raJ i iT .

NEW AND LITTLE KNOWN TIPULIDAE
FROM THE MARQUESAS*By
Cnenr.EsP. Ar,txeNDER
DEpenrrvrENToF ENrolrorccv. Zoor.ocy,AND GEolocv, MessecsusETts Srertr Cor.recn.

INTRODUCTION

The speciesdiscussedin this paper were submittedto me for determination by the collectors of most of the specimens,E. P. Mumford andA. M. Adamson, of the Pacific Entomological Survey.
TRIBE ERIOPTERINIThe vast majority of the Eriopterine Tipulidae in the more remote
Pacific islands pertain to the subgenusLipolthleps of the genus Gonornyia
Meigen. In this series from the Marquesasare included no fewer thanfour species of this subgenus, all, as known, being restricted to theMarquesas.
Key to MarquesanSpeciesof Lipophlepsl.

General coloration metallic blue-black: wings 1'ellowish,
conspicuouslypatternedwith brorvtr; size large ( rving,..--metallesceneEdwards
male, more than 6 mm.)C,eneralcoloration reddish brown to brown, variegated withyellow; wings unmarked, except for the stigmal area
when present; size small (wing, male, not exceeding
5 mm.).....-..............2ngw species
Tips of all femora broadly and conspicuouslyblackened.........-..adamsoni
Femora uniformly blackened,or pale with a brown subterminal ring ..-.-....

...-.-....--.--...---..3

Antennae,legs, and knobs of halteres brown; wings strongly............_mumfordi
new species
sufiusedwith b1ackish............-.
Antennae with the scapal segmentsconspicuouslylight-yellow; knobs of halteres yellow, femora brownish-yellow,
with a slightly darker subterminalbrown ring; wings-.--.-....marqucsana
new species
brownish gray -.-.--..* Pacific Entomological Survey Publication I, article 7.

l8z l

88

BerniceP. BishopMuseum-Bulletin 98

Edwards 2a has recorded still another species of. Li,ltophleps f.rom Hivaoa.
His specimen was in too fragmentary a condition to determine, but is certainly different from the other speciesknown from the Marquesas.
Gonomyia (Lipophleps) metallescens Edwards (fig. 26, a, e).
Gononcyia(Lipophleps) metallescensEdwards: Ann. M"g. Nat. Hist.
(9) , zo, p. 24o, 1927.
Male
Length about 5.5-6 mm.; wing, 6.5-7.5 mm. Palpi dark-brown. Dorsal sternopleurite shimmering blue-gray, its ventral portion polished black. All legs with femora
chiefly blackened. with a paler annulus beyond nridlength; tibiae and tarsi chiefly
blackened. Wings (frg. 26, a) with the prearcular region broadly and abruptly lightyellow; cubital and anal cells beyond base almost uniformly infuscated; remainder ofwing-pattern about as in the type. Venation: Sc, ending opposite or slightly beyond theorigin of Rs. Abdomen black, the hypopygium a trifle paler. Male hypopygium (fig.
26, e) with the lateral arms of the tergal plate produced, each bearing two branches,
one being a flattened black paddle, the other, an acute spine; oll one side the spine exceeclsthe obtuse arm, and on the opposite side the obtuse arm exceeds the spine. Two
dististyles,the outer heavily sclerotized; inner clististylemuch expanded at base.
The ochreous abdominal rings described for the type female are little apparent in
the material at hand, being most evident on the subterminal tergites.

Marquesas: Eiao, uplands toward north end, east side, elevation 19oo
feet, September2e, rg2g, on Hibiscns tilioceus,2 males (t allotype), tfemale; Vaituha, near sea level, September2Q, rg2g, at light, r female, andOctober2, rg2g, r male; Adamson.
The male sex has never been described,and one of the present specimens
is definedas allotype. This striking crane-fly was describedfrorn two females
taken on Nukuhiva, Marquesas.
The hypopygium is very different in structure from that of the twoother species of the flovida.per group rvhose males are known (dicranuraEdwards and labi.dura Edwards). In these latter species,the basistyles arestrongly produced at apices, the lobes thus formed heavily blackened andterminating in a small terminal spine; a single dististyle. In the present
speciesthe basistyleis unmodified and there are two dististyles,as described.

Gonomyia (Lipophleps) adamsoninew species(fig. 26, b, f)'General coloration of mesonotum light reddish-browu, without distinct markings;
pleura -pale, blackened dorsally, the ventral portion with a silvery longitudinal stripe;with tips broadly and conspicuously blackened; wings pale brownish-yellow,i..or"the stigma and axillary region faintly darkened; a small brown cloud on anterior cord;
Sc short, Sc, ending just before origin of Rs; male hypopygium with the phallosome
symmetrical, each arm bearing a lateral branch beyond midlength, the branches andarms clothed with an abundant delicate pubescence.
grEdwards, F. W,, Diptera Nemafocera frorn the South Pacific:
20, p. 24O, 1927.

Ann. \[ag.

Nat. Hist., (g)')l urquesenInsects-l.

89

Frcunc 26. Marquesan speciesof Lipophleps: a, Gonomyia (Lipophleps) ntctallescens Edwards, wing; b, Gornmyia (L.) a^damsoninew species,wing i c, Gonomyia(L.) mumford,i new species,wing ; d, Gonomyia (L.) m,orquesononew species,wing;e, Gonomyio (L.) metallescezsEdwards, male hypopygium; f , Gonomyia (L.) ad.amsoni
new sp€cies,male hypopygiumt g, Gonomyio (L.) mumfordi new species,male hypopygium ; h, Gonomyia (L.) rnarquesm,onew species,male hypopygium. ba : basistyle;id : inner dististyle, od,-- outer dististyle; I : phallosome; gt : 9th tergite.

9oBerniceP. Bi,shoplltfussum-Bulletin 98

Male: length about 4.3-4-4 mm. ; wing, 4-4.2 mrn. Female : length about 5 mm. ;wing, 4.2 mm.
Rostrum and palpi brownish-black. Antennae with the scapal segments light-yellow,
flagellar
thesegments a little more testaceous,in the male with elongate verticils. Headyellow,lightthe center of the vertex restrictedly darkened.
Pronotum yellow. I\fesonotal praescutum and scutum reddish brown, without distinct markings, the lateral margin of the praescutum paler, centers of scutal lobes
slightly darkened; scutellum testaceous; postnotal mediotergite pale testaceous, thecephalic portion a little darker. Pleura blackened dorsally, the ventral portion with aconspicuous silvery longitudinal stripe extending from behind the fore coxae to thebase of the abdomen, passing beneath the root of the halteres; ventral sternopleurite
reddish-brown; the dorsal darkening includes the dorso-pleural membrane, propleura,anepisternum and pteropleurite, the pleurotergite pale. Halteres light sulphur-yellow.
Legs with the fore and middle coxae yellowish testaceous,the posterior coxae slightly
more darkened; trochanters yellow; femora yellow, the tips broadly blackened, theamount subequalon all legs; tibiae and tarsi yellow, the outer tarsal segments slightly
infuscated. Wings (frg. 26, b) pale brownish-yellow; stignra and axillary region faintly
darkened; a more conspicuousdarker broken cloud on anterior cord; veius pale-brown,
the anterior cord dark-browrr. Venation: Sc' ending just before the strongly arcuatedRs, Sc, close to its tip; anterior branch of Rs nearly straight; cell rsf M: closed; m-cu
close to the fork of. M.
Abdominal tergites aud sternites obscure yellorv, narrorvly blackened laterally.
Male hypopygium (fig. 26, l) with two dististyles, the outer a curved, simple, glabrous,
blackened rod; inner style much smaller, fleshy, with abundant erect setae, including
two stouter, more fasciculate setae at apex. Phallosome symmetrical, each arm slender,
densely clothed with a delicate pale pubescence,the tips acute; on outer margin beyond midlength with an acute lateral spine arising from membrane, the basal half ofthe spine with numerous setulae. Aedeagus terminating in an acute recurved spine.

Marquesas: Eiao, Vaituha, near sea level, October z, rgzg, at light,
holotype male, allotopotype female; September 29, tgzg, paratopotype male,Adamson.
This very distinct speciesol Gonornyiois named in honor of the collectorof the material, A. M. Adamson.

Gonomyia(Lipophleps)mumfordi new species(fig.26, c,g).
General coloration dark-brown, the tholacic pleura rvith a more or less distinct
ventral yellow stripe; rostrum and antenuae black; legs blackerted; knobs of halteresclarkenecl;wings with a uniform strong blackish tirrge; vein rsf I with macrotrichia
distributed throughout the entire length.
l\{ale: length about 3.2 mm.; wing, 4 mm.
Rostrum, palpi, and antennae black, the outer segmetrts of the anteunae slender,
with long conspicuousverticils. Head dark-brown, restrictedly variegated with yellow.
I\fesonotum chiefly dark-brown, the median region of the scutum, together with theadjoirring portions of the praescutum,obscure 1'ellow; scutellum dark brown; postnotal
mediotergite dark-brown. Pleura dark-brown, especially the dorsat half, the ventral
half traversed by a more or less distinct yellow longitudinal stripe ; ventral sternopleurite
paler brown than the dorsal region. Halteres browr. Legs with the coxae andtrochanters pale-brown; remainder of legs dark brorvn to black. Wings (fig. 26, c)r,r'ith a uniform blackish tinge, the oval stigma very slightly darker; veins brown.
Costal f ringe relatively long and conspicuous;macrotrichia on veiu rsf A f.or the entirelength. Venation: Sc, ending just before the origin of Rs, 5c, a short distance f rom

Marqaesan Insects-[.

9l

its tip; Rs very strongly arcuated; basal deflection of Ro long; m-cu close to the fork
ol M.
Abdomen, including the hypopygium, dark-brown. Male hypopygium (fie. 26, g)with the outer dististyle simple, glabrous. Inner dististyle very small, terminating in
two fasciculate setae,with additional normal setae scattered over surface. Phallosome
consisting of two divergent apophyses,pale, with the acute tips narrowly blackened,
before the tips on mesal face with an extensive pale membrane. Aedeagus elongate,
curved, at apex split into two acute, slightly divergent spines; immediately beneaththe aedeagusand closely connectedwith it for about one-half of its length is a second
rod that is drawn out into a long pale point.
elevation 37oo feet, March 4, 1930,
miscellaneous beating, holotype male; March 3, 1930, paratopotype male,
Marquesas:

Hivaoa,Matauuna,

Mumford and Adamson.
I take great pleasurein naming this interesting Gonornry in honor ofthe Director of the PacificEntomologicalSurvey, Mr. Edward P. Mumford.
The speciesis readily distinguished from all other small regional speciesof
Lipophlepsby the almost uniform dark color of the body, legs,halteres,andwings. The paratype is more extensivelydarkenedthan the type, the yellow
pleural stripe being obliteratedor nearly so.
Gonomyia (Lipophleps) marquesananew species(fig. 26, d,,h).
Rostrum black; antennal scape light-yellorv, flagellum black; scutellum dark-brownbasally, the apex broadly yellow; thoracic pleura yellow, conspicuously striped longitudinally with dark-brown: knobs of halteres light-yellow; femora brownish-yellow,
with a subterminal pale brown ring; wings brownish-gray, the prearcular and costal
regions pale yellow; stigma pale-brown, oval ; macrotrichia on vein tst A for entire
length.
Male: length about 3.5-3.7 rnm. ; u'ing, 4.2-1.5 mm. Female: length about 4.5 mm. ;wing, 5 mm.
Rostrum and palpi black. Antennae with the scapal segments clear light-yellow,
flagellum
black; flagellar segments of nralc rvith elongate verticils, as in the subthe
genus. Head light-yellorv.
Pronotum yellow dorsally, darker laterally. [Iesonotal praescutum brown medially,
paling to yellowish brown on sides, the humeral region and lateral pretergites clear
light yellow; pseudosutural foveae reddish-brown, but little conspicuous against thisground-color; scutal lobes dark-brown, obscure yellow on median portion; scutellum
dark brown medially at base, the apex broadly yellow; postnotal mediotergite yellow,
the cephalic margin darkened, the caudal margin with two paler brown spots. Pleura
pale whitish-yellow, conspicuously variegated with darker, including a dorsal longitudinal stripe extending from the cervical sclerites to the abdomen, passing beneaththe halteres; a much pater brown area orr the ventral sternopleurite. Halteres with thestem testaceous, the knobs c\ear \ight ye\\ow. \,egs with the coxae pe\e ye\\ow, thebasal portions of the mid-coxae a trifle darkened; trochanters yellow, femora brownish
yellow, with a slightly darker subterminal browr.r ring ; tibiae brownish black; tarsi
black. Wings (fi9. 26, d) with a brownish-gray tinge, the prearcular and costal regions
pale-yellow; stigma pale-brown, oval, very faintly indicated and ill-defined; veins brown.
i\{acrotrichia on vein rst A for the entire length; costal fringe and macrotrichia ofveins long and conspicuous. Venation: Sc short, Sc, ending opposite origin of Rs.
Abdominal tergites obscure yellowish-brown, the sternites a little paler; hypopygium
yellorv. \,Iale hypopygium (fie. 26, h) with tu'o dististyles, the outer one a simple

IBerni,ceP. Bisrnf Maseut*Bulletin 98

gtotly curycd rod, the inner small and fleshy, terminating in two fasciculate s€tae,with other smatler setae, chiefly marginat in distribution. Elements of pfrallosome
symctrical, consisting of two fattened sword-shapeddusky blades, with two moresleoder divergmt rods. Apcx of aedeagusa strort curved hook.

Marquesas: Hivaoa, Mount Temetiu, elevation3,9oo feet, December30,
rggq at light, holotypemaleand numerousparatopofpes, malesand females;
Decambe,2i, 1930, at light, numerous paratopotypes,males and females,
H. Tauraa; Matauuna,elenation3,7oo feet, March 4, L93a,by miscellaneous
beating, allotype female, Mumford and Adamson.
One paratJrpemale has one wing with cefl f$ M, oprcnby the atropty
of. fit.