FISHESof theINDOCHINESE MEKONG

Nagao Natural Environment Foundation

Fishes of the Indochinese MekongTwo fish species endemic to the Mekong, Mekongina erythrospila (right and left) and Tenualosa thibaudeaui (middle two) (Luang Prabang Province, Laos, photo: KS)

FISHESOF THEINDOCHINESE
MEKONG

Nagao Natural Environment FoundationTOKYOFishes of the Indochinese MekongFirst published and distributed in 2021 by:
Nagao Natural Environment Foundation
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Printed in Japan by Shumpousha Co., Ltd., 9-50 Chofu-ogimachi, Shimonoseki, Yamaguchi 752-0927, Japan.
Editorial board: Yasuhiko Taki*1, Ryutaro Ohtsuka, Makoto Komoda, Yoshihiro Natori*2, Kenzo Utsugi*3, Koichi Shibukawa*2,Tomoko Oizumi, and Stefan Ottomanski (Nagao Natural Enviroment Foundation, Tokyo, Japan); Bounthob Praxaysombath andKoneouma Phongsa (National University of Laos, Vientiane, Lao PDR); Wichian Magtoon (Srinakharinwirot University, Bangkok,Thailand); Prachya Musikasinthorn (Kasetsart University, Bangkok, Thailand); Chaiwut Grudpan and Jarungjit Grudpan (UbonRatchathani University, Ubon Ratchathani, Thailand); Apinun Suvarnaraksha (Maejo University, Chiang Mai, Thailand); Nam Soand Phanara Thach (Inland Fisheries Research and Development Institution, Fisheries Administration, Phnom Penh, Cambodia);
Phuong Thanh Nguyen, Dinh Dac Tran, and Loi Xuan Tran (Can Tho University, Cần Thơ, Vietnam).
*1: passed away on 24 March 2020; *2: resigned in 2015; *3 resigned in 2016.
Text and DTP: Koichi Shibukawa [Nagao Natural Environment Foundation, Tokyo, Japan (to March 2015); Museum of Natural andEnvironmental History, Shizuoka, Japan (April 2015 to present)].
Recommended citation:
Nagao Natural Environment Foundation. 2021. Fishes of the Indochinese Mekong. Nagao Natural Environment Foundation, Tokyo.
xii+546 pages.
Acknowledgment. This book has been prepared based primarily on outcomes of the field surveys performed by the international
collaborative project (1st phase: 2006–2011, 2nd phase: 2011–2016), entitled "Basic Study on the Aquatic Fauna and Flora andConservation Activities Participated in by Local Residents in the Mekong-Chao Phraya Region," sponsored, organized and directedby the Nagao Natural Environment Foundation (NEF), Tokyo. This project would not have been possible without the help of all staffof universities/institutions involved in the project, and many local officers in respective countries (Lao PDR, Thailand, Cambodia,and Vietnam), as well as the NEF administrators. We also thank following persons, who helped identification of various kinds ofthe Mekong fishes: Kiyotaka Hatooka (Osaka Museum of Natural History, Japan); Yusuke Hibino (Mie University, Japan); HsuanChing Ho (National Museum of Marine Biology and Aquarium, Taiwan); Hisashi Imamura (Hokkaido University, Japan); Akihisa
Iwata (Kyoto University, Japan); Yukio Iwatsuki (Miyazaki University, Japan); Zeehan Jaafer (National University of Singapore,Singapore); Seishi Kimura (Mie University, Japan); Hellen K. Larson (Museum and Art Gallery of the Northern Territory, Australia);
Hiroyuki Motomura (Kagoshima University, Japan); Edward O. Murdy (George Washington University, USA); Heok Hee Ng(National University of Singapore, Singapore); Kunio Sasaki (Kochi University, Japan); Hiroshi Senou (Kanagawa PrefecturalMuseum, Japan); David G. Smith (National Museum of Natural History, USA). Following persons helped, and provided the working
space and facilities for examining the fish specimens for our research: Hiroshi Kohno and Masato Moteki (Tokyo University ofMarine Technology and Science, Japan); Keiichi Matsuura, Gento Shinohara, Takashi P. Sato, Masanori Nakae, Satoru N. Chiba,Eri Katayama, and Shinpei Ohashi (National Museum of Nature and Science, Japan); Douglass F. Hoese and Mark A. McGrouther(Australian Museum, Australia). Akihisa Iwata (Kyoto University, Japan), Masanori Nakae (National Museum of Nature andScience, Japan), Toshiyuki Suzuki (Kawanishi-midoridai Senior High School, Japan), and the Japan Wildlife Research Center (Japan)
kindly provided fish photographs taken by them and/or permission for using photographs that they manage. Chavalit Vidthayanon(Mekong River Commission, Laos) gave his time for discussion with us about taxonomy and fauna of the Mekong fishes. Note that
information about belonging places of these persons are those during the period of the project (to 2016).

NEF

NUOL

KU

UBU

MJU

SWU

FiA (IFReDI)

Nagao Natural Environment Foundation (NEF), Tokyo, Japan
National University of Laos (NUOL), Vientiane, Lao PDR
Kasetsart University (KU), Bangkok, Thailand
Ubon Ratchathani University (UBU), Ubon Ratchathani, Thailand
Maejo University (MJU), Chiang Mai, Thailand
Srinakharinwirot University (SWU), Bangkok, Thailand
Inland Fisheries Research and Development Institution (IFReDI), Fisheries Administration (FiA), Phnom Penh, Cambodia
Can Tho University (CTU), Cần Thơ, Vietnam

CTU