini var. cinerea Gagnepain; V. thunbergii var. taiwaniana F. Y. Lu.
Branchlets terete, with longitudinal ridges, sparsely pubescent with arachnoid tomentum; tendrils unbranched or bifurcate. Leaves simple, 3-lobed or inconspicuously divided; stipules brownish, ovate-lanceolate, ca. 2 × 1 mm, membranous,
subglabrate, apex obtuse or acuminate; petiole 1–3 cm, densely
pubescent; leaf blade oval, 3–8 × 3–6 cm, abaxially with densebrown and arachnoid tomentum, adaxially densely pubescent or
glabrescent, basal veins 5, lateral veins 3 or 4 pairs, veinsdensely pubescent and sparsely lanate, base subcordate or subtruncate, margin 5–9-toothed on each side, apex acute. Panicle leaf-opposed, small, 3–6 cm, slender, basal branches undeveloped; peduncle 1.5–2 cm, densely pubescent. Pedicels 1.5–2mm, subglabrate. Buds obovoid-elliptic, 1.5–2 mm, apex
rounded. Calyx cupular, glabrous, subentire. Petals calyptrate.
Filaments filiform, ca. 1 mm; anthers yellow, oblong, ca. 0.5
mm. Pistil abortive in male flowers. Berry purple-black at maturity, 6–10 mm in diam. Seeds obovoid, apex retuse, chalazal
knot elliptic, ventral holes furrowed upward 1/4–1/3 from base.
Fl. May–Jun, fr. Jul–Oct.

● Forests, shrublands, hillsides; 200–2800 m. Fujian, Hubei, Hunan, Jiangsu, Jiangxi, Taiwan, Yunnan, Zhejiang.

36. Vitis bryoniifolia Bunge, Enum. Pl. China Bor. 11. 1833[“bryoniaefolia”].
蘡薁 ying yu
Plants polygamo-dioecious. Branchlets terete, with longitudinal ridges, densely lanate when young, becoming sparsely
so; tendrils bifurcate. Leaves simple; stipules brownish, oval orovate-lanceolate, 3.5–8 × 2.5–4 mm, membranous, glabrous or
subglabrate, entire, apex obtuse; petiole 0.5–4.5 cm, densely
lanate, pubescent, or tomentose, becoming sparsely so; leaf
blade ovate, 2.5–8 × 2–5 cm, basal veins 5, lateral veins 4–6
pairs, veinlets inconspicuous or slightly raised adaxially, basecordate or deeply so, notch rounded, margin with 9–16 obtuse
teeth on each side or pinnatifidly divided. Panicle leaf-opposed,
basal branches well developed or sometimes reduced to tendrils, rarely narrow and branches undeveloped; peduncle 0.5–
2.5 cm, with arachnoid tomentum when young, becoming
sparsely so. Pedicels 1.5–3 mm, glabrous. Buds obelliptic or
globose, 1.5–2.2 mm, apex rounded. Calyx ca. 0.2 mm, glabrous, subentire. Filaments filiform, 1.5–1.8 mm; anthers yellow, oblong, 0.4–0.5 mm. Ovary elliptic-ovoid; style short,
slender; stigma expanded. Berry purple-red at maturity, 5–8
mm in diam. Seeds obovoid, apex retuse, chalazal knot
rounded or elliptic, ventral holes narrow, furrowed upward ca.
3/4 from base. Fl. Apr–Aug, fr. Jun–Oct.
● Forests in valleys, shrublands, streamsides, fields; 100–2500 m.
Anhui, Fujian, Guangdong, Guangxi, Hebei, Hubei, Hunan, Jiangsu,
Jiangxi, Shaanxi, Shandong, Shanxi, Sichuan, Yunnan, Zhejiang.

1a. Leaves simple, 3–5(–7)-parted or lobed,
rarely mixed with undivided ones ..... 36a. var. bryoniifolia
1b. Leaves 3-foliolate or 3–5-parted .............. 36b. var. ternata
36a. Vitis bryoniifolia var. bryoniifolia
蘡薁(原变种) ying yu (yuan bian zhong)Vitis adstricta Hance; V. bryoniifolia var. mairei (H.
Léveillé) W. T. Wang; V. bryoniifolia var. multilobata S. Y.
Wang & Y. H. Hu; V. flexuosa Thunberg var. mairei H. Léveillé; V. novisinensis Vassilczenko; V. thunbergii Siebold &Zuccarini var. adstricta (Hance) Gagnepain; V. thunbergii var.
mairei (H. Léveillé) Lauener.
Leaves simple, 3–5(–7)-parted or lobed, rarely mixed with
undivided ones; leaf blade abaxially with dense arachnoid
tomentum and short hairs, then becoming sparsely so, rarely
mostly pubescent with short hairs and only sparsely lanate, baseconstricted to rounded, apex of midlobes acute or acuminate.
● Forests in valleys, shrublands, streamsides, fields; 100–2500 m.
Anhui, Fujian, Guangdong, Guangxi, Hebei, Hubei, Hunan, Jiangsu,
Jiangxi, Shaanxi, Shandong, Shanxi, Sichuan, Yunnan.

36b. Vitis bryoniifolia var. ternata (W. T. Wang) C. L. Li,
Chin. J. Appl. Environ. Biol. 2: 253. 1996.
三出蘡薁 san chu ying yu
Vitis adstricta var. ternata W. T. Wang, Acta Phytotax.
Sin. 17(3): 76. 1979.

Flora of China 12: 210–222. 2007.

Leaves 3-foliolate or 3–5-parted; leaf blade abaxiallysparsely lanate.
● Zhejiang.

37. Vitis lanceolatifoliosa C. L. Li, Chin. J. Appl. Environ.
Biol. 2: 253. 1996 [“laceolatifoliosa” & “lanceolatifoilosa”].
鸡足葡萄 ji zu pu tao
Vitis piasezkii Maximowicz var. angusta W. T. Wang.
Branchlets terete, with longitudinal ridges, densely rubiginous lanate; tendrils bifurcate. Leaves palmate, 3–5-foliolate;
stipules brownish, elliptic, 1.8–2.5 × 1–1.3 mm, membranous,
glabrous or subglabrate, entire, apex acute; petiole 3–5 cm, with
dense brown and arachnoid tomentum; leaflets abaxially with

dense brown arachnoid tomentum, adaxially with sparse arachnoid tomentum when young, then glabrate, only midvein pilose
with very short hairs, lateral veins 3–5 pairs, veinlets inconspicuous; central leaflet lanceolate, rarely oblong or obovatelanceolate, 3.5–9 × 1.5–2.5 cm, base truncate, margin with 5 or
6 sharp teeth on each side, apex acuminate; lateral leaflets
ovate-lanceolate, 3–8 × 1.2–3 cm, base asymmetric, obliquely
cuneate or rounded, margin finely 6–11-toothed, apex acuminate. Panicle leaf-opposed, loose, basal branches well developed; peduncle 4–8 cm, with dense rubiginous arachnoid tomentum. Pedicel 1–1.3 mm, subglabrate. Buds obovoid, 1.7–
2.8 mm, apex rounded. Calyx teeth inconspicuous. Ovary oval;
style short. Berry globose, 8–10 mm in diam. Seeds obovoid,
apex subrounded, chalazal knot elliptic, raphe raised. Fl. May,
fr. Aug–Sep.
● Forests or shrublands by streams, hillsides; 600–800 m. Guangdong, Hunan, Jiangxi.

Flora of China 12: 210–222. 2007.