Bulletin of the Kagoshima University Museum No. 12

AN ANNOTATED CHECKLIST OF MARINE AND FRESHWATER FISHES OF
OKINOERABU ISLAND IN THE AMAMI ISLANDS,KAGOSHIMA, SOUTHERN JAPAN,
WITH 361 NEW RECORDSHIROYUKI MOTOMURA AND KAZUTOSHI UEHARA

February 2020

The Kagoshima University Museum

Cover photograph: A school of Pseudanthias pleurotaenia (Serranidae) off Okinoerabu Island. Photo by K. Uehara
Back cover photograph: A school of Mulloidichthys vanicolensis (Mullidae) and Gnathodentex aureolineatus (Lethrinidae)
off Okinoerabu Island. Photo by K. UeharaBulletin of the Kagoshima University Museum No.12

ISSN-L 2188-9074

An annotated checklist of marine and freshwater fishes of
Okinoerabu Island in the Amami Islands,
Kagoshima, southern Japan,
with 361 new records

Hiroyuki Motomura1, 3 and Kazutoshi Uehara2

1

The Kagoshima University Museum, 1–21–30 Korimoto, Kagoshima 890–0065, Japan
2

GT Divers Okinoerabu, 471–3 Wadomari, Oshima, Kagoshima 891–9112, Japan
3

Corresponding author, e-mail: motomura@kaum.kagoshima-u.ac.jp

Abstract An annotated checklist of marine and freshwater fishes of
Okinoerabu Island in the Amami Islands, Kagoshima Prefecture, southern
Japan, was compiled from specimen and literature surveys. A total of 637
species (278 genera and 88 families), including 361 species that represent thefirst reliable records from the island on the basis of collected specimens and/or underwater photographs, are listed with citation of literature, registration
numbers, sizes, localities on the island, and some remarks. Two of the 361
newly recorded species from Okinoerabu Island (Hippocampus denise andH. pontohi) are also the first specimen-based records from Japanese waters.
Preliminary comparisons of the ichthyofauna of the island with those of 15
other localities in Japan show that the fish fauna of Okinoerabu Island issimilar to that of Yoron Island in the Amami Islands.

Motomura, H. and K. Uehara. 2020. An annotated checklist of marine and freshwater fishes of Okinoerabu
Island in the Amami Islands, Kagoshima, southern Japan, with 361 new records. Bulletin of the
Kagoshima University Museum, 12: 1–125.
Published on 2 March 2020

2

H. Motomura and K. Uehara

Introduction
Okinoerabu Island, a ladle-shaped island with an area of ca. 94 km2 and a highest point of 240 m above
sea level, is the third largest island in the Amami Islands, which are located at the middle of the Ryukyu Archipelago, southern Japan (Fig. 1). The coastline stretches about 50 km, with well-developed coral reefs, andthere are numerous limestone caves with underground rivers on the island.
Since comprehensive ichthyofaunal surveys of Okinoerabu Island have not been previously completed,
two large-scale Okinoerabu Expeditions were made in June 2016 and October 2018 by more than 50 people
from the Kagoshima University Museum and other institutions to study the island’s ichthyofauna. In addition, several small-scale expeditions led by less than five persons at each survey were made on the island
between 2016 and 2019. About 3,500 specimens of marine and freshwater fishes were collected during the
expeditions. About 1,000 unregistered specimens collected from Okinoerabu Island in 1954–1969 were
found at the Faculty of Fisheries, Kagoshima University, and these were registered in the fish collection of
the Kagoshima University Museum and also used in this study.
Examination of the above-mentioned specimens, published literature (plus examination of specimens usedin the literature), and unpublished underwater photographs taken at Okinoerabu Island confirmed 637 species
of marine and freshwater fishes, including 361 new records, from the island. All species are listed below. The
ichthyofauna of Okinoerabu Island is briefly compared with that of other islands in the Ryukyu, Osumi, andOgasawara islands, and several localities along the Pacific coast of southern Japan.
Material and methods
Sampling methods were described and illustrated in Motomura & Matsuura (2010, 2014). Curatorial procedures for newly collected specimens followed Motomura & Ishikawa (2013). Standard and total lengths
are abbreviated as SL and TL respectively. The systematic arrangement of families generally follows Nelson(2006). Scientific names generally follow Fricke et al. (2020), with some modifications following recently
published or unpublished taxonomic studies. Within families, species are arranged in alphabetical order ofthe scientific name. Standard Japanese names generally follow Nakabo (2013), having been transliteratedusing the Hepburn system. Each species record was compiled from voucher specimens, underwater photographs, and/or published literature. Registration number, number of specimens registered (if more than one),mm in SL or TL, and locality on Okinoerabu Island (Fig. 2) are listed in each species record. Okinoerabu Island has two towns: Wadomari (on the northeast side of the island) and China (southwest side). The localities
of marine and freshwater fishes are indicated as names of “port or beach, area, town (e.g., Kasaishi Beach,
Kibiru, Wadomari)” and “river, area, town (e.g., Amata River, Amata, China)” respectively. The locality isexpressed simply as “Okinoerabu Island” for fish(es) collected far off the island or if the precise locality on
the island is unknown.
Institutional codes used in this paper are as follows: FAKU (Kyoto University Museum, Kyoto, Japan),
HMNH (Hiwa Museum of Natural History, Hiroshima, Japan), KAUM (Kagoshima University Museum,Kagoshima, Japan), KPM (Kanagawa Prefectural Museum of Natural History, Kanagawa, Japan), KUN(Kindai University, Nara, Japan), LICPP (Laboratory of Ichthyology, the Crown Prince’s Palace, Tokyo, Japan), MIKU (Marine Biological Institute of Kyoto University, Wakayama, Japan; specimens currently transferred to FAKU), and OMNH (Osaka Museum of Natural History, Osaka, Japan).
Remarks
The freshwater fish diversity of Okinoerabu Island is very poor because there are only three small rivers
flowing on the ground surface of the island, and most rivers flow underground and/or inside caves. The mostof primary freshwater fishes on the island are introduced species, e.g., three poeciliids Gambusia affinis
affinis (Baird & Girard 1853), Poecilia reticulata Peters 1859 and Xiphophorus maculatus (Günther 1866),
and two cichlids Oreochromis niloticus (Linnaeus 1758) and Tilapia zillii (Gervais 1848). Although Ogawa(1937) and Shibata (1964) confirmed the existence of Carassius sp. (Cyprinidae) in Okinoerabu Island (its

Marine and freshwater fishes of Okinoerabu Island

3

status as native or introduced is unknown), Kano et al. (2016) considered that this species is now extinct
from the island. Ogawa (1937), Shibata (1964) and Yoshigou (2014) reported Misgurnus anguillicaudatus(Cobitidae) from channels at a rice paddy in Kamihirakawa, China and a muddy paddy around the middle
reaches of Amata River, China, but the species was not found from the island during this study. Kano et al.
(2016) suggested it to be an undescribed species native to the island. Macropodus opercularis (Linnaeus
1758) (Osphronemidae) (Fig. 3) is considered to be introduced into Okinoerabu Island in 1937 (Shibata
1964; Yoshigou et al. 2005). Shibata (1964) reported that this species was common around Amata River,
China, although its population in Wadomari decreased due to an agricultural chemical. This species is now
rare in Okinoerabu Island.
No species has been described as new on the basis of a primary type specimen from Okinoerabu Island,
although several new marine species were described from the neighboring islands, suggesting that marine
fishes of Okinoerabu Island have been poorly surveyed and studied.
The number of species in the top 10 most speciose families occurring at Okinoerabu Island are shownin Table 1. Comparable data for 15 localities in southern Japan, including the Sagami Sea, Osezaki (Suruga
Bay), Kushimoto (Kii Peninsula), Kashiwa Island (Shikoku), Iwo and Take islands (Osumi Islands), Yaku Island (Osumi Islands), Amami-oshima Island (Amami Islands), Tokunoshima Island (Amami Islands), Yoron
Island (Amami Islands), Okinawa Island (Okinawa Islands), Ie Island (Okinawa Islands), the Miyako Islands,
Ishigaki Island (Yaeyama Islands), Iriomote Island (Yaeyama Islands), and the Ogasawara Islands, were
presented in Motomura (2016: tables 1, 2), Nakae et al. (2018: table 2) and Mochida & Motomura (2018:table 1). The top three most speciose families in all localities, except for the Sagami Sea, Osezaki (Suruga
Bay), and the Ogasawara Islands, are Gobiidae, Pomacentridae and Labridae. The top three most speciose
families in the Sagami Sea, Osezaki, and the Ogasawara Islands are Gobiidae, Labridae, and Serranidae (thelast family was replaced with Pomacentridae in the other localities) (Motomura 2016). Okinoerabu Island isgeographically located between Tokunoshima and Yoron islands. The ichthyofauna of Okinoerabu Island issimilar to that of the latter two islands among the 15 localities in Japan. The top 10 most speciose families,
and their percentages of number of species in eachfamily to total number of species in Okinoerabu Island
Table 1. Number of species in the top 10 most speciose families
are almost the same with those of Yoron Island (Table
occurring at Okinoerabu Island.
1; Motomura 2016: table 1). The only difference beFamily
No. of species% in total
tween Okinoerabu and Yoron islands is that the family
Gobiidae64
10.1
Syngnathidae (17 species), which is ranked as the 10th
Pomacentridae
54
8.5
most speciose family in Okinoerabu Island, is not presLabridae
42
6.6
ent in Yoron Island where it is replaced with the family
Blennidae
35
5.5
Tripterygiidae (18 species) (Motomura 2016: table 1).
Serranidae
32
5.0
Tripterygiidae (13 species) is ranked as the 15th most
Apogonidae
30
4.7
speciose in Okinoerabu Island. The similarity of ichChaetodontidae
25
3.9
Scorpaenidae
24
3.8
thyofaunas between Okinoerabu and Yoron islands is
Acanthuridae
21
3.3
probably caused by their geographic proximity, historSyngnathidae
172.7
ically similar formative period (formed from a rising
Other families
293
46.0
coral reef) and similar environments, including mainly
Total
637
100.0
covered by corals and lacking large rivers.
List of fishes from Okinoerabu Island
FAMILY ORECTOLOBIDAE
Orectolobus japonicus Regan 1906
[Jpn name: Ose]
Underwater photograph (Pl. 1-A).

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H. Motomura and K. Uehara

FAMILY CARCHARHINIDAE
Triaenodon obesus (Rüppell 1837)[Jpn name: Nemuribuka]
Underwater photograph (Pl. 1-B).
FAMILY SPHYRNIDAE
Sphyrna lewini (Griffith & Smith 1834)[Jpn name: Akashumokuzame]
Underwater photograph (Pl. 1-C).
FAMILY CENTROPHORIDAE
Centrophorus moluccensis Bleeker 1860
[Jpn name: Okinawayajirizame]
Hatooka et al. (2013): Okinoerabu Island.
FAMILY RHYNCHOBATIDAE
Rhynchobatus djiddensis (Forsskål 1775)[Jpn name: Tongarisakatazame]
Underwater photograph (Pl. 1-D).
Remarks: Rhynchobatus djiddensis is in fact endemic to the western Indian Ocean; thus a scientific name ofthis photographed species distributed in Japanese waters is unknown.
FAMILY DASYATIDAE
Neotrygon kuhlii (Müller & Henle 1841)[Jpn name: Yakkoei]
Underwater photograph (Pl. 1-E).
Taeniurops meyeni (Müller & Henle 1841)[Jpn name: Madaraei]
Underwater photograph (Pl. 1-F).
FAMILY MOBULIDAE
Mobula alfredi (Krefft 1868)[Jpn name: Nan-yomanta]
Yamaguchi et al. (2013, as Manta alfredi): Okinoerabu Island.
FAMILY AETOBATIDAE
Aetobatus ocellatus (Kuhl 1823)[Jpn name: Madaratobiei]
Underwater photograph (Pl. 1-G).
FAMILY ANGUILLIDAE
Anguilla marmorata Quoy & Gaimard 1824
[Jpn name: Ounagi]
Ogawa (1937): Okinoerabu Island. Yoshigou et al. (2005b): Ginsuido cave, China. Watanabe et al. (2008):
11 specimens, 261–496 mm TL, Okinoerabu Island. Hatooka (2013): Okinoerabu Island. Yoshigou (2014):
Okinoerabu Island.

Marine and freshwater fishes of Okinoerabu Island

5

FAMILY MURAENIDAE
Echidna nebulosa (Ahl 1789)[Jpn name: Kumoutsubo]
Yoshigou et al. (2005a): HMNH 7820, Shinjo, China. This study: KAUM–I. 90719, 94.4 mm TL, Inobe
Port, Inobe, Wadomari; KAUM–I. 122178, 225.5 mm TL, Beach Rock, Tokudoki, China; KAUM–I. 123513,
412.3 mm TL, Kasaishi Beach, Kibiru, Wadomari; KUN-P 48958, 111.1 mm TL, Ujiji Beach, Ashikiyora,
China.
Echidna polyzona (Richardson 1845)[Jpn name: Shimaarashiutsubo]
Yoshigou et al. (2005a): HMNH 7850, Naikina, Nagamine, Wadomari. This study: KAUM–I. 121906, 151.4mm TL, Okidomari Port, Shimojiro, China.
Enchelycore pardalis (Temminck & Schlegel 1846)[Jpn name: Torautsubo]
KAUM–I. 36355, 77.7 mm TL, off Okinoerabu Island.
Gymnothorax chilospilus Bleeker 1864
[Jpn name: Minamiutsubo]
KAUM–I. 121850, 279.6 mm TL, China Port, Serikaku, China; KUN-P 49116, 180.8 mm TL, Yakomo
Beach, Yakomo, China.
Gymnothorax chlorostigma (Kaup 1856)[Jpn name: Hanabirautsubo]
KAUM–I. 122645, 531.0 mm TL, Yaja, China.
Remarks: Gymnothorax meleagris (Shaw 1795) may be a senior synonym of G. chlorostigma (Kaup 1856).
Gymnothorax fimbriatus (Bennett 1832)[Jpn name: Herigoishiutsubo]
Hibino (2018): KAUM–I. 90720, 141.1 TL mm SL, Inobe Port, Inobe, Wadomari. Hibino (2019): KAUM–
I. 90720, 141.1 TL mm SL, Inobe Port, Inobe, Wadomari. This study: KAUM–I. 90720, 141.1 TL mm SL,
Inobe Port, Inobe, Wadomari; KAUM–I. 123515, 291.7 mm TL, Kasaishi Beach, Kibiru, Wadomari.
Gymnothorax flavimarginatus (Rüppell 1830)[Jpn name: Gomautsubo]
Yoshigou et al. (2005a): HMNH 7912, Okidomari, China. Hibino (2018): KAUM–I. 90550, 125.4 mm TL,
Ujiji Beach, Ashikiyora, China; KAUM–I. 90736, 375.0 mm TL, Okidomari Port, Shimojiro, China. Hibino(2019): KAUM–I. 90550, 125.4 mm TL, Ujiji Beach, Ashikiyora, China; KAUM–I. 90736, 375.0 mm TL,
Okidomari Port, Shimojiro, China. This study: KAUM–I. 90550, 125.4 mm TL, Ujiji Beach, Ashikiyora,
China; KAUM–I. 90736, 375.0 mm TL, Okidomari Port, Shimojiro, China.
Gymnothorax javanicus (Bleeker 1859)[Jpn name: Dokuutsubo]
Yoshigou et al. (2005a): HMNH 7847, Naikina, Nagamine, Wadomari. This study: KAUM–I. 90735, 689.5
mm TL, Okidomari Port, Shimojiro, China.
Gymnothorax melatremus Schultz 1953
[Jpn name: Himeutsubo]
Underwater photograph (Pl. 1-H).

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H. Motomura and K. Uehara

Gymnothorax meleagris (Shaw 1795)[Jpn name: Wakautsubo]
Yoshigou et al. (2005a): HMNH 7883, Okidomari, China. Hibino (2018): KAUM–I. 90134, 227.7 mm
TL, Okidomari Port, Shimojiro, China. Hibino (2019): KAUM–I. 90134, 227.7 mm TL, Okidomari Port,
Shimojiro, China. This study: KAUM–I. 90134, 227.7 mm TL, Okidomari Port, Shimojiro, China; KAUM–
I. 123514, 310.4 mm TL, Kasaishi Beach, Kibiru, Wadomari.
Gymnothorax pictus (Ahl 1789)[Jpn name: Aseutsubo]
Yoshigou et al. (2005a): HMNH 7818, Shinjo, China. This study: KAUM–I. 90737, 184.2 mm TL, Hanzaki,
Wadomari; KAUM–I. 90749, 749.5 mm TL, Naikina Port, Nagamine, Wadomari.
Gymnothorax rueppellii (McClelland 1844)[Jpn name: Kurakakeutsubo]
Yoshigou et al. (2005a): HMNH 7849, 2 specimens, Naikina, Nagamine, Wadomari; HMNH 7911,
Okidomari, China.
Gymnothorax thyrsoideus (Richardson 1845)[Jpn name: Sabiutsubo]
Yoshigou et al. (2005a): HMNH 7848, Naikina, Nagamine, Wadomari; HMNH 7913, 2 specimens,
Okidomari, China. Hibino (2018): KAUM–I. 90714, 226.6 mm TL, Ujiji Beach, Ashikiyora, China. Hibino(2019): KAUM–I. 90714, 226.6 mm TL, Ujiji Beach, Ashikiyora, China. This study: KAUM–I. 90195, 211.6mm TL, Yakomo Beach, Yakomo, China; KAUM–I. 90714, 226.6 mm TL, Ujiji Beach, Ashikiyora, China;
KAUM–I. 121905, 200.5 mm TL, Okidomari Port, Shimojiro, China; KAUM–I. 122084, 117.6 mm TL,
Yakomo Beach, Yakomo, China; KUN-P 49025, 256.7 mm TL, China Port, Serikaku, China.
Rhinomuraena quaesita Garman 1888
[Jpn name: Hanahigeutsubo]
Underwater photograph (Pl. 1-I).
Uropterygius micropterus (Bleeker 1852)[Jpn name: Amikikaiutsubo]
KAUM–I. 36356, 94.8 mm TL, off Okinoerabu Island.
FAMILY CONGRIDAE
Conger cinereus Rüppell 1871
[Jpn name: Kirianago]
Yoshigou et al. (2005a): HMNH 7821, Shinjo, China; HMNH 7851, 2 specimens, Naikina, Nagamine,
Wadomari; HMNH 7914, 4 specimens, Okidomari, China; HMNH 7992, Nishibaru, Wadomari.
Conger jordani Kanazawa 1958
[Jpn name: Kuroanago]
KAUM–I. 36352, 199.1 mm TL, off Okinoerabu Island; KAUM–I. 36353, 114.3 mm TL, off Okinoerabu
Island; KAUM–I. 36354, 94.8 mm TL, off Okinoerabu Island.
FAMILY CYPRINIDAE
Carassius sp.
[Jpn name: Gimbuna]Marine and freshwater fishes of Okinoerabu Island

7

Ogawa (1937): Okinoerabu Island. Shibata (1964): OMNH-P 1148, 79.5 mm TL, China; OMNH-P 1149,
76.5 mm TL, China; OMNH-P 1150, 79.5 mm TL, China; OMNH-P 1151, 77.0 mm TL, China; OMNH-P
1152, 76.0 mm TL, China; OMNH-P 1153, 86.5 mm TL, China; OMNH-P 1154, 75.0 mm TL, China;
OMNH-P 1155, 74.5 mm TL, China; OMNH-P 1156, 71.5 mm TL, China; OMNH-P 1157, 68.0 mm TL,
China; OMNH-P 1158, 49.5 mm TL, China; OMNH-P 1159, 45.0 mm TL, China; uncollected specimens,
Amata River, China.
Remarks: Ogawa (1937) and Shibata (1964) confirmed the existence of this species in Okinoerabu Island,
but Kano et al. (2016) considered that this species is now extinct from the island.
Cyprinus rubrofuscus Lacepède 1803
[Jpn name: Koi]
Yoshigou et al. (2005a, as Cyprinus carpio): HMNH 7806, Ginsuido cave, Tamina, China. Yoshigou et al.
(2005b, as Cyprinus carpio): Ginsuido cave, Tamina, China; Kurago, Kibiru, Wadomari. Yoshigou (2014):
Okinoerabu Island.
FAMILY COBITIDAE
Misgurnus cf. anguillicaudatus (Cantor 1842)[Jpn name: Dojo]
Ogawa (1937, as Misgurnus anguillicaudatus): Okinoerabu Island. Shibata (1964, as Misgurnus
anguillicaudatus): OMNH-P 1145, 1146, 2 specimens, channel at rice paddy in Kamihirakawa, China
and muddy paddy around middle reaches of Amata River, China. Yoshigou (2014, as Misgurnus
anguillicaudatus): Okinoerabu Island. Kano et al. (2016): Okinoerabu Island.
Remarks: Kano et al. (2016) suggested that this species is probably an undescribed species.
FAMILY PLOTOSIDAE
Plotosus japonicus Yoshino & Kishimoto 2008
[Jpn name: Gonzui]
KAUM–I. 121931, 57.8 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 121932, 67.6 mm SL, Inobe Port,
Inobe, Wadomari; KAUM–I. 121933, 54.9 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 121934, 69.9mm SL, Inobe Port, Inobe, Wadomari.
Plotosus lineatus (Thunberg 1787)[Jpn name: Minamigonzui]
KAUM–I. 90400, 18.2 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 90459, 52.4 mm SL, Ujiji Beach,
Ashikiyora, China; KAUM–I. 90460, 54.5 mm SL, Ujiji Beach, Ashikiyora, China; KAUM–I. 90461, 48.3mm SL, Ujiji Beach, Ashikiyora, China; KAUM–I. 90462, 63.2 mm SL, Ujiji Beach, Ashikiyora, China;
KAUM–I. 90463, 55.6 mm SL, Ujiji Beach, Ashikiyora, China; KAUM–I. 90464, 50.0 mm SL, Ujiji Beach,
Ashikiyora, China; KAUM–I. 90465, 51.6 mm SL, Ujiji Beach, Ashikiyora, China; KAUM–I. 90466, 52.9mm SL, Ujiji Beach, Ashikiyora, China; KAUM–I. 90467, 49.7 mm SL, Ujiji Beach, Ashikiyora, China;
KAUM–I. 90468, 53.6 mm SL, Ujiji Beach, Ashikiyora, China; KAUM–I. 90469, 49.1 mm SL, Ujiji Beach,
Ashikiyora, China; KAUM–I. 90470, 45.3 mm SL, Ujiji Beach, Ashikiyora, China; KAUM–I. 90471, 50.6mm SL, Ujiji Beach, Ashikiyora, China; KAUM–I. 90472, 51.4 mm SL, Ujiji Beach, Ashikiyora, China;
KAUM–I. 90473, 52.5 mm SL, Ujiji Beach, Ashikiyora, China; KAUM–I. 90474, 48.3 mm SL, Ujiji Beach,
Ashikiyora, China; KAUM–I. 90475, 46.1 mm SL, Ujiji Beach, Ashikiyora, China; KAUM–I. 90476, 49.7mm SL, Ujiji Beach, Ashikiyora, China; KAUM–I. 90477, 47.6 mm SL, Ujiji Beach, Ashikiyora, China;
KAUM–I. 90478, 43.3 mm SL, Ujiji Beach, Ashikiyora, China; KAUM–I. 90479, 47.9 mm SL, Ujiji Beach,
Ashikiyora, China; KAUM–I. 90480, 49.1 mm SL, Ujiji Beach, Ashikiyora, China; KAUM–I. 90481, 47.1mm SL, Ujiji Beach, Ashikiyora, China; KAUM–I. 90482, 49.6 mm SL, Ujiji Beach, Ashikiyora, China;

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H. Motomura and K. Uehara

KAUM–I. 90483, 38.8 mm SL, Ujiji Beach, Ashikiyora, China; KAUM–I. 90484, 51.0 mm SL, Ujiji Beach,
Ashikiyora, China; KAUM–I. 90663, 30.2 mm SL, Wanjo Beach, Azefu, Wadomari; KAUM–I. 90664, 263
specimens, 14.9–26.9 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 121967, 66.5 mm SL, Inobe Port,
Inobe, Wadomari.
FAMILY SYNODONTIDAE
Saurida gracilis (Quoy & Gaimard 1824)[Jpn name: Madaraeso]
KAUM–I. 1971, 142.8 mm SL, off Okinoerabu Island; KAUM–I. 20752, 90.7 mm SL, off Okinoerabu
Island; KAUM–I. 20753, 93.3 mm SL, off Okinoerabu Island; KAUM–I. 20754, 86.4 mm SL, off
Okinoerabu Island; KAUM–I. 20755, 84.0 mm SL, off Okinoerabu Island; KAUM–I. 20756, 80.8 mm SL,
off Okinoerabu Island; KAUM–I. 20757, 70.9 mm SL, off Okinoerabu Island; KAUM–I. 20758, 56.5 mm
SL, off Okinoerabu Island; KAUM–I. 20759, 60.4 mm SL, off Okinoerabu Island; KAUM–I. 20760, 66.4mm SL, off Okinoerabu Island; KAUM–I. 20761, 69.7 mm SL, off Okinoerabu Island; KAUM–I. 20762,
65.2 mm SL, off Okinoerabu Island; KAUM–I. 20763, 52.0 mm SL, off Okinoerabu Island; KAUM–I.
20764, 46.5 mm SL, off Okinoerabu Island; KAUM–I. 20765, 43.4 mm SL, off Okinoerabu Island; KAUM–
I. 90123, 76.4 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 90590, 39.7 mm SL, Okidomari Port,
Shimojiro, China; KAUM–I. 90690, 75.7 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 121770, 100.5mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 121891, 164.6 mm SL, Okidomari Port, Shimojiro, China;
KAUM–I. 122234, 125.1 mm SL, Wadomari Port, Wadomari.
Synodus jaculum Russell & Cressey 1979
[Jpn name: Oguroeso]
KAUM–I. 122316, 72.6 mm SL, off Okidomari Port, Shimojiro, China; KAUM–I. 122317, 74.5 mm SL, off
Okidomari Port, Shimojiro, China.
Synodus variegatus (Lacepède 1803)[Jpn name: Hitosujieso]
KAUM–I. 90755, 37.0 mm SL, Naikina Port, Nagamine, Wadomari.
FAMILY DINEMATICHTHYIDAEAlionematichthys piger (Alcock 1890)[Jpn name: Hadakaryukyuitachiuo]
Nakabo & Kai (2013): Okinoerabu Island. This study: KAUM–I. 2042, 75.8 mm SL, off Okinoerabu Island;
KAUM–I. 10652, 76.9 mm SL, off Okinoerabu Island; KAUM–I. 10655, 55.8 mm SL, off Okinoerabu
Island; KAUM–I. 10658, 52.8 mm SL, off Okinoerabu Island; KAUM–I. 10663, 34.5 mm SL, off
Okinoerabu Island.
Alionematichthys riukiuensis (Aoyagi 1954)[Jpn name: Ryukyuitachiuo]
Nakabo & Kai (2013): Okinoerabu Island. This study: KAUM–I. 10661, 48.1 mm SL, off Okinoerabu Island.
Diancistrus erythraeus (Fowler 1946)[Jpn name: Togeitachiuo]
KAUM–I. 10656, 55.0 mm SL, off Okinoerabu Island; KAUM–I. 10657, 47.3 mm SL, off Okinoerabu
Island; KAUM–I. 10664, 51.8 mm SL, off Okinoerabu Island.

Marine and freshwater fishes of Okinoerabu Island

9

Diancistrus fuscus (Fowler 1946)[Jpn name: Sangoitachiuo]
KAUM–I. 10653, 65.4 mm SL, off Okinoerabu Island; KAUM–I. 10654, 60.5 mm SL, off Okinoerabu
Island; KAUM–I. 10659, 43.4 mm SL, off Okinoerabu Island; KAUM–I. 10662, 35.2 mm SL, off
Okinoerabu Island; KAUM–I. 10665, 26.3 mm SL, off Okinoerabu Island.
Dinematichthys iluocoeteoides Bleeker 1855
[Jpn name: Akaneitachiuo]
Nakabo & Kai (2013): Okinoerabu Island. This study: KAUM–I. 10660, 40.3 mm SL, off Okinoerabu Island;
KAUM–I. 123563, 51.5 mm SL, Yaja, China.
FAMILY ANTENNARIIDAE
Antennarius pictus (Shaw 1794)[Jpn name: Irokaeruanko]
KAUM–I. 2201, 42.5 mm SL, off Okinoerabu Island.
Antennarius randalli Allen 1970
[Jpn name: Himehiratakaeruanko]
Underwater photograph (Pl. 1-J).
Antennatus coccineus (Lesson 1831)[Jpn name: Urumakaeruanko]
Underwater photograph (Pl. 2-A).
Antennatus dorehensis (Bleeker 1859)[Jpn name: Kasurikaeruanko]
Yoshigou et al. (2005a, as Antennarius dorehensis): HNMH 7949, Nishibaru, Wadomari; HMNH 7993,
Nishibaru, Wadomari.
Antennatus nummifer (Cuvier 1817)[Jpn name: Benikaeruanko]
Senou (2019a): KAUM–I. 122613, 7.3 mm SL, off Yakomo, China. This study: KAUM–I. 122613, 7.3 mm
SL, off Yakomo, China.
Antennatus rosaceus (Smith & Radcliffe 1912)[Jpn name: Enagakaeruanko]
Underwater photograph (Pl. 2-B).
Antennatus tuberosus (Cuvier 1817)[Jpn name: Kaeruankomodoki]
Yoshigou et al. (2005a): HMNH 7854, Naikina, Nagamine, Wadomari. This study: KAUM–I. 122311, 8.2
mm SL, west of Taminazaki, Tamina, China.
FAMILY HOLOCENTRIDAE
Myripristis berndti Jordan & Evermann 1903
[Jpn name: Akamatsukasa]
KAUM–I. 90595, 88.5 mm SL, Okidomari Port, Shimojiro, China; KAUM–I. 101625, 161.9 mm SL, Fucha,
Kunigami, Wadomari; KAUM–I. 121764, 170.1 mm SL, Bishi Beach, Kunigami, Wadomari; KAUM–

10

H. Motomura and K. Uehara

I. 121981, 76.7 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 122188, 115.7 mm SL, Wadomari Port,
Wadomari; KAUM–I. 122189, 120.0 mm SL, Wadomari Port, Wadomari; KAUM–I. 122190, 111.9 mm
SL, Wadomari Port, Wadomari; KAUM–I. 122191, 112.0 mm SL, Wadomari Port, Wadomari; KAUM–I.
122192, 114.0 mm SL, Wadomari Port, Wadomari; KUN-P 48866, 74.2 mm SL, Okidomari Port, Shimojiro,
China.
Myripristis greenfieldi Randall & Yamakawa 1996
[Jpn name: Tsumarimatsukasa]
Nakae (2018): KAUM–I. 101633, 80.4 mm SL, Kasaishi Beach, Kibiru, Wadomari. Nakae (2019): KAUM–
I. 101633, 80.4 mm SL, Kasaishi Beach, Kibiru, Wadomari. This study: KAUM–I. 101633, 80.4 mm SL,
Kasaishi Beach, Kibiru, Wadomari.
Myripristis hexagona (Lacepède 1802)[Jpn name: Marumatsukasa]
Eguchi & Motomura (2016): KAUM–I. 36349, 55.5 mm SL, off Okinoerabu Island. This study: KAUM–I.
36349, 55.5 mm SL, off Okinoerabu Island.
Myripristis kuntee Valenciennes 1831
[Jpn name: Kurobimatsukasa]
KAUM–I. 90628, 137.4 mm SL, Naikina Port, Nagamine, Wadomari; KAUM–I. 121973, 140.5 mm SL,
Inobe Port, Inobe, Wadomari.
Myripristis murdjan (Forsskål 1775)[Jpn name: Yogorematsukasa]
Yoshigou et al. (2005a): HMNH 7885, Okidomari, China. Eguchi & Motomura (2016): KAUM–I. 1906, 99.0mm SL, off Okinoerabu Island. This study: KAUM–I. 1906, 99.0 mm SL, off Okinoerabu Island; KAUM–
I. 90659, 88.8 mm SL, Wanjo Beach, Azefu, Wadomari; KAUM–I. 121979, 96.9 mm SL, Inobe Port, Inobe,
Wadomari; KAUM–I. 121980, 66.2 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 122195, 111.6 mm SL,
Wadomari Port, Wadomari; KUN-P 49086, 113.5 mm SL, Wadomari Port, Wadomari.
Myripristis pralinia Cuvier 1829
[Jpn name: Koganematsukasa]
KAUM–I. 122689, 42.5 mm SL, Wadomari Port, Wadomari.
Neoniphon opercularis (Valenciennes 1831)[Jpn name: Hireguroittodai]
Nakae (2018): KAUM–I. 90034, 239.9 mm SL, Okidomari Port, Shimojiro, China. Nakae (2019): KAUM–
I. 90034, 239.9 mm SL, Okidomari Port, Shimojiro, China. This study: KAUM–I. 1979, 114.6 mm SL, off
Okinoerabu Island; KAUM–I. 90034, 239.9 mm SL, Okidomari Port, Shimojiro, China; KAUM–I. 90443,
171.6 mm SL, Inobe Port, Inobe, Wadomari.
Neoniphon sammara (Forsskål 1775)[Jpn name: Ukeguchiittodai]
Eguchi & Motomura (2016): no specimens retained. This study: KAUM–I. 2090, 41.8 mm SL, off
Okinoerabu Island; KAUM–I. 6106, 73.8 mm SL, off Okinoerabu Island; KAUM–I. 6107, 30.6 mm SL, off
Okinoerabu Island; KAUM–I. 90652, 93.3 mm SL, Wanjo Beach, Azefu, Wadomari; KAUM–I. 90653, 98.5mm SL, Wanjo Beach, Azefu, Wadomari; KAUM–I. 90658, 120.1 mm SL, Wanjo Beach, Azefu, Wadomari;
KAUM–I. 90694, 34.0 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 101634, 89.1 mm SL, Kasaishi

Marine and freshwater fishes of Okinoerabu Island

11

Beach, Kibiru, Wadomari; KAUM–I. 121983, 143.2 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I.
122204, 136.6 mm SL, Wadomari Port, Wadomari; KUN-P 49088, 41.9 mm SL, Kasaishi Beach, Kibiru,
Wadomari.
Sargocentron caudimaculatum (Rüppell 1838)[Jpn name: Kurakakebisu]
KAUM–I. 122058, 172.7 mm SL, off Wadomari; KAUM–I. 122059, 173.5 mm SL, off Wadomari.
Sargocentron diadema (Lacepède 1802)[Jpn name: Nijiebisu]
KAUM–I. 90457, 85.5 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 90650, 74.6 mm SL, Wanjo Beach,
Azefu, Wadomari; KAUM–I. 90651, 75.5 mm SL, Wanjo Beach, Azefu, Wadomari; KAUM–I. 121950, 94.0mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 123540, 115.1 mm SL, Yaja, China.
Sargocentron ittodai (Jordan & Fowler 1902)[Jpn name: Teriebisu]
Yoshigou et al. (2005a): HMNH 7844, Shinjo, China. Eguchi & Motomura (2016): KAUM–I. 36346, 68.1
mm SL, off Okinoerabu Island; KAUM–I. 36347, 72.2 mm SL, off Okinoerabu Island. Nakae (2018):
KAUM–I. 90128, 55.5 mm SL, Okidomari Port, Shimojiro, China. Nakae (2019): KAUM–I. 90128, 55.5mm SL, Okidomari Port, Shimojiro, China. This study: KAUM–I. 6105, 101.1 mm SL, off Okinoerabu
Island; KAUM–I. 36346, 68.1 mm SL, off Okinoerabu Island; KAUM–I. 36347, 72.2 mm SL, off
Okinoerabu Island; KAUM–I. 90128, 55.5 mm SL, Okidomari Port, Shimojiro, China; KAUM–I. 90367,
109.5 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 90450, 42.4 mm SL, Inobe Port, Inobe, Wadomari;
KAUM–I. 90458, 91.0 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 90591, 97.3 mm SL, OkidomariPort, Shimojiro, China; KAUM–I. 90592, 102.0 mm SL, Okidomari Port, Shimojiro, China; KAUM–I.
90593, 99.0 mm SL, Okidomari Port, Shimojiro, China; KAUM–I. 90594, 129.1 mm SL, Okidomari Port,
Shimojiro, China; KAUM–I. 90656, 83.4 mm SL, Wanjo Beach, Azefu, Wadomari; KAUM–I. 90657, 90.0
mm SL, Wanjo Beach, Azefu, Wadomari; KAUM–I. 101621, 118.8 mm SL, Fucha, Kunigami, Wadomari;
KAUM–I. 101627, 126.5 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 121948, 105.5 mm SL, Inobe
Port, Inobe, Wadomari; KAUM–I. 121949, 109.8 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 121951,
62.5 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 121982, 127.0 mm SL, Inobe Port, Inobe, Wadomari;
KAUM–I. 122121, 119.7 mm SL, Masana, China; KAUM–I. 122183, 103.6 mm SL, Wadomari Port,
Wadomari; KAUM–I. 122184, 79.6 mm SL, Wadomari Port, Wadomari; KUN-P 48870, 138.6 mm SL,
Okidomari Port, Shimojiro, China; KUN-P 48992, 64.5 mm SL, Kasaishi Beach, Kibiru, Wadomari.
Sargocentron praslin (Lacepède 1802)[Jpn name: Kurobiebisu]
KAUM–I. 122224, 159.4 mm SL, Inobe Port, Inobe, Wadomari.
Sargocentron punctatissimum (Cuvier 1829)[Jpn name: Hoshiebisu]
Eguchi & Motomura (2016): KAUM–I. 36342, 76.1 mm SL, off Okinoerabu Island; KAUM–I. 36343, 67.9mm SL, off Okinoerabu Island; KAUM–I. 36344, 66.0 mm SL, off Okinoerabu Island; KAUM–I. 36345,
67.1 mm SL, off Okinoerabu Island. Nakae (2018): KAUM–I. 90495, 88.9 mm SL, Ujiji Beach, Ashikiyora,
China. Nakae (2019): KAUM–I. 90495, 88.9 mm SL, Ujiji Beach, Ashikiyora, China. This study: KAUM–I.
36342, 76.1 mm SL, off Okinoerabu Island; KAUM–I. 36343, 67.9 mm SL, off Okinoerabu Island; KAUM–
I. 36344, 66.0 mm SL, off Okinoerabu Island; KAUM–I. 36345, 67.1 mm SL, off Okinoerabu Island;
KAUM–I. 90494, 89.7 mm SL, Ujiji Beach, Ashikiyora, China; KAUM–I. 90495, 88.9 mm SL, Ujiji Beach,

12

H. Motomura and K. Uehara

Ashikiyora, China; KAUM–I. 90542, 69.3 mm SL, Ujiji Beach, Ashikiyora, China; KAUM–I. 90596, 114.1mm SL, Okidomari Port, Shimojiro, China; KAUM–I. 90597, 103.8 mm SL, Okidomari Port, Shimojiro,
China; KAUM–I. 101635, 69.6 mm SL, Kasaishi Beach, Kibiru, Wadomari; KAUM–I. 121884, 42.8 mm SL,
Okidomari Port, Shimojiro, China; KAUM–I. 121984, 88.0 mm SL, Inobe Port, Inobe, Wadomari; KAUM–
I. 121985, 90.8 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 121986, 64.5 mm SL, Inobe Port, Inobe,
Wadomari; KAUM–I. 122185, 104.0 mm SL, Wadomari Port, Wadomari; KAUM–I. 122186, 97.6 mm SL,
Wadomari Port, Wadomari; KAUM–I. 122187, 87.7 mm SL, Wadomari Port, Wadomari; KAUM–I. 122197,
113.7 mm SL, Wadomari Port, Wadomari; KAUM–I. 122199, 66.3 mm SL, Wadomari Port, Wadomari;
KAUM–I. 122205, 94.8 mm SL, Wadomari Port, Wadomari; KAUM–I. 122206, 87.7 mm SL, WadomariPort, Wadomari; KAUM–I. 122233, 107.4 mm SL, Inobe Port, Inobe, Wadomari; KUN-P 48865, 103.8 mm
SL, Okidomari Port, Shimojiro, China.
Sargocentron rubrum (Forsskål 1775)[Jpn name: Ayameebisu]
Eguchi & Motomura (2016): KAUM–I. 36348, 66.9 mm SL, off Okinoerabu Island. This study: KAUM–I.
36348, 66.9 mm SL, off Okinoerabu Island.
Sargocentron spiniferum (Forsskål 1775)[Jpn name: Togariebisu]
KAUM–I. 90627, 278.5 mm SL, Naikina Port, Nagamine, Wadomari.
Sargocentron tiere (Cuvier 1829)[Jpn name: Aosujiebisu]
Nakae (2019): KAUM–I. 122201, 144.2 mm SL, Wadomari Port, Wadomari. This study: KAUM–I.
121762, 210.2 mm SL, Bishi Beach, Kunigami, Wadomari; KAUM–I. 121763, 230.9 mm SL, Bishi Beach,
Kunigami, Wadomari; KAUM–I. 122201, 144.2 mm SL, Wadomari Port, Wadomari.
FAMILY MONOCENTRIDAE
Monocentris japonica (Houttuyn 1782)[Jpn name: Matsukasauo]
Underwater photograph (Pl. 2-C).
FAMILY PEGASIDAE
Eurypegasus draconis (Linnaeus 1766)[Jpn name: Umitengu]
Underwater photograph (Pl. 2-D).
FAMILY AULOSTOMIDAE
Aulostomus chinensis (Linnaeus 1766)[Jpn name: Herayagara]
Yoshigou et al. (2005a): HMNH 7886, Okidomari, China. Kawaji (2018): KAUM–I. 90151, 285.0 mm
SL, Okidomari Port, Shimojiro, China. Kawaji (2019a): KAUM–I. 90151, 285.0 mm SL, Okidomari Port,
Shimojiro, China. This study: KAUM–I. 90151, 285.0 mm SL, Okidomari Port, Shimojiro, China.
FAMILY FISTULARIIDAE
Fistularia commersonii Rüppell 1838
[Jpn name: Aoyagara]
Kawaji (2019b): KAUM–I. 123539, 892.0 mm SL, off Okidomari Port, Shimojiro, China. This study:

Marine and freshwater fishes of Okinoerabu Island

13

KAUM–I. 123538, 1210.0 mm SL, off Okidomari Port, Shimojiro, China; KAUM–I. 123539, 892.0 mm SL,
off Okidomari Port, Shimojiro, China.
Fistularia petimba Lacepède 1803
[Jpn name: Akayagara]
Senou (2013a): Okinoerabu Island. This study: KAUM–I. 2079, 198.0 mm SL, off Okinoerabu Island;
KAUM–I. 14191, 165.0 mm SL, off Okinoerabu Island; KAUM–I. 14192, 154.0 mm SL, off Okinoerabu
Island; KAUM–I. 14193, 129.0 mm SL, off Okinoerabu Island; KAUM–I. 14194, 130.0 mm SL, off
Okinoerabu Island.
FAMILY CENTRISCIDAE
Aeoliscus strigatus (Günther 1861)[Jpn name: Hekoayu]
Underwater photograph (Pl. 2-E).
FAMILY SOLENOSTOMIDAE
Solenostomus cyanopterus Bleeker 1854
[Jpn name: Kamisoriuo]
Underwater photograph (Pl. 2-F).
Solenostomus paradoxus (Pallas 1770)[Jpn name: Nishikifuraiuo]
Underwater photograph (Pl. 2-G).
FAMILY SYNGNATHIDAE
Corythoichthys flavofasciatus (Rüppell 1838)[Jpn name: Kishimaishiyoji]
KAUM–I. 121907, 84.6 mm SL, Okidomari Port, Shimojiro, China; KAUM–I. 121908, 78.5 mm SL,
Okidomari Port, Shimojiro, China; KAUM–I. 122104, 92.3 mm SL, Yakomo Beach, Yakomo, China;
KAUM–I. 122120, 85.6 mm SL, Sumiyoshi Port, Sumiyoshi, China; KAUM–I. 122175, 72.3 mm SL, Beach
Rock, Tokudoki, China; KAUM–I. 122176, 63.0 mm SL, Beach Rock, Tokudoki, China; KAUM–I. 122771,
89.0 mm SL, Kasaishi Beach, Kibiru, Wadomari; KAUM–I. 122772, 76.5 mm SL, Kasaishi Beach, Kibiru,
Wadomari; KAUM–I. 123409, 114.8 mm SL, Ujiji Beach, Ashikiyora, China; KAUM–I. 123410, 103.6mm SL, Ujiji Beach, Ashikiyora, China; KAUM–I. 123411, 101.0 mm SL, Ujiji Beach, Ashikiyora, China;
KAUM–I. 123474, 64.0 mm SL, Kasaishi Beach, Kibiru, Wadomari; KAUM–I. 123475, 92.3 mm SL,
Kasaishi Beach, Kibiru, Wadomari.
Remarks: Scientific name is most likely to be Corythoichthys conspicillatus (Jenyns 1842) because C.
flavofasciatus (Rüppell 1838) is an Indian Ocean species.
Corythoichthys haematopterus (Bleeker 1851)[Jpn name: Ishiyoji]
KAUM–I. 90528, 111.5 mm SL, Ujiji Beach, Ashikiyora, China; KAUM–I. 90676, 126.8 mm SL, Wanjo
Beach, Azefu, Wadomari; KAUM–I. 90677, 116.6 mm SL, Wanjo Beach, Azefu, Wadomari; KAUM–
I. 90678, 117.1 mm SL, Wanjo Beach, Azefu, Wadomari; KAUM–I. 90679, 115.4 mm SL, Wanjo Beach,
Azefu, Wadomari; KAUM–I. 90680, 118.7 mm SL, Wanjo Beach, Azefu, Wadomari; KAUM–I. 90721, 20
specimens, 7.0–7.5 mm SL, Wanjo Beach, Azefu, Wadomari; KAUM–I. 101637, 142.9 mm SL, Kasaishi
Beach, Kibiru, Wadomari; KUN-P 48949, 96.3 mm SL, Ujiji Beach, Ashikiyora, China.

14

H. Motomura and K. Uehara

Corythoichthys schultzi Herald 1953
[Jpn name: Kuchinagaishiyoji]
KAUM–I. 121913, 90.5 mm SL, Okidomari Port, Shimojiro, China; KAUM–I. 122738, 86.7 mm SL, Yaja,
China.
Cosmocampus banneri (Herald & Randall 1972)[Jpn name: Hinayoji]
KAUM–I. 122291, 29.4 mm SL, west of Taminazaki, Tamina, China; KAUM–I. 122626, 50.4 mm SL, off
Yakomo, China; KAUM–I. 122732, 26.6 mm SL, Yaja, China.
Doryrhamphus (Doryrhamphus) excisus excisus Kaup 1856
[Jpn name: Hibashiyoji]
Yoshigou et al. (2005a): HMNH 7853, Naikina, Nagamine, Wadomari; HMNH 7884, Okidomari, China;
HMNH 7915, 2 specimens, Okidomari, China; HMNH 7948, Nishibaru, Wadomari. This study: KAUM–
I. 90137, 39.4 mm SL, Okidomari Port, Shimojiro, China; KAUM–I. 90147, 19.0 mm SL, Okidomari Port,
Shimojiro, China; KAUM–I. 90764, 22.5 mm SL, Naikina Port, Nagamine, Wadomari; KAUM–I. 90765,
19.0 mm SL, Naikina Port, Nagamine, Wadomari; KAUM–I. 122142, 41.1 mm SL, Beach Rock, Tokudoki,
China; KAUM–I. 122326, 21.5 mm SL, north of Taminazaki, Tamina, China; KAUM–I. 122764, 31.7 mm
SL, Kasaishi Beach, Kibiru, Wadomari; KAUM–I. 122765, 30.0 mm SL, Kasaishi Beach, Kibiru, Wadomari;
KAUM–I. 123412, 41.3 mm SL, Ujiji Beach, Ashikiyora, China; KUN-P 48959, 53.6 mm SL, Ujiji Beach,
Ashikiyora, China; KUN-P 48960, 36.4 mm SL, Ujiji Beach, Ashikiyora, China; KUN-P 49133, 43.9 mm
SL, Kasaishi Beach, Kibiru, Wadomari.
Doryrhamphus (Doryrhamphus) japonicus Araga & Yoshino 1975
[Jpn name: Nokogiriyoji]
Underwater photograph (Pl. 2-H).
Doryrhamphus (Dunckerocampus) naia Allen & Kuiter 2004
[Jpn name: Kasumioiran-yoji]
KAUM–I. 122027, 146.7 mm SL, Masana, China; KAUM–I. 122028, 84.6 mm SL, Masana, China;
KAUM–I. 122327, 72.0 mm SL, north of Taminazaki, Tamina, China; KAUM–I. 122361, 68.5 mm SL, off
Okidomari Port, Shimojiro, China; KAUM–I. 122707, 119.7 mm SL, off Yakomo, China.
Halicampus brocki (Herald 1953)[Jpn name: Nokogiriumiyakko]
KAUM–I. 122332, 62.3 mm SL, north of Taminazaki, Tamina, China; KAUM–I. 122362, 75.8 mm SL, off
Okidomari Port, Shimojiro, China.
Halicampus macrorhynchus Bamber 1915
[Jpn name: Tatsuumiyakko]
Underwater photograph (Pl. 2-I).
Hippichthys (Hippichthys) spicifer (Rüppell 1838)[Jpn name: Kawayoji]
Yoshigou et al. (2005a): HMNH 7789, Sawa Beach, Wadomari. Yoshigou (2014): Okinoerabu Island.
Hippocampus bargibanti Whitley 1970
[Jpn name: N/A]
Underwater photograph (Pl. 3-A).

Marine and freshwater fishes of Okinoerabu Island

15

Hippocampus denise Lourie & Randall 2003
[New Jpn name: Kakuretatsunoko](Fig. 4)
Material examined. KAUM–I. 132842, female, 23.9 mm SL, off Sumiyoshi Beach, ca. 900 m southof Sumiyoshi Fishing Port, Sumiyoshi, China, Okinoerabu Island, Amami Islands, Kagoshima, Japan,
27°21ʹ10ʺN, 128°31ʹ31ʺE, 38 m depth, 12 Sept. 2019, hand net, K. Uehara.
Remarks. Counts and measurements followed Lourie et al. (1999) and Lourie & Randall (2003). Meristics:
TrR 12; TaR 28; DF 14; PF 11. Morphometrics (as % of SL): HL 8.9; TrL 27.5; TaL 63.6; SnL 5.3; OD 3.3;
PO 7.7; SnD 4.2; HD 9.2; CH 7.1; TD4 5.7; TD9 6.6; DL 7.7. Characters of the present specimen agreed
with those of H. denise given by Lourie & Randall (2003) in having 14 dorsal-fin rays, 11 pectoral-fin rays,
28 tail rings, a rounded nuchal plate without a raised coronet, a snout without a bulbous tip, the inferior andventral trunk ridges reduced to disconnected star-shaped ossifications, the limited number of tubercles on
the body, and the plain orange body color. This species has been known from tropical western Pacific Ocean
and never been reported from Japan on the basis of voucher specimens (Lourie et al. 2016). Thus, the present
specimen represents the first specimen-based record from Japan. In Okinoerabu Island, the species was foundto live on a gorgonian seafan.
Hippocampus pontohi Lourie & Kuiter 2008
[New Jpn name: Yuritatsunoko](Fig. 5)Material examined. KAUM–I. 132427, male, 22.0 mm SL, KAUM–I. 132428, female, 17.0 mm SL, ca. 48
m off Yagoniyazaki, Tamina, China, Okinoerabu Island, Amami Islands, Kagoshima, Japan, 27°24ʹ19ʺN,
128°32ʹ05ʺE, 15 m depth, 28 Aug. 2019, hand net, K. Uehara.
Remarks. Counts and measurements followed Lourie et al. (1999) and Lourie & Randall (2003). Meristics:
TrR 12; TaR 28; DF 13–14; PF 9. Morphometrics (as % of SL): HL 17.5–19.1; TrL 30.2–32.3; TaL 50.2–
50.7; SnL 4.6–5.2; OD 4.6–4.9; PO 8.7–9.2; SnD 4.1–4.2; HD 12.7–13.9; CH 10.6–12.3; TD4 8.4–8.5;
TD9 11.0–11.9; PL 2.3–2.9; DL 7.3–8.3. Characters of the present specimens agreed with those of H.
pontohi given by Short et al. (2018) in having 28 tail rings, two pairs of bilaterally wing-like protrusions
formed by a pair of large truncate spines projecting laterally on both first and second superior trunk ridges,
the laterodorsal surface flat on the third and fourth trunk rings, and a small eighth lateral trunk-ridge spine.
However, the present specimens differed from specimens of H. pontohi described by Lourie & Kuiter (2008)
and Short et al. (2018) in having 13 or 14 dorsal-fin rays (vs. 12 in the latter) and nine pectoral-fin rays (vs.
10). These meristic differences are regarded here as intraspecific variation. This species has been known fromthe tropical western Pacific Ocean and never been reported from Japan on the basis of voucher specimens(Lourie et al. 2016). Thus, the present specimens represent the first specimen-based records from Japan. In
Okinoerabu Island, the species is more common and inhabits shallower depths than H. denise.
Phoxocampus belcheri (Kaup 1856)[Jpn name: Boyoji]
Matsunuma (2019a): KUN-P 49136, 52.9 mm SL, Kasaishi Beach, Kibiru, Wadomari. This study: KUN-P
49136, 52.9 mm SL, Kasaishi Beach, Kibiru, Wadomari; KUN-P 49137, 40.4 mm SL, Kasaishi Beach,
Kibiru, Wadomari; KUN-P 49166, 37.2 mm SL, Kasaishi Beach, Kibiru, Wadomari.
Phoxocampus diacanthus (Schultz 1943)[Jpn name: Hachijoboyoji]
Underwater photograph (Pl. 3-B).

16

H. Motomura and K. Uehara

Syngnathoides biaculeatus (Bloch 1785)[Jpn name: Togeyoji]
KAUM–I. 122301, 173.0 mm SL, Inobe Port, Inobe, Wadomari.
Trachyrhamphus bicoarctatus (Bleeker 1857)[Jpn name: Wakayoji]
Underwater photograph (Pl. 3-C).
FAMILY MUGILIDAE
Chelon macrolepis (Smith 1846)[Jpn name: Kobora]
Yoshigou et al. (2005a): HMNH 7790, Sawa Beach, Wadomari. Yoshigou (2014): Okinoerabu Island. This
study: KAUM–I. 2076, 46.3 mm SL, off Okinoerabu Island; KAUM–I. 18380, 2 specimens, 23.8–30.0 mm
SL, off Okinoerabu Island; KAUM–I. 18408, 27.8 mm SL, off Okinoerabu Island; KAUM–I. 18409, 25.3mm SL, off Okinoerabu Island; KAUM–I. 18411, 25.1 mm SL, off Okinoerabu Island; KAUM–I. 18412,
27.5 mm SL, off Okinoerabu Island; KAUM–I. 18413, 29.2 mm SL, off Okinoerabu Island; KAUM–I.
18414, 24.8 mm SL, off Okinoerabu Island; KAUM–I. 18465, 26.6 mm SL, off Okinoerabu Island; KAUM–
I. 18466, 26.9 mm SL, off Okinoerabu Island; KAUM–I. 18467, 39.0 mm SL, off Okinoerabu Island;
KAUM–I. 18468, 63.9 mm SL, off Okinoerabu Island; KAUM–I. 18469, 53.6 mm SL, off Okinoerabu
Island; KAUM–I. 18470, 66.7 mm SL, off Okinoerabu Island; KAUM–I. 18471, 51.2 mm SL, off
Okinoerabu Island; KAUM–I. 18472, 59.3 mm SL, off Okinoerabu Island; KAUM–I. 18474, 39.6 mm SL,
off Okinoerabu Island; KAUM–I. 18475, 38.8 mm SL, off Okinoerabu Island; KAUM–I. 90235, 27.5 mm
SL, Sumiyoshi Beach, Sumiyoshi, China; KAUM–I. 90236, 24.1 mm SL, Sumiyoshi Beach, Sumiyoshi,
China; KAUM–I. 90237, 26.0 mm SL, Sumiyoshi Beach, Sumiyoshi, China; KAUM–I. 90238, 25.6 mm
SL, Sumiyoshi Beach, Sumiyoshi, China; KAUM–I. 90240, 26.3 mm SL, Sumiyoshi Beach, Sumiyoshi,
China; KAUM–I. 90241, 29.7 mm SL, Sumiyoshi Beach, Sumiyoshi, China; KAUM–I. 90291, 25.0 mm SL,
Sumiyoshi Beach, Sumiyoshi, China; KAUM–I. 90293, 24.6 mm SL, Sumiyoshi Beach, Sumiyoshi, China.
Crenimugil crenilabis (Forsskål 1775)[Jpn name: Furaibora]
KAUM–I. 1974, 51.7 mm SL, off Okinoerabu Island; KAUM–I. 18381, 47.9 mm SL, off Okinoerabu
Island; KAUM–I. 18454, 33.2 mm SL, off Okinoerabu Island; KAUM–I. 18455, 35.1 mm SL, off
Okinoerabu Island; KAUM–I. 18456, 49.3 mm SL, off Okinoerabu Island; KAUM–I. 18457, 33.0 mm SL,
off Okinoerabu Island; KAUM–I. 18458, 36.6 mm SL, off Okinoerabu Island; KAUM–I. 18459, 37.2 mm
SL, off Okinoerabu Island; KAUM–I. 18460, 49.0 mm SL, off Okinoerabu Island; KAUM–I. 18461, 33.4mm SL, off Okinoerabu Island; KAUM–I. 18462, 35.0 mm SL, off Okinoerabu Island; KAUM–I. 18463,
38.8 mm SL, off Okinoerabu Island; KAUM–I. 18464, 32.8 mm SL, off Okinoerabu Island; KAUM–I.
18477, 36.5 mm SL, off Okinoerabu Island; KAUM–I. 18478, 30.6 mm SL, off Okinoerabu Island; KAUM–
I. 18479, 33.7 mm SL, off Okinoerabu Island; KAUM–I. 18480, 32.1 mm SL, off Okinoerabu Island;
KAUM–I. 18481, 31.1 mm SL, off Okinoerabu Island; KAUM–I. 18482, 31.7 mm SL, off Okinoerabu
Island; KAUM–I. 18483, 48.4 mm SL, off Okinoerabu Island; KAUM–I. 18484, 38.2 mm SL, off
Okinoerabu Island; KAUM–I. 18485, 52.7 mm SL, off Okinoerabu Island; KAUM–I. 18486, 48.8 mm SL,
off Okinoerabu Island; KAUM–I. 18487, 48.8 mm SL, off Okinoerabu Island; KAUM–I. 18488, 33.0 mm
SL, off Okinoerabu Island; KAUM–I. 18489, 32.2 mm SL, off Okinoerabu Island; KAUM–I. 18490, 30.5mm SL, off Okinoerabu Island; KAUM–I. 18491, 32.4 mm SL, off Okinoerabu Island; KAUM–I. 18527,
31.1 mm SL, off Okinoerabu Island; KAUM–I. 18533, 33.5 mm SL, off Okinoerabu Island; KAUM–I.
18534, 47.8 mm SL, off Okinoerabu Island; KAUM–I. 18535, 45.9 mm SL, off Okinoerabu Island; KAUM–
I. 18536, 36.9 mm SL, off Okinoerabu Island; KAUM–I. 18537, 31.4 mm SL, off Okinoerabu Island;Marine and freshwater fishes of Okinoerabu Island

17

KAUM–I. 18538, 31.3 mm SL, off Okinoerabu Island; KAUM–I. 18539, 32.2 mm SL, off Okinoerabu
Island; KAUM–I. 18540, 31.3 mm SL, off Okinoerabu Island; KAUM–I. 18541, 32.0 mm SL, off
Okinoerabu Island; KAUM–I. 18542, 28.0 mm SL, off Okinoerabu Island; KAUM–I. 18543, 34.4 mm SL,
off Okinoerabu Island; KAUM–I. 18544, 35.1 mm SL, off Okinoerabu Island; KAUM–I. 90239, 32.4 mm
SL, Sumiyoshi Beach, Sumiyoshi, China; KAUM–I. 90292, 28.3 mm SL, Sumiyoshi Beach, Sumiyoshi,
China; KAUM–I. 90294, 25.8 mm SL, Sumiyoshi Beach, Sumiyoshi, China; KAUM–I. 90295, 25.0 mm
SL, Sumiyoshi Beach, Sumiyoshi, China; KAUM–I. 121876, 415.7 mm SL, Inobe Port, Inobe, Wadomari;
KUN-P 48874, 75.8 mm SL, Okidomari Port, Shimojiro, China; KUN-P 48875, 68.3 mm SL, OkidomariPort, Shimojiro, China; KUN-P 48876, 76.0 mm SL, Okidomari Port, Shimojiro, China; KUN-P 48877, 65.8mm SL, Okidomari Port, Shimojiro, China; KUN-P 48878, 70.8 mm SL, Okidomari Port, Shimojiro, China.
Crenimugil seheli (Forsskål 1775)[Jpn name: Taiwammenada]
KAUM–I. 18473, 48.8 mm SL, off Okinoerabu Island; KAUM–I. 18476, 44.7 mm SL, off Okinoerabu
Island; KAUM–I. 90290, 22.2 mm SL, Sumiyoshi Beach, Sumiyoshi, China; KAUM–I. 121919, 27.5 mm
SL, Inobe Port, Inobe, Wadomari; KAUM–I. 121940, 27.6 mm SL, Inobe Port, Inobe, Wadomari; KAUM–
I. 121941, 27.4 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 121942, 26.0 mm SL, Inobe Port, Inobe,
Wadomari.
Ellochelon vaigiensis (Quoy & Gaimard 1825)[Jpn name: Onibora]
KAUM–I. 121918, 24.3 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 121921, 20.3 mm SL, Inobe
Port, Inobe, Wadomari; KAUM–I. 121922, 23.5 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 121923,
21.2 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 121925, 20.5 mm SL, Inobe Port, Inobe, Wadomari;
KAUM–I. 121926, 20.5 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 121927, 18.2 mm SL, Inobe
Port, Inobe, Wadomari; KAUM–I. 121928, 15.9 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 121929,
14.5 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 121930, 13.3 mm SL, Inobe Port, Inobe, Wadomari;
KAUM–I. 121936, 16.3 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 121938, 14.7 mm SL, Inobe Port,
Inobe, Wadomari.
Moolgarda pedaraki (Valenciennes 1836)[Jpn name: Kamahirebora]
KAUM–I. 90289, 26.0 mm SL, Sumiyoshi Beach, Sumiyoshi, China; KAUM–I. 121920, 24.0 mm SL, Inobe
Port, Inobe, Wadomari.
Osteomugil engeli (Bleeker 1858)[Jpn name: Monnashibora]
KAUM–I. 121924, 21.2 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 121939, 21.0 mm SL, Inobe Port,
Inobe, Wadomari.
Plicomugil labiosus (Valenciennes 1836)[Jpn name: Waniguchibora]
KAUM–I. 121937, 14.7 mm SL, Inobe Port, Inobe, Wadomari.
FAMILY ATHERINIDAE
Atherion elymus Jordan & Starks 1901
[Jpn name: Mugiiwashi]
Yoshigou et al. (2005a): HMNH 7852, 3 specimens, Naikina, Nagamine, Wadomari; HMNH 7916, 3

18

H. Motomura and K. Uehara

specimens, Okidomari, China. This study: KAUM–I. 90559, 48.0 mm SL, Okidomari Port, Shimojiro,
China; KAUM–I. 90560, 48.0 mm SL, Okidomari Port, Shimojiro, China; KUN-P 48871, 36.4 mm SL,
Okidomari Port, Shimojiro, China.
Hypoatherina tsurugae (Jordan & Starks 1901)[Jpn name: Ginisoiwashi]
Kimura (2019a): KAUM–I. 122668, 89.4 mm SL, Naikina Port, Nagamine, Wadomari. This study: KAUM–
I. 121753, 37.9 mm SL, Bishi Beach, Kunigami, Wadomari; KAUM–I. 121754, 36.2 mm SL, Bishi Beach,
Kunigami, Wadomari; KAUM–I. 121755, 35.7 mm SL, Bishi Beach, Kunigami, Wadomari; KAUM–I.
121756, 28.3 mm SL, Bishi Beach, Kunigami, Wadomari; KAUM–I. 121757, 25.4 mm SL, Bishi Beach,
Kunigami, Wadomari; KAUM–I. 121758, 36.6 mm SL, Bishi Beach, Kunigami, Wadomari; KAUM–I.
121759, 31.4 mm SL, Bishi Beach, Kunigami, Wadomari; KAUM–I. 122668, 89.4 mm SL, Naikina Port,
Nagamine, Wadomari.
FAMILY POECILIIDAE
Gambusia affinis affinis (Baird & Girard 1853)[Jpn name: Kadayashi]
Yoshigou et al. (2005a, as Gumbasia affinis): HMNH 7805, Hanjo, Furusato, Wadomari; HMNH 7808,
Amata River, Amata, China. Yoshigou et al. (2005b): Hanjo, Furusato, Wadomari; Amata River, Amata,China. Yoshigou (2014): Okinoerabu Island. Nakajima & Kano (2014): Amata River, Amata, China.
Motomura (2018): 33 specimens (10.9–40.5 mm SL) deposited at KAUM, Okinoerabu Island. This study:
KAUM–I. 6605, 28.0 mm SL, Amata River, Amata, China; KAUM–I. 6606, 21.5 mm SL, Amata River,
Amata, China; KAUM–I. 6614, 28.7 mm SL, Amata River, Amata, China; KAUM–I. 6615, 22.3 mm SL,
Amata River, Amata, China; KAUM–I. 6616, 22.0 mm SL, Amata River, Amata, China; KAUM–I. 6617,
21.0 mm SL, Amata River, Amata, China; KAUM–I. 88917, 25.6 mm SL, Wanjo Beach, Azefu, Wadomari;
KAUM–I. 88918, 20.4 mm SL, Wanjo Beach, Azefu, Wadomari; KAUM–I. 88919, 20.7 mm SL, Wanjo
Beach, Azefu, Wadomari; KAUM–I. 88920, 21.0 mm SL, Wanjo Beach, Azefu, Wadomari; KAUM–I.
88921, 20.0 mm SL, Wanjo Beach, Azefu, Wadomari; KAUM–I. 88922, 17.3 mm SL, Wanjo Beach, Azefu,
Wadomari; KAUM–I. 90018, 20.6 mm SL, Amata River, Amata, China; KAUM–I. 90019, 31.8 mm SL,
Amata River, Amata, China; KAUM–I. 90020, 25.2 mm SL, Amata River, Amata, China; KAUM–I. 90021,
28.2 mm SL, Amata River, Amata, China; KAUM–I. 90196, 22.1 mm SL, Sumiyoshi Beach, Sumiyoshi,
China; KAUM–I. 90197, 24.5 mm SL, Sumiyoshi Beach, Sumiyoshi, China; KAUM–I. 90198, 25.1 mm
SL, Sumiyoshi Beach, Sumiyoshi, China; KAUM–I. 90199, 25.1 mm SL, Sumiyoshi Beach, Sumiyoshi,
China; KAUM–I. 90200, 24.2 mm SL, Sumiyoshi Beach, Sumiyoshi, China; KAUM–I. 90201, 22.7 mm
SL, Sumiyoshi Beach, Sumiyoshi, China; KAUM–I. 90202, 23.6 mm SL, Sumiyoshi Beach, Sumiyoshi,
China; KAUM–I. 90203, 21.4 mm SL, Sumiyoshi Beach, Sumiyoshi, China; KAUM–I. 90204, 20.3 mm
SL, Sumiyoshi Beach, Sumiyoshi, China; KAUM–I. 90205, 20.0 mm SL, Sumiyoshi Beach, Sumiyoshi,
China; KAUM–I. 90206, 18.8 mm SL, Sumiyoshi Beach, Sumiyoshi, China; KAUM–I. 90207, 17.4 mm
SL, Sumiyoshi Beach, Sumiyoshi, China; KAUM–I. 90208, 18.2 mm SL, Sumiyoshi Beach, Sumiyoshi,
China; KAUM–I. 90209, 17.3 mm SL, Sumiyoshi Beach, Sumiyoshi, China; KAUM–I. 90210, 11.1 mm
SL, Sumiyoshi Beach, Sumiyoshi, China; KAUM–I. 90211, 10.9 mm SL, Sumiyoshi Beach, Sumiyoshi,
China; KAUM–I. 90708, 23.5 mm SL, Wanjo Beach, Azefu, Wadomari; KAUM–I. 90709, 26.7 mm SL,
Wanjo Beach, Azefu, Wadomari; KAUM–I. 90710, 11.9 mm SL, Wanjo Beach, Azefu, Wadomari; KAUM–
I. 90738, 33.4 mm SL, Amata River, Amata, China; KAUM–I. 90791, 19.7 mm SL, Wanjo Beach, Azefu,
Wadomari; KAUM–I. 90792, 31.9 mm SL, Wanjo Beach, Azefu, Wadomari; KAUM–I. 90793, 40.5 mm SL,
Wanjo Beach, Azefu, Wadomari.

Marine and freshwater fishes of Okinoerabu Island

19

Poecilia reticulata Peters 1859
[Jpn name: Guppi]
Yoshigou et al. (2005a): HMNH 7936, 2 specimens, Amata River, Amata, China. Senou (2013b): Okinoerabu
Island. Yoshigou (2014): Okinoerabu Island. Nakajima & Kano (2014): Amata River, Amata, China. Uejo
(2018a): KAUM–I. 90707, 21.0 mm SL, Wanjo Beach, Azefu, Wadomari; KAUM–I. 90026, 20.5 mm SL,
Amata River, Amata, China. Uejo (2019a): KAUM–I. 90707, 21.0 mm SL, Wanjo Beach, Azefu, Wadomari;
KAUM–I. 90026, 20.5 mm SL, Amata River, Amata, China. Motomura (2018): 14 specimens (19.1–43.1mm SL) deposited at KAUM, Okinoerabu Island. This study: KAUM–I. 90017, 43.1 mm SL, Amata River,
Amata, China; KAUM–I. 90022, 23.2 mm SL, Amata River, Amata, China; KAUM–I. 90023, 21.8 mm SL,
Amata River, Amata, China; KAUM–I. 90024, 22.2 mm SL, Amata River, Amata, China; KAUM–I. 90025,
19.1 mm SL, Amata River, Amata, China; KAUM–I. 90026, 20.5 mm SL, Amata River, Amata, China;
KAUM–I. 90027, 19.6 mm SL, Amata River, Amata, China; KAUM–I. 90028, 20.5 mm SL, Amata River,
Amata, China; KAUM–I. 90707, 21.0 mm SL, Wanjo Beach, Azefu, Wadomari; KAUM–I. 90794, 18.5mm SL, Wanjo Beach, Azefu, Wadomari; KAUM–I. 90795, 21.9 mm SL, Wanjo Beach, Azefu, Wadomari;
KAUM–I. 90796, 21.2 mm SL, Wanjo Beach, Azefu, Wadomari; KAUM–I. 90797, 21.6 mm SL, Wanjo
Beach, Azefu, Wadomari; KAUM–I. 90798, 23.1 mm SL, Wanjo Beach, Azefu, Wadomari; KAUM–
I. 122743, 19.2 mm SL, Oku River, Wadomari; KAUM–I. 122744, 19.0 mm SL, Oku River, Wadomari;
KAUM–I. 122745, 17.9 mm SL, Oku River, Wadomari; KAUM–I. 122746, 19.8 mm SL, Oku River,
Wadomari; KAUM–I. 122747, 18.5 mm SL, Oku River, Wadomari; KAUM–I. 122748, 23.1 mm SL, Oku
River, Wadomari; KAUM–I. 122749, 26.3 mm SL, Oku River, Wadomari.
Xiphophorus maculatus (Günther 1866)[Jpn name: Sazampurateifuisshu]
Nakajima & Kano (2014): KPM-NI 35588, 34.2 mm SL, Amata River, Amata, China; KPM-NI 35589, 33.1mm SL, Amata River, Amata, China. Uejo (2018a): KAUM–I. 90032, 22.5 mm SL, Amata River, Amata,China; KAUM–I. 90033, 20.1 mm SL, Amata River, Amata, China. Motomura (2018): 5 specimens (19.0–
22.5 mm SL) deposited at KAUM, Okinoerabu Island. Uejo (2019a): KAUM–I. 90032, 22.5 mm SL, Amata
River, Amata, China; KAUM–I. 90033, 20.1 mm SL, Amata River, Amata, China. This study: KAUM–
I. 90029, 19.0 mm SL, Amata River, Amata, China; KAUM–I. 90030, 21.6 mm SL, Amata River, Amata,China; KAUM–I. 90031, 20.3 mm SL, Amata River, Amata, China; KAUM–I. 90032, 22.5 mm SL, Amata
River, Amata, China; KAUM–I. 90033, 20.1 mm SL, Amata River, Amata, China.
FAMILY EXOCOETIDAE
Parexocoetus brachypterus (Richardson 1846)[Jpn name: Tsumaritobiuo]
Tashiro (2018a): KAUM–I. 90001, 126.8 mm SL, north of Okinoerabu Island. Tashiro (2019a): KAUM–
I. 90001, 126.8 mm SL, north of Okinoerabu Island. This study: KAUM–I. 90001, 126.8 mm SL, north of
Okinoerabu Island.
FAMILY BELONIDAE
Platybelone argalus platyura (Bennett 1832)[Jpn name: Himedatsu]
Morishita (2018a): KPM-NI 29270, 317.5 mm SL, off Okinoerabu Island. Morishita (2019a): KPM-NI
29270, 317.5 mm SL, off Okinoerabu Island.
Strongylura incisa (Valenciennes 1846)[Jpn name: Ryukyudatsu]
KAUM–I. 121915, 88.7 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 121943, 248.6 mm SL, Inobe Port,
Inobe, Wadomari.

20

H. Motomura and K. Uehara

FAMILY SCORPAENIDAE
Caracanthus maculatus (Gray 1831)[Jpn name: Dangokoze]
KAUM–I. 122113, 27.6 mm SL, Sumiyoshi Port, Sumiyoshi, China; KAUM–I. 122357, 41.1 mm SL, off
Okidomari Port, Shimojiro, China; KAUM–I. 122358, 32.6 mm SL, off Okidomari Port, Shimojiro, China;
KAUM–I. 122359, 36.4 mm SL, off Okidomari Port, Shimojiro, China; KAUM–I. 122365, 25.2 mm SL, off
Okidomari Port, Shimojiro, China.
Dendrochirus biocellatus (Fowler 1938)[Jpn name: Hireboshiminokasago]
Matsunuma (2019b): KAUM–I. 121999, 39.7 mm SL, Masana, China. This study: KAUM–I. 121999, 39.7mm SL, Masana, China.
Dendrochirus brachypterus (Cuvier 1829)[Jpn name: Shimahimeyamanokami]
Underwater photograph (Pl. 3-D).
Dendrochirus zebra (Cuvier 1829)[Jpn name: Kirimmino]
Yoshigou et al. (2005a): HMNH 7950, Nishibaru, Wadomari. This study: KAUM–I. 90180, 74.3 mm SL,
Yakomo Beach, Yakomo, China; KAUM–I. 90520, 12.8 mm SL, Ujiji Beach, Ashikiyora, China; KAUM–
I. 90538, 31.0 mm SL, Ujiji Beach, Ashikiyora, China; KAUM–I. 101639, 104.7 mm SL, Kasaishi Beach,
Kibiru, Wadomari; KAUM–I. 122072, 41.6 mm SL, Yakomo Beach, Yakomo, China; KAUM–I. 122073,
26.4 mm SL, Yakomo Beach, Yakomo, China; KAUM–I. 122081, 44.8 mm SL, Yakomo Beach, Yakomo,
China; KAUM–I. 122123, 40.4 mm SL, Beach Rock, Tokudoki, China; KAUM–I. 122164, 39.0 mm SL,
Beach Rock, Tokudoki, China; KAUM–I. 122792, 96.8 mm SL, Kasaishi Beach, Kibiru, Wadomari; KUN-P
48873, 97.3 mm SL, Okidomari Port, Shimojiro, China.
Parascorpaena aurita (Rüppell 1838)[Jpn name: Chiburunettaifusakasago]
Furuhashi & Motomura (2019): KAUM–I. 122789, 108.6 mm SL, Kasaishi Beach, Kibiru, Wadomari;
KAUM–I. 123444, 90.7 mm SL, Kasaishi Beach, Kibiru, Wadomari. This study: KAUM–I. 122789, 108.6mm SL, Kasaishi Beach, Kibiru, Wadomari; KAUM–I. 123444, 90.7 mm SL, Kasaishi Beach, Kibiru,
Wadomari; KUN-P 48938, 26.6 mm SL, west of Taminazaki, Tamina, China; KUN-P 49130, 20.5 mm SL,
Kasaishi Beach, Kibiru, Wadomari; KUN-P 49131, 24.1 mm SL, Kasaishi Beach, Kibiru, Wadomari.
Parascorpaena mossambica (Peters 1855)[Jpn name: Nettaifusakasago]
KAUM–I. 122143, 46.1 mm SL, Beach Rock, Tokudoki, China; KAUM–I. 122171, 45.9 mm SL, Beach
Rock, Tokudoki, China; KAUM–I. 122753, 23.9 mm SL, Kasaishi Beach, Kibiru, Wadomari; KAUM–I.
122759, 48.0 mm SL, Kasaishi Beach, Kibiru, Wadomari.
Pterois antennata (Bloch 1787)[Jpn name: Nettaiminokasago]
Matsunuma (2018): KAUM–I. 90425, 25.1 mm SL, Inobe Port, Inobe, Wadomari. Matsunuma (2019b):
KAUM–I. 90425, 25.1 mm SL, Inobe Port, Inobe, Wadomari. This study: KAUM–I. 90425, 25.1 mm SL,
Inobe Port, Inobe, Wadomari.

Marine and freshwater fishes of Okinoerabu Island

21

Pterois radiata Cuvier 1829
[Jpn name: Kimiokoze]
Yoshigou et al. (2005a): HMNH 7950, Nishibaru, Wadomari. This study: KUN-P 48872, 98.9 mm SL,
Okidomari Port, Shimojiro, China.
Pterois volitans (Linnaeus 1758)[Jpn name: Hanaminokasago]
KAUM–I. 122179, 96.9 mm SL, Beach Rock, Tokudoki, China; KUN-P 48907, 80.0 mm SL, Yakomo
Beach, Yakomo, China.
Scorpaenodes guamensis (Quoy & Gaimard 1824)[Jpn name: Guamukasago]
Yoshigou et al. (2005a): HMNH 7835, Shinjo, China; HMNH 7836, Shinjo, China; HMNH 7855, Naikina,
Nagamine, Wadomari; HMNH 7856, Naikina, Nagamine, Wadomari; HMNH 7857, Naikina, Nagamine,
Wadomari; HMNH 7858, Naikina, Nagamine, Wadomari; HMNH 7881, 2 specimens, Naikina, Nagamine,
Wadomari; HMNH 7887, Okidomari, China; HMNH 7917, 2 specimens, Okidomari, China; HMNH 7952,
2 specimens, Nishibaru, Wadomari. This study: KAUM–I. 90497, 49.6 mm SL, Ujiji Beach, Ashikiyora,
China; KAUM–I. 90498, 36.8 mm SL, Ujiji Beach, Ashikiyora, China; KAUM–I. 90504, 34.2 mm SL,
Ujiji Beach, Ashikiyora, China; KAUM–I. 90691, 41.5 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I.
121892, 34.4 mm SL, Okidomari Port, Shimojiro, China; KAUM–I. 121893, 34.5 mm SL, Okidomari Port,
Shimojiro, China; KAUM–I. 122080, 73.0 mm SL, Yakomo Beach, Yakomo, China; KAUM–I. 122126, 65.5mm SL, Beach Rock, Tokudoki, China; KAUM–I. 122151, 21.4 mm SL, Beach Rock, Tokudoki, China;
KAUM–I. 122260, 11.7 mm SL, west of Taminazaki, Tamina, China; KAUM–I. 122781, 42.7 mm SL,
Kasaishi Beach, Kibiru, Wadomari; KAUM–I. 122782, 57.0 mm SL, Kasaishi Beach, Kibiru, Wadomari;
KAUM–I. 123440, 18.0 mm SL, Ujiji Beach, Ashikiyora, China; KAUM–I. 123445, 60.7 mm SL, Kasaishi
Beach, Kibiru, Wadomari; KAUM–I. 123476, 21.2 mm SL, Kasaishi Beach, Kibiru, Wadomari; KAUM–
I. 123477, 16.9 mm SL, Kasaishi Beach, Kibiru, Wadomari; KUN-P 48868, 67.3 mm SL, Okidomari Port,
Shimojiro, China; KUN-P 48954, 36.0 mm SL, Ujiji Beach, Ashikiyora, China; KUN-P 48956, 71.3 mm SL,
Ujiji Beach, Ashikiyora, China; KUN-P 48957, 72.0 mm SL, Ujiji Beach, Ashikiyora, China; KUN-P 49158,
38.6 mm SL, Kasaishi Beach, Kibiru, Wadomari.
Scorpaenodes varipinnis Smith 1957
[Jpn name: Seboshiisokasago]
KAUM–I. 90159, 38.5 mm SL, Okidomari Port, Shimojiro, China; KAUM–I. 122020, 8.9 mm SL, Masana,
China; KAUM–I. 122257, 22.8 mm SL, west of Taminazaki, Tamina, China; KAUM–I. 122288, 31.2 mm
SL, west of Taminazaki, Tamina, China; KAUM–I. 122336, 13.5 mm SL, north of Taminazaki, Tamina,
China; KAUM–I. 122712, 31.4 mm SL, off Yakomo, China; KAUM–I. 122713, 36.4 mm SL, off Yakomo,
China; KAUM–I. 123548, 18.3 mm SL, Yaja, China.
Scorpaenopsis diabolus (Cuvier 1829)[Jpn name: Niraikasago]
Motomura (2019a): KAUM–I. 121889, 149.3 mm SL, Okidomari Port, Shimojiro, China. This study:
KAUM–I. 90713, 82.5 mm SL, Ujiji Beach, Ashikiyora, China; KAUM–I. 121888, 224.7 mm SL,
Okidomari Port, Shimojiro, China; KAUM–I. 121889, 149.3 mm SL, Okidomari Port, Shimojiro, China;
KAUM–I. 122794, 227.3 mm SL, Kasaishi Beach, Kibiru, Wadomari.
Scorpaenopsis macrochir Ogilby 1910
[Jpn name: Marusubekasago]

22

H. Motomura and K. Uehara

KAUM–I. 90496, 56.5 mm SL, Ujiji Beach, Ashikiyora, China; KAUM–I. 90537, 64.4 mm SL, Ujiji Beach,
Ashikiyora, China.
Scorpaenopsis orientalis Randall & Eschmeyer 2002
[Jpn name: Toyourumakasago]
Motomura (2019a): KAUM–I. 122037, 84.8 mm SL, Masana, China. This study: KAUM–I. 122037, 84.8mm SL, Masana, China.
Scorpaenopsis papuensis (Cuvier 1829)[Jpn name: Urumakasago]
KAUM–I. 122003, 26.1 mm SL, Masana, China; KAUM–I. 122289, 19.9 mm SL, west of Taminazaki,
Tamina, China.
Scorpaenopsis possi Randall & Eschmeyer 2002
[Jpn name: Mimitogeonikasago]
KAUM–I. 121914, 123.3 mm SL, Okidomari Port, Shimojiro, China; KAUM–I. 122212, 143.1 mm SL,
Wadomari Port, Wadomari; KAUM–I. 122310, 11.8 mm SL, west of Taminazaki, Tamina, China; KAUM–I.
122379, 162.0 mm SL, off Okidomari Port, Shimojiro, China; KAUM–I. 122630, 120.6 mm SL, Yaja, China.
Scorpaenopsis venosa (Cuvier 1829)[Jpn name: Hyugakasago]
KUN-P 48946, 72.5 mm SL, west of Taminazaki, Tamina, China.
Scorpaenopsis vittapinna Randall & Eschmeyer 2002
[Jpn name: Kogataonikasago]
Motomura (2019a): KAUM–I. 122329, 40.5 mm SL, north of Taminazaki, Tamina, China. This study:
KAUM–I. 122329, 40.5 mm SL, north of Taminazaki, Tamina, China.
Sebastapistes cyanostigma (Bleeker 1856)[Jpn name: Kasurifusakasago]
KAUM–I. 90077, 42.2 mm SL, Okidomari Port, Shimojiro, China; KAUM–I. 90739, 52.0 mm SL, Naikina
Port, Nagamine, Wadomari; KAUM–I. 122352, 51.1 mm SL, off Okidomari Port, Shimojiro, China; KAUM–
I. 122353, 32.0 mm SL, off Okidomari Port, Shimojiro, China; KAUM–I. 123464, 27.9 mm SL, Kasaishi
Beach, Kibiru, Wadomari; KUN-P 49113, 41.2 mm SL, Yakomo Beach, Yakomo, China.
Sebastapistes fowleri (Pietschmann 1934)[Jpn name: Puchifusakasago]
Motomura (2019a): KAUM–I. 122008, 24.3 mm SL, Masana, China. This study: KAUM–I. 122008, 24.3mm SL, Masana, China; KAUM–I. 122258, 19.5 mm SL, west of Taminazaki, Tamina, China; KAUM–
I. 122259, 18.1 mm SL, west of Taminazaki, Tamina, China; KAUM–I. 123546, 16.4 mm SL, Yaja, China;
KUN-P 49001, 20.0 mm SL, Yaja, China; KUN-P 49013, 21.1 mm SL, Yaja, China.
Sebastapistes mauritiana (Cuvier 1829)[Jpn name: Hachijofusakasago]
Motomura (2019a): KAUM–I. 122079, 44.1 mm SL, Yakomo Beach, Yakomo, China. This study: KAUM–I.
122079, 44.1 mm SL, Yakomo Beach, Yakomo, China.

Marine and freshwater fishes of Okinoerabu Island

23

Sebastapistes strongia (Cuvier 1829)[Jpn name: Madarafusakasago]
KAUM–I. 122078, 31.3 mm SL, Yakomo Beach, Yakomo, China; KUN-P 48930, 28.0 mm SL, Yakomo
Beach, Yakomo, China.
Sebastapistes tinkhami (Fowler 1946)[Jpn name: Niramikasago]
Yoshigou et al. (2005a): HMNH 7940, Okidomari, China.
Taenianotus triacanthus Lacepède 1802
[Jpn name: Hadakahaokoze]
Motomura (2019a): KAUM–I. 122324, 46.4 mm SL, north of Taminazaki, Tamina, China; KAUM–I.
123452, 36.3 mm SL, Kasaishi Beach, Kibiru, Wadomari. This study: KAUM–I. 122303, 62.2 mm SL,
west of Taminazaki, Tamina, China; KAUM–I. 122324, 46.4 mm SL, north of Taminazaki, Tamina, China;
KAUM–I. 123452, 36.3 mm SL, Kasaishi Beach, Kibiru, Wadomari.
FAMILY SEBASTIDAE
Sebastiscus albofasciatus (Lacepède 1802)[Jpn name: Ayamekasago]
KAUM–I. 53946, 184.6 mm SL, off Okinoerabu Island; KAUM–I. 53947, 171.8 mm SL, off Okinoerabu
Island.
FAMILY SYNANCEIIDAE
Synanceia verrucosa Bloch & Schneider 1801
[Jpn name: Onidarumaokoze]
Underwater photograph (Pl. 3-E).
FAMILY TETRAROGIDAE
Ablabys taenianotus (Cuvier 1829)[Jpn name: Tsumajirookoze]
Underwater photograph (Pl. 3-F).
FAMILY PLATYCEPHALIDAE
Sunagocia otaitensis (Cuvier 1829)[Jpn name: Fusakuchigochi]
KAUM–I. 122116, 58.1 mm SL, Sumiyoshi Port, Sumiyoshi, China.
Thysanophrys chiltonae Schultz 1966
[Jpn name: Kuroshimagochi]
Motomura et al. (2019): KAUM–I. 90588, 152.5 mm SL, Okidomari Port, Shimojiro, China; KAUM–
I. 122038, 144.3 mm SL, Masana, China; KAUM–I. 122322, 49.1 mm SL, north of Taminazaki, Tamina,
China; KAUM–I. 122323, 43.3 mm SL, north of Taminazaki, Tamina, China. This study: KAUM–I. 90588,
152.5 mm SL, Okidomari Port, Shimojiro, China; KAUM–I. 122038, 144.3 mm SL, Masana, China;
KAUM–I. 122322, 49.1 mm SL, north of Taminazaki, Tamina, China; KAUM–I. 122323, 43.3 mm SL, northof Taminazaki, Tamina, China.

24

H. Motomura and K. Uehara

FAMILY SERRANIDAE
Aporops bilinearis Schultz 1943
[Jpn name: Yamatotogemegisu]
Yoshida & Motomura (2019a): KAUM–I. 122377, 17.5 mm SL, off Okidomari Port, Shimojiro, China.
Sakurai (2019a): KAUM–I. 122377, 17.5 mm SL, off Okidomari Port, Shimojiro, China. This study:
KAUM–I. 122377, 17.5 mm SL, off Okidomari Port, Shimojiro, China.
Caprodon schlegelii (Günther 1859)[Jpn name: Akaisaki]
Sakurai (2018a): KAUM–I. 90726, 277.4 mm SL, Kunigami, Wadomari. Sakurai (2019a): KAUM–I. 90726,
277.4 mm SL, Kunigami, Wadomari. This study: KAUM–I. 90726, 277.4 mm SL, Kunigami, Wadomari.
Cephalopholis argus Schneider 1801
[Jpn name: Aonomehata]
Yoshigou et al. (2005a): HMNH 7889, Okidomari, China; HMNH 7918, Okidomari, China. This study:
KAUM–I. 90126, 256.1 mm SL, Okidomari Port, Shimojiro, China; KAUM–I. 122134, 43.7 mm SL, Beach
Rock, Tokudoki, China.
Cephalopholis sonnerati (Valenciennes 1828)[Jpn name: Azahata]
KAUM–I. 123535, 244.2 mm SL, Hirase, Tamashiro, Wadomari.
Cephalopholis spiloparaea (Valenciennes 1828)[Jpn name: Akahana]
Sakurai (2018a): KAUM–I. 90790, 182.6 mm SL, Naikina Port, Nagamine, Wadomari. Sakurai (2019a):
KAUM–I. 90790, 182.6 mm SL, Naikina Port, Nagamine, Wadomari. This study: KAUM–I. 90790, 182.6mm SL, Naikina Port, Nagamine, Wadomari; KAUM–I. 122318, 103.8 mm SL, off Okidomari Port,
Shimojiro, China; KAUM–I. 122665, 127.5 mm SL, off Naikina Port, Nagamine, Wadomari; KAUM–I.
122666, 101.8 mm SL, off Naikina Port, Nagamine, Wadomari.
Cephalopholis urodeta (Forster 1801)[Jpn name: Nijihata]
Sakurai (2018a): KAUM–I. 90165, 33.3 mm SL, Okidomari Port, Shimojiro, China. Sakurai (2019a):
KAUM–I. 90165, 33.3 mm SL, Okidomari Port, Shimojiro, China. This study: KAUM–I. 90165, 33.3 mm
SL, Okidomari Port, Shimojiro, China; KAUM–I. 90406, 129.2 mm SL, Inobe Port, Inobe, Wadomari;
KAUM–I. 101606, 151.4 mm SL, Kunigami, Wadomari; KAUM–I. 122667, 105.1 mm SL, off Naikina Port,
Nagamine, Wadomari; KAUM–I. 123505, 143.5 mm SL, off Okidomari Port, Shimojiro, China.
Diploprion bifasciatum Cuvier 1828
[Jpn name: Kihassoku]
KAUM–I. 90729, 140.4 mm SL, Naikina Port, Nagamine, Wadomari.
Epinephelus fasciatus (Forsskål 1775)[Jpn name: Akahata]
KAUM–I. 101622, 163.8 mm SL, Fucha, Kunigami, Wadomari; KAUM–I. 121742, 219.8 mm SL, off
Okidomari Port, Shimojiro, China; KAUM–I. 121743, 204.7 mm SL, off Okidomari Port, Shimojiro, China;
KAUM–I. 121976, 122.5 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 122252, 68.7 mm SL, west of
Taminazaki, Tamina, China; KAUM–I. 122671, 100.0 mm SL, off Naikina Port, Nagamine, Wadomari;Marine and freshwater fishes of Okinoerabu Island

25

KAUM–I. 123498, 175.1 mm SL, off Okidomari Port, Shimojiro, China; KAUM–I. 123499, 160.9 mm SL,
off Okidomari Port, Shimojiro, China; KAUM–I. 123500, 183.2 mm SL, off Okidomari Port, Shimojiro,
China; KAUM–I. 123504, 156.5 mm SL, off Okidomari Port, Shimojiro, China; KAUM–I. 123511, 235.4mm SL, off Okidomari Port, Shimojiro, China; KAUM–I. 123516, 202.7 mm SL, off Okidomari Port,
Shimojiro, China; KAUM–I. 123520, 230.1 mm SL, off Okidomari Port, Shimojiro, China; KUN-P 48999,
181.3 mm SL, off Okinoerabu Island.
Epinephelus hexagonatus (Forster 1801)[Jpn name: Ishigakihata]
Yoshigou et al. (2005a): HMNH 7859, Naikina, Nagamine, Wadomari; HMNH 7860, Naikina, Nagamine,
Wadomari; HMNH 7919, Okidomari, China. Sakurai (2018a): KAUM–I. 90166, 36.7 mm SL, OkidomariPort, Shimojiro, China. This study: KAUM–I. 36321, 100.3 mm SL, off Okinoerabu Island; KAUM–I.
36323, 68.1 mm SL, off Okinoerabu Island; KAUM–I. 90166, 36.7 mm SL, Okidomari Port, Shimojiro,
China; KAUM–I. 90581, 140.0 mm SL, Okidomari Port, Shimojiro, China; KAUM–I. 101618, 115.5mm SL, Kunigami, Wadomari; KAUM–I. 101623, 91.3 mm SL, Fucha, Kunigami, Wadomari; KAUM–I.
121769, 105.9 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 122228, 125.4 mm SL, Inobe Port, Inobe,
Wadomari; KAUM–I. 122229, 149.1 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 132842, 47.9 mm
SL, west of Taminazaki, Tamina, China; KAUM–I. 133460, 108.4 mm SL, Beach Rock, Tokudoki, China;
KAUM–I. 133461, 54.3 mm SL, Beach Rock, Tokudoki, China; KAUM–I. 133462, 50.5 mm SL, Beach
Rock, Tokudoki, China; KUN-P 49101, 187.5 mm SL, off Yakomo, China.
Epinephelus howlandi (Günther 1873)[Jpn name: Hiregurohata]
KAUM–I. 90181, 48.5 mm SL, Yakomo Beach, Yakomo, China; KAUM–I. 90248, 25.1 mm SL, Sumiyoshi
Beach, Sumiyoshi, China; KAUM–I. 122064, 297.7 mm SL, Kunigami, Wadomari.
Epinephelus macrospilos (Bleeker 1855)[Jpn name: Kibirehata]
KAUM–I. 36320, 127.7 mm SL, off Okinoerabu Island; KAUM–I. 36322, 84.8 mm SL, off Okinoerabu
Island; KAUM–I. 122065, 211.0 mm SL, Kunigami, Wadomari.
Epinephelus maculatus (Bloch 1790)[Jpn name: Shirobuchihata]
KAUM–I. 123537, 292.8 mm SL, Hirase, Tamashiro, Wadomari.
Epinephelus melanostigma Schultz 1953
[Jpn name: Sumitsukihata]
Sakurai (2019a): KAUM–I. 122635, 41.0 mm SL, China Port, Serikaku, China. This study: KAUM–I.
122635, 41.0 mm SL, China Port, Serikaku, China.
Epinephelus merra Bloch 1793
[Jpn name: Kammonhata]
Yoshigou et al. (2005a): HMNH 7888, Okidomari, China; HMNH 7920, Okidomari, China; HMNH 7953,
Nishibaru, Wadomari; HMNH 7954, Nishibaru, Wadomari; HMNH 7955, Nishibaru, Wadomari. This study:
KAUM–I. 2003, 65.1 mm SL, off Okinoerabu Island; KAUM–I. 7791, 37.6 mm SL, off Okinoerabu Island;
KAUM–I. 7792, 65.1 mm SL, off Okinoerabu Island; KAUM–I. 7793, 37.3 mm SL, off Okinoerabu Island;
KAUM–I. 7794, 37.9 mm SL, off Okinoerabu Island; KAUM–I. 7795, 34.7 mm SL, off Okinoerabu Island;
KAUM–I. 7796, 35.3 mm SL, off Okinoerabu Island; KAUM–I. 7797, 39.2 mm SL, off Okinoerabu Island;

26

H. Motomura and K. Uehara

KAUM–I. 7798, 36.3 mm SL, off Okinoerabu Island; KAUM–I. 7799, 34.0 mm SL, off Okinoerabu Island;
KAUM–I. 7800, 39.2 mm SL, off Okinoerabu Island; KAUM–I. 7801, 35.1 mm SL, off Okinoerabu Island;
KAUM–I. 7802, 39.9 mm SL, off Okinoerabu Island; KAUM–I. 7803, 37.6 mm SL, off Okinoerabu Island;
KAUM–I. 7804, 39.5 mm SL, off Okinoerabu Island; KAUM–I. 7805, 31.1 mm SL, off Okinoerabu Island;
KAUM–I. 7806, 39.8 mm SL, off Okinoerabu Island; KAUM–I. 7807, 43.4 mm SL, off Okinoerabu Island;
KAUM–I. 7808, 39.1 mm SL, off Okinoerabu Island; KAUM–I. 7809, 36.9 mm SL, off Okinoerabu Island;
KAUM–I. 7810, 38.1 mm SL, off Okinoerabu Island; KAUM–I. 7815, 37.3 mm SL, off Okinoerabu Island;
KAUM–I. 90118, 128.4 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 90191, 33.8 mm SL, Yakomo
Beach, Yakomo, China; KAUM–I. 90368, 143.9 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 90369,
133.3 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 90370, 103.8 mm SL, Inobe Port, Inobe, Wadomari;
KAUM–I. 90517, 130.9 mm SL, Ujiji Beach, Ashikiyora, China; KAUM–I. 90518, 150.2 mm SL, Ujiji
Beach, Ashikiyora, China; KAUM–I. 90613, 80.4 mm SL, Naikina Port, Nagamine, Wadomari; KAUM–I.
90660, 35.2 mm SL, Wanjo Beach, Azefu, Wadomari; KAUM–I. 90661, 88.4 mm SL, Wanjo Beach, Azefu,
Wadomari; KAUM–I. 90662, 128.0 mm SL, Wanjo Beach, Azefu, Wadomari; KAUM–I. 90731, 192.0mm SL, Hanzaki, Wadomari; KAUM–I. 90732, 125.0 mm SL, Hanzaki, Wadomari; KAUM–I. 101626,
155.0 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 121767, 129.0 mm SL, Bishi Beach, Kunigami,
Wadomari; KAUM–I. 121864, 133.6 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 121901, 100.0 mm
SL, Okidomari Port, Shimojiro, China; KAUM–I. 121902, 97.0 mm SL, Okidomari Port, Shimojiro, China;
KAUM–I. 121975, 133.0 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 121991, 140.5 mm SL, Inobe
Port, Inobe, Wadomari; KAUM–I. 122196, 130.8 mm SL, Wadomari Port, Wadomari; KAUM–I. 122203,
170.2 mm SL, Wadomari Port, Wadomari; KAUM–I. 122230, 147.0 mm SL, Inobe Port, Inobe, Wadomari;
KAUM–I. 122231, 125.2 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 122232, 130.9 mm SL, Inobe
Port, Inobe, Wadomari; KAUM–I. 122670, 122.6 mm SL, off Naikina Port, Nagamine, Wadomari; KAUM–I.
122760, 113.9 mm SL, Kasaishi Beach, Kibiru, Wadomari; KAUM–I. 122761, 96.3 mm SL, Kasaishi Beach,
Kibiru, Wadomari; KAUM–I. 122762, 48.7 mm SL, Kasaishi Beach, Kibiru, Wadomari; KAUM–I. 123448,
104.9 mm SL, Kasaishi Beach, Kibiru, Wadomari; KAUM–I. 123449, 93.2 mm SL, Kasaishi Beach, Kibiru,
Wadomari; KAUM–I. 123492, 133.8 mm SL, China Port, Serikaku, China; KAUM–I. 123493, 114.0 mm
SL, China Port, Serikaku, China; KUN-P 48867, 103.6 mm SL, Okidomari Port, Shimojiro, China; KUN-P
49097, 154.5 mm SL, off Yakomo, China; KUN-P 49098, 141.1 mm SL, off Yakomo, China; KUN-P 49099,
113.2 mm SL, off Yakomo, China; KUN-P 49100, 175.0 mm SL, off Yakomo, China.
Epinephelus retouti Bleeker 1868
[Jpn name: Akahatamodoki]
KAUM–I. 123533, 355.2 mm SL, off Okinoerabu Island.
Epinephelus tauvina (Forsskål 1775)[Jpn name: Hitomihata]
Yoshigou et al. (2005a): HMNH 7921, Okidomari, China. This study: KAUM–I. 121851, 161.2 mm SL,
Kunigami, Wadomari; KAUM–I. 122225, 152.7 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 123491,
231.1 mm SL, China Port, Serikaku, China.
Grammistes sexlineatus (Thunberg 1792)[Jpn name: Nunosarashi]
Yoshigou et al. (2005a): HMNH 7861, 2 specimens, Naikina, Nagamine, Wadomari. Sakurai (2018a):
KAUM–I. 90160, 58.5 mm SL, Okidomari Port, Shimojiro, China. Sakurai (2019a): KAUM–I. 90160, 58.5mm SL, Okidomari Port, Shimojiro, China. This study: KAUM–I. 90160, 58.5 mm SL, Okidomari Port,
Shimojiro, China; KAUM–I. 90334, 82.0 mm SL, Okidomari Port, Shimojiro, China; KAUM–I. 90335, 82.6mm SL, Okidomari Port, Shimojiro, China; KAUM–I. 90552, 73.2 mm SL, Ujiji Beach, Ashikiyora, China;

Marine and freshwater fishes of Okinoerabu Island

27

KAUM–I. 90553, 87.0 mm SL, Ujiji Beach, Ashikiyora, China; KAUM–I. 121898, 51.4 mm SL, OkidomariPort, Shimojiro, China; KAUM–I. 122125, 30.6 mm SL, Sumiyoshi Port, Sumiyoshi, China; KAUM–I.
122763, 48.7 mm SL, Kasaishi Beach, Kibiru, Wadomari; KAUM–I. 123450, 60.4 mm SL, Kasaishi Beach,
Kibiru, Wadomari; KUN-P 49080, 40.7 mm SL, China Port, Serikaku, China.
Liopropoma susumi (Jordan & Seale 1906)[Jpn name: Kosujihanasuzuki]
KAUM–I. 90397, 59.2 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 122346, 53.6 mm SL, off
Okidomari Port, Shimojiro, China; KAUM–I. 123552, 27.4 mm SL, Yaja, China; KAUM–I. 123578, 26.5mm SL, Yaja, China.
Plectranthias longimanus (Weber 1913)[Jpn name: Muramomijihanadai]
Kawaji et al. (2019): KAUM–I. 122307, 13.3 mm SL, Taminazaki, Tamina, China. This study: KAUM–I.
122307, 13.3 mm SL, Taminazaki, Tamina, China.
Plectranthias nanus Randall 1980
[Jpn name: Chibihanadai]
Kawaji et al. (2019): KAUM–I. 122280, 24.4 mm SL, west of Taminazaki, Tamina, China; KAUM–
I. 122281, 21.9 mm SL, west of Taminazaki, Tamina, China; KAUM–I. 122282, 17.9 mm SL, west of
Taminazaki, Tamina, China; KAUM–I. 122283, 17.9 mm SL, west of Taminazaki, Tamina, China; KAUM–
I. 122284, 10.6 mm SL, west of Taminazaki, Tamina, China; KAUM–I. 122330, 24.4 mm SL, north ofTaminazaki, Tamina, China; KAUM–I. 122342, 23.0 mm SL, off Okidomari Port, Shimojiro, China;
KAUM–I. 122611, 26.7 mm SL, off Yakomo, China; KAUM–I. 122709, 20.1 mm SL, off Yakomo, China;
KAUM–I. 123547, 26.7 mm SL, Yaja, China. This study: KAUM–I. 122046, 26.1 mm SL, Masana,
China; KAUM–I. 122047, 20.0 mm SL, Masana, China; KAUM–I. 122110, 19.6 mm SL, Sumiyoshi Port,
Sumiyoshi, China; KAUM–I. 122280, 24.4 mm SL, west of Taminazaki, Tamina, China; KAUM–I. 122281,
21.9 mm SL, west of Taminazaki, Tamina, China; KAUM–I. 122282, 17.9 mm SL, west of Taminazaki,
Tamina, China; KAUM–I. 122283, 17.9 mm SL, west of Taminazaki, Tamina, China; KAUM–I. 122284,
10.6 mm SL, west of Taminazaki, Tamina, China; KAUM–I. 122330, 24.4 mm SL, north of Taminazaki,
Tamina, China; KAUM–I. 122342, 23.0 mm SL, off Okidomari Port, Shimojiro, China; KAUM–I. 122611,
26.7 mm SL, off Yakomo, China; KAUM–I. 122709, 20.1 mm SL, off Yakomo, China; KAUM–I. 123547,
26.7 mm SL, Yaja, China.
Remarks: In Japanese waters, this species has treated as P. longimanus by many Japanese ichtyologists, but
Kawaji et al. (2019) regarded it as P. nanus.
Plectranthias winniensis (Tyler 1966)[Jpn name: Deigohanadai]
Kawaji et al. (2019): KAUM–I. 121993, 29.5 mm SL, off Masana, China; KAUM–I. 121994, 24.4 mm SL,
off Masana, China; KAUM–I. 122710, 26.1 mm SL, off Yakomo, China; KAUM–I. 122711, 24.1 mm SL,
off Yakomo, China. This study: KAUM–I. 121993, 29.5 mm SL, off Masana, China; KAUM–I. 121994, 24.4mm SL, off Masana, China; KAUM–I. 122710, 26.1 mm SL, off Yakomo, China; KAUM–I. 122711, 24.1mm SL, off Yakomo, China.
Remarks: Kawaji et al. (2019) reported the first records of this species from Japan.
Plectropomus leopardus (Lacepède 1802)[Jpn name: Sujiara]
Sakurai (2019a): KAUM–I. 123534, 356.7 mm SL, Hirase, Tamashiro, Wadomari. This study: KAUM–I.

28

H. Motomura and K. Uehara

122680, 261.8 mm SL, off Wadomari Port, Wadomari; KAUM–I. 123534, 356.7 mm SL, Hirase, Tamashiro,
Wadomari.
Pogonoperca punctata (Valenciennes 1830)[Jpn name: Agohata]
Sakurai (2019a): KAUM–I. 122239, 119.7 mm SL, north of Taminazaki, Tamina, China; KAUM–I. 122298,
127.1 mm SL, west of Taminazaki, Tamina, China. This study: KAUM–I. 122239, 119.7 mm SL, northof Taminazaki, Tamina, China; KAUM–I. 122298, 127.1 mm SL, west of Taminazaki, Tamina, China;
KAUM–I. 122299, 183.3 mm SL, west of Taminazaki, Tamina, China; KUN-P 48935, 142.6 mm SL, west of
Taminazaki, Tamina, China.
Pseudanthias bicolor (Randall 1979)[Jpn name: Futairohanagoi]
Underwater photograph (Pl. 3-G).
Pseudanthias hypselosoma Bleeker 1877
[Jpn name: Keramahanadai]
Underwater photograph (Pl. 3-H).
Pseudanthias lori (Lubbock & Randall 1976)[Jpn name: Akaboshihanagoi]
Underwater photograph (Pl. 4-A).
Pseudanthias pleurotaenia (Bleeker 1857)[Jpn name: Sumirenagahanadai]
Underwater photograph (Pl. 4-B).
Pseudanthias squamipinnis (Peters 1855)[Jpn name: Kingyohanadai]
Sakurai (2018a): KAUM–I. 90135, 12.0 mm SL, Okidomari Port, Shimojiro, China. Sakurai (2019a):
KAUM–I. 90135, 12.0 mm SL, Okidomari Port, Shimojiro, China. This study: KAUM–I. 90135, 12.0 mm
SL, Okidomari Port, Shimojiro, China; KAUM–I. 122005, 36.4 mm SL, Masana, China; KAUM–I. 122006,
37.2 mm SL, Masana, China; KAUM–I. 122247, 23.1 mm SL, west of Taminazaki, Tamina, China; KAUM–
I. 122261, 16.8 mm SL, west of Taminazaki, Tamina, China; KAUM–I. 122262, 26.9 mm SL, west of
Taminazaki, Tamina, China; KAUM–I. 122263, 32.4 mm SL, west of Taminazaki, Tamina, China; KAUM–I.
122264, 37.5 mm SL, west of Taminazaki, Tamina, China; KUN-P 48890, 38.2 mm SL, off Masana, China;
KUN-P 48899, 16.3 mm SL, off Masana, China; KUN-P 48939, 32.6 mm SL, west of Taminazaki, Tamina,
China; KUN-P 48940, 25.2 mm SL, west of Taminazaki, Tamina, China.
Pseudogramma polyacantha (Bleeker 1856)[Jpn name: Togemegisu]
KAUM–I. 90076, 26.7 mm SL, Okidomari Port, Shimojiro, China; KAUM–I. 90399, 49.5 mm SL, Inobe
Port, Inobe, Wadomari; KAUM–I. 122033, 46.1 mm SL, Masana, China; KAUM–I. 122306, 22.2 mm SL,
west of Taminazaki, Tamina, China; KAUM–I. 122331, 25.9 mm SL, north of Taminazaki, Tamina, China;
KAUM–I. 122356, 35.3 mm SL, off Okidomari Port, Shimojiro, China; KAUM–I. 122372, 13.1 mm SL, off
Okidomari Port, Shimojiro, China; KAUM–I. 122721, 43.3 mm SL, off Yakomo, China; KAUM–I. 122722,
31.7 mm SL, off Yakomo, China; KAUM–I. 123542, 43.4 mm SL, Yaja, China; KAUM–I. 123559, 26.7 mm
SL, Yaja, China; KAUM–I. 123560, 12.8 mm SL, Yaja, China; KAUM–I. 123566, 31.5 mm SL, Yaja, China;
KUN-P 49005, 43.0 mm SL, Yaja, China; KUN-P 49019, 22.1 mm SL, Yaja, China.

Marine and freshwater fishes of Okinoerabu Island

29

Serranocirrhitus latus Watanabe 1949
[Jpn name: Hanagombe]
Sakurai (2018a): KAUM–I. 90152, 77.4 mm SL, Okidomari Port, Shimojiro, China. Sakurai (2019a):
KAUM–I. 90152, 77.4 mm SL, Okidomari Port, Shimojiro, China. This study: KAUM–I. 90152, 77.4 mm
SL, Okidomari Port, Shimojiro, China; KAUM–I. 122714, 20.0 mm SL, off Yakomo, China.
Variola albimarginata Baissac 1953
[Jpn name: Ojirobarahata]
KAUM–I. 122672, 156.6 mm SL, off Naikina Port, Nagamine, Wadomari; KAUM–I. 123503, 235.2 mm
SL, off Okidomari Port, Shimojiro, China; KAUM–I. 123512, 169.2 mm SL, off Okidomari Port, Shimojiro,
China; KAUM–I. 123521, 239.9 mm SL, off Okidomari Port, Shimojiro, China; KUN-P 48998, 209.7 mm
SL, off Okinoerabu Island.
Variola louti (Forsskål 1775)[Jpn name: Barahata]
Sakurai (2019a): KAUM–I. 122240, 74.2 mm SL, west of Taminazaki, Tamina, China. This study: KAUM–
I. 121860, 167.8 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 121992, 329.4 mm SL, China Port,
Serikaku, China; KAUM–I. 122240, 74.2 mm SL, west of Taminazaki, Tamina, China; KAUM–I. 122669,
336.0 mm SL, off Naikina Port, Nagamine, Wadomari.
FAMILY PSEUDOCHROMIDAE
Amsichthys knighti (Allen 1987)[Jpn name: Ometanabatamegisu]
Yoshida (2018a): KAUM–I. 90769, 10.7 mm SL, Naikina Port, Nagamine, Wadomari. Yoshida (2019a):
KAUM–I. 90769, 10.7 mm SL, Naikina Port, Nagamine, Wadomari. This study: KAUM–I. 90769, 10.7 mm
SL, Naikina Port, Nagamine, Wadomari.
Labracinus cyclophthalmus (Müller & Troschel 1849)[Jpn name: Megisu]
Yoshigou et al. (2005a): HMNH 7996, Nishibaru, Wadomari. This study: KAUM–I. 90407, 107.5 mm
SL, Inobe Port, Inobe, Wadomari; KAUM–I. 90408, 92.7 mm SL, Inobe Port, Inobe, Wadomari; KAUM–
I. 90409, 94.7 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 90410, 92.4 mm SL, Inobe Port, Inobe,
Wadomari; KAUM–I. 90411, 103.4 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 122799, 104.7 mm SL,
Kunigami, Wadomari.
Pictichromis porphyreus (Lubbock & Goldman 1974)[Jpn name: Kurenainisesuzume]
Yoshida (2018a): KAUM–I. 90636, 17.7 mm SL, Naikina Port, Nagamine, Wadomari. Yoshida (2019a):
KAUM–I. 90636, 17.7 mm SL, Naikina Port, Nagamine, Wadomari. This study: KAUM–I. 90636, 17.7mm SL, Naikina Port, Nagamine, Wadomari; KAUM–I. 90637, 18.4 mm SL, Naikina Port, Nagamine,
Wadomari; KAUM–I. 122343, 37.6 mm SL, off Okidomari Port, Shimojiro, China; KAUM–I. 122344, 37.3mm SL, off Okidomari Port, Shimojiro, China.
Pseudochromis cyanotaenia Bleeker 1857
[Jpn name: Ryukyunisesuzume]
Yoshigou et al. (2005a): HMNH 7995, 3 specimens, Nishibaru, Wadomari. This study: KAUM–I. 90016,
35.5 mm SL, Okidomari Port, Shimojiro, China; KAUM–I. 90057, 13.7 mm SL, Okidomari Port, Shimojiro,
China; KAUM–I. 123597, 26.4 mm SL, off China; KUN-P 48953, 37.5 mm SL, Ujiji Beach, Ashikiyora,
China; KUN-P 49044, 34.2 mm SL, China Port, Serikaku, China.

30

H. Motomura and K. Uehara

Pseudochromis luteus Aoyagi 1943
[Jpn name: Koganenisesuzume]
KAUM–I. 90140, 23.6 mm SL, Okidomari Port, Shimojiro, China; KAUM–I. 122809, 35.2 mm SL,
Sumiyoshi Port, Sumiyoshi, China; KAUM–I. 122810, 42.0 mm SL, Sumiyoshi Port, Sumiyoshi, China;
KAUM–I. 123588, 42.3 mm SL, off China.
Pseudochromis marshallensis Schultz 1953
[Jpn name: Hoshinisesuzume]
KAUM–I. 90133, 24.8 mm SL, Okidomari Port, Shimojiro, China; KAUM–I. 90402, 19.7 mm SL, Inobe
Port, Inobe, Wadomari; KAUM–I. 90638, 15.5 mm SL, Naikina Port, Nagamine, Wadomari; KAUM–I.
90648, 20.6 mm SL, Naikina Port, Nagamine, Wadomari; KAUM–I. 90758, 26.7 mm SL, Naikina Port,
Nagamine, Wadomari; KAUM–I. 123465, 34.9 mm SL, Kasaishi Beach, Kibiru, Wadomari.
Pseudochromis tapeinosoma Bleeker 1853
[Jpn name: Katsuironisesuzume]
Yoshigou et al. (2005a): HMNH 7864, Naikina, Nagamine, Wadomari; HMNH 7956, Nishibaru, Wadomari.
Pseudoplesiops rosae Schultz 1943
[Jpn name: Tanabatamegisu]
KAUM–I. 90046, 24.1 mm SL, Kasaishi Beach, Kibiru, Wadomari; KAUM–I. 90385, 11.7 mm SL, Inobe
Port, Inobe, Wadomari; KAUM–I. 90622, 9.7 mm SL, Naikina Port, Nagamine, Wadomari; KAUM–I.
122376, 24.3 mm SL, off Okidomari Port, Shimojiro, China.
FAMILY PLESIOPIDAE
Assessor randalli Allen & Kuiter 1976
[Jpn name: Tsubametanabatauo]
KAUM–I. 90741, 42.4 mm SL, Naikina Port, Nagamine, Wadomari; KAUM–I. 122351, 51.6 mm SL, off
Okidomari Port, Shimojiro, China; KAUM–I. 122806, 29.1 mm SL, Sumiyoshi Port, Sumiyoshi, China.
Belonepterygion fasciolatum (Ogilby 1889)[Jpn name: Togetanabatauo]
KAUM–I. 2102, 21.1 mm SL, off Okinoerabu Island; KAUM–I. 14641, 20.0 mm SL, off Okinoerabu Island;
KAUM–I. 14642, 17.3 mm SL, off Okinoerabu Island.
Calloplesiops altivelis (Steindachner 1903)[Jpn name: Shimofuritanabatauo]
KAUM–I. 90059, 82.8 mm SL, Okidomari Port, Shimojiro, China.
Plesiops coeruleolineatus Rüppell 1835
[Jpn name: Tanabatauo]
Yoshigou et al. (2005a): HMNH 7862, Naikina, Nagamine, Wadomari; HMNH 7863, Naikina, Nagamine,
Wadomari; HMNH 7890, Okidomari, China; HMNH 7891, Okidomari, China; HMNH 7892, Okidomari,China. This study: KAUM–I. 90192, 10.0 mm SL, Yakomo Beach, Yakomo, China; KAUM–I. 90448, 50.5mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 90666, 36.0 mm SL, Wanjo Beach, Azefu, Wadomari;
KAUM–I. 90667, 32.0 mm SL, Wanjo Beach, Azefu, Wadomari; KAUM–I. 121903, 62.5 mm SL,
Okidomari Port, Shimojiro, China; KAUM–I. 122074, 28.7 mm SL, Yakomo Beach, Yakomo, China;
KAUM–I. 122075, 24.3 mm SL, Yakomo Beach, Yakomo, China; KAUM–I. 122757, 43.0 mm SL, Kasaishi
Beach, Kibiru, Wadomari; KAUM–I. 122777, 25.5 mm SL, Kasaishi Beach, Kibiru, Wadomari; KAUM–I.

Marine and freshwater fishes of Okinoerabu Island

31

123455, 43.0 mm SL, Kasaishi Beach, Kibiru, Wadomari; KAUM–I. 123467, 34.2 mm SL, Kasaishi Beach,
Kibiru, Wadomari; KAUM–I. 123468, 26.0 mm SL, Kasaishi Beach, Kibiru, Wadomari; KAUM–I. 123486,
21.5 mm SL, Kasaishi Beach, Kibiru, Wadomari; KAUM–I. 123487, 21.1 mm SL, Kasaishi Beach, Kibiru,
Wadomari; KAUM–I. 123488, 20.3 mm SL, Kasaishi Beach, Kibiru, Wadomari; KUN-P 49134, 48.9 mm
SL, Kasaishi Beach, Kibiru, Wadomari.
Plesiops verecundus Mooi 1995
[Jpn name: Amamitanabatauo]
KAUM–I. 122773, 18.8 mm SL, Kasaishi Beach, Kibiru, Wadomari.
FAMILY PRIACANTHIDAE
Cookeolus japonicus (Cuvier 1829)[Jpn name: Chikamekintoki]
KAUM–I. 121761, 376.8 mm SL, off China; KAUM–I. 122684, 295.3 mm SL, off Okinoerabu Island.
Heteropriacanthus carolinus (Cuvier 1829)[Jpn name: Gomahirekintoki]
KAUM–I. 90589, 104.5 mm SL, Okidomari Port, Shimojiro, China; KAUM–I. 121752, 175.9 mm SL,
Sumiyoshi Port, Sumiyoshi, China.
FAMILY APOGONIDAE
Apogon crassiceps Garman 1903
[Jpn name: Akafujitenjikudai]
KAUM–I. 122115, 24.6 mm SL, Sumiyoshi Port, Sumiyoshi, China.
Apogon kominatoensis Ebina 1935
[Jpn name: Kominatotenjikudai]
Yoshigou et al. (2005a, as Apogon coccineus): HMNH 7922, Okidomari, China; HMNH 7923, Okidomari,China; HMNH 7941, Okidomari, China; HMNH 7957, Nishibaru, Wadomari; HMNH 7958, Nishibaru,
Wadomari; HMNH 7959, Nishibaru, Wadomari; HMNH 7960, Nishibaru, Wadomari. This study: KAUM–
I. 90055, 22.1 mm SL, Okidomari Port, Shimojiro, China; KAUM–I. 90066, 27.1 mm SL, Okidomari Port,
Shimojiro, China; KAUM–I. 122241, 39.2 mm SL, west of Taminazaki, Tamina, China; KUN-P 49085, 30.0mm SL, China Port, Serikaku, China.
Apogon semiornatus Peters 1876
[Jpn name: Yamitenjikudai]
KAUM–I. 122292, 43.3 mm SL, west of Taminazaki, Tamina, China; KAUM–I. 122293, 34.6 mm SL,
west of Taminazaki, Tamina, China; KAUM–I. 122294, 31.3 mm SL, west of Taminazaki, Tamina, China;
KAUM–I. 122369, 34.5 mm SL, west of Taminazaki, Tamina, China.
Apogon sp.
[Jpn name: Ryukyuishimochi]
Yoshigou et al. (2005a, as Apogon indicus): HMNH 7893, Okidomari, China; HMNH 7894, Okidomari,China.
Apogon spp. (unidentified)[Jpn name: N/A]
KAUM–I. 2123, 29.2 mm SL, off Okinoerabu Island; KAUM–I. 20806, 30.2 mm SL, off Okinoerabu

32

H. Motomura and K. UeharaIsland; KAUM–I. 20807, 25.3 mm SL, off Okinoerabu Island; KAUM–I. 20808, 26.9 mm SL, off
Okinoerabu Island; KAUM–I. 20809, 25.7 mm SL, off Okinoerabu Island; KAUM–I. 20810, 24.6 mm SL,
off Okinoerabu Island; KAUM–I. 20811, 30.1 mm SL, off Okinoerabu Island; KAUM–I. 20812, 27.7 mm
SL, off Okinoerabu Island; KAUM–I. 20813, 26.1 mm SL, off Okinoerabu Island; KAUM–I. 20814, 26.5mm SL, off Okinoerabu Island; KAUM–I. 20815, 25.3 mm SL, off Okinoerabu Island; KAUM–I. 20816,
26.7 mm SL, off Okinoerabu Island; KAUM–I. 20817, 24.8 mm SL, off Okinoerabu Island; KAUM–
I. 20818, 24.6 mm SL, off Okinoerabu Island; KAUM–I. 20819, 28.6 mm SL, off Okinoerabu Island;
KAUM–I. 20820, 25.8 mm SL, off Okinoerabu Island; KAUM–I. 20821, 28.2 mm SL, off Okinoerabu
Island; KAUM–I. 20822, 24.9 mm SL, off Okinoerabu Island; KAUM–I. 20823, 24.3 mm SL, off
Okinoerabu Island; KAUM–I. 20824, 23.8 mm SL, off Okinoerabu Island; KAUM–I. 20825, 22.4 mm SL,
off Okinoerabu Island; KAUM–I. 20826, 19.3 mm SL, off Okinoerabu Island; KAUM–I. 20827, 19.0 mm
SL, off Okinoerabu Island; KAUM–I. 20828, 15.0 mm SL, off Okinoerabu Island; KAUM–I. 90381, 28.8mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 121996, 20.9 mm SL, Masana, China; KAUM–I. 122011,12.7 mm SL, Masana, China; KAUM–I. 122040, 25.1 mm SL, Masana, China; KAUM–I. 122051, 12.9mm SL, Masana, China; KAUM–I. 122242, 30.9 mm SL, west of Taminazaki, Tamina, China; KAUM–I.
122243, 13.2 mm SL, west of Taminazaki, Tamina, China; KAUM–I. 122367, 29.9 mm SL, off OkidomariPort, Shimojiro, China; KAUM–I. 122368, 18.7 mm SL, off Okidomari Port, Shimojiro, China; KAUM–I.
122375, 11.2 mm SL, off Okidomari Port, Shimojiro, China; KAUM–I. 122612, 22.1 mm SL, off Yakomo,
China; KAUM–I. 123543, 18.0 mm SL, Yaja, China; KAUM–I. 123561, 32.3 mm SL, Yaja, China; KAUM–
I. 123580, 32.4 mm SL, off China; KAUM–I. 123581, 24.6 mm SL, off China; KUN-P 49024, 16.3 mm SL,
China Port, Serikaku, China.
Apogonichthys ocellatus (Weber 1913)[Jpn name: Matoshibori]
KAUM–I. 122039, 20.4 mm SL, Masana, China;
Cercamia eremia (Allen 1987)[Jpn name: Sakuratenjikudai]
KAUM–I. 90382, 33.5 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 90641, 33.4 mm SL, Naikina Port,
Nagamine, Wadomari; KAUM–I. 90642, 26.4 mm SL, Naikina Port, Nagamine, Wadomari; KAUM–I.
90643, 16.6 mm SL, Naikina Port, Nagamine, Wadomari.
Cheilodipterus macrodon (Lacepède 1802)[Jpn name: Ryukyuyaraiishimochi]
KAUM–I. 90744, 32.0 mm SL, Naikina Port, Nagamine, Wadomari; KAUM–I. 121852, 169.5 mm SL,
China Port, Serikaku, China; KAUM–I. 122036, 75.8 mm SL, Masana, China; KAUM–I. 122805, 18.2mm SL, Sumiyoshi Port, Sumiyoshi, China; KAUM–I. 132172, 22.5 mm SL, west of Taminazaki, Tamina,
China.
Cheilodipterus quinquelineatus Cuvier 1828
[Jpn name: Yaraiishimochi]
Yoshida (2018b): KAUM–I. 90428, 16.4 mm SL, Inobe Port, Inobe, Wadomari. Yoshida (2019b): KAUM–
I. 90428, 16.4 mm SL, Inobe Port, Inobe, Wadomari. This study: KAUM–I. 1927, 37.1 mm SL, off
Okinoerabu Island; KAUM–I. 14274, 34.5 mm SL, off Okinoerabu Island; KAUM–I. 14275, 33.6 mm SL,
off Okinoerabu Island; KAUM–I. 14276, 30.5 mm SL, off Okinoerabu Island; KAUM–I. 90092, 23.9 mm
SL, Kasaishi Beach, Kibiru, Wadomari; KAUM–I. 90428, 16.4 mm SL, Inobe Port, Inobe, Wadomari.

Marine and freshwater fishes of Okinoerabu Island

33

Fowleria isostigma (Jordan & Seale 1906)[Jpn name: Nahamatoishimochi]
Yoshida (2018b): KAUM–I. 90768, 7.9 mm SL, Naikina Port, Nagamine, Wadomari. Yoshida (2019b):
KAUM–I. 90768, 7.9 mm SL, Naikina Port, Nagamine, Wadomari. This study: KAUM–I. 90768, 7.9 mm
SL, Naikina Port, Nagamine, Wadomari; KAUM–I. 122716, 16.4 mm SL, off Yakomo, China.
Fowleria vaiulae (Jordan & Seale 1906)[Jpn name: Shiboridamashi]
KAUM–I. 122715, 11.8 mm SL, off Yakomo, China.
Gymnapogon vanderbilti (Fowler 1938)[Jpn name: Komonkudaribozugisu]
Yoshida (2018b): KAUM–I. 90635, 21.7 mm SL, Naikina Port, Nagamine, Wadomari. Yoshida (2019b):
KAUM–I. 90635, 21.7 mm SL, Naikina Port, Nagamine, Wadomari. This study: KAUM–I. 90635, 21.7 mm
SL, Naikina Port, Nagamine, Wadomari.
Nectamia bandanensis (Bleeker 1854)[Jpn name: Bandaishimochi]
KAUM–I. 1918, 24.5 mm SL, off Okinoerabu Island; KAUM–I. 14515, 23.9 mm SL, off Okinoerabu Island;
KAUM–I. 14516, 25.0 mm SL, off Okinoerabu Island; KAUM–I. 14517, 23.4 mm SL, off Okinoerabu
Island; KAUM–I. 14518, 23.8 mm SL, off Okinoerabu Island; KAUM–I. 14519, 20.9 mm SL, off
Okinoerabu Island; KAUM–I. 14520, 23.6 mm SL, off Okinoerabu Island; KAUM–I. 14521, 24.4 mm SL,
off Okinoerabu Island; KAUM–I. 14522, 27.0 mm SL, off Okinoerabu Island; KAUM–I. 14523, 22.9 mm
SL, off Okinoerabu Island; KAUM–I. 14524, 22.1 mm SL, off Okinoerabu Island; KAUM–I. 14525, 25.6mm SL, off Okinoerabu Island; KAUM–I. 14526, 21.4 mm SL, off Okinoerabu Island; KAUM–I. 14527,
22.9 mm SL, off Okinoerabu Island; KAUM–I. 14528, 26.7 mm SL, off Okinoerabu Island; KAUM–I.
14529, 21.7 mm SL, off Okinoerabu Island; KAUM–I. 14530, 20.3 mm SL, off Okinoerabu Island; KAUM–
I. 14531, 23.4 mm SL, off Okinoerabu Island; KAUM–I. 14532, 22.9 mm SL, off Okinoerabu Island;
KAUM–I. 14533, 22.6 mm SL, off Okinoerabu Island; KAUM–I. 14534, damaged, off Okinoerabu Island;
KAUM–I. 14535, 20.6 mm SL, off Okinoerabu Island; KAUM–I. 14536, 21.5 mm SL, off Okinoerabu
Island; KAUM–I. 14537, damaged, off Okinoerabu Island; KAUM–I. 14538, 20.7 mm SL, off Okinoerabu
Island.
Nectamia fusca (Quoy & Gaimard 1825)[Jpn name: Hososujinamidatenjikudai]
KAUM–I. 90132, 26.0 mm SL, Okidomari Port, Shimojiro, China; KAUM–I. 90187, 24.3 mm SL, Yakomo
Beach, Yakomo, China; KAUM–I. 90391, 27.2 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 90485,
20.8 mm SL, Ujiji Beach, Ashikiyora, China; KAUM–I. 90486, 18.8 mm SL, Ujiji Beach, Ashikiyora,
China; KAUM–I. 90487, 21.8 mm SL, Ujiji Beach, Ashikiyora, China; KAUM–I. 90488, 21.9 mm SL, Ujiji
Beach, Ashikiyora, China; KAUM–I. 90489, 22.8 mm SL, Ujiji Beach, Ashikiyora, China; KAUM–I. 90490,
24.5 mm SL, Ujiji Beach, Ashikiyora, China; KAUM–I. 90491, 19.1 mm SL, Ujiji Beach, Ashikiyora,
China; KAUM–I. 90601, 60.4 mm SL, Okidomari Port, Shimojiro, China; KAUM–I. 90602, 83.0 mm SL,
Okidomari Port, Shimojiro, China; KAUM–I. 90603, 81.0 mm SL, Okidomari Port, Shimojiro, China;
KAUM–I. 90626, 24.6 mm SL, Naikina Port, Nagamine, Wadomari; KAUM–I. 121947, 73.6 mm SL, Inobe
Port, Inobe, Wadomari; KAUM–I. 121957, 71.8 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 121958,
67.1 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 121959, 60.7 mm SL, Inobe Port, Inobe, Wadomari;
KAUM–I. 123451, 63.5 mm SL, Kasaishi Beach, Kibiru, Wadomari.

34

H. Motomura and K. Uehara

Nectamia savayensis (Günther 1872)[Jpn name: Namidatenjikudai]
KAUM–I. 90507, 9.5 mm SL, Ujiji Beach, Ashikiyora, China; KUN-P 48918, 65.7 mm SL, Yakomo Beach,
Yakomo, China.
Nectamia sp.
[Jpn name: N/A]
KAUM–I. 90509, 15.4 mm SL, Ujiji Beach, Ashikiyora, China; KAUM–I. 90536, 25.5 mm SL, Ujiji Beach,
Ashikiyora, China; KAUM–I. 90687, 10.9 mm SL, Ujiji Beach, Ashikiyora, China.
Ostorhinchus angustatus (Smith & Radcliffe 1911)[Jpn name: Usujimaishimochi]
Yoshigou et al. (2005a): HMNH 7924, Okidomari, China; HMNH 7997, Nishibaru, Wadomari. This
study: KAUM–I. 90531, 25.7 mm SL, Ujiji Beach, Ashikiyora, China; KAUM–I. 122001, 53.5 mm SL,
Masana, China; KAUM–I. 122002, 50.5 mm SL, Masana, China; KAUM–I. 122007, 30.4 mm SL, Masana,
China; KAUM–I. 122101, 11.1 mm SL, Yakomo Beach, Yakomo, China; KAUM–I. 122132, 24.4 mm SL,
Sumiyoshi Port, Sumiyoshi, China; KAUM–I. 122246, 30.0 mm SL, west of Taminazaki, Tamina, China;
KAUM–I. 122266, 39.1 mm SL, west of Taminazaki, Tamina, China; KAUM–I. 122268, 35.9 mm SL,
west of Taminazaki, Tamina, China; KAUM–I. 122370, 36.4 mm SL, west of Taminazaki, Tamina, China;
KAUM–I. 122718, 21.7 mm SL, off Yakomo, China; KAUM–I. 123549, 18.1 mm SL, Yaja, China.
Ostorhinchus apogonoides (Bleeker 1856)[Jpn name: Aohanatenjikudai]
Underwater photograph (Pl. 4-C).
Ostorhinchus aureus (Lacepède 1802)[Jpn name: Aosujitenjikudai]
Underwater photograph (Pl. 4-D).
Ostorhinchus cookii (Macleay 1881)[Jpn name: Sujiishimochi]
KAUM–I. 90534, 63.1 mm SL, Ujiji Beach, Ashikiyora, China; KAUM–I. 90535, 62.0 mm SL, Ujiji Beach,
Ashikiyora, China; KAUM–I. 90649, 61.2 mm SL, Wanjo Beach, Azefu, Wadomari; KAUM–I. 90654, 75.3mm SL, Wanjo Beach, Azefu, Wadomari; KAUM–I. 90655, 64.2 mm SL, Wanjo Beach, Azefu, Wadomari;
KAUM–I. 121883, 43.2 mm SL, Okidomari Port, Shimojiro, China; KAUM–I. 121960, 75.8 mm SL, Inobe
Port, Inobe, Wadomari; KAUM–I. 121961, 76.1 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 121962,
75.0 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 121963, 72.2 mm SL, Inobe Port, Inobe, Wadomari;
KAUM–I. 121964, 73.5 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 121965, 78.9 mm SL, Inobe
Port, Inobe, Wadomari; KAUM–I. 121966, 58.6 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 121987,
57.6 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 121988, 68.0 mm SL, Inobe Port, Inobe, Wadomari;
KAUM–I. 122129, 65.7 mm SL, Beach Rock, Tokudoki, China; KAUM–I. 122141, 38.3 mm SL, Beach
Rock, Tokudoki, China; KAUM–I. 122198, 66.3 mm SL, Wadomari Port, Wadomari; KAUM–I. 122244,
45.0 mm SL, west of Taminazaki, Tamina, China; KAUM–I. 122380, 54.9 mm SL, Ujiji Beach, Ashikiyora,
China; KAUM–I. 122381, 63.4 mm SL, Ujiji Beach, Ashikiyora, China; KAUM–I. 122382, 69.6 mm
SL, Ujiji Beach, Ashikiyora, China; KAUM–I. 122754, 23.9 mm SL, Kasaishi Beach, Kibiru, Wadomari;
KAUM–I. 123404, 64.4 mm SL, Ujiji Beach, Ashikiyora, China; KUN-P 49083, 45.5 mm SL, China Port,
Serikaku, China; KUN-P 49084, 36.7 mm SL, China Port, Serikaku, China.

Marine and freshwater fishes of Okinoerabu Island

35

Ostorhinchus doederleini (Jordan & Snyder 1901)[Jpn name: Osujiishimochi]
KAUM–I. 121944, 55.3 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 132336, 14.6 mm SL, Okidomari
Seaside Park, Shimojiro, China; KAUM–I. 132337, 14.8 mm SL, Okidomari Seaside Park, Shimojiro, China.
Ostorhinchus endekataenia (Bleeker 1852)[Jpn name: Kosujiishimochi]
KAUM–I. 2063, 28.2 mm SL, off Okinoerabu Island; KAUM–I. 12938, 32.3 mm SL, off Okinoerabu Island;
KAUM–I. 12939, 29.3 mm SL, off Okinoerabu Island; KAUM–I. 12940, 26.9 mm SL, off Okinoerabu
Island; KAUM–I. 12941, 25.5 mm SL, off Okinoerabu Island; KAUM–I. 12942, 27.9 mm SL, off
Okinoerabu Island; KAUM–I. 12943, 24.3 mm SL, off Okinoerabu Island; KAUM–I. 12944, 25.1 mm SL,
off Okinoerabu Island; KAUM–I. 12945, 22.9 mm SL, off Okinoerabu Island; KAUM–I. 12946, 21.7 mm
SL, off Okinoerabu Island; KAUM–I. 12947, 21.3 mm SL, off Okinoerabu Island; KAUM–I. 12948, 21.6mm SL, off Okinoerabu Island; KAUM–I. 12949, 17.9 mm SL, off Okinoerabu Island; KAUM–I. 12950,
28.0 mm SL, off Okinoerabu Island; KAUM–I. 12951, 29.5 mm SL, off Okinoerabu Island; KAUM–I.
12952, 35.7 mm SL, off Okinoerabu Island; KAUM–I. 12953, 27.7 mm SL, off Okinoerabu Island; KAUM–
I. 12954, 27.1 mm SL, off Okinoerabu Island; KAUM–I. 12955, 22.5 mm SL, off Okinoerabu Island;
KAUM–I. 12956, 24.9 mm SL, off Okinoerabu Island; KAUM–I. 12957, 27.7 mm SL, off Okinoerabu
Island; KAUM–I. 12958, 21.8 mm SL, off Okinoerabu Island; KAUM–I. 12959, 25.1 mm SL, off
Okinoerabu Island; KAUM–I. 12960, 21.6 mm SL, off Okinoerabu Island; KAUM–I. 12961, 18.4 mm SL,
off Okinoerabu Island; KAUM–I. 12962, 20.0 mm SL, off Okinoerabu Island; KAUM–I. 12963, 27.4 mm
SL, off Okinoerabu Island; KAUM–I. 12964, 32.3 mm SL, off Okinoerabu Island; KAUM–I. 12965, 25.2mm SL, off Okinoerabu Island; KAUM–I. 12966, 31.6 mm SL, off Okinoerabu Island; KAUM–I. 12967,
34.3 mm SL, off Okinoerabu Island; KAUM–I. 12968, 29.6 mm SL, off Okinoerabu Island; KAUM–I.
12969, 31.6 mm SL, off Okinoerabu Island; KAUM–I. 12970, 35.1 mm SL, off Okinoerabu Island; KAUM–
I. 12971, 31.9 mm SL, off Okinoerabu Island; KAUM–I. 12972, 27.9 mm SL, off Okinoerabu Island;
KAUM–I. 12973, 29.2 mm SL, off Okinoerabu Island; KAUM–I. 12974, 28.8 mm SL, off Okinoerabu
Island; KAUM–I. 12975, 26.3 mm SL, off Okinoerabu Island; KAUM–I. 12976, 29.5 mm SL, off
Okinoerabu Island; KAUM–I. 12977, 30.4 mm SL, off Okinoerabu Island; KAUM–I. 12978, 28.5 mm SL,
off Okinoerabu Island; KAUM–I. 12979, 29.7 mm SL, off Okinoerabu Island; KAUM–I. 12980, 27.9 mm
SL, off Okinoerabu Island; KAUM–I. 12981, 28.7 mm SL, off Okinoerabu Island; KAUM–I. 12982, 30.4mm SL, off Okinoerabu Island; KAUM–I. 12983, 24.4 mm SL, off Okinoerabu Island; KAUM–I. 12984,
26.5 mm SL, off Okinoerabu Island; KAUM–I. 12985, 29.6 mm SL, off Okinoerabu Island; KAUM–I.
12986, 31.9 mm SL, off Okinoerabu Island; KAUM–I. 12987, 29.9 mm SL, off Okinoerabu Island; KAUM–
I. 12988, 24.6 mm SL, off Okinoerabu Island; KAUM–I. 12989, 23.8 mm SL, off Okinoerabu Island;
KAUM–I. 12990, 32.8 mm SL, off Okinoerabu Island; KAUM–I. 12991, 23.3 mm SL, off Okinoerabu
Island; KAUM–I. 12992, 31.8 mm SL, off Okinoerabu Island; KAUM–I. 12993, 29.6 mm SL, off
Okinoerabu Island; KAUM–I. 12994, 29.9 mm SL, off Okinoerabu Island; KAUM–I. 12995, 29.9 mm SL,
off Okinoerabu Island; KAUM–I. 12996, 26.8 mm SL, off Okinoerabu Island; KAUM–I. 12997, 30.0 mm
SL, off Okinoerabu Island; KAUM–I. 12998, 29.8 mm SL, off Okinoerabu Island; KAUM–I. 12999, 28.9mm SL, off Okinoerabu Island; KAUM–I. 13000, 22.3 mm SL, off Okinoerabu Island; KAUM–I. 13001,
25.6 mm SL, off Okinoerabu Island; KAUM–I. 13002, 23.7 mm SL, off Okinoerabu Island; KAUM–I.
13003, 22.5 mm SL, off Okinoerabu Island; KAUM–I. 13004, 20.5 mm SL, off Okinoerabu Island; KAUM–
I. 13005, 23.6 mm SL, off Okinoerabu Island; KAUM–I. 13006, 25.4 mm SL, off Okinoerabu Island;
KAUM–I. 13007, 23.8 mm SL, off Okinoerabu Island; KAUM–I. 13008, 23.0 mm SL, off Okinoerabu
Island; KAUM–I. 13009, 22.4 mm SL, off Okinoerabu Island; KAUM–I. 13010, 25.1 mm SL, off
Okinoerabu Island; KAUM–I. 13011, 26.5 mm SL, off Okinoerabu Island; KAUM–I. 13012, 22.2 mm SL,
off Okinoerabu Island; KAUM–I. 13013, 22.8 mm SL, off Okinoerabu Island; KAUM–I. 13014, 20.5 mm

36

H. Motomura and K. UeharaSL, off Okinoerabu Island; KAUM–I. 13015, 24.5 mm SL, off Okinoerabu Island; KAUM–I. 13016, 28.3mm SL, off Okinoerabu Island; KAUM–I. 13017, 22.5 mm SL, off Okinoerabu Island; KAUM–I. 13018,
21.9 mm SL, off Okinoerabu Island; KAUM–I. 13019, 21.4 mm SL, off Okinoerabu Island; KAUM–I.
13020, 21.9 mm SL, off Okinoerabu Island; KAUM–I. 13021, 24.7 mm SL, off Okinoerabu Island; KAUM–
I. 13022, 24.3 mm SL, off Okinoerabu Island; KAUM–I. 13023, 29.9 mm SL, off Okinoerabu Island;
KAUM–I. 13024, 20.4 mm SL, off Okinoerabu Island; KAUM–I. 13025, 20.7 mm SL, off Okinoerabu
Island; KAUM–I. 13026, 22.4 mm SL, off Okinoerabu Island; KAUM–I. 13027, 22.7 mm SL, off
Okinoerabu Island; KAUM–I. 13028, 24.3 mm SL, off Okinoerabu Island; KAUM–I. 13029, 26.6 mm SL,
off Okinoerabu Island; KAUM–I. 13030, 21.7 mm SL, off Okinoerabu Island; KAUM–I. 13031, 21.5 mm
SL, off Okinoerabu Island; KAUM–I. 13032, 23.3 mm SL, off Okinoerabu Island; KAUM–I. 13033, 29.3mm SL, off Okinoerabu Island; KAUM–I. 13034, 21.5 mm SL, off Okinoerabu Island; KAUM–I. 13035,
23.9 mm SL, off Okinoerabu Island; KAUM–I. 13036, 18.3 mm SL, off Okinoerabu Island; KAUM–I.
13037, 20.8 mm SL, off Okinoerabu Island; KAUM–I. 13038, 19.4 mm SL, off Okinoerabu Island; KAUM–
I. 13039, 20.4 mm SL, off Okinoerabu Island; KAUM–I. 13040, 22.9 mm SL, off Okinoerabu Island;
KAUM–I. 13041, 21.4 mm SL, off Okinoerabu Island; KAUM–I. 13042, 12.9 mm SL, off Okinoerabu
Island; KAUM–I. 122168, 30.3 mm SL, Beach Rock, Tokudoki, China.
Ostorhinchus nigrofasciatus (Lachner 1953)[Jpn name: Minamifutosujiishimochi]
Yoshigou et al. (2005a): HMNH 7961, Nishibaru, Wadomari; HMNH 7962, Nishibaru, Wadomari; HMNH
7963, Nishibaru, Wadomari; HMNH 7964, Nishibaru, Wadomari. Yoshida (2018b): KAUM–I. 90067, 30.3mm SL, Okidomari Port, Shimojiro, China. Yoshida (2019b): KAUM–I. 90067, 30.3 mm SL, OkidomariPort, Shimojiro, China. This study: KAUM–I. 90043, 17.7 mm SL, Kasaishi Beach, Kibiru, Wadomari;
KAUM–I. 90067, 30.3 mm SL, Okidomari Port, Shimojiro, China; KAUM–I. 90068, 36.7 mm SL,
Okidomari Port, Shimojiro, China; KAUM–I. 90069, 37.6 mm SL, Okidomari Port, Shimojiro, China;
KAUM–I. 90070, 21.2 mm SL, Okidomari Port, Shimojiro, China; KAUM–I. 90088, 29.0 mm SL, Kasaishi
Beach, Kibiru, Wadomari; KAUM–I. 90089, 23.1 mm SL, Kasaishi Beach, Kibiru, Wadomari; KAUM–I.
90090, 23.9 mm SL, Kasaishi Beach, Kibiru, Wadomari; KAUM–I. 90091, 32.5 mm SL, Kasaishi Beach,
Kibiru, Wadomari; KAUM–I. 90116, 68.6 mm SL, Wadomari Port, Wadomari; KAUM–I. 90117, 59.6 mm
SL, Wadomari Port, Wadomari; KAUM–I. 90492, 24.5 mm SL, Ujiji Beach, Ashikiyora, China; KAUM–I.
90506, 17.4 mm SL, Ujiji Beach, Ashikiyora, China; KAUM–I. 90511, 14.7 mm SL, Ujiji Beach, Ashikiyora,
China; KAUM–I. 90512, 14.4 mm SL, Ujiji Beach, Ashikiyora, China; KAUM–I. 90513, 23.1 mm SL, Ujiji
Beach, Ashikiyora, China; KAUM–I. 90514, 20.5 mm SL, Ujiji Beach, Ashikiyora, China; KAUM–I. 90529,
31.1 mm SL, Ujiji Beach, Ashikiyora, China; KAUM–I. 90530, 33.1 mm SL, Ujiji Beach, Ashikiyora,
China; KAUM–I. 90532, 26.5 mm SL, Ujiji Beach, Ashikiyora, China; KAUM–I. 90533, 32.6 mm SL, Ujiji
Beach, Ashikiyora, China; KAUM–I. 90604, 58.6 mm SL, Okidomari Port, Shimojiro, China; KAUM–I.
90605, 54.8 mm SL, Okidomari Port, Shimojiro, China; KAUM–I. 90606, 59.1 mm SL, Okidomari Port,
Shimojiro, China; KAUM–I. 90607, 63.3 mm SL, Okidomari Port, Shimojiro, China; KAUM–I. 90675, 13.9mm SL, Wanjo Beach, Azefu, Wadomari; KAUM–I. 90688, 31.4 mm SL, Ujiji Beach, Ashikiyora, China;
KAUM–I. 90782, 26.3 mm SL, Hanzaki, Wadomari; KAUM–I. 90783, 22.6 mm SL, Hanzaki, Wadomari;
KAUM–I. 90784, 12.4 mm SL, Hanzaki, Wadomari; KAUM–I. 90785, 12.2 mm SL, Hanzaki, Wadomari;
KAUM–I. 90786, 14.6 mm SL, Hanzaki, Wadomari; KAUM–I. 90787, 13.3 mm SL, Hanzaki, Wadomari;
KAUM–I. 90788, 15.0 mm SL, Hanzaki, Wadomari; KAUM–I. 90789, 14.3 mm SL, Hanzaki, Wadomari;
KAUM–I. 122366, 22.3 mm SL, off Okidomari Port, Shimojiro, China; KAUM–I. 122371, 21.0 mm SL, off
Okidomari Port, Shimojiro, China; KAUM–I. 132171, 21.8 mm SL, north of Taminazaki, Tamina, China;
KUN-P 48891, 29.9 mm SL, off Masana, China.

Marine and freshwater fishes of Okinoerabu Island

37

Ostorhinchus novemfasciatus (Cuvier 1828)[Jpn name: Tasujiishimochi]
KAUM–I. 1919, 52.7 mm SL, off Okinoerabu Island; KAUM–I. 14099, 44.6 mm SL, off Okinoerabu Island;
KAUM–I. 14100, 48.5 mm SL, off Okinoerabu Island; KAUM–I. 14101, 42.8 mm SL, off Okinoerabu
Island; KAUM–I. 14102, 39.5 mm SL, off Okinoerabu Island; KAUM–I. 14103, 34.8 mm SL, off
Okinoerabu Island; KAUM–I. 14104, 26.5 mm SL, off Okinoerabu Island; KAUM–I. 14105, 29.1 mm SL,
off Okinoerabu Island; KAUM–I. 14106, 29.4 mm SL, off Okinoerabu Island; KAUM–I. 14107, 32.7 mm
SL, off Okinoerabu Island; KAUM–I. 14108, 27.3 mm SL, off Okinoerabu Island; KAUM–I. 14109, 25.6mm SL, off Okinoerabu Island; KAUM–I. 14110, 24.5 mm SL, off Okinoerabu Island; KAUM–I. 14111,
27.1 mm SL, off Okinoerabu Island; KAUM–I. 14178, 21.6 mm SL, off Okinoerabu Island; KAUM–I.
14179, 21.3 mm SL, off Okinoerabu Island; KAUM–I. 14180, 24.9 mm SL, off Okinoerabu Island; KAUM–
I. 14181, 27.7 mm SL, off Okinoerabu Island; KAUM–I. 14182, 24.1 mm SL, off Okinoerabu Island;
KAUM–I. 14183, 22.1 mm SL, off Okinoerabu Island; KAUM–I. 14184, 20.0 mm SL, off Okinoerabu
Island; KAUM–I. 14185, 20.6 mm SL, off Okinoerabu Island; KAUM–I. 14186, 22.1 mm SL, off
Okinoerabu Island; KAUM–I. 14187, 21.4 mm SL, off Okinoerabu Island; KAUM–I. 14188, 18.3 mm SL,
off Okinoerabu Island; KAUM–I. 14189, 19.0 mm SL, off Okinoerabu Island; KAUM–I. 14190, 16.9 mm
SL, off Okinoerabu Island; KAUM–I. 121954, 79.4 mm SL, Inobe Port, Inobe, Wadomari.
Ostorhinchus properuptus (Whitley 1964)[Jpn name: Kinsenishimochi]
KAUM–I. 90379, 32.7 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 90380, 30.4 mm SL, Inobe Port,
Inobe, Wadomari; KAUM–I. 90389, 14.6 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 90455, 51.9 mm
SL, Inobe Port, Inobe, Wadomari; KAUM–I. 90456, 52.0 mm SL, Inobe Port, Inobe, Wadomari; KAUM–
I. 121945, 52.9 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 122811, 18.8 mm SL, Sumiyoshi Port,
Sumiyoshi, China; KAUM–I. 123583, 52.1 mm SL, off China; KAUM–I. 123584, 47.7 mm SL, off China.
Ostorhinchus sp.
[Jpn name: N/A]
KAUM–I. 90527, 9.9 mm SL, Ujiji Beach, Ashikiyora, China; KAUM–I. 122173, 15.5 mm SL, Beach Rock,Tokudoki, China; KAUM–I. 122254, 17.9 mm SL, west of Taminazaki, Tamina, China; KAUM–I. 122388,
40.0 mm SL, Ujiji Beach, Ashikiyora, China; KAUM–I. 122648, 20.1 mm SL, China Port, Serikaku, China;
KAUM–I. 123405, 60.7 mm SL, Ujiji Beach, Ashikiyora, China.
Ostorhinchus taeniophorus (Regan 1908)[Jpn name: Misujitenjikudai]
Yoshigou et al. (2005a): HMNH 7865, Naikina, Nagamine, Wadomari. This study: KAUM–I. 101613, 66.6mm SL, Kunigami, Wadomari; KAUM–I. 122076, 38.0 mm SL, Yakomo Beach, Yakomo, China; KAUM–
I. 122077, 38.3 mm SL, Yakomo Beach, Yakomo, China; KAUM–I. 122122, 26.5 mm SL, Beach Rock,Tokudoki, China; KAUM–I. 122131, 23.8 mm SL, Beach Rock, Tokudoki, China; KAUM–I. 133537, 28.5mm SL, Beach Rock, Tokudoki, China; KAUM–I. 133538, 22.9 mm SL, Beach Rock, Tokudoki, China;
KAUM–I. 133539, 23.3 mm SL, Beach Rock, Tokudoki, China; KAUM–I. 133540, 16.4 mm SL, Beach
Rock, Tokudoki, China; KAUM–I. 133541, 14.9 mm SL, Beach Rock, Tokudoki, China; KUN-P 48882,
52.9 mm SL, Okidomari Port, Shimojiro, China; KUN-P 48915, 37.4 mm SL, Yakomo Beach, Yakomo,
China; KUN-P 48916, 39.7 mm SL, Yakomo Beach, Yakomo, China; KUN-P 49065, 14.9 mm SL, China
Port, Serikaku, China; KUN-P 49071, 14.6 mm SL, China Port, Serikaku, China; KUN-P 49077, 21.0 mm
SL, China Port, Serikaku, China; KUN-P 49118, 63.6 mm SL, Kasaishi Beach, Kibiru, Wadomari; KUN-P
49119, 52.3 mm SL, Kasaishi Beach, Kibiru, Wadomari; KUN-P 49120, 48.6 mm SL, Kasaishi Beach,
Kibiru, Wadomari; KUN-P 49121, 42.2 mm SL, Kasaishi Beach, Kibiru, Wadomari; KUN-P 49122, 37.6 mm

38

H. Motomura and K. Uehara

SL, Kasaishi Beach, Kibiru, Wadomari; KUN-P 49123, 43.9 mm SL, Kasaishi Beach, Kibiru, Wadomari;
KUN-P 49124, 46.4 mm SL, Kasaishi Beach, Kibiru, Wadomari.
Pristiapogon exostigma (Jordan & Starks 1906)[Jpn name: Yukataishimochi]
Yoshida (2018b): KAUM–I. 90094, 35.5 mm SL, Kasaishi Beach, Kibiru, Wadomari. Yoshida (2019b):
KAUM–I. 90094, 35.5 mm SL, Kasaishi Beach, Kibiru, Wadomari. This study: KAUM–I. 90093, 25.1mm SL, Kasaishi Beach, Kibiru, Wadomari; KAUM–I. 90094, 35.5 mm SL, Kasaishi Beach, Kibiru,
Wadomari; KAUM–I. 90600, 80.0 mm SL, Okidomari Port, Shimojiro, China; KAUM–I. 90608, 87.9mm SL, Okidomari Port, Shimojiro, China; KAUM–I. 90609, 95.1 mm SL, Okidomari Port, Shimojiro,
China; KAUM–I. 90610, 83.6 mm SL, Okidomari Port, Shimojiro, China; KAUM–I. 90611, 89.9 mm SL,
Okidomari Port, Shimojiro, China; KAUM–I. 90646, 32.0 mm SL, Naikina Port, Nagamine, Wadomari;
KAUM–I. 121946, 83.7 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 121955, 61.0 mm SL, Inobe
Port, Inobe, Wadomari; KAUM–I. 121956, 77.3 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 122222,
71.3 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 122223, 84.4 mm SL, Inobe Port, Inobe, Wadomari;
KAUM–I. 122355, 28.8 mm SL, off Okidomari Port, Shimojiro, China; KAUM–I. 122685, 84.3 mm SL,
Wadomari Port, Wadomari; KAUM–I. 122686, 88.8 mm SL, Wadomari Port, Wadomari; KAUM–I. 122687,
86.5 mm SL, Wadomari Port, Wadomari; KAUM–I. 122720, 23.3 mm SL, off Yakomo, China; KAUM–
I. 90424, 28.5 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 90598, 89.0 mm SL, Okidomari Port,
Shimojiro, China; KAUM–I. 90599, 86.7 mm SL, Okidomari Port, Shimojiro, China; KAUM–I. 121900,
105.5 mm SL, Okidomari Port, Shimojiro, China; KAUM–I. 121972, 92.9 mm SL, Inobe Port, Inobe,
Wadomari; KAUM–I. 121989, 88.0 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 121990, 95.8 mm SL,
Inobe Port, Inobe, Wadomari; KAUM–I. 122265, 39.1 mm SL, west of Taminazaki, Tamina, China; KAUM–
I. 122267, 41.8 mm SL, west of Taminazaki, Tamina, China; KAUM–I. 122354, 44.2 mm SL, off OkidomariPort, Shimojiro, China; KAUM–I. 123582, 39.4 mm SL, off China.
Pristicon rufus (Randall & Fraser 1999)[Jpn name: Akahireishimochi]
Yoshida (2018b): KAUM–I. 90179, 26.5 mm SL, Okidomari Port, Shimojiro, China. Yoshida (2019b):
KAUM–I. 90179, 26.5 mm SL, Okidomari Port, Shimojiro, China. This study: KAUM–I. 90179, 26.5 mm
SL, Okidomari Port, Shimojiro, China.
Pristicon trimaculatus (Cuvier 1828)[Jpn name: Misujiakahireishimochi]
KAUM–I. 90175, 18.5 mm SL, Okidomari Port, Shimojiro, China.
FAMILY MALACANTHIDAE
Hoplolatilus cuniculus Randall & Dooley 1974
[Jpn name: Okinawasangoamadai]
Underwater photograph (Pl. 4-E).
Hoplolatilus starcki Randall & Dooley 1974
[Jpn name: N/A]
Underwater photograph (Pl. 4-F).
Remarks: No specimens of this species have never been collected from Japanese waters. In Okinoerabu
Island, two individuals have been observed at depths of 30–40 m off Cape Tamina for recent four years.

Marine and freshwater fishes of Okinoerabu Island

39

Malacanthus brevirostris Guichenot 1848
[Jpn name: Yaseamadai]
KAUM–I. 122657, 204.1 mm SL, off Naikina Port, Nagamine, Wadomari; KAUM–I. 122658, 206.0 mm SL,
off Naikina Port, Nagamine, Wadomari.
Malacanthus latovittatus (Lacepède 1801)[Jpn name: Kitsuneamadai]
Morishita (2018b): KAUM–I. 90141, 75.7 mm SL, Okidomari Port, Shimojiro, China. Morishita (2019b):
KAUM–I. 90141, 75.7 mm SL, Okidomari Port, Shimojiro, China. This study: KAUM–I. 90141, 75.7 mm
SL, Okidomari Port, Shimojiro, China.
FAMILY CORYPHAENIDAE
Coryphaena hippurus Linnaeus 1758
[Jpn name: Shiira]Morishita (2019c): KAUM–I. 122662, 881.2 mm SL, off Naikina Port, Nagamine, Wadomari. This study:
KAUM–I. 122208, 634.0 mm SL, southeast of Okinoerabu Island; KAUM–I. 122662, 881.2 mm SL, off
Naikina Port, Nagamine, Wadomari.
FAMILY CARANGIDAE
Alectis ciliaris (Bloch 1787)[Jpn name: Itohikiaji]
KAUM–I. 90585, 52.8 mm SL, Okidomari Port, Shimojiro, China.
Carangoides orthogrammus (Jordan & Gilbert 1882)[Jpn name: Nan-yokaiwari]
KAUM–I. 122663, 344.6 mm SL, off Naikina Port, Nagamine, Wadomari.
Caranx lugubris Poey 1860
[Jpn name: Kappore]
KAUM–I. 122678, 363.3 mm SL, off Wadomari Port, Wadomari.
Caranx melampygus Cuvier 1833
[Jpn name: Kasumiaji]
Kimura (2018): KAUM–I. 101616, 483.0 mm SL, Kunigami, Wadomari. Kimura (2019b): KAUM–I.
101616, 483.0 mm SL, Kunigami, Wadomari. This study: KAUM–I. 90437, 105.0 mm SL, Inobe Port,
Inobe, Wadomari; KAUM–I. 101616, 483.0 mm SL, Kunigami, Wadomari; KAUM–I. 101617, 463.0 mm
SL, Kunigami, Wadomari; KAUM–I. 101629, 180.0 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I.
122219, 101.0 mm SL, Inobe Port, Inobe, Wadomari.
Caranx sexfasciatus Quoy & Gaimard 1825
[Jpn name: Gingameaji]
KAUM–I. 2014, 58.5 mm SL, off Okinoerabu Island.
Decapterus macarellus (Cuvier 1833)[Jpn name: Kusayamoro]
KAUM–I. 123497, 282.4 mm SL, off Okidomari Port, Shimojiro, China; KAUM–I. 123525, 408.0 mm SL,
40 km east of Okinoerabu Island.

40

H. Motomura and K. Uehara

Elagatis bipinnulata (Quoy & Gaimard 1825)[Jpn name: Tsumuburi]
KAUM–I. 90551, 354.5 mm SL, south of Okinoerabu Island.
Seriola rivoliana Valenciennes 1833
[Jpn name: Hirenagakampachi]
Kimura (2019b): KAUM–I. 122660, 572.0 mm SL, off Naikina Port, Nagamine, Wadomari. This study:
KAUM–I. 122660, 572.0 mm SL, off Naikina Port, Nagamine, Wadomari.
FAMILY LUTJANIDAE
Aphareus furca (Lacepède 1801)[Jpn name: Ishifuedai]
Hata (2018): KAUM–I. 90355, 282.2 mm SL, off Okidomari Port, Shimojiro, China. Hata (2019): KAUM–
I. 90355, 282.2 mm SL, off Okidomari Port, Shimojiro, China. This study: KAUM–I. 90355, 282.2 mm SL,
off Okidomari Port, Shimojiro, China; KAUM–I. 122802, 264.5 mm SL, Kunigami, Wadomari; KAUM–
I. 123496, 243.2 mm SL, off Okidomari Port, Shimojiro, China; KAUM–I. 123502, 258.1 mm SL, off
Okidomari Port, Shimojiro, China.
Aphareus rutilans Cuvier 1830
[Jpn name: Oguchiishichibiki]
KAUM–I. 122103, 611.0 mm SL, southeast of Okinoerabu Island.
Etelis carbunculus Cuvier 1828
[Jpn name: Hachijoakamutsu]
KAUM–I. 121750, 248.7 mm SL, off China.
Lutjanus bengalensis (Bloch 1790)[Jpn name: Bengarufuedai]
KAUM–I. 122055, 167.3 mm SL, off Wadomari; KAUM–I. 122056, 189.7 mm SL, off Wadomari; KAUM–
I. 122057, 160.8 mm SL, off Wadomari.
Lutjanus fulviflamma (Forsskål 1775)[Jpn name: Nisekurohoshifuedai]
KAUM–I. 90247, 20.5 mm SL, Sumiyoshi Beach, Sumiyoshi, China; KAUM–I. 90274, 34.8 mm SL,
Sumiyoshi Beach, Sumiyoshi, China; KAUM–I. 90275, 22.0 mm SL, Sumiyoshi Beach, Sumiyoshi, China.
Lutjanus fulvus (Forster 1801)[Jpn name: Okifuedai]
KAUM–I. 2060, 71.9 mm SL, off Okinoerabu Island; KAUM–I. 14209, 52.4 mm SL, off Okinoerabu Island;
KAUM–I. 14210, 54.1 mm SL, off Okinoerabu Island; KAUM–I. 14211, 52.6 mm SL, off Okinoerabu
Island; KAUM–I. 14212, 50.0 mm SL, off Okinoerabu Island; KAUM–I. 14213, 52.7 mm SL, off
Okinoerabu Island; KAUM–I. 14214, 52.0 mm SL, off Okinoerabu Island; KAUM–I. 14215, 44.5 mm SL,
off Okinoerabu Island; KAUM–I. 14216, 45.3 mm SL, off Okinoerabu Island; KAUM–I. 14217, 46.6 mm
SL, off Okinoerabu Island; KAUM–I. 14218, 40.0 mm SL, off Okinoerabu Island; KAUM–I. 14219, 38.1mm SL, off Okinoerabu Island; KAUM–I. 14220, 32.9 mm SL, off Okinoerabu Island; KAUM–I. 14221,
40.0 mm SL, off Okinoerabu Island; KAUM–I. 14222, 35.1 mm SL, off Okinoerabu Island; KAUM–I.
14223, 36.1 mm SL, off Okinoerabu Island; KAUM–I. 14224, 33.5 mm SL, off Okinoerabu Island; KAUM–
I. 14225, 32.0 mm SL, off Okinoerabu Island; KAUM–I. 14226, 32.7 mm SL, off Okinoerabu Island;Marine and freshwater fishes of Okinoerabu Island

41

KAUM–I. 14227, 26.7 mm SL, off Okinoerabu Island; KAUM–I. 14228, 29.5 mm SL, off Okinoerabu
Island; KAUM–I. 14229, 34.1 mm SL, off Okinoerabu Island; KAUM–I. 14230, 30.4 mm SL, off
Okinoerabu Island; KAUM–I. 14231, 27.6 mm SL, off Okinoerabu Island; KAUM–I. 14232, 25.3 mm SL,
off Okinoerabu Island; KAUM–I. 101630, 186.0 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 121952,
90.1 mm SL, Inobe Port, Inobe, Wadomari.
Lutjanus gibbus (Forsskål 1775)[Jpn name: Himefuedai]
Hata (2018): KAUM–I. 90182, 49.5 mm SL, Yakomo Beach, Yakomo, China. Hata (2019): KAUM–
I. 90182, 49.5 mm SL, Yakomo Beach, Yakomo, China. This study: KAUM–I. 1990, 58.2 mm SL, off
Okinoerabu Island; KAUM–I. 14237, 32.8 mm SL, off Okinoerabu Island; KAUM–I. 90182, 49.5 mm SL,
Yakomo Beach, Yakomo, China; KAUM–I. 121953, 96.0 mm SL, Inobe Port, Inobe, Wadomari; KAUM–
I. 121977, 96.5 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 121978, 99.3 mm SL, Inobe Port, Inobe,
Wadomari.
Lutjanus kasmira (Forsskål 1775)[Jpn name: Yosujifuedai]
KAUM–I. 2142, 59.5 mm SL, off Okinoerabu Island; KAUM–I. 20844, 43.5 mm SL, off Okinoerabu Island;
KAUM–I. 101624, 128.0 mm SL, Fucha, Kunigami, Wadomari; KAUM–I. 121856, 83.5 mm SL, Inobe Port,
Inobe, Wadomari.
Lutjanus monostigma (Cuvier 1828)[Jpn name: Ittenfuedai]
KAUM–I. 1993, 67.6 mm SL, off Okinoerabu Island; KAUM–I. 2773, 3 specimens, 22.5–50.2 mm SL, off
Okinoerabu Island; KAUM–I. 90558, 49.0 mm SL, Okidomari Port, Shimojiro, China; KAUM–I. 121853,
314.7 mm SL, China Port, Serikaku, China.
Macolor niger (Forsskål 1775)[Jpn name: Madaratarumi]
Hata (2018): KAUM–I. 90767, 15.8 mm SL, Naikina Port, Nagamine, Wadomari. Hata (2019): KAUM–
I. 90767, 15.8 mm SL, Naikina Port, Nagamine, Wadomari. This study: KAUM–I. 90767, 15.8 mm SL,
Naikina Port, Nagamine, Wadomari.
Paracaesio caerulea (Katayama 1934)[Jpn name: Aodai]
KAUM–I. 122180, 249.7 mm SL, Sumiyoshi, China; KAUM–I. 123536, 360.1 mm SL, Sumiyoshi, China.
Paracaesio xanthura (Bleeker 1869)[Jpn name: Umeiro]
KAUM–I. 121746, 242.2 mm SL, off Wadomari; KAUM–I. 122061, 269.5 mm SL, southeast of Okinoerabu
Island.
Pristipomoides argyrogrammicus (Valenciennes 1832)[Jpn name: Hanafuedai]
KAUM–I. 53948, 232.5 mm SL, off Okinoerabu Island; KAUM–I. 121744, 280.9 mm SL, off China;
KAUM–I. 121745, 223.6 mm SL, off China.

42

H. Motomura and K. Uehara

Pristipomoides filamentosus (Valenciennes 1830)[Jpn name: Ohime]
KAUM–I. 122063, 385.0 mm SL, southeast of Okinoerabu Island.
Pristipomoides flavipinnis Shinohara 1963
[Jpn name: Kimmehimedai]
KAUM–I. 122181, 307.2 mm SL, Sumiyoshi, China; KAUM–I. 123527, 207.7 mm SL, Sumiyoshi, China.
Pristipomoides sieboldii (Bleeker 1855)[Jpn name: Himedai]
Hata (2018): KAUM–I. 90727, 225.0 mm SL, Kunigami, Wadomari. This study: KAUM–I. 90727, 225.0mm SL, Kunigami, Wadomari; KAUM–I. 123526, 311.4 mm SL, Sumiyoshi, China.
FAMILY CAESIONIDAE
Caesio teres Seale 1906
[Jpn name: Umeiromodoki]
KAUM–I. 122300, 203.8 mm SL, off Okidomari Port, Shimojiro, China.
Pterocaesio marri Schultz 1953
[Jpn name: Nisetakasago]
KAUM–I. 90577, 217.9 mm SL, Okidomari Port, Shimojiro, China.
FAMILY GERREIDAE
Gerres oyena (Forsskål 1775)[Jpn name: Minamikurosagi]
KAUM–I. 122235, 171.8 mm SL, Wadomari Port, Wadomari.
FAMILY HAEMULIDAE
Plectorhinchus gibbosus (Lacepède 1802)[Jpn name: Kurokoshodai]
KAUM–I. 121916, 15.0 mm SL, Inobe Port, Inobe, Wadomari.
Plectorhinchus lessonii (Cuvier 1830)[Jpn name: Hiregurokoshodai]
KAUM–I. 90674, 23.1 mm SL, Wanjo Beach, Azefu, Wadomari.
Plectorhinchus picus (Cuvier 1828)[Jpn name: Ajiakoshodai]
KAUM–I. 122066, 421.4 mm SL, Kunigami, Wadomari.
FAMILY NEMIPTERIDAE
Pentapodus aureofasciatus Russell 2001
[Jpn name: Yakushimakitsuneuo]
Underwater photograph (Pl. 4-G).
Scolopsis lineata Quoy & Gaimard 1824
[Jpn name: Yokoshimatamagashira]Nakamura (2019): KAUM–I. 121890, 161.1 mm SL, Okidomari Port, Shimojiro, China. This study:
KAUM–I. 121890, 161.1 mm SL, Okidomari Port, Shimojiro, China.

Marine and freshwater fishes of Okinoerabu Island

43

FAMILY LETHRINIDAE
Gnathodentex aureolineatus (Lacepède 1802)[Jpn name: Nokogiridai]
KAUM–I. 90587, 133.8 mm SL, Okidomari Port, Shimojiro, China; KAUM–I. 121971, 137.3 mm SL, Inobe
Port, Inobe, Wadomari; KAUM–I. 122202, 128.1 mm SL, Wadomari Port, Wadomari; KAUM–I. 122214,
216.1 mm SL, Wadomari Port, Wadomari; KAUM–I. 122688, 104.2 mm SL, Wadomari Port, Wadomari.
Gymnocranius euanus (Günther 1879)[Jpn name: Shirodai]
KAUM–I. 122664, 292.6 mm SL, off Naikina Port, Nagamine, Wadomari.
Gymnocranius griseus (Temminck & Schlegel 1843)[Jpn name: Meichidai]
Muto (2019): KAUM–I. 122200, 356.3 mm SL, Sumiyoshi, China. This study: KAUM–I. 122200, 356.3mm SL, Sumiyoshi, China.
Lethrinus atkinsoni Seale 1910
[Jpn name: Isofuefuki]
KAUM–I. 121857, 94.4 mm SL, Inobe Port, Inobe, Wadomari;Lethrinus genivittatus Valenciennes 1830
[Jpn name: Itofuefuki]
Muto (2019): KAUM–I. 121863, 80.5 mm SL, Inobe Port, Inobe, Wadomari. This study: KAUM–I. 121855,
92.7 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 121863, 80.5 mm SL, Inobe Port, Inobe, Wadomari.
Lethrinus harak (Forsskål 1775)[Jpn name: Matofuefuki]
Muto (2018): KAUM–I. 90121, 235.2 mm SL, Inobe Port, Inobe, Wadomari. Muto (2019): KAUM–I. 90121,
235.2 mm SL, Inobe Port, Inobe, Wadomari. This study: KAUM–I. 90121, 235.2 mm SL, Inobe Port, Inobe,
Wadomari.
Lethrinus miniatus (Forster 1801)[Jpn name: Amamifuefuki]
Muto (2019): KAUM–I. 123530, 384.0 mm SL, off Okinoerabu Island. This study: KAUM–I. 123530, 384.0mm SL, off Okinoerabu Island.
Lethrinus obsoletus (Forsskål 1775)[Jpn name: Tateshimafuefuki]
Muto (2018): KAUM–I. 90734, 125.0 mm SL, Naikina Port, Nagamine, Wadomari. Muto (2019): KAUM–
I. 90734, 125.0 mm SL, Naikina Port, Nagamine, Wadomari. This study: KAUM–I. 90734, 125.0 mm SL,
Naikina Port, Nagamine, Wadomari.
Lethrinus rubrioperculatus Sato 1978
[Jpn name: Hoakakuchibi]
Muto (2018): KPM-NI 29273, 145.2 mm SL, off Okinoerabu Island. Muto (2019): KPM-NI 29273, 145.2mm SL, off Okinoerabu Island. This study: KAUM–I. 121854, 145.5 mm SL, Inobe Port, Inobe, Wadomari.

44

H. Motomura and K. Uehara

Lethrinus xanthochilus Klunzinger 1870
[Jpn name: Muneakakuchibi]
KAUM–I. 123495, 241.6 mm SL, off Okidomari Port, Shimojiro, China.
FAMILY MULLIDAE
Mulloidichthys flavolineatus flavolineatus (Lacepède 1801)[Jpn name: Montsukiakahimeji]
Yoshigou et al. (2005a, as Mulloidichthys flavolineatus): HMNH 7791, 2 specimens, Hanjo, Furusato,
Wadomari. Yoshigou (2014): Okinoerabu Island. This study: KAUM–I. 90339, 103.3 mm SL, off OkidomariPort, Shimojiro, China.
Mulloidichthys pflugeri (Steindachner 1900)[Jpn name: Ryukyuakahimeji]
Bandai (2019): KAUM–I. 122315, 278.0 mm SL, off Okidomari Port, Shimojiro, China. This study:
KAUM–I. 122315, 278.0 mm SL, off Okidomari Port, Shimojiro, China.
Mulloidichthys vanicolensis (Valenciennes 1831)[Jpn name: Akahimeji]
KAUM–I. 121969, 168.0 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 122213, 201.0 mm SL,
Wadomari Port, Wadomari.
Parupeneus barberinus (Lacepède 1801)[Jpn name: Osujihimeji]
Bandai (2018): KAUM–I. 90155, 192.7 mm SL, Okidomari Port, Shimojiro, China. Bandai (2019): KAUM–
I. 90155, 192.7 mm SL, Okidomari Port, Shimojiro, China. This study: KAUM–I. 2045, 63.0 mm SL, off
Okinoerabu Island; KAUM–I. 14233, 46.4 mm SL, off Okinoerabu Island; KAUM–I. 90155, 192.7 mm SL,
Okidomari Port, Shimojiro, China.
Parupeneus ciliatus (Lacepède 1802)[Jpn name: Horaihimeji]
KAUM–I. 1897, 11 specimens, 49.4–60.7 mm SL, off Okinoerabu Island; KAUM–I. 90580, 210.5 mm SL,
Okidomari Port, Shimojiro, China; KAUM–I. 121882, 114.0 mm SL, Okidomari Port, Shimojiro, China;
KAUM–I. 122062, 285.0 mm SL, Kunigami, Wadomari.
Parupeneus cyclostomus (Lacepède 1801)[Jpn name: Marukuchihimeji]
Bandai (2019): KAUM–I. 121858, 114.1 mm SL, Inobe Port, Inobe, Wadomari. This study: KAUM–I.
121858, 114.1 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 121896, 132.3 mm SL, Okidomari Port,
Shimojiro, China; KAUM–I. 122659, 118.7 mm SL, off Naikina Port, Nagamine, Wadomari.
Parupeneus heptacanthus (Lacepède 1802)[Jpn name: Takasagohimeji]
KAUM–I. 123531, 273.8 mm SL, Hirase, Tamashiro, Wadomari; KAUM–I. 123532, 286.0 mm SL, Hirase,
Tamashiro, Wadomari.
Parupeneus multifasciatus (Quoy & Gaimard 1825)[Jpn name: Ojisan]
KAUM–I. 90099, 138.1 mm SL, Wadomari Port, Wadomari; KAUM–I. 90119, 191.6 mm SL, Inobe Port,

Marine and freshwater fishes of Okinoerabu Island

45

Inobe, Wadomari; KAUM–I. 90413, 124.0 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 90414, 94.5 mm
SL, Inobe Port, Inobe, Wadomari; KAUM–I. 90415, 86.3 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I.
121768, 169.7 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 121869, 109.4 mm SL, Inobe Port, Inobe,
Wadomari; KAUM–I. 121870, 115.4 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 121871, 100.0 mm
SL, Inobe Port, Inobe, Wadomari; KAUM–I. 121872, 80.1 mm SL, Inobe Port, Inobe, Wadomari; KAUM–
I. 121877, 80.3 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 121878, 91.6 mm SL, Inobe Port, Inobe,
Wadomari; KAUM–I. 121879, 76.5 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 122237, 80.6 mm
SL, Wadomari Port, Wadomari; KAUM–I. 122673, 163.5 mm SL, off Naikina Port, Nagamine, Wadomari;
KAUM–I. 122674, 175.5 mm SL, off Naikina Port, Nagamine, Wadomari; KAUM–I. 123507, 165.4 mm
SL, off Okidomari Port, Shimojiro, China; KAUM–I. 123522, 170.6 mm SL, off Okidomari Port, Shimojiro,
China.
Parupeneus pleurostigma (Bennett 1831)[Jpn name: Ryukyuhimeji]
Bandai (2019): KAUM–I. 123501, 108.1 mm SL, off Okidomari Port, Shimojiro, China. This study:
KAUM–I. 90412, 150.4 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 121859, 101.1 mm SL, Inobe
Port, Inobe, Wadomari; KAUM–I. 123501, 108.1 mm SL, off Okidomari Port, Shimojiro, China; KAUM–I.
123508, 143.0 mm SL, off Okidomari Port, Shimojiro, China.
FAMILY PEMPHERIDAE
Pempheris adusta Bleeker 1877
[Jpn name: Ryukyuhatampo]
KAUM–I. 2047, 62.7 mm SL, off Okinoerabu Island; KAUM–I. 87246, 87.7 mm SL, off Okinoerabu
Island; KAUM–I. 121974, 118.0 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 122209, 139.3 mm SL,
Wadomari Port, Wadomari; KAUM–I. 122210, 117.2 mm SL, Wadomari Port, Wadomari; KAUM–I. 122211,
114.4 mm SL, Wadomari Port, Wadomari.
Pempheris oualensis Cuvier 1831
[Jpn name: Yumehatampo]
Koeda (2018a): KAUM–I. 90157, 188.3 mm SL, Okidomari Port, Shimojiro, China; KAUM–I. 90158, 140.0
mm SL, Okidomari Port, Shimojiro, China. Koeda (2019a): KAUM–I. 90157, 188.3 mm SL, OkidomariPort, Shimojiro, China; KAUM–I. 90158, 140.0 mm SL, Okidomari Port, Shimojiro, China. This study:
KAUM–I. 90157, 188.3 mm SL, Okidomari Port, Shimojiro, China; KAUM–I. 90158, 140.0 mm SL,
Okidomari Port, Shimojiro, China.
Pempheris schwenkii Bleeker 1855
[Jpn name: Minamihatampo]
Yoshigou et al. (2005a): HMNH 7988, Nishibaru, Wadomari; HMNH 7998, Nishibaru, Wadomari; HMNH
7999, Nishibaru, Wadomari.
FAMILY CHAETODONTIDAE
Chaetodon argentatus Smith & Radcliffe 1911
[Jpn name: Kagamichochouo]
Fujiwara (2018): KAUM–I. 90396, 37.8 mm SL, Inobe Port, Inobe, Wadomari. Fujiwara (2019): KAUM–I.
90396, 37.8 mm SL, Inobe Port, Inobe, Wadomari. This study: KAUM–I. 1925, 2 specimens, 37.5–38.5 mm
SL, off Okinoerabu Island; KAUM–I. 90396, 37.8 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 132180,
49.7 mm SL, west of Taminazaki, Tamina, China; KUN-P 48888, 60.8 mm SL, off Masana, China.

46

H. Motomura and K. Uehara

Chaetodon auriga Forsskål 1775
[Jpn name: Togechochouo]
Yoshigou et al. (2005a): HMNH 7839, Shinjo, China. Fujiwara (2018): KAUM–I. 90701, 18.0 mm SL,
Inobe Port, Inobe, Wadomari. Fujiwara (2019): KAUM–I. 90701, 18.0 mm SL, Inobe Port, Inobe, Wadomari.
This study: KAUM–I. 2099, 26.6 mm SL, off Okinoerabu Island; KAUM–I. 19927, 23.5 mm SL, off
Okinoerabu Island; KAUM–I. 19928, 24.3 mm SL, off Okinoerabu Island; KAUM–I. 19929, 22.8 mm SL,
off Okinoerabu Island; KAUM–I. 19930, 21.1 mm SL, off Okinoerabu Island; KAUM–I. 90701, 18.0 mm
SL, Inobe Port, Inobe, Wadomari.
Chaetodon auripes Jordan & Snyder 1901
[Jpn name: Chochouo]
KAUM–I. 2130, 44.1 mm SL, off Okinoerabu Island; KAUM–I. 3948, 41.3 mm SL, off Okinoerabu Island;
KAUM–I. 3949, 39.9 mm SL, off Okinoerabu Island; KAUM–I. 3950, 26.0 mm SL, off Okinoerabu Island;
KAUM–I. 90583, 137.1 mm SL, Okidomari Port, Shimojiro, China; KAUM–I. 90716, 14.0 mm SL, Inobe
Port, Inobe, Wadomari; KAUM–I. 121968, 141.3 mm SL, Inobe Port, Inobe, Wadomari.
Chaetodon bennetti Cuvier 1831
[Jpn name: Umizukichochouo]
Fujiwara (2018): KAUM–I. 90394, 140.1 mm SL, Inobe Port, Inobe, Wadomari. Fujiwara (2019): KAUM–I.
90394, 140.1 mm SL, Inobe Port, Inobe, Wadomari. This study: KAUM–I. 90394, 140.1 mm SL, Inobe Port,
Inobe, Wadomari.
Chaetodon citrinellus Cuvier 1831
[Jpn name: Gomachochouo]
Yoshigou et al. (2005a): HMNH 7966, Nishibaru, Wadomari. Fujiwara (2019): KAUM–I. 122146, 25.0mm SL, Beach Rock, Tokudoki, China. This study: KAUM–I. 36329, 90.6 mm SL, off Okinoerabu Island;
KAUM–I. 121894, 93.3 mm SL, Okidomari Port, Shimojiro, China; KAUM–I. 122035, 28.3 mm SL,
Masana, China; KAUM–I. 122146, 25.0 mm SL, Beach Rock, Tokudoki, China; KAUM–I. 122633, 25.1mm SL, China Port, Serikaku, China; KAUM–I. 132185, 30.0 mm SL, west of Taminazaki, Tamina, China;
KUN-P 48955, 91.0 mm SL, Ujiji Beach, Ashikiyora, China; KUN-P 49082, 54.2 mm SL, China Port,
Serikaku, China.
Chaetodon ephippium Cuvier 1831
[Jpn name: Segurochochouo]
Fujiwara (2018): KAUM–I. 90153, 155.0 mm SL, Okidomari Port, Shimojiro, China. Fujiwara (2019):
KAUM–I. 90153, 155.0 mm SL, Okidomari Port, Shimojiro, China. This study: KAUM–I. 90153, 155.0 mm
SL, Okidomari Port, Shimojiro, China.
Chaetodon guentheri Ahl 1923
[Jpn name: Kokutenkatagi]
Underwater photograph (Pl. 4-H).
Chaetodon kleinii Bloch 1790
[Jpn name: Mizorechochouo]
Fujiwara (2018): KAUM–I. 90156, 69.5 mm SL, Okidomari Port, Shimojiro, China. Fujiwara (2019):
KAUM–I. 90156, 69.5 mm SL, Okidomari Port, Shimojiro, China. This study: KAUM–I. 90156, 69.5 mm
SL, Okidomari Port, Shimojiro, China.

Marine and freshwater fishes of Okinoerabu Island

47

Chaetodon lunula (Lacepède 1802)[Jpn name: Chohan]
Yoshigou et al. (2005a): HMNH 7967, Nishibaru, Wadomari; HMNH 7968, Nishibaru, Wadomari; HMNH
7969, Nishibaru, Wadomari; HMNH 7970, Nishibaru, Wadomari. This study: KAUM–I. 13798, 41.9 mm SL,
off Okinoerabu Island; KAUM–I. 13799, 40.5 mm SL, off Okinoerabu Island; KAUM–I. 13800, 28.9 mm
SL, off Okinoerabu Island; KAUM–I. 13801, 26.6 mm SL, off Okinoerabu Island; KAUM–I. 13802, 24.9mm SL, off Okinoerabu Island; KAUM–I. 13803, 24.0 mm SL, off Okinoerabu Island; KAUM–I. 36326,
34.9 mm SL, off Okinoerabu Island; KAUM–I. 36327, 59.7 mm SL, off Okinoerabu Island; KAUM–I.
36328, 70.7 mm SL, off Okinoerabu Island; KAUM–I. 90446, 139.6 mm SL, Inobe Port, Inobe, Wadomari.
Chaetodon lunulatus Quoy & Gaimard 1825
[Jpn name: Misujichochouo]
Fujiwara (2018): KAUM–I. 90743, 21.9 mm SL, Naikina Port, Nagamine, Wadomari. Fujiwara (2019):
KAUM–I. 90743, 21.9 mm SL, Naikina Port, Nagamine, Wadomari. This study: KAUM–I. 90393, 75.5 mm
SL, Inobe Port, Inobe, Wadomari; KAUM–I. 90743, 21.9 mm SL, Naikina Port, Nagamine, Wadomari.
Chaetodon melannotus Bloch & Schneider 1801
[Jpn name: Akebonochochouo]
KAUM–I. 90582, 99.9 mm SL, Okidomari Port, Shimojiro, China.
Chaetodon mertensii Cuvier 1831
[Jpn name: Beniochochouo]
Underwater photograph (Pl. 4-I).
Chaetodon meyeri Bloch & Schneider 1801
[Jpn name: Ougichochouo]
Underwater photograph (Pl. 5-A).
Chaetodon ornatissimus Cuvier 1831
[Jpn name: Hanagurochochouo]
KAUM–I. 90098, 151.7 mm SL, Okidomari Port, Shimojiro, China.
Chaetodon rafflesii Anonymous [Bennett] 1830
[Jpn name: Amichochouo]
Fujiwara (2018): KAUM–I. 90716, 14.0 mm SL, Inobe Port, Inobe, Wadomari. Fujiwara (2019): KAUM–
I. 90716, 14.0 mm SL, Inobe Port, Inobe, Wadomari. This study: KAUM–I. 90716, 14.0 mm SL, Inobe Port,
Inobe, Wadomari.
Chaetodon reticulatus Cuvier 1831
[Jpn name: Hakutenkatagi]
Underwater photograph (Pl. 5-B).
Chaetodon semeion Bleeker 1855
[Jpn name: Remonchochouo]
Underwater photograph (Pl. 5-C).
Remarks: The underwater photograph represents the first record of C. semeion from the Amami Islands andthe northernmost record for the species. This species is rarely observed in Okinoerabu Island around depthsof 5 m.

48

H. Motomura and K. Uehara

Chaetodon trifascialis Quoy & Gaimard 1825
[Jpn name: Yarikatagi]
Fujiwara (2018): KAUM–I. 90614, 29.5 mm SL, Naikina Port, Nagamine, Wadomari; KAUM–I. 90615,
19.3 mm SL, Naikina Port, Nagamine, Wadomari. Fujiwara (2019): KAUM–I. 90614, 29.5 mm SL, Naikina
Port, Nagamine, Wadomari; KAUM–I. 90615, 19.3 mm SL, Naikina Port, Nagamine, Wadomari. This study:
KAUM–I. 90614, 29.5 mm SL, Naikina Port, Nagamine, Wadomari; KAUM–I. 90615, 19.3 mm SL, Naikina
Port, Nagamine, Wadomari; KAUM–I. 90616, 18.8 mm SL, Naikina Port, Nagamine, Wadomari; KAUM–I.
90647, 29.0 mm SL, Naikina Port, Nagamine, Wadomari.
Chaetodon unimaculatus Bloch 1787
[Jpn name: Ittenchochouo]
Fujiwara (2018): KAUM–I. 90131, 40.0 mm SL, Okidomari Port, Shimojiro, China. Fujiwara (2019):
KAUM–I. 90131, 40.0 mm SL, Okidomari Port, Shimojiro, China. This study: KAUM–I. 90131, 40.0 mm
SL, Okidomari Port, Shimojiro, China; KAUM–I. 122726, 139.7 mm SL, off Yakomo, China.
Chaetodon vagabundus Linnaeus 1758
[Jpn name: Furaichochouo]
Yoshigou et al. (2005a): HMNH 7965, Nishibaru, Wadomari. Fujiwara (2018): KAUM–I. 90044, 22.8 mm
SL, Kasaishi Beach, Kibiru, Wadomari; KAUM–I. 90184, 20.5 mm SL, Yakomo Beach, Yakomo, China.
Fujiwara (2019): KAUM–I. 90044, 22.8 mm SL, Kasaishi Beach, Kibiru, Wadomari; KAUM–I. 90184,
20.5 mm SL, Yakomo Beach, Yakomo, China. This study: KAUM–I. 90038, 102.3 mm SL, OkidomariPort, Shimojiro, China; KAUM–I. 90044, 22.8 mm SL, Kasaishi Beach, Kibiru, Wadomari; KAUM–
I. 90183, 18.9 mm SL, Yakomo Beach, Yakomo, China; KAUM–I. 90184, 20.5 mm SL, Yakomo Beach,
Yakomo, China; KAUM–I. 90505, 31.4 mm SL, Ujiji Beach, Ashikiyora, China; KAUM–I. 121895, 112.3mm SL, Okidomari Port, Shimojiro, China; KAUM–I. 122069, 35.7 mm SL, Yakomo Beach, Yakomo,
China; KAUM–I. 122750, 36.3 mm SL, Kasaishi Beach, Kibiru, Wadomari; KUN-P 48869, 117.0 mm SL,
Okidomari Port, Shimojiro, China.
Chaetodon xanthurus Bleeker 1857
[Jpn name: Amimechochouo]
Underwater photograph (Pl. 5-D).
Forcipiger flavissimus Jordan & McGregor 1898
[Jpn name: Fueyakkodai]
Fujiwara (2018): KAUM–I. 90395, 98.0 mm SL, Inobe Port, Inobe, Wadomari. Fujiwara (2019): KAUM–
I. 90395, 98.0 mm SL, Inobe Port, Inobe, Wadomari. This study: KAUM–I. 90395, 98.0 mm SL, Inobe Port,
Inobe, Wadomari; KUN-P 48887, 104.2 mm SL, off Masana, China.
Hemitaurichthys polylepis (Bleeker 1857)[Jpn name: Kasumichochouo]
Underwater photograph (Pl. 5-E).
Heniochus chrysostomus Cuvier 1831
[Jpn name: Minamihatatatedai]
Fujiwara (2018): KAUM–I. 90445, 123.6 mm SL, Inobe Port, Inobe, Wadomari. Fujiwara (2019): KAUM–
I. 90445, 123.6 mm SL, Inobe Port, Inobe, Wadomari. This study: KAUM–I. 2026, 40.4 mm SL, off
Okinoerabu Island; KAUM–I. 90445, 123.6 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 101636, 63.0mm SL, Kasaishi Beach, Kibiru, Wadomari.

Marine and freshwater fishes of Okinoerabu Island

49

Heniochus singularius Smith & Radcliffe 1911
[Jpn name: Shimahatatatedai]
Fujiwara (2018): KAUM–I. 90493, 219.2 mm SL, Inobe Port, Inobe, Wadomari. Fujiwara (2019): KAUM–I.
90493, 219.2 mm SL, Inobe Port, Inobe, Wadomari. This study: KAUM–I. 90493, 219.2 mm SL, Inobe Port,
Inobe, Wadomari; KAUM–I. 122727, 224.0 mm SL, Yaja, China.
FAMILY POMACANTHIDAE
Apolemichthys trimaculatus (Cuvier 1831)[Jpn name: Shiten-yakko]
KAUM–I. 123402, 107.6 mm SL, north of Taminazaki, Tamina, China; KUN-P 48947, 150.9 mm SL, westof Taminazaki, Tamina, China.
Centropyge ferrugata Randall & Burgess 1972
[Jpn name: Akaharayakko]
KAUM–I. 122108, 42.2 mm SL, Sumiyoshi Port, Sumiyoshi, China.
Centropyge fisheri (Snyder 1904)[Jpn name: Chairoyakko]
Underwater photograph (Pl. 5-F).
Centropyge heraldi Woods & Schultz 1953
[Jpn name: Herarudokoganeyakko]
Wibowo (2018): KAUM–I. 90447, 32.1 mm SL, Inobe Port, Inobe, Wadomari. Wibowo (2019): KAUM–I.
90447, 32.1 mm SL, Inobe Port, Inobe, Wadomari. This study: KAUM–I. 90447, 32.1 mm SL, Inobe Port,
Inobe, Wadomari.
Centropyge shepardi Randall & Yasuda 1979
[Jpn name: Daidaiyakko]
Underwater photograph (Pl. 5-G).
Remarks: The underwater photograph represents the second record of this species from the Ryukyu Islands[first Ryukyu specimen recorded by Koeda & Motomura (2016) from Amami-oshima island].
Centropyge tibicen (Cuvier 1831)[Jpn name: Abrayakko]
KAUM–I. 132186, 27.6 mm SL, west of Taminazaki, Tamina, China.
Centropyge venusta (Yasuda & Tominaga 1969)[Jpn name: Sumireyakko]
Wibowo (2018, as Paracentropyge venusta): KAUM–I. 90127, 65.7 mm SL, Okidomari Port, Shimojiro,
China. Wibowo (2019, as Paracentropyge venusta): KAUM–I. 90127, 65.7 mm SL, Okidomari Port,
Shimojiro, China. This study: KAUM–I. 90127, 65.7 mm SL, Okidomari Port, Shimojiro, China; KAUM–I.
122034, 25.4 mm SL, Masana, China; KAUM–I. 122604, 63.8 mm SL, off Yakomo, China.
Remarks: This species has sometimes been treated as a member of Paracentropyge.
Centropyge vrolikii (Bleeker 1853)[Jpn name: Namerayakko]
Wibowo (2018): KAUM–I. 90056, 23.0 mm SL, Okidomari Port, Shimojiro, China. Wibowo (2019):
KAUM–I. 90056, 23.0 mm SL, Okidomari Port, Shimojiro, China. This study: KAUM–I. 90056, 23.0

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mm SL, Okidomari Port, Shimojiro, China; KAUM–I. 90384, 18.2 mm SL, Inobe Port, Inobe, Wadomari;
KAUM–I. 122607, 24.5 mm SL, off Yakomo, China; KAUM–I. 122705, 42.8 mm SL, off Yakomo, China;
KAUM–I. 123564, 21.8 mm SL, Yaja, China.
Pomacanthus imperator (Bloch 1787)[Jpn name: Tatejimakinchakudai]
Yoshigou et al. (2005a): HMNH 7898, Okidomari, China.
Pomacanthus semicirculatus (Cuvier 1831)[Jpn name: Sazanamiyakko]
Wibowo (2018): KAUM–I. 90519, 90.9 mm SL, Ujiji Beach, Ashikiyora, China. Wibowo (2019): KAUM–
I. 90519, 90.9 mm SL, Ujiji Beach, Ashikiyora, China. This study: KAUM–I. 36325, 46.4 mm SL, off
Okinoerabu Island; KAUM–I. 90519, 90.9 mm SL, Ujiji Beach, Ashikiyora, China; KUN-P 48903, 186.0
mm SL, Beach Rock, Tokudoki, China; KUN-P 49104, 28.9 mm SL, Kasaishi Beach, Kibiru, Wadomari.
Genicanthus melanospilos (Bleeker 1857)[Jpn name: Yaitoyakko]
Underwater photograph (Pl. 5-H).
FAMILY CIRRHITIDAE
Cirrhitichthys aprinus (Cuvier 1829)[Jpn name: Minamigombe]
Koeda (2018b): KAUM–I. 90129, 53.4 mm SL, Okidomari Port, Shimojiro, China. Koeda (2019b): KAUM–
I. 90129, 53.4 mm SL, Okidomari Port, Shimojiro, China. This study: KAUM–I. 90129, 53.4 mm SL,
Okidomari Port, Shimojiro, China.
Cirrhitichthys falco Randall 1963
[Jpn name: Sarasagombe]
Koeda (2018b): KAUM–I. 90082, 36.9 mm SL, Okidomari Port, Shimojiro, China. Koeda (2019b): KAUM–
I. 90082, 36.9 mm SL, Okidomari Port, Shimojiro, China. This study: KAUM–I. 90012, 32.5 mm SL,
Okidomari Port, Shimojiro, China; KAUM–I. 90082, 36.9 mm SL, Okidomari Port, Shimojiro, China;
KAUM–I. 122253, 43.3 mm SL, west of Taminazaki, Tamina, China; KUN-P 48937, 30.7 mm SL, west of
Taminazaki, Tamina, China.
Cirrhitichthys oxycephalus (Bleeker 1855)[Jpn name: Himegombe]
Koeda (2018b): KAUM–I. 90130, 66.1 mm SL, Okidomari Port, Shimojiro, China. Koeda (2019b): KAUM–
I. 90130, 66.1 mm SL, Okidomari Port, Shimojiro, China. This study: KAUM–I. 90130, 66.1 mm SL,
Okidomari Port, Shimojiro, China; KAUM–I. 123550, 37.3 mm SL, Yaja, China; KAUM–I. 123592, 59.3mm SL, off China.
Cirrhitus pinnulatus pinnulatus (Forster 1801)[Jpn name: Isogombe]
Koeda (2018b): KAUM–I. 90733, 192.7 mm SL, Hanzaki, Wadomari; KAUM–I. 101628, 114.3 mm
SL, Inobe Port, Inobe, Wadomari. Koeda (2019b): KAUM–I. 90733, 192.7 mm SL, Hanzaki, Wadomari;
KAUM–I. 101628, 114.3 mm SL, Inobe Port, Inobe, Wadomari. This study: KAUM–I. 90733, 192.7 mm
SL, Hanzaki, Wadomari; KAUM–I. 101628, 114.3 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 122226,
125.9 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 122227, 128.8 mm SL, Inobe Port, Inobe, Wadomari;
KAUM–I. 122800, 104.7 mm SL, Kunigami, Wadomari; KUN-P 49087, 149.6 mm SL, off Yakomo, China.

Marine and freshwater fishes of Okinoerabu Island

51

Oxycirrhites typus Bleeker 1857
[Jpn name: Kudagombe]
Underwater photograph (Pl. 5-I).
Paracirrhites arcatus (Cuvier 1829)[Jpn name: Meganegombe]
KAUM–I. 122004, 24.8 mm SL, Masana, China; KAUM–I. 122304, 54.1 mm SL, west of Taminazaki,
Tamina, China; KAUM–I. 122333, 26.0 mm SL, north of Taminazaki, Tamina, China; KAUM–I. 122616,
28.3 mm SL, off Yakomo, China; KAUM–I. 122699, 30.0 mm SL, off Yakomo, China; KUN-P 48889, 48.6mm SL, off Masana, China; KUN-P 48936, 33.7 mm SL, west of Taminazaki, Tamina, China; KUN-P 49021,
22.0 mm SL, Yaja, China.
Paracirrhites forsteri (Schneider 1801)[Jpn name: Hoshigombe]
KAUM–I. 90630, 81.6 mm SL, Naikina Port, Nagamine, Wadomari; KAUM–I. 122305, 34.6 mm SL, westof Taminazaki, Tamina, China; KAUM–I. 122618, 22.3 mm SL, off Yakomo, China.
FAMILY CICHLIDAE
Oreochromis niloticus (Linnaeus 1758)[Jpn name: Nairuteirapia]
Yoshigou et al. (2005a): HMNH 7989, Wadomari; HMNH 7990, Wadomari; HMNH 7991, Wadomari.
Hatooka & Yagishita (2013): Okinoerabu Island. Yoshigou (2014): Okinoerabu Island. Nakajima & Kano
(2014): Amata River, Amata, China.
Tilapia zillii (Gervais 1848)[Jpn name: Jiruteirapia]
Yoshigou et al. (2005a): HMNH 7807, Amata River, Amata, China. Hatooka & Yagishita (2013): Okinoerabu
Island. Yoshigou (2014): Okinoerabu Island. Uejo (2018b): KAUM–I. 6607, 45.1 mm SL, Amata River,
Amata, China. Uejo (2019b): KAUM–I. 122093, 36.0 mm SL, Oku River, Wadomari. This study: KAUM–I.
6607, 45.1 mm SL, Amata River, Amata, China; KAUM–I. 6608, 40.1 mm SL, Amata River, Amata, China;
KAUM–I. 6609, 42.2 mm SL, Amata River, Amata, China; KAUM–I. 6610, 35.5 mm SL, Amata River,
Amata, China; KAUM–I. 6611, 40.6 mm SL, Amata River, Amata, China; KAUM–I. 6612, 41.7 mm SL,
Amata River, Amata, China; KAUM–I. 6613, 43.5 mm SL, Amata River, Amata, China; KAUM–I. 122093,
36.0 mm SL, Oku River, Wadomari.
FAMILY POMACENTRIDAE
Abudefduf caudobimaculatus Okada & Ikeda 1939
[Jpn name: Shiritensuzumedai]
Yoshigou et al. (2005a, as Abudefduf sp.): HMNH 7944, Okidomari, China.
Abudefduf notatus (Day 1870)[Jpn name: Isosuzumedai]
Iwatsubo (2018): KAUM–I. 90007, 111.5 mm SL, Inobe Port, Inobe, Wadomari. Iwatsubo (2019a): KAUM–
I. 90007, 111.5 mm SL, Inobe Port, Inobe, Wadomari. This study: KAUM–I. 90007, 111.5 mm SL, Inobe
Port, Inobe, Wadomari; KUN-P 49149, 14.0 mm SL, Kasaishi Beach, Kibiru, Wadomari.
Abudefduf septemfasciatus (Cuvier 1830)[Jpn name: Shichisensuzumedai]
KAUM–I. 90700, 19.2 mm SL, Inobe Port, Inobe, Wadomari.

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H. Motomura and K. Uehara

Abudefduf sexfasciatus (Lacepède 1801)[Jpn name: Rokusensuzumedai]
Iwatsubo (2018): KAUM–I. 90699, 18.5 mm SL, Inobe Port, Inobe, Wadomari. Iwatsubo (2019a): KAUM–
I. 90699, 18.5 mm SL, Inobe Port, Inobe, Wadomari. This study: KAUM–I. 90102, 70.0 mm SL, WadomariPort, Wadomari; KAUM–I. 90696, 22.2 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 90697, 20.4 mm
SL, Inobe Port, Inobe, Wadomari; KAUM–I. 90698, 21.0 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I.
90699, 18.5 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 101607, 90.7 mm SL, Kunigami, Wadomari;
KAUM–I. 101608, 102.7 mm SL, Kunigami, Wadomari; KAUM–I. 101609, 99.4 mm SL, Kunigami,
Wadomari; KAUM–I. 101610, 117.5 mm SL, Kunigami, Wadomari; KAUM–I. 101631, 91.6 mm SL,
Kasaishi Beach, Kibiru, Wadomari; KAUM–I. 101632, 97.3 mm SL, Kasaishi Beach, Kibiru, Wadomari;
KAUM–I. 122236, 66.6 mm SL, Wadomari Port, Wadomari; KAUM–I. 122801, 113.6 mm SL, Kunigami,
Wadomari.
Abudefduf sordidus (Forsskål 1775)[Jpn name: Shimasuzumedai]
Yoshigou et al. (2005a): HMNH 7793, Hanjo, Furusato, Wadomari; HMNH 7866, Naikina, Nagamine,
Wadomari; HMNH 7867, Naikina, Nagamine, Wadomari. Yoshigou (2014): Okinoerabu Island. This study:
KAUM–I. 36609, 59.0 mm SL, off Okinoerabu Island; KAUM–I. 36610, 55.8 mm SL, off Okinoerabu
Island; KAUM–I. 36611, 60.8 mm SL, off Okinoerabu Island; KAUM–I. 36612, 57.7 mm SL, off
Okinoerabu Island; KAUM–I. 36613, 66.3 mm SL, off Okinoerabu Island; KAUM–I. 36614, 61.8 mm SL,
off Okinoerabu Island; KAUM–I. 36615, 52.2 mm SL, off Okinoerabu Island; KAUM–I. 36616, 55.3 mm
SL, off Okinoerabu Island; KAUM–I. 36617, 59.6 mm SL, off Okinoerabu Island; KAUM–I. 36618, 59.3mm SL, off Okinoerabu Island; KAUM–I. 36619, 58.8 mm SL, off Okinoerabu Island; KAUM–I. 36620,
65.3 mm SL, off Okinoerabu Island; KAUM–I. 90006, 120.4 mm SL, Inobe Port, Inobe, Wadomari; KAUM–
I. 90229, 18.6 mm SL, Sumiyoshi Beach, Sumiyoshi, China; KAUM–I. 90230, 20.5 mm SL, Sumiyoshi
Beach, Sumiyoshi, China; KAUM–I. 90231, 19.4 mm SL, Sumiyoshi Beach, Sumiyoshi, China; KAUM–
I. 90232, 10.9 mm SL, Sumiyoshi Beach, Sumiyoshi, China; KAUM–I. 90233, 13.8 mm SL, Sumiyoshi
Beach, Sumiyoshi, China; KAUM–I. 90234, 9.2 mm SL, Sumiyoshi Beach, Sumiyoshi, China; KAUM–
I. 90283, 15.0 mm SL, Sumiyoshi Beach, Sumiyoshi, China; KAUM–I. 90284, 15.2 mm SL, Sumiyoshi
Beach, Sumiyoshi, China; KAUM–I. 90285, 15.3 mm SL, Sumiyoshi Beach, Sumiyoshi, China; KAUM–I.
90286, 12.3 mm SL, Sumiyoshi Beach, Sumiyoshi, China; KAUM–I. 90287, 11.7 mm SL, Sumiyoshi Beach,
Sumiyoshi, China; KAUM–I. 90288, 10.8 mm SL, Sumiyoshi Beach, Sumiyoshi, China; KAUM–I. 122221,
148.4 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 122395, 29.4 mm SL, Ujiji Beach, Ashikiyora,
China.
Abudefduf vaigiensis (Quoy & Gaimard 1825)[Jpn name: Oyabitcha]
Yoshigou et al. (2005a): HMNH 7841, Shinjo, China; HMNH 7842, Shinjo, China; HMNH 7897, Okidomari,China; HMNH 7925, Okidomari, China; HMNH 7926, Okidomari, China; HMNH 7927, Okidomari, China;
HMNH 7974, Nishibaru, Wadomari. This study: KAUM–I. 36621, 79.4 mm SL, off Okinoerabu Island;
KAUM–I. 36622, 70.4 mm SL, off Okinoerabu Island; KAUM–I. 36623, 69.0 mm SL, off Okinoerabu
Island; KAUM–I. 36624, 73.1 mm SL, off Okinoerabu Island; KAUM–I. 36625, 67.3 mm SL, off
Okinoerabu Island; KAUM–I. 36626, 66.5 mm SL, off Okinoerabu Island; KAUM–I. 36627, 63.7 mm SL,
off Okinoerabu Island; KAUM–I. 36628, 54.3 mm SL, off Okinoerabu Island; KAUM–I. 36629, 52.3 mm
SL, off Okinoerabu Island; KAUM–I. 36630, 49.0 mm SL, off Okinoerabu Island; KAUM–I. 36631, 48.2mm SL, off Okinoerabu Island; KAUM–I. 36632, 45.4 mm SL, off Okinoerabu Island; KAUM–I. 36633,
48.1 mm SL, off Okinoerabu Island; KAUM–I. 36634, 44.9 mm SL, off Okinoerabu Island; KAUM–
I. 36635, 46.5 mm SL, off Okinoerabu Island; KAUM–I. 36636, 48.3 mm SL, off Okinoerabu Island;Marine and freshwater fishes of Okinoerabu Island

53

KAUM–I. 36637, 41.1 mm SL, off Okinoerabu Island; KAUM–I. 36638, 35.4 mm SL, off Okinoerabu
Island; KAUM–I. 36639, 36.8 mm SL, off Okinoerabu Island; KAUM–I. 122383, 28.7 mm SL, Ujiji Beach,
Ashikiyora, China; KAUM–I. 122755, 32.1 mm SL, Kasaishi Beach, Kibiru, Wadomari; KAUM–I. 152373,
14.1 mm SL, Okidomari Seaside Park, Shimojiro, China; KUN-P 49043, 33.7 mm SL, China Port, Serikaku,
China; KUN-P 49148, 27.5 mm SL, Kasaishi Beach, Kibiru, Wadomari.
Amblyglyphidodon aureus (Cuvier 1830)[Jpn name: Yamabukisuzumedai]
Underwater photograph (Pl. 6-A).
Amblyglyphidodon ternatensis (Bleeker 1853)[Jpn name: Nisekurakaosuzumedai]
Underwater photograph (Pl. 6-B).
Amphiprion clarkii (Bennett 1830)[Jpn name: Kumanomi]
KAUM–I. 26759, 84.2 mm SL, off Okinoerabu Island; KAUM–I. 26793, 75.5 mm SL, off Okinoerabu
Island; KAUM–I. 90054, 30.8 mm SL, Okidomari Port, Shimojiro, China; KAUM–I. 90387, 66.6 mm SL,
Inobe Port, Inobe, Wadomari; KAUM–I. 90388, 64.9 mm SL, Inobe Port, Inobe, Wadomari.
Amphiprion frenatus Brevoort 1856
[Jpn name: Hamakumanomi]
Iwatsubo (2018): KAUM–I. 90429, 29.2 mm SL, Inobe Port, Inobe, Wadomari. Iwatsubo (2019a): KAUM–
I. 90429, 29.2 mm SL, Inobe Port, Inobe, Wadomari. This study: KAUM–I. 90429, 29.2 mm SL, Inobe Port,
Inobe, Wadomari.
Amphiprion ocellaris Cuvier 1830
[Jpn name: Kakurekumanomi]
Underwater photograph (Pl. 6-C).
Amphiprion perideraion Bleeker 1855
[Jpn name: Hanabirakumanomi]
KAUM–I. 90423, 66.6 mm SL, Inobe Port, Inobe, Wadomari.
Amphiprion sandaracinos Allen 1972
[Jpn name: Sejirokumanomi]
Underwater photograph (Pl. 6-D).
Chromis albomaculata Kamohara 1960
[Jpn name: Shiroboshisuzumedai]
Underwater photograph (Pl. 6-E).
Chromis atripes Fowler & Bean 1928
[Jpn name: Hiregurosuzumedai]
KAUM–I. 122730, 19.3 mm SL, Yaja, China; KAUM–I. 123554, 21.4 mm SL, Yaja, China; KAUM–I.
123587, 38.5 mm SL, off China.

54

H. Motomura and K. Uehara

Chromis chrysura (Bliss 1883)[Jpn name: Amamisuzumedai]
KAUM–I. 90728, 99.1 mm SL, Naikina Port, Nagamine, Wadomari.
Chromis lepidolepis Bleeker 1877
[Jpn name: Sasasuzumedai]
KAUM–I. 122111, 51.2 mm SL, Sumiyoshi Port, Sumiyoshi, China; KAUM–I. 122719, 13.6 mm SL, off
Yakomo, China.
Chromis leucura Gilbert 1905
[Jpn name: Fukamisuzumedai]
Underwater photograph (Pl. 6-F).
Remarks: Underwater photograph represents the first record of this species from the Amami Islands.
Chromis margaritifera Fowler 1946
[Jpn name: Shikokusuzumedai]
Iwatsubo (2018): KAUM–I. 90048, 23.7 mm SL, Kasaishi Beach, Kibiru, Wadomari; KAUM–I. 90052,
17.6 mm SL, Okidomari Port, Shimojiro, China. Iwatsubo (2019a): KAUM–I. 90048, 23.7 mm SL, Kasaishi
Beach, Kibiru, Wadomari; KAUM–I. 90052, 17.6 mm SL, Okidomari Port, Shimojiro, China. This study:
KAUM–I. 26753, 101.3 mm SL, off Okinoerabu Island; KAUM–I. 90013, 17.2 mm SL, Okidomari Port,
Shimojiro, China; KAUM–I. 90048, 23.7 mm SL, Kasaishi Beach, Kibiru, Wadomari; KAUM–I. 90049,
25.6 mm SL, Kasaishi Beach, Kibiru, Wadomari; KAUM–I. 90052, 17.6 mm SL, Okidomari Port, Shimojiro,
China; KAUM–I. 90173, 16.9 mm SL, Okidomari Port, Shimojiro, China; KAUM–I. 121998, 18.0 mm SL,
Masana, China; KAUM–I. 122248, 17.4 mm SL, west of Taminazaki, Tamina, China; KAUM–I. 122328,
20.5 mm SL, north of Taminazaki, Tamina, China; KAUM–I. 122606, 18.0 mm SL, off Yakomo, China;
KAUM–I. 123541, 23.1 mm SL, Yaja, China; KAUM–I. 123551, 41.6 mm SL, Yaja, China; KAUM–I.
123573, 32.4 mm SL, Yaja, China; KUN-P 48892, 28.6 mm SL, off Masana, China; KAUM–I. 132173, 18.0mm SL, west of Taminazaki, Tamina, China; KUN-P 49110, 19.8 mm SL, Yakomo Beach, Yakomo, China.
Chromis ovatiformes Fowler 1946
[Jpn name: Marusuzumedai]
Iwatsubo (2018): KAUM–I. 90174, 16.2 mm SL, Okidomari Port, Shimojiro, China. Iwatsubo (2019a):
KAUM–I. 90174, 16.2 mm SL, Okidomari Port, Shimojiro, China. This study: KAUM–I. 90174, 16.2 mm
SL, Okidomari Port, Shimojiro, China; KAUM–I. 122349, 57.1 mm SL, off Okidomari Port, Shimojiro,
China; KAUM–I. 122735, 14.6 mm SL, Yaja, China.
Chromis vanderbilti (Fowler 1941)[Jpn name: Himesuzumedai]
Iwatsubo (2018): KAUM–I. 90051, 18.2 mm SL, Okidomari Port, Shimojiro, China; KAUM–I. 90053, 27.7mm SL, Okidomari Port, Shimojiro, China. Iwatsubo (2019a): KAUM–I. 90051, 18.2 mm SL, OkidomariPort, Shimojiro, China; KAUM–I. 122009, 37.7 mm SL, Masana, China. This study: KAUM–I. 90051, 18.2mm SL, Okidomari Port, Shimojiro, China; KAUM–I. 90053, 27.7 mm SL, Okidomari Port, Shimojiro,
China; KAUM–I. 122009, 37.7 mm SL, Masana, China; KAUM–I. 122014, 19.2 mm SL, Masana, China;
KAUM–I. 122360, 36.2 mm SL, off Okidomari Port, Shimojiro, China; KAUM–I. 123557, 34.4 mm SL,
Yaja, China; KAUM–I. 123567, 33.0 mm SL, Yaja, China; KAUM–I. 123585, 21.9 mm SL, off China;
KUN-P 49008, 36.0 mm SL, Yaja, China; KUN-P 49009, 21.5 mm SL, Yaja, China.

Marine and freshwater fishes of Okinoerabu Island

55

Chromis xouthos Allen & Erdmann 2005
[Jpn name: Himawarisuzumedai]
Underwater photograph (Pl. 6-G).
Chromis yamakawai Iwatsubo & Motomura 2013
[Jpn name: Kihoshisuzumedai]
KAUM–I. 26762, 74.3 mm SL, off Okinoerabu Island; KAUM–I. 90341, 84.0 mm SL, off Okidomari Port,
Shimojiro, China; KAUM–I. 90342, 78.5 mm SL, off Okidomari Port, Shimojiro, China; KAUM–I. 90343,
80.1 mm SL, off Okidomari Port, Shimojiro, China; KAUM–I. 90345, 90.3 mm SL, off Okidomari Port,
Shimojiro, China; KAUM–I. 90346, 89.0 mm SL, off Okidomari Port, Shimojiro, China; KAUM–I. 90347,
89.3 mm SL, off Okidomari Port, Shimojiro, China; KAUM–I. 90348, 83.4 mm SL, off Okidomari Port,
Shimojiro, China; KAUM–I. 90349, 78.5 mm SL, off Okidomari Port, Shimojiro, China; KAUM–I. 90350,
71.4 mm SL, off Okidomari Port, Shimojiro, China; KAUM–I. 90351, 82.5 mm SL, off Okidomari Port,
Shimojiro, China; KAUM–I. 90352, 83.5 mm SL, off Okidomari Port, Shimojiro, China; KAUM–I. 90353,
84.2 mm SL, off Okidomari Port, Shimojiro, China; KAUM–I. 90354, 76.7 mm SL, off Okidomari Port,
Shimojiro, China; KAUM–I. 90357, 72.6 mm SL, off Okidomari Port, Shimojiro, China; KAUM–I. 90358,
70.2 mm SL, off Okidomari Port, Shimojiro, China; KAUM–I. 122808, 48.8 mm SL, Sumiyoshi Port,
Sumiyoshi, China.
Chrysiptera biocellata (Quoy & Gaimard 1825)[Jpn name: Sujibuchisuzumedai]
Iwatsubo (2018): KAUM–I. 90702, 20.2 mm SL, Inobe Port, Inobe, Wadomari. Iwatsubo (2019a): KAUM–I.
90702, 20.2 mm SL, Inobe Port, Inobe, Wadomari. This study: KAUM–I. 90522, 18.5 mm SL, Ujiji Beach,
Ashikiyora, China; KAUM–I. 90702, 20.2 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 122099, 28.5mm SL, Yakomo Beach, Yakomo, China.
Chrysiptera cyanea (Quoy & Gaimard 1825)[Jpn name: Rurisuzumedai]
KAUM–I. 90008, 129.5 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 90085, 34.1 mm SL, Inobe Port,
Inobe, Wadomari; KAUM–I. 90086, 21.4 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 90087, 11.4 mm
SL, Kasaishi Beach, Kibiru, Wadomari; KAUM–I. 90547, 35.8 mm SL, Ujiji Beach, Ashikiyora, China;
KAUM–I. 90548, 36.4 mm SL, Ujiji Beach, Ashikiyora, China; KAUM–I. 90665, 54.6 mm SL, Wanjo
Beach, Azefu, Wadomari; KAUM–I. 90779, 15.1 mm SL, Hanzaki, Wadomari; KAUM–I. 90780, 11.1 mm
SL, Hanzaki, Wadomari; KAUM–I. 121899, 43.1 mm SL, Okidomari Port, Shimojiro, China; KAUM–I.
122082, 26.0 mm SL, Yakomo Beach, Yakomo, China; KAUM–I. 122177, 12.5 mm SL, Beach Rock,Tokudoki, China; KAUM–I. 122770, 40.3 mm SL, Kasaishi Beach, Kibiru, Wadomari; KAUM–I. 123419,
26.5 mm SL, Ujiji Beach, Ashikiyora, China; KAUM–I. 123473, 27.1 mm SL, Kasaishi Beach, Kibiru,
Wadomari; KAUM–I. 132300, 14.4 mm SL, Okidomari Seaside Park, Shimojiro, China; KAUM–I. 132301,
11.8 mm SL, Okidomari Seaside Park, Shimojiro, China; KAUM–I. 132302, 13.0 mm SL, Okidomari
Seaside Park, Shimojiro, China; KAUM–I. 132303, 12.3 mm SL, Okidomari Seaside Park, Shimojiro, China;
KAUM–I. 132304, 11.6 mm SL, Okidomari Seaside Park, Shimojiro, China; KAUM–I. 132305, 14.4 mm
SL, Okidomari Seaside Park, Shimojiro, China; KAUM–I. 132306, 11.2 mm SL, Okidomari Seaside Park,
Shimojiro, China; KAUM–I. 132307, 12.1 mm SL, Okidomari Seaside Park, Shimojiro, China; KAUM–I.
132308, 10.9 mm SL, Okidomari Seaside Park, Shimojiro, China; KAUM–I. 132309, 11.3 mm SL,
Okidomari Seaside Park, Shimojiro, China; KAUM–I. 132310, 11.9 mm SL, Okidomari Seaside Park,
Shimojiro, China; KAUM–I. 132311, 11.3 mm SL, Okidomari Seaside Park, Shimojiro, China; KAUM–I.
132312, 11.0 mm SL, Okidomari Seaside Park, Shimojiro, China; KAUM–I. 132313, 11.7 mm SL,
Okidomari Seaside Park, Shimojiro, China; KAUM–I. 132314, 11.6 mm SL, Okidomari Seaside Park,

56

H. Motomura and K. Uehara

Shimojiro, China; KAUM–I. 132315, 11.9 mm SL, Okidomari Seaside Park, Shimojiro, China; KAUM–I.
132316, 11.1 mm SL, Okidomari Seaside Park, Shimojiro, China; KAUM–I. 132317, 10.3 mm SL,
Okidomari Seaside Park, Shimojiro, China; KAUM–I. 132318, 11.5 mm SL, Okidomari Seaside Park,
Shimojiro, China; KAUM–I. 132319, 11.3 mm SL, Okidomari Seaside Park, Shimojiro, China; KAUM–I.
132320, 11.5 mm SL, Okidomari Seaside Park, Shimojiro, China; KAUM–I. 132321, 11.4 mm SL,
Okidomari Seaside Park, Shimojiro, China; KAUM–I. 132322, 10.5 mm SL, Okidomari Seaside Park,
Shimojiro, China; KAUM–I. 132323, 11.0 mm SL, Okidomari Seaside Park, Shimojiro, China; KAUM–I.
132324, 10.6 mm SL, Okidomari Seaside Park, Shimojiro, China; KAUM–I. 132325, 11.0 mm SL,
Okidomari Seaside Park, Shimojiro, China; KAUM–I. 132326, 10.5 mm SL, Okidomari Seaside Park,
Shimojiro, China; KAUM–I. 132327, 10.9 mm SL, Okidomari Seaside Park, Shimojiro, China; KAUM–I.
132328, 8.2 mm SL, Okidomari Seaside Park, Shimojiro, China; KAUM–I. 132329, 1.01 mm SL,
Okidomari Seaside Park, Shimojiro, China; KAUM–I. 132330, 11.1 mm SL, Okidomari Seaside Park,
Shimojiro, China; KAUM–I. 132331, 10.8 mm SL, Okidomari Seaside Park, Shimojiro, China; KAUM–I.
132332, 11.6 mm SL, Okidomari Seaside Park, Shimojiro, China; KAUM–I. 132333, 11.7 mm SL,
Okidomari Seaside Park, Shimojiro, China; KAUM–I. 132334, 10.9 mm SL, Okidomari Seaside Park,
Shimojiro, China; KAUM–I. 132335, 10.3 mm SL, Okidomari Seaside Park, Shimojiro, China; KAUM–I.
152374, 11.0 mm SL, Okidomari Seaside Park, Shimojiro, China; KAUM–I. 152375, 11.7 mm SL,
Okidomari Seaside Park, Shimojiro, China; KAUM–I. 152376, 11.6 mm SL, Okidomari Seaside Park,
Shimojiro, China; KAUM–I. 152377, 10.6 mm SL, Okidomari Seaside Park, Shimojiro, China; KAUM–I.
152378, 11.1 mm SL, Okidomari Seaside Park, Shimojiro, China; KAUM–I. 152379, 10.6 mm SL,
Okidomari Seaside Park, Shimojiro, China; KAUM–I. 152380, 8.3 mm SL, Okidomari Seaside Park,
Shimojiro, China; KAUM–I. 152381, 8.4 mm SL, Okidomari Seaside Park, Shimojiro, China; KAUM–I.
152382, 11.6 mm SL, Okidomari Seaside Park, Shimojiro, China; KAUM–I. 152383, 10.9 mm SL,
Okidomari Seaside Park, Shimojiro, China; KAUM–I. 152384, 10.5 mm SL, Okidomari Seaside Park,
Shimojiro, China; KAUM–I. 152385, 10.4 mm SL, Okidomari Seaside Park, Shimojiro, China; KAUM–I.
152386, 10.6 mm SL, Okidomari Seaside Park, Shimojiro, China; KAUM–I. 152387, 10.7 mm SL,
Okidomari Seaside Park, Shimojiro, China; KAUM–I. 152388, 11.5 mm SL, Okidomari Seaside Park,
Shimojiro, China; KAUM–I. 152389, 11.6 mm SL, Okidomari Seaside Park, Shimojiro, China; KAUM–I.
152390, 9.5 mm SL, Okidomari Seaside Park, Shimojiro, China; KAUM–I. 152391, 10.3 mm SL,
Okidomari Seaside Park, Shimojiro, China; KAUM–I. 152392, 10.4 mm SL, Okidomari Seaside Park,
Shimojiro, China; KAUM–I. 152393, 10.8 mm SL, Okidomari Seaside Park, Shimojiro, China; KAUM–I.
152394, 9.1 mm SL, Okidomari Seaside Park, Shimojiro, China; KAUM–I. 152395, 11.0 mm SL, Okidomari
Seaside Park, Shimojiro, China; KAUM–I. 152396, 11.0 mm SL, Okidomari Seaside Park, Shimojiro, China;
KAUM–I. 152397, 10.6 mm SL, Okidomari Seaside Park, Shimojiro, China; KAUM–I. 152398, 8.6 mm SL,
Okidomari Seaside Park, Shimojiro, China; KAUM–I. 152399, 10.6 mm SL, Okidomari Seaside Park,
Shimojiro, China; KAUM–I. 152400, 8.5 mm SL, Okidomari Seaside Park, Shimojiro, China; KAUM–I.
152401, 10.8 mm SL, Okidomari Seaside Park, Shimojiro, China; KAUM–I. 152402, 10.1 mm SL,
Okidomari Seaside Park, Shimojiro, China; KAUM–I. 152403, 7.7 mm SL, Okidomari Seaside Park,
Shimojiro, China; KAUM–I. 152404, 10.3 mm SL, Okidomari Seaside Park, Shimojiro, China; KAUM–I.
152405, 9.1 mm SL, Okidomari Seaside Park, Shimojiro, China; KAUM–I. 152406, 10.7 mm SL,
Okidomari Seaside Park, Shimojiro, China; KAUM–I. 152407, 11.4 mm SL, Okidomari Seaside Park,
Shimojiro, China; KAUM–I. 152408, 10.8 mm SL, Okidomari Seaside Park, Shimojiro, China; KAUM–I.
152409, 11.0 mm SL, Okidomari Seaside Park, Shimojiro, China; KAUM–I. 152410, 13.3 mm SL,
Okidomari Seaside Park, Shimojiro, China; KAUM–I. 152411, 11 mm SL, Okidomari Seaside Park,
Shimojiro, China; KAUM–I. 152412, 12.2 mm SL, Okidomari Seaside Park, Shimojiro, China; KAUM–I.
152413, 10.8 mm SL, Okidomari Seaside Park, Shimojiro, China; KAUM–I. 152414, 10.6 mm SL,
Okidomari Seaside Park, Shimojiro, China; KAUM–I. 152415, 9.7 mm SL, Okidomari Seaside Park,
Shimojiro, China; KAUM–I. 152416, 9.6 mm SL, Okidomari Seaside Park, Shimojiro, China; KAUM–I.

Marine and freshwater fishes of Okinoerabu Island

57

152417, 13.0 mm SL, Okidomari Seaside Park, Shimojiro, China; KAUM–I. 152418, 10.2 mm SL,
Okidomari Seaside Park, Shimojiro, China; KAUM–I. 152419, 10.7 mm SL, Okidomari Seaside Park,
Shimojiro, China; KAUM–I. 152420, 11.3 mm SL, Okidomari Seaside Park, Shimojiro, China; KAUM–I.
152421, 11.6 mm SL, Okidomari Seaside Park, Shimojiro, China; KAUM–I. 152422, 11.2 mm SL,
Okidomari Seaside Park, Shimojiro, China; KAUM–I. 152423, 12.0 mm SL, Okidomari Seaside Park,
Shimojiro, China; KAUM–I. 152424, 14.4 mm SL, Okidomari Seaside Park, Shimojiro, China; KAUM–I.
152425, 10.0 mm SL, Okidomari Seaside Park, Shimojiro, China; KAUM–I. 152426, 12.2 mm SL,
Okidomari Seaside Park, Shimojiro, China; KAUM–I. 152427, 9.5 mm SL, Okidomari Seaside Park,
Shimojiro, China; KAUM–I. 152428, 11.3 mm SL, Okidomari Seaside Park, Shimojiro, China; KAUM–I.
152429, 9.4 mm SL, Okidomari Seaside Park, Shimojiro, China; KAUM–I. 152430, 10.9 mm SL,
Okidomari Seaside Park, Shimojiro, China; KAUM–I. 152431, 11.0 mm SL, Okidomari Seaside Park,
Shimojiro, China; KAUM–I. 152432, 10.3 mm SL, Okidomari Seaside Park, Shimojiro, China; KAUM–I.
152433, 10.4 mm SL, Okidomari Seaside Park, Shimojiro, China; KAUM–I. 152434, 10.3 mm SL,
Okidomari Seaside Park, Shimojiro, China; KAUM–I. 152435, 9.7 mm SL, Okidomari Seaside Park,
Shimojiro, China; KAUM–I. 152436, 10.6 mm SL, Okidomari Seaside Park, Shimojiro, China; KAUM–I.
152437, 9.8 mm SL, Okidomari Seaside Park, Shimojiro, China; KAUM–I. 152438, 11.7 mm SL, Okidomari
Seaside Park, Shimojiro, China; KAUM–I. 152439, 10.8 mm SL, Okidomari Seaside Park, Shimojiro, China;
KAUM–I. 152440, 11.5 mm SL, Okidomari Seaside Park, Shimojiro, China; KAUM–I. 152441, 11.7 mm
SL, Okidomari Seaside Park, Shimojiro, China; KAUM–I. 152442, 9.4 mm SL, Okidomari Seaside Park,
Shimojiro, China; KAUM–I. 152443, 10.1 mm SL, Okidomari Seaside Park, Shimojiro, China; KAUM–I.
152444, 8.5 mm SL, Okidomari Seaside Park, Shimojiro, China; KAUM–I. 152445, 10.6 mm SL,
Okidomari Seaside Park, Shimojiro, China; KAUM–I. 152446, 10.3 mm SL, Okidomari Seaside Park,
Shimojiro, China; KAUM–I. 152447, 12.9 mm SL, Okidomari Seaside Park, Shimojiro, China; KAUM–I.
152448, 9.5 mm SL, Okidomari Seaside Park, Shimojiro, China; KAUM–I. 152449, 10.3 mm SL,
Okidomari Seaside Park, Shimojiro, China.
Chrysiptera glauca (Cuvier 1830)[Jpn name: Nezusuzumedai]
Yoshigou et al. (2005a): HMNH 7822, 2 specimens, Shinjo, China. Iwatsubo (2018): KAUM–I. 90176, 24.4mm SL, Okidomari Port, Shimojiro, China; KAUM–I. 90554, 21.6 mm SL, Yakomo Beach, Yakomo, China.
Iwatsubo (2019a): KAUM–I. 90176, 24.4 mm SL, Okidomari Port, Shimojiro, China; KAUM–I. 90554, 21.6mm SL, Yakomo Beach, Yakomo, China. This study: KAUM–I. 2012, 75.1 mm SL, off Okinoerabu Island;
KAUM–I. 13509, 60.6 mm SL, off Okinoerabu Island; KAUM–I. 13510, 53.3 mm SL, off Okinoerabu
Island; KAUM–I. 13511, 49.1 mm SL, off Okinoerabu Island; KAUM–I. 90176, 24.4 mm SL, OkidomariPort, Shimojiro, China; KAUM–I. 90554, 21.6 mm SL, Yakomo Beach, Yakomo, China; KAUM–I. 122392,
38.3 mm SL, Ujiji Beach, Ashikiyora, China; KAUM–I. 122643, 14.0 mm SL, China Port, Serikaku, China;
KAUM–I. 122644, 12.9 mm SL, China Port, Serikaku, China; KAUM–I. 122779, 12.0 mm SL, Kasaishi
Beach, Kibiru, Wadomari; KUN-P 49006, 63.2 mm SL, Yaja, China; KUN-P 49028, 41.8 mm SL, China
Port, Serikaku, China; KUN-P 49067, 13.1 mm SL, China Port, Serikaku, China; KUN-P 49095, 11.6 mm
SL, Kasaishi Beach, Kibiru, Wadomari.
Chrysiptera leucopoma (Cuvier 1830)[Jpn name: Miyakokisensuzumedai]
Yoshigou et al. (2005a): HMNH 7868, Naikina, Nagamine, Wadomari. This study: KAUM–I. 122639,
13.8 mm SL, China Port, Serikaku, China; KAUM–I. 122640, 13.3 mm SL, China Port, Serikaku, China;
KAUM–I. 122641, 13.7 mm SL, China Port, Serikaku, China; KAUM–I. 122783, 12.1 mm SL, Kasaishi
Beach, Kibiru, Wadomari; KAUM–I. 133463, 45.1 mm SL, Beach Rock, Tokudoki, China; KAUM–
I. 133464, 47.3 mm SL, Beach Rock, Tokudoki, China; KAUM–I. 133465, 42.2 mm SL, Beach Rock,58

H. Motomura and K. Uehara

Tokudoki, China; KAUM–I. 133466, 45.2 mm SL, Beach Rock, Tokudoki, China; KAUM–I. 133467, 30.6mm SL, Beach Rock, Tokudoki, China; KAUM–I. 133468, 27.8 mm SL, Beach Rock, Tokudoki, China;
KAUM–I. 133469, 16.2 mm SL, Beach Rock, Tokudoki, China; KAUM–I. 133470, 19.4 mm SL, Beach
Rock, Tokudoki, China; KAUM–I. 133471, 15.7 mm SL, Beach Rock, Tokudoki, China; KAUM–I. 133472,
14.2 mm SL, Beach Rock, Tokudoki, China; KAUM–I. 133473, 15.8 mm SL, Beach Rock, Tokudoki, China;
KAUM–I. 133474, 16.9 mm SL, Beach Rock, Tokudoki, China.
Remarks: This species has been regarded as Chrysiptera brownriggii (Bennett 1828), but Allen et al. (2015)
recognized that it is an Indian Ocean species and Chrysiptera leucopoma (Cuvier 1830) should be applied toa western Pacific species.
Chrysiptera rex (Snyder 1909)[Jpn name: Remonsuzumedai]
Yoshigou et al. (2005a): HMNH 7975, Nishibaru, Wadomari. This study: KAUM–I. 2080, 56.8 mm SL, off
Okinoerabu Island; KAUM–I. 13631, 49.1 mm SL, off Okinoerabu Island; KAUM–I. 13632, 40.6 mm SL,
off Okinoerabu Island; KAUM–I. 13633, 39.3 mm SL, off Okinoerabu Island; KAUM–I. 13634, 43.5 mm
SL, off Okinoerabu Island; KAUM–I. 13635, 39.5 mm SL, off Okinoerabu Island; KAUM–I. 13636, 35.8mm SL, off Okinoerabu Island; KAUM–I. 13637, 37.1 mm SL, off Okinoerabu Island; KAUM–I. 13638,
53.6 mm SL, off Okinoerabu Island; KAUM–I. 13639, 42.3 mm SL, off Okinoerabu Island; KAUM–I.
13640, 47.7 mm SL, off Okinoerabu Island; KAUM–I. 13641, 49.1 mm SL, off Okinoerabu Island; KAUM–
I. 13642, 53.3 mm SL, off Okinoerabu Island; KAUM–I. 13643, 53.3 mm SL, off Okinoerabu Island;
KAUM–I. 13644, 29.3 mm SL, off Okinoerabu Island; KAUM–I. 13645, 27.9 mm SL, off Okinoerabu
Island; KAUM–I. 13646, 30.9 mm SL, off Okinoerabu Island; KAUM–I. 13647, 30.6 mm SL, off
Okinoerabu Island; KAUM–I. 13648, 29.7 mm SL, off Okinoerabu Island; KAUM–I. 13649, 32.8 mm SL,
off Okinoerabu Island; KAUM–I. 13650, 29.7 mm SL, off Okinoerabu Island; KAUM–I. 13651, 26.5 mm
SL, off Okinoerabu Island; KAUM–I. 13652, 26.8 mm SL, off Okinoerabu Island; KAUM–I. 13653, 26.9mm SL, off Okinoerabu Island; KAUM–I. 13654, 27.4 mm SL, off Okinoerabu Island; KAUM–I. 13655,
25.6 mm SL, off Okinoerabu Island; KAUM–I. 13656, 26.8 mm SL, off Okinoerabu Island; KAUM–I.
13657, 28.2 mm SL, off Okinoerabu Island; KAUM–I. 13658, 28.6 mm SL, off Okinoerabu Island; KAUM–
I. 13659, 28.4 mm SL, off Okinoerabu Island; KAUM–I. 13660, 24.5 mm SL, off Okinoerabu Island;
KAUM–I. 13661, 25.0 mm SL, off Okinoerabu Island; KAUM–I. 13662, 24.0 mm SL, off Okinoerabu
Island; KAUM–I. 13663, 26.7 mm SL, off Okinoerabu Island; KAUM–I. 13664, 22.2 mm SL, off
Okinoerabu Island; KAUM–I. 13665, 24.9 mm SL, off Okinoerabu Island; KAUM–I. 13666, 24.1 mm SL,
off Okinoerabu Island; KAUM–I. 13667, 23.9 mm SL, off Okinoerabu Island; KAUM–I. 13668, 23.4 mm
SL, off Okinoerabu Island; KAUM–I. 13669, 25.2 mm SL, off Okinoerabu Island; KAUM–I. 13670, 22.8mm SL, off Okinoerabu Island; KAUM–I. 13671, 22.7 mm SL, off Okinoerabu Island; KAUM–I. 13672,
22.2 mm SL, off Okinoerabu Island; KAUM–I. 13673, 23.2 mm SL, off Okinoerabu Island; KAUM–I.
13674, 20.9 mm SL, off Okinoerabu Island; KAUM–I. 13675, 22.3 mm SL, off Okinoerabu Island; KAUM–
I. 13676, 22.7 mm SL, off Okinoerabu Island; KAUM–I. 13677, 23.7 mm SL, off Okinoerabu Island;
KAUM–I. 13678, 21.1 mm SL, off Okinoerabu Island; KAUM–I. 13679, 22.6 mm SL, off Okinoerabu
Island; KAUM–I. 13680, 23.2 mm SL, off Okinoerabu Island; KAUM–I. 13681, 21.1 mm SL, off
Okinoerabu Island; KAUM–I. 13682, 20.2 mm SL, off Okinoerabu Island; KAUM–I. 13683, 23.3 mm SL,
off Okinoerabu Island; KAUM–I. 13684, 21.4 mm SL, off Okinoerabu Island; KAUM–I. 13685, 20.4 mm
SL, off Okinoerabu Island; KAUM–I. 13686, 18.9 mm SL, off Okinoerabu Island; KAUM–I. 13687, 21.0mm SL, off Okinoerabu Island; KAUM–I. 13688, 24.2 mm SL, off Okinoerabu Island; KAUM–I. 13689,
20.0 mm SL, off Okinoerabu Island; KAUM–I. 13690, 18.5 mm SL, off Okinoerabu Island; KAUM–I.
90050, 20.5 mm SL, Okidomari Port, Shimojiro, China; KAUM–I. 122044, 26.5 mm SL, Masana, China;
KAUM–I. 122384, 30.8 mm SL, Ujiji Beach, Ashikiyora, China; KAUM–I. 123586, 30.4 mm SL, off China;
KAUM–I. 123591, 32.9 mm SL, off China; KAUM–I. 133533, 38.4 mm SL, Beach Rock, Tokudoki, China;
KAUM–I. 133534, 35.3 mm SL, Beach Rock, Tokudoki, China.

Marine and freshwater fishes of Okinoerabu Island

59

Chrysiptera sp.
[Jpn name: N/A]
KAUM–I. 85698, 21.7 mm SL, off Okinoerabu Island; KAUM–I. 85699, 34.8 mm SL, off Okinoerabu
Island; KAUM–I. 85750, 22.3 mm SL, off Okinoerabu Island; KAUM–I. 85751, 23.7 mm SL, off
Okinoerabu Island; KAUM–I. 85752, 21.9 mm SL, off Okinoerabu Island; KAUM–I. 85753, 19.8 mm SL,
off Okinoerabu Island; KAUM–I. 85754, 24.1 mm SL, off Okinoerabu Island; KAUM–I. 85755, 22.5 mm
SL, off Okinoerabu Island; KAUM–I. 85756, 26.1 mm SL, off Okinoerabu Island; KAUM–I. 85757, 18.4mm SL, off Okinoerabu Island; KAUM–I. 85758, 13.0 mm SL, off Okinoerabu Island; KAUM–I. 85759,
14.6 mm SL, off Okinoerabu Island; KAUM–I. 85760, 22.7 mm SL, off Okinoerabu Island; KAUM–I.
85761, 18.6 mm SL, off Okinoerabu Island; KAUM–I. 85762, 14.8 mm SL, off Okinoerabu Island; KAUM–
I. 85763, 12.8 mm SL, off Okinoerabu Island; KAUM–I. 85764, 19.4 mm SL, off Okinoerabu Island;
KAUM–I. 85765, 18.0 mm SL, off Okinoerabu Island; KAUM–I. 85766, 19.4 mm SL, off Okinoerabu
Island; KAUM–I. 85767, 16.4 mm SL, off Okinoerabu Island; KAUM–I. 85768, 22.3 mm SL, off
Okinoerabu Island; KAUM–I. 85769, 17.0 mm SL, off Okinoerabu Island; KAUM–I. 85770, 24.5 mm SL,
off Okinoerabu Island; KAUM–I. 85771, 22.1 mm SL, off Okinoerabu Island; KAUM–I. 85772, 13.0 mm
SL, off Okinoerabu Island; KAUM–I. 85773, 19.4 mm SL, off Okinoerabu Island; KAUM–I. 85774, 20.9mm SL, off Okinoerabu Island; KAUM–I. 85775, 21.8 mm SL, off Okinoerabu Island; KAUM–I. 85776,
15.4 mm SL, off Okinoerabu Island; KAUM–I. 85777, 24.2 mm SL, off Okinoerabu Island; KAUM–I.
85778, 25.3 mm SL, off Okinoerabu Island; KAUM–I. 85779, 34.4 mm SL, off Okinoerabu Island; KAUM–
I. 85780, 23.2 mm SL, off Okinoerabu Island; KAUM–I. 85781, 19.9 mm SL, off Okinoerabu Island;
KAUM–I. 85782, 24.2 mm SL, off Okinoerabu Island; KAUM–I. 85783, 18.1 mm SL, off Okinoerabu
Island; KAUM–I. 85784, 23.6 mm SL, off Okinoerabu Island; KAUM–I. 85785, 20.3 mm SL, off
Okinoerabu Island; KAUM–I. 85786, 16.7 mm SL, off Okinoerabu Island; KAUM–I. 85787, 14.9 mm SL,
off Okinoerabu Island; KAUM–I. 85788, 31.3 mm SL, off Okinoerabu Island; KAUM–I. 85789, 20.2 mm
SL, off Okinoerabu Island; KAUM–I. 85790, 21.6 mm SL, off Okinoerabu Island; KAUM–I. 85791, 20.1mm SL, off Okinoerabu Island; KAUM–I. 85792, 21.1 mm SL, off Okinoerabu Island; KAUM–I. 85793,
22.6 mm SL, off Okinoerabu Island; KAUM–I. 85794, 21.3 mm SL, off Okinoerabu Island; KAUM–I.
85795, 36.7 mm SL, off Okinoerabu Island; KAUM–I. 85796, 19.4 mm SL, off Okinoerabu Island; KAUM–
I. 85797, 22.5 mm SL, off Okinoerabu Island; KAUM–I. 85798, 16.8 mm SL, off Okinoerabu Island;
KAUM–I. 85799, 23.8 mm SL, off Okinoerabu Island; KAUM–I. 85800, 15.3 mm SL, off Okinoerabu
Island; KAUM–I. 85801, 21.3 mm SL, off Okinoerabu Island; KAUM–I. 85802, 12.7 mm SL, off
Okinoerabu Island; KAUM–I. 85803, 22.3 mm SL, off Okinoerabu Island; KAUM–I. 85804, 14.5 mm SL,
off Okinoerabu Island; KAUM–I. 85805, 13.6 mm SL, off Okinoerabu Island; KAUM–I. 85806, 17.6 mm
SL, off Okinoerabu Island; KAUM–I. 85807, 15.8 mm SL, off Okinoerabu Island; KAUM–I. 85808, 14.8mm SL, off Okinoerabu Island; KAUM–I. 85809, 16.2 mm SL, off Okinoerabu Island; KAUM–I. 85810,
13.7 mm SL, off Okinoerabu Island; KAUM–I. 85811, 13.7 mm SL, off Okinoerabu Island.
Chrysiptera starcki (Allen 1973)[Jpn name: Senakirurisuzumedai]
Iwatsubo (2018): KAUM–I. 90171, 34.4 mm SL, Okidomari Port, Shimojiro, China. Iwatsubo (2019a):
KAUM–I. 90171, 34.4 mm SL, Okidomari Port, Shimojiro, China. This study: KAUM–I. 90170, 30.1 mm
SL, Okidomari Port, Shimojiro, China; KAUM–I. 90171, 34.4 mm SL, Okidomari Port, Shimojiro, China;
KAUM–I. 122704, 36.6 mm SL, off Yakomo, China.
Chrysiptera tricincta (Allen & Randall 1974)[Jpn name: Misujisuzumedai]
Underwater photograph (Pl. 6-H).

60

H. Motomura and K. Uehara

Chrysiptera unimaculata (Cuvier 1830)[Jpn name: Ichimonsuzumedai]
Iwatsubo (2018): KAUM–I. 90668, 13.0 mm SL, Wanjo Beach, Azefu, Wadomari; KAUM–I. 90705, 20.2mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 90774, 15.4 mm SL, Hanzaki, Wadomari. Iwatsubo(2019a): KAUM–I. 90668, 13.0 mm SL, Wanjo Beach, Azefu, Wadomari; KAUM–I. 90705, 20.2 mm SL,
Inobe Port, Inobe, Wadomari; KAUM–I. 90774, 15.4 mm SL, Hanzaki, Wadomari. This study: KAUM–I.
2041, 17.5 mm SL, off Okinoerabu Island; KAUM–I. 7790, 19.2 mm SL, off Okinoerabu Island; KAUM–I.
90549, 20.0 mm SL, Ujiji Beach, Ashikiyora, China; KAUM–I. 90668, 13.0 mm SL, Wanjo Beach, Azefu,
Wadomari; KAUM–I. 90705, 20.2 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 122088, 15.7 mm
SL, Yakomo Beach, Yakomo, China; KUN-P 48967, 11.5 mm SL, Ujiji Beach, Ashikiyora, China; KUN-P
49153, 12.8 mm SL, Kasaishi Beach, Kibiru, Wadomari.
Dascyllus aruanus (Linnaeus 1758)[Jpn name: Misujiryukyusuzumedai]
Iwatsubo (2018): KAUM–I. 90669, 13.5 mm SL, Wanjo Beach, Azefu, Wadomari; KAUM–I. 90671, 6.4mm SL, Wanjo Beach, Azefu, Wadomari. Iwatsubo (2019a): KAUM–I. 90669, 13.5 mm SL, Wanjo Beach,
Azefu, Wadomari; KAUM–I. 90671, 6.4 mm SL, Wanjo Beach, Azefu, Wadomari. This study: KAUM–I.
90499, 7.0 mm SL, Ujiji Beach, Ashikiyora, China; KAUM–I. 90500, 14.8 mm SL, Ujiji Beach, Ashikiyora,
China; KAUM–I. 90501, 16.8 mm SL, Ujiji Beach, Ashikiyora, China; KAUM–I. 90669, 13.5 mm SL,
Wanjo Beach, Azefu, Wadomari; KAUM–I. 90670, 9.0 mm SL, Wanjo Beach, Azefu, Wadomari; KAUM–
I. 90671, 6.4 mm SL, Wanjo Beach, Azefu, Wadomari; KAUM–I. 122756, 31.5 mm SL, Kasaishi Beach,
Kibiru, Wadomari; KAUM–I. 123457, 43.8 mm SL, Kasaishi Beach, Kibiru, Wadomari; KAUM–I. 123458,
39.5 mm SL, Kasaishi Beach, Kibiru, Wadomari; KAUM–I. 123459, 30.1 mm SL, Kasaishi Beach, Kibiru,
Wadomari.
Dascyllus reticulatus (Richardson 1846)[Jpn name: Futasujiryukyusuzumedai]
KAUM–I. 122694, 51.6 mm SL, off Yakomo, China.
Dascyllus trimaculatus (Rüppell 1829)[Jpn name: Mitsuboshikurosuzumedai]
Iwatsubo (2018): KAUM–I. 90432, 19.7 mm SL, Inobe Port, Inobe, Wadomari. Iwatsubo (2019a): KAUM–
I. 90432, 19.7 mm SL, Inobe Port, Inobe, Wadomari. This study: KAUM–I. 90063, 12.3 mm SL, OkidomariPort, Shimojiro, China; KAUM–I. 90432, 19.7 mm SL, Inobe Port, Inobe, Wadomari.
Neoglyphidodon nigroris (Cuvier 1830)[Jpn name: Hirenagasuzumedai]
Iwatsubo (2018): KAUM–I. 90430, 20.7 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 90431, 13.7 mm
SL, Inobe Port, Inobe, Wadomari. Iwatsubo (2019a): KAUM–I. 90430, 20.7 mm SL, Inobe Port, Inobe,
Wadomari; KAUM–I. 90431, 13.7 mm SL, Inobe Port, Inobe, Wadomari. This study: KAUM–I. 90430,
20.7 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 90431, 13.7 mm SL, Inobe Port, Inobe, Wadomari;
KAUM–I. 90754, 23.2 mm SL, Naikina Port, Nagamine, Wadomari; KAUM–I. 122803, 83.3 mm SL,
Kunigami, Wadomari; KUN-P 48912, 30.1 mm SL, Yakomo Beach, Yakomo, China.
Plectroglyphidodon dickii (Liénard 1839)[Jpn name: Ishigakisuzumedai]
Yoshigou et al. (2005a): HMNH 7942, Okidomari, China. Iwatsubo (2018): KAUM–I. 90761, 22.8 mm
SL, Naikina Port, Nagamine, Wadomari. Iwatsubo (2019a): KAUM–I. 90761, 22.8 mm SL, Naikina Port,

Marine and freshwater fishes of Okinoerabu Island

61

Nagamine, Wadomari. This study: KAUM–I. 90761, 22.8 mm SL, Naikina Port, Nagamine, Wadomari;
KUN-P 48934, 11.4 mm SL, west of Taminazaki, Tamina, China
Plectroglyphidodon imparipennis (Vaillant & Sauvage 1875)[Jpn name: Iwasakisuzumedai]
KAUM–I. 122609, 36.9 mm SL, off Yakomo, China; KAUM–I. 122642, 38.3 mm SL, China Port, Serikaku,
China; KUN-P 49114, 38.7 mm SL, Yakomo Beach, Yakomo, China; KAUM–I. 133554, 45.0 mm SL, Beach
Rock, Tokudoki, China; KAUM–I. 133555, 32.2 mm SL, Beach Rock, Tokudoki, China; KAUM–I. 133556,
47.6 mm SL, Beach Rock, Tokudoki, China; KAUM–I. 133557, 38.5 mm SL, Beach Rock, Tokudoki, China;
KAUM–I. 133558, 30.0 mm SL, Beach Rock, Tokudoki, China; KAUM–I. 133559, 29.4 mm SL, Beach
Rock, Tokudoki, China; KAUM–I. 133560, 27.2 mm SL, Beach Rock, Tokudoki, China; KAUM–I. 133561,
30.1 mm SL, Beach Rock, Tokudoki, China; KAUM–I. 133562, 42.2 mm SL, Beach Rock, Tokudoki, China;
KAUM–I. 133563, 42.5 mm SL, Beach Rock, Tokudoki, China; KAUM–I. 133564, 32.7 mm SL, Beach
Rock, Tokudoki, China; KAUM–I. 133565, 26.6 mm SL, Beach Rock, Tokudoki, China; KAUM–I. 133566,
28.2 mm SL, Beach Rock, Tokudoki, China; KAUM–I. 133567, 27.0 mm SL, Beach Rock, Tokudoki, China;
KAUM–I. 133568, 27.2 mm SL, Beach Rock, Tokudoki, China; KAUM–I. 133569, 17.3 mm SL, Beach
Rock, Tokudoki, China; KAUM–I. 133570, 26.4 mm SL, Beach Rock, Tokudoki, China; KAUM–I. 133571,
26.8 mm SL, Beach Rock, Tokudoki, China; KAUM–I. 133572, 26.2 mm SL, Beach Rock, Tokudoki, China;
KAUM–I. 133573, 27.4 mm SL, Beach Rock, Tokudoki, China; KAUM–I. 133574, 22.7 mm SL, Beach
Rock, Tokudoki, China; KAUM–I. 133575, 30.4 mm SL, Beach Rock, Tokudoki, China; KAUM–I. 133576,
34.8 mm SL, Beach Rock, Tokudoki, China; KAUM–I. 133577, 28.3 mm SL, Beach Rock, Tokudoki, China;
KAUM–I. 133578, 37.2 mm SL, Beach Rock, Tokudoki, China; KAUM–I. 133579, 40.0 mm SL, Beach
Rock, Tokudoki, China; KAUM–I. 133580, 34.9 mm SL, Beach Rock, Tokudoki, China; KAUM–I. 133581,
33.1 mm SL, Beach Rock, Tokudoki, China; KAUM–I. 133582, 30.5 mm SL, Beach Rock, Tokudoki, China;
KAUM–I. 133583, 26.2 mm SL, Beach Rock, Tokudoki, China; KAUM–I. 133584, 27.3 mm SL, Beach
Rock, Tokudoki, China; KAUM–I. 133585, 24.4 mm SL, Beach Rock, Tokudoki, China; KAUM–I. 133586,
23.7 mm SL, Beach Rock, Tokudoki, China; KAUM–I. 133587, 25.8 mm SL, Beach Rock, Tokudoki, China;
KAUM–I. 133588, 26.9 mm SL, Beach Rock, Tokudoki, China; KAUM–I. 133589, 21.3 mm SL, Beach
Rock, Tokudoki, China; KAUM–I. 133590, 24.1 mm SL, Beach Rock, Tokudoki, China; KAUM–I. 133591,
28.4 mm SL, Beach Rock, Tokudoki, China; KAUM–I. 133592, 26.8 mm SL, Beach Rock, Tokudoki, China;
KAUM–I. 133593, 22.4 mm SL, Beach Rock, Tokudoki, China; KAUM–I. 133594, 21.7 mm SL, Beach
Rock, Tokudoki, China; KAUM–I. 133595, 23.8 mm SL, Beach Rock, Tokudoki, China; KAUM–I. 133596,
14.6 mm SL, Beach Rock, Tokudoki, China.
Plectroglyphidodon johnstonianus Fowler & Ball 1924
[Jpn name: Rurimeishigakisuzumedai]
Iwatsubo (2019a): KAUM–I. 122733, 54.0 mm SL, Yaja, China. This study: KAUM–I. 122733, 54.0 mm
SL, Yaja, China; KAUM–I. 122734, 39.2 mm SL, Yaja, China; KAUM–I. 123562, 40.5 mm SL, Yaja, China.
Plectroglyphidodon lacrymatus (Quoy & Gaimard 1825)[Jpn name: Rurihoshisuzumedai]
KAUM–I. 90386, 25.0 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 90421, 74.3 mm SL, Inobe Port,
Inobe, Wadomari; KAUM–I. 90422, 75.1 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 90617, 13.0 mm
SL, Naikina Port, Nagamine, Wadomari; KAUM–I. 122363, 28.3 mm SL, off Okidomari Port, Shimojiro,
China; KAUM–I. 123408, 21.5 mm SL, Ujiji Beach, Ashikiyora, China; KAUM–I. 26769, 75.8 mm SL, off
Okinoerabu Island; KAUM–I. 26774, 72.0 mm SL, off Okinoerabu Island; KAUM–I. 90106, 83.0 mm SL,
Wadomari Port, Wadomari; KAUM–I. 90703, 14.4 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 90704,
12.4 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 122636, 27.4 mm SL, China Port, Serikaku, China;

62

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KAUM–I. 122637, 20.8 mm SL, China Port, Serikaku, China; KAUM–I. 122638, 11.7 mm SL, China Port,
Serikaku, China; KAUM–I. 122652, 24.8 mm SL, China Port, Serikaku, China; KAUM–I. 122653, 19.8 mm
SL, China Port, Serikaku, China.
Plectroglyphidodon leucozonus (Bleeker 1859)[Jpn name: Hakusensuzumedai]
Yoshigou et al. (2005a): HMNH 7869, Naikina, Nagamine, Wadomari; HMNH 7870, Naikina, Nagamine,
Wadomari; HMNH 7871, Naikina, Nagamine, Wadomari. Iwatsubo (2019a): KAUM–I. 122169, 16.7 mm
SL, Beach Rock, Tokudoki, China. This study: KAUM–I. 122169, 16.7 mm SL, Beach Rock, Tokudoki,
China; KAUM–I. 133548, 52.1 mm SL, Beach Rock, Tokudoki, China; KAUM–I. 133549, 41.9 mm SL,
Beach Rock, Tokudoki, China; KAUM–I. 133550, 42.3 mm SL, Beach Rock, Tokudoki, China; KAUM–
I. 133551, 30.2 mm SL, Beach Rock, Tokudoki, China; KAUM–I. 133552, 29.3 mm SL, Beach Rock,Tokudoki, China; KAUM–I. 133553, 26.3 mm SL, Beach Rock, Tokudoki, China.
Pomacentrus amboinensis Bleeker 1868
[Jpn name: Nisenettaisuzumedai]
KAUM–I. 90107, 53.0 mm SL, Wadomari Port, Wadomari; KAUM–I. 90108, 67.6 mm SL, Wadomari Port,
Wadomari; KAUM–I. 90109, 62.2 mm SL, Wadomari Port, Wadomari; KAUM–I. 122087, 42.1 mm SL,
Yakomo Beach, Yakomo, China.
Pomacentrus bankanensis Bleeker 1854
[Jpn name: Meganesuzumedai]
Iwatsubo (2018): KAUM–I. 90161, 54.4 mm SL, Okidomari Port, Shimojiro, China. Iwatsubo (2019a):
KAUM–I. 90161, 54.4 mm SL, Okidomari Port, Shimojiro, China. This study: KAUM–I. 90161, 54.4mm SL, Okidomari Port, Shimojiro, China; KAUM–I. 90162, 26.1 mm SL, Okidomari Port, Shimojiro,
China; KAUM–I. 122048, 15.0 mm SL, Masana, China; KAUM–I. 122054, 13.5 mm SL, Masana, China;
KAUM–I. 122127, 37.1 mm SL, Beach Rock, Tokudoki, China; KAUM–I. 122128, 33.2 mm SL, Beach
Rock, Tokudoki, China; KAUM–I. 122136, 31.0 mm SL, Beach Rock, Tokudoki, China; KAUM–I. 122144,
23.4 mm SL, Beach Rock, Tokudoki, China; KAUM–I. 122145, 41.6 mm SL, Beach Rock, Tokudoki,
China; KAUM–I. 122216, 27.8 mm SL, Wadomari Port, Wadomari; KAUM–I. 122364, 16.0 mm SL,
off Okidomari Port, Shimojiro, China; KAUM–I. 122397, 38.3 mm SL, Ujiji Beach, Ashikiyora, China;
KAUM–I. 122752, 23.9 mm SL, Kasaishi Beach, Kibiru, Wadomari; KAUM–I. 122776, 31.0 mm SL,
Kasaishi Beach, Kibiru, Wadomari; KAUM–I. 122788, 29.5 mm SL, Kasaishi Beach, Kibiru, Wadomari;
KAUM–I. 123421, 25.1 mm SL, Ujiji Beach, Ashikiyora, China; KAUM–I. 123469, 26.8 mm SL, Kasaishi
Beach, Kibiru, Wadomari; KAUM–I. 123470, 24.7 mm SL, Kasaishi Beach, Kibiru, Wadomari; KAUM–I.
123471, 24.7 mm SL, Kasaishi Beach, Kibiru, Wadomari; KAUM–I. 123478, 31.3 mm SL, Kasaishi Beach,
Kibiru, Wadomari; KAUM–I. 123479, 28.3 mm SL, Kasaishi Beach, Kibiru, Wadomari; KAUM–I. 123480,
26.4 mm SL, Kasaishi Beach, Kibiru, Wadomari; KAUM–I. 123481, 26.8 mm SL, Kasaishi Beach, Kibiru,
Wadomari; KAUM–I. 123482, 23.0 mm SL, Kasaishi Beach, Kibiru, Wadomari; KAUM–I. 123489, 18.9mm SL, Kasaishi Beach, Kibiru, Wadomari; KUN-P 48987, 34.9 mm SL, Ujiji Beach, Ashikiyora, China;
KUN-P 49066, 14.0 mm SL, China Port, Serikaku, China.
Pomacentrus chrysurus Cuvier 1830
[Jpn name: Ojirosuzumedai]
Iwatsubo (2018): KAUM–I. 90672, 23.9 mm SL, Wanjo Beach, Azefu, Wadomari. Iwatsubo (2019a):
KAUM–I. 90672, 23.9 mm SL, Wanjo Beach, Azefu, Wadomari. This study: KAUM–I. 36640, 58.2 mm
SL, off Okinoerabu Island; KAUM–I. 90502, 22.7 mm SL, Ujiji Beach, Ashikiyora, China; KAUM–I.
90503, 16.2 mm SL, Ujiji Beach, Ashikiyora, China; KAUM–I. 90672, 23.9 mm SL, Wanjo Beach, Azefu,

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63

Wadomari; KAUM–I. 90673, 19.1 mm SL, Wanjo Beach, Azefu, Wadomari; KAUM–I. 90718, 56.7 mm SL,
Inobe Port, Inobe, Wadomari; KAUM–I. 101620, 67.7 mm SL, Kunigami, Wadomari; KAUM–I. 122083,
25.7 mm SL, Yakomo Beach, Yakomo, China.
Pomacentrus coelestis Jordan & Starks 1901
[Jpn name: Sorasuzumedai]
Iwatsubo (2018): KAUM–I. 90426, 22.0 mm SL, Inobe Port, Inobe, Wadomari. Iwatsubo (2019a): KAUM–I.
90426, 22.0 mm SL, Inobe Port, Inobe, Wadomari. This study: KAUM–I. 2009, 42.0 mm SL, off Okinoerabu
Island; KAUM–I. 13627, 41.8 mm SL, off Okinoerabu Island; KAUM–I. 90426, 22.0 mm SL, Inobe Port,
Inobe, Wadomari; KAUM–I. 122249, 19.5 mm SL, west of Taminazaki, Tamina, China; KAUM–I. 122270,
15.7 mm SL, west of Taminazaki, Tamina, China; KAUM–I. 122698, 14.6 mm SL, off Yakomo, China;
KUN-P 48895, 14.9 mm SL, off Masana, China; KUN-P 49090, 17.0 mm SL, Kasaishi Beach, Kibiru,
Wadomari; KUN-P 49091, 15.1 mm SL, Kasaishi Beach, Kibiru, Wadomari; KUN-P 49109, 35.1 mm SL,
Yakomo Beach, Yakomo, China; KUN-P 49141, 36.7 mm SL, Kasaishi Beach, Kibiru, Wadomari; KUN-P
49143, 26.5 mm SL, Kasaishi Beach, Kibiru, Wadomari; KUN-P 49144, 30.5 mm SL, Kasaishi Beach,
Kibiru, Wadomari; KUN-P 49145, 26.1 mm SL, Kasaishi Beach, Kibiru, Wadomari; KUN-P 49146, 26.1 mm
SL, Kasaishi Beach, Kibiru, Wadomari; KUN-P 49150, 34.0 mm SL, Kasaishi Beach, Kibiru, Wadomari;
KUN-P 49151, 28.1 mm SL, Kasaishi Beach, Kibiru, Wadomari; KUN-P 49152, 25.0 mm SL, Kasaishi
Beach, Kibiru, Wadomari.
Pomacentrus lepidogenys Fowler & Bean 1928
[Jpn name: Asadosuzumedai]
Iwatsubo (2018): KAUM–I. 90434, 21.7 mm SL, Inobe Port, Inobe, Wadomari. Iwatsubo (2019a): KAUM–
I. 90434, 21.7 mm SL, Inobe Port, Inobe, Wadomari. This study: KAUM–I. 90434, 21.7 mm SL, Inobe Port,
Inobe, Wadomari.
Pomacentrus moluccensis Bleeker 1853
[Jpn name: Nettaisuzumedai]
Iwatsubo (2018): KAUM–I. 90433, 13.8 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 90766, 10.3mm SL, Naikina Port, Nagamine, Wadomari. Iwatsubo (2019a): KAUM–I. 90433, 13.8 mm SL, Inobe
Port, Inobe, Wadomari; KAUM–I. 90766, 10.3 mm SL, Naikina Port, Nagamine, Wadomari. This study:
KAUM–I. 90433, 13.8 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 90766, 10.3 mm SL, Naikina Port,
Nagamine, Wadomari.
Pomacentrus sp.
[Jpn name: N/A]
KAUM–I. 2132, 26.1 mm SL, off Okinoerabu Island; KAUM–I. 14627, 24.5 mm SL, off Okinoerabu Island;
KAUM–I. 14628, 23.4 mm SL, off Okinoerabu Island; KAUM–I. 14629, 20.7 mm SL, off Okinoerabu
Island; KAUM–I. 90047, 92.4 mm SL, Kasaishi Beach, Kibiru, Wadomari; KAUM–I. 121904, 60.0 mm SL,
Okidomari Port, Shimojiro, China; KAUM–I. 122769, 57.0 mm SL, Kasaishi Beach, Kibiru, Wadomari.
Pomacentrus vaiuli Jordan & Seale 1906
[Jpn name: Kuromeganesuzumedai]
KAUM–I. 90392, 48.5 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 90436, 64.6 mm SL, Inobe Port,
Inobe, Wadomari; KAUM–I. 132184, 16.8 mm SL, west of Taminazaki, Tamina, China.
Pomachromis richardsoni (Snyder 1909)[Jpn name: Okinawasuzumedai]
Yoshigou et al. (2005a): HMNH 7930, Okidomari, China.

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Stegastes albifasciatus (Schlegel & Müller 1840)[Jpn name: Kiobisuzumedai]
KAUM–I. 121885, 13.7 mm SL, Okidomari Port, Shimojiro, China; KAUM–I. 122089, 13.9 mm SL,
Yakomo Beach, Yakomo, China.
Stegastes altus (Okada & Ikeda 1937)[Jpn name: Sedakasuzumedai]
Yoshigou et al. (2005a): HMNH 7840, Shinjo, China; HMNH 7943, Okidomari, China; HMNH 7971,
Nishibaru, Wadomari; HMNH 7972, Nishibaru, Wadomari; HMNH 7973, Nishibaru, Wadomari. This study:
KAUM–I. 26780, 92.0 mm SL, off Okinoerabu Island; KAUM–I. 36602, 60.9 mm SL, off Okinoerabu
Island; KAUM–I. 36603, 71.3 mm SL, off Okinoerabu Island; KAUM–I. 36604, 75.4 mm SL, off
Okinoerabu Island; KAUM–I. 36605, 73.5 mm SL, off Okinoerabu Island; KAUM–I. 36606, 92.5 mm SL,
off Okinoerabu Island; KAUM–I. 36607, 91.6 mm SL, off Okinoerabu Island; KAUM–I. 36608, 103.0 mm
SL, off Okinoerabu Island; KAUM–I. 90103, 94.6 mm SL, Wadomari Port, Wadomari; KAUM–I. 90104,
89.6 mm SL, Wadomari Port, Wadomari; KAUM–I. 90105, 85.0 mm SL, Wadomari Port, Wadomari;
KAUM–I. 133489, 23.1 mm SL, Beach Rock, Tokudoki, China.
Stegastes fasciolatus (Ogilby 1889)[Jpn name: Fuchidorisuzumedai]
KAUM–I. 121760, 38.5 mm SL, Bishi Beach, Kunigami, Wadomari; KAUM–I. 122614, 17.3 mm SL, off
Yakomo, China; KAUM–I. 122634, 27.0 mm SL, China Port, Serikaku, China.
Stegastes nigricans (Lacepède 1802)[Jpn name: Kurosorasuzumedai]
KAUM–I. 101638, 43.3 mm SL, Kasaishi Beach, Kibiru, Wadomari; KAUM–I. 122387, 79.0 mm SL, Ujiji
Beach, Ashikiyora, China.
FAMILY TERAPONTIDAE
Terapon jarbua (Forsskål 1775)[Jpn name: Kotohiki]
KAUM–I. 121917, 12.6 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 121935, 11.1 mm SL, Inobe Port,
Inobe, Wadomari.
FAMILY KUHLIIDAE
Kuhlia marginata (Cuvier 1829)[Jpn name: Yugoi]
Yoshigou et al. (2005a): HMNH 7788, Kaikodo cave, Furusato, Wadomari. Yoshigou et al. (2005b): Hanjo,
Furusato, Wadomari; Kaikodo cave, Furusato, Wadomari. Yoshigou (2014): Okinoerabu Island.
Kuhlia mugil (Forster 1801)[Jpn name: Gin-yugoi]
KAUM–I. 90242, 13.0 mm SL, Sumiyoshi Beach, Sumiyoshi, China; KAUM–I. 121970, 171.5 mm SL,
Inobe Port, Inobe, Wadomari; KAUM–I. 122140, 19.4 mm SL, Beach Rock, Tokudoki, China; KUN-P
48879, 46.7 mm SL, Okidomari Port, Shimojiro, China; KUN-P 48880, 59.0 mm SL, Okidomari Port,
Shimojiro, China; KUN-P 48881, 42.2 mm SL, Okidomari Port, Shimojiro, China.

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65

FAMILY OPLEGNATHIDAE
Oplegnathus punctatus (Temminck & Schlegel 1844)[Jpn name: Ishigakidai]
KAUM–I. 122679, 317.7 mm SL, off Wadomari Port, Wadomari.
FAMILY MICROCANTHIDAE
Microcanthus strigatus (Cuvier 1831)[Jpn name: Kagokakidai]
Underwater photograph (Pl. 6-I).
FAMILY GIRELLIDAE
Girella mezina Jordan & Starks 1907
[Jpn name: Okinamejina]
Yoshigou et al. (2005a): HMNH 7792, Hanjo, Furusato, Wadomari. Yoshigou (2014): Okinoerabu Island.
This study: KAUM–I. 90221, 37.2 mm SL, Sumiyoshi Beach, Sumiyoshi, China; KAUM–I. 90222, 49.0mm SL, Sumiyoshi Beach, Sumiyoshi, China; KAUM–I. 90223, 44.4 mm SL, Sumiyoshi Beach, Sumiyoshi,
China; KAUM–I. 90224, 37.0 mm SL, Sumiyoshi Beach, Sumiyoshi, China; KUN-P 49092, 11.9 mm SL,
Kasaishi Beach, Kibiru, Wadomari.
FAMILY LABRIDAE
Bodianus anthioides (Bennett 1832)[Jpn name: Hiodoshibera]
Underwater photograph (Pl. 6-J).
Bodianus bilunulatus (Lacepède 1801)[Jpn name: Kitsunebera]
KAUM–I. 123528, 303.3 mm SL, Hirase, Tamashiro, Wadomari.
Bodianus perditio (Quoy & Gaimard 1834)[Jpn name: Takibera]
KAUM–I. 123529, 225.2 mm SL, off Okinoerabu Island.
Cheilinus trilobatus Lacepède 1801
[Jpn name: Mitsubamochinouo]
Yoshigou et al. (2005a): HMNH 7987, Nishibaru, Wadomari. This study: KAUM–I. 123472, 35.5 mm SL,
Kasaishi Beach, Kibiru, Wadomari; KUN-P 49135, 25.7 mm SL, Kasaishi Beach, Kibiru, Wadomari.
Cirrhilabrus cyanopleura (Bleeker 1851)[Jpn name: Kuroheriitohikibera]
KAUM–I. 90376, 23.4 mm SL, Inobe Port, Inobe, Wadomari.
Cirrhilabrus exquisitus Smith 1957
[Jpn name: Nishikiitohikibera]
Underwater photograph (Pl. 7-A).
Cirrhilabrus katherinae Randall 1992
[Jpn name: Goshikiitohikibera]
Underwater photograph (Pl. 7-B).

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H. Motomura and K. Uehara

Cirrhilabrus lunatus Randall & Masuda 1991
[Jpn name: Tsukinowaitohikibera]
Underwater photograph (Pl. 7-C).
Cirrhilabrus melanomarginatus Randall & Shen 1978
[Jpn name: Tomoshibiitohikibera]
Underwater photograph (Pl. 7-D).
Cirrhilabrus rubrimarginatus Randall 1992
[Jpn name: Benihireitohikibera]
Underwater photograph (Pl. 7-E).
Coris aygula Lacepède 1801
[Jpn name: Kammuribera]
KAUM–I. 1899, 2 specimens, 62.1–67.9 mm SL, off Okinoerabu Island; KAUM–I. 1915, 117.1 mm SL, off
Okinoerabu Island; KUN-P 48917, 69.9 mm SL, Yakomo Beach, Yakomo, China.
Coris gaimard (Quoy & Gaimard 1824)[Jpn name: Tsuyubera]
KAUM–I. 90377, 29.9 mm SL, Inobe Port, Inobe, Wadomari.
Gomphosus varius Lacepède 1801
[Jpn name: Kugibera]
KAUM–I. 90143, 23.5 mm SL, Okidomari Port, Shimojiro, China.
Halichoeres hortulanus (Lacepède 1801)[Jpn name: Tokarabera]
KAUM–I. 122389, 28.2 mm SL, Ujiji Beach, Ashikiyora, China.
Halichoeres margaritaceus (Valenciennes 1839)[Jpn name: Akanijibera]
Yoshigou et al. (2005a): HMNH 7846, Shinjo, China. Motomura (2019b): KAUM–I. 122091, 29.6 mm SL,
Yakomo Beach, Yakomo, China. This study: KAUM–I. 2001, 53.2 mm SL, off Okinoerabu Island; KAUM–I.
14238, 54.2 mm SL, off Okinoerabu Island; KAUM–I. 14239, 38.8 mm SL, off Okinoerabu Island; KAUM–
I. 14240, 35.2 mm SL, off Okinoerabu Island; KAUM–I. 122091, 29.6 mm SL, Yakomo Beach, Yakomo,
China; KAUM–I. 122135, 40.6 mm SL, Beach Rock, Tokudoki, China.
Halichoeres marginatus Rüppell 1835
[Jpn name: Kanokobera]
Yoshigou et al. (2005a): HMNH 7823, Shinjo, China; HMNH 7895, Okidomari, China; HMNH 7896,
Okidomari, China. This study: KAUM–I. 90781, 8.9 mm SL, Hanzaki, Wadomari; KAUM–I. 122098, 12.9mm SL, Yakomo Beach, Yakomo, China; KAUM–I. 122399, 39.6 mm SL, Ujiji Beach, Ashikiyora, China;
KAUM–I. 123407, 31.3 mm SL, Ujiji Beach, Ashikiyora, China; KUN-P 49107, 10.9 mm SL, Yakomo
Beach, Yakomo, China.
Halichoeres melanochir Fowler & Bean 1928
[Jpn name: Munatembera]
KAUM–I. 90640, 23.2 mm SL, Naikina Port, Nagamine, Wadomari.

Marine and freshwater fishes of Okinoerabu Island

67

Halichoeres trimaculatus (Quoy & Gaimard 1834)[Jpn name: Mitsuboshikyusen]
KAUM–I. 2757, 86.6 mm SL, off Okinoerabu Island; KAUM–I. 90543, 33.9 mm SL, Ujiji Beach,
Ashikiyora, China; KAUM–I. 122090, 30.0 mm SL, Yakomo Beach, Yakomo, China; KAUM–I. 122094,
20.7 mm SL, Yakomo Beach, Yakomo, China; KAUM–I. 122766, 57.5 mm SL, Kasaishi Beach, Kibiru,
Wadomari; KAUM–I. 122767, 38.4 mm SL, Kasaishi Beach, Kibiru, Wadomari.
Hemigymnus fasciatus (Bloch 1792)[Jpn name: Shimatarekuchibera]
KAUM–I. 90435, 70.8 mm SL, Inobe Port, Inobe, Wadomari.
Hemigymnus melapterus (Bloch 1791)[Jpn name: Tarekuchibera]
KAUM–I. 90770, 11.5 mm SL, Naikina Port, Nagamine, Wadomari.
Iniistius melanopus (Bleeker 1857)[Jpn name: Monhirabera]
Motomura (2019b): KAUM–I. 122655, 142.8 mm SL, off Naikina Port, Nagamine, Wadomari. This study:
KAUM–I. 122655, 142.8 mm SL, off Naikina Port, Nagamine, Wadomari.
Labroides bicolor Fowler & Bean 1928
[Jpn name: Somewakebera]
Motomura (2019b): KAUM–I. 123553, 24.8 mm SL, Yaja, China. This study: KAUM–I. 123553, 24.8 mm
SL, Yaja, China.
Labroides dimidiatus (Valenciennes 1839)[Jpn name: Honsomewakebera]
KAUM–I. 2040, 64.6 mm SL, off Okinoerabu Island; KAUM–I. 90644, 22.5 mm SL, Naikina Port,
Nagamine, Wadomari; KAUM–I. 122256, 14.9 mm SL, west of Taminazaki, Tamina, China; KAUM–I.
122337, 11.3 mm SL, Taminazaki, Tamina, China; KAUM–I. 122386, 28.1 mm SL, Ujiji Beach, Ashikiyora,
China; KAUM–I. 122702, 10.2 mm SL, off Yakomo, China; KAUM–I. 122703, 13.2 mm SL, off Yakomo,
China; KUN-P 49093, 12.0 mm SL, Yakomo Beach, Yakomo, China.
Labropsis manabei Schmidt 1931
[Jpn name: Manabebera]
KAUM–I. 122053, 12.3 mm SL, Masana, China.
Labropsis xanthonota Randall 1981
[Jpn name: Miyakebera]
Fukui (2018): KAUM–I. 90401, 24.9 mm SL, Inobe Port, Inobe, Wadomari. Motomura (2019b): KAUM–
I. 90401, 24.9 mm SL, Inobe Port, Inobe, Wadomari. This study: KAUM–I. 90401, 24.9 mm SL, Inobe Port,
Inobe, Wadomari.
Novaculichthys taeniourus (Lacepède 1801)[Jpn name: Obitensumodoki]
KAUM–I. 90444, 83.2 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 122804, 13.6 mm SL, Sumiyoshi
Port, Sumiyoshi, China.

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H. Motomura and K. Uehara

Oxycheilinus unifasciatus (Streets 1877)[Jpn name: Hitosujimochinouo]
KAUM–I. 123506, 143.2 mm SL, off Okidomari Port, Shimojiro, China.
Pseudocheilinus hexataenia (Bleeker 1857)[Jpn name: Nisemochinouo]
KAUM–I. 90383, 33.3 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 90390, 14.4 mm SL, Inobe Port,
Inobe, Wadomari; KAUM–I. 90620, 7.5 mm SL, Naikina Port, Nagamine, Wadomari; KAUM–I. 123556,
24.3 mm SL, Yaja, China; KUN-P 49103, 35.2 mm SL, Yakomo Beach, Yakomo, China.
Pseudocheilinus ocellatus Randall 1999
[Jpn name: Yokoshimanisemochinouo]
Underwater photograph (Pl. 7-F).
Pseudocheilinus octotaenia Jenkins 1901
[Jpn name: Yasujinisemochinouo]
KAUM–I. 122045, 14.1 mm SL, Masana, China; KAUM–I. 122285, 11.4 mm SL, west of Taminazaki,
Tamina, China; KAUM–I. 122605, 56.6 mm SL, off Yakomo, China; KAUM–I. 122619, 13.2 mm SL, off
Yakomo, China; KAUM–I. 123577, 14.3 mm SL, Yaja, China.
Pseudocoris aurantiofasciata Fourmanoir 1971
[Jpn name: Shiratakiberadamashi]
Underwater photograph (Pl. 7-G).
Stethojulis bandanensis (Bleeker 1851)[Jpn name: Akaobibera]
KAUM–I. 1969, 81.2 mm SL, off Okinoerabu Island; KAUM–I. 2035, U mm SL, off Okinoerabu Island;
KAUM–I. 3342, 81.4 mm SL, off Okinoerabu Island; KAUM–I. 3343, 88.5 mm SL, off Okinoerabu Island;
KAUM–I. 3344, 71.7 mm SL, off Okinoerabu Island; KAUM–I. 3345, 70.0 mm SL, off Okinoerabu Island;
KAUM–I. 3346, 83.2 mm SL, off Okinoerabu Island; KAUM–I. 3347, 76.2 mm SL, off Okinoerabu Island;
KAUM–I. 3348, 60.7 mm SL, off Okinoerabu Island; KAUM–I. 3349, 71.8 mm SL, off Okinoerabu Island;
KAUM–I. 3350, 49.1 mm SL, off Okinoerabu Island; KAUM–I. 3351, 72.4 mm SL, off Okinoerabu Island;
KAUM–I. 3352, 67.4 mm SL, off Okinoerabu Island; KAUM–I. 3353, 53.0 mm SL, off Okinoerabu Island;
KAUM–I. 3354, 61.8 mm SL, off Okinoerabu Island; KAUM–I. 3355, 81.3 mm SL, off Okinoerabu Island;
KAUM–I. 3356, 72.8 mm SL, off Okinoerabu Island; KAUM–I. 3357, 58.0 mm SL, off Okinoerabu Island;
KAUM–I. 3358, 74.5 mm SL, off Okinoerabu Island; KAUM–I. 3359, 81.8 mm SL, off Okinoerabu Island;
KAUM–I. 3360, 80.2 mm SL, off Okinoerabu Island; KAUM–I. 3361, 73.1 mm SL, off Okinoerabu Island;
KAUM–I. 3362, 78.4 mm SL, off Okinoerabu Island; KAUM–I. 3363, 76.9 mm SL, off Okinoerabu Island;
KAUM–I. 3364, 68.7 mm SL, off Okinoerabu Island; KAUM–I. 3365, 39.2 mm SL, off Okinoerabu Island;
KAUM–I. 3366, 60.8 mm SL, off Okinoerabu Island; KAUM–I. 14195, 50.5 mm SL, off Okinoerabu
Island; KAUM–I. 14196, 49.4 mm SL, off Okinoerabu Island; KAUM–I. 14197, 47.5 mm SL, off
Okinoerabu Island; KAUM–I. 14198, 43.3 mm SL, off Okinoerabu Island; KAUM–I. 14199, 44.2 mm SL,
off Okinoerabu Island; KAUM–I. 14200, 41.1 mm SL, off Okinoerabu Island; KAUM–I. 14201, 37.5 mm
SL, off Okinoerabu Island; KAUM–I. 14202, 42.0 mm SL, off Okinoerabu Island; KAUM–I. 14203, 38.9mm SL, off Okinoerabu Island; KAUM–I. 14204, 35.5 mm SL, off Okinoerabu Island; KAUM–I. 14205,
36.3 mm SL, off Okinoerabu Island; KAUM–I. 14206, 33.1 mm SL, off Okinoerabu Island; KAUM–I.
14207, 33.9 mm SL, off Okinoerabu Island; KAUM–I. 14208, 28.2 mm SL, off Okinoerabu Island; KAUM–
I. 122147, 58.8 mm SL, Beach Rock, Tokudoki, China; KAUM–I. 123466, 37.3 mm SL, Kasaishi Beach,

Marine and freshwater fishes of Okinoerabu Island

69

Kibiru, Wadomari; AUM–I. 133524, 31.5 mm SL, Beach Rock, Tokudoki, China; KAUM–I. 133525, 30.8mm SL, Beach Rock, Tokudoki, China; KAUM–I. 133526, 20.2 mm SL, Beach Rock, Tokudoki, China;
KAUM–I. 133527, 10.0 mm SL, Beach Rock, Tokudoki, China; KUN-P 49142, 38.7 mm SL, Kasaishi
Beach, Kibiru, Wadomari.
Stethojulis strigiventer (Bennett 1833)[Jpn name: Harasujibera]
Yoshigou et al. (2005a): HMNH 7929, Okidomari, China. Fukui (2018): KAUM–I. 90096, 20.2 mm
SL, Kasaishi Beach, Kibiru, Wadomari; KAUM–I. 90544, 39.9 mm SL, Ujiji Beach, Ashikiyora, China.
Motomura (2019b): KAUM–I. 90096, 20.2 mm SL, Kasaishi Beach, Kibiru, Wadomari; KAUM–I. 90544,
39.9 mm SL, Ujiji Beach, Ashikiyora, China. This study: KAUM–I. 90096, 20.2 mm SL, Kasaishi Beach,
Kibiru, Wadomari; KAUM–I. 90544, 39.9 mm SL, Ujiji Beach, Ashikiyora, China; KAUM–I. 90545, 21.5mm SL, Ujiji Beach, Ashikiyora, China; KAUM–I. 90689, 30.1 mm SL, Ujiji Beach, Ashikiyora, China;
KAUM–I. 122768, 74.3 mm SL, Kasaishi Beach, Kibiru, Wadomari; KAUM–I. 123490, 19.6 mm SL,
Kasaishi Beach, Kibiru, Wadomari.
Thalassoma amblycephalum (Bleeker 1856)[Jpn name: Kogashirabera]
KAUM–I. 2051, 102.9 mm SL, off Okinoerabu Island; KAUM–I. 133478, 28.8 mm SL, Beach Rock,Tokudoki, China; KAUM–I. 133479, 19.3 mm SL, Beach Rock, Tokudoki, China; KAUM–I. 133480, 20.3mm SL, Beach Rock, Tokudoki, China; KAUM–I. 133481, 15.3 mm SL, Beach Rock, Tokudoki, China;
KAUM–I. 133482, 14.9 mm SL, Beach Rock, Tokudoki, China.
Thalassoma cupido (Temminck & Schlegel 1845)[Jpn name: Nishikibera]
Yoshigou et al. (2005a): HMNH 7874, Naikina, Nagamine, Wadomari; HMNH 7928, Okidomari, China.
This study: KAUM–I. 1978, 62.0 mm SL, off Okinoerabu Island; KAUM–I. 11019, 22.7 mm SL, off
Okinoerabu Island; KAUM–I. 11020, 15.2 mm SL, off Okinoerabu Island; KAUM–I. 11021, 17.4 mm SL,
off Okinoerabu Island; KAUM–I. 11022, 17.7 mm SL, off Okinoerabu Island; KAUM–I. 11023, 23.8 mm
SL, off Okinoerabu Island; KAUM–I. 11024, 27.3 mm SL, off Okinoerabu Island; KAUM–I. 11025, 60.0
mm SL, off Okinoerabu Island; KAUM–I. 11026, 44.7 mm SL, off Okinoerabu Island; KAUM–I. 11027,
40.0 mm SL, off Okinoerabu Island; KAUM–I. 11028, 39.4 mm SL, off Okinoerabu Island; KAUM–I.
11029, 42.7 mm SL, off Okinoerabu Island; KAUM–I. 11030, 15.2 mm SL, off Okinoerabu Island; KAUM–
I. 11031, 44.6 mm SL, off Okinoerabu Island; KAUM–I. 11032, 41.4 mm SL, off Okinoerabu Island;
KAUM–I. 11033, 45.1 mm SL, off Okinoerabu Island; KAUM–I. 11034, 43.2 mm SL, off Okinoerabu
Island; KAUM–I. 11035, 49.9 mm SL, off Okinoerabu Island; KAUM–I. 11036, 51.7 mm SL, off
Okinoerabu Island; KAUM–I. 11037, 37.8 mm SL, off Okinoerabu Island; KAUM–I. 11038, 18.7 mm SL,
off Okinoerabu Island; KAUM–I. 11039, 51.6 mm SL, off Okinoerabu Island; KAUM–I. 11040, 50.6 mm
SL, off Okinoerabu Island; KAUM–I. 11041, 43.1 mm SL, off Okinoerabu Island; KAUM–I. 11042, 13.2mm SL, off Okinoerabu Island; KAUM–I. 11043, 40.7 mm SL, off Okinoerabu Island; KAUM–I. 11044,
52.6 mm SL, off Okinoerabu Island; KAUM–I. 11045, 46.4 mm SL, off Okinoerabu Island; KAUM–I.
11046, 39.6 mm SL, off Okinoerabu Island; KAUM–I. 11047, 50.8 mm SL, off Okinoerabu Island; KAUM–
I. 11048, 48.4 mm SL, off Okinoerabu Island; KAUM–I. 11049, 46.7 mm SL, off Okinoerabu Island;
KAUM–I. 11050, 37.2 mm SL, off Okinoerabu Island; KAUM–I. 11051, 23.4 mm SL, off Okinoerabu
Island; KAUM–I. 11052, 41.1 mm SL, off Okinoerabu Island; KAUM–I. 11053, 47.8 mm SL, off
Okinoerabu Island; KAUM–I. 11054, 49.8 mm SL, off Okinoerabu Island; KAUM–I. 11055, 51.4 mm SL,
off Okinoerabu Island; KAUM–I. 11056, 48.3 mm SL, off Okinoerabu Island; KAUM–I. 11057, 51.2 mm
SL, off Okinoerabu Island; KAUM–I. 11058, 47.2 mm SL, off Okinoerabu Island; KAUM–I. 11059, 53.7

70

H. Motomura and K. Uehara

mm SL, off Okinoerabu Island; KAUM–I. 11060, 45.9 mm SL, off Okinoerabu Island; KAUM–I. 11061,
50.0 mm SL, off Okinoerabu Island; KAUM–I. 11062, 49.6 mm SL, off Okinoerabu Island; KAUM–I.
11063, 50.1 mm SL, off Okinoerabu Island; KAUM–I. 11064, 41.8 mm SL, off Okinoerabu Island; KAUM–
I. 11065, 45.6 mm SL, off Okinoerabu Island; KAUM–I. 11066, 49.4 mm SL, off Okinoerabu Island;
KAUM–I. 11067, 44.0 mm SL, off Okinoerabu Island; KAUM–I. 11068, 49.7 mm SL, off Okinoerabu
Island; KAUM–I. 11069, 43.0 mm SL, off Okinoerabu Island; KAUM–I. 11070, 50.8 mm SL, off
Okinoerabu Island; KAUM–I. 11071, 49.0 mm SL, off Okinoerabu Island; KAUM–I. 11072, 47.6 mm SL,
off Okinoerabu Island; KAUM–I. 11073, 48.8 mm SL, off Okinoerabu Island; KAUM–I. 11074, 50.6 mm
SL, off Okinoerabu Island; KAUM–I. 11075, 54.2 mm SL, off Okinoerabu Island; KAUM–I. 11076, 55.5mm SL, off Okinoerabu Island; KAUM–I. 11077, 52.7 mm SL, off Okinoerabu Island; KAUM–I. 11078,
64.5 mm SL, off Okinoerabu Island; KAUM–I. 11079, 57.9 mm SL, off Okinoerabu Island; KAUM–I.
11080, 56.6 mm SL, off Okinoerabu Island; KAUM–I. 11081, 48.7 mm SL, off Okinoerabu Island; KAUM–
I. 11082, 56.1 mm SL, off Okinoerabu Island; KAUM–I. 11083, 56.7 mm SL, off Okinoerabu Island;
KAUM–I. 11084, 51.1 mm SL, off Okinoerabu Island; KAUM–I. 11085, 60.0 mm SL, off Okinoerabu
Island; KAUM–I. 11086, 57.9 mm SL, off Okinoerabu Island; KAUM–I. 11087, 60.8 mm SL, off
Okinoerabu Island; KAUM–I. 11088, 58.5 mm SL, off Okinoerabu Island; KAUM–I. 11089, 60.7 mm SL,
off Okinoerabu Island; KAUM–I. 11090, 54.1 mm SL, off Okinoerabu Island; KAUM–I. 11091, 60.4 mm
SL, off Okinoerabu Island; KAUM–I. 11092, 61.2 mm SL, off Okinoerabu Island; KAUM–I. 11093, 48.5mm SL, off Okinoerabu Island; KAUM–I. 11094, 52.3 mm SL, off Okinoerabu Island; KAUM–I. 11095,
50.8 mm SL, off Okinoerabu Island; KAUM–I. 11096, 54.2 mm SL, off Okinoerabu Island; KAUM–I.
11097, 61.2 mm SL, off Okinoerabu Island; KAUM–I. 11098, 62.3 mm SL, off Okinoerabu Island; KAUM–
I. 11099, 58.8 mm SL, off Okinoerabu Island; KAUM–I. 11100, 55.5 mm SL, off Okinoerabu Island;
KAUM–I. 11101, 64.4 mm SL, off Okinoerabu Island; KAUM–I. 11102, 19.5 mm SL, off Okinoerabu
Island.
Thalassoma hardwicke (Bennett 1830)[Jpn name: Senasujibera]
KAUM–I. 1976, 58.8 mm SL, off Okinoerabu Island; KAUM–I. 2016, 60.9 mm SL, off Okinoerabu Island;
KAUM–I. 14235, 74.7 mm SL, off Okinoerabu Island; KAUM–I. 14236, 49.1 mm SL, off Okinoerabu
Island; KAUM–I. 20766, 68.3 mm SL, off Okinoerabu Island; KAUM–I. 20767, 54.8 mm SL, off
Okinoerabu Island; KAUM–I. 20768, 59.8 mm SL, off Okinoerabu Island; KAUM–I. 122174, 13.1 mm SL,
Beach Rock, Tokudoki, China.
Thalassoma jansenii (Bleeker 1856)[Jpn name: Yansennishikibera]
Motomura (2019b): KAUM–I. 122796, 98.0 mm SL, Kunigami, Wadomari. This study: KAUM–I. 2139,
76.0 mm SL, off Okinoerabu Island; KAUM–I. 122796, 98.0 mm SL, Kunigami, Wadomari; KAUM–
I. 133535, 18.1 mm SL, Beach Rock, Tokudoki, China; KAUM–I. 133536, 17.3 mm SL, Beach Rock,Tokudoki, China.
Thalassoma lutescens (Lay & Bennett 1839)[Jpn name: Yamabukibera]
Yoshigou et al. (2005a): HMNH 7876, Naikina, Nagamine, Wadomari; HMNH 7976, Nishibaru, Wadomari.
Motomura (2019b): KAUM–I. 122042, 16.0 mm SL, Masana, China. This study: KAUM–I. 90110, 91.3mm SL, Wadomari Port, Wadomari; KAUM–I. 90111, 93.8 mm SL, Wadomari Port, Wadomari; KAUM–I.
90112, 88.0 mm SL, Wadomari Port, Wadomari; KAUM–I. 90113, 77.3 mm SL, Wadomari Port, Wadomari;
KAUM–I. 90114, 82.0 mm SL, Wadomari Port, Wadomari; KAUM–I. 90115, 90.5 mm SL, Wadomari Port,
Wadomari; KAUM–I. 90122, 103.1 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 90142, 14.9 mm SL,

Marine and freshwater fishes of Okinoerabu Island

71

Okidomari Port, Shimojiro, China; KAUM–I. 90371, 151.5 mm SL, Inobe Port, Inobe, Wadomari; KAUM–
I. 90416, 134.0 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 90417, 132.5 mm SL, Inobe Port, Inobe,
Wadomari; KAUM–I. 90418, 139.9 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 90419, 101.9 mm
SL, Inobe Port, Inobe, Wadomari; KAUM–I. 90451, 104.1 mm SL, Inobe Port, Inobe, Wadomari; KAUM–
I. 101611, 93.6 mm SL, Kunigami, Wadomari; KAUM–I. 122041, 15.7 mm SL, Masana, China; KAUM–
I. 122042, 16.0 mm SL, Masana, China; KAUM–I. 122107, 17.6 mm SL, Yakomo Beach, Yakomo, China;
KAUM–I. 122656, 132.7 mm SL, off Naikina Port, Nagamine, Wadomari; KAUM–I. 122797, 82.5 mm SL,
Kunigami, Wadomari; KAUM–I. 122798, 99.3 mm SL, Kunigami, Wadomari; KAUM–I. 123485, 22.3 mm
SL, Kasaishi Beach, Kibiru, Wadomari; KUN-P 49002, 15.1 mm SL, Yaja, China.
Thalassoma purpureum (Forsskål 1775)[Jpn name: Kinubera]
Yoshigou et al. (2005a): HMNH 7875, Naikina, Nagamine, Wadomari. This study: KUN-P 49138, 16.2 mm
SL, Kasaishi Beach, Kibiru, Wadomari; KUN-P 49139, 23.2 mm SL, Kasaishi Beach, Kibiru, Wadomari;
KUN-P 49140, 22.0 mm SL, Kasaishi Beach, Kibiru, Wadomari.
Thalassoma quinquevittatum (Lay & Bennett 1839)[Jpn name: Hakobera]
Motomura (2019b): KAUM–I. 122290, 18.8 mm SL, west of Taminazaki, Tamina, China; KAUM–I.
122795, 100.5 mm SL, Kunigami, Wadomari. This study: KAUM–I. 1987, 73.7 mm SL, off Okinoerabu
Island; KAUM–I. 14647, 52.8 mm SL, off Okinoerabu Island; KAUM–I. 101612, 86.4 mm SL, Kunigami,
Wadomari; KAUM–I. 122290, 18.8 mm SL, west of Taminazaki, Tamina, China; KAUM–I. 122795, 100.5mm SL, Kunigami, Wadomari.
Thalassoma sp.
[Jpn name: N/A]
KAUM–I. 123422, 18.7 mm SL, Ujiji Beach, Ashikiyora, China; KAUM–I. 123423, 18.6 mm SL, Ujiji
Beach, Ashikiyora, China.
Wetmorella nigropinnata (Seale 1901)[Jpn name: Hashinagabera]
KAUM–I. 90449, 55.9 mm SL, Inobe Port, Inobe, Wadomari.
FAMILY SCARIDAE
Cetoscarus ocellatus (Valenciennes 1840)[Jpn name: Irobudai]
Sakurai (2018b): KAUM–I. 90167, 22.7 mm SL, Okidomari Port, Shimojiro, China. Sakurai (2019b):
KAUM–I. 90167, 22.7 mm SL, Okidomari Port, Shimojiro, China. This study: KAUM–I. 90167, 22.7 mm
SL, Okidomari Port, Shimojiro, China; KAUM–I. 122736, 6.5 mm SL, Yaja, China.
Remarks: This species has been treated as Cetoscarus bicolor (Rüppell 1829), but it is a species endemic to
the Res Sea.
Chlorurus japanensis (Bloch 1789)[Jpn name: Shijukara]
KAUM–I. 44394, 276.5 mm SL, off Okinoerabu Island; KAUM–I. 44395, 270.8 mm SL, off Okinoerabu
Island; KAUM–I. 44396, 289.2 mm SL, off Okinoerabu Island; KAUM–I. 59453, 261.1 mm SL, off
Okinoerabu Island.

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Chlorurus microrhinos (Bleeker 1854)[Jpn name: Nan-yobudai]
Sakurai (2018b): KAUM–I. 90725, 279.5 mm SL, Nishibaru Beach, Nishibaru, Wadomari. Sakurai (2019b):
KAUM–I. 90725, 279.5 mm SL, Nishibaru Beach, Nishibaru, Wadomari. This study: KAUM–I. 90003,
231.3 mm SL, north of Okinoerabu Island; KAUM–I. 90725, 279.5 mm SL, Nishibaru Beach, Nishibaru,
Wadomari; KAUM–I. 122067, 425.0 mm SL, Kunigami, Wadomari.
Scaridae, undetermined genus and species[Jpn name: N/A]
KAUM–I. 90015, 16.0 mm SL, Okidomari Port, Shimojiro, China; KAUM–I. 90041, 15.8 mm SL, Kasaishi
Beach, Kibiru, Wadomari; KAUM–I. 90042, 40.4 mm SL, Kasaishi Beach, Kibiru, Wadomari; KAUM–
I. 90186, 19.6 mm SL, Yakomo Beach, Yakomo, China; KAUM–I. 90188, 19.6 mm SL, Yakomo Beach,
Yakomo, China; KAUM–I. 90193, 24.2 mm SL, Yakomo Beach, Yakomo, China.
Scarus forsteni (Bleeker 1861)[Jpn name: Ichimonjibudai]
Sakurai (2018b): KAUM–I. 90344, 319.0 mm SL, off Okidomari Port, Shimojiro, China. Sakurai (2019b):
KAUM–I. 90344, 319.0 mm SL, off Okidomari Port, Shimojiro, China. This study: KAUM–I. 90344, 319.0
mm SL, off Okidomari Port, Shimojiro, China.
Scarus globiceps Valenciennes 1840
[Jpn name: Daidaibudai]
Sakurai (2018b): KAUM–I. 90002, 213.8 mm SL, north of Okinoerabu Island. Sakurai (2019b): KAUM–
I. 90002, 213.8 mm SL, north of Okinoerabu Island. This study: KAUM–I. 90002, 213.8 mm SL, north of
Okinoerabu Island.
Scarus niger Forsskål 1775
[Jpn name: Buchibudai]
KAUM–I. 90724, 255.4 mm SL, Nishibaru Beach, Nishibaru, Wadomari.
Scarus rivulatus Valenciennes 1840
[Jpn name: Sujibudai]
KAUM–I. 101641, 95.2 mm SL, Kasaishi Beach, Kibiru, Wadomari.
Scarus rubroviolaceus Bleeker 1847
[Jpn name: Nagabudai]
KAUM–I. 122182, 341.6 mm SL, off Wadomari.
Scarus schlegeli (Bleeker 1861)[Jpn name: Obibudai]
KAUM–I. 90452, 187.8 mm SL, Inobe Port, Inobe, Wadomari.
Scarus xanthopleura Bleeker 1853
[Jpn name: Akabudai]
Photograph (Pl. 7-H).

Marine and freshwater fishes of Okinoerabu Island

73

FAMILY PINGUIPEDIDAE
Parapercis cylindrica (Bloch 1792)[Jpn name: Dandaratoragisu]
KAUM–I. 90124, 114.0 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 90125, 99.0 mm SL, Inobe Port,
Inobe, Wadomari; KUN-P 48913, 61.9 mm SL, Yakomo Beach, Yakomo, China.
Parapercis millepunctata (Günther 1860)[Jpn name: Wanuketoragisu]
KAUM–I. 121867, 118.4 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 121868, 133.7 mm SL, Inobe
Port, Inobe, Wadomari; KAUM–I. 122148, 59.7 mm SL, Beach Rock, Tokudoki, China.
Parapercis pacifica Imamura & Yoshino 2007
[Jpn name: Ogurotoragisu]
KAUM–I. 90420, 150.2 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 90454, 165.1 mm SL, Inobe Port,
Inobe, Wadomari; KAUM–I. 121865, 102.1 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 121866, 92.7mm SL, Inobe Port, Inobe, Wadomari; KUN-P 49010, 114.8 mm SL, Yaja, China.
FAMILY CREEDIIDAE
Limnichthys fasciatus Waite 1904
[Jpn name: Tobigimpo]
Yoshigou et al. (2005a): HMNH 7978, Nishibaru, Wadomari.
FAMILY TRIPTERYGIIDAE
Enneapterygius etheostoma (Jordan & Snyder 1902)[Jpn name: Hebigimpo]
Yoshigou et al. (2005a, as Enneapterygius etheostomus): HMNH 7931, Okidomari, China; HMNH 7932,
Okidomari, China. This study: KAUM–I. 90194, 24.9 mm SL, Yakomo Beach, Yakomo, China; KAUM–I.
90523, 26.0 mm SL, Ujiji Beach, Ashikiyora, China; KAUM–I. 122649, 38.3 mm SL, China Port, Serikaku,
China; KAUM–I. 123415, 38.8 mm SL, Ujiji Beach, Ashikiyora, China.
Enneapterygius flavoccipitis Shen 1994
[Jpn name: Erimakihebigimpo]
KAUM–I. 122309, 20.3 mm SL, west of Taminazaki, Tamina, China; KAUM–I. 122625, 20.1 mm SL, off
Yakomo, China; KAUM–I. 123569, 20.5 mm SL, Yaja, China; KAUM–I. 123570, 19.5 mm SL, Yaja, China;
KAUM–I. 123571, 19.7 mm SL, Yaja, China.
Enneapterygius philippinus (Peters 1868)[Jpn name: Kusagimpo]
Yoshigou et al. (2005a): HMNH 7830, 7 specimens, Shinjo, China; HMNH 7977, Nishibaru, Wadomari.
This study: KAUM–I. 123428, 20.0 mm SL, Ujiji Beach, Ashikiyora, China; KAUM–I. 133510, 20.7 mm
SL, Beach Rock, Tokudoki, China; KAUM–I. 133511, 19.9 mm SL, Beach Rock, Tokudoki, China; KAUM–
I. 133512, 16.7 mm SL, Beach Rock, Tokudoki, China; KAUM–I. 133513, 18.0 mm SL, Beach Rock,Tokudoki, China; KAUM–I. 133514, 20.6 mm SL, Beach Rock, Tokudoki, China; KAUM–I. 133515, 21.3mm SL, Beach Rock, Tokudoki, China; KAUM–I. 133516, 16.8 mm SL, Beach Rock, Tokudoki, China;
KAUM–I. 133517, 15.9 mm SL, Beach Rock, Tokudoki, China; KAUM–I. 133518, 18.4 mm SL, Beach
Rock, Tokudoki, China; KAUM–I. 133519, 18.2 mm SL, Beach Rock, Tokudoki, China; KAUM–I. 133520,
12.1 mm SL, Beach Rock, Tokudoki, China; KAUM–I. 133542, 20.4 mm SL, Beach Rock, Tokudoki, China;
KAUM–I. 133543, 18.3 mm SL, Beach Rock, Tokudoki, China; KAUM–I. 133544, 18.8 mm SL, Beach

74

H. Motomura and K. Uehara

Rock, Tokudoki, China; KAUM–I. 133545, 18.7 mm SL, Beach Rock, Tokudoki, China; KAUM–I. 133546,
19.8 mm SL, Beach Rock, Tokudoki, China; KAUM–I. 133547, 19.6 mm SL, Beach Rock, Tokudoki, China;
KUN-P 48923, 19.6 mm SL, Yakomo Beach, Yakomo, China; KUN-P 48924, 12.9 mm SL, Yakomo Beach,
Yakomo, China; KUN-P 48925, 12.5 mm SL, Yakomo Beach, Yakomo, China; KUN-P 48926, 16.5 mm
SL, Yakomo Beach, Yakomo, China; KUN-P 48927, 18.6 mm SL, Yakomo Beach, Yakomo, China; KUN-P
48928, 16.3 mm SL, Yakomo Beach, Yakomo, China; KUN-P 48929, 14.9 mm SL, Yakomo Beach, Yakomo,
China; KUN-P 48965, 21.3 mm SL, Ujiji Beach, Ashikiyora, China; KUN-P 49078, 19.3 mm SL, China
Port, Serikaku, China; KUN-P 49132, 20.3 mm SL, Kasaishi Beach, Kibiru, Wadomari.
Enneapterygius signicauda Fricke 1997
[Jpn name: Ogurohebigimpo]
KAUM–I. 132168, 21.5 mm SL, east of Taminazaki, Tamina, China; KAUM–I. 132169, 18.3 mm SL, east of
Taminazaki, Tamina, China; KAUM–I. 132170, 14.1 mm SL, east of Taminazaki, Tamina, China; KAUM–
I. 133488, 23.3 mm SL, Beach Rock, Tokudoki, China; KAUM–I. 133522, 27.1 mm SL, Beach Rock,Tokudoki, China.
Enneapterygius similis Fricke 1997
[Jpn name: Somewakehebigimpo]
KAUM–I. 122624, 25.3 mm SL, off Yakomo, China; KAUM–I. 133486, 20.3 mm SL, Beach Rock,Tokudoki, China; KAUM–I. 133487, 17.8 mm SL, Beach Rock, Tokudoki, China.
Enneapterygius sp.
[Jpn name: N/A]
KAUM–I. 121995, 17.2 mm SL, Masana, China; KAUM–I. 122096, 19.7 mm SL, Yakomo Beach, Yakomo,
China; KAUM–I. 122124, 23.0 mm SL, Beach Rock, Tokudoki, China; KAUM–I. 122159, 22.8 mm SL,
Beach Rock, Tokudoki, China; KAUM–I. 122160, 17.5 mm SL, Beach Rock, Tokudoki, China; KAUM–
I. 122161, 24.4 mm SL, Beach Rock, Tokudoki, China; KAUM–I. 122162, 22.4 mm SL, Beach Rock,Tokudoki, China; KAUM–I. 122163, 19.9 mm SL, Beach Rock, Tokudoki, China; KAUM–I. 122731, 23.0mm SL, Yaja, China; KUN-P 48943, 20.0 mm SL, west of Taminazaki, Tamina, China.
Enneapterygius tutuilae Jordan & Seale 1906
[Jpn name: Segurohebigimpo]
KAUM–I. 90427, 19.7 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 122623, 17.0 mm SL, off Yakomo,
China.
Enneapterygius unimaculatus Fricke 1994
[Jpn name: Sedakahebigimpo]
KAUM–I. 122095, 28.1 mm SL, Yakomo Beach, Yakomo, China; KAUM–I. 122097, 14.9 mm SL, Yakomo
Beach, Yakomo, China; KAUM–I. 122158, 28.6 mm SL, Beach Rock, Tokudoki, China.
Helcogramma aquila Williams & McCormick 1990
[Jpn name: Momijihebigimpo]
Tashiro & Motomura (2013): HMNH-P 8003, 37.3 mm SL, Nishibaru, Wadomari.
Helcogramma inclinata (Fowler 1946)[Jpn name: Ayahebigimpo]
Yoshigou et al. (2005a, as Helcogramma nesion): HMNH 7945, Okidomari, China. This study: KAUM–
I. 2002, 49.5 mm SL, off Okinoerabu Island; KAUM–I. 90081, 23.4 mm SL, Okidomari Port, Shimojiro,

Marine and freshwater fishes of Okinoerabu Island

75

China; KAUM–I. 90168, 28.0 mm SL, Okidomari Port, Shimojiro, China; KAUM–I. 122154, 38.8 mm SL,
Beach Rock, Tokudoki, China; KAUM–I. 122155, 38.1 mm SL, Beach Rock, Tokudoki, China; KAUM–
I. 122156, 31.3 mm SL, Beach Rock, Tokudoki, China; KAUM–I. 122157, 25.4 mm SL, Beach Rock,Tokudoki, China; KAUM–I. 122817, 38.9 mm SL, Sumiyoshi Port, Sumiyoshi, China; KAUM–I. 122818,
35.5 mm SL, Sumiyoshi Port, Sumiyoshi, China; KAUM–I. 123595, 36.4 mm SL, off China; KAUM–
I. 133475, 30.1 mm SL, Beach Rock, Tokudoki, China; KUN-P 49020, 22.7 mm SL, Yaja, China; KUN-P
49072, 34.0 mm SL, China Port, Serikaku, China.
Helcogramma ishigakiensis (Aoyagi 1954)[Jpn name: Benimonhebigimpo]
KAUM–I. 122338, 33.8 mm SL, north of Taminazaki, Tamina, China.
Helcogramma striata Hansen 1986
[Jpn name: Tatejimahebigimpo]
Tashiro (2018b): KAUM–I. 90144, 16.4 mm SL, Okidomari Port, Shimojiro, China. Tashiro (2019b):
KAUM–I. 90144, 16.4 mm SL, Okidomari Port, Shimojiro, China. This study: KAUM–I. 90139, 20.3 mm
SL, Okidomari Port, Shimojiro, China; KAUM–I. 90144, 16.4 mm SL, Okidomari Port, Shimojiro, China;
KAUM–I. 122308, 29.7 mm SL, west of Taminazaki, Tamina, China; KAUM–I. 122345, 26.0 mm SL, off
Okidomari Port, Shimojiro, China; KAUM–I. 122708, 25.9 mm SL, off Yakomo, China; KAUM–I. 132189,
25.7 mm SL, west of Taminazaki, Tamina, China.
Norfolkia thomasi Whitley 1964
[Jpn name: Nisehebigimpo]
KAUM–I. 122050, 31.2 mm SL, Masana, China; KAUM–I. 123572, 32.8 mm SL, Yaja, China.
FAMILY BLENNIIDAE
Alticus saliens (Forster 1788)[Jpn name: Tamakaeruo]
Kawama (2018): KAUM–I. 90556, 55.0 mm SL, Okidomari Port, Shimojiro, China. Kawama (2019):
KAUM–I. 90556, 55.0 mm SL, Okidomari Port, Shimojiro, China. This study: KAUM–I. 90555, 75.0mm SL, Okidomari Port, Shimojiro, China; KAUM–I. 90556, 55.0 mm SL, Okidomari Port, Shimojiro,
China; KAUM–I. 90557, 54.5 mm SL, Okidomari Port, Shimojiro, China; KAUM–I. 90575, 48.6 mm SL,
Okidomari Port, Shimojiro, China.
Atrosalarias holomelas (Günther 1872)[Jpn name: Indokaeruo]
KUN-P 48993, 54.9 mm SL, Ujiji Beach, Ashikiyora, China.
Blenniella chrysospilos (Bleeker 1857)[Jpn name: Montsukikaeruo]
KAUM–I. 132187, 33.8 mm SL, west of Taminazaki, Tamina, China.
Blenniella bilitonensis (Bleeker 1858)[Jpn name: Hohogurogimpo]
Yoshigou et al. (2005a): HMNH 7831, Shinjo, China; HMNH 7832, Shinjo, China; HMNH 7877, 2
specimens, Naikina, Nagamine, Wadomari. This study: KAUM–I. 122278, 17.7 mm SL, west of Taminazaki,
Tamina, China; KAUM–I. 122646, 37.1 mm SL, China Port, Serikaku, China; KAUM–I. 123414, 46.8 mm
SL, Ujiji Beach, Ashikiyora, China; KUN-P 48920, 47.4 mm SL, Yakomo Beach, Yakomo, China; KUN-P

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48989, 58.6 mm SL, Ujiji Beach, Ashikiyora, China; KUN-P 49033, 38.2 mm SL, China Port, Serikaku,
China; KUN-P 49125, 65.2 mm SL, Kasaishi Beach, Kibiru, Wadomari; KUN-P 49147, 42.1 mm SL,
Kasaishi Beach, Kibiru, Wadomari.
Blenniella periophthalmus (Valenciennes 1836)[Jpn name: Hanakaeruo]
Yoshigou et al. (2005a, as Blenniella periophthalma): HMNH 7833, Shinjo, China. Kawama (2019, as
Blenniella periophthalma): KAUM–I. 122133, 61.0 mm SL, Beach Rock, Tokudoki, China. This study:
KAUM–I. 122133, 61.0 mm SL, Beach Rock, Tokudoki, China; KAUM–I. 133507, 33.4 mm SL, Beach
Rock, Tokudoki, China; KAUM–I. 133508, 22.9 mm SL, Beach Rock, Tokudoki, China; KAUM–I. 133509,
20.1 mm SL, Beach Rock, Tokudoki, China; KUN-P 48921, 46.1 mm SL, Yakomo Beach, Yakomo, China.
Blenniidae, undetermined genus and species[Jpn name: N/A]
KAUM–I. 90315, 28.3 mm SL, Sumiyoshi Beach, Sumiyoshi, China; KAUM–I. 90325, 19.8 mm SL,
Sumiyoshi Beach, Sumiyoshi, China; KAUM–I. 90561, 9.4 mm SL, Okidomari Port, Shimojiro, China;
KAUM–I. 90562, 18.7 mm SL, Okidomari Port, Shimojiro, China.
Cirripectes castaneus (Valenciennes 1836)[Jpn name: Tategamikaeruo]
Kawama (2018): KAUM–I. 90058, 21.3 mm SL, Okidomari Port, Shimojiro, China; KAUM–I. 90075, 52.3mm SL, Okidomari Port, Shimojiro, China; KAUM–I. 90078, 34.3 mm SL, Okidomari Port, Shimojiro,
China; KAUM–I. 90742, 36.5 mm SL, Naikina Port, Nagamine, Wadomari. Kawama (2019): KAUM–
I. 90058, 21.3 mm SL, Okidomari Port, Shimojiro, China; KAUM–I. 90075, 52.3 mm SL, OkidomariPort, Shimojiro, China; KAUM–I. 90078, 34.3 mm SL, Okidomari Port, Shimojiro, China; KAUM–
I. 90742, 36.5 mm SL, Naikina Port, Nagamine, Wadomari. This study: KAUM–I. 2219, 30.1 mm SL,
off Okinoerabu Island; KAUM–I. 90014, 24.0 mm SL, Okidomari Port, Shimojiro, China; KAUM–I.
90058, 21.3 mm SL, Okidomari Port, Shimojiro, China; KAUM–I. 90071, 25.5 mm SL, Okidomari Port,
Shimojiro, China; KAUM–I. 90072, 65.2+ mm SL, Okidomari Port, Shimojiro, China; KAUM–I. 90075,
52.3 mm SL, Okidomari Port, Shimojiro, China; KAUM–I. 90078, 34.3 mm SL, Okidomari Port, Shimojiro,
China; KAUM–I. 90079, 65.7 mm SL, Okidomari Port, Shimojiro, China; KAUM–I. 90080, 34.1 mm SL,
Okidomari Port, Shimojiro, China; KAUM–I. 90742, 36.5 mm SL, Naikina Port, Nagamine, Wadomari;
KAUM–I. 122086, 53.0 mm SL, Yakomo Beach, Yakomo, China; KAUM–I. 122130, 50.5 mm SL, Beach
Rock, Tokudoki, China; KAUM–I. 122137, 25.8 mm SL, Beach Rock, Tokudoki, China; KAUM–I. 122150,
53.4 mm SL, Beach Rock, Tokudoki, China; KAUM–I. 122153, 21.6 mm SL, Beach Rock, Tokudoki, China;
KAUM–I. 122608, 49.3 mm SL, off Yakomo, China; KAUM–I. 123544, 30.5 mm SL, Yaja, China; KAUM–
I. 132188, 32.4 mm SL, north of Taminazaki, Tamina, China; KUN-P 49000, 43.0 mm SL, Yaja, China;
KUN-P 49007, 42.8 mm SL, Yaja, China; KUN-P 49014, 25.0 mm SL, Yaja, China; KUN-P 49015, 29.0 mm
SL, Yaja, China; KUN-P 49016, 24.0 mm SL, Yaja, China.
Cirripectes imitator Williams 1985
[Jpn name: Amimeminokaeruo]
Yoshigou et al. (2005a): HMNH 7838, Shinjo, China. This study: KAUM–I. 2023, 41.7 mm SL, off
Okinoerabu Island; KAUM–I. 2075, 69.8 mm SL, off Okinoerabu Island; KAUM–I. 11956, 41.8 mm SL, off
Okinoerabu Island; KAUM–I. 11957, 33.9 mm SL, off Okinoerabu Island; KAUM–I. 11958, 32.5 mm SL,
off Okinoerabu Island; KAUM–I. 11959, 36.3 mm SL, off Okinoerabu Island; KAUM–I. 11960, 36.8 mm
SL, off Okinoerabu Island; KAUM–I. 11961, 35.7 mm SL, off Okinoerabu Island; KAUM–I. 11962, 30.3mm SL, off Okinoerabu Island; KAUM–I. 11963, 32.9 mm SL, off Okinoerabu Island; KAUM–I. 11964,

Marine and freshwater fishes of Okinoerabu Island

77

32.4 mm SL, off Okinoerabu Island; KAUM–I. 11965, 35.6 mm SL, off Okinoerabu Island; KAUM–I.
11966, 33.6 mm SL, off Okinoerabu Island; KAUM–I. 11967, 31.5 mm SL, off Okinoerabu Island; KAUM–
I. 11968, 33 mm SL, off Okinoerabu Island; KAUM–I. 11969, 31.3 mm SL, off Okinoerabu Island; KAUM–
I. 11970, 29.3 mm SL, off Okinoerabu Island; KAUM–I. 11971, 29.6 mm SL, off Okinoerabu Island;
KAUM–I. 11972, 30.4 mm SL, off Okinoerabu Island; KAUM–I. 11973, 30.5 mm SL, off Okinoerabu
Island; KAUM–I. 11974, 31.1 mm SL, off Okinoerabu Island; KAUM–I. 11975, 28.8 mm SL, off
Okinoerabu Island; KAUM–I. 11976, 28.4 mm SL, off Okinoerabu Island; KAUM–I. 11977, 28.5 mm SL,
off Okinoerabu Island; KAUM–I. 13204, 76.3 mm SL, off Okinoerabu Island; KAUM–I. 13205, 72.1 mm
SL, off Okinoerabu Island; KAUM–I. 13206, 78.8 mm SL, off Okinoerabu Island; KAUM–I. 13207, 74.5mm SL, off Okinoerabu Island; KAUM–I. 13208, 79.4 mm SL, off Okinoerabu Island; KAUM–I. 13209,
70.8 mm SL, off Okinoerabu Island.
Cirripectes polyzona (Bleeker 1868)[Jpn name: Minokaeruo]
KAUM–I. 122350, 50.0 mm SL, off Okidomari Port, Shimojiro, China; KAUM–I. 122647, 35.4 mm SL,
China Port, Serikaku, China; KAUM–I. 123406, 46.1 mm SL, Ujiji Beach, Ashikiyora, China; KAUM–I.
123590, 52.2 mm SL, off China.
Cirripectes quagga (Fowler & Ball 1924)[Jpn name: Makasuminokaeruo]
Yoshigou et al. (2005a): HMNH 8000, Nishibaru, Wadomari.
Cirripectes stigmaticus Strasburg & Schultz 1953
[Jpn name: Inazumatategamikaeruuo]
Underwater photograph (Pl. 7-I).
Crossosalarias macrospilus Smith-Vaniz & Springer 1971
[Jpn name: Erigurogimpo]
Kawama (2018): KAUM–I. 90138, 26.6 mm SL, Okidomari Port, Shimojiro, China. Kawama (2019):
KAUM–I. 90138, 26.6 mm SL, Okidomari Port, Shimojiro, China. This study: KAUM–I. 90138, 26.6 mm
SL, Okidomari Port, Shimojiro, China.
Ecsenius bicolor (Day 1888)[Jpn name: Futairokaeruo]
Kawama (2018): KAUM–I. 90060, 20.4 mm SL, Okidomari Port, Shimojiro, China. Kawama (2019):
KAUM–I. 90060, 20.4 mm SL, Okidomari Port, Shimojiro, China. This study: KAUM–I. 90060, 20.4 mm
SL, Okidomari Port, Shimojiro, China; KAUM–I. 122109, 18.6 mm SL, Sumiyoshi Port, Sumiyoshi, China;
KAUM–I. 122118, 20.1 mm SL, Sumiyoshi Port, Sumiyoshi, China; KAUM–I. 122250, 54.2 mm SL,
west of Taminazaki, Tamina, China; KAUM–I. 122251, 50.5 mm SL, west of Taminazaki, Tamina, China;
KAUM–I. 122269, 52.0 mm SL, west of Taminazaki, Tamina, China; KAUM–I. 123579, 47.8 mm SL, off
China; KAUM–I. 123589, 42.4 mm SL, off China.
Ecsenius lineatus Klausewitz 1962
[Jpn name: Hitosujigimpo]
Kawama (2019): KAUM–I. 122029, 49.9 mm SL, Masana, China. This study: KAUM–I. 122029, 49.9 mm
SL, Masana, China.

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Enchelyurus kraussii (Klunzinger 1871)[Jpn name: Kurogimpo]
KAUM–I. 122775, 28.2 mm SL, Kasaishi Beach, Kibiru, Wadomari; KAUM–I. 123427, 24.5 mm SL, Ujiji
Beach, Ashikiyora, China.
Entomacrodus decussatus (Bleeker 1858)[Jpn name: Kikaikaeruuo]
KUN-P 49029, 63.8 mm SL, China Port, Serikaku, China.
Entomacrodus striatus (Valenciennes 1836)[Jpn name: Sujigimpo]
Kawama (2018): KAUM–I. 90574, 49.0 mm SL, Okidomari Port, Shimojiro, China. Kawama (2019):
KAUM–I. 90574, 49.0 mm SL, Okidomari Port, Shimojiro, China. This study: KAUM–I. 90574, 49.0 mm
SL, Okidomari Port, Shimojiro, China.
Entomacrodus thalassinus thalassinus (Jordan & Seale 1906)[Jpn name: Gotenkaeruo]
KAUM–I. 2131, 43.2 mm SL, off Okinoerabu Island; KAUM–I. 4102, 23.7 mm SL, off Okinoerabu Island;
KAUM–I. 4103, 36.6 mm SL, off Okinoerabu Island; KAUM–I. 4104, 40.6 mm SL, off Okinoerabu Island;
KAUM–I. 4105, 35.3 mm SL, off Okinoerabu Island.
Exallias brevis (Kner 1868)[Jpn name: Sedakagimpo]
Kawama (2018): KAUM–I. 90009, 32.2 mm SL, Okidomari Port, Shimojiro, China; KAUM–I. 90618, 34.8mm SL, Naikina Port, Nagamine, Wadomari. Kawama (2019): KAUM–I. 90009, 32.2 mm SL, OkidomariPort, Shimojiro, China; KAUM–I. 90618, 34.8 mm SL, Naikina Port, Nagamine, Wadomari. This study:
KAUM–I. 90009, 32.2 mm SL, Okidomari Port, Shimojiro, China; KAUM–I. 90618, 34.8 mm SL, Naikina
Port, Nagamine, Wadomari.
Istiblennius edentulus (Forster & Schneider 1801)[Jpn name: Nisekaeruo]
Yoshigou et al. (2005a): HMNH 7794, Hanjo, Furusato, Wadomari. Yoshigou (2014): Okinoerabu Island.
This study: KAUM–I. 33603, 131.9 mm SL, off Okinoerabu Island; KAUM–I. 33604, 119.3 mm SL, off
Okinoerabu Island; KAUM–I. 33605, 113.5 mm SL, off Okinoerabu Island; KAUM–I. 33606, 117.5 mm SL,
off Okinoerabu Island; KAUM–I. 33607, 105.8 mm SL, off Okinoerabu Island; KAUM–I. 33608, 100.0 mm
SL, off Okinoerabu Island; KAUM–I. 33609, 102.0 mm SL, off Okinoerabu Island; KAUM–I. 33610, 93.7mm SL, off Okinoerabu Island; KAUM–I. 33611, 90.5 mm SL, off Okinoerabu Island; KAUM–I. 90084,
99.3 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 90253, 68.2 mm SL, Sumiyoshi Beach, Sumiyoshi,
China; KAUM–I. 90254, 36.5 mm SL, Sumiyoshi Beach, Sumiyoshi, China; KAUM–I. 90255, 34.5 mm SL,
Sumiyoshi Beach, Sumiyoshi, China; KAUM–I. 90258, 20.8 mm SL, Sumiyoshi Beach, Sumiyoshi, China;
KAUM–I. 90330, 17.1 mm SL, Sumiyoshi Beach, Sumiyoshi, China; KAUM–I. 90515, 31.8 mm SL, Ujiji
Beach, Ashikiyora, China; KAUM–I. 90576, 62.7 mm SL, Okidomari Port, Shimojiro, China; KAUM–
I. 90722, 39.9 mm SL, Okidomari Port, Shimojiro, China; KAUM–I. 90723, 31.9 mm SL, OkidomariPort, Shimojiro, China; KAUM–I. 90775, 27.1 mm SL, Hanzaki, Wadomari; KAUM–I. 122631, 70.8 mm
SL, China Port, Serikaku, China; KAUM–I. 122632, 75.5 mm SL, China Port, Serikaku, China; KAUM–
I. 123429, 39.7 mm SL, Ujiji Beach, Ashikiyora, China; KAUM–I. 123430, 11.4 mm SL, Ujiji Beach,
Ashikiyora, China; KAUM–I. 132338, 19.8 mm SL, Okidomari Seaside Park, Shimojiro, China; KUN-P
48883, 74.2 mm SL, Okidomari Port, Shimojiro, China; KUN-P 48884, 64.4 mm SL, Okidomari Port,

Marine and freshwater fishes of Okinoerabu Island

79

Shimojiro, China; KUN-P 48885, 73.1 mm SL, Okidomari Port, Shimojiro, China; KUN-P 48886, 34.5 mm
SL, Okidomari Port, Shimojiro, China; KUN-P 48961, 76.0 mm SL, Ujiji Beach, Ashikiyora, China; KUN-P
48962, 56.0 mm SL, Ujiji Beach, Ashikiyora, China; KUN-P 48981, 50.2 mm SL, Ujiji Beach, Ashikiyora,
China; KUN-P 48982, 41.8 mm SL, Ujiji Beach, Ashikiyora, China; KUN-P 48983, 36.9 mm SL, Ujiji
Beach, Ashikiyora, China; KUN-P 48996, 85.8 mm SL, Ujiji Beach, Ashikiyora, China.
Istiblennius lineatus (Valenciennes 1836)[Jpn name: Senkaeruo]
Yoshigou et al. (2005a): HMNH 7795, Hanjo, Furusato, Wadomari; HMNH 7796, Hanjo, Furusato,
Wadomari; HMNH 7797, Hanjo, Furusato, Wadomari; HMNH 7837, Shinjo, China. Yoshigou (2014):
Okinoerabu Island. Kawama (2018): KAUM–I. 90250, 61.5 mm SL, Sumiyoshi Beach, Sumiyoshi, China.
Kawama (2019): KAUM–I. 90250, 61.5 mm SL, Sumiyoshi Beach, Sumiyoshi, China. This study: KAUM–
I. 90250, 61.5 mm SL, Sumiyoshi Beach, Sumiyoshi, China; KAUM–I. 90251, 32.1 mm SL, Sumiyoshi
Beach, Sumiyoshi, China; KAUM–I. 90252, 34.4 mm SL, Sumiyoshi Beach, Sumiyoshi, China; KAUM–I.
122394, 48.3 mm SL, Ujiji Beach, Ashikiyora, China; KUN-P 48997, 71.9 mm SL, Ujiji Beach, Ashikiyora,
China.
Istiblennius sp.
[Jpn name: N/A]
KAUM–I. 90331, 18.4 mm SL, Sumiyoshi Beach, Sumiyoshi, China; KAUM–I. 90332, 16.3 mm SL,
Sumiyoshi Beach, Sumiyoshi, China; KAUM–I. 90333, 16.5 mm SL, Sumiyoshi Beach, Sumiyoshi, China.
Meiacanthus kamoharai Tomiyama 1956
[Jpn name: Kamoharagimpo]
KAUM–I. 90150, 66.1 mm SL, Okidomari Port, Shimojiro, China; KAUM–I. 101640, 26.8 mm SL, Kasaishi
Beach, Kibiru, Wadomari; KUN-P 48919, 44.1 mm SL, Yakomo Beach, Yakomo, China.
Nannosalarias nativitatis (Regan 1909)[Jpn name: Hinagimpo]
Kawama (2019): KAUM–I. 122340, 24.5 mm SL, north of Taminazaki, Tamina, China. This study: KAUM–
I. 122340, 24.5 mm SL, north of Taminazaki, Tamina, China.
Omobranchus loxozonus (Jordan & Starks 1906)[Jpn name: Kumogimpo]
Kawama (2019): KAUM–I. 123426, 29.6 mm SL, Ujiji Beach, Ashikiyora, China. This study: KAUM–I.
123426, 29.6 mm SL, Ujiji Beach, Ashikiyora, China; KAUM–I. 132339, 19.8 mm SL, Okidomari Seaside
Park, Shimojiro, China.
Parenchelyurus hepburni (Snyder 1908)[Jpn name: Karasugimpo]
Kawama (2018): KAUM–I. 90692, 29.1 mm SL, Inobe Port, Inobe, Wadomari. Kawama (2019): KAUM–
I. 90692, 29.1 mm SL, Inobe Port, Inobe, Wadomari. This study: KAUM–I. 90692, 29.1 mm SL, Inobe Port,
Inobe, Wadomari.
Petroscirtes breviceps (Valenciennes 1836)[Jpn name: Nijigimpo]
KAUM–I. 36350, 40.0 mm SL, off Okinoerabu Island; KAUM–I. 36351, 35.6 mm SL, off Okinoerabu
Island; KAUM–I. 122654, 34.1 mm SL, off Naikina Port, Nagamine, Wadomari.

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Plagiotremus rhinorhynchos (Bleeker 1852)[Jpn name: Minamigimpo]
Underwater photograph (Pl. 7-J).
Plagiotremus tapeinosoma (Bleeker 1857)[Jpn name: Tenkurosujigimpo]
KAUM–I. 1989, 48.5 mm SL, off Okinoerabu Island; KAUM–I. 14277, 47.7 mm SL, off Okinoerabu Island;
KAUM–I. 14278, 44.3 mm SL, off Okinoerabu Island; KAUM–I. 14279, 45.6 mm SL, off Okinoerabu
Island; KAUM–I. 122085, 56.4 mm SL, Yakomo Beach, Yakomo, China; KUN-P 48994, 44.5 mm SL, Ujiji
Beach, Ashikiyora, China.
Praealticus bilineatus (Peters 1868)[Jpn name: Tamagimpo]
KAUM–I. 90569, 45.8 mm SL, Okidomari Port, Shimojiro, China; KAUM–I. 90570, 25.8 mm SL,
Okidomari Port, Shimojiro, China; KAUM–I. 122398, 50.0 mm SL, Ujiji Beach, Ashikiyora, China;
KAUM–I. 123413, 38.6 mm SL, Ujiji Beach, Ashikiyora, China; KAUM–I. 123418, 34.6 mm SL, Ujiji
Beach, Ashikiyora, China; KAUM–I. 123435, 35.6 mm SL, Ujiji Beach, Ashikiyora, China; KAUM–I.
123436, 34.7 mm SL, Ujiji Beach, Ashikiyora, China; KUN-P 48990, 54.5 mm SL, Ujiji Beach, Ashikiyora,
China; KUN-P 49026, 64.0 mm SL, China Port, Serikaku, China; KUN-P 49027, 76.0 mm SL, China Port,
Serikaku, China; KUN-P 49034, 42.5 mm SL, China Port, Serikaku, China; KUN-P 49035, 44.1 mm SL,
China Port, Serikaku, China; KUN-P 49031, 31.7 mm SL, China Port, Serikaku, China; KUN-P 49036, 37.7mm SL, China Port, Serikaku, China; KUN-P 49037, 41.9 mm SL, China Port, Serikaku, China; KUN-P
49038, 37.2 mm SL, China Port, Serikaku, China; KUN-P 49040, 29.1 mm SL, China Port, Serikaku, China.
Praealticus tanegasimae (Jordan & Starks 1906)[Jpn name: Tanegimpo]
Kawama (2018): KAUM–I. 90259, 52.6 mm SL, Sumiyoshi Beach, Sumiyoshi, China; KAUM–I. 90564,
67.6 mm SL, Okidomari Port, Shimojiro, China; KAUM–I. 90567, 45.7 mm SL, Okidomari Port, Shimojiro,
China. Kawama (2019): KAUM–I. 90259, 52.6 mm SL, Sumiyoshi Beach, Sumiyoshi, China; KAUM–
I. 90564, 67.6 mm SL, Okidomari Port, Shimojiro, China; KAUM–I. 90567, 45.7 mm SL, OkidomariPort, Shimojiro, China. This study: KAUM–I. 33612, 61.5 mm SL, off Okinoerabu Island; KAUM–
I. 90256, 27.4 mm SL, Sumiyoshi Beach, Sumiyoshi, China; KAUM–I. 90257, 27.4 mm SL, Sumiyoshi
Beach, Sumiyoshi, China; KAUM–I. 90259, 52.6 mm SL, Sumiyoshi Beach, Sumiyoshi, China; KAUM–
I. 90260, 51.1 mm SL, Sumiyoshi Beach, Sumiyoshi, China; KAUM–I. 90261, 59.5 mm SL, Sumiyoshi
Beach, Sumiyoshi, China; KAUM–I. 90262, 57.0 mm SL, Sumiyoshi Beach, Sumiyoshi, China; KAUM–
I. 90263, 52.9 mm SL, Sumiyoshi Beach, Sumiyoshi, China; KAUM–I. 90264, 53.7 mm SL, Sumiyoshi
Beach, Sumiyoshi, China; KAUM–I. 90265, 50.5 mm SL, Sumiyoshi Beach, Sumiyoshi, China; KAUM–
I. 90266, 49.0 mm SL, Sumiyoshi Beach, Sumiyoshi, China; KAUM–I. 90267, 49.6 mm SL, Sumiyoshi
Beach, Sumiyoshi, China; KAUM–I. 90268, 30.0 mm SL, Sumiyoshi Beach, Sumiyoshi, China; KAUM–
I. 90269, 33.6 mm SL, Sumiyoshi Beach, Sumiyoshi, China; KAUM–I. 90270, 34.3 mm SL, Sumiyoshi
Beach, Sumiyoshi, China; KAUM–I. 90271, 34.1 mm SL, Sumiyoshi Beach, Sumiyoshi, China; KAUM–
I. 90272, 30.2 mm SL, Sumiyoshi Beach, Sumiyoshi, China; KAUM–I. 90273, 23.8 mm SL, Sumiyoshi
Beach, Sumiyoshi, China; KAUM–I. 90279, 56.9 mm SL, Sumiyoshi Beach, Sumiyoshi, China; KAUM–
I. 90280, 21.1 mm SL, Sumiyoshi Beach, Sumiyoshi, China; KAUM–I. 90281, 13.7 mm SL, Sumiyoshi
Beach, Sumiyoshi, China; KAUM–I. 90282, 11.6 mm SL, Sumiyoshi Beach, Sumiyoshi, China; KAUM–
I. 90316, 28.2 mm SL, Sumiyoshi Beach, Sumiyoshi, China; KAUM–I. 90317, 27.5 mm SL, Sumiyoshi
Beach, Sumiyoshi, China; KAUM–I. 90318, 23.4 mm SL, Sumiyoshi Beach, Sumiyoshi, China; KAUM–
I. 90319, 29.4 mm SL, Sumiyoshi Beach, Sumiyoshi, China; KAUM–I. 90320, 24.4 mm SL, Sumiyoshi

Marine and freshwater fishes of Okinoerabu Island

81

Beach, Sumiyoshi, China; KAUM–I. 90321, 27.3 mm SL, Sumiyoshi Beach, Sumiyoshi, China; KAUM–
I. 90322, 30.3 mm SL, Sumiyoshi Beach, Sumiyoshi, China; KAUM–I. 90323, 20.6 mm SL, Sumiyoshi
Beach, Sumiyoshi, China; KAUM–I. 90324, 19.0 mm SL, Sumiyoshi Beach, Sumiyoshi, China; KAUM–
I. 90326, 19.5 mm SL, Sumiyoshi Beach, Sumiyoshi, China; KAUM–I. 90327, 26.4 mm SL, Sumiyoshi
Beach, Sumiyoshi, China; KAUM–I. 90328, 20.5 mm SL, Sumiyoshi Beach, Sumiyoshi, China; KAUM–I.
90329, 23.3 mm SL, Sumiyoshi Beach, Sumiyoshi, China; KAUM–I. 90563, 71.1 mm SL, Okidomari Port,
Shimojiro, China; KAUM–I. 90564, 67.6 mm SL, Okidomari Port, Shimojiro, China; KAUM–I. 90565,
58.5 mm SL, Okidomari Port, Shimojiro, China; KAUM–I. 90566, 54.6 mm SL, Okidomari Port, Shimojiro,
China; KAUM–I. 90567, 45.7 mm SL, Okidomari Port, Shimojiro, China; KAUM–I. 90572, 28.6 mm
SL, Okidomari Port, Shimojiro, China; KAUM–I. 122393, 31.0 mm SL, Ujiji Beach, Ashikiyora, China;
KAUM–I. 123437, 26.5 mm SL, Ujiji Beach, Ashikiyora, China.
Rhabdoblennius nitidus (Günther 1861)[Jpn name: Rosokugimpo]
Yoshigou et al. (2005a): HMNH 7834, Shinjo, China. This study: KAUM–I. 90773, 48.9 mm SL, Hanzaki,
Wadomari; KAUM–I. 123400, 28.5 mm SL, Ujiji Beach, Ashikiyora, China; KAUM–I. 123401, 26.1mm SL, Ujiji Beach, Ashikiyora, China; KAUM–I. 123438, 30.4 mm SL, Ujiji Beach, Ashikiyora, China;
KAUM–I. 133597, 45.7 mm SL, Beach Rock, Tokudoki, China; KAUM–I. 133598, 32.7 mm SL, Beach
Rock, Tokudoki, China; KAUM–I. 133599, 25.4 mm SL, Beach Rock, Tokudoki, China; KAUM–I. 133600,
26.5 mm SL, Beach Rock, Tokudoki, China; KAUM–I. 133601, 30.3 mm SL, Beach Rock, Tokudoki, China;
KAUM–I. 133602, 36.2 mm SL, Beach Rock, Tokudoki, China; KAUM–I. 133603, 26.3 mm SL, Beach
Rock, Tokudoki, China; KAUM–I. 133604, 33.1 mm SL, Beach Rock, Tokudoki, China; KAUM–I. 133605,
31.7 mm SL, Beach Rock, Tokudoki, China; KAUM–I. 133606, 25.5 mm SL, Beach Rock, Tokudoki, China;
KAUM–I. 133607, 42.2 mm SL, Beach Rock, Tokudoki, China; KAUM–I. 133608, 34.4 mm SL, Beach
Rock, Tokudoki, China; KAUM–I. 133609, 39.8 mm SL, Beach Rock, Tokudoki, China; KAUM–I. 133610,
37.8 mm SL, Beach Rock, Tokudoki, China; KAUM–I. 133611, 43.7 mm SL, Beach Rock, Tokudoki, China;
KAUM–I. 133612, 34.9 mm SL, Beach Rock, Tokudoki, China; KAUM–I. 133613, 40.2 mm SL, Beach
Rock, Tokudoki, China; KAUM–I. 133614, 32.2 mm SL, Beach Rock, Tokudoki, China; KAUM–I. 133615,
31.8 mm SL, Beach Rock, Tokudoki, China; KAUM–I. 133616, 36.7 mm SL, Beach Rock, Tokudoki, China;
KAUM–I. 133617, 34.1 mm SL, Beach Rock, Tokudoki, China; KAUM–I. 133618, 25.6 mm SL, Beach
Rock, Tokudoki, China; KAUM–I. 133619, 30.4 mm SL, Beach Rock, Tokudoki, China; KAUM–I. 133620,
32.3 mm SL, Beach Rock, Tokudoki, China; KAUM–I. 133621, 28.1 mm SL, Beach Rock, Tokudoki, China;
KAUM–I. 133622, 26.8 mm SL, Beach Rock, Tokudoki, China; KAUM–I. 133623, 26.5 mm SL, Beach
Rock, Tokudoki, China; KAUM–I. 133624, 23.1 mm SL, Beach Rock, Tokudoki, China; KAUM–I. 133625,
23.9 mm SL, Beach Rock, Tokudoki, China; KAUM–I. 133626, 26.2 mm SL, Beach Rock, Tokudoki, China;
KAUM–I. 133627, 22.6 mm SL, Beach Rock, Tokudoki, China; KAUM–I. 133628, 21.7 mm SL, Beach
Rock, Tokudoki, China; KAUM–I. 133629, 18.0 mm SL, Beach Rock, Tokudoki, China; KAUM–I. 133630,
29.6 mm SL, Beach Rock, Tokudoki, China; KAUM–I. 133631, 24.6 mm SL, Beach Rock, Tokudoki, China;
KAUM–I. 133632, 22.3 mm SL, Beach Rock, Tokudoki, China; KAUM–I. 133633, 30.0 mm SL, Beach
Rock, Tokudoki, China; KAUM–I. 133634, 26.1 mm SL, Beach Rock, Tokudoki, China; KAUM–I. 133635,
21.8 mm SL, Beach Rock, Tokudoki, China; KAUM–I. 133636, 22.9 mm SL, Beach Rock, Tokudoki, China;
KAUM–I. 133637, 20.7 mm SL, Beach Rock, Tokudoki, China; KAUM–I. 133638, 20.2 mm SL, Beach
Rock, Tokudoki, China; KAUM–I. 133639, 23.3 mm SL, Beach Rock, Tokudoki, China; KAUM–I. 133640,
22.3 mm SL, Beach Rock, Tokudoki, China; KUN-P 48933, 23.2 mm SL, Yakomo Beach, Yakomo, China;
KUN-P 48964, 43.9 mm SL, Ujiji Beach, Ashikiyora, China; KUN-P 48984, 41.6 mm SL, Ujiji Beach,
Ashikiyora, China; KUN-P 49032, 27.1 mm SL, China Port, Serikaku, China; KUN-P 49039, 27.0 mm SL,
China Port, Serikaku, China; KUN-P 49064, 21.6 mm SL, China Port, Serikaku, China; KUN-P 49075, 22.4mm SL, China Port, Serikaku, China; KUN-P 49076, 32.2 mm SL, China Port, Serikaku, China..

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Salarias fasciatus (Bloch 1786)[Jpn name: Yaeyamagimpo]
Kawama (2018): KAUM–I. 90706, 30.7 mm SL, Inobe Port, Inobe, Wadomari. Kawama (2019): KAUM–I.
90706, 30.7 mm SL, Inobe Port, Inobe, Wadomari. This study: KAUM–I. 2114, 62.7 mm SL, off Okinoerabu
Island; KAUM–I. 7783, 33.9 mm SL, off Okinoerabu Island; KAUM–I. 7784, 31.6 mm SL, off Okinoerabu
Island; KAUM–I. 7785, 30.5 mm SL, off Okinoerabu Island; KAUM–I. 7786, 23.9 mm SL, off Okinoerabu
Island; KAUM–I. 7787, 21.3 mm SL, off Okinoerabu Island; KAUM–I. 7788, 19.6 mm SL, off Okinoerabu
Island; KAUM–I. 7789, 20.2 mm SL, off Okinoerabu Island; KAUM–I. 90706, 30.7 mm SL, Inobe Port,
Inobe, Wadomari; KAUM–I. 90715, 47.5 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 122780, 73.4 mm
SL, Kasaishi Beach, Kibiru, Wadomari; KAUM–I. 123453, 54.6 mm SL, Kasaishi Beach, Kibiru, Wadomari;
KAUM–I. 123454, 73.6 mm SL, Kasaishi Beach, Kibiru, Wadomari; KAUM–I. 133476, 25.9 mm SL, Beach
Rock, Tokudoki, China; KAUM–I. 133477, 17.9 mm SL, Beach Rock, Tokudoki, China; KUN-P 48985,
41.3 mm SL, Ujiji Beach, Ashikiyora, China; KUN-P 48995, 76.5 mm SL, Ujiji Beach, Ashikiyora, China.
Salarias luctuosus Whitley 1929
[Jpn name: Shimagimpo]
KAUM–I. 90546, 28.7 mm SL, Ujiji Beach, Ashikiyora, China.
Salarias sinuosus Snyder 1908
[Jpn name: Hirenagakaeruo]
KAUM–I. 90568, 52.0 mm SL, Okidomari Port, Shimojiro, China; KAUM–I. 90571, 25.6 mm SL,
Okidomari Port, Shimojiro, China.
FAMILY GOBIESOCIDAE
Aspasmichthys sp.
[Jpn name: N/A]
KAUM–I. 122015, 16.4 mm SL, Masana, China.
Discotrema lineatum (Briggs 1966)[Jpn name: Tasujiumishidaubauo]
Underwater photograph (Pl. 8-A).
Lepadichthys misakius (Tanaka 1908)[Jpn name: Misakiubauo]
KAUM–I. 121912, 36.4 mm SL, Okidomari Port, Shimojiro, China.
Lepadichthys sp.
[Jpn name: Minamiubauo]
Hayashi & Hagiwara (2013): Okinoerabu Island. This study: KAUM–I. 123483, 18.3 mm SL, Kasaishi
Beach, Kibiru, Wadomari.
Pherallodus indicus (Weber 1913)[Jpn name: Hosoubauo]
Fujiwara & Motomura (2017): KAUM–I. 90045, 10.2 mm SL, Kasaishi Beach, Kibiru, Wadomari.
Motomura (2018): KAUM–I. 90045, 10.2 mm SL, Kasaishi Beach, Kibiru, Wadomari. This study: KAUM–
I. 90045, 10.2 mm SL, Kasaishi Beach, Kibiru, Wadomari; KAUM–I. 122119, 22.9 mm SL, Masana, China;
KAUM–I. 122784, 14.2 mm SL, Kasaishi Beach, Kibiru, Wadomari; KAUM–I. 122785, 16.4 mm SL,
Kasaishi Beach, Kibiru, Wadomari; KAUM–I. 122786, 19.0 mm SL, Kasaishi Beach, Kibiru, Wadomari;Marine and freshwater fishes of Okinoerabu Island

83

KAUM–I. 122787, 17.5 mm SL, Kasaishi Beach, Kibiru, Wadomari; KAUM–I. 133505, 11.2 mm SL, Beach
Rock, Tokudoki, China; KAUM–I. 133506, 10.8 mm SL, Beach Rock, Tokudoki, China; KUN-P 49154,
18.0 mm SL, Kasaishi Beach, Kibiru, Wadomari; KUN-P 49155, 20.7 mm SL, Kasaishi Beach, Kibiru,
Wadomari.
FAMILY CALLIONYMIDAE
Dactylopus dactylopus (Valenciennes 1837)[Jpn name: Ipponteguri]
Underwater photograph (Pl. 8-B).
Neosynchiropus morrisoni (Schultz 1960)[Jpn name: Sesokoteguri]
KAUM–I. 122341, 13.3 mm SL, north of Taminazaki, Tamina, China.
Neosynchiropus moyeri (Zaiser & Fricke 1985)[Jpn name: Miyaketeguri]
Iwatsubo (2019b): KAUM–I. 122325, 37.5 mm SL, north of Taminazaki, Tamina, China; KAUM–I. 122737,
17.4 mm SL, Yaja, China. This study: KAUM–I. 90148, 10.3 mm SL, Okidomari Port, Shimojiro, China;
KAUM–I. 122325, 37.5 mm SL, north of Taminazaki, Tamina, China; KAUM–I. 122737, 17.4 mm SL, Yaja,
China.
Neosynchiropus ocellatus (Pallas 1770)[Jpn name: Kowanteguri]
Yoshigou et al. (2005a): HMNH 7979, Nishibaru, Wadomari. This study: KAUM–I. 122774, 27.3 mm SL,
Kasaishi Beach, Kibiru, Wadomari; KAUM–I. 123484, 20.1 mm SL, Kasaishi Beach, Kibiru, Wadomari.
FAMILY ELEOTRIDAE
Eleotris acanthopoma Bleeker 1853
[Jpn name: Chichibumodoki]
Yoshigou et al. (2005b): Kaikodo cave, Furusato, Wadomari. Yoshigou (2014): Okinoerabu Island. This
study: KAUM–I. 122092, 40.0 mm SL, Oku River, Wadomari.
Eleotris sp.
[Jpn name: N/A]
Yoshigou et al. (2005b): Okinoerabu Island. Yoshigou (2014): Okinoerabu Island.
Remarks: This species probably represents the same species recorded as Eleotris sp. from the anchialine caveon Tokunoshima Island (Mochida & Motomura 2018), and from caves on Miyako Island and the Yaeyama
Islands (Yoshigou 2014).
FAMILY GOBIIDAE
Amblyeleotris guttata (Fowler 1938)[Jpn name: Yamabukihaze]
Underwater photograph (Pl. 8-C).
Amblyeleotris wheeleri (Polunin & Lubbock 1977)[Jpn name: Kubiakahaze]
Underwater photograph (Pl. 8-D).

84

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Amblygobius phalaena (Valenciennes 1837)[Jpn name: Sarasahaze]
KAUM–I. 2764, 58.0 mm SL, off Okinoerabu Island; KAUM–I. 90693, 42.0 mm SL, Inobe Port, Inobe,
Wadomari.
Asterropteryx semipunctata Rüppell 1830
[Jpn name: Hoshihaze]
KAUM–I. 90189, 19.6 mm SL, Yakomo Beach, Yakomo, China; KAUM–I. 90681, 28.3 mm SL, Wanjo
Beach, Azefu, Wadomari; KAUM–I. 122778, 19.2 mm SL, Kasaishi Beach, Kibiru, Wadomari; KUN-P
49159, 25.9 mm SL, Kasaishi Beach, Kibiru, Wadomari.
Bathygobius coalitus (Bennett 1832)[Jpn name: Kuroyahazuhaze]
Yoshigou et al. (2005a): HMNH 7935, Okidomari, China. Hagiwara (2018a): KAUM–I. 90215, 55.9 mm
SL, Sumiyoshi Beach, Sumiyoshi, China; KAUM–I. 90573, 51.8 mm SL, Okidomari Port, Shimojiro, China.
Hagiwara (2019): KAUM–I. 90215, 55.9 mm SL, Sumiyoshi Beach, Sumiyoshi, China; KAUM–I. 90573,
51.8 mm SL, Okidomari Port, Shimojiro, China. This study: KAUM–I. 90215, 55.9 mm SL, Sumiyoshi
Beach, Sumiyoshi, China; KAUM–I. 90217, 44.5 mm SL, Sumiyoshi Beach, Sumiyoshi, China; KAUM–I.
90218, 50.1 mm SL, Sumiyoshi Beach, Sumiyoshi, China; KAUM–I. 90219, 19.4 mm SL, Sumiyoshi Beach,
Sumiyoshi, China; KAUM–I. 90573, 51.8 mm SL, Okidomari Port, Shimojiro, China; KAUM–I. 122396,
31.1 mm SL, Ujiji Beach, Ashikiyora, China; KAUM–I. 123416, 26.6 mm SL, Ujiji Beach, Ashikiyora,
China; KAUM–I. 123417, 24.4 mm SL, Ujiji Beach, Ashikiyora, China; KAUM–I. 123434, 39.7 mm SL,
Ujiji Beach, Ashikiyora, China.
Bathygobius cocosensis (Bleeker 1854)[Jpn name: Sujikumohaze]
Akihito & Meguro (1980): LICPP 1964085, 31 mm SL, China. Yoshigou et al. (2005a): HMNH 7798, 3
specimens, Hanjo, Furusato, Wadomari. Yoshigou (2014): Okinoerabu Island. This study: KAUM–I. 123442,
23.0 mm SL, Ujiji Beach, Ashikiyora, China; KAUM–I. 133528, 30.1 mm SL, Beach Rock, Tokudoki,
China; KAUM–I. 133529, 28.1 mm SL, Beach Rock, Tokudoki, China; KAUM–I. 133530, 25.2 mm SL,
Beach Rock, Tokudoki, China; KAUM–I. 133531, 29.0 mm SL, Beach Rock, Tokudoki, China; KAUM–I.
133532, 26.6 mm SL, Beach Rock, Tokudoki, China; KUN-P 48931, 23.4 mm SL, Yakomo Beach, Yakomo,
China; KUN-P 48932, 14.1 mm SL, Yakomo Beach, Yakomo, China; KUN-P 48991, 28.4 mm SL, Ujiji
Beach, Ashikiyora, China; KUN-P 49030, 23.8 mm SL, China Port, Serikaku, China; KUN-P 49079, 23.0mm SL, China Port, Serikaku, China.
Bathygobius cyclopterus (Valenciennes 1837)[Jpn name: Yahazuhaze]
Akihito & Meguro (1980): LICPP 1962056, 35 mm SL, Wadomari; LICPP 1964086, 38 mm SL, Wadomari.
Yoshigou et al. (2005a): HMNH 7799, Hanjo, Furusato, Wadomari; HMNH 7800, Hanjo, Furusato,
Wadomari; HMNH 7801, Hanjo, Furusato, Wadomari; HMNH 7801, Hanjo, Furusato, Wadomari; HMNH
7802, Hanjo, Furusato, Wadomari; HMNH 7803, Hanjo, Furusato, Wadomari. Yoshigou (2014): Okinoerabu
Island. This study: KAUM–I. 132340, 15.9 mm SL, Okidomari Seaside Park, Shimojiro, China; KAUM–I.
132341, 11.9 mm SL, Okidomari Seaside Park, Shimojiro, China; KAUM–I. 132342, 8.0 mm SL, Okidomari
Seaside Park, Shimojiro, China; KUN-P 49073, 33.6 mm SL, China Port, Serikaku, China.
Bathygobius fuscus (Rüppell 1830)[Jpn name: Kumohaze]
Hagiwara (2018a): KAUM–I. 90695, 25.0 mm SL, Inobe Port, Inobe, Wadomari. Hagiwara (2019):

Marine and freshwater fishes of Okinoerabu Island

85

KAUM–I. 90695, 25.0 mm SL, Inobe Port, Inobe, Wadomari. This study: KAUM–I. 2007, 42.1 mm SL,
off Okinoerabu Island; KAUM–I. 14280, 40.7 mm SL, off Okinoerabu Island; KAUM–I. 14281, 38.5 mm
SL, off Okinoerabu Island; KAUM–I. 14282, 41.4 mm SL, off Okinoerabu Island; KAUM–I. 14283, 38.8mm SL, off Okinoerabu Island; KAUM–I. 14284, 36.1 mm SL, off Okinoerabu Island; KAUM–I. 14285,
31.3 mm SL, off Okinoerabu Island; KAUM–I. 14286, 26.7 mm SL, off Okinoerabu Island; KAUM–
I. 90214, 59.2 mm SL, Sumiyoshi Beach, Sumiyoshi, China; KAUM–I. 90216, 47.2 mm SL, Sumiyoshi
Beach, Sumiyoshi, China; KAUM–I. 90220, 21.2 mm SL, Sumiyoshi Beach, Sumiyoshi, China; KAUM–
I. 90296, 30.5 mm SL, Sumiyoshi Beach, Sumiyoshi, China; KAUM–I. 90297, 30.2 mm SL, Sumiyoshi
Beach, Sumiyoshi, China; KAUM–I. 90298, 32.5 mm SL, Sumiyoshi Beach, Sumiyoshi, China; KAUM–
I. 90299, 22.6 mm SL, Sumiyoshi Beach, Sumiyoshi, China; KAUM–I. 90300, 20.0 mm SL, Sumiyoshi
Beach, Sumiyoshi, China; KAUM–I. 90301, 22.7 mm SL, Sumiyoshi Beach, Sumiyoshi, China; KAUM–
I. 90302, 28.9 mm SL, Sumiyoshi Beach, Sumiyoshi, China; KAUM–I. 90303, 27.4 mm SL, Sumiyoshi
Beach, Sumiyoshi, China; KAUM–I. 90304, 24.4 mm SL, Sumiyoshi Beach, Sumiyoshi, China; KAUM–
I. 90305, 24.0 mm SL, Sumiyoshi Beach, Sumiyoshi, China; KAUM–I. 90306, 28.7 mm SL, Sumiyoshi
Beach, Sumiyoshi, China; KAUM–I. 90307, 21.0 mm SL, Sumiyoshi Beach, Sumiyoshi, China; KAUM–
I. 90308, 14.9 mm SL, Sumiyoshi Beach, Sumiyoshi, China; KAUM–I. 90309, 17.8 mm SL, Sumiyoshi
Beach, Sumiyoshi, China; KAUM–I. 90310, 16.4 mm SL, Sumiyoshi Beach, Sumiyoshi, China; KAUM–
I. 90311, 17.3 mm SL, Sumiyoshi Beach, Sumiyoshi, China; KAUM–I. 90312, 17.2 mm SL, Sumiyoshi
Beach, Sumiyoshi, China; KAUM–I. 90313, 17.8 mm SL, Sumiyoshi Beach, Sumiyoshi, China; KAUM–
I. 90314, 20.8 mm SL, Sumiyoshi Beach, Sumiyoshi, China; KAUM–I. 90695, 25.0 mm SL, Inobe Port,
Inobe, Wadomari; KAUM–I. 123432, 28.9 mm SL, Ujiji Beach, Ashikiyora, China; KAUM–I. 123433, 20.4mm SL, Ujiji Beach, Ashikiyora, China; KUN-P 48963, 60.2 mm SL, Ujiji Beach, Ashikiyora, China.
Callogobius hasseltii (Bleeker 1851)[Jpn name: Okinawahaze]
KUN-P 49156, 22.3 mm SL, Kasaishi Beach, Kibiru, Wadomari.
Callogobius okinawae (Snyder 1908)[Jpn name: Namerahaze]
Akihito & Meguro (1975): LICPP 1964066, 23 mm SL, China; LICPP 1968208, 2 specimens, 39–56 mm
SL, Okinoerabu Island. This study: KUN-P 49157, 25.3 mm SL, Kasaishi Beach, Kibiru, Wadomari.
Callogobius sclateri (Steindachner 1879)[Jpn name: Futasujihaze]
KAUM–I. 90146, 20.2 mm SL, Okidomari Port, Shimojiro, China; KAUM–I. 90440, 24.2 mm SL, Inobe
Port, Inobe, Wadomari.
Callogobius sp.
[Jpn name: N/A]
KAUM–I. 90249, 10.0 mm SL, Sumiyoshi Beach, Sumiyoshi, China.
Cryptocentrus nigrocellatus (Yanagisawa 1978)[Jpn name: Kurohoshihaze]
Underwater photograph (Pl. 8-E).
Ctenogobiops tangaroai Lubbock & Polunin 1977
[Jpn name: Hatatateshinobihaze]
Underwater photograph (Pl. 8-F).

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H. Motomura and K. Uehara

Eviota abax (Jordan & Snyder 1901)[Jpn name: Isohaze]
KUN-P 49074, 23.6 mm SL, China Port, Serikaku, China; KUN-P 49161, 23.7 mm SL, Kasaishi Beach,
Kibiru, Wadomari; KUN-P 49162, 21.0 mm SL, Kasaishi Beach, Kibiru, Wadomari; KUN-P 49164, 22.9mm SL, Kasaishi Beach, Kibiru, Wadomari.
Eviota afelei Jordan & Seale 1906
[Jpn name: Shimamidorihaze]
Fujiwara et al. (2018): KAUM–I. 123574, 9.5 mm SL, Yaja, China. Hagiwara (2019): KAUM–I. 123574, 9.5mm SL, Yaja, China. This study: KAUM–I. 123574, 9.5 mm SL, Yaja, China.
Eviota amamiko Fujiwara, Suzuki & Motomura 2019
[Jpn name: Amamikoisohaze]
Mochizuki & Matsunuma (2019): KUN-P 48948, 10.1 mm SL, west of Taminazaki, Tamina, China.
Eviota fasciola Karnella & Lachner 1981
[Jpn name: Toranokoisohaze]
KAUM–I. 132167, 11.9 mm SL, east of Taminazaki, Tamina, China.
Eviota flavipinnata Suzuki, Greenfield & Motomura 2015
[Jpn name: Kibireisohaze]
Fujiwara et al. (2018): KAUM–I. 123545, 17.8 mm SL, Yaja, China; KAUM–I. 123576, 15.5 mm SL, Yaja,
China. Hagiwara (2019): KAUM–I. 123545, 17.8 mm SL, Yaja, China. This study: KAUM–I. 123545, 17.8mm SL, Yaja, China; KAUM–I. 123576, 15.5 mm SL, Yaja, China.
Eviota guttata Lachner & Karnella 1978
[Jpn name: Shiroisohaze]
KAUM–I. 90083, 14.8 mm SL, Okidomari Port, Shimojiro, China; KAUM–I. 122022, 13.0 mm SL, Masana,
China; KAUM–I. 122348, 13.5 mm SL, off Okidomari Port, Shimojiro, China; KAUM–I. 122815, 11.7 mm
SL, Sumiyoshi Port, Sumiyoshi, China; KAUM–I. 132165, 13.8 mm SL, east of Taminazaki, Tamina, China;
KUN-P 48945, 13.9 mm SL, west of Taminazaki, Tamina, China; KUN-P 49003, 14.0 mm SL, Yaja, China;
KUN-P 49004, 15.4 mm SL, Yaja, China; KUN-P 49061, 11.7 mm SL, China Port, Serikaku, China.
Eviota japonica Jewett & Lachner 1983
[Jpn name: Minamiisohaze]
Fujiwara et al. (2019): KAUM–I. 90776, 17.3 mm SL, Hanzaki, Wadomari. This study: KAUM–I. 90516,
15.4 mm SL, Ujiji Beach, Ashikiyora, China; KAUM–I. 90525, 12.7 mm SL, Ujiji Beach, Ashikiyora,
China; KAUM–I. 90776, 17.3 mm SL, Hanzaki, Wadomari.
Eviota korechika Shibukawa & Suzuki 2005
[Jpn name: Shiboriisohaze]
KAUM–I. 122816, 11.5 mm SL, Sumiyoshi Port, Sumiyoshi, China.
Eviota melasma Lachner & Karnella 1980
[Jpn name: Akahoshiisohaze]
KUN-P 49096, 15.1 mm SL, Yakomo Beach, Yakomo, China.

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Eviota prasina (Klunzinger 1871)[Jpn name: Nan-yomidorihaze]
Yoshigou et al. (2005a): HMNH 7828, 5 specimens, Shinjo, China; HMNH 7880, 2 specimens, Naikina,
Nagamine, Wadomari; HMNH 7983, Nishibaru, Wadomari. Hagiwara (2018a): KAUM–I. 90095, 20.2mm SL, Kasaishi Beach, Kibiru, Wadomari. Hagiwara (2019): KAUM–I. 90095, 20.2 mm SL, Kasaishi
Beach, Kibiru, Wadomari. Fujiwara et al. (2019): KAUM–I. 90095, 21.0 mm SL, Kasaishi Beach, Kibiru,
Wadomari; KAUM–I. 122100, 12.3 mm SL, Yakomo Beach, Yakomo, China. This study: KAUM–I. 90095,
21.0 mm SL, Kasaishi Beach, Kibiru, Wadomari; KAUM–I. 90521, 9.5 mm SL, Ujiji Beach, Ashikiyora,
China; KAUM–I. 90777, 14.9 mm SL, Hanzaki, Wadomari; KAUM–I. 90778, 12.5 mm SL, Hanzaki,
Wadomari; KAUM–I. 122100, 12.3 mm SL, Yakomo Beach, Yakomo, China; KAUM–I. 122152, 16.4mm SL, Beach Rock, Tokudoki, China; KAUM–I. 123441, 14.7 mm SL, Ujiji Beach, Ashikiyora, China;
KAUM–I. 133490, 21.7 mm SL, Beach Rock, Tokudoki, China; KAUM–I. 133491, 17.4 mm SL, Beach
Rock, Tokudoki, China; KAUM–I. 133492, 20.0 mm SL, Beach Rock, Tokudoki, China; KAUM–I. 133493,
15.0 mm SL, Beach Rock, Tokudoki, China; KAUM–I. 133494, 16.6 mm SL, Beach Rock, Tokudoki, China;
KAUM–I. 133495, 17.1 mm SL, Beach Rock, Tokudoki, China; KAUM–I. 133496, 19.2 mm SL, Beach
Rock, Tokudoki, China; KAUM–I. 133497, 17.5 mm SL, Beach Rock, Tokudoki, China; KAUM–I. 133498,
15.1 mm SL, Beach Rock, Tokudoki, China; KAUM–I. 133499, 15.9 mm SL, Beach Rock, Tokudoki,
China; KAUM–I. 133500, 20.8 mm SL, Beach Rock, Tokudoki, China; KAUM–I. 133501, 15.6 mm SL,
Beach Rock, Tokudoki, China; KAUM–I. 133502, 17.5 mm SL, Beach Rock, Tokudoki, China; KAUM–I.
133503, 9.9 mm SL, Beach Rock, Tokudoki, China; KAUM–I. 133504, 9.7 mm SL, Beach Rock, Tokudoki,
China; KUN-P 49062, 16.7 mm SL, China Port, Serikaku, China; KUN-P 49068, 19.1 mm SL, China Port,
Serikaku, China; KUN-P 49069, 14.0 mm SL, China Port, Serikaku, China; KUN-P 49070, 17.0 mm SL,
China Port, Serikaku, China; KUN-P 49163, 17.3 mm SL, Kasaishi Beach, Kibiru, Wadomari.
Eviota smaragdus Jordan & Seale 1906
[Jpn name: Kurohoshiisohaze]
KUN-P 49081, 6.5 mm SL, China Port, Serikaku, China; KUN-P 49160, 19.1 mm SL, Kasaishi Beach,
Kibiru, Wadomari; KUN-P 49165, 16.6 mm SL, Kasaishi Beach, Kibiru, Wadomari.
Eviota toshiyuki Greenfield & Randall 2010
[Jpn name: Midorihaze]
Akihito (1969, as Eviota zonura): 24 uncataloged specimens, 16–25 mm SL, China; 18 uncataloged
specimens, 15–24 mm SL, Wadomari; 9 uncataloged specimens, 17–26 mm SL, Wadomari. This study:
KAUM–I. 2115, 24.3 mm SL, off Okinoerabu Island; KAUM–I. 14404, 21.9 mm SL, off Okinoerabu Island;
KAUM–I. 14405, 24.2 mm SL, off Okinoerabu Island; KAUM–I. 14406, 24.6 mm SL, off Okinoerabu
Island; KAUM–I. 14407, 22.8 mm SL, off Okinoerabu Island; KAUM–I. 14408, 23.1 mm SL, off
Okinoerabu Island; KAUM–I. 14409, 24.0 mm SL, off Okinoerabu Island; KAUM–I. 14410, 22.1 mm SL,
off Okinoerabu Island; KAUM–I. 14411, 22.0 mm SL, off Okinoerabu Island; KAUM–I. 14412, 24.4 mm
SL, off Okinoerabu Island; KAUM–I. 14413, 22.8 mm SL, off Okinoerabu Island; KAUM–I. 14414, 21.8mm SL, off Okinoerabu Island; KAUM–I. 14415, 21.2 mm SL, off Okinoerabu Island; KAUM–I. 14416,
21.7 mm SL, off Okinoerabu Island; KAUM–I. 14417, 18.7 mm SL, off Okinoerabu Island; KAUM–I.
14418, 19.4 mm SL, off Okinoerabu Island; KAUM–I. 14419, 19.9 mm SL, off Okinoerabu Island; KAUM–
I. 14420, 24.7 mm SL, off Okinoerabu Island; KAUM–I. 14421, 22.4 mm SL, off Okinoerabu Island;
KAUM–I. 14422, 20.4 mm SL, off Okinoerabu Island; KAUM–I. 14423, 23.0 mm SL, off Okinoerabu
Island; KAUM–I. 14424, 20.4 mm SL, off Okinoerabu Island; KAUM–I. 14425, 20.5 mm SL, off
Okinoerabu Island; KAUM–I. 14426, 19.2 mm SL, off Okinoerabu Island; KAUM–I. 14427, 21.2 mm SL,
off Okinoerabu Island; KAUM–I. 14428, 20.0 mm SL, off Okinoerabu Island; KAUM–I. 14429, 16.3 mm
SL, off Okinoerabu Island; KAUM–I. 14430, 16.2 mm SL, off Okinoerabu Island; KAUM–I. 14431, 20.6

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mm SL, off Okinoerabu Island; KAUM–I. 14432, 18.7 mm SL, off Okinoerabu Island; KAUM–I. 14433,
16.8 mm SL, off Okinoerabu Island; KAUM–I. 14434, 17.4 mm SL, off Okinoerabu Island; KAUM–I.
14435, 18.7 mm SL, off Okinoerabu Island; KAUM–I. 14436, 15.3 mm SL, off Okinoerabu Island; KAUM–
I. 14437, 17.2 mm SL, off Okinoerabu Island; KAUM–I. 14438, 16.9 mm SL, off Okinoerabu Island;
KAUM–I. 14439, 16.6 mm SL, off Okinoerabu Island; KAUM–I. 14440, 16.4 mm SL, off Okinoerabu
Island; KAUM–I. 14441, 16.1 mm SL, off Okinoerabu Island; KAUM–I. 14442, 17.2 mm SL, off
Okinoerabu Island; KAUM–I. 14443, 15.8 mm SL, off Okinoerabu Island; KAUM–I. 14444, 17.2 mm SL,
off Okinoerabu Island; KAUM–I. 14445, 13.4 mm SL, off Okinoerabu Island; KAUM–I. 14446, 14.0 mm
SL, off Okinoerabu Island; KAUM–I. 14447, 14.1 mm SL, off Okinoerabu Island; KAUM–I. 14448, 15.0mm SL, off Okinoerabu Island; KAUM–I. 14449, 14.9 mm SL, off Okinoerabu Island; KAUM–I. 14450,
16.2 mm SL, off Okinoerabu Island; KAUM–I. 14451, 12.4 mm SL, off Okinoerabu Island; KAUM–I.
14452, 12.7 mm SL, off Okinoerabu Island; KAUM–I. 14453, 16.5 mm SL, off Okinoerabu Island; KAUM–
I. 14454, 12.8 mm SL, off Okinoerabu Island; KAUM–I. 14455, 10.9 mm SL, off Okinoerabu Island;
KAUM–I. 14456, 11.2 mm SL, off Okinoerabu Island; KAUM–I. 14457, 11.3 mm SL, off Okinoerabu
Island; KAUM–I. 122052, 13.3 mm SL, Masana, China; KAUM–I. 123599, 16.0 mm SL, off China;
KAUM–I. 132166, 12.7 mm SL, east of Taminazaki, Tamina, China; KUN-P 49022, 19.5 mm SL, Yaja,
China; KUN-P 49023, 12.6 mm SL, Yaja, China; KUN-P 49117, 20.3 mm SL, Kasaishi Beach, Kibiru,
Wadomari.
Fusigobius duospilus Hoese & Reader 1985
[Jpn name: Sehoshisankakuhaze]
Yoshigou et al. (2005a): HMNH 7905, Okidomari, China. Hagiwara (2018a): KAUM–I. 90178, 18.0 mm
SL, Okidomari Port, Shimojiro, China. Hagiwara (2019): KAUM–I. 90178, 18.0 mm SL, Okidomari Port,
Shimojiro, China. This study: KAUM–I. 90145, 26.4 mm SL, Okidomari Port, Shimojiro, China; KAUM–
I. 90178, 18.0 mm SL, Okidomari Port, Shimojiro, China; KAUM–I. 122043, 27.7 mm SL, Masana, China;
KAUM–I. 122117, 33.1 mm SL, Sumiyoshi Port, Sumiyoshi, China; KAUM–I. 122295, 28.0 mm SL,
west of Taminazaki, Tamina, China; KAUM–I. 122339, 33.8 mm SL, north of Taminazaki, Tamina, China;
KAUM–I. 122602, 36.7 mm SL, off China; KAUM–I. 122603, 38.5 mm SL, off China; KAUM–I. 122615,
28.8 mm SL, off Yakomo, China; KAUM–I. 122620, 22.4 mm SL, off Yakomo, China; KAUM–I. 122627,
38.0 mm SL, off Yakomo, China; KAUM–I. 122628, 33.6 mm SL, off Yakomo, China; KAUM–I. 122700,
28.6 mm SL, off Yakomo, China; KAUM–I. 122701, 34.6 mm SL, off Yakomo, China; KAUM–I. 122717,
17.3 mm SL, off Yakomo, China; KAUM–I. 122729, 39.3 mm SL, Yaja, China; KAUM–I. 122742, 25.0 mm
SL, Yaja, China.
Fusigobius gracilis (Randall 2001)[Jpn name: Sesujisankakuhaze]
KAUM–I. 90441, 37.4 mm SL, Inobe Port, Inobe, Wadomari.
Fusigobius inframaculatus (Randall 1994)[Jpn name: Hatatatesankakuhaze]
Yoshigou et al. (2005a): HMNH 7899, Okidomari, China; HMNH 7900, Okidomari, China; HMNH 7901,
Okidomari, China; HMNH 7902, Okidomari, China; HMNH 7903, Okidomari, China.
Fusigobius neophytus (Günther 1877)[Jpn name: Sankakuhaze]
Yoshigou et al. (2005a): HMNH 7980, Nishibaru, Wadomari.

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Gnatholepis cauerensis (Bleeker 1853)[Jpn name: Kataboshiomonhaze]
Hagiwara (2018a): KAUM–I. 90398, 37.4 mm SL, Inobe Port, Inobe, Wadomari. Hagiwara (2019):
KAUM–I. 90398, 37.4 mm SL, Inobe Port, Inobe, Wadomari. This study: KAUM–I. 1889, 46.6 mm SL, off
Okinoerabu Island; KAUM–I. 10277, 42.4 mm SL, off Okinoerabu Island; KAUM–I. 10278, 41.4 mm SL,
off Okinoerabu Island; KAUM–I. 10279, 40.1 mm SL, off Okinoerabu Island; KAUM–I. 10280, 36.4 mm
SL, off Okinoerabu Island; KAUM–I. 10281, 23.7 mm SL, off Okinoerabu Island; KAUM–I. 90398, 37.4mm SL, Inobe Port, Inobe, Wadomari.
Gobiodon aoyagii Shibukawa, Suzuki & Aizawa 2013
[Jpn name: Akatenkobanhaze]
Underwater photograph (Pl. 8-G).
Gobiodon axillaris De Vis 1884
[Jpn name: Seakakobanhaze]
Underwater photograph (Pl. 8-H).
Gobiodon fuscoruber Herler, Bogorodsky & Suzuki 2013
[Jpn name: Mujikobanhaze]
KAUM–I. 90745, 28.1 mm SL, Naikina Port, Nagamine, Wadomari; KAUM–I. 90746, 25.0 mm SL, Naikina
Port, Nagamine, Wadomari.
Gobiodon prolixus Winterbottom & Harold 2005
[Jpn name: Awairokobanhaze]
Underwater photograph (Pl. 8-I).
Gobiodon quinquestrigatus (Valenciennes 1837)[Jpn name: Futairosangohaze]
KAUM–I. 90011, 19.6 mm SL, Okidomari Port, Shimojiro, China; KAUM–I. 90163, 16.3 mm SL,
Okidomari Port, Shimojiro, China; KAUM–I. 90404, 19.6 mm SL, Inobe Port, Inobe, Wadomari; KAUM–
I. 90405, 23.0 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 90442, 25.7 mm SL, Inobe Port, Inobe,
Wadomari; KAUM–I. 90623, 12.5 mm SL, Naikina Port, Nagamine, Wadomari; KAUM–I. 90624, 19.3mm SL, Naikina Port, Nagamine, Wadomari; KAUM–I. 90631, 15.0 mm SL, Naikina Port, Nagamine,
Wadomari.
Gobiodon rivulatus (Rüppell 1830)[Jpn name: Tasujikobanhaze]
KAUM–I. 90010, 23.5 mm SL, Okidomari Port, Shimojiro, China; KAUM–I. 90169, 20.0 mm SL,
Okidomari Port, Shimojiro, China; KAUM–I. 90403, 20.4 mm SL, Inobe Port, Inobe, Wadomari; KAUM–
I. 90619, 24.9 mm SL, Naikina Port, Nagamine, Wadomari; KAUM–I. 90632, 18.0 mm SL, Naikina Port,
Nagamine, Wadomari; KAUM–I. 90633, 21.8 mm SL, Naikina Port, Nagamine, Wadomari; KAUM–I.
90639, 19.6 mm SL, Naikina Port, Nagamine, Wadomari; KAUM–I. 90748, 20.1 mm SL, Naikina Port,
Nagamine, Wadomari; KAUM–I. 122334, 21.9 mm SL, north of Taminazaki, Tamina, China; KAUM–I.
122335, 21.6 mm SL, north of Taminazaki, Tamina, China; KUN-P 48944, 22.6 mm SL, west of Taminazaki,
Tamina, China.

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Gobiodon sp.
[Jpn name: Futasujikobanhaze]
KAUM–I. 122012, 21.3 mm SL, Masana, China; KAUM–I. 122013, 20.9 mm SL, Masana, China; KAUM–
I. 122255, 23.1 mm SL, west of Taminazaki, Tamina, China; KAUM–I. 122277, 12.5 mm SL, west of
Taminazaki, Tamina, China; KAUM–I. 123568, 18.2 mm SL, Yaja, China; KAUM–I. 123575, 8.7 mm SL,
Yaja, China; KUN-P 49017, 20.5 mm SL, Yaja, China; KUN-P 49018, 20.1 mm SL, Yaja, China; KUN-P
49115, 23.3 mm SL, Yakomo Beach, Yakomo, China.
Gobiopsis arenaria (Snyder 1908)[Jpn name: Isagohaze]
Yoshigou et al. (2005a): HMNH 7829, Shinjo, China.
Kelloggella cardinalis Jordan & Seale 1906
[Jpn name: Akahirehadakahaze]
KAUM–I. 122312, 13.0 mm SL, Kasaishi Beach, Kibiru, Wadomari; KAUM–I. 122313, 15.0 mm SL,
Kasaishi Beach, Kibiru, Wadomari; KAUM–I. 122314, 12.4 mm SL, Kasaishi Beach, Kibiru, Wadomari;
KAUM–I. 122650, 23.7 mm SL, China Port, Serikaku, China; KUN-P 48966, 25.0 mm SL, Kasaishi Beach,
Kibiru, Wadomari; KUN-P 48969, 8.6 mm SL, Kasaishi Beach, Kibiru, Wadomari; KUN-P 48970, 13.2 mm
SL, Kasaishi Beach, Kibiru, Wadomari; KUN-P 48971, 15.0 mm SL, Kasaishi Beach, Kibiru, Wadomari;
KUN-P 48972, 12.0 mm SL, Kasaishi Beach, Kibiru, Wadomari; KUN-P 48973, 14.0 mm SL, Kasaishi
Beach, Kibiru, Wadomari; KUN-P 48974, 10.5 mm SL, Kasaishi Beach, Kibiru, Wadomari; KUN-P 48975,
11.2 mm SL, Kasaishi Beach, Kibiru, Wadomari; KUN-P 48976, 10.3 mm SL, Kasaishi Beach, Kibiru,
Wadomari; KUN-P 48977, 10.1 mm SL, Kasaishi Beach, Kibiru, Wadomari; KUN-P 48978, 9.1 mm SL,
Kasaishi Beach, Kibiru, Wadomari; KUN-P 49045, 23.0 mm SL, China Port, Serikaku, China; KUN-P
49046, 22.1 mm SL, China Port, Serikaku, China; KUN-P 49047, 17.8 mm SL, China Port, Serikaku,
China; KUN-P 49048, 18.8 mm SL, China Port, Serikaku, China; KUN-P 49049, 17.8 mm SL, China Port,
Serikaku, China; KUN-P 49050, 13.8 mm SL, China Port, Serikaku, China; KUN-P 49051, 14.5 mm SL,
China Port, Serikaku, China; KUN-P 49052, 14.0 mm SL, China Port, Serikaku, China; KUN-P 49053, 13.2mm SL, China Port, Serikaku, China; KUN-P 49054, 11.5 mm SL, China Port, Serikaku, China; KUN-P
49055, 12.1 mm SL, China Port, Serikaku, China; KUN-P 49056, 10.5 mm SL, China Port, Serikaku, China;
KUN-P 49057, 9.6 mm SL, China Port, Serikaku, China; KUN-P 49058, 10.0 mm SL, China Port, Serikaku,
China; KUN-P 49059, 8.9 mm SL, China Port, Serikaku, China; KUN-P 49060, 7.5 mm SL, China Port,
Serikaku, China.
Lotilia klausewitzi Shibukawa, Suzuki & Senou 2012
[Jpn name: Odorihaze]
Underwater photograph (Pl. 8-J).
Paragobiodon kasaii Suzuki & Randall 2011
[Jpn name: Kasaidarumahaze]
Hagiwara (2018a): KAUM–I. 90756, 16.2 mm SL, Naikina Port, Nagamine, Wadomari. Hagiwara (2019):
KAUM–I. 90756, 16.2 mm SL, Naikina Port, Nagamine, Wadomari. This study: KAUM–I. 90756, 16.2mm SL, Naikina Port, Nagamine, Wadomari; KAUM–I. 90757, 16.6 mm SL, Naikina Port, Nagamine,
Wadomari.
Paragobiodon lacunicolus (Kendall & Goldsborough 1911)[Jpn name: Pandadarumahaze]
KAUM–I. 90747, 12.9 mm SL, Naikina Port, Nagamine, Wadomari.

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Paragobiodon modestus (Regan 1908)[Jpn name: Yogoredarumahaze]
Hagiwara (2018a): KAUM–I. 90625, 18.8 mm SL, Naikina Port, Nagamine, Wadomari. Hagiwara (2019):
KAUM–I. 90625, 18.8 mm SL, Naikina Port, Nagamine, Wadomari. This study: KAUM–I. 90625, 18.8mm SL, Naikina Port, Nagamine, Wadomari; KAUM–I. 90634, 18.3 mm SL, Naikina Port, Nagamine,
Wadomari.
Pleurosicya coerulea Larson 1990
[Jpn name: Hiraumitakehaze]
Wada et al. (2018): KAUM–I. 122102, 10.4 mm SL, Kasaishi Beach, Kibiru, Wadomari; KAUM–I. 122106,
13.2 mm SL, Kasaishi Beach, Kibiru, Wadomari. Hagiwara (2019): KAUM–I. 122102, 10.3 mm SL,
Kasaishi Beach, Kibiru, Wadomari. This study KAUM–I. 122102, 10.4 mm SL, Kasaishi Beach, Kibiru,
Wadomari; KAUM–I. 122106, 13.2 mm SL, Kasaishi Beach, Kibiru, Wadomari.
Pleurosicya mossambica Smith 1959
[Jpn name: Seboshiumitakehaze]
Yoshigou et al. (2005a): HMNH 7981, Nishibaru, Wadomari.
Priolepis cincta (Regan 1908)[Jpn name: Benkeihaze]
Akihito (1969, as Quisquilius naraharai): 3 uncataloged specimens, 27–41 mm SL, Wadomari. Yoshigou
et al. (2005a): HMNH 7825, 5 specimens, Shinjo, China; HMNH 7878, Naikina, Nagamine, Wadomari;
HMNH 7906, Okidomari, China; HMNH 7933, 2 specimens, Okidomari, China. This study: KAUM–
I. 90073, 18.0 mm SL, Okidomari Port, Shimojiro, China; KAUM–I. 90074, 17.0 mm SL, OkidomariPort, Shimojiro, China; KAUM–I. 122010, 24.5 mm SL, Masana, China; KAUM–I. 122031, 28.7 mm SL,
Masana, China; KAUM–I. 122114, 24.2 mm SL, Sumiyoshi Port, Sumiyoshi, China; KAUM–I. 122271,
32.9 mm SL, west of Taminazaki, Tamina, China; KAUM–I. 122272, 20.3 mm SL, west of Taminazaki,
Tamina, China; KAUM–I. 122273, 9.9 mm SL, west of Taminazaki, Tamina, China; KAUM–I. 122275, 20.5mm SL, west of Taminazaki, Tamina, China; KAUM–I. 122276, 20.5 mm SL, west of Taminazaki, Tamina,
China; KAUM–I. 122279, 13.1 mm SL, west of Taminazaki, Tamina, China; KAUM–I. 122600, 17.6mm SL, off China; KAUM–I. 122621, 15.0 mm SL, off Yakomo, China; KAUM–I. 122758, 25.4 mm SL,
Kasaishi Beach, Kibiru, Wadomari; KAUM–I. 132175, 19.1 mm SL, north of Taminazaki, Tamina, China;
KAUM–I. 132176, 16.0 mm SL, west of Taminazaki, Tamina, China; KAUM–I. 132177, 15.3 mm SL, westof Taminazaki, Tamina, China; KUN-P 48897, 23.3 mm SL, off Masana, China; KUN-P 48942, 27.5 mm SL,
west of Taminazaki, Tamina, China; KUN-P 49126, 30.3 mm SL, Kasaishi Beach, Kibiru, Wadomari.
Priolepis fallacincta Winterbottom & Burridge 1992
[Jpn name: Kobenkeihaze]
KAUM–I. 90771, 11.2 mm SL, Naikina Port, Nagamine, Wadomari.
Priolepis inhaca (Smith 1949)[Jpn name: Amimebenkeihaze]
Hagiwara (2019): KAUM–I. 122622, 19.7 mm SL, off Yakomo, China. This study: KAUM–I. 122622, 19.7mm SL, off Yakomo, China.
Priolepis latifascima Winterbottom & Burridge 1993
[Jpn name: Futosujiirezumihaze]
Yoshigou et al. (2005a): HMNH 7827, Shinjo, China. This study: KAUM–I. 90508, 23.6 mm SL, Ujiji

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Beach, Ashikiyora, China; KAUM–I. 121886, 16.6 mm SL, Okidomari Port, Shimojiro, China; KAUM–I.
121909, 20.9 mm SL, Okidomari Port, Shimojiro, China; KAUM–I. 121910, 10.6 mm SL, Okidomari Port,
Shimojiro, China; KAUM–I. 122601, 11.0 mm SL, off China; KAUM–I. 123439, 11.4 mm SL, Ujiji Beach,
Ashikiyora, China; KAUM–I. 132174, 17.6 mm SL, north of Taminazaki, Tamina, China; KAUM–I. 133483,
15.7 mm SL, Beach Rock, Tokudoki, China; KAUM–I. 133484, 14.3 mm SL, Beach Rock, Tokudoki, China;
KAUM–I. 133485, 15.8 mm SL, Beach Rock, Tokudoki, China.
Priolepis semidoliata (Valenciennes 1837)[Jpn name: Irezumihaze]
Akihito (1969, as Zonogobius semidoliatus): 7 uncataloged specimens, 16–39 mm SL, Wadomari. Yoshigou
et al. (2005a): HMNH 7826, 9 specimens, Shinjo, China; HMNH 7879, Naikina, Nagamine, Wadomari;
HMNH 7934, Okidomari, China. This study: KAUM–I. 90149, 16.3 mm SL, Okidomari Port, Shimojiro,
China; KAUM–I. 90510, 20.9 mm SL, Ujiji Beach, Ashikiyora, China; KAUM–I. 90524, 6.8 mm SL, Ujiji
Beach, Ashikiyora, China; KAUM–I. 90683, 11.9 mm SL, Wanjo Beach, Azefu, Wadomari; KAUM–I.
122172, 12.5 mm SL, Beach Rock, Tokudoki, China; KAUM–I. 122274, 17.8 mm SL, west of Taminazaki,
Tamina, China; KUN-P 48922, 22.2 mm SL, Yakomo Beach, Yakomo, China; KUN-P 49063, 16.8 mm
SL, China Port, Serikaku, China; KUN-P 49094, 12.1 mm SL, Kasaishi Beach, Kibiru, Wadomari; KUN-P
49108, 16.3 mm SL, Yakomo Beach, Yakomo, China; KUN-P 49127, 25.8 mm SL, Kasaishi Beach, Kibiru,
Wadomari; KUN-P 49128, 21.1 mm SL, Kasaishi Beach, Kibiru, Wadomari; KUN-P 49129, 16.3 mm SL,
Kasaishi Beach, Kibiru, Wadomari.
Rhinogobius brunneus (Temminck & Schlegel 1845)[Jpn name: Kuroyoshinobori]Shibata (1964): OMNH-P 1138–1144, 7 specimens, channel from Amata River, Kamihirakawa, China.
Yoshigou (2014): Okinoerabu Island. Hagiwara (2018a): KAUM–I. 6604, 48.6 mm SL, Amata River, Amata,China. Hagiwara (2019): KAUM–I. 6604, 48.6 mm SL, Amata River, Amata, China. This study: KAUM–
I. 6604, 48.6 mm SL, Amata River, Amata, China; KAUM–I. 122024, 35.1 mm SL, Oku River, Wadomari;
KAUM–I. 122025, 42.3 mm SL, Oku River, Wadomari; KAUM–I. 122026, 33.7 mm SL, Oku River,
Wadomari.
Rhinogobius sp. YB[Jpn name: Kibarayoshinobori]Kishino & Yonezawa (2003): Okinoerabu Island. Tachihara (2009): Okinoerabu Island. Yoshigou (2014):
Okinoerabu Island. Nakajima & Kano (2014): Amata River, Amata, China. This study: KAUM–I. 122023,
39.4 mm SL, Oku River, Wadomari; KUN-P 48910, 49.6 mm SL, Oku River, Wadomari; KUN-P 48911,
54.3 mm SL, Oku River, Wadomari.
Sicyopterus japonicus (Tanaka 1909)[Jpn name: Bozuhaze]
Yoshigou (2014): Okinoerabu Island. Hagiwara (2018a): KAUM–I. 6603, 55.8 mm SL, Amata River, Amata,China. Hagiwara (2019): KAUM–I. 6603, 55.8 mm SL, Amata River, Amata, China. This study: KAUM–I.
6603, 55.8 mm SL, Amata River, Amata, China.
Stonogobiops yasha Yoshino & Shimada 2001
[Jpn name: Yashahaze]
Underwater photograph (Pl. 8-K).

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Trimma caesiura Jordan & Seale 1906
[Jpn name: Benihaze]
Yoshigou et al. (2005a): HMNH 7907, 4 specimens, Okidomari, China; HMNH 7982, Nishibaru, Wadomari.
This study: KAUM–I. 122049, 14.9 mm SL, Masana, China; KAUM–I. 122165, 25.6 mm SL, Beach Rock,Tokudoki, China; KAUM–I. 122812, 21.8 mm SL, Sumiyoshi Port, Sumiyoshi, China; KAUM–I. 122813,
15.6 mm SL, Sumiyoshi Port, Sumiyoshi, China; KUN-P 48968, 25.8 mm SL, Ujiji Beach, Ashikiyora,
China.
Trimma caudomaculatum Yoshino & Araga 1975
[Jpn name: Aogihaze]
KAUM–I. 90621, 8.3 mm SL, Naikina Port, Nagamine, Wadomari; KAUM–I. 90750, 27.5 mm SL, Naikina
Port, Nagamine, Wadomari; KAUM–I. 90751, 25.4 mm SL, Naikina Port, Nagamine, Wadomari; KAUM–I.
122610, 20.4 mm SL, off Yakomo, China.
Trimma macrophthalmum (Tomiyama 1936)[Jpn name: Omehaze]
Hagiwara (2019): KAUM–I. 122286, 17.0 mm SL, west of Taminazaki, Tamina, China. This study: KAUM–
I. 122286, 17.0 mm SL, west of Taminazaki, Tamina, China.
Trimma maiandros Hoese, Winterbottom & Reader 2011
[Jpn name: Aobenihaze]
KAUM–I. 90763, 20.2 mm SL, Naikina Port, Nagamine, Wadomari; KAUM–I. 122814, 12.7 mm SL,
Sumiyoshi Port, Sumiyoshi, China.
Trimma okinawae (Aoyagi 1949)[Jpn name: Okinawabenihaze]
KAUM–I. 90136, 13.7 mm SL, Okidomari Port, Shimojiro, China; KAUM–I. 90172, 21.9 mm SL,
Okidomari Port, Shimojiro, China; KAUM–I. 90177, 11.5 mm SL, Okidomari Port, Shimojiro, China;
KAUM–I. 90439, 17.7 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 122374, 19.6 mm SL, off
Okidomari Port, Shimojiro, China; KAUM–I. 122728, 13.3 mm SL, Yaja, China; KAUM–I. 122740, 18.7mm SL, Yaja, China; KAUM–I. 122741, 16.5 mm SL, Yaja, China; KAUM–I. 123565, 12.2 mm SL, Yaja,
China; KAUM–I. 132178, 14.6 mm SL, west of Taminazaki, Tamina, China.
Trimmatom pharus Winterbottom 2001
[Jpn name: Sujishimaisohaze]
KAUM–I. 123598, 12.5 mm SL, off China.
Valenciennea longipinnis (Lay & Bennett 1839)[Jpn name: Sazanamihaze]
Akihito (1969, as Eleotriodes longipinnis): uncataloged, 141 mm SL, Okinoerabu Island.
Valenciennea puellaris (Tomiyama 1956)[Jpn name: Otomehaze]
Hagiwara (2019): KAUM–I. 122347, 73.6 mm SL, off Okidomari Port, Shimojiro, China. This study:
KAUM–I. 122347, 73.6 mm SL, off Okidomari Port, Shimojiro, China.

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Valenciennea strigata (Broussonet 1782)[Jpn name: Akahachihaze]
Akihito (1969, as Eleotriodes strigatus): MIKU 1791, 43 mm SL, Wadomari. Hagiwara (2019): KAUM–
I. 122697, 72.3 mm SL, off Yakomo, China. This study: KAUM–I. 90100, 60.3 mm SL, Okidomari Port,
Shimojiro, China; KAUM–I. 122695, 89.4 mm SL, off Yakomo, China; KAUM–I. 122696, 71.3 mm SL, off
Yakomo, China; KAUM–I. 122697, 72.3 mm SL, off Yakomo, China; KAUM–I. 123596, 83.1 mm SL, off
China.
FAMILY KRAEMERIIDAE
Kraemeria cunicularia Rofen 1958
[Jpn name: Ryukyunaminokohaze]
Yoshigou et al. (2005a): HMNH 7804, 6 specimens, Hanjo, Furusato, Wadomari. Yoshigou (2014):
Okinoerabu Island.
FAMILY MICRODESMIDAE
Gunnellichthys curiosus Dawson 1968
[Jpn name: Nishikioomewarasubo]
Underwater photograph (Pl. 9-A).
FAMILY PTERELEOTRIDAE
Nemateleotris decora Randall & Allen 1973
[Jpn name: Akebonohaze]
Underwater photograph (Pl. 9-B).
Nemateleotris magnifica Fowler 1938
[Jpn name: Hatatatehaze]
Hagiwara (2018b): KAUM–I. 90061, 26.0 mm SL, Okidomari Port, Shimojiro, China. This study: KAUM–
I. 90061, 26.0 mm SL, Okidomari Port, Shimojiro, China; KAUM–I. 90062, 18.2 mm SL, Okidomari Port,
Shimojiro, China; KAUM–I. 90064, 18.6 mm SL, Okidomari Port, Shimojiro, China; KAUM–I. 90378, 29.0mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 121997, 32.4 mm SL, Masana, China; KAUM–I. 122739,
26.5 mm SL, Yaja, China; KUN-P 48896, 20.7 mm SL, off Masana, China; KUN-P 48941, 34.9 mm SL,
west of Taminazaki, Tamina, China.
Nemateleotris helfrichi Randall & Allen 1973
[Jpn name: Shikonhatatatehaze]
Underwater photograph (Pl. 9-C).
Parioglossus formosus (Smith 1931)[Jpn name: Yosujihaze]
Yoshigou et al. (2005a): HMNH 7824, 9 specimens, Shinjo, China. Akihito et al. (2013): Okinoerabu Island.
Yoshigou (2014): Okinoerabu Island.
Ptereleotris zebra (Fowler 1938)[Jpn name: Zeburahaze]
KUN-P 49111, 74.3 mm SL, Yakomo Beach, Yakomo, China; KUN-P 49112, 45.8 mm SL, Yakomo Beach,
Yakomo, China.

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FAMILY EPHIPPIDAE
Platax boersii Bleeker 1853
[Jpn name: Mikazukitsubameuo]
Underwater photograph (Pl. 9-D).
Platax orbicularis (Forsskål 1775)[Jpn name: Nan-youtsubameuo]
Underwater photograph (Pl. 9-E).
Platax teira (Forsskål 1775)[Jpn name: Tsubameuo]
Underwater photograph (Pl. 9-F).
FAMILY SIGANIDAE
Siganus argenteus (Quoy & Gaimard 1825)[Jpn name: Hanaaigo]
KAUM–I. 121861, 82.5 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 121862, 86.0 mm SL, Inobe Port,
Inobe, Wadomari.
Siganus fuscescens (Houttuyn 1782)[Jpn name: Aigo]
KAUM–I. 36337, 108.7 mm SL, off Okinoerabu Island; KAUM–I. 36338, 94.5 mm SL, off Okinoerabu
Island; KAUM–I. 36339, 101.9 mm SL, off Okinoerabu Island; KAUM–I. 36340, 98.8 mm SL, off
Okinoerabu Island; KAUM–I. 36341, 95.5 mm SL, off Okinoerabu Island; KAUM–I. 90185, 34.1 mm SL,
Yakomo Beach, Yakomo, China.
Siganus spinus (Linnaeus 1758)[Jpn name: Amiaigo]
Yoshigou et al. (2005a): HMNH 7843, Shinjo, China. This study: KAUM–I. 121747, 194.0 mm SL,
off Wadomari; KAUM–I. 121748, 165.6 mm SL, off Wadomari; KAUM–I. 121749, 178.6 mm SL, off
Wadomari.
FAMILY ZANCLIDAE
Zanclus cornutus (Linnaeus 1758)[Jpn name: Tsunodashi]
KAUM–I. 36324, 80.1 mm SL, off Okinoerabu Island; KUN-P 48904, 129.5 mm SL, Beach Rock, Tokudoki,
China.
FAMILY ACANTHURIDAE
Acanthurus achilles Shaw 1803
[Jpn name: Akatsukihagi]
Underwater photograph (Pl. 9-G).
Remarks: The underwater photograph represents the first record of A. achilles from the Amami Islands andthe northernmost record for the species. Only two individuals were found in Okinoerabu Island; the two havebeen observed by the second author around 3 m depth off Cape Tamina for eight years and no additional
individuals have been found from the island.

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Acanthurus dussumieri Valenciennes 1835
[Jpn name: Nisekanranhagi]
KAUM–I. 90578, 309.6 mm SL, Okidomari Port, Shimojiro, China; KAUM–I. 121874, 310.4 mm SL, Inobe
Port, Inobe, Wadomari.
Acanthurus japonicus (Schmidt 1931)[Jpn name: Namidakurohagi]
Sakurai (2018c): KAUM–I. 90036, 144.3 mm SL, Okidomari Port, Shimojiro, China. Sakurai (2019c):
KAUM–I. 90036, 144.3 mm SL, Okidomari Port, Shimojiro, China. This study: KAUM–I. 90036, 144.3 mm
SL, Okidomari Port, Shimojiro, China.
Acanthurus lineatus (Linnaeus 1758)[Jpn name: Nijihagi]
KAUM–I. 90584, 183.0 mm SL, Okidomari Port, Shimojiro, China; KAUM–I. 122390, 41.9 mm SL, Ujiji
Beach, Ashikiyora, China; KAUM–I. 133523, 42.9 mm SL, Beach Rock, Tokudoki, China.
Acanthurus nigrofuscus (Forsskål 1775)[Jpn name: Naganiza]
KAUM–I. 2065, 45.7 mm SL, off Okinoerabu Island; KAUM–I. 3792, 42.7 mm SL, off Okinoerabu Island;
KAUM–I. 3793, 43.1 mm SL, off Okinoerabu Island; KAUM–I. 3794, 45.0 mm SL, off Okinoerabu Island;
KAUM–I. 3795, 40.2 mm SL, off Okinoerabu Island KAUM–I. 3796, 41.5 mm SL, off Okinoerabu Island;
KAUM–I. 3797, 44.1 mm SL, off Okinoerabu Island; KAUM–I. 3798, 43.1 mm SL, off Okinoerabu Island;
KAUM–I. 3799, 42.3 mm SL, off Okinoerabu Island; KAUM–I. 3800, 38.1 mm SL, off Okinoerabu Island;
KAUM–I. 3801, 37.8 mm SL, off Okinoerabu Island; KAUM–I. 3802, U mm SL, off Okinoerabu Island;
KAUM–I. 3803, 39.5 mm SL, off Okinoerabu Island; KAUM–I. 3804, 37.3 mm SL, off Okinoerabu Island;
KAUM–I. 3805, 38.2 mm SL, off Okinoerabu Island; KAUM–I. 3806, 42.7 mm SL, off Okinoerabu Island;
KAUM–I. 3807, 44.2 mm SL, off Okinoerabu Island; KAUM–I. 3808, 37.8 mm SL, off Okinoerabu Island;
KAUM–I. 3809, 40.8 mm SL, off Okinoerabu Island; KAUM–I. 3810, 41.6 mm SL, off Okinoerabu Island;
KAUM–I. 3811, 35.5 mm SL, off Okinoerabu Island; KAUM–I. 3812, 37.5 mm SL, off Okinoerabu Island;
KAUM–I. 3813, 39.8 mm SL, off Okinoerabu Island; KAUM–I. 3814, 41.8 mm SL, off Okinoerabu Island;
KAUM–I. 3815, 38.6 mm SL, off Okinoerabu Island; KAUM–I. 3816, 39.7 mm SL, off Okinoerabu Island;
KAUM–I. 3817, 37.9 mm SL, off Okinoerabu Island; KAUM–I. 3818, 40.0 mm SL, off Okinoerabu Island;
KAUM–I. 3819, 41.5 mm SL, off Okinoerabu Island; KAUM–I. 3820, 42.0 mm SL, off Okinoerabu Island;
KAUM–I. 3821, 42.9 mm SL, off Okinoerabu Island; KAUM–I. 121880, 97.9 mm SL, Okidomari Port,
Shimojiro, China; KAUM–I. 122030, 35.9 mm SL, Masana, China; KAUM–I. 122751, 61.6 mm SL,
Kasaishi Beach, Kibiru, Wadomari; KAUM–I. 123460, 68.4 mm SL, Kasaishi Beach, Kibiru, Wadomari;
KAUM–I. 123461, 52.7 mm SL, Kasaishi Beach, Kibiru, Wadomari; KAUM–I. 123462, 41.1 mm SL,
Kasaishi Beach, Kibiru, Wadomari; KAUM–I. 123593, 40.2 mm SL, off China; KAUM–I. 123594, 50.7 mm
SL, off China.
Acanthurus olivaceus Bloch & Schneider 1801
[Jpn name: Montsukihagi]
KUN-P 48914, 35.0 mm SL, Yakomo Beach, Yakomo, China; KUN-P 49105, 21.7 mm SL, Yakomo Beach,
Yakomo, China; KUN-P 49106, 22.7 mm SL, Yakomo Beach, Yakomo, China.
Acanthurus pyroferus Kittlitz 1834
[Jpn name: Kuroguchiniza]
KAUM–I. 90154, 177.2 mm SL, Okidomari Port, Shimojiro, China.

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97

Acanthurus sp.
[Jpn name: N/A]
KAUM–I. 90717, 28.2 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 122021, 27.4 mm SL, Masana,
China.
Acanthurus thompsoni (Fowler 1923)[Jpn name: Ohagurohagi]
Sakurai (2018c): KAUM–I. 90336, 152.8 mm SL, off Okidomari Port, Shimojiro, China. Sakurai (2019c):
KAUM–I. 90336, 152.8 mm SL, off Okidomari Port, Shimojiro, China. This study: KAUM–I. 90336,
152.8 mm SL, off Okidomari Port, Shimojiro, China; KAUM–I. 90337, 171.7 mm SL, off Okidomari Port,
Shimojiro, China; KAUM–I. 90338, 148.4 mm SL, off Okidomari Port, Shimojiro, China; KAUM–I. 90356,
91.1 mm SL, off Okidomari Port, Shimojiro, China; KAUM–I. 90359, 140.1 mm SL, off Okidomari Port,
Shimojiro, China; KAUM–I. 90360, 129.7 mm SL, off Okidomari Port, Shimojiro, China; KAUM–I. 90361,
136.7 mm SL, off Okidomari Port, Shimojiro, China; KAUM–I. 90362, 146.8 mm SL, off Okidomari Port,
Shimojiro, China; KAUM–I. 90363, 135.3 mm SL, off Okidomari Port, Shimojiro, China.
Acanthurus triostegus (Linnaeus 1758)[Jpn name: Shimahagi]
Yoshigou et al. (2005a): HMNH 7882, Naikina, Nagamine, Wadomari. This study: KAUM–I. 36330, 83.9mm SL, off Okinoerabu Island; KAUM–I. 36331, 78.5 mm SL, off Okinoerabu Island; KAUM–I. 36332,
71.8 mm SL, off Okinoerabu Island; KAUM–I. 36333, 67.4 mm SL, off Okinoerabu Island; KAUM–I.
36334, 65.2 mm SL, off Okinoerabu Island; KAUM–I. 36335, 65.6 mm SL, off Okinoerabu Island; KAUM–
I. 36336, 55.1 mm SL, off Okinoerabu Island; KAUM–I. 90225, 22.5 mm SL, Sumiyoshi Beach, Sumiyoshi,
China; KAUM–I. 90226, 22.2 mm SL, Sumiyoshi Beach, Sumiyoshi, China; KAUM–I. 90227, 22.1 mm
SL, Sumiyoshi Beach, Sumiyoshi, China; KAUM–I. 90228, 22.5 mm SL, Sumiyoshi Beach, Sumiyoshi,
China; KAUM–I. 90276, 25.4 mm SL, Sumiyoshi Beach, Sumiyoshi, China; KAUM–I. 90277, 25.7 mm SL,
Sumiyoshi Beach, Sumiyoshi, China; KAUM–I. 90278, 24.6 mm SL, Sumiyoshi Beach, Sumiyoshi, China;
KAUM–I. 121765, 75.2 mm SL, Bishi Beach, Kunigami, Wadomari; KAUM–I. 122138, 22.5 mm SL, Beach
Rock, Tokudoki, China; KAUM–I. 122139, 24.0 mm SL, Beach Rock, Tokudoki, China; KAUM–I. 122385,
38.3 mm SL, Ujiji Beach, Ashikiyora, China; KAUM–I. 122391, 35.0 mm SL, Ujiji Beach, Ashikiyora,
China; KAUM–I. 122651, 22.9 mm SL, China Port, Serikaku, China; KAUM–I. 123420, 23.7 mm SL, Ujiji
Beach, Ashikiyora, China; KUN-P 48905, 141.0 mm SL, Beach Rock, Tokudoki, China; KUN-P 48950,
39.0 mm SL, Ujiji Beach, Ashikiyora, China; KUN-P 48951, 38.0 mm SL, Ujiji Beach, Ashikiyora, China;
KUN-P 48952, 38.0 mm SL, Ujiji Beach, Ashikiyora, China; KUN-P 49041, 30.8 mm SL, China Port,
Serikaku, China; KUN-P 49042, 23.1 mm SL, China Port, Serikaku, China.
Ctenochaetus binotatus Randall 1955
[Jpn name: Kokutensazanamihagi]
KAUM–I. 90065, 34.0 mm SL, Okidomari Port, Shimojiro, China; KAUM–I. 90730, 34.0 mm SL, Naikina
Port, Nagamine, Wadomari; KAUM–I. 122723, 46.5 mm SL, off Yakomo, China.
Ctenochaetus hawaiiensis Randall 1955
[Jpn name: Meikyusazanamihagi]
Underwater photograph (Pl. 9-H).
Ctenochaetus sp.
[Jpn name: N/A]
KAUM–I. 1986, 35.4 mm SL, off Okinoerabu Island; KAUM–I. 14630, 34.6 mm SL, off Okinoerabu Island;
KAUM–I. 14631, 29.7 mm SL, off Okinoerabu Island.

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Ctenochaetus striatus (Quoy & Gaimard 1825)[Jpn name: Sazanamihagi]
Yoshigou et al. (2005a): HMNH 7984, Nishibaru, Wadomari; HMNH 7985, Nishibaru, Wadomari; HMNH
7986, Nishibaru, Wadomari. This study: KAUM–I. 1972, 41.8 mm SL, off Okinoerabu Island; KAUM–I.
6440, 39.9 mm SL, off Okinoerabu Island; KAUM–I. 6441, 38.8 mm SL, off Okinoerabu Island; KAUM–
I. 6442, 38.1 mm SL, off Okinoerabu Island; KAUM–I. 6443, 40.1 mm SL, off Okinoerabu Island; KAUM–
I. 6444, 40.4 mm SL, off Okinoerabu Island; KAUM–I. 6445, 34.7 mm SL, off Okinoerabu Island; KAUM–
I. 6446, 37.4 mm SL, off Okinoerabu Island; KAUM–I. 6447, 36.8 mm SL, off Okinoerabu Island; KAUM–
I. 6448, 40.5 mm SL, off Okinoerabu Island; KAUM–I. 6449, 37.2 mm SL, off Okinoerabu Island; KAUM–
I. 6450, 37.1 mm SL, off Okinoerabu Island; KAUM–I. 6451, 38.8 mm SL, off Okinoerabu Island; KAUM–
I. 6452, 34.7 mm SL, off Okinoerabu Island; KAUM–I. 6453, 39.8 mm SL, off Okinoerabu Island; KAUM–
I. 6454, 37.8 mm SL, off Okinoerabu Island; KAUM–I. 6455, 37.4 mm SL, off Okinoerabu Island; KAUM–
I. 6456, 36.3 mm SL, off Okinoerabu Island; KAUM–I. 6457, 39.2 mm SL, off Okinoerabu Island; KAUM–
I. 6458, 37.6 mm SL, off Okinoerabu Island; KAUM–I. 6459, 35.2 mm SL, off Okinoerabu Island; KAUM–
I. 6460, 37.1 mm SL, off Okinoerabu Island; KAUM–I. 6461, 36.2 mm SL, off Okinoerabu Island; KAUM–
I. 6462, 30.4 mm SL, off Okinoerabu Island; KAUM–I. 6463, 35.3 mm SL, off Okinoerabu Island; KAUM–
I. 6464, 36.0 mm SL, off Okinoerabu Island; KAUM–I. 6465, 32.0 mm SL, off Okinoerabu Island; KAUM–
I. 6466, 36.8 mm SL, off Okinoerabu Island; KAUM–I. 6467, 32.6 mm SL, off Okinoerabu Island; KAUM–
I. 90039, 142.0 mm SL, Okidomari Port, Shimojiro, China.
Naso brachycentron (Valenciennes 1835)[Jpn name: Onitenguhagi]
Sakurai (2018c): photo only, off Okinoerabu Island.
Naso lituratus (Forster 1801)[Jpn name: Miyakotenguhagi]
KAUM–I. 90340, 243.0 mm SL, off Okidomari Port, Shimojiro, China; KAUM–I. 121897, 84.7 mm SL,
Okidomari Port, Shimojiro, China.
Naso thynnoides (Cuvier 1829)[Jpn name: Bozuhagi]
Sakurai (2018c): KAUM–I. 90005, 245.9 mm SL, north of Okinoerabu Island. Sakurai (2019c): KAUM–
I. 90005, 245.9 mm SL, north of Okinoerabu Island. This study: KAUM–I. 90005, 245.9 mm SL, north of
Okinoerabu Island.
Naso unicornis (Forsskål 1775)[Jpn name: Tenguhagi]
Sakurai (2019c): KAUM–I. 121881, 120.7 mm SL, Okidomari Port, Shimojiro, China. This study: KAUM–
I. 121875, 253.5 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 121881, 120.7 mm SL, Okidomari Port,
Shimojiro, China.
Paracanthurus hepatus (Linnaeus 1766)[Jpn name: Nan-yohagi]
Sakurai (2019c): KAUM–I. 122706, 53.6 mm SL, off Yakomo, China. This study: KAUM–I. 122706, 53.6mm SL, off Yakomo, China; KAUM–I. 122724, 146.1 mm SL, off Yakomo, China; KAUM–I. 122725,
157.0 mm SL, off Yakomo, China; KUN-P 49102, 51.2 mm SL, Yakomo Beach, Yakomo, China; KAUM–I.
132179, 53.6 mm SL, west of Taminazaki, Tamina, China.

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Prionurus scalprum Valenciennes 1835
[Jpn name: Nizadai]
Sakurai (2019c): KAUM–I. 121873, 340.2 mm SL, Inobe Port, Inobe, Wadomari. This study: KAUM–I.
121873, 340.2 mm SL, Inobe Port, Inobe, Wadomari.
Zebrasoma scopas (Cuvier 1829)[Jpn name: Gomahagi]
Sakurai (2018c): KAUM–I. 90645, 20.6 mm SL, Naikina Port, Nagamine, Wadomari. Sakurai (2019c):
KAUM–I. 90645, 20.6 mm SL, Naikina Port, Nagamine, Wadomari. This study: KAUM–I. 90037, 154.3 mm
SL, Okidomari Port, Shimojiro, China; KAUM–I. 90040, 23.9 mm SL, Kasaishi Beach, Kibiru, Wadomari;
KAUM–I. 90579, 123.6 mm SL, Okidomari Port, Shimojiro, China; KAUM–I. 90645, 20.6 mm SL, Naikina
Port, Nagamine, Wadomari.
FAMILY SPHYRAENIDAE
Sphyraena barracuda (Edwards 1771)[Jpn name: Onikamasu]
Senou (2018): KAUM–I. 101614, 729.0 mm SL, Kunigami, Wadomari. Senou (2019b): KAUM–I. 101614,
729.0 mm SL, Kunigami, Wadomari. This study: KAUM–I. 101614, 729.0 mm SL, Kunigami, Wadomari;
KAUM–I. 101615, 512.0 mm SL, Kunigami, Wadomari.
Sphyraena helleri Jenkins 1901
[Jpn name: Hosokamasu]
Senou (2019b): KAUM–I. 122320, 425.0 mm SL, off Okidomari Port, Shimojiro, China. This study:
KAUM–I. 65293, 435.0 mm SL, off Okinoerabu Island; KAUM–I. 65294, 426.0 mm SL, off Okinoerabu
Island; KAUM–I. 122319, 440.2 mm SL, off Okidomari Port, Shimojiro, China; KAUM–I. 122320, 425.0mm SL, off Okidomari Port, Shimojiro, China; KAUM–I. 122321, 440.5 mm SL, off Okidomari Port,
Shimojiro, China.
FAMILY SCOMBRIDAE
Acanthocybium solandri (Cuvier 1832)[Jpn name: Kamasusawara]
Inaba (2018): KAUM–I. 90000, 596.8 mm SL, south of Okinoerabu Island. Inaba (2019): KAUM–I. 90000,
596.8 mm SL, south of Okinoerabu Island. This study: KAUM–I. 90000, 596.8 mm SL, south of Okinoerabu
Island; KAUM–I. 122661, 811.0 mm SL, off Naikina Port, Nagamine, Wadomari.
Grammatorcynus bilineatus (Rüppell 1836)[Jpn name: Nijosaba]
Underwater photograph (Pl. 10-A).
Gymnosarda unicolor (Rüppell 1836)[Jpn name: Isomaguro]
Underwater photograph (Pl. 10-B).
Katsuwonus pelamis (Linnaeus 1758)[Jpn name: Katsuo]
KAUM–I. 122194, 434.4 mm SL, southeast of Okinoerabu Island; KAUM–I. 122682, 376.8 mm SL,
southeast of Okinoerabu Island.

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H. Motomura and K. Uehara

Scomber australasicus Cuvier 1832
[Jpn name: Gomasaba]
KAUM–I. 123524, 406.0 mm SL, 40 km east of Okinoerabu Island.
Thunnus albacares (Bonnaterre 1788)[Jpn name: Kihada]
KAUM–I. 122193, 397.7 mm SL, southeast of Okinoerabu Island; KAUM–I. 122683, 311.8 mm SL,
southeast of Okinoerabu Island.
FAMILY OSPHRONEMIDAE
Macropodus opercularis (Linnaeus 1758)[Jpn name: Taiwankingyo]
Ogawa (1937): Okinoerabu Island. Shibata (1964): Amata River and its vicinity, China; OMNH-P 1147, 31
specimens, rice paddies at Kamihirakawa, China. Yonezawa (2003): Okinoerabu Island. Hosoya (2013):
Okinoerabu Island. Yoshigou (2014): Okinoerabu Island. Wada (2019): KAUM–I. 115926, 50.2 mm SL,
Tamajiro, Wadomari; KAUM–I. 115927, 53.4 mm SL, Tamajiro, Wadomari. This study: KAUM–I. 115926,
50.2 mm SL, Tamajiro, Wadomari; KAUM–I. 115927, 53.4 mm SL, Tamajiro, Wadomari.
Remarks: This species is considered to be introduced into Okinoerabu Island in 1937 (Shibata 1964;
Yoshigou et al. 2005). Shibata (1964) reported that this species was common around Amata River, China,
although its population in Wadomari decreased due to an agricultural chemical. This species is now rare in
Okinoerabu Island and needed to be protected.
FAMILY BOTHIDAE
Bothus mancus (Broussonet 1782)[Jpn name: Mondarumagarei]
Underwater photograph (unnumbered fig. at bottom of last page).
FAMILY BALISTIDAE
Balistidae, undetermined genus and species[Jpn name: N/A]
KAUM–I. 122207, 28.3 mm SL, southeast of Okinoerabu Island.
Balistoides conspicillum (Bloch & Schneider 1801)[Jpn name: Mongarakawahagi]
KAUM–I. 90453, 144.5 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 101601, 246.4 mm SL, Kunigami,
Wadomari.
Balistoides viridescens (Bloch & Schneider 1801)[Jpn name: Gomamongara]
KAUM–I. 90004, 251.3 mm SL, north of Okinoerabu Island; KAUM–I. 121887, 195.9 mm SL, OkidomariPort, Shimojiro, China; KAUM–I. 123523, 323.0 mm SL, off Okidomari Port, Shimojiro, China.
Melichthys niger (Bloch 1786)[Jpn name: Soroimongara]
Kawama & Motomura (2018): KAUM–I. 101600, 254.0 mm SL, Kunigami, Wadomari. Matsuura (2018a):
KAUM–I. 101600, 254.0 mm SL, Kunigami, Wadomari. Matsuura (2019a): KAUM–I. 101600, 254.0 mm
SL, Kunigami, Wadomari. This study: KAUM–I. 101600, 254.0 mm SL, Kunigami, Wadomari.

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Melichthys vidua (Richardson 1845)[Jpn name: Kuromongara]
KAUM–I. 90035, 174.4 mm SL, Okidomari Port, Shimojiro, China; KAUM–I. 90364, 148.4 mm SL, Inobe
Port, Inobe, Wadomari; KAUM–I. 90365, 159.0 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 90366,
198.9 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 101602, 170.0 mm SL, Kunigami, Wadomari;
KAUM–I. 122016, 167.9 mm SL, Masana, China; KAUM–I. 122675, 196.0 mm SL, off Naikina Port,
Nagamine, Wadomari; KUN-P 48894, 182.7 mm SL, off Masana, China; KUN-P 49011, 210 mm SL, Yaja,
China.
Odonus niger (Rüppell 1836)[Jpn name: Akamongara]
KAUM–I. 101604, 180.4 mm SL, Kunigami, Wadomari; KAUM–I. 122017, 139.6 mm SL, Masana, China;
KUN-P 48893, 171.4 mm SL, off Masana, China.
Pseudobalistes flavimarginatus (Rüppell 1829)[Jpn name: Kiherimongara]
Matsuura (2019a): KAUM–I. 123463, 42.0 mm SL, Kasaishi Beach, Kibiru, Wadomari. This study: KAUM–
I. 123463, 42.0 mm SL, Kasaishi Beach, Kibiru, Wadomari.
Pseudobalistes fuscus (Bloch & Schneider 1801)[Jpn name: Isomongara]
Matsuura (2019a): KAUM–I. 123494, 225.6 mm SL, off Okidomari Port, Shimojiro, China. This study:
KAUM–I. 123494, 225.6 mm SL, off Okidomari Port, Shimojiro, China.
Rhinecanthus aculeatus (Linnaeus 1758)[Jpn name: Murasamemongara]
Matsuura (2018a): KAUM–I. 90684, 21.8 mm SL, Ujiji Beach, Ashikiyora, China. Matsuura (2019a):
KAUM–I. 90684, 21.8 mm SL, Ujiji Beach, Ashikiyora, China. This study: KAUM–I. 2046, 32.9 mm SL,
off Okinoerabu Island; KAUM–I. 90120, 166.8 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 90190,
28.5 mm SL, Yakomo Beach, Yakomo, China; KAUM–I. 90539, 24.4 mm SL, Ujiji Beach, Ashikiyora,
China; KAUM–I. 90540, 26.6 mm SL, Ujiji Beach, Ashikiyora, China; KAUM–I. 90612, 92.0 mm SL,
Naikina Port, Nagamine, Wadomari; KAUM–I. 90684, 21.8 mm SL, Ujiji Beach, Ashikiyora, China;
KAUM–I. 90685, 22.0 mm SL, Ujiji Beach, Ashikiyora, China; KAUM–I. 90686, 26.0 mm SL, Ujiji Beach,
Ashikiyora, China; KAUM–I. 90712, 78.4 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 121766, 155.9mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 122217, 154.7 mm SL, Inobe Port, Inobe, Wadomari;
KAUM–I. 122218, 109.3 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 122793, 131.4 mm SL, Kasaishi
Beach, Kibiru, Wadomari; KAUM–I. 123446, 94.5 mm SL, Kasaishi Beach, Kibiru, Wadomari; KAUM–
I. 123447, 97.8 mm SL, Kasaishi Beach, Kibiru, Wadomari; KUN-P 48906, 172.0 mm SL, Yakomo Beach,
Yakomo, China.
Rhinecanthus rectangulus (Bloch & Schneider 1801)[Jpn name: Tasukimongara]
KAUM–I. 90101, 102.8 mm SL, Wadomari Port, Wadomari; KAUM–I. 101619, 132.7 mm SL, Kunigami,
Wadomari; KAUM–I. 122677, 137.9 mm SL, China Port, Serikaku, China.
Sufflamen bursa (Bloch & Schneider 1801)[Jpn name: Musumehagi]
Matsuura (2019a): KAUM–I. 122000, 34.5 mm SL, Masana, China. This study: KAUM–I. 90375, 147.5 mm
SL, Inobe Port, Inobe, Wadomari; KAUM–I. 122000, 34.5 mm SL, Masana, China.

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Sufflamen chrysopterum (Bloch & Schneider 1801)[Jpn name: Tsumajiromongara]
KAUM–I. 90372, 125.1 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 90373, 147.0 mm SL, Inobe Port,
Inobe, Wadomari; KAUM–I. 90374, 150.5 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 101603, 154.8mm SL, Kunigami, Wadomari; KUN-P 49012, 97.5 mm SL, Yaja, China.
Sufflamen fraenatum (Latreille 1804)[Jpn name: Meganehagi]
KAUM–I. 101605, 166.7 mm SL, Kunigami, Wadomari; KAUM–I. 122676, 183.5 mm SL, off Naikina Port,
Nagamine, Wadomari; KAUM–I. 123509, 195.5 mm SL, off Okidomari Port, Shimojiro, China; KAUM–
I. 123517, 155.8 mm SL, off Okidomari Port, Shimojiro, China; KAUM–I. 123518, 168.8 mm SL, off
Okidomari Port, Shimojiro, China; KAUM–I. 123519, 155.9 mm SL, off Okidomari Port, Shimojiro, China.
Xanthichthys auromarginatus (Bennett 1832)[Jpn name: Hoshimongara]
Matsuura (2019a): KAUM–I. 122296, 125.5 mm SL, west of Taminazaki, Tamina, China; KAUM–I. 122297,
116.8 mm SL, west of Taminazaki, Tamina, China. This study: KAUM–I. 122296, 125.5 mm SL, west of
Taminazaki, Tamina, China; KAUM–I. 122297, 116.8 mm SL, west of Taminazaki, Tamina, China.
Xanthichthys caeruleolineatus Randall, Matsuura & Zama 1978
[Jpn name: Aosujimongara]
Matsuura (2019a): KAUM–I. 123510, 229.6 mm SL, off Okidomari Port, Shimojiro, China. This study:
KAUM–I. 123510, 229.6 mm SL, off Okidomari Port, Shimojiro, China.
FAMILY MONACANTHIDAE
Cantherhines dumerilii (Hollard 1854)[Jpn name: Hakuseihagi]
KAUM–I. 122302, 280.3 mm SL, west of Taminazaki, Tamina, China; KUN-P 48986, 73.0 mm SL, Kasaishi
Beach, Kibiru, Wadomari.
Cantherhines pardalis (Rüppell 1837)[Jpn name: Amimeumazurahagi]
Yoshigou et al. (2005a): HMNH 7845, Shinjo, China.
Oxymonacanthus longirostris (Bloch & Schneider 1801)[Jpn name: Tengukawahagi]
KAUM–I. 90759, 22.3 mm SL, Naikina Port, Nagamine, Wadomari.
Paraluteres prionurus (Bleeker 1851)[Jpn name: Nokogirihagi]
KAUM–I. 90752, 34.4 mm SL, Naikina Port, Nagamine, Wadomari.
Pervagor janthinosoma (Bleeker 1854)[Jpn name: Nishikikawahagi]
KAUM–I. 123555, 47.4 mm SL, Yaja, China.
Thamnaconus modestoides (Barnard 1927)[Jpn name: Kibirekawahagi]
KAUM–I. 122681, 292.9 mm SL, off Okinoerabu Island.

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FAMILY OSTRACIIDAE
Lactoria cornuta (Linnaeus 1758)[Jpn name: Kongoufugu]
Underwater photograph (Pl. 10-C).
Lactoria diaphana (Bloch & Schneider 1801)[Jpn name: Umisuzume]
Underwater photograph (Pl. 10-D).
Lactoria fornasini (Bianconi 1846)[Jpn name: Shimaumisuzume]
Underwater photograph (Pl. 10-E).
Ostracion cubicum Linnaeus 1758
[Jpn name: Minamihakofugu]
Matsuura (2018b): KAUM–I. 90682, 15.4 mm SL, Wanjo Beach, Azefu, Wadomari. Matsuura (2019b):
KAUM–I. 90682, 15.4 mm SL, Wanjo Beach, Azefu, Wadomari. This study: KAUM–I. 90541, 18.9 mm SL,
Ujiji Beach, Ashikiyora, China; KAUM–I. 90682, 15.4 mm SL, Wanjo Beach, Azefu, Wadomari; KAUM–I.
122245, 10.5 mm SL, west of Taminazaki, Tamina, China.
Ostracion immaculatum Temminck & Schlegel 1850
[Jpn name: Hakofugu]
Underwater photograph (Pl. 10-F).
Remarks: This species is very rare in Okinoerabu Island. The second author observed two individuals of
this species at a depth of 30 m off Cape Tamina in 2018; since 2018, no individuals have been found in theisland.
Ostracion meleagris Shaw 1796
[Jpn name: Kurohakofugu]
Matsuura (2018b): KAUM–I. 90438, 95.1 mm SL, Inobe Port, Inobe, Wadomari. Matsuura (2019b):
KAUM–I. 90438, 95.1 mm SL, Inobe Port, Inobe, Wadomari. This study: KAUM–I. 2043, 62.7 mm SL, off
Okinoerabu Island; KAUM–I. 90438, 95.1 mm SL, Inobe Port, Inobe, Wadomari; KAUM–I. 122166, 59.1mm SL, Beach Rock, Tokudoki, China; KAUM–I. 122170, 50.0 mm SL, Beach Rock, Tokudoki, China;
KAUM–I. 133521, 30.0 mm SL, Beach Rock, Tokudoki, China.
FAMILY TETRAODONTIDAE
Arothron hispidus (Linnaeus 1758)[Jpn name: Sazanamifugu]
KAUM–I. 90711, 111.2 mm SL, Inobe Port, Inobe, Wadomari; KUN-P 48988, 296.6 mm SL, Kasaishi
Beach, Kibiru, Wadomari.
Arothron manilensis (Marion de Procé 1822)[Jpn name: Sujimoyofugu]
Underwater photograph (Pl. 10-G).
Arothron meleagris (Anonymous 1798)[Jpn name: Mizorefugu]
Matsuura (2019c): KAUM–I. 123403, 190.8 mm SL, north of Taminazaki, Tamina, China. This study:

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KAUM–I. 123403, 190.8 mm SL, north of Taminazaki, Tamina, China; KUN-P 49089, 15.6 mm SL,
Kasaishi Beach, Kibiru, Wadomari.
Arothron nigropunctatus (Bloch & Schneider 1801)[Jpn name: Kokutenfugu]
KAUM–I. 90629, 189.6 mm SL, Naikina Port, Nagamine, Wadomari; KAUM–I. 122378, 149.7 mm SL, off
Okidomari Port, Shimojiro, China; KAUM–I. 122629, 167.9 mm SL, off Yakomo, China; KAUM–I. 123443,
166.6 mm SL, Kasaishi Beach, Kibiru, Wadomari.
Canthigaster amboinensis (Bleeker 1864)[Jpn name: Gomafukinchakufugu]
Yoshigou et al. (2005a): HMNH 7908, Okidomari, China. This study: KUN-P 48901, 96.5 mm SL, Beach
Rock, Tokudoki, China; KUN-P 48902, 69.2 mm SL, Beach Rock, Tokudoki, China.
Canthigaster axiologus Whitley 1931
[Jpn name: Hanakinchakufugu]
Yoshigou et al. (2005a, as Canthigaster coronata): HMNH 7909, Okidomari, China.
Canthigaster janthinoptera (Bleeker 1855)[Jpn name: Shiborikinchakufugu]
KAUM–I. 90740, 53.0 mm SL, Naikina Port, Nagamine, Wadomari; KAUM–I. 122032, 23.6 mm SL,
Masana, China; KAUM–I. 122373, 20.0 mm SL, off Okidomari Port, Shimojiro, China; KAUM–I. 122807,
20.3 mm SL, Sumiyoshi Port, Sumiyoshi, China; KAUM–I. 123558, 56.0 mm SL, Yaja, China; KUN-P
48979, 68.0 mm SL, Ujiji Beach, Ashikiyora, China.
Canthigaster valentini (Bleeker 1853)[Jpn name: Shimakinchakufugu]
KAUM–I. 1991, 30.9 mm SL, off Okinoerabu Island; KAUM–I. 90753, 27.3 mm SL, Naikina Port,
Nagamine, Wadomari; KAUM–I. 123431, 11.5 mm SL, Ujiji Beach, Ashikiyora, China.
Torquigener hypselogeneion (Bleeker 1852)[Jpn name: Namidafugu]
Underwater photograph (Pl. 10-H).
FAMILY DIODONTIDAE
Diodon holocanthus Linnaeus 1758
[Jpn name: Harisembon]
KAUM–I. 122215, 125.7 mm SL, Wadomari Port, Wadomari; KAUM–I. 122220, 161.0 mm SL, Inobe Port,
Inobe, Wadomari; KAUM–I. 122238, 80.6 mm SL, Wadomari Port, Wadomari; KAUM–I. 122690, 128.7mm SL, Wadomari Port, Wadomari; KAUM–I. 122691, 133.6 mm SL, Wadomari Port, Wadomari; KAUM–
I. 122692, 133.6 mm SL, Wadomari Port, Wadomari; KAUM–I. 122693, 123.6 mm SL, Wadomari Port,
Wadomari; KAUM–I. 122790, 144.9 mm SL, Kasaishi Beach, Kibiru, Wadomari; KAUM–I. 122791, 135.5mm SL, Kasaishi Beach, Kibiru, Wadomari; KUN-P 48909, 150.1 mm SL, Yakomo Beach, Yakomo, China;
KUN-P 48980, 145.1 mm SL, Kasaishi Beach, Kibiru, Wadomari.
Diodon hystrix Linnaeus 1758
[Jpn name: Nezumifugu]
KAUM–I. 90586, 290.1 mm SL, Okidomari Port, Shimojiro, China.

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Diodon liturosus Shaw 1804
[Jpn name: Hitozuraharisembon]
KAUM–I. 122060, 271.6 mm SL, Kunigami, Wadomari; KAUM–I. 122070, 184.3 mm SL, Beach Rock,Tokudoki, China; KAUM–I. 122071, 191.4 mm SL, Beach Rock, Tokudoki, China; KUN-P 48908, 144.1
mm SL, Yakomo Beach, Yakomo, China.
Acknowledgments
We are especially grateful to T. Uehara (GT Divers Okinoerabu), M. Matsuoka and D. Uyeno (Kagoshima
University), M. Matsunuma (Kindai University), Y. Kawama and his family (Okinoerabu), staff of Wadomari
Training Center (Okinoerabu) and Okinoerabu Fisheries Cooperative Association, K. Matsuura (National
Museum of Nature and Science), I. Fukuchi (University of the Ryukyus), G. Short (California Academy of
Sciences), and students, post-doc fellows and volunteers of the Kagoshima University Museum for providing identifications and their assistance with collecting fishes in the island. We greatly appreciated commentson the manuscript by G. Yearsley (Australia). This study was supported in part by JSPS KAKENHI Grant
Numbers JP19770067, JP26241027, JP24370041, JP23580259, and JP26450265; the JSPS Core-to-Core
Program: B Asia-Africa Science Platforms; the “Biological Properties of Biodiversity Hotspots in Japan”
project of the National Museum of Nature and Science, Tsukuba, Japan; “Establishment of Research and
Education Network on Biodiversity and Its Conservation in the Satsunan Islands” project of Kagoshima University adopted by the Ministry of Education, Culture, Sports, Science and Technology, Japan; and the “Island
Research” project of Kagoshima University.
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Yokosuka, Kanagawa Prefectural Museum of Natural History, Odawara, and National Museum of Nature andScience, Tsukuba.
Uejo, T. 2018b. Cichlidae. P. 215 in Motomura, H., K. Hagiwara, H. Senou and M. Nakae (eds.). Identification guideto fishes of the Amami Islands, Japan. Kagoshima University Museum, Kagoshima, Yokosuka City Museum,
Yokosuka, Kanagawa Prefectural Museum of Natural History, Odawara, and National Museum of Nature andScience, Tsukuba.
Uejo, T. 2019a. Poeciliidae. P. 64 in Motomura, H., K. Hagiwara, H. Senou and M. Nakae (eds.). Identification guideto fishes of the Amami Islands in the Ryukyu Archipelago, Japan. The Minaminippon Shimbun Kaihatsu Center, Kagoshima.
Uejo, T. 2019b. Cichlidae. P. 218 in Motomura, H., K. Hagiwara, H. Senou and M. Nakae (eds.). Identification guideto fishes of the Amami Islands in the Ryukyu Archipelago, Japan. The Minaminippon Shimbun Kaihatsu Cen-

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H. Motomura and K. Ueharater, Kagoshima.
Wada, H. 2019. Osphronemidae. P. 400 in Motomura, H., K. Hagiwara, H. Senou and M. Nakae (eds.). Identification
guide to fishes of the Amami Islands in the Ryukyu Archipelago, Japan. The Minaminippon Shimbun Kaihatsu Center, Kagoshima.
Wada, H., K. Hagiwara and H. Motomura. 2018. Northernmost records of Pleurosicya coerulea (Perciformes: Gobiidae) from Okinoerabu-jima island, Kagoshima, Japan, with ecological notes. Nature of Kagoshima 45:
103–107.
Watanabe, S., J. Aoyama, M. J. Miller, S. Ishikawa, E. Feunteun and K. Tsukamoto. 2008. Evidence of population
structure in the giant mottled moray eel, Anguilla marmorata, using total number of vertebrae. Copeia, 2008:
680–688.
Wibowo, K. 2018. Pomacanthidae. Pp. 205–210 in Motomura, H., K. Hagiwara, H. Senou and M. Nakae (eds.). Identification guide to fishes of the Amami Islands, Japan. Kagoshima University Museum, Kagoshima, Yokosuka
City Museum, Yokosuka, Kanagawa Prefectural Museum of Natural History, Odawara, and National Museumof Nature and Science, Tsukuba.
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Museum of Nature and Science, Tsukuba.
Yoshida, T. 2018b. Apogonidae. Pp. 113–142 in Motomura, H., K. Hagiwara, H. Senou and M. Nakae (eds.). Identification guide to fishes of the Amami Islands, Japan. Kagoshima University Museum, Kagoshima, Yokosuka
City Museum, Yokosuka, Kanagawa Prefectural Museum of Natural History, Odawara, and National Museumof Nature and Science, Tsukuba.
Yoshida, T. 2019a. Pseudochromidae. Pp. 105–110 in Motomura, H., K. Hagiwara, H. Senou and M. Nakae (eds.).
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Shimbun Kaihatsu Center, Kagoshima.
Yoshida, T. 2019b. Apogonidae. Pp. 116–145 in Motomura, H., K. Hagiwara, H. Senou and M. Nakae (eds.). Identification guide to fishes of the Amami Islands in the Ryukyu Archipelago, Japan. The Minaminippon Shimbun
Kaihatsu Center, Kagoshima.
Yoshida, T. and H. Motomura. 2019. A yellowish-green colored juvenile of Aporops bilinearis (Serranidae) from Okinoerabu-jima island, Amami Islands, Kagoshima, Japan. Nature of Kagoshima 45: 189–191.
Yoshigou, H. 2014. Annotated checklist and bibliographic records of inland water fishes of the Ryukyu Archipelago,
Japan. Fauna Ryukyuana 9: 1–153.
Yoshigou, H., M. Ichikawa and S. Nakamura. 2005a. Catalogue of fish specimens preserved in Hiwa Museum for
Natural History (4). Hiwa Museum for Natural History Material Reports 5: 1–51.
Yoshigou, H., H. Tamura, M. Iwao and N. Shimada. 2005b. The cave fauna of Okinoerabu-jima Island, Amami
Group, Ryukyu Islands, Japan. Miscellaneous Reports of Hiwa Museum for Natural History 44: 37–59.

Marine and freshwater fishes of Okinoerabu Island

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Fig. 1. Maps of the Ryukyu Archipelago (upper) and the Amami Islands (lower). The island marked in red is
Okinoerabu Island.

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H. Motomura and K. UeharaFig. 2. Colored map of Okinoerabu Island. Green and blue indicate towns of Wadomari and China
respectively. Names of places, areas (red) and rivers (italic) used in text are shown here.

A

B

Fig. 3. Macropodus opercularis (Osphronemidae) from Okinoerabu Island (northernmost population for the
species). A, KAUM–I. 115926, male, 50.2 mm SL; B, KAUM–I. 115927, female, 53.4 mm SL.

Marine and freshwater fishes of Okinoerabu Island

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Fig. 4. Hippocampus denise (Syngnathidae) from Okinoerabu Island (all same specimen: KAUM–I. 132842,
female, 23.9 mm SL). Right photograph shows H. denise and gorgonian seafan, both being collected together.

A

C

B

D

E

Fig. 5. Hippocampus pontohi (Syngnathidae) from Okinoerabu Island. A, KAUM–I. 132428, female, 17.0 mm
SL; B, KAUM–I. 132427, male, 22.0 mm SL; C–E, underwater photographs taken by K. Uehara (specimens
not collected).

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A

H. Motomura and K. Uehara

F

G
B

H
C

I
D

J

E

Pl. 1. A, Orectolobus japonicus; B, Triaenodon obesus; C, Sphyrna lewini; D, Rhynchobatus djiddensis;
E, Neotrygon kuhlii; F, Taeniurops meyeni; G, Aetobatus ocellatus; H, Gymnothorax melatremus; I,
Rhinomuraena quaesita; J, Antennarius randalli. All photos by K. Uehara.

Marine and freshwater fishes of Okinoerabu Island

A

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F

B

G

C

H
D

I
E

Pl. 2. A, Antennatus coccineus; B, A. rosaceus; C, Monocentris japonica; D, Eurypegasus draconis; E,
Aeoliscus strigatus; F, Solenostomus cyanopterus; G, S. paradoxus; H, Doryrhamphus (Doryrhamphus)japonicus; I, Halicampus macrorhynchus. All photos by K. Uehara.

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A

H. Motomura and K. Uehara

E

F
B

G
C

D
H

Pl. 3. A, Hippocampus bargibanti; B, Phoxocampus diacanthus; C, Trachyrhamphus bicoarctatus; D,
Dendrochirus brachypterus; E, Synanceia verrucosa; F, Ablabys taenianotus; G, Pseudanthias bicolor; H, P.
hypselosoma. All photos by K. Uehara.

Marine and freshwater fishes of Okinoerabu Island

A

119

E

F

B

G

C

H

D

I

Pl. 4. A, Pseudanthias lori; B, P. pleurotaenia; C, Ostorhinchus apogonoides; D, O. aureus; E, Hoplolatilus
cuniculus; F, H. starcki; G, Pentapodus aureofasciatus; H, Chaetodon guentheri; I, C. mertensii. All photos
by K. Uehara.

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A

H. Motomura and K. Uehara

E

F

B

G

C

D

H

I

Pl. 5. A, Chaetodon meyeri; B, C. reticulatus; C, C. semeion; D, C. xanthurus; E, Hemitaurichthys polylepis;
F, Centropyge fisheri; G, C. shepardi; H, Genicanthus melanospilos; I, Oxycirrhites typus. All photos by K.
Uehara.

Marine and freshwater fishes of Okinoerabu Island

A

B

121

F

G

H
C

I

D

E

J

Pl. 6. A, Amblyglyphidodon aureus; B, A. ternatensis; C, Amphiprion ocellaris; D, A. sandaracinos; E,
Chromis albomaculata; F, C. leucura; G, C. xouthos; H, Chrysiptera tricincta; I, Microcanthus strigatus; J,
Bodianus anthioides. All photos by K. Uehara.

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A

H. Motomura and K. Uehara

F

B
G

C
H

I
D

E
J

Pl. 7. A, Cirrhilabrus exquisitus; B, C. katherinae; C, C. lunatus; D, C. melanomarginatus; E, C.
rubrimarginatus; F, Pseudocheilinus ocellatus; G, Pseudocoris aurantiofasciata; H, Scarus xanthopleura;I, Cirripectes stigmaticus; J, Plagiotremus rhinorhynchos. Photo H by H. Kobayashi; all other photos by K.
Uehara.

Marine and freshwater fishes of Okinoerabu Island

A

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F

G
B

H

I

C
J

K

D

E

Pl. 8. A, Discotrema lineatum; B, Dactylopus dactylopus; C, Amblyeleotris guttata; D, A. wheeleri; E,
Cryptocentrus nigrocellatus; F, Ctenogobiops tangaroai; G, Gobiodon aoyagii; H, G. axillaris; I, G.
prolixus; J, Lotilia klausewitzi; K, Stonogobiops yasha. Photo B by H. Minami; all other photos by K.
Uehara.

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A

H. Motomura and K. Uehara

E

B
F

C
G

D
H

Pl. 9. A, Gunnellichthys curiosus; B, Nemateleotris decora; C, N. helfrichi; D, Platax boersii; E, P.
orbicularis; F, P. teira; G, Acanthurus achilles; H, Ctenochaetus hawaiiensis. All photos by K. Uehara.

Marine and freshwater fishes of Okinoerabu Island

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E

F

B

C

D

G

H

Pl. 10. A, Grammatorcynus bilineatus; B, Gymnosarda unicolor; C, Lactoria cornuta; D, L. diaphana; E, L.
fornasini; F, Ostracion immaculatum; G, Arothron manilensis; H, Torquigener hypselogeneion. All photos
by K. Uehara.

Ichthyological Publications by the Kagoshima University Museum2009 (March)
Motomura, H. (ed.). Fish collectionbuilding and procedures manual. Japaneseedition. Kagoshima University Museum,Kagoshima. 70 pp.

2010 (March)
Motomura, H. and K. Matsuura (eds.).
Fishes of Yaku-shima Island - A World
Heritage island in the Osumi Group,
Kagoshima Prefecture, southern Japan.
National Museum of Nature and Science,
Tokyo. viii + 264 pp., 704 figs.

2011 (Nobember)
Matsunuma, M., H. Motomura, K.
Matsuura, N. A. M. Shazili and M. A.
Ambak (eds.). Fishes of Terengganu east coast of Malay Peninsula, Malaysia.
National Museum of Nature and Science,
Tokyo, Universiti Malaysia Terengganu,
Terengganu, and Kagoshima University
Museum, Kagoshima. viii + 251 pp., 678
figs.

2013 (February)
Motomura, H. and S. Ishikawa (eds.). Fishcollection building and procedures manual.
English edition. Kagoshima University
Museum, Kagoshima and Research Institutefor Humanity and Nature, Kyoto. 70 pp.

2013 (March)
Motomura, H., S. Dewa, K. Furuta and K.
Matsuura (eds.). Fishes of Iou-jima andTake-shima islands, Mishima, Kagoshima,
Japan. Kagoshima University Museum,Kagoshima and National Museum of Natureand Science, Tsukuba. 390 pp., 883 figs.

2013 (September)
Yoshida, T., H. Motomura, P. Musikasinthorn and K . Matsuura (eds.). Fishesof northern Gulf of Thailand. National
Museum of Nature and Science, Tsukuba,
Research Institute for Humanity and Nature,
Kyoto, and Kagoshima University Museum,Kagoshima. viii + 239 pp., 621 figs.

2014 (March)
Motomura, H. and K. Matsuura (eds.). Field
guide to fishes of Yoron Island in the middleof the Ryukyu Islands, Japan. Kagoshima
University Museum, Kagoshima andNational Museum of Nature and Science,
Tsukuba. 648 pp., 1808 figs.

2016 (July)
Iwatsubo, H., S. Kato and H. Motomura(eds.). Field guide to fishes of Ei insouthern Kyushu, Japan. Kagoshima
Museum of Aquatic Biodiversity, Makurazaki, Kagoshima University Museum,Kagoshima, and Seahorse Ways, Minamikyushu. 80 pp., 192 figs.

Most publications listed above can be downloaded fromhttp://www.museum.kagoshima-u.ac.jp/staff/motomura/dl.html

Ichthyological Publications by the Kagoshima University Museum2016 (August)Matsunuma, M., Y. Fukui and H. Motomura(eds.). Freshwater fishes of Kagoshima
City, southern Kyushu, Japan. Kagoshima
University Museum, Kagoshima. 86 pp.,
221 figs.

2016 (September)
Koeda, K., Y. Hibino, T. Yoshida, Y.
Kimura, R. Miki, T. Kunishima, D. Sasaki,
T. Furukawa, M. Sakurai, K. Eguchi, H.
Suzuki, T. Inaba, T. Uejo, S. Tanaka, M.
Fujisawa, H. Wada and T. Uchiyama.
Annotated checklist of fishes of Yonagunijima island, the westernmost island inJapan. Kagoshima University Museum,Kagoshima. v + 120 pp., 523 figs.

2017 (January)
Motomura, H., U. B. Alama, N. Muto, R.
Babaran, and S. Ishikawa (eds.). Commercial and bycatch market fishes of Panay
Island, Republic of the Philippines.
The Kagoshima University Museum,Kagoshima, University of the Philippines
Visayas, Iloilo, and Research Institute for
Humanity and Nature, Kyoto. 246 pp., 911
figs.

2017 (February)
Motomura, H. and S. Harazaki. Annotatedchecklist of marine and freshwater fishesof Yaku-shima island in the Osumi Islands,
Kagoshima, southern Japan, with 129
new records. Bulletin of the Kagoshima
University Museum, 9: 1–183.

2017 (October)
Iwatsubo, H. and H. Motomura (eds.).
Field guide to fishes of Kagoshima Bay
in southern Kyushu, Japan. Kagoshima
Museum of Aquatic Biodiversity, Kagoshima and the Kagoshima University
Museum, Kagoshima. 302 pp., 618 figs.

2017 (December)
K i m u r a , Y. , Y. H i b i n o , R . M i k i , T.
Minetoma and K. Koeda (eds.) Field guideto fishes of Kuchinoerabu-jima island in
the Osumi Group, Kagoshima, southern
Japan. Kagoshima University Museum,Kagoshima. 200 pp.

2018 (March)
Koeda, K., H. Hata, M. Yamada and
H. Motomura (eds.). Field guide tofishes landed at Uchinoura Fishing Port,
Kagoshima, Japan. Kagoshima University
Museum, Kagoshima. 520 pp., 1059 figs.

2018 (April)Motomura, H., K. Hagiwara, H. Senou
and M. Nakae (eds.). Identification
guide to fishes of the Amami Islands,
Japan. Kagoshima University Museum,Kagoshima, Yokosuka City Museum,
Yokosuka,Kanagawa Prefectural Museumof Natural History, Odawara, and National
Museumof Nature and Science, Tsukuba.
414 pp., 3290 figs.

Most publications listed above can be downloaded fromhttp://www.museum.kagoshima-u.ac.jp/staff/motomura/dl_en.html

Ichthyological Publications by the Kagoshima University Museum2018 (September)
Mochida, I. and H. Motomura. An annotatedchecklist of marine and freshwater fishes ofTokunoshima island in the Amami Islands,
Kagoshima, southern Japan, with 202
new records. Bulletin of the Kagoshima
University Museum, 10: 1–80.

2020 (February)
Sonoyama, T., K. Ogimoto, S. Hori, Y.
Uchida and M. Kawano. An annotatedchecklist of marine fishes of the Sea ofJapan off Yamaguchi Prefecture, Japan, with74 new records. Bulletin of the Kagoshima
University Museum, 11: 1–152.

2019 (February)
Motomura, H., K. Hagiwara, H. Senou and
M. Nakae (eds.). Identification guide tofishes of the Amami Islands in the Ryukyu
Archipelago, Japan. The Minaminippon
Shimbun Kaihatsu Center, Kagoshima. 436
pp., 3370 figs.

2020 (March)
Motomura, H. and K. Uehara. An annotatedchecklist of marine and freshwater fishes of
Okinoerabu Island in the Amami Islands,
Kagoshima, southern Japan, with 361
new records. Bulletin of the Kagoshima
University Museum, 12: 1–125.

Bothus mancus (Bothidae) off Okinoerabu Island (photo by K. Uehara)A total of 637 species, including 361 species that represent
the first reliable records from Okinoerabu Island on thebasis of collected specimens and/or underwater photographs,
are listed with citation of literature, registration numbers,
sizes, localities on the island, and some remarks.