Frorm the ProceeiXingsol the Linneon Societg ol Nep Soutlt,Wales,Vol. lai, Parts 8-4, 1986.

CONTRIBUTION TO A KNOWLEDGE OF PAPUAN TIPULIDAE(DTPTERA).
By CEenrns P. ALExAlrorB, Massachusetts State College, Amhergt, MasB., U.S-4.
(Oonmunicoted, by Franle H, Taglor, F,R.E,B,, F-2.5.,(Seven Texbflgures.)lRead 2{th June, 1986.1

llssued 15th September, 1936.1

From the Proceeilings of the I'innean Society af New South Wales,Vol. lni, Parts 3-4, 1936.

CONTRIBUTION TO A KNOWLEDGE OF PAPUAN TIPULIDAE(DIPTERA).

By Csenr,rs P. Ar,nxennnn, Massachusetts State College, Amherst, Mass., U.S.A.
(Contmunicateil,by Erank H, Tagl^or,F.R.E.S., E.Z.S.)(Seven Text-flgures.)lRead 24th June, 1936.1
The material included in the present paper is chiefly from the mountains of
the Central Division of Papua nolth of Port Moresby, and was collected by Mr.
K. J. Clinton of the School of Public Health and Tropical Medicine, The
University of Sydney. The collection contained eight species,six being describedas new. The subgenus Paramongonla is recorded for the first time from Papua.
I express my very sincere thanks to Professor Harvey Sutton, Director of theSchool of Public Health and Tropical Medicine, for submitting this material to mefor study. All types and uniques are preserved in the collection of the School.
LrrltoxrrNep.
LrlroNrrNr.
LruoNra(Lrlrolt..L)

T.oNGEANTENN'\'IA,n. sp.

Figs.

1, 3.

Size small (wing, 3, 4 mm.) ; general coloration of thorax dark brown,
praescutum
more reddish-brown; femora yellow, with a narrow, subterminal,
thedark ring; wings with a uniform brown tinge, the stigma very small, subclrcularin outline; male terminalia with the basistyle armed at tip with three groups ofmodified setae; dististyle profoundly divided into a dusky clavate outer lobe andan oval inner lobe that is produced into a slender spinous point.
J. Length about 3'5 mm.; wing, 4 mm.; antenna about 2'7 mm.
Rostrum black, about one-third the length of remainder of head; palpi black.
Antennae (d) of unusual length for a member of this subgenus,being only a little
shorter than the entire body, black throughout; flagellar segments long-cylindrical,
with short glabrous apical pedicels; tet'minal segment subequal in length to thepenultimate, strongly narrowed and pale at apex. Head gl'eY, mol'e silvet'y'grey onfront; eyes large, rather broadly contiguous on vertex.
Pronotum dark brown. Mesonotal praescutum more reddish-brorffn,somewhat
darkened behind; scutal lobes dark brown, the median area and the scutellum
more testaceous-brown; mediotergite dark brown. Pleura uniformly dark brown,
the dorso-pleural region and meron somewhat brighter. Halteres dark brown,
the base of stem narrowly pale. Legs with the fore coxae darkened, the remaining
coxae more yellowish; trochanters yellow; femora yellow, with a narrow
subterminal dark ring, the actual tip a little narrower, yellow; tibiae and tarsi
brown; claws small. Wings (Fig. f) with a uniform brown tinge; stigma verysmall, subcircular in outline, darker brown; veins brown, macrotrichia
conspicuous,black. Venation: Sc, ending shortly before midlength of the long Rs,Sc, at tip; free tip of Sc, aud R, in transverse alignment; m a little longer andBY C. P. ALEXANDER.

L23

more arcuated than basal section of Mr; m-cu just beyond fork of 1\I; anal veinsparallel for almost the entire length of 2nd A.
Abdominal tergites dark brown, the basal sternites more yellowish-brown;
terminalia obscure yellow, the outer lobe of dististyle more infuscated.
Male terminalia (FiS. 3) with the tergite, gt, large, the caudal margin
gently emarginate, the low, broad, lateral lobes with long, conspicuous setae.
Basistyle, b, short, the ventro-mesal lobe conspicuous, at apex armed with threegroups of splnes and setae, including a low mass of tubercles shortly before
apex, an outer apical lobe set with long setae, and an inner apical row of four
flattened black fasciculate setae. Dististyle, d, single, produced into a longclavate lobe, with very long erect setae; main body of style elongate-oval,atapex drawn out into a straight, slender, spinous point, Mesal-apical lobe ofgonapophysis,g, a long flattened blade, the apex obtuse.
Holotyrre,J, onoba, Papua,altitude about 2,000feet, July, lg3b (K. J. clinton).
The present fly is readily distinguished from similar small species of thesubgenusby the unusually elongate antennae of the male, pattern of the femora
and, especially,the structure of the male terminalia.
LrMoNr.r (Lrnrvorus) olvoseNa,n. sp. X''ig. 4.
General coloration obseureyellow, the praescutum with a black median stripethat spreads out in front to cover the humeral and lateral portions of thesclerite; halteres yellow; femora obscure yellow, the tips very narrowly clearer
yellow, the fore femora with a narrow brown subterminal ring; wings tinged
with yellowish-grey, margined with dusky, cells c and Sc more infumed; sc,ending just before fork of Rs; Rs gently arcuated; m-cu just beyond one-third
the length of cell 1st mr; anal veins slightly convergent at bases; abdomen black.
3. Length about 6.5 mm.; wing, 7.5 mm.
Rostrum and palpi black. Antennae black, the outer segmentsa little paler;
flagellar segments oval, with short apical pedicels; terminal segment elongate,
about one-half longer than the penultimate. Head dark gley.
Pronotum black. Mesonotal praescutum with the ground-colour obscureyellow, with a median black stripe that is widened in flont and extends lateradaeross the humeral and lateral portions of the sclerite, restricting the groundcolour to broad sublateral areas; at about midlength of the segment, the black
stripe sends cephalad a small lateral point or spur, isolating a linear ground
dot on either side of the midline; scutum with the median area yellow, the lobes
black; scutellum black, paler basally, the parascutella yellow; mediotergite dark,
the cephalic portion paler. Pleura obscure yellow, the propleura and a uarrow
longitudinal stripe across the anepisternum and pteropleurite to and including thepleurotergite, blackened; fore coxae and a small vague area on sternopleurite
darkened. Halteres pale yellow throughout. Legs with the fore coxae darkened,
as described; remaining coxae obscure yellow; trochanters obscure yellow; fore
femora obscure yellow, with a very narrow, brown, subterminal ring, the yellow
apical portion a trifle broader than the ring; middle and posterior femora obscureyellow, becoming a little darker outwardly, but without distinct dark annuli, thetips narrowly clearer yellow; tibiae and tarsi yellowish-brown,the terminal tarsal
segments blackened. Wings tinged with yellowish-grey; cells C and Sc more
infumed, passing into brownish-yellow outwardly; bases of cells R, M, Cu andlst A slightly darkened; a narrow dusky border around the wing, somewhat
more extensive on the apical portions; a tiny cloud at origin of Rs; stigma small,124

PAPUAN TIPUI,IDAE.

oval, darker brown; veins dark. Venation: Sc t'elatively short, Sct ending justbefore fork of Rs, Sc, at tip of Scr; Rs gently arcuated; free tip of Sc, and R, intransverse alignment; inner ends of cells 2nd M, and M. in transverse alignment;
m-cu just beyond one-third the length of cell 1st l\tr; anal veins slightly convergent
at bases.
Abdomen black, the terminalia only a little brightened. Male terminalia(FiS. 4) with the tergite, 9t, relatively nat't'ow apically. Basistyle, D, long andslender, the ventro-mesal lobe basal in position, provided with long coarse setae.
Dorsal dististyle a short curved blackened horn, the tip acute. Ventral dististyle'od,,a small oval lobe, with long setae; rostral blade long and conspicuous,pale,compressed,at apex narrowed into a small spinous point.
Holotype, 3, Onoba, Papua, altitude about 2,000 feet, July, 1935 (K. J.
Clinton).
By Edwards's key to the species of.Libttotes (Journ. Fed. Malay Bt. Mus., t4,1928, ?4-80) the present fly runs more or less directly to Li,monia (Libnotes)
trifasciata (Edwards) of the Federated Malay States, a very different fly. The
peculiar structure of the male terminalia is somewhat as in an otherwise entirely
distinct species to be described, in a later paper of the series, from Kavieng,
New lreland.
Lnrowre (Lrnwores) cr,rwtonr, n. sp. Fig. 2.
General coloration grey; praescutum with a brownish crossband behind theanterlor margin, together with two comma-shapedbrown spots beyond midlengthof the sclerite; fore legs entirely black; wings faintly tinged with yellow, sparsely
patterned with brown; free tip of Sc, in virtual transverse alignment with Rr;
abdomen dark brown, the terminalia brownish'black.
3. Length about 11 mm.; wing, 15 mm.
Rostrum brownish-black; palpi black. Antennae with scape black, pale atbase; pedicel brownish-yellow; flagellum yellow; flagellar segments long'oval, thelongest verticil of each subequal in length to the segment; terminal segment
about one-fifth longer than the penultimate. Head light grey; anterior vertex
reduced to a very uarrow strip that is only a little wider than the diameter of asingle ommatidium.
Pronotum reddish-brown, darker on posterior border. Mesonotal praescutum
chiefly greyish-pruinose, with a brownish crossband behind anterior margin,
continued backward aeross the humeral and lateral borders to the suture; a pairof intermediate brown dashes,shaped like commas, placed just beyond midlengthof praescutum; scutum whitish, the lobes with more greyish centres; scutellum
and mediotergite whitish, the latter with a brownish-black lateral line, widened
posteriorly. Pleura whitish-grey, with narrow, broken, dark brown, longitudinallines on anepisternum and pteropleurite, and again across fore coxae and ventral
sternopleurite. Halteres with stem yellow, its base infuscated, the knob blackened.
Legs with the coxae whitish, the fore coxae marked as above described;
trochanters whitish; fore femora entirely black; middle and hind femora brownbasally, passing into black at or near midlength; all tibiae and tarsi black;vestiture of legs relatively short and inconspicuous,short and spinous on femora,
longer and more appressedon distal segments. Wings (Fi8. 2) long and narrow,faintly tinged with yellow, the costal cells more saturated; a restricted dark
pattern that is conflned to the veins and distributed as small areas as follows:
Arculus; origin of Rs; at intervals along cord; outer end of cell 1st Mr; tip ofSe, and Rr; at distal ends of veins M., Cu' lst A and 2nd A; veins bright yellow,

126

BY C. P. ALEXANDER.

black in the clouded al'eas. Venation: Free tip of Sc, lying immediately basadof Rr, so as to appear virtually in alignment with it; cell 2nd M, about one-thirdlonger than cell M.; m-cu just beyond one-flfth the' length of lower face of celllst Mr; anal veins convergent basally.
Abdomen dark brown, the basal tergites darker laterally, the intermediateones further darkened medially at base; terminalia small, brownish-black;sternites paler.
Holotype, 3, Onoba, Papua, altitude about 2,000 feet, July, 1936 (K. J.
Clinton)This interesting and very disi.inct Librtotes is named in honour of theeollector, Mr. K. J. Clinton. It belongs to the notata complex of species,being
most leadily distinguished from all described allies by the extensively blackened
legs.

nL tltti
t\v\F'igures 1-?.
D, basistyle; d, dlstistyle:, g, gonapophysis; t. tergite; ud, ventral dististyle.
1. Litrtonia (Lim.onia) longeantentr'ata, n. sp., venation'
2. Limonia (Libnotes\ cl,intoni, r. sP., venation.
3. Limonia (Lirnort'ia) longeanten'nata, n. sp., terminalia.
4. Lirnonio (I'ibnotes) onobana, n. sp., terminalia.
I'ucrilera, n. sP., venation'
5. Trentepohl,ia (Paramongotna)
6. Trentepohl,ia (Mon'gorna) longisetoso, n. sp., venation.
7. Glonomyia (I'ipophleps) iwrata, n. sp., venation.

Entoprrnrwr.
n. sp.
Tnrnrnponr,re (Plnentorooua) LUcBTFERA,Fig. 5.
General coloration of thorax uniformly brownish-yellow; antennae with scapeyellow, pedicel and flagellum brown; legs pale brownish testaceous; wings paleyellorvish-grey, unmarked; veins pale; macrotrichia of veins very reduced innumber; Rr*r*. subequal to Rr*n alone, both a little longer than Rr.
9. Length about 3'8 mm.; wing, 3'? mm.
Rostrum and palpi brownish-yellow. Antennae with the scape yellow;
pedicel and flagellum brown; flagellar segments suboval, the verticils short.
Head brownish-yellow.

126

PAPUAN

TIPULIDAE.

Mesonotum and pleura almost uniformly brownish-yellow, rvithout distinct
markings. Halteres pale. Legs with the coxae and trochanters yellow; remainderof legs pale brownish testaeeous, the tarsi a little paler, dirty white. Wings(F''ig.5) pale yellowish-Br€y, unmarked; veins pale. Macrotrichia of veinsvirtually lacking, there being two or three on vein R, near stigmal region andtwo or three scattered trichia on distal section of vein RE. Venation i Rr*s*.
subequal to Rs*a,both a little longer than Rr; cell 1st M, elongate, exceedingvein Mr*, beyond it; m-cu just before fork of M; distal section of Cu, subequal
to nr-cu and about equal to the spaee along wing-margin between veins Cu, andlst A.
Abdominal tergites brown; sternites paler brown.
H o l o t y p e ,Q , O n o b a ,P a p u a ,a l t i t u c l ea b o u t 2 , 0 0 0f e e t , J u l y , 1 9 3 5 ( K . J . C l i n t o n ) .
Trentepohlio, (Pararnongoma) lueri.fera is allied to species such as 7. (P.)
hanoha,oensisAlexander, ?. (P.) chionopoda Alexander and T. (P.) pusilla
Edwards, differing from all in the venation, especially the subequal veins Rr*r*.
And R.r,, the latter exceeding in length vein R, alone.
TnrNrmonlrl(Morvconr,r) r.oNcrsorosA,n. sp.

Fis. 6.

General coloration dark brown; legs brownish-black, the outer tarsal
segments paling to dirty yellow; tarsi short, especially the posterior pair in male;
all femora with short spines at base; wings strongly tinged with dusky; costal
fringe (d) long and conspicuous; vein Rr oblique, cell R, at margin exceeding
one-half the extent of cell Rr.
3. Length, 5'5-6 mm.; wing, 6-7 mnt. 9. Length, 6'5-8 mm.; wing, 6'5-8 mm.
Rostrum and palpi black. Antennae black throughout; flagellar segments
subcylindrical; longest verticils subequal in length to the segments. Headbrownish-black.
Tholacic dorsum and pleura uniformly dark brown, the dorsopleural region
and pronotum even darker in colour. Halteres dusky. Legs brownish'black, theouter tarsal segments paling to dirty yellow; tarsi short, especially the posteriortarsi in male (femora,8 mm.; tarsi, S'3 mm.), which are much shorter than thefemora; femora with several (12-15) short black spines at base. Wings (Fig. 6)
with a strong dusky tinge, the wing-tip narrowly more darkened in outer radial
field; stigma small and inconspicuous; cells C and Sc more yellowish-brown;veins brownish-black. Costal fringe (J) unusually long aud conspicuous,exceedingin length the width of cell 1st trI, at base; in female, costal fringe dense but
short and normal. Venation: Basal section of Ru relatively long, exceeding two'thirds of Rs; Rr*.*. gently sinuous, exceeding Rs; vein R, much more obliquethan in breuipes, cell R, at margin exceeding one-half the extent of cell Rri Ro*.
only about one'third to one'half Rr; cell 1st 1\{. short, the veins and cells beyond
it correspondingly lengthened.
Abdominal tergites and terminalia brownish-black; sternites brown, thecaudal borders of the segments narrowly darkened.
Holotype, d, Maini, Papua, alt.itudeabout 6,300feet, July, 1935 (K.J. Clinton).
Allotopotype, 9. ParatopotyDes,4 6, ?. Paratypes, L 6, 1 t, Kone, Papua, altitude
about 5,000feet, July, 1935 (K. J. Clinton).
By my key to the regional species of Trentepolr,lia (Pnoc. Lrnrv. Soc. N.S.W.,lx, 1g35,66), the Bresent fly runs to Trentepohl,ia (Mongoma) breuipes Alexander(New Britain, New Guinea), which shows several features in common, as thedarkened femora and tibiae and the shortened tarsi, but is very different in theBY C. P. ALEXANDEB.

L27

pattern and venation of the wings and in the short costal fringe of the male sex.
In the long costal fringe of the male, the present fly agrees with ?. (M.) costofimbriata Alexander (New Britain), a species with entirely difrerent leg-patteru.
TsnNrEpoELrA (Pr.nsrouonooue) NovAE-BBrrrANrAE
Alexander.
Pnoc. Lrnn. Soc. N.S.W., lx, 1935, 66-6?.
1 t , M ai n i , P a p u a ,a l ti tu d e a bout 6,300feet, July, 1935 ( K.J. Clinton) .
TnsNrEroELrA (Tnrnrneosrre) prcrrprNNrc Bezzi.
Philipptitte Journ. ,Sci.,xii, D, 1917, 115.
Inawi, Papua, July, 1935 (K. J. Clinton).
Gorvouvre (Lnorrr,rrs)
JUBATA,n. sp. F''ig. ?.
General coloration of mesonotum dark gr€y, the scutellum light yellow, witha dark spot medially at base; antennae with pedicel large, bright orange; mesopleura dark blown, scarcely variegated by brighter; knobs of halteres dusky;
legs black; wings with a strong brown suffusion, the costal border narrowly
yellow; macrotrichia of veins numerous! Sc, ending opposite origin of Rs;
abdomen black.
9. Length about 4.5 mm.; wing, 4.2 mm.
Rostrum and palpi black. Antennae with scape obseure orange; pedicel
enlarged, bright ol'ange; flagellum black throughout; flagellar segments oval, theouter ones more elongate-oval; terminal segment shorter than the penultimate.
Head grey.
Pronotum above obscure brownish-yellow, the pronotal scutellum brighter
yellow. Mesonotal praescutum and scutum dark gr€y, the pseudosutural foveae
tlark; humeral and lateral portions of praescutum not or scarcely brightened:scutellum lisht yellow, with a darkel median spot at base; mediotergite grey.
Pleura dark brown, scarcely variegated by brighter. Halteres obscure yellow,
the knobs dusky. Legs with the coxae yellowish-brown; trochanters a little
briglrter; remainder of legs black. Wings (FiS. 7) with a strong brownish
suffusion; cells C and Sc yellow. llacrotlichia of veins relatively abundant,
including a series on distal two-thirds of vein lst A and a few on outer end ofvein 2nd A. Venation: Scl ending opposite origin of Rs, Sc, close to its outer
end; Rs long, arcuated; basal section of vein Ro long, approximately one-half r-m;
m-cu shortly before the fork of M.
Abdomen black, the bases of the terminalia brownish-black; cerci horn-yellow.
Holotype, t, Onoba,Papua, altitude about 2,000feet, July, 1935 (K. J. Clinton).
Gonont.yia(Lipophleps) iurato is generally similar to speclessuch as G. (L.)
fijiensis Alexander and G. (L.) kertesziarroAlexander, yet is very distinct in alldetails of coloration, especially of the legs, wings and abdomen.