UNDESCRIBEDSPECIESOF NEARCTIC TIPULIDAE(DIPTERA), XICharles P. Alexanderl
New specie and subspecies of Tipulidae from -C4r,fornia andAnsrnacr.(TriTipula- (Trichotipila)
Alaska are described, iricluding
frommeri, .Tipula(Lunatipula)hastingsae- .d.ipe-rona,- f.rom
chotipula)sdnctaecruzae, and. Tipula(Pterelachisus)byersi frommacleani and, Limnophilaand. Tipula
California,
Alaska.

The preceding paper in this series was published in rhe GreatBasin N aturalist 29:l-70.
The species discussed at this time are derived frqm three sources:
two from California. the third from Alaska. The first Californianseries is from the California Channel Islands in the Channel Islands
National Monument west of Los Angeles including Santa Cruz, San
Miguel, and San Nicolas islands, the"materials hav'ing been collected
chieflv by Dr. Charles L. Remington, his son? Eric E. Remington,
and Ij. S6rge Matlovsky. The naried materials are preserved in the
Peabodv Museu-. Yal'e University. New Haven. Connecticut. The
collection is fiom various canyons in and nearsecond taHfornian
Death Vallev" Panamint Mountains. in Riverside and San Bernardino
counties, chietly in conjunction with the P. L loyd Deep Car]y-on
Desert Research Center. The survey is under the direction of Dr.
Saul I. Frommer, presently assisted by Messrs. Irwin and LePre.
The materials are io be pieserved in ihe Uttiversity of California,
Riverside, Research Colleclion. The Alaskan materials are from Dr.
George W. Byers of the University of Kansas, Lawrence; the specimens-havingbeett taken by Stephdn M. I\Iaclean, made in coniuncat Prudhoe'Bay,tion with th; United Statei Biome Tundra Program"ofthe above for t[eAlaska. I express my sincere thanks to all
privilege of eiamininf these particularly interesting species.
Tipula (Trichotipula)frommeri, n. sp.
General coloration of thorax yellow, praescutal stripes borderedbv nale brown. in cases the median area paler; less v6llow. tips of
f6mb.u narrowly brown; wings weakly darkened,"c6ll ,S" an<l thestigma light brown, no trichia in outer wing cells; abdominal tergites
coispicuiusly bicolored, bases broadly fellow, posterior bor:dersbrown, sterriites and hypopygium mor"e riniformly yellowed; malehypopygium with the b'erat"o"f the inner dististylb iong, with conspicuous setae.
Length about 12 mm; wing 10 mm; antenna about 5.5
Mer.r.mm.
FBuar-n.- Length about 13 mrn; wing 11 mm; antenna about
2.5 mm.
lAmherst, Massachusetts.

189

190

GREAT

BASIN

NATURALIST

Vol. 33, No. 3

Sept.1973

Described from alcoholic materials. Frontal prorongation of headslightly .shorter than the remainder,,yellow;fr".;, b;;;;;-.[;d;;;palpi, with proximal two.segments'Erown,'third sligh'tly paier, ter_.uq-ent elongate, light yellow. Antennae oimale'elongate;
Tllll
scape and pedicel
flagellum dark brown, flagellar seqments
wlth consprc-uous-yellow,
basal enlargements, verticils shorter than t[e segments. Head brownish yellow.
Pronotum yellow, center of scutum restrictedly darker. Meso_
notal praescutum with four yellow- stripes bordered by pare brown,
in cases median area paler t-o produce'a vague centrdl itripe. as indorsolineata and some'others; icutal lobes y?ilo*, ri"g"A #iiil p"r;brown; scutellum and,postnoium brownish yellow, *iitt
Pleura and pleurotergife yellowed, restricte,ity au.'t"."rlr"tuu.
o" ll."i."lregron, pr_opleura,and pleurotergite. Halteres with stem
"trJ yellow. knob
lntuscated. Legs wrth coxae and trochanters light yellow: femora
yellow, appearing darker from abundant short iarkened vestit,re,trps narrowly brown; tibiae obscure yellow, tarsi passing into darkbrown; ctaws.small,.simpJe. Wings 1Fig. 1y wealily daikened, cell
Jc and the stlgma trght brown; veins darker brown. stisma with
two or three microscopic trichia, outer cells wirhout trichii. venation: Petiole of cell IlZ, Irom about one-half to two_ti;iar- i; *_""
on Mn.
,^__-Abl"1lilal,tergites,conspicuously bicolored, bases broadly yelborders.blogn, on the proximal segments inciuclingll:,arso f-o:t".ror
rne narrow bases ot the,segments.;.sternitesanrl hypopygium
more,uniformly yellow. Abdorien wiih conspi.ror-,r lili.li'setae.
Ivlale hypopygium (Fig. a) with vestiture of tergal lobes. /. restricted
rn numbers, setae of emargination longer thanl group of about 15
small blackened points nea-r apex of eich robe. Bisistir", lr, *itt
group ot"with more sl)arse.strong setae on.oute{.in?rgip ofbristles on inner sid,e. Dististyles_,.d,-apex
asshown;
outerstylel9lg"f v.ery
rong.blact- setae_,ylln
longest exceeding transverse diametei ofstyle; rnner style wlth.beak long, setaeof
base long and conslit.uous,o^uter ones progressively shorter. apex of beak wiih ,rr-".nr..
u".ylong, more dellcate,yellow setae.
HABr.r.A.r.- California.
HolgtloE,.d,a,lcoholic,,P._L. Boyd Desert Research Cenrcr,
ttrversrde
County, Thunderbolt, Horsethief Creek, gagins station in-.
trrst permanent pool, Malaise, 15 May 1920 (sairl"I.'Frommer).
Ar-r,oropo_rypp,_ eeep Canyon, 4 mi. south of palm Desert. 56g0-g-,t:., 24 May 1969
(M. E. frwin and Saul I.'Frommer) ; 24_4T1.
PARAToporylE,_J, in Alexander Collection, 3.8 mi. soutir'of palm
L,esert,at Marker 58, 15 May 1969 (Frommer and Lapre):24_472.
This inte_restlng fly is named for Dr. Saul I. Frommer. capableglud.ent oj the Diptera, in charge of the Deep Canyon iles6arch
Prolect. -By_meansof the author's key to the Californian species oftne genus I tpula (Crane flies of California, pp. 56_5T; 1962;, thepresent fly runs to couplets-11_and12, including three speciesin thesubgenus 'l'richotipula that lack trichia in the oiter wing cells, these

ALEXANDER:

sr

NEW

5b

5,

CRANE

FL I E S

191

W,"t,ry40

Wry)nhr0
Fies. 1-3. Wing venation:7, Tipul! (Trichotipula) frommeri,-I1.sp'.;.?'
ri. sp; 3' Limnophila bvergi,y.sp' Figs' '+-10'
ripu'ti"'Qi"i"lorn;ii") ii"i"ii;,ei|r" 5vi"iv"si*'i,-rip"to-(iiiliot;p'"ta11'9mmeri, n sp'; 5a, Tipula (Trichoti(r*notipuia) cip:istranoAlexander;5c,pula) sanctaecruzae, ,J.,'.su,-iiiii
hast".u"trilti-birci Su.i."n, 6, ibove.Tipu.la,(Lunatipula)"fri,iti1il;iioiipitb
inisiehastinssae Alexander;

6" below. Tipula;:";;;;""'i;"diiti-<'i''7itl,ii'ij)';,;i-it;;iRl"*ir,d".,' s, ii;;,i"(Lunatipul!).

hasttngsa?. dtperona-io"t'"ani.-n..sp';8' riputa (vestiptez\ber(vest:iplez)aldrichiana; 1o^,Limnophila.bversi'(S y mbots : a' aeoeagus :i . sp. Fi s. 11. M al e antenna oI Li mnophi l a by ers .t, n' s p'd, dististyles; g, gonapophysis; s, sternite; t, tergite)'A. lliirt"i",

192

GREAT

BASIN

NATURALIST

Vol. 33, No. 3

being Tipula (Trichotipula)desertorum Alexander, T. (7.) dorsolineata.Doane, and {. (7.) kennedyana Alexander, all diifering evidently in hypopygial structure, especially the dististyles.
Tipula (Trichotipula)sanctaecruzae, n. sp.

Allied to beatula and capistrano; head and thorax patterned withdark brown-and yellow, mesonotal-mediotergitewith'a narrow yellow centra.lline; male hypopygium with terga-llobes truncate; appendage of ninth sternite iriclirding a long ou"ter lobe, the laterai lobe
Iow.
Mar-r,.- Length about 13 mm; wing 13.5 mm; antenna about
3 mrn.
FEMALE.- Length about 14-15 mm; wing 13 mm.
Frontal prolongation of head blackened. narrowly vellow onsides, ventrally more reddish yellow; palpi black. Aitennae withscape light. brorvn,_t_ippaler, iemainder of organ black; flegellar
segmentselongate. H-eadorange on sides,cerrtral part of vertei andocciput brownish black.
Pronotum brownish black. Mesonotal praescutum on disk withfour brownish gray stripes broadly-bordered'by black, lateral margins
of praescutum light orange; scutal lobes dark brownish gray, wilh anarrow ce-ntral p-olished black line, median region lighi yeilow;
scutellum light yellow with a narrow central blac"k line. "paraicutella
black; postnbtal mediotergite brownish black with a rrurio* medianyellow vitta on anterior half, in casesthis less distinct, anapleurotergite yellowed, katapler+rotergite with central part restricfedly yellowed. Pleura brownish gray, patterned with yellow, more e"vident
in certain individuals, including the dorsal iternopleurite, pteropleurite, and dorsal meral regioi. Halteres with stein yellow, basenarrowly orange, knob brownish black, apex restrictediy yellowed.
Le-gswith coxae brownish gray, trochanteri yellow; femoia brovmish-tibiae-browniih
yellow,dark brown;
yeliow, tarsi passing
fpile_s_.
into black. Wings
asinbeatulaandcapist'rano. differin[.generally
onlY rn mlnor detarls.
Abdomen with segments conspicuously patterned with brownish
black and- yellow, on tergites the dark" cblor including posteriorborder and a broad centralltripe. Male hypopyqium (FiI. 5a) with
ninth tergite, /,_having lobes-truncate, spinbid s6iie small ind sparse,
as_shown; median region bordering emargination with abuirdant
microscopic setulae. Appendage of ninth- stbrnite, s, shaped asshown, including a.majoi-outerlobe provided with numerorr .hiefly
marginal long [wisted' yellow setae, lateral lobe low. In beatula(Fig. Sg) tergite gen-erally similar, appendage of ninth srernite, s,
distinctive, wlth a slendtir lateral lo'be, as "shown. In capistrano(Fig. 5b) tergal lobes, t, more rounded; appendage of ninth lternite,s, distinc-tive, obtuse, outer lobe with abundant] very conspicuous,
long, yellow setae.
H4s11a1.- California.

Sept.1973

ALEXANDER:NEWCRANI]FL I E S

193

i9/u rm' to
Hororxrn, d, Santa Cruz Island', Channel [slands'
1970 (EricApril
4
Devil's peak,2/3t"i. i" U'5'-C'S' 1941''woodland'
I'
PenarYPE'
1970'
March
E. Reminston). arrt"rJ"ottf,-g-,?.5 University of California
lt"aio'iuf
C";;;i;.3tSanta Cruz Island,il;;;..h s;^ti;;,lb ep'iirszo lEricE' Remington)'beatula
(Trichot.i.pula.)
The mosr ,i-ilur"^ rf".iu, ur" Tiputa
amons
t. fi.5- iii ;tt i ono Alexander,all diff erinsand aport"r'5";i-;;the^;-Jespecialll',tergitethemselves i., hypopygiil structure,
andfigured'th;;i'itii-;6'nite, asdiscussed
ilft#;ithe crane flies'' cited underIn the California Bulletin conc-ernin-g
u-iE* tp"ti"s recordedfrom thethe preceding.p".i",itl;;';-;""Jt[
tttt"it *"t" idd"d in 'the collections;;;;;fChannel Islandsin the Peabodv Museum'""dmade bv the Reminit;;;' i;;P;"i"*"dwulpiana (BergNe.phrotoma
iitt"d'tt"t"utL,p".i".
The det-ermir,"dTiiuta (Tripli"iiroth). Ti puta (B"ti;i;;";;i'A;;;.'"'{lu*u"d"''pl anicornis
pula)
iciti
pit" (T r ipl
citiiut a)' occi denta li't*';t;;;,puIa)(tma.totiapiit'iii,
iit
Doine.r iputa Q r ipiic;ii;;i;)-i u a"nomy ia)'d.re itn"'to(W llnc
inna'
a
co
c
t
f
f uI ui I i neat aDoan e, ii ii [; ;. 1b'i

iiston). Limonia ftiliiili'niiQ -ry"fi?;ha-'(osten(7'rimicra) pilipes ('F'abrrctus'1'

Sacken)'and

Erioptera

hastingsae diperona' n' subsp'
antenna about
Length about 23 mm; wing 25 mrn;

Tipula (Lunatipula)Mer-e.5 mm.
20 mm'
FBryrer-n.- Length about 23 mm; winghastings'ae-Alexander
General coloration much as ! ^!ypi"ut
described from sev1951)"+OlaZ.gOi'(Bulletin Brooklyn'["'''S"",
aitttiU"tion being given ineral station, i., Curi?orr;; iit"' .d"L.iiJ
pases 43-44'
earlier'cited
asCutifotnia,
irt"'cr"""]li"rttandsizerarger(-"i;;a;;;h?;iii;"Li r6-tgmm)' Bodv

ii
u"'ttt subspecies.c!:l differwins coloratior,g"rrE#i,iy:ilii;;-i"*utu tt-ynopygium'especiallythe-iigq"tencelfound.in the ;ffi;i'"t;;i- ttt"
(iistii{sae' Fig' 6-'above;phallosome,u, ,n#'i"l;+;f.{t' in"the-preif':ill
dipnrona.Figure6 below)' Gonapophy-1":;
in hasttnssae;
Tateral spine at near mrdlength,^lacking
l"i"*".i"flong"andwith posterior spine ata,aeileagus,"p""*',lt^rrgutt"'tn"tyslender.
California'
Hearrar.Islan4 Channel Islands' at University
Hor.orvrn, d, 6;;;'t-'Canada del Medio' 18 April 1970
of Californiu n"ruui"h*St"li"",
9 i5 April i970; AlexanderAr-r-oropot"tu,(Eric E. Remington).
'
Collection'sp'
Tipula (Pterelachisus) macleani' n'sraY' praescutum meGeneral coloration of head and thorax
black; I"gt--Y]*
u"t"t"u"darker;*ithdially patterr,"a"ttitJ*"tvcla-wstoothed';
black'-wingsfemdrabrownish;;ii"*^iilJ""*"*1y*ii[,''n".y'pale brown; abdomen
iiiluiii,"u,i",',*i:iJrlfiy"p;ti;.;;J

194.

GREAT BASIN NATURALIST

Vol. 33, No. 3

dark brown, in cases tergites 2-4 reddened; male hypopyeium withcentral outer portion of lergite depressed tb form .i ttiirS*v"rro*
posterior margin with two,approximated triangular points;:3I::.,
elgnth sternrte wrth posterior border truncate, with inconspicuoui
pale setae.
Length about 12.5-13 mm; wing 12.5-13 mm; antenna, Y^,r".aoout + mm.
Fewrer.r,.- Length about 17 mm; wing l2.S mm; antenna about_.
c.c rnrn.
. Frontal prolongation of head black, gray pruinose; nasus con_sprcuous' wrth a terminal tuft of light yeilow setae;'palpi black.
Antennae black throuqhouj, scape sp"arseiy pruinos";'tt!g"iiu.
rugments with small basalie"gtfi.enlargeinents,-uertl.iis'i""*.6"di.tg
light Cr?y;.anterior velrtexmore
elevatEa,l"*,-.figl,tiy"i"_11-:11
tuscated; dorsal setae short, black, genae with long'light'yell"ow setae.
Pronotal scutum gray, sides of scutellum lighiveilow.
Mesonotal
praescutum light gray, with two intermediai" 5.o*r, stripei thatnarrow strongly behind, separated by u vaguely differentiatedground vttta, lateral stripes very pale to scarcely evident: median
p$rt 91 transverse suture dark br6wn; posterior"t.tu.it"i 6r-rrot,.rn
clear light gray; s-cutum with a narrow iinpressed median tirr". toru,
slrghtly variegate-dwith very pale brown; scutellum lightsray witha vague central darkening, paiascutella darkened anteiiorfv; 6ehindnarrowly light yellow; poitriotum uniforml.y cteu. tiehis.a+] Ft"*ul€ht gray, dorsop-leuralmembrane restrictedly obscire ieliow. Hal_
teres wrth stem obscure_Yellow,.brighter basaily, J<nobsrnall, brown,

yellow. L-egswith c6xaeand #;t;;;";;'lfiir;'s."y;?lT_:b'ple
temora obscure brownish yellow, tips narrowly black; tiSiae"lig"hibrown, tarsi darker p1owL; claw' with a sm"li,r"".
midlength.. Wings (Fig. Z) subhyaline,very restricteilv";;;;"iltl"t palierned
wrth-.

pate brown,
stigma darker; outer two_thirdj <if cell M
chretly very pale -oval
brown, this with a whitened area at near mid_length; verns brown. Trichia on certain veins beyond cord, lacking
on. most of
It-?, Rs, Mn and .Zst .4 except at base] vei.n 2id ,4 withexrensrve trrchra. In paratype spe-cimen, veins_that comprise longthe dxcepiibnoi.M"*,o,puf" to ,"Uoilroi"tu,fi:f\,!:!
Y" withthe, holotype"".ythese ele_mentsnormally darkened-.
venation: petiole ofcetl IVIl about one-halI m;
distal section of Cu, strongly recurved atouter end narrowing the cell.
Abdomen of holotype with basal segment and outer five tergites
dark brown,-their po_sierio-rborders .,r".f ,,uoo*ly y;ii;;;;i".[it".

sublaterally;all "sternites
ati't<b.o*-rirh groy,?-^1.::1{"t9,,_grayish
posterror borders very- n_arrowly yellow. paratype male witli'aL_!^"li:: I"1f"rmly dark brown. Female*iit-i"igit"r-d z'.ii"iiyouter.segments.darkened, sternite, a, in the male, thl
I-eldened?
extremebases of intermediate segments yellowed. ovipositor'with
cerci long, straight, very
-slende.;"hyporu[uae stouter, pioximal half
Dlackened,,aprces rounded.
Male hypopygium (Fig. 7) with ninth
tergrte' t, dark brown, posterior median portion slightly depressedto

S cpt. l 1)ZI

A I , I . )X A N D I i I i :NI']W

CRANIiFL I D S

195

subfonrr it llirttcrrtxl yellow saucer, margin produced into two low
rrrr.rl i i url ri i rttqulit r 1, , iint s;iet ae of iet gile pale, on- . dar kenedpar t sver yf' rrrr l i ttv v,' i i ,* put t t t ur es, vest it ur e of saucersm all and-pale'
shown; beak of inner style long andd,biiiiivt"t,i,',',ut,.i,i;1,,,',.-.
^,
with posterior border trunbl;rtkcned. Eighth sternite,-8s''1,'rr,L,i,,.,,t,:,'",;.iii;.u pit", ai.?ut tutuu small, ipu.se marginal bristles only
slightly longer.
Alaska.
Hesrrar.l l or-orveo, d, Pr udhoe Bay, 70o 18'! '{'Lat ', 1480 211W'! o"g''1921 (Sti:phen F. Maclean); Field Catalog=:,,ftit"Ju f l *,'{+'1""iwlth
No. 162; University of Kansas Collection' Ar-r-oroporvPE, -?,
j"Iv
No'
1971;
July
Pen'ttoPorYPE,d,t"OZt; No.338.
ifr" tto", t3"Alexander -Collection.
228*"hit;;
to. the collector,-stephen F' I4aclean' of
it;i;ii"t"u-"a Tundra Program' In rts-general.appearance?
the United StatesBiome
including the hypopygium, the fly suggestsvarrous l"9t9lt1t specres

Tipula .(Vbstipler)beri,iU*.""{ V iifp[i'Bezzi, lspedi-utty"nr."ii";o#Ti;;;"d"'"i"if,"r.
d"a'a)$.) -atd,ichianaAlexander?Fig.
?1ri-.'gi.U"iil'i."* et"rt?. fit" drawingsof the malehypopygia.of
the type ipecirnenspreservedin.theU"r"J,'h*%i#""fii;r-;;""united States National Museum. other less similar Arctrc Amerrcan
Alexander, T' i-ohanseni Alexander'
irr.i,rd" Tip"ti i"iiit;
tr";i;;
Alexander, of the Canadian Northwest Territory't"iiitorit;;;^i.
From the structure of the ovipositor
ei!*""d"..
;;; i: nrtriiiiiit
lt; pi"i""t fly is correctlv placed in the subgenus
i;';ip;;;t";ilP tei elachi sus Rondani.
Limnophila byersi, n. sP'
gray, head light gray'- anGeneral'"".tL* coloration of body brownish
afproximately, , onc-thirdmale"ofatttun.rLe
U.oua;
tu.io-.
Iensth of bodv; legs with femora yellow, tips brovunish black'-tarsr;LjBk' i"i"nr'rj.".,'?"ishvellow, st"igma ovai, dark brown; cell R'
one-half"cell R,; cell M,vety small' fromr"tt *i';;;;;';Tinhtlv
outer
u6o"t 8""-fifih to one-sixth its petiole; male hypopygitln] wrth
tf basistyle with numLrous very lon!- seiie, lobe. of rnesal*"*i"rnface"oval; outer dististyle narrowed o-n outer fourth, termmatlng
api*" .*"fi uppro*i-"t6d points; phallosome with gonaPophysest3-a^n acutep"i.i"g as t'imple slendei sickles, outwardly.lyrowe$times
sprne; aedeagus .t*pi", Iong and slender, striright, nearly three
the apophyses.
Mer,p.- Length about 9 mm; wing 7'8 mm; antenna about 3'2
mm.
Fnuer-e.- Length about 12 mm; wing 9 mm'
Rostrum short and broad, transversely oval,dark brown, sparselyo.,rirror"" with lonq-whitish'setae; palpi dark brown' Antennae ofbody.;scapeand.pedicelot"-ih^i.d^theil#'Glg: ir l^tofie,""utlybt".,uo"ishblack;. flage]lar.iegments withd^;;[ f;"";", it"g"tTn*;h;;i ;;;.; i.t"'r"tu" and spa.relongerbrisiles;intermediatefla-

196

GREAT BASIN NATURALISTVoI. 33, No. 3gellar segments about 4-5 times as long as broad, verticils shorter
fhan the" segments; outer segments ab6ut three iimes as long as
broad, subequal to their longest verticils; outer segment oval. Head'anterior"very
light gray;vertexbroad, about thiee times exposed
diameter of eyes.
Pronotum- brown, sparsely pruinose. Mesonotum almost uniformly brownish bHck, pruiieose, praescutal stripes not differentiated; a single median black pit on praescutum, pseudosutural
foveae larger, black; parascutella obscure yellow. Pleura gray;dorsopleuri'l membrane'obscure yellow. Halt6res with stem yejtow,
knob small, weakly darkened. Legs with coxae and trochanters yellow; femora yellow, tips brownisli black, on foreleg including a6out
one-third the"length, on posterior pair about one-fifth; tibiaeirownish yellow, tips very narrowly darkened; tarsi black. Wings (Fig. 3)
brownish yeliow, base and c6stal region'clearer yellow; stig-u 6rrui,dark brown; veins brown. Longitudinal veins beyond cbrd withblack trichia, lacking on veins-comprising cell 7st M"; further-oftrichia on Rs, outer lhird of basal seition
Cur, and on most of
2nd A. Venation: cell R, at margin sliehtlv less than one-half cell
Rr; cell M, very small, about one--fifth t-o one-sixth its petiole; rn-cu
shortly beyond one-third Ms*n.
Abdomen brownish black, sparsely pruinose, including the hypopygium. Ovipositor with valv'es stoit,^cerci obscure yellSw, hypovalvae black. Male hypopygium (Fig. 10) with basistyle, b, stout,
outer margin thickened, with numerous very long setae, longest
about one-Lalf outer disiistyle; cephalic end of mesil face'witlian
oval lobe provided with long pale setae. Outer dististyle, d, glabrous,
gently curved, outer fourth narrowed, terminating in two small approximated points, axial one stouter; inner style with base stou1,
with conspicuous setae? outer half narrowed, setae microscopic.
Phallosome with gonapophyses,g, appearing as slender simple curved
sickles, narrowed gradually into an acute spine; aedeagus simple,
very long and slender.
fl4s11'41.- Alaska.
_ Ifor-orypr, 6f , Prudhoe Bay, 70o 78'N. Lat., 74.8"2t'W. Long.,
altitude 15 m, June 1971 (Stephen F. Maclean); No.345. Ar,r,oroporypE, 9, with tlpe; No. 298. PeneroporypE, 6l , with types.
The species is named for my long tirne friend and fellow studentof the Tipulidae, Dr. George W. By6.r, of the University of Kansas.
The closest regional ally is'LimnoTihila'poetica Osten Sa6ken, widelydistributed in northern North America, from Massachusetts to British
Columbia, thence north to Alaska. This is readily told from the-antennae,present fly by the yellow body coloration, longer
vena- .
tion, espetialiy the" normal c6lI Mr, and' in ietails of the male I
hypopygium, including the basistyle, and the gonapophyses andaededgul of ihe phalloiome. The microscopically"bidd"tate apex of
the ou-ier dististvie is found in both snecies.

\

I

1
I