ğƇʊϮЕ 28 (3) 2018

光合成研究

О 28 Ȱ О 3 ƅׄԵȰ 83 ƅ‫ׅ‬2018 Ⱥ 12 ̀
NEWS LETTER Vol. 28 NO. 3 December 2018
THE JAPANESE SOCIETY OF PHOTOSYNTHESIS RESEARCH
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MGDG Iƍՙ)ΪϊϖG΀ȕ4B$c<Z

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I̩Ǭ̸Է)ȥǉ4PLB J:IWW DGDG ]̕G'$BǑǓFcӺΰg$<=*W4<0IáI®¦®ҀԊG`bʑȹFҥ̸҅ԷGǍˀ2ǂgċbBű,ɡϸπ4¼/W6̊ϮЕJ

dcI=e%4(4:IɜJ LHCII FEIJSPS ЄϮԂׄ16J10176 2671101626440170‫ׅ‬ğƇʊ҅´vԊ)ңА87W<s®vҀIœʊgż.BұfdW4<ԊƇʊIʙń[ь,ICҡёôɒʊJձȀ2d
Received Nov 15, 2018; Accepted Nov 20, 2018; Published<WWGFcDʺ΃2dc·˝dgd1 Ǎϊô
CJ DGDG Iՙ)֣ҢG΀ȕ4B$c<ZPLB

Dec 31, 2018.

CJ MGDG IŎƇ)ϊȶGָ$ΫʃGF?B$cğΠȐɜLPOR JĻӊ2d PLB I̩Ǭ̸ԷJȥdc)͏2d< MGDG JūέC®¦®

ŵѮ˙ΰ
1.

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theRole of galactolipids in etioplast development and etioplast-to-chloroplast differentiation

1

2

1

1

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Sho Fujii , Noriko Nagata , Tatsuru Masuda , Hajime Wada and Koichi Kobayashi *1

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8.

͑ռWC Ycf4 §ª¶°)խ¿6c˿ɜIĻǬօƇԿЊJLHCI8 IцƇgǵǺŠ6c PsaG
9.

c0IԿЊG`b PSI-LHCI )Ƕʊ4Ycf4 §ª¶°(aӊֈ6cPSI ҿƇôGJziv

WdB$F$0IԿЊIǓՍĻJ̈ӊ˦IWW

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C#b:Iӊ˦JÞɜIϮЕIԺȜĞɚ4<

11.

Received Dec 11, 2018; Accepted Dec 13, 2018; PublishedAug 31, 2018.

12.

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Sreedhar Nellaepalli, Shin-ichiro Ozawa, and Yuichiro Takahashi*
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156ğƇʊϮЕ 28 (3) 2018

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1

1

2,

Atsuhiro Hirohata , Kyohei Uemoto , Motomu Endo *1

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Bʫ/adB$<4(4σĖӊ̕D$%ӅΛʜǻֶCϷ4<δƱ0%4<ǻӗϮЕGխ6(aXcD։˧ɃJñ,ˡG$,@(IÅFcǁƏJW=ȕF$Þɜ0%4<ǻӗϮЕG¶°)ժϒ2dB$c 33–38)ϒҒϟɜ#c$J`by¤v±v~¬´I¢´~¨°gЈ̰ϟɜIǻλI HTP JǻֶǽīCI»͆Ĝ

ĨôϖGӘþ4B$,ɨӂ)#c

ɒΫѧҠ̸ԷƣǹIɒΫµ|lIӊ̕ 39–42)
ƭǂCIϒҒβӧ̢Ʋ¼²¡ 43–45)]²¶

3.

l°¶nl´wׄHTP‫ׅ‬´ 46,47)gν$<ԩʹˉɕG`cևҜӱŁFEҿ

 HTP JָŖβµָеɃG̰ΩIҶδƵgӎևFɒʃg[@÷ΩGȌ4B[́ŖF»͆ĜĮ

΃6c0DgϞ̻D4B$c)ԩȺIʗҲIԺ̸Ъӊ̕ʒͪɤȔćôIӟӱĻ֥FEI։͌G`b:d)ƀѿDFb@@#cHTP J֠Gʪʌ4<ϮЕû)ЋǁƏ2dB$cW<̣֭ʊէ̆ḬΩGȌ6c HTP G@$BJÅЋǬIՇŁ(aƭǂCIźГƲ¼Ս(aƲ½ՍWC̹FɒԊgȌӷG̹Fx¶°Cұ

G̰Ҹβ 48)ѧҠIָ2 49, 50)ϐȀ]±I

fdB$cׄƪ 1‫̰ׅ‬ΩIҶδƵӎ΃gl̱Ĺ 51–56))ϮЕȌӷD2dB'bǓƵțīn°¶Š6cGJσĖ´~´wḒ͂Ǳl´w˞Ӕ]՘ǐIƭǂCIՅνû)ȿ

ѬgфXƇf8<ʗҲ)֏ȶǴŖC#c

29–32)

,ǁƏ2dB$cW<λ͐ʊէ̆ḬΩGȌǑ̹FσĖ´|¶gϊFc͂̍ׄƫǺ®

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ӎ˕ 57–65)FEQIՅν)ûD4Bʫ/adcä6c0DCǽīµ՘ǐgƛf7ǓՙIσĖ¶½՘ǐIƭǂG'.cӎ΃QIՅνûgÃɧGgŻɠµӊ̕C*cû&KЋǬIҶδƵӎϊFcλѹ̆լGȌ6c HTP ʗҲg·û7@т

΃I<ZIσĖˉɕJÅGǽīCұfdc)ğß4F)a:IδΫDƛ֠ΛG@$BԬRc

·IǍŠGȌ6c֛ĎɲDÓ$Gџ)b]՗Fƪ 1. ϊFcx¶°G'.c HTP Iû(a) ԎԩʹˉɕGƼA, HTP IûָӊĖɃσĖ]»͆Ĝɻǁ)ָ$֞ɃCŻɠC*c<ZҜºҡ֐А̰Ҹβlq£Iхˮϖӎ΃)ƀѿ(b) ƫǺr¦®]²¡gν$<ˉɕGƼA, HTP IûƭǂCӈ΃4<$ȕ˕ḬΩ]²G@$BˮзľCˉɕ6c̰ҸβʊէԶɃʊէ˻ѓIʺǺ)ƀѿ(c) ñЖ²¶´gν$<ˉɕGƼA, HTP Iû²¶´I҈ŗҍұG`bƭǂĤôgȌӷGˉɕgұ$˼Gҿ˕IıϤ`b»͆ĜƇʊĤƭǂI£´w²ĻŎ)C*c(b)IǺΛӈ΃`b˕Ǒ,ḬΩ]²G@$B̰ҸβʊէԶɃʊէ˻ѓIʺǺ)ƀѿ

184ğƇʊϮЕ 28 (3) 2018
3.1. ЋǬI HTP

3.2. ϒҒµЈ̰ϟɜIȽ̰ΩI HTP̢IտGłν2dcӎ΃I·ҎϖFͱdJσ

Ǔ*,Ļ.Bǽī̪ƻ#c$Jƭǂ̪ƻgƼGĖŻɠЋǬDѻ˲IĻֈЋǬIʡĹЋǬI

ұ%[IGĻ.adcǽī̪ƻgƼG4< HTPɒʃӎ΃ц̘ĹŏDFc0I·ԸIĸζIÃ

CJ÷ΩI»͆Ĝ̸Է 67)Iӎ΃)ұfdc0D

CJˉɕˮIğ̐ç)ǍŠ6cÃCЋǬDѻ˲)Ǒ,·ҎϖFͱdD4BJ»͆ĜΛѧIĮgĻֈ6cDЋǬƈǊ)џ)b]՗F̸Ъ̰Ω֜ƹIΛѧIʡĹƆƣǹIʡĹƆ

bg[@ÃCćIЋǬgʡĹ6c)ƣǹIӘþ)ұFfdćҡI̗˕֐АλѹӦ֠DFcF'0dagƛ֠F,ǻ¶´ 40)̙IĻȲ 42)FEӝсFҶδƵ)

˞C*dK|l]ɒΫFEIΪɥIǺՙϖӎӎ ΃ 2 d c  » ͆ Ĝ Λ ѧ I Į ̸ Ъ C J  ToF ЋǬI HTP JÅGźГɜIźՙ]ƗԊIӧ ϒҒϟɜ#c$JЈ̰ϟɜIȽ̰ΩI HTP Jׄtime-of-flight‫ׅ‬LiDAR ׄLight Detection and΃]Әþgұ%<ZImnjJˡGʾüF'0dWCJÅGźГ

Ranging‫ׅ‬Structured-lightStereo visionSfMɜGǽīCˉɕµӊ̕gұ?B$<)˿ԩCJ

ׄStructure-from-motion‫ׅ‬FEI̹FʗҲ)#՘ǐCIӊ̕gƀѿG6c¶°[ǁƏ2dB

c0daIʗҲJӎ΃еɃӎ΃zɫ

2dB$c

33,66)$cû&KMakanza aνǂ֐)ϊFc 67))˿ԩJȱӼI RGB r¦®

38)

Jm§²z~gȌ

g 1 Ɓν$<=.C[ָ$еɃCI»͆ĜΛѧ

ӷG4B՘ǐC[ʒԝGˉɕgұ$Џ]ЎвIĮ̸Ъ)C*c SfM )˿[ͬϞ2dB'b 68‫ׅ‬I˕]ɒFEgӎ΃C*c¶°g÷ʊ4<ׄƪ2‫ׅ‬Þɜ2aFcϒȜ[̆ɚC*c̰Ω֜ƹIΛ
ѧIʡĹDƆƣǹIʡĹG@$BJϊFcʗҲƪ 2. ՘ǐCЋǬIӎ΃gұ?<û 38‫ׅ‬(a) ńɡğD҈Οğgν$<σĖIŻɠ(b) σĖӊ̕gν$<ЎвIʡĹDЏDЎвIɒʃΪɥIӎЧ(c) ̹FҐv¨|lg[@ЏG@$BϊFcğ·I[DCЏDЎвgʡĹ4<ц̘(d) σĖӊ̕gν$B՗F?<ЎвgĻֈ4ָеɃGӎ˕6c

185

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3.3. ̣֭ʊէ̆I HTP:d;dIʗҲGȌ4BŁGȌФgӯ7cɨ ̣֭ʊէ̆ḬΩGȌ6c HTP Jǽī̪ƻӂ)#cW<Ƚ̰ΩC[ˡGҿևFɒΫg[IǂƇGJÅGćô±°ƭǂ̪ƻIǂƇGJ

@<ZÓ$G՗F?<ՍĻIʡĹFE)˿Ǔ

ѧҠ±°Cұfdc·ҎϖFͱdD4BJ

IӦ֠DF?B$cׄƪ 3‫ׅ‬ƭǂ̪ƻgƼG4σĖŻɠ̰Ω֜ƹIʡĹȌӷɒԊIӎ΃µӘ< HTP IǂƇGJƲ¼ƁԘ/²¡ׄ҈ɛԌþDFcȌӷɒԊD4B̰ҸβϐȀ]

ұ]ʒʠ4ɋ‫]ׅ‬ΝÜ֧ұ͂głν4BÑ͆Ĝ±IЊɃG@$BˡGǑ,Iʒͪ)ժϒ2d

]Ŀ̆ʊէ

HTP ÍŖɲ)Ϸ2dB$c 48,51-56)4(4̰Iͯɲ 73)FEIӎ΃)Ǒ,ǁƏ2dB$c·ҎΩѧҠIָ2Iӎ΃J̈=ժϒ͑ռC#b͏2

ϖFӊ̕IͱdJЋǬI HTP DǓ*,Ǎfa

d<Ӧ֠)#cû&KWang a 74)J°s

F$)ćôgĻֈ6cտGğ̐çIǍŠIá

¥IѧҠIָ2Iӎ΃gȌӷGϊFcσĖŻɠσĖӊ̕`bϒҒβćôӎ˕47,69–72)72‫ׅ‬]ćôDևҜլI՗Fb)Ǔ*F

˝ͪG@$B΃ՙеɃ'`NzI͖Ԟg4

Ӧ֠DF?B$cΪG÷ΩDևҜIţŁG@$<ׄҶ 1‫ׅ‬Madec a 49)JLiDAR ΃ՙDΝÜҍ

GćôլЖ͂Gˆԟ4< RGB r¦®G`c SfM ΃ՙI͖

BJ̈=ǴŖFʒͪ)F$F'ƭǂ̪ƻ
gƼG4< HTP CI»͆ĜΛѧIĮ̸ЪDӝс

Ԟgұ?<W<Hu50) JΝÜҍЖ͂Gˆԟ4

Fӎ΃ʗҲ[ǁƏ2d@@#c)ϊFc΃ՙ͂< RGB r¦®G`c SfM ΃ՙIXIǂƇGѧ

ƣլCIц̘I͖Ԟ]θǆIɕ֕ΪG֦G`

ҠָIӎЧ˝ͪI͖Ԟgұ?<Ʋ¼ƁԘ/²

cɕ֕QIȌФ)Ӧ֠DF?B'bÞɜIӎ΃¡IǂƇJE>aIʒ͑gν$B[ҏ$ʗҲIϒȜĞɚ4<$еɃ)ɠadc0D)Ļ(?<)ΝÜҍЖ͂IǂƇGJr¦®IӊĖɃG[Ǔ*,ýǯ6c

ƪ 3. (a) z¥uIϒҒɜ 2 ԹլŉɜIσĖ 72)ƪÃIΛJćôDƲ֐IʹΛIðѣgϷ4B$cćôլI՗Fb)֏ȶGָ$0D)Ļ(c(b) lIЈ̰ɜ 3 ԹլŉɜIσĖևҜDl'`Nlƈ
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SfM gǻұ6cտIƼ϶®¦¶DFcȗʐ

4Bӟӱ28ӎ˕28F.dKFa7:0GΪɥՙӎЧͪDĿ̆ӔǺׄΪɥΛI˕ΪɥΛ

ָɃF̭͂ǱѬIʗҲ)ɨӂDFcGuo a 59)£´wI˝ͪFE‫ׅ‬r¦®īՍ®¦¶J2 ͑ռI̭͂ǱѬͪgν$cʒͪgʾ̬4<Ŀ̆ӔǺȅFΛѧgλʊ6c<ZIj°{¯δƱCJɱԶGϒȜ4@@#cͻȞǱѬ[̱¥FEG@$Bәұՠӣ6c0DC˿ՅFӔǺĹµӱŁ§°̸ЪIʒͪD4BͬϞ2dB$c

gʸ4Błν4B*<)ÞɜJ0%4<ӔǺͻȞǱѬCJťĻF˕I±¶´w¶)I҈ŗŠG@$B[2aFcϮЕ)ɨӂDѮ&#dK`bָ$еɃCI̱ĹDӎ˕)ƀѿD

adc

F?B$cׄû&KĹЏ 65,75,76)̘ǻ 64)‫ׅ‬

3.4. λ͐ʊէ̆I HTP

4. y¤v±v~¬´'`N HTP gν$<

 λ͐ʊէ̆I HTP IÅFȌӷɒԊJժґˮՇʜIŖβլЏ]̘ǻI˕ʊΣɃFE)ʫ/adB$c
̣֭ʊէ̆I HTP DϊFbȌӷΩIʡĹCJ ѹЋG'.cՇʜIŖβgӘþ6cʩ̻D4F,ȌӷΩI̱ĹDӟӱ)ƛ֠GFb̱ĹD̻)#c 77)ՇʜŹɫ𝑅𝑅JՇʜ2d<Ӈ(aɠBՇʜŹɫׄresponse to selection‫`ׅ‬Kdcʩ

ӟӱI<ZI§°̸Ъ)Ӧ֠DFcW<ć

ad<ǬüIҶδƵIȹƴ𝑦𝑦C DՇʜŉIӇօƧôլРI՗Fb[ƛ֠DFcû&KѹЋƭ

G'.cҶδƵIȹƴ𝑦𝑦D IȮ6Ff>

ǂG'$B°s¥IĹЏIӟӱḎĹgұ?

𝑅𝑅 = 𝑦𝑦C − 𝑦𝑦D<ǂƇƪ 4 I`%GϊFcğ̐çĹЏ̆լ

D4BǺѩ2dcÞՇʜȌӷD4B$cɒԊƗЋµзщЏƈǊI՗FbРG`bσĖgԵ
4Bʰ&adcЏIҐ|lɒ)ǍŠ6c

IՈìβgℎF ՇʜȮׄselection differential‫𝑆𝑆ׅ‬gӇօƧG'.cҶδƵI̻ΈčȮσD DՇʜɐɃ𝑖𝑖

4(4z´ª¶GJ:dagƈ·I[IDIА𝑖𝑖σD D4BҶ6D

ƪ 4. ѹЋƭǂCˉɕ2d<̹FзщI°s¥IĹЏ 59‫ׅ‬187ğƇʊϮЕ 28 (3) 2018
𝑅𝑅 = 𝑖𝑖ℎF 𝜎𝜎D

DFc2 @IɒԊIՇʜɐɃ)ƈ5ǂƇׄ𝑖𝑖U =𝑖𝑖O ‫ׅ‬y¤v±v~¬´IÏ΃ČDɒԊ𝑋𝑋I

D$%խĀGFc0IɋJѹЋȁI˝Њɋׄbreeder’s equation‫ׅ‬ՈìǬƵČIϠխ𝑟𝑟OU )ɒԊ𝑋𝑋IՈìβIȹ˝̨

D`KdcѹЋ¼՗ӂFℎO g¼ƥcDy¤v±v~¬´G`cՇ

ɋC#bȌӷɒԊIՈìβℎ F DӇօƧG'.

ʜIŖβ)ɒԊ𝑋𝑋IϟʹՇʜIŖβg¼ƥc

𝑖𝑖I[DC̆ɚ2dcՇʜŹɫ𝑅𝑅gӎЧC*cÞɒԊ𝑌𝑌gν$<լʹՇʜID*Dƈ̹𝑖𝑖U > 𝑖𝑖O C

cҶδƵI̻ΈčȮ𝜎𝜎D )Ļ(dK#cՇʜɐɃՈìβℎF = 𝜎𝜎JF /𝜎𝜎DF głν6cD
𝑅𝑅 = 𝑖𝑖ℎ𝜎𝜎J#dKy¤v±v~¬´IÏ΃Č𝑋𝑋V DɒԊ𝑋𝑋IՈìǬƵČIϠխ𝑟𝑟OU )ɒԊ𝑋𝑋IՈìβ
g¼ƥaF$ǂƇC[ϟʹՇʜg¼ƥcŖβCDҶ2dcF'𝜎𝜎J JӇօƧG'.cՈìĻՇʜC*cƀѿɲ)#c

˒Iȹ˝̨C#c y¤v±v~¬´CJ¼Ԭ4<`%G

 00CˌҏȌӷD6cɒԊ𝑋𝑋gHTP Cӎ΃ØȮ̱ӗG`?BÏ΃ČDҶδƵČ𝑃𝑃IϠխg

2dcɒԊ𝑌𝑌gν$BլʹϖGՇʜ6cǂƇgӎЧ4еɃIʩ̻D6cǂƇ)Ǒ$0IϠխѮ&c0ID*ɒԊ𝑌𝑌gν$<ɒԊ𝑋𝑋Iլʹ

J

ϖFՇʜŹɫ𝐶𝐶𝑅𝑅O J𝐶𝐶𝑅𝑅O = 𝑖𝑖P ℎP 𝑟𝑟OP 𝜎𝜎JO

𝑟𝑟XU =

DҶ2dc00C𝑖𝑖P JɒԊ𝑌𝑌IՇʜɐɃℎP JɒԊ𝑌𝑌IՈìβIȹ˝̨𝑟𝑟OP JɒԊ𝑋𝑋DɒԊ𝑌𝑌I

𝑐𝑐𝑐𝑐𝑐𝑐ׄ𝑃𝑃, 𝐼𝐼‫𝑋𝑋ׄ𝑐𝑐𝑐𝑐𝑐𝑐 ׅ‬, 𝐼𝐼‫ׅ‬=𝜎𝜎DO 𝜎𝜎U
𝜎𝜎DO 𝜎𝜎U=ՈìϠխ𝜎𝜎JO JɒԊ𝑋𝑋IӇօƧG'.cՈìĻ˒Iȹ˝̨C#c0d)ɒԊ𝑋𝑋gϟʹՇʜ4

𝜎𝜎JO 𝑐𝑐𝑐𝑐𝑐𝑐ׄ𝑋𝑋, 𝐼𝐼‫ׅ‬∙= ℎO 𝑟𝑟OU
𝜎𝜎DO
𝜎𝜎JO 𝜎𝜎UDҶ2d4<)?By¤v±~¬´g<ǂƇIՇʜŹɫ𝑅𝑅O = 𝑖𝑖O ℎO 𝜎𝜎JO `b[Ǔ*,Fν$<ǂƇĞɚ2dcՇʜŹɫJØȮ̱ӗgc6Ff>ՇʜIŖβ)ָWc̐çJν$Bɠadc𝑟𝑟XU gν$B

𝑖𝑖P ℎP 𝑟𝑟OP > 𝑖𝑖O ℎO

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Wei Guo, Hiroyoshi Iwata*Graduate School of Agricultural and Life Sciences, The University of Tokyo193ğƇʊϮЕ 28 (3) 2018ƻƚ0Ɲ

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Antarctica, during 2013. JARE Data Rep., Terrestrial Biol. 8, 1–99.
2. Kosugi, M., Maruo, F., Inoue, T., Kurosawa, N., Kawamata, A., Koike, H., Kamei, Y., Kudoh, S. andImura, S. (2018) A comparative study of wavelength-dependent photoinhibition of drought-tolerant
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1. Berthold, D.A., Babcock, G.T. andYocum, C.F. (1981) A highly resolved,
oxygen-evolving
photosystemIIpreparation from spinach thylakoid
membranes. EPR and electron-transport
properties. FEBS Lett. 134, 231–234.
2. Nanba, O. and Satoh, K. (1987) Isolationof a photosystem II reaction center
consisting of D-1 and D-2 polypeptides
and cytochrome b-559. Proc. Natl. Acad.
Sci. U.S.A. 84, 109–112.

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3. Diner, B.A. and Babcock, G.T. (1996)
Structure,dynamics,
andenergyconversion efficiency in photosystem II,
in Oxygenic Photosynthesis: The Light
Reactions (Ort, D.R. and Yocum, C.F.,
Eds.) pp 213–247, Kluwer, Dordrecht,
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