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I

UNDESCRIBED TIPULIDAE
FROM TEE ETEIOPIAN REGION
By Charles P. ALEXANDER
Amherst, Nlassachusetts,
U. S. A.
The nelv speciesof crane-fliesdiscusserlherewith \Merereceivetl flom various soulces. An important seriesfrcm CapeColony,
now prLrservedin tlre Sou"l,hAfrican Museum, was senl,to me someveals ag'o bv the iate I)r. L. Peringuey. Oth'er speciesw.eretaken
in Cape Colony, SoutherrrRhoclesiaand PortugueseEast Africa |yN{essrs.;\lexander Cuthber:tson,}I. K. ftIunro and R. E. Turner.
Trvo very distinct speciesof Trenlepoltlzcfrorn the slopesof Came_roon \Iountain were taken b-v the Reveren.l\tr. Jacob A. Lieis.
I rvish to express my verv deep thanks to all the entomologists
above mentioned for their appreciateclcooperation in makihg
kttowtt the Tipulid fatrna of the Ethiopian Region. Except rvherc
noted to the contlary, the tvpes of the noveltiesare preservedirr
mv collection.

Limonia (Limonia) cuthbertsoni sp. n.

IuGeneral coloration black; mesonotal praescutum *vith three
reddish ochreousstripes;scutal lobes extensivelvreddish ochreous;
rvings rvit,ha st,rongbrown suffnsion,heavilv patternedwith darker brown; wing-surfacewith abundant macrotrichia.
M c t l c .- L e n g t h , a b o u t 7 m m . ; w i n g , 8 . 4 m m .
i\louthparts greatlv reduced, pale, the maxillary palpi black,
single-segmented.
Antennaeblack'throughout; basal flagellar segrneuts suboval, with short apical pedicels, the outer segments
becoming more.elongate,the basesnarrowed, not wider than the
short glabrous apical pedicels; terminal segment very long andslender, about one-third longer than the penultimate; verticils
long atttl cotrspicuous,mnch exceedirrgthe segments. Heaclblack,the occipital region paler.
\Iesonotal praescutum black with three conspicuo.usreddish
ochreous stripes, the lateral stripes produced caudad across the
suture, covering most of the scutal lobes, the mesal gdge of the'latter trarrowlv blackened; scutellum reddish-ochreous;mecliotergite black, paler caudally. Pleura back. Halteresinfuscated,'theD i p t e ra ,t. V II 1 1 9 3 1 1p, . 4 9.

50

C. P. ALEXATIDER

knobs more heavily so, the base of stem narrowly pate yellow.
Legs with the coxaeand trochanters teslaceousbrown; femora andtibiae brownish yellow; tarsi brorn'nishyellow, the two terminal
segmentsdark brown; claws (C) elongate,simple, with the exception of a single slendertooth at extremebase. Wings with a strong
brown tinge; a conspicuousheavy brou'n pattern, including extensive seamsat origin of Rs, fork of sc, stigma, along cord and outer end of cell lst M" and a conspicuousmarginal clouding in ends
of cells R, and frr; whitish areasbeforeand beyond stigma;centersof cellsR, NI, Ist fulr,1!I.and Cu pale; veins dark brown. Numerous
macrotrichia in all cells of wing' exceptingthe proximal ends ofbasal 'cells. Venation : Sc, ending opposite four-fif[hs the lengthof Rs, Sc, at its tip; Rs angulatedand weaklv spurred at origin;
R, and free tip of ,Sc"subequaland in transversealignment; basal
sectionof R, + . subequalto or a little longer than r-m; nl-cu justbefore fork of.M.
Abdom,enblack, the basal sternitesringed with yellow, especially the secondand third sternites;hypopygium dark. Male hypopygium much as in holotrtcha in the shape and structure of
the dististyle.Basistylewith the ventro-mesallobe producedsubapicallv into a slenderlobule. Gonapophyses
with lateral shoulders
more conspicuousthan in holotricha.
Hab. PortugueseEast Africa.
Holotype, C, Border Farm, R. Jardin, April Lg29(A. Cuthbertson);Coll. N" 3897.
I name this fine speciesin honor of my old-tirne friend andco-u'orkeron the Tipulidae, Mr. AlexanderCuthbertson. It is mostclosely allied to Limonia holotrichcrAlexander (Cameroon) and
L. bequ.aertiAlexander (Liberia), differing especiallyin the heavily-patternedwings and details of venation.
Limonia (Libnotes) libnotina sp. n.
Generalcoloration yellow ; scapeand pedicel black, flagellum
paler ; praescutumwith four brovvn stripes ; mediotergite pale incentral portion; pleura vellowish white with two narrow, darkbrown, longitudinal stripes; wings pale yellow, with a rather
sparsebrown pattern that is confinedto the vicinity of the veins;a gray clouding in the apical cells; abdomen yellow, tlie lateral
margins of the tergites narrowly dark brown.
Iulale.- Length, about 12.5 mm.; wing, LG mm.
Female.- Length, about 12 mm.; wing, 15.4 mm.
Rostrum moderately elongate, dark brown above, more red-

c

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I

TIPULIDAE

FROM TEE ETEIOPIANREGION51

dish ventrally, the surface Inore or less pruinose ; palpi obscureyellow. Antennae with scapeand pedicel brownish black, sparsely
pruinose; flagellar segments pale brown, the base of each segment,
narrowly yellow to produce an indistinct bicolorous appea-rance.
Eyes large, the vertex between them reduced to a mere line; headbrown, covered with a yellowish gray pollen ; a rufous area on
midline, immediately behind eyes; a deep median furrow.
Pronotum yellow. Mesonotalpraescutum vellow; a small brownarea on lateral margin of humeral region ; fou'r brown stripes
behind, very faintlv indicated on the anterior half of the segrnent,
the intermediate pair ending far before the suture, the lateral paira little broader, crossing the suture and suffusing the scutal Iolles, *'ith the exception of the caudal-lateral angles which arealmost vvhite and produced into conspicuousblunt lobes ; scutellum almost white ; mediotergite yellow medially, the sides broadly
dark brown, fainter on the cephalic portion. Pleura yellowish
white, with narrow, dark brown, longitudinal stripes; a dorsal
stripe extends from the cervical scleritesabove the fore coxae extencling to the . cephalic margin of the mesepimeron; a broader
stripe occupies the lateral portions of the mesosternum between
the fore and middlq coxae; a narrow brown line immediately ventrad of wing-root; a narrow line surrounding the posterior spiracle. Halteres pale, the base of knobs faintly darkened. Legs withcoxae _vellow,posterior apical margin of fore coxae narrowly darkened; trochanters' vellow, the margin narrowly blackened andtoothed; rest of legs detachedbut undoubtedly belong to this species, having femora yellow with a narrow brown subterminal
ring, the subequal apex yellow; tibiae and basal trvo tarsill segments brownish' yellow, the tips narrowly darkened; remainderof tarsi dark brovvn. Wlngs rvith a faint yellorv tinge; small
brown spots and dots along the veins, as follorvs : On R beyond
arculus; large spots at origin of Rs, fork of Sc, R, and tip of
Rr + : i series of smaller brown dots at the forks of the veins,
along radial and medial veins beyond cord, there being about
six each on dibtal sectionsof .Rn+ s, Mt and Cu,; marginal brown
clouds at cnds of most of the longitudinal veins; a brown seamiust before midlength of vein lst A and another at base of vein
2nd A; vein Cu. uniformly seamedrvith brown; longitudinal grayclduds in the outer end of cell R, centersof cells R, and Rr, and in
the outer ends of cells NIr, Cur, Cu andthe Anal cells; veins yellovr',
corrspictrouslvdarkened in the infuscated a,reas.Venation : R. + 2

52

C. P. ALEXANDER

fully three times R, alone and longer lhan tn-cu; Rs from three
to four times the basal deflectionof Ro * , and in alignment u'ith
it; cell lst x'1,long and narrow; basalcleflectionof lf, elongate,arcuated,nearly trn'iceul-cu, the latter at about tvl'o-fifthsthe length
gf the lou-er faceof cell lst l/r; veins beyondcord long and slencler,
as rrorrnal for Libnotes.
{bdomen vellow, the lateral margins of tergites narrou-lv but
conspicuouslydark brown, the color slightlv interrupted at thecauclalmargin of each segment;ovipositorreddish horn-color, thellasesof hvpovalvaeblackened;cenciverv short, strongly upcurved.
Hab. Cape Colony.
Holotype, C, receivedfrom H. K. Munro, without closer clata.
in author's collection.
Allotype, L Oudebosch,CaledonDivision, December fg20
(R. Lightfoot); South African Museum.
Tlre first true Libn,olesto be discoverediu the Ethiopian Regiotr.
Pseudolimnophila
albocoxalis sp. n.
General coloration of praescul,umbrownish black, with three
retldish orange stripes that are confluent in front; pleura striped
longitudinallv u'ith black and yellowish gray;"legswith the coxae
whitish, the basesnarrowly and conspicuously
blackened;remainclerof legs vellou'; wings brorvnishyellow, wjth a heavv llrown
pattern, the areassolidly darkened;a supernumerarvcrossveinincell Sc,' m-cu shortlv before midlength of cell lst x[ri abdomen
brorvnish black, the caudal marg'ins of segments conspicuouslv
ringed w-ith yellow.
Female.- Length, about 10 mm.; wing, 9,b mm.
Rostrum and palpi black. Antennae with scape, pedicel ancl
Irasalthree segmentsof flagellum pale yellow; flagellar segments
elongate-cylindrical,the verticils shorter than the segments.Head
brorunish yellow-; anberior vertex at, narrowest point nearlv threetimes the diameterof scape.
Pronotum above -vellow, the anterior notum more orange-vellow. Mesonotalpraescutumalmost coveredb,vthree reddish orange
stripes that are confluent in front, restricting the brownish hlack
ground-colort; lhc lateral borders and narro\/, strip-like interspaces;scutum and scutellum chiefly dark brown, the latter morepruinose; mediotergite with basal half vellowish sray, posterior
half and narrow lateral bordersbrownish black. Pleura alternately
striped with black and vellowish gray, arranged as narrow lon-

TIPULID.{E

FROU TIIE ETIIIOPIAN REGION

53'

sritudinal lirtes, the most conspicuous ones occupving the clorsollleural reg'lon, propleut'a, anepisterrlum, pteropleurite and ventral
sternoplettrite, ltterol) and coral bases; pleuroterg'ite brovr-nishhlack. Halteres olrscure vellorv, knobs dark brovr,!'r.Legs rvithco\ae chieflv atrd couspicuottslvu'hitened, thc basesuarrowlv and'abrr-r1:tlvblack; trochatrters clalk brown; femora yellow, scarcelv'
if at all clarkenecltovvarclstips; tilliae ancl tarsi yellorv, the terminalof latter brou'rt. Nings llrorvnish vellorr., rn'ith atrl'o -qegnterrts
lreavv llrou'u pattem, the markings much more intense ancl soliclly
clarkerredthan in g.v/clenstolpei,
not with pale centers ancl forming
c'celliform areasas in the last-named; a dark cloud at arculus; fi.rne
dark costal areas, the fir*stn1 1h. srrperr)umerarvcrossvein in cellSc, the seconclat origirt of Rs, the fourth at stigma; other long.itrrclinal't'eittsr,r'ith clark clorrclsat rnargin, these largiestat the Analveins; areasa[ rving-tip confluent, leaving small isolated vellow
s p o t s i n o u t e r e n d s o f c e l l s R r , . R . ,a n d l 1 r ; c o r d a n d o u t e r e n d o ft:ell l.sf.1/. narrc.l'r'lvsearnecl;a series of clouds along vein Cu,, theoutel otres srr-raller;veins brorvn. in ihe costal and radial fields
llecomirrg' more yellovu in the interspaces. Venation : A strong
sllperlrtllllerarvct'ossveitrin cell Sr-.,placed at mid-distaneebetrveen
arculus ancl orig'in of Fs,' tls lorrg, sqlrare and short-spumecl at
o r i g ' i n ; R " + 3 + . , g e n t l v a r c r r a t e d .s u l l e q u a lt o R , + , ; n r - c u j u s t
befole micllength of cell lst II"; cell .4,I,about one-thir.dlonger tfian
its lletiole; anterior arculus lacking..
-{llclomen brovvrrish black, ,!.,hecaudal margins of segments
c o n s p i c t t o r t s l vl i n g e c l r v i t h v e l l o r , v . O v i p o s i t o r w i t h c e r c i b l a e k e rred llasallv, the upcurvecltips tlarh red; hypovalvaepolished blackHab. Southenr Rhodesia.
Irolc,tvpe, 9, Chirinda Forest, nlelsetter Disti'ict, altitrrde
3.600 feet, ){ovember 1930 lA . Cutltberfsr,rn); British l{useum,through l'[r. Cuthbertson.
The onlv othel Ethiopian Pseudolinutoltlilu rvith a supemuInerary crossvein in cell Sc is P. gvldenstolpeiAlexancler, which
is r:eadilv tolrl by the paler, more diffuse, ocelliform areas on therl'ittgs, the venation, the leg-pattern, especially of the coxae, andthe verv clifferent coloration of the thorax and abdomen.

Limnophila(Elaeophila) venaguttula

sp. n.

Gerreralcoloration lrrow-nish -vellow; antennae short, scape ancl
peclicel 1:ellou', flagelltrm dark brorvn: a capillarv bro,R'n medianline on heacl,pronotum and mediotergitei pleura black, u'ith three
narrow vellor,r'longitudinal lines; u'ings yellorn', the veins

5 /r.

C. P. ALEXAIIDER

rvith numerous small brown spots and dots; tn elongate, longer
tlrarr m-clr; a supernumerary crossvein in cell il'f.
trIale.- Length, about 6.2 - 6.4 mm.; w-ing, 7.2 - 7.3 mm.
Length (excluding head), about 6.8 mm. ; wing,
F emale.
8.4 mm.
Rostrum dark brorn'n,obscureyellow above; palpi dark brown.
Antennae short; scape and pedicel light yellow-, flagellum dark
brorvn; flagellar segments cylindrical, the outer face slightlv convex, terminal segments more elongate. Head uniformly brownish
vellon'; a capillary dark brown line o.n occiput and posterior portion of verte,x.
Pronotum obscure brownish yellow, with a capillarv darkbrown median line. Mesonotal praescutum light brownish yellow,
w-ith four narrow dark brown stripes, the intermediate pair approximated; a brow-n median spot near cephalic margin of praescutum; lateral stripes less distinct; remainder of mesonotum obscLrre brorvnish yellotv, the mediotergite with a consp!'cuous capillary dark brown line. Pleura dark brorvn with three very narrow light yellovv longitudinal stripes on mesopleur'a,the two more
dorsal ones extending cephalad into the propleura, the ventral
stripe lying between the fore and middle coxae, in some cases
including the mesosl,erllum. Halt"eresyellow. Legs vr'ith coxae dark
brovr'n, apex of fore coxa traversed by the ventral pleural stripe,
apices of other coxae more narrowlv pale; trochanters obscureyellow; remainder of legs broken. \Yings light yellow, with anabundant pattern of brown spots and dots on all the veins; thespots are but slightly larger than the dots, occupYing the originof .,Rs, tips o[ Sc and R, *,, alorrg cord, and at bhe ends of thelongituclinal veins ; the dots occupy all the veins, more abundant
beyond the cord ; in cell C there are but four sttch spots ; alongvein 2rtcl A fbur dots, additional to the larger apical spot ; veins
vellow, dark brown where they traverse.the intuscated areas. Macrotrichia of disk small, longer and more abtrndant in the darkened areas. Venation : Sc long Sc, extending to a short distancebeyond fork of R, * , * o, Sc, at its tip ; Rs long, arcuated atorigin i R, * , * o sltort, about equal to m-cu i R" subequal toR, *r, R, * , to R.; inner ends of cellsRn and R, in alignment;
cell Ist IV, elongate, irregularly pentagonal ; n" very long, arcuated, longer than rn-cu.: basal section of vein M, shorter than rrn and onl,v.about one-third m; petiole of cell M' about one-halfthe cell ; m-cu just before midlen glh of cell lst M2 ; sometimes

TIPULIDAEt
I

FROM THE ETIIIOPIAN REGION

rrp

aa

rvith a small spur near t,ip of vein 2nd A ; anculusbroken or nearlyso ; a supernumerary crossveiu in eell [W, opposite or slightly
lteyond the origin of Rs ; in one wing of allotype, this crossYein
is incomplete.
Abdomen dark brou'n, with a $parseobscure yellow pollen ;hypop-vgium scarcelv brightened. Ovipositor with valves elongate
but, fleshv, the verv compressedcerci deep, u'ith the tips obtusely
roundedHab. Cape Colonv.
.
1921
Llototype,C, ColdstreamHumansdorpDivision, JanuarylR. Tucker). Allotopottpe, 9. Paratopotypes,2 dd.
The referenceto the subgenusEl.ueophilois somelvhatdoubtful.
The peculiar structure of the ovipositor seemsto place it in a rather isolateclposition.
(Elephantomyia)
luteipennis
sp. n.
Elephantomyia
General coloration' yellow, the mesonotal praescutum Inore
or..rg" yellorv, with three narrow brown stripes; legs.black ;wings yellow ; Rs elongale, angulateclat origin ; deflection of Ro
long ; tn-cu trot far from baseof cell Ist fu[".
Length (excluding rostrum), about 10-11 mm. ;F emaleu ' i n g , 9 . 4 - 9 . 8m m .
Rostrum elongate, black, about threlepfourthsthe .,leqgth of
bocty. Antennae black, u,ith elongateverti'cils. Head light gray.
rn'ith three narrow'X{esonotalpraescutum light orang'e-vellowindistinct brown stripes; m,edianstripe distinct anteriorlv, becoming obsoletebeyond midlength of the sclerite; lateral stripes
crossing the suture on to the mesal portions of the scutal lobes;
remainder of mesonotum arrd .the pleura pale vellow. Halteresvellorv, the hnobs dark brou.n. Legs with the coxae and trochantersvellow, the apicesof the latter blackened;femorablack, basepaler, extreme tip narrow'lv paler ; tibiae and tarsi black ; tibial
spurs apparently lackirtg'. Wings w-ith a strong yellow tinge,
slightly more saturatedalong costal margin ; stigma faintly indicated; veils brownish black. Velation : Sc long, ending about
oppositethree-fourthsthe length of Rs, Sc, al, tip of Sc, ; Rs elongate, angulated and sometimesspurred at origin ; deflection ofR, about two and one-half times r-m ; r-m and nt subequalor thelatter longer;cell lst il/, pentagonal,rn about one-halflonger thanthe deflection of ll4r, irr the type provided w-ith about five macro.trichia ; nl-cu,a short distance beyond fork of M, in casesthe dis-

56

C. P. ALEXANDER

tarrce almost as long as r-nr ; vein lst il rvithout macrotrichia,delicate, its extreme tip entirelv atrophied.
Abdomcn vellovr'. Yirlves of ovipositor elor-rgate,very slentlcr
arrd acicular, horn-colored.
LIab. Cape Colonv.
Ilol,ot'ype,?, Coldstream, Htrrnatrsdorp Division, Jattuary
1921 (R. Tuckcr). .4llot-opotype,9.
Elcphantontyia luteipennis is very different from the othcr
Ethiopian specicsso far described.

Elephantomyia
tana subsp.n.
XIal,e
6.4 mm.

(Elephantomyia)

wahlbergi

mon-Length (excltrding tos[r'um), abottt 5 nrm. ; wing,

Rostrum lllack, bloken before midlengl.h. Iutenuae black.
Eves verv lalge, only rrarrowlv separatetldo:sallv, broacllv contig'uous ventrally. l-Ieatl dark gray.
N'lesonobal
praesctrir,rnobscure brou'nislt vellou', with a broatl,rlark brown, median stripe exlcntling'the length of the sclerile;
scutal lobes, scutellum atrd base of mediotergite clull retldish
lllow-n, lhe posteriot matgin of ttre last dark brovr'tr. Pleura uni*
form ollscure ycllow. Haltercs pale, the base of stem narrorvly
yellow, the knob brown. Legs with coxae and trochanters obscure-1.ellow; remaincler of legs broken. Wings sttbhvaline; stigma
elongate-oval, dark browt) ; pale ltt'ou'tt seams along cor{ andotrl.erentl of cell Lst trl"; darker cloudings in cell Sc, distal endso[ cells 1[1.and Cu, along vein Ctr, antl the basesof cells Cu, lst A
a:ld Zntl.4; veins dark brown. Venation : Ag in uestuoocli, differing as follows : Ils a little longer, more gently arcuated; basal deflectation of R, a little shorter than r-nI; cell lst M" lotlger, the basal
section of.tr[, + ,, angttlate; m arcuated or angulated at near midlcrrgt,h, nearly as long as the basal deflection of IVI";m-cu at nearl.hree-{ifthsthe length of cell lst.l/r; vein lst.4 with macrotrichia
on distal two-thirds.
Adbomen with tergite olre dark brou'n ; strcceeding tergites,
dark brovvn, the caudal and lateral margins obscure yellow, narrower on the posterior segmeuts vvhere most of the segmenf isdark; segment eight black to form a conspicuous subterminal
rirrg; hypopygium yellow; sternites similar but the dark brorvn'of dorsum replaced by pale yellowish brow-n. n{ale hypopygium

TIPULIDAE

FROM THE ETEIOPIAN REGIol\a)/v y i l l r o r r t e r r l i s t i s l v l es l e n d e r ' ,b i f i d a t a p e r , t h e s r r l l t e r m i r t a ls p i n e
s m a l k r b t r t m o r e a e t r t et h a r t t l t e a c t u a l t i p .
llal->. Cape Colonv.
Ilolctvpe, d, Cuclebosch, Caledon Division, Decemllcr ,192t-)(R. l.ioht.lottt).
Tlre rlescription ot lllrphctttortr.'yfott'cltll>ergi, Bergroth (Caffralia) rliffers in the details of cololntiou of the bodv and u'irtgs,
arrrl in tht' venatiott.

Gnophomyia capicola sp.n.
Ceneral coloralion hrou'n : heacl gravish prtrinosc: tlroracic
a lrrolvuish
1 , . l c u r :rrv i t h a p a l e l t ' r r r g i t u t l i n asl t l i p c ; u ' i n ; l s w i i hv e l l r ; r v l i n g e ; s l i g m a e l o r r g a t e ,d a r k b r o r v r r : c o r t l f a i r t t l y s e a m e dn - i l . hl t l o n t t , r - r n a [ o r b e f o r e t h c f o r k o f / t . s ; v a l v e s o f o v i p o s i 'loi'ficshv.
i l a l e . - L e n g t h , a l l o r r t5 . i r m m . ; w i t r g , 7 . 4 m m .
Fpnmle. - Lengllr, about (i mm. ; rvitrg, 8.5 mm.
Rosl,r'um and pall;i llrowu. Anteurtae with the seape ancl
p e d i c e l c l a r k l t r o w n ; f l a g e l l a r- s e g m e r r l s r l a r k b r o r t l r , e l o n g a t e cvlindrical. Head gralish prttinose.
Mesottotttm[)rorrottrm browrr, nArro,rvlv g'rayislt mediallv.
rlalk llron'n, the praescutalinterspacesonlv a. lit,tlc paler than therrcarlr, conflrreul,stripes ; humeral region artd latetal margins ofI'rilescuttrm oltscttre ;:elloruish ; scrrtal lohes l]alc laterally ; sctrtellum with the cauclalhalf paler than base; metliotergitc sparselv
prtrirrose. Plerrra rlark brolvrr ; a t,utts11'pflle longitudinal slripe
a('ross l.he ve.nl,ralplerrral reg'itltt. I'Ialteres<lark browtl, the exlrr:me llase of stem pale. Legs with coxae pale testaceous; trocharrlers le'-stacco,us; remaintler of leg= brownish vellow, darker
irr male. Wings with a brorn'nish yellorv, [inge, prearcular atltl
cos:talreg'i<tnssomew'hat lltig'ht,er; stigma conspictlorls, elongate, ; , v a l .r l a r k l r r o r v l r ; a f a i r r I a n d i n d i s t i n c t l r r o w n s e a m a l o n g c o r d
a r r r fo r r t e r e n t l o f c e l l / s f i l [ " ; v e i n s t l a r k t l r o r v r t . V e r r a t i o l r ; ^ S c ,cntling oltJxtsite folk of ll.s ; Sr:, a[ about orte,lhird the distancc
h e t r v e e no r i g i n o f R . sa n c l [ i p o f S c r , ' R . sl o n g a n d s t r a i g h t i F g * , * os t r o n g l v a r c u a t e t l ; R , c l o s et , o f o r k o f R r * . , ; c e l l F , t l a r r o w b a sallv ; r-tn connecting' vvith F.s at or beforc (in female) its fork;
I?.sirr aligrtment r,vitlr ^Rr;cell l.st tr[, long ancl llarrow, closed;
rrr-cu al, or irrst llevonrl the fork of. IV; veln 2nd. il gentlv cttrvetl
to margin.
Valves of ovipositor short, bltrrtt and,\bclomen dark brorvn.
flc'shv, hvpovalyae clull orange.

58

C. P. ALEXANDER

Ilctb. Cape Colony.
altitude 3.000 feetn
Holotype, C, Keiskamahoek,Lenye lrorest, altitude3.000
Allotopotype, 9, in Mr Munro's
June 15, 1921 (H. K. Munro).
collection.
The fly is verv different from all other knorvn speciesof thegenus. The fleshy cerci and hypovalvaeof the ovipositor are notnormal for the g'enusand it may later be assignedto some olher
group.
(Trentepohlia)
sp.Ir.
clitellaria
Trentepohlia
Genelal coloration pale yello\M; lhorax with a brownish black
meclian shield, narro*'est in frortt, more widened behind ; median region of sctttum very restrictedly pale ; pleura pale yellorv ; lcgs yellow ; rvings whitish subhyalinr:,with a restricteddark pattern; abdominal tergites brown, the sternites bicolovvith less than the apical halves of thc
rous, their bases_vellovr',
segmentsdark brow-n.
MaIe. - Length, about 5.5-6 mm. ; u'ing, atlout 4'r:-4.8mm.
h'enmle.-Length, abotrt,6.5-7 mm. ; w-ing, 5-5.2 mm.

Describedfrom alcoholic specitnerrs.
Rostrum and labial palpi pale, maxillarv palpi darker. Arrtennae dark brown throughout. Pr6thorax and mesothorax yellow. the notum wilh a brownish black median area, beginningon pror)otum, becoming wider on praescutum, extending to baseof abdomen, on sctttttm leavinil a verv natrow, pale, median line.
Pleura errtirelv pale yellow. I{alteres pale. Legs 1-ellow; femora near bases set rvith small spinous setae; posterior tiSiae withtwo conspictroussetae near outer encl ; proximal end of posterior
basitarsi rvith tlre setac more abundant, somewhat brush-like.
Wings r,vhitish subhvaline, with a restricted dark pattenr ; stigma
small, dark brorn,tt; wing-apex extensivelv paler brown ; a nar'rov\r scam on Rr, leavitrg'tr pale atea i1 crell /1, i l)arrolv dusky
seams along posterior cord and vein Ctt ; veins chiefly brown.
Venation : Sc" obsolete or.nearlv so ; R"+r+o diverging widel.v
from R., at origin i F, sullequal to or a little longer than. F.*, i
Rr*n more yellovvish, about one-half to lhree-fourths longer thanRr*r*oi fusion of Cu, and,lst.4 very slight; cell Znd A narrow.
Abdominal tergites brown, sternites bicolorous, pale yellorv,
with less than the ap,ical half of each segment dark brown, llte
amount becoming somelvhat more extensive on the outer seg-

TIPULIDAE

FROM TEE ETEIOPIAN REGION

59

ments ; subterminal segments and genitalia of male more uniformly darkened.
Hab. Cameroon.
Holotype, alcoholic d, above Buea, CameroonMountain, altitude 6.500 feet, October1929 (J. A.'Reis). Allotopotype 9. Paru
topotypes,d I .
By *y key to the Ethiopian speciesof Trentepctlilia(Reu.Zool.
Alricaine, 14 : 177-180; 1926), T. (7.) clitellaria runs Io exornata
Bergroth, a very different fly. In reality, it is closer to T. pallidipleura Alexander (Cameroon),differing in the thoracic and leg.
pattern and in t,hevenation.

Trentepohlia(Trentepohlia)

jacobi sp. n.

Generalcoloration of mesonotalpraescutumlight yellow, witha restrictedbrown pattern ; dorsal pleurites dark brown, the sternopleurite light yellow ; coxaeand trochanters dark brorru-n; wings
broad, wtritish, with a moss-like brown pattern ; Sc and R, * ,widely separatedat ma,rgin, the distance on costa exceedingvein
R, i macrotrichia of veins verv sparse.
FenmleLength, about 5.5 mrn. ; wing, 5 mm.
Describedfrom alcoholic material.
Rostrum pale ; palpi black. Antennae with the first segment
black, the pedicel yellow to brownish vellow; flagellum paleyellow, the outer segmentsbroken. Head o'chreous.
,Cervicalscleritesbrown. iVlesonotalpraescutum light yellow,
with a sparsebrorn'n pattern, including a median dash on cephalic portion of sclerite and tw-o dark dashesjust before the suture,
representingthe usual interspaces; lateral margins of scleritemore
broadly brown ; scutum vellow, the lobes extensivelydark brown;
scutellum chieflv dark brown ; mediotergitedark brown. Pleura
vvith dorsal scleritesand membrane brown, the broad sternopleurite conspicuotrslvtight yellow. Halteres pale, the knobs infuscated. Legs rvith coxae and trochantersdark brown ; remainderof legs broken. \Yings relatively broad, rvhitish, the base andcostal region slightly m,ore yellow ; disk with a heavy brot-n
moss-like pattern, appearing chiefly as two broad bands that aremuch dissectedbv white areas; wing-apex with three major areasof the ground-color ; a gray suffusion in outer encl of cell Ist A
and beyondmidlength of Cu; four small scatteredspotsjust beyond
wing-origin ; veins dark brown, pale yellow in the ground areas.
Macrotrichia of veins almost lacking, beyond cord with a scatrtv
series on distal section of R, and a few others on distal end of

I

{;0

C. P.

ALEXANDEIT

Rr, and on llr. Yenation : Sc, ending far befole R, * ", the distance olr costa greater than lt, i Rs long, gentlv sinuous ; celll?, large ; apical fusion of veins Crr, and lsf ,l slight.
-\lldomiual tergites clark brou-rr, the'mol'e basal scleriteswith
a pale transversecentral ring ; sternitessimilarlv dark brown, on
segments t.rvo to four u.ith bases chiefl_vrvhiteuecl; subterminal
segments trniformly clarkenecl; ovipositor horu-rellou-.
Hab. Cameroon.
lTttlottpe, alcoholir) 9, Above Brrea, Cameroon N{otrnbain,altitucle 6,500 feet, October 19?9 (/. A. Reis). Parcdopotype,alcoholic ?.
I-take the greatesl pleastrrein naming this erceptionallv distinct Trerttepohliain honor of ttre collector, Dly'friend, the Reverend \'[r', Jacob A. Reis. The speciesis arnply clifferent from all
knovvn Ethiopian members of the genus. By my k.y to the
Ethiopian member-sof the gentrs (1. c., L4 : 1?7-180; 1926), thespecies tur)s to couplet 20, flisagreeing widely rvith all species
bevond this poirrt. The moss-like or stellatepattern of the rvingS
is somevvhat, suggestive of that found in certain Trvpetidc, as
Ltrelliu.

Gonomyia (Progonomyia) nigrobimbo

sp. n.

Coloraticn black, the pronotum, scutellum and a ventral longituclinal line on pleura obscure nrfous: legs entirelv black;wings tinged rvith black ; distal seetion of vein M" lying free inthe membrane ; abdominal sternites rlnged eatrdallv with rufous-v-ellow.
Length, about 4 mm. ; rviug, 3.6 mm.
trlale.
Fentctle.- Length, aboul 5.4 mm. ; rving, 4.3 mm.
Rostrum ancl patpi black. Antennae black throughou[ ; flagellacl segments short-c1,-lindrical. I{eacl black.
Pronottrm black, the scutellrrm conspictrous,obscure rufous.
\'fesonotrrmblack, t.hc extreme lai;eralmargins of praescutum andthe postcrior lateral margins of the scutal lobes narrowlv obscurerufous. Pletu'a lllack, sparsely gra-v pruinose, with the dorsopleural memhrane ollscure nrfotrs ; a pale rttfous longitudinal
strille alrove the coxae, extending frorn mesepisternum to immecliately above ;losterior co,.x?.e.Halteres black. Legs with coxae
black, sylarselvgray pruinose ; rernainder of legs black. Wings
with a strong blackish suffusion, deepest in the radial field andalorrg vein Cu, ; veins black. Venation : Sc long, Sc,. ending
approrimately at, midlength of Rs, Sc" far from the tip of Sc., im-

TIPULIDAE

FROM TEE ETEIOPIANREGION

6l

mediately beyoncl origin of Rs ; R, very faint but indicated, immediatelv before fork of R, * n i deflection of Ru angulated.at midlelgth ; distal section of M, lying free in the membrane as inpulchrfssima; rrL-cuimmediately beyond fork of ll:I.
Abdomen black, the posterior' margins of the segments very
narrorvlv pale, t,his color scarcely evident on the tergites, a little
broacler and more obscure rufous yellow on the sternites where
it appears as narrow bands. lltale hypopysium black, including
the appendages. Ninth tergite narrowed apicallv, the caudal margin with a V-shaped notch, the lobes formed slightly diverge.t.
6vipositor with nul,n.. long and slender, brownish black'H a b . ' C a p eC o l o n Y .
Holotype, d, Oudebosch, Caledon Division, December 1920
iR. Lightfoot). AllotoPotYqe, 9.
The speciesis readily told from its only near allv, GonomYia( Progortomiyia) pulclvissima Alexander, by the diagnostic features listed above.

Dasymolophilus liliPutanus sP' n'darksize very small (wing, d,2.2 mm.); general coloration
brorvn ; wings grayish brown ; no macrotrichia in cells of wing ;R, and R, * , in almost transversealignment, both shorter thann_ + , ; male hypopygium with the dististyle flattened, glabrous.
AIaIe.- Length, about 1.5 mm' ; wing, 2'2 mm'
Rostrum, paipf and antennae dark brown, the lat'ter short'
Ilead dark brown
Thorax dark brown, slightly pruinose. Legs brown. wings
with a grayish brown tinge, the veins darker brown ; macrotrichia rlark brorn'n. No macrotrichia in cells of wing. Wings appain therently with the anal lielcl reduced in size, this area injured.onlv preservedwing of the uniqtre t-vpe. venation : sc relatively
origin of Rs ; Rs
shorti Sc, encli.g-utopp.ositeR", Sc, jtrst be-vondo,ii two-tfiirds R, * , i R, * ' and R2 suberelativelv short,qual in length ancl almost in lransversealignment i Ro * , frommone and one-half to tw-o times Rr.* r, itr alignment with Rs;
about
cu beyond fork of 1t1,.petiole of cel.l M, relatively short,
trrice n7-ctr.
posiI,Iale hypop)-giurn with the single dististyle termina-l in
is tiption, glabrous, flattened, on face bearing a small lobe that
pea Gtn two tinv setae. Phallosome a depressedplate, the cau,.lal *a"gin slightlv concave, each caudo-lateral angle terminating
in a slender, smooth, darkened spine'
Diptera, t. VII [1934].

o:

62

C. P. ALEXANDER

I
I

Hab. C"p" Colony.
Holotype, 6/, ceres, cape Province, April Lgzb (R. E. Tarner):
British Museum 1925-2L0, sent rne through the friendly interest
tlf Mr. Fred W. Edwards.
Allied to the larger Dasymolophilus eriopteroides (Alexander),likewise from the mountains of southwest i"pu ,Colonv, differi.g
conspicuously in the venation, especiallv the more elongate veins
beyond the cord, Rs being shorter than R, *, , Rn + 5 distinct
and longer than either R, or R, + 3. In eriopteroides,Rs is longer than R, * r, cell Ro at margin being considerably widened:
R, * s and R. arise exactlv at the fork of Rs, there being no element Rn * ,. I regard the ?'aseoceraminutissima Edwards (Trans.
Linn. soc. Lo'don, zool. (z) lb : zL}-zLL ; lg12) as likewise belonging to this genus rather than to Tosjocera. However it is beooming increasingly difficult to separate the species with shortantennae from the typi-cal Tas.iocerrewith elongate org-ans, ancl
clear-cut distinctions between t.he two groups are still lackingThe Australasian species of Tasiocera, including the genotype,have the wings more cuneiformly narrowed than in these tSree
Ethiopian species of. Dasyrnolopltilus, resulting in the shortening
to total loss of vein 2nd A.

Diptera, tome VII [1934](Paris, Paul Lechevalier gt fiIs).

t