សៀវភៅមគ្គុទេ្ទសក៍ ស្តីពី

ប្រភេទត្រីរស់នៅក្នុងដែនទឹកសាប
នៃប្រទេសកម្ពុជា
Field Guide to

Fishes of the CambodianFreshwater BodiesiPublished by the Inland Fisheries Research and Development Institute of the Fisheries
Administration, Phnom Penh, Cambodia©2019 Nagao Natural Environment Foundation, Tokyo, JapanFirst publication in 2019
All right reserved. No part of this publication may be reproduced, stored in a retrieval
system or transmitted, in any form or by any means, electronic, mechanical, photocopying,recording or otherwise, without the prior written permission of the publishers and copyrightholders.
Printed by TSPH Printing, Phnom Penh, Cambodia
For bibliographic purposes, this book should be cited as follows:
So, N., K. Utsugi, K. Shibukawa, P. Thach, S. Chhuoy, S. Kim, D. Chin, P. Nen. and P.
Chheng. 2018. Fishes of Cambodian Freshwater Bodies. Inland Fisheries Research andDevelopment Institute, Fisheries Administration, Phnom Penh, Cambodia. 197 pp.
Acknowledgements
We would like to thank all the local fisheries officers who helped us in our field surveys over
the Kingdom of Cambodia. Many thanks go to the former Director General of the Fisheries
Administration Dr. Nao Thuok, his successor Mr. Eng Cheasan, and FiA colleagues who
have provided very generous support for the successful implementation of Nagao Natural
Environment Foundation (NEF) and IFReDI/FiA joint project in Cambodia. Our sincere
and special thanks are given to Dr. Makoto Komoda and Ms. Kumiko Suzuki of the Nagao
NEF for their insightful comments and edits on the draft field guide book. We are gratefulto Professor Dr. Hiroshi Kohno of Tokyo University of Marine Science and Technology for
his kindly reviewing the field guide book. We thank Mr. Tomoyuki Sato for providing several
photographs of fish specimens. Part of the surveys was financially supported by Japan
Fund for Global Environment (JFGE) and Keidanren Nature Conservation Fund (KNCF).
We would express our gratitude for all.
List of researchers in NEF-IFReDI/FiA joint project
Nagao Natural Environment Foundation, Tokyo, Japan
Yasuhiko Taki
Ryutaro Ohtsuka
Makoto Komoda
Yoshihiro Natori
Kenzo Utsugi
Koichi Shibukawa
Tomoko Oizumi
Inland Fisheries Research and Development Institute of the Fisheries
Administration, Kingdom of Cambodia
Nam SoPhanara Thach
Sopheap Kim
Deth Chin
Samol Chhuoy
Phanna Nen
ii
Chheng Phenសៀវភៅមគ្គុទេ្ទសក៍ ស្តីពី

ប្រភេទត្រីរស់នៅក្នុងដែនទឹកសាប
នៃប្រទេសកម្ពុជា
Field Guide toiii

Fishes of the Cambodian Freshwater Bodies

Fishes of the CambodianFreshwater Bodies

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ែផ�កទី ១ : ខ��ន�តី / SECTION 1: Fish Bodyេ���ះែផ�កៃនដងខ��ន�តី / Name of​ body partsចុងគល់្រព�យគូទ

ចុងគ្រមប្រសកី

End of anal fin base

End of gill coverគល់កន�ុយ

Caudal peduncle្រចមុះ

Snout

ក្បោល
ចុង្រចមុះ

Headដងខ�ួន
Trunkរន�គូទ
Anus

Tip of snout

កន�ុយ
Tail

ចុងឆ�ឹងសំ ែប៉តEnd of hypuralbones

ក��ល: ែផ�កចុង្រចមុះដល់ជាយខងេ្រកយៃនគ្រមប្រសកី
Head: Region from the tip of snout to the posterior margin of the gill cover�ចមុះ: ែផ�កពីចុងក្បោលដល់ជាយខងមុខៃនែភ�ក
Snout: Region from the tip of head to the anterior margin of the eyeដងខ��ន: ែផ�កពីជាយខងេ្រកយៃនគ្រមប្រសកីេទរន�គូទ
Trunk: Region from the posterior margin of gill cover to the anusកន��យ: ែផ�កពីរន�គូទេទជាយខងេ្រកយឆ�ឹងសំ ែប៉តែដល្រទ្រព�យកន�ុយ(ឆ�ឹង hypural)។
Tail: Region from the anus to the posterior margin of hypural boneគល់កន��យ:ែផ�កពីចុងគល់្រព�យគូទេទជាយខងេ្រកយឆ�ឹងសំ ែប៉តែដល្រទ្រព�យកន�ុយ(ឆ�ងឹ hypural)
Caudal peduncle: Region from the end of anal fin base to posterior marginof the hypural bone.

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185

INDEXA
Acantopsis dinema Boyd & Page, 2017
Acantopsis ioa Boyd & Page, 2017
Acantopsis rungthipae Boyd, Nithirojpakdee & Page, 2017
Acantopsis thiemmedhi Sontirat, 1999
Acentrogobius caninus (Valenciennes, 1837)
Acentrogobius janthinopterus (Bleeker, 1853)
Achiroides melanorhynchus (Bleeker, 1850)ADRIANICHTHYIDAEAILIIDAEAKYSIDAE
Akysis bilustris Ng, 2011
Akysis ephippifer Ng & Kottelat, 1998
Akysis varius Ng & Kottelat, 1998
Albulichthys albuloides (Bleeker, 1855)
Allenbatrachus grunniens (Linnaeus, 1758)
AMBASSIDAE
Ambassis interrupta Bleeker, 1853
Ambassis kopsii Bleeker, 1858
Ambassis sp.
Ambassis vachellii Richardson, 1846
Ambastaia sidthimunki (Klausewitz, 1959)
Amblyceps caecutiens Blyth, 1858
Amblyceps serratum Ng & Kottelat, 2000
AMBLYCIPITIDAE
Amblypharyngodon chulabhornae Vidthayanon & Kottelat, 1990
Amblyrhynchichthys micracanthus Ng & Kottelat, 2004
ANABANTIDAE
Anabas testudineus (Bloch, 1792)ANGUILLIFORMES
Annamia normani (Hora, 1931)
Aperioptus delphax (Siebert, 1991)
Aperioptus hapalias (Siebert, 1991)
APLOCHEILIDAE
Aplocheilus panchax (Hamiton, 1822)
ARIIDAE
Arius maculatus (Thunberg, 1792)
ATHERINIFORMES
Auriglobus nefastus (Roberts, 1982)

84
83
84
84
161
161
179
125
105
112
112
112
112
63
120
135
137
138
137
137
87
111
111
111
46
59
172
172
24
77
83
83
128
128
118
119
123
185

B
Bagarius bagarius (Hamilton, 1822)
Bagarius yarrelli (Sykes, 1839)
Bagrichthys majusculus Ng, 2002
Bagrichthys obscurus Ng, 1999

114
114
94
94

186

BAGRIDAEBALITORIDAE
Balitoropsis zollingeri (Bleeker, 1853)
Barbichthys laevis (Valenciennes, 1842)
Barbodes aurotaeniatus (Tirant, 1885)
Barbodes rhombeus (Kottelat, 2000)
Barbonymus altus (Günther, 1868)
Barbonymus gonionotus (Bleeker, 1849)
Barbonymus schwanefeldii (Bleeker, 1854)
BATRACHOIDIDAEBATRACHOIDIFORMES
Batrachomoeus trispinosus (Günther, 1861)
Belodontichthys truncatus Kottelat & Ng, 1999
BELONIDAEBELONIFORMES
Betta prima Kottelat, 1994
Betta siamorientalis Kowasupat, Panijpan, Ruenwongsa & Jeenthong, 2012
Boesemania microlepis (​​​Bleeker, 1858)
Boraras micros Kottelat & Vidthayanon, 1993
Boraras urophthalmoides (Kottelat, 1991)
BOTIIDAE
Brachirus harmandi (Sauvage, 1878)
Brachirus orientalis (Bloch & Schneider, 1801)
Brachirus siamensis (Sauvage, 1878)
Brachygobius cf. aggregatus
Brachygobius mekongensis Larson & Vidthayanon, 2000
Brevitrygon walga (Müller & Henle, 1841)
Butis butis (Hamilton, 1822)
Butis gymnopomus (Bleeker, 1853)

94
77
78
64
55
56
51
51
51
120
120
120
99
126
125
172
173
150
45
45
85
179
180
180
164
163
19
160
160

C
CALLIONYMIDAECARANGIDAE
Carinotetraodon lorteti (Tirant, 1885)
Catlocarpio siamensis Boulenger, 1898
Cephalocassis borneensis (Bleeker, 1851)
CHANIDAE
Channa gachua (Hamilton, 1822)
Channa lucius (Cuvier, 1831)
Channa striata (Bloch, 1793)
Channa​ micropeltes (Cuvier, 1831)
Channa cf. marulius
CHANNIDAE
Chanos chanos (Forsskål, 1775)
CHARACIFORMES
Chaudhuria caudata Annandale, 1918

158
141
183
48
118
32
176
177
177
178
177
176
32
93
132

187

CHAUDHURIIDAE
Chitala blanci (d’Aubenton, 1965)
Chitala lopis (Bleeker, 1851)
Chitala ornata (Gray, 1831)
CICHLIDAE
Cirrhinus cirrhosus (Bloch, 1795)
Cirrhinus jullieni Sauvage, 1878
Cirrhinus microlepis Sauvage, 1878
Cirrhinus prosemion (Fowler, 1934)
Clarias cf. batrachus
Clarias macrocephalus Günther, 1864
Clarias meladerma Bleeker, 1846
Clarias nieuhofii Valenciennes, 1840
CLARIIDAE
Clupeichthys aesarnensis Wongratana, 1983
Clupeichthys cf. perakensis
CLUPEIDAE
CLUPEIFORMES
Clupeoides borneensis Bleeker, 1851
Clupisoma sinense (Huang, 1981)
COBITIDAE
Coilia lindmani Bleeker, 1857
Corica laciniata Fowler, 1935
Cosmochilus harmandi Sauvage, 1878
Crossocheilus atrilimes Kottelat, 2000
Crossocheilus oblongus Kuhl & van Hasselt, 1823
Crossocheilus reticulatus (Fowler, 1934)
Ctenopharyngodon idella (Valenciennes, 1844)
Cyclocheilichthys apogon (Valenciennes, 1842)
Cyclocheilichthys armatus (Valenciennes, 1842)
Cyclocheilichthys heteronema (Bleeker, 1854)
Cyclocheilichthys lagleri Sontirat, 1985
Cyclocheilichthys repasson (Bleeker, 1853)
Cyclocheilos enoplos (Bleeker, 1849)
CYNOGLOSSIDAE
Cynoglossus feldmanni (Bleeker, 1854)
Cynoglossus microlepis (Bleeker, 1851)
Cynoglossus puncticeps (Richardson, 1846)
CYPRINIDAECYPRINIFORMESCYPRINODONTIFORMES
Cyprinus rubrofuscus Lacepéde, 1803

132
23
22
22
156
67
66
67
67
117
117
117
118
117
30
29
29
26
29
105
80
27
30
59
73
73
74
47
60
61
61
60
60
59
181
181
181
181
32
32
128
63

D
Dagetichthys commersonnii (Lacepéde, 1802)
Danio albolineatus (Blyth, 1860)
Danio cf. pulchra

180
39
39

188

DASYATIDAE
Datnioides pulcher (Kottelat, 1998)
Datnioides undecimradiatus (Roberts & Kottelat, 1994)
DATNIOIDIDAE
Dendrophysa russellii (​​​Cuvier, 1829)
Dermogenys siamensis Fowler, 1934
Desmopuntius pentazona (Boulenger, 1894)
Dichotomyctere nigroviridis (Marion de Procé, 1822)
Dichotomyctere ocellatus (Steindachner, 1870)
Discherodontus ashmeadi (Fowler, 1937)
Doryichthys boaja (Bleeker, 1850)
Doryichthys contiguus Kottelat, 2000
Doryichthys deokhatoides (Bleeker, 1854)
Drepane punctata (Linnaeus, 1758)
DREPANEIDAE
Drombus globiceps (Hora, 1923)

19
144
144
144
151
127
57
183
183
63
130
130
131
148
148
162

E
ELEOTRIDIDAE
Eleotris melanosoma Bleeker, 1853
Eleutheronema tetradactylum (Shaw, 1804)
Ellochelon vaigiensis (Quoy & Gaimard, 1825)
ELOPIDAEELOPIFORMES
Elops hawaiensis Regan, 1909
ENGRAULIDAE
Epalzeorhynchos frenatum (Fowler, 1934)
Epinephelus bleekeri (Vaillant, 1878)
Epinephelus coioides (Hamilton, 1822)
Esomus longimanus (Lunel, 1881)
Esomus metallicus Ahl, 1924
Eugnathogobius illota (Larson, 1999)
Eugnathogobius kabilia (Herre, 1940)
Eugnathogobius siamensis (Fowler, 1934)
Eugnathogobius variegatus (Peters, 1868)

159
159
149
121
23
23
23
27
73
139
139
40
40
162
162
163
163

G
Garra cambodgiensis (Tirant, 1884)
Garra fasciacauda Fowler, 1937
Garra fuliginosa Fowler, 1934GASTEROSTEIFORMESGASTROMYZONTIDAEGERREIDAE
Gerres erythrourus (Bloch, 1791)
Gerres filamentosus Cuvier, 1829
Gibelion catla (Hamilton, 1822)
Glossogobius aureus Akihito & Meguro, 1975
Glossogobius bicirrhosus (Weber, 1894)

75
75
74
129
77
146
146
146
48
164
165

189

Glossogobius giuris (Hamilton, 1822)
Glyptothorax fuscus Fowler, 1934
Glyptothorax lampris Fowler, 1934
Glyptothorax laosensis Fowler, 1934GOBIIDAE
Gobiopterus chuno (Hamilton, 1822)
GONORYNCHIFORMES
Gymnostomus lineatus (Smith, 1945)
Gymnostomus lobatus (Smith, 1945)
Gymnostomus siamensis (Sauvage, 1881)
GYRINOCHEILIDAE
Gyrinocheilus aymonieri (Tirant, 1884)
Gyrinocheilus pennocki (Fowler, 1937)

164
115
114
115
161
166
32
66
66
65
76
76
76

H
HAEMULIDAE
Hampala dispar Smith, 1934
Hampala macrolepidota Kuhl & van Hasselt, 1823
Helicophagus leptorhynchus Ng & Kottelat, 2000
Hemiarius verrucosus (Ng, 2003)
Hemibagrus filamentus (Fang & Chaux, 1949)
Hemibagrus spilopterus Ng & Rainboth, 1999
Hemibagrus wyckii (Bleeker, 1858)
Hemibagrus wyckioides (Fang & Chaux, 1949)
Hemiculter leucisculus (Basilewsky, 1855)
Hemigobius hoevenii (Bleeker, 1851)
Hemigobius melanurus (Bleeker, 1849)
HEMIRAMPHIDAE
Hemisilurus mekongensis Bornbusch & Lundberg, 1989
Hemitrygon laosensis (Roberts & Karnasuta, 1987)
Herklotsichthys dispilonotus (Bleeker, 1852)
Hilsa kelee (Cuvier, 1829)
Homaloptera confuzona Kottelat, 2000
Homalopteroides smithi (Hora, 1932)
Homalopteroides tweediei (Herre, 1940)
Hypophthalmichthys molitrix (Valenciennes, 1844)
Hypophthalmichthys nobilis (Richardson, 1845)
Hyporhamphus limbatus (Valenciennes, 1847)
Hypsibarbus lagleri Rainboth, 1996
Hypsibarbus malcolmi (Smith, 1945)
Hypsibarbus suvattii Rainboth, 1996
Hypsibarbus vernayi (Norman, 1925)
Hypsibarbus wetmorei (Smith, 1931)

147
55
55
106
118
98
98
99
98
36
165
165
127
100
19
31
31
77
79
79
47
47
127
53
53
52
52
52

I
Incisilabeo behri (Fowler, 1937)
INDOSTOMIDAE
Indostomus spinosus Britz & Kottelat, 1999

69
129
129

190

J
Johnius belangerii (​​​Cuvier, 1830)

151

K
Kryptopterus bicirrhis (Valenciennes, 1840)
Kryptopterus dissitus Ng, 2001
Kryptopterus geminus Ng, 2003
Kryptopterus paraschilbeides Ng, 2003
Kryptopterus vitreolus Ng & Kottelat, 2013

101
102
101
102
101

L
Labeo barbatulus (Sauvage, 1878)
Labeo chrysophekadion (Bleeker, 1849)
Labeo pierrei (Sauvage, 1880)
Labeo rohita (Hamilton, 1822)
Labiobarbus leptocheilus (Valenciennes, 1842)
Labiobarbus siamensis (Sauvage, 1881)
Laides longibarbis (Fowler, 1934)
Lates calcarifer (Bloch, 1790)
LATIDAE
Laubuka caeruleostigmata Smith, 1931
Laubuka siamensis Fowler, 1939
LEIOGNATHIDAE
Leiognathus equula (Forsskål, 1775)
Lepidocephalichthys hasselti (Valenciennes, 1846)
Lepidocephalichthys kranos Havird & Page, 2010
Lepidocephalichthys sp.
Lepidocephalichthys zeppelini Havird & Tangjitjaroen, 2010
Leptobarbus rubripinna (Fowler, 1937)
Lobocheilos rhabdoura (Fowler, 1934)
Luciosoma bleekeri Steindachner, 1878
LUTJANIDAE
Lutjanus fulviflamma (Forskål, 1775)
Lutjanus russellii (Bleeker, 1849)
Lycothrissa crocodilus (Bleeker, 1850)

69
68
68
68
65
65
105
138
138
33
33
140
140
81
82
81
82
48
72
37
145
145
145
27

M
Macrochirichthys macrochirus (Valenciennes, 1844)
Macrognathus circumcinctus (Hora, 1924)
Macrognathus semiocellatus Roberts, 1986
Macrognathus siamensis (Günther, 1861)
Macrognathus sp.
Macrotrema sp.
MASTACEMBELIDAE
Mastacembelus armatus (Lacepéde, 1800)
MEGALOPIDAE
Megalops cyprinoides (Broussonet, 1782)
Mekongina erythrospila Fowler, 1937

37
133
133
133
134
132
133
134
24
24
69

191

Micronema cheveyi (Durand, 1940)
MONODACTYLIDAE
Monodactylus argenteus (Linnaeus, 1758)
Monopterus albus (Zuiew, 1793)
Moolgarda pedaraki (Valenciennes, 1836)
Moolgarda perusii (Valenciennes, 1836)
MUGILIDAE
MUGILIFORMES
Mugilogobius chulae (Smith, 1932)
Mugilogobius rambaiae (Smith, 1945)
Mugilogobius tigrinus Larson, 2001
MURAENESOCIDAE
Muraenesox cinereus (Forsskål, 1775)
MYLIOBATIFORMES
Mystacoleucus atridorsalis Fowler, 1937
Mystacoleucus greenwayi Pellegrin & Fang, 1940
Mystacoleucus obtusirostris (Valenciennes, 1842)
Mystus albolineatus Roberts, 1994
Mystus atrifasciatus Fowler, 1937
Mystus bocourti (Bleeker, 1864)
Mystus gulio (Hamilton, 1822)
Mystus multiradiatus Roberts, 1992
Mystus mysticetus Roberts, 1992
Mystus singaringan (Bleeker, 1846)
Mystus velifer Ng, 2012

102
155
155
131
122
122
121
121
166
167
166
24
24
19
62
62
62
96
95
95
97
96
97
96
97

N
NANDIDAE
Nandus nebulosus (​​​Gray, 1835)
Nandus oxyrhynchus Ng, Vidthayanon & Ng, 1996
NEMACHEILIDAE
Nemacheilus cf. masyae
Nemacheilus longistriatus Kottelat, 1990
Nemacheilus masyae Smith, 1933
Nemacheilus platiceps Kottelat, 1990
Neolissochilus soroides (Duncker, 1904)
Neosalanx tangkahkeii (Wu, 1931)
Neostethus bicornis Regan, 1916
Neostethus lankesteri Regan, 1916
Nibea soldado (Lacepéde, 1802)
NOTOPTERIDAE
Notopterus notopterus (Pallas, 1769)
Nuchequula gerreoides (Bleeker, 1851)

154
154
154
88
90
89
89
89
54
119
124
123
152
22
22
140

192

O
Ompok siluroides Lacepéde, 1803
Ompok urbaini (Fang & Chaux, 1949)
Onychostoma meridionale Kottelat, 1998
Onychostoma sp.
OPHICHTHIDAE
Ophichthus rutidoderma (Bleeker, 1852)
Ophiocara ophicephalus (Valenciennes, 1837)
Opsarius koratensis (Smith, 1931)
Opsarius pulchellus (Smith, 1931)
Oreochromis mossambicus (Peters, 1852)
Oreochromis niloticus (Linnaeus, 1758)
Oryzias haugiangensis Roberts, 1998
Oryzias minutillus Smith, 1945
OSMERIFORMESOSPHRONEMIDAE
Osphronemus exodon Roberts, 1994
Osphronemus goramy Lacepéde, 1801
Osteochilus lini Fowler, 1935
Osteochilus melanopleura (Bleeker, 1852)
Osteochilus microcephalus (Valenciennes, 1842)
Osteochilus schlegelii (Bleeker, 1851)
Osteochilus vittatus (Valenciennes, 1842)
Osteochilus waandersii (Bleeker, 1853)
OSTEOGLOSSIDAE
OSTEOGLOSSIFORMES
Otolithes ruber (Bloch & Schneider, 1801)
Oxyeleotris marmorata (Bleeker, 1852)
Oxygaster pointoni (Fowler, 1934)

103
103
72
72
25
26
160
38
38
156
156
125
125
119
172
176
175
71
70
71
70
70
71
21
21
152
159
34

P
Pangasianodon gigas Chevey, 1931
Pangasianodon hypophthalmus (Sauvage, 1878)
PANGASIIDAE
Pangasius bocourti Sauvage, 1880
Pangasius conchophilus Roberts & Vidthayanon, 1991
Pangasius elongatus Pouyaud, Gustiano & Teugels, 2002
Pangasius krempfi Fang & Chaux, 1949
Pangasius larnaudii Bocourt, 1866
Pangasius macronema Bleeker, 1850
Pangasius mekongensis Gustiano, Teugels & Pouyaud, 2003
Pangasius sanitwongsei Smith, 1931
Pangio cf. anguillaris
Pangio cf. fusca

107
107
106
108
108
110
109
110
110
109
108
80
80

193

Pangio myersi (Harry,1949)
Pangio oblonga (Valenciennes, 1846)
Pao ​​abei (Roberts, 1998)
Pao baileyi (Sontirat, 1985)
Pao cambodgiensis (Chabanaud, 1923)
Pao fangi (Pellegrin & Chevey, 1940)
Pao turgidus (Kottelat, 2000)
Papuligobius ocellatus (Fowler, 1937)
Parachela maculicauda (Smith, 1934)
Parachela oxygastroides (Bleeker, 1852)
Parachela siamensis (Günther, 1868)
Parachela sp.
Parachela williaminae Fowler, 1934
Paralaubuca barroni (Fowler, 1934)
Paralaubuca typus Bleeker, 1864
PARALICHTHYIDAEParambassis apogonoides (Bleeker, 1851)
Parambassis siamensis (Fowler, 1937)
Parambassis wolffii (Bleeker, 1850)
Paraplagusia bilineata (Bloch, 1787)
Parasikukia maculata Doi, 2000
PERCIFORMESPhalacronotus sp.1
Phalacronotus sp.2
PHALLOSTETHIDAE
Phenacostethus smithi Myers, 1928
Piaractus brachypomus (Cuvier, 1818)
Pisodonophis sp.
Pisodonophis​ cancrivorus (Richardson, 1848)
Planiliza subviridis (Valenciennes, 1836)
PLATYCEPHALIDAE
Platycephalus indicus (Linnaeus, 1758)
Plectorhinchus gibbosus (Lacepéde, 1802)
PLEURONECTIFORMESPLOTOSIDAE
Plotosus canius Hamilton, 1822
Plotosus lineatus (Thunberg, 1787)
Poecilia reticulata Peters, 1859POECILIIDAEPOLYNEMIDAE
Polynemus aquilonaris Motomura, 2003
Polynemus melanochir Valenciennes, 1831
Pomadasys kaakan (Cuvier, 1830)
Poropuntius normani Smith, 1931
PRISTOLEPIDIDAE
Pristolepis fasciata (​​​Bleeker, 1851)

194

80
81
184
185
184
184
185
167
36
35
35
36
35
34
34
178
136
136
136
182
58
135
100
100
123
124
93
25
25
121
135
135
147
178
116
116
116
129
129
149
149
150
147
53
153
153

Probarbus jullieni Sauvage, 1880
Probarbus labeamajor Roberts, 1992
Pseudobagarius filifer (Ng & Rainboth, 2005)
Pseudobagarius inermis (Ng & Kottelat, 2000)
Pseudogobius avicennia (Herre, 1940)
Pseudogobius gastrospilus (Bleeker, 1853)
Pseudogobius javanicus (Bleeker, 1856)
Pseudohomaloptera leonardi (Hora, 1941)
Pseudolais pleurotaenia (Sauvage, 1878)
Pseudomystus siamensis (Regan, 1913)
Pseudorhombus arsius (Hamilto, 1822)
Puntigrus partipentazona (Fowler, 1934)
Puntioplites bulu (Bleeker, 1851)
Puntioplites falcifer Smith, 1929
Puntioplites proctozysron (Bleeker, 1864)
Puntius brevis (Bleeker, 1849)
Puntius masyai Smith,1945

64
64
113
113
168
169
168
78
106
95
178
57
49
49
49
56
56

R
Raiamas guttatus (Day, 1870)
Rasbora amplistriga Kottelat, 2000
Rasbora aurotaenia Tirant, 1885
Rasbora borapetensis Smith, 1934
Rasbora cf. daniconius
Rasbora myersi Brittan, 1954
Rasbora paviana Tirant, 1885
Rasbora rubrodorsalis Donoso-Buchner&Schmidt, 1997
Rasbora tornieri Ahl, 1922
Rasbora trilineata Steindachner, 1870
Rasbosoma spilocerca (Rainboth & Kottelat, 1987)
Redigobius bikolanus (Herre, 1927)
Redigobius chrysosoma (Bleeker, 1875)
Repomucenus sagitta (Pallas, 1770)
Rhinogobius sp.

37
42
43
41
42
43
42
41
43
44
44
167
168
158
169

S
SALANGIDAE
Sardinella gibbosa (Bleeker, 1849)
Scaphognathops bandanensis Boonyaratpalin & Srirungroj, 1971
Scaphognathops stejnegeri (Smith, 1931)
SCATOPHAGIDAE
Scatophagus argus (Linnaeus, 1766)
Schistura cf. bolavenensis
Schistura cf. daubentoni
Schistura cf. sokolovi
Schistura coruscans Kottelat, 2000

119
31
50
50
148
148
91
93
92
90

195

Schistura imitator Kottelat, 2000
Schistura khamtanhi Kottelat, 2000
Schistura kohchangensis (Smith, 1933)
Schistura namboensis Freyhof & Serov, 2001
Schistura sp.
SCIAENIDAE
Scleropages formosus (Müller & Schlegel, 1840)
Scomberoides commersonnianus Lacepéde, 1801
Scomberoides lysan (Forskål, 1775)
Scomberomorus sinensis (Lacepéde, 1800)
SCOMBRIDAESCORPAENIFORMESSERPENTICOBITIDAE
Serpenticobitis octozona Roberts, 1997
SERRANIDAESERRASALMIDAE
Setipinna melanochir (Bleeker, 1849)
SIGANIDAE
Siganus fuscescens (Houttuyn, 1782)
Siganus guttatus (Bloch, 1787)
Siganus javus (Linnaeus, 1766)
Siganus vermiculatus (Valenciennes, 1835)
Sikukia gudgeri (Smith, 1934)
Sikukia stejnegeri Smith, 1931SILLAGINIDAE
Sillago aeolus Jordan & Evermann, 1902
Sillago sihama (Forskål, 1775)
Silurichthys schneideri Volz, 1904
SILURIDAESILURIFORMESSISORIDAE
SOLEIDAE
Stenogobius mekongensis Watson, 1991
Stolephorus dubiosus Wongratana, 1983
Stolephorus indicus (van Hasselt, 1823)
Strongylura incisa (Valenciennes, 1846)
Strongylura leiura (Bleeker, 1850)
SUNDASALANGIDAE
Sundasalanx mekongensis Britz & Kottelat, 1999SYNBRANCHIDAESYNBRANCHIFORMES
Syncrossus helodes (Sauvage, 1876)
Syncrossus lucasbahi (Fowler, 1937)
SYNGNATHIDAE
Systomus rubripinnis (Valenciennes, 1842)

196

91
91
92
90
92
150
21
141
141
142
142
135
88
88
139
93
27
170
171
170
171
171
58
58
143
143
143
103
99
94
114
179
170
28
28
126
126
26
26
131
131
85
85
130
57

T
Tenualosa thibaudeaui (Durand, 1940)
Terapon jarbua (Forsskål, 1775)
TERAPONIDAETETRAODONTIDAETETRAODONTIFORMES
Thryssa hamiltonii (Gray, 1835)
Thryssocypris tonlesapensis Roberts & Kottelat, 1984
Thynnichthys thynnoides (Bleeker, 1852)
Tonlesapia tsukawakii Motomura & Mukai, 2006
Tor tambra (Valenciennes, 1842)
Tor tambroides (Bleeker, 1854)
Toxotes chatareus (​​Hamiton, 1822)
Toxotes jaculatrix (Pallas, 1767)
TOXOTIDAE
Trichopodus microlepis (Günther, 1861)
Trichopodus pectoralis Regan, 1910
Trichopodus trichopterus (Pallas, 1770)
Trichopsis pumila (Arnold, 1936)
Trichopsis schalleri Ladiges, 1962
Trichopsis vittata (Cuvier, 1831)
Trigonopoma pauciperforatum (Weber & de Beaufort, 1916)
Trigonostigma espei (Meinken, 1967)
Trypauchen pelaeos Murdy, 2006
Tuberoschistura cambodgiensis Kottelat, 1990

30
155
155
182
182
28
38
46
158
54
54
157
157
157
174
174
175
173
173
174
46
45
170
88

U
Urogymnus polylepis (Bleeker, 1852)

20

W
Wallago attu (Bloch & Schneider, 1801)
Wallago micropogon Ng, 2004

104
104

X
Xenentodon sp.

126

Y
Yasuhikotakia caudipunctata (Taki & Doi, 1995)
Yasuhikotakia eos (Taki, 1972)
Yasuhikotakia lecontei (Fowler, 1937)
Yasuhikotakia modesta (Bleeker, 1864)
Yasuhikotakia morleti (Tirant, 1885)

86
87
86
87
86

Z
ZENARCHOPTERIDAE
Zenarchopterus ectuntio (Hamiton, 1822)

127
128

197

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