ᕝ༤⣖せࠉྕࠉ㸸ᕝཱྀᕷ࡛᥇㞟ࡉࢀࡓほ㈹⏝࣓ࢲ࢝ࡢグ㘓ෆ⏣኱㈗㸦ओ⎔ቃᣦᶆ⏕≀࣭ᇸ⋢┴❧⮬↛ࡢ༤≀㤋እ㒊◊✲⪅㸧ࡣࡌࡵ࡟ࠉ ࣓ ࢲ ࢝㸦 ᅜ ෆ ࡟ ⏕ ᜥ ࡍ ࡿ ✀ ࡢ ⥲ ⛠ ࡜ ࡋ࡚ グ ࡍ 㸧 ࡣ ࢲ ࢶ ┠ Beloniformes ࣓ ࢲ ࢝ ⛉Adrianichthyidae ࣓ ࢲ ࢝ ᒓ Oryzias ࡟ ᒓ ࡍ ࡿῐ Ỉ 㨶 ࡛ ࠶ ࡿࠋ ඖ ᮶ ᅜ ෆ ࡟ ⏕ ᜥ ࡍ ࡿ ✀ ࡣ ✀ ࡜ ࡉ ࢀ ࡚ ࠸ ࡓ ࡀࠊ ㏆ ᖺ ࢟ ࢱ ࣀ ࣓ ࢲ ࢝
Oryzias sakaizumii Asai, Senou & Hosoya, ࡜࣑ࢼ࣑࣓ࢲ࢝ Oryzias latipes㸦Temminck &Schlegel, 㸧ࡢ  ✀࡟ศ㢮ࡉࢀࡓ㸦Asai etal., 㸹 ୰ ᆓ ⦅㸪㸧࠙ ࡇ ࢀ ࡽ  ✀ ࡣ ⣽㇂㸦㸧࡟ᚑ࠸ࠊ௨ୗࠕ㔝⏕࣓ࢲ࢝ࠖ࡜⾲グࡍࡿࠚࠋࡶ࡜ࡶ࡜ࠊỈ⏣ࡸỈ㊰ࠊࡓࡵụ࡞
࡝ேࠎࡢ㌟㏆࡞ሙᡤ࡟ࡶᖜᗈࡃ⏕ᜥࡋ࡚࠸ࡓ
ࡀࠊᅡሙᩚഛࡸ㛤Ⓨ➼ࡢᙳ㡪࡟ࡼࡾ⏕ᜥᩘࡀῶᑡࡋ࡚࠸ࡿࡓࡵࠊ⎔ቃ┬ࣞࢵࢻࣜࢫࢺ࡟࠾࠸࡚㔝⏕࣓ࢲ࢝୧✀ࡣࠊ⤯⁛༴᝹ II 㢮࠙VUࠚ࡟ࡑࢀࡒࢀ㑅ᐃࡉࢀ࡚࠸ࡿ㸦⎔ቃ┬⮬↛⎔ቃᒁ㔝⏕⏕≀ㄢᕼᑡ✀ಖ඲᥎㐍ᐊ㸪Online㸧ࠋࠉ ࣓ ࢲ ࢝ ࡣࠊ࢟ ࣥ ࢠ ࣙ Carassius auratus
㸦Linnaeus 㸧ࡸࢥ࢖ Cyprinus carpio LinnaeusࡢᨵⰋရ✀࡞࡝࡜࡜ࡶ࡟ᅜෆ࡛ྂࡃ࠿ࡽ
ほ㈹㨶࡜ࡋ࡚ࡢᩥ໬ࡀ࠶ࡾࠊࡇࢀࡲ࡛ேᕤⓗ࡞஺㞧ࡸ㑅ᢤࠊ㣫⫱࡟ࡼࡾከࡃࡢᨵⰋရ✀ࡀసฟࡉࢀ࡚࠸ࡿ㸦໭ᕝ㸪㸹ࡵࡔ࠿ࡢ㤋㸪㸧ࠋ㏆ᖺࡣࠊᑓ㛛ᴗ⪅࡟ࡼࡿ኱㔞㣴Ṫࡶ࠶ࡾࠊẚ㍑ⓗᏳ౯࡟㈍኎ࡉࢀࡿࡇ࡜࠿ࡽேࠎ
ࡢ⏕ά࡟ࡼࡾ㌟㏆࡞Ꮡᅾ࡜ࡋ࡚ከࡃᬑཬࡍࡿ
ࡼ࠺࡟࡞ࡗ࡚࠸ࡿࠋࡋ࠿ࡋࠊࡇࢀ࡟క࠸㣫⫱ࡋࡁࢀ࡞ࡃ࡞ࡗࡓಶయࡢ㑇Რࡸࠊඖࠎ㌟㏆࡛࠶ࡗࡓ㔝⏕࣓ࢲ࢝ࡢᏑᅾ࠿ࡽㄗࡗࡓ⎔ቃಖ඲ࡢㄆ㆑࡟ࡼࡾࠊேᕤᨵⰋရ✀ࡢ㔝እᨺὶࡀ࠾
ࡇ࡞ࢃࢀࡿ஦౛ࡀᩓぢࡉࢀࠊᐇ㝿࡟㔝እ࡛ࡢ⛣ධグ㘓ࡶ」ᩘሗ࿌ࡉࢀ࡚࠸ࡿ㸦୰஭࡯࠿㸪㸹℩⬟㸪㸪㸪㸹Nakao et al.,㸹໭ᕝ࡯࠿㸪㸹ఀ⸨࣭ᒣ⏣㸪㸹ᇼỤ࣭ఀ⸨㸪㸧ࠋࡇࢀࡽࡢேᕤᨵⰋရ✀ࡢከࡃࡣࠊ㑇ఏⓗ࡟ᆒ㉁࡞ሙྜࡀከࡃࠊ⛣ධඛࡢ㔝⏕࣓ࢲ࢝࡜ࡢ஺㞧࡟ࡼࡿ㑇ఏⓗከᵝᛶࡢపୗࡀᘬࡁ࠾ࡇࡿࣜࢫࢡࡀ㧗࠸㸦໭ᕝ㸪㸧ࠋࡇࡢࡓࡵࠊࡇࢀࡽࡣ➨  ࡢእ᮶✀࡜࿧ࡤࢀࠊ௒ᚋࡢ኱ࡁ࡞ᙳ㡪ࡀᠱᛕࡉࢀ࡚࠸ࡿ≧ἣୗ࡟࠶ࡿ㸦໭ᕝ㸪㸧ࠋࠉ௒ᅇ➹⪅ࡣࠊᇸ⋢┴ࡢⲨᕝὶᇦෆ࡟఩⨨ࡍࡿᕝཱྀᕷࡢỈᇦ࡟࠾࠸࡚ほ㈹⏝࡟ရ✀ᨵⰋࡉࢀࡓ࡜⪃࠼ࡽࢀࡿ≉␗࡞Ⰽᙬࢆᣢࡘ࣓ࢲ࢝ࢆ᥇㞟ࡋࡓࠋ௒ᚋࡢ㔝እỈᇦ࡬ࡢ᪂ࡓ࡞㑇Რࡸᨺὶ㜵ṆࡢࡓࡵࡢᬑཬၨⓎࡢほⅬࡶྵࡵࠊࡇࡇ࡟ሗ࿌ࡍࡿࠋ᪉ࠉࠉἲࠉ᥇㞟ㄪᰝࡣᖺ  ᭶᪥࡟ᇸ⋢┴ᕝཱྀᕷᱜ⏫࡟఩⨨ࡍࡿࡓࡵụ࡟࠾࠸࡚ࠊᮌ㐨࠿ࡽ┠ど࡛ụෆࢆほᐹࡋ࡞ࡀࡽࠊᓊ㝿ࡢỈ㠃࡟㐟Ὃࡋ࡚ࡁࡓಶయࢆࠊᑠᆺᐜჾࢆ⏝࠸࡚ᥒ࠸ྲྀࡾ
࡟ࡼࡾ⾜ࡗࡓࠋ᥇㞟ಶయࡣࠊFAࡢᕼ㔘Ỉ࡛㯞㓉ᚋࠊ㸣࣐࣍ࣝࣜࣥ⁐ᾮ࡛ᅛᐃࡋࡓࠋྛ㒊ࡢィ ィᩘࡣࠊ୰ᆓ࣭୰ᒣ㸦㸧࡟ࡋࡓࡀࡗࡓࠋィ ࡣࣀࢠࢫࢆ⏝࠸࡚ ੈࡢ⢭ᗘ࡛⾜ࡗࡓࠋᶆ‽య㛗㸦Standard length㸧ࡣSL ࡜⾲グࡋࡓࠋᶆᮏࡣࠊᇸ⋢┴❧⮬↛ࡢ༤≀㤋ࡢ㨶㢮ᶆᮏ㈨ᩱ㸦SMNH-Pi㸧࡜ࡋ࡚ಖ⟶ࡉࢀ࡚࠸ࡿࠋࠉ࡞࠾ࠊ᥇㞟ࡋࡓ࣓ࢲ࢝ࡣヲ⣽࡞✀ุูࡀᅔ㞴࡛࠶ࡗࡓࡓࡵࠊ᭱⤊ⓗ࡟ရ✀ྡࢆุูࡋ
ࡓࠋရ✀ྡࡣࠊఀ⸨࣭ᒣ⏣㸦㸧࡟࠾࠸ุ࡚᩿ᮦᩱ࡜ࡋ࡚⏝࠸ࡽࢀ࡚࠸ࡿ᪥ᮏ࣓ࢲ࢝༠఍ရ✀ศ㢮㒊㛛㸦㸧࡟ᇶ࡙࠸ࡓᙧ
㉁㸦యⰍࠊ㫣ࠊ┠ࠊ⹿Ⰽ⣲⬊ࡢⓎ⌧㒊఩ࠊᶍᵝࠊ㫅ࠊయᆺ࡞࡝㸧ࡢほᐹ࡜ࠊࡵࡔ࠿ࡢ㤋㸦㸧ࡢᥖ㍕ᅗࢆཧ⪃࡜ࡋุู࡚ࡋࠊᙧ㉁࡟ᇶ࡙࠸ࡓయ⣔ⓗရ✀ྡ⛠ࢆグ㍕ࡋࡓࠋᇸ⋢┴ᕝཱྀᕷ࡛☜ㄆࡉࢀࡓほ㈹⏝࣓ࢲ࢝ࠊෆ⏣኱㈗᧜ᙳ⤖ᯝ࡜⪃ᐹࠉᶆᮏ ࣓ࢲ࢝ᒓࡢ  ✀ Oryzias sp.ࠊSMNH-Piࠊ  ಶ యࠊSL ੈࠊᖺ ᭶᪥ࠊᇸ⋢┴ᕝཱྀᕷᱜ⏫ࠊෆ⏣኱㈗࣭㜿㒊┾኱᥇㞟ࠉ᥇㞟ࡉࢀࡓ࣓ࢲ࢝ࡣ  ಶయࡍ࡭࡚࡛ࠊ  㸧యⰍࡣ▖Ꮝࡸ⹿ᙬࠊྛ㫅᮲ྵࡴ඲㌟ࡀ㯮Ⰽ࡛ࠊⓑⰍࡢỈᵴෆ࡟᫬㛫㐟Ὃࡉࡏࡓࡀ⫼ᆅ཯ᛂࡣ࡞࠸ࠊ  㸧㫣ࡣᬑ㏻㫣࡛࠶ࡿࠊ  㸧⫼㫅㌾஌ᩘࡣ  ᮏࠊ 㸧⮌㫅㌾஌ᩘࡣᮏࠊ 㸧యᙧ࡟≉␗ⓗ࡞ᙧ≧ࡣࡳࡽࢀ࡞࠸≉ᚩࢆ᭷ࡋ࡚࠸ࡓࠋグ㘓ಶయࡣࠊࡍ࡭࡚ᅜෆ࡟⏕ᜥࡍࡿ࣓ࢲ࢝ᒓ  ✀࡟ヱᙜࡋࡓᙧែࢆ᭷ࡋ࡚࠸ࡓࡀࠊⰍᙬ࡟᫂ࡽ࠿࡞㐪࠸ࡀࡳࡽࢀࠊࡍ࡭࡚͆ࣈࣛࢵࢡ࣓ࢲ͇࢝࡜࠸࠺㯮Ⰽ⣔⤫ࡢほ㈹⏝ᨵⰋရ✀㸦᪥ᮏ࣓ࢲ࢝༠఍ရ✀ศ㢮㒊㛛㸪㸫㸫ෆ⏣኱㈗ࠉᕝཱྀᕷ࡛᥇㞟ࡉࢀࡓほ㈹⏝࣓ࢲ࢝ࡢグ㘓㸧࡟ヱᙜࡋࡓࠋࠉ ᥇ 㞟 ᫬ࠊ ᮏ ✀ ࡢ ࡯ ࠿ ࡟ ࣔ ࢶ ࢦ
Pseudorasbora parva㸦Temminck & Schlegel㸧ࠊ ≉ ᐃ እ ᮶ ⏕ ≀ ࡢ ࢝ ࢲ ࣖ ࢩ Gambusia
D৽QLVD৽QLV㸦Baird & Girard 㸧ࠊ ࢜ ࢜ ࢡࢳ ࣂ ࢫ Micropterus nigricans㸦Cuvier, 㸧ࡀ☜ㄆࡉࢀࡓࠋ࢝ࢲࣖࢩࡀ᭱ࡶඃ༨ࡋ࡚⏕ᜥࡋ࡚࠾ࡾࠊᮏ✀ࡶࡑࢀ࡟ΰࡌࡿࡼ࠺࡟㐟Ὃࡋ࡚࠸ࡓࠋࠉ௒ᅇ᥇㞟ࡉࢀࡓ࣓ࢲ࢝ࡣࠊ୍⯡ࡢࢽࢵࢡࢿ࣮࣒ࡀࠕࣈࣛࢵࢡࣃࣥࢲࠖࡸࠕ࢜ࣟࢳࠖ࡜࿧ࡤࢀ㈍኎ࡉࢀ࡚࠸ࡿရ✀࡜⪃࠼ࡽࢀࡿࠋࡇࡢရ✀ࡢ≉ᚩ࡜ࡋ࡚ࠊ㯮Ⰽࡢᙉࡉࡀ኱ࡁ࠸࡯࡝㧗౯್࡛࠶ࡿࡼ࠺ࡔࡀࠊᮏሗ࿌࡛᥇㞟ࡉࢀࡓ  ಶయࡣᇶᮏⓗ࡟඲㌟㯮Ⰽ࡛࠶ࡗࡓࠋࡋ࠿ࡋࠊࡵࡔ࠿ࡢ㤋㸦㸧࡟ᥖ㍕ࡉࢀࡓᅗ࡜ẚ㍑ࡍࡿ࡜㯮Ⰽࡀῐ࠸ࡇ࡜࠿ࡽࠊရ✀౯್ࡣప࠸࡜⪃࠼ࡽࢀࡿࠋほ㈹㨶㣴Ṫ࡛ࡣࠊࡇࡢࡼ࠺
࡞పရ㉁ಶయࡣࠊⰍᙬࡸᙧែࢆ⥔ᣢࡍࡿ㑅ู
⾜Ⅽ࡟ࡼࡾ㝖ཤࡀ⾜ࢃࢀࡿࡀࠊఀ⸨࣭ᒣ⏣㸦㸧࡛ࡣࡇࡢ㝖ཤಶయࡢᨺὶࠊ㑇Რࡀほ㈹㨶ࡢ㔝እ⛣ධせᅉࡢ୍ࡘ࡜࡞ࡗ࡚࠸ࡿࡇ࡜
ࡀᣦ᦬ࡉࢀ࡚࠸ࡿࠋ㏆ᖺࠊ࣓ࢲ࢝ࡣಶேឡዲ
ᐙࡢ㣫⫱࡟ࡼࡿရ✀సฟࡸ㣴Ṫ࡟ࡼࡿ㈍኎ࡶ
┒ࢇ࡟࡞ࡗ࡚࠾ࡾ㸦ࡵࡔ࠿ࡢ㤋㸪㸧ࠊᮏሗ࿌ࡢ᥇㞟ᆅࡣఫᏯ⾤ࡢ୍ゅ࡟఩⨨ࡋ࡚࠸ࡿࡇ࡜࠿ࡽࡶࠊಶே㣴Ṫ᫬ࡢ㝖ཤಶయࡀ㑇Რࡉࢀࡓྍ⬟ᛶࡀ㧗࠸ࠋᅜෆ࡛ࡣࠊほ㈹⏝ᨵⰋရ✀࣓ࢲ࢝ࡢ௦⾲᱁࡛࠶ࡿࠕࣄ࣓ࢲ࢝ࠖ⏤᮶ࡢ㑇ఏᏊࡀᗈ࠸⠊ᅖࡢᆅᇦࡢ㔝⏕࣓ࢲ࢝࠿ࡽ᳨ฟࡉࢀ࡚࠾ࡾࠊ㑇ఏᏊởᰁࡀᠱᛕࡉࢀ࡚࠸ࡿ㸦Nakao et al., a, b㸧ࠋࡇࡢࡓࡵࠊ௒ᚋࡣほ㈹㨶㣴Ṫಶయࡢ㑇Რ㐓⬺㜵Ṇ࡬ࡢࡓࡵࡢ㈍኎
᫬ࡢᬑཬ➼ࢆ୍ᒙ㐍ࡵࠊ㔝እ࡬ࡢ౵ධண㜵ࢆᙉࡵ࡚࠸ࡃᚲせࡀ࠶ࡿࡔࢁ࠺ࠋࡲࡓࠊᅾ᮶Ỉᇦ⏕ែ⣔ࡢಖ඲ࡢࡓࡵ࡟ࡶࠊᨵⰋရ✀ࡢ࣓ࢲ࢝ࢆ㣫⫱ࡍࡿឡዲᐙࡢⓙᵝ࡟ࡣࠊᏳ᫆࡞㔝እᨺὶࡣ᥍࠼࡚࠸ࡓࡔࡁࡓ࠸ࠋㅰࠉࠉ㎡ࠉᮏ✏ࢆᇳ➹ࡍࡿ࡟࠶ࡓࡾࠊ㜿㒊┾኱Ặ㸦ࡉ࠸ࡓࡲᕷ㸧࡟ࡣ᥇㞟࡟ࡈ༠ຊ࠸ࡓࡔ࠸ࡓࠋᕝཱྀ㈗ගẶ㸦ओ⎔ቃᣦᶆ⏕≀㸧࡟ࡣᩥ⊩ࢆࡈᥦ౪࠸ࡓࡔ࠸ࡓࠋ༙⏣ᏹఙᏛⱁဨ㸦ᇸ⋢┴❧⮬↛ࡢ༤≀㤋㸧࡟ࡣࠊᶆᮏ཰ⶶࡢ㝿࡟౽ᐅࢆࡣ࠿ࡗ࡚࠸ࡓࡔ࠸ࡓࠋࡇࡢሙࢆ೉ࡾ࡚ཌࡃᚚ♩⏦ࡋୖࡆࡿࠋᘬ⏝ᩥ⊩
Asai, T., H. Senou and K. Hosoya () Oryzias
sakaizumii, a new ricefish from northern Japan(Teleostei: Adrianichthyidae). Ichthyological
Exploration ofFreshwaters, (): 
ᇼỤ┿Ꮚ㸪ఀ⸨ࠉ⋞㸦㸧ᒱ㜧┴ࡢ㔝እỈᇦ࡟࠾ࡅࡿయእග࣓ࢲ࢝㸦ᖿஅ࣓ࢲ࢝㸧࡞࡝ࡢほ㈹㸫㸫㨶࣓ࢲ࢝ࡢᶆᮏ࡟ࡶ࡜࡙ࡃึグ㘓㸬ఀ㇋἟࣭ෆ
἟◊✲ሗ࿌㸪㸸㸬⣽㇂࿴ᾏ㸪ᑠᯘ∾ே㸪໭ᕝᛅ⏕㸦㸧㔝⏕࣓ࢲ࢝ಖㆤ࡬ࡢᥦゝ㸬Ჷ᪉᭷᐀࣭໭ᕝᛅ⏕࣭ᑠᯘ∾ே㸦⦅ⴭ㸧᪥ᮏࡢ㔝⏕࣓ࢲ࢝ࢆᏲࡿ̿ṇࡋࡃ▱ࡗ࡚ṇࡋࡃᏲࡿ㸬pp㸪⏕≀◊✲♫㸪ᮾி㸬ఀ⸨ࠉ⋞㸪ᒣ⏣⏤⣖Ꮚ㸦㸧ឡ▱┴ࡢỈ㊰࠿ࡽᚓࡽࢀࡓࣄ࣓࢝ࣜࢲ࢝➼ࡢほ㈹㨶࣓ࢲ࢝㸬Ichthy,
Natural History of Fishes of Japan.㸪㸸㸬⎔ቃ┬⮬↛⎔ቃᒁ㔝⏕⏕≀ㄢᕼᑡ✀ಖ඲᥎㐍ᐊ
㸦Online㸧⎔ቃ┬ࣞࢵࢻࣜࢫࢺ㸬https://www.
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disturbances in wild medaka populations (Oryzias
latipes species complex) in Japan. IchthyologicalResearch, : Nakao, R., Kano, Y., Iguchi, Y. and T. Kitagawa(b) Genetic disturbance in wild Minami-medaka
populations in the Kyushu region, Japan. International
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