On new or little-known Tipulid~e.

17

Holotype ~ : Belgian Congo, N.W. Kivu, 4000 feet,
Upper Oso River, forest with some grass, (wet season)
February 1924 (T. A. Barns).
Allotype 9: Uganda, Kampala, 28. iii. 1926 (G. L. R.
Hancock).
Paratypes : 2 ~ , Belgian Congo, N.W. Kivu, Mokoto
Lakes District, 5000-7500 feet, September 1921 (T. A.
Barns).
E X P L A N A T I O N OF T H E PLATES.
PLATE I.

Fig. 1. Opisthodontia spodopasta, Tams, sp. n., 9.
Fig. 2. Lechriolepis nephopyropa, Tams, sp. n., ~.
Fig. 3. Pseudolyra rnegis~a, Tams, sp. n., 9.
Fig. 4. Streblote dysimata, Tams, sp. n., 9.
Fig. 5. Bombycopsis hyatti, Tams, sp. n., 9.
Fig. 6. Leipoxais dolichoprygrna, Tams, sp. n., ~.
Fig. 7. ieipoxais typodes, Tams, sp. n., ~.
Fig. 8. Streblote callizona, Tams, sp. n., 9.
Fig. 9. Streblo~ coUenettei, Tams, sp. n., ~.
Fig. 10. Streblote daz, Hering, ~.
PLATE II.

Fig.
Fig.
Fig.
Fig.
Fig.
Fig.
Fig.
Fig.
Fig.

l. BomSycopsis lepta, Tams, sp. n., ~.
2. Odontocheilopteryx ungemachi, Tams, sp. n., ¢~.
3. Odontocheilopteryx eothina, Tams, sp. n., ~.
4. Chilena scotti, Tams, sp. n., ~.
5. Chilena cardinaUi, Tams, sp. n., ~.
6. Streblotefini~orum, Tams, sp. n., ~.
7. Chrysopsyche pyrodes, Tams, sp. n., c~.
8. Lechriolepis cryptognoma, Tams, sp. n., c~.
9. Trabala prasinoaphena, Tams, sp. n., ~,Fig. 10. Pseudolyra parva, Tams, sp. n., c~,

I I . ~ N e w or little-known Tipulid~ (Diptera).--XLVII.
Australasian Species. By CHARLES P. A L E X A N D E R ,Ph.D., F.E.S., Massachusetts Agricultural College,
Amherst, Massachusetts, U.S.A.

THE crane-flies herein described as new are all from
Australia, with the sole exception of a species of Trentepohlia from Papua. I am greatly indebted to Messrs.
Davidson, Ferguson, Heron, Mj6bergl Perkins, Taylor, andWilson for their continued co-operation in making known
this interesting fauna. Except where stated to thecontrary, the types of the novelties are preserved in th~author's coUection.
Ann. & Mag. N. Hist. Set. 10. Vol, vii.
2

18

Dr. C. P. Alexander on

Trentepohlia (Mongoma) brevipes, sp. n.
General coloration reddish brown, the dorsal pleurites
darkened; halteres short, dark b r o w n ; legs unusuallyshort, the tarsi considerably less than the tibim; legs
brownish black, the tarsi paling to dirty white ; wings witha brownish tinge, the stigma very small; abdominal
segments dimidiate, dark brown and obscure yellow.
Female.--Length about 7 ram. ; wing 5.8 mm. Hindleg, femur 7.7 mm. ; tibia 7.5 ram. ; tarsus 5.3 ram.
Rostrum yellowish brown, the palpi darker. Antennee
dark brown ; flagellar segments elongate, with relatively
short scattered verticles. Head brownish ochreous, darker
above.
Pronotum brownish black. Mesonotal prmscutum reddish brown, darker laterally and sublaterally on theposterior half, the latter areas crossing the suture andsuffusing the scutal lobes ; median area of scutum whitish ;scutellum pale testaceous, darker laterally; postnotal
mediotergite dark, paler medially, the total effect ofcoloration of the mesonotum being a vague pale medianstripe. Pleura yellowish testaceous, the dorsal pleurites
dark brown. Halteres short, dark brown. Legs with thecoxm and trochanters yellowish testaceous; femora
brownish black, the extreme bases obscure yellow ; tibiae
brownish black; tarsi brown at base, paling to dirty
white; legs short for a member of this genus, the tarsi
shorter than the tibim; tarsal segments decreasing inlength outwardly, the terminal three combined being
shorter than the second ; fore legs broken. Wihgs witha faint brownish suffusion, the small oval stigma darkerb r o w n ; cells C and Sc light yellow, the coloration continued as a narrow line along the anterior margin to thewing-apex, in cells /~e and R 3 being vaguely borderedbehind b y a dusky line ; other veins of wing, especially Cu,
less distinctly seamed with darker; veins darkened,
the costal veins more yellowish. Macrotrichia of costal
margin relatively dense and conspicuous. Venation : Sc 1
ending opposite the cephalic end of R e, the latter subequM
to R~+4; cell 1st M~. relatively small, the inner ends ofcells Ra, 2nd M:, and M a nearly in Mignment, the seconda trifle more distad ; apical fusion of ~Y'~I and 1st A slight.
Abdomen conspicuously dimidiate, the tergites darkbrown, the apicM half of each just before the posterior

newor llttle-krwwn Tipulld~e.

19

margin with a transverse obscure yellow b a n d ; on thesternites the pale coloration is more extensive, theposterior margins of the segments being less evidently
darkened; ovipositor reddish horn-colour, the tergal
valves strongly upcurved.
Hab. Papua.
Holotype, ~_, Suali, Vailala River (H. O. C. Littlechild).
Type in the collection of the Australian Institute ofTropical Medicine.

Trentepohlia (Trentepohlia) taylori, sp. n.
Gerneral coloration yellowish brown, variegated withdarker brown ; femora obscure yellow at base, deepening
to dark brown before tip, the apex narrowly and abruptly
pale yellow ; tibim and tarsi pale yellow, the outer" tarsal
segments brownish black; wings greyish yellow, with alongitudinal dusky washing at near mid-width.
Female.--Length about 9 mm. ; wing 6 × 1.2 mm.
Rostrum short, obscure yellow ; palpi brownish black.
Antennm black, the base of the first segment obscureyellow. Head brown.
Mesonotum yellowish brown, the prmscutum with threedark brown stripes, the median stripe ending just before
the suture ; scutum yellowish brown, the lobes darkened ;scutellum and postnotum dark brown, pruinose. Pleura
pale ochreous brown, the dorso-pleural region and ventral
sternopleurite somewhat darker. Halteres pale. Legs
with the cox~e and trochanters yellowish testaceous;femora obscure yellow, darkened outwardly, the coloration
deepening to a broad dark brown ring before apex, thelatter narrowly and abruptly pale yellow ; tibise and tarsi
pale yellow, the outer tarsal segments passing into brownish black ; legs relatively long and slender, especially thefemora and tibise ; no especial armature on legs. Wings
greyish yellow, the stigmal region scarcely darkened; avague brown wash appearing as a longitudianl line at near
mid-width of the wing, in the bases of the anal cells, incell M adjoining vein Cu, across the fork of M into thebase of ceil RE; veins pale, darker in the clouded area.
Venation : as in fijiensis, differing in the details : Sc
longer, Sc 1 extending to opposite one-fourth the lengthof R2+a+4 ; basal section of M 1+2 shorter, subequal to thesecond section of M.
2*

20

Dr. O. P. Alexander on

Abdomen relatively elongate, dark brown, the sternites
a little paler ; ovipositor very small, brownish horn colour.
Hab. North Queensland.
Holotype, ~_, Townsville, April 1927, at light(F. H. Taylor).
Type in the collection of the Australian Institute ofTropical Medicine.
I take great pleasure in naming this species in honour ofm y friend Mr. Frank H. Taylor. The species is closest toT. (T.)fijiensis (Alexander) in the general coloration of
the wings and body, differing in the coloration of the legsand details of venation. The brown longitudinal washon the wings of the present species is indicated in fijiensis
by a more diffuse wash in the same cells. The venationof fijiensis as originally figured (Ann. Ent. Soc. America,vii. pl. xxxiv, fig. 6, 1914) is erroneous in the length ofSc, which ends about opposite the fork of Rs.
Gonomyia (Lipophleps) mesoneura, sp. n.
General coloration dark brown; rostrum yellow;
antennae black; halteres with the knobs dark brown;
wings tinged with brownish yellow; Sc of moderate
length, Sc I ending just before the origin o f / / s , the latternearly straight.
Female. Length about 5 mm. ; wing 5-2-5-3 ram.
Rostrum light yellow; palpi black. Antennse black
throughout. Head grey, the front light yellow.
Pronotum and anterior lateral pretergites light sulphuryellow. Mcsonotal prsescutum and scutum dark brown,
the lateral margins of the former restrictedly yellow;
median region of scutum a trifle brightened; seutellum
brown, the caudal margin dirty yellow; postnotal
mediotergite brownish grey, the cephalo-lateral region
yellow. Pleura dark brown, with a broad silvery-whitelongitudinal stripe extending from the fore coxm to thebase of the abdomen ; a more yellowish area involves the
dorso-pleural membrane, wing-root, and dorsal portion of
the pleurotergite, ttalteres dusky, the knobs dark brown.
Legs with the fore coxm dark brown, the basal half whitened;
middle and posterior cox~e yellow, their bases infuscated ;trochanters obscure brownish yellow; remainder of legs
brown, the tarsi passing into black. Wings relatively
narrow, strongly tinged with brownish yellow, the stigmal
region slightly darker brown ; veins brown, more yellowish

new or llttle-bnown Tipulidm.

21in the basal and costal portions. Anterior branch of Rs
with macrotrichia the entire length. Venation: Sc ofmoderate length, S~ 1 ending just before the origin ofRs, S% a short distance from its tip ; Rs long and straight,
much as in skusei, but somewhat shorter; cell 1st M 2
smaller, the veins beyond it correspondingly elongated;cell 2nd A narrower.
Abdomen brownish black, the caudal margins of thesternites vaguely paler; ovipositor with the valves hght
horn-yellow.
Hab. Victoria.
Holotype, 9, Balnarring, April 28, 1929 (F. E. Wilson).
Paratopotype, ~_.
Type returned to Mr. Wilson.
Gonomyia mesoneura is most closely allied to G. skusei,
Alexander (gracilis, Skuse, preoccupied), and G. fuscohalterata, Alexander, in the relative length and courseof Rs, differing in the darkened halteres and relatively
short Sc.
T eucholabis (T eucholabis ) regince, sp. n.
General coloration bright yellow and shiny black ;prsescutum with three broad black stripes; thoracic
pleura black, with a transverse yellow line occupying thepteropleurite and meral region; a broad longitudinal
stripe of silvery pubescence on the ventral pleurites;
wings pale yellow, the costal region brighter ; sparsebrown seams along the cord, outer end of cell 1st Ms, andwing-apex ; cell 1st M S very long and narrow, exceedingvein MI+ z beyond it.
Male.--Length about 5.5 mm. ; wing 6 × 2.15 mm.
Female.--Length about 7 ram. ; wing 6-3 mm.
l~ostrum approximately as long an the remainder ofhead, dark brown, brighter at base ; palpi brownish black.
Antennm with the first scapal segment brown, the remainder of organ black ; flagellar segments oval. Head darkgrey, the anterior vertex more silvery grey.
Pronotum obscure yellow, darker in front. Anterior
lateral pretergites bright yellow. !Viesonotal prmscutum
bright yellow, with three shiny black stripes, the medianstripe ending some distance before the suture; lateral
stripes very broad, almost semioved in outline, reaching
the lateral margin of the sclerite ; median area of seutum
broadly yellow, the lobes almost covered b y shiny black
triangular marks, the posterior lateral callosities yellow ;22

Dr. C. P. Alexander on

scutellum bright yellow; postnotal mediotergite black,the lateral margins on cephalic half restrictedly yellow.
Pleura chiefly black, this including the extensive sternopleurite and anepisternum and the anterior portion of thepleurotergite ; dorso-pleural membrane, pteroplcurite, andmeral region light yellow, the latter with an appressed
silvery pubescence; dorsal sternopleurite with a broad
longitudinal stripe of silvery-grey pubescence. Halteresdusky, the knobs orange. Legs with the coxm darkbrown, their apices yellow ; trochanters obscure yellow ;femora yellow, the tips of the fore femora narrowly
blackened, of the other femora scarcely darkened ; tibiae
brownish yellow, the extreme base and broad apices
blackened; posterior tibim obscure yellow, the tips very
narrowly infumed; basal tarsal segments brownish
yellow, their apices darkened ; terminal tarsal segments
dark brown. Wings with a pale yellow tinge, the base andcostal region brighter yellow ; stigma oval, dark brown ;broad pale brown seams along the cord and outer end ofcell 1st M2; wing-apex very narrowly dark brown;veins brown, darker in the infuscated area along the cord ;basal and costal areas yellow. Venation: Sc I ending
about opposite two-thirds the length of Rs, Sc~ somedistance from its tip, placed at near one-third the lengthof / i s ; Re+8+4 a little shorter than Rl+e; cell 1st M eunusually long and narrow, gently widened outwardly,
longer than any of the veins beyond it ; m-cu about its
own length beyond the fork of M.
Abdominal segments brownish black, the bases of thesegments broadly yellow; hypopygium dark brown;basal sternites more extensively yellow.
Hab. South Queensland.
Holotype, ~, " Wilmont," Mr. Tamborine, in house,
altitude about 2000 feet, November 12, 1927 (Win. David-

8on).
Allotopoty~oe, ~_, November 30, 1927.
The only other species of Teucholabis described from
Australia is T. meridiana, Skuse (Victoria). The present
species is closely allied and very similar in its generalcoloration, differing especially in the details of colorationand venation. Skuse describes the venation of meridiana
as being exactly like that of the genotype, complexa, O.-S.,
with certain exceptions that do not involve the medial
field. In T. regince, cell 1st .M e is very long and narrow,notably more so than in complexa.

new or llttle-know~ Tipulid~.

23

Tasiocera hiemalis, sp. n.
General coloration dark brown ; antennm (~) elongate ;wings with vein 2nd A not distinct from the anal margin ;male hypopygium with the dististyle subterminal in position, long and slender, the apex strongly bent into an acutespine; phallosomic plate darkened, its caudal margin
very broadly emarginate.
Male.--Length about 3-2-3-3 ram. ; wing 4-4.2 mm.
Female.--Length about 3 mm. ; wing about 4.2 ram.
Rostrum and palpi brown. Antennm (~) elongate,considerably longer than the b o d y ; "outer flagellar
segments with glabrous necks, the outer ones very slender ;setm of flagellar segments angularly bent beyond base.
Antennae (9) short. Head dark brown.
Mesonomm dark brown, the lateral pretergites a little
paler.
Pleura a little paler than the notum. Haltereselongate, dusky, the knobs a little darker. Legs darkbrown, the femoral bases a little brightened. Wings witha dusky suffusion, the veins slightly darker; macrotrichia dark brown. Venation: 2nd anal vein notdistinct from the anal margin of wing.
Abdomen dark brown, the hypopygium paler. Male
hypopygium with the apical lobe of the basistyle relatively
long, but'stout. Dististyle subterminal, pale in colour,
long and slender, gradually narrowed to the outer end,
the apex strongly bent, prolonged into an acute apicalspine. Phallosome appearing as a broad chitinized plate,the caudal margin very broadly emarginate, each caudallateral a.ngle produced into a spine ; in some cases with asmaUer subtermlnal spine on either side of the phallosomic plate.
Hab. Victoria.
Holotype, ~, Belgrave, in dark fern gully, altitude
900 feet, June 4, 1928 (F. E. Wilson).
Allotopotype, 9, pinned with a paratype ~.
Paratopotypes, 12 ~ .
Type returned to Mr. Wilson.
Tasiocera terrce-regince, sp. n.
General coloration dark brown; wings with R~+arelatively long, strongly arcuated; cell 1st M 2 closed ;vein 2nd A short, nearly confluent with the anal margin ;male hypopygium with the dististyle subterminal in position, slender; phallosome consisting of flattened plat~a2~

Dr. C. P. Alexander onon either side of the aedeagus, each terminating in a conspicuous spine.
Male.--Length about 3.4 mm. ; wing 4 ram.
Rostrum brown; palpi black. Antennae (g) elongate,
the first flagellar segment very long, approximately twice
the second; outer flagellar segments with conspicuous
glabrous apical necks that become longer and more
conspicuous on the outer segments; scapal segments
obscure yellow, the flagellum dark brown. Head darkbrown.
Mesonotum dark brown, the humeral region of thepraeseutum testaceous-yellow. Pleura testaceous-yellow.
Halteres dusky. Legs pale, clothed with abundant
appressed black setae that conceal the ground-colour.
Wings with a greyish suffusion, the veins somewhat
darker ; macrotrichia brownish black. Venation : R2+3
relatively long, strongly arcuated to weakly angulated ;R2+3+4 a little longer than the basal section of R5 ; cell
1st M 2 closed, elongate; m-cu less than its own length
beyond the fork of M ; vein 2nd A very short, nearly
confluent with the anal margin, ending just before thelevel of the posterior arculus.
Abdomen dark brown. Male hypopygium with thedististyle subterminal in position, long and slender,
narrowed outwardly, the tip a slightly decurved spine,
before the apex with microscopic pale denticles. Phallosome consisting of a powerful flattened plate on eitherside of the aedeagus, the two together forming an arch,these plates relatively narrow, the apical-cephalic angle
produced into a conspicuous spine. In primaveris,
Alexander, which has a somewhat similar structure of thephallosome, the lateral plates are broader, widely dilated
outwardly, the apical-cephalic spine shorter and moreacute, the remainder of this dilated portion with microscopic appressed spinulae.
Hab. South Queensland.
Holotype, a broken ~, National Park, McPherson Range,
altitude 3000 feet, May 26, 1928 (F. A. Perkins).
Type in the collection of the University of Queensland.
Taslocera acanthophallus, sp. n.
Size large (wing, c~, 5 ram.); vein 2nd A present;
male hypopygium with the dististyle subterminal innew or llttle-known Tipulid~.

25

position, slender, provided with microscopic denticles;
a pale spine on mesal face of basistyle just caudad of the
dististyle ; mdeagus covered with microscopic spinulm.
Male.--Length about 4 mm. ; wing 5 ram.
General coloration dark brown. Antennae (c~) much
longer than the entire body, dark throughout. Mesonotum dark brown, the seutellum beneath and the anterior
lateral pretergites paler, tIalteres dusky. Legs pale,
clothed with dark set~e. Wings subhyaline, the veins
pale brown ; maerotrichia dark brown. Venation : R2+8+4
subequal to the basal section of R5 ; cell 1st M 2 closed ;m-cu nearly its own length beyond the fork of M ; vein
2nd A distinct, extending to slightly beyond mid-distance
between the arculus and origin of Rs ; cell 2nd A relatively
wide for a member of this genus.
Abdomen dark, the hypopygium somewhat paler.
Male hypopygium with the basistyle elongate, considerably
produced beyond the point of insertion of the dististyle,terminating in three or four powerful set0e ; mesal faceof style just caudad of the dististyle produced into a palespine. Dististyle long and slender, gradually narrowed
outwardly, the outer half on inner margin with several
tubercles set with microscopic peg-like spines, the apexof t h e style a decurved pale tooth. Phallosome appearing
as a depressed brown plate, broadest on basal half, theapex more narrowed, the caudal margin very gently
concave; phallosome without evident spinous armature.
,ZEdeagus densely set with microscopic spinulm.
Hab. New South Wales.
Holotype, c~, Brooklana, Eastern Dorrigo, altitude2000 feet, August 8, 1928 (W. Heron).
Tasiocera acanthophallus agrees only with T. tenuicornis,
Skuse, in the subterminal dististyle of the male hypopygium, with an acute pale spine on the basistyle immediately eaudad of its insertion. It differs conspicuously
in the slender dististyle, simple phallosome, and spinous
mdeagus.
Tasiocera cyatheti, sp. n.
Male.--Length about 2.6 mm. ; wing 3.8 mm.
Allied to T. primavera, Alexander (Encyel. Ent. Diptera,iv. p. 122, 1928), of north-eastern New South Wales,differing especially in the details of structure of the malehypopygium.

26

Dl.. O. P. Alexander on

General coloration of the body dark brown, the humeral
region of prmscutum, the lateral pretergites, and thepleura more testaceous. Antennae black, the segments
elongate-cylindrical. Wings dusky, the veins darker.
Venation: m-cu about its own length beyond the forkof M ; vein 2nd A becoming confluent with the anal
margin of wing shortly beyond the level of the arculus.
Male hypopygium (fig. 1) with the dististyle subterminalin postition, unusually slender, much as in angustistylus,
gradually narrowed to the decurved spinous t i p ; lower
margin of style on distal half with a series of pale denticles.
Apex of basistyle beyond the insertion of the dististyle
narrowed to the blunt tip, which is armed with four orfive powerful set~e in a row, in addition to scattered smaller
setm. Phallosome with the powerful lateral arms appearing as flattened blades, the apical portions dilated, inslide mounts superimposed to appear as a bridge or arch
like portion across the median line.
Hab. Victoria.
Holotype, c~, Belgrave, in fern gully, altitude 900 feet,J a n u a r y 13, 1929 (12. E. Wilson).
Type returned to Mr. Wilson.
Tasiocera bucephala, sp. n.
General coloration testaceous-brown ; wings with vein
2nd A relatively long, ending shortly beyond the origin
of R s ; male hypopygium with the dististyle nearlyterminal in position, irregular in form, the lower angle
produced into a very slender pale rod; phallosomic
structure with the caudal-lateral angles produced into
blackened horns that are directed mesad.
Male. Length about 3 mm. ; wing 4-4 mm.
Rostrum obscure yellow ; palpi dark brown. Antennae
elongate, exceeding the body. Head dark grey.
Mesonotum testaceous-brown, the lateral margins of thesclerites more yellowish. Pleura testaceous-yellow. Halteres dusky. Legs with the coxm and trochanters concolorous with the pleura; remainder of legs densely
clothed with long black set~e. Wings greyish subhyaline ;veins a little darker than the ground-colour; macrotrichia elongate, dark brown. Venation: R2+3+a inalignment with Rs, subequal to the basal section of R5 ;cell 1st M 2 elongate, closed; m-cu close to fork of M ;n e w or little-kaown Tipulid~e.

27vein 2 n d A relatively elongate, ending shortly b e y o n d theorigin of R s .
A b d o m e n d a r k brown, the sternites somewhat brighter ;h y p o p y g i u m pale. Male h y p o p y g i u m (fig. 2) with thebasistyle stout, terminating in two flattened-compressed
blades, one slender, with s m o o t h margins, its a p e x obtuse ;

o*¢¢

~

i!

d

h-

h

3
Male hypopygia of Australian species of Tasiocera, Skuse.
[Explanation of symbols : b, basistyle ; d, dististyle ; p, phallosome ;t, tergite.]Fig. 1. Tasiocera cyatheti, sp. n.
Fig. 2. Tasiocera bucephala, sp. n.
Fig. 3. Tasioccra dicksonice, sp. n.
Fig. 4. Tasiocera taylori, sp. n.
t h e second blade is wider, the m a r g i n microscopically
serrulate. Dististyle nearly terminal, arising between thet w o blades of the basistyle, the base stout, e x t e n d e d intoa d u s k y blade, the lower angle f u r t h e r produced into aDr. C. P. Alexander on

28very slender pale rod. Phallosomid structure appearing
as a roughly rectanglular plate, the caudal-lateral angles
produced into black incurved horns. Cephalad of thisstructure is a long framework of slender pale feebly
chitinized rods extending into the body.
Hab. Victoria.
Holotype, ~, Ben Cairn, above Millgrove, April 7, 1928

(F. E. Wilson).
Paratopotypes, 1 fragmentary c~ with t y p e ;2 5`~,
altitude 2900-3200 feet, in beech gully, February 9, 1929

(F. E. Wilson).
Type returned to Mr. Wilson.

Tasiocera dicksonic~, sp. n.
General coloration dark brown ; outer flagellar segments(5`) binodoluse ; wings dusky, the veins seamed with still
darker brown ; vein 2rid A barely indicated, ending opposite the origin of the arculus ; male hypopygium with thedististyle terminal in position, narrowed at base, more orless eapitate, spinous and roughened.
Male.--Length about 3.2 mm. ; wing 4-2 mm.
Rostrum and palpi brown. Antennm (~) nearly twice
the length of the body, brownish black, the scape brighter;
outer flagellar segments conspicuously binodulose, withbasal and medial whorls of vertieils. Head grey, moreyellowish behind.
Mesothorax dark brown, the pleura more testaeeousbrown. Halteres elongate, pale brown. Legs dark brown,
the coxm and trochanters slightly paler. Wings with adusky tinge, the veins seamed with still darker brown;veins pale. Venation: cell 1st M~ closed; vein 2nd A
barely indicated, ending opposite the origin of the areulus.
Abdomendark brown, the hypopygium
only a littlebrighter. Male hypopygium(fig. 3) with the dististyleterminal in position, narrow at base, broadly expanded andmore or less capitate distally, armed with two spines anda serrulate flange. Phallosomie structure relatively simple,
ehitinized, the caudal margin of the median plate emarginate; a slender blackened median spine, the margins
entirely smooth.
Hab. Victoria.
Holotype, 8', Belgrave, in dark fern gully, March 31, 1929

(F. E. Wilson).
Type returned to Mr. Wilson.

new or little-known Tipulid~e.

29

Tasiocera taylori, sp. n.
Size small (wing, ~, 3 mm.) ; distal segments of flagellum
strongly nodulose ; cell M 2 open by the atrophy of thebasal section of M 3 ; male hypopygium with the dististyleterminal in position, before apex on mesal margin produced
into a long pale spine.
Male.---Length about 2-2 ram. ; wing 3 ram.
Antennm much longer than the body, apparently about
twice this length ; flagellar segments strongly nodulose, theapical pedicel of each segment exceeding the strongly
enlarged basal portion. Head dark brown.
Mesonotum and pleura brown. Halteres with darkened
knobs. Wings uniformly dusky, the veins d a r k e r ;macrotrichia dark. Venation: R~. subequal to R2+3;
cell M s open by the atrophy of the basal section of M a ;vein 2nd A very reduced or lacking.
Male hypopygium (fig. 4) with the basistyle short andstout. Dististyle terminal in position, constricted beyond
base, thence gradually expanded, on mesal margin before
apex produced into a long pale spine. Phallosome adepressed plate, the caudal margin deeply notched medially, the lo~bes obtuse, their margins slightly roughened.
ZEdeagus smooth.
Hab. North Queensland.
Holotype, ~, Eungella, 45 miles west of Mackay, altitude2300 feet, March 1929 (F. H. Taylor).
Type in the collection of the Australian Institute ofTropical Medicine.
This interesting and very distinct species of Tasiocera
is named in honour of the collector, 1VIr. Frank t{. Taylor.
Cryptolabis (Cryptolabis) convoluta, sp. n.
General coloration brown, variegated with yellow;
wings creamy, with a pale brown fascia along the cord ;macrotrichia of cells a b u n d a n t ; male hypopygium withthe dististyle fleshy, at tip provided with spinous setae ;aedeagus convoluted; phallosomic structure massive, its
caudal margin with a broad V-shaped notch.
Male.--Length about 4-4.2 mm. ; wing 4.8-5 mm.
• 'emale.--Length about 5 mm. ; wing about 5-6 mm.
Rostrum obscure yellow, the palpi a little darker.
Antennae dark brown, slightly paler in some specimens.
Head obscure yellow to brownish yellow.

30

Dr. C. P. AlexaiMer on

Mesonotum dark brown, in some cases much paler,
obscure yellow ; lateral margins of the prsescutum broadly
pale yellow ; scutellum pale. Pleura dark brown, variegated with obscure yellow and paler brown, the dorsopleural region and a short ventral stripe being yellow, theextensive meron pale brown. Halteres pale yellow.
Legs with the coxoe and trochanters yellowish testaceous ;remainder of legs obscure yellow, the outer tarsal segments
dark brown ; segments of legs with conspicuous pale setm.
Wings with a creamy ground-colour, the caudal margin
more greyish ; a pale brown fascia extends along the cordfrom R 2 to vein Cu 1 ; veins obscure yellow, dark brownin the infuscated portions. Macrotrichia of cells relatively
extensive, including most cells with the exception of thebasal portions of R and the anal cells, and about thebasal half of cells M and Ca. Venation : r-m before or
close to the fork of Rs ; m-cu nearly its own length beyond
the fork of M.
Abdominal tergites brown, the sternites and hypopygium
obscure yellow ; in some specimens the whole abdomen ismore uniformly yellow. Male hypopygium most nearly
as in tasmanica, Alexander, in the unarmed dististyle andmassive unpointed phallosomic structure. Basistyles
short and stout. Dististyle fleshy, provided with long
setm, those at the apex more spinous, with one or two
similar stout setse on the lateral face. 2Edeagus elongate,
convoluted, beyond the base directed cephalad, thence
bent on itself and directed caudad, narrowed apically.
Phallosomie structure massive, the p o s t e r i o r portion
microscopically roughened, the caudal margin with abroad V-shaped notch, not evenly truncated as intasmanica.
Hab. New South Wales.
Holotype, c~, Brooklana, Eastern Dorrigo, along stream,
October 16, 1927 (W. Heron).
Allotopotype, ~_.
Paratopotypes, 2 ~ .

Amphineurus (Amphineurus) fergusoni, sp. n.
General coloration brown, the pr~escutum with threelight grey stripes ; halteres dusky ; femora yellow, withthree brown rings; wings yellowish subhyaline, withsmall brown dots produced b y dark patches of macrotriehia.

new or little-known Tipulidae.

31

M a l e . - - L e n g t h about 3-8-4 mm. ; wing 5-5-5 mm.
F e m a l e . - - L e n g t h about 5-6 ram. ; wing 6.6-7-5 mm.

Rostrum and palpi black. Antennm short, brown, thescape and base of the first flagellar segment more yellow ;fiagellar segments long-oval. Head dark grey.
Anterior lateral pretergites restrictedly whitish. Mesonotal prmscutum brown with three light grey stripes, insome cases the median stripe more buffy grey ; humeral
region china-white; scutum brown, each lobe and the
median area variegated with grey; scutellum obscureyellow; postnotum dark grey. Pleura pale brownish
grey, the dorsal sclerites darker brown ; sternites restricredly darkened. Halteres dusky, the base of stem
narrowly obscure yellow. Legs with the coxae and trochanters yellow; femora yellow, each with three brownrings, the last terminal in position ; tibiae obscure yellow,
the tips a little darkened ; tarsi brown, passing into darkbrown outwardly. Wings yellowish subhyaline, withsmall patches of dark macrotrichia scattered over themembrane to produce a dotted appearance; when thewing is denuded, only the stigma and a vague seam on thecord appear darker than the ground-colour ; veins pale
brown. Venation : all medial veins arising from end ofvein M, or nearly so; m - c u about one-half its length
before the fork of M.
Abdomen dark brown, the massive hypopygium obscurebuffy w i t h conspicuous black styli. Male hypopygiumwith the ninth tergite extensive, narrowed outwardly, thecaudal margin with a deep V-shaped notch, the lobes
formed dusky, their margins serrulate. Ventral lobe ofbasistyle a relatively slender pale fleshy lobe, provided withconspicuous setae. Base of style with a long ribbon-likeblade that terminates in a long black spine. Of the three
dististyles, one is a blackened scabrous rod that terminates
in an acute spine, its distal half with conspicuous appressed
spines, the inner margin before mid-length with a slender
black recurved lobe; inner dististyle a slender pale rod
from an enlarged base, the apex microscopically roughened;
third style a shorter flattened yellow blade, the apex
bifid, its outer spine acute and blackened, its inner a little
longer and broader, the tip obtuse. Gonapophyses
appearing as black flattened blades that terminate in threespines, the outer lateral one largest. Ovipositor powerful,horn-eoloured, the genital segment obscure yellow.

32

Dr. C. P. Alexander on

Hab. West Australia.
Holotype, (~, Perth, August 22, 1926 (E. W. Ferguson).
Allotopotype, ~2.
Paratopotypes, 2 (~_ ; paratypes, 2 ~ , Pemberton,
August 28, 1926; 1 9, Bridgetown, August 29, 1926;
1 c~, Donnybrook, August 29, 1926 (E. W. Ferguson).
Type returned to Dr. Ferguson.
This very distinct crane-fly is named in honour of thememory of the collector, the late Dr. Eustace W. Ferguson,
whose untimely death removed from the lists one of themost capable of Australian entomologists.

Amphineurus (Amphineurus) spinitergatus, sp. n.
General coloration grey, variegated with darker;
thoracic pleura with two narrow dark brown stripes;
knobs of halteres brownish yellow ; wings narrow, greyish
yellow, maculated with patches of black and yellow setse ;R2+~+4 long, more than twice the basal section of Rs;
m-cu at fork of M ; male hypopygium with the ninth
tergite bearing a conspicuous black spine on either sideof the median notch.
Male.--Length about 4.8 mm. ; wing 5.5 mm.
Rostrum and palpi black. Antennae much paler,brownish ochreous, the outer flagellar segments somewhat darker. Head dark grey, the orbits narrowly paler
grey.
Posterior pronotum and lateral pretergites pale yellow.
Mesonotal pr~escutum nearly covered by three reddishgrey stripes, the interspaces a little darker ; lateral stripes
margined externally b y a narrow dark brown line;
humeral region and lateral margins of sclerite obscureyellow; posterior selerites of mesonotum light grey,
the seutellum more yellowish, especially behind. Pleuragrey, with two narrow dark brown longitudinal stripes,
the dorsal stripe including the dorso-pleural membrane.
Halteres with the stem dusky, its base light yellow, theknobs obscure brownish yellow. Legs with the eoxm pale,the fore eoxm somewhat brighter; femora yellow, withdarker setm, especially outwardly; remainder of legs
yellow, the terminal tarsal segments passing into brown.
Wings narrow, with a greyish-yellow suffusion, the base andcostal region elearer yellow; membrane covered with
scattered patches of stout black maerotriehia and morenew or llttle.known Tipulidse.

33

slender yellow hair-like setm, to produce a maculated
appearance. Venation: Sc relatively short, ending
distinctly before R~, Sc~ shortly beyond origin of R s ;~ s long, arcuated at origin, the base pale; R~+a+4 moret h a n twice the basal section of Rs; m-cu at fork of M,
somewhat oblique in position and insensibly arcuated.
Abdomen chiefly dark brown, variegated medially with
pale; hypopygium obscure yellow. Ninth tergite withthe apex heavily blackened, the margin with a U-shaped
median notch, the lateral lobes truncate, the mesal angleof each lobe further produced into a long, straight, powerfulspine. Outermost dististyle longest, gently expanded at
tip into a paddle-like portion, the margin with appressed
spines and a few yellow setm. Phallosome broadly dilated,
the lateral margin on either side with a gently curved
black spine that is a little longer than the tergal spine.
Hab. New South Wales.
Holotype, ~, Brooklana, Eastern Dorrigo, altitude
about 2000 feet, September 20, 1928 (W. Heron).
Amphineurus spinitergatus is apparently closest to A.
maculosus (Skuse), differing especially in the coloration of
the antennee and halteres, and the details of venation.
The male of maculosus appears to be still undiscovered.
Erioptera (Erioptera) diplacantha, sp. n.
General coloration pale brown; wings with a faintgreyish tinge ; vein 2nd A gently sinuous ; male hypopygium with the dististyles terminal in position, the innerstyle a broad flattened blade, its apex split into two acutespines.
Ma/e.--Length about 2.5-3-3 mm. ; wing 3-4 mm.
Rostrum testaeeous, darker above; palpi brownish black.
Antennae dark brown t h r o u g h o u t . Head dark brown.
Mesonotum almost uniform pale brown, the pr~escutum
a little darker medially in front. Pleura more testaceous.
Halteres pale, the knobs broken. Legs with the coxm
and trochanters yellowish testaceous ; remainder of legs
pale brown, the terminal tarsal segments darker brown.
Wings with a faint greyish tinge, the costal region a littlebrighter ; veins pale brown, the macrotrichia dark brown.
Venation: R~+a+4 relatively long, exceeding m - c u ; R 2
about one-half R2+a; m-cu at fork of M ; vein 2nd A
gently sinuous, ending just before m-cu.
Ann. & Mag. N. Hist. Ser. 10. Vol. vii.
3

34

Dr. (I. P. Alexander on

Abdominal tergites brownish testaceous, the sternites
paler. Male hypopygium with both dististyles terminal
m position; outer style a httle longer than the inner,
relatively slender, broadest just before mid-length, thence
narrowed to the obtuse dusky a p e x ; inner dististyle abroad flattened blade that is divided by a deep apical split
into two acute spines, the lateral spine shorter, more
slender, gently sinuous ; inner spine broader at base, thelong ~apex acute. Gonapophyses appearing as slenderdusky horns, at base more expanded, with a laterallobule, before tip with a tooth-like flange.
Hab. South Queensland, New South Wales.
Holotype, ~, Mt. Tamborine, South Queensland, altitude
about 2000 feet, February 20, 1928 (W. H. Davidson).
Paratype, (~, Brooklana, Eastern Dorrigo, New South
Wales, altitude about 2000 feet, December 4, 1928
(W. Heron).

Paragymnastes riedeli, sp. n.
1920.

Gnophomyiasp., Riedel, A r k i v fSr Zoologi, xiii. no. 14, p. 4.

General coloration brownish black, handsomely variegated with yellow; halteres with orange knobs; legswith the femora obscure brownish yellow, the tips brownish
black; tibim and tarsi black; wings strongly d u s k y ;R2 present ; cell 1st 2112 open by the atrophy of the outerdeflection of M a.
Male.--Length about 5.5 ram. ; wing 5.7-6 mm.
Female.--Length about 6 ram. ; wing 5.5 mm.
Rostrum, palpi, and antennae black t h r o u g h o u t ;flagellar segments fusiform, with a dense erect pubescenceand longer verticils that slightly exceed the segments inlength. Head black, heavily grey pruinose.
Pronotum obscure yellow, the sides blackened, theposterior notum conspicuously light yellow, the narrow
median portion darker. Lateral pretergites yellow. Mesonotal prmscutum chiefly covered by three confluent
black stripes, the humeral region and small interspacial
areas before the suture obscure yellow, the lateral marginsof the sclerite clearer yellow ; scutum black, the posteriorcallosities and anterior median portion obscure yellow;
scutellum yellowish orange, the base narrowly blackened ;postnotal mediotergite black, with a yellowish spot oneither side. Pleura light yellow, conspicuously variegatedwith black, this including most of the propleura andanepisternum, the ventral half of sternopleurite, dorsal

new o~' littte-l~nown Tipulid,oe.

35edge of pteropleurite, and the sclerites surrounding the baseof the halteres ; dorso-pleural membrane sulphur-yellow.
Halteres short, pale brown, the large knobs orange. Legs
with the coxm blackened; trochanters dark reddish
krown ; femora obscure brownish yellow, the tips brownish
black; tibim and tarsi brownish black; femora withflattened scales on basal portion. Wings with a strong
dusky tinge, the small stigmal area darker brown; cells
C, Sc, and 1st A~1 clearer, subhyaline. Venation: Sc
elongate, Sc 1 ending at near three-fourths the length of_Rs, Sc~ some distance from its tip, placed just beyond
mid-length of that portion of Sc beyond the origin of
Rs ; Sc 1 longer than m - c u ; Rs long and straight ; _R2+a+4
a trifle longer than the basal section of ~ 5 ; RI+z fromthree to four times R 2 alone ; R~+a very reduced, approximately one-half _R2 or less, in some cases entirely lacking ;elements of cord in tranverse alignment; cell 1st M e
open by the atrophy of outer deflection of M3; cell
2nd M e deep, nearly twice its petiole; rn-cu shortly
beyond the fork of M ; vein 2nd A relatively straight,
the apex curved, cell 2nd A wide.
Abdominal tergites brownish black to black ; sternites
obscure yellowish brown, the outer sternites somewhatbrighter; hypopygium dark. Male hypopygium small:the basistyles short and stout, the dististyle single,
subterminal in position, flattened, yellow in colour.
2Edeagus powerful, broad-based.
//ab. Queensland, New South Wales.
Holotype, (~, Glen Lamington, Queensland (E. M j berg).
Allotylae, ~_, Brooklana, Eastern Dorrigo, New South
Wales, altitude 2000 feet, November 5, 1927 (W. Heron).
Paratopotype, (~; paratype, 1 c~, with the allotype,
November 6, 1927 ; 1 c~, November 15, 1927 (W. Heron).
Type in the collection of the Riksmuseet, Stockholm.
The type had earlier been placed in Gnophomyia by
l~iedel. [Ihe present species agrees with P. cyanoceps(Alexander) in the open cell 1st Me, differing most notably
in t h e retention of R~. I take great pleasure in naming
this interesting crane-fly in honour of m y old friend andco-worker on the Tipulidse, Postamtrat M. P. Riedel.
Ceratocheilus davidsoni, sp. n.
General coloration light brown, the pr~escutal stripes
ill-defined ; wings with a faint brown tinge, chiefly pro3*

36

On new o~" llttle.known Tipulid~a.

dueed b y vague seams along the veins ; maerotrichia ofveins relatively sparse, the anterior branch of Rs withfrom two to five ; abdominal segments bieolorous.
Male.--Length, excluding rostrum, about 5-5 ram. ;wing 5 ram. ; rostrum about 5.2 mm.
Rostrum nearly as long as the remainder of body,brown. Antennae with scapal segments darker than theflagellum. Head brownish grey.
Pronotum dark brown. Mesonotal pr~escutum lightbrown, the usual stripes scarcely evident, the medianregion darkest, the humeral region brightened; seutM
lobes dark brown, the posterior margin of each narrowly
yellow; scutellum more reddish brown, with a conspicuous dark spot medially at base ; postnotum pruinose.
Pleura light brown, more or less distinctly striped longitudinally with greyish brown, the latter coloration
including the dorsal pleurites and ventral sternopleurite.
Halteres pale, the knobs obscure yellow. Legs with theeoxa~ and trochanters testaceous-yellow; femora pale
brown, the outer ends somewhat darker; tibia~ and tarsi
pale brown. Wings with a faint brown tinge, more evident
as vague seams along the veins, the latter darker brown.
Macrotrichia of veins relatively sparse, the anterior
branch of Rs with only from two to five such trichia ;posterior branch with a dense series the entire length;_M4 with from five to six on the central portion. Venation :anterior branch of Rs relatively short and only moderately
sinuous, about one-fifth longer than _Rs ; cell R somewhat
narrowed at costa.
Abdomen bieolorous, the tergites dark brown, the bases
and apices of the segments broadly darkened, leaving
the central two-fifths yellow ; subterminal segments more
uniformly darkened ; hypopygium obscure yellow ; sternites similar, the basal darkening slightly paler.
Hab. South Queensland.
Holotype, ~, Mr. Tamborine, altitude about 2000 feet,
November 1928 (W. H. Davidson).
I dedicate this crane-fly to the memory of the late
Mr. Will H. Davidson, to whom I express m y indebtedness
for kindly co-operation in a study of the Australian
Tipulid~e. Ceratocheilus davidsoni is most similar to
C. tasmaniensis, Alexander, differing especially in the
diagnostic features listed above.