Annotated checklist of fishesof Yonaguni-jima island,
the westernmost island in Japan
Keita Koeda, Yusuke Hibino, Tomohiro Yoshida, Yuki Kimura, Ryohei Miki, Taiga Kunishima,
Daichi Sasaki, Takanao Furukawa, Momoko Sakurai, Keisuke Eguchi, Hiromu Suzuki,
Tomoki Inaba, Takuya Uejo, Saki Tanaka, Misaki Fujisawa, Hidetoshi Wada, and Teppei Uchiyama

The Kagoshima University Museum

Copy Right © 2016 by the Kagoshima University Museum, Kagoshima
All rights reserved. No part of this publication may be reproduced or transmitted in any form or by any means without
prior written permission from the publisher. Copyrights of the photographs are held by the photographers.
For bibliographic purposes this book should be cited as follows:
Keita Koeda, Yusuke Hibino, Tomohiro Yoshida, Yuki Kimura, Ryohei Miki, Taiga Kunishima, Daichi Sasaki,
Takanao Furukawa, Momoko Sakurai, Keisuke Eguchi, Hiromu Suzuki, Tomoki Inaba, Takuya Uejo, Saki Tanaka,
Misaki Fujisawa, Hidetoshi Wada, and Teppei Uchiyama (2016) Annotated checklist of fishes of Yonaguni-jima island,
the westernmost island in Japan. The Kagoshima University Museum, Kagoshima. vi + 120.
ISBN 978-4-905464-07-5

DTP by Keita Koeda, the Kagoshima University MuseumChecklist of fishes of Yonaguni-jima island<Contents>
Introduction...........................................................................1
Materials and methods..........................................................2
Cheklist
Family Hemiscylliidae テンジクザメ科 ................................4
Family Rhincodontidae ジンベエザメ科 ...............................4
Family Lamnidae ネズミザメ科 ............................................4
Family Carcharhinidae メジロザメ科 ....................................4
Family Sphyrnidae シュモクザメ科 ......................................5
Family Pristiophoridae ノコギリザメ科 ................................5
Family Dasyatidae アカエイ科 ..............................................5
Family Myliobatidae トビエイ科 ...........................................5
Family Albulidae ソトイワシ科 ............................................6
Family Megalopidae イセゴイ科 ...........................................6
Family Anguillidae ウナギ科 .................................................6
Family Muraenidae ウツボ科 .................................................6
Family Synaphobranchidae ホラアナゴ科 ............................9
Family Ophichthidae ウミヘビ科 ..........................................9
Family Congridae アナゴ科 ...................................................9
Family Clupeidae ニシン科 ..................................................10
Family Engraulidae カタクチイワシ科 ...............................10
Family Chanidae サバヒー科 ...............................................10
Family Cyprinidae コイ科 ....................................................10
Family Cobitidae ドジョウ科 ...............................................11
Family Aulopidae ヒメ科 .....................................................11
Family Synodontidae エソ科 ................................................11
Family Polymixiidae ギンメダイ科 .....................................12
Family Antennariidae カエルアンコウ科 ............................12
Family Chaunacidae フサアンコウ科 .................................12
Family Mugilidae ボラ科 .....................................................12
Family Atherionidae ムギイワシ科 .....................................14
Family Exocoetidae トビウオ科 ..........................................14
Family Hemiramphidae サヨリ科 ........................................14
Family Belonidae ダツ科 ......................................................14
Family Poeciliidae カダヤシ科 ............................................15
Family Holocentridae イットウダイ科 ...............................15
Family Pegasidae ウミテング科 ..........................................18
Family Solenostomidae カミソリウオ科 ............................18
Family Syngnathidae ヨウジウオ科 ....................................18
Family Aulostomidae ヘラヤガラ科 ....................................19
Family Fistulariidae ヤガラ科 ..............................................19
Family Centriscidae ヘコアユ科 ..........................................19
Family Scorpaenidae フサカサゴ科 ....................................20
Family Tetrarogidae ハオコゼ科 ..........................................23
Family Synanceiidae オニオコゼ科 .....................................23
Family Platycephalidae コチ科 .............................................23
Family Ambassidae タカサゴイシモチ科 ...........................24
Family Serranidae ハタ科 .....................................................25
Family Pseudochromidae メギス科 ......................................30
Family Plesiopidae タナバタウオ科 ....................................31
Family Opistognathidae アゴアマダイ科 ............................31
Family Priacanthidae キントキダイ科 ................................31
Family Apogonidae テンジクダイ科 ...................................31
Family Malacanthidae キツネアマダイ科 ...........................35
Family Coryphaenidae シイラ科 ..........................................36
Family Rachycentridae スギ科 .............................................36
Family Echeneidae コバンザメ科 ........................................36
Family Carangidae アジ科 ....................................................36

Family Bramidae シマガツオ科 ..........................................39
Family Lutjanidae フエダイ科 .............................................39
Family Caesionidae タカサゴ科 ..........................................43
Family Gerreidae クロサギ科 ..............................................44
Family Haemulidae イサキ科 ...............................................44
Family Nemipteridae イトヨリダイ科 ................................45
Family Sparidae タイ科 .......................................................46
Family Lethrinidae フエフキダイ科 ....................................46
Family Mullidae ヒメジ科 ....................................................48
Family Pempheridae ハタンポ科 .........................................50
Family Monodactylidae ヒメツバメウオ科 ........................51
Family Kyphosidae イスズミ科 ...........................................51
Family Chaetodontidae チョウチョウウオ科 .....................52
Family Pomacanthidae キンチャクダイ科 ..........................59
Family Pentacerotidae カワビシャ科 ..................................61
Family Teraponidae シマイサキ科 ......................................61
Family Kuhliidae ユゴイ科 ..................................................61
Family Oplegnathidae イシダイ科 .......................................62
Family Cirrhitidae ゴンべ科 .................................................63
Family Cichlidae カワスズメ科 ...........................................64
Family Pomacentridae スズメダイ科 ..................................64
Family Polynemidae ツバメコノシロ科 .............................72
Family Labridae ベラ科 ........................................................72
Family Scaridae ブダイ科 ....................................................80
Family Pinguipedidae トラギス科 .......................................82
Family Tripterygiidae ヘビギンポ科 ...................................83
Family Trichonotidae ベラギンポ科 ....................................84
Family Blenniidae イソギンポ科 .........................................84
Family Callionymidae ネズッポ科 ......................................86
Family Eleotridae カワアナゴ科 ..........................................87
Family Gobiidae ハゼ科 .......................................................88
Family Ptereleotridae クロユリハゼ科 ................................96
Family Ephippidae マンジュウダイ科 ................................97
Family Scatophagidae クロホシマンジュウダイ科 ..........98
Family Siganidae アイゴ科 ..................................................98
Family Zanclidae ツノダシ科 ..............................................99
Family Acanthuridae ニザダイ科 .........................................99
Family Sphyraenidae カマス科 ..........................................105
Family Gempylidae クロタチカマス科 ............................106
Family Scombridae サバ科 .................................................106
Family Istiophoridae マカジキ科 .......................................107
Family Nomeidae エボシダイ科 ........................................107
Family Ariommatidae オオメメダイ科 .............................108
Family Caproidae ヒシダイ科 ............................................108
Family Bothidae ダルマガレイ科 ......................................108
Family Soleidae ササウシノシタ科 ..................................108
Family Balistidae モンガラカワハギ科 ............................109
Family Monacanthidae カワハギ科 ....................................113
Family Ostraciidae ハコフグ科 ..........................................113
Family Triodontidae ウチワフグ科 ....................................113
Family Tetraodontidae フグ科 ............................................114
Family Diodontidae ハリセンボン科 ................................115
Discussion............................................................................116
Acknowledgments................................................................118
References............................................................................118

V

Checklist of fishes of Yonaguni-jima island

Annotated checklist of fishes of Yonaguni-jima island,
the westernmost island in Japan
Keita Koeda1*, Yusuke Hibino2, Tomohiro Yoshida1, Yuki Kimura3, Ryohei Miki4, Taiga Kunishima5,
Daichi Sasaki2, Takanao Furukawa6, Momoko Sakurai5, Keisuke Eguchi1, Hiromu Suzuki2,
Tomoki Inaba1, Takuya Uejo1, Saki Tanaka3, Misaki Fujisawa3, Hidetoshi Wada7, and Teppei Uchiyama8
1

The Kagoshima University Museum, 1–21–30 Korimoto, Kagoshima 890–0065, Japan
Fisheries Research Laboratory, Mie University, 4190–172 Wagu, Shima, Mie 517–0703, Japan
3
Laboratory of Biology of Aquatic Resources, Hiroshima University, 1–4–4 Kagamiyama, Higashi-Hiroshima 739–0046, Japan
4
Nobeoka Marine Science Station, Field Science Center, University of Miyazaki, 376–6 Akamizu, Nobeoka, Miyazaki 889–0517, Japan
5
Graduate School of Engineering and Science, University of the Ryukyus, 1 Senbaru, Nishihara, Okinawa 903–0213, Japan
6
Graduate School of Bioresources, Mie University, 1577 Kurima-machiya, Tsu, Mie 514–8507, Japan
7
Division of Fisheries Sciences, University of Miyazazki, 1–1 Gakuen-kibanadai-nishi, Miyazaki 889–2192, Japan
8
Diving Service MARLIN, 4652–1, Yonaguni, Yonaguni, Okinawa 907–1801, Japan*Corresponding author: e-mail: hatampo@gmail.com
2

Abstract An annotated checklist of fishes of Yonaguni-jima island, Okinawa Prefecture, southern Japan, was compiled mainly from a fieldsurvey which was carried out at September 2015 by the authors belonging to five universities, as well as from results from two previous smallscale surveys. Additionally, all registered specimens previously recorded from Yonaguni-jima island were re-examined as much as possible. A
total of 603 fish species (104 families) are listed based on the collected specimens during our expedition, museum specimens, literatures andunderwater photographs. Each species in the list is stated with citations of literature, registration numbers, sizes, localities around the island,collection methods, collected dates, nomenclatural and taxonomic remarks, and color photographs if available. 334 species (39 families)
represent the first reliable records from Yonaguni-jima island on the basis of the collected specimens. The zoogeographical implications of
Yonaguni-jima island ichthyofauna are discussed.
Key words: ichthyofauna, Okinawa, Yaeyama Islands, Ryukyu Archipelago, new record, Kuroshio Current.

Introduction
In 1927, Drs. David S. Jordan and Shigeho Tanaka reported four freshwater fish species, Carassius sp., Chanos chanos(Forsskål, 1775), Eleotris fusca (Bloch and Schneider, 1801),
and Kuhlia rupestris (Lacepède, 1802), from Yonaguni-jimaisland on the basis of specimens obtained by Mr. H. Kuroiwa(Jordan and Tanaka, 1927). These represent the oldest available fish specimens collected from Yonaguni-jima island. After
Jordan and Tanaka (1927), several pieces of literature reported
the same four species and Misgurnus anguillicaudatus (Cantor, 1842) without deposited specimens (Okinawa Prefecture
Experimental Fisheries Station, 1934; Ogawa, 1937; Aoyagi,
1948; Shibata, 1964; Gushiken, 1972; Nakamura, 1974).
No comprehensive reports on fishes from Yonaguni-jimaisland were published until the 21st-century. Yoshigou (2000)
is the first report that focused on the freshwater and brackish
ichthyofauna of this island. 63 species of 26 families werereported based on specimens collected in three days in August
1999 by the author. These specimens are currently deposited
at Hiwa Museum for Natural History (HMNH). Subsequently,
Yoshigou et al. (2001) and Yoshigou and Nakamura (2003)
provided the specimen list of HMNH, and the specimens of 35
additional species collected at the tide pools of Yonaguni-jimaisland were included in this list. However, no comprehensive
reports including marine water fishes from Yonaguni-jimaisland have been published up to the present date, and no integrated surveys have been conducted on or around the island.
To clarify the importance and interesting zoogeographicalfeatures of Yonaguni-jima island, all authors of this study tothe exception of the last author T. Uchiyama (who provided anumber of underwater photographs) conducted a large-scale
expedition at Yonaguni-jima island in 17–26 September 2015
(Fig. 1). In the main survey and two additional small-scale
surveys, one by K. Koeda (23–25 July 2013) and one by D.
Sasaki (10–16 March 2015), 1,120 specimens of 399 species(211 genera, 72 families) of marine, brackish and freshwater
fishes were collected, with 334 representing the first records
from the island based on the collected specimens.
This paper provides a list of 603 species of marine, brackish and freshwater fishes (306 genera and 104 families) occurring around Yonaguni-jima island on the basis of collected
specimens along with color photographs where available,
museum specimens, published literatures, and underwaterphotographs. Specimens collected from Yonaguni-jima island
deposited at HMNH and National Museum of Nature and Science, Tsukuba (NSMT) were re-identified during this study.
Finally, the zoogeographical implications of Yonaguni-jimaisland ichthyofauna are discussed.
Yonaguni-jima island
Yonaguni-jima island is located at 24°27′N, 122°57′E (Fig.
2A), ca. 70 km and ca. 120 km west from Iriomote-jima (Fig.
2B) and Ishigaki-jima (Fig. 2C) islands, respectively. This
small island belongs to the Yaeyama Islands, and is the westernmost island in Japan. The distance between Yonaguni-jimaisland and Taiwan is only ca. 110 km, and the mountains of1

K. Koeda et al.

Fig. 1. Photographs from ichthyofaunal expedition in Yonaguni-jima island. A and B, SCUBA Diving; C, line-fishing from coast; D, line-fishing from boat; E,
hand net at intertidal zone; F, hand nets at freshwater river; G, cast net at mangroves estuary; H, gill net at brackish water.

Taiwan are sometimes visible from westernmost Cape Irizaki
on Yonaguni-jima island. The shape of island is elliptical,
with an area of ca. 28 km2 and 28 km of coastline (Fujimoto,1972). Most of the coast-line of the island consists of sheer
rocky cliffs. Thus, in-reef environments such as shallow
waters, coral reefs, and sandy beaches are extremely limited(Fig. 3). Hikawa Bay on the southern side of the island (Fig.
3, number 23; Table 1) is the only large in-reef environmentof the island. Two fishing ports, Kubura and Sonai, located on
the north coast, compose inner bay environments with muddy
bottoms. Two small rivers, the Tabaru and Kubura rivers, run
into the north coast. Tabaru River starts from a spring, ca. 1.5
km upstream from the mouth, and consists of ca. 600 m offreshwater and mangroves estuary (Yoshigou, 2000). Kubura
River starts from a brackish lake, Kubura-mit, which is located ca. 300 m from the mouth of the river. Although such monotonous environments in coastal areas of an island generally
result in a decrease in fish species diversity, Yonaguni-jimaisland has a relatively high diversity as the island faces thewarm and strong Kuroshio Current, which brings tropical fishes from the south.
The Kuroshio Current flows from east of the Philippines
to the Pacific coast of southern Japan, via the channel between
Taiwan and Yonaguni-jima island [Fig. 1 in Motomura et al.
(2010)]. Yonaguni-jima island is sometimes surrounded by the
Kuroshio Current, and thus the current of the sea can become
furiously strong, and the sea conditions around the island areoften unsettled. For this reason, Yonaguni-jima island wascalled “Donan” in traditional Okinawan dialect, which means“difficult to pass to the island”.
Materials and methods
Fishes were collected by using SCUBA diving or snorkeling with hand nets and spears, line-fishing from coast or
boats, and gill nets, cast nets, fishing lights, and tube traps inshallow waters. Only a handful of specimens were purchased
at the fish landing port at Yonaguni-jima island. Curatorial
procedures for collected specimens followed Motomura andIshikawa (2013).

2

The systematic arrangement of families generally follows
Nelson (2006). Scientific names and taxonomical attributions
generally follow Nakabo (2013) with some modifications byrecent published or unpublished taxonomic studies. Authorship and published year follow Eschmeyer et al. (2016). Species in families are arranged in alphabetical order by species
name. Standard Japanese names generally follow Nakabo(2013), and are arranged in Japanese “katakana”. Local names
in Yonaguni-jima island generally follow Ikema (2003), andare arranged in Japanese “hiragana” if available. Each species
record in the list was compiled from voucher specimens andliterature sources. Each voucher specimen in the list includes
registration number, number of specimens registered if morethan one, standard length [abbreviated as SL; total length(TL) are used in the order Anguilliformes], sex, locality on
Yonaguni-jima island (Fig. 3, Table 1), collection method,and collection date. The list of specimens used in previous

Fig. 2. Map of Ryukyu Archipelago and adjacent islands, Japan. Charactersin the map show major islands mentioned in the text: A, Yonaguni-jimaisland; B, Iriomote-jima island; C, Ishigaki-jima island; D, Miyako group;
E, Tokashiki-jima island; F, Ie-jima island; G, Yoron-jima island; H, Yakushima island; I, Iou-jima and Take-shima islands.

Checklist of fishes of Yonaguni-jima island

Fig. 3. Map of Yonaguni-jima island. Numbers in the map show major collection localities at ichthyofaunal expedition. Name, lat/long, depth, andcollection methods in each localities are shown at Table 1. Yellow, land; dark green check, forest; orange stripe, town area; light green stripe, inreef habitat.
Table 1. Name of point, lat/long, maximum depth, and collection methods at major collection localities at ichthyofaunal expedition in Yonagunijima island. Map numbers correspond with the numbers on Fig. 3.

Map no. Point
1
Off Sonai
2
Sonai channel
3
Sonai Fishing Port
4
Tabaru River
5
Mouth of Tabaru River
6
Off Yonaguni Airport
7
Umabana8
Off Northern Pasture9
Danno Beach
10
Danno-dropoff
11
Kubura-bari
12
Mouth of Kubura River
13
Kubura Fishing Port
14
Levee at Kubura
15
Na-ma Beach
16
Kubura Hinan Port
17
Off Kubura Hinan Port
18
Anchor's graves
19
Rakuda Rock
20
Southern Pasture
21
Kataburu Beach
22
Hikawa23
Off Hikawa
24
Arakawa Bay

Latitude24°26ʹ07ʺN
24°28ʹ18ʺN
24°28ʹ18ʺN
24°27ʹ39ʺN
24°28ʹ11ʺN
24°28ʹ32ʺN
24°28ʹ21ʺN
24°28ʹ26ʺN
24°27ʹ55ʺN
24°27ʹ56ʺN
24°28ʹ26ʺN
24°27ʹ06ʺN
24°27ʹ05ʺN
24°27ʹ09ʺN
24°26ʹ52ʺN
24°26ʹ44ʺN
24°26ʹ35ʺN
24°26ʹ29ʺN
24°26ʹ27ʺN
24°26ʹ14ʺN
24°26ʹ18ʺN
24°26ʹ28ʺN
24°26ʹ00ʺN
24°26ʹ15ʺN

Longitude
Environment
Max. depth Methods
123°00ʹ25ʺE
Marine55 m
Lf
123°00ʹ08ʺE
Marine5m
Sd, Hn, Lf, Gn
123°00ʹ08ʺE
Marine10 m
Lf, Cn, Fl, Tt
123°00ʹ17ʺE Brackish, freshwater
1m
Hn, Cn122°59ʹ57ʺE Brackish, freshwater
1m
Hn, Cn, Gn
122°59ʹ22ʺE
Marine55 m
Lf
122°57ʹ49ʺE
Marine26 m
Sd, Lf122°57ʹ36ʺE
Marine55 m
Lf
122°57ʹ16ʺE
Marine5m
Sd, Hn122°57ʹ04ʺE
Marine42 m
Sd, Lf123°00ʹ16ʺE
Marine10 m
Lf
122°56ʹ35ʺE Brackish, freshwater
1m
Hn, Cn, Gn
122°56ʹ34ʺE
Marine3m
Sd, Hn, Lf, Cn, Gn, Fl, Tt
122°56ʹ12ʺE
Marine10 m
Lf
122°56ʹ23ʺE
Marine1m
Cn
122°56ʹ20ʺE
Marine3m
Lf, Tt122°56ʹ22ʺE
Marine30 m
Sd, Lf122°56ʹ51ʺE
Marine28 m
Sd, Lf122°57ʹ02ʺE
Marine30 m
Sd, Lf122°58ʹ17ʺE
Marine10 m
Hn, Lf122°58ʹ28ʺE
Marine10 m
Hn, Lf122°59ʹ06ʺE
Marine4m
Sd, Hn, Cn
123°00ʹ25ʺE
Marine50 m
Lf
123°00ʹ33ʺE
Marine12 m
Sd, LfSd, SCUBA diving; Hn, hand net; Lf, line fishing; Cn, cast net; Gn, gill net; Fl, fishing light; Tt, tube trap.

literatures (Yoshigou, 2000; Yoshigou et al., 2001; Yoshigou
and Nakamura, 2003) were re-identified in this study, andinformation on SL and locality of the re-examined specimens
are provided.
Specimens collected during the main survey at Yonaguni-jima island in 2015 and two previous small surveys byauthors are deposited at the following institutions: Fisheries
Research Laboratory, Mie University (FRLM); the Kagoshima
University Museum (KAUM); Division of Fisheries Sciences,
Faculty of Agriculture, University of Miyazaki, Miyazaki Uni-

versity (MUFS); Department of Marine Sciences, Universityof the Ryukyus (URM: currently deposited at the Okinawa
Churashima Foundation).

3

K. Koeda et al.
Checklist
ORDER ORECTOLOBIFORMES
テンジクザメ目
FAMILY HEMISCYLLIIDAE
テンジクザメ科
Chiloscyllium punctatum Müller and Henle, 1838
イヌザメ [Marine] (Fig. 4)

ORDER LAMNIFORMES
ネズミザメ目FAMILY LAMNIDAE
ネズミザメ科
Carcharodon carcharias (Linnaeus, 1758)
ホホジロザメ [Marine] (Fig. 6)

Fig. 6. Carcharodon carcharias ホホジロザメ (Diving Service MARLIN).
Fig. 4. Chiloscyllium punctatum イヌザメ (Diving Service MARLIN).

FAMILY RHINCODONTIDAE
ジンベエザメ科
Rhincodon typus Smith, 1828
ジンベエザメ [Marine] (Fig. 5)

ORDER CARCHARHINIFORMES
メジロザメ目FAMILY CARCHARHINIDAE
メジロザメ科
Carcharhinus brevipinna (Müller and Henle, 1839)
ハナザメ [Marine] (Fig. 7)

Fig. 7. Carcharhinus brevipinna ハナザメ (Diving Service MARLIN).

Carcharhinus falciformis (Bibron, 1839)
クロトガリザメ [Marine]
Shimose et al. (2008): stomach content of Istiompax indica.
Triaenodon obesus (Rüppell, 1837)
ネムリブカ [Marine] (Fig. 8)

Fig. 5. Rhincodon typus ジンベエザメ (Diving Service MARLIN).

Fig. 8. Triaenodon obesus ネムリブカ (Diving Service MARLIN).

4

Checklist of fishes of Yonaguni-jima island
FAMILY SPHYRNIDAE
シュモクザメ科
Sphyrna lewini (Griffith and Smith, 1834)
アカシュモクザメ [Marine] (Fig. 9)
かしさば
Ikema (2003).

Taeniurops meyeni (Müller and Henle, 1841)
マダラエイ [Marine] (Fig. 11)

Dasyatidae sp.
アカエイ科の一種 [Marine] (Fig. 12)

Fig. 11. Taeniurops meyeni マダラエイ (Diving Service MARLIN).

Fig. 9. Sphyrna lewini アカシュモクザメ (Diving Service MARLIN).

ORDER PRISTIOPHORIFORMES
ノコギリザメ目
FAMILY PRISTIOPHORIDAE
ノコギリザメ科
Pristiophorus japonicus Günther, 1870
ノコギリザメ [Marine]
Okada and Isa (1959).

FAMILY MYLIOBATIDAE
トビエイ科
Manta birostris (Walbaum, 1792)
オニイトマキエイ [Marine] (Fig. 12)
かまんた
Ikema (2003); Nakabo (2013).

ORDER MYLIOBATIFORMES
トビエイ目
FAMILY DASYATIDAE
アカエイ科
Neotrygon kuhlii (Müller and Henle, 1841)
ヤッコエイ [Marine] (Fig. 10)
Fig. 12. Manta birostris オニイトマキエイ (Diving Service MARLIN).

Fig. 10. Neotrygon kuhlii ヤッコエイ (Diving Service MARLIN).

5

K. Koeda et al.
ORDER ALBULIFORMES
ソトイワシ目FAMILY ALBULIDAE
ソトイワシ科
Albula sp.
ソトイワシ属の一種 [Marine, brackish, freshwater] (Fig. 13)
FRLM 51436, 53.7 mm SL, Kubura Fishing Port, fishinglight, 20 Sept. 2015.

FAMILY MURAENIDAE
ウツボ科
あがうど
Echidna nebulosa (Ahl, 1789)
クモウツボ [Marine] (Fig. 16)
FRLM 51468, 415.0 mm TL, FRLM 52078, 273.5 mm
TL, Kubura Hinan Port, tube trap, 21 Sept. 2015; FRLM
51575, 118.8 mm TL, mouth of Tabaru River, hand net, 24
Sept. 2015; KAUM–I. 80382, 319.2 mm TL, Kubura Hinan
Port, tube trap, 21 Sept. 2015.

Fig. 13. Albula sp. ソトイワシ属の一種 (FRLM 51436, 53.7 mm SL).

FAMILY MEGALOPIDAE
イセゴイ科
Megalops cyprinoides (Broussonet, 1782)
イセゴイ [Marine, brackish, freshwater]
Yoshigou (2000).

ORDER ANGUILLIFORMES
ウナギ目
FAMILY ANGUILLIDAE
ウナギ科
なだ
Anguilla japonica Temminck and Schlegel, 1846
ニホンウナギ [Marine, brackish, freshwater] (Fig. 14)
Fujimoto (1972); Yoshigou (2000); Yoshigou et al. (2001).
HMNH-P53, 323.2 mm TL, Tabaru River, hand net, 9Aug. 1999.
Fig. 16. Echidna nebulosa ク モ ウ ツ ボ (upper: FRLM 51468, 415.0 mm
TL; lower: KAUM–I. 80382, 319.2 mm TL, head).

Gymnomuraena zebra (Shaw, 1797)
ゼブラウツボ [Marine]
Ito (2009).

Gymnothorax breedeni (McCosker and Randall, 1977)
ウツボ属の一種 [Marine] (Fig. 17)
Fig. 14. Anguilla japonica ニホンウナギ (HMNH-P53, 323.2 mm TL)

Anguilla marmorata Quoy and Gaimard, 1824
オオウナギ [Brackish, freshwater] (Fig. 15)
Fujimoto (1972); Yoshigou (2000); Yoshigou et al. (2001).
HMNH-P50, 205.2 mm TL, HMNH-P52, 85.1 mm TL,
Tabaru River, hand net, 9 Aug. 1999.

Fig. 15. Anguilla marmorata オオウナギ (HMNH-P52, 205.2 mm TL).

6

Fig. 17. Gymnothorax breedeni (Diving Service MARLIN).

Checklist of fishes of Yonaguni-jima island
Gymnothorax chlorostigma (Kaup, 1856)
ハナビラウツボ [Marine] (Fig. 18)

Fig. 18. Gymnothorax chlorostigma ハ ナ ビ ラ ウ ツ ボ (Diving Service
MARLIN).

Gymnothorax flavimarginatus (Rüppell, 1830)
ゴマウツボ [Marine] (Fig. 19)
KAUM–I. 78607, 1087.0 mm TL, Kubura Fishing Port,line-fishing, 21 Sept. 2015; NSMT-P 55476, 2 specimens,
217.5–290.5 mm TL, Agarizaki, hand net, 28 Mar. 1997.

Gymnothorax isingteena (Richardson, 1845)
ニセゴイシウツボ [Marine] (Fig. 20)
Ito (2009).

Fig. 20. Gymnothorax isingteena ニ セ ゴ イ シ ウ ツ ボ (Diving Service
MARLIN).

Gymnothorax javanicus (Bleeker, 1859)
ドクウツボ [Marine] (Fig. 21)
FRLM 52029, 892.0 mm TL, Kubura Fishing Port, linefishing, 20 Sept. 2015.

Fig. 21. Gymnothorax javanicus ドクウツボ (FRLM 52029, 892.0 mm TL).

Fig. 19. Gymnothorax flavimarginatus ゴ マ ウ ツ ボ (KAUM–I. 78607,
1087.0 mm TL).

Gymnothorax intesi (Fourmanoir and Rivaton, 1979)
ミナミミゾレウツボ [Marine]
NSMT-P 71449, 618.0 mm TL, southeast of Yonagunijima island, 13 July 2005; NSMT-P 76158, 558.0 mm TL,
southeast of Yonaguni-jima island, 3 July 2006.

Gymnothorax pictus (Ahl, 1789)
アセウツボ [Marine]
Ito (2009).

Gymnothorax pseudothyrsoideus (Bleeker, 1853)
アミメウツボ [Marine] (Fig. 22)
KAUM–I. 80218, 825.0 mm TL, Kubura Fishing Port,
tube trap, 17 Sept. 2015.

7

K. Koeda et al.
Gymnothorax thyrsoideus (Richardson, 1845)
サビウツボ [Marine] (Fig. 24)
FRLM 50705, 477.2 mm TL, Kubura Fishing Port, linefishing, 13 Mar. 2015; FRLM 52077, 294.5 mm TL, Kubura
Hinan Port, tube trap, 21 Sept. 2015; KAUM–I. 80381,
459.3 mm TL, Kubura Hinan Port, tube trap, 21 Sept. 2015;
NSMT-P 55475, 3 specimens, 127.3–274.8 mm TL, Agarizaki,
hand net, 28 Mar. 1997.

Fig. 24. Gymnothorax thyrsoideus サ ビ ウ ツ ボ (KAUM–I. 80381, 459.3
mm TL).
Fig. 22. Gymnothorax pseudothyrsoideus アミメウツボ (KAUM–I. 80218,
825.0 mm TL).

Gymnothorax richardsonii (Bleeker, 1852)
モバウツボ [Marine; brackish] (Fig. 23)
FRLM 51574, 409.0 mm TL, mouth of Kubura River,
hand net, 23 Sept. 2015; FRLM 51968, 344.0 mm TL, FRLM
51969, 303.6 mm TL, FRLM 51970, 277.5 mm TL, FRLM
51971, 339.0 mm TL, Kubura Fishing Port, tube trap, 19 Sept.
2015; KAUM–I. 78310, 295.7 mm TL, KAUM–I. 80257,
238.6 mm TL, Kubura Fishing Port, tube trap, 17 Sept. 2015;
MUFS 46879, 331.0 mm TL, Kubura Fishing Port, tube trap,
16 Sept. 2015.

Gymnothorax undulatus (Lacepède, 1803)
ナミウツボ [Marine] (Fig. 25)
FRLM 51573, 826.0 mm TL, Kubura Fishing Port, linefishing, 25 Sept. 2015.

Fig. 25. Gymnothorax undulatus ナミウツボ (FRLM 51573, 826.0 mm TL).

Gymnothorax zonipectis Seale, 1906
ヒレオビウツボ [Marine] (Fig. 26)

Fig. 23. Gymnothorax richardsonii モ バ ウ ツ ボ (upper: FRLM 51968,
344.0 mm TL; lower: KAUM–I. 80257, 238.6 mm TL).

8

Fig. 26. Gymnothorax zonipectis ヒ レ オ ビ ウ ツ ボ (Diving Service
MARLIN).

Checklist of fishes of Yonaguni-jima island
Rhinomuraena quaesita Garman, 1888
ハナヒゲウツボ [Marine] (Fig. 27)

FAMILY OPHICHTHIDAE
ウミヘビ科
Muraenichthys schultzei Bleeker, 1857
ミナミミミズアナゴ [Marine]
NSMT-P 55494, 89.7 mm TL, Agarizaki, hand net, 28
Mar. 1997.

Myrichthys maculosus (Cuvier, 1816)
モヨウモンガラドオシ [Marine]
NSMT-P 68223, 267.0 mm TL, Yonaguni-jima island, 5
July 2004.

FAMILY CONGRIDAE
アナゴ科
Conger cinereus Rüppell, 1830
キリアナゴ [Marine]
URM-P 1536, Sonai Fishing Port, 27 Oct. 1959.
Heteroconger hassi (Klausewitz and Eibl-Eibesfeldt, 1959)
チンアナゴ [Marine] (Fig. 30)
Fig. 27. Rhinomuraena quaesita ハ ナ ヒ ゲ ウ ツ ボ (Diving Service
MARLIN).

Uropterygius micropterus (Bleeker, 1852)
アミキカイウツボ [Marine; brackish] (Fig. 28)
FRLM 51579, 151.7 mm TL, mouth of Tabaru River, handnet, 24 Sept. 2015.

Fig. 28. Uropterygius micropterus ア ミ キ カ イ ウ ツ ボ (FRLM 51579,
151.7 mm TL).

Fig. 30. Heteroconger hassi チンアナゴ (Diving Service MARLIN).

FAMILY SYNAPHOBRANCHIDAE
ホラアナゴ科Dysomma anguillare Barnard, 1923
アサバホラアナゴ [Marine] (Fig. 29)
FRLM 50529, 106.5 mm TL, northwest of Yonaguni-jimaisland, stomach content of Pristipomoides argyrogrammicus(FRLM 50748), 16 Mar. 2015.

Fig. 29. Dysomma anguillare アサバホラアナゴ (FRLM 50529, 106.5 mm
TL, preserved specimen).

9K. Koeda et al.
ORDER CLUPEIFORMES
ニシン目
FAMILY CLUPEIDAE
ニシン科
Herklotsichthys quadrimaculatus (Rüppell, 1837)
ミズン [Marine; brackish] (Fig. 31)
FRLM 51444, 99.6 mm SL, mouth of Kubura River, castnet, 21 Sept. 2015; FRLM 51561, 108.0 mm SL, Kubura
Fishing Port, spear, 24 Sept. 2015; FRLM 51982, 104.4 mm
SL, FRLM 51983, 105.0 mm SL, FRLM 51984, 108.2 mm
SL, FRLM 51985, 110.2 mm SL, FRLM 51986, 105.1 mm
SL, mouth of Kubura River, cast net, 21 Sept. 2015.

ORDER GONORYNCHIFORMES
ネズミギス目FAMILY CHANIDAE
サバヒー科
Chanos chanos (Forsskål, 1775)
サバヒー [Marine; brackish] (Fig. 34)
はねまつ
Jordan and Tanaka (1927); Okinawa Prefecture
Experimental Fisheries Station (1934); Aoyagi (1948);
Yoshigou (2000); Yoshigou et al. (2001).
FRLM 51441, 130.5 mm SL, FRLM 51981, 114.8 mm SL,
mouth of Kubura River, cast net, 21 Sept. 2015; HMNH-P29,
85.2 mm SL, HMNH-P30, 55.3 mm SL, Kubura River, handnet, 8 Aug. 1999.

Fig. 31. Herklotsichthys quadrimaculatus ミズン (FRLM 51444, 99.6 mm SL).
Fig. 34. Chanos chanos サバヒー (FRLM 51441, 130.5 mm SL).

Spratelloides atrofasciatus Schultz, 1943
リュウキュウキビナゴ [Marine] (Fig. 32)
FRLM 51532, 21.8 mm SL, Sonai Fishing Port, fishinglight, 24 Sept. 2015.

Fig. 32. Spratelloides atrofasciatus リュウキュウキビナゴ (FRLM 51532,
21.8 mm SL).

Spratelloides delicatulus (Bennett, 1832)
ミナミキビナゴ [Marine] (Fig. 33)
FRLM 51505, 41.8 mm SL, Kubura Fishing Port, fishinglight, 20 Sept. 2015; KAUM–I. 78452, 43.1 mm SL, Kubura
Fishing Port, fishing light, 20 Sept. 2015.

Fig. 33. Spratelloides delicatulus ミ ナ ミ キ ビ ナ ゴ (upper: KAUM–I.
78452, 43.1 mm SL; lower: FRLM 51505, 41.8 mm SL).

FAMILY ENGRAULIDAE
カタクチイワシ科
Engraulis japonicus Temminck and Schlegel, 1846
カタクチイワシ [Marine]
URM-P 42131, off Yonaguni-jima island, 14 Mar. 2004.

10

ORDER CYPRINIFORMES
コイ目
FAMILY CYPRINIDAE
コイ科
Carassius sp.
ギンブナ [Freshwater]
Jordan and Tanaka (1927); Kuroiwa (1927); Okinawa
Prefecture Experimental Fisheries Station (1934); Ogawa(1937); Aoyagi (1948); Shibata (1964); Fujimoto (1972);
Nakamura (1974); Kouchi (1991).
Remarks. Jordan and Tanaka (1927), Kuroiwa (1927),
Okinawa Prefecture Experimental Fisheries Station (1934),
and Aoyagi (1948) reported Carassius auratus (Linnaeus,
1758) (Japanese name “Funa”) from Yonaguni-jima island.
Fujimoto (1972) noted that “Funa” and “Hibuna” (= C.
auratus or Carassius sp.) are occurred in Yonaguni-jima island
but he did not identify it as a species level. Kouchi (1991)
recorded “Ginbuna” from Yonaguni-jima island. Yoshigou(2000) did not collect Carassius sp. but he speculated that C.
auratus in those reports are misidentification of “Ginbuna”(Carassius auratus langsdorfii Temminck and Schlegel,1846). Takada et al. (2010) analyzed mitochondrial DNA of
Carrassius auratus-complex collected from China, Taiwan,
and Japan, and they reported three generic clades were foundfrom Ryukyu Islands including Ishigaki-jima island. Eachclade can be considered as a species or subspecies level, but
the taxonomic decision has not been concluded. Although
Hosoya (2013) applied Carassius sp. for “Ginbuna” without
detail explanation, we provisionally follow Hosoya (2013)
on the scientific name of “Ginbuna” in our species list, andexclude C. auratus.
Cyprinus carpio Linnaeus, 1758
コイ [Freshwater]
Fujimoto (1972); Ministry of the Environment (2010b).

Checklist of fishes of Yonaguni-jima island
FAMILY COBITIDAE
ドジョウ科
Misgurnus anguillicaudatus (Cantor, 1842)
ドジョウ [Freshwater] (Fig. 35)
Gushiken (1972); Nakamura (1974); Yoshigou (2000);
Okinawa Prefecture (2005).
FRLM 51537, 37.7 mm SL, male, Yonaguni-jima island(extensive locality not shown for conservation), hand net, 24
Sept. 2015.
Remarks. Although Misgrnus anguillicaudatus (Cantor,
1842) was reported by Gushiken (1972) firstly, no additional
collected reports have been presented the species has been
considered as an extinct species (Ministry of the Environment,
2010b). It was rediscovered in 2007 by Dr. J. Nakajima andhis research team (J. Nakajima, pers. comm.). After that the
population of M. anguillicaudatus has been monitored bythem and it is also collected during our survey.

Synodus dermatogenys Fowler, 1912
ミナミアカエソ [Marine; brackish] (Fig. 37)
Yoshigou et al. (2001).
HMNH-P125, 117.4 mm SL, Sonai, hand net, 9 Aug.
1999; KAUM–I. 78309, 118.3 mm SL, Kataburu Beach,line-fishing, 17 Sept. 2015; NSMT-P 55492, 36.1 mm SL,
Agarizaki, 28 Mar. 1997.

Fig. 37. Synodus dermatogenys ミナミアカエソ (KAUM–I. 78309, 118.3mm SL).

Synodus sp.
アカエソ属の一種 [Marine] (Fig. 38)
FRLM 51535, 33.8 mm SL, Sonai Fishing Port, fishinglight, 24 Sept. 2015.

Fig. 35. Misgurnus anguillicaudatus ドジョウ (FRLM 51537, 37.7 mm SL).

ORDER AULOPIFOEMES
ヒメ目
FAMILY AULOPIDAE
ヒメ科
Aulopus sp.
ハタタテヒメ [Marine]
NSMT-P 109704, 140.2 mm SL, southeast of Yonagunijima island, 1 Aug. 2006.
FAMILY SYNODONTIDAE
エソ科
Saurida gracilis (Quoy and Gaimard, 1824)
マダラエソ [Marine; brackish] (Fig. 36)
Yoshigou (2000); Yoshigou et al. (2001).
FRLM 51562, 183.5 mm SL, Kubura Fishing Port, spear,
24 Sept. 2015; HMNH-P81, 115.7 mm SL, Kubura River,
hand net, 9 Aug. 1999; KAUM–I. 78362, 71.1 mm SL, off
Kubura Hinan Port, hand net, 18 Sept. 2015.

Fig. 38. Synodus sp. (FRLM 51535, 33.8 mm SL).

Synodus variegatus (Lacepède, 1803)
ヒトスジエソ [Marine] (Fig. 39)
FRLM 51466, 122.2 mm SL, off Hikawa, line-fishing, 21
Sept. 2015; KAUM–I. 80280, 171.1 mm SL, Sonai Fishing
Port, line-fishing, 20 Sept. 2015; KAUM–I. 80422, 103.0 mm
SL, KAUM–I. 80423, 123.6 mm SL, off Hikawa, line-fishing,
21 Sept. 2015; MUFS 47022, 192.0 mm SL, MUFS 47023,
113.0 mm SL, Sonai Fishing Port, line-fishing, 23 Sept. 2015.

Fig. 39. Synodus variegatus ヒトスジエソ (upper: KAUM–I. 80280, 171.1mm SL; lower: MUFS 47023, 113.0 mm SL).

Fig. 36. Saurida gracilis マダラエソ (upper: FRLM 51562, 183.5 mm SL;
lower: KAUM–I. 78362, 71.1 mm SL).

11

K. Koeda et al.
ORDER POLYMIXIIFORMES
ギンメダイ目FAMILY POLYMIXIIDAE
ギンメダイ科Polymixia berndti Gilbert, 1905
アラメギンメ [Marine]
NSMT-P 71451, 150.0 mm SL, southeast of Yonagunijima island, 12 July 2005.

ORDER LOPHIIFORMES
アンコウ目
FAMILY ANTENNARIIDAE
カエルアンコウ科
Antennarius maculatus (Desjardins, 1840)
クマドリカエルアンコウ [Marine] (Fig. 40)

ORDER MUGILIFORMES
ボラ目
FAMILY MUGILIDAE
ボラ科
ぶらいゆ
Chelon macrolepis (Smith, 1846)
コボラ [Marine; brackish] (Fig. 42)
Yoshigou (2000); Yoshigou et al. (2001).
FRLM 51442, 89.8 mm SL, mouth of Kubura River, castnet, 21 Sept. 2015; FRLM 51546, 67.1 mm SL, mouth of
Kubura River, cast net, 24 Sept. 2015; FRLM 51978, 81.1 mm
SL, Kubura Fishing Port, cast net, 21 Sept. 2015; FRLM
52126, 64.0 mm SL, mouth of Kubura River, cast net, 24 Sept.
2015; HMNH-P31, 47.8 mm SL, Kubura River, hand net, 8Aug. 1999.

Fig. 42. Chelon macrolepis コボラ (FRLM 51442, 89.8 mm SL).

Chelon melinopterus (Valenciennes, 1836)
ヒルギメナダ [Marine; brackish] (Fig. 43)
FRLM 51545, 77.9 mm SL, mouth of Kubura River, castnet, 24 Sept. 2015.
Fig. 40. Antennarius maculatus クマドリカエルアンコウ (Diving Service
MARLIN).

FAMILY CHAUNACIDAE
フサアンコウ科
Chaunax sp.
フサアンコウ属の一種 [Marine] (Fig. 41)
FRLM 50528, northwest of Yonaguni-jima island, stomach
content of Pontinus macrocephalus (FRLM 50749), 16 Mar.
2015.

Fig. 43. Chelon melinopterus ヒルギメナダ (FRLM 51545, 77.9 mm SL).

Crenimugil crenilabis (Forsskål, 1775)
フウライボラ [Marine; brackish] (Fig. 44)
Yoshigou et al. (2001); Ito (2009).
HMNH-P126, 81.9 mm SL, HMNH-P127, 64.1 mm SL,
Sonai, hand net, 9 Aug. 1999.
Fig. 41. Chaunax sp. (FRLM 50528).

Fig. 44. Crenimugil crenilabis フウライボラ (HMNH-P126, 81.9 mm SL,
preserved specimen).

12

Checklist of fishes of Yonaguni-jima island
Ellochelon vaigiensis (Quoy and Gaimard, 1825)
オニボラ [Marine; brackish] (Fig. 45)
Yoshigou (2000); Yoshigou et al. (2001).
FRLM 51452, 58.3 mm SL, FRLM 52013, 66.8 mm SL,
mouth of Kubura River, cast net, 21 Sept. 2015; HMNH-P37,
17.2 mm SL, Kubura River, hand net, 8 Aug. 1999;
HMNH-P94, 32.8 mm SL, Tabaru River, hand net, 10 Aug.
1999; HMNH-P129, 29.5 mm SL, Sonai, hand net, 9 Aug.
1999; KAUM–I. 78443, 21.0 mm SL, Kubura Fishing Port,fishing light, 20 Sept. 2015.

Fig. 47. Moolgarda seheli タイワンメナダ (FRLM 51445, 45.0 mm SL).

Mugil cephalus cephalus Linnaeus, 1758
ボラ [Marine; brackish; freshwater]
Fujimoto (1972); Yoshigou (2000); Yoshigou et al. (2001).
HMNH-P89, 94.3 mm SL, Tabaru River, hand net, 10Aug. 1999.

Oedalechilus labiosus (Valenciennes, 1836)
ワニグチボラ [Marine] (Fig. 48)
FRLM 51491, 42.1 mm SL, Sonai Fishing Port, cast net,
22 Sept. 2015; HMNH-P128, 32.7 mm SL, Sonai, hand net,
9 Aug. 1999; KAUM–I. 78444, 19.9 mm SL, Kubura Fishing
Port, fishing light, 20 Sept. 2015.

Fig. 45. Ellochelon vaigiensis オ ニ ボ ラ (upper: FRLM 51452, 58.3 mm
SL; lower: KAUM–I. 78443, 21.0 mm SL).

Moolgarda pedaraki (Valenciennes, 1836)
カマヒレボラ [Marine; brackish] (Fig. 46)
FRLM 51544, 86.2 mm SL, mouth of Kubura River, castnet, 24 Sept. 2015.

Fig. 48. Oedalechilus labiosus ワ ニ グ チ ボ ラ (upper: KAUM–I. 78444,
19.9 mm SL; lower: FRLM 51491, 42.1 mm SL).
Fig. 46. Moolgarda pedaraki カマヒレボラ (FRLM 51544, 86.2 mm SL).

Moolgarda seheli (Forsskål, 1775)
タイワンメナダ [Marine; brackish] (Fig. 47)
Yoshigou (2000); Yoshigou et al. (2001).
FRLM 51445, 45.0 mm SL, mouth of Kubura River,
cast net, 21 Sept. 2015; FRLM 51979, 47.5 mm SL, Kubura
Fishing Port, cast net, 21 Sept. 2015; HMNH-P32, 39.2 mm
SL, Kubura River, hand net, 8 Aug. 1999.

13

K. Koeda et al.
ORDER ATHERINIFORMES
トウゴロウイワシ目
FAMILY ATHERIONIDAE
ムギイワシ科Atherion elymus Jordan and Starks, 1901
ムギイワシ [Marine] (Fig. 49)
Yoshigou et al. (2001).
FRLM 51465, 24.7 mm SL, Sonai Fishing Port, hand net,
20 Sept. 2015; HMNH-P119, 31.6 mm SL, Agarizaki, handnet, 9 Aug. 1999.

Zenarchopterus dunckeri Mohr, 1926
コモチサヨリ [Brackish] (Fig. 51)
Yoshigou (2000); Yoshigou et al. (2001).
FRLM 51513, 56.7 mm SL, Tabaru River, hand net, 23
Sept. 2015; HMNH-P49, 54.4 mm SL, Tabaru River, handnet, 9 Aug. 1999; KAUM–I. 78412, 102.2 mm SL, Tabaru
River, hand net, 20 Sept. 2015; KAUM–I. 78430, 73.0 mm
SL, KAUM–I. 78456, 23.2 mm SL, Tabaru River, cast net, 20
Sept. 2015.

Fig. 51. Zenarchopterus dunckeri コモチサヨリ (KAUM–I. 78412, 102.2mm SL).

Fig. 49. Atherion elymus ムギイワシ (FRLM 51465, 24.7 mm SL).

ORDER BELONIFORMES
ダツ目
FAMILY EXOCOETIDAE
トビウオ科
とぅびいゆ
Hirundichthys oxycephalus (Bleeker, 1852)
ホソアオトビ [Marine]
URM-P 39583, north of Yonaguni-jima island, 23 Nov.
1998.

FAMILY HEMIRAMPHIDAE
サヨリ科
Hyporhamphus dussumieri (Valenciennes, 1847)
マルサヨリ [Marine] (Fig. 50)
FRLM 51506, 61.9 mm SL, Kubura Fishing Port, fishinglight, 20 Sept. 2015.

FAMILY BELONIDAE
ダツ科
Platybelone argalus platyura (Bennett, 1832)
ヒメダツ [Marine] (Fig. 52)
NSMT-P 101228, 352.3 mm SL, Kubura Fishing Port, linefishing, 11 Mar. 2010; FRLM 50706, 302.5 mm SL, Kubura
Fishing Port, line-fishing, 13 Mar. 2015; FRLM 51470, 358.5mm SL, Kubura-bari, line-fishing, 22 Sept. 2015.

Fig. 52. Platybelone argalus platyura ヒメダツ (FRLM 51470, 358.5 mm SL).

Strongylura incisa (Valenciennes, 1846)
リュウキュウダツ [Marine] (Fig. 53)
FRLM 51469, 273.5 mm SL, Kubura Hinan Port, handnet, 21 Sept. 2015.

Fig. 53. Strongylura incisa リュウキュウダツ (FRLM 51469, 273.5 mm SL).
Fig. 50. Hyporhamphus dussumieri マルサヨリ (FRLM 51506, 61.9 mm SL).

Hemiramphus far (Forsskål, 1775)
ホシザヨリ [Marine]
だる
Ikema (2003).

Tylosurus crocodilus crocodilus (Péron and Lesueur, 1821)
オキザヨリ [Marine] (Fig. 54)
FRLM 51437, 256.0 mm SL, Sonai Fishing Port, hand net,
20 Sept. 2015; FRLM 51592, 265.6 mm SL, FRLM 51945,
256.5 mm SL, Sonai Fishing Port, hand net, 19 Sept. 2015;
MUFS 46950, 86.0 mm SL, Sonai Fishing Port, line-fishing,
20 Sept. 2015.

Fig. 54. Tylosurus crocodilus crocodilus オキザヨリ (FRLM 51437, 256.0
mm SL).

14

Checklist of fishes of Yonaguni-jima island
ORDER CYPRINODONTIFORMES
カダヤシ目
FAMILY POECILIIDAE
カダヤシ科
Gambusia affinis (Baird and Girard, 1853)
カダヤシ [Freshwater] (Fig. 55)
Fujimoto (1972); Takahara et al. (1997); Yoshigou (2000);
Yoshigou et al. (2001).
HMNH-P28, 20.0 mm SL, Tabaru River, hand net, 8 Aug.
1999; KAUM–I. 78431, 19.5 mm SL, KAUM–I. 78432, 15.1mm SL, Tabaru River, hand net, 20 Sept. 2015.
Remarks. Fujimoto (1972) reported “Medaka” [=Oryzias
latipes (Temminck and Schlegel, 1846)] from Yonagunijima island without specimens or photos. However, no later
publications mentioned about his report (e.g., Senou, 2013).
Nonetheless, this species known as not be distributed in the
Yaeyama Islands (Kuroiwa, 1927; Senou, 2013), and canconsider that was misidentification of Gambusia affinis whichis also collected during our survey. From 1924, the species
was widely introduced to Yaeyama Islands to exterminate
malarial mosquito (Takahara et al., 1997). It was firstly
introduced to Yonaguni-jima island in 1934 at first from
Ishigaki-jima island (Takahara et al., 1997). Therefore, it isconsidered that it is widely and numerously distributed in
Yonaguni-jima island when the survey by Fujimoto (1972).
Gambusia affinis is sometime confused with Oryzias spp.
because of the similar size and the body shape (Saitoh andUchiyama, 2015). Consequently, we exclude it from our list.

ORDER BERYCIFORMES
キンメダイ目FAMILY HOLOCENTRIDAE
イットウダイ科
Myripristis adusta Bleeker, 1853
ツマグロマツカサ [Marine] (Fig. 56)
Eguchi and Motomura (2016).
URM-P 48354, 149.8 mm SL, Kubura Fishing Port, linefishing, 25 July 2013.

Fig. 56. Myripristis adusta ツマグロマツカサ (URM-P 48354, 149.8 mm SL).

Myripristis berndti Jordan and Evermann, 1903
アカマツカサ [Marine] (Fig. 57)
Tatsuma (1999); Eguchi and Motomura (2016).
FRLM 51453, 166.5 mm SL, Kubura-bari, line-fishing,
20 Sept. 2015; FRLM 51493, 208.8 mm SL, mouth of Tabaru
River, gill net, 23 Sept. 2015; FRLM 52065, 161.8 mm SL,
Kubura Fishing Port, line-fishing, 24 Sept. 2015; KAUM–I.
78301, 80.1 mm SL, KAUM–I. 78302, 79.5 mm SL, Kubura
Fishing Port, line-fishing, 16 Sept. 2015; KAUM–I. 78373,
162.3 mm SL, Kubura Fishing Port, line-fishing, 18 Sept.
2015; MUFS 46880, 113.0 mm SL, Kubura Fishing Port, linefishing, 17 Sept. 2015; MUFS 46934, 178.0 mm SL, Sonai
Fishing Port, line-fishing, 18 Sept. 2015; URM-P 48351,
142.1 mm SL, Kubura Fishing Port, line-fishing, 24 July 2013.

Fig. 55. Gambusia affinis カダヤシ (KAUM–I. 78431, 19.5 mm SL).

Fig. 57. Myripristis berndti アカマツカサ (upper: KAUM–I. 78373, 162.3mm SL; lower: MUFS 46880, 113.0 mm SL).

15

K. Koeda et al.
Myripristis hexagona (Lacepède, 1802)
マルマツカサ [Marine] (Fig. 58)
FRLM 51433, 37.0 mm SL, Kubura Fishing Port, linefishing, 20 Sept. 2015.

Myripristis violacea Bleeker, 1851
セグロマツカサ [Marine] (Fig. 61)
Eguchi and Motomura (2016).
FRLM 52063, 93.7 mm SL, FRLM 52064, 105.6 mm SL,
Kubura Fishing Port, line-fishing, 24 Sept. 2015; URM-P
48350, 150.4 mm SL, URM-P 48352, 82.9 mm SL, URM-P
48353, 93.5 mm SL, Kubura Fishing Port, line-fishing, 24
July 2013.

Fig. 58. Myripristis hexagona マルマツカサ (FRLM 51433, 37.0 mm SL).

Myripristis kuntee Valenciennes, 1831
クロオビマツカサ [Marine] (Fig. 59)
Tatsuma (1999); Eguchi and Motomura (2016).
MUFS 46929, 95.0 mm SL, Sonai Fishing Port, linefishing, 18 Sept. 2015.

Fig. 61. Myripristis violacea セグロマツカサ (URM-P 48350, 150.4 mm SL).

Myripristis vittata Valenciennes, 1831
ベニマツカサ [Marine]
Tatsuma (1999); Nakabo (2013).
Myripristis sp. 1
アカマツカサ属の一種 [Marine] (Fig. 62)
KAUM–I. 78554, 44.4 mm SL, Umabana, hand net, 22
Sept. 2015.

Fig. 59.
kuntee クロオビマツカサ (MUFS 46929, 95.0 mm SL,
preserved specimen).

Myripristis murdjan (Forsskål, 1775)
ヨゴレマツカサ [Marine] (Fig. 60)
Eguchi and Motomura (2016).
KAUM–I. 80283, 117.6 mm SL, Sonai Fishing Port, linefishing, 20 Sept. 2015.

Fig. 62. Myripristis sp. 1 (KAUM–I. 78554, 44.4 mm SL).

Myripristis sp. 2
アカマツカサ属の一種 [Marine] (Fig. 63)
KAUM–I. 80463, 98.3 mm SL, Umabana, hand net, 22
Sept. 2015.

Fig. 60. Myripristis murdjan ヨゴレマツカサ (KAUM–I. 80283, 117.6 mm SL).

Fig. 63. Myripristis sp. 2 (KAUM–I. 80463, 98.3 mm SL).

16

Checklist of fishes of Yonaguni-jima island
Neoniphon aurolineatus (Liénard, 1839)
ホホベニイットウダイ [Marine]
Tatsuma (1999); Eguchi and Motomura (2016).

Neoniphon opercularis (Valenciennes, 1831)
ヒレグロイットウダイ [Marine] (Fig. 64)
Nakabo (2013); Eguchi and Motomura (2016).
KAUM–I. 78327, 164.0 mm SL, Anchor's graves, spear,
17 Sept. 2015; KAUM–I. 80272, 197.1 mm SL, Sonai
Channel, hand net, 21 Sept. 2015; MUFS 46940, 180.0 mm
SL, Kubura Fishing Port, line-fishing, 19 Sept. 2015.

Sargocentron diadema (Lacepède, 1802)
ニジエビス [Marine] (Fig. 66)
Eguchi and Motomura (2016).
FRLM 52016, 92.3 mm SL, FRLM 52017, 89.9 mm SL,
FRLM 52018, 80.2 mm SL, FRLM 52019, 53.7 mm SL,
Kubura Fishing Port, line-fishing, 19 Sept. 2015; KAUM–
I. 78349, 94.3 mm SL, off Kubura Hinan Port, hand net, 18
Sept. 2015; URM-P 48355, 104.8 mm SL, URM-P 48356,
104.4 mm SL, Kubura Fishing Port, line-fishing, 25 July 2013.

Fig. 66. Sargocentron diadema ニジエビス (KAUM–I. 78349, 94.3 mm SL).

Sargocentron ensiferum (Jordan and Evermann, 1903)
ハナエビス [Marine]
Tatsuma (1999); Eguchi and Motomura (2016).

Fig. 64. Neoniphon opercularis ヒレグロイットウダイ (upper: KAUM–I.
80272, 197.1 mm SL; lower: Underwater Ruin, K. Koeda).

Neoniphon sammara Forsskål, 1775
ウケグチイットウダイ [Marine] (Fig. 65)
Eguchi and Motomura (2016).
KAUM–I. 80275, 113.2 mm SL, Sonai Channel, handnet, 21 Sept. 2015; NSMT-P 101119, 115.7 mm SL, Sonai
Channel, hand net, 10 Mar. 2010.

Fig. 65. Neoniphon sammara ウケグチイットウダイ (KAUM–I. 80275,
113.2 mm SL).

Sargocentron ittodai (Jordan and Fowler, 1902)
テリエビス [Marine] (Fig. 67)
Eguchi and Motomura (2016).
FRLM 50704, 107.6 mm SL, FRLM 50712, 70.4 mm
SL, Kubura Fishing Port, line-fishing, 13 Mar. 2015; FRLM
51948, 91.7 mm SL, FRLM 51949, 96.4 mm SL, Sonai
Fishing Port, line-fishing, 19 Sept. 2015; FRLM 52022, 90.1
mm SL, FRLM 52023, 93.8 mm SL, Kubura Fishing Port,line-fishing, 19 Sept. 2015; FRLM 52072, 126.2 mm SL,
FRLM 52073, 90.1 mm SL, FRLM 52074, 89.3 mm SL,
FRLM 52075, 90.6 mm SL, FRLM 52076, 86.5 mm SL,
FRLM 52080, 85.5 mm SL, Sonai Fishing Port, line-fishing,
24 Sept. 2015; KAUM–I. 78644, 118.3 mm SL, Kubura
Fishing Port, line-fishing, 18 Sept. 2015; KAUM–I. 80388,
122.8 mm SL, KAUM–I. 80389, 114.0 mm SL, Umabana,hand net, 22 Sept. 2015; MUFS 46881, 112.0 mm SL, Kubura
Fishing Port, line-fishing, 17 Sept. 2015.

Fig. 67. Sargocentron ittodai テリエビス (KAUM–I. 80389, 114.0 mm SL).

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K. Koeda et al.
Sargocentron punctatissimum (Cuvier, 1829)
ホシエビス [Marine]
Eguchi and Motomura (2016).
NSMT-P 55455, 12 specimens, 73.3–109.9 mm SL,
Agarizaki, hand net, 28 Mar. 1997.

ORDER GASTEROSTEIFORMES
トゲウオ目
FAMILY PEGASIDAE
ウミテング科Eurypegasus draconis (Linnaeus, 1766)
ウミテング [Marine] (Fig. 70)

Sargocentron spiniferum (Forsskål, 1775)
トガリエビス [Marine] (Fig. 68)
みぶた
Ikema (2003); Ito (2009); Nakabo (2013).

Fig. 70. Eurypegasus draconis ウミテング (Diving Service MARLIN).

Fig. 68. Sargocentron spiniferum ト ガ リ エ ビ ス (fish landing port at
Yonaguni-jima island, K. Koeda).

Sargocentron tiere (Cuvier, 1829)
アオスジエビス [Marine] (Fig. 69)
Eguchi and Motomura (2016).
KAUM–I. 78372, 180.3 mm SL, Kubura Fishing Port,line-fishing, 18 Sept. 2015.

FAMILY SOLENOSTOMIDAE
カミソリウオ科
Solenostomus cyanopterus Bleeker, 1854
カミソリウオ [Marine] (Fig. 71)

Fig. 71. Solenostomus cyanopterus カ ミ ソ リ ウ オ (Diving Service
MARLIN).

Solenostomus paradoxus (Pallas, 1770)
ニシキフウライウオ [Marine] (Fig. 72)

Fig. 69. Sargocentron tiere アオスジエビス (KAUM–I. 78372, 180.3 mm SL).
Fig. 72. Solenostomus paradoxus ニシキフウライウオ (Diving Service
MARLIN).

FAMILY SYNGNATHIDAE
ヨウジウオ科
Corythoichthys haematopterus (Bleeker, 1851)
イシヨウジ [Marine] (Fig. 73)
KAUM–I. 78406, 115.2 m SL, Sonai Channel, hand net,
20 Sept. 2015.

Fig. 73. Corythoichthys haematopterus イ シ ヨ ウ ジ (KAUM–I. 78406,
115.2 mm SL).

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Checklist of fishes of Yonaguni-jima island
Doryrhamphus (Dunckerocampus) naia Allen and Kuiter, 2004
カスミオイランヨウジ [Marine] (Fig. 74)
KAUM–I. 78538, 90.6 mm SL, Umabana, hand net, 22
Sept. 2015.

Fig. 74. Doryrhamphus (Dunckerocampus) naia カスミオイランヨウジ(KAUM–I. 78538, 90.6 mm SL).

Microphis (Oostethus) brachyurus brachyurus (Bleeker, 1854)
テングヨウジ [Brackish; freshwater] (Fig. 75)
Yoshigou (2000); Yoshigou et al. (2001).
HMNH-P48, 103.2 mm SL, Tabaru River, hand net, 9Aug. 1999.

FAMILY AULOSTOMIDAE
ヘラヤガラ科
Aulostomus chinensis (Linnaeus, 1766)
ヘラヤガラ [Marine] (Fig. 78)
FRLM 52105, 652.5 mm SL, Kubura Fishing Port, spear,
24 Sept. 2015.

Fig. 78. Aulostomus chinensis ヘラヤガラ (FRLM 52105, 652.5 mm SL).

FAMILY FISTULARIIDAE
ヤガラ科
Fistularia commersonii Rüppell, 1838
アオヤガラ [Marine] (Fig. 79)
FRLM 51536, 152.8 mm SL, Sonai Fishing Port, fishinglight, 24 Sept. 2015; MUFS 46937, 927.0 mm SL, Sonai
Fishing Port, line-fishing, 19 Sept. 2015.

Fig. 75. Microphis (Oostethus) brachyurus brachyurus テ ン グ ヨ ウ ジ(HMNH-P48, 103.2 mm SL, preserved specimen).

Phoxocampus belcheri (Kaup, 1856)
ボウヨウジ [Marine] (Fig. 76)
Yoshigou et al. (2001).
HMNH-P122, 41.8 mm SL, Agarizaki, hand net, 9 Aug.
1999.

Fig. 76. Phoxocampus belcheri ボウヨウジ (HMNH-P122, 41.8 mm SL).

Fig. 79. Fistularia commersonii アオヤガラ (MUFS 46937, 927.0 mm SL).

FAMILY CENTRISCIDAE
ヘコアユ科
Aeoliscus strigatus (Günther, 1861)
ヘコアユ [Marine] (Fig. 80)
KAUM–I. 78258, 140.9 mm SL, Kubura Fishing Port,
hand net, 16 Sept. 2015.

Trachyrhamphus bicoarctatus (Bleeker, 1857)
ワカヨウジ [Marine] (Fig. 77)
Fig. 80. Aeoliscus strigatus ヘコアユ (KAUM–I. 78258, 140.9 mm SL).

Fig. 77. Trachyrhamphus bicoarctatus ワ カ ヨ ウ ジ (Diving Service
MARLIN).

19

K. Koeda et al.
ORDER SCORPAENIFORMES
カサゴ目
FAMILY SCORPAENIDAE
フサカサゴ科
Caracanthus maculatus (Gray, 1831)
ダンゴオコゼ [Marine] (Fig. 81)

Parascorpaena mcadamsi (Fowler, 1938)
トゲイッテンフサカサゴ [Marine] (Fig. 83)
KAUM–I. 78540, 49.0 mm SL, Umabana, hand net, 22
Sept. 2015.

Fig. 83. Parascorpaena mcadamsi トゲイッテンフサカサゴ (KAUM–I.
78540, 49.0 mm SL).

Fig. 81. Caracanthus maculatus ダンゴオコゼ (Diving Service MARLIN).

Pontinus macrocephalus (Sauvage, 1882)
ヒオドシ [Marine] (Fig. 84)
FRLM 50749, 208.2 mm SL, northwest of Yonaguni-jimaisland, line-fishing, 16 Mar. 2015.

Dendrochirus zebra (Cuvier, 1829)
キリンミノ [Marine] (Fig. 82)
FRLM 51591, 130.7 mm SL, Kubura Fishing Port, handnet, 20 Sept. 2015; KAUM–I. 78487, 25.8 mm SL, Sonai
Channel, hand net, 21 Sept. 2015; KAUM–I. 80364, 116.0 mm
SL, Sonai Channel, gill net, 20 Sept. 2015; MUFS 47035, 115.0mm SL, Kubura Fishing Port, line-fishing, 25 Sept. 2015.

Fig. 84. Pontinus macrocephalus ヒオドシ (FRLM 50749, 208.2 mm SL).

Pterois antennata (Bloch, 1787)
ネッタイミノカサゴ [Marine] (Fig. 85)
あばいゆ・はにあばFig. 85. Pterois antennata ネッタイミノカサゴ (Umabana, K. Koeda)

Fig. 82. Dendrochirus zebra キリンミノ (upper: KAUM–I. 80364, 116.0mm SL; lower: MUFS 47035, 115.0 mm SL).

20

Pterois radiata Cuvier, 1829
キミオコゼ [Marine]
あばいゆ・はにあば
NSMT-P 55493, 109.8 mm SL, Agarizaki, hand net, 28
Mar. 1997.

Checklist of fishes of Yonaguni-jima island
Pterois volitans (Linnaeus, 1758)
ハナミノカサゴ [Marine] (Fig. 86)
あばいゆ・はにあば
FRLM 51590, 150.8 mm SL, Kubura Fishing Port, linefishing, 20 Sept. 2015; FRLM 51946, 138.4 mm SL, Sonai
Fishing Port, line-fishing, 19 Sept. 2015; KAUM–I. 78482,
41.6 mm SL, Sonai Channel, hand net, 21 Sept. 2015.

Fig. 87. Scorpaenodes guamensis グアムカサゴ (upper: KAUM–I. 78505,
40.1 mm SL; lower: KAUM–I. 78473, 19.2 mm SL).

Scorpaenodes parvipinnis (Garrett, 1864)
ヒメサンゴカサゴ [Marine] (Fig. 88)
KAUM–I. 78548, 27.5 mm SL, Danno-dropoff, hand net,
22 Sept. 2015.

Fig. 86. Pterois volitans ハナミノカサゴ (upper: FRLM 51590, 150.8 mm
SL; lower: Diving Service MARLIN)

Scorpaena onaria Jordan and Snyder, 1900
フサカサゴ [Marine]
NSMT-P 76155, 165.5 mm SL, southeast of Yonagunijima island, 1 Aug. 2006.

Scorpaenodes guamensis Quoy and Gaimard, 1824
グアムカサゴ [Marine] (Fig. 87)
Yoshigou et al. (2001).
FRLM 51424, 58.0 mm SL, Kubura Fishing Port, linefishing, 19 Sept. 2015; HMNH-P96, 94.8 mm SL, Kubura
River, hand net, 10 Aug. 1999; HMNH-P130, 22.4 mm SL,
Sonai, hand net, 9 Aug. 1999; KAUM–I. 78473, 19.2 mm SL,
Sonai Channel, hand net, 21 Sept. 2015; KAUM–I. 78504,
40.3 mm SL, KAUM–I. 78505, 40.1 mm SL, Hikawa, handnet, 21 Sept. 2015.

Fig. 88. Scorpaenodes parvipinnis ヒメサンゴカサゴ (KAUM–I. 78548,
27.5 mm SL).

Scorpaenodes varipinnis Smith, 1957
セボシイソカサゴ [Marine] (Fig. 89)
HMNH-P783, 8.3 mm SL, Sonai, hand net, 9 Aug. 1999;
KAUM–I. 78360, 42.7 mm SL, off Kubura Hinan Port, handnet, 18 Sept. 2015.

Fig. 89. Scorpaenodes varipinnis セ ボ シ イ ソ カ サ ゴ (KAUM–I. 78360,
42.7 mm SL).

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K. Koeda et al.
Scorpaenopsis diabolus (Cuvier, 1829)
ニライカサゴ [Marine] (Fig. 90)
Ito (2009).

Scorpaenopsis possi Randall and Eschmeyer, 2001
ミミトゲオニカサゴ [Marine] (Fig. 93)
FRLM 51447, 102.4 mm SL, Kubura-bari, line-fishing, 20
Sept. 2015.

Fig. 93. Scorpaenopsis possi ミミトゲオニカサゴ (FRLM 51447, 102.4mm SL).
Fig. 90. Scorpaenopsis diabolus ニライカサゴ (Diving Service MARLIN).

Scorpaenopsis neglecta Heckel, 1839
サツマカサゴ [Marine] (Fig. 91)

Sebastapistes cyanostigma (Bleeker, 1856)
カスリフサカサゴ [Marine] (Fig. 94)
KAUM–I. 78545, 40.6 mm SL, Danno-dropoff, hand net,
22 Sept. 2015; KAUM–I. 78576, 54.6 mm SL, Danno Beach,hand net, 23 Sept. 2015.

Fig. 94. Sebastapistes cyanostigma カスリフサカサゴ (KAUM–I. 78545,
40.6 mm SL).
Fig. 91. Scorpaenopsis neglecta サツマカサゴ (Diving Service MARLIN).

Scorpaenopsis papuensis (Cuvier, 1829)
ウルマカサゴ [Marine] (Fig. 92)

Fig. 92. Scorpaenopsis papuensis ウ ル マ カ サ ゴ (Diving Service
MARLIN).

22

Sebastapistes fowleri (Pietschmann, 1934)
プチフサカサゴ [Marine] (Fig. 95)
KAUM–I. 78523, 23.0 mm SL, Danno-dropoff, hand net,
22 Sept. 2015.

Fig. 95. Sebastapistes fowleri プ チ フ サ カ サ ゴ (KAUM–I. 78523, 23.0mm SL).

Checklist of fishes of Yonaguni-jima island
Sebastapistes strongia (Cuvier, 1829)
マダラフサカサゴ [Marine] (Fig. 96)
KAUM–I. 78503, 44.4 mm SL, Hikawa, hand net, 21
Sept. 2015.

Fig. 96. Sebastapistes strongia マダラフサカサゴ (KAUM–I. 78503, 44.4mm SL).

Taenianotus triacanthus Lacepède, 1802
ハダカハオコゼ [Marine] (Fig. 97)

Fig. 99. Synanceia verrucosa オニダルマオコゼ (upper: HMNH-P7190,
22.7 mm SL; lower: Diving Service MARLIN).

Fig. 97. Taenianotus triacanthus ハ ダ カ ハ オ コ ゼ (Diving Service
MARLIN).

FAMILY TETRAROGIDAE
ハオコゼ科
Ablabys taenianotus (Cuvier, 1829)
ツマジロオコゼ [Marine] (Fig. 98)

FAMILY PLATYCEPHALIDAE
コチ科
Sunagocia arenicola (Schultz, 1966)
スナゴチ [Marine] (Fig. 100)
KAUM–I. 78557, 63.3 mm SL, Umabana, hand net, 22
Sept. 2015.

Fig. 100. Sunagocia arenicola スナゴチ (KAUM–I. 78557, 63.3 mm SL).

Thysanophrys chiltonae Schultz, 1966
クロシマゴチ [Marine] (Fig. 101)
KAUM–I. 78481, 28.0 mm SL, Sonai Channel, hand net,
21 Sept. 2015.

Fig. 98. Ablabys taenianotus ツマジロオコゼ (Diving Service MARLIN).

FAMILY SYNANCEIIDAE
オニオコゼ科
いちあば
Synanceia verrucosa Bloch and Schneider, 1801
オニダルマオコゼ [Marine] (Fig. 99)
HMNH-P7190, 22.7 mm SL, Sonai, hand net, 9 Aug. 1999.

Fig. 101. Thysanophrys chiltonae クロシマゴチ (KAUM–I. 78481, 28.0mm SL).

23

K. Koeda et al.
ORDER PERCIFORMES
スズキ目
FAMILY AMBASSIDAE
タカサゴイシモチ科
Ambassis buruensis Bleeker, 1856
トゲナガタカサゴイシモチ [Brackish; freshwater] (Fig. 102)
Yoshigou (2000).
FRLM 51547, 48.5 mm SL, FRLM 52111, 47.1 mm SL,
mouth of Kubura River, cast net, 24 Sept. 2015; HMNH-P
4542, 54.8 mm SL, HMNH-P4543, 45.4 mm SL, Tabaru
River, hand net, 9 Aug. 1999; KAUM–I. 78422, 64.0 mm SL,
Tabaru River, hand net, 20 Sept. 2015; KAUM–I. 78424, 69.1mm SL, Tabaru River, cast net, 20 Sept. 2015.

Fig. 102. Ambassis buruensis ト ゲ ナ ガ タ カ サ ゴ イ シ モ チ (KAUM–I.
78424, 69.1 mm SL).

Ambassis interrupta Bleeker, 1853
ナンヨウタカサゴイシモチ [Brackish; freshwater] (Fig. 103)
Yoshigou (2000); Yoshigou and Nakamura (2003).
HMNH-P4538, 65.7 mm SL, Tabaru River, hand net, 10Aug. 1999; HMNH-P4539, 69.3 mm SL, Tabaru River, handnet, 8 Aug. 1999; HMNH-P4540, 78.8 mm SL, HMNH-P
4541, 41.1 mm SL, Tabaru River, hand net, 9 Aug. 1999.

Fig. 104. Ambassis miops セスジタカサゴイシモチ (FRLM 51548, 46.0mm SL).

Ambassis urotaenia Bleeker, 1852
タカサゴイシモチ [Brackish; freshwater] (Fig. 105)
Yoshigou (2000); Yoshigou and Nakamura (2003).
HMNH-P4546, 47.9 mm SL, HMNH-P4547, 44.6 mm
SL, mouth of Kubura River, cast net, 8 Aug. 1999; HMNH-P
4548, 47.8 mm SL, HMNH-P4549, 45.3 mm SL, HMNH-P
4550, 45.1 mm SL, Kubura River, hand net, 8 Aug. 1999;
FRLM 52112, 44.6 mm SL, FRLM 52113, 42.7 mm SL,
Kubura River, hand net, 24 Sept. 2015; KAUM–I. 78414, 60.7mm SL, KAUM–I. 78415, 49.0 mm SL, KAUM–I. 78417,
58.5 mm SL, KAUM–I. 78418, 61.1 mm SL, KAUM–I.
78419, 59.9 mm SL, KAUM–I. 78420, 67.9 mm SL, KAUM–
I. 78421, 64.7 mm SL, KAUM–I. 78423, 39.0 mm SL, Tabaru
River, hand net, 20 Sept. 2015; KAUM–I. 78416, 55.7 mm
SL, Tabaru River, gill net, 20 Sept. 2015; KAUM–I. 78425,
69.4 mm SL, KAUM–I. 78426, 64.5 mm SL, KAUM–I.
78427, 41.0 mm SL, KAUM–I. 78428, 40.3 mm SL, KAUM–I.
78457, 56.1 mm SL, KAUM–I. 78458, 51.4 mm SL, KAUM–
I. 78459, 53.5 mm SL, Tabaru River, cast net, 20 Sept. 2015.

Fig. 105. Ambassis urotaenia タカサゴイシモチ (KAUM–I. 78414, 60.7mm SL).

Fig. 103. Ambassis interrupta ナンヨウタカサゴイシモチ (HMNH-P4539,
69.3 mm SL).

Ambassis miops Günther, 1872
セスジタカサゴイシモチ [Brackish; freshwater] (Fig. 104)
Yoshigou (2000); Yoshigou and Nakamura (2003).
FRLM 51548, 46.0 mm SL, FRLM 52109, 47.1 mm SL,
mouth of Kubura River, cast net, 24 Sept. 2015; HMNH-P
4544, 62.0 mm SL, HMNH-P4545, 60.7 mm SL, Tabaru
River, hand net, 8 Aug. 1999.

24

Checklist of fishes of Yonaguni-jima island
FAMILY SERRANIDAE
ハタ科にーばい
Cephalopholis argus Schneider, 1801
アオノメハタ [Marine] (Fig. 106)
MUFS 46906, 209.0 mm SL, off Kubura Hinan Port,line-fishing, 18 Sept. 2015; NSMT-P 55464, 57.1 mm SL,
Agarizaki, hand net, 28 Mar. 1997; NSMT-P 101125, 223.0mm SL, Hikawa, 11 Mar. 2010.

Fig. 106. Cephalopholis argus アオノメハタ (MUFS 46906, 209.0 mm SL).

Cephalopholis urodeta (Forster, 1801)
ニジハタ [Marine] (Fig. 109)
FRLM 52093, 118.4 mm SL, off Hikawa, line-fishing,
21 Sept. 2015; KAUM–I. 78646, 173.4 mm SL, KAUM–
I. 78647, 169.8 mm SL, KAUM–I. 78648, 147.2 mm SL,
KAUM–I. 80221, 155.0 mm SL, Arakawa Bay, line-fishing,
18 Sept. 2015; KAUM–I. 80395, 150.5 mm SL, KAUM–
I. 80420, 118.2 mm SL, off Hikawa, line-fishing, 21 Sept.
2015; KAUM–I. 80455, 119.5 mm SL, north of Yonagunijima island, line-fishing, 22 Sept. 2015; MUFS 46895, 134.0
mm SL, MUFS 46896, 155.0 mm SL, MUFS 46897, 158.0
mm SL, MUFS 46899, 161.0 mm SL, off Kubura Hinan Port,line-fishing, 18 Sept. 2015; MUFS 46980, 98.0 mm SL, off
Hikawa, line-fishing, 21 Sept. 2015; MUFS 47013, 114.0 mm
SL, off Northern Pasture, line-fishing, 22 Sept. 2015.
Remarks. Caudal fin of Cephalopholis urodeta normaly
rounded, however, upper most caudal-fin ray of MUFS 46897
elongated.

Cephalopholis sonnerati (Valenciennes, 1828)
アザハタ [Marine] (Fig. 107)
MUFS 47005, 258.0 mm SL, off Northern Pasture, linefishing, 22 Sept. 2015.

Fig. 107. Cephalopholis sonnerati アザハタ (MUFS 47005, 258.0 mm SL).

Cephalopholis spiloparaea (Valenciennes, 1828)
アカハナ [Marine] (Fig. 108)
FRLM 51458, 113.7 mm SL, off Hikawa, line-fishing, 21
Sept. 2015; KAUM–I. 80385, 154.5 mm SL, Umabana, linefishing, 22 Sept. 2015.

Fig. 109. Cephalopholis urodeta ニ ジ ハ タ (upper: KAUM–I. 80455,
119.5 mm SL; middle: MUFS 46897, 158.0 mm SL; lower: Umabana,K. Koeda).

Fig. 108. Cephalopholis spiloparaea ア カ ハ ナ (KAUM–I. 80385, 154.5mm SL).

25

K. Koeda et al.
Epinephelus bruneus Bloch, 1793
クエ [Marine] (Fig. 110)
Fujimoto (1972).

Fig. 110. Epinephelus bruneus ク エ (fish landing port at Yonaguni-jimaisland, K. Koeda).

Fig. 112. Epinephelus hexagonatus イシガキハタ (KAUM–I. 78377, 128.4mm SL).

Epinephelus lanceolatus (Bloch, 1790)
タマカイ [Marine] (Fig. 113)
Epinephelus fasciatus (Forsskål, 1775)
アカハタ [Marine] (Fig. 111)
FRLM 52081, 158.2 mm SL, off Hikawa, line-fishing, 21
Sept. 2015; KAUM–I. 79712, 219.7 mm SL, Arakawa Bay,
spear, 18 Sept. 2015; KAUM–I. 79740, 214.4 mm SL, off
Kubura Hinan Port, spear, 18 Sept. 2015; KAUM–I. 80419,
147.1 mm SL, off Hikawa, line-fishing, 21 Sept. 2015; MUFS
47036, 202.0 mm SL, levee at Kubura, line-fishing, 25 Sept.
2015.

Fig. 113. Epinephelus lanceolatus タマカイ (Diving Service MARLIN).

Epinephelus maculatus (Bloch, 1790)
シロブチハタ [Marine] (Fig. 114)
KAUM–I. 78606, 364.0 mm SL, off Hikawa, line-fishing,
21 Sept. 2015.

Fig. 111. Epinephelus fasciatus アカハタ (upper: KAUM–I. 79740, 214.4mm SL; lower: MUFS 47036, 202.0 mm SL).

Epinephelus hexagonatus (Forster, 1801)
イシガキハタ [Marine] (Fig. 112)
いちにーばいFRLM 50747, 105.5 mm SL, Kubura-bari, line-fishing,
14 Mar. 2015; FRLM 51961, 135.7 mm SL, Kubura-bari,line-fishing, 20 Sept. 2015; KAUM–I. 78377, 128.4 mm SL,
Kubura Hinan Port, hand net, 19 Sept. 2015; NSMT-P 55458,
6 specimens, 116.8–143.7 mm SL, Agarizaki, hand net, 28
Mar. 1997.

26

Fig. 114. Epinephelus maculatus シロブチハタ (upper: KAUM–I. 78606,
364.0 mm SL; lower: Diving Service MARLIN).

Checklist of fishes of Yonaguni-jima island
Epinephelus melanostigma Schultz, 1953
スミツキハタ [Marine] (Fig. 115)
FRLM 51499, 234.5 mm SL, Sonai Fishing Port, linefishing, 22 Sept. 2015; KAUM–I. 80226, 220.1 mm SL,
Rakuda Rock, spear, 17 Sept. 2015; MUFS 47020, 199.0 mm
SL, Kubura-bari, line-fishing, 23 Sept. 2015; NSMT-P 55457,
153.0 mm SL, Agarizaki, hand net, 28 Mar. 1997.

Epinephelus octofasciatus (Griffin, 1926)
マハタモドキ [Marine] (Fig. 117)

Fig. 117. Epinephelus octofasciatus マ ハ タ モ ド キ (fish landing port at
Yonaguni-jima island, K. Koeda).

Epinephelus polyphekadion (Bleeker, 1849)
マダラハタ [Marine] (Fig. 118)
Fig. 115. Epinephelus melanostigma スミツキハタ (MUFS 47020, 199.0mm SL).

Epinephelus merra Bloch, 1793
カンモンハタ [Marine] (Fig. 116)
FRLM 52060, 205.0 mm SL, FRLM 52061, 167.0 mm SL,
Hikawa, line-fishing, 24 Sept. 2015; KAUM–I. 78305, 127.9mm SL, Kataburu Beach, line-fishing, 17 Sept. 2015; KAUM–
I. 80279, 177.9 mm SL, Sonai Fishing Port, line-fishing, 20
Sept. 2015; MUFS 47034, 182.0 mm SL, Sonai Fishing Port,line-fishing, 24 Sept. 2015.

Fig. 118. Epinephelus polyphekadion マ ダ ラ ハ タ (Diving Service
MARLIN).

Epinephelus septemfasciatus (Thunberg, 1793)
マハタ [Marine] (Fig. 119)Fig. 119. Epinephelus septemfasciatus マハタ (fish landing port at Yonagunijima island, K. Koeda).

Epinephelus tauvina (Forsskål, 1775)
ヒトミハタ [Marine] (Fig. 120)
FRLM 51954, 245.5 mm SL, Kubura-bari, line-fishing, 20
Sept. 2015; NSMT-P 55459, 103.6 mm SL, Agarizaki, handnet, 28 Mar. 1997.

Fig. 116. Epinephelus merra カ ン モ ン ハ タ (upper: KAUM–I. 78305,
127.9 mm SL; middle: MUFS 47034, 182.0 mm SL; lower: KAUM–I.
80279, 177.9 mm SL).

Fig. 120. Epinephelus tauvina ヒトミハタ (FRLM 51954, 245.5 mm SL).

27

K. Koeda et al.
Grammistes sexlineatus (Thunberg, 1792)
ヌノサラシ [Marine] (Fig. 121)
Yoshigou et al. (2001).
HMNH-P115, 32.2 mm SL, Agarizaki, hand net, 9 Aug.
1999; KAUM–I. 78378, 89.3 mm SL, off Kubura Hinan Port,line-fishing, 19 Sept. 2015; KAUM–I. 78578, 12.9 mm SL,
Danno Beach, hand net, 23 Sept. 2015; NSMT-P 55481, 3
specimens, 21.9–26.0 mm SL, Agarizaki, hand net, 28 Mar.
1997.

Plectropomus laevis (Lacepède, 1801)
コクハンアラ [Marine] (Fig. 123)

Fig. 123. Plectropomus laevis コクハンアラ (Diving Service MARLIN).

Pogonoperca punctata (Valenciennes, 1830)
アゴハタ [Marine] (Fig. 124)
Yoshigou et al. (2001).
HMNH-P116, 11.8 mm SL, Agarizaki, hand net, 9 Aug. 1999.

Fig. 124. Pogonoperca punctata ア ゴ ハ タ (HMNH-P116, 11.8 mm SL,
preserved specimen).

Fig. 121. Grammistes sexlineatus ヌ ノ サ ラ シ (upper: KAUM–I. 78378,
89.3 mm SL; lower: KAUM–I. 78578, 12.9 mm SL).

Liopropoma latifasciatum (Tanaka, 1922)
ツルグエ [Marine]
Ito (2009).

Pseudanthias dispar (Herre, 1955)
アカネハナゴイ [Marine] (Fig. 125)
FRLM 51478, 20.9 mm SL, Umabana, hand net, 22 Sept.
2015; KAUM–I. 78337, 48.2 mm SL, KAUM–I. 78338, 44.7mm SL, KAUM–I. 78339, 31.3 mm SL, off Kubura Hinan
Port, hand net, 18 Sept. 2015; KAUM–I. 78542, 21.5 mm SL,
Umabana, hand net, 22 Sept. 2015.

Plectranthias kamii Randall, 1980
イズハナダイ [Marine]
Yoshino (1972); Nakabo (2013).

Plectranthias longimanus (Weber, 1913)
チビハナダイ [Marine] (Fig. 122)
KAUM–I. 78365, 23.5 mm SL, KAUM–I. 78366, 26.6mm SL, KAUM–I. 78367, 22.1 mm SL, off Kubura Hinan
Port, hand net, 18 Sept. 2015; KAUM–I. 78531, 21.4 mm SL,
Umabana, hand net, 22 Sept. 2015.

Fig. 122. Plectranthias longimanus チビハナダイ (KAUM–I. 78365, 23.5mm SL).

28

Fig. 125. Pseudanthias dispar アカネハナゴイ (upper: KAUM–I. 78338,
44.7 mm SL; lower: Umabana, K. Koeda).

Checklist of fishes of Yonaguni-jima island
Pseudanthias pascalus (Jordan and Tanaka, 1927)
ハナゴイ [Marine] (Fig. 126)

Serranocirrhitus latus Watanabe, 1949
ハナゴンベ [Marine] (Fig. 129)
KAUM–I. 78311, 67.7 mm SL, Anchor's graves, spear, 17
Sept. 2015.

Fig. 129. Serranocirrhitus latus ハナゴンベ (KAUM–I. 78311, 67.7 mm SL).

Fig. 126. Pseudanthias pascalus ハナゴイ (Umabana, K. Koeda).

Pseudanthias squamipinnis (Peters, 1855)
キンギョハナダイ [Marine] (Fig. 127)
FRLM 51473, 46.5 mm SL, Umabana, hand net, 22 Sept.
2015; KAUM–I. 78335, 20.4 mm SL, KAUM–I. 78336, 41.8mm SL, off Kubura Hinan Port, hand net, 18 Sept. 2015.

Variola albimarginata Baissac, 1953
オジロバラハタ [Marine] (Fig. 130)
あがにーばいFRLM 52010, 208.0 mm SL, FRLM 52085, 161.6 mm
SL, off Hikawa, line-fishing, 21 Sept. 2015; KAUM–I. 79748,
216.8 mm SL, Yonaguni-jima island, line-fishing, 18 Sept.
2015; KAUM–I. 80421, 155.1 mm SL, off Hikawa, linefishing, 21 Sept. 2015; MUFS 46976, 164.0 mm SL, off
Hikawa, line-fishing, 21 Sept. 2015.

Fig. 130. Variola albimarginata オジロバラハタ (KAUM–I. 79748, 216.8mm SL).
Fig. 127. Pseudanthias squamipinnis キ ン ギ ョ ハ ナ ダ イ (KAUM–I.
78336, 41.8 mm SL).

Pseudogramma polyacantha (Bleeker, 1856)
トゲメギス [Marine] (Fig. 128)
KAUM–I. 78321, 37.2 mm SL, Rakuda Rock, hand net,
17 Sept. 2015; KAUM–I. 78347, 35.7 mm SL, KAUM–
I. 78348, 26.5 mm SL, KAUM–I. 78357, 31.2 mm SL, off
Kubura Hinan Port, hand net, 18 Sept. 2015.

Fig. 128. Pseudogramma polyacantha トゲメギス (KAUM–I. 78347, 35.7mm SL).

Variola louti (Forsskål, 1775)
バラハタ [Marine] (Fig. 131)
あがにーばいFRLM 52087, 149.4 mm SL, FRLM 52096, 163.3 mm
SL, off Hikawa, line-fishing, 21 Sept. 2015; KAUM–I. 78603,
278.2 mm SL, Kubura Fishing Port, line-fishing, 21 Sept.
2015; MUFS 46905, 228.0 mm SL, off Kubura Hinan Port,line-fishing, 18 Sept. 2015.

Fig. 131. Variola louti バラハタ (KAUM–I. 78603, 278.2 mm SL).

29

K. Koeda et al.
FAMILY PSEUDOCHROMIDAE
メギス科
Amsichthys knighti (Allen, 1987)
オオメタナバタメギス [Marine] (Fig. 132)
KAUM–I. 78526, 19.7 mm SL, Danno-dropoff, hand net,
22 Sept. 2015.

Fig. 132. Amsichthys knighti オオメタナバタメギス (KAUM–I. 78526,
19.7 mm SL).

Cypho zaps Gill, 2004
ドナンメギス [Marine] (Fig. 133)
Gill (2004); Yoshida et al. (in press).
KAUM–I. 78334, 35.6 mm SL, male, Rakuda Rock, handnet, 17 Sept. 2015; KAUM–I. 78364, 38.2 mm SL, male,off Kubura Hinan Port, hand net, 18 Sept. 2015; KAUM–I.
78543, 31.6 mm SL, male, Umabana, hand net, 22 Sept. 2015.
Remarks. The species was reported by Yoshida et al. (inpress) as the first Japanese record on the basis of collected
specimens on the present expedition.

Labracinus cyclophthalmus (Müller and Troschel, 1849)
メギス [Marine] (Fig. 134)
FRLM 51525, 131.4 mm SL, Hikawa, line-fishing, 23
Sept. 2015; FRLM 51570, 86.6 mm SL, Kubura Fishing
Port, line-fishing, 24 Sept. 2015; FRLM 51576, 97.0 mm
SL, FRLM 51577, 87.1 mm SL, FRLM 51578, 106.9 mm
SL, Hikawa, line-fishing, 23 Sept. 2015; FRLM 52024, 68.6mm SL, FRLM 52025, 45.5 mm SL, Kubura Fishing Port,line-fishing, 19 Sept. 2015; KAUM–I. 78281, 105.0 mm SL,
Kubura Fishing Port, line-fishing, 16 Sept. 2015; KAUM–
I. 78577, 76.0 mm SL, Hikawa, hand net, 20 Sept. 2015;
KAUM–I. 80269, 104.2 mm SL, Kubura Fishing Port, linefishing, 19 Sept. 2015; MUFS 46886, 83.0 mm SL, Kubura
Fishing Port, line-fishing, 17 Sept. 2015; MUFS 46916, 93.0mm SL, MUFS 46917, 106.0 mm SL, MUFS 46922, 115.0mm SL, levee at Kubura, line-fishing, 18 Sept. 2015; NSMT-P
55461, 3 specimens, 66.2–93.1 mm SL, Agarizaki, hand net,
28 Mar. 1997; NSMT-P 101233, 85.9 mm SL, Kubura Hinan
Port, line-fishing, 16 Mar. 2010.

Fig. 134. Labracinus cyclophthalmus メ ギ ス (upper: KAUM–I. 78577,
76.0 mm SL; lower: KAUM–I. 78281, 105.0 mm SL).

Pseudochromis cyanotaenia Bleeker, 1857
リュウキュウニセスズメ [Marine] (Fig. 135)
Yoshigou et al. (2001).
HMNH-P117, 21.3 mm SL, Agarizaki, hand net, 9 Aug.
1999; KAUM–I. 78294, 34.8 mm SL, Kubura Fishing Port,
hand net, 16 Sept. 2015.

Fig. 133. Cypho zaps ドナンメギス (upper: KAUM–I. 78334, 35.6 mm SL;
middle: KAUM–I. 78364, 38.2 mm SL; lower: Diving Service MARLIN).

30

Fig. 135. Pseudochromis cyanotaenia リュウキュウニセスズメ (KAUM–
I. 78294, 34.8 mm SL).

Checklist of fishes of Yonaguni-jima island
Pseudochromis luteus Aoyagi, 1943
コガネニセスズメ [Marine] (Fig. 136)

Fig. 136. Pseudochromis luteus コ ガ ネ ニ セ ス ズ メ (Diving Service
MARLIN).

FAMILY PRIACANTHIDAE
キントキダイ科
Heteropriacanthus cruentatus (Lacepède, 1801)
ゴマヒレキントキ [Marine] (Fig. 139)
FRLM 50711, 116.9 mm SL, Kubura Fishing Port, linefishing, 13 Mar. 2015.

FAMILY PLESIOPIDAE
タナバタウオ科
Acanthoplesiops psilogaster Hardy, 1985
フチドリタナバタウオ [Marine]
NSMT-P 77369, 6.4 mm SL, Agarizaki, hand net, 28 Mar. 1997.
Plesiops coeruleolineatus Rüppell, 1835
タナバタウオ [Marine] (Fig. 137)
KAUM–I. 78463, 26.1 mm SL, Sonai Channel, hand net,
21 Sept. 2015.

Fig. 139. Heteropriacanthus cruentatus ゴマヒレキントキ (FRLM 50711,
116.9 mm SL).

Priacanthus hamrur (Forsskål, 1775)
ホウセキキントキ [Marine] (Fig. 140)
Ito (2009).
KAUM–I. 80210, 231.0 mm SL, Arakawa Bay, spear, 18
Sept. 2015; MUFS 46890, 225.0 mm SL, Rakuda Rock, spear,
17 Sept. 2015.

Fig. 137. Plesiops coeruleolineatus タナバタウオ (KAUM–I. 78463, 26.1mm SL).

Plesiops verecundus Mooi, 1995
アマミタナバタウオ [Marine]
NSMT-P 55489, 40.1 mm SL, Agarizaki, hand net, 28
Mar. 1997.
FAMILY OPISTOGNATHIDAE
アゴアマダイ科
Opistognathus variabilis Smith-Vaniz, 2009
シシダマオオクチアマダイ [Marine] (Fig. 138)
Hibino et al. (2016).
FRLM 51432, 103.2 mm SL, male, Sonai Fishing Port,line-fishing, 20 Sept. 2015.
Remarks. The species was reported by Hibino et al. (2016)as the first Japanese record on the basis of the collected
specimen by the present expedition.

Fig. 138. Opistognathus variabilis シシダマオオクチアマダイ (FRLM
514322, 103.2 mm SL).

Fig. 140. Priacanthus hamrur ホ ウ セ キ キ ン ト キ (MUFS 46890, 225.0mm SL).

FAMILY APOGONIDAE
テンジクダイ科
Apogon sp.
コミナトテンジクダイ属の一種 [Marine] (Fig. 141)
KAUM–I. 78535, 26.7 mm SL, Umabana, hand net, 22
Sept. 2015.

Fig. 141. Apogon sp. (KAUM–I. 78535, 26.7 mm SL).

31

K. Koeda et al.
Apogonichthys ocellatus (Weber, 1913)
マトシボリ [Marine] (Fig. 142)
Yoshigou et al. (2001).
HMNH-P118, 14.8 mm SL, Agarizaki, hand net, 9 Aug. 1999.

Cheilodipterus quinquelineatus Cuvier, 1828
ヤライイシモチ [Marine] (Fig. 145)
FRLM 51423, 91.7 mm SL, Kubura Fishing Port, linefishing, 19 Sept. 2015; KAUM–I. 78259, 44.7 mm SL, Kubura
Fishing Port, hand net, 16 Sept. 2015; KAUM–I. 78492, 59.6mm SL, Sonai channel, hand net, 21 Sept. 2015.

Fig. 142. Apogonichthys ocellatus マトシボリ (HMNH-P118, 14.8 mm SL,
preserved specimen).

Cheilodipterus artus Smith, 1961
カスミヤライイシモチ [Marine] (Fig. 143)
FRLM 50710, 104.4 mm SL, Kubura Fishing Port, linefishing, 13 Mar. 2015.

Fig. 145. Cheilodipterus quinquelineatus ヤライイシモチ (upper: KAUM–
I. 78259, 44.7 mm SL; lower: KAUM–I. 78492, 59.6 mm SL).

Fig. 143. Cheilodipterus artus カ ス ミ ヤ ラ イ イ シ モ チ (FRLM 50710,
104.4 mm SL).

Fowleria variegata (Valenciennes, 1832)
シボリ [Marine] (Fig. 146)
KAUM–I. 78510, 40.0 mm SL, Hikawa, hand net, 21
Sept. 2015.

Cheilodipterus macrodon (Lacepède, 1802)
リュウキュウヤライイシモチ [Marine] (Fig. 144)
FRLM 51438, 132.5 mm SL, Sonai Fishing Port, linefishing, 20 Sept. 2015; KAUM–I. 78552, 97.2 mm SL,
Umabana, spear, 22 Sept. 2015.

Fig. 146. Fowleria variegata シボリ (KAUM–I. 78510, 40.0 mm SL).

Nectamia bandanensis (Bleeker, 1854)
バンダイシモチ [Marine] (Fig. 147)
FRLM 51440, 75.5 mm SL, Sonai Fishing Port, linefishing, 20 Sept. 2015.

Fig. 144. Cheilodipterus macrodon リュウキュウヤライイシモチ (upper:
KAUM–I. 78552, 97.2 mm SL; lower: Umabana, K. Koeda).

32

Fig. 147. Nectamia bandanensis バンダイシモチ (FRLM 51440, 75.5 mm SL).

Checklist of fishes of Yonaguni-jima island
Ostorhinchus angustatus (Smith and Radcliffe, 1911)
ウスジマイシモチ [Marine] (Fig. 148)
KAUM–I. 78340, 58.3 mm SL, KAUM–I. 78341, 50.9mm SL, KAUM–I. 78342, 44.3 mm SL, off Kubura Hinan
Port, hand net, 18 Sept. 2015; KAUM–I. 78547, 39.6 mm SL,
Danno-dropoff, hand net, 22 Sept. 2015; KAUM–I. 80387,
42.5 mm SL, Umabana, hand net, 22 Sept. 2015.

Fig. 150. Ostorhinchus ishigakiensis ミヤコイシモチ (upper: KAUM–I.
78282, 30.2 mm SL; lower: KAUM–I. 78283, 35.0 mm SL).

Fig. 148. Ostorhinchus angustatus ウスジマイシモチ (upper: KAUM–I.
78341, 50.9 mm SL; lower: KAUM–I. 78340, 58.3 mm SL).

Ostorhinchus apogonides (Bleeker, 1856)
アオハナテンジクダイ [Marine]
Ito (2009).

Ostorhinchus nigrofasciatus (Lachner, 1953)
ミナミフトスジイシモチ [Marine] (Fig. 151)
Ito (2009).
FRLM 50725, 62.9 mm SL, Kubura Fishing Port, linefishing, 12 Mar. 2015; FRLM 51422, 68.7 mm SL, Kubura
Fishing Port, line-fishing, 19 Sept. 2015; KAUM–I. 78534,
46.0 mm SL, Umabana, hand net, 22 Sept. 2015; KAUM–I.
80238, 67.7 mm SL, Sonai Fishing Port, line-fishing, 20 Sept.
2015.

Ostorhinchus cookii (Macleay, 1881)
スジイシモチ [Marine] (Fig. 149)
Yoshigou et al. (2001).
HMNH-P131, 46.8 mm SL, Sonai, hand net, 9 Aug. 1999.

Fig. 151. Ostorhinchus nigrofasciatus ミナミフトスジイシモチ (KAUM–
I. 78534, 46.0 mm SL).
Fig. 149. Ostorhinchus cookii スジイシモチ (HMNH-P131, 46.8 mm SL,
preserved specimen).

Ostorhinchus ishigakiensis (Ida and Moyer, 1974)
ミヤコイシモチ [Marine; brackish] (Fig. 150)
FRLM 51521, 22.6 mm SL, mouth of Kubura River, handnet, 23 Sept. 2015; FRLM 52145, 26.0 mm SL, FRLM 52146,
26.8 mm SL, Kubura Fishing Port, hand net, 24 Sept. 2015;
KAUM–I. 78282, 30.2 mm SL, KAUM–I. 78283, 35.0 mm
SL, Kubura Fishing Port, hand net, 16 Sept. 2015; KAUM–I.
78511, 38.6 mm SL, Hikawa, hand net, 21 Sept. 2015.

Ostorhinchus novemfasciatus (Cuvier, 1828)
タスジイシモチ [Marine] (Fig. 152)
Yoshigou et al. (2001); Ito (2009).
HMNH-P114, 3 specimens, 22.1–33.0 mm SL, Agarizaki,
hand net, 9 Aug. 1999; KAUM–I. 78284, 43.0 mm SL,
Kubura Fishing Port, hand net, 16 Sept. 2015; KAUM–I.
78483, 42.2 mm SL, KAUM–I. 78484, 34.5 mm SL, KAUM–I.
78485, 22.2 mm SL, Sonai channel, hand net, 21 Sept. 2015;
KAUM–I. 78499, 49.2 mm SL, Hikawa, hand net, 21 Sept.
2015.

33

K. Koeda et al.
Pristiapogon fraenatus (Valenciennes, 1832)
ヒトスジイシモチ [Marine]
Ito (2009).
Pristiapogon kallopterus (Bleeker, 1856)
カスリイシモチ [Marine] (Fig. 155)
FRLM 50708, 90.8 mm SL, Kubura Fishing Port, linefishing, 13 Mar. 2015; FRLM 52055, 114.6 mm SL, Sonai
Fishing Port, line-fishing, 24 Sept. 2015; KAUM–I. 78304,
83.7 mm SL, Kubura Fishing Port, line-fishing, 16 Sept. 2015;
KAUM–I. 78491, 98.4 mm SL, Sonai channel, hand net, 21
Sept. 2015; KAUM–I. 78527, 32.7 mm SL, KAUM–I. 78528,
27.1 mm SL, KAUM–I. 80458, 91.1 mm SL, KAUM–I.
80459, 95.0 mm SL, KAUM–I. 80460, 98.1 mm SL, KAUM–
I. 80461, 101.9 mm SL, Umabana, hand net, 22 Sept. 2015;
KAUM–I. 80235, 105.9 mm SL, KAUM–I. 80236, 71.2 mm
SL, KAUM–I. 80237, 89.4 mm SL, Sonai Fishing Port, linefishing, 20 Sept. 2015.
Fig. 152. Ostorhinchus novemfasciatus タスジイシモチ (upper: KAUM–I.
78284, 43.0 mm SL; lower: KAUM–I. 78483, 42.2 mm SL).

Ostorhinchus taeniophorus (Regan, 1908)
ミスジテンジクダイ [Marine] (Fig. 153)
FRLM 51528, 47.4 mm SL, Sonai Fishing Port, linefishing, 24 Sept. 2015; NSMT-P 55460, 9 specimens, 51.9–
74.7 mm SL, Agarizaki, hand net, 28 Mar. 1997.

Fig. 153. Ostorhinchus taeniophorus ミスジテンジクダイ (FRLM 51528,
47.4 mm SL).

Pristiapogon exostigma (Jordan and Starks, 1906)
ユカタイシモチ [Marine] (Fig. 154)
FRLM 50709, 92.0 mm SL, Kubura Fishing Port, linefishing, 13 Mar. 2015; FRLM 50768, 95.6 mm SL, Sonai Fishing
Port, line-fishing, 12 Mar. 2015; FRLM 51439, 88.5 mm SL,
Sonai Fishing Port, line-fishing, 20 Sept. 2015; KAUM–I. 80239,
92.2 mm SL, Sonai Fishing Port, line-fishing, 20 Sept. 2015.

Fig. 154. Pristiapogon exostigma ユカタイシモチ (KAUM–I. 80239, 92.2mm SL).

34

Fig. 155. Pristiapogon kallopterus カスリイシモチ (upper: FRLM 50708,
90.8 mm SL; lower: KAUM–I. 78527, 32.7 mm SL).

Pristicon trimaculatus (Cuvier, 1828)
ミスジアカヒレイシモチ [Marine] (Fig. 156)
KAUM–I. 78303, 112.0 mm SL, Kubura Fishing Port,line-fishing, 16 Sept. 2015.

Fig. 156. Pristicon trimaculatus ミ ス ジ ア カ ヒ レ イ シ モ チ (KAUM–I.
78303, 112.0 mm SL).

Checklist of fishes of Yonaguni-jima island
Sphaeramia orbicularis (Cuvier, 1828)
ホソスジマンジュウイシモチ [Marine] (Fig. 157)
KAUM–I. 78288, 43.0 mm SL, KAUM–I. 78289, 45.9mm SL, KAUM–I. 78290, 29.8 mm SL, KAUM–I. 78291,
25.4 mm SL, KAUM–I. 78292, 26.4 mm SL, KAUM–I.
78295, 26.9 mm SL, KAUM–I. 78296, 27.0 mm SL, KAUM–I.
78297, 28.4 mm SL, Kubura Fishing Port, hand net, 16 Sept.
2015.

FAMILY MALACANTHIDAE
キツネアマダイ科
Hoplolatilus starcki Randall and Dooley, 1974
サンゴアマダイ属の一種 [Marine] (Fig. 159)

Fig. 159. Hoplolatilus starcki (off Kubura, Diving Service MARLIN).

Fig. 157. Sphaeramia orbicularis ホソスジマンジュウイシモチ (upper:
KAUM–I. 78288, 43.0 mm SL; lower: KAUM–I. 78292, 26.4 mm SL).

Malacanthus brevirostris Guichenot, 1848
ヤセアマダイ [Marine] (Fig. 160)
Ito (2009).
FRLM 52082, 180.5 mm SL, FRLM 52104, 150.8 mm
SL, off Hikawa, line-fishing, 21 Sept. 2015; KAUM–I.
78470, 168.5 mm SL, off Hikawa, line-fishing, 21 Sept. 2015;
KAUM–I. 80383, 191.2 mm SL, KAUM–I. 80384, 184.9 mm
SL, Umabana, line-fishing, 22 Sept. 2015; KAUM–I. 80451,
170.9 mm SL, KAUM–I. 80452, 183.3 mm SL, KAUM–I.
80453, 175.0 mm SL, KAUM–I. 80454, 152.2 mm SL, northof Yonaguni-jima island, 22 Sept. 2015; KAUM–I. 80486,
171.1 mm SL, KAUM–I. 80487, 159.2 mm SL, off Sonai,
line-fishing, 22 Sept. 2015; KAUM–I. 80491, 151.0 mm SL,
north of Yonaguni-jima island, 22 Sept. 2015; MUFS 46979,
174.0 mm SL, off Hikawa, line-fishing, 21 Sept. 2015.

Yarica hyalosoma (Bleeker, 1852)
カガミテンジクダイ [Brackish] (Fig. 158)
Yoshigou (2000); Yoshigou et al. (2001); Ministry of theEnvironment (2010a).
HMNH-P104, 83.3 mm SL, Tabaru River, hand net, 10Aug. 1999.

Fig. 158. Yarica hyalosoma カガミテンジクダイ (HMNH-P104, 83.3 mm SL).
Fig. 160. Malacanthus brevirostris ヤセアマダイ (upper to lower: FRLM
52082, 180.5 mm SL; KAUM–I. 78470, 168.5 mm SL; KAUM–I.
80384, 184.9 mm SL; MUFS 46979, 174.0 mm SL).

35

K. Koeda et al.
FAMILY CORYPHAENIDAE
シイラ科
Coryphaena hippurus Linnaeus, 1758
シイラ [Marine] (Fig. 161)
Ito (2009).
KAUM–I. 78609, 566.5 mm SL, Yonaguni-jima island(purchaced at Kubura Fish Landing Port), line-fishing, 23
Sept. 2015.

Carangoides orthogrammus (Jordan and Gilbert, 1882)
ナンヨウカイワリ [Marine] (Fig. 163)
FRLM 51494, 216.0 mm SL, mouth of Tabaru River, gill
net, 23 Sept. 2015; FRLM 52015, 210.5 mm SL, Kubura
Hinan Port, 20 Sept. 2015; KAUM–I. 80483, 244.7 mm SL,
off Sonai, line-fishing, 22 Sept. 2015.

Fig. 161. Coryphaena hippurus シイラ (KAUM–I. 78609, 566.5 mm SL).

FAMILY RACHYCENTRIDAE
スギ科
Rachycentron canadum (Linnaeus, 1766)
スギ [Marine]
Ito (2009).
FAMILY ECHENEIDAE
コバンザメ科
Remora osteochir (Cuvier, 1829)
ヒシコバン [Marine]
URM-P 42214, off Yonaguni-jima island.
Remora remora (Linnaeus, 1758)
ナガコバン [Marine]
NSMT-P 101229, 30.0 mm SL, off Irizaki, collected frombody of Istiophorus platypterus, 17 Mar. 2010.
Remora sp.
ナガコバン属の一種 [Marine]
URM-P 42130, off Yonaguni-jima island.

Fig. 163. Carangoides orthogrammus ナンヨウカイワリ (upper: FRLM
51494, 216.0 mm SL; lower: KAUM–I. 80483, 244.7 mm SL).

Carangoides plagiotaenia Bleeker, 1857
インドカイワリ [Marine]
Ito (2009).
Caranx ignobilis (Forsskål, 1775)
ロウニンアジ [Marine; brackish] (Fig. 164)
がーらいゆ
FRLM 52100, 404.4 mm SL, Sonai Fishing Port, linefishing, 21 Sept. 2015.

FAMILY CARANGIDAE
アジ科
Alectis ciliaris (Bloch, 1787)
イトヒキアジ [Marine] (Fig. 162)

Fig. 162. Alectis ciliaris イトヒキアジ (Diving Service MARLIN).

36

Fig. 164. Caranx ignobilis ロウニンアジ (upper: FRLM 52100, 404.4 mm
SL; lower: Diving Service MARLIN).

Checklist of fishes of Yonaguni-jima island
Caranx lugubris Poey, 1860
カッポレ [Marine] (Fig. 165)
がーらいゆ

Caranx papuensis Alleyne and Macleay, 1877
オニヒラアジ [Marine; brackish] (Fig. 167)
がーらいゆ
Yoshigou (2000); Yoshigou et al. (2001).
HMNH-P75, 55.7 mm SL, HMNH-P98, 46.8 mm SL,
Tabaru River, hand net, 9 Aug. 1999.

Fig. 165. Caranx lugubris カッポレ (Diving Service MARLIN).

Caranx melampygus Cuvier, 1833
カスミアジ [Marine; brackish] (Fig. 166)
がーらいゆ
FRLM 51467, 100.5 mm SL, mouth of Kubura River,
cast net, 21 Sept. 2015; HMNH-P97, 134.8 mm SL, Kuburamit, hand net, 10 Aug. 1999; KAUM–I. 78352, 289.1 mm
SL, Arakawa Bay, line-fishing, 18 Sept. 2015; MUFS 47019,
288.0 mm SL, Kubura-bari, line-fishing, 23 Sept. 2015;
NSMT-P 101247, 422.4 mm SL, Danno Beach, 19 Mar. 2010.

Fig. 166. Caranx melampygus カスミアジ (upper: KAUM–I. 78352, 289.1mm SL; middle: MUFS 47019, 288.0 mm SL; lower: Diving Service
MARLIN).

Fig. 167. Caranx papuensis オニヒラアジ (HMNH-P98, 46.8 mm SL).

Caranx sexfasciatus Quoy and Gaimard, 1825
ギンガメアジ [Marine; brackish] (Fig. 168)
がーらいゆ
Yoshigou (2000).
FRLM 52079, 300.6 mm SL, Sonai Fishing Port, linefishing, 19 Sept. 2015.

Fig. 168. Caranx sexfasciatus ギンガメアジ (upper: FRLM 52079, 300.6mm SL; middle: Diving Service MARLIN; lower: K. Koeda).

37

K. Koeda et al.
Elagatis bipinnulata (Quoy and Gaimard, 1825)
ツムブリ [Marine] (Fig. 169)
ながいゆ
FRLM 52108, 554.0 mm SL, Yonaguni-jima island(purchaced at Kubura Fish Landing Port), line-fishing,
25 Sept. 2015; KAUM–I. 79710, 660.4 mm SL, south of
Yonaguni-jima island, line-fishing, 18 Sept. 2015.

Seriola dumerili (Risso, 1810)
カンパチ [Marine] (Fig. 172)

Fig. 172. Seriola dumerili カ ン パ チ (fish landing port at Yonaguni-jimaisland, K. Koeda).

Trachinotus baillonii (Lacepède, 1801)
コバンアジ [Marine; brackish] (Fig. 173)
FRLM 51485, 46.0 mm SL, Sonai Fishing Port, cast net,
22 Sept. 2015.
Fig. 169. Elagatis bipinnulata ツ ム ブ リ (upper: KAUM–I. 79710, 660.4mm SL; lower: Diving Service MARLIN).

Naucrates ductor (Linnaeus, 1758)
ブリモドキ [Marine] (Fig. 170)

Fig. 173. Trachinotus baillonii コバンアジ (FRLM 51485, 46.0 mm SL).

Trachinotus blochii (Lacepède, 1801)
マルコバン [Marine; brackish] (Fig. 174)
Yoshigou (2000); Yoshigou et al. (2001).
HMNH-P88, 78.9 mm SL, Tabaru River, hand net, 10Aug. 1999; HMNH-P106, 5 specimens, 45.9–54.8 mm SL,
Hikawa, hand net, 9 Aug. 1999.
Fig. 170. Naucrates ductor ブリモドキ (Diving Service MARLIN).

Scomberoides lysan (Forsskål, 1775)
イケカツオ [Marine]
Ito (2009).
Selar crumenophthalmus (Bloch, 1793)
メアジ [Marine] (Fig. 171)
FRLM 51500, 109.0 mm SL, off Kubura Fishing Port
(24°27ʹ25ʺN, 122°56ʹ17ʺE), hand, 22 Sept. 2015; NSMT-P
101118, 163.7 mm SL, Kubura Fishing Port, 10 Mar. 2010.

Fig. 171. Selar crumenophthalmus メアジ (FRLM 51500, 109.0 mm SL).

38

Fig. 174. Trachinotus blochii マ ル コ バ ン (HMNH-P88, 78.9 mm SL,
preserved specimen).

Checklist of fishes of Yonaguni-jima island
FAMILY BRAMIDAE
シマガツオ科
Eumegistus illustris Jordan and Jordan, 1922
チカメエチオピア [Marine] (Fig. 175)
FRLM 50769, 477.0 mm SL, north of Yonaguni-jimaisland, line-fishing, 16 Mar. 2015.

Etelis coruscans Valenciennes, 1862
ハマダイ [Marine]
あがいゆFujimoto (1972); Ikema (2003); Fukuda and Ebisawa(2001).
Lutjanus argentimaculatus (Forsskål, 1775)
ゴマフエダイ [Marine; brackish] (Fig. 178)
Yoshigou (2000); Yoshigou et al. (2001).
HMNH-P71, 33.3 mm SL, Tabaru River, hand net, 9 Aug.
1999.

Fig. 175. Eumegistus illustris チカメエチオピア (FRLM 50769, 477.0 mm SL).

Fig. 178. Lutjanus argentimaculatus ゴマフエダイ (HMNH-P71, 33.3 mm SL,
preserved specimen).

FAMILY LUTJANIDAE
フエダイ科
Aphareus furca (Lacepède, 1801)
イシフエダイ [Marine] (Fig. 176)
KAUM–I. 78314, 218.0 mm SL, off Hinan Port, linefishing, 17 Sept. 2015; MUFS 46903, 230.0 mm SL, MUFS
46904, 270.0 mm SL, off Kubura Hinan Port, line-fishing, 18
Sept. 2015.

Lutjanus bohar (Forsskål, 1775)
バラフエダイ [Marine] (Fig. 179)
あがな
Ikema (2003).
FRLM 51502, 176.4 mm SL, off Northern Pasture, linefishing, 22 Sept. 2015; KAUM–I. 78604, 308.7 mm SL,
Kubura Fishing Port, line-fishing, 21 Sept. 2015; MUFS
46902, 184.0 mm SL, off Kubura Hinan Port, line-fishing,18
Sept. 2015; MUFS 47003, 138.0 mm SL, off Northern
Pasture, line-fishing, 22 Sept. 2015.

Fig. 176. Aphareus furca イシフエダイ (KAUM–I. 78314, 218.0 mm SL).

Aprion virescens Valenciennes, 1830
アオチビキ [Marine] (Fig. 177)
KAUM–I. 80276, 224.6 mm SL, Sonai Fishing Port, linefishing, 20 Sept. 2015; MUFS 46938, 310.0 mm SL, Sonai
Fishing Port, line-fishing, 19 Sept. 2015.

Fig. 177. Aprion virescens アオチビキ (MUFS 46938, 310.0 mm SL).

Fig. 179. Lutjanus bohar バ ラ フ エ ダ イ (upper: KAUM–I. 78604, 308.7mm SL; lower: MUFS 47003, 138.0 mm SL).

39

K. Koeda et al.
Lutjanus decussatus (Cuvier, 1828)
アミメフエダイ [Marine] (Fig. 180)
MUFS 46951, 95.0 mm SL, Sonai Fishing Port, linefishing, 20 Sept. 2015.

Fig. 180. Lutjanus decussatus アミメフエダイ (MUFS 46951, 95.0 mm SL).

Lutjanus fulvus (Forster, 1801)
オキフエダイ [Marine; brackish] (Fig. 182)
Yoshigou (2000); Yoshigou et al. (2001).
FRLM 50713, 190.5 mm SL, Kubura Fishing Port, linefishing, 13 Mar. 2015; HMNH-P70, 25.7 mm SL, HMNH-P
74, 55.3 mm SL, HMNH-P77, 63.6 mm SL, Tabaru River,
hand net, 9 Aug. 1999; KAUM–I. 78264, 41.4 mm SL,
KAUM–I. 78265, 25.2 mm SL, Kubura Fishing Port, handnet, 16 Sept. 2015; KAUM–I. 78496, 48.2 mm SL, Hikawa,
hand net, 21 Sept. 2015; MUFS 46878, 91.0 mm SL, Kubura
Fishing Port, line-fishing, 16 Sept. 2015; MUFS 46918, 99.0mm SL, MUFS 46919, 61.0 mm SL, Kubura Fishing Port,line-fishing, 18 Sept. 2015; MUFS 46962, 146.0 mm SL,
Kataburu Beach, line-fishing, 20 Sept. 2015; NSMT-P 101120,
132.0 mm SL, NSMT-P 101121, 127.6 mm SL, Kubura
Fishing Port, 10 Mar. 2010.

Lutjanus fulviflamma (Forsskål, 1775)
ニセクロホシフエダイ [Marine; brackish] (Fig. 181)
Yoshigou (2000); Yoshigou et al. (2001).
HMNH-P78, 46.4 mm SL, Kubura River, hand net, 9 Aug.
1999; MUFS 47028, 69.0 mm SL, mouth of Kubura River,line-fishing, 24 Sept. 2015.

Fig. 181. Lutjanus fulviflamma ニ セ ク ロ ホ シ フ エ ダ イ (MUFS 47028,
69.0 mm SL).

Fig. 182. Lutjanus fulvus オキフエダイ (upper: KAUM–I. 78264, 41.4 mm
SL; lower: MUFS 46878, 91.0 mm SL).

Lutjanus gibbus (Forsskål, 1775)
ヒメフエダイ [Marine] (Fig. 183)
FRLM 50714, 143.0 mm SL, Kubura Fishing Port, linefishing, 13 Mar. 2015; FRLM 50731, 109.7 mm SL, FRLM
50732, 100.6 mm SL, Kubura Fishing Port, line-fishing,
12 Mar. 2015; FRLM 51950, 152.0 mm SL, FRLM 51951,
163.2 mm SL, FRLM 51997, 85.9 mm SL, FRLM 51998,
57.7 mm SL, FRLM 51999, 90.7 mm SL, FRLM 52000, 84.3mm SL, FRLM 52001, 83.5 mm SL, FRLM 52002, 89.4 mm
SL, FRLM 52003, 88.3 mm SL, FRLM 52004, 57.8 mm SL,
FRLM 52005, 88.5 mm SL, Sonai Fishing Port, line-fishing,
19 Sept. 2015; FRLM 52026, 85.4 mm SL, Kubura Fishing
Port, line-fishing, 19 Sept. 2015; FRLM 52027, 74.2+ mm SL,
Kubura Fishing Port, line-fishing, damaged, 19 Sept. 2015;
KAUM–I. 78293, 32.4 mm SL, Kubura Fishing Port, handnet, 16 Sept. 2015; KAUM–I. 78307, 155.8 mm SL, Kataburu
Beach, line-fishing, 17 Sept. 2015; MUFS 46882, 94.0 mm
SL, MUFS 46885, 89.0 mm SL, Kubura Fishing Port, linefishing, 17 Sept. 2015; MUFS 46928, 92.0 mm SL, Sonai
Fishing Port, line-fishing, 18 Sept. 2015.

40

Checklist of fishes of Yonaguni-jima island

Fig. 184. Lutjanus kasmira ヨスジフエダイ (upper: FRLM 51947, 106.6mm SL; lower: MUFS 47001, 122.0 mm SL).

Lutjanus monostigma (Cuvier, 1828)
イッテンフエダイ [Marine; brackish] (Fig. 185)
Yoshigou (2000); Yoshigou et al. (2001).
FRLM 51455, 173.0 mm SL, Sonai Fishing Port, linefishing, 19 Sept. 2015; HMNH-P79, 57.8 mm SL, Kubura
River, hand net, 9 Aug. 1999.

Fig. 183. Lutjanidae gibbus ヒ メ フ エ ダ イ (upper: FRLM 50714, 143.0mm SL; middle: MUFS 46885, 89.0 mm SL; lower: KAUM–I. 78293,
32.4 mm SL).

Lutjanus kasmira (Forsskål, 1775)
ヨスジフエダイ [Marine] (Fig. 184)
Ito (2009).
FRLM 50727, 130.2 mm SL, FRLM 50733, 106.9 mm
SL, Kubura Fishing Port, line-fishing, 12 Mar. 2015; FRLM
50766, 136.3 mm SL, Sonai Fishing Port, line-fishing, 12 Mar.
2015; FRLM 51947, 106.6 mm SL, Sonai Fishing Port, linefishing, 19 Sept. 2015; FRLM 52028, 72.2 mm SL, Kubura
Fishing Port, line-fishing, 19 Sept. 2015; KAUM–I. 80240,
63.1 mm SL, Sonai Fishing Port, line-fishing, 20 Sept. 2015;
KAUM–I. 80466, 209.5 mm SL, Kubura Fishing Port, linefishing, 20 Sept. 2015; MUFS 46927, 105.0 mm SL, Sonai
Fishing Port, line-fishing, 18 Sept. 2015; MUFS 47001, 122.0mm SL, off Northern Pasture, line-fishing, 22 Sept. 2015.

Fig. 185. Lutjanus monostigma イッテンフエダイ (FRLM 51455, 173.0mm SL).

Lutjanus rivulatus (Cuvier, 1828)
ナミフエダイ [Marine] (Fig. 186)
つばき
FRLM 52101, 423.0 mm SL, Kubura Fishing Port, linefishing, 21 Sept. 2015.

Fig. 186. Lutjanus rivulatus ナミフエダイ (FRLM 52101, 423.0 mm SL).

41

K. Koeda et al.
Macolor macularis Fowler, 1931
ホホスジタルミ [Marine] (Fig. 187)
MUFS 46939, 168.0 mm SL, Kubura Hinan Port, linefishing, 19 Sept. 2015.

Fig. 189. Paracaesio caerulea アオダイ (MUFS 47037, 316.0 mm SL).

Paracaesio sordida Abe and Shinohara, 1962
ヨゴレアオダイ [Marine] (Fig. 190)
KAUM–I. 78558, 279.4 mm SL, off Yonaguni Airport,line-fishing, 22 Sept. 2015.

Fig. 187. Macolor macularis ホホスジタルミ (upper: MUFS 46939, 168.0mm SL; lower: Diving Service MARLIN).

Macolor niger (Forsskål, 1775)
マダラタルミ [Marine] (Fig. 188)
KAUM–I. 78464, 29.6 mm SL, Sonai Channel, hand net,
21 Sept. 2015.

Fig. 190. Paracaesio sordida ヨゴレアオダイ (upper: KAUM–I. 78558,
279.4 mm SL; lower: Diving Service MARLIN).

Pristipomoides argyrogrammicus (Valenciennes, 1832)
ハナフエダイ [Marine] (Fig. 191)
FRLM 50748, 184.5 mm SL, northwest of Yonaguni-jimaisland, line-fishing, 16 Mar. 2015.

Fig. 188. Macolor niger マダラタルミ (KAUM–I. 78464, 29.6 mm SL).

Paracaesio caerulea (Katayama, 1934)
アオダイ [Marine] (Fig. 189)
Fukuda and Ebisawa (2001).
MUFS 47037, 316.0 mm SL, Yonaguni-jima island(purchaced at Kubura Fish Landing Port), Sept. 2015.

42

Fig. 191. Pristipomoides argyrogrammicus ハナフエダイ (FRLM 50748,
184.5 mm SL).

Checklist of fishes of Yonaguni-jima island
Pristipomoides filamentosus (Valenciennes, 1830)
オオヒメ [Marine] (Fig. 192)
Fukuda and Ebisawa (2001).
MUFS 47039, 365.0 mm SL, MUFS 47040, 315.0 mm
SL, Yonaguni-jima island (purchaced at Kubura Fish LandingPort), Sept. 2015.

FAMILY CAESIONIDAE
タカサゴ科
Caesio teres Seale, 1906
ウメイロモドキ [Marine] (Fig. 195)
FRLM 52058, 210.0 mm SL, off Northern Pasture, linefishing, 22 Sept. 2015; KAUM–I. 78490, 258.8 mm SL, off
Hikawa, line-fishing, 21 Sept. 2015; KAUM–I. 80488, 227.4mm SL, north of Yonaguni-jima island, line-fishing, 22 Sept.
2015; MUFS 46981, 257.0 mm SL, off Hikawa, line-fishing,
21 Sept. 2015; MUFS 47014, 241.0 mm SL, off Northern
Pasture, line-fishing, 22 Sept. 2015.

Fig. 192. Pristipomoides filamentosus オオヒメ (MUFS 47040, 315.0 mm SL).

Pristipomoides flavipinnis Shinohara, 1963
キンメヒメダイ [Marine] (Fig. 193)
MUFS 47038, 395.0 mm SL, Yonaguni-jima island(purchaced at Kubura Fish Landing Port), Sept. 2015.

Fig. 195. Caesio teres ウメイロモドキ (KAUM–I. 78490, 258.8 mm SL).

Pterocaesio marri Schultz, 1953
ニセタカサゴ [Marine] (Fig. 196)
ぐるくん
KAUM–I. 78488, 248.4 mm SL, off Hikawa, line-fishing,
21 Sept. 2015; MUFS 46977, 200.0 mm SL, off Hikawa, linefishing, 21 Sept. 2015.
Fig. 193. Pristipomoides flavipinnis キンメヒメダイ (MUFS 47038, 395.0mm SL).

Pristipomoides sieboldii (Bleeker, 1855)
ヒメダイ [Marine] (Fig. 194)
Fukuda and Ebisawa (2001).
FRLM 51564, 207.8 mm SL, Yonaguni-jima island(purchaced at Kubura Fish Landing Port), line-fishing, 25
Sept. 2015.

Fig. 196. Pterocaesio marri ニセタカサゴ (KAUM–I. 78488, 248.4 mm SL).

Pterocaesio tile (Cuvier, 1830)
クマササハナムロ [Marine] (Fig. 197)
ぐるくん
FRLM 51501, 213.0 mm SL, Kubura-bari, line-fishing,
22 Sept. 2015; KAUM–I. 80457, 222.7 mm SL, Kubura-bari,line-fishing, 22 Sept. 2015; MUFS 47004, 210.0 mm SL,
Kubura-bari, line-fishing, 22 Sept. 2015.
Fig. 194. Pristipomoides sieboldii ヒメダイ (FRLM 51564, 207.8 mm SL).

Fig. 197. Pterocaesio tile クマササハナムロ (KAUM–I. 80457, 222.7 mm SL).

43

K. Koeda et al.
FAMILY GERREIDAE
クロサギ科
Gerres filamentosus Cuvier, 1829
イトヒキサギ [Marine; brackish] (Fig. 198)
MUFS 46960, 191.0 mm SL, Kubura Fishing Port, gill
net, 20 Sept. 2015.

FAMILY HAEMULIDAE
イサキ科
Plectorhinchus gibbosus (Lacepède, 1802)
クロコショウダイ [Marine; brackish] (Fig. 200)

Fig. 200. Plectorhinchus gibbosus ク ロ コ シ ョ ウ ダ イ (Diving Service
MARLIN).

Fig. 198. Gerres filamentosus イトヒキサギ (MUFS 46960, 191.0 mm SL).

Plectorhinchus lessonii (Cuvier, 1830)
ヒレグロコショウダイ [Marine] (Fig. 201)
KAUM–I. 79713, 273.4 mm SL, Arakawa Bay, spear, 18
Sept. 2015.

Gerres longirostris (Lacepède, 1801)
ツッパリサギ [Marine; brackish]
URM-P 34392, Tabaru River, 5 Aug. 1995.
Gerres oblongus Cuvier, 1830
ナガサギ [Marine; brackish]
URM-P 34393, Tabaru River, 5 Aug. 1995.
Gerres oyena (Forsskål, 1775)
ミナミクロサギ [Marine; brackish] (Fig. 199)
Yoshigou (2000); Yoshigou et al. (2001).
FRLM 51549, 49.4 mm SL, mouth of Kubura River, castnet, 24 Sept. 2015; HMNH-P33, 73.3 mm SL, HMNH-P34,
62.0 mm SL, HMNH-P35, 42.8 mm SL, Kubura River, handnet, 8 Aug. 1999; HMNH-P76, 76.5 mm SL, Kubura River,
hand net, 9 Aug. 1999; HMNH-P84, 66.2 mm SL, HMNH-P
85, 64.3 mm SL, HMNH-P86, 60.6 mm SL, HMNH-P87, 69.7mm SL, Tabaru River, hand net, 10 Aug. 1999.

Fig. 201. Plectorhinchus lessonii ヒレグロコショウダイ (upper: KAUM–
I. 79713, 273.4 mm SL; lower: Diving Service MARLIN).

Plectorhinchus lineatus (Linnaeus, 1758)
アヤコショウダイ [Marine] (Fig. 202)

Fig. 199. Gerres oyena ミナミクロサギ (FRLM 51549, 49.4 mm SL).

Fig. 202. Plectorhinchus lineatus ア ヤ コ シ ョ ウ ダ イ (Diving Service
MARLIN).

44

Checklist of fishes of Yonaguni-jima island
Plectorhinchus picus (Cuvier, 1828)
アジアコショウダイ [Marine] (Fig. 203)

Fig. 205. Pentapodus aureofasciatus ヤクシマキツネウオ (upper: FRLM
51975, 116.5 mm SL; lower: MUFS 46988, 138.0 mm SL).

Fig. 203. Plectorhinchus picus ア ジ ア コ シ ョ ウ ダ イ (Diving Service
MARLIN).

Scolopsis bilineata (Bloch, 1793)
フタスジタマガシラ [Marine] (Fig. 206)
KAUM–I. 80391, 125.5 mm SL, Umabana, spear, 22 Sept.
2015; MUFS 47002, 144.0 mm SL, off Northern Pasture,line-fishing, 22 Sept. 2015; NSMT-P 101126, 153.7 mm SL,
Danno Beach, 20 Mar. 2010.

Plectorhinchus vittatus (Linnaeus, 1758)
ムスジコショウダイ [Marine] (Fig. 204)
Ito (2009).
MUFS 46887, Rakuda Rock, spear, 17 Sept. 2015;
KAUM–I. 80227, 216.1 mm SL, Rakuda Rock, spear, 17
Sept. 2015.

Fig. 206. Scolopsis bilineata フタスジタマガシラ (MUFS 47002, 144.0
mm SL).

Fig. 204. Plectorhinchus vittatus ムスジコショウダイ (KAUM–I. 80227,
216.1 mm SL).

Scolopsis lineata Quoy and Gaimard, 1824
ヨコシマタマガシラ [Marine] (Fig. 207)
Yoshigou et al. (2001); Ito (2009).
HMNH-P138, 33.8 mm SL, Sonai, hand net, 9 Aug. 1999;
KAUM–I. 80350, 112.3 mm SL, Sonai Channel, spear, 21
Sept. 2015.

FAMILY NEMIPTERIDAE
イトヨリダイ科
Pentapodus aureofasciatus Russell, 2001
ヤクシマキツネウオ [Marine] (Fig. 205)
FRLM 51975, 116.5 mm SL, FRLM 51976, 137.5 mm SL,
FRLM 52103, 138.0 mm SL, off Hikawa, line-fishing, 132.0
mm SL, 21 Sept. 2015; MUFS 46986, 129.0 mm SL, MUFS
46987, 121.0 mm SL, MUFS 46988, 138.0 mm SL, MUFS
46989, 143.0 mm SL, MUFS 46990, 131.0 mm SL, MUFS
46991, MUFS 46992, 132.0 mm SL, off Hikawa, line-fishing,
21 Sept. 2015.

Fig. 207. Scolopsis lineata ヨコシマタマガシラ (KAUM–I. 80350, 112.3mm SL).

45

K. Koeda et al.
FAMILY SPARIDAE
タイ科
Acanthopagrus sp.
クロダイ属の一種 [Marine; brackish]
くるいゆ
Ikema (2003).
FAMILY LETHRINIDAE
フエフキダイ科
Gnathodentex aureolineatus (Lacepède, 1802)
ノコギリダイ [Marine] (Fig. 208)
あがむてぃ
Ikema (2003); Ito (2009).
FRLM 50730, 100.7 mm SL, Kubura Fishing Port, linefishing, 12 Mar. 2015; FRLM 50767, 118.7 mm SL, Sonai
Fishing Port, line-fishing, 12 Mar. 2015; FRLM 51503, 189.5mm SL, off Northern Pasture, line-fishing, 22 Sept. 2015;
FRLM 52066, 149.8 mm SL, Kubura Fishing Port, line-fishing,
24 Sept. 2015; KAUM–I. 78489, 189.2 mm SL, off Hikawa,line-fishing, 21 Sept. 2015; KAUM–I. 78645, 180.4 mm SL,
Arakawa Bay, line-fishing, 18 Sept. 2015; KAUM–I. 80241,
144.2 mm SL, Sonai Fishing Port, line-fishing, 20 Sept. 2015;
KAUM–I. 80263, 168.7 mm SL, Anchor's graves, spear, 17
Sept. 2015; MUFS 46900, 183.0 mm SL, MUFS 46901, 189.0mm SL, off Kubura Hinan Port, line-fishing, 18 Sept. 2015;
MUFS 46933, 95.0 mm SL, Sonai Fishing Port, line-fishing, 18
Sept. 2015; MUFS 47009, 194.0 mm SL, off Northern Pasture,line-fishing, 22 Sept. 2015; URM-P 48359, 137.6 mm SL,
Kubura Fishing Port, line-fishing, 23 July 2013.

Gymnocranius euanus (Günther, 1879)
シロダイ [Marine] (Fig. 209)
しろいゆ
FRLM 52011, 135.4 mm SL, off Hikawa, line-fishing,
21 Sept. 2015; KAUM–I. 80465, 268.8 mm SL, north of
Yonaguni-jima island, line-fishing, 22 Sept. 2015; MUFS
46915, 301.5 mm SL, Yonaguni-jima island, line-fishing, 18
Sept. 2015; MUFS 46971, 253.4 mm SL, MUFS 46972, 240.8mm SL, MUFS 46973, 319.7 mm SL, MUFS 46974, 225.4mm SL, off Hikawa, line-fishing, 21 Sept. 2015.

Fig. 209. Gymnocranius euanus シロダイ (upper: KAUM–I. 80465, 268.8mm SL; lower: FRLM 52011, 135.4 mm SL).

Gymnocranius sp.
メイチダイ属の一種 [Marine] (Fig. 210)
MUFS 47335, 221.8 mm SL, off cape Umabana,
Yonaguni-jima island, line-fishing, 22 Sept. 2015.

Fig. 210. Gymnocranius sp. (MUFS 47335, 221.8 mm SL).

Fig. 208. Gnathodentex aureolineatus ノ コ ギ リ ダ イ (upper: URM-P
48359, 137.6 mm SL; middle: MUFS 47009, 194.0 mm SL; lower:
Diving Service MARLIN).

46

Checklist of fishes of Yonaguni-jima island
Lethrinus amboinensis Bleeker 1854
ヨコシマフエフキ [Marine] (Fig. 211)
KAUM–I. 80207, 173.5 mm SL, west of Yonaguni-jimaisland, line-fishing, 18 Sept. 2015.

Fig. 211. Lethrinus amboinensis ヨ コ シ マ フ エ フ キ (KAUM–I. 80207,
173.5 mm SL).

Lethrinus harak (Forsskål, 1775)
マトフエフキ [Marine; brackish] (Fig. 212)
KAUM–I. 78285, 198.5 mm SL, Kubura Fishing Port,line-fishing, 16 Sept. 2015.

Lethrinus rubrioperculatus Sato, 1978
ホオアカクチビ [Marine] (Fig. 214)
FRLM 51973, 146.9 mm SL, FRLM 52097, 204.6 mm
SL, FRLM 52102, 146.8 mm SL, off Hikawa, line-fishing, 21
Sept. 2015; KAUM–I. 79716, 240.2 mm SL, Yonaguni-jimaisland, line-fishing, 18 Sept. 2015; KAUM–I. 80207, 173.5mm SL, off Hikawa, line-fishing, 18 Sept. 2015; KAUM–I.
80392, 258.1 mm SL, off Hikawa, line-fishing, 21 Sept. 2015;
KAUM–I. 80484, 239.0 mm SL, KAUM–I. 80485, 205.4mm SL, off Sonai, line-fishing, 22 Sept. 2015; MUFS 46963,
MUFS 46964, 236.0 mm SL, MUFS 46965, 186.0 mm SL,
MUFS 46966, 272.0 mm SL, MUFS 46967, 185.0 mm SL,
MUFS 46968, MUFS 46969, 269.0 mm SL, MUFS 46970,
MUFS 46975, 172.0 mm SL, MUFS 46982, 165.0 mm SL,
MUFS 46983, 167.0 mm SL, MUFS 46984, 153.0 mm SL,
MUFS 46985, 185.0 mm SL, off Hikawa, line-fishing, 21
Sept. 2015; MUFS 46997, 253.0 mm SL, MUFS 46998, 249.0mm SL, MUFS 47006, MUFS 47007, 254.0 mm SL, MUFS
47008, 279.0 mm SL, MUFS 47010, 170.0 mm SL, MUFS
47011, 157.0 mm SL, off Northern Pasture, line-fishing, 22
Sept. 2015.

Fig. 212. Lethrinus harak マトフエフキ (KAUM–I. 78285, 198.5 mm SL).

Lethrinus microdon Valenciennes, 1830
オオフエフキ [Marine] (Fig. 213)
FRLM 51427, 204.2 mm SL, Kubura Fishing Port, linefishing, 19 Sept. 2015.

Fig. 214. Lethrinus rubrioperculatus ホオアカクチビ (upper: KAUM–I.
80484, 239.0 mm SL; lower: MUFS 46983, 167.0 mm SL).

Fig. 213. Lethrinus microdon オオフエフキ (FRLM 51427, 204.2 mm SL).

Lethrinus xanthochilus Klunzinger, 1870
ムネアカクチビ [Marine] (Fig. 215)
MUFS 46961, 121.0 mm SL, levee at Kubura, line-fishing,
20 Sept. 2015.

Lethrinus olivaceus Valenciennes, 1830
キツネフエフキ [Marine]
うむな・むるみ
Ikema (2003).
Lethrinus ornatus Valenciennes, 1830
ハナフエフキ [Marine]
むるみ
Ikema (2003).

Fig. 215. Lethrinus xanthochilus ム ネ ア カ ク チ ビ (MUFS 46961, 121.0
mm SL).

47

K. Koeda et al.
Monotaxis grandoculis (Forsskål, 1775)
ヨコシマクロダイ [Marine] (Fig. 216)
たるみ
FRLM 50755, 96.1 mm SL, Sonai Fishing Port, linefishing, 14 Mar. 2015; FRLM 51496, 291.0 mm SL, Sonai
Fishing Port, line-fishing, 22 Sept. 2015; MUFS 46995, 290.0mm SL, Sonai Fishing Port, line-fishing, 22 Sept. 2015.

Fig. 216. Monotaxis grandoculis ヨコシマクロダイ (upper: MUFS 46995,
290.0 mm SL; lower: FRLM 50755, 96.1 mm SL).

FAMILY MULLIDAE
ヒメジ科
Mulloidichthys flavolineatus flavolineatus (Lacepède, 1801)
モンツキアカヒメジ [Marine; brackish] (Fig. 217)
Yoshigou et al. (2001).
FRLM 51498, 209.0 mm SL, Sonai Fishing Port, linefishing, 22 Sept. 2015; FRLM 52054, 169.7 mm SL, Sonai
Fishing Port, line-fishing, 24 Sept. 2015; HMNH-P82, 76.6mm SL, HMNH-P83, 75.1 mm SL, Tabaru River, hand net, 10Aug. 1999; KAUM–I. 78402, 129.4 mm SL, Kubura Fishing
Port, gill net, 20 Sept. 2015; MUFS 46959, 134.0 mm SL,
Kubura Fishing Port, gill net, 20 Sept. 2015.

Fig. 217. Mulloidichthys flavolineatus flavolineatus モンツキアカヒメジ(upper: KAUM–I. 78402, 129.4 mm SL; lower: MUFS 46959, 134.0
mm SL).

Mulloidichthys pflugeri (Steindachner, 1900)
リュウキュウアカヒメジ [Marine] (Fig. 218)
Koeda et al. (in press).
MUFS 47000, 187.0 mm SL, Umabana, line-fishing, 22
Sept. 2015.

Fig. 218. Mulloidichthys pflugeri リ ュ ウ キ ュ ウ ア カ ヒ メ ジ [MUFS47000, 187.0 mm SL (upper: fresh specimen; lower: live specimen)].

48

Checklist of fishes of Yonaguni-jima island
Mulloidichthys vanicolensis (Valenciennes, 1831)
アカヒメジ [Marine] (Fig. 219)
Yoshigou et al. (2001).
FRLM 50738, 159.3 mm SL, FRLM 50739, 159.2 mm SL,
Sonai Fishing Port, line-fishing, 15 Mar. 2015; FRLM 51497,
205.0 mm SL, Sonai Fishing Port, line-fishing, 22 Sept. 2015;
FRLM 51588, 131.4 mm SL, mouth of Kubura River, gill
net, 20 Sept. 2015; FRLM 51960, 195.5 mm SL, Kuburabari, line-fishing, 20 Sept. 2015; HMNH-P124, 109.7 mm SL,
Agarizaki, hand net, 9 Aug. 1999; KAUM–I. 80222, 163.5mm SL, Arakawa Bay, spear, 18 Sept. 2015; MUFS 46958,
162.0 mm SL, levee at Kubura, line-fishing, 20 Sept. 2015.

Parupeneus multifasciatus (Quoy and Gaimard, 1825)
オジサン [Marine] (Fig. 221)
FRLM 50724, 125.7 mm SL, Kubura Fishing Port, linefishing, 12 Mar. 2015; FRLM 51962, 129.8 mm SL, off
Northern Pasture, line-fishing, 22 Sept. 2015; FRLM 51963,
112.5 mm SL, off Northern Pasture, line-fishing, 23 Sept.
2015; FRLM 51964, 156.3 mm SL, off Northern Pasture,line-fishing, 24 Sept. 2015; FRLM 51965, 154.5 mm SL, off
Northern Pasture, line-fishing, 25 Sept. 2015; FRLM 51966,
153.0 mm SL, off Northern Pasture, line-fishing, 26 Sept.
2015; FRLM 51987, 166.6 mm SL, Sonai Fishing Port, linefishing, 19 Sept. 2015; KAUM–I. 79736, 164.4 mm SL,
KAUM–I. 79737, 99.1 mm SL, off Kubura Hinan Port, linefishing, 18 Sept. 2015; KAUM–I. 80219, 133.5 mm SL,
Arakawa Bay, line-fishing, 18 Sept. 2015; KAUM–I. 80228,
161.0 mm SL, Yonaguni-jima island, line-fishing, 17 Sept.
2015; KAUM–I. 80393, 113.7 mm SL, KAUM–I. 80424,
150.7 mm SL, KAUM–I. 80426, 93.1 mm SL, off Hikawa,line-fishing, 21 Sept. 2015; KAUM–I. 80490, 148.0 mm SL,
off Yonaguni Airport, line-fishing, 22 Sept. 2015; MUFS
46884, 115.0 mm SL, Kubura Fishing Port, line-fishing, 17
Sept. 2015; MUFS 46913, 141.0 mm SL, off Kubura Hinan
Port, line-fishing, 18 Sept. 2015; NSMT-P 101230, 133.4 mm
SL, Kubura Hinan Port, 16 Mar. 2010.

Fig. 219. Mulloidichthys vanicolensis ア カ ヒ メ ジ (upper: KAUM–I.
80222, 163.5 mm SL; lower: MUFS 46958, 162.0 mm SL).

Parupeneus barberinus (Lacepède, 1801)
オオスジヒメジ [Marine]
んまぶる
Ikema (2003).

Parupeneus ciliatus (Lacepède, 1802)
ホウライヒメジ [Marine] (Fig. 220)

Fig. 221. Parupeneus multifasciatus オジサン (upper: KAUM–I. 80228,
161.0 mm SL; lower: KAUM–I. 79737, 99.1 mm SL).

Parupeneus pleurostigma (Bennett, 1831)
リュウキュウヒメジ [Marine] (Fig. 222)
FRLM 51460, 124.1 mm SL, off Hikawa, line-fishing, 21
Sept. 2015.

Fig. 220. Parupeneus ciliatus ホウライヒメジ (Diving Service MARLIN).

Fig. 222. Parupeneus pleurostigma リュウキュウヒメジ (FRLM 51460,
124.1 mm SL).

49

K. Koeda et al.
FAMILY PEMPHERIDAE
ハタンポ科
かたちら
Parapriacanthus ransonneti Steindachner, 1870
キンメモドキ [Marine] (Fig. 223)

Pempheris oualensis Cuvier, 1831
ユメハタンポ [Marine] (Fig. 225)
KAUM–I. 78328, 171.8 mm SL, KAUM–I. 78329, 148.9mm SL, male, Rakuda Rock, spear, 17 Sept. 2015; KAUM–
I. 78354, 154.9 mm SL, female, KAUM–I. 78355, 168.1 mm
SL, female, Arakawa Bay, spear, 18 Sept. 2015; KAUM–I.
78550, 186.1 mm SL, Umabana, spear, 22 Sept. 2015; URM-P
47551, 169.1 mm SL, Kubura Fishing Port, line-fishing, 23
July 2013.

Fig. 223. Parapriacanthus ransonneti キ ン メ モ ド キ (Diving Service
MARLIN).

Pempheris adusta Bleeker, 1877
リュウキュウハタンポ [Marine] (Fig. 224)
FRLM 51543, 111.5 mm SL, Sonai Fishing Port, linefishing, 24 Sept. 2015; KAUM–I. 78465, 130.3 mm SL, Sonai
Channel, spear, 21 Sept. 2015; URM-P 47552, 127.2 mm SL,
Sonai Fishing Port, line-fishing, 23 July 2013; URM-P 47553,
124.6 mm SL, URM-P 47554, 97.4 mm SL, URM-P 47555,
128.0 mm SL, URM-P 47556, 121.4 mm SL, URM-P 47557,
125.4 mm SL, URM-P 47558, 129.3 mm SL, URM-P 47559,
107.2 mm SL, URM-P 47560, 121.9 mm SL, URM-P 47561,
130.9 mm SL, URM-P 47562, 123.3 mm SL, URM-P 47563,
132.8 mm SL, URM-P 47564, 122.0 mm SL, URM-P 47565,
117.6 mm SL, Kubura Fishing Port, line-fishing, 23 July
2013; URM-P 47566, 129.0 mm SL, URM-P 47567, 123.0mm SL, URM-P 47568, 116.5 mm SL, Kubura Fishing Port,line-fishing, 24 July 2013.

Fig. 225. Pempheris oualensis ユメハタンポ (upper: URM-P 47551, 169.1mm SL; middle: KAUM–I. 78328, 171.8 mm SL; lower: K. Koeda).

Pempheris vanicolensis Cuvier, 1831
キビレハタンポ [Marine] (Fig. 226)
URM-P 47569, 139.0 mm SL, Kubura Fishing Port, linefishing, 24 July 2013.

Fig. 224. Pempheris adusta リュウキュウハタンポ (upper: FRLM 51543,
111.5 mm SL; lower: URM-P 47554, 97.4 mm SL).

50

Fig. 226. Pempheris vanicolensis キビレハタンポ (URM-P 47569, 139.0mm SL).

Checklist of fishes of Yonaguni-jima island
FAMILY MONODACTYLIDAE
ヒメツバメウオ科
Monodachtylus argenteus (Linnaeus, 1758)
ヒメツバメウオ [Marine; brackish] (Fig. 227)
Yoshigou (2000); Yoshigou et al. (2001).
HMNH-P23, 69.3 mm SL, Tabaru River, hand net, 8 Aug.
1999; KAUM–I. 78250, 142.6 mm SL, KAUM–I. 78251,
146.3 mm SL, KAUM–I. 78263, 151.0 mm SL, Kubura
Fishing Port, spear, 16 Sept. 2015; KAUM–I. 78429, 13.0 mm
SL, Tabaru River, cast net, 20 Sept. 2015.

FAMILY KYPHOSIDAE
イスズミ科
Kyphosus cinerascens (Forsskål, 1775)
テンジクイサキ [Marine; brackish] (Fig. 228)
FRLM 50707, 124.3 mm SL, Kubura Fishing Port, handnet, 13 Mar. 2015; FRLM 51558, 187.5 mm SL, Hikawa, linefishing, 24 Sept. 2015; FRLM 51589, 135.0 mm SL, mouthof Kubura River, gill net, 20 Sept. 2015; FRLM 51958, 222.0mm SL, Kubura-bari, line-fishing, 20 Sept. 2015; HMNH-P
80, 140.5 mm SL, 9 Aug. 1999, Kubura River, hand net;
KAUM–I. 78252, 226.2 mm SL, Kubura Fishing Port, spear,
16 Sept. 2015; KAUM–I. 80365, 157.4 mm SL, Kubura
Fishing Port, gill net, 20 Sept. 2015.

Fig. 228. Kyphosus cinerascens テンジクイサキ (KAUM–I. 80365, 157.4mm SL).

Kyphosus pacificus Sakai and Nakabo, 2004
ミナミイスズミ [Marine] (Fig. 229)
FRLM 51429, 152.9 mm SL, Sonai Fishing Port, linefishing, 19 Sept. 2015; KAUM–I. 80468, 229.0 mm SL,
Kubura Fishing Port, line-fishing, 20 Sept. 2015; MUFS
46921, 226.0 mm SL, levee at Kubura, line-fishing, 18 Sept.
2015.

Fig. 227. Monodactylus argenteus ヒ メ ツ バ メ ウ オ (upper: KAUM–I.
78250, 142.6 mm SL; lower: KAUM–I. 78429, 13.0 mm SL).

Fig. 229. Kyphosus pacificus ミナミイスズミ (upper: KAUM–I. 80468,
229.0 mm SL; lower: FRLM 51429, 152.9 mm SL).

51

K. Koeda et al.
Kyphosus vaigiensis (Quoy and Gaimard, 1825)
イスズミ [Marine] (Fig. 230)
FRLM 50757, 286.0 mm SL, FRLM 50758, 243.3 mm SL,
Kubura-bari, line-fishing, 13 Mar. 2015; KAUM–I. 78608,
209.6 mm SL, Yonaguni-jima island, line-fishing, 18 Sept.
2015.

Chaetodon auriga Forsskål, 1775
トゲチョウチョウウオ [Marine; brackish] (Fig. 232)
FRLM 50721, 151.2 mm SL, Kubura Fishing Port, handnet, 12 Mar. 2015; FRLM 51567, 143.2 mm SL, Kubura
Fishing Port, line-fishing, 24 Sept. 2015; KAUM–I. 80278,
124.3 mm SL, Sonai Fishing Port, line-fishing, 20 Sept. 2015;
URM-P 48357, 134.2 mm SL, Kubura Fishing Port, linefishing, 23 July 2013.

Fig. 232. Chaetodon auriga トゲチョウチョウウオ (FRLM 50721, 151.2mm SL).

Fig. 230. Kyphosus vaigiensis イスズミ (upper: FRLM 50757, 286.0 mm
SL; lower: KAUM–I. 78608, 209.6 mm SL).

Chaetodon auripes Jordan and Snyder, 1901
チョウチョウウオ [Marine; brackish] (Fig. 233)
Ito (2009).
FRLM 50697, 125.4 mm SL, Sonai Fishing Port, linefishing, 13 Mar. 2015; FRLM 50722, 124.6 mm SL, Kubura
Fishing Port, hand net, 12 Mar. 2015; FRLM 51566, 140.1
mm SL, Kubura Fishing Port, line-fishing, 24 Sept. 2015;
KAUM–I. 78260, 132.6 mm SL, Kubura Fishing Port,
spear, 16 Sept. 2015; KAUM–I. 80362, 132.1 mm SL, Sonai
Channel, gill net, 20 Sept. 2015.

FAMILY CHAETODONTIDAE
チョウチョウウオ科
かびっつぁ・はびるいゆ
Chaetodon argentatus Smith and Radcliffe, 1911
カガミチョウチョウウオ [Marine] (Fig. 231)
KAUM–I. 78325, 81.1 mm SL, Anchor's graves, spear, 17
Sept. 2015.

Fig. 233. Chaetodon auripes チョウチョウウオ (KAUM–I. 80362, 132.1
mm SL).

Chaetodon bennetti Cuvier, 1831
ウミヅキチョウチョウウオ [Marine] (Fig. 234)
Fig. 231. Chaetodon argentatus カ ガ ミ チ ョ ウ チ ョ ウ ウ オ (KAUM–I.
78325, 81.1 mm SL).

52

Checklist of fishes of Yonaguni-jima island
Chaetodon lineolatus Cuvier, 1831
ニセフウライチョウチョウウオ [Marine] (Fig. 237)
KAUM–I. 80464, 205.1 mm SL, Umabana, spear, 22 Sept.
2015.

Fig. 234. Chaetodon bennetti ウミヅキチョウチョウウオ (Diving Service
MARLIN).

Chaetodon citrinellus Cuvier, 1831
ゴマチョウチョウウオ [Marine] (Fig. 235)
Ito (2009).
KAUM–I. 78495, 36.1 mm SL, Hikawa, hand net, 21
Sept. 2015.

Fig. 237. Chaetodon lineolatus ニセフウライチョウチョウウオ (KAUM–
I. 80464, 205.1 mm SL).

Chaetodon lunula (Lacepède, 1802)
チョウハン [Marine; brackish] (Fig. 238)
Yoshigou et al. (2001).
HMNH-P136, 25.6 mm SL, Sonai, hand net, 9 Aug. 1999;
FRLM 51980, 85.5 mm SL, Hikawa, spear, 21 Sept. 2015.

Fig. 235. Chaetodon citrinellus ゴマチョウチョウウオ (KAUM–I. 78495,
36.1 mm SL).

Chaetodon ephippium Cuvier, 1831
セグロチョウチョウウオ [Marine] (Fig. 236)
KAUM–I. 78497, 90.3 mm SL, Hikawa, hand net, 21
Sept. 2015.
Fig. 238. Chaetodon lunula チョウハン (FRLM 51980, 85.5 mm SL).

Fig. 236. Chaetodon ephippium セ グ ロ チ ョ ウ チ ョ ウ ウ オ (KAUM–I.
78497, 90.3 mm SL).

53

K. Koeda et al.
Chaetodon lunulatus Quoy and Gaimard, 1825
ミスジチョウチョウウオ [Marine] (Fig. 239)
KAUM–I. 78471, 22.9 mm SL, Sonai Channel, hand net,
21 Sept. 2015.

Chaetodon meyeri Bloch and Schneider, 1801
オウギチョウチョウウオ [Marine] (Fig. 241)

Fig. 241. Chaetodon meyeri オウギチョウチョウウオ (Diving Service
MARLIN).

Chaetodon ornatissimus Cuvier, 1831
ハナグロチョウチョウウオ [Marine] (Fig. 242)
KAUM–I. 79745, 140.3 mm SL, Rakuda Rock, spear, 17
Sept. 2015.

Fig. 239. Chaetodon lunulatus ミ ス ジ チ ョ ウ チ ョ ウ ウ オ (KAUM–I.
78471, 22.9 mm SL).

Chaetodon melannotus Bloch and Schneider, 1801
アケボノチョウチョウウオ [Marine] (Fig. 240)
KAUM–I. 80260, 113.9 mm SL, Anchor's graves, spear,
17 Sept. 2015.

Fig. 242. Chaetodon ornatissimus ハナグロチョウチョウウオ (KAUM–I.
79745, 140.3 mm SL).

Chaetodon ornatissimus × C. reticulatus
ハナグロチョウチョウウオ × ハクテンカタギ [Marine](Fig. 243)
Remarks. The individual has intermediate color pattern ofChaetodon ornatissimus and C. reticulatus, and formed a pairwith C. reticulatus.

Fig. 240. Chaetodon melannotus アケボノチョウチョウウオ (upper: KAUM–
I. 80260, 113.9 mm SL; lower: Diving Service MARLIN).

54

Fig. 243. Chaetodon ornatissimus × C. reticulatus ハナグロチョウチョウ
ウオ × ハクテンカタギ (Diving Service MARLIN).

Checklist of fishes of Yonaguni-jima island
Chaetodon punctatofasciatus Cuvier, 1831
シチセンチョウチョウウオ [Marine] (Fig. 244)
KAUM–I. 78326, 82.6 mm SL, Anchor's graves, spear, 17
Sept. 2015.

Chaetodon reticulatus Cuvier, 1831
ハクテンカタギ [Marine] (Fig. 247)
Ito (2009).
KAUM–I. 80259, 103.4 mm SL, Anchor's graves, spear,
17 Sept. 2015.

Fig. 244. Chaetodon punctatofasciatus シ チ セ ン チ ョ ウ チ ョ ウ ウ オ(KAUM–I. 78326, 82.6 mm SL).

Chaetodon quadrimaculatus Gray, 1831
シテンチョウチョウウオ [Marine] (Fig. 245)

Fig. 245. Chaetodon quadrimaculatus シテンチョウチョウウオ (Diving
Service MARLIN).

Fig. 247. Chaetodon reticulatus ハクテンカタギ (upper: KAUM–I. 80259,
103.4 mm SL; lower: Diving Service MARLIN).

Chaetodon rafflesii Anonymous [Bennett], 1830
アミチョウチョウウオ [Marine] (Fig. 246)
URM-P 48358, 127.9 mm SL, Kubura Fishing Port, linefishing, 23 July 2013.

Chaetodon semeion Bleeker, 1855
レモンチョウチョウウオ [Marine] (Fig. 248)
KAUM–I. 78353, 164.9 mm SL, Arakawa Bay, spear, 18
Sept. 2015.

Fig. 246. Chaetodon rafflesii ア ミ チ ョ ウ チ ョ ウ ウ オ (URM-P 48358,
127.9 mm SL).

Fig. 248. Chaetodon semeion レ モ ン チ ョ ウ チ ョ ウ ウ オ (KAUM–I.
78353, 164.9 mm SL).

55

K. Koeda et al.
Chaetodon speculum Cuvier, 1831
トノサマダイ [Marine] (Fig. 249)
KAUM–I. 80462, 109.3 mm SL, Umabana, spear, 22 Sept.
2015.

Fig. 249. Chaetodon speculum トノサマダイ (KAUM–I. 80462, 109.3 mm SL).

Chaetodon trifascialis Quoy and Gaimard, 1825
ヤリカタギ [Marine] (Fig. 250)

Fig. 251. Chaetodon ulietensis スダレチョウチョウウオ (upper: FRLM
50698, 131.9 mm SL; lower: KAUM–I. 78494, 23.4 mm SL).

Fig. 250. Chaetodon trifascialis ヤリカタギ (M. Sakurai).

Chaetodon unimaculatus Bloch, 1787
イッテンチョウチョウウオ [Marine] (Fig. 252)
MUFS 46888, 142.0 mm SL, Rakuda Rock, spear, 17
Sept. 2015.

Chaetodon ulietensis Cuvier, 1831
スダレチョウチョウウオ [Marine] (Fig. 251)
FRLM 50698, 131.9 mm SL, FRLM 50699, 121.5 mm
SL, Sonai Fishing Port, line-fishing, 13 Mar. 2015; KAUM–I.
78494, 23.4 mm SL, Hikawa, hand net, 21 Sept. 2015.

Fig. 252. Chaetodon unimaculatus イッテンチョウチョウウオ (MUFS
46888, 142.0 mm SL).

56

Checklist of fishes of Yonaguni-jima island
Chaetodon vagabundus Linnaeus, 1758
フウライチョウチョウウオ [Marine] (Fig. 253)
FRLM 51552, 35.6 mm SL, Kubura Fishing Port, handnet, 24 Sept. 2015; KAUM–I. 78253, 99.8 mm SL, Kubura
Fishing Port, spear, 16 Sept. 2015.

Hemitaurichthys polylepis (Bleeker, 1857)
カスミチョウチョウウオ [Marine] (Fig. 256)
KAUM–I. 78315, 111.7 mm SL, KAUM–I. 80261, 113.0mm SL, Anchor's graves, spear, 17 Sept. 2015.

Fig. 253. Chaetodon vagabundus フウライチョウチョウウオ (KAUM–I.
78253, 99.8 mm SL).

Forcipiger flavissimus Jordan and McGregor, 1898
フエヤッコダイ [Marine] (Fig. 254)
KAUM–I. 78574, 136.0 mm SL, Danno Beach, spear, 23
Sept. 2015.

Fig. 256. Hemitaurichthys polylepis カ ス ミ チ ョ ウ チ ョ ウ ウ オ (upper:
KAUM–I. 78315, 111.7 mm SL; lower: KAUM–I. 80261, 113.0 mm SL).

Fig. 254. Forcipiger flavissimus フエヤッコダイ (KAUM–I. 78574, 136.0mm SL).

Forcipiger longirostris (Broussonet, 1782)
オオフエヤッコダイ [Marine] (Fig. 255)

Fig. 255. Forcipiger longirostris オ オ フ エ ヤ ッ コ ダ イ (Diving Service
MARLIN).

57

K. Koeda et al.
Heniochus acuminatus (Linnaeus, 1758)
ハタタテダイ [Marine] (Fig. 257)
FRLM 52032, 131.6 mm SL, Kubura-bari, line-fishing, 22
Sept. 2015; KAUM–I. 78261, 122.0 mm SL, Kubura Fishing
Port, spear, 16 Sept. 2015; NSMT-P 101234, 147.6 mm SL,
Danno Beach, 19 Mar. 2010.

Heniochus chrysostomus Cuvier, 1831
ミナミハタタテダイ [Marine] (Fig. 258)
KAUM–I. 79747, 112.6 mm SL, KAUM–I. 80229, 105.5mm SL, Rakuda Rock, spear, 17 Sept. 2015; MUFS 46892,
109.0 mm SL, Rakuda Rock, spear, 17 Sept. 2015.

Fig. 258. Heniochus chrysostomus ミ ナ ミ ハ タ タ テ ダ イ (KAUM–I.
79747, 112.6 mm SL).

Heniochus singularius Smith and Radcliffe, 1911
シマハタタテダイ [Marine] (Fig. 259)
KAUM–I. 80234, 181.9 mm SL, Rakuda Rock, spear, 17
Sept. 2015.

Fig. 257. Heniochus acuminatus ハタタテダイ (upper: KAUM–I. 78261,
122.0 mm SL; lower: Diving Service MARLIN).

Fig. 259. Heniochus singularius シ マ ハ タ タ テ ダ イ (KAUM–I. 80234,
181.9 mm SL).

Heniochus varius (Cuvier, 1829)
ツノハタタテダイ [Marine]
NSMT-P 101235, 141.6 mm SL, Danno Beach, 19 Mar. 2010.

58

Checklist of fishes of Yonaguni-jima island
FAMILY POMACANTHIDAE
キンチャクダイ科
Apolemichthys trimaculatus (Cuvier, 1831)
シテンヤッコ [Marine] (Fig. 260)
KAUM–I. 79735, 117.5 mm SL, off Kubura Hinan Port,
spear, 18 Sept. 2015; MUFS 46907, 138.0 mm SL, off Kubura
Hinan Port, line-fishing, 18 Sept. 2015; NSMT-P 101239,
103.6 mm SL, Danno Beach, 19 Mar. 2010.

Centropyge ferrugata Randall and Burgess, 1972
アカハラヤッコ [Marine] (Fig. 261)
KAUM–I. 78320, 43.1 mm SL, Rakuda Rock, hand net, 17
Sept. 2015; KAUM–I. 78346, 58.0 mm SL, KAUM–I. 78358,
25.1 mm SL, off Kubura Hinan Port, hand net, 18 Sept. 2015.

Fig. 261. Centropyge ferrugata アカハラヤッコ (upper: KAUM–I. 78346,
58.0 mm SL; lower: KAUM–I. 78358, 25.1 mm SL).

Centropyge heraldi Woods and Schultz, 1953
ヘラルドコガネヤッコ [Marine] (Fig. 262)
Ito (2009).
KAUM–I. 78343, 70.0 mm SL, KAUM–I. 78344, 64.2
mm SL, off Kubura Hinan Port, hand net, 18 Sept. 2015.

Fig. 260. Apolemichthys trimaculatus シ テ ン ヤ ッ コ (upper: KAUM–
I. 79735, 117.5 mm SL; middle: MUFS 46907, 138.0 mm SL; lower:
Diving Service MARLIN).

Fig. 262. Centropyge heraldi ヘラルドコガネヤッコ (KAUM–I. 78343,
70.0 mm SL).

59

K. Koeda et al.
Centropyge vrolikii (Bleeker, 1853)
ナメラヤッコ [Marine] (Fig. 263)

Pomacanthus semicirculatus (Cuvier, 1831)
サザナミヤッコ [Marine] (Fig. 265)
Yoshigou et al. (2001).
HMNH-P135, 33.3 mm SL, Sonai, hand net, 9 Aug. 1999;
KAUM–I. 80233, 130.5 mm SL, Rakuda Rock, spear, 17
Sept. 2015; NSMT-P 55482, 36.8 mm SL, Agarizaki, 28 Mar.
1997.

Fig. 263. Centropyge vrolikii ナメラヤッコ (Diving Service MARLIN).

Pomacanthus imperator (Bloch, 1787)
タテジマキンチャクダイ [Marine] (Fig. 264)
KAUM–I. 78468, 23.1 mm SL, Sonai Channel, hand net,
21 Sept. 2015; KAUM–I. 79714, 212.9 mm SL, off Kubura
Hinan Port, spear, 18 Sept. 2015; NSMT-P 101236, 173.1 mm
SL, Danno Beach, 19 Mar. 2010.

Fig. 265. Pomacanthus semicirculatus サザナミヤッコ (KAUM–I. 80233,
130.5 mm SL).

Pomacanthus sexstriatus (Cuvier, 1831)
ロクセンヤッコ [Marine] (Fig. 266)
Ito (2009).

Fig. 266. Pomacanthus sexstriatus ロクセンヤッコ (Diving Service MARLIN).

Pomacanthus xanthometopon (Bleeker, 1853)
アデヤッコ [Marine]
Ito (2009).

Fig. 264. Pomacanthus imperator タ テ ジ マ キ ン チ ャ ク ダ イ (upper:
KAUM–I. 79714, 212.9 mm SL; lower: KAUM–I. 78468, 23.1 mm SL).

60

Checklist of fishes of Yonaguni-jima island
Pygoplites diacanthus (Boddaert, 1772)
ニシキヤッコ [Marine] (Fig. 267)
KAUM–I. 79744, 111.6 mm SL, Rakuda Rock, spear, 17
Sept. 2015; MUFS 46891, 108.0 mm SL, Rakuda Rock, spear,
17 Sept. 2015.

FAMILY TERAPONIDAE
シマイサキ科
Terapon jarbua (Forsskål, 1775)
コトヒキ [Marine; brackish] (Fig. 268)
もんたな
Yoshigou (2000); Yoshigou et al. (2001).
FRLM 51446, 84.2 mm SL, FRLM 52012, 60.0 mm SL,
mouth of Kubura River, cast net, 21 Sept. 2015; HMNH-P91,
43.8 mm SL, Tabaru River, hand net, 10 Aug. 1999; MUFS
47025, 105.0 mm SL, Kubura Fishing Port, line-fishing, 24
Sept. 2015.

Fig. 267. Pygoplites diacanthus ニシキヤッコ (upper: KAUM–I. 79744,
111.6 mm SL; lower: MUFS 46891, 108.0 mm SL).

Fig. 268. Terapon jarbua コ ト ヒ キ (upper: FRLM 51446, 84.2 mm SL;
lower: MUFS 47025, 105.0 mm SL).

FAMILY PENTACEROTIDAE
カワビシャ科Pentaceros japonicus Steindachner, 1883
ツボダイ [Marine]
NSMT-P 76169, 130.0 mm SL, southeast of Yonagunijima island, stomach content, 1 Aug. 2006.

FAMILY KUHLIIDAE
ユゴイ科
Kuhlia marginata (Cuvier, 1829)
ユゴイ [Brackish; freshwater] (Fig. 269)
Fujimoto (1972); Yoshigou (2000); Yoshigou et al. (2001).
FRLM 51487, 95.5 mm SL, Tabaru River, cast net, 20
Sept. 2015; HMNH-P90, 2 specimens, 23.5–47.1 mm SL,
Tabaru River, hand net, 10 Aug. 1999; KAUM–I. 78407, 57.5mm SL, Tabaru River, cast net, 20 Sept. 2015.

Fig. 269. Kuhlia marginata ユゴイ (KAUM–I. 78407, 57.5 mm SL).

61

K. Koeda et al.
Kuhlia mugil (Forster, 1801)
ギンユゴイ [Marine] (Fig. 270)
FRLM 51571, 184.3 mm SL, FRLM 51572, 186.0 mm
SL, FRLM 52068, 178.7 mm SL, Kubura Fishing Port,line-fishing, 24 Sept. 2015; KAUM–I. 78381, 21.5 mm SL,
KAUM–I. 78382, 18.8 mm SL, Southern Pasture, hand net, 19
Sept. 2015; MUFS 46944, 32.8 mm SL, Danno Beach, handnet, 19 Sept. 2015; URM-P 48360, 169.2 mm SL, Kubura
Fishing Port, line-fishing, 23 July 2013.

Fig. 271. Kuhlia rupestris オオクチユゴイ (upper: FRLM 51563, 205.8mm SL; middle: KAUM–I. 78442, 129.0 mm SL; lower: KAUM–I.
80282, 145.3 mm SL).

Fig. 270. Kuhlia mugil ギ ン ユ ゴ イ (upper: URM-P 48360, 169.2 mm SL;
middle: KAUM–I. 78381, 21.5 mm SL; lower: MUFS 46944, 32.8 mm SL).

Kuhlia rupestris (Lacepède, 1802)
オオクチユゴイ [Brackish; freshwater] (Fig. 271)
Jordan and Tanaka (1927); Kuroiwa (1927); Okinawa
Prefecture Experimental Fisheries Station (1934); Ogawa(1937); Yoshigou (2000); Yoshigou et al. (2001).
FRLM 51563, 205.8 mm SL, Tabaru River, line-fishing,
25 Sept. 2015; FRLM 52062, 225.0 mm SL, Tabaru River,line-fishing, 24 Sept. 2015; HMNH-P24, 60.3 mm SL, Tabaru
River, hand net, 8 Aug. 1999; KAUM–I. 78442, 129.0 mm
SL, mouth of Tabaru River, line-fishing, 20 Sept. 2015;
KAUM–I. 80282, 145.3 mm SL, Tabaru River, line-fishing,
20 Sept. 2015; KAUM–I. 80448, 197.9 mm SL, Tabaru River,gill net, 22 Sept. 2015.

62

FAMILY OPLEGNATHIDAE
イシダイ科
Oplegnathus punctatus (Temminck and Schlegel, 1844)
イシガキダイ [Marine] (Fig. 272)

Fig. 272. Oplegnathus punctatus イシガキダイ (Diving Service MARLIN).

Checklist of fishes of Yonaguni-jima island
FAMILY CIRRHITIDAE
ゴンベ科
Amblycirrhitus bimacula (Jenkins, 1903)
フタホシゴンベ [Marine] (Fig. 273)
KAUM–I. 78544, 55.4 mm SL, Umabana, hand net, 22
Sept. 2015.

Cirrhitus pinnulatus (Forster, 1801)
イソゴンベ [Marine] (Fig. 276)
FRLM 50746, 157.2 mm SL, Kubura-bari, line-fishing,
14 Mar. 2015; FRLM 51450, 107.3 mm SL, Kubura-bari,line-fishing, 20 Sept. 2015; KAUM–I. 80355, 125.1 mm SL,
Kubura-bari, line-fishing, 20 Sept. 2015; MUFS 47021, 95.0mm SL, Kubura-bari, line-fishing, 23 Sept. 2015; NSMT-P
55456, 111.4 mm SL, Agarizaki, hand net, 28 Mar. 1997.

Fig. 273. Amblycirrhitus bimacula フ タ ホ シ ゴ ン ベ (KAUM–I. 78544,
55.4 mm SL).

Amblycirrhitus unimacula (Kamohara, 1957)
クロホシゴンベ [Marine]
Nakabo (2013).
Cirrhitichthys aprinus (Cuvier, 1829)
ミナミゴンべ [Marine] (Fig. 274)

Fig. 276. Cirrhitus pinnulatus イソゴンベ (upper: KAUM–I. 80355, 125.1mm SL; lower: MUFS 47021, 95.0 mm SL).

Neocirrhites armatus Castelnau, 1873
ベニゴンベ [Marine] (Fig. 277)

Fig. 274. Cirrhitichthys aprinus ミナミゴンベ (Diving Service MARLIN).

Cirrhitichthys falco Randall, 1963
サラサゴンベ [Marine] (Fig. 275)
Ito (2009).
KAUM–I. 78363, 55.9 mm SL, off Kubura Hinan Port,
hand net, 18 Sept. 2015.

Fig. 275. Cirrhitichthys falco サラサゴンベ (KAUM–I. 78363, 55.9 mm SL).

Fig. 277. Neocirrhites armatus ベニゴンベ (Diving Service MARLIN).

Oxycirrhites typus Bleeker, 1857
クダゴンベ [Marine] (Fig. 278)

Fig. 278. Oxycirrhites typus クダゴンベ (Umabana, K. Koeda).

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K. Koeda et al.
Paracirrhites arcatus (Cuvier, 1829)
メガネゴンベ [Marine] (Fig. 279)
FRLM 51476, 39.3 mm SL, Umabana, hand net, 22 Sept.
2015; KAUM–I. 78345, 47.9 mm SL, off Kubura Hinan Port,
hand net, 18 Sept. 2015.

FAMILY CICHLIDAE
カワスズメ科
Oreochromis niloticus (Linnaeus, 1758)
ナイルティラピア [Brackish; freshwater] (Fig. 281)
Yokogawa (2000); Yoshigou (2000); Yoshigou et al.
(2001); Ministry of the Environment (2010b); Ishikawa et al.
(2013); Nakabo (2013).
KAUM–I. 80277, 232.0 mm SL, Tabaru River, linefishing, 20 Sept. 2015.

Fig. 281. Oreochromis niloticus ナ イ ル テ ィ ラ ピ ア (KAUM–I. 80277,
232.0 mm SL).

Fig. 279. Paracirrhites arcatus メ ガ ネ ゴ ン ベ (upper: KAUM–I. 78345,
47.9 mm SL; lower: Diving Service MARLIN).

Paracirrhites forsteri (Schneider, 1801)
ホシゴンベ [Marine] (Fig. 280)
FRLM 51449, 84.9 mm SL, Kubura-bari, line-fishing,
20 Sept. 2015; FRLM 51474, 63.6 mm SL, Umabana, handnet, 22 Sept. 2015; KAUM–I. 78316, 105.3 mm SL, Anchor's
graves, spear, 17 Sept. 2015; KAUM–I. 80358, 107.4 mm SL,
Kubura-bari, line-fishing, 20 Sept. 2015; MUFS 46996, 83.0mm SL, Sonai Fishing Port, line-fishing, 22 Sept. 2015.

Fig. 280. Paracirrhites forsteri ホシゴンベ (upper: FRLM 51449, 84.9 mm
SL; lower: KAUM–I. 78316, 105.3 mm SL).

64

FAMILY POMACENTRIDAE
スズメダイ科ひってぃ
Abudefduf notatus (Day, 1870)
イソスズメダイ [Marine]
NSMT-P 55479, Agarizaki, hand net, 49.4 mm SL, 28
Mar. 1997.

Abudefduf septemfasciatus (Cuvier, 1830)
シチセンスズメダイ [Marine] (Fig. 282)
FRLM 52059, 123.2 mm SL, Hikawa, line-fishing, 24
Sept. 2015; FRLM 52067, 152.0 mm SL, Kubura Fishing
Port, line-fishing, 24 Sept. 2015; HMNH-P 132, 38.5 mm SL,
Sonai, hand net, 9 Aug. 1999.

Fig. 282. Abudefduf septemfasciatus シチセンスズメダイ (FRLM 52059,
123.2 mm SL).

Checklist of fishes of Yonaguni-jima island
Abudefduf sexfasciatus (Lacepède, 1801)
ロクセンスズメダイ [Marine] (Fig. 283)
FRLM 51512, 59.1 mm SL, Kubura Fishing Port, linefishing, 23 Sept. 2015.

Abudefduf vaigiensis (Quoy and Gaimard, 1825)
オヤビッチャ [Marine] (Fig. 285)
FRLM 51952, 139.3 mm SL, Kubura-bari, line-fishing,
20 Sept. 2015; FRLM 51953, 139.8 mm SL, Kubura-bari,line-fishing, 20 Sept. 2015; FRLM 52033, 134.9 mm SL,
FRLM 52034, 127.6 mm SL, Kubura-bari, line-fishing, 22
Sept. 2015; KAUM–I. 80354, 123.2 mm SL, Kubura-bari,line-fishing, 20 Sept. 2015; NSMT-P 55478, 7 specimens,
32.9–60.0 mm SL, Agarizaki, hand net, 28 Mar. 1997.

Fig. 283. Abudefduf sexfasciatus ロクセンスズメダイ (FRLM 51512, 59.1mm SL).

Abudefduf sordidus (Forsskål, 1775)
シマスズメダイ [Marine] (Fig. 284)
Yoshigou et al. (2001).
FRLM 50736, 143.8 mm SL, FRLM 50737, 128.1 mm
SL, Sonai Fishing Port, line-fishing, 15 Mar. 2015;
HMNH-P133, 28.1 mm SL, Sonai, hand net, 9 Aug. 1999;
KAUM–I. 80271, 145.8 mm SL, Sonai Channel, spear, 21
Sept. 2015; NSMT-P 55480, 2 specimens, 39.5–49.9 mm SL,
Agarizaki, hand net, 28 Mar. 1997.

Fig. 285. Abudefduf vaigiensis オヤビッチャ (upper: FRLM 51952, 139.3mm SL; lower: KAUM–I. 80354, 123.2 mm).

Amblyglyphidodon curacao (Bloch, 1787)
クラカオスズメダイ [Marine] (Fig. 286)
KAUM–I. 80353, 94.7 mm SL, Sonai Channel, spear, 21
Sept. 2015.

Fig. 284. Abudefduf sordidus シ マ ス ズ メ ダ イ (KAUM–I. 80271, 145.8mm SL).

Fig. 286. Amblyglyphidodon curacao ク ラ カ オ ス ズ メ ダ イ (KAUM–I.
80353, 94.7 mm SL).

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K. Koeda et al.
Amphiprion clarkii (Bennett, 1830)
クマノミ [Marine] (Fig. 287)
FRLM 52086, 84.8 mm SL, FRLM 52094, 90.4 mm
SL, male, off Hikawa, line-fishing, 21 Sept. 2015; KAUM–
I. 78351, 73.2 mm SL, male, off Kubura Hinan Port, handnet, 18 Sept. 2015; MUFS 47012, 92.0 mm SL, female, off
Northern Pasture, line-fishing, 22 Sept. 2015.

Amphiprion ocellaris Cuvier, 1830
カクレクマノミ [Marine] (Fig. 289)
KAUM–I. 78267, 57.8 mm SL, KAUM–I. 78268, 29.3mm SL, KAUM–I. 78269, 39.4 mm SL, KAUM–I. 78270,
22.4 mm SL, KAUM–I. 78271, 25.0 mm SL, KAUM–I.
78272, 15.6 mm SL, KAUM–I. 78273, 12.0 mm SL (sevenspecimens collected from a single symbiotic anemone),
Kubura Fishing Port, hand net, 16 Sept. 2015.

Fig. 287. Amphiprion clarkii クマノミ (KAUM–I. 78351, 73.2 mm SL).

Amphiprion frenatus Brevoort, 1856
ハマクマノミ [Marine] (Fig. 288)
FRLM 50734, 39.9 mm SL, FRLM 50735, 30.6 mm SL,
Sonai Fishing Port, hand net, 15 Mar. 2015; FRLM 51559,
93.2 mm SL, Sonai Fishing Port, line-fishing, 24 Sept. 2015;
KAUM–I. 78330, 72.4 mm SL, Rakuda Rock, hand net, 17
Sept. 2015.

Fig. 289. Amphiprion ocellaris カクレクマノミ (upper: KAUM–I. 78267,
57.8 mm SL; middle: KAUM–I. 78268, 29.3 mm SL; lower: KAUM–I.
78273, 12.0 mm SL).

Amphiprion perideraion Bleeker, 1855
ハナビラクマノミ [Marine] (Fig. 290)
KAUM–I. 78546, 36.2 mm SL, Umabana, hand net, 22
Sept. 2015.

Fig. 288. Amphiprion frenatus ハマクマノミ (upper: FRLM 51559, 93.2mm SL; lower: KAUM–I. 78330, 72.4 mm SL).

66

Fig. 290. Amphiprion perideraion ハナビラクマノミ (KAUM–I. 78546,
36.2 mm SL).

Checklist of fishes of Yonaguni-jima island
Chromis acares Randall and Swerdloff, 1973
コビトスズメダイ [Marine] (Fig. 291)
KAUM–I. 78350, 31.2 mm SL, off Kubura Hinan Port,
hand net, 18 Sept. 2015.

Fig. 291. Chromis acares コビトスズメダイ (KAUM–I. 78350, 31.2 mm SL).

Chromis atripes Fowler and Bean, 1928
ヒレグロスズメダイ [Marine] (Fig. 292)
KAUM–I. 78324, 46.3 mm SL, Rakuda Rock, hand net,
17 Sept. 2015.

Chromis ovatiformes Fowler, 1946
マルスズメダイ [Marine] (Fig. 294)
KAUM–I. 78322, 58.6 mm SL, Rakuda Rock, hand net,
17 Sept. 2015.

Fig. 294. Chromis ovatiformis マ ル ス ズ メ ダ イ (KAUM–I. 78322, 58.6mm SL).

Chromis vanderbilti (Fowler, 1941)
ヒメスズメダイ [Marine] (Fig. 295)
KAUM–I. 78323, 18.6 mm SL, Rakuda Rock, handnet, 17 Sept. 2015; KAUM–I. 78524, 35.2 mm SL, Dannodropoff, hand net, 22 Sept. 2015; KAUM–I. 78536, 29.4 mm
SL, Umabana, hand net, 22 Sept. 2015.

Fig. 292. Chromis atripes ヒレグロスズメダイ (KAUM–I. 78324, 46.3mm SL).

Chromis margaritifer Fowler, 1946
シコクスズメダイ [Marine] (Fig. 293)
KAUM–I. 78317, 53.6 mm SL, KAUM–I. 78318, 16.2mm SL, KAUM–I. 78319, 17.6 mm SL, Rakuda Rock, handnet, 17 Sept. 2015.

Fig. 295. Chromis vanderbilti ヒメスズメダイ (upper: KAUM–I. 78536,
29.4 mm SL; lower: KAUM–I. 78323, 18.6 mm SL).

Chromis yamakawai Iwatsubo and Motomura, 2013
キホシスズメダイ [Marine] (Fig. 296)

Fig. 293. Chromis margaritifer シコクスズメダイ (KAUM–I. 78317, 53.6mm SL).

Fig. 296. Chromis yamakawai キ ホ シ ス ズ メ ダ イ (Diving Service
MARLIN).

67

K. Koeda et al.
Chrysiptera biocellata (Quoy and Gaimard, 1825)
スジブチスズメダイ [Marine] (Fig. 297)
Ito (2009).
MUFS 47026, 84.0 mm SL, Kubura Fishing Port, linefishing, 24 Sept. 2015.

Chrysiptera cyanea (Quoy and Gaimard, 1825)
ルリスズメダイ [Marine] (Fig. 299)
KAUM–I. 78466, 43.4 mm SL, Sonai Channel, hand net,
21 Sept. 2015.

Fig. 297. Chrysiptera biocellata スジブチスズメダイ (MUFS 47026, 84.0mm SL).

Chrysiptera brownriggi (Bennett, 1828)
ミヤコキセンスズメダイ [Marine] (Fig. 298)
KAUM–I. 78575, 31.9 mm SL, Danno Beach, hand net,
23 Sept. 2015.
Fig. 299. Chrysiptera cyanea ルリスズメダイ (upper: KAUM–I. 78466,
43.4 mm SL; lower: Diving Service MARLIN)

Chrysiptera glauca (Cuvier, 1830)
ネズスズメダイ [Marine]
NSMT-P 55485, 7 specimens, 33.9–50.9 mm SL,
Agarizaki, hand net, 28 Mar. 1997.

Fig. 298. Chrysiptera brownriggi ミヤコキセンスズメダイ (KAUM–I.
78575, 31.9 mm SL).

Chrysiptera rex (Snyder, 1909)
レモンスズメダイ [Marine]
NSMT-P 55484, 51.8 mm SL, Agarizaki, hand net, 28
Mar. 1997.
Chrysiptera unimaculata (Cuvier, 1830)
イチモンスズメダイ [Marine; brackish] (Fig. 300)
MUFS 47027, 71.0 mm SL, mouth of Kubura River, linefishing, 24 Sept. 2015.

Fig. 300. Chrysiptera unimaculata イチモンスズメダイ (MUFS 47027,
71.0 mm SL).

68

Checklist of fishes of Yonaguni-jima island
Dascyllus aruanus (Linnaeus, 1758)
ミスジリュウキュウスズメダイ [Marine] (Fig. 301)
KAUM–I. 78476, 31.4 mm SL, KAUM–I. 78477, 28.0mm SL, Sonai Channel, hand net, 21 Sept. 2015.

Dascyllus trimaculatus (Rüppell, 1829)
ミツボシクロスズメダイ [Marine] (Fig. 303)
FRLM 51988, 98.5 mm SL, FRLM 51992, 59.5 mm SL,
FRLM 51993, 80.9 mm SL, FRLM 51994, 85.1 mm SL,
FRLM 51995, 56.9 mm SL, FRLM 51996, 92.8 mm SL,
Sonai Fishing Port, line-fishing, 19 Sept. 2015; FRLM 52049,
74.7 mm SL, FRLM 52050, 59.5 mm SL, Sonai Fishing
Port, line-fishing, 24 Sept. 2015; FRLM 52069, 92.3 mm SL,
Kubura Fishing Port, line-fishing, 24 Sept. 2015; KAUM–I.
78274, 26.7 mm SL, Kubura Fishing Port, hand net, 16 Sept.
2015; KAUM–I. 78277, 103.5 mm SL, KAUM–I. 78278, 94.9mm SL, KAUM–I. 78279, 93.5 mm SL, Kubura Fishing Port,line-fishing, 16 Sept. 2015.

Fig. 301. Dascyllus aruanus ミ ス ジ リ ュ ウ キ ュ ウ ス ズ メ ダ イ (upper:
KAUM–I. 79476, 31.4 mm SL; lower: Diving Service MARLIN)

Dascyllus reticulatus (Richardson, 1846)
フタスジリュウキュウスズメダイ [Marine] (Fig. 302)
KAUM–I. 78478, 17.4 mm SL, Sonai Channel, hand net,
21 Sept. 2015.

Fig. 302. Dascyllus reticulatus フ タ ス ジ リ ュ ウ キ ュ ウ ス ズ メ ダ イ(KAUM–I. 78478, 17.4 mm SL).

Fig. 303. Dascyllus trimaculatus ミ ツ ボ シ ク ロ ス ズ メ ダ イ (upper:
KAUM–I. 78277, 103.5 mm SL; middle: KAUM–I. 78274, 26.7 mm
SL; lower: piebald individual, Diving Service MARLIN).

69

K. Koeda et al.
Dischistodus prosopotaenia (Bleeker, 1852)
ダンダラスズメダイ [Marine]
Ito (2009).

Neoglyphidodon melas (Valenciennes, 1830)
クロスズメダイ [Marine] (Fig. 304)
KAUM–I. 78517, 14.3 mm SL, Hikawa, hand net, 21
Sept. 2015.

Fig. 306. Plectroglyphidodon lacrymatus ル リ ホ シ ス ズ メ ダ イ (FRLM
51425, 79.6 mm SL).

Fig. 304. Neoglyphidodon melas クロスズメダイ (KAUM–I. 78517, 14.3mm SL).

Plectroglyphidodon imparipennis (Vaillant and Sauvage, 1875)
イワサキスズメダイ [Marine] (Fig. 305)
KAUM–I. 78521, 35.2 mm SL, KAUM–I. 78522, 33.6mm SL, Umabana, hand net, 22 Sept. 2015; NSMT-P 55483,
2 specimens, 30.9–34.4 mm SL, Agarizaki, hand net, 28 Mar.
1997.

Plectroglyphidodon leucozonus (Bleeker, 1859)
ハクセンスズメダイ [Marine]
NSMT-P 55486, 8 specimens, 23.7–48.2 mm SL,
Agarizaki, hand net, 28 Mar. 1997; NSMT-P 55491, 52.5 mm
SL, Agarizaki, hand net, 28 Mar. 1997.

Pomacentrus amboinensis Bleeker, 1868
ニセネッタイスズメダイ [Marine] (Fig. 307)
FRLM 50728, 88.5 mm SL, Kubura Fishing Port, linefishing, 12 Mar. 2015; FRLM 51428, 89.6 mm SL, Kubura
Fishing Port, line-fishing, 19 Sept. 2015; KAUM–I. 78356,
81.7 mm SL, Kubura Fishing Port, line-fishing, 18 Sept. 2015.

Fig. 307. Pomacentrus amboinensis ニセネッタイスズメダイ (KAUM–I.
78356, 81.7 mm SL).
Fig. 305. Plectroglyphidodon imparipennis イワサキスズメダイ (KAUM–
I. 78521, 35.2 mm SL).

Pomacentrus coelestis Jordan and Starks, 1901
ソラスズメダイ [Marine] (Fig. 308)

Plectroglyphidodon lacrymatus (Quoy and Gaimard, 1825)
ルリホシスズメダイ [Marine] (Fig. 306)
FRLM 51425, 79.6 mm SL, Kubura Fishing Port, linefishing, 19 Sept. 2015.
Fig. 308. Pomacentrus coelestis ソ ラ ス ズ メ ダ イ (Diving Service
MARLIN).

70

Checklist of fishes of Yonaguni-jima island
Pomacentrus philippinus Evermann and Seale, 1907
フィリピンスズメダイ [Marine] (Fig. 309)
Ito (2009).
FRLM 50696, 73.3 mm SL, Sonai Fishing Port, linefishing, 13 Mar. 2015; FRLM 51431, 77.2 mm SL, FRLM
52006, 81.8 mm SL, Sonai Fishing Port, line-fishing, 19 Sept.
2015; FRLM 51486, 86.2 mm SL, Sonai Fishing Port, castnet, 22 Sept. 2015.

Pomacentrus sp. 2
ソラスズメダイ属の一種 [Marine] (Fig. 311)
KAUM–I. 78559, 67.8 mm SL, off Hikawa, line-fishing,
22 Sept. 2015.

Fig. 311. Pomacentrus sp. 2 (KAUM–I. 78559, 67.8 mm SL).

Fig. 309. Pomacentrus philippinus フィリピンスズメダイ (FRLM 50696,
73.3 mm SL).

Pomacentrus sp. 1
ミナミイソスズメダイ [Marine] (Fig. 310)
Ito (2009).
FRLM 52020, 68.7 mm SL, FRLM 52021, 76.7 mm SL,
Kubura Fishing Port, line-fishing, 19 Sept. 2015; FRLM
52051, 75.7 mm SL, FRLM 52052, 76.4 mm SL, FRLM
52053, 65.4 mm SL, Sonai Fishing Port, line-fishing, 24 Sept.
2015.

Stegastes altus (Okada and Ikeda, 1937)
セダカスズメダイ [Marine]
NSMT-P 55488, Agarizaki, 28 Mar. 1997.
Stegastes fasciolatus (Ogilby, 1889)
フチドリスズメダイ [Marine] (Fig. 312)
FRLM 51482, 83.3 mm SL, Southern Pasture, line-fishing,
22 Sept. 2015; KAUM–I. 78560, 60.7 mm SL, off Hikawa,line-fishing, 22 Sept. 2015.

Fig. 310. Pomacentrus sp. 1 ミナミイソスズメダイ (FRLM 52020, 68.7mm SL).

Fig. 312. Stegastes fasciolatus フチドリスズメダイ (upper: FRLM 51482,
83.3 mm SL; lower: KAUM–I. 78560, 60.7 mm SL).

71

K. Koeda et al.
Stegastes nigricans (Lacepède, 1802)
クロソラスズメダイ [Marine] (Fig. 313)
Ito (2009).
FRLM 51541, 108.9 mm SL, FRLM 52047, 108.8 mm SL,
FRLM 52048, 112.3 mm SL, Sonai Fishing Port, line-fishing,
24 Sept. 2015; KAUM–I. 78257, 102.6 mm SL, KAUM–I.
78262, 114.0 mm SL, Kubura Fishing Port, spear, 16 Sept.
2015; KAUM–I. 78506, 17.4 mm SL, Hikawa, hand net, 21
Sept. 2015.

Stegastes punctatus (Quoy and Gaimard, 1825)
ハナナガスズメダイ [Marine; brackish] (Fig. 314)
FRLM 51426, 91.5 mm SL, Kubura Fishing Port, linefishing, 19 Sept. 2015; FRLM 51523, 77.2 mm SL, mouthof Kubura River, hand net, 23 Sept. 2015; FRLM 51533,
9.7 mm SL, Sonai Fishing Port, fishing light, 24 Sept. 2015;
FRLM 51534, 2 specimens, 8.9–9.2 mm SL, Sonai Fishing
Port, fishing light, 24 Sept. 2015; FRLM 51553, 20.0 mm SL,
Kubura Fishing Port, hand net, 24 Sept. 2015.

Fig. 314. Stegastes punctatus ハナナガスズメダイ (FRLM 51426, 91.5mm SL).

FAMILY POLYNEMIDAE
ツバメコノシロ科
Polynemidae sp.
ツバメコノシロ科の一種 [Marine; brackish]
はにまち
Ikema (2003).

FAMILY LABRIDAE
ベラ科
くっつぁび
Anampses caeruleopunctatus Rüppell, 1829
ブチススキベラ [Marine] (Fig. 315)
KAUM–I. 80223, 208.5 mm SL, Arakawa Bay, spear, 18
Sept. 2015.

Fig. 313. Stegastes nigricans クロソラスズメダイ (upper: FRLM 51541,
108.9 mm SL; middle: KAUM–I. 78257, 102.6 mm SL; lower:
KAUM–I. 78506, 17.4 mm SL).

Fig. 315. Anampses caeruleopunctatus ブチススキベラ (KAUM–I. 80223,
208.5 mm SL).

72

Checklist of fishes of Yonaguni-jima island
Anampses twistii Bleeker, 1856
ホシススキベラ [Marine] (Fig. 316)
FRLM 50702, 135.0 mm SL, Sonai Fishing Port, linefishing, 13 Mar. 2015.

Bodianus perditio (Quoy and Gaimard, 1834)
タキベラ [Marine] (Fig. 320)
FRLM 51457, 202.5 mm SL, off Hikawa, line-fishing, 21
Sept. 2015; KAUM–I. 80397, 378.9 mm SL, off Hikawa, linefishing, 21 Sept. 2015.

Fig. 316. Anampses twistii ホシススキベラ (FRLM 50702, 135.0 mm SL).

Bodianus axillaris (Bennett, 1832)
スミツキベラ [Marine] (Fig. 317)

Fig. 320. Bodianus perditio タキベラ (FRLM 51457, 202.5 mm SL).

Cheilinus chlorourus (Bloch, 1791)
アカテンモチノウオ [Marine] (Fig. 321)
KAUM–I. 78514, 28.3 mm SL, Hikawa, hand net, 21
Sept. 2015; KAUM–I. 80281, 113.5 mm SL, Sonai Fishing
Port, line-fishing, 20 Sept. 2015.

Fig. 317. Bodianus axillaris スミツキベラ (Umabana, K. Koeda).

Bodianus bilunulatus (Lacepède, 1801)
キツネベラ [Marine] (Fig. 318)
FRLM 51456, 148.6 mm SL, off Hikawa, line-fishing, 21
Sept. 2015; KAUM–I. 80418, 182.8 mm SL, off Hikawa, linefishing, 21 Sept. 2015.

Fig. 321. Cheilinus chlorourus ア カ テ ン モ チ ノ ウ オ (upper: KAUM–I.
80281, 113.5 mm SL; lower: KAUM–I. 78514, 28.3 mm SL).
Fig. 318. Bodianus bilunulatus キツネベラ (FRLM 51456, 148.6 mm SL).

Bodianus dictynna Gommon, 2006
モンツキベラ [Marine] (Fig. 319)
KAUM–I. 78551, 138.0 mm SL, Umabana, spear, 22 Sept.
2015.

Fig. 319. Bodianus dictynna モンツキベラ (KAUM–I. 78551, 138.0 mm SL).

Cheilinus trilobatus Lacepède, 1801
ミツバモチノウオ [Marine] (Fig. 322)
FRLM 51568, 162.2 mm SL, Kubura Fishing Port, linefishing, 24 Sept. 2015.

Fig. 322. Cheilinus trilobatus ミ ツ バ モ チ ノ ウ オ (FRLM 51568, 162.2mm SL).

73

K. Koeda et al.
Cheilinus undulatus Rüppell, 1835
メガネモチノウオ [Marine] (Fig. 323)

Coris aygula Lacepède, 1801
カンムリベラ [Marine] (Fig. 327)
Yoshigou et al. (2001).
FRLM 50762, 232.5 mm SL, Kubura-bari, line-fishing, 13
Mar. 2015; FRLM 52041, 206.5 mm SL, FRLM 52042, 265.0
mm SL, Kubura-bari, line-fishing, 22 Sept. 2015; HMNH-P
112, 41.8 mm SL, Agarizaki, hand net, 9 Aug. 1999; KAUM–I.
78605, 234.0 mm SL, off Hikawa, line-fishing, 21 Sept. 2015.

Fig. 323. Cheilinus undulatus メ ガ ネ モ チ ノ ウ オ (Underwater Ruin, K.
Koeda).

Cheilio inermis (Forsskål, 1775)
カマスベラ [Marine; brackish] (Fig. 324)
KAUM–I. 78509, 38.4 mm SL, Hikawa, hand net, 21
Sept. 2015.

Fig. 327. Coris aygula カンムリベラ (FRLM 50762, 232.5 mm SL).

Coris batuensis (Bleeker, 1856)
シチセンムスメベラ [Marine] (Fig. 328)
Yoshigou et al. (2001).
HMNH-P111, 24.2 mm SL, Agarizaki, hand net, 9 Aug. 1999.
Fig. 324. Cheilio inermis カマスベラ (KAUM–I. 78509, 38.4 mm SL).

Choerodon jordani (Snyder, 1908)
クラカケベラ [Marine] (Fig. 325)
FRLM 51459, 130.7 mm SL, off Hikawa, line-fishing, 21
Sept. 2015.

Fig. 328. Coris batuensis シチセンムスメベラ (HMNH-P111, 24.2 mm
SL, presereved specimen).

Coris dorsomacula Fowler, 1908
スジベラ [Marine] (Fig. 329)
FRLM 51461, 114.1 mm SL, FRLM 51462, 103.8 mm
SL, off Hikawa, line-fishing, 21 Sept. 2015; KAUM–I. 80456,
86.7 mm SL, KAUM–I. 80492, 112.1 mm SL, KAUM–I.
80493, 112.1 mm SL, north of Yonaguni-jima island, linefishing, 22 Sept. 2015.
Fig. 325. Choerodon jordani クラカケベラ (FRLM 51459, 130.7 mm SL).

Cirrhilabrus cyanopleura (Bleeker, 1851)
クロヘリイトヒキベラ [Marine] (Fig. 326)

Fig. 326. Cirrhilabrus cyanopleura ク ロ ヘ リ イ ト ヒ キ ベ ラ (Diving
Service MARLIN).

74

Fig. 329. Coris dorsomacula スジベラ (upper: KAUM–I. 80456, 86.7 mm
SL; lower: KAUM–I. 80493, 112.1 mm SL).

Checklist of fishes of Yonaguni-jima island
Epibulus insidiator (Pallas, 1770)
ギチベラ [Marine] (Fig. 330)
Ito (2009).
FRLM 51569, 164.8 mm SL, Kubura Fishing Port, linefishing, 24 Sept. 2015; KAUM–I. 78275, 159.9 mm SL,
KAUM–I. 78276, 134.8 mm SL, Kubura Fishing Port, linefishing, 16 Sept. 2015; KAUM–I. 78312, 192.2 mm SL,
KAUM–I. 78313, 242.0 mm SL, Rakuda Rock, spear, 17
Sept. 2015; KAUM–I. 78507, 21.4 mm SL, KAUM–I. 78508,
10.2 mm SL, Hikawa, hand net, 21 Sept. 2015.

Halichoeres hortulanus (Lacepède, 1801)
トカラベラ [Marine] (Fig. 331)
FRLM 50751, 225.0 mm SL, Sonai Fishing Port, linefishing, 14 Mar. 2015; HMNH-P110, 26.6 mm SL, Agarizaki,
hand net, 9 Aug. 1999; KAUM–I. 78368, 12.0 mm SL,
Arakawa Bay, hand net, 18 Sept. 2015; NSMT-P 55470, 65.1
mm SL, Agarizaki, hand net, 28 Mar. 1997.

Fig. 331. Halichoeres hortulanus トカラベラ (upper: FRLM 50751, 225.0mm SL, inverted; middle: KAUM–I. 78368, 12.0 mm SL; lower:
Diving Service MARLIN).

Halichoeres margaritaceus (Valenciennes, 1839)
アカニジベラ [Marine]
NSMT-P 55469, 19.6 mm SL, Agarizaki, hand net, 28
Mar. 1997.

Fig. 330. Epibulus insidiator ギ チ ベ ラ (upper to lower: FRLM 51569,
164.8 mm SL; KAUM–I. 78313, 242.0 mm SL; KAUM–I. 78312,
192.2 mm SL; KAUM–I. 78507, 21.4 mm SL).

Halichoeres marginatus Rüppell, 1835
カノコベラ [Marine] (Fig. 332)
Yoshigou et al. (2001).
FRLM 51560, 128.9 mm SL, Sonai Fishing Port, linefishing, 24 Sept. 2015; KAUM–I. 78254, 170.8 mm SL,
Kubura Fishing Port, spear, 16 Sept. 2015; NSMT-P 55471,
76.3 mm SL, Agarizaki, hand net, 28 Mar. 1997.

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K. Koeda et al.
Halichoeres trimaculatus (Quoy and Gaimard, 1834)
ミツボシキュウセン [Marine] (Fig. 334)
Ito (2009).
KAUM–I. 80352, 127.5 mm SL, Sonai Channel, spear, 21
Sept. 2015.

Fig. 334. Halichoeres trimaculatus ミ ツ ボ シ キ ュ ウ セ ン (KAUM–I.
80352, 127.5 mm SL).

Hemigymnus fasciatus (Bloch, 1792)
シマタレクチベラ [Marine] (Fig. 335)
KAUM–I. 78556, 115.0 mm SL, Umabana, hand net, 22
Sept. 2015.

Fig. 332. Halichoeres marginatus カノコベラ (upper: FRLM 51560, 128.9mm SL; lower: KAUM–I. 78254, 170.8 mm SL).

Halichoeres melanochir Fowler and Bean, 1928
ムナテンベラ [Marine] (Fig. 333)
FRLM 50740, 170.1 mm SL, Sonai Fishing Port, linefishing, 15 Mar. 2015.

Fig. 335. Hemigymnus fasciatus シ マ タ レ ク チ ベ ラ (KAUM–I. 78556,
115.0 mm SL).

Hemigymnus melapterus (Bloch, 1791)
タレクチベラ [Marine] (Fig. 336)
KAUM–I. 78469, 44.2 mm SL, Sonai Channel, hand net,
21 Sept. 2015.

Fig. 333. Halichoeres melanochir ムナテンベラ (FRLM 50740, 170.1 mm SL).

Halichoeres nebulosus (Valenciennes, 1839)
イナズマベラ [Marine]
URM-P 34364, Danno Beach, 2 Aug. 1995.
Halichoeres orientalis Randall, 1999
ツキベラ [Marine]
Ito (2009).

Fig. 336. Hemigymnus melapterus タレクチベラ (KAUM–I. 78469, 44.2mm SL).

Hologymnosus doliatus (Lacepède, 1801)
シロタスキベラ [Marine] (Fig. 337)
FRLM 50743, 226.5 mm SL, Sonai Fishing Port, linefishing, 15 Mar. 2015.

Halichoeres prosopeion (Bleeker, 1853)
ムナテンベラダマシ [Marine]
Ito (2009).
Halichoeres richmondi Fowler and Bean, 1928
ゴシキキュウセン [Marine]
Ito (2009).

76

Fig. 337. Hologymnosus doliatus シロタスキベラ (FRLM 50743, 226.5mm SL).

Checklist of fishes of Yonaguni-jima island
Iniistius celebicus (Bleeker, 1856)
ホウキボシテンス [Marine] (Fig. 338)

Fig. 338. Iniistius celebicus ホウキボシテンス (Diving Service MARLIN).

Iniistius pavo (Valenciennes, 1840)
ホシテンス [Marine] (Fig. 339)
Fig. 341. Novaculichthys taeniourus オビテンスモドキ (KAUM–I. 78369,
12.8 mm SL).

Oxycheilinus orientalis (Günther, 1862)
カタグロホホスジモチノウオ [Marine] (Fig. 342)
FRLM 51464, 106.2 mm SL, off Hikawa, line-fishing, 21
Sept. 2015.

Fig. 339. Iniistius pavo ホシテンス (Diving Service MARLIN).

Labroides dimidiatus (Valenciennes, 1839)
ホンソメワケベラ [Marine] (Fig. 340)
KAUM–I. 78332, 23.0 mm SL, Anchor's graves, handnet, 17 Sept. 2015; KAUM–I. 78479, 21.7 mm SL, KAUM–I.
78480, 24.0 mm SL, Sonai Channel, hand net, 21 Sept. 2015;
NSMT-P 55473, 41.3 mm SL, Agarizaki, hand net, 28 Mar.
1997.

Fig. 342. Oxycheilinus orientalis カタグロホホスジモチノウオ (FRLM
51464, 106.2 mm SL).

Oxycheilinus unifasciatus (Streets, 1877)
ヒトスジモチノウオ [Marine] (Fig. 343)
FRLM 50744, 176.9 mm SL, Sonai Fishing Port, linefishing, 15 Mar. 2015; KAUM–I. 79741, 183.5 mm SL,
KAUM–I. 80232, 183.2 mm SL, Rakuda Rock, line-fishing, 17
Sept. 2015; KAUM–I. 80231, 162.4 mm SL, Arakawa Bay, linefishing, 17 Sept. 2015; KAUM–I. 80478, 148.2 mm SL, Danno
Beach, spear, 23 Sept. 2015; MUFS 46894, 155.0 mm SL, off
Kubura Hinan Port, line-fishing, 18 Sept. 2015; MUFS 46978,
196.0 mm SL, off Hikawa, line-fishing, 21 Sept. 2015; NSMT-P
101127, 132.6 mm SL, Danno Beach, 20 Mar. 2010.

Fig. 340. Labroides dimidiatus ホ ン ソ メ ワ ケ ベ ラ (upper: KAUM–I.
78480, 24.0 mm SL; lower: KAUM–I. 78332, 23.0 mm SL).

Novaculichthys taeniourus (Lacepède, 1801)
オビテンスモドキ [Marine] (Fig. 341)
KAUM–I. 78369, 12.8 mm SL, Arakawa Bay, hand net,
18 Sept. 2015.

Fig. 343. Oxycheilinus unifasciatus ヒトスジモチノウオ (upper: KAUM–
I. 80232, 183.2 mm SL; lower: MUFS 46978, 196.0 mm SL).

77

K. Koeda et al.
Pseudocheilinus hexataenia (Bleeker, 1857)
ニセモチノウオ [Marine] (Fig. 344)
KAUM–I. 78462, 37.1 mm SL, KAUM–I. 78486, 15.4mm SL, Sonai Channel, hand net, 21 Sept. 2015.

Fig. 347. Thalassoma amblycephalum コガシラベラ (FRLM 51463, 80.5mm SL).

Fig. 344. Pseudocheilinus hexataenia ニセモチノウオ (KAUM–I. 78462,
37.1 mm SL).

Thalassoma hardwicke (Bennett, 1830)
セナスジベラ [Marine] (Fig. 348)
FRLM 50741, 136.7 mm SL, FRLM 50742, 138.6 mm
SL, Sonai Fishing Port, line-fishing, 15 Mar. 2015; FRLM
51542, 91.1 mm SL, Sonai Fishing Port, line-fishing, 24 Sept.
2015; FRLM 52014, 147.1 mm SL, Kubura Fishing Port, linefishing, 21 Sept. 2015; NSMT-P 55466, 4 specimens, 50.1–
80.1 mm SL, Agarizaki, hand net, 28 Mar. 1997.

Stethojulis bandanensis (Bleeker, 1851)
アカオビベラ [Marine] (Fig. 345)
FRLM 51540, 12.0 mm SL, Kubura Fishing Port, handnet, 24 Sept. 2015.

Fig. 348. Thalassoma hardwicke セナスジベラ (FRLM 51542, 91.1 mm SL).

Fig. 345. Stethojulis bandanensis アカオビベラ (FRLM 51540, 12.0 mm SL).

Stethojulis strigiventer (Bennett, 1833)
ハラスジベラ [Marine]
NSMT-P 55467, 23.9 mm SL, Agarizaki, hand net, 28
Mar. 1997.

Thalassoma jansenii (Bleeker, 1856)
ヤンセンニシキベラ [Marine] (Fig. 349)
Ito (2009).
FRLM 51448, 119.5 mm SL, Kubura-bari, line-fishing, 20
Sept. 2015.

Stethojulis trilineata (Bloch and Schneider, 1801)
オニベラ [Marine] (Fig. 346)
KAUM–I. 78398, 26.4 mm SL, Hikawa, hand net, 20
Sept. 2015.
Fig. 349. Thalassoma jansenii ヤンセンニシキベラ (FRLM 51448, 119.5mm SL).

Fig. 346. Stethojulis trilineata オニベラ (KAUM–I. 78398, 26.4 mm SL).

Thalassoma amblycephalum (Bleeker, 1856)
コガシラベラ [Marine] (Fig. 347)
Yoshigou et al. (2001).
FRLM 51463, 80.5 mm SL, off Hikawa, line-fishing, 21
Sept. 2015; HMNH-P107, 23.0 mm SL, Agarizaki, hand net,
9 Aug. 1999; NSMT-P 55474, 32.0 mm SL, Agarizaki, handnet, 28 Mar. 1997.

78

Thalassoma lutescens (Lay and Bennett, 1839)
ヤマブキベラ [Marine] (Fig. 350)
Yoshigou et al. (2001).
FRLM 51989, 149.5 mm SL, FRLM 51990, 87.4 mm
SL, Sonai Fishing Port, line-fishing, 21 Sept. 2015; KAUM–
I. 78280, 129.3 mm SL, Kubura Fishing Port, line-fishing, 16
Sept. 2015; KAUM–I. 80213, 113.6 mm SL, Kubura Fishing
Port, line-fishing, 18 Sept. 2015; KAUM–I. 80489, 131.5 mm
SL, north of Yonaguni-jima island, line-fishing, 22 Sept. 2015;
MUFS 46893, 123.0 mm SL, off Kubura Hinan Port, linefishing, 18 Sept. 2015; MUFS 47024, 120.0 mm SL, Sonai
Fishing Port, line-fishing, 23 Sept. 2015; NSMT-P 55468, 9
specimens, 16.0–65.9 mm SL, Agarizaki, hand net, 28 Mar.
1997; NSMT-P 101232, 94.6 mm SL, Kubura Hinan Port,
hand net, 16 Mar. 2010.

Checklist of fishes of Yonaguni-jima island
Thalassoma quinquevittatum (Lay and Bennett, 1839)
ハコベラ [Marine] (Fig. 352)
FRLM 51539, 18.3 mm SL, Kubura Fishing Port, hand net,
24 Sept. 2015; FRLM 52038, 104.8 mm SL, FRLM 52039,
100.6 mm SL, FRLM 52040, 97.3 mm SL, Kubura-bari, linefishing, 22 Sept. 2015; FRLM 52070, 97.0 mm SL, FRLM
52071, 109.9 mm SL, Kubura Fishing Port, line-fishing, 24
Sept. 2015; KAUM–I. 80212, 118.5 mm SL, Kubura Fishing
Port, line-fishing, 18 Sept. 2015; KAUM–I. 80356, 98.5 mm
SL, Kubura-bari, line-fishing, 20 Sept. 2015; MUFS 46920,
101.0 mm SL, levee at Kubura, line-fishing, 18 Sept. 2015.

Fig. 352. Thalassoma quinquevittatum ハコベラ (KAUM–I. 80212, 118.5mm SL).

Fig. 350. Thalassoma lutescens ヤマブキベラ (upper: FRLM 51989, 149.5mm SL; middle: KAUM–I. 80213, 113.6 mm SL; lower: MUFS 46893,
123.0 mm SL).

Thalassoma purpureum (Forsskål, 1775)
キヌベラ [Marine] (Fig. 351)
Yoshigou et al. (2001).
FRLM 50763, 196.0 mm SL, Kubura-bari, line-fishing, 13
Mar. 2015; FRLM 51955, 138.0 mm SL, Kubura-bari, linefishing, 20 Sept. 2015; FRLM 52044, 134.6 mm SL, Kuburabari, line-fishing, 22 Sept. 2015; FRLM 52083, 203.0 mm SL,
Kubura Fishing Port, line-fishing, 21 Sept. 2015; KAUM–
I. 78384, 29.0 mm SL, Southern Pasture, hand net, 19 Sept.
2015.

Thalassoma trilobatum (Lacepède, 1801)
リュウグウベラ [Marine] (Fig. 353)
FRLM 51956, 194.5 mm SL, Kubura-bari, line-fishing, 20
Sept. 2015; FRLM 52037, 180.5 mm SL, Kubura-bari, linefishing, 22 Sept. 2015; KAUM–I. 80359, 198.1 mm SL,
Kubura-bari, line-fishing, 20 Sept. 2015; KAUM–I. 80450,
192.2 mm SL, Kubura-bari, line-fishing, 22 Sept. 2015.

Fig. 353. Thalassoma trilobatum リュウグウベラ (KAUM–I. 80450, 192.2mm SL).

Thalassoma sp.
ニシキベラ属の一種 [Marine]
NSMT-P 55472, 17 specimens, 13.6–42.8 mm SL,
Agarizaki, hand net, 28 Mar. 1997.
Wetmorella nigropinnata (Seale, 1901)
ハシナガベラ [Marine] (Fig. 354)
KAUM–I. 78333, 47.7 mm SL, Rakuda Rock, hand net,
17 Sept. 2015.

Fig. 351. Thalassoma purpureum キ ヌ ベ ラ (upper: FRLM 52083, 203.0mm SL; lower: KAUM–I. 78384, 29.0 mm SL).

Fig. 354. Wetmorella nigropinnata ハシナガベラ (KAUM–I. 78333, 47.7mm SL).

79

K. Koeda et al.
FAMILY SCARIDAE
ブダイ科
Bolbometopon muricatum (Valenciennes, 1840)
カンムリブダイ [Marine] (Fig. 355)
Ito (2009).

Fig. 355. Bolbometopon muricatum カ ン ム リ ブ ダ イ (Diving Service
MARLIN).

Calotomus carolinus (Valenciennes, 1840)
タイワンブダイ [Marine] (Fig. 356)
MUFS 46923, 195.0 mm SL, levee at Kubura, line-fishing,
18 Sept. 2015.

Fig. 357. Cetoscarus bicolor イロブダイ (upper: KAUM–I. 78460, 47.5mm SL; middle: KAUM–I. 78461, 12.9 mm SL; lower: Diving Service
MARLIN).

Chlorurus bowersi (Snyder, 1909)
オオモンハゲブダイ [Marine] (Fig. 358)
KAUM–I. 78331, 200.9 mm SL, Rakuda Rock, spear, 17
Sept. 2015.

Fig. 356. Calotomus carolinus タイワンブダイ (MUFS 46923, 195.0 mm SL).

Cetoscarus bicolor (Rüppell, 1829)
イロブダイ [Marine] (Fig. 357)
Senou et al. (1997): piebald individual.
KAUM–I. 78460, 47.5 mm SL, KAUM–I. 78461, 12.9mm SL, Sonai Channel, hand net, 21 Sept. 2015.

Fig. 358. Chlorurus bowersi オ オ モ ン ハ ゲ ブ ダ イ (KAUM–I. 78331,
200.9 mm SL).

Chlorurus microrhinos (Bleeker, 1854)
ナンヨウブダイ [Marine]
Ito (2009).

Scarus dimidiatus Bleeker, 1859
カワリブダイ [Marine]
Ito (2009).

80Checklist of fishes of Yonaguni-jima island
Scarus festivus Valenciennes, 1840
ツキノワブダイ [Marine] (Fig. 359)
KAUM–I. 80475, 282.0 mm SL, Danno Beach, spear, 23
Sept. 2015.

Fig. 361. Scarus niger ブチブダイ (FRLM 50752, 255.5 mm SL).

Fig. 359. Scarus festivus ツキノワブダイ (KAUM–I. 80475, 282.0 mm SL).

Scarus forsteni (Bleeker, 1861)
イチモンジブダイ [Marine] (Fig. 360)
Ito (2009).
FRLM 50745, 205.6 mm SL, Kubura-bari, line-fishing, 14
Mar. 2015; FRLM 52098, 207.7 mm SL, Southern Pasture,line-fishing, 21 Sept. 2015; KAUM–I. 80267, 214.7 mm SL,
Anchor's graves, spear, 17 Sept. 2015; KAUM–I. 80476,
266.0 mm SL, KAUM–I. 80477, 214.5 mm SL, Danno Beach,spear, 23 Sept. 2015.

Scarus psittacus Forsskål, 1775
オウムブダイ [Marine] (Fig. 362)
FRLM 50695, 138.4 mm SL, Sonai Fishing Port, linefishing, 13 Mar. 2015; FRLM 50723, 139.1 mm SL, Kubura
Fishing Port, line-fishing, 12 Mar. 2015; FRLM 51551, 42.9mm SL, Kubura Fishing Port, hand net, 24 Sept. 2015; FRLM
51556, 145.0 mm SL, Kubura Fishing Port, line-fishing, 24
Sept. 2015; FRLM 52009, 140.8 mm SL, Sonai Fishing Port,line-fishing, 19 Sept. 2015; MUFS 46935, 147.0 mm SL,
Sonai Fishing Port, line-fishing, 19 Sept. 2015.

Fig. 362. Scarus psittacus オウムブダイ (upper: FRLM 50695, 138.4 mm
SL; lower: MUFS 46935: 147.0 mm SL).

Scarus rubroviolaceus Bleeker, 1847
ナガブダイ [Marine] (Fig. 363)
KAUM–I. 78371, 11.8 mm SL, Arakawa Bay, hand net, 18
Sept. 2015.

Fig. 360. Scarus forsteni イ チ モ ン ジ ブ ダ イ (upper: KAUM–I. 80476,
266.0 mm SL; middle: KAUM–I. 80267, 214.7 mm SL; lower:
KAUM–I. 80477, 214.5 mm SL).

Scarus niger Forsskål, 1775
ブチブダイ [Marine] (Fig. 361)
FRLM 50752, 255.5 mm SL, Sonai Fishing Port, linefishing, 14 Mar. 2015.

Fig. 363. Scarus rubroviolaceus ナガブダイ (KAUM–I. 78371, 11.8 mm SL).

81

K. Koeda et al.
Scarus schlegeli (Bleeker, 1861)
オビブダイ [Marine] (Fig. 364)
Yoshigou et al. (2001).
FRLM 50753, 235.5 mm SL, Sonai Fishing Port, linefishing, 14 Mar. 2015; HMNH-P113, 33.4 mm SL, Agarizaki,
hand net, 9 Aug. 1999; KAUM–I. 78395, 33.7 mm SL,
KAUM–I. 78396, 27.4 mm SL, KAUM–I. 78397, 15.3 mm
SL, Hikawa, hand net, 20 Sept. 2015; KAUM–I. 78513, 38.7mm SL, Hikawa, hand net, 21 Sept. 2015; KAUM–I. 80270,
198.5 mm SL, Kubura Fishing Port, line-fishing, 19 Sept.
2015; MUFS 46924, 182.0 mm SL, levee at Kubura, linefishing, 18 Sept. 2015.

FAMILY PINGUIPEDIDAE
トラギス科
Parapercis cylindrica (Bloch, 1792)
ダンダラトラギス [Marine] (Fig. 367)
KAUM–I. 78561, 81.6 mm SL, KAUM–I. 78562, 64.1
mm SL, off Hikawa, line-fishing, 22 Sept. 2015.

Fig. 367. Parapercis cylindrica ダ ン ダ ラ ト ラ ギ ス (upper: KAUM–I.
78561, 81.6 mm SL; lower: Diving Service MARLIN).

Fig. 364. Scarus schlegeli オ ビ ブ ダ イ (upper: KAUM–I. 80270, 198.5mm SL; lower: KAUM–I. 78396, 27.4 mm SL).

Scarus xanthopleura Bleeker, 1853
アカブダイ [Marine] (Fig. 365)
あがいらぶたIkema (2003).

Fig. 365. Scarus xanthopleura アカブダイ (Diving Service MARLIN).

Scaridae sp.
ブダイ科の一種 [Marine] (Fig. 366)
KAUM–I. 78300, 12.2 mm SL, Kubura Fishing Port, handnet, 16 Sept. 2015; KAUM–I. 78512, 27.4 mm SL, Hikawa,
hand net, 21 Sept. 2015.

Fig. 366. Scaridae sp. (KAUM–I. 78512, 27.4 mm SL).

82

Parapercis millepunctata (Günther, 1860)
ワヌケトラギス [Marine] (Fig. 368)
Ito (2009).
KAUM–I. 78380, 88.4 mm SL, Kubura Fishing Port, linefishing, 19 Sept. 2015; NSMT-P 101231, 112.7 mm SL,
Kubura Hinan Port, line-fishing, 16 Mar. 2010.

Fig. 368. Parapercis millepunctata ワ ヌ ケ ト ラ ギ ス (KAUM–I. 78380,
88.4 mm SL).

Parapercis pacifica Imamura and Yoshino, 2007
オグロトラギス [Marine] (Fig. 369)
FRLM 50726, 128.4 mm SL, male, Kubura Fishing Port,line-fishing, 12 Mar. 2015; FRLM 51565, 161.5 mm SL, male,
Kubura Fishing Port, line-fishing, 24 Sept. 2015; KAUM–I.
78306, 176.4 mm SL, Kataburu Beach, line-fishing, 17 Sept.
2015; KAUM–I. 78379, 169.5 mm SL, Kubura Fishing Port,line-fishing, 19 Sept. 2015; KAUM–I. 78441, 163.0 mm SL,
Kubura Fishing Port, line-fishing, 20 Sept. 2015.

Checklist of fishes of Yonaguni-jima island
Enneapterygius tutuilae Jordan and Seale, 1906
セグロヘビギンポ [Marine] (Fig. 372)
FRLM 51522, 17.3 mm SL, mouth of Kubura River, handnet, 23 Sept. 2015; KAUM–I. 78361, 17.5 mm SL, off Kubura
Hinan Port, hand net, 18 Sept. 2015; KAUM–I. 78549, 13.4mm SL, Umabana, hand net, 22 Sept. 2015.

Fig. 369. Parapercis pacifica オグロトラギス (upper: KAUM–I. 78441,
163.0 mm SL; lower: Diving Service MARLIN).

Parapercis tetracantha (Lacepède, 1801)
マダラトラギス [Marine] (Fig. 370)
FRLM 51480, 151.0 mm SL, male, off Northern Pasture,line-fishing, 22 Sept. 2015.

Fig. 372. Enneapterygius tutuilae セ グ ロ ヘ ビ ギ ン ポ (upper: KAUM–I.
78549, 13.4 mm SL; lower: KAUM–I. 78361, 17.5 mm SL).

Enneapterygius sp.
ヘビギンポ属の一種 [Marine] (Fig. 373)
KAUM–I. 78532, 19.2 mm SL, Umabana, hand net, 22
Sept. 2015.

Fig. 370. Parapercis tetracantha マ ダ ラ ト ラ ギ ス (FRLM 51480, 151.0mm SL).
Fig. 373. Enneapterygius sp. (KAUM–I. 78532, 19.2 mm SL).

FAMILY TRIPTERYGIIDAE
ヘビギンポ科
Enneapterygius philippinus (Peters, 1869)
クサギンポ [Marine]
URM-P 28692, Hikawa, 10 Oct. 1992.
Enneapterygius rhabdotus Fricke, 1994
クレナイヘビギンポ [Marine] (Fig. 371)
Yoshigou and Yoshino (2002); Yoshigou and Nakamura(2002); Nakabo (2013).
HMNH-P3077, 16.1 mm SL, HMNH-P3078, 17.2 mm SL,
Agarizaki, hand net, 9 Aug. 1999; NSMT-P 106821, 5
specimens, 13.6–21.0 mm SL, Agarizaki, hand net, 28 Mar.
1997.

Fig. 371. Enneapterygius rhabdotus クレナイヘビギンポ (HMNH-P3077,
16.1 mm SL, presereved specimen).

Helcogramma fuscipectoris (Fowler, 1946)
クロマスク [Marine] (Fig. 374)
Yoshigou et al. (2001).
HMNH-P1996, 6 specimens, 12.2–21.8 mm SL,
Agarizaki, hand net, 9 Aug. 1999.

Fig. 374. Helcogramma fuscipectoris クロマスク (HMNH-P1996, 21.8 mm
SL, presereved specimen).

Helcogramma hudsoni (Jordan and Seale, 1906)
ナナメヘビギンポ [Marine] (Fig. 375)
Yoshigou and Nakamura (2003).
HMNH-P7189, 2 specimens, 29.7–31.8 mm SL,
Agarizaki, hand net, 9 Aug. 1999.

Fig. 375. Helcogramma hudsoni ナナメヘビギンポ (HMNH-P7189, 31.8mm SL, presereved specimen).

83

K. Koeda et al.
Helcogramma inclinata (Fowler, 1946)
アヤヘビギンポ [Marine] (Fig. 376)
KAUM–I. 78370, 19.4 mm SL, Arakawa Bay, hand net,
18 Sept. 2015.

Fig. 379. Alticus saliens タマカエルウオ (KAUM–I. 78520, 63.0 mm SL).

Fig. 376. Helcogramma inclinata アヤヘビギンポ (KAUM–I. 78370, 19.4mm SL).

Helcogramma striata Hansen, 1986
タテジマヘビギンポ [Marine] (Fig. 377)
KAUM–I. 78537, 19.1 mm SL, Umabana, hand net, 22
Sept. 2015.

Andamia tetradactylus (Bleeker, 1858)
ヨダレカケ [Marine] (Fig. 380)
FRLM 51434, 82.7 mm SL, FRLM 51435, 83.2 mm SL,
Kubura Hinan Port, hand net, 20 Sept. 2015; KAUM–I.
78519, 85.4 mm SL, Sonai Fishing Port, hand net, 21 Sept.
2015; MUFS 47015, 87.0 mm SL, MUFS 47016, 37.0 mm
SL, MUFS 47017, 31.0 mm SL, MUFS 47018, 28.0 mm SL,
Sonai Fishing Port, hand net, 22 Sept. 2015.

Fig. 380. Andamia tetradactylus ヨダレカケ (KAUM–I. 78519, 85.4 mm SL).
Fig. 377. Helcogramma striata タテジマヘビギンポ (KAUM–I. 78537,
19.1 mm SL).

Blenniella interrupta (Bleeker, 1857)
ジュズダマギンポ [Marine]
Yoshigou and Yoshino (2002); Nakabo (2013).
FAMILY TRICHONOTIDAE
ベラギンポ科Trichonotus sp.
ベラギンポ属の一種 [Marine] (Fig. 378)

Cirripectes castaneus (Valenciennes, 1836)
タテガミカエルウオ [Marine] (Fig. 381)
KAUM–I. 78555, 21.1 mm SL, Umabana, hand net, 22
Sept. 2015; KAUM–I. 78571, 51.1 mm SL, Danno Beach,hand net, 23 Sept. 2015; MUFS 47029, 51.0 mm SL, Danno
Beach, hand net, 24 Sept. 2015.

Fig. 378. Trichonotus sp. (Diving Service MARLIN).

FAMILY BLENNIIDAE
イソギンポ科
Alticus saliens (Forster, 1788)
タマカエルウオ [Marine] (Fig. 379)
KAUM–I. 78520, 63.0 mm SL, Sonai Fishing Port, handnet, 21 Sept. 2015.

84

Fig. 381. Cirripectes castaneus タテガミカエルウオ (upper: KAUM–I.
78571, 51.1 mm SL; lower: KAUM–I. 78555, 21.1 mm SL).

Checklist of fishes of Yonaguni-jima island
Cirripectes sp.
タテガミカエルウオ属の一種 [Marine] (Fig. 382)
KAUM–I. 78541, 35.9 mm SL, Umabana, hand net, 22
Sept. 2015.

Glyptoparus delicatulus Smith, 1959
ヤイトギンポ [Marine] (Fig. 386)
MUFS 46883, 35.0 mm SL, Kataburu Beach, hand net,
17 Sept. 2015; MUFS 46932, 27.8 mm SL, Kataburu Beach,hand net, 18 Sept. 2015.

Fig. 386. Glyptoparus delicatulus ヤイトギンポ (MUFS 46932, 27.8 mm SL).
Fig. 382. Cirripectes sp. (KAUM–I. 78541, 35.9 mm SL).

Ecsenius lineatus Klausewitz, 1962
ヒトスジギンポ [Marine] (Fig. 383)
KAUM–I. 78570, 45.0 mm SL, Danno Beach, hand net,
23 Sept. 2015.

Istiblennius edentulus (Forster and Schneider, 1801)
ニセカエルウオ [Marine] (Fig. 387)
Yoshigou et al. (2001).
HMNH-P120, 76.3 mm SL, Agarizaki, hand net, 9 Aug.
1999; HMNH-P134, 76.6 mm SL, HMNH-P782, 52.1 mm
SL, Sonai, hand net, 9 Aug. 1999; NSMT-P 94374, 5
specimens, 63.1–96.1 mm SL, Agarizaki, hand net, 28 Mar.
1997.

Fig. 383. Ecsenius lineatus ヒトスジギンポ (KAUM–I. 78570, 45.0 mm SL).

Ecsenius oculus Springer, 1971
ゴイシギンポ [Marine] (Fig. 384)
KAUM–I. 78569, 26.7 mm SL, Danno Beach, hand net,
23 Sept. 2015.

Fig. 387. Istiblennius edentulus ニセカエルウオ (HMNH-P782, 52.1 mm
SL, preserved specimen).

Meiacanthus grammistes (Valenciennes, 1836)
ヒゲニジギンポ [Marine]
Ito (2009).
Fig. 384. Ecsenius oculus ゴイシギンポ (KAUM–I. 78569, 26.7 mm SL).

Entomacrodus niuafoouensis (Fowler, 1932)
ケショウギンポ [Marine] (Fig. 385)
FRLM 51483, 92.1 mm SL, FRLM 51488, 72.6 mm SL,
FRLM 51489, 56.7 mm SL, Southern Pasture, hand net, 22
Sept. 2015; KAUM–I. 78518, 90.6 mm SL, Southern Pasture,
hand net, 21 Sept. 2015.

Nannosalarias nativitatis (Regan, 1909)
ヒナギンポ [Marine] (Fig. 388)
MUFS 46941, 29.9 mm SL, MUFS 46942, 28.0 mm SL,
Danno Beach, hand net, 19 Sept. 2015; MUFS 46943, Danno
Beach, hand net, 19 Sept. 2015; MUFS 47030, 32.0 mm SL,
MUFS 47031, MUFS 47032, 38.0 mm SL, Danno Beach,hand net, 24 Sept. 2015.

Fig. 388. Nannosalarias nativitatis ヒナギンポ (MUFS 46942, 28.0 mm SL).

Fig. 385. Entomacrodus niuafoouensis ケショウギンポ (upper: KAUM–I.
78518, 90.6 mm SL; lower: FRLM 51483, 92.1 mm SL).

85

K. Koeda et al.
Plagiotremus rhinorhynchos (Bleeker, 1852)
ミナミギンポ [Marine] (Fig. 389)

Rhabdoblennius nitidus (Günther, 1861)
ロウソクギンポ [Marine] (Fig. 392)
MUFS 46931, 43.8 mm SL, Kataburu Beach, hand net, 18
Sept. 2015; MUFS 46945, 40.1 mm SL, Danno Beach, handnet, 19 Sept. 2015.

Fig. 392. Rhabdoblennius nitidus ロ ウ ソ ク ギ ン ポ (MUFS 46931, 43.8mm SL).
Fig. 389. Plagiotremus rhinorhynchos ミ ナ ミ ギ ン ポ (Diving Service
MARLIN).

Praealticus bilineatus (Peters, 1868)
タマギンポ [Marine] (Fig. 390)
KAUM–I. 78383, 24.0 mm SL, Southern Pasture, handnet, 19 Sept. 2015; MUFS 46947, 35.0 mm SL, Danno
Beach, hand net, 19 Sept. 2015; MUFS 47033, 33.0 mm SL,
Yonaguni-jima island, hand net, 24 Sept. 2015; NSMT-P
94376, 3 specimens, 43.2–52.1 mm SL, Agarizaki, hand net,
28 Mar. 1997.

FAMILY CALLIONYMIDAE
ネズッポ科
Neosynchiropus moyeri (Zaiser and Fricke, 1985)
ミヤケテグリ [Marine] (Fig. 393)

Fig. 390. Praealticus striatus タマギンポ (KAUM–I. 78383, 24.0 mm SL).

Praealticus striatus (Bath, 1992)
カブキギンポ [Marine] (Fig. 391)
FRLM 51484, 52.0 mm SL, FRLM 51490, 50.2 mm SL,
Southern Pasture, hand net, 22 Sept. 2015; KAUM–I. 78385,
61.5 mm SL, male, KAUM–I. 78386, 50.6 mm SL, female,Southern Pasture, hand net, 19 Sept. 2015; MUFS 46946, 54.7mm SL, male, Danno Beach, hand net, 19 Sept. 2015.
Fig. 393. Neosynchiropus moyeri ミヤケテグリ (Diving Service MARLIN).

Fig. 391. Praealticus tanegasimae タネギンポ (upper: MUFS 46946, 54.7mm SL; lower: KAUM–I. 78386, 50.6 mm SL).

86

Checklist of fishes of Yonaguni-jima island
FAMILY ELEOTRIDAE
カワアナゴ科Eleotris acanthopoma Bleeker, 1853
チチブモドキ [Brackish; freshwater] (Fig. 394)
Yoshigou (2000); Yoshigou et al. (2001).
FRLM 51514, 27.9 mm SL, FRLM 51515, 50.4 mm SL,
Tabaru River, hand net, 23 Sept. 2015; FRLM 51527, 50.0mm SL, mouth of Kubura River, hand net, 23 Sept. 2015;
HMNH-P45, 62.4 mm SL, HMNH-P46, 69.0 mm SL, Tabaru
River, hand net, 9 Aug. 1999; HMNH-P102, 33.6 mm SL,
HMNH-P103, 80.8 mm SL, HMNH-P105, 41.5 mm SL,
Kubura River, hand net, 10 Aug. 1999; KAUM–I. 78413, 45.3mm SL, KAUM–I. 78454, 48.5 mm SL, KAUM–I. 78455,
30.7 mm SL, Tabaru River, hand net, 20 Sept. 2015.

Fig. 394. Eleotris acanthopoma チチブモドキ (upper: KAUM–I. 78413,
45.3 mm SL; lower: HMNH-P45, 62.4 mm SL).

Ophieleotris sp. 2
ゴシキタメトモハゼ [Brackish; freshwater]
Ministry of the Environment (2010a).
Ophiocara porocephala (Valenciennes, 1837)
ホシマダラハゼ [Brackish; freshwater] (Fig. 396)
Yoshigou (2000); Yoshigou et al. (2001); Japan Wild Life
Research Center (2002); Senou et al. (2004); Ministry of theEnvironment (2010a); Ministry of the Environment (2010b).
FRLM 51507, 23.1 mm SL, Tabaru River, hand net, 20
Sept. 2015; HMNH-P54, 81.1 mm SL, Tabaru River, hand net,
9 Aug. 1999; KAUM–I. 78451, 28.3 mm SL, Tabaru River,gill net, 20 Sept. 2015; KAUM–I. 78453, 10.9 mm SL, Tabaru
River, hand net, 20 Sept. 2015.

Fig. 396. Ophiocara porocephala ホ シ マ ダ ラ ハ ゼ (upper: HMNH-P54,
81.1 mm SL; lower: FRLM 51507, 23.1 mm SL).

Eleotris fusca (Bloch and Schneider, 1801)
テンジクカワアナゴ [Freshwater] (Fig. 395)
どろ
Jordan and Tanaka (1927); Kuroiwa (1927); Okinawa
Prefecture Experimental Fisheries Station (1934); Ogawa(1937); Hayashi (1995); Yoshigou (2000); Yoshigou et al.
(2001); Konishi (2011)
FRLM 51529, 72.7 mm SL, FRLM 51530, 52.2 mm SL,
Tabaru River, hand net, 24 Sept. 2015; HMNH-P47, 53.0 mm
SL, Tabaru River, hand net, 9 Aug. 1999.

Fig. 395. Eleotris fusca テンジクカワアナゴ (FRLM 51529, 72.7 mm SL).

Ophieleotris sp. 1
タメトモハゼ [Brackish; freshwater]
Yoshigou (2000); Japan Wild Life Research Center (2002);
Ministry of the Environment (2003); Okinawa Prefecture(2005); Ministry of the Environment (2010b); Nakabo (2013).

87

K. Koeda et al.
FAMILY GOBIIDAE
ハゼ科
Acentrogobius janthinopterus (Bleeker, 1853)
カスミハゼ [Brackish] (Fig. 397)
Yoshigou (2000); Yoshigou et al. (2001); Nakabo (2013).
HMNH-P56, 40.6 mm SL, HMNH-P63, 38.7 mm SL,
HMNH-P64, 16.4 mm SL, Tabaru River, hand net, 9 Aug.
1999; HMNH-P95, 23.4 mm SL, Tabaru River, hand net, 10Aug. 1999.

Amblygobius hectori (Smith, 1957)
キンセンハゼ [Marine]
Ito (2009).
Amblygobius nocturnus (Herre, 1945)
ホホベニサラサハゼ [Marine] (Fig. 400)
FRLM 51479, 20.7 mm SL, Hikawa, hand net, 22 Sept.
2015; KAUM–I. 78565, 18.3 mm SL, Kataburu Beach, handnet, 22 Sept. 2015.

Fig. 397. Acentrogobius janthinopterus カスミハゼ (HMNH-P56, 40.6 mm SL).

Amblyeleotris guttata (Fowler, 1938)
ヤマブキハゼ [Marine] (Fig. 398)

Fig. 400. Amblygobius nocturnus ホ ホ ベ ニ サ ラ サ ハ ゼ (upper: FRLM
51479, 20.7 mm SL; lower: KAUM–I. 78565, 18.3 mm SL).

Amblygobius phalaena (Valenciennes, 1837)
サラサハゼ [Marine] (Fig. 401)
Ito (2009).
FRLM 51554, 64.4 mm SL, male, FRLM 51555, 76.5mm SL, male, Kubura Fishing Port, hand net, 24 Sept. 2015;
KAUM–I. 78255, 83.0 mm SL, Kubura Fishing Port, spear,
16 Sept. 2015; KAUM–I. 78498, 88.3 mm SL, Hikawa, handnet, 21 Sept. 2015.

Fig. 398. Amblyeleotris guttata ヤマブキハゼ (Diving Service MARLIN).

Amblyeleotris wheeleri (Polunin and Lubbock, 1977)
クビアカハゼ [Marine] (Fig. 399)

Fig. 399. Amblyeleotris wheeleri クビアカハゼ (Diving Service MARLIN).

88

Fig. 401. Amblygobius phalaena サ ラ サ ハ ゼ (upper: KAUM–I. 78498,
88.3 mm SL; middle: FRLM 51555, 76.5 mm SL; lower: KAUM–I.
78255, 83.0 mm SL).

Checklist of fishes of Yonaguni-jima island
Asterropteryx semipunctata Rüppell, 1830
ホシハゼ [Marine; brackish] (Fig. 402)
KAUM–I. 78399, 29.6 mm SL, Hikawa, hand net, 20
Sept. 2015; KAUM–I. 78515, 21.3 mm SL, Hikawa, hand net,
21 Sept. 2015.
Fig. 405. Bathygobius cyclopterus ヤハズハゼ (KAUM–I. 78439, 18.1 mm SL).

Fig. 402. Asterropteryx semipunctata ホ シ ハ ゼ (KAUM–I. 78399, 29.6mm SL).

Bathygobius fuscus (Rüppell, 1830)
クモハゼ [Marine; brackish] (Fig. 406)
FRLM 51508, 25.7 mm SL, mouth of Kubura River, handnet, 23 Sept. 2015; KAUM–I. 78566, 33.8 mm SL, Kataburu
Beach, hand net, 22 Sept. 2015.

Awaous ocellaris (Broussonet, 1782)
ミナミハゼ [Brackish; freshwater] (Fig. 403)
Yoshigou (2000); Yoshigou et al. (2001); Nakabo (2013).
HMNH-P38, 74.0 mm SL, Tabaru River, hand net, 9 Aug.
1999.
Fig. 406. Bathygobius fuscus クモハゼ (KAUM–I. 78566, 33.8 mm SL).

Fig. 403. Awaous ocellaris ミナミハゼ (HMNH-P38, 74.0 mm SL).

Bathygobius coalitus (Bennett, 1832)
クロヤハズハゼ [Marine; brackish] (Fig. 404)
Yoshigou (2000); Yoshigou et al. (2001).
FRLM 51510, 48.0 mm SL, Tabaru River, hand net, 23
Sept. 2015; FRLM 51516, 48.6 mm SL, mouth of Kubura
River, hand net, 23 Sept. 2015; HMNH-P101, 2 speciemens,
35.5–41.5 mm SL, Kubura River, hand net, 10 Aug. 1999;
HMNH-P250, 60.5 mm SL, Sonai, hand net, 9 Aug. 1999;
KAUM–I. 78393, 13.9 mm SL, KAUM–I. 78394, 12.2 mm
SL, Southern Pasture, hand net, 19 Sept. 2015; KAUM–I.
78400, 43.2 mm SL, Hikawa, hand net, 20 Sept. 2015.

Bathygobius sp.
クモハゼ属の一種 [Brackish]
KAUM–I. 78435, 16.3 mm SL, KAUM–I. 78436, 15.8mm SL, KAUM–I. 78437, 16.1 mm SL, Tabaru River, handnet, 20 Sept. 2015.
Callogobius tanegasimae (Snyder, 1908)
タネハゼ [Marine; brackish] (Fig. 407)
Yoshigou (2000); Yoshigou et al. (2001); Senou et al.
(2004).
HMNH-P67, 22.9 mm SL, Tabaru River, hand net, 9 Aug.
1999.

Fig. 407. Callogobius tanegasimae タネハゼ (HMNH-P67, 22.9 mm SL).

Fig. 404. Bathygobius coalitus ク ロ ヤ ハ ズ ハ ゼ (KAUM–I. 78400, 43.2mm SL).

Bathygobius cyclopterus (Valenciennes, 1837)
ヤハズハゼ [Marine; brackish] (Fig. 405)
Yoshigou et al. (2001).
HMNH-P251, 2 specimens, 21.4–26.6 mm SL, Sonai,
hand net, 9 Aug. 1999; KAUM–I. 78434, 20.4 mm SL, Tabaru
River, hand net, 20 Sept. 2015; KAUM–I. 78439, 18.1 mm
SL, mouth of Tabaru River, hand net, 20 Sept. 2015.

Callogobius sp.
オキナワハゼ属の一種 [Marine] (Fig. 408)
KAUM–I. 78359, 15.0 mm SL, off Kubura Hinan Port,
hand net, 18 Sept. 2015.

Fig. 408. Callogobius sp. (KAUM–I. 78359, 15.0 mm SL).

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K. Koeda et al.
Cryptocentroides insignis (Seale, 1910)
ハスジマハゼ [Marine; brackish] (Fig. 409)
Yoshigou (2000); Yoshigou et al. (2001); Senou et al.
(2004); Nakabo (2013).
HMNH-P100, 35.4 mm SL, male, Kubura River, hand net,
10 Aug. 1999.

Eviota abax (Jordan and Snyder, 1901)
イソハゼ [Marine] (Fig. 413)
KAUM–I. 78563, 16.2 mm SL, KAUM–I. 78564, 13.9mm SL, Kataburu Beach, hand net, 22 Sept. 2015.

Fig. 409. Cryptocentroides insignis ハスジマハゼ (HMNH-P100, 35.4 mm SL).

Cryptocentrus caeruleomaculatus (Herre, 1933)
タカノハハゼ [Marine; brackish] (Fig. 410)
FRLM 51519, 27.3 mm SL, Tabaru River, hand net, 23
Sept. 2015; FRLM 51520, 7.6 mm SL, mouth of Kubura
River, hand net, 23 Sept. 2015.
Fig. 413. Eviota abax イ ソ ハ ゼ (upper: KAUM–I. 78563, 16.2 mm SL;
lower: KAUM–I. 78564, 13.9 mm SL).

Fig. 410. Cryptocentrus caeruleomaculatus タカノハハゼ (FRLM 51519,
27.3 mm SL).

Eviota korechika Shibukawa and Suzuki, 2005
シボリイソハゼ [Marine] (Fig. 414)
KAUM–I. 78529, 13.3 mm SL, Umabana, hand net, 22
Sept. 2015.

Cryptocentrus nigrocellatus (Yanagisawa, 1978)
クロホシハゼ [Marine] (Fig. 411)

Fig. 414. Eviota korechika シボリイソハゼ (KAUM–I. 78529, 13.3 mm SL).

Eviota prasites Jordan and Seale, 1906
アオイソハゼ [Marine] (Fig. 415)
KAUM–I. 78493, 19.5 mm SL, Sonai Channel, hand net,
21 Sept. 2015.
Fig. 411. Cryptocentrus nigrocellatus ク ロ ホ シ ハ ゼ (Diving Service
MARLIN).

Ctenogobiops tangaroai Lubbock and Polunin, 1977
ハタタテシノビハゼ [Marine] (Fig. 412)

Fig. 415. Eviota prasites アオイソハゼ (KAUM–I. 78493, 19.5 mm SL).

Fig. 412. Ctenogobiops tangaroai ハタタテシノビハゼ (Diving Service
MARLIN).

90

Eviota saipanensis Fowler, 1945
ムスジイソハゼ [Marine] (Fig. 416)
KAUM–I. 78390, 18.6 mm SL, KAUM–I. 78391, 16.0mm SL, KAUM–I. 78392, 13.8 mm SL, Southern Pasture,
hand net, 19 Sept. 2015.

Checklist of fishes of Yonaguni-jima island
Favonigobius reichei (Bleeker, 1854)
ミナミヒメハゼ [Brackish] (Fig. 420)
Yoshigou (2000); Yoshigou et al. (2001).
FRLM 51550, 36.8 mm SL, mouth of Kubura River, handnet, 24 Sept. 2015; HMNH-P93, 47.1 mm SL, male, Tabaru
River, hand net, 10 Aug. 1999; KAUM–I. 78433, 38.7 mm
SL, Tabaru River, hand net, 20 Sept. 2015.

Fig. 416. Eviota saipanensis ム ス ジ イ ソ ハ ゼ (upper: KAUM–I. 78391,
16.0 mm SL; lower: KAUM–I. 78390, 18.6 mm SL).

Eviota sp.
イソハゼ属の一種 [Marine] (Fig. 417)
Yoshigou et al. (2001).
HMNH-P249, Agarizaki, hand net, 9 Aug. 1999.
Fig. 420. Favonigobius reichei ミ ナ ミ ヒ メ ハ ゼ (upper: FRLM 51550,
36.8 mm SL; lower: KAUM–I. 78433, 38.7 mm SL).

Fig. 417. Eviota sp. (HMNH-P249, preserved specimen).

Exyrias belissimus (Smith, 1959)
オバケインコハゼ [Marine; brackish] (Fig. 418)
KAUM–I. 78256, 89.8 mm SL, Kubura Fishing Port,
spear, 16 Sept. 2015; KAUM–I. 78516, 53.1 mm SL, Hikawa,
hand net, 21 Sept. 2015.

Favonigobius sp.
ヒメハゼ属の一種 [Brackish] (Fig. 421)
HMNH-P69, 6 specimens, 17.9–19.4 mm SL, Tabaru
River, hand net, 9 Aug. 1999.

Fig. 421. Favonigobius sp. (HMNH-P69, 19.4 mm SL).

Fig. 418. Exyrias belissimus オバケインコハゼ (KAUM–I. 78516, 53.1mm SL).

Exyrias puntang (Bleeker, 1851)
インコハゼ [Marine; brackish] (Fig. 419)
Yoshigou (2000); Yoshigou et al. (2001).
FRLM 51443, 59.9 mm SL, mouth of Kubura River, castnet, 21 Sept. 2015; HMNH-P55, 32.3 mm SL, Tabaru River,
hand net, 9 Aug. 1999.

Fusigobius inframaculatus (Randall, 1994)
ハタタテサンカクハゼ [Marine] (Fig. 422)
KAUM–I. 78472, 37.0 mm SL, Sonai Channel, hand net,
21 Sept. 2015.

Fig. 422. Fusigobius inframaculatus ハタタテサンカクハゼ (KAUM–I.
78472, 37.0 mm SL).

Fig. 419. Exyrias puntang インコハゼ (FRLM 51443, 59.9 mm SL).

91

K. Koeda et al.
Fusigobius neophytus (Günther, 1877)
サンカクハゼ [Marine] (Fig. 423)
Ito (2009).
KAUM–I. 78401, 41.2 mm SL, Hikawa, hand net, 20
Sept. 2015; KAUM–I. 78475, 33.0 mm SL, Sonai Channel,
hand net, 21 Sept. 2015

Istigobius decoratus (Herre, 1927)
ホシカザリハゼ [Marine; brackish] (Fig. 427)

Fig. 427. Istigobius decoratus ホシカザリハゼ (Diving Service MARLIN).

Fig. 423. Fusigobius neophytus サンカクハゼ (KAUM–I. 78475, 33.0 mm SL).

Glossogobius biocellatus (Valenciennes, 1837)
ヒトミハゼ [Brackish] (Fig. 424)
Yoshigou (2000); Yoshigou et al. (2001).
FRLM 50756, 51.0 mm SL, Tabaru River, hand net, 14
Mar. 2015; HMNH-P57, 45.0 mm SL, HMNH-P62, 25.1 mm
SL, Tabaru River, hand net, 9 Aug. 1999.

Istigobius ornatus (Rüppell, 1830)
カザリハゼ [Marine; brackish] (Fig. 428)
Yoshigou et al. (2001).
FRLM 51517, 68.1 mm SL, mouth of Kubura River, handnet, 23 Sept. 2015; HMNH-P99, 40.7 mm SL, Kubura River,
hand net, 10 Aug. 1999; KAUM–I. 78568, 19.3 mm SL,
Kataburu Beach, hand net, 22 Sept. 2015.

Fig. 424. Glossogobius biocellatus ヒトミハゼ (FRLM 50756, 51.0 mm SL).

Glossogobius celebius (Valenciennes, 1837)
イワハゼ [Freshwater] (Fig. 425)
Yoshigou (2000); Yoshigou et al. (2001).
HMNH-P39, 69.8 mm SL, Tabaru River, hand net, 9 Aug.
1999.

Fig. 425. Glossogobius celebius イワハゼ (HMNH-P39, 69.8 mm SL).

Fig. 428. Istigobius ornatus カ ザ リ ハ ゼ (upper: FRLM 51517, 68.1 mm
SL; lower: KAUM–I. 78568, 19.3 mm SL).

Kelloggella cardinalis Jordan and Seale, 1906
アカヒレハダカハゼ [Marine] (Fig. 429)
KAUM–I. 78374, 26.0 mm SL, KAUM–I. 78375, 21.2mm SL, KAUM–I. 78376, 23.4 mm SL, KAUM–I. 78389,
18.9 mm SL, Southern Pasture, hand net, 19 Sept. 2015.

Gnatholepis cauerensis (Bleeker, 1853)
カタボシオオモンハゼ [Marine] (Fig. 426)
FRLM 51477, 37.9 mm SL, Umabana, hand net, 22 Sept.
2015; KAUM–I. 78474, 26.1 mm SL, Sonai Channel, handnet, 21 Sept. 2015.

Fig. 426. Gnatholepis cauerensis カ タ ボ シ オ オ モ ン ハ ゼ (KAUM–I.
78474, 26.1 mm SL).

92

Fig. 429. Kelloggella cardinalis アカヒレハダカハゼ (upper: KAUM–I.
78389, 18.9 mm SL; lower: KAUM–I. 78374, 26.0 mm SL).

Checklist of fishes of Yonaguni-jima island
Lotilia klausewitzi Shibukawa, Suzuki and Senou, 2012
オドリハゼ [Marine] (Fig. 430)

Fig. 433. Oligolepis acutipennis ノボリハゼ (HMNH-P59, 46.2 mm SL).

Oligolepis stomias (Smith, 1941)
クチサケハゼ [Brackish] (Fig. 434)
Yoshigou (2000); Yoshigou et al. (2001).
HMNH-P60, 32.8 mm SL, Tabaru River, hand net, 9 Aug.
1999.
Fig. 430. Lotilia klausewitzi オドリハゼ (Diving Service MARLIN).

Macrodontogobius wilburi Herre, 1936
マダラハゼ [Marine] (Fig. 431)
KAUM–I. 78287, 39.5 mm SL, Kubura Fishing Port, handnet, 16 Sept. 2015.

Fig. 434. Oligolepis stomias クチサケハゼ (HMNH-P60, 32.8 mm SL).

Oplopomus oplopomus (Valenciennes, 1837)
ケショウハゼ [Marine] (Fig. 435)
FRLM 51518, 16.4 mm SL, Hikawa, hand net, 23 Sept.
2015.

Fig. 431. Macrodontogobius wilburi マダラハゼ (KAUM–I. 78287, 39.5mm SL).

Mugilogobius cavifrons (Weber, 1909)
ホホグロハゼ [Brackish] (Fig. 432)
Yoshigou (2000); Yoshigou et al. (2001); Senou et al.
(2004); Ministry of the Environment (2010a); Nakabo (2013).
FRLM 51504, 18.6 mm SL, mouth of Kubura River, hand,
23 Sept. 2015; FRLM 51511, 21.6 mm SL, mouth of Kubura
River, hand net, 23 Sept. 2015; HMNH-P36, 2 specimens,
21.3–30.2 mm SL, Kubura River, hand net, 8 Aug. 1999.

Fig. 435. Oplopomus oplopomus ケショウハゼ .(FRLM 51518, 16.4 mm SL).

Oxyurichthys visayanus Herre, 1927
ミナミサルハゼ [Brackish] (Fig. 436)
Yoshigou (2000); Yoshigou et al. (2001).
HMNH-P58, 49.7 mm SL, Tabaru River, hand net, 9 Aug.
1999.

Fig. 436. Oxyurichthys visayanus ミナミサルハゼ (HMNH-P58, 49.7 mm SL).

Fig. 432. Mugilogobius cavifrons ホ ホ グ ロ ハ ゼ (upper: FRLM 51511,
21.6 mm SL; lower: HMNH-P36, 30.2 mm SL).

Palutrus meteori (Klausewitz and Zander, 1967)
ヒメクモハゼ [Brackish] (Fig. 437)
KAUM–I. 78438, 18.1 mm SL, mouth of Tabaru River,
hand net, 20 Sept. 2015.

Oligolepis acutipennis (Valenciennes, 1837)
ノボリハゼ [Brackish] (Fig. 433)
Yoshigou (2000); Yoshigou et al. (2001).
HMNH-P59, 46.2 mm SL, HMNH-P65, 32.1 mm SL,
HMNH-P66, 17.5 mm SL, Tabaru River, hand net, 9 Aug.
1999.

Fig. 437. Palutrus meteori ヒメクモハゼ (KAUM–I. 78438, 18.1 mm SL).

93

K. Koeda et al.
Pandaka trimaculata Akihito and Meguro, 1975
ミツボシゴマハゼ [Brackish] (Fig. 438)
Yoshigou (2000); Yoshigou et al. (2001); Nakabo (2013).
HMNH-P68, 11.1 mm SL, Tabaru River, hand net, 9 Aug.
1999; KAUM–I. 78445, 11.7 mm SL, KAUM–I. 78446, 12.1mm SL, KAUM–I. 78447, 13.4 mm SL, KAUM–I. 78448,
10.7 mm SL, KAUM–I. 78449, 12.6 mm SL, KAUM–I.
78450, 12.4 mm SL, Tabaru River, hand net, 20 Sept. 2015.

Priolepis cincta (Regan, 1908)
ベンケイハゼ [Marine] (Fig. 440)
KAUM–I. 78533, 22.4 mm SL, Umabana, hand net, 22
Sept. 2015.

Fig. 440. Priolepis cincta ベンケイハゼ (KAUM–I. 78533, 22.4 mm SL).

Priolepis latifascima Winterbottom and Burridge, 1993
フトスジイレズミハゼ [Marine] (Fig. 441)
KAUM–I. 78539, 13.1 mm SL, Umabana, hand net, 22
Sept. 2015.

Fig. 438. Pandaka trimaculata ミ ツ ボ シ ゴ マ ハ ゼ (upper: KAUM–I.
78445, 11.7 mm SL; lower: KAUM–I. 78449, 12.6 mm SL).

Periophthalmus argentilineatus Valenciennes, 1837
ミナミトビハゼ [Marine; brackish] (Fig. 439)
Yoshigou (2000); Yoshigou et al. (2001).
FRLM 51526, 89.7 mm SL, Tabaru River, hand net, 23
Sept. 2015; HMNH-P61, 2 speciemsn, 18.5–32.8 mm SL,
Tabaru River, hand net, 9 Aug. 1999.

Fig. 441. Priolepis latifascima フトスジイレズミハゼ (KAUM–I. 78539,
13.1 mm SL).

Pseudogobius sp.
コクチスナゴハゼ [Brackish] (Fig. 442)
FRLM 51509, 34.3 mm SL, male, Tabaru River, hand net,
23 Sept. 2015.

Fig. 439. Periophthalmus argentilineatus ミナミトビハゼ (FRLM 51526,
89.7 mm SL).

Fig. 442. Pseudogobius sp. コクチスナゴハゼ (FRLM 51509, 34.3 mm SL).

Rhinogobius giurinus (Rutter, 1897)
ゴクラクハゼ [Brackish; freshwater]
Okada and Isa (1959).

Rhinogobius nagoyae Jordan and Seale, 1906
シマヨシノボリ [Brackish; freshwater]
Yoshigou (2000).

94Checklist of fishes of Yonaguni-jima island
Sicyopterus lagocephalus (Pallas, 1770)
ルリボウズハゼ [Freshwater] (Fig. 443)
Yoshigou (2000).
HMNH-P40, 47.2 mm SL, Tabaru River, hand net, 9 Aug.
1999.

Fig. 443. Sicyopterus lagocephalus ル リ ボ ウ ズ ハ ゼ (HMNH-P40, 47.2mm SL).

Sicyopus leprurus (Sakai and Nakamura, 1979)
カエルハゼ [Freshwater] (Fig. 444)
Yoshigou (2000); Yoshigou et al. (2001); Ministry of theEnvironment (2003); Senou et al. (2004); Okinawa Prefecture(2005); Nakabo (2013).
HMNH-P42, 34.6 mm SL, Tabaru River, hand net, 9 Aug.
1999.

Fig. 444. Sicyopus leprurus カエルハゼ (HMNH-P42, 34.6 mm SL).

Stenogobius sp.
タネカワハゼ [Brackish] (Fig. 445)
Yoshigou et al. (2001).
HMNH-P25, 65.1 mm SL, HMNH-P26, 59.3 mm SL,
Tabaru River, hand net, 8 Aug. 1999; HMNH-P43, 88.7 mm
SL, HMNH-P44, 78.3 mm SL, Tabaru River, hand net, 9 Aug.
1999.

Fig. 445. Stenogobius sp. タネカワハゼ (HMNH-P25, 65.1 mm SL).

Stiphodon percnopterygionus Watson and Chen, 1998
ナンヨウボウズハゼ [Freshwater] (Fig. 446)
Yoshigou (2000); Yoshigou et al. (2001).
HMNH-P27, 25.7 mm SL, male, HMNH-P41, 2
specimens, 25.4–26.5 mm SL, male, Tabaru River, hand net,
8 Aug. 1999; KAUM–I. 78408, 33.4 mm SL, male, KAUM–
I. 78409, 25.2 mm SL, male, KAUM–I. 78410, 29.3 mm SL,
female, KAUM–I. 78411, 18.1 mm SL, female, Tabaru River,
hand net, 20 Sept. 2015.

Fig. 446. Stiphodon percnopterygionus ナ ン ヨ ウ ボ ウ ズ ハ ゼ (upper:
KAUM–I. 78408, 33.4 mm SL; middle: KAUM–I. 78409, 25.2 mm SL;
lower: KAUM–I. 78410, 29.3 mm SL).

Trimma macrophthalmus (Tomiyama, 1936)
オオメハゼ [Marine] (Fig. 447)
KAUM–I. 78525, 17.3 mm SL, Umabana, hand net, 22
Sept. 2015.

Fig. 447. Trimma macrophthalmus オ オ メ ハ ゼ (KAUM–I. 78525, 17.3mm SL).

Trimma okinawae (Aoyagi, 1949)
オキナワベニハゼ [Marine] (Fig. 448)
KAUM–I. 78530, 20.0 mm SL, Umabana, hand net, 22
Sept. 2015.

Fig. 448. Trimma okinawae オ キ ナ ワ ベ ニ ハ ゼ (KAUM–I. 78530, 20.0mm SL).

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K. Koeda et al.
Valenciennea sexguttata (Valenciennes, 1837)
ミズタマハゼ [Marine] (Fig. 449)
FRLM 51492, 36.5 mm SL, Sonai Fishing Port, handnet, 22 Sept. 2015; KAUM–I. 78567, 36.8 mm SL, Kataburu
Beach, hand net, 22 Sept. 2015.

Nemateleotris helfrichi Randall and Allen, 1973
シコンハタタテハゼ [Marine] (Fig. 452)
Nakabo (2013).

Fig. 449. Valenciennea sexguttata ミズタマハゼ (KAUM–I. 78567, 36.8mm SL).

Valenciennea strigata (Broussonet, 1782)
アカハチハゼ [Marine] (Fig. 450)
KAUM–I. 78573, 57.6 mm SL, Danno Beach, hand net,
23 Sept. 2015.

Fig. 452. Nemateleotris helfrichi シ コ ン ハ タ タ テ ハ ゼ (Diving Service
MARLIN).

Nemateleotris magnifica Fowler, 1938
ハタタテハゼ [Marine] (Fig. 453)
KAUM–I. 78572, 45.8 mm SL, Danno Beach, spear, 23
Sept. 2015.

Fig. 450. Valenciennea strigata アカハチハゼ (upper: KAUM–I. 78573, 57.6mm SL; lower: Diving Service MARLIN).

FAMILY PTERELEOTRIDAE
クロユリハゼ科
Nemateleotris decora Randall and Allen, 1973
アケボノハゼ [Marine] (Fig. 451)

Fig. 451. Nemateleotris decora アケボノハゼ (Diving Service MARLIN).

96

Fig. 453. Nemateleotris magnifica ハタタテハゼ (KAUM–I. 78572, 45.8mm SL).

Ptereleotris evides (Jordan and Hubbs, 1925)
クロユリハゼ [Marine] (Fig. 454)

Fig. 454. Ptereleotris evides クロユリハゼ (Diving Service MARLIN).

Checklist of fishes of Yonaguni-jima island
Ptereleotris heteroptera (Bleeker, 1855)
オグロクロユリハゼ [Marine] (Fig. 455)

FAMILY EPHIPPIDAE
マンジュウダイ科
Platax boersii Bleeker, 1853
ミカヅキツバメウオ [Marine] (Fig. 457)
Ito (2009).
KAUM–I. 80274, 132.0 mm SL, Sonai Channel, spear, 21
Sept. 2015.

Fig. 455. Ptereleotris heteroptera オグロクロユリハゼ (Diving Service
MARLIN).

Ptereleotris zebra (Fowler, 1938)
ゼブラハゼ [Marine] (Fig. 456)
Nakabo (2013).

Fig. 456. Ptereleotris zebra ゼブラハゼ (Diving Service MARLIN).

Fig. 457. Platax boersii ミ カ ヅ キ ツ バ メ ウ オ (KAUM–I. 80274, 132.0
mm SL).

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K. Koeda et al.
Platax orbicularis (Forsskål, 1775)
ナンヨウツバメウオ [Marine; brackish] (Fig. 458)
KAUM–I. 78403, 98.2 mm SL, Kubura Fishing Port,
gill net, 20 Sept. 2015; MUFS 46877, 25.3 mm SL, Kubura
Fishing Port, hand net, 16 Sept. 2015.

FAMILY SCATOPHAGIDAE
クロホシマンジュウダイ科
Scatophagus argus (Linnaeus, 1766)
クロホシマンジュウダイ [Marine; brackish] (Fig. 459)
Yoshigou (2000).
FRLM 51538, 8.0 mm SL, mouth of Kubura River, handnet, 24 Sept. 2015.

Fig. 459. Scatophagus argus クロホシマンジュウダイ (FRLM 51538, 8.0
mm SL).

FAMILY SIGANIDAE
アイゴ科
Siganus argenteus (Quoy and Gaimard, 1825)
ハナアイゴ [Marine] (Fig. 460)
FRLM 52045, 222.5 mm SL, Kubura-bari, line-fishing, 22
Sept. 2015; KAUM–I. 78440, 195.0 mm SL, Kubura Fishing
Port, line-fishing, 20 Sept. 2015.

Fig. 460. Siganus argenteus ハナアイゴ (KAUM–I. 78440, 195.0 mm SL).

Siganus woodlandi Randall and Kulbicki, 2005
セダカハナアイゴ [Marine]
Ito (2009).

Fig. 458. Platax orbicularis ナンヨウツバメウオ (upper: KAUM–I.
78403, 98.2 mm SL; lower: MUFS 46877, 25.3 mm SL).

98

Siganus sp.
アイゴ属の一種 [Marine]
あいぬいゆ
Ikema (2003).

Checklist of fishes of Yonaguni-jima island
FAMILY ZANCLIDAE
ツノダシ科
Zanclus cornutus (Linnaeus, 1758)
ツノダシ [Marine] (Fig. 461)
FRLM 50720, 90.4 mm SL, Kubura Fishing Port, handnet, 12 Mar. 2015; KAUM–I. 79746, 133.6 mm SL, Rakuda
Rock, spear, 17 Sept. 2015; KAUM–I. 80366, 104.1 mm SL,
Sonai Channel, gill net, 20 Sept. 2015; NSMT-P 101124,
121.0 mm SL, Hikawa, 11 Mar. 2010.

FAMILY ACANTHURIDAE
ニザダイ科
Acanthurus bariene Lesson, 1831
カンランハギ [Marine] (Fig. 462)
MUFS 46926, 185.0 mm SL, levee at Kubura, line-fishing,
18 Sept. 2015.

Fig. 462. Acanthurus bariene カンランハギ (MUFS 46926, 185.0 mm SL).

Acanthurus blochii Valenciennes, 1835
オスジクロハギ [Marine] (Fig. 463)
FRLM 52030, 204.0 mm SL, FRLM 52031, 202.5 mm
SL, Kubura-bari, line-fishing, 22 Sept. 2015; KAUM–I.
78404, 128.8 mm SL, KAUM–I. 78405, 132.0 mm SL,
Kubura Fishing Port, gill net, 20 Sept. 2015; MUFS 46954,
295.0 mm SL, Kubura Fishing Port, gill net, 20 Sept. 2015.

Fig. 463. Acanthurus blochii オ ス ジ ク ロ ハ ギ (upper: KAUM–I. 78404,
128.8 mm SL; lower: FRLM 52030, 204.0 mm SL).

Fig. 461. Zanclus cornutus ツノダシ (upper: KAUM–I. 80366, 104.1 mm
SL; lower: M. Sakurai).

Acanthurus dussumieri Valenciennes, 1835
ニセカンランハギ [Marine]とぅがだ
Ikema (2003).

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K. Koeda et al.
Acanthurus japonicus (Schmidt, 1931)
ナミダクロハギ [Marine] (Fig. 464)
KAUM–I. 80262, 136.5 mm SL, Anchor's graves, spear,
17 Sept. 2015; NSMT-P 101238, 138.5 mm SL, Danno Beach,
19 Mar. 2010.

Acanthurus lineatus (Linnaeus, 1758)
ニジハギ [Marine] (Fig. 466)
FRLM 51585, 126.2 mm SL, mouth of Kubura River, gill
net, 20 Sept. 2015; KAUM–I. 80230, 173.0 mm SL, Rakuda
Rock, spear, 17 Sept. 2015; NSMT-P 55462, 2 specimens,
80.8–150.2 mm SL, Agarizaki, hand net, 28 Mar. 1997.

Fig. 464. Acanthurus japonicus ナミダクロハギ (KAUM–I. 80262, 136.5mm SL).

Acanthurus leucopareius (Jenkins, 1903)
スジクロハギ [Marine] (Fig. 465)
FRLM 52099, 201.8 mm SL, Southern Pasture, linefishing, 21 Sept. 2015; KAUM–I. 80225, 130.8 mm SL,
Arakawa Bay, spear, 18 Sept. 2015.
Fig. 466. Acanthurus lineatus ニジハギ (upper: FRLM 51585, 126.2 mm
SL; lower: Diving Service MARLIN).

Acanthurus maculiceps (Ahl, 1923)
イレズミニザ [Marine] (Fig. 467)
FRLM 51454, 200.8 mm SL, Kubura-bari, line-fishing, 20
Sept. 2015.

Fig. 467. Acanthurus maculiceps イレズミニザ (FRLM 51454, 200.8 mm SL).

Fig. 465. Acanthurus leucopareius ス ジ ク ロ ハ ギ (upper: FRLM 52099,
201.8 mm SL; lower: KAUM–I. 80225, 130.8 mm SL).

100

Acanthurus mata (Cuvier, 1829)
ヒラニザ [Marine] (Fig. 468)とぅがだ
FRLM 51557, 255.3 mm SL, Kubura Fishing Port, linefishing, 24 Sept. 2015; FRLM 51957, 190.1 mm SL, Kuburabari, line-fishing, 20 Sept. 2015; KAUM–I. 79742, 161.7 mm
SL, Rakuda Rock, spear, 17 Sept. 2015.

Checklist of fishes of Yonaguni-jima island
Acanthurus nigrofuscus (Forsskål, 1775)
ナガニザ [Marine] (Fig. 471)
Ito (2009).
FRLM 50700, 122.8 mm SL, Sonai Fishing Port, linefishing, 13 Mar. 2015; FRLM 52056, 114.7 mm SL, Sonai
Fishing Port, line-fishing, 24 Sept. 2015; KAUM–I. 80361,
125.9 mm SL, Kubura Fishing Port, gill net, 20 Sept. 2015;
MUFS 46957, 127.0 mm SL, Kubura Fishing Port, gill net, 20
Sept. 2015; NSMT-P 55463, 12 specimens, 66.6–125.5 mm
SL, Agarizaki, hand net, 28 Mar. 1997.

Fig. 468. Acanthurus mata ヒラニザ (upper: FRLM 51957, 190.1 mm SL;
lower: Diving Service MARLIN).
Fig. 471. Acanthurus nigrofuscus ナガニザ (MUFS 46957, 127.0 mm SL).

Acanthurus nigricans (Linnaeus, 1758)
メガネクロハギ [Marine] (Fig. 469)

Acanthurus olivaceus Bloch and Schneider, 1801
モンツキハギ [Marine] (Fig. 472)
Ito (2009).
KAUM–I. 80273, 174.1 mm SL, Sonai Channel, spear, 21
Sept. 2015.

Fig. 469. Acanthurus nigricans メガネクロハギ (M. Sakurai).

Acanthurus nigricauda Duncker and Mohr, 1929
クロモンツキ [Marine] (Fig. 470)
KAUM–I. 80224, 183.7 mm SL, Arakawa Bay, spear, 18
Sept. 2015; KAUM–I. 80480, 207.6 mm SL, Sonai Channel,
gill net, 20 Sept. 2015; MUFS 46955, 210.0 mm SL, Kubura
Fishing Port, gill net, 20 Sept. 2015.

Fig. 472. Acanthurus olivaceus モ ン ツ キ ハ ギ (KAUM–I. 80273, 174.1
mm SL).

Fig. 470. Acanthurus nigricauda クロモンツキ (KAUM–I. 80480, 207.5mm SL).

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K. Koeda et al.
Acanthurus pyroferus Kittlitz, 1834
クログチニザ [Marine] (Fig. 473)
MUFS 46936, 121.0 mm SL, Sonai Fishing Port, linefishing, 19 Sept. 2015.

Fig. 475. Acanthurus triostegus シマハギ (KAUM–I. 80363, 114.8 mm SL).

Fig. 473. Acanthurus pyroferus クログチニザ (MUFS 46936, 121.0 mm SL).

Acanthurus xanthopterus Valenciennes, 1835
クロハギ [Marine; brackish] (Fig. 476)
FRLM 51586, 118.2 mm SL, mouth of Kubura River, gill
net, 20 Sept. 2015; KAUM–I. 78299, 25.1 mm SL, Kubura
Fishing Port, hand net, 16 Sept. 2015; KAUM–I. 80482, 215.0mm SL, Kubura Fishing Port, gill net, 20 Sept. 2015.

Acanthurus thompsoni (Fowler, 1923)
オハグロハギ [Marine] (Fig. 474)
KAUM–I. 80266, 106.6 mm SL, Anchor's graves, spear,
17 Sept. 2015.

Fig. 474. Acanthurus thompsoni オ ハ グ ロ ハ ギ (KAUM–I. 80266, 106.6mm SL).

Acanthurus triostegus (Linnaeus, 1758)
シマハギ [Marine; brackish] (Fig. 475)
FRLM 51584, 136.0 mm SL, mouth of Kubura River, gill
net, 20 Sept. 2015; KAUM–I. 80363, 114.8 mm SL, Kubura
Fishing Port, gill net, 20 Sept. 2015; MUFS 46956, 137.0 mm
SL, Kubura Fishing Port, gill net, 20 Sept. 2015; NSMT-P
55477, 2 specimens, 19.3–21.1 mm SL, Agarizaki, hand net,
28 Mar. 1997.

Fig. 476. Acanthurus xanthopterus ク ロ ハ ギ (upper: KAUM–I. 80482,
215.0 mm SL; middle: KAUM–I. 78299, 25.1 mm SL; lower: Diving
Service MARLIN).

102

Checklist of fishes of Yonaguni-jima island
Acanthurus sp.
クロハギ属の一種 [Marine; brackish]
Yoshigou (2000).

Ctenochaetus binotatus Randall, 1955
コクテンサザナミハギ [Marine] (Fig. 477)
KAUM–I. 80264, 102.4 mm SL, Anchor's graves, spear,
17 Sept. 2015.

Fig. 477. Ctenochaetus binotatus コ ク テ ン サ ザ ナ ミ ハ ギ (KAUM–I.
80264, 102.4 mm SL).

Ctenochaetus striatus (Quoy and Gaimard, 1825)
サザナミハギ [Marine] (Fig. 478)
FRLM 52057, 106.1 mm SL, Sonai Fishing Port, linefishing, 24 Sept. 2015; KAUM–I. 78286, 33.3 mm SL, Kubura
Fishing Port, hand net, 16 Sept. 2015; MUFS 46889, 125.0mm SL, Rakuda Rock, spear, 17 Sept. 2015.
Fig. 479. Naso hexacanthus テ ン グ ハ ギ モ ド キ (upper: FRLM 50764,
280.0 mm SL; middle: KAUM–I. 80357, 206.0 mm SL; lower: Diving
Service MARLIN).

Fig. 478.

サザナミハギ (KAUM–I. 78286, 33.3 mm SL).

Naso lituratus (Forster, 1801)
ミヤコテングハギ [Marine; brackish] (Fig. 480)
FRLM 51583, 213.5 mm SL, FRLM 51587, 212.0 mm
SL, mouth of Kubura River, gill net, 20 Sept. 2015; KAUM–
I. 80467, 227.7 mm SL, KAUM–I. 80481, 215.0 mm SL,
Kubura Fishing Port, line-fishing, 20 Sept. 2015; MUFS
46953, 163.0 mm SL, Kubura Fishing Port, line-fishing, 20
Sept. 2015.

Naso hexacanthus (Bleeker, 1855)
テングハギモドキ [Marine] (Fig. 479)
FRLM 50764, 280.0 mm SL, FRLM 50765, 257.3 mm SL,
Kubura-bari, line-fishing, 13 Mar. 2015; FRLM 51959, 215.5mm SL, Kubura-bari, line-fishing, 20 Sept. 2015; FRLM
52035, 205.8 mm SL, FRLM 52036, 235.7 mm SL, Kuburabari, line-fishing, 22 Sept. 2015; KAUM–I. 80357, 206.0 mm
SL, Kubura-bari, line-fishing, 20 Sept. 2015.
Fig. 480. Naso lituratus ミヤコテングハギ (KAUM–I. 80467, 227.7 mm SL).

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K. Koeda et al.
Naso unicornis (Forsskål, 1775)
テングハギ [Marine] (Fig. 481)
Fujimoto (1972).
KAUM–I. 80479, 266.6 mm SL, Sonai Channel, gill net,
20 Sept. 2015.

Paracanthurus hepatus (Linnaeus, 1766)
ナンヨウハギ [Marine] (Fig. 483)
FRLM 51495, 187.5 mm SL, Danno-dropoff, spear, 22
Sept. 2015; KAUM–I. 80386, 150.0 mm SL, Umabana, spear,
22 Sept. 2015.

Fig. 481. Naso unicornis テングハギ (KAUM–I. 80479, 266.6 mm SL).

Naso vlamingii (Valenciennes, 1835)
サザナミトサカハギ [Marine] (Fig. 482)
FRLM 50694, 119.1 mm SL, Sonai Fishing Port, linefishing, 13 Mar. 2015; FRLM 51471, 300.4 mm SL, FRLM
51472, 248.2 mm SL, Kubura-bari, line-fishing, 22 Sept.
2015; KAUM–I. 80449, 261.0 mm SL, male, Kubura-bari,line-fishing, 22 Sept. 2015.

Fig. 483. Paracanthurus hepatus ナ ン ヨ ウ ハ ギ (upper: FRLM 51495,
187.5 mm SL; middle: KAUM–I. 80386, 150.0 mm SL; lower: Diving
Service MARLIN).

Prionurus scalprum Valenciennes, 1835
ニザダイ [Marine] (Fig. 484)
KAUM–I. 80209, 364.2 mm SL, Arakawa Bay, spear, 18
Sept. 2015.

Fig. 482. Naso vlamingii サ ザ ナ ミ ト サ カ ハ ギ (upper: FRLM: 51471,
300.4 mm SL; middle: KAUM–I. 80449, 261.0 mm SL; lower: FRLM
50694, 119.1 mm SL).

104

Fig. 484. Prionurus scalprum ニザダイ (KAUM–I. 80209, 364.2 mm SL).

Checklist of fishes of Yonaguni-jima island
Zebrasoma scopas (Cuvier, 1829)
ゴマハギ [Marine] (Fig. 485)
KAUM–I. 78467, 26.0 mm SL, Sonai Channel, hand net,
21 Sept. 2015; KAUM–I. 80265, 121.0 mm SL, Anchor's
graves, spear, 17 Sept. 2015.

Zebrasoma veliferum (Bloch, 1795)
ヒレナガハギ [Marine] (Fig. 486)
KAUM–I. 80268, 173.6 mm SL, Kubura Fishing Port,line-fishing, 19 Sept. 2015.

Fig. 486. Zebrasoma veliferum ヒ レ ナ ガ ハ ギ (KAUM–I. 80268, 173.6mm SL).

FAMILY SPHYRAENIDAE
カマス科
かまさ
Sphyraena barracuda (Edwards, 1771)
オニカマス [Marine; brackish] (Fig. 487)
MUFS 46952, 413.0 mm SL, Kubura Fishing Port, linefishing, 20 Sept. 2015.

Fig. 487. Sphyraena barracuda オニカマス (MUFS 46952, 413.0 mm SL).
Fig. 485. Zebrasoma scopas ゴマハギ (upper: KAUM–I. 80265, 121.0 mm
SL; lower: KAUM–I. 78467, 26.0 mm SL).

Sphyraena helleri Jenkins, 1901
ホソカマス [Marine] (Fig. 488)

Fig. 488. Sphyraena helleri ホソカマス (Diving Service MARLIN).

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K. Koeda et al.
Sphyraena putnamae Jordan and Seale, 1905
オオカマス [Marine] (Fig. 489)

Gymnosarda unicolor (Rüppell, 1836)
イソマグロ [Marine] (Fig. 492)とぅがぎんIto (2009).
KAUM–I. 80425, 525.0 mm SL, Danno-dropoff, linefishing, 22 Sept. 2015.

Fig. 489. Sphyraena putnamae オオカマス (Diving Service MARLIN).

Sphyraena sp.
カマス属の一種 [Marine] (Fig. 490)

Fig. 490. Sphyraena sp. (Diving Service MARLIN).

FAMILY GEMPYLIDAE
クロタチカマス科
Thyrsitoides marleyi Fowler, 1929
ナガタチカマス [Marine] (Fig. 491)

Fig. 492. Gymnosarda unicolor イ ソ マ グ ロ (upper: KAUM–I. 80425,
525.0 mm SL; middle: Underwater Ruin, Diving Service MARLIN;
lower: fish landing port at Yonaguni-jima island, K. Koeda).

Katsuwonus pelamis (Linnaeus, 1758)
カツオ [Marine] (Fig. 493)
KAUM–I. 79749, 349.0 mm SL, south of Yonaguni-jimaisland, line-fishing, 18 Sept. 2015; NSMT-P 100671, 390.3mm SL, south of Yonaguni-jima island, line-fishing, 17 Mar.
2010.

Fig. 491. Thyrsitoides marleyi ナガタチカマス (D. Sasaki).

FAMILY SCOMBRIDAE
サバ科
Acanthocybium solandri (Cuvier, 1832)
カマスサワラ [Marine]
さーら
Fujimoto (1972); Ikema (2003).

106

Fig. 493. Katsuwonus pelamis カツオ (KAUM–I. 79749, 349.0 mm SL).

Scomberomorus commerson (Lacepède, 1800)
ヨコシマサワラ [Marine]
Ito (2009).

Checklist of fishes of Yonaguni-jima island
Thunnus albacares (Bonnaterre, 1788)
キハダ [Marine] (Fig. 494)
しび
Ikema (2003).
KAUM–I. 79711, 320.4 mm SL, KAUM–I. 79715, 329.3
mm SL, south of Yonaguni-jima island, line-fishing, 18 Sept.
2015; MUFS 46914, 324.0 mm SL, south of Yonaguni-jimaisland, line-fishing, 18 Sept. 2015.

Fig. 494. Thunnus albacares キハダ (upper: KAUM–I. 79711, 320.4 mm
SL; lower: KAUM–I. 79715, 329.3 mm SL).

Thunnus obesus (Lowe, 1839)
メバチ [Marine]
しび
Ikema (2003).
Thunnus sp.
マグロ属の一種 [Marine]
だいしび
Ikema (2003).

Makaira nigricans Lacepède, 1802
クロカジキ [Marine] (Fig. 496)
あてぃぬいゆShimose et al. (2006, 2009).

Fig. 496. Makaira nigricans クロカジキ (Diving Service MARLIN).

FAMILY NOMEIDAE
エボシダイ科
Psenes arafurensis Günther, 1889
クラゲウオ [Marine] (Fig. 497)

FAMILY ISTIOPHORIDAE
マカジキ科
Istiompax indica (Cuvier, 1832)
シロカジキ [Marine]
Shimose et al. (2008).
Istiophorus platypterus (Shaw, 1792)
バショウカジキ [Marine] (Fig. 495)
はにあてぃ

Fig. 497. Psenes arafurensis クラゲウオ (Diving Service MARLIN).

Psenes cyanophrys Valenciennes, 1833
スジハナビラウオ [Marine]
URM-P 30224, May 1993.

Psenes pellucidus Lütken, 1880
ハナビラウオ [Marine]
Ito (2009).

Fig. 495. Istiophorus platypterus バ シ ョ ウ カ ジ キ (Diving Service
MARLIN).

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K. Koeda et al.
FAMILY ARIOMMATIDAE
オオメメダイ科
Ariomma luridum Jordan and Snyder, 1904
オオメメダイ [Marine]
NSMT-P 76148, 214.6 mm SL, southeast of Yonagunijima island, 1 Aug. 2006.
FAMILY CAPROIDAE
ヒシダイ科
Antigonia capros Lowe, 1843
ヒシダイ [Marine] (Fig. 498)
FRLM 50750, 141.2 mm SL, west of Yonaguni-jimaisland, line-fishing, 16 Mar. 2015.

ORDER PLEURONECTIFORMES
カレイ目
FAMILY BOTHIDAE
ダルマガレイ科
Bothus pantherinus (Rüppell, 1830)
トゲダルマガレイ [Marine; Brackish] (Fig. 499)
Yoshigou et al. (2001).
HMNH-P73, 31.7 mm SL, Tabaru River, hand net, 9 Aug.
1999; FRLM 51531, 30.4 mm SL, Sonai Fishing Port, fishinglight, 24 Sept. 2015.

Fig. 498. Antigonia capros ヒシダイ (FRLM 50750, 141.2 mm SL).

Fig. 499. Bothus pantherinus ト ゲ ダ ル マ ガ レ イ (upper: FRLM 51531,
30.4 mm SL; middle: HMNH-P73, 31.7 mm SL, preserved specimen;
lower: Diving Service MARLIN).

FAMILY SOLEIDAE
ササウシノシタ科
Aseraggodes cheni Randall and Senou, 2007
ムシクイササウシノシタ [Marine] (Fig. 500)
KAUM–I. 80402, 57.9 mm SL, Na-ma Beach, cast net, 22
Sept. 2015.

108

Checklist of fishes of Yonaguni-jima island
ORDER TETRAODONTIFORMES
フグ目
FAMILY BALISTIDAE
モンガラカワハギ科
くるびBalistapus undulatus (Park, 1797)
クマドリ [Marine] (Fig. 502)
FRLM 50761, 134.0 mm SL, Kubura-bari, line-fishing,
13 Mar. 2015; FRLM 51972, 200.1 mm SL, off Hikawa, linefishing, 21 Sept. 2015; FRLM 52043, 109.0 mm SL, Kuburabari, line-fishing, 22 Sept. 2015; FRLM 52092, 171.0 mm SL,
off Hikawa, line-fishing, 21 Sept. 2015; KAUM–I. 80208,
202.5 mm SL, Yonaguni-jima island, line-fishing, 18 Sept.
2015; KAUM–I. 80394, 139.6 mm SL, female, off Hikawa,line-fishing, 21 Sept. 2015; MUFS 46908, 190.0 mm SL,
MUFS 46909, 125.0 mm SL, off Kubura Hinan Port, linefishing, 18 Sept. 2015.

Fig. 500. Aseraggodes cheni ムシクイササウシノシタ (KAUM–I. 80402,
57.9 mm SL).

Aseraggodes xenicus (Matsubara and Ochiai, 1963)
オトメウシノシタ [Brackish] (Fig. 501)
Yoshigou (2000); Yoshigou et al. (2001).
HMNH-P72, 59.0 mm SL, Tabaru River, hand net, 9 Aug.
1999.
Fig. 502. Balistapus undulatus クマドリ (KAUM–I. 80208, 202.5 mm SL).

Balistoides conspicillum (Bloch and Schneider, 1801)
モンガラカワハギ [Marine] (Fig. 503)
Fujimoto (1972).
FRLM 50759, 204.5 mm SL, Kubura-bari, line-fishing, 13
Mar. 2015; KAUM–I. 80220, 217.0 mm SL, Arakawa Bay,
spear, 18 Sept. 2015.

Fig. 501. Aseraggodes xenicus オトメウシノシタ (HMNH-P72, 59.0 mm SL).

Fig. 503. Balistoides conspicillum モンガラカワハギ (upper: KAUM–I.
80220, 217.0 mm SL; lower: Diving Service MARLIN).

109

K. Koeda et al.
Balistoides viridescens (Bloch and Schneider, 1801)
ゴマモンガラ [Marine] (Fig. 504)
Ito (2009).
FRLM 50754, 251.6 mm SL, Sonai Fishing Port, linefishing, 14 Mar. 2015; KAUM–I. 78501, 41.6 mm SL,
Hikawa, hand net, 21 Sept. 2015; KAUM–I. 80396, 303.5 mm
SL, off Hikawa, line-fishing, 21 Sept. 2015.

Canthidermis maculata (Bloch, 1786)
アミモンガラ [Marine]
NSMT-P 76151, 254.2 mm SL, southeast of Yonagunijima island, spear, 1 Aug. 2006.
Melichthys vidua (Richardson, 1845)
クロモンガラ [Marine] (Fig. 505)
FRLM 52089, 121.5 mm SL, off Hikawa, line-fishing,
21 Sept. 2015; KAUM–I. 79743, 161.3 mm SL, Rakuda
Rock, spear, 17 Sept. 2015; KAUM–I. 80258, 174.9 mm SL,
Anchor's graves, spear, 17 Sept. 2015; MUFS 46910, 145.0mm SL, off Kubura Hinan Port, line-fishing, 18 Sept. 2015.

Fig. 505. Melichthys vidua クロモンガラ (KAUM–I. 79743, 161.3 mm SL).

Odonus niger (Rüppell, 1836)
アカモンガラ [Marine] (Fig. 506)
Ito (2009).
FRLM 52090, 160.8 mm SL, FRLM 52095, 118.6 mm
SL, off Hikawa, line-fishing, 21 Sept. 2015; KAUM–I. 79734,
161.2 mm SL, KAUM–I. 79738, 162.2 mm SL, off Kubura
Hinan Port, spear, 18 Sept. 2015; MUFS 46912, 117.0 mm
SL, off Kubura Hinan Port, line-fishing, 18 Sept. 2015; MUFS
46993, 120.0 mm SL, off Hikawa, line-fishing, 21 Sept. 2015.

Fig. 504. Balistoides viridescens ゴ マ モ ン ガ ラ (upper to lower: FRLM
50754, 251.6 mm SL; KAUM–I. 80396, 303.5 mm SL; KAUM–I.
78501, 41.6 mm SL; Diving Service MARLIN).

110

Fig. 506. Odonus niger ア カ モ ン ガ ラ (upper: FRLM 52095, 160.8 mm
SL; lower: KAUM–I. 79738, 162.2 mm SL).

Checklist of fishes of Yonaguni-jima island
Pseudobalistes flavimarginatus (Rüppell, 1829)
キヘリモンガラ [Marine] (Fig. 507)
KAUM–I. 78502, 32.6 mm SL, Hikawa, hand net, 21
Sept. 2015.

Fig. 507. Pseudobalistes flavimarginatus キ ヘ リ モ ン ガ ラ (KAUM–I.
78502, 32.6 mm SL).

Pseudobalistes fuscus (Bloch and Schneider, 1801)
イソモンガラ [Marine] (Fig. 508)

Fig. 508. Pseudobalistes fuscus イ ソ モ ン ガ ラ (Underwater Luin, K.
Koeda).

Rhinecanthus aculeatus (Linnaeus, 1758)
ムラサメモンガラ [Marine] (Fig. 509)
FRLM 50701, 90.8 mm SL, Sonai Fishing Port, linefishing, 13 Mar. 2015; FRLM 52046, 119.3 mm SL, Sonai
Fishing Port, line-fishing, 24 Sept. 2015; KAUM–I. 80284,
110.0 mm SL, Sonai Fishing Port, line-fishing, 20 Sept. 2015;
KAUM–I. 80351, 115.0 mm SL, Sonai Channel, spear, 21
Sept. 2015.

Fig. 509. Rhinecanthus aculeatus ム ラ サ メ モ ン ガ ラ (upper: FRLM
50701, 90.8 mm SL; middle: KAUM–I. 80284, 110.0 mm SL; lower:
Diving Service MARLIN).

Rhinecanthus rectangulus (Bloch and Schneider, 1801)
タスキモンガラ [Marine] (Fig. 510)
FRLM 50760, 136.6 mm SL, Kubura-bari, line-fishing,
13 Mar. 2015; FRLM 51451, 91.2 mm SL, Kubura-bari,line-fishing, 20 Sept. 2015; KAUM–I. 78649, 137.5 mm SL,
Arakawa Bay, spear, 18 Sept. 2015; NSMT-P 101237, 135.0
mm SL, Danno Beach, 19 Mar. 2010.

Fig. 510. Rhinecanthus rectangulus タ ス キ モ ン ガ ラ (KAUM–I. 78649,
137.5 mm SL).

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K. Koeda et al.
Rhinecanthus verrucosus (Linnaeus, 1758)
クラカケモンガラ [Marine] (Fig. 511)
KAUM–I. 78500, 47.8 mm SL, Hikawa, hand net, 21
Sept. 2015.

Fig. 513. Sufflamen chrysopterum ツマジロモンガラ (KAUM–I. 78308,
113.0 mm SL).

Fig. 511. Rhinecanthus verrucosus クラカケモンガラ (KAUM–I. 78500,
47.8 mm SL).

Sufflamen fraenatum (Latreille, 1804)
メガネハギ [Marine] (Fig. 514)
FRLM 52091, 150.3 mm SL, male, off Hikawa, linefishing, 21 Sept. 2015.

Sufflamen bursa (Bloch and Schneider, 1801)
ムスメハギ [Marine] (Fig. 512)
FRLM 52008, 134.8 mm SL, off Hikawa, line-fishing,
21 Sept. 2015; KAUM–I. 79739, 100.3 mm SL, off Kubura
Hinan Port, spear, 18 Sept. 2015; KAUM–I. 80390, 110.8 mm
SL, Umabana, spear, 22 Sept. 2015.

Fig. 514. Sufflamen fraenatum メガネハギ (FRLM 52091, 150.3 mm SL).

Xanthichthys auromarginatus (Bennett, 1832)
ホシモンガラ [Marine] (Fig. 515)
FRLM 51481, 120.3 mm SL, male, off Northern Pasture,line-fishing, 22 Sept. 2015.

Fig. 512. Sufflamen bursa ムスメハギ (KAUM–I. 79739, 100.3 mm SL).

Sufflamen chrysopterum (Bloch and Schneider, 1801)
ツマジロモンガラ [Marine] (Fig. 513)
FRLM 50729, 104.0 mm SL, Kubura Fishing Port, linefishing, 12 Mar. 2015; FRLM 52084, 102.8 mm SL, FRLM
52088, 110.8 mm SL, off Hikawa, line-fishing, 21 Sept. 2015;
KAUM–I. 78308, 113.0 mm SL, male, Kataburu Beach,line-fishing, 17 Sept. 2015; MUFS 46911, 139.0 mm SL, off
Kubura Hinan Port, line-fishing, 18 Sept. 2015; MUFS 46925,
129.0 mm SL, levee at Kubura, line-fishing, 18 Sept. 2015;
MUFS 46994, 109.0 mm SL, off Hikawa, line-fishing, 21
Sept. 2015.

112

Fig. 515. Xanthichthys auromarginatus ホ シ モ ン ガ ラ (FRLM 51481,
120.3 mm SL).

Checklist of fishes of Yonaguni-jima island
Xanthichthys lineopunctatus (Hollard, 1854)
スジナメモンガラ [Marine] (Fig. 516)
FRLM 51974, 141.6 mm SL, FRLM 52007, 146.9 mm
SL, off Hikawa, line-fishing, 21 Sept. 2015.

FAMILY OSTRACIIDAE
ハコフグ科
Ostracion cubicus Linnaeus, 1758
ミナミハコフグ [Marine] (Fig. 518)
KAUM–I. 80214, 178.0 mm SL, Kubura Fishing Port,line-fishing, 18 Sept. 2015.

Fig. 516. Xanthichthys lineopunctatus スジナメモンガラ (FRLM 51974,
141.6 mm SL).

FAMILY MONACANTHIDAE
カワハギ科
くるび
Paraluteres prionurus (Bleeker, 1851)
ノコギリハギ [Marine] (Fig. 517)
KAUM–I. 78266, 17.9 mm SL, Kubura Fishing Port, handnet, 16 Sept. 2015.

Fig. 518. Ostracion cubicus ミ ナ ミ ハ コ フ グ (upper: KAUM–I. 80214,
178.0 mm SL; lower: Diving Service MARLIN).

Ostracion meleagris Shaw,1796
クロハコフグ [Marine] (Fig. 519)

Fig. 519. Ostracion meleagris クロハコフグ (Diving Service MARLIN).

FAMILY TRIODONTIDAE
ウチワフグ科
Triodon macropterus Lesson, 1829
ウチワフグ [Marine]
NSMT-P 78544, 296.8 mm SL, southeast of Yonagunijima island, line-fishing, 21 July 2007.

Fig. 517. Paraluteres prionurus ノコギリハギ (upper: KAUM–I. 78266,17.9 mm SL; lower: Diving Service MARLIN).

113

K. Koeda et al.
FAMILY TETRAODONTIDAE
フグ科
Arothron caeruleopunctatus Matsuura, 1994
アラレフグ [Marine]
Nakabo (2013).
Arothron hispidus (Linnaeus, 1758)
サザナミフグ [Marine; brackish] (Fig. 520)
KAUM–I. 78298, 62.2 mm SL, Kubura Fishing Port, handnet, 16 Sept. 2015.

Canthigaster amboinensis (Bleeker, 1864)
ゴマフキンチャクフグ [Marine]
URM-P 34361, Danno Beach, 2 Aug. 1995.
Canthigaster compressa (Marion de Procé, 1822)
ヒメキンチャクフグ [Marine]
Ito (2009).
Canthigaster janthinoptera (Bleeker, 1855)
シボリキンチャクフグ [Marine] (Fig. 523)
FRLM 51475, 46.3 mm SL, Umabana, hand net, 22 Sept.
2015; HMNH-P139, 51.2 mm SL, Sonai, hand net, 9 Aug.
1999; KAUM–I. 78553, 45.5 mm SL, Umabana, hand net, 22
Sept. 2015.

Fig. 523. Canthigaster janthinoptera シボリキンチャクフグ (KAUM–I.
78553, 45.5 mm SL).
Fig. 520. Arothron hispidus サザナミフグ (upper: KAUM–I. 78298, 62.2mm SL; lower: Diving Service MARLIN).

Arothron nigropunctatus (Bloch and Schneider, 1801)
コクテンフグ [Marine] (Fig. 521)
Yoshigou et al. (2001).
HMNH-P121, 40.1 mm SL, Agarizaki, hand net, 9 Aug.
1999; KAUM–I. 80519, 175.4 mm SL, Arakawa Bay, spear,
18 Sept. 2015.

Canthigaster valentini (Bleeker, 1853)
シマキンチャクフグ [Marine] (Fig. 524)
FRLM 50703, 65.4 mm SL, Sonai Fishing Port, linefishing, 13 Mar. 2015; FRLM 51430, 78.4 mm SL, FRLM
51991, 93.1 mm SL, Sonai Fishing Port, line-fishing, 19 Sept.
2015.

Fig. 521. Arothron nigropunctatus コクテンフグ (KAUM–I. 80519, 175.4 mm).

Arothron stellatus (Anonymous, 1798)
モヨウフグ [Marine] (Fig. 522)

Fig. 524. Canthigaster valentini シ マ キ ン チ ャ ク フ グ (upper: FRLM
50703, 65.4 mm SL; lower: Diving Service MARLIN).
Fig. 522. Arothron stellatus モヨウフグ (Diving Service MARLIN).

114

Checklist of fishes of Yonaguni-jima island
Sphoeroides pachygaster (Müller and Troschel, 1848)
ヨリトフグ [Marine]
NSMT-P 71452, 230.1 mm SL, southeast of Yonagunijima island, 13 July 2005.

FAMILY DIODONTIDAE
ハリセンボン科とぅとぅ
Diodon hystrix Linnaeus, 1758
ネズミフグ [Marine] (Fig. 525)
FRLM 51967, 308.4 mm SL, Sonai Fishing Port, hand net,
23 Sept. 2015.

Fig. 525. Diodon hystrix ネズミフグ (FRLM 51967, 308.4 mm SL).

Diodon liturosus Shaw, 1804
ヒトヅラハリセンボン [Marine; brackish] (Fig. 526)
FRLM 51977, 203.1 mm SL, mouth of Tabaru River, handnet, 21 Sept. 2015.

Fig. 526. Diodon liturosus ヒトヅラハリセンボン (upper: FRLM 51977,
203.1 mm SL; lower: Diving Service MARLIN).

115

K. Koeda et al.
Discussion
In this study, 603 fish species (306 genera and 104
families) were confirmed to occur around Yonaguni-jimaisland based on the examination of 1,314 specimens, 146
photographs, and previous literature. 334 species and 78
species are representing the first records from the island
based on collected specimens and photographs, respectively.
1,120 specimens of 399 species (211 genera, 72 families) ofmarine, brackish and freshwater fishes were collected during
our expedition at Yonaguni-jima island, with 299 species
representing the first records from the island. Two species,
Cypo zaps (Pseudochromidae) and Opistognathus variabilis(Opistognathidae), were collected for the first time fromJapanese waters in our expedition. These have been separately
reported with morphological descriptions and taxonomic
information (Hibino et al., 2016; Yoshida et al., in press).
The most speciose families of fishes in marine, brackish,and freshwater of Yonaguni-jima island are listed in Table
2. Consequently, a total of 94 species of 33 families and 28
species of 12 families were confirmed from brackish andfresh waters of Yonaguni-jima island in the present study,
respectively (Table 2). It is dramatically fewer than theadjacent islands [290 in Ishigaki-jima and 493 in Iriomotejima islands (Yoshigou, 2014)] because of fewer and smaller
rivers on Yonguni-jima island. As stated above, Yoshigou(2000) recorded 63 fresh and brackish waters species (26families) from Yonaguni-jima island in three days survey
including 50 species (22 families) from Tabaru River.
However, despite focused effort collecting fresh and brackish
water specimens survey in our expedition, only 27 speciesof 12 families were collected from the Tabaru River. In
1999, the tideland of Tabaru River in the mangrove area
had a highly speciose environment with some numerous
small gobies, e.g., Acentrogobius janthinopterus (Bleeker,1853) and Callogobius tanegasimae (Snyder, 1908) (Mr. H.

Yoshigou, pers. comm.). In contrast, this area was a very
poor environment in the present survey in September 2015
, and there were few macro-benthos or gobies [Eleotris
acanthopoma Bleeker, 1853, Pandaka trimaculata Akihito
and Meguro, 1975, Periophthalmus argentilineatus (Bleeker,1854), and Pseudogobius sp.]. The sluice gate of Tabaru Riverand its mouth area were repaired and modified after the survey
in 1999 by Yoshigou (2000). From personal communications
with Mr. H. Yoshigou, it is inferred that the salinity level ofTabaru River became extremely much lower than in 1999
due to the repair work. These results indicate environmental
deterioration has been occurred in the Tabaru River.
The presence or absence of inshore fishes represents
biogeographical features more clearly than the presenceor absence of deepwater fishes, which usually have abroader distribution (sometimes even circumglobal). In
total, 565 marine fish species of 95 families are recorded
from Yonaguni-jima island. The top 15 marine fish families
comprise 64% of the total ichthyofauna; in those, Labridae(47 species), Pomacentridae (39 species), and Gobiidae(37 species) are the dominant families, with a total of 123
species representing more than 20% of total ichthyofauna.
In the Ryukyu Archipelago, Senou et al. (2006; 2007),
Watai et al. (2009), and Motomura et al. (2010) reported
ichthyofaunal checklists which mainly focused on marine
fishes of the Miyako Group (Fig. 2D), Ie-jima island (Fig.
2F), Tokashiki-jima island (Fig. 2E), and Yaku-shima island(Fig. 2H), respectively, and ranked the most speciose families.
Additionally, field guides of Iou-jima and Take-shima islands(Fig. 2I), and Yoron-jima island (Fig. 2G) were publishedby Motomura et al. (2013) and Motomura and Matsuura(2014), respectively, on the basis of collected specimens andunderwater photographs. Although, some reports include
freshwater and brackish water fish in their rank, their results
are herein compared at Table 3. Our expedition was carried

Table 2. Most speciose 15 families of fishes in marine, brackish, and freshwater of Yonaguni-jima island in the present study.

Rank
1
2
3
4
5
6
7
8
9
10
11
12
13
14
15

All
FamilyNS
Gobiidae57
Labridae46
Pomacentridae 39
Chaetodontidae 29
Serranidae
28
Acanthuridae 26
Lutjanidae
20
Apogonidae 20
Scorpaenidae 19
Holocentridae 17
Blenniidae
16
Balistidae
16
Muraenidae
16
Carangidae
15
Scaridae
14
Subtotal
378
Total
603

%TIF
9.5
7.6
6.5
4.8
4.6
4.3
3.3
3.3
3.2
2.8
2.7
2.7
2.7
2.5
2.3
62.7

MarineFamilyNS %TIF
Labridae46 8.1
Pomacentridae 39 6.9Gobiidae
38 6.7
Chaetodontidae 29 5.1
Serranidae
28 5.0Acanthuridae 26 4.6Lutjanidae
20 3.5
Apogonidae 19 3.4
Scorpaenidae 19 3.4
Holocentridae 17 3.0
Blenniidae
16 2.8Balistidae
16 2.8
Muraenidae
16 2.8
Carangidae
15 2.7
Scaridae
14 2.5
Subtotal
358 63.4
Total
565

NS, number of species; %TIF, % of total ichthyofauna.

116

Brackish
FamilyNS %TIF
Gobiidae28 29.8
Mugilidae
7
7.4
Carangidae
6
6.4
Ambassidae
44.3
Lutjanidae
4
4.3
Eleotridae
4
4.3
Acanthuridae
44.3
Gerreidae
44.3
Chaetodontidae 3
3.2
Anguillidae
22.1
Muraenidae
2
2.1
Aulopidae
2
2.1
Apogonidae
2
2.1
Kuhliidae
2
2.1
Pomacentridae 2
2.1
Subtotal
76 81.1
Total
94

Freshwater
FamilyNS
Gobiidae7
Eleotridae
5
Ambassidae
4
Anguillidae
2
Cyprinidae
2
Kuhliidae
2
Elopidae1
Cobitidae
1
Mugilidae
1
Poecliidae
1
Syngnathidae
1Cichlidae
1

Subtotal
Total

28
28

%TIF
25.0
17.9
14.3
7.1
7.1
7.1
3.6
3.6
3.6
3.6
3.6
3.6

Checklist of fishes of Yonaguni-jima island
Table 3. Most speciose 15 families of fishes of the islands in the Ryukyu Archipelago in the previous studies.

Rank
1
2
3
4
5
6
7
8
9
10
11
12
13
14
15

Miyako Group*

Tokashiki-jima island*Ie-jima island*

Senou et al. (2007)
FamilyNS%TIF
Gobiidae146
16.9
Labridae97
11.2
Pomacentridae
83
9.6
Apogonidae
45
5.2
Serranidae
34
3.9
Chaetodontidae
33
3.8
Blenniidae
32
3.7
Scaridae26
3.0
Acanthuridae
25
2.9
Syngnathidae
19
2.2
Pomacanthidae
16
1.9
Holocentridae
15
1.7
Muraenidae
14
1.6
Ptereleotridae
14
1.6
Scorpaenidae
14
1.6
Subtotal
613
71.0
Total
863
100.0

Watai et al. (2009)
FamilyNS%TIF
Pomacentridae
52
12.7
Labridae43
10.5
Gobiidae39
9.5
Blenniidae
31
7.5
Chaetodontidae
22
5.4
Apogonidae
20
4.9
Scorpaenidae
122.9
Muraenidae
11
2.7
Acanthuridae
11
2.7
Serranidae
10
2.4
Holocentridae
92.2
Pomacanthidae
81.9Tripterygiidae
81.9Syngnathidae
7
1.7
Balistidae
7
1.7
Subtotal
290
70.6
Total
411
100.0

Senou et al. (2006)
FamilyNS%TIF
Gobiidae112
12.6
Labridae97
10.9
Pomacentridae
78
8.8
Apogonidae
48
5.4
Serranidae
45
5.1
Blenniidae
37
4.2
Chaetodontidae
32
3.6
Acanthuridae
27
3.0
Scaridae25
2.8
Syngnathidae
212.4
Muraenidae
19
2.1
Pomacanthidae
18
2.0
Scorpaenidae
18
2.0
Holocentridae
17
1.9
Carangidae
16
1.8
Subtotal
610
68.6
Total
889
100.0

NS, number of species; %TFF, % of total ichthyofauna. *only marine fish species are listed
Table 3. continue.

Rank
1
2
3
4
5
6
7
8
9
10
11
12
13
14
15

Yoron-jima island
Motomura and Matsuura (2014)
FamilyNS%TIF
Gobiidae69
9.9
Labridae56
8.0
Pomacentridae
56
8.0
Apogonidae
36
5.2
Blenniidae
29
4.2
Serranidae
28
4.0
Scorpaenidae
21
3.0
Chaetodontidae
21
3.0
Acanthuridae
21
3.0
Tripterygiidae
18
2.6
Carangidae
15
2.2
Balistidae
15
2.2
Muraenidae
14
2.0
Holocentridae
14
2.0
Scaridae14
2.0
Subtotal
427
61.3
Total
697
100.0

Iou-jima and Take-shima islands
Motomura et al. (2013)
FamilyNS%TIF
Labridae42
10.1
Gobiidae34
8.2
Pomacentridae
29
7.0
Serranidae
27
6.5
Scorpaenidae
26
6.3
Apogonidae
26
6.3
Chaetodontidae
18
4.3
Blenniidae
16
3.9
Scaridae13
3.1
Carangidae
12
2.9
Holocentridae
10
2.4
Acanthuridae
10
2.4
Muraenidae
7
1.7
Lutjanidae
7
1.7
Balistidae
7
1.7
Subtotal
284
68.6
Total
414
100.0

Yaku-shima island**Motomura et al. (2010)
FamilyNS%TIF
Gobiidae110
11.6
Labridae98
10.3
Pomacentridae
66
6.9
Apogonidae
45
4.7
Serranidae
43
4.5
Blenniidae
38
4.0
Chaetodontidae
34
3.6
Acanthuridae
28
2.9
Holocentridae
24
2.5
Scaridae24
2.5
Muraenidae
23
2.4
Lutjanidae
21
2.2
Scorpaenidae
20
2.1
Tripterygiidae
181.9
Syngnathidae
161.7
Subtotal
608
63.9
Total
951
100.0

NS, number of species; %TIF, % of total ichthyofauna. **only marine and brackish fish species are listed

out over only 20 days (10 days for main survey at September2015), and was obviously shorter than the other previous
research (Motomura et al, 2010, 2013; Motomura andMatsuura, 2014). However, the number of collected species
has no large difference among those reports. The list of marine
fishes of Yonaguni-jima island does show some differencesfrom other reports. Gobiidae which was ranked as 1st in
most of other islands, are ranked as 3rd, and Labridae andPomacentridae are 1st and 2nd, respectively. In addition, thepercentages of these three families are generally lower than atother islands. Chaetodontidae, Serranidae, and Acanthuridae
are ranked 4th to 6th with small differences in Yonagunijima island, but those were lower (Chaetodontidae: 5th to 7th;
Serranidae: 4th to 10th; Acanthuridae: 7th to 11th) in otherislands. Lutjanidae ranked 7th in Yonaguni-jima island, higher(12th to 18th) than the other islands. Balistidae (11th) is also
a higher rank than at the other islands. On the other hand, theranks of Apogonidae (8th) and Blenniidae (11th) in Yonagunijima island are obviously lower than those of the other islands(Apogonidae: 4th to 6th; Blenniidae: 4th to 8th). The present
results from Yonaguni-jima island can be characterized byhigh species diversity in middle-sized swimming taxa (e.g.
Chaetodontidae, Acanthuridae, Lutjanidae, and Balistidae),and low species diversity in small-sized benthic taxa (e.g.
Gobiidae, Apogonidae and Blenniidae). Based on our resultsof the unusual species composition, it is speculated that theichthyofauna of Yonaguni-jima island is influenced by the
combination of environmental factors, such as few and small
rivers, narrow shallow reef areas with few inner bays, andespecially strong water currents surrounding the island.

117

K. Koeda et al.
Acknowledgments
We are especially grateful to the supervisors of theauthors, H. Motomura (KAUM), S. Kimura (FRLM), Y.
Sakai (Hiroshima University), Y. Iwatsuki (MUFS), and K.
Tachihara (University of the Ryukyus) for their kindness
and support throughout the whole study. We deeply thankH. Yoshigou (Chugai Technos Co., Ltd.) for providing
environmental information of Tabaru River and photographsof fresh specimens collected at Yonaguni-jima island. We
also thank S. Takenishi (Yonaguni Fishery Association),
H. Muramoto (Education Board of Yonaguni Town), Y.
Shichijyo and H. Nakamura (Fisheries Division of Okinawa
Prefecture), and T. Yoshino (URM) for giving advice andsupport for our expedition. We greatly appreciate Y. Nishioka(Okinawa), many other anglers in Yonaguni-jima island,
and the staff of Diving Service MARLIN for providing alarge number of valuable fish specimens and underwaterphotographs, and J. Nakajima (Fukuoka Institute of Healthand Environmental Sciences) for providing information on
M. anguillicaudatus. We also sincerely thank K. Matsuura
and E. Katayama (NSMT), H. Senou (Kanagawa Prefectural
Museum of Natural History), and H. Iwatsubo (Kagoshima
Museum of Aquatic Biodiversity) for providing identificationsof Balistidae, Trichonotidae, Gobiidae and Mugilidae, andPomacentridae, respectively. We also thank G. Shinohara
and M. Nakae (NSMT), S. Nakamura (HMNH), K. Sato, S.
Oka, and K. Miyamoto (Okinawa Cyurashima Fundation)
for their support in examining deposited specimens. We are
deeply indebted to T. Shimose (Seikai National Fisheris
Reseach Institute, Fisheries Research Agency), F. ZiadiKünzli (University of the Ryukyus), students and volunteersof KAUM, especially S. Koeda, B. Jeong, S. Tashiro, H. Hata,
and Y. Fukui, for curatorial assistance, providing references,and processing specimen photographs. J.D. Reimer (Universityof the Ryukyus) improved the English of the manuscript. This
study was supported in part by a Grant-in-Aid from the Japan
Society for the Promotion of Science for JSPS Fellows (PD:26-477; DC1: 23-2553) (DC2: 15J02820) to the first authorand second author, respectively, and the Sasakawa Fund for
Scientific Research to the fourth and fifth authors.
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Authors at Hikawa Bay (left to right): Tomohiro Yoshida, Keita Koeda, Yuki Kimura, Saki Tanaka, Misaki Fujisawa, Momoko Sakurai (first row),
Taiga Kunishima, Keisuke Eguchi, Tomoki Inaba, Yusuke Hibino, Ryohei Miki, Hiromu Suzuki, Takanao Furukawa, Daichi Sasaki (second row),
Hidetoshi Wada, Takuya Uejo (upper windows)