fJ -lt" ~ ± i6J ~ ~~%·!c.1JTransactions and Proceedingsof thePalaeontological Society of Japan
New SeriesNo. 29Palaeontological Society of Japan
April 10, 1958

CONTENTS
TRANSACTIONS
Page

330.

Liassic Volsella. JlJylilus and some other Dysodont Species in Japan ..... .
.......................................................................................... Itaru HAvAl'.n 155

331.

Mesozoic Plants from the Tetori Series, Central Honshu, Japan ........... .
.. . .. ... ... . ........ ... . .. . .. .. . . .. ... . . ... . . .. ... . . . .. . . .. ... . .. . . . .. . ... Tatsuaki KIMUHA 161i

332.

On the Occurrence of Cercis in Japan................................. Keiji SuzuKI 168

333.

A Short Note on Lonsdaleoides toriyamai MINATo .................................. ..
............................................................ 1\Jasao MINATo and i\Iakoto KATo 172

334.

On some Cephalaspid Opisthobranchia from the Japanese l\Iiocene....................................................................................... Junji horGAIVA 175

PROCEEDINGS ................................................................................................ 184

President:

H isakatsu Y Aim

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IIaruyoshi FuJIMOTo (Publication). Shoshiro HANzAwA. Kotara IIATAr, Ichiro
IIAYASAKA (General Affairs). Kazuo HuzioKA, Teiichi KoBAYAsm (Editor),
Jiro MAKIYAMA, Tatsuro MATsuMoTo, Masao MINATo, Tokio SmKAMA, Fuyuji
TAKAr (General Affairs), Ryuzo ToRIYAMA.
Secretary:

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Assistant Secretary:

Takeo lcmKAWA.

All Communications relating to this journal should be addressed to the
PALAEONTOLOGICAL SOCIETY OF JAPAN
Geological Institute, Faculty of Science, University of Tokyo, japan

Trans. Proc. l'alacont. Soc. Japan, N. S., No. 29. pp. 155-165, pis. 23, 2·1, April 10, 1958

330.

LIASSIC VOLSELLA, MYTILUS AND SOl\IE OTIIER
DYSODONT SPECIES IN .JAPAN*(Studies on the Liassic Pelecypods in japan, ti)

ITARU HAYAMI
Geological Institute. University of Tokyo:<f>:fllfl( 7 { 7 "· q"J Volsella. :llytilus -t-q"J(ll!<7.>ftN•r=:.:.t!{JU;ft: .;j~J.t,;;-.; ..t V:;.tiltJII/mgfi.:lifN· 7.>
Volsella. ;lrcomytifus. Jllytilus. "Ostrea ·•. PI£· ria. Oxyloma (?) 1.:"")\, '"C.J;lt:1·t L t.:t'i-'il! 5 irfrRZ.~
tr 12 ~~l:o:,:f'&8UttJLt..:, Mytilus (11.·-?JL\.@Z.?\.~. Volsella I'J·I?"'JnnJB·c, ;; ~ 7~l.:'rl'iiit:Jt;:
Fakimytilus !IE~.i.~l.:.tif~"t 7., o Arcomytilus (tflf:JIE Mytilus Xl:t Erachidontcs q"J.illiA~ t!ii!.(Mc tt ~L -c)J<:t..:il:. ~.\£Lt. J.r~t L"C.V..t'J "'~-c~.Q.

The l\lytilidac and Ostreiclae arc fair·ly common in the Liassic Kuruma
group. KouAYAsm (1935) reported theoccurrence of Mytilus and ~lrcomytilus
from the group, but they remain unde·scribed. Lately the writer added a large
amount of fossils to the Kuruma-Shizu·
kawa collection including some Vulsella
and Pteria (s. I.) in the Lower Hettangian
Geratrigouia llosourensis bed of the Shizu·
kawa group. In the present paper aredescribed the following species:-

1\/ytilus (1-ltldmytilus) lieranirus IIAYAMI,
subsp.
,\lytilus (Fakimytilus) sp. i ndct.
"Os/r('tl ·· a sp. i ndet.
.. Ostn·a•· b sp. indct.
Pteria (s.l.) kilakamiensis lL~.YAMI, newspecies.
Oxytoma (?) sp. i ndct.

Family l\Iytilidac FtEMIN(iThe classification of the family isprincipally based on the anatomy whichis. however. inapplicable to fossils. In
consequence l he classificatory criteria
arc inevitably restricted to the hinge·
st ructurc. I igament -structure, umbonal
septum. external outline, umbonal posi·tion and surface ornamentation.
The earliest of this family is Volsellain the Devonian. Since then. the genus
persisted through the long geologicalages without any striking modification.
It is most reasonable to regard thegenus to be the trunk of the Mytilidae.
whence Mytilus and many other relatedVolsella bakcrc/loides 11.-\.YAMl. new spe·
cics.
Volsella sp. indet.
:lrcomytilus dairensis KouAYASHI arttl HAYAMI, new species.
Arcomytilus sp. indet.
Mytilus (Falcimytilus) stricapillatus IIA Y AMI,
new species.
ftlytilus (Falcimytilus) stricapi/latus l-L\. Y AMI.
suhsp.
ilzl'l ifus (Fa lei mytilus) he rani rus HAYAM 1.
new species.

* Received April 5, 1957; read June 15, 1957.
155

156

ltaru HAYAMI

genera h:n·e branched off. Jl!yl ilus has
been produced from Folse/la by I heshifting of the umbo from subterminal
to terminal.
Living Jfylilus has some umbonal
teeth and weak anterior adductor, while
the hinge-area is edentulous in Volsella.
Nevertheless it is not always easy to
distinguish these two as for Jurassic
forms. because many forms are foundto be intermediate between Volsella andJHytilus with regard to outline andumbonal position. Therefore. the former is considered by some authorities asa subgenus of the latter.
Cox (1931) established Falcimy/ilus asa subgenus of !1h•tilus for some .Jurassic
species with terminal umbones andedentulous hinge areas. Ilc is of opinion that true A!ytilus is absent in the
pre-Jurassic. all hough there are manyso-called .llylilus species in the Triassic
and several in the Permian. But it ispossibly a little prior to Lias that Fahilll)'filus was derived at Jirst from Volsella, if considered that J1JyUus (Falcimytilus) nasai KoBAY Aslll and kmKAW A
and its varieties (1950; Icn1 KAw A. 19!i-l)
are known from the Carnic in the
Kochigalani series in Japan. Putting
aside this problem. Falcimytilus is really closer to Folse/la than Mytilus s. str.
in the edentulous hinge, although it
agrees better with Mytilus s. str. in theterminal umbo. As suggested by Cox(1940), the subgenus may include mostJurassic species of Mytilus, because no
Jurassic species has umbonal teeth. The
present five Liassic forms are probably
included in its category.
Arcomytilus AcAssJz (184:.!) is characterized by the modiol iform out! ine andradial markings on the surface. Cux(1937, 1940) considered it as a subgenus
of Braclzidoutes SwAINsoN (1840). In flraclzidontes s. str., however, the shell isovate and the anterior area more
expanded than in Arcom_rlilus. As mentioned by Cox. dysodont teeth arefrequently ubsC'rvahle in Recent Braclti·
donfes but absent in Arcomytilus, although J\<;AssJz mistook the marginal
impression of radial ribs for them. In
the Middle and Upper Jurassic therearc many radially ribbed species ofArcomytilus. Some of them have modioliform or even mytilifonn outlines andnearly terminal umbones. I~adial markings are commonly met with also in
Sept ifer, Musculus a ncl Creuella.
But,·lrcomylilus differs essentially from
Septifer in the absence of the umbonal
septum for adherence of the anterior
adductor, and from two others in thetrigonal outline.
Although certain phylogenetical relationship may exist between Arcomytilus
and Brachidolll!'s, the writer thinks
plausible to regard ilrcomytilus to be a
distinct genus derived from the main
trunk of the 1\Iytilidac comprising Volsella and *'ome related genera.
So far as the writer is aware, Arcomytilus is scarcely known from the Lias.
The two forms in this paper which have
explicit radial ornaments may be its old
representatives, but the materials are
too poor to give their descriptions indetail.
Genus Volsella ScoPoJ.I. 1777.
( =Mr1diolus LA MAReK, 179~;]IJodiofll L.utARCK, )80);
Emnodiolus humrNG, 1900)
TyjJe sPet·ies :-Mylilus modiolus LINNi·:,
Recent.
V olsclla ballc/Jeiloides 1-1 AY Al\11,
new species.
Plate 23, Figures 1-3.

Liassic Vulsella, J1!ytilus and some other Dysodonl Species in Japan

Desrriptiou :-Shell medium, equivalve.
modioliform, elongated postero-ventrally, non-carinated. strongly inflated,slightly to fairly longer than high;test thin; hinge-margin slightly convex.
fairly long, occupying about two-thirdsof shell-length: posterior margin gently
arcuate: anterovcntral one of shell-bodyalmost straight. but sinuated at its
junction with anterior margin in frontof de,·eloped anterior area; umbonal
angle hetwcen hinge-margin and anteroventral one ahoul 4~ degrees; anterior<trea wide, well inflated, clearly definedfrom shell-body by a shallow groove.
somewhat similar to anterior wing in

157

Ba!?evellia. rounded at the extremity:
anterior slope of shell-surface steep,nearly vertical; the greatl.'st convexity
lying close to antero-ventral margin,
and posterior area comparatively flattened; whole surface marked with moreor less regular coarse concentric Jines.
Internally. ligament probably subinternal. occupying greater part of hingeline. supported by an internal ridge
which is narrow and subparallel tohinge-line. running behind umbonal
area to postero-dorsal corner: umbonal
septum absent: hinge edentulous: adductors unknown.

LengthHeightThickness

Holol YJll' (1\IM ~719) Right vah·e

H.O

30.5

39.0
29. 0
39. 0

8. 0

I' a ra I y pl' (!\I :\1 ~7:!0) Ldl valve
l'aralypc (Ml\1 2721) Left internal mould

J\lr'tlSIIIPI/le/1{ ill 111111.

0/J.w'rNtfion andCnmj){tri~on :-The holotypc (Fig. I) ig a well preserved specimen, showing the complete outline ofright Yalve. In the paratypc (Fig. 3)
umbonal and dygodont teeth are absent.
Internally, the anterior area seems not
so well dctittecl as extl.'rnally. The smallparatype (Fig. 2) is fairly different inoutline from the holotype; its umbonal
angle much larger. Seeing the occurrence from the same f ossi I bed and thesimilar surface-markings, however. suchdifferences arc considered indi\•iduality.
This species. though its outline isvariable to this extent. is safely referable to i'olsrlla by the edentulous
hinge and dl.'veloped anterior area.
Volsella persisted from Devonian to
Recent without any striking evolution.
and there arc a great number of comparable species to this. If comparedwith most Upper Triassic and Jurassic
species. this is characterized by a welldetined and inflated anterior area and

42.0

7. 0
7. G

strong concentric line on the shellsurface which are fairly regular at theintervals and \Yidely spaced.

Modiolus imbrira/us (Sow Ellll\') (1818.
Vol. 3: l\loRIHs and LYct-:TT. 1853:
LoRIOL. 1883: Cox, 193!i, 19,10) is a wellknown iVIicldle Jurassic species with awide distribution. It has a more expanded and flattened anterior area.
weaker concentric lines and a smallerumbonal angle than the present species. JVvtilus jurensis RonmR (Cox, 1935.
pl. 15. figs. lS-11) from British Somaliland seems to vary to some extent in
the development of anterior area. andCox (1937. 1940) regarded it as an intermediate form bet ween Volsdla and"11ytilu~. Among the Somaliland specimens the one in figure 16 is fairly
similar to this species in the outline
and surface-markings, but the anteriorarea of jurensis is generally not so welldeveloped. A!ytilus arbenzi l~EN7. (193S)
from the Upper jurassic "Mytilus-Schi-ltaru HAYAMI

158

chten" of Alps differs from this also
in the less 'developed anterior area. although the concentrics and generaloutline arc very similar to those of
the holotype. Modiola lwffmanni NILssoN
and Jl1. /usofauira in BoEHM ( l!l03) fromthe Lias in Portugal may he related
forms to this. But the concentrics are
more delicate than in the present
species.
Occurrenrr:-Al1 specimen procured
from a black shale of lower Hettangian Niranohama formation of Shizukawa group at Niranohama in Utatsumura, ~liyagi Prefecture (Province ofRikuzen).

Volsella sp. indct.
Plate 23, Figure -1.

Only a fragmentary left valve (l\Hvl
This form is considerably similar to the pre-ceding species in the modioliform outline anddevelopment of anterior area. Hut theconcentric lines are by far weak thanin balwl'e!loides. and there are numerous
fine radial ri blets on the an tero-ventral
slope of shell-body.
Occurrence :-Black shale of DomerioToarcian Shinatani formation of Kuruma group at the upper stream of Teradani in Daira. Asahi-machi. Toyama
Pre f.
2722) is at hand.

Genus Arcomytilus AGAssiz. lR-12.

Type species :-Mytilus j>rrlinatus Sowr:RIW (lR21). Upper Jurassic.
:lrcomytilus dairensis KonAY Asmand l-IAYAMA. new species.
Plate 23. Figures 5 and 6.

Description :-Shell small, mytiliform,
well inflated. much longer than high(holotype. MM 2734. 25.0 mm. long; 18.5
mm. high; !1.0 mm. thick): test thin:umbo terminal and sharply pointed
with an umbonal angle of about 50
degrees; hinge-margin nearly straight.
short. passing gradually into evenly
arcuate posterior margin: anterior
margin almost straight; anterior carination weak. marginal, leaving noanterior area; surface ornamented withmany radial ribs which are fairly
prominent in central area and crossedby conspicuously wide-spaced concentric lines of growth; nothing is knownof internal structure.
0/Jsrrl'rtlion and Comparison :-Only two
imperfect left valves arc giYen.
If
compared with ever-described speciesof .1rromylilus from Middle and Upper
Jurassic, its anterior area is undeveloped and its umbo very terminal.
The radial ribs are comparatively weakin the posterior area of I he holotype
and neither bifurcated nor divaricated.
Orrurrcnce :-·Procured at the lower
stream of Daira River in Asahi·machi.
Toyama Pref .. but its exact horizon is
uncertain.

/\rcomytilus sp. indet.
Plate 23, Figure 7.

Only a small right valve is before
hand. Shell subtrigonal. modioliformwith a developed anterior area. much
longer than high (l\il\1 2736. 19.5 mm.
long; 12.0 mm. high); hinge-margin
fairly convex; anterior area rounded.
elongated along anterior margin of
slwll-body. defined from it by a shallow groove; umbo not terminal. locatedat about a fifth of hinge-line from theanterior end; anterior carination very
weak: surface ornamented with roughly spaced radial ribs which become
fairly strong in ventral area.

Liassic Volsella, Jl{ytilus and sume other Dysodont Species in JapanThe radial ornaments of this form
remined one of .4rcom_riilus, although
the anterior area is similar to that ofVolsella. Tts specific identification isdeferred until sufficient materials will
be procured.
Occunetrce:- Sandstone of DomerioToarcian Shinatani formation at Shinatani in Omi-machi. Niigata Pref.
Genus M_rtilus LINNi,, 1758.

rrpe species ·-Myti!us edu!is

LINNE( 17:18). Recent.

Subgenus Fa/cimytilus Cox, 1937.
Type speciPs :-Mytilus supmjureusis Cox(1935), Upper Jurassic.
.l{!'li!us (Falcimyt ifus) sf ricapi!!at us
HAYAMI. new species.
Plate 23, Figures 8-ll.

159

Description :-Shell large, equivalvc,
sickle-shaped. weakly carinated, moderately convex, more or less longer thanhigh; test thick for genus; hingemargin slightly sinuated. forming anapical angle of about 50 degrees withhinge-line; umbo terminal; anteriorarea very narrow, ill-defined; weakcarina running close to anterior margin. incurving behind beak; anterior
slope in front of carination very sleep
and nearly vertical, while posteriorarea is comparatively flattened; surface marked with tine concentric linesof growth and numerous dense radialcapillae ,,·hich are persistently perpendicular to growth-lines and fairly
prominent in early stage. Internally.
a narrow ridge for adherence of ligament elongated close and subparallel
to hinge-margin; umbonal and dysodont teeth absent.

Jlfeasuremelll in mm.

LengthHeightThickness

Ilolotype (MM 2726) Bivalved specimen
Paratypc (MM 2727) Hight internal mould
Paratypc (MM 2728) Left valve

72.5
75. 5

68.5
74.5
19.5

29.0

ObsPrvation and Comparisol/ :-In a juvenile left valve (Fig. 8) anterior area is
not developed, if compared with theadult holotypc (Fig. 9). The numerous
radial capill:le, which arc charateristicof this species, are seemingly weakened out through growth.
The sickle-shaped outline and thicktest are similar to those of Lycettia Cox(193/), especially of Lycrtlia lwwlaris(LYcF.rr). Faint radial capillae arealso discerni blc in the lectotype ofthat species (Cox, 1937, pl. 17. fig ..1).
although he mentioned nothing of thecapillae. This species, however. cannot
be referred to Lycettia, because of thenarrow ligament area and absence of

24.0

?
3.5

urnbonal septum. The edentulous hingearea is more likely to that of Falci·
my! ilus.
Occurrence :-Common in sandstones ofDomcrio-Toarcian Shinatani formation
at Shinatani in Omi-machi. Niigata
Pref. and rare in black shales of Liassic" Tsuchizawa formation" at Kuruma
in Kitaotari-mura. Nagano Pref. (Prov.
of Shinano.)
illy! ilus (Fa!cimyt ilus) stricapi!latus
1-JAYAMI SUbSp.

Plate 24. Figures 1 and 2.

There are two similar specimens to
the preceding species, which are bro-

Jtaru IIAYAMI

160

ken and secondarily deformed.
If
compared with typical stricapil/atus.
this form has a more elongated hingemargin and a more gently curved
ventral one.
Occurrenrc:-Rare in sandstones ofthe Middle Liassic Negoya form<ttion
at Neiriclani in Kurobe National Forest,
Toyama Pref.

Mylilus (Falcimyl ilus) hemnirus
HAYAMr, new species.
Plate 24. Figures 3 and 4.

Description :-Shell medium for genus.
subtriangular, mytiliform, weakly carinate, not strongly in llated; test I h in;

hinge-line nearly straight, very long,occupying more than two-thirds ofshell-length; anterior margin straight.
forming an umbonal angle of about 50
degrees with hinge-line; umbo terminaland pointed; anterior carination becorning obscure towards ventral side.
leaving a narrow anterior area in its
front, \Vhich is scarcely defined fromanterior slope; the greatest convexity
lying close to anterior margin. andposterior area well flattened; surface
smooth except. for fine concentric line~of growth. Internally, umbonal anddysodont teeth absent; ligament borneby an internal ridge which runs subparallel to hinge-margin.

Jl!casuremcnt i11 111111.

LenglhHeightThickness

Holotype (MJ\f 2723) Right vah·eParalypc (MM 272·1) Right valve

43. 0+
49.0

31. 0+

4.5+
9.5

Obser/Ja/ ion and Comf>arison :-The I ribal name in KonAYAsnt's manuscript is
applied in the specific denomination.
The holotype (Fig. 4) is possibly moreor less l"ompressed secondarily, while
the paratype (fig. 3) is slightly broken
in postero-dorsal area.
judging from the terminal umbo,
distinct carination and edentulous
hinge-area, this species may be refer·
red to subgenus Falcimytilus Cox. The
subtriangular outline. narrow anteriorarea and long hinge-margin arc distinguishing characters of this species.
illy/ ifus mirabilis LEPSI us va r. I imorensis
KRul\mEcK (1923) from the middle Liassic J{rti/us-hank of Timor is somewhat
similar to this. but in that variety thehinge-margin is much ~hortcr and theanterior margin deeply sinuated. Shellconvexity is very strong in timorensis.
but rather weak in the present species.
iv!ytilus (Falcimytilus) sub/aevis Sow ERDY

4~.5

( 1R~3. Vol. 5: J\1oRRis and hcETT, 18:1:{)
from the Bathonian in England also
resembles to this in the triangular outline. In the British species, howeYer.
the antero-vent raJ margin is more orless sinuated. while it is nearly straightin this species.
This species differs from the proceding stricapil/atus in the more trigonal
outline, elongated hinge-line and absence of radial capillae on the surface.
Occurrence :-Common in black shalesof Liassic .. Tsuchizawa formation" atTsuchizawa and Kuruma in Kitaotari·mura. Nagano Pref. (Prov. of Shinano).

Mytilus ( Falcimylilus) llt'rauirus
J·IAYAJ\11 subsp.
Pl:t1'" :!1. Figure :i.

There are two large right valves
which are fairly similar to the preceding species. But the shell is moreLiassk l'olse/la, .Aiytilus and some other Dysodont Species in Japanstrongly inflated with a sharp anteriorcarination than in typical heranirus.
The anterior margin is almost straightin lwranirus. but fairly sigmoidal inthis form. On the other hand. thisform is somewhat similar to J11ytilus(Falcimytilus) suprajurensis Cox in thestrong carina and broad anterior slope
before it. but the anterior margin ismore abruptly sinuated below the umbothan in that species.
Occurrence:- Rare in black shales oflower (?) Liassic Kitamatadani formation of 1\uruma group at Kitamatadaniin Asahi·machi. Toyama Prcf.
Myti!us (Fa!cimytilus) sp. indet.
Plate :.!-1, Figurl's 6 and 7.

Only two internal moulds arc given.
Shell medium. crescentic in outline,
strongly inflated, almost as long as high(Mi\I 2733, ·12.0 mm. long. :mo mm. high.
J:W mm. thick); hinge-margin rather
short, convex, passing gradually into
posterior margin without any angulation; antero-ventral margin gently
sinuated; carination very weak; anterior area small. ill-defined from anterior slope of shell body in internalmould; umbo terminal. pointed; internal ridge running subparallel andclose to hinge-margin; umbonal anddysodont teeth absent.
This form resembles Mytilus mirabilis
L~-:Psrus var. limorensis K•wMnEcK (19~3)in the general outline and strong
shell-convexity. But judging from thegrowth-lines of I imorensis, that variety
seems more trigonal than this form.
Occurrence :-Sandstone of Negoya formation at Nciridani.
Family Ostreidae LAMARCKGenus Ostrea LINNf:, 1758.

161

Type species :-Os/rea edulis LINNE. Recent.
"Ostrea" a sp. indct.
Plate :!4, Figure 8.

Represented by an internal mould of
a left valve. Shell large. inequilateral.
vertically elongated, strongly convex,
thick ClVll\1 2742, 4·1.0 mm. long. Sl.S mm.
high, 15.5+ mm. thick); I igament area
wide, striated. provided with a large
deep opisthocline acute-triangular central pit, which is bordered on eachby a convexity; adductor monomyarian.
located slightly posteriorly to center;inner surface irregularly folded, lacking any radial plications.
Judging from tile irregular outline
and ligament structure. this is safely
placed in the wide sense of "Ostrea ".
DouvJLI.i·: (190.!) established genus Liostrea on the basis of Osfl·ea sublamellusa
DuNKEl~. The genus has been said tohe distinguished from Os!rea (s. s.) by
being more equivalve and by lacking
radial ribs on the left valve. The
name has been hitherto applied for thegreater part of jurassic non-coiled"Ostrea ·· by most authors. The present form belongs probably also to
Liostrm. But the specific determination is impossible at present. because
the right valve and the exterior of theleft valve of this form arc as yet
unknown.
Occurrence :-Procured from a sandstone of Domerio-Toarcian Shinatani
formation at Shinatani.
" Ostrea " b sp. indet.
Plate 2•1. Figure !l.

There are several smaller Osirea-likeshells which were collected from theKuruma group at various localities. It

162

ltaru IL'\YAMJ

is difficult to say, however, which ismost typical as a species. because its
outline is very variable.
The illustrated specimen is an internal mould of a right valve. Shell small.
pyriform, fairly convex in early stage
but later it becomes concave; ligamentarea narrow, having a nearly <!Cline
obtuse-trigonal central pit bordered oneach side by a weak convexity; adduc·
tor single, circular, posterior to center;inner surface smooth.
This form resembles '"Oslrea" sp.
(NAKAZAWA.1955) from the Carnic Nabae
group in the pyriform outline and thereversing of shell-convexity in the
middle stage, but the central pit iswider than in the Carnic form.
Occurrence:- The illustrated specimen
procured from the Liassic" Tsuchizawa
formation" at Tsuchizawa in Kitaotarimu ra.
Family Pteriidae MEEKGenus Pteria ScoPoLI, 1777.

(=.·lvicula BJwGuJEJ{E, 17!:ll)1)Pe sPecies :-Mylilus hinmdo
Recent.

LINNE,

Explanation of Plate 23
Volsella ba!.•et•cl/oidcs IIAYAl\11. new species. .. ............................................................... p. 156
Fig. 1. Right valve (Ml\•I 2719) X l. holotype. Loc. black shale of Lower Hettangian
Niranohama formation at Nira:10hama in Utatsu·mura, l\liyage Prefecture (Province of Rikuzen).
Fig. 2. Left valve (!VII\1 2730) X I. paratypc. Loc. ditto.
Fig. 3. Internal mould of a lett valve (!VIM 2721) x1, paratype. Loc. ditto.
Volsella sp. i ndct. ......................................................................................................... p. 158
Fig. 4. Clay cast from the external mould of a left valve (.MM 2722) x 1. Lac. blackshale of Toarcian (?) Shinatani formation at the upper stream of Tcradani in
Kurobc National Forest, Toyama Pref.
;lrromytilus dairensis KoDAYASIIl and IIAYAMI. new species. .. ..................................... p. 158
Fig. 5. Left valve (Mi\'1 2734) X l. holotypc. Loc. line sandstone at the lower stream of
Daira River in Asahi-machi. Toyama !'ref. KoBAYASHI coil.
Fig. 6. Left valve (1\IM 2735) xI. Loc. the same as fig. 4 .
.·lrcomytilus sp. i ndet. ................................................................................................... p. 158
Fig. 7. Left valvc (M.M 2736) x 1, Loc. sandstone of Toarcian Shinatani formation at
Shinalani in Omi-machi. Niigata Prcf.
Mytilus (Falcimyl ilus) striwpillatus IIA Y AMI, new spcci cs. .. ........................................... p. 159
Fig. 8. Left valve (MM 2728) x2. paratype. Lac. black shale of Liassic "Tsuchixawa
formation" at Kuruma in Kitaotari·mura, Nagano Pref. (Prov. of Shinano).
Fig. 9. Bivalved specimen (MM 2726) x l. holotype. Loc. the same as fig. 7.
Fig. 10. Left valve (1\Il\1 2729) x 1. Lac. ditto.
Fig. 11. Internal mould of a right valve (M!>I 2727) X 1, paratype. Loc. ditto.
Oxytoma (?) sp. i ndet ................................................................................................... p. 163
Fig. 12. Left valve (MM 2740) x 1.5. Loc. line sandstone of Domcrio-Toarcian Shi natani
formation at the upper stream of Karwyama-dani, in Omi-machi, Niigata Pref.
Fig. 13. Left valve (MM 2741) x\.5. Loc. ditto.
All illustrated specimens arc kept in the Geological Institute, University of Tokyo.
Photo by UEKI.

HAYAMI:

5

Liassic Dysodonts

Pl ate 23

163

Liassic Volsella, Mytilus and some other Dysodont Species in Japan

Pteria (s. 1.) kitalwmiensis
HAYAMI,

new species.

Plate 2.J, Figures 10 and 11.
? 190.J. Gen·illia trigo11a YoKOYAMA, four.
Col!. Sci. Imp. Unir. Tollyo. Vol. 18,!lrt. 6. p. 1:!, pl. :.!, ftg. R (non fig. 7)

Description :-Shell highly incquivalve.
trigonal, pterifonn. compressed; test
thin; ldt valve moderately inJlated
but right one is nearly flat; hingcmargi n straight, almost as long as
whole shell-length; anterior margin
long, almost straight. forming an angleof 40 degrees or so with hinge-margin;
ventral margin short. abruptly curved;
posterior one slightly sinuated in dor·sal half. forming a small pointed pos·terior auricle; anterior wing large.
elongated along anterior margin. de·fined from shell-body by an obscure
groove in right valve; umbo fairly
protruded above hinge-line in left
vah'e. but not in right; surface mark·ed wilh more or less fluctuated con·cent ric lamellae; inner surface of left
valve undulated by some broad radialplications; hinge and ligament struc·tures unknown.

J\leasurement in mm.

LengthHeightIlololype (M1'vl 2737) Right external mould
Paralypc (l\IM 2738) !.eft intt.:rnal mould

38.5
26. 0+

24.0

l.S

21. 5-J·.., r:

Obserl'at ion and Comparison: -Six speci ·mens arc at hand. but all are more orless broken or deformed sccondari I y.
Though the hinge structure is unknown.
the present species is referable to Ptrria(s.l.) in view of the general outline andhorizontal ridge along hinge-margin.
The ridge appears in Gerl'illia praecursor
QuENSTF.OT (HEALEY. 1908) and Several
species of the Bakevelliidae, bul morecommonly in lVIesozoic "Pleria ''.
This species is so similar to thespecimen of Gerl'i!lia trigona YoKoYAMA(190-t. p. 12. pl. 2. fig. 8, non fig. 7) inthe trigonal outline and concentriclamellae. that that specimen possibly
is conspecific with this. But the specific name. trigona, is applied to theform as represented by the other typespecimen (fig. 7) which has an explicit
ligament structure of Ballevellia·type.
The writer designated already it as
the lectotype of /Jalwvellia trigona (Ymw·
YAMA) (1-fAY... :\11, 1957. p. 51, pJ. 2, fig. 2).
Occurrence :-Rare in black shales ofThicknessi . •)

lower Hettangian Niranohama forma·tion at Niranohama.
Genus Oxytomn l\lmK. 186-t.
Type species :-A1•icula inequh·all'is So·(18]8, Vfl{. 3. p. 78. pJ. 2-ll, Jig.
~. non fig. 3). Middle Jurassic (=A1•icula
miinsleri BRoNN in GoLnFuss, 1836.)
WEIHW

O:ryl oma (?) sp. i ndet.
Plate ~4. Figures 1~ and 13.

Several ill·preservecl left valves areat hand. Shell small, prosocline. con·vex ; hinge long. straight with apointed posterior auricle. below which
posterior margin is fairly sinuatecl;
umbo more or less rising abo\·c hinge·
margin. located at about a third ofshell-length from front: surface sculp·
tured with about 15 radial ribs of asingle order of prominence.
Nothing is known of the hinge·
structure and the right valve. Judging
from the general outline and ornamen·

164

Itaru HAYAMI

tation. this seems a member of O.lyfoma.
but the specific identification is defer·
red until sufficient material will be
procured.
Occurrence :-Rare in fine sandstoneof the Lower Shinatani formation(Domerio-Toarcian) at the upper streamof Kanayamadani in Omi-machi. Niigata Pref.

-------------------------------

Explanation of Plate ~4
Jl1ytilus (Fa!cimylilus) slricapiflatus IIAY.-\Ml subsp. . .................................................... p. 159
Fig. l. Internal mould of a left valve (MM 2731) xI. Loc. sandstone of rvliddle Liassic
Ncgoya formation at N<.:iritlani in Kurobe National Forest. Toyama Pref.
Fig. ''
Internal moult! of a right valve (MM 2732) X 1. Loc. ditto.
Jl!ytilus (Falcimytifus) heranirus 1-IAYAI\H, new species.
.. .................................... p. 160
Fig. 3. ]{ight valve (i'vHvl 2723) x 1.:1. holotype. Loc. black shale or Liassic "Tsuchizawa
formation .. at Kuruma in Kitaotari-mura, Nag;:mo Pref. KoBAYASHI coli.
Fig. 4. Right valve (l\11\I 2724) X 1. paratypc. Loc. eli Ito.
li!ylifus (Flilcinn·ti/us) herauirus H:\YAMI subsp.
..................
.. ............ p. 160
Fig. 5. Left vah·e (I\11\1 2730) X I. Loc. blacJ; shale or Lower (?) Liassic Kitamr~tadaniformation at the upper stream of 1\itamataclani in Kurobe National Forest!llyti/us (fit!cimylilus) sp. inclL-t. ..
..................................................................
p. 161
Fig. 6. Internal mould of a right ,·alve (l\IJVT 2733) xL Loc. the same as fig. I.
Fig. 7. Internal mould or a ldt vahe (M:\1 2n5) xl. Loc. ditto.
"Oslrea" :1 sp. indct. ................................................................................................... p. 161
Fig. 8. Gypsum cast from the internal mould of a left valve (MM 2742) xl. Loc.
sandstone of Toarcian (?) Shinatani formation at Shin:'ltani in Omi-machi.
:--ii igat:1 Prcf .
.. Ostrea" b sp. inckt.................................................................................. p. 161
Fig. 9. Internal mould of a right valve ().11\1 21-13) xi. Loc. black shale or Liassi~.:"Tsuchizawa formation" at Tsuchiznwa in Kitaotari-mura.
Plaia (s. 1.) kitakamiensi.~ !Lw AMI. new species.
.. ........................................... p. 163
Fig. 10. Internal mould of a left valve (MM 2738) xI. pr~ratype. Loc. bbck shaleof Lower Hettangian ;..;iranohama formation at Niranohama in Utatsu-mura.
l\liyagi Pref.
Fig. 11. External mould of a right yal\·e (Mi\1 2737) xl, holotype. Loc. ditto.
All illustrated specimens are kept in the Geological Institute, University of Tokyo.
Photo. by UEKIHAYAMI :Liassic Dysodonts

Plate 24

4

9

Liassic Volsella. il!ytilus and some other Dysodont Species in japan

Hcferem·c8]. (1!103), Description de Ia F:tune
des CoLil'hL·s de Perei ros. Sen•. ;;cui.
Portugal, Tom. 5.
Cox. L. R. (1935). jurassic Gastropoda :1ndLamellibranchia in Mesozoic Palaeont·ology of British Somalilanu.
- - (1937). Notes on Jurassic L:unellibran·chia. V. On a new Subgenus of Mytilus
and a new Jl{l'lilus-likc Genus. }'roc.
illalacol. Soc. London. Yo!. 22.
- - (19·10). The Jurassic Lamcllibralll:hFauna of Kachh (Cutch). Pat. Jmlir:a.
Ser. 9. Vol. 3, Part. 3.
llAYAlll!. I. (1957), Liassic Bakerc/lia injapan, ]ajxm. ]our. Geol. Gcogr., Vol. 28,
Nos. 1-3.
lcmKAWA, 1\. (1954). Late Triassic Pelecypods from the 1\ochigatani Group in
the Sakuradani and Kilo Areas. Tol-:ushima Prefecture. Shikoku. Japan-Part
I. four. brst. Polytec. Osaka City Uuiv.,
Vol. 1, No. 1.
KoBAYAsm, T. and lcmKAWA, K. (l\150),
BoEHM.

165

Late Triassic Jllylilus, l'olsclla. Pleura·phorus and :\lyrlCOIIclw from the Sakawa
Basin in Japan. ]our. Fac. Sci. Univ.
Tokyo, Sec. 2, Vol. 7. Part :;, No. 2.
KRUliiBECK. L. (W~a), Zur Kcnntnis desJuras der lnsel Timor sowie des Aucel·
len·Horizontes von Seran und Buru.
Jlufiwnt. 1'011 Timor, Lief. 1:!.
1\loJWIS. ]. and I. YCETT, ]. (1853), A .tvlono·
graph or the Mollusca from the Great
Oolite. /'alaeunl. Soc. Loudmr.
1\AKAZAWA. K. (1955), A Study on the
Pelecypod-fauna of the Upper Triassic
Nabae Group in the Northern Part of
Kyoto Prefecture, japan. Part 3. Halohi ids and others. ,U£'111. Col/. Sci. Uni11.
Kyoto, Scr. n. Vol. 22. No. :!.
RENZ. II. H. (1935), Zur Stratigraphie und
Pal:iontologic dt:r Mytilus-Schichtcn imiistlichen Tci I dcr Pr~alpcs romandcs.
!:'dog. Geo/. llell•etiae. Vol. 28.
Smn;IWY. ]. (llHS-~3). l\Hneral Conchology
of Great Britain, Yo/s. 3, ·1. 5.
YoKoYAMA, !\1. (1!!04), On some Jurassic
fossils from Rikuzcn. ]our. Coil. Sci.
Imp. Unit•. Tokyo, Vol. 1/:i, Art. 6.

Trans. Proc. Palacont. Soc. japan. N. S., No. 2!1, pp. 166-168. pl. 25, ;\pri I 10, 1958

331.

.MESOZOIC PLANTS FRO.M THE TETORI SElUES,CENTRAL HONSHU, JAPAN (Part 1)*
TATSUAKI KIMURA
Senior High School att. Tokyo Univ. of Education.

'¥Jiii/t!ltlfl7)flfit'Jftfi. :t.:-q-;-: L-iimi'Jiffi89llf ('f-Jfl(~4llf) q-;J:$:1J.r,, Cladophlebis slzinslmellsis
TATEIWA i!l:,mUH... t~q-;·r:, O::.;lL~!'3cii&L'I::.

:<fi.:ffiltEJ:<f\:'i'(:l:, (t{;V>-(rl)~[ilf1-<::JbJ.,o ;,f; t:i' ~IV]

Introdm•tion

In this paper the writer describes amarked species collected from the Tc
and Tel horizons in the section of Tamodani1>. (T. KIMURA, 1951)
In describing this short report. thewriter expresses his sincere gratitude
for the helps of Dr. II. FuJIMOTo andS. E1-mo who kindly guided him during
this study.
Description of species
Pteridophyta
Fi I icales Incertae Sed isForm-genus Cladop!tlebis BIWNGNIART
C/adop!tlebis slzinslmcnsis TATE I w A
Plate 2!3. Figures 1, 2; Text-figure 1.
1929. C/adojJ/ilebis shiuslzw·usis TATEIWA, fig.
24.
19-10. Cladophlebis shiusluwusis 0JSIII, p. 285,
pl. XX, figs, 5-6; pl. XXI. figs. 5-7.

Diagnosis :-Frond: Hi pinnate. probably large in size, rachis, :2-3 mm. l hickmeasured on impression and traversed* Recci ved !\lay 10, 1957; read Apri I 3, 19531)83£€-ft166

by ·1 weak ridges on its surface.
Pinnae; oblong or obovate in shape,
attached to the rachis at a low angle(about 30 degrees) suboppositely, flexible in habit, tapering gradually towards the acuminated apex and overlapping each other laterally.
l'innules; thin in character, setclosely together, attached by the lower
half of the base or whole of the base,the upper basal edge often making a
deep sinus. Variable in shape, mostly
long in length and narrow in width,with deeply serrated or lobed margin,
each lobe with subacutely or acutely
pointed apex.
l~cproductive organ is not preserved.
!Jrscript ion of specimens:- Pl. ~5. fig.
1 shows three imperfect pinnae which
arc posterior portion of a frond. Pinnae are very flexible in habit, andbending backward at their midcourse.
Pinnules are long and narrow, themargin is mostly lobed. each lobe
acutely or sub-acutely pointed andacutely directed forward just likethe pinnules themselves. Nerves arc
simple. the mid-nerve is considerably
marked. but the secondary nerves aremostly obsolete.
Text-fig. 1 shows a part of pinna of
this specimen.

•

!vfesozuic l'bnts from tlte Tl'lori Series, Central Honshu, japan

J(j7tion regarded as Lower Cretaceous inage.
Characteristic features are recognizable, that is being smaller in the sizeof pinnules, having strongly serrated
or lobed margin and showing simple
nervation as seen in the present species,
in Japanese Upper Jurassic to Lower
Cretaceous fern-like species. namely,

C/aclopltlebis e/cgaul is sima 01sm
C.
lwlwit!llSis 01slll
C.
pan>ula 01sm etc.

Text-lig. 1. Cladop/ilcbis shinslmensis
TAin-.JwA: a part of pinna, x2. (Td-OUOI)

Pl. 2:i, fig. 2 shows the· middle portionof frond.
In the ultimate portion of frond, pinnae are very closely set and the margin of pinnule is lobed in the posterior
portion and is getting entire 10\vard~
the ultimate portion.
Remarks :-The writer's specimens arc
referable to the species figured originally by TATEIWA and described by OJsm (19-10) on the same specimens fromthe Shinshu'l formation in Korea which
is younger than the Naktong 2> forma1)
2)

~:lfi

**

It is dillicult to distinguish this species from above mentioned in cases ofimperfect specimens and only an isolated pinna.
The occurrence of this species inJapan had been unknown before thewriter's present paper.
Horizons:- Tc and Td, in the sect ionof valley Tamodani, the Ryogadani 3lalternating bed of sandstone and shale.
the Upper ltoshiro 1> sub-group. Fukui
Prcfectu re.
Sample monber:-Td-9!73, Td-9151. Td0001. Td-oom. Td-0003, Td-9159. Td-9143.
Td-9160, Td-9161. Td-9164, Td-9149 andothers.
Hcfcrent~'S

T. (1!157). On the stratigraphicalhorizons uf Podozamilcs Reinii GEvu:J(in the Tdori Sl!rics. japan. Studr·inJ:
report of I lu: Senior /ligfl Scltool attachedto the Tokyo Unh·ersily of Eduction. iVo. 1.
Otsm. S. (19-10). The Mesozoic Floras ofjapan. founr. Fac. Sci., lloklwido Imp.
[initl., Series IV. Nos. 2-·1.
'L>\.TEIWt\, I. (1929). GeologicalAtlas of
Chosen (Korea). No. 10, Keisyu-EiscnTaikyu and Wakwan Sheets. 1/50000.
Ceo/. Surv. Chosen (Korea).

KIMUtc\.

3)
4)~7-ft-:cirats

168

Tatsuaki KIMURA and Kciji SuzuKI

Explanation of Plate 25
Figs. I.~. Cfadophfebis shiuslweusis TATEI\\' A ..................................................................... 166
I. Posterior pi nnac (Td, Reg. no. Td-000 I):.!. Middle portion of frond (Ttl. Reg. no. Td. 000:.!)(Photo. by T. Kll'.ll.!RA, all in natural size.)------------------

Figs. a. b. Cen·is Endoi Suzui·u. new speci.:s ................................................................. 170
Collected by 1\. SuzuKI from the Middle part of Fujitogc formation exposed along
upper of the Hara River, about !JOO m. northeast of Kobusegawa. Yamato Town.
Yama·gun, Fukushima Prefecture, Honshu, japan.
Fig. c. Cercis sp ............................................................................................................. 170
Collected by K. SuzuKI. from the lower part of the Fujitoge formation exposedat Shirokozawa Lingite·mine, Yarnato Town. Yama-gun, Fukushima Prefecture,
Honshu, japan.
Figures are all natural size.
(The specimens of figs. a-c arc stored in the Institute of Earth Sciences, Department ofArts and Sciences, Fukushima University.)KIMURA:

Cladophlebis shinshuensis TATEIWA

SuzuKI: Cercis in Japan

Plate 25

Trans. Proc. Palaeont. Soc. Japan, N. S., No. 29, pp. 168-171. pl. 25, April 10. 1958

ON THE OCCURRENCE OF CERCIS IN .JAPAN*

332.

KEI.J I SUZUKI
Department of Arts and Sciences, Fukushima Universityn;;j.:l.::t.>it.; Cacis ~q)i~HUI.:"JI. ··c:

rH:.:··e(t. Cl'l'cis (:l:li!~fJS (!JlftJc-:4~fi:?.·(!(:t) f!.i:J;, ;-_q).w~q)f~Bc L-r 1926 1f.l.: KRYsH-rovovrTc'' tJ:r&:g:.rR.t ,,,tl..!il!XLt.:tq)tJ:;t;,.;, L:tr·L· ::.ttltmf
Jtlt:lft*'!i.: 1{, t -?\ •'CI. ·~q)<:'. Cc>rcis l.:..:f ·t ~ ·c I) :i'J· ·') t.:i:J:;h L 1. •• ~[ffi. ~JiJJJf.JAdlillitJtmq)!'ilitR'i ( l:$i'flW!Utf.i:\ · L "F$~tfillt) J: CJ. Cc>rcis l:::.7i -r ~1t;sm'~.!'·1J.\ L/.:(7)·;:: · ;OOL'L<.:Cq)-Ffl~: "Ccrris

1\'ndoi"

n

<

·t.:

t (,jr~ L t:iJ: . .: (i'J.ffll(:J:, m'J:fofi · ft;s"fol1t q)Jt•liQJ:. c ~) :.:. rp[I~,{(l)~1{, q) J.:l(W;··e:Ji,-.,.

* lii!itfi

cics, it is doubtful whether C. iapnnimm
really belongs lo the genus Cerris. because it was described upon poorly
preserved material.
Eight living species are known of
the genus in the Northern Hemisphere.
Of them, two occur in North America.
five in China and one grows in the
I'v1editcrranean region. The contrasl ing
living and fossil distribution of thegenus is interesting. particular!~ fromthe present discovery of additional undoubted material from japan. which isthought to have bearing on the problem relating to the remote distributionof the known species.
The distributions of the known liv·ing species are as follows. Cercis cana·
densis LJNNAEus is restricted in distri·bution to along the l\Iississippi Riverand C. reniformis ENGJ.EI~ from Texas to
the Sierra i\Iadre. both in the warm
temperate region. Of the five Chinese
species. Cercis rhinensis BnNGE is widelydistributed in Central and Northern
China and C. mcemosa OuvER at altitudeof 4000-5500 feet in Central China. while
the other three species are known only
from Central and Northern China.

The genus Cercis is one of the oldgenera of dicotyledonous plants. bein_gknown to date back to the Paleogene
and is readily recognized by its wellmarked foliar characters such as outline and nervation. The fossils of thisgenus arc rather common in the Paleo·
gene of North America but show adecline in the Neogene. which is moremarked in the l~ccent.
In North America there arc three
species recorded from the Paleogene
deposits. but on! y one or two from thel'vl ioccnc and PI ioccne. Besides these
there is one species known from Europe(BEER. 1856) and two from Asia. Thus.
so far as literature is concerned, thegenus seems to be confined in its gco·
logical and geographical distribution
to the Northern Hemisphere.
Among the Asiatic species. Cerds
miochinensis 1-Iu et CnANEY (IIJclO. p. !'i].
pl. ~6. figs. 3-5) has been reported from
the Miocene of Shantung. China. aud
C. japonicum KRYSJITOFOVITCII ( 1926, p. 13,
pl. 3, fig. 2) from the l\1iocenc of Echi·
go. Japan. However. of these two spc·

* Received June 20. l~l57: read june 15, 1957
169

Keiji SuzUKI

170

As a result of the writer's geologicalobservations in the Tertiary terrain oi
the Aizu Basin, Fukushima Prefecture,
well preserved specimens referable to
the genus Ccrcis have been found.
These described in the present article
arc from the l\Iiocene or Lower Pilocene Fujitoge formation. This formation is rich in plant remains andamong them such as the following arecharacteristics. name! y : Fagus j>alaeo·
crenata 0KuTsu, Zellwva Ungcri Ko\'ATs
and Liquidambar formosana HANCE.
Descriptions of Crrcis species
Cercis Endoi, sp. nov.
PI:Jte 25 Figures a. b.

Description :-Leaves broadly ovate
but asymmetric in outline. ca. 9 em.
long and ca. 8.5 em. wide; asymmct ric
form especially distinct at the apicalpart, margin entire. base slightly cordate, apex acuminate. Seven palmative
primary nerves stout, lateral primaries
on each side of midrib asymmetric.
inner ones diverging from lhe midrib
at angles of ca. 30 degrees and outerones from the midrib at angles of ca.
75 degrees, respectively reached near
the margin upcurving toward the apical part. Secondary nerves numerous.
the ones diverging from the midrib atangles of 40-50 degrees. and not foundin the (O\Yer half, the others diverging
from 1he lateral primaries at angles oi
ca. 40 degrees, making camptoclrome
ncar the margin with the primary
nerves. Lateral tertiary nerves distincl.
making the polygonal meshes with theother certiary nerves. Texture rather
thin. Petiole not preserved.
Remarlls and Compmison :-The pre·sent materials are all asymmetric inform being especially distinct at theapical part. as shown in the annexed
figures. The leaves at the tip of theyounger twigs of Cercis cl1ilzensis BuNGE
is the most similar to the present one
among the living species, and the leavesof C. mccmosa 0LJVF.I~ resemble the present one in the leaf form and the nervation. However. the leaves at the tipof the young twigs of Cere is chinensis
BuNGE. never have very acuminate apex
and seven obvious palmative nerves as
seen in the present materials.
Of the known fossil species. my specimens resemble certain form of Ccrcis
mioclzine11sis Hu et CnANEY (1940, p. 51.
Pl. 2G. Fig. 3) in the leaf form andnervation, but differ from it in theapical form of leaf and in the detailsof the nervation. Of the American
fossil species, none is similar to thepresent specimens. The present specimens are somewhat similar to Ccrcis
cyc/ojJhyl!a A. BRAUN (in 1-IEF.r~. 185G, p.
107, Taf. CXXXIII, Fig. 35) from thel\1iocene of Switzerland.
Locality:- The upper course of Hara
Riyer, about 900 m. Northeast of Obusegawa. Yamato Town, Yama-gun. Fukushima Prefecture, japan.
Geological l10rizon :-Middle part of
the Fujitoge formation. Upper l\Iio·cene or Lower Pliocene.
Cercis sp. inclet.
Plate 25, figure c.

i\ single fragmental leaf. This leaf
is small. and it differs from Cercis
Endoi SuzuKI by both cordate apex andnervation. The leaves with thesecharacters are sometimes recognizable
at the lower part of the twigs of Cercis

chinensis BuNGE.
Locality :-Shirokozawa

lignite-mine.
Yamato Town. Yama-gun. Fukushima
Prefectiure, japan.

On the Occurrence of Ccrcis in Japan

Geological horizon:- Lower part of the
Fujitoge formation; L'pper 1\Iioccne.
The writer wishes to offer his 'var·
mcst thanks to Dr. S. ENno for his kindencouragement to my palaeobotanicalworks. The writer is also indebted to
Dr. K. HuzwK.\ for his kind advices.
References
IIEcR, 0. (1856), Flora Tertiaria Helvatiac.

Bd. III.
llu, H. H. and CnANEY, W. It (1940). A l'vliocene Flora from Shantung Provinn·.
China. Carne. lust. Wash., Pub/. No. 507,

171

Pt. 1, pp. l-1-11, pis. 1-50.

A. (1926), Conlribution to
the Tertiary Flora of Kwannonzawa.
l'ruv. Echigo, Japan. Ann. Russ. Pal. Soc.,
Vol. 6, pp. 1-:!·l, pis. 1-3.
HuzioiC'\, K. and SuzuKI, K. (195-l), The
Flora of the Shiotsubo Formation of the
Aizu Lignite-Field, Fukushima Pref ..
japan. Trans. l'ruc. Palaeont. Soc. Japan,
JV.S., No. 1-1. pp. 133-1·12, pl. lti.
SuzuKI, K. (1951). The Geology of the West
Rq::ion of Aizu Basin. Fukushima Prefecture. Pt. I. Cc·ntral Area. (in Japa·ncsc). }ourn. Gcol. Soc. japan, Vol. 57,
No. tj/:J. pp. 4 l!l-456.
KRYSIITOFOVJCH.

Trans. Pruc. Palacunt. Soc. japan, N. S., No. 29, pp. 112-174, April 10, 1.958

333.

A SHORT NOTE ON LONSDALEOIDES TOR!YilMAI MINA'l'O*
l\IASAO J\IlNATO and MAKOTO KATO
Department of Geology nnd Mineralogy, llol<kaido University.

Lonsdaleoides /oriyamai i\IINATo 1::.-J\ '"L: !J1=lfJHllllt/ll Lousdaleoides loriwmllli MJNATO CT>t!,'i;~jllJ~o J: 0:11!1!\lii:."':>\ '"C.\f\~~ ;!Lt:g ~i)l{.ll~. ~iml:ltlJ•I) ~ Q:>):.v:>\ ··Cfii'il:)ifJ:;lJi(&~.fj-? !.:..

~

Very recently the senior author received several specimens of a fossil coral which were sent by Prof. E. TAJ<A·
uAsHI of Yamaguchi Univ. for study.
All these specimens arc specifically
identifiable to J.onsdaleoides toriyamai
l\1JNATo with certainly by microscopical
study, as will be later described in thisiE$·1lflli~

phical horizon of this coral is, for certain his so-called Cm a=l'rofusuliucllazone. Accordingly it is proposed here
to revise the former description in respect to the locality and the geologicalhorizon of the holotype of /,onsdalroides
loriyamai MINATo.
The new specimens lately sent by
Prof. TAI<AIIAsm to the s1•nior author
were collected, according to his information by letter. from two localities:
one is quite the same place, where theholotype specimen of this species wasfound. while the other locality is at alimestone quarry at !sa. l\Iine City. Yamaguchi Prefecture.
Although the exact place of the second locality is unknown today to thewriters in more exact detail. the speci·
mens must have been discovered at a
locality where the Lower Permian limestone should be developed, according
to the geological map recently published by Prof. ToRJYAMA.
However, it is almost impossible toimagine this coral to show such a long
geological range as from the Middle
Carboniferous to the Permian in age,
therefore inquiry was made of Prof.
ToRJYAMA, whether the Middle Carbonifeous deposits are really developed
near lsa .or not.

nolt>.

This species was first established by
the senior author based on materials
presented br Prof. R. ToHIYA~JA, Kyushu Univ .. which were. according to his
verbal information. collected by himfrom the Fusuliuclla limestone developing at 1\yowa-Mura. Mine-Gun, Yamaguchi Prefecture.
· This was reported by the seniorauthor in a former paper (l'vi. MINATO,l!l55).

1\leanwhile. information has recently
been received through letter from l'rof.
ToRJYAMA that the hololype specimenof this species was not collected from1\yowa-l\iura. but was trucly derived
from the limestone developing at ashort distance northeastward from Ohkubo. Ouda (Ohta)-1\Iachi. (now MitoMachi}, Mine-Gun, Yamaguchi Prefecture, and further. that the stratigra--------• Received Feb. 22, 1951; read Sept. 28. 1957.

172

A Short Note on Loustf,rfcoidcs /oriyamai 1\'IJNATO

In this regard Prof. ToRIYA!'.L\ h; kindenough to inform that there develops
limestone which perhaps is correlable
to his l'rofusulinclla zone even in thearea near Isa, especially in the northor northeast of !Vlaruyama, although
the existence of such formal ion was
not indicated in his geological map
lately published. So it is now suppos·
ed that thi::; species lll:Jy have been also
derived from the Middle Carboniferous
deposits at the second locality near Isa.
Before going into a description of
new specimens, the writers wish to ex·press sincue thanks to Professors TA·
KAIIASHI and ToRIL'\.1\IA for their kind
help in this study.
Description of species

Ccitlls Lonsdafcoides IIERITSCII, 1936
em. ~hl"'ATo, 19:i5
Lonsda/eoides loriyamai l'I'IJNATu
Text-fig.
llJ:i5.

Lonsdalcoid<'s /oriyamui, MJNATO: Japanese Carboniferous and l'ermian
Corals. jour. Fac. Sci. lloldwido Unil· ..
ser. Ill, rol. 9. no.:!, pp. 165-Hii', pl. 3.

Text-fig.

113

fig. 6; pl. 16. fig. i': pl. 2:>. tigs. 1. 2. 3:
tcxt-tigs. 21, 22.

Corallurn fasciculate. Corallites cylindrical, usually closely disposed andrarely in contact with each other, whilesometimes they are dendroid and newcorallites seem to be arisen from the;Jeripheral part of the old ones.
Cal icular diameter ranging from 10
I o 20 mm. in I he ephebic stage.
Outer wall verv thick, the inner sur·:·ace of which r~presents considerable
convexity towards the central area in
the interseptal space, as is shown in
Text-tig.
Outer zone provided with lonsdaleoid
dissepimcnts in generally very wide,
but its width of is not uniform
throughout the corallitc in a cross sec·lion, even in the mature stage; in one
side this is more wide than the otherand it is often disappearing in one por·tion. In an immature stage such Ions·
daleoid disscpiments are a little deve·
loped and in a still younger stage they
are completely lacking.
Septa in two orders. .tvlajor septa
thick. numbering ~4 to 3,1 in the ephebic stage. extend into the columella

Lonsdaleoides /oriJ·amai Jl.hNATO (X 4)17-1

l'vlasao MINATO and Makoto KATO

but never directly unite with it in theephebic stage. although only the counter septum is firmly united with thecolumella in the early stage of the ontogeny. The major septa start to growfrom the outer wall in the younger
stage and there is never left an area
entirely without septa near the outerwall. As the corallite grows larger,
there appears an area occupied bylonsdaleoid dissepimcnts and thus the
major septa become not connectedwith the outer wall. Minor septa alternating with the major ones arc fairly
long, usual I y about two-thirds thelength of the latter. Both of them arestrongly thickened at the thecal region
to form a distinct stcreotheca.
There is no marked fossula.
The axial complex is much complicate in the mature stage, the outlineof which is sub·elliptical in cross section. lt is composed of a thick medianplate, axial tabellae and septal lamellae; all of them are strongly thickend
by stereoplasmic deposits, and thus itreminds one of the columella of thegenus Carrinopllyllum.
In ncanic stage, the septal lamellae
and axial tabellae arc, however, moreclosely disposed with respect to eachother and in still younger stage, theaxial structure of this species is observed to be a compact. solid columella,
I ike the genus Cionodendron or Lop/wp!ty/1idiwn.
Localities and Regislratio11 /lumbers:Ohkubo, Oda, Mito-Cho (Machi). MineGun, Yamaguchi Prefecture. U. H. R.
12708 (i-v).
lsa quarry. lsa-Cho(l'vlachi), Mine-City,
Yamaguchi Prefecture. U. H. R. 1270!1(i-iv).

Heferenccs
W. N. and SMITH, S. (1923), Rugose
Corals from the Burindi Series (Lower
Carboniferous) of New South Wales.
(Juart. ]our. Geol. Soc., l'ol. 79, pp. 156-

BENsoN.

171.
HJ,RITSCH, F. (Hl36). Korallcn der Moskaucr·,

Gshel· und Schwagerinen-Stufe dcr
Karnischcn Alpen. Po/aeontogr. Bd. 83,
Abt. A. S. 99-162.
JEFFORDS. R. M. (19-17), Pennsylvanian Lo·
phophyllidid Corals. Uuiv. Kansas, Pole·on/. Con/ rib. 110. 1.
l\hNATO. M. (1955), Japanese Carboniferous
and Permian Corals. ]our. Fac. Sci. llok·
lwido Uni1•., ser. IV. vol. 9, 110. 2, p. 202.
TORIYAMA. R. (1954), Geology of Akiyoshi.
part. I. Jl!em. Fac. Sci. Kyuslm Univ., ser.
D, vol. 4, no. 1, pp. 39-97.

Trans. Proc. Palacont. Soc. Japan, N. S., No. 29, pp. 175-183, pl. 26, April 10, HJ58

331.

ON SOJ\TE CEPIIALASPID OPISTHOBRANCHIAFROM THE JAPANESE l\IIOCENE*
JUNJI ITOIGA\V A
Institute of Earth Science, Nagoya U11i\"Crsitr

I=?Wflttmili/H&&Bi:

Jll\iJ~·Vil'+·Jili¥.0)3~tl'fn·GimL-t.:liJilrn~i'i lGfiHJ::'"':)\'1: ~·fil!O)

tli:tiH:a-:f]\ ·. l'uji(J hiyoshicnsis, Ret usa (CoelnPh.rsis) slmlmboremis, T'nlrulella millnl'ilsis, V.
yammu:hii. Eocyliclma habei, E. loldensis. /)ecor~(er ena <7> 7 '$JffillO);li:JiiCH:iJ., t.:, f:/.(l Jll ~="Acleon" sp. indd.
Pupa lliyoshiensis n. sp.
Rin{!imla (Ringiculina)

fntroduction and Acknnwledgcmcnt!-1Man)' Recent and fossi I species of the
Cephalaspicl Opisthobranchia of theseas around and the Pliocene and laterrocks in Japan have been described andreported by various authors. However.
the present knowledge of the l\Iiocene
species of this subclass is very meagre.
but for some contributions made by
i\l YoKoYAMA. T. TAKEYAMA and Y.
Onm:A. This scarcity of criteria ischiefly attributed to the minute andfragile shells. so as good preservation
be rare and as aptly be overlooked.
The present writer is studying thestratigraphy and paleontology of the
Miocene rocks that are called the First
Paleo-Setouchi supergroup (IKF.BE. 1951;
lToiGAWA, l!l55a. b. l!l56). He has already reported his works in part on
the Mizunami and Tsuzuki Miocene
groups in the mid-west region of thiscountry.
In this paper, 16 forms of the opisthobranch mollusks of the Cephalaspid
group are dC'scribed. These are collected from the Mizunami, Tsuzuki and
Fujiwara groups. They are:milwensis TAKE·

YAMA
R. (Ringiculina) sp. indet.
Cyliclmatys sp. i ndct.
Retusa (Coe!ophysis) slmkuborcnsis n. sp.
R. (Cor.loj>hysis) sp. i ndct.
Vo/1'1/lella minoensis n. sp.
V. _\Yimauchii n. sp.
F. sp.
Eoscaj>hander corpulenta (YoJWY AM A)
Eocylic/ma habei 11. sp.
E. /okicnsis n. sp.
E. a.(fabilis (YoJ<oYAMA)
Cyliclma sp. i ndct.
Dccorifer ena 11. sp.
The writer is indebted to Prof . .T.

l\fAKIYAMA for instruction in preparation of this paper and to Drs. T. KuRoDA and T. l-L-wE for informations onmalacology. Thanks are clue to l\1r.
T. SAKAMoTo who kindly offered thespecimens of the Fujiwara group andalso to 1\lr. Katsumi YAMAUCHI who assisted the writer in the fields.
Descriptinn of SpecieR

Family Pupidae
Genus Acleo11 MoNTFORT, 1810

* HeceiYcd June 22, 1957; read June 15, 1957.

"Acteon " sp. inclet.
175

Junji lTOIGAWA

176

Plate 26. Figure 1.

Shell small. ovate; spire short. coni~cal; protoconch unknown, remaining
whorls !1. with deep suture forming ashoulder at the upper edge of thewhorls; body-whorl large. occupying
about three-fifths of the shell length;surface with about 30 spiral cords inbody-whorl (unknown on the upper
whorls). separated by narrow sulci;
sulci finely lalliccd by raised lines ofgrowth; aperture ovate. narrow andbluntly pointed above. rounded below;
columella part unknown.
Length. 8
mm .. diameter, '' mm.
Many cast specimens are at hand.
All the columella parts arc broken. sothat it is clillicult to determine thegeneric status whether Acteou or Puj>a.
Apparently it is resembled to Acteon
uij>ponensis !YAMAKAWA. 1911), but thelatter has a longer and narrower bodywhorl.
Gcologiral occurrence:- Toyoda f onnat ion, Fujiwara group.
Lorality :-SF 6 (Fujiwara Range, Pujiwara. Nara City).

I!olotype :- JC1500012, Para type:- JC
1500013.

Remarks :-This species is closely all icc! to Pupa strigosa (GouLn, 1859) ingeneral feature, but it is distinguishedfrom the latter by the features of columella. Acteon ozawai 0TuJ<A. 1!!;)8, fromthe Shohara lVIiocene. is another species
allied to this, but present species has
a more ovate shell with 2 columellafolds.
'J)•pe Locality:-M 20. River-side cliffof Shukubora valley about 100m. S\Vof the bridge at S of Shukubora, IIiyoshi-cho. i\Iizunami City.
(;Mfogical occurrence :-Shukunohora
sandstone, l'vlizunami group.
:lssociated fossils :-Same with the oneof Retusa shukubon'ltsis n. sp.
Family Ringiculidae
Genus Ringicula DEsrrAYEs. 1838
Subgenus Ringiculina MoNTERosATo. 18811

Ringicula (Ringiculina) minoettsis
TAKF.YAMA

Plate 26, Figures 3, 4.

Genus PlijJa RtiorNG. 1798
1935.

Pupa hiyosltiensis n. sp.

Ring/Cilia (Riugicufe/la) minoensis T A·
The Venus. z·ol. 5. p_ 8"1. pl.
6, figs. 36-39.
KEYAMA,

Plate 2!i, Figures 2a, b.

Shell small, ovate; spire conical withnarrowly canaliculated suture; whorls
7, somewhat convex; surface sculptur·
cd with 5 punctured spiral grooves except smooth embryonal whorl; bodywhorl large, occupying about five·sevenths of shell length; aperture narrow. \viclened and rounded in front;
outer lip arcuated; columella somewhat callus with 2 folds; lower foldstrong, upper one weak. narrow.
Dimensions :-Length, 7.0 mm., Diameter. 3.2 mm.

( ;eolomcal occmnmce :-Shukunohora
sanclst one. l\Iizu na m i group.
[,ocality:-"Nl20 (Shukubora. lliyoshicho, Mizunami City, Gifu Prefecture).
Ringicula (Ringiculina) sp. indet.
Plate 2fi, Figure 5.

Shell small, ov:uely globose, withmoderately short spire; protoconch unknown, remaining whorls 4. slightly
convex, separated by impressed suture;
sculpture consisting of fiat-topped spiral ribs separated by narrower inter-On some Ccphalaspid Opist hobranchia from the japan.:se Miocene

spaces; ribs 9 on the penultimate
whorl; last whorl large, slightly lessthan 3/4 the total shell height; complete aperture, lip and columella unknown except the trace oi 2 columellafolds. Length, 3mm, diameter, 2.lmm.
Ill-preserved ·I specimens are at hand.
This species is distinguishable fromthe allied 1\Iiocene species such as R.
ninohensis 0TuKA, 1934, R. ninohensis
kiiensis T AKEYAMA, 1935, R. minoensis'L'\.KEY AMA. 1935 and R. fragilis TAKEYAM,,, 1935 in havin~ the globular shellwith lower spire and large body-whorl.
Investigation will be postponed until
discovery of sufficient material.
Geological occurrence :-Toyoda formation. Fujiwara group.
Locality :-SF fi (Fujiwara Range, Fujiwara, Nara City).
Family Atyidac
Genus G~\'liclmatys KuRODA et 1-IABE, 195~

Cyliclmatys sp. i ndet.
Plate 26, Figure 6.

Shell small. rather thin, ovate. slightly truncated at the apical end. roundedat the lower end; spire concealed, concave. but not perforated; surface withfine spiral striae <tnd longitudinal
growth line; aperture narrow. arched.
dilated below; outer lip thin. somewhat arched; columella margin short.
faintly thickened with a fold; umbilicus narro\v, slit-like. Length. 5.0mm., diameter, 2.7 mm.
Only two specimens were obtained.
The ovately subcylindrical shell form
is a remarkable feature not known
among the japanese Recent and fossil
species. Complete description is postponed.
Geological occurrences :-(1) Kubohara
sandstone, l\Iizunami group, (2) Tsuki-

177

yosh i mem bcr. Ivl izunami group.
Localities :-(l) I 111 (Kamigiri. Iwamura-cho, Ena-gun, Gifu Prefecture).
(~) M 17'-0 ( .1\nabora. Akiyo-cho, l\Iizunami City).
Family Retusidae(;enus Retusa BRowN, 1827
Subgenus Coe!op/iysis. FiscHEl~. 1833
Ret usa tCue/ophysis) slmlwboreusis n. sp.
Plate ~G. Figures 7a, b. 10.

Shell small. subcylindrical, constricted in the middle. t rune a ted at the apicalend, rounded at the lower end; spire
deeply sunl;en; body-whorl equal towhole length of the shell; surface \vithsomewhat coarse longitudinal grO\vthlines; aperture narrow. as long as theshell length. linear above. abruptly
widened below; outer lip straight orslightly constricted in the middle;
columella lip oblique without fold;umbilicus more or less opened.
Dimensions :-Length, 2mm., diameter.
L2mm.
J!olotype:-JC 1!>00002. ParatyjJe:-JC
1Sll0003.
Rcmar/ls :-This shell is distinguishable from R. minima YA~IAKAWA, 1911 inhaving the shorter shell. more deeply
sunken spire. opened umbilicus andnot having any columella fold.
Type Loca!ity:-l\1 20, River-side cliffof Shukubora valley about 100m. SWof the bridge at S of Shukubora, Hiyoshi·cho. Mizunami City, Gifu Prefecture.
Geological occurrence: - ShuK unohora
sandstone, l\1izunami group.
Associated fossils :-Miogypsiua kotoi
HANZAWA, Opcrculina romp!cmata japonica
HANZAW A.
Aloidis nisataiensis 0TuKA,
Cavilucina llitamurai (HATAI et Ntsn·AMA),

liSjunji lTOIGAWA

Turbo ozmrai (0TUKA) etc.
Retusa (Coe/ophysis) sp. indct.
Plate 26. Figure II.

Shell small. subcylindrical. stuggy,rounded at the lower and upper parts;
spire sunken. immersed in later whorls;
body-whorl equal to shell length; surface with coarse growth lines; aperture
narrow, as long as shell height. linear,slightly expanded at the lower onethird: outer lip straight. more or lessconstricted in the middle: inner lip
oblique, with a fold. Length. 2.(i mm ..
diameter. 1.8 mm.
Only one specimen is under examination. But stuggy shell is so characteristic as a more material will show
the ne\v species. This form differsfrom R. s/mlwborensis n. sp. in posessing a more stuggy shell with shallow
sunken spire and a columella fold.
Geologicaloccurrence:- Tsukiyoshi
member. lVIizunami group.
Low/ily :-M ·100 (Tsukiyoshi. Akiyocho, i\Iizunami City. Gifu Prefecture).
Cenus Vo!oulc/la NEwToN, 18!11
Foh•ulcl/a minucnsis n. sp.
l'latc 2li. Figures ~a. b. 9.

Shell small, solid. clongatcly subfusi·form. centrally ventricose, with asharply pointed apex and narrowly
rounded base; whorls convolute, only
the last visible, produced to a sharp
point above. apical part provided witha feeble split; surface polished. sculptured with weak, fine spiral striae on
the terminals; aperture as long as theshell, very narrow and sharply arcuate.
thin, extending above to the apex andbase; inner I ip covered by a narrow
callus; columella short, oblique; um·

bilicus narrow. slit-like.
Dimensions :-Length, 3. 'i mm., diameter. 1..! nun.
l!o!otype :-]C 150000·1, Paratype :-]C
1500005.
Remarks :-This shell is easily distinguished from r·. mdiola (A. ADAMS,
1802) by the more narrowly fusiform

shape with s!Jarp apex and absence ofcolumella fold. V. tokunagai 1\'L'\KIYAMA,
19~7 is closely related to this species.
bul the former has a subcylindricalshell with more blunt apex and columella fold.
Type Localily:-M 211, Shukubora. Hiyoshi·cho, Mizunami City. Gifu Prefecture.
Geological occurrence :-Shukunohora
sandstone, Mizunami group.
:1ssociall'dfossifs :-Same with the oneof Ret usa shulwboreusis n. sp.
l'ull•ulel/a ywJumcllii n. sp.
l'latc 2G, Figures 13. 1·1.

Shell small. elongate. subcylindrical.
tapering to a pointed apex and narrowly rounded base; whorls convolute;surface polished. smooth in the middle,
spirally striated on the terminal portions; striae indistinct, weak; .aperture
as long as the shell length, narrow.
linear, slightly expanded below; outer
lip arcuate. thin; inner lip covered bya narrow callus; columella oblique,smooth. without a fold; umbilicus narrow, slit-like.
Dimensions :-Length. ca. 5 mm., diameter, 1.5 mm.
1-/olatype:-]C 1500006, J>arat:ype:-]C1500007.
Remarks :-The elongate form of thisnew species is the conspicuous featureand is distinguished from the anotherspecie!-l. The preceding species is likethe present species, but the latter has

On some Ccphalaspid Opisthobranchia from the Japanese Miocene

the more elongate and narrower shellwith the blunt apex. This species isnamed in honor of l\1r. Katsumi Y.>\1\.tAucHI who helped the writer in thefields.
Tvpe Loca/il_r :-f.1 20, I~ i ver-sicle cliffof Shukubor:1 valley about 100m. SWof the bridge at S of Shukubora. Hiyoshi-eho. l\Iizunami City. Gifu Prefect u rc.
G1•o/ogical occurrences :-(1) Shukubora
sandstone. Mizunami group. (~) Toyoda
formation, Fujiwara group.
Localities :-(1) lVI 20 (type locality),(2) SF () \Fujiwara Range. Fujiwara.
Nara City).
.:hsocia/{'d fossils:-( I) Same with theone of Retusa slmlwlwrensis n. sp., 12\
Ch/amys lwkmt•ai SAI'AMoTo tl\!S). Crassatelliles mal?iyamai SAKAMoTo (l\·IS).
Nassarius simizui (}ruK A, Protorot ella
yuantaniensis MAKJYAMA. Ca/yptraea tuburrr (huKA.

Volwlel/a sp.
Plate :26. Figure 17.

Only two imperfect specimens arc
under examination. It resembles to V.
minol'llsis n. sp. but has more elongatedand cylindrical shell. Specific determination is postponed.
Geologicaloccurrence:- Tsukiyoslli
member, Mizunami group.
uJcalilies :-1\1 31-1 and l\I 31-2 (Kujiri. Toki City. Gifu Prefecture).
Family Triclidac(;en us Eoscap!umdcr HAnE. 1!lS~
Eoscap/iall{/(•r corpu/ntla (YoKoY A:-.tA \Plate 26. Figure 21.
19:2G.

Cyliclma corpu!enla YoKoYAMA. ]our.
Fac. Sci. Imp. (]nit'. Tokyo. sec. 11. mi.
1, pt. 7. p. 217, pl. 28. figs. 3, 3a.

179

Shell large, thin, ovate. tapering toapical end; whorls convolute. only thelast visible; vertex concave but details
unknown; surface with spiral threads
and growth lines; spirals fine. numerous, broader than interspaces, intercalated by fine, weak striae; growth
I ine fine. cancellate at the i nterspacesof spirals; aperture large. widened on
the lower half; outer lip arched; inner
lip short. slightly arched; no umbilicus.
Length. 34.3 mm .. diameter. 1!1.5 mm.
Only few ill-preserved specimens
were obtained. This is a characteristic
species as noted by YoKoYAMA. This
species falls to the genus Eoscap/wnrkr
in general feature, i.e. shell form.
sculpture etc.
Geological occurrences :-(1) Kubohara
member. 1\Iizunami group, (~) Yamanouchi member. Mizunami group, (3) Tsu·kiyoshi member, l\lizunami group, (4)Toyoda formation. Fujiwara group.
l,ocalities :-(1) I Ill (Kamigiri. lwamura-cho, Ena-gun. (;ifu Prefecture).
(2) 1\I 5:.! (Sakurado. l\fizunami City), (:~)
M 17'-2 (Ana bora. J\kiyo-cho. i\lizunamiCity), (4) SF 6 (Fujiwara Range. Fujiwara. Nara City).
Genus Eocy!idma KuimDA et HABE, 1952£'oryliclma habei n. sp.
Plate 26, Figures l5a. h.

Shell small, solid. elongated, cylindrical. slightly tapering at both ends.
more or less constricted in the middle
on the immature specimens; whorls
convolute. only the last visible; vertex
truncated and deeply excavated; surface polished. smooth except the fine
growth lines; aperture narrow, linear.
widened below; upper margin produced
upwards; outer lip nearly straightdescending rounded base; columellajunji ITOTGAWA

180

margin short. oblique. with a obtuse
fold; umbilicus almost closed.
Dimensions :-Length. 2.7 mm., d ia·
meter, 0.8 mm.
Holotypc:-JC 1500008, Pamtypc:-.TC] 500009.
Remark~:-This shell is similar to
Eocyliclma mu.wrslliensis (ToKUNAGA, 1906)
but the former has a larger and moreelong:fted shell with the wider dilation
at the base of the aperture. Focy/iclma
brmmsi (YoKoYAMA. 1920) closely related
to this new species in general form
but the transverse line in the former
species is not visible in the latter.
Type Locality :-'fi·I 20, R i\·er-sidc cliffof Shukubora valle~' about 100m. SWof the bridge at S of Shukubora, Hiyo·shi·cho. l\Iiwn:11ni City, Gifu Prefec·
lure.
r.(•o{of!iral occurrence:- Shu ku nohora
sandstone. i\lizunami group.
.·lssocialed fossils :-Same \Vi th l he associated fossils of Re/11sa (Coelojl/1ysis)
slwlmborensis n. sp.

Eocyfidnw Iokirnsis n. sp.
Plate :?6. Figures 16a. b.

Shell small. solid. cylindrically ovate,
centrally ventricose. spire convolute:vertex depressed. concave with a small
pit at the bottom: surface polished.
smooth. in the middle, spirally striatedon the terminal portion; sculpture con·sisting of 10-J:·l subequal. distant striae
on the base and 1-9 similar ones on
the lop; growth line visible; aperture
narrow. widened at the both ends. up·
per margin produced upwards andlower margin rounded below; outer lip
rather straight; columella somewhat
callus with a obtuse fold; umbilicalchink narrow.
Dimensions :-Length, 3.9 mm., dia·met cr, 1.9 mm.

llolotype:-JC 1500010,

Paratype:-JC

1500011.

Re.marhs :-This shell resembles to
Hocy!ir/ma brmmsi (YoKOYAMA, l!l20) but
differs in its short and ovate shell. It
is easy to distinguish the present spe·
cies from !•.:. soyoae HAnE, 1954 hy thespiral ornamentation on the terminal
portions.
Type Locality :-M 20. River-side cliffof Shukubnra valley about 100m. S'\Vof the bridge at S of Shukubora. Hiyo·shi·cho. Mizunami City. Gifu Prefec·
t ure.
Geological occurrences :-(1) Shukuno·
hora sandstone. (2) Tsuki~roshi mem·ber. (3) Kubohara member. ~lizunami
group, (4) i\Iiyamura member, Tsuzuki
group.
J.ocalities :--(1) M 20 (Type Locality),(2) M 17'-0. l\I 17-1 (Anabora, Akiyocho, l\Iizunami City), (3) I 2·16 (Naka·nishi. Ye~maoka-cho, Ena-gun, Cifu Pre·
fecture), (4) 0 13-1 (6fuku. Ujidawaracho. Tsuzuki-gun, Kyoto Prefecture).
:lssociated fossils :-(1) Same with theone of Retusa (Coe/op/1ysis) shulwborensis
n. sp., (2) Felmzie!la usia (GouLll), Tapes
siratoriensis (OTuKA), Turritella s-hataii
NoMURA, Euspira mrisensis .1\'IAKIYAMA.
Nassarins simizui Chmu. (3) Nippono·martia nalwmurai I rmim, Tapes sirafori·
cnsis (0TUKA). Pillucina yokoyamai (0Ttr·
KA), Nassarius simizui ClruKA. Protorotella
rlrprcssa MAKTYAMA, (4) "Os/r('a ·· sp ..
Nassarius simizui 0TUKA, Cerithium an·
tismn (YoKoYAMA). C. cfr. otukai NoMURA.
EO!:yliclma affabi!is (YoKoYA~IA)
Plate 26. Figures 12, 20.
1926. C)·lic/111a af(abilis YoKOYAMA.]nur. Fac.
Sci. Imp. U11h•. Tokyo. sec. 1!, z•o!. !, pt.
7. p. 216. pl. 28. figs. 1. 2.

Only ill-preserved cast specimens areat hand. But it seems to fall withinOn some Ccphalaspid Opisthobranchia from the Japanese Miocene

the limit of ·· Cyliclma ·· affabilis YoKo·
YAMA seeing the general features. The
deeply excavated but not perforate
vertex indicates that this species be·longs to the Genus Fnrylic/ma.
Gen!ogicnl occurrences :-(1) l\laki siltstone. (2) Tsukiyoshi member, Mizunami
group, (3) Miyamura sandstone. Tsuzu·
ki group.
Locn/ities:-(1) I 37?.. I 185 (Agi. Agimura, Ena-gun. Gifu l'ref.}, I 124. I 533,
I 383 (NE of Nouchi. Agi-mura). l 158(Shinbara. Agi-mura), (2) .M 32-1 (Kujiri. Toki City. C;ifu Pref.), (3) 0 113(Chaya. Ujidawara-cho, Tsuzuki·gun.
Kyoto Prefecture.).
Genus C_r!idma Lm· EN. I S.ti

Cyliclma sp. indet.
Plate ~fi. Figure 18.

Shell rather medium. thick, cylindrical; spire concealed; vertex truncated.
only showing a very Rmall excavation.
but not perforated: sculpture reticulated consisting spirals and longitudinal
growth lines: spiral threads fine. dense.
coarser to the upper part; aperture
dilated anteriorly. apertural marginand columella part unknown; no umbilicus. Length. ca. 8.5 mm., diameter.
3.2mm.
Only two broken specimens arc athand. However. it is evident that thisshell belongs to the genus Cyliclma be·cause of the features of the apex. It
appears as a promising unique species.
(;m{ogicnl ocmrrences :-(1) Kubohara
sandstone. i\lizunami group, (2) Toyoda
formation. Fujiwara group.
Localitil•s :-(1) l 113 (Kamigiri. Iwamura·cho. Gifu Prcf.) (2) SF G(Fujiwara
Range, Fuji,..mra, Nara City).
Genus Decorzfer lnrrDALE. 1937

Decorifer ena n. sp.

181

Plate 26, Figures 19a, b.

Shell small. solid. sul>cylindrical.
pointed above and rounded below:
whorls five: spire conical, occupying
about two-sevenths of the shell length.
somewhat terraced but not shouldered;apex prominent. obliquely immersed inlater whorls; suture distinct, shallo\\·,not channeled; surface smooth except
for tine growth line; aperture nearly
equal to the height of the body-whorl.
thin, narrowed above, dilated androunded below; outer lip straight;
inner lip with a callus: columella foldobsolete.
Dimensions :-Lengl h. !i.~ mm.. diameter. 2.0 mm.
1/olo/yj>c : -]C 13001()3, !'amlyjw : -]C
130016:1.
Remarks:-This shell is similar to D.
imignis ( Pn.snnY. 190·1). but it d iff crs
from the living species in having thehigher spire but no columella fold. D.
!o 11gispirafa (YAMAKAW A. 191 I) is anotherallied species. but it has narrow shellwith the terraced higher spire and thefolded columella.
Type J.ocali/y:-1 2·1fi. Nakanishi. Yamaoka·cho. Ena-gun. Gifu Prefecture.
Geo/of.[iml occurrences: -(1) Ku hnhara
sandstone. (2) l\laki siltstone. (3) Shukunohora sandstone and (4) Tsukiyoshi
member, 1\lizunam i group.
Loca/ities:-(1)-a. 1246 (type Joe.). I
4~Hi (Nakanishi, Yamaoka-cho. Ena-gun.
Gifu Pref.). I 58 (Agi-mura. Ena·gun.
Gifu Prcf.). I 110 (Kamigiri. Iwamuracho. Ena-gun. Gifu Pref.). (1)-b. I 111.
I 113 (Kamigiri. Iwamura-cho. Ena·gun.
Gifu Prcf.). (2) I 1. I 121 (Agi. Agimura. Ena·gun. Gifu Prcf.). (3) l\I
20 (Shukubora. Hiyoshi-clw. 1\Jir.unamiCity), (4)-a. 1\I 2. M 5-2 (Tsukiyoshi.
Akiyo-cho. l'VIizunami City). (4)-b. M
17-2. J\117'-0(Anabora. Akiyo-cho. l\1izu·
nami City), (4)-c. M ?.2-1. M 31-1

182

Junji IToJGAWA(Kujiri, Told City).
.1ssocia!ed fossils :-(1)-a. Same withthe associated fossils (3) of Eocy!iclma
tolliensis n. sp., (1)-b. Dosinia 110/llltrai
OTuK A, Nipponomarcia ualwmurai I K EBE,
Euspim meiscnsis i'viAKJYAMA. Turrile!las·hal aii Noll.lURA. G~vcymeris minocusis
hmGAWA. Nassarius simizui 0TuKA, (2)Nipponomarcia ualwmumi IKEBE, Tapes
siratoriensis (OTuKA), Euspira meisensis
l\IL~KJ YA MA. Nassarius simizui OTuK A, (3)same with the one of Ret usa (Coclopl!ysis) slmlwborensis n. sp., (4)-a. Cycliua japonica lC'\MAnA, Sanguinolaria
minoensis (YoKOYAMA), Saxolucina k·hataii(0-ruJ<A). Vicmya yokoyamai TAKEYAMA,

Explanation of Plate 26
Page
Fig. 1. ",k/emz" sp. i ndct .. X 3, Loc. SF 6(Fuji wara Range. Fujiwara, Nara City) .......... 175
Figs. 2a. b. Pupa hiyo.~Jiieusis n. sp., Ilololypc, Reg. No. JCl 500012. X 3. Loc. J\1 20(Shukubora.
IIiyoshi-cho, .1\Iizunami City. Gifu Prcf.). ..
. ............................................... 176
Figs. 3. 4. Riugimla (Riugimlina) minoensis TAKF.YAMA. x5. Lo,·. M 20(Shukubora, Hiyoshicho. l\1iwnami City. Gifu Pref.). ....................
. ......................................... 176
Fig. 5. Ringicula (Ringiculina) sp. indet .. X 5. Loc. SF 6(Fujiwara Range. Fujiwara. Nara
City).
. ............................................................. 176
Fig. 6. Cylidma/ys sp. indct .. x3. Loc. I lll(Kamigiri. Iwamura·cho. Ena·gun. Gifu Pref.).
............................. .......................................................................... ........
.. .. 171
Fig~. 7a. b. Retusa (Coelopltysis) slmlmborrnsis n. sp.. ITolotype, Reg. No. JC1500002, Loc. l\1
20 (Shukubor:J, IIiyoshi-cho. Mizunami Cit~·. Gifu Prcf.) ....................................... 177
Figs. 8:1. h. Vo!l'lilel/a minoensis n. sp .. !Iolotype. Reg. No. JCI50000~. x5. Loc. Same as above......................... ........................ .............................. ......
. .............................. 178
Fig. 9. Volntlella minocnsis n. sp .. Paratypc, Reg. No. JC150U005. X 5. Loc. Same as above.
Fig. 10. Retusa (Coe/ophysis) slwlmborensis n. sp. Paratype. Reg. No. JC1500003 x7. Loc. Sameas above.
Fig. 11. Ret usa (Corloj>liysis) sp. indet.. xri, Loc. l'vl 400 (Tsuk i yoshi. Akiyo-cho, 1\IizunamiCity). ............ . .... .. ... ........
.. .......................................... 178
Fig. 12. Eocyliclmtt ajJabilis (YoKOYAMA). x2. Lac. r 373(Agi. Agi-mura. Ena-gun, Gifu Pref.).
................................... ............. ..................................
.. ............................. 180
Fig. 13. l'oh:ulella yamaucliii n. sp .. Ilolotype. Reg. No. JC1500006, x3, Loc. M 20(Shukubara.
lliyoshi-cho. Mizunami City)...........................................
. ............................. 178
Fig. 1<1. Volmle//a :yamauchii n. sp., Paratype, Reg. No. JC1500007, X3. Loc. Same as abo\·e.
................... .................................................................
...............................
178
Pigs. 15a, b. l~'ocyliclma lmbei n. sp .. 1-lolotypc. Reg. No. JCEiOOOOS. x7. Loc. Same as abo\·c .
......................... ................................................................ ...................... ................. 179
Figs. Hia, b. Eocy!iclma loldeiisis n. sp .. llololype. Reg. No . .TC15110010. x5, Loc. Same as above.
................................................................................
. ...................... 180
Fig. 17. Voh·uM!a sp .. X 5, Lo..:. 1\'1 31-1 (Kamigiri. Toki City, Gufu Pref.) ..................... 119
Fig. 1.3. Cylidma sp. incl<:t .. x3. Loc. I 113(Kijiri. lwamura-cho. Gifu Pref.). ...............
181
Figs. 19a. b. Decortfcr ena n. sp. Holotypc, Reg. No. JCI3001G3. X 5, Lac. I 2·16(Nakanishi.
Yamaaka·cho, Ena-gun, Gifu Prcf.). .....................................
.. ................ 181
Fig. 20. Eocyliclma affabi/is (YoKOYAMA), X 3, Loc. M 32-1 (Kujiri, Toki City, Gifu Pref.).
Fig. 21. Eosmplumder corpulenta (YoKOYAMA). X 1. Loc. I. 111 (Kamigiri. Iwamura-cho, Enagun. Gi fu Pref.) ....................................................................................................... 179

On some Cephalaspid Opisthobranchia from the Japanese 1\liocenc

T·icarye/la ishiimw (YoKoYAMA). Ceritlzium
lwn1'1111rai Fu.11TA and 0GosE, (4)-b. same\vith the associated fossils of ,..'ocyliclma
tokil'nsis n. sp., (4)-c. Tuniiella s·halaii
NoMURA. Euspira mciscnsis M AKIYA:o.tA,
Prolorotella d1•pressa l\IAKIYAMA, Tapes
sirat oriensis (Chu KA), Nipponomarcia ua·
lwmurai I KEOE.

Refcrenees
Ft'JITA. K. and OGosE, 5. (1!150. 1!151). Lithologic CI<Jssi tication of the Cenozoic
Strata in the Northern Area of Mizu.
nami-machi. Gifu Prcfecturl'. japan.
}our. Geol. Soc. Japan, 1'01. 56, pp. -181'i!l:.!; vul. 57, pp. 9!:1-llU.
!·lAUE, T. (l!l5U), Ringiculidae and Retusidae(I) in japan. Ill. Cal. japan. Shells. 110. :!,pp. 7-16.
- - (1950). l'upiJac in Japan. Ibid., uo. (j,pp. 39--1-1.
- - (195~). Atyidae in Japan. Ibid .. uo. :!0,
pp. 137-15~.
- - (195·1). l~eporl on the l\lollusca chietly
collected by the S. S. Soyii-Maru of theImperial Fisheries Experimental Stationon the Continental Shelf bordering.Japan during the Years 19~:.!-l!l:JO. part 1,
Ccphalaspidea. !'ubi. Seto Mar. /Jiol. Lab.
!'Of. 3, 110 ••1, pp. 301-318.
- - (1!155), A List of the CephalaspiJ Opist hobranchia of japan. Bull. /Jiogeogr. Sm·.
.fap:Jn. t•ols. 16-19, pp. 54-79.
IsHII!.-\, S. and others (1954), Cenozoic For·mat ions in Okuyamada. Kyoto Prcf eel ure. jour. Geo/. Soc. japan. 1111/. 60. pp.
160-166.
lTott;Aw.-\, J. (1955a), The Cenowil: Strata
in the lwamura Basin. Gifu l'rcfccturl:.

183

Japan. Ibid., r·ol. 61, pp. 511-517.
- - (1955b), Molluscan Fauna of tlw 1\lizunami (;roup in the Iwamura Uasin.
1\Jem. Col!. Sd .. Uni1•. /\.yolo, ser. JJ. !•ul.
:!:!, pp. 1:?7-143.
- - (1956). l'vlolluscan Fauna of the Tsuzuki Group in Kyoto Prefecture, japan.
Ibid., Pol. 23, no. 2, pp. 179-l!l:.!.
Kui<oDA, T. and liABE, T. (1954). On Somejapanese l'vtollusca Described by A.
AuAMS. whose Specimens arc lkposi kd
in the Redpath 1\luseum of Canada (No.
1).japan.}our. 1llal.. uol. 18, IW./, pp. 1-lli.
NoMUI{A, S. (1939). Noles on soml! Opisllto·
branchiala based upon thl' collo.;clion of
the SAITo Hii-on Kai Museum. chietly
collected from Northern Honshu, japan.
japan. ]our. Geol. Geogr .. l'O!. ]1;, pp. 11-:.!7.
0Go>.E, S. (l!l5~). On the Mode of Occurrence of Fossils in the Mizunami Groupin the Eastern M i no, japan. jour. (:eo/.
Soc. japan, 1'0[. 58, pp. 477--IS!i.
0TUKA. Y. (1!138), Mollusca from thl· 1\liocene of Tyugoku, japan. jour. Fuc. Sci.
Imp. Unit•. Tokyo, sec. II, vv/. :), pt. 2, pp.
:!1-45.
SAKAMOTO. T. (1955).
Cenozoi~.: Formationsin the Southern !Jistrict of Nara City .
.four. Gcol. Soc. fajJllu. !Jot. (j I, pp. li~-72.
TAKEYAMA, T. (1935). Review of the Ringi.
culidae of japan (Fossil anJ Recent).
The Venus, vo/. 5, pp. 69-90.
YAI\IAI<AWA, G. (1911). Description of somefossil Opisthobranchiata from the Dilu.
vial Deposits of Japan. ]our. Geol. Soc.
1iJkyo, r•ol. 18, pp. :19-5:!.
YoKoYAMA. 1\1. (l!l2li), Molluscan Fossils
from the Tertiary of 1\lino. jour. N1c.
Sci. Imp. Univ. Tokyo, ~ec. II, I'OI. 1, pp.
213-:.!27.

i\11 of the type spcdmcns arc preserved in the collection of the Institute of (;eulogy and
1Vlincra1ogy. Kyoto University.

n::.JPROCEEDINGS OF THE Pi\LAEONTOLOGlCAL SOCIETY
OF JAPAN

19!:i71f IJ R 30 II.................................... -rw~J~. ilfi: ~1t.mJ..:,.i'=Jll!''i~tlt.lt!!rr-i2<¥t£:.tn:1f::l. --c r~Milli t... t.:< b~

5. 'E:;~iltiJtrlf!W Cardiidae ............... a~iX!J:lti. ::;.,·;,:j/rdtlf¥11 Oliviuac ..................... -m-JJiiiX!JJ
7. ili!=ll!,t~qm~.,liiF. Lobatammlaria 1.:"0\ ··c. ..

riJ;J,;:-d; 1 l.'qW,}·~::~~.fi6Sf~I~J~J:fi· 13 ~).

f71J~I.:1f~1t;:, ,t/I!Ui! /'i:.Wr.H:,]IIii!<imll (.i;{j\~')ji/i I)"( Ji,o"1. On the phylogend ic problem of the socalled Fusulina . .................. Ken ichi ls!III~- Fusulinids from the upper part of the
Yayamadake Limestone in the Kuma
Massif. .................. Kameloshi J\,\NMEI~A
3. Upper Permian and Triassic l\lyophu·
riidae from the Maizuru Zone, South·
west japan ({~~1~) ......... 1\.eiji NAKAZAWA-1. :!6-¥!J.U'~J.>iRit'1i!E LuciiUt 1:."0\ '"L (i\;.liG)................................. q.!)!fpjjt. ~!li~::.tl'illti. On the New Nymphaeaccan Plants fromtlw Omichidani Beds (Cretaceous Sys·tcm). Jshikawa Prefecture. Innerside ofCentral Japan (1\::,b~) ... Hidckuni l'vfATSUo
9. Occurrence of some Crustacean Foot·
print. Semlaia talszmolmclliensis gen. et
sp. nov. from the Neogene Tertiary ofjapan (R,\-,"G) .................. Rikizo lMAIZUMIITorGAw A:

Cephalaspid Opisthobranchia

Plate 26

3

5

2a

2b

1

9

6

7a

7b

Sb

Sa

14
13

11

10

12

17

15a

16b

15b
16a

20
18

19a

19b

21

CONSTITUTION

of the
PALAEONTOLOGICAL SOCIETY OF JAPAN
Article 1. The Society shall be known as the Palaeontological Society of Japan.
Article 2. The object of the Society is to promote the study and popularization of palaeontology and related sciences.
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Meeting, establish within it research committees.
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Addendum
1) Voting in the Council shall he by unsigned bnllot.
2) This Constitution become effective on February I. 1958.
3) Fellows indicated under Article 8 shall he determined this year (only) by the present Council.

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