សៀវភៅមគ្គុទេ្ទសក៍ ស្តីពី

ប្រភេទត្រីរស់នៅក្នុងដែនទឹកសាប
នៃប្រទេសកម្ពុជា
Field Guide to

Fishes of the CambodianFreshwater BodiesiPublished by the Inland Fisheries Research and Development Institute of the Fisheries
Administration, Phnom Penh, Cambodia©2019 Nagao Natural Environment Foundation, Tokyo, JapanFirst publication in 2019
All right reserved. No part of this publication may be reproduced, stored in a retrieval
system or transmitted, in any form or by any means, electronic, mechanical, photocopying,recording or otherwise, without the prior written permission of the publishers and copyrightholders.
Printed by TSPH Printing, Phnom Penh, Cambodia
For bibliographic purposes, this book should be cited as follows:
So, N., K. Utsugi, K. Shibukawa, P. Thach, S. Chhuoy, S. Kim, D. Chin, P. Nen. and P.
Chheng. 2018. Fishes of Cambodian Freshwater Bodies. Inland Fisheries Research andDevelopment Institute, Fisheries Administration, Phnom Penh, Cambodia. 197 pp.
Acknowledgements
We would like to thank all the local fisheries officers who helped us in our field surveys over
the Kingdom of Cambodia. Many thanks go to the former Director General of the Fisheries
Administration Dr. Nao Thuok, his successor Mr. Eng Cheasan, and FiA colleagues who
have provided very generous support for the successful implementation of Nagao Natural
Environment Foundation (NEF) and IFReDI/FiA joint project in Cambodia. Our sincere
and special thanks are given to Dr. Makoto Komoda and Ms. Kumiko Suzuki of the Nagao
NEF for their insightful comments and edits on the draft field guide book. We are gratefulto Professor Dr. Hiroshi Kohno of Tokyo University of Marine Science and Technology for
his kindly reviewing the field guide book. We thank Mr. Tomoyuki Sato for providing several
photographs of fish specimens. Part of the surveys was financially supported by Japan
Fund for Global Environment (JFGE) and Keidanren Nature Conservation Fund (KNCF).
We would express our gratitude for all.
List of researchers in NEF-IFReDI/FiA joint project
Nagao Natural Environment Foundation, Tokyo, Japan
Yasuhiko Taki
Ryutaro Ohtsuka
Makoto Komoda
Yoshihiro Natori
Kenzo Utsugi
Koichi Shibukawa
Tomoko Oizumi
Inland Fisheries Research and Development Institute of the Fisheries
Administration, Kingdom of Cambodia
Nam SoPhanara Thach
Sopheap Kim
Deth Chin
Samol Chhuoy
Phanna Nen
ii
Chheng Phenសៀវភៅមគ្គុទេ្ទសក៍ ស្តីពី

ប្រភេទត្រីរស់នៅក្នុងដែនទឹកសាប
នៃប្រទេសកម្ពុជា
Field Guide toiii

Fishes of the Cambodian Freshwater Bodies

Fishes of the CambodianFreshwater Bodies

FiscC)$Zc9`<  ,QWURGXFWLRQ

Ǭ3Yǔ>  &217(176cǜõiǓ £3Zcș$(© cCb@cȝc7i )LVKQDPHVLQWKLVERRN

L LL
LLLLY

ǔ>c£8`£ǨCme7cCb@cȝc7i  +RZWRXVHWKLVERRN

Y

d9$5Z %¤7« £3Z  6HFWLRQ )LVK%RG\ɳƻƗɹɴɇƒȲɵɅȳƚɯɅƙɁɪ 1DPHRIERG\SDUWVƳɌǏɑɽɴɎȶȳƚɯɅƙɁɪ 0HDVXUHPHQWVRIILVKERG\ɃƙɊȶɽǋɁɽ 3RVLWLRQRIPRXWKɈɭȲǋɁɽ  %DUEHOVɳƻƗɹƙɈȩɋ )LQVƙɃɅɭȶƙɈȩɋ  )LQUD\VǍȶƙɈȩɋȲɅƐɭɋ &DXGDOILQVKDSHɳƙƵȶȸơɫȶȲɅƐɭɋƙɁɪ &DXGDOVNHOHWRQƳɌǍɆɽƙɑƳ 6FDOHVWREHFRXQWHGɑƒɩɁƙɑȲɪ ͬ*LOOUDNHUVd9$5Z +h7_=$©(ǔ>dC¥(cǜõǓi£3Z 6HFWLRQ )LVK6HDUFK$LG$QJXLOOLIRUPHV²&OXSHLIRUPHVɍɸ ƽɆɽ 2UGHU 0\OLREDWLIRUPHV²2VWHRJORVVLIRUPHV²(ORSLIRUPHV²ɍɸ ƽɆɽ 2UGHU *RQRU\QFKLIRUPHV²&\SULQLIRUPHVɍɸ ƽɆɽ 2UGHU&KDUDFLIRUPHV²6LOXULIRUPHVɍɸ ƽɆɽ 2UGHU 2VPHULIRUPHV²%DWUDFKRLGLIRUPHV²0XJLOLIRUPHV²$WKHULQLIRUPHV²%HORQLIRUPHV

6FRUSDHQLIRUPHV²3HUFLIRUPHVɍɸ ƽɆɽ 2UGHU &\SLURGRQWLIRUPHV²*DVWHURVWHLIRUPHV²6\QEUDQFKLIRUPHV²ɍɸ ƽɆɽ 2UGHU 3OHXURQHFWLIRUPHV²7HWUDRGRQWLIRUPHV



d9$5Z £8c;5£3Z 7Y(ǔ>c6¥`F3CǽʲǓ2 6HFWLRQ )LVK6SHFLHVDQG
,GHQWLILFDWLRQ



ン

i

,QWURGXFWLRQ&DPERGLD HQFRPSDVVHV D JUHDW SDUW RI WKH 0HNRQJ ULYHU V\VWHP LQFOXGLQJ ODUJHWULEXWDULHV VXFK DV WKH 7RQOH 6DS 5LYHU ZKLFK UXQV IURP WKH *UHDW /DNH WKH %DVVDF
5LYHU WKH ORZHU UHDFKHV RI WKH 6HNRQJ 5LYHU DQG WKH 6HVDQ DQG WKH 6UHSRN ULYHUVZKLFK UXQ IURP PRXQWDLQV RI WKH QRUWKHDVWHUQ SDUW ,Q DGGLWLRQ LQ WKH VRXWKZHVWHUQ SDUWVHYHUDO VKRUW VWUHDPV UXQ IURP WKH &DUGDPRP PRXQWDLQV DQG IORZ LQWR WKH *XOI RI
7KDLODQG ,Q WKH UDLQ\ VHDVRQ WKH ZDWHU VXUIDFH H[SDQGV WR IRUP SURPLQHQW IORRGSODLQVHQDEOLQJ WKH KDELWDWV RI D JUHDW YDULHW\ RI ILVKHV DQG LQFUHDVLQJ WKH \LHOGV RI PDQ\ ILVKHV1RQHWKHOHVV WKH ULFKQHVV RI ILVK UHVRXUFHV KDV EHHQ WKUHDWHQHG ODUJHO\ EHFDXVH RI WKH
UDSLG HQYLURQPHQWDO H[SORLWDWLRQ DQG GHWHULRUDWLRQ RI WKH ILVK KDELWDWV 7R DOOHYLDWH WKHORVV RI ILVK UHVRXUFHV HQYLURQPHQWDO DZDUHQHVV LV QRZ TXLWH LPSRUWDQW
³)LHOG *XLGH WR )LVKHV RI )UHVKZDWHU %RGLHV LQ &DPERGLD´ LV WKH ILUVW VWHS WRZDUGVDZDUHQHVV LW DLPV WR GHYHORS PDQDJHPHQW DQG FRQVHUYDWLRQ DZDUHQHVV DQG WR
DGYDQFH FRQVHUYDWLRQ SHUIRUPDQFH DPRQJ WKH ORFDO SHRSOH ,W FRYHUV  ILVK VSHFLHVZKLFK ZHUH FROOHFWHG DQG UHFRUGHG IURP WKH 0HNRQJ 5LYHU DQG &DUGDPRP PRXQWDLQVULYHU EDVLQV LQ &DPERGLD GXULQJ H[WHQVLYH ILHOG VXUYH\V FRQGXFWHG MRLQWO\ E\ 1DJDR
1DWXUDO (QYLURQPHQW )RXQGDWLRQ 1()  D -DSDQHVH 1*2 DQG ,QODQG )LVKHULHV
5HVHDUFK DQG 'HYHORSPHQW ,QVWLWXWH ,)5H', RI WKH )LVKHULHV $GPLQLVWUDWLRQ RI&DPERGLD ,Q DGGLWLRQ WKH MRLQW UHVHDUFK WHDP KDV EHHQ HQJDJHG LQ SURJUDPV RISDUWLFLSDWRU\ ILVK ELRGLYHUVLW\ PRQLWRULQJ VXUYH\V VXSSRUWHG E\ -DSDQ )XQG IRU *OREDO(QYLURQPHQW -)*( )LQDOO\ ZH KRSH WKLV KDQG\ ILHOG JXLGH ERRN LV XVHIXO IRU HYHU\RQH ZKR DJUHHV ZLWK RXU
DFWLYLWLHV DQG HDJHUO\ FRQWULEXWHV WR WKH PDQDJHPHQW DQG FRQVHUYDWLRQ RI ILVKELRGLYHUVLW\ LQ &DPERGLD

Ϲ
ii

ヵ

iii

)LVK QDPHV LQ WKLV ERRN
6LQFH  ZKHQ 5DLQERWK SXEOLVKHG µ)LVKHV RI WKH &DPERGLDQ 0HNRQJ¶ QXPHURXVWD[RQRPLF VWXGLHV RI ILVKHV LQ WKH 0HNRQJ UHJLRQ KDYH EHHQ FRQGXFWHG FRQVHTXHQWO\YDULRXV VSHFLHV KDYH EHHQ QHZO\ UHFRUGHG DQGRU GHVFULEHG WR GDWH 0RUHRYHU PDQ\
VSHFLHV KDYH EHHQ WD[RQRPLFDOO\ UHH[DPLQHG DQG DV D UHVXOW WKH VFLHQWLILF QDPHV RI D
QXPEHU RI VSHFLHV KDYH EHHQ FKDQJHG RYHU DQG RYHU DJDLQ 7KH UHFHQW ZRUN RI .RWWHODW RQ WKH QRPHQFODWXUH RI 6RXWKHDVW $VLDQ LQODQG ILVKHV HQFRPSDVVHV DOPRVW DOO RIWKH VSHFLHV IRXQG LQ WKH UHJLRQ GLVFXVVHV WKDW PDQ\ ILVK VSHFLHV DQG JURXSV DUH VWLOO LQ
WD[RQRPLF RU QRPHQFODWXUDO DUJXPHQW
³)LHOG *XLGH WR )LVKHV RI )UHVKZDWHU %RGLHV LQ &DPERGLD´ DGRSWV VFLHQWLILF QDPHV RIILVKHV LQ &DWDORJ RI )LVKHV D ZHEVLWH FRPSLOHG E\ (VFKPH\HU HW DO   )LVK QDPHVLQ WKH .KPHU ODQJXDJH KDYH QRW EHHQ VWDQGDUGL]HG \HW .KPHU QDPHV LQ WKLV ERRN DUH
WKHUHIRUH LQ WKH VWDWXV RI µSURSRVHG VWDQGDUG QDPHV RI ILVKHV LQ &DPERGLD¶
5HIHUHQFHV(VFKPH\HU:15)ULFNHDQG5YDQGHU/DDQ HGV &$7$/2*2)),6+(6*(1(5$63(&,(65()(5(1&(6(OHFWURQLFYHUVLRQDFFHVVHG 0DUFK
7KLVYHUVLRQZDVHGLWHGE\%LOO(VFKPH\HU
KWWSUHVHDUFKDUFKLYHFDODFDGHP\RUJUHVHDUFKLFKWK\RORJ\FDWDORJILVKFDWPDLQDVS .RWWHODW0  7KHILVKHVRIWKHLQODQGZDWHUVRI6RXWKHDVW$VLDDFDWDORJXHDQG
FRUHELEOLRJUDSK\RIWKHILVKHVNQRZQWRRFFXULQIUHVKZDWHUVPDQJURYHVDQG
HVWXDULHVLQ7KH5DIIOHV%XOOHWLQRI=RRORJ\1R 
5DLQERWK:-  )$2VSHFLHVLGHQWLILFDWLRQILHOGJXLGHIRUILVKHU\SXUSRVHV)LVKHVRIWKH&DPERGLDQ0HNRQJ)$25RPH SS

ϻ

iv

А
v

ែផ�កទី ១ : ខ��ន�តី / SECTION 1: Fish Bodyេ���ះែផ�កៃនដងខ��ន�តី / Name of​ body partsចុងគល់្រព�យគូទ

ចុងគ្រមប្រសកី

End of anal fin base

End of gill coverគល់កន�ុយ

Caudal peduncle្រចមុះ

Snout

ក្បោល
ចុង្រចមុះ

Headដងខ�ួន
Trunkរន�គូទ
Anus

Tip of snout

កន�ុយ
Tail

ចុងឆ�ឹងសំ ែប៉តEnd of hypuralbones

ក��ល: ែផ�កចុង្រចមុះដល់ជាយខងេ្រកយៃនគ្រមប្រសកី
Head: Region from the tip of snout to the posterior margin of the gill cover�ចមុះ: ែផ�កពីចុងក្បោលដល់ជាយខងមុខៃនែភ�ក
Snout: Region from the tip of head to the anterior margin of the eyeដងខ��ន: ែផ�កពីជាយខងេ្រកយៃនគ្រមប្រសកីេទរន�គូទ
Trunk: Region from the posterior margin of gill cover to the anusកន��យ: ែផ�កពីរន�គូទេទជាយខងេ្រកយឆ�ឹងសំ ែប៉តែដល្រទ្រព�យកន�ុយ(ឆ�ឹង hypural)។
Tail: Region from the anus to the posterior margin of hypural boneគល់កន��យ:ែផ�កពីចុងគល់្រព�យគូទេទជាយខងេ្រកយឆ�ឹងសំ ែប៉តែដល្រទ្រព�យកន�ុយ(ឆ�ងឹ hypural)
Caudal peduncle: Region from the end of anal fin base to posterior marginof the hypural bone.

1

1

ǔ>ǰCm£8d@(.(%¤7« £3Z 0HDVXUHPHQWVRIILVK௙௙ERG\ǏɑɽƙɆɴɎȶɌǏȶɈɌɪ ȷɸɀɭȷ
0HDVXUHDGLVWDQFHEHWZHHQWZRSRLQWVȲɊƕɑɽȳɯ Ʌ
ƚ
%RG\GHSWK %'ƙɆɴɎȶȼȶƙȷɊɭɹ
6QRXWOHQJWK 6Q/

ƙɆɴɎȶȲǙɍ+HDGOHQJWK +/ƙɆɴɎȶɑɌɭɆ
7RWDOOHQJWK 7/

ƙɆɴɎȶȼȶȳƚɯɅ
6WDQGDUGOHQJWK 6/ɃɃɫȶȳƚɯɅ'LVFZLGWK ':C(Ǟm £8d@(.(%¤7«  ƙɆɴɎȶȼȶȳƚɯɅɈɪȷɭȶƙȷɊɭɹȼɍɽȷɭȶȸơɫȶɑɸ ɴɆɻɁƙɃƙɈȩɋȲɅƐɭɋ ȸơɫȶK\SXUDO
6WDQGDUGOHQJWK 6/ 'LVWDQFHEHWZHHQWLSRIVQRXWDQGHQGRIK\SXUDO ERQH £8d@(.(%¤«7C>]8 ƙɆɴɎȶȼȶȳƚɯɅɈɪȷɭȶƙȷɊɭɹȼɍɽȷɭȶƙɈȩɋȲɅƐɭɋ
7RWDOOHQJWK 7/ 'LVWDQFHEHWZHHQWLSRIVQRXWDQGWLSRIGHSUHVVHGFDXGDOILQ$<C.
m (%¤7«  ȲɊƕɑɽɆȥƆɌɈȷɪ ɸɀɭȷɳɍˊɆɸɇɭɁɅȶɩ ȷɸɀɭȷɳƙƳɊɆɸɇɭɁɵɅȼȶȳƚɯɅ%RG\GHSWK %' 9HUWLFDOGLVWDQFHEHWZHHQWRSPRVWDQGERWWRPPRVWSRLQWVRIWKHERG\£8d@($ǓǺ? ƙɆɴɎȶȼȶȳƚɯɅɈȷɪ ɭȶƙȷɊɭɹȼɍɽƺɋƴȶɳƙƳɋȴƙɊɆƙɑȲɪ
+HDGOHQJWK +/ 'LVWDQFHEHWZHHQWLSRIVQRXWDQGSRVWHULRUPDUJLQRIJLOOFRYHU£8d@(£)<]i ƙɆɴɎȶȼȶȳƚɯɅɈɪȷɭȶƙȷɊɭɹȼɍɽƺɋƴȶɊɭȳɴɉƒȲ
6QRXWOHQJWK 6Q/ 'LVWDQFHEHWZHHQWLSRIVQRXWDQGDQWHULRUPDUJLQRIH\H£8d@(55[(%¤7«  ƙɆɴɎȶȼȶȳƚɯɅɈɪƙɈȩɋƙɃȪȶƴȶɳȸƛȶɆɸɇɭɁɳǵƙɈȩɋƙɃȪȶƴȶǒƉɸɆɸɇɭɁɵɅƙɁɪɆɴɆɍ
'LVFZLGWK ':  'LVWDQFHEHWZHHQOHIWPRVWDQGULJKWPRVWRISHFWRUDOILQVRIVWLQJUD\V

25£<(mǬ3m£3Z 3RVLWLRQRIPRXWKƙɁȶɽɳɑƗˊ
7HUPLQDOɳȶˊɋ
6XSHULRUɳǷƴȶɳƙƳɊ
6XEWHUPLQDOȷɮɍɳƙǫ,QIHULRU:]$Ǭ3m %DUEHOV:]$£)<]iǊƅɆɽɳǵɅȶɫ ɌɅƑƙȷɊɭɹ
1DVDOEDEHODWWDFKHVWRQRVWULO:]$Ǭ3mǕ(c£ǔ=ǊƅɆɽɳǵȸơȶɫ ǃƀɊƴȶɳɍˊ
0D[LOODU\EDUEHODWWDFKHVWRXSSHUMDZ:]$Ǭ3mǕ(<]% ɴȼɍɳǷȷɳǆƚɹɌɅƑƙȷɊɭɹɅɩȶǃƀɊɴɇƒȲƴȶɳɍˊ
5RVWUDOEDEHOORFDWHGEHWZHHQQRVWULODQGXSSHUMDZ:]$)ǘâǓ ǊƅɆɽɳǵǃƀɊƴȶɳƙƳɊ
0DQGLEXODUEDEHODWWDFKHVWRORZHUMDZ

3£:]=£3Z )LQVƙɈȩɋƙɃȪȶ3HFWRUDOILQƙɈȩɋɳǉɹ
1FMWJDGJOƙɈȩɋȳƒȶ%PSTBMGJOƙɈȩɋȲɅƐɭɋ$BVEBMGJOƙɈȩɋƙɃȪȶ
1FDUPSBM ILQƙɈȩɋȴɮɃ
"OBMGJOƙɈȩɋɳǉɹ
1FMWJDGJOƙɈȩɋȳƒȶɃɊɯɪ ɋ'JSTUEPSTBMGJOƙɈȩɋƴƚȻɽ"EJQPTFGJOƙɈȩɋȳƒȶɃɈɪ Ɍɪ
4FDPOEEPSTBMGJOƙɈȩɋȲɅƐɭɋ$BVEBMGJOƙɈȩɋȴɮɃ
"OBMGJO4£57](£:]=  )LQ UD\VƙɃɅɭ ȶɌȶɫ
TQJOFTƙɃɅɭ ȶɃɅɽ
TPGUSBZTƙɃɅɭ ȶɃɅɽ
TPGUSBZTƙɃɅɭ ȶɌȶɫ
TQJOFT)LQUD\V)LVKILQVDUHXVXDOO\VXSSRUWHGE\ILQUD\V)LQUD\VDUHFRPSRVHGRI W\SHVKDUGUD\V VSLQHV DQGVRIWUD\V+DUGUD\VDUH
²LIH[LVW² DOZD\VORFDWHGEHIRUHVRIWUD\VZKLFKDUHIOH[LEOH
£57](£:]=57md8$
ƙɃɅɭȶƙɈȩɋɃɅɽƺƙɆɳɉɃƙɈȩɋɴȼɍɴɆȲƺɈɌɪ ɴɇƒȲɞɳƙȷˊɅʆƙɁȪɎǍɆɽƙɃɅɭȶƙɈȩɋɃɅɽɳǷȴɍɽƙɈȩɋ ᶬ ʆ ɳǷɌɮɆɂɁƴȶɳȸƛȶǋɅƙɃɅɭȶƙɈȩɋɃɅɽɴɆȲƙǇɸɆʆ
ɪ








£57](£:]=
ƙɈȩɋƙɁɪ ƺɄɊƗǂƙɃɳƽɋƙɃɅɭȶƙɈȩɋʆ ƙɃɅɭȶƙɈȩɋǋɅɈɪɌƙɆɳɉɃȴɬƙɃɅɭȶƙɈȩɋɌȶɅɫɩȶƙɃɅɭȶƙɈȩɋɃɅɽʆƙɃɅɭȶƙɈȩɋɌȶɳɆɫˊɑɩɅƺǋɅƙɁȪɎɳǷƴȶɊɭȳƙɃɅɭȶƙɈȩɋɃɅɽɴȼɍǕȷɆɁɽɴɆɅǇɅʆ%UDQFKHGUD\%UDQFKHGUD\LVDNLQGRIVRIWUD\VWKDWGLYHUJHVLQWRWZRRUPRUHSDUWV%DVHRIUD\V ᶬ VKRXOG
EHXVHGZKHQFRXQWHG(LJKWEUDQFKHGUD\V
DUHFRXQWHGLQWKHSKRWRRQWKHOHIW

£57](£:]=57m<7
Y d8$
ƺƙɆɳɉɃɊɯɋɵɅƙɃɅɭȶƙɈȩɋɃɅɽɄɊƗǂɴȼɍɊɅɩ ɴɆȲʆ
ƙɃɅɭȶƙɈȩɋɃɅɽɊɅɩ ɴɆȲȷɭȶɳǷɳƙƳɋɳȴɄɊƗǂɴɎȶƺȶɳȴɆɸɇɭɁʆȹɯɅƳɍǏɌȶɅȶɫɩ ɄɸʆȲƒɭȶȲɌɀɪɴɆɆɳɅɹǏƙɁȪɎǇɅɳȴƸɁɽɃɭȲǃƺ§ƙɃɅɭȶƙɈȩɋɌȶʆɫ
8QEUDQFKHGUD\ VLPSOHUD\
2QHNLQGRIVRIWUD\VWKDWLVVLPSOH QRWEUDQFKHG /DVWXQEUDQFKHGUD\LVXVXDOO\ORQJHVW,WLVVRPHWLPHVVWLIIHQHGDQGHQODUJHG
,QVXFKFDVHLWLVUHJDUGHGDVµDVSLQH¶

5Ǯ(£:]=$7=© £3Z &DXGDOILQVKDSHƳɁɽɳɑƗˊ

7UXQFDWHɇɁ(PDUJLQDWH

Ɋɮɍ

ƙɑȫȷ

5RXQGHG

3RLQWHGɴȸȲɴȸȲɳƙǫ

)RUNHG/XQDWHc£ǖ(*¨([ $7=© £3Z &DXGDOVNHOHWRQȸơɫȶɑɸ ɴɆɻɁǋɅǍȶɆɅƐɹƙɃƙɈȩɋȲɅƐɭɋ ȸơɫȶIZQVSBM ʆ ɳɋȶˊ ɊɅɩ ǕȷɳɊɍ
ˊ ɳȵȻ
ˊ Ɉƴ
ɪ ȶ
ɳƙǤǇɅɳɃɆɻɭɴɅƎɳɍȷɳȷȻȲɴɅƚȶȴǆƚȲɽȷɭȶȸơɫȶɑɸ ɴɆɻɁɈɪɴɑƓȲɳɈɍɴȼɍƙɈȩɋɆɁɽʆ+\SXUDO ERQHVDUHSODWHOLNHERQHVVXSSRUWLQJFDXGDOILQ,WLVQRWYLVLEOH
H[WHUQDOO\EXWLWVHQGLQJSRLQWLV
DSSUR[LPDWHGIURPWKHSRVLWLRQRIVNLQIROGDSSHDUVZKHQWKHILQLV
EHQW
ȸơɫȶɑɸ ɴɆɻɁ ȸơɫȶK\SXUDO



6

ǔ>Ǯ8m£Cǔ£3Z 6FDOHVWREHFRXQWHGƙɑƳƳɁɽɃɃȶɫ ȳƚɯɅƴȶɳɍˊɴȳƞȸƒɮɁȷɸɳɒȢȶȳƚɯɅ
7UDQVYHUVHVFDOHVDERYHODWHUDOOLQHƙɑƳɊɭȳƙɈȩɋȳƒȶ
3UHGRUVDO VFDOHVƙɑƳǂɊɴȳƞȸƒɮɁȷɸɳɒȢȶ/DWHUDOOLQHVFDOHVȹɯɌƙɑƳǂɊɆɳǁƉ ɋȳƚɯɅ
/RQJLWXGLQDOVFDOHURZƙɑƳȹɭɸɎȻȴɍɽȲɅƐɭɋɩ
&LUFXPSHGXQFODU VFDOHVƙɑƳƙɃɅɭȶɳǉɹƙɑȫȷ
6FXWHVd%Ǔ¦*ª3)hcDb( ȹɯɌɵɅɌɅƑɎȦɩ ƈ ɀɳǷȷɸɳɒȢȶȳƚɯɅƙɁɪ/DWHUDOOLQH // VHULHVRIVHQVRU\SRUHVRQVLGHRIERG\£Cǔd%Ǔ¦*ª3)hcDb(ƙɑƳǋɅɌɅƑǂɊɆɳǁƎ ɋɴȳƞȸƒɮɁȷɸɳɒȢȶ/DWHUDOOLQHVFDOHV //V QFSGPSBUFETDBMFTSVOOJOHBMPOHMBUFSBMMJOF+_>£Cǔǣ<8cǢǾǓ=%¤«7 ȹɯɌƙɑƳɳǷǂɊɆɳǁƎ ɋȲǁƎ ɍȷɸɳɒȢȶȳƚɯɅǍɆɽɳǷɳɈɍɴȳƞȸƒɮɁȷɸɳɒȢȶɊɅɳɈȻɞɔɁɽǋɅɩ
/RQJLWXGLQDOVFDOHURZVFDOHURZDORQJPLGODWHUDOOLQH
FRXQWHGZKHQ//LQFRPSOHWHRUDEVHQW£Cǔ<]%£:]=%(ȹɯɌƙɑƳǂɊɆɳǁƎ ɋȲǁƎ ɍȳƒȶɳǷƴȶɊɭȳƙɈȩɋȳƒȶ
3UHGRUVDO VFDOHVVFDOHURZDORQJPLGGRUVDOOLQHEHIRUHGRUVDOILQ£Cǔ+]h@Â-&?m$7©= ȹɯɌƙɑƳȹɭɸɎȻȴɍɽȲɅƐɭɋɩ
&LUFXPSHGXQFODU VFDOHVVFDOHURZVDURXQGFDXGDOSHGXQFOH£Cǔ£57](cǪi£C_)ȹɯɌƙɑƳɌȶɅƎ ɋȲǁƎ ɍɳǉɹ
ɫɩȶƙɑȫȷȼɮȷɆǆƚǂɊɆɳǁ
6FXWH KDUGHQHGDQGWKRUQHG VFDOH URZDORQJPLGYHQWUDOOLQH

7

C3
Y £C$Z *LOO UDNHUVɑƒɩɁƙɑȲɪ
JLOOUDNHUVɄƒɮƙɑȲɪ ɴɇƒȲɳɍˊXSSHUOLPERI
JLOODUFKɄƒɮƙɑȲƙƒ Ȳɪ ɴɇƒɇȲɇƒƒƴȶɳƙƳɊ
ORZHUOLPE
RIJLOODUFKɑƒɩɁƙɑȲɪɳɌȢɆǂɊɆɳǁƎ ɋɴȴɊƴȶȲƒɭȶɵɅɄƒɮƙɑȲɪǌɻȶɳɑƗˊʆ ɑƒɩɁƙɑȲɪǋɅɃƙɊȶɽɳɇƞȶƵƒǄɸȶǍȶɳǾ ɅɩȶȷɸɅɅɯ ɳǵǂɊƙɆɳɉɃƙɁɪʆ ƙɆɳɉɃƙɁɪȳɹƚ ǋɅɑƒɩɁƙɑȲɪɴɎȶ ɅɩȶȻɫȲʆ ƙɆɳɉɃɳɇƞȶɳɃȢɁ ǋɅɑƒɩɁƙɑȲȼɪ ɸɳɌȢɆƵƒǌɻȶɍơ Ʌȶɩ ȼɮȷȼɭɸɈȲ ɞȲʁƵƗɅʆɍȲſɀɺɵɅɑƒɩɁƙɑȲɪ ȴɬɊɅɯ ʉɳɃ ȼɮɳȷƒɹɳɒˊɋɳɃˊɆɳȴɩ ɴƙɆƙɆȫɍɳǷȲƒɭȶƙɆɳɉɃƙɁɪɅɪɊɋ
ɴɁȶɴɁɳƙɆˊǏɑƙǋɆɽȲɸɀɁɽƙɆɳɉɃƙɁɪʆ ɄƒɮƙɑȲɪɊɋɯ ǋɅɈɪɌɴɇƒȲ ȴɬɴɇƒȲɳɍˊǋɅɑƒɩɁƙɑȲɪɁɩȷ ɅɩȶɴɇƒȲƴȶɳƙƳɊǋɅɑƒɩɁƙɑȲɪɳƙȷˊɅʆ ɳɋˊȶɴɁȶɴɁǍɆɽȷɸɅɅɯ ɑƒɩɁƙɑȲɪǄɸȶɈɌɪ ɴɇƒȲ ɳɆɊ
Ʌ
ǋɅɔƛ
Ɇ
Ȧ
ƅ
Ȳɽ
ʆ
ˊ ɩ
ɪ
*LOO UDNHUV DUH XVXDOO\ DUUDQJHG DORQJ LQQHU HGJH RI JLOO DUFKHV UHJXODUO\*LOO UDNHUV YDU\ ERWK LQ VKDSH DQG QXPEHU DFFRUGLQJ WR VSHFLHV ,Q VRPH
VSHFLHV WKH\ DUH YHU\ ORQJ DQG FORVHVHW ,Q RWKHU VSHFLHV WKH\ DUH ZHOOVSDFHG DQG NQREOLNH RU DEVHQW 6WDWXV RI JLOO UDNHUV LV VWDEOH LQ D
VSHFLHV WKHUHIRUH LW LV RIWHQ XVHG IRU ILVK VSHFLHV LGHQWLILFDWLRQ $ JLOODUFK LV FRPSRVHG RI  SDUWV XSSHU OLPE ZLWK OHVV JLOO UDNHUV DQG ORZHU
OLPE ZLWK PRUH JLOO UDNHUV :H XVXDOO\ FRXQW QXPEHU RI JLOO UDNHUV RQ ERWKOLPE XQOHVV VSHFLILHG

8d9$5Z  +h7=
_ $(
© ǔ>dC¥(>$£3Z 6(&7,21)LVK6HDUFK$LG
2UGHU0</,2%$7,)250(6S
Order
MYLIOBATIFORMES-------- p. 19)DPLO\'$6<$7,','$(S
Family DASYATIDIDAE-------- p. 19
2UGHU267(2*/266,)250(6S 
Order OSTEOGLOSSIFORMES----p. 21)DPLO\267(2*/266,'$(S
Family OSTEOGLOSSIDAE--- p. 21)DPLO\127237(5,'$(S 
Family NOTOPTERIDAE------ p. 22
2UGHU(/23,)250(6S
Order ELOPIFORMES---------------- p. 23)DPLO\(/23,'$(S
Family ELOPIDAE-------------- p. 23)DPLO\0(*$/23,'$(S
Family MEGALOPIDAE------- p. 24
2UGHU$1*8,//,)250(6S
Order ANGUILLIFORMES----------- p. 24)DPLO\085$(1(62&,'$(S
Family MURAENESOCIDAE---- p. 24)DPLO\23+,&+7+,'$(S
Family OPHICHTHIDAE------- p. 25
2UGHU&/83(,)250(6S 
Order CLUPEIFORMES-------------- p. 26)DPLO\681'$6$/$1*,'$(S
Family SUNDASALANGIDAE--- p. 26)DPLO\(1*5$8/,'$(S
Family ENGRAULIDAE-------- p. 27
S---------------------------------------- p. 279)DPLO\&/83(,'$(S
Family CLUPEIDAE------------ p. 29
S---------------------------------------- p. 30
2UGHU*2125<1&+,)250(6S )DPLO\&+$1,'$(S
Order
GONORYNCHIFORMES---- p.32
Family CHANIDAE-------------- p. 32
2UGHU&<35,1,)250(6S
Order CYPRINIFORMES------------ p. 32)DPLO\&<35,1,'$(S
Family CYPRINIDAE----------- p. 33
S---------------------------------------- p. 34
S---------------------------------------- p. 37
S---------------------------------------- p. 37
S---------------------------------------- p. 37
S
---------------------------------------- p. 38
S---------------------------------------- p. 39
S---------------------------------------- p. 40
S
---------------------------------------- p. 41
S
---------------------------------------- p. 42
S---------------------------------------- p. 44
S ---------------------------------------- p. 4510)DPLO\&<35,1,'$(S (Continued)
Family CYPRINIDAE----------- p. 46
SS
---------------------------------------- p. 47
S---------------------------------------- p. 48
S---------------------------------------- p. 48
S---------------------------------------- p. 49
S---------------------------------------- p. 50
S---------------------------------------- p. 51
S---------------------------------------- p. 52
S---------------------------------------- p. 54
SS---------------------------------------- p. 55
S---------------------------------------- p. 56
S
---------------------------------------- p. 57
S---------------------------------------- p. 57
S---------------------------------------- p. 58
S---------------------------------------- p. 59
S---------------------------------------- p. 60
S---------------------------------------- p. 62S---------------------------------------- p. 63---------------------------------------- p. 63---------------------------------------- p. 6411)DPLO\&<35,1,'$(S (Continued)
Family CYPRINIDAE----------- p. 64
SS---------------------------------------- p. 65
S---------------------------------------- p. 66
S---------------------------------------- p. 67
S---------------------------------------- p. 68
S---------------------------------------- p. 69
S---------------------------------------- p. 70
S---------------------------------------- p. 72
S---------------------------------------- p. 72
S---------------------------------------- p. 73
S)DPLO\*<5,12&+(,/,'$(S---------------------------------------- p. 73)DPLO\*$67520<=217,'$(S---------------------------------------- p. 75)DPLO\%$/,725,'$(S
Family GYRINOCHEILIDAE----- p. 76)DPLO\&2%,7,'$(S
Family GASTROMYZONTIDAE----p. 77
S
Family BALITORIDAE--------- p. 77
S)DPLO\%27,,'$(S
Family COBITIDAE------------- p. 80S---------------------------------------- p. 83---------------------------------------- p. 84
Family BOTIIDAE--------------- p. 85---------------------------------------- p. 8712)DPLO\ 6(53(17,&2%,7,'$(S
FamilySERPENTICOBITIDAE--- p. 88)DPLO\1(0$&+(,/,'$(S
Family
NEMACHEILIDAE----- p. 8833---------------------------------------p. 89---------------------------------------- p. 90
2UGHU&+$5$&,)250(6S)DPLO\6(55$6$/0,'$(S
Order CHARACIFORMES---------- p. 93
Family SERRASALMIDAE-----p. 93
2UGHU6,/85,)250(6S
Order SILURIFORMES--------------p. 94)DPLO\%$*5,'$(S
FamilyBAGRIDAE-------------- p. 94
SS
---------------------------------------p. 96---------------------------------------- p. 98)DPLO\6,/85,'$(S
Family SILURIDAE------------- p. 99
S S
---------------------------------------p. 100
S
---------------------------------------- p. 103
S
---------------------------------------- p. 103---------------------------------------p. 104)DPLO\$,/,,'$(S)DPLO\3$1*$6,,'$(S
FamilyAILIIDAE----------------- p. 105
S
Family PANGASIIDAE--------- p. 106)DPLO\$0%/<&,3,7,'$(S
---------------------------------------- p. 109
Family AMBLYCIPITIDAE-----p. 111

13)DPLO\$.<6,'$(S
S
Family AKYSIDAE---------------p. 112-----------------------------------------p. 113)DPLO\6,625,'$(S
S
Family SISORIDAE--------------p. 114
)DPLO\3/2726,'$(S
-----------------------------------------p. 114)DPLO\&/$5,,'$(S
Family PLOTOSIDAE-----------p. 116)DPLO\$5,,'$(S
Family CLARIIDAE--------------p. 117
2UGHU260(5,)250(63
Family ARIIDAE------------------p. 118)DPLO\6$/$1*,'$(S 
Order OSMERIFORMES--------------p. 119
2UGHU%$75$&+2,',)250(6S
Family SALANGIDAE-----------p. 119)DPLO\%$75$&+2,','$(S
Order BATRACHOIDIFORMES---- p. 120
2UGHU08*,/,)250(63
Family BATRACHOIDIDAE----- p. 120)DPLO\08*,/,'$(S
Order MUGILIFORMES-------------- p. 121
2UGHU$7+(5,1,)250(6Family MUGILIDAE------------- p. 121)DPLO\3+$//267(7+,'$(S
Order ATHERINIFORMES----------- p. 123
2UGHU%(/21,)250(6²S
Family PHALLOSTETHIDAE--- p. 123)DPLO\$'5,$1,&+7+<,'$(S
Order BELONIFORMES------------- p. 125)DPLO\%(/21,'$(S
Family ADRIANICHTHYIDAE--- p. 125)DPLO\+(0,5$03+,'$(S
Family BELONIDAE------------ p. 126)DPLO\=(1$5&+237(5,'$(S
Family HEMIRAMPHIDAE---- p. 127
Family ZENARCHOPTERIDAE----p. 127

14

2UGHU&<35,12'217,)250(6S

Order
CYPRINODONTIFORMES----- p. 128)DPLO\$3/2&+(,/,'$(S
Family APLOCHEILIDAE----- p. 128)DPLO\32(&,/,'$(S
Family POECILIDAE----------- p. 129
2UGHU*$67(5267(,)250(6S
Order GASTEROSTEIFORMES--- p. 129)DPLO\,1'26720,'$(S
Family INDOSTOMIDAE------ p. 129
Family SYNGNATHIDAE------ p. 130)DPLO\6<1*1$7+,'$(S
2UGHU6<1%5$1&+,)250(6S
Order SYNBRANCHIFORMES----- p. 131)DPLO\6<1%5$1&+,'$(S
Family SYNBRANCHIDAE--- p. 131)DPLO\&+$8'+85,,'$(S
Family CHAUDHURIIDAE---- p. 132)DPLO\0$67$&(0%(/,'$(S
Family MASTACEMBELIDAE--- p. 133
Order
SCORPAENIFORMES------- p. 135
2UGHU6&253$(1,)250(6S
Family PLATYCEPHALIDAE---- p. 135)DPLO\3/$7<&(3+$/,'$(S
2UGHU3(5&,)250(6S
Order PERCIFORMES--------------- p. 135)DPLO\$0%$66,'$(S
Family AMBASSIDAE--------- p. 135)DPLO\/$7,'$(S
Family LATIDAE----------------- p. 138)DPLO\6(55$1,'$( S
Family SERRANIDAE---------- p. 139)DPLO\/(,2*1$7+,'$(S
Family LEIOGNATHIDAE----- p. 140)DPLO\&$5$1*,'$(S
Family CARANGIDAE--------- p. 141)DPLO\6&20%5,'$(S
Family SCOMBRIDAE--------- p. 14215)DPLO\6,//$*,1,'$(S
Family SILLAGINIDAE-------- p. 143)DPLO\'$71,2,','$(S
Family DATNIOIDIDAE-------- p. 144)DPLO\/87-$1,'$(S
Family LUTJANIDAE----------- p. 145)DPLO\*(55(,'$(S
Family GERREIDAE------------ p. 146)DPLO\+$(08/,'$(S
Family HAEMULIDAE---------- p. 147)DPLO\6&$723+$*,'$(S
Family SCATOPHAGIDAE----- p. 148)DPLO\'5(3$1(,'$(S
Family DREPANEIDAE-------- p. 148)DPLO\32/<1(0,'$(S
Family POLYNEMIDAE-------- p. 149)DPLO\6&,$(1,'$(S
Family SCIAENIDAE----------- p. 150)DPLO\35,672/(3,','$(S
Family PRISTOLEPIDIDAE-----p. 153)DPLO\1$1','$(S
Family NANDIDAE-------------- p. 154)DPLO\0212'$&7</,'$(S
Family MONODACTYLIDAE----p. 155)DPLO\7(5$3217,'$(S
Family TERAPONTIDAE------ p. 155)DPLO\&,&+/,'$(S
Family CICHLIDAE------------- p. 156)DPLO\72;27,'$(S
Family TOXOTIDAE------------ p. 157)DPLO\&$//,21<0,'$(S
Family CALLIONYMIDAE----- p. 158)DPLO\(/(275,','$(S
Family ELEOTRIDIDAE------- p. 159

16)DPLO\*2%,,'$(S
Family GOBIIDAE--------------- p. 161
S
---------------------------------------- p. 162
S
---------------------------------------- p. 162
S
---------------------------------------- p. 163
S
---------------------------------------- p. 164
S
---------------------------------------- p. 165
S
---------------------------------------- p. 166
S
---------------------------------------- p. 166
S
---------------------------------------- p. 167
S
---------------------------------------- p. 167
S
---------------------------------------- p. 168
S
---------------------------------------- p. 169
S
---------------------------------------- p. 170
S
---------------------------------------- p. 170)DPLO\6,*$1,'$(S
Family SIGANIDAE------------- p. 170

17)DPLO\$1$%$17,'$(S
Family ANABANTIDAE-------- p. 172)DPLO\263+521(0,'$( S
FamilyOSPHRONEMIDAE---- p. 172
S
S
---------------------------------------p. 173
S
---------------------------------------p. 174)DPLO\&+$11,'$(S
---------------------------------------p. 176
Family CHANNIDAE------------ p. 176
2UGHU3/(8521(&7,)250(6 S
Order PLEURONECTIFORMES---- p. 178)DPLO\3$5$/,&+7+<,'$(S
Family PARALICHTHYIDAE----p. 178)DPLO\62/(,'$(S
Family SOLEIDAE-------------- p. 179
S ---------------------------------------- p. 179)DPLO\&<12*/266,'$(S
S
FamilyCYNOGLOSSIDAE---- p. 181---------------------------------------- p. 182
2UGHU7(75$2'217,)250(6S
Order TETRAODONTIFORMES---- p. 182)DPLO\7(75$2'217,'$(S
Family TETRAODONTIDAE----p. 182
3
---------------------------------------p. 183
S
S
---------------------------------------p. 184---------------------------------------- p. 185


18