Vol. XIV' Nos.10-12'1926'][Reprintedfrom InsecutorInscitie Menstruus,

UNDESCRIBED SPECIES OF TIPULINE CRANEF'LIES F'ROM TROPICAL AMERICA(Diptero,Tipulidae)
BV CHARLES P. ALEXANDER
The majority of the speciesdescribedin this paper are fromthe State of Tachira, Venezuela,where they were collectedby
Messrs. E. B. and J. H. Williamson and W. H. Ditzler, towhom I am very grateful for the gift of this valuablematerial.

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161blotch immediately beneath the second blotch in the radial cell
but barely confluent with it; outer end of cell M darkened;
stigma brown; veins beyond the cord in the radial field conspicuously seamed with brown, almost obliterating the groundcolor; other cells beyond the cord and the distal portions ofcells Ca and Anal brownish gray, the basesof the latter broadly
pale, this including more than the basal half of cell Cz, thebasal quarter of rsf A and the basal half of znd, A; veins darkbrown. Venation: Veins .R3 and Ra15uousuolly sinuous, constricting cell R3 near mid-length; cell rst Mz relatively smail,pentagonal, the basal section ol My2 longest, that of Ms,amore than twice the length
shortest; petiole of cell Mtalittleof m.
Abdominal tergites light brown, trivittate with brownish
black; segments six to eight blackened; hypopygium brown;basal sternites pale.
Habi,tat: Peru.
Holotype, J, Yurimaguas, April I2,1920 (H. S. Parish).

One additional Holorusia was included in Peruvian material
collected by Parish and another in the extensive series of Tipulidae belonging to the Paris Museum and kindly loaned to me
for study by my friend, M. Dr. Eugene S6guy. The types of
all the speciesexcept this last-mentioned are preserved in thewriter's collection.
Holorusia sinuosa, new species.
General coloration light brown; antennae with the flagellar
segaents indistinctly bicolorous; mesonotal praescutum withthree reddish brown stripes; wings subhyaline with a very heavy
and extensive brown pattern, including three blotches in cells
R and M ; vein Raa5verlr sinuous; abdominal tergites trivittate
with dark brown.
Male: Length, 1? mm.; wing, 20.5 mm.
Frontal prolongation of the head brown, slightly darker dorsomedially; palpi brownish black. Antennae short; first scapal
segment dark brown except at tip; second scapal segment andfirst flagellar segment obscure yellow; remaining flagellar segments brown, the incisures very indistinctly paler. Head darkbrown, the posterior orbits of the eyes paler.
Mesonotal praescutum testaceous brown with three reddish
brown stripes, the interspaces darker brown behind; remainderof the mesonotum light brown, the scutellum and postnotumwith an indistinct capillary darker line; a darker brown Ushaped area bordering the postnotal pleurotergite before thehaltere. Pleura testaceous, indistinctly variegated with darker.
Halteres dark brown, the base of the stem restrictedly yellowish. Legs with the coxae pale buff; trochanters obscure yellow;
femora dark brown, the tips broadly blackened; tibiae darkbrown, the tips narrowly still darker; tarsi very long, much exceeding the remainder of the legs, dark brown. Wings subhyaline with a very heavy and extensive brown pattern; prearcularcells pale; cell C obscure yellow, cell Sc brown; three brown
blotches in cell R, the last at origin of Rs, narrowly connected .
with the second along vein M ; the first of these blotches likewise includes the base of cell M: in cell M there is a second

Holorusia guatemalensis, new species.
General coloration pale brown, the praescutal stripes r:bscureyellow; wings pale brown, heavily variegated rvith darl<erbron,nand subhyaline; cell tst Mz small; Rs long, paralleling R1.
Male: Length, 13.5 mm.; wing, 18.5 mm.
Frontal prolongation of the head obscure brownish yellow,
the nasus infuscated; palpi dark basally, brightened outwardly.
Antennae small; scapal segmentsobscureyellow; flagellum darl,brownish black; first scapal segment very long. Head lightbrown, paler anteriorly.
Mesonotal praescutum pale brown rvith four obscure yellow
stripes; scutum similarly marked, the centers of the lobes being
pale; remainder of mesonotum obscure brownish yellow. Pleura
obscure yellow. Halteres brown, the base of the stem obscureyellow. Legs with the coxae and trochanters obscure yellorv;
legs long and slender, brown, the tips of the femora and tibiae
dark brown. Wings with a pale brown tinge, variegated withdarker brown and subhyaline; prearcular region and cells C andIIII

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Sc more yellowish; darker brown ateas at the stigma; origin ofRs,' in bases of cells R and M; at about one-third the length ofcell ft,' at near midlength and at distal end of cell M; wingapex darkened; the subhyaline blotches appear as interspaces
between the darkened areas in cells R and M, in cell rst Ry
base of cell Rz and in the bases of cells Cu to znd l; vgins
dark brown. Venation: Sc2 ending before the fork of Rg thelatter vein very long, strongly arcuated at origin, running parallel to R1; Rz*s straight; Rs and Ra*6 &rc[ated, constricting cellRs near midlength; cell rst Mz small, pentagonal, the basal section of Ma,+the shortest element of the cell, the other elements
more nearly equal in length; cell Mt deep, twice as long as itspetiole.
Abdomen pale brown, the incisures narrowly pale; a subterminal brownish black ring, including tergites six and sevenand sternite seven; hypopygium pale. Male hypopygium oftype with the adeagus long exserted.
Habi,tat: Guatemala.
Holotype, J, In the Paris Museum, labelled " L9 - 55 ", Angrand.
The body of the type is somewhat discolored and the coloration is discussedin general terms only.
Holorusia strangalia, new species.
General coloration brown; antennal flagellum bicolorous;
mesonotal praescutum with four reddish brown stripes that arenarrowly margined with darker brown; pleura yellow; wings
gray, with a heavy brown pattern that includes the broad costal
margin and apex; a large brown blotch in cell M ; aMomen of
male with a black subterminal ring, the hypopygium yellow.
Male : Length, 15 - 15.5 mm.; wing, 18 - 18.5 mm.
Female: Length, 19 mm.; wing, 20 mm.
Frontal prolongation of the head brownish yellow; nasus
elongate, black; palpi brownish black, the terminal segments
passing into obscure yellow. Antennae (male) of moderate
length, if bent backward extending about to the wing-root;
scapal segments yellow, the second brighter; basal flagellar

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segments bicolorous, the basal half of each dark brown, thedistal portion obscure yellow, the terminal segments more uniformly dark brown. Head light brown, more yellowish on thefront; a capillary brown median line; orbits paler.
Mesonotal praescutum brown with four conspicuous reddish
brown stripes, each of which is margined with dark brown;
scutum brown, each lobe with a reddish brown blotch near theanterior margin; scutellum and postnotum dark brown, thelatter sparsely pruinose; posterior margin of scutum, the scutellum and postnotal mediotergite with an appressed yellow
pubescence;pleurotergite pale, concolorous with the pleura, thecaudal half darker. Pleura yellow. Halteres brown, pale basally. Legs with the coxae yellow; trochanters strongly tingedwith greenish; femora and tibiae brownish testaceous,the tips
darker; tarsi brown; claws toothed. Wings gray with a heavy
brown pattern; costal region, including cells C, Sc and anterior
half of R dark brown; wing-apex in cells znd Ry Rz and Rs
dark brown; stigmal spot still darker brown; a seam along Cu
and m-cu; a large rectangular blotch near midlength of cell l\[brown ; cell M and a small area before the stigma subhyaline;
irr some specimensa subhyaline spot in the outer ends of cells
Rz and R3; veins dark brown. Venation: Vein Ra*5 strongly
and irregularly sinuous, constricting cell R3; second section of
M3*a longer than the basal section of the same.
Abdomen brown, the tergites indistinctly marked sublaterallywith dark brown; segmentssix and seven blackenedto produce
a subterminal ring; hypopygium large, compressed, conspicuously yellow.
The female is similar to the male but the antennaeare shorter:
no dark subterminal ring on abdomen. Ovipositor with thevalves conspicuous, straight, compressed.
Habitat: Venezuela.
Holotype, {,T-,aFria, Tachira, April 14, 1920 (J. H. & E. B.
Williamson and W. H. Ditzler).
Allotopotype,Q, April 77,7920.
Paratopotype, {, April 17, 7920; paratlfe, 3, Tachira,Tachira, April ?, 1920.

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rNsEcuToR rNscrTrd MENsrRUus

Tipula zonalis, new species.
General coloration orange-yellow; wings faintly tinged withbrownish, the costal region, apex and veins beyond the cordconspicuously seamed with brown; abdomen with a narrow
black subterminal ring.
MaIe: I*ngth, 11 mm.; wing, 11.2 rnm.; antenna, ?.3 mm.
Female: Length, 14 mm.; wing, 12.2 mm.
Frontal prolongation of the head shiny obscure yellow; palpi
testaceous, the terminal segment darker. Antennae (male)elongated; scape obscure yellow; first flagellar segment yellow,
dark brown subterminally; remaining flagellar segments black"the extreme base and the apex of each segment conspicuously
yellow to produce an annulate appearance. Head shiny obscure yellow.
Thorax shiny orange-yellow, without rnarkings. Pleura
yellowish testaceous, immaculate. Halteres obscure yellow, theknobs brown. Legs with the coxae and trochanters concolorouswith the pleura; femora and tibiae brownish yellow, the tips
narrowly brownish black; tarsi dark brown. Wings with afaint brownish tinge, the base, costal region, apex and the veins
beyond the cord conspicuously seamed with darker brown, thedarkened areas including cells C, Sc, distal half of cell ls/ R1,
all of cells znd Rr and R2 and all but the base of R3,' the cells
beyond the cord have pale centers; veins dark brown. Venation: Sc comparatively short, Sc2 ending about opposite onethird to one-fifth the length of Rs,' R2*3 longer than R2; petioleof cell Ml about one-half longer than m; cell rst M2 elongatepentagonal; basal section of M*z and,w subequal; m-cu at or
immediately before the fork of Ms,a.
Abdomen shiny obscure yellow; segment seven and the baseof eight deep black. In the female the lateral margins of thetergites are distinctly blackened, the mid-dorsal line less distinctly darkened. Male hypopygium with the adeagus tremendously exserted, about one-half longer than tlrc entire body.
Ovipositor with the tergal valves somewhat fleshy, as in Tsmithi. Alexander, to which the prusent species is obviouslyallied.

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Habitat: Venezuela.
Holotype, J, Tachira, Tachira, April 4, 1920 (J. H. & E. B.
Williamson and W. H. Ditzler).
Allotopotype, Q.
Ti.pula zonalis is allied to T. pwishi Alexander (Brazil) andT. smithi Alexander (Brazil).
Tipula detecta, new species.

IIGeneral coloration brown and yellow; antennal flagellum entirely dark brown; mesonotalpraescutumwith four darker brown
stripes; wings with a strong brownish tinge, more suffused along
the costal region; tip of R2 preserved; abdominal tergites reddish brown, the lateral margins conspicuously blackened; tergite seven entirely blackened; sternites uniformly obscureyellow.
Female: Length, 1? mm.; wing, 16 mm.
Frontal prolongation of the head obscure yellow, a little
darker dorso-medially and on nasus; basal two segments ofpalpus brown, the terminal segmentsconspicuouslyyellow. Antennae with the scape yellow; flagellum dark brown. Headbrown, very narrowly and indistinctly paler on the posterior
orbits; a darker brown median vitta.
Mesonotal praescutum rather bright yellow, darker behind;four darker brown stripes, the intermediate pair narrowly separated for the most of their lengths, connectedposteriorly, theirinner margins narrowly bordered with darker; scutum obscure
blownish yellow, the centers of the lobes dark brown; scutellum
and postnotum obscure brownish yellow. Pleura yellow witha faint whitish pruinosity. Halteres rather long and slender,
dark brown. Legs with the coxae and trochanters concolorouswith the pleura; remainder of the legs dark brown, the femoral
basesa little paler; basitarsi much longer than the tibiae. Wings
with a strong brownish tinge, cells C and Sc darker brown;
stigma oval, still darker brown; small obliterative areas before
the stigma and across the base of cell rst M2, extending fromcell R to the base of cell Mg,' veins brownish black. Venation:
Sc2 ending about opposite one-third the length of Rs,. distal sec-

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tion of R2 preserved; cell M1 deep, its petiole about one-halflonger than w; cell rsf M2 elongate-pentagonal, the basal section of My2 and ru subequaland parallel to one another; second
section of Ms*a relatively long; cell znd A long and narrow.
Abdominal tergites reddish brown, the lateral margins broadly and conspicuously brownish black; on segment seven andthe posterior half of segment six producing a subterminal ring;
genital segment light brownish orange; sternites obscure yellow.
Valves of the ovipositor dark brown, only the tips horncolored; tergal valves comparatively deep, straight, the tips
truncated; sternal valves much shorter but similar in shape.
Habitat: Venezuela.
Holotype, t, Tachira, Tachira, April 4, 1920 (J. H. & E. BWilliamson and W. H. Ditzler).
Tipula detecta belongs to the group of T. effeta Alexanderof Peru. For a discussion of this group of species,see ProcAcad. Nat. Sci. Philadelphia,1921, Pt.1: 100-102; 1927.
Tipula tecta, new species.
General coloration light brown; male antennae elongate;wings brown, the stigma darker brown; distal section of Rz
atrophied; abdominal sternites yellow, the incisures heavily
marked with brown.
MaIe: Length, 11 mm.; wing, 11 mm.
Frontal prolongation of the head comparatively short, obscureyellow above, darker laterally; nasus short and broad; palpiblack, the terminal segment fading into yellow. Antennae withthe scape yellow; flagellum dark brown; antenna elongate, if
bent backward extending to beyond midlength of the third
abdominal segment; flagellar segmentswith a well-marked basal
swelling. Head grayish brown, the front yellow.
Mesonotal praescutum light brown with three darker brown
stripes, the broad median stripe very indistinctly split anteriorly
by a pale median line; remainder of the mesonotum brown.
Pleura pale buffy yellow. Halteres long, brown. Legs withthe coxae concolorous with the pleura, the fore coxae darker,
the coxae provided with long pale setae; trochanters pale;rNsEcuToR rNscrTrd MDNSTRUUS

167

legs very long and slender, brownish black, the femoral bases
paler. Wings with a strong brownish tinge, cells C and Sc
slightly darker; stigma oval, dark brown; conspicuous subhyaline areas before and beyond the stigma; a faint obliterative
area in cell rst M2, passing into cell Me; veins dark brown.
Venation: Sc2 ending just before midlength of Rs, the latter
about one-half longer than R2,s, gently arcuated at origin;distal section of R2 entirely atrophied; cell rsf M2 narrowed
distally, m being only a little longer than r-m; petiole of cell
Mr about as long as tlie basal deflection of Rl.r,' m-cu about at
the fork of Ms*a, the latter vein short, only about one-fifth thelength of the first section of Ms; cell znd,l of moderate width.
Macrotrichiae on veins Sc, R, t, Rz*s, R3 and Ra15.
Abdomen yellowish brown, the tergites indistinctly trilineate
with dark brown; segments seven and eight black; sternites
yellow, the base and apex of each segment conspicuously darkbrown medially. Male hypopygium conspicuously yellowish
brown. Ninth tergite with a deep U-shaped notch, the laterallobes small and narrow; from the base of the notch arise two
conspicuous chitinized lobes directed caudad, separated from one
another only by a small oval notch, their apices obliquely truncated and microscopically denticulate.
Habitat: Venezuela.
Holotype,6f, Tachira, Tachira, April 10, 1920 (J. H. & E. B.
Williamson and W. H. Ditzler).
Tipula tecta is a doubtful member of the genus, since it possessessome characters of the subtribe Dolichopezaria.