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NEW TIPULIDAE FROItr THE SOCIETY ISLANDS *By
Cnenr,Es P. AlrxeNDER
MessecnusHTs SrATE CoLLEGEINTRODUCTION
The material on which this paper is based was collected by A. M.
Adamson,of the Pacific EntomologicalSurvey, and Dr. Andr6 L. Tonnoir.
I wish to extend my deepestthanks to them, as well as to E. P. Mumford,
Director of the Pacific EntomologicalSurvey, for the opportunity to study
this material. The Tonnoir material is preservedin my collectionthroughthe kindnessof the collector.
TRIBE ERIOPTERINI

Five speciesof Gonomyia,,subgenusLipophlep.r,are now known fromTahiti, all endemic. These speciesmay be separatedby means of the following key:
1. Wings whitish, with a restricted brown pattern, including spots at arculus,
origin of Rs, end of anterior branch of Rs and the cord; no macrotrichia
on vein rst 4...-......-..punctlgcra
Wings either unicolorous, except for the stigmal darkening, or darkcolored with a restrictedwhitish and yellow pattern; a seriesof from thirty
to fifty macrotrichiaon vein rct A...-......
--...-..--------....-.2
2. Wings dark, with whitish discal spots and a lunate yellow apex; legs
yellow and black, the tibiae black with a broad yellow subbasalring; size
very large (wing, female,over 7 mm.)......-...
-flavldapexEdwards
Wings subhyalineor with a brownish tinge, unvariegatedexcept for thestigma when this is present; legs yellowish brown to brown, the tibiae uniform in color; size small (wing not exceeding4 mm.)...............-3
pleuramesonotumcolorationoflightbrown,
Generaltheonly
indistinctly
3.
striped with pale; cell rst Mt of. wings small, the lower face not exceeding
one-halfof veinsMr beyondit.-.-.............
.....tahltiensir
General coloration of mesonotumdark brown, the pleura distinctly striped
longitudinally with yellow; cell rsl Mz of wings retatively large, its lower
face subequalto vein Mt beyondit.................
...-...........4
light yellow........
.-tonnolrella
4. Scutellumconspicuously
Scutellumdark, concolorouswith remainderof notum............-.-...........--fuscoscutcllata

Gononyia (tipophleps) punctigera,new species(fig. r).
General coloration of mesonoturn brownish gray; pseudosutural foveae black,scutellumbrown; knobs of halteres dark brown; wings whitish with a very restricteddark-brown pattern, including clouds at arculus, origin of Rs, cord, outer end of cell
rst Mz and end of anterior branch of Rs,' vein rs, I without macrotrichia; abdomen* Pacific Entomological Survey Publication 6, article ro. Issued March 8, 1933.

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BerniceP. Bishop Museum-Bulletin 113

dark brown, the caudal margins of the segmentsconspicuouslyyellow, broadest on thetergites.
FernaleIrngth, excluding head, about 4.5 mm.; wing, 4.2 mm. Head broken. Mesonotum
brownish gray, the anterior lateral scleritesmore silvery; pseudosuturalfoveae large,blackened; scutal lobes slightly pruinose, especiallymedially; scutellum brown; postnotal mediotergite blue-gray pruinose. Pleura very dark colored, pruinose, with arestricted paler longitudinal streak, crossingthe dorsal sternopleurite. Halteres yellow,
the knobs infuscated. Legs with the coxae reddish brown, pruinose; trochantersbrownish yellow; remainderof legs broken. Wings (fig. r) whitish, including the broad wingtip, the cells before the cord somewhatmore grayish; stigma pale brown; a restricteddark-brown pattern, distributedas follows : arculus; end of Sc and origin of Rs; cord;
outer end of cell Bt Mg tip of anterior branch of Rs,' veins pale, much darker in theinfuscated areas. No macrotrichia on vein Bt A. Venation: Sc short, Sc' ending opposite the origin of Rs, Sc, at its tip; veins beyond cell rst Mt long, the distal sectionof Mr+z longer than Rs,' m-cu about two-thirds its length before the fork of. M.
AMominal tergites brown, the caudal margins of the segmentsbroadly yellow;
sternitessimilarly colored,but the pale margins narrower and less conspicuous.

Tahiti: Fautaua Valley, altitude 5o feet, r mile from sea, November
8, 1928, holotype female, Adamson.

FrcunE r. Wing of Gonomyia (Li|o|hlefs)

functigero, new species.

The closestally of the present speciesis Gonomyra (Lipophleps) ddgitifera Alexander (Fiji) which differs in the body-coloration,as the colorof the scutellum and postnotal mediotergite, and in slight details of venation, as the shorter veins issuing f rom cell lsf M r, which are here subequalin length to, or shorter than Rq whereasin punctigera the outer section ofM r*, is longer than Rs. The presentgroup of flies, with a restricteddarkened wing-pattern, may be called the cairnensisgroup, from the earliest
describedspecies,G. (L.) cairnensisAlexander (North Queensland). Oneof the most evidentcharactersfor the separationof this group from related
regional aggregationsof the subgenuslies in the entire lack of macrotrichia
on vein lst A.
Gonomyia (Lipophleps) tahitiensis' new species.
General coloration of mesonotum light brown, the scutellum only vaguely brightened; pleura very indistinctly striped with pale; knobs of halteres light yellow; wings
nearly hyaline, the veins darker.
Malelrngth, about 2.6 mm.; wing, 3.5 mm. Rostrum and palpi dark brown. Antennae
entirely dark brown. Head obscure orange, the center of the vertex slightly darkened.

Society Islonds Insects

55

Mesonotum almost uniformly light brown, the scutellum only vaguely brightened, thepostnotumslightly pruinose. Pleura brownish testaceous,very indistinctly striped longitudinally with paler. Halteres with the knobs light yellow. Legs chiefly y.tto*irtt broin,
the outer tarsal segmentsdarkened. Wings nearly hyaline, the costal region not brightened; stigma lacking; veins a little darker than the ground-color. Venation: Sc, ending
just before the origin of Rs, the latter relatively long and only gently arcuated;veins
Rr and Rr strongly divergent; cell tst Mt unusually small, the-lowir face less thanone-half of vein M,beyondit; m-cu just before the fork of.M.
Abdomen brown,. the hypopygium somewhat brightened. Male hypopygium constructed much as in G. (L.) tonnoirello,new species,but the phallosomioiodi difierently
formed, two of the elementsbeing heavily chitinized and blackenedat tips.

Tahiti: Mataiea, August, rg28, holotype male, Tonnoir.

Gonomyia(Lipophleps)tonnoirella,new species.
General coloration of mesonotum dark brown, the scutellum conspicuouslylightyellow, pleura yellow, striped longitudinally with dark brown; scapal segmentsorange;knobs of halteres light yellow; Iegs chiefly dark brown; wings bioad, dusky, the biseand costal margin more yellowish; male hypopygium with the outer dististyie a simple
gently curved rod.
MaleLength, about 2.8 mm.; wing, 3.3 mm. Rostrum and palpi dark brown. Antennae
with the scapalsegmentsorange; flagellum dark brown, the segmentswith very elongate
verticils. Head light yellow, the center of the vertex slightly darkened. Pronotum andanterior lateral pretergites light sulphur-yellow. Mlesonotal praescutum and scutum
dark brown, the lateral portions of the former and posterolateralangles of the scutal
lobes light yellow, scutellum light yellow, infuscated medially at base,postnotal mediotergite dark brown, more or less pruinose. Pleura tight yellow, striped longitudinallywith brown, the dorsal stripe more interrupted on the pteropleurite; ventral stripe including the sternopleuriteand meron. Halteres dusky, the knobs light sulphur yellow. lgs
with the coxae yellow; trochantersbrownish yellow; femora brown to light brown, thetips darkened; tibiae and tarsi dark brown. Wings relatively broad, tinged with dusky,the prearcular and costal regions more yellowish; stigma lacking; veins pale brown,
those in the yellowish areas brighter. Venation: Sc relatively short, Sc, inding justbefore the origin of Rs, Sc, apparently lackingi Rs strongly arcuated at origin; cell
rst Mz strongly widenedoutwardly, longer than vein M. beyond it; m-cu at fork of M.
Abdominal tergites dark brown, the posterior lateral anglesof the segmentslight yellow;
hypopygium chiefly yellow. Male hypopygium with the outer dististyle a simple, long,gently curved rod; inner dististyle entirely pale and fleshy, the base enlarged and conspicuouslyhairy, the outer end narrowed and terminating in a powerful fasiiculate seta.
Gonapophysesappearingas long, slender,chitinized rods that terminate in acute points.

Tahiti: Mataiea, August, 1928,holotype male, Tonnoir.
This interesting Gonomyfo,is named in honor of the collector,my friendand colleague,Dr. Andr6 L. Tonnoir.
Gonomyia(Lipophleps) fuscoscutellata,new species.
General coloration of mesonotumdark brown, including the scutellum, the surface
pruinose; basal segmentof antennalscapeobscureorange; knobs of haltereslight yellow;
wings strongly suffused with brown, the small stigma a little darker; cortal- region
narrowly light yellow; cell rsf Mz relatively large.

56

BerniceP. BishopMuseum-Bulletin 113
FemaleLeneth, about 3.8 mm.; wing, 3.6 mm. Rostrum and palpi dark brown. Antennae
with the first scapal segment obscure orange, the remainder of the organ black. Headorange, the center of the vertex restrictedly darkened. Pronotum, anterior lateral pretergites, lateral margins of praescutum and dorsopleural region light yellow I entire mesonotum dark brown, pruinose, including the scutellum; psandosutural foveae black, conspicuous; lateral margins of postnotal mediotergite obscure yellow. Pleura dark brown,with a yellow longitudinal stripe across the ventral sclerites. Halteres with the knobs
conspicuously light yellow. Legs chiefly dark brown. Wings with a strong browntinge, the small oval stigma a little darker; base and costal region narrowly lightyellow, a vague darkening in the axillary region; veins dark brown. Venation: Scshort, Sc' ending some distance before the origin of Rs, the distance being about equalta*cu;,fcr
faint, close to tip of Scr; Rs angulatedat origin; veins Rr and Ro stronglydivergent; cell rst Mt relatively large, its lower face subequal to vein Mt beyond it;
m-cu iast before the fork of. M. Abdomen blackened,the genital segment and valves ofthe ovilnsitor yellowish horn-colored.

Tahiti: Mataiea,August, 1928,holotypefemale,Tonnoir.
Styringomyia ilidyna Grimshaw.
Styringomyia didrya Grimshaw: Fauna Hawaiiensis,Diptera, p. to,'rgor ; Edwards:Ent. Soc.London,Trans.,pp. 222-223,figs.38,39, 76, r9r4.
Idiophlebio pallida Griinberg: Zool. Anzeig., 26, pp. 524-528,5 fi9s.,
1903'Tahiti: Mataiea,August, 1928,Tonnoir; FautauaValley, altitude go feet,r mile from sea,September6, L928,I male, Adamson.
Widely distributedin the Pacific islands: Hawaii, Fanning,Marquesas,Tahiti, Samoa,Tonga,Fiji, New Hebrides,the Carolinesand New Guinea.