Proceedings of the 7th and 8th Symposia on Collection Building and Natural History Studies in Asia and the Pacific Rim,
edited by Y. Tomida et al., National Science Museum Monographs, (34): 239–272, 2006.

Faunistic Review on the Pselaphine Species Known from New Zealand(Insecta, Coleoptera, Staphylinidae)
Shûhei Nomura1 and Richard A. B. Leschen2
1

Department of Zoology, National Science Museum, Hyakunincho 3–23–1, Shinjuku, Tokyo, 169–0073 Japan(e-mail: nomura@kahaku.go.jp)
2
New Zealand Arthropod Collection, Landcare Research, Private Bag 92170, Auckland, New Zealand(e-mail: leschenr@LandcareResearch.co.nz)

Abstract A checklist for the 42 genera and 385 known pselaphine species of New Zealand is provided with data on their original descriptions, type localities and original species numbers by Thomas
Broun, the author of most species. Systematic notes on each genus were given and all taxa are placedin the current pselaphine classification. Euplectus iracundus (Broun) is transferred to the genus Euglyptus. A list of the “syntypes” of Broun’s species from New Zealand contained in Deutsches Entomologisches Institut, Muncheberg are also provided.
Key words:

Pselaphinae, Staphylinidae, fauna, systematics, New Zealand.

Introduction
The subfamily Pselaphinae is one of the most diverse subfamilies of Staphylinidae and in
New Zealand (NZ) they occur at sea level to the alpine zone, in tree holes and under rocks, inmoss and rotten wood, and are especially abundant in leaf litter, making them ideal for ecologicalstudy. And despite their relatively small size, these beetles have extraordinary body forms thatbear foveae, pits, sulci and grooves that facilitate their identification and make them a fascinatinggroup for taxonomic study. Another feature of the NZ fauna is its richness with endemic taxa, especially Faronitae and Euplectitae, while Goniaceritae and Clavigeritae are completely absent, aclassic case of disharmony indicative of islands. The only drawback to the study of NZ pselaphines is that the group has never been revised, there are perhaps hundreds of undesribed
species, and the biology is poorly known.
The first pselaphine species were described in the 19th Century, first by David Sharp (1874)
who described 30 species on the basis of a collection sent to him by Thomas Broun. Most of the
New Zealand species of Pselaphinae were described by Broun from 1883 to 1922, and he named
334 species, 86.8% of the existing names. During Broun’s time, Raffray (1893, 1904b) and Reitter (1880, 1885) also named several species. Following these early workers, only a few species
have been described during following 80 or so years. Park and Pearce (1962) defined a new genus
Simkinion with descriptions of two new species from Northland. Park (1964) also described a
new genus and a new species, Pselaphotheseus hippolytae from Campbell Island, which is thefirst record of pselaphine species from the NZ subantarctic islands. Recently, Carlton andLeschen (2001) added the second species of Pselaphotheseus, P. ihipuku also from Campbell Is-

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land. Here we provide a checklist of genera and species described and recorded in NZ; we also
provide data on their original descriptions, type localities and original species numbers by Broun.
This is largely based on examination of specimens in the New Zealand Arthropod Collection,
Auckland (NZAC) by S. Nomura.
This is not the first checklist for pselaphines; this group was included by Hutton (1904) in his
catalogue of NZ insects. A more recent checklist of the NZ staphylinid genera that included pselaphines, was published by Klimaszewski et al. (1996): these authors listed taxa that were endemic and introduced and enumerated the number of species for each genus. In a large revisionof Australian pselaphines, Chandler (2000) also recorded additional taxa from NZ that he recorded from both areas. Newton and Thayer (2003) presented a comprehensive on-line database of all
staphylinid species known from the Southern Hemisphere, and the NZ pselaphine checklist we
provide is based on their work and modified further to include additional information.
Materials and Methods
Systematic notes in the following were based on the identified material of pselaphines contained in the NZAC and recently collected material obtained by the authors during fieldwork in
NZ. Most data (species names, original descriptions, the type localities and distributional ranges)
are extracted from the on-line database by Newton & Thayer (2003). The higher classification
system of Pselaphinae proposed by Chandler (2000) is adopted for this checklist. We also provide a list of Broun material inappropriately labeled as types in the DEI (Deutsches Entomologisches Institut, Müncheberg, Germany) examined by S. Nomura.
Systematic Notes
In this chapter, all scientific names and synonyms of pselaphine genera and species known
from NZ are given. The genera are listed according to systematic position by family-group andlisted alphabetically: genera that were not placed prior to this study are located based on theirsimilarity to related taxa. For each species, the data of original description are followed, authornames, years and the following symbol (a, b, c, ...) of which are coincident with the papers cited.
The type localities are added and the distributional ranges in NZ are indicated by the geographical area codes developed by Crosby et al. (1998) (Plate 8) in square brackets. Though geographic
information for all species is limited we indicate those species recorded in Lynfield, Auckland, by
Kuschel (1990) by an asterisk (*). For each of Broun’s species, his reference numbers are indicated by angle brackets behind the page numbers. Junior primary homonymns are indicated byacronym (JPH).
Supertribe FARONITAE
Tribe FARONINI
Genus Exeirarthra Broun (Figs. 1–3)
Exeirarthra Broun, 1893a: 1054. Type species: Exeirarthra enigma Broun by monotypy.

Remarks. This genus is very similar to the more diverse genus Sagola and is separated
from it by having the enlarged second segment of the protarsus in the male (Fig. 3) and the very
short and scarcely visible palpal spine on the fourth palpal segment. While these characters are
distinct in the type species, they vary among the other species of the genus and may not be re-Faunistic Review on the Pselaphine Species Known from New Zealand241

garded as diagnostic for the genus. Sagola planicula Broun was transferred to Exeirarthra by
Kuschel, 1990, though it should be Sagola, because it does not have key characters of this genus.
List of species:
E. angustula Broun, 1917: 374 3824. Glenhope, near Nelson [NN].
E. enigma Broun, 1893a: 1054 1885. Near Howick [AK]* (Figs. 1–3)
E. longiceps Broun, 1917: 375 3825. Hollyford, north of Lake Wakatipu [OL].
E. pallida Broun, 1893a: 1424 2474. Maketu, Hunua Range [AK].
E. parviceps Broun, 1921a: 486 3992. Mount Hope, near Nelson, nearly 4000 ft. [NN].
Genus Sagola Sharp (Figs. 4–14, 97–102)
Sagola Sharp, 1874c: 506. Type species: Sagola misella Sharp, designated by Oke, 1928: 5.

Remarks. This is a very large and morphologically diversified genus distributed in NZ andAustralia. It includes 131 species known from NZ. This genus is characterized by the short median longitudinal sulcus on the frons and the clearly transverse head.
List of species:
S. acuminata Broun, 1921a: 498 4009. Routeburn [OL].
S. aemula Broun, 1921a: 496 4006. Mistake Basin, Canterbury [MC].
S. affinis Broun, 1921a: 500 4012. Mount Algidus [MC].
S. angulifera Broun, 1911b: 491 3363. Waimarino, 2700 ft. [TO].
S. anisarthra Broun, 1893a: 105 1884. Moeraki [DN].
S. arboricola Broun, 1921a: 502 4015. Hollyford, 3500 ft. [OL].
S. auripila Broun, 1911b: 500 3371. Erua, 2400 ft. [TO].
S. basalis Broun, 1911b: 496 3368. Paparoa Bush, near Howick [AK].
S. bifida Broun, 1915: 290 3706. Woodhill, Kaipara Railway [AK].
S. bifoveiceps Broun, 1912b: 629 3489. Greymouth [BR].
S. biimpressa Broun, 1912b: 630 3490. Greymouth [BR].
S. bilobata Broun, 1921a: 486 3993. Mounts Dick, Alfred & Earnslaw, near Lake
Wakatipu, over 1000 ft. [OL] (Figs. 4, 97, 98).
S. bipunctata Broun, 1886: 887 1580. Near Whangarei Harbour [ND].
S. bipuncticeps Broun, 1921a: 499 4011. Mount Algidus, Canterbury [MC].
S. bituberata Broun, 1914b: 160 3525. Greymouth [BR].
S. brevicornis Raffray, 1893b: 27. Auckland [AK].
S. brevifossa Broun, 1921a: 501 4014. Routeburn [OL].
S. brevisternis Broun, 1915: 284 3699. Pudding Hill, near Methven [MC].
S. brevitarsis Broun, 1886: 887 1579. Paparoa, south of Auckland [AK].
S. carinata Broun, 1912b: 622 3482. Greymouth [BR].
S. castanea Broun, 1886: 884 1573. West Taieri bush [DN].
S. cilipes Broun, 1921a: 491 3999. Waitakerei Reservoir, near Auckland [AK].
S. citima Broun, 1893b: 177 none. Wellington [WN].
S. clavatella Broun, 1912b: 631 3491. Greymouth [BR].
S. cognata Broun, 1911b: 494 3365. Tairua [CL].
S. colorata Broun, 1914b: 156 3519. McClennan’s Bush, near Methven [MC].
S. concolorata Broun, 1915: 292 3709. Tairua [CL].
S. confusa Broun, 1915: 286 3701. Hunua Range, Clevedon [AK].
S. convexa Broun, 1886: 889 1583. Tuakau, Waikato [AK].

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S. cordiceps Broun, 1921a: 493 4002. Mount Alfred [OL].
S. crassulipes Broun, 1915: 283 3697. Epsom, base of Mount Eden [AK].
S. deformipes Broun, 1880: 138 252. Tairua [CL].
S. denticollis Broun, 1880: 138 253. Tairua [CL].
S. dickensis Broun, 1917: 377 3827. Mount Dick, near Kingston [OL] (Figs. 5, 99, 100).
S. dilucida Broun, 1914b: 157 3520. Epsom, base of Mount Eden [AK].
S. disparata Broun, 1914b: 160 3524. Old Man Range, Otago [CO].
S. dissonans Broun, 1921b: 601 4165. Tauherenikau Bush, near Featherston [WA].
S. distorta Broun, 1921b: 602 4166. Tauherenikau [WA].
S. diversa Broun, 1911b: 495 3366. Midhurst, near Mount Egmont [TK].
S. duplicata Broun, 1886: 888 1581. Parua, near Whangarei Harbour [ND]*.
S. electa Broun, 1914a: 91 3400. Erua, 2500 ft. [TO].
S. elevata Broun, 1886: 886 1578. Woodhill, near Helensville [AK].
S. elongata Broun, 1893a: 1423 2472. Karaka, near Drury [AK].
S. eminens Broun, 1895: 75 none. Tarukenga, near Rotorua & Mount Pirongia [BP,WO](Fig. 6).
S. excavata Broun, 1886: 884 1574. Paparoa district, south of Auckland [AK]*.
S. fagicola Broun, 1921a: 494 4004. Mount Dick, Lake Wakatipu, 1100 ft. [OL].
S. fasciculata Broun, 1921a: 497 4007. Glenhope, near Nelson [NN].
S. flavipes Broun, 1893a: 1422 2471. Maketu [AK].
S. fovealis Broun, 1886: 886 1577. Waitakerei Range [AK].
S. foveiventris Broun, 1921a: 492 4000. Hollyford, north of Lake Wakatipu [OL].
S. frontalis Raffray, 1893b: 23. NZ.
S. fulva Broun, 1893a: 1052 1881. Clevedon [AK].
S. fulvipennis Broun, 1915: 289 3705. Tairua, Auckland [CL].
S. furcata Broun, 1921a: 495 4005. Belgrove, near Nelson [NN].
S. fuscipalpis Broun, 1914b: 159 3523. Mount Hutt, near Methven [MC].
S. genalis Broun, 1881: 663 1157. Wellington? [WN]*.
S. grata Broun, 1912b: 628 3488. Picton [SD].
S. guinnessi Broun, 1911b: 502 3373. Erua [TO].
S. halli Broun, 1914b: 155 3518. Pudding Hill, near Methven [MC] (Figs. 7, 8, 101, 102).
S. hectorii Broun, 1917: 378 3829. Belgrove, near Nelson [NN].
S. hirtalis Broun, 1893a: 1050 1876. Near Howick [AK].
S. ignota Broun, 1921a: 495 4004. Routeburn, north of Lake Wakatipu [OL].
S. immota Broun, 1893a: 1422 2470. Maketu [AK].
S. indiscreta Broun, 1915: 288 3703. Howick, near Auckland [AK].
S. insignis 1893a: 1049 1875. Mokohinou Island [ND].
S. insolens Broun, 1893a: 1051 1879. Near Howick [AK].
S. insueta Broun, 1914b: 157 3521. Curiosity Gully, near Methven [MC].
S. laetula Broun, 1915: 291 3708. Hunua Range, near Clevedon [AK].
S. laminata Broun, 1893a: 1421 2469. Maketu, Hunua Range [AK]*.
S. laticeps Broun, 1911b: 490 3362. Greymouth [BR].
S. latistriata Broun, 1911b: 495 3367. Ligar’s Bush, Papakura [AK].
S. latula Broun, 1912b: 633 3493. Greymouth [BR].
S. lawsoni Broun, 1912b: 632 3492. Auckland [AK].
S. lineata Broun, 1893b: 175 none. Mount Pirongia [WO] (Fig. 9).

Faunistic Review on the Pselaphine Species Known from New Zealand243

S. lineiceps Broun, 1921a: 504 4018. Glenhope [NN].
S. longicollis Broun, 1911b: 498 3369. Mahuia, near Mount Ruapehu [TO].
S. longipennis Broun, 1911b: 504 3375. Erua, 2400 ft. [TO]*.
S. longipes Broun, 1915: 287 3702. Woodhill, Kaipara Railway [AK].
S. longula Broun, 1912b: 625 3485. Auckland [AK].
S. macronyx Broun, 1893a: 1418 2466. Maketu, Hunua Range [AK].
S. major Sharp, 1874: 507 246. NZ*. (Fig. 10)
S. mimica Broun, 1893a: 1419 2467. Hunua Range, Clevedon [AK].
S. minuscula Broun, 1921a: 497 4008. Heaven’s Gate, near Mount Earnslaw; Mount Alfred& Staircase, southern end of the Remarkables [CO,OL] (Fig. 11).
S. misella Sharp, 1874: 508 248. New ZealandS. monstrosa Reitter, 1880: 168. Type locality is not given (Greymouth implied). [BR?] (Figs.
12, 13).
S. monticola Broun, 1912a: 402 3195. Mount Ngauruhoe [TO]*.
S. nitida Broun, 1911b: 492 3364. Greymouth [BR].
S. notabilis Broun, 1880: 137 251. Tairua [CL]*.
S. occipitalis Broun, 1912b: 624 3484. Greymouth [BR].
S. opercularis Broun, 1915: 284 3698. Rakaia Gorge, near Methven [MC].
S. osculans Broun, 1886: 885 1576. Woodhill [AK].
S. pallidula Broun, 1912b: 626 3486. Greymouth [BR].
S. parallela Broun, 1893a: 1053 1883. Clevedon [AK].
S. parva Sharp, 1874: 508 249. NZ.
S. pertinax Broun, 1893b: 176 none. Ohaupo, Waikato, near Mr. Kusab’s saw-mill [WO].
S. planicula Broun, 1921a: 503 4017. Mount Te Aroha, Auckland, 2000 ft. [BP]*.
S. planipennis Broun, 1921a: 500 4013. Mount Oakden, Canterbury [MC].
S. posticalis Broun, 1915: 291 3707. Wairiri, Kaikoura [KA].
S. prisca Sharp, 1874: 507 247. New ZealandS. pulchra Broun, 1880: 137 250. Tairua [CL].
S. punctata Broun, 1893a: 1052 1880. Tairua [CL].
S. puncticeps Broun, 1911b: 489 3361. Tairua, Auckland [CL].
S. puncticollis Broun, 1911b: 499 3370. Timaru [SC].
S. punctulata Raffray, 1893b: 21. NZ.
S. rectipennis Broun, 1921a: 489 3997. Mount Alfred, north of Lake Wakatipu [OL].
S. rectipes Broun, 1893a: 1051 1878. Tairua [CL].
S. remixta Broun, 1921a: 502 4016. Mount Owen [NN].
S. robusta Broun, 1893a: 1420 2468. Hunua Range, near Drury [AK] (Fig. 14).
S. robustula Broun, 1917: 377 3828. Routeburn, north of Lake Wakatipu [OL].
S. rotundiceps Broun, 1915: 289 3704. Stratford, base of Mount Egmont [TK].
S. rufescens Broun, 1921a: 499 4010. Staircase, nearly ten miles north of Kingston [CO].
S. ruficeps Broun, 1893a: 1053 1882. Clevedon [AK].
S. rugifrons Broun, 1895: 73 none. Mount Pirongia [WO].
S. rustica Broun, 1915: 285 3700. Rakaia Gorge [MC].
S. setiventris Broun, 1915: 282 3696. Erua [TO].
S. sharpi Raffray, 1893b: 26. NZ.
S. sobrina Broun, 1893a: 1050 1877. Mokohinou Island [ND].
S. socia Broun, 1915: 281 3695. Pudding Hill, near Methven, 3500 ft. [MC].

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S. spinifer Broun, 1895: 75 none. Mount Pirongia [WO].
S. spiniventris Broun, 1912b: 627 3487. Picton [SD].
S. strialis Broun, 1921a: 489 3996. Moa Hill, Canterbury, 4000 ft. [MC].
S. striatifrons Broun, 1921a: 492 4001. Harris Saddle & Hollyford, etc., 1100–4200 ft.
[OL].
S. subcuneata Broun, 1921a: 488 3995. Moa Basin [MC].
S. sulcator Broun, 1886: 885 1575. Woodhill, on the Kaipara Railway, near Helensville[AK].
S. suturalis Broun, 1914b: 158 3522. Rakaia Gorge, near Methven [MC].
S. tenebrica Broun, 1921a: 487 3994. Moa Basin, Canterbury [MC].
S. tenuis Broun, 1886: 888 1582. Tairua [CL].
S. terricola Broun, 1886: 832 1480. Waitakerei Range [AK].
S. unicalis Broun, 1917: 376 3826. Scarcliff, near Mt. Algidus & Moa Basin [MC].
S. valida Broun, 1921a: 490 3998. Mount Owen & Woodhen Bend, near Glenhope [NN].
S. ventralis Broun, 1912b: 623 3483. Greymouth [BR].
Genus Stenosagola Broun (Figs. 15, 16)
Stenosagola Broun, 1921a: 504. Type species: Stenosagola planiocula Broun designated by Newton & Chandler, 1989:
18.

Remarks. According to the original description of this genus, Stenosagola is separated
from the genus Sagola by the following characters: body very narrow and slender; pronotum oviform; elytra subquadrate, each with a short and undivided dorsal sulcus; abdomen twice as longas elytra; antennae not longer than combined length of the head and pronotum. Two unidentified
species of this genus are in NZAC, and their unique characters, the slender and very narrowed
body, the oviform pronotum and some characters indicated above are distinct as shown in Figs.
15, 16.
List of species:
S. connata (Broun, 1911b): 503 3374. Erua [TO].
S. crassicornis (Broun, 1911b): 501 3372. Raurimu, Main Trunk Line, 1900 ft. [TO].
S. gracilis (Broun, 1893a): 1424 2473. Mount Pirongia [WO].
S. griseipila Broun, 1921a: 506 4021. Makatote, near Erua [TO].
S. oblongiceps Broun, 1921a: 505 4020. Glenhope [NN].
S. planiocula Broun, 1921a: 505 4019. Glenhope [NN].
Supertribe EUPLECTITAE
Tribe EUPLECTINI
Genus Euplectus Leach (Figs. 17, 18)
Euplectus Leach, 1817: 82. Type species: Euplectus reichenbachii Leach by monotypy.

Remarks. Euplectus is a taxonomically rich genus distributed mainly in the Holarctic,
though it is not so well-represented in Australia (1 species) and NZ. Some species in the following list should be transferred to other genera. Here we tranfer Euplectus iracundus to the genus
Euglyptus (see below). Euplectus sculpturatus (Fig. 19) was identified Plectomorphus by Kuschel(1990), though it should be placed in a group of true Euplectus and allied genera. Three species,
E. opacus, E. ovocollis and E. tuberigerus were combined with the genus Euplectopsis by

Faunistic Review on the Pselaphine Species Known from New Zealand

245

Kuschel (1990), but they are reclassified into this genus by Newton & Thayer (2003). There arean additional seven undescribed species contained in the NZAC.
List of species:
E. auripilus Broun, 1886: 945 1701. Near Howick [AK].
E. caviceps Broun, 1904: 48 none. Thompson’s Gorge [DN].
E. cephalotes Reitter, 1880: 171; preoccupied, not Motschulsky, 1845 (JPH). Type locality is
not given (Greymouth implied). [BR?].
E. frontalis Broun, 1880: 142 260. Tairua [CL].
E. incomptus Broun, 1884: 239; also 1886: 921 1652. Tairua [CL].
E. lepiphorus Broun, 1893a: 1056 1887. Clevedon [AK].
E. longulus Broun, 1880: 141 257. Tairua [CL].
E. opacus Sharp, 1874: 509 258. Auckland [AK]*.
E. ovicollis Broun, 1880: 143 262. Tairua [CL]*.
E. personatus Broun, 1893a: 1059 1894. Near Howick [AK].
E. scruposus Broun, 1893a: 105 1889. Near Clevedon [AK].
E. sculpturatus Broun, 1880: 142 259. Tairua [CL]*.
E. semiopacus Broun, 1895: 81 none. Maketu, Hunua Range [AK].
E. sulciceps Broun, 1904: 49 none. Thompson’s Gorge, Otago [DN].
E. tuberigerus Broun, 1882: 289; also 1886: 760 1348. Near Whangarei Harbour [ND]*.
E. unicus Broun, 1893a: 1060 1895. Stratford, near Mount Egmont [TK].
E. vacuus Broun, 1884: 239; also 1886: 921 1651. Paparoa (Howick). [AK].
E. verticalis Broun, 1893a: 1061 1897; preoccupied, not Reitter, 1884 (JPH). Near Howick[AK].
Genus Leptoplectus Casey (Fig. 20)
Leptoplectus Casey, 1908: 266. Type species: Euplectus pertenuis Casey designated by Lucas, 1920: 371.

Remarks. This genus is broadly distributed in Holarctic and Oriental regions. Klimaszewski et al. (1996) recorded this genus based on two undetermined species in the Lynfield collectionof NZAC (Fig. 20). The genus was not considered to be adventive species in that paper, but it isprobably introduced. This genus is very similar to Euplectus; however, it is separated from it byhaving the labrum with a large and arcuate emargination on the anteromedian area.
Tribe TRICHONYCHINI
Subtribe TRICHONYCHINA
Genus Macroplectus Raffray (Fig. 21)
Macroplectus Raffray, 1898: 255. Type species: Macroplectus calcaratus Raffray designated by Lucas, 1920: 389.

Remarks. The tribe Trichonychini was redefined by Chandler (2000) to include four subtribes (Trichonychina, Panaphantina, Bibloporina and Trimiina) including tribe-level synonymies. The genus Macroplectus was revised in the same work by Chandler (2000) and two
Gabata and a Vidamus species known from NZ were transferred to this genus. Macroplectus ischaracterized by the large and posterolaterally expanded head and the pronotum without a distinct median longitudinal sulcus; it is difficult to distinguish from the genus Vidamus ofPanaphantina.

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List of species:
M. bifoveata (Broun, 1921a): 514 (as Gabata) 4033. Routeburn [OL].
M. parallela (Broun, 1921a): 515 (as Gabata) 4034. Routeburn [OL].
M. spinipes (Broun, 1910): 21 (as Vidamus) 3048. Broken River [MC] (Fig. 21).
Subtribe PANAPHANTINA
Genus Adalmus Reitter
Adalmus Reitter, 1882c: 210. Type species: Adalmus velutinus Reitter by subsequent monotypy.

Remarks. No material of this genus is contained in the NZAC. This genus is similar to
Raffrayia (South Africa), Dalmina (South Africa) and Faronoma (Chile), all of which have thenormal head without an antennal tubercle and a simple prothorax. Adalmus is separated fromthese genera by the normal segments of antennal flagellum, the well demarkated antennal club
formed by three terminal segments and abdominal tergites 4–7 subequal.
List of species:
A. velutinus Reitter, 1885: 338. Greymouth [BR].
Genus Dalma Sharp (Fig. 22)
Dalma Sharp, 1874c: 504. Type species: Dalma pubescens Sharp by monotypy.

Remarks. This genus is distinguished from other members of the subtribe by its stout andflattened body. In some species, antennal segment 9 is swollen in the male. Three known species
and additional undescribed species are contained in the NZAC.
List of species:
D. gigantea Broun, 1914b: 164 3530. Mount Hutt, 2500 ft. & Scott’s Gully [MC] (Fig. 22).
D. graniceps Broun, 1921b: 605 4170. Near south of Lake Rotoiti, Nelson [BR].
D. pubescens Sharp, 1874: 505 244. Hokitika [as Hokatika] [WD].
Trichonyx sordidus L. W. Schaufuss, 1880a: 32; also 1880b: 508. NZ.
D. tuberculata Broun, 1880: 134 245. Near Parua, Whangarei Harbour [ND].
Genus Dalmisus Sharp (Fig. 23)
Dalmisus Sharp, 1886: 381. Type species: Dalmisus batrisodes Sharp by monotypy.

Remarks. No identified specimens of this genus are contained in the NZAC. It is characterized by the elongate first antennal segment (Sharp, 1886). Some specimens with these characteristics were collected by the authors in the South Island (Fig. 23).
List of species:
D. batrisodes Sharp, 1886: 382 1874. Greymouth [BR].
Genus Eleusomatus Raffray (Figs. 24, 25)
Eleusomatus Raffray, 1904a: 548. Type species: Euplectus allocephalus Broun designated by Lucas, 1920: 259.

Remarks. This is a special form of euplectine genus with a small, narrow, and more or lessflattened body with a posteriorly narrowed abdomen. We have examined Eleusomatus allocephalus, E. acuminatus, E. caudatus and E. rotundicollis that are contained in the NZAC.
List of species:
E. acuminatus (Broun, 1893): 1058 (as Euplectus) 1891; preoccupied, not Schaufuss, 1882

Faunistic Review on the Pselaphine Species Known from New Zealand

247(JPH). Clevedon [AK] (Fig. 24).
E. allocephalus (Broun, 1893a): 1057 (as Euplectus) 1890. Clevedon? [AK]*.
E. caudatus (Broun, 1893a): 1429 (as Euplectus) 2482. Maketu, Hunua Range & Clevedon[AK] (Fig. 25).
E. oculatus Broun, 1921a: 516 4036. Mistake, near Mount Algidus [MC].
E. ovicollis Broun, 1915: 305 3725. Bell Rock, near Methven [MC].
E. subcaecus Broun, 1921a: 516 4037. Mount Hope, Nelson [NN].
E. vidamoides Broun, 1921a: 516 4035. Routeburn [OL].
Genus Euglyptus Broun (Figs. 26, 27)
Euglyptus Broun, 1893a: 1411. Type species: Euglyptus elegans Broun by monotypy.
Microtyrus Broun, 1893a: 1413. Type species: Microtyrus punctatus Broun by monotypy.

Remarks. The genus Euglyptus is distinct in having the compact and thick body, long andslender legs, and slender antenna with three-segmented club. Euglyptus punctatus was described
as Microtyrus which was considered to be allied to Tyrus by Broun (1893a); however, it was synonymized and classified into Euplectini (not Trichonychini) by Raffray (1904a). Euplectus iracundus Broun should be placed in this genus based on the following characters that it shares withthe type species as snown in Fig. 27: body narrowed anteriorly, legs long and slender; antennae
long and slender, each with three-segmented club; pronotum with broad and deep longitudinalsulci and antebasal depressions.
List of species:
E. abnormis Broun, 1921b: 604 4169. Rotoiti & Mount Slips, Nelson [BR].
E. costifer Broun, 1893b: 166 none. Mount Pirongia [WO].
E. elegans Broun, 1893a: 1412 2460. Maketu, Hunua Range [AK].
E. foveicollis Broun, 1912a: 403 3196. Retaruke, near Erua [TO] (Fig. 26).
E. iracundus (Broun, 1893a): 1429 2481; new combination. Maketu, Hunua Range [AK]*(Fig. 27).
E. longiceps Broun, 1921b: 604 4168. Waimarino, Main Trunk Railway [TO].
E. longicornis Broun, 1912a: 404 3197. Raurimu [TO].
E. punctatus (Broun, 1893a): 1413 (as Microtyrus) 2461. Maketu, Hunua Range [AK]*.
E. sublaevis Broun, 1921b: 603 4167. Lake Rotoiti, Nelson [BR].
Genus Euplectopsis Raffray (Figs. 27–31, 103, 104)
Euplectopsis Raffray, 1890a: 101. Type species: Trichonyx microcephalus Reitter by monotypy.

Remarks. This genus is the most commonly collected euplectine genus in NZ. It is distinguished from the other allied genera by having the following combination of characters: pronotum ovoid or cordiform, with a complete median longitudinal sulcus or shallow depression;
prosternum with a median carina just before fore coxae; elytra each with four basal foveae. Eighteen identified species are contained in the NZAC.
List of species:
E. antennalis Broun, 1912a: 407 3205. Mount Ngauruhoe [TO].
E. antiquus (Broun, 1893a): 1061 (as Euplectus) 1898. Mokohinou Island [ND]*.
E. biimpressus Broun, 1912a: 410 3208. Raurimu [TO].
E. blandiatus Broun, 1915: 297 3715. Scott’s Gully & Pudding Hill, near Methven [MC].
E. brevicollis (Reitter, 1880): 170 (as Trichonyx) 3200. Type locality is not given (Grey-

248

S. Nomura and R. A. B. Leschen

mouth implied). [BR?].
E. bryocharis Broun, 1915: 296 3714. Bell Rock, near Methven [MC].
E. carinatus Broun, 1912a: 407 3204. Mount Te Aroha [BP].
E. clavatulus Broun, 1913: 199 3496. Greymouth [BR].
E. crassipes (Broun, 1884): 238 (as Euplectus) 1646; also 1886: 919. Howick (Auckland).
[AK].
E. crassulus Broun, 1921a: 506 4022. Glenhope [NN] (Fig. 28).
E. cuneiceps Broun, 1915: 296 3713. Epsom, base of Mount Eden [AK].
E. curvipennis Broun, 1914b: 164 3529. Greymouth [BR].
E. dorsalis Broun, 1915: 299 3717. Rakaia Gorge [MC].
E. duplex Broun, 1915: 295 3712. Epsom, base of Mount Eden [AK].
E. duplicatus Broun, 1913: 197 3495. Greymouth [BR].
E. elongellus Broun, 1915: 299 3718. Rakaia Gorge, near Methven [MC].
E. eminens (Broun, 1886): 945 (as Euplectus) 1700. Near Howick [AK].
E. eruensis Broun, 1912a: 408 3206. Erua [TO].
E. fastigiatus Broun, 1911a: 687 3377. Mount Pirongia [WO].
E. femoralis Broun, 1914b: 163 3528. Pudding Hill, near Methven [MC].
E. granulatus Broun, 1911a: 686 3376. Erua, 2400 ft. [TO].
E. heterarthrus Broun, 1912a: 409 3207. Erua, Raurimu, Makatote & Retaruke [TO] (Figs.
29, 30).
E. impressus Broun, 1915: 298 3716. Rakaia Gorge, near Methven & Mount Hutt [MC].
E. inscitus (Broun, 1893a): 1428 (as Euplectus) 2480. Maketu, Hunua Range [AK].
E. longicollis (Reitter, 1880): 168 (as Trichonyx) 3198. Type locality is not given (Greymouth implied) [BR?].
E. microcephalus (Reitter, 1880): 169 (as Trichonyx) 3199. Type locality is not given(Greymouth implied) [BR?].
E. mirificus (Broun, 1884): 239 (as Euplectus) 1653; also 1886: 922. Near Howick [AK].
E. modestus (Broun, 1895): 78 (as Euplectus) none. Invercargill [SL].
E. monticola (Broun, 1884): 239 (as Euplectus) 1648; also 1886: 920; preoccupied, not
Wollaston 1864 (JPH). Waitakerei Range [AK].
E. mucronellus Broun, 1911a: 689 3378. Southland [SO].
E. nitipennis Broun, 1915: 294 3711. Auckland Domain [AK].
E. obnisus Broun, 1884: 239 (as Euplectus) 1650; also 1886: 921. Waitakerei Range [AK].
E. ovithorax (Broun, 1884): 239 (as Euplectus) 1649; also 1886: 920. Woodhill, near
Kaipara Railway [AK].
E. parvulus (Broun, 1895): 77 (as Euplectus) none. Maketu, Hunua Range [AK].
E. patruelis (Broun, 1884): 238 (as Euplectus) 1647; also 1886: 919. Paparoa (Howick)[AK].
E. perpunctatus Broun, 1915: 293 3710. Papakura Bush, near Auckland [AK].
E. pusillus (Broun, 1895): 82 (as Euplectus) none; preoccupied, not Denny, 1825 (JPH).
Tarukenga, near Rotorua [BP].
E. rotundicollis (Reitter, 1880): 170 (as Trichonyx) 3201. Type locality is not given (Greymouth implied) [BR?].
E. sanguineus Broun, 1913: 196 3494. Greymouth [BR].
E. schizocnemis Broun, 1912a: 406 3203. Retaruke, near Erua [TO].
E. terrestris Broun, 1914b: 162 3527. Bell Rock, near Methven [MC] (Fig. 30).

Faunistic Review on the Pselaphine Species Known from New Zealand249

E. tibialis Broun, 1914b: 161 3526. Rose Hill, near Methven [MC].
E. trichoniformis (Reitter, 1880): 171 (as Trichonyx) 3202. Type locality is not given (Greymouth implied) [BR?].
E. tumidus Broun, 1911a: 690 3379. Erua, Waimarino & Makatote [TO] (Figs. 103, 104).
E. tumipes (Broun, 1895): 76 (as Euplectus) none. Hunua Range, Drury [AK].
Genus Kenocoelus Broun
Kenocoelus Broun, 1911a: 700. Type species: Kenocoelus dimorphus Broun by monotypy.

Remarks. Specimens of this genus are not contained in the NZAC. According to Broun(1911a) it is a very distinct genus bearing the elongate pronotum with shallow antebasal and ashallow longitudinal depressions and the elytra have humeral depressions. It is also characteristicby having diagnostic sexual dimorphism of the eye which is well developed in the male and reduced in the female.
List of species:
K. dimorphus Broun, 1911a: 700 3387. Greymouth [BR].
Genus Paraplectus Raffray (Fig. 32)
Philiopsis Raffray, 1898: 269. Type species: Paraplectus punctulatus Raffray designated by Lucas, 1920: 482.

Remarks. This genus has been known from Australia with five species. Kuschel (1990)
recorded this genus from Lynfield with an undetermined species. Two specimens of this species
are preserved in the Lynfield Collection of NZAC.
Genus Patreus Broun
Patreus Broun, 1904: 47. Type species: Patreus lewisi Broun by monotypy.

Remarks. The NZAC does not have specimens of this genus. In the original description by
Broun (1904), this genus was thought an allied group of Exeirarthra of Faronitae because of theslender body and the broadened basal segment of mid tarsus (note that in Exeirarthra the foretarsi have the basal segment is broadened). Raffray (1911) agreed with Broun, though this genus
was reclassified to Euplectitae by Newton & Chandler (1989).
List of species:
P. lewisi Broun, 1904: 47 none. Ida Valley [CO].
Genus Philiopsis Raffray (Fig. 32)
Philiopsis Raffray, 1893a: 471. Type species: Philiopsis exigua Raffray by monotypy.

Remarks. This is an Oriental genus that is also known from Japan. In NZAC, there are six
specimens of an undetermined Philiopsis species collected from Lynfield, Auckland (Fig. 32).
Klimaszewski et al. (1996) recorded this genus as an adventive taxon probably on the basis of thematerial in NZAC.
Genus Placodium Broun
Placodium Broun, 1893a: 1431. Type species: Placodium zenarthrum Broun by monotypy.

Remarks.

Specimens of this genus were unavailable for study and are not contained in the

250

S. Nomura and R. A. B. Leschen

NZAC. According to Raffray (1904a, 1908) Placodium is allied to Euglyptus (NZ) andPanaphantus (Europe and Africa) by sharing well developed antennal tubercles on the frons; but,
Placodium differs by having the antennal club formed only by the eleventh segment (formed by
the last three segments in Eugluptus and Panaphantus).
List of species:
P. zenarthrum Broun, 1893a: 1431 2484. Maketu, Hunua Range [AK].
Genus Plectomorphus Raffray (Figs. 33–36, 105, 106)
Plectomorphus Raffray, 1898: 265. Type species: Euplectus spinifer Broun by monotypy.

Remarks. This is an endemic NZ genus, is relatively common and is similar to Microplectus (Singapore and New Guinea) and Sagolonus (NZ) by having the normally developed antennal
tubercles; but, is separated from the former by the anteriorly narrowed head and from the latterby having a very strong median longitudinal sulcus on the pronotum. Four species are containedin the NZAC.
List of species:
P. anguliferus Broun, 1921a: 508 4024. Routeburn; Hollyford; Staircase, near Lake
Wakatipu; Moa & Mistake Basins; & Mount Algidus, Canterbury [CO, MC, OL] (Figs. 33,
34).
P. brevicornis Broun, 1913: 201 3498. Auckland [AK].
P. collinus Broun, 1921a: 510 4027. Staircase, southern part of the Remarkables, 3500 ft.
[CO].
P. egenus Broun, 1913: 202 3499. Tairua [CL].
P. insignis Broun, 1921a: 509 4026. Oakden, near Mount Algidus [MC].
P. laminifer Broun, 1915: 300 3719. Pudding Hill, about nine miles from Methven [MC].
P. longiceps Broun, 1913: 200 3497. Greymouth [BR].
P. longipes Broun, 1912a: 412 3213. Greymouth [BR].
P. munroi (Broun, 1893a): 1426 (as Euplectus) 2477. Clevedon, Hunua Range [AK].
P. optandus Broun, 1912a: 412 3212. Erua [TO].
P. rugiceps Broun, 1921a: 509 4025. Glenhope [NN].
P. scitiventris Broun, 1921a: 507 4023. Glenhope, Nelson [NN] (Figs. 105, 106).
P. spinifer (Broun, 1893a): 1426 (as Euplectus) 2476; preoccupied, not Casey, 1884 (JPH).
Maketu, Hunua Range [AK].
P. trisulcicollis (Broun, 1880): 140 255 (as Euplectus). Tairua [CL]* (Figs. 35, 36).
Genus Sagolonus Raffray (Figs. 37, 38)
Sagolonus Raffray, 1898: 265. Type species: Euplectus patronus Broun by monotypy.

Remarks. Two species of this genus, S. impressus and S. patronus, are contained in the
NZAC; however, S. patronus specimens (Fig. 38) are difficult to distinguish from Vidamus, while
S. impressus specimens (Fig. 37) are similar to Plectomorphus. Raffray (1904a) considered thisgenus as a member of a group that includes Plectomorphus, Whitea, Dalmisus and other genera
in having the large and nearly rectangular head, the pronotum with a short and shallow medianlongitudinal sulcus and the elytra each with a short longitudinal sulcus.
List of species:
S. arohaensis (Broun, 1895): 79 (as Euplectus) none. Mount Te Aroha [BP].

Faunistic Review on the Pselaphine Species Known from New Zealand251

S. impressus Broun, 1910: 21 3049. Broken River, Canterbury [MC] (Fig. 37).
S. patronus Broun, 1893a: 1060 (as Euplectus) 1896. Near Howick [AK] (Fig. 38).
Genus Vidamodes Broun
Vidamodes Broun, 1921b: 606. Type species: Vidamodes fulvus Broun by monotypy.

Remarks. Material of this genus is not contained in the NZAC. This genus is placed between Vidamus and Plectomorphus according to the original description by Broun (1921b) and isdistinct by having the pronotum with three antebasal foveae connected by transverse sulci and ashallow median longitudinal depression separated from basimedian fovea.
List of species:
V. furvus Broun, 1921b: 606 4171. Near Lake Rotoiti, Nelson [BR].
Genus Vidamus Raffray (Figs. 39–41, 107, 108)
Vidamus Raffray, 1898: 252. Type species: Euplectus incertus Reitter designated by Newton & Chandler, 1989: 27.

Remarks. This endemic genus is diverse and common in NZ, especially abundant in leaf
litter. It is characterized by the posteriorly broadened body, the large and posterolaterally swollen
head, and the moderately convex pronotum with V-shaped basal transverse sulci. Eight known
species and many unidentified specimens are contained in the NZAC.
List of species:
V. armiferus Broun, 1911a: 692 3381. Greymouth [BR].
V. brevitarsis (Broun, 1880): 143 261 (as Euplectus). Tairua [CL].
V. bryophilus Broun, 1914b: 166 3531. Rakaia Gorge, near Methven [MC].
V. calcaratus Broun, 1912a: 4113210. Makatote [TO] (Fig. 39).
V. cereus (Broun, 1884): 239 (as Euplectus) 1655; also 1886: 923. Waitakerei Range [AK].
V. clavipes Broun, 1915: 302 3721. Mount Hutt [MC].
V. congruous Broun, 1915: 301 3720. Scott’s Gully & Pudding Hill, near Methven [MC].
V. convexus (Sharp, 1874): 509 254 (as Euplectus). Auckland, [AK].
V. fossalis Broun, 1921a: 513 4032. Greymouth [BR].
V. gracilipes Broun, 1917: 379 3830. Mount Alfred, Heaven’s Gate & Mount Earnslaw, allnorth of Lake Wakatipu [OL].
V. incertus (Reitter, 1880): 172 (as Euplectus) 3211. Type locality is not given (Greymouth
implied) [BR?].
V. modestus Broun, 1913: 203 3500. Tairua [CL].
V. muscicola Broun, 1921a: 512 4030. Routeburn [OL].
V. nitidus Broun, 1921a: 513 4031. Mount Algidus, Canterbury [MC].
V. ovicollis Broun, 1921a: 511 4029. Routeburn [OL].
V. punctulatus Broun, 1915: 302 3722. Invercargill [SL].
V. simplex Broun, 1921a: 511 4028. Routeburn [OL].
V. sternalis Broun, 1913: 204 3501. Greymouth [BR].
V. trochanteralis Broun, 1911a: 691 3380. Erua & Waimarino, 2400–2700 ft. [TO] (Fig. 40,
41, 107, 108).
V. u-impressus (Broun, 1884): 239 (as Euplectus) 1654; also 1886: 923. Woodhill (Kaipara
Railway). [AK].
V. validus (Broun, 1893a): 1056 (as Euplectus) 1888. Waitakerei Range, Near Howick &252

S. Nomura and R. A. B. Leschen

near Clevedon [AK].
Genus Whitea Hutton (Figs. 42, 43)
Whitea Hutton, 1904: 179 (new name for Brounia Raffray, published by May 1904).
Brounia Raffray, 1898: 266 (preoccupied, not Sharp, 1878). Type species: Euplectus laevifrons Broun by monotypy.
Brouniella Raffray, 1904a: 584 (new name for Brounia Raffray, unnecessary, published September 1904).

Remarks. This is an endemic genus characterized by having a stout body, the head withprominent antennal tubercles, the very large and transverse pronotum, and the male fourth abdominal ventrite with a small posteromedian fringe. It is closely allied to the genus Plectomorphus. The monotypic species W. laevifrons is commonly collected in leaf litter.
List of species:
W. laevifrons (Broun, 1893a): 1425 (as Euplectus) 2475. Maketu, Hunua Range [AK] (Figs.
42, 43).
Genus Zelandius Raffray (Figs. 44, 45)
Zelandius Raffray, 1898: 272. Type species: Euplectus obscurus Broun by monotypy.

Remarks. This genus name given by Raffray (1898) has been misspelled elsewhere as
Zealandius. This genus is easily distinguished from the other euplectine genera by the small
body, the coarsely punctate head, and the elytra with prominent longitudinal carinae. Twelve
known species are preserved in the NZAC.
List of species:
Z. asper (Broun, 1880): 140 256 (as Euplectus). Tairua [CL].
Z. basalis (Broun, 1914b): 167 (as Zealandius) 3532. Curiosity Gully, near Methven [MC].
Z. brookesi (Broun, 1915): 303 (as Zealandius) 3723. Titirangi, Waitakerei Range [AK].
Z. clevedonensis (Broun, 1893a): 1058 (as Euplectus) 1892. Clevedon [AK]*.
Z. coxalis (Broun, 1893a): 1428 (as Euplectus) 2479. Maketu, Hunua Range [AK].
Z. fovealis Broun, 1913: 205 3502. Greymouth [BR].
Z. foveiceps (Broun, 1895): 80 (as Euplectus) none. Ligar’s Bush, Papakura [AK].
Z. fulgens (Broun, 1911a): 696 (as Zealandius) 3384. Greymouth [BR].
Z. illustris (Broun, 1911a): 695 (as Zealandius) 3383. Mount Pirongia [WO].
Z. moerens (Broun, 1893a): 1055 (as Euplectus) 1886. Tairua [CL].
Z. obscurus (Broun, 1893a): 1427 (as Euplectus) 2478. Maketu, Hunua Range [AK]* (Fig.
45).
Z. raffrayi (Broun, 1911a).?: 693 (as Zealandius) 3382. Erua, 2400 ft. [TO] (Fig. 44).
Z. sandageri (Broun, 1893a).?: 1059 (as Euplectus) 1893. Mokohinou Island [ND]*.
Z. spinifer (Broun, 1914b): 167 (as Zealandius) 3533. Pudding Hill, near Methven [MC].
Z. tuberalis (Broun, 1915): 304 (as Zealandius) 3724. Mount Hutt & Pudding Hill [MC].
Z. usitatus (Broun, 1910): 22 (as Zealandius) 3050. Broken River [MC].
Subtribe TRIMIINA
Genus Alloplectus Broun (Fig. 46)
Alloplectus Broun, 1911a: 697. Type species: Euplectus claviger Broun by original description.

Remarks. The trimiine genus Alloplectus is endemic to NZ. It is similar to some Nearctic
genera, for example Actiastes; however, it is distinct by having a small body, a swollen secondFaunistic Review on the Pselaphine Species Known from New Zealand

253

antennal segment, the pronotum without longitudinal sulcus or depression, and the elytron with
two large basal foveae. One specimen of A. claviger is contained in the NZAC (Fig. 46).
List of species:
A. claviger (Broun, 1893a): 1430 (as Euplectus) 2483. Maketu, Hunua Range [AK]* (Fig.
46).
A. picipennis Broun, 1911a: 698 3385. Howick, near Auckland [AK].
A. subcaecus Broun, 1911a: 699 3386. Retaruke Forest, near Erua [TO].
Tribe TROGASTRINI
Subtribe TROGASTRINA
Genus Neosampa Broun (Figs. 47, 48)
Neosampa Broun, 1921a: 518. Type species: Neosampa granulata Broun by monotypy.

Remarks. This is a distinctive myrmecophilous genus in NZ, and is similar to Asian
myrmecophilous batrisines in the habitus. It is distinctive by having long and slender antenna,
thickened and transverse head with eyes located posteriorly. Several specimens of N. granulata
are contained in NZAC.
List of species:
N. granulata Broun, 1921a: 519 4039. Boulderstone Cleek [sic]. and Scarcliff, near Mount
Algidus [MC] (Figs. 47, 48).
Supertribe GONIACERITAE
Tribe BRACHYGLUTINI
Subtribe BRACHYGLUTINA
Genus Anabaxis Raffray (Figs. 49, 50)
Anabaxis Raffray, 1908: 252. Type species: Bryaxis lunatica King designated by Newton & Chandler, 1989: 42.

Remarks. This genus is distributed in NZ and Australia. It is middle-sized brachyglytine
genus characterized by the visible and demarcated third abdominal ventrite between hind coxae
and the elytron with three basal foveae connected by a transverse sulcus according to the originaldescription. Kuschel (1990) recorded two species shown below from Lynfield, Auckland, andsynonymized A. minor Broun with A. electrica (King). He considered this species to be an introduced species from Australia. Two species collected from Lynfield are contained in the NZAC.
List of species:
A. electrica (King, 1863): 48. Swan River, Tasmania (Australia)* (Fig. 49).
Anabaxis minor Broun, 1921a: 519 4040. Mount Hope, near Nelson [NN].
A. foveolata (Broun, 1880): 143 263 (as Euplectus). Tairua [CL]* (Fig. 50).
Bryaxis euplectoides Broun, 1893a: 1045 1869. Near Howick [AK].
Genus Eupines King (Figs. 51–61)
Eupines King, 1866: 309. Type species: Bryaxis clavatula King designated by Jeannel, 1952b: 84.
Byraxis Reitter, 1880a: 166 (subgenus). Type species: Bryaxis monstrosa Reitter by monotypy.

Remarks. The genus Eupines is broadly distributed in the Oriental and Australian Regions,
including Japan. It is highly diverse in NZ with 44 known species. In many species, the male antenna is modified. Twenty-five described species are contained in the NZAC.

254

S. Nomura and R. A. B. LeschenThis genus is classified to two subgenera, nominotypical subgenus and Byraxis. The subgenus Byraxis is characterized by the ten-segmented antenna usually with sexual character in themale; the female has slender antenna with eleven segments. In the nominotypical subgenus, theantenna is slender, normally consisting of eleven-segments in both sexes.
Subgenus Eupines s. str.
List of species:
E. (E.) calcarata (Broun, 1886): 831 (as Bryaxis) 1479. Tuakau, Waikato [AK].
E. (E.) grata (Sharp, 1874): 500 240 (as Bryaxis). NZ.
E. (E.) micans (Sharp, 1874): 497 231 (as Bryaxis). NZ. (Figs. 53, 54).
E. (E.) nasuta (Broun, 1880): 132 242 (as Bryaxis). Tairua [CL] (Fig. 61).
Bryaxis naso C. Schaufuss, 1888: 29 (new name for nasuta Broun, 1880, unnecessary).
E. (E.) nesobia Broun, 1914a: 92 3401. Great Barrier Island [CL].
E. (E.) piciceps (Broun, 1880): 127 233 (as Bryaxis). Tairua [CL].
E. (E.?) platynota (Broun, 1893a): 1338 (as Bryaxis) 2346. Mangawhare, Northern Wairoa[AK].
E. (E.) simplex Broun, 1913: 206 3503. Tairua, Auckland [CL].
E. (E.) sternalis (Broun, 1893b): 171 (as Bryaxis) none. Ligar’s Bush, Papakura & Mount
Pirongia, Waikato [WO] (Figs. 59, 60).
Subgenus Byraxis Reitter
Byraxis Reitter, 1880: 2. Type species: Byraxis monstrosa Reitter by monotypy.

List of species:
E. (B.) acceptus (Broun, 1923): 682 (as Byraxis) 4277. Rangiriri, Waikato [WO].
E. (B.) allocera (Broun, 1893b): 172 (as Bryaxis) none. Mount Pirongia [WO].
E. (B.) anisarthra (Broun, 1914b): 169 (as Byraxis) 3535. Broken River [MC].
E. (B.) bisulcifrons (Broun, 1914b): 170 (as Byraxis) 3536. Mount Hutt, 3000 ft. [MC].
E. (B.) clemens Broun, 1921b: 607 4172. Glenhope; near Lake Rotoiti, Nelson [BR, NN].
E. (B.) conspicua (Broun, 1893a): 1415 (as Bryaxis) 2464. Hunua Range, Maketu [AK].
E. (B.) costata (Broun, 1893a): 1416 (as Byraxis) 2465. Riccarton Bush, Christchurch[MC].
E. (B.) crassicornis (Broun, 1880): 129 237 (as Bryaxis; preoccupied, not Motschulsky,
1851 (JPH)). Tairua [CL].
E. (B.) decens (Broun, 1893a): 1046 (as Bryaxis) 1870. Paparoa, Near Howick [AK].
E. (B.) deformis (Sharp, 1874): 499 238 (as Bryaxis). New ZealandE. (B.) dispar (Sharp, 1874): 498 234 (as Bryaxis). Auckland, New Zealand [AK].
Bryaxis ovalipennis Schaufuss, L.W., 1880a: 25 (also 1880b: 501). Nov. Zealandia
E. (B.) diversa (Broun, 1893b): 174 (as Bryaxis) none; preoccupied, not Raffray 1887
(JPH), Sharp 1887 (JPH). Hunua Range, Drury [AK].
E. (B.) forficulida (Broun, 1890): 232 (as Bryaxis); also 1893a: 1047 1872. Neighborhoodof Clevedon, southern Wairoa [AK].
E. (B.) foveatissima (Broun, 1890): 233 (as Bryaxis); also 1893a: 1047 1873. Neighborhoodof Clevedon, southern Wairoa [AK].
E. (B.) fraudulenta (Broun, 1886): 944 (as Bryaxis) 1699. Near Howick [AK].
E. (B.) glabrata (Broun, 1886): 830 (as Bryaxis) 1476. Woodhill, near the Kaipara railway

Faunistic Review on the Pselaphine Species Known from New Zealand

255[AK]*.
E. (B.) halli (Broun, 1921b): 608 (as Byraxis) 4173. Mount St. Arnaud, Nelson [BR].
E. (B.) hectori (Broun, 1895): 73 (as Bryaxis) none. Tarukenga, near Rotorua [BP].
E. (B.) ignotus (Broun, 1881): 661 (as Bryaxis) 1155. near Whangarei Harbour [ND].
E. (B.) illustris (Broun, 1914b): 168 (as Byraxis) 3534. Broken River, Canterbury [MC].
E. (B.) impar (Sharp, 1874): 500 (as Bryaxis) 239. Auckland [AK]* (Figs. 51, 52).
E. (B.) impressifrons (Broun, 1880): 128 235 (as Bryaxis). Tairua [CL].
E. (B.) lewisi Broun, 1910: 24 3052. Broken River, Canterbury [MC].
E. (B.) longiceps Raffray, 1904b: 124. NZ.
E. (B.) monstrosa (Reitter, 1880): 167 (as Byraxis) 3214. Not given (Greymouth implied).
[BR?] (Figs. 55, 56).
E. (B.) munda (Broun, 1880): 129 (as Bryaxis) 236; preoccupied, not Sharp, 1874 (JPH).
Tairua [CL].
Bryaxis mundula C. Schaufuss, 1888: 29 (6907; new name for mundus Broun, 1880).
Bryaxis mundulus Broun, 1893: 1417 (new name for mundus Broun, 1880); preoccupied,
not Schaufuss, 1888 (JPH)).
E. (B.) munroi (Broun, 1890): 231 (as Bryaxis) ; also 1893a: 1046 1871. Neighborhood ofClevedon, southern Wairoa [AK].
E. (B.) nemoralis (Broun, 1886): 831 (as Bryaxis) 1477. Woodhill [AK].
E. (B.) paganus (Broun, 1881): 660 (as Bryaxis) 1154. Near Whangarei Harbour [ND].
E. platyarthra (Broun, 1893): 1417 (as Bryaxis) (new name for Bryaxis clavata Broun,
1880). [AK,CL].
Bryaxis clavatus Broun, 1880: 126 232; preoccupied, not Motschulsky, 1851 (JPH), Peyron, 1858 (JPH), Brendel 1865 (JPH). Tairua.
E. (B.) rhyssarthra (Broun, 1912a): 413 (as Byraxis) 3215. Mount Pirongia [WO].
E. (B.) rudicornis Broun, 1882: 288 1347; also 1886: 759. Wellington? [WN] (Figs. 57,
58).
E. (B.) sanguinea (Broun, 1880): 132 243 (as Bryaxis; not preoccupied by Leach, 1817 (unavailable), Linnaeus, 1790 (unavailable?)). Tairua [CL].
Bryaxis vae Schaufuss, C., 1888: 34 (new name for sanguinea Broun, 1880 (unnecessary)).
Bryaxis fulvitarsis Broun, 1893: 1418 (new name for sanguinea Broun, 1880 (unnecessary)).
E. (B.) setifera (Broun, 1893b): 173 (as Bryaxis) none. Mount Pirongia [WO].
E. (B.) sylvicola (Broun, 1884): 238 (as Bryaxis); also 1886: 919 1645. Paparoa bush, Near
Howick [AK].
Genus Eupinogitus Broun (Figs. 62, 109, 110)
Eupinogitus Broun, 1921: 517. Type species: Eupinogitus sulcipennis Broun by monotypy.

Remarks. This monotypic endemic genus resembles the Holoarctic genus Rybaxis in having the pronotum with strong antebasal transverse sulci connecting the antebasal foveae; but, isdifferent in the frons that lacks the anteromedian fovea (usually present in Rybaxis). Many specimens are contained in the NZAC.
List of species:
E. sulcipennis Broun, 1921a: 518 4038. Staircase, southern end of the Remarkables,
3500 ft. [CO] (Figs. 62, 109, 110).

256

S. Nomura and R. A. B. LeschenEupinogitus picescens Broun, 1921a: 518 4038, var.. Staircase, southern end of the Remarkables, 3500 ft. [implied].
Genus Eupinolus Oke
Eupinolus Oke, 1928: 11. Type species: Eupinolus leana Raffray designated by Lucas, 1920.

Remarks. This genus is common in NZ and shared with Australia. It is closely allied toEupines by having a small and compact body, and the smooth surface of the frons lacking a median fovea: it is diagnostic by having the pronotum with three subequal antebasal foveae and theelytron with two basal foveae or depressions and an adsutural sulcus. Its two species were originally described as Bryaxis species, but were later moved to Eupines by Chandler (2000). Bothspecies are contained in the NZAC.
List of species:
E. altulus (Broun, 1880): 131 241 (as Bryaxis). Near Whangarei Harbour [ND]*.
E. punctatus (Broun, 1886): 831 (as Bryaxis) 1478. Paparoa, near Howick [AK]*.
Genus Gastrobothrus Broun (Figs. 63, 64)
Gastrobothrus Broun, 1882: 287; 1886: 758. Type species: Bryaxis abdominalis Broun by monotypy.

Remarks. This is a relatively large-bodied and an endemic brachyglutine genus containing
two species. It is similar to Physobryaxis by having the very robust body and the strongly narrowed elytra at the base; but, is distinct in the robust and subglobose pronotum that lacks foveae,
depressions nor sulci. The two species are contained in the NZAC, and a special type of large excavation is present on the abdominal ventrites 5–7 in the male of G. abdominalis (Fig. 63).
List of species:
G. abdominalis (Broun, 1880): 125 230 (as Bryaxis; preoccupied, not Aubé, 1833 (JPH);
not now congeneric). Tairua [CL]* (Fig. 63).
G. sharpi (Broun, 1880): 124 229 (as Bryaxis). Tairua [CL]* (Fig. 64).
Genus Physobryaxis Hetschko (Figs. 65, 66)
Physobryaxis Hetschko, 1913: 182 (new name for Physa Raffray). Type species: Bryaxis inflata Sharp by monotypy.
Physa Raffray, 1890a: 122 (preoccupied, not Draparnaud, 1801).
Achiraffraya Navas, 1925: 29 (new name for Physa, unnecessary).

Remarks. The endemic genus Physobryaxis was previously synonymized with Gastrobothrus by Newton & Chandler (1989) though we separate it as an independent genus after Kuschel(1990) and the later authors due to the presence of a median antebasal fovea on the pronotum
connected with the lateral foveae by a pair of antebasal transverse sulci, a character not present in
Gastrobothrus.
List of species:
P. inflata (Sharp, 1874): 497 (as Bryaxis/Physobryaxis) 228. Auckland [AK]* (Figs. 65,
66).
Genus Simkinion Park et Pearce (Fig. 68)
Simkinion Park et Pearce, 1962: 251. Type species: Simkinion prelaticum Park et Pearce by original designation.

Remarks.

This is a very distinct and endemic brachyglutine genus bearing a robust body,Faunistic Review on the Pselaphine Species Known from New Zealand257

and having long antenna with strongly thickened terminal segments, a subglobose prothorax
without distinct sulcus or fovea, and long and slender legs. The systematic position is unknown.
Type material of S. prelaticum are deposited in the NZAC (a female paratype is shown in Fig.
68).
List of species:
S. bimanum Park et Pearce, 1962: 255. Dobbie’s Parl, Whangarei, North Island, New Zealand[ND].
S. prelaticum Park et Pearce, 1962: 252. Near Russell, North Island, New Zealand [ND] (female paratype is shown in Fig. 68).
Genus Startes Broun
Startes Broun, 1886: 829. Type species: Startes sculptulata Broun by monotypy.

Remarks. Two species of this Australasian genus are known from NZ (Chandler, 2000).
This genus is similar to Anabaxis by sharing with it the robust and subglobose prothorax without
antebasal transverse sulci, trifoveate elytra, and the fourth abdominal tergite with a pair of shortbasimedian carinae. Startes differs from Anabaxis by the smooth frons that lacks a pair of postantennal pits on the antennal bases (present in Anabaxis, see Chandler, 2000). The NZAC does notcontain specimens of this species.
List of species:
S. foveata Broun, 1893b: 170 none. Hunua Range [AK].
S. sculpturata Broun, 1886: 830 1475. Waitakerei Range [AK]*.
Supertribe PSELAPHITAE
Tribe PSELAPHINI
Genus Pselaphogenius Reitter (Figs. 69–72, 111, 112)
Pselaphogenius Reitter, 1910: 155. Type species: Pselaphus quadricostatus Reitter designated by Jeannel, 1950: 389.
Pselaphodinus Jeannel, 1950: 389. Type species: Pselaphus longipalpis Kiesenwetter designated by original description.

Remarks. The genus Pselaphogenius is known from Europe, East Asia (including Japan),
Australia, and NZ. It is defined by having an elongate head without scales on the postgenae, thepronotum without antebasal sulcus, and the elytron with two basal foveae and a broad longitudinal carina between the basal foveae. Chandler (2000) commented that approximately half of thespecies of “Pselaphus” from New Zealand should be placed in this genus, and three species
shown below were transferred from Pselaphus in the on-line database of Newton & Thayer(2003). The species occurring in NZ are different from typical Pselaphogenius species known
from the Palearctic Region by the elytron with only inner basal fovea and indistinct longitudinalcarina. This character is shared not with Pselaphogenius, but with the European genus Pselaphostomus Reitter (Pselaphopsis Jeannel).
List of species:
P. citimus (Broun, 1893a): 1043 (as Pselaphus) 1867. Howick [AK]* (Figs. 69, 70).
P. delicatus (Broun, 1886): 943 (as Pselaphus) 1697. Near Howick [AK]* (Figs. 71, 72,
111, 112).
P. ventralis (Broun, 1895): 72 (as Pselaphus) none. Mount Pirongia [WO].

258

S. Nomura and R. A. B. Leschen

Genus Pselaphophus Raffray (Figs. 73, 74)
Pselaphophus Raffray, 1890a: 139. Type species: Bryaxis atriventris Westwood designated by Lucas, 1920: 544.

Remarks. The genus Pselaphophus is known from Australia, and the type species, P.
atriventris was recorded from NZ as an introduced species (Martin, 1983, Klimaszewski et al.,
1996, Chandler, 2000). It is easily distinguished from the allied genera by having the non-pedunculate fouth segment of the maxillary palpus. Pselaphophus atriventris in NZ is concerned to be
introduced after Chandler (2000). More than forty specimens of this species are preserved in
NZAC, some of which were collected from Lynfield, Auckland.
List of species:
P. atriventris (Westwood, 1856): 270. Melbourne, Nova Hollandia (Australia) [AK] (Figs. 73,
74).
Genus Pselaphotheseus Park (Figs. 81–84)
Pselaphotheseus Park, 1964: 391. Type species: Pselaphotheseus hippolytae Park by original designation.

Remarks. This is a highly specialized genus of Pselaphini presently known only from subantarctic Campbell Island with two species. It is distinct by having a short maxillary palpus witha simple fusiform fourth segment, and an ovoid head with a large vertexal concavity and posterolaterally expanded postgenae. Type material of both species is deposited in the NZAC.
List of species:
P. hippolytae Park, 1964: 392. Campbell Is. [CA] (Figs, 81, 82).
P. ihupuku Carlton et Leschen, 2001: 387. Campbell Is. [CA] (Figs. 83, 84).
Genus Pselaphus Herbst (Figs. 75–80)
Pselaphus Herbst, 1792: 106. Type species: Pselaphus heisei Herbst designated by Shuckard, 1839.

Remarks. The genus Pselaphus is known from all over the world, is highly variable andshould not be recognized as a monophyletic group. Even the NZ species known as Pselaphus
contains various forms that are most probably separate genera. For example, P. pauper is similar
to typical Pselaphaulax known from the Palearctic Region in having the pronotum with antebasal
sulcus. The Pselaphus species require careful and detailed study for correct classification andidentification.
List of species:
P. caecus Broun, 1886: 943 1696. Near Howick [AK].
P. cavelli Broun, 1893a: 1414 2462. Capleston [BR].
P. cavidorsis Broun, 1923: 681 4276. Oruru, North Auckland [ND].
P. dulcis Broun, 1881: 660 1153. Pataua, north of Whangarei Harbour [ND]* (Fig. 75).
P. fuscopilus Broun, 1886: 944 1698. Near Howick [AK].
P. meliusculus Broun, 1893: 1044 1868. Mount Maungatua, Otago [DN].
P. oviceps Broun, 1917: 380 3832. Routeburn & Hollyford, north of Lake Wakatipu,
1000–3500 ft. [OL].
P. pauper Sharp, 1874: 492 226. Hokitika [WD]* (Fig. 76).
P. pilifrons Broun, 1914a: 93 3402. Tisbury, Invercargill [SL].
P. pilistriatus Broun, 1880: 123 227. Near Whangarei Harbour [ND]..
P. sulcicollis Broun, 1893a: 1415 2463; preoccupied, not Reichenbach, 1816 (JPH). MountFaunistic Review on the Pselaphine Species Known from New Zealand

259

Pirongia [WO].
P. trifoveatus Broun, 1914b: 172 3538. McClennan’s Bush & Rakaia Gorge, near Methven[MC] (Fig. 77).
P. urquharti Broun, 1917: 379 3831. Oakden, near Mount Algidus [MC].
Tribe TYRINI
Subtribe TYRINA
Genus Agatyrus Broun (Figs. 85, 86)
Agatyrus Broun, 1917: 382. Type species: Agatyrus fulvihirtus Broun by monotypy.

Remarks. This endemic and monotypic genus is characterzed by the robust and stout bodyand legs. In the male of A. fulvihirtus, the metaventrite bears a transverse projection just in frontof the hind coxae, and the abdomen is strongly excavated on the ventromedian side throughout its
entire length. More than ten specimens of this species are deposited in NZAC.
List of species:
A. fulvihirtus Broun, 1917: 383 3834. Gordon’s Knob, Nelson [NN] (Figs. 85, 86).
Genus Gerallus Sharp (Fig. 87)
Gerallus Sharp, 1874: 493. Type species: Gerallus nanus Sharp by monotypy.

This genus is known from Australia and New Guinea (Chandler 2002) and was recorded
from NZ by Kuschel (1990) from Lynfield, Auckland, represented by the introduced species, G.
punctipennis Schaufuss. Several specimens were examined in the NZAC.
List of species:
G. punctipennis L. W. Schaufuss, 1880b: 509. Eastern Creek et Nov. S. Wales, Nov. Holl.
(Australia) [AK]* (Fig. 87).
Genus Hamotulus Schaufuss (Figs. 88–92)
Hamotulus Schaufuss, 1887: 108. Type species: Bryaxis chamaeleon Schaufuss by monotypy.
Tychotyrus Broun, 1893b: 168. Type species: Tychotyrus sternalis Broun designated by Newton & Chandler, 1989.

Remarks. The genus Hamotulus is known from NZ and Australia. It is characteristic byhaving the pronotum smooth and shiny with three large foveae connected by antebasal transverse
sulci. In many NZ species, the male has a short spine near the apex of hind tibia. It had beenplaced in the subtribe Hamotina (preoccupied by Somatipionina Jeannel, 1949 after Chandler(2000)) in Newton & Chandler (1989); however, it was moved to Tyrina by Chandler (2000)
probably because it has the simple fourth segment in the maxillary palpus (longitudinally carinated or sulcate in Somatipionina). Eight described species are contained in the NZAC.
List of species:
H. angulipes Broun, 1914b: 171 3537. Curiosity Gully, Rakaia Gorge, near Methven [MC].
H. armatus (Broun, 1893a): 1043 (as Tyrus) 1866. Near Howick [AK] (Fig. 88).
H. cornutus Broun, 1915: 307 3728. Mount Hutt [MC] (Fig. 89).
H. curvipes (Broun, 1893b): 170 (as Tychotyrus) none. Hunua Range [AK].
H. frontalis Broun, 1914a: 94 3403. Broken River, Canterbury [MC].
H. fuscipalpis Broun, 1915: 306 3727. Mount Hutt [MC].
H. mutandus (Sharp, 1874): 487 225. Auckland, New Zealand [AK]*.
H. robustus Broun, 1915: 305 3726. Rose Hill, near Methven [MC] (Figs. 90, 91).

260

S. Nomura and R. A. B. Leschen

H. spinipes (Broun, 1893a): 1411 (as Tyrus) 2459. Riccarton Bush, Christchurch [MC](Fig. 92).
H. sternalis (Broun, 1893b): 168 (as Tychotyrus) none. Hunua Range, three different localities [unnamed]. [AK].
Genus Phormiobius Broun (Figs. 93, 94)
Phormiobius Broun, 1917: 381. Type species: Phormiobius halli Broun by monotypy.

Remarks. This monotypic genus is known only from NZ. It is morphologically distinct andcan be differentiated from other tyrine genera by the strongly shortened and basally narrowed
elytra and the very large abdomen with the large sixth segment. Many specimens are contained in
the NZAC.
List of species:
P. halli Broun, 1917: 382 3833. Belgrove, near Nelson [NN] (Figs. 93, 94).
Genus Plesiotyrus Broun
Plesiotyrus Broun, 1914a: 94. Type species: Tyrus crassipes Broun by original designation.

Remarks. The NZAC does not contain specimens of this monotypic genus, but according
to the original description (Broun, 1914a), it is closely allied to Tyrogetus, but separated from itby the reduced maxillary palpus, the larger eyes and the thickened legs, and different shape of theantenna.
List of species:
P. crassipes (Broun, 1893a): 1337 (as Tyrus) 2345. Moeraki [DN].
Genus Tyrogetus Broun (Fig. 95)
Tyrogetus Broun, 1893b: 165. Type species: Tyrogetus optandus Broun by monotypy.

Remarks. This endemic genus is characterized by the large and elongate maxillary palpus,
and the predominantly large fourth abdominal tergite with a large and setose basimedian depression. Several specimens of T. palpalis are contained in the NZAC.
List of species:
T. optandus Broun, 1893b: 166 none. Mount Pirongia [WO].
T. palpalis Broun, 1910: 23 3051. Waimarino [TO] (Fig. 95).
Genus Zeatyrus Sharp (Fig. 96)
Zeatyrus Sharp, 1881: 48. Type species: Zeatyrus lawsoni Sharp by monotypy.
Palmipalpus Broun, 1881: 662. Type species: Parmipalpus montivagus Broun by monotypy.

Remarks. This endemic and monotypic genus is very distinctive and can be distinguishedfrom other tyrine genera by the presence of a large and thickened maxillary palpus with a strongly swollen fourth segment, the subglobose and smooth pronotum without fovea or sulcus, theconvex elytron with a basal fovea, and the long and stout legs. Several specimens of the typespecies are contained in the NZAC.
List of species:
Z. lawsoni Sharp, 1881: 48. Auckland [AK]* (Fig. 96).
Parmipalpus montivagus Broun, 1881: 662 1156. Mount Manaia, Whangarei Harbour

Faunistic Review on the Pselaphine Species Known from New Zealand

261

[ND].
Broun’s “Syntypes” in the DEI
One of us (Nomura) visited DEI in May 2005 and examined pselaphine material from NZ
that were labeled as syntypes. Type material studied by Broun are deposited in the Natural History Museun, London (NHML) as well as NZAC, but some may have been traded or exchanged
with other collections (like the DEI) and care is required for correct lectotypification. True Brountypes can be recognized by a label written in Broun’s handwriting with the type locality as a single word followed by a period (some specimens may have his species number also followed by aperiod) and easily matched with the locality or localities listed in the original descriptions. The“types” in the DEI have labels that are not in Broun’s handwriting and have been carelessly relabled and are no doubt part of an exchange or a gift. The red “syntypus” label pinned to thespecimens were probably placed there by a DEI worker, because these labels are the same styleas those pinned to other type material from different geographic areas contained in the DEI. It is
unclear if the DEI specimens were used by Broun for his original descriptions, though they
match type localities cited in his descriptions.
The DEI specimens are listed below and illustrations of the labels and specimens are provided in Figs. 97–112. Specimens were previously listed by Gaedike (1984) as type specimens. In
the list, Bryaxis sanguineus was not examined and the data we list were cited from Gaedike(1984).
Bryaxis sanguineus Broun, 1880 (Eupines in the present study): 3 syntypes, Tairua (not examined).
Eupinogitus sulcipennis Broun, 1921: 3 syntypes (cotypes) Staircase, 3,500 ft., Otago (Figs.
109, 110).
Euplectopsis tumidus Broun, 1911: 3 syntypes (cotypes), Erua, King Country (Figs.
103–104).
Plectomorphus scitiventris Broun, 1921: 3 syntypes, Glenhope, Nelson (Figs. 105, 106).
Pselaphus delicatus Broun, 1886 (Pselaphogenius in the present study): 4 syntypes (cotypes), Howick (Figs. 111, 112).
Sagola bilobata Broun, 1921: 3 syntypes (cotypes), Mt. Dick, Otago (Figs. 97, 98).
Sagola dickensis Broun, 1917: 1 syntype (paratype), Mt. Dick, Otago (Figs. 99–100).
Sagola halli Broun, 1914: 3 syntypes (cotypes), Methven, Canterbury (Figs. 101–102).
Vidamus trochanteralis Broun, 1911: 2 syntypes (cotypes), Erua, King Country (Figs. 107,
108).
Acknowledgements
This work was supported in part by the National Science Museum project “Cooperation with
the Museums in Asia and the Pacific Rim for Collection Building and Natural History Studies”
that funded travel for S. Nomura to study specimens in the NZAC and to undertake field work
and R. Leschen to work in Japan during November–December 2005. Additional support to R.
Leschen was provided by Foundation for Research, Science and Technology (Contract Number
C09401). We thank Dr. Shun-Ichi Ueno and comments on the manuscript and Dr. Trevor K.
Crosby for his kind arrangement for this cooperative project.

262

S. Nomura and R. A. B. Leschen

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Park, O., & E. J. Pearce, 1962. Simkinion, a new genus of pselaphid beetles from New Zealand. Proceedings of the Biological Society of Washington, 75: 251–258.
Park, O., 1964. Insects of Campbell Island, Coleoptera: Pselaphidae. Pacific Insects Monograph, 7: 391–394.
Raffray, A., 1890. Étude sur les Psélaphides. V. Tableaux synoptiques. —Notes et synonymie. Revue d’Entomologie,
Caen, 9: 81–172.
Raffray, A., 1893a. Révision des Psélaphides de Sumatra. Annales de la Société Entomologique de France, 61: 463–504.
Raffray, A., 1893b. Essai monographique sur la tribu des Faronini. Revue d’Entomologie, 12: 259–260.
Raffray, A., 1898. Notes sur les Psélaphides. Révision générique de la tribu des Euplectini. Descriptions d’espèces nouvelles. Revue d’Entomologie, Caen, 17: 198–273.
Raffray, A., 1904a. Genera et catalogue des Psélaphides. Annales de la Société Entomologique de France, 72: 484–604.
Raffray, A., 1904b. Genera et catalogue des Psélaphides. Annales de la Société Entomologique de France, 73: 1–400.
Raffray, A., 1905. Genera et catalogue des Psélaphides. Annales de la Société Entomologique de France, 74: 401–476.
Raffray, A., 1908. Coleoptera. Fam. Pselaphidae. In Wytsmann, P. (ed.), Genera Insectorum, 64th fascicule. 487 pp.
Rome.
Reitter, E., 1880. Beiträge zur Käferfauna von Neu-Zeeland. Verhandlungen des Naturforschenden Vereines in Brünn, 18:
165–183.
Reitter, E., 1882c. Versuch einer systematischen Eintheilung der Clavigeriden und Pselaphiden. Verhandlungen desNaturforschenden Vereines in Brünn, 20: 177–211.
Reitter, E., 1885. Abbildungen und Bemerkungen zu wenig gekannten Pselaphiden–Gattungen mit Beschreibungen neuer
Arten. Deutsche Entomologische Zeitschrift, 29: 333–339.
Reitter, E., 1910. Neue Coleopteren aus den Familien der Pselaphiden und Scydmaeniden nebst Bemerkungen zu verschiedenen bekannten Arten. Wiener Entomologische Zeitung, 29: 151–163.
Schaufuss, C., 1888. Catalogus synonymicus Pselaphidarum adhuc descriptarum. Tijdschrift voor Entomologie, 31:
1–104. (Not seen.)

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Schaufuss, L. W., 1880a. Beschreibung sechzig neuer Pselaphiden. In Der Societe entomologique de Belgique zu Brussel
zur Feier Ihres fur fundzwanzigsten Stiftungstages die herzlichsten Festgrusse aus dem Museum Ludwig Salvatorin Oberblasewitz–Dresden., pp. 5–35. (Not seen.)
Schaufuss, L. W., 1880b. Beschreibung sechzig neuer Pselaphiden. Nunquam Otiosis, 3: 481–511.
Schaufuss, L. W., 1887. Beschreibung neuer pselaphiden aus der Sammlung des Museum Ludwig Salvator. Ein Beitragzur Fauna Brasiliens, der Kgl. Niederlandischen Besitzungen in Indien und Neuhollands. (Fortsetzung.) Tijdschrift
voor Entomologie, 30: 91–165.
Sharp, D., 1874. Descriptions of new genera and species of Pselaphidae and Scydmaenidae from Australia and New
Zealand. Transactions of the Entomological Society of London, 1874: 483–518.
Sharp, D., 1881. Some new species and genera of Coleoptera from New Zealand. Entomologist’s Monthly Magazine, 18:
46–51.
Sharp, D., 1886. On New Zealand Coleoptera, with descriptions of new genera and species. Scientific Transactions of theRoyal Dublin Society, 3(2): 351–454.

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Plate 1 (Figs. 1–16). 1. Exeirarthra enigma Broun, female?; 2. E. enigma?, male; 3. ditto, fore leg enlarged; 4.
Sagola bilobata Broun, male; 5. S. dickensis Broun, male paratype; 6. S. eminens Broun, male; 7. S. halli
Broun, male; 8. ditto in ventral side, enlarged; 9. S. lineata Broun, male; 10. S. major Sharp, male; 11. S. minuscula Broun, male; 12. S. monstrosa Reitter, male; 13. ditto, right hind leg enlarged; 14. S. robusta Broun“var.”, male; 15. Stenosagola sp., female; 16. ditto, male.

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Plate 2 (Figs. 17–32). 17. Euplectus? sp., female; 18. E? sculpturatus Broun, male?; 19. ditto, labels; 20. Leptoplectus sp., male; 21. Macroplectus spinipes (Broun), male; 22. Dalma gigantea Broun, male; 23. Dalmisus
batrisodes Sharp?, male; 24. Eleusomatus acuminatus (Broun), male; 25. E. caudatus (Broun), male; 26. Euglyptus foveicollis Broun, male; 27. E. iracundus Broun (from Euplectus, see remarks), male; 28. Euplectopsis
crassulus Broun, male in ventral view; 29. E. heterarthrus Broun, male; 30. ditto, labels; 31. E. terrestris
Broun, male; 32. Philiopsis? sp., male.

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267

Plate 3 (Figs. 33–48). 33. Plectomorphus anguliferus Broun, male; 34. ditto, in ventral view; 35. P. trisulcicollis(Broun), male; 36. ditto, in ventral view; 37. Sagolonus impressus Broun, male; 38. S. patronus Broun “var.”,male; 39. Vidamus calcaratus Broun, male; 40. V. trochanteralis Broun, male; 41. ditto, in ventral view; 42.
Whitea laevifrons (Broun), male; 43. ditto, in ventral view; 44. Zelandius raffrayi (Broun), male; 45. Z. obsculus (Broun), female; 46. Alloplectus claviger (Broun), female?; 47. Neosampa granulata Broun, male; 48.
ditto, head to elytral base enlarged.

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Plate 4 (Figs. 49–64). 49. Anabaxis electrica (King), female; 50. A. foveolata (Broun), female; 51. Eupines
impar (Sharp), male; 52. ditto, left antenna enlarged; 53. E. micans (Sharp), male; 54. ditto, left antenna enlarged; 55. E. monstrosa (Reitter), male; 56. ditto, left antenna enlarged; 57. E. rudicornis Broun, male; 58.
ditto, left antenna enlarged; 59. E. sternalis (Broun), male; 60. ditto, left antenna enlarged; 61. E. nasuta(Broun), male; 62. Eupinogitus sulcipennis (Broun), female?; 63. Gastrobothrus abdominalis (Broun), male inventral view; 64. G. sharpi (Broun), male.

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Plate 5 (Figs. 65–80). 65. Physobryaxis inflata (Sharp), female; 66. ditto, male; 68. Simkinion prelaticum Park et
Pearce, female paratype; 69. Pselaphogenius? citimus (Broun), sex unknown; 70. ditto, head to elytral base enlarged; 71. P?. delicatus (Broun), sex unknown; 72. ditto, haed and pronotum enlarged; 73. Pselaphophus
atriventris (Westwood), female?; 74. ditto, labels; 75. Pselaphus? dulcis Broun, sex unknown; 76. P?. pauper
Sharp, sex unknown; 77. P?. trifoveatus Broun, male; 78. P?. sp. 1, male; 79. P?. sp. 2, sex unknown; 80. P?.
sp. 3, male.

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Plate 6 (Figs. 81–96). 81. Pselaphotheseus hippolytae Park, male?; 82. ditto, head enlarged; 83. P. ihupuku Carlton et Leschen, male paratype; 84. ditto, head to elytral base enlarged; 85. Agatyrus fulvihirtus Broun, male;
86. ditto, in ventral view; 87. Gerallus punctipennis L. W. Schaufuss, male?; 88. Hamotulus armatus (Broun),
male; 89. H. cornutus Broun, male; 90. H. robustus Broun, male; 91. ditto, in ventral view; 92. H. spinipes(Broun), male; 93. Phormiobius halli Broun, male; 94. ditto, in ventral view; 95. Tyrogetus palpalis Broun, female; 96. Zeatyrus lawsoni Sharp, male.

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Plate 7 (Figs. 97–112). 97. Sagola bilobatus Broun, “syntype”; 98. ditto, labels; 99. S. dickensis Broun, “syntype”; 100. ditto, labels; 101. S. halli Broun, “syntype”; 102. ditto, labels; 103. S. tumidus Broun, “syntype”;
104. ditto, labels; 105. Plectomorphus scitiventris Broun, “syntype”; 106. ditto, labels; 107. Vidamus
trochanteralis Broun, “syntype”; 108. ditto, labels; 109. Eupinogitus sulcipennis Broun, “syntype”; 110. ditto,
labels; 111. Pselaphus delicatus Broun (Pselaphogenius), “syntype”; 112. ditto, labels

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Plate 8.

Geographic area codes of New Zealand by Crosby et al. (1998).