:

1rz"):

Ptom the Proceeil,ings of th,e Linnean Society of New Boutlt,Walee,Vol. la, Parts 1-2, 1935.

TEE DIPTERA OF THE, TERRITORY OF' NEW GUINEA.
TAMILYII.

TIPULIDAE.

By Cnenr.ss P. ALoxANDEB,Massachusetts State Coilege, Amherst, Mass., U.S.A.
(Com.nunicated,by Frank H. Taylor,)(Twenty Text-flgures.)lRead 2?th Nlarch, 1935.1

lfssued 15th May, 1935.1

a-i .r-.

: t,':' ';" r'$

Front, the Proceed,ingsol the Linnean Society ol New Boutlt Wales,
Vol. lr, Parts 1-2, 1985.

THE DIPTERA OF' THE TERRITORY OF NEW GUINEA.
FAMILYII.

TIPULIDAE.

By Cuenrps P. Ar,nxanpnn, Illassachusetts State College, Amherst, Mass., U.S.A.
(Comm,u,nicateilbU Eranlc H. Taglor.)(Twenty Text-flgures.)[Read 27th March, 1935.]

The very interesting series of Tipulidae discussed herewith was collectedpart by Mr. Frank H. Taylor, a small number of specimens being
greaterintaken by Mr. S. V. Bayley and by Dr. H. Champion Hosking. The majority ofthe specimens were taken at Keravat, Rabaul, and at an altitude of about 1,000
feet on Mount Toma, all located on the Gazelle Peninsula of New Britain. Fewer
specimens were taken on Makada Island, of the Duke of York Group, east of
the Gazelle Peninsula. The types and uniques have been returned to Mr. Taylor
for incorporation in the collection of the School of Public Health and Tropical
Medicine of the University of Sydney. I wish to express my very deep gratitude
to the Director, Professor Harvey Sutton, of the School of Public Health, and to
my good friend and co-worker on the Diptera, Mr. Frank H. Taylor, to whom f
express my deepest thanks for many appreciated favours.
TrpurlNer.
Crnx--rcnoscnlrscoNsprcABrlrsANGUSTTLTNE.\TA,
n. subsp.
?. Length about 25 mm.; wing, 30 mm.
Characters as in typical conspicabilis Skuse (North-eastern Australia),differing in slight details of coloration, especially the conspicuous brown medianpraescutal vitta, which is yellowish in the typical form; in the present case,it ismuch darker than the brownish-grey intermediate stripes; lateral praescutal
stripes narrowly bordered by yellow. Pleura yellow, without dorsal brown stripe;
fore coxae and ventral sternopleurite restrictedly and vaguely darkened. Abdomen'with the tergites unbrightened, except laterally.
Holotype, 9, Rabaul, January, 1933 (F. H. Taylor).
It is possible that the present fly represents a valid species, rather than arace. The antennae have the first flagellar segment elongate, exceeding one-halfthe second; lower faces of flagellar segments but little protuberant, with a pairof short setae at near midlength.
CrnrecnoscElrs cLoBULrcoRNrs,
n. sp.
Belongs to the untbrin zs group; flagellar segments, J, very conspicuously
produced on ventral side into nearly globular lobes, each tipped rvith a long seta;
pleural stripe distinct; ventral pleurites and coxae spotted and streaked with pale
brown.

52

DIPTERA OF THE TERRITOIIY OF :{EW GUINEA.

1I,

3. Length about 21 mm.; wing, 25 mm.
Frontal prolongation of head brownish-yeilow above, darker brown beneathand on sides; nasus long and conspicuous; palpi brolvnish-black. Antennae with
scaDedark brown; pedicel pale yellow; flagellum black; flagellar segmentsbeyond
the flrst very strongly produced to appear almost globular, in degree exceeding
tha.t of Ib.e serrafas group; a long conspicuous seta at apex of protubel'ance.
Head brownish-black,sparsely pruinose behind; front and anterior orbits nat'rowly
ochleous.
Pronotum blackish-grey medially, pale on sides. Mesonotal praescutum withthe -tour usual stripes gr€y, bordered by blackish, the latter especially clear andevident as internal borders to the intermediate stripes; lateral bordet's ofpraescutum broadly brownish-black, margined internally by a narrow yellow linethat forms an outer border to the lateral praescutal stripes; a capillary yellow
median vitta on about the anterior two-thirds of praescutum; posterior inter.
spacesdarkened, concolorouswith the stripes; posterior sclerites of notum chiefly
blo'vn; median region of scutellum yellowish; posterior and lateral margins of
n:ediotergite bloadly and conspicuously ochreous. Pleura ochreous, with alongitudinal brown stripe from cervical region to beneath the wing-r'oot,
becoming more difruse behind; ventral pleurites and coxae with paler brownspots and streaks. Halteres brown, the knobs brownish-black. Legs with coxae
as described; trochanters yellow; femora brownish-yellow, paler ventrally, thetips broadly blackened; tibiae and tarsi uniformly pale yellow. Wings tinged
with blownish; cells C and Sc, together with the stigma, darker brown; outet'
ends of radial cells slightly infumed; m-cu and adjoining parts of Cur, together
with the anterior cord, narrowiy seamed with brown; obliterative at'eas befot'e
stigma and across cell lst Mr; veins brown, except in obliterative areas. Venatiotr:
rn-cu sinuous at posterior end, placed just before midlength of Mr*n.
Abdominal tergites chiefly dark brown, each slightly brightened medially albase; latelal borders broadly, the caudal margins more narrowly pruinose;
stelnites pale yellorv, the intermediate segments a little darkened mediall]', theouter segments uniformly darkened except for narrow pale margins; terntiualia
and its appendrges paler, especially outwardly.
Holotype, J, Rabaul, January, 1933 (F. H. Taylor).
This fly is readily told from regional allies of the untbrinus group by theunusually produced flagellar segntents, which here are fully as developed ns int\e serrat?rsgroup.
Tllull.

Linnaeus.

P,\PUATIPULA, n. Subgen.

Flontal plolongation of head elongate, subequal to the remainder of head;nasus distinct. Antennae l3-segmented; flagellar segments with verticils ilrat
greatly exceed the segments in length. Tibial spurs long and conspicuous;
fot'mula L-2-2. Wings (Fig. 1) rvith Rs unusually short but not tLansver.se,
appt'oximately two-thirds m-cu; Rr*, very long and straight, exceeding trvice m-cu;
Rr*, entilely atrophied or represented only by a short basal spur; vein Rs
elongate, lying unusually close to the costal border of wing, subequal in length
to Rrrr; cell lst M, elongate, its inner end strongly pointed; cell \{, deep; m-cu
uniting with M'*. some distance before its fork, usually at near midlength of thevein. Mact'otrichia of veins beyond cord unusually sparse and scattered, there

BY C. P. ALEXANDEN.

53

being a loose series on Rn*uand Mr; squama naked. Male terminalia. with thetergite separated by a suture from the sternite, fused only at extreme cephalicportion; basistyle fused with sternite. Tergite notched medially. Outer dististyle(Fig. g) armed r*'ith a spinous apical point. Eighth sternite unarmed. Ovipositor
with pointed, sclerotized valves.
(Papuatipula) noaae-brittaniae, n. sp. (Aus'Type of subgenus.-Tipula
tralasian: Papuan subregion).
Other inclutled species: Tiputa (Papuatipu,ta) tliuergens de MeijerQ, T. (P.)
leucosticta Alexander, ?. (P.) meiiereano, new name (for dentata de Meijere,
preoccupied), and f. e.) omissi.nercis(de Meijere).
Most nearly allied to the subgenera Acu,tipu,laAlexander, Indotipulo Edwards,
and Tiptttodila Enderlein; most readily told by the tibial'spur formula, thevenation, and the fundamentals of structure of the male terminalia. This groupof the genus Tipula has proved to be the most widespread and characteristic of
those occurring in New Guinea and will probably be found to include numerous
species when the Papuan fauna is better known. In their general appearance'
the various forms bear a marked resemblance to one another and are best
separated by the characters of the male terminalia.
n. sp.
rvov:rE-BBrrrANrAE,
General coloration of mesonotum obscure yellow, the praescutttm with threesligSgy more fulvous stripes that are indistinctly bordered by darker; wings
with a greyish tinge; both cells C and Sc dark brown, concolorous with thestigma an6 narrow apical border in outer radial cells; seams at anterior cordand on m-cu scarcely evident; petiole of cell M, very short, about one-half m;male hypopygium rvith the outer dististyle a flattened blade, near outer end
produced into a powerful blackened spine.
wing,
J. Length about 15 mm.; wing, 16.5 mm. Q. Length about 22 mm.;
19 mm.
Frontal prolongation of head orange; nasus with conspicuous black setae;
palpi brown, paler at incisures and at outer end of terminal segment. Antennae.wi6 scape and pedicel yellow; flagellum bicolorous, the bases of the individual
segments black, the remainder yellow, the bicolorous nature eontinued to thevel.y end of the organ. Head pale brown, the inner bolder of eye narrowly
yellorv; midline of vertex vrith a narrow dark vitta.
Ground-colour of mesonotal praescutum obscure yellow, with three slightly
more fulvous stripes that are narrowly and indistinctly bordered by darker; medianstripe u'ithout an evident dark median vitta, as in other species of the subgenus;
central portions of scutal lobes chiefly obscttre orange, the mesal portions more
infuscated; scutellum a little infuscated; mediotergite chiefly yellow. Pleura
yellorv. Halteres elongate, brown, the basal fifth of stem yellow. Legs with thecoxae anrl trochanters yellow; remainder of legs dark brorvn; femoral bases
rather narr-o'wly more obscure yellow, the intermediate portions yellowish-brown'
the tips clarti brorvn. Wings (Fig. 1) r-ith a greyish tinge; both cells C and Se
dark brovr-n, concolorous v'ith the stigma and a narrow apical border in cells
Se, and R. to the wing-tip; anterior cord and m-cu very narrowly and incon'sfricuonsllr seamed rvith darker; veins brorvnish-black. Venationi Rr*r entirely
atrophied; petiole of cell M, very short, about one-half m; inner end of cell
1st M, strongly pointed; m-cn at or shortly before M'*,.
Trpur,,r, (Peru.rrrrur,.r)

64

DrprERA oF THE TEnnrrony oF NEw curNEA.

ii,

Abdomen with intermediate tergites having a glabrous basal ring of a greyish
colour, contrasting with the fulvous-orange ground-colour of the tergites; auarrow, transverse yellowish ring immediately caudad of the glabrous ring, thisin turn followed by a darker brown band; sternites more uniformly yellow.
Male terminalia with the tergite (Fig. 9, 9f) bearing a flattenert lobe on eitherside of a small median notch; apices of lobes obliquely truncated and set withsmall blackened spines. Outer dististyle (Fig. g, oil) unusually broad andflattened, the cephalic margin near outer end produced into a powerful blackened
spine. Inner dististyle with the main blade yellow, somewhat reniform, the outermargin with a single to partly double row of long yellow setae that are angularly
bent near tips; at base of blade with two blackened lobes, as shown (Fig. g, id).
Holotype, J, Toma, altitude about 1,000feet, February, lgBB (F.H. Taylor).
Allotopotype, 9.
The nearest allied species is Tipula (Papuatipula) nteijereana, n. name(dentoto de Meijere, preoccupied by Tiputa d,entata Meigen, lg3g), of Southwestern New Guinea, which is somewhat similarly coloured yet differs in thedetails of venation and structure of the male terminalia.
NnpgnotoMA

FUMrscurELL,\TA,

D. sD.

General coloration yellow; occipital brand brown; pronotum entirely paleyellow; mesonotal praescutum yellow, with three dull velvety-black stripes, themedian one with the posterior portion and a median extension on to anterior
portion shiny plumbeous; scutellum of a medium brown, the base narrowly
yellow; femora brownish-yellow; wings strongly tingecl with yellowish-brown;basal five abdominal segments almost uniformly reddish-yellow, the extreme
caudal borders blackened, the outer abdominal segments black.
J. Length, Lz-Lg mm.; wing, 11.8-12.8 mm. Q. Length, 1?-1g mm.; wing,
13-14 mm.
Head yellow; occipital brand brown. Antennae with scape and pedicel yellow;
flagellum black throughout.
Pronotum entirely pale yellow. Mesonotal praescutum yellow, with three dull
velvety-black stripes, the median one with the posterior half and a further medianextension cephalad almost to front border of stripe shiny plumbeous; lateral
stripes dull velvety-black, the extreme antero-lateral ends more plumbeous;scutum pale yellow, each lobe chiefly covered by two confluent dull black areas;
scutellum chiefly median brown, the base narrowly yellor-; mediotergite yellow,
the posterior border with a rectangular brown area that is narrowly bordered bydarker brown. Pleura yellow, variegated by more reddish on the ventral
anepisternum and ventral sternopleurite. Halteres faintly dusky, the knobs
chiefly pale yellow. Legs with the coxae and trochanters reddish-yeliow; femora
brownish-yellow; tibiae brown, somewhat darker outwardly; tarsi passing into
black. Wings (Fig. 2) strongly tinged with yellowish-brown, especially before
the cord, the outer portions of wing without the yellow tints; stigma oval, brorvn;
veins brown, more yellowish-brow-n in the flavous areas. Venation: Cell XI,
sessile.
Basal five abdominal segments almost uniformly reddish-yello*', the extreme
caudal borders of the basal four tergites narrowly darkened, more broadly so infemale; more than outer half of tergite flve blackened; remaining segments ofabdomen black, the dististyles yellowish.

BY C. P. ALEXANDER.

66

Holotype, J, Toma, altitude about 1,000 feet, February, 1933 (F'. H. Taylor).
Paratypes, 1J, with the
Allotype, t, Rabaul, December,1932 (F. H.Taylor).
allotype; 3 i, Keravat, January, 1933 (F. H. Taylor).
Ne.phrotomafumiscutellata is allied to a number of Papuan species that havea somewhat similar pattern on the mesonotal praescutum, these including
lf. d,imid,iota (de Meijere), N. melanura (Osten Sacken) and ff. speculota(de Meijere), all four speciesdiffering among themselvesin details of coloration.
The present fly is closest to .lf. speculata, differing in the strongly tinted wings
and the brown coloration of the scutellum. Of the three species listed above,
N. d,imidiata tlas the scutellum entirely black; ntelanrlra entirely yellowish'red;
speculata dull black, with the base yellow.
MncrsrocnBAFUscaNr (Wiedemann).
1821. Nematocera fuscana Wiedemann, Dipt. Eaot., i, 29.
Widely distributed throughout the Indo-Malayan Islands, as far north as
Luzon, Philippine Islands, eastward into north-eastern Australia.
1 t, Rabaul, December,1932 (F. H. Taylor).
Lraroxrrx-rn.
Lru.oxrrlrt.
(L,rosE)
n. sP.
LruonreFALoATA,
Antennae yellow, the scape black; legs yellow, the extreme bases of tibiae
black; wings with apex narrowly falcate, whitish subhyaline, with two broadcrossbandsand abundant brown dots in most cells; supernumerary cross-veinsin
cells R. and Rr.
6. Length about 9 mm.; wing, 11 mm.
Rostrum black; palpi with basal segments black, the small terminal twosegments pale brorvn. Antennae with scape black; pedicel and flagellum orangeyellow, the narrowed apical half of the terminal segmentbrown; flagellar segments
oval, the outer ones becoming more elongate; longest verticils subequal to thesegments. Head with front silvery; posterior vertex brownish-Brey, with acapillary dark brown median line; eyes contiguous at a single point onanterior vertex.
Pronotum brown. Mesonotal praescutum with four orange-brown stripesthat are narrowly bordered by dark brown, the intermediate pair confluent infront, euding squarely some distance before the suture; lateral stripes withanterior ends bent laterad to margins; humeral region and posterior portion ofpraescutum more whitish pruinose, the latter traversed by two orange-brown
extensions of the intermediate stripes; scutum blackened medially, the lobes
with two dull orange areas that are narrowly bordered by brown; scutellum
rvhitish, with a median brown triangle, the point directed cephalad; mediotergite
chiefly covered.by a dark triangle, its point directed caudad, the lateral angles
more intensely darkened; a greyish area on each posterior portion of mediotergite.
Pleura light grey, extensively variegated by brown, involving all of anepisternum,with smaller areas on sternopleurite, pteropleurite, pleurotergite and meron; afew yellorv setae on sternopleurite. Halteres pale yellow, the knobs dark brown.
Legs with coxae brownish-yellow, the mid-coxae more pruinose; trochanters
yellow; femora yellow, the tips insensibly darker; tibiae yellow, the bases very
narrowly but conspicuorislyblackened; tarsi yellow, the terminal segmentsdarker,
especially the fourth; claws with basal spine and mieroscopic denticles. Wings

DIPTERA OF THE TERRITORY OF NEW GUINEA.

ii,(Fig. 3) whitish subhyaline, with two broad broken brown fasciae and numerous
browq dots in most cells; basal band across proximal fourth of wing, beginningnarrowly at R, widened behind, broadest in cell M, again narrowed where crossing
the cubital and anal cells, reaching posterior margin in cell 2nd A and again
at end of vein 2nd A; outer band very broad, virtually traversing the wiug andextending from before cord to beyond the level of outer end of cell lst M', incell R6 extending to beyond the supernumerary cross-vein; the latter bandvariegated by ground-areas in base of eell Ro and more extensively in cell 1st M';
a more fulvous arm extends from the outer band across the radial field to cellRr; tips of longitudinal veins at margin extensively seamed with fulvous-yellow;
all interspaces of wing, except near apex, with abundant brown dots, in cases
more crowded and becoming confluent; veins yellow, more reddish-yellow incertain places, especially the outer radial field. Wings of unique conformation in
the genus, falcate, the tip narrowed to a point at end of vein Rr*r, thence strongly
incised. Costal fringe relatively short; macrotrichia of veins abundant, lacking on
1st A excepting a small group at apex. Venation: Sc, ending about opposite
Ro*., Sc, at its tip; free tip of Sc, and R. both pale, the former more than its
length beyond Rr, Rr*, continued beyond it as a conspicuous spur provided withmacrotrichia; Rs straight, oblique; supernumerary cross-veinsin cells R. and Ro,
the former more oblique and lying more distad; cell 1st M, widened outwardly,
m straight, transverse, a little shorter than the arcuate basal section of Mt;
m-cu more than its own length beyond fork of M; anal veins divergent, converging
only at bases.
Abdomen with basal tergite pale orange; remainder of tergites dark brown,
the caudal margins slightly paler; basal half of segment two more testaceousyellow. Male terminalia (FiS. 10) of the usual type of the subgenus; tergite,
9f, with caudal margin nearly transverse; ventral dististyle, od, with prolongation
long and slender, the two spines from a pale tubercle at base of prolongation;
face of ventral dististyle with a long pale lobe that is tipped with elongate
yellow setae.
Holotype, J, Rabaul, January, 1933 (F'. H. Taylor).
I refer this very distinct fly to the subgenus Looso Edwards (Ettcltcl. Ertt.,
Diptera, 3, L926,48), where it is most closely allied to Lintonia (Laosa) ntanobo
Alexander (Mindanao), differing in the abundantly dotted, falcate rt-ings,colorationof tlre antennae and legs, and numerous other characters. lf. Laosa is to be
maintained as distinct from Libnotes, its characters will have to be slightly
modified so as to include all those speciesof the last-named subgenus iraving one
or two supernumerary cross-veins in the outer radial field of the r','ing. Besides
the subgenot5'pe,gloriosa Edwards, and the trvo species abor-e mentioned, thesubgenus 'vgill include the following: L. (L.) dipltragnta Alexander' (western
C h i n a ) ; L . ( L . ) f t t s c i n e r c * i s( B r u n e t t i ) ( H i m a l a y a s , W e s t e r n C h i n a ) ; Z . ( L . )regalis (Edwards) (Formosa); Z. (L.) ried,elellaAlexander (Nes' Guinea); andL. (L.) transaersalis (de Meijere) (Formosa to Java). The strongly falcate
wings of the present fly are quite unique in the genus, but mat- rvell prove to bea male character only. The wing-tip of the unique type is slightly injured, but
it is believed that the figure as given is nearly eorrect.
Lrnroxr.r (Lrnsorrs) surroxr, D. SF.
General coloration brownish-yellow, the mesonotum unmarked except for adark area on either side of the mediotergite; knobs of halteres blackened; legs

BY C. P. ALEXANDER.

57

yellow, the outer tarsal segments and very narrorv tips of tibiae dark brown;
wings yellow, sparsely variegated with brown; free tip of Sc, more than twice
its length before Rr; inner ends of cells 2nd M, and M, about on a level; analveins convergent at bases; male terminalia with the ventral dististyle small,oval, the rostral prolongation elongate, without definitely modified spines.
3. Length about 14 mm.; wing, 15'5 mm.

1.
2.
3.
4.
5.
6.
7.
8.

1-8.
Text-figs.
Tiytula (Papuatitrru,la) ttouae-brittattiae, D. SD., venation'
D. SfJ., venation.
Nephrotonta futniscutellata,
Lir,tonia (Laosa) f alcata, n. sp., venation.
(Libnotes)
Limonia
sttttoni, tr. SF., venation.
(I'ibttotes) eboracensis, D. sP,, venation.
Lintonia
Litttonia (Libnotes)
erythront,ero, n. sp., venation.
(Lintonia)
Lirnonia
distitetta, D. SF,, venation.
(Dicranontyia)
Limoniotnatlnistyla, n. sp., venation.

Rostrum brorvnish-1-ellorv,dalker medialll-; palpi pale yellow, the first
segment darker basalll-. Antennae rvith scape and pedicel yellowish-brown,
flagellum brown; basal flagellar segments globular, beyond the third becoming
more oval, the outer segments rapidly passing into long-oval, the outer ones
elongate; terminal segment long ancl slencler, nearly twice the penultimate;verticils of intermediate segmentsabout trvice the segmentsalone. Head fulvous;
anterior vertex reduced to a narro\v stlip that is only about one-fourth thediameter of the scape.
rvithout markings, except for a brown
Mesothorax uniformly bro'*-nish-1'ellorv,
spot on either side of the mediotergite. Halteres pale yellow, the knobs blackened.
Legs yellorv, the outer tarsal segments and very narro\\r apices of the tibiae darkbrown; apices of femora with a densegroup of setae. Wings (FiS. 4) tinged withyellow-,very sparsely variegated rvith pale brown, as follows: Origin of Rs; along
cord and outer end of cell lst 1\[r; basal section of Sc,; free section of Sc, and Rr;outer portion of vein Cu, and tip of vein 2nd A; a narrow infuscation at rving-apex
in medial field; veins pale brown, darker in the clouded areas, in prearcular andcostal fields clearer yellow. Costa incrassated to apex of R., denselJ set with shorttrichia; vein R, (betr';eenfree apex of Sc, and R.) with about nine macrotrichia.

DIPTERT OF THE TERRITOBI OT NEW GUINEA.

II,

Venation: Free apex of Sc, more than twice its length before Rri Rr*z projecting
slightly beyond R, as a small spur bearing a single trichium; R, bent very strongly
caudad, terminating beyond wing-apex; r-m short; inner ends of cells 2nd M,
and M" about on a level; m-cu at about one-third the length of cell lst Mr; analveins converging near bases.
Abdorninal segments obscure orange, the caudal borders of the intermediatesegments very narrowly pale, preceded by a more blackish crossband. Maleterminalia (FiS. 11) rvith the median area of caudal border of tergite, gt, very
gently emarginate, the setae chiefly distributed along border. Ventral dististyle,ud', very small, its body subglobular to short-oval, prolonged into a slender, gently
curved rostral prolongation; face of style with a small tubercle, the whole surface
with unusually long, coarse setae, none of which is modified into a spine.
Gonapophyses,gt, with mesal-apical lobe straight, narrowed to an acute point.
Holotype, J, Keravat, January, lg3g (F. H. Taylor).
I take great pleasure in naming this distinct species in honour of Professor
Harvey Sutton, Director of the School of Public Health and Tropical nledicine.
By Edwards's key to the species of. Libnotes (Jou,rn. Fed,. Matay Str. I/rrs., xiv,
1928,?4-80), the present fly runs to couplet 28, where it disagrees with all species
beyond by the nature of the wing- and leg-pattern, and, especially, the venationof tlre radial field of the wing. It agrees in some regards with Litnonia(Libnotes) sphagnicola (Edwards), but is very distinct from this and all otherdescribed species known to me.
(Lrnxorns) EsoB.rcnNsrs,n. sp.
General coloration of thorax dark grey; palpi and antennal flagellum black;eyes of J contiguous on anterior vertex; posterior vertex with a narrow black
median line; legs reddish-brown; wings pale yellowish subhyaline, with threevery pale brown, diffuse crossbands; m-cu at midlength of cell lst Mri malehypopygium with the rostral prolongation of ventral dististyle bearing a seriesof four slender spines.
6. Length about 11 mm.; rving, 12.5 mm.
Rostrum pale brown; palpi black, apparently only 3-segmented. Antennae
with scape and pedicel dark brorvn, flagellum black; basal flagellar segments oval,
with long, unilaterally arranged verticils that exceed the segments in length;
outer flagellar segments becoming very long and slender. Head brownish-gr€y,
the posterior vertex with a nalrow blackish median line; anterior vertex moresilvery; eyes contiguous for a short space on vertex.
Pronotum and mesonotum dark 8r€y, somewhat clearer grel- on lateral
portions of praescutum and on pleura; praescutum with vague indications ofstripes; pleurotergite somewhat paler and less pruinose. Halteres yellow, theknobs dark brown. Legs lvith the fore coxae darkened, the remaining coxae
chiefly pale; remainder of legs reddish-brown, the outer tarsal segments
blackened. Wings (Fig. 5) with the ground-colour pale yellorvish-subhyaline,
the prearcular and costal regions clear luteous; three diffuse but conspicuous,
pale brown crossbands, the flrst very extensive, postarcular, ocOupying most ofregion before origin of Rs; second band at and just beyond cord, extending fromtip of Sc, to posterior border, the centre of cell 1st M, more or less pale; thethird band is palest and least evident of all, occupying the wing-apex; seam at Rs
heavier and more distinct; veins pale yellow, darker in the clouded areas.
Lruoxu.

BI C. P. ALEXINDEB.

Venation: R, and R. forming an evenly arcuated element beyond the free tip ofScr, the latter pale, erect; cells beyond cord long and narrow, the veins parailel;
inner ends of cells 2nd M, and M. about in transverse alignment; m-cu at midlength of cell 1st M"; anal veins convergent at bases.
Abdomen with basal tergite yellow, the remaining segments deep orangefulvous; centres of discs of individual tergites a little darkened; terminalia
yellow. Male terminalia (Fig. 12) with the tergite, 9f, narrowly transverse, themedian region of caudal margin very restrictedly and shallowly emarginate;setae of tergite chiefly along caudal border, including a linear row of about eighton either side of median line. Basistyle, b, of moderate size. Ventral dististyle,
cd, considerably smaller in area than basistyle, produced into a darkened rostralprolongation that bears near its base a linear row of four (or possibly flve)
slender spines with obtuse tips. Aedeagus, o, broad.
Holotype, J, Duke of York Island (Dr. Hosking).
Li'rnonia (Libnotes) eboracensisis allied to speciessuch as L. (L.) sublasciata
Edwards (Buru) and L. (L.) trifasciata Edwards (Pahang) in the presence otpale but distinct crossbands on the wing. From the former species it is readily
told by the coloration, together with the position of the m-cu crossvein; fromthe latter it is told by the grey coloration of the thorax, coloration of the head,
and, especially, the structure of the male hypopygium. From the unique type, I
am uncertain as to whether there are four or flve rostral sBines.
Lruoxn (Lrexorns) enrrunoMEBA, n. sp.
General coloration reddish-yellow, the mesonotum, pleura and abdomen
unmarked; rostrum, palpi and antennae black; knobs of halteres and legs
chiefly brorvnish-black; wings subhyaline, the prearcular and costal regions moreyellowish; stigma and a narrow apical border brorvn; Rs unusually straightand oblique for a member of this group; anal veins parallel at origin; maleterminalia with two very unequal rostral spines.
6. Length about 8.5 mm.; wing, 7.8 mm.
Rostrum and palpi black. Antennae black throughout; flagellar segments
oval, becoming more elongate outwardly, th.e terminal segment about one-third
longer than the penultimate; longest verticils unilaterally arranged, about onehalf longer than the segments. Head black; eyes contiguous on anterior vertex.
Extreme cephalic portion of pronotum darkened; remainder of thorax,
including pleura, reddish-yellow, only the ventral sternopleurite restrictedly
blackened. Halteres short, pale, the knobs blackened. Legs with the coxae andtrochanters reddish-yellow; femora with bases obscure yellow, more extensively
so on posterior legs, soon passing into brownish-black; tibiae brownish-black, thetarsi more reddish-brown; claws with basal spine. Wings (X'ig. 6) subhyaline,
the prearcular and costal regions more yellowish; stigma small, subcircular,
brown; wing-apex and margin as far as Cu, narrowly sufrused with blackish;veins black, more brownish-yellow in the costal and prearcular regions. Venation:h apparently lacking; Sc, ending opposite midlength of Rr*r, Sc, far from its
tip, Se, alone about two-thirds Rs; free tip of Sc, and R, subequal, pale; Rsunusually straight and oblique for this group, about four times the basal sectionof Rn*u; veins R. and Rn*, extending parallel to one another to margin; m-cu at
midlength of cell 1st Mr; anal veins parallel at origin.
Abdonen reddish; terminalia with the ventral dististyles dusky. Maleterminalia (Fig. 13) rvith tergite, 9t, only gently notched, the margins of the

60

DIPTERA OF THE TERRITORYOF NEItr GUINEA.

II,

lobes thickened and provided with numerous setae. Ventral dististyle, od, muchlarger than the basistyle; rostral prolongation stout, with two very unequal
spines, the innermost a mere seta that is about two-thirds the length of the outer.
Gonapophyses,gr, with mesal-apical lobe blackened, the margin microscopicallyroughened into pale spinous points.
Holotype, J, Makada Island, Duke of York Group, tr'ebruary, 1933 (F. H.
Taylor).
By Edwards's key to the species of Libnotes (1.c.), the present fly runs tocouplet 61, disagreeing with the species involved and all others since described,
in the uniform reddish coloration of the thorax and abdomen. The generalappearance,course of the anal veins, and structure of the male terminalia, are
all much as in Lintonia (Libnotes') noaae-brittaniae (Alexander), L. (L.)
paraistigmo (Alexander), L. (L.) semitristis (Alexander) qnd L. (L.) ta'llloriana
Alexander, all of the Australian and Papuan subregions, but the coloration of thebody, wing-pattern, straight Rs, and shallowly emarginate tergite of the terminalia
readily serve to distinguish the present fly from other similar forms.
Lrltoxr.c, (Lrnnorns) rrxr,onreNe Alexander.
Attn. Mag. I{at. Hist., (10) v, 1930,149.
Known before only from the male sex, taken at Mossman, North Queensland,l,Iarch, L927,by F. H. Taylor.
One t, Keravat, January, 1933 (F. H. Taylor).
Lruoxre (Lrnnorns) onrreu.l (Alexander), var.
Libnotes obliqua Alexander, Rec. Sou,thAustralian Mus., ii, L922, 232.
Three J, Rabaul, January, 1933 (F.H. Taylor). One 9, Keravat, January, 1933
(F'. H. Taylor).
J. Length about 5.5-5.8 mm.; wing,6.4-6.8 mm. 9. Length about 5'5 mm.;wing, 6 mm.
These specimens differ from the type (Cairns district, North Queensland)in the distinct, dark broln, median praescutal stripe, rvhich contlasts con'spicuously with the yellorv to golden-yellorvremainder of praescutum,the lateral
strfpes being obsolete or nearll' so.
Much uncertainty still exists as to the exact limits of the speeiesor races
that centre about the fly formerly called nertosa de lleijere. This pt'oblem has
5ssn sr.rtlinedbriefly by Edwards (Insects of Samoa,Nematocera,part r-i, fasc. 2,
192E, 80) and by the present writer (Arch. liir H!firobiol.. suppl. Band 9,
Tropische Binnengewlsser, ii, 1931, 160). Members of the complex lange from
Ceylon (intmaculi,pennisSenior-White) throughout the Greater Sunda Islands(nerrosa de Meijere) into North Queensland (obliqua Alexander, subaequalis
Alexander) and thence to certain of the major oceanic island groups (manni
Alexander, Solomons; santoe.nsisAlexander, Samoa).
The name neruosa de Meijere (Tid,j. tsoor Ent.. lir', 1911, 36) unfortunately
preoccupied
by a prior usage of the name in the same paper (1.c.,p. 26) and,isin my opinion, both names are homonyms of a still earlier use of the namenerDosa in the comprehensivegeneric concept Litnonia (Limnobia). The next
available name for a member of the group is santoensis Alexander (Bull.
Brooklyn Ent. Soc., xvi, t921,9), eommon and well known in the Samoangroup. Of this species,or race, Edwards had rather abundant material for study,
and this shorved a surprising range in character of size, coloration and venation,

61

BY C. P. ALEXANDER.

together with marked differences between the sexes. Small males tended toresemble the females in venation and stigmal size rather than the larger andmore highly developed males. This unusual range in size and coloration makes
it doubtful whether, ln the above complex, we are dealing with a group ot
closely allied and generally similar species or whether these are nothing but
subspecies or forms of a single species of great geographic range. The type ofstructure of the male terminalia is generally similar in all of these species inwhich the organ has been studied, but the same statement holds true for mostother members of.Libnotes lvhere, on terminalian characters, the abundant species
fall in only a few groups, showing a curlous monotony of structure. This lattercase especially holds true in the tyBical form of Libnote.s, where a peculiar typeof terminalia is found, and which, moreover, recurs in two other supposedly valid
If it is found thatsubgenera (Laosa Edwards, typical Limonia, Meigen).
group
now under discussion represent but one or few
members of the samoensis
species, then it seems certain that there will occur a similar and even more
drastic consolidation of species names in other groups of Libnotes. Males of thespecies or subspecies of the present group in the Australasian region may beseparated by the accompanying key.
praescutal stripe obsolete, the laterals continued across the suture on to
1. lledianthe scutal lobes .
.. . sqnloensis AlexanderMedian praescutal stripe distinct, the laterals present or obsolete
2
2. Radial and subcostal cells basad of stigma strongly suffused with brown . . .
monni Alexanderr
No infuscation in the cells basad of stigma
3
3. Ali elements of anterior cord of wings subequal in length and lying subtransverseto the length of wing
, , subaequolds Alexanderr
Elements of cord more oblique, Rs either distinctly longer or lying more proximad
tlran the other elements. . , , oblicluo Alexanderr

Lrlroxn(Lrnivorns) solouoxrs (Alexander).

Libnotes solotttortisAlexander, Ann. Mag. Irlat. Hist., (9) xiii, L924,39.
Described from the Solomon Islands. Rabaul, trvo d, one ?, December, L932,
to January, 1933 (F. H. Taylor).
This species is allied to notata (van der Wulp), but is readily told from thisspecies and all others in the gl'oup by the broad, conspicuous, black bases of alltibiae, in conjunction with the undarkened apices of the same. The vestitureof the femora is reduced to very short spinous setae. Apices of cerci bidentate.
Lruoxr--r (Lntoxr.r)

suB-\LBrrrnsrs Alexander.

Ph,ilippine Journ. Sci., xli, 1930,2gg.
Described from Luzon, Philippine Islands.
(f'. H. Taylor).

One J, Keravat, January, 1933

Lruoxru (Lrrroxr.r) Drsrr!'ENA,tr. sp.
General coloration dark brown, the pleura somewhat paler than the central
portions of the mesonbtum; antennae black throughout, the flagellar segments
moniliform, each with short but conspicuous necks; wings tinged with brown, thesmall stigma a little darker; macrotrichia of veins long and conspicuous; innerend of cell 1st M, arcuated; m-cu unusually far distad, lying more than one-half
its length beyond the fork of M.
tLibnotes mantni Alex., Ann. Mag. Not. Eist., (9) xiii, tg}4,
z L . s r t b o e q z { o l i sA l e x , , I b i d . , ( 9 ) v i i i , L 9 Z L , 5 5 4 .
s L, obliqua Alex., Rec. South Austrolian MuE., ii, L922, ZBZ.

4I.

62

DrprERA oF THE TEnBrroRy oF NEw curNEA.

ii,

9. Length about 8.2 mm.; wing, 4 mm.
Rostrum and palpi black. Antennae black throughout; flagellar segments
very strongly moniliform, the segments globular, with short glabrous black necks;
outer segments more oval, but similarly pedicellate; longest verticils unilaterally
arranged and exceeding the segments. Head dark grey
Mesonotum relatively gibbous, projecting cephalad over the small pronotum.
Pronotum dark brown. Mesonotum dark brown, the praescutum somewhat paler
laterally. Pleura brown, the propleura slightly darker. Halteres with stem
black, the base narrowly pale, the knobs broken. Legs with the coxae brown;
trochanters testaceous; remainder of legs broken. wings (Fig. 7) tinged
with brown, the small, short-oval stigma a little darker brown; veins darker
brown. Macrotrichia of veins long and conspicuous, including all longitudinalveins beyond cord; all of Rs excepting the extreme base; distal half of mainstem of M; distal third of basal section of Cu, and extreme tips of both analveins. Venation: Sc, ending about opposite two-thirds to three-fourths the lengthof Rs, sc, at its apex; free apex of Sc, pale, lying a litile basad of level of R,;
inner end of cell lst M, arcuated; m-eu more than one-half its length beyond
the fork of M; cell 2nd A narrow.
Abdominal tergites brownish-black; sternites paler, brownish-yellow. Cerci
slender, the tips acute.
Holotype, $, Keravat, January, lgBB (F'. H. Taylor).
The present fly is most nearly allied to the Philippine Lintonia (Limonia)
retrusa Alexander, being told from this and other small similar members of thesubgenus by the distal position of m-cu.
Lrlroxle (Gnn.rnourre) uenc.r (Alexander.), var.
Geranomllia (Geranomyia) nxancaAlexander, Ann. nlag. Nat. Hist., (g) xiii,
1 9 2 4 ,1 9 0 .
One J, Toma, altitude about 1,000feet, February, lggg (F. H. Taylor).
Close to the typical form (North Queensland) but difiering in having theouter field of wing darkened to the wing-tip; m-cu,fully its own length beyond
the fork of M, exceeding in length the distal section of Cu,; vein 2nd A at near
midlength, gently but distinctly concave. Male terminalia with the tergite
narrow' transverse. Ventral clististyle very large and fleshy, much larger thanin argentifera and allies; apex of rostral prolongation beyond the spines shortand stubby. Dorsal dististyle long and unusually slender, strongll- arcuated.
Gonapophyseswith the mesal-apical lobe slender, but provided wiilr setae. as inall members of t}l.e sorbiHorzsgroup.
Ltuonre (Iorocrocurne) near NovocALEDoNrc-.\
Alexander.
Limonia (Iclioglochina) noaocaled,onica
Alexander, Encycl. Entomol., Dipt., v,
1 9 2 9 ,9 0 .
One broken J, Duke of york Island (Dr.. Hosking).
Lrlroxre (DrcneNolrrra) MAGNrsr-r-L.\1
n. sp.
Belongs to the punctulato group; nearest to kulitt; thorax dark grey; thepraescutum with an ill'defined median stripe; femora brown, the tips
very narrowly
and abruptly yellow; wings of J with long, conspicuous costal fringe; maleterminaha with ventral dististyle very large and fleshy; rostral spines two,
unusually short.
3. Length about 8.8 mm.; wing, 4.8 mm.

BY C. P. ALEXANDEB.

63

Rostrum and palpi black. Antennae black throughout; pedicel enlarged;
basal flagellar segments globular, the outer ones passing into oval. Headbrownish-grey; anterior vertex reduced to a linear strip that is about one-halfwider than a single row of ommatidia.

t('g
/\9t

r3

t4

Text-figs. 9-1{.
9 . T i p u l a ( P a p u , a t i p u l a ) n o t a e - b r i t t a n i a e . D . S I r . ,m a l e i l y p o p y g i u m , d e t a i t s .
10. Limonio (Loosa) Jalcota, n. sp., male hypopygium.
11. Limonia (Libtr,otes) su.ttoni,n. sp., male hypopygium.
12. Limonia (Libtr,otes) eboracensis, D. sp., male h]'popygium.
1 3 . L i m o n i a ( L i b t w t e s ) e r y t h t o n r " e r c r .n . s p . , m a l e h y l t o p y g i u m .
74. Iti,monia (Dicranom.yia) ntagttistyla., n. sp., male hypopygium.

Thorax uniformly dark grey or dark plumbeous, the praescutum with a very
ill-defined brown median stripe that is divided behind. Halteres pale, the baseof knobs restrictedly darkened. Legs with the coxae brown; trochanters obscureyellow; femora dark brown, the bases restrictedly obscure yellow, the apices very
narrowly and abruptly obscure yellow; tibiae paler brown; tarsi chiefly obscureyellow, the outer segments darkenecl. Wings (FiS. 8) with the ground-colour
greyish-white, the prearcular and costal portions to apex clear pale yellow; arestricted grey spotted pattern, arranged as in th.e punctulata group; about
three such spots in cell C beyond the humeral area; a dark spot at midlength ofcell Rr; veins pale brown or yellowish-brown, darker in the infuscated areas.
Costal fringe (at least io d) very long and conspicuous.Venation as in group.
Abdomen dark brown. Male terminalia (FiC. 14) with the tergite, gf, transverse, the caudal margin very shallowly emarginate. Basistyle, D, small. Ventral
dististyle, ad, very large and fleshy, its greatest length nearly equal to two andone-half times that of the dorsal dististyle; rostral prolongation small, with two
unusualll- small spines placed on the side of the prolongation, near the lower
margin; spines shorter than the diameter of the prolongation at point of insertion;

oF NEw curNEA.
DrprEnr oF TrrE TERRTToRY

ii,

distance between spines shorter than the diameter of either. Gonapophyses,,/,with the mesal-apicalangle a slender curved hook.
Holotype, J, Rabaul, February, 1933 (f'. H. Taylor).
The present species is readily told from all allies in the small size, long
costal fringe and the structure of the male terminalia, especially the ventral
dististyle and its rostral armature. The species having the terminalia and costal
fringe most similar is Lirnonia (Dicranomyia) kutin Alexander (Victoria and
New South Wales), a much larger fly with unvariegated costal region and withthe spines of the rostral prolongation conspicuously more elongate.
Lruoxrl (PsuupoorocurNA) EosKrNGr,n. sp.
Mesonotum dark brown, the ventral thoracic pleura blackened; posterior
femora uniformly blackened; all tibiae white, with two dark rings, the morebasal one narrower on fore and irind tibiae, broader on mid-tibae; wings withcell 2nd M. deep, nearly twice its petiole; vein 2nd A unusually short and alched.
9. Length about 6 mm.; wing, 6'3 mm.
Head broken.
Mesonotum dark brown, not or scarcely variegated by paler. Pleura chiefly
blackened,including the entire sternopleurite, the dorsal pleurites lnore testaceousbrown. Halteres broken. Legs with the coxae obscure yeilowish-testaceous;
trochanters a little darker; fore femora obscure brownish-yellow, narrowly tippedwith dark brown; mid-femora chiefly infuscated on basal half, thence passing
into dirty white, becoming restrictedly pure white iust betore the narrow darkbrown apices; posterior femora entirely blackened; all tibae pure white, thefore and hind pair each with two narrow black rings, the more basal one nal'rolYer
and somewhat paler than the outer or post-medial band; on mid'tibiae, the basal
blaek ring is mueh more extensive than the outer, being fully twice as wide,
about two-thirds as extrnsive as the white enclosed annulus; tarsi white. Wings(Fig. 15) greyish-subhyaline; stigma oval, vely distinct; veins brorvnish-black.
Venation: Sc relatively shoi't, Sc, ending shortly before oligin of Rs, Sc, somedistance from its tip, Sc, alone ionger than Rs; cell 2nd M, deep, nearly ts'ice
its petiole; vein 2nd A unusually short and arched.
Abdominal tergites dark brorvn; sternites pale yellow.
Holotype, !, Rabaul, February, 1933 (F. H. Taylor).
I take pleasure in naming this species after Dr. Hosliing, rvho collected
several Tipulidae near Kokopo, Nerv Britain. This is the lirst species of thesubgenus from the Australasian region having two dark tibial rings, the twoiaticittcta (Edwalds), offorms hitherto made known, Litnottia (Pseu,d,oglocltina)
Samoa,and L. (P.) gtu,lchripes(Alexander), of North Queensland,having a single
darkened ring. From l{alayan species having a single tibial annulus, as
L. (P.) lcobusi (de Meijere), of Java, and L. (P.) 'unicittctipes(Alexanderl ofBorneo and the Philippines, the present fly is ampll- distinct. The Oriental
specieshaving two tibial rings, as L. (P.) bilatior Alexander',.L. (P.) bilatissirna
Alexander, Z. (P.) pictipes (Brunetti), L. (P.) r'iukiuensis Alexander, andothers, are readily told by the details of wing- and leg-pattern and the venation.
Lruorre (Trrnrrrrcoutre) An'\crrNoPrrrr,r(Alexander).
DicranomAia (Thrgpticontyia) arachnophilo Alexander, Ph,ilippine Journ. Sci.,
xxxiii, 192?,301.
Described from the Philippines. Keravat, January, 1933, February, 1933
(F. H. Taylor).

BY C. P. ALEXANDEB.

66

yoaKnNgrg, n. subsp.
(Euor,ocurne) noveE-GUTNEAE
Similar to typical not:ae-guineaein the large size (wing, d, more than 11mm.) ;thorax reddish-brown; halteres and abdomen black; proximal third of basitarsus
black, the remainder of tarsi yellowish-white.
3. Length about 13 mm.; wlng, 11.5 mm.
Rostrum and palpi brownish-black. Antennae black throughout; flagellar
segments elongate-oval, very slender. Head dark brown.
Thorax reddish-brown. Halteres black. Legs with the coxae and trochantersreddish-brown; femora brownish-black, obscure yellow basally; fore tibiae brown,
middle and hind tibiae brownish-black to black; basitarsi with a little more than'Wingsthe proximal third black, the remainder of tarsi yellowish-white.
(Fig. 161
hyaline, the tip weakly infumed; stigma oval, darker brown; veins brownishblack. Venation as shown.
Abdomen black.
Holotype, J, Duke of York Islands (Dr. Hosking).
The typlcal form of Limonio (Euglochina) nwae-guineae (d,e Meijere) , Tiid.
ooor Ent., lviii, 1915,101, Pl. 1, fl1.7, wing-apex) difiers in the uniform brownlshyellow coloratlon of the body, including the abdomen; yellowish tibiae andbasitarsi, and the yellow halteres. The type of this form was from "Ilollandla',,on the west coast of Humboldt Bay, Northern New Guinea (2" gz, 2g,, s. Lat.,740"44'L2" E. Long.).
LruonreHnrrug (Rg.rupsor,rMNoBrA) p.{prr-{NUBAlexander.
Philippine Journ. Sci., liv, 1gB4 (in press).
A part of the type-material of this species was from Laup, New Britaln(Dr. Hosking).
Hnxetournr.
.
Er,npgersroMyrA (Er.npg.lnrouroons) TAvLoBTANA,
n. gp.
General eoloration black; halteres and legs black throughout; wings strongly
sufiused with blacktsh; cell 2nd A very long and narrow; abdomen black, thebases of the segments broadly greyish-nacreous.
3. Length, excluding rostrum, about 6.5 mm.; wing, ? mm.; rostrum, 3.b mn.
Length,excluding rostrum, about 6 mm.; wing, b.b mm.; rostrum,2 mm.
9.
Rostrum black, about one-half the length of body in male, much shorter infemale, as shown by measurements. Antennae black, the verticils very longand conspicuous. Head blackish.
Thorax entirely polished black. Halteres black. Legs with the coxae
blackish on outer faces, the inner faces and all trochanters more brownishtestaceous; remainder of legs, including all tarsi, black. wings (Fis. LT,
strongly sufiused with blackish, especially the basal region and cells C and Sc;
origin of Rs and cord seamed with darker brown, more broadly so in male; veins
black. Venation: Rs perpendicular at origin; anterior branch of Rs runningvery close to main stem; m-cu from about one-half to three-fourths its length
beyond fork of M; cell Znd A very long and narrow.
Abdomeu conspicuously ringed with black and greyish-nacreous, the lattercolour occupying the bases of the segments and involving a little less than onehalf the segments; terminalia black. Ovipositor with the cerci black, only theertreme apices pale; hypovalvae black on basal half, the distal end bright hornyellow.

66

DIPTER.{ OF THN TNERITORY OF NEW GUINEA.

ii,

Holotype, 3, Keravat, January, 1933 (F. H. Taylor).
Allotype, ?, Toma,
altitude about 1,000 feet, tr'ebruary, 1933 (F. H. Taylor).
ElephantomAia (Elephantomyodes) tayloriana is named in honour of thecollector of this valuable series of Tipulidae, Mr. F'. H. Taylor. It is strikingly
difierent from the other species with black feet in the body*oloration andunusually long and narrorv cell 2nd A of the wings. The Australian .o. (8.)
lumicosta Alexander (Queensland-Northern New South Wales) is readily told by
the snowy-white tarsi.
Enrosrnnrnr.
(Wiedemann).
Coxosr,l TRBoRATA
Limnobia irrorata Wiedeman\, Aussereur. zweifl. Ins., i, 1828,bZ4.
One example of this widespread crane-fly, Keravat, January, lggg (F. H.
Taylor).
Tnnnrnporrlre (Tnnnrnrour.re) prcrrpENNrcBezzi.
Trentepoh,lia pictipennis Bezzi, Pltilippine Jou.t'n.Sci., xii, D, 191?,115.
The present specimens greatly extend the known range of the species to theeast. One i, Rabaul, January, 1933 (F. H. Taylor). One J, Kelavat, February,1933 (F. H. Taylor).
Thb rather numerous species of. Trentepohlia now known from New Britain
may be separated by the following key:l . \ \ ' i n g s w i t h c e l l l s t M , c l o s e d ; t h r e e o u t e r m e d i a l v e i n s , M r , " , ) I . a n d trf.' ..
2
2.
3.

4.

5.
6.

i.

\ Y t n g s w i t h c e l l l s t M , o p e n b y t h e a t r o p h y o f m a n d the t$'o tlistal sections of M r ltu'o outer medial veins, Mr., and M, (Treu,tepoltlio Bigot )aVein R, atrophied (Plesiornottgonle Brunetti).. notae-brittaniae, n. sp.
Vein R, preserved (llottgotntr, lVestrvood)3
Apices of femora l.rlack or only vaguely brightened4
Apices of femora and Jtases of tibiae broadly and conspicuousll' whiteau,stralasiae Skuse
Tlbiae entirely dark, not dilated at tips
brettipes Alexander
A p i c e s o f t i b i a e r v h i t e n e d ; m i d - t i b i a e c o n s p i c u o u s l y d i l a t e d a t t i p s i n p e n n i p e s a ndsnbpettnota
D
Basitarsi not darkenecl ...
pennipes (osten sacken)
Basitarsi blackened, at least on proximal ends
Costal fringe of wings long and conspicuous, especially in male; cell R, at wingmargin less than one-half as extensive as cell R,; tibiae not dilated at tips;
basitarsi darkened only at extreme bases; mid and hind femora $'ith spines atbases
costofirttbriata, n. sp.
Costal fringe short in both sexes; cell R, at rving-margin very wide, rnore extensive
than cell R.; tibiae, especially the middle pair, conspicuously clilated at outer
ends; basitarsi extensively blackened; mld and htnd fenrora without spines,. . ., su,bpnnota, n. sp.
Basal al,rdominal segments orange, the terminal segments blaek.q,raomen';i;;k'i;;;,;;;;;; . . . . . . . . . . . . . . . . . . . . . . .

":"u:o.|u;,"[y#"tr:TH]t

Tnnnrnporrr-rl (Pr.nsrouoNcouA) NovAE-BRrrrANrAE,
n. sp.
General coloration pale yellow, the mesonotum a little darker; legs brown,
the narrow tips of tibiae and all tarsi snowy-white; wings whitish-subhyaline,
unmarked.
6. Length about 6 mm.; wing, 6 mm.
Rostrum pale yellow, whitish-pruinose; palpi pale yellow. Antennae withbasal two segmgnts yellow; flagellum broken. Head very pale grey; anteriorvertex reduced to a strip.

BY C. P. ALEXANDER.

Pronoturn pale yellow. Mesonotum chiefly pale reddish-brown, the scutal
lobes and posterior portions of mediotergite darker brown; scutellum and medianarea of scutum more testaceous-yellow. Pleura pale yellow. Halteres yellow
throughout. Legs (mlddle legs broken) with coxae and trochanters pale yellow;
femora and tibiae pale brown, the apices of tibiae (distal eighth or less) and alltarsi snowy-white; fore femora with a linear group of four black getae nearbase, the outermost more powerful; in addition to these, with scattered, semi.
erect black setae distributed throughout the length of the segment; posterior
femora with a series of about 15 short, spine-like setae in a linear row near base,with additional, gradually more elongated setae distributed at increasing intervals
throughout the length of the segment. Wings (F''ig. 1S) whitish-subhyaline,
unmarked; veins pale brownish-yellow. Venation: Inner ends of cells Ru and M,
lying somewhat nlore basad than that of 2nd Mr; m-cu shortly before fork of M.
Abdominal tergites pale brownish-yellow; sternites somewhat paler.
Holotype, J, Rabaul, January, 1933 (F. H. Taylor).
The occurrence of. Trentepohlia (Mongonr,a)pennipes (Osten Sacken) in New
Britaln makes it advisable to compare the two flies critically, since it seems very
obvious that the present species has been derived directly from species likepennipes through the total loss of vein R, of the wings. Edwards (Journ. Fed,.
Malay Sfr. Mus., xvi, 1931, 499) has recorded a specimen of pennipes from
Borneo in which vein R, is entirely lacking, making the specimen confolm tothe characters of Plesiotn,ongoma. I have likewise seen specimens of. pennipes
with this abnormality, either on a single wing or on both wings. The present fly
dlffers from pennipes in the pale wing veins, vel'y narrow white tibial tips, andthe evenly areuated vein R,, this vein in pennipes showing a slight angulation at
the point of departule of vein R,. The known speciesof the subgenusmay beseparatedby the follou'ing key:!
1. Wings rvith cell I st 11, open ll1' atrophy of nr ; mid-tibiae near apex rvith long,conspicuous fringes ()r paddles of black and white setae ( Pahang, Sumatra,
Borneo).... nigropennata Edwards\\'ings rvith eell lst M" closed: legs rvithout modified hair fringes ....
2
2. \\-ings u'ith veins pale. stignra lacking ; halteres yellorv throu5;hout; legs pale brown,
the tarsi and narroli: tifis of tibiae u'hite; no ltrightening of genua (NewBritain) ...
nouae-brittaniae, n. sp.
\tr-ings with veins blac'k, distinct ; cord more or less searned with darker; halteres
rvith darkenecl knobs ; legs not patterned as above3
3. Legs pale brown ; femoral tips and tibial bases broarll]' $'hite; tibiae chtefly white,in cases a little clarkened beyond base ; tarsi snorvy-white (Selangor, Borneo)
cand.itlipes Edwards.
Legs bright -vellorv; tiDs of fore fenrora ltlacliened (Assam) .... pelt,oao(Brunetti).

Tnrxrrpour.rr (Moscour) pnxxrprs (Osten Sacken).
Mongonta pennipes Osten Sacken,Berlin. Ent. Zeitscltr., xxxi, 1887,204.
Keravat, January, February, 1933 (S. V. Bayley and F. H. Taylor). Toma,
altitude about 1,000feet, February, 1933 (F. H. Taylor).
Tnnxrnposr,re (Morvcour) cosrorruBRrArl, n. sp.
I\[esonotumand pleura dark cinnamon-brown, the praescutum with a capillarybrownish-black vitta on anterior half; femora black; tibiae black, the outer half
paler, the apices narrowly pale yellow; tarsi yellowish-white, the extreme proximal
ends of basitarsi darker; spines on bases of middle and hind femora; hindbasitarsi with three or four long black setae at base; wings with a dusky tinge,

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DIPTERA oF TEE TEBRIToBY oF NEw GUINEA.

ii.

eells C and Sc blackish; wing-apex narrowly darkened; veins black; costal fringe(male) long; all elements closing cell 1st M, subequal.
J. Length about 5.5-6 mm.; wing,6-6.b mm. g. Length about 6 mm.; wing,
6.5 mm.
Rostrum and palpi brownish-black. Antennae black throughout; flagellar
segments cylindrlcal, the verticils shorter than the segments. Head brownishblack; anterior vertex reduced to a narrow strip.

15.
16.
17.
18.
19.
20.

Text-figs. 15-20.
Lintonia (Pseud,oglochina) hoskitr,gi, n. sp., venation.
(Eu'glochina)
Limonia
noooe-gu,ineae got kensis, n. subsp., venation.
(El'ephantomyod,es) tayloriana, n. sp., venation.
Ele|thantomgia
Tren'tepoh'lia (Plesiamongoma)noaae-brittanioe, tr. Sp., venation.
Trentepohli,a (Itongorna) costofi,mbriata, n. sp., venation.
Trentepohlia (Il[gtr.gom,a) subpennato, r. sp., venation.

Cervical sclerites and pronotum dark brown. Mesonotum and pleura dark
cinnamon-brown, the praescutum with a capillary brownish-black vitta on anterior
half ; scutal lobes slightly more darkened; scutellum dark br.own; anterior
pleurites slightly darker than the posterior ones. Halteres brownish-black, thebase of stem a little paler. Legs with the fore coxae dark brown, the remaining
eoxae and all trochanters yellowish-testaceous; femora black; tibiae black basally,
the outer half paling to dirty brownish-white, the extreme tips narrowly andconspicuously pale yellow; tarsi yellowish-rvhite, the basitarsi very restrictedly
darkened at proximal ends; middle femora in both sexes with a group of from
16 to 17 short black spines at base; hind femora with about 10 to 12 such spines;
fore femora without spines; posterior basitarsi of both sexes with a group of
about 3 or 4 long black setae at extreme base. Wings (Fig. 19) with a dusky
tinge, cells C and Sc, except basally, strongly sufrused with blackish; stigma
not otherwise darkened; wing-apex narrowly more darkened; a dusky area
between anal veins near bases; veins black. Costal fringe of male very long andconspicuous, short only on basal sixth or so; in female, fringe shorter but still
longer than usual in the genus. Venation i Rz*a*rlonger than Rs; vein R, oblique,eo cell R, is wide at margin; all elements closing cell lst M, subequal; m-cu Just
before fork of M; apical fusion of veins cu, and lst A distinct.
Abdominal tergites dark brown; sternites obscure brownish-yellow; terminalia
dark. Oviposltor with hypovalvae darkened at bases, the remainder yellowish
horn-colour.

BY C. P. ALEXANDER.

69

Holotype, J, Keravat, January, 1933 (F. H. Taylor). Allotopotype, t, pinned
with type, paratopotype d.
Trentepohlia (Mongoma) costofimbriata is readily told from allied regional
species by the long costal fringe of the wings. T. (M.\ fimbriata Edwards(Borneo) likewise shows this character, but is otherwise a very different fly,
without spines on mid-femora and with a very difrerent venation of the radial
fleld. By Edwards's key to the Australasian species of Mongonxa (Ins. Samoa,
Diptera Nematocera, vi, fasc. 2, 1928, g4), the present fly runs to couplet 5,
disagreeing with both included species.
Tnsnrnposlre (MoNoolr.r) sunpnnnATA, n. sp.
General coloration of mesonotum dark brown to brownish-black, the pleura
obscure yellow; femora black, the apiees very vaguely paler; tibiae black, thetips broadly snowy-white; basitarsi chiefly blackened, the remaining segments
paling to yellowish-white; tips of middle and posterior tibiae dilated and fringed
with conspicuoui white setae; wings relatively long and narrow, faintly tinged
with blackish, cell C and the stigma black; veins Rs and R. only slightly
divergent; cell R, at margin more extensive than cell R..
3. Length about 7.8-8 mm.; wing, 7-7.5 mm.
Rostrum and labial palpi pale yellow; maxillary palpi black. Antennae black
throughout; flagellar segments cylindrical, with verticils that are shorter than
the segments. Head black; anterior vertex reduced to a linear strip or evenslightly interrupted by approximation of eyes.
Cervical sclerites and pronotum above dark brown, the latter yellow on sides
beneath. Mesonotum dark brown to brownish-black, the humeral region ofpraescutum restrictedly and very vaguely brightened. Pleura obscure yellow, thepropleura and anepisternum vaguely suffused with dusky. Halteres dark brown,
the extreme base of stem pale. Legs with coxae yellowish-testaceous, the fore
coxae a trifle darker; femora black, the extreme bases obscure yellow; tips offemora very vaguely paler, obseure yellowish-brown; tibiae black, the tips broadly
snowy-white, slightly narrower on fore legs; basitarsi blackened, their extreme
tips and the outer tarsal segments paling to yellowish-white; fore tibiae scarcely
expanded at tips; hind, and especially the middle tibiae, conspicuously flattened
and dilated, fringed with white setae that are shorter than the expanded portion;fore femora with three or four long erect black setae at some distance from base.
Wings (FiS.20) relatively long and narrow; ground-colour faintly blackish; cell
Sc and the confluent stigma black; wing-tip insensibly darkened; veins black,very conspicuous. Costal fringe relatively short. Venation: R, shortly before
fork of Rr*., the distance variable, longest in the holotype; R, and Rr only
slightly divergent, so cell R. is narrow and parallel-sicled for more than one-half
its length; cell R, at margin considerably more extensive than cell Rr; vein R.
strongly sinuous; inner end of cell M, slightly more basad than the other outcr
cells; m-cu close to fork of M ; apical fusion of Cu, and 1st A relatively extensive,
nearly equal to m-cu; cell 2nd A long and narrow.
Abdominal tergites dark brown, the incisures of the intermediate segments
vaguely paler; terminalia dark; sternites obscure yellow.
Holotype, d, paratopotype, J, Keravat, February, 1933 (F'. H. Taylor).
By Edwards's key to the Australasian species of Mongoma (1.c., 1928,94),
present
fly runs to tarsalis Alexander. The present fly is very difrerent fromtheall other species known to me in the dilated tips of the middle and hind tibiae,

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DIPTEiIA OF TEE TEBBIToRY Or. NEw GI'INEA.

ii.

the blaek basitarsi, and the venation, as the only sltghtly dtvergent velns R, and
R.. The key given on a preceding page will readily separate the species fromothers in New Britain.
Tnnnrrposr,re(Monooue)

ausrBALAsrAE Skuse.

Trentepoltlia australasiae Skuse, Pnoc. Lrun. Soc. N.S.W., xiv, 1890, 834.
Toma, altitude about 1,000 feet, February, 1933 (E.. H. Taylor).
MakadaIeland, Duke of York Grodp, February, 1933 (F. H. Taylor).
The latter specimen has the thoracic dorsum paler, more reddish-bro\rn, but
I can detect no other difierences.
Trpnrsporr,r.l(MoncoMA) BREvrpEsAlexander.

A n n . M a g . N a t . H i s t , ( 1 0 ) v i i , 1 9 3 1 ,1 8 .
The type, a female, was from Suali, Vailala River, Papua, collected by
Littlechild. One J, Toma, altitude about 1,000feet, February, 1933 (F.H. Taylor).
The costal fringe is short.
SrvnrucouYrA cEYLoNrcAEdwards.
Ann. Mag. Nat. Hist., (8) viil, 62, 1911.
One J, Karavat, December, 1932 (F. H. Taylor).
Edwards had previously (Ann. Mag.Nat. Hist, (9) xiii, 1924, 270) recorded
this species from Madang (Friedrich Wilhelmshafen), Seleo Is., ofi Aitape, andAitape Roadstead (Berlinhafen), in north-eastern New Guinea, a notable eastward extension of the known range.