On New or little-known Tipulidse.

639(I5) K),U~'MANN, R. 1943. " Aquatic Coleoptera of' N o r t h a n d SouthD e v o n . " E . M . M . lxxix, pp. 185-198 : 228-230,(I6) L I ~ E L ' , , J. lStiT. " Nat. Hist. o1" Reigate and its V i c i n i t y . - Coleoptera," H:ohnes(lale N~,L. l:[ist. Club, l{eigate.
(I7) M'L]~oD, W . J .
1912, " .Fur~iler Notes on the Aqm~tie Coleopteraof lVIonklands (Lanarkshire)." Glasgow Naturalist, iv. p. 117.
(18) ~[ONClCEAFF. 1840--1880.
' A Catalogue of the Coleoptera of ,theIsland of Portsea a n d Distriet,' MS. list.
(I9) NI,:WBErcY, E . A .
1907. " ttydr~vna longior LRey a n d Ochthebb~sviridis Peyron, additions to the British List of Coleoptera."
E. M. M. xliii, pp. 172, 173.
(2o) PEA~C~., E . J .
1926-1929. " Coleoptera of Dorset a a d Stipplemerits." Dorset Nat. Hist. mid Antiquarian FieId Club &e.
(21) SCHAUM,H. 1947. " l~evisiolt of British H y d r o e a n t h a r i d m . " Zoologist~ vi. pp. 1887-1897.
(22) SHAaP, D. 1914. " The British Species of Gyrinus." E . M . M . 1.
pp. 128-138.
(23) STJ~P~:~NS, J. F. 1828. ' Illustratiolls of British E n t o m o l o g y . - 5Iandibula, ta,' ' ii.
(24) - - - .
1829. ' (!at. of British insects.'(25) STO'rT, C. E, I927, " A S u p p l e m e n t a r y Note on the List of the
Coleoptera of D o r s e t . " E . M . M, lxiii, pp. 91, 92.
(26) TONLIN, J . g .
1907. " Caqa'mb.us9-lineatus Steph. in B e r k s h i r e . "E. M:. 1Yr. xlv. p.15
• 1913.
" The Coleoptera of Glamorgan." Trans. Cardiff(27)
Nat. See. vl.
(28) - - .
1919.
" New localities for Hydrovat~,s cly2)ealis S h a r p . "E. NL ~i. Iv. p. 159.

X L V . - - N e w or little-known Tipulidse (Diptera).--LXXXII.
Oriental-Australasian Species.
By CRaRLES P.
ALEXANDER, Ph.D., F.I~.E.S., University of Massachusetts, Amherst, Massachusetts, U.S.A.

THE Tipulid~e discussed herewith were taken by Mr. P.
Susai Nathan, chiefly at Cinchona, in tile Anamala,i Hills,
in South India. I am greatly indebted to Mr. Susai
Nathan for the privilege of retaining the types of thenovelties in my extensive personal collection of theseflies.
Ctenacroscelis inclytus, sp. n.
Belongs to the umbrinus group; femoral tips broadly
black, preceded by a clearer yellow ring ; wings strongly
infilseated, patterned with more yellowed areas, including
large ones in outer end of cell M and near outer end ofcell 1st A ; abdomen dark reddish brown, the tergites
m o r e blackened on sides.
44 ~

640

Dr. C. P. Alexander on

Fe't~ude.--Length about 26 ram. ; wing 26 ram. ;antenna about 3 ram.
Frontal prolongation ot" head dark brown; ltaStts
elongate; palpi black, relatively short. Antennae withscape infnscated, corrugated; pedicel and first flagellar
segment brownish yellow, the succeeding segments passing
into cinnamon brown; flagellar segments with ventral
face only moderately produced, each segment with ~bouttwo short black vertieils on this face. Head above
brown, broadly light grey pruinose on front and vertex.
Pronotum brown. 5lesonotal pr~escutum with fourbrownish grey stripes that are conspicuously borderedby dark brown, including the posterior interspaces;
lateral and humeral borders of prmscutum dark brown,
restricting the huffy yellow ground to delicate lines on
the outer and anterior edges of the laterM stripes and as acapillary median v i t t a ; posterior sclerites of notum
chiefly dark grey, the scutal areas more or less distinctly
bordered by brown ; scutellum with indications of a palecentral vitta, the mediotergite with a corresponding blackline; posterior border of mediotergite and much of the
katapleurotergite brownish black. Pleura with a verybroad and conspicuous dark brown longitudinal stripe
extending from the ventral cervical region across thedorsal selerites to the postnotum; dorsopleural region
broadly pale yellow; ventral pleurites more greyish
yellow, more or less patterned with brown, most distinct
on the ventral sternopleurite. Italteres infuscated, thestem somewhat paler. Legs with the coxm light grey,
the fore and hind pairs patterned with brown ; trochantersreddish yellow ; femora obscure brownish yellow, the tips
very broadly and conspicuously blackened, preceded by aslightly clearer yellow ring; tibiae and tarsi brownish
yellow, the tips of the former narrowly darkened. Wings
with a strong brownish suffusion, somewhat darker in the
costal area and only slightly less so in cells R and M ;preareular, apical and anal fields a little less darkened;
conspicuous yellowed areas at and above arculns ; before
stigma, continued caudad as a narrow line along the cord
through cell 1st M e ; paler, more diffuse yellow areas inouter end of cell M and as marginal markings in cell
1st A ; small brown seams at posterior end of m - c u ,outer end of cell 1st M 2 and fork of M1+2 , and as smMler

New or littIe-k,nown Tipulid~e.

641

marginal spots at ends of the veins; veins light brown,darker in the patterned portions, pale yellow in the obliterated areas. Veins beyond cord glabrous, including
R4+ 5. Venation : m-su at midlength of Ms+~; petioleof cell M 1 about two-thirds m.
Abdominal tergites dark reddish brown, more brownish
black on sides, with fainter indications of a mediandarkening ; sternites paler reddish brown.
tlab. South India.
Ilolotype, 9, Cinchona, Anamalai Hills, altitude 4000-5000 fbet, August 30, 1946 (P. Susai Nathan).
The present fly is readily told from the other membersof the group, including Ctenacroscelis ochripe8 (Brunetti)
and C. infindens (Walker), by the pattern of the body,legs and wings. In the unusually short antennal vertieils
the species agrees with the members of the albovittatu~.
group, but the position in the umbrinus group seems
correct.

Tipula (Indot@ula ) b~uchycantha, sp. n.
Thorax uniformly orange yellow; antennm (male)
relatively long, scape and pedicel yellow, flagellum black;abdomen reddish brown with a broad black subterminal
ring; rome hypopygium with the tergite terminatingin two rounded spieulose lobes; outer dististyle widest
at near midlength ; outer basal lobe of inner dististyle aslender curved spine; gonapophyses terminating in
flattened blades.
Male.--Length about 12-13 mm. ; wing 13.5-14 ram. ;antenna about 4.5 ram.
Frontal prolongation of head yellow, relatively short ;nasus elongate; palpi brown, with paler incisures.
Antennm (male) relatively long, as shown by the measurements ; scape and pedicel yellow, flagellum black ; flagellar
segments very gently incised, subequal to or longer thanthe verticils. Head with front yellow ; posterior portion
greyish brown, with a capillary black median vitta.
Thorax uniformly orange yellow, only the pleura atrifle paler. Halteres brown, the base of stem yellow.
Legs with all coxm and trochanters yellow ; remainder oflegs brown, femoral bases restrictedly brightened ; claws(male) with a large tooth at near midlength and with asmM1 basal spur. Wings subhyaline, the prearcular and642

Dr. C. P. Alexander on

costal fields more yellowed: stigma ovM, brown; veinsdark brown. Macrotriehia on most of distal section ofR~+5 and on vein M 1. Venation: Rs short, subequal
to or shorter than m-cu; m subequM to petiole of cell
M1; cell 2nd A narrow.
Abdominal tergites reddish brown, darker laterally;
tergites six to eight, inclusive, black, the lateral borders
broadly pale; eighth sternite similarly blackened, remainder of hypopygium brownish yellow. Male hypopygium with the tergite terminating in two rounded lobes
that are provided with numerous blackened spicules but
without accessory lobes. Outer dististyle widest at near
midlength, on lower margin dilated and with a groupingof black setm. Inner dististyle with the outer basal lobe
a small slender curved spine; beak blackened, simple,
completely cut off from the dorsal crest by a membranous
incision, the dorsal row of set~e continuous across thebreak ; lower beak lying tkr basad, appearing as a small
blackened knob. Gonapophysis appearing as an expanded
pale blade, the tip obtusely rounded.
Hab. South India.
Holotype, ~, Cinchona, Anamalai Hills, altitude 40005000 feet, September 2, 1946 (P. Susai Nathan). Parato29otypes, 4 ~c~, September 2-October 5, 1946 (P. Susai

Nathan).
Among the numerous regional species, the present
fly is closest to forms such as Tipula (Indoti2pula) elegantula Brunetti, differing in the details of coloration andin the structure of the rome hypopygium, especially theouter and inner dististyles.

Tipula (lndot@~da) dil~tistyta, sp. n.
:Belongs to the leucopyga (sulaica) group~

generalcoloration yellow, the pr~escutum with four nearly
confluent reddish brown stripes ; femora obscure yellow,
the tips darker; wings brownish yellow, the prearcularand costal fields more saturated; male hypopygiumwith outer angle of tergal plate produced into a strong
incurred black spine, with a more ventral inner lobe;
outer dististyle strongly dilated outwardly, weakly
bilobed at apex, the lower lobnle with a concentration ofstrong black setse.

N e w o~" little-known Tipulid~e.

643

Male. Length about 15 mm. ; wing t6.2 ram. ; antenna about 3 ram.
Frontal prolongation of head obscure yellow, relatively
short, only about one-half as long as remainder of head ;nasus long and slender, nearly one-half the prolongation ;palpi with basal two segments obscure yellow, the outerones brownish black. Antennae small; basal threesegments obscure yellow, the remaining segments uniformly black; flagellar segments strongly binodose, as
in the group, the outer node larger and more triangularly
produced than the basal swelling; verticils very long.
Head brownish grey, with a capillary brown median line ;a small dark spot on either side of the anterior vertex
opposite the narrowest point.
Cervical region dark brown. Pronotum brownish grey,
paling to yellow on sides. Mesonotum chiefly yellow,
the prsescutum with four reddish brown stripes that are
relatively conspicuous, the various stripes almost confluent ; seutal lobes with reddish brown areas ; posteriorsclerites of notum and the pleura more whitened ; verysmall and vague darkenings on the cephalic portion of theanepisternum and sternopleurite. Halteres dark brown,
the base of stem restrietedly brightened. Legs with thecoxm and trochanters yellow ; femora obscure yellow, thetips darker brown; tibim brown, the tarsi passing into
black; claws (male) with a basal and a medial tooth.
Wings with a brownish yellow tinge, the prearcular andcostal fields more saturated yellow; stigma medium
brown; veins brown, more brownish yellow in the
fiavous fields. Venation: R s a n d m - c a subequal inlength; R~+ 2 preserved, relatively short, about one-halflonger than vein R~ ; m and petiole of cell M 1 subequal ;m - c u a t fork of M ; cell 2nd A relatively narrow.
Abdominal tergites chestnut brown, the outer segments
more darkened to form a ring ; pleural membrane infuscared ; basal sternites yellow ; male hypopygium yellow.
Male hypopygium with the outer angles of the tergal
plate produced into a strongly incurred black spine;
median region of tergite slightly produced, the margin
narrowly blackened ; beneath this rim on either side witha stotlt lobe that terminates in two or three strong
blackened spines. Outer dististyle strongly dilated outwardly, weakly bilobed at apex, the upper lobe ~ith

644

Dr. C. P. Alexander on

delicate setse, the lower one with a concentration ofstrong black setse. Inner dististyle with the beal~ slender,
lower beak reduced; outer basal lobe simple, obtuse,
with relatively sparse setse; on its face with a small
finger-like lobule that is provided with scattered bristles.
[lab. South India.
Holotype, ~, Cinchona, Anamalai Hills, altitude 40005000 feet, September 17, 1946 (P. Susai Nathan).
This fly is most sinfilar to Tipula (Indotipula) palnica
Edwards, having the tergite of the male hypopygium
somewhat similar but with the outer dististyle quite
different.

Tipula ([ndotipula) oreoctistes, sp. n.
Allied to .flavithorax; thorax chiefly reddish yellow,
unpatterned; wings with a weak brownish tinge, thecostal border and stigma more brownish yellow; veins
beyond cord vaguely more seamed with brown; vein
RI+ ~ glabrous, usually entire, its tip not bent strongly
cephalad; abdomen reddish yellow, with a brownish
black subterminal ring ; male hypopygium with the outerbasal lobe of the inner dististyle produced into a longsinuous spine, the remainder unusually glabrous ; eighthsternite bearing a median depressed lobe.
Male.--Length about 13-14 ram. ; wing 17-18 ram. ;antenna about 5.1-5.2 mm.
Female.--Length about 19-20 ram. ; wing 17-18 ram.
Frontal prolongation of head yellow, relatively short,nasus elongate; palpi brown, the incisures restrictedly
pale. Antennse with basal three segments yellow=, thesucceeding ones bicolo'-red, black, their apices narrowly
yellow, the amount of pale colour decreasing on the outer
segments, becoming obsolete beyond midlength of organ ;flagellar segments weakly incised, longer than the verticils.
Front and anterior vertex yellow, the posterior vertex
grey; vertical tubercle very low, entire.
Pronotum light brown, paler on sides. Mesonotum
chiefly reddish yellow, the prsescutal stripes not or
scarcely indicated ; prsescutum with vestiture exceedingly
sparse and small to virtually lacking. Pleura andplenroteNite more yellowed. Halteres with stem brownish yellow, the knob darker. Legs with coxse andNew or little-known Tipulidm.

645

trochm~ters yellow; femora and tibise brownish yellow,
the tips rather narrowly blackened, those of the mid.
tibise even more restricted; tarsi brown; claws (male)slender, toothed. Wings with a weak brownish tinge,
the prearcular and costal fields, with the stigma, morebrownish yellow to light brown; veins brownish black,
those beyond cord vaguely more seamed with brown;veins basad of cord, especially in the brightened fields,much paler. Veins beyond cord with abundant trichia;
R~+e glabrous and pale, in eases with the tip atrophied.
Venation: g~" longer than R2+a; cell M 1 about twice
its petiole; cell 1st M 2 small, pentagonal, with rn-cu ator beyond the fork of M3+4.
Abdomen reddish yellow, the tergites more or lessdarkened; segments seven and eight brownish black toform a subterminal ring; hypopygium fulvous. In thefemale, abdomen almost uniformly darkened but thisseems due, in part at least, to internal discoloration.
Male hypopygium with the caudal border of tergite witha deep U-shaped notch, the lateral lobes appearing as
slender glabrous horns; at base of notch with a low
obtuse lobe. Dististyle parallel, on lower face with asmall setiferous lobule that represents the usual outerdististyle ; outer basal lobe a long slender sinuous spine,
approximately two-thirds as long as the entire style.
Eighth sternite bearing a profoundly divided depressed
lobe, either half relatively narrow, the outer margin withrelatively few long setse.
Hab. South India.
Holotype, c~, Cinchona, AnamalM Hills, altitude 40005000 feet, September 16, 1946 (P. Susai Nathan). Allotopotype, ~, September 21, 1946. Paratojootypes, df¢.
The onlytclose relative appears to be Ti2pula (lndotil)ula)
flavithorax Brunetti, of Which I possess a homotypet~ma.le that had been compared with the type by Edwards
and which certainly appears to be distinct. Edwards(Rec. Indian Mus., 26: 306; 1924) has provided
additional notes on the type male of flavithorax, further
indicating a different species.

fl'richo~eura (X@holimnobia) umbr.@enni,% sp. n.
General coloration of body dark brown, the dorsoplenral
region of pleura and a more ventral pleural stripe narrowly

646

Dr. C. P. Alexander onwhitened; legs brown; wings with a strong blackish
tinge, cell C light yellow; male hypopygium with the
apex of the arm of the outer dististyle terminating in astrong gently curved spine.
Male.--Length about 2.5 ram. ; wing 2.9-3-2 ram.
Female.--Length about 3 mm. ; wing 3 ram.
t~ostrum and palpi black. A n t e n n a short; scapeblack, pedicel brown; flagellum medium to pale brown.
Head grey.
Pronotum brown, the lateral portions, pretergites andlateral pr~escutal border whitened, the eolour reaching thewing root ; remainder of mesonotum dark brown. Pleura
brownish black with a very narrow but conspicuous palelongitudinal stripe extending from behind the ibre coxa~,
passing above the bases of the other eoxee, to the abdomen.
Halteres with stem obscure whitish, knob blackened,
Legs with the cox~e and trochanters yellow, the fore pairsomewhat darker ; remainder of legs dark brown. Wings
with a strong blackish tinge, ceil C to or beyond the levelof origin of Rs light yellow; veins brown. Venation:
Sc 1 ending about opposite four-fit~hs the length of ~ ' ,Sc 2 a short distance from its tip; m-cu approximately
its own length or less before the fork of M3+4. In aparatype, vein R1, apposite the atrophy of the free tip ofSc2, bends sharply down to R2+ 3 to produce a short fusion
of RI+,+. 3.
Abdomen, including the hypopygium, brownish black.
Male hypopygium having the tergal region with a very
deep and narrow median split, the adjoining lobes broad,truncated at tips. Outer dististyle complex, as in thegroup, the apex of the non-spieulose arm terminating in astrong, gently curved spine that is not fasciculate.
Hab. South India.
Holotype, ~, Coimbatore, September 21, 1939 (P. Susai~Vathan). Allotopotype, ~, September 8, 1939. Parstopotypes, 10 ~ , August 16-September 29, 1939 (P.
S,tesai Ncbtha~).
The present fly differs most evidently from Trichoneura(Xipholimnobia) nepalensis (Brunetti), of Nepal, in thedarkened legs, unusually darkened wings, and in otherdetails of coloration. Unfortunately the latter species
is still known only from the unique type female. By a
key to the Asiatic species of Trichoneura (Xipholimnobia)

New or little-known Tipulid~e.

647

by the writer (Philippine Journ. Sci., 53: 440; 1934),
the present fly runs to T. (X.) bontocensis Alexander ofnorthern Luzon, a distinct species.
Limon,~a (Geranomyia) fimbria,rum, sp. n.
General coloration of mesonotum plumbeous, pleuraand pleurotergite yellow; femora brown, the bases
restrictedly yellow; wing's subhyaline, very restrictedly
patterned with dark brown, most evident as the stigma ;costal fringe (male) very long and conspicuous; Sc
relatively long, Sc 1 ending just beyond midlength of Rs ;male hypopygium with the rostral prolongation of thev e n t r a l dististyle large and complex, bearing two very
unequal, widely separated spines from elongate tubercles,
the outer spine from a tubercle that is approximately
three times its own length.
Male.--Length, .excluding rostrum, about 6 ram. ; wing
6.7 ram. ; rostrum about 2-25 ram.
Rostrum relatively short, less than one-half the lengthof wing, black throughout. Antennm black throughout ;flagellar seglnents subcylindrical or truncate-oval; verticil~
relatively inconspicuous. Head with a narrow silvery
median stripe, the sides of the posterior vertex infuscated ;anterior vertex very nmTow, less than the diameter of
two rows of ommatidia.
Pronotum infuscated above, paling to yellow on thesides. Mesonotal pr~escutum with the dorsum chiefly
brown, sparsely pruinose to present a plumbeous appearance, the broad central stripe somewhat darker ; humeraland lateral portions more testaceous yellow; posterior
seterites of notum yellow, the scutal lobes and posterior
half of medioterg%e dark brown. Pleura and pleurotergite
yellow. Halteres brownish black, relatively short, theouter part of stem with conspicuous setse. Legs with allcox~e yellow ; trochanters more greenish ; femora brown,
the bases restictedly yellow; tibiae and tarsi greenish
brown, the outer tarsal segments darker. Wings subhyaline, very restrietedly patterned with dark brown,
most evident as the oval stigma; scarcely evident
darkenings at the supernumerary cross-vein in cell Sc,
Sc2, and the tips of veins /73 and R4+a; veins brown.
Costal fringe (male) very long and conspicuous, the648

Dr. C. P. Alexander onlongest setm fully equal to the combined width of cells C
and St. Venation: Sc relatively long, Nc~ ending justbeyond midlength of Rs, Sc 2 a short distance from its tip ;Rs long, about three times the basal section of R4+a;cell 1st M 2 long-rectangular, only a little shorter than the
distal section of vein MI+ 2 ; m-cu at or very close to thefork of M.
Abdominal tergites dark brown, sternites more yellowed.
Male hypopyghml with the tergite very shallowly emarginate, the sides nearly parMlel. Basistyle small, theventro-mesal lobe very reduced. Dorsal dististyle a curved
hook, its tip acute. Ventral dististyle relatively small,exceeding the basistyle in area; rostral prolongation
very large and conspicuous, sclerotized, with two widely
separated spines of unequal size; outer spine from an evenlonger basal tubercle, the spine exceeding in length therostral prolongation; basal spine about three times aslong as its tubercle. Gonapophysis with the mesal-apicallobe a slender darkened rod, its tip acute, strongly curved.
Hab. South India.
Holotype, c~, Cinchona, Anamalai Hills, altitude 40005000 feet, September 16, 1946 (P. Susai Nathan).
The most similar species is Limonia (Geranomyia)
longifimbriata Alexander, which is readily distinguishedby the open cell M 2 of the wings and especially by thedetails of structure of the male hypopygium, particularly
the rostral prolongation and the gonapophyses.
Eupilaria guttulifera, sp. n.
Mesonotum chiefly light brown, tile prmscutum vaguely
patterned with darker; humeral and lateral portions ofpr~escutum more darkened ; a large black spot on side ofeach scutal lobe behind suture ; pleura yellow, with two
narrow longitudinal black stripes; femora yellow, thetip and a subterminal ring darkened, enclosing a broad
yellow annulus; wings brownish fulvous, conspicuously
patterned with darker brown; abdomen chiefly yellow,
the outer segments vaguely bicoloured.
Male. -Length about 8-9 ram. ; wing 7.5-8.5 mm. ;antenna about 1-1-2 ram.
Female.~Length about 10 mm. ; wing 9 ram.
Rostrum very short, brown, palpi black. Antennze
with scape, pedicel and first segment of flagellum yellow,

N e w or little-l~nown Tipuli&e.

649

the remaining flagellar segments passing into brownish
black; basal tt~gellar segmeuts transverse subglobular,
crowded, the outer ones passing into elongate; outer
segments with very long conspicuous verticils. Headabove dark brownish grey, the front, posterior portion ofvetex and lower surface obscure yellow; anterior vertex
broad, approximately four times the diameter of scape.
Pronotum weakly infuscated medially above, moreyellowed on sides. Mesonotum chiefly almost uniformly
light brown or brownish yellow, in cases more pruinose,
the pr~escutum vaguely patterned with darker ; humeraland lateral portions of pr~escutum weakly infuscated ; aconspicuous black spot on side of scutal lobe, just inside
the wing root and behind the transverse suture. Pleura
ydlow, with two longitudinal black stripes, the more
dorsal one extending from the cervical region across the
propleura, ventral anepisternum and pteropleurite, notreaching the pleurotergite ; ventral stripe extending fi'om
the fore cox~e, crossing the middle of the sternopleurite,
more intense over the meron and metapleura, thence
merging with the lateral abdominal stripe ; remainder of
pleura and sternum light yellow. I-Ialteres with the stem
pale, knob weakly darkened. Legs with the fore coxm
darkened, as described, the other coxse clear light yellow ;trochanters yellow ; fbmora obscure yellow, the narrow tip
and a somewhat more extensive but paler subterminal
ring brownish black, enclosing a much broader yellow~nnulus ; tibiae obscure yellow, the tips blackened ; tarsi
black, the proximal portion of basitarsi paler. Wings
with a strong brownish fulvous tinge, conspicuously
patterned with darker brown, the small oval stigma
darker brown ; paler clouds at origin of Ra, over the cordand outer end of cell 1st 3¢/2, and as marginal clouds overthe ends of the longitudinal veins with the exception of
R.~ ; veins yellowish brown, darker in the patterned areas.
Venation : R~ shorter than R2+ a ; cell 1st M 2 longer thanM ~ ; m-.cu nearly its own length beyond the fork of M.
Basal abdominal tergites obscure yellow, the outer ones
weakly bicoloured, brown, wi~h the incisures somewhatt)nler; lateral borders brownish black, being a continuation of the ventral thoracic pleural stripe ; sternit.es
yellow, hypopygimn brownish yellow.
Hab. South India.

650

Dr. C. P. Alexander on

Holo~ype, ..~, Cinchona, AnamMt~i Hills, altitude 40005000 foot, September 17, 1946, (P. Susai Nathan).
Allotapotgpe, ~, J u l y 16, i946. Paratopotgpes , 7~',
August 29-September 24, 1946, (P. Susai Nathan).
The genus Eupilaria Alexander was originally proposed
as a subgenus of Pilaria Sintenis (Philippine Journ. Sci.,
48: 39; 1932), chiefly becuase of the conspicuously
verticillate antenna~. I now regard it as representing anentirely distinct genus. The earliest described species was
Eupilaria opaca (de Meijere), of Java. Three further
forms, E. auranticolor (Alexander), E. leucopoda (Alexander), and E. phoe~osoma (Alexander), are known fromthe Philippines, and E. albicans (Edwards), from northern
Borneo. Five further species are now known fromsouthern India, three being described at this time. The
present fly is close to E. annuIipes (Brunetti), of southwest India, having the leg pattern approximately thesame, but differing in the conspicuously patterned wings.
The Indian species m a y be separated by the following
key :-l. Thoracic pleura uniformly yellow . . . . . . . .
Thoracic p l e u r a s t r i p e d longitudinally w i t hb r o w n i s h black . . . . . . . . . . . . . . . . . . . . . .
2. Legs b r o w n i s h yellow . . . . . . . . . . . . . . . . . .
Legs black . . . . . . . . . . . . . . . . . . . . . . . . . . .
3. Wings u n p a t t e r n e d except for the s t i g m a . ,Wings with a conspicuous d~rk p a t t e r nadditional to the stigma . . . . . . . . . . . . . . .
4, F e m o r a with two b r o w n rings t h a t enclosea broader yellow one ; g r o u n d colour ofthoracic pleura yellow ; posterior p o r t i o nof vertex pale . . . . . . . . . . . . . . . . . . . . . . .
F e m o r a yellow, the tips n a r r o w l y black~ned, s u b t e r m i n a l dat~k ring l a c k i n g ;tbora~ie pleur~ and v e r t e x tmlformly
grey . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .

2.
3.
inconsequens(Brunetti).
8auvis. sp. n.
annul@es (Bmmetti).
4.

g~ttvlifera, sp. n.

incana, sp. n.

Eupilaria incana, sp. n.
Male.---Length about 6.5-7 mm. ; wing 7.2-8 ram. ;antenna about 1-1.1 ram.
Female.--Length about l0 ram. ; wing 9 ram.
Characters as in guttulifera, sp. n., differing as follows : Head above uniformly dark grey; rostrmn brown.
AntennM itagellum black. PreescutM disk with four
distinct brown stripes; postnotum and pleura distinctly
pruinose, in cases heavily so, completely obscuring theNew or little-known Tipulidse.

651

yellow. Legs with the ibmora yellow, ~he tips conspicuously blackened, but with no dark subterminal ring.
Wings with cell 1st M.~ relatively smM1, with m-cu slightly
nearer the base.
The grey coloration and the pattern of the femora
should readily separate this fly from g~ttulifera.
Hab. South India.
Holotype, ,~, Cinchona, AnamMai Hills, altitude 40005000 feet, September 16, 1946 (P. Susai Nathan,).
Allotoptype, 9, August, 27, 1946. Paratopotype, I c?,
August 29, 1946.

Eupilaria sauvis, sp. n.
Allied to inconsequens; mesonotum almost uniformly
eastaneous, the pleura and pelurotergite unitbrmly yellow ;legs black; wings fulvous brown, the stigma darker
brown ; Sc 1 ends only a short distance beyond the forkof Rs.
MaIe.--Length about 7-9 mm. ; wing 7.5-9 ram.
~¥male.--Length about 7 ram. ; wing 7 ram.
l%ostrum yellow; palpi black. Antennse brownish
black throughout. Head pale grey, the centre of vertex
extensively infuscated.
Pronotum brownish yellow. Mesonotum almost uniformly light brown or castaneous, unpatte~aed, thepleura and pleurotergite uniformly yellow. I-Ialteres
infuscated, the base of stem narrowly yellow. Legs withall coxse and trochanters yellow ; remainder of legs black,the femoral bases restrietedly brightened. Wings lightbrown to fulvous brown, the prearcular and costal fields
more yellowed ; no pattern except the oval brown stigma ;veins brown, somewhat lighter in the yellowed areas.
Venation : Sc relatively short, Sc~ ending a short distancebeyond the fork of Rs and distinctly before the fork of
R2+a+4, the latter subequal to or shorter than the basal
section of Rs; m-cu about two-thirds to three-fourths
its length beyond the fork of M.
Abdominal tergites brown to dark brown, darkest on thesubterminal segments, in cases weakly bicoloured ; sternites
yellow ; hypopygium brownish yellow.
Hab. South India.
Holotype, (~, Cinchona, Anamalai Hills, altitude 4000-5000 feet, August 29, 1946 (P. Susai Nathan).

652

Dr. C. P. Alexander on

Allotopol:qpe, 9, September 24, 1946. Paratol)otypes ,8everM c~, August 29-Septelnber 24, 1946.
Although close to E'~pilari(t inoons~¢g~tens (Brunetti), !
must regard tlte present fly as being distinct. The
smallest measurements given above are those of three
specimens that differ markedly from the larger type andvarious paratypes in this regard.
Hexatoma (E,rioeera) peenulatoides, sp. n.
Allied to p~nulata; size relatively smMt (wing, rome,
11 mm.); antenn~ (mMe) 8-segmented; mesonotum
plumbeous grey, the humerM region of W~eseutum obscureyellow with ~ velvety black spot at the eephMie portionof the interspace and a much larger more laterM one of thesame eolom' ; antennm black, the scape yellow ; pleura andpleurotergite yellow ; legs black, the femorM bases and all
eoxoe yellow; wings with a strong brownish tinge, thepresrealar and eostM borders more brownish yellow; Scvery long, Sc 1 ending shortly beyond the fork of R2+a+,,the latter abo~lt three times the basM section of R 5 ; m-cu
at near two-thirds the length of cell Ist M2; abdomen
opaqne orange, the basal tergite darkened medially above.
Male.--Length about 11 nnn. ; wing l 1 ram. ; antenna
about 2-2 ram.
Rostrum yellow; pslpi black. Antennze (male) 8-segmented, short, as shown by the measurements; scapelight yellow, pedicel and ttagellum brownish black;flagellar segments gradually decreasing in length outwardly, provided with long setee, the longest on flagellar
segment three a little shorter than the segment. Headblack.
Mesonotum chiefly brownish grey or plumbeous, thehumeral region obscure yellow, with a velvety black spotat eephalio portion of the interspace, with a second muchlarger circular black marginal spot opposite the lateral
stripes ; laterM borders of mediotergite and all of plenrotergite yellow. Pleura similarly yellow, with small inconspicuous darkenings' below and in front of wing root.
Halteres brownish black, the base of stem narrowly yellow.
Legs with eox~e and troehanters yellow ; femora black, thebases yellow, somewhat more conspicuous on the fore
pair ; tibim and tarsi black, Wings with a strong brownish

New or little&nown Tipulidw.

653

tinge~ the prearcular and costal ileitis more brownish
yellow : stigma exceedingly ~alTow ~O scarcely iudicated ;vague to scarcely evident darke~fings over the cord andouter ends of cell 1st M e ; veins brown, more yellowed in
the brightened portions. Abundant macrotrichia on veins
beyond cord, much more sparse basad of this point.
Venation : Sc very long, ending shortly beyond the lbrk
of Rz+.~+4, vein Ro oblique, its origin lying slightly moredistad than its cephalic portion, the vein subequal tovein/~1+~ ; R2+3+~ about three times the basal section of
R 5 ; cell 1st M 2 rectangular, with m-cu at about two-tl~irds
the length, the cell subequal in length to vein M a.
Abdomen almost uniformly opaque orange, the firsttergite infuscated medially above; on the individual
tergites the posterior third of the segments somewhat
more intensely ihlvous orange or brownish orange.
Hab. South India.
Holotype, ~, Cinchona, Anamalai Hills, altitude 40005000 feet, September 17, 1946 (P. Susai Nathan).
The most similar described species include Hexatoma(Erioce~'a) pwnulata (Enderlein), H. (E.) subpwnulata(Edwards) and H. (E.) rufiventris (Brunetti), all of which
are represented in m y collection. All differ in the larger
size and in distinct details of coloration and venation.

Hexatoma (Eriocera) anamaliana, sp. n.
Size relatively small (wing, female, 10 ram.) ; mesonotal
praescutum with three polished dark stripes, the interspaces concolorous, opaque; antennae (female) 7-segmented; head brownish black, more pruinose on front,
anterior vertex with a pair of small conical tubercles ; legs
black ; wings with a strong brownish suffusion, the basevaguely more yellowed; R2+.~+.~ about one-half longer thanthe basal section of R 5 ; cell M 1 lacking ; abdomen withbasal two segments obscure brownish yellow, the succeeding ones dark brown, subnitidons, the extreme posteriorborders of the segments obscure yellow ; genital shield(female) obscure orange.
FemaIe.--Length about 11 ram.; wing I0 ram.;antenna ~/bout ].6 ram.
l~ostrmn light brown; palpi dark brown. Antennm(female) short, 7-segmeuted; scape and pedicel (l~rl~
Ann, &Mag. Nat. Hist. Ser. 12. Vol. i.
45

654

Dr. C. P. Alexander onbrowl), flagellunl brownish black; first ttagellar segment
subequM to the combined segments two and three,stouter ; flagellar segment four a little longer than segment
three and nearly twice as long as five. Front pale brown,
prey pruinose; head above brownish black; anteriorvertex with a small but conspicuous tubercle oil eitherside, immediately behind the pruinose field.
Pronotum opaque dark brown. MesonotM pr~esoutum
with three polished dark brown stripes, the interspaoes
restriotedly opaque brown of the same eolour; seutal
lobes similarly polished brown; seutellum brownish
yellow, postnotum uniformly yellow ; vestiture of mesonoturn very sparse, of moderate length only. Pleura lightbrown, the dorsopleural region more brownish black.
I-Ialteres brown, the knobs darker brown. Legs with theeoxre light brown; troehanters yellow; remainder oflegs black, the femorM bases obscure yellow. Wings witha strong and almost uniform brown suffusion, the Wearoular field and cell Sc more yellowed ; no stigma ; veinsbrown, paler in the brightened fields. Maerotriehia onveins beyond cord, with a few others basad of this point.
Venation: Sc~ ending about opposite the fork of R s ;~+3+~ about one-hMf longer than the basal section of
R~ ; R2+:3and RI+~. subequM or the former slightly longer,
either one a trifle less than twice as long as vein t~ 2 ; cellM t lacking; m-cu about opposite midlength to cell
1st M 2.

Abdomen with the basal two segments obscure brownish
yellow, the succeeding segments dark brown with theextreme caudal borders obscure yellow; entire surface
subnitidous; genital shield obscure orange; valves ofovispositor elongate, cerci gently upeurved.
Hab. South India.
Holotype, ~_, Cinchona, AnamMai Hills, altitude 40005000 feet, September 17, 1946 (P. Susai Nathan).
By Edwards' key (1921) to the Old World species of thesubgenus, the present fly runs without serious complications to Hexatoma (Eriocera) caliginosa (Brunetti), withwhich it disagrees in many details of coloration, especiallyof the thorax and abdomen. The latter fly has the thorax
wholly black~ the abdomen black, patterned with yellow
on the third and fourth segments.

New or little-known Tipulid~e.

655

Hexatoma (Eriocera) pur,pu~nta, ~p. n.
General coloration of thoracic dorsum and abdomen
purplish black; head dull black; antennse very shortin both sexes; halteres and legs black; wings with astrong blackish suffusion, slightly paler on basal half;
Sc short, S q ending some distance before the fork of Rs ;vein 2nd A slightly concave.
Male.---Length about 9.5-11 ram. ; wing 8-9 ram. ;antenna about 1.1-1.2 ram.
Female.--Length about 11 mm. ; wing 10 ram.
Rostrum and palpi black. Antennae very short in both
sexes, black throughout ; in male, 7-segmented, the firstflagellar segment stout, about as long as the second andthird taken -together; fiagellar segments two to fivegradually decreasing in length and diameter, all provided
with conspicuous verticils; in female, antennm 8-segmented, the terminal one small. Head dull black, with
abundant long black set~e.
Thoracic dorsum pm'plish black, very sparsely pollinose, the vestiture long and black. Pleura black,pruinose, heavily so on the metapleura and pleurotergite.
Halteres short, black. Legs with the coxm black, pruinose; remainder of legs uniformly black. Wings witha strong blackish suffusion, on basal half somewhat moreyellowed, this especially evident in the female, including
the prearcular field and slightly beyond, as well as cell
C% ; veins brownish black, paler on proximal half of wing.
Veins beyond cord with abundant trichia, basad of cordthese becoming more sparse and finally lacking. Venation : Sc tmusually short, Sc, ending some distance before
fork of Rs, S% near its tip; R~+a+4 nearly twice R2+aand in direct alignment with i t ; RI+ ~ about one-halflonger than R~+a; cell M~ lacking, m short; m-ca atnear one-third to one-half the length of cell ]st M~. Ina single paratype, vein R., lies just before the fork of Ra+ ~.
Abdomen uniformly black, the segments more or lessshiny and with strong purplish reflexions that are scarcely
obscured by a sparse pollen. In cases, abdominal segments five to seven inclusive, in the male, expanded intoa spatula, this not apparent in other individuals that seem
evidently to be conspecific. In female, genital shieldblack, the long straight valves of the ovipositor oranue.
Hab. South india,
45 ~

656

Dr. C. P. Alexander on

ltolotype, ~, Cinchona, AnamMai Hills, altitude 40005000 feet, September ~0, 1946 (P. Susai Natlu~n).
Alloto2otype~ ~ !~, Septembcr 17, 1946. Paralopotypee, 4 ~ ,September 16-20, 1946.
In the short vein So, the present insect agrees with agroup of about ten species of the subgenus known from
Ceylon, all of-which are otherwise quite distinct flies.
However, in other venational characters it disagrees withthese flies and by Edwards' key (1921) to the Old World
species of Eriocera it runs to couplet 35 where it cammt be
traced beyond this point. Evidently the characters of a
spattflate abdomen, found in the males of a few regional
species, is not as constant a character as had formerly
been believed.
Hexatoma (Eriocera) susai-nathani, sp. n.
General coloration of thorax black, the pr~escutum withthree polished stripes ; antennae (roMe) elongate, approximately as long as body, basal three segments of flagellum
light yellow, the outer ones black; in female, antennm
uniformly blackend ; halteres and legs black, the femora]
bases obscure yellow ; wings brownish black, handsomely
variegated with pale yellow and white, the former including
the broad wing base, the latter a wide band before cord
and a subterminal row of spots, as well as a small areaat origin of Rs ; abdomen with segments one and six toeight black, two to five orange, male hypopygium yellow.
Male.--Length about 8-9 ram. ; wing 9-9.5 ram. ;antenna about 8.5-9 ram.
Female.--Length about 14-15 ram. ; wing 12--13 ram. ;antenna about 2.8- 3 ram.
l~ostrum brownish black ; palpi dark brown. Antennm(male) elongate, nearly equal in length to the wing,much shorter in the femMe, asshown by the measurements;
in male, scape and pedicel light chestnut, basal threesegments of flagellum light yellow, the remainder abruptly
brownish black ; 7-segmented, the last flagellar segment
very small; flagellar segments long-cylindricM, the firstand fourth longest, second and third slightly shorter andsubeqnal to one another ; flagellar segments with a dense
erect white pubescence that is nearly as long as thediameter of the segment ; basal flagellar segments with

New or little-kno'une Tipulidee.

657

relatively few short verticils, on the fourth segment
becoming much more numerous; in female, antennae
9-segmented, dark throughout. Head dull black.
Pronotum black, more pruinose on sides. Mesonotumwith the ground colour velvety black, the pr~escutumwith three polished black or plumbeous black stripes;
scutal lobes similarly polished. Pleura and pleurotergitc
black, more pruinose on the propleura and much of themesopleura, the anterior border blackened to form atransverse darkening ; meron blackened. Halteres black.
Legs with the cox~e grey pruinose ; trochanters brown;
remainder of legs black, the femoral bases obscure yellow,
somewhat less extensively so on the fore legs. Wings
brownish black, variegated with pale yellow and whiteor yellowish white; prearcular field and cells C to M
immediately beyond yellow; the white areas include a
broad band before cord, extending from R to the posterior
margin ; wing tip slightly paler than the remainder ofground, its inner portion with a broken band or series ofwhite spots in cells R 3 to M~ inclusive, more obscured
behind and the posterior one often lacking; a small
white spot at origin of Rs ; axilla whitened ; veins brown,
more yellowed in the pale fields. Macrotrichia on veins
beyond cord. Venation : Re+3+a and Re+3 subequal ; R e
shorter than Rl+e; cell M1 lacking; m-cu at nearmid]ength of cell 1st M e.
Abdomen with the first segment velvety black;segments two to five, inclusive, orange ; segments six tocight black ; hypopygium yellow ; none of the segmen~with polished rings ; in female, genital shield orangc.
Hab. South India.
Holotype, ~, Cinchona, Anamalai Hills, altitude 40005000 feet, September 17, 1946 (P. Susai _Nathan).
Allotopotype, ~, September 18, 1946.
Paratopotypes,
several 3~, August 27-October 2, 1946 ; taken on plants
near mountain torrent.
I take great pleasure in naming this very distinct fly for
Mr. P. Susai Nathan, to whom I am indebted for numerous
Tipulidm from South India. B y Edwards' key (1921) to
the Old World species of the subgenus the species runs to
Hexatoma (Eriocera) basilaris (Wiedemann), of Java~ which
has approximately the same wing pattern but differs inthe short antennm of both sexes and in details of coloration,

658

Dr. C. P. Alexander o~

All other generally similar East Indian species likewise
differ in the short antennee.

Elepha~gomyia (Elephantomyodes) a~fluens, sp. n.
General coloration of thorax dark grey, tile pr0escutumwith four dark brown stripes ; head dark grey ; rostrum
elongate, being approximately two-thirds the wing ; legs
black, the tarsi Mfitened before the tips; wings with aweak brownish tinge, the costal border narrowly andconspicuously dark brown; narrow brown seams overcord and the outer end of cells 1st M~.
Male.--Length, excluding rostrum, about 6.5-10 ram. ;wing 7-10 mm. ; rostrum about 4-6 ram.
t~ostrum black, elongate, approximately two-thirds aslong as the wing. Antennm black throughout ; flagellar
segments elongate-oval, with very elongate verticils.
Head dark grey, clearer grey on front; anterior vertex
moderately wide, a little exceeding the diameter of thescape.
Pronogum dark brown, sparsely pruinose. Mesonotum
dark grey, the prmseutum with four dark brown stripes,
the intermediate pair narrowly confluent in front, very
narrowly separated behind; scutellum obscure brownish
yellow to dark brown, bordered by pale; mediotergite
dark brown. Pleura and pleurotergite dark plumbeous
grey, restrictedly patterned with yellow, including thedorsal pleurotergite and the entire region of the metapleura. IlMteres dark brown, the base of the stem
restrictedly yellow. Legs with the cox~e brown; trochanters brownish yellow; femora, tibim and basitarsi
black, the tips of the latter narrowly snowy white;segments two and three white, four and five black. Wings
with a weak brownish tinge, the costal border narrowly
and conspicuously dark brown, occupying cells C and Sc
and continued outwardly about to the wing tip, involving
the outer half of cell R~; narrow brown seams overorigin of Rs and cord, more narrowly so at outer end ofcell 1st M 2 ; veins brown. Venation : Rs relatively long,
fully one-half longer than cell 1st M~, squ~re at origin;cell 1st 2~I~ abont equM in length to vein M4; m-cu atnear one-third the length of celt l st M:, : cell 2nd A narrow.
Abdomen relatively long, basal segments weakly
bicoloured, chiefly brownish black, the bases of each

New or little-known Tipulidse.

659

segment very restrictedly pale; outer segments more
uniformly blackened.
Hub. South India.
ttolotype, <~, Cinchona, Anama/ai Hills, altitude 40005000 feet, September 30, 1946 (P. Susai Nathan).
Paratopotypes, several c~c~,September 16-October 4, 1946.
Elephantomyia (Elephantomyodes ) ajfluens is very
different in all regards from the only other regional memberof the subgenus so far made known, E. (E.) aurantia(Brunetti), of Assam.
Riedelomyia chionopus, sp. n.
Mesonotum chiefly reddish brown, the prmscutum
narrowly margined with brownish black ; antennae black,the four-parted fusion segment yellow ; legs dark brown,
the outer tarsal segments snowy white ; wings brownish
yellow, with a heavy brown spotted pattern.
Female.~Length about 5.5 ram. ; wing 6 ram.
t~ostrum black, sparsely pruinose ; palpi black.
Antennae with the scape black, grey pruinose; pedicel
brownish black ; fusion segment of flagellum yellow, theremainder of organ brownish black, the segments withlong verticils ; fusion segments apparently invoNing four
segments, but with all sutures incomplete; terminal
flagellar segments broken.
Head brownish grey ;anterior vertex broad.
Pronotum buffy above. Mesonotum chiefly reddish
brown, the prsescutum narrowly margined with brownish
black and with faint indications of two paler brownintermediate stripes ; pseudosutural fovem large, pale ;tuberculate pits not evident ; posterior sclerites of notum
brown, more or less yellow pollinose, the scutellum
darkest. Pleura brownish yellow, conspicuously striped
longitudinally wi~h brownish black, the dorsal stripe
broader, extending from the cervical region across thedorsal plenrites to the anterolateral portions of theinediotergite; ventral stripe narrower, beginning more
faintly on the fbre cox,, becoming darker behind, crossing
the central portion of the sternopleurite, passing beneaththe halteres to the abdomen ; dorsopleural membrane buffy
yellow. Halteres infuscated, the base of stem narrowly
pale. Legs with the fore eox~e yellow basally, brown
beyond, us described ; remaining coxm and all trochanters

660

Dr. C. P. Alexander onpale yellow ; remMnder of legs dark brown, the distM fifthof basitarsi and remainder of tarsi white (terminal twosegments broken). Wings with the ground brownish
yellow, the costal border narrowly clearer yellow; arelatively heavy brown spotted pattern, as follows: Atand beyond arculus in bases of cells R and M ; origin ofRs, extending from vein R backward, not reaching vein
M ; a comparable large spot at tbrk of Rs, not or scarcely
connected with the seam along cord; outer end of cell
185 Me; stigma darker brown, relatively small; veins
beyond cord, especially near wing tip, narrowly seamedwith brown ; a somewhat larger area at end of vein 2nd A,
with a further infuscation in the wing axilta; veinsbrownish yellow, clearer in the, brightened po~"~ions, darker
in the clouded area. Venation: Sc~ ending opposite
fork of Rs, Sc~ far from its tip ; veins I£~ and R 5 slightly
approximated at outer ends so cell R~ is somewhat wider
at its base ; cell 1st M~ about as long as distal section ofvein Ma; m-cu from one-fourth to one-third its length
beyond the fork of M.
Abdominal tergites uniformly dark brown or brownish
black; sternites, especially the outer ones, somewhat
paler with darkened posterior borders ; ceroi only gently
curved, obscure yellow, darker basally ; hypovalv~e very
slender, darkened.
ttab. South India.
Holotype, ~, Cinchona, Anamalai Hills, altitude 40005000 feet, September 21, 1946 (P. Su~'ai Nathan).
P~t'ratopotfpe, ~_, September 2, t946.
The present fly is quite different from the two previously
described regional species, both of which have the tips ofthe femora narrowly snowy white. The genus still is
Wry insufficiently known. To the recorded genericcharacters may be added: Legs with long simple set~e
only, no scales. Merci region greatly reduced~ the middle
and hind cox~e nearly contiguous.

Rhabdomastix (Rhabdomastix) nilgirica, sp. n.
General coloration of thorax medium brown, sparsely
pruinose, the more ventral thoracic pleurites reddish
brown; antenn~ (male) greatly lengthened, exceeding
three times the length of the wing ; wings brownish grey,N e w or little-lcnown Tipulid~c.

661

the stigma and outer radial cells more darkened; veins
R:~ erect to virtually perpendicular ; abdomen bicoloured,
the segments obscure yellow with the caudal margins
broadly infuscated ; hypopygium brownish black.
M a l e . - - L e n g t h about 6 ram. ; wing 6-8 ram. ; antenna
about 25 ram.
l~ostrnm brown; palpi reduced, black. Antennae(male) greatly lengthened, as shown by the measurements,
exceeding three times the length of the wing; scapeenlarged, chestnut brown ; pedicel very smM1, testaceous ;flagellar segments yellow, the incisures narroMy inthscared ; outer segments more uniformly darkened ; flagellar segments elongate-cylindrical, with a short dense
pubescence on the outer face, this longer than the very
sparse darkened verticils. Head dark brown, grey
pruinose ; anterior vertex broad.
Pronotum brownish grey. Mesonotum almost uniformly medium brown, sparsely grey pruinose, thehumeral region of prmscugum yellow; pseudosuturat
fove0e and tuberculate pits Mack, conspicuous. Pleura
chiefly reddish brown, the dorsal pleurites and dorsopleural membrane darker brown. Halteres infuscated,the base of stem restrictedly yellow. Legs with the coxm
yellowish testaceous; remainder of legs obscure yellow
or brownish yellow ; fore and middle trochanters elongate,
exceeding the cox~e in length, posterior trochantersnormal. Wings brownish grey, cells C and Sc more
infuscated; outer radial cells conspicuously darkened,
involving'all of cell Ra and the outer half of R 4 ; stigmaoval, dark brown, separated from the outer radialdarkening by a pale spot before vein R a : cord and outerend of cell 1st M~ very narrowly seamed with dark brown ;veins brown, the Anal veins paler. Wings broad, anala @ e conspicuous. Venation : Sc 1 ending nearly opposite
five-sixths the length of Rs ; vein R:~ erect to virtuallyperpendicular, subuequM in length to the space on costa
between veins RI+ ~ and Ra; vein R 4 only slightly
arcuated, subequM to R.~+a+, ; m much longer than thebasal section of M a ; vein 2nd A sinuous.
Abdomen bicoloured, basal tergite brown; remaining
segments obscure yellow with their caudal margins broadly
infuscated ; hypopygium brownish black.
Hub. South India.

662

Dr. C. P. Alexander on

Holotype, c~, Singara, Nilgiri Hills, altitude 3400 feet,
May 21, 1948 (P. Susai Nathan).
This is the first record of occurrence of the genus from
India. From the other Oriental species having cell 1st M,~
closed, including Rhabdoma,~tix flavidula Edwards, ofBorneo, and R. trochanterata Edwards, of Pahang,Federated Malay States, the present fly is readily told b y
the coloration of the body and wings. The closest ally
and the only one of the two species above listed, that iscertainly a member of the typical subgenus, is t~'ochanterata.
Styringomyia thetis, sp. n.
General coloration yellow, the pronotum and lateralborders of prmscutum broadly bordered b y black;pleura and pteurotergite yellow ; fore and middle femora
yellow, each with two brown rings, posterior femora
yellow with a narrow incomplete subterminal brown
annulus ; wings yellow, unpatterned ; vein 2nd A simple ;male hypopygium with two unequal spines on basistyte ;dististytes complex, the intermediate and inner styles orarms with long black set~e.
Male.~Length about 7.5 ram. ; wing 4.8 ram.
Female.--Length about 6 ram. ; wing 5 ram.
Rostrum and palpi black. Antennse with scape andpedicel blnck; flagellum yellow, the outer segments
darker; in female the bases of the segments narrowly
clearer yellow to produce a weak bicoloured appearance.
Head testaceous brown above, darker brown on sides.
Pronotum and mesonotum obscure yellow dorsally, thelateral border continuously and conspicuously black, thepattern more expanded over the humeral region ; disk ofproescutum with two na~Tow blackish lines before suture,
narrowed in front ; posterior sclerites of notum somewhat
more pruinose and less evidently patterned with brown,
more distinctly so on the lateral borders of the scutellum
and mediotergite; set~e of mesonotum unmodified.
Pleura and pleurotergite yellow. Halteres with stem
pale, knob weakly darkened. Legs with all cox~e andtrochanters yellow; femora yellow, the fore and middle
pairs with two dark rings, the outer one darker, about
one-half its own length t}om the tip, narrower than the

New or little-known ~Iipulid~e.

663

yellow subterminal annulus; tibize yellow, with twoalternate black rings, the tip black, the dark and lightrings subequal in extent, only the basal yellow one
narrower; basitarsi yellow, the tips blackened, more
broadly so on the fore legs, remainder of tarsi black;posterior femora yellow with a single narrow and in~complete dark subterminal spot; tibim yellow, the tip
narrowly darkened; basal three tarsal segments yellow,
their tips narrowly' infnscated, tile terminal segments
darkened. Wings yellow, unpatterned; veins brownish.
yellow; costal fringe relatively long. Venation: Cell
2nd M e narrowly sessile; m-cu at near one-third thelength of cell 1st M~ ; vein 2nd A simple.
Abdominal tergites with central portion almost uniformly pale to medium brown, more yellowed laterally;sternites and hypopygium yellow. Male hypopygiumwith the apical lobe of the ninth tergite hemispherical,
transverse, the margin fringed with abundant yellow
setm ; ninth sternite narrowed to the truncated tip whichbears two elongate unmodified setse. Basistyle with theouter lobe bispinous, the spines unequal, the longest at
apex of a tubercle, the second about one-half as long,subapicM. Outer dististyle or arm long and pale, with
two apical setse, one very long: intermediate style anarrow flattened blade that is densely covered with tong
black setse ; inner style or branch triangular in outline,
narrowed outwardly, at tip with a strong blackened
spine, the surface with abmldant black setse, much as in
the case of the intermediate style: at extreme base ofstyle, on mesal face, with a slender curved arm that
ternfinates in a long black spine of nearly equal !et,*gtt.t ;at base of this arm with a comb of long black spi,les.
Phallosome with very sparse darkened areas or points.
Hub. South India.
Holotyl)e, ~, Cinchona, Anamalai Hills. altitude 40005000 feet, September 17, 1946 (P. Susai Nathan).
Allot potypc, ~, in copnta with the type.
The most similar regional species is Ntyri'ngomyia/cempiana Alex~mder, which differs in the coloration of
the body and legs and in the details of structure of themale hypopygium, particularly the basistyle and in alldetails of structure of the dististyles.