Bull. Natl. Mus. Nat. Sci., Ser. B, 39(2), pp. 61–85, May 22, 2013

Distribution of the Thelypteris japonica Complex(Thelypteridaceae) in Japan
Atsushi Ebihara1,* and Narumi Nakato2
1

Department of Botany, National Museum of Nature and Science, Amakubo 4–1–1,
Tsukuba, Ibaraki 305–0005, Japan
2
Narahashi 1–363, Higashiyamato-shi, Tokyo 207–0031, Japan* E-mail: ebihara@kahaku.go.jp(Received 12 February 2013; accepted 25 March 2013)

Abstract The distribution of the four taxa, Thelypteris japonica forma japonica, T. japonica
forma formosa, T. musashiensis and T. japonica×T. musashiensis, was reassessed by observationof spore morphology of 1984 herbarium specimens deposited in the National Museum of Natureand Science. As a result of frequent changes of identiﬁcation especially between T. japonica forma
formosa and T. musashiensis, the range of each taxon has been drastically updated. The hybrid wasrecorded in 32 prefectures in total, including newly added 25 prefectures.
Key words : distribution, hybrid, spore, Thelypteris.

The Thelypteris japonica complex (Thelypteridaceae) is common understory ferns ranging
throughout Japan except in the Ryukyu Islands.
Japanese members of the complex were recircumscribed by Nakato et al. (2004) based on
cytological and morphological characters, andtwo species, one forma and one interspeciﬁc
hybrid are presently accepted (Fig. 1). Nakato etal. (2004) examined 77 specimens, and indicatedthe approximate geographical and altitudinal distribution of the taxa. Takahashi and Hori (2008)
complemented the distribution data of the taxa
with detailed distribution records in Gifu Prefecture where no species was recorded by Nakato etal. (2004) by observation of spores and grossmorphology on 157 herbarium specimens. However, a complete picture of the distribution of thecomplex in Japan has not yet been provided byprevious studies.
In this study, we reidentﬁed voucher specimens deposited in the herbarium of National
Museum of Nature and Science (TNS) for the
distribution maps created by Kurata and Nakaike(1983) which are probably the largest collectionsin terms of geographical coverage. The most reliable distinguishing character for the species of
the complex is spore morphology as demonstrated by Nakato et al. (2004) (Fig. 2), andtherefore sterile specimens and/or specimens
with only immature spores are difﬁcult to identify.
Materials and Methods
We examined spores of all the specimens of
the complex deposited in TNS. All the specimens
were geocoded manually based on place-namesin their labels. For specimens with castaneous
stipes most of which have been identiﬁed as
T. japonica, spores were preliminary observedwith a microscope in 99.5% ethanol. Specimens
producing only irregular spores were identiﬁed
as the hybrid (T. japonica×T. musashiensis) andthose producing normal-shaped spores withmembraneous ornamentation were identiﬁed as
T. japonica forma japonica. Permanent mountsof spores were prepared with Bioleit (Oken-shojiCo.) and deposited in TNS only for the hybrid.

62

Atsushi Ebihara and Narumi Nakato

Fig. 1. Specimens of the Thelypteris japonica complex in Japan. A. Thelypteris japonica forma japonica (TNSVS-432942, Aichi Pref., Seto-shi, M. Muramatsu 80366); B. T. japonica forma formosa (TNS-VS-1157138,
Yamagata Pref., Kaminoyama-shi, N. Sagawa s.n.); C. T. musashiensis (TNS-VS-1152616, Shizuoka Pref.,Fujinomiya-shi, T. Sato 4091); D. T. japonica×T. musashiensis, the castaneous stipe form (TNS-VS-831722,
Kyoto Pref., Kitakuwada-gun, Miyama-cho, S. Masuda s.n.).

Distribution of the Thelypteris japonica Complex (Thelypteridaceae)

63

Fig. 2. Spores of the Thelypteris japonica complex in Japan (scale bars＝50 μm). A. spores with membraneous
ornamentation produced by T. japonica forma japonica (TNS-VS-397231, Yamaguchi Pref., Iwakuni-shi, A.
Minami 16677); B. spores with membraneous ornamentation produced by T. japonica forma formosa (TNSVS-438920, Fukushima Pref., Onuma-gun, Showa-mura, Z. Kaneda s.n.); C. spores with echinate ornamentation produced by T. musashiensis (TNS-VS-1159828, Oita Pref., Naoiri-gun, Kuju-machi, M. Arakane s.n.);D. irregular-shaped spores of T. japonica×T. musashiensis (TNS-VS-438936, Fukushima Pref., Kitakata-shi,
Z. Kaneda 6517).

For specimens with stramineous stipes (stramineous at least in the upper half of the stipes) whichhave been identiﬁed as T. japonica var.
musashiensis (T. musashiensis), var. glabrata, f.
viridescens and forma formosa, those producing
only irregular spores were identiﬁed as the
hybrid and those producing normal-shaped
spores were separated into two taxa dependingon the morphology of the spore ornamentation(membraneous for T. japonica forma formosaand echinate for T. musashiensis). Specimens,even with abundant irregular spores, were not
identiﬁed as the hybrid when a few regularspores were mixed with the irregular ones. All
the spores collected from specimens with stramineous stipes were mounted with Bioleit andthe permanent mounts were deposited in TNS.

Results and Discussion
Among ca. 2500 specimens examined, mature
spores were observed in 1984 specimens. As aresult, 1193 specimens (60.1%) were identiﬁed
as T. japonica forma japonica, 275 (13.9%) as T.
japonica forma formosa, 344 (17.3%) as T.
musashiensis and 172 (8.7%) as the T. japonica×
T. musashiensis (Appendix 1). It is notable thatthe hybrid consists of individuals with castaneous stipes (120, 69.8% of the hybrid) and thosewith stramineous stipes (52, 30.2%).
Thelypteris japonica forma japonica is themost widespread and most common taxon in thecomplex, ranging throughout Japan except in
Hokkaido and Okinawa Prefectures (Fig. 3A).
Thelypteris japonica forma formosa is also widespread, extending to the southern part of Hok-

64

Atsushi Ebihara and Narumi Nakato

Fig. 3. Distribution maps of the Thelypteris japonica complex in Japan based on the specimens examined by thepresent study. A. Thelypteris japonica forma japonica; B. T. japonica forma formosa.

Distribution of the Thelypteris japonica Complex (Thelypteridaceae)

65

Fig. 4. Distribution maps of the Thelypteris japonica complex in Japan based on the specimens examined by thepresent study. A. T. musashiensis; B. T. japonica×T. musashiensis.

66

Atsushi Ebihara and Narumi Nakato

kaido, but less common than the type forma (Fig.
3B). Thelypteris musashiensis prefers rocky habitats at rather high altitudes (Nakato et al., 2004),
and tends to be distributed in inland areas fromsouthern Hokkaido to Kyushu; it has not yet beenfound in 8 prefectures, i.e. Chiba, Kanagawa,Shimane, Yamaguchi, Kagawa, Nagasaki, Kumamoto and Okinawa (Fig. 4A). The hybrid (T.
japonica×T. musashiensis) has been found in
most of the prefectures where distributions of T.
japonoica s.l. and T. musashiensis overlap, but
not yet been found in Hokkaido, Mie, Osaka,
Nara, Wakayama, Tokushima, Fukuoka, Saga,
Kumamoto, Miyazaki and Kagoshima Prefectures (Fig. 4B). It is also notable that the hybrid
occurs in Chiba, Kanagawa and Shimane Prefectures where one of the parents, T. musashiensis
has not yet been recorded. Takahashi and Hori
(2008) already pointed out that hybrids were
often recorded in the localities lacking one orboth of their parents, and our present results frequently showed a similar phenomenon whenlocalities smaller than prefectures were taken
into account. Although insufﬁcient sampling isprobably a cause of this phenomenon, some of
the localities of the hybrid are apparently not preferred by T. musashiensis (e.g. TNS-VS-517192,
Tokyo, Ikegami — located in the city center ofTokyo, almost at sea level). Variation in stipe
color of the hybrid is presumably due to the stipe
color of one of the parents (i.e. T. japonica forma
japonica vs. forma formosa). The hybrid withstramineous stipes has been collected in thenortheastern half of Honshu so far, which seems
to be inﬂuenced by the distribution of T. japonica
forma formosa.
Acknowledgments
We are grateful to K. Hamasaki for her help
with specimen databasing.

Tokyo (in Japanese).
Nakato, N., Sahashi, N. and Kato, M. 2004. Cytotaxonomy of the Thelypteris japonica complex (Thelypteridaceae). Acta Phytotaxonomica et Geobotanica. 55:
89–105.
Takahashi, H. and Hori, S. 2008. Distribution of the
Thelypteris japonica complex (Thelypteridaceae) in
Gifu Prefecture, Japan. Bulletin of the Gifu BotanicalSociety 24: 89–94 (in Japanese).

Appendix 1. Specimens identiﬁed based on spore
morphology. Municipal-level locality information isprovided for Thelypteris japonica forma japonica, andlocality details are provided for the remaining taxa.
Thelypteris japonica (Baker) Ching forma japonica [Harigane-warabi]
Honshu - Aomori Pref.: Higashitsugaru-gun, Kanita-machi, 1977-822, N. Saitoh s.n. (TNS-VS-344145); Iwate Pref.: Shiwa-gun,Tonan-mura, 1980-9-15, Y. Nakajima s.n. (TNS-VS-440259);
Miyagi Pref.: Sendai-shi, 1912-9-28, E. Iishiba s.n. (TNSVS-254666), 1980-9-23, M. Takeichi s.n. (TNS-VS-575758); Ishinomaki-shi, 1972-10-15, Y. Ueno 8550 (TNS-VS-575754);
Shiogama-shi, 1977-8-22, M. Takeichi s.n. (TNS-VS-575762);
Shibata-gun, Shibata-machi, 1980-10-11, K. Shoji s.n. (TNSVS-575751); Igu-gun, Marumori-machi, 1980-8-10, K. Okazakis.n. (TNS-VS-576835), 1980-7-22, K. Okazaki s.n. (TNSVS-576834), 1981-8-23, K. Matsunaga s.n. (TNS-VS-1156160);
Miyagi-gun, Matsushima-machi, 1976-8-19, W. Takahashi s.n.
(TNS-VS-576832); Rifu-cho, 1980-10-26, M. Takeichi s.n. (TNSVS-575756); Shida-gun, Matsuyama-machi, 1977-?-?, M. Takeichis.n.

(TNS-VS-575761);

Tamatsukuri-gun,

Iwadeyama-machi,

1972-10-1, W. Takahashi s.n. (TNS-VS-576833); Mono-gun,Yamoto-cho, 1977-10-15, M. Takeichi s.n. (TNS-VS-575763);
Oshika-gun, Onagawa-cho, 1981-9-23, M. Takeichi s.n. (TNSVS-575767); Inai-cho, 1967-10-22, M. Kikuchi s.n. (TNSVS-177138); Oshika-cho, 1979-11-4, K. Matsunaga s.n. (TNSVS-1156161); Motoyoshi-gun, Shizugawa-cho, 1980-9-19, M.
Takeichi s.n. (TNS-VS-575760); Motoyoshi-cho, 1980-7-26, Z.
Kaneda s.n. (TNS-VS-575759); Akita Pref.: Akita-shi, 1978-9-5,
J. Takada s.n. (TNS-VS-420027); Omagari-shi, 1980-10-10, J.
Takada s.n. (TNS-VS-420029); Yuri-gun, Kisakata-machi, 1980-928, J. Takada s.n. (TNS-VS-420026); Senboku-gun, Kyowamachi, 1976-10-5, J. Takada s.n. (TNS-VS-420028); Ogachi-gun,Ogachi-machi, 1981-9-26, J. Takada s.n. (TNS-VS-420111);
Yamagata Pref.: Yonezawa-shi, 1980-8-28, Z. Sato 326 (TNSVS-437984); Tsuruoka-shi, 1958-10-9, S. Kato s.n. (TNSVS-437817), 1981-8-6, A. Yamamoto s.n. (TNS-VS-437980),
1970-8-14, S. Kato s.n. (TNS-VS-437612); Murayama-shi, 19808-24, N. Sagawa s.n. (TNS-VS-437982); Mogami-gun, Tozawa-

Referencesmura, 1964-9-12, S. Kato s.n. (TNS-VS-437819); Higashitagawa-

Kurata, S. and Nakaike, T. 1983. Illustrations of Pteridophytes of Japan, vol. 3. The University of Tokyo Press,

Nishitagawa-gun, Tagawa-mura, 1892-9-11, T. Nagasawa s.n.

gun, Haguro-machi, 1980-10-1, T. Nakaike s.n. (TNS-VS-437981);(TNS-VS-54786), 1903-8-24, T. Nagasawa s.n. (TNS-VS-517186,

Distribution of the Thelypteris japonica Complex (Thelypteridaceae)

67

517188); Fukushima Pref.: Iwaki-shi, 1976-8-8, H. Ito s.n. (TNS-s.n. (TNS-VS-799754), 1987-9-28, T. Kamiyama s.n. (TNS-

VS-1092657), 1973-10-16, Y. Yuzawa s.n. (TNS-VS-438917),

VS-799741), 1987-7-26, T. Kamiyama s.n. (TNS-VS-799753),

1975-9-28, Y. Yuzawa s.n. (TNS-VS-438919), 1976-7-3, Y. Yuzawa

1987-8-17, T. Kamiyama s.n. (TNS-VS-799740), 1980-9-17, M.

s.n.

(TNS-

Usui s.n. (TNS-VS-422705), 1980-9-17, M. Usui s.n. (TNS-

VS-438939), 1976-8-9, T. Nakaike s.n. (TNS-VS-435492), 1984-

VS-422706); Oyama-shi, 1980-9-12, M. Usui s.n. (TNS-

11-14, M. Origasa s.n. (TNS-VS-1157005), 1975-9-28, Z. Kaneda

VS-422713); Otawara-shi, 1957-9-22, K. Ogawa s.n. (TNS-s.n. (TNS-VS-1172237); Shinobu-gun, Tsuchiyu-mura, 1933-8-21,

VS-1042048);

H. Koidzumi 100833 (TNS-VS-912107); Onuma-gun, Aizutakada-(TNS-VS-431947),

machi, 1980-9-21, A. Baba s.n. (TNS-VS-439165); Higashishi-

VS-1157006); Kawachi-gun, Minamikawachi-machi, 1979-10-20,

rakawa-gun, Tanagura-machi, 1977-10-20, Z. Kaneda 3503 (TNS-

M. Usui s.n. (TNS-VS-422717); Kamikawachi-mura, 1981-10-2,

VS-1172238); Futaba-gun, Naraha-machi, 1975-10-20, Z. Kaneda

S. Waku s.n. (TNS-VS-438798); Kawachi-machi, 1987-9-13, H.

s.n. (TNS-VS-438933), 1969-9-15, Y. Yuzawa s.n. (TNS-

Ogawa s.n. (TNS-VS-799755), 1987-10-4, H. Ogawa s.n. (TNS-

VS-1172236); Tomioka-machi, 1980-12-2, Z. Kaneda 6512 (TNS-

VS-799744); Kamitsuga-gun, Nishikata-mura, 1981-9-8, M. Usui

VS-438935), 1980-12-3, Z. Kaneda 6505 (TNS-VS-438934);

s.n. (TNS-VS-427386); Haga-gun, Mashiko-machi, 1979-9-20, M.

Ibaraki Pref.: Mito-shi, 1979-8-23, M. Yasu s.n. (TNS-

Usui s.n. (TNS-VS-422716); Motegi-machi, 1987-9-13, M.

VS-422597); Ishioka-shi, 1974-10-10, H. Fujita s.n. (TNS-

Yoshikawa 1521 (TNS-VS-799746), 1980-10-2, M. Usui s.n.

VS-422809); Ryugasaki-shi, 1975-8-27, Y. Nakazaki & N. Wada

(TNS-VS-422708); Shimotsuga-gun, Fujioka-machi, 1981-8-17,

s.n. (TNS-VS-422586); Hitachiota-shi, 1980-9-7, M. Yasu s.n.

M. Usui s.n. (TNS-VS-518255); Shioya-gun, Shioya-machi, 1980-(TNS-VS-422594); Kitaibaraki-shi, 1974-10-5, N. Wada 1051

9-8, T. Tashiro s.n. (TNS-VS-422694); Ujiie-machi, 1987-9-13, H.

(TNS-VS-595307), 1975-11-18, T. Waku 4295 (TNS-VS-595305);

Ogawa s.n. (TNS-VS-799739); Kitsuregawa-machi, 1981-9-2, M.

Kasama-shi, 1975-8-8, H. Fujita s.n. (TNS-VS-422811); Toride-

Usui s.n. (TNS-VS-427390), 1987-10-28, S. Waku s.n. (TNS-shi, 1975-8-27, M. Yasu s.n. (TNS-VS-422805), 1975-8-22, H.

VS-1160528); Nasu-gun, Karasuyama-machi, 1977-8-28, T. Waku

Fujita s.n. (TNS-VS-422808); Tsukuba-shi, 1995-11-11, N. Nakato

7418 (TNS-VS-422690); Bato-machi, 1981-9-16, M. Usui s.n.

2009 (TNS-VS-1168523), 1995-11-11, N. Nakato 2014 (TNS-(TNS-VS-427389); Kurobane-machi, 1981-9-30, S. Waku s.n.

VS-1168524), 1997-9-22, S. Fujimoto s.n. (TNS-VS-9508642a/b);(TNS-VS-438797); Nasu-machi, 1968-8-22, H. Ito s.n. (TNS-

Higashiibaraki-gun, Ibaraki-machi, 1974-10-5, M. Yasu s.n. (TNS-

VS-1092234), 1980-10-15, M. Usui s.n. (TNS-VS-422703), 1981-

VS-422588); Nishiibaraki-gun, Iwama-machi, 1958-10-31, E.

8-16, T. Tashiro s.n. (TNS-VS-427392), 1980-8-13, M. Usui s.n.

Sugiyama s.n. (TNS-VS-1042044); Naka-gun, Ogawa-mura, 1974-(TNS-VS-422704); Nishinasuno-machi, 1987-10-8, T. Yamamura

11-8, H. Fujita s.n. (TNS-VS-422589); Kuji-gun, Satomi-mura,s.n. (TNS-VS-799756), 1988-11-13, T. Yamamura s.n. (TNS-

1979-9-5, M. Yasu s.n. (TNS-VS-422580); Daigo-machi, 1970-10-

VS-799738), 1968-8-22, H. Ito s.n. (TNS-VS-1042053), 1979-9-8,

18, Y. Nakazaki s.n. (TNS-VS-422585), 1977-10-30, M. Yasu s.n.

H. Kato s.n. (TNS-VS-422688); Gunma Pref.: Ota-shi, 1973-10-(TNS-VS-422815); Kashima-gun, Kashima-machi, 1974-11-30,

23, S. Suto s.n. (TNS-VS-435554); Seta-gun, Ogo-machi, 1980-11-

M. Yasu s.n. (TNS-VS-422807); Niihari-gun, Yasato-machi, 1973-

4, T. Waku 10466 (TNS-VS-435551); Gunma-gun, Haruna-machi,

10-31, M. Yasu s.n. (TNS-VS-422574), 1979-10-25, M. Yasu s.n.

1980-6-23, S. Suto s.n. (TNS-VS-435553); Kitagunma-gun, Ono-

(TNS-VS-422582); Tsukuba-gun, Tsukuba-machi, 1980-12-18, T.

gami-mura, 1985-10-2, S. Waku s.n. (TNS-VS-1166304); Tano-

Nakamura & A. Nakamura s.n. (TNS-VS-422802), 1980-11-20, T.

gun, Onishi-machi, 1979-9-17, S. Suto s.n. (TNS-VS-435552);

Nakamura & A. Nakamura s.n. (TNS-VS-422801); Makabe-gun,Kanra-gun, Shimonita-machi, 1956-9-9, T. Satomi 16 (TNS-A.

(TNS-VS-

VS-1042065); Usui-gun, Matsuida-machi, 1986-8-3, S. Ishiguro

9508621a/b); Yuuki-gun, Yachiyo-machi, 1975-8-22, Y. Nakazakis.n. (TNS-VS-1156206); Ora-gun, Chiyoda-mura, 1957-11-0, T.

& O. Shimizu s.n. (TNS-VS-422587); Sashima-gun, Goka-mura,

Matsuzawa s.n. (TNS-VS-1042047); Tatara-mura, 1954-8-22, T.

1975-8-31, N. Wada s.n. (TNS-VS-422599); Tochigi Pref.: Utsu-

Wakana 27 (TNS-VS-1042124); Saitama Pref.: Urawa-shi, 1947-

nomiya-shi, 1987-8-5, F. Matsumura s.n. (TNS-VS-799751),

9-25, T. Kawasaki 6767 (TNS-VS-675379), 1980-7-12, T. Waku

1981-9-5, M. Usui s.n. (TNS-VS-427387), 1981-8-29, M. Usui s.n.

10032 (TNS-VS-514508); Omiya-shi, 1980-9-5, T. Waku 10243(TNS-VS-427385), 1980-8-28, M. Usui s.n. (TNS-VS-422709),(TNS-VS-514507); Chichibu-shi, 1994-10-25, T. Iwata 40606

1980-9-28, M. Usui s.n. (TNS-VS-422711); Ashikaga-shi, 1980-(TNS-VS-1133844),

11-3, M. Usui s.n. (TNS-VS-422698), 1980-10-27, M. Usui s.n.

VS-1133835), 2000-8-10, T. Iwata 61054 (TNS-VS-1133820),(TNS-VS-422699); Tochigi-shi, 1980-10-24, M. Usui s.n. (TNS-

1987-11-17, T. Iwata 8334 (TNS-VS-1133723), 1989-10-29, T.

VS-422718); Kanuma-shi, 1980-10-23, K. Satoh s.n. (TNS-

Iwata 9712 (TNS-VS-1133871), 1991-8-25, T. Iwata 10515 (TNS-(TNS-VS-438918),

Makabe-machi,

1980-10-26,

1998-8-18,

Satoh

S.

Saito

TKB0021s.n.

Kuroiso-shi,

1981-9-15,

1987-9-12,

1995-10-1,

S.

T.

T.

Ishiguro

Iwata

Waku

s.n.

s.n.

(TNS-

42159

(TNS-

VS-799737), 1980-10-9, M. Usui s.n. (TNS-VS-422712), 1980-9-

VS-1133860), 1991-9-15, T. Iwata 10566 (TNS-VS-1133859),

1, M. Usui s.n. (TNS-VS-422714), 1980-10-29, M. Usui s.n.

1991-10-6, T. Iwata 10660 (TNS-VS-1133858), 1983-9-15, T.

(TNS-VS-422700); Nikko-shi, 1987-8-9, T. Kamiyama s.n. (TNS-

Iwata 2088 (TNS-VS-1133788), 1983-9-15, T. Iwata 2094 (TNS-

VS-799743), 1987-8-9, T. Kamiyama s.n. (TNS-VS-799742),

VS-1133787), 1983-10-28, T. Iwata 2338 (TNS-VS-1133782),

1956-9-10, M. Ishihara s.n. (TNS-VS-132660), 1980-10-22, M.

1983-10-28, T. Iwata 2339 (TNS-VS-1133781), 1964-8-18, T.

Usui s.n. (TNS-VS-422707); Imaichi-shi, 1987-8-20, T. Kamiyama

Iwata 289 (TNS-VS-1133810), 1981-8-25, S. Kobayashi s.n.

68Atsushi Ebihara and Narumi Nakato(TNS-VS-514514); Hanno-shi, 1993-10-3, T. Yamada s.n. (TNS-

VS-1133881), 1991-7-31, T. Iwata 10355 (TNS-VS-1133861),

VS-737856), 1986-11-9, T. Iwata 7295 (TNS-VS-1133731), 1985-

1969-11-24, T. Iwata 293 (TNS-VS-1133806); Higashichichibu-

11-20, T. Iwata 5909 (TNS-VS-1133745), 1983-10-9, T. Iwatamura, 1996-9-16, T. Iwata 46861 (TNS-VS-1133833), 1988-9-8, T.

2183 (TNS-VS-1133785), 1983-10-9, T. Iwata 2166 (TNS-

Iwata 8886 (TNS-VS-1133902), 1988-9-8, T. Iwata 8883 (TNS-

VS-1133783), 1984-11-10, T. Iwata 3885 (TNS-VS-1133765),

VS-1133903), 1985-9-23, T. Iwata 5523 (TNS-VS-1133754),

1958-11-16, H. Ito s.n. (TNS-VS-1042042), 1975-9-28, I. Ueda

1985-9-23, T. Iwata 5514 (TNS-VS-1133753), 1985-11-1, T. Iwata

s.n. (TNS-VS-1172270); Honjo-shi, 1981-7-13, S. Kojima s.n.

5784 (TNS-VS-1133749), 1982-11-5, T. Iwata 1442 (TNS-(TNS-VS-514513); Higashimatsuyama-shi, 1990-7-14, T. Iwata

VS-1133793); Kodama-gun, Kamiizumi-mura, 1983-11-14, S.

9834 (TNS-VS-1133869), 1990-10-29, T. Iwata 10081 (TNS-

Kojima & K. Mashimo s.n. (TNS-VS-1156149); Osato-gun, Yorii-

VS-1133863), 1981-8-24, S. Kobayashi s.n. (TNS-VS-514502);

machi, 1990-10-7, T. Iwata 10037 (TNS-VS-1133864), 1981-11-

Iwatsuki-shi, 1980-7-12, T. Waku 10051 (TNS-VS-514496);

22, S. Kobayashi s.n. (TNS-VS-425195); Kitasaitama-gun,

Sayama-shi, 1981-11-20, S. Kobayashi s.n. (TNS-VS-425193);

Okuwa-mura, 1952-8-?, I. Nagano s.n. (TNS-VS-1042072); Kita-

Iruma-shi, 1981-8-2, S. Kobayashi s.n. (TNS-VS-514499);

katsushika-gun, Washimiya-machi, 1964-9-12, H. Ito s.n. (TNS-

Sakado-shi, 1980-8-28, S. Kobayashi s.n. (TNS-VS-514504);

VS-1045693); Showa-machi, 1958-10-15, F. Nagasu s.n. (TNS-

Iruma-gun, Moroyama-machi, 2002-12-5, T. Iwata 67378 (TNS-

VS-162717); Chiba Pref.: Kisarazu-shi, 1987-10-7, M. Ono s.n.

VS-1133814), 1988-11-28, T. Iwata 9118 (TNS-VS-1133896),(TNS-VS-1023798); Matsudo-shi, 1979-11-12, Y. Kimizuka s.n.

1988-11-28, T. Iwata 9110 (TNS-VS-1133895), 1984-10-19, T.

(TNS-VS-424253); Sawara-shi, 1981-8-16, K. Ito s.n. (TNS-

Iwata 3673 (TNS-VS-1133770), 1965-11-7, H. Ito s.n. (TNS-

VS-560436), 1981-8-6, Y. Koike s.n. (TNS-VS-560434); Togane-

VS-1045364); Ogose-machi, 1989-9-14, T. Iwata 9554 (TNS-shi, 1976-9-20, I. Ueda s.n. (TNS-VS-424245); Katsuura-shi,

VS-1133879), 1989-9-14, T. Iwata 9560 (TNS-VS-1133877),

1981-8-11, Y. Kimizuka s.n. (TNS-VS-560438); Ichihara-shi, 1978-

1989-9-14, T. Iwata 9542 (TNS-VS-1133878); Umesono-mura,

8-22, Y. Kimizuka s.n. (TNS-VS-424251), 1980-8-28, T. Kawana

1952-11-?, H. Ito s.n. (TNS-VS-1042113), 1952-11-?, H. Ito s.n.

s.n. (TNS-VS-424254); Abiko-shi, 1980-10-10, K. Aida s.n. (TNS-(TNS-VS-1042075, 1042076, 1042078); Naguri-mura, 1989-9-1,

VS-424255), 1979-10-14, T. Kawana s.n. (TNS-VS-424242);

T. Iwata 9438 (TNS-VS-1133880), 1985-11-20, T. Iwata 5918

Kimitsu-shi, 1983-9-15, T. Iwata 2266 (TNS-VS-1133789), 1972-

T.

Iwata

3750

(TNS-

11-12, T. Nakaike s.n. (TNS-VS-288360), 1977-12-3, H.

VS-1133769), 1984-10-19, T. Iwata 3740 (TNS-VS-1133768),

Kobayashi s.n. (TNS-VS-424249), 1980-8-20, Y. Kasuya s.n.

(TNS-VS-1133746),

1984-10-19,

1964-10-25, H. Ito s.n. (TNS-VS-1045361), 2002-9-1, N. Nakato(TNS-VS-424239), 1980-8-18, T. Kawana s.n. (TNS-VS-427514),

2467 (TNS-VS-1168489), 1976-10-17, H. Kobayashi s.n. (TNS-

1980-8-20, Y. Kasuya s.n. (TNS-VS-936021); Yotsukaido-shi,

VS-1172268); Agano-mura, 1953-9-?, H. Inoue s.n. (TNS-

1983-12-12, J. Haginiwa JH014529 (TNS-VS-964529), 1983-12-

VS-1042073); Yamane-mura, 1938-9-18, S. Okuyama 17992

12, J. Haginiwa JH014530 (TNS-VS-964530); Higashikatsushika-H.

gun, Sekiyado-machi, 1981-10-11, Y. Kimizuka s.n. (TNS-

Ohashi, H. Ohba, Y. Tateishi & J. Murata 540 (TNS-VS-337052);

VS-560437); Katori-gun, Kurimoto-machi, 1976-9-19, Y. Koike

Tokigawa-mura, 1984-11-17, T. Iwata 3917 (TNS-VS-1133764),

283 (TNS-VS-424247); Chosei-gun, Chosei-mura, 1966-8-15, M.

1993-9-26, T. Yamada s.n. (TNS-VS-737854); Tamagawa-mura,

Furuse 44360 (TNS-VS-773906); Awa-gun, Amatsukominato-

1989-9-14, T. Iwata 9526 (TNS-VS-1133876), 1989-9-14, T. Iwata

machi, 1951-11-0, H. Ito s.n. (TNS-VS-1042120); Tokyo Pref.:9515 (TNS-VS-1133875), 1989-9-14, T. Iwata 9527 (TNS-Tokyo-shi, 1932-10-0, K. Hiyama 457 (TNS-VS-1166299); Bun-

VS-1133873); Chichibu-gun, Yokoze-machi, 1987-8-23, T. Iwatakyo-ku, 1980-7-12, Y. Kimizuka s.n. (TNS-VS-423948); Kita-

7956 (TNS-VS-1133718), 1981-8-19, T. Iwata s.n. (TNS-toshima-gun, Iwabuchi-machi, 1929-7-?, Y. Jotani 16 (TNS-

VS-1133802), 1982-11-3, T. Iwata 1397 (TNS-VS-1133794);

VS-831884), 1897-8-17, K. Watanabe s.n. (TNS-VS-63268);

Minano-machi, 1985-11-6, T. Iwata 5870 (TNS-VS-1133747);

Hachioji-shi, 2008-8-11, S. Fujimoto 080811-2 (TNS-VS-776368),(TNS-VS-281549);

Hiki-gun,Ogawa-machi,

1974-11-1,

Nagatoro-machi, 1996-11-6, T. Iwata 48175 (TNS-VS-1133830),

1970-9-29, H. Miyazaki 1679 (TNS-VS-831717), 1969-10-6, S.

2001-11-9, T. Iwata 64425 (TNS-VS-1133816), 1988-10-9, T.

Serizawa 11274 (TNS-VS-311877); Ome-shi, 1985-9-29, T. Iwata

Iwata 9017 (TNS-VS-1133900), 1983-8-26, T. Iwata 2031 (TNS-

5587 (TNS-VS-1133752), 1984-10-23, T. Iwata 3773 (TNS-

VS-1133790), 1984-7-23, T. Iwata 3071 (TNS-VS-1133777);

VS-1133766), 1976-9-5, C. Ohkawa s.n. (TNS-VS-435957), 2003-

Yoshida-machi, 1988-7-28, T. Iwata 8676 (TNS-VS-1133907),

7-20, N. Nakato 2507 (TNS-VS-1168490), 1980-9-14, H.

1982-8-18, T. Iwata 1220 (TNS-VS-1133795); Ogano-machi,

Kobayashi s.n. (TNS-VS-424054), 1980-10-4, H. Endo s.n. (TNS-

1995-9-18, T. Iwata 42059 (TNS-VS-1133840), 1988-9-25, T.

VS-423942); Fuchu-shi, 1980-9-19, M. Mitsuhashi s.n. (TNS-

Iwata 8990 (TNS-VS-1133901), 2002-11-17, T. Iwata 67354

VS-424053), 1980-9-30, M. Mitsuhashi s.n. (TNS-VS-423950);(TNS-VS-1133813); Ryokami-mura, 1993-9-25, T. Iwata 11332

Akishima-shi, 1980-10-6, H. Endo s.n. (TNS-VS-424055);

T.

Iwata

41811

(TNS-

Kitatama-gun, Jindai-mura, 1949-11-19, Y. Hiratsuka s.n. (TNS-

VS-1133841), 1990-7-22, T. Iwata 9868 (TNS-VS-1133868);

VS-513626); Chofu-shi, 1955-11-10, T. Kawasaki s.n. (TNS-

Otaki-mura, 1950-11-15, H. Ito s.n. (TNS-VS-1042067); Arakawa-

VS-1042081), 1980-10-31, M. Mitsuhashi s.n. (TNS-VS-427147);

mura, 1989-8-1, T. Iwata 9338 (TNS-VS-1133890), 1989-8-1, T.

Machida-shi, 1980-9-30, A. Yamamoto s.n. (TNS-VS-423949),

Iwata 9340 (TNS-VS-1133886), 1989-8-4, T. Iwata 9319 (TNS-

1980-10-17, C. Azegami s.n. (TNS-VS-936406, 936477); Higashi-(TNS-VS-1133848),

1995-8-31,

Distribution of the Thelypteris japonica Complex (Thelypteridaceae)
yamato-shi, 2002-8-16, N. Nakato 2828 (TNS-VS-1168488);
H.

Endo

s.n.

69

VS-1156201); Nakakanbara-gun, Muramatsu-machi, 1962-8-29, F.

(TNS-

Konta 911 (TNS-VS-929589), 1948-10-?, T. Uchikawa 41 (TNS-

VS-424052); Akiruno-shi, 2003-7-20, N. Nakato 2513 (TNS-

VS-89864), 1978-9-7, M. Saito 2554 (TNS-VS-543169), 1979-10-

VS-1168491); Nishitama-gun, Itsukaichi-machi, 1980-9-1, H.

15, T. Takahashi s.n. (TNS-VS-543179), 1983-8-30, M. Saito s.n.

Endo s.n. (TNS-VS-424058), 1974-10-20, Y. Kimizuka s.n. (TNS-(TNS-VS-1156157), 1967-8-30, S. Iwano s.n. (TNS-VS-1172235),

VS-424057); Hinohara-mura, 1988-8-20, K. Inoue s.n. (TNS-

1978-9-23, M. Saito 2177 (TNS-VS-1172234), 1978-9-23, M.

VS-831721), 1935-9-3, T. Nomura s.n. (TNS-VS-1017544), 1966-

Saito 2181 (TNS-VS-1172233); Nishikanbara-gun, Yahiko-mura,

7-26, K. Saito s.n. (TNS-VS-1046702), 1980-11-15, H. Kobayashi

1978-8-3, M. Saito 2089 (TNS-VS-543166); Bunsui-machi, 1987-

s.n. (TNS-VS-424056); Okutama-machi, 1972-8-18, M. Sakakibara

10-22, M. Saito s.n. (TNS-VS-1156125); Minamikanbara-gun,s.n. (TNS-VS-309785), 1957-8-18, T. Takano s.n. (TNS-

Shitada-mura, 1979-8-27, M. Saito 3192 (TNS-VS-543173);

VS-134105), 1967-11-5, H. Ito s.n. (TNS-VS-1045687), 1971-8-

Sakae-machi, 1987-10-23, M. Saito s.n. (TNS-VS-1156133); Higa-

29, T. Iwasaki 2702 (TNS-VS-831889), 1977-9-23, K. Inoue s.n.

shikanbara-gun, Mikawa-mura, 1943-8-18, S. Iwano s.n. (TNS-(TNS-VS-831894, 831895), 1981-10-19, M. Mitsuhashi s.n. (TNS-

VS-543178); Santo-gun, Koshiji-machi, 1985-10-21, I. Ohzu s.n.

VS-427148), 1980-10-2, M. Mitsuhashi s.n. (TNS-VS-424059),(TNS-VS-1166307); Teradomari-machi, 1987-10-23, M. Saito s.n.

1984-10-13, E. Fukushi s.n. (TNS-VS-1157034); Mita-mura,(TNS-VS-1156121); Minamiuonuma-gun, Shiozawa-machi, 1958-

1932-10-19, T. Satow s.n. (TNS-VS-40798); Hikawa-mura, 1940-

11-3, S. Iwano s.n. (TNS-VS-543177); Kariwa-gun, Takayanagi-

8-26, R. Inoue s.n. (TNS-VS-89887); Mikurajima-mura, 1967-7-

machi, 1953-10-19, S. Iwano s.n. (TNS-VS-543174), 1985-8-14, I.

28, Y. Jotani & al. s.n. (TNS-VS-831703); Kamitsuki-mura, 1934-

Ohzu s.n. (TNS-VS-1166306); Nakakubiki-gun, Kakizaki-machi,

8-9, Y. Jotani 5495 (TNS-VS-831903), 1934-8-8, Y. Jotani 5497

1962-11-20, E. Sakuma s.n. (TNS-VS-543190); Kubiki-mura,(TNS-VS-121089); Toyotama-gun, Iogi-machi, 1931-9-?, F.

1984-10-20, S. Tomitori s.n. (TNS-VS-1157004); Iwafune-gun,

Maekawa s.n. (TNS-VS-831868); Kanagawa Pref.: Yokohama-Sekikawa-mura, 1987-9-29, M. Saito s.n. (TNS-VS-1156124);

shi, 1952-8-9, O. Deguchi s.n. (TNS-VS-92880), 1980-8-24, T.

Sanpoku-machi, 1987-9-28, M. Saito s.n. (TNS-VS-1156126);

Sato s.n. (TNS-VS-441341), 1980-9-7, E. Fukushi s.n. (TNS-

Sado-gun, Aikawa-machi, 1983-10-28, M. Saito s.n. (TNS-

VS-441345), 1981-10-11, A. Yamamoto s.n. (TNS-VS-427169);

VS-1156159), 1986-9-9, M. Saito s.n. (TNS-VS-1157043); Kanai-

Kawasaki-shi Tama-ku, 1979-10-14, Y. Kimizuka s.n. (TNS-

machi, 1983-10-10, s.n. (TNS-VS-1156156); Mano-machi, 1987-

VS-441339); Asao-ku, 1982-11-21, E. Fukushi s.n. (TNS-

11-15, M. Saito s.n. (TNS-VS-1156123); Hamochi-machi,

VS-1156204); Hiratsuka-shi, 1971-8-24, H. Komaki s.n. (TNS-

1995-10-24, N. Nakato 1991 (TNS-VS-1168525); Akadomari-VS-287216), 1970-7-26, J. Moriya s.n. (TNS-VS-280517),

mura, 1984-8-9, M. Saito s.n. (TNS-VS-1157037); Toyama Pref.:1974-8-29, J. Moriya s.n. (TNS-VS-441340), 1971-9-25, J.

Toyama-shi, 1978-7-6, T. Ohshima s.n. (TNS-VS-528751); Taka-

Moriya s.n. (TNS-VS-441342); Odawara-shi, 1980-11-10, M. Aki-oka-shi, 1977-8-9, T. Ohshima s.n. (TNS-VS-528748); Uozu-shi,yama s.n. (TNS-VS-545685), 1982-11-20, K. Tanaka s.n. (TNS-

1981-8-20, T. Ohshima s.n. (TNS-VS-528744), 1977-9-23, T.

VS-1156215); Ashigarakami-gun, Yamakita-machi, 1983-8-27, K.

Ohshima s.n. (TNS-VS-528746); Kurobe-shi, 1978-8-10, T.

Tanaka s.n. (TNS-VS-1156164); Ashigarashimo-gun, Hakone-Ohshima s.n. (TNS-VS-528752); Tonami-shi, 1976-8-21, T.

machi, 1981-9-15, Y. Kimizuka s.n. (TNS-VS-427156); Yugawara-

Ohshima s.n. (TNS-VS-528755); Oyabe-shi, 1979-8-9, T. Ohshimamachi, 1981-11-15, T. Sato s.n. (TNS-VS-438991), 1981-11-3, H.

s.n. (TNS-VS-528753); Kaminiikawa-gun, Oyama-machi, 1983-8-

Kobayashi s.n. (TNS-VS-1166219); Aiko-gun, Aikawa-machi,

13, T. Ohshima s.n. (TNS-VS-1156187); Nakaniikawa-gun, Kami-

1981-11-22, M. Akiyama s.n. (TNS-VS-545686); Tsukui-gun, Sag-

ichi-machi, 1978-10-8, T. Ohshima s.n. (TNS-VS-528704); Tat-amiko-machi, 1982-10-14, H. Kobayashi s.n. (TNS-VS-594944);

eyama-mura, 1929-8-31, Z. Tashiro s.n. (TNS-VS-35380);

Tsuzuki-gun, Tsuda-mura, 1930-8-30, K. Matsushima s.n. (TNS-

Tateyama-machi, 1976-10-20, T. Ohshima s.n. (TNS-VS-528745);

VS-663233); Niigata Pref.: Takada-shi, 1960-11-6, K. Sakai 11

Shimoniikawa-gun, Asahi-machi, 1981-9-15, T. Ohshima s.n.

(TNS-VS-1042038); Kashiwazaki-shi, 1964-10-11, S. Iwano s.n.

(TNS-VS-528743),(TNS-VS-543176), 1986-8-26, I. Ohzu s.n. (TNS-VS-1166311);

VS-1156181); Nei-gun, Yatsuo-machi, 1980-7-8, T. Ohshima s.n.

Shibata-shi, 1962-10-28, F. Konta 1085 (TNS-VS-929625), 1963-(TNS-VS-528756); Yamada-mura, 1978-10-7, T. Ohshima s.n.

10-11, collector unknown s.n. (TNS-VS-929599); Niitsu-shi, 1978-(TNS-VS-528749); Higashitonami-gun, Kamitaira-mura, 1983-8-

8-1, M. Saito 2058 (TNS-VS-543167); Ojiya-shi, 1953-10-8, S.

6,Iwano s.n. (TNS-VS-543175), 1980-7-30, T. Yoshizawa s.n. (TNS-

Fukuoka-machi, 1988-8-21, T. Ohshima s.n. (TNS-VS-1157007);

VS-424026); Kamo-shi, 1964-9-13, T. Takahashi 2457 (TNS-

Ishikawa Pref.: Kanazawa-shi, 1973-9-9, M. Honda s.n. (TNS-

VS-543180), 1966-10-10, T. Takahashi 2846 (TNS-VS-1172232),

VS-424527), 1975-8-20, M. Honda s.n. (TNS-VS-424528), 1976-

1948-10-17, T. Uchikawa s.n. (TNS-VS-89863); Kitakanbara-gun,

10-10, M. Honda s.n. (TNS-VS-424515); Nanao-shi, 1979-8-11, S.

Sasakami-mura, 1963-11-7, F. Konta 3405 (TNS-VS-929610),

Komaki s.n. (TNS-VS-424523), 1978-11-3, S. Komaki s.n. (TNS-

1963-11-7, F. Konta 3426 (TNS-VS-929600), 1978-11-14, M.

VS-424513); Komatsu-shi, 1980-8-18, M. Honda s.n. (TNS-

Saito 2391 (TNS-VS-543168), 1986-10-21, M. Saito s.n. (TNS-

VS-424533), 1980-8-9, M. Honda s.n. (TNS-VS-424532), 1979-8-

VS-1157042, 1166312), 1978-11-14, M. Saito s.n. (TNS-11, K. Yoneyama s.n. (TNS-VS-424525); Wajima-shi, 1975-10-5,

VS-1172239); Toyoura-machi, 1987-9-4, M. Saito s.n. (TNS-

S. Komaki s.n. (TNS-VS-424535), 1977-10-11, S. Komaki s.n.

Musashimurayama-shi,

1980-9-20,

T.

Ohshima

1982-11-6,

s.n.

T.

Ohshima(TNS-VS-1156183);

s.n.

(TNS-

Nishitonami-gun,

70

Atsushi Ebihara and Narumi Nakato(TNS-VS-424509), 1978-8-21, S. Komaki s.n. (TNS-VS-424522);

VS-1166255); Otsuki-shi, 1977-8-31, T. Kobayashi s.n. (TNS-

Suzu-shi, 1978-11-15, S. Komaki s.n. (TNS-VS-424511), 1977-11-

VS-573491), 1968-8-12, T. Kobayashi s.n. (TNS-VS-573487);

13, S. Komaki s.n. (TNS-VS-424544), 1976-11-13, S. Komaki s.n.

Nishiyatsushiro-gun, Kamikuishiki-mura, 1981-8-31, F. Miyamoto(TNS-VS-424510); Kaga-shi, 1972-7-30, H. Ito s.n. (TNS-

7176 (TNS-VS-831734), 1981-8-29, E. Fukushi s.n. (TNS-

VS-1042058, 1042059), 1976-11-3, M. Honda s.n. (TNS-

VS-1166254); Minamikoma-gun, Minobu-cho, 1981-10-24, M.

VS-424516); Hakui-shi, 1975-11-16, S. Komaki s.n. (TNS-

Nakagawa s.n. (TNS-VS-1166259), 1968-8-17, T. Kobayashi s.n.

VS-424540); Enuma-gun, Yamanaka-machi, 1976-11-3, M. Honda(TNS-VS-573488),s.n. (TNS-VS-424529); Nomi-gun, Tatsunokuchi-machi, 1977-11-

VS-262042), 1951-10-2, S. Okuyama 14389 (TNS-VS-262059);

3, K. Yoneyama s.n. (TNS-VS-424530); Ishikawa-gun, Kawachi-

Nanbu-cho, 1985-11-24, A. Ozaki s.n. (TNS-VS-1166305);

mura, 1980-9-28, M. Honda s.n. (TNS-VS-424534); Kahoku-gun,Tomizawa-cho,

Tsubata-machi, 1976-10-1, S. Komaki s.n. (TNS-VS-424541);

VS-1166253), 1987-8-10, T. Hosokura s.n. (TNS-VS-1156192);

Unoke-machi, 1968-11-20, S. Komaki s.n. (TNS-VS-424547);

Minamitsuru-gun, Nakano-mura, 1956-7-0, K. Suenaga s.n. (TNS-

Hakui-gun, Togi-machi, 1978-10-15, S. Komaki s.n. (TNS-

VS-1042077); Narusawa-mura, 1981-8-29, E. Fukushi s.n. (TNS-

VS-424520), 1973-8-21, S. Komaki s.n. (TNS-VS-424508), 1973-

VS-1166260); Kitatsuru-gun, Uenohara-machi, 1981-9-15, H.

9-2, S. Komaki s.n. (TNS-VS-424507); Shika-machi, 1978-10-8, S.

Endo s.n. (TNS-VS-1166261); Tabayama-mura, 1981-8-24, A.

Komaki s.n. (TNS-VS-424514); Oshimizu-machi, 1975-8-15, S.

Yamamoto s.n. (TNS-VS-1166258); Nagano Pref.: Iida-shi, 1981-

Komaki s.n. (TNS-VS-424543); Kashima-gun, Tatsuruhama-

10-24, H. Tsutsumi s.n. (TNS-VS-1157112); Ina-shi, 1981-10-26,

machi, 1978-9-12, S. Komaki s.n. (TNS-VS-424505), 1977-9-17,

H. Tsutsumi s.n. (TNS-VS-574301); Koshoku-shi, 1981-10-7, H.

S. Nakada s.n. (TNS-VS-424519); Nakajima-machi, 1978-9-3, S.

Tsutsumi s.n. (TNS-VS-574326); Kamiina-gun, Iijima-machi,

Komaki s.n. (TNS-VS-424518), 1978-10-24, S. Komaki s.n. (TNS-

1981-8-11, H. Tsutsumi s.n. (TNS-VS-574294); Miyada-mura,S.

1951-10-2,

1981-10-17,

OkuyamaM.

Nakagawa

14394

s.n.

(TNS-

(TNS-

VS-424512); Kashima-machi, 1969-11-19, S. Komaki s.n. (TNS-

1981-10-13, H. Tsutsumi s.n. (TNS-VS-574303), 1981-8-13, H.

VS-424539), 1977-10-12, S. Nakada s.n. (TNS-VS-424524);

Tsutsumi s.n. (TNS-VS-574295); Shimoina-gun, Takamori-machi,

Notojima-machi, 1978-9-27, S. Komaki s.n. (TNS-VS-424521);

1981-11-7, H. Tsutsumi s.n. (TNS-VS-574291); Kamisato-mura,Fugeshi-gun, Anamizu-machi, 1977-10-10, S. Komaki s.n. (TNS-

1932-10-2, M. Muramatsu 3530 (TNS-VS-298523); Achi-mura,

VS-424503), 1975-11-8, S. Komaki s.n. (TNS-VS-424542), 1969-

1981-10-10, H. Tsutsumi s.n. (TNS-VS-574305); Neba-mura,

11-10, S. Komaki s.n. (TNS-VS-424502); Noto-machi, 1977-10-

1981-10-12, H. Tsutsumi s.n. (TNS-VS-574313); Shimojo-mura,S.

Komakis.n.

Yanagida-mura,

1930-9-20, M. Furuse s.n. (TNS-VS-909807); Urugi-mura, 1981-

1976-10-12, S. Komaki s.n. (TNS-VS-424546); Suzu-gun, Uchi-

10-12, H. Tsutsumi s.n. (TNS-VS-574312); Tenryu-mura, 1981-11-

ura-machi, 1968-10-18, S. Komaki s.n. (TNS-VS-424538); Fukui

16, H. Tsutsumi s.n. (TNS-VS-1157069), 1974-7-27, K. Otsuka

Pref.: Fukui-shi, 1977-6-4, Y. Saito s.n. (TNS-VS-1166214), 1987-

1857 (TNS-VS-1157064), 1968-10-10, Y. Hara s.n. (TNS-

7-26, Y. Saito s.n. (TNS-VS-1156134); Tsuruga-shi, 1976-7-18, S.

VS-1157068); Yasuoka-mura, 1981-7-25, H. Tsutsumi s.n. (TNS-

Watanabe s.n. (TNS-VS-1166202); Takefu-shi, 1978-9-13, S.

VS-574307), 1981-10-14, H. Tsutsumi s.n. (TNS-VS-574324);

Watanabe s.n. (TNS-VS-1166200), 1984-8-19, Y. Saito s.n. (TNS-

Kiso-gun, Agematsu-machi, 1981-10-31, H. Tsutsumi s.n. (TNS-

VS-1156993); Ono-shi, 1979-8-13, S. Watanabe s.n. (TNS-

VS-574290), 1980-8-25, T. Nakaike s.n. (TNS-VS-1157071),

VS-1166197), 1971-10-3, Y. Saito s.n. (TNS-VS-1166209), 1979-

1981-8-26, H. Tsutsumi s.n. (TNS-VS-574304); Nagiso-machi,

8-31, S. Watanabe s.n. (TNS-VS-1166196), 1974-11-9, Y. Saito

1976-7-30, K. Otsuka 5483 (TNS-VS-431605), 1976-7-31, T.

s.n.

Nakaike s.n. (TNS-VS-1157070), 1981-10-19, H. Tsutsumi s.n.

10,

(TNS-VS-1166190),(TNS-VS-424545);

1975-8-9,

Y.

Saito

s.n.

(TNS-

VS-1166213), 1975-7-28, Y. Saito s.n. (TNS-VS-1166225), 1975-(TNS-VS-574300),

8-3, Y. Saito s.n. (TNS-VS-1166226); Katsuyama-shi, 1978-8-29,

VS-574308); Hiyoshi-mura, 1981-10-30, H. Tsutsumi s.n. (TNS-

Y. Saito s.n. (TNS-VS-1166210), 1978-10-8, Y. Saito s.n. (TNS-

VS-574292); Okuwa-mura, 1973-7-28, H. Miyazaki 2819 (TNS-

VS-1166211), 1974-10-?, Y. Saito s.n. (TNS-VS-1166191); Sabae-

VS-831863), 1980-11-4, K. Otsuka 7634 (TNS-VS-431604),

shi, 1961-9-5, S. Watanabe s.n. (TNS-VS-1166199); Yoshida-gun,

1981-8-24, H. Tsutsumi s.n. (TNS-VS-574309), 1980-11-3, M.

Eiheiji-cho, 1979-8-5, S. Watanabe s.n. (TNS-VS-1166195); Ono-

Fukuhara 8318 (TNS-VS-1157073), 1974-8-1, K. Otsuka 2017

gun, Izumi-mura, 1979-8-8, S. Watanabe s.n. (TNS-VS-1166203);(TNS-VS-1157076),

Sakai-gun, Mikuni-cho, 1992-11-3, S. Watanabe 36713 (TNS-

VS-1157079), 1981-8-25, H. Tsutsumi s.n. (TNS-VS-574306,

1981-8-24,

1980-8-25,

H.

T.

Tsutsumi

Nakaike

s.n.

s.n.

(TNS-

(TNS-

VS-1156207); Kanazu-cho, 1980-8-24, Y. Saito s.n. (TNS-574310); Yamaguchi-mura, 1981-10-19, H. Tsutsumi s.n. (TNS-

VS-1166192); Imadate-gun, Ikeda-cho, 1963-8-13, S. Watanabe

VS-574299), 1954-8-15, H. Okuhara s.n. (TNS-VS-1157110),s.n. (TNS-VS-1166201), 1964-7-28, S. Watanabe s.n. (TNS-

1967-10-26, H. Okuhara s.n. (TNS-VS-1157111), 1981-8-29, H.

VS-1166198); Nyu-gun, Asahi-cho, 1978-7-15, Y. Saito s.n. (TNS-

Tsutsumi s.n. (TNS-VS-574302); Minamiazumi-gun, Horigane-

VS-1166207); Mikata-gun, Mikata-cho, 1975-7-19, Y. Saito s.n.

mura, 1981-9-11, H. Tsutsumi s.n. (TNS-VS-574311); Kamimino-(TNS-VS-1166212); Onyu-gun, Natasho-mura, 1980-10-9, S.

chi-gun, Shinano-machi, 1981-9-25, H. Tsutsumi s.n. (TNS-

Watanabe s.n. (TNS-VS-1166194), 1978-9-13, S. Watanabe s.n.

VS-574297), 1981-9-25, H. Tsutsumi s.n. (TNS-VS-574296); Gifu

(TNS-VS-1166223), 1976-7-27, Y. Saito s.n. (TNS-VS-1166224);

Pref.: Gifu-shi, 1957-10-31, K. Inami s.n. (TNS-VS-285244),

Yamanashi Pref.: Kofu-shi, 1981-8-17, A. Yamamoto s.n. (TNS-

1967-9-13, S. Serizawa 4731 (TNS-VS-1042057), 1976-9-4, Y.

Distribution of the Thelypteris japonica Complex (Thelypteridaceae)

71

Yamamoto s.n. (TNS-VS-1157130), 1984-7-15, K. Inukai 7057

1981-9-18, A. Yamamoto s.n. (TNS-VS-1157084); Miyagawa-(TNS-VS-1156146); Takayama-shi, 1981-9-19, A. Yamamoto s.n.

mura, 1979-9-15, T. Waku 9596 (TNS-VS-1157114); Kamioka-(TNS-VS-1157086); Tajimi-shi, 1978-12-10, M. Fukuhara 7397

cho, 1977-8-12, T. Ohshima s.n. (TNS-VS-528747); Shizuoka(TNS-VS-1157081); Seki-shi, 1977-9-28, Y. Yamamoto s.n. (TNS-

Pref.: Shizuoka-shi, 1970-10-3, K. Takeno 365 (TNS-VS-929626),

VS-1157080), 1976-8-18, Y. Yamamoto s.n. (TNS-VS-1157120);

1966-10-6, F. Konta & M. Wakabayashi 263 (TNS-VS-929574),Y.

Yamamoto

s.n.

(TNS-

1967-9-30, F. Konta 6554 (TNS-VS-929592), 1981-9-27, T.

VS-1157149, 1157150), 1980-11-23, T. Ogiyama 1011 (TNS-

Shimakawa s.n. (TNS-VS-509170), 1979-9-22, M. Nakagawa s.n.

VS-545769), 1987-8-15, T. Ogiyama 7150 (TNS-VS-1157008);(TNS-VS-509171),

Mizunami-shi, 1977-7-27, Y. Yamamoto s.n. (TNS-VS-1157124);

VS-509173), 1979-11-17, M. Nakagawa s.n. (TNS-VS-509174),

Ena-shi, 1991-8-8, S. Masuda s.n. (TNS-VS-831861), 1978-11-25,

1980-8-18, H. Suzuki s.n. (TNS-VS-509158), 1980-8-2, T. Sugino

Y. Yamamoto s.n. (TNS-VS-1157151), 1983-11-13, K. Inukai 6647

s.n. (TNS-VS-1166235), 1977-5-14, H. Kobayashi s.n. (TNS-(TNS-VS-1156203), 1984-10-28, Y. Yamamoto s.n. (TNS-

VS-1166251), 1985-8-17, K. Tanaka s.n. (TNS-VS-1166303);

VS-1166309); Kakamigahara-shi, 1976-9-25, Y. Yamamoto s.n.

Hamamatsu-shi, 2000-8-8, T. Nakamura s.n. (TNS-VS-1019336),

Nakatsugawa-shi,

1978-11-15,

Y.

Kimizukas.n.

1979-9-8,

M.

Nakagawa

s.n.

(TNS-

(TNS-

1980-11-3, T. Nakura s.n. (TNS-VS-509186), 1980-9-15, collector

VS-1157113); Kaizu-gun, Nanno-cho, 1981-10-29, I. Awata 3417

unknown s.n. (TNS-VS-1157103), 1980-9-15, I. Yamashita s.n.

(TNS-VS-1157131),

1979-9-14,

Y.

Yamamoto

s.n.

(TNS-

(TNS-VS-1166247); Numazu-shi, 1983-9-25, T. Sato 4714 (TNS-

VS-435842); Ibi-gun, Tanigumi-mura, 1984-9-15, H. Miyazaki

VS-929629), 1982-8-13, T. Sato 2143 (TNS-VS-929620), 1977-7-

6585 (TNS-VS-831713), 1984-9-15, H. Miyazaki 6571 (TNS-3, H. Kobayashi s.n. (TNS-VS-509126), 1983-9-25, T. Sato 4713

VS-831712), 1984-9-15, H. Miyazaki 6586 (TNS-VS-831711);(TNS-VS-1156112); Shimizu-shi, 1993-10-11, T. Hosokura s.n.

Kasuga-mura, 1976-11-7, Y. Yamamoto s.n. (TNS-VS-435843),(TNS-VS-1156989); Atami-shi, 1980-10-21, H. Suzuki s.n. (TNS-

1979-11-9, Y. Yamamoto s.n. (TNS-VS-1157132); Motosu-gun,VS-1157099); Fujinomiya-shi, 1974-8-29, F. Konta 10474 (TNS-

Neo-mura, 1981-9-6, F. Miyamoto 7116 (TNS-VS-831733), 1981-

VS-929617), 1969-9-17, F. Konta 8170 (TNS-VS-929604), 1983-

10-4, Y. Yamamoto s.n. (TNS-VS-435846), 1977-10-19, Y. Yamamoto

9-18, T. Sato 4455 (TNS-VS-929616), 1983-8-22, T. Sato 3940

s.n. (TNS-VS-1157121), 1978-9-8, Y. Yamamoto s.n. (TNS-(TNS-VS-929613), 1984-9-16, T. Sato 6156 (TNS-VS-929585),

VS-1157122), 1978-9-1, Y. Yamamoto s.n. (TNS-VS-1157139),

1984-9-24, T. Sato 6426 (TNS-VS-929583), 1983-8-28, T. Sato

1984-11-4, Y. Yamamoto s.n. (TNS-VS-1166308); Yamagata-gun,

4022 (TNS-VS-929575), 1979-9-4, Y. Sakuma s.n. (TNS-(TNS-VS-435847),

1978-10-15,

Miyama-cho, 1982-9-14, H. Takahashi & al. 6650 (TNS-

VS-820712, 820713), ????-9-3, T. Nakamura s.n. (TNS-

VS-929623), 1978-7-25, Y. Yamamoto s.n. (TNS-VS-1157128),

VS-820245, 820582), 1985-9-2, T. Miyazawa 142 (TNS-

1979-9-9, M. Fukuhara 7598 (TNS-VS-1157082); Mugi-gun, Ita-

VS-929627), 1979-8-30, M. Nakagawa s.n. (TNS-VS-508988),

dori-mura, 1964-8-11, K. Fukui 7 (TNS-VS-929587), 1964-8-16,

1983-9-18, T. Sato 4454 (TNS-VS-1156113), 1983-8-22, T. Sato

N. Fukuoka & K. Yamashita 272 (TNS-VS-929586), 1978-10-13,

3939 (TNS-VS-1156114), 1983-9-24, T. Sato 4701 (TNS-

Y. Yamamoto s.n. (TNS-VS-1157116), 1977-10-21, Y. Yamamoto

VS-1156116), 1982-10-16, M. Nakagawa s.n. (TNS-VS-1156165),

s.n. (TNS-VS-1157115); Mugegawa-cho, 1976-11-28, Y. Yamamoto

1984-9-16, T. Sato 6153 (TNS-VS-1157035), 1980-9-23, T. Sugino

s.n. (TNS-VS-1157127); Mugi-cho, 1976-10-10, F. Miyamotos.n. (TNS-VS-1166236); Ito-shi, 1981-7-26, T. Shimakawa s.n.

1320 (TNS-VS-831737), 1976-10-10, F. Miyamoto 1321 (TNS-(TNS-VS-509169); Fuji-shi, 1983-8-22, T. Sato 3796 (TNS-

VS-831738); Gujo-gun, Shiratori-cho, 1966-8-8, K. Sakurai s.n.

VS-929622), 1983-7-24, T. Sato 3474 (TNS-VS-929582), 1983-9-(TNS-VS-170619); Minami-mura, 1977-12-8, Y. Yamamoto s.n.

15, T. Sato 4364 (TNS-VS-1156162); Iwata-shi, 1962-11-4, K.

(TNS-VS-1157126); Wara-mura, 1981-8-10, Y. Yamamoto s.n.

Satake s.n. (TNS-VS-311833, 311845, 318001); Kakegawa-shi,

Y.

Yamamoto

s.n.

(TNS-

1980-10-26, T. Sato s.n. (TNS-VS-509167), 1981-10-11, T. Sugino

VS-1157148); Kamo-gun, Yaotsu-cho, 1979-9-23, Y. Yamamotos.n. (TNS-VS-509292), 1977-6-4, H. Kobayashi s.n. (TNS-s.n. (TNS-VS-1157125), 1984-10-1, K. Inukai 7264 (TNS-

VS-1166237); Fujieda-shi, 1981-7-29, T. Sugino s.n. (TNS-

VS-1156143); Shirakawa-cho, 1978-10-31, Y. Yamamoto s.n.

VS-509123),(TNS-VS-1157146), 1978-10-31, Y. Yamamoto s.n. (TNS-

Gotenba-shi, 1975-10-26, T. Nakaike s.n. (TNS-VS-335679);

VS-1157145), 1977-8-16, Y. Yamamoto s.n. (TNS-VS-1157140);

Fukuroi-shi, 1980-9-23, T. Sato s.n. (TNS-VS-1166246); Tenryu-

Toki-gun, Kasahara-cho, 1984-9-16, K. Inukai 7201 (TNS-shi, 1976-9-2, H. Suzuki s.n. (TNS-VS-509128), 1979-10-5, T.

VS-1156144); Ena-gun, Kawaue-mura, 1980-9-19, Y. Yamamoto

Sato s.n. (TNS-VS-509127), 1980-9-13, T. Sato s.n. (TNS-s.n. (TNS-VS-435845); Kashimo-mura, 1977-8-23, Y. Yamamoto

VS-509193), 1981-10-18, T. Sugino s.n. (TNS-VS-1166232);s.n. (TNS-VS-1157135); Kushihara-mura, 1984-9-24, T. Ogiyama

Hamakita-shi, 1980-9-8, T. Sato s.n. (TNS-VS-509192); Shimoda-

5530

T.

shi, 1981-7-29, T. Sato 339 (TNS-VS-425126); Susono-shi, 1976-

Ogiyama 5473 (TNS-VS-1157057); Mashita-gun, Hagiwara-cho,

11-7, C. Ohkawa s.n. (TNS-VS-388970), 1983-9-17, A. Suzuki s.n.

1981-9-19, A. Yamamoto s.n. (TNS-VS-1157087); Gero-cho,(TNS-VS-1156984); Kosai-shi, 1980-9-15, T. Sugino s.n. (TNS-

1978-11-17, Y. Yamamoto s.n. (TNS-VS-1157134), 1982-9-26, N.

VS-509185), 1980-9-20, I. Yamashita s.n. (TNS-VS-509124,

Futamura s.n. (TNS-VS-1157046); Kanayama-cho, 1978-9-26, Y.

1166249); Kamo-gun, Higashiizu-cho, 1982-9-5, T. Sato 2415

Yamamoto s.n. (TNS-VS-1157142); Yoshiki-gun, Kokufu-cho,(TNS-VS-929618); Kawazu-cho, 1969-10-3, F. Konta 1157 (TNS-(TNS-VS-435844),

1978-10-17,(TNS-VS-1157056);

Kamiyahagi-cho,

1984-9-23,

1981-8-6,

T.

Suginos.n.

(TNS-VS-509121);

72

Atsushi Ebihara and Narumi Nakato
VS-929628), 1956-11-16, K. Satake s.n. (TNS-VS-9512669),

Sakuma-cho, 1980-9-10, T. Sato & H. Suzuki s.n. (TNS-

1965-10-16, H. Ito s.n. (TNS-VS-1045362); Matsuzaki-cho, 1981-

VS-509180), 1981-8-15, T. Sugino s.n. (TNS-VS-509183), 1971-

10-7, T. Sato 833 (TNS-VS-425127); Nishiizu-cho, 1976-10-30, A.

8-17, H. Suzuki s.n. (TNS-VS-1157105), 1969-7-27, I. Yamashita

Yamamoto s.n. (TNS-VS-509177); Kamo-mura, 1975-8-13, A.

s.n. (TNS-VS-1166242), 1976-8-18, I. Yamashita s.n. (TNS-

Suzuki s.n. (TNS-VS-509160); Tagata-gun, Shuzenji-cho, 1983-

VS-1166245); Misakubo-cho, 1976-9-10, F. Miyamoto 1169 (TNS-

10-8, A. Suzuki s.n. (TNS-VS-1156985), 1983-11-17, Y. Kimizuka

VS-831719, 831720a/b), 1978-9-3, F. Miyamoto 3223 (TNS-VS-

s.n. (TNS-VS-1157049); Nirayama-cho, 1979-12-8, H. Kobayashi

831725a/b),s.n. (TNS-VS-1166238); Kamikano-mura, 1931-11-7, T. Tahara

1980-9-11, T. Sato s.n. (TNS-VS-509179), 1979-8-15, T. Sugino

s.n.

s.n.

(TNS-VS-831688),

1938-8-25,

Y.

Jotani

s.n.

(TNS-

T.

1977-8-16,(TNS-VS-509190),

Suginos.n.
T.

1978-8-20,(TNS-VS-509184),
Sato

s.n.

(TNS-

VS-831689, 831690), 1929-11-23, Y. Jotani 16 (TNS-VS-831687);

VS-1157106), 1977-10-11, I. Yamashita s.n. (TNS-VS-1166241);

Amagiyugashima-cho, 1966-7-23, F. Konta & M. Wakabayashi 76

Hamana-gun, Arai-cho, 1980-9-23, T. Nakura s.n. (TNS-(TNS-VS-929578), 1970-8-2, M. Ichikawa & al 1233 (TNS-

VS-1166228); Inasa-gun, Hosoe-cho, 1980-10-9, T. Sato s.n.

VS-929594), 1972-10-30, H. Miyazaki 2294 (TNS-VS-831718),(TNS-VS-1157101); Inasa-cho, 1981-11-22, T. Sugino s.n. (TNS-

1961-9-27, J. Sugimoto s.n. (TNS-VS-247756), 1971-10-10, M.

VS-509182), 1981-11-23, T. Sugino s.n. (TNS-VS-509181); Mikk-

Sakakibara s.n. (TNS-VS-285563), 1970-8-3, M. Ichikawa, T.

abi-cho, 1980-10-9, T. Sato s.n. (TNS-VS-1157100); Aichi Pref.:Fujimura & F. Konta 1338 (TNS-VS-929621), 1983-9-5, T.

Nagoya-shi, Moriyama-ku, 1980-7-6, K. Inukai 3372 (TNS-

Miyazawa 14 (TNS-VS-929633), 1979-8-26, M. Nakagawa s.n.

VS-432968); Chikusa-ku, 1980-9-29, K. Inukai 3905 (TNS-(TNS-VS-509176), 1980-9-14, A. Suzuki s.n. (TNS-VS-509161),

VS-432932); Showa-ku, 1980-?-?, M. Fukuhara 7995 (TNS-

1977-10-23, A. Suzuki s.n. (TNS-VS-509159), 1977-10-10, H.

VS-432935);

Kobayashi s.n. (TNS-VS-1157102); Nakaizu-cho, 1965-11-6, C.

(TNS-VS-432944); Nakagawa-ku, 1982-11-14, K. Inukai 5783

Ohkawa s.n. (TNS-VS-457332), 1977-7-24, A. Suzuki s.n. (TNS-(TNS-VS-1156202); Toyohashi-shi, 1977-11-13, M. Fukuhara

VS-1166230), 1986-9-1, T. Hosokura s.n. (TNS-VS-1166310);

6736 (TNS-VS-432925), 1980-10-26, K. Inukai 4071 (TNS-

Sunto-gun, Nagaizumi-cho, 1948-10-3, Y. Jotani 25248 (TNS-

VS-432964); Okazaki-shi, 1980-9-7, K. Inukai 3787 (TNS-

VS-831683); Susono-cho, 1947-10-18, Y. Jotani 25249 (TNS-

VS-432931), 1980-6-29, K. Inukai 3344 (TNS-VS-432969); Seto-

VS-831684); Oyama-cho, 1985-11-4, T. Hosokura s.n. (TNS-shi,

VS-1156128), 1984-9-24, T. Hosokura s.n. (TNS-VS-1157009,

1980-8-17, K. Inukai 3644 (TNS-VS-432958); Handa-shi, 1978-9-

1157018); Fuji-gun, Shibakawa-cho, 1984-12-1, T. Sato 7115

1, J. Kimura s.n. (TNS-VS-432917); Tsushima-shi, 1978-10-7, M.

(TNS-VS-929584), 1985-9-26, T. Sato 8530 (TNS-VS-929579);

Muramatsu 78526 (TNS-VS-432955); Toyota-shi, 1978-10-8, M.

Abe-gun, Ikawa-mura, 1967-9-10, H. Ito s.n. (TNS-VS-1045379),

Muramatsu 78563 (TNS-VS-432954), 1980-10-6, K. Inukai 3962

1967-9-12, S. Serizawa 4583 (TNS-VS-1042056); Ihara-gun, Yui-

(TNS-VS-432966); Anjo-shi, 1982-7-28, K. Inukai 5433 (TNS-cho, 1983-9-11, T. Hosokura s.n. (TNS-VS-1156171); Tagata-gun,VS-1156210); Nishio-shi, 1979-11-23, K. Inukai 2718 (TNS-

Kamiohmi-mura, 1939-9-2, Y. Jotani s.n. (TNS-VS-831692),

VS-432948); Gamagori-shi, 1978-7-16, M. Fukuhara 6831 (TNS-

1943-11-19, S. Okuyama 5122 (TNS-VS-89892); Shida-gun,

VS-432926);

Okabe-cho, 1962-8-23, I. Yamashita s.n. (TNS-VS-1166240); Hai-

(TNS-VS-831729), 1980-8-30, K. Inukai 3750 (TNS-VS-432956),

T.

Suginos.n.

Midori-ku,

1980-8-28,

M.

Muramatsu

Inuyama-shi,M.

1980-7-27,

80366

1979-7-13,

F.

Fukuhara

7935(TNS-VS-432942),

Miyamoto

6809

(TNS-

1979-11-3, K. Inukai 2655 (TNS-VS-432951); Tokoname-shi,

VS-509132), 1981-11-28, T. Sugino s.n. (TNS-VS-509131); Goka-

1982-10-3, K. Inukai 5645 (TNS-VS-1156991); Komaki-shi,bara-gun,Haibara-cho,G.

1981-7-12,
Hashimotos.n.

(TNS-VS-44317);

1973-7-27, M. Fukuhara 4744 (TNS-VS-432918); Shinshiro-shi,

Kawane-cho, 1980-9-16, T. Sugino s.n. (TNS-VS-509118);

1974-9-13, K. Mimoro, S. Tsugaru & K. Nishiyama 2178 (TNS-

Honkawane-cho, 1970-7-18, K. Nakayama 21 (TNS-VS-929614),

VS-335954), 1980-8-12, K. Inukai 3592 (TNS-VS-432959);

1967-7-23, H. Takahashi 129 (TNS-VS-929596), 1972-11-23, Y.

Chiryu-shi, 1980-9-7, K. Inukai 3758 (TNS-VS-432929); Owar-mura,

1933-8-27,

Saiki & F. Konta 1616 (TNS-VS-929602), 1977-10-30, I.

iasahi-shi, 1980-10-12, M. Muramatsu 80593 (TNS-VS-432943);

Yamashita s.n. (TNS-VS-509125), 1980-9-16, T. Sugino s.n. (TNS-

Ama-gun,VS-509122), 1980-8-19, H. Suzuki s.n. (TNS-VS-1166227), 1980-

VS-432940); Chita-gun, Agui-cho, 1980-8-2, K. Inukai 3494

9-16, T. Sugino s.n. (TNS-VS-1166234); Ogasa-gun, Hamaoka-(TNS-VS-432961); Mihama-cho, 1979-9-16, K. Inukai 2506

cho, 1963-12-15, T. Sugino s.n. (TNS-VS-509157); Shuchi-gun,(TNS-VS-432946), 1981-11-1, K. Inukai 5121 (TNS-VS-436787);

Mori-machi, 1980-8-11, H. Suzuki s.n. (TNS-VS-509129), 1981-

Hazu-gun, Isshiki-cho, 1976-11-1, M. Fukuhara 6373 (TNS-

11-15, T. Sugino s.n. (TNS-VS-509195); Haruno-cho, 1980-10-17,

VS-432923); Hazu-cho, 1974-12-15, M. Fukuhara 5503 (TNS-

H. Suzuki & T. Sato s.n. (TNS-VS-509166), 1964-8-14, I.

VS-432921); Nukata-gun, Nukata-cho, 1990-10-27, Y. Fukuoka

Yamashita s.n. (TNS-VS-1166243), 1981-10-21, T. Sugino s.n.

1895 (TNS-VS-683600), 1990-9-24, Y. Fukuoka 1483 (TNS-(TNS-VS-1166252); Iwata-gun, Asaba-cho, 1981-10-10, T. Sugino

VS-683702), 1990-10-4, Y. Fukuoka 1585 (TNS-VS-683704),

s.n. (TNS-VS-509194); Toyoka-mura, 1982-10-21, Y. Kurosawa

1990-9-24, Y. Fukuoka 1473 (TNS-VS-683705), 1990-9-24, Y.

176 (TNS-VS-637724), 1981-10-10, T. Sugino s.n. (TNS-

Fukuoka 1482 (TNS-VS-683697), 1980-10-18, K. Inukai 4056

VS-1157104), 1981-10-18, T. Sugino s.n. (TNS-VS-1166231); Tat-

(TNS-VS-432963),

suyama-mura, 1968-10-?, Y. Yokoe 243 (TNS-VS-929608);

VS-683735); Nishikamo-gun, Obara-mura, 1981-10-10, K. Inukai

Miwa-cho,

1981-7-22,

1990-10-2,

Y.

K.

InukaiFukuoka

4583

1544

(TNS-

(TNS-

Distribution of the Thelypteris japonica Complex (Thelypteridaceae)

735037 (TNS-VS-436786); Higashikamo-gun, Asuke-cho, 1978-8-

Sonoda s.n. (TNS-VS-268409); Gamo-gun, Hino-cho, 1981-8-16,

12, M. Fukuhara 7060 (TNS-VS-432927), 1980-7-26, K. Inukai

T. Shiraiwa 10267 (TNS-VS-435453), 1982-10-29, T. Tatebe s.n.

3398 (TNS-VS-432934); Shimoyama-mura, 1980-9-23, K. Inukai(TNS-VS-1156209); Ryuo-cho, 1979-9-14, Y. Kimizuka s.n. (TNS-

3895 (TNS-VS-432933), 1979-12-9, K. Inukai 2817 (TNS-

VS-533922);

VS-432947); Asahi-cho, 1980-7-29, K. Inukai 3442 (TNS-

Muranaga s.n. (TNS-VS-533920); Echi-gun, Hatasho-cho, 1977-

VS-432962); Kitashitara-gun, Toei-cho, 1981-8-7, K. Inukai 4698

10-22, A. Muranaga s.n. (TNS-VS-533921); Takashima-gun, Kut-

(TNS-VS-432938); Toyone-mura, 1978-8-23, M. Fukuhara 7182

suki-mura, 1981-9-13, T. Tatebe s.n. (TNS-VS-533919), 1982-10-(TNS-VS-432928); Tomiyama-mura, 1981-9-20, K. Inukai 4953

25, K. Nakamura & al. s.n. (TNS-VS-1156217); Kyoto Pref.:(TNS-VS-432937); Tsugu-mura, 1980-10-10, K. Inukai 3968

Kyoto-shi, 1955-11-3, M. Tagawa 7157 (TNS-VS-132766);(TNS-VS-432967); Inabu-cho, 1979-9-24, K. Inukai 2521 (TNS-

Sakyo-ku, 1982-10-25, F. Miyamoto 7581 (TNS-VS-831727),

VS-432949); Minamishitara-gun, Horai-cho, 1967-11-19, H. Ito

1951-9-24, T. Kodama s.n. (TNS-VS-1042039), 1982-10-25, E.

s.n. (TNS-VS-1045686), 1981-8-22, M. Fukuhara 8634 (TNS-

Fukushi s.n. (TNS-VS-1156214), 1963-9-19, F. Konta 2746 (TNS-

M.

(TNS-

VS-929611), 1963-8-18, K. Iwatsuki 5590 (TNS-VS-248677);

VS-432952), 1980-9-29, K. Inukai 3933 (TNS-VS-432965), 1981-

Higashiyama-ku, 1965-9-5, F. Konta 5313 (TNS-VS-929591);

VS-432920),
8-12,

K.

1980-11-23,
Inukai4743

Muramatsu(TNS-VS-432914);

80614

Tsukude-mura,Ukyo-ku,Kanzaki-gun,

1981-9-9,Y.

Gokasho-cho,

Kimizukas.n.

1977-10-21,

A.

(TNS-VS-1157091);

1974-9-13, K. Mimoro, S. Tsugaru & K. Nishiyama 2195 (TNS-

Kameoka-shi, 1991-9-26, S. Tsugaru 15396 (TNS-VS-575493),

VS-335961), 1981-8-25, M. Fukuhara 8714 (TNS-VS-432919);

1991-8-17, T. Takahashi 2104 (TNS-VS-594806), 1973-9-21, K.

Hoi-gun, Ichinomiya-cho, 1981-9-28, K. Inukai 5024 (TNS-

Mimoro & S. Tsugaru 904 (TNS-VS-337605), 1991-9-17, S.

VS-432936); Otowa-cho, 1978-11-19, M. Fukuhara 7363 (TNS-

Tsugaru 15332 (TNS-VS-571255); Tsuzuki-gun, Yawata-cho,VS-432945); Atsumi-gun, Tahara-cho, 1981-11-29, K. Inukai 5127

1932-11-6, H. Yamamoto s.n. (TNS-VS-227978); Tanabe-cho,(TNS-VS-545669); Atsumi-cho, 1976-7-18, M. Fukuhara 6052

1975-9-28, Z. Tsujimoto 2236 (TNS-VS-423315); Kitakuwada-

K.

Inukai

4211

(TNS-

gun, Keihoku-cho, 1981-8-15, N. Yoshikawa 8923 (TNS-

VS-432953); Mie Pref.: Kitamuro-gun, Owase-cho, 1953-8-15, K.

VS-1157093); Miyama-cho, 1981-8-15, N. Yoshikawa 8979 (TNS-

Seto 4154 (TNS-VS-129853); Ueno-shi, 1981-9-15, S. Yamauchi

VS-1157090), 1981-8-15, N. Yoshikawa 8952 (TNS-VS-1157089);

s.n. (TNS-VS-1166269); Suzuka-shi, 1983-10-28, K. Kada 10578

Hosono-mura, 1935-10-20, Y. Araki 13815 (TNS-VS-194864);(TNS-VS-1156148); Nabari-shi, 1980-8-30, S. Yamauchi s.n.

Funai-gun, Mizuho-cho, 1983-8-8, T. Nakaike s.n. (TNS-(TNS-VS-437418); Owase-shi, 1959-10-23, H. Ito s.n. (TNS-

VS-1157023, 1157025); Amata-gun, Fukuchiyama-cho, 1928-9-

VS-1045388); Toba-shi, 1976-10-14, Y. Kimizuka s.n. (TNS-

27, Y. Araki 6167 (TNS-VS-196020); Otagi-gun, Shizuichino-

VS-1167448); Kumano-shi, 1975-11-20, K. Hanajiri s.n. (TNS-

mura, 1933-9-17, H. Yamamoto 1899 (TNS-VS-227977); Osaka

VS-437425), 1980-11-23, H. Manago s.n. (TNS-VS-1166218);

Pref.: Suita-shi, 1981-9-16, A. Komaki s.n. (TNS-VS-1166266);

Inabe-gun, Hokusei-cho, 1969-10-10, M. Kawazoe s.n. (TNS-

Takatsuki-shi, 1981-10-11, A. Komaki s.n. (TNS-VS-1166264),

VS-1046239); Mie-gun, Komono-cho, 1980-10-10, S. Yamauchi

1981-9-18, A. Komaki s.n. (TNS-VS-1166263); Kaizuka-shi,s.n. (TNS-VS-437417); Suzuka-gun, Seki-cho, 1979-7-15, M.

1978-10-22, H. Manago s.n. (TNS-VS-549171); Toyono-gun, Toy-

Fukuhara 7539 (TNS-VS-437411); Age-gun, Ano-cho, 1980-7-17,

ono-cho, 1978-11-2, T. Shiraiwa 9188 (TNS-VS-1166267), 1978-

S. Yamauchi s.n. (TNS-VS-437421); Ichishi-gun, Hakusan-cho,

11-12, T. Shiraiwa 9210 (TNS-VS-1166221); Nose-cho, 1981-9-

1974-9-21, Y. Higuchi s.n. (TNS-VS-437409); Ureshino-cho,

27, A. Komaki s.n. (TNS-VS-1166262); Minoo-mura, 1932-10-31,

1979-11-11, S. Yamauchi s.n. (TNS-VS-437413), 1980-11-24, S.

H. Yamamoto 1877 (TNS-VS-227967); Minamikawachi-gun, Chi-

Yamauchi s.n. (TNS-VS-437416); Misugi-mura, 1970-10-4, H. Ito

hayaakasaka-mura, 1976-8-16, Y. Inoue s.n. (TNS-VS-1166265);s.n. (TNS-VS-1042054, 1045520), 1976-11-21, T. Nakaike s.n.

Hyogo Pref.: Kobe-shi, 1979-9-15, T. Shiraiwa 9892 (TNS-(TNS-VS-437403), 1984-9-2, K. Kada s.n. (TNS-VS-1157014);

VS-509264), 1977-12-3, T. Shiraiwa 7649 (TNS-VS-563791);

Iinan-gun, Iitaka-cho, 1978-11-18, Y. Higuchi s.n. (TNS-

Sumoto-shi,

VS-437408), 1980-9-15, S. Yamauchi s.n. (TNS-VS-437419);

Toyooka-shi, 1981-7-29, Y. Hayashi s.n. (TNS-VS-563478);

Taki-gun, Miyagawa-mura, 1979-8-28, Y. Higuchi s.n. (TNS-

Kawanishi-shi, 1981-9-27, A. Komaki s.n. (TNS-VS-576886);

VS-437427), 1961-9-23, Y. Higuchi 3614 (TNS-VS-437426);

Sanda-shi, 1977-11-20, T. Shiraiwa 7605 (TNS-VS-563473); Kaw-

Watarai-gun, Futami-cho, 1980-12-14, S. Yamauchi s.n. (TNS-

abe-gun, Inagawa-cho, 2001-10-14, K. Ushijima 21355 (TNS-

VS-437414); Ochiyama-mura, 1980-10-28, Y. Higuchi 13366

VS-756413); Taka-gun, Kurodasho-cho, 1980-11-9, M. Inada(TNS-VS-437422), 1976-11-21, T. Waku 5448 (TNS-VS-437404);

40782 (TNS-VS-563776); Sayo-gun, Sayo-cho, 1978-8-29, T.

Watarai-cho, 1980-11-30, S. Yamauchi s.n. (TNS-VS-437415);

Shiraiwa 8693 (TNS-VS-509261); Nanko-cho, 1963-8-2, H. Ito

Naga-gun, Aoyama-cho, 1979-10-21, S. Yamauchi s.n. (TNS-s.n. (TNS-VS-1045387); Shiso-gun, Yamasaki-cho, 1977-11-3, T.

VS-437412); Kitamuro-gun, Kiinagashima-cho, 1972-8-24, Y.

Shiraiwa 7581 (TNS-VS-563783), 1978-11-19, T. Shiraiwa 8949

Higuchi s.n. (TNS-VS-437405); Minamimuro-gun, Mihama-cho,(TNS-VS-509263); Yasutomi-cho, 1978-9-23, T. Shiraiwa 8796

1967-8-11, K. Hanajiri s.n. (TNS-VS-437424); Shiga Pref.: Otsu-(TNS-VS-509262); Kinosaki-gun, Hidaka-cho, 1973-10-18, Y.

shi, 1908-8-0, I. Sono s.n. (TNS-VS-17586); Shiga-gun, Shiga-

Hayashi s.n. (TNS-VS-1042052), 1978-8-10, Y. Hayashi s.n.

cho, 1977-9-24, T. Waku 7640 (TNS-VS-533923), 1932-11-13, Y.

(TNS-VS-576887); Izushi-gun, Tanto-cho, 1981-8-19, Y. Hayashi(TNS-VS-432922),

1980-11-23,

1980-8-15,

K.

Mune

326

(TNS-VS-563481);

74

Atsushi Ebihara and Narumi Nakato
s.n. (TNS-VS-563476); Mikata-gun, Mikata-cho, 1977-9-15, T.

1981-9-23, M. Iwanaga s.n. (TNS-VS-433510), 1981-9-20, K.

Shiraiwa 7337 (TNS-VS-563781); Yabu-gun, Yabu-cho, 1978-8-

Yasuhara s.n. (TNS-VS-433511); Hino-gun, Nichinan-cho, 1980-

30, Y. Hayashi s.n. (TNS-VS-576889); Oya-cho, 1981-9-1, Y.

9-15, M. Aoto s.n. (TNS-VS-433507), 1981-9-22, M. Aoto s.n.

Hayashi s.n. (TNS-VS-563480); Asago-gun, Ikuno-cho, 1981-9-(TNS-VS-433515), 1980-10-21, M. Aoto s.n. (TNS-VS-433505);

14, Y. Hayashi s.n. (TNS-VS-563472); Wadayama-cho, 1977-8-4,

Hino-cho, 1981-8-14, M. Aoto s.n. (TNS-VS-433513); Shimane

Y. Hayashi s.n. (TNS-VS-576890); Taki-gun, Sasayama-cho, 1954-

Pref.: Kanoashi-gun, Kakinoki-mura, 1958-9-2, S. Okuyama & H.

10-24, T. Kodama s.n. (TNS-VS-129840); Nara Pref.: Nara-shi,

Utsumi 11788 (TNS-VS-258853), 1958-9-1, S. Okuyama & H.

1937-8-20, Y. Matsumura s.n. (TNS-VS-66718), 1975-7-5, Z.

Utsumi 11794 (TNS-VS-258838); Okayama Pref.: Okayama-shi,

Tsujimoto 738 (TNS-VS-423314), 1979-9-16, Z. Tsujimoto 5203

1981-10-14, H. Kobatake s.n. (TNS-VS-503146), 1980-9-2, H.

(TNS-VS-423331); Yamatokoriyama-shi, 1979-10-15, Z. Tsujimoto

Kobatake s.n. (TNS-VS-423258), 1980-12-10, H. Kobatake s.n.

5235 (TNS-VS-423333); Sakurai-shi, 1979-8-19, A. Komaki s.n.

(TNS-VS-423243),(TNS-VS-1166189); Gojo-shi, 1977-10-30, Z. Tsujimoto 3788

VS-1156982), 1980-10-22, H. Kobatake s.n. (TNS-VS-1172267);(TNS-VS-423332); Gose-shi, 1975-9-22, K. Mimoro, S. Tsugaru &Kurashiki-shi, 1981-8-10, H. Kobatake s.n. (TNS-VS-503144);

K. Mizoguchi s.n. (TNS-VS-371109); Yamabe-gun, Tsuge-mura,

Tsuyama-shi, 1980-11-6, H. Kobatake s.n. (TNS-VS-423239),

1951-8-7, T. Kodama s.n. (TNS-VS-247334); Uda-gun, Muro-

1981-8-14, T. Saito s.n. (TNS-VS-1172273); Kasaoka-shi, 1982-

mura, 1979-8-24, Z. Tsujimoto s.n. (TNS-VS-423316); Kitakat-

11-23, K. Yasuhara s.n. (TNS-VS-1157029); Soja-shi, 1981-11-14,

suragi-gun, Taima-cho, 1981-6-7, Z. Tsujimoto 5773 (TNS-

H. Kobatake s.n. (TNS-VS-503138); Takahashi-shi, 1977-7-27, T.

VS-573810); Yoshino-gun, Yoshino-cho, 1902-10-15, J. Nikai

Watanabe s.n. (TNS-VS-1172264); Niimi-shi, 1982-9-5, T. Saito

1605

s.n. (TNS-VS-1157031); Bizen-shi, 1980-11-30, H. Kobatake s.n.

(TNS-VS-254664);

Nosegawa-mura,

1979-9-23,

O.

1983-10-20,

H.

Kobatake

s.n.

(TNS-

Yanagawa s.n. (TNS-VS-573812); Totsukawa-mura, 1976-10-24,(TNS-VS-423242),

Z. Tsujimoto 3622 (TNS-VS-423317); Wakayama Pref.: Hashi-

VS-503142), 1987-8-4, T. Saito s.n. (TNS-VS-1172274); Akaiwa-

moto-shi, 1978-10-29, H. Manago s.n. (TNS-VS-549150); Shingu-

gun, Seto-cho, 1956-8-19, I. Okubo 9 (TNS-VS-1042125);shi, 1976-10-22, A. Nakajima s.n. (TNS-VS-549158); Kaiso-gun,

Kumayama-cho, 1981-8-4, T. Saito s.n. (TNS-VS-503137); Wake-

Shimotsu-cho, 1976-8-31, H. Manago s.n. (TNS-VS-549137);

gun, Hinase-cho, 1980-9-15, T. Saito s.n. (TNS-VS-423259);

Naga-gun, Uchida-cho, 1981-11-3, Sh. Yamamoto s.n. (TNS-

Yoshinaga-cho, 1980-9-21, T. Saito s.n. (TNS-VS-423245), 1977-

VS-431504); Ito-gun, Katsuragi-cho, 1974-6-2, H. Manago s.n.

10-16, T. Saito s.n. (TNS-VS-423249), 1978-9-24, T. Saito s.n.

(TNS-VS-549147); Koya-cho, 1942-10-19, Y. Satake & S.

(TNS-VS-1172266); Wake-cho, 1983-8-28, H. Kobatake s.n.

Okuyama 24953 (TNS-VS-294052), 1979-8-5, M. Matsuoka s.n.

(TNS-VS-1156225); Oku-gun, Ushimado-cho, 1980-11-17, H.

T.

Kodamas.n.

1981-10-22,

H.

Kobatake

s.n.

(TNS-

(TNS-

Kobatake s.n. (TNS-VS-423244); Oku-cho, 1989-11-26, H.

VS-129843), 1952-10-19, K. Seto 3290 (TNS-VS-130347), 1973-

Kobatake s.n. (TNS-VS-1156178); Asakuchi-gun, Konko-cho,

8-25, H. Manago s.n. (TNS-VS-549146); Arita-gun, Yuasa-cho,

1984-8-16, H. Kobatake s.n. (TNS-VS-1156168); Kamogata-cho,

1977-10-10, H. Manago s.n. (TNS-VS-549157); Shimizu-cho,

1957-12-2, I. Akazawa 39 (TNS-VS-1042049); Satosho-cho, 1980-

1977-10-9, H. Manago s.n. (TNS-VS-549140), 1971-10-24, H.

11-3, K. Yasuhara s.n. (TNS-VS-423241); Jobo-gun, Kayo-cho,Manago s.n. (TNS-VS-549138); Hidaka-gun, Nakatsu-mura,

1980-10-27, H. Kobatake s.n. (TNS-VS-423263); Takahashi-cho,

1974-5-3, H. Manago s.n. (TNS-VS-549152); Inami-cho, 1976-11-

1905-10-?, Z. Yoshino s.n. (TNS-VS-517187); Kawakami-gun,

28, H. Manago s.n. (TNS-VS-549161); Nishimuro-gun, Shira-Nariwa-cho, 1975-9-10, T. Watanabe s.n. (TNS-VS-334509),

hama-cho, 1977-8-31, H. Manago s.n. (TNS-VS-549129); Naka-

1978-8-22, T. Saito s.n. (TNS-VS-1172263); Atetsu-gun, Osa-cho,hechi-cho, 1977-7-17, H. Manago s.n. (TNS-VS-549163);

1961-8-18, T. Watanabe s.n. (TNS-VS-423253); Tessei-cho, 1976-

Oto-mura, 1973-9-11, H. Manago s.n. (TNS-VS-549164), 1967-9-

8-6, T. Watanabe s.n. (TNS-VS-1172265); Maniwa-gun, Ochiai-7, H. Manago s.n. (TNS-VS-549162); Susami-cho, 1974-8-10, H.

cho, 1974-8-25, T. Saito s.n. (TNS-VS-423250), 1981-11-5, H.

Manago s.n. (TNS-VS-549132, 549133); Kushimoto-cho, 1977-

Kobatake s.n. (TNS-VS-1156179), 1989-11-5, H. Kobatake s.n.

12-3, H. Manago s.n. (TNS-VS-549127); Higashimuro-gun,(TNS-VS-1156180); Kuse-cho, 1984-11-2, H. Kobatake s.n.

Nachikatsuura-cho, 1980-8-2, K. Ohora s.n. (TNS-VS-549172);(TNS-VS-1157020),

Kozagawa-cho, 1956-10-16, H. Ito s.n. (TNS-VS-1042128), 1979-

VS-1157021); Tomata-gun, Aba-son, 1980-8-21, K. Takayama s.n.

9-24, H. Manago s.n. (TNS-VS-549134); Kumanogawa-cho,(TNS-VS-423256); Kagamino-cho, 1981-8-14, T. Saito s.n. (TNS-

1973-8-13, H. Manago s.n. (TNS-VS-549166); Nachi-cho, 1942-

VS-503145); Katsuta-gun, Shoboku-cho, 1984-11-5, H. Kobatake

10-13, Y. Satake & S. Okuyama 24952 (TNS-VS-294051); Tottoris.n. (TNS-VS-1157019, 1157022); Aida-gun, Nishiawakura-son,

Pref.: Tottori-shi, 1980-9-23, A. Tanaka 28254 (TNS-VS-433508),

1984-8-24, H. Kobatake s.n. (TNS-VS-1156167); Mimasaka-cho,

1981-9-23, A. Tanaka s.n. (TNS-VS-433516); Iwami-gun, Kokufu-

1982-9-16, H. Kobatake s.n. (TNS-VS-1157010); Kume-gun,cho, 1981-10-25, A. Tanaka s.n. (TNS-VS-433514), 1981-8-15, A.

Kumenan-cho, 1981-11-12, H. Kobatake s.n. (TNS-VS-503140);

Tanaka s.n. (TNS-VS-433504); Yazu-gun, Wakasa-cho, 1955-9-12,

Hiroshima Pref.: Hiroshima-shi, 1978-10-19, M. Matsumura s.n.

Y. Ikoma & M. Kawamoto 32 (TNS-VS-1042112); Mochigase-cho,(TNS-VS-547831); Kure-shi, 1977-11-3, T. Takeda 10901 (TNS-

1981-10-18, M. Iwanaga s.n. (TNS-VS-433509); Saji-son, 1955-9-

VS-547854); Takehara-shi, 1990-8-17, M. Matsumura s.n. (TNS-6, Y. Ikoma & M. Kawamoto 62 (TNS-VS-1042051); Chizu-cho,VS-1156139); Mihara-shi, 1977-12-11, T. Takeda 10963 (TNS-(TNS-VS-1053154),

1950-8-25,

1984-11-2,

H.

Kobatake

s.n.

(TNS-

Distribution of the Thelypteris japonica Complex (Thelypteridaceae)
1071

H. Inoue 1721 (TNS-VS-1042136); Kagawa Pref.: Takamatsu-shi,(TNS-VS-1165538); Higashihiroshima-shi, 1977-11-19, T. Takeda

1980-7-6, S. Mitani s.n. (TNS-VS-527149); Marugame-shi, 1980-

10955 (TNS-VS-1165529); Aki-gun, Kamagari-cho, 1995-8-24,

7-13, S. Mitani s.n. (TNS-VS-527148); Kanonji-shi, 1969-10-22,

M. Matsumura s.n. (TNS-VS-1157039); Saeki-gun, Yuki-cho,

Y. Maki 3 (TNS-VS-257032); Okawa-gun, Shirotori-cho, 1980-10-

1974-9-10, T. Takeda s.n. (TNS-VS-1165540), 1976-10-10, M.

19, S. Mitani s.n. (TNS-VS-511283); Nagao-cho, 1974-12-1, S.

Matsumura s.n. (TNS-VS-547833); Saeki-cho, 1978-7-24, M.

Mitani s.n. (TNS-VS-527147); Ayauta-gun, Ryonan-cho, 1976-9-

Matsumura s.n. (TNS-VS-547830); Yoshiwa-mura, 1989-9-17, M.

16, S. Mitani s.n. (TNS-VS-527142); Nakatado-gun, Kotonami-

Matsumura s.n. (TNS-VS-1156990); Asa-gun, Asa-cho, 1969-11-cho, 1976-10-16, S. Mitani s.n. (TNS-VS-527144); Kotohira-cho,2, H. Okada 312 (TNS-VS-1053174), 1969-11-5, H. Okada 395

1974-11-17, S. Mitani s.n. (TNS-VS-527141); Mitoyo-gun,(TNS-VS-1053175); Yamagata-gun, Tsutsuga-son, 1977-10-22, T.

Takase-cho, 1976-11-21, S. Mitani s.n. (TNS-VS-527143); Ehime

Takeda 10736 (TNS-VS-547851); Togochi-cho, 1974-9-25, T.

Pref.: Matsuyama-shi, 1979-10-21, H. Ishikawa s.n. (TNS-

Takeda 2858 (TNS-VS-1165533); Geihoku-cho, 1977-8-6, T.

VS-1166275), 1979-11-1, H. Ishikawa s.n. (TNS-VS-1166278);

Takeda 10151 (TNS-VS-547852); Oasa-cho, 1976-8-8, T. Takeda

Uwajima-shi, 1978-8-16, C. Aoki s.n. (TNS-VS-1166290), 1987-9-

7721 (TNS-VS-547846); Toyohira-cho, 1976-8-8, T. Takeda 7749

20, M. Hyodo s.n. (TNS-VS-1156198); Niihama-shi, 1959-11-5,(TNS-VS-1165535); Kamo-gun, Daiwa-cho, 1977-10-31, T.

H. Ishikawa 49 (TNS-VS-1042127), 1959-11-1, H. Ishikawa 75

Takeda 10874 (TNS-VS-547858); Kochi-cho, 1978-11-16, T.

(TNS-VS-1042121); Ozu-shi, 1978-7-18, H. Ishikawa s.n. (TNS-

Takeda 13371 (TNS-VS-547862), 1977-10-12, T. Takeda 10575

VS-1166279), 1979-8-31, H. Ishikawa s.n. (TNS-VS-1166272),

VS-1165528);

Fukuyama-shi,T.

1973-10-4,

Takeda75(TNS-VS-547848); Jinseki-gun, Jinseki-cho, 1973-10-23, T.

1978-6-13, H. Ishikawa s.n. (TNS-VS-1166277); Iyomishima-shi,

Takeda 1202 (TNS-VS-1165534); Futami-gun, Kisa-cho, 1973-8-

1987-10-7, M. Hyodo s.n. (TNS-VS-1156197); Iyo-shi, 1981-11-3,

31, T. Takeda 286 (TNS-VS-547860); Miwa-cho, 1975-9-15, T.

H. Ishikawa s.n. (TNS-VS-1166286); Hojo-shi, 1980-9-30, H.

Takeda 4672 (TNS-VS-547853); Hiba-gun, Hiwa-cho, 1993-9-25,

Ishikawa s.n. (TNS-VS-1166270); Uma-gun, Doi-mura, 1953-8-4,

M. Matsumura s.n. (TNS-VS-1156176); Yamaguchi Pref.:H. Inoue 1050 (TNS-VS-1042119); Ochi-gun, Tamagawa-cho,Tokuyama-shi, 1980-10-12, A. Minami 17337 (TNS-VS-424218);

1978-10-10, H. Ishikawa s.n. (TNS-VS-1166273); Onsen-gun,

Hofu-shi, 1991-11-5, H. Masaki s.n. (TNS-VS-1156136), 1991-11-

Shigenobu-cho, 1980-9-15, H. Ishikawa s.n. (TNS-VS-1166274);

3, H. Masaki s.n. (TNS-VS-1156138), 1991-10-15, H. Masaki s.n.

Kawauchi-cho, 1981-10-25, M. Hyodo s.n. (TNS-VS-1166292),(TNS-VS-1156140); Iwakuni-shi, 1980-9-21, A. Minami 16677

1979-10-5, H. Ishikawa s.n. (TNS-VS-1166276); Kamiukena-gun,(TNS-VS-397231); Kuga-gun, Shuto-cho, 1969-10-6, H. Masaki

Omogo-mura, 1965-9-28, S. Masuda 4504 (TNS-VS-831696),

16997 (TNS-VS-262904); Miwa-cho, 1956-10-28, M. Wada 4287

1953-8-17, H. Inoue 1445 (TNS-VS-1042114), 1953-8-17, H.

A.

Minami

17732

(TNS-

Inoue 1443 (TNS-VS-1042131); Mikawa-mura, 1981-7-13, H.

VS-424217); Tsuno-gun, Kano-cho, 1980-9-23, A. Minami 16766

Ishikawa s.n. (TNS-VS-1166284), 1979-11-5, H. Ishikawa s.n.

(TNS-VS-397215); Saba-gun, Tokuji-cho, 1980-9-23, A. Minami(TNS-VS-1166285); Yanadani-mura, 1981-9-15, H. Ishikawa s.n.

16799 (TNS-VS-397206); Yasaka-mura, 1920-9-20, J. Nikai s.n.

(TNS-VS-1166271), 1979-8-14, H. Ishikawa s.n. (TNS-VS-(TNS-VS-254665), 1920-9-20, J. Nikai 2811 (TNS-VS-50385);

1166283), 1981-9-12, M. Hyodo s.n. (TNS-VS-1166294); Oda-

Toyora-gun, Toyora-cho, 1982-10-10, G. Imada 6373 (TNS-cho, 1978-10-15, H. Ishikawa s.n. (TNS-VS-1166281); Iyo-gun,

VS-569259); Toyotashimo-mura, 1919-8-23, J. Nikai s.n. (TNS-

Nakayama-cho, 1981-11-7, M. Hyodo s.n. (TNS-VS-1166291);

VS-254663); Mine-gun, Mito-cho, 1981-9-13, S. Miyake s.n.

Higashiuwa-gun, Nomura-cho, 1980-11-9, M. Hyodo s.n. (TNS-(TNS-VS-424048); Shuho-cho, 1975-8-30, S. Miyake s.n. (TNS-

VS-1166288); Shirokawa-cho, 1981-9-15, C. Aoki s.n. (TNS-

VS-732958); Abu-gun, Asahi-son, 1981-8-2, S. Miyake s.n. (TNS-

VS-1166289); Kitauwa-gun, Matsuno-cho, 1983-11-27, M. Hyodo

VS-424047).

s.n. (TNS-VS-1156208); Hiyoshi-mura, 1982-10-10, M. Hyodo(TNS-VS-251199),

1980-10-30,

Shikoku - Tokushima Pref.: Anan-shi, 1961-8-14, C. Abe 17752

s.n. (TNS-VS-1156132); Tsushima-cho, 1981-10-6, M. Hyodo s.n.

(TNS-VS-596025); Naka-gun, Kaminaka-cho, 1965-8-16, H. Ito(TNS-VS-1166293); Nii-gun, Ofuki-mura, 1953-8-5, H. Inoue

s.n.

(TNS-

1388 (TNS-VS-1042043); Kochi Pref.: Kochi-shi, 1975-11-23, Y.

VS-1045380); Kito-son, 1980-8-16, T. Nakayama s.n. (TNS-

Koukami s.n. (TNS-VS-938069); Nankoku-shi, 1981-8-17, K.

VS-596020); Kamodani-son, 1955-8-3, T. Inobe 40 (TNS-

Yamaoka & Y. Yamamoto s.n. (TNS-VS-503760), 1981-8-17, K.

VS-1042040); Kaifu-gun, Yuki-cho, 1970-11-29, C. Abe 43427

Yamaoka & Y. Yamamoto s.n. (TNS-VS-503756); Aki-gun, Umaji-(TNS-VS-596027); Hiwasa-cho, 1960-8-23, C. Abe 17751 (TNS-

mura, 1955-8-23, S. Iwano s.n. (TNS-VS-938070); Kami-gun,

VS-596026); Mugi-cho, 1979-9-23, T. Nakayama s.n. (TNS-

Tosayamada-cho, 1974-10-18, T. Nakaike s.n. (TNS-VS-423544);

VS-596018); Kainan-cho, 1959-8-18, C. Abe 17757 (TNS-

Makiyama-mura, 1954-8-24, H. Inoue 2097 (TNS-VS-1042117),(TNS-VS-1045391),

1965-8-18,

H.

s.n.

s.n.

1954-8-24, H. Inoue 2096 (TNS-VS-1042123); Nagaoka-gun,(TNS-VS-596017); Oe-gun, Kamojima-cho, 1962-8-30, C. Abe

Otoyo-cho, 1981-9-15, K. Yamaoka 19605 (TNS-VS-503761),

17756 (TNS-VS-596021); Kawata-son, 1911-8-27, J. Nikai 2104

1978-10-21, K. Yamaoka s.n. (TNS-VS-938079); Gomen-cho,(TNS-VS-50384); Mima-gun, Waki-machi, 1963-8-3, C. Abe

1913-8-26, Z. Tashiro s.n. (TNS-VS-23346); Hongawa-mura,

17755 (TNS-VS-596022); Miyoshi-gun, Sammyo-son, 1956-8-21,

1950-9-2, S. Okuyama 11437 (TNS-VS-279146); Takaoka-gun,

T. Inobe 97 (TNS-VS-1042041); Higashiiyayama-son, 1954-7-20,

Nakatosa-cho, 1979-9-?, T. Miyazaki s.n. (TNS-VS-938160);

VS-596029);

Shishikui-cho,

1977-8-3,

T.

Ito

Nakayama

76

Atsushi Ebihara and Narumi Nakato
Ochi-cho, 1974-10-25, T. Nakaike s.n. (TNS-VS-333285), 1949-8-

1977-11-7, Y. Ishizaka s.n. (TNS-VS-1172272); Yatsushiro-gun,

13, H. Inoue 310 (TNS-VS-1042132); Niida-mura, coll. dateIzumi-mura, 1974-10-10, N. Beppu s.n. (TNS-VS-550091), 1973-

unknown, H. Inoue 475 (TNS-VS-1042122); Kubokawa-cho,

8-18, N. Beppu s.n. (TNS-VS-550089, 580175); Ashikita-gun,2002-9-15, K. Yamaoka & N. Nakato 2480 (TNS-VS-1168487),

Ashikita-machi, 1978-12-29, N. Beppu s.n. (TNS-VS-550082);

2002-9-15, K. Yamaoka & N. Nakato 2478 (TNS-VS-1168486),

Kuma-gun, Mizukami-mura, coll. date unknown, S. Hattori s.n.

2002-9-15, K. Yamaoka & N. Nakato 2479 (TNS-VS-1168485);(TNS-VS-784684),

K.

Yamaokas.n.

T.

1978-8-1,

Nakamura

s.n.

(TNS-

(TNS-

VS-1016993, 1016994); Itsuki-mura, 1981-8-15, N. Beppu s.n.

VS-938072); Hata-gun, Saga-cho, 1993-10-19, T. Miyazaki 2022

(TNS-VS-550086); Oita Pref.: Beppu-shi, 1960-9-25, M. Arakane(TNS-VS-1156175); Nishitosa-mura, 1981-8-22, Y. Akasawa, T.

s.n. (TNS-VS-437313); Usa-shi, 1976-9-4, M. Arakane s.n. (TNS-

Miyazaki & Y. Yamamoto s.n. (TNS-VS-938076).

VS-437311); Oita-gun, Shonai-cho, 1960-7-29, M. Arakane s.n.

Higashitsuno-mura,

1978-9-24,

Kyushu - Fukuoka Pref.: Kitakyushu-shi, 1976-9-23, S. Kobayashi(TNS-VS-437319), 1959-11-8, M. Hadano s.n. (TNS-VS-437325);

6534 (TNS-VS-431748); Yahatahigashi-ku, 1977-8-15, K. Miura

Minamiamabe-gun, Ume-machi, 1963-1-6, Y. Takaoka s.n. (TNS-

177 (TNS-VS-428991); Fukuoka-shi, 1966-9-15, S. Tsutsui 2919

VS-437310), 1959-8-3, M. Hadano s.n. (TNS-VS-437324); Ono-

(TNS-VS-428993); Kurume-shi, 1969-8-22, S. Kobayashi 2376

gun, Ogata-machi, 1975-8-3, M. Arakane s.n. (TNS-VS-437322),(TNS-VS-429000); Chikushi-gun, Nakagawa-machi, 1973-8-19, S.

1955-8-?, M. Hadano s.n. (TNS-VS-437308, 1157098); Naoiri-

Tsutsui 9550 (TNS-VS-428995); Kasuya-gun, 1947-10-17, S.

gun, Kuju-machi, 1968-8-9, K. Shono 1072 (TNS-VS-286022);

Toyama 408 (TNS-VS-1042068); Umi-machi, 1967-9-17, S. Tsut-Kusu-gun, Kokonoe-machi, 1955-8-?, M. Hadano s.n. (TNS-sui 4605 (TNS-VS-428992); Yame-gun, Hoshino-mura, 1976-8-9,

VS-1157096); Kusu-machi, 1973-9-24, M. Arakane s.n. (TNS-

S. Tsutsui 13139 (TNS-VS-428998); Tagawa-gun, Soeda-machi,

VS-437312); Hita-gun, Kamitsue-mura, 1978-8-3, N. Beppu s.n.

1967-7-25, H. Ito s.n. (TNS-VS-1045375), 1952-10-29, J. Ouchi 4(TNS-VS-435065); Shimoge-gun, Yamakuni-machi, 1984-8-19, T.

(TNS-VS-1042063), 1952-8-7, J. Ouchi 59 (TNS-VS-1042079);

Yamanaka s.n. (TNS-VS-734111), 1977-8-26, Y. Takaoka s.n.

Saga Pref.: Tosu-shi, 1975-9-14, Y. Kuranari s.n. (TNS-(TNS-VS-437315); Usa-gun, Innai-machi, 1960-8-20, M. Arakane

VS-579330); Saga-gun, Fuji-cho, 1980-7-26, M. Sadamatsu s.n.

s.n. (TNS-VS-437320), 1960-8-26, M. Hadano s.n. (TNS-(TNS-VS-579333); Kanzaki-gun, Mitsuse-mura, 1981-11-8, H.

VS-437306); Miyazaki Pref.: Miyakonojo-shi, 1926-8-0, A.

Inoue s.n. (TNS-VS-425079); Ogi-gun, Ogi-machi, 1977-6-25, T.

Noguchi s.n. (TNS-VS-666824); Nobeoka-shi, 1981-8-24, S.

Baba s.n. (TNS-VS-579336); Higashimatsuura-gun, Nanayama-

Tsutsui 21405 (TNS-VS-588348); Kobayashi-shi, 1976-8-15, T.

mura, 1978-9-21, M. Koike s.n. (TNS-VS-579334); Ochi-cho,

Minamitani 23561 (TNS-VS-427808); Saito-shi, 1959-8-27, I.

1975-8-17, Y. Tanno s.n. (TNS-VS-579335); Nishimatsuura-gun,Taki s.n. (TNS-VS-588346); Ebino-shi, 1979-7-22, T. Minamitani

Arita-machi, 1944-10-23, S. Toyama s.n. (TNS-VS-100268);

31841 (TNS-VS-427816), 1979-10-20, T. Minamitani 31832

Nishiarita-cho, 1975-8-24, Y. Tanno s.n. (TNS-VS-579715);(TNS-VS-427815), 1979-7-15, T. Minamitani 31743 (TNS-

Fujitsu-gun, Tara-cho, 1961-8-14, B. Matsubayashi s.n. (TNS-

VS-427817);

VS-1166217); Nagasaki Pref.: Nishisonogi-gun, Kinkai-cho,

Minamitani 26957 (TNS-VS-427812); Kobayashi-machi, 1918-7-

1981-11-19, B. Matsubayashi s.n. (TNS-VS-579384), 1981-8-20,

24, collector unknown s.n. (TNS-VS-33211); Koyu-gun, Taka-

B. Matsubayashi s.n. (TNS-VS-579383); Higashisonogi-gun,nabe-cho, 1980-12-27, K. Nakamura s.n. (TNS-VS-588347); Shin-

Hasami-cho, 1976-7-30, F. Ueno s.n. (TNS-VS-579388); Minami-

tomi-cho, 1973-9-23, T. Minamitani 9331 (TNS-VS-427805);

takaki-gun, Obama-cho, 1961-10-15, B. Matsubayashi s.n. (TNS-

Tsuno-cho, 1977-10-16, T. Minamitani 28437 (TNS-VS-427810);

VS-579386); Minamimatsuura-gun, Ohama-cho, 1934-8-4, T.

Higashiusuki-gun, Nango-son, 1967-5-6, T. Minamitani 1277

Baba s.n. (TNS-VS-423524); Kumamoto Pref.: Kumamoto-shi,(TNS-VS-427803); Kitagawa-cho, 1986-8-9, A. Minami & U.

1973-10-10, N. Beppu s.n. (TNS-VS-550092); Hitoyoshi-shi,

Fujiyama 51575 (TNS-VS-1166301); Kagoshima Pref.: Kanoya-

1968-11-5, H. Ito s.n. (TNS-VS-1045607, 1045631), 1964-7-11,

shi, 1980-10-10, K. Takesako s.n. (TNS-VS-434213); Izumi-shi,

H. Ito s.n. (TNS-VS-1045692); Minamata-shi, 1962-10-7, T.

1963-8-18, T. Yamanaka s.n. (TNS-VS-735651), 1974-8-31, M.

Yamanaka s.n. (TNS-VS-779514), 1960-10-25, M. Kido s.n. (TNS-

Kido s.n. (TNS-VS-433985); Okuchi-shi, 1960-11-4, T. Yamanaka

VS-550103), 1960-10-23, M. Kido 3473 (TNS-VS-550084), 1962-

s.n. (TNS-VS-734241), 1960-9-4, T. Yamanaka s.n. (TNS-

11-18, M. Kido s.n. (TNS-VS-1172269); Tamana-gun, Gyokuto-

VS-511389, 734321), 1981-10-4, T. Yamanaka s.n. (TNS-

machi, 1981-10-10, Y. Ishizaka s.n. (TNS-VS-533580); Aso-gun,

VS-741296), 1963-8-16, T. Yamanaka s.n. (TNS-VS-1112895);

Ichinomiya-machi, 1972-8-23, N. Beppu s.n. (TNS-VS-550093);

Isa-gun, Hishikari-cho, 1980-9-28, K. Takesako s.n. (TNS-

Aso-machi, 1976-11-21, N. Beppu s.n. (TNS-VS-550098); Mina-

VS-434212); Aira-gun, Mizobe-cho, 1980-10-5, K. Takesako s.n.

mioguni-machi, 1977-7-23, N. Beppu s.n. (TNS-VS-550095);(TNS-VS-434215); Kurino-cho, 1980-9-27, K. Takesako s.n.

Oguni-machi, 1977-8-14, N. Beppu s.n. (TNS-VS-550075); Haku-(TNS-VS-434209); Makizono-cho, 1966-8-9, T. Yamanaka s.n.

sui-mura, 1978-8-12, N. Beppu s.n. (TNS-VS-550100); Kugino-(TNS-VS-1112247),

mura, 1978-8-12, N. Beppu s.n. (TNS-VS-550073); Choyo-mura,

VS-434208), 1980-10-4, K. Takesako s.n. (TNS-VS-434214);

1977-8-25, Y. Ishizaka s.n. (TNS-VS-1172271); Kamimashiki-gun,

Kimotsuki-gun, Uchinora-cho, 1974-7-20, K. Takesako s.n. (TNS-

Mifune-machi, 1976-12-5, N. Beppu s.n. (TNS-VS-550102); Yabe-

VS-434216);

machi, 1978-8-27, N. Beppu s.n. (TNS-VS-550101); Seiwa-son,

Kawabata s.n. (TNS-VS-939698).

Nishimorokata-gun,

1979-8-11,

Kumage-gun,Nojiri-cho,K.

1977-7-29,

TakesakoKamiyaku-cho,

s.n.

T.

(TNS-

1964-7-29,

M.

Distribution of the Thelypteris japonica Complex (Thelypteridaceae)

77

Thelypteris japonica (Baker) Ching forma formosa (C.Chr.)
Nakato, Sahashi et M.Kato [Ao-harigane-warabi]

Y. Takahashi s.n. (TNS-VS-419802); Yuri-gun, Nikaho-machi,

Hokkaido - Hokkaido Pref.: Tomakomai-shi, Nishiki-onuma, 1981-

machi,

Toshi, 1978-9-23, J. Takada s.n. (TNS-VS-419859); ChokaiKoshiki-yama,

1981-9-27,

T.

Kikuchi

s.n.

(TNS-

9-20, M. Ishizuka s.n. (TNS-VS-545559); Chitose-shi, Monbetsu-

VS-419792); Senboku-gun, Kamioka-machi, Hogaku-zawa, 1980-

gawa, 1984-10-8, M. Ishizuka s.n. (TNS-VS-1157052); Shiraoi-

10-10, J. Takada s.n. (TNS-VS-419794); Nishisenboku-machi,

gun, Shiraoi-cho, Shadai-gawa, 1982-10-19, M. Ishizuka s.n.

Teramura, 1980-9-25, J. Takada s.n. (TNS-VS-419809); Kyowa-(TNS-VS-1157054).

machi, Sakai, 1977-10-9, J. Takada s.n. (TNS-VS-419860); Nan-

Honshu - Aomori Pref.: Aomori-shi, Goshi-zawa, 1984-9-4, N.

gai-mura, Yatate Pass, 1937-10-22, G. Koie s.n. (TNS-VS-88735);

Saitoh s.n. (TNS-VS-1156142); Mutsu-shi, Kabayama, 1980-9-15,

Hiraka-gun, Sannai-mura, Kurosawa, 1976-9-13, T. Nakaike s.n.

T. Sudo s.n. (TNS-VS-570344); Usori-gawa, 1980-9-14, E. Fukushi(TNS-VS-420025); Ogachi-gun, Ugo-machi, Tazawa, 1978-9-17,

s.n. (TNS-VS-570343); Higashitsugaru-gun, Yomogita-mura, Seh-

J. Fujiwara s.n. (TNS-VS-419795); Yamagata Pref.: Yamagata-eji, 1983-8-28, N. Saitoh s.n. (TNS-VS-1152598); Shimokita-gun,shi, Chitose-yama, 1953-9-21, I. Igura s.n. (TNS-VS-1042064);

Kawauchi-machi, Tozawa, 1980-9-14, E. Fukushi s.n. (TNS-

Hirashimizu, 1976-10-16, N. Sagawa s.n. (TNS-VS-437941);

VS-570292); Iwate Pref.: Morioka-shi, 1975-9-15, K. Sato s.n.

Yonezawa-shi, Torimachi, 1980-8-28, Z. Sato s.n. (TNS-(TNS-VS-440277); Mizusawa-shi, Ofukasawa, 1978-9-3, M.

VS-437960); Asagawa, 1980-8-28, Z. Sato s.n. (TNS-VS-);

Suzuki s.n. (TNS-VS-440275); Esashi-shi, Hitokabe, 1983-10-8, C.

Shinjo-shi, Torigoe, 1977-9-17, N. Sagawa s.n. (TNS-VS-437939);

Komizunai s.n. (TNS-VS-1152578); Iwate-gun, Shizukuishi-cho,

Sagae-shi, Nagaoka-yama, 1981-8-6, N. Sagawa s.n. (TNS-

Nyuto-san, 1977-9-25, Y. Takano s.n. (TNS-VS-440268); Iwate-

VS-437972); Kaminoyama-shi, Yusaka, 1981-8-5, N. Sagawa s.n.

machi, Yashiki, 1980-8-9, E. Fukushi s.n. (TNS-VS-440283);(TNS-VS-1157138); Iwa-kannon, 1980-8-28, N. Sagawa s.n.

Takizawa-mura, Koiwai Farm, 1945-9-?, Y. Jotani s.n. (TNS-(TNS-VS-437955); Obanazawa-shi, Tokura-ko, 1980-8-24, N.

VS-831871); Waga-gun, Towa-cho, Tase, 1977-7-19, K. Inoue s.n.

Sagawa s.n. (TNS-VS-437942); Ginzan, 1981-9-29, N. Sagawa(TNS-VS-440516); Waga-cho, Geto Spa, 1977-10-6, Y. Takano s.n.

s.n. (TNS-VS-437979); Nanyo-shi, Nakagawa, 1980-9-7, N.

(TNS-VS-440266); Isawa-gun, Kanegasaki-cho, Rokuhara, 1977-

Sagawa s.n. (TNS-VS-437956); Higashimurayama-gun, Chitose-

8-17, M. Suzuki s.n. (TNS-VS-440273); Hosono, 1980-9-22, M.

mura, Chitose-yama, 1931-8-9, S. Okuyama 3710 (TNS-

Suzuki s.n. (TNS-VS-440271); Maesawa-cho, Seibo, 1980-9-15,

VS-663626); Nishimurayama-gun, Nishikawa-machi, Ubasawa-

M. Suzuki s.n. (TNS-VS-440276); Isawa-cho, Keshozaka, 1980-8-

goya, 1981-10-2, N. Sagawa s.n. (TNS-VS-437971); Naganuma,

12, M. Suzuki s.n. (TNS-VS-440274); Koromogawa-mura, Mina-

1981-8-9, N. Sagawa s.n. (TNS-VS-437973); Kitamurayama-gun,mimata, 1980-8-12, M. Suzuki s.n. (TNS-VS-440272); Higashi-

Oishida-machi, Ashizawa, 1977-10-5, N. Sagawa s.n. (TNS-

iwai-gun, Higashiyama-cho, Natsuyama, 1980-7-23, M. Suzuki s.n.

VS-437940); Mogami-gun, Mogami-machi, Nabekura, 1981-10-(TNS-VS-440269); Shimohei-gun, Tanohata-mura, Matsumae-

20, N. Sagawa s.n. (TNS-VS-437975); Mamurogawa-machi,zawa, 1980-8-9, E. Fukushi s.n. (TNS-VS-440282); Miyagi Pref.:Showa, 1963-8-23, S. Kato s.n. (TNS-VS-437614); Tashiro, 1981-

Sendai-shi, Hachiman, 1980-10-27, M. Takeichi s.n. (TNS-

10-27, N. Sagawa s.n. (TNS-VS-437976); Higashiokitama-gun,VS-1172251); Natori-shi, Kitagama, 1981-9-30, K. Shoji s.n.

Kawanishi-machi, Inukawa, 1981-9-8, N. Sagawa s.n. (TNS-(TNS-VS-576831); Kurokawa-gun, Taiwa-cho, Tamagaike, 1981-

VS-437966); Nishiokitama-gun, Shirataka-machi, Ayukai, 1980-9-

8-30, K. Matsunaga s.n. (TNS-VS-1172248); Kami-gun, Nakani-

21, N. Sagawa s.n. (TNS-VS-437957); Narita, 1980-9-21, N.

ida-machi, Jou, 1981-9-6, W. Takahashi s.n. (TNS-VS-1172252);

Sagawa s.n. (TNS-VS-437958); Iide-machi, Hagyu, 1981-9-8, N.

Onoda-machi, Suge-numa, 1976-9-15, W. Takahashi s.n. (TNS-

Sagawa s.n. (TNS-VS-437963); Higashitagawa-gun, Asahi-mura,VS-1172246); Nabekoshi Pass, 1976-8-29, W. Takahashi s.n.

Otori, 1980-10-2, T. Nakaike s.n. (TNS-VS-437954); Akumi-gun,(TNS-VS-1172244); Shikama-cho, Ojo-ji, 1982-9-5, M. Takeichi

Yuza-machi, Onoso, 1973-10-8, S. Kato s.n. (TNS-VS-437613);

s.n. (TNS-VS-1157033); Tome-gun, Hasama-cho, Tatebayashi, alt.

Matsuyama-machi, 1975-9-6, S. Kato s.n. (TNS-VS-437620);

20 m, 1975-8-15, Y. Ueno 13855 (TNS-VS-1172249); Mono-gun,

Fukushima Pref.: Fukushima-shi, Machiniwasaka, Shida, 1976-

Kanan-cho, Wabuchi-yama, 1975-8-24, Y. Ueno 19014 (TNS-

10-10, Z. Kaneda 2574 (TNS-VS-438923); Aizuwakamatsu-shi,

VS-1172247); Asahiyama, 1971-10-29, Y. Ueno 5896 (TNS-

Iimori-yama, 1977-10-22, Z. Kaneda 3555 (TNS-VS-438925);

VS-1160529); Motoyoshi-gun, Shizugawa-cho, Sodehama, 1980-

Koriyama-shi, Myoken-zan, 1977-11-10, Z. Kaneda 3714 (TNS-

9-20, M. Takeichi s.n. (TNS-VS-1172250); Akita Pref.: Akita-shi,

VS-439167); Yatagawa, 1977-10-2, Z. Kaneda 3364 (TNS-

Kana-ashi, 1977-10-5, J. Takada s.n. (TNS-VS-419804); Tegata-VS-438922); Iwaki-shi, Taira, Kamiarakawa, alt. 80 m, 1977-8-11,

yama, 1981-10-19, S. Okita s.n. (TNS-VS-419786); Kazuno-shi,

Y. Yuzawa s.n. (TNS-VS-438914); Uchigo-koyamachi, alt. 250 m,

Towada-kogen, 1975-9-28, J. Takada s.n. (TNS-VS-419803);

1972-10-9, Y. Yuzawa s.n. (TNS-VS-439539); Kitakata-shi, Maki,

Kazuno-gun, Kosaka-machi, Ginzan, 1976-8-25, J. Takada s.n.

1976-10-11, Z. Kaneda s.n. (TNS-VS-438926); Yama-gun, Atsush-(TNS-VS-419805); Kitaakita-gun, Takanosu-machi, Kakarido,

iokano-mura, Sakurayashiki, 1981-9-8, Z. Kaneda 6982 (TNS-

1981-10-18, M. Hatakeyama s.n. (TNS-VS-419789); Hinai-machi,

VS-439168); Atsushio, 1977-10-6, Z. Kaneda 3424 (TNS-

Takko-mori, 1976-10-11, T. Kikuchi s.n. (TNS-VS-419790); Min-

VS-438941); Shiokawa-machi, Baba-shinden, 1980-9-14, S. Saito

amiakita-gun, Showa-machi, Motoki-yama, 1981-10-25, S. Okita

s.n. (TNS-VS-1172241); Inawashiro-machi, Numagami, 1975-11-

s.n. (TNS-VS-419787); Ikawa-machi, Manaita-yama, 1978-8-20,

4, Z. Kaneda s.n. (TNS-VS-438927); Kawanuma-gun, Aizubange-

78

Atsushi Ebihara and Narumi Nakato
machi, Todera, 1977-9-1, Z. Kaneda 3330 (TNS-VS-438924);

toshima-gun, Shi-mura, 1929-9-23, F. Maekawa 1212 (TNS-

Onuma-gun, Showa-mura, 1979-9-10, Z. Kaneda s.n. (TNS-

VS-831867); Musashino-shi, Gotenyama, 1980-9-27, H. Endo s.n.

VS-438920); Higashishirakawa-gun, Tanagura-machi, Tennouchi,(TNS-VS-423916); Kitatama-gun, Yamato-mura, Shimizu, 1937-

1983-10-8, H. Kobayashi s.n. (TNS-VS-1159876); Tsutsukowake

9-26, F. Maekawa s.n. (TNS-VS-831866); Niigata Pref.: Kashiwa-

Shrine, 1980-10-6, A. Baba s.n. (TNS-VS-439537); Nakanozawa,zaki-shi, Hirai, 1985-10-10, I. Ohzu s.n. (TNS-VS-1153097); Shi-

1980-10-7, A. Baba s.n. (TNS-VS-439536); Futaba-gun, Namie-

bata-shi, Sugatani, 1978-10-1, M. Saito 2379 (TNS-VS-543148),

machi, Kitasawa, alt. 25 m, 1975-8-21, Y. Yuzawa s.n. (TNS-

ibid., 1978-10-1, M. Saito 2385 (TNS-VS-543192), Nioji Shrine,

VS-438921); Ibaraki Pref.: Tsuchiura-shi, Shishitsuka, 1975-8-

1962-10-28, F. Konta 280 (TNS-VS-929631); Niitsu-shi, Akiba-

22, M. Yasu & N. Wada s.n. (TNS-VS-422804); Katsuta-shi,

yama, 1952-8-1, M. Saito 2062 (TNS-VS-543147); Joetsu-shi,

Kanaage, 1975-8-24, M. Yasu s.n. (TNS-VS-422812); Takahagi-

Atagokokubu, 1962-11-18, T. Kuramata s.n. (TNS-VS-543193);

shi, Takahagi, 1974-9-11, M. Yasu s.n. (TNS-VS-422575); Kitai-

Nakakanbara-gun, Muramatsu-machi, Takaishi, 1978-9-23, M.

baraki-shi, Hanakawa-cho, 1972-9-15, N. Wada 1164 (TNS-

Saito 2187 (TNS-VS-1172253); Nishikanbara-gun, Yahiko-mura,

VS-596378); Tsukuba-shi, Amakubo, Tsukuba Botanical Garden,

Yu Shrine, 1978-8-3, M. Saito 2085 (TNS-VS-543161); Minami-

2000-9-1, S. Matsumoto, H. Ebata & H. Uemura 12-13 (TNS-

kanbara-gun, Shitada-mura, Yagihana, 1979-8-27, M. Saito 3194

VS-9512072); Higashiibaraki-gun, Uchihara-machi, Shimizu-(TNS-VS-543151); Komyo-zan, 1979-8-28, M. Saito 3168 (TNS-

numa, 1975-8-20, M. Yasu s.n. (TNS-VS-422813); Nishiibaraki-

VS-543160); Higashikanbara-gun, Tsugawa-machi, 1978-9-23, M.

gun, Nanakai-mura, Keisoku-zan, 1977-9-10, M. Yasu s.n.

Saito 2233 (TNS-VS-543142); Mikawa-mura, Tsunagi, 1981-10-(TNS-VS-422579, 423407); Tsukuba-gun, Tsukuba-machi, Tsu-

29, M. Saito 3696 (TNS-VS-436143); Santo-gun, Koshiji-machi,kuba-san, 1970-9-6, M. Yasu s.n. (TNS-VS-423409); Sashima-gun,

Tsukayama Pass, 1985-10-21, I. Ohzu s.n. (TNS-VS-1153098,

Y.

Kimizuka

s.n.

(TNS-

1153099); Koshi-gun, Yamadori-mura, Hachibuse, 1909-10-3, M.

VS-596379); Tochigi Pref.: Utsunomiya-shi, Kogashi-yama,

Nakamura s.n. (TNS-VS-89889); Kariwa-gun, Oguni-machi,

1979-9-28, T. Sato s.n. (TNS-VS-1172245); Kanuma-shi, Kawa-

Hachioji, 1966-10-30, S. Iwano s.n. (TNS-VS-543171); Higashi-

bake-yama, 1977-9-10, T. Waku 7468 (TNS-VS-570761); Oyama-

kubiki-gun, Matsunoyama-machi, Usagiguchi, 1981-10-11, M.

shi, 1987-11-6, M. Ono s.n. (TNS-VS-1023725, 1023727); Kawa-

Saito 3704 (TNS-VS-436142); Iwafune-gun, Arakawa-machi,

chi-gun, Kaminokawa-machi, coll. date unknown, Y. Jotani 197

Hanadate, 1982-10-5, T. Nakaike s.n. (TNS-VS-433610); Asahi-(TNS-VS-831877), 1930-?-?, Y. Jotani 179 (TNS-VS-831878);

mura, Shinbo-dake, 1987-9-27, M. Saito s.n. (TNS-VS-1152603);

Minamikawachi-machi, Nishitsubo-yama, 1979-10-20, M. Usui

Sarusawa, 1987-9-27, M. Saito s.n. (TNS-VS-1152604); Nakano,Goka-mura,

Egawa,

1980-11-2,

s.n. (TNS-VS-422691); Kamikawachi-mura, Miyayamada, 1981-

1976-11-8, M. Saito 3344 (TNS-VS-543149); Sanpoku-machi,

10-2, S. Waku s.n. (TNS-VS-438799); Kawachi-machi, Kami-Odai, 1982-10-4, T. Nakaike s.n. (TNS-VS-433608, 433609);

tahara, alt. 160 m, 1987-9-13, H. Ogawa s.n. (TNS-VS-799735);

Toyama Pref.: Toyama-shi, Kureha-yama, 1979-7-17, T. Ohshima

Kamitsuga-gun, Nishikata-mura, Manago, 1981-9-7, M. Usui s.n.

s.n. (TNS-VS-543547); Uozu-shi, Toradani, 1977-8-3, T. Ohshima(TNS-VS-427401); Haga-gun, Mashiko-machi, Kitahara, alt.

s.n. (TNS-VS-543545); Himi-shi, Oura, 1977-8-29, T. Ohshima

120 m, 1987-8-2, Y. Sugita s.n. (TNS-VS-799731); Shimotsuga-

s.n. (TNS-VS-343999); Namerikawa-shi, Minowa, 1976-10-6, S.

gun, Kokubunji-machi, Koganei, 1979-10-20, M. Usui s.n. (TNS-

Matsumoto 761006-44 (TNS-VS-9506763); Hongo, 1976-10-6, S.

VS-1172277); Tomiyama-mura, Ohira-san, 1937-9-23, Y. Jotani

Matsumoto 761006-63 (TNS-VS-9506782); Gyoden Park, 1982-9-

15008 (TNS-VS-831879); Nasu-gun, Kurobane-machi, Kitano-

11, T. Ohshima s.n. (TNS-VS-1156184); Tonami-shi, Seridani,gami, 1981-9-10, M. Usui s.n. (TNS-VS-427397); Nasu-machi,

1976-8-21, T. Ohshima s.n. (TNS-VS-343915); Higashitonami-

Nasu Imperial Villa, 1966-9-5, H. Ito s.n. (TNS-VS-1045368);

gun, Inami-machi, Tojoji, 1982-10-10, T. Ohshima s.n. (TNS-

Kuroiso-machi, Kutsukake, 1969-9-1, T. Waku 1169 (TNS-

VS-1156182); Ishikawa Pref.: Suzu-shi, Hojuji Temple, 1969-9-8,

VS-422689); Nishinasuno-machi, Senbonmatsu, 1967-9-7, H. Ito

S. Komaki s.n. (TNS-VS-424537); Enuma-gun, Higashitanioku-

s.n. (TNS-VS-1045378, 1045381); Gunma Pref.: Gunma-gun,

mura, Taki - Aratani, 1953-9-27, S. Yoshimaru 1 (TNS-Haruna-machi, Haruna-san, 1968-7-19, Y. Jotani 1191 (TNS-

VS-1042109); Nomi-gun, Kaneno-mura, Gokoji, 1953-11-3, S.

VS-831880); Kanra-gun, Shimonita-machi, Kozu Farm, 1955-8-7,

Yoshimaru s.n. (TNS-VS-1042089); Ishikawa-gun, Tsurugi-machi,

T. Satomi s.n. (TNS-VS-1042087); Tone-gun, Minakami-machi,

Okushishiku, 1974-9-23, M. Honda s.n. (TNS-VS-424526); Fukui

S.

Ishiguro

s.n.

(TNS-

Pref.: Fukui-shi, Shigetate, 1984-6-29, Y. Yoshimura s.n. (TNS-

VS-1153084); Saitama Pref.: Iruma-gun, Naguri-mura, Hitomi,

VS-1152599); Tsuruga-shi, Iwagomori-yama, 1964-9-20, S. Wata-

1985-10-13, S. Ishiguro s.n. (TNS-VS-464003); Chichibu-gun,nabe s.n. (TNS-VS-1159835); Ono-shi, Tsukahara, 1977-9-15, Y.

Okununobiki-zawa,

1985-9-23,

Otaki-mura,Taiyoji,

1988-10-15,

T.

Iwata

9054

(TNS-

Saito s.n. (TNS-VS-1159796); Katsuyama-shi, Heisenji, 1977-10-

VS-1025581);

ibid.,

1988-10-15,

T.

Iwata

9055

(TNS-

23, N. Saito 7 (TNS-VS-415179); Oyatani, 1975-8-24, Y. Saito s.n.

VS-1025582); Higashichichibu-mura, Dodaira-san, 1983-9-25, T.

(TNS-VS-384610); Toritate-yama, 1967-8-9, S. Watanabe s.n.

Iwata 2112 (TNS-VS-1133786); Kodama-gun, Kamikawa-mura,(TNS-VS-1159798); Sakai-gun, Mikuni-cho, Jingaoka, alt. 30 m,

Niisato, 1980-10-12, S. Kojima s.n. (TNS-VS-514484); Chiba

1992-11-8, S. Watanabe 36725 (TNS-VS-1152602); Hamaji, 1978-

Pref.: Higashikatsushika-gun, Kamagaya-machi, 1961-8-6, H.

8-, Y. Saito s.n. (TNS-VS-1159833); Yamanashi Pref.: Nishiyatsu-

Ohba & M. Ohsawa s.n. (TNS-VS-147806); Tokyo Pref.: Kita-

shiro-gun, Kamikuishiki-mura, Aokigahara, alt. 1000 m, 1967-8-

Distribution of the Thelypteris japonica Complex (Thelypteridaceae)

79

17, F. Konta & H. Takahashi 643 (TNS-VS-929605); Shojiko,

Hirayu, 1973-8-10, H. Ishikawa 388 (TNS-VS-1045384); Shi-

1981-8-31, F. Miyamoto 11 (TNS-VS-831730a/b, 831732a/b);

zuoka Pref.: Fujinomiya-shi, Inokasira, 1981-10-10, M. Nakagawa

Nagano Pref.: Nagano-shi, Hikage, alt. 750 m, 1981-10-7, H.

s.n. (TNS-VS-1152333); Fukuroi-shi, Yamamoto, 1984-9-23, T.

Tsutsumi s.n. (TNS-VS-574420); Iida-shi, Tenryukyo, 1974-11-1,

SuginoK. Otsuka 2334 (TNS-VS-1157062); Matsukawa, alt. 1050 m,

Kogane-sawa, 1977-10-23, T. Sugino s.n. (TNS-VS-591599);

1980-8-3, H. Tsutsumi s.n. (TNS-VS-1159827); Komagane-shi,

Aichi Pref.: Okazaki-shi, Iwazu-cho, 1980-9-7, K. Inukai 3770

Kochi, alt. 650 m, 1981-10-13, H. Tsutsumi s.n. (TNS-(TNS-VS-432930); Nakashima-gun, Sobue-cho, Sobue, 1983-9-

VS-574418); Omachi-shi, Otari, alt. 680 m, 1981-9-8, H. Tsutsumi

26, K. Inukai 6338 (TNS-VS-1156981); Hazu-gun, Hazu-cho,s.n. (TNS-VS-574427); Kizaki-ko, alt. 800 m, 1981-9-7, H. Tsutsumi

Nishihazu, 1980-7-25, M. Muramatsu s.n. (TNS-VS-432970);

s.n. (TNS-VS-574439); Takagari-yama, 1981-8-30, T. Waku s.n.

Nukata-gun, Nukata-cho, Natsuyama, alt. 250 m, 1990-7-28, Y.

(TNS-VS-1157075); Iyari, 1980-8-21, F. Miyamoto 6710 (TNS-

Fukuoka 970 (TNS-VS-683758); Higashikamo-gun, Asuke-cho,VS-831767); ibid., 1980-8-21, F. Miyamoto 6709 (TNS-

Korankei, 1971-9-24, M. Fukuhara 3723 (TNS-VS-432916); Mie

Iwata-gun,Misakubo-cho,Pref.: Age-gun, Kawage-cho, Ueno, 1980-8-5, S. Yamauchi s.n.

(TNS-VS-831769); ibid., 1980-8-21, F. Miyamoto 6708 (TNS-(TNS-VS-437420); Minamimuro-gun, Kiho-cho, Tashiro, 2004-6-

ibid.,

1980-8-21,

Miyamoto(TNS-VS-458072);

6711

VS-831768);

F.

s.n.

VS-831770); ibid., 1951-9-8, T. Nakamura 15 (TNS-VS-831762);

15, S. Fujimoto s.n. (TNS-VS-764365); Shiga Pref.: Koka-gun,ibid., 1951-9-8, T. Nakamura 26 (TNS-VS-831761); ibid., 1951-9-

Konan-cho, Iwao-yama, 1981-9-2, S. Yamauchi s.n. (TNS-

8, T. Nakamura 27 (TNS-VS-831680); Koshoku-shi, Obasute, alt.

VS-533925); Sakata-gun, Santo-cho, Manganji, 1982-10-23, M.

750 m, 1981-10-7, H. Tsutsumi s.n. (TNS-VS-574419); Saku-shi,

Mase s.n. (TNS-VS-1152559); Ika-gun, Nishiazai-cho, Yamakado,

Yoseishi-yama, alt. 1250 m, 1981-9-22, H. Tsutsumi s.n. (TNS-

1977-10-18, T. Waku 7871 (TNS-VS-558889); Takashima-gun,

VS-574440); Minamisaku-gun, Yachiho-mura, Yachiho Lake, alt.

Imazu-cho, Sanamidera, 1979-11-3, T. Tatebe s.n. (TNS-

1050 m, 1981-9-21, H. Tsutsumi s.n. (TNS-VS-574435); Kitasaku-

VS-558891); Kyoto Pref.: Kyoto-shi Sakyo-ku, Daimonji-yama,

gun, Karuizawa-machi, Mine-no-chaya, alt. 1300 m, 1981-9-24, H.

1979-8-29, N. Yoshikawa 76560 (TNS-VS-594321); Soraku-gun,Tsutsumi s.n. (TNS-VS-574442); Yagasaki, 1980-8-19, T. Nakaike

Kamo-cho, Tono, 1977-7-17, Z. Tsujimoto s.n. (TNS-VS-594322);

s.n. (TNS-VS-1159817); Chiisagata-gun, Wada-mura, Wada Pass,

Wazuka-cho, Jubu-san, alt. 300–600 m, 1978-9-17, S. Tsugaru

1980-10-4, K. Otsuka 7368 (TNS-VS-431618); Shimoina-gun,

5128 (TNS-VS-489993); Amata-gun, Miwa-cho, Miharu Pass,

Seinaiji-mura, Kamiseinaiji, 1980-11-1, B. Sasaki s.n. (TNS-

1983-8-8, T. Nakaike s.n. (TNS-VS-1157024); Osaka Pref.: Shi-

VS-431608); Urugi-mura, Iwakura, alt. 800 m, 1981-10-12, H.

jonawate-shi, Muroike, 1982-8-16, A. Komaki s.n. (TNS-

Tsutsumi s.n. (TNS-VS-574444); Chausu-yama, 1980-10-10, K.

VS-1156220);

Otsuka 7440 (TNS-VS-1159822); Kiso-gun, Okuwa-mura, Shimo-

tenno-ryo, 1950-8-17, T. Koyama 2 (TNS-VS-1042106); Hyogo

zaigo, 1980-8-25, T. Nakaike s.n. (TNS-VS-1159814); Kitaazumi-

Pref.: Kasai-gun, Shimozato-mura, Hokke-san, 1952-10-27, T.

gun, Hokujo-mura, Sarukura, 1953-8-14, T. Nakamura s.n. (TNS-

Kodama 3644 (TNS-VS-129845); ibid., alt. 150 m, 1952-10-27, T.

VS-831742); Shirouma-dake, 1953-8-29, T. Nakamura 698

Kodama 3659 (TNS-VS-129844); Izushi-gun, Tanto-cho, Mushu,(TNS-VS-831741); Shimoina-gun, Oshima-mura, Sakurayama,

1981-8-19, Y. Hayashi s.n. (TNS-VS-563475); Nara Pref.: Nara-

1955-10-9, M. Muramatsu 1304 (TNS-VS-127296); Kitaazumi-

shi,gun, Miasa-mura, Minami-mura, alt. 840 m, 1981-9-6, H. Tsutsumi

VS-311232); Uda-gun, Oda-cho, 1950-10-15, T. Kodama 1241

s.n. (TNS-VS-574422); Hakuba-mura, Sanosaka, alt. 800 m, 1981-(TNS-VS-129839); Yoshino-gun, Higashiyoshino-mura, Myojin-

9-4, H. Tsutsumi s.n. (TNS-VS-574421); Happo, 1980-8-23, Z.

dake, 1982-8-22, Z. Tsujimoto s.n. (TNS-VS-1156219); Tottori

Kaneda s.n. (TNS-VS-431613); Kamiminochi-gun, Toyono-machi,

Pref.: Tottori-shi, Kyusho-zan, 1980-10-12, A. Tanaka s.n. (TNS-

Okura, alt. 500 m, 1981-9-25, H. Tsutsumi s.n. (TNS-VS-574443);

VS-433501); Bogaki, 1981-10-24, A. Tanaka s.n. (TNS-

Shimominochi-gun, Sakae-mura, Mori, alt. 300 m, 1981-10-1, H.

VS-433499); Yazu-gun, Wakasa-cho, Wakasa, 1976-8-17, A.

Tsutsumi s.n. (TNS-VS-574437); Gifu Pref.: Gifu-shi, Kinka-zan,

Tanaka 23146 (TNS-VS-424991); Hino-gun, Nichinan-cho,

1938-8-15, Y. Jotani s.n. (TNS-VS-831681); ibid., 1935-9-19, T.

Shoyama, 1981-9-22, M. Aoto s.n. (TNS-VS-433122); Okayama

Nomura s.n. (TNS-VS-1017515); Takayama-shi, Shiroyama Park,

Pref.: Okayama-shi, Yoshii, 1984-8-1, H. Kobatake s.n. (TNS-

1987-9-18, A. Yamamoto s.n. (TNS-VS-1157085); Mizunami-shi,

VS-1152625); Shinogose, 1980-10-22, H. Kobatake s.n. (TNS-

Sue, 1976-9-16, Y. Yamamoto s.n. (TNS-VS-1159803); Ena-shi,

VS-423262); Niimi-shi, Yatani, 1942-9-27, T. Watanabe s.n. (TNS-

Agi-gawa, 1981-10-11, Y. Yamamoto s.n. (TNS-VS-435839); Toki-

VS-423261); Mitsu-gun, Kamogawa-cho, Hongu-zan, 1981-10-18,

shi, Nakagiri - Ohira, 1985-7-29, T. Ogiyama s.n. (TNS-

T. Saito s.n. (TNS-VS-503836); Jobo-gun, Kayo-cho, Yoshikawa,

VS-1159851); Kakamigahara-shi, Nakanagatsuka, 1979-10-23, Y.

1980-10-27, H. Kobatake s.n. (TNS-VS-1172276); Hiroshima

Yamamoto s.n. (TNS-VS-1159809); Gujo-gun, Hachiman-cho,Pref.: Kure-shi, Haigamine, 1977-10-19, T. Takeda 10672 (TNS-

Nishibora - Kajika, 1978-9-19, Y. Yamamoto s.n. (TNS-

VS-545891); Takehara-shi, Tamari, 1977-10-16, T. Takeda 10619

VS-1157117); Shirotori-cho, Tochibora, 1980-9-21, Y. Yamamoto(TNS-VS-545905); Fukuyama-shi, Nanadani, 1975-10-14, T.

s.n. (TNS-VS-435838); Ono-gun, Takane-mura, Hiwada, 1988-9-

Takeda 4946 (TNS-VS-547864); Kaya, 1974-8-19, T. Takeda 2384

4, H. Miyazaki 7647 (TNS-VS-831751); ibid., 1988-9-4, H.

(TNS-VS-545893); Fuchu-shi, Kusa, 1976-11-9, T. Takeda 8587

Miyazaki 7648 (TNS-VS-831750); Yoshiki-gun, Kamitakara-mura,(TNS-VS-545904); Miyoshi-shi, Kamikawatachi, 1975-9-20, T.

Minamikawachi-gun,Kasuga-yama,

1951-11-4,

M.

Furuichi-machi,

Tagawa4217

Ankan-(TNS-

80

Atsushi Ebihara and Narumi Nakato
Takeda 4746 (TNS-VS-545889); Toyota-gun, Yasuura-cho, Ichi-(TNS-VS-439527); Chikatsu-mura, Yamamoto-fudo, 1975-8-26, Z.

bara, 1976-9-15, T. Takeda 8100 (TNS-VS-547847); Jinseki-gun,Kaneda s.n. (TNS-VS-1172256); Futaba-gun, Namie-machi, Seki-

Jinseki-cho, Taishakukyo, 1972-9-25, T. Takeda 293 (TNS-

mori, 1979-8-23, Y. Kanno s.n. (TNS-VS-439541); Ibaraki Pref.:VS-545888); Sanwa-cho, Age, 1975-9-15, T. Takeda 4684 (TNS-

Hitachi-shi, Takasuzu-yama, 1970-8-28, M. Yasu s.n. (TNS-

10938

VS-422578); Hitachiota-shi, Takasuzu-yama, 1969-9-15, M. Yasu

(TNS-VS-545903); Futami-gun, Kisa-cho, Okada-yama, 1973-8-

s.n. (TNS-VS-423404); Takahagi-shi, Hananuki Valley, 1973-8-17,

31, T. Takeda 286 (TNS-VS-1152799); Hiba-gun, Kuchiwa-cho,

M. Yasu s.n. (TNS-VS-423405); Kitaibaraki-shi, Hanazono-yama,

1975-8-24, T. Takeda 4361 (TNS-VS-545902).

1971-9-15, M. Yasu s.n. (TNS-VS-423399); Higashiibaraki-gun,

VS-545897);

Tokiyasu,

1977-11-11,

T.

TakedaKyushu - Saga Pref.: Taku-shi, Kitataku-machi, 1964-8-?, Y. Inoue

Katsura-mura, Gozen-yama, 1971-9-12, M. Yasu s.n. (TNS-s.n. (TNS-VS-579337); Kumamoto Pref.: Kamimashiki-gun,

VS-423401); Kuji-gun, Satomi-mura, Yokokawa, 1971-9-19, M.

Yabe-machi, Yamaguchi, 1978-8-27, N. Beppu s.n. (TNS-

Yasu s.n. (TNS-VS-423403); Kozuma Farm, 1971-9-24, M. Yasu

VS-550076); Oita Pref.: Beppu-shi, Myoban Spa, 1977-9-18, Y.

s.n. (TNS-VS-423402); Niihari-gun, Sakura-mura, Tennodai,

Takaoka

1978-10-12, M. Michikawa 10010 (TNS-VS-1080164); Makabe-s.n.

(TNS-VS-1159829);

Naoiri-gun,Kuju-machi,

Akagawa, 1963-8-18, Y. Takaoka s.n. (TNS-VS-420030).

gun, Makabe-machi, Kaba-san, 1975-9-28, M. Yasu s.n. (TNSVS-423400); Tochigi Pref.: Kanuma-shi, Futamata-yama, 1980-9-

Thelypteris musashiensis (Hiyama) Nakato, Sahashi et M.
Kato [Iwa-harigane-warabi]
Hokkaido - Hokkaido Pref.: Hiroo-gun, Hiroo-cho, Rakko-dale,
1984-8-11, Y. Goda s.n. (TNS-VS-1152564); Usu-gun, Otakimura, Kitayuzawa, 1972-9-15, M. Hara s.n. (TNS-VS-545560).
Honshu - Aomori Pref.: Mutsu-shi, Arakawa-dake, alt. 750 m, 19809-15, M. Neichi s.n. (TNS-VS-570348); Iwate Pref.: Morioka-shi,
Izari-sawa, 1979-9-24, Y. Nakajima s.n. (TNS-VS-440270); Iwate
Park, 1980-8-14, E. Fukushi s.n. (TNS-VS-935895); Hanamaki-

1, M. Usui s.n. (TNS-VS-570767); Ozaku-san, 1979-9-24, H.
Kobayashi s.n. (TNS-VS-570765); Kawabake-yama, coll. dateunknown, F. Maekawa 80216 (TNS-VS-831864); Ozaku-san,
1981-10-12, M. Usui s.n. (TNS-VS-427402); Nikko-shi, Takino-o,
1980-10-22, M. Usui s.n. (TNS-VS-427399); Hoso-o, alt. 1250 m,
1987-8-7, S. Nemoto s.n. (TNS-VS-799764); Uwano, alt. 520 m,
1987-8-9, T. Kamiyama s.n. (TNS-VS-799767); Imaichi-shi,
Tokawa River, alt. 500 m, 1987-8-6, T. Kamiyama s.n. (TNSVS-799769); Kuroiso-shi, Momura-yama, 1967-9-30, T. Waku 336

shi, Shitodaira, 1966-7-9, K. Inoue s.n. (TNS-VS-440278); Iwate-(TNS-VS-570764); ibid., 1987-9-13, S. Ishiguro s.n. (TNS-

gun, Shizukuishi-cho, Taigakura, 1977-9-16, S. Tamakawa s.n.

VS-1152588); Kamitsuga-gun, Awano-machi, Ode-yama, 1957-9-(TNS-VS-440264); Takinoue Spa, 1981-8-21, K. Inoue s.n. (TNSVS-1156213); Taigakura, 1977-9-15, T. Waku 7513 (TNSVS-935894); Miyagi Pref.: Natori-gun, Akiu-machi, Okunikkawa,
1980-9-20, N. Sagawa s.n. (TNS-VS-575764); Sakaino, 1981-820, K. Shoji s.n. (TNS-VS-576840); Miyagi-gun, Miyagi-machi,
Okunikkawa, 1976-10-11, W. Takahashi s.n. (TNS-VS-1172231);
Kurokawa-gun,Taiwa-cho,Ochiai-aikawa,

1976-10-2,

W.

Takahashi s.n. (TNS-VS-1172223); Akita Pref.: Kitaakita-gun,
Kamikoani-mura, Haginari, 1976-10-15, J. Takada s.n. (TNSVS-419808); Senboku-gun, Tazawako-machi, Tazawako-kogen,
1973-8-11, J. Takada s.n. (TNS-VS-419806), 1976-9-10, T.
Nakaike s.n. (TNS-VS-419785); Yamagata Pref.: Mogami-gun,
Sakegawa-mura, Dakigaeri Valley, 1981-9-15, N. Sagawa s.n.
(TNS-VS-437978); Tozawa-mura, Shinjo Spa, 1980-9-13, S. Katos.n. (TNS-VS-437625); Nishiokitama-gun, Oguni-machi, Nukumidaira, 1979-9-24, M. Saito 3 (TNS-VS-543152); Chojahara Nukumidaira, 1969-8-27, S. Serizawa 10813 (TNS-VS-1042037);
Higashitagawa-gun, Haguro-machi, Toge, 1980-10-1, T. Nakaikes.n. (TNS-VS-437953); Fukushima Pref.: Iwaki-shi, Obamamachi, 1984-11-14, M. Origasa s.n. (TNS-VS-1152593); Ohisamachi, 1984-11-12, K. Nishimaki s.n. (TNS-VS-1152596); Tabitomachi, alt. 400 m, 1972-9-17, Y. Yuzawa s.n. (TNS-VS-439160);

8, K. Ogawa s.n. (TNS-VS-1042094); Kaminagano, 1978-8-27, T.
Waku 8993 (TNS-VS-1172226); Iriawano, 1957-9-8, K. Ogawa
s.n. (TNS-VS-1172225); Ashio-machi, Mochigase, 1957-8-19, K.
Ogawa s.n. (TNS-VS-570756); Haga-gun, Motegi-machi, Yakimori-yama, 1982-9-15, T. Tashiro s.n. (TNS-VS-1152580);
Shioya-gun, Shiobara-machi, Fuji-san, 1977-8-15, T. Waku 7285
(TNS-VS-570760); Kuriyama-mura, Nokado, 1982-9-5, T. Tashiro
s.n. (TNS-VS-1157012); Fujihara-machi, Ryuo-kyo, 1984-9-15,
M. Usui s.n. (TNS-VS-1152565); Kosagoe, 1981-9-29, M. Usuis.n. (TNS-VS-427398); Nasu-gun, Bato-machi, Torinoko-san,
1982-8-24, H. Kato s.n. (TNS-VS-1152590); Kurobane-machi,
Unganji Temple, 1967-9-6, H. Ito s.n. (TNS-VS-1045685); Nasumachi, Kita Spa, 1965-8-29, H. Ito s.n. (TNS-VS-1045367);
Nigato-gawa, alt. 620 m, 1985-8-27, H. Ogura s.n. (TNSVS-799770); Ashino, 1968-8-22, H. Ito s.n. (TNS-VS-1042034);
Aso-gun, Tanuma-machi, Oto, 1981-9-23, T. Waku s.n. (TNSVS-431951); Horai-san, 1963-10-6, K. Ogawa s.n. (TNSVS-1172228); Gunma Pref.: Kiryu-shi, Nemoto-yama, 1973-113, S. Suto s.n. (TNS-VS-435558); Seta-gun, Miyagi-mura, Chuji
Spa, 1971-10-11, S. Suto s.n. (TNS-VS-435559); Azuma-mura,
Sori - Otaki, 1987-9-24, S. Waku s.n. (TNS-VS-1153090); Gunmagun, Haruna-machi, 1977-9-8, H. Suzuki s.n. (TNS-VS-435556);

Adachi-gun, Otama-mura, Endo-taki, 1977-8-23, Z. Kaneda s.n.

Kurabuchi-mura, Tsukinami, 1987-8-20, Y. Terao s.n. (TNS-(TNS-VS-439535); Yama-gun,VS-1156194); Tano-gun, Nakasato-mura, Tofukuji-gawa, 1984-8-

Nishiaizu-machi,

Oyamazumi

Shrine, 1980-9-23, Z. Kaneda 6099 (TNS-VS-1172255); Nishishirakawa-gun, Nishigo-mura, Kasshi-yama, 1984-8-21, A. Yamamoto s.n. (TNS-VS-1152566); Higashishirakawa-gun, Yamatsurimachi,

Yamatsuri-yama,

1980-10-6,

A.

Baba

s.n.

29, T. Iwata 3372 (TNS-VS-1025521); Ueno-mura, Suwa-yama,
1985-10-20, T. Iwata 5741 (TNS-VS-1025496); Tenmaru-yama,
1984-10-10, T. Iwata 3595 (TNS-VS-1025517); ibid., 1984-10-10,
T. Iwata 3613 (TNS-VS-1025518); Hamadaira, 1965-8-11, T.

Distribution of the Thelypteris japonica Complex (Thelypteridaceae)

81

Satomi s.n. (TNS-VS-1045369); Kanra-gun, Myogi-machi, Myogi-(TNS-VS-1025561); ibid., 1993-9-11, T. Iwata 11236 (TNS-

san, 1973-5-19, S. Suto s.n. (TNS-VS-435561); Shimonita-machi,

VS-1025562); Hakuta-san, 1994-7-24, T. Iwata 37908 (TNS-

1986-10-18, M. Ono s.n. (TNS-VS-831860); Usui-gun, Matsuida-

VS-1025554); Matanosawa, 1994-7-25, T. Iwata 37987 (TNS-

machi, Myogi-san, 1951-10-13, N. Adachi s.n. (TNS-VS-92803);

VS-1025553);

Agatsuma-gun, Nakanojo-machi, Yamada, 1985-9-19, S. Waku s.n.

(TNS-VS-1025552); Owakazawa, 1995-7-13, T. Iwata 41484(TNS-VS-1159877); Agatsuma-machi, Agatsuma Valley, 1972-10-(TNS-VS-1025480); Nanten-yama, 1995-8-7, T. Iwata 41633

10, S. Suto s.n. (TNS-VS-435560); Tone-gun, Kawaba-mura, Mae-

(TNS-VS-1025479); Akaiwa Pass, 2002-7-9, T. Iwata 65777

hotaka, 1975-8-21, S. Kurata s.n. (TNS-VS-333438); Akakura,(TNS-VS-1025468); Ochigawa, 1981-8-26, S. Kobayashi s.n.

1985-9-6, S. Waku s.n. (TNS-VS-1153086); Minakami-machi,(TNS-VS-514477); Ochigawa-nishitani, 1996-7-27, T. Iwata

Yubiso-gawa, 1980-10-22, T. Waku 10367 (TNS-VS-435562);

44816 (TNS-VS-1025477); ibid., 2002-7-26, T. Iwata 66079

Okura-zawa, 1981-10-18, T. Waku s.n. (TNS-VS-435557);(TNS-VS-1025466); Ochigawa-higashitani, 1948-8-17, T. Nakai

Saitama Pref.: Chichibu-shi, Urayama, Baramo-sawa, 1993-10-

s.n. (TNS-VS-78194, 79320); Myohogatake - Taiyoji Temple,

31, T. Iwata 11430 (TNS-VS-1025558); Shiratono-sawa, 1994-8-

1988-10-15, T. Iwata 9084 (TNS-VS-1025583); ibid., 1988-10-15,

11, T. Iwata 38863 (TNS-VS-1025551); Sobatsubu-yama, 1987-9-

T. Iwata 9085 (TNS-VS-1025584); Mameyaki-sawa, 1998-8-15, T.

1, T. Iwata 8074 (TNS-VS-1025590); Hirogawara-dani, 1987-9-1,

Iwata 55199 (TNS-VS-1025472); ibid., 1981-8-26, S. Kobayashi

T. Iwata 8058 (TNS-VS-1025591); ibid., 1987-9-1, T. Iwata 8076

s.n. (TNS-VS-514480); Hirogawara, 1981-8-26, S. Kobayashi s.n.

(TNS-VS-1025592); Urayama, 1981-9-17, T. Iwata s.n. (TNS-(TNS-VS-425191); Urayama-mura, 1952-8-, H. Ito s.n. (TNS-

Irigawa,

1994-7-26,

T.

Iwata

38297

VS-1025537); Okubo-dani, 1973-9-30, T. Iwata s.n. (TNS-

VS-1042091); Baramo-sawa, 1953-8-9, S. Kurata 1455 (TNS-

VS-1025541); Ohira-yama, 1969-8-21, T. Iwata 290 (TNS-

VS-831739); Arakawa-mura, Anya-gawa, 1993-9-5, T. Iwata

VS-1025544); Hanno-shi, Tenran, 2001-9-19, T. Iwata 64020

11287 (TNS-VS-1025563); Kumakura-san, 1994-6-5, T. Iwata(TNS-VS-1025469); Higshiagano - Ogami-yama, 1975-9-28, T.

36843 (TNS-VS-1025556); Shibahara, 1991-11-4, T. Iwata 10724

Nakaike s.n. (TNS-VS-332968); Agano - Koburi Pass - Kasasugi(TNS-VS-1025569); ibid., 1991-11-4, T. Iwata 10725 (TNS-

Pass - Kuroyama-santaki Falls, 2001-9-30, A. Yamamoto s.n.

VS-1025570); Saginosu, 1991-7-31, T. Iwata 10379 (TNS-(TNS-VS-9512773); Karibasaka Pass, 1984-11-17, T. Iwata 3931

VS-1025571); Shian-sawa, 1989-8-4, T. Iwata 9316 (TNS-(TNS-VS-1025512); ibid., 1984-11-17, T. Iwata 3930 (TNS-

VS-1025577); ibid., 1989-8-4, T. Iwata s.n. (TNS-VS-1025578);

VS-1025513); Kusumi, 1981-11-1, S. Kobayashi s.n. (TNS-

ibid., 1987-9-20, T. Iwata 8147 (TNS-VS-1025589); Kawauradani,

VS-425190); Iruma-gun, Ogose-machi, Otakatori-yama, 1997-5-

1987-8-9, T. Iwata 7896 (TNS-VS-1025595); ibid., 1987-8-9, T.

23, T. Iwata 49601 (TNS-VS-1025476); Kasasugi Pass, 1984-8-21,

Iwata 7897 (TNS-VS-1025596); Kumakura-san, 1985-9-8, T.

T. Iwata 3282 (TNS-VS-1025522); ibid., 1984-8-21, T. Iwata 3283

Iwata 5429 (TNS-VS-1025500); ibid., 1966-10-23, T. Iwata 300(TNS-VS-1025523); Naguri-mura, Yamanaka, 1989-9-1, T. Iwata(TNS-VS-1025545); ibid., 1971-8-22, T. Iwata s.n. (TNS-9484 (TNS-VS-1025574); ibid., 1989-9-1, T. Iwata 9418 (TNS-

VS-1025542); Higashichichibu-mura, Kasayama, 1998-6-10, T.

VS-1025575); ibid., 1989-9-1, T. Iwata 9424 (TNS-VS-1025576);

Iwata 53493 (TNS-VS-1025470); ibid., 1998-6-10, T. Iwata 53494

Agano-mura, Izugatake, 1935-10-20, S. Okuyama s.n. (TNS-(TNS-VS-1025471); Kodama-gun, Kamiizumi-mura, Jomine-san,

1935-7-7,

S.

Okuyama

VS-1159852);

ibid.,

VS-1152711);

Chichibu-gun, Yokoze-machi,

s.n.

(TNS-

1975-8-17, S. Kojima s.n. (TNS-VS-514483); Osato-gun, Yorii-

Komochi-yama,machi, Kamabuse, 1985-11-1, T. Iwata s.n. (TNS-VS-1133748);

1993-8-7, T. Iwata 11254 (TNS-VS-1025564); Omochi-sawa,Tokyo

1994-9-18, T. Iwata 39920 (TNS-VS-1025550); ibid., 1994-9-18,

Kobayashi s.n. (TNS-VS-423911); Kitatama-gun, Jindai-mura,T. Iwata 39921 (TNS-VS-1025549), 1987-8-23, T. Iwata 7981

Jindaiji, 1949-11-19, Y. Hiratsuka s.n. (TNS-VS-513331); Nishi-

T.

Iwata

7965

Pref.:

Hachioji-shi,

Kamikawa-cho,

1977-5-8,

H.

(TNS-

tama-gun, Hinohara-mura, Mito-taki, 1985-10-21, T. Nakaike s.n.

VS-1025594); Nagatoro-machi, Ido, 1994-5-16, T. Iwata 36344(TNS-VS-570051, 570052, 570056); Okutama-machi, Otaba-(TNS-VS-1025557); ibid., 1994-5-16, T. Iwata 36346 (TNS-

gawa, 1981-8-12, H. Endo s.n. (TNS-VS-570978); Kawanori-tani,

VS-1025547); Yoshida-machi, Onagata, 1985-10-6, T. Iwata 5659

1984-10-23, T. Iwata 3813 (TNS-VS-1025514); ibid., 1984-10-23,(TNS-VS-1025498); Ogano-machi, Haccho Pass, 1993-10-3, T.

T. Iwata 3794 (TNS-VS-1025515); Mitake-san, 1971-8-29, T.

Iwata 11343 (TNS-VS-1025559); Nunosawa, 1994-7-5, T. Iwata

Iwasaki 2690 (TNS-VS-831763); ibid., 1971-8-29, T. Iwasaki

37463 (TNS-VS-1025555); Shiraishi-yama, 1999-7-26, T. Iwata

2691 (TNS-VS-831764); Okumotori-dani, 1980-9-23, H. Endo s.n.

58720 (TNS-VS-1025474); Onouchi-sawa, 2002-8-15, T. Iwata(TNS-VS-423917); Kori, 1969-8-31, A. Yamamoto s.n. (TNS-

66240 (TNS-VS-1025465); Kurio, 1981-8-26, S. Kobayashi s.n.

VS-423912); Toke, 1980-10-10, H. Endo s.n. (TNS-VS-423913);(TNS-VS-514476); Ryokami-mura, Ryokami-san, 1993-7-25, T.

Mito-san, 1981-9-15, H. Endo s.n. (TNS-VS-435241); Mito-san,Iwata 11222 (TNS-VS-1025565); Shiraizasu Pass, 1995-9-8, T.

1980-10-11, T. Yamada 114 (TNS-VS-737859); Mito-san, 1980-

Iwata 41872 (TNS-VS-1025478); Hinata, 1997-8-30, T. Iwata

10-11, T. Yamada 114 (TNS-VS-738506); Kumotori-yama, 1978-(TNS-VS-1025593),

1987-8-23,

51521 (TNS-VS-1025475); Otaki-mura, Wanakura-yama, 1992-9-

8-28, T. Yamada s.n. (TNS-VS-831899); ibid., 1978-10-1, T.

14, T. Iwata 11038 (TNS-VS-1025567); Mameyaki-sawa, 1993-6-

Yoshizawa s.n. (TNS-VS-831898); ibid., 1978-9-30, A. Koyama

6, T. Iwata 11106 (TNS-VS-1025566); Tochimoto, 1993-9-11, T.

s.n. (TNS-VS-831897); Unazawa, 1977-9-25, K. Inoue s.n. (TNS-

Iwata 11233 (TNS-VS-1025560); ibid., 1993-9-11, T. Iwata 11234

VS-831896); Hikawa-machi, Bo-no-ore-yama, 1948-7-27, K.

82

Atsushi Ebihara and Narumi Nakato
Hiyama s.n. (TNS-VS-110131, holotype); Mikurajima-mura,s.n. (TNS-VS-1152605); Kamiina-gun, Tatsuno-machi, Yokokawa

Oyama, 1981-7-13, H. Hatta & S. Matsumoto 15445 (TNS-

Valley, alt. 1000 m, 1981-9-1, H. Tsutsumi s.n. (TNS-VS-574436);

VS-9508268, 9508334); ibid., 1934-7-29, Y. Jotani 10081 (TNS-

ibid., 1977-9-29, M. Suzuki & K. Tamaki s.n. (TNS-VS-1159823);

VS-831701), 1967-7-28, Y. Jotani & al. s.n. (TNS-VS-831704);

ibid., 1977-8-7, T. Waku 7208 (TNS-VS-431615); Iijima-machi,

Niigata Pref.: Shibata-shi, Kamagasawa, 1963-11-10, F. Konta

Yotagiri-gawa Valley, alt. 1350 m, 1981-8-11, H. Tsutsumi s.n.

3541 (TNS-VS-929612); Kitakanbara-gun, Kurokawa-mura, Tai-

(TNS-VS-574434); ibid., 1981-9-20, K. Otsuka 8942 (TNS-nai, 1981-11-3, M. Saito 3 (TNS-VS-436144); ibid., 1981-11-15,

VS-1159826); Shimoina-gun, Chiyo-mura, Mango-gawa, 1934-9-

M. Saito 4033 (TNS-VS-426750); Minamikanbara-gun, Shitada-16, H. Koidzumi 94863 (TNS-VS-909816); Igara-mura, Kuro-mura,

Kasabori,

1967-10-1,

T.

Takahashi

3038

(TNS-

kawa,

1952-9-13,

M.

Muramatsu

307(TNS-VS-114131);

VS-1172240); Kitauonuma-gun, Irihirose-mura, Shiraita Mine,

Kiso-gun, Agematsu-machi, Akazawa, 1980-8-25, T. Nakaike s.n.

1909-8-16, M. Nakamura s.n. (TNS-VS-89877); Iwafune-gun,(TNS-VS-431616); Kiso-mura, Miso-gawa, 1983-9-15, H. Okuhara

Asahi-mura, Suzugataki, 1961-10-16, M. Saito s.n. (TNS-s.n. (TNS-VS-1152568); Kaida-mura, Nagamine Pass, 1978-8-27,

VS-1152573, 1152575); Sarusawa, 1987-9-27, M. Saito s.n. (TNS-

H. Miyazaki 4843 (TNS-VS-831716); Minamiazumi-gun, Hotaka-VS-1153092); Sanpoku-machi, Imagawa, 1987-9-27, M. Saito s.n.

machi, Nakafusa Spa, alt. 1450 m, 1981-9-10, H. Tsutsumi s.n.

(TNS-VS-1153091); Toyama Pref.: Namerikawa-shi, Tofukujino,(TNS-VS-574417); Shinanosaka - Kannon Pass, alt. 1100 m, 1981-

S.

Matsumoto

760810-42(TNS-VS-9506903);

9-10, H. Tsutsumi s.n. (TNS-VS-574416); Azumi-mura, Shi-

Nakaniikawa-gun, Tateyama-machi, Kurobe, 1953-10-13, H. Itomashima - Tokugo Pass, 1976-10-11, F. Miyamoto 659 (TNS-s.n. (TNS-VS-1045689); Shomyo, Hachirozaka, 1983-10-9, T.

VS-831747);

Ohshima s.n. (TNS-VS-1152620); Kurobe Valley, 1953-10-13, H.

1973-7-29, S. Kurata, T. Nakaike & M. Sakakibara 55 (TNS-

Ito s.n. (TNS-VS-1042102, 1042103, 1042107); Shimoniikawa-VS-317322); Ariake-mura, 1930-8-4, N. Hiratsuka 11 (TNS-

gun, Asahi-machi, Kogawa Spa, 1976-10-7, T. Nakaike s.n. (TNS-

VS-513497), 1930-8-4, N. Hiratsuka 113 (TNS-VS-513160); Gifu

VS-510281, 543539); Imizu-gun, Kosugi-machi, Hirano, 1976-7-

Pref.: Ibi-gun, Sakauchi-mura, Ikenomata-dani, 1986-9-21, H.

1976-8-10,-

Iwanadome,alt.

750–900 m,

Pref.:

Miyazaki 7220 (TNS-VS-831756); ibid., 1986-9-21, H. Miyazaki

Ishikawa-gun, Oguchi-mura, Aratani, 1978-10-31, K. Yoneyama

7221 (TNS-VS-831755); ibid., 1986-9-21, H. Miyazaki 7219

s.n. (TNS-VS-424485); Fukui Pref.: Tsuruga-shi, Daguchi, 1978-(TNS-VS-831754); Tokuyama-mura, Kanmuri-yama, 1978-10-22,

8-18, S. Watanabe s.n. (TNS-VS-1159842); Takefu-shi, Hiodani-

Y. Yamamoto s.n. (TNS-VS-1157129); Ojirami-tani, 1984-8-14, T.

cho, 1966-8-20, S. Watanabe s.n. (TNS-VS-1159840); Obama-shi,Ogiyama s.n. (TNS-VS-1157058); Motosu-gun, Neo-mura, Myo-

Hyakuri-dake, 1971-8-24, S. Watanabe s.n. (TNS-VS-1159844);

jin-yama, 1981-10-4, Y. Yamamoto s.n. (TNS-VS-435840); Gujo-

Ono-shi, Arashima-dake, 1979-8-31, S. Watanabe s.n. (TNS-

gun, Hachiman-cho, Kibishima-gawa, 1979-10-10, Y. Yamamoto

VS-1159849); Katsuyama-shi, Hachibuse-yama, 1968-8-2, S.

s.n. (TNS-VS-1159806); Takasu-mura, Dainichi-dake, 1969-8-19,

Watanabe s.n. (TNS-VS-1159837); Ikegahara, 1966-7-29, S.

H. Kaneko 9036 (TNS-VS-254377); Minami-mura, Kappara,

Watanabe s.n. (TNS-VS-1159834); Sakai-gun, Awara-cho, Hama-

1977-12-8, Y. Yamamoto s.n. (TNS-VS-1159807); Ena-gun,saka, 1979-9-30, S. Watanabe s.n. (TNS-VS-1159845); Kanazu-

Kashimo-mura, Akaishi, 1978-11-7, Y. Yamamoto s.n. (TNS-cho, Yubinaka, 1976-10-17, S. Watanabe s.n. (TNS-VS-1159846);

VS-1159808); Higashihondani, 1979-11-29, Y. Yamamoto s.n.

Imadate-gun, Ikeda-cho, Kanmuri-yama, 1979-8-4, S. Watanabe(TNS-VS-1159802); Mashita-gun, Osaka-cho, Ochiai, Karadani,s.n. (TNS-VS-1159838); Heko-san, 1968-7-25, S. Watanabe s.n.

1981-9-19, A. Yamamoto s.n. (TNS-VS-1159813, 1172254); Gero-(TNS-VS-1159836); Mikata-gun, Mihama-cho, Asagase, 1979-10-cho, Hiuchi, 1980-8-17, Y. Yamamoto s.n. (TNS-VS-435841);

9, S. Watanabe s.n. (TNS-VS-1159848); Yamanashi Pref.: Kofu-

Yoshiki-gun, Kokufu-cho, Utsue, 1981-9-18, A. Yamamoto s.n.

shi, Kurobera, 1974-9-9, S. Kurata s.n. (TNS-VS-1152798);(TNS-VS-1159811); Shizuoka Pref.: Shizuoka-shi, Abe Pass,

1,

T.

Ohshimas.n.

(TNS-VS-543546);

Ishikawa

Shimashima

Otsuki-shi, Ubako-yama, 1979-8-9, T. Kobayashi s.n. (TNS-

1981-9-20, H. Endo s.n. (TNS-VS-591596); Numazu-shi, Ashi-

VS-573505); Higashiyamanashi-gun, Mitomi-mura, Nishizawa

taka-yama, 1983-7-28, T. Hosokura s.n. (TNS-VS-1152626); Fuji-

Valley, 1978-8-5, Y. Kimizuka s.n. (TNS-VS-1152332); Kitakoma-

nomiya-shi, Tenshigatake, alt. 1250 m, 1984-10-10, T. Sato 6534

gun, Oizumi-mura, Kawamatagawa-higashizawa, 1987-8-29, H.

(TNS-VS-929221); ibid., alt. 1200 m, 1983-8-28, T. Sato 4091

Kobayashi s.n. (TNS-VS-1153093); ibid., 1987-8-29, T. Nakaike(TNS-VS-1152616); Tagata-gun, Nakaizu-cho, Ikadaba, 1961-7-

s.n. (TNS-VS-535708); Mukawa-mura, Ihsiutoro-gawa, 1980-10-

24, E. Sakuma s.n. (TNS-VS-332285); Kamiohmi-mura, Amagi-5, M. Nakagawa s.n. (TNS-VS-1152328); Minamitsuru-gun,san, 1939-9-2, Y. Jotani s.n. (TNS-VS-831691); Haibara-gun,

Nishikatsura-mura, Mitsutoge-yama, 1935-?-?, T. Nomura s.n.

Nakakawane-cho, Jina, coll. date unknown, K. Tanaka s.n. (TNS-(TNS-VS-1018078); Kitatsuru-gun, Tabayama-mura, Taba-gawa,VS-1152592); Sobatsubu-yama, 1978-10-22, T. Sugino s.n. (TNS-

1981-8-6, H. Endo s.n. (TNS-VS-1152329, 1152330); Nagano

VS-591597); Honkawane-cho, Senzu - Sumata, alt. 500 m, 1969-7-

Pref.: Saku-shi, Kabutoiwa-yama, 1980-9-28, K. Nakayama 35

21, Y. Yokoe s.n. (TNS-VS-929593); Aichi Pref.: Toyota-shi,(TNS-VS-401537); Minamisaku-gun, Kawakami-mura, Jumonji

Sanage-yama,

Pass, 1975-8-16, K. Otsuka 3672 (TNS-VS-431614); Azusa-yama

VS-432915); Kitashitara-gun, Toyone-mura, Chausu-yama, 1981-- Jumonji Pass, 1972-8-11, H. Komaki s.n. (TNS-VS-739074);

8-25, K. Inukai 4853 (TNS-VS-545668); ibid., 1986-8-30, H.

Chiisagata-gun, Wada-mura, Kamiwada, 1985-8-2, T. Hosokura

Miyazaki 7076 (TNS-VS-831753); ibid., 1986-8-30, H. Miyazaki

1980-9-7,

M.

Muramatsu

80-465(TNS-Distribution of the Thelypteris japonica Complex (Thelypteridaceae)

837079 (TNS-VS-831749); ibid., 1986-8-30, H. Miyazaki 7075

(TNS-VS-423236); Katsuta-gun, Nagi-cho, Takiyama, 1979-8-18,(TNS-VS-831752); ibid., 1986-8-30, H. Miyazaki 7077 (TNS-

K. Takayama s.n. (TNS-VS-423235); Aida-gun, Nishiawakura-son,VS-831748); Mie Pref.: Yokkaichi-shi, Nishisakabe-cho, Oike,

Wakasugi Pass, 1977-9-25, T. Saito s.n. (TNS-VS-1172230);

1983-10-29, K. Kada 10552 (TNS-VS-454284); Shiga Pref.:Hiroshima Pref.: Saeki-gun, Saeki-cho, Iiyama, 1971-10-10, T.

Otsu-shi, Hira-san, 1966-10-16, H. Ito s.n. (TNS-VS-1045371,

Tamaru 392 (TNS-VS-545899); Yoshiwa-mura, Nakatsuya, 1978-

1045373); Shiga-gun, Shiga-cho, Uchimi-yama, 1975-8-20, T.

8-29, T. Tamaru 4363 (TNS-VS-545898).

Yahara 4054 (TNS-VS-364353); Koka-gun, Tsuchiyama-cho,

Shikoku - Tokushima Pref.: Myozai-gun, Kamiyama-cho, 1948-8-

Inaga-tani, 1981-8-17, T. Shiraiwa 10283 (TNS-VS-1159830);

15, T. Inobe 48 (TNS-VS-1042090); Ehime Pref.: Niihama-shi,

Kanzaki-gun, Eigenji-cho, Echi-gawa, 1979-9-24, S. Yamauchi s.n.

Sasagamine, 1963-8-15, H. Ishikawa 384 (TNS-VS-1045383);(TNS-VS-558883); Echi-gun, Koto-cho, Gion, Oshitate-yama,Ochi-gun, Tamagawa-cho, Kiji, Shiratsue, 1981-10-29, H.

1980-9-15, T. Tatebe s.n. (TNS-VS-558892); Kyoto Pref.: Kitaku-

Ishikawa s.n. (TNS-VS-1152326); Kamiukena-gun, Kuma-cho,wada-gun, Miyama-cho, Ashu Experimental Forest of Kyoto Uni-

Saragamine, 1980-10-25, H. Ishikawa s.n. (TNS-VS-1152327);

versity, 1987-10-1, S. Masuda s.n. (TNS-VS-831724); Kuroda-

Omogo-mura, Ishizuchi-san, 1965-9-29, S. Masuda 4617 (TNS-mura,

UodaniPass,

1952-9-23,

T.

Kodama2041

VS-831740); Yanadani-mura, Kamikiji, 1990-10-14, M. Hyodo s.n.

(TNS-VS-129837); Haiya, 1951-9-23, T. Kodama s.n. (TNS-(TNS-VS-1152621);

VS-247418); Osaka Pref.: Toyono-gun, Minoo-mura, Minoo,

1982-10-10, M. Hyodo s.n. (TNS-VS-1152558); Kochi Pref.:1913-9-?, U. Faurie 307 (TNS-VS-194774); Minamikawachi-gun,Nagaoka-gun,

Nagano-cho, Iwawaki-san, 1950-7-27, M. Tagawa 3437 (TNS-

unknown, H. Inoue 148 (TNS-VS-1042096).

Kitauwa-gun,

Nishitoyonaga-mura,Hiyoshi-mura,
Kajigamori,Jizo-yama,
coll.

date

VS-311233); ibid., 1950-7-27, T. Kodama 634 (TNS-VS-247431);

Kyushu - Fukuoka Pref.: Fukuoka-shi, Nishi-ku, Ryoshiiwa-yama,

Hyogo Pref.: Kobe-shi Higashinada-ku, Rokko-san, 1979-11-3, T.

alt. 820 m, 1967-10-15, S. Tsutsui 4983 (TNS-VS-431738); Buzen-

Shiraiwa 9919 (TNS-VS-509259); Shikama-gun, Yumesaki-cho,shi, Inugadake, alt. 1000 m, 1969-9-15, S. Tsutsui 6710 (TNS-

Seppiko-san, 1978-9-11, K. Kobayashi s.n. (TNS-VS-831693);

VS-431735); Kasuya-gun, Umi-machi, Goura-tani, alt. 580 m,

Shiso-gun, Ichinomiya-cho, Mizotani, 1977-9-15, T. Shiraiwa

1972-7-30, S. Tsutsui 8612 (TNS-VS-431739); Saga Pref.:7328 (TNS-VS-563484); Yabu-gun, Yabu-cho, Mochikochi, 1972-

Kashima-shi, Kyogadake, 1981-11-3, Y. Kuranari s.n. (TNS-

8-13, T. Shiraiwa 2719 (TNS-VS-563488); Oya-cho, Yokoyuki,

VS-425080); Oita Pref.: Oita-gun, Shonai-cho, Kuro-dake, 1960-

1968-8-12, T. Shiraiwa 1313 (TNS-VS-563482); Asago-gun,

7-30, M. Hadano s.n. (TNS-VS-420031); Naoiri-gun, Kuju-machi,

Ikuno-cho, Kami-ikuno, 1977-8-15, T. Shiraiwa 7176 (TNS-

Hiiji-dake, 1978-8-26, M. Arakane s.n. (TNS-VS-1159828); Kuro-

VS-563486); Tochihara, 1977-9-23, T. Shiraiwa 7399 (TNS-

dake, 1968-8-9, K. Shono 1048 (TNS-VS-285997); ibid., 1968-8-

VS-563485); Hikami-gun, Hikami-cho, Kora, 1976-9-23, T. Shiraiwa

30, K. Shono 1053 (TNS-VS-286002); ibid., 1968-9-8, K. Shono

5863 (TNS-VS-563483); Ichijima-cho, Nago-ike - Godai-san, alt.

1061 (TNS-VS-286010); ibid., 1968-9-8, K. Shono 1070 (TNS-

120–400 m, 1997-11-16, N. Kurosaki 21299 (TNS-VS-697399);

VS-286020); ibid., 1968-9-8, K. Shono 1071 (TNS-VS-286021);

Taki-gun, Sasayama-cho, Koganegadake, 1954-10-24, M. Inada

Miyazaki Pref.: Nishiusuki-gun, Takachiho-cho, Sobo-san, 1972-

3494 (TNS-VS-563489); ibid., 1954-10-24, T. Kodama s.n. (TNS-

8-16, T. Minamitani 7656 (TNS-VS-427819); Kagoshima Pref.:VS-129842); Nara Pref.: [detailed locality unknown], 1913-9-?,

Kanoya-shi, Ontake, 1980-10-10, K. Takesako s.n. (TNS-

U. Faurie 308 (TNS-VS-194773); Yoshino-gun, Tenkawa-mura,

VS-434207).

Sanjo-ga-tsuji, 1981-8-23, Z. Tsujimoto s.n. (TNS-VS-1159799,
1159831); Totsukawa-mura, Okusato, 1976-10-24, Z. Tsujimotos.n. (TNS-VS-423335); Wakayama Pref.: Ito-gun, Koya-cho,
Otaki,

1978-8-12,

H.

Managos.n.

(TNS-VS-549123);

Higashimuro-gun, Nachikatsuura-cho, Okumotori, 1978-11-5, K.
Ohora s.n. (TNS-VS-549124); Kozagawa-cho, 1977-11-3, K.
Ohora s.n. (TNS-VS-549125); Nachi-cho, Nachi-san, 1942-10-13,
Y. Satake 24954 (TNS-VS-294053); Tottori Pref.: Yazu-gun,
Kawahara-cho, Somagoya, 1981-8-12, A. Tanaka s.n. (TNSVS-433502); Saji-son, Yodo, 1979-10-14, A. Tanaka s.n. (TNSVS-424989); Chizu-cho, Okinoyama, 1958-7-27, A. Tanaka 785(TNS-VS-424990); Ashizu, 1980-10-26, A. Tanaka 28440 (TNSVS-433121); Nagi-san, 1981-9-20, K. Yasuhara s.n. (TNSVS-433500); Okayama Pref.: Maniwa-gun, Katsuyama-cho,Taniyama, 1984-9-16, H. Kobatake s.n. (TNS-VS-1152567);
Tomata-gun, Kamo-cho, Kurami, 1976-9-8, T. Saito s.n. (TNSVS-1172275, 1172229); ibid., 1975-9-19, T. Saito s.n. (TNSVS-1172224); Kamisaibara-son, Endo, 1979-9-16, T. Saito s.n.
(TNS-VS-1172227); Aba-son, Osugi, 1980-8-21, K. Takayama s.n.

Thelypteris japonica (Baker) Ching×T. musashiensis(Hiyama) Nakato, Sahashi et M.Kato [Ai-harigane-warabi]*Specimens with stramineous stipes.
Honshu - Aomori Pref.: Kamikita-gun, Towadako-machi, Ouse,
1981-10-13, M. Neichi s.n. (TNS-VS-436671*); Goshogawaratai,
alt. 800 m, 1979-10-27, M. Neichi s.n. (TNS-VS-570290*); Iwate
Pref.: Ichinoseki-shi, Sukawa Spa, 1979-10-11, Y. Nakajima s.n.
(TNS-VS-440279*); Miyagi Pref.: Sendai-shi, Moniwa, Taihakusan, 1979-10-20, K. Shoji s.n. (TNS-VS-575750*); Ido-hama,
1979-9-22, K. Shoji s.n. (TNS-VS-1160530*); Natori-shi, Takadatekumanodo, 1981-10-31, K. Matsunaga 4462 (TNSVS-1152618*); Akita Pref.: Kitaakita-gun, Takanosu-machi,
Yosuketai, 1980-9-12, M. Hatakeyama s.n. (TNS-VS-419749*);
Ryugamori, 1962-8-5, M. Hatakeyama s.n. (TNS-VS-1172243);
Yamagata Pref.: Yonezawa-shi, Nadera-yama, 1991-9-22, Z. Sato
1831 (TNS-VS-1157036*); Murayama-shi, Koshiki-dake, 1974-98, S. Kato s.n. (TNS-VS-437623*); Nishiokitama-gun, Ogunimachi, Jushiga-mori, 1969-9-10, S. Kato s.n. (TNS-VS-437818);

84Atsushi Ebihara and Narumi Nakato
Fukushima Pref.: Kitakata-shi, Hirabayashi, 1980-11-19, Z.
Kaneda 6517 (TNS-VS-438936*); Minamiaizu-gun, Tadami-

VS-1042097*); Kanra-machi, Ueno, 1987-10-10, S. Kojima s.n.

machi, 1980-9-4, Z. Kaneda s.n. (TNS-VS-438928); Kawanuma-

Dainichido, 1996-9-21, T. Iwata 46921 (TNS-VS-1133832);

gun, Yanaizu-machi, Yanaizu, 1980-10-11, A. Baba s.n. (TNSVS-439538*); Onuma-gun, Showa-mura, Yanohara, 1981-10-12,

Urayama, Kawamata, 1981-8-25, S. Kobayashi s.n. (TNSVS-514479*); Hanno-shi, Atera, Nagasawa, 1975-10-10, T.

A. Baba s.n. (TNS-VS-427599*); Higashishirakawa-gun, Tana-

Nakaike s.n. (TNS-VS-333387); Higashiagano - Ogami-yama,gura-machi, Tsutsukowake Shrine, 1980-10-6, Z. Kaneda 6163
(TNS-VS-439164*); Yamamoto-fudo Valley, 1975-8-26, Z.

1975-9-28, T. Nakaike s.n. (TNS-VS-324823); Iruma-gun, Naguri-

Kaneda s.n. (TNS-VS-438932); Yamatsuri-machi, Kamimyoga,

Anazawa, alt. 320 m, 2002-9-1, N. Nakato 2468 (TNS-

1980-10-6, Z. Kaneda 6179 (TNS-VS-1172242); Ibaraki Pref.:VS-1168505); Fujidana-yama, 1985-10-13, S. Ishiguro s.n. (TNS-

Hitachi-shi, Narusawa-cho, 1973-9-16, N. Wada 1053 (TNS-

VS-464002); Agano-mura, Agano, 1953-9-0, H. Inoue s.n. (TNSVS-1042092*); Izugatake, 1934-10-17, S. Okuyama 102175

VS-595306); Takahagi-shi, Yanagisawa, 1972-8-16, M. Yasu s.n.
(TNS-VS-422596*); Kitaibaraki-shi, Hanazono-yama, 1966-8-22,
H. Fujita s.n. (TNS-VS-422600); Tsukuba-shi, Kurakake, 1988-7-(TNS-VS-1156191); Saitama Pref.: Chichibu-shi, Urayama,mura, Shirayazawa, 1964-10-25, H. Ito s.n. (TNS-VS-1045694);(TNS-VS-1166297); Chichibu-gun, Yokoze-machi, Maruyama,
1994-10-21, T. Iwata 40488 (TNS-VS-1025548*); Ubugawa,

14, S. Matsumoto s.n. (TNS-VS-763917); Naka-gun, Tokai-mura,
Shirakata, 1973-11-18, M. Yasu s.n. (TNS-VS-422584*); Miwa-

1987-8-23, T. Iwata 7957 (TNS-VS-1133717); Yokoze-mura,

mura, Torinoko-yama, alt. 350 m, 1985-8-25, H. Kubota s.n. (TNSVS-799566); Kuji-gun, Kanasago-mura, Nakano, 1973-8-20, M.

machi, Kamafuse Pass, 1987-10-18, T. Iwata 8241 (TNSVS-1133720*); Otaki, 1987-10-18, T. Iwata 8245 (TNS-

Yasu s.n. (TNS-VS-422573); Satomi-mura, Inohana Pass, 1972-9-

VS-1133721);

17, N. Wada 1048 (TNS-VS-595342); Daigo-machi, Yamizo-san,(TNS-VS-425194*, 1172258); Ogano-machi, Nunosawa, 1983-12-

1958-6-0, T. Matsumoto s.n. (TNS-VS-1042045); Taga-gun, Juuo-

4, T. Iwata 2400 (TNS-VS-1133780); Nagaru, 1991-10-27, T.

machi, Tatsuware-yama, 1980-10-12, M. Yasu s.n. (TNS-

Iwata 10697 (TNS-VS-1133856); ibid., 1991-10-27, T. Iwata

VS-422593); Niihari-gun, Sakura-mura, University of Tsukuba,

10698 (TNS-VS-1133857); Futtoyama, 1990-9-24, T. Iwata 10004

Matsumi, 1978-11-2, M. Michikawa 10043 (TNS-VS-1080197);(TNS-VS-1133866), Kamihara, 1981-8-25, S. Kobayashi s.n.

Makabe-gun, Makabe-machi, Yubukuro Pass, alt. 150 m, 1998-818, A. Satoh TKB0017 (TNS-VS-9508616*); Tochigi Pref.: Toch-(TNS-VS-514516); Ryokami-mura, 1994-10-8, T. Iwata 40267

igi-shi, Izuru-yama, 1979-10-10, M. Usui s.n. (TNS-VS-422697);(TNS-VS-1133865); Otaki-mura, Taiyoji Temple, 1987-11-6, T.

Akami-cho, 1981-10-15, M. Usui s.n. (TNSVS-1160531*); Karasawa-yama, 1980-10-5, M. Usui s.n. (TNS-

Iwata 8275 (TNS-VS-1133722); Tochimoto, 1993-9-11, T. Iwata

VS-570769*);

H.

VS-1133851); Myohogatake, 1988-10-15, T. Iwata 9063 (TNS-

Kobayashi s.n. (TNS-VS-1172261); Nikko-shi, Konosu-yama,

VS-1133898); Arakawa-mura, Kawauradani, 1984-9-15, T. Iwata

1981-9-21, M. Usui s.n. (TNS-VS-427391); ibid., 1979-10-26, M.
Usui s.n. (TNS-VS-570770*); Moka-shi, Nemoto-yama, alt.

3489 (TNS-VS-1133772); ibid., 1984-9-15, T. Iwata 3490 (TNS-

100 m, 1987-8-2, Y. Sugita s.n. (TNS-VS-799749); Kawachi-gun,Otofuji-yama, 1990-11-19, T. Iwata 10166 (TNS-VS-1133862);

Shiroyama-mura, Kogashi-yama, 1941-10-12, Y. Jotani s.n. (TNSVS-831874*); Kamitsuga-gun, Awano-machi, Shimonagano,

Yadake, 1989-10-1, T. Iwata 9659 (TNS-VS-1133872); Anya-

1978-10-14, T. Waku 9058 (TNS-VS-1172257); Shioya-gun, Shio-chichibu-mura, Hagidaira, 1985-11-1, T. Iwata 5771 (TNS-

bara-machi, Hiruga-dake, 1961-8-5, T. Tashiro s.n. (TNS-

VS-1133750);

VS-422702); Shioya-machi, Tamanyu, 1980-8-8, T. Tashiro s.n.

1979-10-14, K. Nakamura s.n. (TNS-VS-424240); Narita-shi,(TNS-VS-422692); ibid., 1982-9-19, T. Tashiro s.n. (TNSVS-1152581*); Nasu-gun, Bato-machi, Minamidaira, 1981-9-16,

Kamifukuda, 1978-9-29, I. Ueda s.n. (TNS-VS-424244); Tokyo

M. Usui s.n. (TNS-VS-427388); Yamata, 1984-10-26, M. Usui s.n.

VS-517192); Hachioji-shi, Imakuma Shrine, alt. 300–400 m, 1974-(TNS-VS-1152587); Kurobane-machi, Unganji, 1967-9-6, H. Ito

Sano-shi,Kanuma-shi,

Futamata-yama,

1978-9-23,

Bukozan, 1952-8-0, H. Ito s.n. (TNS-VS-1042134); Minano-

Shimotano,

1981-11-4,

S.

Kobayashis.n.

(TNS-VS-1133843); Azumaya-san, 1990-9-30, T. Iwata 10018

11232 (TNS-VS-1133849); ibid., 1993-9-11, T. Iwata 11240 (TNS-

VS-1133771); ibid., 1987-8-9, T. Iwata 7867 (TNS-VS-1133725);

gawa, 1981-11-4, S. Kobayashi s.n. (TNS-VS-1172259); HigashiChiba

Pref.:

Ichikawa-shi,

Kashiwai-machi,

Pref.: Ota-ku, Ikegami, 1886-9-?, K. Shirai s.n. (TNS-(TNS-

9-22, T. Nakaike s.n. (TNS-VS-335120); Kamiange, 1953-9-6, Y.
Tokashiki s.n. (TNS-VS-543245*); Ome-shi, 1981-8-31, H. Endos.n. (TNS-VS-435238*); Nishitama-gun, Itsukaichi-machi, Bon-

VS-1045682*); Takakuhei, alt. 540 m, 1985-8-27, H. Ogura s.n.

bori-gawa, 1978-9-17, T. Yoshizawa s.n. (TNS-VS-831890); Oku-

(TNS-VS-799757); Nasu Imperial Villa, 1963-8-28, H. Ito s.n.

tama-machi, the foot of Bonoore-yama, alt. 440 m, 2003-9-29, N.

(TNS-VS-1045683); Aso-gun, Tanuma-machi, Narabu-san, alt.

Nakato 2546 (TNS-VS-1168504); Atago-yama, alt. 440 m, 2003-730, N. Nakato 2526 (TNS-VS-1168503*); Okumotori-dani, 1980-

s.n. (TNS-VS-1045392, 1045393, 1045394, 1045395, 1045684);
Nasu-machi,

Tarasawa,

1964-8-22,

H.

Ito

s.n.

520 m, 1987-9-14, T. Noguchi & Y. Tamura s.n. (TNSVS-799747); Gunma Pref.: Seta-gun, Fujimi-mura, Akagi-san,
1957-8-31, T. Matsuzawa s.n. (TNS-VS-1042086*); Kanra-gun,

9-23, H. Endo s.n. (TNS-VS-423943); Koizawa, 1967-11-5, H. Ito

Shimonita-machi, Arafune-yama, 1956-9-9, T. Satomi 17 (TNSVS-1042082*); ibid., 1956-9-9, T. Satomi s.n. (TNS-

S. Okuyama 100863 (TNS-VS-1166298); Kanagawa Pref.: Min-

VS-1042088*);

VS-441338); Niigata Pref.: Shibata-shi, Miyayorikami, 1946-10-

ibid.,

1955-9-4,

T.

Satomi

s.n.

(TNS-

s.n. (TNS-VS-1045688); Nariki-mura, Takamizu-san, 1933-9-18,
amiashigara-shi, Kano, 1977-5-22, H. Kobayashi s.n. (TNS-Distribution of the Thelypteris japonica Complex (Thelypteridaceae)

85

3, T. Uchikawa s.n. (TNS-VS-543162); Sugatani, alt. 180 m,(TNS-VS-637704); Egasha, alt. 170 m, 1983-9-18, T. Sato 4578

1962-10-21, collector unknown s.n. (TNS-VS-929607); Nakakan-(TNS-VS-1156115); Saori, 1978-10-12, T. Sato s.n. (TNS-

bara-gun, Muramatsu-machi, Atago-yama, alt. 50 m, 1962-8-29, F.

VS-1157107); Tagata-gun, Amagiyugashima-cho, Joren-no-taki

Konta 910 (TNS-VS-929606); Minamikanbara-gun, Shitada-mura,

Fall, 1971-10-10, H. Ito s.n. (TNS-VS-1092284); Kamiohmi-mura,

Komyo-zan, 1979-8-28, M. Saito 3158 (TNS-VS-543163);

Amagi-san, 1938-8-25, M. Nakaji s.n. (TNS-VS-782508); Aichi

Nakauonuma-gun, Nakasato-mura, Atema-yama, 1986-10-8, M.

Pref.: Higashikamo-gun, Asahi-cho, Komayama, 1979-10-28, K.

Ono s.n. (TNS-VS-831858); Iwafune-gun, Asahi-mura, Saruta

Inukai 2642 (TNS-VS-432950); Kitashitara-gun, Shitara-cho,
Higahsinagura, 1979-9-24, K. Inukai 2536 (TNS-VS-432957*);

Dam, 1986-10-18, M. Saito s.n. (TNS-VS-1157044); Sado-gun,
Aikawa-machi, Funayama, 1979-10-16, M. Saito 3245 (TNSVS-543159*); Toyama Pref.: Kurobe-shi, Oyama, 1978-8-10, T.

Toyone-mura,Ohshima s.n. (TNS-VS-543549*); Shimoniikawa-gun, Unazuki-(TNS-VS-831715); Minamishitara-gun, Horai-cho, 1967-11-19, C.

machi, Kitamata - Iburi-yama - Asahi-goya, 1969-8-27, N. Satomi& R. Mochizuki s.n. (TNS-VS-256057*); Ishikawa Pref.:Ohkawa s.n. (TNS-VS-428527); Shiga Pref.: Hikone-shi, Amat-

Kanazawa-shi, Daimon-zan, 1975-10-10, M. Honda s.n. (TNSVS-424504*); Wajima-shi, Nishide, 1973-10-4, S. Komaki s.n.

Pref.: Kitakuwada-gun, Miyama-cho, Ashu Experimental Forest of(TNS-VS-424536*); Enuma-gun, Yamanaka-machi, 1931-10-20,

831723); Kuroda-mura, Uodani Pass, 1952-9-23, T. Kodama 2042

1995-8-27,

J.

Haginiwa

JH034067(TNS-

VS-984067); Tsugu-mura, Okuwa, 1985-9-1, H. Miyazaki 6970

subo-yama, 1978-10-15, T. Tatebe s.n. (TNS-VS-533916); Kyoto
Kyoto University, 1987-10-1, S. Masuda s.n. (TNS-VS-831722,

N. Hiratsuka 114 (TNS-VS-513158); Ishikawa-gun, Shiramine-(TNS-VS-129836); Hyogo Pref.: Taka-gun, Kami-cho, Toba,mura, Ichinose, 1979-9-15, K. Yoneyama s.n. (TNS-VS-424517);

1976-10-10, T. Shiraiwa 6129 (TNS-VS-563790); Asago-gun,

Fukui Pref.: Ono-gun, Izumi-mura, Shimoyama, Otaru, 1967-10-

Ikuno-cho, Tochihara, 1977-9-23, T. Shiraiwa 7420 (TNS-

1, Y. Saito s.n. (TNS-VS-1166188); Yamanashi Pref.: Kitakoma-

VS-563784); Santo-cho, Awaga, 1976-8-15, T. Shiraiwa 5644
(TNS-VS-563780*); Hikami-gun, Kamonosho-mura, Myoko-san,gun, Sutama-cho, Masutomi, 1963-8-11, K. Akiyama s.n. (TNSVS-573506*); Nagano Pref.: Omachi-shi, Iyari, 1980-8-21, F.

1929-10-27,

Y.

Araki

6175

(TNS-VS-194865);

Taki-gun,(TNS-VS-831765*); ibid., 1980-8-21, F.
Miyamoto 6712 (TNS-VS-831766*); ibid., 1980-8-24, T. Nakaike

Sasayama-cho, Shitsukawakami, 1977-10-10, T. Shiraiwa 7447

s.n. (TNS-VS-1159818*); Kuzu Spa, alt. 1000 m, 1981-9-8, H.
Tsutsumi s.n. (TNS-VS-574425*); Saku-shi, Kabutoiwa-yama,(TNS-VS-563787); Tsuna-gun, Ichinomiya-cho, Yamada, Fudo-

1980-9-28, K. Nakayama 34 (TNS-VS-401536*); Suwa-gun,Pref.: Tottori-shi, Manidera, 1951-3-21, A. Tanaka 981 (TNS-

Miyagawa-mura, 1952-9-5, S. Gomi 11 (TNS-VS-1042070); Shi-

VS-433503); Shimane Pref.: Iishi-gun, Tonbara-cho, Oyorogi-

moina-gun, Takagi-mura, Shimoujinori, alt. 600 m, 1981-10-20, H.

yama, 1977-8-11, H. Yamamoto s.n. (TNS-VS-511261); Okayama

Tsutsumi s.n. (TNS-VS-574298); Kiso-gun, Kaida-mura, Sekiya,
alt. 1300 m, 1981-8-27, H. Tsutsumi s.n. (TNS-VS-574441*);

Pref.: Tsuyama-shi, Kurosawa-yama, 1980-7-11, K. Takayama s.n.

Okuwa-mura, Adera Valley, 1980-8-25, T. Nakaike s.n. (TNS-

T. Saito s.n. (TNS-VS-1172260); Kawakami-gun, Nariwa-cho,VS-1157078); Gifu Pref.: Ena-shi, Kasagi-yama, 1984-9-30, H.

Tenjin-yama, 1975-11-21, T. Watanabe s.n. (TNS-VS-1172262);

Miyazaki 6607 (TNS-VS-831714); Ena-gun, Kashimo-mura,

Hiroshima Pref.: Saeki-gun, Saeki-cho, Iiyama, 1971-10-10, T.

Higashihondani, coll. date unknown, Y. Yamamoto s.n. (TNSVS-435837*); Hirukawa-mura, Nakagiri, 1978-11-25, Y. Yamamoto

Tamaru 391 (TNS-VS-547835); Yoshiwa-mura, Kanmuri-yama,s.n. (TNS-VS-1157147); Yamaoka-cho, Hara, Nakada, 1981-9-27,

Shikoku - Ehime Pref.: Kamiukena-gun, Omogo-mura, Omogo Val-T. Ogiyama s.n. (TNS-VS-545768); Akechi-cho, Sugidaira, 197710-16, Y. Yamamoto s.n. (TNS-VS-435762*); Shizuoka Pref.:ley - Ishizuchi-yama, 1965-9-29, S. Masuda 4656 (TNS-

Atami-shi, Izumi, 1980-11-8, A. Suzuki s.n. (TNS-VS-509163*);

VS-831698); Kita-gun, Nagahama-cho, Kinzansyussekiji Temple,

Mishima-shi, Yamanaka-shinden, 1979-8-19, A. Suzuki s.n. (TNSVS-591592*); Fujinomiya-shi, Asagiri-kogen, alt. 860 m, 1985-10-

Nagaoka-gun,

2, T. Sato 8619 (TNS-VS-929581), ibid., 1985-10-2, T. Sato 8620

unknown, H. Inoue 149 (TNS-VS-1042130).

Miyamoto

6707(TNS-VS-763782); Koganegadake, 1976-10-25, T. Shiraiwa 6213
no-taki Fall, 1981-9-20, K. Mune 554 (TNS-VS-509265); Tottori(TNS-VS-423264); Jobo-gun, Kayo-cho, Higashikario, 1981-9-27,

1971-10-10, T. Tamaru 391 (TNS-VS-547880).

VS-831697);

ibid.,

1965-9-29,

S.

Masuda

4654

(TNS-

1980-7-30, H. Ishikawa s.n. (TNS-VS-1166280); Kochi Pref.:Nishitoyonaga-mura,

Kajigamori,

coll.

date

(TNS-VS-929580); ibid., alt. 170 m, 1983-9-18, T. Sato 4579

Kyushu - Oita Pref.: Naoiri-gun, Kuju-machi, Kuro-dake, K. Shono(TNS-VS-929576); Inokashira, 1961-9-2, J. Sugimoto s.n. (TNS-s.n. (TNS-VS-1168492); Akagawa, 1973-10-14, M. Hadano s.n.

VS-247757); Fumoto, alt. 800 m, 1974-8-29, F. Konta 10474(TNS-VS-1157097).