Bull. Natl. Mus. Nat. Sci., Ser. A, 33(2), pp. 67–83, June 22, 2007

Pamphiliid Sawflies (Insecta, Hymenoptera) from Yamanashi Prefecture,
Central Honshu, Japan
Akihiko Shinohara1, Hirohisa Suda2 and Hideo Takahashi3
1

Department of Zoology, National Museum of Nature and Science,
3–23–1 Hyakunin-cho, Shinjuku-ku, Tokyo, 169–0073 Japan
E-mail: shinohar@kahaku.go.jp
2
2–13–8 Sennari, Sakura-shi, Chiba, 285–0034 Japan3
876–1–203 Higashiasakawa-machi, Hachioji-shi, Tokyo, 193–0834 Japan

Abstract Collection data are given for 36 species of pamphiliid sawflies of the genera Acantholyda, Cephalcia, Onycholyda and Pamphilius from Yamanashi Prefecture, central Honshu. Of
these, one species of Cephalcia has not been determined and two species of Onycholyda and four
species of Pamphilius are newly recorded from Yamanashi Prefecture. Compared with the otherwell-investigated areas in Honshu, the pamphiliid fauna of Yamanashi Prefecture is very rich, having the largest number of species in total (the same number as that of Tochigi Prefecture) and in
the “northern group” of species.
Key words : Hymenoptera, Pamphiliidae, fauna, Yamanashi Prefecture, Honshu, Japan.

Introduction
Yamanashi Prefecture, situated in the middleof Honshu, is not a large prefecture, occupying
an area of 4,465.37 km2, but its natural environments are diversified and fairly well preserved.
About 78% of the area, ranging from 70 to 3,776
m in altitude, is covered with various kinds
of forests, which provide excellent habitats for
phytophagous insects. The sawfly fauna of Yamanashi Prefecture should be very rich, but ourknowledge on this subject is rather poor and only
scattered collection records published in various
taxonomic works are available for the moment.
The first record of the sawfly family Pamphiliidae from Yamanashi Prefecture dates back to
Yano’s (1919, 1920) reports on the infestation oflarch by “Cephaleia kaebelei [sic] Rohwer”.
Since then, 28 species of the family were added
to the Yamanashi fauna by Takeuchi (1938)
and Shinohara (1985a, etc.; see under respective
species).
The present work is an attempt to get together
all the information on the family Pamphiliidae in
Yamanashi Prefecture. All the published records

are summarized and new collection records are
given mainly based on the study of the collectionof the National Museum of Nature and Science,
Tokyo (NSMT), and the private collection of H.
Suda (HSC). Other depositories of the material
examined are H. Kumamoto collection (HKC),
H. Nagase collection (HNC), and Kobe University collection (KU).
We wish to thank H. Kumamoto (Hirakata), H.
Nagase (Kamakura), and T. Naito (Kobe University, Kobe) for the loan of the material.
Results and Discussion
We have recognized 36 species of four generaof Pamphiliidae occurring in Yamanashi Prefecture. They are seven species of Acantholyda,
three species of Cephalcia, seven species of
Onycholyda, and 19 species of Pamphilius(see Table 1 and the enumeration below). Of
these, 29 species are already known to occur
in Yamanashi Prefecture, while six species are
newly recorded in this work. One remaining
species, a Cephalcia, has not been identified.

x
x
x
x
x
x
x
x

x

x

x

x
x
x
x

x
x

x

x

x
x
x

x

x

x

x

Hyogo Pref.
Ishikawa Pref.
[0–1510 m alt.] [0–2702 m alt.]

x
x
x

x
x
x
x

x

x
x
x
x
x
x

x
x
x
x

x
x
x
x

x

x

x

x
x
x

x

x
x

x

Tochigi Pref.
[0–2578 m alt.]

x
x
x
x
x
x

x

Kamiange[ca. 400 m alt.]xxx

Yamanashi Pref.
[70–3776 m alt.]

Pamphiliidae recorded in six areas in Honshu (undetermined species not included).

Subfamily CEPHALCIINAE
Genus ACANTHOLYDA
Subgenus Acantholyda
A. nipponica
Subgenus Itycorsia
A. albomaculata
A. alpina
A. iwatai
A. kojimai
A. mizunoi
A. posticalis posticalis
A. sasakii
A. tsuyukii
A. laricis
Genus CEPHALCIA
C. lariciphila japonica
C. koebelei
C. variegata
Subfamily PAMPHILIINAE
Genus NEUROTOMA
Subgenus Neurotoma
N. iridescens
Subgenus Gongylocorsia N. atrata
N. harai
Genus CHRYSOLYDA
C. leucocephala
Genus ONYCHOLYDA
O. amplecta groupO. decorata
O. esakii
O. lucidaO. minomalis
O. moriutii
O. similis
O. tenuis
O. viriditibialis
O. luteicornis group
O. kumamotonis

Table 1.

x
x

x
x
x

x

x

x

x
x

x

Aomori Pref.
[0–1625 m alt.]

68
Akihiko Shinohara, Hirohisa Suda and Hideo Takahashi

Heavily shadowed: Species of the “basic group”

Main references

Genus PAMPHILIUS
P. basilaris groupP. basilaris
P. alternans group
P. komonensis
P. takeuchii
P. sulphureipes group
P. ishikawai
P. zhelochovtsevi nipponicus
P. histrio groupP. brevicornis ibukii
P. tricolor
P. vafer groupP. confusus
P. flavipectus
P. nakagawai
P. lobatus
P. venustus
P. kamikochensis
P. archiducalis
P. naokoae
P. opacus
P. hortorum
P. itoi
P. varius
P. togashii
P. sylvaticus groupP. alnicola
P. benesi
P. daisenus
P. gracilis
P. japonicus
P. volatilis
P. inanitus group
P. hilaris
Total number of species
B/T ratio (see text for explanation)
N/B ratio (see text for explanation)
x
x

x
x

x
x
35
0.46
1

Shinohara(2002)

x
x
x
19
0.84
0

x
x
x
x
x
x

x

x
x
x
x

Kamiange[ca. 400 m alt.]

x

x
x
x
x
x
x
x
x
x
x

x
x
x

32
0.59
0.53
Togashi (1998)
Naito et al.
Shinohara andpresent work
(2004)
Yamada (2005)
Lightly shadowed: Species of the “northern group”

x

x
x

24
0.75
0.17

x
x
x

x

x

x
x
x

x

x
x
x
x
x

x
x
x

x

x
x
x
x
x

Yamanashi Pref.
[70–3776 m alt.](Continued).

Hyogo Pref.
Ishikawa Pref.
[0–1510 m alt.] [0–2702 m alt.]

Table 1.

x
x
35
0.51
0.78
Nakamura (2003)
Shinohara and
Yamada (2005)

x
x

x
x
x
x

x
x
x
x
x
x

x
x
x

Tochigi Pref.
[0–2578 m alt.]

Shinohara and
Yamada (2005)

26
0.62
0.56

x
x

x
x

x

x
x
x
x
x
x

x
x
x
x

Aomori Pref.
[0–1625 m alt.]

Pamphiliid Sawflies from Yamanashi Prefecture69

70

Akihiko Shinohara, Hirohisa Suda and Hideo Takahashi

Noteworthy species and probable additions in the
futureThe occurrence of three rare species of Pamphilius in Yamanashi Prefecture is noteworthy.
Pamphilius naokoae Shinohara, 1999, which isendemic to Yamanashi Prefecture, is known only
from the type series collected at Masutomi-kosen
in the northern part of the prefecture (for more
information, see comments under the species).
Pamphilius brevicornis ibukii Shinohara, 1995,
and P. tricolor Beneš, 1974, are also very rare
species. In Honshu, the former is known only by
one specimen each from Ishikawa and Yamanashi
Prefectures and the latter by one specimen eachfrom Niigata and Yamanashi Prefectures.
On the other hand, some relatively common or
widespread pamphiliid species have not beenfound in Yamanashi Prefecture. Among these are
Cephalcia variegata Takeuchi, 1930, Neurotoma
iridescens (André, 1882), Onycholyda kumamotonis (Matsumura, 1912), Pamphilius zhelochovtsevi nipponicus Shinohara, 1993, P. japonicus
Shinohara, 1985, P. togashii Beneš, 1977, and P.
varius (Audinet-Serville, 1823). Of these, C. variegata will certainly be found on Pinus pumila
stands in the subalpine zones of Yatsugatake or
Akaishi Mountains, and P. varius on Betula trees
on the same mountain ranges. Onycholyda kumamotonis will be found on Filipendula in theareas where P. venustus, a species also associatedwith this plant, has been collected. Neurotoma
iridescens is rather common and widespread incentral Honshu. Other three species of Pamphilius are not common but widely distributed in
Honshu, including prefectures around Yamanashi. Further collectings will certainly reveal
the occurrence of these species in Yamanashi
Prefecture. A very rare species, Chrysolyda leucocephala (Takeuchi, 1938), has been collected
at Shiratsuka-rindo on the southwestern slope ofMt Fujisan in Shizuoka Prefecture (1/ , 21. V.
1978, S. Tsuyuki, NSMT); this species is also
likely to be found in Yamanashi Prefecture.
Comparison with other prefectures and nature of
local pamphiliid faunae in Honshu

Table 1 shows the pamphiliid species known tooccur in five prefectures and one area in Honshu.
Besides the entry of Yamanashi Prefecture, thetable was already published by Shinohara and
Yamada (2005). These are the best-investigatedareas in Honshu in terms of pamphiliid fauna.
The pamphiliid fauna of Yamanashi Prefecture isvery rich, comparable only to that of Tochigi Prefecture, according to the present study. In Honshu, a little more species of the Pamphiliidae
may have been recorded or collected in Nagano
Prefecture, but the collection data from the prefecture have not been worked out.
Shinohara and Yamada (2005), based on distribution patterns, roughly classified the pamphiliid
species in Honshu into three categories, the“basic group”, the “northern group”, and the“others”, and proposed to use two different ratios, the B/T ratio and the N/B ratio, for comparing the species compositions of the local pamphiliid faunae in Honshu. The species of the“basic group” occur widely in Honshu (ideally inall prefectures), whereas the species of the“northern group” occur mainly on mountains ofcentral to northern Honshu. The “others” include
rare species without sufficient information on
distribution; possibly, they may eventually be allocated into “basic” or “northern” groups. The
B/T ratio is the ratio of the species number of the“basic group” to the total number of species occurring in the area, whereas the N/B ratio is theratio of the number of species of the “northerngroup” to that of the “basic group” found in thearea. The use of these ratios may help to understand the characteristics of the pamphiliid faunae
at the prefecture or lower levels (see Shinohara
and Yamada, 2005, for more details).
The number of the species included in the“basic group” is expected to increase as ourknowledge on the pamphiliid fauna of Honshu
grows (Shinohara and Yamada, 2005). After thedata of the Yamanashi Prefecture are added, we
propose to treat 20 species (heavily shadowed inTable 1) as belonging to the “basic group”. The
representatives of the “basic group” in this work
are widespread species recorded in four or morePamphiliid Sawflies from Yamanashi Prefectureareas out of the six areas treated and are expected
to be found eventually in all prefectures in Honshu. Pamphilius confusus Shinohara, 2005, P.
nakagawai Takeuchi, 1930, and P. venustus(Smith, 1874) were recorded from four areas, butthey are not likely to be found in western Honshu
because their known host plants are not distributed or rarely found there (Shinohara and Morita,
2003; Shinohara, 2005); therefore, these species
are not included in the “basic group”. The“northern group” (lightly shadowed in Table 1) isalso redefined here to include the species of the
Cephalciinae, the Pamphilius vafer group, and
the Pamphilius histrio group, excluding thespecies included in the “basic group”, andOnycholyda kumamotonis (Matsumura, 1912),
which is very similar to Pamphilius venustus, amember of the P. vafer group, in biology anddistribution.
Figure 1 shows the number of species, allocated into three categories, for each prefecture andarea, and the B/T and N/B ratios calculated foreach prefecture and area are given in Table 1.
Among the six areas, Kamiange is exceptional. It
is not a prefecture but a small area about 400 m
high in the west of Tokyo (Shinohara, 2002). It
shows the highest B/T ratio of 0.84 and the lowest N/B ratio of 0, meaning that its pamphiliid
fauna is mostly composed of the species of the“basic group” and no species of the “northerngroup” have been found there (see also Fig. 1).
This is probably a typical species composition of
the Pamphiliidae in the lowlands of central Honshu. On the other hand, Yamanashi Prefecture
shows the lowest B/T ratio (0.46) and the highest
N/B ratio (1) among the six areas considered.
The low value of the B/T ratio, shared by Tochigi
Prefecture (0.51), simply shows the relatively
small share of the “basic group” and consequently high species diversity of the pamphiliid
sawflies in the prefecture. The very high value of
the N/B ratio, also shared by Tochigi Prefecture(0.78), indicates a very rich representation of thespecies of the “northern group” in Yamanashi
Prefecture.
As shown in Fig. 1, the species of the “basic

71

Fig. 1. Diagram showing the number of speciesof pamphiliid sawflies, classified into three categories, for each of Hyogo, Ishikawa, Yamanashi, Tochigi, and Aomori Prefectures andKamiange area. B (heavily shadowed): The“basic group”. N (lightly shadowed): The“northern group”. O (not shadowed): Others.
T: Total number of species known to occur inthe area. See text for more explanation.

group” may occur in all prefectures in Honshu(thus “basic”) and additional occurrence of the“northern group” species in each prefecture will
determine the total species diversity of the Pamphiliidae in the prefecture. Yamanashi Prefecture,having the largest number of the species of the“northern group” (16), apparently has the mostdiverse pamphiliid fauna in Honshu.

List of Pamphiliidae from Yamanashi Prefecture(The numerals in brackets after the locality
names in the collection data refer tothe numerals in the map, Fig. 2)
Acantholyda albomaculata Shinohara, 1985
[Japanese name: Shiromon-matsu-hiratahabachi]

Distribution.
shu, Shikoku).

Japan (central to western Hon-

72

Akihiko Shinohara, Hirohisa Suda and Hideo Takahashi

Fig. 2. Map of Yamanashi Prefecture. Numerals refer to those given in brackets in each collection data and show
approximate locations of the sites.

Material examined. 1? from Hatchodaira[15], Mt. Kimpu-zan, 1/ (paratype) from Aokigahara [48], 1/ from Fuji-rindo recorded by Shinohara (1985b, 2001a)
Host plant. Unknown.
Remarks. This species has been recorded
from Honshu (Yamanashi, Nagano, Mie, Naraand Hyogo Prefectures) and Shikoku (Kochi Prefecture) (Shinohara, 2001a). In Yamanashi Prefecture, three collection records were publishedby Shinohara (1985b, 2001a).

from Mt. Kimpu-zan [17], all recorded by Shinohara (2001a).
Host plant. Unknown.
Remarks. This species is known to occur in
Honshu (Aomori, Tochigi, Yamanashi, Nagano,
Aichi Prefectures), Shikoku (Tokushima Prefecture), and Kyushu (Kagoshima Prefecture). It
was first recorded from Yamanashi Prefecture by
Shinohara (2001a).
Acantholyda laricis (Giraud, 1861)[Japanese name: Kiberi-hiratahabachi]

Acantholyda iwatai Takeuchi, 1938
[Japanese name: Iwata-hiratahabachi]

Distribution. Japan(Honshu,
Shikoku,
Kyushu).
Material examined. 1/ and 2? from Fujirindo, 1? from Masutomi-kosen [13], and 1/

Distribution. Europe to Siberia and China.
Japan (central Honshu).
Material examined. 1/ from Hoo-goya [8],Mt. Hoo-zan, recorded by Shinohara (1997).
Host plants. Larix spp. (Shinohara, 1997).
Remarks. This larch-feeding species is rarePamphiliid Sawflies from Yamanashi Prefecture

in Japan, known so far only from five specimens(3/ , 2?) from Nagano Prefecture and one
female from Yamanashi Prefecture, central
Honshu.
Acantholyda mizunoi Shinohara, 2001
[New Japanese name: O-tsuga-hiratahabachi]

Distribution. Japan (central Honshu).
Material examined. 1? from Masutomikosen [13] and 1/ from Mt. Misaka-yama [42],both paratypes.
Host plant. Tsuga diversifolia (Maxim.)
Masters (see Shinohara, 2001a).
Remarks. This species was described on thebasis of seven specimens from central Honshu(Tochigi, Yamanashi and Nagano Prefectures),including two paratypes from Yamanashi Prefecture (Shinohara, 2001a).
Acantholyda nipponica Yano and Sato, 1928
[Japanese name: Nihon-akazu-hiratahabachi]

Distribution. Japan (Hokkaido, northern tocentral Honshu).
Material examined. 1/ from Tokusa-toge[18] recorded by Shinohara (1997).
Host plants. Larix leptolepis (Sieb. et Zucc.)
Gord., Pinus strobus L., Pinus pumila (Pallas)
Regel (see Shinohara, 2001a).
Remarks. This species is known to occur in
Hokkaido and Aomori, Miyagi, Tochigi, Saitama, Tokyo, Yamanashi, Nagano, and Ishikawa
Prefectures (Shinohara, 2001a; Nakamura, 2003).
From Yamanashi Prefecture, this species was firstrecorded by Shinohara (1997). This is usually an
uncommon species but it occurred in larch plantations of Hokkaido in outbreak proportions in
the last decade.
Acantholyda sasakii (Yano, 1916)[New Japanese name: Haraaka-matsu-hiratahabachi]

Distribution. Japan (Honshu).
Material examined. 1/ from Sasago-toge[29], 1/ from Saruhashi [36], and 1/ from73

Aokigahara [48] recorded by Takeuchi (1938)
and Shinohara (1995 b). Additional material. 1/ ,Mt. Tenjo-yama [46], Kawaguchi-ko, 22. V.
1979, H. Suda (HSC); 1?, Koyodai [49], 3. V.
2002, H. Takahashi (NSMT).
Host plants. Pinus densiflora Sieb. et Zucc.,
Pinus parviflora var. pentaphylla (Mayr) Henry,
Pinus koraiensis Sieb. et Zucc. (see Shinohara,
2001 a).
Remarks. This species is known from
Aomori, Yamagata, Miyagi, Fukushima, Tochigi,
Gunma, Ibaraki, Saitama, Tokyo, Kanagawa,
Yamanashi, Nagano, Ishikawa, and Tottori Prefectures (Shinohara, 1995b, 2001a). Takeuchi(1938) and Shinohara (1995b) recorded thisspecies from Yamanashi Prefecture.
Acantholyda tsuyukii Shinohara, 2001
[New Japanese name: Tsuga-hiratahabachi]

Distribution. Japan (central Honshu, Shikoku).
Material examined. 1/ and 1? from Kitazawa [5], Mt. Kitadake and 1/ and 1? from Mt.
Senjodake [6], South Alps, 1/ from Okanbazawa[7], 1/ from Gozaishi [9]/Hoo-goya [8], 1/ from
Gozaishi-kosen [9], and 2/ from Hatchodaira[15], Mt. Kimpu-zan, all paratypes.
Host plant. Tsuga diversifolia (Maxim.)
Masters (see Shinohara, 2001a).
Remarks. This species was described on
the basis of the material from central Honshu(Tochigi, Yamanashi, Nagano and Gifu Prefectures) and Shikoku (Ehime Prefecture) (Shinohara, 2001a).
Cephalcia koebelei (Rohwer, 1910)[Japanese name: Karamatsu-hiratahabachi]

Distribution. Japan (Hokkaido, Honshu),
Russia (Baikal region).
Material examined. 1/ , 14? from Mt. Hoozan recorded by Shinohara (1997). Additional
material. 1?, Mitsutoge-guchi [43], Kawaguchiko Town, 27. VIII. 1986, Y., T. and H. Suda
(HSC).
Host plant. Larix leptolepis (Sieb. et Zucc.)

74

Akihiko Shinohara, Hirohisa Suda and Hideo Takahashi

Gord. (see Shinohara, 1997).
Remarks. Yano (1919, 1920) reported on themass occurrence of “Cephaleia kaebelei [sic]Rohwer” in larch forests in Yamanashi Prefecture. This old record may actually refer to theforegoing species, C. lariciphila japonica (seeShinohara, 1997). In Honshu, this species has
been recorded from Niigata, Yamanashi, andNagano Prefectures (Shinohara, 1997).
Cephalcia lariciphila japonica Shinohara, 1997
[Japanese name: Nihon-karamatsu-hiratahabachi]

Distribution. Japan (Hokkaido, Honshu).
Material examined. 2/ and 1? from Mt.
Hoo-zan [8] recorded by Shinohara (1997). Additional material. 1/ , Mt. Mitsutoge-yama [44],Kawaguchi-ko Town, 22. V. 1990, H. Suda
(HSC); 1?, Mt. Tenjo-yama [46], Kawaguchi-ko
Town, 10. VI. 1986, H. Suda (HSC); 1/ 2?, Shibokusa–Toriiji Pass [51], 31. V. 1988, H. Suda
(HSC).
Host plant. Larix leptolepis (Sieb. et Zucc.)
Gord. (see Shinohara, 1997).
Remarks. In Honshu, this species is known
to occur in Tochigi, Yamanashi, and Nagano Prefectures (Shinohara, 1997). In artificial larch
forests, C. lariciphila japonica and C. koebelei
sometimes occur in great numbers and are regarded as pests.
Cephalcia sp.
Material examined. 1/ , Kagosaka Pass [55],
Yamanaka-ko Village, 25. V. 1972, T. and H.
Suda (HSC).
Remarks. Japanese species of the genus
Cephalcia are badly in need of revision, and thisfemale specimen has not been identified withcertainty.
Onycholyda decorata Shinohara, 1985
[Japanese name: Wamon-hiratahabachi]

Distribution. Japan (Hokkaido, Honshu).
Specimens examined. 1/ , Tokusa Spa, Suta-

ma Town, 18. VII. 1968, H. Suda (HSC); 1/ ,Oshino [39], Saruhashi Town, Otsuki City, 10. V.
1977, H. Suda (HSC); 2/ , Tateno [40], Yanagawa Town, Otsuki City, 23. V. 1975, H. Suda
(HSC); 1/ , Koyodai [49], 11. VIII. 1993, H.
Takahashi (NSMT); 1/ , Minobu [56], Minobu
Town, 18. V. 1964, H. Suda (HSC).
Host plant. Agrimonia pilosa Ledeb. var.
japonica (Miq.) Nakai (see Shinohara, 2006).
Remarks. In Honshu, O. decorata has beenrecorded from Miyagi, Fukushima, Tochigi,
Saitama, Chiba, Tokyo, Kanagawa, Ishikawa,
Nagano, Osaka, and Hyogo Prefectures (Shinohara, 1985c), and this is the first record from Yamanashi Prefecture.
Onycholyda esakii (Takeuchi, 1937)[Japanese name: Esaki-hiratahabachi]

Distribution. Japan(Honshu,
Shikoku,
Kyushu).
Material examined. 1/ from Yanagisawatoge [23] and 1/ from Fuji-rindo recorded by
Shinohara (1986a). Additional material. 1/ ,Koyodai [49]–Sankodai, 3. IX. 1992, H. Takahashi (NSMT).
Host plant. Unknown.
Remarks. In Honshu, this species is known
to occur in Aomori, Iwate, Yamagata, Fukushima, Tochigi, Tokyo, Kanagawa, Fukui, Yamanashi, Nagano, Gifu, Aichi, Nara, Hyogo, andHiroshima Prefectures (Shinohara, 1986a; Enoki
and Nakamura, 1993; Nagase, 2004). From Yamanashi Prefecture, it was first recorded by Shinohara (1986a).
Onycholyda lucida (Rohwer, 1910)[Japanese name: Tsuya-hiratahabachi]

Distribution. Japan(Honshu,
Shikoku,
Kyushu).
Material examined. 1? from Masutomikosen [13] recorded by Shinohara (1986b). Additional material. 6/ 4?, Utsukushinomori [1],1500–1700 m, Yatsugatake Mts., 5–8, VI. 2000,
A. Shinohara (NSMT); 1?, Kiyosato [2], Takane,Pamphiliid Sawflies from Yamanashi Prefecture

Kitakoma, 20. V. 1980, H. Suda (HSC); 1/ , Masutomi [13], 8. V. 1994, H. Hamaji (NSMT); 1/ ,Kanayama-kogen [14], Sutama Town, 28. V.
1979, H. Suda (HSC); 2/ , Tokusa-toge [18],Kurohira, 19–20. V. 1987, H. Yamazaki (NSMT);
3/ , Kitahara [19], Makioka Town, 24. V. 1976,
H. Suda (HSC); 1/ , Nishizawa-keikoku [21],Mitomi Village, 28. V. 1980, H. Suda (HSC); 1/ ,Sakeishi [24], Enzan, 23–24. V. 1987, H. Yamazaki (NSMT); 1/ , Daibosatsu-toge [25], 19–
22. V. 1961, H. Takenaka (NSMT); 1/ , same locality, 27–28. VII. 1981, T. Niisato (NSMT); 1/ ,Daibosatsu road, Enzan City, 18. V. 1966, T. and
H. Suda (HSC); 1/ , Hikawa-rindo [26], Daibosatsu, 9. VI. 1986, Y. Kurosawa (NSMT); 1/ ,Hikawa Path [26], Daibosatsu, 14. VI. 1992, S.
Tsuyuki (NSMT); 2/ , Hikage [28], Yamato Village, 16. V. 1978, H. Suda (HSC); 1/ , Yoshikubo[30], Sasago Town, Otsuki City, 17. V. 1988, H.
Suda (HSC); 1/ , Yunosawa-guchi [31], Otsuki
City, 21. V. 1998, H. Suda (HSC); 4/ , Magi [33],Otsuki Town–Okuyama [35], Nigioka Town, Otsuki City, 11. V. 1976, H. S Suda (HSC); 2/ ,same locality, 2. V. 1980, H. Suda (HSC); 3/ ,Ogatayama [34], Tsuru City, 26. IV. 1988, H.
Suda (HSC); 3?, Ogisan [38], 9. IV. 1990, H.
Takahashi (NSMT); 3/ , Torisawa [39], Otsuki
City, 7. V. 1974, H. Suda (HSC); 2/ 1?, same locality, 23. IV. 1979, H. Suda (HSC); 2/ , Oshino[39], Saruhashi Town, Otsuki City, 10. V. 1977,
H. Suda (HSC); 4/ , Kawai [41], Uenohara, Kitatsuru District, 24. IV. 1990, H. Suda (HSC);
5/ , same locality, 16. IV. 1991, H. Suda (HSC);
1/ 2?, Mt. Mitsutoge [44], Kawaguchi-ko Town,
22. V. 1990, H. Suda (HSC); 1?, Tenjo-san [46],Kawaguchi-ko, 4. V. 1993, H. Takahashi (NSMT);
1/ , same locality, 28. IV. 1987, T. and H. Suda
(HSC); 1/ , Misaka road, Kawaguchiko Town,
22. V. 1990, H. Suda (HSC); 1/ , Panoramadai[47], Kamikuishiki, 22. V. 1978, H. Suda (HSC);
1/ , Koyo-dai [49], Narusawa-mura, 6. V. 1991,
H. Takahashi (NSMT); 4/ 5?, Oshino-mura[52], 1. V. 1990, H. Nagase (HNC).
Host plants. Rubus spp. (Okutani and Fujita,
1955).
Remarks. Onycholyda lucida is widespread75

in Honshu from Aomori to Hiroshima Prefectures (Shinohara, 1986b; Nakamura and Enoki,1997). Shinohara (1986b) was the first to record
the species from Yamanashi Prefecture.
Onycholyda minomalis (Takeuchi, 1930)[Japanese name: Komon-hiratahabachi]

Distribution. Japan (Hokkaido, Honshu, Shikoku, Kyushu).
Specimens examined. 1/ 1? from Kawaguchiko [45] recorded by Shinohara (1987). Additional
material. 2?, Kamisasao [3], Kobuchizawa, 19.
V. 1980, H. Suda (HSC); 2?, Kitahara [19],Makioka Town, 24. V. 1976, H. Suda (HSC); 2/ ,Nishizawa-keikoku [21], Mitomi Village, 28. V.
1980, H. Suda (HSC); 1?, Kuwasai [32], Otsuki
City, 21. V. 1998, H. Suda (HSC); 1?, Koyodai[49]–Ashiwada-yama [50], 21. V. 1978, H. Suda
(HSC); 1/ , Oshino-mura [52], 6. VI. 1996, H.
Nagase (HNC); 1/ , Kagosaka Pass [55], Yamanaka-ko Village, 25. V. 1972, T. and H. Suda
(HSC).
Host plant. Rubus parvifolius L. (see Okutani, 1967).
Remarks. In Honshu, this species has beenrecorded from Miyagi, Saitama, Tokyo, Kanagawa, Niigata, Fukui, Ishikawa, Yamanashi,
Nagano, Gifu, Mie, Kyoto, Osaka, Hyogo, Tottoriand Hiroshima Prefectures (Shinohara, 1987;
Taguchi, 1988). It was first recorded from Yamanashi Prefecture by Shinohara (1987).
Onycholyda similis Shinohara, 1987
[Japanese name: O-komon- hiratahabachi]

Distribution. Japan (Honshu, Kyushu).
Specimens examined. 1/ from Kawaguchiko [45] and 1/ from Yanagawa [40] (bothparatypes) recorded by Shinohara (1987). Additional material. 1?, Yoshikubo [30], Sasago, Otsuki, 17. V. 1988, H. Suda (HSC); 2/ , Yunosawa-guchi [31], Otsuki City, 21. V. 1988, H.
Suda (HSC); 1?, Kuwasai [32], Otsuki City, 21.
V. 1998, H. Suda (HSC); 1?, Tateno [40], Yanagawa Town, Otsuki City, 23. V. 1975, H. Suda

76

Akihiko Shinohara, Hirohisa Suda and Hideo Takahashi(HSC); 2/ 3?, Kawai [41], Uenohara Town, 14.
V. 1991, H. Suda (HSC); 1/ , Toriiji Pass[51]–Ohasumi, Fujiyoshida City, 31. V. 1988, H.
Suda (HSC); 1/ , Kagosaka Pass [55], Yamanaka-ko Village, 25. V. 1972, T. and H. Suda
(HSC).
Host plant. Unknown.
Remarks. This species was described on thebasis of the specimens from Akita, Gunma,
Tokyo, Kanagawa, Niigata, Fukui, Ishikawa, Yamanashi, Nagano, Gifu, Shizuoka, Shiga, Kyoto,and Hyogo Prefectures (Shinohara, 1987), andwas later recorded from Tochigi Prefecture(Nakamura, 2003).
Onycholyda tenuis (Takeuchi, 1937)[Japanese name: Hoso-haraaka-hiratahabachi]

Distribution. Japan (Honshu).
Specimens examined. 1?, Sensui-toge [4],South Alps, 9. VIII. 1985, A. Shinohara(NSMT); 1/ , Masutomi-kosen [13], Sutama, 6.
VIII. 1986, A. Shinohara (NSMT); 1/ , Tokusa
Pass [18], Sutama Town, 18. VII. 1968, H. Suda
(HSC); 1/ , Marukawa Pass–Sakeishi [24],Enzan, 18. VII. 1978, H. Suda (HSC); 1/ , Yunosawa-guchi [31], Otsuki City, 21. V. 1998, H.
Suda (HSC); 1/ , Kuwasai [32], Otsuki City, 21.
V. 1998, H. Suda (HSC).
Host plant. Rubus microphyllus L. f. var.
subcrataegifolius (H. Lev. et Vaniot) Ohwi (seeShinohara, 1987).
Remarks. This species is known to occur onhigher mountains in Tochigi, Gunma, Saitama,
Niigata, Toyama, Ishikawa, Nagano, Gifu, andHyogo Prefectures (Shinohara, 1985e). The specimens listed above represents the first distribution
record from Yamanashi Prefecture. The record
from Hiroshima Prefecture (Taguchi, 1988) may
need confirmation.
Onycholyda viriditibialis (Takeuchi, 1930)[Japanese name: Aosune-hiratahabachi]

Distribution. Japan (Hokkaido, Honshu,
Shikoku, Kyushu), Korea, Russia (Primorskijkraj).
Material examined. 1/ from Yashajin-toge[10] recorded by Shinohara (1986a). Additional
material. 1/ , Utsukushinomori [1], 1500–1700m, Yatsugatake Mts., 5–8. VI. 2000, A. Shinohara(NSMT);
1?1/ ,
Tokuwa-keikoku[22],
Mitomi Village, 12. VIII. 1993, H. Suda (HSC);
1/ , Hikawa-rindo [26], Daibosatsu, 19. VI. 1982,
S. Tsuyuki (NSMT); 1/ 1?, Yunosawa-guchi[31], Otsuki City, 21. V. 1998, H. Suda (HSC);
1?, Saihara [37], Uenohara Town, 11. VI. 1991,
H. Suda (HSC); 1/ , Kawai [41], Uenohara Town,
9. V. 1979, H. Suda (HSC); 1/ , Kawai [41],Uenohara Town, 14. V. 1991, H. Suda (HSC); 1?,
Panoramadai [47], Kamikuishiki Village, 22. VII.
1975, Y., T. and H. Suda (HSC); 1?, Koyodai[49]–Ashiwada-yama [50], 29. VII. 1986, H.
Suda (HSC); 1?, Sankodai–Mt. Ashiwada-yama[50], 3. VIII. 1995, H. Takahashi (NSMT); 1/ ,Mt. Hirao-yama–Hirano [54], 12. VII. 2002, H.
Takahashi (NSMT); 1/ , Kagosaka Pass [55]–Azamidaira, 20. V. 1998, H. Takahashi (NSMT)
Host plant. Rubus crataegifolius Bunge (seeOkutani and Fujita, 1956).
Remarks. This species occurs widely in Honshu from Aomori to Hiroshima Prefectures (Shinohara, 1986 a; Nagase, 2004). It was firstrecorded from Yamanashi Prefecture by Shinohara (1986a).
Pamphilius alnicola Ermolenko, 1973
[Japanese name: Hannoki-shima-hiratahabachi]

Distribution. Japan (Hokkaido, southern
Kuriles, Honshu), Russia (Primorskij kraj).
Material examined. 1/ from Masutomikosen [13] and 1? from Fuji-rindo recorded by
Shinohara (1985 a, 2002).
Host plant. Alnus hirsuta Turcz. (see Shinohara and Hara, 1993).
Remarks. This is an uncommon species
recorded in Honshu only from Tokyo, Nagano,
and Yamanashi Prefectures (Beneš, 1977; Shinohara, 1985a, 2002).

Pamphiliid Sawflies from Yamanashi Prefecture

Pamphilius archiducalis Konow, 1897
[Japanese name: Yumimon-hiratahabachi]

Distribution. Japan (Hokkaido, Honshu),
Shikotan Is., Sakhalin.
Material examined. 1/ , Ochiai–Ichinose[11], 16. VII. 1977, H. Suzuki (NSMT); 1/ , Masutomi Spa [13], 22. VII. 1978, A. Shinohara(NSMT).
Host plants. Alnus hirsuta Turcz., A. matsumurae Call. (see Shinohara and Hara, 1999;
Shinohara, 2005).
Remarks. This species was described on thebasis of a female specimen from “Japan”(Konow, 1897) and later recorded in Honshu
from Ishikawa Prefecture only (Shinohara andOkutani, 1983). We have examined specimens of
this species from Gunma Prefecture (1/,Marunuma, 1,400 m, 8–9. VI. 2001, A. Shinohara, NSMT), Kanagawa Prefecture (1/ , Mikuni-toge, Yamakita, 1,100 m, 1. VI. 2005, H. Nagase, HNC), Nagano Prefecture (1?, Shichimionsen, 3. VI. 1985, A. Shinohara, NSMT; 2/ ,same data except 4–5. 1988, NSMT; 2/ , Yarisawa, Kamikochi, 1,600–1,900 m, 18–22. VII.
1989, A. Shinohara, NSMT; 1/, same data except 30. VII. 1990, NSMT; 1/ , Makuiwa, Shigakogen, 1,550 m, 7–9. VIII. 1996, A. and T. Shinohara, NSMT; 1/ , Nakanoyu, Mt. Ontake,
1,600 m, 21. VII. 1981, H. Hara, NSMT; 2?,
Shimashima-dani, 1,700–1,800 m, 28. VI. 1976,
A. Shinohara, NSMT; 1?, Kamikochi, 24–25.
VII. 1957, R. Ishikawa, NSMT; 1/ , Karuizawa,
12. VII. 1959, R. Ishikawa, NSMT; 1/ ,Akadake-kosen, Yatsugatake Mts., 2. VII. 1972,
A. Shinohara, NSMT; 1/ , Minoto, 1,800–
2,000 m, Yatsugatake Mts., 26. VII. 1980, A. Shinohara, NSMT; 1/ , same data except 7. VII.
1982, NSMT; 1/ , same data except 6. VIII.
1982, NSMT; 1/ , same data except 3. VIII.
1984, NSMT; 7/ , same data except 29. VII.–3.
VIII. 1986, NSMT; 1/1?, same data except 4–8.
VIII. 1987, NSMT; 4/ , same data except 4–8.
VIII. 1988, NSMT; 6/ 2?, same data except
23–26. VII. 1996, NSMT; 2/ , same data except
31. VII.–1. VIII. 1997, NSMT; 3/ , same data ex-77

cept 27–31. VII. 1999, NSMT; 1/ , Niiyamatoge, 1,200 m, 35°46N 138°02E, Ina, 23–24. V.
2003, A. Shinohara, NSMT), and Gifu Prefecture(1?, Takayama, 20. V. 1979, A. Shinohara,
NSMT; 1/ , Oshirakawa, 6. VII. 1979, A.
Shimizu, NSMT).
Pamphilius benesi Shinohara, 1985
[Japanese name: Hashibami-hiratahabachi]

Distribution. Japan (Hokkaido, Honshu,
Shikoku, Kyushu); Korea.
Material examined. 1? from Utsukushinomori [1] and 1/ from Hikawa-rindo [26] listedby Shinohara (1988, 2001 b). Additional material. 1/ , Shibokusa [52]–Toriiji Pass [51], 31. V.
1988, H. Suda (HSC).
Host plant. Corylus sieboldiana Blume (seeShinohara and Hara, 1997).
Remarks. In Honshu, this species has beenrecorded from Aomori, Iwate, Tochigi, Tokyo,
Niigata, Ishikawa, Yamanashi, Nagano, Shizuoka, Kyoto, Hyogo, Okayama, Nara, and Tottori
Prefectures (Shinohara, 2001b; Nakamura, 2003;
Shinohara and Yamada, 2005). We have examined two females of this species collected in
Hiroshima Prefecture (Yokogo, Togouchi Town,
14–16. VI. 2002, S. Tsuyuki, NSMT).
Pamphilius brevicornis ibukii Shinohara, 1995
[Japanese name: O-tsuyazu-hiratahabachi]

Distribution. Japan (Hokkaido, Honshu).
Material examined. 1/ , a paratype, from
Utsukushinomori [1] (Shinohara, 1995 a).
Host plant. Unknown.
Remarks. Only two females, the holotype
from Mt. Hakusan, Ishikawa Prefecture, and aparatype from Utsukushinomori, Yamanashi Prefecture, are known from Honshu (Shinohara,
1995a).
Pamphilius confusus Shinohara, 2005
[Japanese name: Hannoki-hiratahabachi]

Distribution.

Japan(Hokkaido,

Honshu,

78

Akihiko Shinohara, Hirohisa Suda and Hideo Takahashi

Shikoku).
Material examined. 1/ (paratype) from
Hoo-goya [8] (Shinohara, 2005).
Host plant. Alnus maximowiczii Call. (seeShinohara, 2005).
Remarks. This species is usually found on
higher mountains where the host plant grows. In
Honshu, it has been recorded from Aomori,
Miyagi, Fukushima, Tochigi, Gunma, Niigata,
Tokyo, Yamanashi, Nagano, and Ishikawa Prefectures (Shinohara, 2005).
Pamphilius flavipectus Shinohara, 2005
[Japanese name: Kimune-hiratahabachi]

Distribution. Japan(Honshu,
Shikoku,
Kyushu).
Material examined. 17 paratypes listed by
Shinohara (2005): 1/ 8? from Utsukushinomori[1], 2/ from Kiyosato [2], 1/ from Hoo-goya[8], 1/ 1? from Shinshu-toge [12], and 1/ 2?
from Masutomi-kosen [13].
Host plants Alnus hirsuta Turcz., A. matsumurae Call. (see Shinohara and Kojima, 2006).
Remarks. Shinohara (2005) described thisspecies on the basis of the material from Aomori,
Iwate, Yamagata, Fukushima, Tochigi, Gunma,
Niigata, Yamanashi, Nagano, and Yamaguchi
Prefectures in Honshu, besides the material from
Shikoku and Kyushu. Yoshida (2006) recorded it
also from Osaka Prefecture.
Pamphilius gracilis Shinohara, 1985
[Japanese name: Nanakamado-hiratahabachi]

Distribution. Japan (Hokkaido, Honshu),
Korea.
Material examined. 1? from Utsukushinomori [1] recorded by Shinohara (2001b).
Host plant. Sorbus commixta Hedl. (see Shinohara and Hara, 1992).
Remarks. This species is known to occur
only in Nagano, Niigata, Yamanashi, and Tottori
Prefectures in Honshu (Shinohara, 2001b).

Pamphilius hilaris (Eversmann, 1847)[Japanese name: Bara-hiratahabachi]

Distribution. Russia (Urals to Kamchatka
and to Primorskij kraj), Japan (Honshu), China(Shaanxi).
Material examined. 38/ , 25? from Kawaguchi-ko [45] and 1? from Aokigahara [48]recorded by Shinohara (2002). Additional material. 1/ , Hirano–Mt. Hirao [54], 18. V. 2001, H.
Takahashi (NSMT).
Host plant. Rosa multiflora Thunb. (see Shinohara and Okutani, 1983).
Remarks. This species is uncommon in
Japan, recorded only from Tochigi, Gunma,
Saitama, Tokyo, Yamanashi, and Nagano Prefectures (Shinohara, 2002; Nakamura, 2004). Besides Saitama and Yamanashi Prefectures, only
one or a few specimens were collected in eachprefecture. Close to the northeastern coast of
Kawaguchi-ko Lake, there was a large open fieldcovered with bushes of Rosa multiflora, where P.
hilaris was abundant, when Shinohara visited
there during the end of the 1970s.
Pamphilius ishikawai Shinohara, 1979
[Japanese name: Tsuyakuro-hiratahabachi]

Distribution. Japan (Honshu).
Material examined. 1/ 1? (paratypes) from
Mitsutoge [44] cited by Shinohara (1979). Additional material. 1?, Kanayama-kogen [14], Sutama Town, 28. V. 1979, H. Suda (HSC); 1/ ,Tateno [40], Yanagawa Town, Otsuki City, 23. V.
1975, H. Suda (HSC); 1/ , Oshino-mura [52], 6.
VI. 1996, H. Nagase (HNC).
Host plant. Unknown.
Remarks. This is not a common species but
is fairly wide spread in Honshu. It is known tooccur in Aomori, Akita, Fukushima, Tochigi,
Gunma, Saitama, Tokyo, Kanagawa, Niigata,
Toyama, Ishikawa, Fukui, Yamanashi, Nagano,
Aichi, Hyogo, and Tottori Prefectures (Shinohara, 2002; Shinohara and Yamada, 2005).

Pamphiliid Sawflies from Yamanashi Prefecture

Pamphilius itoi Shinohara, 1985
[Japanese name: Hanno-haraaka-hiratahabachi]

Distribution. Japan (Hokkaido, southern
Kuriles, Honshu).
Material examined. 1/ from Masutomikosen [13] and 1/ from Yanagisawa-toge [23],both paratypes, treated by Shinohara (1985 d).
Additional material. 2/ 2?, Utsukushinomori[1], Yatsugatake Mts., 1,500–1,700, 5–8. VI.
2000, A. Shinohara (NSMT); 1/ , Masutomikosen [13], 6. VIII. 1986, A. Shinohara (NSMT);
2?, Kanayama-kogen [14], Sutama Town, 28. V.
1979, H. Suda (HSC); 1/ , Yunosawa-guchi [31],Otsuki City, 21. V. 1998, H. Suda (HSC); 1/ , Mt.
Tenjo-yama [46], Kawaguchi-ko Town, 10. VI.
1986, H. Suda (HSC).
Host plant. Alnus hirsuta Turcz. (see Shinohara and Hara, 1995).
Remarks. This species has been recorded
from Aomori, Iwate, Tochigi, Tokyo, Kanagawa,Niigata, Ishikawa, Fukui, Yamanashi, Nagano,Gifu, and Hyogo Prefectures (Shinohara, 1985d;
Naito et al., 2004; Shinohara and Yamada,2005).
Pamphilius kamikochensis Takeuchi, 1930
[Japanese name: Ko-haraaka-hiratahabachi]

Distribution. Japan (Hokkaido, Honshu).
Material examined. 1? from Masutomi [13]recorded by Shinohara (1998).
Host plant. Prunus ssiori Fr. Schm. (see Shinohara and Okutani, 1983).
Remarks. In Honshu, this species has beenrecorded from Aomori, Tochigi, Nagano, and Yamanashi Prefectures (Shinohara, 1998).
Pamphilius komonensis Takeuchi, 1930
[Japanese name: Kaede-hiratahabachi]

Distribution. Japan(Honshu,
Shikoku,
Kyushu).
Material examined. A series of specimensfrom Kanayama-kogen [14], Nishizawa-keikoku[21], Yoshikubo [30], Torisawa [39], Yanagawa

79[40], Kawai [41], and Koyodai [49], all listed by
Shinohara and Zhou (2006).
Host plant. Acer mono Maxim. (see Shinohara and Okutani, 1983).
Remarks. Shinohara and Zhou (2006) fully
discussed this and allied species. Pamphilius
komonensis is rather common in lower mountains
in Honshu.
Pamphilius lobatus Maa, 1950
[Japanese name: O-haraaka-hiratahabachi]

Distribution. Japan (Hokkaido, Honshu),
Korea, China (Jilin), Russia (Primorskij kraj).
Material examined. 1?, Kanayama-kogen[14], Sutama Town, 28. V. 1979, H. Suda (HSC);
1/ , Mt. Kimpu-zan [17], 1. VII. 1963, T. Naito(KU); 1?, Mitsutoge [44], 30. IV. 1974, K.
Kimura (NSMT); 1/ , Azamidaira [55], 17. VI.
1990, H. Takahashi (NSMT).
Host plant. Unknown.
Remarks. In Honshu, this species has beenrecorded from Aomori, Akita, Miyagi, Tochigi,
Kanagawa, and Hyogo Prefectures, besides thetype locality, Kirizumi, Gunma Prefecture (Maa,
1950; Nakamura, 2003; Nagase, 2004; Naito etal., 2004; Shinohara and Yamada, 2005). This is
the first distribution record from Yamanashi Prefecture. We have also examined a female specimen from Fukushima Prefecture (Fudosawabashi–Takayu-onsen, 27. VI. 1993, H. Takahashi,
NSMT).
Pamphilius nakagawai Takeuchi, 1930
[Japanese name: Nakagawa-hiratahabachi]

Distribution. Japan(Honshu,
Shikoku,
Kyushu).
Material examined. Long series of specimens from Utsukushinomori [1], Shinshu-toge[12], Masutomi-kosen [13], Biwakubozawa [16],Otome-onsen [20], Tenmoku [27], Saruhashi[36], Tenjo-yama [46], Koyodai [49], all listedby Shinohara (2005). Additional material. 2/ 1?,
Kanayama-kogen [14], Sutama Town, 28. V.
1979, H. Suda (HSC); 1/ , Yunosawa-guchi [31],80

Akihiko Shinohara, Hirohisa Suda and Hideo Takahashi

Otsuki City, 21. V. 1998, H. Suda (HSC); 1?, Mt.
Mitsutoge [44], Kawaguchi-ko Town, 22. V.
1990, H. Suda (HSC); 1/ 3?, Tenjo-yama[46], Kawaguchi-ko, 4. V. 1992, H. Takahashi(NSMT); 16?, Koyodai [49]–Ashiwada-yama[50], 21. V. 1978, H. Suda (HSC); 3?, same locality, 7. V. 1980, H. Suda (HSC); 1?, Ode-yama[53], Yamanaka-ko Village, 23. V. 1980, H. Suda
(HSC); 1/ , Kagosaka Pass [55], Yamanaka-ko
Village, 25. V. 1972, T. and H. Suda (HSC).
Host plant. Alnus hirsuta Turcz., A. matsumurae Call. (see Ishii, 1952; Shinohara, 2005).
Remarks. In Honshu, this species is known
to occur in Aomori, Yamagata, Miyagi, Fukushima, Tochigi, Gunma, Tokyo, Kanagawa, Yamanashi, Nagano, Shizuoka, Gifu, Toyama,
Ishikawa, and Fukui Prefectures (Shinohara,2005). This species is sometimes found commonly on or around the young leaves of the hostplants.
Pamphilius naokoae Shinohara, 1999
[New Japanese name: Hime-kurozu-hiratahabachi]

Distribution. Japan (Honshu).
Material examined. Holotype (/ ) andparatypes (2/ , 21?), all from the type locality,
Masutomi-kosen [13].
Host plant. Unknown.
Remarks. This species was described on thebasis of a series of specimens collected at Masutomi-kosen, Yamanashi Prefecture, in 1978–1979(Shinohara, 1999). So far as we know, these are
the only specimens ever collected. The type series was swept from the foliage of Alnus hirsuta
stands along the stream at the type locality. The
vegetation of this site was completely destroyed
by a flood caused by a typhoon a few years
later.
Pamphilius takeuchii Beneš, 1972
[Japanese name: Usumon-hiratahabachi]

Distribution. Japan (Hokkaido, Honshu,
Shikoku, Kyushu).
Material examined. 4/ 1? from Yashajin[10], Saihara [37], Narusawa, Koyodai [49], andSankodai–Mt. Ashiwada-yama [50], listed by
Shinohara and Zhou (2006).
Host plant. Acer mono Maxim. (see Shinohara and Okutani, 1983).
Remarks. This species is widely distributedin Japan from lowlands to high mountains. See
Shinohara and Zhou (2006) for more discussion.
Pamphilius tricolor Beneš, 1974
[Japanese name: Miiro-hiratahabachi]

Distribution. Japan (Hokkaido, Honshu),
Ural to the Russian Far East, Korea.
Material examined. 1/ , Kitahara [19],Makioka Town, 24. V. 1976, H. Suda (NSMT).
Host plant. Unknown.
Remarks. In Honshu, this species is very
rare, so far known only from one male specimen
from Niigata Prefecture (Shinohara, 1995a). The
first female specimen from Honshu is not distinguishable from the females collected in Hokkaido, Korea and Primorskij kraj.
Pamphilius venustus (Smith, 1874)[Japanese name: Haraaka-hiratahabachi]

Distribution. Japan (Hokkaido, Honshu,
Shikoku).
Material examined. 1/ , Okanbazawa [7],Mt. Kitadake, 12. VII. 1979, N. Takeuchi(NSMT); 1/ , Okanbazawa [7], 15. VII. 1978, H.
Kumamoto (HKC); 1?, Shinshu-toge [12], 4. VI.
1984, A. Shinohara (NSMT).
Host plant. Filipendula camtschatica (Pall.)Maxim. (see Shinohara and Okutani, 1983)
Remarks. Shinohara and Morita (2003) already recorded this species from Yamanashi Prefecture but did not give any collection data. In
Honshu, this species is known to occur in
Aomori, Iwate, Tochigi, Niigata, Yamanashi,
Nagano, Gifu, Ishikawa, and Shiga Prefectures(Shinohara and Yamada, 2005).

Pamphiliid Sawflies from Yamanashi Prefecture

Pamphilius volatilis (Smith, 1874)[Japanese name: Shima-hiratahabachi]

Distribution. Japan (Hokkaido, Honshu,
Kyushu), Korea, Russia (Primorskij Kraj).
Material examined. 1/ , Utsukushinomori[1], Yatsugatake Mts., 1,500–1,700 m, 19. VI.
1999, A. Shinohara (NSMT); 2?, same locality,
5–8. VI. 2000, A. Shinohara (NSMT); 9?, Shinshu-toge [12], 4. VI. 1984, A. Shinohara(NSMT); 2?, Kanayama-kogen [14], Sutama
Town, 28. V. 1979, H. Suda (HSC); 2/ 1?,
Tokusa-toge [18], Kurohira, 19–20. V. 1987, H.
Yamazaki (NSMT); 2/ , Ogatayama [34], Tsuru
City, 26. IV. 1988, H. Suda (HSC); 1/ , Tateno[40], Yanagawa Town, Otsuki City, 23. V. 1975,
H. Suda (HSC); 1/ , Oshino [39], Saruhashi
Town, Otsuki City, 10. V. 1977, H. Suda (HSC);
1/ , Kawai [41], Uenohara Town, 9. V. 1979, H.
Suda (HSC); 1?, same locality, 16. IV. 1991, H.
Suda (HSC); 1/ , Mt. Tenjo-yama [46],Kawaguchi-ko Town, 22. V. 1979 H. Suda
(HSC); 1?, Koyodai [49], Narusawa-mura, 3. V.
1992, H. Takahashi (NSMT).
Host plants. Prunus spp., Crataegus
chlorosarca Maxim. (see Shinohara and Hara,
1999).
Remarks. This widely distributed species isoften commonly found in central Honshu but it
has not been recorded from Yamanashi Prefecture. It has been recorded from Aomori, Miyagi,
Tochigi, Gunma, Saitama, Tokyo, Kanagawa,Toyama, Nagano, Ishikawa, Fukui, Gifu, Mie,Nara, Osaka, Hyogo, Okayama, Hiroshima, Yamaguchi Prefectures in Honshu (Tanaka, 1979;
Shinohara, 1985a, 2002; Miyoshi, 1988; Nakamura and Enoki, 1997).
We have examined specimens of this species
also from Fukushima Prefecture (1?, Tsuchiyuonsen, Fukushima-shi, 29. V. 1993, H. Takahashi,
NSMT), Chiba Prefecture (1?, Matsudo, NSMT;
2/ , Mabashi, 24. IV. 1938, J. Yoshioka, NSMT;
1/ , Naritasan Park, Narita City, 18. IV. 1979, H.
Suda, HSC; 1?, Shibayama, Sanbu District, 18.
IV. 1973, H. Suda, HSC; 2?, Koichibe, Kimitsu
City, 24. IV. 1996, H. Suda, HSC), Niigata Pre-81

fecture (1?, Mantaro-dani, Tsuchitaru, 6. VI.
1984, A. Shinohara, NSMT), Shiga Prefecture(3/ 7?, Azai-cho, 8. V. 1984, A. Shinohara,
NSMT), Kyoto Prefecture (2?, Kyoto, 15. V.
1916, NSMT), and Tottori Prefecture (7?,
Yokotemichi, W. slope of Mt. Daisen, 1,000 m,
20–25. V. 2000, A. Shinohara, NSMT).
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