i_ 8[Reprinted from Psvcnn, Yol. XIX, No. 6,]

NEW NEARCTIC TIPULIDAE @IPTERA).\Bv Cnanr,ns P. Ar,nxerrrpnn,
Ithaca, N. Y.

i
I

The following crane-flies, received from various correspondents
during the past few months, are believed to be new to science.
Limnobiaargenteceps
sp. nov.
Antenne l5-segmented; pleura yellow, with a brown band; wings spotted; Sc
short, ending before the origin oI Rs.
d, Length, 4.6 mm.; wing,5.9 mm.
Head: rostrum and palpi black; antenna lS-segmented, with the eegments
all black, flagellar segmentr rounded-ovate; front, vertex and occiput black witha conspicuous silvery-grey bloom; geneedarker,
pronotum light brownish-yellow; mesonotum, preescutum: in front
Ihorax:and on the lateral margin, rather bright yellow; an indistinct broad, brown, medianstripe which spreads out behind and covers the entire sclerite before the suture;scutum brown, paler, yellowish, medially; scutellum and post-notum brown.
Pleurre yellow, with a broad brown band extending from the cervical sclerites backto the halteres and base of the abdomen, Halteres, pale at base; remainder, darker,
brown. Legs: coxe bright yellow; trochanters yellow; femora yello'wish-brown,darker toward the tip; tibie and tarsi brown.
Wings: hyaline, veins brown; a brown spot at the tip of Sc, extending aroundthe base of the sector; a large square stigmal spot; all cross-veins and deflectionsof veins, narrowly marginal with brown; tip of the wing and ends of all of the veins,faintlytinged
with brown. Venation (See fig. 1): Sc short, Sc, at the tip, ending before the base of Rs; Rs rather square at its origin; basal deflection of Cur
before the fork of M.
Abdomen: tergum dark brown, the extreme apice of each sclerite and a broad
median patch on the base of segments I to 6, yellow; sternum mostly pale yellow,
the lateral margins and an indistinct, sub-apical, cross-band, brown. Hypopy'
gium yellowish: margin of the Qth tergite almost straight; pleural pieces, cylindrical, thickly beset with stout hairs, bearing apically, a large pale segment which
is likewise beset with hairs, and armed at its tip with a slender, chitinized projection, curved at its ends; guard of the penis long, prominent, enlarged apically andnotched.
r Contribution

from the Entomological

Laboratory,

Cornell University.

Psgche164

fDecember

Mts., Arizona; Aug. 1905(H. Skinner,Coll.)Type in
Holotype,d, Huachuca
coll.Acad.Nat. Sci.PhiI.
The reference of this insect to Limnobia is merely provisional;
I do not care to add a new genus to our already too long list, nor
do I know of any genus into which this species will accurately fit.
It is remarkable in possessing antenne which are l5-segmented.
The hypopygium is more like that of Limnobia than it is like thatof. Furcomyio, still the appearance of the insect, and its venation,
are strongly suggestive of the last-named genus. It may be wellto compare it with this genus; in Doane's Key (Ent. News, Jan.
'08; p. 5-?) it runs down to couplet 30, but runs out because of its
spotted wings. It is related to Furcarnyia si,gnipennz.rCoq. of
Calif or nia (J .N .Y . E n t. So c ' ; v o l . 1 3 ; p.56; (1905).
ElliPteraastigrtaticasP.nov.
Wingswithouta stigmalspot.
d Length,8.75 mm.; wing, 14 mm'
Head: rostrum and palpi brownish-blaek; antenne, seguent one, elongate-cytindrical; flagellar segments oval, similar to one another in shape, gradually smaller;antenne black; front, vertex and occiput black; the occiput narrowed behind'(It is probable that the head is covered with a grey bloom in fresh specimens; thetype is injured.)Thorax: very convex, prascutum large, greyish-brown, clearer grey along themargins and on the pleure; the prescutum is very large, so that the meso-thoracic
Iegs are very widely separated from the prothoracic pair, but close to the metathoracic; scutum, scutellum and post-notum dark grey, the caudal margin of the scutelIum brown. Halteres light yellow, the knob brown. Legs: fore coxeedark brown,
yellowish at the tip; remainder lacking; middle and hind legs, coxe yellowish'
brown; femora and tibia obscure yellowish-brown; tarsi lacking'Wings: hyaline, with a faint yellow sufiusion in cells C, Sc, Rr and anterior portions of Ra; stigma entirely absent; veins, brown, Rt darker than the others; abrownish sufiusion below Cu and 9nd Anal. Venation (See fig. 9'): Costa invicinity of sc bellied out cephalad and incrarsated; sc rather long, lying closer toC than to R; Scz far retracted, so that Scr is about equal to Rs; R long, ending
before the wing-tip, strongly incrassated to near the origin of Rrst; Rs long aris'
ing at an extremely acute angle, diverging only slightly fromRr; R,1t in a directline with Rs; basal deflection of Rrar short, strongly arcuated, beyond cross-vein
r-m, straight, parallel with Rrar to near the tip when it runs somewhat caudad; M
weak, on a line with Mr and cur; deflection of Mrar almost as long as the cross-vein
r-m1 Mra, proximad of zrzlonger than that portion beyond it, making cell lst Mnvery elongate; Cu strong, Cur about two-thirds as long as Cu,; Cut fuses with M
just before the fork; lst A very weak; gnd A stronger, gently bisinuate; anal angle
rather proninent.

1e121

Alexander-N

eu N earctic Tipulidae (Diptera)

165

Abdomen greyish on dorsum, pleure darker, blackish; sternum testaceous.
Holotype, 6y', Roger's Pass, British Columbia; July 30, 1908. (J. Chester BradIey, coll.)
Type in Cornell Univ. coll.

This species agrees with the other American species, clausa O. S.
of California, in its possessi'nga median cross-yein. It difiers in
the entire absence of a stigmal spot. I have seen the types ofclausa in Cambridge, thanks to the kindness of Mr. Henshaw, andfind that that portion of Rr, just underneath the stigma is bent
caudad, and, beyond the stigma, regains its former level; no suchcondition exists in astigmatica,
Erioptera(Mesoqphona)distinctasp. nov.
Vertex variegated; mesonotum clear light grey; dorsal stripes rather broad,clear; pleural stripes very distinct, clear-cut; Iemora with two dark bands, excepting the middle pair which has one; wings greyish, spotted and dotted, on the veinsand in the cells, with wbite.
d, Length, 3.6-3.8 mm.; wing, 4.6 mm.
Head: palpi brown; antenne light yellowish-brown; rcrtex with a large, clear
grey, oval spot, surrounded by a brown border; the outer margins of the vertex,
nearest the eyes, fawn-colored; occiput dark brown. Cervical sclerites pale yellow,
dorsally with two dark spots.
Thorax: pronotum whitish, the scutellum interrupted medially with a dark pit;mesonotum, prascutum clear light grey, the lateral margins narrowly greyish; abrown stripe on either side, beginning near the caudal end of the pronotum, conPseudotinuing backward; a brown stripe on either side of the grey median vitta.
sutural pitst elongate-oval, pale; the space anterior to this fovea is pale, fawncolored; prescutal, or tuberculate, pits, distinct, black, separated from one anotherby a distance equal to one and one-half the diameter of one; these pits are locatedin the median ground stripe near the proximal edge of the dark ilorsal stripes;
scutum, greyish, the vitte of the prescutum continued backward onto this; scutellum light yellowish-brown, darker anteriorly; post-notum grey, tri-vittate withdark brown. Pleura dark brown; a narrow yellowish-white pale stripe runningfrom the lateral margin of the cervical sclerites, caudad; broadening out above thefore coxa and continuing to the wing basis; a broad,clear,'silvery-white
stripe,narrowest anteriorly, beginning back of the fore coxa, running back to the base of
the abdomen; sternum clear grey. The pleura has the appearance of having threedark and two pale bands. Ilalteres, stem white, knob pale brown. Legs: fore andhind femur, yellow with a broad sub-basal, and a narrower, sub-apical, bron'nishblack ring; tibire and tarsi whitish; extreme tarsal segments darker; middle leg,similar, but femur has only the sub-apical dark band.
Wings greyish, browner on the cephalic half, with numerous white spots anddots, these largest along the costa and on the cord; the cells are speckled withnumerous fine dots.

166

Psyche[December

Abdomen light brownish-yellow; a broad, brown, median stripe'
Holotype, d, Highrolls, New Mexico; May 31, 1902' Paratypes,
Highrolls, New Mexico; May 31, June 2, and June 10' 1902'
Types in coII. Acad. Nat. Sci. Phil.

3

d's;

This species differs from its nearest ally, caloptero Say, in itsvery clearcut pattern of coloration' clear grey with distinct
pleural stripes, not yellowish with indistinct pleural stripes, etc'
Trimicra pygmaasp.nov.
greyishSmall; brown; basal half of antenne yellow, remainder brown; wings
with s short pubescence in all the cells.
9, length, 3.2 nm.: wing, 3.5 mm.
f,Iead: rostrum and palpi dark brown; antennre, basal seven segments light yellow, terminal segments di,rk brown; front, vertex and occiput dark brown'
yellowThorax: dark brown, the lateral margin of the mesonotal prescutum dull
notevident;
stripesdarkbrown,
dark
post-notum
ish; scutum, scutellum andHalteres yellow'
pleurre dark brown, more yellowish near the dorsal margin'
brown.
tarsi
tibia
and
femora,and trochanters yellowish-brown;
L"g.,
"o**
grey; veins brown'
Wings: hyaline with a slight greyish tinge; stigma indistinctly
pubescence.
R *o"" yellowish. lYing covered in all the cells with a thick short
Rr at anvenation: (see fig. 3) sc, retracted far back from the tip of scr; Rs leaves
deflection
acute angle; fork of Rra, deep, Rra, being a little shorter than the basal
ofof Cu,; c-ross-vein r just beyond the fork of R'a' and far removed from the tip
short,
R.. Abdonen : dark brown; in the I with the valves of the ovipositor rather
yellow.
H o l o t y p e :d ,Wo o d wo r th ' sL a ke ,F u lto n Co ' ,N' Y '(al t'1660ft);Lttg'22'79Lo(Ale"aode", coll.). Allotype; 9, with the type' Paratypes: 9 ' Wooster' Ohio;
Houser'd Coy
iept. 20, 1911, on grass-lands, (Houser, coll') In coll' Johu
Shefreld')(Alexander
andL9l.2;
May
28'
N'
Y.;
Glen, Ithaca,

The types are mounted in balsam, in the author,s collection.
Related io T. anomalo O. S. but much smaller and quite difierently colored; Mexican specimens which I have determined as
T. onomalamay, or may not, be conspecific with T' pilipesFabr'
of Europe (Compare Osten Sacken, Western Dipt', p' 200') If
so, it i; easily separated frop pygmaeo' by its non-pubescentwinss.
rl h a ve a p pliedtheter m,Tlseuilowtur alpils' tothedeep i m pr es s i ons outheanter el ater al
Tipulide (humuat prlla o{ Osten Sacken)'i"margins of the prascutum""i"ii"g-u"vof above as the'tubertuIete pits ''double clote'of Osten Sacken are spoken

Therel2lAlexander-New

Nearctic Tipulidae (Diptera)

Limnophila

167

laricicola sp. nov.

Small; antenne of the d elongate; color of the body light yellow; wings subhyaline.
d, Length, uncertain, abdomen broken; wing, 5.4 mm.
Head: antenne elongate, the segments indistinct, twisted, elongate-ovate, nar-'rowed at the ends, brownish in color; front, vertex and occiput light brown, butdisbolored, and possibly difierent in fresh material.
Thorax: pronotum browrr; mesonotum, prascutum brown on the anterior margin; a dark brown spot on the frontal third of the sclerite in a median position;
remainder of the sclerite light yellow, sub-shining, with a sparse pale bloom; scutum and post-notum light yellow, the latter with an indistinct narrow brown medianstripe; pleure yellow, darker near the dorsal margin. Halteres, stem pale, knob
darker. Legs: coxe and trochanters light yellow; remainder of the legs brown,rather darker outwards.
Wings: subhyaline, stigma indistinct, brown. Venation (See fig. 4): Sc rather
long, extending to beyond the fork of Rs; cross-vein r at the tip of Rr; Rs short,less than Mr or Mz in lcngth; Rr4a rather long, as long as Rs, gently arcuated; RrdeflectionofR+raboutaslongascrossratheroblique,cross-veinrnearitsmiddle;
vein r-m; petiole of cell Mr moderately long, about three-fifths of cell Mr; basal
deflection of Cur beyond the middle of cell lst Mr. CeII R, very acute at its proximal angle; cell lst Mz long and narrow.
Abdomen broken.
Holotype, d.
Canada Lake, Fulton Co., N. Y.; June20, 1911 (Alexander).
Occurred in an extensive sphagnum bog, on vegetation growing underneath larchtrees. (altitude 1550 feet)Type in author's collection.

This speciesby its elongate antennre in thed, is aliied to poetica
O. S. I thought that it was a small d of this species until I saw
the types in Cambridge. Poetica is conspicuously larger, withquite different antennal structure. In laricicola, the antennal
segments are twisted, and the sutures between them indistinct,
producing an appearance quite difierent from that which obtains
in the male sex of.poetica, tenuipes,niueitarsis,etc. Venationally,
this new species differs from poetica in its longer Rz+s, shorter
type, and only specipetiole to cell Mr, etc. Unfortunatelymen, was recently accidentally damaged.-yErioceraafbihirtasp. nov.
Antennre short in the d; cell Mr presenU body densely clothed.ivith long whitehairs.
d Length, 15 mm.; wing, 15.6 mm.; antenna, 4.5 mm.
Fore leg, fem. 8 mm.; tibia, l0 mm.
Middle leg, fem. 9.9 mm.; tibia,8.4 mm.
Hind leg, lem.12 mm.; tibia 12.6mm.

I

168

PsgcheIDecember

Head: rostum and palpi dark brown; antennre, first segment large, cylindrical,
brown; remainder of antenne black with black hairs. Front wide, the frontal
tubercle not prominent; {ron! vertex and occiput, medium brown densely clothed
w i t h l o n g p a l eh a ir s; a sp o to n th e fr o n tju stb e h in d thebaseoftheantenne,pal er,
golden-yellow.
Thorax : mesonotal prrescutum dark dull black without apparent stripes (thoughthese may have been destroyed by some means or another), the entire sclerite
densely clothed with elongated white hairs, very conspicuous; those on the dorsum
darker and shorter, those on the lateral margin of tbe sclerite exceedingly conspicuous; scutum dull black; scutellum dark brown, also with numerous pale hairs;
post-notum black. Pleurre light grey with dense white hairs; the light color of
the pleure renders it probable that the dorsum is grey with darker stripes, in living
Halteres, stem light brown, knob darker brown. Legs: coxa brown,material.
with a grey bloom; trochanter brown; fore femora, basal haU brownish-yellow;
apical half dark brown, uniform throughout; tibia and tarsi dark brown; middle
leg, femora with rather more than the basal half light-colored; hind leg, femorawith only the extreme base yellowish.
Ilings: subhyaline or light brown, more brownish near the veins; cells C and
Sc brownish-yellow; stigma small, oval; veins brown. Venation: (See fig. 6);
Sc extending beyond the fork of Rrar; R long, cross-vein r about one-half as longas that portion of Rr beyond it; r before the middle of Rr; cell Ml present. Venation almost exactly Iike the eastern E. brachycera O. S.
Abdomen: tergum uniform dark brown, the lateral margin pale yellowish, andprovided with long pale hairs, these longest on the basal segments; hypopygium
yellowish-orange; sternum more greyisb.
Holotype: Thelabels read'L2 California. No.846. Coll. Hy. Ddwards.' Probably from Marin Co.

Related to E. brachycera O. S. but darker and clothed with long
pale hairs much longer than in brachgcera. The basal segmentsof the antenn@ in brachycerd are very light yellowish-red, notbrown; the front of. branhyeero in the vicinity of the tubercle isuniformly pale; the thoracic dorsum may or may not be similarly colored in the two species (see statement, above, concerning
the condition of the thorax in albihirta). In brachyceraLhelegs
are much lighter-colored with narrow blackish tips to all the femora;
in albihirta the femoral tips are uniform brown and very broad.
Eriocerafultonensissp. nov.
Antenne short in both sexes; cell Mr absent; body coloration dark brown;prescutum four-striped; wings light brown; cross-vein r far before the fork of
Rr+r,i, Length, 9.6-10.2 mm.; wing, 9-10.4 mm.
9, Length, ll.2-11.5 mm.; wing, 10.5-11.3 mm.

rerelAletonde r-N ew N eareti.c T i pulidae (D i pte ra)

169

Head: palpi dark, blackish, rostrum much paler brown; antennre short in both
sexes; scapal segments dark above, reddish-yellow underneath; flagellar segments
black. Front, vertex and occiput black with a grey pruinosity, most distinct behindthe eyes.
Thorax: mesonotal prrescutum light yellorvish-brown with a dark brown stripe
on either side of the paler median vitta, these stripes broadest in front, narrower
behind, ending just before the suture; a shorter, curved, Iateral stripe on the lateraledge of the sclerite, beginning just behind the pseudo-suture, continuing to thetransverse suture; scutum, scutellum and post-notum blackish, scantily grey
pruinose; pleurre unicolorous greyish-black. Halteres, base of stem, pale, yellow;
remainder of stem, and the knob, dark brown.
Legs: coxe light brown; trochanters and basal three-fourths of the femora,
light brownish-yellow; apical fourth ol the femora, brown; in the fore-legs, theapical half of the femur is brown; tibia light brown, the tip black; tarsi brown.
lYings: veins dark bron'n; membrane rather uniformly tinged w-lth light brown;
stigma small, delimited externall;' by the radial cross-vein; extreme tip of thewing rather darker. Venation (See fig. 7): cross-veinrnear the tip of Rr, situatedfarbeforetheforkof
Rrar; RrmuchshorterthanRrar;
cellMrabsent; basaldeflection of Cur slightly before the fork of M.
Abdomen shiny black, including the large hypopygium; in the 9 , the valves ofthe ovipositor are likewise black.
Holotype, d, Sport Is; SacandagaR; Fulton Co.; N. Y. June 28, 1911. (Alexander, coll.) Allotype, 9 , with the type. Paratypes, 5 d, 5 9, with the type.
Types in author's collection.

Nearest related Lo E. fuliginosa O. S. (East. U.S.), from which
it is readily distinguished by the lack of a grey bloom on the thorax,
the shiny jet black abdomen and genitalia, etc. In normal individuals of fultonensis, cross-vein r is far before the fork of Rz+a (Seefig. 7). In rare instances, however, the cross-vein is much nearer
to the fork though never quite at it. Specimens which I refer
Lo fuli,ginosa, provisionally, show a venation similar to that described for the species (See fig. 8); the coloration is very dark,
however, and compa.rison with the type may prove them to be
novelties. E. fultonensfs differs from .O. gibbosa Doanel (Mich.)in the same venational peculiarity.
Eriocera cinerea sp. nov.
CelI Mr absent; color of the thorax light grey; stripes on the mesothoracic
prrescutum indistinct; cross-vein r far beyond the fork of Rzar.
9, Length, 11.4 mm.; wing, 11.2 mm.
9, Headl rostrum yellow; palpi yellowish-brown;r Doane, Joun.

antenne, segments one

N. Y. Ent. Soc.; vol. 8; p. 193; pt. 8, fig, f0. (f900)

170

PsgcheIDecemberand two, light orange-yellow beneath, light brown above; flagellar
segments brown.
Front and clypeus light orange-yellow; vertex and occiput light grey-,
more brownish medially, paler on the vertex behind the eyes, and on the genre.
Thoraxl prothorax, scutum dark brown with a grey bloom
and prominent palehairs; seutellum dark brown medially, the ends conspicuously
orange_y"llow.
Mesothorax yellowish-grey, the prrescutum with four very
indistinct darker rines;
the middle pair long, confluent or nearly so at their cephalic
ends; lateral pairshort, beginning behind the proninent rounded, black pseudo-suturarfoveu;,"rri.rr*
and scutellum brown with a clear light grey bloom; post_notum
dark brown.
Pleura light grey; halteres, stem brownish-yellow, knob brown.
Legs: coxre and.
trochanters rich orange-yellow; femora light brownish-yeilow,
the extreme tip
darker, brown; tibire, extreme base brown, remainder yellowish-brown,
insensibiydarker apically; tarsi brov,n.
Wings sub-hyaline or very pale brown; costal and subcostal cells light yellow;
stigma ill-defined, yellowish-brown; veins brown. venation (Seefig. of sc short,ending before the fork of Rzaa; Rrlr slightly longer than that portionof Rz before
cross-vein r; cross-vein r inserted near the tip of Rr and far
beyond the fork ofRz..r; cell Mr absent; cell lst Mr elonsate.
Abdomen: tergum dark brown, the-rateral margins of the screrite
broadly, thecaudal margin narrowly, brownish-yellow;
vah'es of the ovipositor broilishorange; sternum greyish-brown.
Holotype.
Q, Boston, Massachusetts.
T y p e i n a u t ho r ' s co lle clio n .

Related to fuliginosa o. s. but head and thorax conspicuousry
grey, not brown; abdomen not uniform brown; cross_vein r
sit_uated far beyond the fork of R2_q, etc. The female of.E. wilsoni,
O. S. (East U. S.), of which f have seen the types in Cambridge,
is unknown. This new species cannot be that sex of wilsini
because of its totally different body-color. E. rongicornziswalker
is quite a different species; its venation seems never to have beerr
figured (The figure in Needham,s p3rd Rep,t of the N. y. State
Entomologist, is not longicornis, buL spinosa.) so I include adrawing. (See fig. 10.)
Rhaphidolabisneomexicana
sp.nov.
cross-vein m of the wings absent; wings pearly-white with a distinct
dark brownstigmal spot.
9, Length, 5.4 mm. wing, ?.6 mm.
Ilead: rostrum and palpi dark brownish-black, antenne black;head dark_brown, sparsely grey pruinose. Thorax: mesonotal prascutum dark brown,
very
sparsely grey pruinose without distinct darker stripes, though rather
d""p".colored medially; scutum, scutellum and post-notum dark brown with afaint grey

Pe:cm, lgt2,

Vor. XIX, Prerr 13

I

,i
F

I
I

I

Alwand,u-N

I
I

;
I

aw N earctit TipulidererzlAleaander*New Nearctin Tipulid"aeDiptera

171

dust. Pleure dark brownish-black, grey-dusted. Halteres, stem pale, knob
brown. Legs: coxe brownish-yellow, more greyish anteriorly; trochanters brownish-yellow; femora and tibire uniform medium brown; tarsi gone.
Wings: hyaline or sub-pearly; a prominent elongate dark brown stigmal spot;
veins brown, the longitudinal veins rather paler at the wing-root.
Venation:(See fig. 5): Sc long extending beyond the fork of Rz4r; Rt long, cross-veia r very
near its tip; Rs short, oblique; Rzlr ver;z short, only a little longer tian cross-vei:osr or r-mi cross-vein r beyond the middle of Rz; Mr about as long as Rs; no crossvein mi basal deflection of Cur beyond the fork of M.
Abdomen: dark brown, the pleural sutures lighter brown; valves of the ovipositor light colored, brownish-yellow.
Holotype, 9 , Beulah, New Mexico: June 99, 1909.
Type coll. Acad. Nat. Sci. Phil.

Most closely allied to R. tenuipes O. S. of the Eastern UnitedStates, but differs in its wing coloration, apparent lack of thoracic
stripes, dark stigmal spot and venational details, i, a,, more obliqueRs, shorter and more divergent Mr and Mz, etc.
R. d,ebilis TVill.l appea$ to be a Tricyphona by its venation(Rz+s fused and Ra and Rs separate). It agrees most closely with
T. oitripennis Doane.
The membem of this Lribe, Ped,icinri, require careful study at thehands of some competent student who has accessto an abundanceof Western material.
The West appears to be the center of distribution for the members of this group, which, in other parts ofthe continent, are represented only by a few species, or else, as in
the tropics, quite lacking. I surmise that such a study, based on
sufficient material, would show eongenerousness of Plectromgi,oO. S. and Rhaphidnlabfs O. S., with DicranotaZeII.
Eaplanation oJ tha Plate 73.
1. Wing ol Limnobia argenteceps, sp. \.
9.
" " Ell'iptera astigmatica, sp, n.
3.
" " Trimicra pggmaea, sp. n.
4.
" " Limnophila larieicola, sp. n.
o.
" " Rhaphid,olabis neomesica?ro,sp. n,
6.
" " Eri,ocera albihirta, sp. n.
3 . ..E .
atultonensis, sp, n,g," "E.
Juliginosa O. S, (or related species)g.
" "E.
cinerea, sp. n.
10 .
Iongiccrnis Walker.
" "E.
t Williston, Kans. Univ. Qurt.;

vol. 2i p. 621,Oot. 1893.