soclllrY'
Reprintecl frorn Bur,r,r:rrNoF THE Bnoorr,rN DNTo\roLocICAL
YoI. XXV, No. 5, pp. 276-282' Decernber,1930'

NEW OR INSUFFICIENTLY_KNOWN
F RO M T H E N EA R T IC R E G ION
D IPT E R A ).

CRANE-FLIES(TIP U LID A E ,PART II.
Bv Cn,tnrns l'. Alnx.qNoER,Amherst, Nlass.x
The nerv speciesdescribed at the present time rn'ereincluded inmaterial sent to me for identification by Miss Helen E' Sweet,
taken at ancl near Claremont, California, and discussedin detailin her stucl1.on Artenisia and its ecoiogicalrelationships(Journalof Entomology and Zoology, Pomona College, r93o) ; in a large
and important collection of crane-fliestaken in the highest mountains of Coioracloby Mr. Charles F. Clagg; and in a small but
highly interesting series of Tipulidae sent to me by N'[r. Charles
W-. Johnson. The types of the species,with the exceptionol Oropesa joltnsonella,are preservedin the author's coliection.
Mr. Fred W. Edwards, of the British Museum of Natural History, has made a critical study of the hitherto unrecognizablespecies of Nearctic Tipula describedby Francis Walker in 1848 ancl
1856 ancl as a result the identity and synonymy of almost all ofthese specieshas been settled. This study has resulted in thechanging of name of several of our best-known speciesof thegenus. I am very deeply indebtedto Mr. Edwards for the opportunity of publishing these changesof names at this time. In thefollowing list, the earliest name is the correct one in all instances::a b d o m i ' n a l i s (S ay) (r8z3)'
albi. lat usW alk e r (r8 a 8 )-h ,e b e s L o e w (1863).
(r8
a
8
)
W
alk
e
r
bor ealisA l exander ( r9t5) '- ta rtg l ta n n och
dis it t nc laW al k e r ( l 8 S 6 )dois it t t oc t la W a l k e r (1 3 4 8 ) --a n g u s ti p e rn i s Loew (t86.3)'- ctncticornasDoane-( rgor ), .
dtipler Walker ( r8a8)
mi.nqweAlexander ( lgt5).
:b
e
l
l
a
L o e w ( 1863).
(
1848)
Walkerfu,rca'qlom,erata
Walker ( rSaB) - arcti,ca Curtis ( t83t ) .
- tesselataLoew ( 1863),
flatynrera Walker ( r856)
labradori'cuAlexander ( tgrs).
* Contribution from the Entomological Laboratory, Massachusetts Agricultural College.

Dcc',1930 Bu,lletin of th,e Brooklyn Entontological Society 277
si m ulat aWalker ( t 856)
tripler Walker ( r8a8): t r t ait t at a Say ( t 823) .
- inerm,i,sDoane ( rgor ),( ?wm , br osaLoew, t 863) .

In addition to the above, Tipr,ila resltrgens Walker (t848) isalrnost certainly 7-. latipennis Loer'v (r86a).
Tipula jacobus n. n.
In r848, \\'all<cr tlcscribc<l'l'ifula fili2c.r as neu'. In r9o9,
Johnsorr irlentifre<lJilipcs irs bcing onc of the larger ancl commoner speciesir-rliastern North America and the name being preoccupiedby 7-ipulafli?es trabricius (r8o5), he r-enarnecl
the species Tipula peilongipes. Mr. Edwards now informs me that thecrane-fly describedby Walker and renamed by Johnson is an entirely different speciesfrom the Eastern American speciestl-rathas
been passing uncler the name of perlongipes. I therefore namethis well-knor.n fly as above, the speciesbeing cledicatedto myfriend ancl co-worker, Professor James Speed l{ogers. The present specieshas been earlier discusseclar-rc1
figurerl (r\lexander,
C rane- f liesof Ner v Yor k, I , p. 952,f ig. 268; r 9r 9) .
Tipula mesotergata n. sp.
General coloration reddish brown, gray pruinose; malehypopygiurn of moderate size; ninth tergite large, the lateralangles produced caudad into short broad bispinous plates;
caudal margin of tergite between the lateral lobes nearlytransverse; eighth sternite with the lateral lobes low.
Male.-Length about 16 mm.; wing r4.5 mm.
Frontal prolongation of head long, yellor,v; nasus small ;palpi with the basal segment yellow, the remainder passing
into dark brown. Antennae r,vith the scape yellow; flagellum
broken. Head reddish brown, dusted with gray; a narrow
median brown vitta on anterior oortion of vertex.
Mesonotum reddish brown, with a gray pruinosity; medianregion of scutum behind the suture with a semicircular yellow area; postnotum yellowish gray. Pieura yellowish gray,the dorsopleural region more yellowish. Halteres ye1low,
the knobs infuscated. Legs with the coxae yeilowish gray;trochanters yellow; femora yellow, the tips narrowly andvaguely darkened; terminal tarsal segments blackened.
Wings yellowish gray, the base and costal region more yellorvish; stigma yellowish brown; antestigmal obliterative
areas narrow but extensive, extending from before the stigma,
across cell rsi AI into cell XI"".

i)cr'.,1!)i)0 lJulletin of the BrooA'Lyt't,
EntgntoftLgicalSociety 279

278 Bu,lletin of the l)rooklyn. Entont'ologicalSociety I-ot'xXV
Abdominal tergites yellorv, with indications of three
clarker longitudinal stlipes, the caudal margins of the segbccoming broader aucl more conspicments narrou'l1.1'el1orv,
1louson the sulrtermitralsegnretrts;hypopygiurl of moderater i th t he tcrgi tc l arge, thes iz e, r c r lr li s h . N l a l c h 1 ' p o p y g i u n rv
lateral angles pro<luceclcatl(la(l into short broa<l plates, theapices of u,hich alc obliqucll' truncated aucl ttuequally bispinous, the outer spine stor-rter;caudal margin of tergite
betlveen r.rearly transverse, tire median portion elevatecl into
two balely evident yellorv tubercles. Basistyle entire, therlorsal portion greatly narro'nvecl.Outer dististyle small andsien<ler. Inner clististyle u'ith the basal portion separated
from the main bodv of the style b1' a deep notch, the apex ofthis slenclerbasal portion ternrinating in two or three spinous
points. Ninth sterniteu,ith a quadrate luecliannotch. Eighthsternite lr'ith the cauclalmargin subtransvelseto broadly andweakly erlarginate, the lateral anglesproducetl into lorv lobes
that ale directed mesad ancl caudad; on either side of themedian line of the sternite a small brush of golclen-yellou'
setae,these clecussateacross the midline.

before apex lvith a small black spinous point. Lorr'el dististyle rnuch longer, glabrous.

I
I

Habitat: California.
Holotltpe: &, Clarenront, on Artentisia tri,dentata (H. L,.
S ' u, eet ) ;Collec to r' sN o . 2 8 .
T-ipulantesoterclata
is apparentll'most closelyrelated to 7'. ster'
ttataDoane, cliffering in thc strrrctureof the male hypopvgium.
Tipula sweeta n. sp.
llale.-Length about r3 lnm.; u'ing ro-rr 111111.
F em ale. -L e n g th a b o u t 1 4 mm.;w i n g rr.5 ml l r.
Allied and generally sirnilar to T-i.pulacaliforni,ca (Doane)(Pach1,vJ1ino
californica Doane, r9o8), differing very conspicuouslyin the structttre of the male hypopygium.
Size smalier. First flagellar segmententirely black. Head,thorax, abdomen and r,vingsalmost exactly as in californico.
Male hypopl'gium r'vith the tergite large, the caudal margin
rvith a narrow U-shaped mecliannotch, the lateral angies ofeach lobe further produced into an acute spine. Basistyle
entire. Lower dististyle conspicuouslyhairy, the distal third
narrorvecl into a spine. Eighth sternite produced mediallyinto tr'vo lobes that are separated by a deep V-shaped mediannotch, the mesaleclgesof the lobes conspicuouslyfringed withlong yellow setae. 7'. cali'fornico has the hypopygium conspiCuously larger, the tergite r,r'ith a very broad U-shaped
notch, the lateral lobes thus formecl blunt, on ventral marginI,i

Habitat: California.
Holotype:6/, Claremont, 1929 (..H.E. Szueet'1; Coliector's No.
Collect
or 'sNo. 32.
ot
1'f
e
s,
Par
alr
t
f
,
lllot
opot
t
,
f
,
c;
+
4&;
J,
42.
I talie gr cat lllcast t r ciu t r lt t t r it t gt lr is specicsiu honor of t he collector',I,Iiss ]Jelcrr l:. Su,cct.
Oropeza johnsonella n. sp.
recldishSize small (w-ing, .f, under ro ntm.); ure-sclt'tottttn
brown, the brown pt'aescutal stripes lelatively inclistinct;
halteres dusky ; legs pale brorvn, the tarsi a little paler; rnale
h1-popygium r,vith the inner dististyle a flattened blade, theafex- subtruncate, on outer margin near base with a smali
setiferous tubercle; gonapophyies recurved, tipped rvitha cut e soines.
trIali. Length about 8 mrr.; '"r'it.tg9.8 mni'
Frontal prolongation of head ancl palpi clark brorvn. Antennae (J) relatively'ofelongate,if bent backrvard extendrng
abdon-ren;scape honey-yellou'; firstto beyond.the baseflageliar segment short, the remaining segmentspassing intobrown. Heacl dark brown.
X{esonotum reddish b'rolvn, the praescututl n'ith tl-rreeinclistinct darker brown stripes. Pleura light brown, I'ith
vaguely indicated darker areas on anepisternum,veutral sternopleurite and ventral pleurotergite. Halteres c1usk1',thebaie of stem yellow. Legs r,viththe coxae yellolv, infuscatedat base; trochanters yellotv; a single (posterior) leg remains,
pale brown, the tarsi a trifle paler, more yellorvish brorvn.
Wings tinged r'vith blown, the stigma darker brorvn; veins
brow:n. Venation: Cell ,41' about one-half longer than its
petiole.
Abdominal segments lingecl rvith brown ancl y,ellou', theapices of the segments paler than the bases. Male hypopygi"tr with the.iateral portions of the tergite produced into
conspictlotls setiferous shoulders, the intermediate rnargttl
very gently crenulate; ventro-iateral arms of tergite strongly
curved, slender, not expancled outu'ardly, the apex acute or
subacute. Outer dististyle a little longer than the inner dististyle, cylindrical, not dilated at base.. Inner dististylea flattened blade, near base on outer margin lvith a small tubercle
set with conspictloussetae; apex of style subtruncate. Gonapophysesrecurved, setiferous,the tips set lvith several acutesplnes.

I),1.,t:t,:() llul[,, litt of the Brooklyn Entom,ologicalSociety 287

280 Bulletin of the Brooklyn, Entontological Society vot.xxv

rnented; wings cream-colored,with a restricted brown pattern; cell M, lacking; cell ts/ .4,1,closed.
Male.-Length about 7 mm.; wing 7.5 mm.
Rostrum gray; palpi bror'vnishblack. Antennae r2-segmentecl,black throughout; flagellar segmentsnearly globular, graciuallytlccreasingin size otttrvarclly,the terminal segm ent sm allcst . I lcad light gr ay.
l'ronotum gra,r', thc antcriol notttt-tt irrfuscatecl. Mesonotal praescutum light gral', rvith four conspicuousbrown
stripes,the intermediatepair ending some clistancebefore thesuture; lateral stripes narro\ver, reaching the suture; scutum
gray, the lobeswith barely indicated dark markings; posteriorscleritesof mesonotumgray, the parascutelladarker. Pleuragray. Halteres pale, the knobs infuscated. Legs with thecoxae pruinose; trochanters obscure yellow; femora obscure1,ei1ow,passing into brou''n at tips ; tibiae and tarsi darkbroln, the outer segmentsof the latter blackened. Wings
cream-colored,rvith a restricted brown pattern, including acloud at origin of /?s, the stigma and anterior cord, narrower
seamson posterior cord and outer end of cell rst M"; radial
field beyond cord vaguely suffused rvith brown; veins muchdarker than the ground-color, especially those beyond thecord. Venationi R* * , distinct ; cell ,4/, lacking; cell -rs/ .[1,closed; nx-cltjust before midlength of cell tst lltl,.
Abdomen gray, the hypopl-gium somewhat brighter. Male
hypopygium r'vith the lateral_lppendages 9f th9 tergite appearing as conspicllotiscurved flattenedblades,the tips acute.
Interbasal spines about one-haif the length, slender, strongly
curveo.

Habitat: Eastern North America.
Holotype:.f, Riverton, Nerv Jersey,August, rgrt (C. LV.Ioh,nson). Type returned to Mr. Johnson.
This interesting Oropezo is nameclin honor of Mr. C. \\r. Johnson, distinguishedauthoritl' on the ,\merican speciesof the genus.
In its snrall size anil gcneral appcarancc, Orope:a joh,nsonella
agrees ir.rostcloscll' u,ith O. roclcr.siAlcxancler, cliffering in the
structure of the malc h1'popl.giunr,cslreci:rllvtlic inner clististyle,rvlrich bears a setiferous tr-rbercleon outer margin be1'ondbase.
This type of structure is found in the Nearctic speciesonly inO. subalbi,pes
Johnson,an otherrvisevery different fly.
Cylindrotoma pallescens n. sp.
General coloration pale 1-sll611',
the usual black areas entirely replaced.by very pale r-ufous,scarcely evident against
the ground-color; antennaepale yellolv.
F em ale.-L e n g th rr m m .; u -i n g ro l rm.
Antennae pale 1'ellow, the teirninal segment darliened;^flagellar segmentsa little shorter than in iplendens. Heacl
brownish gray, more yellou'ish behind.
_.Thorax pale 1.ellon',the very pale rufous areas scarcelyinclicated, against the grouncl-color. Halteres pale yeilow.
Legs pale yellow, the femoral tips a little darkened; last tarsal segmentclalkened. \Vings yellorv, stigma barely evident;
veins pale_brown,those in the prearculai and costal regions
clearer yellow. Venation: Rs relatively short, about onefifth longer than R., arcuated at origin i nt at fork of ,4.1,* ,or beyond on II".
Abdomen yellotv.
Habitat: Colorado.
Holotype: Q, Ute Pass, Williams Fork Range, Grand Co., altitude ro,5oo feet, July t6, r9z9 G. F. Clagg).
Cylindrotonta pallescensis apparently most closely allied to C.
splendensDoane,being distinguishedby the almost uniformly paleyellow coloration, the dark areas being replaced by very palerufous. The wings are fu111'colored and the fly is apparentiy nota teneral individuai. The left wing of the type shows a weaktrace of an adventitiouscrossveinin cell R., in alignment n ith ru,
thus suggestingthe condition found in Cyttaronr,yi,a.
Tricyphona claggi n. sp.
_ Belongs to the diaplruna group; general coloration gray,the praescuturn with four brorvn stripes; antennae r2-seg-

l

i ,1.

f"q

Hab'itat: Colorado.
Holotype: d1, Bobtail Creek, Grand Co., altitude rr,ooo feet,
July zo, rgzg (C. F. Clagg).
Tricyplr,ona claggi is narr-redin honor of the collector, Mr.
Charles tr. Clagg, to lvhorn I am greatly indebted for numerous
crane-fliesfrom Colorado and the Philippines. The speciesis obviously allied to T. diaphana (Doane) , T. frigida Alexander, andT. erolom,a(Doane), differing conspicuouslyfrom all allied forms
by the lack of cell ,4'1,of the lr.ings.
Limnophila claggi n. sp.
Coloration of mesonotalpraescutum and scutal lobes polished black; antennae of male elongate; thoracic pleura282 Buttetin of the Brooklyn Entomological Society

vot' xxv

heavily pruinose; male hypopygium r,vith the outer dististyleslender, the apex bidentate.
Male.-Length about 6 mm.; wing 6.2 mm.
Rostrum and palpi black. Antennae (J) black throughout,elongate, if bent backward extending about to the base of abdomen; intermediate flagellar segments with the ventral face
protuberant; flagellar segments gradually decreasing in size
outwardly, the last segment small. Head black, pruinose.
fnterspaces of mesonotal praescutum and the scutal lobes
polished black, the lateral margins of the former and medianarea of the latter pruinose; remainder of mesonotum black,sparsely pruinose. Pleura black, heavily pruinose. Halteres
pale yellow. Legs with the coxae gray pruinose; trochanters
black; femora yellow, the tips blackened, most extensive on
the fore legs where a little more than the outer third is included, narrower on the middle and hind femora; tibiae
brownish black; tarsi black. Wings whitish subhyaline,thebase and costal region more yellowish; stigma oval, pale
brown; vague brown clouds along vein Cu, and the cord;veins brown. Venation: Sc, ending just before the fork of
Rg Sc, at its tip; Rs angulated at origin; veins R. and Rn
strongly divergent, cell .R, at margin wider than cell .Rn; cellM, about twice its petiole i nx-cu at midlength of cell rst Mr.
Abdomen black, pruinose; hypopygium black. Male hypopygium with the outer dististyle a slender blackened rod, thebase wider, the apex bidentate, the lower tooth slightly
larger. Gonapophysesappearing as large flattened plates, themargin with conspicuousappressedteeth. Aedeagus elongate.
Habitat: Colorado.
Holotype: 61, Bobtail Creek, Grand Co., altitude rr,ooo feet,
July zo, rgzg (C. F. Clagg). Paratopotype:6.
Lin+nophila claggi,is dedicated to the collector of the type material, Mr. Charles F. Clagg. The species differs from the small
polished black species resembling ntwnda Osten Sacken by theelongate antennae of the male and the structure of the hypopygium. The fly is doubtfully referable to the subgenus Prionolabis Osten Sacken.