fJ -Et~~±.~~~~~· ic.fTransactions and Proceedingsof thePalaeontological Society of Japan
New Series

l'alaeontological Society oi japan
April lOth, 1963

CONTENTS

TRANSACTIONS
Page

442.

Some Jurassic Trigoniids from the Tetori Group in the Kuzuryu District,
Central Japan443.

............................................ Shiro MAEDA

Additional Notes on Late Paleozoic Corals found in the Southern Part of
the Tamba District .......... Shigeo SAKAGUCHI and Nobuo YAMAGI\\'A

444.

Lower Carboniferous Bryozoa from the Omi Limestone. Japan.

H6.

9Part 2:

Successive Descriptions of Cryptostomata .............. Sumio SAKAGAJIII
445.

1

15

New Species of Gadus from the Pliocene of Japan ..................... .
. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Kotara HAT AI and Tamio KoTAKA

25

Remarks on Some Fossil Dosinia of Japan .............. 1\.oichiro MASUDA

29SHORT NOTE
10.

Lilloetia sp. (Ammonite callovienne) de !vlitarashi au Bassin de Tetori ..
. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Tadashi SATU et Yasumitu KA!'\IE

8

PROCEEDINGS . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .

39

Honorary President:

Hisakatsu YARE

President : Teiichi KOBA YASill
Councillors:

Kiyoshi ASANO (Editor of "Fossils"), Riuji ENDO, llaruyoshi FI!Jil\IOTO,

Shoshiro HANZAWA, Tetsuro I-lANAI (Secretary). Wataru HASHli\<JOTO (Treasurer),
Kotora 1-IATAI. lchiro HAYASAKA, Hiroshi OZAKI (Planning), Teiichi KOBAYASHI,
Tatsuro l\IATSUl\IOTO (Editor of Special Public:1tions), Masao \liNATO, Tokio
SHIKA~IA(rvlembership), Fuyuji 'L\KAI (Editor), Ryuzo TORIL\l\IAAssistant Secretary:

Takeo ICHIKAWA

The fossil on the front page is Helicoprion bessmwtci KARPINSKY, 1899.

All Communications relating to this Journal should be addressed to the
PALAEONTOLOGICAL SOCIETY OF JAPAN
Geological Institute, Faculty of Science. University of Tokyo, japan

Trans. Proc. Palaeont. Soc. Japan. N. S .. No. 49, pp. 1-7. pl. 1. Apr. 10. 1963

442.

SOME JUHASSIC TRIGONIIDS FHOi\1 THE TETORI GROUPIN THE KUZURYU DISTRICT. CENTRAL JAPAN*
SHIRO MAEDA
Geological Institute, Chiba liniversity·fLiiJ'ifigtt!!~I1)'J'-liJtM11lH'E /"' 7 f:C:::fTJ f!.M : :h.Jm~iUI!!ll.!XtJ ·I? u)::::::frl flllfH'i.:: ;h, i ·cJlllij(;!J(
1i.h?i t W-lllli~!M t fJ. I? fx:fJI\il;.k!ll\X ~ ;11, ·n ·t.:.tJ>, .:: 11) t.:.V:1:i-ili1'i- lllii:*I1)1J.HJi0tl1LI\'llJtll1)/iPJ5
fJ.t_,'$1[Gt.:.l.:.~~~t'lt.:., .::;h,l?~ftitfltLt..:t'im. 5 ftl!tJ'~liHJij~;h,. -t'l1)1'j:l, 4 fifi (Lalilrigonia
horii. L lursaii. lbotrigonia tetorieusis. Vaugonia kuzurywmsis) l:t~fru-c·, .::;: r.:.TcJ!iXL

n m ~

~.

Introduction
The J urasso-Cretaceous Tetori groupis well exposed along the upper tributaries of the Kuzuryu river in Fukui
Prefecture (Prov. Ech izen). Describing
the following four species of trigoni ids.
KOHA YASIII (1956. 1957) concluded that
the Yambarazaka alternation of the district is uppermost Jurassic in age.
Vaugonia yambare11sis
KoBA Y .-\SHJ

of the Kuzuryu (jU.l.i'(fiiV river.
Because the alternation yielding f\.ranaosp/zinctes matsus!Jimai. Peltoceratoides
sp. and Ptyc!Jop!Jylloceras sp. is middle
Oxfordian. all of the trigoniids below
listed are Oxfordian.
Latitrigrmia horii MAEOA. new species
Lalitrigrmia kasaii MAEDA. new species
Ibotrigonia tetoriensis MAEDA, new species
Vaugouia kuzrtt)•ttensis lVL'\EDA. new species
Myop/wrella.•....•.•..•

Yambara conglomerate. baseof Itoshiro subgroup.

[.atitrif!Oilia tetoriensis
Kon.-•:YASlll .. -. · · · · · · · · Yambarazaka
Lotilrigonia orbicularis
alternation.
KoBA Y.\SHI . . . . . . . . . . . . uppermost;l{_I'Op/torella (PrmnyoKuzuryu
p!un·etla) orienta/is
subgroup.

1
J

KoBAYASHI and TA!\IU~.\

Lately the writer made a collection of
trigoniids from the Yambarazaka alter·
nation at the east end of Goribashiri on
the Taniyamadani ({iilJ{i). a tributary* Received Jan. 17. 1962: read at the Annual :Meeting of the Society at Sendai. Jan.
20. 1962.

~

KOBAYASHI(Promyophorella)orienta/isand T A~lt:RA

Here the \Vriter expresses his sincere
thanks to Prof. T. KonAYASI-11 of the
University of Tokyo for his kind advices,to Dr. M. TAJ\!URA of the Kumamoto
University and Dr. I. HA YA!IH of the
Kyushu University for the suggestion,
and also to Messrs T. KAWABE, S. KA·
WAMATA. I. SuzuKI. A. OrRA and S. SASAWKA for assistance in fossil collection.
Stratigraphical notes

The group in this district can be clas·
sifted, in descending order, into the fol·lowing formations:Shiro MAEDA

2
Kitadani alternation

Akaiwasubgroup(about 100m thick) (P)fl Akaiwa
sandstOI~e_(about 500 m thtck) lP. WJ
Itsuki alternation(150-250 m thick) (0. C. W)
Ofuchi conglomerate
Itoshiro(about 150m thick)subgroup Ashidani alternation(about 80 m thick) (P. OJ
Yambara conglomerate(50-53 m thick)(TJ
Umagadani unconformity
Yambarazaka alternation(43-120m thick) CA. B. T)
Kaizara black shale(195-500 m thick) [A. B. P)
Tochimochivama sandstone
Kuzuryu(200-1800in thick) [B)subgroup
Oidani alternation(200-600 m thick) [P. B)
Shimoyama conglomerate(30-200 m thick)
P: Plant, C: Corbicula. A: Ammonite.
T: Trigoniid.
W: Silicified wood.
0:
Ostrea.
B: Belemnitejl

The Shimoyama conglomerate containing subangulaled pebbles of gneiss,granite. sandstone, slate, cherty rock,limestone and so forth is well exposedat Shimoyama. Ono and Asahi.
The Tochimochiyama sandstone, Kaizara black shale, Yambarazaka alternation and some parts of the Oidani alternation and Yambara conglomerate exposed at Shimoyama, Nakatatsu andKaizara arc marine sediments, while theothers are brackish or lacustrine deposits.
According to SATO (1960) the Kaizara
black shale is divisible into three ammonite zones in descending order asfollows:3) Oppelia zone: Camphylites aff. delmontamts(OI't'EI.). Oecotrausles sp., Oppelia aff.

subradiala Sow., Oxycerites sp., Bomburites aff. g/obuliformis (GE~t:-.tF.LLARo).
2) Grossou1'reia zone: Grossouneia d. sub·tilis (NEU~t.), G. laet•iradiate SATO. G. sp.,
Kepph·rites (Seymourites) japrmicus KonA-· .b: ·.o.·.

I.
.
~

!

~

Or-.;ehi conp:lo:~~erl'ltr:-

~

g.
f

'l'och111HJCbi;-us Eandetone~.::

0tdan1 !!iltf'l""f',atior:.

!

---t"'~~~~~-tg---~___~L

A

B

c

0

E

Text.fig. 1. Schematic Columnar Sectionof the Tetori Group.
A. conglomerate: B. conglomeratic
sandstone: C. sandstone: D. shale ;E. tuffaceous rook.
Y.-\SIII, /\. (S.) UCIIficoSI/1111 1\.0BAYASIII,
K. (S.) lmzuryuensis KonA Y A Slit, Lilloetia

sp.
1) _Veu:ucuiceras zone: Seuqucnit't'l'as yoko·

442.

Jurassic Triaoni1'ds

yamai (KoiL\YASHJ and FuKADA). N.
yokoyamai var. alticostatum SATO. N.
maedai SA To. Calliphy/loceras sp.. Hof.
cophylfoceras sp.

The ammonites show the Callovian
age of the Kaizara black shale. Though
the shale is extensive from Kaizara toOno, no occurrence of trigoniids hashitherto been known in it.
The Yambarazaka alternation consistsof conglomeratic coarse-grained sandstone and micaceous sandy shale, presumably indicating a regression of theTetori sea. The '{ambara conglomerate
which is typically developed at Yambara
along the Itoshiro river, lies unconformably on the Yambarazaka alternationand contains boulders of gneiss. granitic
rock. quartz-porphyry, sandstone. slate.
limestone and so forth. It is remarkablethat the sandstone boulders derived fromthe Kuzuryu subgroup are abundant.
The Ashidani alteration well exposed
along the ltoshiro river. consists of alternation of fine· to coarse-grained sand·stone and micaceous sandy shale andcontains Ostrea sp. at some horizons.
The Itsuki alternation consists of fineto coarse-grained sandstones and micabearing black shale, and yields Ostreasp., Corbicula (/11l!Socorbicula) tetoriensis
KoBAYAStll and SUZUKI and other nonmarine molluscs and fossil plants suchas Onychiopsis elongata (GEYLER). Cladopltlebis dcnticufala (BRO!\Gl\IART). C.
exiliformis OISHI. Xenoxylon latiporosum(CRAMER) and so on.
The Akaiwa
sandstone in this district consists
mainly of arkosic coarse-grained massive
sandstone. Several plant fossils occur
at some horizons, but no molluscan
fossil has hitherto been found. The
Kitadani alternation is characterized by
the presence of reddish or greenish tuf·
faceous rock facies and yields Ouyd1iopsis
e/ongata (GEYLER). Cladop!:lebis denticu·

lata (BRONGNIART). C. exi/iformis OISHI.
Nilssonia orienta/is BEER. Podozamites/auceolatus (LINDLEY and I I UTTON), P.
Reinii GEYLER and so forth. The alternation is early Cretaceous in age.
Description of Species

Subfamily Trigoniinae
LAMARCK, 1819

Genus Lalitrigonia K013A YASH I. 1957

Latitrigrmia Jwrii ~IAEDA, n. sp.
Pl. l. rigs. 1-6.

Description :-Shell subquadrate. wellconvex. convexity attaining the maximum on marginal carina at about onethird below umbo and nearly as long as
high : marginal carina diagonal ; test
thick. Umbo located at one-fourth or
one-fifth from front: anterior marginrounded: ventral broadly arched: siphona! arcuated ; anterior and ventral margins disposed with an angle of about
100 degrees bet ween them: hinge typicalof Trigoniae: escutcheon carina somewhat distinct.: marginal carina more orless sharp, nearly straight and largely:uberculated : area extraordinarily large.
divided by a finely tuberculated mediancarina. growth lines fine and transversal: ante-carina! sulcus wide and shallow;
disk ornamented with about 8 concentriccostae \Yhich are parallel to ventral margin and thickened into a large node at
the posterior end.
Measurement (in mm) :Rg. Number; Valve I Length I Height I WidthI

R. 61121501 I{~hlf(Holotype)

R. 611215021 Leit
R. 61121503 Left

351 ~ 5-

I 36
[

30+

I

5
5

Shiro MAED:l4

Comparison :-This species most closely
resembles L. mzituberculata (KOBAYASHI
and T A~!UR:\, 1957) from the Bathonian-Callovian Yamagami formation (4thTrigonian zone) of Soma (TA~IIJRA. 1960).
but it is easily distinguishable from thelatter by having anteriorly situatedumbo and less concentric costae on disk
and being much longer in outline of theshell. The present specimens are also
related to L. orhicu/aris described by
KoBAYASHI in 1957 from the sanw Yambarazaka alternation. but they differ
from each other in featuers of the posterior end of costae on the disk andshell-outline. The specific name is de·dicated to 1vlr. Yoshitaka HoRt of the
Fukui Natural llistory i'vluseurn. who
kindly helped the writer in the studyof the Tetori group.
Occurrence :-Rare in sandstone of the
Yambarazaka alternation at the leftbank of the Taniyamadani river and
1 he east of Guribashiri, Izumi village.
Ono county. Fukui Prefecture.
Latitrigania kasaii \hElM.
n. sp.
Pl. 1. ftg. 7.

Description:-- Shell subtrapezoidal,
about 1.6 times as long as high. gently
convex. conyexity attaining the maxi·
mum on the marginal carina at about
two-thirds the shell height; umbo sharp.
located about one-third from front: an·terior margin well rounded; ventral
broadly arched; siphonal slightly arcu·
ate; test thick : hinge typical of Trigo·
niae. Escutcheon carina distinct, sharplYand finely tuberculated: marginal carina
straightened. somewhat sharp, and tub·erculated postero-ventrally: escutcheon
narrow and provided with several con·centric costellae near umbo; area divided

by tuberculated median carina and withfine lines; ante·carinal sulcus distinct
and shallow ; disk ornamented with concentric costae which are narrower thaninterspace. counted about 10, and thickened into a large node at the posterior
end: concentric fine lines seen in interspaces.
Measurement (in mm) :-Rg. Numberj ValYe Length I Height I Width
R 61l2l5ll L~ft -1 about i 20~·~- 5(Holotypc) ,
: 30
!

!

1

Obsermtion :-Among se\'eral deformed
specimens. the holotype is best preserved.
although it is slightly broken in posterior. The posterior gaping is presumable from growth-line.
Comparison:- This species resembles
L. unituberculala (KOBAYASHI and TA~IU­
RA. l957) closely in the shell outline andcostation. but differs from the latter inthe strong and tuberculate marginal
carina. narrower area and stronger
costae. L. zmicarinata (KOBA YASIII andTAMURA. 1957) from the BathonianCallovian of Soma district in North
Japan has a somewhat similar outlineof the shell. nut it is very different
from the new species in the more densely
spaced costae on the disk. The specificname is dedicated to Mr. Keisuke KASAl
who was a teacher of the Yamanashi
normal school.
Occurrence:- Very rare in a sandstoneof the Yambarazaka alternation, theKuzuryu subgroup. at Taniyamadani,
Izumi village. Ono county, Fukui Prefecture.
Cenus Ibvtrigonia KoRA YASHI. 1957

Ibotrigania teloriensis MAEllA,n. sp.
Pl. 1. figs. 8-9.

442.

Jurassic Trigoniids

Description :-Shell sm:lll. subquadrate.
nearly as long as high, gently convex.
convexity attaining the maximum on
marginal carina at about one-third fromumbo; test thick.
Umbo somewhat
small. located at one-third from front:
beak opisthogyrous: anterior margin
gently rounded. joining ventral margin
with an angle of about 110 degrees which
the latter is broadly arched: siphonal
margin '\vel! rounded:
post·umbonal
margin nearly straight; hinge typicalof Trigoniae. Escutcheon carina distinct
and tuberculated: marginal carina arcuated. sharp and tuberculated: escutcseon
narrow: area fairly large. divided by afinely tuberculated median carina andsmooth ; ante-carina! sulcus narrow andshallo\v; disk ornamented with about lO
concentric costae which are parallel toventral margin. broken into tubercles
on the posterial side and thickened intoa large node at the posterior end.
A!easurcme11l !ill m111) : -------

Rg. Number Val\'cLength Height

Width

R 61121512(Holotypc)

Left

2·1

20

!i

R. 61121316

Right

2Z+

18+

5

Comparison:-This species fairly resembles lbotrigonia masatanii KoBA YASill
and TA:\IURA (1957) from the Yamagami
formation ('1th Trigonian zone of Somal.
Bathonian-Callovian. in the Soma dis·
trict, North Japan. but it is easily distinguishable from the latter by the ornament and shell outline.
I. masatanii yar. from the Yamagami
formation (KOBAYASHI and TAl\lulV\,1957)
resembles this species in ornamentationon the disk. but in this species the shell
outline is subquadrate and costae on thedisk are mostly broken into tubercles

5on the posterial side.
Occurrence :-Rare in sandstone of
the Yambarazaka alternation. the Kuzuryu subgroup. at Taniyamadani. Izumi
village, Ono county, Fukui Prefecture.
Subfamily Vaugoniinae
KOBA YASH!, 195,!
Genus Vaugonia CRICK !VIA Y, 1930
Vaugonia kuzuryuensis lVIAEDA,n. sp.
Pl. l. fig. 10.

D?scription :-Shell small. gently con\'ex: convexity attaining the maximum
on the disk at about one-third height
below from umbo: test thick. Umbo alittle large. located submedially :anterior
margin gently rounded: siphonal wellrounded: hinge typical of Trigoniae.
Escutcheon carina distinct and scarcely
pitted : marginal carina distinct. gently
arcuated. pitted and becomes somewhat
obtuse postern-ventrally; area divided
by a finely tuberculated median carina
and provided with tine growth lines andse,·eral costcllac ncar the umbo : antecarina! sulcus wide and fairly shallow;
disk ornamented with about 8 costae;umbonal three of them gently arcuated:
oth::!rs form Vs at about median portion
on disk; accordingly umbonal ones form
an obtuse angle with marginal carir.a
but become acute ventrally.
:11easuremenl (in mm) : Rg. Number/ Val\'e )_Length I H:·ight Width
R. 611~1513 I R(Holotype)

1

h

28
-----Ig t

19+

8Comparison :-Represented by one external mould with the damaged ventral
portion of a right valve. The costae on

()Shiro M.4EDA

the disk are not tuberculated. Vaugonia
uirano}wme11Sis (KODA YASH1 and MORI.
1955) from the Lower Jurassic in Miyagi
Prefecture fairly resembles it in generalcharacters. especially in ornamentation
on tile disk. but in this species the shell·
size is much larger and costae on the disk
are stronger. Ante·carinal sulcus in theshell is distinct, but in niranollamensis it
is indistinct.
Though V. yokoyamai described in 1955
by KoBA YAS Til and .MOJU from the N i ·ranohama sandstone in :rv!iyagi Prefec·
ture shows some resemblance to thisspecies in shell outline. it differs fromthat species in costae on the disk.
Occurrence:- Very rare in' sandstoneof the Yambarazaka alternation, the Kuzuryu subgroup, at Taniyamadani. Izumi
village, Ono county, Ful<ui Prefecture.

Subfamily lV!yophorellinae
KOBAYASHI, 1954
Genus lvfyop!wrella BAYLE. 1878
J\Iyop!wrel/a (Promyop!wrella) orienta/is
KOHA Y ASHl and T A!IIURA
Pl. I. figs. 11-12
1955. Alyophoretla (Prmnyoplwrella) orienta/is
KOBAYASHI and TA~Il1 R.'., japan. jour.
Geol. Geogr., Vol 26. Nos. 1-2, p. 98,

pl. 5. figs. 6a- b.
1957. Myophorella (Promyoplw1·e!la) orienta/isjapan. jour. Geol. Geogr..
Vol. 28. Nos. 1-3, p. 46. pl. 1. figs. 16-

KoBAYAS!II,18.

Represented by several imperfect ex·
ternal moulds. Shell medium in size;
marginal carina distinct and tubercul-

Explanation of Plate 1
Latitrigonia lzorii MAEDA. n. sp.

. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . p. 3

Fig. 1.
Fig. 2.
Fig. 3.
Fig. 4.
Fig. 5.

Clay cast of right val\'e. holotype (R. 61121501) XIInternal mould of holotype X 1
Posterior Yiew of a clay cast of holotype X I. 3
Clay cast of imperfect left valve, paratype (R. 61121502)
Posterior view of a clay cast of imperfect left valve, paratype( R. 61121503) X 1. 3
Fig. 6. Imperfect left valve. para type (R. 61121504) x 1
Latitrigonia kasaii MAEDA, n. sp ........................................................... p. 4
Fig. 7. Modeling cast of left valve. holotype (R. 61121511) x 1. S
lbotrigonia fetoriensis MAEDA. n. sp. . . . . . . . . . . . . . . .
. .......... .
. ........ p. 4
Fig. 8. Modeling cast of left valve, holotype (R. 61121512)
Fig. 9. Posterior view of a modeling cast of holotope X 1. 7

Vaugonia kuzuryuensis MAEDA. n. sp . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . p. 5
Fig. •10.

Clay cast of right valve. holotype (R. 61121513)

xI. 5

Myoplwrelfa (Promyoplwre//a) orienta/is KouAYA::;HI and TAMURA . . . . . . . . . . . . . . . . . . . . . . . . . . p. 6
Fig. 11.
Fig. 12.

Clay cast of imperfect left valve (R. 61121514)
Clay cast of imperfect left valve (R. 61121515)

X 1. 4x 1. 4

All of the illustrated specimens are kept in the Institute of Geology, College of Arts andSciences. Chiba University. Chiba. Loc: Yambarazaka alternation of sandstone and shale in
the Kuzyuryu subgroup, the lower division of the Tetori group, in Izumi village ('ll:lf.R;ft). Ono
county (:k!t:Jll;). Fukui Prefecture (7M:.Il!Jil:).

MAEDA : Jurassic Trigoniids

2
1

4

'p.

8

10

Plate 1

442.

.J u.ra.ssic Trigon.'iids

ated. Area with numerous. somewhat
coarse and sharp costellae of \Vhich about,1 or 5 are in 5 mm. Disk ornamented
with arcuate costae which form an acuteangl£> with the marginal carina. The
costation is similar to that of KoBAYASHI and TAMURA's form (1955) from compact sandstone (9th Trigonian zone) of
the 1\oyamada formation at Umazawa,
Koyamada, Kamimano village, Fukushima
Prefecture. Therefore the present spcci·
mens are identified with orienta/is.
Occurrence :-Rare in sandstone of the
Yambarazaka alternation. the Kuzuryu
subgroup. at Taniyamadani and east of
Goribashiri, Izumi village. Ono county.
Fukui Prefecture.

References
Cox. L. R. (1954): Notes on the Trigoniidae.
with Outlines of a Classification of the
Family. Proc. !11rtl. Soc. London. Vol. 29.
1'/s. 2-3.
CRICK~lAY, C. H. (1930): The Jurassic Rocksof Ashcroft. British Columbia.
Bull.
Dept. Geol. Sci .. Unir•. California. Vol. 19.
GoTTSCIIE. C. (1878): Uber jurassische Versteinerungen aus der argentinischen Cordillere. Palaeontogr .. Supple., Bd. 3, Lief
2, Abt. 3.
HAY AMI. I. (1961): On the Jurassic Pelecypod Faunas in Japan. four. Fac. Sci.,
Univ. Tokyo, Sec. 2, Vol. 13.
KoBAYASHI, T. (19-17): On the Occurrenceof Seymourites in Nippon and its Bearing on the Jurassic Palaeogeography.
Iapan. jour. Geol. Geogr., Vol. 20.
(1954): Studies on the Jurassic Trigonians in Japan. Pt. I. Preliminary Notes.
Ibid .. Vol. 25.
.. _ (1956): Some JurassicTrigonians fromCentral and West japan. Ibid .. Vol. 21.
No. 1.
(1957): Some Trigonians from the llida
Plateau Region. Central Japan. Il>id ..
Vol. 28, l•los. J-3.

7

- - and MoRI. K. (1955): The Vaugoniinae
from the Kitakami Mountains in North
Japan. On the Jurassic Trigonians inJapan. Pt. 3. Ibid., Vol. 26. Nos. 1-2.
--and TAMURA, (1955): The Myophorellinae from North Japan. Ibid .. Vol. 26,
Nos. 1-2.
--and-- (1957): Additional new Genera
and Species of Trigonians from theJurassic of Soma. North Japan. Ibid ..
Vol. 28. Nos. 1-3.
LAMilERT, L. R. (1944): Algnus Trigoniasdel Neuquen. Ret•. Mus. de La Plate, N.
S .. Tom. 2, 1•lo. l.J.
l\IAEDA, S. (1952): A Stratigraphical Studyon the Tetori Series of the upper Kuzuryu District in Fukui Prefecture.
]our. Geol. Soc. japan, Vol. 58. i\fo. 68-J.
- - ( 1955) : Biofacies of the Tetori Group.
]our. Colt . •4rt Sci, Chiba Uni11. Vol. I,
lllo. 4.
- - (1956): Stratigraphy and Geological
Structure of the Tetori Group along the
Uchinami and ltoshiro Rivers. Fukui
Prefecture. jour. Gcol. Sor. japan, Vol.
63, No. 7·11.
- - (1961):
Stratigraphy of the Tetori
Group in the Southern Part of the Kuzuryu River, Fukui Prefecture. Ibid.,
Vol. 61. No. 184.
- - (1961): On the Geological History of
the Mesozoic Tetori Group in Japan.
japan. jour. Geol. Geogr .. Vol. 32, Nos.
3--1.

M6RICKE. W. (1894): Verstcinerungen desLias und Unteroolith von Chile. Seues]ahrb. f. Jl.lin. usw., Beii.-Bd. 9.
NAKANO, M. (1961): On the Trigoniinae.
]our. Sci. Hiroshima Urzil• .. Ser. C. Vol. 4.
.1\ro. 1.
PACKARD. E. L. (1921): The Trigoniae fromthe Pacific Coast of North America.
Pub/. Unit• Oregon, Vol. 1. No. Y.
SATO, T. (1960): Etudes Biostratigraphiquesdes Ammonites du Jurassique du Japun.
These Pt·esentee a Ia f<i1culte des Sciencesde Paris.
TA:>.tURA, M. (1960): A Stratigraphic Studyof the Torinosu Group and its Relati\'Cs.
Jl1em. F'ac. Educ., Kumamoto Unil'., Vol. 8.

Trans. Proc Palaeonl. Soc. Japan, N. S .. No . •19. p. 8, Apr. 10, 1963

SHORT NOTE
10.

LILLOE'J'/A SP. (AMMONITE CALLOVIENNE) DE :MITARASf

AU BASSIN DE 'l'ETORI*T ADASHI SA TO
Institut de Geologie, Faculte des Sciences, Universitc de TokioetYASUl\IITU KANIE
Institut de Geologie. Uni\·ersite Nationale de Yokohama

Un specimen ci'Ammonite fragmcntairemais non usc. du diametre d'environs 50
mm. se provient de Mitarashi dans Ia
region de Shiokawa.
II s'agit d'une espece proche de Lil!oetia
bw:kmani (CRICKMA Y). representee par un
tour qui montre les caracteristiques sui·vants: cnroulement trcs involute. ombilictrL~S etroit et profond aVeC Jc bord peu
accuse et doucement convcxe. et costu·
Iat ion flexueuse dans !'ensemble it cotes
primaires attenuces sur Ia partie interne
des tlancs et <t secondaires plus nom·brcuses et plus cll'vees prenant naissancepar Ia bifurcal ion irreguliere des l'Otes
primaires sur le tier interne des Ilanes.
Le genre Lilloetia. exclusivement nord·
pacifique. n·existe que dans le Callovien.
en particulier aux zones
Kepp/erites
tyclwis et K. mrlearni de I'Alaska et deIa Colombie britanique. Ces zones cor·
responderaienl aUX zones cal/01•iunse et
it jason de l'echelle europeene.
L'cxistence d'un niveau callovicn in·fcrieur est done approuvee dans Ia region
de Shiokawa. au contraire Ia notice dervL\EDA tl952) qui n'admet qu'un !;eul
nivcau oxfordien marin pour Ia region.
IWA YA (1940) a signale. d'ailleurs, deux
horizons marins dans Ia meme region etson i\18 (\11 d'aprcs MAEDA) correspond
au ni,·eau du present specimen.

aaText-fig. l. Ulloetia sp.

Ouvrages consultesaY. (1940): On the Tetori Series inthe Vicinity of Makito in thl· Upper
Tributaries of thL· Syokawa in the Pro·
vince of Hida. ]. Geol. Soc. japan. mi.
.J7. 110. 561.
l\IAEDA. S. (1952): A Stratigraphical Studyon the Tetori Series in the l'pper Shio·kawa District in Gifu Pref. /hid .. rol.
.'iS. 110. 679.
lWAYA.

• Received Apr. 19. 1962.

8

Trans. Proc. Palaeont. Soc. Ja1Jan. N. S., No. 49, pp. 9-H. pl. 2. Apr. 10. J!lG:i443.

ADDITIONAL NOTES ON LATE PALAEOZOIC CORALS FOUND
IN THE SOUTHERN PART OF THE TAl\IBA DISTRICT*
SlllGEO SAKAGUCHIandNOBUO Y Al'v!AGIW A
Institute of Geology. Osaka Gakugei University

B&~mm•~-AA~*R~~~~~~~:J'Clf~L t.:. -=t~r"'J 2 f!Jl~:Hfrfii!-c·~7.>"

The writers previously studied thelate Palaeozoic corals found in the southern part of the Tamba District and resulted in describing twelve species insix genera of the Rugosa (SAKA<aJcm.
S. and YAMACIWA, N.. 1958). Later. they
have obtained several new specimens of
the rugose coral during their field sur·vey of the same district. Resides thesematerials. they have been permitted to
study some specimens of coral in thecollection of the Geological Institute of
the Kyoto University and in SI!IMIZU'spossession. In this article is given the
result of the study on the materials mentioned above.
Faunal evidences indicate the upper
palaeozoic rocks of the district to be of
middle to lower Permian age, but there
are very thick non-fossiliferous strata
above and below the fossiliferous ones.
It is, however. thought that the overlying strata indicate an upper Permian
age and the underlying ones. at least
its lower part. extend into the Carboniferous. The fauna of this district is* Received Apr. 23. 1962; read at the 80th
Meeting of the Palaeontological Society ofJapan. November 18. 1961, at Fukuoka. '~~~aL~~00@0~M~~~4ffi~.t)i. 11 ill k$ • 111 !i!il ~ J::.

characterized by a rather abundantfusulinids as well as a considerable
amount of corals.
The stratigraphical sequence and thefaunal zonation have been already reported in detail by one of the writers(SAKAGlJCIII. S .. 1961).
The zonal subdivision of this districtbased on fusulinids is quoted here, to·gether with accompanying corals of eachzone:IV Yabeiua zone

6. lVaagr'IWfthyllum
indicum subzone
III "Veoschwagerina 5. JVeosclm•u~-:erilla r:razone!iculifera subzone
II Parafusuliluz-1. Parcdusuliuazonekuramensis subzone
3. 11fise/lina sp.
subzoneI Pseuciosch·~. Pseudo}itsulina
1cageriua zone
1111/garis subzone
1. Tritici/es mouliparus
subzone

The Tritir:ites montipams subzone
marks the lowest fossil horizon of this
district and includes such as Clisiop/iyllum area (\11:\"ATO}, Dibunophyllum? omorii
SAKAGCCHI and YA~IAGIWA, Stylidop/iyl·
lum sp. indet. (S. kinlliense SAKAGUCHI
and YAt\IAG!WA. n. sp.). and I!eritschioides

10

Shigeo SAI\AGUCJ!I and ;\fobuo Y.-1J/;JGHV.4

sp. indet. As discussed in the previous
paper, the former two species are of
Carboniferous type of the rugose coral
but are found in strata referred to the
Triticites montiparus subzone. The Pseu·
dofusu!ina tmlgaris subzone contains aconsiderable amount of corals together
with leading forms of fusulinid of the
subzone. As for corals of this subzone,
they include Iferitschioides ojensis SAKA·
GliCHI and YAlv!AGIWA, Iferitsc/zioide.<ozakii SAKAGUCHI and Y AMAGIWA. Herit·schioides? sp. indet .. Polytlzecalis? mean·
droides SAKAGUCHI and Y AMAGIWA. Styli·
dophyllum kameokense SAKAGUCHI andY A:\IAGIWA, and Sty!idophyl!um quadratum
SAKACL!CHI and YAIHAGIW ..!,. None ofcoral is reported from the Parajusulina
kuramensis subzone but a coral. genus
and species indetermined, is found in theMise/tina sp. subzone. The Neosclnuag·
erina cratiruhfera subzone yields Huangia
izuru/iensis (SAKAGUCIII andY AMAGIWA)
and Huangia? sp. indet. The T.Vaageno·
phy/lum indicum subzone includes atypical form of Waagenophyllum indicum(WAAGE:\ and WE:-\TZEL). Waagenophyf.
fum tambense SAKAGUCHI and 'I'AMAGIWA,
n. sp .. is found in a limestone immediate
below this subzone. There is anotherrugose coral. Huangia sp. indet .. which
was found by the late Shintaro NAKA·
MURA of the Kyoto University in theWestern Hills of Kyoto, but unknown
to what horizon to belong.
The writers wish to express theirhearty thanks to the following gentle·
men: to Professor Susumu ~vlATSUSHIL\of the Kyoto University of Kyoto. who
permitted them to study the materials
in his institute, and also to Mr. Teruo
SHIMIZU, the president of the Nippon
Chikagal<usha of Kyoto. for his kindoffer of corals in his collection.

Description of Coral Species
Genus Stylidophyllmn
FROMENTAL,

1861

Sty/idophy/lum llinlliense SAKAGUCHI
and YA:\1AGIWA n. sp.
PI 2. figs. Ia. lb. 2.

Corallum compound, massive; cornposed of prismatic corallites. polygonal,
usually five-sided in transverse section.
Corallite usually 7-9 mm in diameter oftransverse section in mature stage. External wall thick and prominent, bearing coarse denticles on all sides, whichcorrespond to septa in number. Axial
structure less than 2 mm in diameter :composed of a few septal lamellae andaxial tabellae. Median plate relatiYely
distinct. Septa of two orders. major andminor alternating. 1\lajor ones moderately thick, 17 to 22 in number andalmost reach axial structure ; counter
septum connecting with median plate.
l'v1inor ones thinner. usually one-half to
two-thirds length of major. Septa slightly sinuous. thick in medial area, being
gradually thinner towards axis. and mostof them disappearing into meshes ofvesicles towards periphery. but someconnecting with denticles of external
wall.
Dissepiments arranged in lonsdaleoid pattern showing rather regularcysts. all convex towards axis.
In longitudinal section. tabularium
almost occupied by axial structure. but
sometimes short horizontal tabulae obsen·ed. Tabellae more steeply ascend·
ing to axis.
Dissepiments relatively
long. slightly curved, their convex sides
facing inwards and upwards.
Comparison :-This species has beenreported as StylidojJ/iyllmn sp. indet. in
the previous paper (SAKAGUCIII and Y A-

44,'1.

11

Late Palaeozoic Corals

MAGIWA, 1958, pp. 174, 175. pl. 4. figs. 5a,5J), based on a single specimen. Recently, more numerous specimens havebeen obtained, so a new specific nameis proposed in this paper.
The general structure of this species
shows close relationship to OzAWA's
Lonsdaleia jlor((ormis crassiconus McCoY(OZAWA, 1925. p. 69. pl. 8, fig. 1) fromthe Akiyoshi Limestones of Nagata,japan. Since no longitudinal section is
illustrated in his paper. these two species
can not be compared in detail. It differsfrom Stylidophy/lum floriformis crassiCOIIUS (l\lcCOY) (SMITH. 1915, pp. 20. figs.
3-10) of England in having no distinct
tabula in the longitudinal section andmore loose axial structure in the transverse section. This species closely resembles Stylidop/zyllum fioriformis floriformis (em. tVIARTIN) (SMITH. 1915, pp.
2·17-251. pl. 19. figs. 1-5) from the Lower
Carboniferous of England in the general
structure oft he transverse section. These
two species. however. can be distinguishable in the following respect: theformer has no distinct tabula in thelongitudinal section but the latter has
distinct long horizontal ones. Finally,
it can be differentiated from Stylidop/zyllum yokoyamai (OZAWA 1925, p. 112, pl.
8, figs. 5, 6; MINATO, 1955. p. 134, pl. 41,
fig. 2; pl. 43. fig. 5) from Nagato. japan
by its marked development of the lonsdaleoid dissepiments in the transverse
section.
Locality and llorizon :-Specimens werecJIIected by Teruo SHIMIZU from the
Kannontoge Limestone referred to the
Triticites montiparus subzone in the
Sonobe Formation. Sonobe-cho. Funaigun, Kyoto Prefecture. Lower Permian.
Repository :-Deposited in Geological
Institute, Osaka Gakugei University.
Reg. nos. !GOG 62002a (holotype), 62002b(holotype). 6200,1.

Genus IVaagenophyllum
HAYASAKA, 1924
TVaagenophyllum tambense SAKAGUCHI
and YAMAGIWA, n. sp.
Pl. 2. figs. 3a, 3b.

Corallum fasciculate. composed of cylindrical corallites. Corallites usually 4
to 7 mm in diameter in transverse section. External wall relatively thin.
Septa of two orders, major and minor
in alternation. Sometimes tertiary septa
present. Numbering 22 to 28 for major
septa and as many for minor ones, bothbeing more or less sinuous. Major septa
very thick at proximal ends and thin
distally, mostly reaching near axial
structure. Minor ones usually half to
two-thirds length of major and relatively
thinner. Dissepiments mostly arranged
concentrically. Axial structure small insize, with a width of about one-fourth
to one-sixth diameter of corallites. composed of axial tabellae and septal lamellae. l'v!edian plate relatively distinct.
In longitudinal section, tabularium
entirely occupied by axial structure;tabula absent. External part of dissepimentarium composed of small vesicles.
facing upwards and inwards. Internalpart of dissepimentarium composed ofelongate, slightly curved vesicles facing
inwards or inwards as well as upwards.
Axial structure composed of steeply
ascending tabellae and a distinct medianplate.
Tabularium about one-fourth
width of corallites.
Comparison:- This new species is closely related to Waagenophyllum alwsakensis(Y ABE, 1909. p. 4. text-fig. 1 ; Y ABE and
HAYASAKA, 1915, p. 100; OZAWA, 1925,
p. 75. pl. 14, figs. 5, 6; SMITH, 1935, p.
36; M!NATO, 1955, pp. 1011. 105. pl. 37,
figs. 6, 7) described from the 1\'liddle

S/u:ueo SAKAGUCHI and Nobuo 1".4JI.4GIWA
Permian at many places of Japan in sizeof corallites and number of septa. It
differs. however. from the latter in hav·
ing shorter minor septa, tertiary septa
and no tabula in longitudinal section.
lr closely resembles Waagenop/zyllum in·dirum (WAAGEN and WENTZEL, 1866, pp.
897-900, pl. Cl, figs. l-3, pl. CXV, figs.
3. 4; YABE and 1-IAYASAKA, 1915, pp. 97.
98; S~ttTII, 1935, p. 32. pl. 8. figs. 1-6::-.1 1!\A TO. 1943. pp. 52-54, pl. l, figs. 1-4 ;:\•liN A TO. 1955. pp. 102, 103, pl. 21, figs.
1. 2; pl. 26. figs. 2. 4-6. 8; SAKAGUCHI
and YAMAGJWA. 1958, pp. 175. 176, pl. 5.
figs. 4-7). but differs from the latter inhaving tertiary septa. It also resembles
Waagenop!tyllum nngamie
Y AMAGIWA
UviS) from the Atetsu Limestone. Oka·yama Prefecture. Japan. The difference
between the above two species is shownby smaller corallites and smaller axial
structure of llw Tarnba form. and more·
over it has tent-shaped axial tabellae
instead of dome-like ones af the i\tetsu
specimen. It is distinguished from TVaagenop!tyllum po/yseptata 1\II:-.IATO (1955.
pp. 105-107. pl. 21. fig. 3) found in the!\Iiddle Permian of the Kitakami ·Moun·tainland, Northeast Japan in the follow·
ing respects; the present species has
smaller corallites, less numerous septain transverse section, and has no tabulain longitudinal section.
Locality and llorizon :-Specimens werecollected by Professor Susumu .MATSU·
SIIITA of the Kyoto University in a limestone quarry east of Ogonjo. Oharano.
Ukyo·ku. Kyoto City. This limestone
occurs immediate below a limestoneyielding IVaagenop!tyllum indicum (WA·AGEN and \VENTZEL) of the Yabeina zone?
in the upper part of the lzuruha For·mation. Middle Permian.
Repositm:v :-Deposited in Geologicaland Mineralogical Institute. Universityof Kyoto. l{eg. nos . .I PC10030a (holotype),

40030b (holotype).
Genus Heritscl1ioides YABE, 1950

Ileritschinides sp. indet.
Pl. 2. figs. 4a, 4b.

Corallum fasciculate, composed of cy·lindrical corallites.
Corallites usually
11 to 13 mm in diameter in transverse
section. Septa of two orders. major andminor in alternation. Numbering 20 to
23 for major and as many for minor
ones. both being straight or slightly
sinuous. ivlajor ones mostly reach axial
structure. thick in medial area and thin·
ner towards both ends; some of themconnecting with septal lamellae. !\tinor
ones thin, usually one·half to two-thirds
length of major. Dissepiments arranged
in concentric or angulo·concentric pattern. Axial structure occupies a spaceof about one·third diameter of corallite.
1\Iedian plate indistinct.
In longitudinal section, dissepimentarium composed of small vesicles with
their convex sides inwards. Tabulae
incomplete. composed of small vesicles
with their convex sides upwards andoutwards, steeply ascending to axial
structure. Tabellae composed of small
Yesicles with their convex sides outwards as well as upwards, steeply ascending to axis. Median plate indistinct.
Tabularium occupies two-thirds of entire
space of corallite. Axial structure about
one-half width of tabularium.
Compm·isou:- The present species
resembles Heritschioides ojensis SAKAGUClll and YA:O.IAGIWA (1958, pp. 170. 171,pl. 1. figs. '1-6) from Oji and Heril·
scltinides ozakii SAKAGCCIII and YAC\IA·
GIWA (1958. pp. 171. 172, pl. I, figs. 7-9;
pl. 3. figs. 1. 2) from lnukanno, both at
Kameoka City, Kyoto Prefecture.
It
seems to indicate an intermediate form

443.

J,ate Palaeozoic Comls

between the above two species in trans·verse section. but can be distinctly discriminated from them in longitudinal
section. Therefore, if better preserved
material is obtained in future. the pre·sent species will be able to be proposed
as a ne\v species.
As far as the general structure ofcorallite in transverse section concerned,
the present species and Heritsclzioitks
ojensis are almost nearly indistinguish·
able. In longitudinal section, howe\'er,
it has more steeply ascending tabulaethan those of the Oji form. Tt also differs from 1-leri/schioides ozakii in larger
axial structure of the former and inhaving steeply ascending tabellae instead of rather dome·likc ones of thelatler in longitudinal section.
Locality and Horizon :-Specimen was
found by Naoya NASHIKI of the Higashi!\·Iiddle School. lbaragi City in an oldquarry near l(onzoji, Oharano. Ukyo·ku.
Kyoto City.
The limestone of thisquarry is referred to the Trilicites nwu·
tiparus subzone in the lower part of theTano Formation in the Western Hillsof Kyoto (Kyoto-I\ishiyama!.
Lower
Permian.
Repository .·--Deposited in Geological
Institute. Osaka Gakugei lni\-ersity.
Reg. nos. !GOG 62004a. 62004b.
Genus 1/uangia YAHE, 1950
Hua11gia sp. indet.
Pl. 2, figs. 5a. 5b.

Corallum fasciculate. composed ofcylindrical coral lites. usually 7 to 10 mm
in diameter of transverse section in mature stage. i'v!ajor septa 17 to 22 in
number. alternating with an equal number of minor ones. ~V!ajor ones slightly
thick in medial area. thinning bothends. reaching axial structure: some of

1 •:>

•)

them connecting with septal lamellae.
l\linor ones thinner, less than half lengthof major. Septa almost connected withexternal wall. but sometimes disappearing into meshes of vesicles of peripheralarea. Axial structure loose. composedof septal lamellae and axial tabellae.
Median plate usually lacking: it resembles that of the genus Sip/10nodendronin younger stage. Dissepiments arranged
concentrically in younger stage, but
becoming pseudoherring bone pattern
in mature stage.
In longitudinal section. dissepimentarium composed of vesicles with theirconvex sides facing upwards as well as
inwards. Outer tabularium broad, composed of complete or incomplete horizontal tabulae. Tabellae show dome-likestructure. Tabularium occupies· threefifths of entire space of corallite. Axial
structure one-third width of tabularium.
Comparison :-Jt is reported that thisspecimen was collected by the late Shintaro NAKAIV!URA ncar lzuruha. Takatsuki
City. Osaka Prefecture. Howe\'er. the\\Titers' recent studies show that there
is absent such a black limestone
contained the present specimen nearlzuruha but present near Kurama in the
Northern Hills of Kyoto. For this reason. if it were to be obtained in any
other places than lwruha, it would befound in a black limestone belonging to
the Mise/lina sp. subzone at Kurama.
This species closely resembles Huangia!wshimoloi (NAGAO and MJ:-:ATO. 1941.
pp. 102-105. pl. 27. figs. 1-5: Mll\ATO.
1955. pp. 123-125. pl. 2. figs. 1-3; pl. 22.
figs. 3. 4. 6'1 in the general structure of
the transverse section. but the former
is smaller in size and fewer in numberof septa.
Locality and Horizon :-According toSusumu MATSUSHITA of the Kyoto University, this specimen was collected by14

Shigeo SAKAGUCHI ancl Nobuo YAMAGilVA

Shintaro NAKAMURA near Izuruha.
Takatsuki City, Osaka Prefecture but
the exact locality ramains unknown.
Probably Middle Permian.
Repositm)':-Deposited in Geologicaland Mineralogical Institute, Universityof Kyoto Reg. nos. J PC 4003la, 40031 b.

References
MtNATO, M. (1943): Notes on some Permian
Fossils from the Toman Formation inSoutheastern Manchoukuo. four. Fac. Sci.,
Hokkaido Imp. Uni1• .. Ser. 4. Vol. 7, No.
1, pp. 49-58. pis. 1, 2.
- - (1955) : Japanese Carboniferous andPermian Corals. Ibid .. Vol. 9, No. 2, pp.
1-202, pis. 1-43.
NAGAO. T. and Mt:>~ATO. l\l (19·13): Cortcenia
haslzimoloi, a new Tetracoral from theUpper Palaeozoic of Sikoku. Ibid., Vol. 6.
No. 2, pp. 101-105, pl. 27.
OzAWA. Y. (1925):
Palaeontological andStratigraphical Studies on the PermoCarboniferous of Nagato. Part 2, Palaeontology. four. Coli. Sci., Imp. Unit•. Tokyo,---------

Vol. 45, Art. 6, pp. 1-90. pis. 1-1·1.
SAKAGl:CIIt, S. (1961): Stratigraphy andPalaeontology of the South Tamba District. part. 1. 1\Iem. Osaka Unil•. Lib. Art& Educ., B. Nat. Sci., l•lo. 10, pp. 35-76.
- - and Y AMAGIWA, N. (1958): The Late
Palaeozoic Corals from the Southern Partof the Tanba District. Ibid., No. 7, pp.
163-178, pis. 1-5.
SMITH. S. (1915): The Genus Lonsdaleia and/Jilnmophyllum rugosum (McCoY). Quart.
four. Geo/. Soc .. Vol. 71. pp. 218-272, pis.
17-21.
- - (1935):
Anthracolithic Corals from
British Columbia and related Species
from the Tethys. ]mtr. J>aleonl., Vol. 9,
pp. 30 -42, pls. 8, 9.
WAAGEN, \V. and WENTZEl., J. (1886): Salt
Range Fossils. Palaeonl. Indica. Ser. 13,
Vol. 1, pp. 835-924. pls. 97-116.
YABE. H. (1909) : Materials for a Knowledgeof the Anthracolithic Fauna of Japan.
four. Geol. Soc. Tollyo, Vol. 9, pp. 1-5.
- - and IIAYASAK.'<, I. (1915-1916): Palaeozoic Corals from Japan. Korea and China.
four. Geol. Soc. Tokyo. Vol. 22. pp. 55-70,
79-109, 127-142, vol. 23, pp. 57-75.
--------

Explanation of Plate 2
Stylidopflyl/um kinkiense SAKAGt:CHI and YA~IAGIW.o\, n. sp.
Figs. la-b. 2 .... x3.5
la. Transverse section of the holotype (Reg. no. IGOG 62002a)lb. Longitudinal section of the holotype (Reg. no. IGOG 62002b)
2. Transverse section (Reg. no. IGOG 62003)
Locality: Kannontoge, Sonobe-cho, Funai.gun, Kyoto Prefecture.
l¥aagenoph.)'flrmz tambeuse SAt<AGUCHI and YA:-.'IAGIWA, n. sp.
Figs. 3a-b .... x4.0
3a. Transverse section of the holotypc (Reg. no. JPC ·40030a)3b. Longitudinal section of the holotype (Reg. no. JPC 40030b)
Locality: Ogonjo, Oharano. Ukyo-ku, Kyoto City.
/Ieritscflioides sp. indet.
Figs. 4a-b .... x3.5
4a. Transverse section (Reg. no. !GOG li2004a)4b. Longitudinal section (Reg. no. IGOG 620064b)
ll11angia sp. indet.
Figs. 5a-b .... x35
5a. Transverse section (Reg. no. ]PC 4003la)5b. Longitudinal section (Reg. no. ]PC 4003Ib)
Locality: ? Izuruha, Takatsuki City, Osaka Prefecture.

SAKAGUCHI and YAMAGIWA :lsHll and AoKr photo.

Late Palaeozoic Corals

Plate 2

Trans. Proc. Palaeont. Soc. Japan. N. S., No. 49, pp. 15-24, pis. 3-4, Apr. 10, 1963

H4.

LOWER CARBONIFEROUS BRYOZOA FROl\1 THE
OMI LIMESTONE. JAPANPART 2: SUCCESSIVE DESCRIPTIONS OF CRYPTOSTOMATA*
SUr-.110 SAKAGAl'vll
Department of Geology. Hokkaido Gakugei Universityt-:..

i'riftH51iR!ftr>Vr i'ffi1'T~*Cfl'!.1Ht.::U" .:ttl) 2: .:CO) 1 1..:.~~ '-cir.;J.II H 8 ~~ 13 flll~~;t!Jf.H..
;!tj[ J: ~ 1:;

In this paper. the writer describes
thirteen species in eight genera as the
successive study of the lower Carboni·ferous Bryozoa from the Omi limestone.

boidal to obliquely oval at middle levelof branch. Nodes observable. about 0.03
mm in diameter. but their arrangement
uncertain. Stereom of both sides com·posed of fine fibrous striations and closely
arranged fine granules.
Meshwork formula :-10/8/ /17-18/·l* (*4:number of rows)
Remarks:-The present form is nearest
to Polypora ldworot,ae SrrUI.GA·NESTER·
ENKO (1955) which was reported fromthe Kashinskii formation of the middle
Carboniferous. i{ussian Platform in themeshwork formula (11-12(10)/8-9//18/45). but differs by the fan-shaped zoarium
instead of the parallel branched zoariumof the latter. In the general characters,
the present form also resembles Po!ypora
bassleri which CoNDI<A 11902) reported
from the Coal Measures of Nebraska.
the United States of America. However.
the present form can be distinguishedfrom P. bassleri by the narrower widthof branches, wider fenestrules and 17 to
18 zooccia per 5 mm of branch insteadof 19 to 20.
The specific name is dedicated to Mr.
Hiroshi FUJITA who was recently en·
gaged in the stratigraphic survey of theOmi limestone.
Locality :-Nishiyama. Reg. no. 11218A tholotype).

Genus Po!ypora 1\I'CoY. 1844

Po!ypora fujitae SAKAGA:\!1, n. sp.
Pl. 3. tlg. l.

A single tangential section of zoariu111.
fan-shaped. frequently dichotomosing.
consists of nearly straight branches con·nected by dissepiments at regular inter·vals.
Branch 0.52 111111 to 0.60 111m in
width and about 10 in 10 mm horizontal.
Dissepiments strong, about 0.36 mm inwidth. Fenestrules ellipsoidal or rhom·boidal with rounded corners in outline.
their width and length about 0.60 mm
and 0.72 mm to 0.92 mm. respectively:
about 8 in 10 mm length of branch.
Zooecial apertures circular in shape. ar·ranged in alternating longitudinal series.
usually 4 rows on each branch. 17 to 18
in 5 m111 length of one row. and usually/1 apertures per fenestru le. Diametersof apertures 0.08 mm to 0.10 mm andinterspaces between zooecial apertures
0.16 mm to 0.21 m111. Zooecial tubes rhom·• Received Apr. 30. 1962: read June 2. 1962.
]i)

Su mio SAK.4G.tl1Hl](}Genus Pemziretepom D'OJmJG:\Y, 1819

Penniretej,ora cf. irregularis(NEKHOROSHEV)
Pl. 3, fig. 5: text-fig. I.
1948.

Pinnatopora irregu/aris NEKHOROSIIEY.
:lkad. Nauk Kaz. SSR , p. 47. pl. •1, fig. 2.

Two tangential sections. One of thespecimens t)l2l4i, zoarium consists ofbroad straight main branch 1.00 mm to1.10 mm in width. Lateral branches 0.60
mm to 0.68 mm in width. extending al·
ternately at variable angles ('15' to 100')and at interYals of 0.-10 mm to 0.52 mm
with 4.5 branchlets in 5 mm. Zooecial
apertures circular. their diamters 0.10
mm. Zooecial tubes with well developed
hemisepta. kidney-like shaped at middle
level of tangential section. about 13 in
5 mm length of one row, and spaced
regularly in 3 per interval between
lateral branches. Interspaces between
zooecial apertures 0.21 mm to 0.2·1111111
longitudinal. Single straight carina andsome nodes on carina probably present,
but indistinct. Inner part of stereom
composed of rather coarse fibrous tissue.
but the outer part of very fine granules.
Another specimen (11259) shows only
a few zooecial arrangement but thestructure of stereom covered on reverse
side is well observed. Width of mainbranch and lateral branches 1.10 mm andabout 0.80 mm to 1.00 111111, respectively.
Lateral branches extending alternately
at an angle of about 75' and at intervalsof about 0.60 mm with 4 branchlets in 5mm. Stereom composed of many small
spicules arranging in longitudinal rows.
and interspaces between one and adja·
cent rows of spicules about 0.05 mm.
Remarl~s :-The present form agrees
with Penniretepom irregularis (NEKHOIW·SIIEV) which was reported from the mid·die Carboniferous of near I3alkhashia in

Text-fig. 1. l'nmireiepora cf. irre·
gularis (N'EKJJOIWSHEV), tangential
section (Reg. no. 11214).

the general appearance except for thewidth of main branch of the present
form which is wider than that of NEK·IIOROSHEV's specimen. However. NEK·IIOIWSIIEV's description and illustration
are not su!Iicient to compare with thepresent specimens because the detailed
structure of stereom and arrangementof spicules are not shown in his illust·
ration. The angle between the mainbranch and lateral branches is about 60"
in NEKHOIWSHEV's specimen instead ofvariable angles I 15" to lOW) in the pre·sent form.
Locality :-Nishiyama. l~eg. nos. 1121-1,

11259.
Pi'nui rC'lcpora higashiyamensis
SAKAGA:\11, n. sp.
Pl. :l, fig. :l: text .fig. 2.

A single tangential section. Zoarium
consists of broad. straight main branch,
0.91 mm in width. Lateral branches 0.52
mm in width. extending alternately atan angle of 70" and at intervals of 0.78
mm to 0.88 mm, with 4 branchlets in 5mm. Zooecial apertures circular. theirdiameters about 0.10 mm. Zooecial tubes

4H.

1J1·yozoo. from the Omi Limestone. Pt. 2

with well developed hemisepta. elongatedellipsoidal at middle leYel of tangential
section. about 13 in 5 mm length of one
row. and spaced regularly. 3 per interval
between lateral branches. lnterspaces
between zooecial apertures 0.24 mm to0.28 mm longitudinal. Stereom composedof many small spicules arranged in long·itudinal rows. and interspaces betv;een
one and adjacent rows of spicules about
0.05 mm. lntcrspaces between spicules
in one row somewhat \'ariable. 0.05 mm
to 0.08mm.

17

Penni-retepom regularis SAKAGAMI,n. sp.
Pl. 3, fig. 4: text-fig. 3.

A single tangential section. Zoarium
slender, pinnate. straight midrib. Widthof midrib 0.45 mm to 0.48 mm. Lateralbranches about l mm long and 0.32 mm
to 0.37 mm wide. extending alternately
at angle of 80 degrees and at intervalsof 0.45 mm to 0.56 mm with 6 branchlets
in 5 mm. Shape of zooecial aperture
not observed. Zooecial tubes arranged
in alternating longitudinal series. trape·
zoidal or triangular with rounded cor·ners at base and middle levels of tangential section. about 12 in 5mm lengthof one row. and spaced regularly in pairs
per interval between lateral branches.
Relatively thin stereorn covering zoa·
rium. composed of fine fibrous tissue
with spicules.
Thickness of stereom
about O.lO mm. Carina and nodes notobserved.

Text-fig. 2.
Pemziretepora lti[!ashiyameusis SAKAGA~II. n. sp .. tangential
sect ion. holulype (Reg. no. 11285).

Remarks:- The present form agrees
with Pemziretepom cf. irregularis (l\EK·
HOROSHEV) which was described in thispaper in the general measurements.
However. the difference between the two
species is in the arrangements of spi·
cules. The spicules of the present
species is arranged more coarser thanthat of P. cf. irregu!aris (NEKIIOIWSHEV).
Lomlity :-lligashiyama. Reg. no. 11285: holotype).

Text-fig. 3.
Penuiretepora regularisn. sp.. tangential sedion,
holotype (Reg. no. 11215).

SAI<AGAMI,

Remarks :-Slender and straight mainbranch with regularly di\·ided lateral

18

Sumio S.·lKAG.4Jll

branches. strongly alternated arrange·
ment of zooecial tubes and very thickwall between zooecial tubes are the important characters of the present species.
The present form agrees with Pennirete·
pam taxa var. kasallhstanica NEKIIOROSHEV {1948) which was reported fromthe middle Carboniferous near Balkha·
shia in the width of main branch. num·
ber of lateral branches in 5 mm (6) andnumber of zooecia in 5 mm of mainbranch (12-13). but it can be distinguish·
ed therefrom in the angles between
main branch and lateral branches (80'instead of 50'-55"), and by the thicker
wall between zooecial tubes.
Locality :-Nishiyama. Reg. no. 11215(holotype).
Pe111ziretepora sp. i ndet.
Pl. 3, fig. 2.

A single tangential section. Zoarium
consists of broad. straight main branchabout 0.8 mm in width with short lateralbranches. Lateral branches about 0.45mm in width. extending alternately alan angle of about 90 degrees and atintervals of 0.56 mm to 0.72 mm with
4.5 branchlets in 5 mm. Shape of zooeci·
a! aperture not observed. Zooccial tubes
arranged in alternating longitudinal
series. oval or rounded polygonal at baseand middle levels of tangential section.
Hemisepta not observed.
About 17
zooecia in 5 mm length in row and spaced
regularly in 4 per interval bet ween
lateral branches. Stereom relatively
thin. Carina and nodes not observed.
Remarks :-Only a few species of Pe1lniretepom having four apertures in the·interval between the lateral branches
have been described from the Carboni·ferous strata of the world. The present
form may be a new species, but the

specific identification is not made becauseof the ill-preserved section of specimen
and of insufticiency of the material at
hand.
Locality :-Nishiyama. Reg. no. 11239B.
Genus Protoretepora DEKO:-IINCK. 1876
Protoretepom hayasakae SAKAGAMI,n. sp.
Pl. 3, figs. 8, 9. •? 1924.

Phyl/opora sp., llAY:\SAKA. Sci. Rep.,
Tolwku Imp. Uuiz.•., Ser. 2, Vol. 8, No.
1, p. 56. pl. 7. fig. 6.

Zoarium flattened, forming reticulate
expansion.
consisting offlexuousbranches and broad celluliferous dissepi·
ments. Width of branches usually 0.80
mm to 1.20mm but occasionally 060mm.
and 6 to 8 branches in 10 mm horizontal.
Fenestrules circular or oval. their widthand length 0.60 mm to 1.20 mrn and 0.70
mm to 1.20 mm. respecti\'ely; 5 to 8
fenestrules in 10 mm vertical. Zooecial
apertures arranged in alternating longitudinal series. usually 4 to 5 rows oneach branch. Zooecial tube roundedrhomboidal at base and middle levels oftangential section. Zooecial apertures
without peristomes. circular and theirdiameter 0.14 mm to 0.16 mm. Distance
between zooecial apertures along longi·
tudinal row 0.16 mm to 0.20 mm. Dis·
sepimcnts consist of same rows of zooe·
cia as those of branches. but in places.
division into branches and dissepiments
indisti net. On reverse surface. bothbranches and dissepiments rounded andsmooth. but slightly convex at dissepi·
ments. Thick stereom covering reverse
side, but becoming very thin at obverse.
Remarl?s :-Many species of Protm·ete·pora have been known from the Permianof Australia. Salt Range of India and4-H.

Bryozoa !1·om the Omi Limestone. Pt. 2

the other localities under the genericname of Plly!lopora. From the Carboniferous. however. only a few species of
the genus may have been described.
The present form can be easily distinguished from the species previously
described except for Phyllopnm sp. which
HA YASAKA described and illustrated
from an unknown locality of the Omi
limestone. The present form resembles
HAYASAKA's specimen in the meshwork
measu rem en ts.
The specific name is dedicated to Dr.
IChiro IIA YASAKA who is a pioner of the
Paleozoic Bryozoa in .Japan.
Lnca!ity:-Nishiyama. Reg. no. 11279(surface, holotype). 11233. 11265.
Genus Rhabdome.wm YOUNG
and YOUNG, 1874

R!wbdomeson yabei SAI<AGAt\11,
n. sp.
Pl. 3. fig. 7.

Zoarium consisting of straight. slender
cylindrical stem. about 2mm in diameter.
not observed to branch. Central tube
0.32 mm in diameter. Thickness of mature zone about 0.50 mm. Zooecial apertures regularly arranged in longitudinal
series. 3.5 in 2 mm; and diagonally intersecting one another. 5 in same distanceand longer diameter 0.16 mm to 0.19 mm
and shorter one 0.10 mm to 0.13 mm.
Zooecial tube making an angle of 30' tocentral tube in immature zone, but be·comes perpendicular to surface in rna·
ture zone; oval in tangential section of
mature zone. \Vall thickened in maturezone. distinguishable from very thin
immature wall. lnterspaces between
zooecial tubes in mature zone 0.32 mm
to 0.40 mm in longitudinal row, and0.16 mm to 0.19 mm in diagonal row.

19

Superior hemisepta poorly developed
at posterior end of mature zone. andinferior hemisepta disposed at opposite
side of slightly inner parts of superior
hemisepta. Two kinds of acanthopores-mcgacanthopores usually disposed at
distal edge of each zooecium, micracanthopores around zooecial tube arranged
in one row. Diaphragms not present.
Remarks :-Almost all of the species
belonging to the genus Rhabdomesonfrom the Carboniferous rocks ha\·e morelarger diameter of zoarium than thepresent form.
The present form isnearest to Rltabdomesmz rhombijerum
PHTLLIPS which was reported from the
middle to upper Carboniferous of theseveral localities in Soviet Russia in the
diameter of zoarium and some characters of internal structures. However,the present form can be distinguishedfrom R. rhmnbifemm by the weaker developed hcmisepta and larger micracanthopores.
Recently, ELIAS (1957) described three
species of transitional forms between
the genera Rhabdomesmr and Rhombo·
pora. an<! at that time. he stated that"axial tube in this genus could not
possibly be originally hollow, but wasfilled with some kind of soft or liquidorganic substance, and its occasional
filling with surrounding matrix musthave been a secondary phenomenon."The axial tube of the present form isalso filled with the same material as thesurrounding sediments.
The specific name is dedicated to Dr.
Hisal<atsu YABE, the father of paleontology in Japan.
Locality :-Nishiyama. Reg. nos. 11202(holotype). 1120l.
Genus Rlwmbopora J\IEEK. 1872
Rlwmbopom sp. indet.
PL 3. fig. G.

Sunu·o S.4K.:!G.-lMI

20

A single oblique section. Zoarium acylindrical stem. about 1.6 mm in dia·meter. Zooecial tube bending gentlr
outward in longitudinal section. oval,
somewhat irregularly near surface oftransverse section. its diameter 0.16 mm
to 0.18 mm. and shorter one 0.11 mm to0.13 mm. Zooecial tube probably ar·ranged in longitudinal intersecting rows.
Zooecial wall thin in immature region,
becoming thick gradually to surface.
Near surface, interspaces between zooe·
cia! tubes 0.06 n:m to 0.08 mm in long·itudinal trend; 0.02 mm to 0.03 mm indiagonal trend.
One megacanthopore
porous. surrounded by concentric dark
fibrous tissue, usually present in eachpoint of intersection of zooecial walls.
Diameter of megacantbopore measured
from edge to fibrous tissue 0.040 mm to
0.048 mm and its inner diameter 0.006mm
to 0.008 rnm. One or two micracantho·~ores in one row in each interspace of
megacanthopores.
l\Iesopore and dia·phragm lacking.
Remarks:-The present form differsfrom the preyiously described species.
The megacanthopore with large pore, nodiaphragm and no hemisepta are thecharacters of the present species.
Locality :-Higashiyama. Reg. no. 11294.

Genus Streblascopora !3ASSLER. 1952

Strebfascopora amabilis SAKAGA:O.II,n. sp.
Pl. 4, figs. 1-3.

Zoarium slender and ramose. its dia·meter 1.0 mm to 1.2 mm. Central bundleof small parallel tubes surrounded byzooecial tube, circular. and 0.24 mm to0.29 mm. Number of tubes in central
bundle 10 to 20 in transverse section. 4

to 5 rows obsen·ed in longitudinal sec·tion. Zooecial apertures regularly ar·ranged in longitudinal series, 7 in 2 mm,
oval. and their longer diameters 0.11 mm
to 0.13 mm and shorter ones 0.08 mm to0.11 mm. Zooecial tube making an angleof about 30" to central bundle in imma·
ture zone, but tube near opening becomes perpendicular to surface. Inter·spaces between zooecial tubes in maturezone 0.19 mm to 0.26 mm in longitudinalrow. Thickness of mature zone 0.16mm
to 0.24 mm. Mesopores straight. arise
at right angles from base of matur'e
wall. usually 2 rows with 2 to 3 in eachrow longitudinal, one mesopore dis·posed at distal edge of each zooecium
between ordinary rows. and total number of mesopores disposed between zooe·
cia in one row usually 5 or 7. Diameterof mesopore 0.24 mm to 0.29 mm. Sup·erior hemiseptum poorly developed at
posterior end of mature zone, and welldeveloped 2 or 3 inferior hemisepta de··veloped at opposite side of superior
hemiseptum at about same distancesfrom each other.
Remarl<s :- Abou L a dozen species of
the genus Streblascopora have been described from the Permian rocks but they
were reported as of the genus Streblo·
trypa except for two or three species
recently described. On the other hand,also in the Carboniferous rocks. a con·siderable number of species describedas Streblotrypa should be referred to thegenus StreblascoPom. At least, the fol·lowing two species. Streblotrypa densa
MOROZOVA (1955) and Strebloli)'pa uiki·
forovae MoROZOVA (1955). which haveapparenlly the central bundle can be
included into the genus St1·eblasropora.
Some other species including Streblotrypa
may now also be expected to belong toStreblascopora by the reexaminations ofinternal structures.

H4.

Rryozoa from the Omi Limestone, Pt. 2

The present form re3embles Streblas·
copora dellsa (ivlOROZOVA} which was re·ported from the Gzhelian Stage (Triticites
stuckenberKi zone) of Lhe mid-valley of
the Don in its general appearances. but
it can be distinguished by the morestrongly intercalated zooecia in longitudinal series and smaller size of mesopore.
Locality :-Nishiyama. Reg. nos. 11272(holotype), 11205. 11212, 11216-B. 11244A, 11256. 11267-B. 11271-B.

Streblascopora grossa SAKAGA:\11,
n. sp.
Pl. 4, figs. -1. 5.

Zoarium consisting of cylindrical stem,
its diameter 1.7 mm to 2.0 mm, not observed to branch. Proximal part spreading to attach with foreign substance.
Central bundle of small parallel tubes
surrounded by zooecial tube. circular,
about 0.30 mm in diameter. Number oftubes in central bundle about 7 to 10 intransverse section. usually 3 rows observed in longitudinal section. Zooecial
apertures regularly arranged in longitudinal series. but in proximal part, seemto be arranged rather irregularly. Number of zooecia in 2 mm cannot be counted. Zooccial tube ncar surface oval atordinary part. its larger diameter 0.12
mm to O.l6mm and smaller one 0.06mm
to 0.08 mm; circular at proximal part,its diameter usually larger than that ofordinary part, 0.09 mm to 0.13 mm.
Zooecial tube arises from cemral bundleat an angle of 20, to 30" and becomes
perpendicular to surface.
Interspaces
between zooecial tubes in mature zone
about 0.2•1 mm. Wall of adjacent zooecial tubes in immature zone separated
by a dark line. Thickness of maturezone 0.48 mm to 0.56 mm. l'v!esopores

21

straight, arise at right angles from baseof mature \Vall. and regularly arranged
in two rows with 3 or more in each row
longitudinal: but rather irregularly arranged at proximal part.
l\lesopore
variable in form and size. circular toelongated elliptical, 0.016mm to 0.0-!0mm.
Superior hemiseptum poorly developed
at posterior end of mature zone. andwell developed inferior hemiseptum disposed at opposite side of inner part ofsuperior hemiseptum.
Remarks:-The present form is characterized by the relatively large diameterfor the genus. nearly straight and longzooecial tubes in mature zone. rather
narrow central bundle and wall of theadjacent tubes being separated by adark line in immature zone. The present form is similar to Streblascnpom
linea/a SAKAGA:\Il and AKAGI i 1961)
which was reported from the lower
Permian of the Taishaku Plateau, Japanin the wall structure of immature zone,
but differs by the other character above
mentioned.
Locality :-Nishiyama. Reg. no. 112-HA (holotype); lligashiyama, Reg. no.
11285.
Genus Streblotrype/la NIKIFOHOVA, 19 118

Streblotrype!la? sp. indet.
Pl. I, fig. 6.

A single longitudinal section of a fragment of branched zoarium. Diameter ofzoarium 1.8 mm. Zooecial apertures may
be regularly arranged in longitudinal
series. Zooccial tube near surface circular or oval. and its diameter about 0.01
mm to 0.11 mm. The tube seems to arise
from a linear axis at an angle of about
30 degrees, gradually bends outward
and becomes perpendicular to surface inmature zone.
Wall of zooecial tube

22

Sumio SAK11G.4MI

composed of concentric fibrous tissue,
about 0.03 mm thick. Thickness of rna·
ture zone about 0.40 mm. Superior andinferior hemisepta indistinct. rvlesoporesarranged in two rows. and its total
number dispcsed bet ween zooecial tubes
in one row 2 to 5. Inner diameter ofmesoporc 0.032 mm to 0.048 mm. Inter·spaces between zooecial tubes near sur·face about 0.13 mm.
Remarks:- The present form is questionably included in the genus Streblo·
trypel/a: because the central axis isobscure but may be present. The writer
examined the species belonging to thegenera Streb/of1)·pa and Streblotrype!la. andfound that at least Streblotrypa para/lela
CROCKFO!m which was reported fromthe lower Carboniferous of New South
Wales should be referred to Streblo/1)'·
pella. The present form is not identicalwith already described species of thegenus. however. better speciemcns arewanted for determination of the specificname.
Locality :-.'\ishiyama. Reg. no. 11236.

Genus Sulcm·etepora n'ORBIGNY, 1849
Sulcoretepora complicata
SAKAGAMI, n. sp.

Pl. 4, fig. 7.

Zoarium bi foliate, branching in planeof mesotheca. Mesotheca straight, apparently a closely joined double layer,
reaching surface at acute edges of zoarium. \Vidth and thickness of zoarium
about 3 mm and 1.2 mm to 2.0 mm ? (thelarger unknown owing to bad orientation). respectively. Zooecial tubes parallel to mesotheca for a very short distance. and then rapidly making large
angles in mature region. Zooecial tubes
near surface in tangential section circular, 0.14 mm to 0.21 m111 in diameter.
arranged longitudinally, 10 to 11 rowsof zooecia in one side of zoarium. 4 to
5 in 2 111111 longitudinal. and 5.5 to 6 in
1he same space diagonal.
I nterspaces
between adjacent zooecial tubes in mature zone 0.16 m111 to OAO 111111 longitudinal, 0.12 mm to 0.22 mm diagonal.
Vesicular tissue regularly arranged.

Explanation of Plate 3
Fig. I. Po/ypora jujitae S.\EAGAI>II, n. sp.
Tangential section of holotypc,, X 10. Reg. no. 11218-A.
Fig. 2. Pcnuiretepora sp. indet.
Tangential section. x 10. Reg. no. 11239-B.
Fig. 3. Penniretepora higaslziyamensis St\K,\GA:\11, n. sp.
Tangential section of holotypc, X 10. Reg. no. 11285.
Fig. 4. l'emriretepom regu/aris SAEAGA).If, n. sp.
Tangential section of holotypc, X 10. Reg. no. 11215.
Fig. 5. Pemriretepora cf. irregu/aris (NEKHOROSUEv)
Tangential section, x 10. Reg. no. 11214.
Fig. 6. R!rombopora sp. indet.
Oblique section x20. Reg. no. ll294.
Fig. 7. R/wbdomcson yabei 5.-\KACAMI. n. sp.
Longitudinal section of holotype, x20. Reg. no. 11202.
Figs. 8. 9. Proto1·etepora lrayasakae 5AKAGA1>11, n. sp.
s. Obverse surface of holotypc, 9, reverse surface of holotype. x5. ReR. no. 11279

S. SAKAGAMI:

SAKAGAMI

photo.

Bryozoa from the Omi Limestone, Pt. 2

Plate 3

444.

Bryozoa from the Omi Limestone. Pt. 2

usually consisting of one row of vesicles
between longitudinal rows of zooecia.
two or three rows composed of two or
t.hree vesicles in one row between zooe·
cia in one zooecial row. These arrangements become indistinct near surface.
Shape of vesicles rectangular in both
tangential and longitudinal sections.
vesicular tissue becoming gradually com·pressed on approaching surface of zoarium.
About one half near surface
covered by dense fibrous tissue.
Remarks:- The present form rather
resembles the Permian species Sulcon?te·/>ora nipponim SAKAGAMI (1961) which
was reported from Kamiyatsuse andlwaizaki of the Kitakami massif. japan
than the Carboniferous species alreadydescribed in the more complicated inner
structures. The differences between thepresent form and S. 11ipponica can berecognized only in the number of zooecial tubes per 2 mm in longitudinal row;
1 to 5 in the present form instead of 3
in S. nipprmica. The other charactersof the present form seem to be identicalwith that of the latter.
rocality :-Nishiyama. Reg. nos. 11274(holotype). 11237. 11273.
Sulcoretepora? sp. indet.
Pl. 4, figs. 8, 9.

Zoarium bifoliate, straight stem, notobsen·ed to branch.
Mesotheca apparently a closely joined double layer,
reaching surface at edges of zoarium.
Width and thickness of zoarium 1.5 mm
and 1.1 mm. respectively. Zooecial tuhes
parallel to mesotheca for a short dis·
tance. and then gradually bending out·
ward. Zooecium oval in tangential sec·tion, its larger diameter 0.12 mm to 0.16mm and shorter one 0.06 mm to 0.08 mm.
Vesicular tissue well developed in innerparr of zoarium. Vesicles variable in23

size. rounded rectangular or polygonal
in longitudinal section. Stereom welldeveloped but rather thin, its thickness
0.16 mm to 0.20 mm.
Remarhs :-The present form consistsof less complicated inner structures andmore rounded form than the above described species, and is considered to be
new to science.
However, there aresome doubt to include it in the genus
Sulcoretepora although the branchingpart of the zoarium could not be ob·served. because the inner structures of
the genus, especially in the case of thestructures being very complicated. issimilar to that of the hexagonellicl
genera such as Ramipora. Gmziocladia andVolgia. Therefore, the specific name must
wait until more specimens accumulate .
Lowlity :-Higashiyama. Reg. no.ll295.
References Cited
BASSLER, R. S. (1953): Bryozoa, in MooRe's"Treatise on Invertebrate Paleontology",
Hnt G. Geo/. Soc. America mzd Kansas
Unir•. Press, pp. GI-G253.
Co:-o;oi~A. G. E. (1902): New Bryozoa fromthe Coal Measurrs of Nebraska. Amer.
Geo/., Vol. 30. no. 6. pp. 337<l59. pis. 13-25.
CRocKFORD, j. (19.U): Permian Bryozoa ofEastern Australia. Part I. A Descriptionof Some Previously-named Species ofFenestrelliidac (Fenestcllidac). four. Proc.
Royal Soc. N. S. IV., Vol. 74, pp. 397-418.
pis. 18. 19.
- - (1947): Bryozoa from the Lower Carboniferous of New South Wales andQueensland. /'roc. Linn. Soc. N. S. IV.. Vol.
72. pts. 1-2, pp. 1-48, pis. 1-6.
- - (1957):
Permian Bryozoa from theFitzroy Basin, Western Australia. Commonw. Aust.. Dept. Nat. Det•elojmzent,
Bureau of J\li11. Res.. Geo/. Geophysics,
Bull. No. 34, pp. 1-134. pis. 1-21.
ELIAs, M. K. (1957) : LaLe Mississippian
Fauna from the Redork Hollow Formation of Southern Oklahoma. Part I. four.

24

Sumio SAKAG,4Ml

Pa/ermt .. Vol. 31. 110. 2, pp. 370-427, pis.
39-50.
Ft'JITA. H. (1938): Geology of the Omi limestone (Permo-Carboniferous). 2\iigata
Prefecture. (in japanese with English
abstract) Contr. lnst. &col. Palcont.. Tohoku l.hrh•., No. 48, 60 pp.
IIAYASAKA, I. (1924): On the Fauna of the
Anthracolithic Limestone of Omi-murain the Western Part of Echigo. Sci. Rep..
Tolwktt Imp. Univ., Ser. 2, Vol. 8, no. 1,
pp. 1-83. pis. 1-7.
KAWAO,\, S. (195·1):
Stratigraphical and
Palaeontological Studies of the Omi Limestone in the Itagamine District. Niigata
Prefecture. (in Japanese with English
abstract) Studies Geol. Mill. bzst. Tokyo
Uni11. Educ., No. 3, pp. 15-27.
MoRozovA. L 1'. (1955): Kamennougolnyel\Ishanki Srednego Dona. Akad. Nauk
SSSR, Trudy Palconl. !nsf.. Tom 58, 88
pp., 12 pis.
NEKHOROSIIEV, V. P. (1948): Kamennougolnye:Mshanki Scvero-Vostochnogo Pribalkhashia. A.kad. Nauk Daz. SSR., 70 pp., 11 pls.
- - (1953): Niznckamennougolny(' 1\'lshanki
Kazakhstana. Akad. Nauk SSSR .. Trudy
VSEGEI. 235 pp.. 25 pls.
- - (1956): Niznekamennougolnye Mshanki
Altaya i Sibili. !bid., Tom 13, .U9 pp., 57
pis.
Nll<IFOROV,.\, A. I. (19:{:l): The Carboniferous
Deposits of Central Asia. Contributions
to the Knowledge of the Lower Carboniferous Bryo?.oa of 1he Turkestan. Tmns.
United Geo/. Prospecling Serv. USSR. Fasc.
207. pp. 1-77. pis. 1-12.
SAKAGAMI, S. (1960): l-layasakajmra, A New

Permian Bryozoan Genus from Iwaizaki,!\·Iiyagi Prefecture. Japan. Trans. Proc.
PalaeOilf. Soc. japan. N. S., No. 39. pp.
321-323. pl. 37.
- - (1961): Japanese P;;rmian Bryozoa. Palaeon/. Soc. fapau. Spec. P.1p., No. 7. 58 pp.
30 pis.
- - (1962): Lower Carboniferous Bryozoa
from the Hikoroichi Series. Japan. Traus.
Proc. Pa!aeont. Soc. fapcm, N. S., No. ·16.
pp. 227 -2·12. pis. 35-37.
- - and AK,\GI, S. (1961): Lower Permian
Bryozoa from Miharano, Taishaku Plateau, Southwestern Japan. Ibid .. No. 4.1,
pp. 105-112, pl. 15.
Sl-ll!LG;\-NESTERENKO, M. I. (1951): Kamennougolnye fenestcllidy Russkoii Platformy. :1kad. Nauk SSSR., Pa/t'ont. Inst.,
Trans .. Tom 37. 8-l pp .. 3-! pis.
- - (1955): Kamennougolnye I\fshanki Russkoii Platformy. Ibid., Tom 5':'. 204 pp.,32 pis.
Trro:o.IPSON, M. L. (1948): Studies of American
Fusulinids. Uniz•. Kaw;as Paleont. Coulr.,
Protozoa . .tlrt. 1, pp. 1-18-1. pis. 1-38.
ToRI YA~lA, R. (1954) : Geology of Akiyoshi.
Part I. Study of the Akiyoshi LimestoneGroup . .Mem. Fw·. Sci., l(mslm Uuiz• .. Ser.
D, (;col., Vol. -1. uo. 1. pp. 39-97.
- - (1958): Geology of Akiyoshi. Part III,
Fusulinids of Akiyoshi. Ibid., Vol. I, pp.
1-21iL pis. 1--!8.
TRIZl':\, V. B. (1958): Rannekamennougolnye
Mshanki Kuznechkoi Kotloviny. Trudy
VNJGR!. 122. 435 pp., Ii-I pis.
ULRICH. E. 0. (1890): Palaeozoic Bryozoa.
Geol. Sun• Ill .. Vol. 8. pp. 285-688, pls.

29-78

Explanation of Plate ·1
Figs. 1-3. Streb/ast·opora amabilis SAKAG.UII, n. sp.
1, Longitudinal section of hololype, X20. Reg. no. 11272, 2, enlarged part of holotype,x-!0, 3. tangential section of paratypc, X40. Reg. no. 11216-B.
Figs. 4. 5. Stre/Jfascopora grossa SAK:\GA!I-11. n. sp.
4. Longitudinal section of holotype, x20, Reg. no. 112•11-A, 5, longitudinal section ofparatype, x 10. Reg. no. 11285.
Fig. 6. Streblotrypella? sp. indet.
Longitudinal section. x20. Reg. no. 11236.
Fig. 7. Sulcoretepora complicata SAKAGA:O.II, n. sp.
Tangential section of holotype, X 10. Reg. no. 1127·1.
Figs. 8, 9. Sulcoretepom? sp. indet.
8, Longitudinal section, x20, Reg. no. I 1295-A, 9, transverse section x20. Reg. no. 11295-B.

S. SAKAGAMI:

SAKAGAMI

photo.

Bryozoa from the Omi Limestone, Pt. 2

Plate .1,.

Trans. Proc. Palaeont. Soc. Japan, N. S., No. ·19. pp. 25-28, pl. 5, Apr. 10. 1!163

445.

NEW SPECIES OF c;ADUS FRO~f THE PLIOCENE OF JAPAN*
KOTORA HAT AI and T AMIO KOTAKA
Institute of Geology and Palco:1tology, Tohoku University

EI *ft~M:.iliii!ift~ = Wll:~~atM~ J: l..-'lll~~~Jhu&:i'fl J: 'J~t:;t:ll~'P~lfu.:~t*imil'i.:..!)l~-t :Q fmf.f4:tt; J: O:ili:!l.:H~

sp. :t<; J: 0: Gadus maswlai n. sp. c ~filA-:,

l*m t:t-t. t-:IH.Ji.: "'?t. --c.

c .lt~~t-f L., Gadus cllikagatraeusis n.
;:(I(

Jr '1-

Ltc. • 'J'

l:'li

1~

~

shima Straits. off the coast of Yamaguchi and Shimane Prefectures (Nm-;o,1931: KoTAKA, 1951. 1959). the discovery
of a Gadus-specimen showing fossilaspect is not unreasonable.
Gadus macruceplwlus TELJ·:SJL:s fromthe North Pacific Ocean. particularly
the eastern coast of Hokkaido southwards to Rikuzen, Gadus lltsuts LIN·:'\AEUS and Gadus morslma LI:\:\AEUS.
both from around the British Islands inthe eastern North Atlantic Ocean are
all characteristic in the long and narrow
shapes of their ear-bones. These species
have crude tubercular ornamentation on
their outer face anti their smoother inner face show longitudinal sulcus whichis generally only slightly depressed.
Showing resemblance with the ones
just mentioned is Themgra clta/cogramma(PALLAS) from the sea of eastern Hok·kaiclo, the differences between them being the strength and development of
the crude tubercular sculpture of theouter face. details of the longitudinalsulcus of the inner face. degree of curvature of the narrow shafts and otherminute details.
Fromcomparative
morphologicalstudies on the shafts of the Recent andfossil species above mentioned, it isclear that the grade of variation shownby the specimens arc within recognizable
limits. and that the differences between

Introduction
Although the genus Gadus is common
in the waters of the North Pacific. especially along the eastern coast of Hokkaido. there has been no record of its
species from the Cenozoic deposits extensively distributed in the JapaneseIslands.
In the collection at hand there are
two species of the genus Gadus. one
from the Pliocene I lamada formation(HATAI. I\lASUDA and SUZUKI, 1961) exposed along the Chikagawa stream flowing through Tanabu-machi. Shimo-Kitagun, Aomori Prefecture. and anotherdredged from a depth of 52 meters off
Kamo-machi, Yamagata Prefecture. The
latter specimen is water worn, coloreddark reddish brown as if due to ironoxidation and more or less similar tothe specimen from the Hamada formation just mentioned.
Both of these
specimens differ from the Recent speciesof the genus Gadus in features thought
to be worthy of their specific separationfrom the living species.
Since it is known that Pliocene fossils
have been dredged from the sea bottomof the Japan Sea bordering Niigata.
Yamagata, Akita and Aomori Prefectures (IlATAI, 1939), the Korea and Tsu• Received June 13. 1962; read June 2. 1962.

:25

26

1\otom f!;\TAI and Tam:·o KoT.li\.A

the variation-groups of the species do
not appear to overlap one another, forwhich reason, specific classification ofthem is rendered reasonable.
Since the specimen from the Pliocene
Hamada formation can be distinguishedfrom previously known species it is described in the following lines as new toscience.
Description
Gadus cliilw}.[awaensis !lATA! andKoTAKA, n. sp.
Pl. 5, figs. 3, -1.

Shaft about 23.0 mm in length, of outerface. nearly 9 mm in height and moderately thick; outer face gently concave.
inner face convex. Anterior side moreacutely pointed than posterior. extremities narrowly rounded. Dorsal side
more arched than nearly straight ventral. Outer face longitudinally depressedin middle part, depressed area more distinct on posterior half than anterior.
Dorsal and ventral borders with distinct.
bluntly rounded tubercular structures.
numbering about five to six per 5 mmon middle part of dorsal border andabout five per 5 mm on ventral. Eachtubercle gradually becoming narrcnver
in width and meeting each other at central anterior part but not elsewhere
where they terminate in longitudinal
central depression. Ridges broad andwith sharp tops at central anterior partbut elsewhere merely rounded. Tubercle
forming anteriormost and posteriormost
parts or extremities largest and wellrounded. Inner face with longitudinal
depression or sulcus bordered with rounded ridges and measuring about 20 mm
in length and about 4 mm in maximum
width, open anteriorly but not reaching
posterior margin. Dorsal area of innerface with ridges extending to sulcus
margin. comb-like in appearance, butridges much broader nnd gently rounded
compared with valley-like interspaces,
b3coming oblique ncar anterior andposterior extremities. otherwise quite
perpendicular to sulcus. Area dorsal tomedian sulcus much broader than that
ventral to it. Dorsal part of mediansulcus appearing blocky due to quadrangular separated, slightly elevated. flattopped ridges with sharp borders numbering 7 in posterior part of sulcus andonly one longitudinally elongated one inanterior part. Anterior one separated
from posterior ones by rather wide depressed area.
Locality, geological formatio1l. geologicala}.[e and repository :-Cliff of the Chikagawa Stream flowing through Chikagnwa. Tanabu-machi. Shimo-Kita-gun.
Aomori Prefecture. Hamada formation.
Early Pliocene. Preserved in the collecof the Institute of Geology and Paleontology. Tohoku University. Sendai, Japan.
IGPS coli. cat. no. 79164.
Comparison:-The present new species
was compared with the earbones of the
Recent Gadus macrocephalus TILESIUS(pl. 6, figs. 5-10) collected from the
Pacific Ocean of eastern Hokkaido andnortheastern Honshu, Gadus morslma
LJ NNAEuS (figs. 13, 14) and Gadus luscus
LlK:\AEuS (figs. 15. 16) both from the Atlantic Ocean off the British Islands, andwith Thera;:ra chalco,l.!ramma (PALLAS)(figs. 11. 12) from southeastern Hokkaido.
Among these mentioned species of Gadus
only Gadus macrocep/zalus TILESI!JS and
Tlwragra c//(/lcogmmma (PALLAS) are
worthy for comparison because theothers are of either different shape orwith different kind of sculpture on theirouter and inner faces.
Compared with the common living
species, Gadus macrocephalus, the present

44.5. Gadus from the Pliocene
one can be distinguished therefrom by
the much weaker tubercles and ridgesof the outer face, better development of
the terminal tubercle of the anterior andposterior parts and by the details of thesulcus.
The present new species is also distinguishable from Theragra chalco{fram·ma (PALLAS) by having stronger sculpture of both outer and inner faces.
Remarks:- The Recent specimen used
for comparison with the fossil material
in the present paper are those collectedfrom the sea off eastern Hokkaido andnortheastern Honshu, specimens bought
at the market in Sendai City, and thosereceived in exchange from Dr. F. C.
STIRTO:\ of England. to whom the writers express their thanks. It may be
added that considerable difficulty arises
in studying ear-bones because they must
be extracted from Recent material identi·
fled by specialists. and the specimens
are those collected in person from various parts of the Japanese ls'ands.
The specimen dredged from a depthof 52 meters off Kamo, Yamagata Prefecture (IGPS coli. cat. no. 79165) is somewhat water-worn and iron-colored ~fos·sil). measures about 19 mm in lengthand nearly 8 mm in dorso-ventral direction. This specimen differs from Gadus
clzikagmcaensis is having a less numberof stronger tubercles at the dorsal andventral marginal areas. a more defined
longitudinal central depression which is
sharply incised. two outstanding tubercles midway between the anterior andposterior extremities on the outer face.
more rounded anterior and posterior extremities. and more uniformly roundedridges extending towards the medial
longitudinal depression.
From thesedifferences between the present specimen
and the newly described one. it is considered that it must be retained as a27

distinct form.
Compared with Gadus macrocep!talus
TILESIUS which is the most similar
species among the genus Gadus. thepresent one (probably of Pliocene age)differs in the general shape of the earbone, stronger but less elevated andmore rounded ridges extending from thedorsal and ventral borders towards thelongitudinal depression of the outer face.
development of two outstanding tubercles in the middle part of the outerface, and by the more defined longitudinal depression of the outer faces.
From the differences from the twomore closely related species. Gadus chika·
gau·aensis and Gadus macrocejJ!zalus. thepresent specimen dredged from off Kamo. is considered to represent an unde·scribed form. For this specimen thename of Gadus nwsudai I·IATAI and KoTAKA. n. sp. (pl. 5. figs. 1. 2) is p:oposed.
It is hoped that more specimens of thisspecies will be found by future dredge
operation in the japan Sea.
Gadus masudai from its coloration andgrade of preservation is considered to be a
fossil reworked from the Pliocene strata
now submerged (NOMURA and HATAI.
1937) in the Japan Sea. It is known
that Pliocene fossils are not rare in thematerials dredged from the bottom of
the japan Sea (HATAI, op. cit.). and it isof interest that they are generally dis·colored and usually distinguishable from
Recent specimens although sometimes
distinction is diftlcult. Particularly it
should be mentioned that extinct species
of marine molluscs are known to befound in the materials dredged fromthe sea bottom. and brachiopods of thesame category also occur. Also thepresent geographical distribution of theGadus, suggests that the present specimen is a fossil one.

~8

Kotom l!A.TAI and Tamio KOTAKA
References

H.-\.TAI. K. (1939): A geological considerationof the fossils dredged from japanese
Seas. Geologie der Jleere tmd Bimumgetciisser. Vol. 3, No. 3, pp. 366-374.
- - (1950): Cenozoic mollusca from Aomori
Prefecture, japan. jour. Couchy/., Volume
de Centenaire, Vol. 90, No.2. pp. 128-136,
1 text-fig., 1 table.
- - . MASUDA, K. and SuZUKI. Y. (1961): A
note on the Pliocene molluscan megafossil
fauna from the Shimokita Peninsulu,
Aomori Prefecture. Northeast Honshu,
japan. Saito flo-on Kai Jllus., Rt!S. Bull.,
No. 30, pp. 18-38. pis. 1--!, 1 text-fig., 3tables.
KoTAI.;A. T. (1951): Recent Turritella ofjapan. Short Papers IGPS, No. 3. pp. 7086, pis. 11-12. 1 map, 2 tables.
- - (1959): The Cenozoic Turritellidae ofjapan. Sci. Repi., Tohoku Unil•., 2nd Ser.,
Vol. 31, No. 2, pp. 1-135, pis. 1-15, 10
text-figs., 8 charts. 3 tables.
NrrJ\'o, H. (1934):
Discovery of Thyasirabisecta Co:o.:RAD from the sea bottom at
Korea Strait (in japanese). ]our. Geo/.
Soc. ]afxm, Vol. 41. No. 489, pp. 176-182.
2 pis., 1 text-fig., 5 tables.
No:o.tU!lJ\, S. and HATAr, K. (1937): An interesting problem concerning the marine
fauna of the Japan Sea. Rer. Ocean. Wk.~.
japan, Vol. 9, No. 1. pp. 27-35.
TANAKA. S. and ABE, .M. (1955): One thousand useful fish species (illustrated injapanese). J11orillita Pub/. Co., Tol•yo, pp.
1-29·1+1-12, 1000 text-figs.

Explanation of Plate 5
Figs. I. 2. Gadus masudai HATAI and KoTliK:\. n. sp. x2
Submerged Pliocene deposits. off Karno, Yamagata Prefecture. IGPS coli. cat. no. 79165.
Coil. K. MASUDA. 1 ; outer face. 2 : inner face.
Figs. 3. 4. Gadus chikagawaensis I'L'\TAI and KoT AKA. n. sp. x2
Hamada formation, Pliocene. Cliff of the Chikagawa Stream flowing through Chika-gawa,Tanabu-machi. Shimo-Kita-gun, Aomori Prefecture. IGPS coil. cat. no. 79164. Col!. K
II ATAl. 3: outer face. 4: inner face.
Figs. 5,-8. Gadus macrocephalus TILESII..'S x2
Recent, bought at lshinomaki City. Miyagi Prefecture. 5; outer face, 6: inner face, 7:
outer face, 8: inner face.
Figs. 9, 10. Gadus macrocephalus Tu.Esms x5
Recent. bought at Yuriage, 1\liyagi Prefecture. 9: outer face. 10: inner face.
Figs. ]I, 12. Theragra clwlcogramma (PALLAS) x2
l{ccent. Muror:m. Hokkaiclo. 11 : outer face, 12; inner face.
Figs. 13, 14. Gadus morslwa LIN:-;AEUS x5
Recent. Atlantic Ocean. 13: outer face. 14: inner face.
Figs. 15, 16. Gadus luscus LI:-::-;AEUS x5
Recent. Atlantic Ocean. 15: outer face, 16: inner face.
Figs. 17. 18. Mugil cephalus LtN:-;AEIJS x5
Recent, Sagami Bay, Kanagawa Prefecture. 17; outer face, 18; inner face.
Figs. 19. 20. Merlwxius l'lllgaris FLEI>If:"'G X2Recent, Australia. 19; outer face, 20: inner face.
Figs. 21. 22. Neoplatycep/wlus macrodon FLEMING x3
Recent. Australia.. 21 : outer face, 22: inner face.
Figs. 23, 24. Urophycis lmrziliensis KAUP x3
Recent, Brazil. 23; outer face. 2-1: inner face.

H ATAI and KOTAKA:

1

Gadus from the Pliocene

Plate 5

2

3

5

8

9
7

6

11

13

K uMAGAI

photo.

14

Trans. Proc. Palacont. Soc. Japan, N. S.. No. 49. pp. 29-38. pl. 6. Apr. 10, 1963-!46.

RE.MARKS ON SOME FOSSIL DOSINIA OF JAPAN'*
KOICill RO MASUDA
Department of Geology, Faculty of Education, Tohoku Cniversity

c

c 't*

Dosinia fl::rt!O=.:.::.•:. "'?l •-c : ffi:IO{~#~h -c~ • :Q t 10 1R10•:>f..:P3ff!l!O Jfbf*l¥1~~~tt~~~L. I-'3ID!~:.I!lic L-c;le~~ht.:. Dosinia tugaruana ~f!f~lf.tLt.:., -t:"IOM!
Dosinia rtkaisia11a. Dosi11ia hataii, n. sp., !JMinia kwmoi, n. sp. ~:rc ~il( L. ~ k !Oil€/ll!flij![;:."'?t. ··O~~Lt-:.

WI l!l "¥•~fm

times fragments or inner molds.
The numerous specimens, fossil andRecent of Japan and the adjacent regions
and of foreign countries. preserved in
the collections of the Department of
Geology, Faculty of Education. in theInstitute of Geology and Paleontology.
Faculty of Science, both of the Tohoku
University, and in the Saito Ho-on Kai
Museum, all in Sendai City, were studied
by the writer.
In the prL'sent article discussions on
the classification. especially on the rela·tionships between the fossil specimens
with shell materials and those withoutthem, and the descriptions of somespecies including two new forms are
given. Remarks are given on the stratigraphical data of some ~Iiocene dosinid
species because of their biostratigraphical importance in the Neogene deposits of japan.

Introduction
The pelecypod molluscan genus Dosinia
is one of the interesting and most predominant groups among the fossil pelecypods of Japan.
Because the genus
Dosinia from the Cenozoic deposits ofjapan is abundant in individuals. com·prises a number of species. and showsa wide range in surface sculpture, ex·tensive geographical distribution. rather
restricted geological range of its respective species, it should be accepted as
one of the taxons very important bothin stratigraphr and chronology. Further
from the living habitat of the dosinid
species interpretation of the fossil one
can be analysed.
From japan and the adjacent regions
a total of 25 species of Dosinia havebeen described from the Cenozoic strata.
among which 20 arc fossil and extinct
species. and 21 are known from the
Recent seas around the japanese Islands.
The morphological features of Dosi11ia
are so simple that classification of thespecies have often caused confusion.
Confusion has also arised in cliscrimina·
tions because the type specimens offossil Dosinia have been based uponimperfect specimens which are some-Acknowledgements
Acknowledgements are due to Pro·
fessor Kotara HATAI. Institute of Geology and Paleontology. Faculty of Science. Tohoku University. for his conti·
guous guidance and various suggestions.
Thanks are due Lo Professor Yoshio
0:-.UKI, Department of Geology. Facultyof Education. Tohoku University, for* Received Apr. 20. IY62: read at the annual
meeting of the Society at Sendai, Jan. 20. 1962.
29

:30

Koichiro Jl.4SVDA

his encouragement.
Appreciation is
given to Drs. Tamio KoTAKA and Shozo
HA YASAKA of the Institute of Geology
and Paleontology, Tohoku University. fortheir discussions on problems relating
to the paleontology of the genus Dosinia.
Some Notes on Classification
For the sake of convenience the tcr·
minology employed in the present study
is explained below (text·fig. 1).
A. Shell·size
Small ........ less than 35 mm. in length
Medium ...... between 35 mm. and 60 mm.
in l~ngth
Large ........ exceeding 60 mm. in length

B. Convexity of valves
Slight. ....... depth/length less than 0. 2
1vloderatc .... depth/length between 0. 2
and 0. 3
Great ........ depth/length exceeding 0. 3

C.

Pallial sinus
Shalla'\\ ...... depth of sinusjlenglh lessthan 1/3
Medium ..... depth of sinus/length between 1/2 and 1/3
Deep ........ depth of sinus/length exceeding 1/2~---L

Text-fig. I. II-Ilcight, L-Length, DsDepth of sinus, Ll-Length of lunule.

The fossil specimens of Dosinia frequently occur with the original shellmaterials completely dissolved and thus
represent molds. Since the shell of
Dosinia is thicker near the hinge area

than elsewhere,
the morphological·characters of the fossil specimens lacking the shell materials differ consider·ably from the original ones in which
the shell materials are retained. Therefore. it is very often quite difficult todiscriminate the precise position of thespecies in the case of mold specimens.
To overcome such difficulties the writer
has tried to compare the morphologicalcharacters of the specimens preserving
their external shell with their molds.
This procedure was by inserting plasterof paris or modelling compound into theinner shell of the specimens or into theircasts (text-fig. 2).
As the results repeated experiments
the writer observed the following inter·esting facts. In the case of the specimens
consisting of molds. (1) the height anddepth of shell becomes smaller than theoriginal form, (2) the beak tends torecede posteriorly and somewhat up·
wards. so that. the beak shows featuresdifferent from the original form, (3) the
curvature of the postero-dorsal margin
becomes considerably smaller comparedwith that of the original form.· and (4)the lunule tends to become obscure. In
the case of the shells with obscure
lunule the anterior part tends to become
somewhat concave towards antero·dorsal
margin, but in the case of the shells
having the distinct lunule the anterior
part becomes concave considerably. (5)the antero-dorsal and postero·dorsal
margins are frequently broken, because
the shells at such parts are very thincompared with the other parts. (6) thefeatures of the escutcheon become obscure, and ('/) the surface sculptures areusually not observed. very vagul', oronly observed insufficiently.
The above mentioned features canalways be observed in any one species,
but the grade of difference between theHli.

Fossil Dosi:nia:nText-fig. :!. Showing the rdation between shell and its innermold of /Josi11ia (Pluzco.,oma) japoniw (REEVE).

original form and the molds will varyfrom species to species. Therefore. it
is necessary to have adequate knowledgeof the above mentioned features whenstudying fossil specimens in which theoriginal shell materials has been lost.
In the case of water worn shells themorphological characters of the outersurface usually tend to become different
from those of the original form. that is
to say, the concentric growth lines of
the water worn shells are usually lower.
flat-topped and stout compared with theoriginal form and may give the impression of a different species.
Experiments as abo\·e mentioned arebeing continued with other kinds ofbivalves and in addition slicing of theshells and their structures in profile andsurface are also planned for study.
Description

Family Veneridae
Subfamily Dosiniinae
Genus Dosinia ScoPOLI. 1777

Dosinia akaisiana No~IURA, 1935
Pl. 6, figs. 3 -5.
193:5.

Dosi11ia akaisiana No!.tt•t~t\, Saito Ho-rm
Kai Mus., Res. JJu/1 .. .Vo. 6, p. 60, pl.

1961.

S. figs. 1 6.
Dosi11ia akaisic11a, l\\'AI. Trans. Proc.
Palaccmf. Soc. ]apa11. N. S .. ,Vo. 41. pl.
1. fig. 'i.

Holotype :-S:'vl*. Reg. 'lo. 606l.
Description :-Shell small, moderate inthickness. rather greatly inflated. suborbicular in outline, nearly as long as
high. Anterior side well rounded, passing gradually into broadly roundedventral margin: antero-dorsal side short,conca,·e in front of beak: postero-dorsalside long. broadly rounded. gradually
passing into ventral margin. Beak point·
ed. curved forward. situated anteriorly atabout 3/4 of shell length; lunule cordatein shape, rather small. deeply impressed;
escutcheon linear, somewhat impressed
but rather obscurely defined by rather
obscure ridge. Surface sculptured withrather low. close-set. lamellatecl concentric growth lines. about 23 per 10 mm.
at 10 mm. from beak in central part ofdisc; concentric lines asymmetrical inprofile. somewhat more distinct. moreelevated and more slender at anteroand postero-dorsal sides than on central
part of disc. Pallial sinus somewhat
sloping upwards. rather large. medium* abbreviation for Saito Ho-on Kai Museum.
Sendai.

•">'),,_

Kvichiro llf;lSCIJJl

in depth. oblong-triangular in shape.
pointed at end; adductor muscle scars
slightly impressed. oblong in shape; anterior one somewhat smaller than posterior. Anterior lateral tooth of left
valve rather large. somewhat elevated;
median cardinal teeth strong and broad;posterior cardinal tooth rather low andslender.

Dimensions (in mm.) :-1-Iei~ltt
-32*

32. 5

31..

I

1

-~--

1

1

·I

3;:-5·
33
I
32.5

I

Leng-th 1 De-pth
Ds/L-i---. 32
10.5

o. 59

31. 5
31. 5

0. 6

35

12.5

32

10

33.5

9.5

0. 15
0. 15

--

~---1_1~--'---'• Holotype. ** Specimens from the Kurosedani formation, Toyama Prefecture.
Remarl~s:-Tlwugh the present species
is usually small in size, one moderate
sized specimen (about ·13 mm. in length)
was found among the paratype specimens.
The present species is characterized
by its small. rather greatly inflated. orbicular shell with numerous concentric
growth lines. broad. triangular pallial
sinus pointed at end and rather large
anterior lateral tooth of the left ,·alve.
This species resembles Dosinia r:hilm·
zenensis NAGAO from the Oligocene
Ashiya group of Northern Kyushu (NAGAO. 1928) and Dosinia suketoensis OTLJKA from the ~-liocene Shobara formationin Hiroshima Prefecture (0TUKA, 1938b),
but it can be distinguished from theformer by its smaller. more inflatedshell. small and oblong lunule and theshape of pallial sinus. and from the latter by the position of beak. obscurely
defined escutcheon and the shape of

pallial sinus. Another related species is
Dosinia gnmeri (PHILII'PI) (REEVE, 1851),
a Recent species of the China Sea, but
gruneri is distinguishable from the present one by its moderately inflated shell,
distinctly defirwd escutcheon, small anterior lateral tooth and the shape of
pallial sinus which is roundly pointed
at end.
Type localit_v, Geological formation andilge :-Path on the southern side of thevalley Wakinosawa. about 2 km. west ofHitotsumori. Ajigasawa-machi, NishiTsugaru-gun, Aornori Prefecture (Lat.
·10',1l'N .. Long. J.J0'08'05"E.). Tanosawa
formation. Early Miocene.
Described specimens:- Holotype andparatype specimens.
Distribution:- Tanosawa and Ainaigawa formations. Aomori Prefecture;
Kuroseclani formation. Toyama Prefecture; Yanagawa formation, Fukushima
Prefecture: Shira tori member of Kadonosawa formation, !wale PrefeclUre: all
Early Miocene in age.
Associated fauna :-l'edalion tugam('nse No:\IURA, Ostrea gravitesta YoKoYAl\IA.
"Patinopecten" (Kotorapecten)
itcasakicnsis (NoMURA), etc. at the type
locality.
Subgenus Plzacosoma
JUKES-BROWNE. 1912

Dosinia (Piwcosoma) tugaruanal'\01\IURA, 1935
Pl. 6, rigs. 8. 9. lOa-b.
1935.

!Josinia tugaruana NoML'RA, Saito Hoon Kai :Hus.. Res. Bull., So. 5, p. 58.

1935.

!Jo.~inia odosensis No~IURA,

pl. G, fig. G.

l9GI.

Ibid., p. 5!l.
pl. 7. fig. 2 (?).
!Josi11ia tugaruana, IWAI, Tl'lms. Proc.
Palaecmt. Sot.'. japcm, lll. S., No. 41, pl.
1. fig. 4.

Ha.
1961.

Fossil Dosinict

Dosi11ia odose11sis. IwAI, Ibid., pl. l. lig.
11.

1961.

Dosiniu chikuzenensis, IwAI, Ibid., pl. l.
fig. 8 {non NAGAO. 1928).

Holotypc :-SM. Reg. No. 602·l.
Description :-Shell medium to smallin size. moderate in convexity; suborbicular in outline. somewhat longer thanhigh. Anterior side rounded. passing
gradually into broadly rounded ventral
margin; antero·dorsal side rather short.
somewhat concave in front of beak;postero-dorsal side long. broadly rounded,passing somewhat abruptly into ventral
margin. 13eak pointed, curved forward.
situated anteriorly at about 2/3 of shelllength: lunule rather deeply impressed.
rather long, cordate in shape; escutcheon
linear. depressed, defined by somewhat
distinct ridge. Surface sculptured withclose·set. low, lamellated. fine concentric
growth lines, about 30 per 10 mm. at 10mm. from beak in central part of disc:concentric lines asymmetrical in profile,broader than their interspaces. more
distinct. more elevated and more slenderthan those of central part of disc atantero· and postero·dorsal sides. Pallial
sinus somewhat sloping upwards. rather
large. medium in depth, oblong trian·
gular in shape, bluntly pointed at end:
adductor muscle scars somewhat im·pressed; anterior scar somewhat smaller
than posterior one. Characters of hingearea unknown.
Dime11sio11s (in mm.) :----H;>ight

Length
1

-

-~--

41

C::t.45•

i

-

43
47

I Depth

Ds/L

Ll/L

!

0.47

0. 18

0. •14

4·1. 5
32.5

11. 5

42

44

ca.l2

31

38

. Ilolotype .

0. 18
0.45
0. ·1388

Remarks :-Since this species was described by NoMURA based upon one innermold which lacks its anterior side, its
detailed morphological characters re·mained unknown.
Dosinia odosensis
which \vas described from the same formation with tugaruana by NoMURA at
the same time based upon an imperfectspecimen retaining a little shell material
may be a synonym of tugamana withquery. This procedure can be acceptedfrom the relationships between the rr.old
specimen and the specimen with shellmaterials as already stated in the earlier
pages: the characters of odosensis appear
to correspond with those of lugamana.
Though it is considered that further
study is necessary to settle this problem.
the writer considers that odosensis is asynonym of tugaruana.
The above mentioned description was
based upon a plaster-cast specimen lack·ing the ventral part and collected fromthe Tanosawa formation at Odose. Fukaura·machi, with other dosinid specimens.
Dosinia nomm·ai 0TtJKA (1931) described from the Shiratori member of the
1\.adonosawa formation in lwate Prefec·
ture is closely related with the present
one but it can be distinguished by its
lower shell. rather protruded anterior
part and short lunule (Ll/L. 0.15-016).
Dosinia anguloides NOl\!URA (1935b) fromthe Ajiri formation in Shiogama City.
t-.fiyagi Prefecture. which was describedbased upon a mold specimen, is distin·
guishable from the present species by
its larger and higher shell and narro\v
and deep pallial sinus which is roundedat end.
Type locality. Geological formation and
Age :-Hotatezawa, about 3 km. north·west of the Shinyu hot-spring, a tribu·
tary of the Sasanai RiYer. lwasaki·mura,Nishi·Tsugaru·gun, Aomori Prefecture(Lat. 40'35'N .. Long. HO'Ol'E.). Tano·Koichiro ill.-tsUDA

sawa formation. Early l'vliocene.
Described specimens :-IGPS*, coli. cat.
no. 37060. Odose, Tanosawa. Fukauramachi. Nishi-Tsugaru-gun, Aomori Prefecture. Tanosawa formation.
Distribution:-Tanosawa and Ainaigawa formations, Aomori Prefecture;
Kawai formation (?), Shimane Prefecture: all Early Miocene in age.
Associated jauua :-Ciilamys akitana(YOKOYA~IA), "Patinopecten"(Kotorapecten) iwasakiensis (NOMURA), Glycymeris sp., etc. at the type locality: Ostrea
cf. gravitesta YOKOYAMA, Opercu/ina comp!anata japonica HANZAWA, i\Jiogypsillakotoi HANZAWA, etc. at Odose. Fukauramachi.
Dosinia (Pizacosoma) lzataii
l'v[ASUDA, n.

sp.

Pl. 6, figs. la-c. 2.
l!l26.

1936.

Dosiuia troschl'!i. YOKOYA~IA, ]o11r. Fac.
Sci., Imp. Unir•. Tokyo. Sec. 2, Vol. 1,
Pt. ·1, p. 133, pl. 16, fig. 1 (non LISCIIKE.
1873).
Dosiltia odose11sis, No~rUIL\ and lL\TAI,

Saito Ho·on Kai Mus.. Res. Bull., No.
10. p. 128, pl. 1·1, fig. 1 (non NmJURA,
1935).

Holotype :-IGPS, coli. cal. no. 72"176.
Descriptiou :-Shell large, thick, moderately inflated. suborbicular in outline,
nearly as long as high. Anterior siderounded, passing gradually into broadly
rounded ventral margin; antero-dorsalside short, somewhat concave in frontof beak : postero-dorsal side broadly
rounded. passing somewhat abruptly
into ventral margin and forming a broad
angle with postero-ventral margin. Beakpointed, separated from beak of other• abbreviation for Institute of Geology andPaleontology. Faculty of Science, Tohoku
University, Sendai.

valve by deep hollow. somewhat curved
forward, situated anteriorly at about:3;4 of shell length; lunule cordate inshape. rather large. deeply impressed:
escutcheon narrowly elongated, somewhat depressed. defined by not very
distinct ridge. Surface sculptured withlow, close-set, lamellated. rather irregular concentric growth lines which about
22 per 10 mm. at 10 mm. from beak incentral part of disc; surface slightly
undulated from beak to ventral marginwhich may indicate resting stages ofgrowth: concentric lines at posteriorside somewhat distinct, elevated andslender compared with those at otherplaces of disc. Pallial sinus slightly
sloping upwards. rather narrow andsmall, medium in depth, reach to about
central part of disc, somewhat roundly
pointed at end. Adductor muscle scars
large, oblong, distinctly defined. nearly
equal in size. In left valve anterior
lateral tooth rather small and low;
median cardinal teeth rather slenderand elevated: posterior cardinal tooth
elongated and slender.
Dimensions (in mm.) :====:=~~==~Height j Length ! Depth I Ds/L

I LI/L-w---1· 661- 19.5:-0.55 0.15
56. 5
56
60

57
59
62

61

53

5-1

~(i

50

11. 5
18

o. 15

,I

0. 15

0. 5
18
15

• Holotypc.

Remarks :-This new species was firstillustrated by YoKOY.HIA (19261 fromthe .\liocene Kanomatazawa formationin Tochigi Prefecture as Dosinia trosche!i
LISCHKE and subsequently by Nol\ILJI{Aand HATAI (1936) from the 1\liocene

446.

35

F'os!>'il Dosinia

Tanagura formation in Fukushima Prefecture as Dosinia odosensis NoMURA.
1\.s the results of comparative study it
is evident that the specimens of YoKoYAl\IA and NOMURA and HATAI represent
a new form of Dosinia. Therefore. ane,,· name is proposed for the Tanagura
and Kanomatazawa specimens.
The present new species is named inhonor of Professor Kotara HATAI of the
Tohoku University.
The shell at the younger stage of thepresent new species possesses a longer
shell but tends to become orbicular withgrowth. Ilataii is characterized by its
large. orbicular. thick shell, distinct
large lunule and rather shallow. narrow
pallial sinus being bluntly pointed at
end.
This is distinguishable from Dosinia/uganwna (=odosensis) by its large.
thick. orbicular shell. broad angle between postero-dorsal and ventral sides
and shape of pallial sinus which IS small
and narrow. Dosinia a11gu/oides No:>.Wl{A(1935b) also resembles the present one
but an;.;uloides can be distinguishedfrom the present one by its small. lessinflated shell. less protruded anterior
dorsal side and deep pallial sinus which
is rounded at end.
T_,pe locality, Geologiwl formation andAge.-1-lattomaki, Hanawa-machi. Higashi-Shirakawa-gun, Fukushima Prefecture (Lat. 36'58'45"N., Long. 140<25'10"
E.L Kubota formation. Middle Miocene.
Distribution :-Kubota formation. Fukushima
Prefecture; Kanomatazawa
formation, Tochigi Prefecture; Yamatsuda formation, Iwate Prefecture: alll\liddle !\liocenc in age.
.·lssociated fauna :-Chlamys (1\iimadt/amys' kaneharai (YOKOYAMA). "Palillopccten.. parap/ebejus (NorvlllRA and
HAT.\1.1, Jl1iyagiperten matsumoriensis
MASL;DA, Dosinia (Kaueharaia) kmzelwrai

YoKOYA~IA,

Tracltycardium shiobarense(YOKOYAl\IA). etc.
Subgenus Kanelurraia MAKJYAJ\IA, 1936

Dosinia (1\aneharaia) kamwi
MASUDA, n. sp.
Pl. 6. rigs. 6a-b. 7a-b

Dosinia (Kanelwraia) kaneharai. MAKIYAl\IA, 1Hem. Coil. Sci., Kyoto Imp. U11iv.,
Ser. B, Vol. 11, No. -1. Art. 8, p. 214.
pl. 4, fig. 2.
1936. Dosinia kanelwrai. NoMURA and ZINno,
Saito lfo-011 f(ai 1H11s., Res. Bull.. No.

1936.

I 0. p. 339, pl. 20, fig. I.

Dosinia kaneharai. OTVKA, Bull. Eartflq.
Res. lust .. Imp. Unit•. Tokyo, Vol. 14,
Pt. :J, p. 443. pl. 30. fig. 3.
1940. Dosinia kaneharai, No~rvRA, Sci. Rep.,
Tolwlm Imp. Uui11., Ser. 2. Vol. 21, No.

1936.

I. p. 26. pl. 1. figs. 15, 16.

1960.

Dosinia (Kaneharaia) kmwharai, KA;-.;:-:o.
japa11 Soc. Promo/. Sci .. p. 272. pl. 40.
fig. 9.

1961.

Dosi11ia (Ka11eharaia) ka11eharai, lwAI.
Trans. Proc. Palaeonl. Soc. ]apmz. 1\l. S ..
No. 11, pl. 1. fig. 1.

Holotypc :-IGPS, coli. cat. no. 64682.
Description :-Shell medium. moderate
in thickness and convexity, suborbicularin outline. nearly as high as long. An·
terior side moderately rounded. passing
gradually into broadly rounded ventral
margin; antero·dorsal side slightly concave in front of beak; postero-dorsalside broadly rounded, passing gradually
into ventral margin.
Beak pointed,
curved forward, situated anteriorly atabout 3/ 11 of shell length; lunule shallow,
elongate cordate in shape, rather obscurely defined; escutcheon not observed.
Surface sculptured with lo\v, rather re·gular, broad, somewhat rounded concentric growth lines and sometimes withinterstitial lines which are more slenderthan concentric lines; concentric lines

3G

Koichi1·o JL4.SUDA.

about II per LO mm. at 10 mm. from beakin central part of disc and separated byshallow grooves which are much nar·rower than concentric lines; concentriclines at antero- and postero·dorsal sides,
especially at posterior side. more elevated. more rounded and more slenderthan those of central part of disc, nearly
equal to their interspaces; concentriclines near ventral margin usually tend
to become numerous. slender and low;interstitial lines usually appear nearantero- and postero-dorsal sides and tend
to become stout towards central part ofdisc. Pallial sinus slightly sloping upwards, broad, rather shallow, nearly
triangular in shJpe, rounded at end.
Adductor muscle scars obscurely defined,
large. oblong; posterior one somewhat
larger than anterior one. Hinge verybroad ; anterior lateral tooth of left
valve small. rather low; median cardinalteeth distinctly elevated ; posterior one
somewhat elevated, slender; rather deep
and oblong pit at upper end of posterior
lateral socket distinctly defined.
Dimensions (in mm.) :Heigh; . I Length=:-·--I

Depth- ]

-~-

--

-~-

Ds/L

=

----·-

54.5*
55
I
17.5
0.34
45.5
15
57.5
59
17
53
15.5
45
47
13
0.34
- - - - - - - - - - - - - -I - - - • Holotype.

Renurrks :-This new species is named
in honor of Dr. Saburo KAN::\0 of theGeological and Mineralogical Institute.
Tokyo University of Education.
The present species is usually medium
in shell size but very rarely large ones
are found. This species is characterized
by its medium-sized shell, rather regular.
rounded, broad concentric growth lines,

obscurely defined lunule, no escutcheon.
broad, shallow pallial sinus whichrounded at end, and distinct, rather
deep pit at the end of posterior lateralsocket.
Dosinia (Kanelwraia) kaneharai (YOKO·YA~tA} (1926) is much related with thepresent one and the latter has hitherto
been included into the former. However. the present one can be distinguished from kmzeharai by its smaller shell.
regular concentric growth lines andbroad pallial sinus which is rounded at
end.
The present species occurs from ahorizon lower than that of kaneharai
and the former differs from kane/iarai
in the surface sculptures as mentioned
above. The concentric growth lines of
lumnoi are rather regular except for
those near the ventral margin but thoseof the latter are usually irregular except for at the upper part of disc. The
surface sculpture of the adult forms of
kamwi takes the sculpture of theyounger forms of kanelwrai. Therefore.
the writer is inclined to consider thatkamzoi is ancestral to lwnelwrai. Thismay be supported from the relation
between the geological horizon and thesize of shell, that is to say, the older
the geological age (horizon), the smaller
the size of the shells.
Type locality, Geological formation and
Age :-Kinseido, Kinsh6d6, Eihoku·men,
Kisshu-gun. Kankyo-hokudo, North Korea. lleiroku formation. Early Miocene.
Distribution :-Heiroku formation. Korea; Yanagawa formation, Fukushima
Prefecture; !\1oniwa member of the
Hatatate formation. \liyagi Prefecture;
Sugota formation, Akita Prefecture:
Saginosu formation. Saitama Prefecture;
Ainaigawa formation. Aomori Prefecture: all Early 1\liocene in age.
Associated fauna :-Anadara ogatcai

446.

(MAKIYAl\·IA),
Glcycymeris cisshuensis
MAKIYAMA,
Clilamys(Mimachlamys)
meisensis (iVIAKIYAMA), Pilar itoi lVIAKIYA~IA,:37

Fossil Dosinia

etc. at the type locality.
Remarks

Considering from the above mentioned
geographical distribution, geologicalrange and associated fauna of the fossil
species described in the present. article,
Dosinia akaisiana Nol'IIURA, Dosinia(Phacosoma) tugaruana NOt\IURA andDosinia (K.anelwmia) lumnoi i\IASUDA
are considered to be representatives ofthe lower horizon of the japanese l\lio·cene and while Dosinia (Piwcosoma)
ltataii MASUDA of a horizon higher thanthat of the former species. Furthermore,
it is considered that the horizon of
Dosinia akaisiana, Dosi11ia (Piwcosoma)
tugamana and Dosinia (Kaneltaraia)
kamwi may correspond to the Nanao·dzlamys notoensis assemblage (Earlyl\Iiocene) and that of Dosinia (Pitacosoma)
lwtaii to the Miyagipeclen matsumoriensis
assemblage (Middle l'vliocene) in Central
to Northern Japan (IvlASUDA. 1962). Thisview may be supported by the evolu·
tionary trend of Dosinia (Kane!zaraia)
kamwi and Dosinia (Kanelutraia) kanelwrai.
Dosinia matumotoi OTUKA (1938a, p.
11, pl. 1. fig. 10) was described from the
Miocene Shiroyama formation at Kanbara-machi, Ihara-gun, Sh izuoka Prefec·
lure based upon an imperfect specimen.
Howcyer. according to the writer's studyof the type specimen (Geological In·
stitute. Faculty of Science, Tokyo Uni·
versity, Reg. No. 3992) it is evident that
Dosinia matumotoi can not be included
into the genus Dosinia, because it shouldbe placed in the genus Lucinoma andmay be identified to Lucinoma acutili·
neatum (CONRAD).

References
IIATAI, K. (1938): On Some Species of Fossil
Dosinia from rJapan. japan. jour. Gcol.
Geogr .. Vol. 15, Nos. 1-2, pp. 47-65.
IWAI, T. (1961): The Miocene Molluscan
Fossils from the Area Southwest of Birosaki City, Aomori Prcfec:ture. Japan.
Trans. Proc. Palaeout. Soc. japan, N. S ..
No. 41, pp. 1-8. pl. 1, 1 text-fig., 1 tab.
KA;-.;No, S. (1960) : The Tertiary System of
the Chichibu Basin, Saitama Prefecture,
Central Japan. Part 2. Palaeontology.
japan Soc. Promo!. Sci., pp. 123-396, pls.
31-51, text-figs. 21-26. tables 5-18. 5 maps.
MAKIYA~IA, ]. (1936) : The !\1eisen Mioceneof North Korea. 1'vlem. Col! Sci., Kyoto
Imp. Unit•., Ser. B. Vol. 11. 11.:o. '1, !lrt. 8.
pp. 193-228, pis. 4-5. text-figs. 1-2.
MAst:oA. K. (1962) : Notes on the Tertiary
Pectinidae of Japan. Sci. Rep., Tohoku
Unio., Ser. 2. Spec. Vol. No. 5, pp. 159193, text-tigs. 1-9, tables l-9.
NAGAO. T. (1928): Paleogene Fossils of theIslands of Kyushu. Japan. Part 1. Sci.
Rep., To/wku Imp. Unil•., Ser. 2. Vol. 9.
~vo. 3, pp. 97-128. pis. 18-22.
No~tURA, S. (1935a): Miocene Mollusca fromthe Nisi-Tugaru District. .-\omori-ken,
Northeast IIonsyu, Japan. Saito Ho-011 KaiMus., Res. Bufl. 1\lo. 6, pp. 19-81, pis. 28, 1 map.
- - (l935b): Miocene Mollusca from Siogama. Northeast Honsyu, japan. Ibid,
pp. 193-234. pis. 16-17. 1 map.
- - (1940) : Molluscan Fauna of the Moniwa
Shell-Beds Exposed along the !\atori-gawain the Vicinity of Sendai. Miyagi Prefecture. Sci. Rep., Tohoku Imp. Unit•., Ser.
2. Vol. 21, No. 1. pp. 1-46, pis. 1-3.
- - and HATAI. K. (1936): Fossils from the
Tanagura Beds in the Vicinity of the
Town Tanagura. Ilukusima-ken. Northcast Honsyti, Japan. Saito Ho-011 KaiMus., Res. Bull., No. 10, pp. 109-155, pis.
13-17.
- - and Zt!';BO, N. (1936): Additional Fossil
Mollusca from the Yanagawa Shell-Beds
in the Hukusima Basin, Northeast Honsyti, japan. Ibid., No. 10, pp. 335-345. pl.

l\6ichil'6 J.l[ASUDll

38
20.

Y. (1934): Tertiary Structures of
the Northwestern End of the Kitakami
Mountainland. lwate Prefecture, Japan.
Bull. Earlhq. Res. /nsf., Imp. Unit•. Tollyo,
Vol 12, Pt. 3. pp. 566-638. pis. 47-51.
1 map.
- - (1936) : The Takasegawa Green Tuff
Beds of the Yuri District. Akita Pref.,Japan, and Correlation of the Lower Neo.
gene in Japan. Ibid .. Vol. I 4, Pl. 3, pp.
438-452, pl. 30.
- - (1938a): Neogene Fossils of the Ihara
District. Sizuolw Prefecture, Japan. four.
0TL11{A,l·itc. Sci., Imp. Univ. Tokyo. Sec. 2. Vol. 5,
1'1. 1, pp. 1-19. pis. 1-2, text-figs. 1-2.
tables 1-3.
- - (1938b): Mollusca from the Miocene ofTrugoku. Japan. Ibid., Pt. 2, pp. 21-45.
pis. 1-4. text-figs. 1-•l, tables 1-2.
REE\'E, L.A. (1851):
Monograph of theGenus Artemis. Conch. Icon., Vol. 6, pis.
1-10.
YoiWYA~IA, M. (1926):
Tertiary Mollusca
from Shiobara in Shimotsuke. ]our. Fac.
Sci., Imp. Unit•. Tokyo, Sec. 2, Vol. 1, Pt.
·1. pp. 127-138. pis. 16-20.

Explanation of Plate 6(All figures in naLural size unles!; otherwise stated)
Figs. la-c, 2. Dosinia (f'lwcosoma) hataii MASUDA, n. sp.
la, Right valve. b, Left valve. c, A part of the outer surface of 1a, x ca. 3. IIolotype,
IGPS. coli. cat. no. 72476. Loc. Hattomaki. IIanawa-machi. Higashi-Shirakawa-gun, Fukushima Prefecture. Kubota formation.
2, Hinge area of left Yalve. IGPS, coiL cat. no. 2645. Loc. Okada, Tanagura-machi.
Higashi-Shirakawa-gun, Fukushima Prefecture. Kubota format ion.
Figs. 3.. 5. Dosinia alwisiana No~WRA.
3. Right valve. Holotype. Si\I, Reg. No. 6061. 4, Lefl valve. 5. f~ight valve. Panttype,
SM, Reg. No. 6061. Loc. Hitotsumori, Ajigasawa-machi, Nishi-Tsugaru-gun, Aomori Prefecture. Tanosawa formation.
Figs. 6a-b, 7a-b. Dosinia (Kaneharaia) ka11110i MAsUDA, n. sp.
6a, Right valve. 6b, Inner surface of 6a. Holotype, IGPS, coli. cut. no. 64682. Loc. Kinseid6, Kinsh6d(), Eihoku-men. Kisshu-gun Kanky6-hokud6. Korea. lleiroku formation. 7a,Left valve. 7b, a part of the outer surface of 7a. X ca. 3. Paralype, IGPS, coli. cat. no64755. Loc. Same as above.
Figs. 8-lOa-b. Dosinia (Phacosomq) tugama11a No~tUI~A.
8, Inner mold of left va!Ye. Holotype of "Dosinia odosensis NoMURA •·. SM, Reg. No.
6020. Loc. ()dose, Fukaura-machi. Nishi-Tsugaru-gun, Aomori Prefecture. Tanosawa
formation. 9, Inner mold of left valve. Holotypc, SM. Reg. No. fi024. Loc. Hotatezawa.
lwasaki-murn, Nishi-Tsugaru-gun, Aomori Prefecture. Tanosawa formation. LOa. Plastercast of left valve. lOb. Plaster-cast of right val\'c. IGPS. coli. cat. no. 37060 Loc. Sameas fig. 8.
Fig. 11. Dosi11ia (Kattelwraia) kaneltarai YoKoYA~IA.
A part of outer surface. xca. 3. SM. Reg. No. 261·1. Loc. Same as Fig. 2.

MASUDA :MASUDA

Fossil Dosinia

photo.

Plate 6

39

PROCEEDINGS OF TilE PALAEONTOLOGICAL SOCIETY

OF JAPAN

tl :;{,;ti~F.#J~f:~~ 83 !ill~J~I-.l: 1962 iF-12 JJ 1 R
r.tr~ll1;:;k~'f:!1ll ~:1m J1!!ficf: M:¥~3.w::

rtJm.:r·rnPtrr.t;f.;P"Li}ll!ll\~tLf.: (~~-Jli31 £)o

m1

,,

;111~

llii

The Paleontologic and Stratigraphic Con·siderations on the ::\eoschwagerininae
and Verbeekininae, with the Descriptionsof Some Fusulinid Foraminifera fromthe Kitakami Massif ............ Shoshiro
HANZAWA ancl Masabumi MURATA
Permian Brachiopods from Central Thai·
land .................... Juichi YA~ACIDA
Halobia and some other Fossils from Kedah,
Malaya (l~i\J't) ........ Teiichi KollAYASIII
On the Cretaceous Ban Na Yo Fauna of
East Thailand with a Note on the Distribu·
tion of Nippononaia. Trigonioides andP/icalounio (~~) .... Teiichi 1\:0DA YASHI
On some Lower Cretaceous Trigonids fromCentral Japan (~~) ...... Shiro }.L\EDA
Gfycymeris and Cufteflus from the Tertiary
Hioki (Ashiya) Group in the Yuya-wan.
Yamaguchi Prefecture ............... .
.. Kazuo 0KA\IOTo and Mitsuo NAKANO
Pseudamimztis 1:. "'?~ '"( (ft~) ........ :6'1fili~~
Two Species of Taiuocera.~ from the Per·
mian of the Kitakami Mountains ..... .
....................... lchiro liAYASAKA~L1JI~Hlfil€<D(t.:.ti711 ;r. t:~IC-?L'""C .... iltj~'./ / ;f,:) '7 1-'Al¢[qJ1!!"1J.:t:t\J;L'c Lt.:ii1:JfOJ)"jlt1Cl~"T~P.,~~J
rjl~i1l!JJ1:l.9H~ 1:JEJC:6<7J'J' ~.(T .fL!ll. fl::GM 1'))¥-f

f1E · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · ;rjlftztE=
Upper Lower Carboniferous Brachiopods
from Akiyoshi .............. J. YA='AGIDA
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P "L .................... · · · · · · · · · ·l'f*lfi!!STwo Carboniferous Brachiopods from Loci.
Thailand .............. Takashi IlA\IADA
On the Triassic Dao11ella Beds in Central
Pahang. Malaya ...... Teiichi KoBA YASIII
Preliminary Report on the Lower CretaceousMarine Pelecypods in Japan ........... .
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.. Koichiro lolll\.t\W.-\ and Yasuo MAEDA

40
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