Vol.

201

1970

December,

LXXVIII,

ofUndeseribed
HimalayaSpeciesMountains

forAbstract:
Assamin

Five

newIndia

andStyringomyia

S. melanaspis,

XIX1

01002

Publicationof eriopterine
speciesarefrom Thailand

apiculata,

theAlexanderP.

Massachusetts

Amherst,from(Dip ter a: Tipulidae),Charles

Received

Flies

CraneMarch

crane

from

flies

thesedescribed,

1970

19,

Uttar

Pradesh,Lipsothrix
beingand S. tenuispina.

S. nigrobarbata,andSikkim,
orthrotenes,in the Journal of the Newof this series of papers was published1969. At this time I am describing
York Entomological199-203,
Society, 77:
five further new species belonging to the genera Lipsothrix Loew and Styringo?
Part XVIII

and Thailand
Sikkim, Assam)
myia Loew, all from India (Uttar Pradesh,
where they were collected by Dr. Fernand Schmid and the late Deed C. Thurman.
I express my sincere thanks to the collectors for their interest in theseflies.
Lipsothrix
Generalof

coloration
paleextensively

brown;

yellow,asthe basistyle,

bases

wing

7.5 mm;antennabrown,

stigmalacking,nearlystraight,

virtually

as

male:

Lengthabout

7.5 mm;Rostrum
scapeFront

yellow,light
Pronotal

longyellow,
scutumremainderof headcentralandwithyellow,

clear
pleurotergite
light
all coxae
andtrochanters
oftipsfemoranarrowly

darkerslightly

PleuraLegs

notexpanded.

about

3 mm.

Antennae
long, as shown
relatively
by the measurements;
yellow,
palpi black.
withremainder
dark brown;dense white
setulae.
segments
flagellar
elongate,

scutellum
light brown,sides and the humeral
light fulvous,
borderregion of scutum,
posteriorscutumn.sp.

orthotenes,of notumsclerites
praescutum
light fulvous,
posteriorfemoralvery paletips very narrowly
pale brown;legs yellow,
wings
arms of aedeagus
with
male
very
long and slender,
hypopygium
thoraxyellow.
clear

andregion yellow;
of scutellum,Halt?res
lightbrown,

paleyellow.

pretergitesofMesonotal
light yellow.
of notumremainderpale
and the parascutellalightwithyellow,
tibiaeprae?
brown,yellow.
pale, knob
light brown.
of legs moreremainder
obscurenarrow.
still more
very
Wings
stem

costal border very slightly
veins pale brown.
pale brown,
stigma not evident;
darker,Longi?
tudinal veins beyondof Rs with macrotrichia,
level of midlength
also one or two
includingon each Anal vein. Venation:
as long; vein
R2 less than one-half
RU2 and R2+a subequal,Ri weakly
beforedecurved,
terminating
wing
tip.
Abdominal
decurvata

subterminal
sternites
darkerbrown
segments
tergiteslight brown,
yellow,a weakas inbrownishMaleyellow.
generally
ring, hypopygium
hypopygium
and malla,
in the long slender arms of the aedeagus
that arediffering
evidently

virtually
as long

straight
throughout,as the basistyleandtoform

1NewContribution
Yorkfrom

Entomologicalthenotlonger

or withcurved
stronglyexpanded
than their basal enlargement.

EntomologicalSociety,

Laboratory,

LXXVIII:

University201-205.

December,bases;

armsof Massachusetts.
1970.

virtually

202New

holotype:$, Hkayam(Schmid).
The most

similar

Bourn, Manipur,Assam,

York

Entomological

8,500

feet, JuneSociety

23,

1960

decurvata Alexander
and L. malla
species are Lipsothrixindifferdetails
oftheof venation, colorationbody and legs,
Alexander, whichand especiallyin hypopygialstructure, including the shorter curved arms oftheaedeagus.

Styringomyia
ofcoloration

Generalfemoraldarkenedfaintlyrings
wash
in basal halfa conspicuous

apiculata,n.sp.

dorsumpatterned

yellow,

theindicated;of cell M,

wings
yellow,vein 2nd A withthoraxwithbrown;legs yellow,
withbrown,
includingrecurved; abdominal
slightly

patterned

apex
more
sternites
and hypopygium
malewithtergites
light brown,yellowed;
hypopygium
a single seta; dististyleouter arm long and slender, unarmed,
lobe of basistylewithbearinga small acute point.
intermediate
blade withcombs of blackenedpegs, inner arm at base with

male:

about

Lengthfemale:8-8.5

Lengthabout

Rostrum

darkbrown,

andpedicelbrown,

scapebrownishwing

5.3-5.6

6.5 mm;

wing

6 mm.

flagellar

obscuresparselypruinose,narrower,

theirsegments

segmentsmoreyellowed,brown.

Mesonotal

yellow,

sides

dark

patterned

withbrown,

anterior

on sides, medianremainder
Halt?res

ends

area

obscureoflatter

dark

with

stem

yellow,rings,

yellow,prearcularthescutalpostnotum

knob

paler.
theslightly
particularly

Antennae

withincisures.

Headyellow,setae

blackatintermediateyellow,brownish

black
the

scutellum,

Pleurayellow,clearer

ventrally.

coxae

andtrochanters

yellow;

the more

basal

one

outer

posteriorly

withrings,outer
end

ends.
erect

costal

Venation:
toend

the

singlearm

slender,lobules,

beak,
none

abundant

holotype:

long

basal

short

A with2nd

veinsessile;sternites moreyellowed,
hypopygium
brown,
a singleouterwithlobe of basistylesubequal
thickened,
long and slender, outer third slightly morearm

pointedcellbrownish

light

outer

inner

narrowlybrownishsegmentouterterminal

farthersetae unusually

pointed.

Ninthyellowsetae.

intermediateseta;

at basefew but

tergiteNinthblades

long.

semioval,sterniteeach

a rounded

into

produced

Phallosome
withnarrow,anwith

withplateincludingapicaltwoovalnearlyNortheast
Frontier Agency,$, Bhairabkunda,
1961
700 feet, March
9, March(Schmid). Allotopotype,
7,March1961.
topotype, $, 700-1,000
8,feet,yellow.

seta.

stout

with

style withusual

2nd M2tibia

beneath;fieldsR& suberect,

tergites

hypopygium

pegs;

terminal

midbroader,
with
two narrow

recurved.

slightlyAbdominal
Maleandapex,yellow,

onto

pair

posteriorly

black.
Wingsbrown
and outer medial
spots at r-m, m-cu
clearer;
a conspicuous
in basalfork, veins Rz and outer half of 2nd A darkened;
paler brown wash
half of cell M;
at
those
cord
veins
darker,beyondslightly
especiallyremainingyellow,including

tarsi

obscurebroader,

continued

Legs

brown

vague

praescutum

stripesbrown.

paler

brown.

light
two

lobes

the coloryellow,brown,

femora withyellow,
a concentration
withofbrown

sublateralconvergent;remainder
femur

of outerbases

black,

palpi

yellow.

Pronotumandmm.

mm;

unarmed,

combs

ofand

a small

twoDisti?
bearing

blackened
acutelysets of blackened

lobulecontiguous

provided

withapicalsetae.

Kameng, Assam,
6-8, 1961. ParaVol.

LXXVIII,

203

1970

December,

The closest allies of the present fly are Styringomyis
himalayana Edwards,
5. mitra Alexander,and some others, all differing chiefly in coloration of thestructure. The small acute point or beaklegs and wings and in hypopygialof the dististyle of this species is distinctive and has suggested the specific name.
Styringomyia
Thoraxbrownishyellow,brownrings;

two

narrow

noexcepting
darkenings
sternites
yellowed;ashield-shapedmale:hypopygium

mm;

6 mm;

about

brownish

Rostrum

5.5-6

about

Length

5 mm.

wing

light brown.
mesonotum

parallelon praescutumlongitudinal
stripesand pleura more uniformly
andtrochantersterminal

one

basitarsi

yellowed,

with

scape

stem yellow,

withandremainder

of

tarsi

brown.

knob
narrow

twofemora yellow withyellow;; tibiae withapex and a narrower

the apices

remainder

darkened,yellow.

withdarkerpatterned
brown,
includingrestrictedly
on either side of a paler centralarea, postnotum

Halt?resyellowed.

lightbroaderslightly

Antennaebrown.

palpi

yellow,

tobrownyellowish
Thoraxbrownishyellow,

mm.

4-4.5

wing

Head

coxae

with
femora yellow,
brown;and costal fields clearer yellow,
prearcular
mediumabdominalr-m;
uniformly
tergitesin a blackenedwithterminating
phallosome
darkerwithplate.

Lengthfemale:

over

cloud

small
male

brown,

n.sp.

melanaspis,

restrictedly
patterned
brownishyellow,wings

brown.

Legs

withallbrownrings,

thesubring at near midlengthbrown

brownishyellow,

Wings

faintly;

and broadcostalclearerthe veins,
fieldsveinsprearcularyellow,
includingremainingmore
not including
the membraneat r-m. Venation:
Cell 2nd Mt
darkened,
exceptslightly
to moreto margin.
roundednarrowly
broadlysessile; vein 2nd A evenly
Abdominal
mediumsternitesbrownishtergites
uniformly
brown,yellowed,
hypopygiumMalehypopygium
the phallosome.
Outeryellow.

generally

of

lobe

basal

tubercle.

Dististylewith

includingtwoflattenedyellow

blackenedpegs,

nearofbase

asblades,inner blade

shapedas a flattened
appearingyellow
structure
two protrudingwithpointsat posterior

tinctive,end,

slender,
their

outer

bordersa group

of

stem

that

spinesshield

with

the

apex

very

obtuse,ovalyellow

plate,

is tippedwithsetae.

Ninthsternitenarrow,

with

arow

in thestructure

is longer

thanarmature

ofof

dense

its
style

short

dis?
long dark setae. Phallosome
terminatesin a massive
shieldblackened

anteriorly
a smalltheespeciallyseta thatintermediate

unarmed;withbehindbroadly

holotype:

in himalayana,differinga single stout
withof basistyleouter arm

andterminating

terminating

moreseparatedpoint.

Ninthin a small

centralpoint

spinoid

setae.

two

blackenedin twostrong

smaller
atergitethat

Uttar Pradesh,Sar?n, Almora, Kumaon,
India, 7,200 feet,
1958
September 17,(Schmid). Allotopotype,9, pinned with type. Paratopo$ 9, Dhur, Almora, 7,400 feet, September
types, $ 9, with types. Paratypes,?,8, 1958; SS, Git Dabling, West
Bengal,5,100 feet,$ $, Labha, West Bengal, September 11, 1959 (Schmid).
The most

similar species

is Styringomyia

September

15,1959;

himalayana Edwards, which differs
the more conspicuous
darkened pattern,andincludingcoloration,
inthethedifferent
especiallyhypopygial
structure, including
quite
phallosome.
in wing204

New

Styringomyia

York

EntomologicalSocietyn.sp.

nigrobarbata,the darkened
withbrown; legs yellow,
chiefly brownish
yellow,
patterned
areas narrow and not forming
the restrictedbands ;wings
but clearly definedcompleteyellow,areas at r-m, m and m-cu,
brownvein 2nd A withouter
fourth morepatternincludingThoracic

dorsumabdominalweakly
darkened;of segments
darkernarrowly

tergites

witha palebrown;

male

hypopygiumoutersmoothbrownborders
longitudinal
stripe, posterior
withlobe of tergite
apicalcircular;and obtuse,
blade
short
blackenedwithinner bodythe
dististylecompact,darkenedsetae of lower margin
pegs very abundant,
abundant,crowded,
than in related
the ends blackened.
species;phallosome
chiefly yellow,
male:
female:andstouter

5.5 mm.

wing

6.5 mm;about

LengthRostrum

7 mm;

about

Lengthlonger

wing

6 mm.

Antennaewithscape brownbeneath,pedicelyellowed
palpi brown.
above,
Head
flagellum
yellow.
chiefly yellowed.
dorsumof thoracic
end of praescutumremainder
and anterior
light brown,
with
Halt?resto brownish
with
Legs
yellow.
pleura yellow.
yellow,
patterned
brown,

dark

andbrown,
Pronotumyellowcoxae

andtrochanters

twocluding

upper

in?
with
very
brown,
restrictedly
yellow,
patterned
the outertibiae yellow,
tip
subterminal,
spot
only,
a vagueat near midlength;
tarsi yellow,
darkening

femorayellow;onareas

surfacedark brown,fore pair witha restrictedbrown
withblackened.
patternsegment
clearly definedWingsyellow,notouterof vein 2nd A more weakly
fourththat includes
r-m, m and m-cu;
darkened,areas.
Venation:
Vein Rs
the membrane;in the patternedveins yellow
exceptincludingnarrowly
terminalcell 2nd M2
oblique;or recurved,
cell 2ndAbdominal
darkerrowly

himalayensis
outer arm

thanintovery

intoa

2ndveinA

curvedgentlynot

to margin,

stripe,spurredgenerallyas

Dististyle
the outerwith

circularinbladeof marginlonger andend blackened,
Phallosome
upperyellow,
chieflyspecies.
Ninthventral
the more
apex
tergite withpair divergent.
numerous
setae.
is clothed withthat
lobuleyellow

points,nar?

brodersposteriorhypopygiumseta.
spinoid

inner body of style a compact mass,
slender, unarmed,
thoseblackenedpegs very
abundant,
pale brownobtuse,of style abundant,
setae on lower margin
darkenedblack,

in related

short

small

sessile;a pale brownwithlongitudinalyellow
tergitesMalesternites moreyellowed.
uniformly
brown;
a singlewiththe lobe of basistyleand allies,

long andshort andsmooth,short and crowded;
stouternarrowly
A broad.

produced
produced

S, Lathong, Sikkim, 6,560 feet, July 26, 1959 (Schmid). Alloto
potype,5, pinned with type.
S.
from Styringomyia
TheEdwards,
himalayanapresent
fly differs
inthesomeothersandstructure, including
hypopygial
especiallymelanaspis,distinctive dististyle. The unusually abundant blackened setae on this structure
holotype:

name.

the specifichas suggested

Styringomyia
Headtibiae

andyellow,

thoracic

dorsumeach withover

twor-m, m

darkened,

extensively
brownrings;

cloudsrecurved

at outerend;

abdominalposteriorborders

darkerbrown;

tergites
male

wings

andand m-cu,

includingn.sp.

tenuispina,

the pleura
yellow,

the outeryellow,hypopygium

chestnut

restrictedly
thirdof veinyellow;

patterned
2nd A whichlongitudinallypatterned
withlobe of basistylewithfemorawithandbrown,

is strongly
pale

bearingbrown,
a singleVol.

LXXVIII,

spinoidseta;

phallosome
male:

December,

Rostrumdark

Head

Pronotumposteriora central

and

6.5-7

palpi

brown.

wing

4.8-5mm.

Antennae

withscapemm;

andpedicelandandgray,ascaudad

darkbrown.
Pleuraand pleurotergite
chiefly
mediotergite
scutellum,
coxae
stem yellowed,
infuscated.
withHalt?resapex of knob
Legs withtwo brownwithfemora yellow,chestnut
yellow;rings, the outer narrowat neardarkenings
awith
restrictedyellow,tibiae yellow,
with
genua
narrowly
blackened;outer segment
tarsi yellow,darkened.
apex;
Wings
and outerthat includes
clouds over r-m, m, m-cu

incomplete,andmidlength
brown
patternbrown

broken.

flagellum

black;

onlineyellow.
trochantersa paleabout

pruinose.
brown,
sparsely
withblackdark brown,
half of praescutum
and anterior
patterned
area of scutum yellowed,
this extended
and central
part of praescutumchestnut
andfour separate
complex,bearing
lobes, the inner two bladelike;
slender curved
points.
spine and six small blackened

dististyleunusuallya long
at apex with

Length205

1970

in cell M

washadjoining

vein

Cu;

veins

ofvague

2nd A, withof wing
yellowed,
2nd A
Venation:
Vein

third

anteriorof veinthird

near margins.
veins
darkened,
especiallyslightly morea spur.
at outer end, without
recurved
stronglya conspicuous
each withAbdominal
tergites yellow,
longitudinal
pale brown
an almostdarkerthese areas producing
continuous
borders
narrowlybrown,
remainingline, posterior
centralline;

outerMalewith
lobe of basistylebrownishyellow.
hypopygium
a small lobe on inner face provided
seta, at base the lobe withlong spinoid
arm long and slender, at nearsetae. Dististyleoutermostwithunusually
complex,the length with
one-third
three or four long erectin the usual
setae,
terminating
singleinner armature
two separate
flattenedbladesthat terminate
long spinoid
seta;
includingsternitespale,
a single
numerouswithhypopygium

a smaller more
the innermost
blade withbasal point
and a conspicuous
beaks,of unusually
inner blade
its blackened
beakshorterandlong setae,
slightly
smaller,outer margin
as a virtuallywithshort dense blackened
solid line;
pegs that appear
obtuse,betweena fourth extension,
the outer arm of style and the inner blades withlonger and nar?
in blackened

group

rower
at

than

lower

directed
into

the blades,
its inner marginsimilarwithshortend withfour small blackened
and apoints

dorsad;two small

outer
black

marginof phallosome

narrowly

dense

blackened

conspicuous

pegs.

slender
its dorsal

blackened,

Phallosome
curvedend

spine

producedpoints.

holotype:

1952 (D. C. Thurman).
S, Chiengmai, Thailand, May
Paratopo$, pinned with type.
in the wing pattern
resembles S. javana Edwards
Styringomyia
tenuispina
and venation, differing in the body coloration and especiallythe very distinct
type,

The
hypopygium.
ofthespine
phallosome.

malespecific

name wassuggested

bytheslendercurved