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NEW OR LITTLE-KNOWN SPECIESOF CLADURAFROM JAPAN (DIPTERA, TIPULIDAE).
Amherst,Mass.
By C. P. ATBXANDER,
The genusCladura Osten Sackenis representedby " relatively
snrallnumber of forms in the United Statesand Canada,with agreater concentrationof speciesin Eastern Asia, particularly inin Japan,
Japan. Thereare now knownmorethan a scoreof species
the majority occurringin the mountainousareasof the island ofHonshu. All known speciesof the genusare adult in the autumn,
beingmost frequentfrom Septemberinto November. In the present report I am describingthreenoveltiesand furnishingadditional
recordsof distributionfor two of the uncommonand little-known
species.
Clo.dura(Cladura) alpicola .Nlexander
Shinano,Honshu: Nlount Norikura, near Reisentog€,altitude2,400meters,Septernber
8, 1951(H. Hasegawa); Norikura,near8, 1951(Hasegawa);
Kwaigahara,altitude2,m meters,Septenrber
9, 1951 ( Hasealtitude 1,500meters,Septerr-rber
Sirahoneonsen,gawa).
Cladura (Cladura) babai n. sp.
group,allied to daintio; generalcolBelongsto the rnegacouda
oration yellou', including the antennaeand legs, the tips of theunpatterned;abdomfemoranarrowlydarkened;wings subhyaline,
inal tergitesvellow. eachsegmentwith a -|. -shapeddark marking;male hypopygium with the tergite produced into two spatulate
blades; dististyle stout, expandedoutwardly; mesal lobe of basistyle with its inner arm denselycoveredwith microscopicsetulae;
terminatingin a strongrecurvedspine.
eachgonapophysis
Male; Length about7-7.5 mm. ; wing 8.5-9 mm.
yellow; palpi a little darker. AntennaeyelRostrum testaceouslow; flagellarverticils long and conspicuous.Head light gralt
palingto yellow on the occiput.
Thoracicdorsumfulvousyellow, the pleura sonlewhatpaler yellow. Halteres with stem yellow, knob weakly infuscated. Legs
with the coxae and trochanterspale yellow; rernainder of legs;
yellow,the tips of the femoraand tibiaevery narrowly blackened
unpatterned;
iarsi passinginto brownishblack. Wings subhyaline,veinsbro*tt, thosein the prearcularand costalfieldsmore yellowed.
Venation:Ru faintly indicated:R r*e lnd Rr*, subequal;petioleof
cell M r filor€ than twice no.

Feb',1e55 Bulletin of the Brooltlyn,EntomologicotSocietylgAbdominaltergitesyellow,eachsegmentwith a I -shapedbrown
area,includingthe medianstemand the nearly terrninalposterior
margin; ninth tergite darker; sternitesyellow with darkenedposterior borders. Male hypopygiumwith the posteriorborder of thetergite produced into two conspicuousspatulateblades. Apicallobeof basistylemore slenderand a little shorterthan the dististyle.
narrowedto the tip, the major setaegroupedat apex; proximal lobeof basistyledistinctive,the entire major blade and its inner arm
denselycoveredwith microscopicsetulae. Disistylestout,expanded
outwardly, the apex vaguelybilobed. Gonapophysisappearingas
a powerful sclerotizedstructure, the tip bent backward into apowerfulspine.
Most nearly allied to dai,ntiowhich has the tergite, outer lobe ofbasistyleand dististylesomewhatthe same; proximal lobe of basistylewith its inner arm a glabrousoval blade,without setulae,butwith aboutsix strongsetaeon outer half.
Habitat:Japan(Honshu).
Holotype; 3, Kurokawa, Echigo. October 15, 1953 (KintaroBaba). Paratype:3, Mount Chokai,Uzen,altitude600 meters,October17, 1950(Kotaro Shirahata),throughDr. Baba;3, Morioka,Rikuchiu,October6, 1934(T. Kato).
I am pleasedto namethis fly for Dr. Kintaro Baba,to whom I
am indebtedfor numerousTipulidaefrom Echigo. Although it iscloselyalliedto Cladura(Clodu,ro)dairnioAlexander,I regardthe
two fliesas beingdistinct.
Cladura (Cladura) itoi n. sp.
Belongstt-rthe trlegacauda
group,alliedto olficola; sizevery large(wing, tnale,exceeding10 mm.) ; generalcolorationyellow,including the palpi and antennae;legsyellow,the tips of the femoraandtibiae very narrowly and gradually darkened;male hypopygiumvery largeand complicatedin structure; basistyleelongate,its basal
lobe unequallybilobulate,the outer lobe very long and slender,
subequalin length to the style itself; dististylestout, a little morethanone-halfas longas the outerlobeof basistyle,bearinga slender
erect rod near base,tipped with denseblack setae; gonapophysis
appearingas a flattenedblade,the outer margin with microscopic
denticles,the apicalone largest.
Male: Length about9 mm.; wing 10.5mm.
Rostrum and palpi pale yellow. Antennaeyellow, the conspicrnoreyellowed
uousverticilsblack. Head abovefaintly infuscated,behind,with long erectsetae.

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Bulletin of the Brooklgn EntomologicalSociety

Yol. I'Thorax yellow, the dorsum slightly more fulvous. Haltereswhite, the baseof the knob weakly darkened. Legs yellow, the tipsof the femora and tibiae very narrowly and vaguely darkened;tarsi brown, passinginto black. Wings subhyaline,the prearcular,
costaland stigmal regions very faintly more yellowed; veins darkbrown, paler in the brightened fields. Venation: Scr relativelylong, Sc, lying shortly beforefork of Rr*r*rl Ru distinct, R,*u andRr*, subequal; petiole of cell M t about twice m; tn-cu a little lessthan its own length beyondthe fork of.M.
Abdomen obscureyellow, the posterior borders of the segments
narrowly darkened,the midregionof the tergiteslessevidentlyso;
hypopygium very large, yellow. Male hypopygium of the nxegacaudatype but evenmore accentuated,representingthe extreme ofornamentationas far as known in the genus. Ninth segmentlarge
and complex,the tergite and sternite fused into a continuousring ;tergal region producedinto two relatively slenderdorsally directedlobules; cephaladof these with a group of long erect pale setae;
region of the sternite with abundant long erect pale setae. Basistyle elongate,erect, its ventral margin rounded,with a brush ofvery long setae; basallobe of mesalface of style deeplybilobulate,the larger lobule foot-shaped,the appressedsmaller one fingerlike,tipped with a few long setae; apical lobe of style subequalto or alittle longer than the style itself, very slender,terminating in threeor four long setae,with other scatterederectvery long bristlesdown
the outer margin. Dististyle stouterthan the apicallobeof basistyle,a little more than one-halfas long, near its basewith a slendererect
rod that is fully one-fourthas long as the style; apex of style witha densebrush of black setae,with fewer similar darkenedsetaeon
the cephalicand mesalfaces. Gonapophysisabout equal in length
to the aedeagus,appearingas a flattened blade, the outer margin
with microscopicdenticles,the apicalspinelargest.
Habitat; Japan(Honshu).
Holotype: 3, Sigak6gen,Shinano.altitude 1600meters,Septemb e r 1 1 , 1 9 5 3( I t o ) .
This interestingcrane-flyis dedicatedto ProfessorSyusiro Ito,rvho has addedvastly to our knowledgeof the JapaneseTipulidae.
It is the largest regionalmemberof the genus so far discovered.
The nearestalh' is Cladura (Clodura) alpicolaAlexander, which
hasthe detailsof strttctureof the hypopygiumquite distinct.
Cladura (Cladura) nlegacauda'Nlexander
Shinano:Sigakog€n,altitude 1600meters,September11, 1953(Ito).

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Echigo:Kurokawa,October15, 1953(Baba).
Kotsuke: Hatomati Toge, Oze, September22, 1950 (Hasegawa); Hirami-no Tashiro,Oze, September7, 1952 (Fuhuhara).
L|zen: Mount Chokai,altitude 600 meters,October 17, 1950
( Shirahata).
Cladura (Cladura) shirahatai n. sp.
Belongs to the megacaudagroup; size small (wing, tnale, 5.5mm.) ; generalcolorationyellow, the mesonotumand ventral pleuravaguelypatternedwith darker; wings subhyaline,veinspale brown;
Sc relatively short, Sc, ending somedistancebefore level of fork of
Rr*r*t; cell M,long-petiolate, subequalto its stem; male hypopygium with the apical lobe of the basistyle shorter than the clavate
clististyle; gonapophysisfoot-shaped.
Male; Length about5mm.; wing 5.5 mm.
Rostrunr yellow; palpi pale basally,the outer segmentsdarker.
Antennae with the basalsegmentsyellow, the flagellum chiefly pale
brown, the segmentswith very long verticils. Head light brownish
gray.
Pronotum paleyellow. Nlesonotalpraescutuuryellow with threedarker stripes,the lateralpair and posteriorend of the medianstripe
paler,posteriorscleritesyellow,the scutallobesconspicuouslydarkened; scutellumand mediotergitelessevidentlyinfuscated. Pleuraand pleurotergitepale yellow, the ventral sternopleuriteand meronvaguely darkened. Halteres with stem white, knob conspicuously
infuscated. Legs with the coxae and trochanters pale yellow ; renrainderof legs yellow, the tips of the femora very vaguely moredarkened; outer tarsal segrnentsbecomingbrownish black. Wings
subhyaline,the prearcularand costalportionsa trifle more yellowed;
veins pale brown. Venation: Sc relatively short, Sc, ending somedistancebeforethe level of fork of Rr*r*0,Scnabout oppositeonethird the length of the latter vein ; cell M , long-petiolate,the cell
subequalin length to its stem.
Abdominal tergites pale brown, sternitesmore yellowed; ninth
segmentbrownish black. forming a conspicuousring; hypopygium
brownish yellow. Male hypopygiurn with the tergite relatively
large, its posterior border produced into two slender lobes thatare separatedby " U-shaped notch, with two further smaller andu'eakerlobuleslying still rnore ventrad. Apical lobe of basistylea little more than one-halfthe length of the style itself, shorter thanthe dististyle,the latter clavate,broader at apex than at base,provided u'ith sparsesetae:nresallobe of basistylestottt,prodttcedinto22Bulleti%of the Brooklgn EntomologicotSocietg

vot.L

a slenderapicallobuleand a stoutermore basalone. Phallosome
distinctive,the gonapophysis
foot-shaped,the apex or sole microscopicallycorrugated; aedeagusslender,subequalin length to theapophysis.
H abitat: Japan(Honshu).
Holotype: 3, Sakata,Uzen, october 15, 1953 (Kotaro Shirahata); receivedthroughDr. Kintaro Baba.
This smallCladurais namedfor the collector,Mr. Kotaro Shirahata. It is quite distinct from all other membersof the megocaudogroup in lhe venation,as the short Sc and long petioleof cell M r,andin the structureof the malehypopygium,especiallythe basistyle,dististyleand gonapophysis.This is the smallestmemberof thegenusso far discoveredin Japan.