Biodiversity and Biogeography of the Kuril Islands and Sakhalin (2004) 1, 49–80.

Caddisflies (Trichoptera) of the Kuril Archipelago
Noboru Minakawa1§, Tatyana I. Arefina2, Tomiko Ito3, Takao Nozaki4,
Naotoshi Kuhara5, Hiroyuki Nishimoto6, Makoto Uenishi7, Valentina A.
Teslenko2, Daniel J. Bennett8, Robert I. Gara9, Kemper L. Kurowski10,
Pontus B. H. Oberg10, Todd I. Ritchie11 and Lucie J. Weis9
1

Department of Medicine, Saga University, Saga, Japan; 2 The Institute of Biology and Soil
Sciences, Russian Academy of Science, Vladivostok, Russia; 3 Hokkaido Fish Hatchery, Eniwa,
Hokkaido, Japan; 4 Kanagawa Environmental Research Center, Hiratsuka, Kanagawa, Japan;
5
Chitose Board of Education, Chitose, Hokkaido, Japan; 6 1-71-1 Hikarigaoka, Komaki, Aichi,
Japan; 7 Kowata, Okurayama 39-770, Uji, Kyoto, Japan; 8 Department of Ecology and
Evolutionary Biology, University of Kansas, Lawrence, Kansas, USA; 9 College of Forest
Resources, University of Washington, Seattle, Washington, USA; 10 Department of Zoology,
University of Washington, Seattle Washington, USA; 11 School of Aquatic and Fishery Sciences,
University of Washington, Seattle, Washington, USA.

Abstract
We report the complete data of the caddisfly specimens collected during theseven annual expeditions of the International Kuril Island Project, and provide a list of caddisfly
species known from the Kuril Islands. Caddisflies were collected from 21 out of 30 majorislands, and collected from 14 islands for the first time. A total of 19 families, 45 genera and98 species were collected during the expeditions, including 28 new species distribution records
for the archipelago. These records bring the present total numbers to 20 families, 50 generaand 123 species. Glossosoma inops is placed in the junior subjective synonym of G. ussuricum.

Introduction

Despite the long history of research in the Kuril
Islands, the caddisfly fauna of the central Kurils has notbeen reported except for Simushir. This is mainly due tothe remoteness of the islands. During the International
Kuril Island Project (IKIP; 1994–2000), American,
Japanese and Russian scientists surveyed 30 islands
including eight central islands (Pietsch et al. 2001).
Including new data from the earlier IKIP expeditions,over 100 caddisfly species were recorded from the Kuril
Islands (Arefina 1997a, 1997b, 1997c, 1997d, 1997e,
1997f, 1997g; Arefina and Levanidova 1997; Ito 1997;
Levanidova and Arefina 1997a, 1997b; Nimmo et al.
1997; Arefina et al. 1999; Arefina 2001). In this paper,
we report the complete data of the caddisfly specimens
collected during the seven annual expeditions of IKIP,
and provide a list of caddisfly species known from theKuril Islands.

The caddisfly fauna of the Kuril Islands was firstreported for Shumshu, Paramushir and Iturup in 1930’s(Uéno 1933; Miyadi 1933, 1937). These reports werelimited to immature stages, and species level
identification was not available except for Praecosmoecus
kamtshaticus Martynov (present name: Ecclisomyia
kamtshatica) collected from Shumshu (Schmid 1949).
For adult caddisflies, Kuwayama (1936) reported eightspecies from Shumshu, Paramushir, Urup, Iturup andKunashir. Later Tsuda (1939, 1942) added four species
from Shumshu, Paramushir and Kunashir.
In the 1950’s and 1960’s, Schmid (1952, 1953, 1955,
1964, 1965) and Kuwayama (1967a, 1967b, 1973) added
considerably to the knowledge of the Kuril caddisfly
fauna. In the 1970’s and 1980’s, Russian scientists
actively studied the caddisflies of the Kuril Islands(Levanidova and Schmid 1977; Levanidova 1979, 1980,
1982, 1986, 1989; Vshivkova 1986; Botosaneanu 1988;
Botosaneanu and Levanidova 1988; Arefina et al. 2003).
Since the collapse of the Soviet Union, Russian andJapanese scientists have collaborated closely in studying
caddisfly specimens from the Kurils (Ito et al. 1992;
Vshivkova et al. 1994; Levanidova et al. 1995; Kuranishi
2000).

Methods
Sweep nets and canvas beating sheets were mostly
used for collecting adult caddisflies during the
expeditions. Canvas beating sheets were effective insmall streams on the central and northern islands when
the air temperature was low. Although the beating sheets
became wet and heavy on rainy or foggy days, they werestill more effective than sweep nets. On sunny days,
however, adult caddisflies became too active to capture
on beating sheets, and then sweep nets became a bettercollecting method. In swamps with short grass, sweep§

Correspondence: Noboru Minakawa
Mailing Address: Department of Medicine, Saga University,
Saga, Japan
E-mail: minakawa@post.saga-med.ac.jp

49

IKIP records: Simushir*, Urup, Iturup, Kunashir,
Shikotan*, Zelionyi* (Arefina 1997b; Arefina 2001).
5. Rhyacophila kawamurae Tsuda, 1940*; Kunashir(Vshivkova et al. 1994).
IKIP records: Iturup*, Kunashir (Arefina 2001).
6. Rhyacophila lata Martynov, 1918; southern Kurils(Levanidova 1980).
7. Rhyacophila mirabilis Levanidova and Schmid, 1977*;
Urup, Iturup, Kunashir (Levanidova and Schmid
1977; Vshivkova et al. 1994).
IKIP records: Ketoi *, Simushir *, Urup, Iturup,
Kunashir (Arefina 2001).
8. Rhyacophila nipponica Navás, 1933 * ; Kunashir(Vshivkova et al. 1994).
IKIP records: Kunashir, Shikotan *, Zelionyi *(Arefina 1997b; Arefina 2001).
9. Rhyacophila retracta Martynov, 1914 *; Iturup,
Kunashir (Levanidova 1986; Vshivkova et al.
1994).
IKIP records: Urup*, Iturup, Kunashir.
10. Rhyacophila transquilla Tsuda, 1940*; Kunashir(Levanidova 1986).
IKIP records: Kunashir.

nets were used. Sweeping was also used for tree branches
along streams on the southern islands where airtemperature was higher. Light traps and Malaise traps
were used when the collectors were able to return to thetraps next day. Caddisfly larvae were collected by kicknets and dip nets. Some newly emerged caddisflies werecollected by turning rocks in water.
Collected caddisflies were immediately preservedin 80% ethanol in the field. Collections were identified
by T. I. Arefina, T. Ito, V. D. Ivanov, N. Kuhara, I. M.
Levanidova, H. Nishimoto, T. Nozaki and M. Uenishi.
The caddisfly materials were deposited in the Instituteof Biology and Soil Sciences, Russian Academy of
Sciences, Far Eastern Branch, Vladivostok, Russia, theCalifornia Academy of Sciences, San Francisco,California, USA and Clemson University, Clemson, South
Carolina, USA.

Results
Species record
Caddisflies were collected from 21 out of 30 majorislands, and collected from 14 islands for the first time
(Table 1). A total of 19 families, 45 genera and 98 species
were collected during the expeditions, including 28 newly
recorded species (including three undescribed species)
for the archipelago (Arefina 1997a; Arefina et al. 1999).
These records bring the present total numbers to 20
families, 50 genera and 123 species.
Following is a list of the caddisflies recorded fromthe Kuril Islands. Genera and species with an asterisk(*) are those whose presence on the Kuril Islands wasverified by our collections; Kuril Island occurrences forthe remaining taxa are based on a survey of the available
scientific literature. An island name with an asterisk (*)
indicates that the caddisfly species was recorded fromthe island for the first time. Taxa with two asterisks (**)
were recorded from the Kuril Island for the first time.

2. FAMILY HYDROBIOSIDAE ULMER, 1905(2) Genus Apsilochorema Ulmer, 1907*
11. Apsilochorema sutshanum Martynov, 1934*; Iturup(Levanidova 1986).
IKIP records: Iturup, Kunashir * , Shikotan *(Levanidova and Arefina 1997a).
3. FAMILY GLOSSOSOMATIDAE WALLENGREN, 1891
(3) Genus Agapetus Curtis, 1834
12. Agapetus inaequispinosus Schmid, 1970; Kunashir(Levanidova 1989; Vshivkova et al. 1994).
(4) Genus Anagapetus Ross, 1938*
13. Anagapetus schmidi (Levanidova, 1979)*; Kunashir(Levanidova 1989).
IKIP records: Ketoi *, Simushir*, Urup*, Iturup*,
Kunashir, Shikotan * (Arefina and Levanidova
1997).

1. FAMILY RHYACOPHILIDAE STEPHENS, 1836
(1) Genus Rhyacophila Pictet, 1834*
1. Rhyacophila arefini Lukyanchenko, 1993*; Iturup,
Kunashir, Shikotan (Kuwayama 1967a; Vshivkova
et al. 1994).
IKIP records: Urup*, Iturup, Kunashir, Shikotan(Arefina 2001).
2. Rhyacophila brevicephala Iwata, 1927*; Kunashir(Levanidova 1980).
IKIP record: Iturup*, Kunashir, Shikotan* (Arefina2001).
3. Rhyacophila coreana Tsuda, 1940; Iturup (Levanidova
1986).
Remarks: Levanidova (1986) recorded this species
without precise data.
4. Rhyacophila hokkaidensis Iwata, 1927*; Urup, Iturup,
Kunashir (Kuwayama 1967a; Levanidova 1986;
Vshivkova et al. 1994; Arefina 1997b).

(5) Genus Glossosoma Curtis, 1834*
14. Glossosoma altaicum (Martynov, 1914); Kunashir,(Levanidova 1989; Vshivkova et al. 1994).
15. Glossosoma dulkejti (Martynov, 1934) *; Iturup,
Kunashir (Vshivkova 1986).
IKIP records: Iturup, Kunashir, Shikotan* (Arefina
and Levanidova 1997).
16. Glossosoma intermedium (Klapálek, 1892) *;Paramushir (Kuranishi 2000).
IKIP records: Shumshu*, Paramushir.
17. Glossosoma ussuricum (Martynov, 1934)*; Iturup,
Kunashir (Vshivkova 1986; Vshivkova et al. 1994).
IKIP records: Simushir*, Urup*, Iturup, Kunashir,
Shikotan* (Arefina and Levanidova 1997).
See also “synonymic note” (p. 56).
50

Table 1. Numbers of caddisfly species recorded from the Kuril Islands.
IslandPrevious records

New records

Total

IKIP records

Kuril Islands

95

28

123

98

Northern Group:
Atlasova
ShumshuParamushir
Antsiferova
Makanrushi
Onekotan
Kharimkotan
Chirinkotan
EkarmaShiashkotan

0
16
24
0
0
0
0
0
0
0

4
11
8
0
5
9
7
0
2
8

4
27
32
0
5
9
7
0
2
8

4
19
26
0
5
9
7
0
2
8

Central Group:
Lovushki
RaikokeMatua
Rasshua
Ryponkicha
Yankicha
Ketoi
Simushir

0
0
0
0
0
0
0
5

0
0
4
8
0
0
14
11

0
0
4
8
0
0
14
16

0
0
4
8
0
0
14
15

Southern Group:
Broutona
ChirpoiBrat Chirpoev
Urup
Iturup
Kunashir
Shikotan

0
0
0
14
34
66
11

0
0
0
19
26
21
23

0
0
0
33
60
87
34

0
0
0
32
53
59
29

Habomai Group:
Polonskogo
Zelionyi
Iurii
Anuchina
Tanfilyeva

0
0
0
0
0

2
10
1
1
9

2
10
1
1
9

2
10
1
1
9

The genus Glossosoma was also recorded from
Ketoi* for the first time (Appendix I).

(9) Genus Stactobia McLachlan, 1880
21. Stactobia makartschenkoi Botosaneanu andLevanidova, 1988; Kunashir (Botosaneanu andLevanidova 1988).

4. FAMILY HYDROPTILIDAE STEPHENS, 1836
(6) Genus Hydroptila Dalman, 1819*/**18. Hydroptila sp.*/**IKIP records: Iturup*, Polonskogo*.

5. FAMILY PHILOPOTAMIDAE STEPHENS, 1829
(10) Genus Dolophilodes Ulmer, 1909*
22. Dolophilodes (Dolophilodes) japonicus (Banks,1906)*; Kunashir (Levanidova 1982; Vshivkova
et al. 1994).
IKIP records: Kunashir.
23. Dolophilodes (Dolophilodes) kunashirensis
Ivanov, 1996*; Kunashir (Arefina et al. 1996).
IKIP records: Paramushir*.
24. Dolophilodes (Dolophilodes) nomugiensis(Kobayashi 1980)*; Kunashir (Vshivkova et al.
1994).
IKIP records: Shumshu * , Kharimkotan * ,(7) Genus Orthotrichia Eaton, 1873*/**
19. Orthotrichia sp.*/**IKIP records: Zelionyi* (Arefina et al. 1999).
(8) Genus Palaeagapetus Ulmer, 1912*/**
20. Palaeagapetus flexus Ito, 1991*/**IKIP records: Iturup* (Arefina 1997c).
The genus Palaeagapetus was also recorded from
Urup* for the first time (Appendix I).
51

Simushir*, Urup*, Iturup*.

1999).

The genus Dolophilodes was also recorded from
Ketoi* and Shiashkotan* for the first time (Appendix I).

10. FAMILY POLYCENTROPODIDAE ULMER, 1903
(18) Genus Plectrocnemia Stephens, 1836
36. Plectrocnemia levanidovae Vshivkova et al., 2003;
Kunashir (Vshivkova et al. 1994; Arefina et al.
2003).

(11) Genus Kisaura Ross, 1956*
25. Kisaura borealis (Kuhara, 1999)*/**IKIP records: Kunashir*.
26. Kisaura hattorii (Kuhara, 1999)*/**IKIP records: Iturup*, Kunashir*, Shikotan*.
27. Kisaura tsudai (Botosaneanu, 1970); Kunashir(Ivanov, 1997).
Remarks: Arefina et al. (1999) misidentified K.
hattorii as K. tsudai.
28. Kisaura sp.*/**IKIP records: Kunashir*.
Remarks: This species was different from any otherspecies. N. Kuhara and T. I. Arefina will describe
this as a new species.

11. FAMILY ECNOMIDAE ULMER, 1903
(19) Genus Ecnomus McLachlan, 1864*
37. Ecnomus tenellus (Rambur, 1842) * ; Kunashir(Vshivkova et al. 1994).
IKIP records: Iturup*, Kunashir.
12. FAMILY PHRYGANEIDAE LEACH, 1815(20) Genus Agrypnia Curtis, 1835*
38. Agrypnia acristata Wiggins, 1998 * ; Kunashir(Kuwayama 1967a; Wiggins 1998).
IKIP records: Iturup*.
Remarks: Agrypnia ulmeri by Kuwayama (1967a,
1973) is likely misidentification of this species.
39. Agrypnia czerskyi (Martynov, 1924)*/**IKIP records: Iturup*.
40. Agrypnia picta Kolenati, 1848*; Urup, Paramushir(Schmid 1965; Kuranishi 2000).
IKIP records: Shumshu*, Paramushir, Shiashkotan*,
Rasshua*, Iturup*.
41. Agrypnia sahlbergi (McLachlan, 1880)*; Paramushir(Vshivkova et al. 1994).
IKIP records: Shumshu*, Paramushir.
42. Agrypnia sordida (McLachlan, 1871) *; Iturup(Vshivkova et al. 1994; Wiggins 1998).
IKIP records: Urup*, Iturup, Kunashir* (Arefina
1997d).

(12) Genus Wormaldia McLachlan, 1865*
29. Wormaldia sp.*/**IKIP records: Kunashir* (Arefina et al. 1999).
6. FAMILY STENOPSYCHIDAE MARTYNOV, 1924(13) Genus Stenopsyche McLachlan, 1866*
30. Stenopsyche marmorata Navás, 1920 *; Iturup,
Kunashir (Kuwayama 1967a).
IKIP records: Iturup, Kunashir, Shikotan*.
7. FAMILY HYDROPSYCHIDAE CURTIS, 1835
(14) Genus Cheumatopsyche Wallengren, 1891*
31. Cheumatopsyche infascia Martynov, 1934*/**
IKIP records: Paramushir*, Kunashir* (Arefina etal. 1999).
Schmid (1965) also recorded a female of this genus
from Iturup.

(21) Genus Eubasilissa Martynov, 1930*
43. Eubasilissa regina (McLachlan, 1871)*; Kunashir(Schmid 1965).
IKIP records: Kunashir.

(15) Genus Hydropsyche Pictet, 1834*
32. Hydropsyche albicephala Tanida, 1986*/**
IKIP records: Kunashir* (Arefina et al. 1999).
33. Hydropsyche orientalis Martynov, 1934*; Kunashir(Vshivkova et al. 1994).
IKIP records: Iturup*, Kunashir.

(22) Genus Hagenella Martynov, 1924*
44. Hagenella apicalis (Matsumura, 1904)*; Kunashir(Schmid 1965).
IKIP records: Kunashir, Polonskogo*.
45. Hagenella sibirica (Martynov, 1909)*; Shumshu,
Paramushir (Kuwayama 1973; Kuranishi 2000).
IKIP records: Paramushir.

8. FAMILY ARCTOPSYCHIDAE MARTYNOV, 1924(16) Genus Parapsyche Betten, 1934*34. Parapsyche shikotsuensis (Iwata, 1927)*; Kunashir(Vshivkova et al. 1994).
IKIP records: Kunashir.

(23) Genus Oligotricha Rambur, 1842*
46. Oligotricha hybridoides Wiggins and Kuwayama,
1971; Kunashir (Vshivkova et al. 1994).
47. Oligotricha lapponica (Hagen, 1864)*/**IKIP records: Paramushir*, Onekotan* (Arefina etal. 1999).

9. FAMILY PSYCHOMYIIDAE CURTIS, 1835**(17) Genus Lype McLachlan, 1879*/**35. Lype excisa Mey, 1991*/**IKIP records: Iturup*, Kunashir* (Arefina et al.

The genus Oligotricha was also recorded from Urup*52

and Shumshu* for the first time (Appendix I).

IKIP records: Paramushir.
59. Grammotaulius signatipennis McLachlan, 1876*;
Paramushir (Kuranishi 2000).
IKIP records: Shumshu*, Paramushir, Onekotan*.

(24) Genus Phryganea Linnaeus, 1758
48. Phryganea (Colpomera) japonica (McLachlan,1866); Kunashir (Schmid 1965).

(33) Genus Halesus Stephens, 1836*
60. Halesus sachalinensis Martynov, 1914*; Urup, Iturup,
Kunashir (Schmid 1965; Vshivkova et al. 1994).
IKIP records: Urup, Iturup, Kunashir, Shikotan*.

(25) Genus Semblis Fabricius, 1775
49. Semblis melaleuca (McLachlan, 1871); Iturup,
Kunashir (Schmid 1965; Kuwayama 1967a;
Vshivkova et al. 1994; Wiggins 1998).

(34) Genus Hydatophylax Wallengren, 1891*
61. Hydatophylax festivus (Navás, 1920)*; Urup, Iturup,
Kunashir, Shikotan (Vshivkova et al. 1994; Nozaki,1999).
IKIP records: Urup, Iturup, Kunashir, Shikotan,
Zelionyi*.
62. Hydatophylax magnus (Martynov, 1914); Iturup(Vshivkova et al. 1994).
63. Hydatophylax variabilis (Martynov, 1910)*; Kunashir(Vshivkova et al. 1994).
IKIP records: Kunashir, Shikotan*.
64. Hydatophylax sp.*/**IKIP records: Ketoi *, Simushir*, Urup*, Iturup*,
Kunashir*, Shikotan* (Arefina et al. 1999).
Remarks: Arefina et al. (1999) reported this species
as H. soldatovi, however we confirmed that this isdifferent from any other species after carefulreexamination. T. Nozaki and N. Minakawa will
describe this as a new species.

13. FAMILY BRACHYCENTRIDAE ULMER, 1903
(26) Genus Brachycentrus Curtis, 1834*
50. Brachycentrus americanus (Banks, 1899)*/**IKIP records: Kunashir* (Arefina et al. 1999).
51. Brachycentrus sp.*
IKIP records: Shumshu.
Remarks: This species is not B. americanus. See
also Uéno (1933).
(27) Genus Micrasema McLachlan, 1876*
52. Micrasema hanasensis Tsuda, 1942*/**
IKIP records: Ketoi*, Urup*, Iturup*, Kunashir*.
53. Micrasema (gelidum) kurilicum Botosaneanu, 1988;
Kunashir (Botosaneanu 1988).
The genus Micrasema was also recorded from
Simushir* for the first time (Appendix I).

The genus Hydatophylax was also recorded from
Paramushir* for the first time (Appendix I).

14. FAMILY LIMNEPHILIDAE KOLENATI, 1848(28) Genus Asynarchus McLachlan, 1880*
54. Asynarchus sachalinensis Martynov, 1914*; Iturup,
Kunashir (Kuwayama 1967a; Vshivkova et al.
1994).
IKIP records: Urup*, Iturup, Shikotan*.

(35) Genus Lenarchus Martynov, 1914*
65. Lenarchus fuscostramineus Schmid, 1952*; Iturup,
Kunashir (Schmid 1965).
IKIP records: Urup*, Iturup, Shikotan*, Tanfilyeva*.
(36) Genus Limnephilus Leach, 1815*
66. Limnephilus alienus Martynov, 1914*; Shumshu,
Simushir, Urup, Iturup, Kunashir (Schmid 1965;
Vshivkova et al. 1994).
IKIP records: Shumshu, Onekotan*, Shiashkotan*,
Rasshua*, Urup, Iturup, Kunashir.
67. Limnephilus diphyes McLachlan, 1880; Shumshu,
Paramushir (Kuranishi 2000).
68. Limnephilus elegans Curtis, 1834 * ; Shumshu,
Paramushir (Vshivkova et al. 1994; Kuranishi
2000).
IKIP records: Paramushir.
69. Limnephilus femoralis Kirby, 1837 *; Shumshu,
Paramushir (Vshivkova et al. 1994; Kuranishi
2000).
IKIP records: Shumshu, Paramushir.
70. Limnephilus femoratus Zetterstedt, 1840*/**IKIP records: Iturup* (Arefina et al. 1999).
Remarks: The previous record of this species from
Paramushir was an error (Arefina et al. 1999).
71. Limnephilus fenestratus (Zetterstedt, 1840) * ;Paramushir (Vshivkova et al. 1994).
IKIP records: Shumshu*, Paramushir.

*(29) Genus Dicosmoecus McLachlan, 1875
55. Dicosmoecus jozankeanus (Matsumura, 1931) *;Iturup, Shikotan (Kuwayama 1967a; Nagayasu andIto 1993).
IKIP records: Urup*, Iturup, Kunashir*, Shikotan(Nimmo et al., 1997).
(30) Genus Ecclisocosmoecus Schmid, 1964*/**
56. Ecclisocosmoecus spinosus Schmid, 1964*/**
IKIP records: Urup*, Iturup*, Kunashir*, Shikotan*,Tanfilyeva*.
(31) Genus Ecclisomyia Banks, 1907*
57. Ecclisomyia kamtshatica (Martynov, 1913) *;Shumshu (recorded as Simushir), Paramushir (Uéno
1933).
IKIP records: Shumshu, Paramushir, Ketoi * ,Simushir*, Urup*, Iturup* (Nimmo et al. 1997).
(32) Genus Grammotaulius Kolentai, 1848*
58. Grammotaulius inornatus Schmid, 1964*; Paramushir(Kuwayama 1936; Vshivkova et al. 1994).
53

72. Limnephilus fuscovittatus Matsumura, 1904 * ;Shumshu, Paramushir, Simushir, Urup, Iturup,
Kunashir, Shikotan (Schmid 1965; Kuwayama
1967a; Vshivkova et al. 1994).
IKIP records: Shumshu, Atlasova*, Paramushir,
Onekotan *, Kharimkotan *, Matua *, Rasshua * ,Ketoi*, Simushir, Urup, Iturup, Kunashir, Shikotan,
Tanfilyeva*.
73. Limnephilus incisus Curtis, 1834*/**
IKIP records: Paramushir* (Arefina et al. 1999).
74. Limnephilus major (Martynov, 1909)*; Paramushir(Vshivkova et al. 1994).
IKIP records: Shumshu*.
75. Limnephilus nigriceps (Zetterstedt, 1840)*/**IKIP records: Paramushir*.
76. Limnephilus nipponicus Schmid, 1964*; Simushir,
Urup, Iturup, Kunashir (Schmid 1965; Kuwayama
1967a; Vshivkova et al. 1994).
IKIP records: Ketoi *, Simushir, Urup, Iturup,
Kunashir, Shikotan*.
77. Limnephilus orientalis Martynov, 1935*; Urup, Iturup,
Kunashir, Shikotan (Schmid 1965; Kuwayama
1967a; Vshivkova et al. 1994).
IKIP records: Urup, Iturup, Kunashir.
78. Limnephilus ornatulus Schmid, 1965 *; Iturup,
Kunashir (Schmid 1965; Kuwayama 1967a;
Vshivkova et al. 1994).
IKIP records: Kunashir.
79. Limnephilus picturatus McLachlan, 1875*; Shumshu,
Paramushir (Tsuda 1942; Schmid 1955; Vshivkova
et al. 1994).
IKIP records: Shumshu, Paramushir.
80. Limnephilus quadratus Martynov, 1914*; Kunashir(Vshivkova et al. 1994).
IKIP records: Kunashir, Tanfilyeva*, Iurii*.
81. Limnephilus rhombicus (Linnaeus, 1758) * ;Paramushir, Kunashir (Konakov 1956; Vshivkova
et al. 1994).
IKIP records: Iturup*.
82. Limnephilus sericeus (Say, 1824) *; Shumshu,
Paramushir, Simshir, Urup, Iturup (Tsuda 1942;
Schmid 1955, 1965; Kuwayama 1967a; Vshivkova
et al. 1994).
IKIP records: Shumshu, Paramushir, Makanrushi*,
Onekotan*, Kharimkotan*, Shiashkotan*, Matua*,
Rasshua*, Ketoi*, Simushir, Urup, Iturup, Shikotan*.
83. Limnephilus sparsus Curtis, 1834 *; Paramushir,
Simushir, Urup, Iturup, Kunashir, Shikotan (Tsuda
1942; Schmid 1965; Kuwayama 1967a; Vshivkova
et al. 1994).
IKIP records: Atlasova*, Shumshu, Paramushir,M a k a n r u s h i *, O n e k o t a n *, K h a r i m k o t a n *,Shiashkotan*, Matua*, Rasshua*, Ketoi*, Simushir,
Urup, Iturup, Kunashir, Shikotan, Zelionyi * ,Tanfilyeva*.
84. Limnephilus stigma Curtis, 1834 *; Paramushir,
Kunashir (Kuwayama 1967a; Vshivkova et al.
1994).
IKIP records: Shumshu*, Paramushir, Onekotan*,
Kharimkotan*, Iturup*.

85. Limnephilus subcentralis Brauer, 1857*/**
IKIP records: Iturup*.
(37) Genus Nemotaulius Banks, 1906*
86. Nemotaulius admorsus (McLaclan, 1866); Kunashir(Kuwayama 1967a).
87. Nemotaulius miyakei (Nakahara, 1914)*; Iturup,
Kunashir (Kuwayama 1936; Kuwayama 1967a).
IKIP records: Iturup, Kunashir.
88. Nemotaulius mutatus (McLachlan, 1872); Shumshu,
Paramushir (Kuranishi 2000).
(38) Genus Nothopsyche Banks, 1906*/**
89. Nothopsyche sp.*/**IKIP records: Kunashir* (Arefina et al. 1999).
(39) Genus Onocosmoecus Banks, 1943*
90. Onocosmoecus unicolor (Banks, 1897)*; Shumshu,
Shikotan (Kuwayama 1936; Vshivkova et al. 1994).
IKIP records: Atlasova*, Shumshu, Paramushir*,Makanrushi*, Onekotan*, Kharimkotan*, Ekarma*,
Shiashkotan*, Rasshua*, Ketoi*.
15. FAMILY APATANIIDAE WALLENGREN, 1886(40) Genus Allomyia Banks, 1916*
91. Allomyia coronae Levanidova and Arefina, 1995*;
Kunashir (Levanidova et al. 1995).
IKIP records: Iturup*, Shikotan* (Levanidova andArefina 1997b).
92. Allomyia delicatula Levanidova and Arefina, 1995*;
Kunashir (Levanidova et al. 1995).
IKIP records: Simushir*.
(41) Genus Apatania Kolenati, 1848*
93. Apatania aberrans (Martynov, 1933); Kunashir(Vshivkova et al. 1994).
94. Apatania insularis Levanidova, 1979; Kunashir(Levanidova 1979).
95. Apatania parvula (Martynov, 1935)*; Shumshu, Urup,
Iturup, Kunashir, Shikotan (Schmid 1965;
Kuwayama 1967a; Levanidova 1979; Vshivkova etal. 1994).
IKIP records: Rasshua*, Ketoi*, Simushir*, Urup,
Iturup, Kunashir, Shikotan, Anuchina* (Levanidova
and Arefina 1997b).
96. Apatania sinensis Martynov, 1914; Paramushir(Levanidova 1982).
97. Apatania stigmatella (Zetterstedt, 1840); Shumshu,
Paramushir (Kuranishi 2000).
98. Apatania zonella Zetterstedt, 1840 *; Shumshu,
Paramushir (Vshivkova et al. 1994).
IKIP records: Atlasova*, Shumshu, Paramushir,M a k a n r u s h i *, O n e k o t a n *, K h a r i m k o t a n *,S h i a s h k o t a n *, M a t u a *, R a s s h u a *, I t u r u p *(Levanidova and Arefina 1997b).
The genus Apatania was also collected from
Ekarma* for the first time (Appendix I).

54

111. Molanna submarginalis McLachlan, 1872 * ;Paramushir (Vshivkova et al. 1994).
IKIP records: Paramushir.

16. FAMILY GOERIDAE ULMER, 1903(42) Genus Goera Stephens, 1829*
99. Goera japonica Banks, 1906*; Urup, Iturup, Kunashir(Schmid 1965; Kuwayama 1967a; Vshivkova et al.
1994).
IKIP records: Urup, Iturup, Kunashir, Shikotan*(Arefina 1997e).
100. Goera sp.; Shikotan (Vshivkova et al. 1994).
Remarks: This species differs from the other species
of Goera recorded from Russia and Japan(Vshivkova et al. 1994).

The genus Molanna was also recorded from
Shumshu* for the first time (Appendix I).
(46) Genus Molannodes McLachlan, 1866*
112. Molannodes itoae Fuller and Wiggins, 1987;
Kunashir (Vshivkova et al. 1994).
113. Molannodes tinctus (Zetterstedt, 1840)*/**IKIP records: Shumshu*, Paramushir* (Arefina etal. 1999).

17. FAMILY UENOIDAE IWATA, 1927
20. FAMILY LEPTOCERIDAE LEACH, 1815
(43) Genus Neophylax McLachlan, 1871*
101. Neophylax japonicus Schmid, 1964*/**
IKIP records: Kunashir*.
102. Neophylax ussuriensis (Martynov, 1914)*; Urup,
Iturup, Kunashir (Kuwayama 1936).
IKIP records: Urup, Iturup, Kunashir, Shikotan*(Arefina 1997f).

(47) Genus Ceraclea Stephens, 1829*/**114. Ceraclea alboguttata (Hagen, 1860)*/**IKIP records: Kunashir* (Arefina et al. 1999).
115. Ceraclea valentinae Arefina, 1997*/**
IKIP records: Zelionyi* (Arefina 1997a).
(48) Genus Mystacides Berthold, 1827*
116. Mystacides azureus (Linnaeus, 1761)*/**IKIP records: Iturup*, Kunashir* (Arefina et al.
1999).
117. Mystacides pacificus Mey, 1991*; Iturup, Kunashir(Vshivkova et al. 1994).
IKIP records: Urup*, Iturup, Kunashir, Zelionyi*,
Tanfilyeva*.

18. FAMILY LEPIDOSTOMATIDAE ULMER, 1903
(44) Genus Lepidostoma Rambur, 1842*
103. Lepidostoma albardanum (Ulmer, 1906); Kunashir(Ito et al. 1992; Weaver 2002).
104. Lepidostoma complicatum (Kobayashi, 1968)*;
Kunashir (Ito et al. 1992; Weaver 2002).
IKIP records: Iturup*, Kunashir, Shikotan*.
105. Lepidostoma crassicorne (Ulmer, 1907)*; Kunashir(Ito et al. 1992; Weaver 2002).
IKIP records: Urup*, Shikotan*.
Remarks: Vshivkova et al. (1994) suggested that
Neoseverinia sp. recorded from Shikotan by
Kuwayama (1967a) was this species. We confirmedthat L. crassicorne occurs on Shikotan.
106. Lepidostoma hiurai (Tani, 1971)*; Iturup, Kunashir(Ito et al. 1992; Weaver 2002).
IKIP records: Iturup, Kunashir, Shikotan*, Zelionyi*,
Tanfilyeva* (Ito 1997).
107. Lepidostoma naraense (Tani, 1971); Kunashir (Itoet al. 1992; Weaver 2002).
108. Lepidostoma satoi (Kobayashi, 1968); Shikotan,
Kunashir (Ito et al. 1992; Weaver 2002).
109. Lepidostoma stellatum (Ito, 1984)*; Kunashir (Itoet al. 1992; Weaver 2002).
IKIP records : Paramushir * , Makanrushi * ,Shiashkotan*, Ketoi*, Simushir*, Urup*, Iturup* (Itoand Minakawa 1995).

(49) Genus Oecetis McLachlan, 1877*
118. Oecetis brachyura Yang and Morse, 1997; Kunashir(Oecetis sp.: Vshivkova et al. 1994).
M. Uenishi examined the specimens listed as
Oecetis sp. by Vshivkova et al. (1994) andconfirmed that they are O. brachyura.
119. Oecetis morii Tsuda, 1942*/**
IKIP records: Kunashir* (Arefina et al. 1999).
120. Oecetis nigropunctata Ulmer, 1908*; Paramushir,
Kunashir (Kuwayama 1967a; Kuranishi 2000).
IKIP records: Paramushir, Kunashir, Tanfilyeva*.
(50) Genus Triaenodes McLachlan, 1865*
121. Triaenodes pellectus Ulmer, 1908*/**IKIP records: Kunashir* (Arefina et al. 1999).
122. Triaenodes unanimis McLachlan, 1877*; Iturup(Kuwayama 1967a; Vshivkova et al. 1997).
IKIP records: Iturup, Kunashir*, Zelionyi*.
123. Triaenodes sp.; Iturup, Kunashir (Vshivkova et al.
1994).
Distribution patterns

19. FAMILY MOLANNIDAE WALLENGREN, 1891

Caddisflies of the Kurils are roughly divided into
four groups based on their distribution patterns: (1) agroup of species that is distributed toward the north endof the archipelago, (2) a group of species that is distributed
toward the south end, (3) a group of species that is
distributed toward both ends, and (4) a group of species(45) Genus Molanna Curtis, 1834*
110. Molanna moesta Banks, 1906*; Iturup, Kunashir,
Shikotan (Kuwayama 1936; Kuwayama 1967a).
IKIP records: Urup*, Iturup, Kunashir, Zelionyi*,
Tanfilyeva* (Arefina 1997g).
55

because of their close distance to Hokkaido and lack ofvolcanic activities (Kryvolutskaya 1973; Pietsch et al.
2001). All of the Habomai islands are flat, and numerous
swamps on the islands provide suitable habitats forseveral caddisfly species that are adapted for lentic
environments. However, the current known species
numbers on the Habomai islands are low, which is due toinsufficient sampling days. IKIP allotted only six
sampling days for the five islands of the Habomai group,and previously no one had visited to these islands forcollecting.
Higher species numbers are also expected for thelarger islands. Due to lack of transportation on the islands,
sampling caddisflies was mostly limited to the areas near
the shore during the expedition. The collectors had toreturn to the research vessel for safety at the end of theday, which also prevented them from sampling far inland.
By staying on the islands, collectors can cover a larger
area, and operate insect traps. The light traps and Malaisetraps were used a few occasions when the research vessel
stayed at the same locations over night, and the results ofthe traps were remarkable.
The expedition was limited to the period from theend of July through August each year. Although this isthe best season to collect adult caddisflies on the islands,
it is reasonable to consider that several species that emerge
in the other seasons were unrecorded. Their emergence
period is expected to be longer than the period of the
expedition, particularly, on the southern islands. Thus,
the knowledge of the Kuril caddisfly fauna could be
improvement. At a minimum, the distributions of thefollowing species are expected to be corrected in future
research. Ecclisomiya kamtshatica and Limnephilus
sericeus are expected to occur on Kunashir, because these
species were recorded from the other major southern
islands and Hokkaido (Nozaki and Tanida 1996; Nozaki
et al. 2000). Since Lepidostoma crassicorne occurs on
Urup, Shikotan and Kunashir, this species may also occur
on Iturup that lies between Urup and Kunashir. Similarly,
Micrasema hanasensis and Agrypnia picta may occur on
Simushir. Limnephilus alienus and L. fuscovittatus are
distributed on the major islands throughout the island
chain, therefore, the former species may occur on
Paramushir, Matua and Shikotan, and the later species
may occur on Shiashkotan.

distributed throughout the archipelago. Most species (87species) are distributed toward the south end. The
Philopotamidae, Phryganeidae, Limnephilidae andApataniidae include two or more patterns of distribution.
The Molannidae are distributed either toward the northend or south end.
Synonymic note
Glossosoma ussuricum was described as
Mystrophora ussurica by Martynov (1934) from thesouthern Ussuri, and is widely distributed in continental
Russia (Vshivkova 1986; Arefina and Levanidova 1997).
On the other hand, Glossosoma inops was described as
M. inops from Kyoto, Japan by Tsuda (1940), and has
been commonly recorded from Japanese islands (Nozaki
et al. 1994). During this study, we compared specimens
collected from the Kurils with those from the southern
Ussuri and the Japanese islands, and found that these two
species are identical. Thus, we conclude that G. inops isa junior subjective synonym of G. ussuricum although
we could not examine the type specimens of both species(see also footnote).

Discussion
IKIP added considerably to the knowledge of the
caddisfly fauna on the Kuril Islands. The expedition
recorded about 80% and added about 20% of the currently
known species. Caddisflies were recorded from nearly
half of the Kuril Islands for the first time. IKIP made it
possible to sample caddisflies from the remote and small
islands that biologists had not visited.
Species number of each island is affected by several
factors, distance from the mainland, island size, age andhabitat availability (MacArthur and Wilson 1967). The
high species numbers on the major southern islands arelikely due to their close distances to the main land
(Hokkaido) and their large island sizes. This is also true
for Shumshu and Paramushir which are relatively large
and close to Kamchatka. The low species numbers of
the central islands is likely due to the large distance fromthe mainland. The species that occur in the central islands
must have great dispersal ability to reach the islands. The
islands without caddisfly records are probably too small
to provide habitats even for those with great dispersal
ability.
Although the Habomai Islands are also small, a
relatively rich caddisfly fauna is expected for these islands

Acknowledgements
The work described here was supported in part by
the International Programs Division and the Biological
Sciences Directorate (Biotic Surveys and Inventories
Program) of the U.S. National Science Foundation, Grant
Nos. DEB-9400821 and DEB-9505031, Theodore W.
Pietsch, principal investigator; and by the Japan Society
for the Promotion of Science, Grant No. BSAR-401,
Kunio Amaoka, principal investigator.

Synonymic list of Glossosoma ussuricum (Martynov,
1934)
Mystrophora ussurica Martynov, 1934, 79–80.
Glossosoma ussuricum: Vshivkova, 1986, 72–74; Arefina& Levanidova, 1997, 38.
Mystrophora inops Tsuda, 1940, 193–194. New
Synonym.
Glossosoma inops: Nozaki et al., 1994, 299–300.

56

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East. I. Subfamily Glossosomatinae Wall. pp. 58–
75. In: I.M. Levanidova and V.V. Bogatov, eds.,
Bottom Organisms of the Far East Freshwaters.
Academia Nauka, Vladivostok. (In Russian.)
VSHIVKOVA, T.S., NOZAKI, T., KURANISHI, R.B.
AND AREFINA, T.I., 1994. Caddisflies (Insecta,
Trichoptera) of the Kurile Islands. Bulletin of
Biogeographical Society of Japan 49, 129–142.
VSHIVKOVA , T.S., MORSE, J.C. AND YANG, L.,1997. Fam. Leptoceridae. pp. 154–202. In: P.A.
LEHR, ed., Key to the Insects of Russian Far
East, Vol. 5. Trichoptera and Lepidoptera.
Dal’nauka, Vladivostok. (In Russian.)
WEAVER, J.S.III, 2002. A synonymy of the caddisfly
genus Lepidostoma Rambur (Trichoptera:
Lepidostomatidae), including a species checklist.
Tijdschrift voor Entomologie 145, 173–192.
WIGGINS, G.B., 1998. The caddisfly family
Phryganeidae (Trichoptera). University ofToronto Press.

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Appendix I: Material Examined
In the following list of material examined, the island locality is followed by IKIP number (see Appendix II) andthe number of specimens (M = male adult, F = female adult, P = pupa, L = larva) in the lot.
Rhyacophila arefini URUP: UR-95-PO-11 (1M), UR-95-PO-18 (3M), UR-95-PO-36 (2M), UR-95-VAT-11 (1M 2F), UR95-VAT-15 (2M), UR-95-VAT-23 (2M), UR-96-PO-60 (2M), UR-96-VAT-41 (1M 1F); ITURUP: IT-94-RG-02 (1M),
IT-94-RG-11 (1M), IT-94-VAT-33 (1F), IT-94-VAT-34 (3L), IT-94-VAT-36 (1F), IT-94-VAT-37 (1L), IT-94-VAT-45(1M), IT-94-VAT-49 (2L), IT-94-VAT-51 (1L), IT-97-TIR-05 (1M), IT-98-NM-19 (2M), IT-98-NM-24 (2M), IT-98VAT-14 (1L), IT-98-VAT-18 (2M), IT-98-VAT-22 (1M), IT-98-VAT-23 (1L), IT-98-VAT-24 (1M), IT-98-VAT-38 (3M);
KUNASHIR: KU-94-NM-21 (4 M 4F), KU-94-NM-13 (1M), KU-94-VAT-06 (2L), KU-94-VAT-11 (3M 2F), KU-98LJW-40 (1M); SHIKOTAN: SH-98-VAT-42 (1M 1F), SH-98-VAT-50 (1L).
Rhyacophila brevicephala ITURUP: IT-98-LJW-31 (1M), IT-98-VAT-32 (1L); KUNASHIR: KU-95-VAT-01 (1P), KU95-VAT-03 (4P), KU-97-NM-01 (7M), KU-97-VAT-01 (2M), KU-98-LJW-40 (3M), KU-98-VAT-01 (2M 1F), KU98-VAT-02 (11P); SHIKOTAN: SH-98-VAT-49 (1F).
Rhyacophila hokkaidensis SIMUSHIR: SI-95-PO-39 (8L); URUP: UR-95-PO-10 (1F 2L 6P), UR-95-PO-15 (3L), UR95-PO-22 (1L), UR-95-PO-34 (1L), UR-95-PO-95 (1M), UR-95-PO-96 (2M), UR-95-PO-105 (4L), UR-95-VAT-08(2L), UR-95-VAT-09 (1M 1F 5P), UR-95-VAT-10 (3L), UR-95-VAT-13 (2L), UR-95-VAT-15 (3L), UR-95-VAT-18(3L), UR-95-VAT-23 (3L), UR-95-VAT-66 (1M), UR-96-VAT-42 (3L); ITURUP: IT-94-RG-01 (2L), IT-94-RG-02(10L), IT-94-RG-03 (8L), IT-94-RG-04 (6M 4F 17L 1P), IT-94-RG-05 (6L), IT-94-RG-06 (1M 15L), IT-94-RG-08(1L), IT-94-RG-11 (1L), IT-94-RG-14 (1M 1F 49L), IT-94-RG-15 (2L), IT-94-VAT-34 (27L 5P), IT-94-VAT-35 (1F
4L 1P), IT-94-VAT-36 (1M), IT-94-VAT-37 (3L), IT-94-VAT-38 (6L), IT-94-VAT-40 (1M 1F 13L), IT-94-VAT-46(2L), IT-94-VAT-49 (2L), IT-94-VAT-51 (23L 2P), IT-95-PO-06 (3L), IT-95-PO-07 (7L), IT-95-PO-08 (1L), IT-95PO-120 (3M 30L), IT-95-VAT-06 (20L 1P), IT-95-VAT-68 (2L), IT-96-VAT-39 (2F), IT-97-VAT-08 (2M 4F), IT-98LJW-23 (2L), IT-98-LJW-28 (2M 1L), IT-98-LJW-30 (1L), IT-98-LJW-35 (1L), IT-98-NM-16 (1F), IT-98-NM-37(1L), IT-98-VAT-10 (9L), IT-98-VAT-12 (3L), IT-98-VAT-14 (3L), IT-98-VAT-22 (1M), IT-98-VAT-23 (5L 4P), IT-98VAT-26 (2F), IT-98-VAT-27 (1L 1P), IT-98-VAT-29 (4L 2P), IT-98-VAT-31 (1M), IT-98-VAT-32 (24L), IT-98-VAT-34(1M), IT-98-VAT-35 (4L), IT-98-VAT-36 (2L); KUNASHIR: KU-94-NM-04 (1M), KU-94-NM-05 (2L), KU-94-NM07 (1L), KU-94-NM-10 (5L), KU-94-NM-11 (2L), KU-94-NM-13 (1M), KU-94-NM-16 (1L), KU-94-NM-17 (3L),
KU-94-NM-18 (2L), KU-94-VAT-06 (5L), KU-94-VAT-07 (1M 1F), KU-94-VAT-08 (3L), KU-94-VAT-09 (4L), KU94-VAT-10 (1M), KU-94-VAT-12 (3L), KU-94-VAT-14 (1M), KU-94-VAT-15 (2L), KU-94-VAT-16 (4L 5P), KU-94VAT-19 (2F 5L), KU-94-VAT-20 (2M), KU-95-PO-02 (2M), KU-95-PO-128 (14M 7F), KU-95-VAT-02 (1M), KU95-VAT-03 (1L), KU-95-VAT-71 (4F), KU-97-NM-01 (3M), KU-97-TIR-04 (3M), KU-97-VAT-01 (3F), KU-98-LJW06 (5L), KU-98-LJW-10 (1M), KU-98-LJW-41 (2L), KU-98-VAT-01 (9M 1F), KU-98-VAT-02 (6L 6P), KU-98-VAT06 (1M 1F 1L), KU-98-VAT-08 (4P), KU-98-TIA-65 (1M), KU-99-KLK-51 (3L), KU-99-VAT-62 (5L), KU-99-VAT63 (1F 3L); SHIKOTAN: SH-94-RG-01 (6L), SH-94-RG-03 (2L), SH-94-VAT-29 (10L 6P), SH-94-VAT-32 (1L 1P),
SH-98-LJW-46 (1M), SH-98-LJW-54 (1M), SH-98-LJW-55 (1L), SH-98-LJW-56 (3L 2P), SH-98-NM-30 (1M), SH98-NM-33 (1L), SH-98-VAT-47 (1F), SH-98-VAT-50 (1L); ZELIONYI : ZE-94-NM-03 (14L), ZE-94-VAT-25 (1L).
Rhyacophila kawamurae ITURUP: IT-98-NM-06 (1M), IT-98-VAT-18 (2F), IT-98-VAT-22 (1F), IT-98-VAT-23 (2L), IT98-VAT-38 (1F); KUNASHIR: KU-94-NM-11 (2M), KU-97-TIR-04 (1M), KU-98-LJW-01 (1M), KU-98-VAT-01(4F).
Rhyacophila mirabilis KETOI: KE-95-PO-63 (2M), KE-95-PO-64 (7P 1P), KE-95-PO-74 (2M), KE-95-PO-75 (7L 6P),
KE-95-PO-76 (1M), KE-95-VAT-37 (17M 5F), KE-95-VAT-38 (8L 19P), KE-95-VAT-44 (4M), KE-95-VAT-45 (15L2P); SIMUSHIR: SI-95-PO-39 (1M 1L), SI-95-PO-43 (1M), SI-95-PO-45 (11M 8F), SI-95-PO-66 (1M), SI-95-PO69 (2M), SI-95-PO-70 (4M), SI-95-PO-80 (5M), SI-95-PO-81 (3P), SI-95-PO-82 (2M), SI-95-VAT-24 (9M 3F), SI95-VAT-25 (6L 8P), SI-95-VAT-26 (2M 1F 1P), SI-95-VAT-27 (3L), SI-95-VAT-29 (10L 2P), SI-95-VAT-30 (10M
1F), SI-95-VAT-40 (4M 1F), SI-95-VAT-48 (22M), SI-95-VAT-49 (5L 10P), SI-95-VAT-50 (3M 3F), SI-95-VAT-51(2L), SI-99-KLK-41 (1L 1P), SI-99-KLK-42 (2M), SI-99-VAT-40 (1M 4L 5P); URUP: UR-95-MO-03 (3M), UR-95MO-60 (1M), UR-95-PO-11 (1M 4L 6P), UR-95-PO-13 (1P), UR-95-PO-18 (3M 1F), UR-95-PO-19 (3L 3P), UR95-PO-23 (4M), UR-95-PO-34 (1L 4P), UR-95-PO-89 (1M), UR-95-PO-94 (1M 2P), UR-95-PO-95 (3M 1F), UR95-PO-96 (1L), UR-95-VAT-10 (2P), UR-95-VAT-11 (2M), UR-95-VAT-15 (2M 14L 8P), UR-95-VAT-23 (4L 9P),
UR-95-VAT-53 (2L 5P), UR-95-VAT-66 (1M), UR-96-NM-22 (2M), UR-96-NM-25 (1M), UR-96-PO-62 (1M), UR96-VAT-41 (5M 8F), UR-96-VAT-42 (4L 1P); ITURUP: IT-94-RG-02 (1M 1F 1L), IT-94-RG-04 (8M 3F 1L), IT-94RG-09 (1M), IT-94-RG-14 (2F 5L), IT-94-VAT-33 (4M 2F), IT-94-VAT-34 (3L 1P), IT-94-VAT-36 (16M 2F), IT-94VAT-37 (1L 4P), IT-94-VAT-45 (1M), IT-94-VAT-46 (2P), IT-94-VAT-50 (7M 1F), IT-94-VAT-51 (5L 12P), IT-95VAT-07 (1F), IT-97-NM-06 (1M), IT-97-NM-20 (2M), IT-97-TIR-06 (1M), IT-97-VAT-06 (6M 1F), IT-97-VAT-08(3M 3F), IT-98-DJB-69 (2M), IT-98-LJW-11 (31M 5F), IT-98-LJW-13 (1M), IT-98-LJW-15 (14M), IT-98-LJW-22(1M), IT-98-LJW-27 (3M), IT-98-LJW-32 (7M 1F), IT-98-LJW-33 (3P), IT-98-LJW-34 (1M), IT-98-LJW-36 (5M),
IT-98-NM-20 (1M), IT-98-NM-23 (6M 3F), IT-98-NM-24 (7M), IT-98-NM-37 (1P), IT-98-VAT-09 (3M 1F), IT-98VAT-10 (1L), IT-98-VAT-11 (20M 7F), IT-98-VAT-13 (11M 1F), IT-98-VAT-18 (14M 3F), IT-98-VAT-22 (5M 3F), IT98-VAT-23 (7P), IT-98-VAT-24 (11M), IT-98-VAT-26 (1M 1F), IT-98-VAT-27 (1P), IT-98-VAT-29 (2P), IT-98-VAT31 (6M 1F), IT-98-VAT-32 (18L 9P), IT-98-VAT-34 (1M), IT-98-VAT-38 (4M 2F); KUNASHIR: KU-94-NM-05 (1F),
KU-94-NM-10 (6M), KU-94-NM-13 (5M), KU-94-NM-21 (2M), KU-94-VAT-06 (5P), KU-94-VAT-07 (4M 10F),
KU-94-VAT-11 (3M), KU-94-VAT-16 (1P), KU-94-VAT-17 (1M 1F), KU-94-VAT-20 (6M 1F), KU-95-VAT-02 (1M),
KU-97-NM-03 (2M 1F), KU-97-TIR-04 (4M), KU-97-VAT-01 (1M), KU-98-LJW-01 (1M), KU-98-LJW-40 (1M),
KU-98-VAT-01 (12M 5F).

59

Rhyacophila nipponica KUNASHIR: KU-94-NM-21 (2M), KU-95-VAT-03 (2L), KU-99-VAT-63 (1L); SHIKOTAN: SH94-VAT-28 (1M), SH-98-LJW-56 (2M), SH-98-VAT-49 (3M), SH-98-VAT-50 (7L); ZELIONYI: ZE-94-NM-05 (2M).
Rhyacophila retracta URUP: UR-95-PO-11 (1M), UR-95-PO-13B (1M), UR-95-PO-21 (3M 5F), UR-95-PO-33 (1F),
UR-95-PO-36 (1F), UR-95-PO-95 (1F), UR-95-VAT-16 (1F), UR-95-VAT-52 (1F); ITURUP: IT-98-LJW-15 (1M
1F), IT-98-VAT-38 (1M 1F); KUNASHIR: KU-94-NM-10 (1L), KU-94-VAT-06 (1L), KU-98-LJW-40 (1M).
Rhyacophila transquilla KUNASHIR: KU-94-VAT-10 (1M), KU-97-TIR-04 (1M).
Apsilochorema sutshanum ITURUP: IT-98-VAT-13 (1M 1F); KUNASHIR: KU-94-VAT-14 (2L), KU-95-VAT-03 (4L),
KU-97-NM-01 (1M 1F), KU-97-NM-03 (3F), KU-97-VAT-02 (3L), KU-97-VAT-03 (1F), KU-98-VAT-01 (1M), KU98-VAT-02 (8P); SHIKOTAN: SH-94-VAT-29 (1F), SH-98-VAT-50 (1P).
Anagapetus schmidi KETOI: KE-95-VAT-38 (16L), KE-95-VAT-45 (1L); SIMUSHIR: SI-95-VAT-25 (25L), SI-95-VAT26 (1L), SI-95-VAT-43 (13L), SI-95-VAT-49 (4L), SI-99-VAT-40 (1L); URUP: UR-95-VAT-23 (4L), UR-95-VAT-53(11L), UR-96-VAT-42 (7L); ITURUP: IT-94-VAT-49 (1L), IT-94-VAT-51 (1L), IT-95-VAT-04 (9L), IT-95-VAT-06(5L), IT-98-VAT-12 (2L), IT-98-VAT-14 (4L), IT-98-VAT-23 (1L); KUNASHIR: KU-94-VAT-06 (2L), KU-94-VAT19 (2L), KU-95-VAT-69 (5L), KU-98-VAT-02 (1L); SHIKOTAN: SH-94-VAT-29 (3L), SH-98-VAT-50 (2L).
Glossosoma dulkejti ITURUP: IT-94-RG-04 (2M 3F), IT-98-VAT-018 (1M 3F); KUNASHIR: KU-94-NM-04 (5M 8F),
KU-94-VAT-01 (38M 20F), KU-94-VAT-16 (1P), KU-94-VAT-17 (1M); SHIKOTAN: SH-94-RG-01 (1M 2F), SH94-RG-03 (6M 6F 1P), SH-94-VAT-31 (34M 10F), SH-94-VAT-32 (7P), SH-98-LJW-60 (2M), SH-98-VAT-47 (3M4F), SH-98-VAT-49 (23M 5F), SH-98-VAT-50 (6L).
Glossosoma intermedium SHUMSHU: SU-99-VAT-23 (5M 5F), SU-00-ATR-11 (1F); PARAMUSHIR: PA-96-VAT-07(1M), PA-99-KLK-25 (4M 4F), PA-99-VAT-20 (13M 16F).
Glossosoma ussuricum SIMUSHIR: SI-95-PO-66 (4M), SI-95-PO-69 (1M), SI-95-VAT-40 (4M 2F), SI-95-VAT-42 (1M),
SI-95-VAT-43 (1P); URUP: UR-95-VAT-53 (5P); ITURUP: IT-94-VAT-34 (2P), IT-94-VAT-49 (1P), IT-94-VAT-51(2P), IT-95-VAT-06 (1P), IT-98-LJW-25 (1M), IT-98-NM-16 (1M), IT-98-VAT-22 (2M); KUNASHIR: KU-94-VAT19 (2P), KU-95-PO-128 (1M), KU-95-VAT-69 (1P), KU-97-VAT-03 (1M), KU-98-LJW-40 (1M); SHIKOTAN: SH94-VAT-29 (15P), SH-98-LJW-56 (1M).
Glossosoma sp. KETOI: KE-95-PO-64 (2L), KE-95-VAT-38 (20L).
Hydroptila sp. ITURUP: IT-98-TIA-31 (1F); POLONSKOGO: PO-98-TIA-64 (1F).
Orthotrichia sp. ZELIONYI: ZE-94-VAT-27 (2L).
Palaeagapetus flexus ITURUP: IT-94-VAT-35 (1M), IT-97-VAT-08 (1M), IT-98-LJW-18 (1L).
Palaeagapetus sp. URUP: UR-95-PO-23 (1F).
Dolophilodes (Dolophilodes) japonicus KUNASHIR: KU-97-NM-03 (1F).
Dolophilodes (Dolophilodes) kunashirensis PARAMUSHIR: PA-99-VAT-20 (5M).
Dolophilodes (Dolophilodes) nomugiensis SHUMSHU: SU-97-NM-42 (1M); KHARIMKOTAN: KH-96-PO-36 (3M);
SIMUSHIR: SI-95-PO-66 (1F); URUP: UR-95-VAT-16 (1M); ITURUP: IT-94-RG-06 (1M 1F), IT-94-VAT-39 (10M
5F), IT-95-PO-120 (1M), IT-95-VAT-67 (5M 4F), IT-96-PO-58 (7M 6F), IT-96-PO-59 (1F), IT-96-VAT-39 (2M 8F),
IT-97-NM-19 (2F), IT-97-VAT-08 (7M 1F), IT-98-LJW-29 (2F), IT-98-LJW-34 (1F), IT-98-VAT-34 (4M 3F).
Dolophilodes sp. KETOI: KE-95-VAT-38 (7L), KE-95-VAT-38 (7L); SHIASHIKOTAN: SA-96-PO-53 (1L).
Kisaura borealis KUNASHIR: KU-97-NM-01 (29M), KU-97-NM-03 (1M), KU-97-TIR-04 (4M).
Kisaura hattorii ITURUP: IT-98-LJW-11 (4M), IT-98-NM-23 (1M), IT-98-VAT-11 (3M 1F); KUNASHIR: KU-97-NM01 (6M), KU-97-NM-03 (1M), KU-97-TIR-04 (1M); SHIKOTAN: SH-98-LJW-54 (5M), SH-98-VAT-51-(1M).
Kisaura sp. KUNASHIR: KU-97-NM-01 (1M 6F), KU-97-NM-03 (1M), KU-97-TIR-04 (2M 2F).
Wormaldia sp. KUNASHIR: KU-95-VAT-02 (1F).
Stenopsyche marmorata ITURUP: IT-97-VAT-06 (1F); KUNASHIR: KU-94-NM-10 (1M 1L), KU-94-NM-18 (1M 3L),
KU-94-VAT-19 (4L), KU-95-VAT-03 (1L 1P), KU-95-VAT-69 (4L), KU-95-VAT-71 (1M), KU-96-NM-36 (1L), KU97-NM-01 (3M 1F), KU-97-VAT-03 (1F), KU-98-LJW-04 (3L), KU-98-LJW-41 (10L), KU-98-VAT-01 (1F), KU-98VAT-02 (1P); SHIKOTAN: SH-98-VAT-50 (1L).
Cheumatopsyche infascia PARAMUSHIR: PA-97-BKU-73 (2M 1F); KUNASHIR: KU-95-VAT-02 (1M), KU-97-BKU06 (1M), KU-97-NM-01 (151M 308F), KU-97-NM-03 (3M 3F), KU-97-TIR-01 (1M 1F), KU-97-TIR-04 (31M 43F),
KU-98-LJW-01 (1M), KU-98-VAT-02 (2M), KU-99-VAT-56 (4M 11F).
Hydropsyche albicephala KUNASHIR: KU-94-NM-16 (1P), KU-97-NM-01 (4M 23F), KU-97-NM-03 (6M 5F), KU-97TIR-04 (1M), KU-97-VAT-01 (2M 2F), KU-98-LJW-40 (1F), KU-98-VAT-01 (1M).
Hydropsyche orientalis ITURUP: IT-97-NM-20 (1F), IT-99-NM-14 (1F); KUNASHIR: KU-97-NM-01 (15M 50F), KU97-NM-03 (6F).
Parapsyche shikotsuensis KUNASHIR: KU-97-NM-01 (1M).
Lype excisa ITURUP: IT-98-LJW-13 (1M); KUNASHIR: KU-95-PO-02 (1M), KU-98-LJW-01 (2M 1F).
Ecnomus tenellus ITURUP: IT-98-DJB-34 (1M), IT-98-LJW-16 (20M), IT-98-LJW-19 (1M), IT-98-VAT-15 (1M), IT-99VAT-50 (77M 8F); KUNASHIR: KU-94-VAT-01 (1M), KU-99-VAT-56 (2M).
Agrypnia acristata ITURUP: IT-96-BKU-86 (2M 2F), IT-96-BKU-87 (6F), IT-96-MO-52 (10F), IT-96-PO-69 (44M
53F), IT-96-PO-75 (1M 2F).
Agrypnia czerskyi ITURUP: IT-94-VAT-44 (1M), IT-96-VAT-48 (7M 10F).
Agrypnia picta SHUMSHU: SU-97-TIR-29 (2M 3F), SU-99-BKU-36 (1M); PARAMUSHIR: PA-96-PO-03 (1F), PA-96PO-09 (1F), PA-96-KLK-21 (2M 6F), PA-96-KLK-22 (26M 50F), PA-96-KLK-23 (1F), PA-99-VAT-10 (13M 11F);SHIASHIKOTAN: SA-96-PO-50 (5M 3F); RASSHUA: RAS-95-MO-13 (1F), RAS-95-PO-49 (1M 3F), RAS-99KLK-39 (1F), RAS-99-KLK-40 (1M 1F), PA-99-VAT-30 (2F), PA-99-VAT-32 (2M 3F); ITURUP: IT-96-PO-69 (1M),
IT-98-DJB-40 (1M 1F), IT-98-NM-10 (1F), IT-98-VAT-15 (1F), IT-99-NM-15 (1M).
Agrypnia sahlbergi SHUMSHU: SU-97-BKU-50 (2F), SU-97-NM-32 (1M), SU-97-NM-34 (2M 2F), SU-97-NM-35 (8M60

2F), SU-97-NM-46 (7M 3F), SU-97-TIR-27 (14M 3F), SU-97-TIR-28 (45M 7F), SU-97-TIR-29 (27M 5F), SU-97TIR-31 (13M 4F), SU-97-VAT-14 (3M 4F), SU-97-VAT-15 (3M), SU-99-BKU-36 (2F), SU-99-KLK-20 (32M 2F),
SU-99-KLK-30 (1F); PARAMUSHIR: PA-96-BKU-15 (3M 1F), PA-96-MO-09 (8M 6F), PA-96-NM-04 (48M 2F),
PA-96-NM-07 (28M 1F), PA-96-NM-08 (8M 1F), PA-96-PO-01 (1F), PA-96-PO-03 (1M), PA-96-PO-09 (2M 3F),
PA-96-VAT-01 (3M), PA-96-VAT-09 (11M 1F), PA-96-VAT-10 (1M), PA-97-BKU-31 (2F), PA-97-BKU-87 (1F), PA97-NM-22 (52M 47F), PA-97-NM-52 (1M), PA-97-NM-57 (1M 1F), PA-97-NM-65 (2M), PA-97-NM-72 (1M 2F),
PA-97-RLC-15 (1M 1F), PA-97-TIR-15 (5M), PA-97-TIR-19 (2M 33F), PA-97-TIR-48 (1F), PA-97-VAT-09 (5M15F), PA-97-VAT-10 (1M), PA-99-KLK-21 (2M), PA-99-KLK-22 (23M 3F), PA-99-KLK-23 (1M).
Agrypnia sordida URUP: UR-95-EMS-05 (1M); ITURUP: IT-97-NM-20 (1M 1F), IT-99-DJB-93 (1M), IT-99-NM-14(3F); KUNASHIR: KU-94-VAT-02 (1M).
Eubasilissa regina KUNASHIR: KU-95-VAT-02 (1M), KU-98-VAT-01 (1F).
Hagenella apicalis KUNASHIR: KU-98-VAT-06 (1M); POLONSKOGO: PO-98-VAT-61 (1M 2F), PO-98-DJB-135 (2F).
Hagenella sibirica PARAMUSHIR: PA-00-DJB-17 (1F).
Oligotricha lapponica PARAMUSHIR: PA-96-RLC-04 (1F), PA-97-DES-43 (1F), PA-97-VAT-10 (1F); ONEKOTAN:
ON-99-KLK-04 (2F).
Oligotricha sp. URUP: UR-95-PO-108 (4L), UR-95-PO-109 (4L); SHUMSHU: SU-97-NM-41 (1L).
Brachycentrus americanus KUNASHIR: KU-94-NM-05 (1L 1P), KU-94-NM-10 (3L).
Brachycentrus sp. SHUMSHU: SU-97-NM-56 (38L).
Micrasema hanasensis KETOI: KE-95-PO-64 (1P), KE-95-VAT-37 (1M 1F); URUP: UR-95-PO-21 (1M), UR-95-VAT16 (1M), UR-95-VAT-23 (1M); ITURUP: IT-98-BKU-60 (1F), IT-98-LJW-15 (1M), IT-98-NM-23 (1M); KUNASHIR:
KU-97-NM-03 (1M).
Micrasema sp. SIMUSHIR: SI-95-PO-40 (1M).
Asynarchus sachalinensis URUP: UR-95-VAT-57 (2F); ITURUP: IT-94-RG-08 (16P); SHIKOTAN: SH-98-VAT-42 (1M
1F).
Dicosmoecus jozankeanus URUP: UR-95-MO-05 (1L), UR-95-PO-10 (1L 1P), UR-95-PO-11 (3L 1P), UR-95-PO-34(1M 2L 23P), UR-95-PO-37 (25P), UR-95-PO-88 (1L), UR-95-PO-95 (1P), UR-95-PO-106 (1F), UR-95-PO-107(1L 2P), UR-95-PO-110 (1F), UR-95-PO-112 (13M 14F), UR-95-PO-118 (1F), UR-95-VAT-08 (1L), UR-95-VAT-09(1L), UR-95-VAT-10 (3L), UR-95-VAT-14 (1M), UR-95-VAT-15 (3L), UR-95-VAT-23 (9L), UR-95-VAT-53 (4L),UR-95-VAT-56 (1L), UR-95-VAT-57 (1M), UR-95-VAT-58 (2L), UR-95-VAT-59 (2L), UR-95-VAT-60 (12M 4F),
UR-95-VAT-66 (1M), UR-96-VAT-42 (5L 3P); ITURUP: IT-94-RG-02 (2L), IT-94-RG-04 (1L), IT-94-RG-05 (1P),
IT-94-RG-08 (6L), IT-94-RG-12 (2P), IT-94-RG-14 (4P), IT-94-RG-15 (1L), IT-94-VAT-34 (1L), IT-94-VAT-37 (1L),
IT-94-VAT-42 (3L), IT-94-VAT-46 (1L), IT-94-VAT-47 (1L) IT-94-VAT-49 (1L), IT-94-VAT-51 (1L), IT-94-VAT-53(3L), IT-98-LJW-23 (1L), IT-98-LJW-28 (1P), IT-98-LJW-33 (1M); KUNASHIR: KU-94-NM-08 (2L), KU-94-NM10 (11P), KU-94-NM-16 (1P), KU-94-NM-17 (6L), KU-94-VAT-06 (1L), KU-94-VAT-09 (1L), KU-94-VAT-16 (1L),
KU-94-VAT-19 (3L), KU-95-VAT-01 (3L), KU-97-VAT-02 (1L), KU-98-LJW-04 (2L), KU-98-LJW-09 (2L), KU-98LJW-41 (3L), KU-98-VAT-02 (3L), KU-98-VAT-08 (5L); SHIKOTAN: SH-94-RG-01 (2L), SH-94-RG-03 (5L), SH94-TWP-03 (1L), SH-98-LJW-57 (4L), SH-98-LJW-63 (1L), SH-98-VAT-50 (1L).
Ecclisocosmoecus spinosus URUP: UR-95-PO-10 (1M 1L), UR-95-PO-15 (2L), UR-95-PO-23 (1M), UR-95-PO-24(1M), UR-95-PO-93 (1F), UR-95-PO-94 (3L), UR-95-PO-105 (18L), UR-95-PO-107 (5L), UR-95-VAT-09 (1L 4P),
UR-95-VAT-10 (3L), UR-95-VAT-13 (7L), UR-95-VAT-15 (1L), UR-95-VAT-19 (1M 6L), UR-95-VAT-53 (1L 15P),
UR-95-VAT-55 (28L), UR-95-VAT-66 (2M), UR-96-PO-62 (1M), UR-96-VAT-41 (3M), UR-96-VAT-42 (9L 2P),
UR-96-VAT-46 (10L); ITURUP: IT-94-RG-04 (17M 6F), IT-94-RG-05 (2L), IT-94-VAT-39 (2M), IT-94-VAT-40 (3M),
IT-95-PO-120 (35M 8F 26L 2P), IT-95-PO-121 (1M), IT-95-VAT-67 (21M 3F), IT-95-VAT-68 (27L), IT-96-PO-58(36M 3F 1L), IT-96-VAT-39 (8M), IT-97-NM-20 (1M 1F), IT-97-NM-84 (1M), IT-97-NM-85 (1M), IT-97-VAT-08(3M), IT-98-LJW-12 (2L), IT-98-LJW-23 (1L), IT-98-LJW-28 (1L), IT-98-LJW-35 (10L), IT-98-VAT-28 (1M), IT-98VAT-29 (16L), IT-98-VAT-32 (1L), IT-98-VAT-34 (1M 1F), IT-98-VAT-35 (4L); KUNASHIR: KU-98-LJW-06 (2L);
SHIKOTAN: SH-98-LJW-55 (4L), SH-98-NM-33 (21L), SH-98-VAT-53 (29L); TANFILYEVA: TA-98-LJW-66 (2L).
Ecclisomyia kamtshatica SHUMSHU: SU-97-NM-42 (2M), SU-97-VAT-24 (1F), SU-97-VAT-25 (1M); PARAMUSHIR:
PA-96-NM-03 (1P), PA-96-PO-07 (1M 1F), PA-96-VAT-08 (1L), PA-97-BKU-73 (1M), PA-97-NM-50 (1M), PA-97NM-59 (1M), PA-97-VAT-10 (1M), PA-97-VAT-29 (4M 5F), PA-97-VAT-35 (4M), PA-97-VAT-36 (2L 4P), PA-97VAT-41 (1M 1F), PA-97-VAT-49 (3M 3F); KETOI: KE-95-MO-37 (1M), KE-95-PO-73 (1M 1F), KE-95-PO-74 (9M
9F), KE-95-PO-75 (2P), KE-95-PO-76 (1M 1F), KE-95-VAT-37 (1F), KE-95-VAT-44 (3M 8F), KE-95-VAT-45 (4L1P), KE-95-VAT-46 (1F); SIMUSHIR: SI-95-PO-70 (1M), SI-95-PO-82 (6M), SI-95-VAT-43 (2L 7P), SI-95-VAT-51(3L), SI-99-KLK-41 (1P); URUP: UR-95-PO-10 (4M), UR-95-PO-11 (1M 4L), UR-95-PO-18 (2M), UR-95-PO-23(1M), UR-95-PO-93 (1M), UR-95-PO-100 (1M), UR-95-VAT-11 (1M), UR-95-VAT-15 (1L), UR-95-VAT-23 (1L
4P), UR-95-VAT-58 (4L), UR-96-VAT-42 (1P); ITURUP: IT-94-RG-05 (20P), IT-94-RG-06 (1M 2 F 1P), IT-94-VAT34 (1L), IT-94-VAT-38 (18P), IT-94-VAT-46 (1L), IT-95-PO-121 (4M 1F), IT-96-NM-19 (3M 1F), IT-96-PO-59 (1M),
IT-97-BKU-19 (3M 1F), IT-97-NM-06 (4M 2F), IT-97-NM-13 (1M), IT-97-NM-20 (8M 6F), IT-97-NM-84 (1M), IT97-VAT-06 (4M 1F), IT-98-LJW-27 (1M), IT-98-LJW-30 (1M), IT-98-VAT-32 (98P).
Grammotaulius inornatus PARAMUSHIR: PA-97-NM-22 (1M).
Grammotaulius signatipennis SHUMSHU: SU-97-VAT-24 (1F); PARAMUSHIR: PA-99-VAT-19 (1F); ONEKOTAN:
ON-99-KLK-04 (1P).
Halesus sachalinensis URUP: UR-95-PO-106 (1M), UR-95-VAT-10 (5L), UR-95-VAT-23 (1L), UR-95-VAT-53 (1F 1L);ITURUP: IT-94-RG-04 (1P), IT-94-TWP-02 (1L); KUNASHIR: KU-94-NM-17 (1L), KU-97-VAT-02 (1L);
SHIKOTAN: SH-94-RG-03 (1L).
Hydatophylax festivus URUP: UR-96-VAT-46 (2L); ITURUP: IT-94-RG-04 (5L), IT-94-RG-14 (1L), IT-96-VAT-47 (4L),61

IT-98-VAT-22 (1M); KUNASHIR: KU-94-NM-01 (6L), KU-94-NM-04 (1L), KU-94-NM-13 (1L), KU-98-LJW-41(3L), KU-99-KLK-52 (1L); SHIKOTAN: SH-94-RG-01 (5L), SH-94-RG-03 (1L); ZELIONYI: ZE-94-NM-03 (17L).
Hydatophylax variabilis KUNASHIR: KU-94-TWP-07 (1L), KU-98-LJW-41 (6L); SHIKOTAN: SH-98-LJW-50 (1L),
SH-98-LJW-57 (2L).
Hydatophylax sp. KETOI: KE-95-PO-73 (1F), KE-95-VAT-38 (1P), KE-95-VAT-44 (1M), KE-95-VAT-45 (1L 1P);
SIMUSHIR: SI-95-PO-80 (1F), SI-95-PO-82 (3M 1F), SI-95-VAT-41 (4 L 4P), SI-95-VAT-43 (1L 1P), SI-95-VAT-48(4M), SI-95-VAT-51 (3L), SI-99-KLK-42 (1M); URUP: UR-95-PO-18 (1M), UR-95-VAT-60 (1M), UR-95-VAT-42(2L 1P); ITURUP: IT-95-VAT-06 (1L), IT-96-NM-19 (4M 1F), IT-97-NM-84 (1M); KUNASHIR: KU-95-PO-127(1M); SHIKOTAN: SH-94-RG-01 (1L), SH-98-LJW-50 (1L).
Hydatophylax sp. PARAMUSHIR: PA-96-VAT-08 (11L).
Lenarchus fuscostramineus URUP: UR-95-PO-112 (1F); ITURUP: IT-98-DJB-71 (1F); SHIKOTAN: SH-98-LJW-46(1F), SH-98-LJW-61 (7L); TANFILYEVA: TA-98-TIA-57 (1M).
Limnephilus alienus SHUMSHU: SU-99-VAT-25 (1L); ONEKOTAN: ON-96-VAT-14 (2L 3P); SHIASHKOTAN: SA96-VAT-28 (1L); RASSHUA: RAS-95-PO-47 (1M 1F), RAS-95-VAT-31 (5L); URUP: UR-95-BKU-73 (1M 1F),
UR-95-PO-102 (1P), UR-95-PO-106 (1M 1F), UR-95-PO-110 (3M 2F), UR-95-PO-111 (1F), UR-95-PO-112 (1F),
UR-95-PO-113 (1M 2F), UR-95-PO-116 (2P), UR-95-VAT-08 (3L), UR-95-VAT-14 (9L), UR-95-VAT-54 (1L), UR95-VAT-61 (1M 12F), UR-95-VAT-63 (3P), UR-95-VAT-65 (1M), UR-96-PO-67 (3P), UR-96-VAT-43 (5L 4P), UR96-VAT-44 (4M 4F), UR-96-VAT-45 (3M 6F); ITURUP: IT-94-RG-07 (4M 3F), IT-94-RG-09 (1M 1P), IT-96-NM-29(2P), IT-96-PO-69 (1M), IT-96-PO-75 (2M 1F), IT-96-VAT-49 (2L), IT-98-DJB-41 (1F), IT-98-LJW-19 (1M), IT-98VAT-21 (1M), IT-99-KLK-44 (1F), IT-99-KLK-45 (1F), IT-99-NM-14 (2M 3F); KUNASHIR: KU-98-DJB-140 (1M),
KU-99-VAT-57 (1F).
Limnephilus elegans PARAMUSHIR: PA-97-VAT-09 (1F), PA-97-VAT-10 (1F), PA-00-ATR-015 (1F), PA-00-DJB-016(1M).
Limnephilus femoralis SHUMSHU: SU-97-NM-34 (1M 1F), SU-97-NM-37 (1F), SU-97-TIR-28 (1F), SU-97-VAT-24(2M 1F), SU-99-KLK-20 (10M 11F), SU-99-VAT-16 (2M 7F); PARAMUSHIR: PA-96-MO-01 (1M), PA-96-VAT-04(1M), PA-97-NM-52 (1F), PA-97-VAT-10 (3F), PA-99-KLK-21 (2F), PA-99-KLK-22 (2M 5F), PA-99-VAT-19 (2F),
PA-99-VAT-20 (1F), PA-00-ATR-16 (2F), PA-00-DJB-16 (2M), PA-00-DJB-17 (1M 1F).
Limnephilus femoratus ITURUP: IT-96-VAT-48 (2M), IT-96-VAT-54 (1M 1F).
Limnephilus fenestratus SHUMSHU: SU-97-VAT-24 (2M 1F); PARAMUSHIR: PA-97-NM-65 (1M 6F), PA-97-NM-72(2F), PA-97-RLC-60 (2M 1F), PA-97-VAT-48 (1F).
Limnephilus fuscovittatus SHUMSHU: SU-97-NM-46 (2M 4F), SU-99-DJB-40A (1F); ATLASOVA: AL-97-NM-53(1F), AL-99-VAT-05 (1M); PARAMUSHIR: PA-96-NM-04 (1M 1F), PA-96-NM-07 (1F), PA-99-VAT-19 (1F); PA00-ATR-13 (1F), PA-00-ATR-15 (2F); ONEKOTAN: ON-99-VAT-02 (30M 22F); KHARIMKOTAN: KH-00-ATR30 (1M); MATUA: MA-96-NM-17 (1M), MA-99-VAT-19 (1F); RASSHUA: RAS-95-PO-47 (1F), RAS-95-PO-49(1M 1F); KETOI: KE-95-PO-61 (1M), KE-95-PO-63 (1M), KE-95-PO-74 (1M 2F), KE-95-VAT-37 (1M), KE-95VAT-44 (4F); SIMUSHIR: SI-95-PO-45 (1M 1F), SI-95-PO-82 (1M), SI-95-VAT-48 (4M); URUP: UR-95-PO-31(1M 1F), UR-95-PO-106 (2M 2F), UR-95-PO-110 (1M), UR-95-PO-112 (1M 1F), UR-95-PO-113 (2F), UR-95-VAT60 (2M), UR-96-BKU-80 (1F); ITURUP: IT-94-RG-13 (1L), IT-96-PO-69 (10M 3F), IT-96-VAT-48 (3M), IT-98DJB-71 (1F), IT-98-NM-16A (2M); KUNASHIR: KU-94-VAT-56 (1M), KU-98-VAT-06 (1M 1F); SHIKOTAN: SH98-DJB-93 (1F), SH-98-LJW-46 (1F), SH-98-LJW-61 (7L), SH-98-NM-35 (1M); TANFILYEVA: TA-98-TIA-57(1M).
Limnephilus incisus PARAMUSHIR: PA-96-MO-06 (1F), PA-97-NM-74 (2M 3F), PA-97-VAT-44 (4M 9F), PA-97-VAT48 (1F).
Limnephilus major SHUMSHU: SU-99-KLK-26 (3M 1F), SU-99-KLK-27 (25M 16F).
Limnephilus nigriceps PARAMUSHIR: PA-97-NM-74 (2M).
Limnephilus nipponicus KETOI: KE-95-PO-61 (2F), KE-95-VAT-39 (1M); SIMUSHIR: SI-99-VAT-34 (2M 2F); URUP:
UR-95-PO-31 (1F), UR-95-PO-106 (1F), UR-95-PO-112 (2M), UR-95-PO-118 (1M), UR-95-VAT-10 (1M), UR-95VAT-22 (1F), UR-95-VAT-60 (2M), UR-96-NM-26 (1F), UR-96-PO-66 (2M), RU-96-VAT-45 (1M), UR-00-DJB-81(1M 1F); ITURUP: IT-94-RG-09 (1M), IT-96-PO-69 (6M 2F), IT-96-PO-75 (2F), IT-96-VAT-48 (4M), IT-96-VAT-50(2M), IT-97-NM-12 (1M), IT-97-NM-21 (1M 2F), IT-98-BKU-26 (1F), IT-98-LJW-37 (1F), IT-98-NM-13 (1F), IT98-VAT-15A (1F), IT-98-VAT-37 (3M 9F), IT-99-KLK-45 (1M), IT-99-NM-13 (1F), IT-99-VAT-54 (1M 2F);
KUNASHIR: KU-95-PO-123 (1F), KU-98-VAT-06 (3M 1F); SHIKOTAN: SH-98-BKU-91 (2F), SH-98-LJW-46(1M 1F).
Limnephilus orientalis URUP: UR-96-PO-66 (1F), UR-96-VAT-41 (1F), UR-00-DJB-81 (2F); ITURUP: IT-96-NM-19(2M), IT-97-BKU-19 (1F), IT-97-NM-20 (1M 4F), IT-97-VAT-06 (1F), IT-99-NM-14 (1F); KUNASHIR: KU-95-PO128 (1F).
Limnephilus ornatulus KUNASHIR: KU-97-VAT-05 (1F), KU-99-VAT-56 (1M).
Limnephilus picturatus SHUMSHU: SU-97-MO-12A (4M 9F), SU-97-VAT-20 (2F), SU-97-VAT-21 (1M), SU-97-VAT23 (1M), SU-97-VAT-24 (1F), SU-99-BKU-36 (1M), SU-99-KLK-27 (6M 12F), SU-99-KLK-28 (17M 4F), SU-99VAT-25 (1F); PARAMUSHIR: PA-97-VAT-10 (1M 1F).
Limnephilus quadratus KUNASHIR: KU-99-VAT-56 (5M 2F), KU-99-VAT-57 (1M); TANFILYEVA: TA-98-TIA-57(5M 1F); IURII: IU-98-TIA-59 (1M).
Limnephilus rhombicus ITURUP: IT-97-VAT-06 (1F).
Limnephilus sericeus SHUMSHU: SU-97-BKU-51 (1M), SU-97-MO-12A (19F), SU-97-NM-34 (3M 2F), SU-97-NM39 (2M 1F), SU-97-NM-49 (2F), SU-97-RLC-30 (1F), SU-97-TIR-29 (2F), SU-97-TIR-30 (1M), SU-97-VAT-14(1F), SU-97-VAT-21 (1M), SU-97-VAT-22 (2F), SU-97-VAT-23 (2M 5F), SU-97-VAT-24 (8M 8F), SU-99-BKU-35

62

(1M 2F), SU-99-KLK-26 (1M), SU-99-KLK-27 (11M 8F), SU-99-VAT-25 (5M 1F), SU-00-DJB-12 (1F);
PARAMUSHIR: PA-96-NM-08 (1M), PA-96-VAT-03 (1F), PA-96-VAT-11 (2F), PA-97-BKU-33 (1M 1F), PA-97MO-25A (1M 1F), PA-97-MO-26A (1F), PA-97-NM-62 (1F), PA-97-NM-74 (1F), PA-97-RLC-60 (1M), PA-97-TIR36 (1M), PA-97-VAT-39 (2M 1F), PA-97-VAT-44 (27M 21F), PA-97-VAT-45 (2F), PA-97-VAT-46 (1M), PA-99-VAT19 (1M), PA-00-ATR-13 (3M 2F), PA-00-ATR-15 (1M 1F), PA-00-ATR-16 (1M 1F), PA-00-DJB-16 (3M 1F);
MAKANRUSHI: MK-97-BKU-94 (1M 1F), MK-97-VAT-55 (1F); ONEKOTAN: ON-96-PO-14 (1P), ON-96-PO-15(1M 1F), ON-96-PO-16 (1M 1F); KHARIMKOTAN: KH-96-PO-32 (1M), KH-96-PO-34 (3M 7F), KH-96-PO-39(1M), KH-00-DJB-30 (1M), KH-00-DJB-31 (1F); SHIASHKOTAN: SA-96-PO-48 (1M), SA-96-VAT-31 (1F), SA96-VAT-32 (12P); MATUA: MA-96-PO-55 (2M 5F), MA-96-VAT-34 (11M 1F), MA-96-VAT-37 (1M), MA-99-KLK36 (46M 34F), MA-99-KLK-38 (2M 2F), MA-99-VAT-29 (1M); RASSHUA: RAS-95-MO-13 (1M 1F), RAS-95PO-47 (1M 3F), RAS-95-PO- 49 (45M 30F), RAS-95-VAT-33 (1M 2F), RAS-95-VAT-36 (7M 14F), RAS-99-KLK39 (1F), RAS-99-KLK-40 (10M 5F), RAS-99-VAT-30 (9M 11F), RAS-99-VAT-32 (8M 4F); KETOI: KE-95-MO-23(1M 2F), KE-95-PO-61 (10M 10F), KE-95-PO-63 (1M 3F), KE-95-PO-77 (3P), KE-95-VAT-37 (3M 8F), KE-95VAT-39 (1M 4F); SIMUSHIR: SI-99-VAT-34 (1F), SI-00-ATR-73 (1M); URUP: UR-95-MO-08 (2M 1F), UR-95MO-72 (1F), UR-95-PO-31 (1M 1F), UR-95-PO-96 (1M), UR-95-PO-106 (4M 2F), UR-95-PO-112 (2M), UR-95PO-113 (29M 12F), UR-95-PO-115 (1M 3F), UR-95-PO-117 (2M 2F), UR-95-VAT-22 (4M 1F), UR-95-VAT-57(1M), UR-95-VAT-60 (1M), UR-95-VAT-65 (2F), UR-96-VAT-41 (1M), UR-00-DJB-81 (5M 4F); ITURUP: IT-94RG-11 (1M), IT-96-PO-75 (1F), IT-98-NM-05 (1M); SHIKOTAN: SH-98-VAT-45 (5M 2F).
Limnephilus sparsus ATLASOVA: AL-97-NM-54 (1M 5F), AL-97-VAT-30 (1M 3F), AL-97-VAT-33 (1M 1F), AL-99KLK-07 (1M); SHUMSHU: SU-97-BKU-44 (1F), SU-97-BKU-51 (2F), SU-97-MO-12A (1F), SU-97-NM-34 (1M
7F), SU-97-NM-35 (1F), SU-97-NM-37 (1M 1F), SU-97-NM-39 (3F), SU-97-NM-42 (6F), SU-97-RLC-30 (1F),
SU-97-TIR-27 (1F), SU-97-TIR-30 (1F), SU-97-TIR-31 (1M 4F), SU-97-TIR-32 (1F), SU-97-TWP-21 (1M 1F),
SU-97-VAT-13 (1F), SU-97-VAT-14 (9F), SU-97-VAT-15 (1F), SU-97-VAT-16 (2M 1F), SU-97-VAT-17 (1P), SU-97VAT-18 (1M 1F), SU-97-VAT-20 (2M 3F), SU-97-VAT-21 (3M 9F), SU-97-VAT-22 (3F), SU-97-VAT-23 (3F), SU97-VAT-24 (1F), SU-99-KLK-28 (2F), SU-99-VAT-23 (1M 1F), SU-00-ATR-11 (1M 4F); PARAMUSHIR: PA-96BKU-03 (1M), PA-96-NM-04 (2M 1F), PA-96-NM-07 (1F), PA-96-NM-08 (9F), PA-96-PO-07 (1F), PA-96-VAT-06(3M 2F), PA-96-VAT-07 (3M), PA-96-VAT-09 (1M 2F), PA-96-VAT-10 (1F), PA-96-VAT-11 (4M 8F), PA-97-BKU33 (1F), PA-97-BKU-70 (1F), PA-97-BKU-77 (1F), PA-97-MO-06A (2F), PA-97-MO-25A (1F), PA-97-MO-34 (1F),
PA-97-NM-52 (1F), PA-97-NM-62 (1M), PA-97-NM-72 (1F), PA-97-NM-74 (2F), PA-97-TIR-17 (1F), PA-97-TIR37 (1F), PA-97-TIR-45 (1M), PA-97-VAT-09 (4F), PA-97-VAT-10 (3F), PA-97-VAT-35 (1F), PA-97-VAT-37 (2F),
PA-97-VAT-39 (1F), PA-97-VAT-46 (1M 1F), PA-97-VAT-48 (1M 1F), PA-97-VAT-49 (1M), PA-99-BKU-32 (1F),
PA-99-VAT-10 (1M), PA-99-VAT-20 (3M), PA-00-ATR-13 (3M 4F), PA-00-DJB-16 (1M 3F), PA-00-DJB-17 (2M);
MAKANRUSHI: MK-97-BKU-94 (1F); ONEKOTAN: ON-96-PO-13 (1F), ON-96-PO-21 (2M 1F), ON-96-PO-23(6M), ON-96-PO-25 (2M), ON-96-PO-30 (1M), ON-96-PO-43 (1F), ON-96-VAT-15 (5M 6F), ON-96-VAT-20 (1M),
ON-96-VAT-23 (1F), ON-99-DJB-07 (1M); KHARIMKOTAN: KH-96-BKU-40 (1F), KH-96-PO-36 (1M 1F), KH96-PO-38 (1M), KH-96-PO-41 (1F), KH-96-VAT-22 (2M 3F); SHIASHKOTAN: SA-96-NM-11 (1F), SA-96-NM14 (1M 1F), SA-96-VAT-29 (1M 3F), SA-99-KLK-33 (1F), SA-00-ATR-39 (1F), SA-00-ATR-42 (1M), SA-00-DJB33 (1M 4F), SA-00-DJB-36 (1F); MATUA: MA-96-NM-16 (1F), MA-96-VAT-34 (5M 5F), MA-99-KLK-36 (1M
6F), MA-99-KLK-37 (3M 3F), MA-99-KLK-38 (1M 1F), MA-99-VAT-29 (1M 1F); RASSHUA: RAS-95-PO-47(1M), RAS-95-PO-49 (3M), RAS-95-PO-52 (10M 3F), RAS-95-VAT-36 (3M 3F), RAS-99-KLK-39 (3M 7F), RAS99-KLK-40 (5M 11F), RAS-99-VAT-30 (2M 4F), RAS-99-VAT-32 (2M 3F); KETOI: KE-95-PO-61 (1F), KE-95-PO63 (1M 1F), KE-95-PO-65 (1F), KE-95-PO-73 (2F); SIMUSHIR: SI-95-PO-82 (2F), SI-95-VAT-26 (1F), SI-95-VAT50 (1M), SI-99-VAT-40 (1F); URUP: UR-95-PO-11 (2F), UR-95-PO-14 (1F), UR-95-PO-18 (1M), UR-95-VAT-10(1M), UR-95-VAT-57 (1F), UR-95-VAT-61 (1F), UR-00-DJB-81 (1M); ITURUP: IT-96-PO-69 (15M 12F), IT-96PO-75 (1F), IT-96-VAT-48 (4M 4F), IT-97-BKU-14 (1M), IT-97-NM-06 (1F), IT-98-LJW-15 (1F), IT-98-NM-20(1M), IT-99-DJB-97 (1F), IT-99-DJB-99 (1F); KUNASHIR: KU-99-KLK-49 (1F), KU-99-KLK-50 (1F); SHIKOTAN:
SH-98-DJB-93 (1F), SH-98-DJB-99 (1F), SH-98-LJW-62 (1F); ZELIONYI: ZE-98-LJW-70 (1M 1F); TANFILYEVA:
TA-98-TIA-57 (1F).
Limnephilus stigma SHUMSHU: SU-97-RLC-30 (1M), SU-99-KLK-27 (8M 5F); PARAMUSHIR: PA-96-MO-06 (1F),
PA-96-NM-08 (1F), PA-96-VAT-11 (1F), PA-97-MO-06A (2M), PA-97-NM-72 (1F), PA-97-VAT-44 (1M 4F);
ONEKOTAN: ON-96-RLC-26 (1M 1F); KHARIMKOTAN: KH-96-PO-34 (5M 6F), KH-96-PO-41 (2M); ITURUP:
IT-94-RG-15 (1M).
Limnephilus subcentralis ITURUP: IT-99-VAT-54 (1M).
Nemotaulius miyakei ITURUP: IT-99-VAT-50 (1F); KUNASHIR: KU-98-VAT-06 (1M), KU-99-NM-18 (1M 1F).
Nothopsyche sp. KUNASHIR: KU-94-NM-18 (1L), KU-98-LJW-41 (1L).
Onocosmoecus unicolor ATLASOVA: AL-97-VAT-30 (1M), AL-99-KLK-08 (11L); SHUMSHU: SU-97-NM-32 (1F),
SU-97-NM-39 (2M), SU-97-NM-42 (1M 2F), SU-97-TIR-32 (1M), SU-97-VAT-20 (1M), SU-97-VAT-21 (1M), SU97-VAT-22 (1M), SU-97-VAT-25 (2M); PARAMUSHIR: PA-96-NM-02 (1M), PA-96-NM-03 (2P), PA-96-PO-07(1M), PA-96-VAT-03 (1F), PA-96-VAT-04 (1M 1F), PA-97-VAT-29 (1M), PA-97-VAT-35 (1M), PA-97-VAT-36 (2P),
PA-97-VAT-42 (3M 1F), PA-97-VAT-49 (4M); MAKANRUSHI: MK-97-NM-77 (2M), MK-97-VAT-51 (4M 1F),
MK-97-VAT-52 (8P); ONEKOTAN: ON-96-MO-11 (1L), ON-96-VAT-15 (2M), ON-96-VAT-16 (3P), ON-96-VAT18 (3L 10P), ON-96-VAT-23 (4M), ON-96-VAT-24 (2L 7P); KHARIMKOTAN: KH-96-PO-36 (5M), KH-96-PO-37(2L), KH-96-VAT-21 (1L); EKARMA: EK-99-KLK-31 (1L); SHIASHKOTAN: SA-96-NM-13 (1M), SA-96-PO-46(1M 1F), SA-96-PO-47 (1L), SA-96-VAT-28 (1L), SA-96-VAT-29 (2M 1F), SA-96-VAT-30 (4L), SA-99-KLK-32(1L), SA-99-KLK-33 (1M); RASSHUA: RAS-95-PO-52 (5M 1F), RAS-95-PO-55 (1M); KETOI: KE-95-PO-62
(6L).

63

Allomyia coronae ITURUP: IT-94-VAT-39 (2L), IT-98-VAT-35 (3L); SHIKOTAN: SH-98-VAT-50 (1L).
Allomyia delicatula SIMUSHIR: SI-95-PO-70 (1F), SI-95-VAT-25 (6L), SI-95-VAT-41 (3L).
Apatania parvula RASSHUA: RAS-95-PO-47 (2M), RAS-95-PO-49 (2M), RAS-95-PO-51 (1F), RAS-95-PO-52 (2M4F), RAS-95-PO-57 (7M), RAS-95-VAT-31 (1F), RAS-95-VAT-32 (5L 20P); KETOI: KE-95-PO-63 (1F), KE-95PO-73 (12M 6F), KE-95-PO-74 (13M 1F), KE-95-PO-76 (2F), KE-95-PO-79 (2M 1F), KE-95-VAT-37 (1M 1F), KE95-VAT-38 (4L 2P), KE-95-VAT-44 (12M 4F), KE-95-VAT-45 (8L 4P), KE-95-VAT-47 (5M 1F); SIMUSHIR: SI-95BKU-32B (4M 4F), SI-95-MO-10 (4M 4F 1P), SI-95-PO-40 (1M 2F), SI-95-PO-41 (1P), SI-95-PO-42 (30M 24F),
SI-95-PO-43 (45M 18F), SI-95-PO-44 (2P), SI-95-PO-46 (4P), SI-95-PO-66 (4M 2F), SI-95-PO-68 (1M), SI-95-PO69 (2M), SI-95-PO-70 (1M 1F), SI-95-PO-80 (3M 5F), SI-95-PO-82 (6M), SI-95-VAT-24 (4M 10F), SI-95-VAT-25(20P), SI-95-VAT-26 (15M 6F), SI-95-VAT-29 (1M 1F), SI-95-VAT-30 (22M 5F), SI-95-VAT-40 (3M 1F), SI-95VAT-41 (3L), SI-95-VAT-42 (1F), SI-95-VAT-43 (2L), SI-95-VAT-48 (3M), SI-95-VAT-49 (7L), SI-95-VAT-50 (1M),
SI-99-DJB-69 (1F), SI-99-DJB-71 (1M), SI-99-KLK-42 (3M 2F), SI-99-NM-08 (1F), SI-99-VAT-40 (4M 1F 1P);
URUP: UR-95-MO-08 (1F), UR-95-MO-61 (2M), UR-95-MO-67 (2M), UR-95-PO-10 (1M 4P), UR-95-PO-11 (3M),
UR-95-PO-14 (7M 4F), UR-95-PO-15 (6P), UR-95-PO-21 (1M), UR-95-PO-23 (5M 6F), UR-95-PO-25 (7M 9F),
UR-95-PO-31 (7M 3F), UR-95-PO-93 (1F), UR-95-PO-96 (24M 2F), UR-95-PO-99 (3M), UR-95-PO-100 (1F), UR95-PO-104 (6M 5F), UR-95-PO-106 (6M 3F), UR-95-PO- 107 (2P), UR-95-PO-118 (12M 1F),UR-95-VAT-08 (6P),
UR-95-VAT-11 (3M 1F), UR-95-VAT-13 (6M 1L 10P), UR-95-VAT-18 (2M), UR-95-VAT-20 (1M), UR-95-VAT-22(13M 2F), UR-95-VAT-52 (2M 3F), UR-95-VAT-53 (2P), UR-95-VAT-57 (10M 4F), UR-95-VAT-58 (1L), UR-95VAT-60 (1M), UR-95-VAT-66 (12M 3F), UR-95-VR-32 (1M), UR-00-ATR-85 (1F), UR-00-ATR-86 (2M), UR-00DJB-81 (14M); ITURUP: IT-94-RG-06 (8M 4F), IT-94-VAT-36 (1F), IT-94-VAT-40 (2F 1P), IT-94-VAT-45 (1M), IT95-PO-06 (1F), IT-95-PO-119 (3M 2F), IT-95-VAT-05 (15M 3F), IT-95-VAT-06 (1M 5L 2P), IT-96-PO-59 (1M), IT96-VAT-39 (1M 1F), IT-97-VAT-08 (1M), IT-98-LJW-18 (5M 1F), IT-98-LJW-24 (2M), IT-98-LJW-34 (2M 1F), IT98-LJW-36 (3M 1F), IT-98-NM-19 (10M 9F), IT-98-DJB-63 (2M), IT-98-VAT-16 (1M 1F), IT-98-VAT-24 (9M 4F),
IT-98-VAT-26 (3M), IT-98-VAT-28 (1M 1F), IT-98-VAT-34 (2M), IT-99-VAT-46 (4M 1P); KUNASHIR: KU-94-VAT04 (1M), KU-95-PO-127 (38M 21F), KU-95-VAT-69 (3L); SHIKOTAN: SH-98-VAT-42 (12M 8F); ANUCHINA:
AN-98-TIA-55 (2F).
Apatania zonella ATLASOVA: AL-97-NM-54 (4F), AL-97-VAT-33 (1M 5F), AL-99-KLK-07 (1F); SHUMSHU: SU-97BKU-60 (2F), SU-97-MO-17 (5F), SU-97-NM-30 (1F), SU-97-NM-32 (12F), SU-97-NM-39 (90F), SU-97-NM-42(12F), SU-97-NM-44 (4F), SU-97-NM-46 (3F), SU-97-TIR-32 (10F), SU-97-TIR-33 (6F), SU-97-VAT-13 (3F), SU97-VAT-16 (16F), SU-97-VAT-17 (3L 7P), SU-97-VAT-18 (10F), SU-97-VAT-20 (18F), SU-97-VAT-21 (1M 34F),
SU-97-VAT-24 (7F), SU-97-VAT-25 (11F), SU-97-VAT-27 (7F), SU-99-DJB-27 (1F), SU-99-KLK-28 (3M 360F),
SU-99-KLK-29 (2P), SU-99-KLK-30 (8F), SU-99-VAT-23 (1M 29F), SU-00-ATR-07 (2F), SU-00-ATR-08 (2F),
SU-00-ATR-11 (6F), SU-00-DJB-10 (1M 3F), SU-00-DJB-10A (1F); PARAMUSHIR: PA-96-MO-01 (1F), PA-96MO-03 (2F), PA-96-NM-02 (7F), PA-96-NM-04 (8F), PA-96-PO-01 (1M 12F), PA-96-PO-05 (1M 3F), PA-96-PO-07(16F), PA-96-VAT-01 (1F), PA-96-VAT-02 (2L), PA-96-VAT-03 (8F), PA-96-VAT-04 (36F), PA-96-VAT-05 (6L 2P),
PA-96-VAT-06 (16F), PA-96-VAT-07 (8F), PA-96-VAT-08 (4P), PA-96-VAT-10 (27F), PA-97-BKU-33 (1F), PA-97BKU-72 (1F), PA-97-MO-25A (3F), PA-97-MO-26A (2F), PA-97-MO-29 (1M 1F), PA-97-MO-33A (3F), PA-97NM-50 (2F), PA-97-NM-57 (2F), PA-97-NM-59 (1F), PA-97-NM-62 (7F), PA-97-NM-73 (3F), PA-97-RLC-63(5F), PA-97-TIR-17 (1M 7F), PA-97-TIR-45 (11F), PA-97-VAT-09 (20F), PA-97-VAT-10 (3F), PA-97-VAT-29 (8F),
PA-97-VAT-35 (7F), PA-97-VAT-36 (1L), PA-97-VAT-37 (1F), PA-97-VAT-39 (3F), PA-97-VAT-41 (9F), PA-97-VAT42 (8F), PA-97-VAT-46 (103F), PA-97-VAT-48 (1F), PA-97-VAT-49 (20F), PA-99-KLK-23 (1F), PA-99-KLK-25 (3F),
PA-99-VAT-20 (7F), PA-00-ATR-15 (10F); MAKANRUSHI: MK-97-BKU-97 (2F), MK-97-BKU-103 (1F), MK97-MO-35A (15F), MK-97-MO-37A (1F), MK-97-NM-77 (8F), MK-97-TIR-49 (50F), MK-97-VAT-51 (26), MK97-VAT-52 (35L), MK-97-VAT-53 (30F), MK-97-VAT-55 (4F); ONEKOTAN: ON-96-BKU-16 (5F), ON-96-MO-11(30F), ON-96-PO-18 (5P), ON-96-PO-19 (24F), ON-96-PO-20 (3L 16P), ON-96-PO-21 (20F), ON-96-PO-22 (2F1P), ON-96-PO-23 (34F), ON-96-PO-24 (5F 5L), ON-96-PO-25 (42F), ON-96-PO-26 (1F), ON-96-PO-27 (5P), ON96-PO-43 (1F), ON-96-RLC-17 (1F), ON-96-VAT-12 (29P), ON-96-VAT-15 (78F), ON-96-VAT-16 (12L 18P), ON96-VAT-17 (1F), ON-96-VAT-18 (5L), ON-96-VAT-19 (7L), ON-96-VAT-23 (7L), ON-96-VAT-24 (8L 3P), ON-99BKU-04 (1F), ON-99-BKU-05 (2F), ON-99-BKU-07 (1F), ON-99-KLK-05 (1M 34F), ON-99-KLK-06 (1F), ON99-VAT-02 (29F), ON-00-ATR-18 (2F), ON-00-ATR-19 (1F), ON-00-DJB-21C (6F); KARIMKOTAN: KH-96-BKU36 (1M 24F), KH-96-BKU-40 (3F), KH-96-PO-36 (5M 93F), KH-96-PO-38 (35F), KH-96-VAT-21 (21L 10P), KH96-VAT-22 (155F), KH-00-ATR-30 (6F); SHIASHKOTAN: SA-96-NM-02 (5F), SA-96-PO-46 (22F), SA-96-PO-47(4F), SA-96-VAT-28 (8L), SA-96-VAT-29 (13F), SA-99-KLK-33 (1F), SA-99-NM-02 (5F), SA-99-VAT-26 (13F);
MATUA: MA-96-NM-16 (13F), MA-96-NM-17 (13F), MA-96-PO-55 (13F), MA-96-VAT-34 (19F), MA-96-VAT37 (12F), MA-99-KLK-36 (3F), MA-99-KLK-37 (1M 11F), MA-99-KLK-38 (5F), MA-99-VAT-29 (21F); RASSHUA:RAS-95-VAT-36 (1F); ITURUP: IT-97-NM-85 (3M 1F).
Apatania sp. EKARMA: EK-99-KLK-31 (17L).
Goera japonica URUP: UR-95-PO-09 (1L 1P); ITURUP: IT-94-RG-13 (3L), IT-94-VAT-41 (1L 3P), IT-94-VAT-43 (3L),
IT-94-VAT-49 (3L), IT-95-PO-06 (1F), IT-95-VAT-04 (1M 11L), IT-95-VAT-06 (1L), IT-96-PO-69 (1M), IT-97-TIR05 (1F), IT-98-LJW-20 (3F), IT-98-LJW-21 (2F), IT-98-LJW-22 (1M 1F), IT-98-NM-10 (1M), IT-98-NM-11 (1M),
IT-98-NM-13 (2M 2F), IT-98-NM-15 (1F), IT-98-NM-16 (1F), IT-98-DJB-48 (1M), IT-98-VAT-15 (3M 2F), IT-98VAT-21 (11M 5F), IT-98-VAT-33 (4M 2F), IT-98-VAT-46 (1F 1P); KUNASHIR: KU-94-VAT-54 (1L 2P), KU-95-PO03 (3L), KU-95-PO-127 (1M), KU-95-VAT-01 (1L), KU-97-NM-01 (2M 16F), KU-97-NM-03 (1M 1F), KU-97TIR-04 (2M), KU-98-LJW-01 (1M), KU-98-LJW-40 (1M), KU-98-VAT-01 (1M 1F); SHIKOTAN: SH-98-LJW-56(1M), SH-98-VAT-49 (1M).
Neophylax japonicus KUNASHIR: KU-94-NM-08 (1L), KU-97-VAT-02 (1L).

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Neophylax ussuriensis URUP: UR-95-MO-05 (12P), UR-95-MO-68 (1F), UR-95-PO-11 (1M 3L), UR-95-PO-12 (2F),
UR-95-PO-13 (5P), UR-95-PO-37 (2P), UR-95-PO-106 (7M 5F), UR-95-PO-107 (2P), UR-95-PO-112 (5M 5F),
UR-95-PO-118 (2M), UR-95-VT-08 (3P), UR-95-VT-10 (1F), UR-95-VT-20 (1P), UR-95-VT-23 (10P), UR-95-VT52 (1F), UR-95-VT-53 (3P), UR-95-VT-57 (7M 9F), UR-95-VT-58 (1L 6P), UR-95-VT-60 (5M 2F), UR-95-VT-66(3M 1F), UR-96-VT-41 (1P), UR-96-VT-42 (2P); ITURUP: IT-94-RG-11 (4P), IT-94-RG-14 (8P), IT-94-VT-34 (1L
2P), IT-94-VT-38 (1L 5P), IT-94-VT-51 (4P), IT-96-NM-19 (1M), IT-98-VT-14 (7L 1P), IT-98-VT-27 (1L 1P), IT-98VT-30 (1M 1F), IT-98-VT-32 (5L 1P), IT-98-VT-36 (8L 1P); KUNASHIR: KU-94-NM-10 (1L), KU-94-NM-13 (2P),
KU-94-VT-09 (1L), KU-94-VT-10a (15L), KU-94-VT-12 (6L, 2P), KU-94-VT-19 (2L 1P), KU-95-VT-01 (1L), KU95-VT-69 (1L 1P), KU-97-VAT-02 (3L), KU-99-KLK-50 (1F); SHIKOTAN: SH-94-VT-29 (1P), SH-94-VT-32 (5L3P), SH-98-LJW-54 1F), SH-98-VT-49 (1M 2F), SH-98-VT-50 (1P).
Lepidostoma complicatum ITURUP: IT-94-VAT-06 (1M), IT-98-VAT-31 (1M); KUNASHIR: KU-94-NM-04 (2M), KU94-NM-11 (1F), KU-94-NM-16 (1M), KU-94-VAT-07 (1M 1F), KU-94-VAT-10 (1M), KU-94-VAT-11 (1M), KU-94VAT-20 (2F), KU-95-PO-127 (9M 2F), KU-95-PO-128 (1M), KU-95-VAT-02 (17M 1F), KU-96-NM-43 (2F), KU97-NM-01 (8M 6F), KU-97-TIR-01 (1M), KU-97-TIR-04 (8M), KU-97-VAT-03 (2M), KU-98-LJW-03 (1L), KU98-LJW-40 (2M 4F), KU-98-VAT-01 (19M 2F), KU-98-TIA-65 (10M 8F), KU-99-DJB-105 (2M), KU-99-KLK-50(4M); SHIKOTAN: SH-98-LJW-46 (1F), SH-98-LJW-54 (3M 2F), SH-98-VAT-49 (1M).
Lepidostoma crassicorne URUP: UR-96-NM-24 (1M); SHIKOTAN: SH-98-VAT-42 (2M), SH-98-VAT-47 (1M).
Lepidostoma hiurai ITURUP: IT-98-LJW-16 (3M 3F), IT-98-LJW-18 (7M 1F), IT-98-LJW-19 (1F); KUNASHIR: KU94-NM-04 (3M), KU-94-VAT-04 (6F), KU-95-PO-127 (6M 6F), KU-98-LJW-40 (1M), KU-98-TIA-65 (9F), KU-99VAT-56 (8M 2F), KU-99-VAT-57 (1M); SHIKOTAN: SH-94-VAT-31 (1M), SH-98-VAT-42 (2M 2F); ZELIONYI:
ZE-94-NM-03 (1M); TANFILYEVA: TA-98-TIA-58 (7L).
Lepidostoma stellatum PARAMUSHIR: PA-97-VAT-29 (3M 3F), PA-97-VAT-35 (1M 1F); MAKANRUSHI: MK-97NM-73 (1F); SHIASHKOTAN: SA-99-VAT-26 (5M 2F); KETOI: KE-95-PO-63 (1M 2F), KE-95-PO-73 (8M 8F),
KE-95-PO-74 (2M), KE-95-PO-79 (2M 1F), KE-95-VAT-37 (2M 3F), KE-95-VAT-47 (1F); SIMUSHIR: SI-95-MO10 (1F), SI-95-PO-40 (2M), SI-95-PO-42 (8M 6F), SI-95-PO-43 (12M 14F), SI-95-PO-66 (1M), SI-95-PO-80 (1F),
SI-95-PO-82 (1M 1F), SI-95-VAT-24 (2M), SI-95-VAT-26 (7M 3F), SI-99-VAT-40 (2M 1F); URUP: UR-95-PO-11(1M), UR-95-VAT-11 (10M 1F), UR-95-VAT-16 (1M), UR-95-VAT-18 (1F); ITURUP: IT-94-RG-06 (8M 1F), IT-95PO-119 (1M 2F), IT-97-NM-06 (1F), IT-97-VAT-08 (17M 7F), IT-98-LJW-27 (1M 2F), IT-98-LJW-34 (1M), IT-98NM-24 (1M), IT-98-VAT-11 (1M), IT-98-VAT-16 (1M 1F), IT-98-VAT-28 (2M).
Molanna moesta URUP: UR-95-PO-102 (2L), UR-95-PO-113 (19M 2F), UR-95-PO-114 (2L), UR-95-VAT-54 (1M), UR95-VAT-58 (2L); ITURUP: IT-94-RG-07 (23M 21F), IT-94-RG-11 (1M 1F), IT-94-VAT-41 (5M 1F), IT-94-VAT-47(1F), IT-96-MO-52 (1F), IT-98-DJB-32 (1M), IT-98-DJB-34 (1F), IT-98-DJB-42 (1M), IT-98-LJW-16 (16M 3F), IT98-LJW-19 (4M 2F), IT-98-LJW-21 (1M), IT-98-NM-06 (18M 5F), IT-98-NM-10 (2M 2F), IT-98-NM-13 (1M 1F),
IT-98-NM-14 (1F), IT-98-NM-15 (3F), IT-98-NM-16 (1F), IT-98-VAT-15 (11M 3F), IT-98-VAT-17 (14M 1F), IT-98VAT-21 (4M), IT-98-VAT-33 (2M), IT-99-KLK-43 (3M 2F), IT-99-VAT-46 (10M 6F), IT-99-VAT-50 (4M); KUNASHIR:
KU-94-NM-01 (83M 13F 20L 2P), KU-94-NM-04 (1M), KU-94-NM-05 (1M), KU-94-NM-21 (13M 10L), KU-94VAT-01 (54M 7F), KU-94-VAT-14 (1M), KU-94-VAT-55 (12M 2F), KU-95-PO-123 (45M 10F), KU-95-PO-127(15M 4F), KU-96-PO-76 (13M), KU-98-LJW-74 (1M 1F), KU-98-LJW-75 (1F), KU-99-NM-18 (11L), KU-99-VAT56 (44M 3F), KU-99-VAT-57 (1M); ZELIONYI: ZE-94-NM-02 (3M 2F), ZE-94-RLC-04 (2M 1F), ZE-94-VAT-24(2M 1F), ZE-98-LJW-70 (1M 1F); TANFILYEVA: TA-98-TIA-57 (8M 4F).
Molanna submarginalis PARAMUSHIR: PA-99-KLK-22 (12M 9F), PA-99-VAT-19 (12M 6F).
Molanna sp. SHUMSHU: SU-97-NM-47 (5 cases).
Molannodes tinctus SHUMSHU: SU-97-VAT-24 (5M 5F); PARAMUSHIR: PA-96-NM-08 (6M), PA-96-NM-09 (1L).
Ceraclea alboguttata KUNASHIR: KU-95-PO-127 (1F).
Ceraclea valentinae ZELIONYI: ZE-94-VAT-27 (1M).
Mystacides azureus ITURUP: IT-98-VAT-21 (1M); KUNASHIR: KU-98-BKU-73 (1M), KU-98-DJB-79 (1F).
Mystacides pacificus URUP: UR-95-EMS-05 (1F), UR-95-PO-113 (2M 1F), UR-95-VAT-14 (1M), UR-95-VAT-65 (4M1F); ITURUP: IT-94-RG-07 (27M 11F), IT-94-VAT-41 (21M 31F), IT-94-VAT-45 (1M), IT-94-VAT-47 (1M), IT-96BKU-87 (7F), IT-96-NM-29 (43M 16F), IT-96-PO-69 (3F), IT-96-VAT-48 (1M), IT-96-VAT-50 (27M), IT-98-NM16A (1M), IT-98-VAT-15 (17M 5F), IT-98-VAT-17 (36M 1F), IT-98-VAT-21 (14M 2F), IT-99-VAT-46 (20M 20F), IT99-VAT-52 (2M 1F); KUNASHIR: KU-94-NM-01 (1M), KU-94-NM-04 (2M 2F), KU-94-NM-05 (1M), KU-94NM-21 (37M 2F), KU-94-VAT-01 (1L), KU-94-VAT-02 (8M 8F), KU-94-VAT-03 (1L), KU-94-VAT-04 (11M 6F),
KU-94-VAT-55 (29M 3F), KU-95-PO-127 (3M 1F), KU-98-BKU-73 (1F), KU-99-VAT-56 (1M 1F); ZELIONYI:
ZE-94-NM-02 (2M), ZE-94-NM-05 (1F), ZE-94-RLC-04 (2M), ZE-94-VAT-21 (6M 3F), ZE-94-VAT-24 (19M 6F),
ZE-94-VAT-27 (40); TANFILYEVA: TA-98-TIA-57 (1M 1F).
Oecetis morii KUNASHIR: KU-96-PO-76 (1M).
Oecetis nigropunctata PARAMUSHIR: PA-97-NM-74 (3M 3F), PA-97-VAT-29 (3M 5F); KUNASHIR: KU-94-VAT-01(1M 1F), KU-95-PO-123 (7M 2F), KU-95-PO-127 (1M), KU-95-VR-38 (1F), KU-96-PO-76 (4M), KU-99-VAT-56(1M); TANFILYEVA: TA-98-TIA-57 (1M).
Triaenodes pellectus KUNASHIR: KU-94-NM-43 (1M), KU-97-NM-01 (2M 5F), KU-97-NM-03 (2M), KU-97-TIR-04(2M 1F).
Triaenodes unanimis ITURUP: IT-98-VAT-17 (2M 2F), IT-98-LJW-19 (1M), IT-98-NM-10 (27M 37F); KUNASHIR:
KU-98-DJB-140 (2M 2F); ZELIONYI: ZE-94-VAT-24 (1M), ZE-94-VAT-27 (1M), ZE-98-LJW-70 (1M).

65

Appendix II: IKIP Field Numbers and Field Data
Each IKIP field number consists of four parts: an abbreviation for the particular island, the year the collection was made, theinitials of the representative collectors and the station number; for example, KU-94-NM-01 indicates that the specimens in thatparticular lot were collected on Kunashir in 1994 by N. Minakawa and R. I. Gara.

ATLASOVA
AL-97-NM-53: Lake fed by Zaperty Creek, near abandoned settlement, Alaidskaya Bay; 50˚49.90’ N, 155˚39.96’ E; Aug
12, 1997; N. Minakawa.
AL-97-NM-54, 55: Zaperty Creek, near abandoned settlement, Alaidskaya Bay; 50˚50.83’ N, 155˚39.45’ E; Aug 12, 1997;
N. Minakawa.
AL-97-VAT-30: Zaperty Creek near abandoned settlement , Alaidskaya Bay; Aug 12, 1997; V. A. Teslenko.
AL-97-VAT-33: Stream near Zaperty Creek and abandoned settlement, Alaidskaya Bay; 50˚50.69’ N, 155˚39.63’ E; Aug
12, 1997; V. A. Teslenko.
AL-99-KLK-07, 08: Stream near Zaperty Creek and abandoned settlement, Alaidskaya Bay; 50˚50.43’ N, 155˚39.68’ E; Jul
25, 1999; N. Minakawa, K. L. Kurowski.
AL-99-VAT-05: Lake near abandoned settlement , Alaidskaya Bay; Jul 25, 1999; V. A. Teslenko.

SHUMSHU
SU-97-BKU-44: Top of coastal slope, first stream north of Koshkina River, about 2 km south of Pochtareva Cape; 50˚49.32’
N, 156˚29.76’ E; Aug 8, 1997; B.K. Urbain.
SU-97-BKU-50: Environs of Bolshoye Lake, along road running parallel with coastline and Bolshoye Marshland, easternside of Cape Chibuynyi; 50˚46.43’ N, 156˚15.34’ E; Aug 9, 1997; B. K. Urbain.
SU-97-BKU-51: Environs of Bolshoye Lake, eastern side of Cape Chibuynyi; 50˚46.20’ N, 156˚14.44’ E; Aug 9, 1997; B.
K. Urbain.
SU-97-BKU-60: Slope of stream valley between Cape Yudina and Luzhnanka River, Babushkina Bay; 50˚39.74’ N, 156˚24.51’
E; Aug 10, 1997; B. K. Urbain.
SU-97-MO-12A: Bolshoye Lake, eastern side of Cape Chibuynyi; 50˚45.48’ N, 156˚15.92’ E; Aug 9, 1997; M. Ohara.
SU-97-MO-17: Luzhnanka River, Babushkina Bay; 50˚39.46’ N, 156˚25.00’ E; Aug 10, 1997; M. Ohara.
SU-97-NM-30: Small pond about 0.8 km from coastline, environs of first stream north of Koshkina River, about 2 km southof Pochtareva Cape; 50˚49.13’ N, 156˚28.93’ E; Aug 8, 1997; N. Minakawa.
SU-97-NM-32, 35: Small pond about 1 km from coastline, environs of first stream north of Koshkina River, about 2 km
south of Pochtareva Cape; 50˚49.21’ N, 156˚28.70’ E; Aug 8, 1997; N. Minakawa.
SU-97-NM-34: Small stream about 0.4 km from coastline, first stream from north of Koshkina River, about 2 km south ofPochtareva Cape; 50˚49.17’ N, 156˚29.43’ E; Aug 8, 1997; N. Minakawa.
SU-97-NM-37: Small stream about 1.3 km from coastline, first stream from north of Koshkina River, about 2 km south ofPochtareva Cape; 50˚49.22’ N, 156˚28.53’ E; Aug 8, 1997; N. Minakawa.
SU-97-NM-39: Meadow about 1.6 km from coastline, environs of first stream from north of Koshkina River, about 2 km
south of Pochtareva Cape; 50˚50.06’ N, 156˚29.17’ E; Aug 8, 1997; N. Minakawa.
SU-97-NM-41: About 3.2 km upriver along floodplain inland from eastern side of Cape Chibuynyi, environs of Bolshoye
Lake; 50˚45.72’ N, 156˚17.84’ E; Aug 9, 1997; N. Minakawa.
SU-97-NM-42: Headwater stream between Cape Yudina and Luzhnanka River, about 1.5 km inland from Babushkina Bay;50˚40.20’ N, 156˚23.98’ E; Aug 10, 1997; N. Minakawa.
SU-97-NM-44: Stream between Cape Yudina and Luzhnanka River inland from Babushkina Bay; 50˚39.78’ N, 156˚24.45’
E; Aug 10, 1997; N. Minakawa.
SU-97-NM-46, 47: Lake about 1 km inland from Babushkina Bay, between Cape Yudina and Luzhnanka River; 50˚40.03’
N, 156˚24.13’ E; Aug 10, 1997; N. Minakawa.
SU-97-NM-49: Small ponds between Cape Yudina and Luzhnanka River, about 1.6 km inland from Babushkina Bay;50˚40.16’ N, 156˚23.80’ E; Aug 10, 1997; N. Minakawa.
SU-97-NM-56: Stream about 3 km inland from eastern side of Cape Chibuynyi, environs of Bolshoye Lake; 50˚45.70’ N,
156˚17.64’ E; Aug 9, 1997; N. Minakawa.
SU-97-RLC-30: 3 km northeast of Bolshoye Lake at far edge of flat marshy area, beginning of slope up to hills, eastern sideof Cape Chibuynyi; 50˚46.66’ N, 156˚17.00’ E; Aug 9, 1997; R.L. Crawford.
SU-97-TIR-27: A series of pools about 500 m upriver about 2 km south of Pochtareva Cape, environs of first river north of
Koshkina River; Aug 8, 1997; T.I. Ritchie.
SU-97-TIR-28: Small lake about 2 km upriver, first river north of Koshkina River, about 2 km south of Pochtareva Cape;50˚49.20’ N, 156˚28.79’ E; Aug 8, 1997; T.I. Ritchie.
SU-97-TIR-29: Small pond in river valley near Bolshoye Lake, eastern side of Cape Chibuynyi; 50˚45.88’ N, 156˚15.94’ E;
Aug 9, 1997; T.I. Ritchie.
SU-97-TIR-30: Small ponds upriver from Bolshoye Lake at start of foothills inland from eastern side of Cape Chibuynyi;
50˚45.85’ N, 156˚16.55’ E; Aug 9, 1997; T.I. Ritchie.
SU-97-TIR-31: Small ponds upriver from Bolshoye Lake at start of foothills inland from eastern side of Cape Chibuynyi;
50˚45.55’ N, 156˚17.36’ E; Aug 9, 1997; T.I. Ritchie.
SU-97-TIR-32: Small ponds upriver from Bolshoye Lake at start of foothills inland from eastern side of Cape Chibuynyi;
50˚45.70’ N, 156˚17.64’ E; Aug 9, 1997; T.I. Ritchie, N. Minakawa.
SU-97-TIR-33: Luzhnanka River about 2 km Babushkina Bay; 50˚39.84’ N, 156˚22.15’ E; Aug 10, 1997; T.I. Ritchie.

66SU-97-TWP-21: Slope above ponds in environs of first river north of Koshkina River, about 2 km south of Pochtareva
Cape; 50˚49.14’ N, 156˚29.34’ E; Aug 8, 1997; T.W. Pietsch.
SU-97-VAT-13: Stream north of Koshkina River, about 2 km south of Pochtareva Cape; 50˚49.48’ N, 156˚29.48’ E; Aug 7,
1997; V. A. Teslenko.
SU-97-VAT-14: Bog about 2 km south of Pochtareva Cape, environs of first river north of Koshkina River; 50˚49.17’ N,
156˚29.43’ E; Aug 8, 1997; V. A. Teslenko.
SU-97-VAT-15: Bog about 2 km south of Pochtareva Cape, environs of first river north of Koshkina River; 50˚48.44’ N,
156˚29.46’ E; Aug 8, 1997; V. A. Teslenko.
SU-97-VAT-16, 17: First stream north of Koshkina River, about 2 km south of Pochtareva Cape, near bunker; 50˚49.30’ N,
156˚30.16’ E; Aug 8, 1997; V. A. Teslenko.
SU-97-VAT-18: First stream north of Koshkina River, about 2 km south of Pochtareva Cape; 50˚49.48’ N, 156˚29.48’ E;
Aug 8, 1997; V. A. Teslenko.
SU-97-VAT-20: Bolshoye Lake, eastern side of Cape Chibuynyi; Aug 9, 1997; V. A. Teslenko.
SU-97-VAT-21: Stream about 2 km from mouth, eastern side of Cape Chibuynyi, environs of Bolshoye Lake; Aug 9, 1997;
V. A. Teslenko.
SU-97-VAT-22: Bog by road between lake and Vasenya River, inland from eastern side of Cape Chibuynyi, environs ofBolshoye Lake; Aug 9, 1997; V. A. Teslenko.
SU-97-VAT-23: Tributary of Vasenya River inland from eastern side of Cape Chibuynyi, environs of Bolshoye Lake; Aug9, 1997; V. A. Teslenko.
SU-97-VAT-24: Ponds near Luzhnanka River, Babushkina Bay; Aug 10, 1997; V. A. Teslenko.
SU-97-VAT-25: Ponds 2 km from tributary of Luzhnanka River, Babushkina Bay; Aug 10, 1997; V. A. Teslenko.
SU-97-VAT-27: Stream 300 m from mouth north of cape in Babushkina Bay, environs of Luzhnanka River; Aug 10, 1997;
V. A. Teslenko.
SU-99-BKU-35: Coastal meadow, eastern side of Cape Chibuynyi, environs of Bolshoye Lake; 50˚46.32’ N, 156˚15.46’ E;
Jul 31, 1999; B. K. Urbain.
SU-99-BKU-36: Stretch of road that is parallel with coastline and adjacent to Bolshoye Lake, eastern side of Cape Chibuynyi;
50˚46.32’ N, 156˚15.46’ E; Jul 31, 1999; B. K. Urbain.
SU-99-DJB-27: Inland from Baykoua Village, Jul 29, 1999; D. J. Bennett.
SU-99-DJB-40A: Bluff over pond, environs of Bolshoye Lake; 50˚45.88’ N, 156˚14.46’ E; Jul 31, 1999; D. J. Bennett.
SU-99-KLK-20: Swamps and lakes, about 6 km inland from Baykoua Village; 50˚41.07’ N, 156˚15.58’ E; Jul 29, 1999; N.
Minakawa, K. L. Kurowski.
SU-99-KLK-26, 27: Swamps, inland from east of Cape Chibuynyi, environs of Bolshoye Lake and Bolshoye River; 50˚46.68’
N, 156˚16.66’ E; Jul 31, 1999; N. Minakawa, K. L. Kurowski.
SU-99-KLK-28: Stream, inland from east of Cape Chibuynyi, environs of Bolshoye Lake and Bolshoye River; 50˚46.47’
N, 156˚16.92’ E; Jul 31, 1999; N. Minakawa, K. L. Kurowski.
SU-99-KLK-29, 30: Stream, inland from east of Cape Chibuynyi, environs of Bolshoye Lake and Bolshoye River; 50˚45.71’
N, 156˚17.07’ E; Jul 31, 1999; N. Minakawa, K. L. Kurowski.
SU-99-VAT-16: Small lake 8 km southeast from Baykoua Village; 50˚42.29’ N, 156˚12.43’ E; Jul 29, 1999; V. A. Teslenko.
SU-99-VAT-23: Vesennyaya River, about 1.5 km from mouth, Bolshoye Bight; 50˚44.62’ N, 156˚15.30’ E; Jul 31, 1999; V.
A. Teslenko.
SU-99-VAT-25: Western part of Bolshoye Lake; 50˚45.32’ N, 156˚15.05’ E; Jul 31, 1999; V. A. Teslenko.
SU-00-ATR-07: South of Baykovo Bay towards Bol’shoye Lake, upstream from valley in a small ravine with many mine
shafts; 50°42.97’ N, 156°12.29’ E; July 24, 2000; T. R. Anderson.
SU-00-ATR-08: South of Baykovo Bay towards Bol’shoye Lake, above ravine with mine shafts on top of ridge; 50°42.97’
N, 156°12.29’ E; July 24, 2000; T. R. Anderson.
SU-00-ATR-11: In ravine heading south and west towards Baykovo Bay (inland from mine shafts); 50°42.97’ N, 156°12.29’
E; Jul 24, 2000; T. R. Anderson.
SU-00-DJB-10, 10A: Inland from Baykovo Bay, upstream from valley, near stream bank between valley walls; 50°42.97’
N, 156°12.29’ E; Jul 24, 2000; D. J. Bennett.
SU-00-DJB-12: Inland from Baykovo Bay, on terrace, inland from stream valley; 50°42.97’ N, 156°12.29’ E; Jul 24, 2000;
D. J. Bennett.

PARAMUSHIR
PA-96-BKU-13: Coastal grassland, near Russian military watch tower, western base of Vasil’Yeva Peninsula; 50˚02.87’ N,
155˚23.46’ E; Aug 3, 1996; B. K. Urbain.
PA-96-BKU-15: Western shore of Pernatoye Lake, inland from western base of Vasilyeva Peninsula; 50˚02.96’ N, 155˚22.92’
E; Aug 3, 1996; B. K. Urbain.
PA-96-MO-01: Utesnyi River; 50˚37.73’ N, 156˚08.21’ E; Aug 1, 1996; M. Ohara.
PA-96-MO-03: Utesnyi River, 200 m from mouth; 50˚37.65’ N, 156˚08.13’ E; Aug 1, 1996; M. Ohara.
PA-96-MO-06: Pond on Vasilyeva Peninsula; 50˚01.41’ N, 155˚23.92’ E; Aug 3, 1996; M. Ohara.
PA-96-MO-09: Near Pernatoye Lake, Vasilyeva Bay; 50˚02.37’ N, 155˚23.34’ E; Aug 3, 1996; M. Ohara.
PA-96-NM-02, 03: Bolsheva River, Vasilyeva Bay; 50˚02.59’ N, 155˚22.54’ E; Aug 3, 1996; N. Minakawa.
PA-96-NM-04: Southwest shore of Pernatoye Lake, Vasilyeva Bay; 50˚02.50’ N, 155˚23.20’ E; Aug 3, 1996; N. Minakawa.
PA-96-NM-07: Small pond about 100 m south of Pernatoya Lake, Vasilyeva Bay; 50˚02.37’ N, 155˚23.34’ E; Aug 3, 1996;
N. Minakawa.
PA-96-NM-08, 09: Small pond on Vasilyev Peninsula; 50˚02.30’ N, 155˚23.61’ E; Aug 3, 1996; N. Minakawa.

67

PA-96-PO-01, 02: Small pond 200 m north of Utyosnaya River, about 1 km inland from sea; 50˚37.85’ N, 156˚07.46’ E;
Aug 1, 1996; P. Oberg, N. Minakawa.
PA-96-PO-03: Small pond 300 m north of Utyosnaya River, about 1 km inland from sea; 50˚37.87’ N, 156˚07.32’ E; Aug 1,
1996; P. Oberg, N. Minakawa.
PA-96-PO-05: Large pond 100 m north of Utyosnaya River, about 1 km inland from sea; 50˚37.83’ N, 156˚07.37’ E; Aug 1,
1996; N. Minakawa.
PA-96-PO-07: Utyosnaya River, 1.5 km inland from sea; 50˚37.72’ N, 156˚07.29’ E; Aug 1, 1996; P. Oberg, N. Minakawa.
PA-96-PO-09: Large pond, 300m east of Pernatoye Lake on Vasiljeva Peninsula; 50˚02.68’ N, 155˚24.04’ E; Aug 3, 1996;
P. Oberg.
PA-96-RLC-04: Small pond in valley of Utyosnaya River; 50˚37.78’ N, 156˚07.45’ E; Aug 1, 1996; R.L. Crawford.
PA-96-VAT-01: Small pond by Utyosnaya River, 300 - 500 m from mouth; Aug 1, 1996; V. A. Teslenko.
PA-96-VAT-02, 03: Small stream on Utyosnaya Slope, about 1 km from sea; Aug 1, 1996; V. A. Teslenko.
PA-96-VAT-04, 05: Utyosnaya River, about 1.2 km from mouth; 50˚37.87’ N, 156˚07.32’ E; Aug 1, 1996; V. A. Teslenko.
PA-96-VAT-06: Small tributary of Bolshaya River, 500 m from mouth, Vasilyeva Bay; Aug 3, 1996; V. A. Teslenko.
PA-96-VAT-07, 08: Small tributary of Bolshaya River, 600 m from mouth, Vasilyeva Bay; Aug 3, 1996; V. A. Teslenko.
PA-96-VAT-09: Pernatoye Lake, Vasilyeva Bay; 50˚02.37’ N, 155˚23.34’ E; Aug 3, 1996; V. A. Teslenko.
PA-96-VAT-10: Small stream about 300 m west from Pernatoye Lake, Vasilyeva Bay; 50˚02.37’ N, 155˚23.34’ E; Aug 3,
1996; V. A. Teslenko.
PA-96-VAT-11: Small pond on Vasilyeva Peninsula; 50˚02.30’ N, 155˚23.61’ E; Aug 3, 1996; V. A. Teslenko.
PA-97-BKU-31, 33: South of lake fed by Savushkina River, near Putyatino settlement, between coastline and road; 50˚44.19’
N, 156˚08.82’ E; Aug 4, 1997; B. K. Urbain.
PA-97-BKU-70: Near Shelekhoya River and Shelekhoya settlement, on coastal plateau inland from Shelekhoya Bay;50˚22.31’ N, 155˚36.05’ E; Aug 13, 1997; B. K. Urbain.
PA-97-BKU-72, 73: Near Shelekhoya River and Shelekhoya settlement, along coastal slope at steep exposed area, nearstream, with protruding wet rocks, inland from Shelekhoya Bay; 50˚22.26’ N, 155˚36.17’ E; Aug 13, 1997; B. K.
Urbain.
PA-97-BKU-77: Near Krasheninnikova River, along coastal terrace, inland from Krasheninnikova Bay; 50˚16.87’ N,
155˚20.43’ E; Aug 14, 1997; B. K. Urbain.
PA-97-BKU-87: Near small lake next to Lake Pernatoye, around abandoned buildings, inland from western base of Vasil’Yeva
Peninsula; 50˚02.82’ N, 155˚24.04’ E; Aug 16, 1997; B. K. Urbain.
PA-97-DES-43: Coastal meadow, western base of Vasil’Yeva Peninsula; 50˚01.09’ N, 155˚23.71’ E; Aug 16, 1997; D. E.
Stevenson.
PA-97-MO-06A: Grassland near center of town of Severo-Kurilsk; 50˚40.35’ N, 156˚08.47’ E; Aug 4, 1997; M. Ohara.
PA-97-MO-25A: Near Shelekhoya River and Shelekhoya settlement, Shelekhoya Bay; 50˚22.50’ N, 155˚35.50’ E; Aug 13,
1997; M. Ohara.
PA-97-MO-26A: Near Krasheninnikova River, Krasheninnikova Bay; 50˚16.62’ N, 155˚20.72’ E; Aug 14, 1997; M. Ohara.
PA-97-MO-29: Grassland, western base of Vasil’Yeva Peninsula; 50˚03.18’ N, 155˚25.24’ E; Aug 16, 1997; M. Ohara
PA-97-MO-33A: Grassland, inland from eastern Tukharaka Bay; 50˚11.06’ N, 155˚39.15’ E; Aug 17, 1997; M. Ohara.
PA-97-MO-34: Grassland, inland from eastern Tukharaka Bay; 50˚11.06’ N, 155˚38.64’ E; Aug 17, 1997; M. Ohara.
PA-97-NM-22: Lake fed by Savushkina River, near Putyatino settlement on shore; 50˚44.33’ N, 156˚08.71’ E; Aug 4, 1997;
N. Minakawa.
PA-97-NM-50: Utyosnaya River, Utyosnaya Bay; 50˚37.81’ N, 156˚06.93’ E; Aug 11, 1997; N. Minakawa.
PA-97-NM-52: Ponds in valley of Utyosnaya River, Utyosnaya Bay; 50˚37.82’ N, 156˚07.43’ E; Aug 11, 1997; N. Minakawa.
PA-97-NM-57: Stream south of Shelekhoya River and near Shelekhoya settlement, Shelekhoya Bay; 50˚22.19’ N, 155˚36.15’
E; Aug 13, 1997; N. Minakawa.
PA-97-NM-59: Stream south of Shelekhoya River and near Shelekhoya settlement, Shelekhoya Bay; 50˚22.08’ N, 155˚36.47’
E; Aug 13, 1997; N. Minakawa.
PA-97-NM-62: Stream near Krasheninnikova River, Krasheninnikova Bay; 50˚16.06’ N, 155˚19.90’ E; Aug 14, 1997; N.
Minakawa.
PA-97-NM-65: Pond near Krasheninnikova River, inland from Krasheninnikova Bay; 50˚15.81’ N, 155˚20.04’ E; Aug 14,
1997; N. Minakawa.
PA-97-NM-72: Pond 200-300 m inland from western base of Vasilyeva Peninsula; 50˚01.40’ N, 155˚24.03’ E; Aug 16,
1997; N. Minakawa.
PA-97-NM-73: Stream that flows into eastern Tukharaka Bay; 50˚10.88’ N, 155˚38.89’ E; Aug 17, 1997; N. Minakawa.
PA-97-NM-74: Lake about 300 m inland from eastern Tukharaka Bay; 50˚11.28’ N, 155˚38.06’ E; Aug 17, 1997; N.
Minakawa.
PA-97-RLC-15: Lake fed by Savushkina River, near Putyatino settlement on coast; 50˚44.08’ N, 156˚08.10’ E; Aug 4,
1997; R. L. Crawford.
PA-97-RLC-60: Near tributary feeding Bolshaya River from east, inland from western base of Vasil’Yeva Peninsula; 50˚03.20’
N, 155˚21.52’ E; Aug 16, 1997; R. L. Crawford.
PA-97-RLC-63: Flat valley bottom immediately north and slightly upslope of Pernatoye Lake, near meandering stream,inland from western base of Vasil’Yeva Peninsula; 50˚02.86’ N, 155˚22.64’ E; Aug 16, 1997; R. L. Crawford.
PA-97-TIR-15, 17: Lake fed by Savushkina River, near Putyatino settlement on coast, around bridge; 50˚43.92’ N, 156˚08.24’
E; Aug 4, 1997; T. I. Ritchie.
PA-97-TIR-19: Lake fed by Savushkina River, near Putyatino settlement on coast, edge of lake; 50˚44.24’ N, 156˚08.83’ E;
Aug 4, 1997; T. I. Ritchie.

68PA-97-TIR-36: Bog south of Lagernoye plateau in environ of Cape Ozernyi; 50˚35.97’ N, 156˚09.66’ E; Aug 11, 1997; T.
I. Ritchie.
PA-97-TIR-37: About 1 km from mouth of Severyanka River, south of Cape Ozernyi; 50˚35.31’ N, 156˚08.72’ E; Aug 11,
1997; T. I. Ritchie.
PA-97-TIR-45: Tributary of Krasheninnikova River about 200 m inland from Krasheninnikova Bay; 50˚16.88’ N, 155˚20.93’
E; Aug 14, 1997; T. I. Ritchie.
PA-97-TIR-48: Small pond western base of Vasil’Yeva Peninsula; 50˚03.07’ N, 155˚24.66’ E; Aug 16, 1997; T. I. Ritchie.
PA-97-VAT-09, 10: Lake fed by Savushkina River, near Putyatino settlement on shore; 50˚43.92’ N, 156˚08.24’ E; Aug 4,
1997; V. A. Teslenko.
PA-97-VAT-29: Tributary of Utyosnaya River about 1 km from mouth, Utyosnaya Bay; Aug 11, 1997; V. A. Teslenko.
PA-97-VAT-35, 36: Stream with waterfall , Shelekhoya River and Shelekhoya settlement, Shelekhoya Bay; Aug 13, 1997;
V. A. Teslenko.
PA-97-VAT-37: Tributary of Krasheninnikova River, Krasheninnikova Bay; 50˚16.91’ N, 155˚21.47’ E; Aug 14, 1997; V.
A. Teslenko.
PA-97-VAT-39: Small stream between Krasheninnikova River and Alenushkina River, Krasheninnikova Bay; Aug 14,
1997; V. A. Teslenko.
PA-97-VAT-41: Mouth of Krasheninnikova River, Krasheninnikova Bay; 50˚16.55’ N, 155˚20.37’ E; Aug 14, 1997; V. A.
Teslenko.
PA-97-VAT-42: Forelnyi Stream, 800 m from mouth, east of Cape Kapustnyi; Aug 15, 1997; V. A. Teslenko.
PA-97-VAT-44: Small pond near light house, east of Cape Kapustnyi; Aug 15, 1997; V. A. Teslenko.
PA-97-VAT-45: Lake about 1 km north of light house, east of Cape Kapustnyi; Aug 15, 1997; V. A. Teslenko.
PA-97-VAT-46: About 1 km from mouth of Shikshanka River, western base of Vasil’Yeva Peninsula; Aug 16, 1997; V. A.
Teslenko.
PA-97-VAT-48: Pond (drying), western base of Vasil’Yeva Peninsula; Aug 16, 1997; V. A. Teslenko.
PA-97-VAT-49: Small stream near Baklanyi Rocks, eastern Tukharaka Bay; Aug 17, 1997; V. A. Teslenko.
PA-99-BKU-32: Near two lakes northeast of Medvezhiya River, inland from Rifovaya Bay; 50˚29.75’ N, 156˚05.73’ E; Jul
30, 1999; B. K. Urbain.
PA-99-KLK-21, 22: Swamps inland from Rifovaya Bay; 50˚29.57’ N, 156˚05.58’ E; Jul 30, 1999; N. Minakawa, K. L.
Kurowski.
PA-99-KLK-23: Ponds near Medvezhiya River inland from Rifovaya Bay; 50˚29.62’ N, 156˚04.92’ E; Jul 30, 1999; N.
Minakawa, K. L. Kurowski.
PA-99-KLK-25: Stream near Medvezhiya River inland from Rifovaya Bay; 50˚29.62’ N, 156˚06.64’ E; Jul 30, 1999; N.
Minakawa, K. L. Kurowski.
PA-99-VAT-10: Meadow near Severo-Kurilsk; Jul 26, 1999; A. Lelej.
PA-99-VAT-19: Small lake northeast of Melkoye Lake, Pujsharija Bight, Rifovaya Bay; 50˚29.57’ N, 156˚05.43’ E; Jul 30,
1999; V. A. Teslenko.
PA-99-VAT-20: Stream near Rofovyi Cape, Pujshariya Bight, Rifovaya Bay; 50˚29.66’ N, 156˚06.70’ E; Jul 30, 1999; V. A.
Teslenko.
PA-00-ATR-13: Inland from eastern side of Vasil’yeva Bay; 50°01.62’ N, 155°23.82’ E; Jul 25, 00; T. R. Anderson.
PA-00-ATR-15, 16: Inland from eastern side of Vasil’yeva Bay, from the southwest side of Lake Pernatoye around to northside of the lake; 50°02.83’ N, 155°22.84’ E; Jul 25, 00; T. R. Anderson.
PA-00-DJB-16, 17: Inland from eastern side of Vasil’yeva Bay; 50°01.62’ N, 155°23.82’ E; Jul 25, 2000; D. J. Bennett.

MAKANRUSHI
MK-97-BKU-94: Northwestern edge of big coastal marshland, at base of foothills, inland from Aakat Bay; 49˚44.29’ N,
154˚25.09’ E; Aug 18, 1997; B. K. Urbain.
MK-97-BKU-97: Northwestern edge of big coastal marshland, at base of foothills, inland from Aakat Bay; 49˚44.28’ N,
154˚25.20’ E; Aug 18, 1997; B. K. Urbain.
MK-97-BKU-103: Southeastern facing steep slope, inland from Aakat Bay; 49˚44.51’ N, 154˚24.99’ E; Aug 18, 1997; B.
K. Urbain.
MK-97-MO-35A: Grassland, inland from Aakat Bay; 49˚44.21’ N, 154˚25.19’ E; Aug 18, 1997; M. Ohara.
MK-97-MO-37A: Grassland, inland from Aakat Bay; 49˚44.34’ N, 154˚25.71’ E; Aug 18, 1997; M. Ohara.
MK-97-NM-77: Stream near Pit’yevoy River about 2 km inland from Aakat Bay; 49˚44.63’ N, 154˚25.74’ E; Aug 18, 1997;
N. Minakawa.
MK-97-TIR-49: Small stream, base of foothills that flows into Aakat Bay; 49˚44.27’ N, 154˚25.24’ E; Aug 18, 1997; T. I.
Ritchie.
MK-97-VAT-51, 52: Godnyi Creek about 1.5 km above from waterfall, inland from Aakat Bay; Aug 18, 1997; V. A. Teslenko.
MK-97-VAT-53: Godnyi Creek about 1.0 km above from waterfall, inland from Aakat Bay; Aug 18, 1997; V. A. Teslenko.
MK-97-VAT-55: Small pond near Aakat Bay, north of Aakat Bight; Aug 18, 1997; V. A. Teslenko.

ONEKOTAN
ON-96-BKU-16: South slope of a river valley inland from Nemo Bay; 49˚36.68’ N, 154˚48.97’ E; Aug 4, 1996; B. K.
Urbain.
ON-96-MO-11: Coastal margin of Nemo Bay; 49˚36.62’ N, 154˚49.40’ E; Aug 4, 1996; M. Yabe.
ON-96-PO-13, 14: Pond (sink holes) on plateau about 2 km from Nemo Bay; 49˚36.30’ N, 154˚50.33’ E; Aug 4, 1996; P.
Oberg, N. Minakawa.

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ON-96-PO-15, 16: 2 small ponds on the plateau about 2.5 km from Nemo Bay; 49˚36.31’ N, 154˚50.39’ E; Aug 4, 1996; P.
Oberg, N. Minakawa.
ON-96-PO-18: A series of ponds (sink holes) in drainage area, about 4.2 km inland from Nemo Bay, headwaters for the
stream that flows into Nemo Bay; 49˚35.66’ N, 154˚51.55’ E; Aug 4, 1996; P. Oberg, N. Minakawa.
ON-96-PO-19, 20: Stream that flows into Nemo Bay, about 100 m from mouth of the stream, north of Lake Chyornoye;49˚36.61’ N, 154˚49.51’ E; Aug 4, 1996; P. Oberg, N. Minakawa.
ON-96-PO-21, 22: Stream, about 1 km south of Cape Subbotina; 49˚23.84’ N, 154˚38.67’ E; Aug 5, 1996; P. Oberg, N.
Minakawa.
ON-96-PO-23, 24: Waterfall, about 1 km south of Cape Subbotina; 49˚24.33’ N, 154˚38.88’ E; Aug 5, 1996; P. Oberg.
ON-96-PO-25: Stream 1 km south of Cape Subbotina; 49˚24.06’ N, 154˚38.65’ E; Aug 5, 1996; P. Oberg, N. Minakawa.
ON-96-PO-26, 27: Stream just north of Resvyi River, that flows into Mussel Bay, between Mussle Point and Lisiy Point,
about 500 m inland from the bay; 49˚23.56’ N, 154˚49.08’ E; Aug 7, 1996; N. Minakawa.
ON-96-PO-30: Ponds on plateau north of Resvyi River, about 4 km inland from Lisiy Point; 49˚23.89’ N, 154˚48.69’ E;
Aug 7, 1996; N. Minakawa.
ON-96-PO-43: Trudnyi River, about 2 km from mouth, southern Onekotan; 49˚16.76’ N, 154˚45.02’ E; Aug 9, 1996; P.
Oberg, N. Minakawa.
ON-96-VAT-12: Stream that flows into Nemo Bay, 300 m from the mouth; Aug 4, 1996; V. A. Teslenko.
ON-96-VAT-14: Small pond on the right side of the stream that flows into Nemo Bay, about 300 m from the bay; Aug 4,
1996; V. A. Teslenko.
ON-96-VAT-15: Waterfall about 2 km to the south of Subbotina Cape; Aug 5, 1996; V. A. Teslenko.
ON-96-VAT-16: Stream about 2 km to the south of Subbotina Cape; 49˚23.84’ N, 154˚38.67’ E; Aug 5, 1996; V. A. Teslenko.
ON-96-VAT-17, 18: Stream north of Resvyi River, Mussel Bay; 49˚23.56’ N, 154˚49.08’ E; Aug 7, 1996; V. A. Teslenko.
ON-96-VAT-19, 20: Small pond on the trail to Lisiy Cape, from Mussel Bay; Aug 7, 1996; V. A. Teslenko.
ON-96-VAT-23, 24: Trudnyi River, south part of Onekotan; 49˚16.49’ N, 154˚45.04’ E; Aug 9, 1996; V. A. Teslenko.
ON-96-RLC-17: Stabilized dune meadow inland near river, Nemo Bay; 49˚36.83’ N, 154˚49.11’ E; Aug 4, 1996; R. L.
Crawford.
ON-96-RLC-26: Rezvyi River, in river valley bottom near abandoned army base; 49˚23.76’ N, 154˚48.54’ E; Aug 7, 1996;
R. L. Crawford.
ON-99-BKU-04: Northern slopes (via narrow foot-trail) which lead up to Lake Chernoye, Nemo Bay; 49˚36.51’ N, 154˚49.85’
E; Jul 23, 1999; B. K. Urbain.
ON-99-BKU-05, 07: Northern slopes (via narrow foot-trail) which lead up to Lake Chernoye, Nemo Bay; 49˚36.56’ N,
154˚49.70’ E; Jul 23, 1999; B. K. Urbain.
ON-99-DJB-07: Plateau between Lake Chernoye and stream that flows into Nemo Bay; 49˚36.18’ N, 154˚49.67’ E; Jul 24,
1999; D. J. Bennett.
ON-99-KLK-04: Pond on plateau, inland from Nemo Bay; 49˚36.10’ N, 154˚49.71’ E; Jul 24, 1999; N. Minakawa, K. L.
Kurowski.
ON-99-KLK-05: Chernoye Lake, inland of Nemo Bay; 49˚35.50’ N, 154˚48.87’ E; Jul 24, 1999; N. Minakawa, K. L.
Kurowski.
ON-99-KLK-06: 2 ponds on the plateau, inland from Nemo Bay; 49˚36.04’ N, 154˚49.78’ E; Jul 24, 1999; N. Minakawa,
K. L. Kurowski.
ON-99-VAT-02: Northwest part of Lake Chernoye Lake, Nemo Bay; 49˚35.50’ N, 154˚48.87’ E; Jul 24, 1999; V. A. Teslenko.
ON-00-ATR-18: Inland of Nemo Bay, south hillside of river valley; 49°36.61’ N, 154°49.21’ E; July 27, 2000; T. R.
Anderson.
ON-00-ATR-19: Inland and a little south of Nemo Bay, atop ridge on flat terrace overlooking bay; 49°36.57’ N, 154°49.24’
E; July 27, 2000; T. R. Anderson.
ON-00-DJB-21C: Inland from Nemo Bay, near river mouth just inland from beach extending up slopes to terrace margin
over valley; 49°36.62’ N, 154°49.21’ E; July 27, 2000; D. J. Bennett.
ON-00-DJB-22C: Inland and south of Nemo Bay, between west end of Lake Chernoye and coast; 49°36.62’ N, 154°49.21’
E; July 27, 2000; D. J. Bennett.

KHARIMKOTAN
KH-96-BKU-36: Mouth of coastal stream in Severgina Bay; 49˚09.79’ N, 154˚29.25’ E; Aug 8, 1996; B. K. Urbain.
KH-96-BKU-40: Atop first coastal plateau inland from Severgina Bay; 49˚09.65’ N, 154˚29.27’ E; Aug 8, 1996; B. K.
Urbain.
KH-96-PO-32: Lake about 2km inland from Severgina Bay, between bay and Lake Lazurnoye; 49˚09.49’ N, 154˚28.24’ E;
Aug 8, 1996; P. Oberg, N. Minakawa.
KH-96-PO-34: Pond amongst ridges about 3 km inland from Severgina Bay, between bay and Lake Lazurnoye; 49˚09.35’
N, 154˚27.98’ E; Aug 8, 1996; P. Oberg, N. Minakawa.
KH-96-PO-36, 37: Stream that flows into Severgina Bay; 49˚09.79’ N, 154˚29.25’ E; Aug 8, 1996; N. Minakawa.
KH-96-PO-38: Stream that flows into Severgina Bay; 49˚09.55’ N, 154˚29.54’ E; Aug 8, 1996; P. Oberg.
KH-96-PO-39: Stream that flows into northern end of Severgina Bay, north of former settlement in the bay; 49˚09.65’ N,
154˚30.19’ E; Aug 8, 1996; P. Oberg.
KH-96-PO-41: Pond 200 m inland from Severgina Bay, and north of former settlement in the bay; 49˚09.66’ N, 154˚30.08’
E; Aug 8, 1996; P. Oberg, A. Lopez.
KH-96-VAT-21, 22: Stream that flows into Severgina Bay; 49˚09.63’ N, 154˚29.23’ E; Aug 8, 1996; V. A. Teslenko.
KH-00-ATR-30: Near lakeshore of Lake Lazurnoye; 49°08.75’ N, 154°27.64’ E; Jul 28, 2000; T. R. Anderson.

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KH-00-DJB-30: Northwestern part of island, environs surrounding Lake Lazurnoye, on hillside, upslope from coastal lake
towards volcano; 49°08.93’ N, 154°28.09’ E; Jul 28, 2000; D. J. Bennett.
KH-00-DJB-31: Northwestern part of island, environs surrounding Lake Lazurnoye, valley on north side of volcano leading
into coastal lake- transect; Jul 28, 2000; D. J. Bennett.

EKARMAEK-99-KLK-31: Small coastal stream by shore, west of Cape Lyutyi; Aug 1, 1999; K. L. Kurowski.

SHIASHKOTAN
SA-96-NM-11: Pond (sink holes) on the plateau about 1.5 km inland Zakatnaya Bay; 48˚46.45’ N, 154˚01.18’ E; Aug 12,
1996; N. Minakawa.
SA-96-NM-13: Stream on the north slope of Mt. Kuntominter Zakatnaya Bay; 48˚46.40’ N, 154˚01.13’ E; Aug 12, 1996; N.
Minakawa.
SA-96-NM-14: Stream on the north slope of Mt. Kuntominter Zakatnaya Bay; 48˚46.59’ N, 154˚02.41’ E; Aug 12, 1996; N.
Minakawa.
SA-96-PO-46, 47: Small waterfall that flows into Zakatnaya Bay; 48˚46.96’ N, 154˚02.66’ E; Aug 11, 1996; P. Oberg, N.
Minakawa.
SA-96-PO-48: Small ponds (sink holes) on the plateau inland from Zakatnaya Bay; 48˚46.80’ N, 154˚02.07’ E; Aug 11,
1996; P. Oberg, N. Minakawa.
SA-96-PO-50, 51: 3 large ponds on the plateau about 1 km inland from Zakatnaya Bay; 48˚46.69’ N, 154˚01.99’ E; Aug 11,
1996; P. Oberg.
SA-96-PO-53: Stream section about 1.5 km upstream from waterfall, Zakatnaya Bay; 48˚46.46’ N, 154˚02.62’ E; Aug 12,
1996; P. Oberg.
SA-96-VAT-28, 29, 30: Stream section above waterfall, Zakatnaya Bay, about 3 km east of Grotovyi Cape; Aug 11, 1996;
V. A. Teslenko.
SA-96-VAT-31, 32: Small pond by trail to waterfall, west side of Grotovyi cape, Zakatnaya Bay; Aug 12, 1996; V. A.
Teslenko.
SA-99-KLK-32, 33: Stream above coastal waterfall north of Cape Obvalinyi; 48˚46.27’ N, 154˚03.95’ E; Aug 2, 1999; N.
Minakawa, K. L. Kurowski.
SA-99-NM-02: Stream above coastal waterfall north of Cape Obvalinyi; 48˚45.94’ N, 154˚03.92’ E; Aug 2, 1999; N.
Minakawa.
SA-99-VAT-26: Coastal stream north of Cape Obvalinyi; 48˚46.31’ N, 154˚04.06’ E; Aug 2, 1999; V. A. Teslenko.
SA-00-ATR-39: Inland from southern part of Zakatnaya Bay, west of landing site, plateau above beach bounded by mountains
to the south and cliffs to the north; 48°46.83’ N, 154°01.85’ E; Jul 29, 2000; T. R. Anderson.
SA-00-ATR-42: Inland from southern part of Zakatnaya Bay, plateau west of Zakatnay Bay, on the edge of a small ravine;
48°46.77’ N, 154°01.99’ E; Jul 29. 2000; T. R. Anderson.
SA-00-DJB-33: Inland from southern part of Zakatnaya Bay, on terrace west of the landing site; 48°46.83’ N, 154°01.85’ E;
Jul 29, 2000; D. J. Bennett.
SA-00-DJB-36: Inland from southern part of Zakatnaya Bay, on terrace above landing site, to the west; 48°46.80’ N,
154°01.95’ E; Jul 29, 2000; D. J. Bennett.

MATUA
MA-96-NM-16: Small coastal stream that flows into Yamato Bay; 48˚03.17’ N, 153˚15.53’ E; Aug 15, 1996; N. Minakawa.
MA-96-NM-17: Small coastal stream that flows into Yamato Bay; 48˚03.19’ N, 153˚15.49’ E; Aug 15, 1996; N. Minakawa.
MA-96-PO-55: Pond about 500 m inland from Ainu Bay; 48˚03.39’ N, 153˚14.38’ E; Aug 14, 1996; P. Oberg.
MA-96-VAT-34: Small pond about 0.5 km from Ainu Bay; 48˚03.39’ N, 153˚14.38’ E; Aug 14, 1996; V. A. Teslenko.
MA-96-VAT-37: Small stream that flows into the small cove between Klyuv Cape and Orlova Cape; 48˚03.46’ N, 153˚15.85’
E; Aug 15, 1996; V. A. Teslenko.
MA-99-KLK-36, 37: Swamps near Ainu Bay; 48˚02.85’ N, 153˚13.68’ E; Aug 3, 1999; N. Minakawa, K. L. Kurowski.
MA-99-KLK-38: Swamps near Ainu Bay; 48˚02.60’ N, 153˚13.87’ E; Aug 3, 1999; N. Minakawa, K. L. Kurowski.
MA-99-VAT-29: Small pond near Ainu Bay; 48˚02.64’ N, 153˚13.88’ E; Aug 3, 1999; V. A. Teslenko.

RASSHUARAS-95-MO-13: Tikhoye Lake and Beloye Lake; 47˚43.35’ N, 152˚59.34’ E; Aug 12, 1995; M. Ohara.
RAS-95-PO-47: Tikhoye Lake; 47˚43.30’ N, 152˚59.52’ E; Aug 12, 1995; N. Minakawa.
RAS-95-PO-49: Small ponds near Tikhoye Lake; 47˚43.32’ N, 152˚59.78’ E; Aug 12, 1995; N. Minakawa, P. Oberg.
RAS-95-PO-51, 52: Inland from southwest section of island near the “arches”, environs of small stream; 47˚43.18’ N,
152˚59.25’ E; Aug 12, 1995; N. Minakawa, P. Oberg.
RAS-95-PO-55: Stream that flows into Pacific Ocean; 47˚43.05’ N, 153˚00.67’ E; Aug 13, 1995; N. Minakawa, P. Oberg.
RAS-95-PO-57: Near mouth of stream that flows into Pacific Ocean; 47˚42.70’ N, 153˚01.74’ E; Aug 13, 1995; P. Oberg,
N. Minakawa.
RAS-95-VAT-31: Tikhoye Lake; 47˚43.34’ N, 152˚59.80’ E; Aug 12, 1995; E.M. Sayenko.
RAS-95-VAT-32, 33: Stream that flows into Tikhoye Lake; 47˚43.05’ N, 153˚00.67’ E; Aug 13, 1995; V. A. Teslenko.
RAS-95-VAT-36: Bog adjacent to stream that flows into Tikhoye Lake; 47˚43.05’ N, 153˚00.67’ E; Aug 13, 1995; V. A.
Teslenko.

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RAS-99-KLK-39, 40: Tikhoye Lake and Beloye Lake, 2 km north of Malen’kaya Bay; 47˚43.36’ N, 152˚59.56’ E; Aug 4,
1999; N. Minakawa, K. L. Kurowski.
RAS-99-VAT-30: Beloye Lake; 47˚43.36’ N, 152˚59.28’ E; Aug 4, 1999; V. A. Teslenko.
RAS-99-VAT-32: Tichoye Lake; 47˚43.40’ N, 152˚59.70’ E; Aug 4, 1999; V. A. Teslenko.

KETOI
KE-95-MO-23: Coastal margin, east of Cape Storozheva; 47˚22.60’ N, 152˚27.76’ E; Aug 15, 1995; M. Ohara.
KE-95-KE-95-MO-23: Coastal margin, east of Cape Storozheva; 47˚22.60’ N, 152˚27.76’ E; Aug 15, 1995; M. Ohara.
KE-95-MO-37: Coastal margin in the environs of Stochnyi River; 47˚17.93’ N, 152˚30.00’ E; Aug 19, 1995; M. Ohara.
KE-95-PO-61, 62: Swamp, west of Kaskad Waterfall, east of Cape Storozheva; 47˚22.54’ N, 152˚27.47’ E; Aug 15, 1995;
N. Minakawa, P. Oberg.
KE-95-PO-63, 64: Small stream about 2 km west of Kaskad Waterfall, east of Cape Storozheva; 47˚22.48’ N, 152˚27.53’ E;
Aug 15, 1995; N. Minakawa, P. Oberg.
KE-95-PO-65: Waterfall about 2 km west from Kaskad Waterfall, east of Cape Storozheva; 47˚22.53’ N, 152˚27.88’ E; Aug
15, 1995; P. Oberg.
KE-95-PO-73: Stream near Stochnyi River; 47˚17.93’ N, 152˚30.00’ E; Aug 19, 1995; N. Minakawa.
KE-95-PO-74, 75: Stochnyi River; 47˚18.11’ N, 152˚29.85’ E; Aug 19, 1995; N. Minakawa, P. Oberg.
KE-95-PO-76: Puddles in environs of Stochnyi River, about 200 m from mouth; 47˚18.11’ N, 152˚29.85’ E; Aug 19, 1995;
N. Minakawa, P. Oberg.
KE-95-PO-77: Puddles in environs of Stochnyi River, top of coastal plateau; 47˚18.08’ N, 152˚30.21’ E; Aug 19, 1995; N.
Minakawa.
KE-95-PO-79: Stream near Stochnyi River; 47˚17.89’ N, 152˚30.69’ E; Aug 19, 1995; P. Oberg.
KE-95-VAT-37, 38: Waterfall about 2 km west from Kaskad Waterfall, east of Cape Storozheva; 47˚22.48’ N, 152˚27.53’ E;
Aug 15, 1995; V. A. Teslenko.
KE-95-VAT-39: Swamp, west of Kaskad Waterfall, east of Cape Storozheva; 47˚22.54’ N, 152˚27.47’ E; Aug 15, 1995; V.
A. Teslenko.
KE-95-VAT-44, 45: Stochnyi River; 47˚17.93’ N, 152˚30.00’ E; Aug 19, 1995; V. A. Teslenko.
KE-95-VAT-46: Puddles in environs of Stochnyi River, about 1 km upriver from mouth; 47˚18.11’ N, 152˚29.85’ E; Aug 19,
1995; V. A. Teslenko.
KE-95-VAT-47: Stochnyi River; 47˚18.03’ N, 152˚29.96’ E; Aug 19, 1995; V. A. Teslenko.

SIMUSHIR
SI-95-BKU-32B: Coastal margin, Kitoboyknaya Bay; 46˚51.15’ N, 151˚47.83’ E; Aug 10, 1995; B. K. Urbain.
SI-95-MO-10: Coastal margin, Kitoboyknaya Bay; 46˚51.12’ N, 151˚47.75’ E; Aug 10, 1995; M. Ohara.
SI-95-PO-39: Small coastal waterfall and stream that flows into Kitoboyknaya Bay; 46˚51.24’ N, 151˚47.83’ E; Aug 10,
1995; N. Minakawa, P. Oberg.
SI-95-PO-40, 41: Small coastal waterfall and stream that flows into Kitoboyknaya Bay; 46˚51.10’ N, 151˚47.73’ E; Aug 10,
1995; P. Oberg.
SI-95-PO-42: Small coastal stream that flows into Kitoboyknaya Bay; 46˚51.25’ N, 151˚48.14’ E; Aug 10, 1995; N. Minakawa,
P. Oberg.
SI-95-PO-43: Small coastal waterfall and stream that flows into Kitoboyknaya Bay; 46˚51.40’ N, 151˚48.37’ E; Aug 10,
1995; N. Minakawa, P. Oberg.
SI-95-PO-44: Small coastal stream that flows into Kitoboyknaya Bay; 46˚51.25’ N, 151˚48.14’ E; Aug 10, 1995; P. Oberg.
SI-95-PO-45, 46: Stream near Kostochko Meteorological station, Kitoboyknaya Bay; 46˚51’ N, 151˚47’ E; Aug 11, 1995;
N. Minakawa, P. Oberg.
SI-95-PO-66: Stream that flows into Malaya Bay; 47˚05.38’ N, 152˚08.15’ E; Aug 18, 1995; N. Minakawa, P. Oberg.
SI-95-PO-68: Stream about 200 m upstream from mouth, Malaya Bay; 47˚05.24’ N, 152˚08.20’ E; Aug 18, 1995; P. Oberg.
SI-95-PO-69: Stream about 600 m upstream from mouth, Malaya Bay; 47˚05.16’ N, 152˚08.32’ E; Aug 18, 1995; P. Oberg.
SI-95-PO-70: Stream that flows into Malaya Bay; 47˚05.60’ N, 152˚08.29’ E; Aug 18, 1995; N. Minakawa.
SI-95-PO-80, 81: River that flows into Srednaya Bay; 46˚59.01’ N, 152˚01.30’ E; Aug 22, 1995; N. Minakawa, P. Oberg.
SI-95-PO-82: Stream that flows into Srednaya Bay; 46˚58.69’ N, 152˚00.77’ E; Aug 22, 1995; N. Minakawa, P. Oberg.
SI-95-VAT-24, 25: Waterfall in Kitoboyknaya Bay; 46˚51.24’ N, 151˚47.83’ E; Aug 10, 1995; V. A. Kostenko.
SI-95-VAT-26, 27: Waterfall in Kitoboyknaya Bay; 46˚51.24’ N, 151˚47.83’ E; Aug 10, 1995; V. A. Teslenko.
SI-95-VAT-29, 30: Stream near Kostochko Meteorological station, Kitoboyknaya Bay; 46˚51’ N, 151˚47’ E; Aug 11, 1995;
V. A. Teslenko.
SI-95-VAT-40: Stream that flows into Malaya Bay; 47˚05.38’ N, 152˚08.15’ E; Aug 18, 1995; V. A. Teslenko.
SI-95-VAT-41: Small stream about 2 km inland from Malaya Bay; 47˚05.38’ N, 152˚08.15’ E; Aug 18, 1995; V. A. Teslenko.
SI-95-VAT-42, 43: Small stream that flows into Malaya Bay; 47˚05.60’ N, 152˚08.22’ E; Aug 18, 1995; V. A. Teslenko.
SI-95-VAT-48, 49: Small stream near old house, Srednaya Bay; 46˚58.94’ N, 152˚01.39’ E; Aug 22, 1995; V. A. Teslenko.
SI-95-VAT-50, 51: Stream about 2 km west of Cape Cheornyi, Srednaya Bay; Aug 22, 1995; V. A. Teslenko.
SI-99-DJB-69: Inland of Dushnaya Bay, about 1 km south of Cape Neprochka; 47˚04.46’ N, 152˚11.60’ E; Aug 9, 1999; D.
J. Bennett.
SI-99-DJB-71: Plain near bluffs overlooking coast of Dushnaya Bay; 47˚04.49’ N, 152˚11.52’ E; Aug 9, 1999; D. J. Bennett.
SI-99-KLK-41, 42: Stream that flows from Oleniy Range into Dushnaya Bay; 47˚04.24’ N, 152˚11.00’ E; Aug 9, 1999; N.
Minakawa, K. L. Kurowski.
SI-99-NM-08: Ponds and swamps by road, inland from Broutona Bay, Sredniy Isthmus; 47˚06.08’ N, 152˚14.24’ E; Aug 8,72

1999; N. Minakawa.
SI-99-VAT-34: Broutona Bay, Peresheek Sredniy, small swamps on the left side of the road from Broutona Bay to Paletz
Cape; about 2.5 km from the southeastern part of the bay; 47˚06.29’ N, 152˚13.85’ E; Aug 8, 1999; V. A. Teslenko.
SI-99-VAT-40: River that flows from Oleniy Range into Dushnaya Bay; 47˚04.22’ N, 152˚11.08’ E; Aug 9, 1999; V. A.
Teslenko.
SI-00-ATR-73: North end of island, inland from northeast part of Broutona Bay, near roadside in disturbed flat area backed
by hillside with sasa and birch trees; 47°07.82’ N, 152°16.40’ E; Aug 2, 2000; T. R. Anderson.

URUP
UR-95-BKU-73A: Environs of Vstrechnyi River, Negodnaya Bay; 45˚57.74’ N, 150˚10.54’ E; Aug 29, 1995; B. K. Urbain.
UR-95-EMS-05: Small marshland north of Tokotan Lake, Otkrytyi Bay; 45˚51.20’ N, 149˚47.30’ E; Aug 4, 1995; V. V.
Bogatov, E. M. Sayenko.
UR-95-MO-03, 05: Stream near Shabalina River, Otkrytyi Bay; 45˚51.04’ N, 149˚46.12’ E; Aug 4, 1995; M. Ohara.
UR-95-MO-08: Straya River, Novo-Kurilskaya Bay; 46˚12.84’ N, 150˚18.69’ E; Aug 8, 1995; M. Ohara.
UR-95-MO-60, 61: Rybnaya River, Smuglyi Bay; 46˚01.02’ N, 149˚58.42’ E; Aug 25, 1995; M. Ohara.
UR-95-MO-67: Lopukhovaya River, Barkhatny Bay; 45˚48.26’ N, 149˚54.59’ E; Aug 28, 1995; M. Ohara.
UR-95-MO-68: Pond in environs of Lopukhovaya River, Barkhatny Bay; 45˚47.79’ N, 149˚54.01’ E; Aug 28, 1995; M.
Ohara.
UR-95-MO-72: Bog in environs of Vstrechnyi River, Negodnaya Bay; 45˚57.47’ N, 150˚10.41’ E; Aug 29, 1995; M. Ohara.
UR-95-PO-09: Tokotan Lake, Otkrytyi Bay; 45˚51.49’ N, 149˚46.67’ E; Aug 4, 1995; N. Minakawa, P. Oberg.
UR-95-PO-10: Stream about 2 km southwest from Tokotan Lake, Otkrytyi Bay; 45˚51.37’ N, 149˚46.66’ E; Aug 4, 1995;
N. Minakawa, P. Oberg.
UR-95-PO-11, 12, 13: Stream about 3 km southwest from Tokotan Lake, Otkrytyi Bay; 45˚51.04’ N, 149˚46.12’ E; Aug 4,
1995; N. Minakawa, P. Oberg.
UR-95-PO-14, 15: Stream that flows into Otkrytyi Bay; 45˚52.22’ N, 149˚47.79’ E; Aug 5, 1995; N. Minakawa.
UR-95-PO-18, 19: Vesyolaya River, Natalie Bay; 46˚05.18’ N, 150˚08.44’ E; Aug 6, 1995; N. Minakawa, P. Oberg.
UR-95-PO-21, 22: Coastal waterfall and stream in Natalie Bay; 46˚05.35’ N, 150˚06.95’ E; Aug 6, 1995; N. Minakawa, P.
Oberg.
UR-95-PO-23, 24: Coastal stream about 3 km south of Vesyolaya River, Natalie Bay; 46˚05’ N, 150˚06’ E; Aug 6, 1995; N.
Minakawa.
UR-95-PO-25: Obzhitaya River, Natalie Bay; 46˚05.52’ N, 150˚10.07’ E; Aug 7, 1995; N. Minakawa, P. Oberg.
UR-95-PO-31: Straya River, Novo-Kurilskaya Bay; 46˚12.41’ N, 150˚19.05’ E; Aug 8, 1995; N. Minakawa, P. Oberg.
UR-95-PO-33, 34, 36, 37: Straya River, Novo-Kurilskaya Bay; 46˚12.35’ N, 150˚18.81’ E; Aug 8, 1995; N. Minakawa, P.
Oberg.
UR-95-PO-88: Marsh in environs of Rybnaya River, Smuglyi Bay; 46˚01.46’ N, 149˚59.02’ E; Aug 24, 1995; N. Minakawa,
P. Oberg.
UR-95-PO-89: Side pools in environs of Rybnaya River, Smuglyi Bay; 46˚01.46’ N, 149˚59.02’ E; Aug 24, 1995; N.
Minakawa, P. Oberg.
UR-95-PO-93, 94: Tributary of Rybnaya River, Smuglyi Bay; 46˚01.06’ N, 149˚59.64’ E; Aug 24, 1995; N. Minakawa.
UR-95-PO-95, 96: Tributary of Rybnaya River, Smuglyi Bay; 46˚01.02’ N, 149˚58.42’ E; Aug 24, 1995; N. Minakawa.
UR-95-PO-99: Stream that flows into Katayeva Bay; 45˚34.82’ N, 149˚26.25’ E; Aug 26, 1995; P. Oberg.
UR-95-PO-100: Stream that flows into Katayeva Bay; 45˚34.73’ N, 149˚26.21’ E; Aug 26, 1995; P. Oberg.
UR-95-PO-102: Lake, about 3 km inland from Katayeva Bay; 45˚36.25’ N, 149˚29.24’ E; Aug 26, 1995; N. Minakawa.
UR-95-PO-104, 105: Stream that flows into Katayeva Bay; 45˚34.82’ N, 149˚26.25’ E; Aug 26, 1995; N. Minakawa.
UR-95-PO-106, 107: Tributary of Lopukhovaya River, Barkhatny Bay; 45˚47.85’ N, 149˚54.07’ E; Aug 28, 1995; N.
Minakawa, P. Oberg.
UR-95-PO-108: Near ponds in environs of Lopukhovaya River, Barkhatny Bay; 45˚47.78’ N, 149˚54.04’ E; Aug 28, 1995;
P. Oberg.
UR-95-PO-109, 110, 111: Near ponds in environs of Lopukhovaya River, Barkhatny Bay; 45˚47.79’ N, 149˚54.01’ E; Aug
28, 1995; P. Oberg, N. Minakawa.
UR-95-PO-112: Tributary of Lopukhovaya River, Barkhatny Bay; 45˚47.85’ N, 149˚54.07’ E; Aug 28, 1995; N. Minakawa.
UR-95-PO-113, 114: Lake in environs of Vstrechnyi River, Negodnaya Bay; 45˚57.85’ N, 150˚10.80’ E; Aug 29, 1995; P.
Oberg, D.E. Hoekstra.
UR-95-PO-115: Vstrechnyi River, Negodnaya Bay; 45˚57.60’ N, 150˚10.43’ E; Aug 29, 1995; P. Oberg, D.E. Hoekstra.
UR-95-PO-116, 117: Near ponds in environs of Vstrechnyi River, Negodnaya Bay; 45˚57.76’ N, 150˚10.82’ E; Aug 29,
1995; P. Oberg, N. Minakawa.
UR-95-PO-118: Mouth of stream in Negodnaya Bay; 45˚56.64’ N, 150˚10.56’ E; Aug 29, 1995; P. Oberg, N. Minakawa.
UR-95-VR-32: Near shore of Katayeva Bay; 45˚34.89’ N, 149˚26.24’ E; Aug 26, 1995; V. Roth.
UR-95-VAT-08: Tokotan Lake, Otkrytyi Bay; 45˚51.49’ N, 149˚47.67’ E; Aug 4, 1995; V. A. Teslenko.
UR-95-VAT-09: Stream about 2 km southwest from Tokotan Lake, Otkrytyi Bay; 45˚51.37’ N, 149˚46.66’ E; Aug 4, 1995;
V. A. Teslenko.
UR-95-VAT-10, 11: Stream about 3 km southwest from Tokotan Lak, Otkrytyi Bay; 45˚51.04’ N, 149˚46.12’ E; Aug 4,
1995; V. A. Teslenko.
UR-95-VAT-13: Coastal waterfall in Otkrytyi Bay; 45˚51.22’ N, 149˚47.79’ E; Aug 5, 1995; V. A. Teslenko.
UR-95-VAT-14: Lake southwest of Tokotan Lake, Otkrytyi Bay; 45˚51.03’ N, 149˚47.71’ E; Aug 5, 1995; E.M. Sayenko.
UR-95-VAT-15: Vesolaya River about 2 km from mouth, Natalie Bay; 46˚05.19’ N, 150˚08.43’ E; Aug 6, 1995; V. A.

73

Teslenko.
UR-95-VAT-16: Waterfall about 4 km east of Vesolaya River, Natalie Bay; 46˚05.35’ N, 150˚06.95’ E; Aug 6, 1995; V. A.
Teslenko.
UR-95-VAT-18, 19: 4 small streams along coast within 4 km east from Vesolaya River, Natalie Bay; 46˚05.35’ N, 150˚06.95’
E; Aug 6, 1995; V. A. Teslenko.
UR-95-VAT-20: Obzhitaya River about 2 km from mouth, Natalie Bay; 46˚05.85’ N, 150˚09.91’ E; Aug 7, 1995; V. A.
Teslenko.
UR-95-VAT-22: Small tributary within bog near mouth of Straya River, Novo-Kurilskaya Bay; 46˚12.41’ N, 150˚19.05’ E;
Aug 8, 1995; V. A. Teslenko.
UR-95-VAT-23: Straya River, Novo-Kurilskaya Bay; 46˚12.35’ N, 150˚18.81’ E; Aug 8, 1995; V. A. Teslenko.
UR-95-VAT-52, 53: Rybnaya River, about 2.5 km from mouth, Smuglyi Bay; 46˚01.46’ N, 149˚59.02’ E; Aug 24, 1995; V.
A. Teslenko.
UR-95-VAT-54, 55: Lake near Osma River about 3 km inland from Katayeva Bay; 45˚36.28’ N, 149˚29.20’ E; Aug 26,
1995; V. A. Teslenko.
UR-95-VAT-56: Lake near PVO Station about 3 km Inland from Katayeva Bay; 45˚36.28’ N, 149˚29.20’ E; Aug 26, 1995;
V. A. Teslenko.
UR-95-VAT-57, 58: Lopukhovaya River about 4.5 km from mouth, Barkhatny Bay; 45˚48.55’ N, 149˚53.55’ E; Aug 28,
1995; V. A. Teslenko.
UR-95-VAT-59: Lopukhovaya River about 4.5 km from mouth, Barkhatny Bay; 45˚48.55’ N, 149˚53.55’ E; Aug 28, 1995;
A. Balanov.
UR-95-VAT-60, 61: Southern tributary of Lopukhovaya River, Barkhatny Bay; 45˚47.78’ N, 149˚54.04’ E; Aug 28, 1995;
V. A. Teslenko, N. Minakawa, P. Oberg.
UR-95-VAT-63: Small Lake north of Vstrechnyi River, Negodnaya Bay; 45˚57.55’ N, 150˚10.33’ E; Aug 29, 1995; V. A.
Teslenko.
UR-95-VAT-65: Lake about 1 km northeast of Vstrechnyi River, Negodnaya Bay; 45˚57.84’ N, 150˚10.63’ E; Aug 29,
1995; V. V. Bogatov, E. M. Sayenko.
UR-95-VAT-66: Stream near the ‘YOSHI MARU’ shipwreck, Negodnaya Bay; 45˚56.60’ N, 150˚10.44’ E; Aug 29, 1995;
V. A. Teslenko., N. Minakawa, P. Oberg.
UR-96-BKU-80: Environs of Kama River mouth, Tetyaeva Bay; 45˚38.85’ N, 149˚27.97’ E; Aug 21, 1996; B. K. Urbain.
UR-96-NM-22: Ukromnaya River about 1.5 km from mouth, Ukromnaya Bay; 45˚35.57’ N, 149˚31.28’ E; Aug 20, 1996;
N. Minakawa.
UR-96-NM-24: Tributary of Ukromnaya River, Ukromnaya Bay; 45˚35.56’ N, 149˚31.26’ E; Aug 20, 1996; N. Minakawa.
UR-96-NM-25: Waterfall at west side of Chrenoburka Bay; 45˚35.23’ N, 149˚32.56’ E; Aug 20, 1996; N. Minakawa.
UR-96-NM-26: Tributary of Kama River, about 1 km from mouth, Tetyayeva Bay; 45˚38.43’ N, 149˚28.20’ E; Aug 21,
1996; N. Minakawa.
UR-96-PO-60: Stream that flows into Ukromnaya Bay, about halfway between Ukromnaya River and Kuzinoty Point;
45˚34.90’ N, 149˚31.85’ E; Aug 20, 1996; P. Oberg.
UR-96-PO-62: Stream that flows into Ukamnaya Bay, between Ukromnaya River and Kuzinoty Point, just north of stream
in UR-96-PO-60; 45˚34.95’ N, 149˚32.04’ E; Aug 20, 1996; P. Oberg.
UR-96-PO-66, 67: Ponds in wetland in environs (300 m south of) of Kama River, about 3 km inland from Tetyaeva Bay;45˚38.62’ N, 149˚28.72’ E; Aug 21, 1996; P. Oberg, N. Minakawa.
UR-96-VAT-41, 42: Ukromnaya River, main river and tributary 2.5 km from river mouth, Ukromnaya Bay; 45˚35.21’ N,
149˚32.14’ E; Aug 20, 1996; V. A. Teslenko.
UR-96-VAT-43: Lake about 100 m inland from Bajdorocknaya Bay; Aug 20, 1996; V. A. Teslenko.
UR-96-VAT-44, 45: Small lakes in environs of Kama River, 3 km from mouth, Tetyaeva Bay; 45˚38.58’ N, 149˚28.75’ E;
Aug 21, 1996; V. A. Teslenko.
UR-96-VAT-46: Tributary of Kama River,that flows from the hill near lake on the left side, Tetyaeva Bay; 45˚38.43’ N,
149˚28.20’ E; Aug 21, 1996; V. A. Teslenko.
UR-00-ATR-85, 86: Inland from Aleutka Bay, lowland near river surrounded by small hills; 45°56.10’ N, 150°09.70’ E;
Aug 7, 2000; T. R. Anderson.
UR-00-DJB-81: Inland of Aleutka Bay, river valley about1/3 mile inland from coast, broad plain adjacent to river; 45°56.18’
N, 150°09.39’ E; Aug 7, 2000; D. J. Bennett.

ITURUP
IT-94-RG-01: Lake Natasha, Dobroye Nachalo Bay; 44˚46’06"N, 147˚11’18"E; Aug 12, 1994; R. I. Gara, N. Minakawa.
IT-94-RG-02: Yasnyi Creek, about 700 m upstream from mouth, Dobroye Nachalo Bay; Aug 13, 1994; R. I. Gara, N.
Minakawa.
IT-94-RG-03: Peschanyi Creek (near Yasnyi Creek), about 60 m upstream from mouth, Dobroye Nachalo Bay; 44˚68.4 N,
147˚18.8 E; Aug 13, 1994; R. I. Gara, N. Minakawa.
IT-94-RG-04: Godbaza River, about 1km from mouth, Dobroye Nachalo Bay; Aug 14, 1994; R. I. Gara, N. Minakawa.IT94-RG-05: Stream that flows into Konservnaya Bay, near Zapravochyi Waterfall; 45˚19.92’ N, 147˚59.91’; Aug 16,
1994; R. I. Gara, N. Minakawa.
IT-94-RG-06: Zapravochyi Waterfall, Konservnaya Bay; 45˚19.81’ N, 147˚59.83’ E; Aug 16, 1994; R. I. Gara, N. Minakawa.
IT-94-RG-07: Lake Lebedinoye near Kurilsk; Aug 17, 1994; R. I. Gara, N. Minakawa.
IT-94-RG-08: Kitovaya River near Kurilsk; 45˚15’30" N, 147˚52’06" E; Aug 17, 1994; R. I. Gara, N. Minakawa.
IT-94-RG-09: Reidovoye Lake near Reidovo; 45˚16.00’ N, 148˚01.30’ E; Aug 18, 1994; R. I. Gara, N. Minakawa.

74

IT-94-RG-11: Porozhistaya River, Tornaya Bay in Prostor Sound; Aug 19, 1994; R. I. Gara, N. Minakawa.
IT-94-RG-12: Lake Sopochnoye, Tornaya Bay in Prostor Sound; Aug 19, 1994; N. Minakawa, R. I. Gara.
IT-94-RG-13: Slavnaya River, about 1 km upriver from mouth, Slavnaya Bay in Prostor Sound; Aug 20, 1994; N. Minakawa,
R. I. Gara.
IT-94-RG-14: Kumushka River by Aktivnyi settlement, Slavnaya Bay in Prostor Sound; 45˚28.90’ N, 148˚39.07’ E; Aug
21, 1994; N. Minakawa, R. I. Gara.
IT-94-RG-15: Pioner River, about 200 m from mouth, Kuibyshevskii Bay, 45˚04’30" N, 147˚38’00" E; Aug 22, 1994; N.
Minakawa, R. I. Gara.
IT-94-TWP-02: Tichaya River, between 20-300 m from mouth, Dobroye Nachalo Bay; 44˚43’18" N, 147˚11’36" E; Aug
13, 1994; T. W. Pietsch, W. A. Palsson, B. K. Urbain, J.A. Lopez.
IT-94-VAT-33, 34: Yasnyi Creek, about 1.5 km from mouth, Dobroye Nachalo Bay; Aug 13, 1994; V. A. Teslenko.
IT-94-VAT-35: Godbaza River, at 50 and 200 m from mouth, Dobroye Nachalo Bay; Aug 14, 1994; V. A. Teslenko.
IT-94-VAT-36, 37: Godbaza River, about 800 m from mouth, Dobroye Nachalo Bay; Aug 14, 1994; V. A. Teslenko.
IT-94-VAT-38: Stream that feeds into Konservnaya Bay, near Zapravochyi Waterfall, eastern Chirip Peninsula; 45˚19.92’
N, 147˚59.91’; Aug 16, 1994; V. A. Teslenko.
IT-94-VAT-39: Springs near, Zapravochyi Waterfall, Konservnaya Bay; 45˚20.01’ N, 147˚59.91’ E; Aug 16, 1994; V. A.
Teslenko.
IT-94-VAT-40: Zapravochyi Waterfall, Konservnaya Bay; 45˚20.01’ N, 147˚59.91’ E; Aug 16, 1994; V. A. Teslenko.
IT-94-VAT-41: Lake Lebedinoye near Kurilsk; Aug 17, 1994; V. A. Teslenko.
IT-94-VAT-42: Kitovaya River, between 1.5 and 2 km from mouth; Aug 17, 1994; V. A. Teslenko.
IT-94-VAT-43, 44: Reidovoye Lake near Reidovo; Aug 18, 1994; V. A. Teslenko.
IT-94-VAT-45, 46: Porozhistaya River, Tornaya Bay in Prostor Sound; Aug 19, 1994; V. A. Teslenko.
IT-94-VAT-47: Lake Sopochnoye, Tornaya Bay in Prostor Sound; Aug 19, 1994; V. A. Teslenko.
IT-94-VAT-49: Slavnaya River, about 2 km from mouth, Slavnaya Bay in Prostor Sound; Aug 20, 1994; V. A. Teslenko.
IT-94-VAT-50, 51: Kumushka River by Aktivnyi settlement, Slavnaya Bay in Prostor Sound; 45˚28.90’ N, 148˚39.07’ E;
Aug 21, 1994; V. A. Teslenko.
IT-94-VAT-53: Pioner River, Kuibyshevskii Bay; Aug 22, 1994; V.V. Bogatov.
IT-95-PO-06: First stream from north of Nezhnyi River, near Kitovyi Village; 45˚17.63’ N, 147˚52.55’ E; Aug 3, 1995; N.
Minakawa, P. Oberg.
IT-95-PO-07: Stream, north of Kitovyi Village, environs of Nezhnyi River; 45˚16.71’ N, 147˚52.29’ E; Aug 3, 1995; N.
Minakawa, P. Oberg.
IT-95-PO-08: Stream, north of Kitovyi Village, environs of Nezhnyi River; 45˚16.71’ N, 147˚52.30’ E; Aug 3, 1995; N.
Minakawa, P. Oberg.
IT-95-PO-119, 120: Zapravochyi Waterfall, Konservnaya Bay; 45˚20.01’ N, 147˚59.91’ E; Aug 31, 1995; P. Oberg, N.
Minakawa, D.E. Hoekstra.
IT-95-PO-121: Stream that feeds into Konservnaya Bay, near Zapravochyi Waterfall; 45˚19.92’ N, 147˚59.91’; Aug 31,
1995; N. Minakawa.
IT-95-VAT-04: First stream that flows into Nezhnyi River, about 4 km north of Kitovyi Village, environs of Nezhnaya Hill;
45˚17.63’ N, 147˚52.55’ E; Aug 3, 1995; V. A. Teslenko.
IT-95-VAT-05, 06, 07: Nezhnyi River about 4 km north of Kitovyi Village; 45˚16.71’ N, 147˚52.29’ E; Aug 3, 1995; V. A.
Teslenko.
IT-95-VAT-67, 68: Zapravochyi Waterfall, Konservnaya Bay; 45˚20.01’ N, 147˚59.91’ E; Aug 31, 1995; V. A. Teslenko.
IT-96-BKU-86: Inland coastal margin of Dobroye Nachalo Bay, along foot trail; 44˚45.93’ N, 147˚10.91’ E; Aug 22, 1996;
B. K. Urbain.
IT-96-BKU-87: On ship, Akademic Oparin, anchored in Dobroye Nachalo Bay; 44˚45.42’ N, 147˚10.67’ E; Aug 22, 1996;
B. K. Urbain.
IT-96-MO-52: On ship, Akademic Oparin, anchored in Dobroye Nachalo Bay; 44˚44.61’ N, 147˚10.50’ E; Aug 22, 1996;
M. Ohara.
IT-96-NM-19: Stream that flows into Konservnaya Bay, near Zapravochyi Waterfall; 45˚19.94’ N, 147˚59.76’ E; Aug 18,
1996; N. Minakawa.
IT-96-NM-29: Dobroye Lake, Dobroye Nachalo Bay; 44˚44.23’ N, 147˚13.13’ E; Aug 23, 1996; N. Minakawa.
IT-96-PO-58: Stream that feeds into Konservnaya Bay, near Zapravochyi Waterfall; 45˚19.79’ N, 147˚59.75’ E; Aug 19,
1996; P. Oberg, N. Minakawa.
IT-96-PO-59: Stream that feeds into Konservnaya Bay, near Zapravochyi Waterfall; 45˚19.94’ N, 147˚59.76’ E; Aug 19,
1996; N. Minakawa.
IT-96-PO-69, 75: Lake Natasha, about 1 km Dobroye Nachalo Bay; 44˚46.21’ N, 147˚11.40’ E; Aug 22, 1996; P. Oberg, N.
Minakawa.
IT-96-VAT-47: Lake Natasha, Dobroye Nachalo Bay; 44˚46.21’ N, 147˚11.40’ E; Aug 22, 1996; V. A. Teslenko.
IT-96-VAT-48: Lake Natasha, Dobroye Nachalo Bay; 44˚46.27’ N, 147˚11.40’ E; Aug 22, 1996; V. A. Teslenko.
IT-96-VAT-49: Dobroye Lake, Dobroye Nachalo Bay; 44˚44.23’ N, 147˚13.13’ E; Aug 23, 1996; V. A. Teslenko.
IT-96-VAT-50: Dobroye Lake, Dobroye Nachalo Bay; 44˚44.23’ N, 147˚13.13’ E; Aug 23, 1996; V. A. Teslenko.
IT-97-BKU-14: Stream about 4 km east of Kitovyi, along side road (off main road) that leads down to abandoned fish
hatchery along Podoshevka River; 45˚15.90’ N, 147˚55.90’ E; Jul 29, 1997; B. K. Urbain.
IT-97-BKU-19: Coastal margin of Konservnaya Bay; 45˚20.04’ N, 147˚59.84’ E; Jul 30, 1997; B. K. Urbain.
IT-97-NM-06: Podoshevka River about 4 km east of Kitovyi, along side road (off main road) that leads down to abandoned
fish hatchery; 45˚15.87’ N, 147˚55.71’ E; Jul 29, 1997; N. Minakawa.

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IT-97-NM-12: Podoshevka River about 4 km east of Kitovyi, along side road (off main road) that leads down to abandoned
fish hatchery; 45˚15.87’ N, 147˚55.89’ E; Jul 29, 1997; N. Minakawa.
IT-97-NM-13: Stream that flows into Konservnaya Bay; 45˚20.04’ N, 147˚59.80’ E; Jul 30, 1997; N. Minakawa.
IT-97-NM-19: Zapravochyi Waterfall, Konservnaya Bay; 45˚19.84’ N, 147˚59.73’ E; Jul 30, 1997; N. Minakawa.
IT-97-NM-20: Stream that flows into Konservnaya Bay; 45˚20.04’ N, 147˚59.80’ E; Jul 31, 1997; N. Minakawa.
IT-97-NM-21: On ship, Professor Bogorov, anchored in Konservnaya Bay, near Zapravochyi Waterfall; 45˚19.81’ N,
147˚59.83’ E; Jul 31, 1997; N. Minakawa.
IT-97-NM-84: Stream that flows into Konservnaya Bay, near Zapravochyi Waterfall; 45˚20.04’ N, 147˚59.80’ E; Aug 22,
1997; N. Minakawa.
IT-97-NM-85: Zapravochyi Waterfall, Konservnaya Bay; 45˚19.85’ N, 147˚59.68’ E; Aug 22, 1997; N. Minakawa.
IT-97-TIR-05, 06: Podoshevka River, about 4 km east of Kitovyi by road, near abandoned fish hatchery, 0.5 km upriverfrom road bridge; 45˚15.82’ N, 147˚55.69’ E; Jul 29, 1997; T. I. Ritchie.
IT-97-VAT-06: Podoshevka River about 4 km east of Kitovyi by road, near abandoned fish hatchery, about 1-1.5 km
upstream from hatchery; 45˚15.87’ N, 147˚55.71’ E; Jul 29, 1997; V. A. Teslenko.
IT-97-VAT-08: Near Zapravochnyi Waterfall, coastal margin of Konservnaya Bay; Jul 30, 1997; V. A. Teslenko.
IT-98-BKU-26: North shore of Lake Sredney, northern end of Kasatka Bay; 44˚58.83’ N, 147˚44.14’ E; Jul 31, 1998; B. K.
Urbain.
IT-98-BKU-60: Stream at northern end of sandy coastline, Medvezhya Bay; 45˚27.50’ N, 148˚49.96’ E; Aug 5, 1998; B. K.
Urbain.
IT-98-DJB-32, 34: Near Lake Sredney, northern end of Kasatka Bay; 44˚58.53’ N, 147˚44.38’ E; Jul 31, 1998; D. J. Bennett.
IT-98-DJB-40, 41, 42: Near Kasatka Lake, northeastern part of Kasatka Bay; 45˚00.45’ N, 147˚43.68’ E; Aug 1, 1998; D. J.
Bennett.
IT-98-DJB-48: Coastal margin, Sernozavoskaya Bay; 44˚58.10’ N, 147˚53.71’ E; Aug 2, 1998; D. J. Bennett.
IT-98-DJB-63: Near mouth of Medyezh’ya River, Medyezhya Bay; 45˚26.70’ N, 148˚49.68’ E; Aug 5, 1998; D. J. Bennett.
IT-98-DJB-69: Coastal stream that flows into Medvezhya Bay; 45˚25.56’ N, 148˚49.87’ E; Aug 5, 1998; D. J. Bennett.
IT-98-DJB-71: Bog, Slavnaya Bay; 45˚29.47’ N, 148˚37.13’ E; Aug 6, 1998; D. J. Bennett.
IT-98-LJW-11, 12: Coastal stream near Cape Trikhpaly and Usach River; 44˚28.65’ N, 146˚59.92’ E; Jul 29, 1998; L. J.
Weis, N. Minakawa.
IT-98-LJW-13: Usach River, near Cape Trikhpaly; 44˚28.84’ N, 147˚00.11’ E; Jul 29, 1998; L. J. Weis, N. Minakawa.
IT-98-LJW-15: Khvoinaya River near Burevestnik Village; 44˚55.87’ N, 147˚36.33’ E; Jul 30, 1998; L. J. Weis, N. Minakawa.
IT-98-LJW-16: Sredney Lake, eastern Kasatka Bay; 44˚58.89’ N, 147˚44.08’ E; Jul 31, 1998; L. J. Weis.
IT-98-LJW-18: Stream that flows into eastern Kasatka Bay; 45˚00.10’ N, 147˚43.71’ E; Aug 1, 1998; L. J. Weis.
IT-98-LJW-19: Marsh, eastern Kasatka Bay; 45˚00.67’ N, 147˚43.35’ E; Aug 1, 1998; L. J. Weis.
IT-98-LJW-20: Blagodamyo Lake, east side of Kasatka Bay; 45˚01.06’ N, 147˚43.29’ E; Aug 1, 1998; L. J. Weis.
IT-98-LJW-21: Stream that flows into northeastern Kasatka Bay; 45˚01.06’ N, 147˚43.29’ E; Aug 1, 1998; L. J. Weis.
IT-98-LJW-22: Stream that flows into Sernozavoskaya Bay; 44˚58.06’ N, 147˚53.77’ E; Aug 2, 1998; L. J. Weis.
IT-98-LJW-23: Stream that flows into Sernozavoskaya Bay; 44˚58.16’ N, 147˚53.75’ E; Aug 2, 1998; L. J. Weis.
IT-98-LJW-24: Stream that flows into Zorkaya Bay; 45˚16.47’ N, 148˚30.19’ E; Aug 4, 1998; L. J. Weis.
IT-98-LJW-25: Stream that flows into Zorkaya Bay; 45˚16.82’ N, 148˚30.42’ E; Aug 4, 1998; L. J. Weis.
IT-98-LJW-27, 28: Medvezhya River, Medvezhya Bay; 45˚25.96’ N, 148˚49.63’ E; Aug 5, 1998; L. J. Weis.
IT-98-LJW-29, 30: Medvezhya River, Medvezhya Bay; 45˚26.19’ N, 148˚49.64’ E; Aug 5, 1998; L. J. Weis.
IT-98-LJW-31: Medvezhya River, Medvezhya Bay; 45˚25.56’ N, 148˚49.87’ E; Aug 5, 1998; L. J. Weis.
IT-98-LJW-32, 33: Kumushka River by Aktivnyi settlement / hatchery station, Slavnaya Bay; 45˚28.90’ N, 148˚39.07’ E;
Aug 6, 1998; L. J. Weis.
IT-98-LJW-34, 35: Stream that feeds into Konservnaya Bay, near Zapravochyi Waterfall; 45˚19.68’ N, 147˚59.89’ E; Aug7, 1998; L. J. Weis.
IT-98-LJW-36: Springs near, Zapravochyi Waterfall, Konservnaya Bay; 45˚19.77’ N, 147˚59.74’ E; Aug 7, 1998; L. J.
Weis.
IT-98-LJW-37: Lesozavodskoye Lake, Dobroye Nachalo Bay; 44˚46.07’ N, 147˚11.20’ E; Aug 10, 1998; L. J. Weis.
IT-98-NM-05: Tire truck puddles, northern end of Kasatka Bay; 44˚58.77’ N, 147˚44.20’ E; Jul 31, 1998; N. Minakawa.
IT-98-NM-06: Sredney Lake, eastern Kasatka Bay; 44˚58.40’ N, 147˚44.59’ E; Jul 31, 1998; N. Minakawa.
IT-98-NM-10: Kasatka Lake, northeastern Kasatka Bay; 45˚00.72’ N, 147˚43.30’ E; Aug 1, 1998; N. Minakawa.
IT-98-NM-11: Blagodamyo Lake, northeastern Kasatka Bay; 45˚01.36’ N, 147˚43.43’ E; Aug 1, 1998; N. Minakawa.
IT-98-NM-13: Blagodamyo Lake, northeastern Kasatka Bay; 45˚01.78’ N, 147˚43.56’ E; Aug 1, 1998; N. Minakawa.
IT-98-NM-14: Blagodamyo Lake, northeastern Kasatka Bay; 45˚02.15’ N, 147˚43.81’ E; Aug 1, 1998; N. Minakawa.
IT-98-NM-15: Stream between Sredney Lake and Blagodamyo Lake, northeastern Kasatka Bay; 45˚01.06’ N, 147˚43.29’
E; Aug 1, 1998; N. Minakawa.
IT-98-NM-16: Stream that flows into Sernozavoskaya Bay; 44˚58.05’ N, 147˚53.86’ E; Aug 2, 1998; N. Minakawa.
IT-98-NM-19: Zorky River, Zorkaya Bay; 45˚16.38’ N, 148˚30.12’ E; Aug 4, 1998; N. Minakawa.
IT-98-NM-20: Lugovoy River, Zorkaya Bay; 45˚16.93’ N, 148˚30.42’ E; Aug 4, 1998; N. Minakawa.
IT-98-NM-23, 37: Stream that flows into Medyezhya Bay, north of Medyezhya River; 45˚27.88’ N, 148˚50.30’ E; Aug 5,
1998; N. Minakawa.
IT-98-NM-24: Stream that flows into Dozorny Bay; 44˚32.63’ N, 146˚56.93’ E; Aug 10, 1998; N. Minakawa.
IT-98-TIA-31: Unnamed stream inland from Medvezh’ya Bay; 45°25.53’ N; 148°49.76’ E; Aug 8, 1998; T. I. Arefina.
IT-98-VAT-09, 10: Stream with waterfall 1.5 km north from Usach River, near cape Triokhpaly; 44˚28.14’ N, 1; 46˚59.70’
E; Jul 29, 1998; V. A. Teslenko.

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IT-98-VAT-11, 12: Khvoinaya River about 1 km from the bridge; 44˚55.89’ N, 147˚36.01’ E; Jul 30, 1998; V. A. Teslenko.
IT-98-VAT-13: Ponds near Kasatka River mouth, northern end of Kasatka Bay; 44˚58.42’ N, 147˚44.55’ E; Jul 31, 1998; V.
A. Teslenko.
IT-98-VAT-14: Tichiy Creek that flows into Sredney Lake, northern end of Kasatka Bay; 44˚58.39’ N, 147˚44.59’ E; Jul 31,
1998; V. A. Teslenko.
IT-98-VAT-15: Northeast part of Sredney Lake, northern end of Kasatka Bay; 44˚58.40’ N, 147˚44.59’ E; Jul 31, 1998; V. A.
Teslenko.
IT-98-VAT-16, 17: Blagodamoyo River, about 1 km from mouth, near Yoryachie Kluchi settlement, northeastern Kasatka
Bay; 45˚02.49’ N, 147˚45.58’ E; Aug 1, 1998; V. A. Teslenko.
IT-98-VAT-18: Southeastern part of Blagodamoyo Lake, northeastern Kasatka Bay; Aug 1, 1998; V. A. Teslenko.
IT-98-VAT-21: Otkoshaya River, about 1 km from mouth, between Cape Kanonerka and Cape Yevgeniya; 45˚10.00’ N,
148˚12.76’ E; Aug 3, 1998; V. A. Teslenko.
IT-98-VAT-22, 23: Zorky River, 2 km from mouth, environs of Sentyabrskiy, Zorkaya Bay; 45˚16.29’ N, 148˚28.90’ E; Aug
4, 1998; V. A. Teslenko.
IT-98-VAT-24: Lugovoy River, below the waterfall, environs of Sentyabrskiy, Zorkaya Bay; l; Aug 4, 1998; V. A. Teslenko.
IT-98-VAT-26, 27: Stream west side of Medyezhya Bay; 45˚25.53’ N, 148˚49.76’ E; Aug 5, 1998; V. A. Teslenko.
IT-98-VAT-28: Small stream between the stream on the west of Medyezhya Bay and Medyezh’ya River; Aug 5, 1998; S. Y.
Storozhenko.
IT-98-VAT-29, 30: Kumushka River by Aktivnyi settlement / hatchery station, Slavnaya Bay; 45˚28.90’ N, 148˚39.07’ E;
Aug 6, 1998; V. A. Teslenko.
IT-98-VAT-31, 32: Zapravochyi Waterfall, Konservnaya Bay, 45˚19.85’ N, 147˚59.68’ E; Aug 7, 1998; V. A. Teslenko.
IT-98-VAT-33, 34, 35: Small waterfall next to Zapravochyi Waterfall, Konservnaya Bay; 45˚19.77’ N, 147˚59.74’ E; Aug 7,
1998; V. A. Teslenko.
IT-98-VAT-36: Lake Natasha, Dobroye Nachalo Bay, environs of Lesozavodskiy; 44˚46.21’ N, 147˚11.40’ E; Aug 10,
1998; V. A. Teslenko.
IT-98-VAT-37, 38: Stryi Creek, Dozorny Bay; 44˚32.57’ N, 1; 46˚57.09’ E; Aug 10, 1998; V. A. Teslenko.
IT-99-DJB-93: Coastal margin, northern end of Dobroye Nachalo Bay, near military building; 44˚46.01’ N, 147˚11.58’ E;
Aug 14, 1999; D. J. Bennett.
IT-99-DJB-97: Inland from beach along road up to Tichaya River (1.41 km from beach, northern end of Dobroye Nachalo
Bay; 44˚43.38’ N, 147˚12.77’ E; Aug 15, 1999; D. J. Bennett.
IT-99-DJB-99: Inland from beach, along start of road to Lake, Dobroye Nachalo Bay; 44˚43.82’ N, 147˚11.90’ E; Aug 15,
1999; D. J. Bennett.
IT-99-KLK-43: Kuybyshevskiy Lake, Kuybyshevskiy Bay; 45˚04.21’ N, 147˚39.09’ E; Aug 13, 1999; N. Minakawa, K. L.
Kurowski.
IT-99-KLK-44, 45: Kuybyshevskiy Lake, Kuybyshevskiy Bay; 45˚04.51’ N, 147˚41.98’ E; Aug 13, 1999; N. Minakawa, K.
L. Kurowski.
IT-99-NM-13, 14: Southeastern side of Lake Natasha, northern end of Dobroye Nachalo Bay, near Lesozavodskoy; 44˚46.21’
N, 147˚11.40’ E; Aug 14, 1999; N. Minakawa.
IT-99-NM-15: Southwestern side of Dobrye Lake, Dobroye Nachalo Bay; 44˚44.24’ N, 147˚13.44’ E; Aug 15, 1999; N.
Minakawa.
IT-99-VAT-46: Northwestern part of Kuybyshevskiy Lake, Kuybyshevskiy Bay; 45˚04.34’ N, 147˚39.03’ E; Aug 13, 1999;
V. A. Teslenko.
IT-99-VAT-50: Eastern side of Lesozavodskoye Lake, Dobroye Nachalo Bay; 44˚46.43’ N, 147˚12.84’ E; Aug 14, 1999; V.
A. Teslenko.
IT-99-VAT-52: Southwestern side of Dobroye Lake, near Tichaya River flows from the Lake, Dobroye Nachalo Bay;44˚44.24’ N, 147˚13.44’ E; Aug 15, 1999; V. A. Teslenko.
IT-99-VAT-54: Bolshoye Stream, near the bridge over the stream, about 2 km from the mouth, Dobroye Nachalo Bay;44˚43.69’ N, 147˚14.34’ E; Aug 15, 1999; V. A. Teslenko.

KUNASHIR
KU-94-NM-01: Northern and eastern shore of Lake Aliger; 44˚02’48" N, 145˚44’24" E; Jul 31, 1994; N. Minakawa, R. I.
Gara.
KU-94-NM-04: Lower stream section between Lake Bezymaynnoye and Lake Lagunnoye; 44˚03’06" N, 145˚45’48" E;
Jul 31, 1994; N. Minakawa, R. I. Gara.
KU-94-NM-05: About 150 m from mouth of Ilyushina River; 44˚09’18" N, 145˚56’30" N; Aug 1, 1994; N. Minakawa, R.
I. Gara.
KU-94-NM-07: About 300 m from mouth of Ilyushina River; 44˚09’30" N, 145˚56’12" E; Aug 1, 1994; N. Minakawa, R.
I. Gara.
KU-94-NM-08: About 350 m from mouth of Ilyushina River; 44˚09’30" N, 145˚56’12" E; Aug 1, 1994; N. Minakawa, R.
I. Gara.
KU-94-NM-10: About 450 m from mouth of Ilyushina River; 44˚09’42" N, 145˚56’00" E; Aug 1, 1994; N. Minakawa, R.
I. Gara.
KU-94-NM-11, 13: Tributary of River Rikorda, northern bifurcation, just off east side of road; 43˚51’ N, 145˚33’ E; Aug 2,
1994; N. Minakawa, R. I. Gara.
KU-94-NM-16: Stream section above geothermal pools, 14 km west of Yuzhno-Kurilsk; 44˚00’48" N, 145˚41’06" E; Aug
3, 1994; N. Minakawa, R. I. Gara.

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KU-94-NM-17: Prozrachnyi River, near broken coastal road bridge, 44˚05’42" N, 145˚53’18" E; Aug 3, 1994; N. Minakawa,
R. I. Gara.
KU-94-NM-18: Lesnaya River, at roadside entrance for Kislyi Hot Springs, near junction with Kislyi river; 44˚01.4’ N,
145˚44.1’ E; Aug 4, 1994; N. Minakawa, R. I. Gara.
KU-94-NM-21: Western and southern shore of Lake Krugloye; 44˚22’24" N, 146˚25’12" E; Aug 23, 1994; N. Minakawa,
R. I. Gara.
KU-94-TWP-07: Ilyushina River, about 350 m from mouth, in isolated, shallow, south branch; 44˚09’30" N, 145˚56’12" E;
Jan 8, 1994; T. W. Pietsch, W. A. Palsson, B. K. Urbain, J. A. Lopez.
KU-94-VAT-01, 02: Western shore of Lake Aliger; 44˚02’48" N, 145˚44’24" E; Jul 31, 1994; V. A. Teslenko.
KU-94-VAT-03: Northern shore of Lake Aliger; 44˚02’48" N, 145˚44’24" E; Jul 31, 1994; V. A. Teslenko.
KU-94-VAT-04: Eastern shore of Lake Aliger; 44˚02’48" N, 145˚44’24" E; Jul 31, 1994; V. A. Teslenko.
KU-94-VAT-06, 07, 08: Ilyushina River, near mouth; 44˚09’18" N, 145˚56’30" N; Aug 1, 1994; V. A. Teslenko.
KU-94-VAT-09, 10, 11, 12: Tributary of River Rikorda, northern bifurcation, near main road; 43˚51’ N, 145˚33’ E; Aug 2,
1994; V. A. Teslenko.
KU-94-VAT-14, 15: Stream adjacent to geothermal pools, 14 km west of Yuzhno-Kurilsk; 44˚00.48’ N, 145˚41.06’ E; Aug
3, 1994; V. A. Teslenko.
KU-94-VAT-16, 17: Prozrachnyi River, about 1 km from mouth; 44˚05’42" N, 145˚53’18" E; Aug 3, 1994; V. A. Teslenko.
KU-94-VAT-19, 20: Lesnaya River, about 1km upriver from junction with Kislyi River; at roadside entrance for Kislyi Hot
Springs; 44˚01.4 N, 145˚44.8 E; Aug 4, 1994; V. A. Teslenko.
KU-94-VAT-55: Western and southern shore of Lake Krugloye; 44˚22’24" N, 146˚25’12" E; Aug 23, 1994; V. A. Teslenko.
KU-94-VAT-56: Stream that flows into western part of Lake Krugloye; 44˚22.30’ N, 146˚24.90’ E Aug 23, 1994; V. A.
Teslenko.
KU-95-PO-02, 03: Stream adjacent to geothermal pools, 14 km west of Yuzhno-Kurilsk; 44˚00.47’ N, 145˚40.98’ E; Aug 2,
1995; N. Minakawa, P. Oberg.
KU-95-PO-123: Lake Aliger; 44˚02.83’ N, 145˚44.47’ E; Sep 1, 1995; P. Oberg.
KU-95-PO-127: Stream that flows into Lake Lagunnoye; 44˚03.06’ N, 145˚45.48’ E; Sep 1, 1995; N. Minakawa.
KU-95-PO-128: Lesnaya River; about 200 m upriver from Lesnaya-Kislyi River merge point; 44˚00.10’ N, 145˚46.00’ E;
Sep 2, 1995; N. Minakawa, P. Oberg, D. E. Hoekstra.
KU-95-VR-38: Aliger Lake; 44˚03.13N, 145˚44.68E; Sep 1, 1995; V. Roth.
KU-95-VAT-01, 02, 03: Stream adjacent to geothermal pools, 14 km west of Yuzhno-Kurilsk; 44˚00.47’ N, 145˚40.98’ E;
Aug 2, 1995; V. A. Teslenko.
KU-95-VAT-69: Lesnaya River, near junction with Kislaya River; 44˚00.72’ N, 145˚46.28’ E; Sep 1, 1995; V. A. Teslenko.
KU-95-VAT-71: Lesnaya River, near junction with Kislaya River; 44˚00.10’ N, 145˚46.00’ E; Sep 1, 1995; V. A. Teslenko.
KU-96-NM-36, 43: Stream adjacent to geothermal pools, 14 km west of Yuzhno-Kurilsk; 44˚00.25’ N, 145˚40.34’ E; Aug
25, 1996; N. Minakawa, P. Oberg.
KU-96-PO-76: Aliger Lake; 44˚02.59’ N, 145˚44.18’ E; Aug 26, 1996; P. Oberg, N. Minakawa.
KU-97-BKU-06: Environs of geothermal pools, 17 km west of Yuzhno-Kurilsk, along side trail (not main trail up to road)that leads off from the northwest; 44˚00.47’ N, 145˚41.00’ E; Jul 27, 1997; B. K. Urbain.
KU-97-NM-01, 03: 70 m upstream from geothermal pools, 14 km west of Yuzhno-Kurilsk; 44˚00.34’ N, 145˚40.99’ E; Jul
27, 1997; N. Minakawa.
KU-97-TIR-01: 0-100 m upstream from geothermal pools, 14 km west of Yuzhno-Kurilsk; 44˚00.47’ N, 145˚40.94’ E; Jul
27, 1997; T. I. Ritchie.
KU-97-TIR-04: 0-200 m upstream from geothermal pools, 14 km west of Yuzhno-Kurilsk; 44˚00.34’ N, 145˚40.99’ E; Jul
27, 1997; T. I. Ritchie.
KU-97-VAT-01, 02, 03: Lesnaya River, about 1 km upstream from its junction with Kislyi River; 44˚00.52’ N, 145˚45.45’
E; Jul 27, 1997; V. A. Teslenko.
KU-97-VAT-05: About 1 km inland from shore of Yuzhno-Kurilsk; 44˚01.52’ N, 145˚50.23’ E; Jul 27, 1997; A.S. Lelej.
KU-98-BKU-73: Just off road near Lake Aliger; 44˚03.17’ N, 145˚45.00’ E; Aug 11, 1998; B. Urbain.
KU-98-DJB-79: Lake Aliger; 44˚02.88’ N, 145˚44.48’ E; Aug 11, 1998; D. Bennett.
KU-98-DJB-140: Along road leading to Lake Serebryanoye, about 3 km northwest of Yuzhno-Kurilsk; 44˚03.15’ N,
145˚49.14’ E; Aug 22, 1998; D. J. Bennett.
KU-98-TIA-65: Tributary of Kislyi River; Aug 22, 1998; T. I. Arefina.
KU-98-LJW-01: Upstream from geothermal pools, 14 km west of Yuzhno-Kurilsk; 44˚00.41’ N, 145˚40.96’ E; Jul 26,
1998; L. J. Weis, N. Minakawa.
KU-98-LJW-02, 04: Downstream from geothermal pools, 14 km west of Yuzhno-Kurilsk; 44˚00.41’ N, 145˚40.96’ E; Jul
26, 1998; L. J. Weis, N. Minakawa.
KU-98-LJW-06: Stream that flows into Dlinnoye Lake, Lovtsova Peninsula, northern part of Kruglovsky Isthmus; 44˚24.74’
N, 146˚25.62’ E; Jul 28, 1998; L. J. Weis, N. Minakawa.
KU-98-LJW-09, 10: Stream that flows into Dlinnoye Lake, Lovtsova Peninsula, northern Kruglovsky Isthmus; 44˚24.02’
N, 148˚25.33’ E; Jul 28, 1998; L. J. Weis, N. Minakawa.
KU-98-LJW-40, 41: Lesnaya River and Kislyi River near Kislyi Hot Springs; 44˚00.87’ N, 145˚46.02’ E; Aug 11, 1998; L.
J. Weis.
KU-98-LJW-74: Lake Serebryanoye, about 3km northwest of Yuzhno-Kurilsk; 44˚03.21’ N, 145˚48.92’ E; Aug 22, 1998;
L. J. Weis, N. Minakawa.
KU-98-LJW-75: Lake Serebryanoye, about 3km northwest of Yuzhno-Kurilsk; 44˚03.09’ N, 145˚48.95’ E; Aug 22, 1998;
L. J. Weis, N. Minakawa.

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KU-98-VAT-01, 02: Stream adjacent to geothermal pools, 14 km west of Yuzhno-Kurilsk; 44˚00.56’ N, 145˚40.91’ E; Jul
26, 1998; V. A. Teslenko.
KU-98-VAT-06: Stream that flows into western part of Dlinnoye Lake, Lovstova Peninsula, northern Kruglovsky Isthmus;
44˚24.74’ N, 146˚25.62’ E; Jul 28, 1998; V. A. Teslenko.
KU-99-DJB-105: Environs of geothermal pools, 14 km west of Yuzhno-Kurilsk, base of trail in hot springs clearing andsurrounding area; 44˚00.39’ N, 145˚41.01’ E; Aug 17, 1999; D. J. Bennett.
KU-99-KLK-49: Swamp near shore of Alekhina Bay; 43˚55.14’ N, 145˚32.34’ E; Aug 19, 1999; N. Minakawa, K. L.
Kurowski.
KU-99-KLK-50, 51: Alekhina River, about 1 km from the mouth, Alekhina Bay; 43˚55.19’ N, 145˚32.47’ E; Aug 19, 1999;
N. Minakawa, K. L. Kurowski.
KU-99-KLK-52: Side pool of Alekhina River, about 0.7 km from the mouth, Alekhina Bay; 43˚55.19’ N, 145˚32.47’ E;
Aug 19, 1999; N. Minakawa, K. L. Kurowski.
KU-99-NM-18: Stream that flows into Peshcnoye Lake; 43˚55.02’ N, 145˚37.83’ E; Aug 17, 1999; N. Minakawa.
KU-99-VAT-56: Eastern side of Peschanoye Lake; 43˚55.15’ N, 145˚37.91’ E; Aug 15, 1999; V. A. Teslenko.
KU-99-VAT-57: Small stream that flows from Peschanoye Lake; 43˚55.15’ N, 145˚37.91’ E; Aug 15, 1999; V. A. Teslenko.
KU-99-VAT-62: Alekhina River, about 0.8 km from the mouth, Alekhina Bay: 43˚55’ N, 145˚32’ E; Aug 19, 1999; V. A.
Teslenko.
KU-99-VAT-63: Alekhina River, about 1 km from the mouth, Alekhina Bay: 43˚55.06’ N, 145˚32.38’ E; Aug 19, 1999; V.
A. Teslenko.

SHIKOTAN
SH-94-RG-01: Tributary about 3.5 km from mouth of Svobodnaya River, Otradnaya Bay; Aug 8, 1994; N. Minakawa, R. I.
Gara.
SH-94-RG-03: Coastal stream that empties into Posodebari Inlet of Delfin Bay, between 0-75 m upstream from mouth; Aug
11, 1994; R. I. Gara, N. Minakawa.
SH-94-TWP-03: Svobodnaya River, near river mouth in Otradnaya Bay, between 0-200 m upriver from Ostrovnoye Milk
Farm; 43˚51’06" N, 146˚48’36" E; Aug 8, 1994; T.W. Pietsch, W.A. Palsson, B. K. Urbain, J.A. Lopez.
SH-94-VAT-28, 29: Svobodnaya River, about 5 km from mouth, at confluence of two streams, Otradnaya Bay; Aug 8,
1994; V. A. Teslenko.
SH-94-VAT-31, 32: Coastal stream that flows into Posodebari Inlet of Delfin Bay, between 0-75 m upstream from mouth;
Aug 11, 1994; V. A. Teslenko.
SH-98-BKU-91: Meadow, inland from small, unnamed inlet about 2/3 way in, on the south side of Delfin Bay; 43˚45.03’ N,
146˚37.36’ E; Aug 15, 1998; B. Urbain.
SH-98-DJB-93: Inland from Dimitrova Bay, ridges, hillsides, and valleys within 1 km from shore; 43˚47.69’ N, 146˚49.64’
E; Aug 13, 1998; D. Bennett.
SH-98-DJB-99: Inland from small inlet about 2/3 way in, on the south side of Delfin Bay, upland from stream valley;43˚44.96’ N, 146˚37.75’ E; Aug 15, 1998; D. Bennett.
SH-98-LJW-46: Stream that flows into Otradnaya Bay; 43˚50.44’ N,146˚48.89’ E; Aug 12, 1998; L. J. Weis.
SH-98-LJW-50: Stream that flows into Tserkovnaya Bay; 43˚44.55’ N, 146˚42.76’ E; Aug 14, 1998; L. J. Weis.
SH-98-LJW-54, 55: Stream that flows into Zvezdnaya Bay; 43˚46.73’ N, 146˚36.49’ E; Aug 16, 1998; L. J. Weis.
SH-98-LJW-56, 57: Stream that flows into Zvezdnaya Bay; 43˚46.44’ N, 146˚36.58’ E; Aug 16, 1998; L. J. Weis.
SH-98-LJW-60: Stream that flows into Gorobets Bay; 43˚49.18’ N, 146˚42.58’ E; Aug 18, 1998; L. J. Weis, N. Minakawa.
SH-98-LJW-61, 62: Marsh and ponds, inland from Gorobets Bay; 43˚49.05’ N, 146˚42.68’ E; Aug 18, 1998; L. J. Weis, N.
Minakawa.
SH-98-LJW-63: Marsh and ponds, inland from Gorobets Bay; 43˚48.02’ N, 146˚42.95’ E; Aug 18, 1998; L. J. Weis, N.
Minakawa.
SH-98-NM-30, 33: Stream that flows into Tserkovnaya Bay; 43˚44.83’ N, 146˚41.98’ E, Aug 14, 1998, N. Minakawa.
SH-98-NM-35: Near the mouth of Ostrovnaya River, Delfin Bay; 43˚45.11’ N, 146˚37.41’ E; Aug 15, 1998; N. Minakawa.
SH-98-VAT-42: Small stream that flows into Khromova Bay in southern Malokurilsk; 43˚52.09’ N, 146˚48.52’ E; Aug 12,
1998; V. A. Teslenko.
SH-98-VAT-45: Small stream that flows into Dimitrova Bay, west of Solov’Oeva Cape; 43˚47.64’ N, 146˚48.63’ E; Aug 13,
1998; V. A. Teslenko.
SH-98-VAT-47: Stream that flows into northeastern Srezhkova Bay; 43˚47.59’ N, 146˚48.44’ E; Aug 13, 1998; V. A. Teslenko.
SH-98-VAT-49: Small pond, inland from Tserkovnaya Bay, before river from Tomari Mountain, the river flows into thecentral part of the bay; 43˚44.85’ N, 146˚41.61’ E; Aug 14, 1998; V. A. Teslenko.
SH-98-VAT-50: Small drying pond inland from central part of Tserkovnaya Bay; 43˚44.60’ N, 146˚41.50’ E; Aug 14, 1998;
V. A. Teslenko.
SH-98-VAT-51: Small stream that flows into central part of Tserkovnaya Bay; 43˚44.77’ N, 146˚41.49’ E; Aug 14, 1998; V.
A. Teslenko.
SH-98-VAT-53: First tributary on the left side of Ostrovnaya River from the mouth, south-central part of Delfin Bay;43˚44.90’ N, 146˚39.00’ E; Aug 15, 1998; V. A. Teslenko.

POLONSKOGO
PO-98-DJB-135: Inland from western side; 43°38.38’ N, 146°18.58’ E; Aug 21, 1998; D. J. Bennett.
PO-98-TIA-61, 64: Inland from western side, small unnamed lake, Moryakov Bay; Aug 21, 1998; T. I. Arefina.

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ZELIONYI
ZE-94-NM-02: Lake Utinoye, western finger of lake, northern shore; 43˚29’24" N, 146˚06’48" E; Aug 5, 1994; R. I. Gara,
N. Minakawa.
ZE-94-NM-03: Stream that flows into northeastern Lake Utinoye; 43˚29’36" N, 146˚07’54" E; Aug 6, 1994; R. I. Gara, N.
Minakawa.
ZE-94-NM-05: Southeastern shore of Lake Kamenskoye; 43˚30’12" N, 146˚06’12" E; Aug 6, 1994; R. I. Gara, N. Minakawa.
ZE-94-RLC-04: Lake Utinoye, between 0-50 m west off southwestern shore; 43˚48.0 N, 146˚11.5 E; Aug 5, 1994; R. L.
Crawford.
ZE-94-VAT-21: Stream that flows into northwestern Lake Utinoye; 43˚29’12" N, 146˚06’42" E; Aug 5, 1994, 1994; V. A.
Teslenko.
ZE-94-VAT-24: Lake Utinoye, western finger of lake, northern shore; 43˚29’24" N, 146˚06’48" E; Aug 5, 1994; V. A.
Teslenko.
ZE-94-VAT-25: Stream that flows into Lake Utinoye at north northeastern corner; 43˚29’36" N, 146˚07’54" E; Aug 6,
1994; V. A. Teslenko.
ZE-94-VAT-27: Southwestern shore of Lake Kamenskoye; 43˚30’12" N, 146˚06’12" E; Aug 6, 1994; V. A. Teslenko.
ZE-98-LJW-70: Marsh near Lake Utinoye; 43˚29.45’ N, 146˚06.54’ E; Aug 20, 1998; L. J. Weis, N. Minakawa.

IURII
IU-98-TIA-59: Lake near the sea shore, inland from Shyrokaya Bay; Aug 20, 1998; T. I. Arefina.

ANUCHINA
AN-98-TIA-55: Stream, inland from Boshoy Bay, northern part of the island; Aug 19, 1998; T. I. Arefina.

TANFILYEVA
TA-98-LJW-66: Shallow puddles, inland from Tanfilyevka Bay; 43˚26.82’ N, 145˚54.36’ E; Aug 19, 1998; L. J. Weis, N.
Minakawa.
TA-98-VAT-65: Lake near Russian military base, Tanfilyeva Bay; Aug 19, 1998; V. A. Teslenko.
TA-98-TIA-57: Unnamed lake, Tanfilyeva Bay; Aug 19, 1998; T. I. Arefina.
TA-98-TIA-58: Stream that flows into the lake, Tanfilyeva Bay; Aug 19, 1998; T. I. Arefina.

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