On N e w or little-known Tipulidse.

391

the apex, the median lobe is narrow at ttle base, progressively wider to half its length, then sub-parMlel, the apex
transversely truncate ; the whole organ is smaller than in
any other species seen.
The species is fairly easily recognizable by the regularserial punctures of the elytra becoming distinctly irregularposteriorly close to the suture, and the interstices between
Fi~'. l O.

Ps. w,wlbroecki d ' O r e h y m o n t .
2Edeagus, v e n t r a l aspect.
/, ] 50.

the punctures here range up to four or five times thediameter of t h e punctures against twice the diameter ofthe punctures on the disc. There is a very short, shallow
and somewhat indefinite impression at the apex close to
the suture forming the sutural stria.

XXXII.--Newor little-l,mown Tipulid~e (D@tera).-L X X X I . Oriental-Au.~'trala~'ian Npecies. By (IHAI~L~S

D. ALEXANDER, Ph.D., F.R.E.S., University of Massachusetts, Amherst, Massachusetts, U.S.A.
TuE materials considered at this time are chiefly from
New Guinea and Papua where they were collected by Miss
L. Evelyn (~heesman a n d Mr. Harry Hoogstraal;
additional species from New Herbrides and the Solomons
were taken by Mr. Jean Laffoon and Dr. Robert Staples.
Tile types of tile Cheesman materials are preserved in the392

Dr. C,. P. A l e x a n d e ronBritish Museum (Natural History) ; of the other species,in m y personal collection. I am greatly indebted tothese entomologists ff)r this continued co-operation in
making known the Titmlid fauna of this unusuallyinteresting region.

Itabromastix lucix, sp. n.
Me~onotal prwscutum obscure yellow with three brown
stripes, the median one with a fl~rther delicate hlackline on the cephalic p o r t i o n ; antennm (male) longer thanwing, yellow ; wings pale hrown, variegated with darker
brown areas a n d creanl-coloured spots ; Rs short ;Ma+, long, about two-thirds m-ca ;abdominal tergites
yellow, ringed with brown.
Male.--Length about l~imm. : wing IS ram. ; a n t e n n aabout 21 ram.
Female.--Length about 20 mm. ; w i n g 16 ram. ;a n t e n n a about 4"5 ram.
Frontal prolongation of head slender, about three-fourths
the remainder of head, brown: palpi dark b r o w n ;region of nasus with several strong set~e. Antenme(male} l l-segmented, longer t h a n the body or wing,
yellow', the outermost segments darker ; flagellar segments
very elougate-cylindrical, the more basal ones with longerect slender setae, with shorter bristles on the upper
face: on outer segments the vestiture becomes shorterand stouter : in the female, antennm shorter and darker.
Head dark reddish brown, more yellowed on o r b i t s :vertical tubercle low or lacking.
Pronotnm infuscated. Mesonotal pra, scutum with therestricted ground obscure yellow, with three brown
stripes, the median one split by ~ capillary blackened vitta
on about the cephalic fifth; seutum reddish b r o w n ;scutellmn and mediotergite more 1)ufl}', the latter moredarkened t)ehind. Pleura and plenrotergite chiefly greyish yellow, the propleura and mesopleura somewhat
darker. Halteres yellow, knobs infnscated. Legs withthe cox~e buffy yellow-: trochanters yellow; femora
obseut'e yellow or brownish yellow, the tips narrowly
blackene(t, more abruptly so in the female • tibiw yellowish brown, the tips narrowly d a r k e r ; tarsi ])assing into
black ; claws small, simple. Wings with the ground [)alebrown, variegated with cream-coloured spots and darker

New or little-known Tipulidm.

393

brown areas, the latter including marks in the costal
field, the more distinct stigmal darkening, and with about
three comparable darkenings along vein Cu, chiefly incell M, continuous with the costal darkenings across cells
R and M ; cream-coloured areas before cord conspicuous,
a little less extensive than the ground; beyond cord asingle major pale area, this very extensive, occupying the.~ubterminal part of the wing in the medial field, narrowly
broken by seams along the veins ; outer half of cell 1st A
uniformly of the ground colour ; a restricted post-stigmal
brightening ; veins brown. Venation : Rs short., a trit!emore than one-half R2+3; R1+ 2 short, oblique, vein R acorrespondingly long; petiole of cell M~ nearly twicem ; M3+ 4 long, as common in the genus, about two-thirds
the long oblique m-cu, the latter varying in position fromshortly before fork of M~+~ to just beyond the base of M 4.
Abdominal tergites yellow, conspicuously ringed withbrown, on tergite two at near midlength, on the succeeding
segments subbasal to basal in position; sternites more
uniformly yellow; in the female, tbe posterior borders ofsternites narrowly ringed with dark. Male hyp,~pygium(as described from the dry type) with the tergite short ;basistyle long. Outer dististyle elongate, narrowed outwardly. Inner dististyle slender, the dorsal surface or
crest with abundant dense black set~e.
Hab. Papua.
Holotype, ~, Kokoda, altitude 1200 feet, July 1933
(L. E. Cheesman). Allotype, ~, Mafuh~, 4000 feet, January
1934 (L. E. Cheesman).
This unusually interesting fly is named for Miss Lucy
Evelyn Cheesman. It is the first record of any species of
the genus in ~'ew Guinea, all others being Australian.
The fly is most similar to species such as Habromastix~'e~Jwta [Walker), H. terra~-reginx Alexander, and others,
differing in the length and colour of the antennae of themale and in the pattern of the body and wings.

Tipula (Acutipula) pedicioides, sp. n.
Size large (wing, male, 25 mm.) ; mesonotal przescntum
and scutellum buffy yellow, the intervening scutum
mfiformly brown; central part of mediotergite broadly
bt'owu, the sides whitened; pleura and pleurotergite
variegated with brown, buffy and white ; femora yellow,
Ann. d, Mag. Nat. Hist. Ser. 12. Vol. i.
27

394

Dr. C. P. Alexander on

the tips conspicuously blackened ; tibim with bases narrowly
blackened, followed by a yellow ring that is broadest on
the fore legs; remainder of legs darkened; wings lightgrey and subhyMine, with a conspicuous brown patternarranged is in the genus Pedicia, that is, at costa andeubitus, interconnected along the cord; abdominM
tergites dark brown, each with an elongate huffy markon lateral border; hypopygium with the lobes of thetergite low, truncated ; dististyles small and simple.
M a l e . - - L e n g t h about 25 mm. ; wing 25 mm. ; antenna
about 3"5 ram.
FrontM prolongation subequal in length to remainderof head, dark brown ; nasus elongate, blackened at tip ;palpi brownish black. Antennae (male) short, yellow, theouter six or seven segments very slightly more darkened ;vertieils elongate. Head brown, posterior orbits narrowly
yellowish grey; anterior vertex low.
Pronotum dark brown, sparsely pruinose, restrictedly
patterned with darker. Mesonotal prmseutum almost
uniformly buffy yellow, without pattern excepting thenarrowly infuseated humerM border; seutum uniformly
brown; seutellum yellow, the parascutella darker;
mediotergite broadly brown on central portion, the sides
extensively whitened ; pleurotergite chiefly silvery white.
Propleura and cervical region d a r k , mesopleura variegatedhuffy yellow and dark brown, the latter eolour appearing
as a narrow transverse girdle on the mesepimeron(pteropteurite), extending from beneath the wing root tothe sternum;
dorsopleural membrane dark b r o w n ;metapleural region dark brown with a silvery spot above
the hind eoxm. Halteres brownish yellow. Legs with theeoxm grey, the posterior pair more infnseated ; troehantersdark brown, the fore pair paler ; femora yellow, the tips
conspicuously blackened, subequal on all legs, on theposterior pair involving about the distal eighth; tibim
more narrowly blackened basally, followed by a yellow
ring on the proximal lmlf or less, broadest on the fore
legs, narrowest on posterior tibiae, the outer end of thesegment gradually passing into dark b r o w n ; tarsi darkbrown ; tibial spur formula 1-2-2 ; claws (male) toothed.
Wings light grey and whitish subhyaline, eonspieuously
patterned with (lark brown, the markings arranged nmeh
as in the genus Pedicia ; cell U more brownish yellow, ~qc

N e w or little-lcnown Tipulid~e.

395

darker; seam over the very oblique cord, at wing baseand on proximal half of vein Cu broad and conspicuous,
the latter completely crossing cell M on all but its outerp a r t ; stigma and outer radial cells darkened, the areas
separated by a white post-stigmal mark ; the grey colonr
includes most of cell R and the outer medial field, thelatter bordered internally by whitish, this pattern reaching
the wing tip at vein R4+5; dark mark along Cu interrupted shortly before tile comparable seam at cord, thelatter involving m-c,u, and all but the base of cell I st M.~ ;seam on distal section of vein Cu 1 narrow, more expandedat margin; veins yellow, brown in the patterned fields,
whitish subhyaline at cord. Wing outline weakly faleate,
the margin of the outer medial field being more or lessincised. Venation: Rs about one-third longer thanRe+a; R~+e preserved, cell R e relatively large, notconspicuously narrowed at base ; m about two and onehalf times the petiole of cell M , ; cell 1st M 2 diamondshaped ; cord very oblique ; basal section of M,+ e longer
than the second section ; cell 2,~/A broad.
Abdomen elongate; hypopygium moderately large;
tergites dark brown with an elongate buffy mark on
Lateral borders of segments; sides of basal ring more
restrietedly light grey; sternites and hypopygium more
uniformly brownish grey. Male hypopygium with thesuture between tergite and sternite deep ; basistyle smaLL,delimited only by the ventral suture, the lower apicalangle produced into a blackened spine. Ninth tergite
transverse, the caudal border with two low blaekened lobes,
the tips obliquely truncated ; lobes separated by a smaller
U-shaped median notch; margin of lobe near mediannotch with microscopic roughenings. Outer dististyle
broad, the length about one-third more than the greatest
width ; setm of moderate length only, scattered over thesurface. Inner dististyle relatively small and simple, thebeak separated from the large and more heavily blackened
lower beak. Eighth sternite not produced, its caudal
border unarmed.
Hab. Papua.
Holotype, c~, Mafulu, 4000 feet, January 1934 (L. E.
C,heesman).
No similar or allied species is known. The wing
pattern is superficially very like that in Plusiomyia
27 +

396

Dr. C. P. Mexander on

gracilis (Walker) and in various species of the genus
Pedicia LatreilleTip~da (Acutipula) cyclopica, sp. n.
Mesonotal pr~escutum brownish grey with three reddishbrown stripes that are conspicuously bordered by blackish;
antennm relatively short, flagellum weakly bicoloured ;wings moderately broad, infilscated, with a restricted palepattern; Rs suhequM to R2~:~; cell 2nd A b r o a d ;abdomen brownish grey; rome hypopygium with thetergite produced into two low rounded lobes; outerdististyle broadly flattened, terminating in a short
blackened spine; immr dististyle with the main body an
oval yellow disk, the t)eak short; lower beak large,
subglobular in outline, black.
Male.--Length about 22 mm. ; wing 17"5 ram. ;antenna about 3 ram.
Frontal prolongation approximately equal in length toremainder of head, brown : nasus stout, suberect ; pMpi
brownish black. Antennae (roMe) relatively short ; scapedark brown, pedicel reddish brown; flagellar segments
wealdy bicoloured, obscure yellow, the bases more infuseated; flagellar segments short-subcylindrieal, the basal
enlargement poorly indicated; vertieils longer than thesegments. Head above dark brown, including the oval
elevated vertical tubercle.
Pronotum of the type concealed, on sides silvery
prninose. Mesonotal prmscutum with the ground colour
brownish grey, with three reddish-brown stripes that areconspicuously bordered by blackish lines, the medianstripe vaguely divided by a paler vitta ; the broad lateralprwscutal border more buffy grey, at end behind thepseudosutural fovem more blackened ; scutal lobes chestnut brown, darker brown behind ; scutellum brown, theparascutella more reddish brown; mediotergite reddish
brown behind, darker in front and on sides. Pleura oftype crushed, variegated with reddish brown and darker
brown, the posterior sclerites, with the pleurotergite,
more silvery grey pru;nose. Halteres relatively long andslender, dark brown. Legs with the fore coxm reddish
brown, the posterior pair darker brown; troehantersbrownish yellow: femora obscure brownish yellow toNew or little-known Tipulid~e.

397

brown, the tips p~ssing into (lark brown ; remainder oflegs brown to brownish black; tibial spur formula
1-2-2; claws (male) toothed. Wings relatively broad,infuscated, with a restricted pale pattern, as ibllows:
Before and beyond a more darkened spot in cell Cu
before cord; a small post-stigmal brightening; conspicuous spots in bases of cells M 1 and 2nd M 2 ; a further
small marginal spot in cell M4; more restricted paleseams at ends of veins R4+ 5 M1, M 2 and M a ; a further
pale dot on basal section of vein M a ; stigma oval, darkbrown; paler brown areas over the anterior cord andm-cu; veins brown. Wing veins, excepting R and R1,
glabrous or virtually so ; relatively few trichia on distal
section of vein R4+ ~. Venation : Rs and R2+ a subequM ;RI+ 2 pale but preserved ; petiole of cell M~ about twothirds m ; m-cu at near three-fourths the length of
Ma+4 ; cell 2nd A broad.
Abdomen brownish grey, the basal tergites withindications of a darker median stripe or row of spots.
Male hypopygium with the caudal margin of tergite produced into two low rounded lobes t h a t are separated fromone another by a smaller rounded notch ; mesal margins
of lobes with abundant short dense spinous set,e, thoseof the apex and outer portions longer and more normal.
Outer dististyle broadly flattened, especially opposite
midlength, the apex a short blackened spine. Innerdististyle with the main body an oval yellow disk t h a t isdensely clothed with short silken yellow setm ; beak shortand relatively slender; lower beak a large subglobular
blackened structure ; posterior crest of style low, darkened,
provided with short coarse setm.
Hab. Northern Dutch New Guinea.
Holotype, ~, Mt. Cyclops, altitude 3500 t~et, March 1936

(L. E. Cheesman).
Superficially the present fly most suggests Tipula(Papuatipula) artifex Alexander, differing in colorationand in the structure of the male hypopygium. The
distinctions between the subgenera Acutipula Alexander
and Papuatiputa Alexander are not entirely satisfactory,
the latter evidently having been derived from the former
and representing a highly specialized type, best characterized by modifications in the radial field of the wing.

398

Dr. C. P. Alexander on

Tipula (Indotipula) querella, sp. n.
Allied to vilis; mesonotal pr~eseutnm reddish brown,with four poorly indicated obscure yellow stripes ; malehypopygium with the t ergal border produced into two
blackened lobes, each gradually narrowed outwardly, thetips unequally bilobulate; inner dististyle with theposterior lobe small.
Male.--Length, about 14 ram. ; wing, 16 ram.
Female.--Length about 19-20 ram. ; wing 16-17 ram.
Frontal prolongation relatively short, yellow, about onehalf the remainder of head ; nasus very long and slender ;palpi obscure yellow. Antennm with scape and pedicel
obscure yellow, flagellum black ; flagellar segments (male)strongly incised, as in the group; verticils very long.
Head above brownish grey, with a capillary blackish
median v i t t a ; vertical tubercle very low to scarcely
evident.
Pronotmn reddish brown. Mesonotal przescutum reddish brown with four poorly-indicated obscure yellow
stripes; posterior sclerites of notum brownish yellow,
the scutellum clearer yellow. Pleura and pleurotergite
yellow, silvery pruinose. Halteres brown, base of stem
narrowly yellow, knob dark brown. Legs with coxm andtrochanters yellow ; femora dark brown, the bases yellow ;tibiae and tarsi brownish black; legs long and slender;
claws (male) toothed. Wings brownish yellow, theprearcular and costal regions more saturated yellow;
stigma brown, continued basad as far as vein Sc~ ; veinsbrown. Venation: Sc~ preserved, See beyond it longer
than in serritergata; Rs subequal to R~+a; m-c~t somewhat shorter than the distal section of CUl; cell M 4
correspondingly narrower than in serritergata.
Abdomen dark brown, the basal rings more I)ruinose,
paler ; basal tergite more reddish brown ; outer sternites
yellow ; hypopygium and preceding segments blackened.
Male hypopygium with the tergite produced into two
blackened lobes, gradually narrowed outwardly andunequally bilobulate at tip, the outer angle longer; noevidence of serration of these tergal lobes is evident in
the type. Outer dististyle large, expanded outwardly,
unequally divided into two lobes b y pale membrane.
Inner dististyle with the lobule of the beak relatively

New or little-known Tipulidm.

399

short ; lobe on face of style near posterior end smaller
than in serritergata.
Hab. Papua.
Holotype, ~, Kokoda, altitude 1300 feet, in lower rain
forest by river, September 6, 1933 (L. E. Cheesman).
Allotype, ~, altitude 1200 feet, September-0ctober 1933
(L. E. Cheesman). Paratopotype, ~, with the allotype.
Although it is very closely allied to Tipula (Indotipvla)
serritergata Alexander, of nol'th-eastern New Guinea, I
believe the present fly to be distinct in the details of
structure of the male hypopygium, particularly the
tergite and the inner dististyle.

Tipula (Tipulodina) curtissima, sp. n.
Size medimn (wing, male, 15 mm.); wings whitish
subhyaline, the stigma and narrow apex with microtrichia,
producing darkened clouds; unpatterned parts of wing
entirely without microtrichia ; Rs very short, transverse.
only about one:half as long as the basal section of R4+ 5 ;RI+ 2 atrophied; male hypopygimn with the basistyleunarmed ; inner dististyle simple.
Male.--Length about 15 mm. ; wing 15 ram. ; antenna
about 3"5 ram.
Frontal prolongation short, about one-third the remainder of head, obscure yellowish brown, variegatedwith darker ; nasus long and conspicuous ; palpi brownish
black. Antennae with scape obscure brownish yellow,
pedicel reddish brown, flagellum brownish black ; segments
only slightly incised, subc~lindricM; verticils of outerface short, less than the segments, on the lower face
reduced to spinoid setm. Head medium brown ; anteriorvertex relatively narrow, about three times the greatest
diameter of the scape; vertical tubercle lacking.
Pronotum medially broadly dark brown, yellowed onsides.
MesonotM pr~escutum chiefly cove~ed by fournearly confluent brown stripes, the interspaces barely
evident; humeral region of prmscutum extensively andconspicuously yellow ; scutum chiefly brown, the centresof lobes with obscure brownish yellow areas; scutellum
dark brown, the parascutella still darker; mediotergite
chiefly dark brown with a relatively conspicuous obscureyellow area on either side of the central darkening;

40oDr. C. P. Alexander on

pleurotc,'gite with cephalic half dark browu, posterior
part above the root of the halteres couspicuously whitened.
t'leura extensively whitened, including a transverse area
ou the pteropleurite and on adjoining portions of themcsepisternum and dorsal meron; anterior portion of
mesepisternum, especially the anapleurite, more infuscared; propleura and adjoining regions chiefly pale;ventral metal region weakly darkened ; dorsopleural membrane pale. Halteres brown, knob blackened. Legs withcoxse pale yellow, whitish pruinose, especially the middle
pair; trochanters obscure yellow, the outer face slightly
infuscated ; remainder of legs of type chiefly detached ;femora brown, with abundant still darker setse, the tips
conspicuously whitened ; tibiae brown, the tips similarly
and more extensively whitened; basitarsi dark, with
about the outer fourth whitened ; second tarsal segment
dark, the tip whitened ; outer segments broken ; tips offemora with a brush of long black setse; vestiture ofwhitened portions of legs white, like the ground, theremaining vestiture black ; on tips of tibise with a row ofblack setse interspersed among the abundant whitevestiture. Wings whitish subhyaline, the tip in cells Sc.~
to 2111, inclusive, infuscated ; stigma and a narrow seamover the anterior cord weakly infuscated; veins very
delicate, brown ; infilscated areas due to the presence of
microtrichia, lacking elsewhere on wing. Venation : Rs
unusually short, transverse, only about one-half the basal
section of Ra+ 5 and subequal to +'-m, all three elements intransverse alignment; R,+2 atrophied, R a very long,extending parallel to +,he distal section of R4+ ~ for virtually
the entire length, the latter ending beyond the wing-tip ;cell M~ broad, about one-half longer than its petiole ; cell
1st M~ pentagonal, basal section of M 3 and m almost in
oblique alignment ; m-cu at fork of M.
Abdomen chiefly brown, discolonred in the type. Male
hypopygium with the tergite transverse, narrowed outwardly, the caudal border with a very broad and shallow
emargination, the lateral lobes very low and rounded.
Basistyle not produced. ])ististyles simple, the outerstyle a suboval hairy lobe ; inner style a flattened blade,
its lower edge nearly straight, provided with long setoe,
the upper margin with a deep emargination before theNew or liltl(:-/,:noa,n Tipulida,.

40 !

obtuse-trianguhtr beak ; near apex of s~yle, ol)posi~e thebeak, with a low darkened flange.
Hab. Solomon Islands.
Holotype, c?, Mount Austin, Guadalcanal, altitude 1000
feet, November 1944 (Jean LafJbon).
The subgenus had not been recorded fi'om the Australasian Region. Among the described Oriental species,
the present fly is closest to Tipula (T@ulodina) a,~abili~sAlexander, of Java, and T. (T.) gracillima Brunetti, of
Ceylon, both of which have a somewhat comparable great
reduction in the number of microtricia of the wing surface.
In the present fly, R.s represents the stlortest conditionso far found in the entire genus, approximating thecondition found in Scamboneura and in some Papuan
species of Dolichope, za.

Limonia ~Thryptico~t~yia) c,arissa, sp. n.
Mesonotmn dark brown, the lateral przescutal borders
more reddened; terminal tarsM segments snowy white,
involving the distal four-fifths of the basitarsi; wings
subhyaline, the outer third vaguely more darkened ; malehypopygium with the setm of the tergite very restricted,
including a compact group of three very long bristles nearouter lateral angle; rostra] spines two, rather widely
separated, the prolongation long and slender.
Male.--Length about 6-6"5 ram. ; wing 5"5-(i'~S ram.
ltostrum and palpi black.
Ant ennm with scape testaceous, the remainder of organ black ; flagellar segments
long-oval, with conspicuous dark-coloured apical necks;
longest verticils nnilaterally distributed, approximately
twice as long as the segments. Head black, very sparselyi)ruinose, the front and anterior vertex more heavily s(,,to appear ahnost silvery, anterior vertex relatively
broad, nearly three times the diameter of the scape.
Mesonotum dark brown, the lateral borders of theprmscutnm more reddened. Pleura and pleurotergite
reddish brown. HMteres blackened. Legs with coxm
reddish brown : trochanters testaceous yellow ; remainderof legs black, the oauter tarsal segments white, involving
the distal four-fifths of the basitasi and the succeeding
segments. Wings subhyaline, the outer third very

402

Dr. C. P. Alexander onvaguely to scarcely more darkened; stigma brown.
clearly delimited, long-oval; veins brownish black.
Venation: Sc~ ending a short distance beyond origin ofRs, Scj alone from about one-half to nearly two-thirds
the length of Rs ; free tip of See from two to three times
its own length before R e ; Rll e preserved as a spur t h a t
is about as long as vein R 2 ; inner end of celI R 3 only
slightly arcuated, lying just proximad of cell 1st M,~;
m-cu at one-third to two-thirds the length of cell 1st M~.
Abdomen brownish black, sternites a trifle paler. Male
hypopygium with the tergite transverse, the caudal margin
with a very shallow emargination; lobes low, withunusually few set~e on outer portion, including a closegroup of three very long but unmodified ones near outer
lateral angle and a further more isolated lateral bristle.
Basistyle with the ventromesal lobe stout, obtuse at tip,
without modified set~e. Ventral dististyle large andfleshy, its area approximately three times that of thebasistyle ; rostral prolongation long and slender ; spines
two, from small tubercles, rather widely separated, theinnermost on the style, the outer spine at base of the
prolongation. Gonapophysis with mesal-apieal lobe relatively narrow.
2Edeagus with setm on basal half,
including a transverse row of four very strong bristles atnear two-thirds the length.
Hub. Dutch New Guinea.
Holotype, d, Hollandia, in rain forest, October 20, 1944

(Hoogstraal).

Paratopotype, 3.

The most similar species is Limonia (Thrypticomyia)
sparsi~eta Alexander, of the islands of the South-western
Pacific. This likewise had the number of setm of thetergite of the male hypopygimn very reduced in number
but differs in all further details of the hypopygium,
especially the rostral prolongation and its armature.

Limonia (Thrypticomyia) dichzeta, sp. n.
Allied to dichromogaster; basitarsi with about theproximal third blackened, the remainder of tarsi white;~c~ very long, up to three-fourths the length of Rs ; malehypopygium with the tergite transverse, each lateral lobe
with several long unmodified setm; ventral dististylewith the rostral prolongation high, the spines slender,New or little-lcnown Tipulidm.

403

straight, without basal tubercles ; mdeagus with only twostrong set~e.
Male.--Length about 5-5"5 ram. ; wing 5"8-6"6 mm.
Female.--Length about 5 ram. ; wing 5-5"2 mm.
Rostrum light brown; palpi darker. Antennae black
throughout; flagellar segments oval to long-ovM, withunusually long, unilaterally distributed verticils in both
sexes. Head dark.
Mesonotum light brown, the postnotum and pleurapaling to yellow. Halteres with stem brown, knob
brownish black. Legs with the cox~e and trochanters
yellow ; femora and tibiae dark brown to brownish black ;basitarsi with about the proximM third blackened, theremainder of tarsi white. Wings with a strong dusky
tinge, beyond the whitened prearcnlar field uniform
throughout the length; stigma darker brown, oval,
relatively large; veins dark brown. Venation: Sc~
ending opposite origin of Rs, Sc~ very long, approximately
two-thirds to three-fourths _Ra, the latter about one-hMf
longer than R2+a; vein R 2 about three times its length
before free tip of Sc 2 ; inner end of cell R a moderately
arcuated ; m-cu at near one-third to one-hMf the lengthof cell 1st M 2 ; cell 2nd A long.
Abdominal tergites dark brown to brownish black, thesternites paler, brownish yellow; hypopygium brownish
black. Male hypopygium with the tergite transverse,
the caudal margin with a ve W shallow V-shaped em~rgination ; each low lateral lobe with about seven to ten longunmodified setm. Basistyle with the ventro-mesM lobe
stout, not elongate and without modified set~e. Ventral
dististyle large, the rostral prolongation high, darkeoloured, at apex extended into a short slender b e a k ;rostml spines slender, straight, placed near the upperedge of the base of the proiongation without basal
tubercles; spines separated from one another by adistance that is slightly greater than one-hMf the lengthof either spine. Gonapophysis with mesal-apieal lobe
relatively narrow gently curved to the acute tip. ¢Edeagus
with the lateral angles produced eaudad into slenderspines, near the base with only two strong set~e. In
dichromogaster, the tergite has a different conformation,
each lobe low, with about eleven setw; rostrM prolongation with the spines shorter and stouter, from basal

404

Dr. C. P. Alexander on

tubercles ttlat are t)laced itJlnost together; base ofmdeagus with eight strong sctm arranged in a generMly
transverse row.
Hab. New Hebrides.
Holotype, -~, Espirito Santo, Second Channel Area,
August 1944 (,lean LafJbon). Allotopotype, ~_.. Paratopo-tyl)e~., 5 3~.
The most similar described species is Limonia (Thrypticomyia) dichromogaster (Edwards), of Tahiti, which differs
in the bicoloured abdomen,the venation, and in the detailsof structure of the rome hypopygium, as described above.

Limonia (Thrypticomyia) tr@~sca, sp. n.
Mesonotal prascutum pale reddish brown ; legs black,the outer two-thirds of basitarsi and the remainder oftarsi snowy white; wings whitish subhyMine, the tip
narrowly infuscated ; Sc 1 ending opposite origin of Rs,
free tip of Sc 2 far before R~; male hypopygium withthe caudal margin of tergite with a V-shaped notch;
each lateral lobe with a darkened flange bearing three
powerful set~e ; rostrM prolongation of ventral dististyleslender, subpendant, the two spines in a transverse row
near base of prolongation.
Male.--Length about 6 a m . ; wing 6 mm.
Rostrum and palpi black. Antenna black throughout ;flagellar segments oval to long-oval, with short apicalnecks and very long unilaterMly distributed verticils.
Head blackened, sparsely pruinose.
Mesonotum brown, the prwscutum pale reddish brownlaterally ; postnotum darkened medially, the pleurotergite
obscure yellow. Pleura obsc re yellow, the stemopleurite
more infllscated. Halteres blackened, the base of stem
restrictedly paler. Legs with the coxa yellow, the fore
pair somewhat darker; trochanters yellow; remainderof legs black, the outer two-thirds to three-fifths ofbasitarsi and the remaining segments snowy white.
Wings whitish subhyMine, the tip narrowly infuscated,
the inner edge of darkened portion about at outer end ofcell 1st M2; stigma small, still darker brown; veinsbrownish black. Venation: ~b'c~ ending opposite originof Rs ; free tip of Sc 2 far before R,,, vein Rj alone about
three times R e ; R,,+a long, nearly two times Ra andNew or little-known Tipulidm.

405

snbequal to Rs ; m-ca at near three-fifths to three-fourths
the length of cell 1st M2, tile latter exceeding in lengthvein M2+ 2 beyond it.
Abdomen, including hypopygium, black. Male hypopy
gium with the caudal margin of tergite with a V-shaped
notch, each lateral lobe terminating in a darkened flangethat bears three powerfld set,e, with a smaller bristle
lying slightly more mesad; three unmodified setze nearlateral margin of tergite, arranged in a longitudinal
series. Basisty]e with ventromesal lobe long and slender,
constricted neat" base, without a moditled terminal bristle.
Ventral dististyle relatively large and fleshy; rostralprolongation slender, subpendant, the two rostral spines
in a transverse row near base of prolongation. Gonapophyses with mesal-apical lobe stout, dark-eoloure<l. Setm
at base of ~edeagus rather numerous but unmodified andnot in transverse aligmnent; apex of ~edeagus at eachouter angle produced into a slender spine.
Hab. Solomon Islands.
Holotype, c~, Bougainville, Empress Augusta Bay,
altitude 600 feet, December 2~, 1944 (Robert Staples).

Paratopotype, c~.
From the other regional species of the subgenus ha~qng
the tarsi extensively white, including Li~tw'nia (Tl~ryt~ticomyia) apicalis (Wiedemann), L. (T.) arachnophila
Alexander, and others, the present fly is readily told by
the structure of the male hypopygium, particularly the
tergite, as described.

Limonia (Euglochina) invocata, sp. n.
Female.--Length about 5 ram. ; wing 4"5 ram.
Closely allied to glens Alexander, differing in thedetails of coloration and venation.
Rostrum and palpi black. Antennse black throughout ;flagellar segments oval to long-oval, narrowed at ends.
Head brownish grey. Thorax almost uniformly testaceous
yellow, the central region of the dorsum a trifle darker.
Legs blackened, including about the proximal one-third
to two-fifths of the basitarsi, the remainder of tarsi snowy
white. Wings with cells beyond cord distinctly nmre
infl~scated than the basal ones, producing a dusky wingtip, the stigma only a little darker; veins stouter than insilens. Venation : Rs short and strongly arcuated ; free406

Dr. C. P. Alexander on

tip of Sc 2 and vein R 2 in transverse Mignment,; m - c u
about one-third its length beyond the fork of M.
Hab. Dutch New Guinea.
Holotyl~e, ~?, Hollandia, at light rain forest, April 17,1945 (Hoogstraal). Paratopotypes, 2 ~ .
L i m o n i a ( L i m o n i a ) alienata, sp. n.

General coloration reddish yellow, the pr~escutum witha conspicuous black median stripe ; wings with a strong
brown tinge, the small stigma darker brown; Sc long,
S c , ending just before the fork of R s ; m - c u shortly before
the fork of M ; male hypopygium with the caudal marginof tergite broadly notched, the lobes rounded; ventral
dististyle complex in structure, the body small, subglobular, bearing a long flattened bladelike appendage;
rostral prolongation about as long but more slender.
~ l l a l e . - - L e n g t h about 4"5 ram. ; wing 5 ram.
Igostrum and palpi dark brown. Antennee with scapeand pedicel yellowish b r o w n ; flagellum broken. Headwith the broad front silvery, the remainder of head lightgrey.
Pronotum brown medially, obscure yellow on sides.
Mesonotum reddish yellow, unpatterned except for aconspicuous black pr~escutal stripe, beginning near
cephalic border of sclerite, becoming paler behind, ending
before the suture. Pleura and pleurotergite somewhat
clearer yellow. Halteres brown, the knobs dark brown.
Legs with the coxm and troehanters yellow; remainderof legs broken. Wings with a strong brownish tinge, thesmM1 ovM stigma a little darker b r o w n ; veins darkbrown. Venation: Sc long, Sc 1 ending just before thefork of Rs, Sc 2 a short distance from its t i p ; Rs long,very gently curved, about one-half longer than basal
section of R4+ ~ ; free tip of Sc 2 and R,, in alignment ; cell
1st M e small, rectangular, about one-half as long as the
distal section of Ml+2; m - c u shortly before the fork of M ;~tna,1 veins convergent near their bases.
Abdomen with tergites brownish black, sternites
testaeeous yellow. Male hypopygium with the caudal
margin of tergite with a broad notch, each broadly rounded,with about four setm back from margin, tla.sistyleelongate, the ventro-mesal lobe very low and small,subbasal in position. Two dististyles, the dorsal one aNew or little-known Tipulida.

407

slender, nearly straight rod; ventral dististyle complex,
the body a small subglobular structure that bears a long
flattened glabrous bladelike extension or appendage;
rostral prolongation about as long but much more slender,
the tip narrowed. Gonapophysis with mesal-apieal lobe
elongate, truncated at tip. Aedeagus with apex bilobed.
Holotype, c~, Hollandia, in rain forest, January 21, 1945

(Hoogstraal).
In its general appearance the present fly suggests species
such as Limonia (Limonia) longeantennata Alexander and
L. (L.) mouicola Alexander, differing evidently in the
structure of the male hypopygimn.

Limonia (Limonia) hebridicola, sp. n.
Size small (wing 4 a m . or less); general coloration ofmesonotmn brown or yellowish brown; thoracic pleura
dark brown, with a conspicuous silvery white longitudinal
stripe; antenna (male) elongate, about two-thirds aslong as body, the apices of the fiagellar segments strongly
narrowed and glabrous, on outer segments becoming
slender petioles; wings with a strong brown tinge, thesmall oval stigma darker brown ; Sc short, cell M 2 openb y the atrophy of basal section of M 3 ; male hypopygiumwith apex of tergite emarginate, the lobes only slightly
produced; dististyle oval, its rostral prolongation very
long and slender, terminating in an acute spinous point ;mesal-apical lobe of gonapophysis bidentate at apex.
Male.--Length about 3-3"2 a m . ; wing 3"8-4 a m . ;antenna abou?~ 1"8-1"9 a m .
Female.--Length about 3"2-3"3 a m . ; wing 4 mm.
t~ostrum and palpi black. Antenna black throughout,in male of unusual length, approximately two-thirds thelength of b o d y ; flagellar segments long-ovM, strongly
narrowed at outer end which is restrictedly glabrous, onouter segments the tips further narrowed into slenderpetioles ; remainder of segments covered with a dense pale
erect pubescence. In femMe, antenna much shorter andmore nSrmM, the individual segments oval. Headbrownish black, more or less pruinose.
Thoracic dorsum brown or yellowish brown, the posterior selerites somewhat darker. Pleura dark brown, witha conspicuous silvery white longitudinal stripe extending

408

Dr. C. P. Alexander o nfrom behind the fore coxm to ttle base of abdomen. Halteres brownish bl~ck, the base of stem narrowly yellow.
Legs with eox~e brown; troehanters more testaeeous;remainder of legs brownish yellow, the outer tarsal
segments somewhat darker ; claws smMl. Wings with astrong brownish tinge, the smM1 stigma darker brown;
veius brown. Venation: Sc short, Scl ending about
opposite one-fifth to one-sixth the length of Rs, S%
exactly ol?posite this origin ; free tip of Nc~ and R 2 bothpgde, in transverse Mignment; cell M 2 open by theatrophy of basM section of Ma; m-cu at fi)rk of M,
subequM in length to distal section of C%.
Abdominal tergites black, sternites and bases ofhypol)ygim~l paler, obscure brownish yellow. Male
hypopygimn with the tergite narrowed posteriorly, theapex with a small V-shaped emargination ; lateral lobes
having about the same contour as the notch, somewhat
poorly demarked, each with four or five long setw placed
1)a.ck from margin. Basistyle with ventro-mesal lobe
undifferentiated. Dististyle single, the body oval, thesubequal rostrmn strongly narrowed, terminating in asmM1 acute spine ; no rostrM spines. Gonapophysis withmesal-apical lobe blackened, mmqually bidentate at tip,
the lateral point more acute, directed laterad, the small
apical point more oval, obtuse. ]Edeagus broad, eachhalf strongly notched at apex.
Hab. New Hebrides.
HoIotype, c~, Espirito Santo, Second Channel Area,
October 1943-August 1944 (Jean Laffoon). Allotopotype,9,pinned with type. Paratopotypes, ~9-.
Limonia (Limonia) hebridicola is allied to species suchas L. (L.) machidai (Alexander), wide-spread throughout
Asia and eastern Europe, differing in the elongate anteroomof the male, open cell 1st Me, and in the details of structureof the male hypopygium.

Limonia (Limonia) dipinax, sp. n.
Size small (wing, rome, about 3"5 ram.); pra~seutum
strongly gibbous, brown ; pleura yellow, with a brownish
black longitudinal stripe extending from the fore coxm to
the abdomen ; legs brown, the tips of the tibim and thetarsi paling to brownish yellow; wings with a strong
brownish tinge, the oval stigma darker brown ; cell M e

New or little-lcnown Tipulidm.

409

open by the atrophy of m ; rome hypopygium with thetergite narrowed into two slender glabrous lobes that are
separated by a broad U-shaped notch ; dististyle single,
the prolongation long, ending in an acute point.
Male.--Length about 3 ram. ; wing 3"5-:~'6 ram.
Rostrum and palpi black. Antenn0e black throughout :flagellar segments oval, well separated by apical necks.
Head brownish black.
Pronotum brown. Mesonotum with prmscutum strongly
gibbous, brown, somewhat darker mediMly; scutellum
somewhat more testaceous. Pleura yellow with a brownish
black longitudinal stripe extending from the fore coxm to
the abdomen, passing through the base of the hMtere, thelatter brownish black. Legs with eox~ and trochanters
yellow, the fore coxee darkened; femora brown; tibimbrown basally, paling into brownish yellow, the tarsi of
this same eolour ; claws apparently simple. Wings witha strong brownish tinge, the oval stigma darker brown ;veins brown. Venation: S% ending just beyond midlength of Rs, Sc 2 some distance from its tip ; free tip ofSc 2 and R 2 in transverse alignment, the latter a trifle thelonger; inner end of cell R a lying far based of otherelements of cord; cell M.~ open b y the atrophy of m ;m-cu close to fork of M ; anal veins convergent near
origin.
Abdomen, including hypopygium, black, the sternites atrifle paler. Male hypopygimn with the tergite narrowed,
produced into two slender glabrous lobes that are sepi~rated
b y a broad U-shaped notch. Basistyle with ventro-mesal
lobe large. Dististyle single, produced into a long curved
prolongation that narrows to an acute point, the total
length of the prolongation about equal in length to themain body of style ; a single very long seta at near onefourth the length of prolongation, with other smaller
bristles beyond. Gonap0physis with mesal-apieal lobe
stout, its tip obliquely truncate, the edge back from apex
with a low flange. 2Edeagus stout, the apex obtuse.
Hab. Dutch New Guinea.
Holotype, c~, Hollandia, at light in rain forest, April 18,
1945 (Hoogstraal). Paratopotypes, ~d, ,January 14-1S,
1945 (Hoogstraal}.
The most similar regional species is Limonia (Limonia)
hebridicola, sp. n., which has cell M 2 of the wings open
Ann. & May. Nat. Hist. Ser. 12. l'ol, i.
2s

410

Dr. C. P. Alexander onb y the atrophy of the basal section of M 3 rather than by
the loss of m as in the present fly. The two flies differ
further in the coloration and in structure of the antenn0e
and male hypopygia.

Limonia (Limonia) aureolenta, sp. n.
Size medium (wing, male, 6"8 mm.) ; general coloration
golden-yellow, the mesonotum lined with darker ; thoracic
pleura with a very conspicuous brown longitudinal stripe ;
antennae with basal flagellar segments suboval to transverse, the outer segments with very long verticils ; wings
with a strong brownish tinge, restrictedly patterned withdarker brown ; m-cu more than one-half its length beyond
the fork of M ; male hypopygium with the rostralprolongation of the ventral dististyle long and slender,
the two rostral spines elongate, with truncated tips.
Male.~-Length about 6 mm. ; wing 6"8 ram.
I~ostrum and palpi very reduced, black. Antennm
black throughout; basal flagellar segments suboval ortransverse, the outer ones cylindrical to long-cylindrical ;verticils of the more proximal segments short andunilaterally distributed, those of the outer segments
unusually long and conspicuous, exceeding twice thelength of the segments; first flagellar segment with aslender basal petiole ; first and succeeding segments witha stout glabrous apical neck, this condition persisting toabout the fifth or sixth flagellar segment. Head orange.
Pronotum above golden, darkened on sides. Mesonotum golden-yellow, restrictedly patterned with brown,
including conspicuo,~s pr~escutal borders and continuous
narrower brown stripes beginning on the prmscutum
behind the humeral region, extending caudad across themesal portion of each scutal lobe, sides of scutellum, andas sublateral lines on the mediotergite. Pleura ventrally
pale yellow, with a conspicuous dorsal brown longitudinal
stripe, exending from behind the genm across the propleura, anepisternum, dorsal pteropleurite and ventral
pleurotergite to the abdomen, its ventral edge unusuallyclearcut and more intense, the dorsal edge somewhat
paler and merging with the yellow dorsopleural membrane
and the sclerites surrounding the wing base. Halteresdark brown. Legs with the coxm and trochanters paleyellow; remainder of legs brown, the tips of femora,

New or little:known Tipulidm.

411

tibiae and outer segments of tarsi somewhat more darkened ; claws with a slender erect basal spine and a more
appressed one just beyond midlength of the inner face.
Wings with a strong brownish tinge, restriotedly patternedwith darker brown, including the prearcular field andsmall seams at origin of Rs, fork of Sc, cord and outer endof cell 1st M 2 ; veins brown. Venation : Sc moderately
long, Sc 1 ending about opposite midlength of Rs, Sc2 at
its tip ; free tip of Sc2 lying just proximad of level of R2 ;cell 1st M2 about as long as vein M~+ 2 beyond it ; m-cu
more than one-half its length beyond fork of 3//; vein
2rut A sinuous.
Abdominal tergites brownish yellow, the lateral andposterior borders of the segments brownish black, on thesixth and succeeding segments more uniformly blackened ;first sternite yellow, succeeding ones more infuseated,with an obscure yellow central vitta that is more narrowedon outer sternites and finally becomes obsolete; outer
segments and hypopygium more uniformly brownish
black. Male hypopygium with the tergite transverse, its
caudal margin truncate or virtually so ; caudal third ofsclerite with several black setm. Basistyle elongate, theventro-mesal lobe conspicuous, simple. Dorsal dististylea strong slightly curved rod, the tip acute. Ventral
dististyle with the body oval, the rostral prolongation
long and slender ; rostral spines placed at base of face of
prolongation, long and slender, their tips truncated.
Gonapophysis with lnesal-apieal lobe pale, long andstraight, the apex more or less cultriform ; lateral edge of'apophysis restrictedly blackened.
H,~b. Solomon Islands.
Holotype, ~, Mount Austen, Guadaleanal, altitude 1000
feet, November 1944 (Jean Laffoon).
Limonia (L~monza) aureolenta is allied to certain
Oriental species, particularly L. (L.) calianensis Alexander,of the Philippines, differing from these forms in thecoloration of the body and wings and in the details of
structure of the male antennae and hypopygium.

Limonia (Limonia) laffooniana, sp. n.
General coloration of the entire body orange ; rostrum
very reduced; legs black, the fore femora restrictedly
yellow at bases ; wings strongly petiolate, very strikingly
28+

412

Dr. C. P. Alexander onpatterned with whitish fenestrate areas on a blackened
ground; abdomen orange, the basal rings of the intermediate tergites glabrous and polished ; male hypopygiumwith the dorsal dististyle present; rostral prolongation
a long gently curved yellow blade without rostral spines.
Male.--Length about 7 mm. ; wing 9"5 mm. ; antenna
about 1"9 ram.
Rostrum very reduced, yellowish brown ; palpi reduced,
black. Antennae with scape whitish, pedicel abruptly
black, flagellum uniformly black ; basal flagellar segments
nearly globular, with short apical stems, the outer
segments passing through oval to elongate-oval ; terminal
segment about one-fourth longer than the penultimate,
slender, pointed at tip ; verticils unilaterally distributed,
exceeding the segments in length. Head orange ; anteriorvertex broad, approximately four times the diameter ofthe scape.
Thorax yellowish orange, unpatterned, the central
portion of the pr~escutum somewhat more intenselyorange ; setm of pr~escutum black, sparse, but relatively
long and conspicuous. Halteres black, the base of stem
restrictedly yellow. Legs with the coxse orange; trochanters yellow : remainder of legs black, the fore femora
restrictedly yellow at bases; claws with long strong
basal spines and a delicate appressed one at near midlength.
Wings very conspicuously patterned, the ground brownish
black, with large whitish subhyaline fenestrate areas inmost ceils, distributed as follows: At arculus; in cell
R at end of the basal petiole ; a tiny spot in cell 2nd A ; anearly complete band before origin of Rs, extending fromvein R to the posterior margin but interrupted by the
uniformly darkened cell CUl; in cell R 1 near base andagain near outer end, the latter prodnced behind acrosscell R a to vein R~ ~5 : two smaller spots in outer half ofcell R below Rs; major areas in outer ends of each ofcells M and C'u : beyond cord with further spots in cellR a near outer end, two in cell Rs, one at centre of cell
1st Mo ; an irregular mark in cells 2ztd M 2 and M a anda still larger spot iu cell M , : apical margin in outerends of radial cells paling to brown ; veins dark brown,even in the whitene(l areas. Wings mmsually petiolate,the stem extending to some (listance beyond the arculus.
Venation : ~qc~ ending about opposite the fork of Rs, S%

New or little-known Tipulida~.

4t3

at its tip; R~ strongly arcuated ; inner end of cell R alying i~r basad of the other elements of tile cord ; R 2
lying about its own length beyond the fi'ee tip of Sc 2 ;cell lot M 2 longer than vein M1+,, beyond it; m - c u close
to the fork of M, longer than the distal section of Cu I ;cell 2nd A small; both anal veins arcuated.
Abdomen orange, tile basal rings of the intermediate
tergites highly polished and nacreous, those of segments
three and four broadest, oll the succeeding segments
becoming gradually narrower; sternites orange-yellow;
hypopygium chieiiy yellow, the basistyles light brown.
Male hypopygium with the tergite transverse, the caudal
margin gently emarginate ; lobes very low, with about adozen long black set~c on each. Basistyle with theventro-mesal lobe large, its apex cap-like, narrowed andslightly more sclerotized.
Dorsal dististyle yellow,
straight, the apex strongly curved, narrowed to an acutepoint. Ventral dististyle with the body relatively small,less than the ventro-mesal lobe of basistyle, with scatteredelongate set~e and abundant delicate setul~e; rostralprolongation a long gently curved yellow blade, the apex
subobtuse ; a few scattered setul~e at apex and a single
powerful seta on outer margin at near midlength but
without rostral spines. Gonapophyses broad, the mesalapical lobe slender, gently curved.
Hab. Solomon Islands.
Holotype, (~, Mount Austen, Gnadalcanal, altitude 1000
feet, November 1944 (Jean LafJbon).
I am most pleased to name this striking tty for thecollector, Mr. Jean Laffoon, who has added materiallyto our knowledge of the Diptera of the Pacific Theatre.
The species is entirely different from those previously
described. Superficially it more resembles various speciesof the subgenus Dapanopte'm Osten-Sacken, but thesubgeneric reference here given is correct.
Orimerffa (Orimerga) sp,doptera, sp. n.
Anterior portion of thoracic notum yellow, the posteriorhalf, with the pleura, black ; knobs of halteres infuscated ;wings narrow, whitish subhyaline, with a restricted pale
brown pattern ; Sc~ ending about opposite midlength ofRs; RI+ 2 very short; r-m in approximate transversealignment with the basal section of M1+2; abdomen

il 4

Dr. C. P. Alexander o,J~

bieoloured, the broad bases of tile segments yellow, thenarrower apex brown.
Sex !--Wing 4"8 mm.
l~ostrum dark brown ; pMpi brownish t)laek. Antennae,
the scape and pedicel dark brown, flagellum brownish
yellow; flagellar segments short-cylindrical, with shortvertieils ; antenna~ broken beyond the eleventh segment.
unusually long for a member of this genus, if eomlolete
and bent backward extending nearly to the wing root,
Head black.
Pronotum and cephalic portion of proeseutum yellow,
the posterior hMf of the latter, espeeiMly on the sides,
conspicuously blackened; remainder of notum paler,brown or brownish black. Propleura yellow, the remainder of pleura polished black. Halteres with stem
white, knob infllseated. Legs with the eox~e andtroehanters yellow; remainder of legs broken. Wings
narrow, whitish subhyaline, the base and narrow costal
border yellow ; a restricted pale brown pattern, as follows :Origin of Rs, cord, fork of Ma+4, m-cu and a more diffuse
wash near the wing-base; veins pale brown, darker inthe patterned areas, yellow in the costal and preareularfields. Maerotriehia on the veins beyond cord, excepting
Cu. Venation: Sc short, Sc t ending about opposite
midlength of Rs, the latter angulated at origin ; R~+., very
short to virtually lacking ; R2+ a just less than twice R,, ;basal section of R4+5 areuated, lying a little proximad ofr-m, the latter in approximate transverse alignment withthe basal section of 3//1+,, ; Ma+a subequM to or a little
shorter than vein M 4 ; m-cu opposite base of _R~.
Abdomen elongate, conspicuously bieoloured, the broadbases of the segments yellow, the remainder, including the
distal third or more, brown; abdomen broken at. thesixth segment.
Hat). Dutch New Guinea.
Holotype, Sex ? Hollandia, at light in rain tbrest,
J a n u a r y 21, 1945 (Hoogstracd).
The only other Australasian members of the subgenus
that have the wings patterned are Orimarga (Orimarga)
joana Alexander, of New South Wales, and O. (0.) pietula
Edwards, of the Kei Islands and New Guinea. Both ofthese differ markedly in the venation and in the quite
different wing pattern.

New or little-known Tipulidm.

4i5

Orimarga (Orimarga) soluht, sp. n.
General coloration brownish grey, the prmscutum
without clearly defined stripes; wings yellowish grey,veins very pale ; macrotrichia of veins very sparse; RI+ 2
about one-half R2; R2+ a short, approximately one-hMf
longer than R 2 ; m-cu opposite one-third the length of Rs.
Female.--Length about 4"5 ram. ; wing 3"8 mm.
Rostrum black, sparsely pruinose ; palpi black.
Antennm with tile scape and pedicel black, the former
pruinose ; flagellum broken. Head grey ; anterior vertex
broad.
Pronotum brown. Mesonotmn brownish grey, without
clearly defined stripes. Pleura and pleurotergite reddish
brown, sparsely pruinose. HMteres uniformly pale. Legs
with the coxm reddish brown, troehanters more testaceous ;legs long and slender, the femorM bases a trifle brighter.
Wings yellowish grey, unpatterned, the prearcular andcostal fields dearer yellow'; veins very pale brown.
Macrotrichia of veins very sparse, beyond the cord
restricted to scattered series on the outer third of veins
R a, R4+ 5, M~+~ and Ma: Venation: Sc 1 ending about
opposite two-thirds the length of Rs, the latter angulated
at origin; R~+ 2 about one-half R2; R~+ a short, about
one-third to one-hMf longer than R2; basal section of
R4+ ~ long, nearly square at midlength ; r - m nearly intransverse Mignment with the subequM basal section of
M1+2; Ma+ 4 nearly equal in length to vein M 4 ; m-cu
about opposite one-third the length of Rs ; cell 2nd A
relatively broad.
Abdomen uniformly brownish black. Ovipositor withthe cerci smM1 and slender, gently npcurved to the acutetips ; hypovMvm ending about on a level with the cerci.
Hab. Dutch New Guinea.
Holotype, ~, Hollandia, at light in rain tbrest, November
28, 1944 (Hoogstraal).
Of the various species of Ovimarga so t~tr made kl~own
from the AustrMasian Region, the present fly is quitedistinct from all other forms so fnr discovered in New"
Guinea. Among other species it is closest to Orimarga(Orimarga) toala Alexander, from the mountains ofcentral Celebes, a larger form with the venational details
distinct. Unfortunately the tomes of both of these
species are still unknown.

416

On New or tittle-t~nown Tipulid~e.

EtJ.ii~hraflma (E upolyphragma ) staple si, sp. n.
Mesonotmn in front obscure yellow, the posteriorsclerites and the pleura more blackened ; femora yellow
with two broader darkened rings; vestiture of legs
including elongate scattered scales interspersed among theset,e; wings brown, variegated with darker brown andobscure yellow areas ; male hypopygium with the tergite
very reduced; interbase a strong rod, at and near tip
with two erect spines ; outer dististyle hairy.
M a l e . - - L e n g t h about 7-~ ram. ; wing 7"5-8"5 ram.
Female.--Length about 10 ram. ; wing 9"5 ram.
t~ostrum black ; palpi black, the basal segment obscureyellow. Antennae relatively short; scape and pedicel
brownish yellow ; basal three segments of flagellum yellow,
remaining segments black; all flagellar segments distinctly separated, without fusion; outer segments becoming long-cylindrical, the verticils relatively short andinconspicuous. Head grey, the orbits more infuscated ;occiput paling to yellow.
Pronotum narrow, brownish yellow. Mesonotal prmscutmn and scutum obscure yellow, the cephalic borderof the former restrictedly more blackened; scutellum
and postnotum dark, with a yellow pollen. Pleura andpleurotergite chiefly blackened, restrictedly patternedwith paler, more pollinose areas. Halteres long, stem
brownish yellow, knob brownish black. Legs with thecoxm black, the fore pair variegated with brownish yellow
at near midlength; tr0chanters yellow; femora yellow
with two broad darker rings, the outer one subterminM,
the more basal one broader but ill-defined, on the proximal
half of the segment ; tibiae and tarsi light yellow ; vestitm'e of legs with elongate flattened scales interspersed
among the normal setm. Wings with a strong brownish
tinge, variegated with darker brown and obscure yellow
areas; costal border clearer yellow; the darker marks
are at the ends of the longitudinal veins, larger in theradial field and over 2ncl A ; a major dark area at originof Rs ; other seams at arculus, along cord, outer end ofcell 1st M e and at fork of M ~ e ; the yellow spottings arescattered over the surface, usually alternating with thedarker areas, as described; veins brown, yellow in thebrightened areas. Venation : Sc 1 ending opposite fbrk of
R2+.~+4, Sc~ at its tip ; supernumerary cross-veins in cellOn Pgntaliine Cixiid~e, Kinnarid~e and Tropiduchidm. 417
C varying in number u p to seven ; R~.~a+4 fully two to
three times r-m ; petiole of cell M 1 subequal to or longer
than m ; m-cu about its own length or less beyond thefork of M.
Abdomen with tergites brownish yellow to brown,sternites more yellowed; hypopygium brownish yellow.
Male hypopygium with the ninth tergite very reduced, its
position chiefly occupied by the large membranous
proctiger. Interbase appearing as a strong rod, at andnear tip with two spines placed at right angles to theaxis, the lower spine longer and broader. Outer dististyle
hairy, terminating in a single recurved point ; inner stylelonger.
Hab. Solomon Islands.
Holotype, c~, Bougainville, Empress Augusta Bay,
altitude 650 feet, December 31, 1944 (Robert Staples).
Allotopotype, ~, altitude 600 feet, December 28, 1944.
Paratopotypes, 2c~c~, one each with the allotype andholotype.
I am very pleased to name this interesting fly for thecollector, Dr. l~obert Staples, to whom I am greatly
indebted for numerous Tipulidm from BougainvilleIsland. This is the most easterly known species of thesubgenus, the others being from Celebes and the Philippines. Among the somewhat numerous Philippine species,
the fly is closest to Epiphragma (Eupolyphragma) crenulata,
Alexander, differing conspicuously in the structure of themale hypopygium. For a discussion of the subgenus (asPolyphragma Alexander), a paper by the writer may beconsulted *XXXIII.---New Pintaliine Cixiidm, Kinnaridm and Tropiduchid~e from the Lesser Antilles (Homoptera : Fulgoroidea). By R. G. FEllAH.
ThE first species of pintaliine Cixiidm to be recorded fromthe West Indies was Flata variegata described by Fabricius
from material from an unspecified island. In 1881
Lethierry examined a species of Pintalia from Guadeloupe
which he referred to the South American pictipennis St~l,* Philippine J o u r n . Sci., xlix. pp. 257-262, pl. 4, figs. 44-60 (1932).