Bull. Natn. Sci. Mus., Tokyo, Ser. B, 32(4), pp. 171–173, December 22, 2006

Re-designation of Seventeen Lectotypes for Diatoms(Bacillariophyceae), Originally Designated byOhtsuka & Tuji (2002), Tuji (2002) and Tuji (2004)
Akihiro Tuji
Department of Botany, National Science Museum,
4–1–1, Amakubo, Tsukuba, 305–0005, Japan
E-mail: tuji@kahaku.go.jp

Abstract Seventeen lectotypes of diatoms, which were originaly designated by Ohtsuka & Tuji(2002), Tuji (2002) and Tuji (2004), are re-designated in this paper. Because of lacking the phrase“here designated” (hic designatus) or an equivalent in the typification statements, these original
designations are invalid under International Code for Botanical Nomenclature (ICBN: Saint LouisCode) Art. 7.11.
Key words : Lectotype, Caloneis nipponica Skvortzov, Coconeis disculus var. nipponica Skvortzov, Cymbella nipponica Skvortzov Skvortzov, Cymbella turgidula var. nipponica Skvortzov, Gomphonema lingulatum var. elongatum Skvortzov, Navicula costulata var. tenuirostris Skvortzov,Navicula gastrum fo. nipponica Skvortzov, Navicula hasta var. gracilis Skvortzov, Navicula lambda var. nipponica Skvortzov, Navicula lanceolata var. nipponica Skvortzov, Navicula radiosa fo.
nipponica Skvortzov, Navicula rostellata var. biwaensis Skvortzov, Navicula similis var. nipponica
Skvortzov, Navicula undulate Skvortzov, Melosira solida var. nipponica Skvortzov, Fragilaria
rhabdosoma Ehrenb., Fragilaria sepes Ehrenb.

Introduction
Designations for diatoms by Ohtsuka & Tuji(2002), Tuji (2002) and Tuji (2004) are invalid
under International Code of Botanical Nomenclature (ICBN, Saint Louis Code: Greuter et al.
2000), Art. 7.11., because of lacking the phrase“here designated” (hic designatus) or an equivalent in the typification statements. In this paper,
we re-designate these lectotypes.
Designation
Caloneis nipponica Skvortzov, Philipp. J. Sci. 6:
267. pl. 2, f. 7, pl. 3, f. 9, pl. 4, f. 15. 1836;
Ohtsuka & Tuji, Phycol. Res. 50: 244. f. 1–2.
2002.
Lectotype (designated here): Slide 30000136,
Lake Biwa Museum, Japan.
Isolectotype (isotype raw material): TNS-AL-53965m in TNS (Department of Botany, National Science Museum, Tokyo).
Coconeis disculus var. nipponica Skvortzov,Philipp. J. Sci. 6: 262. pl. 4, f. 16. 1836; Ohtsuka & Tuji, Phycol. Res. 50: 244. f. 3–4. 2002.
Lectotype (designated here): Slide 30000142,
Lake Biwa Museum, Japan.
Isolectotype (isotype raw material): TNS-AL53965m in TNS (Department of Botany, National Science Museum, Tokyo).
Cymbella nipponica Skvortzov, Philipp. J. Sci.
6: 283–4. pl. 5, f. 20, 23. 1836; Ohtsuka &Tuji, Phycol. Res. 50: 245. f. 5–6. 2002.
Lectotype (designated here): Slide 30000144,
Lake Biwa Museum, Japan.
Isolectotype (isotype raw material): TNS-AL53965m in TNS (Department of Botany, National Science Museum, Tokyo).
Cymbella turgidula var. nipponica Skvortzov,Philipp. J. Sci. 6: 283. pl. 2, f. 8, pl. 4, f. 4.

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Akihiro Tuji

1836; Ohtsuka & Tuji, Phycol. Res. 50: 245. f.
7–8. 2002.
Lectotype (designated here): Slide 30000136,
Lake Biwa Museum, Japan.
Isolectotype (isotype raw material): TNS-AL53965m in TNS (Department of Botany, National Science Museum, Tokyo).
Gomphonema lingulatum var. elongatum
Skvortzov, Philipp. J. Sci. 6: 286. pl. 3, f. 11.
1836; Ohtsuka & Tuji, Phycol. Res. 50: 246. f.
9–10. 2002.
Lectotype (designated here): Slide 30000143,
Lake Biwa Museum, Japan.
Isolectotype (isotype raw material): TNS-AL53965m in TNS (Department of Botany, National Science Museum, Tokyo).
Navicula costulata var. tenuirostris Skvortzov,
Philipp. J. Sci. 6: 273. pl. 5, f. 16. 1836; Ohtsuka & Tuji, Phycol. Res. 50: 246. f. 11–12.
2002.
Lectotype (designated here): Slide 30000140,
Lake Biwa Museum, Japan.
Isolectotype (isotype raw material): TNS-AL53965m in TNS (Department of Botany, National Science Museum, Tokyo).
Navicula gastrum fo. nipponica Skvortzov,
Philipp. J. Sci. 6: 274. pl. 8, f. 8. 1836; Ohtsuka & Tuji, Phycol. Res. 50: 246. f. 13. 2002.
Lectotype (designated here): Slide 30000144,
Lake Biwa Museum, Japan.
Isolectotype (isotype raw material): TNS-AL53965m in TNS (Department of Botany, National Science Museum, Tokyo).
Navicula hasta var. gracilis Skvortzov, Philipp.
J. Sci. 6: 275. pl. 7., f. 9. 1836; Ohtsuka &Tuji, Phycol. Res. 50: 246. f. 14–16. 2002.
Lectotype (designated here): Slide 30000136,
Lake Biwa Museum, Japan.
Isolectotype (isotype raw material): TNS-AL53965m in TNS (Department of Botany, National Science Museum, Tokyo).
Navicula lambda var. nipponica Skvortzov,Philipp. J. Sci. 6: 271. pl. 6, f. 15. 1836; Ohtsuka & Tuji, Phycol. Res. 50: 246. f. 17. 2002.
Lectotype (designated here): Slide 30000130,
Lake Biwa Museum, Japan.

Isolectotype (isotype raw material): TNS-AL53965m in TNS (Department of Botany, National Science Museum, Tokyo).
Navicula lanceolata var. nipponica Skvortzov,
Philipp. J. Sci. 6: 274. pl. 2, f. 18, pl, 7, f. 5.
1836; Ohtsuka & Tuji, Phycol. Res. 50: 247. f.
20. 2002.
Lectotype (designated here): Slide 30000140,
Lake Biwa Museum, Japan.
Isolectotype (isotype raw material): TNS-AL53965m in TNS (Department of Botany, National Science Museum, Tokyo).
Navicula radiosa fo. nipponica Skvortzov,Philipp. J. Sci. 6: 273. pl. 2, f. 2, pl. 3, f. 20.
1836; Ohtsuka & Tuji, Phycol. Res. 50: 247. f.
22–23. 2002.
Lectotype (designated here): Slide 30000143,
Lake Biwa Museum, Japan.
Isolectotype (isotype raw material): TNS-AL53965m in TNS (Department of Botany, National Science Museum, Tokyo).
Navicula rostellata var. biwaensis Skvortzov,
Philipp. J. Sci. 6: 272. pl. 3, f. 14. 1836; Ohtsuka & Tuji, Phycol. Res. 50: 248. f. 24–25.
2002.
Lectotype (designated here): Slide 30000136,
Lake Biwa Museum, Japan.
Isolectotype (isotype raw material): TNS-AL53965m in TNS (Department of Botany, National Science Museum, Tokyo).
Navicula similis var. nipponica Skvortzov,Philipp. J. Sci. 6: 276. pl. 3. f. 2. 1836; Ohtsuka & Tuji, Phycol. Res. 50: 249. f. 26. 2002.
Lectotype (designated here): Slide 30000141,
Lake Biwa Museum, Japan.
Isolectotype (isotype raw material): TNS-AL53965m in TNS (Department of Botany, National Science Museum, Tokyo).
Navicula undulate Skvortzov, Philipp. J. Sci. 6:
275. pl. 4, f. 2, pl. 7, f. 6. 1836; Ohtsuka &Tuji, Phycol. Res. 50: 249. f. 28–29. 2002.
Lectotype (designated here): Slide 30000141,
Lake Biwa Museum, Japan.
Isolectotype (isotype raw material): TNS-AL53965m in TNS (Department of Botany, National Science Museum, Tokyo).

Re-designation of Seventeen Lectotypes for Diatoms (Bacillariophyceae)

Melosira solida var. nipponica Skvortzov,Philipp. J. Sci. 6: 255. pl. 1, f. 1, 2, 21. 1836;
Tuji, Phycol. Res. 50: 315. f. 19–25. 2002.
Lectotype (designated here): A individual in aslide TNS-AL-53965s in TNS (Department ofBotany, National Science Museum, Tokyo).
The photograph is shown at figure 22 in Tuji
(2002).
Isolectotype (isotype raw material): TNS-AL53965m in TNS (Department of Botany, National Science Museum, Tokyo).
Fragilaria rhabdosoma Ehrenb., Abh. K. Akad.
Wiss. Berlin. 1833: 249, 1835; Tuji, 17th Int.
Diat. Symp. 2002: 415. f. 19–28.
Lectotype (designated here): BHUPM.MB.
ECO-004 in BHUPM (Museum fürNaturkunde in Berlin).
Isolectotype (raw material): EC sample 2533 in
BHUPM (Museum für Naturkunde in Berlin).
Isolectotype (slide): TNS-AL-56153s in TNS(Department of Botany, National Science Museum, Tokyo).
Fragilaria sepes Ehrenb., Mikrogeologie pl.
37/1, f. 8. 1854; Tuji, 17th Int. Diat. Symp.
2002: 419. f. 34–43.
Lectotype (designated here): BHUPM.MB.
ECO-005 in BHUPM (Museum für173

Naturkunde in Berlin).
Isolectotype (isotype raw material): EC sample
2871 in BHUPM (Museum für Naturkunde inBerlin).
Isolectotype (isotype) (slide): TNS-AL-56154s inTNS (Department of Botany, National ScienceMuseum, Tokyo).
References
Greuter, W., Mcneill, J. Barrie, F. R., Burdet, H.-M.,
Demoulin, V., Filgeiras, T. S., Nicolson, D. H., Silva,P. C., Skog, J. E., Trehane, P., Turland, N. J., andHawksworth, D. L. 2000. International Code of Botanical Nomenclature (St Louis Code). Regnum Vegetabile138. Koeltz Scientific Books, Königstein.
Ohtsuka, T. & Tuji, A. 2002. Lectotypification of somepinnate diatoms described by Skvortzow in 1936 fromLake Biwa. Phycol. Res. 50: 243–249.
Tuji, A. 2002. Observations on Aulacoseira nipponica
from Lake Biwa, Japan, and Aulacoseira solida fromNorth America (Bacillariophyceae). Phycol. Res. 50:
313–316.
Tuji, A. 2004. Type examination of the ribbon-forming
Fragilaria capucina complex described by Christian
Gottfried Ehrenberg. 411–422. Seventeenth International Diatom Symposium 2002 Proceedings. M.Poulin(ed.) Biopress Limited, Bristol.