Reprinterl frorn ButtwtIN oF TrTn BRoor<r,vwENToMoLocrcAr,SocrETY,
Vol. XXV, No. 2, pp. 71-77. April, 1930

NEW

CRANE_FLIESOR INSUFFICIENTLY-KNOWNFROM THE NEARCTIC REGION.
(Tipulidae, Diptera.)
Penr I.
Bv Cnanrns P. Aroxewlnn, Amherst, Mass.l

The new speciesdiscussedat this time were for the most part
included in interesting collectionssent to me for identification byDr. Mortimer D. Leonard and Mr. Millard C. Van Duzee. A
further addition to the paper was contained in the Canadian National Collection, sent to me through the kind interest of Dr.
James McDunnough. I wish to express my thanks and appreciation to the above-namedentomologists for their continued cooperation in studying the Tipulid flies.
Limonia (Dicranomyia) erostrata n. sp.
General coloration ochreous; rostrum and first segmentof
antennae yellow; mesonotal praescutum with a median brown
stripe; male hypopygium with the dorsal dististyle a stout
pale rod that narrows gradually to an acute blackened tip;
ventral dististyle a fleshy lobe that lacks a rostral prolongat10n.
Male.-Length about 6.5 mm.; wing 6.6 mm.
Rostrum light yellow; basal segments of palpi yellow, theouter segments infuscated. Antennae with the basal segment and sometimes the second light yellow; flagellum darkbrown; flagellar segments oval, outer segments narrower.
Head ochreous,the vertex more infuscated.
Mesonotum ochreous, with a brown median stripe that is
subobsoleteand more or less bifid behind, the lateial stripes
less evident; scutal lobes brownish gray. Pleura ochreous
yellow. Halteres destroyed by pests. Legs with the coxae
and trochanters yellow; femora yellow, the tips slightly
darkened; tibiae obscureyellow, darkened at tips; tarsi passing into dark brown. Wings light yellow, the stigma lacking; veins darker yellow. Venation: Sc, ending opposite
origin of Rg Sc, some distance from its tip ; m-cu very
oblique, at the fork of M, subequal to the distal section ofCw.t.
l Contribution from the Department of Iintomology, MassachusettsAgricultural College.

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Aprit,1e30 Bu,Lletin of tlrc Llrooltlyn Entonr,ologicalSociety 73

Abdomen, including the hypopygium, ochreous. Male
hypopygium with the caudal margin of the tergite deeply
notched, the lateral lobes conspicuously setiferous. Basistylewith the ventro-mesal lobe relatively small, placed near thebase of the style. Dorsal dististyle a very stout, broad-based
rod, pale at base, gradually narrowed to an acute blackened
tip. Ventral dististyle without a rostral prolongation, appearing as a pale, gently arcuated lobe that is a little longer
than the dorsal dististyle, the surface with numerous scatteredsetae that are longer and more abundant at apex of style.

Habitat: California. Holotype: d1, Mill Valley, Marin Co.
Ma r ch 13, 19z6 ( M . C. Van Dr nee) . Typ. in t he aut hor 's collection.

Habitat: Utah.
Holotype: ;f, Saltair, May zr, 19z6 (M. C. Van Duzee).
Paratopotypes, r male, t sex?, broken. Type returned to Mr.
Van Duzee.
I cannot identify this species with any of those described byDoane. The male hypopygium has the same peculiar structure
found in L. (D.) si,gnipennis(Coq.).
Tricyphona simplicistyla n. sp.
Male.-Length about 14 mm.; wing 13.5 mm.
Close to T. constans (Doane), differing especially in thestructure of the male hypopygium.
Antennal scape dark brown, the flagellum yellowish brown.
Head pale, the vertex with a dusky area on either side of themedian line. Praescutum almost uniform ochreous, thescutum conspicuously darker; scutellum pale; postnotal
mediotergite pale, margined laterally with brown. Legs yellow, only the terminal tarsal segments a little darkened'Wings with the pattern quite as in consta'ns. Venation: Rs
in alignment with Ru; a supernumerary crossvein in cell R'
placed about its own length before Rr; cell rst M, closed.
Male hypopygium with the outer dististyle broad, the apex
with abundant spinous setae. Inner dististyle a slender simple rod. Interbasal processes appearing as elongate, nearlystraight rods, the tips acute, before apex with a small lateralsplne.
T. constans has the outer dististyle very narrow at apex,
with relatively few spines. Inner dististyle conspicuously
bifid, the outer arm set with two powerful black spines.
It is possiblethat the supernumerarycrossveinin cell R, is
not a constant character but it is identical in both wings ofthe type.

Rhaphidolabis (Rhaphidolabis) vanduzeei n. sp.
General coloration gray, the praescutum with three darkbrown stripes ; antennae black throughout; wings whitish
subhyaline, with a conspicuousdark brown stigma; caudal
margins of abdominal segments narrowly pale; malehypopygium with the dorsal interbase bilid at apex.
Male.-Length about 5.5 mm.; wing 7.2 r:nm.
Rostrum and palpi black. Antennae black throughout, r3segmented;flagellar segmentsshort-oval, gradually decreasing in diameter outwardly. Head gray, the center of thevertex slightly infuscatecl.
Mesonotum light gray, the praescutum with three conspicuousdark brown stripes; posterior scieritesof mesonotum
more nearly immaculate. Pleura gray, the dorso-pleuralregion more ochreous. Halteres pa1e,with darkened knobs.
Legs with the coxae gray; trochanters brown; remainder oflegs dark brown, the tarsi passing into black. Wings whitish
subhyaline, the stigma conspicuous, dark brown; a weak
dusky clouding along the cord; veins dark brown. Venation: Rs short, angulated at near midlength; Ro*u subequal
to the basal sectionof Ru; R, oblique, joined to R' near outer
end ; cell M, present ; ?n-cu shortly beyond f.ork of. M.
Abdominal segments dark brown, the extreme caudal
margins of the segments pale; hypopygium dark. Male
hypopygium much as in caywga; dististyle more slender;
dorsal interbase wider than in caywga but narrower than inrubescens.
i{abitat: California. Holotype: d, Mt. St. Helena, May rz,ry26 (Lt. C. Van Dwzee). Typ" returned to Mr. Van Duzee.
I take great pieasure in dedicating this speciesto Mr. Millarcl
C. Van Duzee, to whom I am indebted for many kindly favors
in the past. Rkaphidolabis aondu,zeeimuch resembles.R. stigma
Alexander but is really more closely allied to R. caywga Alexander, from which it is told most readily by the conspicuous darkbrown stigmal spot.
Eriocera alberta n. sp.
General coloration black, light gray pruinose; antennac(?) 7-segmented,black; knobs of halteres white; wings/

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Aytrit,1e30 Bulletin of the Broolilyn Entomological Society 75

white, with grayish brown seamsto the veins; Rs angulatedand spurred at origin; cell R, deep, parallel-sidedfor morethan two-thirds the length ; cell rsf M rectangular; m,-cu at" than the distal secor close to the fork of M ; nt-ctr, shorlertion of Car; ovipositor with fleshy valves.
Female.-Length about Z mrn.; wing 9.2 ntm.
Rostrum black, sparsely pruinose. Antennae 7-segmented,black, the scapal segments slightly pruinose; first flagellar
segment longer than the succeeding two taken together;terminal segment oval, about one-half the length of thepenultimate. Head broad, the vertical tubercle low; frontand sidesof anterior vertex light gray, the remainder of headdark gray.
Mesonotal praescutum light gral, with four blackish
stripes, the intermediate pair only narrowly separated;scutum gray, the centers of the lobes darkened; scutellum
and postnotum dark, gray pruinose. Pleura gray. Halteresdusky, the knobs white. Legs with the coxae and trochanters
black, pruinose; femora brown, the tips broadly blackened,most extensively so on the short fore legs; remainder oflegs black. Wings white, with a distinct grayish brown pattern that appears as narrow seams along the veins, including
the origin of Rs, Scr, cord and outer end of cell rst M";
stigma and fork of Rr*r*n; small paler circular clouds at endsof longitudinal veins; a circular cloud near center of cellznd IVI basal longitudinal veins with dark seams, most evi"; vein Znd l; veins brown. Venation: Sc, endingdent on
shortly beyond the fork of Rs, Sc, a short distance from its
tip ; Rs relatively short, angulated and spurred at origin;
Rr*r*n more than one-half longer than Rr*, I R*z z little longer
than Rr*r; cell R, relatively deep, the enclosing veins parallel
for about three-fourths their length: cell rst M, rectangular;
nt-cu at or just before the fork ol M; m-cw considerably
shorter than the distal sectionof Cur.
Abdomen black, sparsely pruinose, the sternites moreheavily so; subterminal tergites polished black; ovipositor
with fleshy valves. Tergal plate pruinose, with a V-shapednotch; sternal valves sheathing, dusky at base, the tips
broadly light yellow.

(the female of longicornaswith rr segments), coloration of thewings and the venation. It seems highly probable that the maleof the present specieswill be found to have short antennae.

Habi,tat: Alberta. Holotype: S, Lethbridge, June 24, rgzg (1.
II. Pepper). Typ" in the Canadian National Collection.
Eriocera alberta is readily tolcl from E. longicornis (Walker),the only other describedNearctic specieswith the fleshy type ofovipositor above described,by the number of antennal segmentstIi

Il[,'Rhabdomastix (Sacandaga) leonardi n. sp.
General coloration light yellow, handsomely patternedwith biack, including three conspicuous stripes on thepraescutum; vertex with a conspicuousblackenedarea.
Fem,ale.-Length about 7.5 mrn.; wing 6.5 mm.
Described from an alcoholic specimen.
Rostrum yellow, slightly darkened above; palpi very reduced. Antennae with the basal segment light yellow, thesecondand third segmentsblack; remaining segmentsbroken;
first flagellar segment smaller and scarcely longer than thesecond scapal. Head light yellow, the vertex with a broad
black mark extending to the posterior margin, bordered on
the posterior vertex by more reddish.
Mesonotal praescutum light yellow with three conspicuous
black stripes, the median stripe weakly bifid behind and ending far before the suture; lateral stripes crossing the suture
onto the scutal lobes; pseudosutural foveae black; medianregion of scutum blackened; scutellum yellow; postnotal
mediotergite yellow, weakly darkened medially, the posterior
portion with two approximated black oval areas. Pleura
yeilow, handsomely variegated with small dark marks, including the propleura, anepisternum,ventral sternopleurite andmeron, and ventral pleurotergite. Halteres pale, the knobs
light yellow. Legs with the coxae yellowish brown; trochanters pale; remainder of legs broken. Wings grayish
subhyaline, the stigma barely indicated; veins black, theprearcular, costal, subcostal and radial veins light brown.
Venation: Sc, ending about opposite two-thirds the lengthof Rs, Sc, far from its tip, Sc, alone about one-third Rs,' R.
short and nearly perpendicular; distance on costa between
Rr*, and R, a little longer than the latter; Rn gently arcuated,
a little shorter than the petiole of cell R".
Abdomen light broi'n, the tergite; vaguely darkened
basally, the genital segment more darkened medially.
Habitat: Montana. Holotype: alcoholic t, Bozeman, July :'9,t9z9 (M. D. Leonard). Typ. in the author's collection.
Rhabdomastir leonardi is named in honor of my old friend andco-worker on the Tipulidae, Dr. Mortimer D. Leonard. Thespeciesis very distinct from the other describedAmerican species/Society Tot^xxr
Bultetin of the Brooklyn Entom,ologicalAprit,1e30 Bulletin of the Brooklgn Entomological Soci,ety 77of the genus. The highly contrasting yellow and black pattern of
the head and thorax is somewhat suggestiveof that found inmany species of the Tipuline genus I{ephrotom.a. It is possiblethat in fresh specimensthe colors as above described may besomewhat dulled by a pollinosity or pruinosity not eviclent in thealcoholic type.

four small appressed lateral teeth; inner arm short andstocky, set with conspicuous teeth.

76

Erioptera (Ilisia) bispinigera n. sp.
General coloration obscure yellow, the praescutum withthree nearly confluent dark brown stripes; pleura dark,striped longitudinally with pale; halteres pale yellow; wings
with cell M, open, vein znd. A short and straight; malehypopygium with the inner dististyle bidentate on lateral
margin; gonapophyses entirely blackened, very unequaily
bifid.
Male.-Length about 4.5 mm.; wing about 4.5 mm.
Described from an alcoholic specimen.
Rostrum and palpi brownish black. .Antennae with thescapal segments dark brown, the flagellum paler; flagellar
segments oval. Head chiefly dark brown.
Mesonotal praescutum yellow with three nearly confluent
dark brown stripes; pseudosuturalfoveae dark; scutum obcure veliow. each lobe with the center dark brown; scutellum
obscure yeliow; postnotal mediotergitedark brown predially,
the lateral margins narrowly pale yellow. Pleura chiefly
dark brown, with a broad conspicuous yellowish longitudinal
stripe extending from behind the fore coxae to the base of
the abdomen; dorso-pleural region pale. Halteres relatively
elongate, pale yellow throughout. Legs with the fore coxae
brownish yellow, the other coxae pale yellow; trochanterspale yellow; remainder of legs broken. Wings grayish subhyaline, the diffuse stigma vaguely darker; veins brown.
Venation: veins beyond the cord entirely as in the subgenus
Erioptera; cell M, open; vein znd' A short and straight, as inIli,sia.
Abdomen dark brown, the pleural region pale; hypopygium
obscure brownish yellow. Male hypopygium with the outerdististyle an expanded black sttucture, the dilated apicalportion with microscopic denticles. Inner dististyle shorter,blackened, the outer or lateral margin with two conspicuous
black spines, one at near midlength, the second nearly apicalin position. Gonapophyses appearing as entirely blackened
plates, the basal half dilated, very unequally bifid, the outerarrn produceclinto a slender rod, the tip acute, with three or

Habitat: Montana. Holotype: alcoholic S, Bozeman, July tg,rgzg (M. D. Leonard). Type in the author's collection.
Erioptera bispi.nigerais very doubtfully a member of llisia, inwhich subgenus I am placing it merely on the course of vein znd
A. It may be remarked that all of the subgenera of. Erioptera arc
highly artificial and becoming more so with the constant accessionof new material.
Erioptera (Ilisia) laevis n. sp.
Male.-Length about 4.2-4.5 mm.; wing 5-5.5 mm.
Very similar in the pattern of the wings and legs to E.
armillaris, differing especially in the structure of the malehypopygium.
General coloration of body darker, especially the thoracic
notum which is dark grayish brown instead of yellowish
brown. Male hypopygium with the outer dististyle relatively
small, the apex simply clavate, not emarginate on outermargin. Gonapophysesappearing as simple black horns
that are slender, nearly straight, the margins quite smooth.
E. (1.) arncillaris Osten Sacken has the outer dististyle
very large, the head with the outer margin conspicuously
emarginate to produce an obtuse lateral point. Gonapophyses
stout, abruptly narrowed to an acute point, the outer margin
before this apex microscopicallyspinulose.
E. (1.) indiunensisAlexander has the outer dististyle smalland slender, the apex only feebly dilated, entire. Gonapophyses appearing as curved slender hooks, the long tips
acute, the margins entirely smooth.

t,

Habitat: Northeastern North America. Holotype: J, Chesterfield Gorge, Massachusetts,altitude 85o feet, August z, rgz9 (C.
P. Alerand,er). Paratypes, J, Amherst, Massachusetts,July r8,r9z8 (C. F. Clagg); {,Hazleton, Pennsylvania,June zg-July 18,tgro ('tV. G. Dietz). Type in the author's collection.
There can be no doubt but that there are three distinct species
confused under the name of armillaris. The wing-pattern is almost identical in all three but the structure of the male hypopygium is very different in the various species.