富山市科学博物館研究報告第33号，pp39‑45；（2()10）

ANewSpeciesoftheGenusL"c j"火s（Crustacea：Isopoda）fromAshiu，Miyama，Nantan‑shi，Kyoto，CentralJapan＊NobomNunomura

ToyamaScienceMuseuml‑
8‑
31Nishinakano‑machi，
Toyama
9
39‑
8()
84,
Japan

京都大学芦生研究林から発見されたハヤシワラジムシの1新種
布村昇
富山市科学博物館939‑8084富山県富山市西中野町1‑8‑3：京都府南丹市の京都大学フィールド科学教育研究センター森林ステーション芦生研
究林内から渡辺弘之博士により採集された陸生等脚目の1種を調査したところ、オオ
ハヤシワラジムシ属の新種であることが判明し、新種L"casjoj昨sash奴e"sjs(和名：アシュウハヤシワラジムシ：新称)として記載した。本種は宮崎県から知られている

ニチナンハヤシワラジムシL師c方加α"e" Nunomuraと最も類似するが、(1)はっきりした背面の模様があること、(2)第2触角の第2節が第1節より明らかに長いこと、（3）胸肢に多数の2叉剛毛が見られること、(4)第1触角の剛毛数が多いこと、(5)第1〜4胸節の剛毛が側縁からより離れた位置にあること、(6)第1および第2胸節の長節や腕節の剛毛は比較的密生しないことなどで区別される。本種のホロタイプは富山市科学博物館（TOYACr‑l9907）に保管される。
キ ー ワ ード：等脚類、新種、芦生、分類学、アシュウハヤシワラジムシ

Keywords：：Isopod，newspecies，Ashiu，taxonomy，L"c joj昨s 〃"e"sjs

Hitherto，11speciesofthegenusL"c /or昨sintheworld（Shmalfilss、2003）and7speciesinJapan(Nunomura，1987,1991,1992,2000,2003,2008）havealreadvbeenknownasvaledRecently,ProfEmeritusDrWatanabehappened
tocollectstrangeten･estrialisopodcrustaceansinAshiuForestResearchStation，FieldScienceandResearchCenter，Nantan‑shi，KyotoThevweresenttomefbridentiflcation，Atthecloserexaminationsofmine，theyprovedtorepresent

tobeasnewspeciesofthegenusZ,"c joj昨s、
Orderlsopoda

FamilyAgnaridae
L"c妬加i s ん加E" ，n.Sp.

（Japanesename:Ashiu‑hayashi‑warajimushi,new）（Figs，1−3）
Mnre"α／exα'"j" ：2早早(l早holotype，70mminbodylengthandl早paratype，5．4mminbodylength)：
Nodahatadani，Ashiu，Miyama，Nantan‑shi，Kyoto，31July2009,colLHiroyukiWatanabe；1J(allotype，70mmlnbodylength)andl早paratype,55mminbodylength),Kamitani,AshiuMiyama,Nantan‑shi,Kyoto,May26,2009,colL
HiroyukiWatanabeTypeseriesisdepositedasfbllows：holotype（TOYACr‑19907)，allotype（TOYACr‑19908）and＊Contributions廿omtheTovamaScienceMuseum・No.38重
ヅ

qCJ里』

NobomNunomura

aparatype（TOYACr‑l9909）
atToyamaScienceMuseum,paratypesandaparatype（OMNHAr
‐
7776）
atth
運 OsakaMuseumofNaturalHistory、
longltu
‑
Descr
""o〃
げんo/oeた"，α/e･Bodyelli
psoid(Fig.
1A),
2ltimesaslongasw
ide・
Colorblackish,w
ithdinaldar
kerpattems，
symmetrywithrespecttoaline，itlooksasifanlmageofRorschachtest・Medial areao萱
anter
lor
mar
ginofcephalontr
langular(FiglB).
Ante
rolaterallappet
late
ralborder､Nodulilateralisonpereonalsomlte皇l‑4（Fig.
1R）
remote廿omthelateralborder､Posterlor
marginofpleotelsontr
langular．

一一一/ぐずで､#・ルー

I ・一員伽Eﾙｩ
ｰ〆一一ー 1 m m(、一一、､

一一一一一一一一一〜、−一が門別︑８{◎‐〜＝ごー 、工菖P

ござ５４５０４０００３
一・一・・︾一営一0．25

01胃0．05がR】

Ⅱ､ⅣVⅥⅦFig.1．L" s"j此s ん加e"sjsn.§p、A，DorsalviewラB，AnteriorviewofcephalonラC，Antennule；D､AntennaラE，Rightmandible；F，Le仕mandible；
G，Innerlobeofmaxillula；H，OuterlobeofthesameラI，Maxilla；J，Maxilliped；K−N，Pleopodsoffemalel‑4＆O

Endopodofpleopod5P，Exopodofthesame；Q,Uropod;R,PositionofnodulilateraIis（All,Holotypefemale）

4もAnewL"c /o/火sfromAshiuJapan

Antennule(FiglC）
3−segmented；
terminalsegmentwith6aesthetascsatthetipAntenna（FiglD)，reachingtheboundaxybetw
eenpe
reonitesland2・
Tenn
i
nal
Hage
llar
seg
ment
s
1．
6ti
meslongerthanthebasalone・Rightmandible(FiglE):
parsincisiv
a3‑headed；laciniamobilisnotchitinizedw
eakly
3‑
toothedラ
3penicils；
processu
s
molar
isr
epr
e
‐
sentedbyatufiof
setae・
Left
mandible（FiglF):parsincisiva
3‑headed；laciniamobilis3‑toothed；2penicils
ぅprocessus
molar
isr
epr
esentedbya
mfiofsetae・Maxillu
la：
inner
lobe
（FiglG)w
ith
2plu
mosesetaeas
mallacutepr
Q
jectionouterlobe（
FiglH）w
ithl
Osimpleteethondistalmargin
Maxilla（FiglI）
relativelyslender
ラapicallybilobed,themnerwide
randbearingafieldofdenselyspacedse
nsillaMaxillipe。
（FiglJ)：
enditewith3spursondistalmargm；
palpalsegmentlwithastrongtooth

Pereopodl（Fig2A)
:
basi
s3．
6ti
mesaslongasw
ide,
w
ithl
O
‑
l
2setaeonlnner
mar
ginan
d5set
aeonou
ter
mar
‐

gin；
ischiu
m
4
5％
aslon
g
asbasis，w
ith
6setaeonlnnermargi
nand2setaeonoute
r
mar
g
ln；
mer
u
sal
most
as
lon
g
as
i
sch
i
u
m，
w
i
th
l
O
se
t
a
e
on
l
n
n
er
mar
g
i
nan
d
s
o
me
s
hor
t
se
t
a
e
ono
u
t
e
r
mar
g
l
n；
cal
p
u
s
as
l
on
g
as
me
r
u
s，
w
i
t
h
abou
t
20
set
a
e
i
ncl
u
din
g
6
‑
7bi
f
Ur
cat
e
don
e
sonlnnermar
gln；
propodusalittlelongerthancalpus，
wit
h8
1ongsetaeandmor
et
h
an
l
8shor
tsetaeonlnner
mar
g
ln，
andllsetaeonouter
mar
g
ln、
Per
eopo
d2（Fig2B):basis33ti
mesaslongasw
ide,with
7setaeonlnner
mar
ginan
d4setaeonouter
mar
gln
§
ischiumO､6timesaslongasbasis，withlOsetaeonmnermarginand2−3setaeonouterdistalareaぅmerus0．8times
aslon
g
as
i
sch
i
u
m，
w
it
h
8
set
aeon
mner
mar
g
i
n
an
da
se
t
aatou
ter
di
stalar
eaラ
car
pu
s
l
3t
i
meslongerthanmer
u
so
w
it
h
41on
gbi
f
Ur
cat
e
dsetae
an
d
4‑
5shor
t
e
r
se
t
ae
on
lnner
mar
gi
n
ll
‑
l
2se
t
a
e
o
no
u
t
er
ma
r
g
l
n
an
d
4s
e
t
a
e
at
o
u
t
e
r
di
st
a
l
an
g
l
e
う
p
r
o
po
du
sa
lit
t
l
e
l
on
g
e
r
t
h
an
c
a
中
u
s，
w
it
h
2bi
hlr
cat
edsetaeand3‑4setaeonmner
marginand9−
10setaeonoutermargln・

Pereopod3（Fig2C)：basis2．8timesaslongaswide,withlO‑l2setaeonbothmargins；ischium45％aslong
asbasis，withlOsetaeonmnermargmandasetaatouterdistalangle；merus0．65timesaslongasischium，with
9‑lOsetaeonlnnermarginandasetaatouterdistalangleうcalpusl､stimeslongerthanmerus，withaboutl2setae
inclu
din
g
3
‑
5bifUrcatedonesonlnnermargin，2relativelylongandsomeshortsetaemcludingbifnr
catedonesondis
‑
taI
mar
gln，
and8
‑
9setaeonoutermargln；
propodusalittlelongerthancaI
pus，
w
ithlOsetaeonbothmargins・
Pereopod4（Fig2D)：basis27timesaslongaswide,withl3‑l4shoItsetaejischiumhalfthelengthofbasiS
with5setaeonmnermarginand2−3setaeonoutermargln；merus0．7timesaslongasischium，with7setaeonlnner
margln；camus1．3timeslongerthanmerus，withlOsetaeincluding4bifUrcatedonesonlnnermargin，4−5setaeon
distalmarginand6‑8shortsetaeonmnermargm；propodusalmostaslongascalpus，with5setaeonbothmarglns。
Pereopod5（Fig2E)：basiswide,basis2．9timesslongaswide,with9‑lOshortersetaeonmnermarglnanda
longersetaatlnnerdistalangle；ischiumO､7timesaslongasbasis，withlO−12setaeonmnermarginand2setaeon
outermargln；merus0．4timesaslongasischium，with9−lOsetaeonlnnermarglnand2setaeonouterdistalarea』

carpus1．4timeslongerthanmerus，with7‑9setaeonlnnermargin，2−3setaeondistalmarglnandl5‑l6setaeon
outermargln；propodusalittlelongerthancamus，withlOsetaeonmnermarglnand6setaeonoutermargln･
Pereopod6(Fig2F）：basis27timesaslongaswide,withasetaatmnerdistalangleandlOsetaeonlnnermar‐gin；ischium65％aslongasbasis，with8setaeonlnnermarginand3setaeonouterdistalarea；merus0．55times

aslongasischium，with7setaeonbothmargln；carpus1．7timeslongel･thanmelus，withlOsetaeonlnnermargm，
3setaeondistalmarglnandl3‑14setaeonoutermarglnぅpropodusalittlelongerthancarpus，withl31ongerandshortersetaeonlnnermarginll‑12setaeonoutermargln
Pereopod7（Fig2G):basis3．7timesaslongaswide,with7‑8setaeonlnnermarginand3‑4setaeonoutermar‐
ginうischiumS5％aslongaswide，with3setaeonoutermargln；merus0．8timesaslongasischium，with5setae
onlnnermargin，4setaeonmnerdistalarea，2setaeonouterdistalareaand5−6setaeonoutermargl､；carpusl,Ztlmeslongerthanmerus，withllsetaeonlnnermargin，2setaeondistalmarglnand7‑8setaeonoutermargln号
propodusaslongascarpus，with3weakersetaeonbasalareaand3strongersetaeondistalareaofinnermargm，
aboutll‑l2shortersetaeonoutermargln．

41

NobomNunomura（FiglK)：exopodroundedtriangular,with2setaeonoutermargln
Pleopod lPleopod 2（FiglL)：exopodwith3‑4setaeonoutermargl、
Pleopod 3（FiglM):exopodwith2setaeonoutermargln（Fig｣N)：exopodtriangular,with6setaeonoutermargln
Pleopod 4

Fig.2．〃 αsmj火s 〃"e"姉，、｡sp，

A−G；Pereopodsl‑7(All：Holotypefemale)：４AnewL"c jo〃たs廿omAshiuJapan

PleopodS:endopod（FiglO）rectangularうexopod（FiglP）roundedtriangular,with5setaeonoutermargln
Uropod(Fig｣Q）:basistrapezoid；endopodaslongasbasis,withmorethanlOsetaeonbothsidesうexopodl4times
longerthanendopod，withmorethanl5setaeonlnnermarginand6−7setaeonoutermarglnandatuftofsetaeon
apicalmargln．
A姓J/e：Astopereopods，pleopodsandantenna，therearesomedifferenceslnmale・Antenna（Fig3A）terminalnagellarsegments2ltimeslongerthanthebasalone；mumallengthoftenninalsegment
slightlyshorterthanfbmale・
Peropodl（Fig3B)：basis29timesaslongaswide；ischiumhalfthelengthofbasis；merusO8timesaslong

aswide，withdenselvsetaeincludingbihlrcatedandtrifUrcatedsetaeonlnnermargln；carpus1．2timeslongerthan
merus，withdensesetaeincludingsomebihlrcatedandtrifilrcatedonesonmnermargln；propodusO､8timesaslong

ascamus，withaseriesofsetaeonbasalhalfofinnermar2inand2setaeondistalhalfofinnermargln・
Pereopod2(Fig3C):basis33timesaslongaswide;ischiumO,55timesaslongaswide,withl2setaeonmner
marginand2setaeonouterdistalarea；merusabouthalfthelengthofischium，withmanvsetaeincludingbihlrcatedandtrihlrcatedonesonlnnermargln；camuswithmanysetaeincludingbifilrcatedandtrifUrcatedsetaeonlnnermargln

setae；propodusO､8timesaslongascarpus，with9‑lOsetaeonlnnermargln・Peropod3（Fig3D):basis29timesaslongaswide;ischiumQ6timesaslongasbasis,with21ongsetaeondis‐
talareaandabout4setaeonlnnermarglnanddistalangle；merus2/3aslongasischiumwithmorethan20Setae
oninnermarginand2setaeonlnnermarglnぅcarpus1．3timeslongerthanmerus，withmorethanl5setaeincluding

bifilrcatedonesonlnnermarglnand2‑3setaeonlnnermarglnぅpropodus1．2timeslongerthancalpus，with8setae
onlnnermargln、Pereopod4（Fig3E)：basis28timesaslongaswide；ischium45％aslongaswide,with8‑10setaeonlnner

marginand3‑4outerdistalarea；meruswithmorethan20setaeonmnermarginincludingbifUrcatedonesラcamusl,S

timeslongerthanmems，withmorethan20setaeincludingbihlrcatedonesonlnnermargm；propodusalittlelonger
thancarpus，with6‑7bifilrcatedormfilrcatedsetaeonlnnermarginandl7‑l9setaeonoutermargln
Pereopod5（Fig3F):basis32timesaslongaswide;ischiumO6timesaslongaswide,with8‑9setaeonlnner

marginand4‑5setaeonouterdistalarea；merus0．6timesaslongasischium，withl2−13setaeonlnnermarglnand
3setaeonouterdistalareaラcaIpusl・stimeslongerthanmerus withl3‑14setaeonlnnermargln；propodusalittle
longerthanlOsetaeonmnermargmandmorethan20shortsetaeonoutermargin
Pereopod6（Fig3G):basis3.ltimesaslongaswide,withl5‑16setaeonmnermarginand8‑llsetaeonoutermargin：ischiumO,6timesaslongasbasis，withll‑l2setaeonlnnermarginand21onEbihlrcatedsetaeonouterdis‑

talareaうmems2/3aslongasischium，withlO‑l2setaeincludingbifUrcatedonesonlnnermarglnand2setaeon

outerdistalarea；calpus1．2timeslongerthanmerus，withlO‑llsetaeincluding21ongbihlrcatedonesonlnnermar‑
gln；propodusaslongascarpus，withl2−l4setaeonlnnermarginandl3‑l4setaeonoutermargln，Pereopod7（Fig.3H)：basis23timeaslongaswide,with2setaeatlnnerdistalangle；ischiumO､8halfthe
lengthofbasis，with3setaeonlnnermargln；merus85％aslongasischium；carpusaslongasmerus，posterlorparT

protrudedroundly，with8‑91ongerand7‑8sholtersetaeonlnnermargm，andasetaatouterdistalangle；propodus
aslongascalpuswith8setaeonlnnermarglnandl5−l6setaeonoutermargm・
Penes（Fig31）fUsifbnn

Pleopodl：endopod（Fig31）straightandapicalpartslightlybentsouterwards,withaseriesofdenticles；exopod(Fig3J）semicircular,innermarginslightlysinuateandoutermarginwith2setae
Pleopod2：endopod（Fig3K）longandtapermgtowardthetipandslightlyexceedingbeyondthetipofexopod；

exopod(Fig3L）triangular,withroundedtipand5setaeonoutermarginPleopod3（Fig3M):endopodwith3setae
onoutermarginPleopod4（Fig3N）exoopodwith3setaeonoutermarginPleopod5（Fig30)：endopodwith3
setaeonoutermargln．

4苫NobomNunomura

Fig.3．L"casmj火s 〃"e"卵sn.sp．

A，Flagellumofantennaofmale；B−H，Pereopodsl‑7ofmale；I，PenesandPleopodl；J，Exopodofthesame；K，Endopodofmalesecondpleopod;L,Exopodofthesame；M‑O，Exopodofmalepleopods3‑5（All：AⅡotypemale)E〃"20/ogy:ⅧAshiu"isthenameoftypelocality・

姓"' 肺：ThepresentnewspeciesismostcloselyalliedtoL〃jc〃加α"e"sjs（Nunomura，2003)，recorded廿om

Miyazaki，southemKyushu,Butthefbnnerlsseparated廿omthelatterinhefbllowingfeamres:(1)distinguishedcoler

pattem,（2）
longerter
minalnagellar
segment,（3）
presenceofmanybifilrcatedsetaeonpereopods,(4)
moreaesthetascs
ofantennule,（5）remotepositionofpereonalsomitesl−4and（6）lesssetoseonmerusandcamusofpereopodsl‑2
AcknowledgementS

lwouldliketoexpressmysmceregratimdetoDr，HiroyukiWatanabefbrhiskindnesslngivingmeachancetoex−
ammesuchinterestmgmaterials．

44

AnewL"c roi たsけomAshiuJapan

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Nunomura，N,、2000．TelTestrialisopodandamphipodCrustaceansfomthelmperialPalace，Tokyo，Mｾ脚．MJ Sa
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middleJapanB"" 7byα"7aSciM"& 31:3‑llNunomura，N､，2009．TerrestriallsopodCnlstaceans廿omDaitolslands，SouthemJapan．B""．7bva碗αSα，MiJJ，．32：75
87．

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SCIⅡnalhlss，H，2003.WorldCatalogofterrestrialisopod（Isopods：Oniscidea）Sr""垣α'7eγβe/".M""rkSerA,654
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