fromof Crane Flies
SpeciesMountains
Tipulidae),(D?ptera:theUndescribed
HimalayaP. AlexanderCharles

Abstract:

Six

new

species

Sikkim.

These

are

Trentepohlia

fordiperone;

T.

(T.)

of

AugustPublication

craneeriopterine

5, 1968

are

flies

and
all from Assamdescribed,
Teucholabis(Teucholabis)and B. obtusistyla.

fuscogenualis,
B. latibasis;(Mongoma)
Baeouraaka;

eremnopoda;

01002

Massachusetts

Amherst,Received

XVII1

in the Journal of the New
Part XVI of this series of papers was published1968.
York EntomologicalThe
76:
present report continues the
66-71,
Society,in Sikkim and Assam
the
made
Dr.
Fernand
Schmidofrichcollections
bystudy
in 1959 and 1961 and concerns species of the tribe Eriopterini, vastly developed
the region. I again
throughout
Schmid for these materials.
Trentepohlia

thanksexpress my(Mongoma)large

female:palpi
setae.

10 mm;

wing

andantennaeyellow.

Head

uniformly
pale yellow,
coxae and trochanters
lightLegs withtibiae withrather broadly
femorablack,
tarsal segments
passing
rowly so; outer
the

with

femorasparselongerthe outercostalveins

deeper yellow,
Certainlongitudinal

anddistalsections

light

into

2?3 entire

butcontiguousor

midlength.
Abdomendull

yellow.

shortly

1.8 mm.

setae
borderfused,

Ovipositor

with

gray

heavily

stripesdarker.
slightlyof legs darker

withpruinose,

very

Halt?resyellow.
yellow,tips ofnar?
tips still more

vestiturebrown;
over mostof theblackened,of legs shortlength.
brownishmorethickened,
numerous
withcord
beyondsparse on_?i+2, morevery pale; m-cu
shortly

at margin

yellow,androstrum,
palpi
black,
tips of femoraveinsyellow;
deeperbrown,

praescutal

andVenation:light

remainder
yellow;base morenarrowly

veins

of R5

pruinose,legs yellow,

10 mm.

aboutlong yellow
Thoraxalmostous,

graythoraxLengthRostrum,

8 mm

(wing

yellow;

n. sp.

fuscogenualis,or more); headlight brown,
almostuniformly
pale yellow;
bases and tips of tibiae moredarkened;
narrowly
wings
cerci of ovipositor
very long and slender.
male:
8-9 mm;antennaabout 7.5-8.5 mm; wing
LengthSizeantennae

to Dr.

and appreciation

and

Wings

inconspicu?
lightyellow;yellow.

black
R4
trichia,
including-f-2+3+_,and on the fused vein R5 plus _1_f1+2.
fork of M ; veins C?_ and 1st A
beforeclosing

the

cerci

very

cell;

long

cell

2nd

and

A

very

slender,

broad

their

tips

at

near

gentlyupcurved.

holotype:

Sikkim, 4,000 feet, May 7, 1959 (Schmid). Allotype,5, pinned with allo?
1, 1959. Paratypes,
9, Teng, Sikkim, 4,600 feet, August2$
type;$, Tung, Sikkim, 4,500 feet, August 2, 1959; IS, Lathong, Sikkim,
6,560 feet, July 26, 1959 (all Schmid).
1

Contribution$, Lingdok,

fromthe Entomological

Laboratory,
61

Universityof Massachusetts.

62

New

most

York

EntomologicalSociety[Vol.

LXXVII

horianaregional species are Trentepohlia(Mongoma)
the
T.
and
Alexander, having
legs uniformly yellow,(M.) flava Brunetti, withthe femoral tips narrowly and inconspicuously
infuscated, both species having
cell 1st A of the wings narrowly open by the separation of veins Cu\ and 1st A
Thesimilarat margin.

Teucholabis
Generalnotumofcoloration

thoraxarea
medianblackened,of fore femora
yellowed;bases
except

for

dark

brown

the

pale

brown

; sternal

pockets

dististyleconspicuously
male:
about
LengthRostrumdark

Antennaeterminal

stigma;of malesuturefromincisures

yellowed,

abdomen;male

hypopygium

light brown,
well developed
a shorter base.
antennalong,subequal

scutalremainderof head, palpi dark brown.
ones more
the outeroval,elongate,Headdark brown.

withdisk polishedthe central
black,
includingarea before
humeralregion and central
borders,
area andthe medianscutellum
light yellow;
black,

praescutum
the laterallobes

polished
polished
brownishcoxaehypopygium
outer
with

1.8 mm.

about
toblack;
proximalflagellarsegments
shorter than the segments.
oval, verticils

and postnotum
parascutella
stem
Halt?reswithyellow.

yellow;

abdomen

6.8 mm;relatively

segment
Pronotum
Mesonotal
yellow.
toextensions
stripe and broadn. sp.

diperone,the posterior
lobes and post
disk, scutal
yellow,
praescutalof scutum,
scutellum
and pleura
legs blackened,yellow;brownish
unpatterned
yellow,
faintly
tinged withwings

bispinous
8 mm; wingbrown,

brownish(Teucholabis)

black,

black,
of; remainder

Pleuraslightly
paler.
light orange
alllarge knob yellowed.
Legs withblack, bases of fore femora yellow,

pleurotergite
apex of theand trochanters
yellowlegs brownish
more
than
basal
femoradarkened.
third,
Wings
faintlyposterioruniformly
includingveins dark brown.
Venation:brownishSei ending
tinged withyellow,
stigma pale brown;
shorterthan R2]cell 1st M2
far beyond
l_i+2 and I-2+3+4 subequal, muchlong,origin of Rs;
the distal

exceedingAbdomenlight

spicuous
pocketsevenlongitudinal

sectionof Ma.

brown,
on sternite

incisures

yellowed,

five,

including

rows

with

widelyseparatedsclerotizedplate. Malebase

; innerstyle with
before apex with

margins

hypopygium
broad
lateralarea withmediangroups,six
smallersternitebehind;transversebrown
narrowly

Male

with

con?

brown.

setalof microscopic
tubercles,
progressivelyaseven withsetal brushes,
sterniteouter dististylewithdistinctive,
bispinous
hypopygium
rod or ribbon,
lateral arm a long narrowterminating
microscopic

a

dark

from

a shorterin a longspine,

spinulae.

holotype:?\ Teng, Sikkim, 4,600 feet, May
12, 1959 (Schmid).
ofmembersthethegenus having
regional
wings unpatterned or virtually
so include Teucholabis
diversipes Alexander, T. (T.) eremno(Teucholabis)
n.
T.
sp.,
(T.) shanensis Alexander, and T. (T.) solivaga Alexander, all
poda,
Other

with

the details

distinct.
specificTheof coloration, and especiallythe hypopygial
structure, quite
outer dististyle of the present fly has suggested thebispinous

name.

Teucholabis
Headcrossing
posteriorwings(Teucholabis)

eremnopoda,n. sp.

a blackened
withmesonotum
saddleyellow,
praescutum
black;
polished
polished
brownishblack ;and pleurotergite
the posterior; halt?res
yellow
polished
half, pleuraof femora
halflegs withyellowed;
remainingproximalblack,
legs uniformly
yellow,

stigmadark

brown,

Scx

endingabout

opposite

three-fifths

Rs;

abdomen

Alexander:

1969]

March,

tergites
ringed withwithhypopygium
basistyleyellow,

dark

male:

terminating
6 mm.
wing

about
LengthRostrumand palpi
polished

paler;
black,

andtrochanters

legs
and

black,
costalbase

yellowed.

yellowed,sternite

oval,

Antennae

paler basally.
ones
outer

anteriorly.
a majoryellow withblack

smallerblack
suturebeforeregionsaddleyellow;brownishblack.

Wings

stigmasmall,

a

with
dark

faint

yellowtinge,
prearcular
those at wingveins brown,
three-fifths
Ri+2 andRs;

brown;

about
endingoppositeto distal section of Mi+2.

Sct

bases
with
dark
their
brown,
ringedor brownish
with
brown
black. Male
uniformly
setae
and moredelicate
vestiturein centeryellowcoarseincludinglatter directed
inward,six withsternite
about

than
each

black.
half of mediotergite
posterior
coxae
with
brownishblack.
Halt?resshort,
Legsofremainder
basal halvesfemora withyellow,

tergites

five

more

subequalelongate,yellow,
proximaloutersegments morebrownishandacross more

extensively

area;yellow.
middle

long,

the

side,

slightly

segments

polished
the median

and

Sc

Venation:; cell 1st M2

formermale

pruinoseposteriorlegs uniformly
fields clearer
light yellow,

Rz+s+4 subequal
Abdomenfore

yellow;

theflagellar

slightly

Prothorax
Mesonotum
yellow.
the posterior
half of praescutum,a large polishedlobe withscutalPleuraand pleurotergite
polished

more
outer
darkened;segments
uniformly
in a blackened
dististylecomplex.
hook,brown,

in cases

black,

segment

black,

proximalHeadelongate.

7 mm;

63

Flies

Crane

posterior
15 setae

andapicesonpocketon eitherwithfour narrow
blackof pocket
parallelMale
eitherinward.
side, directed
chieflya blackened
twoouter margin
with
hook,

end
on

lines;
inwithbasistyleterminating
hypopygium
or threethat
bearsetae,
axillaryspinouspointsthree
in structure,
with
arms,
includingcomplexbeak

slender.

two

flattened

Dististylesubterminal,
curvedthatliebladesone withsetae on outer margin;
third arm a long
two strong
the shortersuperimposed,on side.
a relativelya smaller
lateralPhallosome
slenderpoint
posteriorspine, withnear base witha veryits outer margin
slenderblade,
large flattenedrod, outer half withfour or five verysetae.

long delicate

Para
12, 1959 (Schmid).
Sikkim, 2,300 feet, August$, Dikchu,
?
10
with
the
fungi.
damagedmostly
by
type,
badly
topotypes,$,
The present fly is most similar to species such as Teucholabis(Teucholabis)

holotype:

annuloabdominalis
blackenedin the extensively
differing most evidently
wings, and in the quite distinct male hypopygium.

Senior-White,legs, unpatterned

Baeouraof bodycoloration

Generaldarkbrown;

n.

aka,

light

legs

with
male
hypopygium
infuscated;strongly
sternalits apex
mediantruncate;
lobe,
glabroussternitesetulae and a few setae;
terminatingeightha boomerang,
third slightly moreouternarrowed,

wings

male:

about

LengthandRostrum
ones

longer,palpi
all withdarkbrown,

orbitsPronotumbrown,

behindstem
brown;

slightlybrownish5-5.2 mm;wing
5.5 mm.
4.2 mm; wing

4 mm;

about

Lengthfemale:brownishlongslightly

black.

conspicuous

sp.

vestiturelong and conspicuous;
brown,
into a depressedninth
tergite produced
dense
with
membranebilobed,deeplyin an obtuse

withmargined
1 mm.
antennaabout

Antennae
verticils,

of

legs

the outersegments
oval,of organlonger. Headblack;
flagellar
those at midlength

paler.

dark brown,
paler. Mesonotum
pretergites
membranedark brown,
Pleuradorsopleuralpaler.
Legs
brightened.
black, base pale, knob vaguely

remainder

likeshapedlobe; dististylea small
blade.
flattenedlightbrown,

outer

tarsalsegments

scutellum
pruinose,
Halt?reswithlight brown.
coxae and trochanterswithsparselydarker;leg vestiture

long

and

64

New

EntomologicalYork[Vol. LXXVIISocietymore
infuscated,stigmalregion vaguelystrongly
Wings
conspicuous.
R.
than
Rs moderately
about
one-halfbrown.
Venation:
longer
long,

dark

veins

darkened;

into a
ninth
with
Male
dark brown.
Abdomencaudally
tergite produced
hypopygium
membranetruncate.
Sternalits apex
medianlonglobe,depressed
glabrous
conspicuous
four oreach lobe withshort dense
surface withand conspicuous,
setulae,bilobed,deeplya conspicuous
lobe. Dististyleinto an obtuse
five long setae. Apex of eighth sternite produced
blade

like

shaped

near

lower

marginslightly widened

a boomerang,
the point

outer

third

slightlywithofnarrowing
into a spur.

and produced

more
a

small

narrowed,
flattenedrounded;obtuselybase of styletip

blade;

S, Lifakpo, Kameng, North East Frontier Agency, Assam, 3,100
3SS.
$. Paratopotypes:
feet, May 29, 1961 (Schmid). Allotopotype:
is moststernite
ninth
and
the
The male hypopygium,
tergiteeighthespeciallyas in Baeoura nilgiriana Alexander and allies, differing in other structures. The

holotype:

BaeouraSizerelatively

sparselypruinosewings withsetae;
borderof malelarge (wingto appear

LengthRostrumaboutyellow,

andto appear

plumbeous,

palpi

wing
brownishnearlythree

conspicuous,
R2+3+1 fromofcoloration

thoraxblackened,borderposterior
restrictedly
coxae
and

with

very

Antennaebrownishsegment.

Head

the

long

heavily

Mesonotum
blackened,yellow.
Pleuraobscure yellow.
scutellum
brownishmembranedorsopleuralbrownishoutspreading

light yellow,
Venation:brown.

brown,

stigmaRi+2yellow;setae.
not
very

Wings

remainder
very

faintly

the moreofverticils

black;

darkofpaler,
trochantersfieldsdarkermm.

black.

times

brownishpretergites

regionmetapleuralbrown.
Legs withsegments
and costalsp.

; general

5 mm.

4 mm;

about

pruinose.
Pronotumyellow,
arcular

5.5 mm)withverylong outspreadingyellow,legs brownish
plumbeous;
with
male
Rs or more;
one-halfjRi+2 long, aboutposteriorhypopygium
at tip, laterals
acute
to nearly
lobe subacutecentralobtuse;trilobed,

segments

proximaln.

latibasis,of tergite
with basal third expanded.
dististylemale:
5.5-5.6
about 4 mm; wing
Lengthfemale:but the tergite

as in B. bilobula Alexander

ismuchdeeply bilobed ninth sternite
is quite distinct.

light

gray

sparselypruinosechiefly

blackened,
Halt?resyellow.
of legs

obscurepreinfuscated,trichialong andveinsbrown,
or longer;
Rs
one-half
about
long,
in
of i_s, i-2+3 obliquesectionthe basal

indicated;times
and one-half
at margincells R2 and Rs
to one another,parallelgenerallym-cuof M3+4.
before midlength
or the latter slightly
in extent
shortlygreater;
subequalofborderwith posterior
still darker. Male
dark brown,
Adornenhypopygium
hypopygium
about
medianwithobtuselobes
lateralprojection
tips,
longertrilobed,
tergite conspicuously
two-thirds
about

about

branches

position;threeemarginate,holotype:

two

to

two

of Rs

as

long,
tip subacuteas thetimes as wide

with

two

strongtoouterspinoidnearlyacute.
which

portionin thesetae

thirdDististylelong, basal
truncate
is slightly
dilated,notch,

North

setae

East

in holotype

Frontier

expanded,
to feeblyslightly

longer.

Agency, Assam,
Kameng,$, Bhairabkunda,
9, pinned with type.
5, 1961 (Schmid). Allotopotype,feet, March
1 $, 1 9 .
Paratopotypes,
likewisetricalcarata Alexander, which
The most similar species is Baeoura
otherin
has the tergite of the male hypopygium
trilobed, differing evidently
details of venation, nature of the leg setae, and male hypopygium.
700-1,000

Alexander:

1969]

March,

Baeourato acustyla;
generalsides of praescutum;

Alliedareasondarkened,stigmabeforethe wing

pale

antennaebrown,

veinsRs

at marginofn.

thoracic

and

65

Flies

obtusistyla,

colorationtip, cell Rs
or four major

Crane

dorsumsp.
including

blackened,

major

oval

legs blackened

very
wings
faintly
throughout;the latter terminating
and R* strongly
justdivergent,
a
with
about
four times cell R2; malehypopygium

setae at outer end of basistylegroup of three; dististyleobtuse.
apex broadlymale:antennaabout 0.9-1.0
about 3.5-4 mm; wing
3.3-3.9 mm;
Lengthfemale:about 4-4.2 mm; wing 4.2-4.3 mm.
Lengthalmostparallel-sided,

mm.

more
Rostrumand palpi
black
black.
Antennaethe pedicelso;throughout,intensely
shortersomewhatthan the longest verticils.
Headflagellarsegments
elongate,light gray.
area plumbeous
withPronotumblackened.
Mesonotal
centralgray, narrowly
praescutuma majorat anterior
blackened
blackenedoval area; posterior
opaqueend, sides withintensely
sclerites

Pleuragray pruinose.
blackened,
blackened,dark brown.
Legs blackened
throughout,Halt?resof

to appear

sparselypruinosecoxae slightly moreplumbeous.
vestiture
plumbeous;and costal
fieldsprearcular

short and inconspicuous.
very
Wings
faintlydarkened,
dark
veins
stigmabrown,
pale
conspicuous.
whitened,brown;
the latter outwardly
deflected
Ra and Ristronglystrongly
divergent,
four times cell R<?; m-cu
aboutjust before wingtip, cell Rs at margin

caudad,terminating
one-fourth
its
about

fork of M.
length beyond
Abdomenbrownishblack.

transverse,

legs

more

bordertruncate,angle withapicaltendingthree

Dististyle

dististyle.
sided,

lateralapex

hypopygium
intostrongly
curved,obtuse.
Phallosome

broadlyapophyses

Maleanglesproducedor four unusually

narrow,their

beyondwithwithtergitesmall

triangularbroadly

slenderaedeagus

Veins

posteriorat inner
Basistyleas long as the
halfpoints.

long setae, approximately
the slightly
expandedtheVenation:

base
long

almostandparallelsub?

straight,

tips obtuse.

holotype:

Kameng, North East Frontier Agency, Assam,
3, Bhairabkunda,? and para topotypes, #, $,700 feet, March 7, 1961 (Schmid). Allotopotype,on a single pin, March 8, 1961.
The most

similar species include Baeoura acustyla Alexander
the venation and general coloration much

both with

Alexander,
ing most evidently

in details of hypopygial

structure.

and B.

sternata

the same, differ?