.

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Wirh fip Autlnr,s Compl,i,mink
Fnou Rrc. Ceur. Mus., Vor. IV, No. 5
PP.219-2n. PrArE XXUII
9fir Mencr, lg3g,

NeWor Little-known
TipulidaefromNew Zealand(Otder Diptera)
PartIII.
Bv Charles P. Alexander, Amherst,
Massachusetts,[.J.S.A.

CHRISTCHURCH,N.Z.
Pnrxtso Bv rHE Cnnrsrcnuncn Pnsss Couprxy LriltrED

r939

:

NEWOR LITTLE-KNOWNTIPULIDAEFROM
NEWZEALAND (ORDERDIPTERA)PARTIII.
tsy cuenrBs P. ATBxaNDER,Amherst, Massachusetts,U.S.A.
The -precedi.ng
parts under this general title were published in
r93o and rg3z.* The speciesdiscussedat this time^are chieflv
containedin the collectionof the Canterbury
Museum,having been
-Othercollectedby Nlessrs.Lindsay anq Tonnoir.
importaniseries
were from the Hen and ChickenIslands,North Aucklind, collected
October -t_925:_by
Hamilton, and from Stewart Island,_n_Ir._Harold
collected9v.Mrs. M. Jensenand her son, Norman J.Jensen, these
two lots being preserved in my collection of Tipulidae throughthe friendly interest of the personsnamed.
f)'-r'oluminous notes on the distribution of the Tipulidae
of New Zealandarc containedsome thousandsof record.sthat well
warrant publication if the geographicaland seasonaldistributionof theseflies is to be better understoodby Dipterologists. At thistime I am publishing lists of the speciesthaf bear on trvo of thelocalities further mentioned in the report.
Hen Island (Hen and Chicken Islands), North Aucklancl,
collectedOct. '25,rgz1 by IIr. Harold Hamilton.
H olorwsia(Zelnndotipu,Ia)
fulaa (Hutton).
L'Iacromastix(Chlorotipula)holoclclora(Nowicky).
Dolichopeza(Dolichopeza)paraicaudaEdwards.
Linr,onia(Discobola)ampla (Hutton).
Limonia (Discobola)
gibbera(Edwards).
Limonia (Discobola)
tesselata(OstenSacken).
Limonia (I dioglochina)
fumipennis (I3utler).
Limonia (Dicranomyia)multispina (Alexander).
Limonia (Dicranomyia)repanda (Edrvards).
Austrolirnnophilacrassipes (Hutton) .
A ntp hirceuruts (A mpItin,eu,nt,s)hudsoni Edwards.
Amphirteurus(I{othormosia)edenhilussp. n.
Erioptera (Empeda)hamiltoni sp. n.
* Alexander,- C. P. ' New or little-lrnown Tipulidae from Ncw ZcalanJ(Order Diptera) . Rec. Cant. lVlu,s.,4' 281-2tttl; 1930.
T h e s a m e ,P a r t I f . I b i d . , 4 : 459-+67; L932.

220

ALEXANDER

Cass,Canterbury,altitud€ r,85o feet, collectionsmadein rgz4
and rgz1 by Dr. Andr6 L. Tonnoir.*
Holorusia (Zetandotipula)fu,laa (Hutton); Nor'. 30, Ig24; F-eb.
r925
Alexander;Nov. 3o, 1924.
grey(uta,
IIacrimastix (X[acrontastix)
L'Iacromastix(M acrornastix)rniruttissimaAlexander; February
r925'
J,Iacromastix (I[ acromastix) rufiuentris Edwards; February
r925'
M acrirnastix (II acromastix)uu,lpinaHutton; February 1925.
paraicau'daEdwards;Nov. 3o, 1924.
(Dolichopeza)
Dotichol>eze
l.irnonia (Zetandoglocltina)
flauidipennis(Edwardt); treb. 1925.
(F-dwardt); Nov. 30,
su,blacteata
Lirnoniu. (Zctandoglochina)rgz4,Iieb. r925.
Linconia (Dicranomyia)hernimelas(Alexander);Feb. 1925.
Limonia (Dicranomyia)Icudsoni(Edrvardt);Feb. 1925.
Limonia (Dicranomyia)h'tteipes(Alexander);Feb. rg25; Nov.
2 8 ,1 9 2 1 .
Liytortia (Dicranontyia)pen'dulifera(Alexander);Nov. 27, 1924'(Edrvardt); Feb. I925.
Limonia (Dicranontyia)tenebrosa.
Limonia (Dicrarcomyia)aicarians (Schiner);Feb. 1925.
Au,strolint'nopltilahdsoroi (Edwardt); Feb. 1925.
(Edwardt); Nov. 3o, 1924.
A1strolimnophilarcigrocitccta
Ii otholimnofhila exclu,sa(Alexander); F-eb. 1925.
Iletalirmrcl>hilapalmala Alexander; F'eb. I8, I925.
(Alexander); Dec. I, T924.
u,nil>u,u,cta
AI etaliru,ttol>lila
pygmaeaAlexander; Feb' 1925'
Zelan'donr,yia
Gynoplistia (Paratimnophila) sku,sei,(Hutton); Feb. 1925.
Gynoptistia(Gyttoplistia)ntagni,ficaE dwards; Iieb. 1925.
Gynol>Iistia(G3trtoplistia)nebu,losaEdwards; Feb. 1925.
GonLrttyict(Lipophtcps\oliueri Alexander; Nov. 30, 1924, Feb'
r925'
(Eclrvardt); Feb. 1925.
^4phrophitaneozelarr,dica
Rhabdornastix(sacandeg&)callosaAlexander; Nov. 30, L924,
Feb. r925.
Rhabdornastix (Sacandaga) otagana u,nilineata sub. sp. n.;
Nov. 30, 1924.

otaganatrilineala sub' sp' n' ; Feb.
(Sacandage)
Rhabd,omastix
r925'"Notes from the Canterbury College lVlountain*'fhe various articles entitled-should be consulted.
Biological Station,"-itt T'rnn's'^l'Z' [ns[."

NEW OR LITTLE.KNOWN TIPULIDAE FROM N.Z.

221

Erioptern(Trimicra)confhwrs(Alexander);treb. rgzs.
Anophineunts(Amj>hineunts)
senetcAlexancler;Dec. r, r 9 2 1 ,F e b .r g z ; .
Il[olophilusbasispinaAlexander;Nov. 27, rg24,Feb. rg, rgz5.
Molophifus.flagelliferAlexander;Nov. 30, rgz.+,Decemberr,r94.
Il[olophilu,sgladiatorsp.n.; Feb. r8, rg2;.

XIolophilu,sloileipygrtsAlexander; Feb. rgz5.
Ilfolophihrc nroros?ts
Alexander; treb. 18, rg,z;.
Molophilus rnulticintus Edrvards; Feb. rgz;.
trIolophiluspictrtratotsAlexander;f)ec. r, rgzl.
Molophilus pulcherrirnus Eclw'ards;I.-eb. rg2;.
Il[olophilus stewarterslisAlexander; Nor'. ,27, rgzl, Feb. rg2S.
Il[olopltilu,ssublateralisAlexancler;Nov. 30, r g,2,+.
Iltoloplti,lrtssttbmoros?ts
Alexander; Nov. z(), rg24.
Tasiocera (Tasioceru)diuaricata Alerander; Nov. zg, rgz4,
Feb. 18, rgz1.

M.T C R OI,T A S T IX (M.I,CNOM ASTIX) ENUNSTRIS SP. N.

Subapterousin both se\es, the rvings being shorter than thehalteres; antennae short, g-:segmentecl;
rnesonotal praescltum
flattened,u'ith four brou'nishblaCkstripes;legsverlr long, especialll,
the tarsi.
x{ale-Length about rs mm.; wing, z.s mm.; antenna about
2 . 3m m .
Fore l.g, femora, r2.S mm.; tibia, r4.5 mm.; tarsus
about 24 mm.

lliddle lee, femora, r-+.r mm.; tibia, r3.8 mm.; tarsus
about z8 mm.
Hind leg,femora,16 mm.; tibia, 16.5mm.; tarsusabout
38 mm.
Frontal prolongationof headmoderatelyelongate,dark brorvn
above,paler laterally, without a distinct nasus;paipi black. Antennaeshort, as shorvnby the measurements,
dark brown throughou!; f]agel-lumonly.z-se-gmented;
sgap9
flagellarsegments- elongate;"tocylindrical, gradually decreasingin length from the firsthefifth, the sixth J9"S9r,_ab9uj
equalto the third, the terminalsegment
again short. Head dark brown, the anterior orbits broadlirrich
chestnut-brorvn,rvider and more conspicuousat thc narrbwest
point oj t[e vertex; posteriororbits narro\\'ly pruinose;no evident
vertical tubercle.

222

ALEXANDER

Mesonotumstrongly flattened, flush rvith the level of the very
narrow, plate-like pronotum, the latter blackened. Praescutum
light yeilbu'ish gray, r,vith four velvety brorvnish-blackstripes, the6roadly separated by a reddish line; lateral andinlermediate pair^intermediate stripes subconfluent; praescutum clothed withrelatively long bu-t subappressedsetae; scutum. yellorv, each lobe
largely brorvnish black; siutellum lar-ge,blackish, -with a yellow
poiiinbsity; postnotum relatively reduced, graf ish bror,vn, thepleurote.fite^with a vague pale stripe. Plgrya light brou'n, varielated rn'it-hpatchesof darkei brown and whitish. Haltereslong andilenclcr, claik brown, the knobs paler bror,vn. Legs very long andfore
slender, especiallythe tarsi, as ihorvn b.v the -measurements;
femora shorter ihan tibiae; middle femora longer than tibiae;
femora and tibiae of hind legs nearlv equal; coxaeelongate,brorvn;
trochanters browtt, darker bepeath; femora dark bror'vn, a little
paler basally, the tibiae conspicttouslyyellorvish brorvn, the tips
narron'l)riniuscatecl;tarsi pale brown, passinginto darker. Wings
very rehucecl,shorter than the halteres,strongly- infuscated, theveiis blackish; costal margin u'ith strong dense black trichia on
basal tu'o-thirds;a large patch of trichia in the radiat field. Venation greatl5rdistorted.
Abclomen light bron'n, the tergites u'ith darker median andlateral stripes that are narrorvll' connectedu'ith one another acrossthc terrninal segmentsmore extensivelydarkthe caudal^margins,
ened; sternites paler.
Habitat.-Neu' Zealand(Stewart Island).
Holot5'pe, $, Tlalfmoon B"y, altitude 6o feet, Jnly 6, 1924
(N. J. Jensen).
Paratopotypes,a ferv additional males at subsequentdates.
This is the first speciesof.IIIacromastixto be describedin rvhich
the male sex is nearly apterous. The fly is so djfferent from allhitherto describedspeciesthat it is difficult to point out affinities.
sunvlrrATA sp. n.
MacnouASTIX(MecnoMASTIx)
Male.-Length about ro mm.; rving, 16 mm.
Fore leg, femora,8 mm.; tibia, 9.4 mm.;tarsus, t8 mm.
Hind l.g, femora, g.2 mm.; tibia, ro.8 mm.; tarsus,
2 3 . 5m m .
Bearing a resemblanceto M . (M.) aittata Edwards, differing in
the body-colorationand details of structure.
Frontal prolongation of head dark Brayr the nas.usconspicuous.
Head light gray, the vertex largely occupi_ed!y u triangular brownish black area that restricts the ground-color to rather narrow

NEW OR LITTLE.KNOWN TIPULIDAE FROI\I N.Z.

223

orbital lines, the dark area further divided bv a scarcelv evident
capillary pale vitta. Anterior vertex broad,fuliy one-haltinc rviclthof head, including the eyes. Antennae short, r2-segmented,theterminal three segmentsvery small, shorter than the preceding
two taken together.
Mesonotal praescutum blue-gray, with a faint bluish tinge,
with four conspicuous
black stripes,the intermediatepair separated
by a narro\\'but conspicuousline of thc ground-color;lateral praescutal stripes interrupted at the suture; scutnm gray, the lateral
portions of the scutal lobes much paler; each scutal lobe with asmall cephalic and a very largc posterior black mark, cntirely
confluent,the marks further confluentposteriorlyacrossthc meclianline; scutellum infuscated;mecliotergitedark gray, with a vague
circular dark mark on either sicleof the rnedianline bchincl. Pleura
clear bltte-gray,the r.entral half of the pleurotergitepaler; dorsopleural meml-rranepalc brow'nish ochreous; a circular velvettr
black spot otr extremedorsalmargin on ancpisternumanclanothernear the positionof the fusion of thc sternopleuriteanclanepisternunl, the suture betu'eenthe latter tu'o entirelSz
lacking. Halteresdark brown, the extreme baseof stem obscureyellou,. Legs withthe coxae concolorousrvith pleura; trochanters brolvnish yellou';femora brou'nish yellorv, the tips conspicuousll'blackened; remainder of legs bror,r'nishl>lack, the tibial basesbroadly paler;
tarsi nearly trvice the length of thc tibiae. Wings almost as inaittata,with the pale prearcttlanregion and very narrow cell znd A ,but differing as follorvs:Dark marking in cell R just before originof Rs continued caudad acrosscell tr'f, interrupting the extensive
rvhite marking in that cell and breaking the vittate appearance;
pale areasin basesof cells ft, and R, less evident.
Abdomen with light pruinositv on segmenttrn'oand the lateral
portions of the tergites beyond the third; abdomenorange-yellou',on the basal ring of tergite two completely obscuredby whitish
pruinosity, darker orange on the posterior ring and on tergite
three; a continuous,black, dorso-median,tergal stripe, broadest
on segment two, suddenly narrowed there at the level of theimpressedfoveae,thence very gradually widened behind, coveringmuch of the posterior segments. Male hypopygium small, darl<.
colored,telescoped.
Habitat.-Nerv Zealand(South Island).
Holotype, d, Lake Brunner, Westland, February 2, rgzz
(A. L. Tonnoir);Collector'sNo. zr.
Ill[acromastix (M acromastix) softaittata agrees rvith ll/t. (lW.)
aittata Edwards in the very narrow cell znd A of the wings and in
the general wing pattern. It differs in the details of coloration,

224

ALEXANDER

especiallythe head and pleural pattern, the differentiatedpruinosity on certain of the abdominal segments,and in structure,especiallyof the legs,rvherethe tarsi are approximatelytwice thelength of the tibiae.
subsp. n.
KUMARENSIS
GvxoplrsrrA (P-l,neurrNopHrra) sKLTSEIGenerally similar to typical skusei,differing as follows:
Basal f lagellar segments with thc incisures conspicuously
yellow, especiallythe apical portions of the segments. Humeral
region of praescutumsuffusedr,r'ithreddish. \\/ing-pattern strongly
suffused, thc dark areas in the radial field solid, not horseshoeshaped;pale areasbeyond the cord restricted to a small post-stigmal
and a subequal nearly apical spot; remaining cells clf wing suffused.
Male hypopygium with the outer dististyleslender,the apex slightlv
hooked. Inner dististylc narrow()rthan in the typical form.
Habitat.-Nerv Zealand (South Island).
Holotype, d, Iiumara, Westland, October 2{'t, 1929 (J W.
Campbell);in m5' collection.
G. (P.) skusei (Hutton) is either highly variable, as indicatedby Edrvards (Trans. Ii.Z. Inst., 51i 3o7i rg'?3)or there are s(iveral
distinct races,or perhapscloselyallied species,confused. It seems
desirableto differentiate such forms so thel' may be discussedin
geographicalor ecologicaldistribution.
sp. n.
(GvNoertsrta) PHILPoTTI
GvNopLISTIA
General coloration of thorax greenish black; antennae r6-segmented, with eleven branched segments;femora yellow, the distal
fourth or less blackened; posterior tarsi yellorv; rvings with thebasal third pale, the remainder almost uniformly darkened.
Male.-Length about 8 mm.; rving, 7.2 mm.i posterior basit a r s u s ,r . 5 m m .
Rostrum black, with long yellow setae;palpi black. Antennaeb l a c k t h r o u g h o u t , l 6 - s e g m e n t e dt,h e f o r m u l a b e i n g 2 + 2 + 9 + : ;longest branch (about flagellar segment four) less than one-half thelen[th of the flagellum and approximately seven or eight times thelen[th of the segmentalone; last branch subequalin length to thesegment;terminal segment elongate,nearly !r1i-cethe length of thepenultimate and lar[er in diarneter. Head black, with purplish
reflections.
Mesonotum polished black, with vague greenish tints. Pleura
Halteres pale yellorv, the base of stemwith a heavy yellow pollen.
narrowly blickened. Legs with the coxae and trochanters black;femora yellow, the apicaf fourth or less blackened;tibiae and tarsi

NEIV OR LITTLE-KNOWNTIPULIDAE

ITROII N.Z.

225

black, thg posterior_tarsi obscure yellow with only the terminal
segmentrt_arkened.
Wings (Pl. XXVIII, fig. r) with ine basal thircl
pale, tlt. distal trvo-thirdsalmost uniforrnl-ydark brown, especiallybeyond_thccord; cell Sc,stigma."lq a seamon anterior corclihgntt!'darker browtt; v€ins brown. Trichiation of veins as in tha uio"lotto
SrguP:.1/ being.u'ithout trichia. Venation: Cell R, sessile;courseof
vein Rp ,n B.as in G. aiolacea.persimilis,that is,-parallelrvith R4,
beyondthe level of R, with veinsR, and Rnmore strongly diverging.
Abdomen violaceousblack. Male hypopygium much as inuiolacea.
Habitat.-New Zealand(South Island).
Holotype, d, Itotoroa, Nelson,Decernberz, rg,z5(A. philpott).
Namedin honor of the late NIr.Alfleq Philpott, whoseuntimely
death \\'as a.grga!]gr: to New ZealandEntomology. The specieJ,
together s'ith G (G.),harniltoni Llexanderand G.\G.) aiolaceaEd,\l'ards,form a natural group that is characterizedby the violaceouscoloration of the abdomen, the extensively yellow posterior tarsi
and the biramous-gonapop_hyses.The present fly is told by the
reducednttmberof antennalsegments,u'ith proporiionately
shorter-andbranches,thepatternof the u'ings,
the uniformly.peculiar
pale posterior basitarsi.
ItHesnoMASTrx (SacaNnace) oTAGANAuNTLTNEATA
subsp. n.

As in the typical form but the mesonotalpracscutumwith abroad, dark brown, median stripe extendingirom the cephalicrnargin of the sclerite,ending abruptly a short distancebefoie thesuture.
Habitat.-Nerv Zealand(South Island).
Holotypg, ?, cass, canterbury, altitude r,85o feet, November
Jo, 1924(A. L. Tonnoir).
RHesDoxIAsrIx (SeceNlaca) oTAGANATRTLTNEATA
subsp. n.

.\s in the typical form but the praescutum rvith three dark
brorvn stripes, the lateral pair crossingtne suture and extensively
suffusingthe scutal lobes,the median area of the scutum broadly
pale; sternopleuritemore or less infuscated.
Habitat.-New Zealand(South Island).

ryglojypf, 6, Cass,.Canterbury,altitude r,85o feet, February
rgz| (A. L. Tonnoir). Allotopotype,Q. Paratopotype,6.
In general appearance,the various forms of Rhabdomastix
otaganadiffer greatly from oneanotherand their exactrelationships
are still not well understood.

226

ALEXANDER

AupstNE,uRUS(NoruoRMosIA)roeNruLUS sp. n.

Size large (wing about 7 --.); male hyPopl'giuT with the
middle branch of the tripartite dististyle nearly smooth.
Male.-Length about 5.5 mm.; u'ing, 7 mm.
Female.-Length about 6 mm.; wing, 7 mm.
Rostrum yellow; palpi brownish black. AntennSe-(d) elongate,
as in the subgenus; flagellar segments weakly bicolored, black,the incisures cbnspicuously yellorn',the degree of the latter more
restricted on outer segments.
Mesonotum reddish brorvn, the humeral triangles yellow.
Pleura reddish brorvn, the dorsopleuralregion dusky. Legs brownish
yellow,the terminal tarsalsegmcntsinfuscated.!-ittg;_fl. XXVIII,
iig. z) almost as rn insulzrs.Male hypopygiuq (P-1.X{YII-I, 49. 5)."ith'the blackenedlobes of the tergite, 9t, slender,slightly divernotch. All branchet ol the tripartite
gent, separatedby a V-shap_ed
5uter dististyle iale in color, slender, the inner branch nearlystraight, the cenlral branch slender, sinuous, the margin nearlymerely microscopically denticulate, not conspicuously
r*oith'ot
Iuner dististyle, id, pale, bifid at apex into an
toothed as in in,stilu,s.
p, longobtusepale lobe and a small blackcnedspine.Gonapophyses,
and slender,not conspicuouslydilated at base as in insttlsus,very
gradually narrowed [o the acute blackened tigs. The elongate
pale outer lobe of the basistyle is not shown in the figure'
Habitat.-New Zealand (North Island).
Holotype, 6, Hen Island, Hen and Chicken Islands, North
Auckland,-October25, rg25 (Harold Hamilton). Allotopotype, 9.
is closestto A. insulstas(Hutton), differAmphirleulccsedenhtltas
ing especially in the smaller size and structure of the malehypopygium.
lrnropruRA (Itltprnn)

HIIIILTONI sp. n.

Generalcoloration dark brown; u,ingsrvith a faint brown tinge;
cell rst M2 closed; male hypopygium rvith the outer dististyle
deeply bifid, the arms expanded into blades; gonapophysestermina{ing in acute sPinoushooks.
Male.-Length about 3.2 mm.; wiug, 4 mm.
F'emale.-Length about S.+mm.; rviug, S.8-+ mm.
Rostrum and palpi dark brown. Antennac relatively .short,dark browl throughout; flagellar segments oval, the verticils (d)relatively elongate. Head brownish black.
Pronotum dark brown, the lateral pretergites testaceous.
Mesonotal praescutum dark brown, the humeral region ,paler;scutal lobes^darker;remainder of mesonotum dark brown. Pleura

NEW OR LITTLE.KNOWN TIPULIDAE FROM N.Z.

227

brown, the dorsal pleurites darker brown. Halteres brown, theextremebaseof stem paler. Legs with the coxaeand trochantersbrownishtestaceous;remainderof legsbrown, the tarsi only a little
d-arker;.t:gr_l:leti:glf tonq and slenderfor an E riopt erine ciane-fly.
Wings (Pl. {xVIII., fig: 3) with a faint brownishtinge,
the stigma-ending
scarcelydarker; veins darker brown. Venation: Sc,
about
gPpositetwo-thirdsthe length of Rs, Sc, a little moie than-itsown
length from tnttip; Rs nearly straight; R, + s + n and R, subequal;ve.r.ns
Ru _an{ Rn-Bglerally parallef to one another, celi Ro d..p;cell tst XI, closed;distal sectionof vein Ai[o strongly sinuoui; rn-cLt,
closeto for, of.II ; vein znd A nearly straight.
Abdomendark brorvn.I\Ialehypopygium(Pl. xxvIII, fig. 6)
with the bastistyle,b, relativelyshort,af apexrvith two smalllJbes.
Outer dististyle,od, very slenderat base,fhe apicalhalf bifid.,eacharm widely^expanded
in_topaleblades.Inner disiistyle,id, a slend.er,
nearly straigh.tt9g. Gonapophyses,p, flattened, terminating insinuouschitinizedhooks,theif trps acute.
Habitat.-New Zealand(North Island).
Holotype, 6, Hen Island, Hen and chicken Islands, North
Auckland,October25, rgzs (Harold Hamilton). Allotopotype,?.
Paratopotype,9.
I take un_usualp_leasurein naming this distinct speciesin
honor of the collector,Mr. Harold Hamilton. Additional specimens
tre prgservedin the Dominion Museum,Wellington. The subgenus
Empedahad not been recordedfrom the AustralasianRegion.
Moropurlus spErcHTr
sp. n.
Belongs to the plagiatws Broup; general coloration darkbrownishgray; ant_epnae
short; halteresdusky, the knobsabruptly
light yellgw; _legsblackened,the femoral basesrestrictedly fald;wingswith a brownishtinge, the costalregionbrighter; vein zicae
lorgi abdomen, including hypopygium, dark brownish grali
basaldististylea simple blackenedrod, the apical fourth a slend6r
glabrousspine,the remainderwith sparsesetulae.
Male.-Length about 2.7-2.8 mm.; wing, 4-4.2 mm.
Female.-Length about S.5mm.; wing, 4 mm.
Rostrum dark gray; palpi black. Antennae short. black
throughgut; flagellars€tments oval, with lgng verticils that greatly
exceedthe segmentsin length. Head dark brownishgray.Mesonotumdark brownish.gray, variegatedonly by the conlpicuous yellow a4terior_l-at9ra!prelergites and a- similar spot^tn.
beneath the -wing-root._Pletra dark gray. Halteres dusky,
knobsabruptly light yellow. Legswith the coxaebrown; trochanters
yellowish brown; remainder of legs blackened,the femoral bases

228

ALEXANDER

restrictedlyobscureyellorv. Wings rvith a brow"i.! .tinge;-prearcular and- costal regions more yellowistr;macrotichia and veinsdark. Venation: R, * , angulatedand sometimesrveaklyspurredat near midlength; vein znd A long, extendingabout to opposite
tr,r'o-thirdsthe length of the petiole of cell III .
Abdomen blackish gray throughout. Male hypopygium (Ill.
XXVIII, fig. 7) with the-apicalbeak of the bastistyle,b,.relatively
stout and strongtydecurved.Basal dististvle, bd,,a simplestraigh.t
rod, moderatelyblackenedthroughoutits length, the distal fourtha simpleacuteglabrousspine;cephalicor mesalmargil of stvle onrvith a seriesof four or five small setaefrom
basal^three-fouiths
notch-likeincisionsof the style.
Habitat.-New Zealand(SouthIsland).
feet,
Holotype, 3, Mount Peel, Canterbury, altitude 3_,_ooo
No-vember
November-i7, I93o (Stuart Lindsay). Allotopotyper_Q,
r.4, r93o. Paratopotype,rJ, altitude 3,5oofeet, November 14,
r93o;paratypes,z$$, PeelForest,Novemberr 5-2o,r93o.
I am very pleasedto name this interesting speciesin honorof my long-time friend, Mr. R. Speight, former curator of theCanterbury l\fuseum,to whom I expressmy 3,eepindebtedness
for many f-avorsduring the past eighteellyears-..Th. species-belongs
to a seition of the gioup having the basal dististyle of tt-re-male
hypopygium a simple slender rod. In the New Zealandfauna
there iie about a score of speciesin this section,these di{ering
amongthemselvesin the colorationof the body, legs"tt+ halteres;wing-dolorationand pattern; relative length of .vein znd A of thewin[s, and the structure of the male hypopygium, especiallythe
shapeand armatureof the basaldististyle. T!. present{I-yis mostneaily allied to MotophilusanalisAlexander,Il[. morosusAlexander,
Aleiander and M. syluicolusAlexander, differing
M. s-ubmorosus
in the uniformly dark coloration,which is so deep 1: t9 appear
almost blackish, and in the structure of the basal dististyle, as
described.
N I o T o p H I L U SG L A D I A T o Rs p . n .

group;dark pleuralstripe yely distinct;
Belongsto the plagiatu.s
wings not conspicuouilyvariegatedwith patchesof dark setae;
malE hypopygium rvith the basal dististyle.very long, slender,
gently irir"ed to the acute tip, the marginsrvith appressedspines.
IVlale.-Lengthabout 2.6 mm.; wing, 3.2-3.4 mm.
Female.-Length about 2.8 mm.; wing, 3.3 mm.
Rostrum and palpi dark brown. Antennae relatively short,pale, the basal segmg-nFslightly darker, testaceousbrown; flagillat segmentswitli relatively elongateverticils. Head light brown.

NE.W OR LITTLE.KNOWN TIPULIDAE FROM N.Z.

229

I\Iesonotalpraescutum pale brown to yellowish brown, thehumeral region and lateral margins obscurd yeilow; scutal iobes
brown, the median area somewhat paler; scutellum brownish
yellow; postnotum dark brown. Pleurb brownish yellow, with avery broad. and conspicuous, dark brown, longiludinal stripe
occupying the dorsopleural Legionand dorsal pleulites, extending
caudad to the postnotum. Halteresdark brow-n,the baseof stem
a little paler. Legs with the coxae and trochanters vellowish
testaceous;femora yellow, lvith darker trichia, the fore femoran'ith the latter lgttg and black; tibiae and tarsi more yellowish,
the terminaltarsalsegments
passinginto brown. \Vings(PI."XXVIII:
lg. +) rvith a grayishtinge,cloudedwith duskl' alongfRs, Ctcand theAnal angle; macrotrichia darker brown; veins brdwn. Venation:
Vein znd.4 endingabout oppositethe caudalend of.m-cu.
Abdomen dark brown, lhe trypopygium obscureyellow. Male
hygopy'gium(Pt X-XVIII, fig. 8; *ittr ihe apical beal of the basistyle, b, heavily bla_cketr9d,-relatively
deep al base,the apical half
slender and djrected dephalad.Outer dististyle, od, with^the stem
long ttt.dslender,-th"apgx bifid, as in the grotip. Basaldististyle, bd,
a very long, slender,chitinizedrod, gently curved to the ac,tt" tip,
the margins-yith conspicuousappressedspines.Aedeagusslender,
about four-fifths as long as the 6asal dististyle.
Habitat.-New Zealand(South Island).
Holotype_,4 cass, cqtgrbury, altitude r,85o feet, February
t8, rg25 (A. L..TolLgit). tllotopotype, p. paratopotypes,S dd;
paratypes,19.d_d,Hilltop,. Banks Peninsula,Canterbuiy, January
14-16,rg2S(A. L. Tonnoir).
. . This .fly is distinguishedfrom allied forms by the structureof the male hypopygium.
MoropHrLUSTMpRocERUS
sp. n.

.plagiatusgroup; mesonotum pale brownish. Belongs to theochreous to testaceous,the dorsal pleurites darh brown; maleh.ylgnygi-um with the basal dististyl-e straight, the apical third
blackened,expandedinto a rounded head.
Male.-Length about 2.6 mm.; wing, 3.8 mm.
Rostrum brownish testaceous;palpi dark brown. Antennae
short, dark brown; flag_ellarsegments-with relatively long and.
conspicuousverticils. Head brownish testaceous.
Mesonotum pale brownish ochreous to testaceous. Pleura
4g* brown, the ventral pleurites passing into obscure yellow.
Halteres dark brown, the base of st-emnairowly obscureyellow.

230

ALEXANDER

Legs denselycovered with dark setaq |tild legs elongate. Wing-s
the.groundwiitr a grayish tinge, the veins pale, ill-de{ined.ag.ainst
color; tiictiia dark brown. Venation: Vein znd A rclatively short,ending about opposite the caudal end of the oblique rn-cu.
Abdomenyellowishbrown, the sternitespalerin somespecimensb, terminMalehypopygiirm(Pl. XXVIII,fig. 9) with the basis.tyle,ating in; it"ia.t beak that is only-moderatelychitinized.Outer distist5ilerelatively stout, the mesil arm obtuse at ?pex. .P?t?l dististfle, bd, a conspicuousstraight 1od,P?leon basaltwo-thirds, theapical'thild blackened,its apex dilateclinto a shiny.roundedhead,
Aedeagus
the lateral edgeproducedinto two small chitiniz-ed_points..
a trifle shortlr- than the basal dististyle. Phallosomicplate, P,with the caudal margin rounded convex, each outer lateral angle
producedinto a short blunt lobe.
Habitat.-New Zealand(South Island).
Holotype, 3, Hilltop, Banks Peninsula,Canterbury,January
14, rgzs (A. L. Tonnoir). Paratopotypes,3 dd.
As usual in the group, the most distinctive specific featuresof the presentfly are foundin the structure of the male hypopygium.
oti Frcunrs rN PrerB xxvlIIL)BscnrprroNb -basist5,lg;bd -basal dististyle; id:inner dististyle; od -outerdististyle; p : phallosome;t :tergite.
Fig. r-Gynoplistia (Gytcoplistia)philpotti sp.n.; wing.
edentttlussp.n.; wing'
Fig. z.-Arnph,ineurus(I'{oth,ormosia)
Iiig. j.-Erioptera (Empeda)lcamiltonisp.n.; wing.
Fig. 4.-Molophilus gladiatorsp.n ; wing.
sp.n.; male hypopyF ig. 5.-Amphineurus (N othormosia)edentul,r,s
glum.
Fig. 6.-Erioptera (Empeda)hamiltoni sp.n.; tnale ltypopygium'
FiS. 7.-Motophilus speighlisp.n.; male hypopygium'
Fig. 8.-Motophil,wsgl,adiatorsp.n; male hypopygium'
trig. g.-Molophilus improcerussp.n.; male hypopygium.

Prern XXUII.

b-

7