-[
UAt-t
t\1

Rev. de Entomologla, vol. lQ faso. 8, dezembrp19if9

Recordsand descriptionsof Tipulidaefrom
tropicalAmerica(Diptera). Part ll..

byCnenunsP. Ar,uxaNDER

Rio de Janeiro - Brasil

fromtropicalAmerica
ol Tiputidae
anddermiptions
Rccords(Diptera).Part llnntt
by Cnlnuns P. At,r,;xalv
MassachusettsState College, Amherst, l\Iass.
tWith 16figures)

The speciesdiscussedin this report were all taken insouthern Costa Rica by my former student, Mr. D e a nn as madeL e o n a r d R o u n d s . T h i s i n t e r e s t i ncgo l l e c t i o w
in the Valley of the General,Talamanca,at altitudesbetween
2000 and 3000 feet, in January 1939. A few specieswere
taken at a slightly higher altitude at San Juan Miramar, incloud forest on the Pacificcoastalrange.I am greatlyindebted
to Mr. Rounds, and to the ornithologistand botanist,
D r . A l e x a n d e r S k u t c h , f o r t h e o p p o r t u n i toyf s t u d y i n g
this collection,which has addedgreatly to our knowledgeof
distributionof thes'efli,esin southernCosta Rica. All species
mentioned in this paper are new to the Tipulid fauna of
the republic.Of specialinterestis the discoveryof a speciesof Dicranoptyclta,not before known from the Neotropicalregion. All material is preservedin my collectionof theseflies.
D i c r a n op t y c h a c o s t a r i c e n s i s , s p . n .
Generalcolorationdull brownishblack,the thoracicpleuravariegatedwith paler;antennae,
halteresand legsblack;u'ings

Rev. de Entomologia,vol. 10, fasc. 3, dezembro1939

623

narrow, very strongly blackened;cell lst M, open by atrophyof basal section of Msi m-cu mor,e than its own length
beyond the fork of M; male hypopygiumwith outer dististyle
narrow, gradually attenuatedto a slenderapical point; outersurface of style roughened by setiferoustub,erclesbut nct
spinous.
M aI e.
Length, about 8.5 mm.; wing, 8.6 tnm.
Rostrum and palpi black. Antennaeblack; flagellar segments oval, with truncated ends, the outer segmentsmore
elongate;verticilslong. Head brownish black; anteriorvertex
wide.
Pronotum yellowish brown, darker on sides.Mesonotum
uniformly dull brownishblack. Pleura paler brorvn,the ventral
anepisternumwith a narrow, r'elvety black, Iongitudinal linethat barely involves the anterior end of the pteropleurite;
posterior sternopleurite,meral region and pteropleuritepaler,more brownish yellow. Halter,eselongate,black, the base ofstem restrictedlypale. Legs with the coxae obscureyellow;
trochantersblack; remainderof legs black, elongated.Wiirgs(Fig. 1) narrow,very stronglyblackened,
the veinsand trichia
still darker.Venation:Rs in longitudinalalignmentwith basal
section of R+*u and only a little longer than this element;
Rz+e running close to Rr, especiallybefore vein Sc,,,thence
gradually diverging; Rs faint, without trichia; cell I st A42
open by atrophy of basal section of A43,cell 2rtd M, deep;
basal section of Mr +t angulatedand u'eak-spurredat nearmidlength; m-cu more than its own length beyond the forkof M; cell 2nd A narro\\'. The usual p.ale fold in outerhalf of cell Cu is indicated in figure by stippling.
Abdominal tergites black,sternitesbrownish yellow. Male
hypopygium (Fig. 9) with the outer dististyle, od, narrow,gradually attenuat'ed
to a sl'enderapical point, the extremetip
decurved;outer surfaceof style feebly roughenedb-ysetiferous
tuberclesand setae but rvithout spines.Inner dististyle,id,
relatively broad, the apex obtuse,the surface with abundantcoarse setae. Apices of lobes of gonapophysesbroadl-vexpanded,th,e margins smooth. Aedeagus,a, narrow, the apex
obtuse.
Holotype,
6 , S a n J u a n M i r a m a r ,i n c l o u d f o r e s t o n
Pacific coastal range, altitude 3,500 feet, January 1939
(Rounds).

62{

A l e xa n d e r,

Tiputidaefr om tr opical Ar ner ica.II.

Dicranoptl'cha rosturicensisis entirely different from all knolvn
speciesof the genus. It is u'ell-distinguishedby the black coloration
and b1' thc narrorv wings, with cell Ist M, open by the atroph-vof theb a s a l s:cti o n o f fu l r.A s indicatedin the intr oductor ypar agr aph,thegenus had not hitherto been discoveredin Tropical America.

T e u c / r . oI a . bi s ( T e u c l r c l a b i st)a l a m a n c A n a , s p . n .
General coloration black, including the praescutumand
postnotum;scutellumyellow; knobs of halteresycllow; femora
yellow, the tips conspicuouslyblackened,on fore legs involving
about the distal two-fifths, on posterior legs much narrower,
including about the outer fifth; tarsi black; u'ings yellow,
sparselyvariegatedwith darker, including a narrow seam at
cord and an interruptedband at level of origin of Rs,' u'ingtip paler brorvn; m-cu beyond fork of M; abdominal tergites
black, the sternitesdimidiate,black, the outer half of individual
segments yellow.
Female.
L e n g t h ,a b o u t 6 m m . ; w i n g , 5 . 4 m m .
Rostrum brown; palpi black. Antennae brownish black;flagellar segmentssubglobular,passingthrough short-ovalto
oval. Head brorvnish black, sparsely pruinose
Pronotum polished yellow. Mesonotal praescutum uniformly polished black, very restrictedlyvariegatedmediallyrvith yellow at suture; scutellum and parascutellayellorv;
postnotumpolishedblack. Pleura chiefly polishedblack, including the entire mesepisternum
and dorsal pteropleurite;dorsopleural membrane, ventral pteropletrriteand meral region
yellow. Halteresbrorvnishblack, knob light yellow. Legs withcoxaeand trochantersyellou'; femora yello,w,the tills conspicuously blackened,on the slightly dilated tips of fore fernora
including about the distal two-fifths of segment,on posteriorlegs narrower,involving about the outer fifth or sixth; tibiae
yellow, the extremebasesand slightll' u'iCer tips black; tarsi
black. !7ings yellow, sparsely patternedu'ith darker; wingtip paler bron,n, including about the outer sixth, the inner
margin of area not reachihg the outer end of cell Ist Mz;
darker brown seamsat stigma, confluentwith a narrow bandat cord, and at outer end of cell lst nlr; a very narrowbroken band at level of origin of Rs, including small clouds
at origin of Rs, tip of 2n:l A, anC a linear rvash in cell Cu;
veins brownish black, more yellowish in the prearcularfield.
Venation: Sc relativelyshort, Sr, ending about oppositetwofifths the length of (s; Re oblique; R2-i-sal short; branches

Rev. de Entomologia, vol. 10, fasc. 3, dezembfo 1939

625

of Rs parallel to one another for more than their proximal
half ; m-cu more than one-third its l,engthbeyond fork of M.
Abdominal tergites black, the lateral posterior angles
slightly paler; sternitesbicolor'ed,the basesblack, the outerhalf of individualsegmentsyellow; subterminalsternitesmore

3

R'

5r

bi.'
Fig. l. Dicranopt!tc\a costai'ieercis, 5p. n. I venation. -- Fig. 2. Gonomyia (Gonomyi'9,).
iir. "r:"p1g._3, Gnopl.tgmlri lGnoph.omyta)pg"tica1a,- 8P. -D,: venation. .-.Iig.,-1.
'C,:ripiothU,ii
7"7iifer,(Cr11ptol7bia7rot,hrlsi, .p. n. vination. -- fig. 5. .Cryptolabis (Cryptolabis) di'r e t s i p c s , s p . n . ; v e n a t i o n . F i g . d - a . T h e s a m e , s h o w i n'-g " v a r i a t i o n i n v e n a t i o n .
4g-'.0,(('nlptolabis)
7.
Crlpl,olabi*(Ciuptot.obis)
Fig.
vbnatiorr.
n.
r;rrtftothtig
sp.
uatlpes,;"'inaaripes,'ve"riation.
Fig.'8. Toiorhiia l,Tororhinu) n{griuena, sp. n. ; r-enation.

extensivelypale; genitalshieldblack; cercistronglyupcurved,
horn-yellow.
H o l o t y p e , ? , S a n J u a n M i r a m a r ,i n c l o u d f o r e s to n
Pacific coastal range, altitude 3,500 feet, january 193q
(Rounds).
Teucholahis('t eucholtbis) talamanranais most generallv similar
to speciessuch as T. (f .) equinigra Alexander, of northertt Panama,
the latter dif fering especiall-vin the coloration, as the narrow, subequal, blackenecltips of all fentora, the pale basitarsi,scarcely evidentI626

A l e xa n d e r,

Tipulidaefr om tr opical Ar ler ica.Il.

dark u'irrg-tip,more nearly entire dark basal bancl of u.ing, lncl theblackeneclscutellum.

Gonomyia( O o n o n t y i ab) i i u r c i l e r ,sp.n.

General coloration dark brown; antennac,halteresandlegs dark,ened;wings with a weak dusky tinge, the stigma
a trifle darker; scr ending opposite origin of Rs; Rs andR r + r a r subequal; m-cu more than one-halfits length beyond
fork of M; male hypopygiunr with the outer dististl'le a
blackenedrod, the long apical spine attenuate;inner dististyle
deeply bifid, the outer arm darkened,widened outwardly, thetruncatedmargin erose; phallosomea powerful central blackened rod, at apex split into two acute spines.
M aI e.
Length, about 4 mm.; wing, 4.2 mm.
Rostrum and palpi black. Antennae black throughout.
Head black, sparsely pruinose.
Mesonotum dark brown, the scutellum a little brighter.
Pleura yellow. Halteres infuscated.Legs with coxae brorvnish
testaceous;trochantersbrown; remainderof legs dark brown
to brownish black. wings (Fig. 2) with a rveak dusky tinge;
stigma a trifle darker; veins dark brown. venation: sc relatively short,Sc, endingoppositeorigin of Rs; Rz+ r.i r Subequalin length to Rs; basal deflectionof R5 short; m-cu.more than
one-half its l,ength beyond fork of ,M.
Abdomen dark brown; hypopygium paler brown. Male
hypopygium (Fig. 10) with the outer dististyle,od, a blackened simple rod, a little expandedat near midlength, thence
drawn out into a long acute point. Inner clististyle,irl, deeply
bifid, the outer arm a darkenedlobe that is widenedoutrvardlr',
the apex truncate; surface with microscopicroughenings,theapex erose;befor'eapex with a single long seta; inner arm alittle longer but narrower,pale,with nLunerous
setae,including
two very long fasciculate apical bristles. Phallosome,p, apowerful central blackenedrod, or two very closely applieclrods, at apex split into two slightly unequal spines.
Holotype,
6 , P e d r e g o s oa,l t i t u d e2 , 0 7 i f e e t , J a r r u a r ' _ y
1 9 3 9( R o u n d s ) .
Gononi.t'iu (uonom),ia) bilurc'iler is entirell' clistinct from otherregional species, dif fering especially in the stru,:ture of the rnale
h-vpogygium.

Go nomvia( L i p o l t h l e p s a) e q u i d e n s

Alexandcr

sart Ramon,altitude2,900feet, Januarylg3g (Rouncls).

Rev. de Entomologia,vol. lQ fase.3, dezerhbro1939

627(Lipophleps)peracutaco nile ra Alexander
Rivas, altitude 2,875 feet, January 1939 (Rounds).

G onomyia

C o no m y ia (Lipopttleps)m a cs w ai ni Alexander
altitude 2,075feet, January 1939 (Rounds).
Pedregoso,( L i p o p h l e p sc)a l v e r t ip l e u r o s t r i a t a ,subsp.n.
Length,about 3 mm.; wing, 3.5 mrn.
M aI e.
Differs from typical calverti Alexander,likewise from
Costa Rica (Journ.N. Y. Ent. Soc.,22: 122; 1914) in theheavily stripedthoracicpleura and in the detailsof structureof the male hypopygium.
Antennaeblack throughout.Thoracicpleura dark brown,s'ith a broad whitish longitudinalstripe extendingfrorn behindthe fore coxaeto base of abdomen;dorsopleuralregion andadjoining pretergiteslight sulphur yellow. Legs medium torelativelydark brown. Wings with a weak brown tinge, thestigma a trifle darker; prearcularand costal fields moreyellowish. Venation: Sc short, the distancebetween tip ofSr, and origin of Rs subequalto the total length of thelatter, Sc, near tip of Sc1; branchesof Rs strongly divergent
at outer ends; m-cu at fork of M. Male irypopygiurn(Fig.
1I ) rvith the dististyle,d, fleshy, not different in texture
P, asfrom the basistyle,D, the latter slender.Phallosotne,
figured, the upper plate consistingof trvo terminal blades,
one long ancl slender,pale, nearly straight,the other a flatteneclswordlike blade; below this plate, on eith'erside witho scooplikestructuremarginedwith about nine powerful'blackspines,the trvo tnore basalones weakerand more slender.
Gon omyia

of calverti Alexander,rvhich
I regard this as being a subsltecies
cliffers especiallyin the scarcelv striped thoracic pleura and in theabruptll' narrott,ecl,lnore sclerotizeddististyle. The remarkablephallosome of the present fly seemsto be entirely honrologizablewith thatof cahlerti, the most peculiar single feature lying in the series ofstrong blackened spines on either side of the lobes of the ventral
plate. Irt order to avoid confusion, I have figured this plate as separate
from the remainder of phallosorne and have shown the spines of
one side onl-r'.

altitude2,200feet, January
Holotype, 6, P'edregoso,
1 q 3 9( R o u n d s ) .

A l e xa n d e r,

Tipulidae fr om tr opical Anr er icn.II.

G o n o m y i a ( I - i p o p h l e p sh) a p l o i d e s A l e x a n d e r
Pedregoso,altitude 2,075-2,500
feet; Qivas,altitude 2,875
feet, January 1939 (Rounds).
G n o p h o m y i a ( A n o p h o m y i ap )o s t i c a t a , s p . n .
Belongs to the luctuos,cgroup; size srnall (rving, male,under 5 mrn.); general colorationblack, the posteriortibiae
paler brown on centralhalf, posteriorbasitarsiobscureyellow;
wing.sblackish,rvith pale longitudinal streaksin cells R andCu; numerousmacrotrichiain outer cells of wingi Rs at orclose to fork of Rz* B-r-{ , male hypopygiumwith the phallosomea widely expandedplate, its basal portion with abundant setula,e;gonapophyses
appearingas slender pale rods.
M aI e.
Length,about 4 mm.; wing, 4.4 mm.
Female.
L e n g t h ,a b o u t 5 . 5 m m . ; w i n g , 5 m m .
Rostrum black, sparselypruinose;palpi black. Antennae
black; flagellar s€gmentsoval, with verticilsof moderatelength.
Head black, sparselypruinose,more heavily so on front.
Thoraxblack,the notumsparselypruinose,the pleuramore
heavily so. Halteres black throughout.Legs black; posterior
tibiae pal,erbrown for about one-halfthe total length before
the blackenedtips; posteriorbasitarsichiefly obscureyellow,
the tips darkened.U7ings(Fig. 3) with a strongblackishtinge,
with pale str'eaksin cell R adjoining vein tr(, crossingcell
Ist /W2 into cell Ms; a second such pale streak in outerhalf of cell Cu; cell C before arculus similarly pale; veins
black. Macrotrichiain centersof cells beyondcord (indicatedin figure by stippling). Venation:Sc, ending nearly opposite
or shortly before fork of Rs, Sr, not far from its tip; Rs
long, exoeedingRs,' Rz at or close to fork of R:i- r * r i cell
Ist A4z relatively short, not exceedingone-third the lengthof vein Mt*o b,eyondit; m-cu about one-thirilits length
beyond fork of /vl.
Abdomen,including g'enitaliaof both sexes,black. Male
hypopygium (Fig. 12) with the basistyle,D, near apex producedinto a long fingerlike lobe directedcaudadand mesad.
Tn'o dististyles,id, od, slightly unequalin length and width,
both a little expandedat outer ends. Phallosome,p, withlateral flanges that ar,e widely expandedinto a depressedplate, this denselysetuliferous(punctur,es
indicatedon figure
by stippling); apex of aedeagusa slender recurved hook.
g, appearingas long slenderpale rods.
Gonapophyses,

Rev. de Entomologia, vol. 10, fasc. 3, dezembror$9629

2,300feet, January
raltitude
Holotype, 6, Pedregoso,
1 9 3 9( R o u n d s ) .
A l l o t o p o t Y P e , ? , a l t i t u d e2 , 0 7 5f u e t ,January1939(Rounds).

l3

r5

l0
-

FiS. 10. Gonomyia-(Go.-.
-calutti"Gonomyiq-y7nonltteps)
Fig. 9. Dicranoptycha costat'ic.cncie,sp. n. ; maleLrypopygiurn.
-.hial-ur"rciier',.p ;:lmal;- i,ti,o-6i.sl:irh.-_r,gl^t\llGitophornvialpostiin;i; uvpopilfi*. - rig.-tz. Gttoythotriy-i.o;i;i;;;;lc,-i"uip.'";"#iti-::i#:{;yifi#:i},,.#'{fii'i{!,vr:i#;ii";:,{::'#!'*,-il;'o;ilt,t',:',"#L*,,9d#1rlif;;r;.Hs1"i?llgili,!
innerdietisrvre;
ltiot"tiiittuj'5y71oit"tlrfri;p.;:;b;-b;fi;6';';mai6 fivp6pygiirur.- tnirrlariritYonol svmbols: rr, adeag.us;

r'a'Anopltontyia (Anophomyia) posticata is quite different from otherclescribecllrembers of the luctuosa group, especially in the small size,
coloration of the legs, and in the details of structure of the malehypopygium.

63,)

Alexander,

Cryptolabis

Tipulidaefronr tropicalArnerica.II.

( C r y p t o l a b i sr)o u r t d s i ,sp. n.

Sizelarge (wing, male, 5 mm.); generalcolorationyellow,
the praescutumwith four dark stripes; pleura yellow witha conspicuousdark brown longitudinal stripe; femora andtibiae obscureyellow, the tips dark,ened;u'ings pale brown,subhyaline,unmarked;Sr very long, Srr. ending nearlv opposite the fork of the long sinuousRs; m-cu just beyond midlength of Mu 14i rnol.' hypopygiumwith the tergite terminat.
ing in two divergentsubmedianlobes; basistylecomplex,atouter end bearing a d,ensebrush of long yellow setae;
dististylecntirelypale,the apex weakly expandedinto a head;aedeagusstrongly convoluted.
Male.
L e n g t h a, b o u t4 m m . ; w i n g , 5 m m .
Female.
L e n g t h ,a b o u t 4 m m . ; w i n g , 4 . 7 - 4 . 8m m .
Rostrum yellorv; palpi dark brown. Antennaervith th:scape and pediceldark brown; flagellum testaceousyellorv;
verticils long and conspicuous.
Head pale yellow.
Pronotumyellorv above,brown on sides.Mesonotalplaescutumyellow u'ith four brown stripes,the lateral pair slightly
paler than the interm'ediateones; scutum yellow, each lobe
rvith two conspicuous
dark brown areas; scutellumbrownish
testaceous;mediotergitedark brown; thorax conspicuously
hairy. Pleura yellow, with a broad, conspicuous,
dark brown,
longitudinalstripe extendingfrom thc cervi-^alregion acrossthe dorsal pleuritesto the base of abdomen,passingabove
the root of the halteres.Halteres dark brown, the base ofstem yellow. Legs rvith the coxae and trochantersyellow;
femora obscureyellow, the tips narrowly and graduall-vdark
brown; tibiae pale yellow, the extremetips darkened:outertarsal segmentsinfuscated.
Wings (Fig. a) \'er,vpale brownish
subhyaline,the pr,earcularand costal f ields clearcr yellow;
veins pale brown, cleareryellow in the flavous portions.Macrotrichia in outer cells of wing, more extensivein female
than in the male (as shown, the punctunesrepresentedby
stippling); in female,the trichia occurin all cellsbeyondcord
as well as in most of cell Rr and outer ends of Cubital andAnal cells. Venation:Sc unusuallylong, Sr,. ending just before outer end of the long sinuousRs, the latter in longitudinalalignment with Ro; Sc, about opposite or before midlengthof Rs; r-m at or just before th: fork of Rs; m-cu just beyond
midlength of M, +,Abdominal tergites brown, the caudal borders narrowly

Rev. de Entomologia, YoI. 10, fase. 3, dezentbro1939

63r

blackened;sternitesand hypopygiumpale yellow. Male hy'
popygium(Fig. 13) complex.Ninth tergite,9t, with two subr.di"n curv,edlobes,directedcaudad,the tips slightly laterad,

the surface with abundantsetae.Basistyle,b, terminatingin
a flattened black lobe, its apex obtuse,at base on inner sidervith .a densegroup of long yellow setae;face of style laterad of the hair-tuft with a strong straight spine. Dististyle,d, subterminal,entirely pal,e,the apex a little expanded,thesurface with abundant delicate setae and setulae, those on
the head retrorse.Aedeagus,a, long, blackened,strongly bent
itself or convoluted.
2 , 9 2 5f e e t ,J a n u a r y1 9 3 9H o l o t y p e , 6 , R i v a s ,a l t i t u d ' e(Rounds).
altitud'e2,075feet, Januarl'
Allot)'pe, Q, Pedregoso,s a l e sa n d f e m a l e sw, i t h
1 9 3 9 .P a r a t o p o t Y P € s , n u m € r o um
feet, January
the type; with the allotype, altitude 2,075-2,300
1 9 3 9( R o u n d s ) .
in
I take great pleasure- in naming this very distinct crane-fly'IheSpe, r. Dean Leonard Rounds.
h o n o r o f t h e c o l l e c t o rM
cies is entirel-y clifferent f rom the other described regional forms havand ining Rs long and sinuotts, especially in the bod,'--colorationt h e s t r u c tu reo f th e In a l e h -vpopvgium 'C r 1 ,p t o t a b i s ( C r v p t o l u l t i sc )li v e r s i p e s , s p ' n 'Generalcolorationgrayish black, the dorsopleuralre$on
yellow; femora black,'thebasesnarrowly paler; tibiae obscuretarsi
1'ello5',the extremebase and much wider tip blackened;
Lladt, the posteriorbasitarsiobscur'eyellow to brownishyellow
on basal two-thirds; wings tinged with brown, the prearcularand costal regions clear light yellou'i Rs moderately long'sinuous; male hypopygiumwith the dististyl'ebilobedat apex;
aedeagusstout, not convoluted.
m m ' ; w i n g , 4 ' 3 - 4 ' 5m m 'L e n g t h ,a b o u t3 . 5 - 3 . 8Male.
L e n g t ha, b o u t4 m m . ; w i n g , 5 m m 'Female.
Rostrum and palpi black. Antennae with scape black,pedicel and f lagellum dark brown; flagellar segmentsoval,
the outer segmentsprogressivelysmaller. Head dark brown'
Pronotum whitened above, grayish black on sides.Mesonotal praescutumgrayish black, with a narrow paired darker
medianstripe, the lateral bordersand dorsopleuralregion restrictedly yellow; scutal lobes blackishgray, the median area
brownish yellow; posterior scleritesof notum blackish gray.
Pleura blackish gray. Halteres pale yellow. Legs with the632Alexander,

Tipulidaefronr tropicarA'rerica.Ir.

coxae black; trochantersobscureyellow; femora black, very
narrowly paler at base; tibiae obscureyellow, the extreme
baseand much wider apexblackened;tarsi black,the posterior
basitarsi obscurey,ellow to brownish yellow on basal twothirds.vings (Fig. 5) with a slight brown tinge,the prearcularand costalregionsclear light yellow; axillary region narrowly
darkened;veins pale brown, cu and those at cord darkei,
insensiblyseamedwith fuscous.Abundantmacrotrichiain cells
beyond cord (their positi,onindicatedin figure by stippling);
in female,th'etrichia equallyabundantbut lackingin the basesof the cellsbeyondcord. venation:sr, endinga ihort distance
befor'efork of the moderatelylong, ,lnuous fts; r-m variable
in length and position, as is also the position of m-ilt in
relation to the fork of tvr+ ai in cases,i-m is at fork ofRs,in others on R51Sorn€distanoebeyondorigin m-cu at forkofiM? * + or some distance before this fork (the extremesof
this conditionare shown in figures 5 and 5 a).
Abdomen,including the hypopygium,brack. Mare hypopygiurn (Fig. 14) with the dististyle,d, terminal in position,
conspicuouslybilobed at apex, one lobe pointed at tip, theother obtuse; surface of style hairy. Aed,eagus,
a, stout, notconvoluted.
H o I o t y p e , 6 , R i v a s ,a l t i t u d e2 , g 7 5f e e t ,J a ' u a r 1 .l g 3 g(Rounds).
Allotyp!,? , P . e d r e g o saol,t i t u d e2 , 1 0 0f e e t , J a n u a r y
1 9 3 9( R o u n d s ) .
Paratopotypeso, f both sexes,with the type; paratypes, both sexes, altitude 2,075-2,875
feet, Januarl, 1q3g
(Rounds,).
Cryptolabis (cryptotabisl tliversipes is well distinguished tr). thecololation of the body, legs ancl rvings, and, especiallv,bythe structureo f t h e rn a l e h 1 'p o p vg i u m .

Cry ptolabis( C r y p t o t a b i sr )a r i p e s ,sp. n.

ceneral coloration of rnesonotumr,eddishbrorvn, r.ariegated rvith white; antennalflagellum pale yellow iho.acic;
pleura with a conspicuous
brownish black, dorso-longitudinal
stripe; halteresyellow; fore femora blackened,remai-ningfemora yellow with the tips narrorvlyblackened;tibiae anclbasitarsi yellow, the tips narrorvlydarkened;*,ings yellowishsubhyaline,cellsc and sc cleaner
yellow; narrow dark seamsalong
cord and vein cu and in the axilla; Rs relativelyshort; veinsRer,.de Entomologia,vol. 10, fase.3, dezenrbro1939

633

M1 .l-zr Ms and M4 all close together at fork of M; malehypopygium with dististyle a simpl,e,subcultrate blade.
Length,about 3.5-3.8mm.; wing, 4-4'2 mm'
M aI e.
L e n g t h ,a b o u t4 m m . ; w i n g , 4 ' 5 m m 'Female.
Rostrumbrown; palpi darker brown. Antennaewith scapeand pedicelbrownish black, flagellum pale yellow, the outer
segmentsa trifl'e mor,e obscured.Head china-white.
Pronotum white, narrowly darkenedmedially. Mesonotal
praescutumwith virtually confluentreddishbrown stripes,theirumeral region and broad lateral borderswhite; scutal lobes
similarly t.idith brown; scutellumpale medially,darkenedonsides, parascutellabrown; mediotergitedark brown, the cephalic lateral angles and adjoining portions of pleurot'ergite
whitened.Pleura yellow, with a broad,bro$'nishblack,dorsolongitudinalstripe. Halteresyellorv. Legs with the coxaeand
the bases
yellow; fore femorablackened,trochanterstestaceous
paler, in caseswith a vaguely brightenedsubterminalring;iiUiu. yellow, the tips narrowly blackened,in casesrvith amore or less distinctdarkeningbeforemidlengthof segment;
tarsi yellow, the terminal segntentsclarker;middle and hindfemora light yellow, with the tips very narrowly blackened;
tibiae and basitarsiy'ellow,the tips narrowly darkened;outertarsal segmentsweakly infuscated.Wings (Fig. 6) yello'*'ish
subhyalini,th,. costaland prearcularregionsclearlight yellow;
narrow brown clouds at cord; cell Ctt, narrou'ly darkened;axilla clourled;veins ligth yellorv, d;rrker in the infuscatedportions. Macrotrichiapale, including the punctures,w'elldistributed in cells beyond cord. Venation: Sc lottg, Stt
endingnearlyoppositethe outer end of R, + s+ rj Rs relatively
short, weakly convex;basalsectionof Rs distinct;arrangementof veins at fork of m'ediapeculiar, with M, +2, MB and /t44
all arising close together at the cord, m-cu thus connectingwith M, m,orethan one-half its length beyond fork of M;
in rarer cases,m-cu is at or closerto fork of Mu+ t In theallotype, the basal section of Rb is lacking.
black subAbdomenobscureyellow, with a conspicuous
lateral stripe;genitaliaof both sexesyellow. Male hypopygium(Fig. 15) with the dististyle,d, d simplesubcultrateblade,thetip acute; surfaceof style only rnoderatelyhairy. Aedeagus,
a, stout, not convoluted.
H o I o t y p e, 6, Rivas,altitude2,875feet, January,1939(Rounds).

63{

Alexander,

Tipulidaefrom tropicalAmerica.II.

A I I o t y p e, ?, Pedregoso,
altitude2,100feet, January
1 9 3 9( R o u n d s ) .
P a r a t o p o t y p € s , m a l , e sp; a r a t y p e bs o
, , t hs e x e sw
, ith
the allotype,altitude2,075-2,300
feet, January1939.
Cryptolabis (Cryptolabis) varipes is u'ell-clistinguishedfrom allother described regional species by the coloration of the body andlegs, the venation, and by the structure of the male hypopygium.

Cryptolabis( C r v p t o t a b i isn) v a r i p e s , s p . n .
Generalcolorationblack,the posteriorscleritesof mesonotummoregray pruinose;haltercsblack;legsbrownishblack,the femoral basespaler; wings strongly tinged with dusky,
the outer veinsbroadly seamedwith still darker; macrotrichia
of cells restrictedto distal third of wing i Rs sh,ort,convex;
cell MB deep; abdomendark gray; genital region brownish
black.
Female.
L e n g t ha
, b o u t4 m m . ; w i n g , 4 . 5 m m .
Rostrum and palpi black. Antennaervith scapeand flagellum black, pedicela little paler; flagellar segmentsoval,
with conspicuousverticils. Head dark gray.
Mesonotumopaqueblack,the pretergitesobscureyellow;
scutellumand postnotumdarker gray pruinose.Pleura black,pruinose;dorsopleuralmembranerestrictedlvpale. Halteresblack, the extremebase of stem pale. Legs rvith fore coxae
black, middle and hind coxae,with all trochanters,yellowish
testaceous;femora obscureyellow basally,passinginto darkbrown; tibiae and tarsi brownishblack.Wings (Fig. 7) witha very strong duskytinge, especiallyadjoiningthe veinswhere
it appearsas conspicuous
seams,the oentersof the cellsremaining paler but still dark,ened;prearcularfield and cell C more
subhyaline;wing-apex narrorvly borderedby rvhitish subhyaline;axillarestrictedly
infuscated;r'einsdarkbrown; whitish
streaksalong r,ein,M in cell R, crossingthe cephalicportionof cell M, to vein Mr; vein Ist A similarlyborder,edby pale.
Macrotrichiaof cells r,estricted
to distal third of wing, beyond
the cord, with th,eexceptionof outer end of cell Ca. Venation:
Rs relativelyshort, convex;cell M, deep; m-cu beyondtrvothirds the l,engthof A[s+ti vein MB angulatedat base.
Abdomen dark brown; genital region brownish black.
Ovipositor with the major valvesfleshy, each providedrvith
about five large spinous s,etae.Abdomen filled with large
black eggs of unusualsize.

Rev. de Entomologia,vol. 10, fasc. 3, dezembro1939

636

o l, t i t u d e2 , 1 0 0f e e t ,J a n u a r y
Holotype, ?, Pedregosa
1 9 3 9( R o u n d s ) .
bY
CryptoluLis (Cr1'ptolabis) invaripes is readily distinguisheddark
the coloration of the body and wings, especiallv the broad
seams of the outer rving veins.

A4otopht!tts(Molophilus)subIictor'sp'n'Belongsto the plagiatusgroup; size small (wing, male,
3.5 mm.); mesonotalpraescutumdark reddish brown, theposterior scleritesof notum and th: pleura dark brown; legs
dark brorvn; wings rvith a brown tilge, the stigmal region
more suffused;r,Jin 2n:l A long; male hypopygiumwith thebasaldististylea flattenedrocl with a little lessthan the apicalhalf slightly more expanded,the inner face near apex with
spinulaeon outerdenseerectspines;a feu' smallerappressed
margin near base of expandedportion; apex of phallosomic
plate very gentlY concave.
M a l e . - L e n g t ha, b o u t2 . 8m m . ; w i n g , 3 . 5m m . ;a n t e n n a ,about 1.3 mm.
Rostrum and palpi black. Antennaeof moderatelength,
about one-halfthe length of body, black throughout;longest
verticils of more basil f lagellar segmentsver,\' long, f ully
three times the segmentsthemselves.Head bro$'nish black.
Mesonotal praescutumand scutum Cark reddish brown,t|re former ,rroo. blackenedlaterally behind the ps'eudosutural
foveae; posteriorscleritesof notum a little darker' Pleuradark
broq'n. Halteres rvith stem obscure yellow, a little darker
outu'ardly. Legs rvith coxae testaceousbrown; trochantersobscureyello$; remainderof legs dark brown. wings witha brown tinge, the stigmal region more suffused; veins andtrichia darker brown. Venation:Rs in approximatetransversealignment rvith r-m; petiole of cell MB about one-third to
oni-f,"lf longer than m-cu; vein 2nd A relativelylong, ending
about opporit. one-third th,e length of the petiole of cell z?13.
Abdomen, including hypopygium,brownish black. Male
hypopygium (Fig. 16) with the apical beak of basistyle,D,
slender,acute at tip. Outer dististyle,otl, with the outer arm
long and slender,narrowedto a slenderattenuateapex; inn'er
ann shorter, flattened. Basal dististyle, bd, a flattened rod,with a little less than the apical half slightly more expanded
than the base;inner faoe near apexset with denseerect spines;
outer face at base of expandedportion with much smaller

636

A l e xa n d e r,

Tipulidaefr onr tr opical Ar uer ica.II.

appressedspinulae; a few scatteredsetigerouspunctureson
outer half. Phallosomicstructure,p, a depnessedglabrousplate, the apex very gently concave.
Holotype,
altitude2,700feet, January
6, Pedregoso,
1 9 3 9( R o u n d s ) .
Most sintilar to Molophilus ( Molophilus) lictor Alr.xancler, ofColombia, which differs in the larger size and in the details of
structure of the male hypopygium,especiall.vof the basal dististvle andphallosomic
theplate.

S l . yr i n g o m v i aame ri cana Alexander

Rivas, altitude 2,875 1eet, January 1939 (Rounds).
7 ' ox o r l t i n a( C e r a t o c h e i l u sa
) nte ric ana (Alexander)

Pedregoso,altitude 2,500 feet, January 1939 (Rounds).
Toxorhina( T o x o r h i n a )n i g r i v e n A ,sp. n.

General coloration of thorax dark plumbeous; rostrum
about one-half the length of body, black throughout; mesonotal
praescutum with narrow dark'er lines on interspaces;pleura
dark, the meron and posterior coxae abruptly whitish; legs
black; wings whitish subhyaline, the cells be1'ond cord
somervhat darker; veins black; Sc short, Sr, cnding justbeyond origin of Rs,. R1 ending opposite midlength of Rs,m-cu at fork of M; cell 2nd A relatively narrow.
Female.
L e n g t h , e x c l u d i n gr o s t r u m ,a b o u t 6 m m . ;wing, 4.5 mm.; rostrum, about 3 mm.
Rostrum about one-halfthe length of body, black throughout. Antennae brownish black. Head gray.
Mesonotum dark plumb,eousgray, the praescutal interspacesnarrowly lined with more blackish.Pleura dark gray,the meron and posterior coxae more n'hitened.Haltenesblack.
Legs with the coxae darkerred basally, the apices pale, theposterior pair 'entirelypale, as above described;all trochantersabruptly black; remainder of legs black. Vings (Fig. 8)
whitish subhyaline,a trifle more darkenedbeyond cord; veins
black, those at wing-base a little paler. Venation: Sc very
short, Sr, ending immediately beyond origin of r(s, Sc, about
twice this distance before origin i Rr ending opposite midlength of Rs; cell Rb narrowed at wing margin, widest atnear one-third the length; cell I st Mz relatively large, m-cu
at fork of M; cell 2nd A relatively narrow.

Rey, de Entomologla,iol. lQ fasc. g, dezembro1939

687

I .mOomenbrownishblack,the more basalsternitespaler;of ovipositor v€ry long, especiallythe cerci.
Holotype, 9, San JuanMiramar,-inclgUdforeston
Pacific coa;dl range' altitude 3,500 feet, January 1939
(Rounds).
(T.) atriPes
Toxorhina (Toxorhinal dgrtvena is allied to T.
espe'
AlOxandernof C.oiombia,differing in the coloration of the body'
shortcially the thorax, and in the venation, especially the very
Sc and Rr.