EVAN P. ECONOMO, Ph.D.
Professor, Biodiversity and Biocomplexity Unit (http://arilab.unit.oist.jp/)
Dean of Faculty Affairs
Okinawa Institute of Science and Technology Graduate University

1919-1 Tancha
Onna-son
Okinawa, 904-0495
Japan+81 (098) 982-3328 (JP)evaneconomo@gmail.com
Citizenship: USA, Canada

EDUCATION
2009 Ph.D. University of Texas at Austin. Ecology, Evolution, & Behavior.
2002 B.S. University of Arizona. Ecology & Evolutionary Biology.

APPOINTMENTS
Okinawa Institute of Science and Technology Graduate University
2023 - present Dean of Faculty Affairs
2019 - 2022 Associate Ombudsperson2020 - present Professor, Biodiversity and Biocomplexity Unit2012 - 2020 Assistant Professor, Biodiversity and Biocomplexity Unit2011 - 2012 Visiting Assistant Professor
Harvard University
2021 - 2022

Mary I. Bunting Fellow, Radcliffe Institute for Advanced Study

University of Michigan
2012 - 2017 Visiting Assistant Prof., Dept. of Ecology and Evolutionary Bio., Museum of Zoology
2009 - 2012 Postdoctoral Fellow - Michigan Society of Fellows
Assistant Prof. (non tenure-track)- Dept. of Ecology and Evolutionary Biology

INSTITUTIONAL LEADERSHIP EXPERIENCEOIST Dean of Faculty Affairs (2023-present)
I work closely with the President, the Provost, the Dean of Research, and the Dean of the Graduate
School, to manage essential functions of the University and to support the well-being and success of
90 faculty and 480 research staff. As DFA I manage the Faculty Affairs Office to administer faculty
tenure and promotion reviews, external reviews of research units, faculty recruitment, professional
development, research misconduct investigations, salary and promotions for research staff, the library,and participate in overall university management and strategic planning.

OIST Associate Ombudsperson (2019-2022)
In my role as Associate Ombudsperson for the campus I assisted OIST members with conflict
resolution issues including harassment complaints, employee issues, etc.

Principal, OIST/Onna Children’s School of Science (2019-2022)
I was one of two Principals and the sole OIST faculty representative on the executive committee of
the Children’s School of Science, a yearly summer science program we run in partnership with thelocal community (Onna village) that has included over 800 grade-school children over the years. The
operations of the school are run by the OIST community relations section.

OKEON Churamori Project & Environmental Research Support Section
Since 2015, I have led development of the Okinawa Environmental Observation Network (OKEON)
Churamori Project. This project is designed to both further ecological science activities at OIST but
also form partnerships with local communities in Okinawa. We established 24 long-term monitoringsites across the island, and in the process made partnerships with over 100 entities including highschools, local museums, local Universities, NGOs and government entities.

OIST Committees
2023- present | Executive Committee
This committee, chaired by the President, meets weekly to share key information, discuss issues, andsupport the President in deciding essential matters for the University.
2023- present | Faculty Awards Committee (Chair)
I recently formed this committee to promote and facilitate faculty to apply for and receive scholarly
awards. Our main focus is to break down language and other understanding barriers for applying forawards in different countries, as well as proactively nominate and/or encourage underrepresented
researchers to apply for awards.
2023- present | Space Allocation Committee (Chair)This committee makes decisions regarding space allocation among all research and administrative
units on campus. In addition, I am responsible for strategic planning of space on campus.
2023- present | Research Staff Appointments Committee (Chair)This committee evaluates all non-faculty research staff for appointment, salary changes, promotions,and continuing (permanent) employment status.
2023- present | Core Facilities Steering Committee
This committee advises the Provost on operation all the core facilities including operations, policies,
and strategic planning.
2022 | Appointments and Promotions Committee
This committee is positioned between hiring / promotion committees and executive leadership, toensure that relevant procedures are followed and recommendations meet criteria.
2021 | Tenure Review Committee
Tenure review committee member for OIST faculty member.
2021-2022 | Conference and Workshop Committee
Responsible for selecting conferences and workshops to receive support from OIST and set policies
for operations.
2020 | Working Group on Procurement
This is an ad hoc committee responsible for revising procurement regulations, particularly
developing a response to some new regulatiosn from the Japan Ministry of Finance.
2019-present | OKEON Steering Committee
Advise and manage OKEON project and Environmental Science Support section.
2019 | Working Group on Travel Regulations
This was an ad hoc committee responsible for revising travel regulations, I was asked by the faculty
council to represent faculty concerns, and this led to substantial revision of travel rules.
2018 | Perspective Council
This was an ad hoc committee of extremely distinguished scientists (4 of the 6 Nobel laureates, chairedby Steven Chu (former Secretary of Energy, USA) tasked with planning the scientific direction of
future growth / faculty hiring at OIST. I was one of 6 individuals chosen to represent the OIST facultyon this committee, where I particularly represented the fields of Ecology/Environment/Marine
Science.
2016 | Task Force on Rank and Tenure
This was an ad hoc committee formed by the faculty to examine policies and procedures for tenure
and promotion reviews. We made recommendations to the president which led to substantial policychanges.
2016-2019 | Curriculum and Examinations CommitteeStanding committee responsible for curriculum and exam related issues for our PhD program,including setting policy, overseeing curriculum, overseeing proposal exams and final thesis defenses,
approving granting of degrees, etc.

2015-2021 | Human Subjects Research Review Committee (IRB)
This committee oversees and approves all human subjects research at OIST. I was included on thiscommittee particularly as a biostatistics expert.
2013, 2014, 2018, 2019 | Faculty Search Committees
I was a member of faculty search committees for Marine Science in 2013 and 2014, and for
Ecology/Environment in 2018, and have been member of a target of opportunity search committee in2019.
2013-2022 | Gender Equality and Diversity Committee
Promoting Gender Equality and Diversity on OIST campus, including promoting family-friendly
environment, advising on tenure (e.g. stop-the-clock) policies, etc. I was a founding member of thiscommittee.
2013-present | Information Technology Service and Support Committee
This committee oversees IT issues at OIST and advises the CIO, including email systems, phonesystems, privacy and security policies, etc.
2013-2015 | Faculty Council
The main governance committee for OIST faculty, elected by faculty peers.
2012-2013 | Admissions Committee
Responsible for admissions to the OIST PhD program

PUBLICATIONS
Book
Sarnat, E., Economo, E.P. (2012) The Ants of Fiji. University of California Press.

Preprints
Aibekova, L.A, Keller, R.A, Allman, D.M., Hita Garcia, F., Labonte, D., Narendra, A., Economo,
E.P. (2023) Parallel and divergent morphological adaptations underlying the evolution of
jumping ability in ants. bioRxiv.
Ross, S.R.P.J., Friedman, N.R., Armitage, D.W., Dudley, K.L., Yoshida, T., Yoshimura, M.,
Economo, E.P., Donohue, I. (2023) Spatial divergence in ecological responses to typhoons
across a subtropical island. bioRxiv.
Kass, J.M., Yoshida, T., Yoshimura, M., Ogasawara, M., Suwabe, M., Hita Garcia, F., Fischer, G.
Dudley, K.L., Donohue, I., Economo, E.P. (2023) Breakdown of ant community seasonalpatterns over a land cover gradient. bioRxiv.
Nathan, P, Economo, E.P., Guénard, B., Simonsen, A., Fredrickson, M.E. (2023) Generalized
mutualisms promote range expansion in both plant and ant partners. bioRxiv.

Warren, D.L., Kass, J.M, Economo, E.P. (2022) Randomization analyses in niche and distribution
modeling. EcoEvoRxiv.
Warren, D.L., Kass, J.M., Casadei-Ferreira, A., Economo, E.P. (2022) Incorporating sampling bias
into permutation tests for niche and distribution models. bioRxiv.
Luo, Y., Taylor, A., Weigelt, P., Guénard, B., Economo, E.P., Nowak, A. Inderjit, Kreft, H. (2022)
Ant richness and environment shape the global distributions of ant-associated plants. bioRxiv.
Frank, E.T., Kesner, L., Liberti, J., Helleu, Q., LeBoeuf, A., Dascalu, A., Milutinovic, B., Azuma, F.,
Economo, E.P., Waridel, P., Engel, P., Schmitt, T., Keller, L. (2022) Infection signaling andantimicrobial wound care in a predatory ant. bioRxiv.
Toulkeridou, E., Gutierrez, C. E., Baum, D., Doya, K., Economo, E.P. (2021) Automatedsegmentation of insect anatomy from micro-CT images using deep learning. bioRxiv.

Published | In Press
122. French, C.M., Bertola, L.D., Carnaval, A.C., Economo, E.P., Kass, J.M., Lohman, D.J., Marske,
K.A., Meier, R., Overcast, I., Rominger, A.J., Staniczenko, P., Hickerson, M.J. (In Press)
Global determinants of the distribution of insect genetic diversity. Nature Communications
121. Wang, C., Chung, F., Lin, C., Katzke, J., Economo, E.P., Billen, J. (2023) Morphology of themetapleural gland and its associated novel atrial cone gland in Strumigenys ants. Micron. 171:103463120. Liu, D., Essl, F., Lenzner, B., Moser, D., Blackburn, T.M., Cassey, P., Biancolini, D., Capinha,C., Dawson, W., Dyer, E.E., Guénard, B., Economo, E.P., Kreft, H., Pergl, J., Pyšek, P., vanKleunen, M., Nentwig, W., Rondinini, C., Seebens, H., Semenchuk, P., Weigelt, P., Winter,
M., Dullinger, S. (2023) The impact of land use on alien species incidence and richness inlocal assemblages worldwide. Nature Communications 14: 2090.
119. Wong, M., Economo, E.P., Guénard, B.G. (2023) The global spread and invasion capacities ofalien ants. Current Biology 33: 566-571.
118. Liu, C., Economo, E.P., Guénard, B.G. (2023) GABI-I: The global ant biodiversity informaticsisland database. Ecology.
117. Richter, A., Boudinot, B., Hita Garcia, F., Billen, J., Economo, E.P., Beutel, R.G. (2023)
Wonderfully weird: The head anatomy of the armadillo ant Tatuidris tatusia (Formicidae:
Agroecomyrmecinae). Myrmecological News 33: 35-75.
116. Wang, R., Kass, J.M., Galkowski, C., Garcia, F., Hamer, M.T., Radchenko, A., Salata, S.D.,
Schifani, E., Yusupov, Z.M., Economo, E.P., Guénard, B. (2023) Geographic and climaticconstraints on bioregionalization of European ants. J. Biogeography 503-514.
115. Overcast, I., Achaz, G., Aguilée, R., Andújar, C., Arribas, P., Creedy, T.J., Economo E.P.,
Etienne, R., Gillespie, R., Jacquet, C., Jay, F., Kennedy, S., Krehenwinkel, H., Lambert A.,
Meramveliotakis E., Noguerales, V., Perez-Lamarque, B., Roderick, G., Rogers, H., Ruffley,
M., Sanmartin, I., Vogler, A.P., Papadopoulou, A., Emerson, B.C., Morlon, H. (2022) Genetic
Theory of Island Biogeography: Inferring Processes from Multi-Dimensional CommunityScale Data. Global Ecology and Biogeography 32: 4-23.

114. Emerson, B.C., Borges, P.A.V., Cardoso, P., Convey, P., deWaard, J.R., Economo, E.P.,
Gillespie, R., Kennedy, S., Krehenwinkel, H., Meier, R., Roderick, G., Strasberg, D., Thébaud,C., Traveset, A., Creedy, T.J., Meramveliotakis, E., Noguerales, V., Overcast, I., Morlon, H.,
Papadopoulou, A., Vogler, A.P., Arribas, P., Andújar, C. (2022) Collective and harmonised
high-throughput barcoding of insular arthropod biodiversity: toward a Genomic Observatories
Network for islands. Molecular Ecology.
113. Liu, C., Fischer, G., Liu, Q., Peng, Y.Q., Economo, E.P., Guénard, B. (2022) Updating thetaxonomy of the ant genus Myrmecina (Hymenoptera, Formicidae) in China with descriptionsof four new species. Zootaxa 5182: 152-164.
112. Wepfer, P., Nakajima, Y., Economo, E.P.†, Mitarai, S.† (2022) The oceanographic isolation of
the Ogasawara Islands and genetic divergence in a reef-building coral. J. of Biogeography 49:
1978-1990. †joint last authors
111. Wang, W., Soh, E.J.Y., Yong, G.W.J., Wong, M.K.L., Guénard, B.G., Economo, E.P., Yamane,S. (2022) Remarkable diversity in a little red dot: a comprehensive checklist of known ant
species in Singapore (Hymenoptera: Formicidae) with notes on ecology and taxonomy. Asian
Myrmecology 15: 015006.
110. Kennedy, S., Calaor, J. Hans, J., Zurápiti, Y., Yoshimura, M., Choo, J., Andersen, J., Callagan,
J., Roderick, G.K., Krehenwinkel, H.†, Rogers, H.†, Gillespie†, R.G., Economo, E.P.† (2022)
Richness and resilience in the Pacific: DNA metabarcoding enables parallelized evaluation ofbiogeographic patterns. Molecular Ecology. †joint last authors
109. Kass, J.*, Guénard, B., Dudley, K., Jenkins, C.N., Azuma, F., Fisher, B.L., Parr, C.L., Gibb, H.,
Longino, J.T., Ward, P.S., Chao, A., Lubertazzi, D., Weiser, M., Jetz, W., Guralnick, R.,
Blatrix, R., Des Lauriers, J., Donoso, D., Georgiadis, C., Gomez, K., Hawkes, P., Johnson,
R.A., Lattke, J., MacGown, J.A., Mackay, W., Robson, S., Sanders, N., Dunn, R.R.,
Economo, E.P.* (2022) The global distribution of known and undiscovered ant biodiversity.
Science Advances 8: eabp9908. *corresponding
108. Arribas, P. Andújar, C., Bohmann, K., deWaard, J., Economo, E.P., Elbrecht, V., Geisen, S.,
Goberna, M., Krehenwinkel, H., Novotny, V., Zinger, L., Creedy, T.J., Emmanouil
Meramveliotakis, E., Noguerales, V., Overcast, I., Morlon, H., Papadopoulou, A., Vogler,
A.P., Emerson, B.C. (2022) Toward global integration of biodiversity big data: a harmonised
metabarcode data generation module for terrestrial arthropods. GigaScience 11: 1-12.
107. Parker, E., Jones, C., Economo, E.P., Zapfe, K.L. Federman, S., Near, T.J. Dornburg, A. (2022)
Periodic environmental disturbance drives repeated ecomorphological diversification in an
adaptive radiation of Antarctic fishes. The American Naturalist.
106. Khalife, A., Peeters, C., Economo, E.P. (2022) Minute workers and large soldiers in thesubterranean ant Carebara perpusilla: functional consequences of muscle allometry in thethorax. Arthropod Structure & Development 69: 101188.
105. Andrade-Silva, J., Baccaro, F.B., Prado, L.P., Warren, D.L., Kass, J., Guenard, B., Economo,
E.P., Silva, R.R. (2022) A large-scale assessment of ant diversity across the Brazilian Amazon
Basin: integrating geographic, ecological, and morphological drivers of sampling bias.
Ecography 16: e0695.

104. Gómez, K., Kouakou, L., Fischer, G., Hita–Garcia, F., Katzke, J., Economo, E.P. (2022)
Pheidole klaman sp. n.: A new addition from Ivory Coast to the Afrotropical pulchella species
group (Hymenoptera: Formicidae: Myrmicinae). Zookeys 1104: 129-157.
103. Aibekova, L., Boudinot, B. E., Beutel, R.G., Richter, A., Keller, R.A., Hita Garcia, F., Economo,
E.P. (2022) The skeletomuscular system of workers of the generalized ant genus Formica(Formicidae). Insect Systematics and Diversity 6: 2.
102. La Richeliere, F., Dunn, R., Economo, E.P., Guénard, B., Sanders, N., Weiser, M., Abouheif, E.,
Lessard, J.P. (2022) Warm and arid regions of the world are hotspots of social complexity.
Proceedings of the Royal Society: B. 289: 20211899.
101. Katzke, J. Puchenkov, P., Stark, H., Economo, E.P. (2022) A roadmap to reconstructing muscle
architecture from CT data. Integrative Organismal Biology 4: obac001.
100. Takashina, N., Jenkins, C., Planck, M., Economo, E.P. (2022) Species-range size distributions:
integrating the effects of speciation, transformation, and extinction. Ecology and Evolution 12:
e8341.
99. Boudinot, B., Richter, A., Katzke, J., Keller, R., Economo, E.P., Beutel, R., Yamamoto, S. (2022)
Evidence for the evolution of eusociality in stem ants and a systematic revision of†Gerontoformica (Hymenoptera, Formicidae). Zoological Journal of the Linnean Society 195:1355-1389.
98. Beutel, R., Friedrich, F., Economo, E.P. (2022) Patterns of morphological simplification andinnovation in the megadiverse Holometabola (Insecta). Cladistics 38: 227-245.
97. Silva, R….(203 authors including Economo, E.P.)…Ribiero, M.A. (2021) ATLANTIC ANTS: adataset of ants in Atlantic Forests of South America. Ecology 103: e03580.
96. Klunk, C.L., Argenta, M.A., Casadei-Ferreira, A., Economo, E.P., Pie, M.R. (2021) Mandibularmorphology, task specialization, and bite mechanics in Pheidole ants (Hymenoptera:
Formicidae). Journal of the Royal Society Interface 18: 20210318.
95. Richter, A., Keller, R. A., Hita Garcia, F., Billen, J., Katzke, J., Boudinot, B. E., Economo, E.P.*,
Beutel, R.G.* (2021) Head anatomy of Protanilla lini (Leptanillinae, Formicidae,
Hymenoptera) and a hypothesis of their mandibular movement. Myrmecological News 31: 85114. *joint supervision
94. Casadei-Ferreira, A., Friedman, N.R., Economo, E.P., Pie, M.R., Feitosa, R.M. (2021) Head andmandible shapes are highly integrated yet represent two distinct modules within and among
worker sub-castes of the ant genus Pheidole. Ecology & Evolution 11: 6104-6118.
93. Booher, D., Gibson, J., Liu, C., Longino, J.T., Fisher, B.L., Janda, M., Narula, N., Toulkeridou, E.,
Mikheyev, A.S., Suarez, A., Economo, E.P. (2021) Functional innovation promotes
diversification of form in the evolution of an ultrafast trap-jaw mechanism in ants. PLoS
Biology 19: e3001031.
92. Suzuki, Y., Economo, E.P. (2021) Dispersal network topology and spatial environmental
autocorrelation mediate the balance of species sorting and mass effects in metacommunities.
Ecography 44: 1-12.

91. Rosas-Mejia, M. Guénard, B. Aguilar-Méndez, M.J., Ghilardi, A., Vásquez-Bolaños, M.,
Economo, E.P., Janda, M. (2021) Introduced ants (Formicidae: Hymenoptera) in Mexico – thefirst database of records. Biological Invasions 23: 1669-1680.
90. Ross, S.R.P-J., Friedman, N.R., Yoshimura, M., Yoshida, T., Donohue, I., Economo, E.P. (2021)
Utility of acoustic indices for ecological monitoring in complex sonic environments.
Ecological Indicators 121: 107114.
89. Takashina, N., Economo, E.P. (2020) Developing generalized sampling schemes with known
error properties: the case of a moving observer. Ecography 43: 1-14.
88. Casadei-Ferreira, A., Economo, E.P., Feitosa, R.M. (2020) Revisions to the Pheidole(Hymenoptera, Formicidae) fauna of the Brazilian southern grasslands. Revista Brasileira de
Entomologia 64: e20200068.
87. Peeters, C., Keller, R.A., Fischer, G., Khalife, A., Economo, E.P. (2020) The evolutionary loss offlight enabled the remarkable strength of ant workers. Frontiers in Zoology 17: 33.
86. Liu, C., Fischer, G., Hita Garcia, F., Yamane, S., Liu, Q., Peng, Y.Q., Economo, E.P., Guénard,
B., Pierce, N. E. (2020) Ants of the Hengduan mountains: a new altitudinal survey andupdated checklist for Yunnan Province highlight an understudied insect biodiversity hotspot.
ZooKeys 978: 1-171.
85. Dias, R.K.S., Guénard, B., Economo, E.P., Akbar, S.A., Udayakantha, W.S., Wachkoo, A.A.A.
(2020). The Ants (Hymenoptera: Formicidae) of Sri Lanka: A taxonomic research summary
and updated checklist. ZooKeys 967: 1-142.
84. Dinets, V., Friedman, N., Yoshimura, M., Ogasawara, M., Economo, E.P. (2020) Acoustic
detection of an unknown bat species in Okinawa. Mammal Study 45: 1-4.
83. Casadei-Ferreira, A., Fischer, G., Economo, E.P. (2020) Evidence for a thoracic crop in theworkers of some Pheidole species (Formicidae: Myrmicinae). Arthropod Structure &Development 59: 100977.
82. Fischer, G., Friedman, N.R., Huang, J.P., Knowles, L.L., Mikheyev, A.S., Fisher, B.L., Economo,
E.P. (2020) Socially parasitic ants evolve a mosaic of host-matching and parasitic
morphological traits. Current Biology 30: 1-8.
81. Wepfer, P., Nakajima, Y., Radice, V., Toonen, R., Richards, Z., Ang, P., Sutthacheep, M., Chen,
A., Sudek, M., Fujimura, A., Mikheyev, A.S., Economo, E.P.*, Mitarai, S.* (2020)
Evolutionary biogeography of the coral genus Galaxea across the Indo-Pacific ocean.
Molecular Phylogenetics and Evolution 151: 106905. *co-last author
80. Friedman, N.R., Lecroq-Bennett, B., Fischer, G., Sarnat, E.M., Huang, J.P., Knowles, L.L.,
Economo, E.P. (2020) Macroevolutionary integration and modularity of phenotypes within
and across ant worker castes. Ecology and Evolution 10: 9371-9383.
79. Aoyama, Y., Yoshimura, M., Ogasawara, M., Suwabe M., Economo E.P. (2020) Potential
economic impacts of invasion of the red imported fire ant in Okinawa, Japan. Japanese
Journal of Ecology 70: 3-14. DOI: https://doi.org/10.18960/seitai.70.1_3 [in Japanese, originalcitation: 青山 夕貴子, 吉村 正志, 小笠原 昌子, 諏訪部 真友子, エコノモ P. エヴァン (2020) 沖
縄県におけるヒアリの侵入・蔓延時に推定される経済的損失. 日本生態学会誌 70: 3-14.]

78. Miao, B.G., Peng, Y.Q., Yang, D.R., Kubota, Y., Economo, E.P., Liu, C. (2020) Climate andland-use interactively shape butterfly diversity in tropical rainforest and savanna ecosystems ofsouthwestern China. Insect Science.
77. Richter, A., Keller, R.A., Hita Garcia, F., Billen, J., Economo, E.P.*, Beutel, R.G.* (2020)
Comparative analysis of worker head anatomy of Formica and Brachyponera (Formicidae,
Hymenoptera, Insecta). Arthropod Systematics and Phylogeny 78: 133-170. *co-last author
76. Beutel, R.G., Richter, A., Keller, R., Hita Garcia, F., Matsumura, Y., Economo, E.P., Gorb, S.N.
(2020). Distal leg structures of the Aculeata (Hymenoptera): a comparative evolutionary studyof Sceliphron (Sphecidae) and Formica (Formicidae). Journal of Morphology 281: 737-753.
75. Liu, C., Sarnat, E.M., Friedman, N., Hita Garcia, F., Booher, D., Mikheyev, A., Economo, E.P.
(2020). Colonize, radiate, decline: unraveling the dynamics of island community assemblywith Fijian trap-jaw ants. Evolution 74: 1082-1097.
74. Yoshimura, M., Suwabe, M., Ikeda, T., Ogasawara, M., Economo, E.P. (2020) Development and
Implementation of a workshop on alien species and Red Imported Fire Ants (RIFA) for
elementary school students. Japanese Journal of Science Communication 26: 39-56. (inJapanese, original citation: 吉村正志, 諏訪部真友子, 池田貴子, 小笠原昌子, エヴァン・エ
コノモ (2020) 小学生向け外来種＆ヒアリ学習ワークショップの開発と実践. 科学技術
コミュニケーション, 26: 39-56.)
73. Cicconardi, F., Gamisch, A., Krapf, P., Wagner, H. C., Nguyen, A. D., Economo, E.P.,
Mikheyev, A. S., Guénard, B., Arthofer, W., Steiner, F.M., Schlick-Steiner, B.C. (2020)
Strong diversifying and relaxed purifying selection are shifting the evolutionary equilibrium of
the alpine ant Tetramorium alpestre (Insecta: Hymenoptera) genome. Molecular Biology and
Evolution 37: 2211-2227.
72. Mao, Y., Qian, H., Shi, J., Economo, E.P. (2020) TREEasy: an automated workflow for theinference of gene trees, species trees, and phylonetworks from molecular sequences.
Molecular Ecology Resources 20: 832-840.
71. Darwell, C., Fischer, G., Sarnat, E.M., Friedman, N., Liu, C., Baiao, G., Mikheyev, A.S.,
Economo, E.P. (2020) Genomic and phenomic analysis of island ant community assembly.
Molecular Ecology 29: 1611-1627.
70. Guo, F., Guénard, B., Economo, E.P., Deutsch, C., Bonebrake, T. (2020) Activity niches
outperform thermal physiological limits in predicting global ant distributions. Journal of
Biogeography 47: 829-842.
69. Wepfer, P., Nakajima, S., Hui, F., Mitarai, S.*, Economo, E.P.* (2020) The metacommunity
ecology of coral-hosted symbionts (Symbiodinaceae). Marine Ecology Progress Series 633:
71-87. *joint last author
68. Choo, J., Gill, B., Zuur, A., Zent, E., Economo, E.P. (2020) Impacts of an indigenous settlement
on taxonomic and functional dung beetle diversity in the Venezuelan Amazon. Biodiversity
and Conservation 29: 207-228.
67. Friedman, N.R., Miller, E., Ball, J., Kasuga, H., Remeš, V.*, Economo, E.P.* (2019) Evolution ofa multifunctional trait: shared effects of foraging ecology and thermoregulation on beak

morphology, with consequences for song evolution. Proceedings of the Royal Society: Series
B 286: 20192474. *joint last author
66. Hita Garcia, F., Lieberman, Z., Audisio, T., Liu, C., Economo, E.P. (2019) Revision of the cryptic
and highly specialized ant genus Discothyrea (Hymenoptera: Formicidae) in the s with X-Ray
microtomography and 3D cybertaxonomy. Insect Systematics and Diversity 3: 5.
65. Sarnat, E. M., Liu, C., Hita Garcia, F., Dudley, K., Economo, E.P. (2019) Ready Species One:
exploring the use of augmented reality for biodiversity discovery with a technology-enhanced
revision of Fijian Strumigenys. Insect Systematics and Diversity 3: 6.
64. Sharaf, M.R., Aldawood, A.S., Economo, E.P., Wachkoo, A.A., Hita Garcia, F. (2019)
Taxonomy of Arabian Temnothorax Mayr (Formicidae: Myrmicinae) with description of a
new species enhanced by micro-CT next-generation morphology. Scientific Reports 9: 11009.
63. Friedman, N.R., Remes, V.R., Economo, E.P. (2019) A morphological integration perspective on
the evolution of dimorphism in sexes and social insect castes. Integrative and Comparative
Biology 59: 410-419.
62. Ross, S.R. Friedman, N.R., Janicki, J., Economo, E.P. (2019) A test of trophic and functional
theories of island biogeography using the avifauna of a continental archipelago. Journal ofAnimal Ecology 88: 1392-1405.
61. Richter, A., Keller, R., Rosumek, F.B., Economo, E.P., Hita Garcia, F., Beutel, R.G. (2019) The
cephalic anatomy of the Wasmannia affinis (Formicidae, Hymenoptera, Insecta) worker casteand its evolutionary implications. Arthropod Structure and Development 49: 26-39.
60. Economo, E.P., Huang, J.P., Fischer, G., Sarnat, E.M. Janda, M., Narula, N., Guénard, B.,
Longino, J., Knowles, L.L. (2019) Evolution of the latitudinal diversity gradient in thehyperdiverse ant genus Pheidole. Global Ecology and Biogeography 28: 456-470.
59. Takashina, N., Kusumoto, B., Kubota, Y., Economo, E.P. (2019) A geometric approach to scaling
individual distributions to macroecological patterns. Journal of Theoretical Biology 461: 170188.
58. Agavekar, G., Agashe, D., Economo, E.P. (2019) Dimensions of ant biodiversity on a tropicalisland. Insect Conservation and Diversity 12: 161-171.
57. Mao, Y. *, Economo, E.P. *, Satoh, N. * (2018) The roles of introgression and climate change inthe diversification and rise to dominance of Acropora corals. Current Biology 28: 3373-3382.
*corresponding
56. Moser, D., Lenzer, B., Weigelt, P., Dawson, W., Kreft, H., Pergl, J., Pysek, P., van Kleunen, M.,
Winter, M., Capinha, C., Cassey, P., Dullinger, S., Economo, E.P., Garcia-Diaz, P., Guénard,B., Maurel, N., Seebens, H., Stein, A., Essl, F. (2018) Remoteness promotes biologicalinvasions on islands worldwide. Proceedings of the National Academy of Sciences 115: 92709275.
55. Khalife, A., Keller, R.A., Billen, J., Hita Garcia, F., Economo, E.P., Peeters, C. (2018)
Skeletomuscular adaptations of head and legs of Melissotarsus ants for tunneling throughliving wood. Frontiers in Zoology 15: 30.
54. Barlow, J., Franca, F., Gardner, T.A., Hicks, C., Lennox, G., Berenguer, E., Castello, L.,
Economo, E.P., Ferreira, J., Guénard, B., Leal, C.G., Isaac, V., Lees, A., Parr, C., Wilson, S.,

Young, P., Graham, N. (2018) The future of hyperdiverse tropical ecosystems. Nature 559:
517-526.
53. Iglesias, T.L., Warren, D.L., Dornburg, A., Wainwright, P.C., Schmitz, L., Economo, E.P. (2018)
Eyes wide shut: The impact of dim-light vision on neural investment in marine teleosts.
Journal of Evolutionary Biology 31: 1082-1092.
52. Staab, M., Hita Garcia, F., Liu, C., Xu, Z.H., Economo, E.P. (2018) Systematics of the ant genus
Proceratium Roger (Hymenoptera, Formicidae, Proceratiinae) in China - with descriptions of
three new species based on micro-CT enhanced next-generation morphology. Zookeys 770:
137-192.
51. Economo, E.P.*, Narula, N., Friedman, N., Weiser, M., Guénard, B.* (2018) Macroecology andmacroevolution of the latitudinal diversity gradient in ants. Nature Communications 9: 1778.
*equal contribution
50. Matos-Maravi, P. Clouse, R. M., Sarnat, E. M., Economo, E. P., LaPolla, J. S., Borovanska, M.,
Rabeling, C., Czekanski-Moir, J., Latumahina, F., Wilson, E. O., Janda, M. (2018) An ant
genus-group (Prenolepis) illuminates the drivers of insect diversification in the Indo-Pacific.
Molecular Phylogenetics and Evolution 123:16-25.
49. Seebens, H. ,..48 authors including Economo, E.P.…Essl, F. (2018) Historical dynamics of
emerging alien species, secondary invasions, and the pool of potential new alien species.
Proceedings of the National Academy of Sciences 115: E2264-E2273.
48. Liu, C., Dudley, K., Xu, Z., Economo, E.P. (2018) Mountain metacommunities: climate andspatial connectivity shape ant diversity in a complex landscape. Ecography 41: 101-112.
47. Ross, S., Friedman, N.R., Dudley, K., Yoshimura, M., Economo, E.P. (2018) Listening toecosystems: data-rich acoustic monitoring through landscape-scale sensor networks.
Ecological Research 33: 135-147.
46. Agavekar, G., Hita Garcia, F., Economo, E.P. (2017) Taxonomic overview of the hyperdiverse
ant genus Tetramorium Mayr (Hymenoptera, Formicidae) in India with descriptions and X-ray
microtomography of two new species from the Andaman Islands PeerJ 5: e3800
45. Hita Garcia, F., Fischer, G., Liu, C., Audisio, T., Economo, E.P. (2017) Next-generation
morphological character discovery and evaluation: an X-ray micro-CT enhanced revision of
the ant genus Zasphinctus Wheeler (Hymenoptera, Formicidae, Dorylinae) in the Afrotropics.
ZooKeys 693: 33-93.
44. Sarnat, E.M., Friedman, N.R., Fischer, G., Lecroq, B., Economo, E.P. (2017) Rise of the spiny
ants: diversification, ecology, and function of extreme traits in the hyperdiverse genus
Pheidole. Biological Journal of the Linnean Society 122: 514-538.
43. Friedman, N., Harmáčková, L., Economo, E.P, Remeš, V. (2017) Smaller beaks for colder
winters: thermoregulation drives beak size evolution in Australian songbirds. Evolution 71:
2120-2129.
42. Dawson, W., Moser, D., van Kleunen, M., Kreft, H., Pergl, J., Pysek, P., Weigelt, P., Winter, M.,
Lenzer, B., Blackburn, T., Dyer, E., Cassey, P., Scrivens, S., Economo, E.P., Guénard, B.,
Capinha, C., Seebens, H., Nentwig, W., Berthou, E., Casal, C., Essl, F. (2017) Global hotspots

and correlates of alien species richness across taxonomic groups. Nature Ecology and
Evolution 1: 0186.
41. Choo, J., Carasco, C., Alvarez-Loayza, P., Simpson, B., Economo, E.P. (2017) Life history traits
influence the strength of distance- and density-dependence at different life stages of two
Amazonian palms. Annals of Botany 120: 147-158.
40. Hita Garcia, F., Fischer, G., Liu, C., Audisio, T.L., Alpert, G.D., Fisher, B.L., Economo, E.P.
(2017) X-ray microtomography for ant taxonomy: an exploration and case study of two new
Terataner (Hymenoptera, Formicidae, Myrmicinae) species from Madagascar. PLoS ONE
12(3): e0172641.
39. Guénard, B., Weiser, M., Gomez, K., Narula, N., Economo, E.P. (2017) The Global Ant
Biodiversity Informatics (GABI) database: a synthesis of ant species geographic distributions.
Myrmecological News 24: 83-89.
38. Economo, E.P., Janda, M., Guénard, B., Sarnat, E.M. (2017) Assembling a species-area curvethrough colonization, speciation, and human-mediated introduction. Journal of Biogeography
44: 1088-1097.
37. Lasky, J., Keitt, T.H., Weeks, B., Economo, E.P.* (2017) A hierarchical model of wholeassemblage island biogeography. Ecography 39: 982-990. *corresponding
36. Jaitrong, W., Guénard, B., Economo, E.P., Buddhakala, N., Yamane, S. (2016) A checklist of
known ant species of Laos (Hymenoptera: Formicidae). Asian Myrmecology 8: 1-32.
35. Fischer, G., Sarnat, E.M., Economo, E.P. (2016) Revision and microtomography of the Pheidole
knowlesi group, an endemic ant radiation from Fiji (Hymenoptera, Formicidae, Myrmicinae).
PLoS ONE 11(7): e0158544.
34. Sarnat, E.M., Fischer, G., Economo, E.P. (2016) Inordinate spinescence: taxonomic revision andmicrotomography of the Pheidole cervicornis group (Hymenoptera: Formicidae). PLoS ONE
11(7): e0156709.
33. Sukumaran, J., Economo, E. P., Knowles, L.L. (2016) Machine learning biogeographic processes
from biotic pattern: a trait-driven dispersal and diversification model with model-choice bysimulation-trained discriminant analysis of Principal Components Classification. Systematic
Biology 65: 525-545.
32. Wepfer, P., Guénard, B., Economo, E.P. (2016) Influences of climate and historical landconnectivity on ant beta diversity in East Asia. Journal of Biogeography 43: 2311-2321.
31. Liu, C., Guénard, B., Blanchard, B., Peng, Y., Economo, E.P. (2016) Reorganization oftaxonomic, functional, and phylogenetic ant biodiversity patterns after conversion to rubber
plantation. Ecological Monographs 86: 215-227.
30. Economo, E.P., Hong, L., Page, S.E.* (2016) Social structure, endogenous diversity, andcollective accuracy. Journal of Economic Behavior and Organization 125: 212-231.
(*corresponding)
29. Janicki, J., Narula, N., Ziegler, M., Guénard, B. Economo, E.P.* (2016) Visualizing andinteracting with large-volume biodiversity data using client-server web-mapping applications:
The design and implementation of antmaps.org. Ecological Informatics 32: 185-193.
*corresponding28. Bharti, H., Guénard, B., Bharti, M., Economo, E.P. (2016) An updated checklist of the ants
(Hymenoptera: Formicidae) of India with their specific distributions in Indian states. ZooKeys
551: 1-83.
27. Sarnat, E.M., Fischer, G., Guénard, B., Economo, E.P. (2015) Introduced Pheidole of the world:taxonomy, distribution, and biology. ZooKeys 543: 1-109.
26. Liu, C., Fischer, G., Economo, E.P. (2015) A rare ant on Samoa: first record of the cryptic
subfamily Proceratiinae (Hymenoptera, Formicidae) and description of a new Proceratium
Roger species. Journal of Hymenoptera Research, 46: 35-44.
25. Guénard, B., & Economo, E.P. (2015) Additions to the checklist of the ants (Hymenoptera:
Formicidae) of Peru. Zootaxa 4040 (2): 225-235.
24. Fischer, G., Azorsa, F., Hita Garcia, F., Mikheyev, A.S., Economo, E.P. (2015) Two newphragmotic ant species from Africa: morphology and next-generation sequencing solve a caste
association problem in the genus Carebara Westwood. ZooKeys 525: 77-105.
23. Guénard, B., Perrichiot, V., Economo, E.P. (2015) Integration of global fossil and modern
biodiversity data reveals dynamism and stasis in ant macroecological patterns. Journal of
Biogeography. 42: 2302-2312.
22. Economo, E.P., Sarnat, E.M., Janda, M., Clouse, R., Klimov, P., Fischer, G., Blanchard, B.D.,
Ramirez, L.N., Andersen, A., Berman, M., Guénard, B., Lucky, A., Rabeling, C., Wilson,
E.O., Knowles, L.L. (2015) Breaking out of biogeographic modules: range expansion andtaxon cycles in Indo-Pacific Pheidole. Journal of Biogeography 42: 2289-2301.
21. Triantis, K.* Economo, E.P.*, Guilhammon, F., Ricklefs, R. (2015) Diversity regulation at themacro-scales: the case of Oceanic archipelagoes. Global Ecology & Biogeography 24: 594605. *equally contributing corresponding authors
20. Liu, C., Hita Garcia, F., Peng, Y., Economo, E.P. (2015) Aenictus yangi sp. n. – a new species of
the A. ceylonicus species group (Hymenoptera: Formicidae: Dorylinae) from Yunnan, China.
Journal of Hymenoptera Research. 42: 33-45.
19. Liu, C., Guénard, B., Hita Garcia, F., Yamane, S., Blanchard, B., Yang, D.R., Economo, E.P.
(2015) New records of ant species from Yunnan, China. ZooKeys 477: 17-78.
18. Hita Garcia, F., Sarnat, E.M., Economo, E.P. (2015) Revision of the ant genus Proceratium
Roger (Hymenoptera, Proceratiinae) in Fiji. ZooKeys 475: 97-112.
17. Economo, E.P., Klimov, P., Sarnat, E., Guénard, B., Weiser, M.D., Lecroq, B., Knowles, L.L.
(2015) Global phylogenetic structure of the hyperdiverse ant genus Pheidole reveals therepeated evolution of macroecological patterns. Proceedings of the Royal Society of London
Series B: Biological Sciences 282: 20141416.
16. Clouse, R. M., Janda, M., Blanchard, B., Sharma, P., Hoffmann, B.D., Andersen, A.N.,
Czekanski-Moir, J.E., Krushelnycky, P., Rabeling, C., Wilson, E.O., Economo, E.P., Sarnat,
E.M., General, D.M., Alpert, G.D., Wheeler, W.C. (2015) Molecular phylogeny of IndoPacific carpenter ants (Hymenoptera: Formicidae, Camponotus) reveals waves of dispersal andcolonization from diverse source areas. Cladistics. 31(4): 424–437

15. Sarnat, E.M., Rabeling, C. Economo, E.P., Wilson, E.O. (2014) First record of a species from the
New World Pheidole flavens-complex (Hymenoptera: Formicidae) introduced to thesouthwestern Pacific. BioInvasions Records. 3: 301-307.
14. Tin, M.Y., Economo, E.P., Mikheyev, A.S. (2014) Sequencing degraded DNA from nondestructively sampled museum specimens for RAD-tagging and low-coverage shotgun
phylogenetics. PLoS ONE 9(5): e96793.
13. Guénard, B., Blanchard, B.D., Liu, C., Yang, D., Economo, E.P. (2013) Rediscovery of the rare
ant genus Bannapone (Hymenoptera: Formicidae: Amblyoponinae) and description of theworker caste. Zootaxa 3734: 371-379.
12. Sarnat, E.M., Economo, E.P. (2013) Pristomyrmex tsujii sp. n. and P. mandibularis Mann
(Hymenoptera: Formicidae). ZooKeys (340): 43-61.
10. Sarnat, E.M., Blanchard, B.D., Guénard, Fasi, J., Economo, E.P. (2013) Checklist of the ants
(Hymenoptera: Formicidae) of the Solomon Islands with new records from Makira Island.
ZooKeys 257: 47-88.
9. Economo, E.P., & Sarnat, E.M. (2012) Revisiting the ants of Melanesia and the taxon cycle:
historical and human mediated invasions of a tropical archipelago. The American Naturalist
180 : E1-E16. (Presidential award from ASN for best paper of 2012 in Am. Nat.)
8. Economo, E.P. (2011) Biodiversity conservation in metacommunity networks: linking pattern andpersistence. The American Naturalist 177: E167-180. (highlighted by Faculty of 1000)
7. Leibold, M., Economo, E.P., Peres-Neto, P. (2010) Metacommunity Phylogenetics: Separating theroles of environmental filters and historical biogeography. Ecology Letters 13: 1290-1299.
6. Economo, E. P., & Keitt, T.H. (2010) Network isolation and local diversity in neutral
metacommunities. Oikos 119: 1355-1363.
5. Cowperthwaite, M., Economo, E.P., Harcombe W., Miller, E., Meyers, L.A. (2008) The ascent of
the abundant: How mutational networks constrain evolution. PLoS Computational Biology 4:
e1000110.
4. Economo, E. P., & Keitt, T.H. (2008) Species diversity in neutral metacommunities: a network
approach. Ecology Letters 11: 52-62.
3. Enquist, B. J., Kerkhoff, A.J., Huxman, T.E., and Economo, E.P. (2007) Adaptive differences inplant physiology and ecosystem paradoxes: insights from metabolic scaling theory. Global
Change Biology 13: 591-609.
2. Economo, E.P., Andrew J. Kerkhoff, Brian J. Enquist. (2005) Allometric growth, life history
invariants, and population energetics. Ecology Letters 8: 353-360.
1. Enquist, B.J., Economo, E.P., Huxman, T.E., Allen, A.P., Ignace, D.D., Gillooly, J. (2003) Scaling
metabolism from organisms to ecosystems. Nature 423: 639-642.

Non-Peer Reviewed:
Economo, E.P. (2016) Understanding western academia: Advice for job-seekers and recommendation
letter-writers [published in Japanese]. Japanese Journal of Ecology 66, 735-742.

Website:antmaps.org. Economo, E.P., Guénard, B., Janicki, J., Ziegler, M., Narula, N. Published online July,
2015.

EXTERNAL ACADEMIC SERVICE
Society Memberships
Ecological Society of America
Entomological Society of America
Ecological Society of Japan
Society for the Study of Evolution
American Society of Naturalists

Society of Systematic Biology
Society of Integrative & Comparative Biology
International Biogeography Society
Japan Society of Mathematical Biology
Inter. Union for the Study of Soc. Insects

Subject/Associate Editor
The American Naturalist (2021-present)
Current Research in Insect Science (2020-present)
Journal of Biogeography (2019-present)
Myrmecological News (2017-present)Peer Reviewer
American Journal of BotanyAquatic Ecology
The American Naturalist
Biodiversity Conservation
Biological Invasions
Biological Reviews
Biology Letters
Bulletin of Math. Biology
Conservation Letters
Current Biology
Diversity and Distributions

Ecology
Ecology Letters
Ecography
Functional Ecology
Global Ecol. & Biogeogr.
Journal of Biogeography
Journal of Ecology
J. of Environmental Manag.
J. of Theoretical Biology
Landscape Ecology
Myrmecological NewsNat. Science Found. (USA)Nature Communications
Oikos
PLoS Biology
PLoS ONEPLoS Comp. Biology
PNAS
Proc. Roy. Soc. B.
Science Advances
Swiss Nat. Science Found.

Symposia, Conferences, and Workshops Organized••••

2024 (upcoming): International Congress of Entomology (ICE 2024), Kyoto, General Affairs
Committee
2023 (upcoming): OIST Workshop on the Evolutionary Analysis of Morphology, primaryorganizer
2020: OIST Mini-symposium: Ant Biodiversity Data Synthesis- Present and Future
2019: SWARM2019: The 3rd International Symposium on Swarm Behavior and Bio-Inspired
Robotics, co-lead organizer and local host (200 participants)

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2018 OIST mini-symposium “Advances in imaging, quantifying, and understanding theevolution of ant phenotypes”
2017 OKEON Chura-Mori Project Symposium2017 Ecological Society of Japan Meeting Symposium, “Big Data in Ecology and Evolution”
2015 East Asia Joint Symposium (EAJS) (co-organizer)
2014 Ecological Society of Japan, “Island Biogeography: Integrating Ecological and
Evolutionary Perspectives”
2014 IUSSI “Island Biology of Social Insects.”

PRESENTATIONS
Invited Talks
2022 International Union Study of Social Insects Congress (symposium)
2022 Concordia University, Department of Biology
2022 McGill University, Department of Biology
2022 Texas A&M University, Department of Entomology
2022 University of Texas at Austin, Section of Integrative Biology
2021 City College of New York, Department of Biology
2021 UMass Lowell, Department of Biology
2021 Yale University, Department of Ecology and Evolutionary Biology
2021 EU-IUSSI annual meeting (Symposium)
2019 Sorbonne University, Institute of Ecology and Environmental Sciences
2019 University of Lausanne, Department of Ecology and Evolutionary Biology
2019 Ecological Society of Japan (Symposium)
2018 Taiwan Entomological Society (Keynote)
2018 UC-Berkeley, Department of Environmental Science, Policy, and Management
2017 Kyoto University, Asia Research Node Symposium2016 International Colloquium on Soil Zoology (Plenary)
2016 International Congress of Entomology (Symposium)
2016 Ludwig Maximilian University Munich (Ants 2016 Symposium)
2015 University of Cambridge, Department of Zoology
2015 Society of Population Ecology Meeting (Symposium)
2014 Kyoto University, Evolutionary Community Ecology Symposium2013 Kyoto University, Symposium in honor of E.O. Wilson and the Kyoto Prize2013 University of the Ryukyus, Japan
2012 Ecological Society of Japan Meeting (Symposium)
2011 Society of Population Ecology Meeting (Symposium)
2011 University of Alabama
2011 Okinawa Institute of Science and Technology
2011 University of Queensland
2011 International Biogeography Society Meeting (Symposium)2010 University of British Columbia
2010 Field Museum of Natural History
2010 UM/SFI Complex Systems Symposium2004 University of New Mexico

GRANTS AND FELLOWSHIPS
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2022- Research Grant Council Hong Kong. “A stage- and trait-based approach to understandglobal ant invasion patterns and invasion mechanisms” ( PI: B. Guénard, Co-I: E. Economo)
2021-2024 Strategic Environmental Research and Development Program ($2.9 million total)“Next Generation Biosecurity Monitoring of Invasive Alien Arthropod Species.” (~$2.9
million total, (PI: G. Roderick, co-PI: Economo, E.)
2021-2022 Radcliffe Fellowship, Radcliffe Institute of Advanced Study, Harvard University
2019-2022 Japan Ministry of Environment Grant (~$1 million), “Developing Countermeasuresfor Invasive Alien Species”
2016-2019 Okinawa Prefectural Government Grant (~$1.2 million), “Developing a system forfire ant surveillance, prevention, and control in Okinawa”.
2017-2019 Japanese Society for the Promotion of Science, Grant-in-Aid of Research(Kakenhi, ~$50K)
2012-2017 National Science Foundation (DEB 1141989), Phylogenetic Systematics Economo,
E. (PI), L. Knowles (co-PI). $378,522. "Evolving hyperdiversity in phenotypic, ecological,
and geographic networks: testing the taxon cycle and alternatives in Indo-Pacific Pheidole"
2009-2012 Michigan Society of Fellows Postdoctoral Fellowship2009 Marion Elizabeth Eason Scholarship, University of Texas
2005-2009 National Science Foundation Graduate Research Fellowship2004-2005 NSF IGERT Graduate Training Fellowship in Computational Phylogenetics
2004 Santa Fe Institute Complex Systems Summer School
2007 EEB Research Fellowship2006 EEB Research Fellowship2005 EEB Research Fellowship2003 University of Texas Dean's Excellence Preemptive Fellowship2004 NSF Graduate Research Fellowship Honorable Mention
2001 Honors College Undergraduate Research Grant, U. of Arizona
2001 Research Training Grant in the Analysis of Diversification, U. of Arizona

AWARDS AND HONORS
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2019 Ecological Research Award (for one of top papers in Ecological Research in 2018)2017 Top ten new species for Pheidole drogon (given by ESF-SUNY)
2016 Biodiversity Action Award (to OKEON project)
2013 Presidential Award from American Society of Naturalists (For best paper published in
The American Naturalist in 2012)2009 Nominated for Distinguished Dissertation Award, University of Texas
2004 Teaching Award, School of Biological Sciences, University of Texas

CLASSROOM TEACHING
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2019 B16 “Ecology & Evolution” Primary instructor, OIST, 9 students
2018 B16 “Ecology & Evolution” Primary instructor, OIST, 8 students
2018 A406, “Independent Study”, OIST, 1 student
2017 B16 “Ecology & Evolution” Primary instructor, OIST. 6 students
2016 B16 “Ecology & Evolution” Primary instructor, OIST, 5 students
2015 B16 “Ecology & Evolution” Primary instructor, OIST, 6 students
2014 A406 “Independent Study”, OIST, 1 student
2014 B16 “Ecology & Evolution” Primary instructor, OIST, 2 students
2013 A302 “Ecology & Evolution” Primary instructor, OIST, 5 students
2011 “EEB Senior Capstone”, primary instructor. U. of Michigan.
2011 “Insect Ecology & Evolution” (graduate seminar) primary instructor. U. of Michigan.
2010 “General Ecology”, primary instructor, U. of Michigan
2009 TA for “Ecology”, 2 guest lectures. U. of Texas at Austin.
2004 TA for “Ecology, Evolution and Society”, 3 guest lectures. U. of Texas at Austin. Won
graduate teaching award from the School of Biological Sciences.
Summer School:• 2013 Co-organized “OIST Integrative Biology Course: Big Data in Biology”(https://groups.oist.jp/oibc)

MENTORSHIPOIST Postdoctoral Fellows & Staff Scientists
2012-2014 Benoit Guénard
2012-2014 Beatrice Lecroq-Bennett
2013-2018 Masashi Yoshimura
2015-2018 Clive Darwell2014-2020 Georg Fischer
2015-2020 Nicholas Friedman
2016-present Francisco Hita Garcia

2017-2020 Nao Takashina (JSPS Fellow)
2018-2021 Susan Kennedy
2019-present Jamie Kass (JSPS fellow)
2021-present Larisa Kiselva
2021-2022 Nurit Eliash2022 Joshua Gibson (JSPS fellow)
2023-present Riou Mizuno

OIST PhD Student Thesis Supervision
2012-2018 Cong Liu
2013-2019 Patricia Wepfer (co-supervised)2014-2019 Yafei Mao (co-supervised)2015-2021 Yuka Suzuki
2016-2021 Evropi Toulkeridou (co-supervised)2017-present Julian Katzke
2017-present Yazmin Zurita Gutierrez
2018-present Gaurav Agavekar
2018-present Lazzat Aibekova
2018-present Shubham Gautam

OIST PhD Rotation Students
2012 Keita Ikegami
2012 Cong Liu
2013 Han Yan
2013 Sandrine Burriel

2014 Yafei Mao
2014 Yuna Hattori
2014 Matti Krueger
2014 Margaret Brisbin

2015 Jason Ball2015 Yuka Suzuki
2016 Menglin Wang
2016 Christopher Campbell
2017 Julian Katzke2017 Otis Brunner2018 Yazmin Zurita Gutierrez

2018 Shubham Gautam
2018 Lazzat Aibekova
2018 Gaurav Agavekar
2019 Mohamed Boubakour
2019 Michael Izumiyama
2020 Kota Ishikawa
2020 Masato Hirota

OIST Research Interns and Visiting Research Students
2012 Sandrine Burriel
2017 Adam Khalife
2013 Patricia Wepfer2017 Tracy Audisio2013 Benjamin Blanchard
2018 Zachary Lieberman2013 Guilherme Baiao
2018 Fumika Azuma
2014 Juliette Martin
2018 Alexandre Ferreira2015 Brett Morgan
2019 Kelvin Chen
2015 Julia Janicki
2019 Adrian Richter2015 Natalie Greenhalgh
2019 April Lamb2015 Matt Ziegler
2022 Lily Tashiro
2016 Haruka Kasuga
2023 Azumi Kudaka
2016 Aina Urano
2023 Arthur Matte2016 Kotaro Fujiyoshi
2023 Jordan Drapin
2016 Victoria McGruer
2023 Karen Kohama
2016 Gaurav Agavekar
2023 Minsoo Dong
2016 Sam Ross
OIST PhD Student Mentor(advisory, not thesis supervisor)
Tomohito Wauke
Maki Kohata Thomas
Christine Guzman
Miles Desforges
OIST PhD Thesis Committee Membership(not including my own PhD students, see above)
Qiong Huang
Tadashi Kazuno
Robert Campbell
Agneesh Barua

Jekaterina Stemmere
Tracy Audisio
Stefano Pascarelli

Menglin Wang
Otis Brunner
Alexsandra Blizina
Tracy Audisio

OIST-Supported Affiliated Researchers(Off-site researchers who have worked with our lab with OIST funding support)
2012-2020 Eli Sarnat2018 Roberto Keller

OIST Research Technicians2012 Adam Lazarus
2013-2014 John Deyrup
2013-2014 Nitish Narula
2015-2017 Julia Janicki
2014-2020 Kenneth Dudley2018-present Fumika Azuma
2019-2022 Kosmas Deligkaris
2021-present Miyuki Suenaga
2023-present Dimitris Petsoupoulos

University of Michigan (2009-2012)
Supervised eleven undergraduate research projects (6 students) for independent study credit or work study(through UROP).
University of Texas at Austin (2003-2009)
Supervised 6 undergraduate students on undergraduate research

OTHER EMPLOYMENT
2002-2003 Research Technician, University of Arizona, Department of Ecology & Evolutionary
Biology, lab of Brian J. Enquist.