Bulletin of the Kagoshima University Museum No. 10
ISSN-L 2188-9074

AN ANNOTATED CHECKLIST OF
MARINE AND FRESHWATER FISHES OFTOKUNOSHIMA ISLAND IN THE AMAMI ISLANDS,KAGOSHIMA, SOUTHERN JAPAN,
WITH 214 NEW RECORDSITSUKI MOCHIDA AND HIROYUKI MOTOMURA

September 2018

The Kagoshima University Museum

Cover photograph: A school of Acanthurus olivaceus (Acanthuridae) with two individuals of
Parupeneus multifasciatus (Mullidae) off Tokunoshima island. Photo by R. Suzuki
Back cover photograph: Rocky reefs off Amagi, Tokunoshima island. Photo by R. Suzuki

Bulletin of the Kagoshima University Museum No.10

ISSN-L 2188-9074

An annotated checklist of marine and freshwater
fishes of Tokunoshima island in the Amami Islands,
Kagoshima, southern Japan, with 214 new records

Itsuki Mochida1 and Hiroyuki Motomura2, 3

1

Kagoshima High School, 1–21–9 Yakushi, Kagoshima 890–0042, Japan

2

The Kagoshima University Museum, 1–21–30 Korimoto, Kagoshima 890–0065, Japan
E-mail: motomura@kaum.kagoshima-u.ac.jp
3

Corresponding author

Abstract An annotated checklist of marine and freshwater fishes ofTokunoshima island in the Amami Islands, Kagoshima Prefecture, southern
Japan, was compiled from specimen and literature surveys. A total of 505
species (257 genera, 88 families and 19 orders), including 214 species that
represent the first reliable records from the island on the basis of collected
specimens and/or underwater photographs, are listed with citation of literature,registration numbers, sizes, localities on the island, and some remarks.
Preliminary comparisons of the ichthyofauna of the island with those of 14
other localities in Japan show that the fish fauna of Tokunoshima island issimilar to that of Yoronjima island in the Amami Islands.

Published on 28 September 2018
Mochida, I. and H. Motomura. 2018. An annotated checklist of marine and freshwater fishes of Tokunoshimaisland in the Amami Islands, Kagoshima, southern Japan, with 214 new records. Bulletin of the
Kagoshima University Museum, 10: 1–80.

2

I. Mochida and H. Motomura

Introduction
Tokunoshima, a gourd-shaped island, with an area of ca. 248 m2 and a highest point of 645 m above
sea level, is the second largest island in the Amami Islands, which is located at the middle of the Ryukyu
Archipelago, southern Japan (Fig. 1). The coastline stretches about 80 km, with well-developed rocky reefs,
and there are numerous limestone caves and a large anchialine cave on the island.
Since comprehensive ichthyofaunal surveys of Tokunoshima island have not been previously completed,
two large-scale Tokunoshima Expeditions were made in September–October 2014 and November 2015 by
the Kagoshima University Museum to study the island’s ichthyofauna. In addition, the first author previously
lived on the island for several years and collected fishes there from 2013 to 2016. About 2,000 specimens ofmarine and freshwater fishes were collected during the expeditions and the first author’s stay on the island.
Sixty six unregistered specimens collected from Tokunoshima island in March–May 1965 were found at theFaculty of Fisheries, Kagoshima University, and these were registered in the fish collection of the Kagoshima
University Museum and also used in this study.
Examination of the above-mentioned specimens, published literature (plus examination of specimens
used in the literature), and unpublished underwater photographs taken in Tokunoshima island confirmed 505
species of marine and freshwater fishes, including 214 new records, from the island. All species are listed
below. The ichthyofauna of Tokunoshima island is briefly compared with that of other islands in the Ryukyu,
Osumi, and Ogasawara islands, and several localities along the Pacific coast of southern Japan.
Material and methods
Sampling methods were described and illustrated in Motomura & Matsuura (2010, 2014). Curatorial
procedures for newly collected specimens followed Motomura & Ishikawa (2013). Standard and total
lengths are abbreviated as SL and TL respectively. The systematic arrangement of families generally
follows Nelson (2006). Scientific names generally follow Eschmeyer et al. (2018), with some modifications
following recently published or unpublished taxonomic studies. Within families, species are arranged inalphabetical order of the scientific name. Standard Japanese names generally follow Nakabo (2013), having
been transliterated using the Hepburn system. Each species record was compiled from voucher specimens,
underwater photographs, and/or published literature. Registration number, number of specimens registered(if more than one), mm in SL or TL, and locality on Tokunoshima island (Fig. 2) are listed in each species
record. “Tokunoshima island” and “Tokunoshima” are the names of the island and the town respectively.
Tokunoshima island has three towns: Tokunoshima (on the east side of the island), Amagi (west side),and Isen (south side). The localities of marine and freshwater fishes are indicated as names of “area, town
(e.g., Kaneku, Amagi)” and “river (e.g., Akirigami River)” respectively. The locality is expressed simply as“Tokunoshima island” if the fish(es) were collected far off the island or if the precise locality on the island is
unknown.
Institutional codes used in this paper are as follows: FAKU (Kyoto University Museum, Kyoto, Japan),
FSKU (School of Fisheries Sciences, Kitasato University, Iwate, Japan), KAUM (Kagoshima University
Museum, Kagoshima, Japan), KPM (Kanagawa Prefectural Museum of Natural History, Kanagawa, Japan),
LICPP (Laboratory of Ichthyology, the Crown Prince’s Palace, Tokyo, Japan), MIKU (Marine Biological
Institute of Kyoto University, Wakayama, Japan; specimens currently transferred to FAKU), SMBL (SetoMarine Biology Laboratory, Kyoto University, Wakayama, Japan; specimens currently transferred to FAKU),
UPM (Faculty of Science, University of the Ryukyus, Okinawa, Japan; specimens currently transferred to
Okinawa Churaumi Aquarium, Okinawa, Japan), and USNM (Museum Support Center of National Museumof Natural History, Smithsonian Institution, Suitland, MD, USA).
Remarks
The freshwater fishes of Tokunoshima island have been well studied. The oldest available paper related tofishes of the island is probably Kuroiwa (1927) who recorded four species of fishes from rivers on the island:Marine and freshwater fishes of Tokunoshima island

3

Anguilla japonica Temminck & Schlegel, 1846 and Anguilla marmorata Quoy & Gaimard, 1824 (family
Anguilidae), Carassius sp. (as Carassius auratus) (family Cyprinidae), and Misgurnus anguillicaudatus(Cantor, 1842) (family Cobitidae). Misgurnus anguillicaudatus is currently rare on the island (Kano et al.,2016) and is probably in danger of extinction from the island.
Opsariichthys platypus (Temminck & Schlegel, 1846) (Fig. 3A, B) was probably introduced into the
Akirigami River (Fig. 3C) from the Amori River on the mainland of Kagoshima along with Plecoglossus
altivelis altivelis (Temminck & Schlegel, 1946) in May 1979 (Nishida et al., 1992; Sawashi, 1995, 1997).
However, the population of O. platypus in the Shikaura River is likely to be established more recently(Sawashi, 1995). Although O. platypus is not native to Tokunoshima island, the southern limit of its
distributional range in Japan is Tokunoshima island. Incidentally, P. a. altivelis has never been observed orrecorded from Tokunoshima island since 1979 (Sawashi, 1997).
Marine fishes of the island have been poorly studied. The only known species described as new on thebasis of a primary type specimen from Tokunoshima island is Kaupichthys japonicus Matsubara & Asano,1960 (family Chlopsidae). This species is currently known from Tokunoshima and Yoronjima islands (bothin the Amami Islands) and southern Taiwan (Hatooka, 2013a; Hibino, 2014a).
There is an anchialine cave (locally called Umbuki) in Amagi, northwestern Tokunoshima island (Fig.
4C). Two species of fishes, Xenoconger fryeri Regan, 1912 (family Chlopsidae) (Fig. 4A) and Eleotris sp. 1(family Eleotridae) (Fig. 4B) inhabit the anchialine cave, the former being known only from this anchialine
cave on Tokunoshima island in Japan.
The number of species in the top 10 most speciose families occurring at Tokunoshima island are shownin Table 1. Comparable data for 14 localities in southern Japan, including the Sagami Sea, Osezaki (Suruga
Bay), Kushimoto (Kii Peninsula), Kashiwajima island (Shikoku), Iwojima and Takeshima islands (Osumi
Islands), Yakushima island (Osumi Islands), Amami-oshima island (Amami Islands), Yoronjima island(Amami Islands), Okinawajima island (Okinawa Islands), Iejima island (Okinawa Islands), the Miyako
Islands, Ishigakijima island (Yaeyama Islands), Iriomotejima island (Yaeyama Islands), and the Ogasawara
Islands, were presented in Motomura (2016: tables 1, 2) and Nakae et al. (2018: table 2). The top three most
speciose families in all localities, except for the Sagami Sea, Osezaki (Suruga Bay), and the Ogasawara
Islands, are Gobiidae, Pomacentridae and Labridae. The top three most speciose families in the Sagami Sea,
Osezaki, and the Ogasawara Islands are Gobiidae, Labridae, and Serranidae (the last family was replacedwith Pomacentridae in the other localities) (Motomura, 2016).
Tokunoshima island is geographically located between Amami-oshima and Yoronjima islands. The
ichthyofauna of the former is most similar to that of Yoronjima island among the 14 localities in Japan. The
top 10 most speciose families, and their percentagesof number of species in each family to total number
Table 1. Number of species in the top 10 most speciose families
occurring at Tokunoshima island.
of species in Tokunoshima island are almost the samewith those of Yoronjima island (Table 1; Motomura,
FamilyNo. of species% in total
Gobiidae75
14.9
2016: table 1). The only difference between TokunoPomacentridae
38
7.5
shima and Yoronjima islands is that the family
Labridae37
7.3
Monacanthidae lanked as seventh speciose family in
Blennidae
23
4.6
Tokunoshima island is not in Yoronjima island and it is
Apogonidae
22
4.4
replaced with the family Tripterygiidae in Yoronjima
Scorpaenidae
20
4.0
island (Motomura, 2016: table 1). High species
Serranidae
18
3.6
diversity of Monacanthidae in Tokunoshima island
Monacanthidae
18
3.6
might be caused by the presence of well-developed
Chaetodontidae
16
3.2
Acanthuridae
15
3.0
rocky reefs along the coast of the island where the
Other families
223
44.2
family members prefer to inhabit.
Total

505

100.0

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I. Mochida and H. Motomura

List of fishes from Tokunoshima island
ORDER CHIMAERIFORMESFAMILY RHINOCHIMAERIDAE
Rhinochimaera africana Compagno, Stehmann & Ebert, 1990
[Jpn name: Kurotenguginzame]
Nakabo et al. (2013): west of Tokunoshima island.
ORDER SQUALIFORMESFAMILY SQUALIDAE
Squalus mitsukurii Jordan & Snyder 1903
[Jpn name: Futotsunozame]
Bandai (2018a): KAUM–I. 82930, 695.0 mm TL, west of Tokunoshima island. This study: KAUM–I. 82930,
695.0 mm TL, west of Tokunoshima island.
ORDER MYLIOBATIFORMESFAMILY DASYATIDAE
Taeniurops meyeni (Müller & Henle, 1841)[Jpn name: Madaraei] (Fig. 5A)
Underwater photograph, off Tete, Tokunoshima.
FAMILY MOBULIDAE
Mobula birostris (Walbaum, 1792)[Jpn name: Oniitomakiei] (Fig. 5B)
Underwater photograph, off Wan’ya, Amagi.
ORDER ANGUILLIFORMESFAMILY ANGUILLIDAE
Anguilla japonica Temminck & Schlegel, 1846
[Jpn name: Nihon-unagi]
Kuroiwa (1927): specimen(s) (probably registered at Imperial University, Tokyo), Tokunoshima island.
Ogawa (1937): specimen lost, Tokunoshima island.
Anguilla marmorata Quoy & Gaimard, 1824
[Jpn name: Ounagi]
Kuroiwa (1927): specimen(s) (probably registered at Imperial University, Tokyo), Tokunoshima island.
Ogawa (1937): specimen lost, Tokunoshima island. This study: KAUM–I. 98763, 228.6 mm SL, Akirigami
River; KAUM–I. 102039, 161.1 mm TL, Akirigami River.
FAMILY MURAENIDAE
Echidna nebulosa (Ahl, 1789)[Jpn name: Kumoutsubo]
KAUM–I. 105532, 186.6 mm TL, Asan, Isen.
Enchelycore lichenosa (Jordan & Snyder, 1901)[Jpn name: Kokeutsubo]Kamohara & Yamakawa (1965, as Aemasia lichenosa): 1 unregistered specimen, 140 mm TL, Matsubara,
Amagi.

Marine and freshwater fishes of Tokunoshima island

5

Enchelycore pardalis (Temminck & Schlegel, 1846)[Jpn name: Torautsubo]Kamohara & Yamakawa (1965, as Muraena pardalis): 2 unregistered specimens, 122–145 mm TL,
Matsubara, Amagi.
Gymnothorax fimbriatus (Bennett, 1832)[Jpn name: Herigoishiutsubo]
KAUM–I. 98130, 244.2 mm TL, Wan-ya River.
Gymnothorax melatremus Schultz, 1953
[Jpn name: Himeutsubo]
KAUM–I. 82211, 122.4 mm TL, Nishiagina, Amagi.
Gymnothorax meleagris (Shaw, 1795)[Jpn name: Wakautsubo]
KAUM–I. 83474, 355.1 mm TL, Isen, Isen; KAUM–I. 83476, 305.8 mm TL, Isen, Isen; KAUM–I. 86770,
241.3 mm TL, Isen, Isen; KAUM–I. 88376, 409.0 mm TL, Inutabu, Isen; KAUM–I. 89400, 373.0 mm TL,
Kaneku, Amagi; KAUM–I. 89885, 421.2 mm TL, Kaneku, Amagi.
Gymnothorax pictus (Ahl, 1789)[Jpn name: Aseutsubo]
KAUM–I. 97939, 322.2 mm TL, Isen, Isen; KAUM–I. 97940, 270.2 mm TL, Isen, Isen.
Gymnothorax pindae Smith, 1962
[Jpn name: Nokogiriutsubo]
Hatooka (1988): FAKU 51352, 358.5 mm SL, Tokunoshima island. Hatooka (2013b): Tokunoshima island.
Remarks: In Japanese waters, this species has been recorded only from Yoron-jima and Tokunoshima islands(Hatooka, 2013b; Hibino, 2014b).
Rhinomuraena quaesita Garman, 1888
[Jpn name: Hanahigeutsubo]
KPM-NR 13119, underwater photograph, Tokunoshima.
Uropterygius nagoensis Hatooka 1984
[Jpn name: Nagokikaiutsubo]
KAUM–I. 102064, 160.0 mm TL, Omonawa, Isen.
FAMILY CHLOPSIDAE
Kaupichthys japonicus Matsubara & Asano, 1960
[Jpn name: Iwaanago]
Matsubara & Asano (1960, as Kaupichthys diodontus japonicus): MIKU 1748, holotype of K. d. japonicus,
238.0 mm TL, MIKU 1749, paratype of K. d. japonicus, 237.0 mm TL, near Kametsu, Tokunoshima.
Hatooka (2013a): Tokunoshima island.
Remarks: In Japanese waters, this species has been recorded only from Yoron-jima and Tokunoshima islands(Hatooka, 2013a; Hibono, 2014a).
Xenoconger fryeri Regan, 1912
[Jpn name: Umbukianago] (Fig. 4A)

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I. Mochida and H. Motomura

Senou et al. (2014): KPM-NI 29424, 344.9 mm SL. Hibino (2018): KAUM–I. 66591, 459.6 mm TL, Asama,
Amagi. This study: KAUM–I. 66591, 459.6 mm TL, Asama, Amagi; KAUM–I. 66599, 295.0 mm TL,
Asama, Amagi.
Remarks: In Japanese waters, this species has been recorded only from anchialine-cave of Tokunoshimaisland.
FAMILY OPHICHTHIDAE
Myrichthys maculosus (Cuvier 1816)[Jpn name: Moyoumongaradoshi]
KAUM–I. 102421, 270.8 mm TL, Asan, Isen.
FAMILY CONGRIDAE
Conger cinereus Rüppell, 1871
[Jpn name: Kirianago]
Kamohara & Yamakawa (1965): 2 unregistered specimens, 277–385 mm TL, Matsubara, Amagi.
ORDER CLUPEIFORMESFAMILY CLUPEIDAE
Herklotsichthys quadrimaculatus (Rüppell, 1837)[Jpn name: Mizun]
KAUM–I. 105694, 36.4 mm SL, San, Tokunoshima; KAUM–I. 105695, 38.7 mm SL, San, Tokunoshima.
Sardinella melanura (Cuvier, 1829)[Jpn name: Oguroiwashi]
Motomura et al. (2001): URM-P 33569–33574, 6 specimens, 105–124 mm SL, mouth of Akirikami River;
Aonuma & Yagishita (2013): Tokunoshima island.
Spratelloides delicatulus (Bennett, 1832)[Jpn name: Minamikibinago]
KAUM–I. 106500, 35.9 mm SL, Inutabu, Isen; KAUM–I. 106567, 42.1 mm SL, Inutabu, Isen; KAUM–I.
106568, 39.6 mm SL, Inutabu, Isen.
Spratelloides gracilis (Temminck & Schlegel, 1846)[Jpn name: Kibinago]
KAUM–I. 105696, 23.6 mm SL, San, Tokunoshima; KAUM–I. 105697, 24.2 mm SL, San, Tokunoshima.
ORDER CYPRINIFORMESFAMILY CYPRINIDAE
Carassius spp.
[Jpn name: Gimbuna and/or “funa”]
Kuroiwa (1927, as Carassius auratus): specimen(s) (probably registered at Imperial University, Tokyo),
Tokunoshima island. Ogawa (1937, as Carassius carassius): specimen lost, Tokunoshima island. Takada etal. (2010a): 4 unregistered specimens, Tokunoshima island. Takada et al. (2010b): 4 unregistered specimens,
70.3–79.0 mm SL, Wase Pond, Tokunoshima. Kano et al. (2016): Tokunoshima island.
Opsariichthys platypus (Temminck & Schlegel, 1846)[Jpn name: Oikawa] (Fig. 3A, B)
Nishida et al. (1992, as Zacco platypus): unregistered specimens, Akirigami River. Sawashi (1995, asMarine and freshwater fishes of Tokunoshima island

7

Zacco platypus): specimens not retained, Akirigami and Shikaura rivers. This study: KAUM–I. 102247,
40.8 mm SL, Akirigami River; KAUM–I. 105617, 53.8 mm SL, Akirigami River; KAUM–I. 105618, 56.7mm SL, Akirigami River; KAUM–I. 105641, 36.9 mm SL, Akirigami River; KAUM–I. 118136, 75.0 mm
SL, Akirigami River; KAUM–I. 118137, 77.5 mm SL, Akirigami River; KAUM–I. 118138, 72.9 mm SL,
Akirigami River; KAUM–I. 118139, 49.8 mm SL, Akirigami River; KAUM–I. 118140, 59.2 mm SL,
Akirigami River; KAUM–I. 118141, 67.8 mm SL, Akirigami River; KAUM–I. 118142, 58.8 mm SL,
Akirigami River; KAUM–I. 118143, 46.0 mm SL, Akirigami River; KAUM–I. 118144, 46.6 mm SL,
Akirigami River; KAUM–I. 118145, 44.5 mm SL, Akirigami River; KAUM–I. 118146, 42.4 mm SL,
Akirigami River; KAUM–I. 118147, 69.3 mm SL, Akirigami River; KAUM–I. 118148, 61.7 mm SL,
Akirigami River; KAUM–I. 118149, 64.4 mm SL, Akirigami River; KAUM–I. 118200, 53.1 mm SL,
Akirigami River; KAUM–I. 118201, 44.8 mm SL, Akirigami River; KAUM–I. 118202, 51.7 mm SL,
Akirigami River; KAUM–I. 118203, 42.5 mm SL, Akirigami River; KAUM–I. 118205, 73.5 mm SL,
Akirigami River; KAUM–I. 118206, 64.0 mm SL, Akirigami River; KAUM–I. 118207, 62.2 mm SL,
Akirigami River; KAUM–I. 118208, 60.4 mm SL, Akirigami River; KAUM–I. 118209, 60.2 mm SL,
Akirigami River; KAUM–I. 118210, 41.2 mm SL, Akirigami River; KAUM–I. 118211, 57.0 mm SL,
Akirigami River; KAUM–I. 118212, 50.4 mm SL, Akirigami River; KAUM–I. 118213, 42.6 mm SL,
Akirigami River; KAUM–I. 118214, 52.9 mm SL, Akirigami River; KAUM–I. 118215, 52.2 mm SL,
Akirigami River; KAUM–I. 118216, 54.2 mm SL, Akirigami River; KAUM–I. 118217, 47.0 mm SL,
Akirigami River; KAUM–I. 118218, 46.6 mm SL, Akirigami River; KAUM–I. 118219, 41.6 mm SL,
Akirigami River; KAUM–I. 118220, 44.3 mm SL, Akirigami River; KAUM–I. 118221, 45.6 mm SL,
Akirigami River.
FAMILY COBITIDAE
Misgurnus anguillicaudatus (Cantor, 1842)[Jpn name: Dojo]
Kuroiwa (1927): specimen(s) (probably registered at Imperial University, Tokyo), Tokunoshima island.
Ogawa (1937): specimen lost, Tokunoshima island. Kano et al. (2016): Tokunoshima island.
ORDER SILURIFORMESFAMILY PLOTOSIDAE
Plotosus lineatus (Thunberg, 1787)[Jpn name: Minamigonzui]
Matsunuma (2018a): KAUM–I. 83714, 121.4 mm SL, Inutabu, Isen. This study: KAUM–I. 83714,
121.4 mm SL, Inutabu, Isen; KAUM–I. 102126, 64.0 mm SL, Kembuku, Isen; KAUM–I. 102127, 70.9mm SL, Kembuku, Isen; KAUM–I. 102128, 67.1 mm SL, Kembuku, Isen; KAUM–I. 102129, 68.2 mm
SL, Kembuku, Isen; KAUM–I. 102130, 64.9 mm SL, Kembuku, Isen; KAUM–I. 102131, 63.0 mm SL,
Kembuku, Isen; KAUM–I. 102132, 63.0 mm SL, Kembuku, Isen; KAUM–I. 105639, 46.3 mm SL,
Kembuku, Isen.
ORDER AULOPIFORMESFAMILY SYNODONTIDAE
Saurida gracilis (Quoy & Gaimard, 1824)[Jpn name: Madaraeso]
Shinohara (2018): KAUM–I. 65935, 227.4 mm SL, Asan, Isen. This study: KAUM–I. 65935, 227.4 mm SL,
Asan, Isen; KAUM–I. 97930, 124.1 mm SL, Omonawa, Isen.
Synodus dermatogenys Fowler, 1912
[Jpn name: Minamiakaeso]
KAUM–I. 65876, 39.4 mm SL, Nishiagina, Amagi; KAUM–I. 102004, 145.0 mm SL, Omonawa, Isen.

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I. Mochida and H. Motomura

Synodus jaculum Russell & Cressey, 1979
[Jpn name: Oguroeso]
Shinohara (2018): KAUM–I. 82164, 47.2 mm SL, Nishiagina, Amagi. This study: KAUM–I. 82164, 47.2mm SL, Nishiagina, Amagi.
Synodus ulae Schultz, 1953
[Jpn name: Akaeso]
KAUM–I. 82280, 40.0 mm SL, San, Tokunoshima.
Synodus variegatus (Lacepède, 1803)[Jpn name: Hitosujieso]
KAUM–I. 82472, 77.5 mm SL, San, Tokunoshima; KAUM–I. 105561, 89.8 mm SL, off Tokunoshima or
Amami-oshima islands; KAUM–I. 105562, 70.6 mm SL, Omonawa, Isen.
ORDER OPHIDIIFORMESFAMILY DINEMATICHTHYIDAEAlionematichthys sp.
[Jpn name: N/A]
KAUM–I. 105606, 27.7 mm SL, Omonawa, Isen.
ORDER LOPHIIFORMESFAMILY ANTENNARIIDAE
Antennarius sp.
[Jpn name: N/A]
KAUM–I. 102324, 15.6 mm SL, Omonawa, Isen.
ORDER BERYCIFORMESFAMILY HOLOCENTRIDAE
Myripristis adusta Bleeker, 1853
[Jpn name: Tsumaguromatsukasa]
KAUM–I. 102303, 41.0 mm SL, Inutabu, Isen; KAUM–I. 105653, 73.9 mm SL, off Tokunoshima or Amamioshima islands.
Myripristis berndti Jordan & Evermann, 1903
[Jpn name: Akamatsukasa]
Eguchi & Motomura (2016): KAUM–I. 66596, 63.9 mm SL, Hetono, Amagi; KAUM–I. 82564, 159.9 mm
SL, Isen, Isen. Nakae (2018): KAUM–I. 82563, 159.5 mm SL, Isen, Isen. This study: KAUM–I. 65911,
100.3 mm SL, Hetono, Amagi; KAUM–I. 65912, 104.3 mm SL, Hetono, Amagi; KAUM–I. 66595, 99.6 mm
SL, Hetono, Amagi; KAUM–I. 66596, 63.9 mm SL, Hetono, Amagi; KAUM–I. 82563, 159.5 mm SL, Isen,
Isen; KAUM–I. 82564, 159.9 mm SL, Isen, Isen; KAUM–I. 86777, 58.0 mm SL, Kametoku, Tokunoshima;
KAUM–I. 88317, 159.8 mm SL, Isen, Isen; KAUM–I. 88318, 164.2 mm SL, Isen, Isen; KAUM–I. 88319,
166.9 mm SL, Isen, Isen; KAUM–I. 88320, 158.1 mm SL, Isen, Isen; KAUM–I. 93752, 144.0 mm SL,
Agon, Isen.
Myripristis botche Cuvier, 1829
[Jpn name: Urokomatsukasa]
KAUM–I. 102304, 47.6 mm SL, Omonawa, Isen.

Marine and freshwater fishes of Tokunoshima island

9

Myripristis greenfieldi Randall & Yamakawa, 1996
[Jpn name: Tsumarimatsukasa]
KAUM–I. 97479, 126.2 mm SL, Omonawa, Isen; KAUM–I. 97480, 117.8 mm SL, Omonawa, Isen; KAUM–
I. 97482, 42.1 mm SL, Omonawa, Isen; KAUM–I. 102302, 65.9 mm SL, Inutabu, Isen.
Myripristis kuntee Valenciennes, 1831
[Jpn name: Kuroobimatsukasa]
Eguchi & Motomura (2016): KAUM–I. 65921, 120.4 mm SL, Yonama, Amagi; KAUM–I. 83437, 129.3mm SL, Isen, Isen. Nakae (2018): KAUM–I. 82281, 49.1 mm SL, San, Tokunoshima. This study: KAUM–
I. 65921, 120.4 mm SL, Yonama, Amagi; KAUM–I. 82281, 49.1 mm SL, San, Tokunoshima; KAUM–I.
83437, 129.3 mm SL, Isen, Isen; KAUM–I. 86767, 99.7 mm SL, Isen, Isen; KAUM–I. 95079, 74.8 mm SL,
Kametoku, Tokunoshima; KAUM–I. 95080, 68.8 mm SL, Kametoku, Tokunoshima.
Myripristis murdjan (Forsskål, 1775)[Jpn name: Yogorematsukasa]
Eguchi & Motomura (2016): KAUM–I. 82486, 120.1 mm SL, Kametoku, Tokunoshima. This study:
KAUM–I. 82486, 120.1 mm SL, Kametoku, Tokunoshima; KAUM–I. 102072, 60.3 mm SL, Omonawa,
Isen.
Remarks: KAUM–I. 82563 (159.5 mm SL, Isen, Isen) was reported by Eguchi & Motomura (2016) as
Myripristis murdjan but has been re-identified as M. berndti by Nakae (2018) and in this study.
Myripristis violacea Bleeker, 1851
[Jpn name: Seguromatsukasa]
KAUM–I. 86773, 113.3 mm SL, Inutabu, Isen; KAUM–I. 86774, 103.6 mm SL, Inutabu, Isen.
Neoniphon sammara (Forsskål, 1775)[Jpn name: Ukeguchiittoudai]
Eguchi & Motomura (2016): KAUM–I. 65931, 154.0 mm SL, Setaki, Amagi; KAUM–I. 66567, 41.2 mm
SL, Yonama, Amagi; KAUM–I. 82371, 44.2 mm SL, San, Tokunoshima. Nakae (2018): KAUM–I. 82371,
44.2 mm SL, San, Tokunoshima. This study: KAUM–I. 65931, 154.0 mm SL, Setaki, Amagi; KAUM–I.
66567, 41.2 mm SL, Yonama, Amagi; KAUM–I. 82371, 44.2 mm SL, San, Tokunoshima; KAUM–I. 93753,
129.4 mm SL, Inutabu, Isen; KAUM–I. 95071, 130.7 mm SL, Kametoku, Tokunoshima.
Plectrypops lima (Valenciennes, 1831)[Jpn name: Ryukyuebisu]
Eguchi & Motomura (2016): KAUM–I. 66592, 98.2 mm SL, Hetono, Amagi. Nakae (2018): KAUM–I.
66592, 98.2 mm SL, Hetono, Amagi. This study: KAUM–I. 66592, 98.2 mm SL, Hetono, Amagi.
Sargocentron ittodai (Jordan & Fowler, 1902)[Jpn name: Teriebisu]
Eguchi & Motomura (2016): KAUM–I. 66537, 67.6 mm SL, Setaki, Amagi; KAUM–I. 66597, 100.7 mm
SL, Hetono, Amagi; KAUM–I. 82575, 94.5 mm SL, San, Tokunoshima. This study: KAUM–I. 66537, 67.6mm SL, Setaki, Amagi; KAUM–I. 66597, 100.7 mm SL, Hetono, Amagi; KAUM–I. 82241, 74.6 mm SL,
San, Tokunoshima; KAUM–I. 82523, 71.9 mm SL, San, Tokunoshima; KAUM–I. 82575, 94.5 mm SL,
San, Tokunoshima; KAUM–I. 83699, 89.8 mm SL, San, Tokunoshima; KAUM–I. 86768, 114.5 mm SL,
Kametoku, Tokunoshima; KAUM–I. 86790, 81.4 mm SL, Inutabu, Isen; KAUM–I. 86791, 107.6 mm SL,
Inutabu, Isen; KAUM–I. 91357, 109.2 mm SL, Inutabu, Isen; KAUM–I. 91358, 106.3 mm SL, Inutabu, Isen;
KAUM–I. 93445, 130.0 mm SL, Agon, Isen; KAUM–I. 93840, 139.0 mm SL, Isen, Isen; KAUM–I. 93841,

10

I. Mochida and H. Motomura

133.0 mm SL, Isen, Isen; KAUM–I. 95070, 66.7 mm SL, Kametoku, Tokunoshima; KAUM–I. 102219, 53.0mm SL, Kinen, Isen; KAUM–I. 105563, 99.1 mm SL, Omonawa, Isen.
Sargocentron punctatissimum (Cuvier, 1829)[Jpn name: Hoshiebisu]
Nakae (2018): KAUM–I. 93754, 109.7 mm SL, Agon, Isen. This study: KAUM–I. 93715, 109.3 mm SL,
Isen, Isen; KAUM–I. 93716, 106.8 mm SL, Isen, Isen; KAUM–I. 93754, 109.7 mm SL, Agon, Isen.
Sargocentron spiniferum (Forsskål, 1775)[Jpn name: Togariebisu]
Eguchi & Motomura (2016): KAUM–I. 65918, 276.0 mm SL, Yonama, Amagi. Nakae (2018): KAUM–I.
65918, 276.0 mm SL, Yonama, Amagi. This study: KAUM–I. 65918, 276.0 mm SL, Yonama, Amagi.
FAMILY AULOSTOMIDAE
Aulostomus chinensis (Linnaeus, 1766)[Jpn name: Herayagara]
Kawaji (2018): KAUM–I. 89410, 149.4 mm SL, Yonama, Amagi. This study: KAUM–I. 89410, 149.4 mm
SL, Yonama, Amagi.
FAMILY FISTULARIIDAE
Fistularia commersonii Rüppell, 1838
[Jpn name: Aoyagara]
KAUM–I. 95084, 393.0 mm SL, Boma, Tokunoshima; KAUM–I. 97483, 595.0 mm SL, Omonawa, Isen;
KAUM–I. 102256, 125.2 mm SL, Boma, Tokunoshima; KAUM–I. 105558, 146.1 mm SL, Omonawa, Isen;
KAUM–I. 105583, 165.0 mm SL, Omonawa, Isen; KAUM–I. 105584, 156.0 mm SL, Inutabu, Isen.
FAMILY CENTRISCIDAE
Aeoliscus strigatus (Günther, 1861)[Jpn name: Hekoayu]
Koeda (2018a): KAUM–I. 82483, 84.4 mm SL, Kametoku, Tokunoshima. This study: KAUM–I. 82483,
84.4 mm SL, Kametoku, Tokunoshima; KAUM–I. 82549, 69.9 mm SL, Kametoku, Tokunoshima; KAUM–
I. 93509, 8.6 mm SL, San, Tokunoshima; KAUM–I. 98368, 58.6 mm SL, Boma, Tokunoshima; KAUM–I.
98369, 47.3 mm SL, Boma, Tokunoshima; KAUM–I. 98370, 34.1 mm SL, Boma, Tokunoshima; KAUM–
I. 98371, 37.5 mm SL, Boma, Tokunoshima; KAUM–I. 98372, 43.0 mm SL, Boma, Tokunoshima; KAUM–
I. 98373, 42.7 mm SL, Boma, Tokunoshima; KAUM–I. 98374, 48.0 mm SL, Boma, Tokunoshima; KAUM–
I. 102027, 28.1-56.9 mm SL, Kametoku, Tokunoshima; KAUM–I. 102161, 89.6 mm SL, Matsubara, Amagi;
KAUM–I. 102162, 10.2 mm SL, Matsubara, Amagi; KAUM–I. 102242, 35.2 mm SL, Matsubara, Amagi;
KAUM–I. 102243, 40.6 mm SL, Matsubara, Amagi; KAUM–I. 102244, 31.4 mm SL, Matsubara, Amagi;
KAUM–I. 102379, 21.1 mm SL, Kametoku, Tokunoshima; KAUM–I. 102380, 27.4 mm SL, Kametoku,
Tokunoshima; KAUM–I. 102381, 25.7 mm SL, Kametoku, Tokunoshima; KAUM–I. 102382, 27.6 mm SL,
Kametoku, Tokunoshima; KAUM–I. 102383, 28.4 mm SL, Kametoku, Tokunoshima.
ORDER GASTEROSTEIFORMESFAMILY SYNGNATHIDAE
Corythoichthys haematopterus (Bleeker, 1851)[Jpn name: Ishiyoji]
KAUM–I. 102264, 94.2 mm SL, Asan, Isen; KAUM–I. 102265, 95.6 mm SL, Asan, Isen; KAUM–I. 102266,
90.3 mm SL, Asan, Isen.

Marine and freshwater fishes of Tokunoshima island

11

Corythoichthys schultzi Herald, 1953
[Jpn name: Kuchinagaishiyoji]
KAUM–I. 82146, 76.6 mm SL, Nishiagina, Amagi.
Doryrhamphus (Dunckerocampus) naia Allen & Kuiter, 2004
[Jpn name: Kasumioiran-yoji]
KAUM–I. 65897, 116.6 mm SL, Nishiagina, Amagi; KAUM–I. 82144, 82.5 mm SL, Nishiagina, Amagi.
Doryrhamphus (Doryrhamphus) excisus excisus Kaup, 1856
[Jpn name: Hibashiyoji]
KAUM–I. 82225, 33.6 mm SL, Nishiagina, Amagi; KAUM–I. 82361, 28.7 mm SL, San, Tokunoshima;
KAUM–I. 102365, 47.8 mm SL, Furusato, Isen; KAUM–I. 102366, 39.0 mm SL, Furusato, Isen.
Doryrhamphus (Doryrhamphus) japonicus Araga & Yoshino, 1975
[Jpn name: Nokogiriyoji]
KAUM–I. 102263, 59.1 mm SL, Yonama, Amagi.
Hippichthys (Hippichthys) spicifer (Rüppell, 1838)[Jpn name: Kawayoji]
Ike et al. (1990): 5 unregistered specimens (not retained), Ose River; 2 unregistered specimens (not retained),
Minato River.
Hippichthys (Parasyngnathus) penicillus (Cantor, 1849)[Jpn name: Ganten-ishiyoji]
Kamohara & Yamakawa (1965, as Syngnathus argyrostictus): 4 unregistered specimens, 88–95 mm TL,
Matsubara, Amagi.
Remarks: The specimens reported by Kamohara & Yamakawa (1965) as Syngnathus argyrostictus are likely
to be Hippichthys (Hippichthys) spicifer, although S. argyrostictus has been synonymized with Hippichthys(Parasyngnathus) penicillus, which does not occur in the Ryukyu Islands.
Hippocampus bargibanti Whitley, 1970
[Jpn name: N/A] (Fig. 5C)
Underwater photograph, off Kaneku, Amagi.
Hippocampus kuda Bleeker, 1852
[Jpn name: Kuroumiuma]
KAUM–I. 106518, 35.8 mm SL, Yonama, Amagi.
Microphis (Oostethus) brachyurus brachyurus (Bleeker, 1854)[Jpn name: Tenguyoji]
Ike et al. (1990, as Coelonotus brachyurus brachyurus): 4 unregistered specimens (not retained), Akirigami
River. Matsunuma (2018b): KAUM–I. 82299, 160.0 mm SL, Asan, Isen. This study: KAUM–I. 82299, 160.0
mm SL, Asan, Isen; KPM-NR 16181, underwater photographs, Tokunoshima.
Microphis (Coelonotus) leiaspis (Bleeker, 1854)[Jpn name: Issen-yoji]
Ike et al. (1990, as Coelonotus leiaspis): 3 unregistered specimens (not retained), Akirigami River.
Matsunuma (2018b): KAUM–I. 82314, 108.7 mm SL, Kametoku, Tokunoshima. This study: KAUM–I.
82309, 124.3 mm SL, Kametoku, Tokunoshima; KAUM–I. 82314, 108.7 mm SL, Kametoku, Tokunoshima.

12

I. Mochida and H. Motomura

Syngnathoides biaculeatus (Bloch, 1785)[Jpn name: Togeyoji]
KAUM–I. 93529, 168.9 mm SL, Boma, Tokunoshima.
Syngnathus sp.
[Jpn name: N/A]
KAUM–I. 105597, 9.3 mm SL, Asan, Isen; KAUM–I. 105598, 8.3 mm SL, Asan, Isen; KAUM–I. 105599,
10.3 mm SL, Asan, Isen; KAUM–I. 105600, 9.2 mm SL, Asan, Isen; KAUM–I. 105601, 11.0 mm SL, Asan,
Isen; KAUM–I. 105602, 9.4 mm SL, Asan, Isen; KAUM–I. 105603, 8.6 mm SL, Asan, Isen.
ORDER MUGILIFORMESFAMILY MUGILIDAE
Chelon macrolepis (Smith, 1846)[Jpn name: Kobora]
Senou (2018): KAUM–I. 82330, 61.7 mm SL, Kametoku River. This study: KAUM–I. 82324, 52.5 mm
SL, Kametoku River; KAUM–I. 82325, 49.6 mm SL, Kametoku River; KAUM–I. 82326, 57.2 mm SL,
Kametoku River; KAUM–I. 82327, 55.0 mm SL, Kametoku River; KAUM–I. 82328, 54.4 mm SL,
Kametoku River; KAUM–I. 82329, 54.2 mm SL, Kametoku River; KAUM–I. 82330, 61.7 mm SL,
Kametoku River; KAUM–I. 82331, 76.7 mm SL, Kametoku River; KAUM–I. 82332, 99.3 mm SL,
Kametoku River; KAUM–I. 82333, 99.9 mm SL, Kametoku River; KAUM–I. 102297, 42.9 mm SL, Wan-ya
River; KAUM–I. 102298, 29.0 mm SL, Wan-ya River.
Crenimugil crenilabis (Forsskål, 1775)[Jpn name: Furaibora]
Kamohara & Yamakawa (1965): 1 unregistered specimen, 98 mm TL, Matsubara, Amagi. This study:
KAUM–I. 83455, 110.5 mm SL, Inutabu, Isen; KAUM–I. 102133, 60.5 mm SL, Kembuku, Isen; KAUM–
I. 102134, 73.5 mm SL, Kembuku, Isen; KAUM–I. 102135, 63.3 mm SL, Kembuku, Isen; KAUM–
I. 102136, 69.5 mm SL, Kembuku, Isen; KAUM–I. 102137, 73.9 mm SL, Kembuku, Isen; KAUM–
I. 102138, 76.4 mm SL, Kembuku, Isen; KAUM–I. 105649, 37.6 mm SL, Omonawa, Isen; KAUM–
I. 105650, 40.3 mm SL, Omonawa, Isen; KAUM–I. 105656, 30.4 mm SL, Omonawa, Isen; KAUM–
I. 105657, 19.0 mm SL, Omonawa, Isen; KAUM–I. 105658, 30.0 mm SL, Omonawa, Isen; KAUM–
I. 105659, 29.6 mm SL, Omonawa, Isen; KAUM–I. 105660, 30.6 mm SL, Omonawa, Isen; KAUM–
I. 105661, 26.2 mm SL, Omonawa, Isen; KAUM–I. 105662, 26.1 mm SL, Omonawa, Isen; KAUM–
I. 105663, 26.1 mm SL, Omonawa, Isen; KAUM–I. 105664, 27.8 mm SL, Omonawa, Isen; KAUM–
I. 105665, 29.0 mm SL, Omonawa, Isen; KAUM–I. 105666, 31.1 mm SL, Omonawa, Isen; KAUM–I.
105667, 28.4 mm SL, Omonawa, Isen; KAUM–I. 105668, 27.6 mm SL, Omonawa, Isen; KAUM–I. 105669,
25.8 mm SL, Omonawa, Isen; KAUM–I. 105670, 27.7 mm SL, Omonawa, Isen; KAUM–I. 105671, 27.6mm SL, Omonawa, Isen; KAUM–I. 105672, 27.6 mm SL, Omonawa, Isen; KAUM–I. 105673, 27.6 mm
SL, Omonawa, Isen; KAUM–I. 105674, 27.8 mm SL, Omonawa, Isen; KAUM–I. 105675, 27.8 mm SL,
Omonawa, Isen.
Crenimugil sp.
[Jpn name: N/A]
KAUM–I. 106527, 27.9 mm SL, Omonawa, Isen; KAUM–I. 106528, 28.9 mm SL, Omonawa, Isen; KAUM–
I. 106529, 31.9 mm SL, Omonawa, Isen; KAUM–I. 106530, 30.0 mm SL, Omonawa, Isen; KAUM–I.
106531, 30.5 mm SL, Omonawa, Isen.

Marine and freshwater fishes of Tokunoshima island

13

Ellochelon vaigiensis (Quoy & Gaimard, 1825)[Jpn name: Onibora]
Senou (2018): 83877, 42.1 mm SL, Omonawa, Isen. this study: KAUM–I. 83876, 39.1 mm SL, Omonawa,
Isen; KAUM–I. 83877, 42.1 mm SL, Omonawa, Isen; KAUM–I. 83878, 23.5 mm SL, Omonawa, Isen;
KAUM–I. 89357, 42.6 mm SL, Omonawa, Isen; KAUM–I. 102285, 48.9 mm SL, Omonawa, Isen.
Mugil cephalus cephalus Linnaeus, 1758
[Jpn name: Bora]
Senou (2018): KAUM–I. 82494, 271.3 mm SL, Kametoku, Tokunoshima. This study: KAUM–I. 82101,
265.5 mm SL, Akirigami River; KAUM–I. 82107, 106.4 mm SL, Akirigami River; KAUM–I. 82108, 124.6mm SL, Akirigami River; KAUM–I. 82109, 116.4 mm SL, Akirigami River; KAUM–I. 82137, 132.2 mm
SL, Akirigami River; KAUM–I. 82138, 106.1 mm SL, Akirigami River; KAUM–I. 82139, 120.3 mm SL,
Akirigami River; KAUM–I. 82213, 113.3 mm SL, Akirigami River; KAUM–I. 82214, 116.6 mm SL,
Akirigami River; KAUM–I. 82215, 124.4 mm SL, Akirigami River; KAUM–I. 82216, 112.1 mm SL,
Akirigami River; KAUM–I. 82217, 115.3 mm SL, Akirigami River; KAUM–I. 82218, 104.3 mm SL,
Akirigami River; KAUM–I. 82494, 271.3 mm SL, Kametoku, Tokunoshima.
Oedalechilus labiosus (Valenciennes, 1836)[Jpn name: Waniguchibora]
KAUM–I. 93874, 13.0 mm SL, Isen, Isen; KAUM–I. 105648, 29.9 mm SL, Asan, Isen.
Planiliza haematocheilus (Temminck & Schlegel, 1845)[Jpn name: Menada]
Kamohara & Yamakawa (1965, as Liza haematocheila): 1 unregistered specimen, 85 mm TL, Matsubara,
Amagi.
Remarks: Probably a mis-identification of another species of the family because this species does not occur
in the Ryukyu Islands.
ORDER ATHERINIFORMESFAMILY ATHERINIDAE
Atherinomorus lacunosus (Forster, 1801)[Jpn name: Yakushimaiwashi]
KAUM–I. 93563, 49.4 mm SL, Boma, Tokunoshima; KAUM–I. 102296, 55.9 mm SL, Boma, Tokunoshima;
KAUM–I. 105680, 24.0 mm SL, Omonawa, Isen; KAUM–I. 105681, 23.3 mm SL, Omonawa, Isen; KAUM–
I. 105682, 23.2 mm SL, Omonawa, Isen; KAUM–I. 105683, 23.8 mm SL, Omonawa, Isen; KAUM–I.
105684, 23.6 mm SL, Omonawa, Isen; KAUM–I. 105685, 22.9 mm SL, Omonawa, Isen; KAUM–I. 105686,
23.3 mm SL, Omonawa, Isen; KAUM–I. 105687, 21.4 mm SL, Omonawa, Isen; KAUM–I. 105688, 23.0mm SL, Omonawa, Isen; KAUM–I. 105689, 24.7 mm SL, Omonawa, Isen; KAUM–I. 105690, 25.2 mm
SL, Omonawa, Isen; KAUM–I. 105691, 24.7 mm SL, Omonawa, Isen; KAUM–I. 105692, 23.5 mm SL,
Omonawa, Isen.
Atherinomorus pinguis (Lacepède, 1803)[Jpn name: Hosoobiyakushimaiwashi]
KAUM–I. 106501, 31.4 mm SL, Kinen, Isen; KAUM–I. 106502, 31.0 mm SL, Kinen, Isen; KAUM–I.
106503, 27.6 mm SL, Kinen, Isen; KAUM–I. 106504, 32.0 mm SL, Kinen, Isen; KAUM–I. 106505, 30.4mm SL, Kinen, Isen; KAUM–I. 106506, 30.4 mm SL, Kinen, Isen; KAUM–I. 106507, 29.4 mm SL, Kinen,
Isen; KAUM–I. 106508, 33.4 mm SL, Kinen, Isen; KAUM–I. 106509, 28.4 mm SL, Kinen, Isen; KAUM–
I. 106510, 28.4 mm SL, Kinen, Isen; KAUM–I. 106511, 29.5 mm SL, Kinen, Isen; KAUM–I. 106512, 20.9mm SL, Kinen, Isen.

14

I. Mochida and H. Motomura

Hypoatherina tsurugae (Jordan & Starks, 1901)[Jpn name: Gin-isoiwashi]
KAUM–I. 98770, 51.4 mm SL, Omonawa, Isen.
ORDER BELONIFORMESFAMILY HEMIRAMPHIDAE
Hyporhamphus dussumieri (Valenciennes, 1847)[Jpn name: Marusayori]
KAUM–I. 85360, 47.0 mm SL, Inutabu, Isen.
FAMILY BELONIDAE
Platybelone argalus platyura (Bennett,1832)[Jpn name: Himedatsu]
Morishita (2018a): KAUM–I. 83717, 186.2 mm SL, Inutabu, Isen. This study: KAUM–I. 83717, 186.2 mm
SL, Inutabu, Isen.
Tylosurus acus melanotus (Bleeker, 1850)[Jpn name: Tenjikudatsu]
KAUM–I. 82140, 523.3 mm SL, Hetono, Amagi.
Tylosurus crocodilus crocodilus (Péron & Lesueur, 1821)[Jpn name: Okizayori]
Morishita (2018a): KAUM–I. 91361, 267.5 mm SL, Inutabu, Isen. This study: KAUM–I. 91359, 248.0 mm
SL, Inutabu, Isen; KAUM–I. 91360, 256.8 mm SL, Inutabu, Isen; KAUM–I. 91361, 267.5 mm SL, Inutabu,
Isen.
ORDER SCORPAENIFORMESFAMILY SCORPAENIDAE
Dendrochirus zebra (Cuvier, 1829)[Jpn name: Kirimmino]
Kamohara & Yamakawa (1965, as Brachirus zebra): 3 unregistered specimens, 44–115 mm TL, Matsubara,
Amagi. Matsunuma (2018c): KAUM–I. 82477, 49.0 mm SL, San, Tokunoshima. This study: KAUM–
I. 82134, 36.2 mm SL, Nishiagina, Amagi; KAUM–I. 82135, 45.7 mm SL, Nishiagina, Amagi; KAUM–
I. 82308, 61.2 mm SL, San, Tokunoshima; KAUM–I. 82477, 49.0 mm SL, San, Tokunoshima; KAUM–
I. 82478, 63.3 mm SL, San, Tokunoshima; KAUM–I. 82480, 108.3 mm SL, San, Tokunoshima; KAUM–I.
83665, 59.4 mm SL, Omonawa, Isen; KAUM–I. 102062, 110.7 mm SL, Omonawa, Isen.
Parascorpaena aurita (Rüppell, 1838)[Jpn name: Chiburunettaifusakasago]
Motomura (2018): KAUM–I. 83658, 35.5 mm SL, Omonawa, Isen. This study: KAUM–I. 83658, 35.5 mm
SL, Omonawa, Isen; KAUM–I. 102258, 36.0 mm SL, Inutabu, Isen.
Parascorpaena mossambica (Peters, 1855)[Jpn name: Nettaifusakasago]
Motomura (2018): KAUM–I. 82479, 75.5 mm SL, San, Tokunoshima. This study: KAUM–I. 82120, 44.4mm SL, Nishiagina, Amagi; KAUM–I. 82121, 38.2 mm SL, Nishiagina, Amagi; KAUM–I. 82124, 35.8 mm
SL, Nishiagina, Amagi; KAUM–I. 82479, 75.5 mm SL, San, Tokunoshima; KAUM–I. 97938, 93.9 mm SL,
Asan, Isen; KAUM–I. 98286, 90.3 mm SL, Isen, Isen.

Marine and freshwater fishes of Tokunoshima island

15

Parascorpaena moultoni (Whitley, 1961)[Jpn name: Ittenfusakasago]
Motomura (2018): KAUM–I. 82275, 16.7 mm SL, San, Tokunoshima. This study: KAUM–I. 82265, 30.5mm SL, San, Tokunoshima; KAUM–I. 82275, 16.7 mm SL, San, Tokunoshima.
Pterois antennata (Bloch, 1787)[Jpn name: Nettaiminokasago]
KAUM–I. 102123, 53.7 mm SL, Omonawa, Isen; KAUM–I. 102124, 49.7 mm SL, Omonawa, Isen;
KAUM–I. 105560, 44.6 mm SL, off Tokunoshima or Amami-oshima islands.
Pterois radiata Cuvier, 1829
[Jpn name: Kimiokoze]
Matsunuma & Motomura (2015): KAUM–I. 65844, 111.3 mm SL, Setaki, Amagi. This study: KAUM–I.
65844, 111.3 mm SL, Setaki, Amagi; KAUM–I. 102122, 50.6 mm SL, Asan, Isen.
Pterois volitans (Linnaeus, 1758)[Jpn name: Hanaminokasago]
KAUM–I. 65859, 128.9 mm SL, Nishiagina, Amagi; KAUM–I. 98800, 128.5 mm SL, Omonawa, Isen;
KAUM–I. 102235, 51.0 mm SL, Omonawa, Isen; KAUM–I. 102236, 65.0 mm SL, Omonawa, Isen;
KAUM–I. 102237, 41.0 mm SL, Omonawa, Isen.
Scorpaenodes albaiensis (Evermann & Seale, 1907)[Jpn name: Furisodekasago]
Motomura (2018): KAUM–I. 66572, 21.9 mm SL, Yonama, Amagi. This study: KAUM–I. 66572, 21.9 mm
SL, Yonama, Amagi.
Scorpaenodes guamensis (Quoy & Gaimard, 1824)[Jpn name: Guamukasago]
KAUM–I. 102019, 70.8 mm SL, Omonawa, Isen; KAUM–I. 102020, 45.8 mm SL, Omonawa, Isen;
KAUM–I. 106555, 42.7 mm SL, Omonawa, Isen.
Scorpaenodes hirsutus (Smith, 1957)[Jpn name: Mamesangokasago]
Motomura (2018): KAUM–I. 66571, 22.9 mm SL, Yonama, Amagi. This study: KAUM–I. 66571, 22.9 mm
SL, Yonama, Amagi; KAUM–I. 66576, 32.7 mm SL, Yonama, Amagi.
Scorpaenodes varipinnis Smith, 1957
[Jpn name: Seboshiisokasago]
Motomura (2018): KAUM–I. 65899, 19.9 mm SL, Nishiagina, Amagi. This study: KAUM–I. 65899, 19.9mm SL, Nishiagina, Amagi.
Scorpaenopsis macrochir Ogilby, 1910
[Jpn name: Marusubekasago]
KAUM–I. 82474, 40.8 mm SL, San, Tokunoshima; KAUM–I. 102021, 48.3 mm SL, Omonawa, Isen;
KAUM–I. 102022, 46.7 mm SL, Omonawa, Isen; KAUM–I. 102220, 60.0 mm SL, Furusato, Isen. KAUM–
I. 105524, 19.9 mm SL, Omonawa, Isen; KAUM–I. 105525, 60.3 mm SL, Yonama, Amagi.

16

I. Mochida and H. Motomura

Scorpaenopsis neglecta Heckel, 1837
[Jpn name: Satsumakasago]
Motomura (2018): KAUM–I. 82516, 42.5 mm SL, San, Tokunoshima. This study: KAUM–I. 82516, 42.5mm SL, San, Tokunoshima; KAUM–I. 98280, 62.8 mm SL, Asan, Isen.
Scorpaenopsis papuensis (Cuvier, 1829)[Jpn name: Urumakasago]
Motomura (2018): KAUM–I. 89349, 126.9 mm SL, Omonawa, Isen. This study: KAUM–I. 65896, 106.9mm SL, Nishiagina, Amagi; KAUM–I. 89349, 126.9 mm SL, Omonawa, Isen.
Scorpaenopsis possi Randall & Eschmeyer, 2002
[Jpn name: Mimitogeonikasago]
Motomura (2018): KAUM–I. 82569, 121.5 mm SL, Isen, Isen. This study: KAUM–I. 82148, 157.0 mm SL,
Nishiagina, Amagi; KAUM–I. 82569, 121.5 mm SL, Isen, Isen.
Scorpaenopsis ramaraoi Randall & Eschmeyer, 2002
[Jpn name: Inukasago]
KAUM–I. 102157, 70.0 mm SL, Inutabu, Isen.
Scorpaenopsis venosa (Cuvier, 1829)[Jpn name: Hyugakasago]
Motomura (2018): KAUM–I. 65874, 38.0 mm SL, Nishiagina, Amagi. This study: KAUM–I. 65865, 39.6mm SL, Nishiagina, Amagi; KAUM–I. 65874, 38.0 mm SL, Nishiagina, Amagi.
Sebastapistes mauritiana (Cuvier, 1829)[Jpn name: Hachijofusakasago]
KAUM–I. 105530, 31.4 mm SL, Omonawa, Isen.
Sebastapistes strongia (Cuvier, 1829)[Jpn name: Madarafusakasago]
Motomura (2018): KAUM–I. 83488, 46.6 mm SL, Omonawa, Isen. This study: KAUM–I. 83488, 46.6 mm
SL, Omonawa, Isen; KAUM–I. 102232, 42.9 mm SL, Yonama, Amagi; KAUM–I. 105535, 36.1 mm SL,
Omonawa, Isen.
Sebastapistes tinkhami (Fowler, 1946)[Jpn name: Niramikasago]
KAUM–I. 1922, 69.2 mm SL, Tokunoshima island; KAUM–I. 13776, 67.5 mm SL, Tokunoshima island;
KAUM–I. 13777, 58.2 mm SL, Tokunoshima island; KAUM–I. 13778, 55.1 mm SL, Tokunoshima island.
FAMILY SYNANCEIIDAE
Synanceia verrucosa Bloch & Schneider, 1801
[Jpn name: Onidarumaokoze]
KAUM–I. 91369, 73.1 mm SL, Yonama, Amagi.
FAMILY TETRAROGIDAE
Ablabys taenianotus (Cuvier, 1829)[Jpn name: Tsumajirookoze]
Chungthanawong (2018): KAUM–I. 83663, 39.6 mm SL, Omonawa, Isen. This study: KAUM–I. 83663,
39.6 mm SL, Omonawa, Isen.

Marine and freshwater fishes of Tokunoshima island

17

FAMILY PLATYCEPHALIDAE
Cymbacephalus beauforti (Knapp, 1973)[Jpn name: Emmagochi]
KAUM–I. 105533, 35.8 mm SL, Matsubara, Amagi; KAUM–I. 105534, 25.1 mm SL, Matsubara, Amagi.
Sunagocia otaitensis (Cuvier, 1829)[Jpn name: Fusakuchigochi]
Matsunuma (2018d): KAUM–I. 66587, 34.1 mm SL, Yonama, Amagi. This study: KAUM–I. 66587, 34.1mm SL, Yonama, Amagi; KAUM–I. 82206, 48.8 mm SL, Nishiagina, Amagi.
Thysanophrys chiltonae Schultz, 1966
[Jpn name: Kuroshimagochi]
Matsunuma (2018d): KAUM–I. 66585, 114.5 mm SL, Yonama, Amagi. This study: KAUM–I. 65879, 36.2mm SL, Nishiagina, Amagi; KAUM–I. 66585, 114.5 mm SL, Yonama, Amagi; KAUM–I. 66586, 74.0 mm
SL, Yonama, Amagi; KAUM–I. 82248, 54.9 mm SL, San, Tokunoshima.
ORDER PERCIFORMESFAMILY SERRANIDAE
Cephalopholis argus Schneider, 1801
[Jpn name: Aonomehata]
Sakurai (2018a): KAUM–I. 82529, 109.4 mm SL, Omonawa, Isen. This study: KAUM–I. 82529, 109.4 mm
SL, Omonawa, Isen; KAUM–I. 93498, 154.1 mm SL, San, Tokunoshima; KAUM–I. 102018, 103.7 mm SL,
Omonawa, Isen; KAUM–I. 102098, 86.0 mm SL, Omonawa, Isen.
Cephalopholis urodeta (Forster, 1801)[Jpn name: Nijihata]
Sakurai (2018a): KAUM–I. 86788, 148.8 mm SL, Kaneku, Amagi. This study: KAUM–I. 86788, 148.8mm SL, Kaneku, Amagi; KAUM–I. 102089, 115.4 mm SL, Isen, Isen; KAUM–I. 102096, 38.0 mm SL,
off Tokunoshima or Amami-oshima islands; KAUM–I. 102097, 47.8 mm SL, off Tokunoshima or Amamioshima islands.
Epinephelus coeruleopunctatus (Bloch, 1790)[Jpn name: Hakutenhata]
Sakurai (2018a): KAUM–I. 83471, 189.0 mm SL, Omonawa, Isen. This study: KAUM–I. 83471, 189.0 mm
SL, Omonawa, Isen.
Epinephelus corallicola (Valenciennes, 1828)[Jpn name: Hoshihiregurohata]
KAUM–I. 99600, 105.1 mm SL, Inutabu, Isen.
Epinephelus fasciatus (Forsskål, 1775)[Jpn name: Akahata]
Sakurai (2018a): KAUM–I. 65920, 120.2 mm SL, Yonama, Amagi. This study: KAUM–I. 65920, 120.2mm SL, Yonama, Amagi; KAUM–I. 83445, 147.4 mm SL, Hetono, Amagi; KAUM–I. 102040, 70.5 mm
SL, Agon, Isen; KAUM–I. 102063, 110.6 mm SL, Hetono, Amagi; KAUM–I. 102099, 100.0 mm SL, off
Tokunoshima or Amami-oshima islands.

18

I. Mochida and H. Motomura

Epinephelus hexagonatus (Forster, 1801)[Jpn name: Ishigakihata]
Sakurai (2018a): KAUM–I. 82581, 105.1 mm SL, Isen, Isen. This study: KAUM–I. 82551, 89.3 mm SL,
Omonawa, Isen; KAUM–I. 82565, 133.4 mm SL, Isen, Isen; KAUM–I. 82566, 119.9 mm SL, Isen, Isen;
KAUM–I. 82581, 105.1 mm SL, Isen, Isen; KAUM–I. 83432, 130.8 mm SL, Isen, Isen; KAUM–I. 88357,
137.0 mm SL, Isen, Isen; KAUM–I. 93444, 120.1 mm SL, Isen, Isen; KAUM–I. 93719, 113.2 mm SL,
Kaneku, Amagi; KAUM–I. 93720, 121.1 mm SL, Kaneku, Amagi; KAUM–I. 93797, 128.5 mm SL, Isen,
Isen; KAUM–I. 93798, 122.2 mm SL, Isen, Isen; KAUM–I. 102113, 111.2 mm SL, Kaneku, Amagi.
Epinephelus melanostigma Schultz, 1953
[Jpn name: Sumitsukihata]
KAUM–I. 88314, 158.5 mm SL, Isen, Isen; KAUM–I. 102215, 34.9 mm SL, Omonawa, Isen; KAUM–I.
102216, 33.2 mm SL, Omonawa, Isen.
Epinephelus merra Bloch, 1793
[Jpn name: Kammonhata]
Sakurai (2018a): KAUM–I. 98677, 80.6 mm SL, Omonawa, Isen. This study: KAUM–I. 51763, 69.0 mm
SL, Tokunoshima island; KAUM–I. 51764, 65.6 mm SL, Tokunoshima island; KAUM–I. 82122, 77.2 mm
SL, Nishiagina, Amagi; KAUM–I. 82353, 114.3 mm SL, Kametoku, Tokunoshima; KAUM–I. 82368, 145.6mm SL, Kametoku, Tokunoshima; KAUM–I. 82512, 142.7 mm SL, San, Tokunoshima; KAUM–I. 82513,
127.6 mm SL, San, Tokunoshima; KAUM–I. 82567, 180.6 mm SL, Isen, Isen; KAUM–I. 82554, 102.8 mm
SL, Omonawa, Isen; KAUM–I. 82555, 105.9 mm SL, Omonawa, Isen; KAUM–I. 83703, 159.2 mm SL,
San, Tokunoshima; KAUM–I. 93733, 111.8 mm SL, Agon, Isen; KAUM–I. 93734, 111.0 mm SL, Agon,
Isen; KAUM–I. 93735, 113.2 mm SL, Agon, Isen; KAUM–I. 93736, 98.8 mm SL, Agon, Isen; KAUM–
I. 93737, 101.6 mm SL, Agon, Isen; KAUM–I. 93738, 120.1 mm SL, Agon, Isen; KAUM–I. 93739, 111.8mm SL, Agon, Isen; KAUM–I. 93866, 108.1 mm SL, Boma, Tokunoshima; KAUM–I. 93867, 104.8 mm
SL, Boma, Tokunoshima; KAUM–I. 93868, 103.2 mm SL, Boma, Tokunoshima; KAUM–I. 95066, 123.7mm SL, Kametoku, Tokunoshima; KAUM–I. 95067, 90.7 mm SL, Kametoku, Tokunoshima; KAUM–I.
98675, 141.8 mm SL, Omonawa, Isen; KAUM–I. 98676, 71.1 mm SL, Omonawa, Isen; KAUM–I. 98677,
80.6 mm SL, Omonawa, Isen; KAUM–I. 98678, 81.2 mm SL, Omonawa, Isen; KAUM–I. 102002, 112.5mm SL, Omonawa, Isen; KAUM–I. 102003, 87.9 mm SL, Omonawa, Isen; KAUM–I. 102005, 97.9 mm
SL, Omonawa, Isen; KAUM–I. 102006, 90.5 mm SL, Omonawa, Isen; KAUM–I. 102007, 62.5 mm SL,
Omonawa, Isen; KAUM–I. 102222, 33.7 mm SL, Isen, Isen.
Epinephelus polyphekadion (Bleeker, 1849)[Jpn name: Madarahata]
KAUM–I. 98680, 111.0 mm SL, Omonawa, Isen; KAUM–I. 102103, 109.4 mm SL, Omonawa, Isen.
Epinephelus tauvina (Forsskål, 1775)[Jpn name: Hitomihata]
KAUM–I. 93449, 119.5 mm SL, Agon, Isen; KAUM–I. 102214, 27.8 mm SL, Omonawa, Isen.
Grammistes sexlineatus (Thunberg, 1792)[Jpn name: Nunosarashi]
KAUM–I. 65860, 78.8 mm SL, Nishiagina, Amagi; KAUM–I. 88427, 86.7 mm SL, Kanami, Tokunoshima;
KAUM–I. 95059, 78.4 mm SL, Boma, Tokunoshima; KAUM–I. 95060, 76.0 mm SL, Isen, Isen; KAUM–I.
102017, 76.2 mm SL, Omonawa, Isen.

Marine and freshwater fishes of Tokunoshima island

19

Grammistops ocellatus Schultz, 1953
[Jpn name: Janomenunosarashi]
Yoshida & Motomura (2015): KAUM–I. 65901, 22.8 mm SL, Nishiagina, Amagi. Sakurai (2018a): KAUM–
I. 65901, 22.8 mm SL, Nishiagina, Amagi. This study: KAUM–I. 65901, 22.8 mm SL, Nishiagina, Amagi.
Liopropoma susumi (Jordan & Seale, 1906)[Jpn name: Kosujihanasuzuki]
KAUM–I. 66577, 21.7 mm SL, Yonama, Amagi; KAUM–I. 82181, 42.1 mm SL, Nishiagina, Amagi.
Plectranthias longimanus (Weber, 1913)[Jpn name: Chibihanadai]
Sakurai (2018a): KAUM–I. 65891, 27.7 mm SL, Nishiagina, Amagi. This study: KAUM–I. 65891, 27.7mm SL, Nishiagina, Amagi; KAUM–I. 65900, 15.2 mm SL, Nishiagina, Amagi; KAUM–I. 65904, 19.3 mm
SL, Nishiagina, Amagi; KAUM–I. 82129, 25.7 mm SL, Nishiagina, Amagi; KAUM–I. 82169, 30.0 mm
SL, Nishiagina, Amagi; KAUM–I. 82170, 18.0 mm SL, Nishiagina, Amagi; KAUM–I. 82187, 32.0 mm SL,
Nishiagina, Amagi; KAUM–I. 82234, 13.2 mm SL, Nishiagina, Amagi; KAUM–I. 82267, 20.8 mm SL, San,
Tokunoshima; KAUM–I. 82298, 18.8 mm SL, San, Tokunoshima.
Pseudanthias pascalus (Jordan & Tanaka, 1927)[Jpn name: Hanagoi]
Sakurai (2018a): KAUM–I. 66508, 42.7 mm SL, Yonama, Amagi. This study: KAUM–I. 66506, 47.6 mm
SL, Yonama, Amagi; KAUM–I. 66507, 39.7 mm SL, Yonama, Amagi; KAUM–I. 66508, 42.7 mm SL,
Yonama, Amagi; KAUM–I. 66509, 33.8 mm SL, Yonama, Amagi.
Pseudanthias sp.
[Jpn name: N/A]
KAUM–I. 102409, 11.3 mm SL, Asan, Isen.
Pseudanthias squamipinnis (Peters, 1855)[Jpn name: Kingyohanadai]
Sakurai (2018a): KAUM–I. 82147, 27.3 mm SL, Nishiagina, Amagi. This study: KAUM–I. 82147, 27.3 mm
SL, Nishiagina, Amagi; KAUM–I. 82166, 25.8 mm SL, Nishiagina, Amagi.
Pseudogramma polyacantha (Bleeker, 1856)[Jpn name: Togemegisu]
KAUM–I. 65890, 35.3 mm SL, Nishiagina, Amagi; KAUM–I. 82142, 41.9 mm SL, Nishiagina, Amagi;
KAUM–I. 82226, 52.9 mm SL, Nishiagina, Amagi; KAUM–I. 82227, 39.1 mm SL, Nishiagina, Amagi;
KAUM–I. 82232, 32.2 mm SL, Nishiagina, Amagi.
FAMILY PSEUDOCHROMIDAE
Labracinus cyclophthalmus (Müller & Troschel, 1849)[Jpn name: Megisu]
Yoshida (2018a): KAUM–I. 82434, 67.5 mm SL, San, Tokunoshima. This study: KAUM–I. 82291, 66.9mm SL, San, Tokunoshima; KAUM–I. 82369, 105.5 mm SL, Kametoku, Tokunoshima; KAUM–I. 82434,
67.5 mm SL, San, Tokunoshima; KAUM–I. 82583, 128.9 mm SL, Isen, Isen; KAUM–I. 83700, 107.9mm SL, San, Tokunoshima; KAUM–I. 89355, 131.5 mm SL, Kaneku, Amagi; KAUM–I. 93794, 99.2 mm
SL, Omonawa, Isen; KAUM–I. 93795, 92.0 mm SL, Omonawa, Isen; KAUM–I. 102093, 75.3 mm SL,
Omonawa, Isen; KAUM–I. 102092, 90.5 mm SL, Omonawa, Isen.

20

I. Mochida and H. Motomura

Pictichromis porphyreus (Lubbock & Goldman, 1974)[Jpn name: Kurenainisesuzume]
KAUM–I. 65875, 31.0 mm SL, Nishiagina, Amagi; KAUM–I. 65881, 15.7 mm SL, Nishiagina, Amagi;
KAUM–I. 82297, 26.0 mm SL, San, Tokunoshima.
Pseudochromis cyanotaenia Bleeker, 1857
[Jpn name: Ryukyunisesuzume]
Kamohara & Yamakawa (1965): 1 unregistered specimen, 42 mm TL, Matsubara, Amagi. Kamohara &Yamakawa (1965, as Pseudochromis kikaii): 3 unregistered specimens, 42–54 mm TL, Matsubara, Amagi.
Yoshida (2018a): KAUM–I. 66581, 27.7 mm SL, Yonama, Amagi. This study: KAUM–I. 2211, 42.3 mm SL,
off Tokunoshima, Amami-oshima or Takarajima islands; KAUM–I. 2212, 48.0 mm SL, off Tokunoshima,
Amami-oshima or Takarajima islands; KAUM–I. 13948, 40.0 mm SL, off Tokunoshima, Amami-oshima or
Takarajima islands; KAUM–I. 13950, 32.7 mm SL, off Tokunoshima, Amami-oshima or Takarajima islands;
KAUM–I. 13951, 32.6 mm SL, off Tokunoshima, Amami-oshima or Takarajima islands; KAUM–I. 14072,
38.4 mm SL, off Tokunoshima, Amami-oshima or Takarajima islands; KAUM–I. 14073, 36.0 mm SL, off
Tokunoshima, Amami-oshima or Takarajima islands; KAUM–I. 14074, 36.9 mm SL, off Tokunoshima,
Amami-oshima or Takarajima islands; KAUM–I. 14075, 34.0 mm SL, off Tokunoshima, Amami-oshima or
Takarajima islands; KAUM–I. 66581, 27.7 mm SL, Yonama, Amagi.
Pseudochromis luteus Aoyagi, 1943
[Jpn name: Koganenisesuzume]
Kamohara & Yamakawa (1965): 1 unregistered specimen, 54 mm TL, Matsubara, Amagi. Hayashi et al.
(2013): Tokunoshima island. This study: KAUM–I. 13947, 41.6 mm SL, off Tokunoshima, Amami-oshima or
Takarajima islands; KAUM–I. 13949, 38.9 mm SL, off Tokunoshima, Amami-oshima or Takarajima islands;
KPM-NI 92669, underwater photographs, Kanami, Tokunoshima; KPM-NR 92670, underwater photographs,
Kanami, Tokunoshima.
Pseudochromis marshallensis Schultz, 1953
[Jpn name: Hoshinisesuzume]
KAUM–I. 2213, 40.1 mm SL, off Tokunoshima, Amami-oshima or Takarajima islands; KAUM–I. 65864,
19.9 mm SL, Nishiagina, Amagi; KAUM–I. 66519, 13.8 mm SL, Setaki, Amagi; KAUM–I. 66525,
22.6 mm SL, Setaki, Amagi; KAUM–I. 66539, 36.3 mm SL, Setaki, Amagi; KAUM–I. 66540, 32.5 mm
SL, Setaki, Amagi; KAUM–I. 82143, 26.1 mm SL, Nishiagina, Amagi; KAUM–I. 82157, 30.0 mm SL,
Nishiagina, Amagi; KAUM–I. 82171, 35.0 mm SL, Nishiagina, Amagi; KAUM–I. 82257, 24.6 mm SL,
San, Tokunoshima; KAUM–I. 82430, 29.4 mm SL, San, Tokunoshima; KAUM–I. 82431, 32.6 mm SL, San,
Tokunoshima.
Pseudoplesiops rosae Schultz, 1943
[Jpn name: Tanabatamegisu]
KAUM–I. 66579, 17.4 mm SL, Yonama, Amagi; KAUM–I. 66580, 27.7 mm SL, Yonama, Amagi.
FAMILY PLESIOPIDAE
Acanthoplesiops psilogaster Hardy, 1985
[Jpn name: Fuchidoritanabatauo]
KAUM–I. 105638, 13.7 mm SL, Omonawa, Isen.
Assessor randalli Allen & Kuiter, 1976
[Jpn name: Tsubametanabatauo]

Marine and freshwater fishes of Tokunoshima island

21

Yoshida (2018b): KAUM–I. 82485, 23.0 mm SL, Kametoku, Tokunoshima. This study: KAUM–I. 82485,
23.0 mm SL, Kametoku, Tokunoshima.
Calloplesiops altivelis (Steindachner, 1903)[Jpn name: Shimofuritanabatauo]
Yoshida (2018b): KAUM–I. 82278, 83.2 mm SL, San, Tokunoshima. This study: KAUM–I. 82278, 83.2 mm
SL, San, Tokunoshima.
Plesiops coeruleolineatus Rüppell, 1835
[Jpn name: Tanabatauo]
Sano et al. (1984): FAKU 58071501–58071504, 4 specimens, 51.0–68.0 mm SL, Tokunoshima island. This
study: KAUM–I. 13894, 52.0 mm SL, Tokunoshima island; KAUM–I. 13895, 53.2 mm SL, Tokunoshimaisland; KAUM–I. 13896, 53.9 mm SL, Tokunoshima island; KAUM–I. 13897, 46.8 mm SL, Tokunoshimaisland; KAUM–I. 13898, 38.9 mm SL, Tokunoshima island; KAUM–I. 13899, 41.0 mm SL, Tokunoshimaisland; KAUM–I. 13900, 36.7 mm SL, Tokunoshima island; KAUM–I. 13901, 32.7 mm SL, Tokunoshimaisland; KAUM–I. 13902, 35.7 mm SL, Tokunoshima island; KAUM–I. 13903, 37.3 mm SL, Tokunoshimaisland; KAUM–I. 13904, 29.2 mm SL, Tokunoshima island; KAUM–I. 13905, 30.3 mm SL, Tokunoshimaisland; KAUM–I. 13906, 27.6 mm SL, Tokunoshima island; KAUM–I. 13907, 26.7 mm SL, Tokunoshimaisland; KAUM–I. 13908, 21.7 mm SL, Tokunoshima island; KAUM–I. 66556, 44.4 mm SL, Setaki, Amagi;
KAUM–I. 66563, 54.5 mm SL, Yonama, Amagi; KAUM–I. 82228, 21.3 mm SL, Nishiagina, Amagi;
KAUM–I. 82450, 38.9 mm SL, San, Tokunoshima; KAUM–I. 82451, 20.8 mm SL, San, Tokunoshima;
KAUM–I. 82504, 26.6 mm SL, San, Tokunoshima; KAUM–I. 82519, 29.5 mm SL, San, Tokunoshima;
KAUM–I. 83718, 42.1 mm SL, Omonawa, Isen; KAUM–I. 102070, 51.1 mm SL, Omonawa, Isen; KAUM–
I. 102071, 60.4 mm SL, Omonawa, Isen.
FAMILY PRIACANTHIDAE
Heteropriacanthus carolinus (Cuvier, 1829)[Jpn name: Gomahirekintoki]
KAUM–I. 65843, 110.8 mm SL, Setaki, Amagi; KAUM–I. 86786, 106.3 mm SL, Inutabu, Isen.
Priacanthus hamrur (Forsskål, 1775)[Jpn name: Hosekikintoki]
KAUM–I. 82192, 210.3 mm SL, Nishiagina, Amagi.
FAMILY APOGONIDAE
Apogon caudicinctus Randall & Smith, 1988
[Jpn name: Togenagaishimochi]
KAUM–I. 102287, 33.7 mm SL, Omonawa, Isen.
Apogon crassiceps Garman, 1903
[Jpn name: Akafujitenjikudai]
Yoshida (2018c): KAUM–I. 82154, 28.4 mm SL, Nishiagina, Amagi. This study: KAUM–I. 65862, 22.5 mm
SL, Nishiagina, Amagi; KAUM–I. 82154, 28.4 mm SL, Nishiagina, Amagi.
Apogon kominatoensis Ebina, 1935
[Jpn name: Kominatotenjikudai]
Yoshida (2018c): KAUM–I. 82185, 41.1 mm SL, Nishiagina, Amagi. This study: KAUM–I. 65877, 36.6mm SL, Nishiagina, Amagi; KAUM–I. 66569, 31.9 mm SL, Yonama, Amagi; KAUM–I. 66570, 22.9 mm

22

I. Mochida and H. Motomura

SL, Yonama, Amagi; KAUM–I. 66575, 44.2 mm SL, Yonama, Amagi; KAUM–I. 82185, 41.1 mm SL,
Nishiagina, Amagi; KAUM–I. 105508, 26.9 mm SL, Omonawa, Isen.
Apogon sp.
[Jpn name: N/A]
KAUM–I. 66549, 12.2 mm SL, Setaki, Amagi; KAUM–I. 66557, 33.1 mm SL, Setaki, Amagi; KAUM–I.
66561, 17.3 mm SL, Setaki, Amagi; KAUM–I. 105510, 35.0 mm SL, Omonawa, Isen.
Apogonichthyoides timorensis (Bleeker, 1854)[Jpn name: Kakuretenjikudai]
KAUM–I. 82149, 19.2 mm SL, Nishiagina, Amagi.
Apogonichthys ocellatus (Weber, 1913)[Jpn name: Matoshibori]
Kamohara & Yamakawa (1965: fig. 4, as Papillapogon ocellatus): 1 unregistered specimen, 45 mm TL,
Matsubara, Amagi.
Cercamia eremia (Allen, 1987)[Jpn name: Sakuratenjikudai]
KAUM–I. 65855, 29.1 mm SL, Nishiagina, Amagi; KAUM–I. 65856, 32.8 mm SL, Nishiagina, Amagi;
KAUM–I. 65870, 35.1 mm SL, Nishiagina, Amagi; KAUM–I. 65871, 30.2 mm SL, Nishiagina, Amagi;
KAUM–I. 65872, 32.4 mm SL, Nishiagina, Amagi; KAUM–I. 65873, 36.3 mm SL, Nishiagina, Amagi;
KAUM–I. 66528, 21.1 mm SL, Setaki, Amagi; KAUM–I. 66541, 30.1 mm SL, Setaki, Amagi; KAUM–I.
66542, 23.1 mm SL, Setaki, Amagi; KAUM–I. 82128, 23.2 mm SL, Nishiagina, Amagi; KAUM–I. 82254,
32.5 mm SL, San, Tokunoshima; KAUM–I. 82255, 26.6 mm SL, San, Tokunoshima; KAUM–I. 82256, 23.4mm SL, San, Tokunoshima; KAUM–I. 82284, 25.1 mm SL, San, Tokunoshima.
Cheilodipterus macrodon (Lacepède, 1802)[Jpn name: Ryukyuyaraiishimochi]
Yoshida (2018c): KAUM–I. 86769, 134.1 mm SL, off Tokunoshima or Amami-oshima islands. This study:
KAUM–I. 82156, 46.1 mm SL, Nishiagina, Amagi; KAUM–I. 86769, 134.1 mm SL, off Tokunoshima or
Amami-oshima islands.
Cheilodipterus quinquelineatus Cuvier, 1828
[Jpn name: Yaraiishimochi]
Yoshida (2018c): KAUM–I. 82506, 40.8 mm SL, San, Tokunoshima. This study: KAUM–I. 82448, 48.9 mm
SL, San, Tokunoshima; KAUM–I. 82506, 40.8 mm SL, San, Tokunoshima; KAUM–I. 89917, 27.0 mm SL,
Yonama, Amagi; KAUM–I. 89918, 21.6 mm SL, Yonama, Amagi; KAUM–I. 89919, 21.0 mm SL, Yonama,
Amagi; KAUM–I. 89920, 17.4 mm SL, Yonama, Amagi; KAUM–I. 89927, 20.7 mm SL, Yonama, Amagi;
KAUM–I. 102339, 26.7 mm SL, Omonawa, Isen.
Foa fo Jordan & Seale, 1905
[Jpn name: Taiwammatoishimochi]
KAUM–I. 65889, 10.6 mm SL, Nishiagina, Amagi; KAUM–I. 89926, 19.8 mm SL, Yonama, Amagi.
Nectamia bandanensis (Bleeker, 1854)[Jpn name: Bandaishimochi]
KAUM–I. 66538, 9.4 mm SL, Setaki, Amagi.

Marine and freshwater fishes of Tokunoshima island

23

Nectamia fusca (Quoy & Gaimard, 1825)[Jpn name: Hososuzinamidatenzikudai]
Yoshida (2018c): KAUM–I. 82245, 44.5 mm SL, San, Tokunoshima. This study: KAUM–I. 82245, 44.5mm SL, San, Tokunoshima; KAUM–I. 82499, 30.8 mm SL, San, Tokunoshima. KAUM–I. 83489, 43.2 mm
SL, Omonawa, Isen; KAUM–I. 88361, 74.2 mm SL, Inutabu, Isen; KAUM–I. 95077, 81.4 mm SL, Boma,
Tokunoshima; KAUM–I. 102125, 77.2 mm SL, Omonawa, Isen; KAUM–I. 102286, 29.1 mm SL, Omonawa,
Isen.
Ostorhinchus cookii (Macleay, 1881)[Jpn name: Sujiishimochi]
Yoshida (2018c): KAUM–I. 82527, 64.3 mm SL, San, Tokunoshima. This study: KAUM–I. 82502, 80.0
mm SL, San, Tokunoshima; KAUM–I. 82527, 64.3 mm SL, San, Tokunoshima; KAUM–I. 82532, 69.1 mm
SL, Omonawa, Isen; KAUM–I. 83452, 63.3 mm SL, Inutabu, Isen; KAUM–I. 83453, 66.5 mm SL, Inutabu,
Isen; KAUM–I. 86782, 61.7 mm SL, Inutabu, Isen; KAUM–I. 86783, 68.0 mm SL, Inutabu, Isen; KAUM–I.
86784, 70.0 mm SL, Inutabu, Isen; KAUM–I. 89350, 46.0 mm SL, Omonawa, Isen; KAUM–I. 93467, 78.7mm SL, Hetono, Amagi; KAUM–I. 93468, 61.9 mm SL, Hetono, Amagi; KAUM–I. 93755, 62.4 mm SL,
Inutabu, Isen; KAUM–I. 105509, 38.5 mm SL, Omonawa, Isen.
Ostorhinchus doederleini (Jordan & Snyder, 1901)[Jpn name: Osujiishimochi]
Yoshida & Motomura (2017): KAUM–I. 65910, 86.7 mm SL, Hetono, Amagi; KAUM–I. 95082, 89.1 mm
SL, Kametoku, Tokunoshima. Yoshida (2018c): KAUM–I. 82125, 62.7 mm SL, Nishiagina, Amagi.This
study: KAUM–I. 65867, 50.6 mm SL, Nishiagina, Amagi; KAUM–I. 65868, 43.1 mm SL, Nishiagina,
Amagi; KAUM–I. 65869, 52.1 mm SL, Nishiagina, Amagi; KAUM–I. 65910, 86.7 mm SL, Hetono,
Amagi; KAUM–I. 66594, 88.7 mm SL, Hetono, Amagi; KAUM–I. 82125, 62.7 mm SL, Nishiagina, Amagi;
KAUM–I. 83492, 66.1 mm SL, Inutabu, Isen; KAUM–I. 95082, 89.1 mm SL, Kametoku, Tokunoshima;
KAUM–I. 95083, 60.1 mm SL, Kametoku, Tokunoshima.
Ostorhinchus nigrofasciatus (Lachner, 1953)[Jpn name: Minamifutosujiishimochi]
Yoshida (2018c): KAUM–I. 82258, 38.8 mm SL, San, Tokunoshima.This study: KAUM–I. 2091, 61.9 mm
SL, Tokunoshima island; KAUM–I. 13846, 60.0 mm SL, Tokunoshima island; KAUM–I. 13847, 62.2 mm
SL, Tokunoshima island; KAUM–I. 13848, 55.3 mm SL, Tokunoshima island; KAUM–I. 66513, 23.9 mm
SL, Yonama, Amagi; KAUM–I. 82258, 38.8 mm SL, San, Tokunoshima; KAUM–I. 82341, 37.0 mm SL,
San, Tokunoshima; KAUM–I. 82393, 45.2 mm SL, San, Tokunoshima; KAUM–I. 82427, 67.9 mm SL,
San, Tokunoshima; KAUM–I. 82528, 55.7 mm SL, San, Tokunoshima; KAUM–I. 82533, 64.5 mm SL,
Omonawa, Isen; KAUM–I. 93744, 76.1 mm SL, Agon, Isen; KAUM–I. 102338, 55.7 mm SL, Omonawa,
Isen.
Ostorhinchus properuptus (Whitley, 1964), lined type
[Jpn name: Kinsen-ishimochi]
Yoshida (2018c): KAUM–I. 82520, 55.5 mm SL, San, Tokunoshima.This study: KAUM–I. 65882, 47.0mm SL, Nishiagina, Amagi; KAUM–I. 66510, 44.2 mm SL, Yonama, Amagi; KAUM–I. 66511, 36.1mm SL, Yonama, Amagi; KAUM–I. 66512, 42.6 mm SL, Yonama, Amagi; KAUM–I. 82126, 49.9 mm
SL, Nishiagina, Amagi; KAUM–I. 82158, 38.5 mm SL, Nishiagina, Amagi; KAUM–I. 82159, 45.5 mm
SL, Nishiagina, Amagi; KAUM–I. 82497, 53.1 mm SL, San, Tokunoshima; KAUM–I. 82498, 29.5 mm
SL, San, Tokunoshima; KAUM–I. 82520, 55.5 mm SL, San, Tokunoshima; KAUM–I. 83664, 33.9 mm
SL, Omonawa, Isen; KAUM–I. 89414, 50.2 mm SL, Yonama, Amagi; KAUM–I. 102336, 53.7 mm SL,

24

I. Mochida and H. Motomura

Omonawa, Isen; KAUM–I. 102337, 33.7 mm SL, Omonawa, Isen; KAUM–I. 105511, 33.6 mm SL,
Omonawa, Isen.
Ostorhinchus sp.
[Jpn name: N/A]
KAUM–I. 82212, 25.2 mm SL, Nishiagina, Amagi; KAUM–I. 93869, 10.2 mm SL, Isen, Isen; KAUM–
I. 93870, 11.0 mm SL, Isen, Isen; KAUM–I. 105500, 18.5 mm SL, Omonawa, Isen; KAUM–I. 105501,
16.8 mm SL, Omonawa, Isen; KAUM–I. 105502, 18.2 mm SL, Omonawa, Isen; KAUM–I. 105503, 17.5mm SL, Omonawa, Isen; KAUM–I. 105504, 18.3 mm SL, Omonawa, Isen. KAUM–I. 105505, 17.6 mm
SL, Omonawa, Isen; KAUM–I. 105506, 12.9 mm SL, Omonawa, Isen; KAUM–I. 105507, 12.8 mm SL,
Omonawa, Isen.
Ostorhinchus sp. L type
[Jpn name: Misujitenjikudai "L-gata"]
KAUM–I. 82131, 37.6 mm SL, Nishiagina, Amagi; KAUM–I. 82155, 42.5 mm SL, Nishiagina, Amagi;
KAUM–I. 82183, 59.2 mm SL, Nishiagina, Amagi; KAUM–I. 82190, 71.0 mm SL, Nishiagina, Amagi;
KAUM–I. 105615, 57.0 mm SL, Isen, Isen; KAUM–I. 105616, 38.0 mm SL, Omonawa, Isen.
Pristiapogon exostigma (Jordan & Starks, 1906)[Jpn name: Yukataishimochi]
KAUM–I. 65863, 13.5 mm SL, Nishiagina, Amagi; KAUM–I. 65883, 25.0 mm SL, Nishiagina, Amagi;
KAUM–I. 65884, 29.9 mm SL, Nishiagina, Amagi; KAUM–I. 82130, 32.0 mm SL, Nishiagina, Amagi;
KAUM–I. 82162, 29.6 mm SL, Nishiagina, Amagi; KAUM–I. 82163, 28.5 mm SL, Nishiagina, Amagi;
KAUM–I. 82172, 23.6 mm SL, Nishiagina, Amagi; KAUM–I. 82205, 33.7 mm SL, Nishiagina, Amagi;
KAUM–I. 86778, 92.7 mm SL, Inutabu, Isen; KAUM–I. 86779, 83.3 mm SL, Inutabu, Isen; KAUM–I.
86780, 83.2 mm SL, Inutabu, Isen; KAUM–I. 86781, 66.5 mm SL, Inutabu, Isen; KAUM–I. 97477, 79.2
mm SL, Omonawa, Isen; KAUM–I. 102251, 91.4 mm SL, Inutabu, Isen; KAUM–I. 102252, 86.3 mm SL,
Inutabu, Isen.
Pristiapogon fraenatus (Valenciennes, 1832)[Jpn name: Hitosujiishimochi]
Yoshida (2018c): KAUM–I. 89923, 27.8 mm SL, Yonama, Amagi (as KAUM–I. 65884, 29.9 mm SL). This
study: KAUM–I. 89923, 27.8 mm SL, Yonama, Amagi.
Pristiapogon kallopterus (Bleeker, 1856)[Jpn name: Kasuriishimochi]
Yoshida (2018c): KAUM–I. 82182, 40.7 mm SL, Nishiagina, Amagi. This study: KAUM–I. 65908, 99.1mm SL, Hetono, Amagi; KAUM–I. 65909, 102.9 mm SL, Hetono, Amagi; KAUM–I. 82182, 40.7 mm SL,
Nishiagina, Amagi; KAUM–I. 83715, 56.1 mm SL, Inutabu, Isen; KAUM–I. 83716, 73.1 mm SL, Inutabu,
Isen; KAUM–I. 93745, 79.8 mm SL, Inutabu, Isen; KAUM–I. 93756, 82.3 mm SL, Inutabu, Isen.
Taeniamia kagoshimanus (Döderlein, 1883)[Jpn name: Sumitsukiatohikitenjikudai]
KAUM–I. 65851, 51.2 mm SL, Nishiagina, Amagi.
FAMILY SCOMBROPIDAE
Scombrops boops (Houttuyn, 1782)[Jpn name: Mutsu]
KAUM–I. 106556, 22.9 mm SL, Omonawa, Isen.

Marine and freshwater fishes of Tokunoshima island

25

FAMILY CORYPHAENIDAE
Coryphaena hippurus Linnaeus, 1758
[Jpn name: Shiira]
KAUM–I. 102193, 30.7 mm SL, Boma, Tokunoshima; KAUM–I. 102299, 44.9 mm SL, Boma,
Tokunoshima.
FAMILY CARANGIDAE
Caranx ignobilis (Forsskål, 1775)[Jpn name: Ronin-aji]
Kimura (2018): KAUM–I. 82495, 220.1 mm SL, Kametoku, Tokunoshima. This study: KAUM–I. 82495,
220.1 mm SL, Kametoku, Tokunoshima; KAUM–I. 102057, 46.0 mm SL, Kametsu, Tokunoshima; KAUM–
I. 102058, 46.8 mm SL, Kametsu, Tokunoshima; KAUM–I. 102059, 30.6 mm SL, Kametsu, Tokunoshima.
Caranx melampygus Cuvier, 1833
[Jpn name: Kasumiaji]
Kimura (2018): KAUM–I. 82366, 149.0 mm SL, Kametoku, Tokunoshima.This study: KAUM–I. 82366,
149.0 mm SL, Kametoku, Tokunoshima; KAUM–I. 82447, 140.8 mm SL, Kametoku, Tokunoshima;
KAUM–I. 82468, 144.7 mm SL, Kametoku, Tokunoshima; KAUM–I. 82469, 143.6 mm SL, Kametoku,
Tokunoshima; KAUM–I. 82511, 146.6 mm SL, Kametoku, Tokunoshima; KAUM–I. 93495, 117.1 mm SL,
San, Tokunoshima.
Caranx papuensis Alleyne & Macleay, 1877
[Jpn name: Onihiraaji]
Kimura (2018): KAUM–I. 82114, 123.9 mm SL, Akirigami River.This study: KAUM–I. 82114, 123.9 mm
SL, Akirigami River; KAUM–I. 82136, 134.0 mm SL, Akirigami River; KAUM–I. 82339, 68.0 mm SL,
Masena River.
Caranx sexfasciatus Quoy & Gaimard, 1825
[Jpn name: Gingameaji]
KAUM–I. 82340, 69.4 mm SL, Masena River; KAUM–I. 102060, 42.7 mm SL, Kametsu, Tokunoshima;
KAUM–I. 102061, 30.3 mm SL, Kametsu, Tokunoshima.
Scomberoides sp.
[Jpn name: N/A]
KAUM–I. 102201, 23.0 mm SL, Omonawa, Isen; KAUM–I. 102202, 24.1 mm SL, Omonawa, Isen.
Scomberoides tol (Cuvier, 1832)[Jpn name: Minamiikekatsuo]
KAUM–I. 105595, 34.2 mm SL, Kametsu, Tokunoshima.
Seriola quinqueradiata Temminck & Schlegel, 1845
[Jpn name: Buri]
KAUM–I. 106554, 25.6 mm SL, Omonawa, Isen.
Seriola rivoliana Valenciennes, 1833
[Jpn name: Hirenagakampachi]
KAUM–I. 102139, 59.2 mm SL, Boma, Tokunoshima; KAUM–I. 102140, 50.7 mm SL, Boma,
Tokunoshima; KAUM–I. 102141, 53.3 mm SL, Boma, Tokunoshima; KAUM–I. 102142, 41.6 mm SL,

26

I. Mochida and H. Motomura

Boma, Tokunoshima; KAUM–I. 102143, 41.6 mm SL, Boma, Tokunoshima; KAUM–I. 102144, 49.9 mm
SL, Boma, Tokunoshima; KAUM–I. 102145, 36.3 mm SL, Boma, Tokunoshima; KAUM–I. 102146, 36.1mm SL, Boma, Tokunoshima; KAUM–I. 102147, 30.1 mm SL, Boma, Tokunoshima; KAUM–I. 102148,
39.9 mm SL, Boma, Tokunoshima.
Trachinotus blochii (Lacepède, 1801)[Jpn name: Marukoban]
KAUM–I. 102047, 74.9 mm SL, Omonawa, Isen; KAUM–I. 105677, 14.3 mm SL, Isen, Isen; KAUM–I.
105678, 13.0 mm SL, Isen, Isen; KAUM–I. 105679, 12.0 mm SL, Isen, Isen; KAUM–I. 106526, 15.8 mm
SL, Isen, Isen.
FAMILY LEIOGNATHIDAE
Aurigequula fasciata (Lacepède 1803)[Jpn name: Shimahiiragi]
Fujiwara & Motomura (2016): KAUM–I. 82490, 149.0 mm SL, Kametoku, Tokunoshima; KAUM–I. 82507,
110.2 mm SL, Kametoku, Tokunoshima. Kimura (2018): KAUM–I. 82490, 149.3 mm SL, Kametoku,
Tokunoshima. This study: KAUM–I. 82490, 149.3 mm SL, Kametoku, Tokunoshima; KAUM–I. 82507,
110.2 mm SL, Kametoku, Tokunoshima.
FAMILY LUTJANIDAE
Lutjanus bohar (Forsskål, 1775)[Jpn name: Barafuedai]
Hata (2018a): KAUM–I. 66502, 66.1 mm SL, San, Tokunoshima. This study: KAUM–I. 66502, 66.1 mm
SL, San, Tokunoshima.
Lutjanus fulviflamma (Forsskål, 1775)[Jpn name: Nisekurohoshifuedai]
KAUM–I. 102231, 57.0 mm SL, Boma, Tokunoshima; KAUM–I. 102354, 23.7 mm SL, Omonawa, Isen.
Lutjanus fulvus (Forster, 1801)[Jpn name: Okifuedai]
Hata (2018a): KAUM–I. 82492, 140.5 mm SL, Kametoku, Tokunoshima. This study: KAUM–I. 82492,
140.5 mm SL, Kametoku, Tokunoshima; KAUM–I. 82493, 92.3 mm SL, Kametoku, Tokunoshima; KAUM–
I. 83454, 122.3 mm SL, Inutabu, Isen; KAUM–I. 83473, 109.9 mm SL, Matsubara, Amagi; KAUM–I.
89914, 25.0 mm SL, Yonama, Amagi; KAUM–I. 93561, 109.5 mm SL, Boma, Tokunoshima; KAUM–I.
93846, 77.1 mm SL, Boma, Tokunoshima; KAUM–I. 95061, 76.9 mm SL, Boma, Tokunoshima; KAUM–
I. 98124, 48.9 mm SL, Wan-ya River; KAUM–I. 98691, 110.3 mm SL, Omonawa, Isen; KAUM–I. 102180,
43.0 mm SL, Omonawa, Isen; KAUM–I. 102283, 54.8 mm SL, Boma, Tokunoshima; KAUM–I. 102306,
53.3 mm SL, Omonawa, Isen; KAUM–I. 102406, 39.6 mm SL, Omonawa, Isen; KAUM–I. 105512, 28.4mm SL, Omonawa, Isen; KAUM–I. 105651, 22.6 mm SL, Isen, Isen; KAUM–I. 105652, 25.0 mm SL,
Minato River; KAUM–I. 106560, 25.8 mm SL, Tokunoshima island.
Lutjanus gibbus (Forsskål, 1775)[Jpn name: Himefuedai]
Hata (2018a): KAUM–I. 98673, 97.8 mm SL, Omonawa, Isen. This study: KAUM–I. 93482, 76.8 mm SL,
Hetono, Amagi; KAUM–I. 93483, 75.1 mm SL, Hetono, Amagi; KAUM–I. 93564, 60.9 mm SL, Boma,
Tokunoshima; KAUM–I. 93565, 58.6 mm SL, Boma, Tokunoshima; KAUM–I. 93566, 63.9 mm SL, Boma,
Tokunoshima; KAUM–I. 93567, 63.2 mm SL, Boma, Tokunoshima; KAUM–I. 93721, 87.0 mm SL, Isen,Marine and freshwater fishes of Tokunoshima island

27

Isen; KAUM–I. 93722, 96.6 mm SL, Isen, Isen; KAUM–I. 93857, 78.2 mm SL, Boma, Tokunoshima;
KAUM–I. 95065, 104.3 mm SL, Asan, Isen; KAUM–I. 95074, 88.1 mm SL, Kametoku, Tokunoshima;
KAUM–I. 98673, 97.8 mm SL, Omonawa, Isen; KAUM–I. 102172, 57.3 mm SL, Boma, Tokunoshima;
KAUM–I. 102175, 64.2 mm SL, Boma, Tokunoshima.
Lutjanus kasmira (Forsskål, 1775)[Jpn name: Yosujifuedai]
Hata (2018a): KAUM–I. 65916, 182.7 mm SL, Yonama, Amagi. This study: KAUM–I. 65892, 34.2 mm
SL, Nishiagina, Amagi; KAUM–I. 65916, 182.7 mm SL, Yonama, Amagi; KAUM–I. 66598, 91.4 mm SL,
Hetono, Amagi; KAUM–I. 93442, 77.0 mm SL, Hetono, Amagi; KAUM–I. 93443, 75.3 mm SL, Hetono,
Amagi; KAUM–I. 93548, 55.1 mm SL, Hetono, Amagi; KAUM–I. 93549, 59.5 mm SL, Hetono, Amagi;
KAUM–I. 93554, 80.0 mm SL, Boma, Tokunoshima; KAUM–I. 93555, 79.5 mm SL, Boma, Tokunoshima;
KAUM–I. 93556, 75.2 mm SL, Boma, Tokunoshima; KAUM–I. 93557, 79.7 mm SL, Boma, Tokunoshima;
KAUM–I. 93558, 68.9 mm SL, Boma, Tokunoshima; KAUM–I. 93559, 73.3 mm SL, Boma, Tokunoshima;
KAUM–I. 93856, 75.6 mm SL, Omonawa, Isen; KAUM–I. 95058, 101.4 mm SL, Kametoku, Tokunoshima;
KAUM–I. 102174, 57.1 mm SL, Boma, Tokunoshima; KAUM–I. 102176, 56.3 mm SL, Boma,
Tokunoshima; KAUM–I. 102227, 70.6 mm SL, Omonawa, Isen; KAUM–I. 102228, 63.7 mm SL, Omonawa,
Isen; KAUM–I. 102229, 84.5 mm SL, Omonawa, Isen; KAUM–I. 102230, 57.3 mm SL, Omonawa, Isen;
KAUM–I. 102284, 68.4 mm SL, Boma, Tokunoshima.
Lutjanus monostigma (Cuvier, 1828)[Jpn name: Ittenfuedai]
Hata (2018a): KAUM–I. 65929, 161.4 mm SL, Nishiagina, Amagi. This study: KAUM–I. 65929, 161.4 mm
SL, Nishiagina, Amagi; KAUM–I. 102152, 38.2 mm SL, Omonawa, Isen; KAUM–I. 102377, 49.0 mm SL,
Omonawa, Isen.
Lutjanus quinquelineatus (Bloch, 1790)[Jpn name: Rokusenfuedai]
Hata (2018a): KAUM–I. 82508, 160.1 mm SL, Kametoku, Tokunoshima (as KAUM–I. 82509, 89.0 mm
SL). This study: KAUM–I. 82508, 160.1 mm SL, Kametoku, Tokunoshima.
Pristipomoides sieboldii (Bleeker, 1855)[Jpn name: Himedai]
Leis & Lee (1994): AMS I. 31531-001, larva, 6 mm SL, 50 km west of Tokunoshima island.
FAMILY LOBOTIDAE
Lobotes surinamensis (Bloch, 1790)[Jpn name: Matsudai]
KAUM–I. 102248, 36.6 mm SL, Boma, Tokunoshima; KAUM–I. 102249, 71.3 mm SL, Boma,
Tokunoshima; KAUM–I. 105526, 17.6 mm SL, Omonawa, Isen; KAUM–I. 105527, 13.4 mm SL, Omonawa,
Isen.
FAMILY GERREIDAE
Gerres oblongus Cuvier, 1830
[Jpn name: Nagasagi]
KAUM–I. 102028, 50.2 mm SL, Omonawa, Isen; KAUM–I. 106521, 25.1 mm SL, Asan, Isen; KAUM–I.
106522, 22.7 mm SL, Omonawa, Isen.

28

I. Mochida and H. Motomura

Gerres oyena (Forsskål, 1775)[Jpn name: Minamikurosagi]
Morishita (2018b): KAUM–I. 82491, 162.0 mm SL, Kametoku, Tokunoshima. This study: KAUM–I. 82491,
162.0 mm SL, Kametoku, Tokunoshima.
FAMILY HAEMULIDAE
Plectorhinchus chaetodonoides Lacepède, 1801
[Jpn name: Chochokoshodai]
KAUM–I. 102194, 23.7 mm SL, Yonama, Amagi.
Plectorhinchus gibbosus (Lacepède, 1802)[Jpn name: Kurokoshodai]
Hata (2018b): KAUM–I. 91367, 33.2 mm SL, Matsubara, Amagi. This study: KAUM–I. 91367, 33.2 mm
SL, Matsubara, Amagi; KAUM–I. 102031, 35.6 mm SL, Omonawa, Isen; KAUM–I. 102032, 26.4 mm SL,
Omonawa, Isen; KAUM–I. 102090, 23.5 mm SL, Omonawa, Isen; KAUM–I. 102292, 32.5 mm SL, Minato
River.
FAMILY NEMIPTERIDAE
Pentapodus nagasakiensis (Tanaka, 1915)[Jpn name: Itotamagashira]Fujiwara et al. (2014): KAUM–I. 27368, 72.6 mm SL, Kametsu, Tokunoshima. This study: KAUM–I.
27368, 72.6 mm SL, Kametsu, Tokunoshima.
Scolopsis lineata Quoy & Gaimard, 1824
[Jpn name: Yokoshimatamagashira]Nakamura (2018a): KAUM–I. 89356, 75.0 mm SL, Omonawa, Isen. This study: KAUM–I. 88744, 76.4mm SL, Inutabu, Isen; KAUM–I. 88745, 81.2 mm SL, Inutabu, Isen; KAUM–I. 89356, 75.0 mm SL,
Omonawa, Isen; KAUM–I. 102163, 23.6 mm SL, Yonama, Amagi; KAUM–I. 102200, 27.2 mm SL,
Kanami, Tokunoshima; KAUM–I. 102246, 25.1 mm SL, Kanami, Tokunoshima; KAUM–I. 102356, 27.5mm SL, Omonawa, Isen; KAUM–I. 102357, 21.8 mm SL, Omonawa, Isen; KAUM–I. 102358, 20.9 mm SL,
Omonawa, Isen.
FAMILY SPARIDAE
Acanthopagrus sivicolus Akazaki, 1962
[Jpn name: Minamikurodai]
KAUM–I. 82509, 89.0 mm SL, Kametoku, Tokunoshima; KAUM–I. 95064, 116.1 mm SL, Omonawa, Isen.
FAMILY LETHRINIDAE
Gnathodentex aureolineatus (Lacepède, 1802)[Jpn name: Nokogiridai]
Muto (2018a): KAUM–I. 91364, 87.1 mm SL, Inutabu, Isen. This study: KAUM–I. 91364, 87.1 mm SL,
Inutabu, Isen; KAUM–I. 93560, 84.6 mm SL, Boma, Tokunoshima; KAUM–I. 93850, 91.7 mm SL, Agon,
Isen; KAUM–I. 93851, 96.0 mm SL, Agon, Isen; KAUM–I. 93852, 87.7 mm SL, Agon, Isen; KAUM–I.
93853, 98.0 mm SL, Agon, Isen; KAUM–I. 93854, 93.0 mm SL, Agon, Isen; KAUM–I. 93855, 92.3 mm SL,
Agon, Isen; KAUM–I. 97476, 96.5 mm SL, Inutabu, Isen; KAUM–I. 102173, 86.2 mm SL, Inutabu, Isen.
Lethrinus miniatus (Forster, 1801)[Jpn name: Amamifuefuki]
KAUM–I. 102250, 98.5 mm SL, Omonawa, Isen.

Marine and freshwater fishes of Tokunoshima island

29

FAMILY MULLIDAE
Mulloidichthys flavolineatus flavolineatus (Lacepède, 1801)[Jpn name: Montsukiakahimeji]
KAUM–I. 98361, 148.0 mm SL, Inutabu, Isen; KAUM–I. 106553, 69.3 mm SL, Omonawa, Isen.
Mulloidichthys vanicolensis (Valenciennes, 1831)[Jpn name: Akahimeji]
Bandai (2018b): KAUM–I. 66593, 131.3 mm SL, Hetono, Amagi. This study: KAUM–I. 66593, 131.3 mm
SL, Hetono, Amagi; KAUM–I. 102091, 141.5 mm SL, Omonawa, Isen; KAUM–I. 105676, 109.9 mm SL,
Omonawa, Isen.
Parupeneus ciliatus (Lacepède, 1802)[Jpn name: Horaihimeji]
Bandai (2018b): KAUM–I. 93793, 120.3 mm SL, Omonawa, Isen. This study: KAUM–I. 93793, 120.3 mm
SL, Omonawa, Isen; KAUM–I. 102166, 112.4 mm SL, Omonawa, Isen.
Parupeneus cyclostomus (Lacepède, 1801)[Jpn name: Marukuchihimeji]
Bandai (2018b): KAUM–I. 93782, 118.0 mm SL, Omonawa, Isen. This study: KAUM–I. 93782, 118.0 mm
SL, Omonawa, Isen; KAUM–I. 102044, 108.5 mm SL, Agon, Isen; KAUM–I. 102045, 120.1 mm SL, Agon,
Isen.
Parupeneus indicus (Shaw, 1803)[Jpn name: Kobanhimeji]
KAUM–I. 65930, 209.7 mm SL, Nishiagina, Amagi.
Parupeneus multifasciatus (Quoy & Gaimard, 1825)[Jpn name: Ojisan]
Bandai (2018b): KAUM–I. 82115, 131.0 mm SL, Hetono, Amagi. This study: KAUM–I. 82115, 131.0 mm
SL, Hetono, Amagi; KAUM–I. 82239, 146.5 mm SL, San, Tokunoshima; KAUM–I. 82242, 111.1 mm SL,
San, Tokunoshima; KAUM–I. 82243, 85.2 mm SL, San, Tokunoshima; KAUM–I. 82250, 92.2 mm SL,
San, Tokunoshima; KAUM–I. 82515, 87.5 mm SL, San, Tokunoshima; KAUM–I. 82556, 113.1 mm SL,
Omonawa, Isen; KAUM–I. 82363, 98.4 mm SL, Kametoku, Tokunoshima; KAUM–I. 82367, 76.4 mm SL,
Kametoku, Tokunoshima; KAUM–I. 82568, 122.5 mm SL, Isen, Isen; KAUM–I. 83447, 121.7 mm SL,
Isen, Isen; KAUM–I. 83448, 110.1 mm SL, Isen, Isen; KAUM–I. 83449, 138.3 mm SL, Isen, Isen; KAUM–
I. 83451, 98.8 mm SL, Hetono, Amagi; KAUM–I. 83477, 117.0 mm SL, Hetono, Amagi; KAUM–I. 83701,
125.3 mm SL, Hetono, Amagi; KAUM–I. 83702, 124.6 mm SL, Hetono, Amagi; KAUM–I. 86789, 154.1
mm SL, Kaneku, Amagi; KAUM–I. 93441, 137.1 mm SL, Isen, Isen; KAUM–I. 93530, 67.3 mm SL, San,
Tokunoshima; KAUM–I. 93531, 77.7 mm SL, Hetono, Amagi; KAUM–I. 93629, 126.3 mm SL, Kaneku,
Amagi; KAUM–I. 93630, 89.4 mm SL, Kaneku, Amagi; KAUM–I. 93631, 163.8 mm SL, Kaneku, Amagi;
KAUM–I. 93632, 92.1 mm SL, Kaneku, Amagi; KAUM–I. 93758, 144.5 mm SL, Agon, Isen; KAUM–I.
93759, 134.2 mm SL, Agon, Isen; KAUM–I. 93760, 111.0 mm SL, Agon, Isen; KAUM–I. 93761, 93.6 mm
SL, Agon, Isen; KAUM–I. 93762, 86.6 mm SL, Agon, Isen; KAUM–I. 93763, 94.2 mm SL, Agon, Isen;
KAUM–I. 93757, 151.4 mm SL, Isen, Isen; KAUM–I. 98376, 184.0 mm SL, Agon, Isen; KAUM–I. 98377,
159.9 mm SL, Agon, Isen; KAUM–I. 98672, 162.5 mm SL, Omonawa, Isen; KAUM–I. 98689, 133.3 mm
SL, Omonawa, Isen; KAUM–I. 98690, 110.4 mm SL, Omonawa, Isen; KAUM–I. 102001, 112.5 mm SL,
Omonawa, Isen; KAUM–I. 102042, 71.1 mm SL, Agon, Isen; KAUM–I. 102043, 82.0 mm SL, Agon, Isen;
KAUM–I. 102121, 65.7 mm SL, San, Tokunoshima.

30

I. Mochida and H. Motomura

Parupeneus sp.
[Jpn name: N/A]
KAUM–I. 102077, 29.7 mm SL, Omonawa, Isen; KAUM–I. 102078, 31.7 mm SL, Omonawa, Isen; KAUM–
I. 102079, 30.5 mm SL, Omonawa, Isen; KAUM–I. 102080, 31.9 mm SL, Omonawa, Isen; KAUM–I.
102081, 29.5 mm SL, Omonawa, Isen; KAUM–I. 102082, 31.2 mm SL, Omonawa, Isen; KAUM–I. 102083,
32.1 mm SL, Omonawa, Isen; KAUM–I. 102088, 37.2 mm SL, Omonawa, Isen.
Upeneus vittatus (Forsskål, 1775)[Jpn name: Minamihimeji]
Bandai (2018b): KAUM–I. 98131, 159.7 mm SL, Boma, Tokunoshima. This study: KAUM–I. 98131, 159.7mm SL, Boma, Tokunoshima; KAUM–I. 102075, 155.0 mm SL, Boma, Tokunoshima; KAUM–I. 102076,
152.3 mm SL, Boma, Tokunoshima.
FAMILY PEMPHERIDAE
Pempheris adusta Bleeker, 1877
[Jpn name: Ryukyuhatampo]
Koeda & Motomura (2015): KAUM–I. 65846, 125.3 mm SL, Setaki, Amagi; KAUM–I. 65847, 57.0 mm
SL, Setaki, Amagi; KAUM–I. 65848, 50.7 mm SL, Setaki, Amagi; KAUM–I. 65849, 52.1 mm SL, Setaki,
Amagi; KAUM–I. 66550, 57.7 mm SL, Setaki, Amagi. Koeda (2018b): KAUM–I. 86785, 124.2 mm SL,
Kametoku, Tokunoshima. This study: KAUM–I. 65846, 125.3 mm SL, Setaki, Amagi; KAUM–I. 65847,
57.0 mm SL, Setaki, Amagi; KAUM–I. 65848, 50.7 mm SL, Setaki, Amagi; KAUM–I. 65849, 52.1 mm
SL, Setaki, Amagi; KAUM–I. 66550, 57.7 mm SL, Setaki, Amagi; KAUM–I. 82276, 107.5 mm SL, San,
Tokunoshima; KAUM–I. 86785, 124.2 mm SL, Kametoku, Tokunoshima; KAUM–I. 89352, 73.8 mm SL,
Omonawa, Isen; KAUM–I. 89353, 73.6 mm SL, Omonawa, Isen.
Pempheris oualensis Cuvier, 1831
[Jpn name: Yumehatampo]
Koeda et al. (2015): KAUM–I. 65845, 181.8 mm SL, Setaki, Amagi. Koeda (2018b): KAUM–I. 66554, 12.2mm SL, Setaki, Amagi. This study: KAUM–I. 65845, 181.8 mm SL, Setaki, Amagi; KAUM–I. 66554, 12.2mm SL, Setaki, Amagi.
Pempheris schwenkii Bleeker, 1855
[Jpn name: Minamihatampo]
KAUM–I. 66551, 59.2 mm SL, Setaki, Amagi; KAUM–I. 66552, 56.2 mm SL, Setaki, Amagi; KAUM–I.
66553, 53.6 mm SL, Setaki, Amagi; KAUM–I. 82501, 98.8 mm SL, San, Tokunoshima.
Pempheris sp.
[Jpn name: N/A]
KAUM–I. 106543, 13.6 mm SL, Omonawa, Isen; KAUM–I. 106544, 14.5 mm SL, Omonawa, Isen; KAUM–
I. 106545, 15.4 mm SL, Omonawa, Isen. KAUM–I; 106546, 15.4 mm SL, Omonawa, Isen; KAUM–I.
106547, 13.9 mm SL, Omonawa, Isen; KAUM–I. 106548, 13.0 mm SL, Omonawa, Isen; KAUM–I. 106549,
12.7 mm SL, Omonawa, Isen; KAUM–I. 106551, 19.3 mm SL, Agon, Isen.
FAMILY CHAETODONTIDAE
Chaetodon argentatus Smith & Radcliffe, 1911
[Jpn name: Kagamichochouo]
KAUM–I. 10579, 67.9 mm SL, Tokunoshima island; KAUM–I. 66555, 47.6 mm SL, Setaki, Amagi;
KAUM–I. 102262, 28.2 mm SL, Omonawa, Isen.

Marine and freshwater fishes of Tokunoshima island

31

Chaetodon auriga Forsskål, 1775
[Jpn name: Togechochouo]
KAUM–I. 82237, 114.7 mm SL, San, Tokunoshima; KAUM–I. 82522, 49.5 mm SL, San, Tokunoshima;
KAUM–I. 82550, 59.9 mm SL, Kametoku, Tokunoshima; KAUM–I. 93465, 127.4 mm SL, San,
Tokunoshima; KAUM–I. 93624, 134.3 mm SL, Kaneku, Amagi; KAUM–I. 93718, 139.4 mm SL, Isen, Isen;
KAUM–I. 95068, 145.2 mm SL, Omonawa, Isen; KAUM–I. 95078, 124.4 mm SL, Boma, Tokunoshima.
Chaetodon auripes Jordan & Snyder, 1901
[Jpn name: Chochouo]
Kamohara & Yamakawa (1965, as Chaetodon collare): 1 unregistered specimen, 85 mm TL, Matsubara,
Amagi. Fujiwara (2018a): KAUM–I. 93750, 131.5 mm SL, Kaneku, Amagi. This study: KAUM–I. 82489,
126.9 mm SL, Kametoku, Tokunoshima; KAUM–I. 93750, 131.5 mm SL, Kaneku, Amagi; KAUM–I.
93751, 137.0 mm SL, Kaneku, Amagi.
Chaetodon citrinellus Cuvier, 1831
[Jpn name: Gomachochouo]
KAUM–I. 82196, 52.0 mm SL, Nishiagina, Amagi; KAUM–I. 102112, 48.0 mm SL, Omonawa, Isen.
Chaetodon ephippium Cuvier, 1831
[Jpn name: Segurochochouo]
KAUM–I. 102120, 28.8 mm SL, Omonawa, Isen.
Chaetodon lunula (Lacepède, 1802)[Jpn name: Chohan]
Fujiwara (2018a): KAUM–I. 82488, 124.9 mm SL, Kametoku, Tokunoshima. This study: KAUM–I. 65926,
90.4 mm SL, Nishiagina, Amagi; KAUM–I. 82488, 124.9 mm SL, Kametoku, Tokunoshima; KAUM–I.
98762, 56.5 mm SL, Omonawa, Isen; KAUM–I. 102116, 43.3 mm SL, Omonawa, Isen; KAUM–I. 102117,
25.0 mm SL, Omonawa, Isen; KAUM–I. 102118, 24.1 mm SL, Omonawa, Isen; KAUM–I. 102238, 53.7mm SL, Omonawa, Isen.
Chaetodon lunulatus Quoy & Gaimard, 1825
[Jpn name: Misujichochouo]
Fujiwara (2018a): KAUM–I. 82357, 27.1 mm SL, San, Tokunoshima. This study: KAUM–I. 82357, 27.1mm SL, San, Tokunoshima.
Chaetodon melannotus Bloch & Schneider, 1801
[Jpn name: Akebonochochouo]
Fujiwara (2018a): KAUM–I. 65927, 114.5 mm SL, Nishiagina, Amagi. This study: KAUM–I. 65927, 114.5mm SL, Nishiagina, Amagi.
Chaetodon ornatissimus Cuvier, 1831
[Jpn name: Hanagurochochouo]
KAUM–I. 65914, 148.9 mm SL, Yonama, Amagi
Chaetodon vagabundus Linnaeus, 1758
[Jpn name: Furaichochouo]
KAUM–I. 65919, 95.6 mm SL, Yonama, Amagi; KAUM–I. 102119, 38.1 mm SL, Omonawa, Isen; KAUM–
I. 102226, 21.1 mm SL, Omonawa, Isen.

32

I. Mochida and H. Motomura

Forcipiger flavissimus Jordan & McGregor, 1898
[Jpn name: Fueyakkodai]
KAUM–I. 65923, 63.4 mm SL, Yonama, Amagi.
Hemitaurichthys polylepis (Bleeker, 1857)[Jpn name: Kasumichochouo] (Fig. 5D)
Underwater photograph, off Kaneku, Amagi.
Heniochus acuminatus (Linnaeus, 1758)[Jpn name: Hatatatedai]
Fujiwara (2018a): KAUM–I. 93547, 67.9 mm SL, Hetono, Amagi. This study: KAUM–I. 82419, 99.8 mm
SL, San, Tokunoshima; KAUM–I. 93547, 67.9 mm SL, Hetono, Amagi.
Heniochus chrysostomus Cuvier, 1831
[Jpn name: Minamihatatatedai]
KAUM–I. 65936, 110.1 mm SL, Setaki, Amagi; KAUM–I. 105520, 53.8 mm SL, Yonama, Amagi.
Heniochus monoceros Cuvier, 1831
[Jpn name: Onihatatatedai]
Fujiwara (2018a): KAUM–I. 82240, 110.7 mm SL, San, Tokunoshima. This study: KAUM–I. 82240, 110.7mm SL, San, Tokunoshima.
Heniochus varius (Cuvier, 1829)[Jpn name: Tsunohatatatedai]
Fujiwara (2018a): KAUM–I. 66500, 141.5 mm SL, Yonama, Amagi. This study: KAUM–I. 66500, 141.5mm SL, Yonama, Amagi.
FAMILY POMACANTHIDAE
Pomacanthus imperator (Bloch, 1787)[Jpn name: Tatejimakinchakudai]
Wibowo (2018): KAUM–I. 65928, 181.9 mm SL, Nishiagina, Amagi. This study: KAUM–I. 65928, 181.9mm SL, Nishiagina, Amagi.
Pomacanthus semicirculatus (Cuvier, 1831)[Jpn name: Sazanamiyakko]
Wibowo (2018): KAUM–I. 65925, 196.4 mm SL, Nishiagina, Amagi. This study: KAUM–I. 65925, 196.4mm SL, Nishiagina, Amagi; KAUM–I. 82372, 35.1 mm SL, San, Tokunoshima.
Pygoplites diacanthus (Boddaert, 1772)[Jpn name: Nishikiyakko]
Wibowo (2018): KAUM–I. 65878, 16.7 mm SL, Nishiagina, Amagi. This study: KAUM–I. 65878, 16.7 mm
SL, Nishiagina, Amagi.
FAMILY PENTACEROTIDAE
Evistias acutirostris (Temminck & Schlegel, 1844)[Jpn name: Tengudai] (Fig. 5E)
Underwater photograph, off Kaneku, Amagi.

Marine and freshwater fishes of Tokunoshima island

33

FAMILY CIRRHITIDAE
Amblycirrhitus bimaculus (Jenkins, 1903)[Jpn name: Futahoshigombe]
KAUM–I. 102074, 23.5 mm SL, Omonawa, Isen.
Cirrhitichthys aprinus (Cuvier, 1829)[Jpn name: Minamigombe]
KAUM–I. 102038, 134.6 mm SL, Isen, Isen.
Cirrhitichthys falco Randall, 1963
[Jpn name: Sarasagombe]
KAUM–I. 65853, 64.1 mm SL, Nishiagina, Amagi; KAUM–I. 82127, 43.8 mm SL, Nishiagina, Amagi.
Cirrhitichthys oxycephalus (Bleeker, 1855)[Jpn name: Himegombe]
KAUM–I. 105545, 63.4 mm SL, Kaneku, Amagi.
Cirrhitops hubbardi (Schultz, 1943)[Jpn name: Sumitsukigombe]
KAUM–I. 105577, 18.5 mm SL, Asan, Isen.
Cirrhitus pinnulatus pinnulatus (Forster, 1801)[Jpn name: Isogombe]
Kamohara & Yamakawa (1965, as Cirrhitus pinnulatus): 1 unregistered specimen, 125 mm TL, Matsubara,
Amagi. Koeda (2018c): KAUM–I. 65934, 183.1 mm SL, Setaki, Amagi. This study: KAUM–I. 65934, 183.1mm SL, Setaki, Amagi; KAUM–I. 83482, 122.1 mm SL, Kaneku, Amagi; KAUM–I. 86787, 141.0 mm SL,
Isen, Isen; KAUM–I. 97481, 135.0 mm SL, Agon, Isen.
Paracirrhites arcatus (Cuvier, 1829)[Jpn name: Meganegombe] (Fig. 5F)
Underwater photograph, off Kanami, Tokunoshima.
FAMILY CHEILODACTYLIDAE
Goniistius zonatus (Cuvier, 1830)[Jpn name: Takanohadai]
Hata & Motomura (2016): KAUM–I. 82500, 283.0 mm SL, west of Tokunoshima island. Inaba (2018a):
KAUM–I. 82500, 283.0 mm SL, west of Tokunoshima island. This study: KAUM–I. 82500, 283.0 mm SL,
west of Tokunoshima island; KAUM–I. 102037, 49.5 mm SL, Omonawa, Isen; KAUM–I. 102094, 60.0 mm
SL, Omonawa, Isen.
FAMILY CICHLIDAE
Oreochromis mossambicus (Peters, 1852)[Jpn name: Kawasuzume]
Sawashi (1995): specimens not retained, Shikaura River.
Tilapia zillii (Gervais, 1848)[Jpn name: Ziruteirapia]
KAUM–I. 102014, 66.7 mm SL, Omonawa River; KAUM–I. 102015, 68.1 mm SL, Omonawa River;
KAUM–I. 102016, 61.1 mm SL, Omonawa River; KAUM–I. 102048, 41.2 mm SL, Akirigami River.

34

I. Mochida and H. Motomura

FAMILY POMACENTRIDAE
Abudefduf notatus (Day, 1870)[Jpn name: Isosuzumedai]
Kamohara & Yamakawa (1965): 1 unregistered specimen, 85 mm TL, Matsubara, Amagi. Iwatsubo (2018a):
KAUM–I. 82438, 109.0 mm SL, Kametoku, Tokunoshima. Tihis study: KAUM–I. 82438, 109.0 mm SL,
Kametoku, Tokunoshima; KAUM–I. 82439, 81.1 mm SL, Kametoku, Tokunoshima; KAUM–I. 88315, 118.6mm SL, Isen, Isen; KAUM–I. 93717, 103.0 mm SL, Isen, Isen.
Abudefduf sexfasciatus (Lacepède, 1801)[Jpn name: Rokusensuzumedai]
Iwatsubo (2018a): KAUM–I. 98761, 79.9 mm SL, Omonawa, Isen. This study: KAUM–I. 83490, 40.2 mm
SL, Omonawa, Isen; KAUM–I. 83491, 33.1 mm SL, Omonawa, Isen; KAUM–I. 93502, 87.1 mm SL, San,
Tokunoshima; KAUM–I. 93503, 78.6 mm SL, San, Tokunoshima; KAUM–I. 93504, 79.7 mm SL, San,
Tokunoshima; KAUM–I. 93505, 76.1 mm SL, San, Tokunoshima; KAUM–I. 93506, 71.5 mm SL, San,
Tokunoshima; KAUM–I. 93507, 85.5 mm SL, San, Tokunoshima; KAUM–I. 93508, 106.6 mm SL, San,
Tokunoshima; KAUM–I. 93519, 77.3 mm SL, San, Tokunoshima; KAUM–I. 93520, 78.8 mm SL, San,
Tokunoshima; KAUM–I. 93521, 94.3 mm SL, San, Tokunoshima; KAUM–I. 93522, 78.1 mm SL, San,
Tokunoshima; KAUM–I. 93523, 84.8 mm SL, San, Tokunoshima; KAUM–I. 93524, 80.3 mm SL, San,
Tokunoshima; KAUM–I. 93525, 74.6 mm SL, San, Tokunoshima; KAUM–I. 93552, 44.0 mm SL, Hetono,
Amagi; KAUM–I. 93553, 50.7 mm SL, Hetono, Amagi; KAUM–I. 98760, 76.5 mm SL, Omonawa, Isen;
KAUM–I. 98761, 79.9 mm SL, Omonawa, Isen; KAUM–I. 102305, 34.5 mm SL, Omonawa, Isen; KAUM–
I. 102416, 17.1 mm SL, Omonawa, Isen; KAUM–I. 102417, 18.0 mm SL, Omonawa, Isen; KAUM–I.
105543, 45.7 mm SL, San, Tokunoshima; KAUM–I. 105544, 44.5 mm SL, San, Tokunoshima.
Abudefduf sordidus (Forsskål, 1775)[Jpn name: Shimasuzumedai]
KAUM–I. 82113, 102.3 mm SL, Hetono, Amagi; KAUM–I. 82198, 105.7 mm SL, Nishiagina, Amagi;
KAUM–I. 82376, 20.5 mm SL, San, Tokunoshima; KAUM–I. 82377, 26.1 mm SL, San, Tokunoshima;
KAUM–I. 82582, 128.9 mm SL, Isen, Isen; KAUM–I. 93732, 21.0 mm SL, Isen, Isen; KAUM–I. 102204,
27.4 mm SL, Omonawa, Isen; KAUM–I. 102414, 22.3 mm SL, Isen, Isen; KAUM–I. 102415, 18.6 mm SL,
Isen, Isen; KAUM–I. 102340, 12.7 mm SL, Omonawa, Isen.
Abudefduf vaigiensis (Quoy & Gaimard, 1825)[Jpn name: Oyabitcha]
Iwatsubo (2018a): KAUM–I. 93446, 93.3 mm SL, Isen, Isen. This study: KAUM–I. 93446, 93.3 mm
SL, Isen, Isen; KAUM–I. 93860, 103.0 mm SL, Agon, Isen; KAUM–I. 93861, 81.0 mm SL, Agon, Isen;
KAUM–I. 98674, 86.2 mm SL, Omonawa, Isen; KAUM–I. 102034, 22.3 mm SL, Kembuku, Isen.
Amphiprion clarkii (Bennett, 1830)[Jpn name: Kumanomi]
Iwatsubo (2018a): KAUM–I. 82249, 91.7 mm SL, San, Tokunoshima. This study: KAUM–I. 82249, 91.7mm SL, San, Tokunoshima; KAUM–I. 83475, 90.7 mm SL, Kaneku, Amagi; KAUM–I. 102023, 37.5mm SL, Omonawa, Isen; KAUM–I. 102024, 28.9 mm SL, Omonawa, Isen; KAUM–I. 102086, 57.1 mm
SL, Hetono, Amagi; KAUM–I. 102182, 29.0 mm SL, Omonawa, Isen; KAUM–I. 102225, 12.8 mm SL,
Omonawa, Isen.
Amphiprion frenatus Brevoort, 1856
[Jpn name: Hamakumanomi]

Marine and freshwater fishes of Tokunoshima island

35

Iwatsubo (2018a): KAUM–I. 89912, 24.0 mm SL, Yonama, Amagi. This study: KAUM–I. 89912, 24.0 mm
SL, Yonama, Amagi; KAUM–I. 102101, 62.4 mm SL, off Tokunoshima or Amami-oshima islands; KAUM–
I. 102195, 22.1 mm SL, Omonawa, Isen; KAUM–I. 102196, 18.9 mm SL, Omonawa, Isen.
Amphiprion perideraion Bleeker, 1855
[Jpn name: Hanabirakumanomi]
Iwatsubo (2018a): KAUM–I. 82283, 74.3 mm SL, San, Tokunoshima. This study: KAUM–I. 82263, 15.6mm SL, San, Tokunoshima; KAUM–I. 82283, 74.3 mm SL, San, Tokunoshima; KAUM–I. 102104, 52.0 mm
SL, Omonawa, Isen.
Cheiloprion labiatus (Day, 1877)[Jpn name: Atsukuchisuzumedai]
KAUM–I. 66588, 8.0 mm SL, Yonama, Amagi.
Chromis atripectoralis Welander & Schultz, 1951
[Jpn name: Aobasuzumedai]
Iwatsubo (2018a): KAUM–I. 82350, 27.0 mm SL, San, Tokunoshima. This study: KAUM–I. 82350, 27.0mm SL, San, Tokunoshima.
Chromis atripes Fowler & Bean, 1928
[Jpn name: Hiregurosuzumedai]
Iwatsubo (2018a): KAUM–I. 82173, 23.6 mm SL, Nishiagina, Amagi. This study: KAUM–I. 82173, 23.6mm SL, Nishiagina, Amagi.
Chromis chrysura (Bliss, 1883)[Jpn name: Amamisuzumedai]
Iwatsubo (2018a): KAUM–I. 82277, 38.2 mm SL, San, Tokunoshima. This study: KAUM–I. 82277, 38.2mm SL, San, Tokunoshima.
Chromis lepidolepis Bleeker, 1877
[Jpn name: Sasasuzumedai]
Iwatsubo (2018a): KAUM–I. 82180, 13.5 mm SL, Nishiagina, Amagi. This study: KAUM–I. 82180, 13.5mm SL, Nishiagina, Amagi.
Chromis margaritifera Fowler, 1946
[Jpn name: Shikokusuzumedai]
KAUM–I. 82176, 20.9 mm SL, Nishiagina, Amagi; KAUM–I. 82177, 18.2 mm SL, Nishiagina, Amagi;
KAUM–I. 82179, 17.7 mm SL, Nishiagina, Amagi.
Chromis xanthura (Bleeker, 1854)[Jpn name: Monsuzumedai]
Iwatsubo (2018a): KAUM–I. 82175, 30.0 mm SL, Nishiagina, Amagi. This study: KAUM–I. 82175, 30.0mm SL, Nishiagina, Amagi.
Chromis yamakawai Iwatsubo & Motomura, 2013
[Jpn name: Kihoshisuzumedai]
Iwatsubo (2018a): KAUM–I. 82560, 87.6 mm SL, Isen, Isen. This study: KAUM–I. 82559, 95.0 mm SL,
Isen, Isen; KAUM–I. 82560, 87.6 mm SL, Isen, Isen; KAUM–I. 82561, 81.9 mm SL, Isen, Isen; KAUM–

36

I. Mochida and H. Motomura

I. 83440, 81.6 mm SL, Isen, Isen; KAUM–I. 93625, 75.7 mm SL, Kaneku, Amagi; KAUM–I. 93626, 73.4mm SL, Kaneku, Amagi; KAUM–I. 93627, 73.6 mm SL, Kaneku, Amagi; KAUM–I. 93628, 73.3 mm SL,
Kaneku, Amagi; KAUM–I. 105591, 73.5 mm SL, Kaneku, Amagi.
Chrysiptera biocellata (Quoy & Gaimard, 1825)[Jpn name: Sujibuchisuzumedai]
Iwatsubo (2018a): KAUM–I. 82404, 18.1 mm SL, San, Tokunoshima. This study: KAUM–I. 82404, 18.1mm SL, San, Tokunoshima; KAUM–I. 83880, 29.6 mm SL, Omonawa, Isen; KAUM–I. 98392, 22.0 mm SL,
Asan, Isen.
Chrysiptera brownriggii (Bennett, 1828)[Jpn name: Miyakokisensuzumedai]
Kamohara & Yamakawa (1965, as Abudefduf albofasciatus): 1 unregistered specimen, 40 mm TL,
Matsubara, Amagi.
Chrysiptera cyanea (Quoy & Gaimard, 1825)[Jpn name: Rurisuzumedai]
Iwatsubo (2018a): KAUM–I. 82476, 34.3 mm SL, San, Tokunoshima. This study: KAUM–I. 10581, 50.9mm SL, Tokunoshima island; KAUM–I. 10582, 51.9 mm SL, Tokunoshima island; KAUM–I. 82349, 21.1mm SL, San, Tokunoshima; KAUM–I. 82476, 34.3 mm SL, San, Tokunoshima; KAUM–I. 102035, 53.1mm SL, Omonawa, Isen; KAUM–I. 102036, 38.4 mm SL, Omonawa, Isen; KAUM–I. 102105, 46.6 mm SL,
Omonawa, Isen; KAUM–I. 102106, 47.1 mm SL, Omonawa, Isen.
Chrysiptera glauca (Cuvier, 1830)[Jpn name: Nezusuzumedai]
KAUM–I. 88358, 74.0 mm SL, Inutabu, Isen; KAUM–I. 88359, 43.5 mm SL, Inutabu, Isen; KAUM–
I. 102420, 60.2 mm SL, off Tokunoshima or Amami-oshima islands; KAUM–I. 105529, 23.6 mm SL,
Omonawa, Isen.
Chrysiptera rex (Snyder, 1909)[Jpn name: Remonsuzumedai]
Iwatsubo (2018a): KAUM–I. 82496, 37.6 mm SL, San, Tokunoshima. This study: KAUM–I. 66535, 43.4mm SL, Setaki, Amagi; KAUM–I. 66584, 18.8 mm SL, Yonama, Amagi; KAUM–I. 82496, 37.6 mm SL,
San, Tokunoshima; KAUM–I. 102033, 40.1 mm SL, Omonawa, Isen; KAUM–I. 105547, 44.8 mm SL, off
Tokunoshima or Amami-oshima islands.
Chrysiptera starcki (Allen, 1973)[Jpn name: Senakirurisuzumedai]
Iwatsubo (2018a): KAUM–I. 82174, 43.0 mm SL, Nishiagina, Amagi. This study: KAUM–I. 82174, 43.0mm SL, Nishiagina, Amagi.
Chrysiptera unimaculata (Cuvier, 1830)[Jpn name: Ichimonsuzumedai]
KAUM–I. 88360, 35.8 mm SL, Inutabu, Isen; KAUM–I. 106520, 19.9 mm SL, Tokunoshima island.
Dascyllus aruanus (Linnaeus, 1758)[Jpn name: Misujiryukyusuzumedai]
Iwatsubo (2018a): KAUM–I. 82342, 54.0 mm SL, San, Tokunoshima. This study: KAUM–I. 82342, 54.0

Marine and freshwater fishes of Tokunoshima island

37mm SL, San, Tokunoshima; KAUM–I. 82343, 16.0 mm SL, San, Tokunoshima; KAUM–I. 82344, 25.6 mm
SL, San, Tokunoshima; KAUM–I. 82345, 22.2 mm SL, San, Tokunoshima; KAUM–I. 82346, 16.0 mm
SL, San, Tokunoshima; KAUM–I. 82347, 16.0 mm SL, San, Tokunoshima; KAUM–I. 89411, 43.1 mm SL,
Yonama, Amagi; KAUM–I. 89412, 28.5 mm SL, Yonama, Amagi; KAUM–I. 102107, 35.3 mm SL, Yonama,
Amagi; KAUM–I. 102293, 15.7 mm SL, Yonama, Amagi.
Dascyllus trimaculatus (Rüppell, 1829)[Jpn name: Mitsuboshikurosuzumedai]
Iwatsubo (2018a): KAUM–I. 82116, 12.5 mm SL, Nishiagina, Amagi. This study: KAUM–I. 82116, 12.5mm SL, Nishiagina, Amagi; KAUM–I. 102102, 40.0 mm SL, off Tokunoshima or Amami-oshima islands;
KAUM–I. 102197, 21.0 mm SL, Omonawa, Isen; KAUM–I. 102198, 20.0 mm SL, Omonawa, Isen;
KAUM–I. 102281, 38.3 mm SL, Shoda, Tokunoshima.
Neoglyphidodon nigroris (Cuvier, 1830)[Jpn name: Hirenagasuzumedai]
KAUM–I. 102025, 28.4 mm SL, Omonawa, Isen; KAUM–I. 102108, 36.0 mm SL, Omonawa, Isen;
KAUM–I. 105614, 44.6 mm SL, Omonawa, Isen.
Neopomacentrus taeniurus (Bleeker, 1856)[Jpn name: Ribonsuzumedai]
KAUM–I. 82294, 21.1 mm SL, San, Tokunoshima.
Plectroglyphidodon imparipennis (Vaillant & Sauvage, 1875)[Jpn name: Iwasakisuzumedai]
KAUM–I. 102055, 37.0 mm SL, Isen, Isen.
Plectroglyphidodon leucozonus (Bleeker, 1859)[Jpn name: Hakusensuzumedai]
KAUM–I. 82471, 90.6 mm SL, San, Tokunoshima; KAUM–I. 82546, 59.9 mm SL, Kametoku,
Tokunoshima; KAUM–I. 83484, 62.2 mm SL, Kaneku, Amagi; KAUM–I. 83879, 31.8 mm SL, Omonawa,
Isen; KAUM–I. 105585, 12.9 mm SL, Asan, Isen; KAUM–I. 105586, 14.6 mm SL, Asan, Isen; KAUM–I.
105587, 13.8 mm SL, Asan, Isen; KAUM–I. 105589, 12.6 mm SL, Isen, Isen.
Pomacentrus amboinensis Bleeker 1868
[Jpn name: Nisenettaisuzumedai]
Iwatsubo (2018a): KAUM–I. 82348, 58.1 mm SL, San, Tokunoshima. This study: KAUM–I. 82348, 58.1mm SL, San, Tokunoshima; KAUM–I. 102288, 39.9 mm SL, Yonama, Amagi; KAUM–I. 102289, 35.4 mm
SL, Yonama, Amagi; KAUM–I. 102290, 29.9 mm SL, Yonama, Amagi; KAUM–I. 105519, 62.4 mm SL,
San, Tokunoshima; KAUM–I. 106519, 23.4 mm SL, Omonawa, Isen.
Pomacentrus bankanensis Bleeker, 1853
[Jpn name: Meganesuzumedai]
Iwatsubo (2018a): KAUM–I. 82454, 35.1 mm SL, San, Tokunoshima. This study: KAUM–I. 66578, 17.4mm SL, Yonama, Amagi; KAUM–I. 82117, 19.2 mm SL, Nishiagina, Amagi; KAUM–I. 82118, 17.4 mm
SL, Nishiagina, Amagi; KAUM–I. 82454, 35.1 mm SL, San, Tokunoshima; KAUM–I. 82475, 18.1 mm SL,
San, Tokunoshima; KAUM–I. 105588, 15.6 mm SL, Omonawa, Isen; KAUM–I. 105590, 63.8 mm SL, Isen,
Isen.

38

I. Mochida and H. Motomura

Pomacentrus chrysurus Cuvier, 1830
[Jpn name: Ojirosuzumedai]
KAUM–I. 102073, 58.4 mm SL, Omonawa, Isen; KAUM–I. 105531, 41.6 mm SL, Omonawa, Isen.
Pomacentrus coelestis Jordan & Starks, 1901
[Jpn name: Sorasuzumedai]
KAUM–I. 102376, 18.0 mm SL, Kanami, Tokunoshima.
Pomacentrus nagasakiensis Tanaka, 1917
[Jpn name: Nagasakisuzumedai]
Iwatsubo (2018a): KAUM–I. 82282, 33.9 mm SL, San, Tokunoshima. This study: KAUM–I. 66524, 17.0mm SL, Setaki, Amagi; KAUM–I. 82282, 33.9 mm SL, San, Tokunoshima; KAUM–I. 105593, 71.6 mm SL,
Kaneku, Amagi.
Pomacentrus philippinus Evermann & Seale, 1907
[Jpn name: Firipinsuzumedai]
Iwatsubo (2018a): KAUM–I. 102294, 38.8 mm SL, Omonawa, Isen (as KAUM–I. 82294, 21.1 mm SL).
This study: KAUM–I. 83439, 67.1 mm SL, Isen, Isen; KAUM–I. 93836, 74.0 mm SL, Agon, Isen; KAUM–
I. 93837, 77.2 mm SL, Agon, Isen; KAUM–I. 93838, 85.4 mm SL, Agon, Isen; KAUM–I. 93859, 72.0 mm
SL, Kaneku, Amagi; KAUM–I. 102294, 38.8 mm SL, Omonawa, Isen; KAUM–I. 105517, 78.2 mm SL,
Isen, Isen; KAUM–I. 105518, 78.2 mm SL, Isen, Isen.
Pomacentrus vaiuli Jordan & Seale, 1906
[Jpn name: Kuromeganesuzumedai]
Iwatsubo (2018a): KAUM–I. 82247, 38.6 mm SL, San, Tokunoshima. This study: KAUM–I. 82247, 38.6mm SL, San, Tokunoshima; KAUM–I. 82293, 20.9 mm SL, San, Tokunoshima.
Stegastes altus (Okada & Ikeda, 1937)[Jpn name: Sedakasuzumedai]
Iwatsubo (2018a): KAUM–I. 98363, 85.9 mm SL, Agon, Isen. This study: KAUM–I. 82195, 99.7 mm
SL, Nishiagina, Amagi; KAUM–I. 93747, 91.6 mm SL, Isen, Isen; KAUM–I. 93748, 77.1 mm SL, Isen,
Isen; KAUM–I. 93749, 78.4 mm SL, Isen, Isen; KAUM–I. 93806, 71.2 mm SL, Omonawa, Isen; KAUM–
I. 95081, 96.2 mm SL, Kametoku, Tokunoshima; KAUM–I. 98363, 85.9 mm SL, Agon, Isen; KAUM–I.
98364, 86.2 mm SL, Agon, Isen.
Stegastes fasciolatus (Ogilby, 1889)[Jpn name: Fuchidorisuzumedai]
KAUM–I. 105592, 60.0 mm SL, Isen, Isen.
Stegastes obreptus (Whitley, 1948)[Jpn name: Aisuzumedai]
Iwatsubo (2018a): KAUM–I. 82547, 86.9 mm SL, Kametoku, Tokunoshima. This study: KAUM–I. 82547,
86.9 mm SL, Kametoku, Tokunoshima.
FAMILY TERAPONTIDAE
Terapon jarbua (Forsskål, 1775)[Jpn name: Kotohiki]
Inaba (2018b): KAUM–I. 98123, 117.0 mm SL, Boma, Tokunoshima. This study: KAUM–I. 93562, 71.4

Marine and freshwater fishes of Tokunoshima island

39mm SL, Boma, Tokunoshima; KAUM–I. 98123, 117.0 mm SL, Boma, Tokunoshima; KAUM–I. 102159,
50.8 mm SL, Boma, Tokunoshima; KAUM–I. 102239, 40.3 mm SL, Boma, Tokunoshima; KAUM–I.
102280, 29.7 mm SL, Wan-ya River; KAUM–I. 102413, 30.3 mm SL, Asan, Isen; KAUM–I. 105513, 40.1mm SL, Wan-ya River.
FAMILY KUHLIIDAE
Kuhlia marginata (Cuvier, 1829)[Jpn name: Yugoi]
Ogawa (1937): specimen lost, Tokunoshima island. Sawashi (1995): specimens not retained, Akirigami
River. Nakamura (2018b): KAUM–I. 82319, 75.2 mm SL, Kametoku River. This study: KAUM–I. 82312,
52.9 mm SL, Kametoku, Tokunoshima; KAUM–I. 82318, 57.5 mm SL, Kametoku River; KAUM–I. 82319,
75.2 mm SL, Kametoku River; KAUM–I. 82320, 68.3 mm SL, Kametoku River; KAUM–I. 82321, 72.9mm SL, Kametoku River; KAUM–I. 82322, 64.4 mm SL, Kametoku River; KAUM–I. 82323, 48.1 mm SL,
Kametoku River; KAUM–I. 105567, 23.7 mm SL, Akirigami River.
Kuhlia mugil (Forster, 1801)[Jpn name: Gin-yugoi]
KAUM–I. 82100, 177.2 mm SL, Hetono, Amagi; KAUM–I. 82102, 209.5 mm SL, Hetono, Amagi; KAUM–
I. 82103, 182.4 mm SL, Hetono, Amagi; KAUM–I. 82104, 195.4 mm SL, Hetono, Amagi; KAUM–I. 82105,
171.0 mm SL, Hetono, Amagi; KAUM–I. 82106, 160.0 mm SL, Hetono, Amagi; KAUM–I. 82112, 200.3
mm SL, Hetono, Amagi; KAUM–I. 102010, 74.2 mm SL, Asan, Isen; KAUM–I. 102114, 50.2 mm SL,
Isen, Isen; KAUM–I. 102245, 38.4 mm SL, Omonawa, Isen; KAUM–I. 102386, 19.6 mm SL, Asan, Isen;
KAUM–I. 102387, 21.1 mm SL, Asan, Isen; KAUM–I. 105655, 13.9 mm SL, Asan, Isen; KAUM–I. 106550,
12.5 mm SL, Omonawa, Isen.
Kuhlia rupestris (Lacepède, 1802)[Jpn name: Okuchiyugoi]
Nakamura (2018b): KAUM–I. 82317, 56.2 mm SL, Kametoku River. This study: KAUM–I. 82316, 91.3 mm
SL, Kametoku River; KAUM–I. 82317, 56.2 mm SL, Kametoku River.
FAMILY OPLEGNATHIDAE
Oplegnathus punctatus (Temminck & Schlegel, 1844)[Jpn name: Ishigakidai]
Kuriiwa (2018a): KAUM–I. 82352, 151.2 mm SL, Kametoku, Tokunoshima. This study: KAUM–I. 82352,
151.2 mm SL, Kametoku, Tokunoshima; KAUM–I. 102013, 48.2 mm SL, Boma, Tokunoshima; KAUM–I.
102391, 16.7 mm SL, San, Tokunoshima.
FAMILY KYPHOSIDAE
Kyphosus bigibbus Lacepède, 1801
[Jpn name: Notoisuzumi]
KAUM–I. 102269, 30.0 mm SL, Boma, Tokunoshima; KAUM–I. 102270, 26.9 mm SL, Boma,
Tokunoshima; KAUM–I. 102271, 24 mm SL, Boma, Tokunoshima; KAUM–I. 102272, 25.2 mm SL, Boma,
Tokunoshima; KAUM–I. 102273, 20.4 mm SL, Boma, Tokunoshima; KAUM–I. 102274, 25.2 mm SL,
Boma, Tokunoshima; KAUM–I. 102275, 26.3 mm SL, Boma, Tokunoshima; KAUM–I. 102276, 24.5 mm
SL, Boma, Tokunoshima; KAUM–I. 102277, 23.9 mm SL, Boma, Tokunoshima; KAUM–I. 102278, 22.0mm SL, Boma, Tokunoshima; KAUM–I. 102279, 23.0 mm SL, Boma, Tokunoshima.

40

I. Mochida and H. Motomura

Kyphosus cinerascens (Forsskål, 1775)[Jpn name: Tenjikuisaki]
Nakamura (2018c): KAUM–I. 86771, 132.9 mm SL, Inutabu, Isen. This study: KAUM–I. 82458, 150.1mm SL, Kametoku, Tokunoshima; KAUM–I. 82459, 145.6 mm SL, Kametoku, Tokunoshima; KAUM–I.
82553, 161.8 mm SL, Omonawa, Isen; KAUM–I. 86771, 132.9 mm SL, Inutabu, Isen; KAUM–I. 86772,
134.6 mm SL, Inutabu, Isen; KAUM–I. 93783, 132.2 mm SL, Omonawa, Isen; KAUM–I. 93784, 123.1mm SL, Omonawa, Isen; KAUM–I. 93785, 95.2 mm SL, Omonawa, Isen; KAUM–I. 93786, 117.0 mm
SL, Omonawa, Isen; KAUM–I. 93833, 130.1 mm SL, Omonawa, Isen; KAUM–I. 93834, 131.0 mm SL,
Omonawa, Isen; KAUM–I. 93835, 144.3 mm SL, Omonawa, Isen.
FAMILY MICROCANTHIDAE
Microcanthus strigatus (Cuvier, 1831)[Jpn name: Kagokakidai]
Haraguchi (2018): KAUM–I. 95062, 90.0 mm SL, Omonawa, Isen. This study: KAUM–I. 95062, 90.0
mm SL, Omonawa, Isen; KAUM–I. 98798, 13.9 mm SL, Kametoku, Tokunoshima; KAUM–I. 102188,
26.0 mm SL, Omonawa, Isen; KAUM–I. 102189, 25.7 mm SL, Omonawa, Isen; KAUM–I. 102190, 20.9mm SL, Omonawa, Isen; KAUM–I. 102367, 19.4 mm SL, Omonawa, Isen; KAUM–I. 102368, 12.8 mm
SL, Omonawa, Isen; KAUM–I. 102369, 11.2 mm SL, Omonawa, Isen; KAUM–I. 102370, 13.7 mm SL,
Omonawa, Isen; KAUM–I. 102371, 12.2 mm SL, Omonawa, Isen.
FAMILY GIRELLIDAE
Girella leonina (Richardson, 1846)[Jpn name: Kuromejina]
Mochida (2018): KAUM–I. 82442, 203.0 mm SL, Kametoku, Tokunoshima. This study: KAUM–I. 82355,
152.4 mm SL, Kametoku, Tokunoshima; KAUM–I. 82442, 203.0 mm SL, Kametoku, Tokunoshima.
Girella mezina Jordan & Starks, 1907
[Jpn name: Okinamejina]
Mochida (2018): KAUM–I. 82461, 148.3 mm SL, Kametoku, Tokunoshima. This study: KAUM–I. 82356,
160.5 mm SL, Kametoku, Tokunoshima; KAUM–I. 82443, 140.5 mm SL, Kametoku, Tokunoshima;
KAUM–I. 82444, 125.2 mm SL, Kametoku, Tokunoshima; KAUM–I. 82445, 149.1 mm SL, Kametoku,
Tokunoshima; KAUM–I. 82461, 148.3 mm SL, Kametoku, Tokunoshima; KAUM–I. 82462, 167.9 mm SL,
Kametoku, Tokunoshima; KAUM–I. 82463, 152.1 mm SL, Kametoku, Tokunoshima; KAUM–I. 82464,
175.7 mm SL, Kametoku, Tokunoshima; KAUM–I. 82465, 152.3 mm SL, Kametoku, Tokunoshima;
KAUM–I. 82466, 147.2 mm SL, Kametoku, Tokunoshima; KAUM–I. 82467, 165.5 mm SL, Kametoku,
Tokunoshima; KAUM–I. 83469, 143.3 mm SL, Omonawa, Isen; KAUM–I. 98796, 17.4 mm SL, Kametoku,
Tokunoshima; KAUM–I. 105553, 15.3 mm SL, Omonawa, Isen; KAUM–I. 105554, 15.3 mm SL, Omonawa,
Isen; KAUM–I. 105555, 25.5 mm SL, Omonawa, Isen; KAUM–I. 105556, 14.6 mm SL, Omonawa, Isen.
FAMILY LABRIDAE
Anampses caeruleopunctatus Rüppell, 1829
[Jpn name: Buchisusukibera]
Fukui (2018): KAUM–I. KAUM–I. 82422, 143.3 mm SL, San, Tokunoshima. This study: 65933, 208.5 mm
SL, Setaki, Amagi; KAUM–I. 82422, 143.3 mm SL, San, Tokunoshima.
Bodianus mesothorax (Bloch & Schneider, 1801)[Jpn name: Kesagakebera]
Fukui (2018): KAUM–I. 82184, 44.3 mm SL, Nishiagina, Amagi. This study: KAUM–I. 82184, 44.3 mm
SL, Nishiagina, Amagi.

Marine and freshwater fishes of Tokunoshima island

41

Cheilinus trilobatus Lacepède, 1801
[Jpn name: Mitsubamochinouo]
KAUM–I. 66523, 13.4 mm SL, Setaki, Amagi.
Cheilio inermis (Forsskål, 1775)[Jpn name: Kamasubera]
KAUM–I. 105552, 45.0 mm SL, Matsubara, Amagi.
Cirrhilabrus cyanopleura (Bleeker, 1851)[Jpn name: Kuroheriitohikibera]
Fukui (2018): KAUM–I. 82271, 22.3 mm SL, San, Tokunoshima. This study: KAUM–I. 82269, 18.0 mm
SL, San, Tokunoshima; KAUM–I. 82271, 22.3 mm SL, San, Tokunoshima; KAUM–I. 82272, 8.3 mm SL,
San, Tokunoshima.
Coris aygula Lacepède, 1801
[Jpn name: Kammuribera]
KAUM–I. 65852, 73.9 mm SL, Nishiagina, Amagi.
Coris batuensis (Bleeker, 1856)[Jpn name: Shichisemmusumebera]
KAUM–I. 102255, 109.5 mm SL, Omonawa, Isen.
Coris dorsomacula Fowler, 1908
[Jpn name: Sujibera]
Fukui (2018): KAUM–I. 82188, 53.8 mm SL, Nishiagina, Amagi. This study: KAUM–I. 82186, 56.6 mm
SL, Nishiagina, Amagi; KAUM–I. 82188, 53.8 mm SL, Nishiagina, Amagi.
Coris gaimard (Quoy & Gaimard, 1824)[Jpn name: Tsuyubera]
Fukui (2018): KAUM–I. 82292, 43.8 mm SL, San, Tokunoshima. This study: KAUM–I. 82292, 43.8 mm
SL, San, Tokunoshima; KAUM–I. 102054, 41.4 mm SL, Omonawa, Isen.
Epibulus insidiator (Pallas, 1770)[Jpn name: Gichibera]
KAUM–I. 82251, 149.9 mm SL, San, Tokunoshima; KAUM–I. 82374, 76.0 mm SL, San, Tokunoshima.
Halichoeres hortulanus (Lacepède, 1801)[Jpn name: Tokarabera]
Fukui (2018): KAUM–I. 83481, 101.2 mm SL, Kaneku, Amagi. This study: KAUM–I. 66526, 12.4 mm SL,
Setaki, Amagi; KAUM–I. 66527, 22.6 mm SL, Setaki, Amagi; KAUM–I. 66582, 27.7 mm SL, Yonama,
Amagi; KAUM–I. 66583, 21.7 mm SL, Yonama, Amagi; KAUM–I. 82449, 35.3 mm SL, San, Tokunoshima;
KAUM–I. 82518, 32.5 mm SL, San, Tokunoshima; KAUM–I. 82526, 15.1 mm SL, San, Tokunoshima;
KAUM–I. 83481, 101.2 mm SL, Kaneku, Amagi; KAUM–I. 102439, 15.5 mm SL, Omonawa, Isen;
KAUM–I. 105538, 142.9 mm SL, Kaneku, Amagi.
Halichoeres margaritaceus (Valenciennes, 1839)[Jpn name: Akanijibera]
KAUM–I. 102301, 37.9 mm SL, Omonawa, Isen; KAUM–I. 105551, 67.0 mm SL, Inutabu, Isen.

42

I. Mochida and H. Motomura

Halichoeres marginatus Rüppell, 1835
[Jpn name: Kanokobera]
Fukui (2018): KAUM–I. 93448, 125.1 mm SL, Hetono, Amagi. This study: KAUM–I. 14287, 41.2 mm SL,
off Tokunoshima or Amami-oshima islands; KAUM–I. 14288, 39.8 mm SL, off Tokunoshima or Amamioshima islands; KAUM–I. 14289, 27.4 mm SL, off Tokunoshima or Amami-oshima islands; KAUM–I.
82525, 19.3 mm SL, San, Tokunoshima; KAUM–I. 93447, 133.1 mm SL, Hetono, Amagi; KAUM–I. 93448,
125.1 mm SL, Hetono, Amagi; KAUM–I. 102257, 26.0 mm SL, off Tokunoshima or Amami-oshima islands;
KAUM–I. 102401, 20.6 mm SL, Isen, Isen; KAUM–I. 102402, 21.8 mm SL, Asan, Isen; KAUM–I. 102403,
30.9 mm SL, Omonawa, Isen; KAUM–I. 102404, 31.7 mm SL, Omonawa, Isen.
Halichoeres melanochir Fowler & Bean, 1928
[Jpn name: Munatembera]
Kamohara & Yamakawa (1965, as Halichoeres prosopeion): 1 unregistered specimen, 71 mm TL, Matsubara,
Amagi. Fukui (2018): KAUM–I. 82514, 145.2 mm SL, San, Tokunoshima. This study: KAUM–I. 66536,
46.6 mm SL, Setaki, Amagi; KAUM–I. 82223, 64.9 mm SL, Nishiagina, Amagi; KAUM–I. 82418, 125.4mm SL, San, Tokunoshima; KAUM–I. 82514, 145.2 mm SL, San, Tokunoshima; KAUM–I. 93788, 130.2mm SL, Omonawa, Isen; KAUM–I. 97931, 83.4 mm SL, Omonawa, Isen; KAUM–I. 102405, 32.0 mm
SL, Omonawa, Isen; KAUM–I. 105515, 51.1 mm SL, Omonawa, Isen; KAUM–I. 105516, 115.0 mm SL,
Omonawa, Isen.
Halichoeres nebulosus (Valenciennes, 1839)[Jpn name: Inazumabera]
Fukui (2018): KAUM–I. 82219, 30.0 mm SL, Nishiagina, Amagi. This study: KAUM–I. 82219, 30.0 mm
SL, Nishiagina, Amagi.
Halichoeres orientalis Randall, 1999
[Jpn name: Tsukibera]
Fukui (2018): KAUM–I. 66566, 78.4 mm SL, Yonama, Amagi. This study: KAUM–I. 66566, 78.4 mm SL,
Yonama, Amagi.
Halichoeres scapularis (Bennett, 1832)[Jpn name: Seitembera]
Kamohara & Yamakawa (1965): 1 unregistered specimen, 64 mm TL, Matsubara, Amagi.
Halichoeres trimaculatus (Quoy & Gaimard, 1834)[Jpn name: Mitsuboshikyusen]
Fukui (2018): KAUM–I. 82390, 28.4 mm SL, San, Tokunoshima. This study: KAUM–I. 82375, 24.9 mm
SL, San, Tokunoshima; KAUM–I. 82390, 28.4 mm SL, San, Tokunoshima; KAUM–I. 82391, 31.1 mm SL,
San, Tokunoshima; KAUM–I. 82505, 26.1 mm SL, San, Tokunoshima; KAUM–I. 82580, 30.3 mm SL, San,
Tokunoshima; KAUM–I. 89884, 116.4 mm SL, Isen, Isen; KAUM–I. 89887, 90.3 mm SL, Inutabu, Isen;
KAUM–I. 93799, 125.0 mm SL, Omonawa, Isen; KAUM–I. 93800, 114.2 mm SL, Omonawa, Isen; KAUM–
I. 93801, 136.4 mm SL, Omonawa, Isen; KAUM–I. 93802, 130.0 mm SL, Omonawa, Isen; KAUM–I.
98359, 104.6 mm SL, Inutabu, Isen; KAUM–I. 102440, 35.6 mm SL, Asan, Isen.
Hemigymnus fasciatus (Bloch, 1792)[Jpn name: Shimatarekuchibera]
Fukui (2018): KAUM–I. 93466, 180.1 mm SL, Agon, Isen. This study: KAUM–I. 93466, 180.1 mm SL,
Agon, Isen.

Marine and freshwater fishes of Tokunoshima island

43

Hologymnosus doliatus (Lacepède, 1801)[Jpn name: Shirotasukibera]
KPM-NR 28906, underwater photograph, Tokunoshima.
Iniistius evides (Jordan & Richardson, 1909)[Jpn name: Hoshihirabera]
KAUM–I. 82132, 60.9 mm SL, Nishiagina, Amagi.
Iniistius pavo (Valenciennes, 1840)[Jpn name: Hoshitensu]
Fukui (2018): KAUM–I. 82178, 68.4 mm SL, Nishiagina, Amagi. This study: KAUM–I. 82178, 68.4 mm
SL, Nishiagina, Amagi.
Labroides bicolor Fowler & Bean, 1928
[Jpn name: Somewakebera]
Fukui (2018): KAUM–I. 82481, 10.0 mm SL, Kametoku, Tokunoshima. This study: KAUM–I. 82481, 10.0mm SL, Kametoku, Tokunoshima.
Labroides dimidiatus (Valenciennes, 1839)[Jpn name: Honsomewakebera]
Fukui (2018): KAUM–I. 82141, 42.5 mm SL, Nishiagina, Amagi. This study: KAUM–I. 82141, 42.5 mm
SL, Nishiagina, Amagi; KAUM–I. 82260, 32.5 mm SL, San, Tokunoshima; KAUM–I. 82429, 26.7 mm SL,
San, Tokunoshima; KAUM–I. 102224, 43.6 mm SL, Asan, Isen; KAUM–I. 102291, 47.1 mm SL, Inutabu,
Isen; KAUM–I. 102389, 26.7 mm SL, Isen, Isen.
Labroides sp.
[Jpn name: N/A]
KAUM–I. 106579, 15.4 mm SL, Asan, Isen.
Labropsis manabei Schmidt, 1931
[Jpn name: Manabebera]
KAUM–I. 82246, 44.1 mm SL, San, Tokunoshima.
Parajulis sp.
[Jpn name: N/A]
KAUM–I. 82268, 27.4 mm SL, San, Tokunoshima.
Stethojulis bandanensis (Bleeker, 1851)[Jpn name: Akaobibera]
Fukui (2018): KAUM–I. 82452, 30.4 mm SL, San, Tokunoshima. This study: KAUM–I. 82452, 30.4mm SL, San, Tokunoshima; KAUM–I. 89921, 52.4 mm SL, Yonama, Amagi; KAUM–I. 89930, 19.6mm SL, Omonawa, Isen; KAUM–I. 89931, 16.2 mm SL, Omonawa, Isen; KAUM–I. 102422, 39.1 mm
SL, Omonawa, Isen; KAUM–I. 102423, 60.4 mm SL, Omonawa, Isen; KAUM–I. 102444, 47.8 mm SL,
Omonawa, Isen; KAUM–I. 102445, 29.3 mm SL, Omonawa, Isen.
Stethojulis sp.
[Jpn name: N/A]
KAUM–I. 106578, 20.5 mm SL, Omonawa, Isen.

44

I. Mochida and H. Motomura

Thalassoma amblycephalum (Bleeker, 1856)[Jpn name: Kogashirabera]
KAUM–I. 102424, 13.0 mm SL, Asan, Isen; KAUM–I. 102425, 13.0 mm SL, Asan, Isen; KAUM–I. 102426,
13.1 mm SL, Asan, Isen; KAUM–I. 102427, 16.9 mm SL, Asan, Isen; KAUM–I. 102428, 14.0 mm SL, Asan,
Isen; KAUM–I. 102429, 14.0 mm SL, Asan, Isen; KAUM–I. 102430, 13.0 mm SL, Asan, Isen; KAUM–I.
102431, 13.9 mm SL, Asan, Isen; KAUM–I. 102432, 16.5 mm SL, Asan, Isen; KAUM–I. 102433, 16.5 mm
SL, Asan, Isen; KAUM–I. 102434, 15.0 mm SL, Asan, Isen; KAUM–I. 102435, 15.0 mm SL, Asan, Isen;
KAUM–I. 102436, 15.3 mm SL, Asan, Isen; KAUM–I. 102437, 55.5 mm SL, Omonawa, Isen; KAUM–I.
102438, 58.4 mm SL, Omonawa, Isen.
Thalassoma hardwicke (Bennett, 1830)[Jpn name: Senasujibera]
Kamohara & Yamakawa (1965): 1 unregistered specimen, 54 mm TL, Matsubara, Amagi. This study:
KAUM–I. 102446, 18.8 mm SL, Asan, Isen; KAUM–I. 102449, 17.3 mm SL, Asan, Isen.
Thalassoma jansenii (Bleeker, 1856)[Jpn name: Yansemnishikibera]
Kamohara & Yamakawa (1965): 1 unregistered specimen, 112 mm TL, Matsubara, Amagi. This study:
KAUM–I. 102359, 106.3 mm SL, Agon, Isen.
Thalassoma lunare (Linnaeus, 1758)[Jpn name: Otomebera]
KAUM–I. 2141, 51.2 mm SL, Tokunoshima island.
Thalassoma lutescens (Lay & Bennett, 1839)[Jpn name: Yamabukibera]
Kamohara & Yamakawa (1965): 1 unregistered specimen, 82 mm TL, Matsubara, Amagi. Fukui (2018):
KAUM–I. 82238, 109.6 mm SL, San, Tokunoshima. This study: KAUM–I. 82238, 109.6 mm SL, San,
Tokunoshima; KAUM–I. 82244, 109.5 mm SL, San, Tokunoshima; KAUM–I. 82295, 86.6 mm SL, San,
Tokunoshima; KAUM–I. 82414, 71.3 mm SL, San, Tokunoshima; KAUM–I. 82415, 95.7 mm SL, San,
Tokunoshima; KAUM–I. 82416, 102.4 mm SL, San, Tokunoshima; KAUM–I. 82552, 96.4 mm SL,
Omonawa, Isen; KAUM–I. 82562, 105.7 mm SL, Isen, Isen; KAUM–I. 83450, 123.8 mm SL, Isen, Isen;
KAUM–I. 93809, 125.0 mm SL, Omonawa, Isen; KAUM–I. 93810, 90.4 mm SL, Omonawa, Isen; KAUM–
I. 93811, 122.0 mm SL, Omonawa, Isen; KAUM–I. 93812, 102.5 mm SL, Omonawa, Isen; KAUM–I. 93844,
131.0 mm SL, Agon, Isen; KAUM–I. 93845, 105.4 mm SL, Agon, Isen; KAUM–I. 93864, 134.0 mm SL,
Kaneku, Amagi; KAUM–I. 93865, 107.2 mm SL, Kaneku, Amagi; KAUM–I. 102441, 14.4 mm SL, Asan,
Isen; KAUM–I. 102442, 14.4 mm SL, Asan, Isen; KAUM–I. 102443, 19.4 mm SL, Asan, Isen; KAUM–
I. 105539, 128.8 mm SL, Kaneku, Amagi; KAUM–I. 105542, 80.3 mm SL, Kaneku, Amagi; KAUM–I.
105546, 82.0 mm SL, Omonawa, Isen.
Thalassoma purpureum (Forsskål, 1775)[Jpn name: Kinubera]
KAUM–I. 83459, 16.3 mm SL, Isen, Isen; KAUM–I. 83460, 16.2 mm SL, Isen, Isen; KAUM–I. 102448,
14.3 mm SL, Asan, Isen.
Thalassoma sp.
[Jpn name: N/A]
KAUM–I. 102447, 14.6 mm SL, Asan, Isen.

Marine and freshwater fishes of Tokunoshima island

45

Wetmorella nigropinnata (Seale, 1901)[Jpn name: Hashinagabera]
Fukui (2018): KAUM–I. 66560, 11.6 mm SL, Setaki, Amagi. This study: KAUM–I. 66560, 11.6 mm SL,
Setaki, Amagi.
FAMILY SCARIDAE
Chlorurus microrhinos (Bleeker, 1854)[Jpn name: Nan-yobudai]
KAUM–I. 82517, 33.1 mm SL, San, Tokunoshima.
Chlorurus sordidus (Forsskål, 1775)[Jpn name: Hagebudai]
KAUM–I. 102160, 43.7 mm SL, Omonawa, Isen; KAUM–I. 106570, 32.5 mm SL, Omonawa, Isen;
KAUM–I. 106573, 30.8 mm SL, Yonama, Amagi.
Chlorurus sp.
[Jpn name: N/A]
KAUM–I. 106574, 17.7 mm SL, Asan, Isen; KAUM–I. 106575, 19.6 mm SL, Asan, Isen; KAUM–I. 106576,
16.1 mm SL, Asan, Isen.
Leptoscarus vaigiensis (Quoy & Gaimard, 1824)[Jpn name: Mizorebudai]
KAUM–I. 65895, 33.9 mm SL, Nishiagina, Amagi; KAUM–I. 82145, 32.7 mm SL, Nishiagina, Amagi;
KAUM–I. 82392, 34.7 mm SL, San, Tokunoshima; KAUM–I. 106566, 38.8 mm SL, Omonawa, Isen.
Scarus prasiognathos Valenciennes, 1840
[Jpn name: Nishikibudai]
KAUM–I. 106571, 38.6 mm SL, Omonawa, Isen.
Scarus rivulatus Valenciennes, 1840
[Jpn name: Sujibudai]
KAUM–I. 82446, 202.2 mm SL, Kametoku, Tokunoshima; KAUM–I. 82530, 149.6 mm SL, Omonawa,
Isen.
Scarus rubroviolaceus Bleeker, 1847
[Jpn name: Nagabudai]
KAUM–I. 89928, 24.0 mm SL, Yonama, Amagi.
Scarus schlegeli (Bleeker, 1861)[Jpn name: Obibudai]
KAUM–I. 106558, 58.4 mm SL, Yonama, Amagi; KAUM–I. 106559, 63.3 mm SL, Yonama, Amagi;
KAUM–I. 106569, 40.3 mm SL, Omonawa, Isen.
Scarus sp.
[Jpn name: N/A]
KAUM–I. 106572, 32.9 mm SL, Omonawa, Isen.

46

I. Mochida and H. Motomura

FAMILY PINGUIPEDIDAE
Parapercis clathrata Ogilby, 1910
[Jpn name: Yotsumetoragisu]
KAUM–I. 82220, 62.1 mm SL, Nishiagina, Amagi.
Parapercis cylindrica (Bloch, 1792)[Jpn name: Dandaratoragisu]
Matsunuma (2018e): KAUM–I. 82435, 59.0 mm SL, San, Tokunoshima. This study: KAUM–I. 82396, 70.4mm SL, San, Tokunoshima; KAUM–I. 82435, 59.0 mm SL, San, Tokunoshima; KAUM–I. 82456, 65.7 mm
SL, San, Tokunoshima; KAUM–I. 82457, 60.8 mm SL, San, Tokunoshima; KAUM–I. 82576, 75.3 mm SL,
San, Tokunoshima; KAUM–I. 82577, 108.7 mm SL, San, Tokunoshima; KAUM–I. 82578, 56.9 mm SL,
San, Tokunoshima; KAUM–I. 89913, 19.5 mm SL, Omonawa, Isen; KAUM–I. 93842, 98.4 mm SL, Boma,
Tokunoshima; KAUM–I. 93843, 108.0 mm SL, Boma, Tokunoshima; KAUM–I. 102046, 69.6 mm SL,
Inutabu, Isen.
Parapercis millepunctata (Günther, 1860)[Jpn name: Wanuketoragisu]
KAUM–I. 102087, 140.0 mm SL, Inutabu, Isen; KAUM–I. 102100, 103.2 mm SL, Inutabu, Isen.
Parapercis pacifica Imamura & Yoshino, 2007
[Jpn name: Ogurotoragisu]
Matsunuma (2018e): KAUM–I. 82365, 131.1 mm SL, Kametoku, Tokunoshima. This study: KAUM–I.
82365, 131.1 mm SL, Kametoku, Tokunoshima; KAUM–I. 93787, 156.4 mm SL, Omonawa, Isen; KAUM–
I. 102333, 41.1 mm SL, Isen, Isen.
Parapercis tetracantha (Lacepède, 1801)[Jpn name: Madaratoragisu]
Matsunuma (2018e): KAUM–I. 82279, 113.6 mm SL, San, Tokunoshima. This study: KAUM–I. 82279,
113.6 mm SL, San, Tokunoshima.
FAMILY CREEDIIDAE
Limnichthys fasciatus Waite, 1904
[Jpn name: Tobigimpo]
Tashiro (2018a): KAUM–I. 82236, 14.8 mm SL, Nishiagina, Amagi. This study: KAUM–I. 82236, 14.8 mm
SL, Nishiagina, Amagi; KAUM–I. 83751, 27.5 mm SL, Omonawa, Isen.
FAMILY TRIPTERYGIIDAE
Enneapterygius etheostoma (Jordan & Snyder, 1902)[Jpn name: Hebigimpo]
KAUM–I. 2097, 43.3 mm SL, Tokunoshima island; KAUM–I. 4324, 44.9 mm SL, Tokunoshima island;
KAUM–I. 4325, 39.8 mm SL, Tokunoshima island; KAUM–I. 102069, 33.5 mm SL, Asan, Isen; KAUM–I.
102396, 42.6 mm SL, Furusato, Isen; KAUM–I. 102397, 36.8 mm SL, Omonawa, Isen; KAUM–I. 105550,
45.5 mm SL, off Tokunoshima or Amami-oshima islands.
Enneapterygius flavoccipitis Shen, 1994
[Jpn name: Erimakihebigimpo]
KAUM–I. 66531, 23.3 mm SL, Setaki, Amagi.

Marine and freshwater fishes of Tokunoshima island

47

Enneapterygius philippinus (Peters, 1868)[Jpn name: Kusagimpo]
KAUM–I. 102398, 20.6 mm SL, Isen, Isen; KAUM–I. 102394, 21.4 mm SL, Omonawa, Isen; KAUM–I.
102395, 19.1 mm SL, Omonawa, Isen.
Enneapterygius phoenicosoma Motomura, Ota & Meguro, 2015
[Jpn name: Akahebigimpo]
Tashiro (2018b): KAUM–I. 82455, 25.2 mm SL, San, Tokunoshima. This study: KAUM–I. 66532, 22.0 mm
SL, Setaki, Amagi; KAUM–I. 66589, 23.9 mm SL, Yonama, Amagi; KAUM–I. 82455, 25.2 mm SL, San,
Tokunoshima.
Enneapterygius sp.
[Jpn name: N/A]
KAUM–I. 82161, 17.0 mm SL, Nishiagina, Amagi.
Enneapterygius tutuilae Jordan & Seale, 1906
[Jpn name: Segurohebigimpo]
KAUM–I. 65905, 16.0 mm SL, Nishiagina, Amagi; KAUM–I. 66590, 18.2 mm SL, Yonama, Amagi.
Helcogramma inclinata (Fowler, 1946)[Jpn name: Ayahebigimpo]
KAUM–I. 86776, 38.8 mm SL, Isen, Isen.
Ucla xenogrammus Holleman, 1993
[Jpn name: Kasurihebigimpo]
Tashiro (2018b): KAUM–I. 66533, 45.7 mm SL, Setaki, Amagi. This study: KAUM–I. 66533, 45.7 mm SL,
Setaki, Amagi.
FAMILY CLINIDAE
Springeratus xanthosoma (Bleeker, 1857)[Jpn name: Asahigimpo]
KAUM–I. 10588, 63.5 mm SL, Tokunoshima island.
FAMILY BLENNIIDAE
Alticus saliens (Forster, 1788)[Jpn name: Tamakaeruuo]
Kawama (2018): KAUM–I. 82542, 62.8 mm SL, Isen, Isen. This study: KAUM–I. 82537, 61.3 mm SL, Isen,
Isen; KAUM–I. 82540, 55.4 mm SL, Isen, Isen; KAUM–I. 82541, 66.7 mm SL, Isen, Isen; KAUM–I. 82542,
62.8 mm SL, Isen, Isen; KAUM–I. 105568, 47.8 mm SL, off Tokunoshima or Amami-oshima islands.
Aspidontus taeniatus taeniatus Quoy & Gaimard, 1834
[Jpn name: Nisekurosujigimpo]
KAUM–I. 102335, 48.7 mm SL, San, Tokunoshima.
Atrosalarias fuscus holomelas (Günther, 1872)[Jpn name: Indokaeruuo]
Kawama (2018): KAUM–I. 66568, 37.8 mm SL, Yonama, Amagi. This study: KAUM–I. 66568, 37.8 mm
SL, Yonama, Amagi; KAUM–I. 82362, 30.2 mm SL, San, Tokunoshima.

48

I. Mochida and H. Motomura

Cirripectes imitator (Valenciennes, 1836)[Jpn name: Amimeminokaeruuo]
KAUM–I. 13891, 68.1 mm SL, Tokunoshima island; KAUM–I. 13892, 70.0 mm SL, Tokunoshima island.
Ecsenius yaeyamaensis (Aoyagi, 1954)[Jpn name: Ishigakikaeruuo]
KAUM–I. 82436, 37.7 mm SL, San, Tokunoshima; KAUM–I. 102334, 41.8 mm SL, Omonawa, Isen.
Enchelyurus kraussii (Klunzinger, 1871)[Jpn name: Kurogimpo]
Kawama (2018): KAUM–I. 82360, 35.6 mm SL, San, Tokunoshima. This study: KAUM–I. 82360, 35.6 mm
SL, San, Tokunoshima.
Entomacrodus decussatus (Bleeker, 1858)[Jpn name: Kikaikaeruuo]
Kawama (2018): KAUM–I. 82538, 30.8 mm SL, Isen, Isen. This study: KAUM–I. 82538, 30.8 mm SL, Isen,
Isen; KAUM–I. 102218, 59.2 mm SL, Isen, Isen.
Entomacrodus striatus (Valenciennes, 1836)[Jpn name: Sujigimpo]
Kawama (2018): KAUM–I. 83708, 54.9 mm SL, Isen, Isen. This study: KAUM–I. 82539, 30.8 mm SL,
Isen, Isen; KAUM–I. 83705, 57.1 mm SL, Isen, Isen; KAUM–I. 83706, 44.3 mm SL, Isen, Isen; KAUM–I.
83707, 40.9 mm SL, Isen, Isen; KAUM–I. 83708, 54.9 mm SL, Isen, Isen; KAUM–I. 83709, 22.2 mm SL,
Isen, Isen; KAUM–I. 83710, 41.2 mm SL, Isen, Isen; KAUM–I. 102068, 42.5 mm SL, Isen, Isen; KAUM–I.
102327, 25.3 mm SL, Isen, Isen.
Exallias brevis (Kner, 1868)[Jpn name: Sedakagimpo]
KAUM–I. 66574, 44.2 mm SL, Yonama, Amagi.
Istiblennius edentulus (Forster & Schneider, 1801)[Jpn name: Nisekaeruuo]
Kawama (2018): KAUM–I. 82305, 45.3 mm SL, Asan, Isen. This study: KAUM–I. 82305, 45.3 mm SL,
Asan, Isen; KAUM–I. 82378, 23.7 mm SL, San, Tokunoshima; KAUM–I. 82379, 22.2 mm SL, San,
Tokunoshima; KAUM–I. 82380, 37.7 mm SL, San, Tokunoshima; KAUM–I. 82381, 39.6 mm SL, San,
Tokunoshima; KAUM–I. 82382, 21.2 mm SL, San, Tokunoshima; KAUM–I. 82383, 74.2 mm SL, San,
Tokunoshima; KAUM–I. 82384, 55.9 mm SL, San, Tokunoshima; KAUM–I. 88363, 92 mm SL, Inutabu,
Isen; KAUM–I. 98768, 31.6 mm SL, Kembuku, Isen; KAUM–I. 98769, 42.7 mm SL, Kembuku, Isen;
KAUM–I. 102008, 84.7 mm SL, Kinen, Isen; KAUM–I. 102009, 94.6 mm SL, Kinen, Isen; KAUM–I.
102388, 23 mm SL, Isen, Isen.
Istiblennius enosimae (Jordan & Snyder, 1902)[Jpn name: Kaeruuo]
KAUM–I. 2069, 104.6 mm SL, Tokunoshima island.
Istiblennius lineatus (Valenciennes, 1836)[Jpn name: Senkaeruuo]
Kawama (2018): KAUM–I. 82385, 50.7 mm SL, San, Tokunoshima. This study: KAUM–I. 82385, 50.7 mm
SL, San, Tokunoshima.

Marine and freshwater fishes of Tokunoshima island

49

Meiacanthus atrodorsalis (Günther, 1877)[Jpn name: Ogonnijigimpo]
Kawama (2018): KAUM–I. 91368, 28.8 mm SL, Omonawa, Isen. This study: KAUM–I. 91368, 28.8 mm
SL, Omonawa, Isen; KAUM–I. 102181, 26.0 mm SL, Omonawa, Isen.
Meiacanthus grammistes (Valenciennes, 1836)[Jpn name: Higenijigimpo]
KAUM–I. 102151, 22.5 mm SL, Omonawa, Isen; KAUM–I. 102332, 39.3 mm SL, Kanami, Tokunoshima;
KAUM–I. 102407, 33.3 mm SL, Kanami, Tokunoshima; KAUM–I. 105514, 51.5 mm SL, Omonawa, Isen.
Meiacanthus kamoharai Tomiyama, 1956
[Jpn name: Kamoharagimpo]
Kawama (2018): KAUM–I. 82370, 62.4 mm SL, Kametoku, Tokunoshima. This study: KAUM–I. 82370,
62.4 mm SL, Kametoku, Tokunoshima.
Petroscirtes breviceps (Valenciennes, 1836)[Jpn name: Nijigimpo]
Kawama (2018): KAUM–I. 82288, 37.0 mm SL, San, Tokunoshima. This study: KAUM–I. 65854, 54.0mm SL, Nishiagina, Amagi; KAUM–I. 82261, 20.7 mm SL, San, Tokunoshima; KAUM–I. 82273, 18.8 mm
SL, San, Tokunoshima; KAUM–I. 82285, 18.6 mm SL, San, Tokunoshima; KAUM–I. 82286, 24.0 mm SL,
San, Tokunoshima; KAUM–I. 82287, 15.5 mm SL, San, Tokunoshima; KAUM–I. 82288, 37.0 mm SL, San,
Tokunoshima; KAUM–I. 102212, 32.0 mm SL, Kanami, Tokunoshima; KAUM–I. 102282, 36.0 mm SL,
Omonawa, Isen.
Petroscirtes mitratus Rüppell, 1830
[Jpn name: Hatatategimpo]
Kawama (2018): KAUM–I. 82548, 49.6 mm SL, Kametoku, Tokunoshima. This study: KAUM–I. 82524,
18.0 mm SL, San, Tokunoshima; KAUM–I. 82548, 49.6 mm SL, Kametoku, Tokunoshima; KAUM–I.
102000, 40.0 mm SL, Omonawa, Isen; KAUM–I. 102066, 33.0 mm SL, Omonawa, Isen; KAUM–I. 102372,
34.7 mm SL, Omonawa, Isen; KAUM–I. 102373, 39.2 mm SL, Boma, Tokunoshima; KAUM–I. 102375,
30.6 mm SL, Omonawa, Isen; KAUM–I. 102385, 30.8 mm SL, Omonawa, Isen; KAUM–I. 102412, 22.0mm SL, Omonawa, Isen; KAUM–I. 102418, 17.4 mm SL, Omonawa, Isen.
Plagiotremus tapeinosoma (Bleeker, 1857)[Jpn name: Tenkurosujigimpo]
Kawama (2018): KAUM–I. 82425, 73.6 mm SL, San, Tokunoshima. This study: KAUM–I. 82425, 73.6 mm
SL, San, Tokunoshima; KAUM–I. 102213, 41.0 mm SL, Omonawa, Isen.
Praealticus bilineatus (Peters, 1868)[Jpn name: Tamagimpo]
Kawama (2018): KAUM–I. 93741, 44.1 mm SL, Isen, Isen. This study: KAUM–I. 93740, 35.6 mm SL,
Isen, Isen; KAUM–I. 93741, 44.1 mm SL, Isen, Isen; KAUM–I. 93742, 43.2 mm SL, Isen, Isen; KAUM–I.
93743, 40.0 mm SL, Isen, Isen; KAUM–I. 93848, 34.0 mm SL, Isen, Isen; KAUM–I. 93849, 27.1 mm SL,
Isen, Isen; KAUM–I. 98785, 45.3 mm SL, Kembuku, Isen; KAUM–I. 98786, 40.2 mm SL, Kembuku, Isen;
KAUM–I. 102154, 49.1 mm SL, Isen, Isen; KAUM–I. 102155, 42.4 mm SL, Isen, Isen.
Praealticus tanegasimae (Jordan & Starks, 1906)[Jpn name: Tanegimpo]
KAUM–I. 102209, 41.7 mm SL, Isen, Isen.

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Salarias fasciatus (Bloch, 1786)[Jpn name: Yaeyamagimpo]
Kawama (2018): KAUM–I. 82428, 108.4 mm SL, San, Tokunoshima. This study: KAUM–I. 82428, 108.4mm SL, San, Tokunoshima; KAUM–I. 83672, 57.2 mm SL, Omonawa, Isen; KAUM–I. 88362, 75.2 mm SL,
Inutabu, Isen; KAUM–I. 102165, 79.0 mm SL, Kembuku, Isen; USNM 297646, 89.9 mm SL, Tokunoshimaisland.
Salarias luctuosus Whitley, 1929
[Jpn name: Shimagimpo]
KAUM–I. 102067, 40.5 mm SL, Omonawa, Isen; KAUM–I. 102352, 21.8 mm SL, Omonawa, Isen.
Salarias sinuosus Snyder, 1908
[Jpn name: Hirenagakaeruuo]
KAUM–I. 10583, 44.1 mm SL, Tokunoshima island; KAUM–I. 10584, 47.1 mm SL, Tokunoshima island;
KAUM–I. 10585, 42.3 mm SL, Tokunoshima island; KAUM–I. 10586, 40.3 mm SL, Tokunoshima island;
KAUM–I. 10587, 40.6 mm SL, Tokunoshima island.
FAMILY GOBIESOCIDAE
Conidens laticephalus (Tanaka, 1909)[Jpn name: Ankoubauo]
Fujiwara (2018b): KAUM–I. 65898, 31.9 mm SL, Nishiagina, Amagi. This study: KAUM–I. 65898, 31.9mm SL, Nishiagina, Amagi.
FAMILY CALLIONYMIDAE
Diplogrammus goramensis (Bleeker, 1858)[Jpn name: Minamikobunumeri]
KAUM–I. 102374, 39.7 mm SL, Omonawa, Isen.
Neosynchiropus morrisoni (Schultz, 1960)[Jpn name: Sesokoteguri]
Iwatsubo (2018b): KAUM–I. 82264, 35.8 mm SL, San, Tokunoshima. This study: KAUM–I. 82264, 35.8mm SL, San, Tokunoshima; KPM-NR 15798, underwater photograph, Tokunoshima.
Neosynchiropus ocellatus (Pallas, 1770)[Jpn name: Kouwanteguri]
Iwatsubo (2018b): KAUM–I. 82230, 49.0 mm SL, Nishiagina, Amagi. This study: KAUM–I. 82230, 49.0mm SL, Nishiagina, Amagi; KAUM–I. 102158, 70.9 mm SL, Inutabu, Isen; KAUM–I. 102419, 29.3 mm SL,
Omonawa, Isen; KPM-NR 35915, underwater photograph, Tokunoshima.
FAMILY ELEOTRIDAE
Eleotris acanthopoma Bleeker, 1853
[Jpn name: Chichibumodoki]Akihito (1967): 5 specimens, 52–86 mm SL, Kametsu, Tokunoshima. Ike et al. (1990): 5 unregisteredspecimens (not retained), Kametoku River; 6 unregistered specimens (not retained), Ose River; 17
unregistered specimens (not retained), Minato River; 1 unregistered specimen (not retained), Akirigami
River. This study: KAUM–I. 98126, 32.2 mm SL, Akirigami River; KAUM–I. 98127, 18.9 mm SL,
Akirigami River; KAUM–I. 98128, 17.8 mm SL, Akirigami River; KAUM–I. 98129, 15.5 mm SL,
Akirigami River; KAUM–I. 102164, 61.3 mm SL, Takou River; KAUM–I. 102206, 44.7 mm SL, Wan-ya

Marine and freshwater fishes of Tokunoshima island

51

River; KAUM–I. 102217, 53.2 mm SL, Omonawa, Isen; KAUM–I. 105569, 51.9 mm SL, off Tokunoshima
or Amami-oshima islands; KAUM–I. 105596, 25.9 mm SL, Takou River; KAUM–I. 105570, 34.7 mm SL,
Kametoku River.
Eleotris fusca (Bloch & Schneider, 1801)[Jpn name: Tenjikukawaanago]
Ogawa (1937): specimen lost, Tokunoshima island. Akihito (1967): 1 specimen, 79 mm SL, Tokunoshimaisland. Ike et al. (1990): 2 unregistered specimens (not retained), Akirigami River. Hagiwara (2018a):
KAUM–I. 82301, 78.2 mm SL, Asan, Isen. This study: KAUM–I. 82300, 81.0 mm SL, Asan, Isen; KAUM–
I. 82301, 78.2 mm SL, Asan, Isen; KAUM–I. 82302, 52.1 mm SL, Asan, Isen; KAUM–I. 82303, 25.4 mm
SL, Asan, Isen; KAUM–I. 82304, 25.9 mm SL, Asan, Isen.
Eleotris melanosoma Bleeker, 1853
[Jpn name: Okamehaze]
Ike et al. (1990): 1 unregistered specimen (not retained), Ose River.
Eleotris sp. 1[Jpn name: N/A] (Fig. 4B)
KAUM–I. 65913, 144.1 mm SL, Asama, Amagi; KAUM–I. 66562, 147.5 mm SL, Asama, Amagi.
Remarks: These specimens were collected from the anchialine cave in Asama along with Xenoconger fryeri.
The specimens probably represent the same species recorded as Eleotris sp. from caves on Okinoerabujima island (Yoshigou et al., 2005: pl. 2, figs. 10, 11), and on Miyakojima island and the Yaeyama Islands(Yoshigou, 2014).
Eleotris sp. 2[Jpn name: N/A]
KAUM–I. 98764, 68.4 mm SL, Akirigami River; KAUM–I. 98765, 31.4 mm SL, Akirigami River; KAUM–
I. 98766, 31.0 mm SL, Akirigami River; KAUM–I. 98767, 43.3 mm SL, Akirigami River; KAUM–I. 98793,
23.3 mm SL, Akirigami River; KAUM–I. 98794, 17.3 mm SL, Akirigami River; KAUM–I. 98795, 16.3 mm
SL, Akirigami River.
Ophieleotris sp.
[Jpn name: Tametomohaze]
Ike et al. (1990): 1 unregistered specimen (not retained), Kametoku River.
FAMILY XENISTHMIDAE
Xenisthmus clarus (Jordan & Seale, 1906)[Jpn name: Yanagihaze]
Akihito (1969): MIKU 1763, 33 mm SL, Kametsu, Tokunoshima.
FAMILY GOBIIDAE
Acentrogobius virgatulus (Jordan & Snyder, 1901)[Jpn name: Sujihaze]
Ike et al. (1990, as Acentrogobius pflaumii): 9 unregistered specimens (not retained), Ose River; 2unregistered specimens (not retained), Minato River; 1 unregistered specimen (not retained), Wan-ya River.

52

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Amblygobius phalaena (Valenciennes, 1837)[Jpn name: Sarasahaze]
Tomiyama (1936): specimen(s) (probably registered in Imperial University, Tokyo), Tokunoshima island.
Hagiwara (2018b): KAUM–I. 82398, 53.1 mm SL, San, Tokunoshima. This study: KAUM–I. 82394, 104.1mm SL, San, Tokunoshima; KAUM–I. 82398, 53.1 mm SL, San, Tokunoshima; KAUM–I. 82579, 89.9 mm
SL, San, Tokunoshima; KAUM–I. 105521, 33.6 mm SL, Yonama, Amagi.
Asterropteryx semipunctata Rüppell, 1830
[Jpn name: Hoshihaze]
Hagiwara (2018b): KAUM–I. 82359, 24.5 mm SL, San, Tokunoshima. This study: KAUM–I. 82359, 24.5mm SL, San, Tokunoshima; KAUM–I. 82399, 20.9 mm SL, San, Tokunoshima; KAUM–I. 82400, 19.3 mm
SL, San, Tokunoshima; KAUM–I. 82401, 27.5 mm SL, San, Tokunoshima; KAUM–I. 82402, 19.8 mm SL,
San, Tokunoshima; KAUM–I. 82403, 22.1 mm SL, San, Tokunoshima; KAUM–I. 82407, 17.8 mm SL, San,
Tokunoshima; KAUM–I. 82408, 14.9 mm SL, San, Tokunoshima; KAUM–I. 82410, 18.3 mm SL, San,
Tokunoshima; KAUM–I. 82411, 20.3 mm SL, San, Tokunoshima; KAUM–I. 82412, 17.3 mm SL, San,
Tokunoshima; KAUM–I. 106523, 22.6 mm SL, Omonawa, Isen; KAUM–I. 106524, 19.2 mm SL, Omonawa,
Isen; KAUM–I. 106525, 19.4 mm SL, Omonawa, Isen.
Awaous melanocephalus (Bleeker, 1849)[Jpn name: Kurominamihaze]
Ike et al. (1990): 1 unregistered specimen (not retained), Ose River; 1 unregistered specimen (not retained),
Akirigami River.
Bathygobius coalitus (Bennett, 1832)[Jpn name: Kuroyahazuhaze]
Hagiwara (2018b): KAUM–I. 93746, 59.4 mm SL, Isen, Isen. This study: KAUM–I. 93746, 59.4 mm SL,
Isen, Isen; KAUM–I. 98787, 49.1 mm SL, Kembuku, Isen; KAUM–I. 98791, 31.2 mm SL, Kembuku, Isen;
KAUM–I. 98792, 25.0 mm SL, Kembuku, Isen; KAUM–I. 102221, 40.1 mm SL, Isen, Isen; KAUM–I.
105564, 41.0 mm SL, Matsubara, Amagi; KAUM–I. 105565, 33.6 mm SL, Matsubara, Amagi; KAUM–I.
105630, 33.4 mm SL, Matsubara, Amagi; KAUM–I. 105631, 32.2 mm SL, Matsubara, Amagi; KAUM–I.
105632, 38.3 mm SL, Matsubara, Amagi.
Bathygobius cocosensis (Bleeker, 1854)[Jpn name: Sujikumohaze]
Akihito & Meguro (1980): LICPP 1967077, 4 specimens, 24–35 mm SL, Hon River; LICPP 1972052,
3 specimens, 17–22 mm SL, mouth of Omonawa River. Ike et al. (1990): 1 unregistered specimen (notretained), Ose River; 1 unregistered specimen (not retained), Minato River. Hagiwara (2018b): KAUM–I.
82306, 29.8 mm SL, Asan, Isen. This study: KAUM–I. 82306, 29.8 mm SL, Asan, Isen; KAUM–I. 82307,
108.8 mm SL, Asan, Isen; KAUM–I. 83881, 30.3 mm SL, Omonawa, Isen; KAUM–I. 86775, 24.8 mm SL,
Yonama, Amagi; KAUM–I. 102191, 61.8 mm SL, Minato River; KAUM–I. 102192, 41.7 mm SL, Minato
River.
Bathygobius cyclopterus (Valenciennes, 1837)[Jpn name: Yahazuhaze]
Akihito & Meguro (1980): LICPP 1972054, 3 specimens, 23–31 mm SL, mouth of Omonawa River.
Hagiwara (2018b): KAUM–I. 82386, 50.4 mm SL, San, Tokunoshima. This study: KAUM–I. 82386, 50.4mm SL, San, Tokunoshima; KAUM–I. 98788, 19.4 mm SL, Kembuku, Isen; KAUM–I. 98790, 23.3 mm SL,
Kembuku, Isen.

Marine and freshwater fishes of Tokunoshima island

53

Bathygobius fuscus (Rüppell, 1830)[Jpn name: Kumohaze]
Ike et al. (1990): 1 unregistered specimen (not retained), Ose River; 8 unregistered specimens (not retained),
Minato River. This study: KAUM–I. 102207, 52.3 mm SL, Omonawa, Isen; KAUM–I. 105566, 40.3 mm
SL, Wan-ya River; KAUM–I. 105635, 17.9 mm SL, Omonawa, Isen; KAUM–I. 105636, 17.3 mm SL,
Omonawa, Isen; KAUM–I. 105637, 11.5 mm SL, Omonawa, Isen.
Bathygobius hongkongensis (Fowler, 1931)[Jpn name: Kurohoshiyahazuhaze]
Akihito & Meguro (1980): LICPP 1971008, 2 specimens, 51–59 mm SL, Kaneku, Amagi; LICPP 1971079,
3 specimens, 46–59 mm SK, Hetono, Amagi. This study: KAUM–I. 82387, 44.1 mm SL, San, Tokunoshima.
Bathygobius sp.
[Jpn name: N/A]
KAUM–I. 106564, 14.6 mm SL, Isen, Isen.
Bryaninops loki Larson, 1985
[Jpn name: Hosogarasuhaze]
Hagiwara (2018b): KAUM–I. 82296, 20.5 mm SL, San, Tokunoshima. This study: KAUM–I. 82296, 20.5mm SL, San, Tokunoshima.
Cabillus lacertops Smith, 1959
[Jpn name: Mikagehaze]
KAUM–I. 89911, 32.4 mm SL, Yonama, Amagi.
Cabillus tongarevae (Fowler, 1927)[Jpn name: Yorimehaze]
KAUM–I. 98789, 15.3 mm SL, Kembuku, Isen.
Callogobius hasseltii (Bleeker, 1851)[Jpn name: Okinawahaze]
KAUM–I. 98379, 24.4 mm SL, Asan, Isen.
Callogobius okinawae (Snyder, 1908)[Jpn name: Namerahaze]
Akihito & Meguro (1975b): LICPP 1967040, 21 mm SL, Hon River. This study: KAUM–I. 82388, 26.4 mm
SL, San, Tokunoshima; KAUM–I. 102153, 52.2 mm SL, Matsubara, Amagi; KAUM–I. 105522, 40.4 mm
SL, Omonawa, Isen. KAUM–I; 105528, 24.5 mm SL, Asan, Isen.
Callogobius tanegasimae (Snyder, 1908)[Jpn name: Tanehaze]
Ike et al. (1990): 3 unregistered specimens (not retained), Minato River.
Cryptocentrus caeruleomaculatus (Herre, 1933)[Jpn name: Takanohahaze]
Yanagisawa (1977, as Mars caeruleomaculatus), SMBL-F 73424, 36.7 mm SL, Kametoku, Tokunoshima;
additional specimen(s) (catalog number and size not mentioned), Kametoku, Tokunoshima.

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Discordipinna griessingeri Hoese & Fourmanoir, 1978
[Jpn name: Homurahaze]
Hagiwara (2018b): KAUM–I. 82266, 18.2 mm SL, San, Tokunoshima. This study: KAUM–I. 82266, 18.2mm SL, San, Tokunoshima.
Drombus sp.
[Jpn name: Kurokohaze]
Ike et al. (1990): 9 unregistered specimens (not retained), Ose River; 4 unregistered specimens (not retained),
Minato River. This study: KAUM–I. 106563, 20.8 mm SL, Akirigami River.
Eviota abax (Jordan & Snyder, 1901)[Jpn name: Isohaze]
Akihito (1969): 5 unregistered specimens, 13–19 mm SL, Tokunoshima island. This study: KAUM–I. 82405,
14.3 mm SL, San, Tokunoshima.
Eviota afelei Jordan & Seale, 1906
[Jpn name: Shimamidorihaze]
KAUM–I. 106561, 19.5 mm SL, Omonawa, Isen.
Eviota melasma Lachner & Karnella, 1980
[Jpn name: Akahoshiisohaze]
KAUM–I. 82270, 18.4 mm SL, San, Tokunoshima.
Eviota punctulata Jewett & Lachner, 1983
[Jpn name: Mitsubaisohaze]
KAUM–I. 89925, 16.0 mm SL, Omonawa, Isen.
Eviota saipanensis Fowler, 1945
[Jpn name: Musujiisohaze]
Hagiwara (2018b): KAUM–I. 82534, 13.3 mm SL, Isen, Isen. This study: KAUM–I. 82534, 13.3 mm SL,
Isen, Isen.
Eviota smaragdus Jordan & Seale, 1906
[Jpn name: Kurohoshiisohaze]
KAUM–I. 89924, 19.0 mm SL, Omonawa, Isen.
Eviota sp.
[Jpn name: N/A]
KAUM–I. 65906, 11.4 mm SL, Nishiagina, Amagi; KAUM–I. 106562, 18.9 mm SL, Isen, Isen.
Eviota toshiyuki Greenfield & Randall, 2010
[Jpn name: Midorihaze]
Akihito (1969, as Eviota zonura): 1 unregistered specimen, 22 mm SL, Kanami, Tokunoshima.
Favonigobius gymnauchen (Bleeker, 1860)[Jpn name: Himehaze]
Ike et al. (1990): 4 unregistered specimens (not retained), Kametoku River; 1 unregistered specimen (notretained), Minato River; 1 unregistered specimen (not retained), Wan-ya River. Hagiwara (2018b): KAUM–I.

Marine and freshwater fishes of Tokunoshima island

55

83875, 49.6 mm SL, Omonawa, Isen. This study: KAUM–I. 83875, 49.6 mm SL, Omonawa, Isen; KAUM–I.
98125, 52.5 mm SL, Wan-ya River; KAUM–I. 102205, 41.7 mm SL, Minato River; KAUM–I. 105575, 43.2mm SL, Wan-ya River; KAUM–I. 105576, 54.5 mm SL, Wan-ya River.
Favonigobius reichei (Bleeker, 1853)[Jpn name: Minamihimehaze]
Ike et al. (1990): 1 unregistered specimen (not retained), Minato River. This study: KAUM–I. 102300, 45.1mm SL, Wan-ya River.
Fusigobius duospilus Hoese & Reader, 1985
[Jpn name: Sehoshisankakuhaze]
Hagiwara (2018b): KAUM–I. 82221, 33.7 mm SL, Nishiagina, Amagi. This study: KAUM–I. 65886, 31.9mm SL, Nishiagina, Amagi; KAUM–I. 65893, 33.5 mm SL, Nishiagina, Amagi; KAUM–I. 66517, 13.4 mm
SL, Setaki, Amagi; KAUM–I. 66518, 17.7 mm SL, Setaki, Amagi; KAUM–I. 66529, 22.9 mm SL, Setaki,
Amagi; KAUM–I. 66530, 39.9 mm SL, Setaki, Amagi; KAUM–I. 66534, 33.1 mm SL, Setaki, Amagi;
KAUM–I. 66545, 32.4 mm SL, Setaki, Amagi; KAUM–I. 82167, 38.8 mm SL, Nishiagina, Amagi; KAUM–
I. 82221, 33.7 mm SL, Nishiagina, Amagi; KAUM–I. 82229, 34.3 mm SL, Nishiagina, Amagi; KAUM–I.
82274, 29.2 mm SL, San, Tokunoshima; KAUM–I. 82437, 29.9 mm SL, San, Tokunoshima.
Fusigobius inframaculatus (Randall, 1994)[Jpn name: Hatatatesankakuhaze]
KAUM–I. 65887, 47.4 mm SL, Nishiagina, Amagi; KAUM–I. 65894, 23.4 mm SL, Nishiagina, Amagi;
KAUM–I. 66522, 38.5 mm SL, Setaki, Amagi.
Fusigobius neophytus (Günther, 1877)[Jpn name: Sankakuhaze]
Kamohara & Yamakawa (1965, as Gobius neophytus): 2 unregistered specimens, 52–53 mm TL, Matsubara,
Amagi. Akihito (1969): 1 unregistered specimen, 50 mm SL, Kametsu, Tokunoshima. This study: KAUM–I.
105573, 32.8 mm SL, Yonama, Amagi.
Fusigobius sp.
[Jpn name: N/A]
KAUM–I. 82168, 24.4 mm SL, Nishiagina, Amagi.
Gnatholepis anjerensis (Bleeker, 1851)[Jpn name: Omonhaze]
Kamohara & Yamakawa (1965, as Gobius knighti): 1 unregistered specimen, 39 mm TL, Matsubara, Amagi.
This study: KAUM–I. 105594, 30.2 mm SL, Omonawa, Isen.
Gnatholepis cauerensis cauerensis (Bleeker, 1853)[Jpn name: Kataboshiomonhaze]
Hagiwara (2018b): KAUM–I. 82207, 33.2 mm SL, Nishiagina, Amagi. This study: KAUM–I. 2059, 45.5mm SL, Tokunoshima island; KAUM–I. 14296, 43.8 mm SL, Tokunoshima island; KAUM–I. 14297, 36.1mm SL, Tokunoshima island; KAUM–I. 14298, 35.7 mm SL, Tokunoshima island; KAUM–I. 14299, 35.2mm SL, Tokunoshima island; KAUM–I. 82133, 26.6 mm SL, Nishiagina, Amagi; KAUM–I. 82207, 33.2mm SL, Nishiagina, Amagi; KAUM–I. 82235, 17.3 mm SL, Nishiagina, Amagi; KAUM–I. 82397, 38.4 mm
SL, San, Tokunoshima; KAUM–I. 82406, 33.0 mm SL, San, Tokunoshima; KAUM–I. 82521, 28.2 mm SL,
San, Tokunoshima.

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Gobiodon quinquestrigatus (Valenciennes, 1837)[Jpn name: Futairosangohaze]
Hagiwara (2018b): KAUM–I. 82358, 25.6 mm SL, San, Tokunoshima. This study: KAUM–I. 82358, 25.6mm SL, San, Tokunoshima.
Gobiodon rivulatus (Rüppell, 1830)[Jpn name: Tasujikobanhaze]
KAUM–I. 66547, 27.4 mm SL, Setaki, Amagi; KAUM–I. 66548, 28.1 mm SL, Setaki, Amagi.
Gobiopsis arenaria (Snyder, 1908)[Jpn name: Isagohaze]
KAUM–I. 2108, 24.2 mm SL, Tokunoshima island; KAUM–I. 14459, 32.0 mm SL, Tokunoshima island;
KAUM–I. 14460, 27.5 mm SL, Tokunoshima island; KAUM–I. 14461, 29.0 mm SL, Tokunoshima island.
Istigobius decoratus (Herre, 1927)[Jpn name: Hoshikazarihaze]
Hagiwara (2018b): KAUM–I. 82484, 69.5 mm SL, Kametoku, Tokunoshima. This study: KAUM–I. 82153,
26.6 mm SL, Nishiagina, Amagi; KAUM–I. 82484, 69.5 mm SL, Kametoku, Tokunoshima.
Istigobius goldmanni (Bleeker, 1852)[Jpn name: Himekazarihaze]
KAUM–I. 105574, 33.6 mm SL, Omonawa, Isen.
Istigobius nigroocellatus (Günther, 1873)[Jpn name: Okikazarihaze]
KAUM–I. 82233, 25.2 mm SL, Nishiagina, Amagi.
Kelloggella cardinalis Jordan & Seale, 1906
[Jpn name: Akahirehadakahaze]
Hagiwara (2018b): KAUM–I. 82544, 22.2 mm SL, Isen, Isen. This study: KAUM–I. 82535, 10.0 mm SL,
Isen, Isen; KAUM–I. 82536, 10.0 mm SL, Isen, Isen; KAUM–I. 82544, 22.2 mm SL, Isen, Isen; KAUM–I.
82545, 19.4 mm SL, Isen, Isen; KAUM–I. 83704, 18.0 mm SL, Isen, Isen; KAUM–I. 93862, 14.0 mm SL,
Isen, Isen; KAUM–I. 93863, 17.3 mm SL, Isen, Isen; KAUM–I. 102167, 22.0 mm SL, Isen, Isen; KAUM–
I. 102168, 20.0 mm SL, Isen, Isen; KAUM–I. 102169, 19.0 mm SL, Isen, Isen; KAUM–I. 102363, 26.5 mm
SL, Isen, Isen; KAUM–I. 102364, 25.4 mm SL, Isen, Isen; KAUM–I. 102378, 22.2 mm SL, Isen, Isen.
Mugilogobius sp.
[Jpn name: Izumihaze]
Ike et al. (1990): 2 unregistered specimens (not retained), Minato River.
Oligolepis acutipennis (Valenciennes, 1837)[Jpn name: Noborihaze]
Ike et al. (1990): 5 unregistered specimens (not retained), Ose River; 1 unregistered specimen (not retained),
Minato River.
Oligolepis stomias (Smith, 1941)[Jpn name: Kuchisakehaze]
Ike et al. (1990): 5 unregistered specimens (not retained), Ose River.

Marine and freshwater fishes of Tokunoshima island

57

Oxyurichthys lonchotus (Jenkins, 1903)[Jpn name: Minamisaruhaze]
Ike et al. (1990, as Oxyurichthys visayanus): 8 unregistered specimens (not retained), Ose River; 7
unregistered specimens (not retained), Minato River; 1 unregistered specimen (not retained), Wan-ya River.
Oxyurichthys ophthalmonema (Bleeker, 1856)[Jpn name: Matsugehaze]
Ike et al. (1990): 1 unregistered specimen (not retained), Ose River. Suzuki & Shibukawa (2004):
Tokunoshima island.
Priolepis cincta (Regan, 1908)[Jpn name: Benkeihaze]
KAUM–I. 82152, 19.5 mm SL, Nishiagina, Amagi; KAUM–I. 82209, 15.6 mm SL, Nishiagina, Amagi.
Priolepis latifascima Winterbottom & Burridge, 1993
[Jpn name: Futosujiirezumihaze]
Winterbottom & Burridge (1993): LICPP 1964055, paratype of P. latifascima, 15.4 mm SL, “China,
Tokunoshima” (maybe “China, Okinoerabujima”). Suzuki & Shibukawa (2004): Tokunoshima island. This
study: KAUM–I. 105640, 15.8 mm SL, Omonawa, Isen.
Priolepis semidoliata (Valenciennes, 1837)[Jpn name: Irezumihaze]
KAUM–I. 65858, 10.8 mm SL, Nishiagina, Amagi; KAUM–I. 82453, 18.3 mm SL, San, Tokunoshima.
Priolepis sp.
[Jpn name: N/A]
KAUM–I. 65888, 12.5 mm SL, Nishiagina, Amagi.
Psammogobius biocellatus (Valenciennes, 1837)[Jpn name: Hitomihaze]
Ike et al. (1990, as Glossogobius biocellatus): 2 unregistered specimens (not retained), Akirigami River.
Redigobius bikolanus (Herre, 1927)[Jpn name: Hinahaze]
Akihito & Meguro (1975a): LICPP 1967036, 21 mm SL, mouth of Boma River.
Rhinogobius brunneus (Temminck & Schlegel, 1845)[Jpn name: Kuroyoshinobori]Mizuoka (1967): 140 unregistered specimens, Sado River (currently called as Nada River). Ike et al. (1990,
as Rhinogobius sp. DA): 8 unregistered specimens (not retained), Kametoku River. This study: KAUM–I.
82311, 35.8 mm SL, Kametoku, Tokunoshima; KAUM–I. 82313, 36.1 mm SL, Kametoku, Tokunoshima;
KAUM–I. 85357, 42.1 mm SL, Akirigami River; KAUM–I. 85358, 40.0 mm SL, Akirigami River; KAUM–
I. 85359, 33.3 mm SL, Akirigami River; KAUM–I. 106538, 33.5 mm SL, Kametoku River; KAUM–I.
106540, 37.3 mm SL, Kametoku River; KAUM–I. 106541, 37.3 mm SL, Takou River.
Rhinogobius nagoyae Jordan & Seale, 1906
[Jpn name: Shimayoshinobori]Mizuoka (1967): 160 unregistered specimens, Ino River (currently called as Takara River). Ike et al.

58

I. Mochida and H. Motomura(1990, as Rhinogobius sp. CB): 2 unregistered specimens (not retained), Akirigami River. Sawashi (1995):
specimens not retained, Akirigami River. This study: KAUM–I. 105629, 34.6 mm SL, Ose River; KAUM–
I. 105633, 68.1 mm SL, Akirigami River; KAUM–I. 105634, 49.9 mm SL, Akirigami River; KAUM–I.
105646, 38.6 mm SL, Takou River; KAUM–I. 105647, 40.4 mm SL, Takou River; KAUM–I. 105654, 42.5mm SL, Takou River; KAUM–I. 106534, 34.7 mm SL, Kametoku River; KAUM–I. 106539, 34.2 mm SL,
Kametoku River; KAUM–I. 106542, 25.9 mm SL, Hon River.
Rhinogobius similis Gill, 1859
[Jpn name: Gokurakuhaze]
Akihito (1969): 17 unregistered specimens, 24–52 specimens, Shimoda River; 23 unregistered specimens,
14–40 mm SL, Kametoku River. Hagiwara (2018b): KAUM–I. 82310, 51.5 mm SL, Kametoku,
Tokunoshima. This study: KAUM–I. 82310, 51.5 mm SL, Kametoku, Tokunoshima; KAUM–I. 82336, 53.1mm SL, Kametoku River; KAUM–I. 82337, 55.4 mm SL, Kametoku River; KAUM–I. 82338, 53.1 mm
SL, Kametoku River; KAUM–I. 93550, 103.5 mm SL, Takou River; KAUM–I. 93551, 79.4 mm SL, Takou
River; KAUM–I. 105619, 67.9 mm SL, Wan-ya River; KAUM–I. 105620, 57.5 mm SL, Wan-ya River;
KAUM–I. 105621, 50.2 mm SL, Wan-ya River; KAUM–I. 105622, 50.2 mm SL, Wan-ya River; KAUM–
I. 105623, 46.7 mm SL, Wan-ya River; KAUM–I. 105624, 45.4 mm SL, Wan-ya River; KAUM–I. 105625,
38.5 mm SL, Wan-ya River; KAUM–I. 105626, 42.2 mm SL, Wan-ya River.
Rhinogobius sp. MO[Jpn name: Ayayoshinobori]Suzuki & Shibukawa (2004): Tokunoshima island. Akihito et al. (2013): Tokunoshima island. This study:
KAUM–I. 105642, 26.8 mm SL, Takou River; KAUM–I. 106533, 43.2 mm SL, Takou River; KAUM–I.
106535, 33.0 mm SL, Shikaura River.
Rhinogobius sp. YB[Jpn name: Kibarayoshinobori]Ike et al. (1990): 6 unregistered specimens (not retained), Kametoku River. Kon & Yoshino (2003):
URM-P 31804, 31805, 41425, 41427, 9 specimens, Uenari River; URM-P 27660, 27661, 17 specimens,
Akirigami River; URM-P 27665, 27668, 27669, 17 specimens, Agon River. Suzuki & Shibukawa (2004):
Tokunoshima island. Ohara et al. (2008): 35 unregistered specimens, Akirigami River. Tachihara (2009):
Tokunoshima island. This study: KAUM–I. 106532, 42.8 mm SL, Takou River; KAUM–I. 106536, 34.7 mm
SL, Kametoku River; KAUM–I. 106537, 29.7 mm SL, Kametoku River; KAUM–I. 118204, 42.6 mm SL,
Akirigami River.
Sicyopterus japonicus (Tanaka, 1909)[Jpn name: Bozuhaze]
Akihito & Meguro (1979): 1 unregistered specimen, 86 mm SL, Shimoda River. Ike et al. (1990): 1
unregistered specimen (not retained), Akirigami River. Sawashi (1995): specimens not retained, Akirigami
River. This study: KAUM–I. 102156, 85.6 mm SL, Akirigami River; KAUM–I. 102223, 54.4 mm SL,
Shikaura River.
Stenogobius sp.
[Jpn name: Tanekawahaze]
KAUM–I. 106565, 18.3 mm SL, Akirigami River.
Tridentiger kuroiwae Jordan & Tanaka, 1927
[Jpn name: Naganogori]Marine and freshwater fishes of Tokunoshima island

59

Ike et al. (1990): 1 unregistered specimen (not retained), Minato River. Sawashi (1995): specimens notretained, Akirigami River. Akihito et al. (2013): Tokunoshima island. Hagiwara (2018b): KAUM–I. 82334,
78.8 mm SL, Kametoku River. This study: KAUM–I. 82334, 78.8 mm SL, Kametoku River; KAUM–I.
82335, 48.1 mm SL, Kametoku River; KAUM–I. 105643, 34.7 mm SL, Takou River; KAUM–I. 105644,
34.6 mm SL, Takou River.
Trimma caesiura Jordan & Seale, 1906
[Jpn name: Benihaze]
Hagiwara (2018b): KAUM–I. 82208, 23.0 mm SL, Nishiagina, Amagi. This study: KAUM–I. 66544, 22.7mm SL, Setaki, Amagi; KAUM–I. 82208, 23.0 mm SL, Nishiagina, Amagi.
Trimma caudomaculatum Yoshino & Araga, 1975
[Jpn name: Aogihaze]
KAUM–I. 66546, 19.1 mm SL, Setaki, Amagi.
Trimma maiandros Hoese, Winterbottom & Reader, 2011
[Jpn name: Aobenihaze]
Hagiwara (2018b): KAUM–I. 66558, 19.9 mm SL, Setaki, Amagi. This study: KAUM–I. 65857, 11.9 mm
SL, Nishiagina, Amagi; KAUM–I. 66558, 19.9 mm SL, Setaki, Amagi.
Trimma naudei Smith, 1957
[Jpn name: Chigobenihaze]
KAUM–I. 66520, 8.9 mm SL, Setaki, Amagi; KAUM–I. 66521, 11.1 mm SL, Setaki, Amagi; KAUM–I.
66543, 13.6 mm SL, Setaki, Amagi.
Trimma okinawae (Aoyagi, 1949)[Jpn name: Okinawabenihaze]
Hagiwara (2018b): KAUM–I. 82289, 22.5 mm SL, San, Tokunoshima. This study: KAUM–I. 65861, 12.9mm SL, Nishiagina, Amagi; KAUM–I. 65880, 13.5 mm SL, Nishiagina, Amagi; KAUM–I. 65885, 11.9 mm
SL, Nishiagina, Amagi; KAUM–I. 65907, 15.0 mm SL, Nishiagina, Amagi; KAUM–I. 82150, 13.1 mm SL,
Nishiagina, Amagi; KAUM–I. 82160, 16.9 mm SL, Nishiagina, Amagi; KAUM–I. 82231, 19.0 mm SL,
Nishiagina, Amagi; KAUM–I. 82259, 23.6 mm SL, San, Tokunoshima; KAUM–I. 82262, 16.5 mm SL, San,
Tokunoshima; KAUM–I. 82289, 22.5 mm SL, San, Tokunoshima.
Trimma sp.
[Jpn name: N/A]
KAUM–I. 82290, 22.5 mm SL, San, Tokunoshima.
Trimmatom sp.
[Jpn name: Shimaisohaze]
Suzuki & Shibukawa (2004, as Trimmatom sp. A): Tokunoshima island. Akihito et al. (2013): Tokunoshimaisland. This study: KAUM–I. 66516, 13.1 mm SL, Setaki, Amagi; KAUM–I. 66559, 8.1 mm SL, Setaki,
Amagi.
Trypauchenopsis intermedia Volz, 1903
[Jpn name: Higewarasubo]
Ike et al. (1990, as Taenioides limicola): 1 unregistered specimen (not retained), Ose River; 16 unregisteredspecimens (not retained), Wan-ya River.

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Valenciennea helsdingenii (Bleeker, 1858)[Jpn name: Kuroitohaze]
KAUM–I. 65902, 26.6 mm SL, Nishiagina, Amagi.
Valenciennea longipinnis (Lay & Bennett, 1839)[Jpn name: Sazanamihaze]
KAUM–I. 105571, 64.8 mm SL, Asan, Isen; KAUM–I. 105572, 70.4 mm SL, Asan, Isen.
Valenciennea puellaris (Tomiyama, 1956)[Jpn name: Otomehaze]
KPM-NR 13120, underwater photograph, Tokunoshima.
Valenciennea strigata (Broussonet, 1782)[Jpn name: Akahachihaze]
Hagiwara (2018b): KAUM–I. 82222, 69.9 mm SL, Nishiagina, Amagi. This study: KAUM–I. 66514, 51.5mm SL, Yonama, Amagi; KAUM–I. 66515, 52.1 mm SL, Yonama, Amagi; KAUM–I. 82222, 69.9 mm SL,
Nishiagina, Amagi; KAUM–I. 88421, 123.5 mm SL, Omonawa, Isen.
Vanderhorstia sp.
[Jpn name: Komon-yatsushihaze]
KPM-NR 16180, underwater photograph, Tokunoshima.
Yongeichthys nebulosus (Forsskål, 1775)[Jpn name: Tsumugihaze]
KAUM–I. 102026, 69.8 mm SL, Matsubara, Amagi.
FAMILY KRAEMERIIDAE
Kraemeria cunicularia Rofen, 1958
[Jpn name: Ryukyunaminokohaze]
Ike et al. (1990): 1 unregistered specimen (not retained), Minato River. Akihito et al. (2013): Tokunoshimaisland. Hagiwara (2018b): KAUM–I. 83891, 27.9 mm SL, Omonawa, Isen. This study: KAUM–I. 83891,
27.9 mm SL, Omonawa, Isen; KAUM–I. 98393, 26.2 mm SL, Omonawa, Isen; KAUM–I. 102328, 29.5 mm
SL, Omonawa, Isen; KAUM–I. 102329, 28.7 mm SL, Omonawa, Isen; KAUM–I. 102330, 28.3 mm SL,
Omonawa, Isen; KAUM–I. 102331, 27.6 mm SL, Omonawa, Isen.
FAMILY PTERELEOTRIDAE
Ptereleotris evides (Jordan & Hubbs, 1925)[Jpn name: Kuroyurihaze]
Hagiwara (2018c): KAUM–I. 82482, 68.4 mm SL, Kametoku, Tokunoshima. This study: KAUM–I. 82482,
68.4 mm SL, Kametoku, Tokunoshima.
Parioglossus raoi (Herre, 1939)[Jpn name: Miyarabihaze]
KAUM–I. 105536, 18.8 mm SL, Isen, Isen. KAUM–I. 105537, 19.8 mm SL, Isen, Isen.
FAMILY EPHIPPIDAE
Platax orbicularis (Forsskål, 1775)[Jpn name: Nan-yotsubameuo]

Marine and freshwater fishes of Tokunoshima island

61

Kuriiwa (2018b): KAUM–I. 82510, 77.7 mm SL, Kametoku, Tokunoshima. This study: KAUM–I. 82510,
77.7 mm SL, Kametoku, Tokunoshima; KAUM–I. 95069, 40.2 mm SL, Boma, Tokunoshima; KAUM–I.
98692, 46.5 mm SL, Omonawa, Isen; KAUM–I. 102095, 30.5 mm SL, Omonawa, Isen; KAUM–I. 102178,
33.1 mm SL, Matsubara, Amagi.
FAMILY SCATOPHAGIDAE
Scatophagus argus (Linnaeus, 1766)[Jpn name: Kurohoshimanjudai]
Kuriiwa (2018c): KAUM–I. 3312, 29.9 mm SL, Tokunoshima island. KAUM–I. 102392, 13.8 mm SL,
Boma, Tokunoshima; KAUM–I. 102393, 14.3 mm SL, Boma, Tokunoshima; KAUM–I. 105645, 80.4 mm
SL, Takou River.
FAMILY SIGANIDAE
Siganus spinus (Linnaeus, 1758)[Jpn name: Amiaigo]
KAUM–I. 2038, 99.6 mm SL, Tokunoshima island; KAUM–I. 82423, 67.5 mm SL, San, Tokunoshima;
KAUM–I. 82424, 83.6 mm SL, San, Tokunoshima; KAUM–I. 83662, 59.8 mm SL, Omonawa, Isen;
KAUM–I. 102341, 34.1 mm SL, Omonawa, Isen; KAUM–I. 102342, 32.8 mm SL, Omonawa, Isen; KAUM–
I. 102343, 33.8 mm SL, Omonawa, Isen; KAUM–I. 102344, 34.3 mm SL, Omonawa, Isen; KAUM–I.
102345, 33.5 mm SL, Omonawa, Isen; KAUM–I. 102346, 30.8 mm SL, Omonawa, Isen; KAUM–I. 102347,
34.7 mm SL, Omonawa, Isen; KAUM–I. 102348, 34.4 mm SL, Omonawa, Isen; KAUM–I. 102349, 31.1mm SL, Omonawa, Isen; KAUM–I. 102350, 32.4 mm SL, Omonawa, Isen; KAUM–I. 102351, 32.0 mm SL,
Omonawa, Isen; KAUM–I. 105559, 90 mm SL, Omonawa, Isen.
FAMILY ZANCLIDAE
Zanclus cornutus (Linnaeus, 1758)[Jpn name: Tsunodashi]
KAUM–I. 65932, 67.6 mm SL, Setaki, Amagi; KAUM–I. 82487, 72.9 mm SL, Kametoku, Tokunoshima;
KAUM–I. 95057, 76.0 mm SL, Omonawa, Isen; KAUM–I. 102056, 66.8 mm SL, Omonawa, Isen.
FAMILY ACANTHURIDAE
Acanthurus dussumieri Valenciennes, 1835
[Jpn name: Nisekanranhagi]
Sakurai (2018b): KAUM–I. 82421, 193.0 mm SL, San, Tokunoshima. This study: KAUM–I. 82421, 193.0mm SL, San, Tokunoshima.
Acanthurus japonicus (Schmidt, 1931)[Jpn name: Namidakurohagi]
Sakurai (2018b): KAUM–I. 66501, 65.7 mm SL, Yonama, Amagi. This study: KAUM–I. 66501, 65.7 mm
SL, Yonama, Amagi.
Acanthurus leucopareius (Jenkins, 1903)[Jpn name: Sujikurohagi]
Sakurai (2018b): KAUM–I. 91366, 36.6 mm SL, Omonawa, Isen. This study: KAUM–I. 91365, 36.7 mm
SL, Omonawa, Isen; KAUM–I. 91366, 36.6 mm SL, Omonawa, Isen.

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Acanthurus nigrofuscus (Forsskål, 1775)[Jpn name: Naganiza]
Sakurai (2018b): KAUM–I. 82440, 120.3 mm SL, Kametoku, Tokunoshima. This study: KAUM–I. 82354,
117.3 mm SL, Kametoku, Tokunoshima; KAUM–I. 82426, 37.7 mm SL, San, Tokunoshima; KAUM–I.
82440, 120.3 mm SL, Kametoku, Tokunoshima; KAUM–I. 82531, 82.0 mm SL, Omonawa, Isen; KAUM–I.
98099, 84.1 mm SL, Omonawa, Isen.
Acanthurus olivaceus Bloch & Schneider, 1801
[Jpn name: Montsukihagi] (cover photo)
KAUM–I. 89922, 37 mm SL, Yonama, Amagi; KAUM–I. 102259, 31.5 mm SL, Asan, Isen; KAUM–I.
102260, 25.9 mm SL, Asan, Isen; KAUM–I. 102261, 25.6 mm SL, Asan, Isen; KAUM–I. 105627, 25.5 mm
SL, Omonawa, Isen; KAUM–I. 105628, 24.5 mm SL, Omonawa, Isen.
Acanthurus sp.
[Jpn name: N/A]
KAUM–I. 105578, 40.0 mm SL, Omonawa, Isen; KAUM–I. 105579, 42.6 mm SL, Omonawa, Isen; KAUM–
I. 105580, 42.7 mm SL, Omonawa, Isen; KAUM–I. 105581, 34.7 mm SL, Omonawa, Isen; KAUM–I.
105582, 41.2 mm SL, Omonawa, Isen.
Acanthurus triostegus (Linnaeus, 1758)[Jpn name: Shimahagi]
Sakurai (2018b): KAUM–I. 82193, 108.4 mm SL, Nishiagina, Amagi. This study: KAUM–I. 82193, 108.4mm SL, Nishiagina, Amagi; KAUM–I. 83467, 143.3 mm SL, Omonawa, Isen; KAUM–I. 83468, 136.3mm SL, Omonawa, Isen; KAUM–I. 88743, 51.7 mm SL, Inutabu, Isen; KAUM–I. 97474, 157.6 mm SL,
Omonawa, Isen; KAUM–I. 98287, 23.5 mm SL, Isen, Isen; KAUM–I. 102115, 24.1 mm SL, Isen, Isen;
KAUM–I. 102240, 44.4 mm SL, Omonawa, Isen; KAUM–I. 102241, 42.8 mm SL, Omonawa, Isen.
Acanthurus xanthopterus Valenciennes, 1835
[Jpn name: Kurohagi]
Sakurai (2018b): KAUM–I. 82503, 127.0 mm SL, San, Tokunoshima. This study: KAUM–I. 82503, 127.0
mm SL, San, Tokunoshima; KAUM–I. 83470, 99.0 mm SL, Matsubara, Amagi; KAUM–I. 93496, 151.6 mm
SL, San, Tokunoshima; KAUM–I. 98098, 91.0 mm SL, Omonawa, Isen.
Ctenochaetus binotatus Randall, 1955
[Jpn name: Kokutensazanamihagi]
Sakurai (2018b): KAUM–I. 82210, 31.1 mm SL, Nishiagina, Amagi. This study: KAUM–I. 82210, 31.1 mm
SL, Nishiagina, Amagi; KAUM–I. 91363, 42.8 mm SL, Omonawa, Isen; KAUM–I. 102233, 29.6 mm SL,
Omonawa, Isen; KAUM–I. 102234, 29.6 mm SL, Omonawa, Isen.
Ctenochaetus striatus (Quoy & Gaimard, 1825)[Jpn name: Sazanamihagi]
Sakurai (2018b): KAUM–I. 82441, 161.6 mm SL, Kametoku, Tokunoshima. This study: KAUM–I. 82441,
161.6 mm SL, Kametoku, Tokunoshima; KAUM–I. 89886, 145.0 mm SL, Isen, Isen; KAUM–I. 98362,
156.6 mm SL, Agon, Isen.
Naso lituratus (Forster, 1801)[Jpn name: Miyakotenguhagi]
Sakurai (2018b): KAUM–I. 82151, 70.0 mm SL, Nishiagina, Amagi. This study: KAUM–I. 82151, 70.0 mm

Marine and freshwater fishes of Tokunoshima island

63

SL, Nishiagina, Amagi; KAUM–I. 82191, 229.9 mm SL, Nishiagina, Amagi; KAUM–I. 83472, 243.0 mm
SL, Omonawa, Isen; KAUM–I. 97475, 94.0 mm SL, Omonawa, Isen.
Naso sp.
[Jpn name: N/A]
KAUM–I. 106552, 23.2 mm SL, San, Tokunoshima.
Naso unicornis (Forsskål, 1775)[Jpn name: Tenguhagi]
Sakurai (2018b): KAUM–I. 83433, 147.4 mm SL, Omonawa, Isen. This study: KAUM–I. 83433, 147.4 mm
SL, Omonawa, Isen; KAUM–I. 95063, 97.5 mm SL, Omonawa, Isen; KAUM–I. 102065, 118.7 mm SL,
Omonawa, Isen.
Zebrasoma scopas (Cuvier, 1829)[Jpn name: Gomahagi]
KAUM–I. 66565, 27.9 mm SL, Yonama, Amagi; KAUM–I. 105548, 27.3 mm SL, off Tokunoshima or
Amami-oshima islands.
Zebrasoma velifer (Bloch, 1795)[Jpn name: Hirenagahagi]
Sakurai (2018b): KAUM–I. 82432, 34.1 mm SL, San, Tokunoshima. This study: KAUM–I. 66564, 25.5 mm
SL, Yonama, Amagi; KAUM–I. 82432, 34.1 mm SL, San, Tokunoshima.
FAMILY SPHYRAENIDAE
Sphyraena barracuda (Edwards, 1771)[Jpn name: Onikamasu]
KAUM–I. 105523, 30.6 mm SL, Asan, Isen.
Sphyraena obtusata Cuvier, 1829
[Jpn name: Taiwankamasu]
KAUM–I. 105604, 64.5 mm SL, Matsubara, Amagi.
FAMILY SCOMBRIDAE
Thunnus albacares (Bonnaterre, 1788)[Jpn name: Kihada]
Inaba (2018c): KAUM–I. 82915, 327.6 mm SL, west of Tokunoshima island. This study: KAUM–I. 82915,
327.6 mm SL, west of Tokunoshima island.
ORDER PLEURONECTIFORMESFAMILY BOTHIDAE
Bothus mancus (Broussonet, 1782)[Jpn name: Mondarumagarei]
KAUM–I. 65924, 239.9 mm SL, Nishiagina, Amagi.
Bothus pantherinus (Rüppell, 1830)[Jpn name: Togedarumagarei]
Muto (2018b): KAUM–I. 98360, 136.8 mm SL, Inutabu, Isen. This study: KAUM–I. 98288, 81.5 mm SL,
Omonawa, Isen; KAUM–I. 98289, 74.7 mm SL, Omonawa, Isen; KAUM–I. 98290, 66.3 mm SL, Omonawa,

64

I. Mochida and H. Motomura

Isen; KAUM–I. 98291, 41.3 mm SL, Omonawa, Isen; KAUM–I. 98292, 32.4 mm SL, Omonawa, Isen;
KAUM–I. 98293, 33.7 mm SL, Omonawa, Isen; KAUM–I. 98294, 30.5 mm SL, Omonawa, Isen; KAUM–
I. 98295, 29.6 mm SL, Omonawa, Isen; KAUM–I. 98360, 136.8 mm SL, Inutabu, Isen; KAUM–I. 105557,
78.1 mm SL, Omonawa, Isen; KAUM–I. 106557, 31.6 mm SL, Omonawa, Isen.
FAMILY SOLEIDAE
Aseraggodes cheni Randall & Senou, 2007
[Jpn name: Mushikuisasaushinoshita]
Muto (2018c): KAUM–I. 82224, 25.7 mm SL, Nishiagina, Amagi. This study: KAUM–I. 82224, 25.7 mm
SL, Nishiagina, Amagi.
Aseraggodes sp.
[Jpn name: N/A]
KAUM–I. 102295, 49.9 mm SL, Omonawa, Isen.
Pardachirus pavoninus (Lacepède, 1802)[Jpn name: Minamiushinoshita]
Muto (2018c): KAUM–I. 84262, 65.5 mm SL, Omonawa, Isen. This study: KAUM–I. 84262, 65.5 mm SL,
Omonawa, Isen; KAUM–I. 84263, 57.8 mm SL, Omonawa, Isen; KAUM–I. 84264, 31.5 mm SL, Omonawa,
Isen; KAUM–I. 89358, 39.1 mm SL, Omonawa, Isen; KAUM–I. 98296, 107.5 mm SL, Omonawa, Isen;
KAUM–I. 98297, 113.3 mm SL, Omonawa, Isen; KAUM–I. 98298, 128.4 mm SL, Omonawa, Isen; KAUM–
I. 98299, 52.5 mm SL, Omonawa, Isen; KAUM–I. 98300, 53.9 mm SL, Omonawa, Isen; KAUM–I. 98301,
48.1 mm SL, Omonawa, Isen; KAUM–I. 98302, 47.1 mm SL, Omonawa, Isen; KAUM–I. 98303, 39.4 mm
SL, Omonawa, Isen. KAUM–I. 102353, 120.0 mm SL, Omonawa, Isen. KAUM–I. 102384, 41.8 mm SL,
Omonawa, Isen.
Soleichthys heterorhinos (Bleeker, 1856)[Jpn name: Sazanamiushinoshita]
Kamohara & Yamakawa (1965): 2 unregistered specimens, 83–93 mm TL, Matsubara, Amagi.
ORDER TETRAODONTIFORMESFAMILY BALISTIDAE
Balistapus undulatus (Park, 1797)[Jpn name: Kumadori]Matsuura (2018a): KAUM–I. 82460, 149.0 mm SL, Kametoku, Tokunoshima. This study: KAUM–I. 82417,
63.0 mm SL, San, Tokunoshima; KAUM–I. 82460, 149.0 mm SL, Kametoku, Tokunoshima; KAUM–I.
83443, 99.5 mm SL, Isen, Isen; KAUM–I. 89883, 133.1 mm SL, Kaneku, Amagi.
FAMILY MONACANTHIDAE
Aluterus monoceros (Linnaeus, 1758)[Jpn name: Usubahagi]
KAUM–I. 85356, 44.0 mm SL, Boma, Tokunoshima.
Aluterus scriptus (Osbeck, 1765)[Jpn name: Soushihagi]
KAUM–I. 85355, 47.8 mm SL, Boma, Tokunoshima; KAUM–I. 102267, 43.0 mm SL, Boma, Tokunoshima;
KAUM–I. 102268, 35.6 mm SL, Boma, Tokunoshima.

Marine and freshwater fishes of Tokunoshima island

65

Balistoides conspicillum (Bloch & Schneider, 1801)[Jpn name: Mongarakawahagi]
Matsuura (2018b): KAUM–I. 82420, 216.4 mm SL, San, Tokunoshima. This study: KAUM–I. 65915, 121.4mm SL, Yonama, Amagi; KAUM–I. 82420, 216.4 mm SL, San, Tokunoshima; KAUM–I. 97478, 134.6 mm
SL, Isen, Isen.
Balistoides viridescens (Bloch & Schneider, 1801)[Jpn name: Gomamongara]
KAUM–I. 95075, 59.5 mm SL, Boma, Tokunoshima; KAUM–I. 95076, 53.1 mm SL, Boma, Tokunoshima;
KAUM–I. 102049, 22.3 mm SL, San, Tokunoshima; KAUM–I. 102050, 27.0 mm SL, San, Tokunoshima;
KAUM–I. 102051, 28.0 mm SL, San, Tokunoshima; KAUM–I. 102052, 25.6 mm SL, San, Tokunoshima;
KAUM–I. 105540, 52.3 mm SL, Boma, Tokunoshima; KAUM–I. 105541, 56.6 mm SL, Boma,
Tokunoshima; KAUM–I. 105549, 32.5 mm SL, off Tokunoshima or Amami-oshima islands.
Cantherhines dumerilii (Hollard, 1854)[Jpn name: Hakuseihagi]
Matsuura (2018b): KAUM–I. 82199, 231.9 mm SL, Nishiagina, Amagi. This study: KAUM–I. 82199, 231.9mm SL, Nishiagina, Amagi.
Cantherhines pardalis (Rüppell, 1837)[Jpn name: Amimeumazurahagi]
KAUM–I. 89354, 160.0 mm SL, Inutabu, Isen.
Canthidermis maculata (Bloch, 1786)[Jpn name: Amimongara]
KAUM–I. 93501, 41.3 mm SL, Boma, Tokunoshima; KAUM–I. 102109, 39.3 mm SL, San, Tokunoshima;
KAUM–I. 102110, 28.0 mm SL, San, Tokunoshima; KAUM–I. 102111, 21.6 mm SL, San, Tokunoshima.
Melichthys vidua (Richardson, 1845)[Jpn name: Kuromongara]
Matsuura (2018b): KAUM–I. 82572, 128.1 mm SL, Isen, Isen. This study: KAUM–I. 82572, 128.1 mm SL,
Isen, Isen.
Odonus niger (Rüppell, 1836)[Jpn name: Akamongara]
KAUM–I. 82110, 39.5 mm SL, Nishiagina, Amagi.
Paraluteres prionurus (Bleeker, 1851)[Jpn name: Nokogirihagi]
Matsuura (2018b): KAUM–I. 82197, 57.5 mm SL, Nishiagina, Amagi. This study: KAUM–I. 82197, 57.5mm SL, Nishiagina, Amagi.
Pervagor janthinosoma (Bleeker, 1854)[Jpn name: Nishikikawahagi]
Matsuura (2018b): KAUM–I. 66573, 57.3 mm SL, Yonama, Amagi. This study: KAUM–I. 66573, 57.3 mm
SL, Yonama, Amagi; KAUM–I. 82433, 43.6 mm SL, San, Tokunoshima.

66

I. Mochida and H. Motomura

Pseudobalistes flavimarginatus (Rüppell, 1829)[Jpn name: Kiherimongara]
KAUM–I. 93847, 64.8 mm SL, Boma, Tokunoshima; KAUM–I. 93858, 58.3 mm SL, Boma, Tokunoshima.
Rhinecanthus aculeatus (Linnaeus, 1758)[Jpn name: Murasamemongara]
Matsuura (2018b): KAUM–I. 82395, 40.9 mm SL, San, Tokunoshima. This study: KAUM–I. 82395,
40.9 mm SL, San, Tokunoshima; KAUM–I. 91362, 24.9 mm SL, Omonawa, Isen; KAUM–I. 98365, 93.0mm SL, Inutabu, Isen; KAUM–I. 98366, 80.9 mm SL, Omonawa, Isen. KAUM–I. 98679, 70.2 mm SL,
Omonawa, Isen; KAUM–I. 102029, 27.6 mm SL, Omonawa, Isen; KAUM–I. 102084, 57.8 mm SL, Boma,
Tokunoshima; KAUM–I. 102203, 25.7 mm SL, Omonawa, Isen; KAUM–I. 102411, 22.2 mm SL, Omonawa,
Isen.
Rhinecanthus rectangulus (Bloch & Schneider, 1801)[Jpn name: Tasukimongara]
Matsuura (2018b): KAUM–I. 82364, 146.4 mm SL, Kametoku, Tokunoshima. This study: KAUM–I. 82364,
146.4 mm SL, Kametoku, Tokunoshima; KAUM–I. 82470, 101.0 mm SL, San, Tokunoshima; KAUM–
I. 82558, 123.4 mm SL, Omonawa, Isen; KAUM–I. 82570, 130.8 mm SL, Isen, Isen; KAUM–I. 82571,
102.1 mm SL, Isen, Isen; KAUM–I. 83485, 62.7 mm SL, Kaneku, Amagi; KAUM–I. 93620, 148.8 mm SL,
Kaneku, Amagi; KAUM–I. 93839, 71.0 mm SL, Agon, Isen; KAUM–I. 102030, 29.1 mm SL, Omonawa,
Isen; KAUM–I. 102085, 50.8 mm SL, Kaneku, Amagi.
Rhinecanthus verrucosus (Linnaeus, 1758)[Jpn name: Kurakakemongara]
Matsuura (2018b): KAUM–I. 82111, 70.0 mm SL, Nishiagina, Amagi. This study: KAUM–I. 82111, 70.0
mm SL, Nishiagina, Amagi; KAUM–I. 82473, 71.1 mm SL, San, Tokunoshima; KAUM–I. 102170, 26.7 mm
SL, Yonama, Amagi.
Sufflamen bursa (Bloch & Schneider, 1801)[Jpn name: Musumehagi]
KAUM–I. 65922, 94.2 mm SL, Yonama, Amagi.
Sufflamen chrysopterum (Bloch & Schneider, 1801)[Jpn name: Tsumajiromongara]
Matsuura (2018b): KAUM–I. 82252, 122.9 mm SL, San, Tokunoshima. This study: KAUM–I. 82252, 122.9mm SL, San, Tokunoshima; KAUM–I. 82315, 115.5 mm SL, San, Tokunoshima; KAUM–I. 82351, 116.5mm SL, San, Tokunoshima; KAUM–I. 82557, 112.4 mm SL, Omonawa, Isen; KAUM–I. 83442, 131.5 mm
SL, Isen, Isen; KAUM–I. 83446, 92.2 mm SL, Hetono, Amagi; KAUM–I. 83483, 127.3 mm SL, Kaneku,
Amagi; KAUM–I. 83486, 153.5 mm SL, Kaneku, Amagi; KAUM–I. 83487, 134.8 mm SL, Kaneku, Amagi;
KAUM–I. 88316, 140.0 mm SL, Isen, Isen; KAUM–I. 95072, 102.3 mm SL, Kametoku, Tokunoshima;
KAUM–I. 95073, 112.4 mm SL, Kametoku, Tokunoshima; KAUM–I. 102041, 65.3 mm SL, Agon, Isen;
KAUM–I. 102171, 34.7 mm SL, Yonama, Amagi.
Sufflamen fraenatum (Latreille, 1804)[Jpn name: Meganehagi]
KAUM–I. 93646, 130.6 mm SL, Kaneku, Amagi.

Marine and freshwater fishes of Tokunoshima island

67

FAMILY OSTRACIIDAE
Ostracion cubicum Linnaeus, 1758
[Jpn name: Minamihakofugu]
KAUM–I. 102177, 26.2 mm SL, Yonama, Amagi; KAUM–I. 102199, 12.5 mm SL, Yonama, Amagi;
KAUM–I. 102208, 18.2 mm SL, Kanami, Tokunoshima; KAUM–I. 102360, 10.1 mm SL, San,
Tokunoshima; KAUM–I. 102361, 11.0 mm SL, San, Tokunoshima.
Ostracion meleagris Shaw, 1796
[Jpn name: Kurohakofugu]
Matsuura (2018c): KAUM–I. 82189, 51.1 mm SL, Nishiagina, Amagi. This study: KAUM–I. 65917, 114.6mm SL, Yonama, Amagi; KAUM–I. 82189, 51.1 mm SL, Nishiagina, Amagi; KAUM–I. 102012, 51.7 mm
SL, Yonama, Amagi.
FAMILY TETRAODONTIDAE
Arothron hispidus (Linnaeus, 1758)[Jpn name: Sazanamifugu]
Matsuura (2018d): KAUM–I. 82574, 211.7 mm SL, San, Tokunoshima. This study: KAUM–I. 82574, 211.7mm SL, San, Tokunoshima; KAUM–I. 91095, 167.8 mm SL, Yonama, Amagi; KAUM–I. 97929, 56.2mm SL, Asan, Isen; KAUM–I. 98391, 22.3 mm SL, Boma, Tokunoshima; KAUM–I. 102399, 19.6 mm
SL, Omonawa, Isen; KAUM–I. 102400, 11.1 mm SL, San, Tokunoshima; KAUM–I. 102408, 8.6 mm SL,
Omonawa, Isen.
Arothron nigropunctatus (Bloch & Schneider, 1801)[Jpn name: Kokutenfugu]
KAUM–I. 82194, 136.3 mm SL, Nishiagina, Amagi; KAUM–I. 82373, 179.2 mm SL, San, Tokunoshima.
Canthigaster amboinensis (Bleeker, 1864)[Jpn name: Gomafukinchakufugu]
Matsuura (2018d): KAUM–I. 82543, 25.2 mm SL, Isen, Isen. This study: KAUM–I. 82543, 25.2 mm SL,
Isen, Isen; KAUM–I. 102053, 42.0 mm SL, Yonama, Amagi.
Canthigaster janthinoptera (Bleeker, 1855)[Jpn name: Shiborikinchakufugu]
Matsuura (2018d): KAUM–I. 82165, 52.4 mm SL, Nishiagina, Amagi. This study: KAUM–I. 82165, 52.4mm SL, Nishiagina, Amagi.
Canthigaster valentini (Bleeker, 1853)[Jpn name: Shimakinchakufugu]
Matsuura (2018d): KAUM–I. 82253, 68.9 mm SL, San, Tokunoshima. This study: KAUM–I. 65850, 81.5mm SL, Nishiagina, Amagi; KAUM–I. 82253, 68.9 mm SL, San, Tokunoshima; KAUM–I. 89413, 48.9 mm
SL, Yonama, Amagi; KAUM–I. 102011, 56.7 mm SL, Omonawa, Isen.
Chelonodontops patoca (Hamilton, 1822)[Jpn name: Okinawafugu]
Matsuura (2018d): KAUM–I. 66503, 43.3 mm SL, San, Tokunoshima. This study: KAUM–I. 66503, 43.3mm SL, San, Tokunoshima; KAUM–I. 66504, 38.3 mm SL, San, Tokunoshima; KAUM–I. 66505, 34.1 mm
SL, San, Tokunoshima; KAUM–I. 102149, 30.0 mm SL, Minato River; KAUM–I. 102150, 17.9 mm SL,
Minato River; KAUM–I. 102183, 41.5 mm SL, San, Tokunoshima; KAUM–I. 102184, 37.8 mm SL, San,

68

I. Mochida and H. Motomura

Tokunoshima; KAUM–I. 102185, 33.5 mm SL, San, Tokunoshima; KAUM–I. 102186, 33.0 mm SL, San,
Tokunoshima; KAUM–I. 102187, 29.4 mm SL, San, Tokunoshima.
FAMILY DIODONTIDAE
Chilomycterus reticulatus (Linnaeus, 1758)[Jpn name: Ishigakifugu]
Matsuura (2018e): KAUM–I. 82413, 350.9 mm SL, Kametoku, Tokunoshima. This study: KAUM–I. 82413,
350.9 mm SL, Kametoku, Tokunoshima.
Diodon holocanthus Linnaeus, 1758
[Jpn name: Harisembon]
Matsuura (2018e): KAUM–I. 82573, 183.6 mm SL, San, Tokunoshima. This study: KAUM–I. 82573, 183.6mm SL, San, Tokunoshima; KPM-NI 38956, underwater photograph, Tokunoshima.
Diodon hystrix Linnaeus, 1758
[Jpn name: Nezumifugu]
Matsuura (2018e): KAUM–I. 98799, 254.3 mm SL, Omonawa, Isen. This study: KAUM–I. 98799, 254.3mm SL, Omonawa, Isen.
Acknowledgments
We are especially grateful to R. Suzuki (Marine Service KAMUI, Tokunoshima) for providing underwaterphotographs and his assiatance with collecting fishes in Tokunoshima island, T. Mochida and M. Mochida(Kagoshima) for their devoted support to the first author during this study, students and volunteers of
KAUM for providing identifications and their assistance with collecting fishes in the island. We greatly
appreciated comments on the manuscript by G. Yearsley (Australia). This study was supported in part by
JSPS KAKENHI Grant Numbers JP19770067, JP26241027, JP24370041, JP23580259, and JP26450265;
the JSPS Core-to-Core Program: B Asia-Africa Science Platforms; the “Biological Properties of Biodiversity
Hotspots in Japan” project of the National Museum of Nature and Science, Tsukuba, Japan; “Establishmentof Research and Education Network on Biodiversity and Its Conservation in the Satsunan Islands” projectof Kagoshima University adopted by the Ministry of Education, Culture, Sports, Science and Technology,
Japan; and the “Island Research” project of Kagoshima University.
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Marine and freshwater fishes of Tokunoshima island

Fig. 1. Maps of the Ryukyu Archipelago(upper) and the Amami Islands (lower).
The island marked in red is Tokunoshima island.

Fig. 2. Colored map of Tokunoshimaisland. Yellow, blue and green indicate
towns of Amagi, Isen and Tokunoshimarespectively. Names of areas and rivers
used in text are shown here.

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A

B

C

Fig. 3. Southernmost population of Opsariichthys platypus (Cyprinidae) in Japan collected from Akirigami
River, Tokunoshima island. A, KAUM–I. 118205, 73.5 mm SL; B, KAUM–I. 118216, 54.2 mm SL;
Akirigami River.

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A

B

C

Fig. 4. Fishes inhabiting an anchialine cave in Amagi, Tokunoshima island. A, Xenoconger fryeri(Chlopsidae), KAUM–I. 66591, 459.6 mm TL; B, Eleotris sp. 1 (Eleotridae), KAUM–I. 65913, 144.1 mm
SL; C, anchialine cave in Amagi.

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A

B

C

D

E

F

Fig. 5. Fishes recorded from Tokunoshima island on the basis of underwater photographs. A, Taeniurops
meyeni (Dasyatidae), Tete, Tokunoshima, 20 m depth; B, Mobula birostris (Mobulidae), Wan'ya, Amagi,
30 m; C, Hippocampus bargibanti (Syngnathidae), Kaneku, Amagi, 30 m; D, Hemitaurichthys polylepis(Chaetodontidae), Kaneku, Amagi, 8 m; E, Evistias acutirostris (Pentacerotidae), Kaneku, Amagi, 32 m; F,
Paracirrhites arcatus (Cirrhitidae), Kaneku, Amagi, 10 m. Photos by R. Suzuki.

Ichthyological Publications by the Kagoshima University Museum2009 (March)
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2010 (March)
Motomura, H. and K. Matsuura (eds.).
Fishes of Yaku-shima Island - A World
Heritage island in the Osumi Group,
Kagoshima Prefecture, southern Japan.
National Museum of Nature and Science,
Tokyo. viii + 264 pp., 704 figs.

2011 (Nobember)
Matsunuma, M., H. Motomura, K.
Matsuura, N. A. M. Shazili and M. A.
Ambak (eds.). Fishes of Terengganu east coast of Malay Peninsula, Malaysia.
National Museum of Nature and Science,
Tokyo, Universiti Malaysia Terengganu,
Terengganu, and Kagoshima University
Museum, Kagoshima. viii + 251 pp., 678
figs.

2013 (February)
Motomura, H. and S. Ishikawa (eds.). Fishcollection building and procedures manual.
English edition. Kagoshima University
Museum, Kagoshima and Research Institutefor Humanity and Nature, Kyoto. 70 pp.

2013 (March)
Motomura, H., S. Dewa, K. Furuta and K.
Matsuura (eds.). Fishes of Iou-jima andTake-shima islands, Mishima, Kagoshima,
Japan. Kagoshima University Museum,Kagoshima and National Museum of Natureand Science, Tsukuba. 390 pp., 883 figs.

2013 (September)
Yoshida, T., H. Motomura, P. Musikasinthorn and K . Matsuura (eds.). Fishesof northern Gulf of Thailand. National
Museum of Nature and Science, Tsukuba,
Research Institute for Humanity and Nature,
Kyoto, and Kagoshima University Museum,Kagoshima. viii + 239 pp., 621 figs.

2014 (March)
Motomura, H. and K. Matsuura (eds.). Field
guide to fishes of Yoron Island in the middleof the Ryukyu Islands, Japan. Kagoshima
University Museum, Kagoshima andNational Museum of Nature and Science,
Tsukuba. 648 pp., 1808 figs.

2016 (July)
Iwatsubo, H., S. Kato and H. Motomura(eds.). Field guide to fishes of Ei insouthern Kyushu, Japan. Kagoshima
Museum of Aquatic Biodiversity, Makurazaki, Kagoshima University Museum,Kagoshima, and Seahorse Ways, Minamikyushu. 80 pp., 192 figs.

Most publications listed above can be downloaded fromhttp://www.museum.kagoshima-u.ac.jp/staff/motomura/dl.html

Ichthyological Publications by the Kagoshima University Museum2016 (August)Matsunuma, M., Y. Fukui and H. Motomura(eds.). Freshwater fishes of Kagoshima
City, southern Kyushu, Japan. Kagoshima
University Museum, Kagoshima. 86 pp.,
221 figs.

2016 (September)
Koeda, K., Y. Hibino, T. Yoshida, Y.
Kimura, R. Miki, T. Kunishima, D. Sasaki,
T. Furukawa, M. Sakurai, K. Eguchi, H.
Suzuki, T. Inaba, T. Uejo, S. Tanaka, M.
Fujisawa, H. Wada and T. Uchiyama.
Annotated checklist of fishes of Yonagunijima island, the westernmost island inJapan. Kagoshima University Museum,Kagoshima. v + 120 pp., 523 figs.

2017 (January)
Motomura, H., U. B. Alama, N. Muto, R.
Babaran, and S. Ishikawa (eds.). Commercial and bycatch market fishes of Panay
Island, Republic of the Philippines.
The Kagoshima University Museum,Kagoshima, University of the Philippines
Visayas, Iloilo, and Research Institute for
Humanity and Nature, Kyoto. 246 pp., 911
figs.

2017 (February)
Motomura, H. and S. Harazaki. Annotatedchecklist of marine and freshwater fishesof Yaku-shima island in the Osumi Islands,
Kagoshima, southern Japan, with 129
new records. Bulletin of the Kagoshima
University Museum, 9: 1-183.

2017 (October)
Iwatsubo, H. and H. Motomura (eds.).
Field guide to fishes of Kagoshima Bay
in southern Kyushu, Japan. Kagoshima
Museum of Aquatic Biodiversity, Kagoshima and the Kagoshima University
Museum, Kagoshima. 302 pp., 618 figs.

2017 (December)
K i m u r a , Y. , Y. H i b i n o , R . M i k i , T.
Minetoma and K. Koeda (eds.) Field guideto fishes of Kuchinoerabu-jima island in
the Osumi Group, Kagoshima, southern
Japan. Kagoshima University Museum,Kagoshima. 200 pp.

2018 (March)
Koeda, K., H. Hata, M. Yamada and
H. Motomura (eds.). Field guide tofishes landed at Uchinoura Fishing Port,
Kagoshima, Japan. Kagoshima University
Museum, Kagoshima. 520 pp., 1059 figs.

2014 (April)Motomura, H., K. Hagiwara, H. Senou
and M. Nakae (eds.). Identification
guide to fishes of the Amami Islands,
Japan. Kagoshima University Museum,Kagoshima, Yokosuka City Museum,
Yokosuka,Kanagawa Prefectural Museumof Natural History, Odawara, and National
Museumof Nature and Science, Tsukuba.
414 pp., 3290 figs.

Most publications listed above can be downloaded fromhttp://www.museum.kagoshima-u.ac.jp/staff/motomura/dl_en.html

ErrataThis page was added on 6 November 2018 after this article was officially published.
The following 12 species of the family Balistidae have mistakenly been placed in the family Monacanthidae in the checklist: Balistoides conspicillum (Bloch & Schneider, 1801), Balistoides viridescens(Bloch & Schneider, 1801), Canthidermis maculata (Bloch, 1786), Melichthys vidua (Richardson, 1845),
Odonus niger (Rüppell, 1836), Pseudobalistes flavimarginatus (Rüppell, 1829), Rhinecanthus aculeatus(Linnaeus, 1758), Rhinecanthus rectangulus (Bloch & Schneider, 1801), Rhinecanthus verrucosus (Linnaeus,
1758), Sufflamen bursa (Bloch & Schneider, 1801), Sufflamen chrysopterum (Bloch & Schneider, 1801),
and Sufflamen fraenatum (Latreille, 1804). Total numbers of the balistid and monacanthid species occurring
off Tokunoshima island are 13 [Balistapus undulatus (Park, 1797) has correctly been placed as a member ofBalistidae] and 6 respectively.
Based on the correct number of species in Balistidae, the last paragraph of Remarks (page 3) and Table
1 (page 3) are revised as follows: Tokunoshima island is geographically located between Amami-oshima
and Yoronjima islands. The ichthyofauna of theformer is most similar to that of Yoronjima island
Table 1. Number of species in the top 10 most speciose families
among the 14 localities in Japan. The top 10 mostoccurring at Tokunoshima island.
speciose families, and their percentages of number ofFamilyNo. of species% in total
species in each family to total number of species inGobiidae75
14.9
Tokunoshima island are almost the same with those
Pomacentridae
38
7.5
Labridae37
7.3
of Yoronjima island (Table 1; Motomura, 2016: table
Blennidae
23
4.61). The major difference between Tokuno-shima andApogonidae
22
4.4
Yoronjima islands is that the family Balistidae lanked
Scorpaenidae
20
4.0
as tenth speciose family in Tokunoshima island is
Serranidae
18
3.6
not in Yoronjima island and it is replaced with the
Chaetodontidae
16
3.2
family Tripterygiidae in Yoronjima island (Motomura,
Acanthuridae
15
3.0
2016: table 1). High species diversity of Balistidae inSyngnathidae
13
2.6
Tokunoshima island might be caused by the presence
Balistidae
13
2.6
Other families
215
42.6
of well-developed rocky reefs along the coast of the
Total
505
100.0
island where the family members prefer to inhabit.

Taeniurops meyeni (Müller & Henle 1841) off Tokunoshima island (photo by R. Suzuki)

A total of 505 species, including 214 species that represent
the first reliable records from Tokunoshima island on thebasis of collected specimens and/or underwaterphotographs, are listed with citation of literature,registration numbers, sizes, localities on the island, andsome remarks.