+{trNEW OR LITTLEKNOWN SPECIS OF WEST
INDIAN ,,TIPULIDAE'(DIPTERA)L. L
by
By CHARLES P. ALEXANDER

Reprint fromIIIEJOURTIT
OFA8RICUTIURE
l|FIIIEUTIIER$ITY
OTPUERII|RICO
Vol. XX,

No. 4, Oct. 1986,p. 577-8Sg

NEW OR LITTLE.KNOWN SPECIESOF WEST INDIAN..TIPULIDAE'' (DIPTTRA) L. I.
By CHnnlps P. Ar,nxnNDER
Massachusetts State College, Amherst, Massachusetts

A few interesting new speciesof crane-flies have been includedin various Jotsof specimenssent to me for identification.
The following sourcesare included: Puerto Rico, through Dr.
William A. Hoffman; Cuba, through Professor Stephen C.
Bruner and 1\'Ir.A. R. Otero; and the Bahamas,through Dr.
John G. Myers. Through the friendly interest of the abovementioned en+"omologists,
I have been privilegecl to retain thet1'pesof the speciesin my private collectionof these flies.
Shannonontyiahoffmanf sp. n.
General coloration gr&y, the praescutumand scutum dark
brorvn; pleura brorvnishblack, sparselypruinose; wings creamyellou', sparsely variegated by tlark brown areas; cell ^R3small,vein R2+3 belng iongerthan R2+3+4; cell 1sf Mz elongate,closed;abdomenblack.
MaIe.- Length about 5.3 - 5.5 mm. ; wing 5 - 5.2 mm.

Rostrum anc. palpi dark J:rown. Antennae short; seape,pedicel and basal one or two flagellar segments light yellow,
the remaining segmentsbrownish black; verticils exceedingthe
segmentsin lengtir. Head gr&y, the front and anterior vertex
clearer and lighter gray.
Pronotum and lateral pretergites clear gr&y, the latter extending to the wing-root. Mesonotalpraescutumand scutum almost uniformly clark brown, the median area of the former
darker, the Iaterai portions more pruinose; scuteilumuniformly
blackened,sparselypruinose; mediotergitegray pruinose. pleura brownish blerck,sparsely pruinose. Haltercs chiefly pale.
Legs with th e coxee brownish testaceous; trociianters obscurey'-ellow;femora obscurebrownish yeilow, the tip.s rveakly darkened; tibiae brown, the tips narrowly more darkenecl; tarsi
blacl<. Wings ( Fig. 1) with the ground-color cream-yellow,J l t

878

THE JOURNAL OF AGRICULTURE OF THE UNIVERSITY OF P. R,.

sparsely variegated by dark brown areas, including spots atarculus; a virtually commonarea at origin of Rs,ScZ and tip of
Sc1; series of spots along cord and outer end of cell lst M2;
small marginal darkenings at tips of all longitudinal veins excepting R5, the largest ai Znd A; veins yellow, darker in theinfuscatedareas. No macrotrichia on Rs or its anterior branch;
R5 with abundant trichia over its entire length. Venation: Sc
short, Scl enclingabout opposit€on€-third to one-fourth lengthof Rs, ScZabout mid-clistancebetween/?s and the tip of Scl ;R2 about one-half shorter than R2+3+4; R2-13 longer tlran
R2+3+4; cell Lst M2 elongate,closed;m-ctt closeto fork o! M.
Abdomenblack, the hypopygium a trifle paler.
Habitat.- Puerto Rico.
Holotype, male, Luquillo National Forest, at light, September 8, 1935 (W. A. Hoffman). Paratopotype,male.
Associatedat light with Shannonomy'ialeonarclfAlex. andErioptera (Mesocyph'ona)portoricensis Alex.
I take great pleasurein naming this distinct crane-fly inhonor of my long.time friend, Dr. william A. Hoffman. The
only closeally in the West Indian fauna is Shunnonomyi,aleonard,i,likewise from the Luquillo National Forest. The latter isreadily told by the uniform pale yellow coloration of the thorax,
together with the brownish yeliow abdomen. By my key to thePuerto Rican speeiesof the genus (Journ. Dept. Agr. Puerto
Rico, 26 : 368; 1932), the speciesruns directly to l'eonardi'
T euch,olabis(T eucltolabis) oterotlsP. n.
General coloration yellow, variegated with black; lateralpraescutalstripes bent laterad to ma,rgin; knobs of halteresiniuscated; femora yellow, eachwith a single,narrow, pale brown
ring immediately before midlength; tibiae uniformly yellow;
wings whitish subhyaline,with three pale brown fasciae,the last
being the naryow 'wing-tip; cell lst M2 closed;abdomenyellorv,
the basal/teryite black, the outer segmentsrestrictedly variegated by brown/areas;the basalareas; uralehypopygiumu'ith themargin of the basistyleproducedinto a thin, siightly blael<enedplate or flange; basistylenot producedat apex.
MaIe.- Length about 8.5 - 9 mm. ; wing 7 - 7.5 mm.

IiEW

OR LITTLE.KNO\\'TIPULIDAE

BYC. P. AIJEXANDER

8?9Rostrum, brown, obscureyellow above,slightly shorter than
the remainder of head; palpi black. Antennal flagellum black, thescapeyellow ; peclicelbrou'nish yellow; flagcllar segmentsoval,
with conspicuousverticils. Front and anterior vertex blackened,the posterior vertex and occiput yellow, the latter with aninconspicuousdarkenedtriangte.
Pronotum obscure yellow, darker anteriorly above. Mesonotal praescutum yellow, with three black stripes, the median
one paling to brown on anterior end, its posterior portion ending
abruptly somedistancebefore the suture, leaving a conspicuous
yellow area behind it ; lateral stripes -with anterior ends continued outward to margin of sclerite, irolating a yellow area between them and the suture; median area of scutum pale yellow,
the lobesextensir,'siydarkened;scutellumweakly darkened;mediotergite obscure brownish yellow. Pleura chiefly darkened,heavily light gray pruinose. Halteres with the stem yellow, theknob infuscated. Legs with the coxae and trochanters obscureyellow; femora y,:llow, unmarked except for a single, narrow,pale brown ring just before midlength of the segment;tibiae andbasitarsi yellow, the outer tarsal segmentsblack. wings (Fig. 1.
2) whitish subhyaline,with three pale brown fasciae, the firstlying before level of origin of Rs, extending from veil ,B to theposterior margin ; secondband chiefly beyoncllevel of cord, €Xtending from the darker brown stigma to the posterior margin,
including all of cell lst M2; third band including the narrow
wing-tip; veins yellow, the costarfield more brighgy so; veins
darkened where tt'aversing brown areas. Venation: Sc1 ending
oppositemidlensh of .Rs,the latter moderatelyarcuatecl; RZ+3+4 a little shorter than the slighilS'oblique R2; branchesof^Rslying approxintatelyparalle! to one another to margin, cellR2 being neariy three times as extensiveat bcrder as is cell R4;
cell \st MZ closecr; m-cu shortly beyond fork of M.
First abdominaltergite black; outer segmentsyellow, veryvaguely and restrictedly marked with clarker,msre extensivelyso on the sternites; hypopygium pale. lvlale hypopygium withthe mesal edgeof basistylethin and weakly blackened,spine ofbasistyle long, siencler,gently curved, the basal half pale anddilated; basistylenot or scarcelyproducedcaudadat outer end.
Outer dististyle a gently curveclrod that terminates in a simple
spinouspoint.

OT' P' R'
THB JOURNAI/ ON AGIiICULTURE OF THE UNIVBRSITY

Habitat.- Cuba (Pinar del Rio) 'Holotype,male, Las Animas, sierra Rangel, altitude 1'500
feet, september4, 1gB4(s. c.Bruner and A. R. otero). Paratopotype, male.
I take great pleasurein dedicatingthis interesting species
to Mr. A. R. Otero, to whom I am indebtedfor severalinteresting speciesof Cuban Tipulidae. The speciesis very different
from the remaining regional speciesof the genus in the diagnostic features listed. The nearest ally is the Jamaican Teuclt'otabi,s (T eucholabis) gowd,eyiAlexander, which differs conspicuously in the opell cell MZ of the wings, the leg-pattern, andthe structure of tl^e male hypopygium.
Teucholabis (Teucholabis) portoricoina,sp. 11.

Closely allieC to T. (T.) myersi Alexander, of Cuba andSouthernFlorida (Journ, N. Y. Ent. Soc.,34 : 228 - 229; L926),
differing chiefly in the structure of the male hypopygium.
lV[61s.- Length about 5 mm. ; wing 5.5 mm.
Fem,ule.- Length about 5 mm. ; wing 5 mm.
Male hypopygium with the apical spine of basistyle astraight pale rod, i-erminatingin a sl,ort acute slrinethat lies inthe sameaxis as the remainder of the rod. Outer dististyle palethroughout,slender,the spine on mesal margin small, suberect.
Inner dististyle with the outer emarginationshallow and much
shorter than the basal olle, the denticle separating them lorv
and obtuse. Aedeagus with the apical point straight, palethroughout.
The male hypopygium of myersi has the lpical spine of
the basal half dilated, thethe basistylelarge ancl conspicuoLis,
distal half abruptly narrowed into a long sinuousspine. Outerdististyle darkeneclthroughout,the spineoI1mesalmargin acute,subappressed.Outer dististyle elongate, the two emarginatious
subequalin size or the outer a trifle more extensive,the lnedian denticle acute. Aedeaguswitir the apical point cttrt'ed,blackened.
Habitat.- Puerto Rico.
Holotype,male, El Semille,Villalba, altitude 1,600feet, atNllw oR LrrrLE-KNo\\- TrpuLrDAE By c. p. ATJExANDEB

881

lighat, January zG, 19Bb (w. A. Hoffman). Allotype,female,Paratopotype, sex?
)
The wing-venartionis shown in Figure B.
Tororhina (Tororhina) dtstalis sp. n.
General coloration grdy, the praescutumwith three brown
stripes; femora brownish yelrow, the tips weakry
darkened;tibiae brownish yellow, the tips narrowly brownish
black;
wings with macrotrichia of veins very sparse; Rs unusu.lly
Iong, exceedingin length vein R5, due to the positionof r,-r/L
closeto outer end of the long cell lst M2; nl-cu about
two- thirds
its length beyond the fork of M.
mm.

Female.- Length, excluding rostrum, g mm.; wingb.b

Rostrum broken beyond the base, which is dark brown.
Antennae with basal segments dark brlorvn; outer flagellar
segments broken. Head gtay.
cervical region dark brorvn. I\[esonotumlight grey,
thepraescutum with three brown stripes, the laterals
somewhatless distinct; posterior interspacesweakly infuscated; centersof scutal lobesdalkened; scutellumancl merliotergiteclear lightgra)" Pleura and extreme lateral margin of praescutum
lighttheventral sternopleurite somewhat clarkened. Halteres
9r&)',
pale' Legs with the coxae weakly infuscated; trochanters
yellowish testaceous;femora brownish yellow. the tips
rveaklydarkened; tibiae obscurebrownish black; tarsi brolvn.
Wings(Fig' ) rvith a grayish tinge, the pearcularregion
light yellow;
veins brown. I\fa.crotrichiaof veins very sparse, intype witha single trichium near outer fourth of ^Rs; iou" or fivescattered along R5,' &h equarnumber near outer end of cell rst
MZ
venation: Rs unusualrylong, exceeclingr?b, due to the positi,nof r-nt far distad, closeto outei' end of cell Ist MZ;
cell Ist M2
present, elongate,exceedingcell ZnclM2; r-rn
lying immediate_
Iy before nL; ?n-cu(.
about two-thirds its leneth teyonct the forkof M.
Abdominal tergites brown, the caudal ancl lateral borders.
together with a more or less distinct crossbandat
midlengthof the segmentsrestrictecllydarker.

SS2TEsJoURNAIJox.AGBIcUI/aUBEox|TEEUNIVDBSITYox'P.R.

Habitat.-Bahamas.

female,New Providence,May Lg32(J. G: Myers) ;Holotyrpe,
Colector'sNo. 2557.
Tororhi,na (Tonorhina) d,i,stalisis very different from allother regional species in the distal pcsition of crossveinsr-rnand In-c'tt,together with the unusually long cell lst M2'2Explanation of figures.
l-Shannonnomyia hoffmnni, sp' n'; venation'
Z-Teucholabi,s(Teucholobis)oteroi sp. n.; venation'
sp' n' ; venatiotr"
3-Teucholabis (Teucltotobi,s)portori,ca'n&A-Tororhina (Tonorhina) di,stalissp. n.; venation.