Bulletin of the Geological Survey of Japan, vol. 70 (1/2), p. 137–161, 2019

Notes and Comments

Radiolarian assemblages from the lower to middle Miocene
at IODP Site U1335 in the eastern equatorial PacificShin-ichi Kamikuri1,*
Shin-ichi Kamikuri (2019) Radiolarian assemblages from the lower to middle Miocene at IODP Site U1335
in the eastern equatorial Pacific. Bull. Geol. Surv. Japan, vol. 70 (1/2), p. 137–161, 1 figure, 1 table, 18 plates.
Abstract: Totally 183 species or species group of radiolarians including 78 spumellarians and 105
nassellarians were identified from the lower to middle Miocene at IODP Site U1335 in the easternequatorial Pacific. Photographs have been illustrated in 18 plates. These taxa include 70 unidentified
forms, particularly within the families Actinommidae, Collosphaeridae, Hexalonchidae, Litheliidae,
Pyloniidae, Spongodiscidae and Stylodictyidae. Some of these taxa might be new species.
Keywords: radiolaria, Miocene, equatorial Pacific

1. Introduction
The eastern equatorial Pacific is one of the major
upwelling areas and supports a significant proportion of
the new biological production (Chavez and Barber, 1987).
The band of cool surface water from about 3°S to 3°N iscalled the equatorial “cold tongue” (Fiedler and Talley,
2006; Kessler, 2006; Pennington et al., 2006). Hence thisarea makes global contribution to biogenic sediment burial
due to equatorial upwelling.
Integrated Ocean Drilling Program (IODP) Expedition
320, “Pacific Equatorial Age Transect (PEAT)”, wasdesigned to recover continuous well-preserved sedimentsof the eastern equatorial Pacific (Pälike et al., 2010).
The primary object was to reconstruct the evolution of
the equatorial climate system throughout the Cenozoic.
Pälike et al. (2012) presented a record of the carbonate
compensation depth (CCD) that covers the past 53 million
years from a depth transect in the eastern equatorial
Pacific, and found large superimposed fluctuations in the
CCD during the middle Eocene.
One of the other aims of this program was to provide
information about rapid biological evolution and turnoverrates during times of climatic stress (Pälike et al., 2010).
IODP Exp. 320 Site 1335 focused on the early andmiddle Miocene. This interval of the time is noted for its
significantly heavy benthic oxygen isotopes, indicating
long-term global cooling (Zachos et al., 2001). The earlyand middle Miocene also marks the beginning of thedeposition of the abundant siliceous microfossil in theeastern equatorial Pacific (Keller and Barron, 1983).
Species components and the faunal turnover ofradiolarians, which is one of siliceous microfossils, havebeen well documented from the Quaternary to late Neogene
in the eastern equatorial Pacific (e.g. Molina-Cruz, 1977;
Moore et al., 1981; Haslett and Funnell, 1996; Kamikuri
et al., 2009). However that of radiolarian assemblages
during the early and middle Miocene are still unknown.
The purpose of this study is to illustrate radiolarianmorphotypes from the lower to middle Miocene at IODP
Site U1335 in order to record the species components inthe eastern equatorial Pacific.

2. Material and methods
Site U1335 was drilled at a water depth of 4,327.5 m(5°18.735’N, 126°17.002’W) in the eastern tropical
Pacific (Fig. 1). The position of this site was estimated to
40°

20°
U1335
0°

20°S

Equatorial upwelling zone

150°W

120°

90°

Fig. 1 Location map of IODP Site U1335 in the easternequatorial Pacific.

1

Faculty of Education, Ibaraki University, Bunkyo 2-1-1, Ibaraki 310-8512, Japan*
Corresponding author: S. Kamikuri, Email: shin-ichi.kamikuri.
@vc.ibaraki.ac.jp− 137 −

Fig. 1

Bulletin of the Geological Survey of Japan, vol. 70 (1/2), 2019

have ranged in latitude from about 0°to 2°
N from the early
to middle Miocene (Pälike et al., 2010). The lithology
from the lower to middle Miocene of this site consistsof nannofossil ooze with well-preserved radiolarians(Pälike et al., 2010). A total of 21 sediment samples from
U1335A-10H-CC to 31H-CC (106.0 to 337.1 meterscomposite core depth below seafloor) were examinedin this study. The samples were treated with standard
methods to isolate radiolarian individuals (Sanfilippo
et al., 1985). The sediment samples were treated with
H2O2 and HCl solution. The residues were sieved with63 µm, and covered with a 24 × 40 mm cover glass with

Norland Optical Adhesive #61 as the mounting medium.
The studied interval was divided into four radiolarianbiostratigraphic zones from RN5 to RN2 (Pälike et al.,2010).

3. Radiolarian fauna
The aim of this study is to illustrate radiolarians of the early
to middle Miocene age collected from IODP Site U1335 in
the eastern equatorial Pacific. A total 183 morphotypes ofradiolarians including 78 spumellarians and 105 nassellarians
were encountered (Table 1, Plates 1 to 18). These taxa

Table 1 Explanation of Plates 1 to 18.
Plate no.
Pl.01, fig. 1
Pl.01, fig. 2
Pl.01, fig. 3
Pl.01, fig. 4
Pl.01, fig. 5
Pl.01, fig. 6
Pl.01, fig. 7
Pl.01, fig. 8
Pl.01, fig. 9
Pl.01, fig. 10
Pl.02, fig. 1
Pl.02, fig. 2
Pl.02, fig. 3
Pl.02, fig. 4
Pl.02, fig. 5
Pl.02, fig. 6
Pl.02, fig. 7
Pl.02, fig. 8
Pl.02, fig. 9
Pl.02, fig. 10
Pl.02, fig. 11
Pl.02, fig. 12
Pl.02, fig. 13
Pl.03, fig. 1
Pl.03, fig. 2
Pl.03, fig. 3
Pl.03, fig. 4
Pl.03, fig. 5
Pl.03, fig. 6
Pl.03, fig. 7
Pl.03, fig. 8
Pl.03, fig. 9
Pl.03, fig. 10
Pl.04, fig. 1
Pl.04, fig. 2
Pl.04, fig. 3
Pl.04, fig. 4
Pl.04, fig. 5
Pl.04, fig. 6
Pl.04, fig. 7
Pl.04, fig. 8
Pl.04, fig. 9
Pl.04, fig. 10
Pl.04, fig. 11
Pl.05, fig. 1
Pl.05, fig. 2
Pl.05, fig. 3
Pl.05, fig. 4
Pl.05, fig. 5
Pl.05, fig. 6
Pl.05, fig. 7
Pl.05, fig. 8
Pl.05, fig. 9
Pl.05, fig. 10
Pl.05, fig. 11
Pl.05, fig. 12
Pl.05, fig. 13
Pl.06, fig. 1
Pl.06, fig. 2
Pl.06, fig. 3
Pl.06, fig. 4
Pl.06, fig. 5
Pl.06, fig. 6
Pl.06, fig. 7

Species and species group

Location

Trisolenia combinata Bjørklund and Goll
13H-2, 104-106 cm, D32/4
Collosphaera macropora Popofsky
15H-CC, B26/3
Trisolenia sp. A
15H-CC, K44/0
Siphonosphaera sp. C
13H-2, 104-106 cm, Z20/0
Collosphaera pyloma Reynolds
13H-CC, Q23/0
Collosphaera reynoldsi Kamikuri
10H-CC, H23/1
Trisolenia megalactis costlowi Bjørklund and Goll
13H-2, 104-106 cm, Y39/4
Polysolenia spinosa (Haeckel) group
15H-CC, R37/2
Polysolenia murrayana (Haeckel)
11H-CC, T33/3
Polysolenia spinosa (Haeckel) group
11H-CC, D20/1
Trisolenia megalactis megalactis Ehrenberg
17H-CC, W48/4
Trisolenia megalactis megalactis Ehrenberg
18H-CC, G40/4
Trisolenia megalactis megalactis Ehrenberg
17H-CC, Y48/1
Siphonosphaera sp. A
17H-CC, G41/3
Siphonosphaera sp. A
17H-CC, D22/1
Siphonosphaera sp. D
18H-CC, R10/1
Siphonosphaera sp. B
13H-2, 104-106 cm, P39/2
Collosphaera glebulenta Bjørklund and Goll
19H-CC, V21/1
Collosphaera glebulenta Bjørklund and Goll
13H-CC, R21/3
Collosphaera sp. A
15H-CC, E36/3
Collosphaera sp. A
13H-2, 104-106 cm, Q37/3
Collosphaera sp. A
13H-CC, J15/4
Collosphaera brattstroemi Bjørklund and Goll
10H-CC, S38/4
Axoprunum bispiculum (Popofsky)
11H-CC, K20/0
Axoprunum bispiculum (Popofsky)
11H-CC, V12/1
Stylosphaera communis (Carnevale)
23H-CC, D38/0
Stylosphaera sp. B
13H-CC, P35/0
Amphisphaera santaeannae (Campbell and Clark)
16H-CC, S38/0
Amphistylus polistylus (Carnevale)
16H-CC, X40/0
Amphisphaera neptunus (Haeckel)
17H-CC, U43/3
Stylosphaera sp. A
23H-CC, L46/4
Amphistylus angelinus (Campbell and Clark)
16H-CC, U45/2
Stylosphaera sp. B
13H-CC, F14/1
Actinomma robusta (Kling)
10H-CC, G38/0
Actinomma robusta (Kling)
10H-CC, W20/1
Thecosphaera sp. A
11H-CC, O35/0
Actinomma robusta (Kling)
13H-2, 104-106 cm, E39/4
Thecosphaera sp. B
13H-CC, U36/0
Thecosphaera sp. C
18H-CC, N15/3
Thecosphaera sp. D
18H-CC, H20/3
Thecosphaera sp. E
25H-CC, T34/2
Hexacontium sp. B
20H-CC, U27/3
11H-CC, E21/2
Hexacontium cf. arachnoidale Hollande and Enjumet
Saturnalis circularis Haeckel
11H-CC, W43/1
Cenosphaera sp. A
15H-CC, Q45/4
Heliodiscus sp. B
15H-CC, X21/1
Hexapyle dodecantha Haeckel
15H-CC, L15/2
Excentrodiscus sp. A
15H-CC, R12/0
Cenosphaera sp. B
15H-CC, E12/0
Heliodiscus sp. A
16H-CC, C46/0
Thecosphaera sp. E
15H-CC, N37/0
Hexacontium sp. C
15H-CC, V37/3
Axoprunum sp. A
15H-CC, Q45/4
Didymocyrtis violina (Haeckel)
18H-CC, P33/2
Didymocyrtis tubaria (Haeckel)
17H-CC, T46/4
Didymocyrtis mammifera (Haeckel)
15H-CC, X21/0
Spongurus cylindricus Haeckel
15H-CC, Y39/3
Cladococcus sp. A
16H-CC, L44/2
Hexacontium pachydermum Jørgensen
13H-CC,− 138− Y40/1
Hexalonche sp. A
21H-CC, S23/3
Didymocyrtis prismatica (Haeckel)
25H-CC, D11/2
Didymocyrtis prismatica (Haeckel)
23H-CC, O31/0
Didymocyrtis tubaria (Haeckel)
16H-CC, G16/0
Didymocyrtis tubaria (Haeckel)
16H-CC, K41/2

Zone Reference
RN5 Bjørklund and Goll, 1979
RN5 Popofsky, 1917
RN5
RN5
RN5 Reynolds, 1980
RN5 Kamikuri, 2010
RN5 Bjørklund and Goll, 1979
RN5 Nigrini and Lombari, 1984
RN5 Nigrini and Lombari, 1984
RN5 Nigrini and Moore, 1979
RN4 Bjørklund and Goll, 1979
RN4 Bjørklund and Goll, 1979
RN4 Bjørklund and Goll, 1979
RN4
RN4
RN4
RN5
RN3 Bjørklund and Goll, 1979
RN5 Bjørklund and Goll, 1979
RN5
RN5
RN5
RN5 Bjørklund and Goll, 1979
RN5 Takemura, 1992
RN5 Takemura, 1992
RN2 Sanfilippo et al. , 1978
RN5
RN5 Petrushevskaya and Kozlova, 1972
RN5 Takemura, 1992
RN4 Petrushevskaya and Kozlova, 1972
RN2
RN5 Campbell and Clark, 1944
RN5
RN5 Morley and Nigrini, 1995
RN5 Morley and Nigrini, 1995
RN5
RN5 Morley and Nigrini, 1995
RN5
RN4
RN4
RN2
RN3
RN5
RN5 Nishimura and Yamauchi, 1984
RN5
RN5
RN5 Haeckel, 1887
RN5
RN5
RN5
RN5
RN5
RN5
RN4 Sanfilippo et al. , 1985
RN4 Sanfilippo et al. , 1985
RN5 Sanfilippo et al. , 1985
RN5 Takahashi, 1991
RN5
RN5 Petrushevskaya and Kozlova, 1972
RN3
RN2 Sanfilippo et al. , 1985
RN2 Sanfilippo et al. , 1985
RN5 Sanfilippo et al. , 1985
RN5 Sanfilippo et al. , 1985

Pl.04, fig. 10
Pl.04, fig. 11
Pl.05, fig. 1
Pl.05, fig. 2
Pl.05, fig. 3
Pl.05, fig. 4
Pl.05, fig. 5
Pl.05, fig. 6
Pl.05, fig. 7
Pl.05, fig. 8
Pl.05, fig. 9
Pl.05, fig. 10
Plate
Pl.05, no.
fig. 11
Pl.01,
Pl.05, fig. 112
Pl.01,
Pl.05, fig. 2
13
Pl.01,
Pl.06, fig. 3
1
Pl.01,
Pl.06, fig. 42Pl.01,
Pl.06, fig. 5
3Pl.01,
Pl.06, fig. 6
4Pl.01,
Pl.06, fig. 7
5Pl.01,
Pl.06, fig. 8
6Pl.01,
Pl.06, fig. 9
7Pl.01,
Pl.06, fig. 10
8
Pl.02,
Pl.06, fig. 1
9Pl.02,
Pl.06, fig. 2
10
Pl.02,
Pl.06, fig. 3
11
Pl.02,
Pl.06, fig. 4
12
Pl.02,
Pl.06, fig. 5
13
Pl.02,
Pl.07, fig. 6
1
Pl.02,
Pl.07, fig. 7
2
Pl.02,
Pl.07, fig. 8
3
Plate
Pl.02,
fig. 9Pl.07, no.
4
Pl.07, fig. 10
5
Pl.02,
Pl.07, fig. 11
6
Pl.02,
Pl.07, fig. 12
7
Pl.02,
Pl.07,
Pl.02, fig. 8
13
Pl.07, fig. 1
9Pl.03,
Pl.07, fig. 2
10Pl.03,
Pl.07, fig. 3
11
Pl.03,
Pl.07, fig. 4
12Pl.03,
Pl.07, fig. 5
13
Pl.03,
Pl.07,
Pl.03, fig. 14
6
Pl.07, fig. 7
15
Pl.03,
Pl.07, fig. 8
16Pl.03,
Pl.07, fig. 917
Pl.03,
Pl.07, fig. 10
18
Pl.03,
Pl.08,
Pl.04, fig. 1
Pl.08, fig. 2
Pl.04,
Pl.08, fig. 3
Pl.04,
Pl.08, fig. 4
Pl.04,
Pl.08, fig. 5
Pl.04,
Pl.08, fig. 6
Pl.04,
Pl.08,
Pl.04, fig. 7
Pl.08, fig. 8
Pl.04,
Pl.08, fig. 9
Pl.04,
Pl.08, fig. 10
Pl.04,
Pl.08, fig. 11
Pl.04,
Pl.08, fig. 1
12
Pl.05,
Pl.08, fig. 2
13
Pl.05,
Pl.08, fig. 3
14
Pl.05,
Pl.08, fig. 4
15
Pl.05,
Pl.09, fig. 5
1
Pl.05,
Pl.09, fig. 62Pl.05,
Pl.09,
Pl.05, fig. 37
Pl.09, fig. 8
4Pl.05,
Pl.09, fig. 95Pl.05,
Pl.09, fig. 10
6
Pl.05,
Pl.09, fig. 11
7
Pl.05,
Pl.09,
Pl.05, fig. 812
Pl.09,
Pl.05, fig. 9
13
Pl.09, fig. 1
10
Pl.06,
Pl.09, fig. 2
11
Pl.06,
Pl.09, fig. 3
12Pl.06,
Pl.09, fig. 4
13
Pl.06,
Pl.09,
Pl.06, fig. 14
5
Pl.09, fig. 6
15
Pl.06,
Pl.09, fig. 7
16
Pl.06,
Pl.10, fig. 8
1
Pl.06,
Pl.10, fig. 92Pl.06,
Pl.10, fig. 10
3
Pl.06,
Pl.10,
Pl.06, fig. 4
11
Pl.10, fig. 12
5
Pl.06,
Pl.10, fig. 13
6
Pl.06,
Pl.10, fig. 1
7
Pl.07,
Pl.10, fig. 2
8Pl.07,
Pl.10, fig. 3
9Pl.07,
Pl.10, fig. 4
10
Pl.07,
Pl.10, fig. 11
Pl.10, fig. 12
Pl.10, fig. 13
Pl.10, fig. 14
Pl.10, fig. 15
Pl.10, fig. 16
Pl.10, fig. 17
Pl.11, fig. 1
Pl.11, fig. 2
Pl.11, fig. 3

11H-CC, E21/2
RN5
Hexacontium cf. arachnoidale Hollande and Enjumet
Saturnalis circularis Haeckel
11H-CC, W43/1
RN5 Nishimura and Yamauchi, 1984
Cenosphaera sp. A
15H-CC, Q45/4
RN5
Heliodiscus sp. B
15H-CC, X21/1
RN5
Hexapyle dodecantha Haeckel
15H-CC, L15/2
RN5 Haeckel, 1887
Excentrodiscus sp. A
15H-CC, R12/0
RN5
in theE12/0
tropical Pacific (Kamikuri)
Cenosphaera sp. BEarly to middle Miocene radiolarians
15H-CC,RN5
Heliodiscus sp. A
16H-CC, C46/0
RN5
Thecosphaera sp. E
15H-CC, N37/0
RN5
Hexacontium sp. C
15H-CC, V37/3
RN5
Axoprunum sp. A
15H-CC, Q45/4
RN5
Table 1 Continued.
Didymocyrtis violina (Haeckel)
18H-CC, P33/2
RN4 Sanfilippo et al. , 1985
Species and species
Location T46/4
Zone Sanfilippo
Referenceet al. , 1985
Didymocyrtis
tubaria group(Haeckel)
17H-CC,
RN4
Trisolenia
combinata
Bjørklundand Goll
13H-2, 104-106
Bjørklund and1979
Didymocyrtis
mammifera(Haeckel)
15H-CC,
X21/0 cm, D32/4 RN5 Sanfilippo, 1985et al.Goll,
Collosphaera
macroporaHaeckel
Popofsky
15H-CC, B26/3
RN5 Popofsky,
Spongurus cylindricus
Y39/3
Takahashi,1917
1991
Trisolenia
sp. sp.
A A
15H-CC, L44/2
K44/0
Cladococcus
16H-CC,
RN5
Siphonosphaera
sp. C
13H-2,
104-106
Hexacontium pachydermum
Jørgensen
13H-CC,
Y40/1 cm, Z20/0 RN5 Petrushevskaya and Kozlova, 1972
Collosphaera
pylomaReynolds
13H-CC,
RN5
Hexalonche sp.
A
21H-CC, Q23/0
S23/3
RN3 Reynolds, 1980
Collosphaera
10H-CC,
RN5
Didymocyrtis reynoldsi
prismaticaKamikuri(Haeckel)
25H-CC, H23/1
D11/2
RN2 Kamikuri,
Sanfilippo 2010
et al. , 1985
Trisolenia
megalactis
costlowi
Bjørklund and Goll
13H-2, 104-106
RN5 Sanfilippo
Bjørklund and1979
Didymocyrtis
prismatica(Haeckel)
23H-CC,O31/0 cm, Y39/4 RN2et al.Goll,, 1985Polysolenia
15H-CC,RN5 Nigrini
andetLombari,
1984
Didymocyrtisspinosatubaria(Haeckel)(Haeckel)group
16H-CC, R37/2
G16/0
Sanfilippo
al. , 1985
Polysolenia(Haeckel)
11H-CC,RN5 Nigrini
andetLombari,
1984
Didymocyrtismurrayana
tubaria (Haeckel)
16H-CC, T33/3
K41/2
Sanfilippo
al. , 1985
Polysolenia(Haeckel)
11H-CC,RN5 Nigrini
andetMoore,
1979
Didymocyrtisspinosalaticonus(Riedel)group
10H-CC, D20/1
L30/2
Sanfilippo
al. , 1985
Trisolenia
megalactis
megalactis
Ehrenberg
17H-CC,
W48/4 cm, D23/1 RN5RN4 Sanfilippo
Bjørklund and1979
Didymocyrtis
mammifera(Haeckel)
13H-2,
104-106
et al.Goll,, 1985
Trisolenia
megalactis
18H-CC, Y43/0
G40/4
RN4 Sanfilippo
Bjørklund and1979
Didymocyrtis
bassaniimegalactis(Carnevale)Ehrenberg
25H-CC,RN2
et al.Goll,, 1985
Trisolenia
megalactis
Ehrenberg
Y48/1
BjørklundGoll,1979
1979
Cladococcus
dentata megalactis(Mast)
17H-CC, O45/1
RN4 Nigrini
andandMoore,
Siphonosphaera
sp. A (Carnevale)
17H-CC,
RN4
Didymocyrtis bassanii
13H-CC, G41/3
V12/0
RN5 Sanfilippo et al. , 1985
Siphonosphaera
sp. A (Carnevale)
17H-CC,RN4 Sanfilippo et al. , 1985
Didymocyrtis bassanii
18H-CC, D22/1
Q37/0
Siphonosphaera
D group
18H-CC, U42/3
R10/1
RN4 Lazarus et al. , 2005
Larcopyle hayesi sp.
(Chen)
15H-CC,
RN5
Siphonosphaera
B group
13H-2, 104-106 cm, P39/2
Larcopyle hayesi sp.
(Chen)
Z24/0 RN5 Lazarus et al. , 2005
Collosphaera
glebulenta
and Goll
19H-CC, V21/1
RN3 Bjørklund
Larcopyle hayesi(Chen) Bjørklund
group
W15/2
Lazarus etand
al. ,Goll,
20051979
Species andspecies
group
Location R21/3
Zone Bjørklund
Reference
Collosphaera
glebulenta
Bjørklundand Goll
13H-CC,
RN5
Larcopylehayesi(Chen)
group
23H-CC,
C26/0
RN2
Lazarusetand
al. ,Goll,
20051979
Larcopyle sp. sp.
C A
13H-2, 104-106
Collosphaera
15H-CC,E36/3 cm, M39/0 RN5Lithelius sp. Asp. A
12H-4, 104-106 cm, Q37/3
X27/2 RN5
Collosphaera
13H-2,
Larcopyle polyacantha(Campbell and Clark)
23H-CC, J15/4
N37/0
RN3 Motoyama, 1996
Collosphaera
sp. A
13H-CC,
RN5
Larcopylesp.brattstroemi
B
17H-CC, S38/4
S14/3
RN4 Bjørklund and Goll, 1979
Collosphaera
Bjørklund and Goll
10H-CC,
RN5
Lithelius sp.bispiculumA
16H-CC,RN5 Takemura, 1992
Axoprunum(Popofsky)
11H-CC, K31/1
K20/0
Lithelius sp.bispiculumA
15H-CC,RN5 Takemura, 1992
Axoprunum(Popofsky)
11H-CC, Q43/3
V12/1
Lithelius sp. Acommunis (Carnevale)
23H-CC, N37/1
RN2 Sanfilippo et al. , 1978
Stylosphaera
D38/0
Lithelius sp. Csp. B
25H-CC, P35/0
J36/2
RN2
Stylosphaera
13H-CC,
RN5
Larcopyle sp. Asantaeannae (Campbell and Clark)
10H-CC, S38/0
F22/2
Amphisphaera
16H-CC,
RN5 Petrushevskaya and Kozlova, 1972
LarcopyleA
15H-CC,RN5 Takemura, 1992
Amphistylussp.polistylus(Carnevale)
16H-CC, X33/4
X40/0
Larcopyle sp. Aneptunus (Haeckel)
15H-CC, U43/3
J12/0
RN5 Petrushevskaya and Kozlova, 1972
Amphisphaera
17H-CC,
RN4
Larcopyle titansp.(Campbell
and Clark)
15H-CC, L46/4
F40/0
RN5 Campbell and Clark, 1944
StylosphaeraA
23H-CC,RN2
Actinomma sp.
B
17H-CC, U45/2
M44/2
RN4 Campbell and Clark, 1944
Amphistylus
angelinus(Campbell and Clark)
16H-CC,
RN5
Larcospira moschkowskii
Kruglikova
10H-CC,
RN4
Stylosphaera
sp. B
13H-CC, V23/0
F14/1
RN5 Nigrini and Lombari, 1984
Stylodictya
sp. A (Kling)
15H-CC, G38/0
R43/2
RN4 Morley and Nigrini, 1995
Actinomma robusta
10H-CC,
RN5
Stylodictya robusta
sp. A (Kling)
11H-CC, W20/1
C14/4
RN4 Morley and Nigrini, 1995
Actinomma
10H-CC,
RN5
Stylodictya sp. sp.
A A
25H-CC,RN2
Thecosphaera
11H-CC, R36/0
O35/0
RN5
Spongurus cylindricus
Haeckel
17H-CC,
J16/0 cm, E39/4 RN5RN4 Morley
Takahashi,
Actinomma
robusta (Kling)
13H-2,
104-106and 1991
Nigrini, 1995
Stylodictya camerina
14H-2,
105-107
Thecosphaera
sp. B Campbell and Clark
13H-CC,
U36/0 cm, N10/2 RN5 Campbell and Clark, 1944
Stylodictya camerina
25H-CC,RN2
Thecosphaera
sp. C Campbell and Clark
18H-CC, T36/2
N15/3
RN4 Campbell and Clark, 1944
Stylodictya
sp. sp.
B D
15H-CC,
RN5
Thecosphaera
18H-CC, U34/1
H20/3
RN4
Stylodictya camerina
15H-CC,
RN5
Thecosphaera
sp. E Campbell and Clark
25H-CC, Y32/3
T34/2
RN2 Campbell and Clark, 1944
Amphymeniumsp.
amphistylium
Haeckel
11H-CC, U27/3
D27/1
RN5 Morley and Nigrini, 1995
Hexacontium
B
20H-CC,
RN3
Stylodictya sp.cf.Carachnoidale Hollande and Enjumet
25H-CC,
RN2
11H-CC, P30/3
E21/2
RN5
Hexacontium
Porodiscuscircularis
circularisHaeckel
Clark and Campbell
10H-CC,RN5 Clark
and Campbell,1942 1984
Saturnalis
11H-CC, J36/1
W43/1
Nishimuraand Yamauchi,
Porodiscus sp.sp.A A
Z39/1
Cenosphaera
15H-CC, Q45/4
RN5
A
16H-CC, X21/1
F38/2
Heliodiscus sp. B
15H-CC,
RN5
Phacodiscus
sp. A
11H-CC,RN5 Haeckel, 1887
Hexapyle
dodecantha
Haeckel
15H-CC, T19/3
L15/2
Dictyocoryne sp.sp.
BA
16H-CC, R12/0
V34/0
Excentrodiscus
15H-CC,RN5
Spongasteriscus
Martin25H-CC,
RN2
Cenosphaera
sp. marylandicus
B
15H-CC, D28/0
E12/0
RN5 Palmer, 1986
C
17H-CC,RN4
Heliodiscus sp. A
16H-CC, E31/0
C46/0
RN5
Tetrapyle
sp. Asp. E
10H-CC,
RN5
Thecosphaera
15H-CC, D40/0
N37/0
Tetrapyle sp. A
17H-CC, V37/3
G37/1
RN4
Hexacontium
sp. C
15H-CC,
RN5
Dictyocorynesp.malagaense(Campbell and Clark)
10H-CC, Q45/4
G46/3
AxoprunumA
15H-CC,RN5 Campbell and Clark, 1944
Spongaster sp.
A
13H-CC,
RN5
Didymocyrtis
violina(Haeckel)
18H-CC, J30/1
P33/2
RN4 Sanfilippo et al. , 1985
Dictyocoryne tubaria
sp. A (Haeckel)
18H-CC, T46/4
U32/0
Didymocyrtis
17H-CC,RN4 Sanfilippo et al. , 1985
Spongopyle osculosa
Dreyer
17H-CC, X21/0
H46/4
RN4 Sanfilippo
Nigrini andetMoore,
1979
Didymocyrtis
mammifera(Haeckel)
15H-CC,
RN5
al. , 1985
Spongodiscus
resurgensHaeckel
Ehrenberg
17H-CC,RN4
Spongurus cylindricus
15H-CC, Q31/4
Y39/3
RN5 Petrushevskaya
Takahashi, 1991 and Kozlova, 1972
Spongodiscus sp.
resurgens
Ehrenberg
23H-CC, L44/2
O44/4
RN2 Petrushevskaya and Kozlova, 1972
Cladococcus
A
16H-CC,RN5
Spongodiscuspachydermum
resurgens Ehrenberg
23H-CC,
RN2
Hexacontium
Jørgensen
13H-CC, O43/4
Y40/1
RN5 Petrushevskaya and Kozlova, 1972
Spongodiscussp.resurgens
Ehrenberg
17H-CC,
RN4
HexaloncheA
21H-CC, R29/2
S23/3
RN3 Petrushevskaya and Kozlova, 1972
Excentrococcus
annulatus
Dumitrica
16H-CC, D11/2
H13/4
RN5 Sanfilippo
Dumitrica, et
1978
Didymocyrtis
prismatica(Haeckel)
25H-CC,
RN2
al. , 1985
Spongodiscus
sp. B
16H-CC,
RN5
Didymocyrtis prismatica(Haeckel)
23H-CC, C29/3
O31/0
RN2 Sanfilippo et al. , 1985
Spongodiscustubaria
sp. C (Haeckel)
15H-CC, G16/0
N21/3
Didymocyrtis
16H-CC,RN5 Sanfilippo et al. , 1985
Spongodiscus tubaria
klingi Caulet
25H-CC,
RN2
1986
Didymocyrtis(Haeckel)
16H-CC, J34/2
K41/2
RN5 Caulet,
Sanfilippo
et al. , 1985
Cyrtocapsella laticonus
tetrapera (Riedel)
HaeckelU23/0
Sakai, 1980et al. , 1985
Didymocyrtis
10H-CC, L30/2
RN5 Sanfilippo
Cyrtocapsella mammifera
tetrapera Haeckel
12H-4, 104-106 cm, D23/1
W44/1 RN5 Sanfilippo
Sakai, 1980et al. , 1985
Didymocyrtis(Haeckel)
13H-2,
Cyrtocapsella bassanii
japonica(Carnevale)(Nakaseko)
17H-CC, Y43/0
X10/1
RN4 Sanfilippo
Sakai, 1980et al. , 1985
Didymocyrtis
25H-CC,
RN2
Cyrtocapsellajaponica(Mast)(Nakaseko)
19H-CC, O45/1
S42/4
RN3 Nigrini
Sakai, 1980
Cladococcus dentata
17H-CC,
RN4
and Moore, 1979
Cyrtocapsella bassanii
cornuta (Carnevale)
Haeckel
12H-4, 104-106
Sakai, 1980et al. , 1985
Didymocyrtis
13H-CC,
V12/0 cm, V18/0 RN5 Sanfilippo
Cyrtocapsella bassanii
cornuta (Carnevale)
Haeckel
10H-CC, Q37/0
Y15/2
RN5 Sanfilippo
Sakai, 1980et al. , 1985
Didymocyrtis
18H-CC,
RN4, 1985
Lithopera baueri
and Riedel
13H-2,
104-106
Larcopylehayesi Sanfilippo(Chen) group
15H-CC,U42/3 cm, X30/4 RN5 Sanfilippo
Lazarus etetal.al.
, 2005
Stichocorysdelmontensis(Campbell and Clark)
13H-CC,
T33/1 cm, Z24/0 RN5 Sanfilippo, 1985
Larcopylehayesi(Chen) group
13H-2, 104-106
Lazarus etetal.al.
, 2005
Lithopera renzae
Sanfilippo
and Riedel
15H-CC, W15/2
L37/0
RN5 Lazarus
Sanfilippoetetal.al.
, 1985
Larcopylehayesi (Chen)group
19H-CC,
RN3, 2005
Stichocorys
wolffii(Chen)
Haeckel
21H-CC, C26/0
R41/1
RN3 Lazarus
Sanfilippoetetal.al.
, 1985
Larcopylehayesigroup
23H-CC,
RN2, 2005
Lithopera thornburgi Sanfilippo and Riedel
10H-CC, U43/0
RN5 Sanfilippo et al. , 1985
Eucyrtidium sp. A
23H-CC, D38/0
RN2
Stichocorys armata (Haeckel)
19H-CC, L40/3
RN3 Sanfilippo et al. , 1985
Phormocyrtis alexandrae O'Connor
25H-CC, Z39/1
RN2 O'Connor, 1997b−
139−
Eucyrtidium sp. C
17H-CC, N12/3
RN4
Eucyrtidium cienkowskii Haeckel
10H-CC, V9/3
RN5 Sakai, 1980
Stichocorys diaphanes (Sanfilippo and Riedel)
27H-CC, C46/1
RN2 Sanfilippo et al. , 1985
Carpocanium cingulatum (Riedel and Sanfilippo)
18H-CC, T15/0
RN4 Nigrini and Lombari, 1984
Carpocanium bramlettei (Riedel and Sanfilippo)
15H-CC, E44/1
RN5 Nigrini and Lombari, 1984
Carpocanium bramlettei (Riedel and Sanfilippo)
10H-CC, Z44/1
RN5 Nigrini and Lombari, 1984

Pl.09, fig. 15
Pl.09, fig. 16
Pl.10, fig. 1
Pl.10, fig. 2
Pl.10, fig. 3
Pl.10, fig. 4
Pl.10, fig. 5
Pl.10, fig. 6
Pl.10, fig. 7
Pl.10, fig. 8
Pl.10, fig. 9
Pl.10, fig. 10
Plate no.
Pl.10,
fig. 11
Pl.01, fig. 12
1
Pl.10,
Pl.01, fig. 13
2
Pl.10,
Pl.01, fig. 14
3
Pl.10,
Pl.01, fig. 15
4
Pl.10,
Pl.01, fig. 16
5
Pl.10,
Pl.01, fig. 17
6
Pl.10,
Pl.01, fig. 17
Pl.11,
Pl.01, fig. 28
Pl.11,
Pl.01, fig. 39
Pl.11,
Pl.01, fig. 410
Pl.11,
Pl.02, fig. 51
Pl.11,
Pl.02, fig. 62
Pl.11,
Pl.02, fig. 73
Pl.11,
Pl.02, fig. 84
Pl.11,
Pl.02, fig. 95
Pl.11,
Pl.02, fig. 10
6
Pl.11,
Pl.02, fig. 11
7
Pl.11,
Plate
Pl.02, no.
8
Pl.11,
fig. 12
Pl.11, fig. 9
13
Pl.02,
Pl.11, fig. 10
14
Pl.02,
Pl.11, fig. 11
15
Pl.02,
Pl.11, fig. 12
16
Pl.02,
Pl.11,
Pl.02, fig. 17
13
Pl.11, fig. 1
18
Pl.03,
Pl.11, fig. 2
19Pl.03,
Pl.11, fig. 3
20
Pl.03,
Pl.11, fig. 4
21
Pl.03,
Pl.12, fig. 5
1
Pl.03,
Pl.12, fig. 62Pl.03,
Pl.12, fig. 7
3Pl.03,
Pl.12, fig. 8
4Pl.03,
Pl.12, fig. 95
Pl.03,
Pl.12, fig. 10
6
Pl.03,
Pl.12, fig. 1
7
Pl.04,
Pl.12, fig. 2
8Pl.04,
Pl.12, fig. 3
9Pl.04,
Pl.13, fig. 4
1
Pl.04,
Pl.13, fig. 52Pl.04,
Pl.13, fig. 6
3
Pl.04,
Pl.13,
Pl.04, fig. 4
7
Pl.13, fig. 85Pl.04,
Pl.13, fig. 96Pl.04,
Pl.14, fig. 10
1
Pl.04,
Pl.14, fig. 11
2
Pl.04,
Pl.14, fig. 1
3
Pl.05,
Pl.14, fig. 2
4
Pl.05,
Pl.14, fig. 3
5Pl.05,
Pl.14, fig. 4
6
Pl.05,
Pl.14, fig. 5
7Pl.05,
Pl.14, fig. 86
Pl.05,
Pl.14,
Pl.05, fig. 9
7Pl.14, fig. 8
10Pl.05,
Pl.14, fig. 9
11Pl.05,
Pl.14, fig. 10
12
Pl.05,
Pl.14, fig. 11
13
Pl.05,
Pl.14,
Pl.05, fig. 14
12
Pl.14, fig. 13
15
Pl.05,
Pl.15, fig. 1
Pl.06,
Pl.15, fig. 2
Pl.06,
Pl.15, fig. 3
Pl.06,
Pl.15, fig. 4
Pl.06,
Pl.15, fig. 5
Pl.06,
Pl.15, fig. 6
Pl.06,
Pl.16, fig. 7
1
Pl.06,
Pl.16, fig. 82Pl.06,
Pl.16, fig. 93Pl.06,
Pl.16, fig. 10
4
Pl.06,
Pl.16,
Pl.06, fig. 5
11
Pl.16, fig. 12
6
Pl.06,
Pl.16, fig. 13
7
Pl.06,
Pl.16, fig. 1
8
Pl.07,
Pl.16, fig. 2
9Pl.07,
Pl.17, fig. 3
1
Pl.07,
Pl.17, fig. 42
Pl.07,
Pl.17, fig. 3
Pl.17, fig. 4
Pl.17, fig. 5
Pl.17, fig. 6
Pl.17, fig. 7
Pl.18, fig. 1
Pl.18, fig. 2
Pl.18, fig. 3
Pl.18, fig. 4
Pl.18, fig. 5

Spongodiscus sp. C
15H-CC, N21/3
RN5
Spongodiscus klingi Caulet
25H-CC, J34/2
RN2 Caulet, 1986
Cyrtocapsella tetrapera Haeckel
10H-CC, U23/0
RN5 Sakai, 1980
Cyrtocapsella tetrapera Haeckel
12H-4, 104-106 cm, W44/1 RN5 Sakai, 1980
Cyrtocapsella japonica (Nakaseko)
17H-CC, X10/1
RN4 Sakai, 1980
Cyrtocapsella japonica (Nakaseko)
19H-CC, S42/4
RN3 Sakai, 1980
Bulletin of the Geological Surveyof Japan,vol.
(1/2), 2019
Cyrtocapsella cornuta Haeckel
12H-4,
104-106
cm,70V18/0
RN5 Sakai, 1980
Cyrtocapsella cornuta Haeckel
10H-CC, Y15/2
RN5 Sakai, 1980
Lithopera baueri Sanfilippo and Riedel
13H-2, 104-106 cm, X30/4 RN5 Sanfilippo et al. , 1985
Stichocorys delmontensis (Campbell and Clark)
13H-CC, T33/1
RN5 Sanfilippo et al. , 1985
Lithopera renzae Sanfilippo and Riedel
15H-CC, L37/0
RN5 Sanfilippo et al. , 1985
Table 1 Continued.
Stichocorys wolffii Haeckel
21H-CC, R41/1
RN3 Sanfilippo et al. , 1985
Species andspecies group
Location U43/0
Zone Sanfilippo
Referenceet al. , 1985
Lithopera
thornburgi
Sanfilippo and Riedel
10H-CC,
RN5
Trisolenia combinata
Bjørklund and Goll
13H-2, 104-106
RN5 Bjørklund and Goll, 1979
Eucyrtidium
sp. A
23H-CC,
D38/0 cm, D32/4 RN2Collosphaeraarmata
macropora
Popofsky
15H-CC, L40/3
B26/3
RN5 Sanfilippo
Popofsky, et
1917
Stichocorys(Haeckel)
19H-CC,
RN3
al. , 1985
Trisolenia sp. alexandrae
A
15H-CC, Z39/1
K44/0
RN5 O'Connor, 1997b
Phormocyrtis
O'Connor
25H-CC,
RN2
Siphonosphaera
13H-2, 104-106
RN5
Eucyrtidium
sp. Csp. C
17H-CC,
N12/3 cm, Z20/0 RN4Collosphaeracienkowskii
pyloma Reynolds
13H-CC, V9/3
Q23/0
Reynolds,
Eucyrtidium
Haeckel
10H-CC,RN5 Sakai,
19801980
Collosphaeradiaphanes
reynoldsi Kamikuri
10H-CC, C46/1
H23/1
RN5 Sanfilippo
Kamikuri, 2010
Stichocorys(Sanfilippo and Riedel)
27H-CC,RN2
et al. , 1985
Trisolenia megalactis
costlowi
Bjørklundand Goll
13H-2, 104-106
RN5 Nigrini
BjørklundGoll, 1979
Carpocanium
cingulatum(Riedeland Sanfilippo)
18H-CC,
T15/0 cm, Y39/4 RN4andandLombari,
1984
Polysolenia spinosa(Haeckel)
R37/2
Carpocanium
bramlettei(Riedelgroup
and Sanfilippo)
15H-CC, E44/1
RN5 Nigrini and Lombari, 1984
Polysolenia murrayana
11H-CC, Z44/1
T33/3
Carpocanium
bramlettei(Haeckel)(Riedel and Sanfilippo)
10H-CC,RN5 Nigrini and Lombari, 1984
Polysolenia spinosa
11H-CC, T42/0
D20/1
RN5 Nigrini and Moore, 1979
Carpacanium
sp. B (Haeckel) group
23H-CC,
RN2
Trisolenia megalactis
megalactis
Ehrenberg
17H-CC, S34/1
W48/4
RN4 Nishimura,
Bjørklund andGoll, 1979
Carpacanium
kinugasense
Nishimura
10H-CC,
RN5
1990
Trisolenia megalactis
megalactis
Ehrenberg
18H-CC, G44/1
G40/4
RN4 Nishimura,
Bjørklund andGoll, 1979
Carpacanium
kinugasense
Nishimura
25H-CC,
RN2
1990
Trisolenia megalactis
Y48/1
Siphocampe
sp. D megalactis Ehrenberg
17H-CC, K18/3
RN4 Bjørklund and Goll, 1979
Siphonosphaera
sp. A
17H-CC, L24/0
G41/3
RN4 O'Connor, 1997b
Siphocampe
grantmackiei
O'Connor
15H-CC,
RN5
Siphonosphaerasp.sp.BA
17H-CC, M43/0
D22/1
RN4
Botryostrobus
10H-CC,
RN5
Siphonosphaeraaquilonaris
sp. D
18H-CC, W38/2
R10/1
RN4 Nigrini and Lombari, 1984
Botryostrobus(Bailey)
15H-CC,
RN5
Siphonosphaera
sp.PetrushevskayaB
13H-2, 104-106 cm, D39/2
P39/2 RN5 Nigrini and Lombari, 1984
Spirocyrtis
subtilis
12H-4,
Species and species
group
Location E20/3
Zone Nigrini
Reference
Collosphaera
glebulenta
Bjørklund and Goll
19H-CC,
V21/1
RN3
BjørklundGoll, 1979
Siphostichartus
corona(Haeckel)
10H-CC,
RN5
andandLombari,
1984
Botryostrobusglebulenta
sp. A
15H-CC, R21/3
L37/0
Collosphaera
Bjørklund and Goll
13H-CC,RN5 Bjørklund and Goll, 1979
Theocamptra ovata
16H-CC,RN5 Nigrini and Lombari, 1984
Collosphaera
sp. A (Haeckel)
15H-CC, Z42/0
E36/3
Siphocampe sp.
13H-CC,
V36/0 cm, Q37/3 RN5Collosphaera
sp.BA
13H-2,
104-106
Siphocampe sp.
15H-CC, J15/4
T38/1
Collosphaera
sp.BA
13H-CC,RN5
Phormostichoartus
fistula Nigrini
15H-CC, S38/4
M34/0
RN5 Bjørklund
Nigrini andand
Lombari,
1984
Collosphaera brattstroemi
Bjørklund and Goll
10H-CC,
Goll, 1979
Phormostichoartus
fistula
Nigrini
11H-CC, K40/0
RN5 Nigrini
and 1992
Lombari, 1984
Axoprunum
bispiculum(Popofsky)
K20/0
Takemura,
Lithocampana
sp. A (Popofsky)
17H-CC,
RN4
Axoprunum
bispiculum
11H-CC, X10/1
V12/1
RN5 Takemura, 1992
Lithocampanacommunis
sp. B
17H-CC,
RN4
Stylosphaera(Carnevale)
23H-CC, Y42/1
D38/0
RN2 Sanfilippo et al. , 1978
Cycladophora sp.
conica25H-CC,RN2
Stylosphaera
B Lombari and Lazarus
13H-CC, F36/4
P35/0
RN5 Lombari and Lazarus, 1988
Calocycletta caepa
Moore (Campbell and Clark)
16H-CC, N38/4
RN5 Nigrini
and Lombari,
Amphisphaera
santaeannae
S38/0
Petrushevskaya
and 1984
Kozlova, 1972
Calocycletta polistylus
virginis (Haeckel)
17H-CC, X40/0
T42/1
RN4 Takemura,
Nigrini and 1992
Lombari, 1984
Amphistylus(Carnevale)
16H-CC,
RN5
Calocycletta serrata
Moore
30H-CC, U43/3
S38/0
RN2 Petrushevskaya
Nigrini and Lombari,
Amphisphaera
neptunus(Haeckel)
17H-CC,
RN4
and 1984
Kozlova, 1972
Calocycletta costata
14H-CC,
RN5
Stylosphaera
sp. A (Riedel)
23H-CC, V39/4
L46/4
RN2 Nigrini and Lombari, 1984
Calocycletta angelinus
robusta Moore
31H-CC, U45/2
X43/4
RN2 Campbell
Nigrini andand
Lombari,
Amphistylus(Campbell and Clark)
16H-CC,
RN5
Clark, 1984
1944
Albatrossidiumsp.
minzok
Sanfilippo and Riedel
10H-CC, F14/1
J23/3
Stylosphaera
B
13H-CC,RN5 Sanfilippo and Riedel, 1992
Anthocyrtidium
sp. A(Kling)
25H-CC, G38/0
Y37/3
RN2 Morley and Nigrini, 1995
Actinomma
robusta
10H-CC,
RN5
Anthocyrtidium
sp. B(Kling)
25H-CC, W20/1
T46/1
RN2 Morley and Nigrini, 1995
Actinomma
robusta
10H-CC,
RN5
Cyrtolagena sp.sp.
AA
19H-CC,
RN3
Thecosphaera
11H-CC, M26/4
O35/0
RN5
Lophocyrtis robusta
tanythorax(Sanfilippo and Riedel)
W25/4 RN5 Morley
Sanfilippo
Riedel,1970
Actinomma(Kling)
13H-2, 104-106 cm, E39/4
andandNigrini,
1995
Lophocyrtis tanythorax(Sanfilippo and Riedel)
15H-CC,RN5 Sanfilippo and Riedel, 1970
Thecosphaera
sp. B
13H-CC, S38/4
U36/0
Cinclopyramis pacifica
18H-CC, N12/4
RN4 Nakaseko, 1963
Thecosphaera
sp. C (Nakaseko)
N15/3
Cinclopyramis(Campbell and Clark)
17H-CC,RN4 Campbell and Clark, 1944
Thecosphaera woodringisp. D
18H-CC, T16/0
H20/3
Lophocyrtis sp.sp.
AE
25H-CC, Y43/4
RN2
Thecosphaera
T34/2
Calocyclura sp.
12H-4,
104-106
Hexacontium
sp.AB
20H-CC,
U27/3 cm, Y24/4 RN5
RN3
Lamprocyclas cf.
maritalis
HaeckelHollande and Enjumet
M20/0
11H-CC, E21/2
RN5 Nigrini and Moore, 1979
arachnoidale
Hexacontium
Lophocyrtiscircularis
aspera (Ehrenberg)
25H-CC,
RN2
1998
Saturnalis
Haeckel
11H-CC, S29/3
W43/1
RN5 Sanfilippo
Nishimura and Caulet,
Yamauchi,
1984
Theocorys spongoconus
Kling
27H-CC, Q45/4
C28/0
RN2 Nigrini and Lombari, 1984
Cenosphaera
sp. A
15H-CC,
RN5
Lamprocyclas
27H-CC, X21/1
H49/3
RN2
Heliodiscus
sp.sp.
BB
15H-CC,
RN5
Theocorysdodecantha
sp. A
17H-CC,
RN4
Hexapyle
Haeckel
15H-CC, J22/0
L15/2
RN5 Haeckel, 1887
Clathrocanium atreta
25H-CC, R12/0
C45/1
RN2 Sanfilippo et al. , 1973
Excentrodiscus
sp. A Sanfilippo and Riedel
15H-CC,
RN5
Lithomelissa sp.
11H-CC, E12/0
D22/0
Cenosphaera
sp. AB
15H-CC,
RN5
Lithomelissasp.
sp.AA
12H-4,
104-106
Heliodiscus
16H-CC,
C46/0 cm, M39/0 RN5Lophophaena
11H-CC,
RN5
Thecosphaera sp.
sp.AE
15H-CC, G19/0
N37/0
Ceratocyrtis sp.
12H-4,
104-106
Hexacontium
sp. AC
15H-CC,
V37/3 cm, K44/1 RN5Lamprocyclas
Campbell and Clark var. A
16H-CC, Q45/4
U9/3
Axoprunum
sp.margatensis
A
15H-CC,RN5 Campbell and Clark, 1944
Lipmanella conica
25H-CC,RN2
Didymocyrtis
violinaPetrushevskaya(Haeckel)
18H-CC, T28/3
P33/2
RN4 Petrushevskaya
Sanfilippo et al. , and1985Kozlova, 1979
Lophophaena tubaria
tekopua(Haeckel)
O'Connor
12H-4,
104-106
1997a
Didymocyrtis
17H-CC,
T46/4 cm, X36/2 RN5RN4 O'Connor,
Sanfilippo etal. , 1985
Lophophaena mammifera
tekopua O'Connor
25H-CC,
RN2
1997a
Didymocyrtis(Haeckel)
15H-CC, L40/4
X21/0
RN5 O'Connor,
Sanfilippo etal. , 1985
Clathrocanium
coarctatum
Ehrenberg
16H-CC,RN5 Takahashi, 1991
Spongurus
cylindricus
Haeckel
15H-CC, Z40/0
Y39/3
Pterocanium audax
14H-2,
105-107
Cladococcus
sp. A (Riedel)
16H-CC,
L44/2 cm, G17/3 RN5 Nigrini and Lombari, 1984
Pterocanium audax(Riedel) Jørgensen
23H=CC,RN2and Lombari,Hexacontium
pachydermum
13H-CC, J20/1
Y40/1
RN5 Nigrini
Petrushevskaya
and 1984
Kozlova, 1972
Lychnocanoma(Vinassa de Regny)
25H-CC,
RN2
Hexalonche
sp. elongataA
21H-CC, W33/4
S23/3
RN3 Sanfilippo et al. , 1985
(Ehrenberg)
13H-2,
104-106
Pterocanium prismatica
aff. tridentatum
Didymocyrtis(Haeckel)
25H-CC,
D11/2 cm, L15/0 RN5RN2 Sanfilippo et al. , 1985
Pterocanium charybdeum(Müller)
16H-CC,
RN5, 1985
Didymocyrtis
prismatica (Haeckel)
23H-CC, J43/0
O31/0
RN2 Lazarus
Sanfilippoetetal.al.
, 1985
Pterocanium tubaria
sp. A (Haeckel)
10H-CC, G16/0
E43/3
Didymocyrtis
16H-CC,RN5 Sanfilippo et al. , 1985
Lychnocanomatubaria
sp. A (Haeckel)
23H-CC,RN2
Didymocyrtis
16H-CC, P42/2
K41/2
RN5 Sanfilippo et al. , 1985
Lychnocanomalaticonus
sp. A (Riedel)
23H-CC,RN2
Didymocyrtis
10H-CC, E29/2
L30/2
RN5 Sanfilippo et al. , 1985
Lychnocanomamammifera
sp. A
27H-CC,
H41/1 cm, D23/1 RN2Didymocyrtis(Haeckel)
13H-2, 104-106
RN5 Sanfilippo et al. , 1985
Lychnocanomabassanii
nodosum(Haeckel)
21H-CC, Y43/0
S28/4
RN3 Sanfilippo
Haeckel, 1887
Didymocyrtis(Carnevale)
25H-CC,RN2
et al. , 1985
Lychnocanoma
nodosum(Haeckel)
18H-CC, O45/1
U41/3
RN4 Nigrini
Haeckel,1887
Cladococcus dentata(Mast)
17H-CC,
andMoore, 1979
Dictyophimus bassanii
splendens(Campbell and Clark)
11H-CC, V12/0
W43/1
Morley andetNigrini,
1995
Didymocyrtis(Carnevale)
13H-CC,RN5 Sanfilippo
al. , 1985
Dictyophimus bassanii
sp. A (Carnevale)
17H-CC, Q37/0
D12/0
Didymocyrtis
18H-CC,RN4 Sanfilippo et al. , 1985
Valkyria pukapuka
O'Connor
W44/1
O'Connor,
Larcopylehayesi (Chen)group
15H-CC, U42/3
RN5 Lazarus
et1997aal. , 2005
Valkyria pukapuka
O'Connor
13H-CC,D41/2 cm, Z24/0 RN5 Lazarus
O'Connor,
Larcopylehayesi (Chen)group
13H-2,
104-106
et1997aal. , 2005
Dorcadospyris
dentata
18H-CC, W15/2
U11/0
RN4 Lazarus
Sanfilippoetetal.al.
, 1985
Larcopylehayesi(Chen)Haeckel
group
19H-CC,
RN3, 2005
Dorcadospyris
alata(Riedel)
16H-3,
149-151, 1985
Larcopylehayesi(Chen)
group
23H-CC,
C26/0 cm, D44/0 RN5RN2 Sanfilippo
Lazarus etetal.al.
, 2005
Dorcadospyris ateuchus (Ehrenberg)
27H-CC, O46/1
RN2 Sanfilippo et al. , 1985
Lamprocyclas margatensis Campbell and Clark var. B 15H-CC, V38/2
RN5 Campbell and Clark, 1944
Lamprocyclas margatensis Campbell and Clark var. B 16H-CC, U27/4
RN5 Campbell and Clark, 1944
Lamprocyclas margatensis Campbell and Clark var. B 15H-CC, V36/3
RN5 Campbell and Clark, 1944−
140−
Cornutella trochus Ehrenberg
15H-CC, N37/0
RN5 Ehrenberg, 1873
Tympanomma binoctonum (Haeckel)
25H-CC, G43/3
RN2 Nigrini and Lombari, 1984
Tympanomma binoctonum (Haeckel)
18H-CC, N15/0
RN4 Nigrini and Lombari, 1984
Tholospyris anthophora (Haeckel)
19H-CC, E42/3
RN3 Nigrini and Lombari, 1984
Tholospyris anthophora (Haeckel)
17H-CC, T13/2
RN4 Nigrini and Lombari, 1984
Dendrospyris pannosa Goll
23H-CC, O44/4
RN2 Petrushevskaya and Kozlova, 1972

Pl.15, fig. 6 Pterocanium sp. A
10H-CC, E43/3
RN5
Pl.16, fig. 1 Lychnocanoma sp. A
23H-CC, P42/2
RN2
Pl.16, fig. 2 Lychnocanoma sp. A
23H-CC, E29/2
RN2
Pl.16, fig. 3 Lychnocanoma sp. A
27H-CC, H41/1
RN2
Pl.16, fig. 4 Lychnocanoma nodosum (Haeckel)
21H-CC, S28/4
RN3 Haeckel, 1887
Pl.16, fig. 5 Lychnocanoma nodosum (Haeckel)
18H-CC, U41/3
RN4 Haeckel, 1887
Early (Campbell
to middle andMiocenein theW43/1
tropical Pacific (Kamikuri)
Pl.16, fig. 6 Dictyophimus splendens
Clark) radiolarians
11H-CC,RN5 Morley and Nigrini, 1995
Pl.16, fig. 7 Dictyophimus sp. A
17H-CC, D12/0
RN4
Pl.16, fig. 8 Valkyria pukapuka O'Connor
15H-CC, W44/1
RN5 O'Connor, 1997aPl.16, fig. 9 Valkyria pukapuka O'Connor
13H-CC, D41/2
RN5 O'Connor, 1997aPl.17, fig. 1 Dorcadospyris dentata Haeckel
18H-CC, U11/0
RN4 Sanfilippo et al. , 1985
Table 1 Continued.
16H-3, 149-151 cm, D44/0 RN5 Sanfilippo et al. , 1985
Pl.17, fig. 2 Dorcadospyris alata (Riedel)
Plate
Species and species
group
Location
Zone
Pl.17, no.
fig. 3 Dorcadospyris
ateuchus(Ehrenberg)
27H-CC, O46/1
RN2 Reference
Sanfilippo et al. , 1985
Pl.01,
combinata
Bjørklundand Goll
13H-2,
104-106
Goll, 1979
Pl.17, fig. 1
4 Trisolenia
Lamprocyclas
margatensis
Campbelland Clark var. B
15H-CC,
V38/2 cm, D32/4 RN5 Bjørklund
Campbell and Clark,
1944
Pl.01,
15H-CC,
RN5 Popofsky,
1917Clark, 1944
Pl.17, fig. 2
5 Collosphaera
Lamprocyclasmacropora
margatensisPopofsky
Campbell and Clark var. B
16H-CC, B26/3
U27/4
Campbell andPl.01,
sp. A
K44/0
Pl.17, fig. 3
6 Trisolenia
Lamprocyclas
margatensis Campbell and Clark var. B 15H-CC, V36/3
RN5 Campbell and Clark, 1944
Pl.01,
sp. Ehrenberg
C
13H-2,
104-106
Pl.17, fig. 47 Siphonosphaera
Cornutella trochus
15H-CC,
N37/0 cm, Z20/0 RN5 Ehrenberg, 1873
Pl.01,
Reynolds
13H-CC,
RN5
Pl.18, fig. 5
1 Collosphaera
Tympanommapyloma
binoctonum(Haeckel)
25H-CC, Q23/0
G43/3
RN2 Reynolds,
Nigrini and1980
Lombari, 1984
Pl.01,
Kamikuri
10H-CC,
RN5
Pl.18, fig. 62 Collosphaera
Tympanommareynoldsi
binoctonum(Haeckel)
18H-CC, H23/1
N15/0
RN4 Kamikuri,
Nigrini and2010
Lombari, 1984
Pl.01,
costlowi
Bjørklund and Goll
13H-2, 104-106
RN5 Nigrini
BjørklundGoll, 1979
Pl.18, fig. 7
3 Trisolenia
Tholospyrismegalactis
anthophora(Haeckel)
19H-CC,E42/3 cm, Y39/4 RN3andandLombari,
1984
Pl.01,(Haeckel)group
15H-CC,
RN5
Pl.18, fig. 84 Polysolenia
Tholospyris spinosa
anthophora(Haeckel)
17H-CC, R37/2
T13/2
RN4 Nigrini and Lombari, 1984
Pl.01,(Haeckel)
11H-CC, O44/4
T33/3
RN5 Petrushevskaya
Nigrini and Lombari,
Pl.18, fig. 95 Polysolenia
Dendrospyrismurrayana
pannosa Goll
23H-CC,
RN2
and 1984
Kozlova, 1972
Pl.01,(Haeckel)group
11H-CC,
RN5
Moore, 1979
Pl.18, fig. 10
6 Polysolenia
Tholospyris spinosa
mammillaris(Haeckel)
19H-CC, D20/1
J49/3
RN3 Nigrini and Lombari,
1984
Pl.02,
megalactis Ehrenberg
17H-CC, K45/2
W48/4
RN4 Nigrini
BjørklundGoll, 1979
Pl.18, fig. 1
7 Trisolenia
Tholospyrismegalactis
kantiana (Haeckel)
10H-CC,
RN5
andandLombari,
1984
Pl.02,
megalactis
megalactis
Ehrenberg
18H-CC, V14/1
G40/4
Bjørklund and Goll,
Pl.18, fig. 2
8 Trisolenia
Tympanomma
tuberosum(Haeckel)
17H-CC,
RN4 Petrushevskaya
and1979
Kozlova, 1972
Pl.02,
megalactis Ehrenberg
17H-CC, D29/0
Y48/1
RN4 Nigrini
BjørklundGoll, 1979
Pl.18, fig. 3
9 Trisolenia
Liriospyris megalactis
mutuaria Goll
25H-CC,
RN2
andandLombari,
1984
Pl.02,
17H-CC,
RN4
Pl.18, fig. 410 Siphonosphaera
Liriospyris sp. A sp. A
25H-CC, G41/3
C43/4
RN2
Pl.02,
17H-CC,
RN4
Pl.18, fig. 511 Siphonosphaera
Dendrospyris sp.sp.
A A
10H-CC, D22/1
Q40/4
RN5
Pl.02,
sp. D(Haeckel)
18H-CC, X37/4
R10/1
RN4 Sanfilippo et al. , 1985
Pl.18, fig. 612 Siphonosphaera
Eucoronis octopylus
14H-CC,
RN5
Plate
Species andspecies
Location
Zone Reference
Pl.02,
fig. 7
sp. Bgroup
13H-2,
104-106
Pl.18, no.
13 Siphonosphaera
Eucoronis
perspicillum
Haeckel
16H-CC,
V22/3 cm, P39/2 RN5Petrushevskaya and Kozlova, 1972
Pl.18,
perspicillum
16H-CC, V21/1
P45/0
RN5 Bjørklund
Petrushevskaya
and1979
Kozlova, 1972
Pl.02, fig. 14
8 Eucoronis
Collosphaera
glebulentaHaeckel
Bjørklund and Goll
19H-CC,
RN3
and Goll,
Pl.18,
circumflexa(Goll) and Goll
25H-CC, R21/3
C45/1
RN2 Bjørklund
Nigrini andand
Lombari,
1984
Pl.02, fig. 15
9 Acanthodesmia
Collosphaera glebulenta
Bjørklund
13H-CC,
RN5
Goll, 1979
Pl.18,
25H-CC,
RN2
Pl.02, fig. 16
10 Acanthodesmia
Collosphaera sp.sp.
AA
15H-CC, X36/0
E36/3
RN5
Pl.18,
13H-2, 104-106 cm, Q30/0
Pl.02, fig. 17
11 Acanthodesmia
Collosphaera sp.sp.
AB
Q37/3 RN5
Pl.18,(Carnevale)
C46/3
Pl.02, fig. 18
12 Dendrospyris
Collosphaera pododendros
sp. A
13H-CC, J15/4
RN5 Petrushevskaya and Kozlova, 1972
Pl.18,
18H-CC, S38/4
U11/0
RN4 Bjørklund
Goll, 1969 and Goll, 1979
Pl.02, fig. 19
13 Giraffospyris
Collosphaera annulispina
brattstroemiGoll
Bjørklund and Goll
10H-CC,
RN5
Pl.18,(Haeckel)
14H-CC,RN5 Goll,
1969 1992
Pl.03, fig. 20
1 Eucoronis
Axoprunumtoxarium
bispiculum(Popofsky)
11H-CC, D16/2
K20/0
Takemura,
Pl.18,
stabilis (Goll)
C11/4
Nigrini and 1992
Lombari, 1984
Pl.03, fig. 21
2 Phormospyris
Axoprunum bispiculum(Popofsky)
11H-CC, V12/1
RN5 Takemura,
Pl.18,
sp. B
13H-CC, D38/0
D41/0
RN5 Sanfilippo et al. , 1978
Pl.03, fig. 22
3 Phormospyris
Stylosphaera communis(Carnevale)
23H-CC,
RN2
Pl.18,
parkerae
17H-CC, P35/0
Q31/0
RN4 Riedel and Sanfilippo, 1978
Pl.03, fig. 23
4 Liriospyris
Stylosphaera
sp. B Riedel and Sanfilippo
13H-CC,
RN5
Pl.18,
stauropora(Haeckel)
17H-CC, S38/0
J16/0
RN4 Petrushevskaya
Riedel and Sanfilippo,
1978 1972
Pl.03, fig. 24
5 Liriospyris
Amphisphaera
santaeannae(Campbell and Clark)
16H-CC,
RN5
and Kozlova,
Pl.18,
15H-CC, X40/0
Z39/1
Pl.03, fig. 25
6 Zygocircus
Amphistylussp.
polistylus (Carnevale)
16H-CC,RN5 Takemura, 1992
Pl.18,
sp.neptunus (Haeckel)
15H-CC, U43/3
S37/0
RN5 Petrushevskaya and Kozlova, 1972
Pl.03, fig. 26
7 Zygocircus
Amphisphaera
17H-CC,
RN4
Pl.18,
D38/3
Pl.03, fig. 27
8 Botryocyrtis
Stylosphaera sp.
sp. AA
23H-CC, L46/4
RN2
Pl.18, fig. 928 Centrobotrys
thermophila
Petrushevskaya
28H-CC, U45/2
S45/2
RN2 Campbell
Sanfilippo andet al.Clark,, 1985
Amphistylus angelinus(Campbelland Clark)
16H-CC,
RN5
1944
Pl.03,
Pl.18,
Sanfilippo and Riedel
28H-CC, F14/1
S45/2
RN2 Sanfilippo et al. , 1985
Pl.03, fig. 29
10 Centrobotrys
Stylosphaera petrushevskayae
sp. B
13H-CC,
RN5
Pl.18,
Haeckel
25H-CC, G38/0
G44/3
RN2 Morley
Haeckel,and
1887
Pl.04, fig. 30
1 Acrobotrys
Actinomma disolenia
robusta (Kling)
10H-CC,
RN5
Nigrini, 1995
Pl.04, fig. 2 Actinomma robusta (Kling)
10H-CC, W20/1
RN5 Morley and Nigrini, 1995
Pl.04, fig. 3 Thecosphaera sp. A
11H-CC, O35/0
RN5
Pl.04, fig. 4 Actinomma robusta (Kling)
13H-2, 104-106 cm, E39/4 RN5 Morley and Nigrini, 1995
Pl.04, fig. 570 unidentified
Thecosphaera sp.
B
U36/0
RN5Samples were provided by thetaxonomic
problems.
includeforms,
particularly within the 13H-CC,
Pl.04, fig. Actinommidae,
6 Thecosphaera Collosphaeridae,
sp. C
N15/3 Ocean Drilling
RN4
Integrated
Program (IODP). This work
families
Hexalonchidae, 18H-CC,
Pl.04, fig. 7 Thecosphaera sp. D
18H-CC, H20/3
RN4
was
also
supportedinpart
by the Japan Society for the
Litheliidae,
Pyloniidae,
Spongodiscidae
andStylodictyidae.
Pl.04, fig. 8 Thecosphaera sp. E
25H-CC, T34/2
RN2
Pl.04, fig.
Hexacontium
sp.described
B
U27/3 of ScienceRN3
Promotion(JSPS) KAKENHI Grant NumberSomeof 9thesetaxa will be
as new species in the 20H-CC,
E21/2
RN5
Pl.04, fig. 10 Hexacontium cf. arachnoidale Hollande and Enjumet
15K17780(S. Kamikuri).
near
future. The radiolarian assemblages included at least 68 11H-CC,
Pl.04, fig. 11 Saturnalis circularis Haeckel
11H-CC, W43/1
RN5 Nishimura and Yamauchi, 1984(e.g.
tropical-subtropicaltaxa sp.
Pl.05, fig. 1 Cenosphaera
A Acanthodesmia circumflexa, 15H-CC, Q45/4
RN5
Pl.05, fig. 2 Heliodiscus
sp. B
RN5
Calocycletta
caepa, Calocycletta
robusta, Carpocanium 15H-CC, X21/1
References
Pl.05, fig. 3 Hexapyle dodecantha Haeckel
15H-CC, L15/2
RN5
Haeckel, 1887
cingulatum,
Centrobotrys
petrushevskayae,
Collosphaera
Pl.05, fig. 4 Excentrodiscus sp. A
15H-CC, R12/0
RN5
brattstroemi,
Dendrospyris
pannosa, Didymocyrtis 15H-CC,
Bjørklund,
K. R. andPl.05, fig. 5 Cenosphaera
sp. B
E12/0
RN5 Goll, R. M. (1979) Internal
Pl.05, fig. 6 Didymocyrtis
Heliodiscus sp. A
RN5of Collosphaera and Trisolenia:
laticonus,
tubaria, Dorcadospyris alata, 16H-CC, C46/0
skeletal structures
Pl.05, fig. 7 Thecosphaera sp. E
15H-CC, N37/0
RN5
Eucoronis
dodecantha, Liriospyris 15H-CC, V37/3
a case of repetitive
Pl.05, fig. 8 octopylus,
HexacontiumHexapyle
sp. C
RN5evolution in the Collosphaeridae(Radiolaria). Jour.
Pl.05, fig. 9 Lithopera
Axoprunumthornburgi,
sp. A
RN5
parkerae,
Lophocyrtis tanythorax, 15H-CC, Q45/4
Paleontol., 53, 1293–1326.
Pl.05, fig. 10 Didymocyrtis violina (Haeckel)
18H-CC, P33/2
RN4 Sanfilippo et al. ,(1985 )Lychnocanoma
elongata,
Polysolenia
spinosa,
Pterocanium
Campbell,A.
S.
and
Clark, B. L. 1944 Miocene
Pl.05, fig. 11 Didymocyrtis tubaria (Haeckel)
17H-CC, T46/4
RN4 Sanfilippo et al. , 1985
audax,
Stichocorysarmata,Stichocorys
wolffii, Tholospyris 15H-CC, X21/0
radiolarian faunas
southernGeol.
mammifera(Haeckel)
RN5from
Sanfilippo
1985
et al. ,California.
Pl.05, fig.
12 Didymocyrtis
Pl.05, fig. 13 Sponguruscylindricus
Haeckel costlowi) and 31 15H-CC, Y39/3
anthophora,
Trisolenia
megalactis
Soc. Amer., Spec.RN5
Pap.,Takahashi,
no. 51, 1991
1–76.
Pl.06, fig. 1 Cladococcus sp. A
L44/2
(
)RN5
cosmopolitan
taxa (e.g. Actinomma
Amphisphaera 16H-CC,
Caulet,Radiolarians
fromandthe
southwest
Pl.06, fig. 2 Hexacontium
pachydermumrobusta,
Jørgensen
13H-CC,
Y40/1J. P. 1986 RN5Petrushevskaya
Kozlova,
1972
neptunus,
Pacific. In Kennett,
Pl.06, fig. 3 Amphymenium
Hexalonche sp. A amphistylium, Axoprunum 21H-CC, S23/3
RN3 J. P., von der Borch, C. C. et
Pl.06, fig. 4 Didymocyrtis
prismatica(Haeckel) Cinclopyramis 25H-CC, D11/2
RN2 Sanfilippo
et al.Washington, 1985(U.S.
bispiculum,
Botryostrobus
aquilonaris,al., eds., Init. Repts.
DSDP, 90,
Pl.06, fig. 5 Didymocyrtis prismatica (Haeckel)
23H-CC, O31/0
RN2 Sanfilippo et al. , 1985)
pacifica,
Cladococcus
dentata,
Cycladophoraconica,Government
Printing
Office,835–861.
Pl.06, fig. 6 Didymocyrtis tubaria (Haeckel)
16H-CC, G16/0
RN5 Sanfilippo et al. , 1985
Pl.06, fig. 7 Didymocyrtis
tubariaDictyophimus(Haeckel)
K41/2F. P. and Barber,
RN5 R.
Sanfilippoet)al.
, 1985
Cyrtocapsellajaponica,
splendens, 16H-CC,
Chavez,T. (1987
Anestimate of new
Pl.06, fig. 8 Didymocyrtis
laticonusLarcopyle(Riedel)Sanfilippo
et al. ,Deep-Sea
1985
Lamprocyclas
margatensis,
polyacantha, 10H-CC, L30/2
production in the RN5equatorial
Pacific.
Res. Pt.
Pl.06, fig. 9 Didymocyrtis mammifera (Haeckel)
13H-2, 104-106 cm, D23/1 RN5 Sanfilippo et al. , 1985
Lipmanella
conica, Lophocyrtis
aspera, Stylodictya 25H-CC, Y43/0
A, 34, 1229–1243.
Pl.06, fig. 10 Didymocyrtis
bassanii (Carnevale)
RN2 Sanfilippo et al. , 1985
Pl.06, fig. 11Theocamptra
Cladococcus dentata
17H-CC,O45/1
RN4 A.
Nigrini
Moore,
1979 radiolarian) Eocenecamerina,
ovata).(Mast)
Clark,
B. L. and Campbell,
S. (and
1942
Pl.06, fig. 12 Didymocyrtis bassanii (Carnevale)
13H-CC, V12/0
RN5 Sanfilippo et al. , 1985
faunasfromtheMt.
Diabloarea,
California.
Geol.
Pl.06, fig. 13 Didymocyrtis bassanii (Carnevale)
18H-CC, Q37/0
RN4 Sanfilippo et al. , 1985
Acknowledgement
Soc. Amer., Spec.RN5
Pap.,Lazarus
no. 39,Pl.07, fig. 1 Larcopyle hayesi (Chen) group
15H-CC, U42/3
et 1–112.
al. , 2005
Pl.07,
2 Larcopylehayesi (Chen)
104-106 cm,P.Z24/0
LazarusRadiolaria
et al. , 2005 from Central(1978)RN5
Thefig.author
is gratefulto Dr.group
Isao Motoyama for 13H-2,
Dumitrica,
Badenian
Pl.07, fig. 3 Larcopyle hayesi (Chen) group
19H-CC, W15/2
RN3 Lazarus et al. , 2005
reviewing
manuscript,
and togroup
Dr. Noritoshi Suzuki 23H-CC, C26/0
Paratethys. In Brestenska,
E., ed.,Pl.07, fig. 4 theLarcopylehayesi (Chen)
RN2 Lazarus
et al.Chronostratigraphie, 2005and Dr. Kaoru Ogane for useful discussions concerning− 141 −

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Recieved December 19, 2017
Accepted September 18, 2018
Published on-line January 18, 2019

東赤道太平洋における統合国際深海掘削計画（IODP）U1335 地点の下部および中部中新統から産出した放散虫群集
上栗伸一
要 旨
東赤道太平洋における統合国際深海掘削計画（IODP）U1335地点の下部および中部中新統から，78種のSpumellaria亜目および105種のNassellaria亜目を含む合計183種の放散虫化石が産出した．その写真を18枚のプレートに示した．これらの分類群にはActinommidae科，Collosphaeridae科，Hexalonchidae科，Litheliidae科，Pyloniidae科，Spongodiscidae科
およびStylodictyidae科 を含む70の未同定種が含まれており，この中のいくつかは新種である可能性がある．

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Bulletin of the Geological Survey of Japan, vol. 70 (1/2), 2019

Plate 1

2a

1a

1b

2b

4a

3a

3b

7a

100 µmPlate 1

9a

7b

5a

4b

5b

6

8a

8b

10a

10b

9b

Illustrations of the encountered radiolarians. 1: Trisolenia combinata Bjørklund and Goll; 2: Collosphaera macropora Popofsky;
3: Trisolenia sp. A; 4: Siphonosphaera sp. C; 5: Collosphaera pyloma Reynolds; 6: Collosphaera reynoldsi Kamikuri; 7:
Trisolenia megalactis costlowi Bjørklund and Goll; 8, 10: Polysolenia spinosa (Haeckel) group; 9: Polysolenia murrayana(Haeckel).
− 144 −

Early to middle Miocene radiolarians in the tropical Pacific (Kamikuri)

Plate 2

3a

1

2a

4a

7a

6a

6b

5

7b

9a

100 µm

2b

4b

3b

8a

8b

10a

10b

13a

13b

9b

12
11a
Plate 211b

Illustrations of the encountered radiolarians. 1–3: Trisolenia megalactis megalactis Ehrenberg; 4, 5: Siphonosphaera sp. A;
6: Siphonosphaera sp. D; 7: Siphonosphaera sp. B; 8, 9: Collosphaera glebulenta Bjørklund and Goll; 10–12: Collosphaera
sp. A; 13: Collosphaera brattstroemi Bjørklund and Goll.
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Bulletin of the Geological Survey of Japan, vol. 70 (1/2), 2019

Plate 3

3

1a

1b

2a

2b
100 µm4

5a

8aPlate 3

5b

8b

6a

6b

9a

9b

7a

7b

10a

10b

Illustrations of the encountered radiolarians. 1, 2: Axoprunum bispiculum (Popofsky); 3: Stylosphaera communis (Carnevale);4, 10: Stylosphaera sp. B; 5: Amphisphaera santaeannae (Campbell and Clark); 6: Amphistylus polistylus (Carnevale); 7:
Amphisphaera neptunus (Haeckel); 8: Stylosphaera sp. A; 9: Amphistylus angelinus (Campbell and Clark).
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Early to middle Miocene radiolarians in the tropical Pacific (Kamikuri)

Plate 4

1a

1b
2a

3a

2b

3b

4a

4b

5a

5b

6a

6b

7a

7b

8a

8b
9a

9b
100 µm10a
Plate 410b

11Illustrations of the encountered radiolarians. 1, 2, 4: Actinomma robusta (Kling); 3: Thecosphaera sp. A; 5: Thecosphaera
sp. B; 6: Thecosphaera sp. C; 7: Thecosphaera sp. D; 8: Thecosphaera sp. E; 9: Hexacontium sp. B; 10: Hexacontium cf.
arachnoidale Hollande and Enjumet; 11: Saturnalis circularis Haeckel.
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Bulletin of the Geological Survey of Japan, vol. 70 (1/2), 2019

Plate 5

2

1a

4a

1b

3

5a

5b

4b

6

7a

7b

8a

8b

12

13

100 µm9a
Plate 5

9b

10

11

Illustrations of the encountered radiolarians. 1: Cenosphaera sp. A; 2: Heliodiscus sp. B; 3: Hexapyle dodecantha Haeckel; 4:
Excentrodiscus sp. A; 5: Cenosphaera sp. B; 6: Heliodiscus sp. A; 7: Thecosphaera sp. E; 8: Hexacontium sp. C; 9: Axoprunum
sp. A; 10: Didymocyrtis violina (Haeckel); 11: Didymocyrtis tubaria (Haeckel); 12: Didymocyrtis mammifera (Haeckel); 13:
Spongurus cylindricus Haeckel.
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Early to middle Miocene radiolarians in the tropical Pacific (Kamikuri)

Plate 6

2a

2b

1

3a

3b

4a

6

5a

8

9

12

13

5b

10
Plate 674b

11100 µmIllustrations of the encountered radiolarians. 1: Cladococcus sp. A; 2: Hexacontium pachydermum Jørgensen; 3: Hexalonche
sp. A; 4, 5: Didymocyrtis prismatica (Haeckel); 6, 7: Didymocyrtis tubaria (Haeckel); 8: Didymocyrtis laticonus (Riedel); 9:
Didymocyrtis mammifera (Haeckel); 10, 12, 13: Didymocyrtis bassanii (Carnevale); 11: Cladococcus dentata (Mast).
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Plate 7

4

3
1

2

6

5

12
9

10

8

7

11

13

14

15

18
16
Plate 717a

17b

100 µmIllustrations of the encountered radiolarians. 1–4: Larcopyle hayesi (Chen) group; 5: Larcopyle sp. C; 6, 9–11: Lithelius sp. A;
7: Larcopyle polyacantha (Campbell and Clark); 8: Larcopyle sp. B; 12: Lithelius sp. C; 13–15: Larcopyle sp. A; 16: Larcopyletitan (Campbell and Clark); 17: Actinomma sp. B; 18: Larcospira moschkowskii Kruglikova.

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Early to middle Miocene radiolarians in the tropical Pacific (Kamikuri)

Plate 8

3
2

4

1

8
7

6

5

11

100 µm

12

10
9

14
15

13
Plate 8Illustrations of the encountered radiolarians. 1–3: Stylodictya sp. A; 4: Spongurus cylindricus Haeckel; 5, 6, 8: Stylodictya camerina
Campbell and Clark; 7: Stylodictya sp. B; 9: Amphymenium amphistylium Haeckel; 10: Stylodictya sp. C; 11: Porodiscus circularis
Clark and Campbell; 12: Porodiscus sp. A; 13: Heliodiscus sp. A; 14: Phacodiscus sp. A; 15: Dictyocoryne sp. B.
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Bulletin of the Geological Survey of Japan, vol. 70 (1/2), 2019

Plate 93

42

100 µm

1

6

5

10

9

7

11

8

12

14
13
15
Plate 916

Illustrations of the encountered radiolarians. 1: Spongasteriscus marylandicus Martin; 2: Heliodiscus sp. C; 3, 4: Tetrapyle sp.
A; 5: Dictyocoryne malagaense (Campbell and Clark); 6: Spongaster sp. A; 7: Dictyocoryne sp. A; 8: Spongopyle osculosa
Dreyer; 9–12: Spongodiscus resurgens Ehrenberg; 13: Excentrococcus annulatus Dumitrica; 14: Spongodiscus sp. B; 15:
Spongodiscus sp. C; 16: Spongodiscus klingi Caulet.
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Early to middle Miocene radiolarians in the tropical Pacific (Kamikuri)

Plate 10

1a

1b

2a

3a

2b

6a

5a

4

7a

6b

7b

5b
9a

8a

12a

3b

8b

10a

9b

11b

14a

14b

16

17

10b

12b

13a

11a

13b

100 µm15a

15b

Plate 10 Illustrations of the encountered radiolarians. 1, 2: Cyrtocapsella tetrapera Haeckel; 3, 4: Cyrtocapsella japonica (Nakaseko);
5, 6: Cyrtocapsella cornuta Haeckel; 7: Lithopera baueri Sanfilippo and Riedel; 8: Stichocorys delmontensis (Campbell and
Clark); 9: Lithopera renzae Sanfilippo and Riedel; 10: Stichocorys wolffii Haeckel; 11: Lithopera thornburgi Sanfilippo andRiedel; 12: Eucyrtidium sp. A; 13: Stichocorys armata (Haeckel); 14: Phormocyrtis alexandrae O'Connor; 15: Eucyrtidium
sp. C; 16: Eucyrtidium cienkowskii Haeckel; 17: Stichocorys diaphanes (Sanfilippo and Riedel).
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Bulletin of the Geological Survey of Japan, vol. 70 (1/2), 2019

Plate 11

4
2a
1a

2b

1b

5a

6a

5b

9a

10a

9b

13a

13b

17a

17b

11a

10b

14a

18a

6b

3b

7

8a

8b

12a

12b

16a

16b

11b

100 µm

14b

15a

19

18b

3a

15b

20

21a

21b

Plate 11 Illustrations of the encountered radiolarians. 1: Carpocanium cingulatum (Riedel and Sanfilippo); 2, 3: Carpocanium bramlettei (Riedel
and Sanfilippo); 4: Carpacanium sp. B; 5, 6: Carpacanium kinugasense Nishimura; 7: Siphocampe sp. D; 8: Siphocampe grantmackiei
O'Connor; 9: Botryostrobus sp. B; 10: Botryostrobus aquilonaris (Bailey); 11: Spirocyrtis subtilis Petrushevskaya; 12: Siphostichartus
corona (Haeckel); 13: Botryostrobus sp. A; 14: Theocamptra ovata (Haeckel); 15, 16: Siphocampe sp. B; 17, 18: Phormostichoartus
fistula Nigrini; 19: Lithocampana sp. A; 20: Lithocampana sp. B; 21: Cycladophora conica Lombari and Lazarus.
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Early to middle Miocene radiolarians in the tropical Pacific (Kamikuri)

Plate 12

2a
1a

2b

3a

1b

3b
4a

6a

4b

6b

7a

5a

5b

7b

100 µm8a8b

9Plate 12 Illustrations of the encountered radiolarians. 1: Calocycletta caepa Moore; 2: Calocycletta virginis (Haeckel); 3: Calocycletta
serrata Moore; 4: Calocycletta costata (Riedel); 5: Calocycletta robusta Moore; 6: Albatrossidium minzok Sanfilippo andRiedel; 7: Anthocyrtidium sp. A; 8: Anthocyrtidium sp. B; 9: Cyrtolagena sp. A.
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Bulletin of the Geological Survey of Japan, vol. 70 (1/2), 2019

Plate 133

100 µm
1a

1b

4
2a

2b

6a
5a

6b

5b

Plate 13 Illustrations of the encountered radiolarians. 1, 2: Lophocyrtis tanythorax (Sanfilippo and Riedel); 3: Cinclopyramis pacifica(Nakaseko); 4: Cinclopyramis woodringi (Campbell and Clark); 5: Lophocyrtis sp. A; 6: Calocyclura sp. A.

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Early to middle Miocene radiolarians in the tropical Pacific (Kamikuri)

Plate 14

3
2a
1a

2b

1b

5a

4a

6a

5b

6b

4b
7

12

11a

100 µm

9

8

13

14

10

15a

15b

11b

Plate 14 Illustrations of the encountered radiolarians. 1: Lamprocyclas maritalis Haeckel; 2: Lophocyrtis aspera (Ehrenberg); 3:
Theocorys spongoconus Kling; 4: Lamprocyclas sp. B; 5: Theocorys sp. A; 6: Clathrocanium atreta Sanfilippo and Riedel; 7,
8: Lithomelissa sp. A; 9: Lophophaena sp. A; 10: Ceratocyrtis sp. A; 11: Lamprocyclas margatensis Campbell and Clark var.
A; 12: Lipmanella conica Petrushevskaya; 13, 14: Lophophaena tekopua O'Connor; 15: Clathrocanium coarctatum Ehrenberg.

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Bulletin of the Geological Survey of Japan, vol. 70 (1/2), 2019

Plate 1521a1b100 µm

4a
3a

4b

3b

6a
5a

6b

5b

Plate 15 Illustrations of the encountered radiolarians. 1, 2: Pterocanium audax (Riedel); 3: Lychnocanoma elongata (Vinassa de Regny);
4: Pterocanium aff. tridentatum (Ehrenberg); 5: Pterocanium charybdeum (Müller); 6: Pterocanium sp. A.
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Early to middle Miocene radiolarians in the tropical Pacific (Kamikuri)

Plate 16
207

1a

215

1b

3

2a

2b

5a
4a

4b

5b
7

6a

6b

8a

100 µm8b

9Plate 16 Illustrations of the encountered radiolarians. 1–3: Lychnocanoma sp. A; 4, 5: Lychnocanoma nodosum (Haeckel); 6: Dictyophimus
splendens (Campbell and Clark); 7: Dictyophimus sp. A; 8, 9: Valkyria pukapuka O'Connor.

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Bulletin of the Geological Survey of Japan, vol. 70 (1/2), 2019

Plate 17

1

2

5a

4a

4b

5b

3

6a

6b

7

100 µmPlate 17 Illustrations of the encountered radiolarians. 1: Dorcadospyris dentata Haeckel; 2: Dorcadospyris alata (Riedel); 3: Dorcadospyris
ateuchus (Ehrenberg); 4–6: Lamprocyclas margatensis Campbell and Clark var. B; 7: Cornutella trochus Ehrenberg.

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Early to middle Miocene radiolarians in the tropical Pacific (Kamikuri)

Plate 18

2

1

8

9

7

6

4

3

5

10
11

14

13

12

17

15

16

18

23
22
21
20

19

24

100 µm

25

26

27

28

29

30

Plate 18 Illustrations of the encountered radiolarians. 1, 2: Tympanomma binoctonum (Haeckel); 3, 4: Tholospyris anthophora (Haeckel);
5: Dendrospyris pannosa Goll; 6: Tholospyris mammillaris (Haeckel); 7: Tholospyris kantiana (Haeckel); 8: Tympanomma
tuberosum (Haeckel); 9: Liriospyris mutuaria Goll; 10: Liriospyris sp. A; 11: Dendrospyris sp. A; 12: Eucoronis octopylus(Haeckel); 13, 14: Eucoronis perspicillum Haeckel; 15: Acanthodesmia circumflexa (Goll); 16: Acanthodesmia sp. A; 17:
Acanthodesmia sp. B; 18: Dendrospyris pododendros (Carnevale); 19: Giraffospyris annulispina Goll; 20: Eucoronis toxarium(Haeckel); 21: Phormospyris stabilis (Goll); 22: Phormospyris sp. B; 23: Liriospyris parkerae Riedel and Sanfilippo; 24:
Liriospyris stauropora (Haeckel); 25, 26: Zygocircus sp.; 27: Botryocyrtis sp. A; 28: Centrobotrys thermophila Petrushevskaya;
29: Centrobotrys petrushevskayae Sanfilippo and Riedel; 30: Acrobotrys disolenia Haeckel.
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