On new o~, llttle-known Tipulid~e.

177

X I X . - - N e w or little-known Tipulidm (Diptera).--XXXIV.
Australasiau Specie.~. By CnartL~s P. AL~XANDEa, Ph.D.,
F.E.S., Massachusetts Agricultural College, Amherst,
Massachusetts, U.S.A.
IN the present instalment the writer has continued his consideration of the Australian and Tasmanian species of thegreat genus Molophi/us. The species herein described belong
for the most part to the ruficollis subgroup of the 5racilisgroup, the typical form of which (ruficollis, Skuse) was
figured in the preceding part under this general title. In
this subgroup, which is very extensively developed in Australfi~
and Tasmania, the ventral lobe of the basistyle is the only
one that is well preserved and it is unarmed at the apex.
The only known species of the verticatis group (verticalis,
sp. n.) is likewise described at this time. This group is
distinguished by the great development of the dorsal lobe of
the basistyle, which is produced at apex into a spinous pointor blade.
As before, the majority of the species discussed at thistime were collected in New South Wales, Victoria, South
Australia, and Tasmania by Dr. Andrd Tonnoir, to whom thetypes have been returned. Additional species were collectedby Messrs. Walter Heron, Ferguson, and Maekerras in New
South Wales, and by Mr. Hardy iu Tasmania. I express
my sincere thanks to the above-named gentlemen for theirkind co-operation in making known this rich crane-fly~auna.

Molophilus amiculus, sp. n.
Belongs to the gracilis group, ruficollis subgroup ; generalcoloration pale brownish yellow; halteres with brown knobs;
wings tinged with greyish yellow, the base and costal region
more saturated yellow ; male hypopygium with the ve,~trallobe of the basistyle unarmed apically; outer dististyleshort-stemmed, the two arms conspicuous, one acutely
pointed; basal dististyle a pale arcuated blade with four
small spinous tubercles near apex.
Male.--Length (excluding head) about 3 ram. ; wing
4"3 ram.
Head of the unique type lacking.
Pronotum brownish testaceous. Mesonotum pale brownish
yellow, shiny, with conspicuous erect black sct~e. Pleura
more yellowish. Haltere, pale, with brown knobs. Legs
Ann. & May. N. Hist. Set. 9. Vol. xix.
12

178

Dr. C. P. Alexander onwith the coxm and troehanters pale yellow ; femora brown,
paler basally ; fore legs broken; tibiae and tarsi passing
into darker brown. Wings tinged with greyish yellow, thebase and costal region more yellowish ; veins a trifle darker,
with relatively sparse and short brown setm. Venation: r
ending about opposite r - m ; vein 2nd A ending about
opposite the caudal end of the oblique m-cu.
Abdomen pat~ brown, the hypopygium somewhat moreyellowish. Male hypopygium with the ventral lobes of haststyle prolo~ged into a slender lobe, the tip unarmed except
for a cluster of four or five setm; outer face of style with
scattered setm, the mesal edge with tiny erect setulm. Outerdististyle bifid, the stem paler than the arms, relatively shortand stout, the outer arm a long, slender, black spine that isabout equal in length to the stem, the tip acute, just beibrc
apex on the mesal face with a microscopic denticle ; innerarm much shorter, dilated into a broadly oval blade, thoouter edge thinner and paler. :Basal dististyle a flattened
arcuated blade, a little dilated beyond the base, the tip narrowly darkened, before the apex with an oblique linear row
of four small tubercles, each set with a stout microscopic
spine. ~]~deagus long, straight, slender, about one-half longer
titan the basal dististyle.
Hub. Tasmania.
Holotype, ~ , King River, altitude 500 feet, February 4,
1923 (-4. 1'onnoir).

Molophilus ampliatus, sp. n.
Male.--Length about 3"3 ram. ; wing 4"2-4"3 ram.
Female.--Length about 3"2 ram. ; wing 4"5 ram.
Closely allied to M. lucidil~enuis, Skuse, differing chiefly
ia the structure of the male hypopygium.
Lateral pretergites of pr~escutum a trifle brightened.
Halteres pale, the knobs faintly infuscated, tile setal covering
dark. Wings with m-cu beyond the fork of M.
Male hypopygium with the ventral tobe of the basistyle
produced into a relatively short blade that is slightly dilated
outwardly and obliquely truncated at apex ; margin of apex
with a delicate fringe of sctul~e and scattered conspicuous~,etm ; outer face of style with a few scattered setm. Outerdististyie smaller, the apex darkened, the apical spine gradual,
gently curved, the basal spine a little smaller, but long and:~.cute~ the margin of the style between these two spines
U-,shaped ; basad of the basal spine~ near mid-length of thestyle, a long, slender, dig~tiform lobule, pale in colour,new or little-knozon Tipulida~.

179

provided with a couple of setigerous punctures. Inner dististyle quite as in lucidipennis, bent upon itself near base, thelong distal portion sinuous~ the tip acute. }Edeagus long,
slender, iu slide-preparations appearing almost straight,
extending caudad to opposite two-thirds the length of thedilated ventral lobe of the basistyle.
The female has the antennte correspondingly shorter, theflagellar segments short-oval. Ovipositor and genital segment reddish horn-eolour; tergal valves of ovipositor long,
slender, the tips bent slightly upwards.
Hab. Tasmania.
Holotype, ~ , Cradle Valley, January 10, 1923 (.4.
Tonnoir).
Allotype, ~, Mt. Field, December 18, 1922 (A. Tonnoir).
Paratopotypes~ 7 ~ ~, 2 ~ ?, January 10-13 and 24-27,
1923; paratypes, 1 ~', with the allotype~ 1 ~', Adventure
Bay, December 31, 1922 (A. Tonnoir).

Molophilus exiguus, sp. n.
Belongs to the gracilis group, ruficollis subgroup; size
small (wing, c~, under 3 ram.) ; general coloration testaceousbrown, tile pleura dark greyish brown; vein 2nd A short
aud straight, ending some distanee before the short transverse m-cu; male hypopygium with the basistyle unspined
at tip ; basal dististyle bifid.
Male.--Length about 2'3 ram. ; wing 2'6 ram.
Rostrum and palpi brownish black. Antennae ( c ~ ) o f
moderate length, if bent backward extending to shortly
beyond the wing-root; seapat segments ochreous brown, theflagellum dark brown; flagellar segments oval with a conspicuous, erect, pale pubescence. Head grey.
Pronotum brownish testaeeous~ the lateral pretergites narrowly yellowish. Mesonotal pr~escutum light testaceousbrown, the median line narrowly greyish; pseudosutural
fovea~ linear, shiny reddish brown; scutum and seutellum
concolorous with the pr~escutum, the postnotum dark greyish
brown. Pleura dark greyish brown. Halteres relatively
short, brownish testaceous, the stem a little darker. Legs
with the cox~e and trochanters brown; femora and tibia~
testaceous brown, with dark set,e, the tarsal segments passing
into brown ; fore legs broken. Wings comparatively l~urrow,with a light yellow tinge, the veins brownish yellow withslightly darker brown maerotrichi~e. Venation: r apt)roximately in alignment with r-m, the latter connecting with
R 5 shortly beyond its origin ; vein 2nd A unusually short and
12"

180

Dr. C. P. Alexander on

straight, ending some distance before the short transverse
Abdomen dark brown, including the hypopygium. Male
hypopygium with the ventral lobe of the basistyle produced
caudad into a broad fleshy lobe, the mesal-apical angle more
conspicuously sctiferous than the remainder, but otherwise
unarmed. Dististyles two, arising close together, the outerwith a long stem, shallowly bifid at apex, the outer arm
gently curved, the apex truncated and blackened ; inner arm
a triangular arched blade. Basistyle a little lorrger andstouter, gently curved to the acute blackened apex, before
the tip with a slender acute spine; apex of style beyond thespine about twice the length of the latter. _/Edeagus relatively
short but slender, arcuated.
Hab. Tasmania.
Holotype, ~ , Zeehan, altitude 200-300 feet, February
1924 (G. H. Hardy); collector's number 372.
Type in the collection of the Queensland University
Museum.
Molophilus adamantinus, sp. n.
Male.--Length about 3"2 ram. ; wing about 4 ram.
Closely allied to M. exiguus, sp. n., from which it differs
especially in the larger size and wing-venation. The structure of the male hypopygium is very similar in the two
species.
Antennae shorter, if bent backward scarcely attaining thewing-root ; scapal segments ochreous, the flagcllar segments
dark brown, oval. Praescutum narrowly pruinose laterally ;scutal lobes each with two darker dashes, the median areaslightly greyish. Pleura reddish brown, sparsely pruinose.
HMtcres pale, the knobs more infuscated. Legs with thefemora pale, darkened toward the tip ; tibiae yellow, the tips
and a conspicuous, dilated, subbasal ring on fore tibia ( ~ )darkened; tarsi pale, passing into brown at tips. Wings
rather broad, with a pale yellowish ti,~ge, the veins darker
yellow; maerotrichae pale brown. Venation: r lying
proximad of r-m, the basal section of R~+s very short andstrongly arcuated ; vein 2nd A long, ending nearly opposite
the cephalic end of the long oblique m-cu. Abdomen darkbrown, tim hypopygium a trifle lighter in colour, Male
hypopygium very similar in colour to that of M. eziguus,
differing only in minor details. Basal dististyle longer andmore slender, the apical spine longer, more slender, and gently
curved at tip, the lateral spine ercc b placed at about twothirds the length of the style.
Hab. Tasmania.

new or little-known Tipulidm.

181

Holotype, ¢), Lake Margaret, altitude 2500 feet, February 3,
1923 (d. ~bnnoir).
The most obvious differences between the present species
and ~[. exiguug lie in the sborter antennae, the venation,
especially the longer vein 2nd A and the long oblique m-cu,
and the hypopygium.
Molophilus apricus, sp. n.
Belongs to the graeilis group, ruficollis subgroup ; generalcoloration dark brown; antennm short, dark throughout;anterior lateral pretergites conspicuously light yellow;
halteres infuseated, the k,obs conspicuously light yellow;
wings with a strong brownish tinge, the veins darker brown ;male hypopygium with the basistyle short and stout, theventral lobe broad and flattened, the dististyles lying at its
base, both heavily blackened, the basal style terminating iu
an acute sickle-shaped point.
Male.--Length about 4,'5-4"6 mm. ; wing 5'5-6 ram.
Female.--Length about 5 ram. ; wing about 5"5 ram.
Rostrum brown, the palpi darker brown. Antennm short,
if bent backward not attaining the wing-root, brownish black
throughout; flagellar segments short-oval to short-cylindrical, in the male appearing ahnost moniliform. Head darkbrownish grey, with relatively short yellow set,e; a more orless distinct ochreous spot behind each antennal fossa.
Pronotum obscure yellow. Anterior lateral pretergites
clear light yellow, the posterior pretergites obscure. Mesonotum dark grey, the prmscutum with a narrow dark brown
median stripe that is slightly narrowed behind; humeral
region strongly tinged with reddish ; scutellum with a reddish cast. Pleura dark leaden-brown, including the dorsopleural membrane ; tegular region restrictedly yellow.
Halteres infu~cated, the knob~ abruptly and conspicuously
light yellow. Legs with the cox~e leaden-brown, the forecoxm a little paler ; trochanters dark brown; remainder oflegs dark brown, the terminal tarsal segments passing into
black. Wings with a strong brown tinge, the veins conspicuously darker brown. Venation : r lying some distancebeyond the level of r - m ; m-cu unusually long, sinuous,
longer than the short petiole of cell M~ ; vein 2nd A gently
sinuous, extending to just beyond the cephalic end of m-cu.
Abdomen dark brown, including the hypopygium, thelatter with conspicuous fringes of yellow setm on the tergum
and ventral lobe of basistyle. Male hypopygium with thebasistyle very short and stout, the ventral lobe relatively
short and wide, flattened~ the apex obtuse and fringed with

182

Dr. C. P. Alexander onyellow setm. The dististyles lie at the base of this ventral
lobe, appearing unusually open and exposed; outer style asmall, arcuated, black rod, the apex shallowly bifid ; innerdististyle longer, the base stouter and nearly straight, theapical half or less more slender, sickle-shaped, narrowed tothe acute apex ; between these two styli lies a small, acute,
tooth-like spine. In a position of rest~ the sickle-shapedends of the basal dististyles lie decussate across the genitalchamber.
In the female that is associated with this species~ thepetiole of cell M3 is somewhat longer and there are otherslight differences from the male.
Hab. Tasmania.
Holotype, c~, Strahan, February 5, 1923 (A. Tonnoir).
Allotype, ~, Adventure Bay, January 1, 1923 (A. Tonnoir).
Paratopotype, c~, February 1924 ( G. H. Hardy), collector's
number 399; paratype, ~ , Geeveston, December 8, 1922(.4. Tonnoir).
The paratopotype is in the collection of the University of
Queensland.

Molophilus cerberus, sp. n.
Belongs to the gracilis group, ruficollis subgroup ; generalcoloration dark brown; antennm relatively short; legs andwings brown; male hypopygium with the outer dististyle
deeply bifid, the outer arm long, slender, sinuous, approximately four times the length of the short straight innerarlTl.

Male.--Length 4"5 mm. ; wing 5"4-5'8 mm.
Female.--Length about 5"5-5"7 mm. ; wing 7"7-7"8 ram.
Rostrum and palpi brownish black. Antennm relatively
short, if bent backward (c~) not attaining the wing-root,
brownish black throughout ; flagellar segments oval, with ashort pale pubescence and relatively short verticils. In thefemale the antennm are a little shorter with the basal segments a trifle paler. Head dark grey, with numerous shortyellow setm.
Pronotum dark brownish grey. Lateral pretergites conspicuously light sulphur-yellow, narrowly interrupted justbeyond the pseudosutural fove~e, becoming more conspicuousat the tegular region. Mesonotum dark plumbeous brown.
Pleura similar, the dorso-pleural membranes conspicuously
light yellow. Halteres pale, the knobs light yellow. Legs
with the coxm concolorous with the pleura; trochantersbrown ; remainder of legs dark brown ; fore tibiae ( c~) with

new or little-known Tipulidze.

183

a slightly enlarged sub-basal ring. In the female the femora
and tibiae are more yellowish, the terminal tarsal segments
passing into dark brown. Wings with a strong brown tinge,
the longitudinal veins more or less seamed with darker; veinsslightly darker brown, especially R 5 ; macrotrichi~e darker
brown. Venation: r lying approximately opposite r - m ;vein 2nd A relatively long, ending opposite one-half to onethird the length of the petiole of cell M3. In the female,vein 2rid A is even longer, in casea extending almost to thebase of cell 21/3.
Abdomen brownish black, with conspicuous yellow setae ;hypopygium dark, with long yellow setae. Male hypopygium
relatively short and stout, the basistyle with the ventral lobe
produced into a stout finger-like appendage. Dististyles
somewhat approximated at base, the outer style with a long
slender stem, bifid at apex, the outer arm long, slender,strongly curved to the acute tip ; inner arm a short bluntspur that is only about one-fourth the length of the outerarm. Basal dististyle a long, slender, gently curved rod, thetip acute, the ventral margin with microscopic appressed
spinules, the base of the style more dilated. _/Edeagus relatively short. Ovipositor with the valves very long and slender,strongly upcurved to the acute tips; sternal valves long andslender, straight.
Hub. Tasmania.
Holotype, c~, Cradle Valley, January 11, 1923 (A. Tonnoir).
Allotopotype, ? ~ January 16, 1923 (A. Tonnoir).
t)aratopotypes, 3 ~ ~ , 1 ?~ January 16-27, 1923 (,4.
Tonnoir) .

Molophilus erebus, sp. n.
Belongs to the gracilis group, ruficollis subgroup ; generalcoloration dark brown ; antennae of male of moderate length ;wings with a strong brown tinge, broadly washed with paler
before the cord ; petiole of cell M 3 short ; male hypopygiumwith the ventral lobe of basistyle slender, tapering; basal
dististyle a simple sinuous rod, the apex dilated into a
blade.
Male.--Length about 4"5-5 ram. ; wing 5"5-6"5 ram.
Female.--Length about 5'5 mm. ; wiug 7"3 mm.
Rostrmn and pa|pi dark brown. Antennae ( ~ ) relatively
elongate, if bent backward extending to about opposite thesecond abdominal segment ; flagellar segments elongate-oval
to fusiform, with a conspicuous, erect, white pubescence andsubequal unilaterally arranged black verticils. In the female

184

Dr. C. P. Alexander on

the antennte are shorter, but still extend to beyond the wingroot. Head dark brownish grey.
Pronotum obscure brownish yellow, the anterior lateralpretergites narrowly obscure yellow. Mesonotum darkbrown, the humeral region of the prmscutum restrictedly
obscure yellow; pseudo-sutural fovem black. Pleura darkbrown, the dorso-pleural membrane a little paler. Haltereswith the extreme base and knobs light yellow, the remainderof the stem weakly infuscated. Legs with the cox~e weakly
infuscated, especially the posterior coxm; trochauters obscureyellow; femora yellowish brown, the tips narrowly infuscated ; tibim light brown, the tips narrowly infuscated ; terminal tarsal segments passing into darker brown ; hind tarsi
shorter than the tibiae. Wings with a strong brown tinge,
with conspicuous paler washes before the cord and in cells
Cu and 1st A ; veins darker than the ground-colour, especiallyR~ and Cul ; macrotrichi~e dark brown. Venation : r lying
some distance beyond the level of r-m ; basal section of R~+~nearly twice as long as R s ; petiole of cell M3 short, about
equal to or a little longer than tim weakly sinuous m-cu ;vein 2rid A relatively elongate, ending about opposite midlength of the petiole of cell M3.
Abdomen dark brown, including the hypopygium. Male
hypopygium with the ventral lobe of basistyle very long,
slender, tapering gradually to the narrowly obtuse apex, themesal face with abundant delicate erect to retrorse setulm ;dorsal lobe of basistyle broadly expanded. Outer dististyleslender, bifid at apex, the outer arm a narrow, flattened,
curved blade, the inner arm a small dilated blade. Basal
dististyle very conspicuous, simple, the basal half or more
slender, sinuous, then dilated into a broad, flattened, curved
blade, the apex gradually narrowed to an acute point, theouter margin of the blade microscopically serrulate ; face of
blade covered with abundant microscopical setul~e.
Ovipositor with the bases darkened, the tergal valves yellowish,
very elongate.
Hab. Tasmania.
Holotype, c~ , Zeehan, February 7, 1923 (A. Tonnoir).
Allotype, ~, Cradle Valley, January 12, 1923 (.4. Tonnoir).
Paratypes, 1 ~', 1 ? , with the allotype, January 12-26,
1923 ( A. Tonnoir).

Molophilus lyratus, sp. n.
Belongs to the gracilis group, ruficollis subgroup ; antennm
short ; head pale brownish yellow, the anterior vertex almost
fulvous ; knobs of halteres yellow; fore tibim ( ~ ) with a~ew or little-known Tipulidae.

185

conspicuous subbasal black ring; male hypopygium witha conspicuous lyrifbrm structure, the arms protruding between
tile large ninth tergite and the long narrow basistyles.
M a l e . - - L e n g t h about 5 ram. ; wing 6 ram.
Rostrum pale brown; palpi dark brown. Antennm short,
if bent backward extending to just beyond tile wing-root ;scapal segments ochreous ; flagellar segments brown, passing
into darker brown outwardly; flagellar segments oval,
clothed with a very short inconspicuous pubescence andlonger conspicuous verticillate setm that are a little longer
than the segments themselves. Head pale brownish yellow,
the anterior vertex almost fulvous, the orbits narrowly paleyellow.
Pronotum yellow, the lateral pretergites paler yellow, thiscolour becoming obsolete near the level of the pseudosutural
foveae. Mesonotal praescutum yellowish grey, with threebroad, subconfluent~ brown stripes, restricting the colour tothe interspaees and laterally; humeral region vaguelybrightened ; scutum brown, the "lobes darker laterally andposteriorly, the posterior lateral region of each produced intoa small yellow tubercle to the wing-base ; scutellum vaguely
more yellow; postnotal mediotergite brown, the lateral
margins with a narrow strip of yellow on either side. Pleura
brown, the sterno-pleurite and meron more reddish brown,
the anepisternum darker. Halteres pale, the knobs yellow.
Legs with the coxae and trochanters reddish brown ; femora
yellow, the tips infuseated; tibiae brownish yellow, thetips infuscated ; fore tibia ( 8 ) with a conspicuous, feebly
dilated, subbasal, black ring that is a trifle more thanits own length beyond the base ; basitarsi pale brown, theremainder of the tarsi darker brown. Wings with a strong
yellowish tinge, the veins darker yellow ; macrotrichiae pale
brown, only moderately long and dense. Venation: vein
2 n d A moderately long, extending to a short distance beyond
the base of the petiole of cell M~.
Male hypopygium relatively long and narrow, of very
unusual structure. Ninth tergite large, arched, narrowed
epically, appearing as a conical plate with the apex narrowly
truncated. From beneath this plate arises a remarkablelyriform chitinized structure that presumably represents aphallosome, the two arms slender, narrowed gradually to theacute tips which jut caudad from between the tergal plate
and the elongate basistyles. Basistyles ventral, slender, lying
parallel and closely approximated, the apex of each obtuse,
unarmed, provided with rather abundant delicate yellow setae.
Outer dististyle relatively small, the apex slender blackened

186

Dr. C. P. Alexander ona subbasal more flattened and obtuse arm of approximately
equal length. Basal dististyle a nearly straight powerful
black rod, the extreme apex a little enlarged and provided
with two or three obtuse teeth on the lower or inner face ;lower face of style with a conspicuous sub-basal spine directed
toward the apex of the style, the axil densely set with black
spinous pegs that extend in a row along the lower face of thestyle ahnost to the tip. zEdeagus slender, extending caudad
to opposite the tips of the lyre.
Hab. Tasmania.
Holotype, (~ , Hobart, January 13, 1923 (A. Tonnoir).

Molophilus perdistinctus, sp. n.
Belongs to the plagiatus group; general coloration darkbrown; antennae of male elongate, the flagellar segments
only moderately enlarged at base ; wings tinged with brown ;male hypopygium with the basal dististyle small, slender,
bent into a boomerang-shaped structure ; gonapophyses fused
into a blackened central mass that terminates in two strong
horns.
Male.--Length about 4 mm. ; wing 5 ram.
JFemale.--Length about 4"5 mm. ; wing 5"2 ram.
Rostrum and palpi dark brown. Antennae ( ~ ) elongate,
in the unique type broken at the eighth segment, if entire
presumably extending to opposite mid-length of the abdomen ;flagellar segments fusiform, the base of each segment only
feebly enlarged, gradually narrowed to the short, shiny,terminal pedicel; antennm dark brown throughout, with aconspicuous erect white pubescence and unilaterally arranged
verticils. Head dark brown.
Thorax dark brown, the humeral region of the przescutum
only narrowly and indistinctly brightened. Halteres dark,
the knobs brown. Legs with tile eoxm yellow, the fore cox~emore infuscated; trochanters yellow; remainder of legs
brownish yellow, the tarsi passing into brown, the segmentswith brown macrotrichi~e, some of which are suberect. Wings
with a uniform brown tinge, the veins darker brown ; veins
R, Rs, and Cul darker; macrotrichi~e darker brown. Venation: r lying a short distance beyond r-m; vein 2nd A
extending to shortly beyond the origin of the petiole of cellMs, the veins only slightly sinuous and closely parallelingthe anal margin of the wing.
Abdomen dark brown, including the hypopygium. ~-~ale
hypopygium with the basistyles produced apically intoa small tubercle that is filrther produced into a small
chitinized beak, the whole structure suggestive of the headnew or fettle.known Tipulid~.

187

and beak of a bird, the tubercle bezel with delicate setm.
Dististyles widely separated at origin by membrane; outerdististyle a powerful blackened rod, gently curved, the apex
terminating in two points, the outer a short straight arm,
the inner a little longer, gently curved. Basal disti~tyle very
slender, small, shaped somewhat like a boomerang, the headat near one-third the length, the distal two-thirds a little
enlarged, the apex blunt, the surface of the style with a few
microscopic punctures. Gonapophyses fused into a massive
blackened central mass, the caudo-lateral angles produced
into conspicuous hort~s; in the unique ~ype one hor~a isgently curved, the other directed caudad. -?Edeagus verysmall. Ninth tergite an oval setiferous lobe. Ovipositor
with the tergal valves slender, gently upcurved to the acutetips.
Hab. Tasmania.
Hololype, c~, Cradle Mt., January 27, 1923 (.4. Tonnoir).
Allolopotype, ?.

Molophilus inelegans, sp. n.
Belongs to the gracilis group, ruficollis subgroup ; generalcoloration dark brown, the pleura conspicuously dark grey ;halteres obscure yellow ; legs light yellowish brown, the tipsof the tarsi darker; wings with R4+~ short; mate hvpopygium with the gonapophyses and ~edeagus very long°and
slender.
Male.--Length about 3"8 ram. ; wing 4"7 ram.
Rostrum and pulp; dark brown. Autennm relatively short,
if bent backward scarcely reaching the wing-root; scapal
segments light brown, the flagellum darker brown ; flagellar
segments oval, with a dense erect pale pubescence. Headdark hrownish grey.
Pronotum dark. Lateral pretergites pale whitish yellow,
becoming darkened beyond the level of the pseudosutural
fovem. 1Y[esonotumuniformly dark brown, the setm of theprmscutum small and inconspicuous; pseudosutural fove~eelongate, shiny reddish brown; humeral region a little
tinged with reddish; scutellum more testaeeous brown;
postnotum dark grey. Pleura uniformly dark grey, thedorso-pleural membrane dark. Halteres obscure yellow.
Legs with the coxm and trochanters obscure yellow ; femora
light yellowish brown, with relatively sparse and inconspicuous darker brown triehim ; tibiae yellowish brown, thetips narrowly darker ; fore tibiae (d') with the basal region
very slightly dilated~ but not darkened ; tarsi passing into
dark brown. Wings slightly greyish, the veins brown;

188

Dr. C. P. Alexander onmacrotrichim darker brown. Venation: r lying almost
opposite or just beyond r-m, the basal section of R2+ashort,arcuated ; R~+~ very short, not much longer than r-m;
vein 2nd A ending opposite the base of the petiole ofcell Ms.
Abdomen dark brown, the hypopygium a little paler,
especially the tips of the styli, which appear bleached.
Basistyles relatively stout, the base with relatively small
setm, the apex rather abruptly narrowed into a pale flattened
lobe with more numerous conspicuous set~e. Outer dististyledusky, a slender rod, the basal half arched into a halfcylinder, the apex gradually narrowed to the subacute tip.
lnner dististyle with the stem strongly curved, the apex
dilated into a conspicuous pate flattened blade that bears two
spines, a larger one near the base on the lower margin, anda small straight spine placed more distally on the upper
margin; disk of the blade with'about fifteen delicate setiferous punctures. Gonapophyscs very powerful, appearing
as slender curved horns, the tips more blackened, acute.
2Edeagus exceedingly elongate, slender, arcuated, in a positionof rest jutting beyond the other elements of the hypopygium,
shortly before its apex a little dilated.
Hab. New South Wales.
Holotype, c~ , Waterfall, November 1921 (A. Tonnoir).
M. inelegans is allied to M.fro#gatti, Skuse, and M. gigas,
Alexander.

MotophiluspolioceThalus, sp. n.
Belongs to the gracilis group, ruficollis subgroup ; generalcoloration reddish brown, the head brown with a conspicuous
blue-grey area on centre of vertex; wings greyish yellow,
the base darkened ; male hypopygium with the inner dlstistyle terminating in a long spine with a small acute spine in
its axit.
Male.--Length about 3 ram. ; wing 3"5 ram.
l#emale.--Length about 3"5 ram. ; wing 3"7 ram.
Rostrum and palpi brownish black. Antennm of malemoderately elongate, if bent backward extending to justbefore the base of the abdomen ; first scapal segment dark,
the second scapal and basal flagellar segments yellow,
the terminal segments passing into pale brown ; flage]larsegments (c~) elongate-fusiform, with a conspicuous erect
white pubescence and with unilaterally arranged verticils.
In the female, the antennm are shorter, pale almost to theend. ttead brown, the vertex with a conspicuous blue-greyarea.

new or tittle-known Tipulidae.

189

Mesonotum pale to dark reddish brown, the lateral marginof the praescutum narrowly yellowisb, the posterior scleritesof the mesonotum slightly pruinoue. Pleura dark brown.
Halteres pale, the knobs testaceous yellow. Legs with thecoxm brownish testaceous ; trochanters yellow ; remainder oflegs pale brown, the apices of the tibiae narrowly darkened,
the terminal tarsal segments darker. Wings with a greyishyellow tinge, the costal region clearer yellow, the base conspicuously darkened, this eolour including the basal third ofcell 2nd A ; veins darker yellow than the ground-colour.
Macrotrichiae relatively dense, brown. Venation: r lying
beyond the level of r-m, the basal section of R~+a thus beingnearly one-half longer than R4+~; vein 2nd A terminating
about opposite the caudal end of m-cu.
Abdominal tergites dark brown, the sternites more reddish
brown, the caudal margins of the segments narrowly paler,more evident in the male ; in the female, the tergites with amore or less distinct longitudinal dark median line. Male
hypopygium with the basistyles relatively stout, the ventral
lobe prolonged caudad beyond the level of the dististyles,
ap.pearing flattened to subclavate, the mesal face with conspmuous erect setm and abundant microscopic setulae.
Dististyles placed close together in the notch at the base of
the lobe of the basistyle ; outer dististyle a massive, subquadrate, fleshy lobe that has two rows of setiferous tubercles
along the mesal face, each of these rows terminating in a
powerful spine ; the entire surface of this style is closely set
with microscopic setulm, in addition to the larger setae.
Inner dististyle longer, the base dilated, a little less than theapical half bent at a right angle and gradually narrowed intoa long chitinized point, the outer margin near the bend witha few microscopic tubercles; in the axil of the bend asmaller, acute, black spine. _/:Edeagus relatively slender andelongate. Ovipositor with the tergal valves nearly straight.
Hub. New South Wales.
Holotype, ~ , Dorrigo, 1922 (Walter Heron).
AUotopotype, ~.
Type in the writer's collection.
Molophilus capitalus~ sp. n.

Closely allied to M. 2oliocephalus, sp. n., differing chiefly
in the structure of the male hypopygium.
Male.--Length about 3"8 mm. ; wing 4"5 ram.
l:Lostrum and palpi brownish black. Antennae with thescapal segments dusky at base, tim tips broadly light yellow,

190

Dr. C. P. Alexander on

the flagellum brownish black; flagellar segments elongateoval, with a conspicuous erect white pubescence and unilaterally arranged black verticils. Head light yello% the centreof the vertex with a conspicuous blue-grey area.
Pronotum light yellow, with a small dorso-median brown
spot, the lateral margins infuscated. Lateral pretergites
narrowly whitish yellow. Mesonotal prmscutum rather lightbrown, the margins paler; scu~cllum aud postnotam moretestaeeous brown ; thoracic set~e relatively sparse and short.
Pleura light brown. Hatteres pale, tim knobs light yellow, tile
stem a little infllscated. Legs with the coxm and trochanters
yellowish testaceous, the posterior coxm brighter; femora
brown, the bases broadly paler ; tibiae pale brown, the tips
darkened; fore tibia ( ~ ) with a relatively narrow, shiny,
black sub-basal ring; tarsi passing into brownish black.
Wings with a faint yellowish-brown tinge, the base notconspicuously darker ; veins and macrotrichim darker brown.
Venation: r almost opposite r-m; vein 2nd A ending
opposite m-ca.
Abdomen brown, the hypopygium more yellowish. Male
hypopygium with the ventral lobe of the basistyle armed
with longer stouter set~e. Outer dististyle similar to thatin M. poliocep]~alus, the outer tooth simple, the lower ormore basal tooth flattened into a serrulate plate ; a curious,
flattened, ribbon-like structure appears to arise from thisstyle, directed caudad, the tip a small spine, before the apex
on either side with small serrations. Inner dististyle withthe axillary spine smaller. The long ribbon-like appendage
on the outer dististyle of the male hypopygium does notoccur in poliocephalus, being replaced by a microscopic spur.
Hub. South Australia, New South Wales.
Holotype, ~, Adelaide, South Australia, October 20, 1921

( A. Tonnoir).
Paratype, (~, Woy Woy, :New South Wales, September 2~
1923 (I. Mackerras); ~pecimen returned to Dr. :E. W.
Ferguson.

Molophilus tortilis, sp. n.
Belongs to the gracilis group, ruficollis subgroup ; closely
allied to M. poliocephMus, sp. n. ; head cream-yellow, thecentre of the vertex with a large oval grey spot; male hypopygium with the ventral lobe of the basistyle with very
long setm; inner dististyle forked to mid-length, the arm~sinuous.
Ma/e.--Length about 3"8 mm. ; wing 4r7 ram.
Rostrum and palpi brownish black. Anteaum dark brown~new or little-k~wwn Tipulidm.

191

the flagellum broken. Head light cream-yellow, the centreof the vertex with a large oval grey spot.
Pronotum light sulphur-yellow. Mesonotum reddish
brown, tim pr~escutum with a faint bluish cast ; pseudozutural
foveae dark reddish brown, but little evident against the
ground-eolour; lateral pretergites light sulphur-yellow;
seutellum a trifle more testaceous. Pleura dark brown.
Halteres dusky, the knobs infuscated. Legs with the coxm
and trochanters testaceous ; remainder of legs dark brown,
the fore legs broken. Wings with a faint dusky tinge, thevehas a little darker, the macrotriehim dark brown. Venation :vein 2ud A eading opposite or a very short distance beyond?Yt--CU.

Abdomeu dark brown, the hypopygium more yellowish.
Male hypopygium with the ventral lobe of the basistyle welldeveloped, slender, the mesal lace with very long, powerful,erect setm, these only a little shorter than the lobe itself ;other lobes of basistyle poorly developed. Outer dististylea short, powerful, ciavate lobe, entirely darkened, the basenarrow, widened to a collar betbre the apex, the conical tip
suddenly produced, sparsely setiferous ; tiJc collar-like ringis produced on the outer margin into a black, gently curved,
spiny lobe. Inner dististvle elongate, very deeply forked,
the longesL arm about as l~ong as the stem, strongly twisted
at base, thence extended into a long straight spine; innerarm shorter than the stem, the apex blackened, the extreme
tip suddenly curved, acute. 2Edeagus relatively long and~lender.
Hab. New Soutb'Wales.
Holotype, d', Balmoral, near Sydney, May 19, 1923
(E. ¢V. Ferguson ) ; Collector's No. 80.
Type returned to Dr. Ferguson.

Molophilus sigma, sp. n.
Belongs to thegracilis group, ruficollis subgroup ; generalcoloration light reddish brown ; basal segments of antennm
yellow;head yellow, the centre of the vertex with a small
grey spot ; ventral pleurites dark brown ; knobs of haltereslight yellow ; male hypopygium with the outer dististyle auoval, densely setiferous lobe, with a powerful, curved, blackspine at base; basal dististyle a long, simple, strongly
sinuous rod, extended at apex into a blackened spine.
Male.--Length about 3"5 mm.
Rostrmn and palpi brownish black. Antennm moderately
elougate~ if bent backward extending to ~hortly beyond the192

Dr. C. P. Alexander onwing-root; scapal segments light yellow, flagellum pale
brown ; flagellar segments long-oval, with a dense erect pale
pubescence and unilaterally arranged setm. Head largely
yellow~ the centre of the vertex with a prainose area ; occiput
darkened ; genm infuscated.
Pronotum light yellow, the propleura blackened. Lateralpretergites yellowish testaceous, becoming narrowed toward
the wing-root ; tegula light yellow. Mesonotum lighi reddish brown, with a vague bluish east, the anterior portion of
the pr~scutum brighter, the posterior prmscutum and thescutum narrowly darkened laterally; pseudosutural fovem
pale ; setm of mesonotum short and relatively inconspicuous ;scntellum and posterior portion of postnotal mediotergite
somewhat brightened. Pleura dark brown ventrally, thedorso-pleural membranes, pteropleurite, and pleuro-tergite
remaining light reddish brown. Halteres pale, the extreme
base of the stem infuseated, the knobs conspicuously lightyellow. Legs with the cox~e and trochanters yellowish testaceous, the fore cox~e darker; of the remaining legs, only asingle incompIete posterior leg remains--this obscure yellow,
the extreme tip of the tibia infuscated. Wings of the uniquetype broken.
Abdomen dark brown, the hypopygium more yellowish.
Male hypopygium of very unusual structure. ]3asistyles
relatively stout, the ventral lobe produced into a shortclavate structure that is provided with microscopic spinuke
and a dense patch of long, powerful, rctrorse setze. The
dististyles arise from the base of this lobe; outer styleterminating in an oval, densely setuliferdus lobe, at the baseof which is a powerful, black, sickle-shaped spine. Basal
dististyle a long, simple, strongly sinuous rod that terminatesm a long, acute, black spine ; before tim apex of the style,provided with numerous microscopic sctul~e. 2Edeagus
relatively long and stout, a little shorter than the basal
dististyle.
Hub. Victoria.
Holotype, ~, Sassafras, Dandenong Range, altitttde 1000
feet, October 21, 1922 (A. ;lbnnoir).

Molophilus rasilis, sp. n.
Belongs to the graeilis group, rufieollis subgroup ; generalcoloration shiny reddish brown ; wings with a greyish-yellow
tinge, the base and costal region more saturated yellow;
abdomen dark brown; male hypopygium with the outerdististyle deeply and unequally bifid, the basal dististylewith two long lateral spines, the space between them withlong erect set~e.

new or tit~le-k~wwn Tipulidm.

193

M a l e . - - L e n g t h about 3"8 ram. ; wing 4"5 ram.
Rostrum and palpi dark brown. Autennm relatively short,
if bent backwards scarcely attaining the wing-root, darkbrown throughout ; flagellar segments oval, with conspicuous
verticils and a short but dense erect white pubescence.
Head dark brown.
Pronotum dark brown. ]Y[esonotumshiny reddish brown,
the margins paler; lateral pretergites narrowly yellowish ;seutum reddish brown, tile caudal margin narrowly yellowish; scuteilum brownish testaceous ; postnotum shiny
brown. :Pleura shiny reddish brown. Haltcres with thestem pale brown, yellow at base, the knobs obscure brownish
yellow, with dark set~e. Legs with the coxm testaceous
yellow; trochanters yellow; femora yellow, the tips narrowly
darkened, more extensively so on the fore legs; tibiabrownish yellow, the fore tibia (c~) darkened but scarcely
dilated at base; tarsi dark brown, the basitarsi somewhat
paler at base. Wings with a greyish-yellow tinge, the baseand costal region more yellowish ; veins yellowish brown,with long, pale brown macrotrichim. Vcnatiou: r lying shortly
beyond the level of r - m ; vein 2rid A moderately elongate,
ending about opposite the caudal end of m-cu.
Abdomen dark brown, the hypopygium a little moreyellowish. Male hypopygium with tim ventral lobe of the{)as[style produced iuto a long slender lobe, the mesa[ facewith abundant setulm and fewer scattered erect setm that
tend to become retrorse at tim apex ; outer face of the lobe
with fewer scattered large setm; dorso-mesal angle of"basistyle produced into a low rounded lobe. :Dististyles
placed close together in the notch between the lobes of thebasistyle ; outer dististyle with a short stout stem, the arms
slender, especially the outer arm which is more than twice
the length of the stem, sinuous, gradually narrowed to theslightly spatulate apex ; inner arm much shorter, about equalin length to the stem, gently arcuated. Basal dististyle apowerful blackened rod, terminating in a slender curved
spine, with a similar straight spine on the face at about twothir(ts the length of the style, the area between these spines
with numerous very long erect setm that are approximately
as long as the spines. JEdeagus long and sleuder~ taperingto the slender apex, in balsam mounts extending caudad tonearly opposite mid-length of the ventral lobe o[ thehas[style.
[Iab. Tasmania.
Hololype, c~, Wilmot, January 8, 1923 (.4. ~bnnoir).
A m , & Mag. 2i. [li,~t. Ser. 9. Vol. xix.
13

194:

Dr. C. P. Alexander on

Molophilus verticalis, sp. n.
Belongs to tile verticalis group ; general coloration reddish
brown to dark brown, the pronotum and lateral pretergites
conspicuously yellow ; head yellow, the centre of the vertex
grey ; halteres yellow ; wings yellow, the veins and macrotrictfim darker; male hypopygium with the dorsal lobe of
the basistyle produced caudad into a stout lobe that terminates in a cbitinized spine ; outer dististyle profoundly bifid,
each arm needle-like.
M a l e . - - L e n g t h 3"8-4"5 mm. ; wing 4"7'-6 ram.
Female.--Length 4"5-4"6 ram. ; wing 6-6"2 mm.
Rostrum and palpi black. Antennm of moderate length,
if bent backward ( g ) extending about to the wing-root;
scape and basal segments of flagellum yellow, the organpassing into brown toward outer end; flagellar segments
long-oval with an erect white pubescence and unilaterallyarranged verticils. Head ochre, the centre of the vertex
largely grey. In the Mr. Field paratype, the head is
unifbrmly grey.
Pronotum yellow, more infuscated laterally. Lateral pretergites light sulphur-yellow, extending caudad to the concolorous tegula. Mesonotal reddish brown to dark brown,with a fait~t grey cast ; pseudosutural fovem light chestnutbrown, inconspicuous; scutellum yellowish to testaceous;postnotum dark brown, the mediotergite with a yellow spoton either side. Pleura dark reddish brown, in some specimens distinctly pruinose ; dorsal portion of the postnotal
pleurotergite paler; dorso-pleural membrane more or lessyellowish. Halteres yellow. Legs with the fore coxm
brown, the other eoxm and the trochanters yellow ; femora
obscure yellow, the tips vaguely darkened ; tibim light brown,
the tips narrowly darker, fore tibia ((~) with a very narrow,dark brown, subbasal ring ; tarsi passing into dark brown.
Wings with a strong yellowish tinge, the veins and macrotrichim darker yellow. Venation : r lying shortly beyondr - m ; m-cu relatively short and straight, oblique; vein
2nd A ending opposite the proximal end of the petiole ofcell M~.
Abdomen clothed with conspicuous yellow setm, thetergites obscure yellow, with an interrupted darker dorsomedian stripe. In a few specimens the abdomen is uniformly
brown, the hypopygium usually more reddish brown. Male
hypopygittm with the basistyles relatively slender, the dorsal
portion produced into a stout lobe, the outer lateral angle ofwhich is further produced into a slender, sinuous, chitinizednew or llttle-known Tipulicke.

195

blade or point, the style at base of this blade a little elevatedand with a group of about eight powerful set~e. Mesal faceof basistyle with a small lobe that is provided with strong
spinous setm. Outer dististyle profoundly bifid, each arm
long and slender, needle-like, the outer arm longer, a little
dilated before the acute blackened apex; inner arm more
slender, a long~ feebly sinuous, acicular black spine. Innerdististyle an elongate blackened rod, a little dilated before
the slightly curved spinulose apex; the dilated portion of
the style with a small tooth-like or wing-like expansion.
~]~d~gus long and slender, straight.
Hab. Tasmania.
Holotype, (~, Fern Tree, Mt. Wellington, November 11,

1922 (A. Tonnoir).
Allotopotype, 9.
Paratopotype, 1 c~; paratypas, 1 ~ , Burnie~ January 31,
1923 ; 2 ~' ~ ~ St. Patrick's River, October 30-November 4,
1922 ; 1 c~ ~ 1 ~, Cradle Valley, January 10-27, 1923 ; I ~',Mt. Field, December 19, 1922 ; 2 ~ ~ , l~ational Park,
December 16, 1922; 1 ~', 1 ~, Eaglehawk Neck, Tasman:Peninsula, November 18, 1922; 5 ~' ~', Mt. Wellington,
November 21-30, 1922; 1 ~ , Hartz M t s , December 10,
1922 (A. Tonnoir).

Molophilus verticalis reductus, subsp, n.
In the mountainous section of North-western Tasmania
there occurs a large form of verticalis that is of greatinterest. Tim subspecies varies almost as widely in its
general coloration as does the typical form, tending to havethe thorax and abdomen more uniformly darker brown.
The male hypopygium differs from the typical form inseveral regards. The dorsal lobe of the basistyle is much
more slender and is not produced into a shoulder at the baseof the narrowed apical portion, the few long setm being
placed along the mesal edge; the lobe narrows gradually tothe very short chitinized apex. Outer dististyle with theouter arm dilated on basal half and here microscopicallyspinulose, the apical half a long straight slender spine ; innerarm stouter and more sinuous. Inner dististyle more slenderapically, without the conspicuous wing before the microscopically spinulose apex.
Holotype, ~, Cradle Valley, January 12, 1923 (A.
Tonnoir).
Paratopotypes, 3 c~ c~, January 24-27, 1923 (A. Tonnoir).
13 ~

196

On new or little-known Tiputid~.

Molophilus furvus, sp. n.
Belongs to the plagiatus group ; general coloration dark
blackish grey; haiteres with sulplmr-yellow knobs; wings
with a strong brownish tinge ; vein 2 n d A short; male hypopygium with the basistyles elongate; basal dististyle aslender rod that is bent strongly ventrad at tile apex intoa blackened spine.
Jllale.--Length about 2"4 ram. ; wing 3"2 mm.
Female.--Length about 2"7 mm. ; wing 3"5 ram.
Rostrum and palpi black. Antennm (c~) with the scapedark brown, the flagellum brownish black; moderately
elongate, in d' if bent backward extending about to the baseof tile abdomen; flagetlar segments fusiform, with conspicuous verticillate hairs. Head dark grey.
Pronotum dark brown. Mesonotum dark blackish grey,
the humeral region of the prmscutum vaguely more reddish.
:Pleura dark grey, the dorso-pleural membrane dark. Halteres pale, the knobs light sulphur-yellow. Legs with thecoxm and trochanters dark brown; remainder of legs
brownish black, the femoral bases a little paler. Wings witha strong brownish tinge, the centres of the cells beyond
the cord somewhat paler ; veins darker brown ; macrotrichim
dark brown. VenaLion : r opposite to just beyond the levelof r-m ; m-cu long and gently arcuated ; vein 2nd A short,a trifle longer in the female, in the male ending abou~ onethird its length before the level of the caudal end of m-cu.
Abdomen dark brown, including the hypopygium. Male
hypopygium with the basistyles very elongate, the apex ofeach with a stout beak surrounded by numerous setm.
Dististyles rather widely separated at origin; outer style
stout, shallowly bifid at apex, the outer arm longer flattened,
the tip obliquely truncated; inner arm small. Basal dististyle a very long and slender, chitinized rod, the distal third
sizmous and blackeHed, narrowed to the acute tip; in aposition of rest, these styli lie on the dorsal face of thebasistyle, directed caudad, the long acute apex vet~trad.
2Edeagus long, slender, straight. Ovipositor with the basal
shields dark, the valves horn-coloured ; dorsal valves very
slender and acute, gently upcurved ; sternal valves a little
longer, nearly straight.
Hab. New South Wales.
Holotype, ~ , Mt. Wilson, altitude 3475 feet, November 19,]921 (,4. "lbnnoir).
Allotopotype, ?, in copula and pinned with type.
Paraloyolype, c~.