On new or little-known Tipulida~.

353

L I . - - N e w or little-known Tipulidm (Diptera).--XXXVI.
Australasian Species. By ChArLES P. AL~xANDm~,Ph.D.,
F.E.S., Massachusetts Agricultural
Massachusetts, U.S.A.

College, Amherst,

IN this instalment, additional descriptions of Australian andTasmanian species of the genus Molophilus are given. The
majority of these species belong to the pervagatus group, themate hypopygium of the typical form (pervagatus, Skuse) of
which was figured in Part X X X I I I . of this series of papers(Ann. & Meg. Nat. Hist. set. 9, vol. xix. p. 17, 1927). In
this group there are three dististyles, the outermost closely
adnate to the rnesal face of the basistyle, only the distal end
being free, the elongate stem reduced to a narrow chitinizedstrip that extends based to the level of the point of insertion of the two basal dististyles. The group is peculiar toAustralia, where it is represented by numerous, often closelyallied species.
As before, the majority of the species described at thistime were collected in Australia and Tasmania by Dr. Andr6
L. Tonnoir, to whom the types have been returned. Other
material was included in large and wduable collections sent
by Dr. E. W. Ferguson, collected by himself and Dr. Mackerras. This material has been returned to Dr. Ferguson.
I express my deep thanks to all of these gentlemen for theirkind co-operation in this study. I also wish to thank Dr.
Maekerras for further notes on certain species that were
described from the Skuse collection (M. gracillimus, sp. n.,
and M. mac~'errasi~ sp. n.).

Molophilus picticeps, sp. 11.
Belongs to the pervagalus group ; antennae of moderate
length ; head yellow, variegated with dark brown ; mesohotel pr~eseutum light reddish brown, the pleura darkbrown ; knobs of halteres yellow ; femora with two darkbrown annuli, the outermost terminal in position ; fore tibia(d') with a swollen blackish r i n g ; male hypopygium withthe apex of the basistyle unchitinised; outer dististyle
simple ; basal dististyles simple, elongate, the inner one paleand filiform.
Male.--Length about 8"1 ram. ; wing 4"2 mm.
Rostrum dark, the palpi dark brown.
Antennae ofmoderate lengttb if bent backwards extending about to theAnn. & Meg. N. Itist. Set. 9. Vol. xx.
24

354

Dr. C. P. Alexander onwing-root; basal segment light yellow, the second infuscared ; flagellum testaceous, the terminal segments passing
into brown; flagellar segments long-oval, clothed with
abundant erect Mlvery verticils. Head yellow, the vertex
with an extensive dark brown area in the centre, the broad
orbits remaining of the ground-colour.
Lateral pretergites sulphur-yellow.
Mesonotum light
reddish brown, the pr~escutum a little darker anteromedially ; scutellum and postnotum darker brown. Pleura
brown, the dorsal pleurites almost blackish, the ventral
pleurites more reddish brown~ the junction between thesternopleurite and anepisternum with an obscure yellow spot.
Halteres pale, the knobs clothed with pale yellow set~e.
Legs with the coxm and trochanters pale testaceous-yellow ;femora yellow, with two brown rings, one apical, the othersubequal, slightly paler brown, placed about its own lengthfrom the terminal ring; fore tibia ((~) with a subbasal
enlarged black annulus ; apex of tibiae and the tarsi brown.
Wings with a pale greyish tinge, the base narrowly yellowish;
veins slightly darker ; macrotrichim dark brown, longer andmore crowded to form black patches along the anterior andposterior cords, near mid-length of vein 2rid .4, and above
this p~int on veins M and Cu ; wing-axil a little infuscated.
Venation : vein 2nd .4 of moderate length, extending to justbeyond the cephalic end of m-cu.
Abdomen brown, the hypopygium more yellowish. Male
hypopygium with the basistyles elongate, the outer dististylelying far distad, the basistyle beyond it restricted, produced
mesad into a weak unchitinised beak, this region of the stylewith numerous erect delicate setm. Outer dististyle simple,
the ba~e broad, the apical portion with a short curved neck,the head oval. Outer basal dististyle a long, simple, gentlycurved, black rod that narrows gradually t'o the acute point,
the distal third provided with delicate subappreased setuleealong the outer face; inner basal dististyle only a trifle
shorter, very slender and pale, the distal portion almost fillform, the apex very feebly expanded. Tergal plate relatively
narrow, setiferous. 2Edeagus large and conspicuous.
Hab. Tasmania.
Holotype, c~, Burnie, October 24, 1922 (A. Tonnoir).

Molophilus pictipes, sp. n.
Belongs to the pervagalus group ; antennae of moderate
length ; head dark grey; mesonotum brown; an obscureorange spot on humeral region of pr~escutum ; knobs of

~ew or little-known Tipulid~e.

355

halteres yellow ; femora yellow with two dark brown annuli,
the outermost terminal ill position; all tibi,e ( ~ ) with anarrow brown subbasal annulus, that of the fore tibia
slightly swollen ; male hypopygium with the basal dististyles
simple, subequaI in size and length: relatively stout.
M a l e . - - L e n g t h about 3 8 mm. ; wing 4"6 ram.
Rostrum dark, the palpi dark brown. Antennae of moderate length, if bent backward barely attaining the wingroot ; basal segment yellow, the remainder of the organ darkbrown ; flagellar segments long-oval, with conspicuous erectpale verticils. Head dark grey.
Lateral pretergites narrowly sulphur-yellow. Mesonotal
prmscutum brown medially, with an obscure orange spot in
the humeral region extending cephalad from the pseudosutural fore,e, but not reaching the lateral margin; scutellum
and postnotum dark brown. Pleura dark greyish brown,
the region between the sternopleurite and anepisternum
variegated with obscure yellow. Halteres pale, the knobs
yellow. Legs with the coxm and trochauters obscure yellow,
with two dark brown annuli, one apical, the other postmedial, a trifle narrower, the yellow ring between about
equal in extent to the darkened tip; tibiae testaceous, theextreme base more yellowish, the tips narrowly infuscated ;all tibim with a narrow subbasal brown annulus that is atrifle narrower than the yellow base, that of the fore tibia( d~) a trifle swollen ; tarsi brown, the basitarsi more or lessbrightened basal]y, especially those of the middle legs.
Wings moderately broad, tinged with yellowish; stigmal
region infuscated ; brown patches of trichim on the anterior
and posterior cords and near mid-length of vein 2 n d A ; prcarcular region and the axilla weakly infuscatcd ; veins pale,
clothed with conspicuous dark macrotrichiae. Yenation:vein 2 n d A ending just beyond the nearly transverse m-ca.
Abdomen brown, the hypopygium paler. Male hypopygium of the same general structure as M . picticeps, sp. n.
bbasistyle not chitinized at apex; outer dististyle simple;
oth basal dististyles simple, slender, pointed at tips), differil,gconspicuously in the structure of the styli. Basistyle only
vaguely produced at apex, here very densely set with yellow
set~e. Outer dististyle with the oval head smaller. Outerbasal dististyle rather stout, the surface with triangularappressed dentieles on the outer two-thirds, giving the stylea feebly roughened appearance ; apex stout, but acute; innerbasal dististyle subequal in length and only a little more
slender, the surface reticulated with elongate-oval plate-likeareas, the apex more acute.
24 *

356

Dr. C. P. Alexander on

Hab. Tasmania.
Holotype, ~, St. Patrick River, November 4, 1922
(A. Tonnoir).
Molophilus graciUimus, sp. n.
Agreeing in the essentials of body-coloration with M.pulchripes, Skuse, differing conspicuously in the structure of
the male hypopygium.
The following comparative notes on the respective typesof pulchripes and gracillimus were made by Or. Maekerras :"Scutum yellowish brown ; dark brown in pulchripes, but thispossibly due to the gum. Wing with two well-defined spots,
one eKtending from r to the fork Rs, the other covering thefork of M. These spots are due to black hair. In pulchripes
the spots are not so clearly defined and tend rather to form
a continuous band, the wing being rather rubbed. The
swollen dark subbasal band on the fore tibia of pulchripes
is not represented on any leg in the present species. The
femora have a dark brown apical band and a pale yellow tocreamy subapical hand. The remainder is pale yellowish
brown, becoming darker towards the subapical pale band,
especially in the mid and hind legs, where the part proximal
to the pale band is dark brown, the femur being yellowish
brown oil its proximal two-thirds, then a fairly broad darkbrown band, followed by an equally broad pale band, andfinally with a brown apical band equal in width to the others.
The legs of pulchripes are similar, but the banding is notnearly so prominent, and the darkening proximal to the paleband is only very faintly indicated. Antenme similar in thetwo species."
Male hypopyglum with the basistyles relativoly slender,
the restricted apical portion densely setiferous ; mesal apicalangle further produced into a small blunt fleshy lobe; theentire mesal face of this apical hood-like portion is densely
setiferous. Outer dististyle a simple blackened arm, strongly
curved to the obtusely rounded apex. Outer basal dististylea long, nearly straight rod, gently curved beyond the base, alittle dilated before the apex, the extreme tip produced intoa long, straight, needle-hke point ; the dilated portion of thestyle is further roughened, more especially by a series ofappressed teeth-like spines on the outer margin before theend, some of the teeth larger and more conspicuous ; innerbasal dististyle a little shorter than the outer, nearly straight,
appearing as a very slender needle-like rod, the surface of

new or little.known Tipulid~e.

357

the distal fifth indistinctly roughened. ~Edeagus straight,
shorter than the ilmer basal dististyle.
Hub. New South Wales.
Holotype, d'~in the Skuse Collection, Macleay Museum,as paratype No. ] of M. pulchripes, Skuse; " n o exact
locality data on the pin."--Mackerras.

Molophilus euuonia, sp. n.
Belongs to the pervagatus group ; general coloration darkbrown, including the head ; basal segment of scape yellow ;tips of femora abruptly and conspicuously dark brown;
wings greyish, clouded and washed with brown ; male hypopygium with the outer dististyle black, truncated at apex;
outer basal dististyle a broad flattened black blade that
extends into a long point ; inner basal dististyle slender,
the apex dilated itlto a small head that is weakly spinulose.
Male.--Length about 4 ram. ; wing 5 mm.
Rostrum and palpi dark brown. Antennm of moderate
length, if bent backward extending to beyond the wing-root ;basal segment of scape yellow, the remainder of the antenna
dark brown ; flagellar segments lo~lg-oval, with a long erectpubescence and conspicuous vertieils. Head brown.
Pronotum brown, narrowly yellowish on either side of themedian line. Lateral pretergites bright sulphur-yellow,
broader and more conspicuous before the pseudosuturalfore,e, thence narrow and inconspicuous back to the wingroot. Mesonotum uniformly dark brown, only the humeral
region restrictedly sulphur-yellow; pseudosutural fove~eblack ; scutum coneolorous with the prmscutum ; scutellum
light brown; postnotum dark brown. Pleura uniformly
dark brown, the dorso-pleural membrane dusky. Halteres
pale at base, the stem a little dusky, the knobs conspicuously
light yellow with golden setm. Legs with the coxm reddish
brown, the fore coxm darker; trochanters obscure yellow;
only a single (posterior) leg remains; femora yellow, alittle infascated subbasaliv, the tips conspicuously andabruptly dark brown; tibiae yellow, the tips passing intobrown; basitarsi with the proximal half pale~ the remainder of the tarsi dark brown. Wings greyish, conspicuously washed and clouded with brown, tMs including
the stigma, wing-tip, cord, a conspicuous transverse wash atlevel of origin of Rs, and a brown cloud at base of cell
2**d .4 ; the pale ground-colour is thus restricted to a small
post-arcular area, with larger discal patches before andbeyond the cord ; wing-base conspicuously yellowish ; veinsdark brown, a little paler in the ground-areas, veins Cul and358

Dr. C. P. Alexander on

R a somewhat darker ; macrotrichi~e conspicuous, dark brown.
Yenation : r lying just proximad of r-m, the basal sectionof R~+a being a little shorter than R4+5 ; m-cu transverse,
about one-half the petiole of cell 21/3; vein 2ndA relatively short, ending about opposite the caudal end of m-cu.
Abdomen dark brown, including the hypopygium. Male
hypopygium with the basistyles relatively short and stout,
the ap~'x not conspicuously produced, armed with a dense
patch of yellow set~e. Outer dististyle ihort, heavily
blackened, apparently simple, but the apex dilated outwardly
and transversely truncated. Outer basal dististyle a broad
flattened blade that tapers gradually to a long straight apicalspine. Inner basal dististyle shorter and more slender, theapex only slightly dilated into an oval head that is about
one-fifth the total length of the style, the base and marginsof the head with conspicuous small erect spinules.
Hub. Tasmania.
Holotype, c~, Hartz Mts.~ altitude 3000 feet, December 9,1922 (A. Tonnoir).

Molophilus pusio, sp. n.
Belongs to the pervagatus group; basal segment ofantenna yellow ; mesonotum light reddish brown, the postnotum and pleura dark brown, the latter with a more or lessdistinct longitudinal pale stripe on the sternopleurite ; knobsof halteres light yellow ; tips of femora narrowly but notvery abruptly darkened ; male hypopygium with the outerbasal dististyle of moderate width; inner basal dististyleslender, the apex spatulate, with a few microscopic spinnles
along the outer margin.
Male.--Length about 3 ram. ; wing 4"2 mm.
Female.--Length about 3"5 mm. ; wing 4"6 mm.
Rostrum and palpi brownish black. Antennm (c~) ofmoderate length~ if bent backward extending almost to theroot of the halteres ; basal segment of scape conspicuously
light yellow; second segment light brown; remainder ofantenna dark brown; flagellar segments (c~)elongate-oval,
with long conspicuous erect white pubescence. In thefemale the antennm are shorter, scarcely attaining the wingroot. Head brown, the front and anterior part of vertex
yellowish; in some specimens the posterior orbits are paler.
Pronotum dark, the posterior notum yellow. Lateralpretergites narrowly light sulphur-yellow. 1YIesonotal prms :utum, seutum, and scutel]um light chestnut-brown, thehumeral triangle yellowish ; postaotum dark brown. Pleuranew or little-known Tipulid~e.

359

dark brown, the sternopleurite with the dorsal portion
extensively pale to produce au indistinct longitudinal pale
stripe. Halteres pale, the knobs light yellow. Legs withthe coxm yellowish testaceous ; trochantcrs yellow; femora
yellow, the tips narrowly darkened; tibim brownish yellow,
fore tibia ( ~ ) with a narrow subbasal dark ring; tarsal
segments passing into dark brown. Wings with alight grey
tinge, the cord and a broad suhbasal cloud pale brown ;wing-tip inconspicuously darkened; wing-base narrowly
yellowish; veins darker than the ground-colour~ provided
with dark macrotrichim that are more dense and conspicuousalong the cord. Venation : r lying just beyond the level ofr-m; vein 2nd A ending a short distance beyond the baseof the petiole of cell Ms.
Abdomen dark brown, the large male hypopygium paler,with conspicuous black dististyles. )/Isle hypopygium withthe basistyles relatively long and slender, the apex pale,truncated, further produced into a small setiferous lobule.
Outer dististyle bifid at apex, the largest lobe truncated attip. Outer basal dististyle a moderately broad blackened
blade, curved into a sickle-shaped structure, the tip acute ;on lower or cephalic margin before apex with a series ofsmall inconspicuous setul~e. Inner basal dististyle relatively
long and slender~ only a trifle shorter than the outer, at apex
dilated into a small dark spatula that has several microscopic
setiferous punctures ; the outer margin before the apex witha series of small appressed teeth or spines, the outermost
largest and placed before mid-length of the spatula. /~Edeagus
relatively long and slender, in mounts extending nearly tothe end of the basistyle. Ninth tergite small, narrowed tothe concave apex, the surface setifcrous.
Hab. Victoria.
Holotype, ~ , Sassafras, Dandenong Range, altitude 1000
feet, October 19, 1922 (A. Tonnoir).
.411otopotype, ~.
Paratopotypes, 4 (~ ?.

Molophilus hastatus, sp. n.
Belongs to the pervagatus group; head dark brown, theposterior orbits paler, more yellowish ; mesonotum reddish
brown to dark brown, the scutellum more yellowish ; femora
tipped with brown ; wings with a pale yellowish tinge, washedand clouded with brown ; male hypopygium with the outerdististyle trilobed; both basal dististyles slender, the outer
acute, the inner smaller, dilated into a small spatulat¢ head.

360

Dr. C. P. Alexander on

M a l e . - - L e n g t b about 3 mm. ; wing 4-4"2 mm.

Rostrum and pa]pi dark brown. Antennae short, if bent
backward not attaining the wing-root ; scapal segments a
trifle paler than tile dark brown flagellum; flagellar segments oval with a short inconspicuous erect pubescence andlong conspicuous whorls of verticils. Head dark brown, theposterior orbits paler, more yellowish.
Anterior pronotum dark, the posterior notum yellow.
Lateral pretergites narrowly sulphur-yellow. Mesonotum
reddish brown to dark brown, in the latter case the humeral
region more reddish brown ; remainder of mesonotum darkbrown, tile scutellum behind broadly yellow. Pleura darkbrown, the dorsal portion of the sternopleurite with a conspicuous yellow spot, this subobsolete in the Wilmot
specimen, the dorso-pleural membrane narrowly light yellow.
Halteres pale, the knobs light yellow. Legs with the coxte
and trocbanters brownish yellow; femora brownish yellow,
the tips conspicuously dark brown, preceded by a subequal
light yellow ring; in the Wilmot specimen the femora arebrownish yellow with the tips narrowly darkened, gradually
so oil the fore femora, more abruptly so on the posterior
femora; tibim brown, fore tibia ( ~ ) w i t h a dilated subbasal ring; tarsi dark brown. Wings with a pale yellowish
tinge, slightly washed with pale brown opposite the originof Rs, at the cord, and more broadly at the apex ; anal angle
darkened ; cord a.d veins M and Cul with dense patches ofbrown macrotrichim, elsewhere on the wing these being moresparse but long. Venation : r lying a short distance proximad of r - m ; vein 2nd A ending about opposite the caudal
end of the short transverse m-cu.
Abdomen dark brown, the hypopygium paler, the abdomenwith conspicuous erect pale yellow setm. Male hypopygiumwith the basistyles relatively long and slender, the apex
flattened and squarely truncated, fringed with delicate setm.
Outer dististyle with the apex flattened and conspicuously
bilobed, with an additional third lobe at the base. Outerbasal dististyle a long, slender, nearly straight spine thatterminates in an acute point, the lower or cephalic margin
with a few delicate setulm; inner basal dististyle about twothirds the length of the outer, slender, sinuous, the apex
expanded into a small spatula that is provided with a few
microscopic setiferous punctures but is not otherwise armed.
JEdeagus long, slender, extending caudad ahnost to the apexof the basistyles, the distal portion notably narrowed.
flab. Tasmania.
Holotype, ~ , Burnie, October 25, 1922 (A. Tonnoir).

new or little-knowa Tipulid~e.

361

Paratopotypes, c~, Jauuary 31, 1923 (A. Tonnoir) ; paratypes, ~ , Wilmot, January 8, 1923 (A. Tonnoir); c~,Eaglehawk Neck, Tasman Peninsula, November 17, 1922
(A. Tonnoir).
Molophilus setuliferus, sp. n.
Belongs to the pervagatus group; antennm short ; headdark brown; mesonotum and pleura dark brown; femora
yellow, the tips only vaguely and gradually darkened; wings
pale greyish, the cord seamed with darker ; male hypopygiumwith the outer dististyle unequally bifid at apex ; outer basal
dististyle a long slender spine with a group of delicate erectsetulm before apex; inner basal dististyle dilated at apex
into a large spatula.
Male.--Length about 3"3-3"4 ram. ; wing 4"4-4"5 mm.
Rostrum and palpi dark brown. Antennae relatively short,
if bent backward extending about to the wing-root; basal
segment of scape yellow, the remainder dark brown ; flagellar
segments long-oval, with a short inconspicuous pubescenceand very long conspicuous whorls of verticils. Head darkbrown.
Prouotum dark brown. Lateral pretergites light yellow
before the level of the pseudosutural fore,e, more infuscatedbehind. Mesonotum dark brown, ttle humeral region of tilepr~escutum with a small yellow triangle. Pleura dark brown.
Halteres pale yellow, the stem dusky, with dark setm. Legs
with the coxm b~ownish testaceous, darker basally; trochanters yellow ; femora yellow, slightly and very graduallydarkened at the tips, provided with appressed dark setae ;tibim brownish yellow, the tips narrowly darkened, tbre
tibia ( ~ ) w i t h an unusually narrow subbasal dark ring;tarsi testaceous-brown passing into darker brown at tips.
Wings with a pale greyish tinge, the cord with a broad,slightly darker seam; a vague brown suffusion across theI)asal and anal cells; extreme wing-base more yellowish.
Venation: r lying immediately proximad of r - m ; vein
2rid A relatively short, ending immediately before the caudal
end of the transverse m-cu.
Abdomen dark brown, including the hypopygium. Male
hypopygium with the basistyle ending in a flattened setiferous lobe. Outer dististyle bifid at outer end, the chief lobe
broad and obtuse, at its base with a small blunt arm. Outerbasal dististyle a slender, smooth, curved rod~ narrowed to
the long, acute, straight apex, before the tip with a linear
row of erect straight setulm ; inner ba~al dististyle slender,
at apex dilated into a conspicuous flattened spatula, the362

Dr. (3. P. Alexander on

margin of the stem before this apex and the surface of the
spatula microscopically spinulose, including a large conspicuous appressed spine on the margin of the spatula justbeyond the base. 2Edeagus relatively long and slender.
Hab. Tasmania.
Holotype, ~, Mt. Wellington, November 26, 1922(,4. Tonnoir).
Paratopotypes, 2 ~ d'~ November 2 4 - 2 8 ,Tonnoir).

1922 (A.

Molophilus diversistylus, sp. n.
Belongs to the pervagatus group; general coloration o[mesonotum varying from light to dark brown, the pleura
dark brown; knobs of halteres yellow; male hypopygiumwith the outer dististyle simple; basal dististyles very
unequal in shape and size.
Male.--Length about 3-3"3 ram. ; wing 3"7-4"7 ram.
Female.--Length about 3"3 ram. ; wing 4 mm.
Rostrum and palpi dark brown. Antennae short;in themale, if bent backward, extending about to the wing-root;
flagellar segments oval; antennae dark brown throughout.
In the female the antennae are slightly shorter. Headbrownish blaek.
Pronotum obscure brownish yellow, paler laterally. Mesonotum of type male and female light brown, the humeral
region and narrow lateral margins of the praeseutum yellowish; seutellum and postnotum passing into darker. In the
Tasmanian paratypes the pr~eseutum is a much darker
brown. Pleura dark brown dorsally, paler ventrally, with asomewhat paler longitudinal stripe between the anepisternum
and sternopleurite ; in the paratypes, pleura more uniformly
dark brown to brownish black. Halteres pale, the knobs
conspicuously light yellow. Legs with tim coxae and trochanters obscure yellow; femora pale brown, the tips alittle darker; remainder of the legs passing into darker
brown ; in the paratypes the legs are more uniformly darkbrown.
Wings greyish subhyaline, variegated with pale
brown along the cord and in the basal and anal cells, thisinfuscation including the stigmal region; veins darker
brown, especially Gul and Rs; macrotrichiae dark brown.
Venation : r lying just beyond the level of r-m ; petiole ofcell M3 relatively short, ending opposite the caudal endof the nearly transverse m-cu.
Abdomen dark brown to brownish black, including the
hypopygium. Male hypopygium with the ninth tergite anero or little-known Tipulida~.

363

rectangular pale plate, slightly narrowed posteriorl.y~ thecaudal margin truncated and fringed with long conspicuous
setae. Basistyle produced caudad into a narrow, flattened,dark lobe, the upper surface of which is subglabrous, themargins densely fringed with setae; outer face of style withsparse strong setae ; dorsal lobe very small. Outer dististyle
simple, the apex dilated into an oval head. Outer basal
dististyle a long powerful flattened blade that is gently
curved, narrowed gradually to the long acute tip; outermargin with conspicuous appressed serrations ; viewed fromthe side this style is seen to be slender, with the stem
glabrous, only the apex dilated into a spinulose head, that isfurther prolonged into the apical spine described. In a
position of rest, this style is directed caudad, the long apicalbeak strongly dorsad. Inner basal dististyle not more thanhalf the length and much more slender than the outer style,a simple smooth gently sinuous rod, the apex obtuse. In a
position of rest, this style is directed caudad. Ovipositor
with the valves relatively slender; tergal valves yellowish
horn-eolour, gently curved to the acute tips.
Hub. Victoria, Tasmania.
Holotype, c~, Sassafras, Dandenong Range~ Victor;%
altitude 1000 feet, October 21, 1922 (A. Tonnoir).
Allotopolype~ ~ ~in copula and pinned with type.
Paratypes, c~, King River, Tasmania, February 4,~ 1923
(A. 7bnnoir); c~, National Park, Tasmania, December 15,
1922 (.4. Tonnoir); d', Mr. Wollington, November 30,
1922 (.4. Tonnoir) ; d', Fern Tree, Mr. Wellington, November 12, 1922 (A. Tonnoir); c~, Eaglehawk Neck, Tasman
Peninsula, Tasmania, November 18, 1922 (A. Tonnoir).
The thoracic coloration of the Victoria types is very
different from the Tasmanian paratypes, but I cannot detect
any correlated .-structural differences, and must consider
them as being identical.

Molophilus flexilis, sp. n.
Belongs to the pervagatus group; general coloration
greyish brown, the head grey ; knobs of halteres somewhat
darkened; wings narrow, pale yellow, washed with pale
brown; male hypopygium with the two basal dististyles
both acute at tips.
Male.--Length 2"6--3 ram. ; wing 3"2-3"8 mm.
Female.--Length about 3 mm. ; wing about 3"8 ram.
Rostrum and palpi dark brown. Antennae of moderate
length, if bent backward extending about to the wing-root ;364

Dr. C. P. Alexander on

sealpal segments obscure oehreous ; flagellum brown, timsegments oval with a short sparse pubescence and conspicuous dark verticils, these longer than the segments.
Head light grey.
Pronotum ochreous. Lateral pretergites narrowly lightyellow'. Mesonotum greyish brown, the humeral region
scarcely or only restrictedly brightened; pseudosutural
fovem narrow, blackened; scutellum paler. Pleura darkbrown, sparsely pruinose. Halteres pale, the knobs faintly
darkened, with pale setm. Legs with the corm testaceous,darker hasally ; trochanters testaccous ; femora pale brown,
paler basally, the tips darkened ; tibiae pale brown, the tips
narrowly darker; fore tibia ( ~ ) narrowly darkened, but
scarcely enlarged at base; tarsi dark brown. Wings relatively narrow, pale yellowish, the base and costal region
brighter yellow; indistinct brown washes along the cord
and in the bases of cells Cu, 1st A, and 2~d A ; veins brown,
those in the flavous areas more yellowish; macrotrichi~edark brown, forming darker patches in the region of thestigma and along the cord. Venation: r approximately
equal to r - m ; vein 2nd A short, ending some distance
before m-cu, celt 2nd A correspondingly narrow, especiallyon its distal half.
Abdomen dark brown, the genital segment of the female
paler. ~V[ale hypopygium with the basistyles relatively
broad, the apex produced into a weak fleshy lobule that isdensely setiferous.
Outer dististyle a simple clava:eblackened rod. Outer basal dististyle stouter, a long,nearly straight rod, the tip acute, the distal half darkened
and provided with microscopic appressed spinules and smaller
setulm. Inner basal dististyle smaller, appearing as a broadly
flattened blade, beyond mid-length narrowed to a slenderacute point. _/Edeagus only a little longer than the outerbasal dististyle. Ovipositor with the valves long, horncoloured, almost straight, the tips acute.
Hab. Tasmania.
tlolotype, ~ , Hobart, November 8, 1922 (.4. Tonnoir).
Allotopotype, 7, Launceston, October 29, 1922 (A.
Tonnoir).
Paralypes, 1 ~ , in copula and pinned with the allotype ;3 3' ~', Burnie, October 25, 1922, and January 31, 1923
( A. Ton~wir).
Molophilus trianguliferus, sp. n.
Belongs to the pervaqatus group ; antennm dark throughout; head dark brown; mesonotum reddish brown; the,~ew or little.known Tipulid~e.

365

postnotum and pleura dark brown; femora with the distal
half darkened; wings with a pale brown tinge, variegatedwith darker seams ; vein 2nd A relatively short ; malehypopygium with the head of the outer dististyle large, triangular, glabrous ; outer basal dististyle a strongly curved
blackened rod, with a few microscopic setulae on outer face
before apex; inner basal dististyle a little shorter andstouter, almost straight, dilated at apex into a small
elongate head.
M a l e . - - L e n g t h about 2"8 ram. ; wing 3"5 mm.
Female.--Lengtb about 2"7 mm. ; wing about 3"2 mm.
Rostrum and palpi brownish black. Antennae short, if
bent backward not extending beyond the wing-root ; brownish black throughout; flagellar segments oval with longconspicuous verticils. Head dark brown.
Mesonotum reddish brown, tile median area of the praescutum broadly darker, tile lateral margins slightly brighter,including a small obscure yellow mark at the humeral angle ;scutum and scutellum dark reddish brown ; postnotum darkbrown. Pleura dark brown, somewhat paler in the allotype
and paratypes. Halteres pale yellow. Legs with the coxae
and trochauters yellowish testaceous; femora brown, tile
distal half somewhat darker than the base, but the apex notconspicuously darkened ; tibize and tarsi dark brown; fore
tibia (d') extensively dilated ,~ear base. Wings with a pale
brown tinge, the stigma, cord, a seam along vein Cul andthe base of cell 2nd A with a somewhat darker tinge ; wingbase narrowly yellowish; veins darker brown, the triehiae
almost black. Venation : rlying proximad of r-m, the hasal
section of R~+~ only about two-thirds the length of R4+~;
vein 2nd A relatively short~ ending just before the caudal
end of m-ca.
Abdomen dark brown, including the hypopygium. Male
hypopygium with the basistyles relatively slender, narrowed
to the apex, the mesal face at apex with a group of conspicuous yellow setae; outer face of style with conspicuous
larger sctae. Outer dististyle dilated at apex into a large,
glabrous, triangular head, tile neck short. Outer basal dististyle a slender chitinized rod, strongly curved to the acuteapex ; a few very short setulae on outer face shortly before
the tip. Inner basal dististyle nearly as long as the outer,stout, at apex dilated into a small elongate head, which issparsely microscopically spinulose and provided with a few
set~ at the apex. 2Edeagus sinuous. Ovipositor with long
slender valves.
Hab. New South Wales.

366

Dr. C. P. Alexander on

Holotype, 6~, Wentworth Falls, Blue Mts., altitude 2844
feet, November 8, 1921 (A. Tonnoir).
Allotype, ~ , Narara, November 3, 1921 (A. Tonnoir).
Paratypes, 2 ~ c~, with the allotype, November 3-6, 1921 ;1 c~, French's Forest, near Sydney, July 15, 1923 (L M.
Mackerras) ; 1 ~ , Sydney, October 14, 1923 (E. //g. Ferguson); No. 71.
Molophilus incomptus, sp. n.
Belongs to the pervagatus group; basal segment ofantenna obscure ochreous ; anterior mesonotum light brown,
the postnotum and pleura darker; halteros yellow; legs
brown, unvariegated ; wings grey with a dense grouping ofmacrotrichim along the cord; male hypopygium with theouter dististyle bilobed at apex ; outer basal dististyle nearlystraight, the base expanded.
Male.--Length about 3"5 ram. ; wing 4"3-4"5 ram.
Rostrum and palpi dark brown. Antennm relatively
short, if bent backward extending about to the wing-root;
basal segment of the scape obscure ochreous ; remainder ofantenna dark brown; flagellar segments oval with a dense
erect white pubescence and unilaterally arranged verticils.
Head dark brown, the anterior part of the vertex and tile
orbits vaguely more ochreous.
Pronotum ochraceous testaceous, with very long setm.
Lateral pretergites narrowly sulphur-yellow, brighter andmore conspicuous anteriorly. Mesonotum dull light brown,with conspicuous erect black set~e ; t)r~escutum with thehumeral region obscure yellow; scutellum brownish testaceous ; postnotum darker brown. Pleura darker brown.
Halteres pale, the knobs light yello% with silken sct~e.
Legs with the eoxm and trochanters yellowish testaceous;femora light brown, unvariegated ; tibiae light brown, fore
tibia (c~) with a dilated, slightly darkened, subbasal ring;tarsi passing into brown at tips. Wings moderately broad,tinged with grey; veins slightly darker; the macrotrichim
dark brown, conspicuous, forming a darker cloud along
the cord; veins Cul and Anals with similar dark patches.
Venation : r lying beyond the level of r-m, the basal deflection of R~+a being longer than R4+~ ; vein 2nd A of moderate length, ending just beyond the origin of the petiole ofcell Ms.
Abdomen dark brown, conspicuously clothed with erectpale setm; hypopygium a little brighter brown. Male
hypopygium with the basistyles relatively slender, the apex

new or little-known Tipulid~e.

367with the oblique mesal face with dense yellow setae. Outerdististy]e with the dilated apical portion trifid, the apex
being bilobed, with a basal triangular tooth separated fromthe apex by a deep U-shaped notch. The two basal disti~tyles appear to he reversed in position to what is normal
in the pervagatus group, the outer or more lateral one being
a little shorter and stouter, narrowest near mid-length, therelatively small apex with microseopie appressed spinulm.
Inner basal dististyle a slender, nearly straight rod from adilated yellow base, the stem narrow, thence slightly dilated,
the long black apex acute, tEdeagusvery long and slender,
in slide-mounts extending eaudad to opposite the tips of thebasistyles.
Hab. New South Wales.
Holotype, ~ , Wentworth Falls, Blue Mts., altitude 2844
feet, November 18, 1921 (A. Tonnoir).
_Paralopotype, ~ .

Molophilus micropteryx, sp. n.
General coloration dark grey ; halteres very small ; wings
subatrophied, less than 1 ram. long.
)'emale.--Length about 2"8 ram. ; wing 0"8 ram.
Rostrum and palpi black. Antennae of moderate length,
black throughout, provided with conspicuous vertieils.
Head grey..
Thorax entirely dark grey. Halteres pale brown, verysmall, tile heads feebly dilated. Legs relatively short andstout ; cox~e brownish black ; troehanters brown ; remainderof legs dark brown. Wings (of ? , at least) very reduced,
if bent backward not extending beyond tile second abdominal
segment, with a pale brown tinge, the base more yellowish ;veins darker brown, stout. Venation distorted, but evident
cell R, barely sessile; vein 1st A ending opposite m-cu ;vein 2rid A short.
Abdomen brownish black ; basal shield of ovipositor dark~
the valves light yellow, the tergal valves more slender, gently
upcurved, the tips acute.
Hab. New South Wales.
Itolotype, ~, Wentworth Falls, Blue :~¢fts., altitude 2844
feet, November 18, 1921 (.4. Tonnoir).
Molophilus mieropteryx does not agree with the known
males of any of the speci¢s described by Skusc orthe writer.
In its general appearance it agrees most closely with M.
furvus, Alexander, which is full-winged in both sexes.

368

Dr. C. P. Alexander on

Molophilus subapterogyne, ~p. n.
Subapterous, at least in the female sex; general coloration testaeeous-brown, somewhat shiny ; wings subatrophied,less than one-half' the length of the body, dimidiate, thedistal portion darker than the base; no marginal triehim
beyond the wing-apex.
Female.~Length about 3 ram. ; wing 1"2 mm.
Rostrum and palpi brownish black.
Antenum darkbrown. Head dark brown.
Mesonotum dark testaeeous-brown, somewhat shiny, thepleura a little paler. Helteres small, pale, the knobs verysmall. Legs relatively short, brownish yellow, the terminal
tarsal segments passing into dark brown. Wings subatrophied (in the ? , at least), dimidiate, the basal threefifths grey, the distal two-fifths (beyond the cord) darkbrown; costal setm relatively few aud scattered ; a row ofsimilar setze along vein R~ for its entire length ; costal
fringe ending at the wing-apex~ the margin beyond vein R~
entirely naked ; veins very indistinct and ill-defined.
Abdomen dark brown, the genital segment ochrcous, thevalves of the ovipositor long but relatively stout, the tergal
valves more slender, gently upcurved to the acute tips.
Hab. Tasmania.
Itolotype, ?, Cradle Valley, altitude 3500 feet, January23~ 1923 (.4. ~Ibnnoir).

Paratopolype, ?.
I cannot reconcile this species with any males known from
Tasmania of which the female sex is not known. The fly
resembles M. micropteryx, sp. n., in its subapterous condition,differing in the body- and wing-coloration and ir~ the slightly
longer wings.

Molophilus indivisus~ sp. n.
Belongs to the plagiatus group, allied to M. flavonotatus,
Skuse ; basal dististyle of male hypopygium a long cylindrical rod, not dilated near mid-length~ but here armed witha conspicuous lateral spine; apex of style simple, curved,
heavily blackened for a short distance back from the tip.
Male.--Length about 3"5 mm; wing 4"5 ram.
Generally similar to M.flavonotatus, Skuse, differing conspicuously in the structure of the male hypopygium.
General coloration of thorax grey, tim lateral pretergites
narrowly sulphur-yellow, the praeseutum with three confluent
brown stripes; pseudosutural revere conspicuous, dark

new or little-known Tipulidse.

369

brown ; scutal lobes brown, the median region grey ; scutellure dark brown, the caudal margin inconspicuously paler ;postnotum dark brown, the anterior lateral angles of themediotergite paler. Pleura grey. Haheres pale. Legs
with the eox~e and trochanters brownish yellow ; femora
dark brown, their bases more yellowish; remainder of thelegs dark brown; fore tibia (d') with an enlarged subbasal
brownish-black ring. Wings subhyaline, the brown spot in
the base of cell Sc well-defined ; veins darker brown.
Male hypopygium brownish yellow, the basistyles more
suffused with brown. Dorsal lobe of basistyle dusky incolour, unarmed with a spine ; apical spine of ventral lobeof basistyle long and slender. Outer dististyle slender, thearms long; outer arm expanded into a spatula, the innerarm slender, blackened. Basal dististyle a slender, cylindrical, sinuous rod, near mid-length bearing a single strong
black spine, the apical portion beyond this curved to thesimple acute black apex.
~deagus short, ending about
opposite the spine of the basal dististyle.
Hub. New South Wales.
Holotype, ¢~, Sydney, September 14, 1923 (E. W. Ferguson) ; No. 69.
Paratype, ¢~, in the Skuse Collection as paratype No. 1of the type-series of M. canus, Skuse ; no exact localitydata.--Mackerras.

Molophilus mackerrasi, sp. n.
Belongs to the plagiatus group ; legs banded quite as in
M. annulipes~ Skuse ; wings narrow, yellowish~ with no dark
spot in the base of cell Sc ; male hypopygium with the basal
dististyle a short powerful rod, the dilated head terminatingin two powerful black spines.
Although confused by Skuse with his species annulipes,
tim present form does not even belong to the same subgroup, but is more correctly referable to the typical divisionof the plagiatus group.
The following comparative notes on the two types ofmackerrasi and M. annulip'es, Skuse, are kindly supphed by
Dr. Mackerras : " M . mackerrasi.--Legs not differing appreciably from those of annulipes. Wings narrow and yellowish ;no dark spot in cell Sc. Antennm uniformly greyish yellow.
M. annulipes.--Wings rather broad, clear, the spot in cell
Sc well-marked. Antennm very pale, with narrow blackbands at the constrictions between the segments, a very
striking feature."
Ann. & Mug. ~ . Hist. Ser. 9. Vol. xx.
25

370

Dr. C. P. Alexander on

Male hypopygium with the apical beak of the basistyle
blackened, relatively large, the apex subacute, the base of
the beak surrounded by numerous set~e. Outer dististyle
pale basally, the stem sinuous, the two arms unequal, thelateral arm much larger than the mesal arm, directed mesad,
both arms of the style blackened. Basal dististyle a shortand very powerful rod, the stem short, soon dilated into ahead that terminates in two powerful black spines, the mesal
spine lying along the axis of the style, gently curved, thelateral spine nearly straight, directed mesad and lying transversely across the end of the style. 2Edeagus relatively longand slender, considerably longer than the basal dististyle.
Hab. New South Wales.
Holotype, c~, in the Skuse Collection, as paratype No. 2of the type-series of M. annulipes, Skuse ; Hogan's Brush,
August.--Mackerras.
This very distinct fly is named in honour of Dr. :[anMackerras, to whom I am vastly indebted for co-operationin making known the identities of the numerous species of
Molophilus described in 1889 by Skuse.

Molophilus in~equidens, sp. n.
Belongs to the plagiatus group; general coloration richbrown, the lateral pretergites conspicuously light sulphuryellow; legs dark brown; wings greyish subhyaline; vein
2nd A short ; male hypopygium with the outer arm of tileouter dististyle sinuous, acutely pointed; basal dististyle ashort, straight, flatfened blade, terminating in two unequal
teeth.
Male.--Length, when entire, probably about 3 ram. ; wing
about 3"8 ram.
Head of the type destroyed by Psocids.
Lateral pretergites conspicuously light sulphur-yellow, thecolour extending eaudad to the wing-root. Mc~onotal prmscutum and scutum rich brown, the humeral region of theformer paler; scutellum and poatnotum vaguely darker
brown, slightly pruinose. Pleura dark. Halteres broken.
Legs with the cox~e and trocha/lters brownish testaceous ;remainder of legs dark brownish black, the femoral bases
only vaguely brightened; fore tibim ( ~ ' ) w i t h a subbasal
black ring. Wings greyish subhyaline, the veins darker.
Yenation : vein 2nd A short, ending before the level ofm-cu.

Abdomen destroyed by Psocids, only the hypopygium
remaining3 this brownish yellow in colour. Apical spine of

new o~" llttle-known Tipulid~.

371

basistyle short and moderately stout, heavily blackened.
Outer dististyle slender, the two arms very unequal~ theouter more than twice the length of the iImer, sinuous to
the acute point; inner arm stout, straight, both arms
blackened. Basal dististyle a short straight flattened blade,the apex bifid, the two teeth straight but very unequal, theouter much larger ; besides these two primary teeth, there
are a few microscopic spines on the blackened distal third ;the basal style is distinctly shorter than the outer dististyle.
j~deagus relatively short, the straight apical portion about
as long as the basal dististyle.
Hab. New South Wales.
Holotype, 6 , Sydney, September 2, 1923 (E. 14r. Fer#uson) ; No. 73.
Molophilus arcanus, sp. n.
Belongs to the plagiatus group ; mesonotum light reddish
brown ; the pleura yellowish testaceous with a broad brown
dorsal longitudinal stripe; knobs of halteres infuscated;male hypopygium with the apical beak of the basistyle
microscopic; outer dististyle with the stem stout, the twoarms blackened ; basal dististyle a small slender rod that isabout one-third the length of the mdeagns.
Male.--Length about 4 ram. ; wing 5"5 mm.
Rostrum and palpi dark brown. Antennae brown throughout, of moderate length, if bent backward extending to ashort distance beyond the wing-root; flagellar segments
long-oval, with conspicuous verticils. Head dark brown.
Mesonotal praescutum light reddish brown, the posteriorseleri~es of the notum somewhat darker. Pleura yellowish
testaceous, with a conspicuous broad dorso-longitudinal
brown stripe, extending from the propleura, passing caudad
above the root of the halteres to the abdomen. Halteres
narrowly pale basally, the stem and knobs dark brown.
Legs with the eoxae and troehanters yellowish testaceous ;femora pale brownish testaeeous, with sparse appressed darker
setae; tibim and tarsi pale, the terminal tarsal segments
darker ; middle tarsi a little longer than the tibiae ; hind tarsi
shorter than the tibi,e. The fore legs of the unique type arebroken, and it cannot be stated whether there is a sexual ringon the tibiae. Wings with a faint yellowish tinge, the veins
pale brown ; macrotrichiae relatively short and sparse, brown.
Venation: basal section of R2+a a trifle longer than R4+~;
vein 2nd A relatively short, ending about opposite the caudal
end of m-cu.
Abdomen dark brown, the hypopygium a very little
25 ~

372

Mr. A. M. Massee onbrighter. Male hypopygium with the ventral lobes of thebasistyles produced, as in the group, terminating in a tiny
feebly chitinized beak, the integument surrounding this beakwith abundant yellow setm. Outer dististyle very large
and powerful, the stem short and broad, the arms large,
blackened, both truncated. Basal dististyle very small,shorter than the stem of the outer dististyle, a feebly chitinized rod that gradually narrows to the slender tip~ thesurface and apex feebly tuberculate. 2Edeagus relatively
long, moderately stout, approximately three times as long as
the basal dististyle.
Hab. Tasmania.
Holotype, c~, Mt. Wellington, November 25, 1922 (.4.

Tonnoir).

LII.--Descriptions of Three new Species of Gall-Mites(Eriophyidse) from Sudan. By A. M. MASSEE, F.E.S.,
East Mailing :Research ~tadon.
THESE notes are the result~ of the examination of a smallcollection of galls, preserved in spirit, which were forwarded
to East Malling Research Station by Dr. G. A. K. Marshall,
Director of the Imperial :Bureau of Entomology.
Although it is indeed very difficult to study these milmte
animals when they have been preserved in spirit for any
length of time, it has been possible to describe accurately andname three species, all belonging to the well-known genus

Zr~ophy~.

Jn one of the vials containing Eriop]tges balanltes, sp. n.,
an inquiline belonging to the genus _Phglloeoptes was foundto be present in small numbers. Utffortunately, it has notbeen possible to describe this species, owing to the lack ofmaterial, but it is hoped to procure further material in orderthat this species may be described at a later date.
The writer is indebted to Dr. G. A. K. :Marshall for much,~aluablo assistance.

.Eriophyes balanites, sp. 11.
:Body moderately large, drawn-out, cylbMrieal. :Nymphal
stages ~orm-shaped. Thoracic shield semicircular, not projecting to a marked extent over rostrum~ 36 ~ in breadth.
Shield ol namented with longitudinal fin'rows, the ~hree central