9lJ

NEW OR IITTIE.KNOWN TIPUIID,€ FROM EASTERNASIA(DIPTERA),[J(IIBy Cuerurs P. Arrrxeronn
Amherct, Mossochusetts

lgE

Separatefrom
PHILIPPNE JOT'RNAL OF SCIENCE

Published by the National Instihrte of Scienceand Technology
Manila, Philippines
Vor.uuo 96, No. 1, Mmcu, 1g6l

NATIONALXAIVIIA
SCIEhICE DD|TTEI{)PUEVT BOAND
PBINTING PNESS
1008

NEW OR LITTLE.KNOWNTIPULID.IEFROM
EASTERN
ASrA (DIPTERA),IJ(IIBy CuanLnsp. Ar.rxeNnrn
Amherst, Massachusetts(Recelved for publication, OctoberS, f966.)

SE'\ZENPTATES

Tttis paperis devotedto a eonsiderationof the Indian speeiesof the Eriopterinegenus GonomyiaMeigenand isbasedarmost
entirely on the very extensivecollectionsthat were
madeby Dr.
Fernandschmid, as discussedin severalearrier partsof thisseries. Gonomyi,ais welr deveropedin this area,
there
known to me presenily some 120 speciesdistributed gbeing
insubgenera. It may be noted that in the earrier
studiesmade by
EnrieoBrunetti (1911-1924)l onryB species
wereknown,pertain
ing to 4 subgenera. The most sitistr.toty aecountof
the strueture and interrelationshipswithin the genushas been provided
by Edwards.z
Gonomyea
is oneof the major generain the Tipurida, incruding
hundredsof sp_eeies
arrangedin ts subgenericgroupsthat arelistedbelow. It may be notedthat whilJcertain of tt
ere groups
are eloselyallie_da1! may prove to be difficult to maintain,sev.
eral are unusuallydistinctand on further studymay proveto
representvalid genera. On the basisof the pr.run.eor loss of
the anterior arculus of the wings, the subgenerathat have retained this element include protogonomEii,Etli,pteroid,es,
Id,i,ocera, and some smaller groups,whereastypical Gonomyiaand-tr'rom
Li'pophlepshave lost the anterior arculus.
the hypopygial
structure,particularly the redeagusand the nature of Uranctringof the dististyle,it appea,rscertain that protogonomyin,Ettifi;
roides and someothers are related to ld,i,ocera,
despitethe very
different generalappearanee.Protogonomyiain the local faunar Alexander, C. P. Classifieaiion and synonymyof the erane-flies describedby Enrieo_Brunetti(Diptera: Families piyctropteride, Triehoeerida,
and Tipulide). Ree.Indian Mus. 59 (1961) 19_84.
2 Edwards, F. W. British short-palpedcraneflies.
Taxonomy of adults.
Trans.'Soe.British Ent., part I 5 (tb3s) l0s_112.

29

30

The PhiltpTtineJourrnl, of Science1967

or dat'l1
irrcludesabouta seoreof species,virtually all beingblack
Iack'
coloredflies of large size,with vein Rg (r of someauthors)
ing anclwith the small fleshy valvesof the ovipositorsetiferousisviituatty to their tips. The generally similar Ellipteroides
much like Protogonomgiaexceptfor the normal elongatesclero'
tized ovipositor. The newly proposedsubgenusRamago'1sPu1irt
is the only one in the local fauna that has vein Rz (r, as above)
preserved,as in the New world and Ethiopian group Praooro''nryto.
THE SUBGENIERA OF GONOMYIA(Amber stu'ElectrogonomgiaAlnxlNonn, Bernstein'Forschungen(Electrogonomyio\.
dtes) 2 (1931)118,figs. 158,159,as gonofrl,ai'aType,pinetorumAlexander'Fossil(BalticAmber'UpperEoccnc)'(190r) 239.Tvpe, piceus
EllipteroidesBncrnn, zeits. Hym. Dipt. 7
Becker (North Afriea). Palearctic,Oriental'
Philip. Jour. sci. 66 (1938) 126-12?, as
Alnx^e,NDER,
Eu,ptilnsteno
Gonun'gin(Euptitosterw);nameomittedthroughatypograplricel
Alexandcr
error, tine 6 tiom bottom of page' TyPel-rett'culato
Neotropical'
inrri.rn patearctie) Holarctic, northern(1818) 146. Tl'pe,
Gonomgw MulcnN, Syst. Beschr' Eur' Dipt' I
tenella Meigen (Europe). Virtually Cosmopolitan'l (1946) 5 - 6 , a s
GonomyiaAlnx^exonn,Almeida Commemorative' Type, dntabi'tts Alexander ( Br azil) .
Gonimuia (Gonomui'na)
NeotroPical; Brazil.
433' as Ltmnobia
I(li,oceroDrtu, Ann. Mag. Nat. Hist' 8 (1842) 431'
BsncnorE,(ldiocero). Type, seriuttata Dale. Sunonvm-Pttlostena
rectt"n:ht'ert:i's
Ann. Mag. Nat. ifitf (S) tl (1913) 575-5?6; type,in Austra'
Bergroth. chiefly Holarctic and oriental, fewer species
lasian, Ethiopian and Oriental regions'
Bsncnors, Psyehe 22 (1915) 55; renaming of Letponetto
Lipoph'tep.s(1913) nerv
skuse (1889), preoccupied. lYpe, skuset,Alexander
inname f.or gradlis skuse, preoccupied. Almost cosmopolitan,cluding many of the remote Pacilic islands'
as Go'
NeolipophlepsAlnxaNDEB,Rev. Entomol. 18 (L947) 98-99,
Nerv
no*olio (Ncolipophl,eps). Type, cinerea Doane. Entirely
World and chieflY NeotroPical'
porotipophteps Ar,px^txpnn, Rev. Entomol. t8 (1947) 9? - 98' (asGoiomAio (Paral.i.pophleps). Type, plewalis Willistcn. Chiefiv
Neotropical, a few in southeastern Neartic'
progonungb Ar,sxANDEB,
cornell univ. Agr. Expt. sta. Mem. 38
af.Gonomyel'
trggrl 938,as Gonomaia(Ptogonomgi,a).Renaming
preocett'(1917)
1'52,
Ia Alexander [Ann. South African Mus. 17
piecl bl- Gonomvellcl\{nuNrun(1S99) 334, fie.l. Tt'pr, :':i,s.qn??de06'I

Alerander:Ti,p,,tid,efrom Eartern Asia,LXn

Bl

Alexander (southernNeartic). Neotropieal,Ncarctie; sguthern
Ethiopian.
ProtogonomanAr,nxarnnn,philip. Jour. sei. 55 ( 1934) 52 - bs, as
Gonomgia(Protogonomaia).Type, confluentaAlexancler(Formosa). Oriental.
PtilostmodesAlnxaNDEB,
Archiv fiir Hydrobiologie,suppl.-Bd,,,Tropische Binnengewasser"
2 (1931) lB2, as Gonom?Jia(ptitosrenod,es), lVpe, ptilostenelloAlexander (Java). Oriental.
Ramagonomyia
subgen.nov. Type, protensasp. nov.
Shtagonontyia
subgen.nov. Type, d,iscotophallos
sp. nov.
T euchogonomgiasubgen.nov. Tlpe, noueboracensisAlexander.

Palneogonomyia
Meuniert(1sgg) Bsgl is now placedas a subgenusof.Rhnbdomasti,x
SkusefseeAlexander,Bernstein-Forschungen 2 (1931) 1111.Fossit(BalticAmber,upper Eoeene).
SugenusPROTOGONOII,IYIA
Alexander
GONOMYIA (PRoToGoNolrrrA)

ACUSTYLATA Atexander.

ptrte r, fig, t0.

Gononaio, (Protogono'tngia) acustglata Ar,rxarnnn, Jrrur. N. y. Ent.
Soe. 70 (1962) rl-r2.

Tlpe-male, Tung,sikkim, altitude4,b00feet.August2, lgbg.
Malehypopygium(Ptate1, fig 10).
GOM)MYIA(PROTOGONOMYIA) APICULATA sp. nov.

Plate l.

t!g.tI.

Generallysimilar to Gonornyin(protogonomYia\nigriTtes( Brunetti), differing espeeiallyin the strueture of *re hypopygrum.
Male.-Length, about 6 millimeters; wing, 7 to 7.2: antenna,
about 2.
Femnle.-r'ength, about 6 milllmeters; wing, 7 to 7.2.
Generaleolorationof the entire body, including also the antenne, halteres and legs, black. Antenna with flagellar segments long-oval,subequalto the longestverticils. wings slight[rblackened,stigmalong and narrorv,d.arkerbrown; veins brownish
btack. Venation: sc relatively long, sc, ending shorfly before
fork of Rs, Sc1,alonemore than one-halfRs.
_ Male hlpopygium (prate 1, fig. 11) with outer lobe of basistyle,b, relatively slender, with a conspicuousglabrous sclerotizedflange on mesalmargin. outer dististyle,d,,-atapex dilated into
a subcircularhead; intermediatestyte longest,beyondmidlengthvery slender, the apex triangularly expanded,teiminating in 1nappressedvery slender spine,apiealmargin of blade mieroseopically notchedto p,rodueetwo very unequaldivisions:inner style

32

The Phttippt'neJournnl of Sctence

1e6?

half with abundant
stout, terminatingin a slenclerspine,outerrelativelynarrow,the seta com'strong spinoids.t*. .4Edeagus
parativelyfew in number.
Habttat.-India ( Assam)'feet, April 25.
Holotype,male,Nongirong,i\{ikir, altitude2,700
1960(Sc|lmid').Allotopotype,female,pinnedwithtype.Para.
topotypes,9 malesand females,with the types'
APOCRYPHA sp' nev'
GONOMYIA OBOTOGONOMYIA)

Platc I'

flgs' I'l3'

generalcolorSizemedium(wing of maleabout? miltimeters); with flange
black;malehypopygium
ationof bodyandappendages
slender'inter'of apicallobe of Uttittyle distinct;outer dististyle style a long'
inner
mediatestyle with terminal lobesvery short,obtuse'oval bro*nirf, yellow blade,its tip narrowly.
wing, 7 to 7.3;
Male._Lungih,about 6.5 to o.o mutimeters;
antenna,about2.
last relatively long'
Rostrum,p"tpi and antenna black, theHeadbrowniihblack,sparselypruinose'
T h o r u r d u l l b l a c k , s p a r s- e l y p r u i n o s e , t h e p rLegs
e s c u black'talstripes
large'
knobs
ntact,
Halteresvaguely indicated.
long narrowstigma
wings (Platei, tig. 1) faintty clarken"d,il:delicate' Venation: Scr
darker brown; veins brownish black.
e n d i n g a l m o s t o p p o s i t e t h e f o r k o f R s ' ' S c r s ucellb e qR:r;u a cellltoR2"'31a;widening
outwardly,
decurvedR+uuincell Ra deep,
2ndMlaboutfourtimesitspetiole;m-cuclosetoforkofM.
A b d o m e n , i n c l u d i n g h y p o p y g i u m , b l a c k , . t hhypopygium
ecentralpartsof
Malebrown'.inn.*ottmore
the segmentsvaguelyb, stout, with a(plate r, tig. rz) *itt apical lobe of basistyle,the apex
semiovalflange.outer dististyle,d, slender'
conspicuousintermediatestyle with
only stightlywider than the stem,obtuse;
setiferousvirtually to the base'apex very
surfaceconspicuously
shortened'inner blade
shallowlybifid, the lobescorrespondingly lateral blade rvith
point,
a litile u.oro.r,- eit.noed into a srrorf
apexobtusetonearlytruncate;inner'tyJualong.ovalbrownish
with relativelyfe.,yellow blade,tip nariowly obtuse,outer halfdelicatesete.
Habitat.-India ( Assam)'Frontier Agency,alHolotype,n,.1", h,upr,Kameng,Northeast
2 males'Paratopotypes'
titude 5,200feet,May2, 1961 6inmm)'
other similar regional
The presentfly is quite distinct from

e6,1

Alerander: Tipulda, from Eastern Asia, LXil

33

speciesin hypopygialcharacters,especiallythe dististyles. The
megalomata
most similar speciesis Gonomyrn(.Protogonomyia)
Alexander,which differs in the apicallobe of the basistyleand in
all three dististyles.
GONOII1YIA (PROTOGONOMYIA) CLISTA Alexander.

Plate I, figs. 2, 13.

Gonotnllia(Protogonom.gia)
clista Ar,uxaxpnn,Jour. N. Y. Ent. Soc.
70 (1962)10-11.

Type-male, Dikchu, Sikkim, altitude 2,300feet, May 9, 1959.
Venation(Plate1, fig.2);male hypopygium(Plate1, fig. 13).
GONOMYIA (PROTOGONOMYIA) CONTOSTYLA sp. nov.

Plate 2, fig. 14.

Size medium (wing of male about 6.5 millimeters); general
black; male hypopygiumwith
eolorationof body and appendages
apical lobe of basistylevery obtuse,without a flange; outer dististyle very small,the length oniy aboutthree times the diameter;sete of inner style spinoid.
Male.-Length, about 6.5 to 6.7 millimeters;wing, 6.5 to 6,8;
antenna,about 1.5 to 1.6.
Female.-Length,about 7 to 7.5 millimeters;wing, 6.8 to 7.5.
Rostrum palpi and antenne black; flagellansegmentselongate, the more proximal orressllorter than their longestverticils.
Hea{ black.
Thorax entirely dull blac:krvith the exceptionof a small red'
dened area on the anterior pretergites;surface more or lesspruinose,especiallyon pleura and the posteriormesonotum.Hal'
teresbrolvnishblack,extremebaseof stemreddened. Legswith
coxa gray pruinose;remainderof legs black. Wings moderately darkened,the very long and narrow stigmal area darker; veinsdark brown,trichia btack. Venation:Sc long, Sc, endingnearly
oppositefork of Rs, Scr alonefrom about one-haUto nearly twothirds as long as Rs; Rz+r+r nearlythree timesthe basalsectionof R.',icell 2nd M: aboutfour timesits petiole;m-cuat fork of M.
Abdomen, including hypopygium, black. Male hypopygium(Plate 2, fig. L4) with apicallobe of basistyle,b, very obtuse,
without a flange, setrelong. Dististyles,d, with outer style very
short, not extendingbeyondapexof basistyle,the length of stem
only aboutthree to four times the diameter,tip obtuse;interme'
diatestylelongest,terminatingin a long spine,the subapicalblade

The Phi,h,pprne
Jourrwl of Scienne

34

triangular, with a small setuliferouslobe on inner margin; innerstyle simple, sausage-shaped,
the obtusetip narrow, outer thfud
with relatively few but very long spinoid set€, the more basal
onesretrorse. .4,deagus
relatively narrow, the sete numerous,
very small and delicate.
Habitat.-India (Assam) .
Holotype, male, Shilliang, Myntang, Khasi-Jaintia,altitude3,500 feet, April 2L, 1960 (Sch,mtd).Allotopotype,female,pinnedwith a paratype. Paratopotypes,
1 male,2 females;paratypes, 2 females,Thangram,Khasi-Jaintia,altitude 3,000 feet,
April 22, 1960 (Schmid).
Gonurnyin(Protogonomytn)contostylais ,readilytotd by the
hypopygialstructure, especiallythe unusually small outer dististyle. Other regionalmembersof the subgenusthat have conspieuousspinoidsete on the inner dististyleincludeG. (P.) orus'
tglnta Alexander,G. (P.) apicul,ata
sp. nov., G. (P.) pakistanira
Alexander,and G. (P.) pellar sp. nov. Specieshavingsuchseta
Alexander,G. (P.)
small and delicateincludeG. (P.) aLomnta,
megalomnta
Alexander,G. (P,\ nigripes (Brunetti), and several
others.
GONOMYTAOROTOGONOMYIA)DTSTITURCAAlexandcr. Plete l, llg. 3; Pletc 2, l$.15.

Jour. N.Y. Ent'
Gonotngin (ProtogonomAta) di,sti'furco ALSXAI.IpER,
Soc. 70 (1962) 13.

Type-male, Dikchu,Sikkim, altitude2,300feet, May 9, 1959.
Venation(Plate1, fig. 3); malehypopygium(Plate2' fig. 15).
coNoMyIA(pRoTocoNoMYH)

GLABRISTYLA sp. nov. Plate lr ltg. 4; Plrte 2, fll. 16.

black;wingstinged
Generalcolorationof body and appendages
with blackish,cells R, and 2nd Mz deep; male hypopygtumwith
apical lobe of basistyleshort and stout, without a flange; inter'
nrediatedististytewith baseshort, the longer arms slender,glabrous,inner style with outer half a long-ovaldusky blade.
Male.-Length, aboutB to 8.5 millimeters;wing, 8.5 to 9.5; an'tenna,about2.3 to 2.5.
Femnte.-Length,about7.5 to I millimeters;wing, 8.8.to 9.
Rostrum black, relatively long, with the labial palpi subequal
to remainderof head;palpi black,terminalsegmentequalto thepenultimate, slender. Antenne blaek; proximal flagellar seg'ments oval, the fourth and fifth cylindrical,terminal segment
slightly longer than the penultimate. Headblack.

e6,1

Alerander: Tlpulidp from Eastern As?n,LXn

35

Thorax,halteresand legs dull black. Wings (Plate L, fig. 4)
tingedwith blackish,the eostalborder and narrow stigmaslightly
darker; veins brownish black. Venation: Sc long, Sct ending
shortly before fork of Rs; ceIl R:r deep, veins Rs and R+ only
slightly divergent,cells Rg and R.qsubequalin extent at margin;
cell 2nd M2 cleep,aboutthree timesits petiole;m-cucloseto forkof M.
Abdomen black. Male hypopygium ( Plate 2, fig. 16) wlth
apicallobe of basistyle,b, short and stout, without a flange,thesete nearly as long as the lobe. Outer dististyle,d, d slendercurved pale rod from a slightly expanded base; intermediatestyle longest,the basevery short.the two unequalarms appressed
to one another,longer than the base,especiallythe stoutenaxialarm, its tip obtuse;lateralarm abouttwo-thirdsas long, very slender, both arms glabrous,shiny, the relatively few short seta of
the style'restrictedto the stembeforethe fork; inner styleyellow,
its outer haif a long-ovaldusky blade with relatively few sete.
,lEdeagus
with abundantdelieatesete.
Habi,tat.-India(Assam) .
Holotype,male,Jhum La, Kameng,NortheastFrontier Ageney,
altitude 7,800feet, June 2, L96t (Sch.mid).Allotype,female,Talung Dzong,I(ameng,altitude ?,800feet, June 6, 1961. Paratopotypes,7 of.both sexes,with the allotype.
glabristELa
is told from the numerGonomyia(ProtogonomAia)
ous regional allies by the hypopygial structure, particularly theintermediatedististylewith its long slenderglabrousbranches.
GONOIYIYIAeROTOGONOMYIA) KHASIANA Alexandor.

Plate Z, llt.

17.

Jour. N. Y. Ent.
Gonomai,a (Protogonomaia) khasiona Ar,exAIYpER,
Soe.70 (1962) 14.

Type-male, Cherrapunji,KhasiHills, August 1935 (S. Sircor).
NIalehypopygium ( Plate 2, fig. L7) .
GONOMYIA (PROTOGONOMYIA) LATEROMACULA

sp. nov. Plate l,flg.

5i Platc 2,flg. lt.

Mesonotumbrownish gray, median prescutal stripe blackgned
anteriorly;pleura brown,variegatedwith yellow and with a large
intenselyblackenedspot on anepisternum;wings weakly tinged
with brown, cell R:rdeep, veins R.r and Rr diverging very gradually; male hypopygiumrvith the outer dististylea broad-based
spine.

36

The Phrli,pprneJourrtal of Science

Male.-Length,about5.5 to 6 millimeters;wing,6 to 6.5; antenna,about1.4 to 1.6.
Female.-Length,about5.5 millimeters;wing, 6.2 to 6.3.
Rostrumbrown; palpi black. Antenna brorvn;flagellar segmentssubequalto their verticles.the outeroneslong anclslender.
Headbrown, anteriorvertex and orbits light gray pruinose;anterior vertex elevated.
honotum yellow, variegatedwith dark brown. Mesonotum
brownishgray, prreseutumwith humeral and lateral regionsyellowed, medianstripe faintly indicated,widenedand blackenecl
anteriorly; posteriorscleritiesof notum brown, seutellumslightly moreyellowed,postnotummorepruinose. Pleurabrown,ventral pleurites paler brown, anepisternumwith a large intensely
blackenedarea,beneathit with a vagueyellowedstripe extending from propleurato metapleura. Halteresbrown. Legs withyellow; remainderof legs obcoxe and trochanterstestaceous
scureyellow,outer tarsalsegmentsslightly darker. Wings (Plate
1, fig. 5) weekly tinged with brown, stigmascareelydifferen'
tiated; veinsbrown. Venation:Scr endingabout oppositeforkof Rs,Sezretracted,Sctexceeding
Rl+l+q veinsR,qandRr diverging very gtadually;cell Rslong, vein R+aboutfour times Rz+r+1,
Rsabouttwo andone-halftimesthis vein; cell M2openby atrophyof basalsectionof Mo;m-cuat or closeto fork of M.
Abdominaltergitesandhypopygiumdark brown,basalsternites
obseureyellow, outer segmentsmore darkened. Male hypopl'grum (Plate2, fig. 18) with apicallobeof basistyle,b, short,without flange,apexobtuse. Outer dististyle,d, a simplepale rod,broadat base,narrowedgraduallyinto a needieiikespine;inter'mediatestyleunequallybifid at apex,axial arm nearlythree times
the lateralbranch;inner styleoutrvardlygraduallydilatedinto an
obtusehead.
Habi,tat.-Thailand.
Holotype,male, Chiengmai,near Dr. Buker's Cabin,Febntary
female,pinnedrvith
24, L953(DeedC. Thu:rmnn). Allotopotype,
6 of both sexes,rvith the type (Thtrmaitt)rye. Paratopotypes,
and Manop').
lateromnculais most similar to G.
Gonomgia(Protogonomgi,a)(P.) wmtokensi,sAlexander,differingevidentlyin the colorationin hypopygialeharacters.
inof the thoracicpleuraand especiallycluding the outer dististyle,e6'1

Alerander:Ttpulida from EasternAsia,LXn

GoNoIltYIa (PRorocoNolurrA)

MEGALOMATA Alexandcr.

g7

plar. z, trg. lt.

Gonomyia (Protogonomuin)megalomatoAr.nxeNDER,
Jour. N. Y. Ent.
Soc.70 (1962)14-15.

Type-male, Dikchu, Sikkim, altitude 2,300feet, May g, lgbg.
I\{ale hypopygium (Plate 2, fig. 19).
GONOIIYIA (PROTOGONOMYIA) PELLAX rp. nov.

plate Z, flg. 20.

Sizerelativelylarge (wing of male to 7 mitlimeters);bodyand
appendagesblack; male hypopygiumwith outer dististyle small,
slender,at apex dilated and twisted to appearfoot-shaped;intermediate style very large, inner style at apex with a concentration of long yellow spinoid sete; edeagus with sete relatively
sparseand weak.
Male.-Length, about 6.b millimeters,wing, 6.2 to Z; antenna,
about1.7to 1.8.
Femnle.-Length, about G to G.bmillimeters;wing, ? to B.
Generalcolorationof the entire body dull black, sparselypruinose to appear dark gray. Mouthparts,palpi, antenne and legs
uniformly black. lvings faintly darkened, stigma very narrow,brown; veins brownishblack. Venation: Scr ending shorily before level of fork of Rs, scs moderatelyremoved,sc, alone fromtwo-fifths to one-halflis; position of m-cu variable,from shorily
beforethe fork of M to about one-halfits length beyondthis fork
(in the holotypeand allotype).
Male hypopygium ( Plate 2, fig. 20) with apex of lobe of basistyle broadlyobtuse,with an ill-definedinner flange. Outer dististyle, d, small, slender,distal end slightly dilated and twisted toappear more or less foot-shaped;intermediatestyle very large
and conspicuous,nearly as long as the basistyle,the outer half
expandedand forking into a gently curved spinelikeblade and amore triangular expandedinner section,the usual sete of thestem and baseof blade abundantbut unusuallysmall and weak;
inner style a very gently arcuatedsimple arm that is longer thanthe outer style, with a concentration of about lz long yellow
spinoid sete at and close to apex. ,4Edeagus
small, only mod.
erately expanded,the sete relatively few and weak.
Habitat.-India ( Assam) .
Holotype, male, Nongjni, Khasi-Jaintia,altitude B,?b0 feet,
April 19, 1960 (sclmtid). Allotopotype,female, pinned wiflr
type. Paratypes, 1 male, Jarain, Khasi-Jaintia, altitude 2,900

38

Jornnal,of Scienre
The Phi,Li,ppr,ne1967

feet, April 13, 1960; 1 female, Umlangshor,Khasi-Jaintia,April
18, 1960.
Other regionalmembersof the subgenushaving eonspicuous
spinoid setre on the inner dististyle of the hypopygiuminclude
GonomEia(Protogonomyia\acustElataAlexander,G. ( P.) apiculatasp. nov.,G. (P.) contostylosp. nov., G. (P.) reiecta sp.
nov., and some others, all readily distinguishedamong theutselvesby the structure of the three dististyles.
GONOMYIA@ROTOGONOMYIA) REJECTA sp. nov.

Plate 2, fis. ll.

Generalcolorationblack,pruinose;antenna,halteresand legs
black; male hypopygiumwith both the inner and outer dististyles
appearingas narrow flattenedblades,at near three-fourthsthe
length with a compactgroup of long spearlikespinoid sete thatare directed basad.
Mate.-I*ngth, about 6 millimeters;wing, 6.5; antenua,about
2.L.
Femnle.-Length,about 6 millimeters;wiug, 6.5.
Rostrum and palpi black. Antenna black; flagellar segments
oval, the outer onesmore elongate. Head dull brownishblack,on the gene.
pruinose,,esPeciallY
Pronotum dull black, scutellum obscureyellow. Mesottotum
black, surfacesubnitidousby a sparsepruinosity,especiallylaterally; humeralregion of prescutum,posteriorcallositiesof scutal lobesand posteriorborder of scutellumobscurebrownishyellow, lessevidentin the female. Pleuradull black,sparselypntinose. Halteresbrownish black, base of stem restrictedlyyellowed. Legsblack. Wingsweaklydarkened,stigmalinear,palebrown;veinsdark brown,macrotrichiablack. Venation:Scrending shorily before fork of Rs; m-cu closeto fork of M.
Abdomen,includingthe genitaliaof both sexes,black. llale
hypopygium(Ptate 2, frg. 2I) with apex of basistyle,b, broad,atmoit tnrncate,the inner anglewith indicationsof a lorv flange;
face of basistylewith an oblique sclerotizedridge extendingto
baseof the apicatemargination. Outerdististyle,d, & slenderyel'
low blade,gently curved,basegreatlydilated,apexobtuse;intermediatestyleblackened,apexbifid, the outer arm a slenderspinethat lies acrossthe face of the expandedinner blade;inner dististyle distinctive,appearingas a flattenedblaclethat is subequalin lengthto the outer style and a little broader,the outer margin
at near three-fourthsthe length with a concentrationof about

e6,1

Alerander: Tipulida frorn Eastern Asin,LXngg

a dozen long spearlike spinoid sete in a compact group that is
directed basad. ,fEdeagus
narrow, with ,relatively sparse seta.
Habitat.-India (Assam) .
Holotype,male, Mynso,Khasi-Jaintia,altitude 9,b00feet, April
20, 1960 (Schmid). Allotopotype,female,pinned with type.
Gonomyia (Protogonomgia)rejecta is a very distinct fly thatis readily told from other speciesby the armature of the innerdististyle of the hypopygium. other regional forms that havemodified spinoid sete on this style include G. (p.) aatstglata
Alexander,G. (P.) apiculatasp. nov., G. (p.) coniostEta
sp. nov.,
G. (P.) pakistani,ca
Alexander,G. (p.) pellnr sp.nov., and someothers, all with distinctive hypopygia.
GONOMYIA OROTOG0NOMYIA) SCOTEINA sp. nov.

Plate l, ng. 6t platc t, 1i1s,
zz.

size large (wing of male over 8.b millimeters); generalcoloration of body and appendagesdull btack; wings with ceu Rg deep,
the veins only slightly divergent; male hypopygium with apicallobe of basistyle short and stout, without a flange; outer distrstyle narrowed to the subacutetip, intermediate style with theouter bladessubequalin length.
Male.-Length, about 10 millinreters;wing, B.T;antenna,about2.
Rostmm slightly produced,about one-third the remainder ofhead, black, pruinose;palpi black. Antennreblack throughout;proximal flagellar segmentsoval, the outer ones more elongate,
subequalto their verticils. Head black, gray pruinose.
Thorax dull black, the pretergites light brown; pseudosutural
fovee black; scutellum, mediotergite and pleura slighfly morepruinose; membrane of the meral region restrictedly yellowed.
Halteres and legs black. wings (plate 1, fig. 6) tinged withblackish,the prearcular and costalfields a trifle more saturated;stigma long and narrow, darker brown, virtually restricted to the
costal cell; veins dark brown. Venation: Sc long, Sc, ending
shortly before fork of Rs; cell Rs deep, the veins only slighfly
divergent, cell Rr at margin about one-half more extensivethancell Rz; anterior branch of Rs more than three times R213.r.a3c€ll
2nd Mz deep, more than three times its petiole.
Abdomen black throughout. Male hypopygium ( ptate B, fig.
22) with apical lobe of basistyle,b, short and stout, without aflange but with abundantlong setre. outer dististyle,d, broadbased,,gradually narrowed outwardly to the subacutetip; inter-

40

The Philipp'meJournal of Scterrce

1967

mediatestyle with the distatbladessubequalin length,the iuuer
more pointed,margin of outer blade with rnoderatelylong seta;
inner dististyleexpandedon outer half.
Habr,tat.-India (Assam) .
Holotype,male, Lifakpo, Kameng,NortheastFrontier Agettcy,
attitude3,100feet, May 15, 1961 (Schnu'd).
scotehwis one of the largestlnemGonomyin(Protogonomytn)
bersof the subgenus. It is mostsimilarto speciessuchas G. (P.)
alomnta Alexander and G. ( P.) nigripes ( Brunetti) , differingfrom theseand othe'rsin the detailsof venationand hypopygial
structure.
GONOMYTA (PROTOGONOMYIA)TIIIORHOPALA Alexander.

Plate I' fis. 7;?,|;:"ri.,Gonomgrn (ProtogonontAin) thi,orhopalo AlnxaporR, Jout. N. Y. Ent.
Soc. 70 (1962) 15-16.

Type-male, Singbeng,Sikkim, April 26, 1959 (Schm'id).
Venation(Plate1, fig. ?); rnalehypopygium(Plate3' fig. 23;.
SubgenusSIVAGONOI,IYIAnovum

Charactersgenerallyas in Progonomyiaand Ramagonomynin
the retention of vein Rg of the wings, with other featuresas inProtogonomgiaand Etti,pterordes.Ovipositss\a/ithcerci long andslender, sclerotized. NIaIehypopygium (Plate 3, fig. 24) witlt
the phallosomedistinctive,appearingas a broadcentralstructure,
the outer third divideclinto two slender spinesthat are directedcaudadand lie generallyparallel to one another.
Tlpe of subgenus: Gonomyta (Si,uagonomgin)discoloplnll'os
sp. nov. Oriental. The only speciespresentlyknorvn. The sub'generic name is in allusion to the two-spinedredeagus,which is
unique in the genus.
GONOMYIA (SMGONOIUYIA)DISCOLOPHALLOS sp. nov. Plate I, fig. 9; Platc 3, lig. A,

pat'
Generalcolorationof thorax polishedblack,,conspicuously
terenedwith yellow;headlight gray; halteresyellorv;legsyellorv,
tips of femoraand tibie broadlyblack; wings yellow,restrictedly
patternedwith pale brown; male hypopygiumwith the charac'
under the subgenus,the apex of the adeagus
ters as desc,ribedbeing divided into two long paraliel spines.
Mal,e.-Length,about5 millimeters;wiug, 6; antcttita,abgut2.
Female.-Length, about I millimeters; wing, 7.

e6'I

Alerand"er:Tipulida from Eastern Asia,LXU

4l

Rostrumbrown; palpi brownishblack. Antenne of male rela.
tively long, approximatelyone-thirdthe wing; scapeand pedicel
brownish yellow, flagellum black, bases of the more proximal
segmentsnarrowly yellowed; verticils slightly longer than thesegments. Head light gray; anterior vertex broad.
Pronotal scutum light yellow, the sides black, scutellum concolorous. Mesonotalprescutum with disk polished black, humeral region and a small medianspot at suture yellow, the latterbeing confluent with a larger area at rnidregion of scutum, thelobes of the latter polished black; scutellum light yellow, baseblackened,parascutellaand postnotumblack. pleura intensely
black on propleura,anepisternum,ventral sternopleuriteand dor,sal pteropleurite;dorsopleurarregion and a conspicuousventral
longitudinalstripe light yetlow; the latter widenedbehind to include the meron. Halteresyellow.knob paler yellow. Legswith
coxre obscureyellow, basesblackenecl,broadly so on forl pair,very narrow on hind eoxE, trochantersyellow; femora light yellow basally,tips broadly black, on fore pair including the outer
two-thi,rds,on the remaining legs the black tips narrower,
involving the outer fifth to sixth; tibia yellow, tips broadly
black, broadeston fore pair, including the outer thir-d,extrenre
base similarly clarkened,tarsi biack. vestiture of legs normal,
inconspicuous.wings (plate 1, fig. g) yellow, the prearcularand costal fields clear light yellorv, ilie extreme base vaguelydarkened; a narrow pale brown seam includes the siigmi
and cord, narrowly interrupted at M31a, veins brownish yellow,
darker alongcord. venation: Sc long, Scr ending about opposite
three'fourthsRs; R2 present,subequalto R1*2,cell Mz open byatrophyof basalsectionof M*; m-cu aboutits own tength ueyono
the fork of M.
Abdomen dull black, ninth segrnentobscureyellow, the styli
black. ovipositor with cerci long and slender,selerotized,gengy
upcurvedto the acute tips. Male hypopygium(plate s, tig. z+jyith the tergite, t, transverse, narrowed outwardly, posteriorborder truncate, Basistyre,b, unusually compact,iuboval, theouter lateral angle produceclinto a stout spine that is directed
inwardly. Three dististyles.d, all simple: outer style a slender
blackenedrocl. the short terminal point reeu,rved,ih. expandecl
baseproducedinto a still smaller prlint with two strong ipinoid
seta; intermediatestyle stoutest,the basebroad,bearinga single
strong seta on its lateral extension,the outer head blacfened,fts

42

T'hePhi,ltpptneJaurnnl of Scienne

shapeas figtrred; inner style a narrow yellow blade,the tip obtuse. PhallosoffiQ,P, distinctive,edeagusa broad centralyellow
structure,the outer third dividedinto two slenderspinesthat lie
generally parallel to one another, their tips slightly incurved;
body of adeagusbefore the fork with three pairs of long sete.
Habi,tat.-India ( Assam) .
Holotype,male,Rupa,Kameng,NortheastFrontier Agency,al'feet, May 2, 1961 (Schmid). Allotopotype,female,titude S,ZOO
1 male,I female.
pinnedwith type. Paratopotypes,
fire presentfly requires comparisonwith no other generally
polishedbody
similar Eriopterinecrane flies. The conspicuous
pattern and leg .s1s1ationseparateit from other apparently
ielated speciesin the subgeneraProgonomEtn.The genusDcsgthairy, is more dismallomgioBtunetti, with the legsconspicuously
tantly related.
SubgenusRAMAGONOMYIAnovum

in the retentionof the
Charactersgenerallyas in Progonomyi,avein Rz,differing in other featuresof venationand especiallyin
hypopygialstruciure. Wings (Plate 1, fig. B) with the anterior
rt.uf*- lacking or only vaguely preserved. Male hypopygium(Plate3, figs. 25,26) with the apexof basistyleproducedinto aera) or slenderspine (protensa). Two
ftattenedptate (bistcul,if
dististytes,ttte inner one complex,as figured. Adeagusextend'
ed dir-eclycaudadinto a point, not bent laterador recurvedasin
ot ProtogonomYia.
Progonomgia
protensoAlexType of subgenus:Gonomyia(Ro':ntagonomyia)
anOei. Oriental.G. (R.) bisiculi,feraLlexanderlikewisebelongs
here,both speciespresentlybeingknownonly from Sikkim. The
male hypopygiumof protensais shown ( Ptate 3, fig. 26); btlsi'
culi,fera(Ptate 3, fig. 25).
are the only
and Ramagonomgrn
Progonomyia,Sioagonomyi,amembersof the genusthat havevein R, ( r of someauthors) preserved,the other closeallies,Ellipterotdesand Protogonomyia,
is representedby
having this elementatrophied. Progonom,Et'anumerousspeciesin the Neotropicaland southe'rnNearctic re'gions [Alexinder,C. P., Rev. Entomol.18 (t947) 83-84]. A
imalt number of further speciesin SouthAfrica that have been
may not belonghere, as has beensug'
referredto Progonomgia
gestedby Edwards;Trans.Soc.BritishEnt.,Part 1 5 (19381106i
ind may be found to pertain to the presentgroup. I lack ade-e6,1

Alerander: Tipulida from Eastern Asi,a,LXfi

43quate materials of the African speciesto decide this question[Alexander,C. P., South African Animal Life, Lund University
Expedition 1950-1951. Diptera (Nematocera) Tipuiide t0(1964) 381-3871. SuchAfrican specieshaveboth the anterior
arculusand vein Rgdistinctly preserved.
GONOIYIYIA (RAMAGONOMYIA)

BISICULTFERA Alexander.

Plate 3r lls,. 25.

GonomAi.a(Progonomgia) bisiculifeto Ar,nxlwunn, Jour. N. Y. Ent.
$sg. 7l (1963) 73-74.

Type-male, Yugang, Sikkim, altitude 5,700 feet, May L4,
1959.
Male hypopygium( Plate 3, fig. 25) .
GONOIvIYIA (RAMAGONOMYIdPROTENSA Alexander.

Plate l, ftg. Ei Plate t, flg. 26.

Gonomgi,a (Progonomgw) protensa ALsxANpEr, Jour. N. Y. Ent.
Soc. 7l (1963) 75.

Type-male, Manu, Sikkim, altitude 4,92Afeet, May 10, 1959.
Venation(Plate1, fig.8);male hypopygium(Plate3, fig.26).
SubgenusIDIOCERADaleGONOMYIA (IDIOCERA) ACAENOPHALLOS sp. nov.

Plate 3, tig. 27i Platc 4, llg. 3E.

Generalcolorationof mesonotumgray, patternedwith browtt;
proximalantennalsegmentsyellow;femoraobscureyellow;wings
light yellow with a restrictedbut conspicuous
brown spottedpattern; male hypopygiumwith the three dististylesof distinctive
conformation;redeagusstout, on the convexouter margin with aconspicuous
blackenedflangethat is extendedinto a strongcurved
black spine.
Male.-Length, about7.5 millimeters;wing, 6.5; antenna,about
L.4.
Rostrum dark brown; palpi black. Antenne with scape andpedicellight yellow,flagellumbrown;flagellarsegmentslong-oval,
the outer onesstill more lengthened,with conspicuous
verticles.
Head brownishgray.
Pronotumbrorvnishgray, yellowedlaterally. Mesonotalprescutumgray,the lateralbordersvaguelymore yellowed,disk rvithvagueindicationsof two diffuse brown stripes on posterior half,
the lateral pair barely irrdicated,pseudosuturalfovea brownish
black; scutum gray, lobes vaguely more darkened; scutellum
brownish gray, posterior border more yellowed; mediotergite
brownishgray, the lateral bordersand pleurotergiteobscureyel-

44

Jownal of Sc;ence
The Philr,pphte

1967low. Pleura light brown, posterior scleritesyellow. Haiteresoutrvardlylight yellow,
with basalhalf of stemweaklyinfuseated,
includingthe knob. Legswith fore coxe dark brown;trochant'
ers brownishyellow,remainingcox€ and trochantersyellow; fe'mora obscureyeltow; tibire obscureyellow, tips very narrowly
darkened;basitarsiyellowed,tips andremainderof tarsi brorvnish
black. Wings (Ptate3, fig. 27) light yellowwith a restrictedbut
brownspottedpattern,the stigmalargest,oval;small
conspicuous
isolatedbrown spotsat origin of Rs, cord,fork of Mr+2, arculus,
and at tips of all outer longitudinalveins exceptingSc and R;;
dark brorvnin the patternedparts.
veins yellow, conspicuously
Venation:Sc short, Scr ending about oppositeone-fifthto onebut notsixthRs;tips of veinsRr+eandR,,at marginapproximatecl
contiguous;eell R. large,cell 2nd [I, aboutone'halflongerthan
its petiole;m-cu aboutone-halftimes its length beforethe forkof M.
Abdomendark brown, the extremeposteriorbordersof themoreproximaltergitesvagueiypaler,of the sternitesmorebroadly so; hypopygiumbrorvn. i\tale hypopygium(Plate 4, fig. 38)b, paie,stout,tip obtuse,the innerwith outer lobe of basisty'ie,,blackenedpoint. Three
apicalangleproducedinto a microscopicd, the outer stylewith stemslender,bifid at apex,thedististytes,
longerslenderblacieproducedinto a point in the axil near base;styie a sol]iet'rliatshorter and stouterrod, at apex
intermecliate
into a longblackspine, with a smallersimilarspineat its
extencled
the two spinesforming a straightbasethat is directedbackward,,
angle;inner style subequalin length and cliameterto the last, at
its tip with a slightly recurvedblack spine, baservith severalp, unique; adeaguslong and conspistrong sete. Phaliosome,cuous,appearingasa stoutgentlycurvedrod, on the convexuppersurfaceat near midlengthbearinga blackenedflange that is extendedinto strong curvedblack spine,the flange more basally
serrulate;apex of edeagusbewith the margin microscopicaliyneathwith two smallacutepointsanda singlevery long paleseta.
Habitat.-India (Assam) .
Kameng,Northeasttr\'ontier'
Holotype,male,,Bhairabkuncla,
Agency,altitude700to 1,000feet, MarchB, 1961(Sclmtid).
(ldiocera)ac0,enophal'l.os
is entirelydifferentfrom all
Gonomyi,ain the hypopygialof the subgenus
otherknownmembers
structure,especiallythe somewhatremarkableedeagus,as described,and
which has suggestedthe specificname.

ee'I

Alexarder: Ti,puli.d,afrom Eastem Asia,LXn

coNoMYIA(IDIOCERA) ACTFURCA Atexander.

45

plate 3, fte. 29i prate 4, fis. 4r.

Gonomai,a(rdiocera) aci,fu,rcaALexaNosn,Ann. Mag. Nat. Hist. (12)7 Gss+)Bee-901.
The types were from Nuristan,East Afghanistan.other recordsincludePakistan,in the NorthwestFrontier province,Le
Vallee,July 28, lgb3 (Schmid,)and Salf-ul-Maluk
Sur, altitude
11,000feet, July 2, lgb3 (scfunicl)and Kumaon,Inclia,at Gangrea,PauriGahrwal,
aititucle10,000
feet,Junelb, lgb8 (schmid,).
venation (Plate B, fig. 2B). Male hypopygium(plate 4, fig.
39) with the basistyle,b, short anclcompaet,the outer apicat
tgbeexceeding
it in length,appearingas a cluskyclub with abundan seta on outer thircl. Three dististyles,d,; the outer withstem slender,unequallybifid into an elongateouter arm and ashortstoutinner one;intermediatestylestigtrttyshorterthan thelobe of the basistyie,appearingas two long slenderblaekened
spinesfrom a short commonbase:inner styte a litile longer,asimpleslenderrod, tip acute,outermarginoi apicalfourth microscopicallyspiculose,
lower edgeof outer half with a row of longerect bristles. adeagusflattened,at apexproducedinto a slender beak,lolverpart of outer harf with numeroussetre.
GONOMYIA ODIOCERA) COHERIANA Atexander.

Plate 4, ftg. 40.

Gonomyia(Idiocera) coherianaAr,rxawnrn,Jour. N. Y. Ent. Soe.66(1958)164-165.

Type-male, Parerva.vir,
Nepal,NIareh26. lgb? (8. I. Coher).
ptate
Male hypopygiunr(
4, fig. 40) .
GONOMYIA (IDIOCERA) FORCOSA sp. nov.

Platc 3r llg. 29i Plate 4, flg. 41.

Generalcoloration of ilrorax gray, p,rascutumwith trvo brownish gray stripes, posterlor selerites of notum and the pleura^variegatedgray anclbrown; proximal antennalsegmentsyellow,
remainderbrown; legs]ellss', outer tarsal segmentsbrownj wings
yellowed,restrietedtypatternecrwith brown in the radial fieti,
with further narrotrvbrorvn seams arong cord and elsewherei
abdomenbrorvn,posteriorborclersof segmentsnarrowly yellowi
malehypopygiumwith outer lobe of basistyleelongate;three dististyles,all bifurcate, the outer style very deeply so, intermediatestyle unequally lobeci,inner style bispinous; edeagus slender,
apexobtuse.
Male.-Length, about 6 to 6.b millimeters;wing,6.8 to T; err.ttenna,about1.2to 1.3.

46

The Phi'tippineJournnl of Science

1e67

7 to B'
Fenmle.-Length,aboutB to 8.5 millimeters;wing,rvith
Rostrumbrownishgray; palpi brownishblaek. Antenne
brorvn;proximal three or fousug*.nts light yellow, remainderandpubescenee
flagellar segmentselongate,with a densewhitegray'tonler verticils. Head yellow, center of vertex light
light yelcervical region dark brown. Pronotum chiefly clear
brorvnishlow. Mesonotalprascutum with disk graY, with two
anctgray intermediatestripes that ils widenedbehind,'humeral
notum brownish
lateralborderslight yettow;posteriorscleritesof
bordergray, scutallobespatterneclr,vlthdarker brown, posterior
laterally rvithof scutellum dull orange, mediotergite borderedpale trown, eS.
yellow, pleurotergiteright yellow variegatedby
above gray'pecially^on the ventral katapleurotergite' Pleuraa broad lightvariegatedwith brown, ventrally pale-brownwith
eoxee'dorsal
yellow longitudinal stripe that incluclesthe fore
and metapleura'sternopleurite, ventral pteropleurite. meron
with eoxE-yellorv'
Haltereswith stem yellow, t<nol brorvn. Legs
brorvn;trochantersbasesof middl. ,nO hind pairs narrowly pale
brorvnyellow;remainderof legs yellow,terminal tarsal segments
plate g, fig 29) yellowed,clearer
to brownish black. wings (dark brorvn; ayellow in the prearcular and costal fielcls; stigma
narrowly inpaler brown ,r.. in outer half of cell R* and more
cord and outercell Ri',inarrow brown seamsat origin of Rs,
da'rkenedareasat
medialfork; veinsyellow,brownishblackin the
oppositemidcord and fork of fu. Venation: Scr enclingabout
to nearly
length of Rs; veins R r+z and Rg strongly approximated.onligoous at margin, virtually closingcell Rr'
pruinose,PosAbdominaltergites dark brown, very sparsely
paler brown rvith
terior borders nairowty light yellow; sternities
obscureyellow'
still narrower yellow outer mirgins; hypopygium
of basistyle'Male hypopygium(Plate 4, fig. 4l) with outer lobe
dististyles'b, very long, aboutthree-fourttrithe remainder' Threedeeplyd, all forked; outer style with basestout,before midlengththat narrows
bifurcate,the outer "i* u blackenedribbonlikebladea trifle short'u.ry graduailyinto a long terminal spine,inner arm
narrorvs"r., ,pp"aring as a slendergently curved black spinethatinvery^graOuaityto the acuiu trp; intermediatestyle subequal
rod that
length,very unequaltybifurcate,the outer arm a slendervery
is n'early as long as the base,tip weakly acute, inner arm
point; inner
short and broaO,tip emarginate,the angle a small
apstyle shortest,appeiring chelate,baseshorter,arlns suhequal'e6,1

Alerander: Tipulida from Eastem Asia,LXU

47

pearing as blaek spines,the mesal one lvith about six scatteredlong setre. Phallosoffi€,p, with redeagusslender,slightly moreconstrictedbeforethe obtusecuplike apex.
Habitat.-India ( Assam) .
Holotype,male,Shergaon,Kameng,NortheastFrontier Agency,
altitude 6,400feet, May B, 1961 (Schmi.d). Allotopotype,female.
Paratopotypes,
5 of both sexes,with the types;paratypes,2 males,
2 females,Bilo La, Kam€Dg,altitude 5,.800feet, June 9, 1961.
Other regionalmembersof the subgenushaving the wings patterned and with the inner dististyleof the male hypopygiumbispinousinclude Gonomyia(Idiocera) paleumnAlexander,G. (I.)
kashongensts
sp. nov., G. (I.) mngra Alexander,G. (I.) maharajah Alexander,G. (I.) terribilis sp. nov., and some others, alldiffering among themselvesin hypopygialcharacters.
GONOIYIYIA (IDIOCERA) KASHONGENSIS !p. Dov.

Plrte 3r flg. 30i Flate {, ng 42,

Sizemedium ( wing of male to 6 millimeters) ; mesonotumgray,preseutumwith two brown intermediatestripes;basalsegmentsof
antenne yellow; femora and tibia brownish yellow; wings yellow,
patternedwith brown, including the costal borderconspicuously
and most of cells Rs and Rr; male hypopygiumwith the outer dististyle pale, irregular in outline; intermediate style profoundly
bifid, the arms unequal,very long, tips pointed;inner style bifid,
the inner arm a long slenderspine.
Male.-Length, about 5 to 5.5 millimeters; wing, 5.5 to 6; antenna,about 1.2 to 1.3.
Female.-Length, about 6 to 6.5 millimeters;wing,,6.5 to ?.
Rostrum brownish black; palpi black. Antenne with proximal
three or four segmentslight yellow, succeedingsegmentsblack,oval, shorter than their verticils. Head light yellow, center ofposteriorvertex gray, posteriorgen€ brown.
Pronotumlight yellow, with a narrow brown central area. Mesonotal prrescutumlight gray with two narrow brown intermediate stripes, paler to virtually obsoleteon anterior half, lateral
stripesbarely indicated;scutumgray, eaehlobe with a darkened
center, posterior callositiesyellow, central area obscureorange
with a medianbrown line that extendscaudadonto the otherwise
yellowish orange scutellum; mediotergitegrey, lateral margins
light yellow, pleurotergiteyellow above,patterned with brown,
ventrally with a circular pale spot. Pleura conspicuouslypatterned, ineluding a broad pale yellow ventral stripe that is bor-

48

Jownnt of Science
The Ph,i,tr,ppine

1e6?

light
dered aboveand below by brorvn; dorsopleuralmembranewithyellow. Halteresclarkbrown. baseof stem yellowed'. LgSs
coxa yellow, anterior faces of fore and hind pairs infuscated;yellow'
troehantersbrownishyellow; femora and tibie brownish
Wings
basitarsibrownish yuilow, outer tarsal segmentsblack'(Plate 3, fig. 30) with the ground yellow, clearer basally,con'and sc, with ex'
spicuoustypatternedwith b,rown,includingcellsc
more restricttensionsinto cellsRnanclR,; stigmadarker brown;
wing mared browncloudsovercorcl,outerhediat forks andouterends
gin from vein Mr+z to beyond2nd A, including the outerparts'
of veins; veins tight yellow, dark b'rownin the patternedwith con'Longitudinalveinsbeyondgenerallevel of origin of Rs
anal veins'
spicuoustrichia, including also the outer endsof both
one-fifth
Venation:Sc relativelyshort, Scr endingaboutopposite
Rg appro'to one-third.Rs, Sc, closeto the origin; veinsRr+z andcostalessthan
ximatedat margin,tft. clistancebetweenthem on
its length be'
one-fourthRa; m-cu about one and one'ha}ftimes
fore the fork of M.
intermediate
Abdomendark brown, the posteriorborders ofhypopygiunt
tergites narrowly yellow, sternites paler brown;(Ptate 4, Iig' 42) dis'brown to dark brown. Male hypopygt
lobes,the lateraltinctive. nasistyre,b, terminatingin t*o fleshly
the length
one much shorter,the larger lobe lnore than one-half a bifid
the outerof the longestdististyle. Three oististyles,d,and a smallyellorvblade,its inner arm with a curvetlblack spine
forkinger bidentatetone; intermediatestyle longest,blackened, outertheshortly beyondbaseinto two flattlned unequalblades'
longer than thearm extendedinto a long point, about one-half
margin with ainner arm; inner style wilfr-a fleshylobe,its outerp, with edeagusunusuallyslen'
long slenderspine. Phallosome,der] straight, the apex bent at a right angle'
Habitat.-lndia (Assam) .
6'q00 feet.
Holotype,male, Sirhoi Kashong,Manipur, altitude
female,pinnedwith type'
June Z, rgOO(Scimid\. AltotopotyPet
7 of'both sexes'with the types'
Paratopotypes,
GonomEia
In the detailsof structureof the male hypopygium, 'regional
is quite distinct -f.romthe other(Id,i,ocera)kashongensis(I.\ mngraAlexspecieshavingthelnner styll bifid,.includingG.
Alex'
ander,G. (l.i maharajohAlexander,G' (I') mEriacantha
in theander, G. (I.i paleumaAlexander,and others describedpresentPaper.

96, I

Alemnder: 'fipulide frorn Eastern Asta,LXil

GONOIIIYIA (IDIOCERA| l.AllIA

sp. nov.

49

Plate 3r flg. 3li Plate 4' fig. 43.

Generalcolorationof mesonotalprascutum and seutumbrown'
ish black, pruinose; pletrra clark brolvn, striped longitudinallywith whitish; femora brownish yellow; wings brownish yellorv,
very restrictedly patterned with brown; male hypopygiumrvith
the outer otle very unequallybifid, intermediate
three ciististyles,.
into a long straightspine;adeaguswith two small
style extendecl
nearlyapicalpoints.
Male.-Length, about 5 millimeters;wings,.5.5.
llead broken. Pronotutn and pretergites yellowed, Mesonotal prrescutuuranclscutumbrownishblack, pruinose,lateral prescutal borders yellorved,pseudosuturalfovee reddish brown;
seutellumbrorvnbasaiiy,apexbroadlyyellow;mediotergitebrown,
the anterolateral angles yellow, pieurotergite yellow. Pleurawith a conspicuouswhitenedlongitudinalstripe ex'
clark brcir,vn,
tending fronr behinclthe lore coxe to baseof abdomen,widened
behind to include the posterior pieurites, dorsal pteropleurite
more yeliowed. Halteres yellow. Legs with coxe darkened
basally,apicesyellowed;trochantersyellolv; remainderof legs
tips of basitarsiand remianingtarsal segments
browuish.v*ellorv,
brorvnishblack. Wings (Plate 3, fig. 31) narrow,brownishyell0\4',baseand eostalregion cleareryellorv;stigma oval, brown;veins yellonr, the cord, including m-cu, dark brown. Longitudinal veinsbeyondgenerallevelof origin of Rs with maerotrichia,
lacking on R:,;basadof ocrd trichia lacking on M and all but thetips of the anals.Venatiorr:Sc relativelyshort, Scr endingabout
veitrRgnearlyerect,
oppositeone-fourthRs,Sc:very fdr retractecl;from Rrn.zby a distanceaboutequalto twoat margin separatecl
thirds its length; cell 2nd Mr aboutone-thirdlonger than its petiole; m-cu nearly twice.its length before fork of M.
z\bclominalsegnrentsbro$'n, their posterior borders narrowly
light yellow; hypop;rgiumbrownish yellow. Male hypopygium(Plate 4, fig.43) rvith apicallobe of basistyle,b, aboutone-halfthe length of longestdististyle. Three dististyles,dj outer styleglabrous,very unequall,vbifid, the long outer arm slender,inner
blade only about one'fifth as long; intermediatestyle longest,appearingas a stout nearly straightrod from a more expandedbase,outer fifth suddenlynarrowedinto a long aeutespine,near outerenclalmostto tip of spinewith severalvery long yellow sete; inner
st1'lesmall, slender.in length exceedingone-fourththe interrnediate str4e,with about eight sete along margin. lEdeagus,a, re-50

Journnl of Scienne
The Phi,li.pprne

lativelyshort but slencler,terminatingin two very smallrecurved
teeth, the stem with a seriesof long clelicatesetreover virtually
the whole length.
Habitat.-Inclia (WestBengal) .
Holotype,male,Git Dabling,altitude4,900to 5,100feet, September13, 1959 (Schmid).
In its hypopygialstructure,Gonomyi'a(Idt'ocera)lnmiais very
differentfrom all other regionalmembersof the typicalsubgenus,
being more as a Gonomyia(Euptilostena\moghalicaAlexander.
In this latter speciesthe dististyleand edeagusare somewhatthesame,differing in details,includingthe obtuseapexof the inter'mediatestyle.
GONOIIIYIAODIOCERA)

LEDA stt. nov.

Platc 3' tlg, ,2i Plate 5' tlt.

aa,General coloration of thorax gr&Y, the prescutum with trvo
pleurastripedbrown and yellow;legsintermediatebrown stripes,.
obscureyellow; wings pale brown with a spotteddarker browrt
pattern, the areasmarginedwith whitish hyaline;Sc long; tnale
hypopygiumwith a long basalspineon outer dististyle,inner stylesimpie;edeagusslender,apexdecurved.
Male.-Length, about 5.5 millmeters;wing, 6.
Rostrumbrownishbtack; palpi black. Antenna brorvn,seapepaler. Head grayish brown.
Pronotum brorvnishgraY, seutum laterally and the scutellum
whitened. Mesonotumbrownish gray, preseutum lvith trvo in'termediatebrownstripes,scutallobessimilarlyclarkened. Pleura
dark brown with a longitudinalyellow stripe extendingfrom thefore coxa acrossthe dorsal sternopleurite,more expandedon
posteriorsclerites. Flaltereswith stemyellow,knob dark brown.
Legswith coxe ycllow, darkeneclbasally;trochantersyellow; re'rnainclerof legs obscureyellow, outer tarsal segmentsdarker.
Wings (Plate3, fig.32) very pale brown with a spotteddarker
brown pattern, the areasmarginedwith whitish hyaline; darker
markingsincludethe stigma,arculus,origin of Rs and Se2,cord,
tip of SC1,and m; more extensivepaler brown marginalcloudsin
inouter radial field, includingtips of Rr and R5,interconnected
ternally, leaving a small ground area in cell Rr; costalcell vir'tually clear, on one wing of type rvith one or two very small spotsat nearmidlengthof eeltC; no darkeningsin eellsCu or the anals;
veinslight brown,darkerbrown in the patternedareas,more yel-

96. rAl,erander:Tipu,lida from Eastern Asia,LXilbllorvedin the costalinterspaees.Venation:Sc long, Sc, ending
abontoppositetrvo-thircls
Rs, Scsoppositeorigin of the long Rs;
vein R,,obliclue,fused apicallywith Rr+z closingcell Rr; tip ofR{ bent very strongly cephaladat three-fourthsthe length; mtt'ansverse,
at junetion rvith free tip of M, with a short spur directedbasad;m-cu abouttwice its length beforefork of M.
Abclominaltergites dark brown, sternites and hypopygium
slightly paler brou'n. Male hypopygium( plate 5, tig. 44) withouter apicalangleof basistyle,b, rounded,not produeedinto alobe as in similior. outer dististyle,d, appearingas a blackenecl
hookwith a long straightblackspineat base,outer marginof thehook in casesproducedinto a low point; intermediatestyle gla.
brous,unequallvbifid. the outer arm a long darkenedribbon,thetip acute.inner arm a short broadblaekenedsubtriangularblade,
its outer marginat nearrnicllengthproclueed
into a long spine,,theapexlessaeute;inner style a simpleslenderrod, narrowedgraduall.yinto a long gently eurvedspine,surfaeeof style with longsetre. Phallosome,p, with edeagusslencler,tip deeurved.theouter half s'ith aboutfour seattereddelieateseta.
Habitat.-Pakistan.
Holotype.male,Kaghan.NorthwestFrontier Provinee,altitude
6,688feet, June27, lgb3 (Sclmtid\.
The ntostsimilar speeiesis Gonomgia ( Idiocera)si,mitiorAlexancler,which likewisehasthe darkeneclareasof the rvingnarrowly borderedby whitish. This differs from the presentity inthewing pattern,ineluclingdarkenedareasin eell Sc anclat endofvein 2ndA, andin all tletairsof struetureof the malehypopygium.
GONOMYIA (IDIOCERA) PETILIS Atexander.

Plate 5, ftg. f5.

Gonomyia (ldiocera) pettlis Ar,nxnxosR,Philip. Jour. Sci. 86 (1957)
434-435.

Type-male, Amlekhgang,Nepal, altitude 1,690feet, May 29,
Ne1956(8. l. Coher). A secondmalervastakenat Nayagaon,
pal, October13, 1957,by Coher.
Nlalehypopygum(Plate 5, fig. 45).
GONOMYIA (IDIOCERA) SERRATISTYLA sp. noy.

Phtc 5, llg. {6,Generalcolorationof mesonotumbrown, including a central
preseutalstripe;femoraand tibie brownishyellow;wingsbrown'
ish yellow, restrictedlypatteined with ctarker;Sc short, Sc, op'
positeorigin of Rs; male hypopygiumwith margin of outer disti-

52

The Phttipinc Journal of Seience1967

style serrate,inner style simple,terminatingin a long spine'
about 5 rnillirneters;rving, 4.5; antelllla' aboulIl1ale.--Length,
0.7.
Rostrumbrorvnishblack;palpi black. Airtennrervith scapeatrci
peclicelwhitened,flagellum brownish black; flagellar segtnettts
oval, shorter than the verticils. Head gray above,front and auterior vertex whitened,gen& more infuscated.
PronotumbrownishyeilOwabove,darker on sides. Mesonotal
prescutum brownish yellow rvith a median brown stripe that is
expandedon posterior half, lateral stripes indistinct; posteriorscleritesof notum dark brown, sparselypruinose. Pleura darkbrown dorsally, broadly reddish brown beneath, dorsopleurai
membranepale. Ilalteres infuseated. l.,egswith coxe reddish
yellow, forspair more Carkened;trochairtersyellow; fenroraandiinie brownishyellow, tarsi darker brown. Wings brorvnishyel'low, cleareryellorvin the preareularand costalfields; a restricteddarker pattern includesthe pale browrrstigmaand narron'brown'
ish black seamsover the veins that comprisethe cord, origin ofveins
Rs, and tip of vein Rr, outer end of all R+weakly cla'rkened;
iight brown, darker in the patternedparts. Venation: Se short,Sc, ending opposite orig:n of Rs; distance on costa between
veins Rr+r and Rr about one-thi'rdthe latter veiu; tip of vein Rt
upcurved;cell 2nd Mg longer than its petiole,in casesnearly totwice the length; m'cu about.one ancl one-halftimes its length
beforefork of M.
Abdominal tergites dark brown, the incisurespaie. sternites
somewhatlighter brown. Male hypopygium (Plate 5, fig. 46)
u'ith apieallobe of basistyle,b, stout, tip obtusewith numerous
setee. Outer dististyle, d, ? simplerotl, eurvedat basalthird. thelong terrninalspineseruulatealonglor',,erfaee;intermeriiatestyleprofoundly bificl, baseshort, the long outer arm a glabrouspaddte-shapedstructure,inner arrn terminating in a penclantblackened blade with a slender curverl spine on outer margin at its
base;inner style a simple rod that narrowsoutwardly,terminating in a long st'raightspine,with a few strong sete. Phallosome,p, lvith edeagusyellow,elongate,near apexbent at abouta rightangle.
Habztat.-lndia ( Assam) .
Holotype,male,.Sihai Khulen, Maniptrr,altitude 4,700feet, June25, 1960 (Schmid). Paratypes,! males,Nongjri, n'Ianipur.alti'
tude 350 feet, April 1, 1960 (Schmid).

96, I

Alenander:Tipulidn from EasternAsia,LXU

53

The paratypesagreewell with the type exceptthat the inneraun of the intennediatedististyleof the hypopygiumis serrulated. other regionalmembersof the subgenusthat have theinner dististylesimpleincludeGonomyin(ldiocera'yabsonaAlexander,G. (1.) coheriana
Alexander,G. (1.) pergracilisAlexander,
G. ( I. ) phallostenaAlexander,G. ( I. ) prori,nw Brunetti, andothers,all clifferingfrom the presentfly in hypopygialstructure,includingthe nonserratedouter dististyle.
GONOIIIYIA flDIOCERil

SITA sp. oov.

plarc 3, lig. 31, plate j, ilr. q7,

Generalcolorationyellow,patternedwith dark brorvn;.rostrum
and proximal antennalsegmentsyellow; knob of halteresobscure
1'ellou':legs yellorved;wiugs light yellorv, unpatterned;male hypopygium rvith outer dististvie bifid, the others simple; redeagus
pale. generally parallel-siderl.tenninating in a slender point.
Il,Iale.-Length,about 6 millirneters;wing, 5.5.
Rostrum yeliorv, mouthpar:tsand palpi black. Antennre withscapeand pedicellight yell.ow.basalsegmentsof flagellum brov,'itish yellow, the outer ones darker brown; verticils exceedingthe
segments,pubescencervhite, relatively long and eonspicuous.
Head light yelioiv, center of vertex slightly darkened.
Pronotal scutum yelIor,v,.
narrowly brown nredially, scutellum
and pretergiteslight yeiiol. Ivlesonotal
prrescutumwith disk covered b,yconfiuentlight brown stripes,hunreraland lateralbo,rders
narrowly yellowed;scutal lobeslight broln, posteriorcallosities
yellou'; scntellumlight br'o\.,-i, parascutellalight yellow; postnotum brolnish yellorv,cer,ii':.ri
legir.lnof rnediotergiteweakly darkt'ned. PJeuralight bror,,,ii.
u'ith a drfiusepale yellow longitudinal
str:ipeextentlingfrotn belrin,rthe fore cox& aerossthe dorsalstcrnopleurite,rvidely expa;ideCllcliinrl; dorsopleuralregiorl brorvn.
Ilalteres rvith stem darkened,ilrob otrscureyellow. Legs wiilr
coxE and trochantersyellow, fore coxreslighily clarker;remainderof iegs yellowed,tips of tibia and outer tarsal segmentsslighily
darker. wings (Plate B, fig. BB) almost uniformly light yellow,
pt'earcularand eostalfields cleareryellorv;stigmadarker yellorv,
scareelyindieated;veins light brown. yellorvedin the brightcnetl
fields. Venation: Sc short, Sc, ending about oppositeone-fifth
Rs. sc.-.far retractediRr+z and R, closelyapproxirnatedat cosir,
virtualiy closingceli R1;vein R,,strcngly obiique,wiilrout trichia;vein R1 long, gently arcuated;cell 2nd Mz longer than its petiole

54

Journnl of Scierce
The Philippi,neAbdomen elongate,tergites brown, sternites more yellowed.
Male hypopygium(Plate 5, fig. 4?) with outer lobe of basistyle,b, pale, terminating in a microscopicpale point. Outer dististyle,d, unequally bifid at tip, the lateral blade large, triangular, extendedinto an acutepoint, inner arm an oval blackenedlobe; in'termediatestyle shortestbut stout, outer end blackened,with trvo
or three smallblack spines;inner style a simplerod, its stem very
slender,outer third slightly dilated,terminatingin a small blackened point. Phallosornewith edeaguspale, the stem generailyparallel-sided,
the tip a slenderpoint.
Habitat.-India (Assam) .
Holotypemale,Leimatak,Manipur,altitude 1,300feet, l\[ay 30,
1960 (Sch,mid).
(Idiocera)
The most similar regionalspeciesinclude Gonom,gin
prorima Brunetti and G. (1.) oayu sp. nov., all being very different amongthemselvesin hypopygialstructure.
GONOIIIYIA (IDIOCEnA) rERRIBILIS sp. nov.

Plate 3, lig. 3f; Phtc t' nt. et.

Generalcolorationof thorax graY,prascutum rvith two inter'mediatebrown stripes.the lateral bordersyellowed;pleura darkwith a yellow longitudinalstripe; wings yellowed,outrvardiypat'
terned with brown; male hypopygiurnwith apical lobes of basi'
styte; three dististyles,all producedinto blackenedspines,innerstyle bispinous;adeaguslong and slender',tip recurved.
Male.-Length, about 7.5 millimeters;lving 7.
Rostrumand patpi black. Antennrervith scapeyeliorv,black'ened beneath.pedicel clear yellow, proximal two flagellar seg'ments pale brown, remainder black; segmentselongate. Head
Bray, anterior orbits broadly light yellow.
Pronotumpale yellow, dorsumof scutum more obscured,pre'tergites yellow. Mesonotalprescutum with lateral borders ancl
humeri yellow, disk gray with two brown stripesthat are faintlr'
indicatedon anteriorhalf, posteriorlydarkerand widened;scutunr
obscureyellow, lobes brownishgray; scutellumbrown, posteriorborder obscureyellow; mediotergitegray, posteriorhalf darker.
antero-lateralareasyellow, confluentwith an area on the dorsal
pleurotergite.remainderof the latter brown, variegatedwith yellow. Pleura dark brown with a conspicuouslongitudinallight yellow stripe that includesthe centralthird of fore coxa.tlorsalsternopleurite,widcnedbehindon ventral pteropleurite,dorsallreronand metapleura;dorsopleuralmembraneyellowed. Halteresyel-

96, I

Alerander: Tipulzdn jrom Eastern Asia,LI.'II

55

iol, baseof stem weakly darkened,knob dark brown. Legs withfore coxe brown, patternedas described;middle and hind coxrc
dark brown basally, apices yellow, the middle pair extensively
so; trochantersobscureyellow; femorayellow,tips vaguelybrown;tibie yellow, tips narrowly brownish black; tarsi brown, darker
outwardly. Wings ( Plate 3, fig. 34) with the ground yellow, baseclearer,disk conspicuouslypatternedwith brown, including especially the darker stigma and broad tips of cells R,rand Ra; narrowbrown areasat arculus,origin of Rs,cord and very inconspicuously
on veins I\[3,1\[,,Cur and 2nd A; veins yellow, clark brown in thepatternedareas. Venation: Sc1endingjust before midlengthof
Rs, Sc2retracted;m-cumore than its length beforethe fork of M.
Abdomen dark blown. Niate hypopygium (Plate 5, fig. 48)
with the basistyle,b, telnrinating iu two short lobes,the inne.r
one somewhatmore slender,with long yellow sete throughoutits
length,the broaderouter lobe with apexglabrousand more broadly obtuse. All three dististyies, d, bearing conspicuousblackspines;outer style longest,the distal half slightly expandedintoa yellow blade,narrolvedat tip iuto an acutespine,inner rnargin
with two subequalbtrackspines,one at near one-fourththe length,
the other about at two-thirds; intermediate style shorter, entirely blackened,narrowed gradualty into a long terminal point,
the inner malgin with two spines,ihe outer one shorterthan theterminal point, the basalspine at near midlength,very long andslender; inner style smallest,ciividedinto two slighfly unequal
spines,near basesof thesewith severallong yellow setre. phal
Iosome,p, with the aedeagus
a long siender rod, its tip slighily
decurved;outer surfacervith llumerouslong yellow sete extending fronr baseahnostto apex.
Habitat.-India (Assarn).
Holotype,lnale,shergaou..Karneng.
.\ortireastFr.ontierAgency,
altitude 6,300feet, May 7, 1961 (schmid).
The very conspicuousmale hypopygiumof this fty readily se.
paratesit from all other known rnembersof ilre subgenus. The
t'egionalspecieshaving patternedn'ings and with the inner dististyle of the hypop"vgiumbispinousinclude Gonomgia(ld,iocera)acifurcaAlexander,G. (1.) furcososp. nov.,G. (I.) kctsltongensis
sp. nov.,G. (I.) magraAlexander,G. (/.) rnaharajoliAlexander,c. {1.\ rnyriacantha
Alexancler,ancla few others,all quite distinct
amongthemselvesin hypopygialcharacters.

56

The Ph,ilippineJaurrtal of Science

GONOII{YIA (IDIOCERA) VAYU r

noY.

Plate 4, tls. 75i Plate 5, fig 5u.

Generalcolorationof mesonotumbrownishgraY,pleura andpleurotergitelight ;'ellow;rostrum bron,nishblack; antenniervith
basalsegmentsyellow; femorayellou';rvingsyellou'ed,costalattdprearcularfields clear light yellorv,Sc relativelyshort; maie h5'popygiumwith apex of outer lobe of basistyleterminatingin a
miscroscopicblack spine; outer dististylebifid; adegeausstout,
parallel-sided,
apex gently emarginate.
Male.-Length, about5 millimeters;wing, 4.8.
Rostrumand palpi brownishblack. Antenne rvith scapeantipedicelyellow; flagellumbroken. Head light yellow, cettterof
vertex pale brown.
Pronotalscutumobscureyellow,indistinctlypatternedwitlt paiebrown, includingcentral and lateral areas;scutellumanclprctergites tight yellow. Mesonotalprescututn browrtish gray tt'ith
vaguelyindicateddarker brown intermediatestripes,pseudosutural fovee reddish brown; posteriorscieritesof notum brorvn.
sparselypruinose. Pleura and pleurotergitelight yeilow, ciorsopl'eura1region slightly darker. Halteresrvith stem light brorvn:
knob broken. Legs with cox& and trochatttersyeliott'; fentora
yellow;remainderof iegsbroken. Wings (Plate4, fig. 35) yellowed,prearcularand costalfields clear tight yeliorv;veius ver)'pale brown, yellow in the brightenedfields. Venatiotl:Sc reRs. Sc- farlatively short, Scr ending about oppositeone-seventh
retracted;veinsR r+z and R'r slightiy separatedat costa.ccll R 'large;cell 2nd M: a lrttle longerthan its petiole;m'cu abouttn.ice
its length befo'refork of I\'I.
Abdorninaltergitesbrown.sternitesanclltypopygiummore 1'ellowed. Male hypopygium(Plate 5, fig. 50) rvith outer lobs .1
blackenedspitte,innerbasistyle,b, terminatingin a microscopic
lobe very slender,with aboutthree long sete. Outcl dististl'le.
d, with outer half expandedinto two oval blatles.tips obtusc,api'
cal angle of outer blade farther extendedinto a ltarrow h1'aiine
iobe;intermediatestyle simple,smooth,gently curved.tlte blackenedapexnalTowedinto an acutespine;inner sti'le lcugcst,simir:to a lorrgblackspine.
ple, a long slenderrod with tip narrowecl
rnner margin of rod rvith ahoutten scattcred{'l'cctsetrc. Phallosome,p, with edeagusstout, parallel-sidcd,altex gently emar'
ginate.

96, 1

Alerander: 7'ipuitdu iro rn'I'jcstct'li,ltsiu,LXII

57

Habitat.-India ( Assam).
Holotype, male, Bhairabkutttla,Katneng, Nr;r'tlteastIirotrtier
Agency,aititude700 feet, i\Iarch3, 1961 (Sclmdd)OthergenerallysiurilarregionalspeciesinciudeGonornyia(Idiocere,)disptosoAlexonder,G. ( I.) petil,isAlcxander,G. (I.) phnl'
LostenaAlexancier,G. (I.) nt;yriocanthaAiexancler,and someothers,all quite distinctin the structureof the male hypop"n*gium.
S u b g e n u sP T IL OSTENODESAlexander
GONOMYIA (PTILOSTDNODIS) CAPITULA

sp. nor'.

Irlatc 4, fig. 36; Plate 5' f-ig, 49.

Generalcgloratioublack, variegatedr','ithyellow r.'nhead andespeciallyon the thoracicpleura;antennie,halteresancllegsblack;wings whitish hyalinei Scr ending oppositeorigin of [ls, Sc1very
iong, subequalto Rs, cell 2nd N{r less than one-half its petiole,cell 2nd A relatively long; male hypopygiumwith inner apicallobeof basistyledilated at outer end.

Mate.-Length,,about4.3millimeters;wings,5.2;antenna,about
1.
Rostruurclarli brorvn;paipi black. Atttennreblack throughoui;
fiagellar segmentsoval, the outer segmentsgradually smaller,siightty exceedingthe verticils. Head biack, the auterior vertex
and broad anterior orbits ligttt yellow.
Pronotumblack,the enlalgedpretergalregiou light yellor'l'.I'Iesouotal prrescuturnu'ith Cisk cialli Br3)', hutneral and iateral regiorrs blackened;scutuiri dalk gI'aY, uiedian region yeliowed;
scutellumgray basall"v,variegatecili'itlt ;'eliol beltincl;postnotum
darli gray. Pieura polisheciblack olr aileiiisiertrttmand ventral
stenropleurite,the rcuraindet'.irrciuriiugpleulotergite,light yellou'. Halteres blackenecl. Legs 1.,'ililccxic clarli gray; tt'ochanters grayishblack; remainderof icgs biack. Wings (Ptate 4, ftg.
36) whitish hyaline; veins bror,vtt. Venatjon: Sc1 ending op'
posite origin of Rs, Sc, Iar retracted, Sct alone subequal toRs; outer section of vein R; iottg, tnore than twice Prs;ceII 2ndi\,[, less than one-halfits petiole; ur-cu lnore than its own length
before fork of N'I;vein Cu: virtually lacking, indicated only by avague fold near base; ccll 2nci A relatively long. Macrotrichia
on outermost raclial and rnediai veins, including also the outerlhird of Cur.
."., ,,".i1,,,1
Abdominal telgites, inciuding hypopygiurri,clull black, sternites
paler. i\fale hypopygiurn(Plate 5, tig. 49) rvith outer apical lobe

5B

Th,ePhili,pplneJourrwl of Science1967

of basistyle,b, broadly obtuse,with denselong curved sete eon'tinued along mesalface of lobe. What is construedas being an
inner apicaltobe of the basistyleis an elongaterod, apex dilated,outer margin of rod with about eight strong bristles. Outer dististyle, d, longest,deeply forked beyondmidlength,outer branchabout one-half longer than the inner; inner style smallest, 8P'
pearing as a gently curved horn that narrows to the acute tip.
p, with edeagus terminating in a small apiculate
Phallosome,.
point, outer end with relatively few setre.
Habitat.-India ( Sikkim).
Holotype, male, Chateng,altitude 8,700 feet, June L2,,1959(Schm,id).
Gonomyin (Ptilostenodes)capitula is most readily told from
G. (P.) stenamerosp. nov., by the hypopygialstructure,particularly the dilatedapexof the inner lobe of the basistyle. G. (P.)
pakr,stanensu
Alexander, the most westerly known member of thesubgenus,has the venational details distinct.
GONOMITA (PTILOSTENODES)STENOIYIERAcp. nov.

Plate f, llg. 77i Plate 5, Ilg il.

Generally as tn capitulo, differing chiefly in hypopygial characters, particula'rly the basistyles.
Mate.-Length, about 4 millimeters; wing, 5; antenna,about 1.
Female.-Length,,about 4.5 millimeters; wing, 5 to 5.2.
Rostrum brown, palpi and antenn& black, the latter relatively
long. Head brownish black, anterior vertex and orbits yellolv.
Pronotumchiefly yellow. Mesonotumblack on anterior borderof prrescutum,the remainder sparselygray pruinose,patternedwith yellow at midregion of scutum,.posterior scutal callosities,
patand the scutellum. Pleurachiefly light yellow, conspicuously
vagueiyextend'
ternedwith black on propieuraand anepisternurn,
ed to the wing root; ventral sternopleuriteblack, gray pruinose,
midsternal region yellow. Halteres and legs brownish black to
black, coxe pruinose. Wings (Piate 4, fig. 37) whitish hyaline;
veins brown. Venation: Scr ending shortly before origin of Rs,
Sczfar retracted; petiole of cell 2nd Mz approximatelythree times
the cell.
Abdomendull brownish black, dorsopieuralmembraneand adloining margins of scleritesyellow. Male hypopygium l Piate 5,
fig. 51) generaliy as in capituln, diff.ering evidently in the conforrnationof the imer lobe of basistyle,b, u'liich is slender,tapering very gradually to the narrowly obtuse apex, at and near tip

96, I

Alerander:'I'ipu,lidnirorruEasternAsia,LmI

59

with about eight sete. with morc numcrouspale puncturesthat
lack seta ahnostto baseof lobe. Other miuor clifferencesin thebasistyleand dististyles,as the fen'er setie at apex of the outer
lobe of the former.
Habitqt.-India (Sikkirn) ,Holotype,male, Shingbo,altituclc10,400feet, June 30, lgbg
t Schmid). Allotopot)'pefemale, pinnecl u'ith type. Paratopotypes,severalof both sexes,mostlyin poor conditionand mounted
or1a single microscopeslide u'ith the t1,pes. The entire series
had been seriouslydamagedb5rfungus action.
S u b g e nusLIPOPHLEPSBer gr oth
GONOIIIYIA(LIPOPHLEPS) AII1BLYSTYLA sp. nov.

Plate 5' llgs. 52, 6'1.

Altied to inconrpleta;mesonotalprescututn and scutal lobes
brownish grsy, pleura yellow. pattemeclwith brown; anteune
with scapeand pedicelyellow, flagellum black; legs dark brown;
wings pale brown, prearcular and costal regions slightly moreyellowed,Sc, endingsomedistancebeforeorigin of Rs, branchesof latter stronglydivergent,cell Ravery extensiveat margin; ntale
hypopygiumwith a single dististyle,its apex obtuseanclunblackened, with a stroug black spine at near midlength; phallosome
large and conspicuous,apex bilobed.
Male.-Length, about 3 to 3.2 millimeters;u'iug, 3.5 to 3.8.
Femnle.-Length.about 4 nriliirncters;rviug,4.
Rostrum brownish yellow, palpi black. Antennreu'ith scapeand pedicel yellow, flagellurn black; flagellar segrneutslongcylind,rical,with abundant very long erect r,r'hitesete. Front
vellowed,vertex gray.
Pronotum ancl pretergitesvellorv. llesonotal priescutumand
scutal lobes brownish gray; rncciianre.gionof scutum and posterior borders of scutal iobes .v-eilolr'ed;
scutellumyellorv,parascutelamore obscure;mediotergiteclark broln, 1'ellorved
laterally, pleurotergiteyellow. Pteui'a.incluciingthe dorsopleuralmelnbrane, yellorv,the mesepisterllunrancl meron more infuscated.
Halterespale brown, baseof stem yellolved. Lgs with coxe andtrochanterspale yellow, remainderof legs dark brown. Wings( Plate 6, fig. 52\ pale brolvn, stigrla scarcelyindicated,preareular and costal fields slightlv ulore yellowed;veins very pale
bt'o$'n.Venation:$c, endiuga iiistancetiefot'eorigin of Rs about
equal to three-fourthsRs. branchesof the latter strongly diver-

60

The PhilippirteJournol of Science196?

the basalsectionof R; regent,cell R* at marginvery extensive,
duced (as shorvn)or longer; cell lst M, stronglynarroweclat
base,n-cu at fork of UI.
Abdomenlight brown above,paler on sternites,hypopygiunt
yellorved. Male hypopygium( Piate 6, fig. 64) with apicallobeof basistyle,b. .long,subequalto or longer than the dististyle.
on the two sides;tip obtuse,d, which are slightly asyrnrnetricalon outer margin at or just beyondmidlengthwith a blaekened
spinethat is stouteron one style than on the other. Phallosome,pale,stemstout,outwardlyexpandp, very largeandconspicuous,ed into broadblades,terminatingin two needlelikeblack spines.
Habi,tat.-India( v!'estBengal) .
7, 1959
ai[itude3,930feet,September
Lingsoka,
male,,
Hoiotype,(Schm,id).Atlotopotype,female.Paratopotypes,
2 males,pinned
femaie.
with allotype;1 sepa'rate
Although it is generallysimilar to GonornytalLipophleps) incom,pieta( Brunetti) -synonyms, G. ( t. ) insulensisAlexander;(1964)726],the presentfly is disL. (G.) pusilla[Lackschewitz
especiallythe pale obtuseapexof
tinct in hypopygialcharacters,
the dististylethat bears the usual two fasciculatesete. In hr
the apex of the ciististyieis a sienderblack spine.
com,pleto
GONOMYIA (LIPOPHLEPS)CURVISTYLA Alexander.

Platc 6, tigs. 53, 0t.

Gonomgi,a(LtpophLe'ps)curuisfglc Aluxaxoun, Jour. N. Y, Ent. Soc.
69 (1961) t26-r27.

Type-male, Teri, Teri Ciarhwal, I(unraon,altitude 2,500 to
5,000feet,April 16, 1958(Scltndd).
Venation( Piate6, fig. 53) ; nralehypopygium( Plate6, fig. 65) .
GONOMYIA (LIPOPIILEPS) HESTTCA sp. nov.

Ptrte 6, llgs. 5d, 06.

Sizesmall ( vring of male 4 miilinieters) ; rostrum, palpi and antenne black; mesonotumbrorvn, posterior border of seutellum
abscureyellow; legs yellowish brown; wings pale brown, costal
border tight yellow; male hypopygium generally as in bicolorata
but with the details distinct.
Male.-Length, about 4.5 rniiiimeters;wing, 4; antenna,about
0.9.
Rostrum and palpi black. Autennre biackened;flagellar segments elongate,the more proximal ones vrith exceedinglylong
verticais. Fiead iight ycilow, central area of posterior vertex
darkened.

96, I

Alerander:Ti,pnltd,efrom EasternAsia,LXll

61

Pronotumand pretergiteslight yellorv.sidesof notum darkbrown. Mesonotalprrescutumlight brown, vaguelymore darkened medially,scutumdarlier brown; scutellumbrown basally,
posteriormargin obscureyellow;mediotergitepale brown, sidesof anterior half yellow. Pleura obseureyellow,patternedwitlr
brownandapparentlywith a narrov/whitenedlongitudinalstripe.
Halteresurith stem yellow, knob light brorvn. Legs with coxac
and trochantersyellow,remainderpale yellowishbrown,the extremetip of femur vaguelypaler. Wings (Plate6, fig. 54) patebroln, costalborde'rnarrou'ly pale yellorv,interruptedby theslightly darker brown stigma;veins pale brorvn,more yellowedin the brightenedfield. Costalfringe long. Venation:Branchesof Rs divergent,R; deflectedstronglytoward the rving tip, cell
Rr at ma,rginvery extensive;m-cushortlybeforefork of M.
Abdominaltergitesand hypopygiumbrown,sternitesmoreyellowed. MaIe hypopygium(Plate 6, fig. 66) with dististyles,d,terminal; outer style a long slendernearly straight blackened
rod, the tip obtuse,,on mesalface at basewith an elongatesetiferous cushion;inner style about one-thirdas long, blackened,
the body elongate,with severalstrong seta at apex and alonglower margin,the more basalof thesesubtendedby a strong
point or tooth; outer margin of styte at near midlengthrvith astrongspinelikelobe,neartip with trvostrongseta. Phallosome,p, entirelypale,the two outer bladesor apophyses
subhyaline,
tips obtuse.entirelyglabrous.
Habitat.-India (Assam).
Holotype,male, Chapai,Kameng,NortheastFrontier Agetrcy,
altitude700feet, February26, 1961(Scltmid).
Other regionalspeciesof the subgenusLipoTthteps
that have
two radialbranches
andthe malehypcpygiumof this generaltype
includeGonomyin(Lipopltleps)antbiensAlexander,G. (L.) bL
colorataAlexanderandG. (L.) ytallicostata
Alexander,all having
the hypopygialdetailsdistinct. It is possiblethat ambi,ensand
palli,costata
may prove to be identical. I havenot been able tostudyan authenticmalespecimenof G. (L.) flauomnrginata
Brunetti but from the descriptionof color charactersof the type it
appearsto be distinctfrom the presentfly.
GONOMYIA OIPOPHLEPS) XAMA Alexandcr. plate 6, ftgs. jj,

J6, 6Ti plate T, ttg. or.

Gonurn'yia(Lipophleps) lcamaAr,rxluonR, Jour. N. y. Ent. soe. Tl(1963)76-77.

The Phi,lippineJownnl of Science62

I(umaon,altitude 7,250feet,Type--male,Sutol,Pauri Gat'ltrval.
August 18, 1958 (Schmid).
Venation(Plate 6, figs.55,56; male hypopygiumPlate 6, fig.
67; Plate7, fig.68).
coNoMyIAolpopHl,Eps)

MIZoENSIS Alexander.

Plate 6. iis. l7z Plate 7, fis. 69.

Ar,sxaNosn,Jour. N. Y' Ent' Soc'
Gonomgia(Lipophleps) mizoensi^s
7l (1963)77-79.

Tlpe-male, Thingsat,Lushai Hills, Mizo District,.Assam,8l'
9, 1960 (Schmid)'
titude 2,000feet, September
Venation( Plate6, fig. 57) ; malehypopygium( Plate7,.fig. 69) .
GONOMYIA (LITOPHLEPS) NTLGIRIANA Alexander.

Plate 6' tlg. 5E; Plate 7' rtg', 7o',

Gonomaia (Lipophleps) nilgiriana Ar,nxlNpun,Jour. N.
69 (1e61) L2g-L29.

Y. Ent' soe'toType-Perumalmalai,Madras,south India, altitude 4'000'
4,5d0feet, December6' 1958(Schmi'd\(Plate7, fig.70)'
venation(Plate6, fig. 5s); malehypopygium
GONODTYIAGIPOPIILEPS) SPERATINA sp. nov.

Plate 6, tlg' 59i Plate 7' llg' ?l'Sizesmall (wing of male lessthan 4 millimeters);mesonotal
browniitrgray,posteriorscleritesof notumyellowed;prreseutum
pt*ur" dark brownwith a bioad silveryyeltowlongituclinalstripe;*ing. very light b,rown,stigmascarcelvdifferentiated,Scrending
distia short distaneebeforeorigin of Rs; malehypopygiumwithouter style a powerfulyellowrod thatstylesterminalin position,.
inner maris extendedinto a l,ongblackspine,at nearmidlengthof
gin with a similar appressedspine;phallosomeincludingthreefurther strong sPines.
Male.-Length,about3.Smillimeters;wing'3'8'
pedicelRostrumanl patpi black. Antenne black, seapeand
yellowedabove;flagellarvertieilsverv long' Headaboveextensively yellowed.
p'rescuPronotum and pretergiteschiefly yellow. Mesonotal
scutaltum, scutume.ndbise of scutellumbrownishgray,posterior
chief'
callositiesandbroadmarginof seutellumyeilow;postnotum
narrowly-pleura posterior border of mediotergite
t brownish yellow,
dark brownwith a broadsilveryyellowlongidark brown.
tudinal stripe extenclingfrom behindthe fore cox& to the baseof abdomen. Halteresyellott" basalhalf of knob light brown'
Legswith eoxa andtrochantersobseureyellow;remainderof legs

e6,I

Alerander: Tipulide from Eastent,Astn,LI.ll

68

broken. (wings (Plate6, fig. bg) very light brown,prearcularand costalfields light yellow;stigmascarcelydifferentiatecl,
very
pale brown; veinslight brown,more yeilowedin the brightened
fields. venation:scr endinga distaneebeforeorigin of Rs about
equalto basalsectionof vein R;, Sc2at its extremetip; branchesof Rs strongly divergent,cell R+at margin very extensive,fully
three times cell R.; m-eu about one-fifth its length beyondfork
of M.
Abdominaltergites and hypopygiumbrown, sternitespaler.
Male hypopygium(Plate T, fig. 7L) unusualiycomplexin structure. what appearsto representthe tergite,,t,includes
two wid+ly separatedfingerlike lobes, their mesal parts with unusuallylong seta. Dististyles,d, terminal;,outer stytea powerfulyellow
rod, strongly curved on outer third, extendedinto a long blackspine,on inner margin at near midlength with a smaller more
appressedblack spine; inner style elongate,with severallong
sete, the two fasciculatebrisiles terminal. phallosoffi€,p, complex, ineludinga y-shapedstructureabout as figured,,eacharm
extendedinto a strongblack-tipped
spine;on inner marginwith
a slendercurved arm, toward tip slighily expandedand withdelicatesete; adclitionalto the abovestructurethe phallosome
includessomeothers,the mostconspicuous
beinga further powerful blackenedspine.
Habitat.-India ( Assam) .
Holotype,male,Khanggai,Manipur,altitude4,428feet,July 1,6,
1960(Schmid).
Gonomyia(Lipophleps)speratinois quitedistinctfrom all othermembersof the subgenusin the hypopygialstrueture,especiallythe outer dististyleand phallosome,togetherrvith the furcate
structurethat is believedto representthe tergite.
GONOIIYIA GIPOPHLEPS) TURRITELLA Atexander.

Plate 7. llS. ZZ,

Gonarwia (Lipophleps) tumitelto Ar.p:caxorn,Jour. N. y. Ent. soc.
66 (1958) 165-166.

tlpe-Parewavir, Nepal,lVlarch26, tgST (Coher).
Male hyopygium( Plate 7, fig. 72).
GONOMYTA (LIPOPHLEPS) VARSHA

Arexander.

ptate 6r ftg. 601 plate 7r fig. 7!.

Gonomgia (Lipophleps) uorsha Ar,exaNosR,Jour. N. y. Eht. soe.
7l (1e63) 79-80.

64

196?

The Phil.ippineJournal,of Science

Type-Pagna Malla,Pauri Garhu'al.Kumaon,altitude5'995feet'
August11, 1958(Se'hnt'id).
Venation(Plate6. fig. 60); malehypopygium(Plate7. fig' 73)'
SubgenusGONOMYIAMeigen
GONOIIIYIA (GONOMYI0FULVIPINNIS sp' nov'Plate

6, fig. 61.

Sizeverylarge (wingof femaleB millimetersor more);rostrum
with darklight yelloy, intennrc blaeli; tnesonotalpraescutum
biown stripes,seutallobesextensivelydarkened,pleura yellorv,
stripedwith brown; halteresyellorv;legs brown;rvingsstrongly
tight fulvous,prearcularand costalregionscleareryellow,stigma
pale brown;veinsSc and Rr long.
Femnle.-Length,about6.5 to ? nrillimeters;wing, B to B'5'
Rostrumlight yellow; palpi black. Antenne relatively long'btackthroughout. Heacllight yellorvon front andanteriorvertex.
behindextensivetydark brolvnishgray'q'ith sidesbroad'prrescutum
light yellorv. Mesonotal
P,ronotum
disl biovrnishgray with three dark stripesthat arely ye11ow,,iiuia.Ovaguelybehind,in casesthe lateralstripeslacking;scutulTyellow,ut.t, tonewith a large triangularbrown areathat is nar'iowed behind,confluentacrossthe suturervith the lateral prre'scutaldarkening;scutellumyellow;mediotergiteyellow,posterior
lr::lfr;ith trvolaige brorvnspots,anteriorborderfaintly darkenecl'
in caseswith a more evident central infuscation. Pleura andpleurotergitewhitish yellolv with a narrow brown longitudinal.tripu trom the cervicairegio,nacrossthe propleura,endingon thepaler grayishbrown.
ventralsternopleurite
veniral anepisternum;
Halterestight yellow. Legswith coxe whitishyellow;troehanters
yellow; femorabrown, obscureyetlow basally,beeomingdarker
brorvnoutrvardly;remainderof legs brown. Wing (Plate 6, fig'61) stronglytigtrt fulvous,the prearcularand costalregionselear
yuilo*, stigmapale brown; veinsbrorvnishyellow,cleareryellow
in the brighteneclparts. Venation: Sc long, Se, ending aboutorcuated,subequalto R.r;vein
opposite*lOt.ngttr of Rs; R21314
r- long, rJecurved,ending shortly before the wing tip, basal
sectronof Rr short; m-cu at from about one'fourth to neariy
one-halfI{r*1.
Abdominaltergitesdark brown, sidesnarrowly paler; pleural
membraneand sternitespalerbroln, posteriorbordersof the latter narowly yellorv.

e6,1

Alerander: Tipu,tidafrom EastemAsin,,LXU

65

Habitat.-India ( Sikkim) .
association,
altitude
Holotype,female,Kalep,in Rhododendron
12,000feet,JuneLB,1959(Schmid). Paratypes,
severalfemales,
Yagtang,in Rhododendro?aassociation,
altitucle11,600to 11,650
feet,June 17, 1959(Schmi.d).
GonomEin(Gonomyia)fuluipennisis the largestregionalmember of the subgenus,
beingreadilytold from othersby the unusual
sizeand stronglyfulvouswings.
GONODTYIA (GONOIIIYIA) HIRSUTISTYLA OITUSISTYLA

subtt). nov.

Differs from typical hirsuttstyla in slight details of the malehypopygium. Outer lobe of basistylenot prolonged,the apicalhalf subobtuse,with very long sete. Outer distisytle slender,
withou'ter half only slightly widened,tip subobtuse.,lEdeagus
outer spine or apophysissmall.
Habitat.-India (Sikkim).
altitudeassoeiation,
Holotype,male,Yedang,in Rhododendrqn
female,pinned
10,600feet,June9, 1959(Schm,id).Allotopotype,with type.
GoNoMYtA (coNoMYra) sulAFrINrS rp' nov'

llate 6' tV' 62i Pler 7' nt' 74'

Allied to affi,ni.s;general eoloration of mesonotummedium
brown, variegatedwith yellow, pleura light yellow; wings sub'hyaline,without a stigmaldarkening;Sc relativelylong, Scr end'
ing about oppositeone-fourthRs; male hypopygiumwith apicalflangeof outer dististylevery low; inner style with two unequalwithblackenedspines.the outer one angularlybent; phallosome
apophysesblaekened,very unequal,one long and ribbonlike;adeaguselongate.pale.apexsuborbicular.
Male.-Length,about4.5 millimeters;wing, 5.3.
Rostrumlight yellow; palpi brown. Antenne with scapeyel'lowed,the remainderbrown; flagellarsegmentslong, the intermediateoneswith long verticils. Front yellowed,vertex brownish g,ray.
pre'Pronotalscutumlight yellorv,scutellumbrown. Mesonotal
scutumand scutallobesalmostuniformlymediumbrown,central
area of scutum,posteriorbrders of scutal callositiesand thescutellumlight yellow; mediotergitetight brown, anterolateralangles,pleurotergiteand pleuralight yellow. Heltereswith stomtight yellow, knb very slightly darker. Legs with coxc yellow,
fore pair darkenedanteriorly;trochantersyellow; remainderof

The PhilippineJournal of Science196?

legs light brown. Wings (Plate 6, fig. 62) subhyaline,unpat'
teined,stigmalacking;veinspalebrown. Veinsbeyondthe cordwith abundantlong trichia. Venation: Sc relatively long, Sct
endingaboutoppositeone-fourthRs, Scznearlyobsolete;R2n'3aa
nearlytwiceRs;cell lst Mz gentlywid'
only moderatetylrcuated,enedoutwardly,subequalin lengthto vein Mr.
Abdomen light brown, sternites paler. Male hypopygium(Plate7,fig. ?4) with apicallobeof basistyle,b, small,aboutoned, the latter virtually simple,
third as long as the outer dististylQ,,with a very low apicalflange on mesalface; inner style unitedbasally with the outer, distally with two blackenedspines,theouter one more angularlybent, with a singlestrongsetain thevery unequal'one an elong'axil. Phallosoil€,P, with apophyses
ate blaekenedribbon, slightly expandedbefore the acute tip;
secondapophysisa very small spine at baseof the longer one;edeaguspite, elongate,the suborbicularapexstightly expanded.
Hobi,tat.-India (Assam) .
Kameng,NortheastFrontierAgency,
Holotype,male,,Shergaon,
altitude6,400feet, May B, 1961(Schmid).
fire elosestatly of this speciesappearsto be Gonomyt'a(Gono'
nyio (affi,nisBrunetti, deserlbedfrom the Darjiling district inttre easternHimalayas.Of this I havea femaleparatypereceived
from Brunetti by exchangeof materials,this agreeingfainly wellwith his speciesdescription,includingthe darkenedwing stigma.
The type of the speciesis a male but Brunetti providedno account of the genitaliestruetures.
GON0MYIA (GoNoMYrA) TANA(rcANTHA

Alexander. Platc 6r ll8' 63t Plttc 7' try' 7t'

GonvrUtia (Gonorryio) tnnaocanthaAr,uxlxonn, Jour. N. Y. Ent. Soc.
7l (1963) 75-76.

Bpe-Chumtang, Sikkim, altitude 5,120feet, July 18, 1959
(Schmid).
(Plate7,fig.75).
Venation(Plate6, fig.63); malehypopygium
novum
3ugcnusTEUCHOGONOTTAYIA

Charaetersgenerallyas in typical Gonomyza.Antenna withpnoximaltwo or three flagellar segmentspartially united to form
outer flagellarsegments
an incompleteelongatefusion-segment;
long, with relatively short verticils. Wings with Rs relatively
long,from aboutone-halfto two-thirdsthe long straightRz+r+c;

96, 1

Alerand,er:Tiputt'd,efrom EastemAsin,LXn

67

Mz gencell Rasmall,vein R:rsuberect,R+r€afty straight;cell 1st
openby atrophyof basalsectionof
erally closed,in percornplefro
notMr; anterior arculuslacking. Male hyopygiumwith basistyle
producedinto apicallobes;three terminal dististyles,all simple;
interouter style longind slenderwith microscopicapicalpoints,
mediatestyle *ittt outer and triangularlyexpanded. Phallosome
distinctive,symmetrical,very large, complicatedby branchingof
the gonapophyses.
-sungenus; Gonornaia(Teuchogonowia) noueboro'
type orcensisAlexander ( 1916) EasternNearctic.
(1919),.G.
Ahexanderother speciesincludeG. (T.) aci.cuti,fera
Alex'), G. (T.) seatere??sis( T.) percompleraAlexander ( 1.946
ander-( 1948i and G. ( ?.) tetonensisAlexander ( 1946) ' of theAlexander ( 1941),
western Nearctic region; G. ( ?.) honi,bi.Lr,seasternpalaearctic( Japan) , and G. ( T.) edwardsiLackschewitz( 1g2b) and G. ( T. ) ithyphallus Lackschewitz( 1935) , western
Palaearctic(EuroPe).
Illustrationsof the male hypopygiuminclude the following:
G. (T.) ednorilsi Llcrscunwrrz, Arbeit Naturforsch. Vereins Riga(n.f.) t6 (1925) 7-8, fig. 4; T.rnonn,Bo-Opusculaentomolgiea6(1941)60-61, fie.2.
Philip. Jour. Sci. 76 (1941) 60-62' fig.
G. (T.) horribclisAr,sxANoEB,
M,
Lacxscnnwlrz, Naturhist. Maandblad (Limburg'
G. (?.) ithgphatfu,cs
Maastrieht)24 (1935) 14, pl. 3' fig. 8.
Alnxlxpnn, Proe. Aead. Nat. Sci. Philadel'G. (?.) nouebmacensis
phia for 1916 ( 1916)523,pl- 30, figs. ?9' 80.

ILLUSTRATIONS
ll.egcnd: b, basistyle; d, distirtyle; p, phallo$mc.l

Plers

I

{rc. 1. Gonomyi,a(ProtogonomAio)apocryphosp.nov.;venation.
2. Gonomyia (Protogonomgin) clista Alexander; venation.
3. Gonomgio (ProtogonomAin\ disti,furu, Alexander; venation.
4. Gonomgia (ProtogonomAia)glabristgla sp.nov.; venation.
a . GonamAia (ProtogonomAia) lateromncula sp.nov.; venation.

6. Gonomyia (Protogonomqia)scoteina sp.nov.; venation.
7. Gonomgin (Protogonomqta) thiorhopala Alexander; venation.
B. Gonomgia (Ramagonomgin)protensa Alexander; venation.
9. Gonomyia (Siaagonomgia)discolophnllossp.nov.; venation.
10. Gonomyin (ProtogonomAia) acustglata Alexander; male hypcpy.
glum.
1 1 .Gonsrnyin (Protogonoryia) api,culata sp.nov.; male hlryopygium.
12. Gonomgia (ProtogonomAia)apougpha sp.nov.;male hypopygium.
13. Gonomyio (ProtogonomAi,a) clisto Alexander; male hypopygium.
Pl,nrs 2

Frc. 14. Gonomyit (Protogonomyia) contostgla sp.nov.; male hypopygium.
15. Gonomgin (ProtogonomAin) distifurca Alexander; male hypopy-

grum.
16. Gonomgi,a(ProtogonomAin)glabri,stylasp.nov.; male hypopygium.
L7. Gonomyin (ProtoganomAin) klwsinna Auxaxorn; male hypopy-

grum.
LB. Gottomgia (Protogonomuia) lateromacula sp.nov.; male hypopy'

gium.
19. Gonornyi'a(Protogonomgin) megalomntoArexander; male hypopygium.
20. Gonomyia (Protogonomyia) pellar sp.nov.; make hypcpygium.
2 1 . Gonom.gia(Protogonomyia) rejecta sp.nov.; male hypopygium.
PLArs 3

.Frc.22. GonomEia(Protogonomaia)scoteinasp.nov.;male hypopygium.

23. Gonomyia (Protogonomyia) thiorhopala Alexander; male hypopy.

gium.

24. Gonomuia (SiuogonomUia) discotophallossp.nov.; male hypopv-

gium.

25. Gonomgio (Ramagonomgia) bisicutifero Alexander; male hl'popy-

gium.

26. Gonomvio (Ronngonomgia) yrotensa Alexander; male hypopy-

gium.

69

70

Journnl of Science
The Phili,ppt'ne

196?

27.Gonomaw (Idiocera) acbenophallossp.nov.; venation.
28. Gonomuia (Idiocera) acifurca Alexander; venation.
29. Gonomgin (Idiocero) furcosasp.nov.;venation.
30. Gonomgia(Idiocera) kashongensissp.nov.; venation.
31. Gonomgia(Idiocera) lamio sp.nov.;venation.
32. Gonomyia (ldiocera) leda sp.nov.;venation.
33. Gonomvia (Idiocero) sito sp.nov.;venation.
34. Gonomgin (Idiocero) terribilis sp.nov.; venation.
Pratn 4
Ftc. 35. Gonomgi,a(Idiocera) uagu sp.nov.; venation.
36. Gonomyi,a(Ptitostenodes)capituln sp.nov.; venation.
37. Gonowit (Ptilostenodes) stenomerosp.nov.;venation.
38. Gonomgin (Idiocero) acaenoplwllossp.nov.; male hypopygtum.
39. GonomAin(ldiocera) acifurco Alexander; male hypopygium.
40. Gonom,gia(ldiocera) cohernno Alexander; male hypopygium.
4L. Gonomgtn (ldiocero) furcosa sp.nov.; male hypopygium.
42. Gonomyio (ldiocero) kashongensissp.nov.; male hypopygium.
43. Gonomyin (ldiocero) lomin sp.nov.;male hypopygium.
Pllrn

5

Ftc. 44. Gonomgi,o(ldiocera) ledn sp.nov.;male hypopygium.
45. Gonomain (Idi.oeera) petilis Alexander; male hypopygium.
46. Gonomgi'a (Idiocera) sertatistgla sp.nov.;male hypopygiurn.
47. Gonomgin (Idiocera) si,tasp.nov.; male hypopygium.
48. GonornAin (Ptilostenodes) capitula sp.nov.; male hypopygium.
49. Gonomaia (Ptilostenodes) capttuLasp.nov.; male hypopygium.
50. Gonowia (Idiocera) uagu sp.nov.; male hypopygium.
5t. GonomAin (Ptilostenodes) stenomera sp.nov.; male hypopygium.
Plarn 6
Frc. 52. Gonomgin (Li,pophleps)amblgstala sp.nov.; venation.
53. Gonomyia (Li,pophleps) curaistglo Alexander; venation.
54. Gonomgio (Li,pophleps)hesticc sp.nov.;venation.
55' 56. GonomAin(Lipophleps) komn Alexander; venation.
57. Gonomyin (Lipophleps) mizoenstsAlexander; venation.
58. Gonomgia (Li,pophleps) nilgiriana Alexander; venation.
59. Gonomgia (Li,pophleps)sperotinn sp.nov.; venation.
60. Gonomyin (Li,pophleps) aarsha Alexander; venation.
61. Gonomgio (Gonomgin) fului.perm,issp.nov.; venation.
62. GonornAi,a(Gonornyin) suboffinis sp.nov.; venation.
63. Gonomgin (Gonomgia) tnnaocontho Alexander; venation.
64. Gonornuia(Lipophleps) amblAstAlnsp.nov.;male hypopygium.
65. Gonomgio (Lipophleps) curuistyla Alexander; male hypopyglurn
66. Gonomgta (Li,pophleps) hesti,casp.nov.; rnale hypopygium.
6?. Gonomyia (Li,poph,leps)korna Alexander; male hypopygium.

96, 1

Ateronder:Tiputi'dufrom EastemAsia,LXn

7L

PI.rm 7
Frc. 68. GmmWia (Lipophleps) komn Alexander; male hypopygium.
69. GonomUin(Lipophteps) mizoensi.sAlexander; male hypopygiun.
70. Gonornun (Lipophleps) nitgiriona Alexander; male hypopygium.
7L. Gonomaio (Lipophleps) speratinosp.nov.;male hypopygium.
72. Gonotrryia(Lipophleps) tunitelln Alexander; male hypopygium.
73. Gonurngin (Lipophleps) aarslw Alexander; male hypopygium.
74. GonurnAin(Gmqrn'yin'l suboffmis sp.nov.; male hypopygium.
75. GonornAin(GonmnUia) tmwocantha Alexander; male hypopygium.

ALExANDER: TrputronE rnou EnsrEnN Aste, tXII.l[Pntun. [OUn. Sct., VOr.96,

No. l.

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PLATE I.

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43
PLATE 4.

No. I,

AT,EXANDEn:
Trpwmer rnou Easrrnx Asrl, LXILI[Pruur. toun. Scr., Vor.. 96, No. l,

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foun. Scr., vor.96,

No. l.

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PLATE 6.

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]oun. scr., yot. 96, No. l.

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PLATE 7.