CRANE FLIES FROM THE
MARITIME PROVINCEOF SIBERIA
loN THB n{SSCT FAITNAOF TSB MARrrIrlr
PROVTNCB
OF SIBERIAI

8.

l;fffl.'.:"[B:'ffI

3Y

Honolulu; Ffawaii g1gl?

CHARLES P. ALEXAI{DER
of thc llurchurttr

No 2605.-FrouAgrieltunl.P. Bf$HOp A,ttSEUI/

Collceer An[srltfc frocceain$ of thc Unitcd Strtcl Nrtioad Mrrc'VoL 6g Art 4r pp l-21, pL I'lWASEINCTONCOVBRNMENT PRINTING OIIICE
l92S

CRANE T'LIES X'ROM THE MARITIMESIBERIAPROVINCE OT'

By Cnenr.rs P. Ar,nxeNnnn
O! the Massochusetts Agricultural CoUege,Amherst

The irane flies that were taken by prof. T. D. A. and Mrs. cockerell on their recent trip to Easternsiberia have been submitted tome for determination. Because of the almost total lack of material
and records from this region, the collection proved. to be of excep:
tional interest and furnished several important adclitions to theincomplete list of the Tipuloidea of Eastern siberia. rn
addition to
the speciesrecorded in the following pa,ges,professor cockerell writes
me that ho saw, but was unablo to capture, a specimen of Ctenatroscel'ismi'kod,o(westwood) at Tsuruga, Japan. idy sincere thanks
are extended to Professor and Mrs. Cockerell, and. to the National
Museum authorities, for the privilege of studying this seriesof ripulidae.
typel and spocimettr of all othlr ircruded species i,reth9
proserved in the National Collection.
rn order that the.known and probable components of the Tipulid
fauna of Eastern siberia may be mo"e cod"ctry understood, theto{g,"i"g list of negimlal Bpeei,es
is suppried. rn addition, it may bewell to indicate the occurrenceof tho European Di"ctetuidi,a'bimacul,ata(Linnaeus) in this coastal province of siberia, on enormous extensionof the range of the species,which, however, has never been taken in
Japan.
REGIONAL SPECIES
Almost th_eonly siberian Tipulidao that may be considered as be.
are the- following: Ti,puta breui,ceps(Motschursky)litg
from "eglooal
Amurland and few Tipulin"
flies bel^ongingto the sub"portschinsky"rao" and 18gz
tribe ctonophoraria, described in lgz3f,y(Ctn oTilnra [,arua Portschinsky, C. pi,cti,Ttenruts
portschinsk y, Tany_ptera grad,Zrs (Portschinsky), T. minuta (portschinsky),"andT.
si'bertfra (Portschinsky), all from various parts of siberia, soms
from the vicinity of Vladivostock.
I Tlpnlitu brr?ic4rrMotschulsky, Bull. Soc.Imp.

Not. Moscou,yol. A2,p. bOA,1ES9.

NO. 26O6._PRooEEDINGSU. s. NATIoNAL MusEUM, VoL. 68, ART.
4.
63198_251_1

P R OCEEDINGS OF ' I' IIE

The Tipuiidae

NA' I' I0 N' \I'

MIIS I']IJM

v01.. orlx'f

soptrrntcd lrom tlrttof the island of Saglur'lion'of rar-siur*iu onry*bv [he nairow GuIf""m"iti""coast province o{ Eas"ttwith the ntoinland' The following
tarv, show notable
and Mrs' cockerellon
fi;Iia bv Proressor
fiJ,ir,#i"r;;i;from JapanesoSaghalien(Karafuto):
their trip a,re&lso k";;AIeximmoiJesfo'ides
pul'Li,monia karafuton'isJ'i"*n"a""' 2i!'y!n*uia
R'
Meigerynnlpa!;" i'iulata
ander, D. sytmicau," ft;ffi;'chraseptent*r,r,',-Al;;;;;"itkiopia"nemoralis(Meigen)'varietv'";;,i";;;"wi,*,ry1i*=J"i'rit,"1,it'::trtgl1#";iXif
i#i"lnparly,,"^i";i:::'btio Alexanaet. This constitutesof the species"ii^i^inl
be noted that allsndemicspeciesttk;;;Uut ii "fto"id
the Japanesethrough
southward
listed have a *id" ;"";;; "itnt" Region'or' in the casesof Bhryislands,throughoutini-p*""-"'ic thriushoui the Holarctic Region'"f"o* f utufuto occur in Eastern
ilin mnculon uoan'i{i'9*"*i"q
rtoo*"
The following Tip;;i;;Siberia or &reregional:

C heitotrichi'aiadda (Meigen)'Dis cobola mar garit a Alexander'
deuiata Alexander'
Ormo sia su'b
D. argzs (SaY)'
albib osis A'lexander'
olophilus
M
Alexander"-prloitt
il,rrino^gio'immoilestoides(Erioptero) iorii Alexander'"
D. megacauila Alexander'
nl. fn; fl,ooohumeralisAlexander'
D. mesosternafuAlexander'
E. @.) aanthoPtera Alexander'
D. sachaldnaAlexander'sachalino Alexander'
E. iliapno"")Alexander'
D. sqorso Alexander'
n. iu iitouals) asiat'ica
Alexander'
D. sPi,wicoud'aAlexander'
satsurnaO, opr"ider'Limonio ePi'semoAlexan
C teiophora big ultat a M atsumura'
Alexander'
utorui's;ij oJ,ana Mat's't:mura' variety'
L. kar aI""iio- " i"cludes nrocro Matsumura'L. ptutonis Alexander'
ipiouufrv
L, iubnubeculoso Alexander'
not Loew.)
Zl ono*a;'o aao cettaAlexander'
btbo Alexander'
TipulaR",hi'ni'ilio mo,culata M eigen'T . bi'Penicitleata Alexander'der'Alexans
oni
sep te nt'ri
iil i"iT. coouilletti Enderlein'"i, "uen'oscAlexander'o'li, inoPta
T . coquillettiano Alexander'"n"Alexander'
j aP onica
ii*ioenli"
T. flauocostalisAlexander'
varietY'T. nemaralit (Meigen),
T. i,nsulicola luscicaudo Alexander'
Alexander'
L. saPPtorensis
T. nipPonensis Alexander'
L. subodusto Alexander'
T . laiemar gin alo Alexander"
L. subqoeticoAlexander'
T. tantula Alexander'
L. utvicoides Alexander'
T. uerecundaAlexander'
P ilari,a d,orsalis Alexander'
T. aariicurnis Schummel't sachalinensis Alexander'iii"rrT. westwooiliano Alexander'
AlexanF' otionno""t gloriosus Alexander'i rpioi,o*o aculeala atricauila
a omalno g ibbera Al exander'ni
der.
' "iniu"d."l'^,i, i ri, ntaa"U sf' auiba si's*t:t
N. iorsalis sachof,ina Alexander'-.
Sacken'osten
uttimaii*i'i*"tii,ii
N, hi,rsulicaud'o Alexander'
varietY.
N. Iamellata (Riedel)' varietY'
Gntophorngia lristrls Alexander'
Alexander'sachalinensis N . minuticornds(Ptilostena)b""tr*uti
N. saghaliensisAlexander'
Alexander.
H elobia hYbrida (Meigen)'

I

j

I

(rl l ^N l ' :t,' t-l l ,l s l moM

S II]U MA _A LE X A N D E R

3'lh. P[yt:[.ptori<1, Ptgchoptera subscutellaris Alexander, likewiseo(:curs.
Matsumura, describedtwo new speciesand a supposedlynew genus
from sakhalin. IJ.is Limnobia saihalinenscs wo,rld upp"u* to bo a
speciesof Limnophila, or at least r{exatomine. The new-genus Metali'mnobi,a(for uittata,new) is certainly close to Li,mnni,ain ih" venationand nature of the claws. Noither of the above specieshavo beenrecognized since their original characterization.
Tho island of llokkaido (Yezo), the most northerly of the four
chief islands of Japan, is separated from sakhalin oniv bn the narrow La Perouse strait and from the southern end of the coast province ofby the relatively narrow northern portion of the-siberia
Japan soa. The Tipulid fauna shows a characteristic rrolarctic
facies. The following cra,neflies are now known from Hokkaido andseveralwill certainly be found to occur also in Easternsiberia.
Ascobola orgus (Say).
D. rnargarito Alexander.
D. moiwana Alexander.
Di,cranonr,yiaglobulithorat Alexander.
D. immo destnidesAlexander.
D. longipennis (Schummel).
D. mesostemala Alexander.
D, spinicauila Alexander.
D. subtristis Alexander.
Limoruia amaUilis Alexander.
L. ang ustistrio Alexander.
L. annulus truncota Alexander.
L. bosispdna Alexander.
L. bifasciata ar'.rsAlexauder.
L. oinitq, Alexander.
L. fusciceps Alexander.
L. inelegans Alexander.
L. machidai, (Alexander).
L, mendar Nexander.
L. monacanthc Alexander.
L. neoind,igenoAlexander.
L, neonebuhtsc Alexander.
L. nigronitiila Alexander.
L. quailrinototc (Meigen).
Libnotes longist;igrno Alexander.
L. nohirai Alexander.
Rhi,pidda n aculato Meigen.
E, pulchra septentrionis Alexander.
Elliptaro zipanguensis Alexander.
H elius tenudroslrrs Ale:iander.
D icr ono pty cha uezrosaAlexander.

Antocho (Proantocha) serricauda AJexander.
A. (Antocha) biTidoAlexander.
A. (4,) brerineraisAlexander.
A. (4.) brevistylaAlexander.
A. (4.) dilatata Alexander.
A. (4.) sdtsuma Alexander.
U Ia perelegans Alexander.
U. cincta Alexander.
E piphragrna subfaseipenni,s Alexander.
E. subinsignis Aiexander.
Limnophila inconcussa Alexander.
L. japonica Alexander.
L. kuwaEomai Aiexander.
L. melanommola Alexander.
L. neomunda AlexanderL. unicoides Alexander.
P ilaria dorsalis Alexander.
Eriocer a fulaibasis Alexander.
E. jozana Alexander.
Elephantomyia hokhaidenyi.s {.lexander.
P ed,icia daimio (Matsumura).
D icr anota yezoensisAlexander.
Rhaphid,olabis subconsorsAlexander.
Conosia irrorata (Wiedemann).
Chionea, sp.
P aratropesa esatfi Alexander.
Teu,cholabiegezoensisAlexander.
(Ptilostena)
Gonomgia
subpruinoso
Alexander.
G. (Gonomgia) superba Alexander.

tErstor Beitreg zur rnsekton-Faunayotr sachalin, Joum. coll.
agric. Tohoku rmpor. univ., vol. 4,
pp. l-45, 191r.

4

^rr,

MUSDUM
PRooEEDTNGSoF TrrE NATIONAI,of the related familY
Two species^Hokkaidoz Ptyclnpftera il'at'rnt'ofound intunar;a Alexander.
Family TIPULIDAE

lr.t,Itts IROM SIBE}I.IA_ALEXANDER

L829, DicranomyiaSrntnmrrs,
DICBANOMYIAStepheneCat. Brit, fns., vol.2,p,
IMMODESTOIDES

b

Z4B.

Aterander

19L9. Dicranorngia immod,estoid,esAr,nxaNonr, Ann. Ent. Soc. Amer., vol.
12, pp. 327-828.

Males from Yladivostock, Siberia, and Olga, Siberia, July 1928.
The specieshas an extensive range in Japan.
DICRANOMyIASPINICAUDA

Alexander

t924. Di*anot gia spinicaud,a ar,nxrNoun, phirippine Journ. sci., vol. 24,
pp. 545-546.

Specimens from Kudia River, Amagu, Siberia, July 1928 (T. D
A. cockerell). The specieshad been recorded only irom sakhalinand Hokkaido, Japan.
DICBANOMIIA

Iikewisesubscu-subfamilY T''Molv[NAE
Tribe IIilONIINI
GenusLIMONIA Meigen
Mag','t-ol'2' p'262'
L803.LimonioMnreuu,Illiger's
Zweifl'Ins'' vol' 1' p' 116'
Bes"lr'18L8.Limnobt"fvruin"i, Sigt'
LIMONIA trABAtrUTOMS Alesedcr
L924. Linowio
55q-551.

(ilt,\Nll

Genus DICRANOMYIA

G. (Lipophteps) flauocostal;i'sAlexander'
japonica(Soconilqa)
Rhabdimosti'r
Alexander.
E elobia hgbrido (Meigen)'M olophilus sericatus Alexander'
lrlexEr;'oitera (Erioptera) elngontula
ander.
E. (8.) horarlAlexander'
E. (8.) orbital;is Alexander'
n. il'iu pno"o) asgmtnetri'co Alexander'
E. (A.) aezoano'Alexander'
E. (H oplotobis) asi,oli,caAlexander'
Oropezo sa,tsumo Alexander'
N esopezo geniculato Alexander'
Cteniphor a bigttttota Matsumura'
Cneion"otis nohi,roe (Matsumura)'C . rni,plagi'atc Alexander'
P selli,iphora septentrionalis Alexander
DiAenidia fascriota CoquilIett, variety'T angptera angrrst;istylus Alexander'
T. iozana (Matsumura)'
Tipulo bubo Alexander'
T. coquill,etti,Enderlein'
T. fumi,ilo AJexander.
T. isshnkni Alexander'
T . lcuwaYomoi, Alexander'{

Sci" vol' 24' pp'
korqfutonis AloxANoun' Philippine Journ'

One fomale,AmaguVillage,Siberia,JuIy 1923.tt' P',t-Cockerthe type from Sakhalin' but-ooeU). This specim;;i;dy;osembles
praescutum'togotherwithiL"
has the lateralpoit-u'oootalcolored'thoracic ploura,somewhatdarkerl

Ii

AMURDNSIS,

new rpecie

General coloration dark gray; pleura yenow, the sternopleurito
darkened; wings yellowish subhyaline, the stigma pale; gc shortl
male hypopygrum with the mesal lobe of the basistyle complexj
dorl{dististylebent in the form of a rectangle, the aplx narrowed.
Male.-Length about 6.8 mm.; wingr g.2 mm.
Rostrum obscure brownish yellow, the palpi brown. Antennao
rolatively short, da,rk brown throughout, tho flagellar segments oval,
the more distal segmonts passing into cylindrical. r{eai dark gray.
Pronotum dark brownish gray medially, yellowish laterally. Mu"onotal praescutum brownish g"ey, a little darker medially, the latoral margins indistinctly paler; scutal robes dark; scutellum yellow;
postnotal mediotergite pale basarly wiih a little more than the aoicai
half infuscated. Pleura yellow, the anepisternum with a o"grr"'",rggestion of dusky; sternopleurito with its vsntral portion lt"o"gl:yinfuscated. rlalteres of moderate length, pale, the knobs infuscated.
Legs with the coxae yeilowish testaceous, the fore coxa,€ a littleda,rker; trochanters brownish testaceousl femora concolorous, thotips infuscated; tibiae pale brown, the tips weakly infuscatod; tarsi
da,rk brown. rvrlss
subhyaline, the base clearer yellow;-yellowish.!ig-"
only faintly indicated;voins brown. venation: BJ short,
8c' enling opposite the origin of Rs, the tip of g, extonding a distance beyond r but not reaching costal Bs relatively long, more than
twico tho basal deflection of Bn*u; basal deflection it cur'at tho forkof. M.
Abdominal tergites da,rk brown, tho caudal ma,rgins of the segments na,rrowly pale; sternites infuscated, the lateral-"rgin,6PR0oEEDTNGS oF 'frrn

NAIIoNAT'

MUSEUMvor,, 08

AD'l'4

basistyles
tergitenanowly yellowish; hypopygrum with the-andnearlytergite(fig' 1) with tho ninth
infuscated. Vf*" Uypi|*illow'
aproducodinto
tr&nsverse, tte mediio'ui"n ""ty slightlv
apex
tho
large,
outy
dark tuberclo. nuJlyr"r small, in" J"*uf tob"
ventral disblackened and set witd numorous setigerous tubercles.
18) that(fig.
rostrumt*tyr" _"aerately li*hy with a long-siender
bristles;delicatemore
boa,rsat its tip u rffiu" rpinous seta'and a fow
apartlengthown
two relatively short spines, placed less than theirvery
B'dististyle
Dorsaluna oot far from the base of th" rostrum'thebeyondapex
theri""GV curved chitinized rocl, the base stout'
thebeforea,Uate$feeblyupp*"*i-utely rectangular curve narrowed,of
angle
apicalwith the mesal
acuto straight
Aedeagus"p"*.--G*upopnyt"slobe'straight;;;h p;"J";ecl caudad into a aui<y nearlystout.
Village, Siberia'
Described from a single malo, collected at Amagu
July 1923 (T. D'A. Cockerell)
fypt.-i,fale, Cat. No. 28358,U'S'N'M'
Amer' Eut' Soc" vol'192O.Dicranomgi'opseuilomorioLt'r;xtNonn' Trans'
46, PP. 3-4.
belolgs here' Kudia
One badly damaged specimen that probably

River,e-ugn, Sibiria,iuly 1923(T'D'A' Cockerell)'Genus RHIPIDIA

Meigen1818.RhiPidio MnrGnN, Syst' Beschr' Zweifl'
BEIPIDIA(BIIIPIDIA)

Ins'' vol' 1' p' 153'

MACULATA

1818. Rhi,pidi'a maculata MurGnN, Syst'
Pl. 5, fig. 11.

MeigenRHIPIDIA

Beschr' Zweifl' fns'' vol' 1' p' 153'

(MONORHIPIDIA)

STBIRICA'

ne* specle

the basal quarter
General coloration darkl fore legs dark brown'
brown spots
darker;cellwith a gra4sn ]ing-e,jtro costal
p"f".lG"Ssout'er end of
andcord
thetip if sc, ulong
and seams o"rgri'Jht,"t Sclong, ending beyond midlength of 'Es"
lrt M;"uU
Fen'rd,e.-Length,7 mm'; wing, 7'8 mm'
througlout'-tho flagelRostrum and palpi black. Ant'ennae black
face of each seginnert'he*itnTi.tio"t Uutut pet'ioles'
fu.r"g-""t,m"otiiigUtty produced. Head da'rk, discolored'
pat'tern that might bo
Thorax uoti""if au"t*olored', uoytloo- or
palo' the knobs
Halteresnormally presont, a""t"oy"a by'molsture'
discolored' thebrown'
Legs with the coxae dark
iigrrtrv"ai*.""a.

FIIOM

SItsEIIIA-ALEXANDEN.

(ARHIPIDIA)

PULCHBASEPTENTAIONISIAtennderL973.Rhi,pidia pulchra septentrionisAr,Ex,lNonn, Can. Ent., vol.4b, pp. 206_
207, p7.3, fig. 1 (wing).

(T' D' A' Cockerell)'
One male, Kongaus, Siberia, August 1923
region'This fly is distribu"ted ihroughout the Holarctic
RHIPIDIAIILIES

7

tips brightened; trochanters obscureyellow; femora obscure yellow
basally, dark brown towards the tips; on the forelogs the femora
are dark brown with the oxception of tho basal fourth; middle andhind femora with tho extreme tips a little paler than bhesubterminal
lykened portions; tibiae and tarsi dark biown. Wiosr with a grayish tinge, the costal cell strongly infuscated; conspicuois brown spotsat origin of Bs and tip of Bc,. stigma brown, sending se&ms caudad
onto r and to the fork of 8s,' posterior cord and outer end of ceil
lst M" more n&rrowly soamed with brown; wing-tip in cells pnd R'
R_u
and.Tnil M, more weakly infuscated, especially in the centers-R,
gf tl9 cells; veins pale brown, the proarcuhr*ins
and.Cu moreyellowish. Venation: Bc, ending beyond mid-length of Rs, Bc, oniy atrifle shorter than ,!cr;'Rs long, arcuated at o"igio; r at tip o,f Br;-lst
inner end of cellMr;
-8, lying a little proximad of that of cell
cell /sf M, gently widened distally, m abott equal to the outerdeflection of. Mr; basal deflection of Curat, the foik of M.
tergites dark brown, the stornites alittie paler, yellow- ,Abdominal
ish brown, especiallythe intermediate segments. ovipositor wiih thetergal valves slender,gently upcurved, the tips acutej sternal yalves
stout, relativoly straight, the base abruptly blackened.
Describedfrom a single female, collected at okeanskaja, siboria,
August 1923 (T. D. A. Cockerell).
Tyyte.-Female, Cat. No. 288b9,U.S.N.M.
Rhi,pid,i,a(Monorh,ipid,i,a)sibi,rina is allied to R- (M.) uniseriata
Schiner (Western Palaearctic) and B. (M.) fd,etts Osten Sacken(Eastern Nearctic), difrering in the venation and coloration of thelegs.

Alerander

qrccles' near PSEIIDOMORIODICBANOMYIA,CIIANIII

I
r.'l_ Specimensfrom Vladivostok and l(udia River, Amagu, Siberia,
JuIy 1923 (T.D.A. cockerell). The northern form of the oriental
pulchra is widely distributed throughout the JapaneseEmpire but
had not previously been recorded from Siberia.
Tribe IIEXATOMINII

Genus PSEUDOLIMNOPHILA

Alexander

1919. Pseud,olimnophilaAr,nx-tNonn, Cornell Univ. Agr. Expt. Sta., Mem.
25, p. 917.
PSSUDOLIMNOPHILAOCIIBACEAASIAT! CA, new subslncles

Female.---l'ength about g mm.; wing, g mm., its width, 2.g mm.
_ Generally similar to typical ochracea (Meigen) of the Tfestern
PalaearcticRegion, difrering as follows:

I

L'Ro olcBl) lN( lsot f 1' lllr : NA' , t "loNAl' M t t s l 'l u M

vot,. otI^u1,yellow' the tip darkenod'
Basal sogmontof the flagellum lurgoly
tle-ta"si infuscated. Yflings
mg" yJf"#, tho tips oi in"-tiUiu. aod'arcuated at origin; 8'*' longorl ceil R'
with Es short and t**tt
of vein B, sinuousl cellcorrespondingly slroii"",-in1 distal section
the length,i*ritu; u*ui J"a""tion of Cu' at two-fifths;;;;ryoI' M;
fork
th9
beyondof the cell lsl Z, uoJ-i",s-1q;'; than its length
typicalt'heincoorpi"oouuly btoader thancellynd,,{ broacler.
form.
Describedfromafem.ale,collectedatokeanskaja,siberia,July
1923 (T. D. A. Cockerell)'
Tgpe.-Female, Cat'.No' 28360,U'S'N'M'
Genus LIMNOPIIILA

Suit' a Buff'' vol' 1' Elist' Nat' Ins'' Dipt'

1834. LimnophdJo Mlcquenr,
p. 9 5.
LIMNOPIIILA

Macquart(LIMNOPHILA)'

gpecies' nsr

PTCTIPENNIS(Mslgm)Beschr' Lweifl" Ins'' vol' I'
t8t8. Li,mnobia pictipenwis Ll[sranN' Syst'
P. 119.
1923 (T' D' A' Cockerell)'A female, Okeanskaja, Siberia,- Augustof this subgenus have not yet been satisfacThe Palaear"ti""p"a"itorily difierentiatuu"*no""r"arNcoNcussA Aleoorlq
Ar'pxe'r'tona'Can'Ent''-vol' 45' pp'313-314'
L9t3. Li,mnoplvilai'nconatsso--pi
12 (malehvpopvgium)'
fig'
pl'
10,
a, ns. f lwing;;
Two spec im ens t ak e n a t' T s u ru g a ,J a p a n ' Au g u s t 19,23' byP rofesThis is-the commooest and most widely
sor and Vf"r. Coctu""it.
into

distributed-u-uo
Taiwan.

oiih" guorrsin Japan, extending southwa'rd

LIMNoPEIILANSMORALIS

18L8- Ldmnobi,o nemoral'is Mnlcnw'(Moigen)'Ysrlelv

Syst' Beschr' %weifl' Ins" vol' 1'

One male specimen, Olga, Siberia' Julyp' 126'

1923 (T' D' A' Cockerell)'Genus PILABIA Slntenis
t888. Pilorio Swrnnrs, Sitzber' Nat'-Gee'
PILARIA

L818. Limnobi'oP. 125.

FUSCIPENI\IIS

fusci'pennds Mrrcou'

Dorpat' vol' 8' p' 398'

(MOig€N}'"6TICIY

Syst' Beschr'

Zweifl' Ins'' vol'l'

JuIy 1923 (T' D' A'
Specimons from Amagu Village, Siberia'
Cockerell).

OllANllrl.Itls].ROM

SIBDR.IA-ALEXANDEng

Tribe PEDfCIilW
Genus PEDICIA Latreile
18O9. Ped,icia Lernorllr,
Hist. Nat. Crust. et Ins., vol. 4, p. 255.
L916. Dai,miotdpula Mtrsvlrune,
Thous. Ins. Japan, add. 2, p. 468.
PEDICIA

COCKERELLI,

rew B0ecl6

General coloration gray and yellow; wings with the costal margin
pale brown, the remaining markings cla,rk brown but restricted inextent I basistyle of male hypopygium produced into a slender acutehorn.
Male.-Length, 20 mm.; wing, l9.b mm.
Rostrum_an{ nalpi dark brown. Antennae brown, the basal segments of the flagellum short and crowded., the terminal segments
elongate and with conspicuous verticils. rread dark gray; vertical
tubercle distinct.
Pronotum obscure yellow, the sides of the anterior notum with abrown spot. Mesonota,l praescutum yeilow with four distinct browrr
spots, the intermediate pair only vaguely separated from one anotherby apalo lino; lateral stripes crossing tho suture onto thescutal-capillary
lobes; scutellum yellow; postnotal mediotergite testaceous,darkened behind. Pleura yellowish testaceous, tho- sternopleurite
darkened ventrally; postnotal pleurotergite with a dark spot at thebase of the halter. Halteres pale, the Inobs obscure yellow. Legs
with the cox&o and trochanters light brown; femora y"llo*, the tiis
conspicuously blackened; tibiae brownish yellow, tho bases,ru""o*iyandindistinctly, the tips more broadly and conspicuously,infuscated;tarsi brown. TV'ingsof the usud, Ped,icio pattern, the costal margin
much paler brown than tho remaining dark markings, the humeral
region almost yellow; brown seam along vein curcJntinued to themargin but becoming less defined outwardly; brown ma,rkings lessextonsive than in ilaimi,o (Matsumura); a pale spot in cell R, opposite the lovel of the cord, as in ilaimi,o.
Yenation: CeIl -lst -try',relatively small; petiole of cell Mrnearly
twice m; basal deflection of cu, a short distance beyoncl tie forkof. M.
_ Abdon+inal segmentsnea"rlyuniform rusty brown, without distinct
markings in the rubbed unique type; hypopygium passing
9""F"into brown. Male hypopygium with the ventrat ali"utLgt"
or eachbasistylo (fig. 2) produced into a shiny, acutely pointed ho'rn that isgently curved, these horns directed dorsad and- slightry mesad.,theapex of each horn deeply grooved on ventral
face, the a,pex
microscopically bifid; dorsal apical angle of basistyle-u*alwith rather
numerous yellow setae. Dististyles elongate, darkened, the apex
shiny, obtuse.
5319&-25i__2

I'lto(iltl{l)lN(ls10o l" ' l' tl l' : NA' l' l( ) NAl'

M IJSIiU M

vol,. ot

^rt,p.lat Olga' Siberia' Jul'v 1923
from a single rnaltr,<lrtlltlctc<lDes<;ril-rcd(T.D.A. Cockcrell)Type.-Male, Cat. No. 28361,U'S'N'M'
of the collector'
Tilis interesting new species is named in honor
Prof. T. D. A. Cockerell.
Genus REAPHIDOLABINA'

BEAPEIDOLABINASIBIRICA'

ft;ion oI---i;;;.-Nrale.

MOLOPEILUS

new spcler,,h !ip- ;1 R, verv short to punctiform;R'u's'N'M'cai. No. 28362,

S IB Ii R .IA -A LE X A N D E F

111833, Molophr,lzs Cunrrs, Brit. Ent., p. 444.

Phila', pp' 540-541'

M'cell'M' a fitlle longer tlan its potiole; cell lst;;;;Iy ;""ated;t'woaboutto this point;-nowith the inner end poi',t*d, r-?r' ctose
deflection of Cu' about twobasalM";oithfit the outer defldction
tne fork of trI' Wings relatively broad';Jrh" its length b"v;;iwidest opPosite the origin of Rs'
bicolorous' the caudal
Abd,ornen with th;'tergites indistinctly
pale yellow;
darker brown tnaln tne bases; basal sternites-;;;;"
hypopYgium dark brown'
at Amagu' Kudia River'
Described t"o* u ti"gf" male, collected
Siberia,JuIy 1923 (T'D'A' Cockerell)'.

l |tl oM

GenusMOLOPHILUSCurtis

the terGeneral coloration pale whitish yellow; legs yollow' olly
with aminal tarsal segmenti infuscated; wings whitish subhyaiine
infuscatedbasheaqipaie brown pattern, the costai region""futi"ay, ptut"otl abdominal fergites indistinctly bicolorous'
uliy;-Male-.-Length
about 5 mm.; wing,6'5 mm'
flesh-color, the palpi pale yrcllow'- Antennae
Rostrumthe torminal"*adi"nshort, the scapal segment, reddish; fi"g"U"* yellow'
segments a litile infuscated. Head yellowish brown'
yellow' without disPronotum and mesonotum very pale whitish
the knobs atinci narLings. Ploura whitish yuUo*- Halteres pale'the.terminal tarsal
iiiliu *o"u yJliowish. Legs whitish yellow, only
a relatively-basal withr"g^""t- infuscated. Wiigt yhitis\ subhyalin-e'
hatf of cell ?nil C
heavypalebrown pattern;"cell /sf 9 Td
Bc" including a large
infuscated; interruptotl areas in cells Bc andns an$ at the tip ofbasal area and spots ut B'r, above the origin 9f
Sc':thesedarkareasincellscextendcaudadintocellRbutdonota; narrow brown seams along the cord and outer;i;""-;;-iai;" rho ourer deflection of vein .Br1 a circular,t-M,;;d]"J
tip of vein R';"r ""iilcloud at tip of vein '8, and a smaller one atbrown
and ?nil A;
spots and dots at ends of voins M", M"' Cuu^Ct4';t;;of veins Cu' lst A'
ipots at fork of Mr* and'along the bases;;;ii;tVenation: 8s strongly
and,2ni1,.4,' p"ou""rl"" "uif" Iargely dark'
uf o"igi";-a, r presentl u: i'' f. ili,*anotoiil,es Alexandsr;;;ri";;J6;;t'l l -tl l s

Tribe ERIOPTERINI

Alexandersci'
L9L6.Rhaphidolabina ALnxeNonn, Proc. Acad. Nat.

(i l t.\N tt

a

LOBIFERUS,

new sp*ies

General coloration brown, including the thoracic pieura; halterespale, the knobs yellowish; male hypopygium with each basistyle
produced mesad and caudad into a fleshy setiferous lobe; apex ofbasistyle and dististyles totalling four chitinized arms.
Male.--T'eneth about 3.6 mm.; wing, 4.b mm.
Female.-Length about 4 mm.; wing, 5 mm.
Rostrum and palpi brown. Antennae of moderate length in thefemale, broken in the unique male, pale brown, the basal segments
a little brighter. Head dark gray.
Mesonotum dark brown, the lateral margins of the praescutum alittle paler. Pleura dark brown, Ifalte'.es pale, the knobs yellowish. Legs with the coxae and rrochanters yollowish testaceous;remainder of legs obscure yellow, the tarsi passing into brown.
Wings with a pale yellowish tinge, the veins pale brown. Venation:
B.*u about twice the basal sectiou of vein Ru alone; basal deflection
ol Cur less than its length beyond the fork M; vein ?nd A ending
about opposite one-fourth the petiole of cell -ly'r.
Abdomen brown, the sternites and hypopygium a little paler.
Malo hypopygium (fig.a) with the basistyles stout, approximatedon the ventral face, each here produced caudad and slightly mesad
into a slender fleshy lobe that is provided with stout setae. Apexof basistyle produced into two long, powerful arms, the outer orlateral one directed caudad, before midlength bent strongly mesad
into a long, nearly straight black spine; inner or mesal &rm connected basally with the outer, shorter, heavily blackened, feebly
sinuous. A thbd chitinized arm is longer than any of the others,
weakly oxpanded on the distal half and here provided with small
subappressedtoeth, the apex acute. Another dististyle is present
and chitinized, but in tho unique type has been broken beyond thebase which is dilated, thence narrowed and blackened; this appendage is not shown in the figure. Aedeaguslong and slender, palo.
Describedfrom a pair taken at Olga, Siberia, July 1923 (T. D. A.
Cockereil).
Tgpe.-Mule, Cat. No. 28363,U.S.N.M.
GenusEBIOPTEBAMeigen"t"1803.

Erioptera Morcuw, llliger's Mag., vol. 2,p.262.
ERIOPTERA(EBIOPTEBA)

TUSCOIIALTERATA,

new qmles

General coloration pale reddish and yellow; halteres pale, tholicrobs conspicuously brorvnish black: legs yellow, the terminal tarsal

L2

P . l t o c u l ' l D lNo s

o ll

1 ' llt' ) NAil' Io NAL

M U S IqU M

A II'T 'I

VOlr 0a

GONOMIIA

L804. Limonia hgbrida Mrrcnx,

onespecimen,Vladivostock,Siberia,lg2s(T.D.A.Cockorell).
Thiscommoncraneflyisverywidelydistributedthroughouttherlolarctic Region'
GenusTBTMTCRAOstensacken
lg6t.Trimicra ostnwSec r nl, Pr oc . Ac ad. Nat . Sc i. Ph i l a . 'p . 2 9 0 .
TRIMICBA

1787. Tipula pilipes F tlntcrus,

PILIPEA(FAbTICiUS).II

Mantissa Ins', vol 2, p' 324'

One male, Kongaus, Siberia, August 1923 (T' D' A' Cockerell)'
If most of the numerous described species of. Tntni'cra are synonJrms(GONOMYIA),

speie,

ner

SUpERBA

AlmderA single badly damaged specimen, Amagu Village, Siberia, July
1923 (T. D. A. Cockerell),that appearsto belong here. The species
is the most,widely distributed Japanesemember of the genus.
GenusTIfABDOMASTIXSkuse
1889. Rha.bdornaslinSrsse, Proc. Linn. Soc. N. S. 'Wales, ser. 2, vol. 4, pp.
828-829.
RAAEDOMASTIXl

Klass', vol' 1, p' 57, pI' 3' fig' 17'

13

L913, Gonomyi.a (Gonomg,ia) superba Ar,nxauonn, Can. Ent., vol, 45, pp.
285-286, pl. 3, flg. 14 (wnre); pl. 10, figs. 1-2 (hypopygium).

Ins., vol. 10, p'
tg25. Helobiasr. F.l.ncsau et snnvrr,r,n, Encyclop. Method.
585.
6' p' 282'
1830. Sgmpteclo Mrrcuu, Syst' Beschr' Zweifl' Ins'' vol'(Melgen)

S IB Iq& IA -A LE X A N D E R ,L8L8, Gonomyia Mnrcor, Syst. Beschr. Zweifl. Ins., vol. L, p. 146.

Genus IIELOBIA St' Fargeau et Serville

IIIBRIDAl ,' I-IIqS Irl i OM

oI pili.pes,as would now eppea,rto be the case, th" p""rurrt fly has
the most oxtended range of any Tipulid, being found in virtuaily allparts of the world, including many of the most remote oceanicislands'
GenusGONOMYTA
Meigenyellow; ma,IesegmentsinfuscatedI vringspale yellow, the veias darker
blackened
acuteio
fslminatiog
goiupophytss
thewithfty}"pygi"horns.
Male.-Letgth about 4 mm.1 wing, 5'4 mm'
aJd palpi brown. Antennae pale reddish bro.w-n, thenort.o{,s1minalsegmentsmore infuscated. Eyes large, contiguousbeneath'
Head yellow
marginsMes'onotum reddish, more or less discolored, the laleral
ofthepraescutumandscut.umlightyellow,thiscolorextending
the postback to the wing-rootl scutellum and lateral margins o{
it isnotal mediotergite similarily yellow. In normal specimens
redp"r.nf" that thls yellow coto" is mueh more extensive. Pleura
black.
Ilalieres pale, the k"ob: conspicuously brownish
ii.n yJf"*.
the terminal tarsal segments infusc?iud' , Wings
only
y"ffo*,
l,"gt"veinsdarker yellow. venation: As in the subgenusl
pJ?y"u"*,',the
9nil Ah ,.', J""t one-half longer t'han the basal section of Rr; vein'very stronglY sinuous.
Urownish yellow. Male hypopygium with the basistyles
afao-"ibase.
rolatively stout. Ouie" dististyle apparently broken off at
theat
ai'*tistyle (fig. 3) a strongiy curved arm, pale,excepti;;";
appea'rsuddenly pointed apex which is blackonod' Gonapophyses-blackened, thel"g * ,lf.iau", o"*iy straight horns, the_apical third
acute. Aedeagus lyriform, eactr half provided with a long,ffi
andc,r^"v"d branch that" is directed caudo-laterad, thenco mesad
finally stronglY cePhalad.
Described from a single male takon at Amagu village, siberia, July
1923 (T. D. A. Cockerell).
Type.*Male, Cat. No' 28364,U'S'N'M'

EELOBIA(i l l ^N l i )I(SACANDAGA)USURIENSIS,

new spctea

General coloration black, light gray pruinose; knobs of halteresconspirruously light yellow; wings with a strong brown tinge; veins.8, and .B, widely separated at wing-margin; Er longer than Br*r.
Male.-Length about 3.5-3.6 mm.; wing, 4.F-4.6 mm.
Rostrum and paipi black. Antennae short, black, the flagellar
segments oval. Head black, heavily light gray pruinose.
Pronotum and mesonotum black, light gray pruinosel pseudosutural foveae black. Pleura gray pruinose. Halteres dark brown, thoknobs conspicuously pale yellow. Legs with the cox&e and trochanters black, gray pruinose; femora and tibiae dark brown, the basesof the former vaguely paler; tarsi black. Wings with a strong browntinge, the veins still darker brown. Venation: Sc, ending Eetween
one-third to two-fifths the length of the long, nearly straight Bs, Sc,
close to its extreme tip but relatively indistinct; ,R, oblique, at thewing-margin separated from the tip of Rrby a distance greator thanits own length; Er longer than Rr*r, cell B, thus being relatirely large;
2 ]ssking. Anal angle of wing moderately developed. No macrotrichiae on veins Rs, Rra, R, ot R"; numerous trichiae on Br+ufor its
entire length.
Abdomen black, gray pruinose; hypopygium black.
DeScribed from three males collected at Okeanskaja, Siberia,
August 1923 (T. D. A. Cockerell).
TEpe.-M*lo, Cat. No. 28BOE,
U.S.N.M.

74

I ' 1 r (X jtltr ) t) lN( ls t) tr , ,l,il l,l NA,t' lo NAI/

Ml .Jsl ttIMvor,.0al rrr. ,lSubfamilyTrculrx-m

GenueDfCTENIDIA Brull6
1833. Dtcten;id,iaBnv:,;.,fi,Ann. Soc.Ent. t'rance, vol. 2, pp. 4OL-402.
BIMACIJLATA(Linreus)

1761. Tipula bi,macul,ata
Lr*wrnus, Fauna Suec.,ed. 2, p. 433.

One male, Kongaus, Siberia, August 1g2A (T. D. A. Cockereil); epair, Kudia Biver, Amagu, Siberia, Juiy 1923 (T. D.A. Cockerell).
A critical comparison with European specimensreveals no differencos
between these flies.
Genus TIPULA Linnaeus
1758. Tipula Lrur.lrus,
IIPULA

Syst. Natur., ed, 10, p. 585.
COCKERDLLIANA,

new srrcies

Allied to T. fulatpenni,s d,eGeer, ?. sh,i,rabi,i,
Edwards, and similar
speciesl general coloration grayish yellow, the praescutum with fourbrown stripes; wings with a strong yellowish brown suffusion, theobliterative band before the cord very broad and conspicuous,
cream-colored.
Female.-Length, 28 mm. ; wing, 23 mm.
tr'rontal prolongation ofhead relatively elongate,brownl n&susconspicuousl palpi dark brown. Antennae with the scapal segments
brownish yellow; flagellar segments feebly bicolorous, the basal
enlargement darker brown than the ground-color; verticils long andconspicuous. I{ead dark brown, grey pruinose.
Mesonotal praescutum grayrsh yellow with four brown stripes, theintermediate pair na,mowly separated from one another by a grayline; scutal lobes dark brown, each with two dark gray markings;
scutellum brownish yellow, the center of the disk grayish brown;
postnotum dark but heavily yellowish pollinose to virtually obliterate the ground-color. Pleura yellowish testaceous, the posterior
margins of the postnotal pleurotergite weakly infuscated. Ilalteresbrown, the bases of the stems and the apices of the lorobs paler.
Legs with the coxae and the trochanters yellow; femora yellow, thetips broadly and conspicuously dark brown, the tibial apices morenarrowly infuscated; tarsal segments one to three brownish yellow,
the tips of the individual segmentsnamowly darkened, the torminal
tarsal segments uniformly infuscated. Wings with a strong yellowish brown suffusion, the stigma da,rker; cells boyond the cord somewhat darker than the basal cellsl a brown cloud near midJength ofcell Cu, preceded by a clearer yellow area, the latter continued into
celJ,M; a, very broad and conspicuous creamy obliterative markr,'l IItS tfROM

SIBEEIA_AIEXANDEn

15crossosthe wing.before the cord, narrowest
in ceil c at the tip ofvoin 8c, and at the. posterior marginof the *i"g;;iotn
otvein Cu,; this are-aincludes *o." ihuo the"iau.
basali"UlstMr;
additional pale blotches in outerhalf of.cell R;irr;;r"-"t ""Uof cell M,
and restrictedly in
gyd M; veins b"o*,"iul""i.,tUu
costal3:f
region and radial field. venation: B,
persistent; rn longer than the"."
petigi-eof.cell Mr; m-at punctiform ; i"i e"A i;;#.
Abdominar tergites brownish y"ilo*, thebase of the abdomenbrighter, the segmentswith ilr-dednedsublateralu"o*rr-"t"ipes; posterior segments with incricationsof a
median vitta; sternites more
uniformly yellowish. ovipositor withthe varves long and straight,
especially the slender tergal valves.
Described frol a. sinlle female, taken
at Okeanskaja, Siberia,
August 1e2B(T. D. A. CJck";;lD.--' "Type.-Female, Cat. No. 28866,U.S.N.M.

Tribe TIPIIIIIIDICTEMDIA(llt^Nll

Anrennaeormare;:Hffiffi;;;";"il1"0"- brack,rhebasarenlargement of eachqegment
lighl, yellow; wings brown_"o.rrpi"ui.rrty
ish yellow: ce,. M. short-petiolate
to ;;il;;;;il;;;;;;;1um
wirh
the eighth sternite produced cautlad
rnto a conspicuousriguliform
blade.
!ale.--Length, 12-tB mm.; wing, 11.2_tt.b mm.
tr'rontal orolongation of head reiatively
short, brownish yellow;
long but relativery stout, concolorous.
Antennae mocreratery'asus
elongate,if bent backward, extending
about to the baseof the abdomon; scapat segments
the]is_ht lr9wnf flagellar ,rg_;;.-;lack,basal enlargements of theindividuar segments conspicuously rightyellow, this greater in amount on thobasal segmonts,becoming more
restrictod in extent, o:outer_segments;verticils'relatively shortth"
but stout.
-o"ult"rreadright brown,th"

indistinctlyil.-"
Pronotum testaceous brown. Mesonotal
praoscutum obscurebrownish yollow with four_slighily d;k;;,ilt-defined uro-*o,i"n stripes;
centers of scutal robesda,rkened;-remainderof the ;".r""tr;brown_

ishtestaceous.pleurawhitirhi"stuc"ouqril" pr.*"t"igir"T-rghier.
rr.a|!.e1os
palobrown,thebaseof thestempaler.' t"g, ;-thiilu
whitish testaceous: trochalters pate y"uow;femora-and tibiae "o"uu
right

i
I

I

brown, the tarsi oassing into dl,rk il;;*":
Wings with a srrong
browaish yellow t-iogu,inu base andcostar region more ye'owish;
ljigma small, oval, pale brown;-a,obliteratiye axeabefore
the stigna, extendi?rg frgm_ve11"oo*pi"uousB;f"f*"and including the cord,crossing thoproximar end of cerr-lst'rt
i,,to th"u*.
otl"fi l.; veio"
brown. Venation: Bs mod.eratelyelongate,arcuated.,about one_halflongorthan-R,*r;B, preserved
thro"ghi"t its tength;'*tiiiiAb"sNAtloNAr't'ttoL:oltDlNcs orr 'l'tltl

16

MusEUM

votr'o8

longer thanmore or less poinled' mbeing&nd n&rrow, its outer end
vein rs very
somtr specimens' the latterthe petiole of ceII !r; in'ld being sessilel nr-ctl
the lattei""i1
short to obliterateo,-in
"""ir
"u"u
broad'

moclerately

hnil'A"^u* ut":?i:"i"#"1"'
ffil;; ; purut,oil'i,'lnu.
:::"l*k ll"fJy]i:fj lX:

A tr,T ,I

orr^Nlt

ITLIU$ FITOM SIBERIA-AI/E)L|NDE&

17

of thr: knobs obscuro yellow, tho base of thestem narrowry parer.
Legs with the coxae, pale, sparsely pruinosel
trochanters yellow;
femora brownish ye'ow, the'tips
bil;k;;;d;tibiaelight brown, the tips narrowry da,rk b"own;"on.pi"uousry
tarsi dark brown. wings
with a grayrsh brown tinge, ihe base and costal
region yellow; stigma
brown;
conspicuour",otnyuri,,e are&fr
l"o*i,wing-apex .dalker
beyond thecord an$ st{qma, appearing as an oblique binJfrom
thecosta-through cell lst M, into-the baie of ceilfin; a smaner palearea before the stigma ,L"ll Ist Bi; a large puf",pot
fut*u the endofcrossing'vein cu into celi' cu; u small pale sirot
on vein.cell -tr[,
lst A near one-third its length, the spaco between
these two latter
subhyaline &re&s & trifle more infuscated trru"
1n" g"ound-color;
anterior cord and origin of Bs_slightly infuscated;
o"io"-b"o*, dark)'ellow in the flavo's areas. venotionr a, p.u.""o"d, *ra"ry divergent from.Br; petiole of cell .I4, shorter iaio*;
irJ M, elongate, its principal faces parallel.
""U
Abdomen with the.basal two tgrgrtes yellow, theremaining ter_gites brown, each with gray laterar uiramargins uoJ-u m"diuoand sublateral dark brown longitudinal""rautstripes;tn"
segment's-,
the pale rateral margins &re .,nerybroad"i and conspicuous,-"tierminal
more yellowish; sternites generaily similar
with p"r;;;rd;I;argins.
Male hypopyglum (fig. 6) as in the hebesgroup; tergite
dark brown,;",];:nr:lffi3l"ltrJ't'f'l{-ii'*:i:il#J'ile#:,-mt
outer dististvreelonnot
3n" t"tgi*" (fig' 7) wit'h a V-shapect

providedwith micrl:":ni9'?i;:i i:"Y1";: in a conspic-uousblack-gate,

terminating
fleshy. JnSer disiistylenarjnth-sterni'"
l::?Y::d."fthin'sDine,d-irectedoo"ui'""^nlgl"
conspicucus'
J""kt" bearing a' very
r-owly incised. Eig;;;
in color'
vellowu"arislrt
il;;it'ioo
liquliform blade,collectJ at Okeanskaja' Siberia'tI"""
Described f"o--;d;"-iul"''(T' D' A' Cockerell)'-e,Ie-riJ1s23-- U'S'N'M'^"ft;;;.:iaie, Cat'No' 28367'
TIPULAAIIIURENSIS'

new sp€cies

elongate' the flagellum uniBelongs to the h'ebesgro'tp; 3ntlPle
formlydarkbrown;g"i""ofcolorationlisht-sray,thelateralprees#t'n tn" eighthsternito

rtyp"pygi;tcut,alstripesitl-dodn?i';"-"r"short-hairY'denselY""
13 mm'; wing' 14 mm'
i"it.-l'ength,y"uo*, with an indistinct darker

i'"*sesFronral p"otoog##ii'i,"J
palpi
1l^u'1t11m'rnal""i;f"oiu" brownith v:lio*''lateral line; nasus";#';;isegments
terminaltheelongatei
ments a little d"";;'
to beyond the basou;;*urd,."*i"oding"firstsegmentof flagelbroken,if entire "#i'""tv"iro*;of the abdomen;##;*i1i',.J"tinto $arf tro11^; roma'rnpassing
haif,of theIum brownirn y"ilo#o" uirrr
dark brown' tni t"tut enlargement
anterior
ing segment'or nugai;thewith
U"i"-brighter' I{eadproximal"*"ot'l"l'bufiy' the remarn"ily
the alntennalbases
bart o{ the vertexls;";;di"gmedianline'brown
an indistinct
der of the vertex it;il;;;;;thMesonotuma trifle darker medially'
Pronotum b"fi;;"fi;#'stripes'brown
pof;i ili-marked
clear gra,v,tn" p"u1""JJi'i*'J'n-t
from one another andintermediat" i" #iti";;o"*"o*ty-teparated
the anterior halJ of thea""toil';iU;odivided"nfurther"iii'
gt.t than the ground-color'theirsclerite; lateral t?"i* Ji".oiign* scutum light*i1l tt anteroUoJt'"ed by brown;'grayanterior
a grayenclosingcirclebrown"od"postero-mesalof slateral brown spot and adir[y ryy yi'h-Tiications
poitnotum
andscutellum
center;ptu*u #tn tn" anepisternumand
capilla,ryau*x""Llttt#;;;'
pt"*it"" andpostnotaI pleurosternopleurit" u;1;;'ilult"'us
J\t n""' "11"1 Utownishyellow' the tips
tergite *or. *niti'fi'"t*i""'

th.",nea,rly
irunJate,*ith u o.ry

i5lT::,p:3:yt"l"i".dilTarginrow,
Dlunr,medran lobo thatis light yellow (fig. b). outer dististyre
flattened, irregularly ov_al, palel fentral'i."utp;;;";of thebasistyle produced mesad into a pare, submembraooirs;l;-.1" that isirregularly bifid at apex on *"d]uo hne, thedorsal a,rm with themargr+ narrowly blackened, overrying its mateof the opposite side;
posterior lobes directed ventrad u" uo'upoo-likestructure that liesin the notch of the eighth srernite. Gonapophys",p""i;"Gventradfrom this]1otch, chitinized, na,rrowed to the iip"'*ni"nu,i" w"uttytoothed.
.Eishth sternite deepry incised, the incision densery fiiledwith relatively short, golden-yeliow setae.
Described from a sin_glemale, collected at Amagu,
Kudia River,
Siberia, July l92B (T. D. A. Coclerell).
Type.-Male, Cat. No. 28868,U.S.N.U.
TIPULA LATEMABGINATAAterands
7921. Tipulolatemarg"inata Ar,nxaxoun, Ann. Ent.
Soc. Amer., vol. 14, pp.
t2&-L29.

O,nemale, Amagu, KudiaRiverrsiberiarJuly 1928 (T.D.
A. Cockereil)' The spocieshas an extensive atutributioni" i;G:-'

18

pnoonDDrNGS orrrnli)
TIPULA

ArrT.{

NATToNAL MUSEUM

YOr r .88

BUBO Al€renderto rhe,r""a*-ilrJ"iipu*. Me_
fl33"ully*l:d B,reaof ninth
ctran
sternitemembranous,on either side of themedianline with a fleshvflartenedtor", aire"tedcaud"d,lh; ;;;"" margin-'eiuito
fringodwith rongyolow r"tu"
sterniteunarmed.
Described*"*.
at Amagu, Kudia River,ljA*t"
Siberia,July 1e2B_"-o-Uu"ted(T.D". A.-ut",
c;;L;;"li).
Tgpe.-Male, Cat. No. 28869,U.S.lV.lt.

One male, Okeanskaja,Siberia,August 1928 (T. D. A. Cockerell)"
The fly had hitherto been known from sagharien, rrokkaido and rfonshiu, Japan.
USUBIENSIS,

newsl)ecles

General coloration gray; wings pale yellowish brown, tho stigma
pale brown; abdominal tergites bilineate with dark brown; malehypopygium with the sclerites fused into a continuous ring; em&rgination of the ninth sternite with two fleshy flattened lob-es,margined with long yellow setae.
Male.-Length about 12 mm.iwing, lB.E mm.
Frontal prolongation of head brownish yelrow above, paler later_ally; palpi dark brown. Antennae moderateiy elongaie, if bent
backward, oxtonding about to the base of the abdomen; black, thesecondlcapaf segment and the base of the first flagellar segment
brownish yollow. Ilead dark-colored, gray pruinoru, th" anterior
part of the vertex obscure yellow.
Thorax of the uniquo type badlydiscolorod; gray, the praescutum-pleurawith four da.rker stripes; scutellum largely palo.
pale, with
spa,r:sebrownish markings; postnotal pleurotergite likewise pale.
r{alteres pale brownish yellow, the base of tne stem brighter. L"g,with the cox&e da,rk, paler apically; trochanters yeilow; femora
browdsh yellow, the tips oxtensively brownish black; tibiae brown,
the tips gradually deepening into black; tarsi long and slender, d.ark
brown. Wings with a pale yellowish brown tinge, the base and.
costal region somewhat brighter coloredl stigma-relatively small,oval, pale brown; obliterative areasbefore the cord reiativeiy indistinct, more evidont before the stigma at the end of gs and acrossthe
proximal ond of cell ,/sl M; vains dark brown, thosein the flavous
areasbrownish yellow. venation: 8s very long and almost straight,
in alignment with Rr*r; tip of .8, atrophied, the subterminal sectionof the vein being in alignment with r which bears macrotrichiae
throughout its length; distal section of vein B, entirely preservedl
cell M, nearly twice its petiole; cell isl Mrlong and narrowi nl-ar
distinct, about twice its length beyond the fork of tr[; cer| pnit, a{reiatively n&rrow.
Abdoninal tergites bufry yellow, with a sublateral brown longitudinal stripe on eithor side; hypopygium darker. Male hypopygium_with the tergite and sternite fused in a continuousring (dg. 11).
Ninth t-trgite (fig. 10) relatively small, tho caudal ,ou"gio with asmall lobe on either side of the median line, the ape* of each lobe
blackened and densely set with black spines; caudo-lateral anglosof the tergito subacute. outer dististyle cylindrical but very shirt,

19bearing conspicuous
black setae. fn_nerdististyle a long-nanowed
flat-l_ong..
btad:,

19t8. Tipula DaboAr,nxenonR, Journ. N. y. Ent. Soc., vol. 26, pp. 69_20.

TIPULA

CIId.NU rLTEg TROM SIBEEIA_AI/EXANDEN,

Generar€;;-t-;;

rffia*shrhebaseandcostat""r"r"rio"regionstronglvyelJw, tt" airt aiffi;;;;e tingo,
brown;
a broadbandof the q:ou1a-colo"b;;o"t th"-;;"di"r"ty,""_o":":r- the_wing; tip of vein .8, atrophied.
Male.-Length about 18 mm.; *irr*,, _*.
prolongation of head yettoii"n gray;nasus distinct; palpi^.,-F"oltuldark brormr.
elongato,
Antennae of modirail leng;L, if -blntback_beyondrhewing_root;
scapal,.g_"ii,II;;?lrt nug"l_:It""dingIar segment
yellow;'ITq
flagellar s"gmeni" yellow, the basal en".:T1i"Tglargements of the individuar;"d";tr
itu"t; on the subterminar andouter segments,the ground-coior becomesmoro infuscated. rlead
dark gray, the anterior pa"' of the o""t"rmore yellowishl vertex
with a vaguely indicated brown medianline.
Pro-notyy gray, the anterior notum i"*ia
d.orso-media'y and more..
tinged with brown. Mesonotut p"a"scutum
light gray with fourvaguely defined brovnish gray stripesthat are better delimited near
the suturel scutum light gray, the ientersof the lobes marked withja.rker; scutellum gray, more or less infuscateJ;;;Uy;;rnotum
light gray. pleura clear Jighr_s"";; th" d"r;":;L;;;iil""_r"uoubufry-yellow. rrarteres pareleilJw, iheknobs int rscateJ. Legs \Eiththe coxae light gray; trochanters da'k ye'ow;
r.-o.u-u"ofri"ny"tIow, ne&rmid leneth passinginto b.own
i tibiu"'U"o*, iilu iiol nu"o^_ing still darker biown; tarsibrownish
biack.- wing" wiih aiaint yellowish- tinge, the base, costal region
uoa th" space behin d, vein Cu
strongiy yellowish; stigma oval, -darkbrown; a smat brown spot at
origin of Bs and conspicuousseamso" tn"b.";;;;ib",o"i*n*_
clouds in the cells, darker at the
restricting the ground_
T;;ip"*,
color to a broad conspicuous cross-baid
beyond.the cord, extending
from the costal margin beyond thesffia,
pasging caudad through'tit
the basesof cells 8.. nu, Bi, anda,,'tai"ghu, t"
the wing-margin;"i?}ril ir"i-gJy "ilp"r"""iI mid_
*rth
a
brown
croid
near
""ir
length and anothera! i!s_o,.tui*a';rargerysufiusedwith
pale brown,interruote{ b{ u putu u""u""ii'zur.4
oear the basal third; veinsdark brown,paler in the huoou" u""*.Venation:Tip of vein g,

20

PB(X)INEDINOSOr'I:IrU

NATIONAL MUSEUM

YOIA 0ll

abruptly atrophied, the basar spur (about one-hau tho total
rength)provided with macrotlo.h-r"j aaand petiole of cell
subequal;il-cupunctiform, near ono-third the lengtl of the oell. -ilr',
Abdomen with the basal segments yelrow, the tergites with arather indistinct dorso-mediaa stripe that is interruptei at theposterior margins of the segmentsl su6terminal stemites darker.
MaIehypopygium (F. Sl with the ninth tergite large, the LJ"ruflobes
produced c-1u_d1d
into flattened, obtuse f,rod.", inl m"sat edge beinggro{ucqd slightly into flattensd chitinized flanges; caudaliotch of
tergite broadly rounded, tho dorso-median pirtion of ihescreriteproduced caudad for a slight distance into thii indentation,
which isthus feebly bifid at its base (fig: 9). outer dististyreleratively-ventrallarge, pale, somewhat flattened.
mesal maxg; oibasistyle"p"o"ia.dan oval, dark-colored fleshy lobe, tho mesar
with
"ag"
spaxse erect yellow setao lying across the genital
Eighthsternito with the caudar margin provideJ with an"hr,mb"".
even fringi ofconspicuous elongate yellow se-tae.Described from a single malo, collected at okeanskaja,siberia,
August 1923 (T. D. A. Cockerell).
Type-Male, Cat. I{o. 28820,U.S.N.M.

'

GenusNEPEROTOMAMeigen
t803. Nephrolorao
Mnrcsu, iliger,s Mag.,vol. 2, p.262.
1834.Pochyrrhi,na
Mi,:aqvenr,Eist. Nai.'Ins., fiipt. vot. 1, p. gg.
NEPEBOTOMA yIncATA (Coqrr||lett)1898. Pu,hgmhinovirgatoCoqurr,rJErr,

proc. U. S. Nat. Mus., vol.

21, p.806.

. several specimonsfrom various siberian rocaritios appa,renilybeLonghore. vladivostock, r92a. okeanskaja,August i&a--roai"
River,4-€",
July 1928(T. D.A. Cockereilj. flo ,p"ciesis common and widely distributed throughout the Japaneoofmpire.
NEPIIBOTOM.I

COBNICINA(rrnh.ous),

vsdGty1768. Tipul,oe,ormdcino
Lruueous, Syst. Natur., ed. 10, p. 6g6.

Threo males from Okeanskaja,Siberia, August 1928 (T. D. A.
Cockerell).
NEPEBOTOMAACITLEATA (Loer), vadsry

1871. Pae;hynhhwuutred,aLolw, Beschr.Eur. Dipt., vol 2, p.20.

_.4 ma,l.e,Sp"gu VjUuegrSiberia,July 1928 (T. D. A. Cockerell).
The typical forn is from the TV'ostempuluuu*ii" Rqgion.

oR,ANE FIJDs lRoM srBEBra_arrExANDEBZL

DXPLANATION OT'PLATE
c:aedeagusliI

D:basistyle; d. d:dorsal dististyle;.g:gonapophyee;t:te,rgrtei u. d._ventral dististyle.

s:sternite;

8ro. 1. Di,crawmAia atnurmsi,\ newspeciesl male hypopygium,.d: Dorsal
dististyle, enlargedl B-Tip oi,ppendage, enlarged.
"o.tfrypopygi,rn.
cockerelli,,,new.specieslnufu"l?. Yjdt
Basistyle.
3. Erioptera fuscohalterota, new speciesl
Jale nypopygium.
4. MolophiLus lobiferus, new species;
male hypopygium.
Enlarged figures
show different aspects of dististge.
5. Tnpula amurmsis,,rrew species; maie
hypopygium.
Ninth tergite.
6. T. amutensisl male nypopvgiun.
i"iJrur*purt.
7. T.ligulifera, o"w rpe"iurl--"f"nypopygl"*.
Ninth tergite.
8. T. sibirienrzs, new species;
Lateral aepect.
'
T";;d;;;i"_.
" -1il-"if,

mate
,1.
:y""szs ; new hypopygium.
Lv- I.
t. uaurteiTa?s,ll.

r"rsit".

speciesl male by_popygiuml Ninthtergite.
T. usuriensr.s; male hypopygium. "i##aspbct.

U. S. NATIONAL MUSEUMPROCEEDIN(IS, VOL. 68, ART.4,

nno|' il )lg r*ry-,ru

CRANE- F LIE s FR oM S IB E R IA
FOR

ExpLAi^Ttotr

o F PL ATE

sEE

p^oE

,l

PL. I