光合成細菌の分類名についての追加情報（2023/4/17 更新）「付録 A．光合成細菌の分類名」の更新（2023 年 4 月）
本書発行以降、門の名称変更を含め多くの分類名の変更が行われ、また新種や新しい分類群の記載も行われた。以下に 2023 年 1 月の時点での階層分類名（学名）のリストを英文の説明とともに示す。
This Appendix lists the taxa of anoxygenic phototrophic bacteria so far described as being culturable. Theselists include bacterial taxa whose photosynthetic functions or chlorophototrophy have been demonstrated only
by genomic data. The taxonomic names are based on information from the LPSN - List of Prokaryotic nameswith Standing in Nomenclature (https://www.bacterio.net/).
Appendix A.1
Taxa of purple nonsulfur bacteria and anoxygenic aerobic phototrophic bacteria (phylumPseudomonadota)
付録 A.1 紅色非硫黄細菌および好気性光合成細菌（Pseudomonatota 門）ClassAlphaproteobacteria

Order
Caulobacterales
Hyphomicrobiales

GenusaFamilyBrevundimonas*2"Photocaulis"*1
Afifella
Fulvimarina*2
Methylocella*2
Blastochloris
Hyphomicrobium*2
Rhodomicrobium
Methylobacterium*2
Methylorubrum*2
Methylocystis*2
Bradyrhizobium*2
Rhodoplanes
Rhodopseudomonas
Agrobacterium*2
Hoeflea*3
Rhodobium
Rhodoblastus
Roseiarcus*1
Roseibium*2
Stappia*2
CereibacterDinoroseobacter*1
Fuscovulum
Neotabrizicola*1
Phaeovulum
Pseudotabrizicola*1
Rhodobaca
Rhodobacter*5
Rhodobaculum
Roseibaca*1"Roseibacula"
Roseicitreum*1
Roseinatronobacter*1
Rubribacterium
Rubrimonas*2
Jannaschia*2
Nereida*1
Planktomarina*1
Rhodovulum
Roseibacterium*2
Roseicyclus*1
Roseisalinus*1
Roseivivax*2
Roseobacter*2
Roseovarius*2

Caulobacteraceae
Maricaulaceae
Afifellaceae
Aurantimonadaceae
Beijerinckiaceae
Blastochloridaceae
Hyphomicrobiaceae
Methylobacteriaceae
Methylocystaceae
Nitrobacteriaceae
Rhizobiaceae
Rhodobiaceae
Roseiarcaceae
Stappiaceae

RhodobacteralesParacoccaceae

Roseobacteraceae

1Type species orphototrophic
representativea
B. subvibrioides"P. sulfatitolerans"
A. marina
F. pelagi(M. silvestris)
B. viridis(H. zavarzinii)
R. vannielii
M. organophilum
M. extorquens(M. rosea)(B. oligotrophicum)
R. roseus
R. palustris"A. albertimagni"(H.olei)R. orientis
R. acidophilus
R. fermentans
R. denhamense
S. stellulata
C. changlensis
D. shibae
F. blasticum
N. shimadae
P. veldkampii
P. algicola
R. bogoriensis
R. capsulatus
R. claviforme
R. ekhonensis"R. alcaliphilum"
R. antarcticum
R. thiooxidans
R. polymorphum
R. cliftonensis(J. cystaugens)
N. ignava
P. temperata
R. sulfidophilum
R. elongatum
R. mahoneyensis
R. antarcticus
R. halodurans
R. litoralis
R. tolerans

No. ofspeciesb
1–
3
1
14
1386
12810–
222
17
1
5
1
1
12
12
15
11–
12
1
12
1
120
12
13
33

Rhodospirillales

Acetobacteraceae

Azospirillaceae
Rhodospirillaceae"
Rhodovibrionaceae
Thalassobaculaceae
RhodothalassialesSphingomonadales

Rhodothalassiaceae
Erythrobacteraceae

Sphingomonadaceae

Betaproteobacteria

Gammaproteobacteria

Burkholderiales

Nitrosomonadales
Rhodocyclales
Cellvibrionales

Burkholderiaceae
Comamonadaceae

Not assigned to family
Sterolibacteriaceae
Rhodocyclaceae
Halieaceae

Salipiger*2
Sulfitobacter*2
Yoonia*2
Acidiphilium*2
Acidisphaera*1
Craurococcus*1
Elioraea*5
Paracaurococcus*1
Rhodopila
Rhodovastum
Roseococcus*2
Roseomonas*2
Rubritepida*1
Rhodocista
Skermanella*2
Aerophototrophica*1
Caenispirillum*2
Magnetospirillum*3
Pararhodospirillum
Phaeovibrio
Rhodospira
Rhodospirillum"Roseospirillum"
Roseospira
Rhodovibrio
Oceanibaculum*2
Thalassobaculum*2
Rhodothalassium"Citrimicrobium"*1
Erythrobacter*1
Novosphingobium*2
Blastomonas*1
Chioneia*1
Sphingomonas*2"Sandaracinobacter"*1
Polynucleobacter*2
Ideonella*2
Limnohabitans*2
Rhodoferax*3
Roseateles*2
Rubrivivax*3
Aquincola*2
Methyloversatilis*2
Rhodocyclus
Chromatocurvus*1

S. mucosus(S. noctilucicola)
Y. vestfoldensis
A. cryptum
A. rubrifaciens
C. roseus
E. tepidiphila
P. ruberR. globiformis
R. atsumiense
R. thiosulfatophilus(R. aestuarii)
R. flocculans
R. centenaria(S. aerolata)
A. crusticola(C. salinarum)(M. fulvum)
P. photometricum
P. sulfidophilus
R. trueperi
R. rubrum"R. parvum"
R. mediosalina
R. salinarum
O. indicum(T. salexigens)
R. salexigens"C. luteum"
E. longus(N. subterraneum)
B. natatoria
C. frigida(S. koreensis)"S. sibiricus"
P. necessarius(I. sakaiensis)(L. planktonicus)
R. fermentans
R. depolymeransR. gelatinosus
A. tertiaricarbonis(M. universalis)
R. purpureus
C. halotolerans

1215113111211221153111–
52111–
82646–
111222113
1Congregibacter*1C. litoralis
L. syltensis
P. rubra

1Luminophilus*1
Pseudohaliea*1aSpecies given in parentheses are not the type but a phototrophic representative.
Number of phototrophic species validly published under the ICNP.
*1 Genera comprising AAnPB (aerobic anoxygenic phototrophic bacteria) species only
*2 Genera comprising AAnPB and chemotrophic species*3 Genera comprising PNSB (purple nonsulfur bacteria) and chemotrophic species*4 Genera comprising PNSB, AAnPB, and chemotrophic species*5 Genus comprising PNSB and AAnPB speciesb211

Appendix A.2 Taxa of purple sulfur bacteria (phylum Pseudomonadota)
紅色硫黄細菌（Pseudomonadota 門）
Class

Order

Gammaproteobacteria

Chromatiales

FamilyChromatiaceae

Ectothiorhodospiraceae
Halorhodospiraceae"
aGenus
Allochromatium
Caldichromatium
Chromatium
Halochromatium
Imhoffiella
Isochromatium
Lamprobacter
Lamprocystis
Marichromatium
Pfennigia
Phaeobacterium
Phaeochromatium
Rhabdochromatium
Thermochromatium
Thioalkalicoccus
Thiobaca
Thiocapsa
Thiococcus
Thiocystis
Thiodictyon"Thioflavicoccus"
Thiohalocapsa
Thiolamprovum
Thiopedia
Thiophaeococcus
Thiorhodococcus
Thiorhodovibrio
Thiospirillum
Ectothiorhodosinus
Ectothiorhodospira"Halorhodospira"Type species orphototrophic
representaive
A. vinosum
C. japonicum
C. okenii
H. salexigens
I. purpurea
I. buderiL. modestohalophilus
L. roseopersicina
M. gracile
P. purpurea
P. nitratireducens
P. fluminis
R. marinum
T. tepidum
T. limnaeus
T. trueperi
T. roseopersicina
T. pfennigii
T. violacea
T. elegans"T. mobilis"
T. halophila
T. pedioforme
T. rosea
T. mangrovi
T. minorT. winogradskyi
T. jenense
E. mongolicus
E. mobilis"H. halophila"

No. ofSpeciesa
71
2
3
2
1
12
4
1
1
1
1
1
1
1
5
1
62–
2
1
1
2
6
1
1
1
7–

Number of phototrophic species described phenotypically and/or genotypically and validly published.

3Appendix A.3 Taxa of green sulfur bacteria (phylum Chlorobiota), chloroflexus and relatives (phylumChloroflexota), heliobacteria (phylum Bacillota), chloracidobacteria (phylum Acidobacteriota), gemmatimons(phylum Gemmatimonadota), and eremiobacteria (phylum "Candidatus Eremibacterota")
緑色硫黄細菌（Chlorobiota 門）、クロロフレクサス（Chloroflexota 門）、ヘリオバクテリア（Bacillota 門）、クロラシドバクテリア（Acidobacteriota 門）、ゲマチモナス（Gemmatimonadota門）、エレミオバクテリア（"Candidatus Eremibacterota"門）、
Class

Order

FamilyChlorobiia

Chlorobiales

Chlorobiaceae

Chloroflexia

Chloroflexales

Chloroflexaceae

Oscillochloridaceae
Roseiflexaceae"Thermofontia"
Clostridia"Phototrophicales"
Eubacteriales"Phototrophicaceae"
Heliobacteriaceae

Terriglobia
Gemmatimonadetes"Candidatus
Eremiobacteria"

Terriglobales
Gemmatimonadales"Candidatus
Baltobacterales"

Acidobacteriaceae
Gemmatimonadaceae"Candidatus
Baltobacteraceae"

ab

Genus
Ancalochloris
Chlorobaculum
Chlorobium
Chloroherpeton"Chloroplana"
Pelodictyon
Prosthechochloris
Chloroflexus
Chloronema
Oscillochloris
Heliothrix
Roseiflexus"Phototrophicus"
Heliobacterium
Heliomicrobium
Heliophilum
Heliorestis"Chloracidobacterium"
Gemmatimonas"Vulcanimicrobium"

Type species orphototrophic
representaivea
A. perfilievii
C. tepidum
C. limicola
C. thalassium"C. vacuolata"
P. clathratiforme
P. aestuarii
C. aurantiacus
C. giganteum
O. chrysea
H. oregonensis
R. castenholzii"P. methaneseepsis"
H. chlorum
H. gestii
H. fasciatum
H. daurensis"C. thermophilum"(G. phototrophica)"V. alpinus"

Species given in parentheses are not the type but a phototrophic representative.
Number of phototrophic species described phenotypically and/or genotypically and validly published.

一部改変 Anoxygenic Phototrophic Bacteria (2023) in press

4

No. ofspeciesb
1251–
3
33
12
1
1–
2
3
1
3–
1–