Appendix 1

Species list of Shiretoko Peninsula
and its coastal area

The Vascular Plants of Shiretoko Peninsula
From Sato, K. et al. (1985)* Recorded in the bibliography, although its presence was unclear** Red List categories of IUCN, details are given on the last page*** Red List Categories of Japanese Ministry of the Environment, details are given on the last page**** Endemic based on Satake et al. (1982; 1989) and Iwatsuki (1992), listed as "endemic to Japan" or "limited toJapan"IUCN RL **Scientific namePTERIDOPHYTA
EQUISETACEAE
Equisetum arvense
Equisetum arvense var. boreale
Equisetum hyemale
Equisetum palustre
Equisetum fluviatile
LYCOPODIACEAE* Lycopodium alpinum
Lycopodium annotinum
Lycopodium annotinum var. pungens
Lycopodium clavatum var. nipponicum
Lycopodium complanatum
Lycopodium obscurum forma strictum
Lycopodium obscurum forma flabellatum
Lycopodium selago var. appressum
Lycopodium selago var. miyoshianum
Lycopodium serratum
Lycopodium sitchense var. nikoense
SELAGINELLACEAE
Selaginella helvetica
Selaginella shakotanensis
Selaginella sibirica
Selaginella tamariscina
ISOETACEAE
Isoetes asiatica
OPHIOGLOSSACEAE
Botrychium lunaria
Botrychium robustum
OSMUNDACEAE
Osmunda asiatica
Osmunda japonica
HYMENOPHYLLACEAE
Mecodium wrightii
PTERIDACEAE
Adiantum pedatum
Coniogramme fraxinea var. intermedia
Dennstaedtia hirsuta* Dennstaedtia wilfordii
Pteridium aquilium var. latiusculum
PLAGIOLOGYRIACEAE
Plagiogyria matsumureana
ASPIDIACEAE
Athyrium alpestre
Athyrium crenulatoserrulatum
Athyrium filix-femina var. longipes
Athyrium melanolepis
Athyrium pterorachis
Athyrium pycnosorum
Athyrium vidalii
Athyrium vidalii var. yamadae
Athyrium yokoscense
Dryopteris amurensis

MoE RL ***

EN

VU

VU
VU

3

Endemic toJapan ****

* Recorded in the bibliography, although its presence was unclear** Red List categories of IUCN, details are given on the last page*** Red List Categories of Japanese Ministry of the Environment, details are given on the last page**** Endemic based on Satake et al. (1982; 1989) and Iwatsuki (1992), listed as "endemic to Japan" or "limited toJapan"IUCN RL **Scientific nameDryopteris austriaca
Dryopteris crassirhizoma
Dryopteris fragans
Dryopteris monticola
Lastrea dryopteris
Lastrea nipponica
Lastrea phegopteris
Lastrea quelpaertensis
Lastrea robertiana var. longula
Lastrea thelypteris
Matteuccia orientalis
Matteuccia struthiopteris
Onoclea sensibilis
Polystichum braunii
Polystichum retroso-paleaceum
Polystichum retroso-paleaceum var. coraiense
Polystichum tripteron
Rumohra miqueliana
Rumohra mutica
Rumohra standishii* Woodsia manchuriensis
Woodsia polystichoides
BLECHNACEAE
Blechnum nipponicum
ASPLENIACEAE
Asplenium incisum
Asplenium scolopendrium
POLYPODIACEAE
Pleopeltis ussuriensis var. distans
Polypodium fauriei
Polypodium virginianum
SPERMATOPHYTA
GYMNOSPERMAETAXACEAE
Taxus cuspidata
PINACEAE
Abies sachalinensis
Larix kaempferi
Picea glehnii
Picea jezoensis
Picea jezoensis var. takedae
Pinus pumila
CUPRESSACEAE
Juniperus chinensis var. sargentii
Juniperus communis var. montana
ANGIOSPERMAEDICOTYLEDONEAECHLORANTHACEAE
Chloranthus japonicus
Chloranthus serratus
SALICACEAE
Populus nigra var. italica
Populus maximowiczii
Populus tremula var. davidiana
Salix hultenii var. angustifolia
Salix pet-susu
Salix reinii

MoE RL ***Endemic toJapan ****

●

EN

4

* Recorded in the bibliography, although its presence was unclear** Red List categories of IUCN, details are given on the last page*** Red List Categories of Japanese Ministry of the Environment, details are given on the last page**** Endemic based on Satake et al. (1982; 1989) and Iwatsuki (1992), listed as "endemic to Japan" or "limited toJapan"IUCN RL **Scientific name
Salix reinii var. eriocarpa
Salix rorida
Salix sachalinensis
Salix taraikensis
Salix vulpina
Salix yezoalpina
Toisusu urbaniana
Toisusu urbaniana var. schneideri
JUGLANDACEAE
Juglans ailanthifolia
BETULACEAE
Alnus hirsuta
Alnus hirsuta var. sibirica
Alnus japonica
Alnus mayrii
Alnus maximowiczii
Betula ermanii
Betula maximowicziana
Betula platyphylla var. japonica
Carpinus cordata
Ostrya japonica
FAGACEAE
Quercus dentata
Quercus mongolica var. grosseserrata
ULMACEAE
Ulmus davidiana var. japonicus
Ulmus davidiana var. japonicus forma suberosa
Ulmus laciniata
MORACEAE
Humulus lupulus var. cordifolius
Morus bombycis
URTICACEAE
Boehmeria tricuspis
Boehmeria tricuspis var. unicuspis
Laportea bulbifera* Urtica angustifolia
Urtica platyphylla
SANTALACEAE
Thesium refractum
LORANTHACEAE
Viscum album var. coloratum forma rubro-auratiacum
ARISTOLOCHIACEAE
Asarum heterotropoides
POLYGONACEAE
Polygonum amphibium
Polygonum aviculare
Polygonum bistorta
Polygonum caespitosum var. laxiflorum
Polygonum convolvulus* Polygonum dentato-alatum* Polygonum dumetorum
Polygonum filiforme
Polygonum lapathifolium
Polygonum longisetum
Polygonum nepalense
Polygonum persicaria
Polygonum sachalinense5

MoE RL ***Endemic toJapan ****

VU
EN

●

* Recorded in the bibliography, although its presence was unclear** Red List categories of IUCN, details are given on the last page*** Red List Categories of Japanese Ministry of the Environment, details are given on the last page**** Endemic based on Satake et al. (1982; 1989) and Iwatsuki (1992), listed as "endemic to Japan" or "limited toJapan"IUCN RL **Scientific name
Polygonum thunbergii
Polygonum viviparum
Polygonum weyrichii
Polygonum weyrichii var. alpinum
Rumex acetosella
Rumex crispus
Rumex longifolius
Rumex montanus
Rumex obtusifolius
CHENOPODIACEAE
Atriplex gmelinii
Atriplex subcordata
Chenopodium album
Chenopodium album var. centrorubrum
Chenopodium glaucum
Kochia scoparia
Salsola komarovii
AMARANTHACEAE
Amaranthus lividus
CARYOPHYLLACEAE
Arenaria merckioides
Cerastium fischerianum
Cerastium holosteoides
Cerastium holosteoides var. hallaisanense
Cucubalus baccifer var. japonicus
Dianthus superbus
Dianthus superbus var. speciosus
Honkenya peploides var. major
Moehringia lateriflora
Sagina maxima forma crassicaulis
Silene alba
* Silene armeria
Silene noctiflora
Spergula arvensis
Spergula arvensis var. sativa
Stellaria fenzlii
Stellaria graminea
Stellaria media
Stellaria radians
Stellaria ruscifolia
NYMPHAEACEAE
Nuphar pumilum
Nymphaea tetragona
CERCIDIPHYLLACEAE
Cercidiphyllum japonicum
RANUNCULACEAE
Aconitum gigas
Aconitum maximum var. misaoanum
Aconitum maximum var. misaoanum forma album
Aconitum yesoense
Aconitum yesoense var. macroyesoense
Actaea asiatica
Actaea erythrocarpa
Adonis amurensis
Anemone debilis
Anemone flaccida
Anemone flaccida forma viridis

MoE RL ***Endemic toJapan ****

VU

VU
VU

●

LR/nt

VU

VU

6

* Recorded in the bibliography, although its presence was unclear** Red List categories of IUCN, details are given on the last page*** Red List Categories of Japanese Ministry of the Environment, details are given on the last page**** Endemic based on Satake et al. (1982; 1989) and Iwatsuki (1992), listed as "endemic to Japan" or "limited toJapan"IUCN RL **Scientific name
Anemone narcissiflora
Anemone yezoensis
Aquilegia buergeriana var. oxysepala
Aquilegia flabellata var. pumila
Caltha palustris var. barthei
Cimicifuga simplex
Clematis fusca
Clematis ochotensis
Coptis trifolia
Glaucidium palmatum
Ranunculus nipponicus var. major
Ranunculus quelpaertensis
Ranunculus repens
Ranunculus reptans
Ranunculus sceleratus
Thalictrum aquilegifolium
Thalictrum foetidum var. glabrescens
Thalictrum minus var. hypoleucum
Thalictrum minus var. stipellatum
Thalictrum sachalinense
Trollius riederianus
BERBERIDACEAE
Caulophyllum robustum
Diphylleia grayi
MAGNOLIACEAE
Magnolia kobus var. borealis
Magnolia obovata
Schisandra chinensis
PAPAVERACEAE
Chelidonium majus var. asiaticum
Corydalis ambigua
Corydalis speciosa
Dicentra peregrina
Dicentra peregrina forma alba
CRUCIFERAE
Arabis glabra
Arabis lyrata var. kamtschatica
Arabis pendula
Arabis serrata var. glauca
Arabis stelleri var. japonica
Barbarea orthoceras
Brassica napus
Capsella bursa-pastoris
Cardamine leucantha
Cardamine scutata
Cochlearia oblongifolia
Draba borealis
Erysimum cheiranthoides var. japonicum
Isatis tinctoria var. yezoensis
Macropodium pterospermum
Rorippa islandica
Rorippa sylvestris
DROSERACEAE
Drosera rotundifolia
CRASSULACEAE
Sedum aizoon
Sedum ishidae

MoE RL ***Endemic toJapan ****

EN

●

EN

VU

EN

CR

7

* Recorded in the bibliography, although its presence was unclear** Red List categories of IUCN, details are given on the last page*** Red List Categories of Japanese Ministry of the Environment, details are given on the last page**** Endemic based on Satake et al. (1982; 1989) and Iwatsuki (1992), listed as "endemic to Japan" or "limited toJapan"IUCN RL **Scientific name
Sedum iwarenge var. boehmeri
Sedum kamtschaticum* Sedum pluricaule
Sedum rosea
Sedum telephium var. purpureum
Sedum verticillatum
SAXIFRAGACEAE
Astilbe thunbergii var. congesta
Boykinia lycoctonifolia
Chrysosplenium flagelliferum
Chrysosplenium grayanum
Chrysosplenium kamtschaticum
Hydrangea paniculata
Hydrangea paniculata var. intermedia
Hydrangea petiolaris
Parnassia palustris
Parnassia palustris var. tenuis
Ribes latifolius
Ribes latifolius forma siretokense
Ribes sachalinense
Ribes triste
Saxifraga cherlerioides var. rebunshirensis
Saxifraga fortunei var. incisolobata
Saxifraga fusca
Saxifraga japonica
Saxifraga merkii
Saxifraga sachalinensis
Schizophragma hydrangeoides
Tiarella polyphylla
ROSACEAE
Agrimonia pilosa
Aruncus dioicus var. kamtschaticus
Filipendula multijuga var. ciliata
Filipendula kamtschatica
Fragaria ananassa
Fragaria iinumae
Fragaria yezoensis
Geum aleppicum
Geum calthaefolium var. nipponicum
Geum macrophyllum
Geum pentapetalum
Potentilla centigranaPotentilla dickinsii
Potentilla egedei var. groenlandica
Potentilla fragarioides var. major
Potentilla matsumurae
Potentilla megalantha
Potentilla miyabei
Potentilla nipponica
Potentilla norvegica
Potentilla palustris
Potentilla stolonifera
Prunus maximowiczii
Prunus maximowiczii forma rotundifolia
Prunus nipponica
Prunus nipponica var. kurilensis* Prunus padus

MoE RL ***
VU

CR

8

Endemic toJapan ****

* Recorded in the bibliography, although its presence was unclear** Red List categories of IUCN, details are given on the last page*** Red List Categories of Japanese Ministry of the Environment, details are given on the last page**** Endemic based on Satake et al. (1982; 1989) and Iwatsuki (1992), listed as "endemic to Japan" or "limited toJapan"IUCN RL **Scientific name
Prunus sargentii
Prunus ssiori
Rosa acicularis
Rosa rugosa
Rubus crataegifolius
Rubus idaeus var. aculeatissimus
Rubus idaeus var. aculeatissimus forma concolor
Rubus parvifolius
Rubus pedatus
Rubus phoenicolasius
Rubus pseudo-japonicus
Sanguisorba stipulata
Sanguisorba tenuifolia var. alba
Sanguisorba tenuifolia var. grandiflora
Sorbaria sorbifolia var. stellipila
Sorbus alnifolia
Sorbus commixta
Sorbus matsumurana
Sorbus sambucifolia
Spiraea betulifolia
Spiraea betulifolia var. aemeliana
Spiraea betulifolia var. aemeliana forma glabra
Spiraea media var. sericea
Waldsteinia ternata
LEGUMINOSAE
Astragalus japonicus
Desmodium oxyphyllum
Hedysarum hedysaroides
Lathyrus japonicus
Lathyrus japonicus var. aleuticus
Lathyrus japonicus var. aleuticus forma albinus
Lespedeza bicolor* Lotus cornicularis var. japonicus
Maackia tashiroi
Melilotus alba
Robinia pseudo-acasia
Thermopsis lupinoides
Trifolium hybridum
Trifolium lupinaster
Trifolium pratense
Trifolium repens
Vicia amoena
Vicia cracca
Vicia cracca forma canescens
Vicia japonica
Vicia unijuga
GERANIACEAE
Geranium erianthum* Geranium eriostemon var. reinii
Geranium krameri
Geranium sibiricum var. glabrius
Geranium thunbergii
Geranium yezoense
Geranium yezoense var. pseudo-palustre
OXALIDACEAE
Oxalis acetosella
Oxalis stricta

MoE RL ***

EN

CR
VU

9

Endemic toJapan ****

* Recorded in the bibliography, although its presence was unclear** Red List categories of IUCN, details are given on the last page*** Red List Categories of Japanese Ministry of the Environment, details are given on the last page**** Endemic based on Satake et al. (1982; 1989) and Iwatsuki (1992), listed as "endemic to Japan" or "limited toJapan"IUCN RL **Scientific name
RUTACEAE
Phellodendron amurense var. sachalinense
Skimmia japonica var. intermedia
Skimmia japonica var. intermedia forma repens
SIMAROUBACEAE
Picrasma quassioides
EUPHORBIACEAE
Daphniphyllum macropodum var. humile
BUXACEAE
Pachysandra terminalis
EMPETRACEAE
Empetrum nigrum var. japonicum
ANACARDIACEAE
Rhus ambigua
Rhus trichocarpa
AQUIFOLIACEAE
Ilex crenata var. paludosa
Ilex rugosa
Ilex sugerokii
CELASTRACEAE
Celastrus orbiculatus var. papillosus
Euonymus alatus forma ciliatodentatus
Euonymus macropterus
Euonymus oxyphyllus
Euonymus sieboldianus
STAPHYLEACEAE
Staphylea bumalda
ACERACEAE
Acer japonicum
Acer monoAcer mono var. mayrii
Acer tschonoskii
Acer ukurunduense
Acer ukurunduense var. pilosum
BALSAMINACEAE
Impatiens noli-tangere* Impatiens textori
VITACEAE
Ampelopsis brevipedunculata
Vitis coignetiae
TILIACEAE
Tilia japonica
Tilia maximowicziana
Tilia maximowicziana var. yesoana
ACTINIDIACEAE
Actinidia arguta var. platyphylla
Actinidia kolomikta
Actinidia polygama
GUTTIFERAE
Hypericum ascyron
Hypericum erectum
Hypericum kamtschaticum
Hypericum yezoense
VIOLACEAE
Viola acuminata
Viola blandaeformis
Viola blandaeformis var. pilosa

MoE RL ***Endemic toJapan ****

●
●
●

VU

VU

10

* Recorded in the bibliography, although its presence was unclear** Red List categories of IUCN, details are given on the last page*** Red List Categories of Japanese Ministry of the Environment, details are given on the last page**** Endemic based on Satake et al. (1982; 1989) and Iwatsuki (1992), listed as "endemic to Japan" or "limited toJapan"IUCN RL **Scientific name
Viola collina* Viola grypoceras
Viola kitamiana
Viola kusanoana
Viola langsdorffii
Viola langsdorffii forma parviflora
Viola mandshurica
Viola repens
Viola sacchalinensis
Viola selkirkii
Viola verecunda
Viola verecunda var. semilunaris
THYMELAEACEAE
Daphne kamtschatica var. jezoensis
Daphne miyabeana
LYTHRACEAE
Lythrum salicaria
ONAGRACEAE
Circaea alpina
Epilobium amurense
Epilobium angustifolium
Epilobium cephalostigma
Epilobium cephalostigma var. nudicarpum
Epilobium dielsii
Epilobium fauriei
Epilobium foucaudianum
Epilobium glandulosum var. asiaticum
Oenothera muricata
ARALIACEAE
Aralia cordata
Aralia elata
Kalopanax pictus
UMBELLIFERAE
Aegopodium alpestre
Angelica anomala
Angelica edulisAngelica genuflexa
Angelica ursina
Anthriscus sylvestris
Bupleurum longiradiatum var. breviradiatum* Bupleurum longiradiatum var. shikotanense
Chamaele decumbens
Cicuta virosa
Coelopleurum lucidum var. gmelinii
Conioselinum filicinum
Conioselinum kamtschaticum
Cryptotaenia japonica* Daucus carota
Glehnia littoralis
Heracleum dulce
Hydrocotyle ramiflora
Ligusticum hultenii
Oenanthe javanica
Osmorhiza aristata
Peucedanum multivittatum
Peucedanum terebinthaceum
Pleurospermum camtschaticum

MoE RL ***Endemic toJapan ****

●
VU

CR

●

●

11

* Recorded in the bibliography, although its presence was unclear** Red List categories of IUCN, details are given on the last page*** Red List Categories of Japanese Ministry of the Environment, details are given on the last page**** Endemic based on Satake et al. (1982; 1989) and Iwatsuki (1992), listed as "endemic to Japan" or "limited toJapan"IUCN RL **Scientific name
Sanicula chinensis* Sium suave var. nipponicum
Spuriopimpinella calycina
Tilingia ajanensis
Tilingia holopetala
Torilis japonica
CORNACEAE
Cornus canadensis
Cornus controversa
DIAPENSIACEAE
Diapensia lapponica var. obovata
PYROLACEAE
Chimaphila japonica
Monotropastrum globosum
Pyrola alpina
Pyrola faurieana
Pyrola japonica forma subaphylla
Pyrola renifolia
Pyrola secunda
ERICACEAE
Andromeda polifolia
Arcterica nana
Arctous alpinus var. japonicus
Bryanthus gmelinii
Cassiope lycopodioides* Gaultheria adenothrix
Gaultheria miqueliana
Harrimanella stelleriana
Ledum palustre var. diversipilosum
Leucothoe grayana var. oblongifolia
Leucothoe grayana var. yezoensis
Loiseleuria procumbens
Menziesia pentandra
Phyllodoce aleutica
Phyllodoce caerulea
Phyllodoce caerulea forma yezoensis
Rhododendron aureum
Rhododendron brachycarpum
Rhododendron camtschaticum
Tripetaleia bracteata
Vaccinium hirtum
Vaccinium japonicum
Vaccinium ovalifolium
Vaccinium ovalifolium var. coriaceum
Vaccinium oxycoccus
Vaccinium praestans
Vaccinium smallii
Vaccinium uliginosum
Vaccinium vitis-idaea
PRIMULACEAE
Glaux maritima var. obtusifolia
Lysimachia thyrsiflora
Lysimachia vulgaris var. davurica
Primula cuneifolia
Primula cuneifolia forma leucantha
Primula modesta var. fauriei
Primula modesta var. matsumurae

MoE RL ***Endemic toJapan ****

EN
●
●

EN

VU

VU

12

* Recorded in the bibliography, although its presence was unclear** Red List categories of IUCN, details are given on the last page*** Red List Categories of Japanese Ministry of the Environment, details are given on the last page**** Endemic based on Satake et al. (1982; 1989) and Iwatsuki (1992), listed as "endemic to Japan" or "limited toJapan"IUCN RL **Scientific namePrimula jesoana var. pubescens
Trientalis europaea
Trientalis europaea var. arctica
OLEACEAE
Fraxinus lanuginosa
Fraxinus mandshurica var. japonica
Ligustrum tshonoskii var. glabrescens
Syringa reticulata
Syringa reticulata var. macrophylla
Syringa reticulata var. tatewakiana
GENTIANACEAE
Fauria crista-galli
Gentiana triflora var. japonica
Gentiana triflora var. horomuiensis
Gentiana triflora var. montana
Gentiana zollingeri
Halenia corniculata
Menyanthes trifoliata
Swertia tetrapetala
Tripterospermum japonicum
APOCYNACEAE
Apocynum venetum var. basikurumon
ASCLEPIADACEAE
Cynanchum caudatum
CONVOLVULACEAE* Calystegia japonica
Calystegia sepium var. americana
Calystegia soldanella
BORAGINACEAE
Mertensia asiatica
Myosotis scorpioides
LABIATAE
Agastache rugosa
Agastache rugosa forma hypoleuca
Clinopodium chinense var. parviflorum
Clinopodium chinense var. shibetchense
Clinopodium gracile var. sachalinense
Dracocephalum argunense
Elsholtzia ciliata
Galeopsis bifida
Lamium album var. barbatum
Lycopus uniflorus
Mentha arvensis var. piperascens
Nepeta subsessilis var. yesoensis
Prunella vulgaris var. lilacina
Scutellaria dependens
Scutellaria pekinensis var. ussuriensis
Scutellaria pekinensis var. transitra
Scutellaria strigillosa
Stachys japonica var. villosa
Teucrium viscidum var. miquelianum* Teucrium teinense
Thymus quinquecostatus
SOLANACEAE
Solanum megacarpum
Solanum nigrum

MoE RL ***

EN

EN

13

Endemic toJapan ****

* Recorded in the bibliography, although its presence was unclear** Red List categories of IUCN, details are given on the last page*** Red List Categories of Japanese Ministry of the Environment, details are given on the last page**** Endemic based on Satake et al. (1982; 1989) and Iwatsuki (1992), listed as "endemic to Japan" or "limited toJapan"IUCN RL **Scientific name
SCROPHULARIACEAE
Linaria japonica
Mimulus nepalensis var. japonica
Mimulus sessilifolius
Pedicularis apodochila
Pedicularis chamissonis var. japonica
Pedicularis respinata
Pedicularis schistostegia* Pedicularis yezoensis
Pentstemon frutescens
Scrophularia grayana
Verbascum thapsus
Veronica americana
Veronica persica
Veronica schmidtiana
Veronica schmidtiana var. lineariloba
Veronica stelleri var. longistyla
Veronicastrum sibiricum
OROBANCHACEAE
Orobanche coerulescens
LENTIBULARIACEAE
Urticularia japonica
PHRYMACEAE
Phryma leptostachya var. asiatica
PLANTAGINACEAE
Plantago asiatica
Plantago camtschatica
Plantago lanceolata
Plantago major var. japonica forma yezomaritima
RUBIACEAE
Asperula odorata
Galium kamtschaticum
Galium kamtschaticum var. acutifolium* Galium mollugo
Galium trifidum var. brevipedunculatum
Galium trifloriforme
Galium verum var. asiaticum
Galium verum var. trachycarpum
Rubia jesoensis
CAPRIFOLIACEAE
Linnaea borealis
Lonicera alpigena var. glehnii
Lonicera caerulea var. edulis
Lonicera caerulea var. emphyllocalyx
Lonicera chamissoi
Lonicera chrysantha
Lonicera sachalinensis
Sambucus sieboldiana var. miquelii
Viburnum furcatum
Viburnum opulus var. calvescens
Viburnum wrightii
Weigela middendorffiana
ADOXACEAE
Adoxa moschatellina
VALERIANACEAE
Patrinia gibbosa
Patrinia scabiosaefolia

MoE RL ***

VU

VU

VU

EN

VU
VU
VU

14

Endemic toJapan ****

* Recorded in the bibliography, although its presence was unclear** Red List categories of IUCN, details are given on the last page*** Red List Categories of Japanese Ministry of the Environment, details are given on the last page**** Endemic based on Satake et al. (1982; 1989) and Iwatsuki (1992), listed as "endemic to Japan" or "limited toJapan"IUCN RL **Scientific name
Patrinia sibirica
Patrinia villosa
Valeriana fauriei
CUCURBITACEAE
Gynostemma pentaphyllum
Schizopepon bryoniaefolius
CAMPANULACEAE
Adenophora pereskiaefolia var. heterotricha
Adenophora triphylla var. japonica
Campanula chamissonis
Campanula lasiocarpa
Lobelia sessilifolia
Peracarpa carnosa var. circaeoides
COMPOSITAE
Achillea millefolium
Achillea ptarmica var. macrocephala
Achillea sibirica
Adenocaulon himalaicum
Anaphalis margaritacea var. angustior
Arnica unalascensis
Artemisia arctica
Artemisia iwayamogi
Artemisia japonica
Artemisia japonica forma resedifolia
Artemisia japonica var. macrocephala
Artemisia keiskeana
Artemisia koidzumii
Artemisia koidzumii var. tsuneoi
Artemisia laciniata
Artemisia montana
Artemisia montana var. shiretokoensis
Artemisia schmidtiana
Artemisia stelleriana
Aster glehnii
Aster scaber
Bidens radiata var. pinnatifida
Breea setosa
Cacalia auriculata var. kamtschatica
Cacalia hastata var. orientalis
Carpesium triste
Chrysanthemum leucanthemum
Chrysanthemum vulgare var. boreale
Chrysanthemum yezoense
Cirsium kamtschaticum* Cirsium pectinellum
Cirsium pectinellum var. alpinum* Cirsium pendulum* Cirsium vulgare
Erigeron acris
Erigeron acris var. amplifolius
Erigeron annuus
Erigeron canadensis
Erigeron thunbergii var. glabratus
Eupatorium chinense var. sachalinense
Gnaphalium sylvaticum
Hieracium aurantiacum
Hieracium umbellatum var. japonicum

MoE RL ***
EN

VU
VU

VU

15

Endemic toJapan ****

* Recorded in the bibliography, although its presence was unclear** Red List categories of IUCN, details are given on the last page*** Red List Categories of Japanese Ministry of the Environment, details are given on the last page**** Endemic based on Satake et al. (1982; 1989) and Iwatsuki (1992), listed as "endemic to Japan" or "limited toJapan"IUCN RL **Scientific name* Hypochoeris radicata
Inula salicina var. asiatica
Ixeris dentata
Ixeris dentata var. albiflora
Ixeris repens
Lactuca indica var. laciniata
Lactuca raddeana var. elata
Leibnitzia anandra
Ligularia hodgsonii
Matricaria inodora
Matricaria matricarioides
Matricaria tetragonosperma
Petasites japonicus var. giganteus
Picris hieracioides var. glabrescens
Picris hieracioides var. alpina
Rudbeckia serotina
Saussurea riederi var. yezoensis
Senecio cannabifolius
Senecio nemorensis
Senecio pseudo-arnica
Senecio vulgaris
Solidago virga-aurea var. leiocarpa
Sonchus asper
Sonchus brachyotis
Sonchus oleraceus
Taraxacum laevigatumTaraxacum officinale
Taraxacum shikotanense
MONOCOTYLEDONEAE
TYPHACEAE
Typha latifolia
SPARGANIACEAE
Sparganium hyperboreum
Sparganium sp.
POTAMOGETONACEAE
Potamogeton fryeri
Potamogeton natans
ZOSTERACEAE
Zostera nana
SCHEUCHZERIACEAE
Scheuchzeria palustris
GRAMINEAE
Achnatherum pekinense
Agrostis alba
Agrostis borealis
Agrostis clavata
Agrostis flaccida
Agrostis scabra
Agrostis stolonifera
Alopecurus aequalis var. amurensis
Anthoxanthum odoratum
Beckmannia syzigachne
Brachypodium sylvaticum
Bromus japonicus
Bromus mollis
Bromus pauciflorus
Brylkinia caudata

MoE RL ***

EN

EN

16

Endemic toJapan ****

* Recorded in the bibliography, although its presence was unclear** Red List categories of IUCN, details are given on the last page*** Red List Categories of Japanese Ministry of the Environment, details are given on the last page**** Endemic based on Satake et al. (1982; 1989) and Iwatsuki (1992), listed as "endemic to Japan" or "limited toJapan"IUCN RL **Scientific nameCalamagrostis epigeios
Calamagrostis hakonensis
Calamagrostis langsdorffii
Calamagrostis neglecta var. aculeolata
Calamagrostis purpurascens
Calamagrostis sachalinensis
Dactylis glomerata
Deschampsia caespitosa var. festucaefolia
Deschampsia flexuosa
Echinochloa crus-galli
Elymus dahuricus
Elymus mollis
Elymus sibiricus
Festuca elatiorFestuca ovina
Festuca rubra
Glyceria alnastretum
Glyceria leptolepis
Glyceria lithuanica
Hierochloe alpina
Hierochloe odorata var. pubescens
Koeleria cristata
Melica nutans
Milium effusum
Miscanthus sinensis
Muhlenbergia curviaristata var. nipponica* Muhlenbergia japonica
Panicum bisulcatum
Phalaris aruundinacea
Phleum pratense
Phragmites communis
Poa annua
Poa hisauchii
Poa macrocalyx
Poa macrocalyx var. scabriflora
Poa macrocalyx var. tatewakiana
Poa pratensis
Poa radula
Poa sachalinensis
Poa trivialis
Poa viridula
Puccinellia pumila
Sasa cernea
Sasa kurilensis
Sasa senanensis
Sasa sendaica
Sasa shiretokoensis
Setaria faberi
Setaria viridis var. pachystachys
Trisetum sibiricum
CYPERACEAE
Carex aphanolepis
Carex augustinowiczii
Carex augustinowiczii var. sharensis
Carex bigelowii
Carex blepharicarpa
Carex breviculmis

17

MoE RL ***Endemic toJapan ****

* Recorded in the bibliography, although its presence was unclear** Red List categories of IUCN, details are given on the last page*** Red List Categories of Japanese Ministry of the Environment, details are given on the last page**** Endemic based on Satake et al. (1982; 1989) and Iwatsuki (1992), listed as "endemic to Japan" or "limited toJapan"IUCN RL **Scientific name
Carex brunnescens
Carex buxbaumii
Carex caryophyllea var. microtricha
Carex conica
Carex curta
Carex curvicollis* Carex dispalata
Carex dissitiflora
Carex echinata
Carex flavocuspis
Carex foliosissima
Carex gmelinii
Carex hakkodensis
Carex hakonensis
Carex humilis var. nana
Carex incisa
Carex insaniae
Carex jacens
Carex japonica
Carex lanceolata
Carex lasiocarpa var. occultans
Carex limosa
Carex longerostrata
Carex lyngbyei
Carex macrocephala
Carex mertensii var. urostachys
Carex michauxiana var. asiatica
Carex middendorffii
Carex mollicula
Carex omiana var. monticola
Carex oxyandra
Carex parciflora
Carex pauciflora
Carex pilosa
Carex puberula
Carex pumila
Carex pyrenaica
Carex remotiuscula
Carex rhynchophysa
Carex sachalinensis
Carex sadoensis
Carex schmidtii
Carex scita var. riishirensis
Carex scita var. scabrinervia
Carex shimidzensis
Carex stenantha var. taisetsuensis
Carex stipata
Carex subspathacea
Carex thunbergii var. appendiculata
Carex traiziscana
Carex vanheurckii
Carex vesicaria
Eleocharis congesta var. thermalis
Eleocharis intersita
Eleocharis margaritacea
Eriophorum vaginatum
Rhynchospora alba

MoE RL ***
EN

VU

EN

VU

CR
NT
VU

EN

18

Endemic toJapan ****

* Recorded in the bibliography, although its presence was unclear** Red List categories of IUCN, details are given on the last page*** Red List Categories of Japanese Ministry of the Environment, details are given on the last page**** Endemic based on Satake et al. (1982; 1989) and Iwatsuki (1992), listed as "endemic to Japan" or "limited toJapan"IUCN RL **Scientific name
Scirpus caespitosus
Scirpus maximowiczii
Scirpus wichurae
ARACEAE
Arisaema angustatum var. penisulae
Lysichiton camtschatcense
Symplocarpus renifolius
COMMELINACEAE
Commelina communis
JUNCACEAE
Juncus beringensis
Juncus effusus var. decipiens
Juncus effusus var. decipiens forma gracilis
Juncus filiformis
Juncus gracillimus
Juncus haenkei
Juncus kamtschatcensis
Juncus krameri
Juncus mertensianus
Juncus prominens
Juncus tenuis
Juncus wallichianus
Luzula capitata
Luzula multiflora
Luzula oligantha
Luzula plumosa var. macrocarpa
Luzula rostrata
Luzula wahlenbergii
LILIACEAE
Allium schoenoprasum
Allium schoenoprasum var. yezomonticola
Allium splendens
Allium victorialis var. platyphyllum
Asparagus schoberioides
Clintonia udensis
Convallaria keiskei
Disporum sessile
Disporum smilacinum
Fritillaria camtschatcensis* Gagea lutea
Gagea vaginata
Heloniopsis orientalis
Hemerocallis middendorffii
Hemerocallis yezoensis
Hosta rectifolia
Lilium cordatum var. glehnii
Lilium maculatum var. dauricum
Lilium medeoloides
Lilium medeoloides var. kurilense* Lloydia serotina
Maianthemum dilatatum
Paris verticillata
Polygonatum humile
Polygonatum odoratum var. maximowiczii
Smilacina japonica
Streptopus amplexifolius var. papillatus
Streptopus streptopoides

MoE RL ***
VU

EN

NT

VU

19

Endemic toJapan ****

* Recorded in the bibliography, although its presence was unclear** Red List categories of IUCN, details are given on the last page*** Red List Categories of Japanese Ministry of the Environment, details are given on the last page**** Endemic based on Satake et al. (1982; 1989) and Iwatsuki (1992), listed as "endemic to Japan" or "limited toJapan"IUCN RL **Scientific name
Tofieldia coccinea
Tofieldia coccinea var. fusca* Tofieldia okuboi
Trillium apetalon
Trillium apetalon var. atropurpureocarpum
Trillium hagae
Trillium kamtschaticum
Trillium tschonoskii
Veratrum grandiflorum
IRIDACEAE
Iris ensata var. spontanea
Iris setosa
ORCHIDACEAE
Calanthe nipponica
Calanthe tricariata* Cephalanthera erecta
Cephalanthera longibracteata* Coeloglossum viride var. bracteatum
Cremastra appendiculata
Ephippianthus schmidtii
Epipactis papillosa
Gastrodia elata
Goodyera maximowiczianum
Goodyera repens
Gymnadenia camtschatica* Gymnadenia cucullata
Liparis makinoana
Listera cordata
Listera pinetorum
Microstylis monophyllos
Myrmechis japonica
Orchis aristata
Oreorchis patens
Platanthera chorisiana
Platanthera metabifolia
Platanthera phrydioides
Platanthera phrydioides var. takedae
Platanthera sachalinensis
Platanthera tipuloides
Spiranthes sinensis

MoE RL ***Endemic toJapan ****

CR

EN
EN

VU

References

Iwatsuki, K. (1992) Ferns and Fern Allies of Japan. HEIBONSHA Ltd., Publishers, Tokyo (in japanese).
Sato, K., Nishikawa, T., Sakai, S., Matsui, K., Kohyama, T., Koike, F., Kobayashi, M. and Ito, K. (1985)
Flora of Vascular Plants of the Onnebetsudake Wilderness area and Shiretoko Peninsula. In: Conservation
Reports of the Onnebetsu-dake Wilderness Area, Hokkaido, Japan, pp.115-172. Nature Conservation
Bureau, Environment Agency, Japan (in Japanese with English summary).
Satake, Y., Hara, H., Watari, A. and Tominari, T. (1989) Wild Flowers of Japan, Wood Plants.
HEIBONSHA Ltd., Publishers, Tokyo (in Japanese).
Satake, Y., Ohwi, J., Kitamura, S., Watari, A. and Tominari, T. (1982) WildFlowers of Japan.
HEIBONSHA Ltd., Publishers, Tokyo(in Japanese).

20

Other Vascular Plants Recorded on Shiretoko
Peninsula
From the Records of Shiretoko Museum (July 2003) except species listed by Sato, K. et al. (1985).
*** Red List Categories of Japanese Ministry of the Environment, details are given on the last page**** Endemic based on Satake et al. (1982; 1989) and Iwatsuki (1992), listed as "endemic to Japan" or "limited
to Japan"
MoE RL ***Scientific namePTERIDACEAE
Pleurosoriopsis makinoi
ASPIDIACEAE
Athyrium neglectum
Dryopteris coreano-montana
Polystichum microchlamys
Polystichum craspedosorum
Leptogramma pozoi ssp. mollissima
Cystopteris fragilis
ASPLENIACEAE
Asplenium trichomanes
CARYOPHYLLACEAE
Silene dichotoma
Stellaria sessilifolia
RANUNCULACEAE
Anemone dichotoma
Ranunculus silerifolius
VIOLACEAE
Viola biflora
Viola patrinii
ONAGRACEAE
Epilobium pyrricholophum
Oenothera erythrosepala
Oenothera biennis
PYROLACEAE
Pyrola japonica
GENTIANACEAE
Pterygocalyx volubilis
BORAGINACEAE
Cynoglossum asperrimum
Mysotis arvensis
Symphytum officinale
LABIATAE
Glechoma hederacea var. grandis
SCROPHULARIACEAE
Veronica tenella
COMPOSITAE
Crepis tectorum
SPARGANIACEAE
Sparganium glomeratum
Sparganium angustifolium
ZOSTERACEAE
Phyllospadix iwatensis
Zostera marina
SCHEUCHZERIACEAE
Triglochin palustre
CYPERACEAE
Carex breviculmis var. discoidea
ARACEAE
Symplocarpus nipponicus
IRIDACEAE
Iris pseudacorus

Endemic toJapan ****

●

VU

EN

VU
VU

VU

21

*** Red List Categories of Japanese Ministry of the Environment, details are given on the last page**** Endemic based on Satake et al. (1982; 1989) and Iwatsuki (1992), listed as "endemic to Japan" or "limited
to Japan"
MoE RL ***Scientific name
ORCHIDACEAE
Amitostigma kinoshitae
Dactylostalix ringens
Neottia asiatica
Neottia japonica
Pogonia japoncia
LEMNACEAE
Spirodela polyrhiza

Endemic toJapan ****

VU
VU
VU

References

Iwatsuki, K. (1992) Ferns and Fern Allies of Japan. HEIBONSHA Ltd., Publishers, Tokyo (in japanese).
Sato, K., Nishikawa, T., Sakai, S., Matsui, K., Kohyama, T., Koike, F., Kobayashi, M. and Ito, K. (1985)
Flora of Vascular Plants of the Onnebetsudake Wilderness area and Shiretoko Peninsula.
In: Conservation Reports of the Onnebetsu-dake Wilderness Area, Hokkaido, Japan, pp.115-172.
Nature Conservation Bureau, Environment Agency, Japan (in Japanese with English summary).
Satake, Y., Hara, H., Watari, A. and Tominari, T. (1989) Wild Flowers of Japan, Wood Plants.
HEIBONSHA Ltd., Publishers, Tokyo (in Japanese).
Satake, Y., Ohwi, J., Kitamura, S., Watari, A. and Tominari, T. (1982) WildFlowers of Japan. HEIBONSHA
Ltd., Publishers, Tokyo(in Japanese).

22

The Seaweeds of Shiretoko Peninsula and its
Coastal Area

From Kurogi et al. (1985) and Yamada and Tanaka (1944)**** Data Book Categories of Japanese Fisheries Agency, details are given on the last page
Scientific name
CHLOROPHYCEAECHLOROCOCCALES
Endosphaeraceae
Chlorochytrium inclusum
Codiolum gregarium
ULOTRICHALES
Ulotrichaceae
Ulothrix flacca
Ulothrix implexa
Ulothrix flexuosa
ULVALES
Capsosiphonaceae
Capsosiphon groenlandicus
Kornmanniaceae
Kornmannia leptoderma
Monostromataceae
Monostroma angicava
Monostroma crassidermum
Protomonostroma undulatum
Ulvaceae
Blidingia minima
Enteromorpha compressa
Enteromorpha intestinalis
Enteromorpha linza
Ulva pertusa
Ulvaria fusca
CLADOPHORALES
Cladophoraceae
Chaetomorpha aerea
Chaetomorpha moniligera
Cladophora arenaria
Cladophora opaca
Cladophora stimpsonii
Rhizoclonium tortuosum
Rhizoclonium lubricum
ACROSIPHONIALES
Acrosiphoniaceae
Spongomorpha heterocladia
Spongomorpha duriuscula var. cartilaginea
Urospora penicilliformis
Urospora wormskioldii
CODIALES
Codiaceae
Codium yezoense
BRYOPSIDALES
Derbesiaceae
Derbesia marina
PHAEOPHYCEAEECTOCARPALES
Pilayellaceae
Pilayella littoralis
RALFSIALES
Ralfsiaceae
Analipus filiformis
Analipus japonicus
Ralfsia fungiformis

23

FA DB****

R

**** Data Book Categories of Japanese Fisheries Agency, details are given on the last page
FA DB****

Scientific name
SPHACELARIALES
Sphacelariaceae
Sphacelaria rigidula
DICTYOTALES
Dictyotaceae
Dictyopteris divaricata
Dictyota dichotoma
CHORDARIALES
Acrotrichaceae
Acrothrix pacifica
Chordariaceae
Chordaria flagelliformis
Saundersella simplex
Sphaerotrichia divaricata
Elachistaceae
Halothrix ambigua
Leathesiaceae
Leathesia difformis
Leathesia sphaerocephala
DICTYOSIPHONALES
Coilodesmaceae
Akkesiphycus lubricum
Coilodesme cystoseirae
Coilodesme japonica
Dictyosiphonaceae
Dictyosiphon chordaria
Dictyosiphon foeniculaceus
Punctariaceae
Punctaria latifolia
SCYTOSIPHONALES
Scytosiphonaceae
Colpomenia bullosa
Colpomenia sinuosa
Petalonia binghamiae
Petalonia fascia
Scytosiphon lomentaria
DESMARESTIALES
Desmarestiaceae
Desmarestia ligulata
Desmarestia viridis
LAMINARIALES
Alariaceae
Alaria angusta
Chordaceae
Chorda filum
Laminariaceae
Agarum clathratum
Costaria costata
Cymathaere japonica
Laminaria diabolica
Laminaria ochotensis
Laminaria sachalinensis
FUCALES
Fucaceae
Fucus distichus subsp. evanescens
Silvetia babingtonii
Sargassaceae
Cystoseira crassipes
Cystoseira hakodatensis
Sargassum confusum
Sargassum miyabei
Sargassum thunbergii

R

R

24

**** Data Book Categories of Japanese Fisheries Agency, details are given on the last page
FA DB****

Scientific nameRHODOPHYCEAEGONIOTRICHALES
Goniotrichaceae
Stylonema alsidii
ERYTHROPELTIDALES
Erythropeltidaceae
Erythrotrichia carnea
BANGIALES
Bangiaceae
Porphyra crassa
Porphyra kurogii
Porphyra occidentalis
Porphyra ochotensis
Porphyra onoi
Porphyra pseudocrassa
Porphyra pseudolinearis
Porphyra variegata
PALMARIALES
Palmariaceae
Halosaccion firmum
Halosaccion yendoi
Palmaria palmata
Rhodophysemataceae
Rhodophysema georgii
CORALLINALES
Corallinaceae
Bossiella cretacea
Clathromorphum reclinatum
Corallina pilulifera
GIGARTINALES
Dumontiaceae
Dumontia contorta
Masudaphycus irregulare
Neodilsea crispata
Neodilsea yendoana
Endocladiaceae
Gloiopeltis furcata
Gigartinaceae
Chondrus ocellatus
Chondrus pinnulatus
Chondrus yendoi
Mazzaella japonica
Halymeniaceae
Grateloupia divaricata
Kallymeniaceae
Callophyllis cristata
Callophyllis rhynchocarpa
Petrocelidaceae
Mastocarpus pacificus
Rhodophyllidaceae
Rhodophyllis capillaris
Tichocarpaceae
Tichocarpus crinitus
GRACILARIALES
Gracilariaceae
Gracilaria vermiculophylla
RHODYMENIALESChampiaceae
Lomentaria hakodatensis
Rhodymeniaceae
Sparlingia pertusa

25

**** Data Book Categories of Japanese Fisheries Agency, details are given on the last page
FA DB****

Scientific name
CERAMIALES
Ceramiaceae
Antithamnion densum
Antithamnion crassa
Antithamnion hypnaeoides
Ceramium cimbricum
Ceramium kondoi
Ptilota filicina
Ptilota phacelocarpoides
Tokidaea corticata
Delesseriaceae
Acrosorium yendoi
Branchioglossum nanum
Congregatocarpus pacificus
Hideophyllum yezoense
Neoholmesia japonica
Neohypophyllum middendorfii
Tokidadendron kurilensis
Rhodomelaceae
Chondria dasyphylla
Enelittosiphonia stimpsonii
Janczewskia morimotoi
Laurencia nipponica
Neorhodomela aculeata
Neorhodomela munita
Neosiphonia japonica
Odonthalia annae
Odonthalia corymbifera
Polysiphonia morrowii
Polysiphonia senticulosa
Pterosiphonia bipinnata
Rhodomela teres
Symphyocladia latiuscula
Symphyocladia marchantioides

References

Kurogi, M., Yamada, I. and Masuda, M. (1985) Marine flora and its vegetation of Rausu
Coastal Area on the East Side of Shiretoko Peninsula. In: Comprehensive Report on
the Kelp in Rausu Sea Area, pp.1-82. Rausu Fishermen's Cooperative, Rausu town,
Menashi-gun, Hokkaido (in Japanese).
Yamada, Y. and Tanaka, T. (1944) Report on Seaweeds of Kitami Coastal Area ofShiretoko Peninsula. Hokusuishi Monthly Report 1 (3): 165-171 (in Japanese).

26Cold Current/Warm Current Species of Seaweed
Based on Kurogi et al. (1985)

Warm Current Species

Cold Current Species

Scientific name
CHLOROPHYCEAEULVALES
Capsosiphonaceae
Capsosiphon groenlandicus
Monostromataceae
Monostroma crassidermum
Ulvaceae
Ulvaria fusca
CLADOPHORALES
Cladophoraceae
Rhizoclonium tortuosum
ACROSIPHONIALES
Acrosiphoniaceae
Spongomorpha heterocladia
Spongomorpha duriuscula var. cartilaginea
Urospora wormskioldii
PHAEOPHYCEAERALFSIALES
Ralfsiaceae
Analipus filiformis
CHORDARIALES
Chordariaceae
Chordaria flagelliformis
Saundersella simplex
DICTYOSIPHONALES
Coilodesmaceae
Akkesiphycus lubricum
Coilodesme cystoseirae
LAMINARIALES
Alariaceae
Alaria angusta
Laminariaceae
Cymathaere japonica
Laminaria diabolica
Laminaria sachalinensis
FUCALES
Fucaceae
Fucus distichus subsp. evanescens
Sargassaceae
Cystoseira crassipes
RHODOPHYCEAEBANGIALES
Bangiaceae
Porphyra kurogii
Porphyra occidentalis
PALMARIALES
Palmariaceae
Halosaccion firmum
Halosaccion yendoi
GIGARTINALES
Dumontiaceae
Masudaphycus irregulare
Kallymeniaceae
Callophyllis cristata
Callophyllis rhynchocarpa
Rhodophyllidaceae
Rhodophyllis capillaris
CERAMIALES
Rhodomelaceae
Odonthalia annae
Odonthalia corymbifera
Pterosiphonia bipinnata

Scientific name
CHLOROPHYCEAE
CLADOPHORALES
Cladophoraceae
Chaetomorpha aerea
BRYOPSIDALES
Derbesiaceae
Derbesia marina
PHAEOPHYCEAEDICTYOTALES
Dictyotaceae
Dictyopteris divaricata
Dictyota dichotoma
CHORDARIALES
Acrotrichaceae
Acrothrix pacifica
Chordariaceae
Sphaerotrichia divaricata
SCYTOSIPHONALES
Scytosiphonaceae
Colpomenia bullosa
Petalonia binghamiae
LAMINARIALES
Laminariaceae
Agarum clathratum
Laminaria ochotensis
FUCALES
Sargassaceae
Cystoseira hakodatensis
Sargassum confusum
Sargassum miyabei
Sargassum thunbergii
RHODOPHYCEAEBANGIALES
Bangiaceae
Porphyra onoi
GIGARTINALES
Halymeniaceae
Grateloupia divaricata
RHODYMENIALESChampiaceae
Lomentaria hakodatensis
CERAMIALES
Ceramiaceae
Antithamnion crassa
Antithamnion hypnaeoides
Ceramium kondoi
Delesseriaceae
Acrosorium yendoi
Branchioglossum nanum
Hideophyllum yezoense
Neoholmesia japonica
Rhodomelaceae
Chondria dasyphylla
Enelittosiphonia stimpsonii
Laurencia nipponica
Neorhodomela munita
Polysiphonia morrowii
Polysiphonia senticulosa
Symphyocladia latiuscula
Symphyocladia marchantioides

References
Kurogi, M., Yamada, I. and Masuda, M. (1985) Marine flora and its vegetation of Rausu Coastal Area on the
East Side of Shiretoko Peninsula. In: Comprehensive Report on the Kelp in Rausu Sea Area, pp.1-82.
Rausu Fishermen's Cooperative, Rausu town, Menashi-gun, Hokkaido (in Japanese)

27

The Mammals of Shiretoko Peninsula and its Coastal
Area
Terrestrial mammals are from Shiretoko Museum (2000), Marine mammals are from Ito (1970), Kawamura(1981), Ohtaishi & Saito (1981), Ohtaishi and Nakagawa (1988), Sato (1998), Ishinazaka and Uni (2000),
and revised with comments by Ohtaishi (September 17, 2003).
Endemic based on Abe et al . (1994)** Red List categories of IUCN, details are given on the last page*** Red List Categories of Japanese Ministry of the Environment, details are given on the last page**** Data Book Categories of Japanese Fisheries Agency, details are given on the last page

Scientific name
INSECTIVORA
Soricidae
Sorex minutissimus hawkeri
Sorex gracillimus
Sorex caecutiens saevus
Sorex unguiculatus
CHIROPTERA
Vespertilionidae
Myotis mystacinus
Myotis ikonnikovi
Myotis macrodactylus
Myotis daubentonii
Myotis frater
Eptesicus nilssonii
Plecotus auritus
Barbastella leucomelas
Murina ussuriensis
LAGOMORPHA
Leporidae
Lepus timidus ainu
RODENTIA
Sciuridae
Sciurus vulgaris orientis
Tamias sibiricus lineatus
Pteromys volans orii
Muridae
Clethrionomys rutilus mikado
Clethrionomys rufocanus bedfordiae
Apodemus argenteus
Apodemus peninsulae giliacus
Apodemus speciosus
Mus musculus
Rattus rattus
Rattus norvegicus
CARNIVORA
Canidae
Vulpes vulpes schrencki
Nyctereutes procyonoides albus
Ursidae
Ursus arctos
Procyonidae
Procyon lotor
Mustelidae
Mustela erminea orientalis
Mustela nivalis nivalis
Mustela itatsi
Mustela vison
Martes zibellina brachura
Otariidae
Callorhinus ursinus
Eumetopias jubatus

IUCN RL**MoE RL

***

FA DB

****

Endemic to
Japan

VU

EN

LR/nt
EN

VU
EN
VU
VU
EN
VU
VU
VU

NT

●
●

NT

VU
EN

28

VU

D
R

** Red List categories of IUCN, details are given on the last page*** Red List Categories of Japanese Ministry of the Environment, details are given on the last page**** Data Book Categories of Japanese Fisheries Agency, details are given on the last page

Scientific name
Phocidae
Phoca largha
Phoca vitulina
Phoca hispida
Phoca fasciata
Erignathus barbatus
ARTIODACTYLA
Cervidae
Cervus nippon yesoensis
CETACEA
Eschrichtiidae
Eschrichtius robustus
Balaenopteridae
Balaenoptera acutorostrata
Megaptera novaeangliae
Balaenoptera borealis
Ziphiiadae
Ziphius cavirostris
Berardius bairdii
Mesoplodon stejnegeri
Mesoplodon carlhubbsi
Mesoplodon ginkgodens
Delphinidae
Lagenorhynchus obliquidens
Orcinus orca
Stenella coeruleoalba
Lissodelphis borealis
Tursiops truncatus
Delphinus delphis
Pseudorca crassidens
Globicephala melas
Globicephala macrorhynchus
Phocoenidae
Phocoena phocoena
Phocoenoides dalli
Physeteridae
Physeter macrocephalus

IUCN RL**MoE RL

EN

***

FA DB

EN

LR/nt
VU
EN

TD
R
D

LR/cd
LR/cd

Endemic to
Japan

TD
VU
DT
TD
D

CR

LR/cd

****

R
D
R
R
R
TD
R
D
TD
TD
DT

LR/cd

D

VU
LR/cd

R
TD

VU

TD

References

Abe, H., Ishii, N., Kaneko, K., Maeda, K., Miura, S. and Yoneda, M. (1994) A Pictorial Guide to theMammals of Japan. Tokai University Press, Tokyo (in Japanese).
Ishinazaka, T. and Uni, Y. (2000) Marine Mammals. In: Mammals of Shiretoko I. (Edited by
Shiretoko Museum), pp.162-225. Shari town and Shari town Board of Education (in Japanese).
Itoo, T. (1970) Survey of the Pinnipeds in Hokkaido. Mammalian Science19: 67-68(in Japanese).
Kawamura, A. (1981) Distribution of Whales on Okhotsk Sea Coast of Hokkaido. In: Report of theSurvey on Vertebrate Communities in Shiretoko Peninsula, Hokkaido, Japan, pp. 182-188. Natural
Conservation Division, Department of Lifestyle and Environment, Hokkaido (in Japanese).
Ohtaishi, N. and Saito, T. (1981) Pinnipedia of Shiretoko Coast and Migration of Steller's sea-lionon the Coast of Hokkaido. In: Report of the Survey on Vertebrate Communities in Shiretoko
Peninsula, Hokkaido, Japan, pp.165-181. Natural Conservation Division, Department of Lifestyleand Environment, Hokkaido (in Japanese).
Ohtaishi, N. and Nakagawa, H. (1988) Animals of Shiretoko. Hokkaido University Press, Sapporo(in Japanese with English summary).
Sato, H. (1998) A Migratory Route for Whales and Dolphins Supported by a Rich Environment.
SCIaS 49: 78-81 (in Japanese).
Shiretoko Museum (2000) Mammals of Shiretoko I. Shari town and Shari town Board of Education(in Japanese).

29

The Birds of Shiretoko Peninsula and its Coastal
From Nagata (1956), Japanese Society for Preservation of Birds (2001) and Shiretoko Museum (unpublished)
Endemic birds based on Wild Birds Society of Japan (1982)** Red List categories of IUCN, details are given on the last page*** Red List Categories of Japanese Ministry of the Environment, details are given on the last page**** Data Book Categories of Japanese Fisheries Agency, details are given on the last page
Scientific name

IUCN RL **GAVIIFORMES
GaviidaeGavia stellata
Gavia arctica
Gavia pacifica
Gavia adamsii
PODICIPEDIFORMES
Podicipedidae
Tachybaptus ruficollis
Podiceps nigricollis
Podiceps auritus
Podiceps grisegena
Podiceps cristatus
PROCELLARIIFORMES
Procellariidae
Fulmarus glacialis
Puffinus carneipes
Puffinus griseus
Puffinus tenuirostris
Hydrobatidae
Oceanodroma furcata
Oceanodroma leucorhoa
PELECANIFORMESPhalacrocoracidae
Phalacrocorax capillatus
Phalacrocorax pelagicus
Phalacrocorax urile
CICONIIFORMES
Ardeidae
Gorsachius goisagi
Ardeola bacchus
Bubulcus ibis
Egretta intermedia
Egretta garzetta
Ardea cinerea
ANSERIFORMES
AnatidaeBranta bernicla
Anser albifrons
Anser fabalis
Cygnus olor
Cygnus cygnus
Cygnus columbianus
Aix galericulata
Anas platyrhynchos
Anas poecilorhyncha
Anas crecca
Anas formosa
Anas falcata
Anas strepera
Anas penelope
Anas acuta
Anas clypeata
Aythya ferina

EN

VU

30

MoE RL ***FA DB ****

CR

EN

NT

NT

R

VU
NT
VU

R

VU

Endemic toJapan

** Red List categories of IUCN, details are given on the last page*** Red List Categories of Japanese Ministry of the Environment, details are given on the last page**** Data Book Categories of Japanese Fisheries Agency, details are given on the last page
Scientific name

IUCN RL **Aythya collaris
Aythya fuligula
Aythya marila
Melanitta nigra
Melanitta fusca
Histrionicus histrionicus
Clangula hyemalis
Bucephala clangula
Mergus albellus
Mergus serrator
Mergus merganser
FALCONIFORMES
Accipitridae
Pandion haliaetus
Milvus migrans
Haliaeetus albicillaHaliaeetus pelagicus
Accipiter gentilis
Accipiter gularis
Accipiter nisus
Buteo lagopus
Buteo buteo
Spizaetus nipalensis
Aquila clanga
Aquila chrysaetos
Circus cyaneus
Circus spilonotus
Falconidae
Falco rusticolus
Falco peregrinus
Falco subbuteo
Falco columbarius
Falco tinnunculus
GALLIFORMES
Tetraonidae
Tetrastes bonasia
Phasianidae
Coturnix japonica
Phasianus colchicus
GRUIFORMES
Gruidae
Grus japonensis
RallidaeRallus aquaticus
Porzana pusilla
Gallinula chloropus
Gallicrex cinerea
Fulica atra
CHARADRIIFORMES
Charadriidae
Charadrius dubius
Charadrius placidus
Charadrius alexandrinus
Charadrius mongolus
Pluvialis fulva
Pluvialis squatarola
Scolopacidae
Arenaria interpres

MoE RL ***

NT
LR/nt
VU

EN
VU
VU
NT

EN
EN
VU

VU

EN

31

VU

FA DB ****

Endemic toJapan

** Red List categories of IUCN, details are given on the last page*** Red List Categories of Japanese Ministry of the Environment, details are given on the last page**** Data Book Categories of Japanese Fisheries Agency, details are given on the last page
Scientific name

IUCN RL **Calidris ruficollis
Calidris subminuta
Calidris alpina
Calidris tenuirostris
Crocethia alba
Limicola falcinellus
Tringa erythropus
Tringa nebularia
Tringa glareola
Heteroscelus brevipes
Actitis hypoleucos
Numenius phaeopus
Scolopax rusticola
Gallinago gallinago
Gallinago stenura
Gallinago megala
Gallinago hardwickii
Gallinago solitaria
Recurvirostridae
Himantopus himantopus
Phalaropodidae
Phalaropus fulicarius
Phalaropus lobatus
Stercorariidae
Stercorarius pomarinus
Stercorarius parasiticus
Stercorarius longicaudus
Laridae
Larus ridibundus
Larus heuglini
Larus argentatus
Larus schistisagus
Larus glaucescens
Larus hyperboreus
Larus glaucoides
Larus canus
Larus crassirostris
Rissa tridactyla
Rhodostethia rosea
Pagophila eburnea
Chlidonias leucopterus
Sterna hirundo
Alcidae
Uria aalge
Uria lomvia
Cepphus columba
Cepphus carbo
Brachyramphus marmoratus
Synthliboramphus antiquus
Aethia cristatella
Aethia pusilla
Aethia psittacula
Cerorhinca monocerata
Fratercula corniculata
Lunda cirrhata
COLUMBIFORMESColumbidae
Streptopelia orientalis

MoE RL ***FA DB ****

NT

32

EN

R

CR

EN

VU
CR

EN
R
EN

CR

EN

Endemic toJapan

** Red List categories of IUCN, details are given on the last page*** Red List Categories of Japanese Ministry of the Environment, details are given on the last page**** Data Book Categories of Japanese Fisheries Agency, details are given on the last page
Scientific name

IUCN RL **MoE RL ***

EN

CR
CR

Sphenurus sieboldii
CUCULIFORMES
Cuculidae
Cuculus fugax
Cuculus canorus
Cuculus saturatus
Cuculus poliocephalus
STRIGIFORMES
Strigidae
Nyctea scandiaca
Bubo bubo
Ketupa blakistoni
Asio otus
Asio flammeus
Otus scops
Otus lempiji
Aegolius funereus
Ninox scutulata
Strix uralensis
CAPRIMULGIFORMESCaprimulgidae
Caprimulgus indicus
APODIFORMES
Apodidae
Hirundapus caudacutus
Apus pacificus
CORACIIFORMES
Alcedinidae
Ceryle lugubris
Halcyon coromanda
Alcedo atthis
UpupidaeUpupa epops
PICIFORMES
PicidaeJynx torquilla
Picus canus
Dryocopus martius
Dendrocopos major
Dendrocopos leucotos
Dendrocopos minor
Dendrocopos kizuki
PASSERIFORMES
Alaudidae
Alauda arvensis
Hirundinidae
Riparia riparia
Hirundo rustica
Hirundo daurica
Delichon urbica
Motacillidae
Motacilla flava
Motacilla cinerea
Motacilla alba
Motacilla grandis
Anthus hodgsoni
Anthus spinoletta

FA DB ****

Endemic to
Japan

EN

VU

●

33

** Red List categories of IUCN, details are given on the last page*** Red List Categories of Japanese Ministry of the Environment, details are given on the last page**** Data Book Categories of Japanese Fisheries Agency, details are given on the last page
Scientific name

IUCN RL **Pycnonotidae
Hypsipetes amaurotis
LaniidaeLanius bucephalus
Lanius cristatus
Lanius excubitor
Bombycillidae
Bombycilla garrulus
Bombycilla japonica
Cinclidae
Cinclus pallasii
Troglodytidae
Troglodytes troglodytes
Prunellidae
Prunella collaris
Prunella rubida
TurdidaeErithacus akahige
Luscinia calliope
Luscinia cyane
Tarsiger cyanurus
Phoenicurus ochruros
Phoenicurus auroreus
Saxicola torquata
Monticola solitarius
Zoothera dauma
Turdus sibiricus
Turdus cardis
Turdus chrysolaus
Turdus pallidus
Turdus obscurus
Turdus naumanni
Turdus pilaris
Sylviidae
Urosphena squameiceps
Cettia diphone
Locustella fasciolata
Locustella ochotensis
Locustella lanceolata
Acrocephalus bistrigiceps
Acrocephalus arundinaceus
Phylloscopus borealis
Phylloscopus borealoides
Phylloscopus coronatus
Regulus regulus
Muscicapidae
Ficedula narcissina
Cyanoptila cyanomelana
Muscicapa sibirica
Muscicapa griseisticta
Muscicapa dauurica
Aegithalidae
Aegithalos caudatus
Paridae
Parus palustris
Parus montanus
Parus ater
Parus varius

MoE RL ***FA DB ****

Endemic to
Japan

NT

LR/nt

●

34

** Red List categories of IUCN, details are given on the last page*** Red List Categories of Japanese Ministry of the Environment, details are given on the last page**** Data Book Categories of Japanese Fisheries Agency, details are given on the last page
Scientific name

IUCN RL **MoE RL ***

VU

NT
NT

Parus major
Sittidae
Sitta europaea
Certhiidae
Certhia familiaris
Zosteropidae
Zosterops japonicus
Emberizidae
Emberiza leucocephalos
Emberiza cioides
Emberiza fucata
Emberiza rustica
Emberiza elegans
Emberiza aureola
Emberiza sulphurata
Emberiza spodocephala
Emberiza variabilis
Emberiza schoeniclus
Calcarius lapponicus
Plectrophenax nivalis
Fringillidae
Fringilla montifringilla
Carduelis sinica
Carduelis spinus
Carduelis flammea
Leucosticte arctoa
Carpodacus roseus
Pinicola enucleator
Loxia curvirostra
Loxia leucoptera
Uragus sibiricus
Pyrrhula pyrrhula
Eophona personata
Coccothraustes coccothraustes
Ploceidae
Passer rutilans
Passer montanus
Sturnidae
Sturnus philippensis
Sturnus cineraceus
Corvidae
Garrulus glandarius
Pica pica
Nucifraga caryocatactes
Corvus corone
Corvus macrorhynchos
Corvus corax

References

FA DB ****

Endemic to
Japan

Japanese Society for Preservation of Birds-Kushiro chapter (2001) 2000 Report on the Shiretoko National
Wildlife Protection Area. Nature Conservation Bureau, Ministry of the Environment (in Japanese).
Nagata, Y. (1956) Shiretoko Peninsula (1). Wild Birds 21: 220-225 (in Japanese).
Nagata, Y. (1956) Shiretoko Peninsula (2). Wild Birds 21: 337-342 (in Japanese).
Nagata, Y. (1956) Shiretoko Peninsula (3). Wild Birds 21: 388-393 (in Japanese).
Wild Bird Society of Japan (1982) A Field Guide to the Birds of Japan. Wild Bird Society of Japan, Tokyo.

35

The Reptiles of Shiretoko Peninsula and its Coastal
Area
From the records of Shiretoko Museum, August 2003
Endemic based on Uchiyama et al . (2002)**** Data Book Categories of Japanese Fisheries Agency, details are given on the last page
FA DB ****

Scientific name
SQUAMATALACERTILIA
Lacertidae
Takydromus tachydromoides
Scincidae
Eumeces latiscutatus
SERPENTES
Colubridae
Elaphe climacophora
Elaphe quadrivirgata
Elaphe conspicillata
Viperidae
Agkistrodon blomhoffii
TESTUDINATA
Dermochelyidae
Dermochelys coriacea

Endemic to
Japan

●

●
●
●
●

EN

The Amphibians of Shiretoko Peninsula
From the records of Shiretoko Museum, August 2003
Endemic based on Uchiyama et al. (2002)
Endemic toJapan

Scientific name
CAUDATA
Hynobiidae
Hynobius retardatus
SALIENTIA
RanidaeRana pirica
Hylidae
Hyla japonica●●

References

Uchiyama, R., Maeda, N., Numata, K. and Seki, S. (2002) A Photographic Guide;
Amphibians and Reptiles in Japan. HEIBONSHA Ltd., Publishers, Tokyo (in Japanese).

36

The Fishes of Shiretoko Peninsula and its
Coastal Area

From Shiretoko Museum (2003)** Red List categories of IUCN, details are given on the last page*** Red List Categories of Japanese Ministry of the Environment, details are given on the lastpage**** Data Book Categories of Japanese Fisheries Agency, details are given on the last page
IUCN RL**

Scientific name
PETROMYZONTIFORMESPetromyzontidae
Lethenteron japonicum
Lethenteron kessleri
CHIMAERIFORMES
Chimaeridae
Hydrolagus barbouri
CARCHARHINIFORMES
Carcharhinidae
Prionace glauca
LAMNIFORMES
LamnidaeLamna ditropis
SQUALIFORMES
Dalatiidae
Somniosus pacificus
Squalidae
Squalus acanthias
RAJIFORMES
RajidaeRaja pulchra
Bathyraja diplotaenia
Bathyraja violacea
Bathyraja matsubarai
Bathyraja minispinosa
Bathyraja aleutica
Bathyraja smirnovi
Bathyraja bergi
Bathyraja lindbergi
Rhinoraja kujiensis
Dasyatidae
Dasyatis matsubarai
ACIPENSERIFORMES
Acipenseridae
Huso dauricus
Acipenser medirostris
ALBULIFORMES
Notacanthidae
Notacanthus chemnitzi
ANGUILLIFORMES
Synaphobranchidae
Synaphobranchus kaupii
Congridae
Conger sp.
CLUPEIFORMESEngraulidae
Engraulis japonicus
Clupeidae
Clupea pallasii
Etrumeus teres
Sardinops melanostictus

MoE RL ***FA DB****

NT

R

TD

DT

EN
EN

EN

D

37

** Red List categories of IUCN, details are given on the last page*** Red List Categories of Japanese Ministry of the Environment, details are given on the lastpage**** Data Book Categories of Japanese Fisheries Agency, details are given on the last page
IUCN RL**

Scientific name
CYPRINIFORMES
Cyprinidae
Tribolodon hakonensis
Tribolodon brandti
Tribolodon ezoe
Phoxinus percnurus sachalinensis
Carassius auratus langsdorfii
Cyprinus carpio
Cobitidae
Noemacheilus barbatulus toni
Misgurnus anguillicaudatus
OSMERIFORMES
Osmeridae
Osmerus eperlanus mordax
Hypomesus japonicus
Hypomesus nipponensis
Salangidae
Salangichthys microdon
SALMONIFORMES
Salmonidae
Hucho perryi
Salvelinus leucomaenis
Salvelinus malma
Oncorhynchus keta
Oncorhynchus tschawytscha
Oncorhynchus gorbuscha
Oncorhynchus nerka
Oncorhynchus masou masou
Oncorhynchus kisutch
Oncorhynchus mykiss
Salmo trutta
Salmo salar
STOMIIFORMES
Stomiidae
Chauliodus macouni
AULOPIFORMES
Notosudidae
Scopelosaurus harryi
Paralepididae
Paralepis atlantica
Anotopteridae
Anotopterus nikparini
Alepisauridae
Alepisaurus ferox
MYCTOPHIFORMESMyctophidae
Nannobrachium regale
Lampanyctus jordani
LAMPRIDIFORMES
Trachipteridae
Trachipterus ishikawae
GADIFORMES
Macrouridae
Coryphaenoides pectoralis
Coryphaenoides acrolepis
Coryphaenoides cinereus
MoridaePhysiculus maximowiczi
Antimora microlepis

MoE RL ***FA DB****

NT

EN

VU

NT

D

D

DT

38

** Red List categories of IUCN, details are given on the last page*** Red List Categories of Japanese Ministry of the Environment, details are given on the lastpage**** Data Book Categories of Japanese Fisheries Agency, details are given on the last page
Scientific name
Laemonema longipes
Gadidae
Theragra chalcogramma
Gadus macrocephalus
Eleginus gracilis
LOPHIIFORMES
Lophiidae
Lophius litulon
MUGILIFORMES
Mugilidae
Mugil cephalus cephalus
Chelon haematocheilus
BELONIFORMES
Scomberesocidae
Cololabis saira
Hemiramphidae
Hyporhamphus sajori
CYPRINODONTIFORMES
Poeciliidae
Poecilia reticulata
STEPHANOBERYCIFORMES
Melamphaidae
Poromitra crassiceps
Cetomimidae
Cetomimus compunctus
ZEIFORMES
ZeidaeZeus faber
Zenopsis nebulosa
Oreosomatidae
Allocyttus verrucosus
GASTEROSTEIFORMES
Hypoptychidae
Hypoptychus dybowskii
Gasterosteidae
Gasterosteus aculeatus (Sea of Japan form)
Gasterosteus aculeatus (Pacific Ocean form)
Pungitius tymensis
Pungitius pungitius (freshwater type)
Pungitius pungitius (brackish-water type)
Syngnathidae
Syngnathus schlegeli
SCORPAENIFORMES
Scorpaenidae
Sebastes glaucus
Sebastes alutus
Sebastes flammeus
Sebastes borealis
Sebastes steindachneri
Sebastes taczanowskii
Sebastes wakiyai
Sebastes minor
Sebastes schlegelii
Sebastes vulpes
Sebastes trivittatus
Sebastolobus macrochir
Triglidae
Lepidotrigla kishinouyei

39

IUCN RL**MoE RL ***FA DB****

TD

NT

D

D
DT

** Red List categories of IUCN, details are given on the last page*** Red List Categories of Japanese Ministry of the Environment, details are given on the lastpage**** Data Book Categories of Japanese Fisheries Agency, details are given on the last page
IUCN RL**

Scientific nameAnoplopomatidae
Erilepis zonifer
Anoplopoma fimbria
Hexagrammidae
Hexagrammos stelleri
Hexagrammos otakii
Hexagrammos octogrammus
Hexagrammos lagocephalus
Pleurogrammus azonus
Pleurogrammus monopterygius
Cottidae
Hemilepidotus gilberti
Icelus cataphractus
Icelus canaliculatus
Icelus mandibularis
Triglops jordani
Triglops scepticus
Cottus hangiongensis
Cottus amblystomopsis
Enophrys diceraus
Taurocottus bergi
Myoxocephalus stelleri
Myoxocephalus brandti
Myoxocephalus polyacanthocephalus
Myoxocephalus jaok
Porocottus allisi
Porocottus tentaculatus
Argyrocottus zanderi
Cottiusculus schmidti
Cottiusculus gonez
Bero elegans
Alcichthys elongatus
Gymnocanthus herzensteini
Gymnocanthus intermedius
Gymnocanthus detrisus
Cottidae gen. sp.
Hemitripteridae
Hemitripterus villosus
Nautichthys pribilovius
Blepsias cirrhosus
Blepsias bilobus
Agonidae
Percis japonicus
Hypsagonus corniger
Hypsagonus proboscidalis
Hypsagonus jordani
Tilesina gibbosa
Occella dodecaedron
Occella iburia
Brachyopsis rostratua
Pallasina barbata
Podothecus sachi
Podothecus sturioides
Podothecus hamlini
Freemanichthys thompsoni
Bothragonus occidentalis
Psychrolutidae
Malacocottus zonurus
Dasycottus setiger

40

MoE RL ***FA DB****

** Red List categories of IUCN, details are given on the last page*** Red List Categories of Japanese Ministry of the Environment, details are given on the lastpage**** Data Book Categories of Japanese Fisheries Agency, details are given on the last page
IUCN RL**

Scientific nameEurymen gyrinus
Cyclopteridae
Eumicrotremus taranetzi
Eumicrotremus pacificus
Aptocyclus ventricosus
Liparidae
Liparis miostomus
Liparis ochotensis
Liparis agassizii
Liparis latifrons
Crystallichthys matsushimae
Squaloliparis dentatus
Careproctus macrodiscus
Careproctus furcellus
Careproctus cypselurus
Careproctus cyclocephalus
Careproctus rastrinus
Careproctus colletti
Careproctus roseofuscus
Careproctus mederi
Careproctus parvidiscus
Careproctus sp.
Paraliparis grandis
PERCIFORMES
Priacanthidae
Cookeolus japonicus
Pomatomidae
Scombrops boops
Echeneidae
Echeneis naucrates
Coryphaenidae
Coryphaena hippurus
Carangidae
Trachurus japonicus
Seriola quinqueradiata
Seriola lalandi
Alectis ciliaris
Naucrates ductor
Emmelichthyidae
Erythrocles schlegelii
Lobotidae
Lobotes surinamensis
Oplegnathidae
Oplegnathus fasciatus
Oplegnathus punctatus
Cichlidae
Oreochromis niloticus
Embiotocidae
Ditrema temmincki
Bathymasteridae
Bathymaster derjugini
Zoarcidae
Neozoarces steindachneri
Zoarces elongatus
Petroschmidtia albonotata
Lycenchelys albomaculata
Lycodes soldatovi
Bothrocara molle
Bothrocara tanakae

41

MoE RL ***FA DB****

** Red List categories of IUCN, details are given on the last page*** Red List Categories of Japanese Ministry of the Environment, details are given on the lastpage**** Data Book Categories of Japanese Fisheries Agency, details are given on the last page
IUCN RL**

Scientific name
Bothrocarina microcephala
Stichaeidae
Chirolophis japonicus
Soldatovia polyactocephala
Opisthocentrus ocellatus
Pholidapus dybowskii
Lumpenopsis pavlenkoi
Ernogrammus hexagrammus
Stichaeopsis nana
Stichaeus grigorjewi
Stichaeus nozawai
Stichaeus fuscus
Lumpenella longirostris
Acantholumpenus mackayi
Lumpenus sagitta
Leptostichaeus pumilus
Pholidae
Rhodymenichthys dolichogasterPholis picta
Pholis fasciata
Pholis crassispina
Anarhichadidae
Anarhichas orientalis
Zaproridae
Zaprora silenus
Trichodontidae
Arctoscopus japonicus
Ammodytidae
Ammodytes personatus
GobiidaeGymnogobius urotaenia
Gymnogobius petshiliensis
Acanthogobius lactipes
Luciogobius guttatus
Gymnogobius castaneus
Trichiuridae
Trichiurus japonicus
Scombridae
Scomber japonicus
Auxis rochei
Thunnus thynnus
Katsuwonus pelamis
Xiphiidae
Istiophorus platypterus
Centrolophidae
Hyperoglyphe japonica
Psenopsis anomala
PLEURONECTIFORMES
Pleuronectidae
Atheresthes evermanni
Reinhardtius hippoglossoides
Hippoglossus stenolepis
Acanthopsetta nadeshnyi
Hippoglossoides pinetorum
Hippoglossoides dubius
Verasper moseri
Pleuronectes mochigarei
Pleuronectes punctatissimus
Pleuronectes asper

MoE RL ***FA DB****

DT

D
D
R

42

** Red List categories of IUCN, details are given on the last page*** Red List Categories of Japanese Ministry of the Environment, details are given on the lastpage**** Data Book Categories of Japanese Fisheries Agency, details are given on the last page
IUCN RL**

Scientific name
Pleuronectes herzensteinii
Pleuronectes schrenki
Pleuronectes obscurus
Pleuronectes quadrituberculatus
Platichthys stellatus
Kareius bicoloratus
Clidoderma asperrimum
Glyptocephalus stelleri
TETRAODONTIFORMES
Monacanthidae
Thamnaconus modestus
Tetraodontidae
Takifugu porphyreus
Takifugu xanthopterus
Diodontidae
Diodon holocanthus
Molidae
Mola mola

References

MoE RL ***FA DB****

Shiretoko Museum (2003) Fishes of Shiretoko. Shari town and Shari-town Board of
Education (in Japanese).

43

The Categories**Categories of IUCN Red List
EX

EXTINCT: A taxon is Extinct when there is no reasonable doubt that the last individual has died.
A taxon is presumed Extinct when exhaustive surveys in known and/or expected habitat, atappropriate times (diurnal, seasonal, annual), throughout its historic range have failed to record
an individual. Surveys should be over a time frame appropriate to the taxon's life cycle and life
form.

EW

EXTINCT IN THE WILD: A taxon is Extinct in the Wild when it is known only to survive incultivation, in captivity or as a naturalized population (or populations) well outside the past range.
A taxon is presumed Extinct in the Wild when exhaustive surveys in known and/or expected
habitat, at appropriate times (diurnal, seasonal, annual), throughout its historic range have failed
to record an individual. Surveys should be over a time frame appropriate to the taxon's life cycle
and life form.

CR

CRITICALLY ENDANGERED: A taxon is Critically Endangered when the best available evidence
indicates that it meets any of the criteria A to E for Critically Endangered, and it is therefore
considered to be facing an extremely high risk of extinction in the wild.

EN

ENDANGERED: A taxon is Endangered when the best available evidence indicates that it meets
any of the criteria A to E for Endangered, and it is therefore considered to be facing a very highrisk of extinction in the wild.

VU

VULNERABLE: A taxon is Vulnerable when the best available evidence indicates that it meets
any of the criteria A to E for Vulnerable, and it is therefore considered to be facing a high risk of
NEAR THREATENED: A taxon is Near Threatened when it has been evaluated against thecriteria but does not qualify for Critically Endangered, Endangered or Vulnerable now, but is close
to qualifying for or is likely to qualify for a threatened category in the near future.

NT

LR(1994 LOWER RISK: A taxon is Lower Risk when it has been evaluated, does not satisfy the criteria forcategory) any of the categories Critically Endangered, Endangered or Vulnerable. Taxa included in theLower Risk category can be separated into three subcategories:
LR/cd Conservation Dependent. Taxa which are the focus of a continuing taxon-specific or habitatspecific conservation programme targeted towards the taxon in question, the cessation of which
would result in the taxon qualifying for one of the threatened categories above within a period offive years.
LR/nt Near Threatened. Taxa which do not qualify for Conservation Dependent, but which are close toqualifying for Vulnerable.
LR/lc Least Concern. Taxa which do not qualify for Conservation Dependent or Near Threatened.

***Categories of Red Data Book published by Environment Agency of Japan
EX
EW
CR

Extinct: A taxon is considered extinct in Japan.
Extinct in the wild: A taxon known only to survive in cultivation or in captivity.

EN

Threatened IB: Endangered. A taxon is not Critically Endangered but is facing a very high risk ofextinction in the wild in the near future.

VU

Threatened II: Vulnerable. A taxon believed likely to move into the upper rank in near future if
the casual factors continue operating.
Near threatened: A taxon which are close to qualifying for vulnerable.

NT

Threatened IA: Critically Endangered. A taxon is facing an extremely high risk of extinction in thewild immediate future.

44

**** Date Book Categories of Japanese Fisheries Agency
EN
VU
R

Endengered: A taxon considered to be facing a very high risk of extinction.
Vulnerable: A taxon considered to be facing a high risk of extinction.
Rare: A taxon that is vulnerable in terms of its base for survival.
It is transferred to the next category depending on environmental conditions.
a. Low population density and scarceb. Very limited habitatc. Isolated distribution biogeographicallyd. Special environmental conditions required in all or part of the life cycle
Judging from the change in habitat, the survival of the species becomes threatened.
a. Decline in populationb. Worsening living conditions
c. Under threat of over-capture or collection pressured. Intrusion of crossable species or subspecies

D

Decreasing: A taxon that is evidently declining in population. It is difficult to recover to the
population in the past because of changes to living conditions, or of continuous collection pressure.

DT

Decreasing Tendency : A taxon considered to be on the decrease in population on a long-termbasis.
The long-term variation seen from the fishery statistics and other documents clearly backs thetrend of gradual decline.

TD

Tenporal Decreasing: A taxon considered to be within the range of natural fluctuations. Recovery
is possible.

References

Environment Agency (2000) Thereatened Wildlife of Japan - Red Date Book 2nd ed., Vascular Plants. 660pp.
Japan Wildlife Research Center, Tokyo.
Environment Agency (2000) Thereatened Wildlife of Japan - Red Date Book 2nd ed., Reptilia/Amphibia.
120pp. Japan Wildlife Research Center, Tokyo.
Environment Agency (2000) Thereatened Wildlife of Japan - Red Date Book 2nd ed., Bryophytes, Algae,
Lichens, Fungi. 429pp. Japan Wildlife Research Center, Tokyo.
Fisheries Agency (1998) Data book of rare aquatic animals and plants in Japan. 437pp. Japan Fisheries
Resource Conservation Association, Tokyo.
IUCN. (1994). IUCN Red List Categories and Criteria: Version 2.3. IUCN Species Survival Commission.
IUCN, Gland, Switzerland.
IUCN. (2001). IUCN Red List Categories and Criteria: Version 3.1. IUCN Species Survival Commission.
IUCN, Gland, Switzerland and Cambridge, UK.
Ministry of the Environment（2002）Threatened Wildlife of Japan - Red Data Book 2nd ed., Aves. 278pp,
Japan Wildlife Research Center, Tokyo.
Ministry of the Environment（2002）Threatened Wildlife of Japan - Red Data Book 2nd ed., Mammalia.
177pp, Japan Wildlife Research Center, Tokyo.
Ministry of the Environment（2003）Threatened Wildlife of Japan - Red Data Book 2nd ed., Pisces-Brackish
and Fresh Water Fishes. 230pp, Japan Wildlife Research Center, Tokyo.
The IUCN Red List of Threatened Species: http://www.iucn.org/themes/ssc/red-lists.htm

45