of Crane

UndeseribedSpeciesMountains(D?ptera:Charles

Six new

Abstract:
Gonomyia
Ormosiaand 0.

(0.)

Alexander

October

Publicationof Eriopterine

speciesn.sp.,

0.

n.sp.;

17, 1967

flies are described,

crane

acucurvata(Protogonomyia)
furcivena(Ormosia)

Tipulidae),

P.

for

the Himalaya
XVI1

Massachusetts

Amherst,Received

from

Flies

(0.)

all from Assam,(Teleneura)n.sp.; 0.
(0.)

Eriopteraidioneurodes

these

being

laetipesn.sp.;

idiostylan.sp.,

n.sp.

neopulchra

Part XVin the Journal of the Newof this series of papers was published1967. As in earlier papers the
183?187,
EntomologicalSociety, 75:
materials were collected by Dr. Fernand Schmid in Assam, representing
theeastern Himalayas
I again express my deepest
and their southern extensions.
York

to Dr.

thankscollection

for the privilege

SchmidGonomyia

of

innerLength

an

style
about

acute
6-6.5

Thoraxblack,

size

heavily

dorsopleural

wing

7-7.5

black.

Headtoantennabrownishtheappeargray,

brown.

Halt?resvery
darkened,stigmalong and narrow,brown;
fork of Rs, Sc-> removed
fromendingnearlyoppositeto nearly
in lengthfrom about
one-third
R2+3+4 variable
m-cu
sive at margin;
at or close to fork of M.
Sei

Male
black.
includingAbdomen,hypopygium,outerlobe of basistyleverylarge, semicirculara slenderrod,
dististyle
stem narrow,setiferous,

withterminal

1.7-1.8mm.

pruinosesparselytoappearcentralpraescutal
stripe darkened
blackened.
black.
Legs
Wings
veins
darkerbrown.
Venation:alone

tip, Sex
one-halfRs;

longer than m-cu;
exten?
cell R.> very

withthe glabrousflangethe lobe. Outer
terminatingnear its base witha conspicuousintermediate
knob;
style
longest,one
at apex withtwo blackened
the more
slenderblades,lying
hypopygium
in outline,

acrossthe face of the second
transversely
triangularpoint;
its apex bent
into an acutelateradthe latter withspine,
Aedeagus

about

black,

vertex.

anterior

pruinosemembranemm;weaklyofn.sp.

of malespine.
mm;

Rostrum,
palpi and antennae
moreonopaque,conspicuous

anteriorly;

personal

to 7.5 mm);large (wingbody blackened,withthe flange of outer
lobe of basistyleverylarge,
hypopygium
a slenderouter dististylea conspicuous
near base,rod withknob

Generallymaleheavilypruinose;in outline;
semicircular
apex

in my

the types

acucurvata(Protogonomyia)

to megalomata;similarmale:of retainingof these flies.

hookstrongly

inner
small

style
weak

narrowed
setae

near

outwardly,
its base.

curved.

holotype:$, Kelang, Kameng, North East Frontier Agency, Assam, 6,000
3 6 $, with the type.
feet, July 2, 1961 (Schmid).
Paratopotypes,
The most similar regional species are Gonomyia(Protogonomyia)
acustylata
Alexander and G. (P.) megalomata
Alexander, which similarly have the outer
1 Contribution

fromthe Entomological

Laboratory,
66

Universityof Massachusetts.

Alexander:

1968]

June,

67

Flies

Crane

flange of the lobe of basistylelarge and flattened, all three species
inofstructure.
details
among themselves
hypopygial(Teleneura)

Erioptera
Generalmoresclerites
andtibiaelightwithyellow;yellowed;

yellowed,
modifiedbrown,

female:andpleuraappearing
scales ofprearcular

LengthRostrumaboutbrown,

dark
chieflymoredarkenedn.sp.

laetipesborders
and posteriorpraescutalmeronfemoralegs withyellowed;
clear
tarsi andvestiture,
tips of tibiae
lateralbrown,

byin sizesubequallegs elongate,
and costal
darker.
fields slightly
5-5.5 mm;
5-6 mm.
wing

black.

palpi

mediumAntennae

black

to

the

verticils

throughout,setae;

wings

tingedelongate.

Head

clear

yellow.

light

Pronotumandof mesonotum

colorationdifferingobscurearea

surrounding
dark
chiefly

PleuraHalt?resbrownishyellow;
includesfemoraelongateandborders
Mesonotum
lateral praescutal
yellow.
brown,
chiefly medium
root broadly
the wing
brownishyellow,
mediotergite
paleryellow.
more
meral
and
brown,
dorsopleuralyellowed.
regions
extensively
coxae and trochantersbrownishbase of stem yellow.
Legs withblack,
thatbut appearing
darkerfrom the abundantvestiture
yellowedin size to the normalthat are subequal
tarsi andsetae,
tips ofand costal
fields slightly
darker;brown,
tinged withWings
prearculartibiaescalestibiaelight yellow.
trichia
darker.
pale
brown,
to one-half
R>2+:;+i from two-fifths

at near
Venation:Cord
the wing
two-fifths
length;veins beyond
cord long and
Rs; R>+.i and R2 subequal;
as in the subgenus;
to one another,vein 2nd A gently
sinuous.
generallyparallel
Abdomenbrownishvalvesof ovipositor
black;
horn-yellow.

veinsholotype:9, Talung Dzong, Kameng, North East Frontier Agency, Assam,
7,800 feet, June 6, 1961 (Schmid). Para topotypes, 29 9, pinned with the type.
mostis distinguishedreadily from otherlaetipes(Teleneura)
Eriopteratarsi,
species with unpatterned wings by the light yellow
and E.
(T.)(T.) ctenophora Alexander
species being E.
was
at
to
the
the
A
writertheal?elaBrunetti.
parkey
provided by
species46:
Sei.
time of defining the subgenus Teleneura286-287;(Philippine Jour.
17: 277; 1933) indicates
States Mus.
1931). Edwards(Jour. Fed. Malay
generally
the most

similarsimilarnigribasis Edwards, placed in Teleneura on venational characters,
scales on the legs as possessed by other species and its
lacks the modified
position here may be in question.
that Erioptera

Ormosia(Ormosia)

furcivenan.sp.

obscurewiththreeof female
7 mm); mesonotal
praescutum
yellow(wing
a dark brown median
one withfemorahalf of central
line;
stripes, posterior
darkenedmediumand subterminal
with
basal
brown,
bands;
wings
conspicuously
yellow
areas
beforethe cord, outerradialfield uniformly
chieflyyellowvariegatedby whitish
Size

light

large
brown

on disk;
small darkened
spots along cord and elsewhereto the wing margin
the cell.
veinbisectingsupernumeraryantennafemale:
6.5 mm;
7 mm;
about
about
wing
Lengthbrown;

Rostrum

black.

Antennaewithbrown;
palpi
bicolored,
pale brown withsegments
large, weakly
setae.
abundantwith
Headdark brown,
yellow

scape

light

yellowed

vein

2nd

A witha strong

1.4 mm.

brown;
apices,six
proximaloutersegments

flagellai
darker.

68

York

New

Pronotumbrown.
halfofposteriorsclerites

of

coxae

brownishnotumbrown.

light

Pleuratrochanters

ring
with

and

yellow

end
dark

of

cell M,
inthe

Halt?resgray.
femoraLegs

witha nearly
withyellow
the extreme
tip yellow;

basal

yellow.

tibiae
band,
dark
tarsi
brown.
mediumbrown,
vestiture;
Wings
cord withcells Ra to 2nd _?_spots before
cord, beyond

2nd

of

base
brown

brownishyellow;
subterminal

a narrower

darkenedyellowby large whitishof the ground;the major
uniformly
and a small post-stigmal
brightening;
outer

with
three light brown
stripes,yellowdark brown medianvitta;
posteriorobscurevariegatedwings
darkera narrowdark

fVoL. LXXVISociety

obscurepraescutum
one with
centralgray;darkeneddiffusely
brownishMesonotal

the broad

Entomologicalareas

whitenedfurtherA,

portions,and vein

andinyellow
2nd A,

andcell Rj

includein bothpale marks
at base and

near

ends

cell Sc,adjacent
cells R and 1st A,of

1st M2;

vestitureareas;

veins

light

apex

the whitenedofofbrown,

ex?
the fork; all longitudinalveinscord, R2, m,
along
includingat margins.
to radialdarkened
Venation:R2 close
cepting Rs and 1st A vaguelyfork;
m transverse;
m-cu
a strong
fork of M;
beforevein 2nd A at near midlengthwithshortlybranchthat attains
the posterior
margin.
Abdomen
the

dark

acute

of

tips

holotype:withbrown,
provided
cerci narrowly

9, Hkayam

yellowsetae.

Ovipositor

Assam,

8,500

feet, Juneabundant

horn-yellow,

blackened.

Bourn, Manipur,

1960

22,

(Schmid).
Ormosia

is most similar to species such as O. (O.)
(Ormosia)
furcivena
idioneura Alexander and 0. (O.) idioneurodes n.sp., being told by the patternof the legs and wings and by the venation,
the supernumerary
especiallyin cell 2nd A.

crossveinThe

uniformlyouterdarkenedradialfield oftheis noteworthy.

wing

Ormosia
to idioneura;idioneurodes(Ormosia)of bodyn.sp.

antennae
and dark brown;
generalyellowbases and tips of femora yellowed;
legs brown,
relatively
long; halt?res
pale yellow;
a supernumerary
in
with
with
crossvein
wings
ground whitened,
light brown;
patternedonetwocell RA; male
with
the
with
acute,
very
largertip
dististyles,
hypopygium
unequalAlliedcoloration

brownishof maleas flattened
blades.
dark-colored
gonapophyses
appearingmale:
antenna
4.6 mm;about
about
4.5 mm;Lengthwing
black.
Rostrumand palpi
Antennae
relatively
long, morebrown;

flagellar
twice

exceedingabundant
Pronotalsegments
long-oval,
the segments,
muchporrect
yellow
scutumdark

the moreshorter

than

ones

proximaloutwardly.

1.4 mm.

Head

one-fourth

with

very

dark

brownishlong

the wing,
verticils
with

gray,

setae.

and pretergites
scutellum
Mesonotal
praescu?
brown,
pale yellow.
a darkerthat is marginedwithcentralwithdarkeryellow
stripenarrowlycon?
sides of sclerites
foveaedarkened;brown,
blackened,similarly
pseudosutural
oval,
area dark brown,
scutum withcentralthe broad
lobes obscurebrownishspicuous;yellow,and postnotum
dark brown
scutellum
dark brown,
the former with
very
long
laterally;tum

yellowbrownishsetae.

pteropleurite
trochanters
half
Wings
with
costalfork;

darker,withlight

Pleura
with
dark

dark
very

brown,

regiondorsopleuralsetae.
Halt?reslong yellow
femora withbrown;
the color produced

groundwhitened,
to produce
brown

base

and

by
especiallybase and costal

a variegated

Venation:Seiinterspaces.
a supernumerarycrossvein

apex
dark
regionappearance;

paler;

slightly

yellow.
withyellowed,
setae;
more

tibiaeveinsbrown,
Sc_ far

and

than

the

central

light
veins

brown.

andyellowed,

about
endingoppositeR-,in cell R>, at near
two-thirds

propleura
with

Legs
more

pale

the

tarsi
most

dorsalcoxae

andseamedmorein theyellowed
R* at radial
retracted;
lengthbeyondwhich

Alexander:

1968]

June,

vein

is strongly
upcurved;
at the angulation;

Ri

short

spur

Abdomen,including

spicuous,

subrectangular

2nd M?
nearlysquareits own

darkbrown.

blackenedhypopygium,in outline,

and stout,
narrows
toshortBasistyie
pale lobe thatemarginate.
one a smallrod,flattened

outer

cell
m-cu

Flies

69

m
origin,beforeattransverse,

being
fork

lengthof M; vein

with

a

2nd

A

sinuous.

strongly

a

Crane

Male

outwardly
outer
setae

very

the obtuse

tip;

posterior

two, very unequal,
long. Dististyles
secondstyle verylarge, a powerfulwitheach gonapophysis
Phallosome

its outerto an acute point.
third curved
turnedapex
blade,
nearlyparallel-sided,a few microscopic
withdenticles;
aedeagus

dark

marginoutward,gradually

tip

con?
tergitebordergentlywithhypopygium

parallel-sided,

lateradinto

ansubequalinlength,acute

point,
narrowed

truncate.

holotype:?, Sirhoi Kashong, Manipur,
7,500 feet, June 9, 1960
Assam,
Schmid().
The larger Ormosia
of northeastern Burma,(Ormosia) idioneura Alexander,
likewise has a supernumeraryin cell R? of the wings, differingcrossveininthe more heavily patterned wings, with m-cu close to the fork of M.
The
sex

male

of

is unknown.

idioneura

Ormosiatogeniculata;

centralstripe,posteriorfemorablack,

tips

brown;

wings

weaklyAllied{Ormosia)

mesonotal
notum

very

praescutum
dark

and

pleuranarrowlypaledarkened,variegatedyellow,n.sp.

idiostyla
light

brown

brown;
tibiae

knobs

by

and

into

Lengthcompact,
long blackspine;
gonapophysesabout
4.8-5.3
4.5-4.8
mm;
mm;
wing
4 mm;
about
4.5 mm.
Lengthwing

female:Rostrum

darkbrown,

with
Antennae
palpi black.
of theindividual
the apices

flagellum

dark

uniformly
two andwithdarkened;segments
proximalflagellarouter
one-half
times the segment,
verticils

Pronotumdarkbrown,

a narrow

dark

pits;

scutumlight

gray, indistinctly
stem yellow,
with

knob

Pleurabrownishdarkened.

Halt?restrochanters
the vestiture;slightly
paler;
tibiae andvariegatedby whitened

includingmajor
one
the posterior
andstigmalandwithpraescutum
tuberculate

dark

of scutellumsidesbrown,

light brown,behindtheregions.
conspicuous.

Abdomen,the outerantenna

scape

about

brown,

segmentsasterminating
inner style

in
in
ex?

apex.
1.5-1.6 mm.

pedicellarge, whitened;outer
six segments

pale,

the

very
long verticils,
shorter.
dark
Headlongestbrown.

about

anteriorbrown

Mesonotum
pretergites
pale yellow.
centralstripe that is slightlyinterrupted
scutellum
dark brown.
and postnotum

brown,
short
lined withbrownishblack,

posterioralmostarranged
outer
style

brown

darkened;tarsi light

weaklyblack marks,

darkened.

Legs

membranedorsopleuralcoxae dark brown,withblack,
tips very narrowly
pale yellow,
tarsi light brown.
Wings
weaklyposteriorincludingdarkened,

as in geniculata,almostcord
beforeexactlyarranged
Cu and anals,
cord witharcuatebands,R, M,
beyonda circle, bisectedveins brown,in the prearcularby vein M<;
paler
Costaland dark,
those of the stigmal
fringe variegatedyellow
regionspots
almostunusually

femoratarsi

black,

areas,

dark

weaklyslender,at near midlength,ateach bispinousatendedmale:

narrowof halt?res

tarsi

whitenedmalewithgeniculata-,long anddististyles
hypopygium
a long acutea small bispinous
blade
spine and bearing

a

with

areas,

in

these

cells

Venation:

m-cu

shortly

before

fork

of M.

from

the

black.
Malewithincludinghypopygium,
hypopygium
withend moretruncate,
expanded,microscopic
pubescence,caudad.
some of thebase of the head
Basistyie
stout, with

very

long,

yellow,

the moreproximalones

smaller,thoseof mesa!

facetransverse,

tergitewithlarger

setae

outer

apical

setae

of

very

small

style

70

New

and

delicate.

Outer

duced

into

a slender

in a

York

EntomologicalSocietyshorterand stouter
than the inner,dististylepro?
long and narrow,
black
which
is a slightlythat terminates
broaderblade
spine, beneathblackened
the lower margin
small
outer
with
point,
setae;
membranous,

marginnarrowlyof stylespines,the outer

near

at

narrowed

holotype:

intointerminating?, Rumkhang,

a

small

blade

that

forks

stouter;a long

inner

style

long

and

with

midlength
shorter
andone

twisted,gradually
short and compact,

two

into two unequal
black
before midlength
slender,with
Phallosome
spine.
gonapophyses
short blackspines.

slenderdivergent

Assam, 2,000 feet, March
20, 1960
March
1960.
24,
feet,
<5,
Paratopotype,

Khasi-Jaintia,(Schmid).
Allotopotype,with the allotype.

9,

5,500

in general appearance
to Ormosia
theinthe blackened(Brunetti),
present
fly differs
in the very different hypopygium,
especially
includingsimilar

While

the

and

styles,

LXXV1

[Vol.

(Ormosia)
geniculata
tibiae and tarsi andthe tergite,

both

disti

gonapophyses.
Ormosia(Ormosia)n.sp.

neopulchra

to pulchra;similara brown
colorationof thorax gray, praescutum
withGenerally
generalcentralantennaecon?
brownishfemorastripe;long,
relatively
flagellum
black;
yellow,withthatwhitened
spicuously
patterned
black; wings
light brown withspots
conspicuous
include a marginal
without
series; malehypopygium
pointedacutely
dististyles.
male:
about
5-5.2 mm;
5.4-5.8 mm;
antennaabout
1.6-1.7 mm.
Lengthwing
Rostrumdark brown,
Antennae
palpi black.
long, as shown
relatively
by the measure?
and pedicel
testaceousbrownishthe incisuresofments;
scapeyellow,
black,
flagellum
the morethesegments
proximalrestrictedly
yellowed;
segments
flagellar
subcylindrical,
verticilsof the intermediate
two and one-halfabouttimes the segments,
longestsegments
the

erect

whitishPronotumpraescutum

Head
gray.
pubescenceconspicuous.
brownish
gray,
paler
laterally;
pretergites
a brownwithcentralthe lateral pair
stripe,vestitureground;ofand scutum
Pleuragray, ventral
long and conspicuous.
coxae
with
trochanters
yellow.
Legs
brown;
yellowed,
femoraon
bandedwithdarkyellow,
conspicuously
brown,

paler.

tips narrowly
middlefemoradarkened;bases broadly
darkbrown,

uniformly
narrowermuchyellow,bases

centralnotdarkenedormarginal
whitenedand M2;

stigma darker
of veins R?, and Ra

than

the

ground;

veins

vein
tips
strongly
upcurved;that narrows
the cell.
beyondmidlength
Abdomenbrownishblack.
Male
hypopygium
shallowly
blackened,

emarginate.
Longestan oval palewithto subtruncate.
tip very obtuse
two slenderwithacute points

holotype:
feet, April
April

brown.
2nd

and

hind

femoratotheAof veinthe bendofcells R

Venation:sinuous,

2nd A ; remaining

and M
R2

with

near

and mostfork

a strong

of

loop

of Ri
i?_+:;++;
at andwitha narrow
dististylelobe on outer margin
Eachgonapophysis
additional

fore

narrow,ring

subterminal
yellowring
brightened,
and tarsi light brown,
outer segments
whitened
the most
evident
spots,
being

scarcelytibiaeapex;

than the restrictedyellowwith
Wings
light brownconspicuousin cells R2 through2nd M2 and in cell 1st A atareas more
extensive
but less distinct,
in basesdarker.

Mesonotumgray,yellow.
fromthedifferentiatedscarcelypraescutum
Halt?ressternopleurite

more

light

medianlobe broad,apex
tergalcurvedsubobtuse,
apex
rod,
gentlyat base;inner style
broader,slightlya broadblackenedsubtriangularplate,

stouterapex.

S, Rahung, Kameng, North East Frontier Agency, Assam,
7,000
25, 1961 (Schmid).
Paratypes,
2??,
7,300 feet,
Chug, Kameng,

17, 1961.

June,The most
0.

Alexander:

1968]

Crane

Flies

71

species are Ormosia (Ormosia) kashmiri Alexander,and 0.
(Brunetti),(0.)
subpulchra Alexander,
differinginstructures. All of the above species
chiefly
hypopygial

similar regional(0.) pulchra
among themselves
have the dististyles

terminatingin spinous points, not obtuse as in the presentfly.