6ot

194h.
SeptenrLcl--()'.:tober,Entrltrtologist,
Excerpt from The Canadiar-r
NEW NEARCTIC CRANE-FLIES (TIPULIDAE, DIPTERA) : PART XXVIIBY CHARLES P. ALEXANDER,
State College,Amherst, Massachusetts
Massachusetts

The preceding part under this title was published in 1945 (Can. Ent.,
77:20a-208). The speiies discussedat this time are chiefly-fro--O_t,"go1: where
they were' collected.^by *y friends, Dr. Jarnes A. Macnab and Mr. Kenneth
M. Fender, to whom I am'indebted for the privilege of retaining the tyPg T1terials. Dr. Macnab and Mr. Fender are engaged in detailed studies of thismost interesting part of the Coast Range and already have added vastly to ourprevious knowledge in man\/ grouPs of insects.
Limonia(Dieranomyia)

anteapicalis

n. sp.

Sizelarge (wing, female, over 8 mm.) ; general coloration of thoracic notum
pale, the ptuJr.trtrrrn-.with three poorly-defined. m9r9 brownish stripes; fem.ora
wings with ayellow wiitr a narrorv but conspiiuous-subterminal brown- Itg;
weak brownish yellow tinge, the prearcular and costal fields somewhat clearer
yellow; a very reitricted biolvn pattern, chiefly evident at origin 9f Rs and outerend of the otherwise pale stigmi; Sc, rnore than one-half .Rs; abdominal tergites
weakly infuscared, the poster"iorlateial angles or even the entire caudal borders
more yellowed; cerci very small and slender, hypovalvae long and straight, darkcolored.
h[ale. Length abottt 6.5 mrn.; wing 7.5 mm..
Female. Lenqth about 7.5 mnl.; rvings 8.5 mm.
Rostrum obscure yellow; palpi with first segment ,vellow, the remainder
brownish black. Antennae rvith's.ipe brownish vellow, peclicel and flagellum
brownish black; flagellar segnrents?rval, passing.into long-.oval;-terminal segment about one-fouith longJr than the p-enultimate; verticils subequal to the
Head light brow"n; anterior ,re.te* (female) slightly exceeding twicer.g-."rt.
the diameter of scaPe.
Pronotum brownish yellow. Mesonotal praescutum with the ground brownishyetto"',-rrith three poorly'definecl more brownish stripes, the rnedian line broader
ind more conspicious, the lateral pair very narrow;.lateral Praescutal borderbehind the humeri vaguely clarkbnecl; posterior sclerites of notum yellow testaceous; scutal lobes fitteitr"d with brorvn. Pleura almost uniformly .1'ellow
testaceous. Halteres weakly infuscated, base of stem yellow' Legs with .thecoxae and trochanrers light'yellow; femora yellow with a narrow but conspicuous brown subterminal iing that is a little shorter than the yellow apgx, somewhat more extensive in the male; tibiae and tarsi obscure yellow, the outertarsal segments brownish black. Wings with a weak brownish yellow -tingc,brown
the preaicplar ancl costal fielclssomewhit clearer yellow; a verv restricted
ofstigmau"tt-indicatect at origin 9f Rf and.over R", the remainder;;;r[;;;uilr
little harker than the ,$o"".,li veins brown, brighteriate-yettowish brown, only ain the flavous areas. Vehation: Sc, ending opposiie origin of Rs, Sc, far flo1
in
its tip, Sc, alone more than one-half'Rs,'{re"etif of Sc" alittle -?t9 proximal
one'rn-cu
M, about as long^ as vein {, beyond it;
position'fourth than Rr; cell lst
its own length before fbrk of M; vein 2nd A gently sinuous.
Abdominal" tergites weakly infuscated, more he-avily so. on outer--segmore yellowmenrs, the posterior la"teral angle b. .uen the entire caudal margin

156

THE CANADIAN ENTOMOLOGIST

Sept.-Oct.,1946ed; sternites and subterminal tergite light yellow. Ovipositor with cerci verysmall and slender, directed ventrad; hypovalvae mucfi longer and nearlystraight, brownish black, restricredly more blackened at bases.
Habitat. Washington, California.
Holotype.
9, Camp lVlanzanar,Inyo Co., California, altitude about
3,900-ft.,Ju_Iy,.
1945^(Jor_Ma19da);rhrotrgh Dr. John A. Cornstock. Allotype.
Moxee,Yakimaco., washington, 1,000ft., in-yakinra valrey, July 15, i94l
6,(8. .Brookman). Paratype. 9, with the allotype, May 24, lg4l (8. nrook,*?r).
I "tT gleltly indebted to Messrs.Brookman and Mayedafor the aboveand
other very desirablespecirnensof Tipulidae.
Limonia (Dicranontyia) anteapicalis is quite distinct from all orher
Nearctic species.
of -the subgenus,differing especiallyin the parrern of the legs
and wings and in the structure of the rlviposiior, as the very unequal cerci an?
hypovalvae.
Chionea macnebeana n. sp.
Belongs to the araneoidesgroup; general coloration light brown; size
large (length, female, about 8 nrm.) ; intennae l2-segmented,
ihere being
nine-shorisegmentsbey'ondthe fusion-sesment;first segnrentbeyond the fusion
cylindrical, abottt-twice as long as its diamercr; succeedingsegmentsgraduallv
longer; verticils of unusual length, those on flagcllar segmehtsiix and ieven exceeding tnree times the lcngth of the scgnlents,on the terminal one evenlonger; vestiture-of legs consistinsof unusually long black bristles; ovipositor
with cerci very long and slender,gentlv upcun'ed ro the acute tips.
Female.Length about 8 nun.; wing rudimentar),;antenna about 1.5 mm.
General coloration of entire b<1d1,and appendageslight brown.
Terminal segmentof palpus o'lindrical, about one-fourth longer than thepe_nultimate.
Antennae lZ-scgmented,relativelylong; fusion-segmentof flagelluma low trtrncatedcone; seconclsegmentshclrt-cy,lindrical,
about twice as long as its
gradually longer, the third about one-thirdlonger
diameter; strcceeding
segrnents
than the second;verticils of segnrentsof untrsual length, at near midlength-oforgan (about flagellar segmentssix and seven,including the fusion-segmenr),rhe
longer verticils exceeding three times the length of the segments; terminal segment about tlvo-thirds the penultimate, the verticils exceedingly lengthened.
Head above with strong erect or slightlv proclinate setae.
Legs with the vestitureconsistingof unusually long powerful black bristles,
stronger on the uPper surface, the sides more nearly destitute of setae.Ovipositor with cerci very long and slender,gently upcurved to rhe acute tips.
Habitat. Oregon.
Holotype. g on slide, Wilson River, Tillamook burn, northwest of Fore s t G r o v e ,a l ti tu d e 2 ,1 2 5ft., F e b rtr ar y8, 1946 ( .1.1. Macnab) ; ar ea badly bur ned
over in 1933, 1939and 1945:-tvpe captured in a snowstormon about 20 inchesof snow.
This interestingspeciesis named for the collectcor,Dr. JamesA. Macnab,haswhosent me numerous Tipulidae frorn the Coast Range. The most similar
Nearctic speciesarc Chionea albertensisAlexander and C. primithta Alexander.
The former, known from the Canadian Rockies,differs in the smaller size andthe I l-scgmentedantennae,with the relative proportions of the basal flagellar
segmentsquite different. The eastern printitiua likewise has l2-segmentedantennae, differing in the smaller size, shorter cerci, and in the longer and moredelicate vestiture of the legs. The present fly is one of the largest speciesof thegenus so far made known.
Ormosia (Rhypholophus)

fugax n. sp.

Allied to oregonica; genetal coloration of mesonotum gray, the humeraland lateral portions of the praescutumyellow; wings brownish yellow, stigma

LXXVIII

THF. CANADIAN ENTOMOLOGIST

r57

and a vague seamalong cord darker brown; male hypopygium with the gonapophysesbispinous,the outer arm stout, sirnple,the inner one much more slender;
apical arms of aedeagusshort, lessthan the outer spine of the gonapophysis.
MaIe. Length about 5.5 mm.; wing 6 mm.
Rostrum and palpi black. Antennae with scapeand pedicel brown, flagellum black; flagellar segmentssuocylindricalto truncate-fusiform;longestveriicils exceedingthe segmenti.Head gray.
Pronotum yellow above, gray on sides. Nlesonotalpraescutllmgray, thehumeral and lateral portions yellow, enciosing the dark brown pseudosutural
foveae;posteriorscleritesof notum brownish gray, the scutellum somewhatpaler
brown; mediotergite on sides restrictedly yellow, the pleurotergite chiefly ofthis color. Pleura obscureyellow, with a very sparsegrayish bloom. Halteresyellow. Legs with the coxae yellow, sparselypruinose; trochanters yellow; rcmainder of legsbrown, the femoral basesobscureyellow; tarsi passinginto black.
Wings brownish yellow, the prearcular and costalfields a trifle more brightene'J;
stigma and a vague seamalong cord darker brown; veins broln, more yellowedin the brightened areas. Venation: Cell lsf lf" relatively small, subequal to or
shorter than the distal section of vein Mr; vein 2nd I strongly sinuouson almost the outer half.
Abdomen brown, the hypopygium more yellowish brown. Male hyPoPygium with the lobes of the outer dististylenot greatly disproportionatein length,
the inner one stouter.Gonapophysisbispinous,the outer arm stout, simple, theinner one much more slender,especiallythe straight outer spine which is abotrt
one-half longer. Apical arms of aedeagusshort, less than the outer spine of thegonapophysis.
Habitat. Oregon (Yamhill Co.)
Holotype. 6, PeavineRidge, near McNlinnville, altitude 605 ft., Septl 6, September
ember 18, 1945 (Fender); Fender Station No. 3. Paratopoty'Pe.
10,1945.
Very close to Ormosia (Rhypholophus) oregonicaAlexander, differing
chiefly in slight details of the male hypopvgium,particularly the_stout_simple
outer arm of the gonapophysis.More material 6f oregonica may show that thepresentfly is a form of this species.
Ormosia (Ormosia) upsilon n. sP.
Allied to perspectabilis;
sizelarge (wing, male, usuallyover 5 mm.);-general coloration ol mesonotumbrown, the pleura yellow; antennae (male) elongate, only a little. shorter than the body; wings with the stigma blow-n,relativeiyconspicuous;vein Znd A strongly sinuous on its outer half; male hypopygiurnwith the inner dististyle narrorvedat apex, on outer margin with a stout blackspine; phallosomic structure large and conspicuous,Y-shaped, each arm bisPrnous.
Male. Length about 4.5-6mm.; wing 4.6-7.5mm.; antennaabout 4-5 mm.
Rostrum brownish testaceous;palpi black. Antennae (male) elongate;scapeand pedicel brown, flagellum black flagellar segmentselongate-subcylindriial, morb enlarged at base,thence gradually narrowed to the apex; verticils
basal,shorter than the segments,subequal in length to the sParseoutsPreading
pale pubescence.Head dark gray.
Pronotum gray,narrowly borderedby paler; pretergiteslight yellow. Mesrestrictonotum chiefly biown, the humeral and lateral regions of prae_scutum
Pleurayelblack.
pl_t-sandtuberculate
foveaeedly obscureyellow; pseudosutural
thewithlow, the more dorsal-scleritesweakly infuscated.Halteres yellow. Legscoxae and trochantersyellow; femora obscureyellow, the lips ,restrictedlyinfuscated; tibiae brown; tarsi passinginto black. \Aringswhitish subhyaline,t-he.pr9arcular and costal fields restrictidly more yellowed; stigma brown, relatively

r58

THE CANADIAN ENTOMOLOGIST

Sept.-Oct.,1946

conspicuous;vague seamsalong cord, Cu and Rr; veins dark brown, paler in the
more brightened fields. Nlacrotrichia of cells abundant, paler in color than theveins.Venation: Ii" a trifle longer than Rrfr; cell open by atrophy of basal
sectionof ,tf ,; nr-cuat fork of M; r'ein 2nd A stronglysinuouson outer half.
Abdomen, including hypopl,gium, dark browtr. I\{ale hypopygium withthe tergite elongate,nal'rowecloutrvardly',the apex with a deep median notch;
lateral lobes long anci conspicuous,densely provided with long scabroussetae.
Basistyleat apex produced into a lobe. Outer dististylervith its outer surface
blackened,microscopicalivscAbrous,the entire lower face paler, with scattercd
microscopicsetae.In-ner dististyle broad basally, narrowed at apex, the uppcr
margin at point of narrorving produced into a stout black spine. Outer gonapophysis appearing as a strong blackened horn, the lorver rnargin with sparsecoarseserrations.Phallosomicstructlrre very conspicuous;centl'al organ more expanded outrvardly and divided into two powerful arnrs,each of rvhich is bispinbus,the spinesacute,nearly alike, the inner or apical one more darkened.
Habitat Oregon (Yamhill Co.)
Holotype. 6, PeavineRidge, near NIcNIinnr,ille,altitude 605 ft., Sept2 3 6 , Octobcr
ember 24, 1945 (Fender); Fender Station No. 3. Paratoltotypes.
ll, 1945.
The presentfly is allied to speciessuch as Ormosia (Orntosia) lanuginosa(Doane) and O. (O.) perspectabilisAlexander,differing very conspicuouslyirr
the structure of male hypopygium, especiallythe phallasonricstlucture.
Molophilus (Molophilus)

oregonicolus n. sP.

Belongsro the gracilis group, ltubipennis subgroup; sizesmall (wing, ntalc,under 3 mrn.) ; general coloration of entire body black, the anterior pretergitcs
obscureyell<lw; antcnrraeshort, flagellar scg.mentsoval,-with long verticils anda very inconspicuouspubescence;halteresr,vithwhitened knobs; legs black; lvings
with-a strong blackish tinge, the prearcular and basal fields rlore whitened;vein 2nd I short, ending solne distance before level of rn-cu; male hypopygium
entirely blackcned;basistyleshort and compact; outer dististylestrongll' curveC,with about fivc strong teeth on the conca\/emargin; inner dististylevery narrow;
phallosornicstructrlre with severalscatteredpale set.aeon apical Portion.
M aIe . L e n g th a b o u t 2 .8 m nr .; wing 2.9 mm .
Rosrrunrand palpi black.Antennac black throughout,short; flagellar segments oval, with long -onspicuousverticils ancl a short scarcelyvisible ptrbescence. Head black, somewhatdiscoloredin the t1'pe.
Pronotum black, the restrictedantcrior pretergitesobscure1,sll6tv.Mesonotum black, more or lessdiscoioredand possibiysomewhatpruinose in frerir
specimens.Pleura unifonnly black. Halteres with stem dirty white, the knobs
cbnspicuouslyclear n'hite. Legs black. \\Iings snrall, with a strong blackish ,i"g:,the prearcular and basal costal fields more lvhitened; veins atrd macrcltrichia
pale-brown.Venation: R" lying oppositeor-immediatelybeyond r-m; petiol..nlLell M. about one-third the cell or more than three tirnes m-cu; vein Znd -4
sinuous,ending sorltedistancebcforc the lcvel of mtu.
only' gently
Abdomen, including hypopygium, black. Male hypopygium with the ba';rstyle short and compact, the apical lobe rclatively stout, about as-long as-thebasalporrion of style; mesallobe at near midlength o{ style, provided rvith about
18 blickened spinosesetae,their tips drawn out into hairlike points. Out3r
dististyle strongiy curved, sornewhatas in spicttlatus,the lower or concaveface
with about five itrong denticles.Inner dististyle much smaller, very narrow, thea-boutone-fourth the length; outer half of style with microwidth at midlength"moreappressedon outer face. Phallosomic structnre oval, thescopic denticles,
ap.i* with several scatteredpale setulae.Aedeagus relatively short, about onchllf longer than the apical lobe of basistyle.

LXXVIIITHE CANADIAN ENTOMOLOGIST

159

Habitat. Oregon (Yamhill Co.)
Holotype. 6 , PeavineRidge, near N'IcMinnville,altitude 605 ft., June 2,1945 (Fender); Fender Station No. 3.
Molophilus (Molophilus) oregonicolusis best told by its almosi uniforrn
black coloration,short antennae,black legsand conspicuouslywhitened knobs of
the halteres.Among the describeddark-c<lloredspecies,it superficially resembles
M. (M.) rostriferu.sAlexander, br:t the hypopygium is quite different.

ilt;

I,f,

llolophilus(Molophilus)

suffalcatus n. sp.

Belongs to the gracilis group, pubipennis subgroup; allied to falcatus;
generai coloration of body dark brownish gray or plumbcous; antennae (male)elongate;halteresuniformly pale yellow; wings pale yellow, conspicuouslystripedwith brown, especiallyas seamsalong veins Cu and rn-cu; male hyopygium withthe apical spineof the outer dististyleshort; inner dististylewith numerouserect
spiculeson the lower face.
Male. Length about 4.5-5mm.; wing 5.5-6rnm.; antennaabout 2.8-3mnr.
Female. Length about 5.5 mm.; wing 6 mm.
Rostrum and palpi black. Antennae (male) unusually long; segments
elongate-fusiform,each rvith a single unilaterally arranged verticil and abundaut
subequalor longer pale erect setae. Head gray.
Pronotum dark brown, the scutellum and pretergitestestaceousyellow.
Mesonotum almost unifornrly dark brolvnish gray, without pattern; pseudosutural foveaeblack; hurneralregion of praescutunrrestrictedlyyellorv.Pleura dark
plumbeousgray; dorsopleuralmembraneclark. Halteres uniformly pale yellow.
Legs with the fore coxae darkened,renraining coxae and all trochantersyellow;
fore femora chiefly blackened,the basesrestrictedly paler, involving about theproximal third or less; rniddle and hind femora obscureyellow, the tips broadly
blackened, including about the outer fourth or fifth; tibiae yellowish brown,
the tips narrowly brownish black; tarsi passing into black. Wings with theground color pale yellolv, the prearcular and costal fields more saturatedyellorv,
a diffuse brown pattern, including washesin outer radial field, a conspicuousand a strong
seamfor almost the rvhole length of vein Cu, including also n1.-c1t,
axillary darkening, including all of cell 2nd A and adjacent parts of lsf l; veinspale, darker in the patterned areas.Venatiotr: R" lying just beyond the level ofr-rn; petiole of cell tlf, nearly three times m-cn; vein Znd I long and sinuous,thecell narrow, cspeciallyon its outer portion, the vein ending somedistancebeyond
rn-cii,about oppositeone-third to one-fourth the iength of the petiole of cell M,.
Abdomen, including h-vpopygium,brownish black to black. Male hypopvgium of the general tvpe of falcatus but differing in details, especiallyof theouter dististyle.Basistvlewith the peglike spinesof mesal lobe congregatedinto
a compact group. Outer dististyle with the narrowed outer portion short, lessthan the enlarged base and usually only from one-half to two-thirds as long.
Inner dististyle much smaller, the spiculosepoints small, more numerous on thelorver face.Phallosomicplate with densepale setulaeover most of the surface.In
falcatus, the outer distist,vleis long-producedinto a slender spine that is much
longer than the enlargedbase,in casestwice this length; the,inner dististyle is ofdifferent shape,with the sprculesrestrictedto the uPPer surface.
Habitat. Oregon,.BritishColumbia.
Holotype. A, Metolius River, JeffersonCo., Oregon, June 14, 1945(Fender). Allotopotype. 9 , pinned with type. Paratypes. 6 L Peavine Ridge,
Yamhill Co., Oregon, on sword fern, May 3-16,1945 (Fender); 16, Steelhead,
British Columbia, May 28, 1933 (Hugh B. Leech).
The nearestally of the present fly is Mclophilus (It[olophilus) falcatus
Bergroth (Alaska, southward along the Rocky Mountains to Wyoming), whicu
differs in the structure of the male hypopygium, as compared above.