a -lt~1;-i.~~~~%-·J2.f
Transactions and Proceedingsof thePalaeontological Society of Japan
Hi:{ .1'1] 13 ~!if; 11-12 ~ll

1938

Nos. 11-12[:!tI!'fitJ}'!~i:~'t,~ 45 ~~; 536 -539~!It:tJ.tAilU)[Reprinted from Jour. Geol. Soc. Japan, Vol. 45. No. 536-539.)

Palaeontological Society of Japan

:k
fi

{5-

CONTENTS
Transactions
lit

x

35

Takeo SEKI, Preliminary Report on the Fauna of Fusulina L imestone from lVIt.

63.

64.

Ibuk i a nti its Adjacent Areas (Resume) (Published Ma y 20) · ·· · . . . .. . " . . . .. . " . .

39l3 *lm1t;SJ~f::.Jtt ~'-C~

40

Tokio SHI KAMA. On Some Japanese Fossil Equids (Resume) (Pub lished Ma y 20) ··· ··· ··46(5 ~ 20 13 ~*)""" .. .. .. ...... .... .. .. · · .... · · · }lU 1m

Wolf

Kiyosi ASARO, On the J ap anese Species of Elphidiu,m a nd Its Alli ed Genera(Published J).lly 20) .,.,., . , ., . . .. . . ...... .. .. .. .. .. .. , ....... .. . .. .......... . ......
El *EIi E lp li:irl-ium Ml\ ~:fL £li ni'i:i~ ) ( 7 }l 20 El !lHO . . .. , . . . . .. . . . . . ... . . . ~

65

!Ill'

47

m 57

Kiyosi ASARO, On tbe Japanese Species of Nonion a nd Its Allied Gen er a (P ublish ed Ju ly 20 ) .. .... .. .. ...... .. .. .. .. .. .. .... .. . ....... .. .. ......... . .. .. ........ ..

58

El *EIi Nonion Ml\1'H Lfi (fi'i:i~) ( 7 }l 20 El ~~ ) . .. ..... . .... , . . ..... . . .. . . ~

G5

illj·

f,lf

66 . ' Kiyosi ASANO, On the J a p a nese Sp ecies of B olivina a nd Its Allied Gene ra (Published July 20) . . : .. . . . . . , . .... . . , . . ... . ... , . . . . . , . . .. . . . . . . . . , . . .. ,. ... .... .. . ..
El*EIi Bolivinn 1j ~:fL £li Gi'i:i~) ( 7 }l

67 .

~O

El ~~ ) .. . ... . . . . . . . . ...... . . . . ' ~lll-J-

66.m 75

K iyosi ASANO, On the J a panese Species of Uv ige1'ina, and Its A ll ied Gener a(Published July 20) " " ... . ...... " " . . .. . . . . . . . , . . ..... . . . .. . . . . . .

75

El*EIi UV'iger'ina ~ ~:fL £li Cfi'i3~) (7 }l 20 El ~~ )" " .. ... .. ..... . .. .

Seido ENDO, On Fossil Pla uts from t.he Euvirons of Sendai (I) (Resume) (Published JUly20) .. .. .. ........ .... .... .. ·· .... .. ...... ............ .. .. .. .. · .. · .......... · · · .. · ......
69.

Tuneteru OINOMIKADO, N eogene Shells f l'om the Vicilllty of t be City of Ta kasaki, Gumma -ken , J apan (Published A ugust 20) " "

70.

SG

.... .... .. . .. " .... .... ... ..

87

Tunetel'u OINOMIKADO and Ki nji KA:-< EII ARA, A Ne w Sp ecies of C(llyp t ogena fromthe H igasiyama Oil Fie ld, N iigata-ken, Jap an (Publis hed Aus ust 20) · ··· · · ·· · · ,·~3$fi!Sl~* ill ifb IIl Eli Calyptogena

94c7.:> l $f:flll (Wd~ ) (8 }l 20 13 ~~ )"" *:I!K.{3'1W'1~lf · 1itffJf~-=

('l'ransactiollS of the Palaeontological Society of' Japan)

1;rt1!7, U-L2iz. v: {- O) r~r·H!I:v.t:l:-tI!S:ljJ'l'IHj!U:.9'I!-[!;j~ib 1.> :f:i!!.:f:i,Jt c L. "Cr~1 <v.t/H~ IJff.±, "IJ~"l'~:&: , lli: <v.t'r'r JlI ~±vc J: Ij j[1lltJ: 1.>ltff'1Li6~* c; fL -c if;) ~ 0 3uH* , IJ,**f>IljlJff.± ~ iWl¥i EI *:l:-l!!.ij!j\,:f,ji~_U:E @IT'" ~:l:i!!.
:/;g(tJ. ~ 4J:f. tr: fm12k c; tL-c l::> ~ 0~~v.t'~-C*;I~1ii *:f:i!!.~15 ~4o/J~~~3f":*fm1)cJm @I c L. -c?JF-:J:&OIJ, ffij!jj7-j:j'/I1J tr: iill~ ~;j»jl tr: filth l¥i~~ c L.;I~v.t:J:if.~)IIm-c:f:i,Jtvc}[ ~:l:-l!!.:f:i,JtO)ti!!.1!ttr: *~\x~:j[jJ1lliWO)T vc jiffYL L.k:o*:l:-l!!.:f:i,JtO):l:-l!!.1~H'fI¥~v.t;ji]1~ -cWl!r~~ L. -c if;) 1.> iJ,~Ji)fft15ij(;fJO) lens tr: JJHiI L., {- O)15trz;fJ"p O)1-t ;fi
1'illiffiR:5EO)J:.VC :x..j;'fI¥i1!tO)jqfH$O) J:.I'C lllW'tJ: 1.>~5:tjl;lj tr:11tr-t' ~ ~ O)-c'ib ~ 0
bEi±I T 1.> 1-t15 v.t~tL iliit,!Im ·!1IOl'~J!Im · ff.\tiliiI.!Im " Jl91i;'~!IJ~ ' )J}lJE!JiJi~$ IIfk:VC}[ ~ ?b lli ~ ~ '8!."VCbE i±I L., ..E\.0
lli Ib 1ll%tl'tJ: ~ v.t0'Jfm11fJJ1WiliiI.gJi-c'ib ~ 0 ~~1-t15O)tF*ln tJ: 1.>1~FYEv.t*tt {- 0)~<1:J vc;(:E ~ ~Jlt~-C·v.t {0)=i::'tJ: ~ ~ 0") tr:7U ~ L., ftt!,:l:-l!!.tJt c O)tilt vc1§t.3l:-C J: 5 c,IIU,o*:l:-l!!.tHt15bE:f:i!!.2kV:·=i::tJ:1.>1t15v.t:j(~2.0)~Jl L. (*~\J: Ij)
1. 11Jtlj!)\tHrr-~;g~ 'lfn'.~U'l"15 I.lI
Ve?·b eek-i.nct ve?'bcebi GEINITZ
Neoschwage?'ina crat'icul'i jem (SCIIWAGER)

G?'omosp-im sp.
Tetmlax-is 5p.
Textula?'ia 9p

N. n'ipponica OZAWA
Schwagel'ina cfr. ?'ichthojeni (SCHWAGER)
S.5p.

Cl'irnacamm'ina sp.
lofizzia -vclebitana SCIIUBERT

Schube?'tella gimudi (DEPRA'r)

Gastropoda

2.

IIJtJjl..**:fJf:~;g~/J \w,~;-.J"~~

Schwage?'ina japonica (GihmEL)

Neoschwage?'ina efr. cmt'icul'i jem SCHWAGER

3. IIfk:-ifl**:fJf:~;g~.0..1Jllilt-j·~tY&~nlli:
1-t150)1~H'f-jUtfL~ffll\<Wc5Evc:tlili.r-...~" ~ ~ O)iJ'~ \.A

J

Schwage·rina. efr. ?'-ichthojeni (SCIl\VAGEU)
S. sp.

.J.Yeoschwage?·ina ? sp.
lof-izzict? 8p.

Neoschwagc?'ina cmt-iculifem (SCIIWAGER)
N. efr. nippon-ica .oZAWA
N. cfr. s·imple:c OZAWA

Schwage?"ina haJ/'t-i (SCHELLWIEN)
S. cfr. -incisa (SCIIELLWlEN)
S. sp.

Schwage?'ina japonica (GUMBEL) .

Sclmbe?·tella sp.
-

35-

426

5. Il&:lfLl\%YF:!&IIIT~~ ill

IH* IJ,lf.±O)f,¥:*IH1X. Q1t1"l5J'E ~ if:, IJ ,Jl":)W~~*tc: 13 5J'0)1*~r[j!1 ~ ~ 1i:jffi L L lJ 1X. \("I O)-C'~fllla-i"" Q 0
6. II&: lfLl\%/GliL/(;¥,li *=f *t:k;;P:1tE. Foraminife ra

Coral

Fusulinellct biconica HAYASAKA

Nagatophyllmn efr. scttoi OZAWA
Chaetetes sp.

Sta:D'ella efr. miille?'i OZAWA
Bmdyinct sp.

Prisrnatophyllurn? sp.

Textuim'ia sp.
Tet mtaxis sp,

Bryozoa

7 • 1'fr I!l<.

Gastropoda

ill=

J:, T- i'C 5J'-:t Q 4J1: iJ'Ifil ~<: Q 0(A) J:~;;P~* ,*, J: IJ ~Yctbeina n. sp.
lVeoschwctge?'ina cmticulifcm (SCHW.t\ GER)

Fu sttlmella 11. sp.
Coclonojus·iellct pamdoxica D UNBAR and SKill'NER

Neoschwagc?'ina ma?'ga?'dae DEPRNl'

Endothy?:a sp.

Neoschwage?'ina efr . colan-iae OZAWA

Tetmtetxis sp.

VC?'beebina ve?'beeki (GEl Nl'l'Z)
Pseudodoliol'ina ozawai YABE a.nd HAKZAWA
Schwage1'ina japonica (GUMBEL)
S. arn~iguct (DEPRAT)

Clirnacamm·ina sp.
Crib1'ogene?'ina, sp.
Te."Ctula?·ict sp.

S. subobsoleta var. okuboens'is (O ZAW.>I.)
S. n. sp. ex
Schubel·tella ibu7;·iensis n. sp.
S. simplex LANGE

Spi?'ill'inct sp.
Mizzict velevitana SCHUBERTlYeoschwage?'ina n:ipponica OZAWA

Schttbertella omiensis n. sp.

N. cmticul-ifem (SCHWAGER)

StaJl'elia ep.
Cl'imCLcamm:ina sp.
Tetm taxis sp.

Glornosp'i m sp.

Coral
Gastropoda

Doliol-ina oval·is DEPRA'£
SchWCtgeTina knrO't-i (SCHELLWIEN)
S. rntLit'iscptatct (SCEIELLWIEN)?
S. tschemyschewi (SCHELLWIEN)

Glomospim sp.
Enclothym sp.

S. gmnurn-a'venae (ROEMER)?

Ycttzengia sp.

Schube?·tella gimudi (DJlPJ~A'r)

Mizzia velebitanet SCHUBER'I'

S, s'irnplex LANGE

Gastropoda

Neoschwage?'ina nipponica OZAWA

Schwage?'ina efr . kozui (D EPRA'r)
S. efr. complicatet (SCUELLWIEN)

N efr. s·irnplex OZAWA

S. multiseptata (SCHELLWIEN)

N . cmt'icul'ifem SCEI\V AGER
Schwagel'ina vulga?'is val'. jLtsifo?'?n'is

T?'itici.tes sp.
Schubel·tella g'imuel·i (DEPRA'r)(SCHELLWIEN)

Bryozoa

S. e?'uca?'ia (SCHWAGER)
S. haJfti (SCHELLWIEN)

Co ra l
-

36-

427

9. 7£H'D\l*:J:&E8%tli~ !!?Jt*~ J1!
~fflli: r.tIlili'M![ffl:bt!!. fJ:.:t, 1£,?E5i- fJ:. :t, :J*~r.t/fnJIl~c' ib -0k

0Schwagel'ina (Pseuclojttsulina ) sp,

Schubel'tella sp.

Schwagel'inct 7~n~tlt'i ( SC ITELLwmN)
Ssp.

Tr'it'icites sp.

11. 7iH:{Jll*:J:& EEl~t~1:~J JJj(lllJ::klliJ'*

6*

Schwagel'inct (Paeuclojttsul·ina.) n. sp, f3

Schubel'tellct sirnplc.'C LA NGE

S. kmJlt'i ( SC RELT"WlEN)

SiaJldla sp.

S jap onica (GthmEL)?

,12. 7il't'-L!Il*:J:& E8 ~tli1:1:J}]f(llIJ ::k~,* ~Lr
Schtt ber'tel/a sp.
Bryozoa

Schwagel'ina hqt!'t·i ( SCAEL-LWIE)<)

S j ctpon-ica (GUMBEL)
S. n. sp, f3

13. w.'Dll*:J:& EEI %tli1:J~}]f(Plr~"J'*-mjj
Schwagel'ina (PsCttClofusuUna ) n . sp. f3
Doliol-ina ctlic-iac DEPHAT
Sclmbcl·tella sp.

Bigenel'ina sp.
CI'ibl'ogenel'inn sp.
Endothym sp.

StqU'cl/Ct sp.

lJfizzia ? sp.

iIli. 7il'i11l!ll*:J:& E8~f~*~~ III ~"J'7J ®'{~
Schwagel'ina hc~t!'t i (SCRELLWlEN).Schwager'ina n , sp. f3
Te:l:tttlal'ia sp.

lJfizzia? sp,

Schwagel'ina vulgar'-is (SCEIELLWlEN)

S . sp.

Schwager-ina vulgar'is val'. jusifol~n'is ( S CEIE LLWIEN)

Pamschu;agel'ina obZonga (OZ AWA )Schwagel'ina tlaZida (LEl~ )?

Schwctgel'inct vulgm'is (SCElEJ"LWlEN)
S andel'ssoni ( SCRE LT"WIEN)
S. tsche1'11.yschewi ( SCRELLWIEN)

Schubel'tella? sarnega·iens·is n. sp.
Pseu(/oschwagel'ina ·sp.

Y,J::0)1-t;fi J: IJ -t-O);;r-;-T n;Ht1!:~~ ,--,J3,,,,?~;fig-wO)iltH[: 1!:~J;,c:t, fJ:. Gr.r , Loc.·No. (6) (*-=f~"t-;k;fi) r.t-t-0)~ ,'±l-T ~ m75ililLiTill~f1J1;K :llJIJJm!JijU IJ Moscovia n J: lJ~iJ'G-P ~ ~-""G.n, *:bIl!±J;H#c)(ti itMO) :fftTf,{I 1!:;;r-;-T ~ 0) C' ib :t, 0
Lot'. No. (9) ....... Loc. No, (18) O) ft;fir.t friJ.n ~~*O) Ul'ali an 1!:tlfi;;r-;-T:t, Ib O)c' ib IJ , :;!;l;~ a.irp~-l'C J:T~mi'*1!:~fi?,c\'):t,~rr.t0S1!)'iftc' ib:t, :M, L oc.
0, (11) (::klli],*O)}IO, 7Jz-a:. Loc. No, (12) (::k~ J11 )1 IJ r.t Schwa,qC1'ina japonica (GihvIBEL) 1!: ~-T :t, 1!: y, L J;[:lli3CF.I':]J::ift 1!:7F-Tlb O)"'C'ib G :5 o · JZ Loc.
-

37 -

428

Fossil Zones
N·

Impor tant Fossils

Locality

Zone of N eoschwagel'ina

Yabe'ina u . sp., Neoschwagel'ina cmt'icu -ffri!k. ilI J:.llil

cmticul~fcm &lifem , Vel'beek-ina 'vel'beeki, Pseuclocloli o-1i\';f!}(tHlill

SchWCtgel'ina

jCtponicct

linaozawai,Schwagel'ina

japonica,

Schl1:agel'ina subobsoleta val'. okuboens·is,
F'u sulinella n . sp. Coclonofusiella pamcloxica , etc.
ill , Zone of NeoschwagerinaJ[.

Neoschwagel'ina

nippon-ica.,

D oliolinct

nipponica & S 9hwagel'ina

oval'is, Schwagel'ina 7cmfft'i, Schubertellakra,O'tisimplex, etc,

Zone of Pamschwagel'ina

Pamschwagerinlt

oblonga,

D oliolina

oblonga & Schwagel'ina vulga1'is al-ic'iae, Schwagerinct vulgari4J, S. h~tlti,I.

n. sp, f3. etc.

{;)} i!k.ilI "fllil, Jfj,IITt*'.t*ti!f7k

1l' .r:v#*i3' , JII JTt.mj[[**111, *lllJ·*Jl;L1f, 3JtJI I,
t:t*1¥i1f, mllfi'i3\ :fl:ilI'l,
1!!j~IHf{# " -v tJ: =," :k/lli ,mH~~ l:ff!iJ

I , Zone of F'ttsulinella biconica

Fusul'inella biconica, Staffella cfr. mollel'i,*'f-#*~

Bmclyina sp.

No. (18) Crnlli&-;I:I) fi ~)lb 'f'C1fE*Q) "Schwage1'ina" horizon (4- El (J) Pse2tdoschwager-ina andParaschwage1'i n ahorizon) c'.t:t ij5( ~I~T{:ll::tt- ~1' t Q) c!:.;}§-~ b,fl"~1t!!.Q)

Locality fi.f Q) 'Nfl'f'C i\,Q t Q) c!:. }~, fi,fl,,Q 0
1~jll~k ill ;:S~*TmlfiaJli» f'C/J \~ i~:±Q)

Kn Zone f'C!Ht c;,Q '>. fauna :tt-'f-f L, Tms=~~tc,~1~ ­m:iL-C Loc. No. (8) C:kYf~7.1<*ift ill ), Lo c. No. (2) ( + ~*,,;Hij~tl), Loc. No . .
(3) C ~llf&:), Loc. No. (4) (J llffD fi1iiJ,fl,t;Z. c!:.m* fjj'] J!t\§c·~,Qo#t1~zill;:S~*J:msfi*:f:i!rr~~M(t1~Ji Q) ~J:ml~~1' t Q)-r\ IJ \~tvJ± Q) Nc, Nm lJz(J..' Ng Q)-mlf'C t i It c;,fl" f1;'fftfi 9='ml="I'iUc. c!:.;}§-~ b,fl,~ 0 Loc. No . (1) C7:if;f.!U".!·;:Sl-U) fiPl(h;z' c!:. fjj'] Ji$c' ~ 7.>0
~J:.~k{;:) l' ,fl,tf*:f:il!~U)j~ iliill;:S~* f'C fi 4 {t;:Sif'j1tJ{ilft, ~ b,fl,,Q 0 *.ft;:S~(J).:E 'It ,Q .ft;:s lJz ltpfil!!. .

~1' ,Q tQ)C'~b?o

HJv:y,fl,tfJ:*(J)~Q <c'~,Q 0

1f It
1. ili'flffl!: Il l:f:il! c!:. Q) ~H It*:fil!~ c!:.:Ift t !&'~tJ.,Q ~~1*f'C;tt ,Q#J1Ii: U.l :f:il!Q)il!!.j!i)k(J.'.j~!H:TIfi:lftili: iJjj*~±f'C J: i? ~* c; ,fl,ko ffi]~± fim {IIJ ~ ~ Artinskian"-,,Saxonian C' ~ G ? c!:. c;,fl, -C b,Q 0 -t ,fl,fi~ b < Pal'aschwage1'ina
oblonga (OZAWA) :illE f'C Mizzia ve lebitana SOHUBERT Q) &i /±\ f'C ~ < ~ Q) c!:. :JiU\~ ;:;,fl,,Q 0 Para- .
schwagel··i n a oblonga fi/H*'I~t±f'C J: i? ;r,*E~ Q) S2tmatl'ina subzoneQ)Tmsf'Cr;7J t6 -c~JiI, c; ,fl,kt Q)c'~,Q iI", *:fil!JliXfJh'll &-:;J.\:f'CJE:--C fi SchwagM'ina vl~lgaris (SCHELLWIEN), Pset~doschwagel'ina .
sp. ~ c!:. 1*1±'11',Q ~ J;J, -C Jifr~I§l~* Q) "Schwa.ge l·ina" horizon ~1t*'T,Q ~ Q) c!:.;}§- ..h~ ~ -C' ~ G? 0 3Z.MHillJEi-fi J:.T ~~ t -c1~rl1jz ill Q) ftQ ~ Ne oschwage1·ina. fauna ~~JiI, l' ,Q*iI','±l*tJ. 0 0 J;J, .
J: J: i? ~1IJl lEffi~jff c!:. L -C*:f:il!~Q) M &-;I:I: ' :k}J[~ c!:. l i lt c;,fl,,Q ~ Q)c'~ b ? 0
m1IiI Ji}5t;{fj:f:il!~Q)*:Jjf'CtE,Q~.ID.1:~lJzlt* ~J.i~tgg Q);:S~*fi*il!!.:J:et0)/H~A"J~;g<D 'i; Q) c!:. tilt c; -

38-

429

.t1:o 1/;, o.yc'ib b :7 a2.

~m* I-Li:hU1 · ;f}(E';g tiF:J.&c O)tHL*:l:-\J!,:!JR 1tlEi~~~ 1 1;!IHi~m*ili:tU10) F7£sttline lla b oclci zone, ;r:Y\E'I

O) C2 Kt-HL L-L7t;-:::rFI'I, ·j&LJ.
tJ.. <0 :i!f~ 2 1;!IHi ~~* 0) S ch1Vag erina vttlga9'is zone, ;r:Y\E'* 0) C3 K t "JJ:L2' fL:O c'ib b ') 0 ~£ 3~fi~Ur.O) N eoschwagerina c1'aticulifM'a and Schw((,gel'ina ambigna zone O)T~IK, ;r:y\E'~:0)

CPg lfzv:. PI o) - ~K , tiF:J.&O) N n zone lfzv:. Nc zone 0) - 1j{IKi::fL-c' ~'l~,JJL2' fL o ~ 4 ?~fi~~*O) N eosch1Vag e1'·i na c1'at·i c'u U!e1'a an d SchwagM'ina am,b-i'guct zone 0) J:.~I, N eoschwagel'·i na

mal'gal'itcte zone lfz v:. Yabeina globosa zone O) - ~IK,--~K)

x;r:y\E'~ 0)PI 0) J:.~I) P 2 ) lfz v:. P 3 0)

tiF:I:&O) Nc O)J:.~\, NIll, llU,.' Ng O)-~\Kf::t.'i"t':o 1/;, O) c' ib b :7 0

Preliminary Report on the F auna of Fusulina Limestone
from Mt, Ibuki and its Adjacent Areas(Resume)By
T a k eo SEKI
I, There are many large and small fusulinid-beal'ing limestone lenses in the Titibu System ofthis reg ion, The fossil localities and important fo ssils are enumerated,
2, The geological ages of the limestone lenses a \' e foun d to range fro m the Moscovian to theMiddle Permian, on the evidence of the F usulinids co ntained in them .
3, Four fossil zones are distinguished,(N) Zone of Neosehwagel"ina emt'iwZ'ijem (SCHWAGER ) and Schwagel'-ina jap on·iea (GUMBEL)(ID) Zone of JVeose!i.wctgerina nippon'ica (OZAWA) and Schwagerina ha,ffti (SCRELLWrEN)(][) Zone of Pamsehwagerina obZonga (OZ AWA ) and Schwagerina vuZgm'is (SCHELLWIEN)( I ) Zone of F-usnZ'inelZa biconiea (HAYASAKA)
4, These fossil zones are correlated with th ose of Nagata, Almsaka, the Kwanto and Snzukal\1onnta-inlands,

-

39-

63.

11'[

,~ Ii14l'~I~ ,!: ~:t: KX.1HI*~ K 1t;s" ,!: L.. -e f?mtgH .. m!lm t f? <~ilL r m~il\Yl ~ fL -e!iS- 0iJ{, :1:2!: K B *~,~5c'li 1ili cD 1t;s"~!lJ.j K .t[: L..11li3~ c'ib 0 ,!: ~.b*]illM lJ. 0 1 f9JJc'ib l? ? 0 1t;s"~K ~ tt11 .~ fLtdF1~.
iJ{L ~cD3:: '!: L.. -e l¥i}Jfq~cD JlDJIjI?,J!.ilil\ ( XlittJlDJ) K/G~c' ib--::J kc'ib l? ? ~ <\') '!: }[I,lifL0 ,1 )1Jf: L..k

C U\I®fJ'i- c' t L iJ{Eii i±\ T 7.> ±&)1fIi*I~ '!: cDj\\'H[:-? ~ tIJ::*~ '£.m:~ lJ.1t'* t 1fT 0 4ij:i=Ii;G;!{L -e lJ. l? lJ.
\/"I 0 ~r~;([ Ii *~ft**1Uf€!f~r5.4:IjI£IJ~~X~ cD l~fl$LJlDJiIo/Jf~*~f{fll1.K ~ L.., 1t;s":& r):"1t;s" l? L.. ~ 21m1cD ,~1tA'it~* t 9;rl Dq~kcDc', :i:"f.K;M'; cDfMJ.!T1lJ. 0 ~[l.~l£t L..jfzV:' K*T cD~~t L- -efii t£" )i:\/"I o ili'fr~lJ. iJ{ G~*~ttl®fjl- s':J lJ. 0~*if.JUI'~·f:U:~;!il:1iliK~m L.. 1®f ~ I:H ~<:q~ lJ. \/"I cDli B 'U t q~ lJ. \/"I 4iJ:j:c 'ib 0 "* liWi t l~i T 0 K t ; IJ i~~!:U~lJ. 0ttlftJcD~~t Jl&\--::Jtd,:trll:$\:;.4:KiW- <~rJJt~ L.. ~ T o

1. t'\EEI ~%l¥if* lII*r\~ )I I *l·.j:l;\!*Eii (:b1~ 1 !; a :&V:':b1~ 2 fill] a )~* Ii J::1Ci)i]t£r cD lfii~=m:tr!I ('C Qi: \/"Itr!l5l' cD lk'.kJi-c', Jl- lll it.zp-JJIl.~:l: cD 1*~ d~c' ib 0 (Reg. No. 36729)0;fl;\!*Wtili 1II±~71fJEi-cbEii , 1tl!.3'U:: D 5 .RT K ib 0 ±~71f 9:1 K -emtJ¥. ~ fLkS:l c 'ib D, ~ 5l'*cD1t;s"~~UJ~ Kli T 0 t cDc'ib l? ? .2)

Palaeoloxoclon na1nad-ic~! s yab ei (lIfATSUJ\WTO)
S us n'i pp onic Lls nip ponicus MA'l.'SUMOTO
Oe?'vus (Si ka) ezoen sis HEUDE
Oanis Sp.3)

f~*tIJl:1: K J: 0 L

IlrJQi:cD')"~JiliPT~JEi- cD Ttr!lt W.f.r~T 0 Vf:\*K ffitr 1iPJit~:lt:f:JEiiJ{:fr*trlIKM-\/"I-e

2 1Jc cD ±~71fRJt '@l 7f. , .Il.tcD 9" KJW(~JHt;s" t '@l1f-t 0 S:1c' ib 00 tJ.J1* cD 1t;S"lIjll$LJlDJIIo/J~Ii ~~~ J: IJ!/mlfLf;f S t egoclon cD lji~L.. k ~ftJ: t>~WhJ~l? < 9:1tr!1-tlt:f:j~'I!tt;FTc' ib l? ? '!:}(l;I,lifL, .Il.tcD~tt
K Ii/J 'JL ~-? J::~I~.4:JWi-~K~lt 3t · i'iWljH* cD JlDJIjI£lJiJ{ l:f:lm L.. -em 0 cDc'ib 0 iJ{, 7J,.1. 0~1tK '11?J7J' ?b.Il.tcD ~trt ~ll\W!.lT 0 f.£f.1 < *~ t1tl!.75 K ~J.jj(·11cD,~ iJ~ l±\m L.. -e!iS-;:' cDlit~lJiI~ ib t> 1I.M~tJ: ~ c'ib;:' 0
rtlm!l~tr!l Ii ~m13t£ L.., \i'iJ-1tl!.Eii nctmadicus ~tir cDlrlIJljli\-nJUtl~ fij~1fL T 0 0 ~*IiWf~K-e3't"JJJ't ~ fL -e!iS- 0 iJ{, lfir%LtrllcD**liliik:i.w. L.. , ;:~J::W J: IJ W!fI.-e 0 1'& 2 Wi] a) 1illK71~:IIk t:fltt ~'HA~0
cD7f.c'ib ;:'o *~ 22 ~oe · ~~ 23 ~oe · ~~ 31 ~oec', 5t*Ii* ~ ~~~:t:Kt--::Jcib--::Jt<:l?L- < , *Utir .
%LWlilE:1.77f5 C:E: J~ J: ~ *:1.7%KQi:iJ, 0 k l? L- \/"1 0 t£r~f~cD~ ~ Ii, Fltt J.2 ~oe · ?1-mrtt · ?'t-Fltt~K
9 ;ftc'ib 0 0 J.jj(Ji p r otocone IiJffiFF-c',~ t£r'!: L -e 1iE:*c' t lJ. <IJ'7f:bc' t tJ: <, PJf~jlj caballus cD/l'lI/fijll'fx : 1t;Q ~(illi}!lU: L. L co EPjqfi'.Q; o ;fi~J?~, ~ 7 1&, ~ 6 lJj,~, 1937, pp. 215- 216.
MA'f SUMOTO, H .: On Some Fossil Mamma1s from Tsukinoki, Ugo. Sci. Rep. Tohoku Imp ..
Univ., Sen dai , Sel'. II (Geol.), Vo l. III, No. ], 19]5, pp. 39-48.
3) ~ il~ iJ.l.: ;Q:JlS:1I'f1~* I7.)fflf;f'H T""'1 >< ~})Ij ;fJ,Yi. I:PI>.;I(:, ~ 5~, ~ 4!JJJi, 1936, p. 14.
1)

2)

-

40 ·-

431

JJJ[Ji: ~/"}V 0:0 y ~~g~Jti,~,1~~T Eqwus pj,z ewalskii POLIAKOFF? O)mtJi:O)7f*K ib1:-c m ~ 0 I1J;IJ'iHIJiIl
Vi l:I:;iI&~I/~rI~lj 1flc'ib~o fiR~I~ caballus ~ E. hemion?,ts O)f~1~KI1:rnX;IJ ~Jl1b1. L--em~0 ~r'i[O)lttr11l\@'P~ p a m st yle, p amefossette, Pl'otoconule

ViiiJ.UJt L--elJj'o~tm()..j)'if€0iJ{ J l:tij&i'1'~Kn1 ~iJ{tl~j~c' ib ~ 0 E q~!t!sleptostyltts l\I[ATSUMOTOl)

~

~

~

E . samneniensis TEIL-

2~ 1 Ililla. E quus sp. 1:ir:iI%ill:in'!kIIIJ\lH.ilI.* illi

1.

:VI-nil]

2.

I"'Jnll]

Equus sp. l~ ir:~~ 3 k iO1: 1I:ir;JI-::Ij&;IIJ,j;Jl)JJII/If:tili:Erb.
3,

I"'Jnll]!lil1

a . E quus sp. f.ill.*iltiliU ii o.:> JITr7IITffib, Eq'uus cCtballus L. :;{:i 1:t!t 4 I j, r:.:1
1l1r O,) ~rt:lH'C ili: \'! '!"iI5} 0') 1jJ] ItiJrTIfJ 0,) 1:nllJ

4. 1:W: fI1l]m;N~~~Jti, @t,~ 1l¥,i O) f~~Vi*1l\'?* J:: IJ f';t 1;wJjl~O)t%~c' ib ~ 0 i~*IJtl.=I::K J:: ~ C, Y:!~TI~-lJto),~ VC Vi*1fi'l2 *ib lJ, E . pliciclens, E.s iissenbo?'?'l.ens is ~t~~~?i1Hx-eJJJfll:: O)*'lli!(j:~'<0 O) c, E. cf. st enonis
E. lepto stylus ~f~~H~J1flK -eJJjfJi:!J \'lli!(j:~ '<0O)cK%~q~ ~ cO) ~Jc'ib ~o *~*Vi*2E . JJjfJi: O) 7f*~,* o f;,j~ O) !,*f:L~~ bl~li§: K An~ c leptosty lus, sa.mneniensis ~O,)]j:t~pgHt;:r:i,~ c~11J , 7if; t;, < przewalskii c '<0 f4 IJ, ]lZ Vi h ernionus iJ,3Z.Vi Celtic P ony O):tzQ ~ IJ \'lli!O) ,~ (:ill:i:iJ
Eq~tuS K -efi)i-~fll caball'U,s oj" K A ~) c' ib ~ iJ, '<0 f nJ/'l,(j: 0 0 3\l K ffJ. ,~O)%~JlVi=BEt o1fi'l;j;jf RI'H i&o~·~·~t=E~~* c L- -em IJ, %JllIE Ltdtf,iO);r)li'I/~ll\'?* O):tm~ t .bl,-e:;JtO)*;mi3':JW:TIitfn~:tzQ ~ Vi
~UO) l:t~&ll\'?*O)lt~-tt t;, J/'l,~" ~ IlR IJ /GI1J1J~ K~lf:00)c' , iiJ:K ViJ=~ Eq'U,us sp. c L- -e1Nt~ Jft0 0~ 2 II b Vi JJ1!. -lit 0) ,~ 0) ti J:~ 4 !J\ /==I1l¥,r0) ll¥,i~:l~ K ~ 0 'M1:5j-0) ~;rjll[ t J:. J:: IJ fIJi! k. II c'WI& K.:f'-1iII Vf-et&: < (Reg. No. 59746).
COCCHI,")0*1) lVL~'l'SUMOTO, H .: On Some Fossil Mammals from Ho-nan, China. Sci. R ep. T6hoku Imp.
Univ., Send ai, Ser. II (Geo1.), Vol. III, No .. 1, 1915, pp. 29-30.
2) T EILElAIW de CElARDTN & PIVE'l'EAU, J.: Les Mammife re Fossiles de Nihowan (Chine). Annales
de Paleont., Bd, XIX, 1930, pp. 33-41.
3) BOULE, M, : Observations sur Quelques Equides Fossiles. Bull. Soc. Geol. F rance, III Ser.,
27 T om. , 1899, p, 536. ·
-

41 -

432

2.

*~Jl*#-1i;!tg~1Ij]JII [!f-Jm:~ (1Jg 1

IlmI b)~*li1CT5f~ 3 * r::lllfff'c-eWtJ2~O)n:~K:b' l..o(Reg. N o. 59739). ~;f~li~n1!~ft.,

JiiIJ'11l~t~I:l:\-lt L~ , ;i'lmll~l31ft ~ 7.k~ t5t vt-e f.S- D, -:g{IK lii1!*au'\\UO)~ftm ..t .0 IJ) t Jt.o t. 1j\GliyffiJrKfl ~ J:Jf Gft.k. ~ Il):b, ~ J Ilft.1'J. v. ~ H?i~ - !iilJlll~~ K }~ r.~-& c' 7\:~K ~ lj..
L -e 1t1:i:b,;;r;.:b'~Ii L v.iJ", #1"*'1'0p - ,,"[lfJili: Monte Sa cro Il) cp'B9\;9~:mMC~.l!!§llfr (Reg. No. 38904)1) 1';t*1~*.J;J.J:.
K~ At L \rIfHIl t £ i"" .0 Il)l'.., ~~* ~ 3Z9i-illll) lj. J: D i""f1Af"1t;SK~f-P t.1i5E l,.-l! .0 ~ ~ 0§j)'ijE-C'~.o:b'G , 1t;SGLl!;f~* t. L-e$jH!fL~v.o :f!Q*~~* Utq[3(~fl~V.o :&~~t-130 ;ffdffi.ff.J 12
Jfe . jl1ij ~ kt-l 43 ;J.-'f;-c' , * ~ •%itJt~~K !j\l;tltlfi.1!!§ Il) 1l'ii t. t~ G1'J. v. 0 )W ~i!i cctballus K [j'I] 5E ~ .0"'-";: l! ~ Il)'*'C' ib G ') 0 .
~~(;!iDDRIJ)1t;S,1!!§ t. B*~1$~~1t;S.l!lfr t.1l)~IDt*:MzU;:K*;FII~{t;S.I!!§Il)~i"" .o~~K~~Jt" .o~~);ji]HRIl){t.T:i.I!!§K;¥;tV. -e - ~ L -etl£ l! ~V.o::9cK*;F~1t.T:i.I!!§K ~ID L,

Equ~ts mlifr.\l'~iltll)m~ ~ ~'I·I K ,'±I !Jil.L, ~m·[ Il) J:.'B9\!~!(~~·lltKIi E . stenonis COCCHI (* '1~IH¥ )'E, siissenbonMnsis WUST ( ~) ~~ D, 3Z.J:.'B9\ Siwalik t-t J: !J Ii E . sivalensis FALC. and CAUT.
(Siwa lik Hills - Rup ar - L eh ri • Pabbi Hills ~) -E . n umacliclls FALC, and CAUT. (Hoshiarpur 'R angra ~) ~1l)*'@Il).l!!§ iJ" l±\B1. L -ef.S-.o .2) xJJj\c'liWt~Il)=I"j*~P ~1JEinrt. ·Reddish Clay . f,!iJ

r:r m~K E . sanm eniensis TEILHARD aDd PrVETEAU iJ{ffl,ft,.o ll)iJ{ Eqttus t. L -e ll):lfkfJ]Il)~2J"J("e~ .0,:1) ;tli sivalensis ~ namaclicus 2kU;:~ l'1H K stenonis K::9(V.c·,'±IB1.i"".o E. ?'obustus POUEL~*m!.I!!§KnL ,3U~* 1l)

E . pacificus LEIDY, E . occidentalis L [<;IDY, E. giganteus GIDLEY ~K ~ ~~%~ D, ~ G <1i~ ~~'I'1 J: D~~ ~~'I'I Ki1ti\ Lk Equtls Ii san m eniensis K m::b '--?k~\f.t~JU(f-It L~.o 0 ~*K1R-vt.o sanmeniensis .bf.iWll).1!!§ lifnIfL ~ Hippa?'ion . -C'~ !J , H iplJa?'ion R ed
Clay Kli H. richthofeni KOKEN 1l)5't- 10 fj[.bf. .l:.il{9;nG ft.-e f.S-.o: 1) 1JEinrifiJ: ~ Ii HilJpct?'ion
Il)£Jjilli~ k.o H. (P?'obo sciclippctrion) sinense (SEFVE) iJ{~1f. L -ef.S-.o 4J\f.il{9;11 /?ft. .oil", .%Kf£J
R eddish Clay A ·B·C ~Wft:iill t-e (cp'B9\f(\l'~.l.7:¥T'B9\}J~{J'lllt) ~;tkll)li E. sanmeniensis -c'~
.0 0 CP'B9\'fJt{j!f'llt K Ii samneniensis Ii~tm L, Ordos Il) Sj ara .. Osso .. Gol Sands K Itj: E . cf
prz ewalski-i POLIAK01!'F C. E. hemiomts PALLAS (~tlll) Cbi g t ai 3Z. fi Dzigetai, Tatar Il)
Roolan) iJ"JI1 i?ft.,s) ""'oj[.,~. ;/ Jmfl~I1!lrt:!.~ fU!, 1l) Sun gal'i Sa nds K ~ [j'I]f~ przewalskii .l!Ifrn1bJ,1l) .l!!§ iJ~-#- <~i"" .o .n ZDANSKY liJJKK~r5 Hallong.. Osso, Tomenl 0 2kU;:inr~ ~ ~' Hsi .. L eang .. Sbang J: b
1)

Equus adamat'icus SCElLOTfT., 18~O c :7" I~ Ki!l';6,;j1.. -C <b 0 a E. caballus L. f1;J Synonym c Yo A

2) F ALCONEH, H . and CAUTLEY, P.: Fanna Antiqua Si va lensis, pt. 9, 1849.
3) TEILHAHD de OHARDIN and PIVETEAU, J.: op. cit ,ZDANSKY, 0.: Equus and andere P eri ssodactyla. Pal. Sin ica, Ser. 0, Vol. VI, Fasc. 5, 1935,
pp.21-45.
4) SEFVE, I. : Die Hipparionen Nord.. Ohinas. Pal. Sinica, Ser . 0, Vol . IV, Fasc. 2, 1927.
5) BOULE, M. and TEILHAHD de OHAHDIN : Paleontologie. in BOULE, M., BHEUIL, H ., LICENT, E .
and TEU.HAHD de OHARDIN, Le Paleolithiqne de la Ohine. Arch. lust. Paleont. Humaine, Mem 4,
1928.

6) r.w. 7k m:Jllt . iili:.5tg~: iitilJlh'i'fW£l1!i*Hnm~IIrt11lt 1 1i'iJ~1~i~ l!fF:?i:$3to 1lt 1 *mJ~~H\ljID~I'ilEl!fF~'E ~$-';!r ,1lt 2 1l11,.1lt 1 ~i;j , 1933, pp. 71-8l.
.\'IJ[~;r111t76.~KYJ: Ordos c "'J~p?'zewals k'i-i c hemionus f1;Jjifj:f.ill:V~<boG L..v'o
-

42-

433

rm :fjl!(7),'i!H~¥R.-'rS- L --em 0 .1) *:fjI!I'i TElJJRARD de CRARDIN

K J: 0 c sctnmeniensis c Vi!jJI~;:fJ:. < ,X:1cf£*%c·fJ:. \tIil{i? *~ 1i,if!§c'ib i?, sanmen ien sis K1-c0--e OIJ1fIl~;j"j~·Ht~1HH.ki±rc·ib]?, 1879 '¥~r!i Kob do K::Jt---e:!t(7)~, ~Jil!. 1i::~.Ji!.~tL, !L1J'~.Ii1fr c L--e:t'filCfJ:.0kt(7)-eib0 o
ANTONIUS Vi P1'z cw (!lslcii ,if!§;!J{*,if!§(7) P r ototype (7) 1 c'ib 0 C L, POCOCK t X~r!i ,if!§ · jllfjff.\'(~ ,l'!§.if'i¥i~(7) ']'arpan ~;!J{ZJ: i?~;O>tLkc L--em0 0 ]l!Kr:PNt(7) Kat hiawal" Pony 'b p 1'zewaZslcii,if!§(7).yfffi c ~ tL --e m0il{ , Z K Vi E. namadictts ;~ ttl!. (7) .rul~;:~m1* t ib 0 c'ib G ? 0 %KfiJ,
J:.=BllilJ<;j"j~·iJl(7)ti!inRfC.m:! 1i::1i Lk(7)Vi p 1'zewalslcii C hemiontts c 'ib 00 ifrJi¥i~'(7)}i{±J: i?;j~* f:VJ±Vi
E . Zeptostyltts MATSUMOTO 1i::¥R. [:: GtLk o JJfi~5:tt~~*Vi1i:J:.~ 4 ' H::/Uf.i 1 11~lc'ib0--e, -If,\~ ±Vi
przewaZslcii c f!l1- 0 ,if!§ c ~ tL --em 0 0 * ~ Vi* ~ '1IIjM~K 29 ~"f; c ~ tL, ZDANSKY (7)'!i:.Z) K J: 0 C,
SALENSKY ¥R.-'rS- ( 1902 '¥) (7) p1,zewalslcii ,if!§(7)~~ 4 ,H=lllfiVi* ~ 'II%~~K 30 ~'f;ib i?, BouLE II-u: T ElLRARD ¥R.-'dr- (1928,¥) (7) hcmionus (7)5f~ 4 + E/ IiffVi'&~ 30 ~'f; 'IIJi% 28 If:ib o (7) C·, lept o.st y l'lts (7)* ~ ViJl:[:;~ C *~1i <, XJiIJ;])~n~~ t IWJl~~~T\:1@'i.lilfr llfi (7);J;!; C *~1i \tI (7)c', ~ G < lepto.styltts Vi p r zewalslcii ,if!§KMl-t 0 K~Ff.if' C t ~{~-t 00 -.lfkill: TElLRARD de CRARDIN C Y OUNG
VijjiTT¥j1~r:1<'~(7) IY.ltllffi:~:fff:rR!, 1i:: 1ifF~t~'!i:. L, ~ <(7)-\lY(ym U~K Eqn~ts caballus L. 1i:: ~ [::, ;j~*tJ,l):±(7) l eptasty l~ts U.lt±JJe.:gUffirfJ!,(7)-',11lc'ib 0 C (7)iW4j\k(7)T K caball us K AtL --e m 0 .') 1fl. L~ltx:Jffi~ii,f$ 1-C(7)~* ,if!§c L--e(7) l eptostyZ~tS 1i0.f=,mVi~f lj c'ib0 c:Zi:0--em00~ --e E. (E qtttts) cabal/u,s L. Vi Pol yphyl etic 1if]l:c'ib!J, ~:W&(7):5li'fl[K5J'ktL --e
0 il>,
LlNNAEUS (7)~c~i\; LkfiidtlJ!.(7)*.Ii1frVi H i ghla n d Po ny .if' Shire Horse J: i? WJII G ~ tL, Jlt~Y;Vi~1;JH
K~vt 0 Protot ype (J) 1 k0 E. 1'obttst,ltS POMEL J: i? ~;O>tLk t (7)c'ib 0 0 :jft-0--e}j:r.1$K~vt
o * ,if!§K1J;J,ktt;;S,if!§Kl't L --e-tRK E . cabalhts tass ilis CUVlER 1i 0ilffl}Hff-t~Vi, .Ji!.)jK J:---::J--e Vir",'Ji\!c'ib G ? L, 9 < c t *t-l:5J'!im~K c 0--e Vi71\JJI~ L~ij~Hlli;O> t g;[l tL~o caball,!ts K1);i,--e~ Ffi 0 JJ U
:fjI!c'ib 0 :iJ> t g;ntLfX caballus Kill: < c t t ossilis c Vi],[5J U~ 0"" ~ t (7);0> t g;n tL~oleptostyltts 1i::~ ~ X:JJf[ ~6'* ,if!§(7)~ilK-ttA,; c -t 0(7) t Jlt(7)~~(7)Jj!Jft!Klli"::\tIc'ib . G? 0 X p1'zewaZ .s7cii ,if!§(7).ytffi~il{~tl · i'mmHW'lf!.\'(:K ib i? c ~ tL04-13 c'Vi , iW~(J) Jj!,f~~ t i{~K f@:-t 0 c'ib G ? 0

PJm

'*

m

N£

fillr:l)glJii~ 1 1'itK 1j~",--C DARWIN l1:*r:I):!m~mlt~-?--CJi!TQo'With respect t o hO "ses, fr om reasons wh ich I cannot here g ive, I am with much doubt inclined tobeliev e, i n opposit ion to seve ra l autho rs, that a ll the races have descen ded from one wi ld stoc k.
J1:1;(7)!l~I1:?1JiMtr* .l0r:1) ilJj:~~T""('~ b ? 0 p1'zewalsMi );!~r:I)!IlJ·;j:J1:fillYlJil,l1:filll7)g'Ji(~~ 6 illHI:llJ&:l;R 7 ~F- 'C'~ <> 7l, b,

D ARWIN ~fJ1jq.ll1: f,,'Jlli c tJ. b tJ. 7l,-? k l'..~ b ? C ,\l1,I1:tt- <> 0 1ifJil;r:I) l't'iliJI g.iEl[KIM~ L---C 11:~*iIDiiifiiiJ!:Hin/ --C Ji!T<> 7l'~t*11: ANTONIUS, POCOCK

K1ft:.rNitv'o Ji'[i'PIJii~f:l'f.i!l~KIMlI L- --CI1:*r:I)jj]) ~ r:I)~JiI,7l,l3t ~ 'SL-?oEquus (Equus ) caballus fossil-is CUYlER
E. (E. ) caball1ts leplostylus MATsuMo'r oE. (E. ) caballtts p1,zewals kii POLIAKO I' F
E . (E .) p1'zewalskii POLIAKOFF

I-lippC!1'ion 11:1it*~ t *K~~ L-, 1l[~/ll l::I2krfw:rfH K;I[jA L-t::: C r:l)5EiitI:""(,~-?kn ~ t;)j~o.) Valent ine beds (Mio"Pliocene) KMlt> --C Hippcwion gmlurn LEIDY 7l,I3W\i L-, zf;t ]'{e1'chipp1ts .l: ~ ill!1-t L- t::: t fD c ~ tt- --C m- <>-iWl:r.li:;i(JtiHcl1:.lfr~m Pontian

Q

llli

K-*;TtK I-I-ippa1'ion 7l'*7~Q7l' , P ikermi · Samos · N. China ~'f;r:I) Ponti an V;t~1) Z DANSKY, 0 .: op . cit. , p. 48.
2 ) _ZDANSKY, 0 .: op. cit ., p . 50.
3) T EILllARD de CllARDIN, P. & YOUNG , e.c.: On the Mammalian Remai ns fr om t h e Archaeo lo:g ica l S ite of Anyan g. Pa l. Sin ica, Ser . C, V ol. XII, FaRe. 1, 1936, p. 19,

-

43-

434

11lf.

Oriental autbors(PILGHDI, TETLf-IAHD,),
STmToN

Siwalik Series

P leistocene

Boulder Conglom erate

American authorse\,L~'l"l'JJEW, OSBOEN,)

GIDLEY, COLBERTUp.

Pl eistocene

Pinjor Zon e
Tatrot Zon e

Pliocen e

Dbok Pa tb an Zone
Mid.

Pliocen e

Nagri Zon e---

Chinji Zon e

Mi oce n e
L ow.

K amli a l Zone
* 1i:R~JfI<v~~~' C Jil1 ;f,1bfn~f.jlff!'MI';JiWU: I..- lft", oMiocen e

r:1l J1[ Siwa lik Seri es K Ii Dho!;: Path an Zone Klf<: I(, t£. < , 5 :fill!I!fl?;Q iO,;ft,;;'o ;;fs:'Iif ~ P onti an ci'" ;;';(T1Jt,r ;;, Jjfffl;!<V- " 'ib;;, 0 COLBEH'l' KJ:;;, c , ~i~ , Chinji Zon e J: ~ t
H ippm'ion theob~ldi (LYDEK KEl,) ;Fi.lt;:i'" ;;' 0 1) Hipp a?'ion ~bfHs il)iIDt;li ll[[~U l.JlI ~~*;¥J,r:j:t lVL\'.rTITEW-OSBORN- llit <v 'lU~T ;;' )Yi"'ib ~ , 1ll1*III1ll1:kIM1<v H ippa j'ion 1!;-/('fJEi- v;t

V a len tine beds c Jil1 11~H-\,; X Ii J: ~ !ilIi I..-i! I(, <v -r: t,rIt ;f1-Vf fJ: iO, /!;l c 1i1ht~ <v 1itfR-r:<v ;;, 0 11t:.. ··::;,-( L Oll-er Siwal ik <V C hinji Zon e Ii~f;f& P onti an -r:ib;;, c ~~ <V
1'!M:ilI 2klflll[;fjU1ll1<v J}~1k li 7.P l.;;' ,." 9" -, f- :;, 7 t,r~J/. KiP;2i1i-~';;' ~ 1% t,r " ' 0 .e,.. EI m*<v Siwa lik Series-r: ib ;;, 0G') 1I~H'\';KlMd T;;' ~J/. Ii~G') :!m <'ilii tc:' I..- dW, p!ii:i.J; ib ;;' 0 jiiSOk K H ippCLl'ion gmtwn j:·IH:J~ 9"~.J H ippcL?'ion~.J )fill:$t;-r:t,r V) ;ft, Vf t,r iO, /!;l G') :i.J' , ~rflJl- K V:l: 7fur1~ c T ;;, )Yi-r:ib ;;, 0 Si walik <V WJ ~~ lir,:i;t Il~H::5/§:[&'i- ;;' ~%- I'i PILGlm[ <V ke

J/.lilOC t tz<.'!:'; "· ib ;;, WJ~!¥f.':j:' cv ~ -1l1S<V t <V K1l1ij. Ho: i! Z ~ 'lli'dvlj -:;j- ;;' ..Jm , - ITii'l'T5f:U-r: t ib ~ , I&: Hti.Lt K li
1i!!5f:Ut,r ll:lit t:-", ;j;, Z rc:. ~iiJ iiti'" ;;, II;YIi , II'If rc:. J: VU: ti L "'Sf mI'C II§ ", ;;, 0 ;(Pf:Ut,ril\:f,lH;t jnJ lI'If K;ffi!f· tJ: Jitf,if -r: t ib ;;, 0~fi* Ii I;iWl <v 1jiPJ rc:. )ji,t ", -c Ii k:l1 ~iMJ'rt'{ 1iU a: ¥.,'L.\, t <v -r: ib;;, 0 H ippa?'ion theouald'i c: 7C H ilJ]Jw 'ion <V l'jlr.!:l G') t <V
fJ: ~ c i - ;;' P ILGR IM' <V J/.1rF: X li~i}·I!!:Jf ~.J Hip Fw'ion Ii U nknown ]'fe?'ch'ip pus (Valentine beels 17.J t <V c
1Mdf;l'i fJ: ;! JYr <V ) J: ~ lii* i'" ;;, t <V tJ. t, AI i: T ;;' TEIJ.HAHD d e GEIAHDIN G') J/.i;.¥K i'f I..- COLBERT cv ttn < illU ~ 1!f
5Ei'";;, ~Jj: I'i'f:(I!:lWi*Ih1~"' o L ower Siwalik = Upp ermost Vindovoni an, M.iddle Siwaliks =Pon tian, Upper
0

Siwaliks=Astian or Upper m ost Pliocene c 1" ;;, TElf.HARD de GIJAIWIN cvk\J/.Ii ;&',W-r:<b iO, 50

JiJ *Siwalik Series Ii EPj~cv l'fIj~LillJJ ~:M,"";fTM c IMJf;f, i~ <, ttnJ:<v5L1f'(. I(, ~~¥T;;' }JIi': k U l},..\i'&:, ft K!&\J/. H l!; .-..:: -c ID': ,,'tc o~ ;fr\ OJ 1t:£ .~f'j:~ r-1f; VC lf!;c' ib Q 0

l!lXlfl ff,!,f:.2f-4t.t I) Anchi th c?'i~Lm h ypo hilJPoiclcs lVIATS Ul\{QTO iJ{ZDANSKY f'j: 1.L1f!!i1~f1*f.#'ff,!,f: Liu-Wa n-Kou OJ Hip pa?'ion R ed Cla y .t l? A nchit h lJ?'ium 5liJ1 1fl,OJt~~ib Qtw?*~¥R* L- -eiiSQiJ', .2f-!&ii'.t i? ~ vcffi!JJ- C' ib Q o~) N:lliJ·J\%~t:t§}'~tI\~Mu!t*i OJfr.\~f:trr1fi1E (?) .t I) Pliohip pu,s (4- 8 OJ
Jifr~t.~(:!f.lPf=j) iJ{1±\ 11: '!: OJ¥Ri!fiJ,ib Q iJ{ , 1if~JtVCr& I) ,!: Tl/'Lf:nml];j~ ib Q l~-c' ib Q o"l #tf:j't/:!tvc ;I~",,-e~, *;f~ f'j:fM'& '!: L- -e .~1t:£ vc. :li!U: l/'LtJ..
~-!J.lf(\!f:J-itrW:j]'[~) I I *~~~ :J:F OJ ~@.~:1I/;f:j~4'!IJ.t i? f'j:, mn ~ vcj;~ ~ l/'LQ iJ>, ;l\'~!1'Mi:ctfii'#*f'j: 1C l:.5f~ 3 IH:/® ,!: 1CT~;; 4 IH~I ll'if vc iill1 ~· tJ.. "" 0""01) COLBER'!', E . : Siwalik Mammals in the American Museum of Nat ural Hist ory. Trans. Amer.
Phil. Soc., N.S. , Vol. XXVI, 1935.
2) lVL~'.r S(JMOTO, H.: Description of Some New F ossil Mamma'ls from Kani District, Provoof Mino,wi th R evision of Some Asiatic Rbinocerotids. Sci. R ep. T6hoku Imp. Univ., Sendai, Ser. II (Geo\.}Vol. V, N o. 3, 1921, pp. 77- 78.
ZDANSKY, 0.: . op. cit., pp. 17- 20.
3) {~7k1l1}!rt: IIjIl~Li/;'i'i, Wilk: rU~ r,l , !Il!. 'i:t,!;J~2k vt ti'.1:. ~M~, 1934, p. 58.

- 44-

435-

f.'I1\7kIW:r:K J:: i? E. caballu,s j'o ssilis C UVIER (7)ftJ:.5f~ 1 :& U'5f~ 2 * 1::1 1if!;6;: j~~ t. GfL-eJlS· .Qol) :f!;k*~ 31 · 32 ~"tc·*lJ~(7) )!g 1~U:. ~J/'l , I~lJfil)t1Il ~fi~" EI (7)* )!gc~ GIi\/"lC ~J/'lk o flJ~K lmewaZs7di

J:: i? *l" ib.Q ;6;:, 7l~ G < sanrnenien sis ~f- C ~ffii* ib .Q K~r-~".Q ~ C }t',fiJ/'l.Q 0 f.w'7ki'W± K J:: .Q c,
II'iJ t. < E. caballu8 j'o ss iZis CUVlER C ~ fL.Q 't (7) iJ':%J!I[JI&bJ(J:.~f[\~I~j:':~"J· J:: i? $ t. GJ/'l-em .Qo ;!'-t(7).
;rMJEa~utlE fi 7F HJj c · ib .Q 0
gpi? :;)t:tt [aJfJ'la~l" ib.Q ;6'*;P1\ 8~fkUto).~ fi li1iiWtZ,c; fL, fJfi~(7) cabazz,us K AfL GJ/'l.Q ;6' gJ K fi t],
7f~(7) t,(7) t,1'f.T.Q o Z.~;6' sanm f!n iensis Klli:\/"Ij), p1'z ewaZskii ~ h f'mionus Klli:\/"Ij)'fi , !'~*1q¥
jk L.fJt\/"lJiJfl" ib .Q 0 VEfriJ~jt (7) j[}JiII?,J;w('j)v .. Jv 1::." y 5:t (7)j[}Jtj&)}~~j)'friJ J/'lj» Km'J T .Q 4j~:t- ffi Tc' ib G J 0~ K 1&~(7)WIJfi*j,jl* J:: i? 9;11 GtLtc: o*;PI\(7)1rj~!lVJif'?,Jl¥t K t, )~ fi ~ J ~ <~Yl!. ~ J/'l li \/"1 0 ti <*~~mljJo\t:l::(7) $il-j- ~ tLk M'K J:: .Q L fifb~.R~*~EEI ?iJj~1rj~ ql~~&:±f,[~3t1tlt'r Kfi S us nipponictts miyae MA'l'S UMOTO ib i?, .~fi~i:lj ~5:t:I::.
@3t1tJ!, Kfi T .Q Ill ) j), G 1::I1~H"J l~i· ;;{:jJJfj)jxJ!f:t'~· j)>I±l l'· , ~Jf-filllIJjiJ:: i? mrf.'~·· ~1J·· ~:'WJ· · 81~i~:t- ,':1:\
L., )J~1~~~O: ±*~JI11tl'BEt1rj~ J:: b i=:1 ~r t , iiili~~1±\7k*~ 1±\ 7klliJi!1r!ii'1rj~ J:: i? 1::J1l!i· f'lrlfr · 11t~·~:t­
I±\ L.k i3:J c', £:(7) ~1 )jX'~j'1JffYEK J:: .Q C*;Pil1j<;'j'K.7Lj'I"(7)::Ef,if.n;lf1-tilllWliK ft!. 1±\ ~ fL .Q.~ fi~~!J \ l" ib0tc:
Eb C' ib.Q o~) £:fi r-5Rfi 1±l 71<.1rJ~(7))]#t~· tt& L. -e J:.i1!];(7):ftj:1 ~ tj'!H;J'{Hf'tV7 , 3Z.:ft~ -r!:i1-t (7) )!giJ'~~!J\
Ii i? L. c c :t- t~~ L.~~td', :f!;1JBi:!-H b III *-1:[- .Q j»(7) fl-l'] ffi1K:lz: A .Q c c fi c ' ~ tQ o mH "foJt::Ef,if.f1;J.1-t(7)~*j)'W:m!; E.(7) ::Ef,#~;lf1-t;j(WJ.!),f&~IJ;lj-1tilljtJli J:: i? ~Yl!. ~ J.:, ~ :l:$ji,: c II'iJ -f:ill:~j c ~PJ C6 -eTtJfJ. .Q K,
JJfj~ (7)5K. -efi.< 1&~(7)W.;lfft*WJ~.K~-e~C6-effilli:Wffi:t-t,c~~ .Q ~~ ~~ ~~~~~tT'i ;tc: c .FG', J.J

c ~0-emGJ/'lJ.:, %Kffj, .~(7)lli'j~l· fiiN EI *(7)::ER#f1;J.tt;ti1llJ,K± c L. -ell! GfL
0.Q*~c'ib .Q ;6" ~JJfj~1jli~H\5\(IIi*J~"J'*ln mJ1rj~H€(7) ,~(7)ID'~'j~' iJqill:Z;f(7)~t~K ib b (Re,'?;, N o. 59879). ~;j~ffIU\:K J:: .Q c ~ L. -e 1Eli1liK1rj~ffj),:;r;.j),0tfi L. \/"I c (7)4jI--:c' ib J.:, iJ', 1*ffAA!l~iJ'1U!. (7)ill·j~· K 1];J,-em.Q (7) c'i:t K ~-',Jr L. -e fi3: <0::E@f1;J.tt (7)*;PB.~iJ'!J \*c· ib0k c :j\:K,

gJ'l5iIj:A1:J'J(7) namacZicms ~K 1'l' 0k.~ t, 3Z. h cmion~ts
K 1];J, -e ;J,;}]~ c' ib0k4j~fi!f\lI* ib.Q 4JI- c,FG', fiJ/'l.Qo S2~S nipp onic'us M ATS UMOTO j)'''IJ'8'11 #tf:l1-lJtJ:: i?
;Pf,if.f1;J.1tK][ i? ~ff. L.k:ftj:1 <, *1J1*~(7).~c 1l'iJi1;1r-:M K ~1-J. L..k(7) c 'ib J.:, i 0'/J'o h emion2tS (7):ftj:1 <7G~VN~ (7) t, (7)j)'3Z.fi p 1' z ewctZslcii ~(7) .~iJ'~iWa~'MZ1Li=K J:: D1£1t L.tc 't (7)c' t, ib G J. j)'[JfI)t:(7)1Mt:J;tI! ib J.:, M
JjJi\j[c' ib .Q 0i!!.~i!

;fi1l-'t-.~ K*}] iJ1~~il~-tt ~ tl-::5 '7 '" '7 """ Equus sp. V;t'l3''',!~r, co 1t;o!o!~ r.j:' fC V;t , J1:~ co;fill 0.0 ~ Q) co :{j:;(£ V iJl@. K1iiL1' -3,(0i'll'i;{fC J: ~ ~YJ.r!I'H ~ ft)\. ~ ~1.. t;: ~ <7) <?ytUl < -C, ~Yl
T1if i3 \/'.1 tl"m -C <b ::5 co -C1:'kc, fhl:l~.lc~ V;t~ -? 1:" m~;f1.. Q

0

K I'f;t ~c. l' ::5 0

1) TOKUNAG A, S.:
Vol. 5, No. 3, '1929.

2):&:~f.i\~A:

Mammalian Fossi ls found in Limeston e Caves in Korea.

;o:(,S: I1'IH-I(;I7J~~K lhfd 1..-1:" , XI!MJ"\~f; );f"

-

45 -

1lt 40 {(&, ~1 4 }'J}c, 1925

Proc. Imp. Acad.,

436

fm

1!'If

On Some Japanese F O'3sil E quids(Resume )By
Tokio SHIKAMA
Fossil equ id remains are very r are in the J apanese Islands, probably owi ng to t he particularlaud condition that ptevailed during t,he later Cenozoic, which is adverse to the life of eq uids andon the contrary favourable to that of Proboscidea. In Middle Pleistocene, ~ome Manchurian or
Nor tb Chinese elements evidently migrated into the l and. as such remains a re foun d in 1be U pper
Kuzuii beds of T otigi pl'efect ure, the sand and gravel bed of the Inland sea, and the Tukinoki
as phalt bed of Akita prefectUre. The last mentioned deposit, yielded the eq uine tooth now underconsideration; other mammalian remain s previously descri bed from the same deposit are Palaeoloxodon namadictts yabei (MATSUMOTO), S(( S nipponiCtts nipp onictts MATSUMOTO, Ce?"lI'lIS (Sika) ezoensis
H EUDE, and Canis sp. This occurren ce, the writer t,hinks, indicate th at some steppe an imals mighth ave invaded into the J apanese Islands in assoc iation with the other Manchurian elements,
The very fragmenta l eq uine tooth f rom the aspha lt bed, about 22 mm long, 23 mm wide and 31
mm high, probab ly belonged t o a median-sizet! eq uid ; it has a flat protocone as in E. pj'zewalsl,;i'i
P OLIAKOFF or E . caballus LINNE, and relatively weak e namel plication, weaker than in R. sanmeniensis
TEILIIARD and PIVE'r EAu or in E . leptostylus MATSUMOTO, The fossil remain may represe nt a species
perhaps alliet! with E. hemionus or ot her small sized equids,
Another tooth, shown in Fig, 1 b is a left lower last molar, about 30 mm long, J2 m m w ide and
43 mm high ; it is q,uite simila r to the corresponding one of E. caballus, It was found lying on the
sandy sh ore near Hitati-mati, Ibaraki prefectme; it may be a fossi l or subfossil, or even its being aquite recent one is by no means exclude d.
In connection of the r espective ant iquity and sequence of E. sanmf.niens'is, E, p1"zewalsk'i'i and E.
cabctllu s, it is questioned whether the eq uine tooth from Honan described under the name of E .
leptostylu s MNrSUMO'l'O is not referrable to E , p jO
zewalskii, because tIle correspo ndent tooth of the latter
species differs not milch from the for mer in dimensions !tnd enamel plication, Which of t he various
names-E, (E.) caballusjoss-il-is Cuvllm, E. (E, ) cctballus hptostylus MATSUMOTO, E. ,( E. ) caballus pj,zewalsk-ii(POLIAKOFF) and E, (E.) p j,zewalsk-ii POLIAKOl'F -- should properly be applied to the Anyan equid which
is sometimes regarded to be a domestic form, is a question d iffic ult to answer.
At present wide divergence of opinion exists be"tween the American an d oriental authors as toth e geological age of the H ipPa?'ion bed of the Siwalik series ; t he present writer rather ten ds tothe view held by PILGRIM or TEILIIARD de Ca.illDIN, as t he Americans seems to bim to be t oo
arbitrary in rega.rding the :New World as the n ative la nd of Hippa?'ion. For intercontinental correlation of fa una, the use of certain single group of animals instead of the whole fmma m:}y pro ve
to be of considerable value, though sometimes quite dangerous. ,Beside the two specimens cited above, there a re some others which have previousl y been r eported
by Dr. TOKUNAGA, namely severa l specimens of E, cabal/us from Keisei, Tyosen, and Kuroi vil lage inH y6go prefecture; farther t here are several remains from Neolithic sites i'eport ed by Dr. H ASEBE,
especially of Kyushu, which specially attracted bis atte nt ion to the small size of them. Hence one<:an at least safely express that some small sized horses li ved in Japan from the Middle Pleistoceneof the Neolithi c age, contemporaneons with Sus n'ipponictts MA'rSU~IOToo The small size may beinterpreted either as its specific characte r or as due to island life,

-

46-

64.

On the Japanese Species of Elphidium and
Its Allied Genera.
By

Kiyosi ASANO(Contribution from t,he Institute of Geology and Palaeontology, T6hoku Imperial Univers·i ty.
Sendai, Japan: Read December 18th; received December 20th, ]937.)

A study of the various collections of foraminifei'a now stored in the I nstituteof Geo logy and Palaeontology, Sendai, Japan, has rendered the writer possible to·review the Japanese species of Elphiclium and its allied genera, both fossil andrecent.
Of many different species of Elpidium distinguished up to the present timefrom the Japanese material, valid are the following eleven species, namely:
Elphidit~m c 1"l;spt~m

E.
E.
E.
E.
E.

(LINNE).

subgranttlosum n . sp.
cf. f abum (FICHTE and MOLL)
aclvenum (CUSHMAN)
jenseni (CUSHMAN)
c?·nticuZatttm (FrcHTEL and MOLL)

yabp,·i n . sp.
E.
E.
yezoense ASANO .
E.
lcusi1'lJense n . sp .
Elphidiella nagaoi n. sp.
Polystomellina discorbinoides YABE andHANZAWA.

Of the mention ed forms, 1) E. c1'ispum, E. subg1'(~mdos~~m and E. d. jabum
are widely distributed in Japan, both r ecent and fossil, 2) E. ad~'em~m and E
jenseni occur commonly in the r ecent or Plio-P leistocene mater ial from the Pacificside of Japan, but are very rare in those from the Japan Sea side, 3) E. cmtictdatum is only known from the Pacific side of Japan, both fo ssil and recent, 4) E .
yabei and Elphi diella nagaQi seem to be confined to the Pliocene of Setana,
Hokkaido, and their living representatives are still unknown, 5) E. yezOe1?Se andE. h~si1'oense are only known in the fossil materia l from Hokkaido aud Karahuto,
and 6) Poly.stomellina disco1'binoides is a characteristic Pliocene form of Etigo,
Japan.
Further, the species belonging to 1) and 5) find their closely related form s in
the Plio-Pleistocence of the W est Coast of North America, and those belongin g to2) and 3) are now living in th(:) Indo-Pacific region .
As to the distributions of species, these ceems to exist. a very interesting relationship between Elphidit~m and Oassidulina, as previously stated by the writer;
-

47-

-582

K . .8.SA.c'lO

namely some of the extinct fo r ms of Elph'idium treated in this article show an
intimate co-existence with certain extinct forms of Cassidulina. Thus, Elphidi1Lm
yabei and Elphidiella ?wgcwi are found in aSEOciation with Cassidulina yabei
ASANO and NAKAMURA and C. setanaensis ASANO aud NAKAMURA, an d Polystomellina
disco?'binoides associated with Cassidttlina yab ei, fur ther, E. yezo ense is found tobe always associated with Pecton ta7cahushii YOKOYAMA, a characteristic Pliocene
mullusc of Japan.
The species dealt with iu thil: i article are derived from many geographica1.ly
isolated localities, which are enumerated bel ow in order from north to south , asfollows:I) . Karahuto (a) Kal'agai, T~l'ie- gu n. ColI. Y . SASA,(b) Rutaka-m at i, Rutaka-gun. CoIL Y. INA!.
(e) Minami-Rokusenzawa, l'oyobara-mat i, Toyohara-gun. ColI. Y. I NA!.
II). Hokkaido (a) Zarigawa, Numata, Uryu-gun . ColI. S . ENDO .
(b) Kaigarasawa, Kuromatunai-m m a, Suttu-gun. Co1l. IV. I-IASI:M:OTO.
(e) Nakanosawa, ditto. Coli. W. HASI:M:OTO.
(d) Nakan okawa, Yubetu-mUl'a, Suttu-gun. ColI. T. NAGAO aud Y . SASA.
(e) Pirika-E ki, Tosibetu-mUl'a, Setana-gun . CoIL T. NAGAO an d Y . SASA.
(f) Hanaisi-Ek i, ditto . ColI. T. NAGAO and Y. SA SA.
(g) Omagari, d itto. CoIl. 'r. NAGAO and Y. SASA.
(h) Maruyama, ditto. ColI. T. NAGAO and Y. SASA.
(i) Minami -Kanehara, ditto. ColI. T. NAGAO and Y. SASA.
(j) Hal"Utori, neal' Kusil'O City. Coll. Y . SASA.
III). Aomori, (a) Ut imanbegawa, Okuna i-mul'a, Higasi-TugaJ"u-gun . ColI. K . ASANO .
(b) Taya, Iwamisannai-nml'a, Kawabe-gun. ColI. K . ASANO.
Akita,
Niigata, (e) Sa wane, Sado Island. ColI. Y. OZAWA.
Toyama (d) Kutta, Misima-gun. CoIl. S. HANZAWA.
(e) Natukawadan i, Kitadyo-mura, Kariha-gun . ColI. S . H ANZAWA.
andIsikawa. (f) Simotaka-m ati, Kariha-gun. ColI. S . HANZAWA.
(g) Tagawa, KOllade-lllUl'a; Nisitollami-gun. ColI. M . NAKA~WRA.
(h) ilorakuzi , ditto . ColI. K. ASANO .
(i) Amaku zi. ditto . ColI. K. A SAl'W.
(j ) Hiradoko, Syoin-mura, SUZll-gun . ColI. T. l\IATUSI:M:A.
(k ) Jagaya, Kosaka-mura; Isikawa-gun. ColI. M. NAKAnfURA.
(1) Onma, S'akiura-mm'a, I sikawa-gun. ColI. M. NAKLU1:URA.
IV). Tiba(a) Sematanoseki, SitO-mura, Itihara-gull. ColI. M. NAKA~WRA .
(b) Olluki-mati, Kirnitu-gun . ColI. F. UEDA.
(e) Sanuki-inati, ditto. ColI. F. UEDA.
(d) Moroiti, ittu-mura, Itihal'a-gun. ColI. F . UEDA.
(e) Tosaki, Tomioka-m ura, Kimitu-gun. ColI. F . UEDA.
(f) Higasi-Knniyosi, SitO-mul'a, Itihara-gul1 . ColI. F . UEDA.
(g) Mandano, Makita-mura, Kimitu-gun. ColI. F . UEDA.
(h ) Ititoku, Koito-mura, Kimitu-gun . ColI. F . UEDA.
(i) Nisi-Nagata, Toyohusa-mura, Awa-gu n. ColI. F . UEDA.
(j ) N uma, Awa-gul1 . ColI. S. NO~WRA.
V). K anagawa (a) Tomioka, Kanazawa-mati , Kuraki-guil. ColI. 1VI:. NL\K"\:M:URA.
-

48 -On the J apa.nese Species of ElphicUum and Its Allied Genera.

583(b) Hitol'izawa, ditto. ColI. M. NAKAMURA.
Col I. lVr. NAKAUURA.
(d) Teramae, ditto. CoIl. M. N.LI.KAlIWRA.
(e) Nozima, ditto. Coll. H . YABE.
(f) Ueki, Tatna nawa-mul'a, Karnakura-gun. ColI. M. N.LI.KAMURA.
(go) T a kaya, MUl'l1oka-rnu1'a, K amakura-gun. ColI. F. UEDA.
(h) Osiki1'i , Hong6-mura, Kamakura-g un. Coll. M. NAK.LI.lIWRA.
(i) .Asahina, K a makura-mati, K amakur a-gun. ColI. 1\1. NAK.LI.lIWR.LI..
(j) Utik osidani, Toyota-m ur a, Kam akura-gun. ColI. lIf. N.LI.KAMUR.LI..
(k) N agamuma, Toyota-mura, Kamakura-gu n. CoIl. F. UEDA.
(1) N i sinota ni, Hon g6-mura, Katnakura-g uu. ColI. l\,f. NAK.LI.MUR.LI..
(m) Simo-lVIiyata, Hatuse-rnura, MiuTa-gUll. ColI. K. ASL\NO.
(n) Okine, Hatuse-mura, Miura-gun. ColI. M. NAK.LI.lIWRA.
(0) K ami-JHiya t a, Si rnoura-mur a, Miura-gun . ColI. lVI. N.LI.K.LI.lIIURA.
(a) Dain iti, U kal"i-mu1'a, Sy uti-gun. Coll. R. AO KI.
(b) Asuka, T aruki-mma, Ogasa-gun. CoIl. R. AOK I.
(c) Hosoya, Haranotani-mura, Ogasa-gun. ColI. R. AOKI.
(d) Ketienzi, Nan g6-mura, Ogasa-guu. Co il. R. AOKI.
(a) loki, Aki-g un . ColI. II. YABE.
(b) T6nohama, Aki-g un . Coll. H . Y.LI.BE.
(c) One, Ak:-gun . ColI. II. Y.LillE.
(c) Yadu, ditto.

VI).

Sizuoka

VII). K6ti

The geological distribution of the species is lobulated below:~
eSe~~:Ci~~dLocalities

~~

~ ~

~

~~

~§

&::

~

.~ J

. ~~

.~ ~

...::: '-"~"'9~t~ .....

~'B

~~

.~ ~

~ ~ .........
~~ ~~ '"\j~_ __ __ _ __ +_~-'---_ ~ '" ~ ~

~

~

~.~

~
~
-~ ~

..§;.~
i:;:;l

~ t;

~~

~

~
~
~ ~

~.~
~ ~

~
~ ~~ 6u~ Ij"i:;:;l~;g.~. ~ f,E~t........ <j

~~

~
~
~ Ij

.~ .:s

..&~

~~

A
R

(b)

,,~~

~~

.~~

(a)

Ka rahuto (!)

~

.~:5
R,~

~ ' C;~ "C; Geological
G:~

~ I;:

R. ~

age

~ --.:...:"''---I----Pliocene
1/

II
(cl
R
- -- - - - - - 1- - - - - - - - - - - - - - - - - - - -- - -- - - - - -- - - -1 - - -- (a)Pliocen e
A
1/
C
(b)
F
R
F
F
R
1/
(c)
C
F
R
R
R
1/
(d)
R
R
C
//
Hokkairl6 (II) (e)
R
1/
(f)
R '
C
R
1/
(g)
R
F
1/
(11)
R
R
1/
(i)
R
(i)
C
F
C
PleistoceneF- - - - - - --1--- - - - - - - - - - -- - - - - - - - - - - -- - - - - - - - 1 - - -- (a)eRR
Pliocene

R

~)

(c)

C

F

(d)

C

R

Aomori,(e )Akita, Niigata, fToyama a nd ()I sikawa. (III) (g )

A
A
F

(h)
(i)

C
F

R

'
R

C

R
R

R

F

R

R

R

R

//

C
C

R
?

o

R

F

II
II
II

R
R

II

?
?

R

R
-

49 -

II
II

K. ASANo

584

Geologicalage
Pleistocene
Pliocene

(j)

(III)

R

(k)

R
R

(1)

II

- - - - - - --1- -- - - - - -- - - - - - - - - - - - - - - - - -- - - - -- -1--- - F(a)Tiba (IV)

(h)

R

(c)

F

R

R
R

(d)

C

(e)

F

(f)
(g )

R
R

(11)

R

R
R
R

Pleistocene
Pliocene( ?)

R
R

II
II

R
R
R

II

II

R

II

II

R
R

(c)

F
F
F

(e)
(f)

R

(g)

C

R
R
R
R

Kanagawa (V) (b)
(il

R
F

R
R

(el)

(j)

(k)
(1)
(m)
(n)
(0)

F

R

R
R

F
R
F

F

R

II

R

R

R

II

R

II

R

II

R
R
R
R
R
C
R
F

II

R

R

II
II

II

R
R

II
II
II

R

II

R
R

II

II

R

- - - - - - - - 1 - - - - - - -- - - - - - - - - - - - - - - - - 1 - -- - Pliocene
(a)
R
R
II
(b)
R
R
Sizuoka (VI)

(c)
(d)

- - - - - -Koti (VII)

(a)
(b)
(c)

II

R
R

R
R

II

- - - - - --_·· _ - - -- - -R
F
FF

Pliocene

R

II

R

R

R

R

- - - - -- -1----II(A · ... Abundant; C·· .. Ccommon; F·· '" Few; R · · · · R are)

Before going into the descriptions of the species, the writer wish es to offer
his warmest thanks to Prof. H . YAB E of the Institute of Geology and Palaeontology, T ohoku Imperial University, Sendai, Japan, under whose supervision thepresent work was carried out, for kind suggestions and corrections of this article
before publication; he is also indebted to those who kindly submitted their specimens to him for study.
-

50 -On the J apanese Species of Elphidium and Its Allied Genera.

585

Family N onionidae REUSS, 1860
Nonionidea REuss, Si'z. k. Ak Wiss. Wien, Math-Naturw., Cl., vol. 40, 1860, p. 221.

Subfamily E lphidinae GALLOWAY, 1933
A Manua1 of Foraminifera, 1933, p. 269.

Genus Elphidi~t?n MON'J.'FoRT, 1808
E lphidium MONTFORT, Conch . Syst., vol. 1, J80S, p. 14; (Genotype, Naut'ilus macellus FICElTEL and
MOLL, in part) ; CUSHMA~, Contr. Cushman Lab. Foram . Res., vol. 3, pt, 1, ]927, p . 49, pI. 10, fig. 5.

Test free, pl a nispiral, bilaterally ' symmetrical, mostly involute; chambers numerous wit h
distinct sutures, either depressed or raised an d limb ate with sept al bridges a nd c1epressio'ns;
wall calcar eous, finely perforate; aperture a curved slit or row of p ores a t base of septal
face on inner periphery, 01' num erous por es on septal f a ce.

The genus first appears in the Middle Jurassic of Russia (Ornatent.hon), later
it found its best development in the Tertiary, especially of the Vienna or Paris
Basin, an d became gradually common in succeeding formations up to r ecent times.
I t is very abundant in some you nger Neogene deposits of W estern North America
and .Japan. Most of the ' recent species h au nt in cold ' to warm, shallow waters.
Elphidi~vrn is the largest genus in the family Nonionidae as to the numberof species and individuals. The genus seems to have been evolved from Nonion
which is th e most prim itive member of the family, by developing pores along thesuture lines. It is sometimes very difficult to distinguish a simple Elphidiu?11.
f.rom Nonion owing to the very small size of the "eptal pores in the former, buthigher species of the former are easi ly distinguished from the latter by t heircharacteri stic septal bridges.
Elphidi~t?ncl'ispwn (LINNE)

PI. 14 (3) figs. l a, b .

Compare with: CARPE~TER, W. B ., Introduction to the Study of Foraminifera, 1862, p. 27S.
CUSElMAN and LEAvIT'r, On Elphid'ium macellwn (FrcHTEL and MOLL), E . stria,to-]J"unctatum (FICElTEL and MOLL) and E. crispum (L.), Contr. Cushman Lab. Foram. Res., vol. 5, pt.. 1,1929,
pp. 1S-22.

The species orig inally recorded by LINNE from the Mediterranean, seem to be
widely spread in th e I ndo-Pacific region. Con siderable variations exist in the
shape of chamber, umbilicus and septal bridges ; and its typical form has a thickbiconvex tef't with sharp periphery. Septal brjdges are conspicu.ously developed
and the umbilical region is fi lled with clear shell material in which there are a
few large, rounded pores. The writer h as exam ples from th e ¥iocene of Vienna
Basin and th e Pliocene of California, as well as numerous specimens from theI ndo-Pacific region. It suggest s that the Japanese specimens, especially those fromthe Pliocene of Japan Sea side, are hardly to be di stin guish ed from the Californian
form, they are more biconvex and more strongly umbonate than those from the
Miocene .of Vienna Basin. I n spite of these apparent differences, the two types
are similar with one another in their essential feature; and it is perhaps better
-

51 -

586

K. ASANO

to include both the types to the same species until more IS known of theircharacter s and geographical distribution.
Distribution in Japa n:
Recent: J a pan Sea and P acific, especially common in the shallow waters of vVakasa,l\IIutu and S iogama bays a nd coast of Onc1yuku, Tiba prefecture.
F ossil: Pliocene of Setana, Hokkaido (h, c, d, e, f, g , h ); Aomori, Akita, Niigata,
Toyama and I sikawa prefectures (a, c, d, e, f, · g, h, i, j, k, 1); Tiba prefecture(h, c, d, e, f, g, h, j ); K anagawa prefectu r e (a . b~ c, d, e, f, g, h, i, k, 1, rn, 0);
Si zuoka prefecture (a, c, d); and Koti pl'efectUl'e (a, b, c) .

Elphidium subgmnuloswn n. sp.
PI. 14(3) figs. 4a, h .

Compa re with: Pol ystomella si1"iato-pwn ctata BRADY (not Na,ui'ilus st?'iato-par1ctaia F ICEITEL andMOLL), Rep. Voy. Challenger Zoo l., vol. 9, 1884, p. 733, p I. 109, fig. 22.
Themeon g?'C£n'u/osa GALLOWAY and IVISSJJER, J our. PaJ. , vol. 1, No.1, 1927, p. 83, pI, 12,
figs. 15, 16.
Elphidill?n hughesi COSm[AM and GRANT, Trans. San Diego Soc., Nat. Rist. vol 5, No.6,
1927, p. 75, pI. 7, fig. 1.
Elphirlitlm decipiens HAD A (not COSTA), Sci. Rep. Tohokn I mp. Univ., Ser. 4, 1931, vol . 6,
No. ], p, ]26, Text-fig. 83.
Test rather small, bilaterally symmetrical, peripher y broadly rounded throughout, veryslightly lobulated in last few ch ambers, diameter about threc times thickness; chambers
infiatetl, 8-11 in last whorl, umbilical i'egion slightl y depressed, ty p ically with gr anula rmaterial; sutures depressed, sligbtly cu rved and each provided with small indistinct, roundedpores; wall fairly thin, fin ely perfora te ; aperture a series of small rOllllded open ,ngs atbase of opertural face. Diametei' 0,3-0.5 mm.; thickn ess 0.1-0.2 mm.
Holotype (Reg. No . 21418, Institute of Geology and Palaeon tology, Tohoku Im p ~rial
University, Sendai , J apa n), from the Pliocene of Seta n n, Hokkaido.

Up to the present time, this .'pecies has been referred by the writer t o E .
st1'iato-punctat'Lvrn (FICII'l.'EL and MOLL), but it may r eadily be separated from
FrcIl'.rEL and MOLL'S origil al (Test. Micr., 1798, p. 61, pI. 9, figil. a-c). Forms
more or less similar to the present species have been recorded by the authors
enumerated above. E. h'LLghesi CUCIlMAN and GRANT has sutures of different
characters and E. g1'ant~losa (GALLOWAY and ,VrssLER) a more granu lar umbilicus.
There are two varieties in the present form; one is ovate, and similar, in theoutline of test, to E. 91'antLlosa (GALLOWAY and ,VrsSLER), while the other isnearly circu lar and similar to E. h'LLghesi CuSIlMAN and GRAN'l', both forms occur
generally in association in . the material examined.
Distribution in J apan:
Recent: Shallow to deep watel:s of J apan Sea and P a cific sides.
Fossil: Pliocene of Setana, Hokkaido (b, c) and Pleistocen e of Kusil'o, Hokka ido(j); Pliocene of Aomori, Akita, Niigata, Toyama und I sikawa (a, b, c, d, e, f ,g, h. j , ki; Plio-Pleistocene of Tiba (c, el, e) a nd Early Holocene of N uroa (j );P lio-Pk stocene of Kanag-awa (cl, e, f, g, h, i, k, 1) ; Pliocene of S izu oka (b)ancI' Koti prefecture (b).
-52 -On the Japanese Species of Elphichurn and Its Allied Genera.
Elphidi~£rn

587

cf. jaburn (FlCH'PEL and MOLL)
Pl. 14(3) figs. 7a, b.

Compare with: Polystorne7la faba HERON-ALLEN and EART,AND, Trans. Linn. S'oc. vol. 11, 1916, pt.
13, p. 281, p1. 43, figs. 11-19.
Elph'id-it,m jaburn HADA, Sci . Rep. Tahoku Imp. Ulliv., sel'. 4, vol. 6, 1931, No.1, p. 125,
tex t-fig. 82.

HERON-ALLEN and EARLAND state that two distinctive forms are included
under FrcHTEL and MOLL'S species. The first or compressed type, which is nearest
to E. jah~£rn, has somewhat inflated chambers in involute coils, six or seven visible in the final coil; sutural lines curved and stro ngly depressed, filled, with finegranular matter radiati.ng from the umbilicus and giving a ::;tellate appearance tothe test, owing to the whi.tish granulation s in strong contrast with the hyaline
surface of the chambers; septal bridges few in number, are marked by secondary
sh ell-material which becomes visible when wet. The second or turgid t.ype, ismuch larger, the chambers being less inflated and more nL1merous, ranging up to
8 or 9 in the final coil; sutural lines depressed, but less so than in the compressed type, and filled with the Eame granular matter radiating from the depressedumbilicus, but to a lesser degree; septal bridges much more numerous than in
the compressed type, and come out strongly when wet., although rarely viRible in.dry state.
Both forms are present in the collections at the writer' s . possesion. But ac.curate iden tification is impossible without a stud y of the original flpecimen.
Distribution in Japan:
R ecent: Common in the northern Jap a n Sea, but very ra1'e in the Pac ific coast of
Jap an.
Fossil: Plio cene of Setana, Hokka ido (b, c) a nd Pleistocene of Kusiro, Hokka ic1o(j); Pliocene of Aomori, Akita, Nii gata, To yama, and lsikawa prefoctUl'es (a ,el, e, f, h, i); Plio-Pleistocene of Kanagawa prefecture (g) .

Elphidi n?n adventmi (C USHMAN)
PI. 14(3) figE. 3a, b.

1884. PulystorneUa subnodosa BRADY (not l\1UN~TER), Rep. Voy. Challenger Zool. vol. 9, p. 734, pJ.
lIO, fig. 1.
1904. Polystomella subnodosa l\11LLE'f'f , Jour. Roy. Micr. Soc. pt. 16, p. 604.
1922. . Polystomella aclvenct CUSHMAN, Carnegie lnst. Washington, Publ. 311, p. 56, pI. 9, figs. 11,12.
1924. Poly.,tornella aclvena CUSEIMAN, Ibid ., PubI. 342, p. 4S.
1928. PolystcrndZa advena CusmIAN, Ibid., P ubl. 344, p_ SO.
1930. Elphidium ctdvlmttrn CUSHMAN, U. S. Nat. Mus., Bull. 104, pt. 7, p. 25. pl. 10, figs. 1, 2.
]930. Elph-idium acivenum Cus maN, Florida St. Geo l. Sm·v., Bull. 7. p . 40, pI. 7, fig. 7.
J933. Elph'irZiurn aclt'enum CUSHMAN, U. S. Nat. Mus., Bull. 161, pt . 2, p. 50, pl. 12, figs. 1-3.

According to CUSHMAN, MUNS'l'ER'S Rob~£lina s~£bnodosa h,as a rhomboid test
apertural view, with the greatest width at the umbilicus. But the specimens
referred by BRADY and MILLETT to ;MUNSTER'S species are very far from the type;these 'may be ap~ig'n ed ' to CUSHMAN'S adVenurn.
. CUSHMAN describes this as follows : '.
III.- .53 .-588

K. ASANO"Test equally biconvex, periphery acute, often with a nal'l'O"IV keel, and in the last.
formed portion somewhat lobulate in side view, umbili cal region usually with a small bo ss;.
chambers numerous, somewhat inflated , esp ecially in the last-form ed portion; sutures de-·'pressed, the r etra( processes in t he later port ion ouly about one-f ourth the width of thechamber, in depressed channels above the s utures, the · intermediate portions raised; wallsmooth, t ran sluce nt, very finel y perfo r a te ; ap er ture a series of r ounded pores, at the baseof t he ap ertural f aee of t he chamber."

Our specimen figured is . quite identical with CUSHMAN'S species from thePacific.
Distribution in J.ap an :R ecent: Common in the sh allow waters of the Pacific side of Jap an, but rare in
J ap an Sea .
F ossil: Pliocene of Set an a, H okk aido (b, c) and Pleistocen e of Kusiro, Hokkaido,(j); Plio cene of Tiigata, Toyama and Isikaw a pref ectures (e, f, g, h, j, 1) andP leist ocene of Hiradoko, Isikawa prefecture (k); Plio-Pleistocene of 'riba pre-·fecture (a , b, c, d, f , g, i, j ); Plio-Pleistocene of K anagawa prefe cture (a, c, cl,.
e, f , g, b, i, j, k, 1, m, n, 0); P liocene of Sizuoka p ref~ c ture (a, b, c, d) and
Aki -gun, Koti pl'efe cture (a, b).

Elphidium j enseni (CUSHMAN)
PI. 14(3) fi b'S ' 5a, b.
1904. Polystomella macella (F. and M.l var. J ENSEN, Proe. Linn . Soc., "01. 29, p. 817, pl . 23, fig. 4 ..
1924, Polystomella j enseni Cusmr.nf, Cal'll egie Inst. 'Washington, Pub], 342, p. 49, pl. ]6, fi g. 6..
1933. E lp hidium jensen'i CUSHMAN, U . S. Nat . 1\1us., Bull. 161 pt. 2, p. 48, pI. 1], fi gs. 6, 7.

JENSEN origillally described this species under th e name macella from off the·coast of Au stralia in 100 fathoms. Its distribution is in shallow seas of the IndoPacific. The writer previously r ecorded this form likewise under the name of E.
mace ll~t?n (FICHTEL aDd MOLL), but th e typical mace llurn has a mor e convex andcompletely involute test . The Japanese material m ay be described as follows :T est . much compressed, p er ip hery slightly keeled, som etimes lobul ated; chamb er s numerous, n ot inflat ed, latel' t end ing to uncoil; sut ures dist in ct, slightly ra ised, septal bridges
alm ost ent irely occup yin g' ar ea bet wee n sut \lres ; umbilical region finely papillat e. Diamet el:
up to 0.6 mm. ; t hi ckness ca 0.15 mm.
Distributio n i n Jap an :R ecent ': Shallow wat er s of the P acific sid e and West ern Jap an Sea .
F ossil: Pliocene of Setana, Hokk aido (b, c, d) and P leistocen e of Kusiro, H okkaidO:.
(j ) ; Pliocene of Ni!gata and I sikawa pref ectures (c, cl, h, i) and Pleistocene of
Hir!3 ko ko, Isikaw a prefectu re (j); Plio-Pleist ocene of Tiba prefec tUre (a, b, e,
f , j ) ; Plio-Pleistocene of Kanagawa prefecture (g , j, k, m, 0); and Pliocene of 'Koti (b) .

Elphidiwn c1'atic'ulatum (FICHTEr. and MOLL)
PI. 14( 3) figs. 6a, b.
1862. Polystomella cmticulata CARPENTER, Intr. Foram., p. 279, pl. 16, figs. 1, 2.
1884. Polystomella cmticulata BRADY, Rep. Voy. Chalienger Zool., vol. 9, p. 739, pI. no, fi ge. 17, 16 ~1893. Polystomella cm t·iculata EGG ER, Abh. Kiin, bay, Akad. Wiss., 01. 2, vol. 1.8, p. 433, pl. 20,.
figs. 24, 25.
-54 -On the J apan ese Species of Elphidiwn and I ts Allied Genera.

589

1914. PolystomeZla c?'at'iculata CusEmAN, U. S. Nat. Mus .. Bull. 71, pt. 4, p. 43, pJ. 19, fig. 4.
1933. Elphidium cmt'iculattom CUSEIMAN, Ibid., Bnll. 161, pt. 2, p. 48, pl. 11, figs. 5a, b.

Test subg;obose, composed of more than 20 chambers in last form ed coil, 'broadly lenticular in spertural face, pcriphery ent ire, not keeled ; umbilic al area filled with clear shellmaterial with numerous por es, but not distinctly raisen, rest of su rfacc r eticula te ; aperture
distinct, at base of apertul'al face. Di amet er up to '1.5 mm . ; thickness ·ca. 0.5 mm.

The specimen from the ' Pliocene of K amakura, Kanagawa prefecture, h er efigured is not typical; those from the Byoritu Beds, Taiwan are composed of as
may as 50 chambers in the last coil. This species is a typicalJy shallow water
form of the Indo-Pacific.
Distribution in Japan:
Recent: Only known from the coast of K yusyil and Sikoku.
Fossil: E arly Holocene of Numa , Tiba prefecture; Plio-Pleistocene of Kanaga wa
prefecture (g) and Pliocene of Koti pref ecture (a, b) .

ElphidiU?f1 yabe·i n. sp.
PI. 14(3) fi gs. ga, b; lOa , b.

T est nearl y ci:vcular in side view, p eriphery broadl y rounded, lobulated, umbilical region
somewhat depressed, occupied by a large :!lat boss ; chambers numerous, 11 01' 1 2 in last<coil, of rather uniform shape, lat er ones slightly in:!lat ed; sutures distinct, depressed and·curved; septal pores often indistinct, very small ; aperture in early stages represented bynumerous pores at base of a pertm:al f ace, in adul t by many rounded pores in t erminal face{)f the last chamber. Diameter up to 1.5 mm.; thickness ca. 0.5 mm.
Holotype (Reg . No. 21419) from t he Plio ce ne of Setana, Hokkaido.

E. simple?; CUSHMAN which is abundant in the Indo-Pacific region somewhat
resembles the present fossil, but may easily be distinguish ed by the different characters of. aperture.
Dist ribution in .J apan :R ecent: unknown.
Fossil: Pliocene of Setana, Hokkaido (d, f, g).

Elphidium yezoense ASANO
Pl. 14(3 ) fig 11.

1937. Elph'id'iu'm yezoense ASANO, Trans. P';ll. Soc. J apan, No. 4<1, p . 120, pI. 24, figs. 1, 2; text figs. 1,2.

Test comparatively large, compressed, diameter a hout 4 ·times thickness, pe ;'ipher yrounded, usually lobulated, umbilical region str ongly umbonate, much raised with a large
boss of clear shell materia1, chamb ers as many as 20 in last fOl'lned coil; later chambers
slightly in:!lated with wavy p eriphery, earlier oue3 not in:!lated and entire along p eriphery ;sutures gently curved, depressed, retral pro cesses 'num erous and short, somewhat indistinct
in earlier ones; wall thick; aperture consisting of a low broad openings at ba se ofapertuml face; . occasionall y with circular pores 011 central portion of ap ertural fa ce.
Length up to 2.5 mm.; breadth ca. 0.6 mm.
Holotype (Reg. No. 21433) from the Pliocene of Takika wa, Hokkaido .

. Thi s species is allied to E. o?'egonense CUSHMAN and GRAN'!.', but is distinguishable by .t he different character of umbol1al region. It is apparently variable-55 -K. ASANO

590

in number and shape of chambers. The species is very common in the Plioceneof Takikawa, Hokkaid6 and Karagai, Karahuto, in both cases found solely v\Tithout
associ~tion of any other species of foraminifera; it is also noteworthy that it occurs
always in association with Pecten ta7cahashii YOKOYAMA, a characteristic Pliocene
Mollusca of Japan.
Distribution in Japan:
Recent: unknown.
Fossil: Pliocene of Takikawa beds, Rokkaido (a), Zintaki beds, K arahuto (a), andMaru yama sandy sha le, Karahuto (b, c).

Elphidi'LLm hlsi7'oense n. sp.
PI. 14(3) £g. 2.

Test rather small, compressed, periphery rounded, not keeled; umbilical r egion depressed, wit hout a definit e boss; chambers distinct, very slightly inflated, usually 9 to 11 in
number; sutures slightly depressed, septal processes very distinct, but occupying only an alTOW band above sutures; wall translucent, very finely perforate; aperture consisting of
one or more openings at base of apertural face. Diameter O.3- 0Amm.; thickness 0.1- 0.15mm.
Rolotype (Reg. No. 21420) from tl:e Pleisto cene of Rarutori, Kusiro City, Rokkaido .

This is a small species, but hardly to be confused with any other in theregion.
Distribution III Japan:
Recent: Nor t hern Jap an.
Fossil: Pleistocene of Kusiro, Rokkaido.

Genus Elphicliella CUSHMAN, 1936
Elphidiella CUSHMAN, Conk Cnshman L ab. Foram. Res., vol. 12, pt. 4, 1936, p. S9.

The genus is distinguishable from Elphidium by two rows of openings along
the sutures. The species of this genus are generally arctic and r each fairly largesize; they are:
E. aJ"ctica (PARKER and JONES)
E. hannai (CUSHMAN and GR.A.J."\TT)E . nagaoi n. sp.

E . sibil';ica (GoEs)
E. g·ro enla,ndica (CUSH:M:A:,)

Elphidiella nagaoi n. sp.
PI. 14(3) £ gs. Sa, b.

Test composed of about 10 chambers in last formed coil, in face view about 3 times aslong as wide, periphery broadly rounded" . umbilical al:ea slightly depressed, papillate; chambers somewhat inflated; septal lines with openings in two rows, but irregularly arrangedin earlier ones; a perture divided into. a number of small openings at base of apertural
fac e. Diameter up to 2 mm .
Holotype (Reg . No. 21421) from the Pliocene of Setana, Rokkaido.

This large and striking species is unlike any of the others recor ded. The
species is usually found in association with extinct forms, such as Elphidiurn
yabei D. sp., Cassidulina yabei ASANO and NAKAMURA, and Cassidulina setanaensis- 56 -

On the J apanese Species of Elph-iclium and Its Allied Genera.

591

ASANO and N AKAl\WllA.
Distribution in Japan:
R ecent: unknown.
Fossil: Pliocene of Setana, H okkaico. (b, d, f, h, i) .

Genus Polystomellina YABE and HANZAWA, 1923
Polystomella (PoZ ystomeZlinal clisc01'binoides Y ABE and HA NZAW,I, J ap. J OUl'. GeoJ. Geogr., vol. 2, 1923, p.95

T est similar t o Elphidiu,m in general structure but trochoid, pla no-convex, ventral side.
fla t tened, dors al side convex.

The m embers of this genus ar e now known from the Natukawa beds of theoil-bearin g N eogene T ertiary of Etigo, Japan and from the Miocene of Filter
Quarries, Batesford, Victori a, Australia (P. rniocenica CUS HMAN), and from 10
fathoms, Van Dieman f:i Inlet, Gulf of Carpentaria, Queen sland, Australia (P. a'U8tmlis CUSHMAN).

Polystomellina disco?'binoides Y ABE and HANZA WA
Polystomell'ina clisco1'binoides YABE and HAi'ZAWA, Jap. J our. GeoJ. Geogr., voJ. 2, p. 95, textfigs. a-c.

1923,

Foraminifera. are very common in the Natukawa beds, Etigo, Japan an d thepresent species is very common and distinct from other s. Living r epresentative
unknown.

EI*~ E l phidium J.% 1if fL $01~O!~ El ;<j~El't Elphidium Jlli 17.)1f:fL#.<A K IMJ l" "C 1'1 , 3::·"CTtFfY'i:: ~ tL "C .Q tc:V', ;w;':{R (/,)wld'llEK1t{ ~,Uiii :W)c l" "C BRADY (/,) "Challenger Foraminifera " K£ II'!J'I. ;(£;<j~MJ1Vc (/,) ili:(J;J.J~·U (/,) 1!, (/,) K
11 ;fill:v'1f$lc c Wd. t\? btL Q 0 ~~(/,) 1!, (/') K)ll.\: II' L 17.) Syn onym, geological lieU:: geographical distribution a:llX.:J1t.::.:> t~ 1!, (/,) "(' ib:O 0 Z ~(/,) 5 '!> "(', # K Elph:idium yabei, E . yezoen.'e, Rlph-icl'iella nagaoi, Polystomell-ina[&1(i LtJ < "C 1'1 tJ b !IJ.1" 5 K tJ"::':>fc. o

disco1'binoic7es 1'1, ~~~ El * ' ~~1f!fjJ;t · ;f1i'ij.\:(/,)ll!r'~-I!t (/,) ;,;", K El't L., -EI.-::.:>fn]'tL 1!, ~~;fill ,,(,ib:O i: c 1'1 , fll!'l't~Ll:tH! j"-.:.
~;fillc:3/f "" btL:o o

-

57-

65.

On the Japanese Species of Noniona nd Its A llied Genera.
By
Kiyosi ASANO(Contribution from tbe Institute of Geology and Palaeonto:ogy, Taboku Imperial University,
Sendai, Japan ; Read December 18th; received December 20th, ]937.)

A preliminary .study of the Japanese species referable to the four genera,
Nonion, Nonionella, Pse~6dononion and Ast1'ononion was undertaken and the r esults obtained are here given as the basi,' for farther research.
Family Nonionidae REUSS, 1860
Subfamily Nonioninae SCHULTZE, 1854
Genus NONION MONTFOR'l', 1808
Non'ion MONTFORT, Concb. Syst., vol. ], 1808, p. 210. (Genotype, Non·ion 'incl'as~atus (FICHTEL andMOLL); CUSElMAN, Special Pub]" "0 . 1, Cushman, Lab: Foram. Res., 1928, p. 204.
,
Test free, planispiral, bilaterally symmetrica l, periphery bro adly ronnded to acute;chamb ers numerous, closely appressed; wall calcal'eous, h yaline, finely pe :'forate, smooth,
sometimes with umbilical thickening; aperture, an arched, narrow opening at base ofapertural face on inner p eriphery.

The genu s Nanion is the most primitive member of the family Nonionidaewith the earliest appearance in the Jurassic. The f08sil s of Nanian and its allied
genera found in the Neogene rocks of Japan mostly belong to recent specie8, butthere are several unrecorded ones from the seas surroundin g our country. There
are 9 species of N()nian in Japan, namely, N. japanicum n. ,p., N. manp~dcujiense
ChURA, N. balf,ean~Lm (d'ORBIGNY) var., J.V. scaph~Lm (FrcHTEL and MOLL), N. subtU1'gid'u m (CUSHMAN), N. gmtelaupi (d'ORBIGNY), N. pampiliaides (FrcHTEL andMOLL), N. pampiliades etigaense n. subsp., and N. cf wnbilicat~d~Lm (MON1'AGU).
Of these forms, N. manptdcujiense OTURA and N. pompilioides etigaense n . subsp.
are known only as fossil. The former seems in its distribution to be confin ed to
the Plio-Pleistocene of the Pacific side of Japan, and the latter is always found inassociation with Palystamelli na disc01'binoid es Y ABE and I-IANZAWA, a characteristic Pliocene Foraminifera of Etigo, Japan. N. pompiliaides (FrCH'I'EL and MOLL)
is fairly common in the Poliocene of Kakegawa, Tot6mi and Aki.-gun, Tosa, but
apparently absent in the Japan Sea side, both fossil and recent. Other forms arewell adapted to have a geographic range as those of Elphiditm~. Compared to
Elphidi~Lm, Nanian and its allied genera are, as a rule, more abundant inslightly deeper water of thf> present seas.
-58 -On the Japanese Species of Non-ion and Its Allied Genera.

593

Nonion japoniwm n. sp.
Pl. 15(4) figs. la, b; 2a, b .
.'fest oval, bilatera lly symmetrical, composed of 10-13 'chamb ers in last formed coil;
umbilical region covered with a granular shell m atrial, fl'om which curved, limbate sutures
radiate; periphery ac ute but not keeled; surface smooth, very finely perforate; aperture
forming a n:lrl'OW curved slit at basal margin of ap ertural face. L ength up to 1 mm.;thickness ca. D.4 mm.
Holotype (Reg. No . 21422, Institute of Geology and Palaeontology, Tohoku Imperial
University) from the Pliocene of Muraoka-mum, Kamakura-gun, Kanagawn prefect ure.

This new form is one of the best know)) of the Plio-Pleistocene Foraminifera
of Japan, and has hitherto been treated by the writer as .LV. bo~wanum (d'ORBIGNY);
but it may better be separated by ·much granular umbo and almost entire marginof test, since d'ORBIGNY'S' original figure r epresents a specimen with a compressed
test, sharp lobulated in peripheral margin (Foram. Foss. Vien., 1864, p. 108, pI.
5, figs. 11, 12). CUSHMAN treats .LV. b016eam6'1n (d'ORBIGNY) as synonym.ous with
.LV. aste?·izans (FICH'.rEL and MOLL) (D. S. Nat. Mus., Bull. 104, pt. 7, 1930, p~ 6,
pI. 2, figs. 5-7).
Distribution in J a pan:
Recent: Common i n the P acific side and r are III the Japan Sea side.
Fossil: Pleistocene of Kusir o, Hokkaido j Pliocene of Sa wanc, Sndo I sla nd, andSimotaka-mati, Kariha-gun, Niigata prefecture; Atuki, Totuka-mati, and Okine,
Hatuse-mur3, Kanagawa prefecture; Sanuki-mati, Onuki-mati, and Tosaki, Tiba
prefecture; Asuka and Kamiyasiki, Taruki-mura, Hosoya, Haranotani-mura,Sugiya and Ketienzi, Nango-mUl'a, Sizuoka prefectme ; loki, Tonohama andOno, Aki-g un, Koti prefect Ul'e.

Nonion rnanp~dcujiense O'l'UKA
PI. 15(4) figs. 3, 8.

]932. Nonion 7nr.J,npuku.i'iens'is OTUKA, J our. Geol. Soc. Tokyo, vol. 39, No. 469, p. 654 (in Japa-

nese).

The present form originally dealt with by Y. 01'UKA as a distin ct species
differs from N. japo nic~6m only by having fine striae which obliquely crosss.uturesnear umbilical r egion.
Distribution in J apan:
Recent: unknown.
Fossil: Plio-Pleistocene of Naganuma, Toyota-mum, Utikosidani anti Nisinotani,
Hongo-mura; Y azu and Nozima, Ka nazawa-mati, and rrakaya, Muraoka-mura,all in K anagawa pl'efecture ; Hosoya, Haranotani-ll1ura and Sug iya Nango-mura,Sizuoka prefecture j loki, Tonohama and Ono, Aki-gUD, K6ti prefecture.

Nonian scaphum (FrcH'rEL and MOLL)
PI. 15(4) figs. 4a. b.

Compare with: 1884. Nonionina scapha BRADY, Rep. Challenger Zool., vo l. 9, p. 730, pI. 109,
figs. 14-16.
1930. Non·ion scaphum CUSHMAN, U. S. Nat . Mus., Bull. 104, ·pt. 7, p. 5, pI. 2, figs. 3, 4.
- 59-

594

K ..AsANO

Test planispiral, bila terally symmetrical,' not completely involute, composed of about 10
chambel's in last formed coil, periphery broadly l'olmded; umbilical l'egion depressed, notornamented; latel' cbambers much inflated, broadening rapidly; sutUl'es distinct, depl'essed,
not limbate; wall smootb, finely peI'forate ; apel'ture narrow at base of ap ertul'al face.

CUSHMAN treats Nonion communis (d'ORBIGNY) (For. Foss. Vien., 1864, p. 108,
pI. 5, figs. 7, 8) as synonymous with N. scaphum; REUSS, on the other hand,assigns the former to N. boueanum (Sitz, d.i<:. Ale. vViss. Wien, 1864, vol. 1, p.
479, No.5), but BRADY states that Nonion communis forms an intermediate linkalmost equally r elated to both varieties. Farther, CUSHMAN takes "Na1.Ltilus jaba "of FWH'l'EL and MOLL (Test. Micr ., 1798, p. 103, pI. 19, figs. a-c) as a synonymof N. scaplnLm, whiJe HERON-ALLEN and EARLAND say, "FrcHTEL and MOLL'S description of jaba indicates a pauperate form intermediate between Nonion andElphidium, but referable to the latter genus on account of its r eb'al processes."(Trans. Linn. Soc. L ondon, vol. 11, pt. 13, 1916, p. 281).
Dist:(ibution in Japan:
Recent: Common in the nOl't hern Jap an Sea, but l'are in the Pacific side of Japan.
Fossil: Pliocene of TUl'ihasi, and Pirika-eki, Tosibetu-mura, Setana-gun, .Hokka iqo ;Pliocene of \ ,V nkimoto, Oga Peninsula, Akita prefecture; Simotaka-lllati, Karibagun, Niigata prefect ure; Horakuzi, Konade-mul'a. Nisitonami-gun, Toyama prefecture; Plio-Pleistocene of Sanuki-mati, Kimitu-gun and l\I[oroiti, Sittu-gun,Tiba prefecture i Plio-Pleistocene of Tomioka, K anazawa-mati, Kuraki-gun, andl\I[uraol;:a-mura, Kamakura-gun, Kanagawa prefecture; Farly Holocene ofNuma, Awa-gun, Tiba prefecture,

Nonion s1.Lbturgid~Lm (CUSHMAN)
PI. 15(4) figs. 611., b.
1924. Non-ian-ina subtul'gida CUSH~[AN, Carnegie lost. Washington, Publ. 342, p. 47, pl. 16, fig. 2.
1933. Nonion subturgicltlin CUSffiIlli"", U . S. "at. Mus., Bull. 16], pt. 2, p. 43, pl. 10, figs. 4-7.

Test bilaterall y symmetrical, composed of 8-9 chambers, increasing in leng th as added,in last-formed coil; periphery rounded, somewhat edged in elll'lier chambers ; suture di stinct,n early straight, very slightly depressed; apel'tul'ul face rounded triangular, aperture narrow,at base of la.st-fo rmed chamber.

This species d iffers from N. japonicum by less number of chambers andabsence of secondary matErials at the umbilicu. '.
Distribution in J apau :Recent : Pacific side of Japan.
Fossil: Pliocen e of Wakimoto, Oga P enins1.lla, .Akita prefecture; Pliocene of Natukawad ani, Kitadyo-mura, Kariha-gun, Niigata pr efcture; Plio-Pleistocene of
Sanuki-mati, and Ititoku, Koito-mura, both in Kimitu-gun, Tiba prefecture;and of Munwka-l11ura, K amakura-gun, Kanagawa prefecture; Pliocene of
Rosoya ROl'anotani-mura, Ogasa-gun, Sizuoka prefecture.

Nonion g1·atelO1.Lpi (d'ORBIGNY)
PI. 15(4) fig. 14.

Compare wit h: 1930. Nonion gmteloupi CUSHMAN, U . S. Nat. Mus., Bull. 104, . pt. 7, .p. 10, pI. 3,. fi$s. 9-11, ; pI. 4, figs. ~-4.
~60 -On the Japanese Species of Nunian and Its Allied Genera.
1933. Nonion gmtcloup'i CUSHMAN, Ibid., Bull. 161 pt. 2, p. 43, pl. 10, figs. 8a-c.

Test small somewhat elongate, oval, planispiral, nea rly parallel-sided, rounded at periphery; chambers numerous, usu ally 10-12 in last formed coi l, chambers increasing rapidly
in length, especially in last few chambers; sutures distin ct, slightly depressed; wall smooth"finely perforate; aperture small at base of ape'r tural face.

Several specimens are found in the PI io-Pleistocen
Miura Peninsulas.

materials from Boso andDistribution in Japan:
ReceD t : Pacific.
Fossil: Pliocene of Simotaka'mati, Kariha-g un, N iigata prefecture; Plio .. Pleistoceneof lVIa ndano, lVIakita-ll1ul'a, Kimitu -gun, Tiba prefecture; and of Utikosidani,
Hong6:nnll'a, and 'rakaya, "'furaoka-mura, both in Kamakura-gun, Kanag'awa
prefecture.

Nanian bOtLean·(tm, (d'ORBIGNY) val'.
PI. 15(4) figs. 5a, b.
Compare with: 1864. Nonionina bOlteana d'ORBlGI>.'Y, For. Foss. Vien., p. 108, pI. 5, figs . 11, 12.

The present form is much compressed and nearly circular in side view, but
the peripheral margin of chambers is not so acute .as in N. bO~Le an~Lrn .
Distribution in Japan.
R ec ent: unknown.
Fossil: Pliocene of Kai garazawa, Kuromatunai-mul'a, Su ttu-gun, Hokkaido.
Pleistocene of Muraoka-mul'a, Kall1akUl'a~guu, Kanagawa prefecture.

Plio-

Nanian cf. 'umbilicatlLlurn (MONTAGU)
PI. 15(4) figs. 7a, b.
Compare with: 1884. Nonionina t,lf1bii'icatula BRADY, Rep. Voy. Challenger Zool., vol. 9, p. 726,
pI. 109, figs. 8 . 9.

BRADY says, ".LV. wnbilicatula forms a good quasi-specifi c type, embodying
characters about midway b etween those of .LV. dep?'ess'l.da (WALKER and J ACOB) and.LV. parnpiliaides (FICH'J.'EL and MOLL). From the former ' it is distinguished by its
more compact built and deeply sunK umbilici, from t.he latter by its relatively
depressed contour . and larger number of chambers."
HERON-ALLEN and EARLAND statr, "There i, ' a certain amount of variationdue (1) to the degree of turgidity in the growth of sh ell, and (2) to the degreeof envelopment of th e chambers of the final whorl. In young specimens there isno depression at the umbilicus. With increase in size if the shell continues of
the non-turgid type, the whorlR are almost entirely embracing, so that the umbilical r egi.on r emain s either almost flu sh or very slightly clepret=sed, If thechamber s are of the turgid type approaching .LV. parn.piliaides (FrcHTEL and MOLL),
each successive convolution becomes less enveloping, so that the umbilicus becomes deep." (Tran s. Linn. Soc. London, vol. 11, pt. 13; 1916, p . 297).
If the above noted statements are accepted, ',t his species is a ' cosmopolitan
form, ranging from Smith Sound, about lat. SOON., to' the south of Patagonja~- 61 ' -K . A S A~O~96

about lat. 51°8. The bathymetrical range extends from 30 or 40 fathoms· down
to 3125 fathoms.
Distribution in Japan:
R ecent: R are in Japan Sea and P acific.
Fossil: Pliocene of Sawan e, Sado Island, Niig ata prefecture ; Plio-Pleist ocene of
Sanuki.- mati, Kimitu-gun, Tiba prefecture, Muraoka-mura, Kamakura-guu, Kanag awa prefecture; Pliocene of Hosoya, H aranotani-mura, Ogasa-gun, Sizuoka
prefecture.

Nanian pampili aides (FICHTEL and MOLL)
PI. 15(4) fi gs. 13a, b.

Compare with: 1933. N on-iun pompil'io'ides GUSEIMAN, U. S. Nat . Mus., Bull. 161, pt. 2, p. 41, pI
10, fi gs. 1, 2.

The present form originally described by FrcHTEL aDd MOLL on recent speciillellS of the Mediterranean and Pliocene one of Coronica, Italy, seem. to be an
essentially deep-water bpecies. Our specimens, both fossil and recent, are quite
similar to those from the Albatross stations recorded by J. A . . CUSHMAN. H edescribes this specie. as follows :"Test planispiral, bilaterally symmetrical, entirely involute, except that the un.bilici a redeeply excavated, periphery very broadly l'ounded, entire; chambers 8 to 10 in the lastformed coil of the same general sbap e, not inflat ed; sutures flush with the surfac e, generall y limbate, fusing along the umbilicus to f orm a slight t hickening about it but n~t r aised
above the surface; wall smooth, coal'sely p erforate; aper ture slit-lik e, between the base of
the ap ertuml face and the previous coil, apertural f ace broad and low."This species d iffers from N. ~~mbilicat~~l'L~m by much thicker test and rather
fewer chambers of the last formed coil.
Distribution in J apan:
Recent: Pacific.
Fossil: Pliocene of Asuka, and Kamiyasiki, Taruki-mul'a, Og asa-g un, Sizuoka pref ecture; Pliocene of ']'onohama and Ono, Aki-gun, Koti prefecture.

Nanion pampilioides etigaense n. subsp .
PI. 15(4) figs. lOa, b.

This fo rm differs from the t ypical one wi th less number of inflat ed ch amb ers and nearlystraight su tures ; it is known only from t he Pliocen e of Etigo, where it is found always
in associat ion with Po~y stom e lli mi discO?'binoicZes YABE and ' HANZAWA, and CassicZ2blina yabei
ASANO and NAKAMURA, two chanl cteristic Pliocene Foraminifera of Japan.
Holotype (Reg . No. 21423, Institute of Geology a nd P alaeontology, Tohoku Imperial
University), fr om the Pliocene of Nat ukawadani, Kitadyo-mUl'a, Kariha-g un, iig ata prefecture, Japan.

Genus Pseudonanion ASANO, 1936
Poe2tdononion A S ANO, Trans. Pal. Soc., Japan, 1936, No. 15, p. 347.
N onionellcb (Part) CUSIUlAN, ContI'. Cushman Lab. Foram. Res., vol. 2, 1926, p. 64.

Test free, bila terally asymmetrical, dorsal side partially involute, leaving previous whorls'Uncovered, ventral side completely involute, having no elongate lobe at umbilicus; chambers-

62 -On the Japanese Species of Non'i~n and Its Allied Genera,

597

numerous, eadier ones only visible on Olle side of test; wall calcareous, finely perforate;
apel'tl11'e a nanow slit at base of apertural face,This genus was derived from a planispirally involute form, probably Nonion~
by an asymmetrical arrangement of the later chambers; already in Nonionella, adistinct, elongate lobe is developed at · the umbilicus 011 the ventral side of thetest. It is now r epresented in Japan by two species, p, japonic~£rJ~ ASANO aud P.
i1'edecum ASANO which are common in the late T ertiary and recent material ofJapan.
Pse~(, dononion japoniw?n ASANO
Pl. 15(4) figs. Ha-c.
1936. Paettdononion japonicum ASANO, Trans. Pal. Soc. J apan, No. 15, p. 347, Text-figs. a-c.
Test asymmetrical, depressed, dorsal side slightly convex with all coils visible, ven'tral
side with only last-formed coil visible, periphery subacute ; chambers distinct, 10-12 inadult, having no elongate lobe extending over umbilicus on ventral side; sutures distinct,
slightly depressed, gently curved; wall finely perforate; aperture a narrow slit at base ofapertural face .
Distribution in Japan:
R ecent: Few in the Japan Sea and rare in the Pacific,
Fossil: Pliocene of Kaigar azawa and Nakanosawa, Kuromatunai-mUJ"a, Suttu-gun,
Hokkaido; Plei~tocene of Kusiro, Hokkaicl6; Pliocene of Wakimoto, Oga
Peninsula, .Akita prefectlll'e ; Plio-Pleistocene of Sanuki-mati, Kimitu-gun, andOnuki-mati, lGmitu-gun, Tiba prefecture; Plio-Pleistocene of Okine, Hatusemura, Miura-gun, and Takaya, Mluaoka-mul'a, Kamakura-gun, Kanagawa pre-·f ecture ; Pliocene of loki and Ono, .Aki-gun, K6ti prefecture,

Pseudononion t?'edecum A SANO
1936.

P6euclonon'ion t1'edecum ASANO, J ourn. Geol. Soc. Japan, vol. 43, No. 515, p. 622, pI. 33, figs.

7a-c.
Test depressed, bilaterally asymmetl'ical, ventral side completely involute, dorsal sidesomewhat evolute, periphery subacute, umbilicus of ventral side distin ct, filled with granularshell-substance ; chamber s numerous, usually] 3-14 in last formed coil. in adult; sutures distinct, slightl y depressed: wall smooth, finely p erforate; apertlue small, at base of lastchamber.

This species, originally described by the writer from the Pliocene of Setana,
Hokkaido, differs from P. japonicum by nearly circular test with more cham ben:: ,Distribution in Japan:
Recent: Ral'e in Japan Sea and Pacific.
Fossil: Pliocene of K aiga razawa and Nakanosawa, Kuromatunai-mUl'a, Suttu-gun,
IIokkaido; Plio-Pleisto cene of Sanuki-mati, Kimitu-gun, Tiba prefecture.

Genus Nonionella CUSHMAN, 1926
Nonionella CUSHMAN, Contr. Cushman Lab. Foram. Res., vol. 2, 19 ~6, p. 64,"Test subtrochoicl, the dorsal sicle only partially involute, ventral side completely so,
close colled ; chambers especia lly in the adult inequilatel'al, the ventral side developing a
distinct elongate lobe at the umbilical end, which covers the umbilicus itself; wall calacare-

63 -K. ASANO

598

ous, fin ely p erfor a te; ape rture at the apertural fa ce of the chamber, low and elongate, extending from the peripheral bonIer toward the ventr al side." (CUSHM~N)

Nonionella ptdchella H ADA
PI. 15(4) fig3. 12a, b.

1931. Nonionella pulch dla liADA, Sc i. Rep. Tohoku I mp. Univ., Sendai, J apan, 4th ser., \' 01. 6,
No. I, p. I~O, Text-fig. 79.

Test oval, somewhat compressed, composed of about two convolutions, peri pheral marginrounded; chambers nalTOW, cm ved, increasing rapidly in size as added, all chambers inouter coi l visible, umbilical end of last cha mber gr owing in a peculiar manner to coverumbilicus in ventral side; sutures distin ct, depressed slight ly; wall smooth, som ewhat translucent or opaque, finely perforate; aperture fOJ'min g a nalTOW, arched slit.

This species is distinguish able from N . miocenica CUSHM AN by tho margin oft h e umbilical lobe of chamber, having five distinct lobelets or cr enation,'.
Distribution in J apan :Recent: Common in J apan Sea and North-Eastern Pacific side of Jap an .
Fossil: Pliocene of Kaigarazawa and Nakanosawa, Sutt u-gun, Hokkaido; Pliocene
of ULimanbe-gawa, Okunai-mura , Higasi-TugaTu-gun, Aomori prefecture; PlioPleistocene of Man dano, l\fakita-mma, Kimitu-gun, Tiba prefecture; PlioPleistocene of Okine, Hatuse-mura, :Miura-gun, and Takaya, Murao lm -muTa,
Kamakura-gun" Kanagawa pref ectUr e.

Genus. Ast?'ononion CUSHMAN aDd Ed~ua?'ds, 1937
fl st?'ononion CUSHMAN and EDWARDS, Contr. Cushman, Lab. FOl'am. Res., vo1. 13, pt. I, W37, p. 30.
N onionina and Nonion (part) of authors.

" Test hee, planispiral, coi led, bilaterally sy mmetrical or n early so, periphery bToadly
Tounded; ch ambers numerous, distinct, usually somewhat inflated, with supplementar y t ubularor rhomboid chambers on both sides, altemating with the primary ch am bers, wall calCal'eOllS
at the base of the last formed ch amber in the median li ne, a low arc hed opening, which insome species may be somewhat subdivided, the supplementary chambers with either rounded·op enings at the p eripheral end, or, in those species wi t h clistinctly rhomboid supplementarychambers, with the a perture elongate alo ng th e peripheral posterior margin." (CUSHMAl.'< andEDWARDS),

Asl?'ononion 8te ll ige?'~Lrn (d'ORBIGNY)
Pl. 15(4) figs. 9a, b.

Compare with: 1937. .Ast?·onon·ion slelligerurn CUSIH[AN and EDWARDS, Contr. Cushman, Lab .
Foram. Res., vol. 13, pt.. I, p. 31, figs. 7a, b.

This is a species comparatively .small m SIze, and hardly to be coufused with.any other.
Distribution in J apa n:
R ecent: R are in Japan Sea and Pacific.
Fossil: Pho-Pleisto cene of Takaya, Muraoka-mura, Kamakura-gun, Kanagawa prefecture; Pliocene of Natukawada ni, Kitadyo-mu l a, Kariha-gun, Niigata prefecture .
-

64 -

599

On the Japanese Species of Non'ion and Its A llied Genera.

Summary
Of the 12 forms described, 2 are known on ly as fossil, namely, Noninn rnanpttkttjiense 01'UKA and N pornpilioides etigoense n . subsp. Of the other 10
forms, N. pornpilioides (FICHTEL aDd MOLL) aDd N. subttt1·gidttrn (CUSHMAN) areinhabitants of the 'warm waters southward from Japan at the present time, while
N scaphurn (FICHTEL and MOLL), N g1'atelovpi (d'ORBIGNY), N. cf. t6'J'1'i bilicattdttrn(MON1'AGU) and Ast1'ononion stellige1'wm (d'OHBIG:s'Y) have a world wide distribution, both fossil and r ecent. N. japonicttrn n . sp. occurs commonly in the surrounding walen ; of Japan, as well in its Plio-Pleistocene materials.
Among t he fossil forms, it is noteworthy thl1t N .. pornpilioides etigoense n.
subsp. is always found in association with Polystornellina disc07'binoides YABE andHANZAWA and Cassidt6lina yabei ASANO and NAKAMURA, which are two characteristic P liocene Foraminifera of Japan. N. rnanpttk'Ltjiense 01'UKA is common in
the Plio-P leistocene of Tiba, Kanagawa, Sizuoka and K6ti prefectures.
Finally I wish to offer my warmest thanks to P rof. H. YABE of the Instituteof Geology and Palaeontology, T 6hoku I mperial University, Sendai, Japan, underwhose direction I am studying the Neogene Foraminifera faunas of Japan, forkind ly correcting and recollstructing this Dote before publication.

EI*~ Nonion .1m 1'f i L ili'ill

ctil'd ~)1'ft'£3/HI;;j'-E1E lVonion Maco'!'f=tL;f,',hK!Mil L."Cr.t, 1!lii3>:1&'fill I7') ij. L-i?>j;n fd·1.."CJi!f1':i:7»-::::>td', ~ IE!],~JUI:.'fill,

1t;:Q'filli: 1l--lt"C 12 'filli:li!l?, .lO ~ c. l:. i3' I:l~*1t o 51:> 2 'fill@P1:> N. manpukuj-iense, 1ft.. lY. pompil'io'icles etigoense f;t
1t;:Q co ij. L- i3>j;n ~ .tJ.. "C ,Q tJ:\l' o ~~1l!!. 17) 10 'fill r.t1iiJ.tJ.. 'PYI.;tE 1 "('1;. ~ "C ,Q ~ 'b co -C'<h ~, lV. subtu1'giclurn 1ft.. N.
EJ* I7.)r*1K ~ <, EJ;;jqrff,Kr.t*ti~~ ~ .tJ..tJ: \/'0 ~:9 t- co 'b cor.t , m-I!t, 1t;:QoJtK, :$-};{fi*,l!¥>"CR;:r<, "fF.,t: cosmopolitan "('<ho o ;;js:.3'(Kr.t, z~ co 'fill co EJ;;j'-Klj~~o:$-i'-;lj)l:.;f1llco Synonym K/!IJT~

pompilioicles co 2 'fillr.t,;;J§~ i:rr-::::>t:: 'b co,,(,<h ~ 0

-

65 -

66.

On the Japanese Species of Bolivina and
Its Allied Genera .
By

Kiyosi ASANO(Contribution from the Institute of Geology and Palaeontology, T6boku Imperial University,
Sendai, Japan: Received January 18th; read February 12th, 1938)

Of many different species of Bolivina and its allied genera distinguish ed up
to the present time from the Japanese material, valid are the following species,
namely:1. Bolivina hant7ceninnn BRADY
n . Bolivi n ci st?·.(a,t~lla CUSHMAM
2.
3.
4.
5.

12. Lo'Xostoma, 7cal'1'el'i(~ml1J~ (BRADY)
13. L. amygdalaefonne (BRADY)
14. Lox ostoma amygclalaefonn e io7ciens e n.

B. j'Ob?,lstCI BRADY
B. sem imlda CUSHMAN
B. hanzclwcli ASANO
B. s?,lbang~l la?'is ogasaen sis

subsp.
15. B ifa?' ina japonica n. sp.
16. R ectobolivin a bi[?'on s (BRADY)
17. B olivinita quad?'ila tem (SCHWAGER)
18. B. q ~tacll'ilatel' a C~tn eata n. subsp ,19. Bolivinella foz.£um (PARKRR and JONES)
20. Ge?nina?'icta pacifir:a n. sp ,ASANO
6. B, bmdy·i n. sp .
7. B. psmlclodijJo?'mis n. sp.
S, B . aenal'iensis (COSTA)
9. B. cilata (SEGUENZA)
10. B. sp·i nescen s CUSRMAN

I n the present paper these forms in the collection of the Iustitute of Geology
and Palaeontology, Tohoku Imperial University, Sendai, are to be r ecorded. The
material comprises the collections of Professors H , YABE, R. AOKI and S. HANZAWA, and several members of the In stitute, Messr s.
S. NOMURA, F. UEDA, H .
NIlNO and M, Nakamura, and of myse1f, made at various places in Japan. The
localities exend over the prefectures, from north to south, of Aomori, Akita, Niigata,
I sikawa, Tiba, Kanagawa, Sizuoka and K6ti,
Of the deccribed species, Bolivina hanzawai, B. subangula?'is ogasaensis, B ..
pse~tdocl~tJo?'rnis, Loxostorna arnygclalapfo?'rne iolciense, and B olivinita quacl?'ilate?Yk
C1meata are not found .in the adjacent seas of Japan, while the others are living.
As to the generic names, I have followed CUSHMAN'S work entitled, "A Monograph of the Subfami ly Virgulininae of the Foraminiferal Family Buliminidae."
Genus Bolivina d'ORBIGNY, 1839.
Bolivina d'ORBIGNY, Amer. Merid. Vol. 5, pt. 5, 1839, p. 61 (Genotype, Bol-ivina plicata d'ORBIGNY);CUSInIAN Lab. Foram. Res., Spec. Publ, No.9, 1937, p, 36.

"Test elongate, usually compressed, tapering, initial end and often whole test twisted;chambers typically biserial; wall calcareous, finely or coarsely perforate, smooth or variously-

66-

On the Japanese Species of Bolivhw and. Its Allied Genera

601

ornamented; apert ure elongate, usually oblique, somewhat loop-shaped, often with ~n internal
tube." (CUSHMAN)

According to CUSHMAN, the foraminifera of the genus, with the earliest appearance in the Jurassic, are widely distributed in the present oceans, and mostabundant in about 50-150 fms. on the continental shelves or even in somewhat
deeper water. The species as a rule are not long lived, and have a rather limited
geographic distribution.
Bolivina is considered to be derived from Vi?'g~dina by the reduction of thetriseri.al portion in early stages.
Bolivina hantlceniana BRADY
PI. 16 (5) figs. 1, 15.

1884. Bolivina lwntkeniana BJ~ADY, Rep. Voy. Challenger Zool., Vol. 9, p. 424, pI. 52, figs. 16-18.
1900. Bol'ivina hantkeniana MrLLE'l'T, JOUI'. Roy. Micr. Soc., pt. 9 p. 546, pl. 4, fig. 9.
1911, Bolivina hantkeniana CUSHMAN, U. S. Nat. Mus., Bul l. 71, pt. 2, p, 42, text-fig. 68.
1921. Bolivina hantkeniana CU SHMAN, Ibid., Bull. 100, p. 132, pI. 27, fig. 2,
1924. Bol'ivina hantkeniana CUSHMAN, Carnegie Inst, Washington, Publ. 342, p. 16, pI. 6, figs. 1,2.
1937. Bolivina hantkeniana CUSHMAN, CUSHMAN Lab. Foram. Res., Spec. Publ. No.9, p. 127, pI.
16, figs. 18- 20.

The species, widely distributed in the Pacific, also occurs in the Plio-Pleistocene deposits of Japan, though lacking in those of the Japan Sea side, both fossill
and recent. Ther e are two varieties, test very broad with a distinct, transparent
keel about the entire periphery in one variety and narrow without keel or withthat g'reatly reduced in the other. Both forms are found together, having in common the characteristic costae on the test .
. Distribution in Japan:
R ecent : Pacific.
Fossil : Plio-Pleistocene of Tiba, Kallagawa, and Sizuoka prefecture:;;.

Bolivina ?'ObtLsta BRADY
PI. 16 (5) figs . 5a, b .
. 1884, Bolivina 1'obusia BRADY, Rep. Voy. Challenger Zool., Vol. 9, p. 421, pI. 53, figs. 7-9.
1893. Bolivina 1'obusta EGGER, Abh. kon. bay. Akad. W·iss, Miinchen, Cl. II, Vol. 18, p. 294, pI. 8,
figs. 31, 32.
1911. Bolivina 1'obusta CUSHMAN, U. S. Nat. Mus., Bull. 71, pt. 2, p. 36, text-figs. 59, 60, .
1931. Bolivina 1'Obusta HADA, Sci. Rep. T6hoku Imp. Univ. Sendai, Japan, Ser. 4, Vol. 6, p, 131,t ex t~fig. 88,
1937. Boliv-ina 1'obt!sta CUSHMA.c~, CUSHMAN Lab. Foram. Res., Spec, Publ. No. 9, p. 131, pI. 17, figs.
.
1-4.

There are many records for this species ranging from the Eocene to recent.
Distribution in Japan:
Recent: P acific, and Japan Sea.
Fossil: Plio-Pleistocen e of Tiba, Kanagawa and Sizuoka prefectures.
-

67 -

602

K. As,rno

Bolivina seminuda CUSHMAN
PI. 16 (5) fig. 10:1911. Bol'i vina serninucla CusEmAN. U. S. Nat. Mus., Bull. 71, pt. 2, p. 34, text-fig. 55.
1927. Bol-iv·ina serninnda CUSHMA,,'<', Scripps In st. Oceanogr. Tech. Ser., VOl . 1, p. 157, pl. 3, fig. 6,
1931. Bolivina sern'imula HAllA, Sci. Rep. T6hoku Imp. Univ. Sendai, J apan, Ser. 4, Vol. 6, p. 132,

text-fig. 89.
1937.

Bolivina~erninncla

CUSHMAN, CUSHMAN Lab. Foram. Res. Spec. Publ., No.9, p. 142. pl. 18,

figs. 13, 14.

CUSHMAN states, "The species ranges southward alo ng the Pacific coa, t as far
at least as Panama, and westward across the Pacific to Japan, where it is recorded
by HADA from shallow water of Hokkaido aDd Mutsu Bay."The species occurs commonly in the Plio-Pleistocene of California aDd Japan.
Distribution in Japan:
Recent : J pan Sea and Pacifi c.
Fossil: Pliocene of Akita and Nii gata prefectlll'es aud Plio-Plestoeene of Tiba andKanagawa prefectures.

Bolivina hanzawai ASA"OPI. 16 (5 ) figs. 8, 13a, b.
Bbl-ivincb hanZCbWCbi ASANO, Jap, JOUl'. Geo1. Geogr., Vol. 13, No. 3- 4. p. 330. pI. 37 . figs, 9a, b.
1937. Bolivina hanzawai CUSHMAN, CUSHMAN Lab, Foram. Res., Spec. Publ. No. 9, p . Ill, pI. 13,
fi g. 1.

1936.

Test broad, co mpressed, apex bluntly pointed, edges except at aperture surrounded by athin wing-like flange; chambers almost flatte ned; sutUl'es distinct; 2- 3 short longit udinalcostae at earlier portion of test; wall thin, with mthcl' coal'se pel'forations; apertul'e o\'al,terminal.

T he types are from the P liocene of Asuka, Taruki-mura, Ogam-gun, Sizuoka
prefecture. The species differs from B. hantlceniana BRADY by the few costae on
earlier portion of test .
Distribution in Japan:
R ecent: unknown.
Fossil : P liocene of Sizuoka and K6ti pl'efeGtures,

Bolivina subang1~la?'is ogasaensis ASANO• PI. 16 (5) fig. 16.
1936.

Bol'ivina subangulm'is ogasaensis ASANO, Jap. Jour. Geol. Geogr., Vol. 13, Nos. 3-4, p. 334, pI.

37, fig. 8.

1937. Bol'ivina subangulal'is ogasaensis CUSH~fAN, CUSHMAN Lab. Foram. Rel , Spec. publ. No.9, p . .
114, pl. 13, fig. 2.

This subspeeies differs from the typical subang1~la?'is BRADY in the ornamentation of the test whieh consists of two prominent and never irregularly curved
costae.
The types are from th e P liocene of Hosoya, Haranotani-mura, OgaEa-gun,Sizuoka prefecture.
- 68-

On the J apanese Species of Bolivina and Its Allied Genera

603

Distribut ion in J a p an :R ecent: un known.
Fossil: Pliocene of Sizuoka prefecture.

Bolivina bmclyin. sp.
Pl. ]6 (5 ) fig3. 2, 14.
1884.

Boliv'i na beyrich'i BMDY, Rep, Voy. Challenger Zool. Vol. 9, p, 422, pl. 53, fig' 1.

Test elongate, a bout 4 time as long as broad, r at her narrow, much compressed, apicalend bluntly pointed, apertural end evenly rounded; chambers numerous, increasing rapidly
in hei g ht as added, each distinctly prolonged backward to a point project ing at outer edge;
sutures distinct, oblique or gent ly curved; wall smooth, punctuate; ape:t ure elongate, terminal. L ength up 0.8 mm.
Holotype (R eg. No. 21424) from the Soya-maru statio n 352, west coast of Kii peninsula,
JD 154 meter s.

The present form, once r eferred to B. beY?'ichi REUSS by BRADY, may be distingui .. hed from the latter by the shape of chambers and by more elongate test.
According to CUSHMAN, B. veY?'ichi REUSS occurs at num erous localit ies in Central
Europe in the middle Oligocene, but the specimens from later and earlier T ertiary
formations and also the R ecent specimens refen ed to the same species are decidedly
distinct from it.
Distribution in J apan :Recen t : P acitic .
Fossil: . Pliocene of Taya, Kawabe-gun , Aki ta prefecture, and P lio-Pleistocene of
Kanagawa prefecture.

Bolivinct pseuclocl~lJo?'mis n . sp.
PI. ] 6 (5) figs. 4a, b,
Test similar to that of B. cl·i ffonnis I ,V ILI,JAMSO N (R ec. F ormn . Gt. Britain, 1858, p, 6,::figs. 166, 167), but much more elongated and nal'ro,,'er test .
Distribution in Japan:
R ecent: unknown .
Fossil: Pliocene of T iba prefecture.
Holotype (R eg. No. 21425) from the ·Pliocene of Sanuki-mat i; Kimitu-g un Tiba prefecture.

B olivinct alata (SEGUENZA)
1884.

Bol'i1lina beyj'icl!'i REUSS val', alata BRADY, Rep. Voy. Challenger Zool., Vol. 9, p, 422, pI. 53,

figs, 2-4,
1910. Bol·i?;·ina be./}j·ichi REUSS val'. a/(Lta SIDEBOTl'O~f, Mem, Proc. Manchester, Lit. Phi los. Soc"
Vol. 54, pt. 3, p, 13.
1937, ' Bolivina alat(L CUSHMAN, CUSFIMAN Lab. FO\'aill, Res., Special Publ., No.9, p. 106, pl. 13,
figs, 3-11.

Several specimens referrable to SEGUENZA'S alata were found in th e recen tmaterial from the Pacific side of Japan.
Distribution in Japan:-

69-

K . AsA...'<o604

Recen t : Fair ly common 1ll the P acific side of J a pan, but absent in th e Japan S ea_
Fossil: unknown.

Bolivina spinescens CUSHMAN
Pl. 16 (5) fig. 5.

B olil1ina textilarioides BRADY (not REUSS), Rep. Voy. CballengeJ' Zool., Vol. 9, p. 419, pl. 52~
fi gs. 24, 25.
1900. B olivin a textilal-1oides MILLETT, J om . Roy. Micr. Soc., p. 542, pI. 4, fi g. 5.
1911. B ol'ivina spinescens CUSBMAN, U . S. Nat. Mus .. Bull. 71, pt. 2, p. 46.
1932. B ol'iv'ina spinescens HERON-ALLEN & EARLAND, Discovery Re p., Vol. 4, p. 354, pI. 9, figs. 7,8.
1934. B olivina spinescens E.mLAND, Ibid., Vol. 10, p. 132, pl. 6, fi gs. 8-10.
1937. Bol'iv-ina spinescens C USB ~IAN, CUSHMAN Lab. F oram, Res., Special Publ. No, 9, p. 142, pI. 18,
fi gs. 17-19.

1884.

The present species is widely di stributed in deep water of the Pacific. CUSHMAN already recorded it from the southern coast of Japan an d from the area between Yokohama and Guam. Recently H ERON - A LLEN a nd EARLAND record it from
off th e Falkland I slands.
Distribu t i on in J a p a n:
Re cent ; Pacific.
Fossil; unkn'o wn .

B olivina st1'iatula C USHMAN
PI. 16 (5) fig. 18.
Bolivina stl-iatula CUSIHIAN, Carnegie Inst.it. Washingt on, Pub!. 311, p. 27, pl. 3, fig. 10.
B ol'iv'ina stl'iatula CUSHMAN, U . S. Nat . Mus. Bull. 104, pt. 3, p. 43.
B ol·iv ina stl'iatula CUSIIMAN and PARKER, Ibid. Proc., Vol. 80, Art, 3, p. 16, pl. 3, figs, 21a, b.,
B olivina st1'iatula COLE, Florida St. Geol. SunT., Bull. 6, p, 41, pI. 2, fig. 9.
B olivi17a st1'iatula CUSHMAN, CUSHMAN "Lab. Foram. Res. Special Publ. TO. 9, p. 154, pI. 18"
fi gs. 30, 31.

1922.
1922.
1931.
1931.
1937.

CUSHM AN' states that the 'pecies IS fairly in the 'Western Indian r egion southward to the coast of Brazil. The occurrence of this species in the V\Testern Pacific:
I S noticeable in its distribution.

Distribution in J a p a n:
R ecen t : Pacific sid e of J apan.
Fossil: unknown.

B olivina aena1'iensis (COS1'A)
PI. 16 (5) fi g, 21.
Compare with: 1937. B olivina aenwiensis CUS AMAN, CUSHMAN La b. FOl'am. R es" Special Publ. No. 9"
p. 105, pl. 12, fi gs, 21-26.

Several specimens referrable to the present species were obtained from thePacific side of Japan:
Distribution in J ap a u :Recent: P acific.
Fossil: unknown.
-70-

On the J apanese Species of Bol'ivina and Its Allied Genera

605

Genus L oxostoma EHRENBERG, 1854
Loxostomttm EmmNBERG, Mikro. Geo1., 1854, p1. 17, fig. 1!J (Genotype, Loxostoma SUbl'ostratttm

EHRENBERG ).
LO.'Costoma HOWE, Jour. Pal.. Vol. 4, 1930, p. 329 ; OUSEI~[AN, OusmIAN Lab. Foram. Res., Special

Publ. No.9, 1937, p . .168.
Boliv'i na (part) of authors.

Test in the earlier stages similar to Bolivina, in adult tending to become uniserial withterminal aperture.

This genus with its early memb er s, appearing in tho upper Cretaceous, continues to the present ocean s; it, greatest development seems to be in the present
tropical waters at moderate depth.
Loxostoma amygdalaejol'1ne (BRADY)
PI. 16 (5) fig. 7.
1884. BoUvinn amygcZalapjol'mis BRADY, Rep. Voy. Ohallenger Zool. Vol. 9, p. 426, p1. 53, figs. 28,29.
1911. Bolivhw amygcZalaejol"1nis OusmlAN, U . S. Nat. Mus., Bull . 71, pt. 2, p. 42, text-figs. 69a, b.
1921. BoUvi.n a amygcZalaejol-rr1:is OUSm(AN, Ibid., Bull. 100, p. 133. pI. 26, fig. 3.
1937. Luxostomet arnygdala~fol'me OUSHMAN, OUSHMAN Lab. Foram Res., Spec ial Publ. No.9, p .- 183.

pI. 21, figs. 21-23.

The types are from a Challenger statio n off the Philippines in 95 fathoms.
There are also many records of thif:i species from the late Tertiary and recent
material of the Indo-Pacific region .
Distribution in Japan:
Recent: Pacific side of Japan.
Fossil: unknown.

Loxostoma lca?Te?'ianum (BRADY)
PI. 16 (5) figs. Ga, b; l~a, b.
1884.
1893.

Bolivina kW'/'el'ietna BRADY, Rep. Voy. Ohallenger Zool.. Vol. 9, p. 424, pI. 53, figs. 19-21.
Bolivina karre'''iana EGG ER, Abh. kOL1. bay. Akad., Wiss ., Mbnchen, 01. II, Vol. 18, p. 299,

p1. 8, figs. 38, 39.
1911. Bol·ivina l.·ct1'1·el·iana OUSHMAN, U. S. Nat . Mm., Bull. 71, pt. 2, p. 40, text-fig. 65.
1921. Bol'ivina kWTe"iana OUSII~IAN, Ibid., Bull. 100, p. 131, pI. 26, fig. 4.
1937. Loxostoma lcCtl'j'~I-ial'ltLm CusmIAN, Ous mIAN, Lab. Foram. Res., Spec. Pub1., No.9, p. 184, pI.

21, fig. 17.

The types are , from the Challenger station 232, in 345 fms., south of Japan.
The species is widely distributed in the Pacific in fairly deep water. It is mostabundant off southern Japan and off New Zealand.
Distribution in Japan:
Recent: Pacific and W estel'l1 Japan Sea, bu t doubtful in til e Northem Japan Sea.
Fossil : Plio-Pleistocene of Tiba, Kanagawa, and Sizuoka prefectures.

Loxostoma amygclalaejo?'me 'iolciense n. subsp.
PI. 16 (5) figs. 3a, b.
The present fossil differs from the typical amygcl~laefo?'me by the prominent raised cos-

71-

606

K. ASANO

tae whi ch ornament the whole t est .
H oloty p e (Reg. No. 21426) f rom the Plio cene of loki, Aki-g un, K6 ti p r efecture.

•

OUSHMAN records L . am yg dalaefo?'me (BRADY) from the Albatross stations D
4875, 'Korea Strait, in 59 fms., D L1964, off southeastern Japan , in 37 fm s., and D
4883, Eastern Sea of J apan, in 53 fms. The present subspecies has not yet beenfound in th e adj acent seas of Japan .
Genus Bi/ a'l'ina PARKER and J ONES, 1872
Bija1' in a P ARKER & J ONES, Ann. Mag. Nat. R ist., sel". 4, Vol. 10, 1872. p. 198 (Genotype, D imo1ph'i na
saxipam E mmN BERG); CUSEIMAN, Cusml AN Lab. Foram. Res., Special Publ. No.9, 1937, p. 196
D im01"p h'ina EEIRENBERG , 1854,"Test with the earlier chamb ers biserial, later ones uniseria l, uniserial por tion making
up most of t he t es t in mos t s pecies ; waH calcareous, p erf ora te : aperture in t h e young as
i n B oliv ina. later t erm inal an d ro unded." ' ( CUS H ~UN )According to O USH~f AN , the genus appearing fi rst in the Jurassic, found its
well developped f'pecies in th e Eocene, b ut the best characterized species ar e th e
recent ones of the . I ndo-Pacific.
Bifm'·i na japonica n. sp.
Pl. 16 (5) fig. 12.
Test elongate, early port ion biser ia l ; somewhat compressed, later portion uniserial, n earlyrounded in t ransve rse section; chambe rs distin ct, excavated on low er side, often with a broadcollar-like proj ecti on in a band about m iddle portion of un iserial chamber s ; sutmes strongly
depressed ; waH smooth, excep t for raised band on ea ch cha mber; aperture ter mi nal, elliptical with a disti nc t lip . Len g th ca. 1 mm.
Holotyp e (Reg . N o. 21444 ) from the S 6yo-maru station 347, Kii Channel, in 126 f a thoms .

Th e present form more or less resembles B. fimb?'iata (MILLET1'), but may beeasily dd inguish ed from it by the differen t characters of chambers.
. Distribution in J apan .:Recent: P a cific side of Japan.
F ossil : u nkno wn .

Genus R ectobolivina OUSIIlIIAN, 1927
R ectobolivina CUSEIMAN, COotI". CUSEIMAN Lab. Foram. Res., Vol. 3, 1927, p. 68. (Genotype, Sagrinab'ij1'ons BRADY)
Sag1'ina (par t) and S iphogene1'ina (part) of anthors.

Test elon gate, somewha t compr essed, ear ly chambers bisel'i aI, later unis erial ; wall calcare ous, p ei'for atc ; ap erture t erminal, round ed with a slight lip .

Some of th e forms have been placed in Sip hogene?'ina, and th e generIC position of som e of the species is confusing.
R ectobolivina bif?'ons (BRADY)
P I. 16 (5) figs . lla, b .
1884.

Sag1'ina biJ1'ons BRADY, Rep. Voy. Challenger Zool., Vol. 9, p. 582 . pI. 75, figs. 18-20.

-72 -On the Ja panese Spel'ies of Bolivina and Its Allied Genera
1893.

607

SiphogenerincL b'ijj·ons EGGER, Abh. kon. bay. Akad. Wiss., Miillchen, CJ. II, VoL, 18 p. 317,

pI. 9, figs. 25, 26, 29.
1913. Siphogenej'ina bi/j'ons CUSIH!AN, U. S. Jat. Mus., Bull. 71, pt. 3, p.105, pl. 45, figs. 1, 2, 5-7.
1921. Siphogenerinct b~rj'ons CUSIH:!AN, Ibid., Bull. 100, p. 277, pI. 36, figs. 2, 3.
1926. Siphogene j'inct b'ijj'ons CUSIH!AN, Ibid., Froc., Vol. 67, art. 25, p. 16, pI. 3, figs. 7-9 ; pJ. 4, fig. 4.
19~7. Rectobol'i vina b'i jj'ons CUSm!AN, Conk CUSHMAN Lab. Foram. Res., Vol. 3, p. 68.
1937. Rectoboliv'incL b'i/1'Ons CUSIHIAN, Ibid., Spec. PubJ., N. 9, p. 204, pI. 23, figs. I3, 14.

This species is best developed in the r eg IOn from Japan !Southward to thePhilippin~s and Malay Archipelago.

Distribution in Japan :Recent: Pacific unknown in the Japan Sea.
Fossil : Plio-Pleistocene of Tiba and Sizuoka prefectures.

Genus Boli'Vinella CUSHMAN, 1927
Rol'ivinella CUSHMAN, ContI'. CUSfDIAN, Lab. Foram. Res. Vol. 2, pt. 4, 1927, p. 79 (Genotype, Tex tulm'icL foliurn PARKER and JONES.)"Test much compressed, the pl'oloculum in the megalospheric form rectangular, in the
microspheric for111 the young i s apparently planospiral, later chambers bisel'ial, chambers longand recurved, not overlapping; wall calcal eo us, perforate; aperture tr ansverse to the compression of the test with numerous papillae at t h e base of the opening." (CUSllllU,N)Thi;:; . genus occurs in the Eocene and continues to the present sea, especiallyof the Indo-Pacific.

Bolivinella foli'um (PARKER and JONES)Pl. 16 (5) fig. 20.
1884. Te~:tulm'icL foliurn BRADY, Rep. Voy. Challenger Zool., Vol. 9, p. 357, pl. 42, figs. 1-5.
1936. Bolivinf'lla "zegans ASANO, Jour. Geo1. Soc. Japan. Vol. 43, No. 515, p. 619, pI. 33, fig. 3.

There are considerable variations lD this species.
similar to BRADY'S figure 3.

The present form is very

Distribution in Japan:
Recent : Soyo Maru station 345, off Mmotozaki, Sikoku, in 199 meters.
Fossil: Pliocene of Nakanosawa, Kuromatunai-mura , Suttu-gun, Hokkaido,

Genus Bolivinita CUSHMAN, 1927
Bolivinita CUSJUfAN, Contr. CUSHMAN Lab. Forum. Res., Vol. 2, pt. 4, 1927, p. 90 (Genot,ype, Textulm'ia quadj'ilatera SCHW,IG ER.)"T est with chambers biserial, periphery and broader sides all concave with strongly
developed angl es giving' a qnadrate end view to th e test; wall calcareous, perforate; aperturelarge at base of inner margin in the median line." (CUSHMAN)

The lJl.embers of this genus seem to be common in shallow, warm waters of
the present Ol:ean s.

Bolivinita quad1'ilate1'a (ScmVAGER)
Pl. 16 (5) figs. 17, 18.

1866.

Te.'l'tv.laj·ia quadj"ilatl'1'a SCHWAGER, Novar a Exp. Geol. Theil., Vol. 2, p. 253, pl. 7, fig. 10.

-73-

608
1884-. Tex tuZm'ia guad'riZatem BRADY,. Rep. Voy. Challenger Zool., yol. 9, p. 358, pl. 42, figs. 8-12
1897. Te.'ctuZm··ia guacl?'ilatem F LINT, U. S.. Nat. Mus., Misc. Rep., p. 283. pI. 28, fig. 3.
1911. Tex tuZm'ia guad?'ilatei'a CUSH~[AN, I bid., Bull. 71, pt. 2, p. 24, text-figs. 42-44.

The present form seems to be wid ely distributed in the northwestern Pacific .
The types are from the Pliocene of Kar Nikobar . .
Distribution in J apan :Recent : Pacific; unknown in J apan Sea .
Fossil: Pleistocene of Tibll, Kanagawa, Sizuoka and Koti prefectures . .

Bolivinita q'l.6ad1·ilatem ctmeata n. subsp.
PI. 16 (5) figs. 19a,. b.
Test nearly wedge-shaped, similar to that of B . qu acZ?'-iZatem (SCHWA.GER), but differingfrom it by curved ch ambers which overlap the predecessors.
Holotype (Reg . No. 21427) from the Plio-Pleistocene of Muraoka-mura, Kamakura-gun,
Kanagawa prefecture,
Distribution iuJapan:
R ecent: unknown.
Fossil: Plio-Pleistocen e of Kan agawa prefecture.

Genus G~mina?'ic ta Cusm1AN, 1936
Gem;naricta CUSlnlAN, CusH~rAN Lab. Foram. Res., Special Publ. No.9, p. 61.
BoZ'ivinella CGSIDItIN (part), Cont)'. CUSHMAN Lab. Foram. Res., Vol. 5, 1929, p. 33.

"'fest in tb e early stages biserial, compressed, in the a dult uniserial; waU calcar eous,perforate ; apert ur e in the adult consisting of a small rounclecl openin gs well separatecl fromone another towarcl t he encls of an elongat e, elliptical depress ion in the terminal wall."(CUSHMAN)The genus is a specialized one and has been confined to the Miocene of thesouthern France. But CUSHMAN says, "It is possible th at this genus may be foundin the I ndo-Pacific or Australian region.', as other forms of the Miocene of F rance
show this range."

Gemina1'icta pacifica n. sp.
PI. 16 (5) fi gs. 22a, b.
Test nearly triangular, earl y portion biserial, ancl tapering, later portion uniseria l, sides
n early parallel, periph ery in eady portio n acute, later rounclecl ; chambers distinct , not inflatecl; sutlll'es dis tinctly raised, very slightly curved; wall l'oug henecl; aperLure terminal,
elongate, with t wo sman openings at both ends. L ength up to 0.5 mm .
Holoty pe (Reg . No. 21445) from the Soyil-Maru station 578, Toy am~ Bay, in 187 fat homs.
:rhis is tbe first recorcl of the recent form of t his genus.
Distribution in Jap an:
Recent: 'l'oyama Bay, J apan Sea.
F ossil: un known.

Finally I wish to offer my warmest thanks to Prof. I-I. Y ABE of the Instituteof .Geology and Palaeontology, Tohoku Imperial Un iversity, Sendai, Japan, uuder- 74 -

609

On the Japanese Species of Bol'ivina and Its Allied Genera-whose direction I am studying the Neogene Foraminifera faunas of Japan , fork indly correcting and reconstructing this note befor e publication.

Bol'ivi?w ;&;!tll[(PJ,)%c7..)1'f:fU,':Jilz rMd L-Lr.t. J'&ll[ CUSHMAN c7..) monographic study j)rib ~ , TifJilZ ~ .l: <?<I1Gtt~ K.?e -:::> Pc. 0

EI;,j;:E/ic7..)"4, c7..) K f£tIt' L Zli:: ftJIJ 'l!E-li- L- K, 20:fillli:: iffi!t5lU-t-:o c c. j)r /1:1 *Pc.

0

;It~c7..) r.f:,'

Bolivinct han-:zawai, B.. subangulal'is ogasaensis, B, pseudod{ffo1'rnis, Loxostoma amygdaiaejoTl1w 'iokiense, Bolivin-ita quadTilatem cuneala r.t EI;,j;:c7..)*2JS.fjoallj c7..)f!ttJi;li-llt (7.)"'n> G1JlUiL ~ tt-, ;lt11!! c7..) JU!..7Jn>G r.t, ffl:llt, 1t-1ioltK?<11 Gtt- -c J,;,

11 It, C c. r.ti1/3 -t--< ~"1:ib:O0f,l-j'1'I Ge??~ina?'icta Mjc7..)1'HLi!?,r.t , '1iOI~~:1:IHc7..) Miocene c7..)", K liR G tt- L J,;, Pc. "4, c7..)"1:ib:O j)r, 1'f!I~jL:J!R~c7..)~Ii1:llt;f;~;,j;:1'f fL£}'c7..) 5 i> K , *M} c7..) "4, c7..) c.:3/5~ G~c:O 1 Ji;If:fillli::!iifJiL L-tc 0 ;,j;:3t.K r.t Z ~c7..)~J{(:fillK)jj,tIt'Lc7..):3/5*li::~-<tc."4, c7..)"1:ib:O o

67.

On the Japanese Species of Uvigerinaand Its Allied Genera
By

Kiyosi ASANO(Contribution from the Institute of- Geology and Palaeontology, Tohoku Imperi al University,
Sendai. Japan : . Received January 18th, 1937; read February 12th. 1938)

In the collection of fossi l foraminifera from Japrn of the Institute of geology.and Palaeontology, there are numerous specimens of the three genera, Uvige?'ina,
AngulogM'ina and T?'ija?'ina, their localities are;
1. vVakimoto, Oga Pe'n insula, Akita prefecture; Pliocene; ColI. K. ASANO.
2· Taya, Iwamisannai-mUI'a, Kawabe-gun, ditto; Pliocene; ColI. K . ASANO .
3. Sade Island, Niigata prefecture; P liocene; ColI. Y . OZAWA.
4. Natukawa-dani, Kitadyo-mul'a, Kariha-gun, ditto; Pliocene; ColI. K . HANZAWA.
5. Tagawa, Konado-mul'a, Nisi-tonami-g un, Toyama pl'efectl1l'e; Pliocene; ColI. K .
ASANO.
6. Sema tanoseki, Sito-mlll'a, Jtihara-gu n, Tiba p r efect ure; Pleistocene; Coll. ]VI. NAKAMURA.

7.. Sanuki-mati, Kimitu-g un, ditto; P liocene; ColI. F. DEDA.
S. Kamo, Toyato-mura, Awa-gun, ditto; P liocene; Coli. F. DEDA.
9. Hiroseitiba, Kok ubunzi-mlll'a, ditto; Pliocene; ColI. F . DEDA.
-75 -K. ASANO

610

10. Nisi-Nagat a, 'l 'oyohusa-mura, di tto; Pliocene ; ColI. F. UEDA.
11. T akaya, Tomioka-mul'a, K amakura -gun, Kanagawa prefedure; P lio-Pleist ocen e;.
ColI. F. UEM.
12. Yazu, Kanazawa-mati, Kuraki-gun, ditto; Plio cene; Ccll. M . NAKAMURA..
13. Xozima, ditto, Pliocen e; ColI. M . NAKAMURA..
14. Osikiri, Hongo-mul"a, Kamakura-g un, dit to ; Plio-Pleistocene; ColI. M . NAKAlIiUR.tI..
15· Naganuma, Toyota-mul"a, ditto; Plio-P l eisto cen e; CoIl. F. UEDA.
16. Asuka, Taruki-mura, Ogasa-gun, Sizuoka pref ecture; Pliocene; ColI. R . AOKI.
17. Kamiyasiki, ditto; P iocene; ColI. R. AOKI.
18. Hosoya, Haranotani-mul"a, ditto; Pliocen e; Col I. R. AOKI.
19. Sugiya, Na n go-m u ra, ditto; Co lI. R. AOKI.
20.

Keiienzi, ditt o; ColI. R. AOKI.

Most of the8e fossils belong to r ecent species, but several are not known fromthe adjacent seas of Japan.
Ther e are 12 species of Uvige?'ina and its alli ed genera in Japan, namely :Geological range in Japan
Pliocen e-Pleist ocene-Recen t

Uv-ige?'-ina pygmaea d'ORBIGNY
U.
cf. b if~w cata d'ORBIG1'<--YU.
t enUl:st?'-iata REUSS.
U.
m edit e?Tane££ HOFKER
U.
schwage?'i BRADYU.
aw leata d'ORBIGNY
U.
cf. pe"eg?'-ina CUSHlILA..l.'T
U.
yabe·i n. sp .
s~£b st?'-iat(£ n. sp .
U.

U.

ps e~£doa?np~£llacca

"

"
"
.,

"
P liocene""
Recentn. sp.

Anguloge?'ina japonicc£ .n. sp.
l'dfa?'ina b'r aclyi CUSHMAN""

Of these forms, U. yabei, U. st6bst?'iata and U. cf. pe?'eg?'ina are not yet known
living, while U. medite?'?'anea, U. tenuist?'iata and U. schwage?'i are fairl y common
in the Plio-Pleistocene depodits of Tiba, Kanagawa and Sizuoka prefectures, but
apparently absent in those of the J apan Sea side. Other forms a re widely distributedin Japan, both fossil and recent.
As a rule, Uvig eTina and its allied genera are abu ndant ill fairly deep waterof the present seas.
Before going into the descriptions of the species, t he writer wishes to offer
his warmest thank' to Prof. H . YABE of the Institute of Geology and Palaeontology,
T ohoku Imper ial Univer sity, Sendai, Japan, under whm,e supervision th e present
work was carried out, for kind suggestions and correction of this ar ticle before
publication; he is indebted to those who kindly submitted their specimens to him
for study.
Genus Uvige?'ina d'ORBIGNY, 1826
UO'i ge?'ina d ' ORBIGNY, Ann. Sci. Nat .. vol. 7 , 1826, p. 269. (G enotype, U. pigmafa cl' ORBlGNY)
Uhlig'ina SCHUBER'.r, Sitz deut. naturw. med . V el'. Bohmen, 1890, p. 222.

-

76 -

On the Japanese Species of Uvige9'ina and Its Allied Genera

611'"Test generally triserial, elongate, fusiform, rounded in transverse section; chamb ers
distinct and inflated; wall calcareous. perforate; apertlll'e terminal, rounded, with a neckand phi aline lip. oftell with a spiral tooth and an intel'l1al twisted tube." (CTJSHM:AN)

Uvige1'ina seems to have been derived from Uvige?'inella with a virguline
aperture. The m embers of this genn s are, in general, specifically distinguishedfrom one another by the difference in the form and arrangement of chambers andin the ornamentation of wall. Although some . pecies of Uvige'l 'ina, have short
ge010gic ran ges aud arc good horizon makers, most of the species are difficult todistingui sh, and too rare in occurrence to be of much stratigraphic value.
Uvig e1'incL pygmaea d'ORBIGNY
Compare with: Uv ige9'ina pygmaea CUSHMAN, On Uvi:le9"ina p ygmaea d'OIWIG NY, ContI'. Cushmnn.
Lab. Foram. Res ., vol. 6, pt. 3, 1930, p. 62, pl. 9, figs. 14-20.

Variou s form s of costate Uvigerin ae have been referred to this species bymany anthors. In 1927, OUSHMAN visited the type locality, OOl'oncina, near Siena,.
Italy and collected abundant material. According to his study, the test of thisspecies is fairly small and elongate, with the width of initial end differing muchin the microspheric and megalospheric form,~ . All but the last one or two chambers have rather sharp longitudinal costae, usually discontinuous over sutures ~costae are replaced by numerou, ' short spines on the last one or two chamber s.
The apertural end has a distinct neck, elongate and slender, and in well preserved specimens thers is a distinct lip. The ~pecimens at my dispoEal from the
Plio-Pleistocene of Japan, as shown on Plate 17 (6), figs . 5-7 varys considerably inthe shape of chambers, but otherwise are quite similar to d'ORBIGNY', ' pygmaea,
illustrated by OUSHMAN.
Distribution ill J apau :Recent: Pacific and J a pan Sea ..
Fossil: Pliocene of Akita, Niigata, and Toyama prefectures (1, 2, 4, 5); Plio-Pleistocene of Tiba prefecture (6, 7).

Uvige?'ina d. bif~wcata d'ORBIGNY
PI. 17 (6) fi gs. 3, 4.
Compare with: 1839. Uvige9'inCL biftt9'cata d'ORBTGNY, Foram. Amer. Merid ., p. 53, pI. 7, fig. 17.
1858. Uvige9'ina pygmaea WILLIAMSON, R pc. Foram. Gt. ·Britain, p. 66, pI. 5, figs. 138, 139.
1884. Uvig(winct pygmam BRADY (part), Rep. Voy. Challenger Zool., vol. 9, p. 575, pI. 74, figs. 13,14.

d'Or:BIGNY distiIlguish es U. bijw'cata frOll U. pygrna ea by the arran gementof costae, which" au lieu de se continuer sur tout I'hauteur de chaque loge, sout interrompups, 01.1 bifurquees, caractere constant chez tous les individus." But "VILLIAMSON and BRADY Eay that the difference is not sufficient to consititute a specific distinction. OUSHMAN, emphasizes that BRADY'S figures of U. pygrnaea d'ORBIGNY in the Ohallenger Report are of two distinct species. The writer considers
that BRADY'S figures 13, and 14 on Plate 74, may be separated from the others
-77-

{l12

K. As.mo(Figs. 11, 12) by the well-defined raised costae and more slender and elongate test.
These forms are perhaps referrable to d'ORBIGNY'S biftwcata, but au accurate
identi fi cation is impo sible without a study of the original specimen.
Distribution in J apan:
Recent : J apan Sea a nd Pacific.
Fossil: Pliocene of Akita, and Niigata prefectmes (1,3,4) ; Plio-Pleistocene of Tiba
and Kanagawa pref ectures (7, 10, 12, 13, 15).

.Uvi.ge?·ina m edite?'1'anea HOFKER
Pl. 17 (6) figs. 8, 10, 11, 14, 15.
U!32.

Uvigerina mediterranea HOFKER, Publ. Staz. Zool. Napoli, vol. 12, Fasc. 1, p. 118, text-fig. 32.

The present form, once r eferred by many authors t.o U. pygmaea d'ORBIGNY,
was fir st separated by HOFJ<ER from the typical ' d'ORBIGNY'S pygmaea. According
to HOFKER there are three forms in this species; one is larger than the two others,
which are transitionally variable to one another, and generally slender , taperingto the initial end. These three forms explained to be due to " Trimorphism" areTespectively named as B, Al and A2 forms.
Distribution in J apan :Recent: Pacific.
Fossil: Plio-Pleistocene of 'l'iba a nd Kanagawa prefectmes (7; 10, 11, 14, 15) and
Pliocene of S izuoka pref ecture (17, 20)

Uvige?'ina tenuist?'iata REUSS
Pl. 17 (6) figs. 12, 13.
Uvigel'in a tem tistl'iatct BRADK, Rep. Voy. Ch allenger Zool, vol 9, p. !574, pl. 74, figs . 4- 7.
1893. Uvigel'ina tenwistl'iata EGGER, Abh. kon. bay. Akad. ··Wiss .. C1. II, vol. 18, p. 315, pI. 9, figs
44, 52.
1913. UV'igel'ina,tr.nuist7·iata CUSHMAN, U. S. Nat. Mus., Bull. 71, pt. 3, p. 95, pI. 42, fig. 4.
1921. Uvigcl'ina temtistl'iata CUSIDLlN, Ibid., Bull. 100, p. 269, pl. 55, fig. 2.
1926. Uvigel'ina tenu.ist7·iata CRAPMAN, New Zealand. Geol. Sury., Pal. Bull. II, p. 70, pI. 14, fig. 9.
1884.

Test elongate, subcylindl'ical, tapering to apical en'd, comppsed of numerous ch a mb ers{)rnamen ted wit h numerous fine, lon gitudinal costae; sutures dist in ct; apertme with a ShOl·ttubular n eck and a distinct phialine lip .

The species may easily be distinguishable from U. m edite?'1'anea HOFKER by'much elongate test with very fiuely striated ornamentation.
Distribution in J ap a n:
R ecent: P acific, and Wakasa Bay, Japan Sea.
Fossil: Plio-Pleistocene of Takaya, Muraoka-mul'a, Kamakura-gun, Kanagawa prefecture.

Uvige?' ina schwage?'i BRADY
PI. 17 (6) figs. 19, 20, 24, 30.
1884. Uvigel'ina schwagel'i BHADY, Rep. Voy. Ch'l.llenger Zool., vol. 9, p. 575, pI. 74, fig. 8-10.

-

78 -

On the Japanese Species of Uvigel'ina and Its Allied Genera
1913.
1921.

613:

Uvigel"ina schwagel'i CUSJnIAN, U. S. Nat. Mus., Bull. 71, pt. 3, p. 97, pI. 37, figs. 3, 4.
Uvigel'ina schwageri CUSHMAN, Ibid., Bull. 100, p. 270, pI. 55, figs. 3-5.

The species originally described by BRADY from the Philippines is very common in the r egion. CUSHMAN Eays " Apparently the region south of J apan , where·so many species reach their northern limitE, is too cold for this species, for it was
not noted there." The species is also absent in our collection of recent Foraminifera from the adjacent seas of Japan, but fairly common in the Plio-Pleistocene·of Tiba and Sizuoka prefectures.
Distribution in Japan:
R ecent: unknown.
Fossi l: Pliocene of Tiba and Sizuoka prefectures (8, .9, 10, 18, 19).

Uvige?'ina aC1deata d'ORBIGNY
Pl. 17 (6) fig. 23.
1846.
1884.
1913.
1921.

Uvigel'ina aculeata d'ORBIGNY, Foram. Fos~. Vienne, p. 191, pI. n, figs. 27, 28.
Uvigel'ina aculeata BRADY, Rep. Voy. Challenger Zool., vol. 9, p. 578, pI. 75, figs. 1, 2.
Uv·igel·ina aC1.lleata CUSHMAN, U. S. Nat. Mus., Bull. 71, pt. 3, p. 100, pI. 43, fig. 4.
Uvigerina aculeat(£ CUSHMAN, Ibid ., Bull. 100, p. 273, pI. 55, fig. 6.

Next to UvigM'ina schwage?'i BRADY this is a COmmon species in the Philip-.
pine regions. CUSHMAN records it from the Albatross Station D 4957, off J apan,_in 437 fathom s.
Distribution in Japan:
Recent: Pacific.
Fossil: Plioc ene of Tiba prefecture (8, 9, 10).

[}vige?'ina pse1Ldoamp1Lllacea n. sp.
PI. 17(6) figs. 28, 29.
Compare with: 1913 UV'igel-ina ampul/acea CUSHMAN (not BRADY) U. S. Nat. Mus., Bull. 71, pt ..
3, p. 102, pI. 42, figs. 3n , b.
Te5t elongate, composed ' of a more or less compact spiral of chambers in early portion·
followed by two 01' more chambers uniserially al'l'anged; wall finely hispid; aperture with
an elon gate neck and pbialine lip. Length up to 0.8 mm.
Holotype (Reg. No. 21428) from the Soyo-Maru station 345, near Muroto-zaki, Siko.ku, .
in 199 meters. Paratypes are from the Soyo-Maru station 297, near Tanegasima in 516·
meters.
Distribution in J apal1 :Recent: Pacific alon g tbe coast of Southwestern Jap an.
Fossil: unknown .

UvigM'ina yabei n. sp.
PI. 17(6) figs. 1, 2.
T est elongate, subcylindrical, tapering at apical end, composed of numerous cbamberswith several distinct longitudinal costae except last formed-chambers; aperture with a short:n eck and distinct phialine lip. Lengtb up to 1.5111m.
-

79 -

'614

K. ASANO

Holotype (Reg . No . 21429) from fhe P liocene of Wakimoto, Oga Peninsulu, Akita prefecture.

The present form is similar to U. tenuist1'icLta REUSS, but differs in less number of costae. Furthermore the last few chambers are not ornamented with costae.
Distl'ibution i n Japan:
Recent: unknown.
Fossil: Pliocene Wa kimoto sandy shale of Oga P eninsula, Aki ta pref ecture.

Uvige1'i na subst1"iata 11 . sp.
Pl. 17(6) figs. 21. 22.
Test subcylindri cal, comp osed of 5 or 6 whorl s ; ch amb er s num erous, inflated; sutures·distinct, gently curved, depressed; wall striated only neal' sutures ; aperture at end of ashori. neck and sUITounded by a phialine lip. Length up to 1.8 mm.
Holot ype (Il,eg. N o. 21430) f rom the Plio-Pleisto cen e of Sanuki-mati, Kimit u-gull, Tiba
pref ecture.

This new species IS very distinct in sutures crossed by numerous fin e striae.

Uvige1'incL cf. pe1'eg1'ina CUSHMAN
Pl. ]7(6) fi g. 18.
Compare wit·b: 1923. Uvige?'ina pe?'eg?'ina CUSIIMAN, U. S. Nat. Mus., Bull. 104, pt. 4, p. 166, pI.
42. figs. 7-10.
]927. Uvige·rina p e?'eg?'inCL G ALLOWAY and VVISELER, J our. P al., vol. ], No.1, p. 76, pl. 12, figs. 1,2.

The specimen s are very close to CUSHMAN'S p e?'eg1'ina in short, high plate-like-costae, ,vhich ar e granula r ou surface, spinose or brocken near apel'tu;ral andinitial ends. But the last few chambers ar e considerably larger in the present
fossil which is wedge-shaped in general outline of test.
Distribution in Japan:
R ecent: doubtful.
Fossil : Pliocen e W akimoto sandy shale of Oga P eninsula, Akita p r efecture.

Uvige?'ina sp.
P I. 17(6) fig. 16.

Test very rmall, f usifo rm, composed of about 3 whorls; chambers few, very sligbtly.inflated; sutures distinct, depr e~s ed; wall smooth, no t ornamented; aperture terminal witha thickened lip and short neck.

The form differs from U. ca17CL1'iensis 'ORBIGNY by less number of chamber s-which are leES inflated.
It is ,poEsible that this is merely a young stage of smooth surface Uvigerinae,.lSuch as U. cana1'iensi.~ d'ORBIGNY or U. jCL1'inosa HAN'l.'KITN.
Genus Anguloge1"ina CUSSM AN, 1927
.Angulogo'Tina CusmrA.'T.
J. Contr. CUSHMAN L ab. Foram. Res., vol. 3, 1927. p. 69 . (Genotype, Uviger'ina,
angnlosa WILLrAMSON)
Ut-igerinn (pm·t) of authors.

"Test h iseria J, elongate, t h e whole test angled, with three flatten ed sides and disti n ct
- 80 -

On the J apallese Species of Uvigel'ina and Its Allied Genera.

615

angles; wall calcareous,. perforate; aperture at the end of a short neck, with a phialinelip." (CUSHlIf.AN)
Angt~loge?'ina is a specialized form not far removed from Uvige?'ina, in which,besides being trihedral, the later chambers tend to become uniserial.

Anguloge?'ina japonica n. sp.
PI. 17(6) fig. 17.

Test generally trian gular in section, both ends sharpl y pointed, the angles with sharp
carinae; chambers few, sometimes indistinct only slightly inflated; sutures indistinct, slightly
depressed; wall ornamented with several longitudinal costae; apertme terminal, with ashort neck and distinct lip. Length up to 0.8 mm.

The present species differs from A. angulosa (WILLIAMSON) by the sharply
pointed apical end and distinctly cari~ated edges.
H olotype (Reg. No . 21431) from Urasimasyo, "Vakasa Bay, J apan Sea, in 190 meters.
Distribution in ,Japan:
Recent: Japan Sea.
Fossil: Pliocene of Ketienzi, Nango-mura, Ogasa-gun, Sizuoka pl'efectme.

Angtdoge1'ina sp.
Pl. 17( 6) fig. 26.

In this form chamber,' are very indistinct due to promi.nent costae; apicalend is sharply pointed and edges are strongly carinated.
Distl'ibution in Japan:
Recent: Soyo-rnaru sta tion 352, "Vest coas ' of Kii P eninsula, ll1 154 maters.
Fossil: unknown.

Genus T1'ija1'ina CUSHMAN 1923
TrijCtl"ina CGSIHIAN, U. S. Nat. lYIus., Bull. 104, ]923, pt. 4, p. 99 (Genotype, T l'ifw'ina bmdyi

CUSHMAN).
Rlwbdogonium BRADY (not REUSS) Rep. Voy. Challenger Zool., vol. 9, 188·1, p. 524.
Tl"ipla,soia CusmfAN (not REUSS ) U. S. Nat. lYIus., Bull. 71. 1913, pt . 3, p. 62.

"Test elongate, triangular in transverse section j the early chambers in an il'l'egularspil'al, later ones very loosely so or even uniserial; " 'aU t hin, translucent, finely punctate;
aperture terminal n ot J'adiute, at t he end of a shOl't, often phialine lip." (CUSHlIfAN)

T1'ija1'ina is a homeomorph of Siphogene1'ina, although it is not known tohave a siphon.
'l'1-ija1'ina bmclyi CUSHMAN
1923.

Tl'ifar'inCt bmcly·i CusmlAN, U. S. Nat. Mus., Bllli. 104, pt. 4, p. 99, pI. 22, fig. 3-9.

There are numerous records for this species, both Pacific and Atlantic.
Distribution in Japan:
Recent : Soyo-ma l'U station 485, off the coast of S'imune prefecture, Japan Sea
93 meters.
Fossil: doubt.ful.

-

81 -

lU

616

. 13

*

~ Uvig c?"ina,m ~ f L !GL

Oi'i:i~)EPfs:&lf1t ;fi3ltK:m;tE U1igelina, Angulogel'ina, Tl'ifmina 3 ~f7.):till, 12 K%\: It'-C 5{f~~~ L ko ;f*EFIl~~Jlll;4',Q;f7.)Etf~·I!t1f=IL i!1f7.)- "? ,:j@t~ Q t f7.)"(';b Q 7.1',

U. yabei n . sp. ]{('(Fm~4'&bf7.) U. wbstr'i ata n . sp . f'j::Kkf7.):it!!.7JKlj~ It ,-C ,~1i!! f7.) 10 :ell f'j:, ±I!!W(Jl]lk lftl!! 'iitrll])-]-;(fj' 7.I,1tIt(1I] rrHi[f'j:z. L..1t'oExplanation of Plate 14 (3)Figs. la, b.
Fig. 2.
Figs. 3a, b .
Figs. 4a, b.
Figs. 5a. b.
Figs. 6a, b.
Figs. 7a, b.
Figs. Sa, b.
Figs. 9a, b ; lOa, b .
Fig. 11.

Elphid-I:um cl'ispum (LINNE) x 25.
Elphid'ium kusil'oense n . sp. x 60.
E lphid'i um advenum ( CUSETM AN ) x 25.
Elphidium subgmnulosurn n. sp. x 50.
Elplvidiurn jenseni ( CUSH~[AN) x 25.
EI!Jhidium cmticulatwn ( F I C HTE L an d MOLL) x 20.
E lphidium cf. jabum ( F I CETT E L ·an d. MOLL) x 25.
ElphirJ)iella nagao'i n . sp. x 20.
Elphidittm yabei n . sp·. x 25.
Elphicl-ium yezoense ASANO, (Photo.) x 5.

E xplanation of Plate 15 (4)Figs. l a, b; 2a, b. Non'ion japoniCttm n. sp. x 25.
Figs. 3, 8.
.
Non'ion manpukujiensc O'£UKA x25.
Figs. 4a, b.
Nonion scap lmm (FICEITEL an d M OLL) x 30.
F igs. 5a, b .
Non'ion boueanum (d'ORBlGNY) var . x 25.
Figs. 6a., b
Nunion sttbtm'gidum (CUSHM AN) x 30.
Figs. 7a, b .
Non'ion cf. umbilicottdum (MoNTAGu) x 30.
Figs. 9a, b.
Astrononion stelligel'um (d'ORBIG NY) x 30.
Figs. lOa, b .
N onion pompilioides etigoense n . subsp. x 30.
Figs. II a-c.
Pseudononion japonicurnAsANo x 40.
Figs, 12a, b.
N on'ionella pulchella H ADA x 50.
Figs. 13a, b.
Nonion pomp il'ioidc8 (FICWI.'EL and MO LL ) x 25. .
Fig. 14
Nonion grateloupi (d' ORIlIGNY) x 30.

Explanation of Plate 16 (5)Figs. 1, 15.

Fig. 2.
Figs. 3a, b.
Figs. 4a, b.
Figs. 5a, b.
Figs. 6a., b.

Bolivina hantkeniana B RADY
F ig. 1. R ecen t, off MUl'otozaki, Sikoku. x 40.
F ig. 15. Pliocen e of l oki, Aki-gan, K 6ti prefecture. ,;, 30.
B ol·ivina bl'adyi n . sp.
R ecent, west coas t of Kii P eninsul a. x 40.
Loxostoma amygdalaejol'rne iok'ir~nse n. subsp.
Pliocene of l oki. Aki-gun, K 6ti prefecture. x 35.
Boli'Oina pseudocl'ijfo1'1nis n. s p.
Pliocene of Sanuki-mati. K imit u-gun, Tiba p r efecture. x 40.
Bol'ivina robusta B RADY
Pliocene of H osoya. H a ranotani-m ura, Ogasa-gun. Si~uoklt. prefe cture.
L oxostoma 7,el'l'ej'ianurn (BRADY)
-

82-

*

<, 11."? &lf lfl t ~ <f7.) t f7.)f'j: 'flii:'P' -r: ;b !) , M1i'L

x 45.

Vol. XLV, P l. 14 (3)V o 1. XLV, PI 15 (4)Vol. XLV, pi ~ 16 (5)Vo l. XLV, P l. 17 (6)617

Fig. 7
Figs. 8, 13a, b.

Fig. 9.
Fig. 10.
Figs. lla, b.
Fig. 12.
Fig. 14
Fig. 16.
Fig. 17.
Fig. ]8.
Figs. 19a, b.
Fig. 20.
Figs. 21a, b.

Plioce ne of Muraoka-mura, KamaInna-gun, K anagawa prefecture. 35.
x 40.
Recent. T osa Bay, Si];:oku. 40.
B olivina hanzawai A SA NO
Fig. 8. Pliocene of loki, AId -gun, K6ti prefecture. x 30.
Fig. 13. Pliocene of Hosoya, H aran ot.a ni-m ura, Sizuoka prefectUre. x 30.
Bol'ivina spinescens CUSHMAN
B ecent, Eastern J apan. x 40.
B oliv'ina seminucla C US HMAN
Pliocene of Muraoka-mura , Kam akura-gun. Kanagawa prefecLure. x40.
R ectobol'i vina b'! frons (BR,\DY)
Pliocene of Hosoya, H aranoi;ani-mUJ'a., Ogasa-gun, Sizlloka pr efect.llre. x 30.
Bym'ina japonica n. sp.
Recent, Kii Chann eL x 40.
B ol-iv'i na aena?'iensis (COSTA)
Recent" Kii Channel. x 30.
B ol-ivina .~u bangulaj·is ogasaensis A SANO
Pliocene of Hosoya, Hara notani-mura, Ogasa-gun. Sizlloka prefectUre. x 25 ..
B olivin'ita quaclj'ilatel'cL (ScFlWAGlm)
Recent, off Murot-ozaki, Sikoku. x 45.
Boliv'i na stj'iatula CusmBN
Recent, Eastern J apan. x 35.
Bol'ivin'ita quaclj"ilcLtem cuneata n . subsp.
Pli ocen e .of Muraolm-m ura, K amakm a-gun. Kanagawa prefecture. x 45"
B ol-ivinella folium (PARKER anr! J ONES)
R ecent, off Murotoza.ki, Sikoku. x 50.
Gem'in aricta paciJica n . sp.
Recen t , Toyama Bay, J apan Sea. x 40.

L O.Tostoma arnygdalaejonne (BRADY)

Explallation of Plate 17 (6)Figs. 1, 2.

Uvigcj'ina yabei n. sp.

Figs. 3, 4, 9.

(?) UvigerincL cf . biJtwcata d'ORBIGNY

Figs. 5, 6, 7.

(?) Uvigerina pygrnaecL cl'OmnG NY

Wakimoto, Oga Peninsula, Ak ita prefecture.

x 20.

vV akirnoto, Oga Peninsula, Akita prefecture. 20.
'\Vakiriloto, Oga Pen in sul a, Akita prefecture. x20.
Figs. 8,10,11,14,15. Uvigej'ina rned'it~j'ranea HOFKER
Fig. 8. Muraoka-rnura. K:magawa prefecture x20.
Figs. 10, 11. Osikiri , Hongo-mura, K anagawa prefecture. x 20.
Figs. 14, 15. K etienz i, Nang6-mura, Sizuoka prefec ture. 20.
F jgs. 12, 13.
Uvigej'~n a temL'istj"iata REUSS.
Muraoka-mura, Kanagawa prefecture. 20.
Fig. 16.
UVI:gej'ina sp.
Wakasa Bay, Japan Qea, 190 m eters. x20.
Fig. 17.
Angulogej"ina japonica D. sp.
Wakasa Bay, J apan Sea. 190 meter s. x30.
Fig. ]8.
Uvigej'ina .cf. pej:egrincL CUSElMAN.
W akimoto, Oga Peninsula, Akita prefect ure. 20.
Figs. 19; 20, 24, 30. UV'i gerina schwageri Bl~i\DY
Figs. 19, 20. H osoya, Haranotani-m ura, Sizuoka prefecture. xl/O.
Figs. 24, 30. Hiroseitiba, Kol. . ubunzi-mura, Tiba prefecture. x20.
Figs. 21, 22.
Uvigej·in a. substj·jata n. sp.
-

83 -

618

Fig. 23.
Fig. 25.
Fig. 26.
Fig. 27.
Figs. 28, 29.

Fig. 20. Sanuk i-mati, R imitu-gun, Tiba prefecture. x 20.
Fig. 22. R amo, Toyota-mara, Awa-gull, Tiba prefecture. x 20.
Uvige'/'ina actdeata d ' O HBIGl\"Y
Hiroseitiba, Kokubunzi-mura, 'l'iba prefecture. x 20.
T1"ijm'ina bmdyi C USHMAN
Soyo-mam station 485, Japan Sea, 93 meters. x 20.
Anguloge1"ina sp.
Soya-mara station 3~ 2, west coast of K ii Peninsul a, 154 meters. x 30.
Uvige1'inct sp.
D it t o. x 40.
Uviye1'ina pseucloampullacea n . sp.
F ig. 28. Soyo-mara stat.ion 345, neal' Muroto-zaki, Silw ku, 119 m. x 40.
Fig. 29. Soyo-mal'u station 297, neal' Tallegasima, 516 meters. x 40.

68 .

(I)

1111

(IItlfll 13 ~p. 2 f-J 12 El i;'j1'Hlil, 2 f-J 12 El §l:3illD1111 ~rlffQUL1iHtT 7.Jirr~:=~:DJi~~~I:'iL ilJ 7i IJ ~i:I'L .ft P"ti!if'£lJ1c ~T 7.J:J;U!.m-tJ"ib 7.J 0 ;!{ - Vi ~ff~tJ"
i'-c ¥lX15- L-k 1) 'B:vk~%'B:9£:;rt~~~ O) s;fi~:j·~J':fI\ffi [i7;J fX. 7.J ~~0) S ;fi '@;-.ft15;j:rRiJ'Wm-'C· , 1: ;fL J: IJ ~1*.f*1'f.ai;.-} fX. 7.J {t;fiti!if'£IJ 1c ~ ~ 1'L1*~ L-k o J1.{;j~O) s ;fi,@;-.ft P"ti!if'£lJJi,l: fj'I] [I-~ft· U'l!. G;fL7.J'@;-{t;fi;j:ffi[if'£lJJiI'L ,'B 9t\l%'B:vk!~*t!~:j'~iEt±, ]l lAfj'I]*'<j' s t!$, fj'I] \l%45 Jr:l:.;rtl\f)( f*~:j'1mt!]l lA fj'I]~:j' S t&IIIf.~tJ"ib 7.J 0~rf~ Vi~~ltifu-"E.4:~:I:O) :J2t JljJ I'L J: -::::>-c :&~0)~:f;U!.};.i.J' G1*~ L-kMflf 1c~3Jll L--c 37 Jm 52 f:j[ (~ 1*~!ff:O H~ ,':I:\ i- 7.> 1: ,l: tJ" .'±\*ko (~0)9'*TVi~~I:t*.4:~±tJ"Qi: <~MH I'L~c.~ L- -C ~*T 7.J 1:,l: I'L fX. -::::>-C -=5 )J1.{;~ 1 * 9'B * ;MiM~~1cO) . FOI'L:ffl.1::.L-7iv>bO)tJ" 7 ~G{'J 19%),13 flIt G:q 259-6)WJ ~G;fL 7.J 0 ]I! I'L JI.{;.ft;fiti!if'£IJ~~ O) ~:J:i!!. ~fJ Qi:7i 7.J *~·lnR:l tJU!.::15I'L:ffl..4: L-7i V> lb 0) tJ" 3 )~ ( Ci n n a momum,
l 'e tmclenia" Ficus). 3 flIt ( F icus sp ., C innam o m~tm camp h om , l 'e t?'acZ enict gla~t ca) ib 7.J O) 'C'iJU~,l:,g. -tt-C 10 ~ (*'-1 27%), 16 flIt U{~J 30% ) l't1lttJ"*~'lnR~tJU!.::15I'L Jl!. G;fLfX. v> bO) 'C·ib7.J o~ B *~lt,~I'L ~ vt 7.J~7m<flIto):; i?,'*L i1'ioclenclj'on honsyuensis E ND04J

Glyptost?'obus eU?'op aeU3 HEEP'2)
Sequoia

semperv i'r en~ E NDTJ3 ) .

SctSBajm s ya bei E NDO et OKU'1' SU 5)

mif~fJ£:IT;t : !irr~1t ".J1t::r:iti([~ (.wilJrj jy;mo. P. 20, 1931.
E NDO, S. an d OKu'r su, H. : Glyptost1'obus Cone fr om t he L i?'ioclen cl?'on Bed near Sellda i. Proc.
I mp. Acad . Tokyo. XII, pp. 138-140, Figs. 1-3, 1936.
3) ENDO, S. : A ~eoge lle Species of Sequoia from J apan. Bot. Gazette. XCI V. pp. 605-610, Figs.
1-13, 1933 ; New F ossil Species of Sequ o'ia fr om tbe F ar-E ast. Proc. Imp. Acad. 'J'okyo, X II, pp. 172175, F igs. 1- 13, 1936.
4) E NDO, S. : D iscovery of L i1'ioclencl1'on Leaves from t be Neogen e of J a pan. Proc. I mp. Aead.
Tokyo, X , pp. 590- 593, Figs. 1, 2, 1934.
5) E NDO, S. and OKUT SU, H . : A Neogene Species of Stusa/ras from J a pa n. Proe. I mp. Acad.
Tokyo, XII, pp. 47-49, Figs. 1- 4 . 1936.

1)

2)

-

84 -

619

V) 4 ,®) 4 {llil'Ct*'" -e vt~I'C¥R{5- LkiJ, G;!t1l!!.V) t V) I'C--:::J'" -e P L O!!;~ -eJj!,k '" c!:. ,1[I, ~~0Taxodittm vt~ t illi}I~5f:1j 1JQ V)*',11\:&, V::;) '1'- C/ ::1 I'CV\!.~-t J., t V) "c1t::r5 c!:. L -e vtiJ}t <~t ~~:t;;RV)jf~.=mc.
liiJ, G¥R{5-~ l/'L, 1It~1 1'C~-e t r!ljlg.=mc.V)?I(~'ZIiI'C~ <.ffi-t J., t V)c'ib J., iJ>:Jirr:1$.=mc.f@}.ffi c!:. Vc vt
1III ~rltJm:.V)1iliI'CW$j5, S) II IIH~iJ' G1*ff-'&~}1t t V)iJ>:tR3t ~ l/'L-c m- J., 0
Catalpa vt~tilli7¥:5f:1j 1m*=m\:&.lJ:':3t~Jlll'CfJi.~-t J.,~ c' ib J., iJ>::JJtil I'C~-c vtV\!.;ffL@*c!:. L -cWi <~~~ l/'L-C m- J., I'C -t ~'Ji.
1Ili;grltJm:.iJ, GI±\k1t::r5 vtx1~\m~{lli1'C [j'iJ5l:': ~ t1k~
Fi?'mianc! platanif olia vt iff~I'C :!J¥,~-t J., t V)c,1Itil~:bU!.I'CL@* c!:. L -c~:l:fi ~ l/'L -C m- J.,lft t 31,ffml
V)W*c'ib J., 0 1t::r5 c!:. L -C vt 4-lBJiJ>:lftt;7JV)iW.{5-c' ib J., 0
Ilex C01'n2tta vt!j;tPrfJi.%V)~~lc~-t J., t v)T1ItiJ{jjt11'I'I,ifb:&.v::x1}Bl'CfJi.~T J.,lfl'!.'*ib J.,{llic', '=*3JI1.~±:MZ 1c ~]!Il!Wt\V) r:1)j::r5~ftJ1fiJ' GM; C6 -C iW.-#:r L k Oli) 1IIli.HfJm:.vt;!tjf:t; 2 V)1t::r5.ffi:l:-I1!. C' ib J., 0 Llct1'
tnmcatttm :&. v:: A ce1' f 1'ancheti vt :J~: I'C x~}BV\!.~{llic' ~ F agtl.s f e1'1'ttginea vt:1t illi~~5f: Ij:fJn V) !llfl5'IUU!.
151'C~T-iTJ., Fagt!s c!me?'iccma 1'Cm:.1b1,c'ibJ.,0
PopttlttS bctlsctmoides c!:. Fic'l!S 1'ttminianct c!:. vt:J~I'C I&\ff.lt E V) J:f:tl¥JT-litiJ' G~c.~~ ~ l/'Lk1t::r5{llic'iW::f
vtl8\V) 93 l¥JT-litl'CfrU., 'Mfmll'C~T J., iJ>:, 1JtilTvt '8:9'&;!Wt\·(fr JU1l\:l t~gl\tt*iIm.l ~ ~Jj!, ~ l/'Lk ~ V) c!:..lIt
1111~rlftm:..ffic!:.-g--lt"-C 2 fi!lffifrLiJ>9:r1 Gt1-Cm-1t"',JLtm:.1b1,{llivt:ltilli~~5f:1j1m V)~t1f\lc!:. y", 1) 7:bU!.151'CV\!.
~-tJ., Pop1!l'lis balsamifera L, c'ibJ.,o FiCt!s 1't!1ninianCl vt Oswald HEER ~~iJ>: Schweiz V)"'0OS1nunda sp_" Glyptostrubus eUl'opaeus H EE R
" Sequoia sernpe1'v'il'ens ENDL," Taxocl-itt1n clist'ichmn RICU,
Taxv.s cusp-ielata S, et Z ,Tht/ja japon'iea MEXlM,
Acel' cl-iabolicwn BL,
A, pictum TUUNB,
A , 1'!lfine1've S, et Z.
A, eupal-rn.atwn {(omz.
t A, t1'uncat1.l1n Bunget A , jmnchet'i PAX.
A. s'ieboldianwn MIQ.
Alnus fi1'ma S. et Z. val'. s'ieboldiana WINKLE..-I,
mnxlmow·icz# CALL.
A,
h'i r;ttta TURcz.
Betula japonica 5mB.
B.
max-imo7lI'icz-iana REGEL
B,
ermann·i CUAM.
Castanea c'rena~a BL.
" Catalpa Ol'ata G, DON.
c,·lt-is sp.
CletMa bal'bine7'vis S. et Z.
Cinnamomum carnphom NEES, et EnERM .
Fagus sp.
cl'enata BI..
F.

t Fagus jer7'ttgines An.
t Ficus l'umin'i ana HEERF.
sp.
" Fil'1n'i ana platan'!fo1ia SCHOTT et ENDL," L-i7'iorlend1'on honsyuens'is ENDO
t llex c07'nuta LINDLEY et PAXTON
Juglans sp.
Hydrangea sp,
Kalopanax l'icinijoliwn MIQ.
lofeliosma sp.
lofagnolia sp.
Ostl'?}a japon'ica SARG .
Pints sinensis L,
t Popnius balsamo'ides GAEPP.
Pnmus ssiori FR. SCUoIl .
Quel'cus sp.
Q.
cr·is}.uln. BL.
Q,
glauca TUU NB," 8assafl·as yauo:i ENDO et OKUTSU
Sap'iu1n japonicum PAX et K . HOFF)I ,Stewa7·t'ia pseudocamell-ia MAXIM.
T etmclenic£ glauca MA'l'SU~I.
Ulmus .iaponica SARG _'Vitis sp,
Vibumu1n fUl'catum BL,
Zell.;owa sel'mta MAKINO

Table I . List of the Fossil Flora found in the Nenosiraishi P lant bed etc.
{* (Genera) t (Species,) ex tinct in J apanese Island, except Ta iwan.}6) MIKI, S,: ·Plant fossils from the Stegoclc,n Beels a n d the Elephas Beds near Akashi. Jap,
Jour. Bot. T okyo, V III, pp,3::!O- 322, pI. IX, B. D, Fig. 7, F-I-I, 1937,
-85-

620

i1:l~

m.?fcl(q:,~-tI!:)j}> GMI,,\)L~Cn-~ L-kM7)T@ ~ 0?cKJit;{~O)s/Eitt/Eift[tjMiltO):flfr~T ~:J;l!!.J;t~tt:a::=!3Cf ~ K:&~*1-t:SftlIf£IJm;j,-~bf~*K W(f,tt'"t n~ -n{l~*m:&V:'W[ I I::I 1r-~ft:smO):til!.1!t~tt ~Jlt*1t15ft[f£IJmf~ltO)T-K 1lL-T ~f£~ft/EiftlItJMJl;,Hlt
EEft[f£IJl.-~;;&v:'ilJtJM1lt ~ iJ~tlfr~-T ~:J;l!!.1!t~Mt ~ :a::,g.-1t:?1~ L,

J:$q:,miit :a::tlfr~T ~ t 0) ~ 5i!, G;fL~

On Fossil Plants from the Environs of Sendai (I).
(Resume)By

Seido ENDO
In the present article is treated a Neogene flora of the Nenosiraisi plant bed. The plant bed is
one of the prolific plant bearing deposits hitherto found in the environs of Sendai, its type locality
being Nenosiraisi, about 12 kln. NW. of th e City; the plant bed seems to occupy a considerable areaaround the type locality .
The material were collected by Mr. H . Okutsu and the writer during the several years pastfrom the type horizon and from some others thought to be almost contemporaneous with it.
The plant species di scriminated of the material are listed . The fl ora consists of 52 species in 37
genera, and has 7 genera (about 19.% of the whoJe) ann 13 species (about 25.% of the whole) exotic
ToO the present flora of the islands of Japan proper. Some of them, however, are now living in
Taiwan. Fmthermore. 10 genera (about 27.% of the whole) and 16 species (30.% of the whole) are no
more living in northern Honsyu. The plant bed is Upper Miocene in age, judged on the mammalianand molluscan remain s found in the underlying Siogama and overlying Tatunokuti group.

7) HEER, 0.: Die tertiare Flora de Schweiz. III, p. 183, pI. CLII, Figs. 11, 12, 1859.
-86 -

69.

N eogene Shells from the Vicinity of the Cityof Takasaki, Gumma-ken, Japan
By
Tuneteru OINOMIKADO(Read February 12f;h; received March 26th, 1938)

At the 6 tho Ordinary Meeting of the Palaeontological Society of Japan (1937),
the occurrence of Neogene mollusks from the Tomioka-Simonita distriet, Gummaken was repo rted by Mr. K . SUZUKI. In 1938, Dr. I-IUZfll10TO and Mr. KOBAYASr1 )described the geology of the district, and according to them the geological sequenceof the Neogene deposits is as follows (in descending order).
1. Akima bod: Oonglomerate, agglomerate, and lava flow.
2. Itahana bed: Alternation of sandstone, sh ale and conglomerate; lignite
beds containing pla nt leaves at the upper.
3. Tomioka bed: Alternation of sandstone and sha le, and conglomerate ..
4. Usibuse bed: Sandstone.
5. K anoh ara bed: Oonglomerate.
Dr. I-IuZIlI10TO and Mr. KOBAYASI collected L epidocyclina a nd lVliogyp ~ina fromthe Tomioka bed and many mollusks from the Italuina bed. They cancluded thatthe age of the Tomioka bed is referred to Miocene, and the Itahana bed, which
unconformably covers the T omioka bed, is Pliocene in age.
Th e writer collected some interesting species of Moll usca from the Itahanab ed. The distinguished forms are as follows:1

2

3

PELECYPODA
Nuculana con/usa (ILI NLE Y) ..... .
e Anadam o.qawai lYI.~KIYAMA . . . . ..••• . ..................•••..........• • .
L imopsis spo
e Glycyme1'is k-suzuki·i n. sp . ... .
lJfytillls spo...
e Chlarnys kanehami (YOKO YA~fA) ..
o.
o.
Chlarnys sp. .
0

0

0

0

0

1)

•

0

•

0

0

•

0

0

••••

•

0

•••

0

0

0

•

0

0

•

0

0

•

0

0

•

0

••

0

••

•

0

0

0

0

•

•

•

0

•

0

HUZlMOTO and KOBAYASI:

•

•••••

0

••••••••

0

•

0

•

0

•

••

•

•••

•

0

0

0

0

0

0

0

0

0

••

0

0

••

0

•

0

•

0

••

•

0

•

0

•

•

•••••

•

0

0

0

•

0

0

0

0

0

0

0

••••

••

0

0

•••

0

•

0

0

0

••••

0

0

0

0

0

0

0

0

0

0

0

0

0

••

0

0

0

0

0

0

•••

0

•

"

•

0

•

•

•

•

0

•

•••

0

••

0

0

•

0

0

0

0

•

•

•

••

0

•

0

0

0

0

0

•

•

0

•

•

••

•

••

•

•

••

•

•

•

•

0

0

0

0

•

•

••

••

0

',

•••••

••

•

•

0

0

•

•

•

0

•

•

R

F

R
R
F
R
F
R

Jour. Geol. Soc. Japan, Vol. 45, No. 533, pp. 205-226. 1938.
-

87 -

R

672

T. OINOMIKADO

1

2

3

~~-

Crassatellites suyamensis n. sp. . .. . ........ .. .......... .. .. . . . . . .... . ... .
Can/ita (.ilfio dont·iscus) nakamur.ai anna7;aensis n. subsp. . ... . . .. .... . .. .
Venericard'ia sp . ...... . ... . ... ... . . . .... . ..... .. . . . . .. . .... .... . ..... .. .
e Diplodunta (Felaniella) usta GOULD ............. . . . .. . .. .. . . . . ... . ... . .. .
e Cm'dium (Tra chyca?'d'i um) shiobm'ense YOKOYAMA ........ . ...... . .. . ... . .
C(Lj'dium (Cemstor/e?'11w) sp ........ . ...... .. ............... . . . .. ......... .
e Cm'd'i um (Nem ocm'dhtm) .'omamngae Wr.~KIYAnrA . .. . ............ .. .. . . . .. .
e Venus (Me?'cena7'ia) yo7;OYCL7nwi MAKIYAMA ........................ . . . . . .. .
kIact?'a (ltfa ct7'omi'?"is ) polynyma voyi (GABB) . . ......... . . ....... . .... . •..
e .Jfacoma tokyoens·is MAKlYAMA .. . ... . . . . .. .. .. .. .. . ... ..... . .... . . . .. .. .
e Aloiclis venusta (GOULD) . ... . . . . ... . . .
.. . ........ .. ........ .
Thmcict higasinorZonoensis n. sp.. . . ...... ... .......... . .. . .. . ... . ...... . . .

GASTROPODA
e So/a?'iella sa.7;ya yOKOyAMA ... .... .................. . ... . .. . . .. ... . .. . .. .
TU?'cicula (Lj'genteonitens (LrSCHKE) . . . ... . .... . . .......... . .......... . . . .
e TU?'?'itella kadonosawensis OTUKA .. . . . .. . . . ............. .... . . . ... . .... . .
Actae~pymm'is. (~xi7nia (LrSCHKE) ? ......... , ... . . .. . .. .. . . . . .. .... ...... .
CaptLlus sp. . . . ..... . .. .. ............ . ... . . . .... . ...... .. . .... . ........ .
C7'epidula sp ......... . ... . ... . .... '. . . .. ... ..... .. . .. .. .. .. . ..... . .. . . . .. .
e Natica (Tectonat'ica) .iantho.~toma DESHAYES . ... .. . . ... . . . . ... . ...... . .. ,.
e Natica (Neverita) didymo. BOLTEN .. . . . . . ............. . .. .. .. . .. . .... . ... .
e B1.L ?·sa yabei NOMURA et RATA! ..... ... . . ...... . . .... . '.' .... . .. .. . .. .... .
e Phos (Comeophos) iwctl.:iantIS (YOKOYAMA) . . .. ...... .. ...... . ...... .. .... .
e OZ'i:vellct iwakiensis NOMURA et RATA! . . .......... . .. . . .. .. . ........ . .. . . .
Olivella consobl'ina (LrSC LIKE) .... . ...... . .. . .... . . '.................... . . .
Fulgoraria (Psephaea) p7'evostiana (CROSSE) ....... . ... . ...... . ... . .... . . .
e Cancella?'ia hu ku.limana NOMURA et RA'l'AI ........ . .. . ... .. ... . .... . . . . .
e Surcuz.ites (ltfegasU?'cula) yo7coyamai O'l'i :KA . . .......... . .. . . . . . . . . .. . ... .
Oylich na kozukensis n. sp. . .. ... .. . . .... . .... ... .. . .. . .... . . . . . . .... . .. .

F
F

R

R
R
R
R
F
F
F

R
F

R

R
R
R

R
R
F
R
R

F

F
F

R

R
F
F
R

R
R
R

R

(R. " . rare; F .... frequent)Loc. 1. Rigasinodono, Iwanoya·mura, Usni·g un.
Loc. 2. Suyama, Iwanoya·mnra, Usu.i'gun.
Loc. 3. Annaka·mati, Usui'gl1ll.

It is interesting that ther e are many common species between the fossil faunaof the Itahana bed and that of the shell beds of TaDagura, Hukusima-ken, described by Dr. YOKOYAMA l ) and Messrs. NOMURA and HATAI2). The geological ageof the Tanagura Beds is considered by the latter author s as the Middle Miocene.
The mollui::ican fauna of the Itahana bed is also related to that of the Kadonosawa
series in I wate-ken. Now Messrs. OMURA, SAWATARI, and SA:J'O of the Nippon Oil
Company are studying the geology of the above titled region. After the completionof their geological survey, the age of the Itahana bed will be diEcus:=:ed.
T he type specimens of Glycyme?'is 7c-8u zu7cii n. sp. collected by Mr. MURAKOSI
are p reserved in the Geological I nstitute, Faculity of Science, Imperial Universityof Tokyo. · The other specimens described in this paper wer e presented by the·e Common species between the Tanagura Beds and the Itahana bed.
1) YOKOYAMA: Jonr. Fac. Sci. Imp. Univ. Tokyo, Sec. 2, Vol. 3, Pt. 4, pp. 197- 203,1931.
2) NOMURA and I-lATAI: Saito Ro·on Kai Mus., Res. Bull.,' No. 10, pp. 109-155, 1936.

-88-

Neogene Shells from the Vicinity of the City of Takasaki, Gumma-ken, Japan

673

Nippon Oil Company to the I mperial Geological Survey of Japan .
Description of the Species

Glycyme?'is 7c-suz'L67cii n . sp.
Pl.. 20 (7), figs. 1, 2, 3.
Shell fairly large, thick and solid, suborbicular in outline, somewhat longer than high,greatly inflated, nearly equilatera I. .A n terior and p csterior ends rounded; an tero andpostero-dorsal margins decending, slightly Convex; vel1tml margin regularly arcuated. Beakmther swollen, prominent, located almost at the center of the dorsal border. Surface ornamented with many mdiating impressed lines crossed by concentric growth lines which are
numerous and irregular. Interior of the type specimen unknown . I nterior of pamtype :Ligamental area mther broadly triangular, ' equilateJal, flattened, tmDsversely finely striated.
Cardinal plate large and thick, the upper margin nearly stmig'ht, the lower one broadly
curved, obtusely angulated at the middle. Teeth about 10 on each side. Inner surface with
faintly radiating striae. Muscular impression distict, thickened.
Holotype (in Geol. Inst. Fac. Sci. Imp . Univ. Tokyo.) 73.4mm long; 63.3 mm h igh;
29.1 mm thick.
OCcul'l'ence : I-Iatonosukake, Iwa daira-mum, Kita-kanra-gun (type locality) ; Namerisawa, Annaka-mati, Usui-gun; Higasinodono, Iwa noya-mura, Usui-g un; Suyama, Iwanoya-mura, Usui-gun.

This species is closely allied to G. cisslmensis MAKIYAMAJ ), but it differs fromtbat species in that the shell is more equi lateral, more inflated, the beak is larger
and moi'e swollen. G. k-suzulcii also r esembles G. nalcarm6?'ai MARIYAMA2 ) and G:
OinO'L6yei NOMuRA 31• Compared with G. nakamv.mi, the new species has a nlO1'econvex shell with larger beak, more numerous radiating striae on the surface, andbroader ligamental area. It also differs from G. oinouyei in having a more dilatedshell with more swollen bealc The present new species has the most convex shell as
seen in the followiJJg tab le.
G. c:sshuensis
G. na7~amunti
G. oinouyei
G. k-StlztGkii

Length
67mm
78
61.5
62
73.4

I-Ieight
62 mm
71
51.5
61.7
63.3

Thickness
22 mID
26
22.5
19
29.1

(3 : 2.8 : 1)

(3 : 2.7 : 1)
(2.7. : 2.'l : 1)
(3.2 : 3.2 : 1)
(2.5 : 2.2 : 1)

The new species differs from G. impericdis KmwDA·1), a living species of Japan.
in that the shell is much larger and longer. G. 1c-.S1.mdcii is easily distinguishedfrom G. cdbolineata (LrscHKE)5), a common recent species, in being more circu lar inoutline and in possessing a more inflated shell.
Tho specimen from the Tallagura Beds, which was referred to G. albolineata
by Dr. YOKOYAMA 6) , is probably th.e present species. It was dedicated to Mr. K.
1)
2)
3)
4)
5)
6)

MAKIYAMA: Mem. Col I. Sci. Kyoto Imp. Univ., SeL B, Vol. 2, No.3, p. 155, pl. 13, f . 3, 3,1926
MAKIYAlIfA: Op. cit. , Vol. 3, No.1, p. 30, pl. 1, f. 5, 6, 1927.·
NOMU RA: Op . cit., No. 5, p. 74, pI. 3, f. 1, 1935.
KURODA: Ven us, Vol. 4, No.4, p. 201, pl. 4, 1934.
LISCEIKE : J ap. Meer. Conch., Vol. 3, p. 108, pI. 9, f. 11, 12, 1874.
YOKOYAMA: Op . cit., Sec. 2, Vol. 3, pt. 4, p. 199, 1931.
- 89-

T. OINOMIKADO674

SUZUKI, who first reported at the meeting of the Palaeont0logical Society of Japan(1937) that the Glycyrne?'is from Hatonosukake, I wac1aira-mura is new to science,but he did not give any descriptions. ,Cm8satellites s~6y(6?nensis n . sp.
PI. 20 (7), figs . 9, 10.
Shell small, trigonal, son~ ewhat longer than high. Anterior end rounded; posterior end
obtusely truncated. Antero-dol'sal margin straight, sloping, even slightly excavated near the
beak. Postero-dorsal marg in somewbat convex . Ventr al margin broadly arcuated, finely
crenated. Posterior l'idg e obsol ete. Sculpture consisting of n:any , concentric rounded ribs,
obsolete on the postero-dorsal border. Beak small, pointed, sli ghtly inclined forward . Lunule distinct, longly ovate. Escucheon lanceolate, longer tban the ·lunule.
Measurement:
HeightThickness
Length
4.7 II m
Holotype (in Imp. Geo1. SUl'v. Japan)
15.6mm
IS.5mmParat ype
14.8
unknown
IS.1
Occurl'e~('e :Suyama, Iwanoya-mura, Usui-gun.

This specie[' resembles C. nanus (ADAMS et REEVEr), a species found in theJapanese waters. It differ s from the latter species by its more steeply sloping
convex poster'o-dorsal margin, and. less arched ventral margin. C. pauxila (YOKOYAMA)2) from the Neogene deposits of Titibu is an allied species. But this new
species has a more acute apical angle and finer concentric ribs.
CaTdita (J11iodontisw s) nalcamvmi annakensis n . subsp.
Pl. 20 (7), figs. 7, 8.
Tbe shell is closely allied to C. n akarntwa·i ( YOKOYA~{;A)3) but differs in its more numerous rediating ribs. 'i'be species has ten to twelve ribs, while the r ibs of the subspecies are
fourteen to sixteen in number.
Measurement :Thickness
L engthHeightHolotype (in Imp. Geo1. SUl've. Japan)
4.Smm
1 .6mm
4 .2 =
Paratype No . 1 .
1.2
3.7
3.7
unknown
Paratype N o.2.
4.5
4.0
Occurrence: Annaka-mati, U sui- gun.

The species is closely allied to C. pTolongata (CARPENTER) 4) . I n a former
paper5) the writer combined th em into a single species. Dr. SCHENCK has examined
the specimens of C. nalcarnumi (YOKOY AlIfA), which were forwarded by M~. OTUKA,
and answer ed him as follows "Comparison of J.11iodontisc~6s p?'olongata and' Vene?'ica?'dia' nalcarnu?Y6i shows them to be congeneric but not conspecific".
Th?'acia higasinodonoe'};l,sis n. sp.
PI. 20 (7), figs. 11, 12.
1)
2)
3)
4)
5)

ADAMS and REEVE: Voy. Samarang Moll., p. 8+, pl. 23, f. 2, 1850.
YOKOYAMA: Op. cit., Sec. 2, Vol. 1, Pt. 3, p. 122, pI. 15, f. 8-11, 1925.
YOKOYAMA: Jom. Coll. Sci. Imp. Univ. Tokyo, Vol. 44, Art. 7, p. 5, pI. 1, f. 9, ]923.
CARPEN1.'ER: Brit. Assn . Ad\7. Sci., Rept. for 1863. p. 642, 1864.
OINo~nKADo: Jour. Geo1. Soc. J apan, Vo1.44, No. 520, p. 61'1, 1937.
-

90-

Neogene Shells from the Vicinity of the City of Takasaki, Gumma-ken, Japan

675

Shell rather small, ovately transverse, swollen in the central and umbonal area, compressed in the postero-dorsal area, rounded in f ront, and truncated behind. Antero-dorsal
margin n early convex ; postero-dorsal margin stra ight and horizontal. Anterior end broad,. r ounded; posterior end truncated, narrower than the anterior. Ventral margin broa dl ya rched, rounded in front , slopin g straight along t he p osterior border, slightly exca vat ed
neal' the p ostero-ventral cornel'. Surf ace sculptured with irregular concentric growth l ines.
Posterior r i dge oblique, r a tb er distin ct. Beak not prominent, located neal' tbe middle ofthe sbell length .
JVL asurement :Leng thHei ght 'l'hickness (Both valves.
9.1mm
Holotype (in Imp . Geol. SUl"v. Jap a n )16.3mmclosed)26.9mmOccurrence: Higasinodono, Iwanoya-mura, U sui-gun.

This species resembles T. conci na GOULd) ( = T. papy?'acea of YOKOYAMAr),but differs from that species in that the sh ell is hig her; the ventral margin ismore arched; the surface behind the posterior ridge is more depressed in the
middl e ; the postero-ventral angle m ore acute. T. be?'ingi DALL3) ( = T elline lcalm.('?ncma YOROY AMA)") is also an allied species. But Dall's sp ecies has a larger
shell and more steepl y sloping dorsal margin . T. pe?'tmpezoides NOMURA 5) is a.
sh ell closely allied to thi s new species. But the former has a larger sh ell, moreprominent beak, and less oblique posterior ridge.
Cylichna lcOz'i(,lcen si s n . sp.
PI. 20 (7), figs. 4, 5, 6.
Shell rather la rge f or the Genus, thin, cylindrical, convolute, t apering at both ends, broadest
at one-third of the length from the a nterior end; post erior end h'uncatecl, anterior end
rounded; surfa ce with obtuse lines of growth and fin e impressed t ransver se striation s, thelatter more distinct neal' the upp er and lower ends a nd a fin e' inter strial line lying between
the striations nea r the a nterior end; ap erture as long as the shell-height, na rr owed in the·posterior por t ion, dilated in the anterior; outer lip thin.
Measuremen t :D ia meter
H ei ght
17.4mm ;7.7 mmHolotype (in Imp. Geol. Slll'v. J3'pan )Ma ximum diameter of a perture
5.8mm
Diametel' of a perture at apical portion
2mm
OCCUrrence: Annalea-mati, Usui-gun .

O. ?7w sashiensis TOKUNAGA6) is an allied allied apecies. But the new species is.
easily distinguished by its surface sculpture and more dilated aperture. I n the
sculpture the present species resembles C. cop1('lenta YOKOYAMA7\ but it differs.
1) GOULD: Otia Conch., p. ]61.
2) YOKOYAMA: Op. cit., Vol. 44, Art . 1, p. 171, pl. 14, f. 12, 1922.
3) DALL: Proc. U.S. Nat. Mus., Vol. 49, p. 442, 1915.
4) YOKOYAMA: Op. cit., Sec. 2, Vol. 2, P t. 4, p. 177, pI. 67, f. 14, 1927.
5) NOMURA: Op. cit., No . 6, p. 50, pl. 7, f. 4, 1935.
6) TOKUNAGA : Jour. Cal L Sci. Imp. Uni v. Tokyo, Vo l. 21, Art. 2,32, pI. 2, f. 12, 1906.
7) YOKOYAMA: Op. cit., Sec. 2, Vol. 1, Pt. 7, p. 217, pI. 28, f. 3, 1926.
- 91-

676

from the latter
front.

T. OmOMI KADO
III

that th e sh ell is slender, and th e aperture I S less dilated lD

Explanation of P late 20 (7)Fig. 1. Glycyme1'-is 7c-suzttlcii n. sp . Specimen from Namel'isawa, Annaka-mati, Usui-gun,
Gmnma-ken. (x 1).
F igs. 2, 3. D itto. (Holotype). ( x 1).
F igs. 4, 5, 6. Cylichna 7c6zu7c en sis D. sp . (Holotype). (X 1.5).
Figs. 7, 8. Cl.wc7itct (llliodontis cus) ?Ht7camtwa-i anna7cen sis n. subsp . (Holotype) (x 3)
F igs. 9, 10. Cmssatellites sttyum ens is n. sp . (Holotype) (x 1.5).
Figs. 11 , 12. · 17wacict higasi?loclonoensis J;l . sp. (Holotype) (X 1.5)

;jl:f.I!!§!Wf.~III~·nl~t·j-jlI: O)m:5i~-=i1;effit 1t;qj(

1* 1f!IlF~~(100W)*U!

Ep f>:.!S& I/¥4J;:1-W:tfs 6 le}Wtl* vclj~ -cit.*P.t-&JH fr.-1ft ~ ~fU~ ~\f\ l~ 1= EEl, 1:\l1IJ 111]'1~;j j[ CJ.)~jf> ~*i3 Jl.1t;fi :>l>$llJ ;! ;f1.
t t:. o filJJ:Il!.1i·CJ.) Jll!.rt vclMdL -C r.tjf},i~j'f:t?f.k:1W±2f("'f/H*~~ ~ CJ.)WFrc:>l><b Q
lJU),tK flClliZ U::: Jl.1t;fir.tiiilPl>:., /H;jqqtj~CJ.)
1P.!j,\{(ii-CJ.) t CJ.)-"(., 1P.!j,I.f(ii-CJ.)1t;fi 7 ,..- -;1- r.t ii\'ii .Q;Y.\f\t!mfllnr l~j·j[ CJ.) ~i:M;~~i3Jl.1t ;fi'Wn: ~ 1..- <1JJ'ifW. L., 3Z 1fWo-=r=-1%CJ.)
I"] / 1~1l;f%CJ.)~) CJ.) 2: t IMd'fW<7.5'<b Qo f:I -F 13:<1>:.;fiiur\'Ht!:CJ.)*tf ~~H, 1tllltiJJ:~¥±, 1H~~~±;:l~~jll!.1fCJ.)jll!.'I1 K
JlX ~ WFrc 1ft c:-, ill!. '11%'1-1;; VC)f'\:v' -c f';tJ:ftcm\i·~ CJ.) T!tm;:l>:71:1iX T Q ~1*l1!-j- Q
0

0

-

92 -

Vol. XLV, PI. 20(7)

4

11
7

5

6

·

~

'p

70. A New Species of Calyptogena from theHigasiyama Oil Field, Niigata-ken, Japan* 'By

Tuneteru OINOMIKADO and Kinji KANEHARA(Received lYIay 27th; read June 11th, 1938)

In 1937, one of the present writers (OINOMIKADO) made a large collection offossil sh ells from the Usigakubi bed of the Higasiyama Oil Field, Niigata-ken .
Some specimens of a peculiar shaped vivalve, Calyptogena nipponica n. sp. havebeen founded among the fossil fauna. In the same year, the another writer (KANETIARA) vi::;ited the oil field, and obtained two speclmens of this new species froma water well sunk into the black shale of the Kubiki series at Nakanosawa, Higasiyama-mura, Niigata-ken.
•
There are three known species of the genus Calyptogena,
including C. nippo nica, which seems to be exti nct. C. pacifica DALL' ) is living in the Clareuce Strait,
Alaska to the Santa Barbara Chann el, California, and is also known as a fossil
from the Pliocene of California. R ecently it was r eported by OTUKA~) from the3
Pliocene of the Oga P eninsula, Akita-ken, Japan. C. elongatct DALL ) is anotherr epresentative of the genus, and now living in the waters of Santa Barbara I slands
to San Diego, California.
The specimens from the U sigakubi bed were contributed to the Imperial
Geological Survey of Japan from the Nippon Oil Company.
Genus Cctlyptogenct DALL, 1891
Type (by monot.ype), Ca7.yptogena pac ifica DAI.I., Pl'oc. U. S. Nat. lYIus., Vol. 14, p. 190, 1891.

Calyptogenct nipponiw n . sp.
Holotype: A both valves closed specimen from the Usigakubi bed (Pliocene); preservedt he Impel'ial Geol ogical Survey of J apan . .
Desc?'iption: Shell model'ately. large, rather solid, transvel'sely elongate, strongly inequ ilateral. Beaks low , tumid, sit uated ab out one-third 01' one-forth of the length from theanterior end. Anterior end rounded; posterior end more broadly I'ounded. A nter0.-dorsal
mal'gin short, slight ly ex cavated in front o.fthe beaks; postero-do.rsal margin lon g, gently
arcuate. V cntr1l1 margin nearly straight, somewhat co.ncave near the middle. Surface sculphued with co.ncentric incremental l ines, slightly depressed neal' the centel' of the ventral
portion. N o. lunule. Ligament long and strong. E scutcheon lanceolate, bounded by eleva ted
ridges, long, extending to the half o.f the po.stel'o-dorsal marg in. Hinge normal.

lU* By the permission of Dr. S. YA~rANE, Director of the Imperial Geological Survey of Japan.
1) Proc. U. S. Nat. lYIus., Vol. 14, p. 190. 18~1.
2) Jom . Geol. Soc. Japan, Vol. 44, No.. 522, p. 231, 1937.
3) Proc. U. S. Nat. lYIus, Vol. 52, p. 408, 1916.
-

93 -T. OINOMI K AD O and K . KANER ARA

678
Dim. ent·i ons :Holotyp e.

Height46.1 mill

L ength115.4 mmThickness
15mm (one valve)Compa?'isons : At fir st sight this shell resembles somewhat T?·apizi 'l.('m japoni cum P ILSBRY,l) but th e escutch eon and the hinge arc entir ely differ ent. Th e new
species is distinguished from Calyptogena pacifica DALL by a larger and m ore
elongate sh ell. C. nippom'ca is closely allied to C. elongata DALL, but differs fromt nat species in th at the . h ell is ver y much lar ger and more solid ; th e ant erior
end is llarrower; and the surface of the ventral portion somewhat depressed.
Paphi a hachi yai NOMURA2) of th e Miocene of the Nisi-Tugaru District, Aomori-ken,.
is also an allied species. But NOMURA'S species h as a high er sh ell and broad ly
arch ed ventral border .
T ype locali ty : Usigakubi bed (Lower P liocene). In a g ray shale, on theeastern bank of the Mae-kawa, about 1.2 km . south of th e village offi ce of Nisitanimura at Nakamura, Ni.'itani-mura, Kosi-gu n, Niigata-ken.
R ema?'!cs : The occurrence of the species in the type Jocality is not rare, AsaJready stated , K ANEBA RA obtained this . pecies from the K u biki series (Miocene)_
MITUTI, geologist of the I mperial Geological Surv8Y, colleted this Calyp togena fromhis K atsuura Beds (Upper Miocene or L ower P liocene) in t he Boso P enin. u Ja ..
T h e geological range of the new species, h ithrto known, is Miocene-Pliocene.

ExpJanation of P late 21 (8)Calyp togena nipponica n .. p.
Fig . 1.
Fig. 2.
Figs. 3, 4.
F ig . 5.

H olotype. ( x 5/7)
Di tt o, dor ·a.l vie ,,·. ( x 5/ 7)
Specimens from Nakanosawa, H ig-asiyama-l11 ura, N iigata-ken.
P a r atype. (x 6/7)

( x l)= ;f:lcft. Calyptogtna Mijl';t D AJ.L (1891) 1'L1t{ ~ fJlj1l)t ~:11-, :m:~;fillV;t C. pacifica D ALL ""('ib ~ ~;;.:fillI';tM;{£ Alaska.
:h. ; Californi a i'C:7};{fj L. , Z>F1t;J:i1';t Cn liforni a Q.)Et$Wr*X:h>; r-n ;:I1-, :}:!tw.m:h>; 'b Joc ili::T~Jg:l±K1t{ H.;ic EEl kl!l\,lg0)llI4' .I:h Q.)Et'''Wr~~ :h> ; $"* ~ :h-t::o :id'tK{l'!J I"JV;t:m:.illiib EEl K1llt~-,::; LI~ 7 -m'/E} J: ~Calyptogena Q.)jffr:fill C. nipp onica;.

B:=l*1I~ l..Jc ii', ~}jf( r.tlllt-!IlIS]f! ill ;j:;j 9' 1~-Ik Q.) /J ,JJ}ttJiiHq~! G:I1- tdLFi (ljk;j47M fl ) il, G=l*~ ii :11- k ~;::f]ll Q.)~~;;'B: ~ tc
J1;t G1)/N i: }*IIHili: K l';t'i'i"iEI/Erii'1llt@ L-, 1t;(:J I1::lI1! ~ T 62 R G1)i$s, :h> G=l*:I1-t.: :ttt:i\ ;f;"N~;;'K 1';t *1f-G1)lh, .tI~)T -m-/E}.i%lEQ.)
00'b G1) B: ffil5t L- t.: 0 C. nipponicct Q.) fl1!'L'tII'!f1-~r.t<tEI:itr-n ;:11- -C J,;, ~ i$s,""('1';t o:p~i'lltJ:'lliS:h> ; .@foWr-l!tl~~S""('</:> ~ 0

1) P roc. Acd . Nat. Sci. P bila ., Vol. 57, p. 119, pl. 5, f. 34-36, 1905.
2) Saito HO- Oll K a i Mus. Res. Bull ., No. 6, p. 55, pI. 3, f. 5, 6, 1935.
-:)4-

Vol. XLV, Pl. 21(8)3(.'... ~.

:~ -. ,.).

1. *~~, 8 *JIl!~~if / $ ir .: :/ T 8 *tr~%J~if 1· fm.7.
2,*ir~,tr~%J~:&. e'Z 1/ .:rMl7, J l.-~~fJf / ~2l7'7 DJJ'rWr~/ "/If:&. '7 !mllJv '7 ~T ~B'-.J 1·.7.

5.

* 1Wr / ifil!t ~'1F1m3l1ID 1,:/, 8 *jll!'i~~if,di'!e'1F:ifjP ~ 1,.7., @:/-fl.'if.:~ 100 ~.l;J.J:. '7 1!lr~ft -!Ul.- :1!f'7'lll:
D:JJlWr f!l. .=. :fIE 7,

1

B

WI
;m;
ii'fll;

:R

:R

~

ffil

131'

*

*

J5 4 tto/J "~r 1ilr i5t ~
1=1

JJj

~

rJ;X

:;.1:1: J:: ffifl ;L lJ)]

JEt

X

1i£

h'& *i!&\ :1k m:**JJi* fEl *i' (OJ vij

~13

WF

*

*~l

-

~

;1111

ijg{

gj'f

1L

~~

*

/ '\

~"J' ~i'r!:J(~:t< Il Ii'3\~'* 7.( Jilll :R* *t!if:;j(*~~ !JW ~*

I.Li *[~ ~~i'

J::.

13

1~

*7dx fW rH~ J\.1Ft**~ 81Ji7 80 1Ii= )'Pill :R

*7( j~ ~p.ffi ;L DjJ
~

.NII

*/)'

~*

JJj

*..!f-~

5~(* 1j;'~~ffiU

Jil~

Constitution of the Palaeontological Societyof Japan.
Article 1. The Society shall be known as the Palaeontolog ical Society of Japan. It forms a sectionof the Geological Society of Japan.
Article 2. The object of the Society is the promotion of palaeontol ogy and related sciences.
Article 3. This Society to execut e the scheme outlined under Article 2. shall hold annual meetings anddiscussions.
Article 4.

Proceedings of the Society and articles for pnblication shall be published tlll"ough the
Journal of t.he Geological Society of Japan. Separates and circulations will be sent tomembers of the Palaeontological Society who are not members of the Geological SOCiety
of Japa n .

Article 5. The annual dues of this SOCiety is two dollars for the fore ign members of the Society.
Article 6.

This Society sball hold the following executives. President one person, Councillors several
persons.

Article 7. The President and Councillors shall be elected annually. The President and Councillors
shall be elected from the Society body by vote of its members. All elections shall be ballot.

President

Takumi NAGAO

Councillors Ichiro HAYASAKA

Haruyosi HUZI1IlOTO*

T s unen aka IKI

K in osuke INOUYE

Nobuyas ll KANEHARA

rrakeo KA'J'o

Rokuro KniURA

Teiiehi KOBAYASHI*

Hanzo MURAKAMI

Takumi NAGAO

Shintaro NAKAJVIURA

TUnetel'U OINOMIKADO*

Ichizo OMURA

Yanosuke 01'UKl\ *I wao T ATEIWA

Shigeyasu TOKUNAGA *Hisakatsu Y ABE"

Shinji Y AMANE*(" Executive committee)

All communications relating to this Journal should be add ressed to the
PALAEONTOLOGICAL SOCIETY OF JAPAN

Geological Institute, Faculty of Science, Imperial University of Tokyo, Japan