「付録 A．光合成細菌の分類名」の更新 （2024 年 4 月）
本書発行以降、門の名称変更を含め多くの分類名の変更が行われ、また新種や新しい分類群の記載も行われている。以下に 2023 年末の時点での光合成細菌（ゲノム上で光合成遺伝子を持つものも含む）の階層分類名（学名）のリストを英文の説明とともに示す。学名は LPSN - List ofProkaryotic names with Standing in Nomenclature (https://lpsn.dsmz.de/) に基づいているが引用符（” ”）の付いているものは未公認名である。
This Appendix lists the taxa of anoxygenic phototrophic bacteria so far described as being culturable. Theselists include bacterial taxa whose photosynthetic functions or chlorophototrophy have been demonstrated onlyby genomic data. The taxonomic names are based on information from the LPSN - List of Prokaryotic nameswith Standing in Nomenclature (https://www.bacterio.net/).
付録 A.1 紅色非硫黄細菌および好気性光合成細菌（Pseudomonadota 門）
Appendix A.1 Taxa of purple non-sulfur bacteria (phylum Pseudomonadota)Class
Alphaproteobacteria

Order
Caulobacterales
Hyphomicrobiales

GenusaFamilyCaulobacteraceae
Maricaulaceae
Afifellaceae
Aurantimonadaceae
Beijerinckiaceae
Blastochloridaceae
Hyphomicrobiaceae
Methylobacteriaceae

Brevundimonas*2
Photocaulis*1
Afifella
Fulvimarina*2
Methylocella*2
Methylocapsa*1
Blastochloris
Hyphomicrobium*2Rhodomicrobium
Methylobacterium*2
Methylorubrum*2
Methylocystis*2
Bradyrhizobium*2
Rhodoplanes

Methylocystaceae
Nitrobacteriaceae

Rhodopseudomonas

Rhizobiaceae“Peteryoungia”*2
Hoeflea*3

Rhodobiaceae
Roseiarcaceae

Rhodobium
Rhodoblastus
Roseiarcus*1
Salinarimonas*1
Roseibium*2
Stappia*2
Cereibacter

Salinarimonadaceae
Stappiaceae
Rhodobacterales

Paracoccaceae

Dinoroseobacter*1
Fuscovulum
Gemmobacter*2
Halovulum*2
Neotabrizicola*1
Phaeovulum“Pseudotabrizicola”*1
Rhodobaca
Rhodobacter*5

1Type specie orphototrophic
representativea
B. subvibrioides
P. sulfatitolerans
A. marina
F. pelagi(M. silvestris)(M. palsarum)
B. viridis(H. zavarzinii)
R. vannielii
M. organophilum
M. extorquens
M. rhodinum(M. rosea)(B. oligotrophicum)
R. roseus
R. azumiensis
R. elegans
R. pokkaliisolii
R. tepidamans
R. palustris
R. pentothenatexigens
R. telluris
R. thermotolerans"P. albertimagni"(H.olei)H. phototrophica
R. orientis
R. acidophilus
R. fermentans
R. rosea
R. denhamense
S. stellulata
C. changlensis
C. azotoformans
C. johrii
C. sphaeroidesD. shibae
F. blasticum(G. aestuarii)
H. dunhuangense
N. shimadae
P. veldkampii“P.formosa”
R. bogoriensis
R. capsulatus
R. calidifons

No. ofAnP
speciesb
12
3
1
1
14
138
6
188

10

1
22
2
1
3
7
1
5

1
1
1
1
12
1
2
15

Rhodobaculum
Roseibaca*1"Roseibacula"*1
Roseicitreum*1
Roseinatronobacter*1
Rubribacterium
Rubrimonas*2
Tabrizicola*2
Jannaschia*2
Litoreibacter*2
Kangsaoukella*1
Mameliella*2
Marivita*2
Nereida*1
Palleronia*2
Planktomarina*1
Planktotalea*2
Pseudaestuariivita*1
Puniceibacterium*2
Rhodovulum

Roseobacteraceae

Roseibacterium*2
Roseicyclus*1
Roseisalinus*1
Roseivivax*2
Roseobacter*2

Rhodospirillales

Roseovarius*2
Salipiger*2
Shimia*2
Sulfitobacter*2
Yoonia*2
Acidiphilium*2

Acetobacteraceae

Acidisphaera*1
Craurococcus*1
Elioraea*5
Humitalea*1
Lichenicoccus*1
Paracaurococcus*1
Rhodopila
Rhodovastum
Roseococcus*2
Roseomonas*2
Rubritepida*1
Rhodocista
Niveispirillum*2
Skermanella*2
Geminicoccus*2
Aerophototrophica*1
Caenispirillum*2
Magnetorpirillum*3

Azospirillaceae
Geminicoccaceae
RhodospirillaceaePararhodospirillum
Phaeovibrio
Rhodospira
Rhodospirillum"Roseospirillum"
Roseospira
Rhodovibrio
Oceanibaculum*2
Thalassobaculum*2
Rhodothalassium"Citrimicrobium"*1
Erythrobacter*1"
Rhodovibrionaceae
Thalassobaculaceae
Rhodothalassiales
Sphingomonadales

Rhodothalassiaceae
Erythrobacteraceae

Sphingomonadaceae

2

Aurantiacibacter*2
Novosphingobium*2
Blastomonas*1
Chioneia*1

R. claviforme
R. ekhonensis"R. alcaliphilum"
R. antarcticum
R. thiooxidans
R. polymorphum
R. cliftonensis
T, aquatica(J. cystaugens)(L. roseus)
K. pontilimi
M. albaM. cryptomonadis
N. ignava(P. rufa)
P. temperata
P. frisia
P. atlantica
P. atlanticum
R. sulfidophilum
R. iodosum
R. robiginosum
R. strictum
R. elongatum
R. mahoneyensis
R. antarcticus
R. halodurans
R. litoralis
R. denitrificans
R. tolerans
S. mucosus(S. ponticola)(S. noctilucicola)
Y. vestfoldensis
A. cryptum
A. acidophilum
A. multivorum
A. rubrum
A. rubrifaciens
C. roseus
E. tepidiphila
E. tepida
H. rosea
L. roseus
P. ruberR. globiformis
R. atsumiense
R. thiosulfatophilus(R. aestuarii)
R. flocculans
R. centenaria(N. cyanobacteriorum)(S. aerolata)
G. roseus
A. crusticola(C. salinarum)(M. fulvum)
M. molischianum
P. photometricum
P. sulfidophilus
R. trueperi
R. rubrum"R. parvum"
R. mediosalina
R. salinarum
O. indicum(T. salexigens)
R. salexigens"C. luteum"
E. longus
E. tepitarius(A. marinus)(N. subterraneum)
B. natatoria
C. frigida1211211382
11214112
12032
1333112251131111125122211153111152111182264

Sphingomonas*2"Sandaracinobacter"*2
Sandarakinorhabdus*1
Polymorphobacter*2
Polynucleobacter*2
Piscinibacter*2
Limnohabitans*2
Rhodoferax*3

Sphingosinicellaceae
Betaproteobacteria

Gammaproteobacteria

Burkholderiales

Burkholderiaceae
Comamonadaceae

Nitrosomonadales
Rhodocyclales

Not assigned to family
Sterolibacteriaceae
Rhodocyclaceae

Cellvibrionales

Halieaceae

Roseateles*2
Rubrivivax*4
Aquincola*2
Methyloversatilis*2
Rhodocyclus
Chromatocurvus*1Congregibacter*1
Luminophilus*1
Pseudohaliea*1(S. koreensis)"S. sibiricus"
S. limnophila
P. multimanifer
P. necessarius(P. sakaiensis)(L. planktonicus)
R. fermentans
R. antarcticus
R. depolymeransR. gelatinosus
A. tertiaricarbonis(M. universalis)
R. purpureus
R. tenuis
C. halotolerans
C. litoralis
L. syltensis
P. rubraa61445112231131111

Species given in parentheses are not the type but a phototrophic representative.
基準種が光合成細菌でない場合は光合成細菌の代表種を括弧内に示す。また、それ以外に本書に出ている種があれば 2 行目以下に記載した。
b
Number of phototrophic species validly published under the ICNP.
ICNP に正式名として記載されている光合成細菌の種数*1 Genera comprising AAnPB (aerobic anoxygenic phototrophic bacteria) species only
AAnP(好気性光合成)細菌のみが知られている属*2 Genera comprising AAnPB and chemotrophic species
AAnP 細菌と化学栄養細菌が混在する属
3* Genera comprising PNSB (purple nonsulfur bacteria) and chemotrophic species
紅色非硫黄細菌と化学栄養細菌が混在する属*4 Genera comprising PNSB, AAnPB, and chemotrophic species
紅色非硫黄細菌、AAnP 細菌と化学栄養細菌が混在する属*5 Genus comprising PNSB and AAnPB species
紅色非硫黄細菌と AAnP 細菌が混在する属

付録 A.2 紅色硫黄細菌（Pseudomonadota 門）
Appendix A.2 Taxa of purple sulfur bacteria (phylum Pseudomonadota)
ClassOrder

Gammaproteobacteria

Chromatiales

FamilyChromatiaceae

Genus
Allochromatium
Caldichromatium
Chromatium
Halochromatium
Imhoffiella
Isochromatium
Lamprobacter
Lamprocystis
Marichromatium
Pfennigia
Phaeobacterium
Phaeochromatium
Rhabdochromatium
Thermochromatium
Thioalkalicoccus
Thiobaca
Thiocapsa
Thiococcus
Thiocystis
Thiodictyon"Thioflavicoccus"
Thiohalocapsa
Thiolamprovum

3Type specie orphototrophic
representaive
A. vinosumC. japonicum
C. okenii
H. salexigens
I. purpurea
I. buderiL. modestohalophilus
L. roseopersicina
M. gracile
P. purpurea
P. nitratireducens
P. fluminis
R. marinum
T. tepidum
T. limnaeus
T. trueperi
T. roseopersicina
T. pfennigii
T. violacea
T. elegans"T. mobilis"
T. halophila
T. pedioforme

No. ofSpeciesa
7
12
3
2
1
12
4
1
1
1
11115162
121

Thiopedia
Thiophaeococcus
Thiorhodococcus
ThiorhodospiraThiorhodovibrio
Thiospirillum
Ectothiorhodosinus
EctothiorhodospiraHalorhodospira

EctothiorhodospiraceaeaHalorhodospiraceae

T. rosea
T. mangrovi
T. minorT. sibirica
T. winogradskyi
T. jenense
E. mongolicus
E. mobilis
H. halophila

126131184

Number of phototrophic species described phenotypically and/or genotypically and validly published.
ICNP に正式名として記載されている光合成細菌と光合成遺伝子をもつ種数

付録 A.3 緑色硫黄細菌（Chlorobiota 門）、クロロフレクサス（Chloroflexota 門）、ヘリオバクテリア（Bacillota 門）、クロラシドバクテリア（Acidobacteriota 門）、ゲマチモナス（Gemmatimonadota門）、エレミオバクテリア（"Candidatus Eremibacterota"門）、Appendix A.3 Taxa of green sulfur bacteria (phylum Chlorobiota), chloroflexus and relatives (phylumChloroflexota), heliobacteria (phylum Bacillota), chloracidobacteria (phylum Acidobacteriota), gemmatimons(phylum Gemmatimonadota), eremiobacteria (phylum "Candidatus Eremibacterota")
Phylum
Class

Order

FamilyChlorobiota
Chlorobiia

Chlorobiales

Chlorobiaceae

Chloroblexota
Chloroflexia

Chloroflexales

Chloroflexaceae

Genus

Type specie orphototrophic
representaivea
A. perfilievii
C. tepidum
C. limicola
C. thalassium"C. vacuolata"
P. clathratiforme
P. aestuarii
C. aurantiacusC. giganteum
O. chrysea
H. oregonensis
R. castenholzii“K. aurantiaca"“C. asiatica”“V. halotolerans”"P. methaneseepsis"
H. chlorum
H. gestii
H. fasciatum
H. daurensis"C. thermophilum"

No. ofspeciesb"Thermofontia"
Bacillota
Clostridia"Phototrophicales"
Eubacteriales

Not assigned to family
Not assigned to family
Not assigned to family"Phototrophicaceae"
Heliobacteriaceae

Acidobacterota
Terriglobia
Gemmatimonadota
Gemmatimonadetes"Candidatus
Eremiobacterota"
Ca. Eremiobacteria

Terriglobales

Acidobacteriaceae

Ancalochloris
Chlorobaculum
Chlorobium
Chloroherpeton"Chloroplana"
Pelodictyon
Prosthechochloris
Chloroflexus
Chloronema
Oscillochloris
Heliothrix
Roseiflexus“Kouleothrix”“Chloroploca”“Viridilinea”"Phototrophicus"
Heliobacterium
Heliomicrobium
Heliophilum
Heliorestis"Chloracidobacterium"

Gemmatimonadales

Gemmatimonadaceae

Gemmatimonas(G. phototrophica)2"Candidatus
Baltobacterales""Candidatus
Baltobacteraceae"

Vulcanimicrobium

V. alpinum

1

Oscillochloridaceae
Roseiflexaceae

a125113331211122123133

Species given in parentheses are not the type but a phototrophic representative.
基準種が光合成細菌でない場合は光合成細菌の代表種を括弧内に示す。また、それ以外に本書に出ている種があれば 2 行目以下に記載した。
b
Number of phototrophic species described phenotypically and/or genotypically and validly published.
ICNP に正式名として記載されている光合成細菌と光合成遺伝子をもつ種数4